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1933 PHYTOLOGIA ios, 


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STUDIES OF NEW AND NOTEWORTHY TROPICAL AMERICAN PIANTS --- I 


Harold N. Moldenke 


Among the recent plant collections of G. Klug, B. A. 
Krukoff, A. E. Lawrance, Y. Mexia, and E. J. Valeur in trop- 
ical America which have come to the New York Botanical Gard- 
en for identification, there have been found a number of new 
and noteworthy species, some of which will be very briefly 
discussed on the following pages. The type specimens of all 
new species herein described are deposited in the herbarium 
of the New York Botanical Garden. All of the Lawrance plants 
mentioned hereinafter were collected in the western part of 
the department of Boyaca, Colombia, 50--100 km. northwest of 
Bogota, in 1932, 

Langsdorffia hypogaea Mart. -- A number of complete 
plants collected June 19 in a thin forest, altitude about 
1075 m., lewrance 251. The collector describes this exceed- 
ingly scarce plant as 5—-8 cm. tall with red non-odorous 
flowers, and remarks that it is very rare where found. 

Celosia argentea var. linearis Sw. -- Flowering stems 
and roots of this very attractive plant collected Jume 17 in 
a high forest front, altitude about 2100 m., lawrance 235. 
Field notes indicate the plant as about 1 m. tall, witha 
stemdiameter of 1,5--% cm. and purple slightly odorous 
flowers. 

Raimondia quinduensis var. latifolia R. E. Fries -- 
Flowering stems collected June 7 in thick forest fringes, 
altitude about 1075 m., Lawrance 170. It is described by the 
collector as a shrub 2,.5--3 m tall, with a stemdiameter of 
about 5.5 cm. and dirty creamcolored non-odorous flowers. 

GYNANDROPSIS HIRSUTA sp. nov. Frutex 1--1.5 m altus; 
caulibus simplicibus vel paullum furcatis, superne et prae- 
sertim ad nodos hirsutis, inferne glabrescentibus; innova- 
tionibus dense purpureo-hirsutis; petiolis 1.5--19 cm. 
longis, summis brevissimis, juventute purpureo-hirsutis, 
senectute parciore vel sparse hirsutis, apice plerumque per- 
sistente hirsuto; foliis trifoliolatis vel summis simplici- 
bus; petiolulis centralibus 2.5--10 mm. longis dense pur- 
pureo-hirsutis; petiolulis lateralibus ad 3 mm longis vel 
obsoletis; foliolo centrali elliptico 9.5--17 cm. longo, 3-- 
5.2 cm. lato integerrimo acuminato, ad basin longe attenuato 
vel cuneato, utrinque dense hirsuto (praesertim subtus et 
juventute), pilis maturis 2--3 mm. longis et 1m, dissitis, 
juventute perspicue purpureis, subtus in costa et venis 
secundariis densissimis; foliolis lateralibus plusminus 


5 


6 Pl Yo? 0: 0: Gatce No. 1 


asymmetricis in omnibus aliis notis foliolo centrali con- 
similibus; paniculis 10--16 em. longis sparse hirsutis; 
bracteis ut videtur nmullis (vel caducis et cicatricium opp- 
ositorum vel suboppositorum paria 20 plusve in rhachide re- 
linquentibus); floribus ut videtur ad terminatioem rhachid- 
is 10--20 subremotis laxe patentibus; pedicellis filiform 
ibus ad 2 cm. longis sparse hirsutis; sepalis 4 ovatis ca. 7 
mm. longis et ad basin 2.4 mm. latis, ad apicem argute acu- 
tis, dorso plusminus carinatis et sparse hirsutis; petalis 4 
rubris vel coccineis ovato-ellipticis ca. 9 mm longis et 
5.2 mm. latis obtuse acutis glabris; toro in glandulam 
longam binariam secundum androphorum positam prolongato; 
androphoro crasso ca. 12 mm longo et ad basin 1.3 mm. lato 
glabro; staminibus 6 apice androphori emergentibus et gyno- 
phorum circumvallantibus; filamentis crassis ca. 5 mm. 
longis glabris; antheris elongatis lineari-oblongis ca. 5.2 
mn. longis et 2 mm. latis; gynophoro brevissimo ca. 0.4 m, 
longo; ovario oblongo ca. 2 mj. longo, ad apicem truncato; 
stylo nullo. 

Type, lawrance 73, collected May 18 along the side of a 
brook in a thick forest, altitude about 1050 m, in the Mt. 
Chapon region. The species is related to G. brachycarpa DC., 
from which its 3-foliolate leaves and characteristic purple- 
hirsute pubescer.ce at once separate it. The collector des- 
cribes the root-system of the type specimen as "very dainty? 
Lawrance 2533 collected at an altitude of 2100 m. in the 
Alto Chapon region, June 17, differs from the type only in 
its much denser and more pronouncedly purple pubescence. 
Concerning the latter specimen the collector says: "A de- 
lightful plant -- would make a fine garden plant." 

DIOCLEA PULCHRA sp. nov. Liana alte scandens; ramlis 
junioribus plusminus dense longeque pilosis vel hirsutis, 
pilis brumeis; stipulis trianguleri-ovatis ca. & mm longis 
villosis, margine p lerumque argute incisis; petiolis 14,5-- 


=) "ET oO Olle longis dense brunneo-villosis; rhachide 4--4.5 cm. 


supra foliola lateralia producta, petiolo consimili; sti- 
pellis subulatis 1,5--2 mm. longis dense villosis; petiolu- 
lis crassiusculis 8--9 mm. longis dense villosis; foliolo 
centrali elliptico 13--14 cm. longo 7—~8.5 om. lato symmet- 
rico, ad basin rotundato, margine integerrimo in siccitate 
leviter revoluto, ad apicem in cuspidem ca. 1 cm. longam 
acuminato, supra densiuscule subtus dense strigoso-villosis, 
venis secundariis utroque latere 10--12 subrectis ad margin- 
em arcuatis subtus valde prominentibus; foliolis lateralibus 
foliolo centrali consimilibus sed perspicue asymmetricis; 
inflorescentiis axillaribus singulis validis cernuis ad an- 
thesin ca. 23 cm. longis densifloris, glomerulis florigeris 
subsessilibus ca. 18 cm. longitudinis racemi possidentibus; 
calyce firme membranaceo extra breviter piloso intus glabro, 
tubo ca. 6 m. longo, lobo superiore late ovato ca. 7 mm, 


195) Moldenke, Tropical American Plants 7 


longo et 10 mm. lato obtuso, dorso sellaformi, inferiore an- 
guste ovato vel lanceolato ca. 12 um. longo acuminato, lobis 
lateralibus falcato-ovatis ca. 7 mm. longis acutissimis; 
vexilli limbo ca. 14 ma. longo et 18 mm lato, ad apicem e- 
marginato, supra basin cartilagineo-incrassato, ad basin 
valde auriculato, ungue curvato ca. 5 mm longo et 1.5 m, 
lato; alis ovato-oblongis, limbo ca. 14 mn, longo et 10 m. 
lato, ad apicem obtuso, ad basin subtruncato in unguem ca. 

8 ma. longum cuneato, margine integro; carinae limbo ca. 15 
mm. longo et 10 m, lato, ad apicem profunde 4~lobato, ungue 
ca. 6 mm longo; staminibus usque ad 7 mm. monadelphis, 
supra diadelphis, alternatim longioribus et stylum aequant- 
ibus; stylo ca. 20 mm. longo, usque ad 15 m,. longitudinis 
densissime longeque strigoso-villoso, superne glabro; ovario 
sesgili. 

Type, Lewrance 528, collected in a high thick forest, alt- 
itude about 925 m, in the El Umbo region, October 12. The 
plant is described by the collector as a creeper attaining a 
height of 100--120 feet and a stem—diameter of 2--4 inches, 
with sticky sap and odorous flowers which are dark helio- 
trope with a y ellow blotch at the top of the throat. The 
species is a member of the section Pachylobium Benth. and is 
related to D. coriacea (Grah.) Rusby and D. densiflora Hu- 
ber. The former differs pronouncedly in its long linear 
bracts, its mostly glabrous upper leaf-surfaces, and its 
ve lutinous-tomentose pubescence on branchlets, peduncles, 
petioles, and lower leaf-surfaces. The latter differs in its 
semisagittate stipules, shorter petioles, longer secondary 
stipules, erect racemes which are flower-bearing almost to 
the base, and its log linear-subulate and ciliate bracts. 

MUCUNA HOLTONI (Kuntze) comb. nov. Stizolobium Holtoni 
Kuntze, Rev. Gen. Pl. 1: 207. 181. -- Flowering and fruit- 
ing stems collected May 13 in impassable thickets of a low 
forest, altitude about 1075 'm., in the Chapdon region, lew- 
rance 44, The collector describes the plant as a creeper 6-- 
10 m. in length, with a stem-diameter of 1.5--3 cm. and yel- 
low non-odorous flowers; “very rare hereabouts." The genus 
Mucuna as recognized by Bentham & Hooker and by Engler & 
Prantl clearly consists of two distinct genera, as pointed 
out by David Prain [Journ. As. Soc. Bengal, new series, 66: 
404. 1897] and by K. S. Bort |U. 3S. Dept. Agr. Bur. Pl. Ind. 
Bull. 141: 31. 1909]. According to these authors, the genus 
Mucuna was first published by Adanson [Fam. Pl. 2: 325. 
1763] and the type species is Mucuna urens (L.) DC., while 
the genus Stizolobium was first published by Patrick Browne 
(Hist. Jam. 290. 1756] and the type species is Stizolobium 
pruriens (L,) Pers. The two genera are easily distinguished 
by the seed. In Stizolobium the hilum is linear, elevated, 
and oblong-crateriform, extending from one-fifth to nearly 
one-fourth the circumference of the seed, and germination is 


8 PHYTO LCG as No. 1 


not hypogeous. In Mucunea [in the restricted sense] the hilum 
is mich elongated and band-like, extending nearly all the 
way around the seed, and germination is hypogeous. According 
to these criteria, our species is very plainly a member of 
Mucuna. 

Roucheria humiriifolia Planch. -- Flowering stems col- 
lected in December, 1929, in a forest at Mishuyacu, near 
Iquitos, altitude about 100 m., Loreto, Peru, Klug 683. It 
is described by. the collector as a tree 22 m tall and with 
orange flowers. The species was known previously from French 
Guiana. 

CASTELARIA GALAPAGEI (Hook. f.) comb. nov. Castela gal- 
apagei Hook. f., Trans. Lim. Soc. 20: 229. 1851, (1) 

I1EX UTILIS sp. nov. Arbor; ramlis glebris; petiolis 
crassis 11--12 mn. logis glabris in siccitate perrugatis; 
laminis membranaceis oblongis 8.5--14.5 em. longis, 3.2=-5.5 
em. latis, ad anicem acuminatis, margine fere usque ad basin 
obtuse dentato vel .sinuato in siccitate subrevoluto, ad bas- 
in cuneatis, supra glabris nitidisque, subtus glabris ob- 
scurisque, venis venulisque utroque prominulis; inflores- 
centiis glomeratis, glomerulis 4—7-floris; pedicellis 
crassiusculis cae 5 mm. longis glabris; calyce subpatelli- 
formi vel perlaxe campanulato incrassato, lobis 4 vel 5 tri- 
angulatis ca. Oe«7 mm longis et 2 m,. latis integris hyali- 
nis, ad apicem obtusis; corollae albae lobis 4 vel 5 ob= 
longo- vel subobovato-linzulatis ca. 3.8 mm. longis et 2.6 
mm. latis, ad apicem rotundatis, ad basin connatis; stam- 
inibus 4 vel 5 6 sinubus petalorum emergentibus; filamentis 
crassis ca. 1.3 mm. longis glabris; antheris triangulari- 
ovatis ca. 1.1 mm. longis et ad basin 1m. latis; ovario 
subg loboso vel ovoideo ca. 2 mm longo et lato glabro, ad 
apicem in stigmatem abeunto. 

Type, lewrance 559, collected November 4 in forest fronts, 
altitude about 1200 m., in the El Umbo region. The species 
is described by its collector as a tree 20--30 m. tall with 
a stem—diameter of 1--1.5 m., whose wood is used extensively 
by the natives in hut construction. It superficially re- 
sembles I, guianensis (Aubl.) Kuntze, which, however, dif- 
fers pronouncedly in its more acute leaf-b lades whose mar- 
gins are entire and whose petioles are mich shorter and not 
wrinkled and its pedunculate inflorescences and smaller 
flowers, 

Cissus rhombifolia var. glabrescens Planch. -- F lowering 
and fruiting stems collected June 28 in the Mt. Chapon 
region, altitude about 1000 m., lewrance 263. The collector 
describes the plant as rambling and growing over vegetation 
horizontally, with ite stems 1.5--4 cm. in diameter, 

"square, ribbed on the top and bottom only and with fins 
{=wings?] at all 4 edges." This specimen is placed here only 
tentatively. It does not agree well with any of the identi- 


1933 Moldenke, Tropical American Plants 9 


fied specimens of this variety available for comparison, nor 
with the description of Planchon [in DC. Monog. Phan. 5: 
544--546. 1883]. Planchon states "ramis....se.ee.non alatis" 
and "ramilis hirtis," but in our specimen both the branches 
and the branchlets are very decidedly and conspicuously 4- 
winged and glabrous. He likewise. states "foliolo intermedio 
rhombeo- vel subrhombeo-ovato v. ovato," but in our speci- 
men the central leaflets are elliptic-oblong, 10--15 cn, 
long, 4--8.5 cm. wide, rather abruptly long-acuminate at the 
apex, and rounded at the base, and the two lateral leaflets 
are likewise elliptic-oblong, somewhat smaller, rather ab- 
ruptly long-acuminate at the apex, decidedly inequilateral 
and obliquely rounded at the base. The fruit is oblong- 
elongate, 10-—-14 mm. long, 4—5.5 mm. wide, and sharply 
acute to a mucronate apex. The specimen my very possibly 
represent an undescribed variety. 

VITIS NOVOGRANATENSIS sp. nov. Frutex scandens 10--13 nm. 
longus; ramis 1/2 poll. in diametro; ramlis gracilibus laxe 
breviterque villosis, pilis mollissimis et irregulariter 
dispositis; petiolis gracilibus elongatis 4--10 cm. longis 
plusminus dense villosis praesertim ad apicem; laminis mem 
branaceis oblongo-rotundatis 8,5--15 cm. longis 8--11.8 cm. 
latis, ad basin cordatis, ad apicem saepe plusminus 3- 
lobatis, lobo centrali acuminato uno et lobis lateralibus 
brevibus acuminatis 2, margine irregulariter angulato- 
dentatis (dentibus mucronatis, quoque dente venam lateralem 
unam terminante, dentibus majoribus cum venis majoribus 
occurrentibus), supra glabris vel minute ad costam et venas 
majores puberulentibus, subtus praesertim secundum costam 
et venas majores leviter villoso-pubescentibus; inflores- 
centiis vel capreolis folia oppositis et subaequantibus; in- 
florescentiis 10--19 cm. longis plusminus pyramidali- 
paniculatis ubique dense breviterque villosis, ramulis dense 
multifloris subremotis; pedunculis gracilibus 6--8 cm 
longis sparse breviterque villosis vel subglabratis, capreo- 
lum non-furcatum unum 4--6 cm. supra basin gerentibus; ped- 
icellis gracillimis ca. 1 mm longis villosis; floribus vir- 
idantibus suaveolentibus; calyce parvissimo plusminus pat- 
elliformi ca. 1 mm in diametro et 0.4 mm, longo glabro, 
margine subtruncato vel irregulariter undulato; disco plano 
ca. 0.8 mm. in diametro; corolla glabra, petalis 5 praeter 
basin coherentibus ca. 2 mm. longis, apicibus petalorum in- 
volutis ut corolla calyptroides, deorsum visa, habitum 4- 
umbonatum assumat, a disco sub anthesi decidua, petalis tum 
ad 2/3 longitudinis fissis et senectute partibus libris ex- 
trinsecus volventibus; staminibus 5 aequalibus in discum 
insertis; basibus filamentorum filiformium hyalinoruwm ca. 
0.8 mm. longorum stricte in canaliculos ovarii adpressis; 
antheris ovato-ellipticis prope basin percordatam amplis- 
simis, ad apicem emarginatis, ca. 0.5 mm longis et 0.4 m. 


10 Pe ¥°T0 LOO eA No. 1 


latis 2-locularibus; ovario subgloboso plusminus tetragono 
ca. Of mm. longo et lato 4-lobato glabro; stylo brevissimo 
crasso cae 0-5 mm. longo truncato-conico glabro plusminus 
striato; stigmate terminali ca. 0.2 mm. lato levi. 

Type, lawrance 162, collected June 6 at the edge of a 
thick low forest, altitude about 1075 m., in the central 
Chapoén region. The species seems to be related to the North 
American V. cordifolia Michx. and pessibly V. aestivalis 
Michx. It is quite distinct from the common tropical Ameri- 
can V. tiliaefolia H. & B. and V. caribaea DC., which dif- 
fer pronouncedly in pubescence, leaf-shape, and general 
habit, but most especially in the character and abundance 
of the pubescence on the leaves. 

TRIUMFETTA BOYACANA sp. nov. Frutex 3m altus; ramlis 
et sarmentis teretibue dense (praesertim in innovationibus) 
breviterque tomentosis, pilis fuscis; petiolis gracilibus 
1.5--4 cm. longis dense breviterque tomentosis; laminis ova- 
tis supra atroviridis, subtus pallidioribus, 8--12 cm. 
longis 4,5--6.5 cm. latis, ad apicem acuminatis, ad basin 
rotundatis, secundum marginem copiose et irregulariter 
"ile ese oan naming ima basi glandulis magnis sessilibus 

-6 ornatis, supra densiuscule stellatis praesertim ad cos= 
tam et venas secundarias, subtus densissime ste llato-tomen- 
tosis; venis secundariis 4 (magnis 2 et parvis 2) e basi 
folii orientibus, aliis 6 vel 8 e parte 2/3 superiori costae 
in paribus approximatis divergentibus, arcuatis ad marginem 
anastomosantibus; venis tertiariis multis subrectis et sub- 
parallelis; paniculis terminalibus ad 10 cm. longis brachia- 
tis sublaxe miltifloris ubique dense tomentosis; bracteis 
foliaceis linearibus vel oblongis vel angusto-ellipticis 
breve stipitatis 3.5--6.4 cm. longis 4—19 mm. latis, foliis 

consimi libus; pedicellis gracilibus 2—4 mm. longis dense 
tomentosis; sepalis 5 slongato-linearibus, ad basin et api- 
cem amplietis, petala paullum excedentibus ca. 145 mm longis 
ad basin ca. 2.6 mm latis, ad medium ca. 1.5 mm, latis et 
prope apicem ca. 2. mm latis, ad apicem obtuse acutis et 
percucullatis, dorso dente erecto subapicali plusminus 0.7 
mm. longo ornatis, extra sparse villosis, intus glabris; 
petalis 5 flavis spathulatis venosis ca. 11.5 m. longis, e 
basi conspicuiter papilliformi et longissime pilosa emerg- 
entibus, prope basin ca. 1.5 m,. latis, in 5 m. sequentibus 
longitudinis ad 2 mm. ampliatis, tum ad ca. 5 m,. abrupte 
ampliatis, ad apicem rotundatis et subcrenatis, praeter 
basin glabris; staminibus ca. 30; filamentis filiformibus 
8—9.3 mm. longis glabris; antheris oblongis flavis ca. 1 
mm. longis et 0.5 mm. latis, ad basin et apicem emarginatis, 
dorsifixis; pistillo stamina et lobos perianthi paullum ex- 
cedente; stylo crasso ca. 13 mm longo glabro; lobis stig- 
matis 5-lobati 0.2 mm. longis; ovario subgloboso ca. 1.3 m. 
longo et lato dense tomentoso 4-sulcato ut videtur 4-locu- 


1933 Moldenke, Tropical American Plants 1l 


lari; ovwuwlis 2 in quoque loculo. 

Type, Lawrance 7, collected May 7 in an open forest, alt- 
itude about 1200 m., in the Chapon region. The species is 
apparently related to T. althaeoides Lam. and T. semitriloba 
L., but differs jn its pubescence and much larger flowers. 
Its collector states that it is "rare where found" and that 
its flowers are not odorous. In the size of its flowers it 
approaches T. mollissima H.B.K., but in the latter the pub- 
escence and leaf-shape are entirely different. 

PARATHESIS RUBELLA sp. nov. Frutex 5--6 m. altus; hor- 
notinis crassiusculis dense puberulentibus glabrescentibus; 
petiolis crassis 18--25 mm longis inconspicue breviterque 
subalatis minutissime puberulentibus glabrescentibus; lamin- 
is leviter membranaceis oblogis ca. 25 cm. longis, 9.5--11 
cm. latis, ad basin subacuminatis, margine integris, ad api- 
cem perspicue acuminatis, supra glabris subnitidis, subtus 
dense glanduloso-punctatis et costa prominente nervisque 
secundariis numerosis parce breviterque pilosis; inflores- 
centiis terminalibus multifloris pyramidato-paniculatis ca. 
10 cm. latis parce puberulentibus; pedunculis crassiusculis 
cae 7 cm. longis; rhachide florifero ca. 9 cm. longo; pedi- 
cellis gracilibus 4--5 mm. longis glanduloso-puberulentibus; 
sepalis 5 angusto-triangularibus ca. 1m. longis, ad basin 
ca. 0.3 mm. latis, ad apicem argute acutis, glanduloso- 
puberulentibus; corolla rubella 5--5.4 m. longa, tubo 1 mm 
longo, lobis 5 triangulato-ovatis 4—4.3 mm longis, ad 
basin ca. 1.8 mm. latis, ad apicem acutis, utrinque dense 
pubescentibus et lineolis purpureis parallelis punctisque 
paucis pictis, senectute recurvis; staminibus 5 ad imam 
basin tubae corollae insertis; filamentis maturis ca. 2.8 
mm. longis glabris; antheris ovatis acutis 0.8--0.9 mn. 
longis, ad basin ca. 0.7 mm latis, dorso punctis 1--4 nig- 
ris pictis; stylo crasso ca. 3.8 mm. longo plusminuse piloso 
praesertim ad basin; stigmate punctiforme; ovario ovoideo 
infra glabrato, supra parce piloso; owalis paucis summis in 
placentam discoideam immersis. 

Type, Lawrance 372, collected July 31 in a high thick for- 
est along the side of a stream in the Mt. Chapon region. The 
flowers are described by the collector as pink and the plant 
as very beautiful. It is apparently related to P. Eggersiana 
Mez from Ecuador and P. serrulata (Sw.) Mex from Mexico and 
the West Indies to northern South America. The former dif-~ 
fers in its densely ferruginous-villose pubescence, its 
smaller and coarsely crenate leaf-blades which are densely 
ferruginous-tomentose on the midrib and margins and other- 
wise densely pilose beneath, and its sepals being rounded 
at the apex and villose along the margins. The latter dif- 
fers in its ferruginous-lepidote branchlets, its lepidote 
lower leaf=-surfaces, its smaller leaf~blades and flowers, 
its petals being narrowly rounded at the apex and only 


12 PHY TO LD OMe Te No. 1 


slightly punctate, and its more densely ferruginous- 
tomentellous pubescence in the inflorescence. 

Stylogyne sp. -- Fruiting stems collected June 14 in 
shade on the south front of a precipice, altitude about 1050 
m., in the Chapén region, Lawrance 220. It is described by 
its col lector as a very attractive shrub 2=-3 m. tall and 
said to be "worth propagating.” This probably represents a 
new species, but since no flowers are available, it has been 
thought best not to describe it as such. The leaves are much 
like those of 8. micans Mez, only in our plant they are much 
larger, being to 31.4 cm. long and 12.8 om. wide, abruptly 
short-acuminate at the apex, revolute along the margins, 
attenuate into the extremely short and stout petiole, and 
beautifully reticulate-veined. The inflorescence is about 
one-half as long as the subtending leaves and appears to be 
terminal, its branches being thick and glabrate. The fruits 
are borne in subumbellate clusters, on pedicels which are 
very slender and 7—11 mm. in length. 

WEIGELTIA LAWRANCEI sp nov. Frutex 2.5 m altus; ramlis 
et sarmentis gracilibus glabris; petiolis gracilibus:7--10 
mm. longis glabris sublate alatis; laminis tenuiter membran= 
aceis vel chartaceis oblongis 9--12 cm. longis 2.7--4.5 ecm - 
latis argute acuminatis, ad basin acuminatis vel tuneatis, 
margine superne remote dentato, dentibus latis obtusis 4-6 
(acuminatione ipsa integra et saepe subcurva), utrinque gla- 
bris, subtus dense punctulatis; venatione gracili et vems- 
ta; venis secundariis utrinquesecus 6 vel 7, subtus 3--6 m, 
& margine conspicuiter anastomosantibus; paniculis axillar- 
ibus solitariis numerosis. ca. 6 cm longis rectis, ramulis 
lateralibus 6 vel 7 brevibus 4--5-floris; pedunculis gracil- 
ibus 10—14 mm. longis glabris; rhachide gracili, pedunculo 
consimili, plerumque minutissime puberulente; pedicellis 
filiformibus ca. 1.5 mm longis puberulentibus; lobis caly- 
cis 4-partiti ca. 1.5 mm. in diametro patelliformis ovatis 
cae O.7 ma. longis et ad basin 0.5 m. latis obtuse acutis 
glabris; lobis corollae 4—partitee ca. 5 m. in diametro 
virido-albae ovatis ca. 1.5 m. longis et latis patentibus 
et plusminus reflexis obtusissimis, margine scarioso, in 
loco medio dense papillosis et gramlosis; staminibus 4 
rectis in centro basis loborum corollae insertis; filamentis 
cae 1 mm. longis glabris; antheris late ovatis vel triangu=- © 
latis ca. 0.5 mm. longis et latis ad basin bilobatis 2- 
locularibus; ovario non viso. 

Type, lewrance 32, collected May 9 in a thick forest, al- ' 
titude about 1075 m, in the Mt. Chapon region. The flowers 
are said by the collector to possess no odor. The species is 
apparently a member of the subgenus Euweigeltia Mez and re- 
lated to W. bogotensis Mez, which differs in its ferrugin- 
ous-tomentellous branchlets, longer petioles, larger leaves 
with very conspicuously lighter margins, and larger inflor- 


1953 Moldenke, Tropical American Plants 13 


escences which surpass the leaves. 

Echites microcalyx A. DC. -- Flowering stems collected 
May 13 in impassable thickets of a low forest, altitude 
about 1100 m., Lawrance 51. The collector describes the 
plant as a creeper 3--6 m. in Length, with a stemdiameter 
of 0.4—-0.9 cm. and yellow non-odorous flowers. The leaves 
on our plant are less pubescent than is ordinarily seen in 
this species, but in all other respects it agrees well with 
typical material. 

Odontadenia sp. -- Flowering stems and one fruit col- 
lected June 8 in thick forest fringes, altitude about 900 
me, lawrance 182. Field notes indicate the plant as a 
creeper 12=--15 m in length, with flat stems 1.5--3 cm in 
diameter and very pretty slightly odorous pink flowers which 
are fringed with heliotrope. This my represent a new spe- 
cies since it does not match any of the available herbarium 
material at this institution nor does it agree well with any 
published description. The stems are bi-alate and glabrous; 
leaves opposite; petioles elongate, 6--7.5 cm. long, glab- 
rous; blades broadly rotund, to 12 cm long and 10.5 om. 
wide, rounded to an acuminate apex (the acumination being 
about 15 m, long), entire, very deeply cordate at base, 
glabrous above, very lightly puberulent or subglabrate be- 
neath; inflorescence to 28 cm. long; peduncles 6--7 cm. 
long; pedicels slender, 2.5 cm. long; calyx about 4 m. long 
and 6 mm. wide; corolla-tube to 5 m. long, 1.5 mm. wide 
at the apex. 

FISCHERIA VIRIDIS sp. nov. Frutex scandens; ramis et 
ramulis crassis fistulatis dense brevissimeque furfuraceo- 
pubescentibus, pilis brunneis, et sparsim setosis, setis 
multicellularis usque ad 4 m. longis; petiolis crassis 1--2 
cm. longis dense brunneo-furfuraceis et plusminus setosis; 
laminis firme membranaceis oblongo-ellipticis 10-12 cm. 
longis, 5-6.5 em, latis, ad apicem in caudam 1 cm, longam 
acuminatis, margine integerrimis in siccitate plusmimus rev- 
olutis, ad basin cordatis vel subauriculatis, supra dense 
subtus densissime strigoso-setosis, pilis brevibus ad basin 
tumidis; costa venisque secundariis (utroque 6 vel 7) et re- 
ticulo venulorum subtus insigniter perspicuis et. densissime 
brunneo-furfuraceis; cymis multifloris foliam subtendentam 
subaequantibus vel paullo brevioribus; pedunculo crasso 2-— 
7-5 cm. longo dense brunneo-furfuraceo et sparsim setoso; 
pedicellis crassiusculis 1--3.5 cm. longis in pubescentia 
' pedunculo consimilibus; calycis segimentis lanceolatis ca. 6 
ma. longis ad basin ca. 1.6 mm latis in apicem acutam vel 
subacuminatam longe attenuatis, reflexis, dorso dense 
gramtloso-pubescentibus et sparsim longeque setosis, intus 
glabris; corolla campanulata viridi usque supra basin 5- 
partita, lobis subovato-oblongis ca. 18 mm. longis et 6.5 
m. latis, ad apicem acutam angustatis, margine e medio ad 


14 PHYTOLOGIA No. 1 


apicem duo latere valde crispis, perspicue reticulato- 
venosis, extra parce pilosis, pilis brevissimis ad basin 
tumidis, intus glabris; corona exteriore basi corollae arcte 
adnatis, lobis 11.5 mm. longis et latis carnosis crispis 
glabris; coronae interioris foliolis carnosis, dorso anther- 
arum adnatis, apice amliatis cucullatisque, summis incras- 
satis bullato-subpapillosis, antice in rostrum porrectum 
productis; antheris curvatis, appendicibus hyalinis ca. 0.7 
mm. summam antheris excedentibus; polliniis ca. 1.3 mm 
latis, translatoribus perbrevibus, retinaculo rhomboideo 
brunneo ca. 0-42 m. longo, mssis polliniis late obtuseque 
rotundo-lingulatis ca. 0.7 mm. longis et (ad apicem) latis; 
stigmate ca. 2.6 mm. in diametro. 

Type, Lawrance 396, collected August 8 in high forest 
fronts in the upper Chapén region. It is described by its 
collector as a creeper 75--100 feet in height, with a stem- 
diameter of 1/2--1 inch and inodorous green flowers which 
are mottled with tiger-like stripes of moss-green. The 
species is very closely related to F. columbiana Schultr. 
The latter, however, differs in the following respects: (1) 
its pubescence throughout (exclusive of the cetaa is mich 
finer and shorter gramzlose-velutinous-puberulent, (2) its 
branches and branchlets are "filiform", (3) its leaf-bases 
are merely "subcordate," (4) the secondary veins are 10--12 
in mumber on each side of the midrib and arranged close to- 
gether, (5) the cymes exceed the leaves in length, (6) the 
pedicels are slender and only to 2 cm. long, (7) the calyx- 
segments are oblong-lanceolate, to 4 mm. long, patent, and 
obtuse at the apex, (8) the corolla is subrotate, (9) the 
corolla-lobes are only to 7 mm. long, crisped on only one 
side and only to just below the apex, and densely puberu- 
lent within, (10) the exterior corona is annular and puber- 
ulent, and (11) the retinaculum is many times shorter than 
the pollen-masses, 

Ipomoea cardiosepala Meisn. -- Flowering stems collect- 
ed June 7 at the fringe of a thick forest, altitude about 
1100 m., Lewrance 167. The plant is described by the col- 
lector as a creeper 6--12 m in length, with a stem 
diameter of 0.7--1.5 cm. and deep pink non-odorous flowers; 
"very scarce hereabouts." The species was known hitherto 
from British Guiana. The sepals in our specimen are except- 
ionally long-ciliate and the leaves larger than any hither- 
to seen. 

Ipomoea denticulata (Desr.) Choisy -- Flowering stems 
collected May 17 in a thin forest, altitude about 1200 m, 
Lawrance 59. The plant is described by its collector as a 
vine about 1m. in length, with a stem-diameter of 0.8 cm. 
and white non-odorous flowers with a purple throat. Our 
specimen is much more pubescent on the stems, petioles, 
blades, peduncles, and sepals than is usually seen in this 


1933 Moldenke, Tropical American Plants 15 


species, but agrees well in all other respects, 

Ipomoea Nil (L.) Roth -- Flowering and fruiting stems 
collected June 9 in open spaces in fringes of a forest, 
altitude about 1100 m., in the Chapon region, Lawrance 193. 
Field notes indicate the plant as a rambling creeper with a 
stem-diameter of about 4 mm. and pretty deep heliotrope non- 
odorous flowers. Our plant is remarkable in having entire 
and unlobed leaf-blades, beautifully rounded to an acuminate 
apex and deeply cordate at the base. The characteristic 
narrowly-elongate sepals and the characters of the corolla 
and of the pubescence on the stems and leaves, however, 
leave no doubt as to the identity of this plante 

TASSADIA APOCYNELLA Gleason & Moldenke, sp. nov. Frutex 
scandens; ramis gracilibus striatis parce puberulis; ramulis 
gracilibus dense puberulis; petiolis pergracilibus 5--11 mm. 
longis minute puberulis vel subglabratis; laminis oblongo- 
ellipticis supra mdiam amplissimis 4.7—6.3 cm. longis, 
2.1—43 cm. latis, ad apicem acuminatis et in cuspidem cae 9 
mn. longam prolongatis, margine integris in siccitate valde 
revolutis, ad basin cuneatis, supra minutissime puberulis 
vel glabris, subtus glabris punctatisque; inflorescentiis 
paniculatis ramilos axillares 2—0-foliatos terminantibus, 
umbellis sessilibus 3--8-floris; rhachide et ramlis flor- 
iferis dense puberulis; pedicellis pergracilibus 2--3 mn, 
longis puberulis; sepalis ovato-oblongis ca. 1mm. longis et 
0-5 mm. latis, ad apicem acutis vel subacutis, margine tenu- 
ibus scariosisque, tantummodo ad basin connatis, dorso pil- 
osulis; corolla ante anthesin suburceolata ad anthesin cam 
panulata, tubo ca. 1.6 mm. longo, lobis ovato-lanceolatis 
ca. 1.8 mm longis et 0.7 mm latis valde recurvatis vel 
circinatis dense pilosis, ad apicem obtusis vel subacutis; 
coronae lobis ex parte latissima gynostegii emergentibus 
semicircularibus ca. 0.5 mm. longis et latis glabris, ut 
videtur simplicibus, ad apicem subacutis; gynostegio pyri- 
formi ca. 1.3 mm longo et lato; stigmate vix convexo cae 
0.35. mm lato; antherarum appendicibus hyalinis truncato- 
triangularibus ca. 0.1 mm longis; polliniis ca. 0.2 mm. 
latis; corpusculo lineari-oblongo ca. 0.15 mm longo brun- 
neo; massis pollinis anguste obovoideis cae 0.15 mn. longis; 
caudiculis curvatis. 

Type, lawrance 584, collected November 13 in forest 
fringes along a brookside, altitude about 1100 m., in the 
El Umbo region. The plant is described by its collector as a 
creeper 15--20 feet long, with a stem-diameter of 1/4—1/2 
inch and slightly odorous pale yellow star-shaped flowers. 
It is closely related to T. recurva Rusby, which, however, 
differs in its leaf-blades being widest at or below the 
middle, more or less pubescent=-pilose above, especially 
when immature, and non=-punctate beneath, its: longer and 
denser pubescence on branchlets, petioles, rachis, and ped- 


16 PY PO L:O-geea No. l 


icels, its calyx-lobes equaling or somewhat exceeding the 
corolla-tube, and other more minute characters of the floral 
parts. 

CERDANA GERASACANTHUS (L.) comb. nov. Cordia Gerasacanth- 
us L, Syst., ed. 10, 936. 17. -- Collected March 28-30, 
1920, in the vicinity of Montego Bay, Jamaica, W. Re Maxon & 
E. P. Killip 1441. 

Varronia guianensis Desv. — Flowering stems collected 
May 25 in a thick forest, altitude about 1200 m., Lawrance 
117. It is described by its collector as a shrub about 2 m. 
tall, with a stemdiameter of 3—6 cm. and greenish-white 
non-odorous flowers. The species was known previously from 
British Guiana. 

LYCIANTHES NOVOGRANATENSIS sp. nov. Frutex scandens; 
ramis et ramulis glabratis vel parcissime pilosis brumneis, 
innovationibus et interdum senectute ad nodos plusminus set- 
osis, pilis simplicibus vel furcatis; petiolis 2--11 m. 
longis glabratis; laminis subchartaceis oblongo-ellipticis 
6.3--12.2 cm. longis 4-7--6.2 em. latis ad apicem acuminatis, 
margine integris in siccitate subrevolutis, ad basin inae- 
qualiter acutis, supra nitidis et in siccitate nigrescentibus 
glabris, subtus pallidioribus viridibus glabratis vel ad 
costam et venas secundarias parcissime setosis; venis secund- 
ariis utroque 4--7 utrinque prominulis, venulorum reticulo 
subtus insigniter perspicuo sed mon elevato; pedicellis 7--25 
mm. longis, sub fructu incrassatis; calyce campanulato ca. 7 
mm. longo et 9 mm. lato parce piloso, margine scarioso eroso- 
truncato, dentibus 10 linearibus inter se 2.5—6 mm. in lon- 
gitudine variis constanter 0.7—1 mm. latis ad apicem ob- 
tusis, ca. 2 mm. infra marginem calycis abeuntibus; corolla 
campanulata alba, tubo cae 2.5 m. longo, lobis 5 lanceolatis 
ca. 2 cm longis et 4.5 mm. latis ad apicem subacutis, fere 
usque ad apicem membranis subhyalinis interpetalariis glabris 
conjunctis; staminibus ad oram tubae corollae insertis ae- 
qualibus; filamentis crassis aequilongis ca. 1.5 mm. longis; 
antheris ovatis cae 7.5 mm. longis, ad basin 3 m. latis et 
cordatis, ad apicem attenuatis, poris minutis apicalibus; 
stylo ca. 13 mm. longo; stigmate obtuse capitato; ovario 
ovato ca. 2.8 mm. longo et ad basin 2.5 mm lato glabro; 
calyce fructifero indurato ca. 5 nim. longo et 15 mm, lato, 
dentibus ad 11 mm. longis. 

Type, lawrance 478, collected in forest fringes at a 
streamside, altitude about 1100 m, in the El Umbo region. 
The plent is described by its collector as a woody vine 6—- 
10 feet in height, with a stem-diameter of 1/3 inch and pure 
white slightly odorous flowers whose stamens are cream 
colored. He reports further that the species flowers only at 
night. Lawrance 362 is a fruiting specimen from the Mt. 
Chapon region and is described as a vine attaining a height 
of 20—30 feet and a stem-diamter of 1/4—-1 inch. It was 


1933 Moldenke, Tropical American Plants 17 


collected in a thin. low forest at an altitude of about 1200 
m. The species is apparently a member of the subgemus Poly- 
meris (Dun.) Bitt. 

Mendoncia Lindavii Rusby — Flowering stems collected 
May 18 in an open forest, altitude about 1100 m., Lawrance 
69. Field notes indicate the plant as a creeper 6--9 m. in 
length, with a stem-diameter of 1--2.5 cm. and red non= 
odorous flowers. The species was known previously from Bo- 
livia. 

CITHAREXYLUM FRUTICOSUM var. SUBSERRATUM (Sw.) stat. 
nov. Citharexylum subserratum Sw. Prod. Veg. Ind. Occ. Yl. 
1788. 

CITHAREXYLUM INTEGERRIMUM (Kuntze) stat. nov. Citharexy- 
lon villosum var. integerrimum Kuntze, Rev. Gen. Pl. 2: 504. 
1891. 

CITHAREXYLUM MONTEVIDENSE (Spreng.) comb. nov. Ehretia 
monteviderisis Spreng. Syst. 1: 647. 1825. 

CITHAREXYLUM RIGIDUM (Briq.) stat. nov. Citharexy lum 
myrianthm var. rigidum’ Briq., Ann. Conserv. & Jard. Bot. 
Geneve 7-8: 317. 1904, -- Although the binomial Citharexylum 
rigidum appears in the article by Briquet cited above, it 
appears there merely as a herbarium name given in synonyny. 
It has therefore not been validly published hitherto. 

Citharexylum fruticosum var. villosum (Jacq.) 0. E. 
Schulz — Fruiting specimens collected May 30, 1951, ina 
semi-arid pine region at Moncion, altitude 300--400 m, dis- 
trict of Moncién, province of Monte Cristy, Dominican Repub- 
lic, Valeur 662. Field notes indicate the plant as a tree 
with white fruit and record the vernacular name "penda." 
This collection is a splendid example of the typical form of 
this variety. 

Lantana Chamissonis (D. Dietr.) Benth. & Hook. -— Flow- 
ering specimens collected March 15 and 17, 1930, at the edge 
of and in openings in cut-over woods, altitude 650--700 m., 
on Agricultural Hill of the Agricultural College lands and 
near the boundary of Sha-Sha Valley, Vigosa, state of Minas 
Geraes, Brazil, Mexia 4468 and 4474, The plant is described 
by its collector as a shrub l--1 1/2 m. tall, with flowers 
either white or red to orange and leaves with a verbena-like 
odor, and is said to be so abundant as to be a feature in 
the landscape. She reports the vernacular name "cambara." 

Petrea Schomburgkiana Schau. -- Flowering specimens 
collected in December, 1931, in an old clearing in the Ma- 
chado River region, state of Matto Grosso, Brazil, Krukoff 
1575. Field notes describe the plant as a woody vine. 

Vitex cymosa Bert. -- Flowering specimens collected in 
September, 1931, on varzia land near Cassipa, Tapajos River 
region, state of Para, Brazil, Krukoff 1259. The plant is 
described by its collector as a tree 10 m tall, witha 
stem-diameter of about 22 cm. at 1.5 m from the base; vern- 


18 PHYTOLOGIA No. 1 


acular name, "taruma."” -—- Fruiting specimens collected in 
December, 1931, on varzia land along a river shore near Tab= 
ajaza in the upper Machado River region, state of Matto 
Grosso, Brazil, Krukoff 1472. It is described as a tree 25 
m. tall and 6 dm in diameter at 1.5 m. from the base, with 
violet flowers and the vernacular name "taruman." 
Monographic studies of the genus Bouchea, carried out by 

the present writer during the past year with the cooperation 
and assistance of botanical institutios in America and in 
Europe, have convinced him of the correctness of the conclu- 
sions advanced by Myrle Grenzebach in her excellent treatise 
on this group [Ann. Mo. Bot. Gard. 15: 80--100. 1926] in re- 
gard to the distinctness of the South African genus Chasca= 
num from the genus Bouchea. This matter will be discussed in 
detail in the present writer's forthcoming monograph of the 
latter genus. In the meantime, it has been found necessary 
to publish the following new binomials: | 

BOUCHEA BOLIVIANA (Kuntze) comb. nov. Valerianodes boliv- 
jana Kuntze, Rev. Gen. Pl. 3: 254. 1898. 

CHASCANUM .INCISUM (Pearson) comb. nov. Bouchea incisa 
Pearson, Trans. S. Afr. Phil. Soc. 15: 180. 1904. 

CHASCANUM INTEGRIFOLIUM (Pearson) comb. nov. Bouchea in- 
tegrifolia Pearson, Trans. 8. Afr. Phil. Soc. 15: 179. 1904. 

CHASCANUM KROOKII (Guerke) comb. nov. Bouchea Krookii 
Guerke, Ann. Nat. Hofms. 20: 45. 1905. 

CHASCANUM IATIFOLIUM (Harv.) comb. nov. Bouchea latifolia 
Harv. Thes. Cap. 2: 57. 1863. 

CHASCANUM LONGIPETALUM (Pearson) comb. nov. Bouchea longi- 
petala Pearson in Thiselton-Dyer, Fl. Cap. 5: 199. 1901. 

CHASCANUM NAMAQUANUM (Bolus) comb. nov. Bouchea namaquana 
Bolus ex Pearson in Thiselton-Dyer, Fl. Cap. 5: 204. 1901. 

CHASCANUM SCHLECHTERI (Guerke) comb. nov. Bouchea Schlech- 
teri Guerke, Notiz. K. Bot. Gart. Berlin 3: 75. 1900. 

CHASCANUM SESSILIFOLIUM (Vatke) comb. nov. Bouchea sessil- 
ifolia Vatke, Linnaea 43: 529, 1882. 

CHASCANUM WILMSII (Guerke) comb. nov. Bouchea Wilmsii 
Guerke, Notiz. K. Bot. Gart. Berlin 3: 74. 1900. 


** KK K KK K K 


(1) For discussion of the genus Castelaria and the related 
genus Neocastela [Castela] see Small in N. Am. Fl. 25: 227 
and 240. 1911. 


* KK KK OK K OK K 


The New York Botanical Garden 


A NEW SPECIES OF MASCAGNIA FROM COLOMBIA 


C. V. Morton and Harold N. Moldenke 


MASCAGNIA DISSIMILIS sp. nov. Subg. Mesogynixa, ser. Zy- 
gandrae Liana alte scandens; rami crassi, aetate glabri, 
paullum angulati, grisei, internodiis longis, lenticellis 
nullis; lamina foliorum oblonga, ca. 20 cm longa, 8 cme 
lata, basi rotundata, apice abrupte et breviter acuminata 
(ca. 1-5 cm.), coriacea, utrinque pallido-viridis, glabra, 
Margine integra perspicue cartilagineo-incrassata, eglandu- 
lifera, nervis primariis 7 vel 8 supra obscuris, subtus 
prominentibus arcuatis infra marginem anastomosantibus, 
nervis secundariis tertiariisque supra obscuris, subtus 
prominulis reticulatis; petiolus crassus, ca. 1-5 cm. long-= 
us, 3 mm diametro, glaber, supra valide canaliculatus, 
eglanduliferus; inflorescentia paniculata axillaris, ca. 11 
cm. longa, pedunculo ca. 4 cm. longo compresso glabro, 
pedunculis racemorum usque ad 13 m. longis puberulis, 
pedunculis floriferis usque ad 4 m. longis puberulis, ped- 
icellis ca. 12 m. longis tenuibus puberulis, apice leviter 
incrassatis; flores désunt; sepala glandulifera; samarae 
glabrae; nux ovoidea, ca. 5 mm. longa, areola ventrali 
ovata, ca. 4 mm. longa, 3 mm, lata; ala leteralis basi con- 
_tinua, apice emarginata, 18--22 mm. alta, ca. 25 mm, lata, 
hyalina, membranacea, nervis promimulis; ala dorsalis semi- 
oblonga, 3--9 mm. longa, nucem milto superans; alae inter- 
mediae nullae. 

Type, A. E. Lawrance 546, collected October 21, 1932, at 
El Umbo, altitude 950 m, Boyaca&, about 130 miles north of 
Bogota, Colombia, and deposited in the herbarium of the New 
York Botanical Garden. 

The present species is most closely related to M. nervosa 
Niedenzu, but differs as indicated in the following key: 


Leaves oblong, about 20 cm. long, the margins conspicuously 
cartilaginous, the secondary nerves obscure above; 
branches and petioles very thick; mit glabrous, the 
ventral areole about 4 m,. long ‘and 3 mm. broadeceeeees 
ta Be ce xe bei alee ee OS Reh OS wih bisa & ab eels dissimilis. 

Leaves ovate, up to 15 cm. long (usually leas), the margins 
not cartilaginous, the secondary nerves conspicuously 
reticulate above; branches and petioles slender; nut 
usually puberulous, the ventral areole about 3 mm. long 
and 2 me DrOoaderecececcesseces coccccscceceede NErVOSAe 


19 


THREE NEW SPECIES FROM THE A. E. LAWRANCE COLLECTION 


Albert C. Smith 


The specimens hereinafter described were collected by A. 
E. Lawrance in 1932 in the western part of Boyaca, 50--100 
km. northwest of Bogota, Colombia, and the types are depos- 
ited in the herbarium of the New York Botanical Garden. 

PSITTACANTHUS DILATATUS sp. nov. Frutex parasiticus gle- 
ber; ramis ramlisque crassis teretibus, maturitate rugosis 
fuscescentibus, ad nodos conspicue incrassatis continuis; 
petiolis incrassatis nigrescentibus rugosis 4--8 mm longis, 
superne anguste alatis; laminis coriaceis viridibus ob long- 
is, 9--14 cm, longis, 3--5.5 cm. latis, basi attenuatis, 
apice rotumdatis, margine integris, utrinque dense stomatif- 
eris, pinnatinerviis, costa crassa utrinque prominente, 
nervis secundariis 4 vel 5 in quoque latere adscendentibus, 
utrinque elevatis; inflorescentiis 2--5 in axillis foliorun, 
plerumque 4-floris, floribus binis; ramilis inflorescentias 
stramineis subrugosis, pedunculo primario ca. 6 mm. longo, 
pedunculis secundariis 2--4 m. longis, bracteis parvis; ped- 
icellis 2—-4 mm. longis; cupula patelliformi submembranacea 
margine integra apice 2--3 mm. diametro; calyculo carnoso 
cylindrico apice crenato-serrato, sub anthesi 4--5 mi. longo 
et 2.5 mm, diametro; perigonio carnoso coccineo distaliter 
luteo, maturitate 3.5 cm. longo, ad medium 1.5--2.5 m. dia- 
metro, basi conspicue dilatato, saspe ad basin 6-lobato, 
lobis linearibus recurvatis, ca. 1.6 mm. latis, 3 mm supra 
basin intus ligulas submembranaceas deltoideas ca. 1.5 mm. 
longas gerentibus, apice subacutis, margine ventrali minute 
hamilosis; filamentis carnosis ca. 12 mm longis, prope med- 
ium perigonii adfixis, apice gracilibus; antheris dorsifix- 
is, oblongis, 3--4 mm. longis, apice productis et obtusis; 
stylo perigonium aequante, stigmate ellipsoideo papilloso. 

Type, Lawrance 108, collected May 24 in a dense tropical 
forest, altitude about 1200 m, in the Mt. Chapon region, It 
is a species related to the Amazonian P. biternatus (Hoff- 
mgg.) Blume, from which it differs by having the leaves 
larger and attenuate rather than obtuse at the base, the 
flowers in pairs rather than ternate, and the perigonium 
glabrous rather than cinereous-pulverulent. 

SECURIDACA CRISTATA sp. nov. Frutex scandens; ramis ram 
ulisque teretibus glabris vel juventute pilosulis; petiolis 
rugosis canaliculatis mox glabris 6--9 mm longis; laminis 
papyraceis viridibus glabris oblongis vel ovato-ob longis, 
10--20 cm. longis, 7--11 cm. latis, basi subcordetis vel ro- 


20 


1933 Smith, Three New Species 21 


tundatis, apice breviter acuminatis (apice ipso obtuso), 
margine integris et leviter revolutis, costa supra imressa 
subtus prominente, nervis lateralibus primariis 5--7 in” 
quoque latere arcuato-adscendentibus utrinque leviter ele- 
vatis, venulis reticulatis utrinque prominulis; inflores- 
centiis racemosis, rhachidibus 1 vel 2 in axillis foliorum 
8-—-15 mm. lorfeis minute puberulis, pedicellis mox glabris 
gracilibus 4—7 m, longis, bracteis deciduis pilosis acutis 
1.5 mm. longis subtentis; sepalis 4 exterioribus deltoideo- 
ovatis 4 mm longis et latis, utrinque minute puberulis, 
Margine membranaceis; sepalis 2 interioribus membranaceis 
miltinerviis oblongo-orbicularibus, ca. 7 mm longis et 
latis, glabris vel margine ciliatis, apice rotundatis vel 
emarginatis, unguiculo angusto; carina cucullata glabra 
ecristata 7 mm longa, basi subito angustata; petalis sup- 
erioribus spathulato-oblongis 7 mm longis, intus basi 
pilosulis; filamentis prope ad apices connatis, tubo intus 
piloso, antheris oblongis 0.8 mm. longis; ovario glabro 
anguste alato, stylo superne curvato; fructo sublignoso, 
absque ala subgloboso 22--25 mm. diametro, conspicue retic- 
ulato-venoso, ala obliqua unilaterali, ca. 20 mm. longa et 
basi 20 m. lata, venis elevatis parallelis striata. 

Type, Lawrance 174, collected June 7 on edge of a high 
forest in the Mt. Chapon region. The type bears flowers 
which are said to be yellow. Lawranoe 259 is another col- 
lection from the same locality, of which the fruits are 
above described. It is a species closely related to the 
Amazonian 8. Corytholobium A. W. Benn., from which it dif- 
fers by having the inflorescence racemose rather than sub- 
fasciculate, the pedicels only about one-third as large, 
and the filaments more highly connate. The fruit of the new 
species is more decidedly globose and bears a larger spur 
than that of 8. Corytholobium. In leaf character, S. crista- 
ta has the base subcordate or rounded rather than subacute, 
the apex short-acuminate rather than merely acute, and the 
costa impressed rather than plane. 

ESCHWEILERA SESSILIS sp. nov. Arbor pulchra glaberrima 
18-25 m. alta, trunco ad 1 m. diametro; ramlis cinereis 
subteretibus striatis versus apices lenticellatis; petiolis 
crassis nigrescentibus rugosis canaliculatis 5--8 m,. 
longis; laminis rigide coriaceis viridibus opacis oblongis 
vel ovato-oblongis, 8--ll cm, longis, 3--5 cm. latis, basi 
acutis, apice breviter acuminatis (apice ipso obtusis), 
margine cartilagineis et obsolete serrulatis, utrinque parce 
punctato-granulosis, pinnatinerviis, costa crassa utrinque 
prominente, nervis lateralibus primariis 8—10 in quoque 
latere arcuato-adscendentibus prope marginem anastomosant- 
ibus, venulis copiosissimes reticulatis utrinque elevatis; 
racemis axillaribus vel terminalibus saepe paniculas simu- 
lantibus; rhachidibus angulatis vel subteretibus lenticel la- 


22 PB YT Oek OrGer as No. l 


tis 2--8 om. longis 6--20-floris; floribus majusculis sub= 
sessilibus, pedicellis crassis ad 1 mm longis; hypanthio 
carnoso rugoso dense lenticellato sub anthesi 4—5 m. 
longo; sepalis aequalibus carnosis verrucosis ovato-= 
deltoideis, 2 m. longis, 2--2.5 mm. latis, apices obtusis, 
margine membranaceis; petalis tenuiter carnosis reticulato- 
venosis ovatis leviter falcatis, maximis 20 m. longis et 13 
mi. latis, apice rotundatis, basi angustis; androphoro ex- 
planato ca. 35 mm. longo, ligula 11--12 m, lata, galea 
carnosa 12--18 mm. diametro dense echinata, appendiculis 
anantheris imbricatis linearibus acutis 37--5 m. longis ob- 
tecta, staminibus 150—200, filamentis carnosis ca. 1.5 mm 
longis distaliter incrassatis apice gracilibus, antheris 

ob longo-ovoideis 0.5-——-0.6 mm. diametro basifixis; ovario 
semisupero (vertice rugoso conico sub anthesi supra disco 2 
mm, exstato), biloculari, septo fragili, ovulis 3 vel 4 ob- 
ovoideis sessilibus erectis. 

Type. laewrance 239, collected gune 17 in a forest, alti- 
tude about 2100 m, on Mt. Chapon. The collector notes that 
the flowers are purple with yellow stamens. It is a species 
related to the Venezuelan E. Fendleriana Miers, f rom which 
it differs by having the leaves somewhat larger, the vena- 
tion more prominent on the upper surface, and the flowers 
subsessile rather than long=pedicellate. The latter charac- 
ter is very consistent, and in addition the flower-parts, 
especially the sepals, of the new species are smaller than 
those of E, Fendleriana. 


The New York Botanical Garden 


NEW SPECIES OF COLOMBIAN MIMOSACEAE AND CAESALPINIACEAE 


Nathaniel L. Britton and Ellsworth P. Killip 


INGA LAWRANCEANA sp. nov. Arbor 7--9 m. alta; sarmentis 
glabris verrucosis; petiolo et folii rhachide glabris 4——5 
em. longis; foliolis 2-jugis;, rhachide inter bases cujusque 
jugi glandula sessilem orbicularem vel oblongem concavam 
1--1.5 m. latam gerente; foliolis chartaceis oblongis ad 

oblongo= lanceolatis 9--15 cm. longis 3——5 cm. latis, ad api- 
cem acutis vel acuminatis, ad basin angustatis, supra pal- 
lidis et glabris, subtus lucido=viridibus furfuroso0- 
papillosis, juventute puberulentibus senectute glabris; 
venis pinnatis adscendentibus, subtus prominentibus, costa 
utrinque prominente; pedunculis gracilibus puberulentibus 
2--4 cm. longis; floribus 10--15 umbellatis; bracteolis 
lineari-oblongis obtusis dense puberulentibus -ca. 2 mm. 
longis; pedicellis paene filiformibus puberulentibus 6—12 
mm. longis; calyce anguste campanulato puberulente ca. 4 m, 
longo, dentibus triangularibus acutjs ca. 1 mm longis; 
lobis corollae ca. 7 mm. longae oblongo-ovatis acutis; 
staminibus cae 3 mm. longis ad 1/3 longitudinis conjunctis. 

Type, lawrance 260, collected June 28 in a forest, alti- 
tude about 1000 m., in the Chapon region, Boyaca, Co lente 
and deposited in the herbarium of the New York Botanical 
Garden. 

MIMOSA LAWRANCEANA sp. nov. Liana gracilis usque ad 4.5 
me longa, ramis quadrangularibus glabratis; petiolus cum 
rhachidi gracilis puberulus 6--8 cm. longus, aculeis brev- 
ibus recurvatis armatus, basin versus glandulam sessilem or- 
bicurem paulo elevatam gerens; pinnae 2-jugae, rhachilla 2-- 
4.5 cm. longa, apicem versus glandula consimili notata; 
foliola 37--5 membranacea late ovata obliqua valde reticula- 
ta, supra nitidula, subtus opaca pallida et puberula, jugo 
supremo 5-—9 cm. longo, aliis multo brevioribuss; inflores- 
centia paniculata puberula et breviter aculeata, folia 
subaequans vel longior, pedunculis filiformibus 0.5-—3 cm. 
longis; flores breviter spicati pallide flavescentes, spicis 
multifloris 9-15 mm. longis; calyx minutus truncatus; cor- 
olla ca. 1.2 mn. longa 4-lobata; stamina 7 vel 8, ca. 4 m. 
longa. 

Type, lewrance 600, collected November 26, 1932, at an 
altitude of about 1250 m., in the El Umbo region of western 
Boyaca, Colombia, and deposited in the herbarium of the New 
York Botanical Garden. A common name for the species record- 
ed by the collector is "iguana." 


23 


24 PHYTOL OG ais No. 1 


SENEGALIA PODADENIA sp. nov. Liana alte scandens, cauli 
ca. 7 cm. diametro, annotinis rhachidibus foliorum et in- 
florescentia dense breviterque pilosis, annotinis aculeis 
brevibus recurvatis armatis; petiolus cum rhachidi gracilis 
aculeatus 8-~14 cm. longus, inferne et hinc inde inter juga 
glandulosus; stipitibus glandularum 1—1.5 mm, longis, 
glandulis ca. 0.5 m. latis; aculeis numerosis ca. 1 ma. 
longis; pinnae 20-—-30-jugae 2-—3.5 cme longae approximatae, 
rhachilla subfiliformi; foliola 20—-40-juga linearia approx- 
imata glabra acuta 2—-4 mm. longa ca. 0.5 mm. lata, costa 
inconspicua paulo excentrica; inflorescentia magna termin- 
alis paniculata, pedunculis ca. 15 mm. longis vel brevior- 
ibus; capitula parva globosa miltiflora; calyx canescens 
cae 1-5 mm longus, dentibus obtusis; corolla ca. 2 m. 
longa, lobis acutis; stamina numerosa ca. 4 mm. longa pall- 
ide flava. 

Type, lewrance 346, collected July 21, 1932, in a high 
thick forest, altitude about 1100 m., in the Mt. Chapén 
region of western Boyacaé, Colombia, and deposited in the 
herbarium of the New York Botanical Garden. 


STUDIES ON THE FLORA OF NORTHERN SOUTH AMERICA - XVIII 
PLANTAE LAWRANCEANAE COLOMBIANAE 
H. A. Gleason 


The following notes and descriptions are based on herbar- 
ium material collected by A. E. Lawrance in Department Boy- 
acé, Colombia, and now deposited in the herbarium of the New 

York Botanical Garden. 

HORTIA COLOMBIANA sp. nov. Arbor excelsa; ramis supremis 
crassis glabris atropurpureis longitudinaliter rugosis; fol- 
jis chartaceis oblongo-oblanceolatis subacutis basi in peti- 
olum brevem cuneatis utrinque glabris et reticulato-venulo- 
sis; inflorescentia corymbiformi verisimiliter glabra ramis 
crassis rugulosis confertis; pedicellis brevissimis crassis; 
calycis campanulati in sicco rugosi lobis depresso-semicirc- 
ularibus late rotundatis; petalis carnosis ovato-oblongis 
apice inflexis infra medium pulvinato-incrassatis et dense 
brunneo-barbatis; filamentis crassis; ovario ovoideo in sty- 
lum conicum brevem 5-sulcatum abrupte contracto. 

A tree 22-30 m. high and 6-20 dm. in diameter; upper 
branches fleshy, dark purple, longitudinally wrinkled, gla- 
brous; petioles semiterete, narrowly margined, about 15 mm. 
long; leaf-blades chartaceous, narrowly oblong-oblenceolate, 
as mch as 26 cm. long and 5.5 cm. wide, subacute, very nar- 
rowly revolute, et least when dry, long-cuneate at base, 
glabrous, all veins and veinlets prominulous and reticulate 

on both sides; inflorescence corymbiform, densely branched, 
many-flowered, its axes probably glabrous (actually covered 
with ea fungus mycelium), very stout, longitudinally wrinkled 
and bractless; pedicels stout, rugose, about 2 mm. long; 
calyx campanulate, fleshy, wrinkled when dry, its lobes 
broadly rounded and depressed-semicircular; petals fleshy, 
rose-color, ovate-oblong, 5.5 mn. long, 2.4 mm. wide, tipped 
with a linear inflexed tip 1 mm. long, pulvinate below the 
middle and densely bearded on the pulvinus with brown hairs; 
stamens 5, inserted in the lobes of the disk; filaments very 
stout, 3 mn. long; anthers oblong, 2 mm. long; disk low, ob- 
scurely 5-lobed; overy ovoid, 5-celled, abruptly narrowed 
into the short conic style; ovules 2 in each locule. 

Type, Lawrance 392, collected in thick forest in the 
Upper Chapon region, alt. 1500-1800 m. The genus Hortia in- 
cludes eight other described species, essentially Amazonian 
in their distribution.. One of them has globose heads; most 

of them have far larger leaves than our species, and in sev- 
eral the veins and veinlets are evanescent. H. colombiana 
apparently resembles H. brasiliana Vand. in most features, 


25 


iit Sage 
* a 


26 PH YT O baiee- 1. £ No. l 


but differs in its larger flowers, its petiolate leaves, and 
the absence of bracts. The extension of the generic range to 
the northern Andes is most unexpected. 

SYMPHYLLANTHUS PLICATUS sp. nov. Liana; rami lenticellis 
atris sparse notati; folia subrigida subsessilia ovato-ob- 
longa breviter cuspidata ad basin inaequileteralem rotundata 
vel subcordata, supra praeter costam puberulam glabra, sub- 
tus brevissime pilosa praecipue ad venas; venae laterales 
utrinsecus circa 7 curveto-adscendentes et arcuatim aonnect-— 
ee, supra impressse subtus valde prominentes; inflorescentia 
parva cinereo-tomentosa; sepala oblongo-ovata extra canesc- 
entia; petala sordide albida ad medium biloba, lobis repli- 
catis; ovarium dense longeque villosum. 

A vine 3-4 m. high, the younger branches densely ferrugi- 
nous-tomentulose, soon glabrescent and marked with large 
black lenticels; petioles stout, puberulous, 2-4 mm. long; 
leaf-blades subcoriaceous, obovate-oblong, 9-15 cm. long, 


4.53 om. wide, abruptly short-cuspidate, entire, inequilet- — 


eral at base, the one side rounded, the other somewhat sub- 
cordate, above glabraus on the surface and puberulent on the 
midvein, beneath inconspicuously pilosulous, especially on 
the veins; principal veins impressed above, very prominent 
beneath and with the veinlets conspicuously reticulate, the 
lateral veins about 7 on each side, curved-ascending and 
arcuately connected; inflorescence 4 em. long, cinereous- 
tomentose, the pedicels very short; sepals distinct, oblong- 
ovate, 2.5 mm. long, 1.5 mm. wide, rounded above, canescent 
on the back, dark purple within; petals distinct, erect, ob- 
ovate, 2 mn. long, 2-lobed to the middle, the lobes repli- 
cate and approximate, each 1.2 mm. wide; filaments distinct, 
2.5 mm. long; anthers broadly reniform, notched at the end, 
0.9 mn. wide, 0.7 mm. long; scales of the disk oblong, 0.3 
mn. long, obscurely emarginate; ovary concealed in a mass of 
villous: tomentum 1 mm. thick; styles 2, filiform, 2 mm.long. 

Type, Lawrance 414, from. the Alto Chapén region, alt. 
about 2100 m. It is apparently most closely related to S. 
Donnell-Smithii (Engl.) Gl., in which the leaves are more 
prominently acute and softly hirsute beneath and the petels 
spatulate, involute, and cucullate at the apex. So far as 
known to me this is the first species of the genus recorded 
from, the Colombian Andes. 

MATISIA LONGIFLORA sp. nov. Arbor ramis supremis ciner- 
eis; folia anguste oblonga vel elliptica acuminata integra 
basi inaequaliter rotundseta solemniter penninervia utrinque 
glabra reticulato-venosa; flores oppositifolii pedicello 
tenuissime brunneo-tomentello'superne sensim ampliato; calyx 
elongetus in sicco rugosus ad basin sensim angustetus tenu- 
issime tomentellus; petala lineari-spathulata obtusa extra 
tomentella intra tubum calycis rigida supra calycem reflexa 


ee 


1933 Gleason, Plantae Lawranceanse 27 


et calyce subdimidio longiorsa; staminum tubus longe exsertus 
tomentellus cruribus patulis stigma paulo excedentibus. 

A tree 12-18 m. high; upper brenches cinereous, glabrate; 
petioles slender, glabrate 25-35 mm. long; leaf-blades firn, 
narrowly oblong or elliptic, 16-27 cm. long, 6-10 cm. wide, 
acuminate, entire, inequilaterally rounded at base, glabrous 
on both sides, pinnately veined, the lateral veins 9 or 10 
on each side, arcuately connected near the margin, the vein- 
lets prominently reticulate; flowers solitary opposite the 
leaves, very thinly cinereous-tomentulose; pedicels 5-7 cm. 
long, stout, nearly straight, slightly enlarged upward; 
calyx narrowly tubular-cempanulate, leathery, 38 mm. long, 
gradually tapering to the base, rugose when dry, the erect 
lobes broadly depressed-ovate, obtuse, 4-5 mm. long; petals 
narrowly spatulate, the part within the celyx stiff, the ex- 
posed portion 2 em. long, reflexed at anthesis, rounded at 
the summit; staminal tube 6 cm. long, more or less curved to 
one side, ‘thinly tomentulose, its branches 17 mm. long, as- 
cending; loculi 6 on each arm, linéar-oblong, about 6 m. 
long; style exserted 16 mm. beyong the staminal tube, the 
stigma capitate, not lobed. 

Type, Lawrance 538, collected at the margin of a forest, 
El Umbo region, alt. about 750 m. Twelve species of this 
poorly known genus have been described. Of these, M. cord- 
ata and M. bicolor Ducke have cordate leaves; M. obliquifo- 
lia Standley bears its flowers on the trunk; in M. Dowdingii 
Sprague the leaves are pubescent beneath; in M. Castano Tr. 
& Kareat., M. alchornaefolia Tr. & Pl., M. lasiocalyx Schun., 
M. glandifera Tr. & Pl. the leaves are 3-nerved; in M. ob- 
longifolia P. & E. the calyx is split down one side; in M. 

' paraensis Huber and M. ochrocalyx Schum. the flowers are 
very much smaller. Oyr species most closely resembles the 
remaining one, M. Cornu-copiee Tr. & Pl., in which the 
leaves are 4-nerved, acute at base and much smaller than in 
ours, the pedicels longer, and the calyx strongly curved. 

‘ GODOYA (§ Rutidenthera) MAGNIFICA sp. nov. Arbor; folia 
imperipinnata 4(rerissime 5)-juga; foliola anguste oblonga 
inaequilatere grosse incurvo-dentata; inflorescentia pania- 
ulato-racemosa, rachidis basi cicatricibus bractearum de- 
lapsarum confertis notata; sepala inaequalia exteriora ante 
anthesis, interiora ad anthesin decidua; petala magna alba 
subpersistentia; stamina circa 50 mox decidua. 

A tree 15-18 m. high, the twigs not seen; leaves pinnate- 
ly compound, about 4 dm. long; petioles glabrous, about 5 
em. long; leaflets usually 9, or rarely 11, or the terminal 
leaflet deeply 2-parted, varying in shape from marrowly 
ovate to linear-oblong or nerrowly oblanceolate, the lowest 
as small as 6.5 by 2 cm., the upper as large as 20 by 5 cm., 
acute or acuminate, coarsely dentate with incurved teeth, 


28 PHY. T 0 oO:@- 1s No. l 


all except the terminal inequilateral at base, glabrous on 
both sides, conspicuously pinnately veined, the veins light- 
ly impressed above, prominent and reticulate beneath; inflo- 
rescence probably terminal, composed of numerous racemes on 
an elongate axis, glabrous; peduncle crowded with the annu- 
lar or nearly annuler scars of fallen bracts; racemes 10-15 
em. long, with mostly 10-15 flowers on pedicels 10-15 mn. 
long; buds ovoid-conic, 15 mm. long; outer 2 sepals about ~~ 
equal, broadly orbicular, 8 mm. long, imbricate; inner 5 se- 
pals orbicular, each one completely encircling the bud with 
its lateral margins overlapping; petals white, obovate-ob- 
long, 20 mm. long, 12 mm. wide, rounded and somewhat inequi- 
lateral at the summit; stamens about 50 (49 and 55 in two 
unopened buds); filaments 3-4.5 mn. long; anthers linear, 
straight, 7-11 mm. long; ovary fusiform, about 1 em. long at 
anthesis. 

Type, Lawrance 599, collected in the El Umbo region, alt. 
about 1200-1500 m., "on precipitous barranca fronts in well- 
aired positions," Although the thirteen specimens examined 
probably came from one tree, the leaves show a remarkable 
variation. The basal leaflets sre always shorter and pro- 
portionately broader. The upper leaflets are elongate, and 
either lanceolate, narrowly oblong, or oblanceolate; the 
base varies from rounded or even subcordate to obtuse or 
cuneate on the distal side of each leaflet, while the prox- 
imal side ranges from obtuse to narrowly cuneate. The term- 
inel leaflet is regularly oblanceolate, but on one leaf it 
is cleft nearly to the base into two divergent oblanceolate 
segments. The axis of the inflorescence appears to be about 
20 cm. long and its divergent branches give it a width of 
20-30 cm. The basal sterile portion (5-8 cm. long) is 
closely covered with the scars of the fallen bracts, which 
sometimes completely encircle the stem. The sepals fall 
very early. The two outer may fall even before the buds 
- have opened and at anthesis the flowers are quite asepalous. 
The stamens are declined to one side of the ovary and also 
fall early, so that the number visible in an open flower is 
much less than those counted in the buds. According to the 
collector, its local name is asuseno and its wood is used 
' for making the gears of the small native sugar-mills. 

The section Rutidanthera was separated by Planchon in 
1846 to include the then recently discovered G. splendida. 
Van Tieghem discussed the group in 1904, raised it generic 
rank under the emended spelling Rhytidanthera, and described 
two other species. Gilg, in 1925, mentioned but one species 
which he retained in the genus Godoya. Our species differs 
from the three hitherto described in the mech greater number 
of.stamens; fron G. splendida in its. straight anthers; from 
G. sulcata in its deciduous bracts, and apparently stands 
neerest to G. fragrans. 


1933 Gleason, Plantae Lawranceanse 29 


ANTHODISCUS MONTANUS sp. nov. Arbor excelsa glabra, ra- 
mis floriferis teretibus gracilibus; folia digitatim trifol- 
iata; petioli breves; leminae firmulee sessiles spathulato- 
obovatae, apice rotundatae, crenatae, ad basin longe cuneat- 
ae, inter se omnes aequales; racemi terminales foliis termi- 
nalibus paullum breviores, floribus apicem versus pedunculi 
confertis, pedicellis quam alabastris paullum longioribus; 
celyx cupuliformis patens obscurissime 5-angulatus; petala 
coriacea flava inter se cohaerentia ad anthesin decidua; 
stamine numerosissima inaequalia in alabastro in phalanges 
cohserentia ad anthesin distincta radiatim patentia; ovsrium 
subglobosum; styli 12 lineares adscendentes. 

A tree "60-70 ft. high;" flowering branches slender, gla- 
brous throughout; leaves digitately 3-foliolate; petioles 
slender, 10-15 mm. long, slightly dilated at the suit; 
leaflets equal in each leaf, sessile or nearly so, firm, 
spatulate-obovate, 4-7 cm. long, 2-3 cm. wide, rounded at 
summit, shallowly crenate above the middle, cuneate from 
above the middle to the base and at the base with slightly 
involute margins; racemes 4-6.5 cm. long, sterile below, the 
flowers crowded and corymbiform; pedicels escending, 6-8 mm. 
long; calyx spreading, 4.5 mm. in diameter, very obscurely 
S-angled; petals 5, closely coherent, falling at anthesis 
without separating, thick and leathery, yellow, 5.5 m. 
long; stamens very numerous, filaments loosely cohering in 
phalanxes before anthesis and the anthers inflexed, 
straicht, separate, and radiately spreading at anthesis, 3-6 
mn. long; anthers broadly oblong, 0.5-0.6 mm. long; ovary 
globose, nearly 2 mm. in diameter; styles 12, linear, as- 
cending and divergent, 2.5-3 mm. long, tapering to the min- 
ute terminal stigme, deciduous shortly after anthesis. 

Type, Lawrance 474, collected in thick forest in the El 
Umbo region, alt. about 1050 m. The collector notes that a 
"yéllow cap consisting of five petals completely covers the 
stamens and falls to the ground when the flower is fully 
matured." Only three species of this poorly known genus have 
been described. Of these A. trifoliatus Mey. and A. obova- 
tus Benth. have much lerger leaves, petioled leaflets, and 
elongete recemes; in the former the leaves are acuminate and 
in the latter they are entire. Our plant is much more 
closely related to A. peruanus Baill., in which the receme 
is elongate, the stanens united into phalenxes, and the ter- 
minal leaflet much larger than the leteral ones. 

DENDROSTIGMA gen. nov. Flacourtiacearum. Sepala 2 vel 3, 
valde imbricata. Petala 10 spiraliter imbricata. Stamina 
numerosa (circa 407). Ovarium superum muricatum l-loculare. 
Styli 5 trichotome ramosi in stigmata innumerabilia soluti. 
Capsula immature globosa densissime muricato-spinosa. Arbor 
foliis alternis stipuleatis, floribus 1-43 axillaribus speci- 


530 P-B.Y-t.0 L.0.62°4 No. l 


osis albidis. 

DENDROSTIGMA HYSTRICINA sp. nov. Petioli pilosuli; lam- 
inae magnae oblongae breviter ecuminatae repando-dentatae 
basi obtusae vel rotundatese supre glabrae subtus ad costam 
venasque majores parce pilosulae venis subtus discoloribus 
pulchre reticulatis; sepala late rotundata extra pilosula; 
petala triangulari-obovata; filamenta crasse teretia anth- 
eras anguste ellipsoideas subaequantia; ovarium globosum 
muricatum et villosum; styli 5 ovarium subaequantes ramos- 
issimi, stigmatibus conicis. 

Small tree, the younger branches drying striate and some- 
whet sulcate, brown, sparsely puberulous; petioles pilosul- 
ous, 15-30 mm. long; leaf-blades thin but firm, oblong or 
elliptic-oblong, 12-24 cm. long, 5-9 cm. wide, short-acumin- 
ate, repand-serrate with low blunt teeth 5-10 mm. apart, 
somewhat narrowed from neer the middle to a rounded or ob- 
tuse base, glabrous above or minutely pilosulous on the 
costa, beneath glabrous on the surfece and pilosulous on the 
costa and principal veins; lateral veins about 10 on each 
side, curved-ascending; venation conspicuous beneath, browne 
ish and beautifully reticulate; pedicels axillary, about l 
cm. long; sepals thick and leethery, rotund or depressed, 
strongly concave around the flower, 10-12 mm. long, as mich 
as 16 mm. wide, rounded above, sparely pilosulose without; 
petals triangular-obovate, neerly white, 13-16 mm. long, 12- 
13 mn. wide, broadly rounded above, somewhat crisped or 
slightly erose on the margin, cuneate or subunguiculate at 
base; filaments contiguous but distinct, 1.8-2.5 mm. long, 
subulate, thinly villous; anthers narrowly ellipsoid, 2 mm. 
long, setose in a narrow strip on each side, elsewhere gla- 
brous; ovary 4 mm. long, densely covered with villous fleshy 
projections 0.5-1 mm. long, l-celled with numerous ovules on 
parietal placentae; styles 4-4.5 mm. long, villous on the 
lower third, above this point repeatedly trichotomously 
brenched and terminating in about 300 minute conical stig- 
mas; fruit globose, 3 cm. in diameter, densely spiny. 

Type, Lawrance 406, collected in the Alto Chapén region, 
alt. about 2100 m. Tre collector notes it as a tree 20 feet 
high and 6-8 inches in diameter, with sweet-scented greenish 
white flowers and brownish stamens. He was impressed by the 
remarkable stigmas and refers to them in his notes: "Flower 
is surmounted by a green parsley-like tuft growing out of a 
round green-colored ball." Dendrostigma is related to May- 
na, five species of which were described from Colombia by 
Kersten and Triana under the generic name Dendrostylis. They 
state that the ovary is 3-carpellate with three bipartite 
styles which are miltipartite at the stigmas; the flowers 
are completely unisexual. The same characters ere assigned 
to Mayna by Melchior in the second edition of the Pflanzen- 


1943 Gleason, Plantae Lawranceanse 31 


familien and it is unbelievable that any of these authors 
would have neglected the remarkable features of our genus if 
they had been observed. In superficial appearance of the 
fruit and leaf, except the venation, our plant is much like 
an Apeiba. 

HENRIETTELLA LAWRANCEI sp. nov. § Euhenriettella; frutex 
2-5 m. altus; ramli teretes gracili, primum hirsuti, pilis 
subuletis basi incrassatis recurvatis, supra basin patenti- 
bus, mox glabri; petioli graciles 8-15 mm. longi sicut ram- 
uli villosi; laminae tenues opacae lanceolato-ellipticae vel 
oblanceolatee, 5-8 cm. longae 16-25 mm. latae, breviter acu- 
minatae, integrae, ad basin cuneatae, 5-pli-nervisae, nervis 
exterioribus inconspicuis, supra planis subtus leviter prom- 
inentibus, nervis secondariis subhorizontaliter patentibus 
supra obscuris, supra ad costam et nervorum lateralium basin 
breviter pubescentes, ad paginam scabrae tuberculis depresso- 
conicis albidis in pilum brevissimum abeuntibus, subtus hir- 
suteae praecipue ad venas venulasque, pilis circiter 1 m. 
longis erectis; flores 5-meri in fasciculis paucifloris ad 
nodos ramorum vetustiorum infra folia oriundi; bracteae lan- 
ceolatae subulatee 0.8-1 mm. longae; pedicelli 3-6, 1.2 mm. 
longi; hypanthium campanulatum 10-costatum 1.6 mm. longum 
parce strigosum, pilis crassis 0. 4 mm. longis; calycis tubus 
paullo patens O.1] mm. latus; sepala late triangularia, a 
sinibus rotundatis lateribus concavis, 0.2 mm. longa; petala 
ad anthesin reflexa late ovata obtusa integra, 1.8-2 mn. 
longa, 1.6 mm. lata; stamina isomorpha; filamenta gracilia 
2.2 mm. longa, O.7 mm. infra apicem: geniculata; antherae ob- 
longo-lineares 1.7 mm. longae poro apicali dehiscentes, con- 
nectivo dorso elevato; ovarium breve inferum 5-loculere, 
summo plano vel paullo depresso; stylus crassus teres 5.5 
mm. longus, stigmate capitato 0.8 mm. diametro. 

Type, Lawrance 265, from thick forest in the Chapon reg- 
ion. The flowers are described as white and very fragrant; 
the stamens are also white. In general aspect the plant is 
reminiscent of H. Goudotiana Naud., but in its structure it 
shows a much closer relationship to H. verrucosa Triana. The 
latter is a much stouter plant, with thicker and broader 
leaves lacking the white tubercles of our hel: and much 
longer pedicels. 

MICONIA PLENA sp. nov. § Cremanium (? ); arbuscula 5-6 m. 
alta; ramuli acute quadrangulares et ad nodos incrassatos 
sat compressi, dense ferrugineo-tomentelli, vetustioribus 
glabrescentibus; petioli graciles, 15-35 mm. longi sicut 
ramuli tomentelli; laminae elliptico-oblongae, firmulae, 10- 
17 cm. longee, 3-6 cm. latae, acuminatae, minutissime repan- 
dae vel integrae, basi acutae, 5-nervieae, nervis 2 marginal- 
ibus tenuioribus et infre apicem evanescentibus, venis sec- 
underiis rectis 3 mm. dissitis sub angulo 80 orientibus 


52 PLB: T.0. LOS No. l 


supra obscuris subtus prominulis, supra subnitentes subtus 
arcte stellato-tomentellae more M. chrysophyllee; inflores- 
centia late paniculata ramosa miltiflora, 1-2 cm. stipitata, 
axibus arcte tomentellis; flores S-meri, sessileg, ad ramos 
paniculae singuli, oppositi, vel fasciculati et terminales 
glomerati; hypanthium late campanulatum ecosteatum, usque 
torum 1.7 mm. longum, arcte rufescenti-tomentellum, calycis 
tubus 0.4 mm. longus, margine subtruncato, sepalis vix evol- 
utis, dentibus exterioribus late triangularibus sepala vix 
excedentibus, sicut calyce rufescentibus; petala obovato- 
oblonga, 2.2 mm. longa inaequilatera oblique truncata; stam- 
ina 20 isomorpha, inter se in magnitudine variabilia; fila—- 
menta 1.6 vel 1.8 mm. longa gracilia, 0.3 mm. infre apicem 
geniculata; antherae oblongae, 0.9 vel 1.1 mm. longae, ad 
basin subcuneatae, supra medium latiores, poro lato ventro- 
terminali dehiscentes, connectivo simplici; ovarium inferum 
4-loculare, ovulis in quoque loculo 4-6, elongato-ellipsoi- 
deis, a placenta basali erectis; stylus teres glaber 4 m. 
longus, stigmate punctiformi. 

Type, Lawrance 152, from the Alto Chapon region. The as- 
signment of the species to § Cremanium is questionable. The 
distally widened anthers are too broad for § Eumiconia or § 
Glossocentrum, while both anther and pore are narrower than 
the usual proportions in § Cremanium. The geniculate fila- 
ment suggests § Cremanium. The anthers are probably 4- 
celled, as in M. theaezans, but this could not be verified. 
In general habit the species greatly resembles \i. chryso- 
phylla (Rich.) Urb. (= M. fulva DC.), but differs from it in 
numerous structural features. 

MICONIA TUBULOSA sp. nov. § Tamonea; M. Kraenzlinii af- 
finis differt foliis multo majoribus subtus arctissime cin- 
erascentibus supra glabris caudato-acuminatis, panicula maj- 
ore, floribus numerosioribus, ovario 5-loculare superne vil- 
loso, styli basi villoso. 

A shrub 2.5-4 m. tall, younger branches lightly flattened 
and densely tomentose with deep brown stellate hairs, the 
same indument covering the petioles and inflorescence; 
leaves conspicuously uneaual in each pair; petioles slender, 
strongly furrowed, 15 or 35-50 mn. long; bledes thin, ellip- 
tic, 10-15 or 20-25 cm. long, 5-6 or 8-9 cm. wide, tapering 
to a caudate tip 2 cm. long, conspicuously denticulate, ob- 
tuse to subrotund at base, 5S-nerved with an additional pair 
of marginal veins, glabrous above, very closely and minutely 
cinereously stellate-tomentulose on the surface beneath; 
primary and secondary veins lightly impressed above, the 
latter 4-7 mm. apart, diverging at an angle of about 80 ; 
primaries and secondaries prominent beneath, roughly brown- 
stellate, tertiaries prominulous and longitudinally: paral- 
lel, the quaternaries obscure; panicle compact and thyrsoid, 


| 
f 


1933 Gleason, Plantae Lawranceanse ay 


12 cm. long, including the peduncle of 3 cm., its axis and 
branches roundly 4-angled and shallowly sulcate; flowers 5- 
merous, sessile in small terminal glomerules; hypanthium 
nerrowly tubular, 4.5 mm. long to the torus, obscurely 10- 
ribbed, densely cinereous-tomentose; calyx-tube erect, pro- 
longed 1.2 mn., the sepals depressed semicircular, 0.3-0.4 
mm. long, pubescent externally like the hypanthium; exterior 
teeth none; petals obovate, 5 mm. long, 3 mm. wide, strongly 
inequilateral, acute on one side, silvery-stellate extern- 
ally; stamens nearly isomorphic; filaments slightly flatten- 
ed, glabrous, immature; anthers (immature) subulate, 3 mn. 
long, apparently 2-celled; connective flat on the back, di- 
lated toward the base and prolonged into an ovate rounded 
lateral lobe on each side and a somewhat longer dorsal lobe, 
which is ovate in one series of stamens and truncate and wi- 
der in the other series; ovary nearly inferior, 5-celled, 
prolonged above into a short beak terminating in rounded 
villous lobes; style stout, straight, terete, 10 mm. long, 
sparsely villous near the base; stigma truncate, 1.1 mm. in 
diameter. es 

Type, Lawrance 484, collected in thick forest in the £1 
Umbo region, alt. about 900 m.-The collector notes that the 
flowers are sea-green in color. Tpe silvery petals, the 
shape of the sepals, the beak of the ovary, the shape of the 
stigma, and the character of the connective indicate its 
close relationship to M. Kraenzlinii Cogn., from which it 
differs in the characters mentioned in the diagnosis. 

MICONIA MEGALANTHA sp. nov. § Adenodesma; arbor excelsa, 
ramis supremis rachideque inflorescentiae acute 4-angulatis 
arctissime stellato-tomentellis; folia longe petiolata el- 
liptico-oblonga; hypanthium late campanulatum superne valde 
incrassatum et ovarium 5-loculare inferum fere toto amplect- 
ens; calyx late patens lobis obsoletis; petala 5 pro genere 
longissima; stamina subisomorpha filamentis dense glandulo- 
sis, antheris subulatis 2-locularibus valde curvatis, con- 
nectivo crasso ad medium marginis glanduloso ad basin in 
_lobos 2 laterales dense glandulosos dilatato; stylus longus 
glandulosus stigmate peltato. 

A tree "75-100 ft. high" (Lawrance), upper branches 
sherply 4-angled, somewhat dilated at the nodes, very close- 
ly and finely tomentulose with minute cinereous stellate 
hairs, the same type of indument extending over the inflo- 
rescence, hypanthia, petioles, and veins of the lower leaf- 
surface; petioles slender, 5 cm. long; blades thin, oblong- 
elliptic, about 10 by 25 cm., abruptly acuminate, entire, 
acute at base, S-nerved, including a weaker pair of submar- 
ginal veins, thinly stellate-puéscent above when young, per- 
sistently so on the primaries, sparsely stellate on the sur- 
_ face beneath; principal veins lightly impressed above, prom- 


3h PHYTOLOGIA No. 1 


inulous beneath, the secondaries 6-8 mm. apart, straight, 
diverging at almost right angles; inflorescence a freely 
branched, rather narrow panicle 15-25 cm. long, its brenches 
sometimes in whorls of 3, the pedicels mostly sbout 5 mm. 
long; flowers 5-merous; hypanthiun very thick-walled and 
firm, 2.5 mm. long to the torus, its walls greatly thickened 
inwardly above the ovary and closely surrounding the style; 
calyx widely spreading, 9 mm. in diameter, exactly truncate, 
sparsely pubescent internally, tomentellous like the hypan- 
thium, the exterior teeth minute tubercles only; petals 
widely spreading at anthesis, elliptic, 25 mm. long, 10 mm. 
wide, densely and finely gray-tomentulose on the outer side, 


almost equilateral, several-nerved; stamens almost isomorph- 4 


ic; filaments slender, arcuate above, 11-12 mm. long, dense- 
ly pubescent throughout with minute stipitate glands; anth- 
ers stoutly subulate, 7-8 mm. long, abruptly curved ventral- 
ly above the base, 2-celled, opening by a terminal pore; 
connective stout and thick, glandular along its margin near 
the middle, at base dilated into 2 lateral lobes which curve 
ventrally over the sides of the thecae and are strongly 
glanduler; ovary wholly inferior, 5-celled, prolonged above 
into a stout beak 1 mm. high and dilated at its summit into 
a 5-lobed saucer-shaped organ 1.4 mm. wide; style somewhat 
curved, 14 mm. long, densely glandular-puberulent through- 
out; stigma peltate, 2 mm. in diameter. 

Type, Lawrance 431, collected in forest in the Lower Cha- 
pon region, alt. about 900 m. According to the collector's 
field notes the flowers are "peach-blossom" in color and the 
wood is used for building houses. A second collection, no. 
456, is in fruit; it comes from the El Umbo region at 900 m. 
and is described as a tree 15-18 m. high and 6-12 dm. in 
diameter, with bark "reminding one of the bark of the fir 
tree." The fruiting hypanthium is 8 mm. in diameter, ob- 
scurely 10-ribbed, and surmounted by the persistent calyx. 

The shape and glandulosity of the anthers is precisely 


that of the nine species recently included by me in § Adeno- g 


desma (Bull. Torrey Club 59: 365-370. 1932), and the species 
is accordingly added to that group notwithstanding the long- 
petioled 5-nerved leaves. In M. axinaeoides Gl. the leaves 
are only weakly 5-pli-nerved, the hypanthium-is similarly 
thickened distally, and the lobes of the calyx are also ob- 
solete, while the stamens are almost precisely like those of 


M. megalantha. Our new species may be differentiated from M. 


axinaeoides and from the other members of the section by 


amending the key. presented on p. 366 (l.c.) as follows: Del- 


ete the paragraph leading to M. axinaeoides, beginning 

"Leaves weakly 5-pli-nerved," and substitute the following: 

Leaves 5-nerved or weakly 5-pli-nerved; walls of the hypan- 
thium greatly thickened distally; calyx-lobes obsolete; 


1933 Gleason, Plantae Lawranceanae 55 


style and filaments densely glandular; style 14-21 mn. 
long. 

Petiole 1.5 cm. long; leaf-blades broadly obtuse or subro- 
tund at base; glabrous on the actual surface beneath; 
petals 16 mm. long; stigma small and truncate 

4, M. axinaeoides. 

Petiole 5 cm. long; leaf-blades acute at base, stellate on 
the lower surface; petals 25 mm. long; stigma peltate, 2 
mm. in diameter 4a. M. megalantha. 

It is interesting to note that the flowers of M. megalan-.- 
tha are the largest known in the genus Miconia. 

HENRIETTELLA FISSANTHERA sp. nov. (an gen. nov.?) Frutex; 
ceules juniores 4-angulati et sulcati, mox subteretes glab- 
rescentes; petioli graciles tenuiter furfuracei; laminae 
tenues anguste obovatae, ad apicem obtusam abrupte acuminat- 
ae, ad basin late cuneatae, 5-pli-nerviae, minutissime albo- 
punctatae, ad venas tenuissime furfuracese; flores 5-meri 
graciliter pedicellati in fasciculis paucifloris infra folia 
orientibus; hypanthium late campanulatum parvum; sepala ob- 
soleta; petal triangulari-lanceolata acuta intus supra med- 
ium lobos carnosos 3-4 gerentes; stamina isomorpha, filamen- 
tis complanatis, antheris oblongis crassis rimis 2 tota 
longitudine dehiscentibus, connectivo basi minute calcarato; 
ovarium 5-loculare inferum; stigma ovoideo-oblongum basi 
5-lobatum. 

Shrub 2-3 m. high, the young stems densely brown-furfur- 
aceous, roundly 4-angled and sulcate, soon becoming subter- 
ete and nearly glabrous; petioles slender, 1-2 cm. long, 
thinly furfuraceous, soon glabrescent; blades thin, bright 
green, narrowly obovate, 8-11 cm. long, 3.5-5 cm. wide, ab- 


_ruptly acuminate to a slender obtuse tip, entire, broadly 


cuneate at base, 5-pli-nerved, barely scaberulous above with 
minute whitish punctae, sparsely and minutely fur furaceous 
on the primeries beneath, otherwise glabrous; principal 
veins plane above, prominulous beneath, the secondaries 2-3 
mm. apart, arising at an angle of about 80 , upper primaries 
diverging 10-15 mm. above the base, the outer pair weaker 
and subnarginal; inflorescence of opposite fascicles of 2-6 
flowers from the leafless lower nodes; pedicels filiform, 9 
mm. long, thinly furfuraceous; flowers 5-merous; hypanthium 


very broadly cempanulate, 2.5 mn. long, sparsely pubescent 


— 


with dense but scattered tufts of minute hairs; calyx-tube 
prolonged about 0.2 mm., the sepals nearly obsolete, de- 
pressed-triangular, projecting only 0.1 mm.; petals barely 
imbricate, widely recurved-spreading at anthesis, lance- 
triangular, 4 mm. long, 2 mm. wide at base, acute, on the 
back dark-colored, sepal-like in texture and scabrously fur= 
furaceous, on the inner side greatly thickened just above 
the middle into 4 fleshy oblong protuberances pointing in- 


36 PHYTOLOGIS No. l 


ward and downward; stamens 10, isomorphic; filaments flat, 
2.2 mm. long, triangularly narrowed at the summit; anthers 
stoutly oblong, 1.9 mm. long, 2-celled, soon opening by 2 
longitudinal slits extending from apex to base; connective 
prolonged at base into an ovoid blunt spur 0.2 mm. long; 
ovary wholly inferior, 5-celled, many-ovuled, its upper wall 
concave and extending almost to the torus, radially 10- 
ribbed; style terete, straight, 5 mm. long at maturity; 
etigme ovoid-oblong, 1.3 mm. long, 5-lobed at base around . 
the style. 

Type, Lawrance 606, collected in dense forest in the El 
Umbo region, alt. about 900 m. It differs from every other 
species of the genus known to me, either directly or through 
description, in two important characters, the longitudinal 
dehiscence of the anthers and the elongate 5-lobed stigma. 
Such differences are usually considered of sufficient weight 
in this family to warrant the erection of a genus. It has 
nevertheless been kept in Henriettella because of its entire 
agreement with the other characters of the genus. The -pecul- 
iarly appendaged petals have not been observed by me in the 
genus before, but similar structures have been found in Bel- 
lucia, which is apparently closely related. 

CYPHOSTYLA STRIGOSA sp. nov. Caulibus, petiolis, venis 
gsubtus, inflorescentia, hypanthiis, et calycibus strigosis 
pilis incurvis; alabastris elongatis fusiformibus; petalis 
magnis. 

Shrub about 2 m. high, the younger branches, petioles, 
inflorescence, hypanthium and calyx densely strigose with 
stoutly subulate, incurved hairs about 0.7 mm. long; peti- 
oles slender, 10-18 mm. long; leaf-blades firm, elliptic- 
oblong, 8-14 om. long, 3.5-6 cm. wide, short-acuminate, en- 
tire, rounded to broadly obtuse below and subcordete, the 
suricles more or less overlapping, 5-7-pli-nerved, finely 
strigose above with short brown curved hairs and densely 
strigose on the primaries with long straight hairs, pubes- 
cent beneath, especially on the veins and veinlets; veins 
plane above, prominulous beneath, the secondaries about 3 
mn. apart, the tertiaries finely reticulate; inflorescence 
a emell, few-flowered, terminal panicle, the 5-merous flow- 
ers all sessile, but often solitary and then apparently on 
pedicels 10-15 mm. long; buds fusiform, 20 mn. long, taper- 
ing to both ends; hypanthium 7 mm. long at anthesis, ob- 
conic, its walls greatly thickened distally to the terminal 
annular torus which bears 5 broadly elliptic petal-scers and 
5 trianguler stamen-scare; calyx calyptriform, deciduous at 
anthesis without splitting; petals broadly triangular-obov- 
ete, 36 mn. long, 26 mm. wide, equilateral, rounded above, 
cuneate to the base, freely nerved; stamens 5, isomorphic, 
alternate with the petels; filaments stout, 6 mm. long, 


een me pS ee re 


—— 
?— 


7 tae 


Pie. 


1933 Gleason, Plantae Lawranceanae S7 


rounded on the inner side, flat on the outer, strongly re- 
curved above; anthers oblong, straight or slightly curved, 
44.5 mn.’long, the tip strongly arched over the concealed 
pore; connective simple, gradually expanded basally and con- 
tinuous with the filament; ovary inferior; style terete or 
obscurely ribbed, about 5 mm. long, near the summit greatly 
thickened and bent at right angles to the truncate stigma. 
Type, Lawrance 419, collected in the Alto Ohapén region, 
alt. about 2100 m. The genus Cyphostyla was proposed in 


1929 to include two plants of remarkable floral structure 


from Antioquia. It is distinctly interesting to detect a 
third species among the recent collections of Lewrance. The 
two original species, C. hirsuta and C. villosa, were char- 
acterized by the long reflexed hairs of the hypanthium and 
calyx, while C. strigosa has stout, short, erect and more or 


‘less incurved hairs, and therefore necessitates a small 


emendation in the generic characters as originally stated. 
ALLOMAIETA GRANDIFLORA Gl., originally described from 
Paime in northern Cundimarca, has been re-collected by Law- 


-rance in the adjacent mountains of Boyacd. 


SACCOLENA DIMORPHA Gl. Originally described in 1925 from 
a@ single specimen, this little known species has now been 
collected again by Lawrance, no. 144, in the mountains of 
Boyacé near its type locality at Paime in Cundimarca. Field 
notes indicate that it grows there in sphagnum moss at an 
altitude of about 2300 m. The collector adds that it has 
white flowers and is worth propagating because of the very 
dark green upper surface of the leaves. Certain amendments 
to the original descriptions are now possible. In the gener- 
ic description: petals white, rather than yellow; anthers 
subulate and elongate, not oblong; appendage of the connect- 
ive subulate in the long stamens, ovoid-triangular and sac- 
cate in the short stamens. In the specific description: 
height 8-25 cm.; leaves rounded to cuneate at base; pedunc- 
les stramineous, sharply contrasting with the stem in color; 
petals obovate, entire; anthers subulate, the smaller neerly 
straight, 2.2 mm. long, the connective prolonged 0.6-0.8 mm. 
to the summit of the filament and bearing a flattened ovoid- 
triangular ascending appendage; large anthers dorsally 
curved, 5.3 mm. long, the connective prolonged 1.9 mm. to 


the filament. Examination of the original dissection of the 


type shows that the anthers are all broken, a condition not 
recognized then end explaining the considerable discrepancy 
in the dimensions of the stamens. 

JUANULLOA BICOLOR sp. nov. Liana alta ramis juvenilibus 
tomentosis mox glabrescentibus et irregulariter papyraceo- 
alatis; petioli tomentosi; laminae mediocres oblongae obtu- 
sae basi subrotundatae supre glabrae opacae subtus stellato- 
pubescentes, venis circa 4 in utroque latere arcuato-con- 


38 PHY f0L0 G 3's No. l 


nectis venulis obsoletis; inflorescentia mltiflora corymbi- 
formis; calyx campanulatus 5-angulatus coloratus lobis ob- 
longis acuminatis; corolla anguste tubulosa supra medium vix 
dilatata extra tomentosa, lobis brevibus erectis ovatis; 
stamina inclusa; stylus breviter exsertus. 

A liana, climbing to 15 m., the flowering btanches slend- 
er, thinly tomentose, soon becoming glabrous and irregularly 
wingedwinged with projecting plates of cork; petioles stout, 
10-15 mm. long, densely stellate-tomentose; leaf-blades 
thin, charteceous, oblong, 7.5-10 cm. long, 4.5-5.5 cm. 
wide, broadest slightly above the middle, obtuse, entire, 
subratund at base, opaque and minutely rugulose above, be- 
neath thinly stellate-pubescent on the surface and tomentose 
on the veins; lateral veins plane above, scercely elevated 
beneath, about 4 on each side, curved-ascending and obscure- 
ly ercuately connected, veinlets obsolete; inflorescence 
terminal and from the upper exils, corymbiform, densely fer- 
ruginous-tomentose, the pedicels at anthesis about 6 m. 
long; calyx campanulate, thick and coriaceous, 27 mm. long, 
thinly ferruginous-tomentose externally, 5-lobed about to 
the middle, the lobes oblong, triangularly acuminate; corol- 
le narrowly tubular, 57 mm. long, 6 mm. in diameter at base 
and summit, 7-8 mm. in diameter above the middle, densely 
stellate-tomentose externally, glabrous within, its lobes 
broadly ovate, erect, 4 mm. long; stamens 5, inserted 5 mm. 
above the base of the corolla; filaments slender, 40 mm. 
long, densely white-villous at’ base; anthers linear, 13 mm. 
long, included in the corolla-tube; style filamentous, ex- 
serted about 5 mm., the stigma narrowly clavate. 

Type, Lawrance 445, collected in thick forest in the El 
Umbo region of Boyacé, alt. about 900 m. Juanulloa bicolor 
is apparently a member of § Sarcophysa, differing from the 
previously known J. speciosa (Miers) Dun. in its less pubes- 
cent, thinner leaves, its proportionately longer and. more 
slender corolla, and its mch shorter and erect petals. 


x 
> 
= 
t 
* 
é 


LEITGEBIA AND RORAIMANTHUS 


c H. A. Gleason 

The genus Leitgebia was established by Eichler in 1871 to 
include a small shrub of heathlike aspect, L. guianensis 
Eichl., originally collected on the high savannas near Mount 
Roraima by Schomburgk. Eichler's description is accompanied 
by a plate. His description is accompanied by a”plate and 
the two togetler leave little doubt as to the structure of 
the floral organs. Its characteristic features, at least in 
contrast to other plants to be mentioned below, are the pre- 
sence of five staminodes which are inserted between the sta- 
mens and in a continuous circle with them, clawed at hase, 
spatulate and flattened at summit, and the ovoid ovary, 
blunt at the top, l-celled, with a few (6-8) ovules attached 
at the base, terminated by a very slender style. 
_ Some years later Oliver added a second species, Leitgebia 
imThurniana, collected on the summit and upper: levels of Ro- 
raima by im Thurn and recently re-collected there by Tate. 
Oliver called attention to the specific characters which 
separate the two very clearly, but apparently failed to see 
or to comprehend the generic significance of certain other 
structures. In Oliver's species the five staminodes are 
quite separate from the stamens, but are connate with each 
other, forming a short sheath outside the stamens; each of 
the five is elongate, surpassing the enthers, petaloid, 
spatulate, and more or less involute. The ovary is conic, 


~tapering gradually into the stout style, 3-celled, with sev- 


| 
* Lt 2 


eral ovules in each locule. There. is no doubt that it repre- 
sents a distinct genus, which is here proposed. | 

RORAIMANTHUS gen. nov. Folia conferta praesertim termina- 
lia, stipulis alte fimbriatis. Flores axillares pedicellati. 
Sepale 5 imbricata. Petala 5 obovata rosea vel alba. Stamin- 
odia 5 inter petala et stamina inserta basi connata stamini- 
bus alterna, lobis erectis petaloideis spathulatis involu- 
tis. Filamentia brevia. Antherae lineariae erectae filamen- 
ta excedentes. Ovarium 3-loculare miltiovulatum conicum in 
stylum elongatum angustatum. 

A shrub of ericoid aspect, the small leaves crowded at 
the summit of the branches. Leaf-blades obovate-oblong, 
thickened at the mrgin, crenate near the apex, with ascend- 
ing branched lateral veins. Stipules persistent, deeply 
fringed. Flowers 5-merous, axillary among the upper leaves, 
short-pedicelled. Staminodia 5, connate at base into a short 
membranous tube surrounding the base of the stamens, the 
lobes alternate with the stamens, erect, membranous, peta- 


39 


40 P SY Oo eee No. 1 


loid, spatulate, involute. Stamens 5 with short filaments 
and linear-subulate anthers. Ovary conic, 3-celled with 
numerous ovules, gradually tapering into the subulate 
style. Stigma punctiform. 

RORAIMANTHUS IMTHURNIANUS comb. nov. (Leitgebia im- 
Thurniana Oliver) Endemic, so far as known, to the summit 
end upper levels of Mount Roraima 


A NEW ELEPHANTOPUS FROM BRAZIL 
H. A. Gleason 


ELEPHANTOPUS ERECTUS sp. nov. Herbaceus circa 6 dm. altus, 
caulibus densiter pilosis foliaceis; folia 6 vel 7 utrinque 
densiter pubescentia leviter crenato-serrata, inferiora 15 
cm. longa 4 cm. lata oblongo-oblanceolata subacuta ad basin 
cuneata in petiolum brevem, superiors minora usque ad 5 cm. 
longa 2 cm. lata sessilia basi rotundata; spicae 2 terminal- 
es erectae circa 4 dm. longae densiter pubescentes, infra 
medium steriles bracteis 1 vel 2 parvis ovatis; glomeruli 
sessiles et spicati 9-12; bracteae numerosae, exteriores ov=- 
ato-lanceolatae usque ad 2 cm. longae, interiores gradatim 
angustiores et breviores, intim@e lineari-subulatae capitula 
aequantes, omnes densiter sericese; capitula 4-6 in quoque 
glomerulo; involucrum 1] mm. altum, squamis 4 exterioribus 
5.5 mm. longis ovato-lanceolatis acutis ad apicem mimute 
sericeis, interioribus 4 11 mm. longis limearibus apicem 
versus sericeis; achaenia 4 lineari-clavata 4.5 mm. longa 
densiter hirsuta; pappi setae 5 vel 6, 2-2.5 mm. longae 
laeves planae e basi ad apicem angustatae. 

Type, number 16231, collector not stated (Glaziou?), from 
Este. Fortaleza, Sao Paulo, Brazil, 29 Apr. 1899, in the 
herbarium of the New York Botanical Garden. It is obviously 
related to E. hirtiflorus DC., and differs in its denser 
pubescence, much larger and broader serrate leaves, more 
numerous spicate heads, and shorter psppus. 


NINE SOUTH AMERICAN MELASTOMES 
H. A. Gleason 


Unless otherwise stated, the type specimens are in the 
herbarium of the New York Botanical Garden. 

GRAFFENRIZDA COLOMBIANA sp. nov. Arbor 8 m. alta, ramlis 
supremis profunde bisulcatis glabris; folia paulo inaequal- 
ia; petioli 45-70 mm. longi graciles subteretes minutissime 
furfuracei; laminae late rotundato-ovatae membranacese 15-20 
cm. longae 15-17.5 cm. latae superne late obtusae ad basin 
late cuneatam rotundatae supra virides glabrae subtus fusc- 
escentes prope basin ad venas furfuracese ceterum glabrae, 
5-pli-nerviae, venis 2 marginalibus neglectis, nervi primer- 
ii usgque ad apicem ercuatim producti, vense secundariae in- 
ter costam et venas primariss 2 et 3 sub angulo circa 60 
orientes et infra medium folii distaliter recurvae, 6-8 mm. 
inter se distantes, inter venas 2 et 3% et venas exteriores 
subdirecti; venulae tertiasriae obsoletae; panicula terminal- 
is 15 cm. longa, pedunculo 6 cm. longo fere glabro sustenta, 
ramis paucis distantibus 2-5 cm. longis; flores 4-meri in 
capitulis plurifloris lateralibus termihalibusque subsessil- 
ibus dispositi; pedicelli 1-1.5 mm. longi; hypanthium cam- 
panulatum 2.3 mm. longum obscure nervatum tenuissime et 
sparsissime furfuraceum; calyx in alabastro acutus 1.4 m. 
longus, ead anthesin fere ad torum in lobos 3 vel 4 irregu- 
lares ruptus; petala oblonga 4.7 mm. longa 1.6 mm. lata 
breviter acuminata; stamine isomorpha; filamenta subulata 
2.2 mm. longa glabra; antherae arcuatae aurantiacae subula- 
tae 4.6 mm. longae; connectivum infra thecas 0.2 mm. longun, 
in calcsr erectum subulatum 0.8 mm. longum productum; ovar-- 
ium liberum ellipsoideum 5-costatum glabrum 3-loculare; sty- 

lus grecilis 7 mm. longus, stigmate.punctiformi. 

Type, Klug 1866, in forest at Umbria, Comisaria del Puto- 
mayo, Colombia, alt. 325 m. Twenty-six species of Graffen- 
rieda have been described, of which seventeen are included 
in Cogniaux' monograph. It is quite probable that detailed 

_ study of the genus as a whole will indicate that his classi- 

_ fication of the species is artificial. Our species would be 
- associated with numbers 11 to 15 of the monograph, with 
Which it has apparently no real relation. It is rather to 

be compared with G. emarginata (R. & P.) Triana, which has 
much thicker leaves, cordate at base and conspicuously 

_ veiny, much stouter petals and rounded calyx-lobes, and 

_ which is much more pubescent in all its parts. 

+ MICONIA PACHYDONTA sp. nov. § Jucunda: arbuscula; foliis 

_ oObovato-oblongis mediocris breviter petiolatis apiculetis 


4y 


4 PH YT OLGA No. l 


basi angustatis alternatim 5-pli-nerviis supra glabris sub- 
tus arctissime cinereo-tomentosis; panicula tomentosa, flor- 
ibus 5-meris sessilibus in glomerulis bracteatis ageregatis; 
hypanthio tomentoso tubuloso; sepalis mox deciduis truncato- 
triangularibus, dentibus exterioribus magnis pyramidatis 
petulis; petalis spathuletis; staminibus fere isomorphis, 
antheris subulatis basi bilobis; ovario 4-loculare, stylo 
elongato basi glanduloso-puberulo. 

Small tree 5 m. high, the younger stems nearly terete, 
very finely fulvons-tomentose, soor. glabrescent, the inter- 
nodes 2-5 cm. long; petioles 5-10 mm. long, channeled above, 
thinly tomentulose or furfuraceous; leaf-blades thin, obo- 
vate-oblIong, 12-20 cm. long, 5-8 cm. wide, abruptly apicu- 
late, entire, narrowed or subcuneate at base, alternately 
5-pli-nerved, with an additional pair of marginal veins, 
dull green and glabrous above, very finely cinereous-toment- 
ulose beneath; uppermost primaries srising 3-4 cm. above the 
base, neerly plane above; secondaries 3-8 mm. apart, in the 
basal part of the leaf spreading horizontelly, above at an 
angle of about 70 , obscure above, elevated beneath; terti- 
aries obsolete above, reticulate beneath; panicle erect, 1 
dm. long, its axes strongly angled and compressed, thinly 
cinereous; flowers 5-merous, sessile, the lower solitary at 
the nodes or in 2-3-flowered glomerules, the upper crowded 
in 3-5-flowered glomerules; bracts closely appressed, sub- 
orbicular, about 7 mm. long, thinly stellate; hypanthium 
nearly tubular, thick-walled, 7 mm. long to the torus, 
densely and clogely cinereous-tomentose; calyx-tube not pro- 
longed; sepals connivent or connate in bud, at anthesis 
splitting to the torus and soon deciduous, truncate-triangu- 
lar, 2.4 mm. long with a minute terminal apiculum 0.4 m. 
long, thinly pubescent within, tomentose on the back like 
the hypanthium; exterior teeth subapical, stoutly pyremidal, 
spreading at nearly right angles, the acute upper (inner ) 
edge 1.1 mm. long; petele oblong-spatulate, 6 mm. long, 3.2 
mm. wide, subunguiculate, the basal portion and a triangular 
part of the blade thick and fleshy, nearly symmetrical, not 
retuse; stamens nearly isomorphic; filaments flattened, 7 or 
5.7 mm. long, glabrous; enthers subulate, 8.5 or 7 mm. long, 
2-celled, with strongly convolute thecee, the thecae and 
connective bilobed at base and prolonged 0.8 mm. below the 
sumnit of the filament; connective of the smaller anthers 
simple, of the larger anthers slightly elevated just above 
the filament and bearing a minute rounded protuberance; 
ovary about one-third inferior, 4-celled, with a short beak; 
style slightly curved, 17 mm. long, minutely glandular-pub- 
erulent in the lower fourth; stigma truncate or subcapitate. 

Type, Klug 2143, in dense forest at Florfda, at the mouth 
of Rfo Zubineta into Rfo Putomeyo, Peru, alt. about 180 m.; 


eT a ee et en 


POGlast“ 


1933 Gleason, South American Melastomes 4x 


vernacular name (Huitoto Indian) name Jucaguino-ey. A second 
collection at the same place (Klug 2223) bears the native 
name Chaita-nargu-ey. 

Miconia pachydonta obviously belongs with a group of § 
Jucunda recently discussed by me (Bull. Torrey Club 59: 361- 
364. 1932.) and characterized by unusually large exterior 
calyx-teeth, as well as by other features. The primary div- 
ision of the group in the key depends on the leaf-base; our 
species falls in the first group and is at once contrasted 
with M. gretissima Benth., which has similer pyramidal ex- 
terior teeth, but differs in its much narrower leaves with 
primeries arising nearly from the base, smaller and more 
virgate panicle, more thinly tomentose hypenthium, erect and 
somewhat incurved exterior teeth, and spinulose-tipped ovary. 
Notwithstanding its leaves, M. pachydonta is more closely 
related to M. fissa Gl. and M. megaphylla Gl. In both of 
these the leaves are nearly or quite sessile and rounded to 
amplexicaul at base. In M. megaphylla the leaves are also 
9-13-pli-nerved and the flowers 6-merous; in M. fissa the 
smaller anthers are glandular and the exterior teeth are 
carinate on the outer side. 

In most of these species, including M. pachydonta, the 
enthers are bent back so far at anthesis that they lie hori- 
zontally across the flower with the connective down; that 
is, they turn during the opening of the flower through an 
angle of 270 degrees. 

MICONIA COOKII sp. nov. § Amblyarrhena: arborescens; 
caulibus juvenilibus 4-angulstis et sulcatis; petiolis grac- 
ilibus elongatis; lsminis ovato-lanceolatis membranaceis in- 
tegris abrupte acuminatis basi cunestis 3-pli-nerviis utrin- 
que glsbris; paniculis multifloris; floribus 5-meris; hypan- 
thio subgloboso; sepalis trisngularibus acutis dentibus ex- 
terioribus nullis; petalis rotundato-obovatis; staminibus 
isomorphis connectivo basi truncato ad lateres minute pro- 
ducto; ovario 5-loculare stylo elongato stigmete truncato. 

A tree with glabrous stem and foliage; younger stems 
roundly 4-angled and shallowly sulcate; petioles slender, 
2-3 cm. long; leaf-blades thin, bright green, narrowly 
ovate-lanceolate, or occasionally obovate-lanceolate, as 
much ae 16 cm. long by 5.5 cm. wide, abruptly acuminate to a 
short cusp, entire, somewhat cuneate at base, 3-pli-nerved, 
the primeries lightly impressed above, elevated beneath, the 
secondaries 5-7 mm. apart, ascending at an angle of about 
80" » obscure above, barely elevated beneath, the tertiaries 
obsolete above, plane and reticulate beneath; inflorescence 
a sessile, freely branched panicle 11 cm. long; flowers 5- 
merous, in terminel glomerules of 3, on pedicels O.7 mm. 
long, the terminal and sometimes also the lateral short-ped- 
unculate; hypanthium subglobose, 2.2 mn. long to the torus, 


ak PHYTOLOGTA No. l 


very sparsely end minutely arachnoid-puberulous and brown- 
punctate; calvx-tube erect, prolonged 0.2 mm. to acute sin- 
uses; sepals triangular, acute, 0.9 mm. long, somewhat 
spreading and slightly pubescent at the tip, exterior teeth 
not differentiated; petals round-obovate, 2.2 mm. long, ob- 
scurely inequilateral and retuse; stamens isomorphic; file- 
ments stout, glabrous, 2 mm. long; anthers stout, blunt, 2 
mn. long, opening by a minute dorso-terminal pore, the con- 
nective forming @ conspicuous dorsal ridge, truncate at base 
on the back but prolonged at the sides into lateral lobes 
0.2 mm. long; ovary half-inferior, 5-celled, its summit de- 
pressed-conic, sharply 10-angled and minutely 10-toothed; 
style straight, glabrous, 5.6 mm. long; stigma cepitellate, 
berely expanded. 

Type, Cook & Gilbert 1742, collected at San Miguel, Uru- 


bamba Valley, Peru, alt. about 1800 m., and deposited in the 


United Stetes National Herbarium, no. 604912. The species is 
related to M. elongata Cogn. of Bolivia and M. monzoniensis 
Cogn. of Peru. The latter differs from ours in its crowded 
panicles, much shorter petioles, broader leaves sbruptly 
narrowed to a slender cusp, and 4-celled ovary; the former 
has much smaller flowers in all dimensions, peltate stigma, 
and long-acuminete leaves. 

MICONIA MESMEANA sp. nov. § Amblyarrhena: M. difficili 
Triana habitu et structura arcte affinis differt pubescentia 
stellata furfuracea multo temuiore, hypanthio subglabrato, 
calycis lobis erosis, dentibus exterioribus latissime tri- 
angulsri-subulatis, antheris alternatim subineequalibus con- 
nectivo basi l- vel 2-csalesrato, overio 2-loculari, stigmate 
peulo dilatato. 

4 shrub 2-4 m. high, the younger branches minutely and 
sparsely stellate-furfuraceous and rather densely hirsute 
with spreading hairs 1 mm. long; petioles slender, 3-5 mm. 
long, hirsute above, glabrous beneath; leaf-blades firm, el- 
liptic to oblanceolate, 5-7 cm. long, 1.5-2.5 cm. wide, ab- 
ruptly acuminate to an obtuse tip, slightly revolute after 
drying, minutely ciliate with incurved subulate teeth, acute 
or broadly cuneate at bese, 3-pli-nerved; primary veins 
lightly impressed above, prominent benesth and hirsute near 
the base with horizontally spreading hairs; secondsries 
spreading at nearly right angles, obscure above; tertieries 
obscure and reticulate; both surfaces glabrous, the upper 
minutely rugulose; inflorescence rather densely peniculste, 
6-8 cm. long, its branches pubescent like the stem; flowers 
5-merous, subsessile; hypanthium subglobose, 2.4 mm. long to 
the torus, yellowish, spparently glabrous but very minutely 
stellate-furfureaceous toward the base; celyx-tube prolonged 
0.4 mm.; sepals broadly triengular, O.4 mm. long, obtuse or 
subacute, often slightly erose; exterior teeth broadly tri- 


wi tai Loa) atin 


Co oe 


an 


¥) 


1933 Gleason, South American Melastomes 45 


-angular-subulate, appressed, about half aa long as the se- 


pals; petals (immature) suborbicular, shallowly retuse; fil- 
aments (immature) flat, much broadened below the middle; 
anthers ellipsoid, 4-celled; connective slightly elevated 
toward the base into a dorsal ridge and prolonged in alter- 
nate stamens into one or two short basal spurs; ovary half- 
inferior, 2-celled; stigma truncate, slightly expanded. 

Type, Killip & Smith 20012, collected between Pamplona 
and Toledo, Norte de Santander, Colombia. The general simi- 
larity of the plant to the type specimen of M. difficilis is 
striking, but a closer comparison discloses the various dif- 
ferences enumerated in the diagnosis, which are considered 
to justify the recognition of the plant as a distinct spe- 
cies. 

CLIDEMIA OSTRINA sp. nov. § Sagraea: fruticosa; folia am- 
pla cvata 7-9-nervia supra glabra subtus purpurea ad venas 
venulasque minutissime furfuracea; cymae minimae pauciflo- 
rae; flores 4-meri subsessiles; hypanthiun suburceolatum 
parvum; sepaia minuta obtusa, dentibus exterioribus crassis 
patulis late triangularibus; petala ex basi lata oblonga; 
stamina isomorpha, filamentis sub apicem geniculatis, anth- 
eris anguste obovatis 4-locularibus, connectivo paulo elev- 
ato rubro-lineato vel glanduloso; overium toto inferum 2- 
loculare, stylo glabro, stigmate punctiformi. 

Shrub 2.5 m. high, younger stems roundly 4-angled, pro- 
minently sulcate, thinly furfuraceous, later becoming ter- 
ete and glabrous; leaves approximately equal in each pair; 
petioles slender, 3-6 cm. long, minutely furfuraceous; 
blades membrenous, ovate, 10-18 cm. long, 7-12 cm. wide, ab- 
ruptly short-acuminate, entire, rounded to the cordate or 
subcordate base, 7-9-nerved, green and glabrous above, be- 


neath purple, glabrous on the surface, minutely furfuraceous 


on the veins and veinlets; veins plane and obscure above, 
prominulous and reticulate beneath, the secondaries diverg- 
ing et an angle of about 80 , 3-5 mm. apart; cymes few- 
flowered, about 1 cm. long, minutely furfuraceous; flowers 
4-merous, on pedicels 0.5-1 mm. long; hypanthium urceolate 
to subglobose, 1.4 mm. long to the torus, very minutely fur- 
furaceous; calyx-tube prolonged 0.6 mm., its lobes erect, 
very broadly triangular, scarcely more than 0.1 mm. long; 
exterior teeth thick, spreading, broadly trianguler, 0.5 mm. 
long; petals oblong, 1.5 mm. long, 1 mm. wide at base, ob- 


scurely inequilateral and shallowly retuse; stamens iso- 


morphic; filaments flattened, 1.4 mm. long, geniculete just 
below the apex; anthers narrowly obovoid, 1.1 mm. long, 4- 
celled, opening by a minute terminal pore; connective barely 
elevated into a low dorsal ridge, not prolonged at base, 
red-lineate or possibly glandular; ovary inferior, 2-celled; 
style straight, 3.6 mm. long; stigma punctiform. 


46 PHYTOLOGIA No. 1 


Type, Pittier 11818, collected in humid forest at El 
Portachuelo, ‘ragua, Venezuela. According to the treatment 
of the older species by Cogniaux, ¢. ostrine falls among the 
species numbered 73 to 76, all of which have much larger 
panicles and flowers and denser pubescence. Its real affin- 
ity is apparently with C. amrla Cogn., numbered 88 in Cogn- 
iaux' monocraph, which also has much lerger flowers and dis- 
tinctly stellate pubescence. 

HENRIETTELLA LORETENSIS sp. nov. Arbuscula; remis junior- 
ibus minutissime furfuraceis mox glabris; petiolis fere gla- 
bris; leminis oblongo-ellipticis ecuminatis imtegris basi 
cuneatis. 3-pli-nerviis utringque glabris praeter costam sub- 
tus sparissime stellato-puberulam; floribus 4-meris fascicu- 
latis breviter pedicellatis; hypanthio late campanulato; 
calycis tubo truncato dentibus exterioribus minimis subula- 
tis; petalis triangularibus acutis supra medium valde in- 
cressatis; staminibus isomorphis, antheris crassis obtusis 
connectivo simplici; ovario infero 4-loculare; stylo elong- 
ato, stigmate truncato. 

Small tree 5 m. high; twigs grayish-brown, very minutely: 
furfuracesus and somewhat flattened when young, soon glabre- 
scent and terete; petioles slender, channeled above, about 
15 mm. long; leaf-blades thin, oblong-elliptic, 10-13 cm. 
long, 4-4.5 cm. wide, abruptly short-acuminate, entire, cun- 
eate at base, 3+pli-nerved with.an additional pair of margi- 
nal veins, glabrous above, very sparsely stellate-puberulent 
on the midvein beneath; | secondaries 4-5 mn. apart, arising 
at an angle of about 80° ; flowers 4-merous, numerous in 
sessile fascicles below the leaves, on pedicels 4 mm. long; 
hypanthium broadly campanulate, 1.6 mm. long to the torus, 
glabrous; calyx-tube prolonged 0.3-0.4 mm., truncate; exter- 
ior teeth subulate, projecting less than O.1 mm.; petals 
triangular, 3 mm. long, nearly 2 mm. wide, glabrous, thin at 
the base, above the middle strongly thickened on the inner 
side; stamens isomorphic; filaments flat, l-nerved, 2.5 mm. 
long; anthers stoutly oblong, obtuse, 2 mm. long, the con= 
nective neither appendaged nor prolonged; overy inferior, 4- 
celled; style glabrous, slénder, 7 mn. long; stigma truncate. 

Type, Klug 2215, from the Rfo Putomayo, at the mouth of 
Rfo Zubineta, Dept. Loreto, Peru. The species is apparently 
related to H. fascicularis (Sw.) Triana, which has larger 
flowers and hirsute stems and foliage. 

BLAKEA TRUNCATA sp. nov. Frutex scandens glaberrimus; 
rami juveniles crassi irregulariter angulati internodiis 
brevibus; folia chartacea obovato-oblonga ad apicem rotund- 
atam in acuminem brevem linearem, abrupte contracta, integra, 
ad basin in petiolum crassum brevem subcuneata 3-pli-nervia; 
pedunculi elongati solitarii uniflori; breacteae externae 


} 
foliaceae venosee basi connatae ovatae acuminatae, internse 


} 


1933 Gleason, South American Melastomes 47 


' late ovatae milto breviores obtusae paulo recurvatse; hypan- 


thium campanulatum; calycis limbus erectus truncatus; petala 
magna alba anguste ovata acuminate; antherae semi-ovoideae 
in annulo cohaerentes basi longe calcaratae; ovarium toto 
adhaerens in rostrum conicum productum; stylus gracilis 
stigmate punctiformi. 

§ Pyxidanthus: a climbing vine, glabrous throughout; up- 
per branches stout, irregulerly angled with short inter- 
nodes; leaves apparently equal in each pair, the blades ob- 
ovate-oblong, chartaceous, opaque, about 13 cm. long by half 
as wide, rounded above into a linear acumen 1 cm. long, en- 
tire, cuneate or subcuneate at base into a petiole 2-5 mm. 
long, 3-pli-nerved, %-pli-nerved, the obscure secondaries 
about 1 mm. apart, the lower surfece minutely punctate; ped- 
uncles solitary, 1-flowered, about 3 cm. long; outer bracts 
connate for 9 mm. at base, the free portion broadly ovate, 
36 mm. long, 24 mm. wide, foliaceous, acuminate, about 5- 
nerved, the midvein elevated on the back; inner bracts free, 
c losely appréssed to the hypanthium, broadly ovate, 20 mn. 
long, 16 mm. wide, obtuse, recurved at the summit; hypanth- 
ium campanulate, 12.5 mn. long to the torus; calyx-limb 
erect, truncate, 2.5 mn. wide; petals 6, white, narrowly 
ovate, 40 mn. long, 12 mm. wide, long-acuminate; filaments 
10 mm. long; anthers violet, cohering in a ring, stoutly 
gemi-ovoid, opening by 2 terminal pores, the thick connect. 
ive prolonged basally below the attachment of the filament 
into a triangular ascending spur 5 mm. long; ovary wholly 
inferior, its concave summit produced at the center into a 
slenderly conic beak 5 mm. high; style slender, 18 mn. long, 
Slightly decurved; stigma punctiform. 

Type, Klug 1862, collected in forest at Umbria, Comisaria 
del Putomayo, Colombia, alt. 325 m. Its connate outer bracts 
place it at once into the section Pyxidanthus, within which 
only a few species are known. None of these even approaches 
our species in the size of flowers or bracts. 

MICONIA MUTISELLA sp. nov. § Amblyarrhena; frutex ramosus 
internodiis brevibus; folia coriacea parva ovata denticulata 
utrinque rotundata J3-nervia, supra glabra, subtus tomentel- 
la; paniculae parvae folia vix excedentes; flores 5-meri; 
hypanthium tomentellum; sepala triangularia acuta, dentibus 
exterioribus crasse conicis patulis. 

Apparently a dwarf shrub of high altitudes, the stems 
stout, densely and somewhat fastigiately branched, terete, 
densely brown-tomentulose when young, glabrescent in age, 
the internodes above only 3-6 mm. long, or 15 mm. long on 
the oldest parts; petioles stout, 1-2 mm. long, tomentulose; 
leaf-blades thick, ovate, 7-9 mm. long, 4-6 mm. wide, round- 


_ @d at both ends, denticulate with blunt teeth 0.5-0.8 m. 
apart, 3-nerved with an additional pair of marginal veins, 


ae 


ha 


48 PHYTOLOGIA a ye 


glabrous above with obscure venation, sparsely stellate— 
escent beneath, especially on the prominently elevated f 
maries, the secondaries and tertiaries not differentiated 
panicles terminal, crowded, few-flowered, brown-stell ‘ 
tomentose, scarcely exceeding the leaves; flowers S-nerot 
subsessile; hypanthium campanulate, 2 mm. long to ‘the 1 
densely stellate-tomentose; calyx-tube prolonged 0.3 mm. 
rounded sinuses, pubescent like the hypenthium, the sep ale 
triangular, acute, 0.5 mm. long, exterior teeth stoutly eae 
obliquely conic, spreading 0.3 mm. at right angles to the $4 
sepals; petals oblong-ovate, 2 mm. long; stamens isomorphic 4 
filaments flat, 0.8 mm. broad, abruptly narrowed a be- a 
low the anther; anthers stoutly oblong, 1.3-1-4 . long, >a 
opening by a minute terminal pore, the connective « iéva ed = 
into a low dorsal ridge, neither lobed nor prolonged at th : 
base; ovary half-inferior; style stout, glabrous, 25 i 
long; stigma punctiform. 
Type, Mutis 2457, collected presumably in Colombia and Z 
deposited in the United States National Herbarium, no. © i 2: 
1561606. The structure of the anthers confirms its positior 
in § Amblyarrhena, within which it differs from all known 
species in its minute leaves. met 
MICONIA BUGENIOIDES Triana was originally described ‘from 
the upper Rio Negro. In the same paper Triana eiso deser ibe 
Oxymeris cuspidata from two types, one from Rio de Janeiro 
and the other from San Gabriel on the Rio Negro, Careful ex- 
amination of type material of both species, Spruce 2263 and 
3531, reveals only one small point of difference between ~~ 
them, except differences in dimensions which may well be ex— 
pected from material in quite different stages of develop- 
ment. In the type of M. eugenioides the appendage of the . 
larger stamens is triangular, while in Leandra (Oxymeris) 
cuspidata it is truncate at tip. The two species are accord E 
ingly united. Nomenclatorially their union offers no diffi- 
culty, but some question may be raised on the propriety of — 
uniting two plants which were placed in different genera be 
such distinguished students as Triana and Cogniaux. The re 
nus Leandra differs from Miconia essentially only in its 
acute petals. Most species of the genus have a different — 
habit, and may often be distinguished at a glance, but in 
several cases they approach each other rather closely. The 
anthers of Leandra are regularly unappendaged; in Miconia a 
both simple and appendeged anthers exist. The anthers of our 
species are distinctly miconioid in structure and are pract— 
ically duplicated in numerous other species of § conten 
The habit of our species also resembles Miconia mch more 
than Leandra. The name M. eugenioides Triana therefore 4 
stands for the species. The nearest relative of the plant is 
M. tetrasperma Gl., also with acute petals. e* 


~~ 


a 


~~ 


pam 

‘ef 

~~ ae 
4 


a 
yn 


PHY TOLOGIA 


Designed to expedite botanical publication 


July, 1934 No. 2 


CONTENTS 


Ho. Russy: New Species of Plants of the Ladew Expedition to 


Bolivia By Naat SIE HR CR A A SH A ANI Wis Sees WO ate wot eB 4 49 

B. SmitH: Taxonomic Notes on American Phanerogams—I ..... 2 Sh 
von Remnant: AR Cmce OF) POMBE. v's sie dan tas aie wip ob PAE Rate eh oe ass 83 
ERoy AsrAMs: The Mahonias of the Pacific States ................ 89 


N, Mo.vENKE: Studies of New and Noteworthy Tropical Amer- 


: ican Plants—I I “Ome ae LAAN ee oe Cia ey. ALUM TR, URGE eRe Coa 95 
A. Gueasow: Plantae Krukoviande—IIl .......00000ccveveeeeees 106 
A. Gizason: Note on the Genus Goethalsia Pittier ...........-.. 112 


"Published by H. A. Gleason and Harold N. Moldenke 


1 AN The New York Botanical Garden 
eit Bronx Park, New York, N. Y. i 
rice of, this iicaiers : 75 cents; per volume $5. 00 i in pin Neate nf 
Vol. I, number 1, was issued December 4, 1933. 


vier ARh Aes 
eects 
A ASNT! Lava 


NEW SPECIES OF PLANTS OF THE LADEW 
EXPEDITION TO BOLIVIA. (a) 


Henry Hurd Rusby 


In 1926, through the generosity of Mr. Harvey S. ladew, 
the American Museum of Natural History was able to send a 
small biological expedition to Bolivia. The territory to be 
explored was the Amazonian slopes of the Cordillera Real, 
the principal objectives being the assemblage of representa- 
tive collections of the mammals, birds, and plants of the 


region. The botanical collections were made by Mr. 


o H. He 


Tate and included 1275 numbers, of which the following are 
new. to science. The types of these new species are deposit- 
ed in the herbarium of The New York Botanical Garden. Ex- 
cellent collector's notes accompany all specimens, and it is 
to be regretted that want of space forbids the publication 
of the entire list, as well as Mr. Tate's interesting sketch 


of the journey. 


Mr. Tate has compiled the following table showing the 


geographical distribution of the collections: 


First trip to Yungas, Bolivia 


La Cumbre 15,000 ft. 
Alaska Mine 13,500 ft. 
Pongo 12,000 ft. 


148 
49.102 
105=275 


(mumerous mosses also, but unnumbered). 


Maviri-Tipuani trip, Bolivia 


Altiplano between La Paz 14,000 - 
and Sorata 15,000 ft. 

Sorata 8,000 ft. 

Cocopunco 10,000 ft. 


Road from Cocopunco to Pararani 
(Pararani about 5,000 ft.) 
Road from Pararani to Mapiri 


es Mapiri 2,000 ft. 
=_ Rio Chimate 1,900 ft. 
a 6CY) 

= ©? Guanay 1,800 ft. 
a Road from Guanay to Carahuarani 

rey na Cmaquini 

es Ticunhuaya 4,800 ft. 
© — Simacu 5,800 ft. 
ee on La Joya 5,900 ft. 
¥ Okara 7,500 ft. 


4g 


998-1003 
767-779 
274-385 


386-392 
ie 99 
00-498 
499-552, 
593-613 
553-592 
1213-1215 
1115-1195 
1051-1114 
No numbers 
1004~ 1050 
890-997 


50 PHY TO.b0 Gis Noe 2 


Road from Okara to Ancoma 836-889 
Tipuani trail, from top of divide, 
17,000 ft., down to Sorata 780-835 
Second trip to Yungas, Bolivia 
Pi tiguaya 72 1-766 
Nequejahuira 8,000 ft. 614-689 
Rio Aceramarca 11,500. ft. 690-720 
Peru 
' Arequipa 7;000' £4. 1196-1212 


COMMELINA REFLEXA Rusby, sp. nov. Caules vix ramosi 
erecti crassi glabri vel glabrati; laminae quam vaginis 3—4- 
plo longiores acuminatae et acutae; spatha valde reflexa ad 
angu lam 90° a pedunculo; ramlus floriferus spatham aequans 
densiflorus; capsula ovalis 5 mm longa 3 mm. diametro. 

Nearly glabrous, or the upper stem portion more or less 
puberulent; stems stout, erect, sparsely branched, the 
branches erect; leaves few; sheaths a fourth to a third as 
long as the blades, thick and rigid, about 1l0-ribbed; leaf- 
blade to 12 em. by 2.5 cm., lanceolate, long-acuminate and 
acute, the upper ones erect, the lower spreading; peduncles 
elongated, stout, erect; spathe turned at a right angle to 
its peduncle, in fruit 3.5 by 2.5 em. in the folded state, 
scarcely acuminate, acute or obtusish, about 15-nerved, the 
nerves slender, sharp, connected by numerous fine straight 
veins; lower branch of the cyme about 13 mm. long, naked, 
rigid, the floriferous branch about two-thirds of the length 
of the spathe, in its incurved position, but about equaling . 
it when straightened, densely flowered, the flowers not 
seen; pods 5 mm. long, 3 mm. broad, oval. 

First collected by 0. E. White at Pongo de Quime, Boliv- 
je, alt. 11,500 ft., in July, 1921 (Mulford No. 172). 
"Growing in full sunshine, about 12 inches high, the flowers 
blue." Collected by G. H. He. Tate on the road from Okara to 
Ancona, alt. 11,000 ft., April 29, 1926 (No. 847). 


COSTUS LONGIFOLIUS Rusby, sp. nov. Caules debiles inter- 
nodiis 2— 3 om. longis 6—7 mm. crassis; vaginae apice brev— 
iter pilosae et ciliatae; petioli breves torti pubescentes; 
laminae lanceolatae utrinque acuminatae acutae tenues; spica 
late ovoidea squamis patulis late ovatis acutis crassis; 
calyx infimdibuliformis dentibus late triangularibus mucron- 
atis. 

Stem rather slender, weak, its height unknowm; internodes 
2 or 4 em. long, 6 or 7 m. thick, the sheaths light brown, 
the summit slightly oblique, sparsely short-pilose and 
short-ciliate; petioles 1 cm. long, light brown, twisted, 
pubescent, nerveds blades to 25 by 4 cm, lanceolate to ob- 
lanceo late, acurrinate at both ends, acute, very thin, bright 


ee ee ee a ne a a ee 


1934 Rusby, New Species from Bolivia 51 


green, glabrous above, softly pubescent beneath, but becom 

ing nearly glabrous with age; spike broadly ovoid, 8 em. 

long by 9 cm. broad (in my specimen), the scales spreading, 
‘ to 5 by 2.5 cm., ovate, with broad base and acute summit, 
very thick and coriaceous, finely puberulent; bractlets 2.5 
| em. long, lanceolate, retuse at the summit, strongly com 
; planate, about half enclosing the fruit, many-nerved; ca lyx 
1.5 cm. long, 8 mm broad at the mouth (as pressed), many- 
nerved, regularly infundibular, the teeth 3 mm long, 
broadly triangular, tipped by a dark-colored mucro; summit 
of ovary and base of calyx sparsely short-pilose; fruit ob- 
ovoid, 1.5 by 1 cm.s3 seeds 3.5 mm long, 2 mm. broad, obov- 
oid, truncate, black, tuberculate, mostly enclosed in the 
thin, white, hyaline aril, which is longer than the seed on 
one side. 

Collected by G. H. H. Tate at Mapiri, Bolivia, alt. 2,000 

ft., March 30 to April 6, 1926 (No. 442). 


COSTUS TATEI Rusby, spe nov. Caulis debilis puberulus; 
ligula auriculata pilosa; lamina obovata micronulata ci lia- 
ta; spica ellipsoidea squamis adpressis subrotundis; calyx 
4mm. longus campanulatus dentibus brevissimis. 

Pubescent or puberulent, the leaves ciliate; stem weak, 
its height unknowm; intemodes 4 or 5 cm. long, the sheaths 
& to 10 mm broad, mimtely puberulent, the summit slightly 
oblique, the Hgule shortly auriculate above the base of 

‘4 the leaf, its margin more or less pilose; leaf thickish, 
sessile, to 17 by 7 em, obovate, with acutish base and a- 
: cutely micronate summit, the margin strongly ciliate with 
hairs similar to those which clothe the upper surface of 

f the midrib, the upper surface otherwise glabrous, the lower 
surface puberulent or pubescent, the transverse lines ex- 
§ ceedingly numerous, almost as wide as the spaces between 

z them; spike ellipsoid, 8 by 3.5 em. (in my specimen), the 
scales puberulent and closely appressed, 2 cm. long and 
broad, subrotund, obtuse; bractlet 1 cm. long, oblong, a- 
cutish, complanate, not enclosing the flower; immature 
fruit 5 mm. long, 2 mm broad at the trumcate summit, tub- 
erculate; calyx i, mn. long, broadly campanulate-turbinate, 
s subtruncate, the teeth extremely short; remainder of 
flower not seen. 

Collected by G. H. H. Tate at Rio Chimate, Bolivia, 
alt. 1900 ft., April 1 to 14, 1926 (No. 523, the type). No. 
Ab1, from Mapiri, alt. 12,000 ft., March 30 to April 9, is 
the same species. This specimen exhibits a cylindric spike 
9 by 3 cm. Its leaves reach 25 by 10 ome No. O47, of the 
Mulford collection, from the Bopi River, alt. 2200 ft., 
August 8, 1921, collected by 0. E. White, is the same spe- 
cies, as is also No. 1808, from Rurrenabaque, November 25. 
These latter specimens are much more pubescent than the 


~~ 


— 


52 Soo P BET Ob 0:6:TA No. 2 


Tate plant. The species is well distinguished by the ciliate 
margins and midribs of the leaves. 


MYRIOCARPA TATEI Rusby, sp. nov. Asperrime denseque pub- 
escens; caules crassi atque debiles; petioli albido-setosi; 
laminae tenues ovatae abrupte acuminatas acutae basi obtusae 
serrataeé; spicae numerosae elongatae saepissime simplices 
densiflorae floribus divaricatis albido-pilosis; ovarium 
late ovoideum stylum subsaequans. 

Densely and shortly rough-hairy throughout; branchlets 
stout, but weak and crooked, reddish-brown and gray—hairy, 
the hairs matted together; stipules early deciduous, 1 cm. 
‘long, ovate, acuminete and acute; petioles (only the upper 
leaves seen) to 4 cm. long, slender, white setose-hairy, the 
hairs divaricate; blades to 10 by 6 cm., ovate, with blunt 
or rounded base and abruptly short-acuminate and acute sum 
mit, finely and regularly serrate, the teeth ovate, broader 
than long, acute with a whitish tip, the sinuses acute, very 
thin, %nerved from a little above the base, the upper sur-— 
face densely and shortly rouch-hairy, dark green, the lower 
surface grayish-green, more densely hairy but less rough, 
the venation finely reticulate, not prominent; spikes very 
numerous, simple or branching near the bass, 15 to 20 cm. 
long, 5 mm. broad, densely flowered, the flowers divaricate, 
all the parts pilose with spreading white hairs; bractlets 2 
and linear, slightly longer than the slender stipe, which is 
half the length of the broadly ovoid ovary, the style about 
equaling the ovary, the stigma mostly recurved. 

Collected by G. H. H. Tate at Nequejahuira, Bolivia, alt. 
8000 ft., May 15 to 24, 1926 (No. 645). 


PHENAX BULIATUS Rusby, sp. nov. Rami elongeti graciles 
adscendentes subglabrati; folia lanceolato-ovata acuminata 
acuta ad basin rotundata, dentibus mucronulatis, pagina sup- 
eriore bullata saepe nitente; bracteae ovales obtusae con- 
cavas; pistillum sessile ovoideum, stylo elongato. 

Subglabrous; branches elongate, very slender, erect or 
strongly ascending, licht red, terete, very leafy; stipules 
3mm. long, triangular-acuminate, scarious, reddish; peti- 
oles to lcm. long, very slender, minutely scurfy; blades to 
3.5 by 1.5 cm, lance-ovate, with rounded base and acuminate 
and acute summit, serrate with mucronulate teeth, deep- 
green, the upper surface mostly bullate, often shining, with 
the venation impressed, very minutely puberulent, the lower 
surface mostly brown or reddish, densely papillose, the ven- 
ation lightly prominent, the three ribs originating in the 
petiole, connected by slender crooked secondaries; heads 
Bypherical, nearly 1 cm. broad, reddish or rich-brown; bracts 
brown, scarious, 1.5 to 2 mm. long end more than half as 
broad, oval, obtuse, conceve; staminate flower very shortly 


1934 Rusby, New Species from Bolivia 53 


stipitate, the calyx parted below the middle, the lobes 2 
mme long, faintly 3-nerved, broadly oval, the summit rounded 
and mucronate; stamens 4, longer than the calyx; rudimentary 
ovary minute, free from the stamens; pistillate flower ses- 
sile, the ovary less than 1 mm. long, ovoid, the style more 
than 3 mm. long, very slender. 

Collected by 0. Buchtien at Cotana, Bolivia, alt. 2700 
m., in November, 1911 (No. 3158). "A shrub about 2 meters 
high." Also collected by G. H. H. Tate at Nequejahuira, alt. 
8000 ft., May 15 to 24, 1926 (No. 649). The epOgeer is very 
near P, ballotaefolius Wedd. 


PHENAX FIAVIFOLIUS Rusby, sp. nov. Rami elongati grac- 
iles setosi; stipulae ovatae acuminatae acutaeque scariosae 
caducae; folia lanceolata acuminata acutaque basi acuta ser- 
rata flavo-brunnea; pistillum oblanceolatum stipitatum pub- 
escens. 

Setose-hairy and slightly hispid; branches elongated, 
slender, reddish-brown, terete, narrowly sulcate, very 
leafy, the hairs white and widely spreading or divaricate; 
stipules caducous, 4 or 5 mm. long, ovate, acuminate and 
acute, brown, scarious, pilose; petioles to 2 cm. long, 
slender, red, setose, deeply channeled above; bladea to 15 
by 4 cm., lanceolate with a short-acuminate and acute base 
and a long-acuminate and acute summit, finely serrate with 
short acute teeth and sinuses; heads to 8 or 10 m.,. broad, 
dense, brown; bractlets minute, hyaline; pistil short- 
stipitate, the ovary 1.5 mm. long, oblanceolate, the base 
tapering into the stipe, pubescent, obtuse, the style 
stout, tapering, pubescent, 2.5 mm. long; staminate flower 
not found. 

Collected by G. H. H. Tate at Pitiguaya, Unduavi, Boliv- 
ia, alt. 5800 ft., May 7 to 15, 1926 (No. 726). 


ALSINE YUNGASENSIS Rusby, sp. nov. Caules elongati deb- 
iles graciles glabri flavi nitentes; folia late sessilia ov- 
ata acuta apiculata tenuia; flores pauci solitarii penduli 
aut recurvi; sepala late ovata acuta; petala quam sepalis 
staminibusque paullo longiora; stamina stylos aequantia. 

Glabrous; the stems elongated, weak and slender; branch- 
es elongated, ascending, yellow, shining; leaves sessile by 
a broad base, 2 to 3.5 cm. long, 1 to nearly 2 em broad, 
ovate, acute, minutely apiculate or mucronulate, light 
green, drying yellowish, very thin, the venation slender, 
crooked, ascending and meeting to form a conspicuous line 
Close to the margin; flowers few, axillary, solitary, most- 
ly recurved or pendulous on long peduncles; sepals 6 m. 
long, broadly ovate, acute, thin, pale green, veiny; petals 
8 mm. long, bifid, the divisions narrow; stamens shorter 
than the petals, about equalling the styles; ovary 2 mm 


54 PHYTOL OG IA No. 2 


long, obovoid, the styles 3, shorter than the ovary, the 
upper part somewhat recurved, the summits slightly thick- 
enede 

Collected by H. H. Rusby in Yungas, Bolivia, alt. 6,000 
ft., in 1885 (Parke, Davis & Co. No. 1185, the type), dis- 
tributed as "S. cuspidata Willd.?", Also collected by G. He 
H. Tate at Pongo, alt. 12,000 ft., February 17 to March 1, 
1926 (No. 159). 


CERASTIUM BREVICARPICUM Rusby, sp. nov. Dense breviter- 
que pilosum; caules crassiusculi ex basi dense ramosi; folia 
crassa, basalia lanceolato-oblonga obtusa inferne angustata 
in basin longum pestioloideum, caulina deinceps minora et 
brevius petiolata; sepala oblanceolata obtusa petala parum 
excedentia; stamina quam petalis 1/3 brevioria pistillum 
subaequantia. 

Densely pilose or subtomentose throughout, with stout 
hairs; stems mostly 43 to 5 cm. long, densely branching from 
the base, thickish; leaves thickish, the radical ones 5 am. 
long, 8 mm. wide, the lower half being a long petiole-like 
base, the limb lance-oblong, obtuse, densely hairy and cil- 
jate, the cauline leaves successively smaller and with 
shorter basal portions; principal veins strongly ascending, 
the venation coarsely reticulate; sepals thickish, 1 cm. 
long, oblanceolate, obtuse, tomentose; petals a little 
shorter than the sepals, shortly 2-lobed, the lobes obtuse; 
stamens two-thirds the length of the petals; ovary brown, 5 
mm. long and more than half as broad, with rounded summit, 
10-ribbed and finely many-nerved; styles 5, half as long as 
the ovary; mature pod not seen, but it opens by 10 equal, 
short, obtuse teeth. 

Collected by G. H. H. Tate at the top of the Tipuani- 
Ancona-Sorata trail, alt. 16,500 ft., April 30, 1926 (No. 
785). Nos. 21 and 820 represent the same species. 


DRYMARIA IADEWII Rusby, sp. nov. In juventute minute 
puberulum; caules ramique gracillimi patuli; stipulee al. 
bidae lacerae; petioli filiformes; laminae ovatae obtusae 
mucronulatae basi truncatae vel subcordatae; flores pauci 
pedicellis filiformitus; sepala lanceolato-ovata rigida ac- 
uminata et acuta staminibus petalisque longiora; capsula 
globosa seminibus magnis. 

Younger portions and inflorescence very mimitely puberu- 
lent; stems elongated, very slender, the branches similar, 
divaricate or widely spreading; stipules whitish, irregular- 
ly lacerate, the divisions filiform; petioles of the largest 
leaves to 5 m,. long, filiform; blades very unequal, succes~ 
sively smaller upward, the largest 3 cm. long and nearly as 
broad, ovate, very obtuse, but minutely mucronate, with a 
truncate or shallowly cordate base, very thin, pale green, 


1934 Rusby, New Species from Bolivia 55 


glabrous, %nerved from the base, the venation slender, 
sparse; inflorescence cymose, very lax and few-flowered, 
the flowers on filiform pedicels; sepals 4 mm long, rigid, 


pale green with whitish margins, lance-ovate, acuminate and 


acute; petals two-thirds as long as the sepals, white, very — 
delicate, bifid, the divisions linear; stamens and pistil 
about equalling the petals; ovary globose, the style 4-fid 
about half-way; capsule splitting into 3 valves, the seeds 
large, brown. 

Collected by G. H. H. Tate at Nequejahuira, Bolivia, alt. 
8,000 ft., May 15 to 24, 1926 (No. 652, the type), and at 
Bara, alt. 7,500 ft., April 26 to 29 29 (No. 982). The species 
is very near D. pauciflora Bartl., but is is not at all pilose 
like that species. 


DRYMARIA STRICTA Rusby, spe nov. Breviter pubescens; 
caules ramique elongati graciles erecti; stipulae setaceae; 
petioli breves lati; folia ovata latioria quam longa basi 
subtruncata apice breviter acuminata et acuta; sepala et 
stamina a petalis excessa; capsula sepala aequans. 

Pubescent with short spreading hairs; stems to 3 dm. 
tall, slender and erect, like the branches; stipules very 


‘small, setaceous; petioles 1 or 2 mm. long, broad; blades to 


12 mm. long, broader than long, ovate with a subtruncate 
base and an abruptly short-acuminate and acute summit, thin, 
drying yellowish, 5—-nerved from the base; inflorescence, 
with its filiform erect branches and pedicels, pilose with 
divaricate hairs like the calyx; sepals 3 mm. long, oval, 
obtuse, pale green, rigid; petals exceeding the sepals, bi- 
fid, white, narrowed to the base; stamens about equalling 
the calyx; style trifid half-way down; capsule equalling the 
calyx. 

Collected by G. H. H. Tate at Pongo de Quime, Bolivia, 
alt. 12,000 ft., February 17 to March 1, 1926 (No. 160). 
The species is very near to D. glandulosa Presl, _ 


GUATTERIA SETOSA Rusby, sp. nov. Setoso-pilosa et sca- 
bra; folia subsessilia oblonga acuminata et attenuata sup- 
erne glabra; flores solitarii axillares, pedunculo calyceque 
setosis; petala exteriora sepala paullo excedentia, inter- 
jora longiora margine ad apicem excavata, omnia obtusa. 

More or less setose-hairy; branchlets much elongated, 
slender, harshly pilose or scabrous; leaves nearly sessile, 
the petiole about as broad as long, the blades to 14 by 3.5 
cm., oblong with an obtuse base, the summit acuminate and 
ending in an attenuate point about 5 mm. long, glabrous a- 
bove, sparsely pilose below with stiff appressed hairs, the 
venation lightly prominent beneath, the secondaries 15 to 18 
on a side, widely spreading, the ends strongly upcurved and 
connecting near the margin; flowers axillary, solitary, the 


56 PHYTOLO GIA No. 2 


peduncles 2 cm. long, setose like the sepals; calyx appar- 
ently valvate, coriaceous, the sepals 1 cm. long, broadly o- 
vate, obtusish; petals coriaceous, the outer slightly larger 


then the sepals, short-pubescent or tomentose, probably val- 


vate, obtuse, the inner longer and much broader than the 
outer, very obtuse, the margins slightly excavated near the 
summit, as though inclined to be 3-lobed, somewhat conni- 
vent; stamens densely massed, but not coherent, 1.5 mm. 
long, broadened upward, with a truncate summit; carpels 
forming a dense head 3 mm. broad. 

Collected by G. H. He Tate at Chuquini, Bolivia, alt. 
3,000 ft., February 17 to March 1, 1926 (No. 1138). 


DIOCLEA ORNITHORYNCHA Rusby, sp. nov. Pubescens pilis 
divaricatis vel retrorsis; petiolus brevissimus; foliola o- 
vales apice in rostrum longum abrupte contracta; calycis 
tubus late campanulatus ferrugineo-hirsutus, dentes longi 
falcati adscendentes. 

Pubescent with divaricate or somewhat retrorse hairs; 
stems rather stout, costate or strongly nerved; stipules 
not seen; petioles 7 or 8 cm. long, slender, narrowly chan- 
neled above; rachis 2 or 4 cme long; petiolules 5 mm. long; 
terminal leaflet 15 by 8 cm., oval with a rounded base and 
a summit abruptly contracted into a curved beak-like acum 
ination which is 2 cme or more long, entire, very thin, the 
very slender midrib and secondaries lightly prominent be- 
neath, the secondaries 5 or 6 on each side, strongly as- 
cending, lightly curved, connected by a loose reticulation, 
the upper surface sparsely, the lower more densely pubes- 
cent, the veins pilose with stiff divergent hairs; lateral 
leaflets about equal, similar but inequilateral; peduncle 
and rachis each about 2 dm long, the latter black-nodose 
from the fallen flowers. A single detached flower is pres- 
ent on my specimen. This has the pedicel 7 mm. long, slen- 
der; calyx-tube campanulate, 5 mm. long, 8 mm. broad,- 5— 
ribbed, hirsute and ferruginous, the long calyx-tooth near- 
ly 1 em. long, falcate-ascending; corolla-bud beginning to 
open, 1 cm. long and a little broader; dissection meterial 
wanting. 

Collected by G. H. H. Tate at Mapiri, Bolivia, alt. 2000 
ft., March 5 to April 9, 1921 (No. 479). The species is 
near to D. rufescens Benth., but the latter is wanting in 
the stiff divergent hairs on the veins of the lower leaf 
surface and has different flowers, 


LUPINUS BUCHTIENII Rusby, sp. nov. Puberulus; caules ex 
basi mumerosi prostrati vel adscendentes, ramosi et folio- 
si; stipulae lanceolatae abrupte attenuatae brunnese; fol- 
iola oblanceolata basi cuneata apice obtusa vel rotundata; 
racemi elongati simplices pedunculati; bracteae lanceolatae 


1934 Rusby, New Species from Bolivia — 57 


acuminatae quam pedicellis filiformibus duplo longiores; 
calycis labia subaequalia, labio inferiore 3-denticulato; 
corolla subcaerulea. 

Closely and finely puberulent; stems numerous from the 
summit of a long tap-root, prostrate or ascending, freely 
branching, 5 to 10 cm. long, slender, leafy, angled and 
nerved; stipules to 5 mm. long, lanceolate, sbruptly con- 
tracted into an attenuate summit, thin, brown, erect; pet— 
ioles very unequal, to 1 cm. long, filiform; leaflets most-— 
ly 7 or 9, sessile, the lower smaller, to 3 mm. long and 1 
mm. wide or less, oblanceolate with cuneate base and blunt 
or rounded summit, entire, gray-puberulent, the venation ob- 
scure; racemes terminating the branches, long and slenderly 
peduncled, simple, mostly 5S to 10-flowered, the pedicels 
about half as long as the lanceolate acute bracts; lower lip 
of calyx 3 mm. long and about as broad, ovate, S-nerved, 
minutely 3-dentate, the upper lip about as long, ovate, ac- 
uminate and acute, entire; corolla light blue, 7 mm. long; 
stamens 5 mm. long, the filaments united more than half-way; 
ovary 2.5 mm. long, lanceolate, the style exceeding the sta- 
mens, lightly curved; pod 1 cm. long, 5 mm wide, oval, com 
pressed, one margin thickened, tomentose, mucronate, 3 
seeded. 

Collected by Otto Buchtien at Unduavi, Bolivia, alt. 3500 
meters, in November, 1910 (No. 2869, the type). Distributed 
as Le prostratus Ag., which it closely resembles. Also col- 
lected by G. H. H. Tate at Pongo, alt. 12,000 ft., February 
17 to March 1, 1926 (No. 174). - 


LUPINUS TATEI Rusby, sp. nov. Cinereo-sericeus; caules 
alti crassi ramis adscendentibus; stipulae varie subulatae 
attenuataeque; foliola oblanceolata acuta basi acuminata; 
bractearum pars basalis late ovata, pars superior setacea; 
lobus calycis major ovatus acuminatus alii angusti acumin- 
ati et acuti. ' 

Densely gray-sericeous throughout; only the upper 
branches seen, these 2 or 3 dm. long, ascending, rather 
stout, floriferous at the summit; stipules more than 1 cm. 
long, narrowly subulate and long-attenuate; petioles to 5 
cm. long, slender, narrowly grooved above; leaflets appar- 
ently 7, sub-petiolulate, the outer successively smaller, 
the central to 6 by 1 or 1.2 cm., oblanceolate, acute with 
an acuminete base, entire, thin, densely sericeous on both 
surfaces; flowers loosely racemed, mostly 2 together; 
bracts in two parts, the basal broadly ovate, about 5 mm. 
long, the upper longer and setaceous; pedicels about 1 cm 
long, slender; calyx-tube 3 mm. long, 5 mm, wide, obtusely 
10-costate, the larger lobe 8 mm. long, ovate, long-acumin- 
ate and acute, the pthers narrower, 7 mm. long, acuminate 
and acute; vexillum orbicular, 1.5 cm. long and even broad- 


¢ 


58 PHY T O04 0-6: 4 No. 2 


er; alae obovate, equally long; keel 13 mm. long, finely 
many=nerved, ovate, obtusish, the summit darker; stamens a— 
bout as long as the keel, connate two-thirds of their 
length, the anthers lanceolate, 2 mm long; style 2 mm lon- 
ger than the stamens, the stigma bearded; pod 3 or 4 cm. 
long, 1 cm. broad, oblanceolate, compressed, hirsute, the 
edges thickened, mostly 5-seeded. 

Collected by G. H. He Tate on the road from Okara to An- 
cona, Bolivia, alt. 11,000 ft., April 29, 1926 (No. 855). 
The species is very near Le paniculatus Benth. and L. sora- 
tensis Rusby. 


MEIBOMIA ADHAERENS Rusby, sp. nov. Dense albo-hispida; 
cavles validi flexuosi srecti; stipulae late ovatae abrupte 
2.tenuatae longe pilosae quam petiolis longiores; foliola 
consimilia ovata obtusa minute apiculata; paniculae axil- 
lares et terminales laxiflorae; pedicelli recurvi; calycis 
labium superius quam inferiore fere duplo longius; legumen 
sub-f-articulatum suturis utrique invaginatis. 

Densely hispid with slender white, shining, divergent 
hairs which are mostly hooked at the tip; stems herbaceous, 
rather stout, flexuous, yellowish, apparently erect; stip- 
ules about 7 mm long, broadly ovate, abruptly contracted 
into an attenuation that ia nearly as long as the body, 
thin, brown, nerved, long-pilose; petiole to 5 cme long, 
slender, spreading, nerved, grooved on the upper surface, 
pilose, the rachis similar, 1 cm. long, the stipellae 4 m. 
long, lance-linear, attenuate; leaflets 3, the petiolules 
shorter than the stipellae; leaflets alike, the lateral 
somewhat smaller, to 4.5 by 3 cm., ovate with a broad base 
and summit, the latter mimitely apiculate, thin, pale green, 
sparsely pilose, the slender venation not prominent, the 
- secondaries 5 or 6 on each side, at an angle of about 45°, 
little curved except at the ends, the venation coarsely re- 
ticulate; panicles axillary and terminal, elongated, little 
branched and very loosely flowered; pedicels filiform, re- 
curved, 4 to 7 mm. long in flower, later slightly elongat— 
ing; upper lip of calyx nearly twice the length of the lower 
which is 3-lobed more than half-way, the lobes triangular, 
acute; vexillum 1 cm. long and broad, the claw short and 
very broad, cuneate; wings and keel shorter than the vexil~ 
lum; filaments united for most of their length; only one im 
mature pod seen, this 7-jointed, both margins indented but 
mot equally, strongly hispid, the joints small, inequilat— 
terally oval, a little longer than broad, 

Collected by G. H. H. Tate at Sorata, Bolivia, alt. 
10,000 ft., May 1, 1926 (No. 769). 


MEIBOMIA ADPRESSA Rusby, sp. nov. Minute puberula; cau- 
les basi radicantes atque adscendentes, ramis gracilibus e- 


1934 Rusby, New Species from Bolivia 59 


longetis, dense foliosis; stipulae adpressae ovatae longe 
attenuatae; petioli basi dilatati; stipellae minutae subu- 
latae; foliola ovalia obovata vel orbicularia; racemi term 
inales stricti pauciflori; legumina pauci-articulata, sutura 
ventrali paullo incisa dorsali alte sulcata. 

Very mimuitely and closely puberulent; stems rooting at 
the base, then ascending, branching from below, the branches 
long and slender, ascending, densely leafy; stipules 4 mm. 
long, appressed, ovate, with a long-attenuate summit; petio- 
le to 1 cm long, stout, mostly erect, dilated at the base, 
the rachis a fourth its length; stipellae subulate, very 
small; leaflets 3, the petiolules 1 mm. long, broad; blades 
varying from orbicular to broadly oval or somewhat obovate, 
with a roundeé summit and base, the lateral ones about two- 
thirds the size of the terminal, all deep green, thin but 
rigid, the margin thinly revolute, grayish-puberulent be- 
neath, the venation prominent beneath, secondaries crooked, 
the finer venation strongly anastomosing; racemes terminal, 
long-peduncled, slender and strict, few-flowered; pedicels 
filiform, about 8 mm. long, erect and incurved; longer lip 
of calyx 4 mm. long and nearly equalling the stamens, ovate 
with a long-acuminate summit, the other three lobes similar 
and only a little smaller; perfect pods not seen, apparent- 
ly 3-jointed, the ventral suture slightly and sharply 
notched, the dorsal deeply and widely intruded, the joints 
about 3 mm. long, 1.5 mm. wide, about twice as broad near 
the summit as near the base. 

Collected by G. H. H. Tate at Mapiri, Bolivia, alt. 2000 
ft., March 20 to April 9, 1926 (No. 468). The species is 
characteristic in its numerous subimbricated leaves, erect 
petioles and pedicels, and appressed stipules. 


MEIBOMIA NUMMULARIA Rusby, sp. nov. Dense pilosa; cau- 
les ex basi lignosa ramosi gracillimi; stipulae divaricatae 
late ovatae attenuatae; petiolis quam rhachide triplo long 
ior; stipellae setaceae; foliola subrotundae supra strigosa 
subtus molliter pilosa; racemi axillares minimi pauciflori. 

Densely pilose with soft white divergent hairs; stems 
branching from a woody base, in my specimen to 5 dm. long 
but probably growing much longer, the branches elongated, 
very slender; stipules 5 to 7 mm. long, divaricate, thin, 
brown, broadly ovate and attenuate, pilose, finely many- 
nerved; petioles to 4 cm. long, slender, thrice the length 
of the rachis; stipellae 3 or 4 m. long, setaceous; leaf- 
lets 3, the petiolules less than 1 mm. long; leaflets very 
irregular in size, from 5 by 5 mm to 2.5 by 2.5 em., the 
terminal one a little larger and relatively broader than 
the lateral ones, all subrotund, thin, bright green, the 
upper surface bearing appressed hairs with enlarged base, 
the lower surface densely soft-pilose; secondaries 4 on each 


60 PRY TOL OS Gls No. 2 


side, strongly ascending, nearly straight; racemes axillary, 
very small and few-flowered, the flowers not yet developed 
in my specimen. 

Collected by G. H. H. Tate at Ia Joya, Bolivia, alt. S00 
ft., April 25 to 26, 1926 (No. 1021). 


MEIBOMIA SIMPLICIFLORA Rusby, spe nov. Pubescens; rami 
graciles elongati; stipulae ovatae acuminatae et acutae pat- 
ulae vel recurvae; vetiolus quam rhachide triplo longior; 
foliola ovata obtusa vel apiculata pubescentia, pilis ad- 
pressis basi bulbosis; recemi elongati simplices vel subsim- 
plices laxiflori; calycis dentes ovati acuminati; legumen 
sutura una alte altera paullo invaginatum. 

Pubescent with short, fine, white hairs; stems slender, 
terete, reddish, the branches elongated and slender; stip- 
ules 5 mm. long, broadly ovate, abruptly long-acuminate and 
acute, brown, spreading or recurved; petiole 2 to 4 em. 
long and thrice the length of the rachis, slender, the stip- 
ellae similar to the stipules, but half as large; leaflets 
3, the terminal one from 4 by 2 cm. to 6 by 3 cm, ovate 
with 2 rounded base and acute or obtusish and minutely apic- 
ulate summit, thin, deep-green, sparingly pubescent on both 
surfaces, the hairs anpressed, with bulbous bases, the sec- 
ondaries 6 to 8 on each side, strongly ascending, lightly 
curved; lateral leaflets similar and about two-thirds as | 
large, on petiolules about 1mm. long; raceme terminal, e- 
longated, simple or nearly so, loosely flowered; longer lip 
of calyx 3 mm. long, ovate, attenuate, the other teeth sim 
ilar, about half as long, less attenuate; corolla absent; 
stamens 6 mm. long; pod mostly 5=- or 6-seeded, nearly 
straight, 2 to >.5 cm. long, 2 mm. wide, shortly stipitate; 
ventral sinus not at all or very little intruded, the dorsal 
sinus deeply so, the joints triangulate, hispid, very small. 

Collected by G. H. H. Tate at Okara, Bolivia, alt. 7,500 
ft., April 26 to 29, 1926 (No. 988). 


MEIBOMIA TETRASPERMA Rusby, sp. nov. Hispidulum et pilo- 
sum; rami elongati graciles patuli; stipulae subulatae att- 
enuatae 5— /-nervae; foliole ovata obtusa apiculata; panic- 
ulae magnae late patulae laxiflorae; pedicelli divergentes 
vel reflexi; calycis dentes aciuminati et acuti; filamenta 
fere ad apicem conjuncta; legumen reflexum saepe 4~articule- 
tum, sutura ventrali paullo, dorsali alte invaginata. 

Minutely and sparsely hispidulous with very short, re- 
curved hairs and a few long straight slender ones; stem her- 
baceous, very slender, apparently very long, with elongated 
and widely spreading branches; stipules 3 or 4 mm. long, : 
subulate and attemate, sharply 5- or 7-nerved; petiole to 3 
cm. long, nearly filiform, the rachis more than half as long 
as the petiole; stipellae very small, the petiolules 2 or 3% 


1934 Rusby, New Species from Bolivia 61 


mm. long, thick, pilose; blades very thin, pale-green, the 
terminal to 6 by 3.5 cm., rhomboidally ovate, blunt, minute- 
ly apiculate, the lateral ones considerably smaller, ovate, 
sparsely pilose with appressed hairs, the venation incon 
spicuous, the secondaries ahout 6 on each side, strongly 
falcate-ascending, rather crooked; panicles large, widely 
and sparsely branching, the branches filiform, loosely- 
flowered; pedicels 1 to 1.5 cm. long in flower, 2 cme long 
in fruit, divergent or lightly reflexed; upper lip of calyx 
1.5 mm. long, deeply 2-toothed, the teeth triangular- 
lanceolate, the lower 3-toothed, the lateral teeth much 
like the upper, the middle tooth a half larger, all acumin- 
ate and acute; vexillum nearly 1 cm. long, rose-colored, 
the claw very short, the keel and wings about two-thirds as 
long and nearly as long as the stamens, which are united 
nearly to the summit, shorter than the pistil; stigma cap- 
itate, flat; pod reflexed on the pedicel, mostly 4 jointed, 
the ventral suture slightly, the dorsal suture deeply in- 
truded and almost meeting, the joints very much flattened, 
With thickened densely ciliate margins, the joints nearly 1 
em. long and half as wide, reticulately veined, the seed 
attached at the middle, the long axis of the seed parallel 
with that of the pod. 

Collected by G. H. H. Tate at Okara, Bolivia, alt. 7500 
ft., April 26 to 29, 1926 (No. 987). 


OXALIS DENSISSIMA Rusby, sp. nov. Pubescens; caules 
dense caespitosi; stipulae brunmneae tenues ovatae obtusae 
inferne ad petiolum adnatae superne divaricatae; petioli 
albo-pilosi; foliola sessilia basi cumeata subtiliter ob- 
cordata; pedunculi erecti uniflori medio bibracteolati; 
calyx corollaque concolores. 

Finely pubescent; stems caespitose, very short, grow- 
ing in dense masses with mosses; stipules 1.5 to 2 mm. 
long, ovate, obtuse, adnate to the base of the petiole, 
divaricate, brown, thin and hyaline; petioles to 1.5 cm. 
long, slender, white-pilose; leaflets 4, sessile, equal, 
2mm. long, 3 mm wide, cuneate at the base, shallowly 
obcordate, pilose, rather thins; peduncles filiform, pilose, 
erect, 3 or 4 cm. long, 2-bracted about the middle, one- 
flowered, the bracts lance-linear, ecuminate; sepals col- 
ored like the corolla, 5 mm long, oblong, with rounded 
summit, finely nerveds; corolla 14 mm, long, purple; stamens 
monadelphous, very unequal, the longest reaching to the 
middle of the corolla, the filaments short-pilose; ovary o- 
val, about 2 mm long; styles 4, as long as the ovary, 
distinct, slender; stigma capitate. 

Collected on the Ladew Expedition by G. H. H. Tate at Co- 
copunco, Bolivia, alt. 10,000 ft., in March, 1926 (No. 383). 
Nos. 363 and 323 grow together and are badly mixed in the 


62 PHYTOLOGIA No. 2 


specimens. No. 383 is well distinguished by its longer peti- 
oles, larger and thinner leaflets, smaller and thinner stip- 
ules of different form, and its non-thickening petiole base. 


OXALIS TATEI Rusby, spe nov. Molliter pubescens, pilis 
purpureo—brunneis vel ferrugineis; caules caespitosi erecti 
uniflori inferne stipulis crassis brunneis ovatis imbricatis 
dense obtectis, his cum basi adnata petioli post lapsum fol- 
iorum persistentibus, petiolo deinde in corpusculum durum 
inter stipularum bases accrescente; corolla calyxque purpur- 
ati. 

Softly pubescent, the indumentum somewhat ferruginous or 
brownish-purple; stems gregarious, one or more cm. tall, the 
lower portions densely imbricated by persistent stipules, 
one-flowered; stipules 2 mm. long, ovate, acute, brown, thick 
and somewhat coriaceous, adnate to the base of the petiole, 
which eventually disarticulates at the point of union, the 
stipules and base of the petiole persisting, the latter en- 
larging and thickening to form a hard body between the stip- 
ules; free portion of petiole 3 mm. long; leaflets 3, sub- 
sessile, the terminal one a little larger, 2 mm. long, 3 mm. 
wide, triangular-obcordate with a shallow sinus, thick and 
somewhat fleshy, deep-green; peduncles about 2 cm. long, 
filiform, pilose, two=bracted about the middle; sepals 4 to 
3.5 m. long, lance-ovate, obtuse, somewhat unequal, colored 
like the corolla; corolla 13 mm. long, apparently purple; 
stamens monadelphous at the dilated bases, 5 of them 4 mm. 
long, the others 6 mm. long and pilose, those of either set 
sometimes umequal among themselves; pistil 2 mm long, the 
oval ovary two-thirds of the length; styles 4, distinct, 
rather stout, spreading; stigmas capitate. 

Collected on the Ladew Expedition by G. H. H. Tate at 
Cocopunco, Bolivia, alt. 10,000 ft., March 24 to 29, 1926 
(No. 323). The peculiar origin of the scales clothing the 
stem-bases sheds light on the relationship of the species 
of "Oxalis" with scaly rhizomes and scaly bulbs. Perhaps 
this species and 0. densissima represent a genus not as yet 
segregated. 


OXALIS VIRGATA Rusby, sp. nov. Puberula et parce ferru- 
ginea; caules lignosi erecti crassi foliosissim; stipulae 
erectae lanceolatae acuminatae et acutae scariosae brun- 
neae; petioli graciles subtus ad apicem glandule magna or- 
nati; foliola 3 consimilia triangulari-obovata sim brevi 
lato lobis rotundatis ad apicem costae glandula magna nota- 
tis; pedunculi in axillis superioribus solitarii saepissime 
uniflori; sepalea viridia; petala flava. 

Puberulent and somewhat ferruginous; stems woody, stout, 
terete, erect, very leafy; stipules 2 or 3 mm. long, erect, 
brown, thin and scarious, lanceolate, acuminate and acute; 


1934 Rusby, New Species from Bolivia 6% 


petioles about 1 cm. long, slender, erect or somewhat 
spreading, puberulent, with a large gland at the summit on 
the lower side; leaflets 4, often reflexed, sessile or near- 
ly so, equal or the terminal one a little larger, 7 mn. 

long (to the summit of the midrib), 6 mm. broad, triangular- 
obovate, the terminal sinus broad and shallow. the summit of 
the lobes rounded, with a large round brown gland at the 
summit of the midrib on the lower surface, the upper surface 
deep-green, puberulent with few minute whitish hairs, the 
venation slightly prominent beneath, ferruginous and pubes- 
cent, the midrib very stout, the secondaries 3 or 4 on each 
side, ascending at about 45°; peduncles few, solitary in the 
upper axils, mostly l-flowered, occasionally 2= or 3—flower- 
ed, filiform, to 4 cm. or more long, bearing linear- 
attenuate brow scarious bracts; sepals 3 mm. long, thin, 
green, lanceolate, with an attenuate summit, the midrib 
strong; corolla yellow, 12 mm. long; stamens very unequal, 
the longest ones half the length of the corolla; dissection 
material and fruit wanting. 

Collected on the Ladew Expedition by G. H. H. Tate at 
Cocopunco, Bolivia, alt. 10,000 ft., March 24 to 29, 1926 
(No. 282). The species is very characteristic because of its 
tall, stout, woody stems, densely clothed with leaves on 
short erect petioles. 


ALCHORNEA MEGALOSTYLIS Rusby, sp. nove Breviter albo- 
pubescens; caules petiolique rubri; folia crassa ovali- 
ovata, basi rotundata, apice abrupte breviter et obtuse ac- 
uminata, obscure sinuato-dentata; spicae foemineae axillar- 
es solitariae breves graciles pauciflorae; styli 2 rubri — 
crassi pubescentes complanati valde recurvi quam ovario mul- 
to longiores; spicae masculae graciles laxe ramosae. 

Pubescent with short white hairs; stem terete, reddish; 
stipules not seen; petioles to 5 cm. long, stout, reddish, 
nerved; blades to 15 by 8 cm, ovate or oval, with rounded 
or blunt base and very abrupt, short, obtuse acumination, 
shallowly sinuate-dentate, thickish and rigid, pale-green, 
above very shortly and sparsely pubescent on the veins, 
which are slightly prominent, more pubescent beneath, with 
the red venation very stout and prominent, 3-nerved, the 
secondaries 4 or 5 on each side, strongly ascending and 
lightly falcate, connected by numerous straightish tert- 
iaries and strongly looped together near the margin, the 
finer venation strongly anastomosing; pistillate spikes 
axillary, shorter than their leaves, simple, slender, very 
loosely flowered; calyx white, tomentose, deeply 4—parted, 
about one-third the length of the white tomentose ovary, 
which is broadly ovoid, about 1 mm. long; styles 2, thick 

and fleshy, red, pubescent, flattened, strongly recurved, 
much longer than the ovary; staminate plant more pubescent 


= 
we 
be 
a 
hy 


i 


a 
4 


64 PHYTOLOGIA No. 2 


and the leaves ciliate, the spikes loosely branched, very 
slender, the flowers glomerate at the nodes; calyx 2-parted; 
stamens 4 or 5, not united. 

Pistillate plant collected by M. Bang near Coroico, Bol- 
ivie, August 6, 1894 (No. 2375, the type). Staminate plant 
collected by  H. H. Tate at Nequejamira, alt. 8000 ft., 
“May 15 to 24, 1926 (No. 646). 


PAULLINIA TATEI Rusby, sp. nov. Glabra; rami graciles 
rubelli Z-anguiati; petioli graciles 7-angulati; foliola 
ovali-ovata basi in petiolum breviter producta apice obtusa 
remote sinuato-dentata; racemi axillares simplices, peduncu- 
lis elongatis recurvatis apice cirrhiferis; capsula obtuse 
triangularis parce lobata. 

Giabrous throughout; branches elongated, slender, reddish 
brown, triangular; leaves trifoliolate, the petioles 4 cm, 
long, slender, triangular; lateral petiolules 2 or 45 mm. 
long, the terminal a little longer; lateral leaflets 7 by 4 
cm., oval or slightly ovate, “ith very slightly produced 
base and obtuse summit, simuately dentate with few very 
short blunt teeth, slightly shining, the venation impressed 
above, prominent beneath, the secondaries about 7 on each 
side; terminal leaflet somewhat larger, lightly obovate; 
racemes axillary, mostly simple, on elongated, slender, re- 
curved peduncles, tendril-bearing at the summit, the rachis 
densely fruited or nodose from the fallen fruits; capsule 
obtusely triangular and lightly 3-lobed, with a broad flat 
summit and a short thecaphore at the base, 11 or l2 m, 
broad and not quite so long, red; seed 8 mm long, obovoid 
with a rounded summit, deep purple, shining, the aril light 
brown, papillose, cup-shaped, the margin truncated and 
sometimes irregularly ruptured, ebout half the length of 
the seed, 

Fruiting specimen collected by G. H. H. Tate at Guanai, 
Bolivia, alt. 1800 ft., April 14 to 16, 1926 (No. 553). 


SERJANIA LYRATA Rusby, sp. nov. Breviter ferrugineo= 
tomentosa; rami graciles 6-costati; laminae foliorum quem 
petiolis sub-6-plo longiores bipinnatae; foliola lanceola- 
to-oblonga basi breviter acuminata apice subacuta argute 
serrata utringue ferrugineo-tomentosa; paniculae graciles; 
carpellae maturae tenues parce puberulae. 

Shortly ferruginous—tomentose; branches rather slender, 
reddish-brown, 6-costate, unarmed; petioles about one- 
seventh the total length of the leaf, about 15 mm. long; 
blade bipimate, 10 or 12 om. long and nearly as broad at 
the base, the lower pinnae on short slender petiolules, 
their lateral pinnules 2 to 2.5 cm. long and half as broad, 
the lower on short petiolules, lance-oblong with an abrupt- 
ly short-acuminate base and acutish summit, serrate with a- 


> 


1934 Rueby, New Species from Bolivia 65 


cute teeth, occasionally the lowest pinnule pinnate, the 
terminal pinnule ovate with abruptly acuminate base, much 
larger than the lateral ones; upper pinnae sessile, occas- 
ionally with a single basal pinmule, similar to the terminal 
pinnules of the lower pinnae; terminal pinna lance-ovate, 
long-acuminate, variously incised, serrate, or pinnatifid; 
leaves thin, ferruginous-tomentose on ‘both sides, the vena- 
tion slender, abundant; panicles narrow, short—peduncled, 
the peduncles angular; fruiting pedicels 4 mm, long, slen- 
der, the carpels nearly 2.5 cm. long, 9 mm. broad, with a 
rounded base, light-green, thin, slightly puberulent, the 
veins horizontal or somewhat depressed, the summit rounded, 
the seed small, near the summit. 

Fruiting apevinen collected by G. H. H. Tate at Okara, 
Bolivia, alt. 7500 ft., April 26, 1926 (No. 991). 


WALTHERIA LADEWI Rusby, sp. nov. Dense et mlliter cin- 
ereo~stellato-tomentella; caules graciles erecti subsimp- 
lices; stipulae angustae basi paullo dilatatae; folia ovata 
_crenato-dentata basi rotundata apice obtusa vel mucronula- 
tas; racemi breves secundi recurvati; calycis dentes a basi 
acuminati. , 

Densely, closely, and softly gray-stellate-tomentel lous 
throughout; stems to 6 dm. high, slender, erect, simple or 
little branched, the branches erect, terete; stipules not 
seen; petioles 3 to 5 mm. long, slender, slightly dilated 
at the base; blades to 30 by 135 mm, ovate with a broadly 
rounded base and a blunt, minutely mucronulate summit, 
finely crenate-dentate with short broad mucronulate teeth, 
the upper surface. gray-green, very densely tomentellous, the 
venation finely channeled, the lower surface gray, with the 
slender midrib and secondaries sharply prominent, the sec- 
ondaries 5 or 6 on each side; flowers secundly racemose in 
the axils, the racemes much shorter than their leaves, 
shortly and slenderly peduncled and recurved; bracts 4A or 5 
mm. long, imbricated, lanceolate, acuminate, hirsute, cil- 
iate, thickish; calyx turbinate, 5 mm. long, toothed nearly 
half-way, the tube 10-nerved, the teeth acuminate from the 
base; petals 2 m. longer than the calyx; filaments 1 mm. 
long, completely coherent, the anthers 1 mn. long, mimtely 
mucronate; style pilose, 1 mm. longer than the stamens, 
plumose at the summit. 

Collected by G. H. He Tate at ar eet Bolivia, alt. 2000 
ft., March 30 to April 9, 1926 (No. 470). The species is 
near W. americana L. 


CAOPIA PARVIFOLIA Rusby, sp. nove Puberula, in inflor- 
escentia glandulosa; folia lanceolata in petiolum brevissi- 
mum abrupte contracta breviter acuminata et acuta supra 
glabra subtus parce ferruginea; pedicelli fructiferi grac- 


66 P.BY T.0-0.0 Gvaek’ No. 2 


iles; sepala reflexa lanceolata acuminata et subacuta;fruct— 
us insigniter glandulosi. 

Sparsely puberulent, the inflorescence glandular; 
branches very slender, angled; petioles very short, margin- 
ed; blades to 5 by 2 em., lanceolate, very abruptly con- 
tracted into a short petiole-like base and an abruptly 
short-acuminate and acute summit, thin, yellowish-green, a- 
bove glabrous, beneath finely puberulent with appressed 
hairs and somewhat ferruginous, the venation slender, light- 
ly prominent beneath, the secondaries 6 or 8 on each side;- 
panicles terminating the branches, small and loosely fruit— 
ed, the pedicels slender, 2 or 3 mm. long; sepals reflexed 
in fruit, 1.5 to 2 mm long, lanceolate, acuminate, acutish; 
fruit 7 mm. long, mostly broader than long, strongly gland- 
ular, mostly bearing the short and slender recurved styles; 
the remains of a flower show a yellow petal, 7 mm long, 
densely covered with red-brown glands. 

Fruiting specimen collected by G. H. H. Tate at Chuquini, 
Bolivia, alt. 3000 ft., April 17 to 19, 1926 (No. 1128). 


PASSIFLORA TATEI Killip & Rusby, sp. nove Pilosula; 
caules trigoni; stipulae lineari-falcati purpurascentes; 
petioli eglandulosi; laminae oblongae et latiores apice 
truncato obsolete 2— %lobato; pedunculis solitarii vel gem 
inati graciles; bracteae setacei 2— 4-fidae; flores circa 3 
em. lati; petala anguste linearia alba; corona biseriata ex- 
terior liguiata interior filiformis. 

Herbaceous vine; stem trigonous, densely pilosulous; 
stipules narrowly linear-falcate, 7 to 10 mm. long, 0.5 mm. 
wide, purplish; petioles 1 to 2.5 cm. long, glandless, pil- 
osulous; leaves oblong or suborbicular in general outline, 
5-5 to 10 cm. long, 3 to 7 cm. wide, obsoletely 2=- or 3- 
lobed at the truncate apex, rounded or shallowly cordate at 
base, entire, 4-nerved, subcoriaceous, lustrous and minute- 
ly puberulous above, dull red, densely pilosulous on the 
nerves and veins beneath; peduncles solitary or in pairs, 
3.5 to 4 cm. long, slender; bracts dissitate, 6 to 9 m. 
long, setaceous, deeply 2= or 3-cleft, purplish; flowers 
2-5 to 3.5 cm. wide; calyx-tube broadly campanulate, 7 to 
10 mm. wide at base; sepals linear or lanceolate, about 1.5 
em. long, 0.4 cm. wide, obtuse, greenish-white; petals nar- 
rowly linear, 7 to 9 mm, long, 1.5 mm wide, obtuse, white; 
corona in two series, the outer narrowly liguliform, about 
8 mm. long, the imer filiform, 2 to 3 mm. long; operculum 
closely plicate, about 1.5 mm high, denticulate, incurved; 
nectary ring annular; limen cupuliform, about 2 mm high; 
ovary globose, densely lanate-villose; fruit globose, seeds 
ovate-orbicular, transversely sulcate. 

Type collected at Nequajahuira, Cordillera Real, Bolivia, 
alt. 2500 meters, May 15 to 24, 1926, by G. H. H. Tate (No. 


. (oye 


1934 Rusby, New Species from Bolivia 67 


654). This species has the general appearance of P. bauhin- 
ifolia H.B.Ke, a species kmowm only from the mountains of 
central Ecuador, and the two sre obviously closcly related. 
In P. tatei the bracts are deeply cleft; in P. bauhinifolia 
entire; the calyx-tube is much broader in P. tatei, and the 
petals are narrowly linear, rather than ovate-lanceolate. 


CAIOPHORA MACROPHYLIA Rusby, sp. nove Breviter setosa 
pilis saepissime reflexis; caules volubiles cinereo-pubes- 
centes; petioli basi abrupte dilatati et inter se connati; 
laminae ovatae cordatae pinnatifidae lobis utrinque circa 4; 
pedunculi volubiles; sepala pinnatifida quam petalis sub- 
triplo brevioribus; stamina inaequalia. 

Setose with short and weak, mostly reflexed prickly 
hairs; stems rather stout, twining, terete, gray-hairy; pet- 
ioles (only the upper ones seen) 4 or 4 cm. long, the bases 
abruptly dilated and cormecting, the upper surface narrowly 
and deeply channeled; blades to 14 by 9 cm., ovate, cordate, 
pinnatifid, the lobes about 4 on each side, ovate, acutish, 
irregularly and shallowly dentate, the sinuses obtuse or the 
upper acute, very thin, bright-green on both surfaces, 
sparsely setose on both surfaces, the setae very unequal, 
the veins beneath densely setose-pilose, the venation weak, 
coarsely reticulate; peduncle terminal, to 4 dm. long, 


stout, twining, retrorsely setose; sepals 1 cm. long, lan- 


ceolate, incisely pinnatifid, some of the teeth gland- 
tipped, bright-green, setose-pilose; corolla light-scarlet, 


_ setose-pilose, the petals 4.5 cm. long, broadly oval, very 


veiny; stamens numerous, unequal, some of them two-thirds 
the length of the petals, the filaments filiform, the anth- 
ers elliptic, about 1 mm. long; pistil not seen. 

Collected by G. H. H. Tate at Pongo de Quine, Bolivia, 
alt. 12,000 ft., February 17 to March 1, 1926 (No. 185). The 
species is notable for its ample bright-green leaves and 
large handsome flowers. 


BEGONIA SUBRECTANGULA Rusby, spe nov. Caules cinerei 
papillosi; stipulae obtusae ovatae ex basi lata; folia ten- 
uia late ovata abrupte et brevissime acutata, minute serru- 
lata supra glabra subtus dense lepidota, costa a petiolo 
fere ad angulam rectam divergente, lobo basali rotundato in 
latitudine laminam propriam fere aequante. 

Stems coarsely angled, gray, papillose; stipules sessile 
by a broad base, to 2 cm. long, 1 cm. wide, ovate, obtuse, 
entire, spreading or reflexed, thin and membranaceous, brow 
and veiny; petiole to 43 cm. long, slender, gray, nerved, 
lightly papillose; total length of blade 6 to 12 cm. by 3 to 
6 cm. wide, the midrib nearly at right angles to the petio- 
le and 4 to 8 cm long, the main body of the leaf broadly o- 
vate, very anruptly contracted into a short and acute point, 


68 PHYTOLOGIA Noe 2 


the basal lobe rounded, nearly as wide as the main portion, 
the sinus deep, broad, acute, the margin finely serrulate, 
the leaf thin, deep-green, glabrous, but densely lepidote on 
the lower surface, the venation slender, sharply prominent 
on both sides, the basal nerves about 9, one or two second— 
aries from the lower half of the midrib, strongly ascending, 
nearly straight; pedicels filiform, nearly 3 cm. long; stam 
inate flower with sepals 2, subequal, broadly ovate, cor- 
date, nearly 1 cm. broad, thin and netaloid, about 20-nerv-— 
ed, the nerves faint; petals none; stamens numerous, dist- 
inct or barely united at the base, the filaments filiforn, 
scarcely 1 mm. long, the anthers linear-oblong, 2 mm. long, 
obtuse. The plant is apparently dioecious, no pistillate 
flowers being found. 

Collected on the ladew Expedition by G. H. H. Tate, May 
15 to 24, at Nequejamira, Bolivia, alt. 8000 ft. (No. 656). 
The species seems to be near B. sanguinea Raddi. . 


BEGONIA UNILATERALIA Rusby, sp. nov. Furfuraceo-tomento— 
sa et ferruginea; stipulae lanceolatae acumineatae in caudam 
longum filiformem; folia oblique ovata, abrupte breviterque 
acuminata, margine irregulariter leviterque crenato-dentato, 
sinu basilari fere obsoleto, costa a petiolo sub angulo 
circa 135°; pedicelli elongati filiformes; petala exteriora 
sepala fere aequantia et valde similantia, ovata et obtusa. 

Scurfy-tomentose with short, thick, ferm’cinous hairs; 
stems very Plemous, coarsely angled, light=-browm; stipules 
brown, thin and scarious, unequal, the longest about 12 m. 
long, lanceolate from a broad base, acuminate and bearing a 
terminal filiform appendage, finely nerved; petiole slender, 
3 or 4 cm long, scurfy-tomentose; blades to 7 by 43 cm., the 
midrib 6 cm. long, meeting the petiole at an angle of about 
135°, the blade obliquely ovate, the basal lobe rounded, the 
sinus almost wanting, the summit abruptly short-acuminate 
and acute, the margin irregularly and shallowly crenate- 
dentate, the teeth mostly acute, the leaf thin, sparsely 
scurfy on the veins above, scurfy-—tomentose on the veins be- 
neath, finely lepidote, the nerves about 9, from the base, 
with 2 or 3 secondaries ascending strongly from the midrib; 
inflorescence (only pistillate flowers seen) scanty and lax, 
the flowers on long filiform pedicels, the 2 outer sepals 
subequal, obliquely ovate, 1 cm. long, 6 mm. wide, obtuse, 
serrate, thin, nerved; petals of similar texture, unequal, 
‘the outer ones nearly equalling the sepals and mich like 
them, the inner 3 lance-ovate, obtuse, entire or slightly 
serrate; ovary and its 2 larger wings ferruginous-tomentose, 
ovoid, & mm. long and nearly as broad, the largest wing 8 
mm. long and broad, its upper margin nearly straight and 
almost horizontal with the surface of the ovary, obtuse at 
the end, whence it curves regularly to a point slightly be- 


1934 Rusby, New Species from Bolivia 69 


low the base of the ovary, the margin lightly dentate, the 
second wing 3 mm. wide at the summit, whence it narrows 
gradually and slightly, its lower end meeting the primary, 
along its inner margin is a narrow, brown, thickened line, 
the third wing is thin and hyaline, irregular and incom 
plete; styles 4, slightly united at the base, 5 mm long, 2- 
or %cleft nearly to the base, the branches again cleft or 
variously branched, the branches nearly smooth, tortuous or 
sinuous, flattened, stout and tough, thickened upward, ob- 
tuse@e 

Collected on the ladew Expedition by G. H. H. Tate at Ne- 
iti Bolivia, alt. 8000 ft., May 15 to 24, 1926 (No. 


657 


PARSONSIA SATUREIOIDES Rusby, sp. nov. Glanduloso-pubes- 
cens, pilis patulis; rami graciles foliosi; folia opposita 
sessilia linearia acuta basi angustata integra glabra; ped- 
icelli axillares solitarii graciles erecti, floribus reflex 
is; calyx infundibulariformis 6-costatus hispidus ‘calcare 
brevi rotundato, dentibus 6 trianguleribus. 

. Glandular—hairy; densely branched, the branches erect or 
ascending, slender, 5 to 10 cm. long, shortly and coarsely 
glandular-hairy, the hairs spreading or divaricate; leaves 
numerous, opposite, sessile, to 2 cm. long and 4 mm. wide, 
linear, acute, narrowed to the base, entire, glabrous, 
thickish, the midrib stout on the lower side, the venation 
obscure; pedicels solitary in the axils, slender, erect, 
about two-thirds the length of their leaves, slightly thick- 
ened upward; flowers more or less reflexed on the summit of 
the pedicel; calyx (in flower) about 5 mm. long, about 1 mm. 
wide at the summit, nearly straight, infundibular, about 6- 
ribbed, hispid, the basal sac very short and rounded, the 
teeth six, about one-sixth of the total length, triangular, 
acute, with acute sinuses, two teeth a little longer and 
each with a smaller tooth at the side; petals 6, red, sub- 
equal, 2 mm longer than the calyx, adnate to its summit, 
oblanceolate, obtuse, the claw rather broad, the basal 
glands very small; stamens somewhat unequal, reaching to the 
base of the calyx-lobes, the anthers oval; seeds 6, 1.5 mm. 
long, irregularly oval, flattened. 

Collected by G. H. He Tate at Rio Chimate, Bolivia, alt. 
1900 ft., April 10 to 14, 1926 (No. 541). 


BRACHYOTUM BARBIFERUM Rusby, sp. nov. Scabrum et levi- 
ter strigosum; rami rubelli anguste 4-alati alis hispidis; 
ramuli infra nodos saepe setas glomeratas gerentes; folia 
lanceolato-oblonga apice obtusiuscula vel saepius apiculata. 

Scabrous and somewhat short-strigose; branches red- 
purple, quadrangular, the angles narrowly winged, the wings 
hispid with short ascending hairs; branchlets bearing, on 


790 PEPE CL Cire i Nos 2 


the internodes or usually just below the petioles, occasion- 
al pairs of tufts or fascicles of coarse unequal setae; pet- 
ioles of the pair often unequal, to 1.5 om. long, slender, 
scabrous with short stout appressed hairs; blades to 6 by 2 
cm., or narrower, lance-oblong with obtuse or acutish base 
and obtusish mimutely apiculate summit, entire, thickish and 
rigid, 3-ribbed from the summit of the petiole, with a pair 
of secondary ribs from near the base of the outer ones, the 
ribs narrowly channeled on the upper surface, which is his- 
pid in four bands between the ribs and near the margins, 
with five glabrous bands at and beside the ribs; ribs and 
seconderies prominent on the lower surface, the secondaries 
about 40, with smaller intermediate ones, widely spreading 
and strongly falcate to meet in a sinuous line between the 
ribs; panicles terminating short branchlets, very slenderly 
pedunc led, their branches and branchlets strictly opposite, 
the rachis dilated at the point of branching; young buds o- 
void, acute, the calyx-teeth acute; flowers not seen. 
Collected by G. H. He Tate at Cocopunco, Bolivia, alt. 
10,000 ft., March 24 to 29, 1926 (No. 362, the type); also, 
without inflorescence, at the same time and place, No. 33l. 


PERNETTYA DENSA Rusby, sp. nov. Caules lignosi ramosis- 
simi juventute leviter pilosi; folia sessilia basi angusta- 
ta, coriacea oblonga obtusa crenulato-dentata; pedicelli 
fructiferi graciles divaricati; sepala rubella rigida quam 
fructu 5-lobato dimidio breviora. 

Young stems sparsely short-pi lose; stems woody, diffusely 
branched, the branchlets mostly 2 to 4 cm. long, densely 
crowded, very leafy, reddish; leaves sessile but with a nar- 
rowed petiole-like base, eoriaceous, thick and rigid, oblong 
and obtuse, to 7 by 2 m., pale-green above, reddish or fer- 
ruginous beneath and on the shallowly crenulate-dentate mar- 
gins, the midrib very stout and prominent beneath, the sec- 
ondaries erect or strongly ascending, crooked; flowers not 
seen; fruiting pedicels 2 to 4 mm. long, slender, spreading 
or mostly recurved, reddish; sepals puberulent, reddish, 
thick and rigid, about half the length of the fruit, ovate, 
acute; fruit above 4 mm. broad and shorter than its breadth, 
S-lobed, glabrous or puberulent when young, the stout per- 
Sistent style less than half the length of the fruit. 

Collected by G. H. He. Tate at Cocopunco, Bolivia, alt. 
10,000 ft., March 24 to 29, 1926 (No. 283). The species is 
apparently a shrub less than a foot high. 


PERNETTYA SCHIZOSTIGMA Rusby, sp. nove Rami mumerosi e- 
recti aut adecendentes nigrescentes breviter setosi; folia 
oblonga obtusa breviter obtuseque serrata; flores in axil- 
lis superioribus solitasii; stylus brevis superne incrasse— 
tus stigmate 5-lobato. 


1934 Rusby, New Species from Bolivia 71 


Much-branched, the branches short, slender, erect or as- 
cending, somewhat crowded, blackish, sparsely clothed with 
short, stout bristles; leaves sessile, but having a very 
short petiole-like base, about 15 mm. long by 3.5 to 4 m. 
wide, oblong, obtuse, the recurved margin serrate with short 
obtuse teeth, thick and rigid, pale-green, glabrous, the 
midrib and sometimes the veins impressed above, very strong 
and prominent beneath, the secondaries 3 or 4 on each side, 
erect or strongly ascending, very crooked, irregularly and 
sparsely anastomosing, sometimes with several bristly hairs 
on the midrib; flowers solitary in the upper axils, the pe- 
duncles about half as long as the leaves, angled, subulate- 
bracted, bristly; sepals 2 mm. long, ovate, spreading, thick | 
and veiny, mucronate and acute; corolla white, 4 or 5 mm 
long, two-thirds as wide, the teeth short, obtuse, recurved; 
filaments 2 mm. long, abruptly dilated and concave at the 
base, tapering upward, attached below the middle of the an- 
ther, which is less than 1 mm. long, broadly ovoid with a 
rounded base, slenderly 4-awned at the summit, the pores 
looking upward and slightly inward; ovary 2.5 mm. long and 
broad, green, the. style a little larger, stout, thickened 
upward, the stigma shortly 5-lobed, the lobes spreading; 
fruit red, papillose, 1 cm. broad and not so long. 

Collected by G. H. He Tate, on the ladew Expedition, at 
Pongo, Bolivia, alt. 12,000 ft., February 17 to March l, 
1926 (No. 198). 

"Gaultheria martaensis Rusby" proves, on re-examination, 
to be a species of Pernettya, perhaps a form of P. Pent- 
landii DC. 


VACCINIUM TATEI Rusby, spe nov. In partibus juvenilibus 
minute puberulum; rami dense ramosi, ramulis adscendentibus 
rubellis; laminae quam petiolis recurvis vel tortis 5-plo 
longiores ovales obtusae coriaceae subtus ferrugineae; ped- 
icelli crassi quam fructu dimidio breviores. 

Young portions minutely downy; stems densely branched, 
the branchlets ascending, slender, ferruginous or reddish, 
mostly 43 to 6 om. long, densely leafy; petioles about 1.5 
mn. long, mostly recurved or twisted, stout; blades 8 by 5 
mm., oval, obtuse, coriaceous, shallowly sinuate-dentate (or 
entire toward the base), pale-green above, ferruginous be- 
neath, the venation faintly impressed above, stout and prom 
inent beneath, the secondaries 4 to 5 on each side, crooked, 
uniting to form a line near the margin; fruits few at the 
summit of the branchlets, purple, the pedicels stout, crook 
ed, less than half the length of the fruit, which, in the 
dried state, is 3 or 4 mm in diameter, globose; calyx-teeth 
short and broad, acute. 

Collected by G. H. H. Tate at Pongo, Bolivia, alt. 12,000 
ft., February 17 to March 1, 1926 (No. 199). The species is 


72 FAY TT .0)4:0-63 2 No. 2 
apparently a low, much-branched shrub. 


RAPANEA DENTICULATA Rusby, sp- nov. Subglabra; ramli 
cicatricibus foliorum asperati; folia sessilia oblonga vel 
oblanceolata margine revoluta et prope basin denticulata, 
glabra rugulosa; sepala quam fructu dimidio brevioria tri- 
angulari-oveta adpressa. 

Subglabrous; branchlets short, stout, coarsely and irreg- 
ularly angled, roughened with sharply projecting leaf-scars; 
leaves sessile, but with narrower petiole-like bases, from 7 
by 3 cm. to 10 by 4 om, oblong or mostly slightly oblanceo- 
late, acute, the margin sharply revolute, minutely denticu- 
late, increasingly so toward and upon the petiole-like base, 
thick and rigid, sometimes slightly scurfy on the midrib a- 
bove, otherwise glabrous, but the extremely fine wrinkles 
resembling trichomes or papillae; midrib slightly impressed 
above, prominent beneath, the rest of the venation lightly 
prominent ebove, the secondaries about 15 on each side, 
slender, crooked; bracts triangular-subulate, thick and rig- 
id; flowers not seen; fruiting-calyx closely appressed, the 
sepals thick and rigid, keeled, nearly half the length of 
the fruit, triangular-ovate, acute; fruit sessile, about 5 
in a cluster, mostly 5 m. long and 4 mm. wide, oval, tipped 
with the strong style-base, brown, faintly nerved, finely 
wrinkled-papillose. 

Collected by G. H. H. Tate, on the Ladew Expedition, at 
Rio Aceromarco, Bolivia, alt. 10,800 ft., May 24 to 28, 1926 
(No. 706). The species is peculiar in its denticulate leaf- 
bases. It is near R. latifolia (R. & P.) Mez, but the fruit 
characters and the denticulation of the leaf are very dif- 
ferent. 


GENTIANA TATEI Rusby, sp. nov. Glabra; caules graciles 
erecti simplices foliosi; folia basalia oblanceolata basi 
angustata obtusa, caulina a basi amplectente sensim ad api- 
cem angustatas cymae sessiles pauciflora vel multiflora, 
pedicellis brevibus confertis anguste 4-alatis; calyx quam 
corolla dimidio brevior, lobis quam tubo duplo longioribus; 
corollae tubus quam lobis latis rotundatis duplo longior. 

Glabrous; stems slender, erect or ascending, simple, 15 
to 30 cm. long, leafy; root-leaves 2 cm. or more long, 3 or 
4 mm. wide, oblanceolate, obtuse, tapering gradually to the 
base, the stem-leaves sessile with clasping base, from 
which they taper gradually and regularly to the summit, 
which is either acute or obtuse, mostly about 2 cm. long, 
narrowly linear, very thin, finely 7- to 9-nerved, those at 
the base of the cymes being relatively broader; cymes simple 
or corymbose, several- to many-flowered, sessile, the pedi- 
cels & to 12 mm. long, slender, slightly thickened upward, 
4-angled, the angles bearing narrow membranous hyaline marg= 


1934 Rusby, New Species from Bolivia 73 


ins; flowers 14 to 17 mm. long, about 1 cme broad at the 
mouth; calyx a little more than helf the length of the cor- 
olla, divided two-thirds of the way, the tube broadly cam 
panmulate, 8-nerved, the sinuses broad and obtuse or round- 
ed, the lobes triangular and tapering regularly from the 
base to the acute summit, apparently pale-green; corolla 
campanulate, apparently white, pink, or possibly yellowish, 
lobed a third of the length, the lobes very broad and round- 
ed, entire, minutely apiculate, without folds in the sinus- 
es; stamens nearly equalling the corolla, attached at its 
base, distinct, the filaments stout, broad and flattened, 
attached broadly to the lower half of the blackish anther, 
which is 1 mm. long, ovate, and obtuse; pistil equalling the 
corolla, the ovary lanceolate, the style very short, the 
stigmas small, strongly recurved. 

Collected by G H. He Tate at Alaska Mine, Bolivia, alt. 
13,700 ft., March 1 to 4, 1926 (No. 65). 


DURANTA RECURVISTACHYS Rusby, sp. nov. Partes juveniles 
pubescentes; ramli divaricati aut recurvi, racemi vel pan- 
iculae terminales recurvi; folia ovalia vel ovata, abrupte 
acuteque angustata, in basin petioloideum abrupte contracta, 
serrata; pedicelli brevissimi reflexi vel torti; corollae 
tubus quam calyce duplo longior. 

Inflorescence, younger portions, etc., pubescent; 
branches stout, terete, the branchlets divaricate and some- 
what recurved, the suprea-—axillary spines very short, stout, 
straight, pungent; stem-leaves to 8 by 5 or 6 cm., oval or 
ovate, very abruptly, shortly, and acutely pointed at the 
summit and very abruptly contracted into a petiole-like base 
about 1 cm. long, coarsely serrate with short and broad me- 
ronate teeth, rather thick, deep-green and sparsely and ob- 
scurely pubescent above, where the slender venation is very 
lightly impressed, the lower surface yellowish-green, pubes— 
cent on the veins, which are yellowish, slender, and promin- 
ent, the secondaries about 5 on each side, falcate and some— 
what crooked; leaves of the branchlets mostly 5 by 2 cm, o- 
val, abruptly short-acuminate at both ends, acute, entire or 
with a few obscure teeth, otherwise like the stem-leaves, but 
more pubescent; inflorescence either simple, slender, loose- 
ly flowered, recurved racemes or panicles terminating the 
branchlets, their branches similarly racemose, nodose where 
the flowers or fruits have fallen, some of the flowers sub- 
tended by very small lance-linear acute bracts; pedicels 
very short, mostly reflexed or twisted; calyx about 6 mm. 
long, the teeth about 1 mm. long, erect, subulate; corolla- 
tube twice the length of the calyx, the exserted portion 
strongly recurved, the limb abrupt, 8 to 10 mm broad, the 
lobes reflexed; fruit 1 cm. long and nearly as broad, ovoid, 
mucronate, brown, somewhat shining. 


74 PHYTOLOGIA No. 2 


Collected by M. Bang in Bolivia, probably near Cochabamba 
(No. 1798) and distributed as D. Plumieri Jacq., but cert- 
ainly not that species. Buchtien's No. 314, collected in 
South Yungas, et an altitude of about 7000 ft., June l2, 
1906, "a shrub of 4 or 5 meters", is probably of this spe- 
cies. Specimens collected by G. H. H. Tate at Okara, alte 
7500 ft., April 26 to 29, 1926 (No. 910), apparently repre- 
sent a distinct and undescribed species, but flowers are 
wanting. The branch-leaves have slender petioles 1 to 1.5 
em. long, the young growth is ferruginous-tomentellous, the 
pedicels are slender, and the fruits are as broad or broad- 
er than long and more strongly and stoutly beaked. 


LANTANA TENUIFOLIA Rusby, sp. nov. Strigosa; aculei 
parvi graciles acuti recurvi; rami graciles adscendentes; 
folia petiolata ovata acuta, utrinque breviter acumineta, > 
pilis albidis adpressis ex basibus bulbosis viridibus ves- 
tita; pedunculi elongati graciles erecti; capitula lata den- 
siflora arcte involucrata; flores strigosi. 

Armed with small, slender, recurved, pungent prickles and 
more or less strigose throughout; stems elongated, irregu- 
larly angled or sulcate above, the branches elongated, 
strongly ascending; petioles to 2 cm. long, grooved above; 
blades (which turn black in drying) to 10 or 12 by 5 cm, 
ovate, very abruptly short-acuminate at both ends, ecute, 
jrregularly and finely crenate-dentate, the teeth short, 
obtuse or acutish, the sinuses narrow and acute, thin, deep- 
green, strigese with stiff, appressed, white hairs, each 
from a lerge bulbous green base, the venation lightly prom- 
inent beneath, the secondaries about 8 on each side, ascend- 
ing at about 45°, little curved; peduncles to 15 cm. long, 
erect or strongly ascending, very slender; heads to 2 or 3 
cm. broad, densely flowered, strongly involucrate by a 
circle of closely appressed reduced leaves; flowers stri- 
gose, about 1 cm. long, the tube contracted to about half 
its diameter near the limb. 

Collected by H. H. Rusby in the Bopi River valley, Boliv- 
ia, alt. 3000 ft., September 11, 1921 (Mulford No. 653, the 
type) and by G. H. H. Tate at Mapiri, alt. 2000 ft., March 
30 to April 9, 1926 (No. 469). The species is very near L. 
tiliaefolia Cham. 


HEDEOMA ADSCENDENS Rusby, sp. nov. Cinereo=puberula et 
asperula, pilis crassis albidis; caules graciles decumbent— 
es, ramis mimerosis erectis; folia ovata obtusa obtuse den- 
tata basi rotundata; pedicelli brevissimi; calyx longe pil- 
osusB. 

Gray-puberulent and roughish, with short, stout, white, 
divaricate hairs; stem slender, more or less decumbent, dif- 
fusely branched, the branches erect, 4 to 5 -m. long, the 


| 
Laat 
ey 

an 


1934 Rusby, New Species from Bolivia 75 


internodes mostly longer than the leaves; petioles 1 to 1.5 
mm. long; blades to 4 by 4 mm., ovate, with rounded base and 
blunt summit, thickish, shallowly and obtusely dentate, the 
venation coarse and stout, the secondaries about 43 on each 
side; flowers mostly solitary in the axils of the upper 
leaves, some of these reduced to bracts; inflorescence 
white-pilose with rigid hairs; pedicels about 1 mm. long; 
calyx-tube 2.5 mm. long, infundibular, the lower half tur- 
gid, strongly ribbed, the limb divided nearly to the base, 
the teeth long—vilose, subulate and very acute, subequal, 
the lower somewhat longer; flowers not seen. 

Collected by G He He Tate on the Tipuani-Ancona-Sorata 
trail, alt. 13,000 ft., April 30, 1926 (No. 783). 


SOLANUM TENUISPINUM Rusby, spe nov. Aculatus setosum et 
glanduloso-pilosum; aculei flavescentes elongati graciles 
acutissimi infra medium glanduloso=pubescentes; setae albi- 
dae graciles; petioli elongati; laminae late ovatae obtusae 
vel subacutae, dentibus megnis vel lobis utroque letere 2 
vel 3, basi truncata vel subcordatas calyx setosus. 

Branches, petioles, peduncles, and pedicels spiny, 
bristly and glandular-hairy; spines yellowish or light- 
brown, elongated, slender and very pungent, mostly divari- 
cate, mostly a littel compressed laterally at the base, and 
bearing, especially below the middle, short—glandular hairs, 
which are also borne on the stem and elsewhere; bristles 
whitish, slender; branches slender, very flemious, green, 
irregularly angled; petioles to 8 cm. long, widely spread- 
ing, green-sulcate or angular; blades to 14 cm. long and 
nearly as wide, ovate, the blade somewhat oblique and trun- 
cate or subcordate, obtuse or acutish, with 2 or 3 large 
triangular teeth or lobes on each side, which are obtuse or 
acutish, the sinuses obtuse or rounded, the margin ciliate, 
very thin, deep-green above, bright-green beneath, the 
veins on both surfaces sparingly spiny, both surfaces more 
or less bristly and minutely and sparsely stel late-hairy 
among the bristles, the venation slender, prominent beneath, 
the secondaries 5 or 6 on each side, the finer venation 
coarsely reticulate; cymes umbelliform, few-fruited, the pe- 
duncle shorter than the pedicels, opposite and below the 
petiole, all the stems long-hispid and glandular, the pedi- 
cels regularly thickened upward; calyx bristly, parted near- 
ly to the base, the lobes about 5 mm. long, narrowly lanceo- 
late, acuminate and acute, subcarinate; fruit spherical, 
glabrous, light-brown, 1 cm. in diameter. 

Collected by G. H. H. Tate, on the ladew Expedition, at 
Nequejamira, Bolivia, alt. 8000 ft., May 15 to 24, 1926 
(No. 662). The species is near S. atropurpureum Schrenk. 


BARTSIA ALTISSIMA Rusby, sp. nov. Albido-hispida; caulis 


76 PHYTOLOGIA No. 2 


altus crassus sed debilis rubellus, ramis numerosis brevi- 
bus; folie amplectentia divaricata oblongo-lanceolata obtusa 
crassa crenato-dentata, ad basin latissima; flores in axil- 
lis superioribus sessiles; celycis tubus late campanulatus 
lobis trianguleri-ovetis obtusis. 

Mispid with short, stout, white, glistening hairs, many 
of which are branched at the base, or 2 or 5 from the same 
base; stem (of my specimen) nearly 7 dm. long, stout, but 
weak, reddish, strongly sulcate and obtusely angular, the 
branches numerous, short; leaves sessile and clasping, di- 
varicate or somewhat deflexed, to 2 cm. long, broadest at 
the base, which is sometimes nearly half the length, lance 
oblong, obtuse, gray-green, densely hispid, thickish, cre- 
nate-dentate with short obtuse teeth and acute sinuses, the 
midrib and secondaries very strong and keel-like on the low- 
er surface, the secondaries 4 or 5 on each side, strongly 
ascending; flowers few, solitary and sessile in the upper 
axils, unexpanded in my specimen, the calyx broadly campan~ 
ulate, the tube 4 mm. long and broad, the lobes triangular- 
ovate, obtuse, the longer about 2 m. long; dissection 
material wanting. . 

Collected by G. H. He Tate, on the ladew Expedition, at 
Pongo, Bolivia, alt. 12,000 ft., February 17 to March 1, 
1926 (No. 222). The species is very near B, mtica Benth. 
and B. hispida Benth., but is mich more hispid than either 
and the hairs are different. 


BARTSIA GUGGENHEIMIANA Rusby, spe nove Cinereo-hispid- 
ula; caules caespitosi ex caudice crasso foliosissimi; 
folia sessilia valde revo lta oblonga obtusa leviter cre- 
nata, basi angustata; pedicelli brevissimi graciles; caly- 
cis tubus cupulatus, dentibus rotundatis carnoso-incrassa= 
tis; corolla recta pubescens, tubo gracili. 

Minutely grayish-hispidulose; stems caespitose, branch= 
ing from a short and stout caudex, the branches 3 or 4 ecm. 
long, stout, ascending, very densely leafy; leaves ses- 
sile, 5 or 6 m. long, about 2 mm. wide when flattened out, 
but revolute and appearing narrower, oblong, obtuse with a 
narrowed base, the margin revolute, shallowly crenate, deep 
green; pedicels very short, slender; calyx-tube cupulate, i 
mm. long, 2.5 mm. wide, the 2 longer lobes 1 mm. long and 
broad, the 2 smller ones about two-thirds as large, all 
rounded, of a deeper green than the tube, fleshy-thickened; 
corolla straight, pink (7), pubescent, the tube slender, 4 
mm. long, the limb slightly shorter, its longer lip more 
then twice the length of the shorter and several times 
lerger; stamens at length slightly exserted, the summit re- 
curved, the anthers broadly oval, the thecae micronate at 
the base; style filiform, about twice the length of the 
calyx, sigmoid-curved, the stigma oval, obtuse, somewhat ob- 


1934 Rusby, New Species from Bolivia 77 


lique; capsule (immature) broadly ovoid, with a rounded sum 
mite 

Collected on the Ladew Expedition by G. H. H. Tate at Al- 
aska Mine, Pongo, Bolivia, alt. 13,500 ft., March 1 to 4, 
1926 (No. 64). 


CALCEOLARIA CRYPTANTHA Rusby, sp. nove Ubique tomentosa; 
rami graciles recurvi; petioli breves graciles; laminae ova- 
tae acutae basi truncatae crenato-dentatae dentibus micronu- 
latis, supra puberulae, subtus cinereo-tomentellae; calyx 
latus sepalibus fere distinctis et aequalibus subtrinervis; 
corolla tomentella labio parviore poculiforme. 

Habit unknown, apparently herbaceous and erect, with 
slender recurved branches; shortly tomentose throughout; 
branches strongly, irregularly, and obtusely angled; petiole 
to 5 mm. long, slender; blades to 3 by 2 cm., ovate with a 
truneate or lightly cordate base, acute, crenate-dentate 
with acute or mucronulate teeth, thin, deep-green and puber- 
ulent above, grsey-tomentellous beneath, with the slender 
venation prominent, lightly falcate-ascending; inflorescen— 
ce somewhat crowded at the ends of the short branchlets, the 
pedicels slender, at length 1.5 to 2 cm. long; calyx nearly 
2 cm. broad, the sepals nearly distinct, broadly oval, muc-— 
ronate, veiny, sub-trinerved; corolla yellow, tomentellous, 
the larger lip 13 mm. long and broad, nearly circuler (as 
pressed), the claw very short, the smaller lip 3 mm. long 
and 4 mm broad, cup-shaped, the stamens partially conceal- 
ed in it; anthers 4.5 mm. broad, the thecae short and broad, 
divaricate from a broad base, confluent; style stout, equal- 
ling the anthers. 

Collected on the Ladew Expedition by G. H. H. Tate at the 
top of the pass Tipuani to Sorata, alt. 14,000 ft., April 


20 5 1926 (No. 819). 


CALCEOLARIA LECHIOIDES Rusby, spe nov. Ubique cinereo- 
tomentello; caules lignosi rubelli ramosissimi, ramis ad- 
scendentibus foliosis; petioli breves crassis; laminae cras= 
siusculae ovatae obtusae crenato-dentatae basi rotundatae; 
pedicelli brevissimi; calyx 1 om. latus lobis late ovatis 
obtusis quam tubo duplo longioribus. 

Gray-tomentellous throughout; stems woody, erect or as— 
cending, reddish, much-branched, the branches short, ascend- 
ing, very leafy; petioles nearly 1 mm. long, rather stout, 
the blades to 6 or 7 m. long, 3 mm. wide, ovate, with a 
rounded base and obtusish summit, strongly crenate-dentate, 
with 4 or 5 teeth on each side, thickish, the venation im 
pressed on the upper surface, stout and prominent under- 
neath, the upper surface gray, the lower somewhat ferrugin- 
ous; flowers racemose at the summit, the pedicels very 
short; calyx 1 cm. broad, divided two-thirds to the base, 


78 PE YT 0 L.0-G 27k No. 2 


the lobes broadly ovate, obtusish, thick, one distinctly 
broader than the others; corolla tomentose, yellow, 12 mm. 
long, 1 em. broad, unopened in my flowers dissection mate 
erial wanting, 

Collected on the Ladew Expedition by G. H. H. Tate at 
Pongo, Bolivia, alt. 12,000 ft., February 17 to March 1, 
1926 (No. 220). 


CALCEOLARIA RIBESIAEFOLIA Rusby, spe nov. Albido=pubes— 
cens; rami angulati et sulcati valde adscendentes; petioli 
suberectis laminae triangulari-ovatae acutae paullo triloba- 
tae grosse dentatae, basi subcordatae; pedicelli filiformes; 
sepala fere aequalia fere 1 cm. longa, ovata, acuta, quam 
corolla dimidio brevior. 

Pubescent, with short white divergent hairs, the upper 
leaf-surfaces strigilloses stems and branches erect or 
strongly ascending, strongly angled and sulcate, my speci- 
men 8 dm. high; leaves opposite, the petioles nearly erect, 
slender, to 5 cm. long, slightly dilated at the base, chan= 
neled above; blades 6 or 7 cm. long and broad, triangular- 
ovate, with a broadly truncate or subcordate base and an 
acute summit, sub-3-lobed, coarsely and unequally dentate 
with triangular acute teeth and acute or obtuse sinuses, 
very thin, the upper surface deep-green and rather sparsely 
short-strigose, the hairs stout, white, and glistening, the 
lower surface pale-green, grayish=pubescent, 5S- to 7=nerved 
from the base or near it, the venation weak and not con- 
spicuous; flowers few, loosely corymbose at the summit, the 
pedicels filiform; sepals sub-equal, the largest nearly 1 
em. long, broadly ovate, acute, densely pubescent; large 
lip of corolla 2 cm. long, the claw 3 mm., the sac 1 cm. 
broad; smaller lip of corolla 5 mm. long and broads; anther 
cells oval, turgid, 2 mm. long, 1.5 m. wide, contiguous, 
nearly horizontal, confluent. 

Collected on the ladew Expedition by G. H. H. Tate at 
Pongo, Bolivia, alt. 12,000 ft., February 17 to March 1, 


1926 (No. 219). 


ARCYTHOPHYLLUM NODOSUM Rusby, sp. nov. Subglabrum; 
caules crasei; rami nmumerosi erecti graciles, nodis incras= 
Satis et e foliis delapsis alte foveolatis; stipulee scaric- 
sae adpressae trianguleres apice 2—%setosae; folia demm 
recurva acuta breviaristata ex basi rotundata lanceolata. 

Nearly glabrous; stems stout, erect, mich-branched, the 
branches erect, slender, swollen at the nodes, which are 
deeply pitted where the leaves have disarticulated, the 
branchlets light-brown, sharply angled and clothed with the 
stipules; stipules scarious, strongly appressed, distinct or 
nearly so, triangular, 2 to 2.5 mm. long and mostly a little 
broader, bearing 2 or 3 slender terminal setae about of 


1934 Rusby, New Species from Bolivia 79 


their own lengths; leaves sessile, densely crowded, spreading 
and at length recurved, deep-green and shining above, appar- 
ently evergreen, at length turning brownish before falling, 
to 9 by 4 mm, lanceolate with a rounded base and an acute 
short-aristate summit, the middle portion much thickened, 
the margin somewhat thickened and deep-green against the ye= 
llowish lower surface; flowers few in the upper axils, sub- 
sessile, the calyx 5 mm long, parted nearly to the base, 
the calyx-lobes closely resembling the leaves; corolla-tube 
7 or & mm. long, 1.5 mm. wide, the lobes 3 mm. long, broad- 
ly ovate, acute, the corolla somewhat coriaceous; stamens 
attached near the summit of the tube, the filament slender, 
the anther 1 mm. long, oblong, slightly exserted; ovary very 
small, the style very short. 

Collected by G. H. H. Tate at Cocopunco, Bolivia, alt. 
10,000 ft., March 24 to 29, 1926 (No. 367). The species is 
very near to A. flavescens, but is well characterized by its 
swollen pitted nodes and its stipules, as well as by the 
crowded foliage and the leaf-characters. 


EVEA RADIATA Rusby, sp. nov. Glabra; rami graciles ni- 
grescentes, nodis incrassatis persistentes imbricatas trun- 
catas gerentibus; petioli graciles; laminae oblmgae vel ob-= 
lanceolatas, basi subacutae, apice breviter acuminatae et a- 
cutae; cyma terminalis, pedunculo gracili nigrescente, ramu- 
li cymae bracteis 4 parvis ovatis acutis subtenti. 

Glabrouss; branchlets slender, blackish, the internodes 
short, the nodes swollen and bearing the persistent stip- 
ules, which are interpetiolar, distinct but mostly imbricate 
at the base, 4 mm. long and broader, thick, truncate and 
sometimes bearing one or more short awns; petioles about 1 
cme long, slender, somewhat dilated at the base; blades to 
15 by 6 cm, oblong or oblanceolate with an acutish base and 
an abruptly short-acuminate or micronate acute summit, rath 
er thin but coriaceous, the slender venation lightly promin-e 
ent beneath, the midrib narrowly margined on the lower side, 
the secondaries very mumerous, the alternate ones smaller, 
divaricate and the outer portions falcate to meet in a 
strictly marginal line, loosely connected by the tertiaries; 
cyme terminal, long and slenderly peduncled, the peduncle 
blackish, angled, the branches subtended by four small o- 
vate acute bracts, the branches (4 in my specimen) about a 
third as long as the peduncle, slender, spreading, each ter— 
minating in a small head of 2 to several flowers, bracted 
like the cyme; fruits sessile or subsessile, about 1 ome 
long and nearly as broad, ovoid, blackish, the persistent 
calyx-limb light-colored, 2 or 3 mm. long and about as 
broad, truncate or obscurely toothed, the summit a little 
broader; flowers not seen, 

Collected by G. H. H. Tate at Ticunhuaya, Bolivia, alt. 


80 PEY T 0-L-0 @ ie No. 2 


5000 ft., April 20 to 24, 1926 (No. 1103). The foliage is 
closely similar to that of E. pectinata, but is thinner, and 
the species is well distinguished by its long peduncle and 
branches of the cyme. 


GALIUM CHAROIDES Rusby, sp. nov. Ubique scabrum pilis 
conicis, verisimiliter annuum; caules graciles adscendentes 
ex basi ramosi 4~anguleti vel anguste alati foliosi; folia 8 
in quoque verticillo sessilia inaequealia linearia acute ale 
bido-mucrenata, margine dense scabra; flores parvi axillares 
solitarii; fructus hispidus. 

Scabrous throughout, with short, conical, divaricate or 
ascending trichomes; apparently anmel, the stems slender, 
to 30 cm. tall, much branched at the base, ascending, 4— 
angled or very narrowly winged, very leafy; verticils mostly 
8-leaved, the leaves sessile, very unequal, the longest to 
15 mm. long, spreading or often recurved, linear and the up- 
pers ones somewhat broader, acutely mucronate with a white 
point, obscurely 3=-nerved, the margins densely scabrous with 
the white trichomes above described; flowers few, very small 
and solitary in the axils, the fruiting pedicels at length 
to 1 cm. longs; fruit 1 mm. long and nearly 2 mm. broad, em- 
arginate at the base and summit, the carpels nearly rotund, 
hispid, the hairs more slender than the others on the plant. 

Collected by G. H. H. Tate, on the ledew Expedition (No. 
234), at Pongo, Bolivia, alt. 12,000 ft. The species is near 


G,. pseudoaparine Griseb. 


(a) A limited number of reprints of this article are a— 
vailable, and will be sent postpaid on receipt of fifty 
cents each. To those ordering copies of my "Three Hundred 
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contribution will be sent without charge. 


TAXONOMIC NOTES ON AMERICAN PHANEROGAMS — I 


Lyman B. Smith 


This series is begun in order to record such notes as are 
necessitated by routine determinative work. Unless otherwise 
stated all material cited is deposited in the Gray Herbarium 
of Harvard University. 

Although many authors have regarded Pleurostachys as a 
synonym of Rynchospora, very few of the species have ever 
been transferred. The following species appear to be valid 
and to merit transfer to Rynchospora: 

RYNCHOSPORA FIUMINENSIS L. B. Smith, nom. nov. Nemochloa 
Beyrichii Nees in Mart. Fl. Bras. ii. pt. 1, 152 (1842), non 

chospora Beyrichii Steud. Syn. Pl. Glumac, ii. 144 

1855). Pleurostachys Beyrichii Steud. Syn. Pl. Glumac. ii. 
12 es P. panicoides Pfeiff. in Bot. Archiv. ix. 240 
1925). ‘ 

RYNCHOSPORA FOLIOSA (Kunth) L. B. Smith, comb. nov. 
Pleurostachys foliosa Kunth, Enum. Pl. ii. 284 (1837). 

RYNCHOSPORA GAUDICHAUDII (Brongn.) L. B. Smith, comb. 
nov. Pleurostachys Gaudichaudii Brongn. Voy. Coquille, 174 
(1834). Nemochloa Gaudichaudii Nees in Mart. Fl. Bras. ii. 
Dts 1,150 (1842). 

RYNCHOSPORA GRAMINIFOLIA (Brongn.) Le Be Smith, comb. 
armen s graminifolia Brongn. Voy. Coquille, 175 

18 ¢ 

RYNCHOSPORA MARTIANA (Nees) L. B. Smith, comb. nov. 
Nemochloa Martiana Nees in Mart. Fl. Bras. ii. pt. 1, 150 
(1842). Pleurostachys Martiana Steud. Syn. Pl. Glumac. ii. 
139 (1855). 

RYNCHCSPORA MILLEGRANA (Nees) Schrad. ex Griseb. Fl. 
Brit. W. Ind. 575 (1864). Nemochloa millegrana Nees in 
Mart. Fl. Bras. i1. pt. 1, 149 (1842). Pleurostachys mi 1- 
legrana Steud. Syn. Pl. Glumac. ii. 199 (1855). 

RYNCHOSPORA ORBIGNIANA (Brongn.) L. B. Smith, comb. nov. 
Pleurostachys Orbigniana Brongn. Voy. Coquille, 175 (1834). 
P. mecrantha Kunth, Enum Pl. ii. oath eal Nomochloa Or- 
bigniana Nees in Linnaea, ix. 299 (1834+). Nemochloa macran- 
tha Nees in Mart. Fl. Bras. ii. pt. 1, 151 (1842), 

RYNCHOSPORA PANICOIDES Schrad. ex Nees in Mart. Fl. 
Bras. ii. pt. 1, 154 (1842), in synonymy. Pleurostachys 
stricta Kunth, Enum. Pl. ii. 286 (1837), non Aumehospors 
atricta Boeckl. in Linnaea, xxxvii. 603 (1873). Nomochloa 
stricta Nees in Hooker's Journ. Bot. ii. 398 (1840). The 
name panicoides has not been used for any other species of 


ee 


81 


82 PRYTto0L Oe Fs No. 2 


Rynchospora, so it may be revived for this one although pub- 
lished in synonyny. 

RYNCHOSPORA PUBERULA (Boeckl.) L. B. Smith, comb. nov. 
Pleurostachys puberula Boeckl. in Flora, 1lxiii. 453 (1880). 

RYNCHOSPORA SCALARIS L. B. Smith, nom. nov. Pleurostachys 
Urvillii Brongn. Voy. Coquille, 173, t. 31 (1834), non R 
chospora Urvillei Steud. Syn. Pl. Glumac. ii. 149 (1855). 
Nemochloa Urvillei Nees in Mart. Fl. Bras. ii. pt. 1, 1H 

1842). 

RYNCHOSPORA SPARSIFLORA (Kunth) L. B. Smith, comb. nov. 
Pleurostachys sparsiflora Kunth, Enum. Pl. ii. 286 (1837). 
Nemochloa sparsiflora Nees in Mart. Fl. Bras. ii. pt. 1, 151 

1842). 

RYNCHOSPORA TENUIFLORA (Brongn.) L. B. Smith, comb. nov. 
Pleurostachys tenuiflora Brongn. Voy. Coquille, 175 (1834). 
Nomochloa tenuiflora Nees in Linnaea, ix. 295 (1834). N. 
turbinata Nees in Meyen, Reise, i. 112 (1843), in synonymy. 
Pleurostachys elegans Kunth, Enum. Pl. ii. 285 (1857). Nemo= 
chloa elegans Nees in Mart. Fl. Bras. ii. pt. 1, 149 (1642). 
Pleurostachys Luetzelburgiana Pfeiff. in Fedde, Rep. Spec. 
Nove xix. 20H (1924). Pe latifolia Pfeiff. ibid. 295. 

Nomochloa turbinseta was published in synonymy to begin 
with, and later authors in separating it have given no 
really distinctive characters to justify such action. The 
inflorescence is mich denser than in mature Rynchospora ten- 
uiflora, but that is only because the type consists of very 
young material. 


Pleurostachys luetzelburgiena and P. latifolia were 
based on minor distinctions which break down on comparison 


with ample material of chospora tenuiflora. 

APHELANDRA PRI SMATICA (Tell.) Hiern in Kjoeb. Vidensk. 
Meddel. 78 (1877). Ruellia prismatica Vell. Fl. Flum. 267 
(1825), Icones, vi. t. 98 (1827). R. quadranguleris Vell. 
ibid. 267, t. 97. Ruellia quadrangularis has been consid- 
ered a synonym of Aphelandra sciophila Mart., yet the lat- 
ter has distinctly dentate floral bracts while Vellozo's 
plate of the former shows the bracts entire. R. quadrangu- 
laris and R. prismatica are simply young and mature stages 
of the same species. In a single collection from Alto da 
Serra in the state of Sao Paulo, Brazil (L. B. Smith 
1934), I have specimens matching each plete perfectly. 

RUELLIA SOLITARIA Vell. Fl. Flum. 266 (1825), Icones, 
vi. t. 95 (1827). Dipteracanthms Schauerianus Nees in 
Linnaea, xvi. 290 (1842). Ruellia Schaueriana Lindau in 
Engl. & Prantl, Net. Pflanzenf. iv. Abt. 3b, %10 (1895). 
That there can be little doubt of the identity of Vellozds 
species, was found in determining A. C. Brade 11015 and 
L. B. Smith 1365, both from the city of Rio de Janeiro. 
Consequently there is no point in maintaining Lindeu's 
much later name for the species. 


r 
- 


A CASE OF "POLLINIA" (a) 


Charles Clinton Smith 


Recently while studying the pollen of Sarcolaena mlti- 
flora Dup. Ehs., a member of the Chlaenaceae, the author no- 
ticed that the grains were large and were deeply sculptured 
in a most curious manner. Between the sculpturings could be 
seen the outlines of a number of spherical objects. 

Upon being shown a specimen, Dr. Paul B. Sears suggested 
that it might be a pollinium of sixteen grains, all ene losed 
within the thick and sculptured wall. Further study proved 
this to be the case. . 

A search into the literature for information or refer- 
ences concerning the pollen of this family revealed the no- 
tation from Ingler and Prantl that the pollen grains are 
comparatively large, that their form is, in all species ex- 
amined, uniform and spherical, and that they have six deep 
furrows crossing their surface. Tetrads are arranged accord- 
ing to the angles of furrows (b). 

The only cases of pollinia recorded ‘in the literature 
were those of the Orchidaceae, Asclepiadaceae, Mimosae, and 
Acaciae. In these cases of pollinia there are usually 8, 16, 
64, or hundreds of pollen grains which are united by a waxy 
substance. There is no mention of a special covering around 
the mass or any consistent numbers of cells concerned. In 
some cases the contents of the whole locule of the anther 
has been united. 

A condition similar to the one found by the writer is the 
so-called "tetrad", a condition in which 4 cells are within 
& common wall or are united with various degrees of firm 
ness. These have been reported by various workers in the 
following plant groups: many species of Orchidaceae, in the 
Ericaceae (Erica, Calluna, Menziesia, and Andromeda), in the 
Epacridacese (Epacris and Leucopogon). Other genera are 
Arctostaphylos, Arbutus, Rhododendron, Ledum, Vaccinium, 
Juncus, Jacquinia, lumla, Anona, Drimys, Jussiaea, Perip lo- 
ca, Fourcroya, and Typha. 

Upon request, Dr. H. Humbert of Paris sent eight addit- 
4onal species of the Chlaenaceae, including six genera. An 
examination of this material revealed that a similar con- 
dition existed in all of these species. In four species the 
pollen grains are in quartets (see figs. 1, 2, 5, and 8), 
the degree of union of the members of the quartet varying 
from a condition where a heavy sculptured wall encloses the 
four cells to a condition where the four cells are only 


83 


Pea EO tO ees No. 2 


$ 


TABLE I. 


Data concerning the pollen of the members of the Chlaenaceae 
examined in this study. 


Pollen of: Diameter Number of Description 
in m cells 
Eremolaena 90 4 Common wall thick, not 
rotundifolia sculptured in regular 
fashion. 
Eremoleena 117 4 Common wall with at 
Humb lotiana least 4 wide deep fur- 
rows, wall papillate 
along margin. 
leptolaena 73 16 Tetrad separated by 
multiflora heavy partition, com 
mon wall sculptured. 
Rhodo laena 109.5 16 Tetrads very distinct, 
Bakeriana common wall very thin, 
not sculptured. Cells 
bulge walls out. 
Rhodo laena 45 4 No outer wall, members 
parviflora of tetrads separated 


Sarco laena 
codonochlamys 127.4 


Sarcolaena 101 
multiflora 


Schi zo laena 40 
cauliflora 


Xerochlanys 80 
Bojeriana 


Xy lolaena 1% 
Richardii 


easily. 


Outer wall heavy. 


like above, tetrads 
very distinct. 


Rather easily separa= 
ted into single grains 


Common wall rather 
thick. Crystals in 
wal ls. 


Members of tetrads 
rether firmly united, 
may be surrounded by a 
common wall. 


1934 Smith, A Case of "Pollinia" 85 


slightly coherent. 

The remaining species have compound grains composed of 16 
cells. The outer wall of these compound structures is quad- 
risected by six deep furrows (fig. lla). Each of the four 
portions of the wall ths formed covers a quartet of cells. 

Such a condition could have resulted if the members of 
the tetrads formed during sporogenesis had undergone two 
more divisions and all the cells thus formed had remained 
within the spore-mother cell wall. The descendants of mem 
bers of the original tetrad would form quartets enclosed 
within the wall which probably formed about each member of 
the original tetrad before the next divisions occurred. Thuis 
the sixteen cells are grouped into four quartets, each quar- 
tet distinct from the other three quartets. 

The members of each quartet are arranged so that three 
cells are against the outer wall and one is in the center 
(fig. llc). The three cells against the outer wall thus form 
in general outline a triangle. The deep furrows mentioned 
above pass between adjacent quartets (fig. lla) and the 
areas between the furrows are also triangular. Fach triang- 
ular portion of the outer wall fits over a quartet like a 
cap (fig. 1lb). 

The compound grain might be said to be quadriseptate, 
since the wall of the cells which divided to form the mem 
bers of each quartet is discrete and definitely separates 
the quartets from one another. 

The outer wall which covers the four quartets probably 
represents the modified spore-mother cell wall, 

The width of the furrows varies with the water content of 
the structure. When the cells are turgid, the furrows are 
wide, when dried, narrow. 

The word pollinia undoubtedly was not coined to designate 
as complex or highly organized structures as have just been 
described. Kemer and Oliver, in their Natural History of 
Plants, give the following definition of a pollinium, "The 
resultant mass of pollen cells formed when the pollen of one 
Archesporium remain united into a tissue". Webster's New In- 
ternational Dictionary definites it simply as, "A coherent 
mass of pollen grains", 

A new term to more aptly describe these structures seems 
desirable. The term, pollen tetraquartet, although a little 
awkward, describes the true condition well, as will be seen 
from the description above (c). 

Table I gives the data concerning the members of the fam 
ily examined in this study. 

The ancestral type of this family apparently arose on the 
island of Madagascar and for some reason remained in a small 
area on the island. Tracing the evolution of the present 
types on the basis of pollen morphology may be done, al- 
though some steps in the process may be missing. If we ar- 


86 PHYTOLOGIA Noe 2 


range the species according to complexity we arrive at this 

series: 

Rhodoleena parviflora, simple quartet, easily separated. 

Schizolaena cauliflora, simple quertet, rather easily separ- 
ated. 

Eremolaena rotundifolia, single quartet, surrounded by a 

common unsculptured wall. 

Eremolaena Humblotiana, single quartet, surrounded by a 
thick sculptured wall. 

Xylolaena Richardii, four quartets, common wall unmodified. 
Leptolaena multiflora, Rhodolaena Bakeriana, Sarcolaena 
codonochlamys, Sarcolaena multiflora, and Xerochlamys Bojer- 
jana would then culminate the series as the most complex on 

the basis of pollen morphology. 


: 
i 
4 
: 
% 


Fig. 8 Fig. 9 


Fig. 10 
f Fig. 11 


if 
q 
Bie 


88 PRY TO b.0- Gis No. 2 


Explanation of Figures 


Figures 1 to 10 inclusive, photomicrographs by the auth- 
or, made with Spencer Microscope Model 44. Figure 11 photo- 
graphed by Theodore Kibby. 

Fig. 1. Pollen quartet of Eremolaena rotundifolia P. Danduy, 


X 180. 
Fig. 2. Pollen quartet of Eremolaena Humblotiana H. Bnse, 
X 115. 


Fig. 4. Pollen tetraquartet of Leptolaena multiflora Du 
Petit Thouars, X 160. 
Fig. 4. Pollen tetraquartet of Rhodolaena Bakeriana H. Bn., 


xX 145. 
Fig. 5. Pollen quartet of Rhodolaena parviflora F. Gerard, 
X 215. 


Fig. 6. Pollen tetraquartet of Sarcoleena codonochlamys 
Baker, X 140. 

Fig. 7. Pollen tetraquartet of Sarcolaena miltiflora Dup. 
Ehs., X 185. 

Fig. 8. Pollen quartet of Schizolaena cauliflora Dup. Ehs., 


X 290. 

Fig. 9. Pollen tetraquartet of Xerochlemys Bojeriana H. Bn., 
X 180. 

Fig. 10. Pollen tetraquartet of Xylolaena Richardii H. Bn., 
X 105. 


Fig. 11. Clay model of pollen tetraquartet of Rhodolaena 
Bakeriana H. Bn.j a. one cell; b. rim of cap; 
c. bulge in cap due to cell beneath. 

Fig. 12. Pollen tetraquartet of Sarcolaena miltiflora Dup. 
Ehe., X 780. 


(a) Contribution from the Botanical Laboratory of the 
University of Oklahoma, No. 29. 

(b) Die naturlichen Pflanzenfamilien, III Teil, 6 Ab- 
teilung. 

(c) Sharp (Lester W., An introduction to cytology. New 
York. Ed. 3. 1934. Page 250), it seems to us wisely, has ad- 
vocated the use of the term quartet to designate four cells 
formed by division of a mother cell and limited the term 
tetrad to the univalent groups of chromosomes present pre- 
vious to the formation of separating cells. 


THE MAHONIAS OF THE PACIFIC STATES 


LeRoy Abrams 


The late Professor Piper has given a scholarly account 
(1) of the taxonomic history of Mahonia aquifolium and M. 
repens, in which he reports the results of critical studies 
of the peculiar differences in the epidermis of the lower 
surfaces of the leaflets. He found that these epidermal 
characters set off M. aquifolium and M. pinata from the 


rest of the species. In these two species the epidermal 


cells of the lower surface of the leaflets are rounded on 
the back, the enlarged rounded portion conforming in outline 
with the cell and not evident as distinct papillae. Whereas 
in M. repens, M. pumile, and related species the lower sur- 
face is beset with minute cylindrical papillae. Accompanying 
this papillete character is the grayish color of the lower 
surface. 

By using this microscopical structure of the epidermis as 
an additional taxonomic character it is possible to delimit 
the species of Mahonia more clearly, and the present paper 
is an attempt to revaluate and realign the species of the 
Pacific States, which is one of the chief geographic centers 
of the gems. 


Bud-scales glumaceous, persistent, 15-40 mm. long; filaments 
unappendaged. 1, Me. nervosa. 
Bud-scales deciduous, 2-5 mm. long; filaments with a pair of 

recurved teeth near the apex. 

Racemes densely many-flowered; floral bracts triangular- 
ovate, obtuse or acute; berry oblong-ovoid, blue- 
black, fleshy. 

Epidermis of the lower surface of the leaflets not pap- 
illate; upper surface bright green and shining. 
leaflets ovate, mostly obtuse, the lowest pair borne 


close to the base. 2. M. pinnata. 
Leaflets ovate-lanceolate, the lowest pair remote from 
the base. 3. M. aquifolium. 


Epidermis of the lower surface of the leaflets with 
minute papillae. 
leaflets brisht-green and shining above. 
Teeth 7-11 on each margin of the leaflets, tipped 
with slender spines; lower surface of leaflets 


gray-green, 4, M. piperiana. 
Teeth 12-16 on each margin, merely bristle-tipped; 
lower surface not gray. 5. M. sonnei. 
89 


90 | PHYTOLOGIA No. 2 


Leaflets dull or gray-green above, grayish beneath. 
Teeth small, bristle-tipped, usually 12 or more ona 
side. 6. M. repens. 
Teeth large, tipped with stout spines, usually 5-9 
on each margin. 
Plants low, 2-5 dm. high; teeth not as long as the 
body portion of the leaflet. 7. M. pumila. 
Plants 1-2 m. high; leaves pale gray-green on both 
surfaces, strongly undulate and rigid, the 
teeth about as long as the body portion of the 
leaflet. 8. M. dictyota. 
Racemes loosely 5-7-flowered; floral bracts lanceolate- 
acuminate; leaflets glaucescent on both surfaces; ber- 
ries globose, becoming dry. 
Leaflets with 2-4 prominent spiny teeth on each margin. 
9. M. fremontii. 
Leaflets narrow, long-acuminate, with 8 or more bristle 
like teeth on each margin. 10. M. nevinii. 


1. MAHONIA NERVOSA (Pursh) Nutt. Gen. Pl. 1: 212. 1818. 


Berberis nervosa Pursh, Fl. Sept. 219, pl. 5. 1814. 
Mahonia glumosa DC. Syst. Veg. 2: 21. 1821. 
Odostemon nervosus Rydb., Bull. Torrey Club 33: 141. 1906. 
Stems simple, 2-6 dm. high, bearing the leaves in a term 
inal tuft, the scales of the terminal bud glumaceous. per- 
sistent, 2-4 em. long. Leaves 25-40 cm. long; leaflets 7-21, 
ovate to ovate-lanceolate, acute, 3-5 cm. long, spinulose- 
serrate, glossy-green, somewhat palmately nerved; racemes 
7-20 em. long, erect; pedicels 5-8 mm. long; berries glau- 
cous-blue, 8-10 mm. in diameter. 
Coniferous forests, Humid Transition Zone; Vancouver 
Island and British Columbia to Monterey, Celifornie. Type 
locality: The Cascades of the Columbia River. 


2. MAHONIA PINNATA (Lag.) Fedde in Engler, Jahrb. 31: 86. 
1901. , 


Berberis pinnata Lag. Elench. Hort. Madr. 6. 1803 (nomen 
nudum); l.c. 14. 1816. 

Mahonia fascicularis DC. Syst. Veg. 2: 19. 1821. 

Odostemon fascicularis Abrams in Bull. N. Y. Bot. Gard. 6: 

- 1910. ‘ 

Stems erect, branching, 3-15 dm. high; leaflets 5-17, 

commonly 7-9, crowded on the rachis, the lowest pair near 

the base, glossy green above, scarcely paler beneath, rather 

thin, spinulose-dentate, the teeth usually 12 or more on 

each margin; racemes dense, fascicled, %-5 cm. long. 
Wooded slopes or thickets, mainly Upper Sonoran Zone; 


aay 


1934 Abrams, Mahonias of the Pacific States 91 


California Coast Ranges from Humboldt to Los Angeles Count- 


ies. Type locality: Monterey, California. 
3, MAHONIA AQUIFOLIUM (Pursh) Nutt. Gen. Pl. 1: 212.1818. 


Berberis aquifolium Pursh, Fl. Sept. 1: 219. 1814. 
Odostemon aquifolium Rydb., Bull. Torrey Club 43: 141. 1906. 


Odostemon nutkamus (DC.) Rydbe 1l.o. 


Stems erect, 8-20 dm. high, or sometimes dwarfed; leaf- 
lets 5-9, the lowest pair distant from the base of the rach- 
is, ovate-lanceolate, 3-7.5 cm. long, with usually 10 or 
more small spinulose teeth on each margin, bright glossy 
green above, light green beneath; racemes fascicled, 4-8 cm. 
long; berries blue and glaucous, on pedicels 10-15 mm. long, 

Coniferous woods, Humid Transition Zone; southwestern 
British Columbia to the Calipooia Mountains, Oregon, also ir 
northwestern Idaho. Type locality: The Cascades of the Col- 
umbie River. 


4, MAHONIA PIPERIANA Abrams, sp. nov. 


Frutex erectus 2-5 dm. altus; foliolis 5-9 frequenter 7, 


‘jugo inferiore a petioli basi distante ovato 2.5-6 cm. long. 


is, dentibus 7-9 spinosis. 

Stems erect, 2-5 dm. high; leaflets 5-9, commonly 7, the 
lower pair usually distant from the base of the rachis, 
ovate, 2.56 cm. long, spinose-dentate, with 7-9 teeth on 
each margin; upper surface glossy green and rather finely 
reticulate, the lower surface gray-green and papillate; ra- 
cemes 3-7 cm. long, densely or rather loosely fascicled; 
berry blue-black, ellipsoid-ovoid. 

Ysually on thinly wooded slopes, Arid Transition Zone; 


Jackson and Josephine Counties, Oregon, to Mendocino and 


nortrern Lake County, California. Type locality: Along the 
Pacific Highway, near the Siskiyou summit, Jackson County, 
Oregon, Abrams and Benson 10176, March 9, 1924. A fruiting 
specimen was collected in the same vicinity in September, 
1927 (Abrams 12136). 

Most California botanists have been referring these 
plants to M. aquifolium, but Piper clearly showed that they 
were not closely related to that species. He tentatively 
placed them with the then little understood M. dictyota, 4 
species we now know to be quite different. M. piperiana is 
fairly common over the range indicated, and is represented 
in the Dudley Herbarium by thirty-five different collect- 
ions. 


92 REY 7 0.8 G4) 2:6 Noe 2 
5. MAHONIA SONNEI Abrams, sp. nov. 


Frutex 3-5 dm. altus; foliolis 5, jugo inferiore a peti- 
oli basi distante ovato-lanceolatis; dentibus utrinque 13-16 
spinulosis; racemis congestis paucis elongatis. 

low shrub, 2-5 dm. high; leaflets 5, ovate-lanceolate, 
4-8 cm. long, glossy green above, pale beneath but not gray- 
ish or very obscurely so, the papillae being rather thinly 
scattered; racemes densely flowered, 4-7 em. long; berries 
blue-black, about 6 mm long. 

The type was collected on rocky banks of the Truckee 
River, Nevada County, California, by C. F. Sonne no. ll, 
August 11, 1884, and April, 1885. It is sheet no. 95828 in 
the Dudley Herbariun. : 

This species has long been confused with M. aquifolium 
which it superficially resembles in the shape of the leaf- 
lets and in their shiny green appearance, but the small num 
erous teeth and the papillate lower surface clearly indicate 
that its closest affinities are with M. repens. 


6. MAHONIA REPENS (Lindl.) G. Don, Gen. Hist. Pl. 1: 118. 
1831. 


Berberis repens Lindl. Bot. Reg. 14: pl. 1176. 1828. 
Odostemon repens Cockerell in Daniels, Fl. Boulder Colo. 

125. 1911. 

Stems creeping and stoloniferous, 1-2 dm. high; leaflets 
3-7, oval or rarely ovate, 3-9 cm. long, plane or nearly 80, 
dull green above, grayish-green beneath and papillate with 
minute microscopic protuberances; teeth usually 12 or more 
on a side, small and merely bristle-tipped; racemes densely 
many-flowered; berry ellipsoid-globose, 7-8 mm. long. 

Open pine forests, Arid Transition Zone; eastern British 
Columbia south along the eastern slopes of the Cascade 
Mountains to northeastern California, and east to Alberta, 
Nebraska, and New Mexico. Type locality: probably Montana. 


7. MAHONIA PUMILA (Greene) Fedde in Engler, Bot. Jahrb. 
31: 82, 1901. 


Berberis pumila Greene, Pittonia 2: 161. 1691. 
Odostemon pumilus Heller, Muhlenbergia 7: 139. 1912. 

Stem erect, 2-4 dm. high, rarely higher, simple or 
branched; leaflets 5-9, broadly oblong-ovate, blunt at the 
apex, dull green and strongly reticulate-veined above, glau- 
cous beneath, undulate and strongly spinose-dentate, the 
teeth 5-9 on each margin; racemes fascicled, many-flowered; 
berries blue-black, with @ bloom, oblong-ovoid, about 6 mm. 
long. 


1934 Abrams, Mehonias of the Pacific States 93 


Wooded slopes, Upper Sonoran and Arid Transition Zones; 
Rogue River, southern Oregon, south in the North Coast 
Ranges and the Sierra Nevada; also the Cuiamaca Mountains, 
California. Type locality: near Waldo, Oregon. 


8. MAHONIA DICTYOTA (Jepson) Fedde in "ngler, Bot. Jahrb. 
51: 89. 1901. 


Berberis dictyota Jepson, Bull. Torrey Club 18: 319. 1891. 

Odostemon dictyota Cockerell, Bull. Am. Mus. Nat. Hist. 24: 
91. 1901. 

Berberis californica Jepson, Fl. Calif. 549. 1921. 

Erect shrub, 5-20 dm. high, rather sparsely leafy; leaf- 
lets 5-7, broadly oblong to suborbicular in outline, strong- 
ly undulate and prominently spinose-toothed with 4-5 teeth 
on each margin, thick-coriaceous and prominently net-veined 
on both surfaces, pale green above and glaucous beneath; ra- 
cemes fasciculate; berries very glaucous, ebout 5 mm. long. 

Dry chaparral ridges, Upper Sonoran Zone; California from 
the dry interior foothills of the upper Sacramento Valley to 
San Diego County. Type locality: Marysville Buttes, Calif- 
ornia. 


9. MAHONIA FREMONTII (Torr.) Fedde in Engler, Bot. Jahrb. 
31: 89. 1901. 


Berberis fremontii Torr. Bot. Mex. Bound. 30. 1859. 
Odostemon fremontii Rydb., Bull. Torrey Club 33: 141. 1906. 

Erect shrub, with rigid branches, 1-3 m. high; leaflets 
3-7, commonly 5, rather narrowly ovate, 15-25 mm. long, cor- 
jaceous, pale gray-green on both surfaces, the spiny teeth 
prominent, usually 3 on each side; racemes terminating the 
numerous short stubby branchlets, usually 3-5-f lowered; 
bracts acuminate; pedicels slender, 10-16 mm. long; berries 
becoming inflated and dry. 

Dry mountain slopes of the desert regions, Upper Sonoran 
Zone; southern Nevada to southern Colorado south to Arizona, 
the deserts of southern California, and adjacent Lower Cali- 
fornia. Type locality: "On the tributaries of the Rio Vir- 
gen", southern Utah. 


10. MAHONIA NEVINII (A. Gray) Fedde in Engler, Bot. Jahrb. 
31: 102. 1901. 


Berberis nevinii A. Gray, Syn. Fl. 1: 69. 1895. 
Odostemon nevinii Abrams, Bull. N. Y. Bot. Gard. 6: 359. 
1910. 
Much branched shrub, 1-2 m. high; leaflets 4 or rarely 
5, lanceolate, the lateral 2-43 cm. long, the terminal often 


94 ae: ie ey Kis Dee Be ee No. 2 


nearly twice as long, gray-green on both surfaces, spinu- 
lose-serrate, the teeth numerous, bristle-like; racemes ter- 
minating the numerous stubby branchlets, loosely 57- 
flowered; bracts acuminate; berries densely covered with 
bloom. 

Dry sandy washes, Upper Sonoran Zone; a rare species, 
known only from the eastern end of San Fernando Valley, 
southern California. 


(1) Piper, Charles V. The identification of Berberis 
aquifolium and Berberis repens. Contr. U. S. Nat. Herb. 20:3 
437-451, pl. 24-26. 1922. 


STUDIES OF NEW AND NOTEWORTHY TROPICAL AMERICAN PLANTS — II 


Harold N. Moldenke 


AEGIPHILA BREVIFLORA (Rusby) Moldenke, comb. nov. 


Pseudaegiphila breviflora Rusby, Mem. N. Y. Bot. Garde 7: 
ah. 1927. 


ALOYSIA LIGUSTRINA var. SCHULZII (Stand1.) Moldenke, 
comb. nov. Lippia ees var. Schulzii Standl., Field 
Mus. Publ. Bote $3 2 e 1929. 


ALOYSIA MACROSTACHYA (Torr.) Moldenke, comb. nove 


Lippia Wrightii var. macrostachya Torr. Bot. Mex. Bound. 
127. 185. 


ALOYSIA PULCHRA (Briq.) Moldenke, comb. nov. 
Lippia pulohra Briq., Arkiv Bot. 2, no. 10: 18. 1904. 


ALOYSIA SCORODONIOIDES var. DETONSA (Briq.) Moldenke, 
comb. nov. Lippia scorodonioides var. detonsa Briq., Bull. 
Herb. Boiss. i 39. 1896. 


ALOYSIA SCORODONIOIDES var. MATHEWSII (Briq.) Moldenke, 
comb. nov. Lippia scorodonioides var. Mathewsii Briq., Bull. 
Herb. Boiss. rs 530. 1896. ; 


ALOYSIA SPATHJLATA (Hayek) Moldenke, comb. nov. 
Lippia spathulata Hayek in Engl. Bot. Jahrb. 42: 165. 1908. 


ALOYSIA VIRGATA var. LAXA (Chod.) Moldenke, comb. nov. 
Lippia urticoides var. laxa Chod., Bull. Herb. Boiss., ser. 
2, 2: 819. 1902. 


ALSEIS MUTISII Moldenke, sp. nov. Frutex vel arbor; ram 
ulis gracilibus plusminus tetragonis brunneis glabratis; no- 
dis anmlatis; internodiis 1.5--1.7 cm. longis; foliis de- 
cussato-oppositis stipulatis; stipulis parvis caducis; pet- 
iolis gracilibus 5--12 mm. longis glabria; leminis firme 
membranaceis utrinque nitidis et prasinis, oblongis vel ob- 
longo-ellipticis 5.5--10.3 cm. longis, 1.7--3.3 cm. latis, 
abrupte acutis vel subacuminatis integris, ad basin acutis, 
supra glebris, subtus juventute levissime breviterque fur- 
furaceo-puberulentibus senectute subpuberulentibus vel gla- 
bris; costa gracili, subtus promimulente; venis secundariis 
gracilibus frequentibus ubique ca. 10 brevibus adscendenti- 


$5 


96 PHYTOLOGIA No, 2 


bus saepe vix arcuatis utringue prominulentibus; reticulo 
venarum venularumque utringue subconspicuo prominulente; in- 
florescentiis paniculatis terminalibus; paniculo 15--22 cm. 
longo et 14--15 cm. lato, racemis spiciformibus ca. 9, ped- 
unculo (2--2.2 cm. longo) et symodiis (1.4--3.5 cm. longis) 
firmis brunneis dense breviterque puberulentibus; bracteis 
foliaceis caducis, 2 quamque jugam racemorum subtentibus, 
foliis consimilibus sed constanter deminuentibus; hypanthio 
cae 1.5 mm longo et 1 mm. lato puberulente; lobis calycis 5 
usque ad summam hypanthii disjunctis, elliptico-—lingulatis 
cae O.7 mm. longis et 0.5 m,. latis glabris, ad apicem ro- 
tundatis; tubo corollas ca. 1.8 mu. longo et constanter 1.35 
mm. diametro, extra glabro, intus villoso, lobis 5 ca. 0.6 
mn. longis et 1 mm latis late ovatis ad apicem rotundatis; 
staminibus 5, ad basin tubae corollae insertis, corollam ae- 
quantibus; filementis crassis conicis, ad basin complanatis, 
densissime longeque villosis, pilis sufflavis adscendenti- 
bus; pistillo longe exserto; stylo ca. 3.6 mm. longo glabro, 
ad apicem bifido, ramlis stigmatiferis divaricatis subre- 
curvatis ca. 0.6 mm. longis; placentis ab apice ovarii pen- 
dulis miltiovuliferis. 

The type of this species was collected by José Celestino 
Mutis (No. 2948), probably in the vicinity of Bogota, Cun- 
dinamerca, Colombia, and is deposited in the United States 
National Herbarium (No. 1,561,457). The specimen was re- 
ceived by the United States National Museum from the Jardin 
Botanico in ‘iadrid in 1942 and the number is an arbitrary 
one assigned by ©. Pe Killip. The inflorescences are com 
posed mostly of 4 pairs of opposite racemes and one termin- 
al one, the individual racemes being 4.5--1l cm. long, the 
longest being the terminal one and the lowermost pairs, 
each raceme about 1 cm. wide in anthesis. The species is re- 
lated to A. leiantha Blake, from Venezuela, but differs in 
its much more slender twigs, its slender retioles, its 
smaller and firmer leaf-blades with fewer secondaries, its 
more paniculate inflorescences and shorter and more slender 
racemes, and its much smaller flowers. 


AVICENNIA NITIDA var. TRINITENSIS Moldenke, var. nove 
Haec varietas a forma speciei typica recedit in omibus 
partibus, praesertim laminis foliorum, glabris vel subglab- 
ris et plerumque nitidis. ; 

The type of this variety was collected by R. L. Brooks 
at Caroni Swamp, Trinidad, May 29, 1932 (Trinidad & Tobago 
Botanical Garden no. 12,656) and is deposited in the herb- 
arium of the New York Botanical Garden. Herb. Trin. & Tob. 
Bot. Gard. no. 12,651, collected at the same locality by R. 
C. Marshall on May 9, 1932, is the same variety, as is also 
Britton, Hazen, & Mendelson 541 from Patos Island, March 
13, 1920. The complete absence of pubescence from the nitid 


1934 Moldenke, Tropical American Plants 97 
lower leaf-surfaces is most characteristic. 


CASSELIA ILICIFOLIA Moldenke, sp. nov. Fruticuius; ramis 
gracilibus 4-angulatis et 4-costatis glabratis, cortice 
griseo vel albido obtectis; ramlis et sarmentis gracil l- 
mis tetragonis brunneis vel brumneo-luteis dense breviter— 
que pubescentibus vel puberulentibus; internodiis abbrevia— 
tis 0.8--3 cm. longis; foliis sessilibus oppositis vel ter- 
natis; laminis ellipticis vel ovatis 7--15 mm. longis 3—14 
mm. latis utrinque puberulentibus, margine dense irregulare 
iterque spinosis, majoribus ad basin amplissimis et trunca- 
tis, minoribus in sarmentis perabbreviatis confertis per- 
angustioribus et ad basin acutis; pedunculis axillaribus 
oppositis vel ternatis gracillimis 1.2-—-2 cm. longis dense 
puberulentibus vel breviter pubescentibus 2—4—floris 
(plerumque 2-floris), ad apicem jugam bracteolorum linearum 
dense puberulentum 2--6 mm. longorum gerentibus; calyce 
tubuloso 6--10 mm. longo 2.5--3 mm. diametro plusminus 5- 
costato dense puberulente vel breviter pubescente, margine 
S-dentato, dentibus linearibus 1—2 mm. longiss; corolla hy- 
pocrateriformi zygomorpha, tubo fere recto, limbo 5~lobato; 
staminibus 4 didynamis prope basin tubae corollae insertis, 
brevisaimis; stylo gracili ca. 2.5 mm. longo glabro, ad a- 
picem stigma peltatum oblique gerente; ovario ca. 1.4 mm. 
longo glabro 2-loculare, quoque loculo l-owlato. 

The type of this species was collected by Frederico 
Schmaedke at la Diana, Ulapes, Rioja, Argentina, February 
24, 1907 (T. Stuckert no. 17,013), and is deposited in the 
herbarium of the Conservatoire et Jardin Botaniques at Gen- 
eva. Stuckert nos. 4724; 4734; 17,014; and 22,502, from the 
same locality, represent the same species. C. ilicifolia 
may be distinguished at once from all other members of the 
genus by its spiny-margined leaf-blades. 


CITHAREXYLUM LINEARIFOLIUM Moldenke, sp. nov. Frutex 
usque ad 4m altus; ramis et ramlis gracilibus stramin- 
eis glaberrimis laevisque, non lenticellatis; nodis non 
anmilatis; internodiis 1--5.5 om. longis; cicatricibus fol- 
iorum delapsorum in sterigmatibus sitis, sterigmatibus in- 
Signibus gracilibus adscendentibus 1--3.5 mm longis gla- 
bris et in latere abaxiali projecturam longam rostelloid- 
eam gerentibus; foliis decussato—oppositis vel ternatis 
(vel interdum suboppositis vel subspiraliter dispersis); 
petiolis gracilibus 3—6 mm. longis glabris; laminis mem 
branaceis utrinque atro-viridibus et pernitidis lineari- 
bus 4,516 em. longis 3—6 mm. latis integris, ad apicem 
subulato-acuminatis, ad basin longe attenuatis, utrinque 
glaberrimis, ima basi jugam glandularum discoidearum nig- 
rarum gerentibus; costa gracili utrinque prominulente; 
venis secunderiis non dilucidis; reticulo venarum venu- 


98 PHYTOLOGIA No. 2 


larumque utrinque plusminus prominulente, praesertim supra, 
plerumque subtus obscuro; racemis terminalibus et sarmenta 
brevia axillaria terminantibus, erectis vel nutantibus 5-- 
10 cm. longis ca. 1.5 cm. latis densiuscule miltifloris 
simplicibus; pedunculis gracillimis 5--9 mm. longis gla- 
bris; rhachide gracillimo glabro; pedicellis gracilibus us- 
que ad 1 mm. longis glabris; bracteis et bracteolis nullis; 
prophyllis lineari-subulatis 1--2 m. longis glabretis; 
calyce tubuloso ca. 3.6 mm. longo st 2 mm disametro glebra- 
to, margine 5-dentato, sinubus i tenuissimis, sinu 1 per- 
profundo (ca. 0.7 mm profundo); corolla hypocrateriformi 
alba glabra suaveolente, tubo cylindrico ca. 6.2 mme longo, 
intus piloso, limbo 5-partito, lobis elliptico-lingulatis 
vel subspathulatis ca. 3.6 mm. longis et 2.3 mm latis ven- 
osis, ad apicem rotundatis; staminibus 4 prope apicem tubae 
corollae insertis inclusis; filamentis filiformibus ca. 0.7 
mm. longis; antheris oblongis ca. 1 mm. longis; pistillo 
incluso; stylo crassiusculo ca. 2.2 mm. longo levissime 
puberulente; stigmate breviter bifido, ramulis complanatis 
cae O.7 mm longis et 0.5 mm. latis; ovario subgloboso ca. 
1 mm. longo et lato glabrato 4-loculari (sed non perfecte, 
ut videtur 2-loculeri); calyce fructifero indurato leviter 
cupuliformi ca. 3 mm. longo et 6.5 mm. diametro glabro, 
margine subprofunde 5-lobato, lobis rotundatis; fructibus 
oblongis usque ad 16 mm. longis et 9 mm. latis carnosis 
nitidis glabris 2-lobatis. 

The type of this remarkable species was collected by 
Edward Palmer in Paraguay during the exploration of the Rio 
la Plata and adjacent territory in the years 1853—1856, in 
charge of Capt. T. J. Page, and is deposited in the United 
States National Herbarium. It has been collected both in 
anthesis and in fruit in October and in March. The species 
is unmistakable because of its long linear leaves. Its 
sterigmata are also unique and its leaf-arrangement note- 
worthy. J 

PHYLA NODIFLORA var. CANESCENS (H.B.K.) Moldenke, stat. 
nov. Lippia canescens H.B.K. Nov. Gen. & Sp. Pl. 2: 263. 
1817. 


PHYLA NODIFLORA var. SERICEA (Kuntze) Moldenke, state 
nov. Lippia nodiflora var. normalis f. sericea Kuntze, Rev. 
Gen. Pl. 2: 508. 1891. 


PINGUICULA PUMILA var. BUSWELLII Moldenke, var. nov. 
Haec varietas a forma speciei typica recedit corolla tota 
butyroideo- vel aureo-flava. 

The type of this very rare variety of butterwort was 
collected by Harold N. Moldenke (No. 98la) in a moist sandy 
pineland along the olf Tamiami Trail, Collier Co., Florida, 


te i = ee | 
Sate 


1954 Moldenke, Tropical American Plants 99 


April 14, 1930, and is deposited in the herbarium of the 
New York Botanical Garden. The variety was first discovered 
by Mr. Walter M. Buswell, of Fort Myers, Fla., in May, 
1914, in whose honor it is named. It is known from only two 
localities: (1) near what is known as the "Welsh Grove" on 
the east side of the Big Cypress Swamp, about 10 or 12 
miles northwest of Deep lake, in Collier County, and (2) 
about 1 1/2 miles south of Bonita Springs on the old Ta- 
miami Trail, also in Collier County (about 1 mile over the 
Lee County line). These two localities are approximately 
opposite each other in an east-west line, one being on the 
east side and the other on the west side of the Big Cypress 
swampe It was collected at the second locality (the type 
locality) by Mies Jeanette P. Standley on October 11, 1916, 
and specimens were distributed by her as No. 307 to the 
United States National Herbarium and the New York Botanical 
Garden. Those who have seen this plant growing have dif- 
ficulty in believing that it is not a totally different 
species from P. pumila Michx. Dr. John Hendley Barnhart, 
however, is convinced that it is a mere color form of the 
latter widely distributed species of the southeastern Uni- 
ted States and the West Indies, and it is so regarded by 
Small in his "Manual of the Southeastern Flora", page 1232 
(1933). In the typical form of P. pumila, however, the 
corolla is white, pale violet, or pale rose, while in the 
variety which we are here describing the corolla is of a 
rich golden-yellow or butter-yellow throughout, even as it 
is in P. lutea Walt. In the opinion of the present writer 
this pretty yellow-flowered butterwort deserves at least 
varietal designation. 


RECORDIA Moldenke, gen. nov. Frutices vel arbores. 
Folia opposita. Inflorescentia racemosa terminalis muilti- 
flora. Calyx tubuloso-campanulatus, in latere inferiore 
(abaxiali) fissus, 5-costatus, costis 4% superioribus in 
apiculationes brevissimes terminantibus, costis 2 infer- 
ioribus in apiculationes nullo modo vel perobsolete term 
inantibus. Corolla hypocrateriformis zygomorpha non valde 
curvata; tubo obconico, lobis 5, 2 supremis (adaxialis) 
minimis, lateralibus 2 mediocriter, lobo infimo in magni- 
tudine maximo. Stamina fertilia 4 didynama infra oram tubae 
corollae inserta, inferiora 2 maxima, stamen quintum ad 
staminodium filiforme reductum. Antherae sagittatae dorsi- 
fixae, thecis ad basin divergentibus. Stylus simplex term 
inalis, ad apicem lateraliter curvatus, arcu convexo com 
planato stigmatifero. Ovarium fusiforme, ad apicem in sty- 
lum abrupte attenuatum, 2-loculare, quoqgue loculo l- 


ovulato. 


RECORDIA BOLIVIANA Moldenke, sp. nov. Frutex vel arbor; 


100 Pro (Y 27:0 L-G-5 a No. 2 


ramis et ramulis crassiusculis subteretibus griseis glabris; 
sarmentis et innovationibus immeturis brunneis et dense brev 
iterque cano~pubescentibus; nodis anmlatis; cicatricibus 
foliorum sessilibus; internodiis abbreviatis 5-16 mm. 
longis; foliis decussato-oppositis plerumque ad apicem sar- 
mentorum juvenilium confertis; petiolis subgracilibus 3--12 
mm. longis dense breviterque pubescentibus; laminis leviter 
membranaceis vel chartaceis atro-viridibus, in siccitate 
nigrescentibus, ellipticis 5.2--10.5 cm. longis, 2,.2--4.4 
cm. latis, breviter acuminatis, argute sed minute serratis 
fere usque ad mediam vel subintegris, ad basin abrupte acu- 
tis vel breviter cuneatis, non glanduliferis, supre parce 
etrigilloso-puberulentibus vel glabratis, subtus (praesertim 
in venis majoribus et in juventute) dense breviterque pubes- 
centibis vel interdum subvelutinis; costa gracili, supra 
plana vel impressa, subtus prominente; venis secundariis 
utroque 5-——7 adscendentibus saepe non valde arcuatis, ad 
marginem anastomosantibus; reticulo venulorum gracili saepe 
obscuro; racemis erectis 7--14 cm. longis et usque ad 2.5 
cm. latis densissime miltifloris ramilos et sarmenta axil- 
laria terminantibus; pedunculo gracili abbreviato 5--8 mm. 
longo dense cano-pubescente; rhachide gracili dense cano= 
pubescente; pedicellis gracillimis 2--4,5 mm. lorngis dense 
pubescentibus; prophyllis minutis setaceis plerumque non 
conspicuis; floribus plerumque in fasciculis 2-, 3, vel 
multifloris secus rhachidem dispositis, numerosissimis; 
calyce tubuloso-campanulato ca. 4.9 mm. longo et 2.5 m. 

(ad apicem) lato, extra densiuscule pubescente, intus gla- 
bro, in latere inferiore (abaxiali) usque ad 1.8 mm. fisso, 
5-costato, costis 4 superioribus in apiculationes brevis- 
simas ca. 0.2--0.3 mm. longas terminantibus (costa centra- 
li saepe paullo majori quem costis 2 lateralibus), costis 2 
inferioribus in apiculationes obsoletas vel millas termin- 
antibus; corolla hypocrateriformi zygomorpha non valde cur- 
vata, tubo obconico ca. 6.5 mm. longo et ad basin 0.5 mm 
lato, supra usque ad 3.6 mm. infra limbum ampliato, extra 
glabro, intus ad oram leviter breviterque pubescente, lobis 
5 extra leviter puberulentibus intus breviter pubescenti- 
bus, lobis 2 supremis (adaxialibus) minimis ovato-lingulatis 
cae 1 mm longis et ad basin 1.5 mm. latis ad apicem rotun- 
datis, lobis 2 lateralibus mediocriter magnis triangulato- 
ovatis cae 2--3 mm. longis et ad basin 2--2.8 mm. latis ad 
apicem rotundatis, lobo infimo triangulato-ovato ca. 3.5 MMe 
longo et ad basin 3 mm. lato; staminibus fertilitus 4 didy-— 
namis ca. 2 um. infra oram tubae corollae insertis, infer- 
ioribus 2 maximis (filamentis ca. 4.2 mm. longis), lateral- 
ibus 2 brevioribus (filamentis ca. 1.2 mm. longis), stamine 
quinto ad staminodium filiforme ca. 1 mm. longum et ca. 1 
mm. infra stamina fertilia insertum reducto; antheris sag- 
ittatis dorsifixis ca. 1 mm. longis, thecis ad basin diverg~ 


a 


1934 Moldenke, Tropical American Plants 101 


entibus; stylo simplici terminali ca. 7.1 mm. longo glabro, 
apicem versus angustato, ad apicem lateraliter flectente, 
supra convexo ovato complanato et stigmatifero; ovario fus- 
iformi ca. 1.3 mm. longo et 0.5 mm. diametro, ad apicem in 
stylum abrupte attenuato, glabro 2-loculari, quoque loculo 
l-ovulato; ovulis in axilla exteriori dissepimenti circa ad 
dimidiam ovarii affixa; calyce fructifero et fructibus non 
visis. Fig. 13. 

Type collected by José Steinbach (No. 7240) at Batiado, 
Rio Surutu, alt. 400 m, Sara, Santa Cruz, Bolivia, October 
1, 1925, and deposited in the herbarium of the Naturhistor- 
iska Riksmuseet at Stockholm. It ascends to 800 m. and seems 
to be endemic to the mountains of Bolivia. 

The genus Recordia is a member of the Verbenoideae - 


Petraceae of Briquet, although in habit it greatly resembles 


the genus Citharexylum. Very superficially the genus also 
resembles the rubiaceous genus Alseis, especially in habit, 
but the epigynous flowers of the latter gems of course at 
once distinguish it from ours. 


VITEX KLUGII Moldenke, sp. nov. Arbor 4m. alta; ramlis 
gracilibus tetragonis plusminus brevissime strigoso-nilosis 
vel puberulentibus; foliis oppositis 3-foliolatis; petiolis 
gracilibus ca. 6 cm. longis puberulentibus, supra sulcatis, 
ad apicem ampliatis; petiolulo centrali ca. 12 mm. longo, 
petiolulis lateralibus 6--7 mm. longis, parce strigoso— 
pilosis; laminis foliolorum membranaceis ellipticis 18--22.5 
em. longis, 8.5=-10.2 cm. latis (centrali quam lateralibus 
paullo majori) integris, ad apicem acuminatis, ad basin acu- 
tis, utrinque glabris vel in nervatione supra sparsissime 
subtus plusminus parce strigillosis; venis secundariis utro- 
que 10--12, ad marginem arcuatim confluentibus; inf lores- 
centiis cymosis supra~axillaribus oppositis usque ad 6.5 cm. 
longis et ca. 4 cm. latis ubique brevissime strigosis ca. 


'20-floris insigniter dichotomis, ramalis cymorum complana- 


tis ad apicem amplissimis; pedicellis filiformibus 1.5--2 
mn. longis dense strigosis; floribus caeruleis et albis 
(teste Klug); calyce tubuloso-campanulato ca. 5.4 mm. longo, 
2.6—2.8 mm. diametro, breviter strigoso (pilis albidis 


arcte adpressis), margine S-dentato 2-labiato, dentibus 2 


majoribus 4 minoribus; corolla zygomorpha, tubo recto cylin- 
drico ca. 7./ mm. longo, extra dense strigoso-pubescente 
(praeter 1.5--2.5 mm. inferiorum longitudinis), intus ad 
oram longe piloso, ad basin 1.5 mm. lato, ad apicem usque 

ad 4.1 mm. ampliato, limbo 5—partito, lobis 4 parvis triang- 
ulari-ovatis ca. 2.6 mm longis et ad basin 1.8 mm. latis 
usque ad apicem acutam angustatis, lobo quinto magno 61 lip- 
tico ca. 4.9 mm longo et 2.8 mm lato, ad apicem acuto; 
staminibus 4 subaequalibus, exsertis, ca. 2.6 ma. supra bas- 
in tubae corollae insertis; filamentis ca. 7 mm. longis ad 


102 PHYTOLOGIA No. 2 


basin longe pilosis, superne glabris; pistillo longe exser- 
to; stylo gracili ca. 14 mm. longo glabros; stigmate bifido, 
ramilis 0.5 mm. longis divergentibus; calyce fructifero et 
fructibus non visis. 

The type of this species was collected by G. Klug (No. 
625) in a forest at Mishuyacu, near Iquitos, alt. 100 m, 
Loreto, Peru, in December, 1929, and is deposited in the 
herbarium of the New York Botanical Garden. 


VITEX LUCIDA Moldenke, sp. nov. Arbuscula usque ad 5 me 
alta; ramlis gracilibus obtuse tetragonis plusminus ad a- 
picem breviter flavo-pubescentibus, internodiis demum gle- 
brescentibus; foliis oppositis 3--S-foliolatis; petiolis 
gracilibus 3--4.5 cm. longis breviter flavo-strigillosis, 
supra complanatis et canaliculatis, ad apicem non amplia- 
tis; foliolis in magnitudine inter se valde variis, plerun 
que foliolo centrali maximo, foliolis 2 lateralibus medio- 
cris, et foliolis 2 basalibus parvis (interdum minimis vel 
mullis!); laminis foliolorum firme membranaceis atro-virid- 
ibus nitidis ellipticis vel oblanceolatis usque ad 8.5 cm. 
longis et 4.2 mm. latis integris, ad apicem obtusis vel 
acutis (interdum in foliolis minimis retusis), ad basin 
acutis vel interdum subrotundatis, utringque praeter costam 
leviter strigillosam glabratis; petiolulis gracilibus 1--10 
mm. longis strigillosis in foliolo centrali longissimis, in 
foliolis basalibus brevissimis; inflorescentiis axillaribus 
pppositis thyrsoideis non divaricatis usque ad 15 cm. long- 
ie et cae 325 cm. latis, laxe sparseque multifloris; pedun= 
culo 4--5.5 cme longo sympodiisque pergracilibus minute 
strigillosis,ramulis trifloris in ca. 4 jugam et 1 termin- 
alem dispositis; pedicellis pergracilibus 1--11 mm. longis 
glabris; calyce cupuliformi zygomorpho ca. 1.9 mm. longo et 
2 mm. diametro glabro, margine plusminus bilabiato 5-denta- 
to, dentibus parvis acuminatis, sinubus 3 parvis et 2 pro- 
fundis; corolla hypocrateriformi violacea, extra minute 
puberulente, intus dense pilosa, tubo infundibulariformi 
ca. 4,1 mm. longo recto, ad basin glabram ca. 1.5 mm. lato, 
ad apicem valde ampliato et 4.6 mm. lato, limbo 5S-partito, 
lobis 4 ovatis parvis ca. 2.3 mm. longis et 1.8 mm. latis 
acutis, lobo quinto permagno late spathulato venoso cae 5e2 
mm. longo, ad basin et usque ad 1.5 mm. longitudinis ca. 
2.8 mm. lato, apicem versus valde dilatato ca. 4.9 mm. 
lato, ad apicem abrupte breviterque acuminato; staminibus 
4, didynamis ca. 5.9 mm longis, fere ad basin tubae cor- 
ollae insertis, exsertis; filamentis gracilibus pilosis, ad 
basin ampliatis complanatisque et densiore longioreque pil- 
osis; antheris obcordatis ca. 0.6 mm. longis et latis; pis- 
tillo exserto; stylo ca. 5.2 mm. longo glabro, ad apicem 
bifido, ramlis stigmatiferis ca. 0.5 mm longis divarica- 
tis; ovario subgloboso ca. 1 mm. longo et diametro subglab- 


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1934 Moldenke, Tropical American Plants 103 


ro; calyce fructifero patelliformi indurato plerumque valde 
bilabiato ca. 2.5 mm longo et 5——6 mm. diametro, margine 
scarioso; fructibus subglobosis flavis, in siccitate levi- 
ter 4~suleatis. 

The type of this hitherto neglected species was collected 
by E. J. Valeur (No. 630) in an open pine forest at Jicomé, 
Loma Bajita, district of San José, prov. Santiago, Dominican 
Republic, at an altitude of 600—700 m., May 13, 1931, and 
is deposited in the herbarium of the New York Botanical 
Garden. Additional specimens are Valeur 695 from the Domin- 
ican Republic and Leon 11,801 and C. Wright 1355 from Orien- 
te, Cuba. Common names recorded by Valeur are "palo perrito" 
and "mata becarro",. 

The species is related to V. umbrosa Sw., of Jamaica, 
with which it has hitherto been confused. The latter, how- 
ever, differs in its much larger and more acute leaflets, 
its mich longer and stronger petioles and petiolules, the 
former being prominently ampliate and disk-shaped at the a- 
pex, its large inflorescences, longer peduncles, puberulent 
calyx, and many other characters. V. divaricata Sw. is at 
once distinguished by its conspicuously divaricate inflor- 
escencese 


VITEX RUSBYI Moldenke, sp. nov. Arbor usque ad 15 m 
alta; ramis crassis tetragonis furfuraceo-pubescentibus; 
ramulis et sarmentis densissime ferrugineo-velutinis; fol- 
iis oppositis 5--7-foliolatis; petiolis crassiusculis us- 
que ad 10.5 cm. longis dense ferrugineo-velutinis, supra 
complanatis et canaliculatis, ad apicem ampliatis; petio- 
lulis 1--6 mm longis velutinis, in longitudine inter se 
variis, petiolulo centrali plerumque maximo; laminis fol- 
iolorum membranaceis supra in siccitate brumnescentibus, 
ellipticis vel oblongis, usque ad 7.7 cm. longis et 2.8 
cm. latis acutis integris, ad basin acutis, supre brev- 
iter pubescentibus, subtus dense ferrugineo=-velutinis; 
inflorescentiis axillaribus numerosis cymosis mitiflor- 
isp 2.5—5.5 cm. longis et 2-—-3.5 cm, latis divaricatis; 
pedunculo crassiusculo complanato 1.2<—3 cm. longo fur- 
furaceo-pubescente vel dense ferrugineo-velutino; pedi- 
cellis 1--1.8 mm longis velutinis; calyce campanulato 
crasso plusminus zygomorpho ca. 3.9 mm. longo et 4.2 mm 
diametro velutino-pubescente, margine variabile plerum- 
que plusminus 2-labiato et 5-lobato (interdum labio uno 
fere elobato), lobis triangularibus acutis interdum re- 
flexis; corolla hypocrateriformi extra dense velutino=- 
pubescente subrecta vel curvata, tubo plusminus urceola- 
to ca. 7.2 mm. longo, intus piloso, ad basin glabram ca. 
1.8 mm. lato, medium versus ca. 4.1 mm, lato, ad apicem 
ca. 363 mm. lato, limbo 5-partito, lobis 4 parvis ovatis 
ellipticis ca. 2.5 mm. longis et 2.6 mm. latis acutis, 


104 PHYTOLOGIA Nos 2 


lobo quinto magno lete spatmlato ca. 3.6 mm longo, ad bas- 
in cae 2.3 mm. lato, apicem versus valde dilatato et ca. 4.6 
mm. lato, ad apicem rotundato vel subacuto; staminibus 4 dts 
tincte didynamis fere ad basin tubae corollae insertis ex 
sertis cae 5.2 mm. longis; filamentis ad basin complanatis 
ampliatisque et dense longeque pilosis, supra basin subglab= 
ris; antheris parvis obcordatis; pistillo exserto; stylo ca. 
7.5 mm longo glabro, ad apicem bifido, ramlis stigmatifer- 
ig ca. 0.5 mm. longis divaricatis; ovario subgloboso ca. lee 
mm. longo et diametro subglabro; calyce fructifero et fruct- 
ibus non visis. 

The type of this handsome species was collected by Henry 
Hurd Rusby and Francis Whittier Pennell (No. 1143) in a 
light forest on a rocky hill at the gorge above Natagaima, 
Huila, Colombia, August 12, 1917, and is deposited in the 
~ herbarium of the New York Botanical Garden. The collectors 
describe the flowers as lavender—blue. Other specimens of 
this species are Rusby & Pennell 221, André 19h2 and 4155, 
and Rose & Rose 23,488. 

VITEX TRIFLORA var. QUINQJEFOLIOLATA Moldenke, var. nov. 
Haec varietas a forma speciei typica et omnibus aliis var—- 
fetatibus recedit foliis 5-foliolatis. 

The type of this distinct variety was collected by B. . 
A. Krukoff a 5765) on terra firma near the mouth of the 
Rio Macauhan (tributary of the Rio Yaco), lat. 9°20' S., 
long. 69° W., Acre Territory, Brazil, September 3, 1933, 
and is deposited in the herbarium of the New York Botanical 
Garden. The collector describes the plant as a tree 45 feet 
tall, with a stem-diameter of 3 inches. 


Explanation of Figure 13 
A. Branchlet X 0.5 
B. Lateral view of one flower X 2 
C. Front view of calyx X 2.6 
D. Cross-section of ovary X 16 
E. Top of style and stigma X 2.5 
F. One of the large stamens X 2.5 


G. One of the emall stamens X 2.5 


H. Oorolla spread open X 2.5 


105 


Moldenke, Tropical American Plants 


1934 


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PLANTAE KRUKOVIANAE - III 


H. A. Gleason 


The species described below were collected by Mr. B.A. 
Krukoff on his fourth expedition to Brazilian Amazonia. The 
types are deposited in the herbarium of the New York Botani- 
cal Garden and are duplicated in several other herbaria. 


RYANIA SAURICIDA Gleason, sp. nov. Arbor parva 6 m. alta 
ramulis gracillimis angulatis glabris; petioli 2-3 mm. longi 
breviter stellato-pilosi; laminae membranaceae, oblongae vel 
elliptico-oblongae, 10-18.5 om. longae, 4-6 cm. latae, acum- 
inateae et breviter mucronatae, integrae, basi acutae, utrin- 
que glaberrimae, venis lateralibus in utroque latere ca. 7 
curvato-adscendentibus, venulis utrinque creberrime reticu- 
latis et paulo elevatis; stipulee persistentes setaceae 5-7 
mm. longae breviter hispidulae; pedicelli in axillis solita- 
rii ca. 5 mm. longi curvati crassi angulati tomentosi, pilie 
stellatis fuscis; flores jam immaturi; calyx in alabastro 
inaperto 10 mm. longus anguste conicus acutus, dense fusco- 
stellato-tomentosus; filamenta nondum elongata; antherae an-— 
guste lineares jam usque ad 5 mm. longae basi sagittatae ap- 
ice mucrone 0.5 mm. longo ornatae; ovarium sessile ovoideum 
densissime hirsutum; stylus crassus glaber 4.5 mm. longus 
(verisimiliter in floribus maturis longior) apice brevissime 
6-fidus, stigmatibus 6 carnosis rotundatis. 

Type, Krukoff 5815, from Sepatini, on the Purus River, 
Amazonas. It inhabits the "restinga alta," just above the 
varzea land above water at most time of the year. 

The genus Ryania apparently contains about ten species. 
In R. Riedeliana and R. Sagotiana the ovary is stipitate; in 
R. canescens and R. Mansoana the leaf is tomentose beneath; 
in R. dentata the short anthers are not mucronate; in R. 
Candollei, chocoensis, speciosa, and stipularis the leaves 
are pubescent beneath and usually rounded at the base. The 
remaining species, R. acuminata Spruce, appears to be most 
closely similar to ours, but differs in its smaller and pro- 
portionately narrower leaves, which are more narrowly acute 
at the base. The bark of R. sauricida is used by Indians for 
poisoning alligators, whence the specific name» 


TRIOHILIA KRUKOVII Gleason, sp. nov. Sect. Moschoxylum: 
arbor excelsa gracilis 25 m. alta; folia alterna breviter 
petiolate tota longitudine usque ad 17 cm. metientia, rhach- 


106 


1934 Gleason, Plantae Krukovianae - III 107 


ide supra canaliculato minutissime puberulo; foliola subcor- 
iacea, anguste elliptico-oblanceolata 8--12 cm. longa 2.5-—4 
em. lata (infima multo breviora), abrupte acuminata, apice 
ipso obtuso, integra, basi longe cuneata, supra glabra sub- 
tus minutissime pubescentia, venis secundsriis utroque late- 
re ca. 12 leviter curvato-adscendentibus supra planis subtus 
elevatis; paniculae confertae ramosae rhachidem foliorum ae- 
quantes vel dimidio excedentes, strigulosae, bracteolis tri- 
angulari-ovatis arcte adpressis strigosulis acutis 0.5--0.8 
mm. longis; pedicelli veri crassi 0.5--1 mm. longi; flores 
4-meri; calyx gemosepalus 3 mm. diam., ad medium in lobos 4 
late rotundatos vel truncatos fissus, extra leviter strigo- 
sulus; corolla ad anthesin 6 mm. diam., extra parce strigos- 
ula, petalis carnosis recurvo-patulis trianguleri-ovatis 
acutis 3.5 mm. longis; staminum tubus erectus 1.6 mm. altus 
5.2 mm. diam., extra tenuissime puberulus intus breviter 
villosus, margine breviter 16-denticulatus, dentibus 8 alt- 
ernis setaceo-subulatis erectis 1 mm. longis parce villosul= 
is; antherae 8 sessiles anguste triangulares 1 mm. longae; 
discus carnosus obscure 8-gibbosus, cum ovario late ovoideo 
0.7 mm. longo dense pubescens; ovarium 2-loculare, ovulis in 
quoque loculo 2 ex apice collateraliter pendentibus; stylus 
crassus erectus glaber 0.8 mm. longus, stigmate subcapitato 
integro3; capsula bivalve atropurpurea (tantum in sicco) ang- 
uste obovoidea 15--18 mm. longa, valvis post maturitatem 
divergentibus recurvatis et arcte involutis; arillus ruber. 

Type, Krukoff 4711, collected on varzea land at the 
mouth of the Embira River, Jurua Basin, Amazonas. 

The species is most closely related to Trichilia Cipo 
(Juss.) 0.DC., as illustrated by Spruce 2237 from the Rio 
Negro. De Candolle's description, in fact, indicates very 
little by which the two might be separated, but a dissection 
- of the Spruce specimen shows that the anthers are distinctly 
ovoid and just half as long (0.5 mm.) and that the subulate 
teeth of the staminal tube are only a fifth as long (0.2 
mm.) as in our plant. The fruits of T. Krukovii are regul- 
arly 2-valved, according to the collector, and we are not 
acquainted with any other species in which the valves become 
recurved and involute. 


BIXA EXCELSA Gleason & Krukoff, sp. nov. Arbor 40-metra- 
lis trunco gracili; petioli graciles glabri longitudine 
inter 25 et 70 mm. variabiles; laminae subcoriaceae utrinque 
glabrae et virides ovateae 10.5--19 om. longae 4.5--9 om, 
latae, sensim longe acuminatae, basi late obtusae subrotund- 
atae vel rarissime subtruncatae minquam cordatae, 5-nerviee 
vemlis utrinque reticulatis parum elevatis; stipulae jam 
delapsae; flores nobis ignoti; panicula fructifera ut videt- 
ur 15 om. longa ramis crassis patulis apice minutissime tom- 


108 PE: T:02L 0 Ge No. 2 


entellis; capsula ferruginea reniformis lateraliter paulo 
compressa 45 mm. leta 30 m. longa, pericarpio extra dense 
ferrugineo-tomentosa pilis apice glanduliferis, aculeis pate 
ulis rigidis 2--8 mm. longis infra medium ferrugineo-toment- 
osis dense obtecto; semina complanata obovato-oblonga 4.5 
mm. longa 3 mn. lata, praeter chalazam flavescentem castan- 
ea, raphe leviter costata, funiculo elongeto apice valde 
dilatato semen ad quartam partem tegente. 

Type, Krukoff 4960, collected June 21, 1934, in a high 
forest on terra firma near the mouth of the Rio Embira, 
Jurua basin, Amazonas. 

Bixe Orellana L., the common species of pantropic culti- 
vation, is so widely distributed and so variable in tropical 
America that ample justification must be sought for the de- 
- scription of a new species. In examination of numerous 
specimens of B. Orellana, the fruit is found to be flattened 
ovoid, distinctly tapering to the summit, with smooth peri- 
carp and thorns; the leaves vary from truncate to subcor date 
at base and are in most cases distinctly marked beneath with 
mimute pits which appear black under the lens. Variations in 
most of these points may be discovered, especially in the 
absence of pits, but no specimens have been seen with the 
least trace of glandular hairs on the pericarp or with such 
a broadly reniform capsule. Our species differs also in the 
strongly flattened seeds and costate raphe. 

Bixa urucurana Willd., considered by Pilger as merely a 
variety of B. Orellana and the same as B. sphaerocarpa Tri- 
ane and B. platycarpa R. & P., has leaves scaly beneath; its 
capsules approach ours in shape but are spherical. Glanduler 
pubescence is not mentioned in its description, while in our 
species it is so conspicuous that it could scarcely be over- 
looked, even in the most casual observation. 

In Bixa arborea Huber the capsules are described as ru- 
gose and minutely papillose under the lens, not aculeate, 
and 4-costate at base. The chalaza is white surrounded by a 
narrow zone of red and the body of the seed is blue. Its 
leaves are rounded at base and firm as in our species. 

Bixa Orellana is regularly a low spreading tree rarely 
more than § m. tall. Bixa arborea is said to reach a height 
of 15 m.$ apparently it is also stout and widely spreading, 
since its trunk is 20-30 cm. in diameter. Bixa excelsa is a 
slender tree with the typical shape of many other species of 
the rain forest, being only 24 om. in diameter near the base 
when neerly 350 meters tall. 


HENRIETTELLA SYLVESTRIS Gleason, sp. nov. Arbor mediocris 
15 m. alta; rami superiores graciles primm dense strigosi, 
pilie conicis subulatis, demm glabrescentes; petioli 1--2 
om. longi graciles subtus strigosi supra breviter hispidis 


y* Lab Sppeymereth 


1934 Gleason, Plantae Krukovianae - III 109 


laminae obovato-oblongae chartaceae, usque ad 14 cm. longae 
7-5 cm. latae, abrupte et obtuse acuminatae, integrae, basi 
acutae vel obtusae, 5-pli-nerviase, subtus ad venas venulas- 
que breviter strigosae, inter venas scabrae pilis erectis 
brevibus conicis, supra scabra, ad costam breviter hispidee 
et sub epidermide lineis brevibus crystallos lineares tegen- 
tibus dense notatae; flores 5-meri ad nodos defoliatos 2--5 
in fasciculis sessilibus, pedicellis gracillimis 5--10 mm. 
longis fere glabris; hypanthium parvum glabrum in fructo im- 
maturo 3.7 mn. longum; calycis tubus 0.2 mm. longum, lobi 
late rotundati, 1.7 mm. lati, a toro 0.5 mm. longi, dentibus 
exterioribus minutis verruciformibus; petala et stamina 
adhuc ignota. 

Type, Krukoff 5272, collected on terra firma at the mouth 
of the Rio Macauhan, Acre Territory. 

More than half of the South American species of the genus 
have sessile or subsessile flowers, and of the seven with 
pedicelled flowers all but one have smooth leaves. This one 
is H. tovarensis Cogn., a Venezuelan species with scabrous 
hypanthium and completely different pubescence. 

H. sylvestris seems to be common and widely distributed 
in western Amazonia. Specimens collected along the Amazon 
River by Williams (1717, 1736, 1864, 2091, 2972) were once 
identified by me as H. verrucosa Triana. It also extends up 
the Andes to at least 600 m. (Macbride 5493), 


BERNOULLIA SWIETENIOIDES Gleason, sp. nove Arbor excelsa 
55 m. alta, 1.35 m. in diametro; ramuli minores cinerei ten- 
uiter rugoso-costeti; folia unifoliolata; petioli graciles 
35-70 mm. longi, basi apiceque paullo incrassati et corruga- 
ti vel transversim incisi; laminae foliolorum firmulee late 
obovato-oblongae vel ellipticae, basi rotundatae, utrinque 
glabree, penninervieae, nervis seaundariis utroque latere 
circa 7 adscendentibus leviter arcuatis, nervulis temiter 
reticulatiss; inflorescentia et flores ignoti; pedunculus 
fructiferus 4 dm. longus ut videtur uniflorus; capsula fusi- 
formis fusca glabra, ad apicem obtusum angustata, basi obtu- 
sa, 20 cm. longa 7 om. in diam., ad maturitatem in valvas 5 
loculicide dehiscens, 5-loculeris; pericarpium crassum lig- 
nosum ab endocarpio papyraceo diremptum et dissepimenta pap- 
yracea ut in Swietenia; columella papyracea 5-alata ad apic- 
em pedunculi persistens; semina in quoque loculo in seriebus 
2 longitudinalibus disposita, in quoque seriei ca. 7, atro- 
castanea triangularia, ala suberosa 6 om. longa 15 mm. lata 
more Swieteniae ornata, quorum 4% basalia superne alata et 
adscendentia sunt et 4 apicalia alis pendentibus gaudent; 
cotyledones valde contorti et plicati. 

Type, Krukoff 5609, collected on terra firma at Foz do 
Macauhan, Purus River basin, Acre Territory. The tree has 


~ 


110 PHY. T-0 L OvG-25 No. 2 


very high buttress roots, according to Mr. Krukoff. 

Few, if any, of the plants recently brought back from 
Brazil by Mr. Krukoff have as great an interest to the taxo- 
nomist as the tree described above. The family to which it 
belongs remains in question. There is little doubt as to the 
genus to which it is most closely related and in which it is 
here placed, but a study of the flowers, as yet unknown, may 
indicate that it deserves the erection of a new gems. 

The genus Bernoullia was proposed by Oliver in 1876, 
based on a specimen from a tall tree in Guatemala. This 
plant, illustrated in Hooker's Icones, pl. 1169, 1170, was 
in bloom; the figure of the fruit was taken from a "careful 
drawing" sent by the collector. The accompanying notes ‘indi- 
cate that the fruit is almost indehiscent and the drawing 
& woody endocarp and dissepiments. The fruit of ours 
Suggests a wooay endocarp and dissrepiments. The fruit of 
ours is so extraordinsrily like that of the mahogany and so 
different from that of the Guatemalan plant that another 
gems is at once suggested. 

Bernoullia was described in the Sterculiaceae; Schumann, 
in the Naturlichen Pflanzenfamilien, classed it in the Bomb- 
acaceae. Bakhuizen keeps it in that family but suggests that 
it belongs in the Sterculiaceae instead. Dr. A. J. Panshin, 
of the New York State College of Forestry, suggests Stercul- 
iaceae on the evidence of its wood structure. 


DICLIDANTHERA OCTANDRA Gleason, sp. nov. Frutex 3 m. alt- 
us; rami superiores recti glabri subteretes; ramuli flori- 
feri valde divergentes primum tenuiter pubescentes angulati 
demam subglabri subteretes; petioli 5-8 mm. longi mox gla- 
bri; laminae firmlse elliptico-oblongae usque ad 14 cm. 
longee 6 cm. latae, superiores minores, integrae, basi api- 
ceque obtusae, glabrae (juniores supra ad costam minutissime 
puberulae), penninerviae, nervis adscendentibus ramosis ca. 
8 utroque latere, venulis arcte reticulatis utrinque promin- 
ulis; spicae ex axillis superioribus soliteariae adscendentes 
vel suberectae temiter pubescentes demum 7 cm. longae; flo- 
res sessiles inferne dissiti (in specimine nostro jam delap- 
si) superne conferti; sepala fere ad basin distincta, oblan- 
seolato-oblonga 6.5-7 mm. longa apice rotundata arcte pube- 
rula vel tomentella, ad anthesin paulo involuta; corollae 
tubus 11-12 mm. longus subcarnosus hine inde pilosulus fere 
cylindraceus superne paulo ampliatus, lobis 5 patulis vel 
subrecurvatis obovatis 4.5-4 mm. longis; stamina 8; fila- 
menta isomorphea ad faucem tubi inserta complanata carnosula 
1 mm. longa; antherae rotundatae basifixae introrsse 4-loc- 
ellatae, thecis 2 exterioribus majoribus, 3 ad letere unum 
corollae 0.8 mm. longae fertiles, 5 gradatim minores usque 


1934 Gleason, Plantae Krukovianse - III 112 


O.4 mm. longae et verisimiliter steriles vel substeriles; 
ovarium superum ovoideum 5-loculare, ovulo in quoque loculo 
1 axialis stylus cylindraceus villoeus 3 mm. longus (in ala- 
bastro; fortasse ad anthesin longior), stigmate capitato 0.9 


mm. diametro. 


Type, Krukoff 5778, collected in terra firma near the 
mouth of the Rio Macauhan, Acre Terrotory. 

The herbarium of the New York Botanical Garden contains 
type or authentic material of all three hitherto known spe- 
cies of Diclidanthera. D. octandra differs distinctly from 
all of them in habit and foliage and most notably in the 
presence of only 8 stamens. In ite general aspect and in 
all other structural features it ie clesrly a Diclidanthera 
Because of the limited material, it has not been 
practicable to determine whether the three large anthers are 
posterior or anterior. The two gaps in the circle of ten lie 
between the petals. 

Barnhartia Gl., referred by me originally, together with 


Diclidanthera, to the Styracacesae, has since been trans- 


ferred to the Polygalaceae. Barnhartia also has eight stam- 
ens, but the petals are separate and the ovary 2-celled. Our 
plant unites the staminal features of this genus with the 
gamopetalous corolla and 5-celled ovary of Diclidenthera and 
undoubtedly represents a connecting link between the two. 

Diclidanthera has been segregated from the Styracaceae 
for apparently good reasons. Bernhartia has been placed in 
the Polygalacesae after careful study by competent botanists. 
Its relationship to Diclidanthera is admitted. The discovery 
of such an intermediate type as D. octandra seems to justify 
the union of the two genera in the same family, whether it 
be Polygalacease or Diclidantheraceae. 


NOTE ON THE GENUS GOETHALSIA PITTIER 
H. A. Gleason 


Study of an herbarium specimen from Colombia, Lawrance 
494, led to the belief that it represented an undescribed 
genus of Flacourtiaceae. Because of the strong superficial 
resemblance of the plant to certain genera of Tiliaceae, 
search was also made for it in that family, where it was 
soon placed in the recently described gems Goethalsia of 
Pittier. Comparison with the type material at Washington, 
through the courtesy of the United States National Museum, 
verified the identification of the Lawrance plant with 
Goethalsia isthmica Pittier. 

Since it was not possible to reconcile my own belief that 
the gems is flacourtiaceous with Pittier's assignment of it 
to the Tiliacese, a more careful study of its structure was 
made and two interesting features unknown to Pittier were 
discovered. 

Pittier had before him young fruits and mature flowers. 
Lacking the younger buds, he never saw the sepals, which are 
deciduous. The organs described by him as sepals are the 
petals and those which he termed petals are the corone. 
Pittier alsc described the ovary as three-celled, while my 
microtome sections show beyond a doubt that it is one-celled 
with three deeply intruded placentae, each of which bears 
two rows of seeds. The adjacent seeds from two placentae 
protrude into the false loculus between them and give the 
appearance of two rows of axile ovules. 

Because of these facts, the genus is now transferred to 
the Flacourtiacese and an emended description is presented. 


Goethalsia Pitt. emend. Gl. 


Sepala 4 valvata ante anthesin decidua; petals 5 valvata 
lanceolato-attenuata extra argenteo-tomentosa; coronae seg- 
menta 5 libera hypogyna e medio ad basin cumeata, quam pet- 
alis mito brevioria, parte superiore erecta obtusa; stamina 
25 libera; ovarium trigonum l-loculare, placentis 3 alte in- 
veaginatis, ovulis paucis in quaque placenta biseriatis. 

The young buds show the sepals distinctly. They are el- 
liptic, 4 mm. long, and thinly stellate. When the bud reach- 
es 4 mm. in length, the sepals separate from their close 
valvate union and fall off, while the petals increase in 
length to 10 mm. Lewrance 494 was collected at 900 m. alti- 
tude in the western mountains of Boyacé, a region which has 
yielded many other new or interesting species. 


112 . 


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; 


PHY TOLOGIA 


Designed to expedite botanical publication 


; January, 1935 No. 3 
i " 
i CONTENTS 


ALC, Sure: Parag) Ry ukovr grag snd he osc: sscssescotnerenccdosconses soos cdvetbhedespeedptneee (22D 


y AC Suhre Studies of South American Plants-—IV sesssssssseesrvsseessesernee 126 


vh if 


138 


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a EP, KILuiP: A New ELanchOcar bus. sie Ue ee eae 
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: Some Necessary Nomenclatural Changes sicccsvecssseeccerccsseeeeee 141 


“Published by H. A. Gleason and Harold N. Moldenke 
Rae The New York Botanical Garden 
Me | _ Bronx Park, New York, N. Y. 
; ce of this liatusber’ 50 cents; per volume $5.00 in advance. 
Nin I, number 2, was issued ites 4, 1934. 
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PLANTAE KRUKOVIANAE - IV GARDEN 
A. CO. Smith 


The plants described below were collected ‘in Brazil by 
Mr. B. A. Krukoff. Types of new species are deposited in the 
herbarium of the New York Botanical Garden and are duplica- 
ted in several other American and European herbaria. 


LORANTHACEAE 


PSITTAOCANTHUS LEPTANTHUS A. C. Smith, sp. nov. Frutex 
parasiticus scandens glaber; ramis ramuilisque teretibus fus- 
cis, ad nodos continuis; petiolis rugosis 3-6 mm. longis, 
superne anguste alatis; laminis coriaceis supra viridis sub- 
tus fuscis oblongo-ovatis, 10-13 cm. longis, 4-8 cm. latis, 
basi apiceque acutis, margine integris et leviter revolutis, 
subtus dense stomatiferis, pinnatinerviis, costa utrinque 
prominente, nervis secundariis 4 vel 4 in quoque latere 
utrinque subplanis; inflorescentiis ut videtur solitariis in 
axillis foliorum, plerumque 4-floris, floribus binis; pedun- 
culis pedicellisque nigrescentibus gracilibus, bracteis mi- 
nutis; pedicellis 8-10 mm. longis; cupula patelliformi mar- 
gine integra; calyculo sub anthesi 3-4 mm. longo et 1.5 mm. 
diametro; perigonio tenuiter carnoso coccineo, mturitate 
7-8 cm. longo, ad mdium 3-4 mm. diametro, 6-lobato, lobis 
linearibue (ad 3 cm. longis) sub anthesi recurvatis, 1-1.5 
mm. latis, apice acutis, margine integris, ligula nulla; 
filamentis gracilibus circiter 10 mm. longis, supra medium 
perigonii adfixis; antheris dorsifixis oblongis 3-4 mm. lon- 
gis, apice obtusis; stylo perigonium aequante, stigmate 
ellipsoideo papilloso. 

Type, Krukoff 4709, collected June 10, 1933, on "varzea" 
land near mouth of Rio Embira ‘tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. It is allied to the 
group of species with united perigonium lobes, lacking basal 
ligules. From P. siphon Eichl., its nearest ally, the new 
species is distinguished by its broader leaves and simpler 
inflorescences, which are branched only twice. The pedicels 
of F. leptanthus are long and slender; the perigonium is . 
flaring from a very slender base. 


MYR ISTICACEAE 


DIALYANTHERA OLACOIDES A. C. Smith, sp. nov. Frutex; ra- 
mis glabris, ramlis teretibus, partibus juvenilibus seri- 
j= ceis mox glabris; petiolis subsericeis gracilibus canalicu- 
SC latis 5-8 mm. longis; laminis subcoriaceis elliptico-oblong- 
> is, 9-14 om. longis, 4.5-5 cm. latis, basi acutis, apice ob- 
tuse acuminatis, margine integris, utrinque glabris demim 


s 113 


22 193 


114 PHYTOLOGIA No. 3 


rugulosis, costa subtus prominente, nervis secunderiis 9-11l- 
jugis patulis prope marginem arcuatis conjunctis, supra 
planis vel impressis subtus elevatis, venulis inconspicuis; 
inflorescentiis ¢ axillearibus vulgo binis ad 8 mn. longis, 
ubique ferrugineo-sericeis; floribus 2-4 in fasciculis, 
bracteis minutis subteretibus; pedicellis 1-3 mm. longis; 
perigonio infundibuliformi 3 mm. longo, basi bracteola parva 
suffulto, intus glabro, lobis 3 ovatis acutis, 1.2 mm. long- 
is, 1.5 m. latis; filamentis in columnam glabram carnosam 
cylindricam 1.8 mm. longam connatis; antheris 3 oblongis 
liberis, 0.8-1 mm. longis, basi affixis, loculis 2 connec- 
tivo carnoso sustentis. 

‘Type, Krukoff 4740, collected June 10, 1933, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. Another collection 
from the same locality is Krukoff 4972. The extremely short 
inflorescences differentiate this species from all others of 
the genus. D. acuminata Standl., of Panama, approaches the 
new species in this respect, but is otherwise very differ- 
ent. The enlarged anther connective of D. olacoides is also 
noteworthy. 


MONIMIACEAE 


MOLLINEDIA KRUKOVII A. C. Smith, sp. nov. Frutex circiter 
4m. altus; ramie ramlisque teretibus striatis, juventute 
cinereo-pilosulosis mox glabris; petiolis oppositis subter- 
etibus canaliculatis 1.5-3.5 em. longis, velut ramlis no- 
vellis pilosulosis; laminis chartaceis ovatis, 20-33 em. 
longis, 11-17 cm. latis, basi acutis, apice acuminatis (acu- 
mine 1 cm. longo), margine integris et leviter revolutis, 
supra glabris, subtus sericeis (pilis viridibus basi 2-3- 
fidis circiter 0.2 mm. longis) demm glebrescentibus, copi- 
ose sed inconspicue pellucido-punctatis, costa supra eleveta 
subtus prominentissima, nervis secundariis 7-9-jugis arcua- 
to-adscendentibus utrinque elevatis, venulis copiose reticu- 
latis subtus prominulis; inflorescentiis & incompletis; 
pedicellis nigrescentibus 8-10 mm. longis, cinereo-pilosulo- 
sis; floribus & flavis extra velut pedicellis; receptaculis 
carnosis intus glabris; tepalis carnosis intus glabris, 2 
exterioribus late ovatis, 8-9 mm. longis, 9-10 mm. latis, 
mrgine integris et submembraneceis, 2 interioribus oblongo- 
orbicularibus, circiter 8 mm. longis et latis, mrgine mem- 
branaceis et irregulariter laciniatis; staminibus circiter 
50, filamentis glabris carnosis circiter 1.5 m. longis, 
antheris oblongis, 3 mm. longis, 1-1.5 mm. latis. 

Type, Krukoff 4819, collected June 14, 19343, on "terra 
firma" near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amezonas. It falls into the 
Section Appendiculatse Perk., being related to M. Selloi 


be 


1935 Smith, Plantae Krukovianae - IV 115 


(Spreng.) A. DO., M. latifolia (P. & E.) Tul., and M. Rusby- 


ana Perk. From these species it is readily distinguished by 


its large entire leaves, mich larger flowers, and more nu- 
merous stamens. 


LAURACEAE 


Mr. Krukoff's present collection is particularly rich in 
large forest trees of this family, of which about 45 species 
are represented. Of these, eleven are apparently new species. 


AIOUEA RUBRA A. C. Smith, sp. nov. Arbor ad 10 metralis, 
trunco 8 cm. diametro; ramulis striatis glabris rubellis; 
petiolis crassis profunde canaliculetis 8-15 mm. longis; 
laminis coriaceis glabris oblongis, 18-25 cm. longis, 4-6 
cm. latis, basi attenuatis, apice acuminatis (apice ipso 
obtuso), margine integris et leviter revolutis, costa rubel- 
la utrinque prominente, nervis secundariis utroque 5-7 ad- 
scendentibus supra prominulis subtus prominentibus, venulis 
reticuleatis supra immersis subtus prominulis; paniculis ru- 
bris glabris gracilibus mltifloris 10-20 cm. longis, 2-4 in 
axillis foliorum; floribus fulvis sub anthesi 2-3 mm. long- 
is; pedicellis gracilibus 6-8 mm. longis; perianthii lobis 
6 aequalibus ovatis membranaceis, minute luteo-glandulosis, 
0.8 mm. longis et laties; staminibus 6 fertilibus glabris, 

1 mm. longis, antheris bilocellatis truncatis filamentas ae- 
quantibus; seriei tertiae staminodiis filiformibus 0.6 mn. 
longis, basi glandulis 2 sessilibus 0.5 mm. diametro auctis; 
seriei quartae staminodiis carnosis ovatis cordatis sessili- 
bus 0.5 mm. diametro; ovario glabro sub anthesi 1 mm. longo, 
quam stylo paullo breviore, stigmate truncato. 

Type, Krukoff 4606, collected June 1, 1933, at Foz do 
Tarauaca, basin of Rio Jurua, State of Amazonas. It is rela- 
ted to A. scandens Ducke, but that species is a liana with 
long-petioled broad leaves. By its filiform staminodes (of 
the third series) and glabrous filaments, the flowers of the 
new species differ from those of the liana. From A. guyanen- 
sis Aubl., the new species differs by the texture of its 
leaves and its larger flowers, as well as by the shape of 
its staminodes. 


ANIBA ELLIPTICA A.C. Smith, sp. nov. Arbor ad 12 metra- 
lis, trunco 6 cm. diametro; ramis ramlisque subteretibus 
glabris; petiolis nigrescentibus canaliculatis 13-20 mm. 


- longis; laminis subcoriaceis glabris ellipticis, 15-20 cm. 
_longis, 7-10 cm. latis, basi cuneatis, apice obtusis, mar- 


gine integris, costa utrinque prominente, nervis secunderiis 
utroque 6-8 patulis prope margines anastomosantibus supra 

saepe planis subtus prominentibus, venulis copiose reticula- 
tis supra immersis subtus prominulis; inflorescentiis ut vi- 


116 PHYTOL OG 2s . No. 3 


detur solitariis e ramlis 5 mm. supra axillis foliorum ori- 
entis, 8-12 cm. longis, ramlis et floribus cinereo-tomen- 
tellis; pedicellis 2-2.5 mm. longis; floribus 4 mm. longis, 
2-2.5 mm. diametro; perianthii lobis 6 inaequalibus late 
ovatis subacutis intus glabris, exterioribus 1 mm. longis et 
1.5 mm. latis, interioribus 1.5 mm. longis et 1.8 mm. latis; 
staminibus 9 fertilibus 1.5 mm. longis, filamentis carnosis 
pilosis quam antheris duplo longioribus, antheris apice ob- 
tusis, seriei tertiae filamentis basi giandulis binis ses- 
silibus 0.3 mm. diametro auctis; gynaecio glabro 4 mm. 
longo, stylo ovarium aequante, stigmate truncato. 

Type, Krukoff 5601, collected Aug. 22, 1933, on "terra 
firma" near mouth of Rio ‘acauhan (tributary of Rio Yaco), 
basin of Rio Purus, Territory of Acre. In having unequal 
perianth segments, the new species resembles A. Brittonii 
. Mez, from which it differs by its longer petioles and dif- 
ferently shaped larger leaves. 


ANIBA JURUENSIS A. C. Smith, sp. nov. Arbor ad 25 metra- 
lis, trunco circiter 18 cm. diametro; ramis ramlisque ni- 
grescentibus striatis, juventute cinereo-puberulis demum 
glabris; petiolis gracilibus 6-12 mm. longis puberulis; la- 
minis chartaceis oblongis, 14-20 cm. longis, 4-7 cm. latis, 
basi cuneatis, apice longe acuminatis (acumine 1.5-2 cm. 
longo), margine integris et leviter revolutis, supra gle- 
bris nitidis, subtus fuscis arcte cinereo-puberulis, costa 
supra saepe impressa subtus prominentissima, nervis secun- 
dariis utroque 3-5 adscendentibus supra prominulis subtus 
prominentibus, venulis reticulatis utringque prominulis; in- 
florescentiis ad apices ramorum in axillis foliorum par- 
vorum congestis, 4-10 cm. longis, 20-50-floris, ramulis et 
floribus cinereo-puberulis; floribus in cymulis parvis 
bracteolis 2 mm. longis mox deciduis subtentis; pedicellis 
ad 3 mm. longis; perianthii tubo 1.5 mm. longo, lobis 6 ae- 
- qualibus oblongis, 1.2 mm. longis, O.7 mm. latis, apice ro- 

tundatis, utrinque minute puberulis, margine leviter incras- 
satis; staminibus 9 fertilibus, serierum exteriorum 0.4 mm. 
longis et latis, antheris subsessilibus, apice emarginatis, 
seriei tertiae filamentis brevissimis, glandulis binis ses- 
silibus auctis, antheris minutis; ovario glabro sub anthesi 
1.5 mn. longo, stylo subnullo, stigmate discoideo, 0.5 mm. 
diametro; bacca juvenili in cupula nigrescente obovoidea 
immersa. 

Type, xrukoff 4775, collected in June, 1933, near mouth 
of Rio Embira (tributary of Rio Tarauaca), basin of Rio 
Jurue, State of Amazonas. Krukoff 4932, from the same local- 
ity, also represents the species; it is froma juvenile 
plant noted as a shrub 5 meters high, with slightly broader 
leaves, which ere silvery-sericeous beneath, and shorter in- 


1935 Smith, Plantae Krukovianae - IV 1i7 


florescences than the type. The species is related to A. 
Muelleriana Mez, from which it differs by the shape and pu- 
berulence of its leaves, its minute subsessile anthers, and 
its discoid stigma. 


ANIBA KRUKOVII A. C. Smith, sp. nov. Frutex 5 m. altus; 
ramulis fuscis glabris striatis; petiolis rugosis semiter- 
etibus glabris 10-16 m. longis; laminis chartaceis glabris 
elliptico-oblongis, 13-20 cm. longis, 5-7 cm. latis, basi 
acutis, apice caudato-acuminatis, margine integris, nervis 
secundsariis utroque 2 vel 3 adscendentibus, cum costa supra 
elevatis subtus prominentibus, venulis reticulatis utr inque 
prominulis; inflorescentiis 1-3 in axillis foliorum, 2-3 cm. 
longis, 15-25-floris, ramlis et floribus cinereo-sericeis; 
floribus 4 mm. longis in cymlis parvis subsessilibus; per- 
ianthii lobis 6 aequalibus oblongis, 1.8 mm. longis, 1.4 mn. 
latis, apice rotundatis, utrinque dense luteo-glandulosis; 
staminibus 9 fertilibus 1 mm. longis, serierum exteriorum 
antherarum connectivis subacutis, filamentis pilosis quam 
antheris paullo brevioribus, seriei tertiae filamentis dila- 
tatis sericeis, glandulis binis sessilibus 0.4 mm. diametro 
auctis; ovario glabro sub anthesi 1.3 mm. longo, stylo bre- 
viore, stigmate membranaceo irregulariter discoideo. 

Type, Krukoff 5023, collected June 26, 193%, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. It is related to A. 


Muelleriana Mez and the above described new species, from 


both of which its few ascending lateral nerves distinguish 
it. The pale sericeous pubescence of the short inflorescence 
and the glandular perianth segments also characterize the 
new species. 


ANIBA FLEXUOSA A. C. Smith, sp. nov. Frutex glaber; ramis 
ramulisque teretibus olivaceis; petiolis angulatis 15-25 mm. 
longis; laminis chartaceis viridis elliptico-oblongis, 20-30 
em. longis, 10-15 cm. latis, basi cuneatis vel subrotunda- 
tis, apice caudato-acuminatis (apice ipso 1-2 cm. longo ob- 
tuso), margine integris, costa utrinque prominente, nervis 
secundariis utroque 8-11 patulis prope margines adscendenti- 
bus anastomosantibus utrinque elevatis, venulis copiose re- 
ticulatis utrinque prominulis; inflorescentiis ut videtur 
solitariis axillaribus ad 15 cm. longis, 15-20-floris, rha- 
chide flexiosa, ramlis secundariis rectis 12-20 mm. longis, 
pedunculis 5-7 mm. longis, pedicellis brevissimis; floribus 
3 mm. longis, tubo obconico quam lobis longioribus; perian- 
thii lobis aequalibus oblongis subacutis, 1.3 mm. longis, 1 
mm. latis; staminibus 9 fertilibus; serierum exteriorum ob- 
longis 0.5 mm. longis et latis, basi parce pilosis, antheris 
subsessilibus, connectivis productis obtusis; seriei tertiae 


118 PHYTOLOGIA | No. 3 


deltoideis, 0.8 mm. longis et latis, antheris minutis, con- 
nectivis productis obtusis, filamentis dilatatis sericeis 
quam antheris duplo longioribus, basi glandulis binis ses- 
silibus 0.3 mm. diametro auctis; ovario glabro sub anthesi 
1.5 mm. longo, stylo subnullo, stigmate discoideo 0.8 mm. 
diametro. 

Type, Krukoff 5030, collected June 21, 1933, on "terra 
firma" near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. From A. citrifolia 
(Nees) Mez, probably its closest relative, the new species 
differs by ites much larger leaves, its lax few-flowered gla- 
brous inflorescence, and its small subsessile outer anthers. 


ENDLICHERIA FORMOSA A.C. Smith, sp. nov. Arbor ad 10 me- 
tralis, trunco 8 cm. diametro; ramlis teretibus lenticel- 
latis glabris vel juventute parce pilosis; petiolis rugosis 
15-25 mm. longis; laminis chartaceis utrinque minute puberu- 
lis mox glabris obovato-ellipticis, 15-22 cm. longis, 5-9 
cm. latis, apice breviter acuminatis, margine integris, ner- 
vis secundeariis utroque 9-11 cum costa supra elevatis subtus 
prominentibus, venulis copiose reticulatis utrinque peracute 
prominulis; inflorescentiis axillaribus solitariis milti- 
floris 7-10 cm. longis, ramulis minute tomentellis, pedicel- 
lis 2 mm. longis basin versus minute bibracteolatis; flori- 
bus o albis subglobosis 1.5 mm. diametro, tubo quam lobis 
minutis majore; perianthii lobis late deltoideis, 0.5 mm. 
longis, 0.8 mm. latis, obscure pellucido-punctatis; stamini- 
bus 9 fertilibus ovatis, 0.6 mm. longis et latis, antheris 
per poros sublaterales dehiscentibus, connectivis productis 
subacutis, filamentis contractis quam antheris brevioribus, 
seriei tertiae basi biglandulosis; gynaecio nullo vel minuto. 

Type, Krukoff 5156, collected July 4, 1933, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauace), 
basin of Rio Jurua, State of Amazonas. In foliage, the new 
species closely resembles E. glaberrima Mez, but has many- 
flowered inflorescences with differently-shaped smaller 
flowers. The perianth of the new species is sharply truncate 
at base, with minute lobes. 


PERSEA INAEQUALIS A. C. Smith, sp. nov. Arbor 30 m. alta, 
trunco 30 cm. diametro; ramis ramlisque subteretibus fuscis 
juventute cinereo-puberulis; petiolis suboppositis mox gla- 
bris canaliculatis 10-15 mm. longie; laminis coriaceis ob- 
longis, 12-16 cm. longis, 5-7 cm. latis, basi cuneatis, 
apice obtusis vel rotundatis, margine integris, supra fuscis 
glabris, subtus glaucescentibus mimutissime pulverulentis 
mox glabris, costa supra leviter impressa subtus prominent- 
issima, nervis secundariis utroque 8-11 rectis adscendenti- 
bus supra planis subtus prominentibus, venulis reticulatis 


— - . ea 


1945 Smith, Plantae Krukovianae - IV 119 
utringque planis vel prominulis; inflorescentiis 1-4 axillar- 
ibus 2-4 cm. longis et 10-20-floris, ramulis minutissime pu- 
berulis; pedicellis circiter 1.5 mm. longis 2- vel 3-fascic- 
ulatis, bracteolis ad 0.8 mm. longis subtentis; perianthii 
tubo subnullo, lobis albis 6 inaequalibus minutissim fer- 
rugineo-tomentellis, 3 exterioribus deltoideis, 0.5 mm. 
longis et latis, 3% interioribus oblongis obtusis, 2.5 mm. 
longis, 1.8 mm. latis; staminibus 9 fertilibus; serierum ex- 
teriorum antheris bilocellatis suborbicularibus, 0.6 mm. 
diametro, apice obtuse apiculatis, introrso-lateraliter de- 
hiscentibus, filamentis gracilibus quam antheris breviori- 
bus; seriei tertiae antheris similibus sed extrorso-lateral- 
iter dehiscentibus, filamentis basi biglandulosis antheras 
aequantibus; staminodiis 3 lineare-spathulatis acutis 0.8 
mm. longis; ovario parce piloso sub anthesi 0.8 mm. diame- 
tro, stylo breve, stigmate truncato. 

Type, Krukoff 4770, collected June 12, 1933, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauacea), 
basin of Rio Jurua, State of Amazonas. It is a species of 
the subgenus Hemipersea Mez, having bilocellate anthers in 
common with P. cuneata Meisen. and P. durifolia Mez. On this 
character the three species would appear to belong to the 
genus Hufelandia, but on other characters, notably the un- 
equal perianth lobes, they are best placed in Persea. The 
new species differs from P. durifolia, its nearest ally, by 
its larger leaves with more numerous secondary nerves, its 
short-pedunculate inflorescences, its shorter filaments and 
ite differently shaped anthers. A few flowers on the type 
specimen have 8 perianth lobes and a correspondingly in- 
creased number of stamens. 


OCOTEA TENELLA A. C. Smith, sp. nov. Arbor 13 m. alta, 
trunco 7-10 cm. diametro; ramlis teretibus dense et arcte 
cinereo-tomentellis demum glabrescentibus; petiolis grecili- 
bus tomentellis 4-7 mm. longis; laminis chartaceis ellipti- 
co-oblongis, 6-9 cm. longis, 3-5 cm. latis, basi acutis, 


epice breviter obtuse acuminatis, margine integris, supra 


glabris et impresso-punctatis, subtus molliter et arcte cin- 
ereo-tomentellis, nervis secundariis utroque 3 vel 4 arcus- 
to-adscendentibus, cum costa supra subplanis subtus eleva- 
tis, venulis reticulatis subtus prominulis; inflorescentiis 
solitariis axillaribus gracilibus 2-4 cm. longis et 10-20- 
floris, rhachide parce tomentella, ramlis glabris 1-2 mm. 
longis flores 2 vel 3 ad apicem gerentibus; pedicellis ad 1 
™m. longis; floribue glabris nigrescentibus mturitate 1.5- 
2 mm. longis, perianthii tubo breviter obconico, lobis ob- 
longis obtusis, 0.8 mm. longis, 0.5 mm. latis; staminibus 9 
fertilibus; serierum exteriorum antheris. sessilibus oblongo- 
orbicularibus, circiter 0.4 mm. longis et latis, apice ov- 


120 PHYTOLOGIA No. 3 


tusis vel minute apiculatis; seriei tertiae similibus sed 
cum filamentis brevissimis biglandulosis; gynaecio glabro, 
ovario sub anthesi 0.8 mm. longo, stylo brevissimo, stigmate 
minute discoideo. 

Type, Krukoff 5406, collected Aug. 11, 1933, on "terra 
firma" near mouth of Rio Macauhan (tributary of Rio Yaco), 
basin of Rio Purus, Territory of Acre. It is a species of 
the Section Mespilodaphne, without close relatives, perhaps 
closest to O. lanata (Nees) Mez, from which it differs by 
its minute flowers, slender inflorescences, leaf shape, type 
of pubescence, etc. 


NECTANDRA SUPERBA A. C. Smith, sp. nov. Arbor 37 m. alta, 
trunco 7 dm. diametro; ramlis subteretibus vel leviter an- 
gulatis juventute dense et arcte fusco-tomentosis; petiolis 
rugosis tomentosis 15-25 mm. longis; laminis coriaceis sic- 
citate olivaceis elliptico-oblongis, 15-20 cm. longis, 5-8 
em. latis, utrinque acutis, margine integris, supra praeter 
costam glabris nitidis, subtus dense et arcte adpresso-pilo- 
sis, nervis secundariis utroque 5-7 adscendentibus cum costa 
supra plus minusve planis subtus prominentibus, venulis 
copiosissime reticulatis utringque prominulis; inflorescen- 
tiis ad apices ramlorum axillaribus, 5-12 cm. longis, ad 
30-floris, ubique arcte tomentosis, ramulis lateralibus 4-10 
mm. longis; floribus subsessilibus (pedicellis ad 1 mm. 
longis) 2-4 in fasciculis, bracteolis minutis deciduis sub- 
tentis; perianthii tubo obconico 1.5-2 mm. longo, lobis 6 
ovatis acutis, 1.5-2 mm. longis, 1-1.5 mm. latis; staminibus 
9 fertilibus; serierum exteriorum antheris subsessilibus 
suborbiculeribus apice obtusis, 0.6-0.7 mm. diametro, dor- 
saliter glendulas 2-4 minutas nigras gerentibus, loculis in 
serie unica recta juxtapositis; seriei tertiae staminibus 
oblongis 0.8 mm. longis, antheris rotundatis glandulosis, 
filamentis carnosis antheras aequantibus glandulis binis 
sessilibus auctis; gynaecio glabro; ovario 1 mm. diametro, 
stylo 0.5 mm. longo, stigmate truncato.— 

Type, Krukoff 5757, collected Sept. 3, 1933, on "terra 
firma" near mouth of Rio Macauhan (tributary of Rio Yaco), 
basin of Rio Purus, Territory of Acre. It is probably most 
closely allied to N. lineatifolia (R. & P.) Mez, from which 
it differs by the prominently reticulate veinlets and the 
subsessile flowers with blunt anthers. 


PLEUROTHYRIUM NOBILE A. 0. Smith, sp. nov. Arbor 25 m. 
alta, trunco 20 cm. diametro; ramlis teretibus crassis ru- 
gosis glabris; petiolis crassis fuscis canaliculatis 2-4 
cm. longis; laminis coriaceis glabris oblongis, 20-30 cm. 
longis, 9-13 cm. latis, basi obtusis vel rotundatis, apice 
cuspidatis vel breviter acuminatis, margine integris, costa 


1945 Smith, Plantae Krukovianae - IV 121 


utrinque prominente, nervis secundariis utroque 9-12 arcua- 
to-adscendentibus supra planis vel leviter elevatis subtus 
prominentibus, venulis reticulatis supra planis subtus pro- 
minulis; inflorescentiis solitariis axillaribus quam foliis 
paullo brevioribus multifloris, ramlis angulatis glabris 
vel minute ferrugineo-tomentellis, ramlis secundariis 6-9 
rectis 3-4 cm. longis; pedicellis 6-10 mm. longis cum flor- 
ibus velut inflorescentiae ramulis tomentellis; perianthii 
tubo obconico 4 mm. longo et diametro, lobis 6 carnosis ob- 
longis subacutis, 5-6 mm. longis, 4-5 mm. letis; staminibus 
9 fertilibus oblongis truncatis ferrugineo-tomentellis 1.4 
mm. longis, locellis 2 introrsis 2 extrorsis vel omnibus 
lateralibus, filamentis antheras aequantibus contractis, 
omnibus manifestissime glandulis binis maximis confluenti- 
bus cinctis; ovario glabro vel minutissime tomentello glo- 
boso sub anthesi 2 mm. diametro, stylo 0.5 mm. longo, stig- 
mate discoideo 0.4 mm. diametro; fructibus juvenilibus glo- 
bosis apice perianthii lobos gerentibus. 

Type, Krukoff 5121, collected July 1, 1933, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauacea), 
basin of Rio Jurua, State of Amazonas. Another specimen 
from the same locality is Krukoff 4800, which is more or 
less juvenile-and bears shorter inflorescences. It is a 
species related to P. bifidum Nees, from which it differs 
by its lack of inflorescence tomentum and its large long- 
pedicelled flowers. From P. Poeppigii Nees it differs by 
its glabrous leaves and less robust, more spreading inflo- 
rescences. P. parviflorum Ducke, another related species, 
has very mch smaller flowers than P. nobile. 


PLEUROTHYRIUM KRUKOVII A. C. Smith, sp. nov. Arbor ad 28 
m. alta; remlis teretibus fistulosis juventute cinereo- 
tomentellis mox glabris; petiolis rugosis nigrescentibus 
canaliculatis 15-30 mm. longis; laminis coriaceis glabris 
oblongis, 25-45 cm. longis, 7-12 cm. latis, basi acutis, 
apice acutis vel breviter acuminatis, margine integris, 
costa supra elevata subtus prominentissima, nervis secund- 
ariis utroque 14-16 rectis supre planis subtus prominenti- 
bus, venulis reticulatis supra obscuris subtus prominulis; 
inflorescentiis solitariis axillearibus quam foliis mlto 
brevioribus (7-12 cm. longis) miltifloris, ramlis (et 
bracteolis floribusque) minute cinereo-tomentellis, ramlis 
secundariis 10-16 gracilibus complanatis 4-10 mm. longis; 
floribus 3-6 in fasciculis, bracteolis 2 vel 4 spathulatis 
acutis 3-4 mm. longis mox deciduis subtentis; pedicellis 
maturitate 3-4 mm. longis; perianthii tubo obconico 1 m. 
longo, lobis 6 oblongie obtusis, 2.5 mm. longis, 1.5 mm. 
latis; staminibus 9 fertilibus 1 mm. longis, antheris ob- 
longis apice emarginatis, locellis plus minusve laterali- 


122 PHYTOLOGIA No. 3 


bus, filamentis gracilibus antheras saequantibus, omnibus 
glandulis binis non confluentibus cinctis; ovario glabro 
globoso sub anthesi 1 mm. diametro, stylo arcte tomentello 
ovarium aequante, stigmate truncato. 

Type, Krukoff 5563, collected Aug. 17, 1933, on "terra 
firma" near mouth of Rio Macauhan (tributary of Rio Yaco), 
basin of Rio Purus, Territory of Acre. Other collections 
from the same locality are Krukoff 5255 and 5722. It is re- 
lated to P. parviflorum Ducke, from which it differs by the 
mich larger leaves, the narrower inflorescences (of which 
P. parviflurum has secondary branches 12-15 mm. long), and 
the more numerous flowers. 


ELAEOCARPAOCEAE 


Sloanea reticulata A. C. Smith - Territory of Acre: up- 
per Rio Jurapary, basin of Rio Jurua, Krukoff 5230. This 
specimen agrees with the Maranhad plant described as S. re- 
ticulata in all details except size of inflorescence. The 
peduncles of the Acre plant are 4-6 cm. long, the pedicels 
2-3 cm. long. Other floral and leaf characters mentioned in 
the description:of $8, reticulata distinguish the species 
from S. Garokeana Schum. 


GUTTIFERAE 


TOVOMITA KRUKOVII A. 0. Smith, sp. nov. Frutex glaber 8 
m. altue; ramlis teretibus cinereis; petiolis gracilibus 
canaliculatis 7-20 mm. longis; laminis subcorieceis oblong- 
is, 7-10 cm. longis, 2.5-4 cm. latis, basi attermatis, 
apice acutis vel breviter acuninatis, margine integris et 
leviter recurvatis, costa utrinque prominente, nervis se- 
cundariis utroque 7-10 (nervis tertisriis interspersis) 
patulis cum venulis utrinque promimulis; inflorescentiis 
axillaribus pauciramosis compsctis 1-1.5 cm. longis, ram- 
lis gracilibus brevissimis bracteis 1-2 mm. longis subten- 
tis; pedicellis 5-5 mm. longis, alabastris $3 mm. diame- 
tro; sepalis 4 tenuiter coriaceis ovato-oblongis, 4 m. 
longis, 4 mm. latis, 2 interioribus angustioribus; petalis 
4 oblongis obtusis papyraceis, 4 mm. longis, 1.5 mm. latis; 
staminibus cireiter 30, 3-4 mm. longis, filamentis cerno- 
sis, antheris globosis obtusis 0.3 mm. longis. 

Type, Krukoff 4887, collected in June, 1933, on “terra 
firma" near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. From its ally, T. 
Riedeliara Engl., the new especies differs by its compact 
rather than ample terminal inflorescence, and by its smaller 
flowers. T. acuminata Engl., another related species, has 
broader conspicuously acuminate leaves and a more ample in- 
florescence than T. Krukovii. 


| 


19435 Smith, Plantee Krukovianae - IV 123 


TOVOMITA MICRANTHA A. C. Smith, sp. nov. Frutex glaber 5 
m. altus; ramlis teretibus fuscis; petiolis crassis leviter 
canaliculatis 8-16 mm. longis; laminis coriaceis oblongo- 
ellipticis, 12-20 cm. longis, 6-12 em. latis, basi acutis, 
apice acutis vel breviter acuminatis, margine integris, 
costa utrinque prominente, nervis secundariis utroque 10-15 
patulis curvatis supra leviter impressis suhtus elevatis, 
venulis reticulatis subtus saepe prominulis; inflorescentiis 
axillaribus pauciramosis 2-3 cm. ante? ramlis crassis, 
bracteis parvis caducis; pedicellis 4-5 mm. longis, alabas- 
tris & albis subglobosis 3-4 mm. diametro; sepalis 4 cori- 
aceis ovato-oblongis subacutis, 3-4 mm. longis, 2-3 mm. la- 
tis; petalis 4 (semper?) velut sepalis sed papyraceis mox 
deciduis; staminibus 25-30 circiter 43.5 mm. longis, fila- 
mentis carnosis, antheris acutis 0.5 mm. longis. 

Type, Krukoff 5071, collected June 28, 1933, on "terra 
firma" near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. It is a species of 
the alliance of T. rubella Spruce and T. Spruceana Pl. & ~ 
Tr., having broader leaves than either. In floral characters 
it more nearly resembles the former, which, however, has 
fewer lateral nerves. The new species resembles T. macro- 
phylla (P. & E.) Walp. in foliage, but has a far less ro- 
bust inflorescence and fewer petals. 


LEOYTHIDACEAE 


LECYTHIS HIANS A. C. Smith, sp. nov. Arbor glabra ad 30 
m. alta; ramlis teretibus fuscis dense lenticellatis; pet- 
iolis gracilibus nigrescentibus 3-6 cm. longis anguste ala- 
tis; laminis chartaceis fuscescentibus oblongis, 7-10 cm. 


longis, 3-4 cm. latis, basi acutis vel obtusis, apice brevi- 


ter acuminatis, margine serrulatis, costa utrinque conspi- 
cua, nervis secundariis utroque 9-12 rectis prope margines 
anastomdsantibus, cum venulis reticulatis utrinque prominu- 
lis; paniculis axillaribus et terminalibus pauciramosis 5-12 
em. longis; rhachidibus rugosis conspicue lenticellatis, 
geniculis incrassatis; pedicellis subnullis vel ad 2 mm. 
longis; sepalis subaequalibus carnosis deltoideo-ovatis, 4 
mn. longis, 4 mm. latis, apice rotundatis, margine membrana- 
ceis; petalis rotundato-oblongis, 15-17 mm. longis, 12-14 
mm. latis; androphoro carnoso explanato 25-35 mm. longo, 
ligula prope basin 15-23 mm. lata distaliter contracta, 
galea 15-25 mm. diametro subtus dense echinata, appendiculis 
anantheris linearibus 6 mm. longis (prope margines plerum- 
que staminiferis) obtecta, staminibus circa annulum numero-— 
sissimis, filamentis carnosis 0.8 mm. longis distaliter in- 
crassatis, antheris subglobosis 0.4 mm. diametro; ovario 


- ‘subinfero, vertice plano, stylo carnoso 1 mm. longo, loculis 


4, ovulis in quoque loculo 10-15 funiculis sustentis; pyx- 


124 PHYTOLOGIA No. 3 


idio vestuto subgloboso rugoso, 14 cm. longo (sine opercu- 
lo), 16 om. lato, supra medium zona calycari rugosa cincto; 
vitta interzonali suberecta 3-4 cm. longa; zona superiore 
integra, ore 11-13 cm. diametro; pericarpio tenui, 6-8 mm. 
crasso, 

Type, Krukoff 4811, collected June 14, 1933, on "terra 
firma" near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Asdisae State of Amazonas. Another specimen, 

a ailmate on “varzea" land in the same vicinity, is Krukoff. 
4630. Both specimens bear flowers, which are said to be Oe 
white and violet," and 4811 is accompanied by an old fruit, 
lacking operculum and seeds. It is apparently most closely 
related to L. usitata Miers, like which it is called "Sapu- 
caya." The new species has uniformly smaller leaves and 
flowers; its fruit has a much thinner pericarp (in thickness 
6-8 mm. instead of 11-18 mm. as in L. usitata) and is only 
slightly contracted at the mouth (diameter of mouth 11-13 
em. rather than 6-10 cm. as in L. usitata). L. paraensis 
Huber, a species not properly published, is not known to m. 


— APICULATA (Miers) A. C. Smith, comb. nov. 
Chytroma apiculate Miers, Trans. Linn. Soc. 430: 245. 1874. 
Maranhad: Maracassumé River region, Froes 1868. Our speci- 
men is similar to type mterial; a local name is "Atiriba." 


ESCHWEILERA RORIDA (Miers) A. C. Smith, comb. nov. Chy- 
troma rorida Miers, Trans. Linn. Soc. 30: 243. 1874. 


ESCHWEILERA (Eueschweilera) KRUKOVII A. C. Smith, sp. 
nov. Arbor glabra 25-35 m. alta, trunco circiter 1.5 m. 
diametro; ramis ramlisque cinereis teretibus lenticellatis; 
petiolis rugosis nigrescentibus 6-11 mm. longis; laminis 
coriaceis oblongis, 8-16 cm. longis, 2.5-6 cm. latis, basi 
acutis vel subtruncatis, apice breviter et obtuse acumina- 
tis, margine integris vel obsolete serrulatis, costa crassa 
utrinque subprominente, nervis secundariis utroque 6-8 prope 
margines adscendentibus utrinque prominulis vel supra immer- 
sis, venulis copiose reticulatis subtus prominulis; panicu- 
lis axillaribus pauciramosis 2-8 cm. longis; rhachidibus ju- 
ventute fusco-puberulis mox glabris; pedicellis rugosis 5-10 
mm. longis; sepalis 6 subaequalibus ovatis obtusis, 2 mm. 
longis, 2-3 mm. latis, margine membranaceis; petalis tenui- 
ter carnosis oblongo-ovatis, 16-18 mm. longis, 12-14 mm. le- 
tis, apice rotundatis; androphoro explanato 30 mm. longo, 
ligula tenuiter coriacea 8-10 mm. lata, galea 12-14 mm. di- 
ametro subtus dense echinata appendiculis anantheris linear- 
ibus 4-5 mm. longis obtecta; staminibus circiter 150 circa 
annulum et ligulae basi, filamentie carnosis 1-1.5 mm. long- 
is, antheris subglobosis 0.5 mm. diametro; ovario subinfero, 


1935 Smith, Plantae Krukovianae - IV A125 


vertice plano, stylo carnoso subconico 2 mm. longo prope 
basin 1 mm. diametro, stigmate truncato, loculis 2, ovulis 
in quoque loculo 4 vel 5 e basi sessilibus. 

Type, Krukoff 4847, collected June 15, 1933, on "varzea" 
land near mouth of Rio Embira (tributary of Rio Tarauaca), 
basin of Rio Jurua, State of Amazonas. Another collection 
from the same locality is Krukoff 4739. It is a species re- 
sembling E. rorida (Miers) A. C. Smith, from which it dif- 
fers by its 2- rather than 4-celled ovary, its pedicelled 
rather than subsessile flowers, and its narrower leaves. The 
number of cells in the ovary (the distinguishing feature of 
the Sections Eveschweilera and Chytroma) is considered of 
secondary importance by Eyma (a), since by this means 
closely related species have been too far separated. 


ESCHWEILERA OBTECTA (Miers) A. C. Smith, comb. nov. 
Jugastrum obtectum Miers, Trans. Linn. Soc. 30: 276. 1874. 
Krukoff 1293 (Madeira River region, State of Amazonas) has 
a fruit resembling that portrayed by Miers, although I have 
not compared the foliage with type material. 


Gouratari macrosperma A. C. Smith, formerly kmown from 
the upper Machado River region, State of Matto Grosso, is 
also represented by Krukoff 5638 and 5687, both collected 
near the mouth of the Rio Macauhan (tributary of Rio Yaco), 
Territory of Acre. Both specimens are taken from trees near- 
ly 60 meters high. 5638 bears fruits similar to those of the 
type, but with seeds slightly different in proportion (about 
10 by 3 cm.). 5687 bears inflorescences, which are here de- 
scribed: 

Racemis axillaribus et terminalibus, 10-20 cm. longis, ut 
videtur simplicibus; rhachide crassa arcte fusco-stellato- 
pubescente 15-20-flora; pedicellis crassis dense pubescenti- 
bus, 5-8 mm. longis, infra medium articulatis et minute 5- 
bracteolatis; sepalis 6 imbricatis oblongo-rotundatis ob- 
tusis, ad 8 mm. longis et latis, extra pubescentibus, mar- 
gine breviter ciliatis; petalis 6 albis obovatis extra pu- 
berulis, circiter 20 mm. longis et latis; androphoro explan- 
ato 40 mm. longo, ligula 10 mm. lata, galea 15-20 mm. diam- 


. etro subtus densissime echinata; staminibus circa annulum 


numerosis, filamentis gracilibus 1 mm. longis, antheris ob- 
longis 1 mn. longis; ovario turbinato pubescente subinfero, 
vertice plano, stylo crasso subtruncato. 


Couratari pulchra Sandwith - Amazonas: near mouth of Rio 
Embira (tributary of Rio Tarauaca), basin of Rio Jurua, 
Krukoff 5086. Our specimen agrees precisely in vegetative 
characters with the type collection of the Guiana species, 
and I note this extension of range with som confidence. The 


126 PAY TO) 667-ios No. 3 


fruits of 5086 agree with those described from Surinam by 
Eyma (b). 


Cariniana domestica (Mart.) Miers appears to be repre- 
sented by Krukoff 5597, collected near mouth of Rio Macau- 
han (tributary of Rio Yaco), basin of Rio Purus, Territory 
of Acre. Our plant, in foliage and fruit, resembles the de- 
scriptions and plates. Mr. Krukoff notes it as a buttressed 
tree nearly 60 meters high. 


Cariniana micrantha Ducke - Amazonas: near mouth of Rio 
Embira (tributary of Rio Tarauaca), basin of Rio Jurua, 
Krukoff 5095. A strongly buttressed tree nearly 60 meters 
high, equalled in the Jurua region only by & species of 
Swietenia. The wood of the two species is occasionally sold 
in mixture. 


(a) Meded. Bot. Mus. en Herb. Rijks Univ. Utrecht 4: 59. 


1932 * 
Cho 2.6. Sz. 


STUDIES OF SOUTH AMERICAN PLANTS - IV 
New Monimiaceae, Trigoniaceae, and Vacciniaceae 
A. C. Smith 


In the accompanying paper specimens from several herbaria 
are cited. These institutions are indicated by the following 
abbreviationst Arnold Arboretum (A); British Museum (BM); 
Grey Herbarium (G); Royal Botanic Gardens, Kew (K); Jardin 
Botanico, Madrid (M); U. $3. National Museum (N); New York 
Botanical Garden (Y). 


MONIMIAOEAE 


SIPARUNA HEMISPHAERICA A. C. Smith, sp. nov. Frutex dio- 
ecus (vel arbor parva?); ramlis elongatis subteretibus fus- 
cis glabris; petiolis oppositis vel suboppositis semi-teret- 
ibus rugosis nigrescentibus glabris 7-20 mm. longis; laminis 
fuscescentibus oblongis, 13-19 cm. longis, 4-5.5 cm. latie, 
basi acutis, apice subacuminatis, margine subintegris et an- 
guste revolutis, utrinque glabris, pinnatinerviis, costa su- 
pra plana subtus prominente, nervis lateralibus 8-11 in quo- 
que latere arcuato-adscendentibus prope margines anastomo- 
santibus, utrinque leviter elevatis, venulis reticulatis 
leviter elevatis; inflorescentiis o axillaribus plerumque 


st eis 


19435 Smith, Studies of South American Plants - IV 127 


binis 2-3.5 cm. longis 6-12-floris; pedunculo 1.5-2.5 cm. 
longo gracillimo nigrescente glabro (vel minute puberulo); 
floribus nigrescentibus ubique glabris (juventute parce pu- 
berulis); pedicellis 2-5 mm. longis; receptaculo cupuliformi 
sub anthesi 2 mm. diametro; tepalis 5 patulis carnosis ob- 
longis rotundetis circiter 1 mm. longis et latis; velo car- 
noso plano circiter 0.7 mm. lato, ore minuto; staminibus 4 
vel 5 carnosis oblongis subacutis, 1-1.2 mm. longis, 0.7-1 
mm. latis, dorsaliter minute luteo-glandulosis, antheris 
filamenta subaequantibus, per poros ovales contiguos 0.4 mm. 
longos dehiscentibus; inflorescentiis @ desideratis. 

Type, Mutis 1843, collected in Colombia and deposited in 
the herbarium of the Jardin Botanico, Madrid. It is a spe- 
cies very similar in general appearance to S. Archeri A. OC. 
Smith, from which it differs by its totally glabrous habit, 
its longer @ inflorescence, its slightly larger flowers with 
spreading tepals, and its reduced number of stamens. The re- 
lationship of the new species to S. nicaraguensis Hemsl. 
should also be noted, but the Central American plant has a 
slightly puberulous habit, shorter inflorescences, and dif- 
ferently shapéd leaves than S. hemisphaerica. The specific 
name refers to the shape of the receptacle. 


SIPARUNA EPISCIAEFOLIA A. C. Smith, sp. nov. Frutex dio- 
ecus; ramulis subteretibus fuscis parce pilosis (pilise his- 
pidis plerumque simplicibus); foliis alternis, petiolis sub- 
teretibus rugosis fuscis parce hispidis 7-17 mm. longis, la- 
minis fuscis papyraceis oblongis, 10-12 cm. longis, 4-4-5 


em. latis, basi acutis vel anguste obtusis, apice acutis, 


margine crassis et crenatis vel calloso-denticulatis, utrin- 
que praecipue ad nervos pilos plerumque simplices 0.4-0.6 
mm. longos gerentibus, pinnatinerviis, nervis secundariis 
utroque 11-14 arcuato-adscendentibus prope margines anasto- 
mosantibus, cum costa supra subplanis subtus leviter eleva- 
tis, venulis subtus prominulis; inflorescentiis So axillari- 
bus saepe binis 1-2 cm. longis; pedunculo subglabro flexuoso 
ad 1.5 cm. longo; floribus nigrescentibus extra parce fusco- 
hispidulis (pilis ad 0.3 mm. longis), 2-3 mm. diametro; ped- 
icellis 1-3 mm. longis; receptaculo obconico- sub anthesi 1.5 
mm. diametro; tepalis 5 (vel 4) carnosis patulis saepe re- 
curvatis oblongis obtusis maturitate O.7-1 mm. longis et la- 
tis; velo tenuiter carnoso vel submembranaceo conico-elevato 
circiter 0.5 mm. lato, extra saepe luteo-glanduloso, ore 1 
mm. diametro; staminibus plerumque 3 carnosis oblongis ob- 
tusis, 1.3-1.7 mm. longis, 0.7 mm. latis, dorsaliter parce 
luteo-glandulosis, antheris quam filamentis paullo breviori- 
bus, per poros ovales contiguos 0.4 mm. longos dehiscenti- 
bus; inflorescentiis @ desideratis. 

Type, Mutis 1901, collected in Colombia and deposited in 


128 PRET OL OaER No. 3 


the U. S. National Herbarium (no. 1,560,008). Duplicate at 
M. It ie a species related to the Peruvian S. hispida A... 
DC., from which it differs by the less dense and less hispid 
pubescence and the narrower leaves. Flowers of the new spe- 
cies consistently have 3 stamens, a number which is conspic- 
uously low for the genus. As o& flowers of S. hispida are not 
known, the plants cannot be compared in regard to staminal 
characters. 


SIPARUNA METENSIS A. C. Smith, sp. nov. Frutex dioecus; 
ramulis teretibus hispido-pilosis (pilis simplicibus vel e 
basi 2-3-fidis O.5-1 mm. longis)s; petiolis suboppositis ter- 
etibus 5-8 mm. longis velut ramlis pilosis; laminis papy- 
raceis obovato-oblongis, 12-17 em. longis, 5-7 cm. latis, 
basi obtusis, apice subito acuminatis, margine crenatis vel 
apiculato-dentatis (dentibus plus minusve callosis 4-5 per 
centimetrum), utringue pilis 1-3-fidis pilosis, pilis secus 
nervos principales densioribus, rervis secundariis utroque 
9-12 arcuato-adscendentibus, cum costa supra prominulis sub- 
tus elevatis, venulis utringque planis vel promimulis; in- 
florescentiis & et 9 cymosis, cymis in foliorum axillis 2-4 
paucifloris 12-30 mm. longis, cum floribus parce hispidis 
(pilis e basi 1-3-fidie 0.3 mm. longis); pedicellis circiter 
4 mm. longis; floribus @ maturitate 3 mm. diametro; recepta- 
culis subglobosis limbo angusto obtuse 5-gono (e tepalis 
connatis constante) circumdatis; velo leviter elevato, ore 
minuto; staminibus 4, antheris minutis, filamentis dilata- 
tis minute luteo-glandulosis 1.5 mm. longis et latis apice 
contractis; floribus ¢g velut o, stylis circiter 15 leviter 
exsertis leviter cohaerentibus sed non connatis; drupis 
juvenilibus obconicis pilosis 6 mm. diametro. 

Type, Brother Apollinaire, collected in Dec., 1929, at 
Villavicencio, Intendencia Meta, Colombia, and deposited in 
the herbarium of the New York Botanical Garden. Duplicate at 
N. The type collection bears flowers. Another collection 
from the same locality, bearing 9 flowers, is Brother Nic- 
éforo Maria (Jan., 1927)(N, Y). It is a species related to 
the preceding, from which it differs by its shorter peti- 
oles, larger leaves, and its minute tepals and anthers. 


TRIGONIACEAE 


TRIGONIA MEMBRANACEA A. C. Smith, sp. nov. Arbor parva; 
ramis violaceis teretibus glabris lenticellatis; ramlis 
fuscis striatis cinereo-puberulis mox glabrescentibus; pet- 
iolis gracilibus 3-4 mm. longis decidue puberulis; laminis 
membranaceis viridibus glabris (subtus costa et axillis ner- 
vorum interdum puberulis) ovato-oblongis, 5-7 cm. longis, 2- 
2.5 cm. latis, basi obtusis vel rotundatis, apice cuspidatis 
saepe calloso-mucronatis, margine integris, nervis secund- 


. ~. 
© a> Dita iat actalialal 


ee 


1935 Smith, Studies of South American Plants - IV 129 


ariis plerumque 4 utroque arcuato-adscendentibus anastomo- 
santibus, cum costa utrinque leviter elevatis subtus pal- 
lidis; inflorescentiis ut videtur terminalibus 4-6 cm. lon- 
gis, rhachidibus puberulis; capsulis breviter pedicellatis 
fuscis glabris subrugosis ovoideo-trilobatis, 18-20 mm. lon- 
gis, circiter 10 mm. laetis, valvis concavis intus flaves- 
centibus apice bifidis, marginibus introflexis; seminibus 
1.5 mm. longis, lana lutescente 10 mm. longa circumvestitis. 

Type, Killip & Smith 14396, collected Nov. 11, 1926, in 
woods at Torrecilla, near Turbaco, Bepartment of Bolivar, 
Colombia, alt. 150-300 m., and deposited in the herbarium of 
the New York Botanical Garden. Duplicates at A, G, N. It is 
distinguishable from other species of the gemus by the size, 
shape, texture, and venation of its leaves. Its relation- 
ship is probably with the Guiana species T. microcarpa Sagot 
and T. laevis Aubl., than both of which its leaves are more 
delicate in texture and with fewer lateral veins, and its 
inflorescences probably shorter. 


TRIGONIA CRASSIFLORA A. C. Smith, sp. nov. Arbor parva; 
ramulis subteretibus striatis fuscis, pilis ferrugineis cir- 
citer 0.5 mm. longis obtectis, demum glabrescentibus et len- 
ticellatie; stipulis interpetiolaribus (mox deciduis) sub- 
coriaceis puberulis oblongo-lanceolatis, circiter 10 mm. 
longis. et 3 mm. latis, apice obtusis; petiolis leviter can- 
aliculatis basi incrassatis, 3-4 mm. longis, decidue strigo- 
sis; laminis papyraceis concoloribus oblongis, 12-17 cm. 
longis, 5-9 cm. latis, basi subcordatis, apice breviter 
acuminatis, margine integris et leviter undulatis, supra 
parce papillosis et scabridulis, utrinque glabris (subtus 
nervis principalibus interdum strigosis), costa subtus pro- 
minente fusca, nervis secundariis utroque 7 vel 8 arcuato- 
adscendentibus superne anastomosantibus utrinque elevatis, 
venulis copiose reticulatis utrinque prominulis; inflores- 
centiis terminalibus anguste paniculatis densifloris, ad 12 
cm. longis et 3 cm. latis, rhachidibus ramisque crassis den- 
sissime ferrugineo-tomentosis (pilis ad 0.8 mma. longis); 
floribus in ramis secundariis brevibus subsessilibus, bract- 
eis ovatis subtentis (bracteis acutis, ad 6 mm. longis et 4 
mm. latis, extra ferrugineo-sericeis intus pallido-puberu- 


lis); calycis lobis oblongo-ovatis 6~7. mm. longis, extra 


ferrugineo- vel cano-tomentulosis intus glebris; petalis 
membranaceis glabris insequalibus; petalo postico 7 mm. 
longo, inferne in gibberem 4 mm. longum excavato, ad faucem 
sacculi breviter ferrugineo-piloso, superne plano rotundato, 
margine undulato; petalis lateralibus obovato-spathulatis 
planis, 7 mm. longis, 4 mm. latis, apice rotundatis; petalis 
anticis obovatis, circiter 5 mm. longis et 4 m. latis, 
medio intus carinatis; staminibus fertilibus 8 glabris, 3-4 


130 PHYTOLOGIA No. 3 


mm. longis, filamentis prope ad apices connatis, antheris 
ovoideis 1 mm. longis; glandulis 2 castaneis ad 2 mm. lon- 
gis; ovario ovoideo, pilis stramineis ad 1 mm. longis dense 
hispido; stylo 4 mm. longo superne incrassato; stigmate albo 
discoideo 1 mm. diametro, medio depresso. 

Type, R. A. Toro 1115, collected June 20, 1928, at Puerto 
Berrio, Department of Antioquia, Colombia, and deposited in 


the herbarium of the New York Botanical Garden. It is a very . 


distinct species on the basis of its large flowers and 
bracts, its robust inflorescence, and its concolored gla- 
brous subcordate-based leaves. The Peruvian T. virens Macbr. 
seems to be its closest ally, but has pedicellate flowers 
and a different type of inflorescence pubescence, as well as 
being less robust throughout. 


VACCINIACEAE 


CERATOSTEMA RIMBACHII A. C. Smith, sp. nov. Frutex par- 
vus; ramulis cinereis subteretibus glabris, petiolorum basi- 
bus~incrassatis; stipulis minutis (1 mm. longis) mox decidu- 
is; petiolis subnullis (ad 1 mm. longis); laminis coriaceis 
glabris oblongis vel obovato-oblongis, 12-18 mm. longis, 5-8 
mm. latis, basi rotundatis vel late cuneatis, apice obtusis, 
margine integris incrassatis, costa utrinque prominula, ner- 
vis secundariis 2 vel 4 e costa prope basin orientibus sed ~ 
obscuris; floribus axillaribus solitariis glabris, bracteis 
minutis 1 vel 2 subtentis; pedicello rugoso 1 mm. longo de- 
cidue minute glanduloso-piloso, cum calyce articulato; caly- 
cis tubo obconico sub anthesi 1.5-2 mm. longo, limbo sube- 
recto papyraceo lobis inclusis 2 mm. longo, lobis deltoide- 
is, 1.5 mm. longis; corolla subcarnosa subcylindrica 10-12 
mm. longa, 3 mm. diametro, lobis oblongis obtusis 1-2 mn. 
longis; staminibus 7-10 quam corolla paullo brevioribus; 
filamentis nigrescentibus glabris 1-2 mm. longis; antheris 
7-8 mm. longis, tubulis flexilibus quam loculis 2-3-plo lon- 
gioribus; stylo corollam aequante, stigmate subcapitato. 

Type, Rimbach 188, collected in February, 1934, on the 
_ @astern slope of Mt. Tunguragua, Province of Tunguragua, 

Ecuador, and deposited in the herbarium of the Field Museum 
(no. 740,295). Duplicate at Y. It is a species allied to OC. 
coronarium Linden, from which it differs by its subsessile 
flowers, differently shaped leaves, and its glabrous and 
less compact habit. The reduced number of stamens is unusual 
in the genus. C. Rimbachii bears a resemblance to certain 
species of Disterigma, but the lack of large pedicellary 
bracts, the presence of intrapetiolar stipules, and the 
branchlets, which are swollen at the base of each petiole, 
indicate the place of the new species in Ceratostema. 


t 


CERATOSTEMA COLOMBIANUM A. O. Smith, ep. nov. Frutex; 


oa oe 


1935 Smith, Studies of South American Plants - IV 131 


petiolis rugosis 2 mm. longie juventute puberulis; laminis 
oblongis vel ovato-oblongis, 20-35 mm. longis, 9-15 mm. la- 
tis, basi cordatis, apice rotundatis vel obtusis, subtus 
pilosis glabrescentibus, obscure 5-nerviis; inflorescentiis 
2-5-floris; pedicellis 5-8 mm. longis pilosis, cum calyce 
articulatis; calycis tubo 3-4 mm. longo, limbo 4 mm. longo, 
lobis deltoideis 2 mm. longis; corolla circiter 15 mm. longa 
et 6 mm. diametro pilosa; staminibus alternatim leviter in- 
aequalibus, filamentis basi subconnatis, antheris 9.5-10 mm. 
longis, tubulis quam loculis 2-3-plo longioribus rimis de- 
hiscentibus. 

Type, Ariste Joseph B.112, collected on the eastern para- 
mos of Guasca, towards Gacheta, Department of Cundinamarca, 
Colombia, and deposited in the U. S. National Herbarium. The 
following specimens from Cundinamarca also represent the 
speciest near Bogoté, Triana 2688 (BM)s Guasca, E. Perez Ar- 
bolaez 1178 (K, N). In my recent monograph (a) the Ariste 
Joseph specimen was erroneously referred to Oeratostema par- 
vifolium Benth., which name was properly referred, elsewhere 
in the same paper, to Thibaudia parvifolia (Benth.) Hoer. My 
description of C. parvifolium (a) was based entirely upon 
Ariste Joseph's specimen; therefore that complete descrip- 
tion may be taken to augment the above. 


CAVENDISHIA NITIDA (HBK.) A. C. Smith, comb. nov. Thi- 
baudia nitidea HBK. Nov. Gen. & Sp. 3: 271. 1818. Psammisia 
nitida Kl. Linnaea 24: 45. 1851. Macleania nitida Hoer. Bot. 
Jahrb. Engl. 42: 269. 1909. Cavendishia obtusa A. O. Smith, 
Contr. U. S. Nat. Herb. 28: 498, pl. 16% 1942. A specimen 
collected in Tolima by Humboldt and Bonpland, deposited in 
the Botanisches Museum at Berlin, is erroneously indicated 
as the type of Thibaudia nitida, and was accepted as that 
species both by Hoerold and the present writer. However, the 
actual type of Thibeudia nitida, to which the description 
more accurately applies, is deposited in the herbarium of 
the Jardin des Plantes, Paris. Consequently the above new 
combination ie necessary. The specimens cited in my recent 
monograph as Cavendishia obtusa should be referred to OC. 


nitida. 


MACLEANIA RUPESTRIS (HBK.) A. C. Smith, comb. nov. Thi- 
baudia rupestris HBK. Nov. Gen. & Sp. 3: 270. 1818. Psam- 
migia rupestris Kl. Linnaea 24: 45. 1851. Other synonyms 
cited in my recent monograph (b). The use of the nam Thi- 
baudia nitida for a species of Cavendishia, as already not- 
ed, makes necessary the above new combination for the common 
Andean species of Macleania, although I regret the necessity 
of discarding Hoerold's name M. nitida for this well-known 
species. The specimens cited by me as M. nitida should be 


132 PRY? OL 0 6-58 No. 3 


referred to M. rupestris. The type collection may be seen at 
both Parie and Berlin. 


MACLEANIA MOLLIS A. C. Smith, sp. nov. Frutex; ramlis 
angulatis pilis pallidis ad 1 mm. longis dense vestitis; 
petiolis rugosis pilosis 2-3 mm. longis; laminis coriaceis 
late ovatis, 4-5 om. longis, 2.5-3.3 om. latis, basi cor- 
datis, apice obtusis, margine integris et leviter revolutis, 
supra juventute pilosis mox glabris, subtus praecipue ad 
nervos molliter albo-pilosis, 5- vel 7-pli-nerviis, nervis 
supra impressis subtus prominentibus, venulis reticulatis; 
inflorescentiis axillaribus fasciculatis 1-4-floris bracteis 
minutis subtentis; floribus ubique pilos albos circiter 0.4 
mm. longos gerentibus; pedicellis 1-3 mm. longis; calycis 
tubo obconico vel obscure 5-angulato, circiter 3 mm. longo 
et diametro, limbo erecto-patente lobis inclusis 3 mm. 
longo, lobis 5 deltoideis apiculatis, 1.5 mm. longis, 3 mn. 
latis; corolla urceolata, 18 mm. longa, prope basin 5 mm. 
diametro, lobis parvis; staminibus 10 aequalibus, 11 mm. 
longis; filamentis glabris nigrescentibus 3 mm. longis; an- 
theris circiter 9 mm. longis, tubulis 2 angustis basi later- 
aliter connatis loculos aequantibus, rimis longis dehiscent- 
ibus; stylo corollam aequante. 

Type, Rimbach 119, collected in 1932 near Riobamba, Pro- 
vince of Chimborazo, Ecuador, and deposited in the herbarium 
of the New York Botanical Garden. It is related to M. hirti- 
flora (Benth.) A. O. Smith, from which it differs by its 
pli-nerved rather than pinnate venation, the more pronounced 
pubescence of all its parts, its fasciculate y te 
and its short pedicels. 


(a) Contr. U. S. Nat. Herb. 28: 346. 1932. 
(b)-1. c. 576. 


BOTANIOAL MISCELLANY 
H. A. Gleason 


TIBOUCHINA ERYTHROPHYLLA Gl. sp. nov. § Pseudopterolepis: 
caules diffusi prostrati ramosi gracillimi, juventute acute 
4-anguleati, demim subteretes, ad angulos sparse pubescentes 
pilis curvato-adscendentibus fere 1 mm. longis, nodis saepe 
setosis, internodiis ramorum brevissimis; petioli 2--} mm. 
longis laminae firmae ovatae, saepe circ. 8 mm. longse et 4 
mm. latae, maximae 11 mm. longae 6 mm. latae, acutae vel 
subacuminatae, integrae, basi late cuneatae, 5-pli-nerviee, 
subtus glabrae rubrae, supra dense albo-villosulae, venae et 
venulae ohsoletae; flores 4-meri pauci solitarii ramos late- 
rales terminantes; pedicelli 1 cm. longi; hypanthium obconi- 
cum 3.8 mm. longum angustiesime 4-alatum, costis intermediis 
4 obscuris, inter alas glabrum et ad alas tenuiter pubescens 
pilis adscendentibus 0.3 mm. longis; sepala suberecta trian- 
gulari-ovate 1.6 mm. longa 1.3 m. lata l-nervia dense cili- 
ata, seta terminali rigida 2 m. longa, setis basalibus et 
intersepalinis paucis, O.2--1 mm. longis; petala non visa; 
stamina isomorphia sed inaequalia; filamenta glabra, 1.8 vel 
1.5 mm. longa; antherae fere rectae lineares 3.3 vel 2.2 mm. 
longae; connectiva 0.8 vel 0.6 mm. longa lobis basalibus mi- 
nutis rotundatis; ovarium liberum ellipsoideum 4-loculare 3 
mm. longum summo setosum; stylus rectus glaber 7 mm. longus 
stigmte punctiformi. 

Type, Standley & Valerio 49048, collected at Yerba Buena, 
northeast of San Isidro, Prov. de Heredia, Costa Rica, alt. 
about 2000 m., deposited in the United States National Herb- 
arium (1305867). Standley & Valerio 50110, of the same loc- 
ality and date, is identical. Tibouchina erythrophylla is 
one of a small group of Central American species of which T. 
Oerstedii (Triana) Cogn. is best known. It differs from the 
other two species of the group in its very small flowers and 
its leaves red and glabrous beneath. It is further differen- 
tiated from T. Oerstedii by its short and broad sepals. 


Tibouchina nana (Standley) Gl. comb. nov. 
Chaetolepis nana Standley, Field Mus. Pub. Bot. 4: 247. 1929 
Zibouchina Oerstedii var. subsessiliflora Cogn. Monogr. Phan. 

7: 269. 1891. 

The types specimens of the two synonyms are identical. It 
is easily distinguished from the related T. Oerstedii by the 
short sepals and dimorphic hairs on the leaves. Both are 
separated from T. erythrophylla, described above, by their 
pubescent foliage. 


sb) 


134 PHYTOLOGIA | No.3 


TIBOUCHINA OLIGANTHA Gl. sp. nov. § Diotanthera: frutex; 


rami superiores gracillimi juventute complanati, spargissime 


strigosi, pilis 0.4 mm. longis; petioli graciles 8--15 m. 
longi minute sed quam rami densius strigillosi; laminae late 
ovato-oblongae vel ellipticae usque ad 60 m. longae 40 m. 
latae (superiores minores), abrupte et breviter acuminatae, 
margine ciliato-serrulatae, inferne late rotunaatae ad basin 
imam acutatam, 5-7-nerviae, supra scaberulae, pilis adpress- 
isis ed dimidium adnatis, parte libera circiter 0.5 mm. lon- 
ga, subtus ad venas primarias sparse strigosae pilis 0.4 mm. 
longis, ad paginam glabrae; venae primariae supra fere plan- 
aeae, jugis intermediis brevissime connatis, subtus elevat- 
is; .venae secundariae obscurae flexuosae; inflorescentia 
terminalis trichotoma pauciflora fere glabra; cymulee 3-flo- 
rae et in quaque cymila flores laterales bracteis 2 caducis 
late rotundatis 3 mm. longis membranaceis ciliatis involu- 
crati; flores 5-meri; hypanthium campanuletum 5 mm. longum 
scabro-strigosum pilis subulatis albidis ca. 0.8 mm. longis; 
sepala oblonga, dense ciliata, 5 mm. longa basi 1.8 mm. la- 
ta, ciliis arcuato-adscendentibus 0.5 mm. longis, in linea 
centrali sicut hypanthium strigosa; petala cuneato-obovata 
14 mm. longa 8.5 mn. lata superne glanduloso-ciliata, ut vi- 
detur flava; filamenta glabra; antherae subulatae, 6 vel 4 
mn. longae, connectivis 2.4 vel 1.2 mm. longis basi in lobos 
2 rotundatos dilatatie; stylus glaber 12 mn. longus. 

Type, Steinbach 8262, collected at Samipata, Dept. Santa 
Cruz, Bolivia, alt. 2200 m., and deposited at the New York 
Botanical Garden. Under the artificial arrangement of the 
species in Cogniaux' Monograph, T. oligantha may be placed 
next to T. latifolia (Naud.) Britt. The latter differs from 
ours in its narrower leaves and mch smaller and more numer- 
ous flowers. 


TIBOUCHINA VENOSA Gl. sp. nov. § Diotanthera: frutex; 
rami minores tenuissime et adpresse pubescentes mox glabrati 
obscure 4-angulati, internodiis majoribus usque ad 6 em. 
longis; petioli 5--10 mm. longi, sparse adpresseque setosi; 
laminae foliorum lanceolatae, maximae quae suppetunt 50 mm. 
longae 17 ma. latae in ramlis leteralibus, verisimiliter in 
ramis usque ad 100 mm. longae et certe 355 mm. latae, acumin- 
atae, integrae vel minutissime serrulatae, tenuiter ciliatae 
setis 0.4 mn. longis, basi acutae saepe paullo inaequileter- 
ae, 7-pli-nerviae, jugo exteriore foliorum minorum saepe 
marginali, supra pubescentes pilis adscendentibus gracilibus 
0.5--0.5 mm. longis basi vix adnatie, subtus in pagina glab- 
rae ad venas venulasque tenuissime minutissime adpresso-set- 
_ o88e pilis 0.1--0.3 mm. longis; costae supra impressae sub- 
_ tus elevatae, venae secundarise supra obsoletae subtus ob- 
scurae planse flexuosae. Cymae-pauciflorae axillares et ter- 


1935 Gleason, Botanical Miscellany 135. 


minales in ramlis lateralibus paniculas parvas formantes, 
ultra internodia saepe et in pedicellis semper sparse gland- 
uloso-pilosae; bractese caducae sessiles rotundatae 6 mm. 
longae; flores 5-meri; hypanthium late campanulatum vel sub- 
hemisphsericum, 3.7 mm. longum, sparse glanduloso-setosun, 
pilis 0.6 mm. longis arcuato-adscendentibus; sepala anguste 
oblongo-triangularia 2.7 mm. longa ciliata, sinubus rotunda- 
tis; petala obovata circiter 12 mm. longa (flava?); filamen- 
ta glabra 6 vel 5 mm. longa; antherae subulatae leviter arc- 
uatae 5.5 vel 4 mm. longae, connectivis subteretibus 2 vel 1 
mn. longis infra insertionem filamenti in lobos 2 rotundatos 
dilatatis. 

Type, Steinbach 8509, collected at Comarapa, Dept. Santa 
Cruz, Bolivia, alt. 2500 m., and deposited in the Gray Herb- 
arium. T. venosa is apparently most nearly related to T. 


citrina (Naud.) Cogn., in which the leaves are notably wider 


and more sharply acuminate and the flowers less than half as 
large. 


TIBOUCHINA SAXOSA Gl. sp. nov. § Pseudopterolepis: frutex 
ramis superioribus gracilibus elongatis rectis castaneis 
subteretibus, juventute dense adpresso-pilosis, demim glab- 
rescentibus, cortice longitudinaliter fisso; internodiis 
majoribus 4--8 cm. longis, in ramis floriferis milto brevi- 
oribus; petioli dense adpresse setosi usque ad 1 cm. longi; 
laminae foliorum oblongo-lanceoletae usque ad 45 mn. longae 
17 mm. latae acuminatae integrae basi acutae vel fere obtus- 
ae, S-nerviae, supra dense pilosae, pilis subadpressis, basi 
vix adnatis, usque ad 1 mm. longis, gracillimis, a venis 
primariis arcuato-adscendentibus et inter venas convergenti- 
bus, subtus pallidiores ad venas densissime strigosae, pilis 
fere 2 mm. longis, et ad paginam molliter pubescentes, pilis 
erectis 1 mm. longis; venae primariae supra valde impressae 
laterales 4--7 m. ultra basin confluentes; rami superiores 
foliosi 4--10 cm. longi valde divergentes ex axillis omi- 
bus, foliolis valde reductis; cymae axillares et terminales 
pauciflorae, pedunculis et pedicellis adpresso-setosis; 
flores 4-meri; hypanthium campamlatum 3.5 mn. longum sparse 
strigoso-setosum; calycis tubus 0.3 mm. longus, lobi ad- 
scendentes 1.3 mm. longi supra basin late triangularem subu- 
lati; petala. obovata valde inequilatera purpurea superne 
ciliata, 7 mm. longa; antherae subulatae, 4.3 vel 3.4 mm. 
longae, connectivis 1.2 vel 0.6 mm. longis, ad faciem anter- 
iorem cenaliculatis, basi in lobos 2 rotundatos vix dilatat- 
is; ovarium 4-loculare superne molliter albo-pilosum. 

Type, Pennell 13992, collected in rocky forest at Pilla- 
huata, Cerro de Gusilluyos, Dept. Cusco, Peru, alt. 2200-- 
2400 m., in the Gray Herbarium. Tibouchina saxosa appears 
to have no related species in the section Pseudopterolepis 


1356 PHYTOLOGI4& No. 3 


with which it may be compared. Superficially it- resembles T. 
asperifolia Cogn., @ Peruvian plant with 5-mrous flowers. 


TIBOUCHINA BRACHYPHYLLA Gl. sp. nov. § Diotanthera: rami 
juveniles angulati sparse setoso-hispidi, pilis 1-2 mm. lon- 
gis, demim glabrescentes subteretes; petioli fere glabri in 
ramis sterilibus 2--4 mm., in ramlis floriferis 1--2 mn. 
longi; laminae variae, ovato-lanceolatae, ovatae, vel late 
ovatae, in remis sterilibus usque ad 25 mm. longae 14 m. 
latee, in ramalis floriferis saepissime 8-10 mm. longae 5-6 
mm. latae, obtusae vel acutae, 3-(vel interdum in majoribus 
5-)nerviee, supra (minores) glabrae, sparse ad apicem setul- 
osee, vel (majores) in zonis 4 longitudinalibus setosae, 
subtus glabrae vel ad costas sparsissime setulosae; flores 
S-meri, verisimiliter ebracteati, in cymlis 1--5-floris ad 
ramilos axilleribus et terminalibus, pedicellis 1--3 mm. 
longis; hypanthium subglobosum 3 mm. longum glabrum vel par- 
cissime strigillosum, pilis 0.1--0.3 mm. longis; calycis 
tubus membranaceus 0.3 mm. longus in medio sinorum late 
truncatorum setas 2 conico-subulatas 0.4 mm. longas patulas 
vel subreflexas gerens; sepala patula anguste trianguleria 
5 mm. longa ciliata, ad apicem acutam breviter setosa caet- 
erum glabra; petala obovata glabra 9 mm. longa; filamenta 
glabra 4.6 vel 35.6 mm. longa; antheree subulatae vix arcua- 
tae 4.1 vel 3.2 mm. longae; connectivum in ser. ext. 1 m. 
longum leviter curvatum basi in lobos 2 dilatatum, in ser. 
int. simile sed vix productum 0.5 mm. longum; ovarium 5-loc- 
ulare setis 10 subuletis 0.4 mm. longis coronatum; stylus 7 
mm «. longus 8 

Type, Steinbach 8432, collected in forest at Comarapa, 
Dept. Santa Cruz, Bolivia, alt. 2500--3000 m., and deposited 
at the New York Botanical Garden. A sheet of the same number ~ 
in the Gray Herbarium is identical and also exhibits sterile 
lower branches with larger leaves, as described above. Tib- 
ouchina brachyphylla has mich smller leaves than any other 
Andean species known to me. 


Rhynchanthera paludicola (Donn.Sm.) Gl. comb. nov. 
Tibouchina paludicola Donn. Sm., Bot. Gaz. 42: 293. 1906. 


Torresia cearensis Allemao. This leguminous tree, origin- 
ally described in an-obscure’ publication about seventy years 
ago, is briefly discussed in the supplements to the Natur- 
lichen Pflanzenfamilien. It is stated to have winged pods 
and seeds, but little information about their structure has 
been available. Krukoff's 5495, recently collected in the 
Acre Territory, is certainly of this gems and apparently 
conspecific. It consists only of opened pods, seeds, and 
detached leaflets. 


1935 Gleason, Botanical Miscellany 137 


Neither pods nor seeds were adequately described in the 
Pflanzenfamilien. In our plant the pods are dark brown, lin- 
ear-oblong, 4.5--7 cm. long, rounded at both ends, glabrous, 
and strongly flattened except over the single seed, where 
they are somewhat swollen. The seed is about 5 mm. shorter 
than the pod, solitary and basal. It consists of a basal 
wing, with a linear hilum across the end, and an apical ker- 
nel. The kernel is dark gray, 10 mm. long, flatterned-ellip- 
soid, thinly tomentose, and exhales a strong odor of cumarin 
when opened. The wing conforms to the dimensions of the pod 
and is generally very thin and almost translucent, but it is 
slightly thicker and opaque over the vascular supply near 
one margin. It apparently represents an expansion of the fu- 
niculus, comparable to the esrilloid structures in iting 
genera of legumes. 

The leaflets of our specimen are linear-oblong, 5--10 Cite 
long, 1--2 cm. wide, acuminate, rounded at base, glabrous, 
on petiolules 2 mm. long, and with secondary veins obsolete. 

According to Mr. Krukoff, the native name in the Acre 
Territory is Cumaru de Cheiro, referring to the odor of the 
wood, that of the true tonka bean being odorless. 


ALSEIS LATIFOLIA Gl. ep. nov. Arbor 8 m. alta ramis ter- 
’ etibus novellis ferrugineo-pubescentibus mox glabrescenti- 
bus; folia ad apicem ramorum dense conferta internodiis cir- 
citer 1 mm. longis; petioli 15--25 mn. longi ferrugineo-pub- 
escentes; laminae firme chartaceae obovatae vel oblanceolat- 
ae 8--12 cm. longae 2.5--6 cm. latae acutae vel obtusae nee 
acuminatae basi subito contractae et secus petiolum breviter 
decurrentes, supra ad paginam breviter scabro-pubescentes ad 
costam densius pilosae, subtus dense pubescentes praecipue 
venas venulasque; spicae complures terminales at ex axillis 
superioribus paniculam laxam formantes, fructiferae 8--l1 
em. longae; capsulae sessiles confertae anguste clavatae 8-9 
mm. longae ferrugineo-pubescentes; semina linearia alata. 

Type, Krukoff 2043, collected on terra firma near Coroat- 
asinho, State of Maranhao, and deposited at the New York 
Botanical Garden. While apparently most neerly related to 
A. floribunda, it is amply differentiated from that and 
other species by the broad leaves with broad bases and dis- 
tinctly pubescent beneath and by the short and slender hairy 
capsule, 


= 


TAXONOMIC NOTES ON AMERICAN PHANEROGAMS -- II 
Lyman B. Smith 


LOZANIA BIPINNATA, spec. nov., arbor, 5-8 m. alta: ram- 
lis gracilibus, subglabris: petiolo foliorum ad 1 om. longo, 
sparse puberulis, lamina oblonga, integerrima, breviter 
acuminata, coriacea, 14 cm. longa, 5 cm. lata: inflorescen- 
tiis 1-3 in axillis foliorum, laxe bipinnatis, ad 8 on. 
longis, dense adpresseque puberulis; ramlis perabbreviatis, 
paucifloris, haud ultra 2 mm. longis, bracteis imbricatis 
omnino obtectis: floribus cum pedicellis 2 mm. longis, viri- 
dibus; sepalis latissime ellipticis, obtusis, 0.8 mm. lon- 
gis; stamini ovarium aequanti; ovario globoso, uniloculare, 
appresse puberulis, ovwulo l. Fig. 14. 

Colombia: Dept. Boyaca: high thick forest, El Umbo 
region, northwest of Bogota, alt. 1500 m., Oct. 1932, A. E. 
Lawrance 524. 
~~ GOZANTA PEDICELLATA (Standl.) L. B. Smith, comb. nov. 
Lacistema pedicellatum Standl. in Journ. Wash. Acad. Sci. 


xvii. 1927). 

LOZANIA PITTIERI (Blake) L. B. Smith, comb. nov. Laci- 
ee Blake in Contrib. U. S. Nat. Herb. xx. 
(1924). 

All three of the above species confirm the characters 
given by Mansfeld when he reérected the genus Lozania in 
Notizblatt, xi. 592 (1932). The following key distinguishes 
the five species now known in the genus: 


1. Racemes or panicles usually fasciculate in the aril of 
each leaf. 
2. Flowers in racemes. 
3. Leaves closely serrulate. 
4. Stamen about equaling the ovary. 
1. L. Mutisiana R. & 8. 
4. Stamen much longer than the ovary. 


2. Le Ki i Mansfeld. 
3. Leaves entire.......3. L. Pittieri (Biake} a B. Smith. 
2. Flowers in short-branched panicles. 


4. L. bipimata L. B. Smith. 
1. Raceme always solitary in the axil of each leaf. 
5. L. pedicellata (Standl.) L. B. Smith. 


SPIGELIA POLYSTACHYA Klotzsch. The first known collec- 
tion of this species since that of the type has appeared in 
Iundell's plants of British Honduras. It is no. 1971, from 
Big Fall, Belize River, collected March 17, 1933. This 
station marks a very large jump from the original one in 
British Guiana, but the habitat is almost identical. 

POLYPOMPHOLYX LONGECILIATA (DC.) L. B. Smith, comb. nov. 
Utricularia longeciliata DC. Prodr. viii. 23 (1844). Poly- 

Gupholys lasinista Benj. in Mart. Fl. Bras. x. 251 (16471. 


138 


1945 159 


Leaf and axillary 
panicles X l. 


Young 
flower 


Xx 10. 


Section of ovary X 10. 


LOZANIA BIPINNATA 
Fig. 14. 


140 PHYTOLOGIA No. 3 


A NEW LONCHOCARPUS 
E. P. Killip 


LONCHOOCARPUS PALLIDUS Killip, sp. nov. Frutex; cortex 
glaber cinereo=brunneus, lenticellis albis; folia alterna 
5-foliolata, rhachide 6--9 cm. longa 4-angulata, flavida, 
glabra, petiolulis 4--6 mm. longis, corrugatis, sulcatis, 
foliolis ovatis vel ovato-lanceolatis, 7--1l1 cm. longis, 
3--5 cm. latis, subabrupte acuminatis, basi rotundatis, cor- 
jaceis, supra viridibus et subnitentibus, glabris, subtus 
pallidissimis, glabrie vel minute pubescentibus, nervis et 
venis supra paullo elevatis, subtus non prominentibus; ra- 
ecemi axilleres:- 4-angulati glebri, laxiflori, pedunculis 1- 
floris, 4--5 mm. longis, bracteolis mox deciduis; calyx late 
campanulatus, 3--3.5 mm. longus, circiter 5 mm. latus, min- 
ute rufo-sericeus; flores flavido-albi (?); vexillum suborb- 
iculeari-unguiculatum, lamine glabra circiter & mm. longa, ad 
unguem parce sericeam circiter 1 mm. longam angustata; alee 
oblongae, circiter 10 mm. longae ungue ineluso; staminum ~ 
tubus 7 mm. longus; ovarium adpresse rufo-hirsutum. 

Type in the herbarium of the New York Botanical Garden, 
collected on low land near river, Oamp de Cabeca, Maracas- 
sume River region, State of Maranhao, Brazil, September 8, 
1932, by R. Froes (no. 1882). 

This species belongs to Bentham's group Laxiflori, or to 
Pittier's Epunctati. It is nearest_L. denudatus, but differs 
in the arrangement of the inflorescence, the glabrous leaf- 
rachis, and other details. 


SOME NECESSARY NOMENCLATURAL CHANGES 
(with one new species) - 
H. A. Gleason 


The generic name Martia was proposed by two authore inde- 
pendently and at about the same time; by Sprengel for a 
group of species which he segregated from Hypericum, and by 
Leandri for a@ species now referred to Clitoria. In both 
cases the name began its history as a synonym, but like oth- 
er generic synonyms liable to be revived during segregation 
of a genus. The Martie of Leandri is also a homonym, since 
it is antedated by the Martia of Sprengel. 

Schultes observed this homonymy almost immediately, and 
only one year later proposed the name Martiueia as a substi- 
tute for Martia Leandri. Bentham considered that Martiusia, 
having no validity as a genus, was still available for use 
as a generic name, and set up the leguminous genus Martiusia 
Benth. in 1840. Having been informed that his name was in- 
correct orthographically, he renamed it Martia- during the 
same year, thereby creating another homonym. 

Bentham's genus remains to this day without a valid name, 
under the homonym rule of the International Code. In the 
meantime, Martiusia Schultes has again been used in the se- 
gregation of Clitoria, demonstrating the value of the homo- 
nym rule. These beautiful legumes of Amazonia, one of which 
was actually collected by Martius, were very appropriately 
dedicated to that eminent botanist and explorer, and in re- 
naming them it appears desirable to continue this wish of 
Bentham. I therefore propose the following substitute name: 


Martiodendron Gleason, nom. nov. 
Martiueia Benth. in Hook. Jour. Bot. 2: 84. 1840. Not Mart- 
iusia Schultes, 1822. 
Martia Benth. op. cit. 146. Not Martia Spreng., 1618. 


Martiodendron excelsum (Benth.) Gleason, comb. nov. 
Martiusia excelsa Benth. in Hook. Jour. Bot. 2: 64. 1840. 


Martiodendron parvifolium (Benth.) Gleason, comb. nov. 
Martiusia parvifolia Benth. op. cit. 103. 1840. 


Martiodendron elatum (Ducke) Gleason, comb. nov. 
Martiusia elata Ducke, Arch. Jard. Bot. Rio 3: 116. 1922. 


/ 


142 


142 PHYTOLOGIA No. 3 


MARTIODENDRON MACROCARPON Gleason, sp. nov. Arbor excelsa 
ueque ad 45 m. alta, ramis juvenilibus temuissime puberulis 
mox glabris, gemmis axilleribus complanatis bivalvis 1 cm. 
longis; rhachis foliorum 15 cm. longa glabra, petiolo libero 
2--3 cm. longo; foliola alterna 9 vel 10, petiolulo crasso 
nigro 3 mm. longo, laminis subcoriaceis anguste oblongis vel 
oblongo-lanceolatis, 7--l1l cm. longis, 20--3% mm. latis, ac- 
uminatis, apice ipso obtusis vel leviter retusis, basi ro- 
tundatis vel leviter subcordatis, utrinque glabris, supra 
subnitentibus, venis fere obsoletis, subtus opacis brunnes- 
centibus, venis lateralibus utroque latere 15--20 subrectis 
prominentibus, venulis reticulatis; inflorescentia panicula- 
ta miltiflora, pedicellis puberulis brevibus; calyx 15 mm. 
longus imbricatus, in alabastro anguste conicus; sepala an- 


guste lanceolata extra aureo-sericea intus densius et long- 


ius argenteo-sericea numquam late patentia, marginibus ex- 
ternis leviter involutis, marginibus tectis 0.5 mm. latis 
glabris leviter revolutis; petala flava mox decidua oblonga 
vel oblongo-elliptica, 14 mm. longa, petalum superum brevis- 
Sime unguiculatum obovatum 8 mm. latum, petala alia ellip- 
tica, 5--6 mm. lata; stamina 5, inter petala inserta, fila- 
mentis crassissimis, 1 mm. longis; anthera 1 superior 10 mm. 
longa, laterales 15 m., inferiores 12.5 mm., omnes anguste 
lineari-subulatae; pistillum 15 mm. longum, ovario paullo 
complanato leviter sericeo in stylum glabrum angustato; leg- 
umina elliptica 16 cm. longa, 4.5 cm. lata, tenuiter aureo- 
sericea arcte reticulato-venosa, ala dorsalis 5 ventralis 10 
mm. lata, nervis 2 (suturalibus) basi 3 cm. coalitis. 

Type, Krukoff 5015 (in flower), collected near the mouth 
of the Rio Embira, basin of the Rio Jurua, on varzea land. 

The description of the fruit is taken from Krukoff 4950, 
collected at the same locality and agreeing with the type in 
foliage characters. A third specimen is Krukoff 5401, col- 
lected on terra firma near the mouth of the Rio Macauhan in 
the Acre Territory. The leaflets are only 5--7 cm. long and 
15-235 mm. wide; the legumes average a trifle longer and are 
5-6 om. wide and broadly rounded at the base. 


M. excelsum differs fromthe other speciés in the broad 


fruit with narrow wings and in ite hairy anthers. M. elatum 
is certainly very close to M. parvifolium. Ducke states that 
ite buds are smaller, its panicles more pyramidal, and its 
pods sericeous. From the lack of further contrasting state- 
ments, we my infer that the leaf-veins are obscure beneath 
and the sutural veins of the pod separate to the base, as in 
M. parvifolium. M. macrocarpon apparently agrees with M. el- 
atum.in the size of flowers and fruits. It differs notably 
from M. parvifolium in its slightly hairy ovary, its conspi- 
Ccuous leaf-veins, and the coalescent sutural nerves of its 
broad pod. a 


es Te, a 


ae 


1935 Gleason, Nomenclatural Changes 143 


Apoleya Gleason, nom. nov. 
Apuleja Mart. Herb. Fl. Bras. 12%. 1837. Not Apuleja Gaertn. 

Fruct. 2: 439. 1791. 
zenkera Arn., Mag. Zool. & Bot. 2: 548, 1638. Not JZenkera 

Trin., Linnaea ll: 150. 1837. 

Although the International Code. provides that names dif- 
fering by even a single letter my be maintained, it is im- 
probable that anyone would insist on a difference between 
Apuleja, the original spelling, and Apuleia, as used in the 
Flora Brasiliensis and on most herbarium specimens. In pro- 
posing a new name, I have followed the original pronuncia- 
tion as nearly as practicable. 


Apoleya leiocarpa (Vogel) Gleason, comb. nov. 
Leptolobium (7?) leiocarpum Vogel, Linnaea 11: 59%. 1837. 
Apuleja praecox Mart. Herb. Fl. Bras. 123. 1837. 

Apuleia leiocarpa Macbr. Contr. Gray Herb. 59: 2%. 1919. 


Apoleya molaris (Spruce) Gleason, comp. nove 
Apuleia molaria Spruce, Fl. Bras. 15-2: 177. 1870. 


It is with regret that I call attention to the change in 
name of two long established species of Miconia, macrophylla 
(Don) Triana and serrulata (Don) Triana. The first of these 
is such a widespread and commonly collected species that it 
early began to accumlate nomenclatural difficulties. Col- 
lected originally by Pavon at the end of the eighteenth cen- 
tury, it first received botanical recognition from Devid Don 
in 1823, who described it briefly under the name of Chitonia 
macrophylla. The Pavon specimen was unknown to De Candolle 
in 1828, who repeated Don's description verbatim in the Pro- 
dromas, but under the name Diplochaeta, on the basis of pre- 
occupation of the generic name Chitonia by Mocino. Although 
stating in his description that the leaves are crenulate, he 
placed the species in a group with entire leaves, andin a 
second group with crenate leaves he again described the same 
species twice, as Diplochaeté. leucotephata and D. serrulata, 
and also recognized a variety latifolia under the latter. He 
also noted two manuscript names which had not been published, 
In 1844 Steudel described the species again, under the name 
Decaraphe Hostmanni, placing it in a genus now merged in Mi- 
conia which had been proposed in 1840 by Miquel for a Guiana 
species. In 1850 Miquel again used the same specific names, 
but expressed doubt on the validity of Diplochaeta. 

Not until 1851 did any of these specific names appear in 
the genué Miconia. Then Naudin recognized the identity of D. 
Hostmanni and Diplochaeta serrulata and formed the new bino- 
mial Miconia serrulata. Diplochaeta leucocephala was at one 
time considered by him as doubtfully belonging to the same 


144 PHYTOLOGIA No. 3 


species, but later in the same year he named it Miconia leu- 
cocephala, as a questionable species perhaps the same as M. 
Berrulata. He did not see Pavon's specimen but realized that 
it was also a Miconia. For it he made the new binomial Mico- 
nia platyhedra, since the name M. macrophylla wes already in ~ 
use for a Surinam plant now referred to M. prasina. Triana 
in 1871 and Cogniaux in 1887 recognized that serrulata, leu- 
cocephala, and macrophylla were identical, and each used the 
name Miconie macrophylla, disregarding the fact that it was 
already in use. : 

We have then the following state of affairs. The oldest 
valid specific name is macrophylla D.Don, but Miconia macro- 
phylla (Don) Triana_can not be used because it is antedated 
by Miconia macrophylla Steud. The next oldest specific names 
are serrulata and leucocephala. Both were transferred to Mi- 
conia, the former definitely and the latter as a doubtful 
apecies. Miconia serrulata (D0.) Naud. is therefore its cor- 
rect name under the International Code. 

Gremanium serrulatum was described by Don in. 1824..Neudin 
transferred it to Miconia in 1851 and re-named it as Miconia 
galactantha, since he had previously used the name M. serru- 
lata.. Triana and Cogniaux both used the name Miconia serr- 
ulata, but Naudin's combinetion mst stand as the valid name 
of the plant. 

In 1887 Cogniaux diagnosed a Brazilian species under the 
name Miconia robusta, and another species from French Guiana 
as M. techudyoides. Soon discovering that the Guiana plant 
had previously been named Tschudya robusta by Sagot, he at- 
tempted to rectify his error in the Addenda to Flora Brasil- 
iensis in 1888. Here he changed his first M. robusta to M. 
robustissima and transferred Sagot's specific name to Micon- 
ia as M. robusta. This procedure is distinctly contrary to 
the accepted rules of nomenclature: the first species, val- 
idly published, can not receive a new name, while the second 
can not be given a homonym. 


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PHY TOLOGIA 


Designed to expedite botanical publication 


Not. Ty; September, 1935 No. 4 
CONTENTS 

‘C. V. Morton: New Species of Plants from Guatemala..............00- 145 

C. V. Morton: The Genus Besleria in British Guiana...........0...008. 151 


H. N. Motpenxe: A Monograph of the Genus Tectona as it Occurs 


eee emeneranye (Cems: S98: | CULLEDOLION 6 OC ies os) hha ee aU is alae Le Una 154 
HOH, WaLKer: An Aid in Botanical Drawing... 0.60.0 cee cece cee ene 165 
_H.N. Morpenxe: Nomenclatural and Taxonomic Notes ............... 167 
HN. Motpenxe: A Monograph of the Genus Recordia............0..: 171 
my & A. Giresson: Two new Melastomes of the Krukoff Collection......... 174 
NEES OEE SAMSON PRE NELLE AESE le ih ec cee Wd SE LOB RON TPES ane : 
+ ce 

Bi. 


Published by H. A. Gleason and Haro!d N. Moldenke 
3 The New York Botanical Garden 
Bronx Park, New York, N. Y. 


5 ; Price of this number: 50 cents; per volume $5.00 in advance. 
Vol. I, number 3, was issued January 21, 1935. 


NEW SPECIES OF PLANTS FROM GUATEMALA (a) es 
Wh YY YOURE 
C. V. Morton "OT ANICAL 


‘PaViitr 


The following new species have been detected in a large 
collection recently made in Guatemala by Dr. Alexander F, 
Skutch. Several additional new species belonging to critical 
groups will be described in a future paper. 


CHUSQUEA LANCEOLATA Hitchc., sp. nov. Culmus erectus, 10 
m. altus, in diametro ca. 6 cm.; vaginae internodiis brev- 
jiores, ore nudo; laminae planae, lanceolatae, basi rotunda- 
tae, acuminatae, supra glabrae, subtus pubescentes, 12-15 cm 
longae, 1.5-2.5 cm. latae; panicula ca. 15 cm. longa, ramis 
5-7, adscendentibus vel patulis, ramulis brevibus adpressis; 
spiculae 6-9 mm. longae, glabrae; glumae 2, obtusae, plus 
minusve inaequales, 0.5-1.5 mm. longae; lemmata sterilia 2, 
dimidiam spiculam aequantia, acuta vel acuminata; lemma fer- 
tile acuminatum, minute apiculatun. 

Main culms erect, about 10 meters tall, 6 cm. thick at 
base; flowering branches leafy, 30 to 40 cm. long, scaberu- 
lous, the sheaths sligntly roughened, longer than the inter- 
nodes, naked at summit; ligule firm, slightly dentate, 1 m. 
long; blades of the flowering branches lanceolate, flat, 
rather thin, rounded into a short pubescent petiole 2 to 4 
mm. long, acuminute to a fine point, scabrous on the marg- 
ins, rather strongly tessellate-veined, pubescent beneath, 
glabrous but stigiitly roughened on the upper surface, 12 to 
15 cm. long, 1.5 to 2.5 cm. wide, the main veins 5 or 6 
pairs; panicle more or less inclosed in the uppermost 
sheath, about 15 cm. long, the main branches 5 to 7, ascend- 
ing or spreading, floriferous from near the base, the lower 
2.5 cm. distant, 7 to 9 cm long, the axis and branches 
angled, scabrous-pubescent; spikelets 6 to 9 mm. long, purp- 
lish, imbricate on short appressed branches, the ultimate or 
lateral pedicels mostly less than 1 mm. long; glumes round- 

ed, obtuse, more or less unequal, glabrous, the first 0.5-1 

mm. long, the second somewhat longer to twice as long; ster- 

ile lemmas about equal, about half as long as the spikelet, 
acute or acuminate, glabrous except the scaberulous keels, 

3- to S-nerved; fertile lemma about 9-nerved, acuminate, 

_ glabrous, minutely apiculate; palea about as long as the 

4 lemma. 

Type in the U. S. National Herbarium, no. 1,587,761, col- 

..Lected at Santa Elena, Dept. Chimaltenango, Guatemala, alt. 

3000 meters, in cypress forest, December 24, 1933, by A. F. 

_ Skutch (No. 768). 

3 145 


146 PHYTOLOGIA No. 4 


The species is known only from the type collection. The 
type specimen consists of two flowering branches 35 to 40 
em. long, with 3 or 4 leaves and a terminal inflorescence. 
The label gives the height and thickness of the main culm. 


PILEA QUERCIFOLIA Killip, sp. nov. Herba monoica, glabra; 
caules erecti, basi radicantes, simplices vel apicem versus 
ramosi; folia inferioria reducta, decidua, superioria oppos- 
ita, aequalia, ovata, acuminata, basi rotundata vel auricu- 
lata vel interdum obliqua, 3—-nervata, membranacea, subtus 
pallida, grosse serrato-crenata; cymae masculae 10-12-florae 
pedunculatae, pedunculis filiformibus, floribus pedicellat- 
is, segmentis oblongo-lanceolatis; cymae foeminae peduncu- 
latae, floris segmentis inaequalibus; achaenia lanceolato- 
ovata, compressa, laevia. 

Plant glabrous throughout; stem erect, rooting at the 
basal nodes, about 30 cm. high, simple or branched near the 
apex, the lower leaves much reduced and soon deciduous, the 
lower internodes elongate, up to 5 cm. long; stipules ob- 
long-lanceolate, 4 to 5 mm. long, 2 to 2.5 mm. wide, suba- 
cute, soon deciduous; leaves of a node subequal and similar, 
the petioles 0.7 to 3.5 cm. long, the blades ovate, 4 to 12 
cm. long, 2 to 5 cm. wide, acuminate at apex, at base round- 
ed or subauricular, or sometimes oblique, coarsely serrate- 
dentate from apex to base (teeth ascending, rounded), 3- 
nerved (lateral nerves extending to the upper tnird of the 
blade), membranous, pale beneath, bearing obscure linear 
cystoliths on both surfaces; plants monoecious, the cymes 
unisexual; staminate cymes with filiform peduncles 2 to 4 
cme long, 1O- to 12-flowered, 8 to 10 mm. wide, tne flowers 
pedicellate, subglobose in bud and 2 to 2.5 mm. in diameter, 
the segments oblong-lanceolate, about 1 mm. long, dark 
green, pale at base; pistillate cymes borne at the same 
axils with the staminate, on peduncles subequal to or sligh- 
ly shorter than those of the staminate, 7 to 8 mm. wide, the 
segments 3, markedly unequal, the middle one about 0.8 mm. 
long, the lateral ones barely 0.2 mm. long; achenes lance- 
ovate, about 1 mm. long, strongly flattened, obscurely cos- 
tate on one face, smooth. 

Type in the U. S. National Herbarium, no. 1,585,999, col- 
lected at Chichavac, Dept. Cnimaltenango, Guatemala, alt. 
2400 to 2700 meters, in a deep ravine in dense forest, Aug- 
ust 19, 1933, by A. F. Skutch (No. 559). 

This species is related to P. gracilipes Killip, a spec- 
ies common in Costa Riea and western Panama, the two having 
very similar floral characters. The proposed species is, how 
ever, readily distinguished by tne la jer, proportionately 
broader leaves, which have much larger teeth, the toothing 
being, indeed, more pronounced than in any otner species 


1935 Morton, Plants from Guatemala 147 


with which the writer is familiar. In general outline the 
leaves resemble those of Quercus prinus. 


DALEA DISPAR Morton, sp. nov. Frutex usque ad 1.6 m. alt- 
us, diffuse ramosus; caules hornotini virides, subteretes, 
2-5.5 mm. diam., glabri, plus minusve longitudinaliter stri- 
ati; folia alterna, imparipinnata, stipulis lineari-subula- 
tis, usque ad 10 mm. longis, pubescentibus, petiolo ca. 1.5 
mm. longo, viridi, fere glabri, foliolis ca. 7-jugis, ob- 
longis, ca 18 mm. longis et 6 mm latis, oppositis vel alt- 
ernis, apice mucronatis, basi obtusis, membranaceis, pallido 
viridibus, glabris, subtus glanduloso-punctatis, stipellat- 
is, stipellis minutissimis, glabris; inflorescentia racemo- 
sa, anguste cylindrica usque ad 17 cm. longa, floribus dense 
confertis, adscendentibus, pedicellatis, pedicellis brevis- 
Simis, ca. 0.6 mm. longis, crassis, apice articulatis, 
bracteis concavis, lanceolatis, apice longe subulatis, gla- 
bris; calycis tubus anguste campanulatus, 3-3.5 mm. longus, 
10-costatus, perspicue glanduloso-punctatus, villosus, lobis 
subulatis, viridibus, ca. 2.2 mm. longis, villosis, margine 
subspinulosis; flores lutei, vetustiores purpurascentes; 
vexillum longe unguiculatum, ungue usque ad 7 mm. longo, 
limbo suborbiculari, ca. 4 mm. longo et 3.5 mm. lato, apice 
obtuso, basi cuneato, non appendiculato, dorso basi villo- 
80; alae longe unguiculatae, ungue tubo stamineo basi adna- 
to, limbo oblongo, ca. 3 mm longo, valde obliquo, basi uno 
latere unguiculato; carinae petala alis similia, ca. 4.5 mm. 
longa, parum connata, apice et basi libera, unzuibus liber- 
is; stamina 9 in vaginam connata, vexillari deficiente; o- 
varium compressum, hirsutum; stylus villosus. 

Type in the U. S. National Herbarium, no. 1,494,921, col- 
lected at Chichavac, Dept. Chimaltenango, Guatemala, alt. 
2400-2700 meters, February 18, 1933, by A. F. Skutch (No. 
259). An additional specimen was collected at the game loc- 
ality, December 2, 1933 (Skutch 725). 

This is apparently an isolated species without a close 
relative, at least among the North American species. In Ryd- 
berg's treatment in the North American Flora it would seem 
to be closest to the Section Leucostomae, differing essent- 
ially nevertneless from all species of that section. 


GERANIUM PULCHRUM Morton, sp. nov. Sect. Incanoidea? 
Herba perennis; caules ex radice numerosi, decumbentes, no- 
dis saepe radicantes, graciles, 1-1.5 ma. diam., angulati, 
strigillosi (pilis retrorsis) vel demum glabrati; folia opp- 
osita, radicslia longissime petiolata, petiolo usque ad 14 
cm. longo, laminam 3-/-plo longiore, tenui, strigilloso, 
stipulis lanceolatis, ca. 5 mm. Longis, brunneo-scariosis; 
lamina membranacea, ambitu plus minusve deltoidea, usque ad 


148 PH, ¥ BR OcLoQve:® & No. 4 


2.7 cm. longa et 4.3 cm. lata, fere usque ad basin quinque- 
partita, segmentis pinnatifidis, ultimis lanceolatis, acu- 
tis, pallido-virides, supra pilis sparsis hyalinis rectis 
acutissimis minutis, sursum spectantibus dissita, subtus 
hirsutula; pedunculus axillaris, longus (usque ad 10 cm), 
pubescens, pilis patentibus vel subadpressis, interdum 
glendulosis, biflorus, pedicellis usque ad 2.5 cm. longis, 
glanduloso-pilosis, bracteis subulatis, ca. 5.5 mm. longis; 
calycis lobi ca. 6.5 mm. longi, ovati, virides vel purpureo- 
tincti, subulato~mucronati, trinervii, extus strigillosi, 
pilis longioribus patulis intermixtis; petala lilacina, pur- 
pureo-venosa, oblanceolata, ce. 13 mm. longa, 4 mm. lata, a- 
pice rotundata, intus basi hirsuta; stamina basi dilatata, 
ciliata; styli et ovaria pubescentes; fructus deest. 

Type in the U. S. National Herbarium, no. 1,587,712, col- 
lected at Santa Elena, Dept. Cnimaltenango, Guatemala, alt. 
re 2850 mters, November 25, 1933, by A. F. Skutch (No. 
109). 

The species is related apparently to G. Palmeri Rose, of 
central Kexico, but that species has much broader petals 
(over 10 mm. in width) anc longer pedicels, these strigill- 
ose rather tnan glandular-pilose. Furthermore, the stems of 
G. Palmeri are not prostrate and do not root at the nodes, 
The latter cheracter seems to place G. pulchrum in Knuth's 
section Diffusa, although it bears little resemblance to the 
other species, all South American. It is more probably to be 
placed in the section Incanoidea, 


BUDDLEIA SKUTCHII Morton, sp. nov. Caules teretes, cae 5 
mm. diametro, stellato-puberuli, demum glabrati; folia opp- 
esita, stipulata, stipulis connatis, semiorbicularibus, rev- 
olutis, crassiusculis, petiolata, petiolo usque ad 27 mm. 
longo, dense stellato-puberulo, supra canaliculato, lamina 
oblongo-laenceolata, usque ad 16 cm. longa et 5.5 cm. lata, 
coriacea, integra, apice acuminata, basi cuneata, obliqua, 
eupre viridi, spyarse stellato-puberula, mox glabreta, venis 
impressis, subtus dense stellato-tomentosa, pilis minutis, 
albidis vel aureis, venis elevatis, reticulatis; capitulae 
ca. 16-florae, flobosae, densae, 3-5 mm. diametro, in panic- 
ulam terminalem sessilem 13 cm. longam et 16 cm. latam dis- 
positae; bracteae minutae, lineares, stellato-tomentosae; 
flores eurei, sessiles; calycis tubus ca. 0.5 mm. longus, 
lobis ce.’l mm longis, deltoideis, extus stellato-tomento- 
sis, epice ecutis; corolla aurea, tubo turbinato, ca. 1.5 
ma. longo, glabro, lobis deltoideis, ca. 1.5 mm. longis, 
obtusis, suberectis, extus stellato-puberulis, intus sparse 
pubescentibus; stamina 4, fauce sffixa, filamentis glabris, 
ca. O.5 ma. longis, entheris oblongis, ca. 0.5 mn. longis, 
introrsis, loculis distinctis, parallelis; ovarium apice 


1935 Morton, Plants from Guatemala 149 


pubescens; stylus indivisus, clavatus; fructus deest. 

Type in the U. S. National Herbarium, no. 1,494,931, col- 
lected at Chichavac, Dept. Chimaltenango, Guatemala, alt. 
2400-2700 meters, February 2, 1933, by A. F. Skutch (No. 
269). An additional specimen was collected at the same place 
in November, 1930 (Skutch 54). 

The species is distinguished at once from all other North 
American svecies of Buddleia by its small pedunculate heads, 
disposed in large terminal sessile panicles. The large per- 
sistent stipules and entire leaves also distinguish B. 


Skutchii from most of the other species. 


SOLENOPHORA PIRANA Morton, sp. nov. Frutex 1.8-2.4 m. alt 
us; caules fistulosi, ca. & mm. diam., subquadrangulati, 
villosi, pilis hyalinis, multiseptatis, flaccidis, demum 
subglabrati; folia opposita, longepetiolata, petiole usque 
ad 10.5 cm. longo, villosulo, lamina elliptica, magna, usque 
ad 30 cm. longa et 14 cm. lata, duplicato-dentata, apice 
breviter acuminata, basi cuneata, inaequali, tenuiter mem- 
branacea, supra viridi, pilis satis sparsis multiseptatis 
obsiti subtus pallidiore, fere glabra, venis albidis per- 
spicue reticulatis pilis paucis minutis instructis; inflores 
centia axillaris, subumbellata, 3-5S-flora, pedunculo com- 
muni elongato, usque ad 8 cm. longo, subglabro, bracteis ob- 
longis cae 16 mm. longis, pedicellis usque ad 37 mm. longis, 
glabratis; calycis tubus (pars adnatus) turbinatus, ca. 5 
mm. longus, villosus, pars liberus subcampanulatus, ca. ll 
mm. longus, ca. 12.5 m. latus, subglaber, viridis, apice 
obliquus, lobis inconspicuis, deltoideis, ca. 4 mm. longis, 
6 mm. basi latis, perspicue denticulatis, apice acutis; 
corolla flava, 4-4.5 cm. longa, anguste cylindrica, erecta, 
basi ecalcarata, 8 mm. lata, sursum paullulum ventricosa, 
villosa, intus villosula, limbo vix 10 mm. diametro, lobis 
parvis, subaequalibus, erectis, ca. 5 mm. latis, 2 mm. lon- 
gis, immaculatis, apice fere truncatis, integris; stamina 4, 
didynama, filamentis liguliformibus, glabris, rectis, corol- 
lae basi adnatis, antheris connatis, exsertis, loculis ob- 
longis, discretis; staminodium evolutum; ovarium omnino in- 


ferum; stylus rectus, pubescens, stigmate dilatato, concavo; 


disci glandulae 5, irregulares, fere distinctae, pubescent- 
es; fructus deest. 

Type in the U. S. National Herbarium, nos. 1,587,684 and 
1,587,685, collected near Chichavac, Dept. Chimaltenango, 


Guatemala, alt. about 2550 meters, along a stream in a deep 


ravine, November 11, 1933, by A. F. Skutch (No. 680). An 
additional specimen was collected at Santa Elena, Dept. Chi- 
maltenango, Guatemala, June 2, 1933 (Skutch 346). 

The species is named at the request of Dr. Skutch in hon- 
or of Sefior Don Axel Pira, his host in Guatemala. 


150 P H-Y-9O LO46tE is No. 4 


The species is most closely related to S. insignis (Mart. 
& Gal.) Hanst., a Mexican species known only from the orig- 
inal collection. The description by Martens & Galeotti is 
short, but a more complete one is given by Fenzl (b). S. 
Pirana differs in having the disk composed of five rather 
than two glands and the corolla pilosulous, ratier than gla- 
brous within. 


(a) Published by permission of the Secretary of the Smith- 


sonian Institution. 
(b) In Otto & Dietr. Allg. Gartenz. 16: 306. 1848, 


THE GENUS BESLERIA IN BRITISH GUIANA (a) 


C. V. Morton 


Up to the present time only one species of Besleria 
(Gesneriaceae) has been known from British Guiana. In con- 
nection with monoszraphic studies of this genus the writer 
has noted three more species, none of which can be identifi- 
ed with any previously described, 


Key to species 


Calyx lobes free. 

Calyx more than half as long as the corolla (Subg. Pseudo- 
besleria); flowers cymose on a common peduncle; cor- 
olla red; leaves and stems avpressed-pubescent..e.eee 
SUES USERS CESE CRE CHES HOR TE REDRESS Co anewS Be laxiflora. 

Calyx one-third as long as the corolla or less (Subg. 
Parabesleria); flowers aggregate in tiie leaf axils; 
corolla yellow; leaves and stems hirsute.....ccccece 
ere c cece ccccccccccccccccccscceecsceece Be SAXiCOlA. 

Calyx lobes united for over one-third their length (Subg. 
Eubesleria); leaves and stems appressed-pubescent,. 

Flowers borne on a common peduncle; calyx lobes equal to 

. or a little Ionger than the calyx tube...ccccevsccce 
eer c recs eer ecccccccccccccesccesoveerte Be Vorecunda. 

Flowers aggregate in the leaf axils; calyx lobes one-half 
as long as the calyx tube.........+ee4. Be. insolita. 


1. BESLERIA LAXIFLORA Benth., Lond. Journ. Bot. 5: 361. 1846. 

Through the courtesy of the Director of the Royal Botanic 
Gardens at Kew, I have been able to examine the type of this 
species, an unnumbered specimen collected by Schomburgk in 
British Guiana. The North American plant which has passed as 
B. laxiflora is found to be quite different and must bear 
the name B. chiapensis Brandeg. A second specimen of B. lax- 
iflora, also in the Kew herbarium, was collected at Surinam, 
Rio Branco, Amazonas, Brazil, by E. Ule (No. 8459). 


2. BESLERIA SAXICOLA Morton, sp. nov. 

Subg. Parabesleria. Frutex 1.5-1.8 m. altus, vix ramo- 
sus; caules teretes, superne hirsutissimi, inferne glabres- 
centes, ca. 6 mm. diametro; lamina foliorum oblique ellipti- 
ca, membranacea, maxime ca. 30 cm. longa et 12 cm. lata, a- 
_ pice breviter acuminata, basi late vel anguste cuneata, re- 
mote serrata, supra sparse hirsuta demum,glabrescens, sub- 
tus imprimis in nervis hirsuta, nervis secundariis 7-9; pet- 

Lox 


152 PHYTO GD O16 A No. 4 


iolus usque ad 10 cm. longus, angulatus, hirsutus; peduncu- 
lus communis nullus; pedicelli numerosi in axillis foliorun, 
vix 1 cm. longi, hirsuti; lobi calycis liberi, lutei, ovati, 
valde imbricati, 6-8 mm. longi, apice acuminati vel subula- 
to-acuminati, parce hirsuti, longe ciliati; corolla lutea, 
ca. 20 mm. longa, utrinque glabra, basi non saccata, sursum 
ampliata et paullulum ventricosa, lobis vix patentibus, 
late rotundatis, glabris; filamenta libera, crassa, glabra; 
antnerae liberae vel connatae, loculis confluentibus; stam 
inodium bene evolutum, ca. 3 mm. longum, glabrum, antheram 
sterilem gerens; ovarium conicum, glabrum; stylus puberulus; 
stigma bilobum; discus semiannularis, crassus, glaber. 

Type in the U. S. National Herbarium, no. 1,056,446, 
collected by brook in forest, Tumatumari, Potaro River, lat. 
5°20' N, British Guiana, January 3-5, 1920, by A. S. Hitch- 
cock (No. 17,375); a duplicate in the herbarium of the New 
York Botanical Garden. 

Additional specimens examined: BRITISH GUIANA: Type loc- 
ality, Gleason 420 (Gray Herbarium, N. Y. Botanical Garden, 
National Herbarium); Moraballi Creek, near Bartica, Essequi- 
bo River, Sandwith 58 (Kew, Ne Y. Botanical Garden); Potaro 
River, Abraham 345 (Kew). 

It is related to Besleria montana Rusby, of Bolivia, but 
that species is widely different in its long-pilose corollas. 
A closer relationship is with a species of Peru as yet un- 
described. 


3. BESLERIA VERECUNDA Morton, sp. nov. 

Subg. Eubesleria. Frutex (7) 1.2 me altus; caules superne 
sparse strigosi, inferne glabri, teretes; lamina foliorum 
elliptica vel elliptico-oblonga, maxima 22 cm. longa et 10 
cm. lata, apice acuminata, basi cuneata, in petiolum decur- 
rens, serrata basi excepta, supra pilis paucis hyalinis ap- 
pressis praedita, subtus imprimis in nervis strigosa, nervis 
secundariis 7-9; petiolus longus (usque ad 7 cm.), parce 
strigosus; pedunculus communis usque ad 4.5 cm. longus, 
strigosus, flores paucos (3 vel 4) simpliciter umbellatos 
vel raro cymosos gerens, pedicellis tenuibus ca. 2 cm. long- 
is, fere glabris, apice vix incrassatis; tubus calycis cam- 
panulatus, sparsissime strigosus, ca. 5 mm. longus, lobis /7- 
8 mm. longis, lanceolatis, integerrimis, apice subulato- 
acuminatis; corolla 20 mm. longa, glabra, basi ecalcarata, 
non saccata, sursum vix ventricosa (ca. 7 mm. lata), lobis 
ca. 2.5 mm. longis, erectis, rotundatis, glabris, aequali- 
bus; antherae connatae, in medio tubae corollae sitae, locu- 
‘lis confluentibus; staminodium basi corollae bene evolutunm, 
ca. 2 mm. longum, antheram sterilem gerens; ovarium ovoid- 
eum, glabrum; stylus longus, glaber; stigma bilobum; discus 
annuleris, crassus, glaber; bacca coriacea, tuberculata,sem-- 


1955 Morton, Besleria in British Guiana 155 


inis rubris minutis in utroque parieti placentarum adspers- 
is. , ‘ 
Type in the U. S. National Herbarium, nos. 57,862 and 
57,863, collected on the Upper Demerara River, British Guia- 
na, in September, 1887, by G. S. Jenman (No. 5156). Dupli- 
cate types are at Kew and the New York Botanical Garden. 

Besleria verecunda is not closely related to other spe- 
cies of Eubesleria. It suggests B. laxiflora Benth. (Subg. 
Pseudobesleria), which occurs in the same region, but that 
is at once distinguished by its nearly free calyx lobes, its 
more numerous flowers which are only 12 to 135 m. long, and 
its smaller short-petiolate leaves. 


4, BESLERIA INSOLITA Morton, sp. nov. 

Subg. Eubesleria. Frutex (7) 0.9-1.2 m. altus; caules 
teretes, superne hirto-strigosi, inferne glabrescentes; 
lamina foliorum elliptica vel elliptico-oblonga, maxima ca. 
25 cm. longa et 10 cm. lata, apice breviter acuminata, basi 
cuneata, serrata basi excepta, supra strigosa mox glabres- 
cens, subtus strigillosa, nervis secundariis 8-12; petiolus 
usque ad 7 cm. longus, strigillosus; pedunculus communis 
nullus; pedicelli numerosi, in axillis foliorum, 12-15 mm. 
longi, hirto-puberuli; calyx flavus, cylindricus 7-16 m. 
longus, omnino hirto-puberulus, lobis quam tubo duplo brev- 
joribus, mucronatis, erectis; corolla ochroleuca (f. Bart- 
lett), calyce paullulum longior, glabra, ecalcarata, vix 
ventricosa, Lobis parvis, rotundatis, glabris; antnerae 
connatae; ovarium glabrum; discus anmnularis, glaber, tenuis, 
altus; fructus deest. 

Type in the Kew herbarium, collected in French Guiana by 
Martin. 

Additional* specimens examined: FRENCH GUIANA: Poiteau 
(Kew). BRITISH GUIANA: Holmia, Potaro River, November, 1907, 
A. W. Bartlett 8743 (N. Y. Botanical Garden). A specimen 
without collector or locality of collection designated is in 
the Copenhagen herbarium. 

The present species is related only to B. lutea, of the 
West Indies, from which it is distinct by its more narrowly 
cylindric, hirto-puberulous calyx, cream-colored flowers, 
and thin high disk. 


Ce i a 


(a) Published by permission of the Secretary of the Smith- 
sonian Institution. 


A MONOGRAPH OF THE GENUS TECTONA AS IT OCCURS IN AMERICA 
AND IN CULTIVATION 


Harold N. Moldenke 


The following is the second in my series of monographic 
studies of the genera of Verbenaceae and Avicenniaceae, the 
first having been my monograph of the genus Aegiphila pub- 
lished in 1934 [Brittonia 1: 245-477]. In the list of cita- 
tions of herbarium specimens the following abbreviations of 
the names of herbaria are employed: A = Arnold Arboretum, 
Jamaica Plain, Mass.; B = Botanisches Miseum, Berlin; Ba = 
Le. H. Bailey Herbarium, Ithaca, N. Y.; Ca = University of 
California, Berkeley, Calif.; Cb = Conservatoire et Jardin 
Botaniaques, Geneva; Cp = Universitetets Botaniske Museun, 
Copenhagen; D = Academy of Natural Sciences, Philadelphia; 
E = Missouri Botanical Garden, St. Louis; Es = Estacion Ex- 
perimental Agronomica, Havana; F = Field Museum of Natural 
History, Chicago; G = Gray Herbarium of Harvard University, 
Cambridge, Mass.; K = Royal Botanic Gardens, Kew; L = Jard- 
in Botanique Principal, Leningrad; N = New York Botanical 
Garden, New York City; P = Miséum National d'Histoire Natur- 
elle, Paris; R = Trinidad & Tobago Botanical Garden, Port- 
of-Spain, Trinidad; S = Naturnistoriska Riksmseet, Stock- 
holm; U = Georgetown Botanical Garden, Georgetown, British 
Guiana; V = Naturhistorisches Myseum, Vienna; W = United 
States National Herbarium, Washington; Y = Yale School of 
Forestry, New Haven, Conn.e; and Z = H. N. Moldenke Herbari- 
um, Watchung, N. J. To the directors and curators of the 
above-mentioned herbaria the writer extends his most sincere 
thanks for their courtesy and kindness in allowing him to 
study their material of this genus and for their continuous 
and very generous cooperation throughout the progress of 
this work. All specimens so studied have been annotated 
with uniform printed annotation labels and mention is made 
on each Label that the specimen is cited in this monograph. 
All New World material and all cultivated material thus far 
received from these 22 herbaria is herein accounted for and 
cited. Thirty-one other herbaria have been canvassed, but 
did not contain any New World or cultivated material of this 
group. 


TECTONA L. f. Suppl. 20 & 151. 1781. (a) 
Theka Adans. Fam. Pl. 2: 445. 1763. (b) 
Nautea Noronha, Verh. Batav. Gen. V, ed. 1, art. 4: 3. 
1790. 


154 


- 1935 Moldenke, The Genus Tectona LS) 


Tall trees with soft bark and more or less tetragonal 
branches and branchlets; leaves deciduous, petiolate or 
subsessile, mostly large and broad, entire, opposite or ter- 
nate; inflorescence cymose; cymes numerous, many-flowered, 
borne in massive terminal panicles (with sometimes smaller 
axillary ones in the upper axils); flowers hypogynous, act- 
inomorphic; calyx gamosepalous, campanulate, shortly 5S-/- 
lobed, persistent, in fruit greatly enlarged and often in- 
flated, including the fruit and closed above it; corolla 
gamopetalous, hypocrateriform, white or blue, its tube cy- 
lindric, short, its limb patent or reflexed, 5-/-parted, the 
lobes subequal, overlapping in bud; stamens 5 or 6, inserted 
in the corolla-tube, equal, exserted; anthers ovate or ellip- 
tic-oblong, 2-celled, dorsifixed, the thecae parallel, open- 
ing by longitudinal slits; pistil single, elongate; style 
terminal, capillary; stigma very shortly bifid, its branches 
subequal; ovary compound, composed of 2 carpels, completely 
4-celled (each carpel 2-celled), each cell l-ovulate; ovules 
lateral or high-lateral, hemianotropous; fruit drupaceous, 
rounded or weakly 4-lobed, completely included by the en- 
larged fruiting-calyx, with a thin subcarneous exocarp and a 
thick bony 4-celled endocarp which has a small central cav- 
ity between the cells; seeds without endosperm. 

The oldest name for this important genus of plants is 
Theka, first proposed by Rheede in 1683 and again by Adanson 
in 1763, but, unfortunately, the much later name, Tectona, 
is included in the list of nomina conservanda of the Inter- 
national Rules of Botanical Nomenclature, edition 3, as a- 
dopted at Cambridge in 1930. The genus, because of the type 
species, T. grandis L. f., source of teak, is the most im- 
portant and valuable genus of the Verbenaceae from a purely 
commercial and economic standpoint. Only three species are 
known, all natives of southern and eastern Asia. Besides the 
species which will be discussed in full below, there are T. 
philippinensis Benth. & Hook. f. (c) [Diospyros tectona ~ 
Blanco; Tectona Hamiltoniana Schau., pep.j, a native of Lu- 
zon, which differs in its leaf-blades being only 8-15 cn. 
long and 3-6 cm. wide, its petioles 5-7 mm. long, and its 
_ tomentum very minute, and T. Hamiltoniana Wall. [Tectona 
ternifolia Buch.-Ham.; Theka ternifolia Hamilt.], a native 
of the Malay Peninsula, Burma, and Ava, which differs in its 
branchlets being subglabrous, its leaves usually ternate and 
their blades about 24 cm. long and 12.5 cm. wide, its petio- 
les about 1.6 mm. long, its panicles 15-30 cm. long, its 
calyx-teeth acute, and its corolla blue with a villous 
throat. 

The generic name Tectona is taken from the Latin tectoni- 
cus in allusion to the use of the wood of these plants in 
construction, especially of ships and vessels. Theka is an 


156 PAY TO LO ats No. 4 


adaptation of the vernacular Dravidian name "teka". The gen- 
us is divided into two sections by Briquet: Sect. Lachnaio- 
carpae Briq., characterized by an inflated fruiting-calyx 
and a densely tomentose exocarp, contains only one species, 
T. grandis; Sect. Leiocarpae Briq., characterized by an en- 
larged but not inflated fruiting-calyx, closely appressed to 
the fruit, dnd a glabrous exocarp, contains the two other 
known species mentioned above. 


TECTONA GRANDIS L. f. Suppl. 151. 1781. (e) 
Jatus s. caju jati Rumph. Herb. Amb. 3: 34, t. 18. 1743. 
(ad) 

Tectona Theka Lour. Fl. Cochinch., ed. 1, 137-6 17%. 

Theka grandis (L. f.) Lam. Illustr. 2: 111. 1793. (f) 

Tectona grandis f. abludens Koorders & Valeton, Bijdr. 

Booms. Java, no. 7, 171. 1900. 

Tree to 50 m. tall; branches and branchlets stout, 4- 
sided, with large quadrangular pith; twigs stout, very med- 
ullose, obtusely tetragonal with rounded angles, usually 
more or less sulcate between the angles in drying, densely 
furfuraceous-tomentose with cinereous or ochraceous toment- 
um; nodes distinctly amulate with usually a corky layer and 
denser tomentum; principal internodes 4.5-16.5 cm. long; — 
leaves drooping, deciduous, decussate-opposite [or ternate?], 
short-petiolate or subsessile with a clasping base; petioles 
very stout, flattened above, rounded beneath, 1-5 (usually 
2.5-5) cm. long, more or less margined or alate, densely 
ochraceous-furfuraceous, or occasionally obsolete; blades 
firmly membranous or subcoriaceous, very dark green and nit- 
id above, very much lighter and not nitid beneath, broadly 
elliptic, 11-85 cm. long (to almost 1 m. long on sprouts; 
mostly 23-55 cm. long), 6-50 cm. wide (mostly 22-37 cm. 
wide), acute or short-acuminate at apex, entire or repand- 
denticulate, abruptly acute or long-acuminate at base and 
prolonged into the alate petiole or clasping at base, dense- 
ly squamose and rugose (or even bullate) above, becoming 
glabrate and smooth, densely tomentose, furfuraceous-tomen- 
tose, or pubescent beneath with ochraceous, yellowish, red- 
dish, or brownish hairs and densely resinous-punctate, the 
tomentum sometimes appearing stellate; midrib very stout, 
somewhat prominent above, very coarsely so beneath; second- 
aries slender, 9-15 per side, ascending, quite straight and 
but little arcuate, arcuately joined at the margins, promin- 
ulent above, prominent beneath; vein and veinlet reticula- 
tion very abundant, the tertiaries numerous, issuing at 
right angles to the secondaries, parallel, usually conspic- 
uous above and prominulent beneath; inflorescence axillary 
(in the uppermost axils only) and terminal, paniculate, mas- 
sive, the terminal panicles often several feet long and 2 


1935 Moldenke, The Genus Tectona 157 


feet or more wide (mostly about 40 cm. long and 35 cm. 
wide), with distant, opposite, widely divaricate, decus- 
sate, many-branched, many-flowered cymes, densely furfura- 
ceous with cinereous or ochraceous tomentum throughout; pe- 
duncles and sympodia continuous with the twigs and similar 
in texture, color, and pubescence, often elongate; pedicels 
stoutish or slender, 1-4 mm. long, furfuraceous; bracts 
large and foliaceous, 2 subtending each pair of cymes, re- 
sembling the leaves in all respects but smaller; bractlets 
numerous, lanceolate-linear, to 15 mm. long and 4 mm. wide 
at base, sessile, ochraceous-furfuraceous, attenuate at a- 
pex; prophylla oblong or linear-lanceolate, to 5 mm. long 
and 1 mm. wide, densely furfuraceous; calyx campanulate, 
light yellow or light green, 3-4.5 mm. long in all, 3-3.5 
(-77) mm. wide, densely furfuraceous—-tomentose, its rim 5- 
7-toothed or lobed, the teeth ovate or ovate-oblong, 1.5- 
2.5 mm. long, often reflexed, blunt or obtuse; corolla 

white (or rosy on the lobes), short-hypocrateriform, glab- 
rous outside and within, its tube broadly cylindric, 1.5-3 
mm. long, about 1.5 mn. wide, its limb 5-7-parted, its lobes 
obovate-elliptic, 2.54 mm. long, about 2.5 mm. wide, round- 
ed at apex, overlapping, erect or reflexed; stamens 5 or 6, 
inserted about 1.3 mm. below the mouth of the corolla-tube 
(or lower), equal, somewhat exserted; filaments white, fili- 
form, 2.5-4 mm. long, glabrous, ampliate and flattened be- 
low; anthers yellow, ovate or oblong, about 1 mm. long and 
0.5 mm. wide, dorsifixed near the base; style yellowish- 
white, slender, 3.6-5.2 mm. long, glabrate or more or less 
pubescent with branched hairs; stigma yellowish-white, bi- 
fid, its branches 0.2-0.5 mm. long; ovary ovate or conical, 
1.5-2 mm. long, about 1.5 mm wide, densely pubescent, dirty 
white or rosy when fresh, 4-celled, each cell l-ovulate; 
fruiting-calyx greatly enlarged and inflated, to 2.5 cm. 
long and wide, glabrate, panyraceous, light brow and brit- 
tle when dry, mostly irregularly plaited or crumpled and 
bladder-like; fruit subglobose or more or less tetrezonally 
flattened, to 1.5 cm. long and wide, densely tomentose with 
irregularly branched, light brown or ochraceous hairs, umb- 
ilicate and 4-lobed at apex, 4-seeded (rarely 1-3-seeded by 
abortion), the exocarp thin, somewhat fleshy when fresh, 
densely pubescent or tomentose, the endocarp thick, bony, 
corrugated, 4-celled, with a small central lumen between the 
cells; seeds oily. 
This species is the very important teak tree whose wood 
is the most valuable of all knowm timbers. It is undoubtedly 
the most important and most valuable member of the entire 
Verbenaceae from the point of view of human economics and 
trade, commanding a higher price on the marlzet than any tim- 
‘ber excent mahogany. It is a native of and is widely distrib- 


158 PHYTOLOGIA No. 4 


uted in tropical and southern Asia and the Malayan region, 
being especially abundant in the mixed forests of India, 
Burma, Siam, and Malaya. Its numbers have been greatly re- 
duced in India and Burma by cutting for the timber, but it 
is now being maintained in special plantations by the gov- 
ernment in numerous suitable parts of the British domain. It, 
has been introduced and become naturalized in the Philipp- 
ines, Java, and the Fiji Islands, as well as in parts of the 
New World. In India its northern limit is 24°40' on the 
western side of the Aravalli hills and 25°30' N. lat. in the 
central part near Jhansi. In Burma it extends to 25°30', In 
Bengal it is not indigenous, but plentations have been made 
in Assam north to the 27th parallel. In Punjab it is culti- 
vated north to the 32nd parallel. 

The tree attains a height of 120-150 feet and a girth of 
20-25 feet in its native habitat. Its trunk is straight and 
often buttressed. It is unmistakable because of its tremend- 
ously large leaves and inflorescences, its clasping-based 
bractlets and prophylla, and inflated fruiting calyces. 
According to Standley and others, the crown of the tree is 
often very large and spreading, especially if it stands a- 
lone. The timber is yellowish-browm in color, straight- 
grained, and easily worked. When once seasoned it does not 
warp or crack. It is very hard and strong and owing to the 
presence of an aromatic resinous oil is extremely durable. 
It has no equal for use in tropical climates and is vastly 
preferred in temperate climates for certain purposess It is 
the most valuable wood known for ship building, being espec- 
ially preferred for armored vessels, since it does not, like 
oak, corrode the iron. It is exported in large quantities to 
Great Britain and other countries for use in the building of 
ships and railway carriages. No satisfactory substitute has 
ever been found. In Burma the oil is extracted from the wood 
and used medicinally and also as a substitute for linseed 
oil and varnish. A tar, used medicinally, is distilled from 
it, and the leaves afford a red dye used in Malabar for col- 
oring silks and cotton. The large leaves are used as plates, 
for thatching, and to wrap parcels by native Burmese. 

The first good figure and description of this species wer 
given by Rheede in his Hortus Malabaricus, vol. 4, p. 57, te — 
27, in 1683. Perhaps the best modern illustration of it ap- 
pears in Brandis, Forest Flora of North-West and Central In- 
dia, vol. 3, t. 44 (1874). The svecies requires a dry trop- 
ical climate and thrives best in India where in the summer 
it receives the heavy rainfall from the southwestern monsoon 
and the winter is nearly rainless. In regions of a mean an=- 
nual rainfall of 5O inches it thrives best; where the mean 
is only 30 inches teak is more scarce. The most favorable 
mean annual temperature is between 75° and 81° F. The tree 


1935 Moldenke, The Genus Tectona 159 


is absent near the coast and inland the most valuable for- 
ests are situated on low hills up to 3000 feet in altitude. 
It can inhabit a variety of soils, but perfect drainage or a 
dry subsoil is the one indispensable condition. Irregularly 
shaped stems are produced when drainage is poor. 

In the dry season the tree is leafless, the leaves fall- 
ing off in January in hot regions or remaining on the tree 
until March in moister climates. When the monsoon begins 
fresh leaves are produced. It is said that on water-sprouts 
the leaf-blades may become 2-3 feet long and resemble tobac- 
co leaves. They are said to be whorled occasionally when the 
tree is young and sometimes subglabrous beneath, but I won- 
der if these statements do not perhaps apply to another spe- 
cies, T. Hamiltoniana. The inflorescences unfold naturally 
during the rains, usually in July or August in the tree's 
native haunts, and the seeds ripen the following January or 
February. Herbarium specimens in anthesis, however, have 
been examined which were collected in February, Jume, July, 
August, September, October, and November, and in fruit in 
every month of the year except December. The tree can be 
distinguished from a distance in the rainy season by its 
large panicles of white flowers held above the leaves and in 
the dry season by its feathery fruit-bearing panicles. Many 
seeds are produced and sown every year by a mature tree of 
this species, but the numerous and devastating forest fires 
of the dry season in March and April after the seeds have 
fallen, greatly impede its spread. Heavy rains wash the 
seeds down the hillsides and thus seedlings and young trees 
may occasionally be found also in the valleys. Seed germina- 
tion is very untertain since so much moisture is needed. 
When teak grows with bamboo, as it often does, the young te- 
ak seedlings are mostly smothered out by the bamboo. 

The bark of the trunk is about 1/2 inch thick, gray or 
brownish-gray. The sapwood is white, the heartwood golden- 
yellow, with a pleasant, strong, aromatic odor. When season- 

ed it darkens into brown, mottled with darker streaks. The 
odor is retained to a great age. Its extraordinary durabil- 
ity is its chief value. In India beams of teak have been 
found in buildings several centuries old -- in many cases 
more than 1000 years old. Some instances have been reported 
where temple beams were more than 2000 years old and still 
in perfect condition! It is still used on ocean liners, al- 
though iron is now employed generally for war vessels. It is 
also extensively used for furniture, door and window frames, 
and railway carriages. Termites eat the sapwood, but rarely 
the heartwood. The teredo bores into it and therefore metal 
sheaths are used to protect teak piles on wharves. It does 
not split, shrink, crack, or warp when once seasoned. It 
takes a beautiful polish, is easily worked, is not heavy, 


160 PHY EO LAO Gk No. 4 


and has great elasticity and strength. It weighs 38-46 lbs. 

per cubic foot, SO cubic feet weighing approximately a ton. 

When it is green it is heavier than water and will not 

float. In-order to float the timber dowm streams to the sea- 

ports it must be well seasoned first. This is accomplished 

by girdling -- making a deep circular cut through the bark 

and sapwood to the heartwood. A tree thus girdled dies and 

is allowed to stand 1 or 2 years. The wind and sun complete- 

ly season the wood of such trees in that time, after which 

they are felled and floated to the seaports. It is not true 

that teak trees are tapped for oil before being felled in 

Burma. Several varieties of teak are distinguished in In- 

dia, Burma, and Java by the color and texture of the wood, 

but these are not important distinctions in the timber 

trade. Soil, elevation, climate, and density of forest in- 

fluence the grain of the wood. An isolated teak tree has 

more side branches and therefore more knotty and heavier 

wood. Fires may scorch or even destroy the bark of young 

trees, causing an irregular crack or hollow in the center 

of large logs, where decay may commence. The growth of the 

tree (in plantations) is in youth very rapid -- a 2-year 

old seedling is 5-10 feet tall. In 15 years the trees are 

60 feet tall with a girth, breast high, of 19 inches. When 

80 years old the girth is 72 inches and this is a market- 

able age. Natural timber grows more slowly because of ad- 

verse environmental conditions. A natural forest tree with a 

trunk 24 inches in diameter is 100-200 years old. The spec- 

ies naturally does not grow in pure stands. It is always 

associated with bamboo and other trees. India uses most of 

the teak grown there and the demand is greater there than 

the supply. It is therefore exported from Burma and some- 

what also from Java and Bangkok. Rangoon is where it is 

mostly used for ship building -- European vessels were form 

erly built here. From 1901 to 1907 an average of 55,994 cub- 

ic tons of teak wood was exported from British India per 

year. In India now government conservation management con- 

trols all teak tracts and a permanent and increasing supply 

is assured, 
Common and vernacular names for the species are of course 

numerous. In Sanskrit it is "g&ka" or "saka", in Arabic and ~ 

Persian "saj", in Mahratti "sag", in Hindu "sa&gin" and "sag- © 

wan", in the Dravidian languages "teka" (Malayan "tekka", : 

Tamil "tekku"), in Portuguese "teke" or “teca", and in Eng- 

lish "teak". In Cuba it is called "teca" and in Guadeloupe 

"bois de tek". In the Philippines it is known as "dalanang", ~ 

"dalandon", "dati", "“djéti", "hadlayéti", "héti", "kalaydéti} 

"sagunyéti", "téca", and "yéti". In parts of Malaya it is P 

called "iattie"” or "djattie". The so-called "African teak" 

is Oldfieldia africana Benth. & Hook. f., "bastard teak" in © 


1935 Moldenke, The Genus Tectona 161 


the East Indies is Pterocarpus Marsupium Roxb. and in Bengal 
is Butea frondosa Roxb., "Ben teak" is Lagerstroemia micro- 
carpa Wight {although this name is also used for low grades 
of true teak], "New Zealand teak" is Vitex littoralis A. 
Cunn. (a tree 50-60 feet tall, yieZding hard fissile timber 
indestructible under water), "white teak" is Flindersia Ox- 
leyana F. Muell., the "teak" of Queensland is Dissitaria ba- 
loghioides F, Muell., and the "teak" or "teakwood" of New 
South Wales is Endiandra glauca R. Br. 

The species is widely cultivated in plantations for tim- 
ber and also as scattered specimens for ornament or because 
of the great interest attached to this tree. It tends to es- 
cape from cultivation and become naturalized in favorable 
_ climates, Lam & Bakhuizen [Bull. Jard. Bot. Suitenz. III, 3: 

28. 1921] report its cultivation in Delhi. Hallier f. [Med- 
ed. Rijks Herb. Leiden 37: 34. 1918] records it as cultiva- 
ted in Java and Ceylon and cites his C. 34a and C.34b from 
Java and C.34c from Ceylon. E. D. Merrill (Enum. Philip. Pl. 
3: 389. 1923] states that it is cultivated in various parts 
of the Philippines, especially in the Sulu Archivelago and 
in parts of Mindanao and that it is now established in the 
southern part of the archipelago. He also notes that "the 
tree at Tanay, Rizal Province, Luzon, from wnich Blanco se- 
cured specimens before 1837 still exists tnere". The same 
author [Enum. Born. Pl. 512. 1921] records it as cultivat- 
ed in Borneo, while Setchell [Univ. Calif. Publ. Bot. 12:: 
204. 1926] found it cultivated in the Society Islends. Tne 
following list of citations will prove how widely it is cul- 
tivated in tropical parts of the New World. In addition to 
these records, Grey & Hubbard [List Pl. Atkins Instit. 195. 
1933] record the species as cultivated in Santa Clara, Cuba, 
from stock received from Vilmorin-Andrieux & Cie. in 1927, 
collected by Barbour, Dorsett, & Higgins, 1928-1929. It is 
there called "teak" or "teakwood". 

The species has become naturalized in at least two places 
in America -- on the islands of Martinique and St. Vincent — 
and probably in other localities not yet recorded. Hahn, who 
collected it an Martinique, found it in the "vallée de S. 
Pierre", blooming in July. G. We. & H. H. Smith, who collect- 
ed it on St. Vincent, say that here it is "a large tree" 
growing in "second growth spurs of Mt. St. Andrews, to 330 
Me, On the western side. The species may nave been introduc- 
ed, but now grows wild in this locality". Fruit was observed 
and collected in May. On another label these collectors say 
that the trees here are about 60 feet tall already in nearby 
Belaire Valley. 

One hundred and thirty-one herbarium specimens and 15 
mounted photosraphs have been examine’. 

Citations: MARTINIQUE: Hahn 1275 (Cb, Cb, D, G, K, L, S, 


162 PY 7 OL OG Bt No. 4 


S). ST. VINCENT: Smith & Smith 1273 (B, N-photo, Z-photo), 
1573 (K). CULTIVATED: Cuba: Santa Clara: Jack 8118 (A), 8174 

A, A, N, N, W, W, W, W); Roig, Acufia, & Malberti s.n. (Herb. 
Roig 6145] (Es). Jamaica: G. N. Collins 126 [photo 6851] 

W); H. A. Lang 616 (D)# St. Croix: T. Thomson 973 (N).* St. 
Kitts: Britton & Cowell 416 (B, N). Montserrat: Ryan s.n. 
(Hort. Rohr] (S). Guadeloupe: R. P. Duss 3786 (E, F, L, N, 
N, N-photo, W, Z-photo). Martinique: Bélanger 1145 (Cb, Ob, 
Cb)*; Collector undesignated s.n. (V, V, V, V)*; Re P. Duss 
sen. (B, B, N-photo, Z-photo); Kohaut [Sieber] s.n. (Cp); De 
S. Martin s.n. [Ex India occid.] (Cb, Cb)*. Barbados: Bailey 
& Bailey 339 (Ba, N-photo, Z-photo); Dash sen. [Bot. Stat. 
Herb. Barbados 578] (F, N, N-photo, W, Z-photo)*. St. Vin- 
cent: Guilding s.n. (K). Grenada: Arnott s.n. (K). Trinidad: 
W. E. Broadway sen. [July; 1907] (N), sen. [Sept. 4, 1916] 
(Ba, D, E, F, G, N-photo, W, Z-photo), sen. [Trin. Bot. Gard 
Herb. 9691] (R); Pinder sen. [Trin. Bot. Gard. Herb. 3362] 
(R, R); Te Thomson 375 (K); Whitford s.n. [April, 1920] (¥); 
Wrbna s.n. [Sieber Fl. Trinit. 27] (B, B, B, B, B, E, K, V, 


V)*. Panama: Canal Zone: Bailey & Bailey 466 (Ba); Johansen 
50 (W)*; P. ©. Standley 26,900 (W, W); Wetmore & Abbe 22 
(A, A). British Guiana: British Guiana Bot. Gard. mie 
uary, 1892] (U, U). French Guiana: Poiteau s.n. [Cayenne] 
(Cb, Cb, K). Brazil: Rio de Janeiro: Bailey & Bailey 43 
(Ba); Guillemin 668 (K, P); A. Richard s.n. [Rio de Janeiro] 
(K); Warming ssn. [Cult. in Hort. Bot.] (Cp); Weddell 425 
[Environs de Rio de Janeiro, 1843] (N, N), 425 [Rio de Jan- 
eiro, 1844] (N, N); Whitford 2 (E, F. G, N, S, W, Y). China: 
Kwangtung: Tsiang 2 (N). India: Bengal: Kurz sen. [Cult. 
Hort. Bot. Calcutt.] (N); Wallich 772 (N); Punjab: R. R. 
Stewart 1107 (N). Ceylon: Baker 126 (Ca, N, N). Philippines: 


Luzon: Merrill Sp. Blanc. 837 (N). Java: Kuntze 5334 (N, N). 


Geographic distribution of the species of Tectona in the 

New World and in cultivation: 

Martinique, St. Vincent: 

T. grandis 

Cultivated: 

T. grandis [Cuba (Santa Clara), Jamaica, St. Croix, St. 
Kitts, Montserrat, Guadeloupe, Martinique, Barba- 
dos, St. Vincent, Grenada, Trinidad, Panama (Canal 
Zone), British Guiana, French Guiana, Brazil (Rio 
de Janeiro), China (Kwangtung), India (Bengal, 
Punjab), Ceylon, Philippines (Luzon), Borneo, Java, 
Society Islands, Fiji Islands] 


Alphabetized list of citations: 
Arnott s.n. (grandis) 
Bailey & Bailey 43 (grandis), 339 (grandis), 466 (grand- 


K 
4 
oY 


1935 Moldenke, The Genus Tectona 163 


is); Baker 126 (grandis); Bélanger 1145 (grandis); Bot. 
Stat. Herb. Barbados 578 (grandis); British Guiana Bot. 
Gard. sen. (January, 1892] (grandis); Britton & Cowell 
416 (grandis); Broadway, W. E., sen. [Sept. 4, 1916] 
(grandis), sen. (July, 1907] (grandis), sen. (Trin. 
Bot. Gard. Herb. 9691] (grandis) 

Collins, G. N., 126 [photo 6851] (grandis); Collector un- 
designated sen. (grandis) 

Dash sen. [Bot. Stat. Herb. Barbados 578] (grandis); 
Duss, R. P., 3786 (grandis), sen. (grandis) 

Guilding s.n. (grandis); Guillemin 668 (grandis) 

Hahn 1275 (grandis); Herb. Roig 6145 (grandis) 

Jack 8118 (grandis), 8174 (grandis); Johansen 50 (grandis) 

Kohaut ‘s.n. [Sieber] (grandis); Kuntze 5334 (grandis); 
Kurz sen. [Cult. Hort. Bot. Calcutt.] (grandis) 

Lang, H. Ae, 616 (grandis) | 

Martin, D. S., sen. [Ex India occid.] (grandis); Merrill 
Sp. Blanc. 837 (grandis) 

Pinder sen. (Trin. Bot. Gard. Herb. 8362] (grandis); 
Poiteau s.n. [Cayenne] (grandis) 

Richard, A., sen. [Rio de Janeiro] (grandis); Roig, Ac- 
ufia, & Malberti sen. [Herb. Roig 6145] (grandis); Ryan 
sen. [Hort. Rohr] (grandis) 

Sieber s.n. (Kohaut] (grandis), Fl. Trinit. 27 (grandis); 
Smith & Smith 1273 (grandis), 1573 (grandis); Standley, 
P. C., 26,900 (grandis); Stewart, R. R., 1107 (grandis) 

Thomson, T., 375 (grandis), 973 (grandis); Trin. Bot. 
Gard. Herb. 8362 (grandis), 9691 (grandis); Tsiang 3322 
(grandis) 

Wallich 772 (grandis); Warming sen. [Cult. in Hort. Bot.] 
(grandis); Weddell 425 [Environs de Rio de Janeiro, 
1845] (grandis), 425 [Rio de Janeiro, 1844] (grandis); 
Wetmore & Abbe 227 (grandis); Whitford 2 (grandis), 
sen. [April, 1920] (grandis); Wrbna sen. [Sieber Fl. 
Trinit. 27] (grandis) 


(a) This generic name is sometimes misspelled "Tectonia" or 
"Tektona". 

(b) This generic name is sometimes accredited to Rheede, who 
first used it in his Hortus Malabaricus, vol. 4, p. 57; 
a e7, An 1685, 

(c) This binomial is sometimes accredited to E. D. Merrill 
because he first validated it by publication of a des- 
crintion in Philip. Journ. Sci. Bot. 5: 227 (1910); how- 
ever, in this reference Merrill distinctly accredits the 
name to Bentham & Hooker f., who first effectively pub- 
lished it as a nomen nudum in their Gen. Pl. 2: 1152 in 
1876. Inasmuch as an author has the right when publishing 


164 PHY $04 0°96 is No. 4 


a new name to accredit it to whomsoever he wishes, it be- 
ing assumed that he has a valid reason for so accrediting 
it, I see no reason why tnis name should not continue to 
stand as Tectona philippinensis Benth. & Hook. f. 

(d) This pre-Linnaean name is often cited merely as “Iatus". 

(e) This binomial is often erroneously accredited to Linnae- 
us the elder or even to Roxburgh. 

(f) This binomial is sometimes erroneously accredited to Lin 
naeus [vid., Fr. de Montholou, Notices sur 1'Indie, p. 


60. 1837]. 


* The specimens followed by an asterisk in this list of cit- 
ations do not definitely state on their labels that they 
were collected from cultivated material, but since the 
svecies is more widely cultivated tnan naturalized in Am- 
erica, the present monographer is assuming that all New 
World specimens not definitely marked as having been tak- 
en from wild plants were from cultivated material. On the 
other hand, in regard to Old World material (i.e., tropi- 
cal Asiatic), all specimens not definitely marked as hav- 
ing been taken from cultivated meterial ere assumed to 
heve come from wild plants. 


AN AID IN BOTANICAL DRAWING 


E. H. Walker 


When taxonomists are asked why they do not publish draw- 
ings of their new species, they usually reply that they have 
no talent in drawing. The writer some years ago was faced 
with the necessity of making fifty or more drawings to ill- 
ustrate a tree manual (1), and having no more skill in draw- 
ing than the average systematist, he devised the mechanical 
aid here pictured. This device was subsequently used in pre- 
paring illustrations to accompany the descriptions of vari+ 
ous new species of plents. Such a drawing can easily be made 
from an herbarium specimen by placing it on a sheet of glass 
over an electric light in a semi-darkened: room with a sheet 
of drawing paper over the specimen. The light casts a sharp 
shadow on the drawing paper, the outline of which can be 
lightly traced, Frequently more veins show than are visible 
in the specimen seen by reflected light. The outline with 
the visible features having been prepared, the drawing is 
removed and additionel details are freely sketched. The 
drawing is finally inked. If illustrations of dissected 
flowers or other parts are to be made, the dissection is 
placed on a millimeter cross-lined glass or paper under a 
lens or binocular microscope. The drawing is made on cross- 
ruled paper, the lines of whith are as many millimeters a- 
part as is the desired scale of enlargement. 

The apparatus here pictured (fig. 15) can be made from 
two shelf-brackets, a strip of lath, two screw hooks, and a 
sheet of glass of any size and shape. Almost any desk light 
can be adjusted to go under the gless; or daylight can be 
used, the apparatus being placed before a window in a semi- 
darkened room. 

The advantages of accurate line-drawings over half-tone 
reproductions of photographs of type specimens are usually 
beyond question, especially in respect to cost. The adop- 
tion of a simple mechanical aid like this should greatly 
reduce the number of newly described species, the identity 
of which is dimmed by the gloom of a tecnnical description. 


The apparatus may be used equally well in drawing from 
photographs and need not be confined to taxonomic illustra- 


' 


: 


ft 
d 


(1) Fifty-one common ornamental trees of the Lingnan Univ- 
ersity campus. Lingnan Sci. Journ. 6: 1-166. 1 pl. 56 
figs. 1928 (issued 1930). 

165 


we © oes a 


oe ir eS S 


- 


—— 


NOMENCLATURAL AND TAXONOMIC NOTES 


Harold N. Moldenke 


Study of newly received material in tne herbarium of the 
New York Botanical Garden has revealed the necessity for the 
following new scientific names and combinations and has dis- 
closed two species new to science. 


ALOYSIA LIGUSTRINA var. PARAGUARIENSIS (Briq.) Moldenke, 


comb. nov. Lippia lipustrina var. paraguariensis Briq., Ann. 
Conserv. & Jard. Bot. Genev. 7-8: 305. 1904, 


ALOYSIA URUGUAYENSIS Moldenke, nom. nov. Lippia affinis 
Briq.e, Bull. Herb. Boiss. 4: 339. 1896 [not L. affinis 
Schau. in DC. Prodr. 11: 576. 1847]. 


BENTHAMIDIA NUTTALLII (Audubon) Moldenke, comb. nov. 
Cornus Nuttallii Audubon, Birds of Am. t. 367 (1837); T. & 


G. Fl. N. Am. 1: 652. 1840, 


CHASCANUM RARIFLORUM (A. Terrac.) Moldenke, comb. nov. 
Hebenstreitia rariflora A. Terrac., Bull. Soc. Bot. Ital. 
1892: 424. 1892. 


CLERODENDRUM KAEMPFERI var. ALBUM (P'ei) Moldenke, comb. 
nov. Clerodendron japonicum var. album P'ei, Mem. Sci. Soc. 
China 1 (3): 144. 1932. 


CROTON. LAMARCKIANUS Moldenke, nom. nov. Croton Cascarilla 
Lam. Encycl. 2: 203. 1786 [not C. Cascarilla L. Sp. Pl., ed. 
2, 1424, 1763]. 


DISPORUM SCHAFFNERI Moldenke, sp. nov. lierba usque ad 5 
dm. alta brachiata; rhizomate luteo usque ad 5 cm. Longo; 
radiculis numerosis in crassitudine longitudineque subuni- 
formibus; caule glabro nitido, parte non subterranea lineis 
minutis numerosis tenuissimis purpureis vel nigris notata, 
dichotomo, ad nodos majores paullo turgido et 2 stipulas 
plusminus vaginatas gerenti; ramulis ultimis plusminus pub- 
escentibus, pilis brevibus (quam 1 m, brevioribus) sub an- 
gulo 90° e ramulis abeuntibus; stipulis lanceolatis, 3-4.5 
cm. longis, ad basin vaginatam ca. 8 mm. latis, sensim usque 
ad apicem acutum vel acuminatum angustatis, intezris, paullo 
puberulentibus vel subglabratis; internodiis primariis 5.5-8 
cm. longis; nodis annulatis; foliis alternis, 2 supremis in 


_quoque ramulo approximatis; petiolis ca. 1 mm. longis vel 


167 


168 Pa PE OLe Ge Le No. 4 


obsoletis, in sectione triangulo-ovatis, supra complanatis 
sulcatisque, subtus 4—-7-striatis; laminis ellipticis vel 
elliptico-ovatis leviter membranaceis, supra atroviridibus, 
subtus valde pallidis pernitidisque, 8-11 cm. longis, 3-5.2 
cm. latis, integris, ad basin rotundatis vel subacutis et 
plerumque inaequilateralibus, ad apicem longe acuminatis, 
supra glabris nitidulisque, subtus in venis majoribus par- 
cissime piloso-pubescentibus; venis primariis palmatis 4-6 e 
basi laminae emergentibus, in textura costae consimilibus, 
paene ad apicem arcuato-adscendentibus, in crassitudine sub- 
uniformibus (apice excluso); venulis numerosis sub angulo 
90° e venis primariis costaque abeuntibus; floribus 1-3; 
fructibus 1 vel 2, ad apicem ramorum dispositis; pedicel- 
lis fructiferis gracilibus 11-17 mm. longis, ad apicem gen- 
iculatis, ubique densiuscule hirsuto-pubescentibus, pilis e- 
rectis rigidis sub angulo 90° e pedicello abeuntibus; fruct- 
ibus juventute pyriformibus, senectute plusminus complana- 
tis, 7-9 mm. longis, 8-10 mm. latis, valde 2- vel 3-lobatis, 
densissime stellato-tomentosis, pilis flavis longitudine 
variis, radio centrali quam lateralibus duplo vel triplo 
longiore; seminibus glabris. 

The type of this species was collected by Delzie Demaree 
(No. 10,749) on wooded north hillsides in the C. C. C. Camp 
Gordon at Friendship, Shawnee State Forest, Scioto Coe, 
Ohio, June 21, 1934, and is deposited in the herbarium of 
the New York Botanical Garden. The species is remarkable for 
its decidedly lobed and densely stellate-tomentose fruits. 
On D. lanuginosum (Michx.) Nichols., a species common 
throughout the eastern states and also collected by Demaree 
(No. 10,748) at the same locality on the same date, the 
fruits are glabrous and nitid or rarely marked with here and 
there a scattered, simple, very short, and obscure hair. On 
the western North American D. oreganum (S. Wats.) W. Miller, 
of British Columbia, Washington, Oregon, Idaho, and Montana, 
tne fruits are puberulent with simple nairs and the leaves 
are decidedly ovate and cordate-clasping at the base. On De 
Smithii (Hook.) Piper, of California, Oregon, and Washing- 
ton, and on D. Hooxeri (Torr.) Nichols., of California, the 
leaves are perfectly glabrous beneath, as also are: the un- 
lobed fruits. On D. trachycarpum (S. Wats.) Benth. & Hook. 
the fruits are densely squamulose. On D. trachyandrum 
(Torr.) Britton, of Oregon and California, the unlobed 
fruits are also glabrous and nitid. D. parvifolium (S. 
Wats.) Britton, of Oregon, has very much smaller leaves and 
is very much more dwarf in stature, with very short inter- 
nodes. 

Our species is most closely related to D. maculatum 
(Buckl.) Britton, a species of the mountains of North Caro- 


4 


lina, Tennessee, Kentucky, and Georgia and recently found in © 


. 
: 
: 

, 
; 
j 
& 
« 
~ 

; 


1935 Moldenke, Taxonomic Notes 169 


abundance by Demaree (No. 10,597) at Camp Gordon along with 
D. lanuginosum and D. Schaffneri. On the twelve specimens of 
D. maculatum in the herbarium of the New York Botanical Gar- 
den, however, the leaves are decidedly chartaceous and frag- 
ile and noticeably smaller (3-8 cm. long and 1.2-3.8 cm. 
wide), more acute at the base, very sparsely puberulent or 
strigillose with scattered hairs above and more densely pub- 
erulent with soft hairs beneath, the stems are not regularly 
marked with purple linear dashes, and the ovaries vary from 
subglabrate to pubescent with long whitish hairs of irregul- 
ar size, but not at all stellate and not at all uniformly 
distributed over the surface; in fact, the hairs are mostly 
borne in 2 or 3 bands on the ovary, with the intervening 
portions subglabrate. 

The speciés is named in honor of Prof. John Henry. 
Schaffner of Ohio State University, who sent the type spec- 
imen to me with a description of its distinguishing charac- 
teristics and the suggestion that it probably represented a 
new species. Flowering specimens of D. Schaffneri and fruit- 
ing specimens of D. maculatum from Ohio, preferably from the 
type locality of D. Schaffneri, are needed in order to set- 
tle definitely the relationship between these two species. 
D. Cahnae Farwell, from Michigan, is also closely related to 
these species, but only a flowering topotype of this species 
has thus far been available for study, on which the ovary, 
while pubescent, was not stellate. 

D. Schaffneri has also been collected by Arthur R. Harper 
at Churn Creek, Adams Co., Ohio, on July 15, 1928. 


GHINIA JUNCEA (Schau.) Moldenke, comb. nov. Tamonea jun- 
cea Schau. in Mart. Fl. Bras. 9: 177. 1851. 


GHINIA SPICATA (Aubl.) Moldenke, comb. nov. Tamonea spic- 
ata Aubl. Pl. Guian. 2: 660, t. 268. 1775. 


HELENIUM DENTICULATUM (Nutt.) Moldenke, combs nov. Lepto- 
oda denticulata Nutt., Trans. Am. Phil. Soc. II, 7: 573. 
1841. Tne name Helenium decurrens published by myself for 
this plant in Bull. Torrey Club 62: 230 (1935) is invalid 
because it is a homonym of H. decurrens Vatke, Ind. Sem. 
Hort. Berol. Appe (1875). 


HUMULUS SCANDENS var. VARIZGATUS (F. Roem.) Moldenke, 
comb. nov. Humulus japonicus var. variegatus F. Roem., Am. 
Florist 8: 489, 1892; Gartenfl. 42: 19, t. 4 & 5. 1893. The 
binomial, Humulus scandens, has been very recently proposed 
for this common species of Asiatic hop by E. D. Merrill 
(Trans. Am. Philos. Soc. 24 (2): 138. 1935] on the basis of 
an older name of Loureiro. 


170 PHYTOLOGIA No. 4 


LANTANA ARISTATA var. ANGUSTIFOLIA (Kuntze) Moldenke, 
comb. nov. Lippia aristata var. angustifolia Kuntze, Rev. 
Gen. Pl. 3: 251. 1898. 


LANTANA ARISTATA var. PLURIFEDUNCULATA (Kuntze) Moldenke, 
stat. nov. Lippia aristata f. pluripedunculata Kuntze, Rev. 
Gen. Pl. 3: 251. 1898. 


PHRAGMITES MAXIMUS var. VARIEGATUS (A. S. Hitche.) Mol- 
denke, comb. nov. Phragmites communis var. variegata A. 5S. 
Hitche. in Bailey, Stand. Cycl. Hort. 5: 2601. 1916. The 
name Phragmites has been considered masculine by many 
authors and feminine by as many other authors. Trinius' or- 
iginal description of the genus gives no hint as to what he 
considered to be its gender -- his only species there pub- 
lished being P. communis. Steudel, however, accredits four 
or five other specific combinations to Trinius "in mss." and 
these are given with the masculine ending. Dioscorides, from 
whom Linnaeus took the name Phragmites in his Arundo Phrag- 
mites, according to classical Greek lexicons considered the 
word as masculine. The binomial Phragmites maxims has re- 
cently been proposed for this common plant by Cniovenda 
[Nuov. Giorn. Bot. Ital. 26: 80. 1919] on the basis of an 
older binomial of Forsk@l [cf., Merrill, Trans. Am. Philos. 
Soce 24 (2): 19% 1935]. 


PHYLA NODIFLORA var. PUSILLA (Briq.) Moldenke, comb. nov. 
Lippia nodiflora var. pusilla Briq., Arkiv Bot. 2, no. 10: 
19. 1904. “ 


STACHYTARFHETA MINIACEA lioldenke, sp. nov. Herba peren- 
nis; ramis ranulisque puberulentibus plusminus obtuse tet- 
ragonis, juventute brunneis, senectute stramineis; foliis 
decussato-opvpositis; petiolis onsoletis; laminis membranac-— 
eis oblenceolatis, 5-10.5 cm. longis, 2-5.5 cm. latis, ad 
apicem abrupte acutis, ad basin integram longe (1.5-4.5 cm. ) 
cuneatis, ultra basin grosse serrato-dentato, supra sparsis- 
sime strigillosis (pilis late dispersis brevissimis saepe 
non conspicuis), subtus dense (sed non nanifesto) puberulen- 
tibus (pilis paullo majoribus in costa et in venis secundar- 
iis); venis secundariis arcuatis utroque ca. 4, non directe 
in dentes excurrentibus; spicis usque ad 36 cm. longis, ca. 
1 cm. latis, rhachide senectute leviter excavato, bracteis 
semper late patentibus; corolla minia. 

The type of this svecies was collected by R. S. Felly 
(No. 14) along roadsides and in clearings in acache bush 
country at the Freshwater Creek Reserve, British Honduras, 
in February, 1933, anc is deposited in the herbarium of the 
Royal Botanic Vardens at Kew. The collector describes the 


1955 Moldenke, The Genus Recordia 171 
plant as four feet tall, with vermillion flowers. 


VERBENA PATAGONICA Molcenke, nom. nov. Verbena bonari- 
ensis Rendle, Journ. Bot. 42: 370. 1904 [not V. bonarien- 
sis L. Cpe Pl. 20. L731 


A MONOGRAPH OF THE GENUS RECORDIA 


Harold N. Moldenke 


The following is the third in my series of monographic 
studies of the genera of Verbenaceae and Avicenniaceae. In 
the list of citetions of herbarium specimens the following 
abbreviations of the names of herbaria are employed: B = 
Botanisches IMuseum, Berlin; E = Missouri Botanical Garden, 
St. Louis; F = Field Museum of Natural History, Chicago; G = 
Gray Herbarium of Harvard University, Cambridge, Mass.; K = 
Royal Botanic Gardens, Kew; N = New York Botanical Garden, 
New York City; S = Naturhistoriska Riksmuseet, Stockholm; 

V = Naturhistorisches Museum, Vienna; X = Herbier Boissier, 
Geneve; and Z = H. N. Moldenke Herbarium, Watchung, N. Je 

To the directors and curators of the above-mentioned herbar- 
ia the writer extends his most sincere thanks for their 
courtesy and kindness in allowing him to study their mater- 
ial of this genus and for their continuous and very generous 
cooperation throughout the progress of this work. All speci- 
mens so studied have been annotated with uniform printed 
annotation labels and mention is made on each label that the 
specimen is cited in this monograph. All material thus far 
received from these ten herbaria is herein accounted for and 
cited. Forty-three other herbaria have been canvassed, but 
did not contain any material of this group. 


RECORDIA Moldenke, Phytoloria 1: 99. 1934. 

Shrubs or trees; leaves deciduous, opvosite, petiolate, 
serrate; inflorescence racemose, terminal, many-flowered; 
flowers hypogynous, zyfomornnic; calyx gamosepalous, tubu- 
lar-camnanulate, irregular, cleft on the lower (abaxial) 
side, S-ribbed, the 3 upper ribs terminating in very short 
apiculations, the 2 lower ribs not ending in apiculations or 
only very obsolete ones; corolla -amopetalous, hypocrateri- 
form, irregular, its tube obconic, not greatly curvate, its 
limb 5-parted, 2-lipped, its lobes spreading, the 2 upper- 
most (adaxial) smallest, the 2 lateral ones medium-sized, 
and the lowest (abaxial) one largest; fertile stamens 4, di- 


172 PHYTOLOGIA No. 4 


dynamous, inserted in the corolla-tube below its mouth, the 
2 lowest being the longer, the fifth stamen reduced to a 
filiform anantherous staminode; anthers sagittate, dorsi- 
fixed, 2-celled, the thecae divergent at the base and open- 
ing by longitudinal slits; pistil single; style simple, ter- 
minal, curvate at apex, the convex arch flattened and stig- 
matiferous; ovary fusiform, compound, suverior, abruptly 
nerrowed into the style-base at apex, 2-celled, each cell l- 
ovulate. 

Only one species, the type of the genus, is known thus 
far, although more may be discovered when South America is 
more thoroughly explored botanically. The genus is named in 
honor of Prof. Samuel James Record, professor of forest pro- 
ducts at Yale University, whose admirable and painstaking 
researches into the vast field of comparative wood anatony 
have contributed so much of inestimable value to the science 
of botany and whose generous cooperation, freely given, has 
proved invaluable so many times to his taxonomic colleagues. 

Although structurally a member of Briquet's Verbenoideae 
Petraeeae, the genus Recordia greatly. resembles in habit and 
general appearance the genus Citharexylum. Very superficial- 
ly the genus also resembles the rubiaceous genus Alseis, es= 
pecially in habit, but the epigynous flowers of the latter 
genus of course at once distinguish it from ours. The genus 


Recordia, insofar as it is now known, is confined to Bolivia, 


ee Moldenke, Phytologia 1: 99-101, fig. 13. 
1994. 

Shrub or tree; branches and branchlets rather stoutish, 
subterete, grayish, glabrous; twigs and immature shoots 
brown and densely cano-pubescent with short hairs; nodes an- 
nulate; leaf-scars sessile, not borne on sterigmata; inter- 
nodes abbreviated, 5-16 mm. long; leaves decussate-opposite, 
mostly clustered at the tips of young twigs; petioles rather 
slender, 3-12 mm. long, densely short-pubescent; blades 
thinly membranous or chartaceous, dark green, blackening in 
drying, elliptic, 5.2-10.5 cm. long, 2.2-4.4 cm. wide, 
short-acuminate, sharply but minutely serrate almost to the 
middle or subentire, abruptly acute or short-cuneate at the 
base, not glanduliferous, sparsely strigillose or puberulent 
above (or sometimes glabrate), densely short-pubescent or 
occasionally subvelutinous beneath, especially on the larger 
veins and when immature; midrib slender, flat or impressed 
above, prominent beneath; secondaries 5-7 per side, ascend- 
ing, often not conspicuously arcuate, anastomosing at the 
margins; veinlet reticulation slender, often obscure; ra- 
cemes erect, 7-13 cm. long and to 2.5 cm. wide, very densely 
many-flowered, terminating branchlets and axillary twigs; 
peduncle slender, abbreviated, 5-8 mm. long, densely cano- 


1935 Moldenke, The Genus Recordia 173 


pubescent; rachis slender, densely cano-pubescent; pedicels 
very slender, 2-5.5 mm. long, densely pubescent; prophylla 
minute, setaceous, mostly inconspicuous; flowers mostly 
borne in 2-, 3-, or many-flowered fascicles along the rach- 
is, very numerous; calyx tubular-campanulate, about 4.9 mn. 
long and 2.5 mm. wide at apex, rather densely pubescent on 
the outside, glabrous within, fissured on the lower (abax- 
ial) side to almost 1.8 mm., S-ribbed, the 3 upper ribs ter- 
minating in very short (about 0.2-0.3 mm. long) apicula- 
tions (the central rib often slightly larger than the two 
lateral ones), the 2 lower ribs not terminating in apicula- 
tions or in very obsolete ones; corolla zygomorphic, hypo- 
crateriform, not noticeably curvate, its tube obconic, about 
6.5 mm. long and 0.5 mm. wide at base, ampliate to 3.6 mm. 
just below the limb, glabrous outside, slightly short-pubes- 
cent in the throat within, its limb S-parted, its lobes 
lightly puberulent outside, short-pubescent within, the 2 
uppermost (adaxial) lobes smallest, ovate-lingulate, about 1 
mm. long and 1.5 mm. wide at base, rounded at apex, the 2 
lateral lobes medium in size, triangular-ovate, 2-3 mm. 

long and 2-2.8 mm. wide at base, rounded at apex, the lowest 
(abaxial) lobe triangular-ovate, about 3.5 mm. long and 3 
mm. wide at base; fertile stamens 4,. didynamous, inserted 
‘about 2 mm. below the mouth of the edie erhantens the 2 lower 
ones largest (with filaments about 3.2 mm. long), the 2 lat- 
eral ones shorter (with filaments about 1.2 mm. long), the 
fifth stamen reduced to a filiform anantherous staminode a- 
bout 1 mm. long and inserted about 1 mm. below the fertile 
stamens; anthers sagittate, dorsifixed, about 1 mm. long, 
the thecae divergent at base; style simple, terminal, about 
7-1 mm. long, glabrous, narrowed toward the apex, at the a- 
pex laterally bent, with the ovate convex upper surface 
flattened and stigmatiferous; ovary fusiform, about 1.5 mm. 
long and 0.5 mm. in diameter, at the apex abruptly attenuate 
into the style-base, glabrous, 2-celled, each cell l-ovul- 
ate; ovules attached in the axil of the exterior dissepiment 
about at the center of the ovary; fruiting-calyx and fruit 
not seen. 

The type of this species, which is the type species of 
the genus, was collected by José Steinbach (No. 7240) at 
Bafado, Rio Surutu, Sara, Santa Cruz, Bolivia, at an alti- 
tude of about 400 m., on October 1, 1925, and is deposited 
- the herbarium of the Naturhistoriska Riksmuseet at Stock- 
holm. The species ascends to 1600 m. according to Troll, and 
seems to be endemic to the mountains of Santa Cruz, Bolivia. 
Seventeen specimens, including the type, and three photo- 
graphs have been examined, 

- Citations: BOLIVIA: Santa Cruz: Steinbach 7240 (B-iso- 
type, B-isotype, E-isotype, F-isotype, G-isotype, N-fragment 


174 PH DAT Orb: Ors & No. 4 


& photo of type, S-type, Z-photo of type), 7296 (B, E, F, G, 
S); Troll 969 (B); Werdermann 2707 (S); Province undetermin- 
ed: Bridges sen. [1846] (K), sen. [1850] (X); Quming 152 
(V). ILLUSTRATIONS: Phytologia l: fig. 13. 19 N). 


Georraphic distribution of the species of Recordia: 
Bolivia: 
R. boliviana [Santa Cruz] 


Alphabetized list of citations: 
Bridges sen. [1846] (boliviana), s.n. [1850] (boliviana) 
Cuming 152 (boliviana) 
Steinbach 7240 (boliviana TYPE), 7296 (boliviana) 
Troll 969 (boliviana) 
Werdermann 2707 (boliviana) 


TWO NEW MELASTOMES OF THE KRUKOFF COLLECTION 


H. A. Gleason 


HORMOCALYX Gl. gen. nov. 

Melastomaceae Miconieae: Flores S-meri axillares; hypan- 
thium campanulatum setosum; calycis tubus prolongatus erect- 
us truncatus, dentibus exterioribus in annulum herbaceum 
horizontaliter patentem coalitis; petala 5 inaequilatera ob- 
ovata magna; stamina 10 isomorpha, intheris crasse linearib- 
us, poro terminali dehiscentibus, connectivo paulo elevato 
Simplici; ovarium inferum 2-loculare, summo plano glabro; 
stylus rectus glaber, stigmate semigloboso. Frutex hirsutus 
anisophyllus; folia formicariis ad basin laminae ornata. 


HORMOCALYX HIRSUTUS Gl. sp. nov. Frutex 5 m. altus; ram- 
uli subteretes juveniles hirsuti demm glabri; folia con- 
ferta subsessilia in quoque jugo valde inaequalia; petioli 
2-5 mm. longi; laminae majores oblongae vel ellipticae, 8-14 
cm. longrae, 4-7 com. latae, breviter acuminateae, integrae, e, 
ciliatae, ad basin paulo angustatae et cordulatae, S-nervi- — 
ae, supre breviter sparseque flavido-setulosae, subtus ad. 
venas setosae vel setulosae ceterum slabrae; formicariae di- 
dyma semi-ovoidea hirsuta, 8-15 mm. longa, in basi lLiminae 
et infra basin plusminus pendentia; laminae minores sine 
formicariis cordato-ovatae, 2-5 cm. longae et fere totidem 
latae, jugo venarum exteriore in auriculas curvato; flores 
in axillis foliorum minorum solitarii subsessiles; hypanthi- © 
um campanulatum herbaceum, ad torum 4 mm. longum, sparse 


1959 Gleason, Two New Melastomes . 175 


glanduloso-hirsutum; calycis tubus erectus truncatus hirsut= 
us, 2-2.5 mm. longus, pilis glandulosis 2-4 mm. longis pat- 
entibus; dentes exteriores in annulum fere 2 mm, latum hir- 
sutum coaliti, pilis 5 marginalibus quam aliis multo valid- 
ioribus; petala ut videtur purpurea obovata retusa glabra, 
11-12 mm. longa, latere breviore truncato, longiore obtuso; 
filamenta complanata glabra 6.5-7 mm. longa; antherae fere 
rectae 5.3 mm. longae; ovuli in quoque loculo circ. 12; sty- 
lus 9-10 mm. longus, stigmate 1.3 mm. diametro. 

Type, Krukoff 7098, collected in "campinarana alta" on 
the plateau between the Livramento and Ipixuna rivers, Ama- 
zonas, Brazil, and deposited in the Britton Herbarium. Tne 
stems and bases of the leaves are mostly covered with the 
black debris used by ants to shelter their runways. This and 
the numerous short leafy axillary branches make it often 
difficult to discern the character of the leaf-bases or the 
position of the flowers. 

The distinctive feature of the genus is, of course, the 
remarkable development of the exterior calyx-teeth, but to 
this feature may be added the solitary sessile axillary 
flowers, the 2-celled ovary, and the small number of ovules. 

Notwithstanding the axillary flowers, the affinity of the 
genus is distinctly with Tococa, as shown by the structure 
of the anthers. These agree almost completely with those of 
Tococa in shane, connective, and general apnearance, but are 
not incurved at the summit and have a strictly terminal 
pore. Axillary flowers are known in a few srecies of Tococa 
also. In inflorescence the genus is reminiscent of Maieta, 
but differs notably in its stamens and hypanthium. In gener- 
al aspect it suggests certein myrmecopnilous species of Cli- 
demia, but avain differs in the anthers and h:nanthiun. 


CLIDEMIA FEROX Gl. sp. nov. Sect. Stanhidium: frutex 5 m. 
altus ubique hirsutus, pilis fuscis vel stramineis basi in- 
crassatis curvatim patentibus 4-8 mm. longis non glandulos- 
is; folie aequalia vel subaequalia; petioli validi 1-2 cn. 
longi; laminae late ellipticae, 10-20 cm. longae, S-11 cm. 
latae, abrupte breviterque acuminatae vel subcaudatee, in- 


_tegrae, ciliatae, basi rotundatae, S-nerviae, venis primari- 
is supra leviter impressis subtus elevatis, secundariis sup- 
Ya obscuris subtus vrominulis, 5-8 mm. dissitis, sub anzulo 
_fere recto divergentibus; flores 6-meri in capitula sessilia 
axillaria pauci densissime conferti; hypanthium complanatum 
10 mm. longum 6-nervium, pilis saepissime versus nervos cur- 
_vatis et lineas 6 longitudinales formantibus, his lineis in 
_siccis vix manifestis sed in aqua maceratis conspicuis; to- 


rus ad basin staminum breviter denseque pilosus; sepala fere 


ad basin distincta, patentia, late oblonza, 3.2 mm. longa, 
obtusa, intus densissime sericea, dentibus exterioribus ter- 


176 | PHYTOLOGIA No. 4 


minalibus tuberculiformibus a pilis obtectis; petala late et 
inaequaliter obovata, 10 mm. longa, 9 mm. lata, ad apicem 
subretusam paucisetosa, ad margines tenuia et saepe fimbria- 
tula; filamenta glabra leviter complanata 6 mm. longa; anth- 
erae compressae lineares 6 mm. longae, poro terminali dehis- 
centes, connective basi brevissime 2-lobato; ovarium inferum 
6-loculare, ovulis numerosis; stylus rectus 15 mm. longus, 
in dimidio inferiore dense tomentosus et faucem hnypanthii 
occludens, pilis basi 4 mm. longis gradatim decrescentibus; 
stigma truncatum. 

' . Type, Krukoff 7262, collected on the shore of the Rio 
Ipixuna between Monte Christo and Santa Victoria, Amazonas, 
Brazil, and deposited in the Britton Herbarium. The extra- 
ordinarily large flowers in very dense clusters give the 
plant a superficial aspect resembling Myrmidone or Myria- 
spora. The number of the ovules and the character of the 
stamens and stigma distinguish it at once from these genera 
and show its position in Clidemia. There is apparently only 
one species in this large genus with which it may be associ- 
ated, C. longisetosa Hoehne, from the Tapajoz River. That 
species has flowers of only half the size, a glabrous style, — 
and a 2-celled ovary. 


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PHY TOLOGIA 


Designed to expedite botanical publication 


Vou. 1 February, 1937 No. 5 


CONTENTS 


C. L. Lunpert: Chamekia, a new Genus in the Lauraceae... . .. 177 


H. N. Moxpenkxe: Additional Notes on the Genus Aegiphila .. . . 182 


ee 


Published by H. A. Gleason and Harold N. Moldenke 
The New York Botanical Garden 


i , ) Bronx Park, New York, N. Y. 


Price of this number: 50 cents; per volume $5.00 in advance. 
Vol, 1, number 4, was issued September 19, 1935 


oe 


‘en walt Bre 


EW YORK 

sSOTANICAL 

GARDEN 
CHANEKIA, A NEW GENUS IN THE LAURACEAE (a) 


C. L. Lundell 


[ In the identification of collections of Lauraceae obtain- 
ed by the Carnegie-Michigan expeditions to the Yucatan Pen- 
insula, and in studying other material from Mexico and Cent- 
ral America, certain species have been found which apparent- 
ly represent an undescribed genus. In the course of the 


fy) 


4 
} 


Field Museum, Arnold Arboretum, and the University of Calif- 
-ornia were examined. 


CHANEKIA Lundell, gen. nov. (b) 


Panniculae axillares vel pseudoterminales; flores in ped- 
-unculorum apice subumbellati vel racemosi, hermaphroditi; 
perstanthii tubus infundibuliformis vel subconicus; Limbi 
segmenta 6, subaequalia, inflexa; staminodia cyclorum I, iL; 
et IV nulla; stamina cycli III fertilia, tubum circum gynae- 
ceum formantia; filamenta crassa, biglandulosa; antherae 2- 
eellulae, extrorsae; ovarium glabrum; hypanthium accrescens, 
dupliciter marginatum; bacca ellipsoidea vel 


leaves alternate; panicles axillary or pseudo- 
the peduncles solitary or rarely fasciculate; 
flowers subumbellate or racemose, perfect; hypanthium con- 
picuous, infundibuliform or subconical; perianth 6-parted, 
ihe segments inflexed, small, unequal, the 3 outer segments 
Argest, enclosing the androecium except for a small apical 
iperture; stamens of series I, II, end IV absent; stamino- 
dia none; the 3 stamens of series III fertile, forming a 
tube surrounding the gynaeceum; filaments thick, biglandu- 
lar; staminal tube obtuse-pyramidal, short; anthers 2- 
celled, extrorse; pores apical, semi-exserted; ovary glab- 
fous; style semi-exserted from the staminal tube or includ- 
Bd; accrescent hypanthium cupuler, double-margined; berry 
@llipsoid or ovoid. 
_ Type species, Chanekia campechiana (Standl.) Lundell. 
_ Ghanekia is intermediate between Acrodiclidium and Mis- 
Bnteca. In Acrodiclidium the stamens of series I and II are 
‘educed to foliaceous staminodia, whereas in Chanekia stam- 
inodia are absent. In Misanteca the anthers are exserted 
Prom the perianth, the stamens of series III are connate to 
he apex, and the staminal tube is obtuse-cylindric, as 
jontrasted with the included anthers, partially connate 
tamens, and obtuse-pyramidal staminal tube in Chanekia. 


177 


178 PHYTOLOGIA No. 5 


Because the floral characters were not given in the orig- © 
inal descriptions, Chanekia campechiana, C. mexicana, and C. 


misantlae are here redescribed. 


CHANEKIA CAMPECHIANA (Standl.) Lundell, comb. nov. 
Ocotea campechiana Standl., Carnegie Inst. Wash. Publ. 
G61: 56. 1935. 


Misanteca campechiana (Standl.) Lundell, Carnegie Inst. 
Wash. Publ. 78: = 1937. 


A tree, height 8 to 25 m., diameter 35 to 45 cm.; branch- ; 


lets minutely sericeous, canescent; petioles 4 to 10 m. 
long, minutely sericeous; leaf-blades oblong-lanceolate or 
linear-lanceolate, 4 to 11 cm. long, 0.8 to 3 cm. wide, apex 
long caudate-acuminate, the acumen obtusish, base acute, 
coriaceous, minutely sericeous above, glabrate with age, 
minutely sericeous beneath; costa elevated, lateral nerves 
inconspicuous; inflorescence axillary, paniculate, 2 to 5.5 
cm. long (including the peduncles), the peduncles solitary, 
minutely sericeous; flowers subumbellate; pedicels 1 to 2.5 
mm. long, minutely sericeous; hypanthium obovoid-conical, 2 
to 2.3 mm. long, about 2 mm. in diameter, fleshy, minutely 
sericeous; perianth 6-parted, the segments squamiform, 
thick, subequal, depressed triangular-ovate, 0.9 to 1.5 m. 


wide, acutish, hairy inside; staminodia of series I, II, and 


IV absent; the 3 stamens of series III fertile, connate at 


base, forming a tube around the gynaeceum, pilose inside and — 
at base outside; anthers 2-celled, extrorse; the pores api- 
cal, semi-exserted from the perianth; glands about equalling 


one-third the length of the stamen, squamiform, connate lat- 
erally at base with those of adjacent stamens; ovary glab- 


rous, l-celled; style glabrous, included or shortly exsert-— 


ed from the apex of the staminal tube. 
Specimens examined: MEXICO: Campeche: Tuxpefa, Lundell 
1295, type collection; Lundell 1580. GUATEMALA: Department 


of Petén: La Libertad, Lundell 3065, 3359, 3409; vernacular @ 
names "“ektit", "dzol". Uaxactun, Bartlett 12339; vernacular ~ 


name "copal-chi". 
The species also occurs in British Honduras. 


CHANEKIA PECKII (I. M. Johnston) Lundell, comb. nov. 
Misanteca Peckii I. M. Johnston, Contr. Gray Herb. 70: 
70. 1924, 


BRITISH HONDURAS: Without locality, Peck 826, type (Gray _ 


Herbarium). 


The writer has not seen the type, but the species is un- 


doubtedly referable to Chanekia. 


CHANEKIA CAUDATA Lundell, sp. nov. 
Arbor, 7-metralis; petiolis 4--7 mm longis, albido- 


1937 Lundell, The Genus Chanekia 179 


pubescentibus; folia lanceolato-oblonga, 5--9.5 cm. longa, 
1.6--3.2 om. lata, apice caudato-acuminata, basi subacuta, 
adulta glaberrima, chartacea; inflorescentia axillaris, sub- 
umbellata-racemosa, 2--3.5 om. longa (pedunculo incluso), 
pauciflora; pedicellis 3.5--5.5 mm. longis, glabris; flores 
albi, hypanthio subgloboso-turbinato, 1.8--2 mm. longo, 2.2 
mm. diametro, carneo, glabro; perianthium 6-partitum, seg- 
mentis squamiformibus, crassis, subaequalibus, ca. 1.1 mm. 
latis; staminodia cyclorum I, II, et IV nulla; staminia cyc- 
li III fertilia, basi connata tubum pilosum circum gynaeceun 
formantia, antheris 2-cellulis, extrorsis; glandulae incon- 
spicuae, squamiformes; ovarium glabrum, l-cellulum, stylus 
glaber, breviter ex tubi apice staminalis exsertus; fructus 
ignotus. 

A small tree, height 7 m., diameter 7.5 cm.; branchlets 
slender, densely pubescent, grayish; petioles 4 to 7 m. 
long, canaliculate, pubescent with grayish hairs; leaf- 
blades lanceolate-oblong, 5 to 9.5 cm. long, 1.6 to 3.2 cm. 
wide, apex long caudate-acuminate, the acumen narrow, ob- 
tusish, oblique, base acutish, glabrous above except for a 
few hairs along the costa, glabrate beneath or with few 
scattered appressed hairs, costa prominent, lateral nerves 
inconspicuous, finely reticulate-veined, chartaceous; in- 
florescence axillary, subumbellate-racemose, 2 to 3.5 cm. 
long (including peduncles), the peduncles solitary, 3- to 
6-flowered, the flowers white, drying brownish-black; pedi- 
cels slender, 5.5 to 5.5 mm. long, glabrous; hypanthium 
subglobose-turbinate, 1.8 to 2 mm. long, 2.2 mm. in diamet- 
er, fleshy, glabrous; perianth 6-parted, the segments 
squamiform, thick, equal or nearly so, ovate-orbicular, 
about 1.1 mm. wide, acutish, hairy inside; staminodia of 
series I, II, and IV absent; the 3 stamens of series III 
fertile, connate at base, forming a short tube around the 
gynaeceum, hairy inside and at base outside; anthers 2- 
celled, extrorse, the pores apical; glands small, squami- 
form; ovary glabrous, l-celled; style glabrous, shortly ex- 
serted from apex of staninal tube. 

Type in the herbarium of the University of Michigan, C. 
L. Lundell 6183, collected in advanced forest on limestone 
along the Arenal-Valentin road, El Cayo District, British 
Honduras, June 21, 1936. 

Additional specimens examined: BRITISH HONDURAS: Toledo 
District: Camp 32, British Honduras-Guatemala boundary sur- 


vey, Schipp 1279. 


CHANEKIA CORIACEA Lundell, sp. nov. 

Arbor, 10-metralis, glabra; petiolis 6--10 mm. longis; 
folia lanceolata vel oblongo-lanceolata, 5.5--11 cm. longa, 
1.9--3.8 cm. lata, apice acuminata, basi subacuta, coriac- 


180 PHYTOLOGTIA No. 5 


ea; infructescentiae axillares, pedunculis 1--3 cm. longis; 
hypanthium accrescens, cupulare, dupliciter marginatunm, 
1.6--2.3 cm. diam., ca. 1 cm. longum; bacca ellipsoidea, ca. 
1.7 cm. longa, 1.2 cm. diam. 

A tree, height 10 m., diameter 22 om., entirely glabrous; 
petioles 6 to 10 mm. long; leaf-blades lanceolate or oblong- 
lanceolate, 5.5 to 11 cm. long, 1.9 to 3.8 cm. wide, apex 
acuminate, the acumen obtusish, base acutish, coriaceous, 
costa prominent, lateral nerves inconspicuous; infructes- 
cence axillary, bearing a single fruit; peduncle 1 to 4 cm. 
long, thick; the accrescent hypanthium very shallow, verru- 
cose, cupular, double-margined, 1.6 to 2.3 cm. in diameter, 
about 1 cm. long; berry ellipsoid, about 1.7 cm. long, 1.2 
cm. in diameter, abruptly short-apiculate. 

Type in the herbarium of the University of Michigan, W. 
A. Schipp 1282, collected at Camp 31, British Honduras- 
Guatemala boundary survey, altitude 700 nm. 

The herbarium label bears the following data: "Small 
tree in shady valley, also on hill top in open places; 
flowers white; fruits black with red pedicel. Occasional." 


CHANEKIA MISANTLAE (Brand.) Lundell, comb. nov. 

Acrodiclidium misantlae T. S. Brandegee, Univ. Calif. 

Bot. 6: 497. 1919. : 

A small tree, entirely glabrous; branchlets, peduncles, 
and pétioles drying black; petioles 5 to 14 mm. long; leaf- | 
blades elliptic or narrowly elliptic-obovate, 4.5 to 12.5 
cm. long, 2 to 4.8 cm. wide, apex obtuse, base ouneate, 
subcoriaceous, costa elevated beneath, lateral nerves in- 
conspicuous, finely but prominently reticulate-veined; in- 
florescence axillary, 1.5 to 35.5 cm. long (including ped- 
uncles), the peduncles solitary, very slender 1- to 3- 
flowered, the flowers drying brownish-black; pedicels slen- 
der, 3.5 to 7 mm. long; hypanthium subglobose-turbinate, 2 
to 2.2 mm. long, 2.5 to 5.6 mm. in diameter, fleshy; peri- 
anth shallowly 6-parted, the segments squamiform, thick, 
subequal, depressed-orbicular, 1.1 to 1.9 mm. wide, round- 
ed or obtuse-apiculate; staminodia of series I, II (7), 
and IV absent; the 3 stamens of series III fertile, con- 
nate at base, forming a short tube around the gynaeceun, 
the tube glabrous inside and outside; anthers 2-celled, ex- 
trorse, the pores apical; ovary glabrous, l-celled; style 
glabrous, apparently included in the staminal tube; cupule 
double-margined, 9 mm. in diameter 

Specimen examined: MEXICO: Vera Cruz: Misantla, Purpus 
8145, type (Herb. Univ. Calif. no. 200935). 

Because of the few flowers present on the type, only 
two were dissected, The character of the glands was not de- 
terminable from the material available. 


1937 Lundell, The Genus Chanekia 181 


CHANEKIA MEXICANA (Brand.) Lundell, comb. nov. 

Acrodiclidium mexicanum T. S. Brandegee, Univ. Calif. 

Bot. 6: 497. 1919. 

A small tree; branchlets, peduncles, and petioles densely 
fulvous-pubescent; petioles thick, 3 to 9 mm. long; leaf- 
blades lanceolate, 4 to 14 cm. long, 2 to 5 cm. wide, apex 
acute or acuminate, the acumen sometimes obtuéish, base ob- 
tuse or acutish, bromish-green above and glabrescent with 
age, paler and pilose beneath, subcoriaceous, costa and lat- 
eral nerves slightly impressed above, prominent beneath; in- 
florescence axillary or subterminal, paniculate, 2 to 5.5 
om. long (including peduncles), the peduncles solitary; 
flowers subumbellate, numerous; pedicels 2 to 3 mm. long, 
glabrous; hypanthium subconical, 2 to 2.2 mm. long, about 
1.5 mm. in diameter, fleshy, glabrous; perianth shallowly 
6-parted, the segments squamiform, thick, subequal, depres- 
sed triangular-ovate, 0.8 to 1 mm. wide; staminodia of 
series I, II, and IV absent; the 4 stamens of series III 
fertile, connate at base, forming a tube around the gynae- 
ceum, pilose inside and at base outside; anthers 2-celled, 
extrorse; the pores apical, semi-exserted from the peri- 
anth; glands one-third the length of the stamen, small, 
squamiform; ovary glabrous, l-celled; style glabrous, in- 
cluded or shortly exserted from the apex of the staminal 
tube. 

Specimens examined: MEXICO: Vera Cruz: Zacuapan, Purpus 
8081, type (Herb. Univ. Calif. no. 200978); Purpus 14335. 


(a) Contribution from the Herbarium of the University of 


Michigan. 
(b) The genus is named for Mercedes Chanek, the Maya plant 


collector. 


ADDITIONAL NOTES ON THE GENUS AEGIPHILA -- I 


Harold N. Moldenke 


Since the publication of my monograph of the genus 
Aegiphila in 1934 (1) considerable additional information 
has come to light and a large number of additional speci- 
mens and photographs have been examined. In addition to the 
abbreviations for the names of herbaria listed on pages 249 
and 250 of the above-mentioned work, the following are em- 
ployed in the present contribution: Ba = L. H. Bailey Herba- 
rium, Cornell University, Ithaca, N. Y.; Bb = Barbados Muse~ 
um and Historical Society, Bridgetown, Barbados; Bc # Barn- 
ard College, New York City; Bg = Bergens Museum, Bergen, 
Norway; Br = Jardin Botanique de 1'Etat, Brussels; Bu = W. 
M. Buswell Herbarium, University of Miami, Miami, Florida; 
Ch = Carey Herbarium, Royal Botanic Gardens, Kew; Cp = Uni- 
versitetets Botaniske Museum, Copenhagen; Dc = De Candolle 
Herbarium, Conservatoire et Jardin Botaniques, Geneva; Dr = 
Botanisches Institut, Dresden; Ed = Royal Botanic Garden, 
Edinburgh; Hb = H. Bassler Herbarium, N. Y. Botanical Gard- 
en, New York City; He = W. G. Herter Herbarium, Montevideo, 
Uruguay; I = Langlois Herbarium, Catholic University of Am- 
erica, Washington; J = Brooklyn Botanic Garden, Brooklyn, N. 
Y.; Le = Rijksherbarium, Leiden; Ls = Linnean Herbariun, 
Linnean Society, London; Ifu = Botanisches Museum, Munich; Ol 
= Universitetets Botaniske Museum, Oslo; Os = Osborn Botani- 
cal Laboratory, Yale University, New Haven, Conn.; Ru = Rut- 
gers University, New Brunswick, N. J.; Th = Thunberg Herbar- 
ium, Botaniska Institutionen, Uppsala; Us = Botaniska Insti- 
tutionen, Uppsala; Ve = Museo Comercial de Venezuela, Carac- 
as; Vu = Botunisches Institut der Universitat, Vienna; and Z 
= H. N. Moldenke Herbarium, Watchung, N. J. All specimens 
hereinafter cited have been annotated with printed and type- 
writteh annotation labels, except those of the Linnean Herb- 
arium. 

In addition to the numerous variations in the spelling of 
the generic name noted on page 250, there have recently come 
to light "Egiphila", "Aegyphilla", and “Aeguephila". It is 
also worth noting that some specimens of Cornutia pubescens 
Gaertn. f. have in the past been confused with Aegiphila, 
the Hahn 920 distributed as "Aegiphila sp." is Bourreria 
ovata L., and the Pohl 2143 distributed as "Aegiphila sp." 
is actually Siparuna guianensis Aubl. In the description and 
discussion of the genus as a whole on pages 251--261 it 
should be noted that as to habit A. cordata is one of the 
most beautiful species, A. Deppeana is normally a tree, but 

182 


Pin 


(1937 Moldenke, Notes on Aegiphila 183 


tends at times to become subscandent, and A. graveolens and 
the so-called "salutaris" form of A. mollis exude a fetid or 
nauseous odor. In regard to habitat, A. A. Her zogii prefers tne 
open pampas, A. Smithii and A. vitelliniflora have been col- 
lected in swamps and marshes, while A. graveolens prefers 
dry hills and fields. As to elevation, A. bogotensis ascends 
to 3660 m. in Colombia, A. ferruginea to 3200 m. in Ecuador, 
and A. Killipii to 35000 m. in Colombia. A. anomala, A. obo- 
vata, and A. ternifolia have very fragrant flowers, while 
those of A. Hassleri are narcotic-odorous after sundown. A. 
Hassleri, A. bracteolosa, A. Candelabrum, and especially A. 
_breviflora have often 2 or 3 distinct types of flowers in 
the same cyme or at least on the same branchlet. The flowers 
of A. Smithii are extremely small for the genus. The branch- 
es of the terminal pyramidal thyrsi in this genus are always 
_decussate-opposite. In A. graveolens and A. plicata the 
leaves are densely impressed glandular-punctate beneath. In 
regard to corolla, the red-purple ones of A. cordata and A. 
_vVillosissima are noteworthy; those of A. Smithii are espec- 
-ially small in size, while those of A. ~ bogotensis and A. 
multiflora are especially large. A. Smithii also has especi- 
ally small calyxes, while those of A. bogotensis are ex- 
tremely large. A. anomala has the largest calyxes in the 
genus. The fruit of A. bogotensis is umbonate at apex and 
that of A. Hassleri is edible. The fact that the fruit of 
this last-mentioned species is edible should be noted under 
the economic uses of the genus, as well as the fact that A. 
_Deppeana and A. Luschnathi have been introduced into culti- 
vation.- Throughout the monograph "Santander do Norte" should 
read "Santander Norte" sand “Santander do Sur" should read 
"Santander Sur" (although, actually, the official titles of 
the Colombian departments in question are “Norte de Santan- 
der" and "Santander" ) -- the "Sur" being here added to dis- 
tinguish this department more readily from Santander Norte. 
The number of contributors to our knowledge of the group 
“should be 503 and the number of publications reviewed 191. A 
list of the several new species and varieties which have re- 
cently come to light and their position in the artificial 
key to species and varieties (pages 265--280), together with 
other corrections to this key, will be published later. In 
order to facilitate reference and increase the usefulness of 
these and all future supplementary notes, the species and 
verieties will be arranged in alphabetic instead of taxonom- 
ic sequence, but each name will be preceded by the number of 
the species or variety according to the taxonomic sequence 
proposed in the original monograph. Of extreme interest is 
the fact that there is no genus cover for Aegiphila in the 
Linnean Herbarium nor any specimen of A. martinicensis, the 
type species. (often erroneously accredited to him), but 


2 
* 


184 PHY: T0602 No. 5 


under the genus Clerodendrum sheet number 8 is A. elata and 
sheet number 9 is A. Deppeana, both regarded as unidentified 
species of Clerodendrum by Linnaeus. 


49a. AEGIPHILA ACULEIFERA Moldenke in Fedde, Repert. 37: 209 

~-210. 1934, 

Shrub; branchlets stoutish, tetragonal, deeply 4-sulcate 
to the apex, densely furfuraceous, abundantly armed with 
short and more or less recurved prickles about 1 mm. long; 
principal internodes 3--4.5 cm. long; leaves decussate-oppo- 
site; petioles stout, 1.8--2.5 cm. long, furfuraceousy 
slightly margined; blades thin-membranous, dark green above, 
somewhat lighter beneath, obovate-elliptic, often very un- 
date in drying, 18--21 cm. long, 7--9.5 cm. wide, acuminate 
at apex, regularly glandulose-apiculate along.the margins 
(with the apiculations 5--13 mm. apart), cuneate at base, 
more or less furfuraceous along the venation on both sur- 
faces (and sparingly or obsoletely on the lamina also); mid- 
rib stout, very prominent beneath and slightly so above 
(very broad, especially toward the base above! ); secondaries 
slender, 7--9 per side, prominent beneath, arcuate-ascend- 
ing, not anastonosing; vein and veinlet reticulation slen- 
der, prominent beneath, often undate in drying; inflorescen- 
ce supra-axillary; cymes solitary, opposite, to 4 cm. long 
and 5 cm. wide, pronouncedly bifurcate, copiously armed with 
recurved prickles (about 1 mm. long) throughout, very lax 
and loosely many-flowered; peduncles stoutish, copiously 
armed, 1--1.7 cm. long, furfuraceous; pedicels very slender, 
furfuraceous, 3--4 mm. long; prophylla minute; bracts and 
bractlets none; calyx obconic, about 7 mm. long and 4 m. 
wide at apex, densely pulverulent, sparsely verruculose, its 
margin 2-lipped, its lips about 2.1 mm. long, equal and 
truncate; corolla hypocrateriform, white, its tube cylin- 
dric, about 8.3 mm. long, glabrous outside, pubescent with- 
in, its lobes 4, oblong-lingulate, about 6.3 mm. long and 
3.1 m. wide, subacute; stamens 4, inserted about 5.6 mn. 
below the mouth of the corolla-tube, long-exserted; fila- 
ments filiform, about 10.8 mm. long, pilose toward the base; 
anthers oblong, about 1.8 mm. long and 0.7 mm. wide; pistil 
glabrous, included; style about 5.2 mm. long; stigma bifid, 
its branches about. 3.9 mm. long, parallel; ovary pyriforn, 
about 1 mm. long and 0.9 mm. wide at apex, obscurely 4- 
lobed, glabrous, 4~-celled; fruiting-calyx and fruit not seen — 

The type of this very remarkable species was collected by 
Friedrich Carl Lehmann (No. 8524) at La Conga, in the west- 
ern Andes of Popayan, at an altitude of 1800--2400 m., El 
Cauca, Colombia, and is deposited in the herbarium of the 
Field Museum of Natural History at Chicago. The collector 
notes that it blooms in June and July. The prickles which 


1937 Moldenke, Notes on Aegiphila 185 


are so abundant on the branchlets, twigs, and peduncles of 
this species characterize it well and separate it at once 
from every other known species in tne genus. The type col- 
lection is erroneously cited on page 349 of my monograph as 
A. novogranatensis, while the Stork collection is erroneous- 
ly cited on pages 352 and 476 as A. odontophylla. The Pitt- 
fer specimen was collected in forests along the Rfo Naranjo, 
altitude 200--250 m.; blooming in March. 

COSTA RICA: San José: H. Pittier 7584 (Br--2); Cartago: 
Stork 2230 (A--photo, B--photo, D--photo, F, G--photo, K-- 
photo, N--photo, P--photo, S--photo, W--photo, Z--photo). 
COLOMBIA: El Cauca: Lehmann 8524 (B--isotype, B--photo of 
type, F--type, K--isotype, K--photo of type, N--fragment of 
type, N--photo of type, S--photo of type, W--isotype, Z-- 
photo of type). 


3a. AEGIPHILA ALBA Moldenke, sp. nov. 

Arbor; ramis ramulisque crassiusculis argute tetragonis, 
ad nodos valde complanatis; foliorum cicatricibus magnis 
suberosis; petiolis crassis dense puberulis; laminis tenu- 
iter membranaceis late ellipticis obtusis vel subacutis in- 
tegris, ad basin attenuatis vel acuminatis, utrinque spars- 
issime puberulis, glabrescentibus; inflorescentiis axillar- 
ibus glomeratis dense multifloris; pedunculis nullis vel 
brevissimis; pedicellis sub fructu valde incrassatis; caly- 
ce fructifero valde incrassato coriaceo, obconico dense 
puberulente valde lenticellato. 

Tree, to 10 m. tall or taller; branches and branchlets 
rather stout, sharply tetragonal, light gray, rather dense- 
ly pulverulent or glabrate, conspicuously flattened and am- 
pliate at the nodes; nodes not annulate; principal inter- 
nodes 2--5 cm. long; leaf-scars large and corky, somewhat 
prominent, with an equally large scar just above on older 
wood left by the falling off of the inflorescences; leaves 
decussate-opposite, very lerge; petioles stout, 2--3 cm. 
long, flattened and canaliculate-margined above, convex 
beneath, densely puberulent with extremely minute grayish 
hairs, not noticeably ampliate at base; blades thin-memb- 
ranous, dark green above, much lighter beneath, broadly 
elliptic, 18--20 cm. long, 8.5--9.5 cm. wide, obtuse or 
subacute at apex, entire, attenuate or acuminate at base, 
very minutely puberulent on both surfaces, becoming sub- 
glabrate in age; midrib rather stout, flattened or sub- 
prominulent-rounded above, rounded-prominent beneath; sec- 
ondaries slender, 14--18 per side, arcuate-ascending, most- 
ly flat above, sharply prominent beneath,’ conspicuously 
arcuate-joined near the margins in many loops; vein and 
veinlet reticulation sparse, mostly obscure on both sur- 
faces or only the larger portions discernible; inflores- 


186 Pon TT Ob eee No. 5 


cence axillary, glomerate, borne in the axils of this seas- 
on's leaves or at the nodes of last’ season's growth, pract- 
ically surrounding the branchlets, the glomerules many- 
flowered; peduncles absent or to 1 or 2 mm. long, deciduous; 
pedicels very slender, 1--3 mm. long, puberulent, greatly 
incrassate in fruit; calyx heavy, leathery, obconic, 5.2-- 
5.5 mm. long, 4-5 m. wide, verruculose, glabrate, the rim 
2-lipped, the sinus about 1.3 mm. deep on one side and 2.4 
mm. deep on the other side, the lips truncate and equal or 
again split into 2 or 5 shorter and truncate teeth; corolla 
infundibular, white, glabrous, its tube slender, about 5 mm. 
long and 1 mm. wide, ampliate above, its limb 5S-parted, its 
lobes obovate, about 6 mm. long and 2.4 mm. wide, rounded at 
apex, venose; stamens 5, inserted about 2.4 mm. below the 
mouth of the corolla-tube, exserted; filaments filiforn, 
11--12 mm. long, glabrous; anthers narrowly oblong or lin- 
ear, 2.4--3 mm. long, auriculate at base, often twisted, 
dorsifixed at about 1/3 their length; style capillary, in- 
cluded, 2.4--43 mm. long, somewhat flattened, glabrous; stig- 
ma bifid, the branches erect, 1.6--35 mm. long, about O.7 m. 
wide; ovary very small, about 1 mm. long and wide, 4-lobed 
at apex, glabrous, 4-celled; fruiting-calyx very heavy and 
coriaceous, obconic, to 1 cm. long and wide, prominently 
lenticellate, densely puberulent; fruit not seen. 

The type of this species was collected by Ynes Mexia (No. 
6656) in second-growth woods near Quevedo, Canton Vinces, 
altitude about 50 n., Los Rfos, Ecuador, between October 22 
and November 6, 1934, and is deposited in the United States 
National Herbarium at Washington. Miss Mexia reports that 
the flowers are white and the tree an abundant one to 10 nm. 
tall, popularly called "lulu". Eggers, who first collected 
it (in fruit, December 31, 1891), describes it as “arbor 
altiss." and records the names "koit tree" and "tutumbo". 

ECUADOR: Guayas: Egvers s.n. [Balao, 31/12/1891] (B, N-- 
fragment); Los Rios: Mexia 6656 (N--isotype, N--fragment of 
type, W--type). 


6. AEGIPHILA ANOMALA Pittier. 

Addenda and errata to the description as published on 
pages 288 and 289: Internodes to 4 cm. long; leaf-scars to 7 
mm. long and 5 mm. wide; petioles to 2.5 om. long; blades 
membranous or subcoriaceous, to 35 cm. long and 13 cm. wide; 
calyx in anthesis to 6 mm. long and 4 mm. wide, puberulent 
or very short-pubescent, often 2-lipped; corolla large, ex- 
serted; fruiting-calyx very large when mature, obvolute and 
cucullate only when immature, to 19 mn. wide, often eventu- 
ally splitting on one side; fruit to 12 mm. long, umbonate 
with a light and corky umbo at the apex. 

The number of the type collection, cited on pages 289 and 


1937 Moldenke, Notes on Aegiphila 187 


475 as H. Pittier 16,711, seems on further investigation to 
be a herbarium number of the Instit. Pnysico-geogr. Nat. 
Costaric. rather than a Pittier number. 

Additional citations: COSTA RICA: Limén: H. Pittier s.n. 
(Herb. Instit. Physico-geogr. Nat. Costaric. 16,711] (A-- 
photo of type, B--photo of type, Cb--2 isotypes, D--photo of 


type, F--photo of type, G--photo of type, K--isotype, N-- 


fragment of isotype, N--photo of type, P--photo of type, S-- 
photo of type, W--photo of type, Z--photo of type); Stork 
2290 (F, N--fragment). 


2la. AEGIPHILA AUSTRALIS Moldenke, sp. nov. 

Frutex; ramulis obtuse tetragonis dense puberulis; horn- 
otinis subgracilibus tetragonis dense breviterque pubescent- 
ibus vel tomentellis; petiolis dense breviterque pubescenti- 
bus; laminis tenuiter membranaceis obovate-ellipticis brev- 
issime acuminatis integris, ad basin longe acuminatis, supra 
densiuscule breviterque pubescentibus et glanduloso~puncta- 
tis, subtus dense tomentellis; inflorescentiis axillaribus; 
cymis dense multifloris. 

Shrub; branchlets medium, obtusely tetragonal, grayish, 
densely puberulent, the leaf-scars usually elevated; twigs 
rather slender, tetragonal, densely short-pubescent or tom- 
entellous; nodes not annulate; principal internodes 1--6 cm. 
long; leaves decussate-opposite; petioles slender, 2--5 mn. 
long, densely short-pubescent; blades thin-membranous, dark 
green above, lighter beneath, obovate-elliptic, 3.5--10.4 
cm. long, 1.7--6.3 cm. wide, very short-acuminate at apex, 
entire, long-acuminate at base, rather densely short-pubes- 
cent and glandular-punctate above, densely tomentellous be- 
neath; midrib slender, slightly prominulent on both sur- 
faces, more so beneath, but mostly hidden by the dense pub- 
escence; secondaries slender, 5 or 6 per side, arcuate- 
ascending, not conspicuously anastomosing; vein and veinlet 
reticulation hidden; inflorescence axillary, solitary, opp- 
osite, abundant, shorter than the leaves; cymes densely 
many-flowered, 1.5--3 cm. long, 1--2.6 cm. wide, more or 
less brachiate; peduncles very slender, 5--15 mm. long, den- 
sely flavescent-pubescent like the twigs; bracts none; 
bractlets and prophylla linear, 2--5 mm. long, densely flav- 
escent-pubescent; calyx light and thin, obconic, 1.6--2 mm. 


long, about 1.6 mm. wide, densely short-pubescent, its rim 


very minutely 4-denticulate, the teeth being merely the ter- 
minations of 4 heavier costae; corolla infundibular or hypo- 
crateriform, its tube slender, about 2 mm. long and 1 m. 


wide, ampliate above, its limb 4-parted, its lobes elliptic, 
about 2 mm. long and 1.2 m. wide, rounded at apex, venose; 
stamens 4, inserted about 1.8 mm. below the mouth of the 


corolla-tube, equalling the lobes or exserted; filaments 


188 POY)? Oe One tk No. 5 


filiform, 2.5--4 mn. long, twisted, glabrous; anthers ob- 
long, about 1 mm. long and 0.5 m. wide, dorsifixed near the 
base, rounded at both ends; style capillary, included, about 
2 mm. long, glabrous; stigma bifid, its branches about 1 mm. 
long, twisted; ovary minute, about 0.5 mm. long and wide, 4- 
lobed, glabrous, 4-celled; fruiting-calyx and fruit not seen, 

The type of this species was collected by Ernst Heinrich 
Ule (No. 1520) at a laguna in the Campo d'Una, Santa Cathar- 
ina, Brazil, in December, 1889, and is deposited in the her- 
barium of the Botanisches Museum at Berlin. 

BRAZIL: Santa Catharina: Pabst 435 (B); Ule 1520 (B-- 
type, N--2 fragments of type, P--isotype). 


70. AEGIPHILA BARBADENSIS Moldenke. 

Additional citations: BARBADOS: Warming 71 (Cp), 101 (A-- 
photo of type, B--photo of type, F--photo of type, G--photo 
of type, K--photo of type, N--photo of type, P--photo of 
type, W--photo of type, Z--photo of type). 


3. ABGIPHILA BOGOTENSIS (H.B.K.) Moldenke. 

The third synonym on page 283 should read “Amerina tomen- 
tosa (H.B.K.) P. DC." 

Additional citations: COLOMBIA: Santander Norte: Killip & 
Smith 19,706 (N); Schlim 306 (Cb, N--photo, Z--2 photos); 
Cundinamarca: Goudot sen. [Bogoté] (P--2); Kareaten s.n. [Bo- 
gota] (N--photo, Z--photo); Linden 798 (Bm--2, Cb, P, V, X); 
Mutis 2332 (W), sen. (P--cotype); Triana 2123 (Bm, Br, Cb, 
Ed, N--photo, P, Z--2 photos), 3743 (Bm); Tolima: Goudot as. 
n. [Quindiu] (Z--2 photos); Purdie sen. [Summit of Quindiu] 
(Kk) » Sen. (A--photo, B--photo, D--photo, F--photo, G--photo, 
K, N--2 photos, S--photo, W--photo, Z--2 photos); Narino: 
Triana 2124 (Bm--2, Cb, Ed, V, Z--2 photos); El Cauca: Pen- 
nell 7097 (N); Department undetermined: Dawe 192 (K). 
ECUADOR: Carchi: Lehmann 6338 (Z-~photo); Pichincha: 
Schimpff 252 (B--2, Cb, N--fragment), 267 (B--2, Cb); Los 
Rios: Rimbach 466 (8). LOCALITY OF COLLECTION UNDETERMINED: 
Herb. Baillon sen. (P). 


80. AEGIPHILA BOLIVIANA Moldenke. 

The Steinbach 5066 cited on page 400 as the type collect- 
ion, should more accurately be referred to as a cotype. In 
all my monographic work I am using the term "cotype" in its 
original meaning (4), viz. “One of two or more specimens to- 
gether forming the basis of a species, no type having been 
selected. No species would have hoth type and cotypes, but 
either the former or two or more of the latter." The cotype, 
therefore, as I employ the term, is synonymous with what 
Bather and Swingle (5, 6) term "syntype". 

Additional citations: BOLIVIA: Santa Cruz: Steinbach 2799 


: 
; 
: 
1937 Uoldenke, Notes on Aegiphila 189 


(Z--photo), 3116 (Z--photo), 3186 (B), 5066 (A--photo of co- 
type, B--photo of cotype, D--photo of cotype, F--cotype, G-- 
photo of cotype, K--photo of cotype, N--photo of cotype, P-- 
photo of cotype, W--photo of cotype, Z--photo of cotype), 
6437 (A--photo of cotype, -B--photo of cotype, Cb--cotype, D- 
-photo of cotype, F--photo of cotype, G--photo of cotype, 
K--cotype, N--photo of cotype, W--photo of cotype, Z--photo 
of cotype), 6554 (Cb, K, Z--photo), 7260 (Bm, Ca, Cb, E, Ed, 
K, N--2 photos, Ut, Z--3 photos), 7289 (B, Bm, Ca, Cb, &, 
Ed, K, N--photo, Z--2 photos). 


23. AEGIPHILA BRACHIATA Vell. 
| It seems probable that further investigation may reveal 
“Glaziou 14,165 and 17,714 to represent a different species, 
as yet undescribed, with mich smaller calyxes and corollas 
and smaller, darker, more entire leaves. It also seems prob- 
able that the Sellow specimens at Berlin may all be A. Hass- 
leri or a variety thereof. 
_ Additional citations: BRAZIL: Rio de Janeiro: Glaziou 
(14,165 (B--photo, Br, Cb, Cp, D--photo, F--photo, K, N--2 
photos, P, S--photo, W--photo, Z--2 photos), 17,714 (Cb, Cp, 
K, N--photo, P, Z--2 photos); Parandé: Jénsson 979a (B, Cb, 
-E, N, 8, W); Santa Catharina: Fr. Miller a.n. [Schwacke 
1465] (Cb); Rio Grande do Sul: Gaudichaud 1801 (P); State 
undetermined: Sellow ee (K), 1269 (B, P), 1322 (B, Bm, N-- 
photo, P, Z--2 photos), 3012 (A--photo, B, G--photo, N-- 
‘photo, Z--2 photos), sen. (Vu). PARAGUAY: Jorgensen 3662 
(Op, D, E, 3, W). 
: 
120. AEGIPHILA BRACTEOLOSA Moldenke. 
Aegiphila solanifolia Mart., in herb. -- The name Pseud- 
Begiphila breviflora Rusby, reduced to synonymy under A. 
bracteolosa with a question on pages 456--458, has proved 
upon more careful examination to represent a distinct spe- 
cies. The Cardenas 16, special, cited under A. bracteolosa 
on page 458, is the type collection of A. breviflora. The 2 
Martius collections cited hereinafter and designated as 
Herb. Monac. 1004 and 1005 are cotypes of A. arborescens 
var. longiflora Schau.; therefore this trinomial is to be 
egarded as a synonym of A. bracteolosa rather than of A. 
integrifolia as stated on page 53/7. 

Additional citations: BRITISH GUIANA: Lang & Persaud 253 
(A--photo of type, B--photo of type, D--photo of type, G-- 
hoto of type, N--photo of type, P--photo of type, S--photo 
of type, W--photo of type, Z--photo of type). BRAZIL: Amaz- 

as: Ducke 131 (Cb--2, N--fragment, N--photo); Jobert 630 
(P); Krukoff (B, Cb, K--2, N--2, 8); Luetzelburg 22,296 
Mu); Martius s.n. [Porto dos Miranhao, Decbr.; Herb. Monac. 
004] (Ii), s.n. [Barra de Rio Negro, Octbr.; Herb. Monac. 


190 PHYTOL 06-2 2 No. 5 


1005] (Iu), sen. (Prov. Rio Negro; Herb. Monac. 1006] (Im); 
Poeppig 2488 (V, Z--2 photos); Schwacke 3635 (Cb); Spruce 
1283 (K), s.n. [In vicinibus Barra] (A--photo, Bm, Cb, D-- 
photo, Ed, F, N--3 photos, P, W--photo, Z--4 photos); Ule 
5435 (B, Cb, Le, N--photo, Z--photo); Pard: Spruce 101 
fHerb. Monac. 1003] (Mu). PERU: Loreto: Tessmann 5 B, 
Hb, S). ILLUSTRATION: Line drawing (N). 


20. AEGIPHILA BRASILIENSIS Moldenke. 

Additional citations: BRAZIL: Rio de Janeiro: United 
States Exploring Exped. [Wilkes] sen. [Rio de Janeiro] (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--isotype, G--photo of type, K--photo of type, 
N--photo of type, P--photo of type, S--photo of type, W-- 
photo of type, Z--photo of type). 


39a. AEGIPHILA BREVIFLORA (Rusby) Moldenke, Phytologia 1: 

95. 1934. 

Pseudaegiphila breviflora Rusby, Mem. N. Y. Bot. Gard. 7 
1927. 

Shrub [or "tall perennial herb" according to Rusby]; 
branchlets acutely tetragonal, more or less sulcate, very 
medullose, densely incanous-strigose with short, closely- 
appressed forward-pointing hairs; internodes 4~--5.5 om. 
long; Leaves decussate-opposite; petioles much reduced, 
rather stout, 3--4 mm. long, densely incanous-strigose, 
mostly contracted at apex, slightly alate-margined, canalic- 
ulate above; blades chartaceous, dark green above (brunnes- 
cent in drying), lighter beneath, oblanceolate, 10--21 cm. 
long, 3--6 cm. wide, acute or short-acuminate at apex, obso- 
letely sinuate-serrate with gland-tipped teeth, long-cuneate 
at base and gradually tapering into the petiole, densely 
brownish-puberulent on both surfaces; midrib stout, slightly 
prominulent above, very prominent beneath; secondaries slen- 
der, 8--10 per side, lightly arcuate- or falcate-ascending, 
prominulent beneath, not conspicuously anastomosing; vein 
and veinlet reticulation sparse, fine and delicate, obscure 
above, not prominulent beneath; inflorescence axillary; ped- 
uncles slender, 1--2 cm. long, densely incanous-strigose 
with very short closely-sappressed hairs; cymes subcapitate, 
solitary, opposite, 2--3 cm. long, 1--3 cm. wide, once fur- 
cate, very densely many-flowered; bracts and bractlets none; 
prophylla hidden in the dense inflorescence; calyx obovoid- 
campanulate, 3.5--4.7 mm. long, 2--2.5 mm. wide, densely in- 
canous-strigose, its rim normally 4-toothed with short, e- 
qual, erect, obtuse, lobe-like teeth, but often irregularly 
4-toothed or more or less 2-labiate with one lip entire and 
the other 2-lobed; corolla accrescent, slightly exceeding 
the calyx, straight, its tube cylindric, slender, about 2 


1937 Moldenke, Notes on Aegiphila 191 


mm. long, somewhat ampliate at apex, its limb slightly 2- 
labiate, the upper lip entire or subentire and to 3 mn. 
long, the lower lip 3-lobed, with the lobes each smaller 
than the upper lip [occasionally the limb is 3-lobed or 4- 
lobed with all the lobes broadly ovate-lingulate, obtuse, 
and subequal]; stamens 5 or 4, inserted about 1 mm. below 
the mouth of the corolla-tube, exserted, mostly didynamous, 
the 2 long ones normally equal and 2.5--5 mm. long, the 2 
short ones frequently unequal and 1--2 mm. long [in the occ- 
asional 3-lobed corollas only 3 subequal stamens are pres- 
ent]; filaments filiform, slightly flattened, glabrous; an- 
thers oval-oblong, about 1 mm. long and 0.5 m. wide, dorsi- 
fixed near the base, lightly cordate, 2-celled, the thecae 
parallel; pistil exserted; style capillary, 4--5 mm. long, 
glabrous; stigma bifid, its branches filiform, strongly and 
widely recurved, 1--2 mm. long; ovary hemispheric-turbinate, 
about 1 mm. long and wide, glabrous, its truncate sumnit 
bearing a marginal, recurved, membranous, annular appendage 
(according to Rusby) or fonehapeudaced and umbilicate and 
more or less 4-lobed; fruiting-calyx and fruit not seen, al- 
though Rusby describes immature fruits as being closely en- 
closed by the base of the calyx, hemispheric-turbinate, with 
the annulus much expanded, coriaceous, lustrous, light 
brown, and recurved so as to conceal the upper third of the 
fruit, and with a concave center, 2- or 3-sulcate, and 2- or 
3-seeded. 

The type of this most remarkable species was collected by 
Martin Cardenas (No. 16, special) at Santa Ana de Yacuma, at 
an elevation of 700 | Sea" El Beni, Bolivia, March 4, 1922, 
and is deposited in the Britton Herbarium of the New York 
Botanical Garden. This plant was referred by me in my mono- 

graph (pages 457--458) with some doubt to A, bracteolosa, 
which it does indeed closely resemble superficially. Ae 
bracteolosa, however, is a native of British Guiana, Brazil, 
and Peru, and is confined to Amazonian forests. A. brevi- 
flora, if it belongs in this genus at all, is a member of 
the ¢ group Cymosae, subgroup Lobatae, while A. bracteolosa is 
very obviously a member of the group Paniculatae, subgroup 
Dentatae. In his original description Dr. Rusby throws out 
the suggestion that this plant may be a natural hybrid be- 
_ tween an Aegiphila species and a Callicarpa, Unquestionably 
_ the remarkable variations exhibited by the flowers on the 
type collection indicate something abnormal, but I am unable 
to discern any characters which point especially to the gen- 
us Callicarpa. The chief characters by which Callicarpa dif- 
fers from Aegiphila are that in Callicarpa the stigma is 
_ capitate or peltate, the flowers are usually polygamous and 
never diclinous (heterostylous ) as in Aegiphila, the stamens 
are inserted at the very base of the corolla-tube, and the 


192 PHYTOL UGT a No. 5 


ovary is mostly pubescent. None of these features is exhib- 
ited by our plant, although it does have the axillary in- 
florescences characteristic of Callicarpa. Whether or not 
its fruiting-calyxes are accrescent is not yet known. The 
fact that the type collection seems unquestionably to be ab- 
normal leads me to attach less importance to the didynamous 
nature of the stamens seen in many of the flowers than does 
Dr. Rusby. Could it be showm that the stamens are regularly 
didynamous on normal specimens of this species and could the 
curously irregular features of the calyx-rim and corolla- 
limb be proved to be regular features of the species, and 
were the remarkable ovary and fruit features mentioned by 
him always exhibited, then I should perhaps be willing to 
maintain the genus Pseudaegiphila for this plant. The fact 
is, however, that there seems very little constancy in these 
features on the four specimens making up the type collection. 
Many of the calyxes are quite regularly 4-toothed and not at 
all labiate, while others are only obscurely so. The corolla 
limb, too, is quite often almost regular, while on the nun- 
erous 3-lobed ones the lobes are all equal. The stamens seem 
to show no constancy at all. The 3-lobed corolla usually 
have but 3 stamens and these may or may not be equal in 
length. The 4—lobed corolla have 4 stamens and these are 
mostly noticeably didynamous, but not regularly or uniformly 
so. Usually the 2 shorter stamens are equal or subequal, but 
the 2 longer stamens mostly are of varying lengths again. 
The characters of the ovary and immature fruit are similarly 
inconstant. The strange annular appendage which Dr. Rusby 
describes is indeed present on a number of the ovaries, but 
I have examined just as many flowers from the same cyme 
where the ovary exhibited no trace of this annular ring, but — 
was the normal, umbilicate, 4-lobed, 4-sulcate ovary so wide ~ 
spread in the genus Aegiphila. The foliar characters of the 
species are precisely those of A. bracteolosa. 

BOLIVIA: El Beni: Cardenas 16, special (K--isotype, Mi-- 
ey N--type, N--photo of type, W--isotype, Z--photo of 
type). 


102, AEGIPHILA BUCHTIENII Moldenke. . 
Additional citations: BOLIVIA: La Paz: Buchtien 1716 (Z-- — 
photo), 1717 (A--photo of type, B--photo of type, D--photo 
of type, F--photo of type, G--photo of type, K--photo of 
type, N--photo of type, P--photo of type, S--photo of type, 
W--photo of type, Z--photo of type & photo of isotype). 


99. AEGIPHILA CANDELABRUM Briq. 

Add to description: Fruiting-calyx campanulate, Leath- 
ery, 4--6 mm. long, 5--9 mun. wide, densely hirtellous, its 
rim shallowly 4-toothed with triangular teeth about 1 mm. 


1937 Moldenke, Notes on Aegiphila 193 


long and 4 mm. wide at base, acute at apex, or obscurely re- 
pand; fruit drupaceous, elliptic-ovate, 7--l12 mm. long, 5--8 
mn. wide, mostly subumbonate at apex, glabrous, not nitid. 

Additional citations: BRAZIL: pro eic aa Hoehne, Com. 
Rondon 4301 (N). PARAGUAY: Fiebrig 4638 (Cb--2, Z--photo), 
4875 (Bm, Cb--2, Ed, Le, Z--3 photos), 5039 (Cb, N--frag- 
ment, N--2 photos, Z--2 photos), 5201 By Hassler 2886 (Bn, 
Z--photo), 7974 (Bm, Cb--2, N--2 hotest P, X, Z--2 photos), 
7974a (Bm, Ob, P, X, yo hsiet: 8120 (bucsepiype: Z--photo 
of type). 


40, AEGIPHILA CAPITATA Moldenke. 

Additional citations: BRAZIL: S&o Paulo: Burchell 3547 (A 
--photo of type & photo of isotype, B--photo of type, D-- 
photo of type & photo of isotype, F--photo of type & photo 
of isotype, G--photo of type, N--photo of type & photo of 
isotype, S--photo of type, W--photo of type & photo of iso- 
type, Z--photo of type & photo of isotype). 


18. AEGIPHILA CASSELIAEFORMIS Schau. 

Additional citations: BRAZIL: Rio de Janeiro: Raben 842 
(Br--isotype); S&o Paulo: Burchell 3669 (A--photo, B--photo, 
D--photo, F--photo, ee i K, N--photo, P--photo, W-- 
photo, Z--2 photos), 3704 fa Sohne; B--photo, D--photo, F-- 
photo, G--photo, P--photo, W--photo, Z--photo). 


87. AEGIPHILA CAUCEYSIS Moldenke. 

Additional citations: COLOMBIA: Caldas: Pennell, Killip, 
& Hazen 8667_ (A--photo of type, B--photo of type, D--photo 
of type, F--photo of type, G-=-photo of type, K--photo of 
type, N--photo of type & 2 photos of isotypes, P--photo of 
type, S--photo of ‘type, W--photo of type, Z--photo of type & 
2 photos of isotypes). 


54. AEGIPHILA CAYMANENSIS Moldenke. 

It appears that the type collection of this species was 
gathered on Grand Cayman on January 17, 1891. It should be 
so written in the citations on page 354 and the New York 
specimen should be described as "fragment of type". 

Additional citations: CAYMAN ISLANDS: A. S. Hitchcock s. 
n. [Grand Cayman, 1-17-'91] (A--photo of type, B--photo of - 
type, D--photo of type, E--isotype, G--photo of type, K-- 
photo of type, N--photo of type, P--photo of type, S--photo 
of type, W--photo of type, Z--photo of type). 


90. AEGIPHILA CEPHALOPHORA Standl. 

L. H. and E. Z. Bailey describe this species as a "tree" 
and record its blooming in July. 

Additional citations: PANAMA: Canal Zone: Aviles 988 (F); 


194 PHYTOLOGIA No. 5 


Bailey & Bailey 662 (Ba, F); Bangham S43x (N--2 photos, Z--2 
photos); Kenoyer 607 (A--photo of type, B--photo of type, 
D--photo of type, F--photo of type, G--photo of type, K-- 
photo of type, N--photo of type, P--photo of type, S--photo 
of type, W--photo of type, Z--photo of type). 


98. AEGIPHILA CHRYSANTHA Hayek. 

The Poeppig 2314 collection, being designated as the type 
by myself on page 426 from the several collections original- 
ly cited by Hayek, ought to be referred to as a logotype. 

Additional citations: ECUADOR: Manabi: Eggers 14,838 
(Herb. Monac. 1842] (Cp, G, K, Le, Mu, N, N--2 photos, P, S, 
Z--2 photos); Guayas: Egpers 14,348 [Herb. Monac. 1840; Mac- 
bride photos 20,349] (A--photo, B, B--photo, D--photo, F-- 
photo, G--2 photos, Le--2, Mu, N--2 photos, P--photo, 5S-- 
photo, Vu--2, W, W--photo, Z--4 photos); Province undeterm- 
ined: Schimpff 279 (B). PHU: Loreto: Klug 2027 (A, B, E, K, 
Mi, W); Poeppig 2314 (N--photo of logotype, P--isotype, Z-- 
photo of logotype). BRAZIL: Pernambuco: Schenck 4074 (N-- 
photo, Z--photo); Bahia: Curran 225 (Z--photo). BOLIVIA: 
Santa Cruz: Kuntze sen. (Z--photo). 


44b, AEGIPHILA CONTURBATA Moldenke in Fedde, Repert. 37: 

210. 1934. 

Shrub; branchlets stout, very deeply 4-sulcate to the a- 
pex, densely ochraceous-villose-tomentose with very short 
hairs; internodes 3--5 cm. long; leaves decussate-opposite; 
petioles stout, 1.5--2.5 cm. long, densely villose-tomen- 
tose; blades membranous, bery dark above (brunmnescent in 
drying), incanous beneath, obovate-elliptic or obovate, 12.5 
--15.5 cm. long, 5.5--7.5 cn. wide, acute at apex, entire, 
cuneate at base, finely short-pubescent above, densely in- 
canous~tomentose with very short hairs beneath; midrib very 
stout and very prominent beneath, sharp and narrowly pro- 
minulent above; secondaries slender, 10--13 per side, arcu- 
ate-ascending, close together, prominent beneath; inflores- 
cence supra-axillary; cymes solitary, opposite, to 5.3 cm. 
long and wide, very dense and almost subcapitate, densely 
many-flowered; peduncles slender, 1--1.7 cm. long, densely 
villose-tomentose; pedicels slender, tomentose; calyx in- 
fundibular, about 6.2 mm. long and 3.6 mm. wide at apex, 
densely villose outside, glabrous within, its rim irregu- 
larly 4-lobed; corolla infundibular, its tube cylindric, 
about 5.2 mm. long, its lobes 4, oblong-lingulate, about 1.8 
mm. long and 1 mm. wide, acute; stamens 4, inserted about 1 
mm. below the mouth of the corolla-tube, long-exserted; fil- 
anents filiform, about 4.1 mm. long, glabrous; anthers not 
seen; pistil glabrous, slightly exserted; style slender, 
about 5.1 mm. long; stigma bifid, its branches about 1.8 mm. 


: 


1937 Moldenke, Notes on Aegiphila 195 


long, not greatly divaricate; ovary oblong, about 0.6 mm. 
long and wide, truncate at both ends, glabrate, 4—celled; 
fruiting-calyx and fruit not seen. 

The type of this very perplexing species was collected by 
John Newman in Maranh@o, Brazil [the label is inscribed 
"Maranham" ], and is deposited in the herbarium of the Con- 
servatoire et Jardin Botaniques at Geneva. The species dif- 
fers from A. villosa in its deeply sulcate branchlets, its 
very short pubescence on the branchlets and leaves (1 mn. 
long or less), its decidedly pubescent upper leaf-surfaces, 
and the pronouncedly tomentose character of the pubescence 
on the lower surface of its mature leaves. Very immature 
leaves in the process of expanding may exhibit a more vil- 
lose pubescence, but mature blades never possess the long, 
straight, ochraceous hairs of A. villosa nor the short, 
straight, strigose ones of A. intermedia. 

BRAZIL: Maranh&o: Newman sen. (B--photo of type, Cb-- 
type, K--photo of type, l+-fragment of type, N--photo of 
type, S--photo of type, Z--photo of type). 


112. AEGIPHILA CORDATA Poepp. 

Additional citations: PERU: Loreto: Poeppig 2158 (A-- 
photo of type, B--photo of type, D--photo of type, pe, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type & photo of isotype, P--isotype, S--photo of type, W-- 
photo of type, Z--photo of ey & photo of isotype). BRAZIL: 
Acre Territory: Ule 9720 (B, re. 


113. AEGIPHILA CORDIFOLIA (Ruiz & Pav.) Moldenke. 

It is of interest to note that although the original des- 
cription of this species gives the locality as "Miffa", the 
specimens in the British Museum herbarium are inscribed 
"Mufia". 

Additional citations: PERU: Amazonas: A. Mathews sen. 
[Chacapoyas] (Bm, Cb, N--fragment); Huanuco: Macbride 3922 
(A--photo, B--photo, D--photo, G--photo, K--photo, N--photo, 
P=-photo, S--photo, W--photo, Z--photo); Department undeter- 
mined: Rufz 188 (B--isotype); Rufz & Pavon s.n. [Mifia, Pana- 
tahua] (A--photo of isotype, B--photo of i isotype, Bm--type, 
Bm--2 isotypes, Cb--isotype, D--photo of isotype, F--photo 
of isotype, G--photo of isotype, K--isotype, K--photo of 
isotype, N--fragment of isotype, N--2 photos of isotypes, 
P--photo of isotype, S--photo of isotype, W--photo of iso- 
type, Z--2 photos of isotypes). 


29. AEGIPHILA CORIACEA Moldenke. 

Additional citations: BRAZIL: State undetermined: Frey- 
reiss sen. (A--photo of type, B--photo of type, D--photo of 
type, F--photo of type, G--photo of type, K--photo of type, 


196 PHYTOLOGIA No. 5 


N--photo of type, P-~photo of type, rane of type, W-- 
photo of type, Z--photo of type). 


10. AEGIPHILA COSTARICENSIS Moldenke. 

Errata and addenda +o description on page 294: Calyx cyl- 
indric or obconic, 3.5 mm. long and 1.8 m. wide, light, 
glabrous, its rim slightly flaring, 5B-toothed, its teeth a- 
cute; corolla hypocrateriform, its tube very slender, to 8.5 
mm. long, its limb 5—-parted, its lobes narrowly oblong, 5.2- 
-6 mm. long; stigma bifid, its branches 2.3--2.6 mm. long, 
parallel; ovary prismatic, 4-lobed, shallowly umbilicate at 
apex, 4-celled, each cell with one ovule attached at the 
base. The type collection is sometimes cited as "Tonduz 
9167". 

Additional citations: COSTA RICA: Guanacaste: Standley & 
Valerio 44,597 (W), 44,606 (W), 45,538 (A--photo, B--photo, 
i nie op F--photo, G--photo, K--photo, N--fragment, N-- 
photo, P--photo, S--photo, W--photo, Z--photo); Puntarenas: 
H. Pittier s.n. [Herb. Instit. Physico-geogr. Nat. Costaric. 
16,034] (N--photo, W, Z--photo). PANAMA: Bocas del Toro: 
Pittier & Tonduz sen. [Herb. Instit. Physico-geogr. Nat. 
Costaric. 9167] (A--photo of type, B--photo of type, D-- 
photo of type, F--photo of type, G--photo of type, K--photo 
of type, N--photo of type & photo of isotype, P--photo of - 
type, S--photo of type, W--isotype, W--photo of type, Z-- 
photo of type & photo of isotype); Tonduz 8564 [Herb. Monac. 
ae Herb. Instit. Physico-geogr. Nat. Costaric. 8564] (Mu, 
W € 


35. AEGIPHILA CRENATA Moldenke. 

It seems rather certain that more detailed study will re- 
veal that the Pernambuco specimens cited on page 327 and 
hereinafter under this name will prove to be distinct from 
the southern Brazilian specimens which typify the species. 
The Pernambuco form inhabits thickets and blooms in July. 

Additional citations: BRAZIL: Pernambuco: Pickel 
(Ba, I, ¥-2, W); Minas Geraes: Schenck 3310 (B, Z--photo); 
Parané: Dusén 9701 (Z--photo), 10,541 (A--photo ‘of type, B-~ 
isotype, ~B=-photo | of type, Cbh--isotype, D--isotype, D--photo 
of type, E--isotype, F--photo of type, G--isotype, G--photo 
of type, K--isotype, K--photo of type, N-~photo of type, P-- 
photo of yl S--photo of type, W--photo of type, Z~-photo 
of type), 16 (Cb, D, E, G, K, N--photo, Z--photo), s.n. 
(May 5, arty z—-photo); Jénsson 403a (A--photo, B, B—— 
photo, D--photo, F--photo, G--photo, K--photo, N--photo, P-- 
photo, W, W--photo, Z--2 photos); State undetermined: Sellow 

5091 (B, Bm, Z--2 photos). 


9. AEGIPHILA CUNEATA Moldenke. 


1937 Moldenke, Notes on Aegiphila 197 


Additional citations: PERU: Loreto: Killip & Smith 28,379 
(N--photo, Z--photo), 28,386 (A--photo of isotype, B--photo 
of isotype, D--photo of isotype, G--photo of isotype, K-- 
photo of isotype, N--photo of isotype, P--photo of isotype, 
W--photo of isotype, Z--photo of isotype). BRAZIL: Acre 
Territory: Ule 9859 (K, Le, N). 


41, AEGIPHILA DENTATA Moldenke. 

Additional citations: BRAZIL: Sa Paulo: Edwall s.n. 
(Herv. Geogr. e Geol. 4362; Inst. Biol. 15,614] (G--photo of 
type, N--photo of type, W--photo of type, Z--photo of type). 


117. AEGIPHILA DEPPBANA Steud. 

A specimen of this species was determined as “Callicarpa 
sp. nov." by Pavon and another was identified as Buddleia by 
Galeotti. Linnaeus had a specimen of A. Deppeana in his her- 
barium. It is filed under the genus Clerodendrum and is 
sheet number 9 in that folder. It is unnamed and bears no 
inscriptions on its front side except the number of the 
sheet. On the reverse side we find in Linnaeus’ own hand- 
writing (the identity of the handwriting verified by Dr. 
Savage!) "Clerodendrum" and then in darker ink and with a 
heavier pen (but also by Linnaeus, according to Dr. Savage) 
"No. 8 a Millero” and in the former light ink and fine pen 
"Cal. amplius 4 fidus obtusus. Cor. 4 fidus. Stam. 4 long- 
iss. Stylus capillaris semi~bifidus." The specimen was prob- 
ably collected for Linnaeus in the Chelsea Garden, although 
Miller may have received it from a correspondent or the "No. 
8 a Millero" might even mean that sheet number 8 was from 
Miller. -- The Von Rohr 97 cited below is inscribed “West 
Indies", but this is certainly erroneous! 

Additional citations: MEXICO: Tamaulipas: Schiede 1165 
(A--photo of type, B--photo of type, D--photo of type, F-- 
photo of type, G--photo of type, N--photo of type, W--photo 
of type, Z--2 photos of type), s.n. [Estero, Jan. 29] (Bm); 
Schiede & Deppe 259 (Z--photo); Nayarit: Maltby 107 (W), s. 
n. (Tres Marias Ials.] (Z--photo); E. W. Nelson 4245 (N-- 
photo, Z--photo), 4254 (Z--photo); Hidalgo: Liebmann 11,302 
(Cp), 11,303 (Cp); Veracruz: Gouin sen. [1867] (P--2); Hahn 
sen. (Medelli] (K, P--2); Liebmann 11,936 (Cp); Orcutt 3057 
(F), 3418 (F); Edw. Palmer 464 (K, Z--2 photos); Seler & 
Seler 3734 (B); Wawra 614 (V); Oaxaca: Galeotti 1268 (P); 
Chiapas: Seler & Seler 2005 (B, N--photo, Z--photo); State 
undete:mined: Herb. Pavon sen. (Bm--2); Kerber 305 (X); 
Liebmann 11,957 (Cp); Pavon sen. [Nueva Espafia] (Z--photo); 
Sartorius sen. (Z--photo); Von Rohr 97 (Bm). COSTA RICA: 
Guanacaste: Standley & Valerio 46,358 (B--photo, D--photo, 
G--photo, N--2 photos, W--photo, Z--photo); San José: 

Sreted 11,179 (Cp) (a); Department undetermined: frsted 


198 PHY TOLOGI<A No. 5 


11,180 (Cp). PANAMA: Chiriquf: Wagner s.n. [Herb. Monac. 
1007] (Mu); Veraguas: Seemann 1202 (Bm, K). COLOMBIA: Magda- 
lena: Allen 150 (Z--photo); Bertero sen. (Do); H. He Smith 
881 (Bm, Br, Cb--2, EB, Ed, K, Le, N, N--photo, P, Ut, Z—- 
photo), 1864 (Bm, Br, Cb, E, Ed, K, Le, P, Ut, Z--photo) 
(b); Bolfvar: Pennell 4543a (Z--photo). FRENCH GUIANA: Von 
Rohr sen. (Cp--2). OULTIVATED: England: P. Miller 8 (Ls). 
LOCALITY OF COLLECTION UNDETERMINED: Ryan s.n. (Cp). 


124. AEGIPHILA ELATA Sw. 

This binomial is sometimes accredited to "Sch." and oc- 
ours written “Aegiphylla elata Sw." The species has been 
collected in fruit in April; it inhabits roadsides and 
thickets, and its vernacular name in Cuba is “guauro". 
Schipp says that it is a “fairly large bush of spreading ha- 
bit; fls. yellow, slightly perfumed; common in secondary 
forests. June". It is said to be cultivated in Cuba [J. G. 
Jack, Buenos Aires, 1930] (2). The Galeotti 7238 mentioned 
by Turczaninow (3) as possibly representing his A. virgata 
has now been examined and is definitely A. elata. In the 
Linnean Herbarium sheet number 8 under Clerodendrum is A. 
elata. It is unnamed, but bears notations in Limaeus’ own 
handwriting (verified by Dr. Savage!) "Knoxia ? scandens" 
and also "Knoxia 2 Browne 140. t.5.f.3." 

Additional citations: CUBA: Pinar del Rfo: Ekman 12 
(B); Roig 1205 (Es); Havana: Acufia sen. [Herb. Roig ey 
(Es); Santa Clara: Britton & Britton 5086 (N--photo, Z-- 
photo); Jack 7029 (A, K, P, W); Oriente: Ekman 2031 (B, B-- 
photo, D--photo, N--photo, W--photo), 6116 (B, 8); Hioram & 
Maurel 4799 (Z--photo); N. Taylor 414 (Z--photo); C. Wright 

Br, Cb--2, E, G, K--2, Os, P, Z--photo), 154 (Bs Cb, 
E, K--2, P); Province undetermined: G. Don sen. (B); ers 
5184 (B, K, Vu), 5184b (B); Linden 1798 (B, Bm, Cb, K); 
a .360 (Bm). CAYMAN ISLANDS: Fawcitt s.n. [May, 1888] 
K); A. 8. Hitchcock sen. [Grand Cayman, 1-17-'91] (E--3); 
C. F. Millspaugh 1281 (B, N--photo, Z--pnoto); Rothrock 158 
(B). JAMAICA: R. 0. Alexander sen. [Moneague] (K--3), sen. 
(Z--photo); Bertero 2104 (B), sen. [S. D.; Herb. Monac. 
1019] (iu); E. G. Britton 2952 (Z--photo); N. L. Britton 
3408 (N); Britton & Harris 10 (2--photo); Britton & 
Hollick 26 B {a sler 1 P Gea Pango ashe (Cb); 
W. Harris B, Bg, Bm), 10,021 (B, Bm, K, P), 11,082 
(Bm), 1,746 (Bm, E, K), BeNe 7-XI.95] (OL); Hart seNe 
(B--photo, D--photo, G--photo, N--photo, W--photo, Z-- 
photo); Herb. Ventenat s.n. (Cb); A. S. Hitchcock sen. (E); 
W. Hooker sen. [1843] (K); March 972 (K), 1461 (K); Masson 
Sen. (Bm); Maxon 8820 (Z--photo); McFadyen s.n. (K, P); 
Nichols “, (E, Os, Z--photo); J. R. Perkins 1271 (K); Pur- 
die s.n. (K, Le, P, Ut); Rehder s.n. (A); Swartz sen. (Bm--- 


ie 


| 


1937 Moldenke, Notes on Aegiphila 199 


isotype, Ch--isotype, Dc--isotype, N--photo of isotype, S--3 
isotypes, Z--photo of type & 4 photos of isotypes); W. J. 
Thompson 6493 (B), 8012 (B); Wiles s.n. (Cb); N. Wilson 224 
(B), Sen. [Jamaica] (P--2); W. Wright e.n. (Bm, Cb, K); 
Wullschlaégel 974 [Herb. Monac. 1017] (Mu), 1364 [Herb. Mon- 
ac. 1018] (Mu). HISPANIOLA: Haiti: Bertero 35, in part (E, 
P), sen. (De); Deschisaux sen. [Herb. Jussieu 5035a] (P); 
Desportes s.n. [Herb. Jussieu 5035b & 5041] (P--2); Ekman 
HeoLol (B 


)3 Leonard & Leonard 13,072 (A, W); Nash 2 (K); 
Dominican Republic: Abbott 1368 (B, Z--photo), 2 B, Z-- 
photo); era 1602 [Herb. Monac. 3833] (B, Bm, Cb--3, K, 
Le, Mu, Vu), 1602b (B), 16020 (B); Ekman H.12,310 (B, N, 3), 
H.135,279 (B, 8, W). PORTO RICO: Plee sen. (P). MARTINIQUE: 
Collector undesigated s.n. [Hb. Portenschlag] (Z--photo). 


TRINIDAD: W. E. Broadwa (B); Trin. Bot. Gard. Herb. 
2384 (R, Z--photo), 2387 (Z--photo). MEXICO: Oaxaca: Gale- 
otti (Cb, N--photo, P); Tabasoo: Rovirosa 421 (Z-- 


photo); State undetermined: Hahn s.n. [Potrero] (P). GUATE- 
MALA: Alta Verapaz: Tiirckheim 7961 (K, W, Z--photo); Izabal: 
P. C. Standley 24,684 (Z--photo). HONDURAS: Santa Barbara: 
Thieme 5412 (K); Yoro: P. Wilson 656 (N, Z--photo); Atlént- 
ida: P. C. Standley 53,746 (Z--photo), 53,758 (Z--photo), 
55,166 (Z--2 photos); Yuncker 4749 (F, Mi). BRITISH HONDU- 
RAS: Burns 10 (F); Sehipp 216 B, Bm, Ca, Cb--2, E, F, J, K,; 
Mi, 3, W). COSTA RICA: Puntarenas: H. Pittier 6782 (Br, X), 
sen. [Herb. Inetit. Physico-geogr. Nat. Costaric. 12,017] 
(B, Z--photo); Pittier & Tonduz 6782 (X--3); Tonduz s.n. 
(Herb. Instit. Physico-geogr. Nat. Costaric. 6762) (B, Br). 
PANAMA: Panama: R. S. Williams 829 (W, Z--photo). COLOMBIA: 
Magdalena: Goudot s.n. (P--2); El Valle: Lehmann 8410 (B, F, 
K); Oundinamarca: Triana 2081 (Bm, Cb), 3713, in part (Bm). 
VENEZUELA: Carabobo: H. Pittier 8806ea (Cb, G); Suringar s. 
Ne [Puerto Cabello] (Le); Aragua: Fendler Fale Z--photo ); 
State undetermined: Moritz 973 (Bm), 1478 (Bm). BRITISH 
GUIANA: De la Cruz O (Ca, BE, Z--2 photos). SURINAM: 
Focke 297 (Ut), Le); Samuels s.n. [Forest of Zandery ] 
(Z--photo). FRENCH GUIANA: W. E. Broadway 421 (Z--photo); 
Von Rohr sen. (Cp--2). BOLIVIA: Santa Cruz: Steinbach 3259 
? (Z--photo). CULTIVATED: Florida: Buswell sen. [Gov't. 
Grounds, July 22, 1934] (Bu, N); Austria: Cult. Hort. 
Schénb. s.n. (Z--photo). ILLUSTRATIONS: Bot. Reg. 11: t. 946 
1826 (B). 


121. AEBGIPHILA ELEGANS Moldenke. 

Specimens of this species have been found in herbaria 
misidentified as Cordia pubescens Willd. 

Additional citations: PERU: Loreto: Killip & Smith 27,055 
(A--photo of isotype, B--photo of isotype, D--photo of iso- 
type, F--photo of isotype, G--photo of isotype, K--photo of 


200 PHY TOLOGOG IA No. 5 


isotype, N~-photo of type & photo of isotype, P--photo of 
isotype, S--photo of isotype, W--photo of isotype, Z--photo 
of type & photo of isotype), 27,562 (B, Z--photo), 27,991 
(G, Z--photo); Ule 6239 (K); Junin: Killip & Smith 26 
(N--photo, Z--3 photos). BOLIVIA: La Paz: Ule 9718 (K). 


101. AEGIPHILA ELONGATA Moldenke. 

Additional citations: BOLIVIA: La Paz: Buchtien 1645 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, P--photo of type, W-~-photo of type, Z--photo of type & 
photo of isotype). 


61. AEGIPHILA FALCATA Donn. Sn. 

A vernacular name for this species in Costa Rica is 
"zorrillo". 

Additional citations: MEXICO: Chiapas: Purpus 6982 (Ca), 
7521 (Ca). GUATEMALA: Quezaltenango: Tonduz & Rojas 148 (Z-- 
photo); Retalhuleu: Rojas 584 (W); J. D. Smith 1479 [Herb. 
Monac. 1724] (A--photo, B--photo, D--photo, F--photo, K, Mu, 
N--photo, P--photo, S--photo, W, W--photo, Z--3 photos); 
Escuintla: J. D. Smith 2111 (A--photo of type, B--photo of 
type, Cb--isotype, D--photo of type, F--photo of type, G-- 
photo of type, K--isotype, N--fragment of isotype, N--photo 
of type & 2 photos of isotypes, P--photo of type, S--photo 
of type, W--photo of type, Z--photo of type & 2 photos of 
isotypes). COSTA RICA: Limén: Biolley 7409 (Br--2); Cartago: 
He Pittier 11,244 (K), 13,216 (K, W, X--2). PANAMA: Bocas 
del Toro: H. Pittier 8643 (Br, Z--photo); Tonduz 8627 (Br-- 


2), 9292 (Br--2, W), 9293 (Br--2, W, Z--photo), 9293b (X--2), 


15. AEGIPHILA FASCICULATA Dom. &n. 

The “Aegiphila fasciculata Donn. Sm." which Standley des- 
cribes in Field Mus. Pub. Bot. 10: 334--335 (1931) and ill- 
ustrates on plate 57 is Dermatocalyx parviflorus fret. and 
the J. A. Stevenson 83 [Mus. Yale School of Forestry 14,490] 
distributed by him under this name is also Dermatocalyx par- 
viflorus, a member of tne Scrophulariaceae. 

Additional citations: GUATEMALA: Alta Verapaz: Tiirckheim 
4013 (Herb. Monac. 4297 & 4298] (A--photo of type, B--photo 
of type, D--photo of type, F--photo of type, G--photo of 
type, Mu--2 isotypes, N--photo of type & photo of isotype, 
P--photo of type, S--photo of type, W--isotype, W--photo of 
type, Z--photo of type & photo of isotype). 


114, AEGIPHILA FENDLERI Moldenke. 

Additional citations: VENEZUELA: Aragua: Fendler 2032 
(A--photo of type, B--photo of type, D--photo of type, F-- 
photo of type, G--photo of type, N--photo of type & 2 photos 


1937 Moldenke, Notes on Aegiphila 201 


of isotypes, P--photo of type, S--photo of type, W--photo of 
type, Z--photo of type & 2 photos of isotypes). 


31. AEGIPHILA FERRUGINEA Hayek & Spruce. 

It seems rather certain that the Spruce 9473 originally 
cited by Hayek as the type collection of this species, is a 
typographical error for 5473 and that the specimens of this 
number cited by me as “cotypes" ought to be referred to as 
"type" and “isotypes”. 

Additional citations: ECUADOR: Carchi: Mexia 7446 (N); 
Imbabura: Lehmann 4700 (B~-2, B--photo, K, ee Z-- 
photo); Pichincha: Firmin 632 (A--photo, B--photo, D--photo, 
F--photo, N--photo, S--photo, W--photo, rah a Sodiro 
125/22 (B, N--photo, Z--photo); Spruce 5473 (A--photo of oo- 
type, B--photo of cotype, Bm--cotype, Cbh--cotype, Cp--co- 
type, D--photo of cotype, Ed--cotype, F--cotype, F--photo of 
cotype, G--photo of cotype, K--cotype, N--photo of cotype, 
P--cotype, S--photo of cotype, V--cotype, W--photo of co- 
type, Z--3 photos of ootypes), sen. [Aug. 1858] (K). LOCALI- 
TY OF COLLECTION UNDETERMINED: Herb. Pierre s.n. (P). 


63. AEGIPHILA FILIPES Mart. & Schau. 

This species has been confused with Cordia bifurcata 
Roem. & Schult. and some herbarium specimens have been so 
identified! The binomial sometimes occurs accredited merely 
to Martius. The cotype collection cited below has been re- 
ferred to as “Martius 1620", in error. The “"L. Williams 
5146" cited for this species on page 477 is an error and 
should be deleted. 

Additional citations: PANAMA: Panamé: P. C. Standley 
26,55 Sahe Z-~photo). COLOMBIA: Magdalena: H. H. 
Smith 1831 (Bm, Br, Cb--2, E, Ed, K, Le, N, Ne--photo, P, Ut, 
Z--6 ey, totes) PERU: Loreto: Killip & Smith 26,882 (N, W); 
Raimondi 578 (B), 981 (B); Tesamann 3705 (Cb, Hb); L. Will- 
We dams 8190 (N--photo, Z--photo). BRAZIL: lit ‘Hanis 

A--photo, B--photo, Bm, Cb, D--photo, FP--photo, G-- 
STs N--photo, WoT Z--photo); Krukoff 5125 (B, Ob, 
K, N--2, 8); Spruce 1761 (Bm, Br, Cb, Ed, F, K, N--photo, P, 
Z--7 Sutton): Ule Sake Cb, K, Z--photo); Parks Martius s.n. 
{Sylvis secus Amazon, Pardé; Herb. Monac. 1020 & 1689; Mac- 
bride photos 20,350] (G--photo of cotype, Mu--2 cotypes), 8. 
n. [Prov. Paraénsis; Herb. Monac. 1021 & 1022] (Mu--2); Acre 
Petectory: Ule 8293 (B), 9723 (B). BOLIVIA: Bl Beni: H. He 

Rusby 2472 (N--2 photos, Z--2 photos). 


66. AEGIPHILA FLORIBUNDA Moritz & Moldenke. 

Additional citations: VENEZUELA: Aragua: Fendler 845 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, N--photo of type & 2 photos of 


202 Peay Tt Oh Case No. 5 


isotypes, S--photo of type, W--photo of type, Z--photo of 
type & 2 photos of isotypes); Karsten sen. (V); Moritz 1765 
(Bm, N--photo, Ol, V--3, Z--3 photos). __ 


28. AEGIPHILA FLUMINENSIS Vell. 

This specific name is sometimes written with a capital 
initial letter. Gardner notes: “Sometimes all the bracts 
fall off when the persistent calyces are all reflected [{re- 
flexed]. Is this when the umbels consist wholly of @ 
flowers?" 

Additional citations: BRAZIL: Bahia: Blanchet 682 (Bm, 
Cb), 1603 (Ob, F), 1740 (Cb--3, P), sen. [1834] (Bm, Cb); 
Guillot s.n. fBahia’ (Z--photo); Rio de Janeiro: Burchell 

1225 (K, N--photo, Z--photo); Chamberlain s.n. [25'Juli 
1617; Herb. Monac. 1023] (Mu); Collector undesignated s.n. 
(Cb); G. Gardner 5574 (Bm, K, l--photo, Z--2 photos); Gaud- 
ichaud 468, in part (P); Glaziou 806 (Br--2, Cp, P), 3067 
(Br--2, Cb, P); Guillemin 248 (Cb, Do, Z~-photo); Lhotzky 8. 
ne OpYephoto Z--photo); Luschnath s.n. [Brasilia] (EB), sen. 
(B); Martius 1039 [Herb. Boas 1114] (Br, De), 1112 (Herb. 
Monac. 1024] (Mu); Miers 3191 (Cb, K), sen. [Laranjeira] 
(Bm), Sen. [Te juco] (Bm); Mikan sen. [Aquoduit] (2~~photo ) 3 
Pohl sen. [Rio de Janeiro] (Br); Riedel 0 “ ar Riedel 
& Luschnath 323 (L--3); Saint-Hilaire A.361 (P), A 665 (P), 
AL.665 (P); Schwacke 5381 (Cb); Sellow B); United States 
Exploring Exped. Wilke] 2 gen. [Rio de Janeiro] (Z--photo); 
Von Seneloh 139 9 (B); Warming 3 Bene rt 1866 ] (cp--2), BeNe 
[Ad Rio de Janeiro] (Bm); Weddell 40 (P), 95 (P); Widgren — 
661 (Z--photo); Parana: Dusén 8405 (A--photo » B, Be-photo, 
D--photo, F--photo, G--photo, N--photo, W--photo, Z--photo); 
State undetermined: Collector undesignated 285 (P), sen. 
(Br, K); Herb. Martius s.n. (Br); Lusohnath s.n. [Capocabo- 
na] (Br); Sellow sa.n. [Brasilia] (B). 


57. AEGIPHILA FOETIDA Sw. 

Aegiphila pubescens W. Wright, in herb. [not A. pubescens 
Willd., 1 

batviauat citations: JAMAICA: R. C. Alexander s.n. ([Jam- 
aica, 1850] (B--3, B--photo, D--photo, E, F--photo, G, N, 
N--2 photos, P--photo, W, W--photo, Z--4 photos), sen. 
(Pleasant Valley, Monette s 10 Apr. 1850] (K); Distin BeNe 
Nie an Maroh 1411 (B, K); Purdie sen. (Mt. Diablo, April, 

J (); Swartz Bene (Jamaica) (A--photo of type, B--photo 
of ees; Me gras Wet pag Cp~-isotype, D--photo of type, De-- 
isotype, F--photo of type, G--photo of type, N--photo of 
type & photo of isotype, P--photo of type, S--photo of type, 
W--photo of type, Z--photo of type & photo of isotype); We. 
Wright SoNe (Bm). 


—— a 


R91 Moldenke, Notes on Aegiphila 205 


68. AEGIPHILA GLABRATA Moldenke. 


Additional citations: PERU: Junin: Killip & Smith 25,503 
(A--photo of type, B--photo of type, D--photo of type, F-- 
photo of type, G--photo of type, K--photo of type, N--photo 
of type & photo of isotype, P--photo of type, S--photo of 
type, W--photo of type, Z--photo of type & photo of isotype} 


62. AEGIPHILA GLANDULIFERA Moldenke. 
Haught describes this species as a small tree with pen- 
dent inflorescences and says it is "not rare". He also 
states that "The entire plant is strongly aromatic". Klug 
says "Shrub 2 m. tall; fls. cream; blooms in April" and re- 
ports the vernacular name in Peru “chirapa sacha". 
Additional citations: COSTA RICA: Alajuela: Brenes s.n. 
(Herb. Instit. Physico-geogr. Nat. Costaric. 14,484] (B, 
K); Hoffmann 826 (B--2, K, N--photo, Z--photo). PANAMA: 
Canal Zone: Hayes 74 (N--photo, Z--photo), 145 (Z--photo), 
253 (N--photo, Z--photo), 269 (Bm, K); H. Pittier 6519 (G, 
K, Z--photo); Province undetermined: Hayes sen. [Dec. 7] 
(Ed); Seemann 335 (Bm). COLOMBIA: Santander Sur: Dawe 472 
(K=-isotype, N--photo of type & photo of isotype, S--photo 
of type & photo of isotype, W--photo of type & photo of iso- 
type, Z--photo of type & photo of isotype); Haught 1629 (N, 
W); Pennell 3865 (N--photo, Z--photo); Tolima: Goudot s.n. 
[Ibagué] (Z--2 photons); Antioquia: Woronow & Jugepezuk 4445 
(Z--photo). ECUADOR: Napo-Pastaza: Diels 942 (B--2). PERU: 
Loreto: Klug 3016 (A, B, Cb, K, N); Tessmann 3508 (Hb). 
BRAZIL: Amazonas: Poeppig 2760 (N--photo, Z--photo). 


62a, AEGIPHILA GLANDULIFERA var. PARAENSIS Moldenke. 

Additional citations: BRAZIL: Para: Burchell 10,060 (A-- 
photo, B-~photo, D--photo, F--photo, G--photo, K, N--photo, 
S--photo, W--photo, Z--photo); Killip & Smith 30,661 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, N--photo of type, S--photo of 
type, W--photo of type, Z--photo of type & photo of iso- 
type); Krukoff 5923 (N, S); Moss sen. [1919] (Bm). 


62b. AEGIPHILA GLANDULIFERA var. PYRAMIDATA L. C. Rich. & 

Moldenke, var. nov. 

Haec varietas a forma speciei typica recedit ramulis pe- 
dunculisque rhachideque inflorescentiarum ramisque pedicel- 
lisque petiolisque dense strigillosis vel breviter strigo- 
sis, pilis adpressis, foliorum laminis subtus et calyce lev- 
iter strigillosis, et costa supra minute strigillosa. 

This variety differs from the typical form of the species 
in that its branchlets, peduncles, rachis, branches of the 
inflorescence, pedicels, and petioles are densely strigil- 
lose or short-strigose with appressed antrorse buff-colored 


204 PHY T:0:L.0-6 1a No. 5 


hairs, the calyx and lower leaf-surface are lightly so, and 
the midrib above minutely so. 

The type of this variety was collected by Louis Claude 
Richard in Paré, Brazil, and is deposited in the herbarium 
of the Muséum National d'Histoire Naturelle in Paris. The 
collector has inscribed the following notes on the label of 
the type specimen: “Aegiphila pyramidata. Caulis fruticosum 
-- ramis debilibus, propendentibus, pubescentibus. Flores 
sublutescentes in racemum amplum terminalem pyramidatun. | 
Fructus luteo-rubentes 4~loculare. In vixit fluvii Para". 
The Venezuelan vernacular name of “tabaquero"has been re- 
corded by Moritz. 

COLOMBIA: Méta: Triana 37135, in part (Bm), VENEZUELA: 
State undetermined: Moritz (B, Bm). FRENCH GUIANA: Sa- 
got 473, in part (Bm). BRAZIL: Paré: L. C. Richard sen. (Ik- 
photo of type, P--type, Z--photo of type)e 


17. AEGIPHILA GLEASONII Moldenke. 

Additional citations: BRITISH GUIANA: Gleason 237 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--2 photos of 
type, P--photo of type, S--photo of type, W--photo of type, 
Z--photo of type). 


108. AEGIPHILA GLOMERATA Benth. 
Additional citations: ECUADOR: Manabi: Barclay 632 (Bm); 
ers 15,088 (N--photo, Z--photo), 15,827 [Herb. Monac. 
3706] (A--photo, B--photo, D--photo, F--photo, G--photo, Le, 
Mu, N--2 photos, P--2, S--photo, W--photo, Z--2 photos); 
Sinclair s.n. [Salango Isl.] (A--photo of type, B--photo of 
type, D--photo of type, F--photo of type, G--photo of type, 
N--photo of type, P--photo of type, S--photo of type, W-- 
photo of type, Z--photo of type). 


116. AEGIPHILA GLORIOSA Moldenke. 

The species has been confused by some with the genus Vi- 
tex. It seems fairly certain that the "13598" on the Label of 
the type specimen is an error for "1998" and that all the 
specimens of the latter number ought to be regarded as iso- 
types. 

Additional citations: BRAZIL: Bahia: Blanchet 1398 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, S--photo of type, W--photo of type, Z--photo of type), 
1998 (Bm, Cb--2, F, N--fragment); Riedel 781 (L, Z--2 
photos). ILLUSTRATION: Line-drawing (N). 


32. AEGIPHILA GOELDIANA Huber & Moldenke. 
Additional citations: BRAZIL: Para&: Goeldi 8166 (A--photo 


1937 Moldenke, Notes on Aegiphila 205 


of isotype, B--photo of isotype, D--photo of isotype, F-- 
photo of isotype, G--photo of isotype, K--photo of isotype, 
N--photo of isotype, P--photo of isotype, S--photo of iso- 
type, W--photo of isotype, Z--photo of isotype). 


2. AEGIPHILA GOUDOTIANA Moldenke. 

Additional citations: COLOMBIA: Cundinamarca: Goudot 58.n. 
{[Pandi] (A--photo of type, B--photo of type, D--photo of 
type, F--photo of type, G--photo of type, K--photo of type, 
_N--photo of type, P--photo of type, S--photo of type, W-- 
photo of type, Z--photo of type). 


4, AEGIPHILA GRANDIS Moldenke. 
. On page 473 Goudot s.n. [Bogoté] is cited as A. Goudotia- 
na. This was due to a typographical error. The specimen is 
. A. grandis. The cheironyn "Aegiphila guyanensis Moldenke® 
occurs on some herbarium specimens, but was never proposed 
for this species by me! It is not at all the true A. guian- 
ensis Moldenke, q. Vv. 
Additional citations: COLOMBIA: Cundinamarca: Goudot s.n. 
[Bogota] (A--photo, B--photo, D--photo, F--photo, G--photo, 
P, S--photo, W--photo, Z--2 photos); Mutis 365 / (s); 4554 
(B--photo, Cb, F--photo, G--photo, K, N--photo, S, Z-- 
photo); Triana 2080 (Cb, l+-photo, Bhat)! 2 (Bm), 8. 
cn. (W); Tolima: Goudot sen. [Portachuelo, Quindiu] (B--photo 
of type, G--photo of type, pe, N--photo of type & photo of iso- 
type, S--photo of type, Z--photo of type & photo of isotyp@. 


= ~~ 


19. AEGIPHILA GRAVEOLENS Mart. & Schau. 

Aegiphila tetragona Mart., in herb. -- A Sear of 
Martius 1934 at Munich bears the annotation "Stigmatococca" 
in a ve very old handwriting, probably that of Martius, and a 
note in Latin that the specimen probably represents a new 
genus in the Rubiaceae or else in the Solanaceae related to 
Cestrum. A generic description in Latin is appended. Stig- 
matococca Willd. [ex Schult. Mant. 3°: 55. 1827] is a genus 
of doubtful systematic position [see Benth. & Hook. f. Gen. 
Pl. 2: 888], but is typified by 8S. solanacea Willd. (1l.c.) 
with alternate leaves and terminal inflorescences, appar- 
ently quite a different thing from Martius' proposed genus 
of the same name. -- The Sellow 608 cited on page 305 as 
having been collected in Sa&o Paulo was actually collected 
in Bahia [the label at Kew reads "Vittoria - Bahia" ]. The 
Blanchet 3451 cited below is undoubtedly typical of the 
species. The material examined for my monograph and therein 
described [pages 304--305] is apparently not typical and 
ought probably be given a varietal designation at least. On 
the basis of the new Blanchet material, the specific des- 
cription as given in my monograph ought to be modified in 


—— a a oe Se no 


206 PHYTOLOGIA. No. 5 


the following respects: petioles to 1 cm. long, very weak, 
often collapsing in drying; blades elliptic, 9.5-~-21 om. 
long, 3--6.6 cm. wide; secondaries to 14 per side; veinlet 
reticulation obscure above, not at all prominulent beneath; 
cymes to 5 cm. long; peduncles to 2 om. long; pedicels in 
anthesis to 18 mm. long; calyx in anthesis sometimes to 5 
mm. long and 6 mm. wide, often densely pulverulent-punctate; 
filaments 1 ma. long; style 2--3 mm. long; stigma branches 
1.5 mm. long. 
Additional citations: BRAZIL: Bahia: Blanchet 231 (P, V), 
3451 (B, C, Cb--2, N--fragment, N--photo), 2651 (Br, N-- 
photo, V, Z--2 photos); Martius 1934 [Herb. Monac. 1091 & 
1692] (Mu--2); Riedel 221 (L--2, N--fragment); Sellow 608 
(B, B--photo, K, N--photo, Z--2 photos); Rio de Janeiro: 
Martius 120 [Herb. Monac. 1025 & 1027] (Mu--2), sen. [Herb. 
Monac. 1026] (Mu); Miers s.n. [Barra de Iguassu] (Bm); Rie- 
del & Luschnath 1473 (L--2); Sao Paulo: Burchell 5019 (A-- 
photo, B--photo, D--photo, F--photo, G--photo, K, N--photo, 
S--photo, W--photo, Z--photo); Gehrt s.n. [Herb. Instit. 
Biol. S. Paulo 30,081] (K--2, N--2, Sp); Lund 796 (Macbride 
photos 7880] (B--photo of type, Dc--type); Sellow 802 (B, 
P, Z--2 photos), 878 (A--photo, B--photo, Bm, D--photo, F-- 
photo, G--photo, N--photo, S--photo, W--photo, Z--photo); 
State undetermined: Sellow 9 (B--2), 2189 (Br). 


43, AEGIPHILA GUIANENSIS Moldenke. 

Aegiphila arborea Spruce, in herb. -- The “Aegiphila 
guyanensis Moldenke”™ which appears on some herbarium spec- 
jmens is a cheironym which I did not propose, but which 
is apparently synonymous with A. grandis. 

Additional citations: COLOMBIA: Cundinamarca: Triana 
2084 (Cb--2, Z--2 photos); Méta: Karsten sen. [Llano de San 
Martin, Villavicensio] (L, N--photo, Z--photo); Triana 
3713, in part [Llano de San Martin, Villavicensio] (Bm). 
VENEZUELA: Amazonas: Spruce 3578 (K, N--photo, Z--photo). 
BRITISH GUIANA: M. R. Schomburgk 404, in part (A--photo of 
type, B--photo of type, D--photo of type, F--photo of type, 
‘G--photo of type, '--photo of type & photo of isotype, S-- 
photo of type, W--photo of type, Z--photo of type & photo 
of isotype). BRAZIL: Para: Spruce 3113, in part (K). 


22. AEGIPHILA HASSLERI Briq. 

The "Hassler 6831" cited on page 473 is an error and 
should be deleted. The Sellow specimens at Berlin and cited 
under A. brachiata may represent A. Hassleri instead: The 
Niederlein 1205 and Arechavaleta B are anomalous in their 
almost completely glabrous leaves! They may well represent 
a distinct variety. Fruiting specimens from Paraguay, how- 
ever, such as Fiebrig 5923, also have the leaves almost 


— —— ee oe oe —_ 


1937 Moldenke, Notes on Aegiphila 207 


completely glabrous} 

Additional citations: BRAZIL: Paran&: Dusén 9382 (S). 
PARAGUAY: Anisits 122 (Z--photo); Balansa 2085 (B--2, Bm, Cb 
--2, P--2, Z--photo), 2085a (Cbh--2, N--fragment, P); Fiebrig 
260 (Herb. Monac. 4033] (B, B--photo, Bm, Cb, D--photo, Ed-- 
3, F--photo, G--photo, K, Le, Mu, N--photo, W--photo, Z-- 
photo), 530 (B, Cb, Ed, l+-photo, Z--photo), 5923 (Z--2 
photos); Hassler 1674 (K--cotype, P--cotype, Z--photo of co- 
type), 3193 (Bn--cotype, Cb--2 cotypes, F--cotype, K--co- 
type, N--fragment of cotype, NW--photo of cotype, P--cotype, 
X--cotype, Z--2 photos of cotypes), 4271 (Bm--cotype, Z-- 
photo of cotype), 6780 (A--photo of cotype, B--photo of co- 
type, Bm--cotype, Cbh--cotype, D--photo of cotype, F--photo 
of cotype, N--2 photos of cotypes, P--cotype, W--photo of 
cotype, X--3 cotypes, Z--photo of cotype), 8632 (Bm, Cb--2, 
K, P). ARGENTINA: Misiones: Ekman 1227 (Z--photo), 1228 (N-- 
photo, Z--photo); Niederlein 1705 (Z--photo), 1725 (Z-- 
photo), 1732 (Z--photo), 2248 (B, Z--photo); Corrientes: 
Bonpland 755 (P). URUGUAY: Arechavaleta 43 (Cb--3, F, N-- 
fragment), B (B), ssn. (Cp--2 photos); Berro 1019 (X), 508 
(X); Felippone 5082 (Z--photo). ILLUSTRATION: Lamina 1, "Ra- 
na florida en tamano", etc. (Dr). 


34a. AEGIPHILA HASTINGSIANA Moldenke, sp. nov. 

Frutex vel arbor; ramlis gracilibus obtuse tetragonis 
densissime breviterque pubescentibus; petiolis dense brev- 
iterque pubescentibus, pilis fulvis vel ferrugineis; foliis 
chartaceis ellipticis breviter acuminatis, ad basin Longius- 
cule acuminatis, supra dense puberulis vel sparse breviter- 
que pubescentibus, subtus dense velutinis et punctatis; in- 
florescentiis axillaribus cymosis; pedunctlis gracilibus; 
margine calyce valde 4~dentato; staminibus 4. 

Shrub or tree; branchlets slender, obtusely tetragonal, 
very densely short-pubescent with fulvous or ferruginous 
hairs; nodes not annulate; principal internodes 2--4.8 cm. 
long; leaves decussate-opposite; petioles slender or stout- 
ish, 3--7 ma. long, densely short-pubescent with fulvous or 
ferruginous hairs, flattened and canaliculate-margined a- 
bove, convex beneath, not conspicuously ampliate at base; 
blades chartaceous, dark green above, lighter beneath, ell- 
iptic (or stunted ones at the base of the branchlets some- 
times almost subrotund), 2.8--10.8 om. long, 2--4.8 cm. 
wide, short-acuninate at apex (or rounded on stunted 
leaves), entire, rather long-acuminate at base, rather 
densely puberulent above or sparsely short-pubescent, very 
densely velutinous-pubescent with fulvous or ferruginous 
hairs beneath and there also densely punctate, marked with 
numerous, black, elliptic or circular, glandular disks a- 
long the midrib toward the base beneath; midrib slender or 


208 Pe YD TO ba a ae No. 5 


comparatively stout, fluttened or subprominulent and very 

narrow above, rounded-prominent beneath; secondaries slend- 

er, 7--9 per side, arcuate-asoending, flattened above, prom- 
inulent beneath, rather obscurely arcuate-joined in many 
loops near the apex; vein and veinlet reticulation obscure 
or indiscernible on both surfaces; inflorescence axillary, 

cymose; cymes solitary, opposite, 1.3--2.5 cm. long, 7--18 

mm. wide, densely many-flowered, brachiate, far shorter than 

the subtending leaves; peduncles slender, 5--10 mm. long, 

densely short-pubescent with fulvous or ferruginous hairs; 
bracts none; pedicels obsolete; calyx campanulate, thin, 
light and herbaceous, 2.8--5 mma. long, 2--3.2 mm. wide, den- 
sely short-pubescent throughout outside, its rim conspicu- 

ously 4-lobed or 4-toothed, each lobe triangular, about 1 

mm. long and 1.6 mm. wide at base, acute at apex; corolla 

infundibular, its tube cylindric, 2.4--2.8 mm. long, about 

O.7 mm. wide at the middle, often slightly swollen to 1 mm. 

at the base and apex, glabrous throughout, its limb 4-part- 

ed, ite lobes elliptic-obovate, about 2 mm. long and 1 m. 

wide, obtuse at apex; stamens 4, inserted at 2 levels, one 

pair about 1.6 mm. and the other pair about 1.8 mm. above 
the base of the corolla=-tube, included; filaments obsolete 
or very abbreviated; anthers oblong, about 0.8 mm. long and 

0.4 mm. wide, 2-celled, dorsifixed near the base; style cap- 

illary, exserted, 4.8--6 mm. long, glabrous; stigma bifid, 

its branches about 2.5 mm. long, twisted; ovary subglobose, 

0.8--1 mm. long and wide, depressed above, glabrous, 4- 

celled; fruiting-calyx and fruit not seen. 

The type of this rare species was collected by René de 
Grosourdy somewhere in Guatemala in 1864 and is deposited 
in the herbarium of the Muséum National d'Histoire Naturel- 
le in Paris. The species is named in honor of George Tracy 
Hastings, noteworthy collector in Chile, Hawaii, and the U. 
S.A., successful botanical educator, editor of "Torreya" 
since 1921, and an ardent student of Nature. 

GUATEMALA: Province undetermined: Grosourdy sen. [1864] 
(N--fragment of type, P--type). 

(a) This specimen may have been collected in Cartago or in 
Puntarenas. The label merely states “Inter San José et 
Puntarenas", 

(b) On page 452, line 41, this number is erroneously cited 
as "1861". 

(1) Brittonia 1: 245--477. 1934. 

‘2 List Pl. Atkins Instit. Arnold Arb. 7. 1933. 

Bull. Soc. Imp. Nat. Mosc. 36°: 220. 1863. 

(4) Oldfield Thomas, Proc. Zool. Soc. 1893: 242. 1893. 

(5) Natural Sei. 4: 57. 1894. 

(6) Science II, 37: 866--867. 1913. 


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PHY TOLOGIA 


Designed to expedite botanical publication 


Vo. 1 June, 1937 No, 6 


CONTENTS 


A. C. Smitu: Three new Species of Vacciniaceae . 1... 16 6 + « 209 


C. L, Lunvewt: Studies of Mexican and Central American Plants--II . 212 


H. N. Moupenxe:, Additional Notes on the Genus Aegiphila--II . . 222 


Published by H. A, Gleason and Harold N. Moldenke 
The New York Botanical Garden 
Bronx Park, New York, N. Y. 
Price of this number: 50 cents; per volume $5.00 in advance. 
Vol. 1, number 5, was issued March 1, 1937 


cr inert eS 

BP RE 
At =) eS ve ees 
pet ee ner es 


SSS foremwacn pass oe 
Se onl 
. < Paz . 


a "EON AIS 

e NEW YORK 

; BOTANICAR 
THREE NEW SPECIES OF VACCINIACEAE GARDEN 


A. C. Smith 


_. Types of the following species are deposited in the herb- 
arium of the New York Botanical Garden, and are duplicated 
in the U. S. National Herbariun. 


THIBAUDIA ALBIFLORA A. ©. Smith, sp. nov. 
ay Frutex scandens; ramulie elongatis glabris teretibus cin- 
j ereis vel fuscescentibus; petiolis incrassatis glabris 3--4 
mm. longis; laminis coriaceis oblongis vel lanceolato- 
‘oblongis, 12--22 cm. longis, 4-7 cm. latis, basi rotundatis 
‘vel obtusis, apice longe acuminatis, margine leviter revolu- 
tis, utrinque glabris, Spli-nerviis, nervis secundariis 
prope basin orientibus, cum costa supra leviter elevatis vel 
planis subtus prominentibus, venulis reticulatis supra ob- 
gcuris subtus prominulis; inflorescentia ramulis infra folia 
_plerumque exoriente, copiose paniculata, 25--75-flora, 8--10 
om. longa, 5-8 cm. lata, rhachide et ramulis secunderiis 
‘gracilibus cinereo-puberulis; pedicellis gracilibus 9~-14 
mm. longis, cum bracteis et bracteolis minutis puberulis; 
_calyce mox glabro sub anthesi 2=-5 mm. longo, tubo brevi 2-- 
3 mm. diametro basi truncato, limbo lobis late deltoideis a- 
cutis inclusis 1.5--2 mm. longo; corolla alba 8--10 m. 
longa 2-—-3 mm. diametro, basi et apice contracta, lobis l 
mm. longis; staminibus circiter 7 mm. longis aequalibus; 
‘filementis membranaceis connatis 3 mm. longis; antheris 5 
mm. longis, tubulis latis flexilibus distinctis per rimas e- 
longeatas dehiscentibus quem loculis brevioribus; stylo fili- 
formi, stigmate late peltato. 
Type, Ynes liexia 6682, collected Nov. 17--20, 1934, in 
forest near stream, Hacienda Solento, near Santa Rosa, Can- 
‘ton Pajili, Province of Leén, Ecuador, alt. 1000 m. The ped- 
‘icels are said to be clear coral red, the corollas waxy 
i white. The occurrence of white flowers in the genus may be 
more frequent than generally believed, but collectors usual- 
‘ly fail to state the color, and in a dried condition the 
corollas appear red. 
The new species extends southward the known range of that 
emall group of the geme with paniculate inflorescences and 
large leaves, comprising the Colombian T. paniculata A. 0. 
Sn. and T. Archeri A. C. Sm. and the Costa Rican T. costari- 
sensis Hoer. From the first of these, T. albiflora is dis— 
tinguished by its shorter flowers and proportionate ly longer 
leaves, which are neither bullate nor abruptly acuminate. 
Tom the second our species differs also by its short flow 


210 PHY 7:0 b-@:6-f°s No. 6 


ers and by its 5- rather than 7=-nerved leaves. From T. cost- 
aricensis, perhaps its closest ally, the new species differs 
by having its leaves proportionately broader, its inflores- 
cence conspicuously larger and with more numerous flowers, 
and its anthers with comparatively short tubules. 


CAVENDISHIA MEXIAE A. C. Smith, sp. nov. 

Frutex scandens; ramulis subteretibus glabris juventute 
striatis; petiolis subteretibus 7--10 mm. longis glabris; 
laminis coriaceis oblongis, 10--13 cm. longis, 4--5.5 cm. 
latis, basi acutis vel cuneatis, apice caudato-acuninatis 
(acumine 1--2 cm. longo), margine integris et anguste revo- 
lutis, utrinque glabris vel subtus parcissime nigro-setulos- 
is, 5 (—7)-pli-nerviis, nervis secundariis supra basin ori- 
entibus, cum costa supra leviter impressis vel subplanis 
subtus prominentibus, venulis reticulatis supra leviter ele- 
vatis subtus immersis; inflorescentia axillari racemosa 10-- 
14-flora ubique glabra ad 7 cm. longa, basi bracteis imbric- 
atis oblongis papyraceis ad 3 cm. longis instructa; pedicel- 
lis rugosis 7--15 mm. longis, basi bractea eis basi rhachid- 
is simili subtentis, prope basin bracteolas ovatas 5 m. 
longas gerentibus, superne incrassatis et cum calyce articu- 
latis; calycis tubo rugoso breviter cylindrico sub anthesi 
circiter 4 mm. longis et 6 mm. diametro, limbo erecto ob- 
scure glanduloso cum lobis circiter 4 mm. longis, lobis late 
deltoideis 1 mm. longis, praeter apicem calloso-marginatis; 
corolla tenuiter carnosa cylindrica, 17--24 mm. longa, 4--6 
mm. diametro; staminibus subaequalibus (filamentis antheris- 
que compensanter inaequalibus), quam corolla leviter brevi- 
oribus; filamentis nigrescentibus glabris circiter 3 mm. et 
6 mm. longis; loculis leviter gramulatis circiter 5 mm. lon- 
gis; tubulis alternatim circiter 9 mm. et 6 mm. longis, per 
rimas elongatas dehiscentibus; stylo filiformi corollam ae- 
quante, stigmate capitato. 

Type, Ynes hiexia 6916, collected Feb. 18--20, 1935, on 
edge of dense forest near Puyo, Province of Napo-Pastaza, 
Ecuador, alt. 360 m. The closest ally of the new species ap- 
pears to be the Colombian C. amalfiensis Mansf., which has 
the leaves different in texture, less obviously pli-nerved, 
nerrower in proportion, and subcordate at base, the inflor= 
escence shorter, and the flower proportions somewhat differ- 
ent. 


CAVENDISHIA CONFERTIFLORA A. C. Smith, sp. nov. 

Frutex; ramulis subteretibus glabris; petiolis teretibus 
glabris 6--9 mm. longis; laminis tenuiter coriaceis vel 
chartaceis, utrinque pallide viridis, oblongis vel oblongo- 
ellipticis, 12—-16 cm. longis, 3.5--5.5 cm. latis, basi ro- 
tundatis vel minute subcordatis, apice acuminatis (acumine 


1937 Smith, New Vacciniaceae 211 


angusto obtuso 7--20 mm. longo), margine leviter recurvatis, 
supra glabris, subtus parce et minute brunneo-pilosis vel 
punctetis, 5 (vel obscure 7)-pli-nerviis, nervis 5--10 m. 
supra basin orientibus, supra leviter impressis vel elevat- 
is, subtus prominentibus, venulis utrinque prominulis; in- 
florescentiis prope apices ramulorum in axillis foliorum 
caducorum solitariis, racemosis, 15--25-floris, glabris, 
basi bracteis deciduis instructis; rhachide robusto angulato 
4-6 cm. longo; floribue confertis in axillis bractearum 
solitariis, bracteis chartaceis vel tenuiter coriaceis in- 
tegris suborbicularibus vel late obovatis, 10--12 mm. long- 
is, 13--14 mm. latis, margine interdum minute nigro~glandu- 
loso-ciliatis; pedicellis teretibus apice dilatatis et saepe 
obliquis, sub anthesi inconspicuis, sub fructu ad 4 mn. 
longis, basi bibracteolatis (bracteolis chartaceis vel mem 
branaceis ovatis vel obovatis, 4--5 mm. longis et latis, sub 
anthesi calycis basem amplectentibus); calyce coriaceo, sub 
anthesi bracteis occulto 6--8 mm. longo, tubo 4—5 mm. dia- 
metro obscure glanduloso=punctato, limbo suberecto quam tubo 


_paullo longiore, lobis 5 oblongis margine apicem versus in- 
terdum obscure glanduloso-ciliatis, 4--5 mm. longis, 5=—4 


mm. latis, apice obtusis vel interdum emarginatis, margini- 


bus basin versus anguste imbricatis; corolla carnosa cylind- 


| 
i 
L 
4 


A 


} 


eer uroeolata, maturitate ad 16 mm. longa, 6=--7 mm. diamet- 
ro, apice ad 3 mm. contracta, lobis 5 parvis acutis; stamin- 
ibus subaequalibus, filamentis ligulatis glabris distaliter 


contractis et nigrescentibus, alternatim circiter 2.5 et 4 


mm. longis, antheris flavis, alternatim circiter 8 et 7 m. 


_longis, tubulis quam loculis paullo longioribus, per rimas 
@longatas dehiscentibus; stylo quam corolla paullo breviore, 


 stigmate truncato; fructis juvenilibus coriaceis subglobosis 


calycis lobis tiene coronatis. 


Type, A. F. Skutch 2991, collected in December, 1936, in 


riverside thicket in the vicinity of El General, Province of 
San José, Costa Rica, alt. 1560 m. The collector notes that 


the bracts are pink and the corollas waxy white. In a dried 


; 


condition, the apex of the corollas appears much paler than 


the lower portion, which is hidden by bracts. 


4 


This very distinct species, in inflorescence characters, 
can be compared among Central American species only with C. 
crassifolia (Benth.) Hemsl. and another undescribed plant 


redently collected by Skutch, but both of these have much 


larger enveloping bracts, comparatively long pedicels, and 


entirely different calyx lobes, in addition to foliage diff- 
erences. A closer ally of the new species is the OJolombian 
GC. compacta A. C. Sm., with which it has in common short 


pedicels and large slightly imbricate calyx lobes. The Col- 


ombian plant, however, has mich larger bracts which give the 
inflorescence a different aspect, longer flowers, and leaves 


ote PHY BO O-G-Es No. 6 


which are proportionately broader and with more highly con- 
fluent primary nerves. 


STUDIES OF MEXICAN AND CENTRAL AMERICAN PLANTS -- II (a) 
C. L. Lundell 


The present paper is a continuation of studies of Mexican 
and Central American plants based primarily on collections 
in the Herbarium of the University of Michigan and the Gray 
Herbarium. Five of the novelties described, species of Tapi- 
rira, Ilex, Clusia, Bauhinia, and Erythroxylon, were col- 
lected by the writer in 1936 on the Carnegie-Michigan exped- 
ition to British Honduras. 

One of the most important collections ever made in south- 
ern Mexico was received recently from Mr. Eizi Matuda. His 
botanical work is being concentrated in Chiapas, a state too 
much neglected by botanists. A number of new and noteworthy 
species are represented, and seven of the novelties are des- 
cribed herein. Probably the outstanding discovery is a new 
species of Podocarpus which the writer takes pleasure in 
naming for the collector. 


PODOCARPUS MATUDAI Lundell, sp. nov. 

Arbor, foliis coriaceis, lanceolatis, 4--8.5 em. longis, 
12--19 mm. latis, apice acuminatis, basi breviter angustat- 
is; costa supra ad basin versus prominente ad apicem versus 
planiuscula. Amenta mascula solitaria, sessilia, cylindrica, 
ca. 3.5 cm. longa, 4 mm. diam., basi bracteata. 

A tree; leaves coriaceous, lanceolate, 4 to 8.5 cm. long, 
12 to 19 mm. wide, apex acuminate, base short-attenuate, 
costa prominent above near the base, becoming plane toward 
the apex, plane or slightly impressed beneath; staminate a- 
ment solitary, axillary, cylindrical, about 3.3 cm. long, 4 
mm. in diameter, subtended at base by 6 bracts, the bracts 
broadly ovate or depressed ovate, cuspidate, 2 to 4.5 m. 
long. 

Type in the Herbarium of the University of Michigan, E. 
Matuda 698, collected on Mt. Pasitar, Chiapas, Mexico, Dec- 
ember 29, 1936. 

This remarkable addition to the Mexican flora is distin- 
guished from closely related Central American species by its 
prominent costa and broad leaves. 


19357 Lundell, Mexican & Central American Plants 213 


COCCOLOBA ESCUINTLENSIS Lundell, sp. nov. 

Arbor. Ramuli glabri, striati. Folia glabra, subchartac- 
ea, lanceolata vel ovato-lanceolata, 10--26 cm. longa, 4=9 
cm. lata, apice acuminata, basi rotundata vel obtusa; petio- 
lis 1--3.2 cm. longis. Inflorescentia racemosa, subpaniculae~ 
ta, vel subfasciculata, axillaris vel terminalis, minute 
puberula, 6--14 cm. longa; nodulis 1-floris; ochreolis ca. 1 
mm. longis, minute puberulis; pedicellis ochreolas duplo 
superantibus. 

A tree; branchlets glabrous, striate; petioles striate, 
drying blackish, glabrous, 1 to 3.2 cm. long; leaf blades 
entirely glabrous, subchartaceous, broadly lanceolate or o- 
vate-lanceolate, 10.5 to 26 cm. long, 4 to 9 cm. wide, apex 
acuminate, base rounded or obtuse, slightly decurrent, vena- 
tion prominent on both surfaces, the lateral veins about 9 
on each side; inflorescence racemose, subpaniculate, or sub- 
fasciculate, axillary and terminal, 6 to 14 cm. long, the 
racemes slender, striate, minutely puberulous; ochreolae 
tubular-cempamulate, about 1 mm. long, minutely puberulous, 
the subtending bract slightly shorter; floral pedicels soli- 
tary, stout, glabrous, about twice as long as the ochreolae; 
perianth tube very short, glabrous, the lobes broadly ovate- 
elliptic, 2 to 3 mm. long, very minutely ciliate, reflexed 
at anthesis; filaments glabrous, about 2 mm. long. 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda 413, collected at Escuintla, Chiapas, Mexico, Decem- 
ber 2, 1936. 

In the subpaniculate inflorescence, the central rachis is 
once or twice branched near the base, forming one or two 
short lateral racemes. 


BAUHINIA GIGAS Lundell, sp. nov. 

Arbor, 40-metralis, trunco 45 cm. diam., ramulis gracili- 
bus, glabris. Folia membranacea, ovata, 7--14.5 cm. longa, 
6--12.5 cm. lata, 7-nervia, glabra, ad mediem biloba, basi 
cordata, lobis obtusiusculis vel acuminatis; petiolis 5——4.7 
em. longis. 

An unarmed tree, 40 meters high; trunk 45 om. in diamet- 
er; twigs slender, glabrous; stipules minute; petioles very 
slender, 3 to 4.7 cm. long, glabrous; leaf blades membran- 
ous, broadly ovate, 7 to 14.5 cm. long, 6 to 12.5 cm. wide, 
strongly 7-nerved, finely reticulate-veined on both surf- 
aces, glabrous, dark green above, paler beneath, apex 2- 
lobed to below the middle, the lobes divaricate, obtusish or 
acuminate, base deeply cordate. 

Type in the Herbarium of the University of Michigan, 0. 
L. Lundell 6298, collected in advanced forest on top of 
limestone hill at Valentin, El Cayo District, British Hondu- 
ras, June 29, 1936. 


214 P.H.Y T-0:1°0:G-i.8& No. 6 


B. gigas is frequent in forests on the limestone hills at 
Valentin, and may be considered important ecologically. It 
is distinct from the few other North American species of 
similar habit. Unfortunately only sterile material was ob- 
tainable. 


The writer interprets Bauhinia and Cassia in the broader 
sense, hence the following nomenclatorial changes are neces— 


sary. 


BAUHINIA CALDERONII (Rose) Lundell, comb. nov. 
Casparea Calderonii Rose, N. Amer. Fl. 25: 21/. 1930. 


BAUHINIA CONGESTA (Britt. & Rose) Lundell, comb. nov. 
Casparea congesta Britt. & Rose, N. Amer. Fl. Qtseles 
1940. 


BAUHINIA JERMYANA (Britt.) Lundell, comb. nov. 
Casparea Jermyana Britton, N. Amer. Fl. 23: 211. 1930. 


BAUHINIA MONANTHA (Britt. & Rose) Lundell, comb. nov. 
Casparea monantha Britt. & Rose, N. Amer. Fl. 253-2203 
1930. 


BAUHINIA PURPUSII (Britt.) Lundell, comb. nov. 
Casparea Purpusii Britton, N. Amer. Fl. 25: 210..193Gq 


CASSIA BRITTONIANA Lundell, nom. nov. 
Chamaefistula chiapensis Britt. & Rose, N. Amer. Fl. 25: 
258. 1930. 


CASSIA COBANENSIS ( Britt.) Lundell, comb. nov. 
Vogelocassia cobanensis Britton, N. Amer. Fl. 23: 259. 
19350. 


CASSIA COMAYAGUANA (Britt. & Rose) Lundell, comb. nov. 
Chamaecrista comayaguana Britt. & Rose, N. Amer. Fl. 235% 
291. 1930. 


CASSIA DOYLEI (Britt. & Rose) Lundell, comb. nov. 
Peiranisia Doylei Britt. & Rose, N. Amer. Fl. 23: 265. 
1930. 


CASSIA FENIXENSIS (Britt. & Rose) Lundell, comb. nov. 
Chamaecrista fenixensis Britt. & Rose, N. Amer. Fl. 25: 
291. 1930. 


CASSIA LONGIROSTRATA (Britt. & Rose) Lundell, comb. nov. 
Peiranisia longirostrata Britt. & Rose, N. Amer. Fl. 25: 


1937 Lundell, Mexican & Central American Plants 215 
- 264. 1930. | 


CASSIA MAYANA Lundell, nom. nov. 
Chamaecrista Tonduzii Britt. & Rose, N. Amer. Fl. 23: 
290. 1930. 


CASSIA YUCATANA (Britt. & Rose) Lundell, comb. nov. 
Chamaecrista yucatana Britt. & Rose, N. Amer. Fl. 23: 
287. 1930. 


CASSIA MONSERRATENSIS Lundell, nom. nov. 
Chamaecrista chiapensis Britt. & Rose, N. Amer. Fl. 23: 
284. 1930. 


CASSIA PINOI (Britt. & Rose) Lundell, comb. nov. 
Chameaecrista Pinoi Britt. & Rose, N. Amer. Fl. 23: 283. 
~ 1930. 


CASSIA SELERI (Rose) Lundell, comb. nov. 
Chamaecrista Seleri Rose, N. Amer. Fl. 23: 287. 1930. 


CASSIA SIMULANS (Britt. & Rose) Lundell, comb. nov. 
Peiranisia simulans Britt. & Rose, N. Amer. Fl. 23: 266. 
1930. 


_CASSIA STENOCARPOIDES (Britt.) Lundell, comb. nov. 
Chamaecrista stenocarpoides Britton, N. Amer. Fl. 23: 
q 295. 1950. 


CASSIA SUBMONTANA (Britt. & Rose) Lundell, comb. nov. 
Adiptera submontana Britt. & Rose, N. Amer. Fl. 23: 241. 
1930. 


 CASSIA VILLOSISSIMA (Britt. & Rose) Lundell, comb. nov. 

Chamaecrista villosissima Britt. & Rose, N. Amer. Fl. 

a 23: 290. 1950. 

_CASSIA XANTHOPHYLLA (Britt. & Rose) Lundell, comb. nov. 

; Isandrina xanthophylla Britt. & Rose, N. Amer. Fl. 25: 
269. 1930. 


é 

_ ERYTHROXYLON BELIZENSE Lundell, sp. nov. 

a Frutex 2 m. alt., ramlis compressis. Folia chartacea, 
@lliptica, 3--6 cm. longa, 1.8--3.1 cm. lata, apice obtusa 
vel emarginata, basi obtusiuscula; petiolis 5--5 mm. long- 
is; stipulis persistentibus, triangulatis, 3.5--4 mm. long- 
is, 3-striatis, apice 3 (--5)-setulosis. Pedicelli fructif- 
eri 8--10 mm. longi. Drupa oblongo-ovata, acuta, ca. 14 m. 
longa, 5--6 mm. dian. 


216 PEY: TOs 0 Gos - “Bowes 


A shrub 2 meters high; twigs brown, somewhat compressed; 
stipules persistent, broadly triangular, 3.5 to 4 mm. long, 
3-ribbed, the apex 3 (--5)=setulose; petioles slender, can- 
aliculate, 3 to 5 mm. long; leaf blades chartaceous, ellip= 
tic, 3 to 6 om. long, 1.8 to 3.1 cm. wide, apex obtuse or 
slightly emarginate, base obtusish, dark green and lustrous 
above, paler green beneath, costa reddish and prominent be- 
neath, finely reticulate-veined on both surfaces; fruiting 
pedicels 1 to 3 in the axils, slender at base, enlarged ton 
ard apex, ribbed, 8 to 10 mm. long; fruiting calyx deeply 5 
lobed, the lobes ovate-triangular, acute, splitting to 
hae of 1.6 mm.; drupes oblong-ovate, dark red, about 13 

. long, 5 to 6 mm. in diameter, apex acute, oblique. 

Tad in the Herbarium of the University of Michigan, C. 
L. Lundell 6810, collected in open secondary forest on top 
of limestone hill at San Agustin, Mountain Pine Ridge, 51 
Cayo District, British Honduras, August 6, 1936. 

Apparently closely related to E. rufum Cav. from which 
it most obviously differs in size of leaves, length of 
fruiting pedicels, and size of drupes. 


TAPIRIRA MACROPHYLLA Lundell, sp. nov. 

Arbor 25 m. alta. Folia magna, glabra, 26-~44 cm. longa; 
foliolis 6 vel 7, subcoriaceis, oblongis vel elliptico- 
lanceolatis, 8--21 cm. longis, 4.5--7 cm. latis, apice brev—- 
iter acuminatis vel acutiusculis, basi acutis. Paniculae 
4 ,5--12 cm. longae. Drupa ovoidea, 2.3--2.8 cm. longa, obli- 
qua, truncata. 

A tree, 25 meters high; trunk 30 cm. in diameter; bark 
thin, black; tips of branchlets thick, striate, covered with © 
short brownish appressed hairs; leaves pinnate, large, 26 to — 
44 cm. long, glabrous; petioles stout, striate, 4 +o 10 cm. 
long; leaflets 6 or 7, the petiolules of lateral leaflets 5 
to 9 mm. long, those of terminal leaflets 2 to 3 cm. long, 
the blades subcoriaceous, oblong or elliptic-lanceolate, 8 
to 21 cm. long, 3.5 to 7 cm. wide, apex short acuminate or 
acutish, base acute and decurrent, costa prominent, lateral 
nerves 9 to 15 on each side, reticulate veined on both surf= 
aces; panicles axillary, long-pedunculate, narrow, few- 
branched, 4.5-to 12 cm. long, glabrate with age; fruits ov- 
oid, 2.53 to 2.8 cm. long, l-celled, oblique, the apex trunc- 
ate. 

Type in the Herbarium of the University of Michigan, C. 
L. Lundell 6841, collected in riparian forest near San Agus- — 
tin, Mounts Mountain Pine Ridge, El Cayo District, British Hondur- 
as, August 7, 1936. 

The species appears to be closely related to T. mexicana 
Marchand, but may be readily distinguished by size of leaves 
and fruits. 


2’ ee poh - 


: 
by 
E 
™ 


6 


1937 Lundell, Mexican & Central American Plants 217 


ILEX BELIZENSIS Lundell, sp. nov. 

Arbor glabra, 40 m. alta, trunco 45 cm. diam. Ramuli nig- 
rescentes. Folia subcoriacea vel subchartacea, glabra, oblo- 
ngo-elliptica vel oblanceolata, 5--11.5 cm. longa, 2.5--4.4 
cm. lata, apice obtuse acuminata, basi rotundata et breviter 
decurrentia vel acutiuscule; petiolis 6--9 mm. longis; stip~ 
ulis subulato-triangularibus, ca. 1.1 mm. longis, persisten- 
tibus. Infructescentiae in foliorum axillis 3--8-fascicula- 
tae, pedicellis 2--4 mm. longis. Oalyce persistenti ca. 4.5 
mm. diam., breviter 4—~partito. Drupa (immatura) ovoidea vel 
ellipsoidea, 6--7.5 mm. longa, 5--6 mm. diam. 

A glabrous tree, 20 to 40 m. high; trunk 30 to 45 om. in 
diam.; bole straight; bark rough, but not fissured; cortex 
discoloring when cut; branchlets slender, striate, drying 
black; internodes 0.5 to 2 cm. long; stipules triangular, 
subulate, about 1.1 mm. long; petioles slender, canalicula- 
te, 6 to 9 mm. long, drying black; leaf blades subcoriaceous 
or subchartaceous, oblong-elliptic or broadly oblanceolate, 
5 to 11.5 cm. long, 2.5 to 4.4 cm. wide, apex acuminate, the 
acumen obtuse, base rounded and slightly decurrent, or acut- 
ish, costa plane or slightly impressed above, prominent be- 
neath, lateral veins slightly elevated, 7 to 10 on each 
side, margin entire, slightly revolute, dark green, turning 
blackish when dried; infructescence fasciculate, axillary, 
consisting of 3 to 8 pedicels; pedicels thick, 2 to 4 m. 
long, minutely puberulous; the persistent calyx about 4.5 
mm. wide, minutely puberulous, shallowly 4-lobed, the lobes 
rounded or somewhat obtuse-triangular; drupe (immature) ov- 
oid or ellipsoid, 6 to 7.5 mm. long, 5 to 6 mm. in dian., 


drying black, containing 4 or 5 mitlets; the persistent 


stigma 4~ or 5-lobed, about 2 mm. in diameter. 

Type in the Herbarium of the University of Michigan, C. 
L. Lundell 6247, collected in advanced forest in limestone 
valley near Valentin, El Cayo District, British Honduras, 
June 26, 1936. 

Additional specimens examined: BRITISH HONDURAS: El Cayo 
District: Valentin, Lundell 6205, 6331, 6332. 

I. belizensis is one of the characteristic large trees 
in the advanced forest on the limestone ‘plateau. Although 
most abundant in the valleys, it is also present in consid- 
erable mmbers on the hills. The species is evidently rele- 
ted to I.guianensis (Aubl.) Kuntze. 

Certain British Honduras collections [Lundell 4237; 
Bartlett 13087; Schipp 66, 507; Gentle 72, 1024, 1190, 


1244, 1271) referred by Standley to I. panamensis Standl. 


agree rather closely with Guiana specimens [Broadway 741, 


895; De la Cruz 2954, 3654] distributed as I. guianensis. 


Further study may show that I. panamensis is a synonym of 


this species. 


218 PHY T.OLOGIA No. 6 


CLUSIA SUBORBICULARIS Lundell, sp. nov. 

Arbor 17-metralis, omnino glabra, trunco 25 cm. diam. Fo- 
lia coriacea, orbiculari-obovata vel suborbicularia, 12.5-~ 
16 cm. longa, 9--12.2 cm. lata, apice rotundata, basi cunea- 
ta, breviter decurrentia, costa subtus elevata, ad apice ob- 
soleta, nervis lateralibus prominulis; petiolis 15--20 m. 
longis, crassis. Inflorescentiae terminales, breves, 2= vel 
3-florae. Bracteae calycinae 6--10, latissime ovato-orbicu- 
lares, 3--9 mm. longae, rotundatae vel obtusae. Sepala 4, 
latissime ovato-orbiculeria, ca. 11 mm. longa, 13 mm. lata, 
rotundata. Capsula obovoidea, ca. 3.5 cm. longa, 6-locularis 
(raro 5= vel 8locularis). 

A glabrous tree 17 meters high; trunk 25 cm. in diam.; 
petioles 15 to 20 mm. long, thick, flettened above, 3 to 4 
mm. wide; leaf blades coriaceous, orbicular-obovate or 
suborbicular, 12.5 to 16 om. long, 9 to 12.2 cm. wide, apex 
rounded, base cuneate, shortly decurrent, costa prominent 
beneath but becoming obsolete 2.5 to 5.5 cm. from apex, pri- 
mary veins conspicuous, 20 to 25 on each side, diverging 
from the costa at a broad angle, merging into a submarginal 
vein; inflorescence terminal, very short, 2- or 3-flowered; 
the fruiting peduncles thick, less than 1.5 cm. long; bracts 
6 to 10, very crowded, depressed ovate-orbicular, 5 to 9 mm. 
long, rounded or obtuse; sepals 4, depressed ovate-orbicular, 
about 11 mm. long, 13 mm. wide, rounded; capsule obovoid, 
about 3.5 cm. long, 6-celled (rarely 5- or 8-celled); the 
persistent stigmas 6 (rarely 5 or 8). 

Type in the Herbarium of the University of Michigan, C. 
L. lundell 6805, collected in open secondary forest on top 
of limestone hill near San Agustin, Mountain Pine Ridge, El 
Cayo District, British Honduras, August 6, 1936. 

Additional specimens examined: GUATEMALA: Department of 
Petén: La Libertad, Aguilar 241; collected on top of lime- 
stone hill; vernacular name "sello". 

In the absence of flowers the exact relationship of C. 
suborbicularis has not been determined. It is clearly un- 
like any American species known to the writer. 

Another interesting and apparently undescribed Clusia, 
resembling C. Lundellii Standl., but differing in its mich 
larger leaves and capsulesl2 cm. long, was collected near 
Valentin on the limestone plateau. 


OALYPTRANTHES BELIZENSIS (Standl.) Lundell, comb. nov. 
Eugenia belizensis Standl., Field Mus. Bot. 11: 157. 
1932. 
The calyx is evidently calyptrate, hence the species 
should be referred to Calyptranthes. 


EUGENIA MATUDAI Lundell, sp. nov. 


1937 Lundell, Mexican & Central American Plants 219 


Arbor glabra, ramis hornotinis crassis, compressis; in- 
ternodiis 5.5--5 cm. longis. Folia lanceolata vel lanceola- 
to-elliptica, 16--15 cm. longa, 7--7.5 cm. late, apice an- 
gustata, obtuse, basi acuta, subchartacea, glabra; petiolis 
1.7--2.2 cm. longis. Pedicelli fructiferi axillares, fascic- 
ulati, crassi, 1--3 cm. longi. Bacca ellipsoidea, 2--2.5 cm. 
longa, 1.7--2.3 om. crassa, glabra, apice basique rotundata. 
Calycis lobi persistentes, rotundati, 6-7 mm. lati, 3--7 
mm. longi, crassi. 

A glabrous tree; branchlets thick, somewhat compressed, 
especially at the enlarged nodes; internodes 3.5 to 5 cm. 
long; leaves subchartaceous, large, entirely glabrous; peti- 
oles canaliculate, 1.7 to 2.2 om. long; leaf blades lanceo- 
late or lanceolate-elliptic, 16 to 18 cm. long, 7 to 7.5 cm. 
wide, narrowed to an obtuse apex, base acute, costa promin- 
ent beneath, the lateral nerves 10 to 15 on each side; 
fruiting pedicels axillary, fasciculate, stout, 1 to 5 cm. 
long, glabrous; fruits ellipsoid, 2 to 2.5 om. long, 1.7 to 
2.53 om. thick, inconspicuously 8= to l2-ribbed, rounded at 
both ends, glabrous, l-seeded; the 4 persistent calyx lobes 
rounded, 6 to 7 mm. wide, 3 to 7 mm. long, thick, glabrous. 

Type in the Herbarium of the University of Michigan, Eizi 
Matude 648, collected on Mt. Madre Vieja, Chiapas, Mexico, 
May 2, 1936. 

The large thin leaves, long stout petioles, long fruiting 
pedicels, and large fruits distinguish E. Matudai. 


_QOLETHRA GLABERRIMA Lundell, sp. nov. 

. Arbor. Ramuli glabri. Folia parva, glabra, coriacea, pet- 
iolis 8-14 mm. longis; laminis lanceolatis vel oblancecla- 
tis, 5--8 cm. longis, 1.5--2.9 cm. latis, apice apiculato- 

_obtusis vel acutis, basi obtusis vel acutis, integris. Rac~ 
emi ca. 10, graciles, 1--7 cm. longi, tomentelli; pedicellis 
1.2--2.7 mm. longis. Calyx tomentellus, ca. 3.6 mm. longus. 

_ Petala oblonga vel oblongo-obovata, 3--3.6 mm. longa, 2--2.2 

mm. lata, emarginata, ciliato-fimbriate. 

; A tree; branchlets rather slender, pale reddish-brown, 

entirely glabrous; leaves small, glabrous, coriaceous; peti- 

oles slender, shallowly canaliculate, 8 to 14 mm. long; leaf 
blades lanceolate or oblanceolate, 5 to 8 cm. long, 1.5 to 

2.9 cm. wide, apex apiculate-obtuse or acute, base obtuse 

and slightly decurrent, or acute, equal or subequal, margin 
entire, drying paler beneath, costa pale reddish-brown, inm- 

a above, prominent beneath, lateral veins obscure, 
about 9 pairs, finely reticulate-veined; inflorescence of 

branched racemes; racemes about 10, slender, 1 to 7 cm. 

long, brownish-tomentulose with stellate hairs; pedicels 1.2 

to 2.7 mm. long, each subtended by a narrow caducous bract 4 

to 5 mm. long; calyx tomentulose, about 3.6 mm. long, the 


220 PHYTOLOGIA No. 6 


segments oblong or ovate-oblong, obtuse, ciliate; petals ob- 
long or oblong-obovate, 3 to 3.6 mm. long, 2 to 2.2 mn. 
wide, emarginate, ciliate-fimbriate; filaments glabrous; 
style short, less than 1 mm. long; ovary hirsute. 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda 520, collected on Mt. Orando, Chiapas, Mexico, April 
25, 1956. 

C. glaberrima evidently is related to 0. suaveolens 
Turcz. It differs from that species in having shorter race- 
mes and pedicels, smaller ciliate-fimbriate emarginate pet- 
als, and shorter styles. 

The writer's interpretation of C. suaveolens is based on 
Britton's description [N. Amer. Fl. 29: 5. 1914] and speci- 
mens of the species from Alta Vera Paz, Guatemala, collected 
by Tuerckheim in February, 1886. 


CLETHRA MATUDAI Lundell, sp. nov. 

Arbor. Remuli glabri. Folia coriacea, glabra, petiolis 
O.7--2 cm. longis; laminis lanceolato-ellipticis, 7--12.5 
em. longis, 1.8--4.7 cm. latis, apice apiculato-obtusis vel 
acutis, basi cuneatis, serrulatis. Racemi 7, breves, fulvo= 
tomentelli. 

A tree; branchlets entirely glabrous, brownish; leaves 
coriaceous, entirely glabrous; petioles canaliculate, 0.7 to 
2 cm. long; leaf blades lanceolate-elliptic, broadest near 
the center, 7 to 12.5 cm. long, 1.8 to 4.7 cm. wide, apex a- 
piculate-obtuse or acute, base cuneate, equal or subequal, 
apical two-thirds of margin irregularly serrulate with low 
incurved teeth, costa and lateral veins impressed above, the 
costa prominent beneath, the lateral veins slightly elevat- 
ed; racemes 7, short, fulvous-tomentulose; pedicels, subten- 
ding bracts, and flower buds tomentulose. 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda » collected at Pasitar, Chiapas, Mexico, December 
29, 1956. 

Although flowering material is not available for compari- 
son, the species appears easily separable from the other 
North American representatives of the genus. C. Matudai may 
be related to C. vulcanicola Standl. of Costa Rica, which is 
described as having Tcoarsely dentate" nearly glabrous 
leaves and tardily glabrate branchlets. 


CLETHRA PARVIFOLIA Lundell, sp. nov. 

Frutex (7?) ramosissimus. Folia subcoriacea, oblanceolato- 
oblonga, +.5--7.5 cm. longa, 1.3—-2.5 cm. lata, apice acum 
inata vel acuta, basi acuta, supre glabra, subtus mimute al- 
bido-puberulas; petiolis 6-12 mm. longis. Racemi 3--8, grac- 
iles, 3--9 cm. longi, minute tomentelli; pedicellis 2.5--4 
mm. longis. Calyx tomentellus, 3.7-~4 mm. longus. Petala ob- 


1937 Lundell, Mexican & Central American Plants 221 


ovata, 4--5 mm. longa, ciliato-fimbriata. 

A shrub (7), much branched; branchlets slender, rough, 
covered with a pale reddish-brown stellate tomentum, glabra- 
te early; leaves clustered at the apices of the branchlets; 
the internodes very short; petioles slender, 6 to 12 m. 
long, sparsely covered with stellate hairs; leaf blades sub- 
coriaceous, oblanceolate-oblong, 4.5 to 7.5 cm. long, 1.3 to 
2.5 cm. wide, apex acuminate or acute, base acute, costa im 
pressed above, costa and lateral veins conspicuous beneath 
and glabrate very early, irregularly serrulate, entirely 
glabrous above, covered beneath (except upon the veins) with 
a very fine whitish tomentum; racemes 3 to 8, slender, erect, 
3 to 9 cm. long, tomentulose; pedicels slender, 2.5 to 4 m. 
long, each subtended by a narrow caducous bract equaling the 
pedicel; calyx tomentulose, 3.7 to 4 mm. long, the segments 
variable in form, ciliate, the 3 outer oblong, obovate, or 
ovate-oblong, acutish, the 2 inner obovate, rounded; petals 
obovate, 4 to 5 mm. long, ciliate-fimbriate; filaments glab- 
rous; style 1.2 mm. long or shorter; ovary hirsute. 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda 452, collected on Mt. Orando, Chiapas, Mexico, Decem 
ber 30, 1936. 


SIDEROXYLON MATUDAI Lundell, sp. nov. 

Arbor. Folia oblanceolata, 9--19.5 cn. longa, 5.5--7 om. 
lata, apice obtusiuscula, basi longe attenuata, acuta; peti- 
olis 1--1.8 cm. longis. Pedicelli 1.3--1.8 cm. longi. Sepala 
5, persistentia, glabra, latissime ovata vel orbiculari- 
ovata, rotundata, inaequalia, exteriora ca. 1.5 mm. longa, 2 
mm. lata, interiora 2--2.7 mm. longa, 2.5--3.2 mm. lata. 
Fructus subglobosus vel ovoideus, ca. 2 cm. longus. 

A tree; older parts of branchlets defoliate, glabrous, 
reddish-brown; new growth leaf-bearing; apical buds covered 
with appressed reddish hairs; leaves thin, crowded at ends 
of branchlets, covered at first on both surfaces with app- 
ressed reddish or silvery hairs, becoming glabrous very 
early; petioles 1 to 1.8 cm. long, glabrous; leaf blades ob- 
lanceolate, 9 to 19.5 cm. long, 3.5 to 7 cm. wide, apex ob- 
tusish, base long-attenuate, acute, costa impressed above, 
prominent beneath, lateral nerves 15 to 19 on each side, 

_ prominent beneath; fruiting pedicels 1 to several, borne on 
branchlets below the leaves, glabrous, rather thick and 
stiff, 1.3 to 1.8 cm. long; calyx persisting at base of 
fruits 5-phyllous, glabrous, the sepals depressed ovate or 
_ orbicular-ovate, rounded at apex, unequal, the outer smaller 
ones less than 1.5 mm. long, about 2 mm. wide, inner ones 2 
to 2.7 mm. long, 2.5 to 3.2 mm. wide; fruits subglobose or 
ovoid, about 2 cm. long, glabrous, sharply apiculate; seeds 
_ ovoid, about 1.8 cm. long; umbilical area ovate-oblong; en- 


222 PHY TOULOG. IA No. 6 


dosperm abundant (in immature seed). 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda 571, collected on Mt. Orando, Chiapas, Mexico, Decem- 
ber 16, 1936. 


(a) Contribution from the Herbarium of the University of 
Michigan. 


ADDITIONAL NOTES ON THE GENUS AEGIPHILA -- II 


Harold N. Moldenke 


The following notes are supplementary to those contained 
in my original monograph of the genus (1) and in the first 
installment of the present series of additional notes (2). 
To the list of herbarium abbreviations should be added the 
following: Pr = Princeton University, Princeton, N. J. The 
total mumber of recorded contributors to our knowledge of 
this genus, as given on page 245 of my original monograph 
and on page 183 of the first part of these additional notes, 
should be 578 and the mumber of publications reviewed 197. 


49a. AEGIPHILA ACULEIFERA Moldenke. 

The species ascends to at least 1300 m. in Costa Rica and 
has been collected in anthesis in November. A vernacular 
name in Costa Rica is "tabaquilla". 

Additional citations: COSTA RICA: Alajuela: Brenes 5709 
[301] (F--2). 


6. AEGIPHILA ANOMALA Pittier. 

The species has been collected in anthesis in May and in 
fruit in February. A vernacular name in Costa Rica is “taba- 
quillo". It has been confused by some with Siparuna and a 
few herbarium specimens have been distributed under that 
name ! 

Additional citations: COSTA RICA: Alajuela: Brenes 6652 
(F), 15,661 (F). 


2la. AEGIPHILA AUSTRALIS Moldenke. 
Additional citations: BRAZIL: Santa Catharina: Ule 1520 
(N—-photo of type, Z--photo of type). 


1937 Moldenke, Notes on Aegiphila 225 


113. AEGIPHILA CORDIFOLIA (Rufz & Pav.) Moldenke. 
Additional citations: PERU: Department undetermined: Rufz 
& Pavon s.n. [Mifia, Panatahmua] (F--isotype). 


55. AEGIPHILA CRENATA Moldenke. 

The Pickel 526, 3030, and 3042 specimens from Pernambuco 
cited by me on page 327 of my monograph and on page 196 of 
the first installment of these additional notes, do not rep- 
resent A. crenata. They represent a new species which will 
be fully described under species 35a in a future installment 
of these notes. 


117. AEGIPHILA DEPPEANA Steud. 

This species has been confused with and some herbarium 
specimens distributed as Callicarpa sp. and Aegiphila mart- 
inicensis. 

Additional citations: MEXICO: State undetermined: Sessé, 
Mocifio, Castillo, & Maldonado 603 (Q), 1074 (Q). CULTIVATED: 
England: P. Miller 8 [Herb. Linnaeus G.810, S.9] (Z--photo). 


‘124. AEGIPHILA ELATA Sw. 


The type of P. Browne's Knoxia 2 is apparently sheet mm- 
ber 8, under genus 810, Clerodendron, in the Linnaean Herb- 
arium. The "Deschisaux" cited by me on page 199 of my first 
supplement is an error for Desclusaux. 

Additional citations: JAMAICA: P. Browne s.n. [Herb. Lin 
naeus G.810, S.8] (Ls, Z--photo). COSTA RICA: Alajuela: 


Brenes 6193 (F). 


121. AEGIPHILA ELEGANS Moldenke. 

The Ule 9718 cited by me in Phytologia 1: 200 (1937) 
should have been written "Ule 9718, in part", since other 
specimens of this number are definitely A. ovata. 

Additional citations: BRAZIL: Amazonas: Krukoff 8701 (N). 


15. AEGIPHILA FASCICULATA Donn. Sm. 

The "Aegiphile fasciculata Donn. Sm." which Standley des- 
cribes in Field Mus. Pub. Bot. 10: 334——335 and illustrates 
on plate 57 (1931) is A. monstrosa, not Dermatocalyx parvi- 
florus Zrst. as erroneously stated by me in Phytologia 1: 
200 (1937). 


31. AEGIPHILA FERRUGINEA Hayek & Spruce. 


: 
a 
“a 
; 
z 
i 


The "Sodiro 152/22" cited in Brittonia 1: 476 (19354) is a 


typographic error for Sodiro 125/22. 


65. AEGIPHILA FILIPES Mart. & Schau. 
Krukoff describes this species as a shrub to 10 feet 
tall, with a stem to 1/2 inch in diameter and orange fruit, 


224 PHYTOLOGOGHIA No. 6 


inhabiting old clearings. 
Additional citations: BRAZIL: Amazonas: Krukoff 8041 (N). 


2. AEGIPHILA GOUDOTIANA Moldenke. 

The "Goudot s.n. [Bogotd]" cited by me as this species in 
Brittonia 1: 473 (1934) is a typographic error. The collect- 
ion actually represents A. grandis. 


4. AEGIPHILA GRANDIS Moldenke. 

The "Goudot s.n. [Bogoté]" cited by me in Brittonia 1: 
473 (1934) as A. Goudotiana, is the result of a typographic 
error. The collection actually represents A. grandis, as 
stated on page 286. 


95. AEGIPHILA HERZOGII Moldenke. 

Additional citations: BOLIVIA: Santa Cruz: Herzog 1369 
(A--photos of type & of isotype, B--isotype, B--photos of 
type & of isotype, Ch--isotype, D--photos of type & of iso-— 
type, F--photos of type & of isotype, G--photo of type, K-- 
photos of type & of isotype, Le--isotype, Mu-~isotype, N-- 
fragment of isotype, N--photos of type & of isotype, P-=- 
photos of type & of isotype, W-photos of type & of isotype, 
Z--photos of type & of isotype). 


109. AEGIPHILA HIRSUTA Moldenke. 

Additional citations: BOLIVIA: La Paz: Buchtien 1715 (A-- 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, P--photo of type, S--photo of type, W-photo of type, 
Z——photos of type & of isotype). 


86. AEGIPHILA HIRSUTISSIMA Moldenke. 

Additional citations: COLOMBIA: Bolfvar: Pennell 4563 (N- 
-photo, Z--photo). VENEZUELA: Miranda: H. Pittier 8257 jie 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--isotype, G--photo of type, K--photo of type, N-- 
photos of type & of isotype, P--photo of type, S--photo of 
type, W--photo of type, Z--photos of type & of isotype). 


115a. AEGIPHILA HOEHNEI Moldenke, sp. nov. 

Frutex subscandens; ramis subgracilibus obtuse tetragon- 
is dense villosis; ramulis densissime ochraceo-villosis, 
pilis rectis antrorsis 1--2 mm. longis ad basin bulbosis; 
petiolis 5--8 mm. longis densissime longeque villosis; lam- 
inis ovatis 8--12.5 cm. longis, 3.7--6 cm. latis, acuminat- 
is integris, ad basin rotundatis vel subcordatis, supra set- 
osis et subbullatis, subtus dense villosis; inflorescentiis 
axillaribus terminalibusque; panicula terminali cymosa; cy- 
mis subabbreviatis multifloris paucis; pedunculis crassis 


* 


a Moldenke, Notes on Aegiphila 225 
_usque ad 1.5 cm. longis dense ochraceo- vel fulvo-villosis; 
calyce infundibuleriformi 8--10 mm. longo, ca. 4.6 mm. lato, 
dense setoso-villoso, margine valde 4~lobato; tubo corollae 
albae ca. 7.2 mm. longo; ovario subtetragono ca. 1.5 mm. 
longo et diametro, ad basin et apicem truncato glabro 4- 
loculeri; calyce fructifero magno incrassato cupuliformi, 
extra setoso-villoso, margine profunde 4-lobato. 
: Subscandent shrub; branches rather slender, obtusely tet- 
Yragonal, densely but often irregularly villose with gray 
_forward-pointing hairs of various lengths; branchlets rather 
slender, obtusely tetragonal, very densely ochraceous-vill- 
ose with straight forward-pointing hairs 1--2 mm. long and 
bulbous at base; principal internodes 2.7--6.5 cm. long; 
leaves decussate-opposite; petioles stout, 5--& mm. long, 
very densely long-villose like the branchlets; blades firmly 
“membranous or subcoriaceous, bright green, somewhat lighter 
beneath, very nitid above, ovate, &--12.5 cm. long, 3.7--6 
em. wide, acuminate, entire, beautifully rounded or subcord- 
ate at base, setose above with stiff hairs 1——2.5 mm. long 
‘and conspicuously bulbous at base, densely villose beneath 
with ochraceous hairs 2--3 om. long and not at all bulbous, 
densest at the venation, densely punctate beneath; midrib 
slender, prominulent in a channel above, rather prominent 
_ beneath; secondaries 7~-10 on each side, arcuate-ascending, 
prominulent above, prominent beneath; vein and veinlet ret- 
iculation abundant and very handsome, decidedly prominulent 
above and prominent beneath, the lamina more or less pressed 
up from beneath in the interstices (at least, when dry), 
thus imparting a bulleate appearance to the upper surface and 
causing the venation beneath to be very prominent; inflores- 
cence axillary and terminal; terminal panicle cymose like 
the axillary cymes; cymes rather abbreviated, about 3 cm. 
long and 2.5 om. wide in anthesis, to 4.5 cm. long and 5 cm. 
wide in fruit, many-flowered, solitary, few; peduncles 
stout, to 1.5 cm. long, usually much shorter, very densely 
-willose with ochraceous or fulvous forward-pointing hairs; 
calyx infundibular, 8--10 mm. long, about 4.6 mm. wide, very 
“narrow to just above the ovary, then ampliate, densely set- 
ose-villose with hairs to 2 mm. long, its rim deeply 4 
lobed, the lobes ovate, about 2.6 mm. long and 2 mm. wide, 
decidedly rounded and subcucullate at apex, setose; corolla 
white, its tube at least 7.2 mm. long, narrowcylindric, 
glabrous; corolla-limb, stamens, and stigma not seen; style 
at least 7 m. long, slender, glabrous; ovary subtetragonal, 
about 1.5 mm. long and wide, truncate at both ends, glab- 
rous, 4—~celled; fruiting-calyx very large and incraseate, 
cupuliform, to 1.5 cm. long and 1.8 cm. in diameter, vill- 
ose-setose outside, its rim deeply 4-lobed, the lobes tri- 
angular-ovate and about 7 mm. long; fruit yellow-orange, ob- 


226 PHYTOLOGIA No. 6 


long, 13~-16 mm. long, 9--10 mm. wide, glabrous or slightly 
verruculose in drying, not nitid, 4-seeded. 

The type of this distinct and extremely handsome species 
was collected by Joo Geraldo Kuhlmann at Manaos, Amazonas, 
Brazil [Commiss&o Rondon 2277], in November, 1918, and is 
deposited in the herbarium of the Instituto Biologico de 
Defesa Agricola e Animal at SHo Paulo. It gives me great 
pleasure to dedicate this species to Dr. Frederico Carlos 
Hoehne, whose indefatigable labors and collections have add- 
ed so very materially to our knowledge of the flora of Bra- 
zil. The species is obviously related to A. cordata, A. vil- 
losissima, and A. racemosa, from all of which species, how 
ever, it differs in many characters, especially in the sub- 
bullate character of its upper leaf-surface, its subcoriace— 
ous blades, its bulbous-based setose hairs on branches, 
branchlets, and upper leaf-surface, etc. A. cordata and A. 
villosissima, which have pubescence most like that of the 
present species, never have the subcoriaceous and subbullate 
blades, while A. racemosa, which has the coriaceous or sub- 
coriaceous blades and strong vein and veinlet reticulation, 
does not have a pubescence anything like that of our present 
species. 

BRAZIL: Amazonas: Kuhlmann, Com. Rondon 2277 (Herb. Inst. 
Biol. S. Paulo 33,425] (N--fragment of type, N--photo of 
type, Sp--type, Z--photo of type). 


106. AEGIPHILA INSIGNIS Moldenke. 

Additional citations: PERU: Ancachs: Tefalla s.n. [Chico- 
Playa] (A--photo of type, B--photo of type, D--photo of 
type, F--photo of type, G--photo of type, N--photos of type 
& of isotype, P--photo of type, S-photo of type, W--photo 
of type, Z--photos of type & of isotype). 


42. AEGIPHILA INTEGRIFOLIA (Jacq.) Jacks. 

On page 339 of my monograph the name of Huggins should 
have been given as the actual collector of Trin. Bot. Gard. 
Herb. 2385; on line 44 of the same page Burchell 8345 should 
be deleted, since more careful study have proved this col- 
lection to be A. paraguariensis. The "Spruce 3115" cited on 
pages 3539 and 476 should have been cited as "3113, in part", 
since some sheets of this number are identical with Spruce 
3578, which is A. guianensis, and the "Schomburgk out cited 
on the same pages should have been cited as "M. R. Schom- 
burgk 404, in part", since some sheets of this mumber are 
also definitely A. guianensis. C. Houard (3) reports that A. 
integrifolia is often infested with the galls of the insect 
Autodiplosis Jherings Tavares. He also records the vernacu- 
lar name "tocaneiro" for the plant and (4) says that the 
galls of Aegiphila merit further study. Klug describes the 


| 


1937 Moldenke, Notes on Aegiphila 227 


species as a slender tree to 10 m. tall, inhabiting dense 
_ cut-over woods and mountain forests, with white flowers and 


the fruit in heavy clusters, blooming in December, fruiting 
in March. The type specimen of Manabea arborescens has been 


- examined by the present writer in the herbarium of the Brit- 


ish Museum (Natural History). The Ryan s.n. specimen from 


_ Trinidad in the British Museum herbarium is inaccurately 


labeled "Type specimen" of Aegiphila arborescens. The actual 
type of Aegiphila arborescens is the same specimen as con- 
stitutes the type of Aublet's Manabea arborescens. The Aeg- 
iphila arborescens ver. longiflora Schau. cited by me on 
page 537 of my monograph as a synonym of A, integrifolia, 
has proved, upon examination of the actual cotypes at Mun- 


ich, to be synonymous with A. bracteolosa. The Rufz & Pavon 


collection from Pentahua and Chichao is the type collection 


of Callicarpa globiflora. The "Burchell 3418" cited in 
Brittonia 1: 339 & 472 (1934) should read "Burchell 3418, in 


part", since at least one sheet of this number is &. Sellow 


dana. 


Additional citations: TRINIDAD: W. E. Broadway 5836 (A, 
Bm--2, E, K), 6682 (E, F, Z--photo), s.n. [Manzanilla, Aug. 
1918] (K), s.n. [Trin. Bot. Gard. Herb. 7164] (R); Crliger 8. 


n. [17 Aug. '46] (K); Eggers 1002 (Herb. Monac. 3831) (B--2, 


se ti, ee i ti al 


Br, Mu, P, Vu), 1364 (Cp--2, K); Fendler 596 (Bm, K); Hug- 
ins s.n. (Trin. Bot. Gard. Herb. 2585] (B, W); Ryan s.n. 
Bm, Cp--7, Le); Swabey s.n. [Trin. Bot. Gard. Herb. 12,252] 


(K); Trin. Bot. Gard. Herb. 4770 (R); Vahl s.n. [1796; Herb. 


A. L. Jussieu 5036] (P); Williams, Freeman, & Cheesman s.n. 
[Trin. Bot. Gard. Herb. 11,329]-(K). COLOMBIA: Boyaca: 
Lawrance 548 (A, B, Cb, E, G, N, S, Ut); Department undeter- 


mined: Triana 375 (Bm, W). VENEZUELA: Zulia: H. Pittier 
10,626 (Cb, P); Tejera 119 (G), 124 (G, Z--photo); Amazonas: 


————— 


Bonpland 956 (B); Spruce s.n. [April, 1650] (Bm, Cb--3, Ed, 
F). BRITISH GUIANA: Gleason 513 (Z--photo); Jenman 1761 (Z—- 
photo), (U, Z--photo), 5948 (U); M. R. Schomburgk 362 
(B, Bu, K), 404, in part (Bm, Ob—2, De, F, K, Le, P, Us), 
679 (B); Waby 8355 (U). FRENCH GUIANA: Aublet s.n. (Bm); 
Herb. Adanson 227 (P). ECUADOR: Napo~Pastaza: Mexia 71 

(N), 7167 (N), 7289 (N). PERU: San Martin: Klug 5468 (0b, E, 
I, N); Loreto: Castelnau s.n. [Rfo Ucayali] (P); Ule 6821 


(Cb, Le); Huanuco: Macbride 5053 (B, Z--photo); Department 


undetermined: Dombey s.n. (P--2); Rufz & Pavon s.n. [Pente- 
hua & Chichao] (Bm, N--2 photos, Z--2 photos), s.n. [Peruvia 


et Chili] (Bm, N--photo, Z--photo). BRAZIL: Amazonas: Ducke 


(Cb); Huber 4253 (Cb); Labroy 9 (P--2); Poeppig 1615 
A--photo, B--photo, Bm, Cb, D-=-photo, Dc, F--photo, G= 
photo, K, N--photo, P, V-=-2, W--photo, Z--3 photos); Ule 
861 (B, Cb, Le, W); Pard: Spruce 691 [Herb. Monac. 1002] 
Mu), 3113, in part (B, Bm, Br, Cb--2, Op, Ed), s.n. [In 


228 PHYTOLOGIA No. 6 


vicinibus Santarem] (K); Bahia: W. Baldwin s.n. [June 3, 
1818] (N-~photo, Z--photo); Blanchet 2121 [Herb. De Candolle 
829] (Cb, Dc); Minas Geraes: Mexia 4205 (B, Bm, Cb, D, E, G, 
I, P, S), 4500 (A, B, Bm, Cb, D, E, G, I, P, 8, W)3 Rio de 
Janeiro: Burchell 2581 (K); S&o Paulo: Curran 9 (N); Matto- 
grosso: Hoehne, Com. Rondon 1341 (B), 5122 (N); State undet- 
ermined: Glaziou "A" (P--2). BOLIVIA: El Beni: Buchtien 3533 
(Mu); H. H. Rusby 1722 (Z--photo), 2459 (Z--photo); La Paz: 
M. Bang 584 [Herb. Monac. 1726] (A--photo, B--photo, Bm, Ob, 
Cp, D-=photo, E, Ed, F-=-photo, G-=photo, K, L, Le, Mi, Mu, 
N--photo, O01, R, Us-~2, Vu, W, W-photo, Z--2 photos), oe 
(Cb); Buchtien 719 (Cb, Cp, E, Ed, K, N--photo, Z--photo); 
Cardenas 2065 (K, Mi, Z--photo); Fleischmann 520 (S3~-2); 
Troll 1704 (B); Santa Cruz: Herzog 1571 (B, Cb, Le, Z— 
photo); Steinbach 5498 (Cb, Z--photo), 7514 (Bm, Cb, E, K). 
CULTIVATED: Hispaniola: Haiti: Mayerhoff s.n. [1859] (B). 
LOCALITY OF COLLECTION UNDESIGNATED: Collector undesignated 
s.n. (P). ILLUSTRATIONS: Aubl. Hist. Pl. Guian. 1: « iTS 
(P) ° 


44. AEGIPHILA INTERMEDIA Moldenke. 

The petioles of this species become up to 2 cm. in length 
and the leef-blades to 24 cm. long and 9 cm. wide. The leaf- 
blades are villose with short straight hairs beneath, not 
tomentose as erroneously stated by me on page 342 of my mo- 
nograph. 

Additional citations: BRAZIL: Pard: Huber 809 (Bm); Mar- 
enhfio: Ducke 544 (Cb, N--fragment); Herb. Gen. Mus. Para. 
2270 [Herb. Mus. Goeldi] (A--photo of type, B--photos of 
type & of isotype, Bm--isotype, Ch--isotype, D--photo of 
type, G--photo of type, K--photos of type & of isotype, Nhe- 
photos of type & of isotype, P--photo of type, S--photos of 
type & of isotype, W--photo of type, Z--photo of type). 


82. AEGIPHILA KILLIPII Moldenke. 

Additional citations: COLOMBIA: Santander Norte: Killip 
& Smith 20,870 (A--isotype, A--photo of type, B--photo of 
type, D--photo of type, F--photo of type, G--photo of type, 
K--photo of type, N--photos of type & of isotype, P--photo 
of type, S--photo of type, W--photo of type, Z—-photos of 
type & of isotype). 


53. AEGIPHILA LAETA H.B.K. 

On page 394 of my monograph I gave the name A. laeta H. 
B.K. as a synonym of A. laevis (Aubl.) Gmel. This was done 
before the opportunity was presented to me of examining the 
type of A. laeta -- the reduction having been made on the 
basis of statements in the literature of the gemis. Since 
1934, however, I have examined the originel type of A. 


1937 Moldenke, Notes on Aegiphila 229 


laeta. It has been found to be a perfect match for the spec- 
ies which I called A. stricta Rusby on pages 552--455 of my 
monograph. Rusby's name, therefore, has to be reduced to 
synonymy and the correct name for species no. 55 now becomes 
A. laeta H.B.K. Mocquerys reports the species as a tree 5--4 
m. tall, with white flowere and orange fruit. The name A. 
stricta is sometimes erroneously accredited to me on herbar-~ 
ium sheets! 

Additional citations: PANAMA: Taboga Island: Barclay 2498 
(Bm, N--fragment). COLOMBIA: Magdalena: ©. Allen 416 CK); H, 
H. Smith 330 (B--photo, Ob, D-=photo, E, G--photo, K--2, N-= 
3 photos, P--3, S--photo, Ut, W, We-photo, Z--4 photos), 
350a (Z--photo); Santander Sur: Killip & Smith 14,971 (N-- 
photo, Z--photo); Caldas: Bonpland 1364 (N=--photo of type, P 
--type & isotype, Z--photo of type). VENEZUELA: Zulia: Plée 
s.n. (Maracaybo] (Z--photo); Anzoategui: Potter 5154 (N); 
Btate undetermined: Mocquerys 1018 (P-=4); Saer 602 (F). 


78. AEGIPHILA LAEVIS (Aubl.) Gmel. 

Aegiphila laeta H.B.K. is not a synonym of A. laevis 
(Aubl.) Gmel. as stated by me on page 394 of my monograph, 
but is the correct name for species no. 535 (which see, a- 
bove). The specific name of the present species is sometimes 
misspelled "levis", both under Aegiphila and under Manabea, 
the former appearing by accident in Brittonia 1: 389 (1934). 
The species has occasionally been confused with A. brachia- 
ta. The misspelling "Aegiphila lavis" also occurs. The Von 
Rohr s.n. from Cayenne at the British Museum is labelled 
WPype specimen". It is indeed a very typical specimen, but 
not the nomenclatural type. The actual type is the Aublet s. 
n. in the same herbarium. The type has the calyx splitting 
into 4 acute lobes or teeth when the flowers are in full an- 
thesis, merely scalloped before anthesis! 

Additional citations: COLOMBIA: Antioquia: Pennell 3696 
(Z--photo). BRITISH GUIANA: Talbot s.n. (Z--photo). SURINAM: 
Berthoud-Coulon 196 (Bm); Collector indig. 82 (Ut); Hostmann 

3 ae Bm—-=2, Ob, xy 21 (K), s.n. (Le); Hostmann & Kappler 
545 [Herb. Monac. 1189] (Mu, Ut, V--2, Z--3 photos), 721 (Vv, 
2-4 photos); Kappler 543 (P), s.n. (Z--photo); Samels 104 
(K), 382 (Z--photo); Wullechltigel 405 (Br--3, V, Z--photo). 
_ FRENCH GUIANA: Aublet s.n. (A--photo of isotype, B--photo of 
isotype, Bm--type, D--photo of isotype, F--photo of isotype, 
_G--photo of isotype, N--photo of isotype, S--photo of iso- 
type, W-photo of isotype, Z--photo of isotype); Collector 

- undesi ted s.n. [Cayenne] (Dc--3, P), s.n. (P--2); Estebom 
ya =F Gabriel s.n. [1802] (Cb--3); Leblond 362 (P); Lep- 
'rieur s.n. [Guian. fr.] (Dc), s.n. (Z--photo); J. Martin 5. 
in. Rudge] (Bm), s.n. (A--photo, B--photo, D--photo, F-— 
photo, G--photo, K, N--photo, S--photo, W-photo, Z——2 pho- 


230 PHYTOLOGIA No. 6 


tos); Mélinon s.n. [1845] (P--2); Perrottet s.n. [1819] (Cb- 
-2), s.n. [1020] (Cb, P); Poiteau s.n. (Cb--2, P); L. C. 
Richard s.n. (Guian. fr.] (P), s.n. (Cp, Z--photo); Sagot s. 
n. [Boura, 1858] (P), s.n. [Cayenne, Fevrier 1859] (P--2); 
Soubirou s.n. (P), 8.n. |Prés Oayenne] (P); Von Rohr s.n. 
[Cayenne] (Bm). BRAZIL: Bahia: Blanchet 862 (Cb), 1600 (Bm, 
Cb); Sellow 1828 (B, Z--2 photos); Rio de Janeiro: Sellow 10 
(B, Ne=2 photos, Z--2 photos); Mattogrosso: Hoehne, Com. 
Rondon 1287 (Sp); State undetermined: Herb. Link s.n. (Z-- 
photo); Sellow s.n. (N--photo, Z--photc). ILLUSTRATIONS: 
Aubl. Hist. Pl. Guian. 1: 66. 1775 (P); Lamarck, Illustr. 1: 
t. 70, f. 3. 1791 (B). 


47, AEGIPHILA LANATA Moldenke. 

Additional citations: BRAZIL: Goyaz: Glaziou 21,917 (A-- 
photos of type & of isotype, B--photos of type & of isotype, 
Br--isotype, Ob--isotype, Op--isotype, D--photos of type & 
of isotype, F--photos of type & of isotype, G--photo of 
type, N--fragment of isotype, N--photo of type & 2 photos of 
isotypes, P--2 isotypes, P--photos of type & of isotype, W-- 
photos of type & of isotype, Z--photo of type & 2 photos of 
isotypes). 


97. AEGIPHILA LANCEOLATA Moldenke. 

E. Z. and L. H. Bailey describe this species as a woody 
vine, with yellowish corollas, blooming in January. The 
Léfgren s.n. cited by me in my monograph as from SHo Paulo, 
Brazil, should have been listed as from cultivation, since 
the specimens of this collection were collected in the Horto 
Botanico at SHo Paulo. The “Instituto Biologico de Defesa 
Agricola e Animal, SHo Paulo, 20,099" and "20,562" cited by 
me on page 474 as A. vitelliniflora are typographic errors. 
These two collections are actually A. lanceolata as stated 
on page 425. 

Additional citations: BRAZIL: Pard: Burchell 9645 (K); 
Minas Geraes: Clausen 630 (P); Espirito Santo: Glaziou 
11 (Cp, K, Z--2 photos); Rio de Janeiro: Burchell 1624 

K, Z--2 photos); Pohl 5535 (L), a.n. [Rio Paranahyba] (A— 
photo, B--photo, D--photo, F--photo, G--photo, N-—2 photos, 
S--photo, V, We~photo, Z-~2 photos); SHo Paulo: F. 0. Hoehne 
s.n. (Herb. Inst. Biol. S. Paulo 20,562] (Z~-photo); State 
undetermined: Herb. Imp. Vien. 1642 (Z--photo), 1643 (Z-- 
photo). PARAGUAY: Balansa 2094 (Cb--2, N--fragment, P--2); 
Fiebrig 6241 (A--photo of type, B--isotype, B--photo of 
type, D--photo of type, Fe-photo of type, G--photo of type, 
N--photo of type, S-~photo of type, W-photo of type, Z-- 
photo of type). CULTIVATED: Brazil: Rio de Janeiro: Bailey & 
Bailey 575 (Ba); So Paulo: Lifgren s.n. (Herb. Inst. Biol. 
S. Paulo 20,099] (N, Z--photo). | 


1937 Moldenke, Notes on Aegiphila 231 


79. AEGIPHILA LAXICUPULIS Moldenke. 

Additional citations: GUATEMALA: Bernoulli 748 (B, Cb, 
K); Santa Rosa: Heyde 191 (A-~photo of type, Be-photo of 
type, D--photo of type, F--photo of type, G-photo of type, 
K--photo of type, N--photo of type, P--photo of type, S-~ 
photo of type, W-photo of type, Z--photo of type); Heyde & 
Lux 4041 [Herb. Monac. 1723] (Cb--2, Ed, Mu, N-=2 photos, Wi, 
‘J--2 photos). SALVADOR: Ahuachap&n: Padilla 80 (E); San Vic~ 
ente: Calderén 1184 (N--2 photos, Z--2 photos). NICARAGUA: 
Matagalpa: Rothschuh 628 (A--photo, Be-photo, D--photo, F-~ 
photo, G--photo, K=--photo, N=-photo, P--photo, S=-photo, W-- 
photo, Z--photo). 


76. AEGIPHILA LAXIFLORA Benth. 

An additional synonym is Aegiphila laeviflora Briq., in 
herb. The species has been confused by some with the genus 
Olerodendrum and some specimens have been distributed under 
that name. 

Additional citations: TRINIDAD: W. E. Broadway 7386 (A, 
Ob, E, F), s.n. [Buenos Ayres] (D, N, P, W, Z-~3 photos); 
Mrs. W. E. Broadway s.n. [St. Ann's] (A--photo, B--photo, 

De-photo, G-photo, K, N--2 photos, S--photo, W-photo, Z-~4 
photos); Orliger s.n. (B, Z--photo); Dannouse s.n. (Z--pho- 
to); Finley s.n. [Trin. Bot. Gard. Herb. 2391] (R, W, Z-=3 
photos); J. R. Johnston 114 (Z--photo); Lockhart s.n. [All 
hills that have been lately cultd.] (K); L. A. M. Riley 64 
(Bn, K); Trin. Bot. Gard. Herb. 427 (Z--photo), 5126 CZ 

_ photo). BRITISH GUIANA: M. R. Schombur 401 (Ob, V, Z-= 
photo), 592 (B, Ne-photo, Z--2 photos), 772 (A--photo of 
type, B--photo of type, Bm--isotype, Br--isotype, Cb--2 iso- 

types, D--photo of type, Dce--isotype, F--isotype, F--photo 
of type, G--photo of type, K--isotype, Le--isotype, N--frag- 
ment of isotype, N--photos of type & of isotype, P--isotype, 
 $--photo of type, Us--2 isotypes, V--isotype, W#-photo of 
type, Z--photos of type & of isotype). VENEZUELA: State un- 
determined: Otto 1092 (Z~-2 photos). 


27. AEGIPHILA LEHMANNII Moldenke. 
| The "Triana 2083" cited by me on pages 316 and 476 of ny 
monograph, should read "Triana 2083, in part". 

Additional citations: COLOMBIA: Boyacdé: Lawrance 156 (A, 
B, Cb, E, K, Mi, 3, W)3 Chocé: Triana 2083, in part (Bm, Cb, 
N--fragment), 3713, in part (Bm); El Cauca: Lehmann B.T.1117 
_ (K--2 isotypes, Le--isotype). 


aaa 


81. AEGIPHILA LEWISIANA Moldenke. 

Additional citations: VENEZUELA: Federal District: Fend- 
ler 844 (A--photo of type, B--photo of type, D--photo of 
type, F--photo of type, G--photo of type, N--photo of type & 


252 PHYTOLOGIA No. 6 


5 photos of isotypes, P--photo of type, S--photo of type, 
phote of type, Z--photo of type & 3 photos of isotypes). 


33. AEGIFHILA LHOTZKIANA Cham. 

Additional synonyms are the following: Aegiphile glandif= 
era Casar., in herb. [not A. glandulifera Moldenke, 1932], 
Aegiphila villosa Lam., in herb. [not A. villosa (Aubl.) 
Gmel., 1789], Aegiphila vestita Mart., in herb., and Vitex 
erythrocarpa Salzm., in herb. [not V. erythrocarpa Girke, 
1928]. The type of A. Lhotzkiana was collected by Johann 
Lhotsky in April, 1831. The species has been confused with | 
the genus Buddleia and with Callicarpa americana and speci- 
mens have been distributed under those names. The "Sellow 
2509" cited on page 476 of my monograph is a typographic er- 
ror for Sellow 2 and the Usteri s.n. [Herv. Esc. Polit. 
8. Paulo "2256" cited on the same page is a typographic er- 
ror for Herv. Esc. Polit. S. Paulo 223b. Some of the leaves 
on Glocker 578b are sparsely and irregularly dentate and are 
marked with glandular disks beneath. The locality of collec- 
tion of Netto e.n., Barbacena, is erroneously spelled 
"Barbacoa” on p. 475. 

Additional citations: BRAZIL: Ceard: Luetzelburg 26,1 
(Mu, N); Parahyba: Pohl 3481 (V); Bahia: Blanchet 4 fine 
286 (Cb, Op), 554 (Ob--2, F), 729 (Mi), 1064 (Gb), 1902 (Bm, 
P), 2064 (De), 5597 (Bm), 3399 [Herb. Monac. 1032 & 1447] 
(Bm, Br--2, Ch--2, De, F, Mu--2, P--2, V, Z--photo), s.n. 
[1857] (Cb); Casaretto 2022 [Macbride photos 24,614] (Cb); 
Glocker 540 (Bm, N--photo, Z--photo); Lhotsky s.n. [Herb. De 
Candolle 892] (Cb--isotype, Dc--isotype, N--fragment of iso 
type, Z--photo of isotype); Lockhart s.n. [Bahia] (Bm); 
Martius 2222 [Herb. Monac. 1028 & 1029] (Mu--2), s.n. [Herb. 
Monac. 1030] (Mu); Moricand s.n. (K); Salzmann 432 (De), 8. 
n. (E, K--2, P, V), s.n. [1631] (Ed); Sellow s.n. (K); Goy- 
az: Ule 335 (P); Minas Geraes: Olausen 552 (Bm, Br, Cb), 631 
(N, P=-2, Us), s.n. [Curvello] (le), s.n. [Aug.--April, 
1840] (Br--2), s.n. (Bm, Cb); Netto s.n. [Near Barbacena in 
1862] (A--photo, G--photo, Z--photo); Regnell 1.310x [1865] 
(N--photo, 8, Us, Z--photo), 1.310x [1867] (Us), 1.310xa 
(A--photo, B--photo, D--photo, F—-photo, N--photo, W-photo, 
Z--photo); Saint-Hilaire B?.2104 (P), s.n. (K, P—-2); Warm 
ing s.n. [Lagoa Santa] (Cp--3, F, N, P, Z--photo); Widgren 
s.n. [in 1845] (S--2); Parand: Dusén 1028a (A--photo, B= 
photo, D--photo, F--photo, G--photo, N--photo, W, W-photo, 
Z--photo); Jénsson 1028a (B, Cb, E, G, K); State undetermin- 
ed: Blanchet 79 (Bm), s.n. (K)3 Glocker 578b (Bm), 2166 
(Bm); Herb. Bernhardi s.n. (E); Herb. Imp. Vien. 154 (K), 
155 (K--2); Herb. Zuccarini s.n. [Herb. Monac. 1034] (mu); 
lund s.n. (De); Poeppig 34 (Z--photo); Pohl s.n. [Herb. Imp. 
Vien.) (N--photo, V, Z--2 photos), s.n. [Brasilia] (Bm, Br); 


1937 Moldenke, Notes on Aegiphila 233 


Sellow 2268 (Br), s.n. [Brasilia] (B--3), s.n. (De). SOUTH 
AMERICA: Country undesignated: Herb. Spruner s.n. (B). 


67. AEGIPHILA LONGIFOLIA Turez. 

Additional citations: COLOMBIA: Santander Sur: els s. 
n. [Ocana] (Z--photo); Schlim 688 (A--photo of isotype, B— 
photo of isotype, Cb--isotype, D--photo of isotype, F-- 
isotype, F--photo of isotype, G--photo of isotype, K-~iso- 
type, N--fragment of isotype, N--2 photos of isotypes, S— 
photo of isotype, W--photo of isotype, X--isotype, Z--2 
photos of isotypes); Department undetermined: Herb. Mus. 
Paris s.n. (P). 


93. AEGIPHILA LONGIPETIOLATA Moldenke. 

Additional citations: PERU: Loreto: Weberbauer 4667 (A-= 
photo of type, Be-photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, P--photo of type, S--photo of type, W--photo of type, 
Z-—-photo of type). 


21. AEGIPHILA LUSCHNATHI Schau. 

The Saint-Hilaire specimens cited below are anomalous in 
being densely velutinous throughout, especially on the 
younger parts, but this condition is approached on the very 
young parts of several other collections cited under this 
species in my original monograph. The species has occasion= 
ally been confused with the genus Pisonia and some herbari- 
um specimens distributed under that name! The Specimens ci- 
ted in my monograph as "isotypes" ought to have been called 
cotypes, since the species is based on several separate 
collections of Luschnath, no particular one of which was 
specifically designated as the type by Schauer. The "Gaudi- 
chaud 468" cited on pages 308 and 473 of my monograph 
should read "Gaudichaud 468, in part", since part of this 
collection is A. fluminensis. 

Additional citations: BRAZIL: Bahia: Blanchet 3273 (Ba— 
2, Br--2, Cb--2, F, Z--photo); Minas Geraes: Clausen 53, in 
part (Ob, P--2); Glaziou 15,326 (Bm, Br, Cb, Op, K, P, Z--2 
photos); Widgren 481 (Z--photo); Rio de Yaneiro: Barboza s. 
n. [1846] (Br); Beyrich s.n. (B); Burchell 734 (K), 1653 
(A--photo, B--photo, D--photo, F--photo, G--photo, K--2, 
photo, Z--photo), 2801 (A--photo, B--photo, D--photo, F-- 
photo, G--photo, K, We-photo, Z--photo); Clausen 53, in 
part (Ob--2, F); G. Gardner 5572 (Bm, K), 5 (Bm, K)3 
Gaudichaud 110 (P), 468, in part (Cb, P--2); Gleziou 865 
(Br=--2, Cp--2, K, N--photo, P, Z--photo), 8832 ET Oe 
--photo); Guillemin 53 (Ob), 630 (Cb--2, P); Lund s.n. [Rio 
de Janeiro] (Bm, Cp--2); Luschnath s.n. [Lagoa Sacaremo] (A 
--photo of cotype, B--2 cotypes, B--photo of cotype, Op-- 


234 PHYTOLOGIA No. 6 


cotype, D--photo of cotype, F--photo of cotype, G--photo of 
cotype, N--photo of cotype, We-photo of cotype, Z-~photo of 
cotype), s.n. [Brasilia] (E--cotype); Martius 1040 [Macbride 
photos 7879] (A--photo of cotype, B--photo of cotype, Br=-=3 
cotypes, Cbh--cotype, Dce--cotype, G--photo of cotype, N-- 
photo of cotype, Z--photo of cotype); Miers 3712 (Bm, K)3 
Miken s.n. (N--photo, Z--photo); Regnell 156 (P); Riedel & 
Luschnath 431 (L--2); Schott 4918 (V, Z--2 photos); Santa 
Catharina: Saint-Hilaire C*.1721 bis (P--2); Ule 4436 (B); 
State undetermined: Freyreiss s.n. (Z--photo); Gaudichaud 
630 (De); Herb. Ventenat s.n. (Cb); Pohl s.n. (Br); Raben 


149 (Br), 8 .n. (Cp). 


30. AEGIPHILA MACRANTHA Ducke. 

An additional synonym is Clerodendrum capitatum Klotzsch, 
in herb. [not C. capitatum (Willd.) Schum. & Thonn., 1827]. 
The species has been confused with Cordia sp. and some herb= 
arium sheets have been distributed under that name} 

Additional citations: TRINIDAD: Trin. Bot. Gard. Herb. 
1899 (R). BRITISH GUIANA: De la Cruz 2836 (D, E, G, 
photo, Z--photo); Sandwith 502 (B, K--2, Ut, W)3; M. R. 
Schomburgk 873 (C, Cb, N--photo, V, X, Z--photo), 1501 (A-- 
photo, Be-5, B--photo, D--photo, F--photo, G--photo, N-- 
photo, P, W-photo, Z--photo). FRENCH GUIANA: Mélinon 330 
(P), s.n. [in 1864] (P--4, Z--photo), s.n. [1877] (P--3)3 
Wachenheim 484 (P--2), 489 (Pe-2). BRAZIL: Pard: Ducke 
22,549 (B, K, P, S, Ut, X), s.n. [Herb. Rio de Janeiro 
18,949] (A--photo of isotype, B--photo of isotype, D--photo 
of isotype, F--photo of isotype, G-photo of isotype, K-- 
isotype, N--%3 photos of isotypes, Ut--isotype, W-photo of 
teh Z--2 photos of isotypes); Poeppig s.n. [Collares] 


129. AEGIPHILA MACROPHYLLA H.B.K. 

The specimen labelled with this name and discussed by me 
on pages 470--471 of my monograph proves to be Cormitia od- 
orata var. calvescens Moldenke. Since this specimen is from 
the Bonpland herbarium, it seems quite probable that Aegiph- 
ila macrophylla H.B.K. will eventually prove to be synony- 
mous with some species of Cormtia. The Berlin specimen, 
however, differs in so many respects from the original des=- 
cription of Aegiphila macrophylla that I do not think that 
it is part of the type collection. The actual type has not 
yet come to light. A specimen collected by H. M. Ridley in 
Porto Rico and distributed under the name "Egiphylla macro= 


phylla" has proved to be Callicarpa ampla Schau. 


75. AEGIPHILA MAGNIFICA Moldenke. 
Additional citations: NICARAGUA: Chinandega: 0. F. Baker 


— 


1937 Moldenke, Notes on Aegiphila 235 


204 (A--photo of type, B--photo of type, Ca--isotype, D-- 
photo of type, E~-2 isotypes, F--photo of type, G-photo of 
type, K--isotype, Mi--isotype, N—photo of type, P--photo of 
type, S--photo of type, W-photo of type, Z--photo of type), 
658 (N--photo, Z~-photo); Chontales: Seemann 87 (Bm, Z-- 
photo). COSTA RICA: Guanacaste: Tonduz s.n. (Herb. Inetit. 

sico-geogr. Nat. Costaric. 13,629] (B--2, Bm, P); San Jo- 
sé: Preted 11,176 (Cp); Tonduz 13,629 (K). PANAMA: Panamé: 
P. CO. Standley 29,797 (N--2 photos, Z--2 photos). 


71. AEGIPHILA MARTINICENSIS Jacq. 

An additional synonym for this species is Aegiphila mini- 
ma W. Wright, in herb. The species has occasionally been 
misidentified as Feramee occidentalis L. C. Rich., Catesbaea 
ep., Boussingaultia sp., Cestrum sp., and Psychotria SPs 
and specimens have been distributed under those names! There 
is no specimen of this species in the Linnean Herbarium, al- 
though the name Aegiphila martinicensis is often erroneously 
accredited to Linnaeus. The Funck & Schlim 81 cited below 

represents the "variabilis" form form of the species. Ramage re- 
cords the vernacular variant "bois cabrit® from St. Lucia. 
Otero says that it fruits in February and Sandwith describes 
the fruit as red in color. The species is often attacked by 
Hobsonia Ackermanni Pat. and by Guignardia prominens Earle 
[Laestadia prominens (Earle) Sacc. & D. Sacc.]. The Legit 
Anonymus ante ann. 1840 s.n., cited below, is anomalous eke 
its coarsely dentate lower leaf-blades and fine pubescence 
throughout. The "Spruce s.n." cited on page 476 of my mono- 
graph as representing this BDeGAOF is an error and should be 
deleted, and the "Grebs s.n." cited on page 482 should read 
"Krebs s.n.". The Britton & n & Cowell 594 cited on pages 383 
and 472 as ver. oligoneura | is better regarded as represent- 
ing a small-leaved form of the true species. The Hort. Kew. 
s.n. is strongly like A. filipes, but the specimen is n is only : & 
_ fragmentary one and cultivated specimens of A. martinicensis 
are in general similarly anomalous in leaf-texture and 
shape. 
fj Additional citations: CUBA: Havana: De la Ossa s.n. 
[1825] (De--2). JAMAICA: Bertero 2046 (B); Swartz e.n. (Z-- 
photo); W. Wright e.n. (Bm). PORTO RICO: Britton, Britton, & 
_ Brown 6786 (0b); Collector undesignated s.n. (P)$ Gook & 
Collins 291 (w); A. A. Heller 4670 (B. B--photo, Cb, D-— 
oT Ed, Es, Le, Mi, N--photo, W, W--photo, Z--photo), 
6444 (Es); Heller & Heller 276 (K, Z--2 photos), 825 (B, Z— 
photo), 964 (K, Z—photo); Hioram s.n. [Rf{o Piedras] (P); 
Ledru s.n. (Ed); Otero 78 (N, N, Ne-seed); Plée 172 (P); H. Ne 
Ridley s.n 8.n. (Ob--5); Riedlé 202 (P); aah (Herb. A. L. Juss- 
feu eg (P); Sintenis 1185 (3). 1295 95b (B), 2674 (B, Bas 
Kk), 6492 (w), a.n. [1895] (01); aa oe (B), oe (B), 995 


236 PY TOs oO Sis No. 6 


(B). ST. JOHN: greted s.n. [1846] (Cp--2). ST. THOMAS: Col- 
lector undesignated s.n. (P); Krebs s.n. (Z--photo); Riédlé 
s.n. (P). ST. EUSTACHE: Boldingh 246 (Le), ra [189] (ut), 
894 [410] (Ut). ST. CROIX: Benzon 127-3713 (Cp); Britton, 
Britton, & Kemp 114 (Cb, Z--photo); Collector undesignated 
s.n. (Z--photo); Herb. Univ. Christian. s.n. (01); Ravn s.n. 
(Cp); L. C. Richard s.n. (P); J. B. Thompson 239 (A—photo, 
Cb, F--photo, G--photo, N, Z—photo), 503 (G--photo), 719 (Z 
--photo). ST. KITTS: Britton & Cowell 116 (Z--photo), 120 (Z 
--photo), 332 (B, K, Z--photo), 5904 (B, K, N); Eggers 133 (2 
--photo), s.n. [Decbr. 1882] (B, Vu); Forsstrém s.n. (S--2, 
Z--photo). ANTIGUA: Ponthieu s.n. [Ind. occid.] (Cb-—2); 
Rose, Fitch, & Russell 5341 (B, Z--photo); Wullschlugel 42 
[Herb. Monac. 1037] (B, Ma), 424 [Herb. Monac. 1039] (Mia). 
MONTSERRAT: Ryan s.n. (Cp--6); Shafer 162 (Cp), 197 (2-- 
photo). GUADELOUPE: Badier 176 (De); Duchassaing s.n. [Gua- 
deloupe] (B, P--3, V, Z--photo); R. P. Duss 303 (P), 2389, 
in part (E, Z--photo); Forsstrim s.n. (S); Funck & Schlim 81 
(Cb, P); Grisebach s.n. (BE, K); Herb. Lamarck s.n. [Guadel- 
oupe] (P); Herminier s.n. (X--4); Perrottet s.n. [18 Juin 
1824] (Cb), e.n. [Juillet 1641] (Bm); Picard s.n. (P); Quen- 
tin 17 (P); Read s.n. (B); L. C. Richard s.n. (N--photo, P, 
Z——photo); Stehlé 1247 (N). DOMINICA: G. P. Cooper 60 (Mi), 
80 (N), 167 (Mi); Eggers 501, in part [Herb. Monac. 3822] (B 
==) 0M, ies i; Ya), 786 (B, Cp); Imray 85 (K), 119 (Z— 
photo), 420 (K); Kraus 273 (Ed); Nicholls 94 (B); Ramage s. 
n. [April 24, 1888] (Bm, K), s.n. (Aug. 22, 1888] (Bm), s.n. 
(B). MARTINIQUE: Bélanger 260 (Ob--2), 290 (P), 571 (P); 
Collector undesignated s.n. (Cp); R. P. Duss 1966 (B, Z--2 
photos), 2389, in part (B--2); Hahn 125 (Bm, Br, Cb--5, K, 
Le, P), 491 [Herb. Monac. 1448] (B, Bm, Gb--2, K--2, Le, Mu, 
P--3, Us); Isert 87 (Op); Jacquin s.n. (Bm--type); Kohaut s. 
n. (B); Plée s.n. [Martinique] (P--3, Z--photo); Sieber Fl. 
Mart. 78 [Herb. Monac. 1035] (B--2, Bm, Cp, E, K, Ma, X)3 
Terrasson 62 aie Be jer ti 5034b ] pire ST. LUCIA: 
Lambert s.n. (Dc); Lee a.n. [Dec. 2, 1887] (W); Ramage s.n. 
(B), s.n. (Sept. 23, 1888] (K). BARBADOS: Eggers 7155 (B, 
Le); Sandwith s.n. [Barbados] (K); Waby 72 (3. F = Warm 
ing 134 (Cp, Z--photo); a. Wright s.n. (Br). ST. VINCENT: 
Caley s.n. (Jan. 1823] (Cb); Collector undesignated s.n. 
[Aug. 1824] (Z--photo); Eggers 66 i» in part (Py; Guild 

s.n. (B); Smith & Smith 336 (B, Ba, Ed, Z--photo), 1601 (B, 
Z-~photo). GRENADA: W. E. Broadway 142 (B), 1410 (B), s.n. 
[St. George's] (A, B--photo, £, G--photo, Z--3 photos), s.n. 
{Mount Parnassus, June, 1906] (E--2), s.n. [Oct. 31, 1905; 
Hort. Thenensis 1.4507] (Br), s.n. [St. Paul's] (Ed—-2, Mi), — 
s.n. [Year 19067] (E); Eggers 6049 (B, P, Z--photo); G. We. 
Smith 93 (B). TRINIDAD: W. &. Broadway s.n. [River Estate] 
(B==photo 3 D--photo, N--photo, W--photo); Sieber Fl. Trinit. 


1937 Moldenke, Notes on Aegiphila 237 


85 [Herb. Monac. 1036 & 1449] (B--2, Cbh--2, De, E, Ed-=2, 

Le, Mu--2, V--2, Z--7 photos). WEST INDIES: Island undesig- 

nated: "B. S." s.n. (K); Collector undesignated 426 (Us), s. 
(Ed); 


n. (Cb, P); Forster s.n. ["India"] Forsyth s.n. (Dc); 
Herb. Adanson s.n. (P=-7); Herb. Puerar s.n. Figoh] (De); 


Ponthieu s.n. [Ind. occid.] (Bm); West s.n. (Dc). PANAMA: 
Bocas del Toro: G. P. Cooper 571 (K, W, Z--photo). COLOMBIA: 
Méta: Pennell 1650 (Z-~photo). VENEZUELA: Zulia: Mocquerys 
910 (P--3); H. Pittier 10 (B, Cb, Z--photo); Mérida: W. 
E. one 19 (G, Z--photo), 479 (G). FRENCH GUIANA: Leb- 


SSeS ee 


Seaforth, West Indies] (B--photo, Ob--2, N--fragment, N-=-2 
photos, S--photo, Z--2 photos), s.n. [A. B. Lambert, 1808; 
fa) Seaforth, West “aan e Hort. Kew. s.n. [1785] 
Bm); Hort. Liverpool s.n. (K); France: Jard. Bot. Paris s. 
n. [de l'isle de Cuba] (Cb); Jard. Malmaison s.n. (P); 
Perrottet s.n. [Jardin des Pl. de Paris, 1518] (Cb); Neth- 
erlands: Herb. Persoon s.n. (Le); Belgium: Collector undes- 
ignated s.n. (Br); Nyst s.n. (Br); Germany: Hort. Monac. s. 
n. iGarb . Monac. 1038] (Mu); Ceylon: Hallier f. 0.240 (Le). 
LOCALITY OF COLLECTION UNDESIGNATED: Baudin s.n. (P); Herb. 
Colsmann s.n. (N--photo, Z--photo); Herb. Hooker a.n. (B); 
Herb. Hornemann s.n. [Ex Ind. occid.?] (Cp--2); Herb. A. L. 
Jussieu 8.n. (P--3); Herb. Liebmamn s.n. (Cp); Herb. Vahl 
Ben. (Cp); Legit Ano 8 ante ann. 1640 s.n. (Ut); Mus. 
Bot. Berol. s.n. ["Bras.?"] (N--photo, Z--photo). ILLUSTRA- 
TIONS: Copy of Andr. Bot. Rep. t. 578 (Ba); Lamarck, Illus- 
tr. 1: t. 70, f. 1. 1791 (B); Unidentified illustration WN). 


7la. AEGIPHILA MARTINICENSIS var. OLIGONEURA (Urb. )Moldenke 
Hahn records the vernacular variant "bois cabrite" from 
Martinique. The Britton & Cowell 594 cited on page 383 of 
my monograph is better considered to represent a form of 
typical A. martinicensis. The "Eggers 6617" cited on pages 
383 and 473 should read "Eggers eat] >» in part", since some 
sheets of this collection are typical A. martinicensis. 
Additional citations: JAMAICA: W. Harris 8166 (A--photo 
of isotype, B--type & isotype, B--photo of isotype, Bn-~ 
isotype, D--photo of isotype, F-—-photo of isotype, G-~-photo 
of isotype, K--photo of isotype, N--isotype, N=--2 photos of 


isotypes, W—photo of isotype, Z--photo of isotype). DOMIN- 
ICA: Eggers 5Ola (Br, P); Lloyd 606 (N--photo, Z--photo); 


* 


- 
} 


} 


MARTINIQUE: Hahn 1236 (B, Bm, Br, Cb--2, N--2 photos, P--2, 


'X, Z--2 photos). ST. VINCENT: Eggers 6617, in part (B); 
Smith & Smith 336a (N--photo, Z-~photo), 


153 (Bm, N--photo, 


. Z--photo). LOCALITY OF COLLECTION UNDESIGNATED: "West Ind- 


ies, Nov. 17--17" (N--photo, Z--photo). 


238 PY €-O:b-O G Ek No. 6 


38. AEGIPHILA MEDITERRANEA Vell. 

This binomial is sometimes erroneously to Turczaninow. 
The type specimen of A. cestrifolia has been examined in the 
herbarium of the British Museum. The Rodriguez specimen cit- 
ed below bears a label inscribed "Misiones: Santa Ana", but 
probably this is meant to be the Santa Anna on Grande Is- 
land, Rio de Janeiro, and not 4 locality in Misiones, Argen- 
tina, which would be far out of the known range of the spe- 
cies. 

Additional citations: BRAZIL: Rio de Janeiro: Boog s.n. 
(Z=--photo ); G. Gardner 100 (Bm--2, Cb—-2, Ed--2, F, K, Ne=3 
photos, P, V, Z--3 photos); Glaziou 1546 (Br, Cp, P), 11,337 
(Br, Cb, Cp, K, Pe=2, Z--photo); Martius 166 [Herb. Monac. 
1040] (Ma); Miers 3096 (Cb); Pohl s.n. (Herb. Imp. Vien. 
151] (K, Z--photo); Riedel 0,31 i; 0,37 (L); Riedel & 
Luschnath 1005 (L--2, N)} Rodriguez s.n. [Lillo 10,448] (G); 
Schenck 2109 (N--photo, oa ameteys Schott 4916 (Cb, V, Z— 
photo); Swainson s.n. (Z--photo); Ule 4854 (B); Widgren 1151 
(Us, Z--photo), 1225 (Br), s.n. (A--photo, B--photo, D-- 
photo, F--photo, G--photo, N--photo, #-photo, Z--photo). 
CULTIVATED: Cameroons: H. Winkler 628 (B); Versuchsanstalt 
Kamerun 270 (B, Us). 


12. AEGIPHILA MEDULLOSA Moldenke. 

The type collection of this species is Saint-Hilaire ¢.50 
and not "Co. 50" as inaccurately written in my monograph. 

Additional citations: BRAZIL: Rio de Janeiro: Riedel & 
Lund 0,30 (L--2, N--fragment, Z--2 photos); Saint-Hilaire 
C.50 (A—photo of type, B--photo of type, D--photo of type, 
F-—-photo of type, G--photo of type, K--photo of type, N— 
photo of type, S--photo of type, W--photo of type, Z-=-photo 
of type). 


65. AEGIPHILA MEMBRANACEA Turcz. 

This species has occasionally been confused with the gen 
us Clerodendrum and some herbarium specimens have been dis- 
tributed under that name! 

Additional citations: COLOMBIA: Chocé: Triana 3713, in 
part (Bm). SURINAM: Hostmann 89 (A--photo of isotype, B— 
isotype, B--photo of isotype, Bm--2 isotypes, Cb--isotype, D 
--photo of isotype, F--photo of isotype, G--photo of iso- 4 
type, K--2 isotypes, N--2 photos of isotypes, P--isotype, Ut 
--isotype, V--2 isotypes, W-photo of isotype, Z--35 photos 
of isotypes); Hostmann & Kappler 89, in pert [Herb. Monac. 
1188] (Ma). FRENCH GUIANA: Benoist 740 (P), 786 (P), 787 (P 
--3); Jelski s.n. [Cayenne] (B); Mélinon 83 (P), 89 (N—— 
photo, Z--photo), 111 (P), 433 (P, Z--photo), 436 (P), 486 
(P), s.n. [in 1864] (Bm, Z--2 photos), s.n. [in 1877] (Bm, 
Br, Ob, F, K, N=-photo); Sagot s.n. (P--2); Wachenheim 100 


1937 Moldenke, Notes on Aegiphila 239 


(Bm, F, P--2), 188 (P), s.n. [environs de Godebert] (K, Z-- 
photo). VENEZUELA: Falcon: Jahn 384 (Z--photo). PERU: Lore- 
to: Tessmann 4666 (B). 


83. AEGIPHILA MOLLIS H.B.K. 

Additional synonyms ere Cornutia velutina Hayek in Engl. 
Jahrb. 42: 172 (1908) and Aegiphile salutaris var. lutea 
Moritz, in herb. The type of Cornutia velutina was collected 
by Friedrich Carl Lehmann (No. 6692) in El Cauca, Colombia, 
and several isotypes have been examined. The collection 
matches typical material of Aegiphila mollis perfectly. The 
type of A. salutaris [Bonpland 983] and another cotype of A. 
Mutisii [Bonpland 8.n.}| have now been examined. Neither of 
these can be separated from A. mollis. Dugand reports that 
the present species is a frequent shrub in second-growth, 
inhabiting clayey and stony soil. He describes it as a shrub 
or shrubby tree, 3--5 m. tall, with a trunk to 15 cm. in di- 
ameter, grayish rugose bark, and abundant white flowers, the 
corolla soon falling off. The Humboldt & Bonpland s.n. spec- 
imen from the Berlin herbarium, cited in my monograph as the 
type, is actually only an isotype. The Venezuelan state An- 
zoategui is misspelled "Anzoatequi" on page 405. 

Additional citations: COLOMBIA: Atldéntico: Dugand 101 (F, 
Y), 469 (F, X), 659 (Y)s Elias 1102 (N--2, W); Magdalena: H. 
H. Smith 868 (Bm, Br, Cb--2, E, Ed, K, Le, N, P, Ut, Z-- 
photo), 870 (E, G-photo, K, Z--photo), 1860 (Bm, Cb—2, E, 
K, P, Z--photo); Purdie s.n. (Santa Marta] (K, N--photo, Z-- 
2 photos); Cundinamarca: André K.1554 (kK); Bonpland s.n.(N-- 
photo, P--2); Hartweg s.n. |Fusagasuga, near Bogota] (K, Z—— 
photo); Mutis 425 (B--photo, F--photo, G--photo, N~-photo, Z 
--photo); Triana 2082 (Bm--2, Br, Cb--2, Ed, P), 3713, in 
part (Bm); Bolivar: Elias 581 (Z--photo); Tolima: André 
K.1556 (K); El Cauca: Karsten s.n. [Papayan] (A--photo, B-- 
photo, D--photo, F--photo, G--photo, N--photo, We-photo, Z-- 
photo); Lehmann 6692 (B, B--descr., F--photo, F, Ke-=2, L, 
W); H. Pittier 626 (Z--photo); Department undetermined: 
Goudot 2 (P). VENEZUELA: Lara: Saer 723 (Ve); Carabobo: H. 
Pittier 7910 (G); Aragua: Fendler 8435 (G, N--photo, Z--2 
photos), 2052 (Z--photo); Vogl 1177 (Mu), 1179 (Mm), 1180 
(Mu), 1181 (Mu--2), 1183 (Mu); Miranda: H. Pittier 7855 (G); 
Guarico: Geay s.n. [Guaritico] (P); Sucre: Funck 645 Ob, P, 
V)3 Moritz 365 (Bm); Anzoategui: Otto 856 (B, K, Z-=2 
photos); Monagas: Moritz 1910 (Bm); Amazonas: Bonpland 
(N--photo, P, Z--photo); State undetermined: Chaffanjon 245 
(P); Eggers 13,464 (Cp, Z--photo); Humboldt & Bonpland s.n. 
[Woods of the Orinoco] (A--photo of isotype, B--photo of 
isotype, D--photo of isotype, F--photo of isotype, G--photo 
of isotype, N--photo of isotype, W-photo of isotype, Z— 
photo of isotype). BOLIVIA: Santa Cruz: D'Orbigny 1086 (Cb); 


240 PHY PfOwoeers No. 6 


Steinbach 3168 (Z-—photo). LOCALITY OF COLLECTION UNDESIGNA- 
TED: Herb. Baillon s.n. (P); Lehmann B.T.705 (Le, N--photo, 
V; Z--photo). ILLUSTRATIONS: H.B.K. Nov. Gen. & Sp. Pl. 23 
t. 130. 1817 (P). 


83a. AEGIPHILA MOLLIS var. INTERMEDIA Moldenke. 

Additional citations: COLOMBIA: Magdalena: H. H. Smith 
329 (A--photo of type, B--photo of type, Bm--isotype, Br-- 
isotype, Cbh--2 isotypes, D--photo of type, E--isotype, Ed— 
isotype, G--photo of type, K--isotype, K=--photo of type, N-= 
photo of type & 2 photos of isotypes, Pe-isotype, We-photo 
of type, Z--photo of type & 2 photos of isotypes), 869 (N—- 
photo, res Bolfvar: Killip & Smith 14,296 (N-=photo, 
Z=—photo). 


13. AEGIPHILA MONSTROSA Moldenke. 

The "Aegiphila fasciculata Donn. Sm." which Standley des- 
cribes in Field Muse. Pub. Bot. 10: 334--335 (1931) and ill- 
ustrates on plate 57 is A. monstrosa, not Dermatocalyx par- 
viflorus Zret. as erroneously stated by myself in Phytologia 
1: 200 (1937). The Cook & Griggs 521 [U. S. Net. Herb. 
408,225] said to be A. fasciculata by J. D. Smith in Bot. 
Gaz. 57: 425 (1914) is actually A. monstrosa. Schipp reports 
that the species under discussion is a small tree growing in 
forest shade, that it is quite common, and that the flowers 
are highly perfumed. The vernacular name "hulub" has been re 
corded from British Honduras. . 

Additional citations: GUATEMALA: El Petén: Lundell 1492 
(Mi); Alta Verapaz: Cook & Griggs 521 (W); H. V. Johnson 520 
(N--photo, Z--photo); Izabal: P. 0. Standley 25,114 (Z-- 
photo); Department undetermined: Skinner s.n. (K). HONDURAS: 
Atldntida: P. C. Standley 52,717 (N--2 photos, Z~-3 photos), 
55,176 (B-photo, D--photo, N--2 photos, P--photo, S--photo, 
We-photo, Z—-2 photos); Cortés: Carleton 422 (A--photo of 
type, B-photo of type, D--photo of type, F--photo of type, 
G=-photo of type, N=--photo of type, P--photo of type, S-= 
photo of type, W-photo of type, Z--photo of type). BRITISH 
HONDURAS: H. H. Bartlett 11,941 (Mi), 15,011 (Mi); Gentle 
2635 (Mi,.N), 404 (Mi, W), ate (A, E, I, Mi, N, S--2); Lund 
ell 146 (K, Mi); Schipp 1085 (A, Bm, Ca, Ob, E, K, Mi, N,S). 


) Brittonia 1: 245--477. 1934. 

) Phytologia 1: 182--208. 1937. 

) Housrd, Zoocéd. Pl. Amer. Sud 551-352. 1933. 
) Houard, Zoocéd. Pl. Amer. Sud 344, 1933. 


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PHY TOLOGIA 


Designed to expedite botanical publication 


VoL. 1 September, 1937 No. 7 


CONTENTS 


C. L. Lunpety: Studies of Mexican and Central American Plauts--L1I . 241 


H. N. Monpenke: Additional Notes on the Genus Aegiphila-- Il] . . 248 


nee ae ee = ae 
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Published by H. A. Gleason and Harold N. Moldenke 
The New York Botanical Garden 
Bronx Park, New York, N. Y. 
Price of this number: 50 cents; per volume $5.00 in advance. 
Vol. 1, number 6, was issued June 14,1937 | 


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STUDIES OF MEXICAN AND CENTRAL AMERICAN PLANTS -- III (a) 


CO. L. Lundell 


Of the six novelties described, two are from the writer's 
1936 collections on the Carnegie-Michigan expedition to 
British Hondures. The other four were discovered in the 
highly interesting and valuable Mexican collections of Mr. 
Eizi Matuda and Professor C. Conzatti. All the specimens 
cited are in the Herbarium of the University of Michigan. 


PICRAMNIA MATUDAI Lundell, sp. nov. 

Arbor, ramulis, petiolis, et pedunculis dense fulvopubes- 
centibus. Folia magna, 25--45 cm. longa; foliolis 3--9, al- 
ternis, ovatis, lanceolato-oblongis, vel lanceolato- 
ellipticis, 10--18.5 cm. longis, 4.2-~6.8 cm. latis, apice 
acuninatis, acumine obtuso vel acutiusculo, basi rotundatis, 
obtusis, vel cuneatis, supra glabris, subtus fulvopubescen- 
tibus; petiolulis crassis, dense pubescentibus, 4—-5.5 mm. 
longis. Inflorescentiae axillares, 15--25 cm. longae. Flores 
masculi glomerato-reacemosi, glabri; pedicellis 1--1.2 mm. 
longis. Sepala 3, raro 4, latissime ovata vel ovato-orbicu- 
laria, 1.3--1.8 mm. longa, integra vel eroso-dentata. Petala 
3, rero 4, oblonga vel oblanceolato-oblonga, 2--2.3 mm. lon- 
ga. Stamina 3, rero 4, 3.5--3.9 mm. longa; filamentis crass—- 
is. Flores femineos non vidi. Fructus obovoideus, ca. 1.5 
cm. longus. 

A tree with fulvous and copiously pubescent twigs, petio- 
les, and peduncles; leaves large, fulvous, as much as 45 cm. 
long; leaflets somewhat inequilateral at base, alternate, 
subcoriaceous, 3 to 9, the lower ones ovate, the upper ones 
lanceolate-oblong or lanceolate-elliptic, 10 to 18.5 cm. 
long, 4.2 to 6.8 cm. wide, apex acuninate, the acumen obtuse 
or acutish, base of lower leaflets rounded, base of upper 
leaflets obtuse to cuneate, glabrous above, persistently 
pubescent beneath, costa and lateral nerves nearly plane a- 
bove, prominent beneath: petiolule thick, densely pubescent, 
4 to 5.5 mm. long; inflorescence axillary on old wood, the 
rachis simple, 15 to 25 cm. long, densely fulvous-pubescent; 
staminate flowers glomerete-racemose, glabrous; pedicels 1 
to 1.2 m. long; sepals 3, rarely 4, broadly ovate or ovate- 
orbicular, 1.3 to 1.8 mm. long, entire or erose-dentate; 
petals 3, rarely 4, oblong or oblanceolate-oblong, 2 to 2.3 
mm. long; stamens 3, rarely 4, 3.5 to 3.9 mm. long, the fil- 
ements thick; fruits glabrous, obovoid, about 1.5 cm. long; 
fruiting pedicels 2 to 3 mm. long, pubescent. 

Type in the Herbarium of Ker University of Michigan, Eizi 

241 


242 PHY! £: 0.i7 O.6-1°.4 No. 7 


Matuda 428, collected on Mt. Orando, Chiapas, Mexico, Decem 
ber 17, 1936. 

P. Matudai is a well-marked species distinguishable at 
once from the other species with three-parted flowers by its 
large pubescent leaves esd axillary inflorescences borne on 
old wood, 


Notes on the species of Exothea 


Heretofore only two species have been know in the genus 
Exothea. The widespread E. paniculata (Juss.) Radlk. occurs 
in Florida, the West Indies, and Central America. E. copal- 
illo (Schlecht.) Radlk. is restricted apparently to southern 
Mexico. E, diphylla (Standl.) Lundell, an endemic tree in 
the Yucatan Peninsula, is also referable to the genus. The 
three species may be satisfactorily separated as follows: 


Leaflets 2; petioles less than 1 cm. long.....1. E. diphylla 
leaflets 2 to 8 (usually 4); petioles generally 
more than 2.5 cm. long. 


Leeflets entirely gleabrous........... .-2. E. paniculata 
Leaflets barbate beneath ir. the axils of 
the nerves....... ec ec seers cecccees «+d. E. copalillo 


1. EXOTHEA DIPHYLLA (Standl.) Lundell, comb. nov. 

Talisia diphylla Standl., Field Mus. Bot. 8: 21. 1930. 

A tree reaching a height of 20 meters; flowers white. Oc- 
casional in sapodilla forest, and in marginal forest sur- 
rounding pine lands; endemic in the Yucatan Peninsula. 

Specimens examined: MEXICO: Campeche: Tuxpefla, Lundell 
1347. BRITISH HONDURAS: Corozal District: Gentle 646. Belize 
District: liaskall Pine Ridge, Gentle 1147; vernacular neme 
"wild kenep". 

The flowers of E. diphylla are very similar in structure, 
size and pubescence to the flowers of E. paniculata and E. 
copalillo. In fact, the flowers of all three species are 
strikingly alike. 


2. EXOTHEA PANICULATA (Juss.) Radlk. 

A medium-sized tree reaching a diameter of 30 cm.; bark 
smooth and gray; flowers cream-colored; fruits reddish 
brown. Occasional in forest; Florida, ‘est Indies, British 
Honduras, and Guatemala. 

Specimens examined: FLORIDA: Ko Name Key, J. H. Simpson 
in 1891. Without locality, A. P. Garber in 1877. WEST ItiD- 
IES: Behemas: New Providence, iim. Cooper ] in 1859. SAITISK 
HONDURAS: Toledo District: Pueblo Viejo, Schipp 1263; Cenip 
32, British Honduras-Guatemala boundary survey, Schipp 1266. 


a 


a ee ee ee 
¥ 


1937 Lundell, Mexican & Central American Plants 243 


GUATEMALA: Department of Petén: Uaxactun, Bartlett 12250, 
12560. ehage of Santa Rosa: Malpais, Heyde and and Lux 
(Donnell Smith 6294). 


3. EXOTHEA COPALILLO (Schlecht.) Radlk. 

A medium-sized tree; flowers white. In forest, Vera Cruz, 
Jalisco, and San Luis Fotosi. 

Specimens examined: MEXICO: Jalisco: Santa Cruz de Vall-_ 
arta, Mexia 1249. Vera Cruz, Purpus 15240. 


MELIOSMA KMATUDAI Lundell, sp. nov. 

Arbor glabra. Folia coriacea, integra, petiolis 6--10 m. 
longis; laminis anguste lanceolato-oblongis, 7--12 cm. long- 
is, 1--1.9 cm. latis, apice acuminatis, base longe acuminat- 
is. Inflorescentiae terminales, paniculatae, 6--11.5 cm. 
longae. Pedicelli 1--3.5 mm. longi. Sepala 5, inaequalia. 
Petala 5, inaequalia, 3 exteriora suborbicularia, ca. 2 m. 
longa, 2--3 mum. lata, 2 interiora minuta, bifida, puberula. 
Staminodia 3. Stamina 2. Discus ovarium subaequans, margine 
denticulis linearibus obsitus. Overium 2-loculare. Stylus o- 
verium subaequans. 

A tree, glabrous throughout; branchlets slender; leaves 
clustered at apices of branchlets, subopposite, coriaceous; 
petioles slender, 6 to 10 mm. long; leaf-blades very narrow 
ly lanceolate-oblong, 7 to 12 cm. long, 1 to 1.9 cm. wide, 
entire, apex acuminate, base long-acuminate, costa prominent 
on both surfaces, reticulate-veined, paler beneath; inflor- 
escence terminal, paniculate, 6 to 11.5 cm. long; flowers 
pedicellate, the pedicels 1 to 4.5 mm. long; prophylla ob- 
long-slliptic, about 1 mm. long, ciliolate, deciduous very 
early; sepals 5, unequal, ciliolate, the outer 2 broadly ov- 
ate, the inner 3 suborbicular, larger, about 1.4 mm. long; 
petals 5, unequal, the 3 outer (opposite the staminodia) de- 
pressed-orbicular, about 2 mm. long, 2 to 3 mm. wide, the 2 
inner (adnate to base of fertile filaments) minute, deeply 
bifid, puberulous; fertile stamens 2; anthers longitudinally 
dehiscent; disk thin, nearly equaling ovary in length, bear- 
ing slender linear teeth; ovary glabrous, 2-celled; style 
glabrous, equaling or slightly exceeding ovary in length; 
Stigmas connate. 

Type in the Herbarium of the University of liichigan, Eizi 
Matuda 526, collected on Mt. Orando, Chiapas, hexico, Decem— 


ber 19, “1936, 


M. liatudai is closely related to M. idiopoda Blake from 
which it differs obviously in having smaller and much nerr- 
ower leaves and glabrous panicles. It appears to be related 
to M. oaxacana Standl. also, but may be distinguished read- 
ily “from thet species by its much smaller flowers, narrower 


244 PHY TO Ls oe No. 7 
leaves, and the absence of pubescence. 


SOUROUBEA TRIANDRA Lundell, sp. nov. 

Frutex epiphyticus, glaber, S-metralis. Folia subcoriac- 
ea, petiolis 4--8 mm. longis; laminis oblanceolatis vel ob- 
lanceolato-obovatis, 5.5--10.5 cm. longis, 2--4 cm. latis, 
apice rotundatis vel obtusis, minute apiculatis, basi cunea- 
tis. Inflorescentiae terminales, racemosae, 8--16 cm. long- 
ae, ca. 2.2 cm. latae, multiflorae, pedicellis 7--9 mm. lon- 
gis; bractea sub calyce inserta, tubuloso-cylindrica, 8--10 
mn. longa, breviter stipitata; ostiolo lateraliter breviter 
2-auriculato; bracteolis 2. Sepala 5, suborbicularia, 1.2=- 
1.8 mm. longa. Corolla ca. 4.5 mm. longa, ad mediam partem 
gemopetala, lobis 3 vel 4, apice obtusis. Stamina 3 vel 4, 
filamentis crassis, ca. 2.1 mm. longis, basi complanatis, 
intus ad corollae basin applicatis, margine inter se contig- 
uis, apice attenuatis; antheris ovato-cordatis, ca. 1 m. 
longis. Ovarium 3- vel 4~loculare. 

An epiphytic shrub 5 meters high, glabrous throughout; 
branchlets slender, 2 to 3 mm. in diameter; leaves subcoria- 
ceous; petioles canaliculate, 4 to 8 mm. long; leaf-blades 
oblanceolate or oblanceolate-obovate, 5.5 to 10.5 cm. long, 
2 to 4 om. wide, apex rounded or obtuse, minutely apiculate, 
base cuneate, costa prominent beneath, the primary nerves 
inconspicuous; inflorescence terminal, erect, racemose, 8 to 
16 cm. long, about 2.2 om. wide, many-flowered; pedicels 
slender, 7 to 9 mm. long; bract tubular-cylindrical, 8 to 10 
mm. long, suspended obliquely from base of calyx, short sti- 
pitate (the stipe about 1.8 mm. long or shorter), the aper= 
ture laterally 2-auriculate, the auricles very short; bract- 
eoles 2, ovate-orbicular, about 1.2 mm. long, closely app- 
ressed to calyx; sepals 5, suborbicular, 1.2 to 1.8 m. 
long, imbricate; corolla about 4.5 mm. long, 3- or 4-lobed 
to the middle, splitting irregularly at anthesis and reflex- 
ed, the lobes obtuse; stamens 3 or 4, attached to base of 
corolla; filaments thick, broad and flattened below with the 
margins touching, narrowed above, about 2.1 m. long (inclu- 
ding portion adnate to corolla); anthers ovate-cordate, ob= 
tuse, about 1 mm. long; ovary 3— or 4—celled, ovoid; stigma 
sessile, 

Type in the Herbarium of the University of Michigan, C. 
L. Lundell 6492, collected in advanced forest on limestone 
hill near Cohune Ridge, El Cayo District, British Honduras, 
July 13, 1936. 

Additional specimens examined: GUATEMALA: Puerto Barrios, 
Deam 6020. 

S. triandra is remarkable in having flowers with corolla 
3- or lobed, 3 or 4 stamens, and ovary 3- or 4-celled. As 
far as the writer has been able to determine from published 


——-— = 


it ——— wr 


1937 Lundell, Mexican & Central American Plants 245 


descriptions and dissections of flowers, all the other 
species knowm in the genus regularly have S-parted flowers. 
In both the type collection and the Deam collection, the 
flowers of S. triandra consistently differ in this respect. 
A majority of the flowers have only 3 corolla lobes, 3 sta- 
mens, am a 3-celled ovary. The bract is typical of the gen- 
us. 

S. triandra is closely related to S. exauriculata Delp. 
Aside from the floral differences already mentioned, it may 
be distinguished further from that species in its much smal- 
ler flowers, glabrous inflorescence, narrower leaves, short- 
stipitate bract attached at base of calyx, and in other 
noteworthy characters. 

The Deam collection from Puerto Barrios, Guatemala, was 
identified by J. Donnell Smith as Ruyschia mexicana Baill., 
a species regarded as a synonym of S. exauriculata by Witt- 
mack (Fl. Bras. 12: 254. 1879) and by Hemsley (Biol. Centr. 
Amer. Bot. 1: 92. 1879). In the original description of R. 
mexicana, the flowers are described as having 5 stamens and 
a S-celled ovary, hence S. triandra differs in those import- 
ant and in other minor characteristics. 


RUYSCHIA ENERVIA Lundell, sp. nov. 

Frutex epiphyticus, glaber, 3—-metralis. Folia coriacea, 
oblanceolata vel oblanceolato-oblonga, 8--12 cm. longa, 2-=- 
4.2 cm. lata, apice obtuso-apiculata, basi obtusiuscula vel 
rotundata, costa planiuscula, nervis lateralibus inconspicu- 
is; petiolis 3-~8 mm. longis. Inflorescentiae terminales, 
racemosae, pauciflorae, ca. 8 cm. longae, glabrae, pedicel- 
lis 3--4.5 mm. longis; bractea sub calyce inserta, intus 
convexa, extus complariata, tumida, apiculata, 3--4.5 mn. 
longa, 3-~4 mm. lata, ca. 2 mm. crassa; bracteolis 2. Sepala 
5, coriacea, imbricata, suborbicularia, 4-4.5 mm. lata, 2-- 
2.5 mm. longa, spice rotundata. Petala 5, imbricata, coriac- 
ea, libera. Stamina 5, filamentis basi complanatis, lanceo- 
latis, intus ad corollae basin adnatis. Ovarium subglobosun, 
5—costatum, 2=-loculare. 

A glabrous epiphytic shrub 5 meters high; branchlets 5 to 
5 mm. in diam.; leaves coriaceous, opaque, oblanceolate or 
oblanceolate-oblong, 8 to 12 cm. long, 2 to 4.2 cm. wide, a- 
pex obtuse-apiculate, base obtusish or rounded, costa plane 
or nearly so, the lateral nerves invisible beneath, absent 
or faintly visible above, glands few and scattered, margin 
revolute; petioles canaliculate, stout, 3 to 8 mm. long; in- 
florescencs racemose, terminal, few-flowered, about 8 cn. 
long, glabrous; pedicels stout, 3 to 4.5 mm. long; bract at- 
tached at base of calyx, subsessile, solid, convex inside, 
flattened or slightly concave outside (resembling bract of 
R. clusiaefolia), apiculate, 3 to 4.5 mm. long, 3 to 4 m. 


246 PHY Tf Ob Oets No. 7 


wide, as much as 2 mm. thick; bracteoles 2, coriaceous, dep= 
ressed-orbicular, about 3 mm. wide, 1.6 mm. long, apex 
rounded, appressed to calyx; sepals 5, coriaceous, imbrica- 
te, suborbicular, 4 to 4.5 mm. wide, 2 to 2.5 mm. long, apex 
rounded; petals 5, imbricate, coriaceous, free, (in mature 
unopened buds ovate-elliptic, about 6 mm. long, 3.9 mm. wi- 
de); stamens 5, adnate at base of corolla; filanuents (in un 
opened buds) flattened, lanceolate, about 1.2 mm. wide at 
base, narrowed above; ovary subglobose, Sribbed, 2-celled, 
the ovules numerous; stigma subsessile, capitate. 

Type in the Herbarium of the University of Michigan, CO. 
L. Lundell 6308, collected in advanced forest on the 
Valentin-Retiro road, El Cayo District, British Honduras, 
June 30, 1936. 

R. enervia is a noteworthy species related to R. clusiae- 
folia Jacq. of the West Indies and South America from which 
it differs in having inconspicuously nerved oblanceolate— 
oblong leaves, much shorter racemes, shorter pedicels, and a 
capitate stigma. 

As far as the writer has been able to determine, this is 
the first Ruyschia know from continental North America. The 
species previously reported as belonging to schia are re- 
ferable to Souroubea. 


MYRTUS MATUDAI Lundell, sp. nov. 

Frutex ramosissimus. Folia coriacea, petiolis 3--5 mm. 
longis; laminis ellipticis, 1.5--3 cm. longis, 0.6--1.3 cm. 
latis, apice acutis vel breviter acuminatis, basi acutis. 
Flores ad axillas solitarii. Pedicelli 10--15 mm. longi. 
Prophylla sub calyce inserta linearia 4--7 mm. longa. Calyx 
ca. 6 mm. longus, S=partitus; lobis lanceolatis, 3--4 m. 
longis, acuminatis. Petala orbicularia, 3.9--4.5 mm. longa. 
Ovarium 3-loculare. 

A much-branched shrub; branchlets slender, covered with 
whitish hairs; leaves coriaceous; petioles 3 to 5 mm. long; 
leaf-blades elliptic, 1.5--3 cm. long, 0.6--1.3 cm. wide, 
apex acute or short-acuminate, base acute, costa impressed 
above, prominent beneath, the lateral nerves inconspicuous, 
glabrous above, paler beneath and at first sparsely pubes— 
cent with appressed whitish hairs, punctate, revolute; 
flowers solitary, axillary; pedicels slender, 10 to 15 m. 
long, hairy; prophyllea linear, 4 to 7 mm. long, sparsely 
pubescent beneath; calyx glabrous or nearly so, about 6 m. 
long, 5-lobed, the lobes lanceolate, 4 to 4 mm. long, acum 
inate; petals orbicular, 3.9 to 4.5 mm. long; ovary 3- 
celled; styles slender, glabrous, about 5 mm. long. 


Type in the Herbarium of the University of Michigan, Eizi 


Matuda 459, collected on Mt. Pasitar, Chiapas, iiexico, Dec- 
ember 29, 1936. 


: 
i 
3 
b, 
% 
‘ 
7 


ee Tee ee 6 ee B® 


1937 Lundell, Mexican & Central American Plants 247 


Additional specimens examined: MEXICO: Chiapas, Nt. Oran- 
do, Matuda S-3. 

M. Miatudai may be distinguished from M. montana Benth. 
and related species by its long linear prophylla and long 
calyx-lobes. 


JACQUINIA CONZATTII Lundell, sp. nov. 

Arbor. Folia subsessilia, coriacea, rugosa, oblanceolata, 
3--5.8 cm. longa, 6--14 mm. lata, apice acutiuscula vel rot- 
undata in aciculum 3 mm. longum desinentia, basi longe att- 
enuata, decurrentia, revoluta. Inflorescentiae terminales, 
abbreviate racemosae, foliis breviores; pedicellis 6--8 mn. 
longis. Sepala glabra, suborbicularia, 3--4 mm. longa, minu- 
te erosa. Corolla ca. 9.5 mm. longa, ad mediam partem gamo- 
petala, lobis oblongo-obovatis, ca. 3.8 mm. latis, apice ro- 
tundatis; staminodiis ovatis, ca. 3.1 mm. longis, 2.1 mn. 
latis. Stamina libera, filamentis intus ad corollae basin 
applicatis. 

A tree; branchlets striate, green, loosely pubescent with 
branched hairs; leaves coriaceous, oblanceolate, 3 to 5.8 
em. long, 6 to 14 mm. wide, apex acutish or rounded, spine- 
tipped, the spine about 3 mm. long, base long-attenuate, de- 
current into a short stipe, revolute, rugose, glabrous above 
except for a few hairs along the costa, pubescent beneath 
with scattered branched hairs and punctate, costa impressed 
above, prominent beneath, lateral nerves inconspicuous; in- 
florescence terminal, short-racemose, shorter than the 
leaves, S- to 12-flowered, loosely pubescent; pedicels 6 to 
8 mm. long, each subtended at base by a lanceolate, acumina- 
te-cuspidate bract 2 to 3 mm. long; sepals glabrous, subor- 
bicular, 3 to 4 mm. long, minutely erose; corolla infundibu- 
liform, orange-red, minutely puberulous inside at base with 
gland-tipped hairs, about 9.5 mm. long, lobed to the middle, 
the lobes oblong-obovate, about 3.8 mm. wide, rounded at a- 
pex; staminodia ovate, about 3.1 mm. long, 2.1 mm. wide, ob- 
tuse; stamens about 7 mm. long (including portion adnate to 
corolla); filaments free, adnate to base of corolla, narrow- 
ed above, flattened, minutely puberulous at base with gland- 
tipped hairs; style short, about 1 mm. long. 

Type in the Herbarium of the University of iiichigan, C. 
Conzatti 5326, collected near Dominguillo, District of Cui- 
catlan, Oaxaca, Mexico, on April 16, 1937, at an altitude of 
700 meters. The species is noteworthy for its narrow oblan- 
ceolate rugose leaves. 


(a) Contribution from the Herbarium of the University of 
Michigan. 


ADDITIONAL NOTES ON THE GENUS AEGIPHILA -- III 


Harold N. Moldenke 


The following notes constitute a continuation of those 
published in Phytologia 1: 182--208 and 222--240 (1937). 


85. AEGIPHILA MONTANA Moldenke. 

Additional citations: COLOMBIA: Huila: Rusby & Pennell 
7O1 (A--photo of type, B--photo of type, D--photo of type, 
F--photo of type, G--photo of type, K--photo of type, N-— 
photos of type & of isotype, P=--photo of type, S--photo of 
type, W-photo of type, Z-—-photos of type & of isotype). 


32a. AEGIPHILA MONTICOLA Moldenke in Fedde, Repert. 57: 210 

--211. 1934. 

Tree; branchlets and twigs slender, obtusely tetragonal, 
furfuraceous; principal internodes 1--2.5 cm. long; leaf- 
scars borne on short appressed sterigmata 2--% mm. long; 
leaves decussate-opposite; petioles slender, 12--15 m. 
long, furfuraceous; blades elliptic or elliptic-obovate, 8—— 
12 em. long, 3--6.1 cm. wide, acute at apex, entire, acute 
at base, densely punctate above, more or less furfuraceous 
(especially on the venation) beneath; midrib stoutish, very 
prominent beneath, slightly so above; secondseries slender, 
about 8 on each side, arcuate-ascending, prominulent beneath 
and arcuate-joined near the margins; vein and veinlet retic- 
ulation rather sparse; inflorescence axillary; cymes solit- 
ary, opposite, to 4 cm. long and 3.5 cm. wide, bifurcate and 
brachiate, -laxly many-flowered; peduncles slender, 1--2 cn. 
long, furfuraceous; pedicels slender, 1--3 mm. long, furfur- 
aceous; bractlets and prophylla linear, 1--4 mm. long, fur- 
furaceous; calyx obconic, about 5.7 mm. long and 4.1 m. 
wide at apex, coriaceous, densely pulverulent, sometimes 
Slightly verruculose, its margin 4~lobed, its lobes trunca- 
te, their margins distinctly scarious; corolla infundibular, 
white, its tube broedly cylindric, about 9.6 mm. long, glab- 
rous, its lobes 4, ovate-lingulate, about 4.5 mm. long and 
3.3 mm. wide, acute; stamens 4, inserted about 1.5 mm. below 
the mouth of the corolla=-tube, included; filaments filiform, 
about 1.3 mm. long; anthers oblong, about 3.6 mm. long and 
1.45 mm. wide; pistil included; style about 3.9 mm. long, 
glabrous; stigma bifid, its branches about 2.6 mm. long and 
slightly divaricate; ovary tetragonal, about 1 mm. long and 
wide, glabrous, 4-lobed, 4—celled. 

The type of this interesting species was collected by Dr. 
August Rimbach (No. 118) aang ein "on the outer slope 

2 


~~ 


: 
| 
| 


1937 Moldenke, Notes on Aegiphila 249 


of the Western Cordillera", Chimborazo, Ecuador, at an alti- 
tude of 2500 m., in November, 1932, and is deposited in the 
herbarium of the Field Museum of Natural History at Chicago. 
The collector notes that it is a middle-sized tree of for- 
ests, with white corollas, 4 broad corolla-segments, and 4 
stamens inserted in and included by the corolla-tube. He al- 
so states that the flowers are faintly scented and that the 
wood is not used by the natives of the region. In its gener- 
al habit this species reminds one greatly of some members of 
the section Amerina, especially A. grandis. The Schimpff 252 
cited below seems to bear strong resemblances to A. bogoten- 
sis, and, indeed, one flower was found in its inflorescence 
with 5 corolla-segments. It would appear that these three 
species need further study. 

ECUADOR: Pichincha: Schimpff 252 (E, N--fragment); Chim 
borazo: Rimbach 118 [Mus. Yale School of Forestry 22,820] 
(A--isotype, B--photo of type, F--type, K--photo of type, 
N--isotype, N--photos of type & of isotype, S--photo of 
type, Y--isotype, Z--photos of type & of isotype). 


7a. AEGIPHILA MORTONI Moldenke, sp. nov. 

Frutex; ramulis dense breviterque pubescentibus; foliis 
chartaceis oblongis vel ellipticis acutis integris, ad basin 
obtusis vel acutis, supra breviter pubescentibus, subtus 
densissime tomentosis barbellatisque non glanduliferis; in- 
florescentiis axillaribus cymosis capitatis dense multiflor- 
is dense albo-tomentosis perspicue bracteolatis. 

Shrub to 7 m. tall; branchlets medium-slender, gray-brown 
in color, obtusely tetragonal, densely short—pubescent 
throughout, slightly flattened at the nodes; nodes not annu- 
late; principal internodes 3--5.5 cm. long; leaves decussate 
-opposite; petioles slender or medium-stoutish, 1.5--1.8 cm. 
long, densely tomentose with whitish hairs, flattened (but 
not margined) above, convex beneath; blades chartaceous, 
very dark green above (brunnescent in drying), much lighter 
beneath, oblong or elliptic, 9.5--13 om. long, 3.3--4.4 om. 
wide, acute at apex, entire, obtuse or acute at base, not at 
all prolonged into the petiole, short-pubescent above (much 
more densely so or tomentellous along the midrib), very 
densely tomentose-pubescent beneath and densely barbellate 
along the midrib and secondaries, not glanduliferous; midrib 
slender and densely tomentellous-pubescent above with whit- 
ish hairs, rounded-prominent beneath; secondaries slender, 
8--12 per side, ascending, only very slightly arcuate, flat- 
tened above, prominulent beneath, rather obscurely arcuate- 
joined at the margins beneath; vein and veinlet reticulation 
indiscernible above, hidden in the tomentum or somewhat con- 
Spicuous beneath and flat or subprominulent; inflorescence ~ 
axillary, cymose; cymes solitary, opposite, capitate, 1l--2 


250 PHY -¥ 0:50 Geek No. 7 


cm. long and wide, densely white-tomentose throughout, den- 
sely many-flowered, shorter than the subtending petioles, 
conspicuously bractesolate; bractlets and prophylla linear, 
elongate, 8--10 mm. long, very densely short-pubescent with 
fulvous hairs, usually 3 clustered at the very base of the 
calyx; calyx obconic, 6--7 mm. long, 3.5--5 mm. wide, very 
densely short-pubescent with fulvous hairs, 5-ribbed, its 
rim 5-toothed, the teeth broadly triangular, about 1 mm. 
long and 2 mm. wide at base, acute; corolla-tube (in bud) 
broadly cylindric, about 4 mm. long, glabrous; corolla-limb 
5-parted, its lobes (in bud) regular, obovate, 2--3 mm. 
long, about 2 m. wide, rounded or subcucullate at apex, 
zlabrous; stamens (in bud) 5, inserted about half way up the 
corolla-tube, included; filaments flattened, about 1.5 mm. 
long, glabrous; anthers oblong, elongate, about 2.5 mm. 
long, 1 mm. wide, dorsifixed near the base; pistil (in bud) 
included; style capillary, 2--4 mm. long, glabrous; stigma 
bifid, its branches erect, about 2 mm. long; ovary subglob— 
ose, about 1.5 mn. long and 1 mm. wide, umbilicate at apex, 
glabrous; fruit in large clusters, green (according to Cook 
& Gilbert). 

The type of this most distinct species was collected by 
Orator Fuller Cook and Grover Bruce Gilbert (No. 1234) at 
Pifiasniocj, Panticalla Pass, altitude 3600 m., Cuzco, Peru, 
June 18, 1915, and is deposited in the United States Natio- 
nal Herbarium. The collectors describe the species as a 
"shrub 15--20 ft. high, with large clusters of green ber-= 
ries". It is named in honor of my esteemed friend and colle- 
ague, Dr. Conrad Vernon Norton, assistant at the United 
States National Herbarium and a recognized authority on the 
Solanaceae and Acanthaceae. 

Saat Cuzco: Cook & Gilbert 1234 (N--fragment of type, 
We-type). 


11. AEGIPHILA MULTIFLORA Ruiz & Pav. 

An additional synonym is Clerodendron bolivianum Britton 
ex Rusby, Bull. Torrey Club 27: 82 (1900). This binomial is 
often inaccurately accredited to Rusby. The type of Clero- 
dendron bolivianum was collected by Henry Hurd Rusby (No. 

2619) a at Unduavi, El Beni, Bolivia, and is deposited in the 
herbarium of Columbia University. It is identical with the 
type collection of the Rufz & Pavon species. Although this 
species superficially greatly resembles a Clerodendrum, its 
floral characters show that it actually is an Aegiphila. 
Steinbach reports that the flowers are red or rose in col- 
or, that it is a bush 2--3 m. tall, and that it ascends to 
an altitude of 3300 m. The second type locality name is 
spelled "Pill&o" on the type sheet in the British ijuseum. 
It was misspelled "Pellao" by me on page 475 of my original 


es 


1937 Moldenke, Notes on Aegiphila 251 


monograph. 

Additional citations: PERU: Huanuco?: Rufz & Pavon s.n. 
(Huassachuass & Pallao] (A--photo of isotype, B--photo of 
isotype, Bm--type & 2 isotypes, Cb--isotype, D--photo of is- 
otype, F--photo of isotype, G--photo of isotype, K--photo of 
isotype, N--fragment of isotype, N--2 photos of isotypes, 
P--isotype, S--photo of isotype, W--photo of isotype, X-- 
photo of isotype, Z-~3 photos of isotypes), s.n. (Bm); Puffo: 
Weddell 4626 (N--photo, Z--photo); Department undetermined: 


Dombey 251 (Le), s.n. [P&son Huare-huari] (P); Rufz 187 (B). 


BOLIVIA: El Beni: Buchtien 2992 (G, Le, N, f-2), an. (XI. 
1910] (B); Julio 454 (W); H. H. Rusby 2619 (C, D, G, W); 
Cochabamba: Steinbach 5809 (B, N--photo, Z—photo). 


125. AEGIPHILA NERVOSA Urb. 

An additional synonym is Aegiphila reticulata L. C. Rich., 
in herb. [not A. reticulata Moldenke, 1933]. The following 
notes made by Dr. A. ©. Smith concerning the specimens of 
this species in Urban's herbarium in February, 1932, are of 
interest: "The Haiti specimens are certainly conspecific. 
Concerning their identity with the fragment from Jamaica I 
am less certain. The young leaf of the fragment is similar 
to those of the Haiti specimens. The remaining calyx of the 
fragment is a trifle less pilose than that of the others; 
however, I conclude that all the above specimens are one 
species." 

Additional citations: JAMAICA: Collector undesignated s. 
n. ["Bois de Calme"] (Cb); Swartz s.n. (A--photo of type, B 
--fragment of type, B--photo of type, D--photo of type, F-- 
photo of type, G--photo of type, K=--photo of type, N--photo 
of type, P--photo of type, S--photo of type, W-—photo of 
type, Z~-photos of type & of fragment of type). HISPANIOLA: 
Haiti: W. Buch 1239 (B), 1949 (B, N--photo, Z--photo); 
Christ 1946 (B), 1992 (B), 2229 (B); Ekman H.472 (A--photo, 
B, B--photo, D—-photo, F--photo, G--photo, K--photo, N-=- 
photo, P--photo, W—photo, Z--photo), H.1518 (B, N--photo, 
Z--2 photos), H.8971 (B, N--photo, Z--photo); Herb. Lamarck 
261 (P); Picarda 171 (B), 1418 (B--2); L. C. Richard s.n. 


fe.ap.) (Py: 


51. AEGIPHILA NOVOFRIBURGENSIS Moldenke. 
Additional citations: BRAZIL: Rio de Janeiro: Clausen 
134 (Cb--isotype, N--fragment of isotype). 


49, AEGIPHILA NOVOGRANATENSIS Moldenke. 

This species has occasionally been confused with Cordia 
8p. and herbarium specimens have been distributed under 
that name. The Lehmann 8524 cited by me on page 349 of ny 
monograph is actually the type collection of A. aculeifera 


252 PHY TO0LjO6 14 No. 7 


and should therefore be deleted. : 

Additional citations: COLOMBIA: Antioquia: Lehmann 4066 
(A--photo of type, B-—2 isotypes, B--photo of type, Bn-- 
isotype, D--photo of type, F--isotype, F--photo of type, G-- 
photo of type, K--isotype, N--photos of type & of isotype, 
P-=-photo of type, S--photo of type, W--photo of type, X-— 
isotype, Z—photos of type & of isotype). 


119. AEGIPHILA OBDUCTA Vell. 

This is said to be a rather large tree at times, with 
creamy-white and very fragrant flowers. The very young 
leaves are often very densely villose-lanuginous above as 
well as beneath, this villosity gradually wearing off as the 
leaves mature. 

Additional citations: pee Amazonas: Ule s.n. (B); 
Minas Geraes: Bunbury 2 K); Clausen 8 (Cb, P), 652 (Us), 
s.n. [Aug.-—April, gene K); Damazio 1786 (X-=2); 
Riedel 452 (L--3); Saint-Hilaire C8774 (P--3); Schwacke 
9556 (cb--3); Sellow 1256 (N--photo, Z--photo), 5930 (B—7, 
Z—-2 photos), 8.n. (B, Z--photo); Rio de Janeiro: Bowie & 
Cunningham s.n. (Bm, N--photo, Z--photo); Brade 10,542 
[Herb. Rio de Janeiro 22,947] (B); Burchell 2018 (G, kK); 
Casaretto 926 (Cb); G. Gardner 5125 (A--photo, B--photo, Bm, 
D--photo, F--photo, G-=-photo, K, N--photo, W--photo, Z— 
photo), 5830 (Bm, K, N--photo, Z--photo); Glaziou 1498 (Br-- 
3, Cp, K, P), 3068 (Br—2, Cp, P), 4160 (Cp--2, P, Z--2 
photos), 4161 a P), 6652 (Cp, K, W-photo, P, Z--2 pho- 
tos), 11,338 (Cp, K, P, Z--photo), 11,539 (Cb, Cp, K, P, Z-- 
photo), 18 (B, Bm, Cp, K, P, Z——-photo), 19,722 (Cp, K, 
P, Z--photo); Guillemin 975 (Cb, Dc, P--2); Houllet s.n. 
(Br); Miers s.n. [Organ Mount.] (Bm); Riedel 0,33 (L--2)3 
Schott 4950 (V--2, Z--photo); Wawra 342 (V); S& Paulo: 
Bowie & Cunningham s.n. [Road to Sta. Amaro, near Sta. 
Paulo] (Bm); Burchell 4015 (K); Gaudichaud 265 (P); Puiggari 

200 (P), 3225 (P); Schwacke 1939 (Cb); Parana: Dusén 379a 
E--2, Z--photo), 8143 (A--photo, B, B--photo, Cb, D--photo, 
E, F--photo, G, G--photo, K, N--photo, S, W, w--photo, Z2-- 
photo); Jénsson 379a (B, Ca, Cb, D, N--photo, S, W, Z-- 
photo); Santa Catharina: Bunbury 463 (K); Fox 463 (Bm); 
Schwacke 13,098 (Cb); Ule 785 (Z--photo); State undetermin- 
ed: Collector undesignated 23 (Z—photo); Sellow s.n. (B). 
CULTIVATED: Brazil: So Paulo: Handro s.n. [iierb. Inst. 
Biol. S. Paulo 33,523] (N); F. C. Hoehne s.n. (Herb. Inst. 
Biol. 8. Paulo 33,523] (K), pl. viv. 442 [Herb. Inst. Biol. 
S. Paulo 33,523] (K, N, Sp). ILLUSTRATION: Line-drawing (N). 


77. AEGIPHILA OBOVATA Andr. 
Synonyms are Aegiphila dubia Moldenke in Fedde, Repert. 
33: 120 (1933) and Aegiphila sylvatica Greg, in herb. [not 


: 
‘ 
/ 


1937 Moldenke, Notes on Aegiphila om 


A. sylvatica hMoldenke, 1935]. At the time when my monograph 
was sent to the printer, I had not yet been able to secure 
the loan of the type collection of Andrews' A. obovate, and 
g0, on page 391 of my monograph, I placed Andrews' name in 
synonymy under A. dubia with a question. From Andrews’ ill- 
ustration and description I had surmised that his species 
and mine were the same, but hesitated to resurrect his name 
without having seen the actual type specimen. Since this 
time Andrews' type has been examined by myself and found to 
be definitely conspecific with the A. dubia of pages 591-- 
394 of my monograph. Andrews' name, being by far the older, 
must therefore be used for this species. The name A. = Os 
ica Greg is supposed to have been published in Sloane's 
Hist. Jamaica, but as yet I have not succeeded in locating 
the reference. The species is said to inhabit rocky vales 
and roadsides. It is described as a shrub or small tree, 
with greenish-white or -yellow corollas. The common name of 
"timber fiddlewood" has been recorded. 


Additional citations: TOBAGO: oe E.. ee 2975 (Herb. 
K-=2 > 


Monac. 4305] (By E, Le, Mu), 9064 1 (ae reg s.n. (Bm); 
Seitz 61 (B), 2 (B); R. O. Williams s.n. ee Bot. Gard. 


Herb. 11,115] (N--photo, Zi -photo). TRINIDAD: W. E. Broadwa: 
2584 (B), 3157 (B, Le), 6349 aah Es K), 6396 Cae E), 6406 
(Bn, E, K, N--photo, Z--photo), 8 (Bm); Finlay s.n. [Trin 
Bot. Gaed:, Herb. 2386] (Z-=-2 Woe os), s.n. [Trin. Bot. Gard. 
Herb. 2389] (A--photo, B--photo, D--photo, F--photo, G-- 
photo, N--3 photos, P--photo, S--photo, W--photo, Z-~2 
photos); Trin. pote Gard. Herb. 2388 (Z--photo), 5668 (N-- 
photo, Z--photo); 2 Ow ~O. Williams sen. (Trin. Bot. “Gard. 
Herb. 12,026] oe B--photo, D--photo, F--photo, G-- 
photo, K, N--photo, P--photo, S=--photo, We-photo, Z--2 
photos), s.n. [Trin. Bot. Gard. Herb. 12,163] (K, Z--photo), 

a.n, (Trin. Bot. Gard. Herb. 12,180] (K, N--photo, Z-- 
photo). CULTIVATED: British Guiana: British Guiana Bot. 
Gard. s.n. [May, 1905] (K, U); England: Hort. . Boyton 8.1 BoNe 
[A. B. Lambert, 1807; Lord Seaforth, West t Indies B—— photo 
of isotype, Cb--3 isotypes, K--photo of isotype, N--fragment 
of isotype, N--photo of isotype, S--photo of isotype, Z-- 
photo of isotype). ILLUSTRATION: Copy of Andr. Bot. Rep. 9: 
pl. 578. 1809 (Ba). 


58. AEGIPHILA OBTUSA Urb. 

Additional citations: JAMAICA: N. L. Britton 3224 (B, 
N=--photo, Z--photo); W. Harris 8996 (A--isotype, A--photo of 
isotype, B--type, B--photo of to of isotype, Bm-~isotype, D--—photo 
of isotype, G--photo of isotype, K--isotype, N--2 photos of 
isotypes, P--photo of isotype, S--photo of isotype, W--photo 
of isotype, Z--2 photos of isotypes). 


254 PHYTOLOGIA No. 7 


52. AEGIPHILA ODONTOPHYLLA Donn. Sm. 

Additional synonyms are Aegiphile furia @rset., in herb., 
and Aegiphila fusca fret., in herb. The Stork 2230 cited by 
me on pages 352 and 476 of my monograph is actually A. acul- 
eifera. The character of the teeth on the margins of the 
leaf-blades serves to distinguish the two species at once. 

Additional citations: COSTA RICA: Guanacaste: grsted 
11,174 [Macbride photos 22,775] (Cp, F=-~photo, N--photo, Z-- 
photo); Heredia: H. Pittier 288 (A--photo of type, B—photo 
of type, Br, D--photo of type, F—photo of type, G--photo of 
type, K—-photo of type, N--photo of type, P--photo of type, 
S--photo of type, #-photo of type, Z--photo of type). 


110. AEGIPHILA OVA’TA Moldenke. 

Additional citations: PERU: Junfn: Killip & Smith 26,520 
(A=-photo of type, Be-photo of type, D--photo of type, F-- 
photo of type, G--photo of type, K=-photo of type, N--photo 
of type, P--photo of type, S=--photo of type, W--photo of 
a Z~-photo of type). BOLIVIA: La Paz: Ule 9718, in part 

B). 


69. AEGIPHILA PANAMENSIS Moldenke. 

An additional synonym is Aegiphila nigrescens frst., in 
herb. Paul records the vernacular neme "hombre grande" from 
Panama, and describes the plant as a shrub 2--3 m. tall, 
blooming in November and December at an altitude of 1050-- 
1075 m. Skutch collected it in a forest at an altitude of 
850 m., blooming in July, and describes it as arborescent, 
to 6 m. tall, with yellowish flowers. It has been thought by 
some herbarium workers to belong in the Rubiaceae, and herb-= 
arium specimens have been thus distributed! 

Additional citations: COSTA RICA: Alajuela: Brenes 3574 
[110] (F), 4320 [105] (F), 4345b [105a] (F), 4395 [180] (F), 
14,315 (F); San José: Skutch 2680 (N); Cartago: @rsted 
11,175 (Cp), 11,178 (Cp). PANAMA: Bocas del Toro: Cooper & 
Slater 117 (N--photo, Z--2 photos); Dunlap 407 (Z--photo), 
438 (Z--photo); Stork 20 (Mi); Canal Zone: Killip 12,171 (N 
--photo, Z--photo); Piper 5616 (N--photo, Z--photo); Colon: 
H. Pittier 4149 (G, Z--photo); Panamdé: Heriberto 163 (Z-- 
photo); Faul 153 (W); H. Pittier 6815 (A--photo of type, B- 
photo of type, D--photo of type, F--photo of type, G--photo 
of type, N--photo of type, P=-photo of type, W--photo of 
type, Z--photo of type); P. C. Standley 28,155 (Z--photo); 
Province undetermined: Duchassaing s.n. fPenama] (P--5). 


74. AEGIPHILA PANICULATA Moldenke. 

Additional citations: NICARAGUA: Segovia?: Englesing 100 
(N--photo, W, Z--photo). PANAMA: Canal Zone: P. C. Standle 
29,232 (Z--photo), 30,246 (N--photo, Z--photo); Panamé: P. 


1937 Moldenke, Notes on Aegiphila 255 


C. Standley 28,871 (N--photo, Z--photo). COLOMBIA: Santander 
Sur: Killip & Smith 14,729 (A--photo of type, B-—photo of 
type, D--photo of type, F--photo of type, G--photo of type, 
K--photo of type, N--photos of type & of isotype, P--photo 
of type, S--photo of type, #-photo of type, Z--photos of 
type & of isotype). 


34. AEGIPHILA PARAGUARIENSIS Brigq. 

The Burchell 8345 cited by me on pages 359 and 472 of my 
monograph as A. integrifolia has proved on further study to 
be A. paragueriensis. 

Additional citations: BRAZIL: Minas Geraes: Lindley 222/ 
222b (Br); Martius s.n. [Herb. Monac. 1031] (hu); Stephan 
s.n. [1843] (Br); Warming s.n. [Lagoa Santa] (Bm, Cp--2, Z-- 
photo); Goyaz: Burchell 8345 (G, K); G. Gardner 3401 (Br, 
K); Mattogrosso: Malme 2473a (B, Cb, G, Us); Rio de Janeiro: 
Glaziou 11,334 (Cp, K, P, Z--photo); S@ Paulo: Léfgren 245 
(B); Mendonga 727 (B); Parand: Dusén 10,472 (B, Oa, E, G, 
N--photo, W, Z--photo), 15,963 (B, Cb, D, E, K, N--photo, W, 
Z--2 photos); Léfgren 95 (Z--photo), AR he State 
undetermined: Collector undesignated 404 (B); Herb. Martius 
379 (Br); Sellow s.n. (B). PARAGUAY: Fiebrig 5631 (B); Hass- 
ler 1921 (Z--photo of cotype), 4498 (B--photo of cotype, Bm- 
-cotype, Cb--cotype, D--photo of cotype, F--cotype, F--photo 
of cotype, G--photo of cotype, K--photo of cotype, N--frag- 
ment of cotype, N--3 photos of cotypes, P--cotype, V--photo 
of cotype, #-photo of cotype, X--cotype, Z--4 photos of co- 
types), 5056 (Bm--cotype, Cb--2 cotypes, F--fragment of co- 
type, N--3 photos of cotypes, P--cotype, X--cotype, Z--3 
photos of cotypes), 6766 (Bm--cotype), 6931 (Bm -cotype, 
Cb--cotype, K--cotype, N--fragment of cotype, N--photo of 
cotype, P--cotype, X--cotype, Z--2 photos of cotypes), 
10,652 (B). 


36. AEGIPHILA PARVIFLORA Moldenke. 

Additional citations: VENEZUELA: Bolfvar: Grosourdy Cat. 
13 s.n. (Z--photo). BRAZIL: Pard: Spruce 342 [Herb. Monac. 
1033] (Mu), 589 (A--photos of type & of isotype, B--photos 
of type & of isotype, Bm--isotype, Cb--isotype, D--photos of 
type & of isotype, F--photos of type & of isotype, G--photos 
of type & of isotype, N--photos of type & of 4 isotypes, W-- 
photos of type & of isotype, Z--photos of type & of 4 iso- 


types). 


25. AEGIPHILA PAUCIFLORA Standl. 

Dr. J. S. Karling of Columbia University states that the 
word "guamil" which occurs on page 314 of my monograph is a 
misspelling of "huamil". 

Additional citations: BRITISii HONDURAS: D. Stevenson 5 


256 re yY tO 222 No. 7 


(A--photo of type, B--photo of type, D--photo of type, i- 
photo of type, K--photo of type, N--photo of type, P—photo 
of type, S--photo of type, *#-photo of type, Z--photo of 


type). 


122. AEGIPHILA PAVONIANA Moldenke. 

The Conserv. Bot. Geneva specimen cited below is labelled 
"Nexique -- Pavon", but Urban claims that specimens thus 
labelled were probably collected by Sessé and liocifio in lex- 
ico and sent to Pavon by them. The species was described by 
myself from a specimen labelled "Peruvia -- Herb. Pavon" and 
was said by me to have been collected somewhere in reru by 
Rufz and ravon (vid., Brittonia 1: 190, 460, snd 461). This 
assertion was made on the basis of the specimen in the uerb- 
ier Boissier, which was the only known specimen at the time 
of that writing and which is the type specimen of the spec- 
ies. The label on the Conservatoire Botanique specimen is 
obviously older and contains longhand inscriptions. It would 
seem, therefore, to be the more authentic. The label of the 
Herbier Boissier specimen seems to be one of a uniform form 
label printed in “eneva for Pavon's herbarium when this was 
received, on the assumption (apparently erroneous) that all 
the specimens in this herbarium came from Peru. The specimen 
in the herbarium of the British Museum (which also claims to 
possess Pavon's original herbarium!) has a label which bears 
the inscription, apparently in Pavon's own handwriting, 
"Huayaquil", It would seem to me that this last-mentioned 
specimen is probably the most authentic of all and that the 
species is actually a native of Ecuador, the type having 
been collected at Guayaquil. 

Additional citations: ECUADOR: Guayas: Herb. Pavon s.n. 
{[Huayaquil] (Bm), s.n. ["Peruvia"] (A--photo of type, B= 
photo of type, D--photo of type, F--photo of type, G--photo 
of type, K--photo of type, N--photo of type, P--photo of 
type, S--photo of type, W-photo of type, Z--photo of type); 
Pavon s.n. ["Mexique"] (Cb, N--fragment, N-—photo). 


72. AEGIPHILA PENDULA Moldenke. 

The Heinrichs specimens cited below were collected in 
fruit in July, the leaf-blades are densely puberulent with 
erect hairs on both surfaces, the fruit is described as yel- 
low, and the collector describes the plant as a rare shrub 
about 2m. tall. 

Additional citations: PANAMA: Canal Zone: P. C. Standle 
25,719 (Z--photo). VENEZUELA: Zulia: H. Pittier 10,645 (A-- 
photo of type, B--photo of type, Cb--2 isotypes, D--photo of 
type, F--photo of type, G--photo of type, K--photo of type, 
N--photos of type & of isotype, P--photo of type, S--photo 
of type, ‘l--photos of type & of isotype, Z--photos of type & 


* 
a 


EE —— = 


1937 Moldenke, Notes on Aegiphila 257 


of isotype); Tejera 157 (N--photo, 2--photo), 168 (Z--photo} 
ECUADOR: Oriente: Heinrichs 499 (Cb--2, N, N--fragment, N-- 
photo, Z--photo). 


48. AEGIPHILA PENNELLII Moldenke. 

Additional citations: COLOMBIA: Tolima: Pennell 3185 (A—— 
photo of type, B--photo of type, D--photo of type, F--photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, P--photo of type, S--photo of type, W--photo of type, 
Z=--photo of type). 


35a. AEGIPHILA PERNAMBUCENSIS Moldenke, sp. nov. 

Frutex vel arbuscula; ramulis plusminus puberulis; sar- 
mentis dense breviterque pubescentibus; petiolis brevibus 
densissime bréviterque pubescentibus; laminis chartaceis ob- 
longo-ellipticis vel obovatis abrupte acutis vel subacumina- 
tis integris, ad basin subcuneatis vel acuminatis, supra 
minute puberulis vel glabratis et nitidis, subtus dense 
furfuraceo-tomentosis; inflorescentiis axillaribus; cymis 
dense multifloris obscure brachiatis; calyce campanulato 
dense tomentello, margine 4~apiculato. 

Shrub or small tree; branchlets tetragonal, gray, medul- 
lose, lightly or densely puberulent, marked with large and 
prominent corky leaf-scars; twigs slender, tetragonal, often 
decussately flattened and ampliate at the nodes, very dense- 
ly short-pubescent with dark brown hairs; principal inter- 
nodes 1.4—-5 cm. long; leaves decussate-opposite; petioles 
rather slender, 2--12 mm. long, very densely short-pubescent 
like the twigs, flattened above; blades chartaceous, oblong- 
elliptic or obovate, dark green above (brunnescent in dry- 
ing), much lighter beneath, 5--16 cm. long, 2.5--5 cm. wide, 
abruptly acute or subacuminate at apex, entire (or on water- 
sprouts and young shoots often coarsely and irregularly 
crenate-dentate with sharp teeth), subcuneate or acuminate 
at base, minutely puberulent above, becoming glabrate and 
nitid, densely furfuraceous-tomentose with rather short cin- 
ereous hairs beneath; midrib comparatively slender, flatten 
ed and usually puberulent above, very prominent beneath; 
secondaries slender, 6--9 per side, arcuate-ascending, flat 
or very slightly impressed above, prominulent beneath, the 
terminations mostly hidden in the tomentum beneath; inflor- 
escence axillary; cymes solitary, opposite, 2--2.5 cm. long, 
1.2--2.2 cm. wide, densely many-flowered, rather obscurely 
brachiate; peduncles slender, 6--10 mm. long, densely short- 
pubescent with brown or cinereous hairs; pedicels slender, 
0.5--2 mm. long, pubescent with cinereous hairs; bractlets 
and prophylla linear, 2--5 m. long, cinereous-short-pubes- 
cent; calyx campanulate, about 3.8 mm. long and 3.1 mm. wide 
at apex, densely tomentellous with hairs about 0.2 mm. long, 


258 Bet YEO 2-0 Geis No. 7 


obscurely 4-ribbed at apex, its rim 4~-apiculate, the apicu- 
lations about 0.2 mm. long; corolla hypocrateriform, actino- 
morphic, glabrous, its tube narrow-cylindric, about 3.1 m. 
long and 0.7 mm. wide, straight, its limb 4~parted, its 
lobes oblong-lingulate, about 4.2 mm. long and 2.1 mm. wide, 
acute or blunt at apex; stamens 4, greatly exserted, insert- 
ed about 0.2 mm. below the mouth of the corolla-tube; fila- 
ments filiform, about 5.2 mm. long, glabrous; anthers ob- 
long, about 1 mm. long and O.5 mm. wide, 2=celled, dorsifix- 
ed just above the base; pistil included; style capillary, 
about 3.6 mm. long, glabrous; stigma bifid, its branches a- 
bout 1.5 mm. long, slightly divergent; ovary subglobose, a- 
bout 0.7 mm. long amd 0.6 mm. wide, glabrous, 4-lobed, 4 
celled, terminated by a flattened glandular 4-lobed disk; 
fruiting-calyx and fruit not seen. 

The type of this species was collected by Don Sento 
Pickel (No. 3042) in a thicket at Tapéra, ‘ernambuco, Braz- 
il, July 14, 1932, and is deposited in the Britton Herbar- 
ium at the New York Botanical Garden. The species was in 
cluded with A. crenata by myself in my monograph and in the 
first installment of these supplementary notes (Brittonia 
1: 327. 1934; Phytolozia 1: 196. 1937) and the Pickel col- 
lections from Pernambuco were all erroneously cited as A. 
crenata. A. crenata is a more southern species, confined to 
Minas Geraes, Sao Paulo, and Parand. One sheet of the type 
collection in the Britton Herbarium is composed of the tips 
of water-sprouts with conspicuously obovate leaf-blades, ‘ 
which are cuneate at the base and with their margins coars- 
ely and very irregularly crenate-dentate. The collector 
states that these twigs were taken from an offshoot of a 
young plant. 

BRAZIL: Pernambuco: Pickel 526 [Herb. Inst. Biol. S. 
Panlo 20,089] (B, N), 3030 (N), 3042 (Ba--isotype, I--iso- 
type, N--type, N--4 isotypes, W--isotype); Ramage s.n. [21 
Jan. 1888] (Bm); Rio de Janeiro: Burchell 9 (K); Swainson 
s.n. (G, K). 


75. AEGIPHILA PERPLEXA Moldenke. 

Additional citations: TOBAGO: W. E. Broadway 4243 (B, 
Cp); Eggers S540 (B, Le, N--photo, P, Vu, Z--2 photos). 
TRINIDAD: D. W. “q®. Alexander s.n. (Trin. Bot. Gard. Herb. 

5022] (B); Fendler 571 (Z--photo), 592 (Bm, Ed--2, N=- 
photo, Z--photo). VENEZUELA: Delta Amacuro: Rusby & Squires 
316 [Herb. Monac. 1843] (B--photo of type, Bu--isotype, Cb- 
-isotype, D--photo of type, E--isotype, Ed--isotype, G-- 
photo of type, K--isotype, Mi--isotype, Mu--isotype, N-- 
photos of type & of 2 isotypes, P--photo of type, S--iso- 
type, S--photo of type, Vu--isotype, W--photo of type, X-- 
isotype, Z--photos of type & of 2 isotypes). 


————— 


1937 Moldenke, Notes on Aegiphila 259 


64. AEGIPHILA PERUVIANA Turcz. 

Klug describes the species as a shrub about 1 m. tall, 
growing in forests, with cream-colored corollas, blooming in 
January. It seems very probable that further study will 
bring to light a sufficient number of differences between 
the Peruvian and Bolivian specimens cited for this species, 
to justify their separation (as Dr. Rusby maintains) and to 
warrant the application of a new name to the Bolivian forn, 
even if only of varietal rank. ; 

Additional citations: PERU: San Martin: Klug 3511 (Cb, E, 
I, N); A. Mathews 1617 (N--photo, Z—-photo), s.n. [Bioyobam- 
ba] (Z--photo); Spruce 4275 [Macbride photos 24,619] (A-- 
photo of isotype, B--photo of isotype, Bm--isotype, Br-- 
isotype, Cb--2 isotypes, Cp--isotype, D--photo of isotype, 
Ed--isotype, F--fragment & photo of isotype, F--photo of 
isotype, G--photo of isotype, K--isotype, N--4 photos of 
isotypes, P--isotype, P--photo of isotype, S--photo of iso- 
type, #-photo of isotype, Z--3 photos of isotypes); L. 
Williams 5383 (N--photo, Z--photo), 5479 (Z--photo), 5480 
(Z--photo), 5588 (Z--photo), 5697 (Z—-photo), 6152 (Z-= 
photo), 6291 (Z--photo), 6304 (Z--photo), 6836 (A--photo, B 
~-photo, D--photo, N--2 photos, P--photo, S=--photo, #-- 
photo, Z--photo), 6860a (Z--photo), 7334 (Z--photo); Loreto: 
Ule 6476 (K). BOLIVIA: El Beni: H. H. Rusby 2473 (Bm, E, Ed, 
K, Pr, W, Z--4 photos). 


100. AEGIPHILA PLATYPHYLLA Briq. 

Additional citations: PARAGUAY: Hassler 8056 (B--photo of 
isotype, Br--isotype, Cb--isotype, D—photo of isotype, F-- 
isotype, F--photo of isotype, G--photo of isotype, K-~-iso- 
type, N--fragment of isotype, N--43 photos of isotypes, P-- 
isotype, S=--photo of isotype, W-photo of isotype, X--iso- 
type, Z--photos of type & of 3 isotypes). 


60. AEGIPHILA PLICATA Urb. 

Additional citations: JAMAICA: Bertero 2105 (B--type, De- 
-isotype, Z--photos of type & of isotype), s.n. [Majo Jun. 
1831] (B). 


88. AEGIPHILA PUBERULENTA Moldenke. 

Additional citations: COLOMBIA: Bolfvar: Elias 617 (A-- 
photo of type, B--photo of type, D-—-photo of type, G--photo 
of type, K--photo of type, N--photos of type & of isotype, 
P--photo of type, S--photo of type, W-photo of type, Z-= 
photos of type & of isotype), 673 (N--photo, Z--photo). 


105. AEGIPHILA PULCHERRIMA Moldenke. 
Additional citations: PERU: Jun{fn: Schunke 400 (N--photo, 
Z--photo), 443 (A--photo of type, B--photo of type, D--photo 


260 PHY TOLOG2i2s No. 7 


of type, G-photo of type, K--photo of type, N--photos of 
type & of isotype, P--photo of type, S--photo of type, #- 
photo of type, Z--photos of type & of isotype). 


91. AEGIPHILA QUINDUENSIS (H.B.K.) Moldenke. 

The "Pittier 8806" cited by me on pages 416 and 475 of ny 
monograph should have been cited as "H. Pittier 8806". Some 
Sheets of this collection are A. elate and have been desig- 
nated as H. Pittier 8806a. 

Additional citations: COLOMBIA: Cundinamarca: Mutis 182 
(W); Tolima: Bonpland 5864 (P); Goudot s.n. [Quindiu] (N-- 
photo, P, Z--photo); Humboldt & Bonplan and 8 .! s.n. [El Noral, 
Quindiu] (A--photo of type, B--photo of type, D--photo of 
type, F--photo of type, G--photo of type, N--photo of type, 
P--photo of type, S--photo of type, W--photo of type, Z-- 
photo of type). VENEZUELA: Federal District: H. Pittier 
10,404 (Ba, Cb--2, P, Z--photo); Carabobo: Karsten s.n. 
[Puerto Cabello] (aL B--photo, D--photo, F=--photo, G@ 
--photo, N=-2 photos, P--photo, S--photo, W-photo, Z--2 
photos); H. Pittier 8806-(Z--photo). 


115. AEGIPHILA RACEMOSA Vell. 
The Sagot 473 cited by me on pages 449 and 476 of ny 
monograph should read "in part", since some sheets of this 


collection seem to be A. glandulifera var. pyramidata. The 
Aegiphila aequinoctialis Mart. cited on page 421 as a syno= 
nym of A. vitelliniflora is actually a synonym of A. racemo= 
sa. This has been aeabegenee through examination of the 
type specimen, Martius 2724, in the Munich herbarium. The 
"Schomburgk 281" cited on a 476 was cited in error and 
should be deleted. The "Dahlgren & Sella 542" cited on page 
473 is a typographic error for Dahlgren & & Sella 524. The 
species has been confused by some herbarium workers with A. 
foetida Sw., and some herbarium specimens have been distri- 
buted under that name! The two Goeldi collections cited be- 
low are very anomalous in their thinly membranous leaf- 
blades, but in all other respects match precisely typical 
specimens of A, racemosa. They may possibly represent a 
hybrid with A. . vitelliniflora. 
Additional citations: VENEZUELA: Zulia: Rutten-Pekelhar- 
40 (Ut). BRITISH GUIANA: Aitken 1082 (kK); Beckett. 8447 
3 U); De la Cruz 946 (A-=photo, B--photo, D--photo, F-- 
photo, N--photo, W--photo, Z--photo), 3627 (A--photo, B- 
photo, agg E, G--photo, N=--photo, W-=photo, Z--2 
Photos), 4292 (E), 4552 (E); Drake s.n. (N--2 photos, Z-- 
ne Jenmen 1993 (N--photo, U, Z-=2 photos), (K, U)s 
. Schomburgk ce eae ee 146 (B; sir (Bm, 
af 5 615 (Z--photo). SURINAM: Ber thoud-Coulon 350 a 


Collector i indig. Suriname 87 (Ut); Department of Forestry 


f 
J 
q 
2 
{ 

y 


1937 Moldenke, Notes on Aegiphila 261 


2850 (Ut); Kappler 1717 (E, P, Ut, V, Z+-photo). FRENCH 
GUIANA: Collector undesignated s.n. (P); Mélinon 174 (2-- 
photo), hak (P, a Os Sagot 473, in part (Cb--3, K, P-- 
4, V, Z--2 photos), s.n. [Karovany, 1854~-58] (P). BRAZIL: 
Amazonas: Martius 0,39 [Herb. Monac. 1012] (Mu), s.n. [R. 
Negro; Herb. Monac. 1011] (Mu); Pardé: Burchell 9726 (K, N-- 
photo, Z--photo); Dahlgren & Sella 524 (N--photo, Z--photo); 
Goeldi 6999 (Cb), 7002 (Cb); Killip & Smith 30,278 (Z—- 
photo); Martius 2724 [Herb. Monac. 1008] (Mu); Snethlege 109 
(B, F); Pernambuco: Pickel 3642 (W); Bahia: Martius s.n. 
[Herb. Monac. 1000 & 1009] (Mu--2); State undetermined: 
Blanchet s.n. (Bu--2). LOCALITY OF COLLECTION UNDESIGNATED: 
Herb. A. L. Jussieu 5039 (P). ILLUSTRATION: Line-drawing (N). 


89. AEGIPHILA RETICULATA Moldenke. 

Additional citations: COLOMBIA: Cundinamarca: Mutis 985 
[857] (F--photo of type, G--photo of type, K--photo of type, 
N--photo of type, S--photo of type, Z--photo of type). 


24. AEGIPHILA RIEDELIANA Schau. 

Schwacke says that the branches of this species are pen- 
dent, the corolla yellow, the fruit green, and that it 
blooms in November, a vernacular name being "cajuja". 

Additional citations: BRAZIL: Bahia: Pohl 4392 (A--photo 
of cotype, B--photo of cotype, D--photo of cotype, F-—-photo 
of cotype, G--photo of cotype, N--photo of cotype, S--photo 
of cotype, W--photo of cotype, Z--photo of cotype); Minas 
Geraes: Mosén 2005 (S); Rio de Janeiro: Burchell 1985 (K); 
Miers 4500 (N-—photo, Z--photo); S&o Paulo: Miers s.n. [Ar- 
raras, Jan. 7, 1838] (Bm); Santa Catharina: Schwacke 12,966 
(Cb, N--fragment); Ule 1537 (A--photo, B--photo, D--photo, 
F--photo, G--photo, N--photo, S--photo, W--photo, Z=--photo), 
1863 (N--photo, Z--photo); Rio Grande do Sul: Malme 898 (S-- 
2), 951 (S); Tweedie s.n. [Porto Alegre] (N--photo, Z-- 
photo). 


14a. AEGIPHILA RIMBACHII Moldenke, sp. nov. 

Frutex vel arbor; ramulis plusminus furfuraceo-puberulis 
glabrescentibus; petiolis crassis furfuraceis submarginatis; 
laminis coriaceis nitidis oblongo-ellipticis vel ellipticis 
vel subobovatis subacutis vel obtusis integris subrevolutis, 
ad basin acutis vel cuneatis, utrinque zlabris (praeter cos- 
ta puberula vel cinereo-pulverulenta), subtus punctulatis; 
inflorescentiis axillaribus glomeratis multifloris; calyce 
campanulato brevissime tomentello, margine truncato integro. 

Shrub or tree; branchlets more or less acutely or obtuse- 
ly tetragonal, stramineous or brownish, sparsely lenticell- 
ate, more or less furfuraceous—puberulent, becoming subglab= 
rate; principal internodes apparently variable and irregu- 


262 PRY TOLOo Gta No. 7 


lear, 0.7--6 cm. long, often a long one followed immediately 
by a very short one and then a very long one again, so that 
the leaves may appear to be whorled; leaves decussate- 
opposite, often approximate; petioles stout, 1.3--2 cm. 
long, flat above, rounded beneath, furfuraceous, slightly 
margined; blades coriaceous, bright green above, much darker 
and blackening beneath only in drying, nitid on both surf- 
aces, oblong-elliptic, elliptic, or tending toward being 
slightly obovate, 10--14 cm. long, 5.5--5.5 om. wide, blunt- 
ly acute or rounded at apex, entire and slightly revolute a- 
long the margins, acute or cuneate at base, glabrous above 
or slightly puberulent along the midrib, glabrous beneath 
except for the cinereous-pulverulent or puberulent midrib, 
somewhat punctate beneath; midrib stout, more or less imp- 
ressed or else prominulent in a groove above, very prominent 
beneath; secondaries slender, 6 or 7 per side, arcuate- 
ascending and strongly joined almost at the margins beneath, 
often subimpressed above, very sharply prominent beneath; 
vein and veinlet reticulation very sparse and obscure on 
both surfaces or a few veins sharply prominent near the mar- 
gins beneath and connecting the secondaries; inflorescence 
axillary, glomerate, the fascicles many-flowered and dense; 
calyx campanulate, about 5.4 mm. long and 5.7 mm. wide at a- 
pex, very short-tomentellous, its rim truncate and entire; 
corolla hypocrateriform, actinomorphic, glabrous outside, 
its tube broadly infundibular, about 5.4 mm. long, ampliate 
to 5.5 mm. at apex, its limb 4~parted, regular, its lobes 
ovate-lingulate, about 3.1 mm. long and wide, rounded at a- 
pex; stamens 4, included, inserted about 1.7 mm. below the 
mouth of the corolla-tube; filaments filiform, about 1.3 mm. 
long, glabrous; anthers oblong, about 3.1 mm. long and 1 mm. 
wide, 2-celled, dorsifixed just above the base, the thecae 
parallel; pistil included; style heavy, about 3.1 mm. long, 
glabrous; stigma bifid, its branches about 1.6 mm. long, di- 
vergent; ovary subglobose, about 1 mm. long and wide, glab- 
rous, 4-lobed, 4-sulcate, 4-celled, 4-ovulate; fruiting- 
calyx and fruit not seen. 

The type of this most distinctive species was collected 
by August Rimbach (No. 234) on the outer slope of the west- 
ern Cordillera ebove Balsapampa, at an altitude of 2600 m., 
Los Rfos, Ecuador, in October, 1934, and is deposited in the 
herbarium of the Field Museum of Netural History at Chicago. 
It is named in honor of its distinguished collector, who has 
made such noteworthy collections in a little-known region of 
Ecuador. “e describes the species as follows: "Middle-sized 
forest tree. Leaves opposite, thickly stiff, glabrous, shin- 
ing. Flowers in axillary glomerations. Flower 15 mn. long. 
K. gamopetalous with brownish pubescence. C. with greenish 
tube and 4 reddish lobes. A. 4, fixed in the corolla-tube. 


1937 Moldenke, Notes on Aegiphila 263 


P. superior, with globular ovary and thin style. Fruit not 
scen. ‘ood YePtoween Cortex 5 mm. thick. Bark gray, rough, 
with round warts," 

ECUADOR: Los Rios: Rimbach 234 [Mus. Yale School of For- 
estry 28,648] (N--isotype, N--fragment of type, N--photo of 
isotype, W-isotype, Y--isotype, Z--photo of isotype). 


118. AEGIPHILA RORAIMENSIS Moldenke. 

Additional citations: BRITISH GUIANA: Appun 1204 (A-- 
photo of type, Be-photo of type, D--photo of type, F-~photo 
of type, G--photo of type, N--photo of type, P--photo of 
type, S--photo of type, iW--photo of type, Z--photo of type). 


4ha, AEGIPHILA SALTICOLA Moldenke in Fedde, Repert. 37: 21l- 

~212. 1934. 

Spreading shrub, to 3 m. tall; branchlets stoutish, tet- 
ragonal, somewhat flattened and ampliate at the nodes, very 
densely and shortly appressed-pubescent; principal inter- 
nodes 4--5.5 cm. long; leaves decussate-opposite; petioles 
stout, 1--3.5 cm. long, densely appressed-pubescent with ex- 
tremely short blackish hairs; blades membranous, oblong- 
elliptic, 12--19 cm. long, S--7 cm. wide, acuminate at apex, 
entire, long-cuneate at base, roughened with abundant minute 
elevated punctae above, densely pubescent or subvillose be- 
neath; midrib very stout, prominent beneath; secondaries 
slender, 9--12 on each side, ascending, not very arcuate, 
prominulent and distinctly arcuate-joined near the margins 
beneath; vein and veinlet reticulation slender, mostly hid- 
den beneath and obscure above; inflorescence supra~axillary; 
cymes solitary, opposite, about 4 cm. long and 3 cm. wide, 
very densely many-flowered, often subcapitate in fruit; ped- 
uncles slender, about 2 cm. long, densely short-appressed- 
pubescent; branches of the cymes and the pedicels slender, 
densely subvillose, the fruiting-pedicels to 6 mm. long; 
flowers not seen; fruiting-calyx cupulirorm, 3--5 mm. long, 
6-~9 mm. wide, densely subvillose with whitish hairs, its 
margin shallowly 4-lobed with broadly triangular lobes; 
fruit drupaceous, oblong, about 7 mm. long and 6 m. wide, 
yellow when mature, smooth, hard, not at all fleshy, 4- 
seeded. 

The type of this neglected species was collected by Ynes 
Mexia (No. 5922) along the edge of cut-over woods in the 
Japanese @ colony at Thomé Assu, district of Acard, altitude 
about 30 m., Pardé, Brazil, July 16, 1931, and is deposited 
in the herbarium of the Field Museum of Natural History at 
Chicago. It is very closely releted to A. integrifolia, 
from which its pubescence, however, at once distinguishes 
» 

BRAZIL: Par&: Mexia 5922 (B--photo of type, F--type, K-- 


264 PHY T Ob 6G 2-8 No. 7 


photo of type, N-~-fragment of type, N--photo of type, S-- 
photo of type, Z--photo of type). 


63a. AEGIPHILA SCANDENS Moldenke in Fedde, Repert. 37: 212. 

1934. 

Scandent shrub; branches rather slender, obtusely tetrag- 
onal, buff, rather densely furfuraceous—puberulent; princip- 
al internodes elongate, 4.5--7.5 cm. long; leaves decussate- 
opposite; petioles stoutish, 7--10 mm. long, densely puberu- 
lent; blades firmly membranous or subcoriaceous, dark green 
and nitid, slightly lighter beneath, ovate or ovate-elliptic, 
14~-19 cm. long, 5--8.5 cm. wide, acute at apex, entire, 
rounded at base, glabrous on both surfaces (except for the 
more or less puberulent midrib beneath), densely impressed- 
punctate beneath; midrib rather slender, sharply prominu- 
lent in a channel above, very prominent beneath; secondaries 
slender, 8--10 per side, arcuate-ascending; inflorescence 
axillary and terminal; axillary cymes very few, apparently 
in the uppermost axils only, to 5 cm. long and 4.5 cm. wide, 
many-flowered, bracteolate; panicle terminal, about 10.5 cm. 
long and 6.5 cm. wide, composed of about 9 cymes, its sym- 
podia short and densely yellowish-puberulent; peduncles 
rather slender, 1.5--45 cm. long, yellowish-puberulent; 
bractlets various, oblong, obovate, lanceolate, or elliptic, 
3--13 mm. long, 1--3 mm. wide, rather long-stipitate, spar- 
sely puberulent; pedicels very slender, 1--5 mm. long, 
densely yellowish=puberulent; calyx campanulate, about 3.1 
mm. long and 3.6 mm. wide at apex, minutely puberulent, its 
margin subtruncate, slightly scarious and repand; corolla 
hypocrateriform, yellow, its tube cylindric, about 9.5 mm. 
long, ampliate above, its lobes 4, lanceolate-oblong, 4--6.5 
mm. long and 1--1.8 mm. wide, subacute at apex; stamens 4, 
inserted about 4.9 mm. below the mouth of the corolla-tube, 
exserted; filements filiform, about 9 mm. long, glabrate; 
anthers and pistil not seen; ovary obovate, tetragonal, a- 
bout 1 mm. long and wide, glabrous, 4-celled; fruiting-calyx 
and fruit not seen. ; 

The type of this handsome species was collected by Ernst. 
Heinrich Georg Ule (No. 9721) at Seringal Auristella on the 
Rio Acre, Acre Territory, Brazil, in March, 1911, and is 
deposited in the herbarium of the Royal Botanic Gardens at 
Kew. The label on the type specimen bears the written anno- 
tation "Peru", but according to that indefatigable collect 
or, Mr. B. A. Krukoff, who has done perhaps more than any 
one person since the days of the pioneer explorers and col- 
lectors in Brazil to advance our knowledge of the flora of 
that country, the Seringal Auristella is definitely in Acre 
Territory, Brazil. 

BRAZIL: Acre Territory: Ule 9721 (B--isotype, K--type, 


— 


1937 Moldenke, Notes on Aegiphila 265 
N--fragment of type, N--photo of type, Z--photo of type). 


108a. AEGIPHILA SCHIMPFFII Moldenke, sp. nov. 

Frutex; ramulis dense puberulis; petiolis dense puberul- 
is; laminis chartaceis fragilibus ellipticis longe acuminat- 
is integris, ad basin acutis, supra sparsiuscule puberulis, 
subtus dense puberulis; inflorescentiis axillaribus termin- 
alibusque; cymis axillaribus subpaucis, ad apicem ramulorum 
congestis; paniculis terminalibus thyrsoideis ubique dense 
puberulis. 

Shrub, to 5m. tall; branchlets slender, obtusely tetrag- 
onal, brownish, densely puberulent, rather sparsely lentic- 
ellate; nodes not annulate; principal internodes 3-7 cm. 
long; leaf-scars large and prominent, borne on elongate 
stout and divergent sterigmata; leaves decussate-opposite; 
petioles slender, 7--8 mm. long, densely puberulent with 
brownish hairs like the branchlets, slightly flattened a- 
bove, convex beneath, not noticeably ampliate at base; 
blades chartaceous, dark green above, lighter beneath, very 
fragile and undate in drying, elliptic, 8--18 cm. long, 3.5 
--6.5 cm. wide, long-acuminate at apex, entire, acute at 
base, rather sparsely puberulent above, densely so beneath, 
the puberulence brownish like that on the branchlets and 
petioles; midrib slender, slightly prominulent above, 
rounded-prominent beneath; secondaries slender, S--15 per 
side, irregular, flat above, prominulent beneath, arcuate— 
ascending, not conspicuously anastomosing; vein and veinlet 
reticulation very slender, obscure above, flat or the larg- 
er portions subprominulent beneath; inflorescence axillary 
and terminal; axillary cymes rather few, crowded toward the 
apex of the branchlets and building up an elongate pyramid- 
al thyrse with the terminal panicle, the individual cymes 
2--5.8 cm. long and 1--3 cm. wide, densely many-flowered; 
terminal panicle 8--135 cm. long, 4--4.5 cm. wide at base, 
composed of 3 or 4 pairs of cymes and a terminal one; ped- 
uncles of cymes 1--3.5 cm. long, of panicles 4--5 cm. long, 
densely puberulent throughout; sympodia densely brown- 
puberulent like the peduncles; bractlets and prophylla fil- 
iform, 1--3 mm. long, short-pubescent-pilose; calyx campan- 
ulate, 2.9--45 mm. long, about 2.5 mm. wide at apex, slight- 
ly contracted just above the ovary, pulverulent and slight- 
ly puberulent, its rim 4-toothed, the teeth triangular, a- 
bout 1 mm. long and wide at base, rounded, short-apiculate 
with a callose point at apex; corolla infundibular, its 
tube nerrow-cylindric, about 3.5 mm. long, glabrous out- 
side, tomentulose in the throat within, its limb 4—parted, 
its lobes elliptic, about 2.7 mm. long and 1 mm. wide, ob- 
tuse at apex; stamens 4, inserted at apex of the corolla- 
tube; filaments obsolete; anthers subsessile, elliptic, a- 


266 PHY oF. O450-G/ 28 Nowy 


bout ©.5 mm. long end 0.5 mm. wide, dorsifixed near the 
base; pistil long-exserted; style capillary, about 4.5 m. 
long, glabrous; stigma bifid, its branches greatly elongate, 
o--5.5 mui. long, twisted; ovary minute, tetra sonal, black, 
about 0.5 mm. long and wide, glabrous, 4sulcate, 4-lobed at 
apex, 4—celled; fruiting-cealyx incrassate, ebout 4 om. long 
and 6 mm. wide, obscurely pulverulent-puberulent or glabra- 
te, its rim subentire or shallowly erose; fruit not seen. 

The type of this recently discovered species was collect- 
ed by H. J. F. Schimpff (No. 1003) -- in whose honor it is 
named -—- in a forest near Biscay, Ecuador, et an altitude of 
300 m., April 1&, 1934, and is deposited in the herbarium of 
the Botanisches Museum at Berlin. 

ECUADOR: Province undetermined: Schimpff 1C03 (B--type & 
isotype, N--fregment of type). 


39. AEGIPHILA SELLOWIANA Cham. 

An additional synonym is Aegiphila Orbignyana Mart., in 
herb. The accepted specific name for the species is also 
sometimes misspelled "Selloviana” or "Schowiana". On page 
475 of my monograph I cited a "Riedel & Luschnath 1815" for 
this species -- this is a typographic error for Riedel & 
Luschnath 1812. The "Severén 188" cited on page 476 is an 
error and should be deleted. On page 334 I cited "Sellow s. 
n. [flowers]" as the type collection; this should be cited 
as a cotype collection, since the species was based by Cham- 
isso on two separate collections. Schwacke states that the 
flowers are odorous. Miss “exia describes the species as a 
conrmon shrub, with long straggling brenches, growing to be a 
tree, with slightly fregrant flowers and greenish-white cor- 
ollas. Saint-Hilaire reports the vernacular name "habiara", 
while "cajug&" has also been reported from Brazil. 

Additional citations: BRAZIL: Bahia: Blanchet 28 (Bm); 
Riedel 222 (L--2); Goyaz: Pohl & Schott 9536/4915 (V--2); 
Mattogrosso: Malme 2473 (S, W); Minas Geraes: Ackermann s.n. 
[1831] (Br--2); Clausen 361 (B), ue (Cp, P—-3, Us), 8.n. 
{Minas Geraes; Herb. De Candolle 869 & 876] (Cb--4, Dc-=2, 
K), s.n. (Aug.--April, 1840] (Br--2, K); Damazio 1926 (X-- 
5); Frambach Le (F, N, 8); Gaudichaud s.n. [1830] (De); 
Henschen s.n. [Herb. Regnell 1.184, 1868] (B, Us, W, Z2--3 
photos); Mexia 5396 (B, Bm, Cb, E, G, I, P, 8); Regnell I. 
184 [1845] (Cp), 1.184 (1866; Herb. Monac. 1658) (Br, Mu, 01 
--53, S—2, Us, W), 1.184 [1877] (Us); Saint-Hilaire 2229 
(P), 2229 ter (P—-2), s.n. [Itabura] (P--3); Schwacke 11,264 
(Cb); Warming s.n. [Lagoa Santa] (Bm, Cp); Espirito Santo: 
Campos Novaes 932 (Vu); Luetzelburg 12,345 (Mu); Rio de Jan- 
eiro: Burchell 2042 (K), 2492 (B--photo, D--photo, G--photo, 
S--photo, W--photo, Z--photo); Glaziou 808 (Br--2, Cp-=2, K, 
Z=-photo), 5956 (Cp, P—2), 14,164 (Bm, Br, Cb--2, Cp, K—=2, 


1937 Moldenke, Notes on Aegiphila 267 


P--2); Langsdorff s.n. [Mandiocca; Riedel] (L--2); Martius 
448 [Herb. Monac. 1042] (Mu); Miers 4592 (P), s.n. [Organ 
Mount., 1828] (Bm), s.n. [Imbuby, Organ Mt., Dec. 1837] (Bm); 
Peckolt 223 (Z--photo), 422 (Br); Pohl s.n. [Herb. Imp. 
Vien. 150] (B, K); Riedel 0,34 (L--2), s.n. [Rio de Janeiro] 
(Us); Schfich a.n. (V); Sellow s.n. [flowers] (K--2 cotypes, 
N--2 photos of cotypes, P--cotype), s.n. [fruit] (A--photo 
of cotype, B--2 cotypes, B--photo of cotype, Bm--cotype, D-- 
photo of cotype, F--photo of cotype, G--photo of cotype, K-- 
cotype, W—-photo of cotype, Z—-4 photos of cotypes); Warming 
a.n. (Cp); So Paulo: Brade 7464 (B); Burchell 3418, in part 
(K), 4604 (K); Collector undesipnated 3653 (Vu); F. C. 
Hoehne s.n. (Herb. Inst. Biol. S. Paulo 8179] (A, W); Kos- 
cinsky 225 [Herb. Ingt. Biol. S. Paulo 31,656] (K, Sp); 
Lofgren s.n. [Herv. Geogr. e Geol. 373] (Cp), s.n. [Herv. 
Geogr. e Geol. 407] (Cp); Mosén 1534 (Z--photo), 3035 (S, 
Us, Z--photo), 4323 (A--photo, B--photo, D--photo, F--photo, 
G--photo, N--photo, P, W--photo, Z--photo); Riedel & Lusch- 
nath 1812 (L--2); Widgren 1177 (Us), 11.1176 (Us), I1.1176 
1/2 (Us); Parand: naa L7he as 2532 (G), 11,228 (Z-- 
photo), 15,855 (B, Ob, Ey &, NW), 16,162 (Cb, D), 16,669 (B, 
N--photo, Z—2 photos), 17,362 (B, E, 8, W, Z--photo); Rio 
Grande do Sul: Malme 799 (B, N--photo, Us, Z--photo); State 
undetermined: Blanchet s.n. (Bm); Herb. Martius s.n. [Herb. 
Monac. 1450] (Mu); Herb. Zuccarini s.n.-[Herb. Monac. 1043] 
(Mu). BOLIVIA: La Paz: M. Bang 1332 [Herb. Monac. 1725] 

(Bm, Cb—-2, E, Ed, K, Mi, Mu, N--photo, S, Vu, Z--photo); 
Santa Cruz: D'Orbigny 1094 (P--2). CULTIVATED: Brazil: Séo 


Paulo: F. C. Hoehne, pl. viv. 192 (Herb. Inst. Biol. S. 
Paulo 28,666] (W--2). 


16. AEGIPHILA SESSILIFLORA Moldenke. 

Additional citations: COLOMBIA: Antioquia: Archer 392 
(A--photo of type, B--photo of type, D--photo of type, F-- 
photo of type, G--photo of type, N--photo of type, P-=-photo 
of type, S--photo of type, W--photo of type, Z--photo of 


type). 


104. AEGIPHILA SETIFORMIS Rusby. 

The "Bang 878a" cited by me as the type collection of 
this species on pages 434, 435, and 472 of my monograph, 
should more accurately be cited as "M. Bang 878a, in part", 
since some sheets of this number are A. spicata. 

Additional citations: BOLIVIA: La Paz: M. Bang 8/8a, in 
part (A--photo of type, B--photo of type, Bm--isotype, D— 
photo of type, E--isotype, Ed--isotype, G—photo of type, 
K--isotype, N--photos of type & of 2 isotypes, P--photo of 
type, S—photo of type, W--isotype, W--photo of type, Z— 
photos of type & of 2 isotypes), 1732 [Herb. Monac. 1727] 


268 PE t.0 ere No. 7 


~photo, S=-photo, V, Vu, W, W--photo, Z=--3 photos). 


94. AEGIPHILA SMITHII Moldenke. 

Additional citations: PERU: San Martfn: Klug 3894 (S); 
Loreto: Killip & Smith 26,957 (A--photo of type, B-—photo of 
type, D—-photo of type, F--photo of type, G--isotype, G-- 
photo of type, K--isotype, K--photo of type, N--photos of 
type & of isotype, P—-photo of type, S-=photo of type, W-= 
photo of type, Z--photos of type & of isotype); Klug 1460 (A 
--photo, B--photo, D=--photo, By Page? i SE Pe-photo, S 
--photo, W--photo, Z--2 photos), 1490 (Z--photo); Melin s.n. 
[Iquitos] (8); Mexia 6499 (N); Tessmann 3591 (Cb, Hb), 3603 
(Hb); L. Williams 3689 (S). 


14. AEGIPHILA SORDIDA Moldenke. 

Additional citations: PERU: Loreto: L. Williams 5054 (A-- 
photo of type, B--photo of type, D--photo of type, G-—photo 
of type, K--photo of type, N--photo of type, P=--photo of 
type, S--photo of type, W--photo of type, Z--photo of type). 


103. AEGIPHILA SPICATA (Rusby) Moldenke. 

Additional citations: BOLIVIA: La Paz: M. Bang 878a, in 
part (A--photo, Be-photo, D--photo, F--photo, G--photo, K-- 
photo, P--photo, W--photo, Z——photo). 


46. AEGIPHILA SPLENDENS Schau. 

Additional citations: BRAZIL: Minas Geraes: Pohl 1022 
{[Herb. Imp. Vien. 149] (A--photo of isotype, B--photo of 
isotype, D--photo of isotype, F--photo of isotype, G--photo 
of isotype, K--isotype, N--fragment of isotype, N--photos of 
type & of 4 isotypes, P--isotype, S--photo of isotype, 
photo of isotype, Z——photos of type & of 4 isotypes). 


92. AEGIPHILA SPRUCEANA Moldenke. 

The Spruce 3017 cited by me on page 417 of my monograph 
as a cotype collection of this species, should not have been 
called a cotype. The species is based solely on Spruce 2296. 

Additional citations: BRAZIL: Amazonas: Koch-Grtinberg 93 
(B); Spruce 2296 (A--photo of type, B--isotype, B--photo of 
type, Bm--isotype, Br--isotype, Cb--isotype, Cp--isotype, D- 
-photo of type, Ed--isotype, F--isotype, F--photo of type, G 
--photo of type, K--isotype, N--isotype, N--photos of type & 
of isotype, W--photos of type & of isotype, Z--photos of 
type & of isotype), 3017 (K, N--3 photos, S, Z—3 photos). 


26. AEGIPHILA STANDLEYI Moldenke. 
Additional citations: COSTA RICA: San José: P. C. Stand- 
ley 37,570 (A—photo of type, B--photo of type, D--photo of 


1937 Moldenke, Notes on Aegiphila 269 


type, F--photo of type, G-—-photo of type, K--photo of type, 
N=--photo of type, P--photo of type, S—photo of type, W-- 
photo of type, Z--photo of type). 


[53]. AEGIPHILA STRICTA Rusby. 

The species described by me on pages 352 and 353 under 
the name of A. stricta must take on the older name of A. 
laeta H.B.K., which see in these supplementary notes for 
details [Phytologia 1: 228--229. 1937]. 


125. AEGIPHILA SUFFLAVA Moldenke. 

Additional citations: PERU: Loreto: Killip & Smith 27,439 
(A--photo of type, B--photo of type, D-=-photo of type, F-- 
photo of type, G--photo of type, K--photo of type, N--photos 
of type & of isotype, P--photo of type, S=--photo of type, 
W--photo of type, Z--photos of type & of isotype); Klug 2076 
(B, E, G, K, Mi, W); Tessmann 5155 (Z--photo); L. Williams 
8225 (Z--photo). 


84. AEGIPHILA SURFACEANA Moldenke. 

Additional citations: BRAZIL: Amazonas: Rob. Schomburgk 
981 (Bm, Cb--3, F, K, Le, N--3 photos, P, Z--4 photos ); 
Spruce 2297 (K—~2, N--photo, Z--photo); Tate 162 (A--photo 
of type, Be-phnoto of type, D-~photo of type, F--photo of 
type, G--photo of type, N--2 photos of type, P--photo of 
type, S--photo of type, We-photo of type, Z--2 photos of 
type); Pard: Dahlgren & Sella 173 (A--photo, 8--photo, D-- 
photo, G--photo, N=-photo, P--photo, S--photo, -photo, Z-- 
ans Huber 2022 (F--photo, Z--photo), 3296 (F--photo, Z-- 
photo). 


55. AEGIPHILA SWARTZIANA Urb. 

The fruiting-calyx is shallowly cupuliform, black in dry- 
ing, about 3 mm. long and 7 mm. in diameter, glabrous, some- 
what striate, its margin subtruncate or slightly and irregu- 
larly erose and subscarious. The fruit is oblong, 4--5 m. 
long and wide, glabrous, not nitid, blackening in drying. . 

Additional citations: JAMAICA: W. Harris 11,716 (A--phota 
B--photo, Bm, D--photo, E, K, N--4 photos, P, W--photo, Z--4 
photos); Swartz s.n. [Jamaica] (A--photo of type, B--frag- 
ment of type, B--photo of type, Bm--isotype, D--photo of 
type, F—photo of type, G--photo of type, N--photo of type, 
W--photo of type, Z--photo of type); Wolle s.n. (G). 


50. AEGIPHILA SYLVATICA Moldenke. 

Additional citations: COLOMBIA: Santander Sur: Killip & 
Smith 14,849 (B--photo of isotype, D--photo of isotype, F-- 
photo of isotype, G--photo of isotype, K--photo of isotype, 
N--2 photos of isotypes, P-~photo of isotype, S--photo of 


270 PH Y-T-O;beO-Gsi-& No. 7 


isotype, W--photo of isotype, Z--photos of type & of 5 iso-= 
types). 


1. AEGIPHILA TERNIFOLIA (H.B.K.) Moldenke. 

The specimen of Goudot s.n. in the Kew herbarium, cited 
below, is anomalous in having its leaves merely opposite in- 
stead of ternate as is usual in this species, The name 
"Amerina ternifolia (H.B.K.) DC." listed as a synonym on 
page 280 of my monograph, should more accurately be written 
"Amerina ternifolia (H.B.K.) P. DC." 

Additional citations: COLOMBIA: Cundinamarca: Goudot s.n. 
[Near Bogota] (A--photo, B--photo, D--photo, F--photo, G—- 
photo, K, N--photo, P=--photo, S--photo, W--photo, Z—photo); 
Department undetermined: Lehmann B.T.690 (Le, N, N--photo, Z 
--photo). LOCALITY OF COLLECTION UNDESIGNATED: Herb. Baillon 


an. (P). 


56. AEGIPHILA TRIFIDA Sw. 

This binomial is occasionally written "Aegiphila 3-fida". 
The species has been confused with Ixora fasciculata and 
Chomelia sp., and herbarium specimens have been distributed 
under those names! 

Additional citations: JAMAICA: N. L. Britton 4053 (K, Z-- 
photo); W. Harris 5892 (B, Bm), 6144 (B, Bu, Z--photo), 6273 
(B, Be-photo, Bm, D--photo, G--photo, N--3 photos, Os, P-== 
photo, t+-photo, Z—-4 photos), 9372 (B, K); March 902 (B); 
McFadyen s.n. (B, K--2, N=--photo, Z-—photo); Orcutt 5622 
(Ca); Purdie s.n. [Port Royal Mtns., Aug. 1843] (K); Swartz 
s.n. [Jamaica] (A—photos of type & of isotype, B-—-pnotos of 
type & of isotype, Bm--isotype, Cp--isotype, D--photos of 
type & of isotype, Dc--isotype, F--photos of type & of iso 
type, G--photos of type & of isotype, N--photos of type & of 
isotype, P--photo of type, S--isotype, W--photos of type & 
of isotype, Z—photos of type & of isotype). 


9a. AEGIPHILA TRIFLORA Moldenke in Fedde, Repert. 37: 212--~ 

213. 1934. 

Tree, 5--6 m. tall; branches rather stout, obtusely tet- 
ragonal or subterete, glabrous, covered with gray flakey 
bark; branchlets slender, short, gray or buff in color, 
slabrous, often ridged lengthwise; principal internodes very 
variable, 1--10 cm. long; leaves decussate—opposite or ap= 
proximate; petioles 11--21 mm. long, glabrous, deeply canal- 
iculate above, greatly incrassate and more or less expanded 
at base; blades rather firmly membranous, gray-green above, 
oright green beneath, oblong-oblanceolate, 9.5--19 cm. long, 
3.4--65 cm. wide, short-acuminate at apex, entire, cuneate at 
base, glabrous and nitid on both surfaces; midrib slender, 
prominulent in a channel above, prominent beneath; secondar= 


1937 Moldenke, Notes on Aegiphila me ih 


ies slender, 7--9 on each side, promimalent in a channel a- 
bove, prominent beneath, usually very obscurely or not at 
all arcuate-joined at the margins; inflorescence axillary, 
3-flowered, about 5.6 cm. long and 2.5 cm. wide; peduncles 
slender, 2--4 cm. long, ampliate at apex, glabrous; pedicels 
cuneate-ampliate, 11--19 mm. long, glabrous, widened into 
the calyx above; bractlets 2, filiform, about 3 mm. long, 
glabrous, borne at the base of the pedicels; calyx campanu-. 
late, about 6 mm. long and 3.6 mm. wide, glabrous, attenuate 
at base into the flattened pedicel, its margin 4dentate, 
its teeth ovate-triangular, about 0.5 mm. long and 1.3 mn. 
wide at base; corolla hypocrateriform, its tube cylindric, 
about 4.9 mm. long, glabrous, its lobes 5, oblong-obovate, 
about 4.6 mm. long and 2.6 mm. wide, rounded or subacute at 
apex; stamens 4, inserted about 1.5 mm. below the mouth of 
the corolla-tube, included; filaments about 0.6 mm. long, 
flattened; anthers sagittate, about 1 mm. long and 0.5 m. 
wide at base; pistil exserted, glabrous; style rather 
stout, about 5.7 mm. long; stigma bifid, its branches about 
0.5 mm. long, divaricate; ovary tetragonal, glabrous, about 
1 mm. long and wide, 4-lobed, 4-celled; fruiting-calyx and 
fruit not seen, 

The type of this remarkable species was collected by 
Ellsworth Paine Killip and Albert Charles Smith (No. 
28,858) in a dense forest at Santa Rosa, on the lower Rio 
Huallaga below Yurimaguas, at an altitude of about 135 m., 
Loreto, Peru, between September 1 and 5, 1929, and is dep— 
osited in the Britton Herbarium at the New York Botanical 
Garden. The fruit is described by the collectors as 
"pink". It seems to be related to A. cuneata from the same 
region. The latter differs conspicuously, however, in its 
short—-petioled or subsessile leaves, setose petioles, char- 
taceous and more or less asymmetrical leaf-blades which are 
setose on their venation, its many-flowered inflorescences, 
and its densely hirsute peduncles and pedicels. 

PERU: Loreto: Killip & Smith 28,858 (N--type, W- 
isotype). 


7. AEGIPHILA TRUNCATA Moldenke. 

Additional citations: COLOMBIA: Cundinamarca: Mutis 3659 
(F--photo, G--photo, K--photo, N--photo, Z-~photo), 5191 
(Cb—-isotype, F--photo of type, G—-photo of type, K-~photo 
of type, N--photo of type, Z--photo of type). 


59. AEGIPHILA UNIFLORA Urb. 

Additional citations: JAMAICA: W. Harris 55353 (A--photo 
of isotype, B-type & 2 isotypes, B--photo of isotype, D-- 
photo -of isotype, G--photo of isotype, K--photo of isotype, 
N——-2 photos of isotypes, P--photo of isotype, S—-photo of 


272 PAY. 2 Ob eo Gian No. 7 
isotype, W--photo of isotype, Z--2 photos of isotypes). 


8. AEGIPHILA VALERII Standl. 

Additional citations: COSTA RICA: Guanacaste: Standley & 
Valerio 45,236 (A--photo, B--photo, D--photo, F--photo, G-— 
photo, K, K--photo, N--photo, P=-photo, S--photo, Y-photo, 
Z--photo), 46,079 (N--photo, Z--photo); J. Valerio 148 (A-- 
photo of type, B--photo of type, D--photo of type, F-=-photo 
of type, G--photo of type, K--photo of type, N--photo of 
type, P--photo of type, S--photo of type, w--photo of type, 
Z--photo of type). ILLUSTRATION: Line-drawing (N). 


107. AEGIPHILA VELUTINOSA Moldenke. 

The "Gook & Gilbert 1362" cited by me on page 472 of my 
monograph is a typographic error for Cook & Gilbert 1382. 

Additional citations: PERU: Cuzco: Cook & Gilbert 1382 
(A--photo of type, Be-photo of type, D--photo of type, F-- 
photo of type, G-=photo of type, K--photo of type, N-—photo 
of type, P--photo of type, S=-photo of type, W--photo of 
type, Z--photo of type). 


5. AEGIPHILA VERRUCOSA Schau. 

Additional citations: VENEZUELA: Aragua: Allart 426 (N-- 
photo, Ve, Z--2 photos); Fendler 841 (Cb, G, K); Jahn 444 
(Ve, Z--photo); Karsten s.n. [Colonia Tovar, 1848] (B--6, Bm 
--2, K, L--3, N, N--2 photos, P, V--2, Z--4 photos), s.n. 
[1847] (B); Moritz 897 (A--photo of isotype, B--photo of 
isotype, Bm--isotype, Cb--2 isotypes, D--photo of isotype, F 
--photo of isotype, G--photo of isotype, K--isotype, N-- 
fragment of isotype, N--photos of type & of 5 isotypes, S-= 
photo of isotype, W--photo of isotype, Z--photos of type & 
of 3 isotypes), s.n. (Bm); H. Pittier 9347 (Ba, Z--photo). 


37. AEGIPHILA VERTICILLATA Vell. 

An additional synonym is Aegiphila lanata Casar., in 
herb. [not A. lanata Moldenke, 1933]. The "Dusén 7982" cited 
by me on page 473 of my monograph is a typographic error for 
Dusén 7980. 

Additional citations: BRAZIL: Mattogrosso: A. Robert 4 
(B, Bm), 434b (Z--photo); kiinas Geraes: Casaretto 2624 (Sb); 
Clausen 15 (Cb), 5.n. [Aug.--April, 1840] (Bm, K--2, Z-- 
photo); Glaziou 13,056 (Br, Cb, Cp, P--3, Z--photo), 20,428 
(B, Br, Cp, K, P, X); F. C. Hoehne, Com. Rondon 6109 (Sp); 
Lindberg 496 (Br); Mosen 1533 (N--photo, P, S, Us, Z-—photo) 
4332 (N--photo, $, Z--2 photos); Regnell 1.310 [1845] (Ut), 
1.310 [1856] (S), 1.310 [1862] (K, Z--photo), 1.310 [1866; 
Herb. Monac. 1657] (A--photo, Be-photo, Br, Cp--2, D--photo, 
F--photo, G--photo, Mu, N--photo, Ol--2, P=-3, Vu, ii--photo, 
Z--photo), II.310 [1856] (Us), I1.310 [1874] (Us). 


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PHY TOLOGIA 


Designed to expedite botanical publication 


Vor. I May, 1938 No. 8 


CONTENTS 


H. N. Moxpenxe: Studies of New and Noteworthy Tropical 
AIETILAU Pages se TLE Eh sie ohesten 4s 1273 


C.L. Lunpert: Two New Species of Plenckia. si ee ee 284 
B. A. Kruxorr: Two New Species of Erythrina from Central America .286 


H. N. Moxpenxe: Additional Notes on the Genus Aegiphila--IV . . . 289 


a a 


Published by H. A. Gleason and Harold N. Moldenke 
The New York Botanical Garden 
Bronx Park, New York, N. Y. 
Price of this number, 50 cents; per volume, $5.00 in advance. 


Vol. 1, number 7, was issued September 21, 1937 


Se ee ace 


iran =a™ = 
new YORE 
BOTANICAL 


(A ro piad 
STUDIES OF NEW AND NOTEWORTHY TROPICAL AMERICAN PLANTS-- III 


Harold N. Moldenke 


AVICENNIA TONDUZII Moldenke, sp. nov. 

Arbor; ramulis sarmentisque gracilibus articulatis dense 
brevissimeque adpresso-tomentellis, pilis cinereis vel sord- 
idis; petiolis gracilibus 0.8--2 om. longis dense brevissi- 
meque adpresso-tomentellis; laminis coriaceis elongato- 
oblongis, 9--17 om. -longis, 1.7~-2.8 cm. latis, ad apicem 
obtusis, integris, ad basin longe attenuatis vel acuminatis 
et in petiolum prolongatis, supra dense minuteque pulveru- 
lentis et impresso-punctulatis glabrescentibus, subtus dense 
brevissimeque adpresso-tomentellis, pilis cinereis vel sord- 
idis; inflorescentiis paniculatis axillaribus terminalibus- 
que 3--6 cm. longis et latis multifloris brachiatis. 

Tree; branchlets and twigs slender, jointed, densely 
matted-tomentellous with grey or cinereous-sordid hairs, 
swollen and annulate at the nodes; leaves decussate-oppcite; 
petioles slender, 0.6--2 cm. long, densely matted-tomentell- 
ous like the twigs, wrinkled-striate in drying; blades cori- 
aceous, elongate-oblong, 9--17 cm. long, 1./--2.8 cm. wide, 
obtuse or blunt at apex, entire, long-attenuate or acuminate 
at base and prolonged into the petiole, densely and very 
minutely pulverulent and impressed-punctate above, becoming 
glabrescent in age, densely matted-tomentellous beneath with 
cinereous or sordid hairs like the twigs; inflorescence cy- 
mose, paniculate, axillary and terminal, 3--6 cm. long and 
wide, usually regularly several-branched from the very base, 
the branches many-flowered; peduncles, axis, and branches of 
the inflorescence densely matted-tomentellous and wrinkled- 
striate in drying like the petioles; bractlets ovate, a pair 
subtending seach pair of inflorescence-branches; flowers ses- 
sile; prophylle 3, ovate, about 2 mm. long and 1.5 mm. wide, 
strigose at the center on the back, villous-tomentose toward 
the margins, rounded or obtuse at apex, simulating sepals 
and closely appressed to them, imbricate; sepals 5, separate, 
imbricate, broadly elliptic or subrotund, about 2.5 mm. long 
and wide, rounded at apex and base, densely villous-—tomen- 
tose on the back; corolla hypocrateriform, its tube broadly 
cylindric, short, straight, about 1 mm. long, glabrous on 
both surfaces, its limb 4—parted, the lobes equal, wide- 
spreading in anthesis, narrowly elliptic-lingulate, about 3 
mn. long and 1.5 mm. wide, rounded at apex, densely toment- 
ose on both surfaces; stamens 4, inserted about 0.5 mm. a- 
bove the base of the corolla-tube, equal, exserted; fila- 
ments slender, about 1.5 mm. long, glabrous; anthers oblong, 


275 


274 PHYTOLOGIA No. 8 


about 0.75 mm. long and 0.375 mm. wide, 2-celled, not appen- 
daged, opening by longitudinel slits; pistil 1, compound, 2= 
carpellary; ovary ovate-subglobose, about 1.25 mm. long and 
wide, densely appressed-villous with antrorse hairs, not 
lobed, incompletely 4-celled; style terminal, comparatively 
stout, about 0.75 mm. long, densely appressed-villous; stig- 
ma bifid, its branches about 0.25 mm. long, unequal; ovules 
borne on a basal 4-winged placenta, pendent. 

The type of this species was collected by Adolfe Tonduz 
(No. 6776) in the littoral zone bordering the Pacific Ocean 
at Punta Mala, Costa Rica, in March, 1892, and is deposited 
in the herbarium of the Jardin Botanique de 1'Etat at Brus- 
sels. A complete discussion of this species, citation of 
specimens, and comments on its relationships will be found 
in my forthcoming monograph of the genus. 


AMBRINA AMBROSIOIDES var. ANTHELMINTICUM (L.) Moldenke, 
comb. nov. Chenopodium anthelminticum L. Sp. Pl., ed. l, 
220. 175. 


BALSAMITA MAJOR var. TANACETOIDES ( Boiss.) Moldenke, comb. 
nov. Pyrethrum Balsamita tanacetoides Boiss. Fl. Orient. 
Sy 346. -1875. ~ 


CISSAMPELOS FASCICULATA Benth. 

Additional specimens representing this species are Tess- 
mann 3183 from Middle Ucayali, Peru, and 5067 from Iquitos, 
Loreto, Peru, and Krukoff 7274, collected on campinarana on 
the plateau between Rio Livramento and Rio Ipixuna, Munici- 
pality Humayta, in the basin of the Rio Madeira, Amazonas, 
Brazil, between November 7 and 18, 1934. 


DISOIPHANIA KILLIPII Diels. 

This species has been hitherto known only from the type 
collection, Killip & Smith 27,041, collected at Iquitos, 
Loreto, Peru, in August, 1929. Recent work on this and re- 
lated groups of Menispermacesae by Mr. B. Ae Krukoff and ny- 
self has brought to light another collection, kKrukoff 8249, 
collected on high land, terra firma, near Palmares, Munici- 
pality of SHo Paulo de Olivenga, Amazonas, Brazil, between 
September 11 and October 26, 1936, described as a vine. 
This constitutes a notable range extension for the species. 


HYPERBAENA DOMINGENSIS (P. DO.) Benth. 

According to Diels in Engler, Pflanzenreich 47%: 200-201 
(1910), this is a very widely distributed species, ranging 
from Cuba to southernmost Brazil. The type was collected by 
Poiteau in H#iti. It seems rather doubtful to me whether 
all of the material commonly regarded as representing this 


: 
5 


1938 Moldenke, Tropical American Plants 275 


species, or even all of thes material cited by Diels, is act- 
ually conspecific. If it is, then the species is certainly 
extremely polymorphic in its floral characters. Blanchet 
2346 and Spruce 3167 (both cited by Diels) have been care- 
fully dissected. In spite of the much more filiform-elongate 
inflorescences of the latter (which doubtless caused hiiers 
to propose for it the name H. graciliflora liers, of which 
it is the type collection), the staminate flowers are pract- 
ically identical. The sepals are conspicuously dark-spotted 
toward the center and base, translucent at the margins, the 
outer ones lanceolate, about 0.75 mm. long and O.5 mm. wide, 
sparsely villous toward the center and base on the back, the 
inner ones obovate, about 1 mm. long and 0.75 mm. wide, 
glabrous. The petals are obovate, not at all rhomboid, about 
O.6 mm. long and 0.3 m. wide, glabrous. The stamens are 0.5 
--0.75 mm. long, separate to the base, spreading, the anth- 
ers large and conspicuously projecting. Blanchet 1556, iden- 
tified as this species by both Eichler and Diels, but not 
cited by Diels, has only pistillate flowers. These differ in 
having the sepals minutely punctulate, but not at all dark- 
spotted, membranous, slightly more incrassate toward the 
center and base, the outer ones ovate, 1.4--1.6 mm. long, a- 
bout 1.2 mm. wide, blunt or subacute at apex, rounded- 
truncate at base, the inner ones obovate-elliptic, about 1.5 
mm. long and 1.3 mm. wide, rounded at both apex and base, 
concave within, and the petals very tiny, obovate, about 
0.25 mm. long and 0.125 mm. wide, rounded at apex, attenuate 
at base. H. H. Smith 1553, from Santa Marta, Colombia, iden- 
tified as "Abuta prob. sp. nov. aff. Selloana" by A. C. 
Smith and as Hyperbaena domingensis by B. A. Krukoff, dif- 
fers in having its sepals membranous, not dark-spotted, 
lighter at the margins, the outer ones lanceolate, about 1 
mm. long and 0.5 mm. wide, obtuse at apex, truncate at base, 
sparsely strigose on the back and appearing ciliate along 
the margins, the inner ones broadly elliptic, about 2.25 m. 
long and 1.25 mm. wide, rounded at apex, subacute at base, 
glabrous but minutely punctulate throughout; petals 6, mem 
branous, obovate, not rhomboid, about 1.3 mm. long and 0.7 
mm. wide, rounded at apex, gradually narrowed to the acute 
base, concave within, glabrous and minutely punctulate 
throughout; stamens 6, separate to the base, subequal, 1-- 
1.25 mm. long, glabrous, incurved below the apex, slightly 
ampliate at apex; anthers small, but rather conspicuous, 2= 
celled, opening by longitudinal slits. Duss 3220, from Gua- 
deloupe, cited by Diels, differs yet again in various de- 
tails of floral characters from the South American specimens 
just described ana nas even smaller flowers. If all these 
collections actually represent the same species -- and in 
foliar characters they are indistinguishable -- then the 


276 PHY T0206 D-8 No. 8 


sheet of Poiteau s.n. from French Suiana, in the Kew herbar- 
ium, and Dusén 12,058 from Paran&, Brazil, and F. C. Hoehne 

pl. viv. 137 137 [Herb. Instit. Biol. S. Paulo 28,450], cultiva- 

ted in the Jardim Botanico at S&o Paulo, Brazil, probably 
also belong here. 


HYPERBAENA HASSLERI Diels. 

Additional synonyms ere Abutea parvifolia Rusby, in herb., 
and Hyperbaena ovalifolia (Mart.) Chod. & Hassler ex lialme, 
in herb. (sphalm.). 

This species was known to Diels only from the type col- 
lection, Hassler 7299. Recent revisionary work in this group 
has brought to light two more Paraguayan collections, Has— 
sler 12,223 and Malme 1058, and likewise H. H. Rusby 1 
from the pampas near near Lake Rogagua, Bolivia, and Jorgensen 
2061 from the Territorio de Chaco, Argentina. The Rusby col- 
lection (in fruit) was originally distributed as Abuta parvi 
folia and the J&rgensen collection (flowers and fruit) as 
Hy per baena domingensis. Thus the range of this species is 
considerably extended and fruits are known for the first 
time. The fruits may be briefly described as follows: drupes 
oblong or slightly obovate, 1.3--1.6 mm. long, 6-—-7 m. wide 
(when dry), the exocarp fleshy, black in drying, glabrous, 
nitid. It is worthy of note that tne characters given by 
Diels in hie key to the species of Hyperbaena (op. cit. p. 
199) as distinguishing H. domingensis from H. Hassleri are 
just reversed. The line reading "Folia glabra" should lead 
to H. domingensis and the lines reading "Folia subtus pilosa” 
should lead to H. Hassleri. It is also worthy of note that 
the inflorescences of H. Hassleri are not always 5.5--/ cm. 
in length and strongly resembling those of H. domingensis, 
as stated by Diels. In the specimens cited above the inflor- 
escences range from 0.4 to 4 om. in length and are not near- 
ly as strong, complex, or elongate as those of H. domingen- 
sis, and yet very much larger and more conspicuous than 
those of H. oblongifolia. 


HYPERBAENA OBLONGIFOLIA (Mart.) Chod. & Hassler. 

An additional synonym is Abuta oblongifolia (Mart.) Miers, 
Ann. Mag. Nat. Hist. III, 14: 258. 1864. 

This species was cited by Diele (op. cit. p. 202) only 
from Paraguay and Rio de Janeiro, Brazil. Recent revisionary 
work on the group has brought to light two more Paraguayan 
collections -- Fiebrig 5946 and 6403 -- and also a collect- 
ion from Minas Geraes, Brazil -- Regnell III.1/720. An emend- 
ed description of the staminateflowers of this species, bas- 
ed on these new specimens, follows: sepals 6, menbranous, 
slightly incrassate toward the center and base, not spotted, 
the outer 3 narrowelliptic, 1.1--1.2 mm. long, 0%5--O.7 mm. 


1938 Moldenke, Tropical American Plants 277 


wide, acute or blunt at apex, blunt at base, rather sparsely 
villous-pubescent on the outside with long appressed yellow 
ish hairs, the inner 3 broadly elliptic, 1.25--1.6 mm. long, 
1--1.25 mm. wide, blunt or rounded at apex, subacute or 
rounded at base, very sparsely appressed-villous in the cen- 
ter (from base to apex) on the outside; petals 6, membran- 
ous, broadly elliptic or ovate-elliptic (or subobovate in 
outline when not spread out), 0.75--1.12 mm. long, 0.5--1 
mme wide, blunt or rounded at apex, acute at base, with two 
lobes below the middle, which are turned in and envelop the 
filaments, glabrous throughout; stamens 6, separate to the 
base, spreading, 0.75--0.875 mm. long, equal, glabrous; fil- 
aments slender, ampliate toward the apex or subgibbous, of- 
ten curvate and humped at the apex; anthers large and prom- 
inent, swollen, 2-celled, plainly 2-lobed, opening by: longi- 
tudinal slits. 


HYPERBAENA SOLIMOESANA Moldenke, sp. nov. 

Frutex scandens; ramis dense elongato-lenticellatis, min- 
ute puberulis vel glabratis; ramulis gracilibus striatis 
glabratis, lenticellis suberosis prominentibus, supra nodos 
saepe in panno elongato angustato velutino-tomentosis; peti- 
olis gracilibus 1.1--7.1 cm. longis glabris, ad apicem curv- 
etis incrassatisque; laminis firme chartaceis vel subcoria- 
ceis ellipticis, 7--25 cm. longis, 3.4--10.4 cm. latis, ad 
apicem acuminatis, integris, ad basin obtusis vel acutis, 
utrinque glabris nitidisque. 

Woody vine; branches medium-slender, densely lenticell- 
ate, minutely puberulent or glabrate, the lenticels elong- 
ate, abundant; branchlets and twigs slender, usually more or 
less longitudinally striate (when dry) and dark, with fewer, 
less elongate, corky=prominent lenticele, glabrous except 
for a patch of densely velutinous tomentum often borne in a 
narrow band between the petiole—base and the base of the 
next succeeding inflorescence; petioles slender, 1.1--7.1 
em. long, curved and thickened at the apex, glabrous; blades 
firmly chartaceous (a) or subcoriaceous, elliptic, 7--25 cm. 
long, 3.4—10.4 cm. wide, acuminate at apex, entire, obtuse 
or acute at base, glabrous and nitid on both surfaces, S- 
pli-nerved, the 2 marginals and 2 laterals all issuing from 
the very base of the blade; midrib and laterals (not the 
marginals) flat or subimpressed above, very prominent be- 
neath; vein and veinlet reticulation rether abundant, the 
ultimate portions mostly obscure above (except on very large 
basal leaves) and flat beneath, the larger portions prominu- 
lent on both surfaces; inflorescence supre-axillary, solit- 
ary or 2--4 superposed, 2--8 cm. long, densely many-flowered; 
peduncle snd rachis rather slender, but not filiform, glab- 
rous except for velutinous-tomentose patches at the base of 


278 PHY T°O:L-0 @ Fa No. 8 


each branch, the branches short and puberulent, several- 
flowered; pedicels obsolete or very short and puberulent; 
staminate flowers: sepals 6, imbricate, membranous, not 
spotted, the 4 outer ones narrowelliptic, 1--1.5 mm. long, 
0.57--0.5 mm. wide, subacute at apex, rounded-obtuse at base, 
rather densely and irregularly strigose throughout on the 
outer surface with appressed yellow hairs which project over 
the margins in ciliate fashion, the inner 3 broadly elliptic 
or elliptic-ovate, 1.7--1.9 mm. long, about 1.2 mm. wide, 
rounded at apex, abruptly acute at base, very densely tomen- 
tellous on both surfaces with short yellow hairs; petals 6, 
membranous, rhomboid-obovate, 0.6--O.7 mm. long and wide, 
sharply acute at apex, attenuate-acute at base, densely tom- 
entellous on both surfaces from about the widest portion to 
the apex with spreading yellowish projecting hairs; stamens 
6, separate to the base, about 0.75 mm. long, equal, free 
from the petals, hardly at all adherent at base, glabrous, 
ampliate at apex; anthers small, not conspicuously project- 
ing nor swollen, 2=-celled, opening by longitudinal slits, 
not prominently 2-lobed; drupes oblong-elliptic, 1.6--1.8 
mn. long and 6--10 mm. wide (when dry), sessile, usually 
paired, glabrous, the exocarp fleshy, black in drying. 

The type of this species was collected by Boris Alexander 
Krukoff (No. 8924) in a high forest on terra firma in the 
basin of the creek Belem, Municipality of So Paulo de Oliv— 
énga, basin of Rio Solimoes, Amazonas, Brazil, between Octo- 
ber 26 and December 11, 1936, and is deposited in the Britt— 
on Herbarium at the New York Botanical Garden. The descrip-— 
tion of the fruit is taken from Krukoff (279, collected on 
Campinarana on the plateau between Rio Livramento and Rio 
Ipixuna, Municipality Humayta, basin of Rio Madeira, Amazon— 
as, Brazil, between November 8 and 18, 1934, since this col- 
lection probably represents the same species. Gleason 827, 
described as a vine 20 feet long, colletted in dense upland 
forest at Rockstone, British Guiana, between July 15 and 
August 1, 1921, probably represents an older basal branch of 
the same species, thus accounting for the much larger 
leaves. This collection was originally identified and dist- 
ributed by H. A. Gleason as a species of Piper and later de- 
termined as "Abutea concolor P. & E. ?" by N. E. Brown. 

H. solimoesana | differe notably from all other knowm spec- 
ies of Hyperbaena Hyperbaena in its flat or indistinct tertiary and 
veinlet reticulum and in the characters of its inflorescence. 
In its foliar and inflorescence characters, in fact, it 


closely resembles some species of Abuta, notably A. Selloana 


Eichl. It certainly differs pronouncedly from all other 
known species of Hyperbaena from central tropical South Am 
erica. Its tomentose petals also seem to plece it in a group 
apart from the rest of the genus. 


Se ee 


1938 Moldenke, Tropical American Plants 279 


LANTANA CORDOBENSIS Moldenke, nom. nov. 


Tamonopsis spicata Griseb., Abh. K. Ges. Wiss. Gdtting. 
19: 246. 1874 [not Lantana spicata Vell. Fl. Flum. 254, 


LANTANA JUNELLIANA Moldenke, nom. nov. 


Lippia lantanifolia var. crenata Griseb., Abh. K. Ges. 
Wiss. G8tting. 19: 243. 1874. 


LIPPIA GRISEBACHIANA Moldenke, nom. nov. 
eaniaee lantanifolia Griseb., Abh. K. Ges. Wiss. Gdtting. 
242 


19: . 1874 [not Lippia lantanifolia F. Mill. Fragm. 6: 
151. 1868]. 


UNGULIPETALUM Moldenke, gen. nov. 

Herbae scandentes. Radices elongatae tuberoso-incrassatae 
filipendulae (an parasitico-offensae?). Caules e basi erecta 
sursum volubiles tenues. Folia alterna longe petiolata, lam- 
ine membranacea vel temiter pepyracea cordata vel aequilat- 
erali-triangulari plinervia. Inflorescentiae cymosse, cymae 
parcae in paniculam compositae longe pedunculateae. Flores 
pedicellati dioeci. Flores @: sepala 9, 6 exteriora in seri- 
ebus duis dispositea lanceolato-ovata extus subvillosa, 3 in- 
teriora multo majora lanceolato-ovata, parte basali non cam- 
panulato— vel tubuloso-conniventia, parte anteriore non re- 
flexa. Petala 6 late elliptica bilobata, ad apicem retusa, 
ad basin valde unguiculatas; lo»is tumidis translucentibus, 
in parte ventrali confluentibus, in parte dorsali non con- 
fluentibus sed secus costam petali in carina dua parallela 
longitudinaliter elevatis, ad basin cordatis, glabris mem- 
branaceis. Stamina 6 usque ad mediam connata, parte libero 
filamentorum erecto incrassato glabro, ad apicem incurvato 
in connectivum non producto. Antherae subimuersae terminales 
biloculares, thecae oblongae, rimi longitudinali dehiscent- 
es, in anthesin plerumque horizontaliter dispositae. Flores — 
9: sepala 12, 9 exteriora conico-ovata imbricata extus sub- 
villosa, 3 interiora multo majora ovata, parte basali non 
campanulato— vel tubuloso-conniventia, parte anteriore non 
reflexa. Petala 6 ovata vel hastato-sagittata incrassata, ad 
basin unguiculata glabra, lobis non tumidis. Staminodia nul- 
la. Carpella 6--9. Ovarium gibboso-semiovatum dense villos- 
um. Stylus teres subfiliformis elongatus, ad apicem tmcine- 
tus. 

Scandent herbs; roots elongate, bearing several to many 
elongate thickened tubers, perhaps parasitic; stems erect at 
base, twining above, slender; leaves alternate, long-petiol- 
ate, the blades membranous or thin-papery, cordate or equi- 
lateral-triangular, pli-nerved; inflorescence cymose, axill- 
ary, the cymes few, disposed in long-pedunculate panicles; 


280 PET F:0 &-0:G-7°4 No. 8 


flowers pedicellate, dioecious; male flowers: sepals 9, the 
6 outer ones in two series of 3 each, lanceolate-ovate, sub- 
villous outside, the 3 inner ones much larger, lanceolate- 
ovate, not campanulate=- or tubular-connivent at the base, 
not reflexed at the apex; petals 6, broadly elliptic, bilob- 
ed, retuse at apex, conspicuously clawed at base, the lobes 
swollen, translucent, confluent on the ventral side, on the 
dorsal side the inner margins not confluent, but raised in 
the form of two parallel longitudinal keels along the midrib 
of the petal, cordate at base, glabrous, thin-membranous; 
stamens 6, firmly connate to about the middle, the free por- 
tion of the filaments srect, thick, glabrous, curved inwards 
at the apex, the connective not appendaged; anthers termin- 
al, 2-celled, the thecae oblong, dehiscing by means of a 
longitudinal slit, usually borne horizontally during anthe- 
sis; female flowers: sepals 12, the 9 outer ones conic= 
ovate, imbricate, subvillous outside, the 3 inner ones much 
larger, ovate, not campanulate- or tubular=connivent at the 
base nor reflexed at the apex; petals 6, ovate or hastate- 
sagittate, incrassate, glabrous, clawed at the base, the two 
lobes not noticeably swollen or inflated or keeled; stamino- 
des absent; carpels 6--9; ovary gibbous, half-ovate, asymet- 
ric, densely villous; style terete, slender, elongate, hook- 
ed at the apex. 

This genus, whose name is taken from two Latin words, 
ungula, a claw, and petalum, a petal, is thus far know only 
from a single species, a discussion of which follows below. 
It is probably a member either of the tribe Tinosporeae or 
the tribe Cocculeae of the family Menispermaceae. It seems 
to be most closely related to the genus Cocculus P. DC.,; 
from which it differs conspicuously in its tuberous filipen- 
dulous roots, its staminate flowers having 6 outer stamens 
in two series of 3 each, inflated clawed and bilobed petals, 
and 6 connate stamens, and its pistillate flowers having 12 
sepals, elongate uncinate styles, and densely villous ovar- 
ies. It also bears resemblances to the genus Odontocarya 
Miers, from which the same characters and the lanceolate- 
ovate shape of its sepals distinguish it. 


UNGULIPETALUM FILIPENDULUM (Mert.) Moldenke, comb. nov. 

Cocculus Filipendula Mart., Flora 24: Beibl. 2: 43. 1641. 
Odontocarya filipendula (Mart.) Miers, Contrib. Bot. 35: 65. 
1871. Ohondodendron filipendulum (Mart.) Diels in Engl. 
Pflanzenreich 447: 81. 1910. 

The type of this species, which is the type species of 
the gems, was collected by Maximilian Alexander Philipp, 
Prinz zu Wied-Neuwied, in woods at Cabo Frio, on September 
8 (or a few days previously), 1615. According to Eichler in 
Mart. Fl. Bras. 15': 184 (1864) it was also collected in 


te ee ee 
. 


aerree: 


1938 Moldenke, Tropical American Plants 281 


southeastern Brazil by Sellow (No. 596) and by Luschnath. 
Eichler separates a variety from the species, to which, how 
ever, he does not assign a formal name. He states that, the 
true species, represented by the Wied-Neuwied and Sellow 
specimens and perhaps a "Mart. Herb. Fl. Bras. 283", has 
petioles only 1--2 inches long, leaf-blades 1--2 1/2 inches 
long and 1—-2 inches wide, and sparse pubescence throughout. 
The variety, on the other hand, represented by the Luschnath 
collection, has petioles 3--4 inches long, leaf—-blades 4—5 
inches long and wide, and denser pubescence throughout. App- 
arently all four of these collections were sterile, since no 
mention is made of floral or fruit characters either by 
Martius, Eichler, or Miers, or by Walpers in his Repert. 2: 
748 (1843), where the species is discussed. Diels saw only 
pistillate flowers, obtained from the SHo Paulo portion of 
Glaziou 13,520. He cites, however, an unnumbered Schwacke 
collection also from Cabo Frio. 

Perhaps Eichler's variety can be maintained, although not 
having seen any of the collections which he cites, I cannot 
commit myself nowe The four sheets of Glaziou's collections 
before me, however, exhibit leaves with the blades varying 
from 2.9 cm. long and 2.5 cm. wide to 9.4 cm. long and 5.6 
cm. wide and petioles from 0.8 to 3.1 cm. in length. 

The species is said to be called "abuta miuda" by the 
natives and to be used medicinally as an antidote. Although 
the specific name was originally spelled with a capital ini- 
tial letter it certainly has no connection with the genus 
Filipendula Tourn. and therefore does not need to be capit- 
alized. Miers was the first to write it with a small initial 
letter, both under Cocculus and under Odontocarya, in Con- 
trib. Bot. 3: 65 (1871). 

The species is quite distinct in its vegetative and flor- 
al characters from all known members of the South American 
Menispermaceae. It has been carefully compared by B. A. Kru- 
koff and myself during the course of our studies of the bot- 
anical ingredients of "curare" with the genera Sciadotenia 
Miers, Hyperbaena Miers, Somphoxylon Eichl., Abuta Barrére, 
Anomospermum Miers, Cissampelos L., Synandropus A. C. Smith, 
Chondodendron Rufz & Pav., Oocculus P. DC., Odontocarya 
Miers, and the other new genus which will be proposed in our 
“curare" paper. The only genera of the list in which it 
could conceivably go are the last two and its floral and 
habital differences from these two have been mentioned abow. 
Martius, Walpers, Eichler, and Glaziou considered it to be- 
long in the genus Cocculus, although Eichler placed it there 
with an interrogation-point, Miers thought it was an Odonto- 
carya, and Diels regarded it as an anomalous Chondodendron. 
These three genera belong in three distinct tribes of the 
family, according to Diels' monograph in Engler, Pflanzen- 


282 PHYTOLOGIA No. 8 


reich 474; 46~-47 (1910), but it must be remembered that all 
of the earlier collections were apparently sterile and not 
even Diels saw flowers of both sexes. He says of it "Quae 
stirps singularis a speciebus ceteris longius distat". 

Since an abundance of material of both staminate and pis- 
tillate flowers has been available to me, thanks to the kind 
courtesy of the curators of the Paris and Kew herbaria, I 
give herewith a detailed description of both types of flor 
ers: staminate flowers: outer sepals 6, in two series of 3 
each, lanceolate-ovate, the smaller ones about O.5 mm. long 
and 0.125 mm. wide, the larger ones about 1.25 mm. long and 
O.5 mm. wide, irregularly long-pubescent on the outer surf- 
acé with subvillous multicellular brownish hairs; inner sep- 
als 3, lanceolate-ovate, thin-textured, 2.2--2.5 mm. long, 
O.8--1 mm. wide, acute at the apex and base, densely short- 
puberulént with more or less glandular or capitate hairs on 
the outer surface; petals 6, broadly elliptic, retuse at the 
apex, conspicuously clawed at the base (the claw heavy, 
firm, and dark and extending as a heavy, firm, and dark mid=- 
rib to the apex of the petal, but rapidly diminishing as the 
apex is approached), with 2 inflated translucent lobes or 
wings on either side, which are confluent on the ventral 
surface and broadly cordate at the base, the inner margins 
not confluent dorsally, but raised in the form of two paral- 
lel longitudinal keels or crests, glabrous throughout; stam- 
ens 6, connate for about half their length into a dark firm 
cylindric or obconic glabrous structure 0.5--0.8 mm. long, 
the free portion of the filaments stoutish, erect, 0.5--0.8 
mm. long, dark, incurved at the apex; anthers oblong, term- 
inal, 2=-celled, dehiscing by longitudinal slits, borne al- 
most horizontally during anthesis by the curving of the fil- 
ament apex, the connective not prolonged or appendaged; pis- 
tillate flowers: outer sepals 9, conic-ovate, 0./5>--1.5 m. 
long, 0.2--0.6 mm. wide, acute at the apex, trumcate at the 
base, densely long-pubescent on the outer surface with sub- 
villous multicellular brownish hairs; inner sepals 3, ovate, 
3--35.1 mm. long, 1.9%-2 mm. wide, acute at the apex, rounded 
at the base, densely short-puberulent on the outer surface 
with more or less glandular capitate hairs; petals 6, ovate 
or hastate-sagittate, thickened, 1.1--1.2 mm. long, about l 
mm. wide, acute at the apex, the two wings with their basal 
lobes patent, glabrous and dark throughout, the claw broad- 
ened at the apex, the wings not noticeably inflated; pistils 
6--9 per flower; ovary gibbous, semi-ovate, asymuetric, 
flattened on adjacent surfaces, O./--O.8 mm. long, densely 
long-villous; style terete, slender or subfiliform, 1.3--1.5 
mn. long, glabrous, uncinate at the apex. The flowers are 
described by Glaziou as being reddish-brown in color, bloom 
ing in November and December. 


4 
; 
j 


1938 Moldenke, Tropical American Plants 283 


The Glaziou 18,129, cited by Diels, is certainly not con- 
specific with the other Glaziou collections which he cites. 
Its inflorescences are still very immature, but careful ex- 
amination indicates that it probably represents a species of 
Disciphania Eichl. 

Citations: Four specimens have been examined. BRAZIL: Rio 
de Janeiro: Glaziou 8563 (K, P); S&o Paulo: Glaziou 15,520, 
in part [o, S&o Vicente] (K). CULTIVATED: Brazil: Rio de 
Janeiro: Glaziou 13,520, in part [¢’, S&o Christovfo] (P). 


XYLOSTEON CANADENSE Duhan. 

In Rev. Sudam. Bot. 5: 3 (1937) I published a superfluous 
binomial, Xylosteon album (L.) Moldenke. This binomial was 
published on the assumption that Asa Gray [Syn. Fl. 2': 201. 
1878] and Britton & Brown [Ill. Fl., ed. 1, 3: 241 (1898), 
ed. 2, 3: 281 (1917 & 1936)] were correct in their reduction 
of Vaccinium album L. [Sp. Pl., ed. 1, 350. 1753] to Lonic- 
era canadensis Marsh. [Arb. 81. 1785], of which Lonicera 
ciliata Muhl. (Cat. 23. 1813] is a synonym. Actually, accor- 


ding to Blake in Rhodora 16: 118 (1914), they were in error 


in making this reduction. Examination of the type in the 
Linnean herbarium by Blake reveals that it is a species of 
Symphoricarpos, which Blake calls Symphoricerpos albus (L.) 
Blake. Druce [Rep. Bot. Exch. Club 1913, 35: 420. 1914] ap- 
parently also feiled to consult the Linnean type when he 
proposed the binomial Lonicera ulba (L.) Druce for what had 
hitherto been know as L. canadensis, and likewise ignored 
the earlier Lonicera alba of Linnaeus, which is a synonym of 
Chiococca alba (L.) A. S. Hitchc. If the genus Xylosteon is 
to be maintained, as is done by Rafinesque, Howell, R&hling, 
Michaux, Decaisne, Eaton, Buchanan-Hamilton, Dumort de Cour- 
set, Goldie, Bonpland, Ruprecht, Richards, Moench, Fischer, 
Maximowiez, Pursh, Loddiges, Webb, Fuss, and Small, to men- 
tion just a few, then it seems that the name Xylosteon cana- 
dense Duheam. is the proper one for the species in question. 
Unfortunately, the last edition of the International Rules 
invalidates all binomials published in Duhamel's work (1755). 
The binomial, however, occurs frequently in literature, ac- 
credited to Duhamel [e.g., P. DC. Prodr. 4: 337. 1830; 
Steud. Nom. Bot., ed. 2, 2: 793. 1840; Jacks. Ind. Kew. 2: 
1241. 1895]. Probably a valid publication of this binomial 
could be found among the various effective publications, if 
diligent search were made. 


(a) The term "chartaceous" in previous publications of mine 
was mis-used in the sense of "thin-membranous" and the term 
“nembranous" was mis-used in the sense of "chartaceous". In 


284 P-RAy Ps) BOR Ez No. 8 


the present contributions and in all future publications, a 
special effort will be made to use these two terms in their 
proper and more correct sense. 


w= oe —- = =e = = = = @Pe we = = = = = = - = = = 


TWO NEW SPECJES OF PLENCKIA (a) 


C. L. Lundell 


PLENCKIA INTEGERRIMA Lundell, sp. nov. | 

Arbor parva, 3--4 m. alta, 20 cm. diam. Folia alterna, 
integerrima, parva, subchartacea, rufo-punctata, breviter 
petiolata, cuneato-oblanceolata, 1.5--3.3 cm. longa, 3-11 
mme lata, apice rotundata, inconspicue emarginata. Cymae 
axillares. Flores parvi, virides. Pedicelli 4--5 mm. longi, 
glabri. Calyx profunde quinquefidus, lobis minute eroso- . 
ciliolatis, ca. 0.6 mm. longis. Petala 5, ovata, 1.5-——-2 m. 
longa, 1.2--1.4 mm. lata, minute eroso-ciliolata. Stamina 5, 
demum patentia. Ovarium in discum subimmersum, biloculare, 
loculis biovulatis, rarius triovulatis. Fructus samaroideus. 

A small tree, 3 or 4m. high, 20 cm. in diameter; branch- 
lets usually slender, elongated, sometimes abortive and 
spine-like, striate, pulverulent. Leaves alternate, small, 
glabrous or slightly pulverulent, subchartaceous, punctate- 
variegated at first, concolorous and faintly variegated with 
age. Petioles 1.5 mm. long or less, subcanaliculate. Leaf- 
blades cuneate-oblanceolate, usually 2.5 to 3.45 cm. long, 
sometimes only 1.5 cm. long, 5 to 11 mm. wide, apex rounded, 
usually slightly emarginate, costa raised above as a fine 
narrow ridge, nearly obsolete below except at base, veins 
obsolete or nearly so, margin entire, slightly revolute. 
Cymes axillary, less than 1 cm. long, much reduced, usually 
2- or 3-flowered. Flowers green. Pedicels slender, usually 4 
to 5 mm. long, sometimes shorter, glabrous, jointed above 
the base. Calyx deeply 5-lobed, apparently glandular-puberu- 
lent, the lobes 0.6 mm. long or less, rounded, minutely 
erose-ciliolate. Petals 5, ovate, 1.5 to 2 mm. long, 1.2 to 
1.4 mm. wide, apex broadly rounded, patent, margin minutely 
erose-ciliolate. Stamens 5, patent at anthesis; filaments 
subulate, 0.6 mm. long or less, inserted on lower edge of 
disk; anthers basally attached. Disk thick, pentagonal, con- 
fluent with ovary. Ovary 2-celled, with 2 or 3 erect ovules 
in each cell, about two-thirds submerged in disk, the free 
part slightly compressed laterally, 2-angled, tapering into 
the very short style. Stigma minutely bifid. Very young im- 
mature fruits samaroid, with a terminal wing, punctate- 


1938 Lundell, New Species of Plenckia 285 


variegated, the rounded or subtruncate apex crowned by the 
persistent stigma and style, l-seeded. 

Type in the Herbarium of Arnold Arboretum, S. Venturi 
1936, young leaves, flowers, and young fruits, collected at 
Vipas, Department of Francas, Province of Tucuman, Argenti- 
na, at alt. of 850 m., December, 1922. 

Additional specimens examined: ARGENTINA: Province of Tu- 
cuman, Department of Francas, Vipas, alt. 800 m., Oct. 1921, 
Venturi 1376, flowers. Province of Salta, Department of Can- 
delaria, alt. 1000 m., Nov. 15, 1929, Venturi 9786, flowers. 

The abortive spine-like branches, small entire leaves, 
and the much reduced intlorescence are among the outstanding 
characteristics which distinguish P. integerrima. All the 
other species in the genus have larger, serrate or serrulate 
leaves, 


PLENCKIA MICROCARPA Lundell, sp. nov. 

Arbor glabra, parva, 5--6 m. alta. Folia parva, alterna, 
chartacea, crenulato-serrulata, 8-20 mm. longe petiolata, 
ovata, late elliptica, vel suborbicularia, 2.5--3.5, raro 
4.6 cm. longa, 1.6-—-2.7, raro 3.7 cm. lata, apice breviter 
acuminata vel acutiuscula, basi rotundata, obscure subcord- 
ata, vel abrupte acuta, utrinque reticulata. Cymae axillar- 
es. Flores parvi. Pedicelli ca. 1 mm. longi. Calyx profunde 
quinquefidus, lobis rotundatis, minute ciliolatis. Petala 5, 
suborbicularia, 1.3--1.6 mm. longa, minute eroso-ciliolata. 
Ovarium in discum subimmersum, biloculare, loculis biovulat- 
is. Fructus samaroideus, oblanceolati-oblongus, 1.4--1.6 cm. 
longus. 

A glabrous tree, 5 to 6 m. high, much-branched; branchlets 
slender, striate, drying reddish or reddish-black. Leaves 
alternate, small, chartaceous, concolorous or slightly paler 
beneath, minutely stipulate. Petioles very slender, shallow- 
ly canaliculate, 8 to 20 mm. long. Leaf-blades ovate, broad- 
ly elliptic, or suborbicular, usually 2.5 to 3.5 cm. long, 
sometimes as much as 4.6 cm. long, usually 1.6 to 2.7 cm. 
wide, sometimes eg much as 4.7 cm. wide, apex short-acumin- 
ate or acutish base rounded, obscurely subcordate, or ab- 
ruptly acute, margin crenulate-serrulate, reticulata-veined, 
costa and veins prominulous oi both surfaces. Cym7s axill- 
ary, much-branched, usually 1.‘ cm. long or less, rarely 2 
em. long. Flowers minute, whitish. Pedicels about 1 m. 
long. Calyx deeply 5-lobed, the lobes rounded, mimitely cil- 
folate. Petals 5, suborbicular, 1.3 to 1.6 mm. long, minute- 
ly erose-ciliolate. Stamens 5; filaments about 0.5 mm. long, 
inserted on margin of disk; anthers attached at base. Ovary 
almost entirely submerged in disk, 2-celled, with 2 erec. o- 
vules in each cell. Stigma cayitate. Samaras oblanceolate- 
oblong, small, 1.4 to 1.6 om. long, including the wing, apex 


286 PY: O.b-0..6 ie No. 8 


rounded, crowned by the persistent stigma, the body 9 to ll 
mn. long, terete, striate, with the wing decurrent along the 
sides to the base, l-seeded; the seed erect, about 12 m. 
long, dark red and minutely verrucose; cotyledons nearly 
flat, lanceolate-linear, about 1 cm. long, 1.2 mm. wide; 
radicle very short, about O.7 mm. long. 

Type in the Royal Botanic Gardens, Kew, B. Balansa 4477, 
fruits, collected on the east slope of Cerro Hu, near the 
Paraguay, Paraguay, April 10, 1883. 

Additional specimens examined: PARAGUAY: in forest, Cerro 
Hu, near the Paraguay, Oct. 1881, Balansa 3079, flowers. 
BRAZIL: Minas, M. A. Glaziou 16741, flowers. ; 

P. microcarpa differs from P. populnes Reisse in its 
small fruits which do not exceed 1.6 cm. in length, and 
small leaves. From P. bahiensis Loes. it may be readily dis- 
tinguished, according to description, by the small ovate 
leaves and longer petioles. 


- =e se =e ws =F = = = =F = |= = = = 


TWO NEW SPECIES OF ERYTHRINA FROM CENTRAL AMERICA 


Bevie Krukoff 


ERYTHRINA FOLKERSII Krukoff & Moldenke, sp. nov. 

Arbor; ramulis glabris; petiolis glabris inermis; petio- 
lulis subgracilibus glabris; laminis foliorum chartaceis 
plerumque late ovatis glabris, ad basin rotundatis vel sub- 
truncatis; inflorescentiis dense tomentosis; pedicellis den- 
se tomentosis; calyce tubuloso—-campanulatis extus adpresso- 
tomentellis, ad apicem rotundatis et saepe emarginatis. 

A small or medium-sized tree; branchlets stout, grayish, 
glabrous; petioles rather stout, striate, yellowish, 21--23 
em. long, glabrous, unarmed; petiolules rather slender, 
0.8--1.0 cm. long, glabrous; leaflet-blades chartaceous, 
green above, slightly paler beneath, usually broadly ovate, 
11--18 om. long, 7.5--13.5 cm. wide, acute or short-acumin- 
ate at apex, rounded or subtruncate at base, glabrous; mid- 
rib prominent above and beneath; secondaries 6 or 7 per 
side, ascending, prominent above and beneath; tertiaries 
prominulous on both surfaces; veinlet reticulation discern- 
ible to the naked eye above and beneath; inflorescences 17-— 
36 cm. long, densely covered with rather loose brown toment- 


a a 


: 
: 
| 
‘ 
| 
: 
1 
; 
; 


—_- - = 


1938 Krukoff, New Species of Erythrina 287 


um; pedicels 0.4~-0.7 cm. long, with tomentum similar to 
that of the inflorescences; calyx coriaceous, tubular-camp- 
anulate, 1.4-—-2.3 cm. long on the ventral side, 1.1--1.7 cm. 
long on the dorsal side, about 0.3 cm. wide at base, gradu- 
ally ampliate to about 0.8 cm. at apex, asymmetric, rounded 
at apex and often emarginate, usually more or less crenulate 
at apex, appressed-tomentellous throughout; standard firmly 
membranous, oblanceolate, to 8.2 cm. long and 2.4 cm. wide, 
broadly rounded at apex, gradually narrowed to the base, 
scarcely or not at all clawed; wings thin-membranous, obov- 
ate, not at all sagittate or hastate ('), 0.8--0.9 cm. long, 


about 0.3 cm. wide at the widest part, rounded at apex, cun- 


eately narrowed to the base; keel-petals thin-membranous, 
strongly asymmetric, subflabelliform-obovate, 0.8--0.9 cm. 
long, straight on the adjacent sides, flaring toward the a- 
pex on the outer side, 0.4--0.5 cm. wide near the apex, sub- 
cuneate at base; stamens to 6.5 em. long, glabrous; pods 
blackish, 11--27 cm. long, usually not coiled, regularly and 
deeply constricted between each seed, tapering below into a 
stipe about 53 cm. long, abruptly terminated at apex by a 
stiff slender acumination about 2 cm. long; sseds scarlet, 
about 1.1 cm. long, with a distinct dark line extending from 
the hilum for approximately 0.2 cm. toward the chalazal end 
of the seed. 

Type, He. H. Bartlett 11515, collected on February 15, 
1931, between El Cayo and Benque Viejo, in British Honduras, 
and deposited in the Herbarium of the University of MMichi- 
gan. Other collections from the same district are W. A. 
Schipp 935, hiercedes Chanek 74 and 102, and J. B. Kinloch 
9163; also Chas. C. Deam 49 collected near Puerto Barrios, 
Guatemala. 

It is ea pleasure to name this species in honor of Dr. 
Karl Folkers, wio succeeded in isolating several useful alk- 
aloids from the seeds of certain species of Erythrina. 

Until J. B. Kinloch 9163 became available this species 
was represented only by flowers amd fruits. When in flower 
it is devoid of leaves. The species is apparently confined 
to the Atlantic coast. From E. rubrinervia H.B.K. and all 
other species which have the calyx somewhat resembling in 
shape that of the present species, it is immediately dist- 
inguished by its inflorescences and pedicels b=ing densely 
covered with rather loose brown tomentum, by its calyx being 
appressed-tomentellous throughout, and by other characters 
indicated in the diagnosis. 


ERYTHRINA NEGLECTA Krukoff & Moldenke, sp. nov. 

Arbor; ramulis glabris plerumque spinosis; petiolis glab- 
ris plerumque spinosis; petiolulis crassiusculis parce pub- 
erulis mox glabrescentibus; laminis foliolorum subcoriaceis 


288 PHY “T.0.L6°G. 2k No. 8 


rhomboideo-ovatis parce puberulis mox glabrescentibus, basin 
versus rotundatie vel subauriculatis, ima basin triangulato- 
cuneatis (rare truncatis); inflorescentiis glabris; pedicel- 
lis glabris; calyce tubuloso, extus glabris, ad apicem ro= 
tundatis et emarginatis. 

A small spreading tree; branchlets stout, grayish, glab- 
rous, often armed with spines; petioles rather stout, stri- 
ate, brownish or yellowish, 10--22 cm. long, glabrous, often 
armed with short curved spines; petiolules rather stout, 0.9 
--1.5 cm. long, sparingly puberulent, soon glabrescent; 
leaflet-blades subcoriaceous, green above, distinctly paler 
beneath, sparingly puberulent when young, soon glabrescent; 
terminal leaflets usually rhombic-ovate, 7--14 cm. long, 7-= 
15 cm. wide, acute or short—acuminate at apex, rounded or 
subauriculate toward the base and then often triangularly 
cuneate, rarely truncate at base; lateral leaflets similar 
in outline to the terminal ones, but inequilateral and 
smaller; midrib flat or prominulous above, prominent be- 
neath; secondaries 6--& per side, ascending, prominulous a- 
bove, prominent beneath; tertiaries prominulous on both sur- 
faces; veinlet reticulation mostly obscure or indiscernible 
above, the larger portion discernible to the naked eye be- 
neath; inflorescences 16--235 cm. long, glabrous; pedicels a- 
bout 0.7 cm. long, glabrous; calyx coriaceous, tubular, 1.7 
--2.2 cm. long on the ventral side, 1.0--1.7 cm. long on the 
dorsal side, 0.6--0.7 cm. wide, approximately equal in width 
throughout, except at the narrowed and asymmetric base, the 
very apex rounded and emarginate, minutely crenulate, not 
surmounted by an acute triangular tooth, glabrous through- 
out; standard firmly membranous, narrowly oblanceolate, to 
8.6 cm. long, to 1.4 cm. wide at the widest part, subacute 
at apex, gradually narrowed to the acuminate base or short- 
clawed, slightly recurved toward apex; wings thin-membranous, 
lanceolate-sagittate, slightly asymmetric, 0.9--1.0 cn. 
long, 0.2--0.25 cm. wide at the widest part, rather oblique- 
ly subacute at apex, very slightly clawed at base; keel- 
petals thin-membranous, strongly asymmetric, 0.9--1.0 cm. 
long, straight to about the middle on the adjacent sides, 
incurved toward the apex, rather sharply acute at apex, 
flaring on the outer side, to 0.25 om. wide near the base, 
short-clawed at base; stamens to 5.5 cm. long, glabrous; 
pode blackish, about 16 cm. long, often coiled, irregularly 
constricted between some of the seeds, tapering below into a 
stipe about 4 cm. long, abruptly terminated at apex by a 
stiff slender acumination about 2.5 cm. long; seeds scarlet, 
about 0.9 cm. long, with a distinct dark line extending from 
the hilum for approximately 0.2 cm. toward the chalazal end 
of the seed, 

Type, R. S. Williams 572, collected February 24 — March 


7 
{ 


th al ees 


1938 Krukoff, New Species of Erythrina 289 


22, 1908, at Penonome and vicinity, at 50--1000 feet eleva- 
tion, in Panama, and deposited in the herbarium of the New 
York Botanical Garden. Other collections are H. Pittier 
6939, collected at the Agricultural Experiment Station at 
Matias Hernandez in Panama, and J. M. Benitez 9159 from El 
Zapote, Escuintla, Guatemala. 

This species is common in Central America and is well 
represented in herbaria. It has been often confused with E. 
rubrinervia H.B.K. in the past, from which species, however, 
it cam be immediately distinguished by the fact that in the 
latter species the very apex of the calyx is also rounded, 
but is surmounted by a sharply acute triangular membranous 
tooth 1.5--2 mm. long (imparting. to the calyx-apex a sharply 
acute appearance when fresh), the seeds are larger and with- 
out a distinct dark line extending from the hilum toward the 
chalaza, and the leaflets are narrower, acuminate or long- 
acuminate at the apex, cuneate or rarely rounded at the 
base, end only slightly paler beneath. 


ERYTHRINA EGGERSII Krukoff & Moldenke, nom. nov. 
Erythrina horrida Eggers, Fl. St. Croix 45. 1879 [not Er- 
ythrina horrida Moc. & Sessé ox P. DC. Prodr. 2: 413. 1825]. 


ADDITIONAL NOTES ON THE GENUS AEGIPHILA -- IV 


Harold N. Moldenke 


The following notes constitute a continuation of those 
published in Phytologia 1: 182--208, 222--240, and 248--272 
(1937). Herbarium abbreviations herein employed, in addition 
to those. published in Brittonia 1: 249--250 (1934) and Phy- 
tologia 1: 182 and 222 (1937) are: Gt = Botanische Anstalt- 
en, GSttingen; hr = B. A. Krukoff Herbarium, New York Botan- 
ical Garden, New York City; and Na = Natal Government Herb- 
erium, Durban, Natal. 

The generic name is mis-spelled "Aegiphita" in Fedde, 
Repert. 42: 248 (1937). To date 205 publications on the gen- 
us have been reviewed and the list of contributors to our 
knowledge of the genus embraces 585 persons. One hundred and 
forty-nine species and varieties are accepted as valid (in- 
cluding 4 doubtful species) and 246 names have been reduced 
to synonymy. A complete alphabetic list of the lattsr will 
be published in the next installment of these notes, with 
the disposition which has been made of each for ready refer- 
ence. The types or original collections on which 140 of the 


290 PHYTOLOGIA No. 8 


accepted names and 207 of the rejected names were based, 
have been examined in the course of this study. In all, 4766 
herbarium specimens of the genus, in 51 herbaria, represent- 
ing 145 accepted species and varieties, and 2708 mounted 
photographs and illustrations, in 18 herbaria, representing 
138 accepted species and varieties, have been examined and 
annotated. In number of specimens the largest collection of 
the genus is at Berlin, where there are 529 specimens at the 
present time; the second largest collection is at New York 
(518 specimens), followed by Kew (467), Paris (450), United 
States National Herbarium (404), and the Conservatoire Bot- 
anique at Geneva (321). In number of species and varieties 
represented the best collection is at New York (115), follow 
ed by the United States National Herbarium (90), Kew (85), 
Berlin (82), Conservatoire Botanique at Geneva (68), and 
Paris (66). 


40a, AEGIPHILA ACULEIFERA Moldenke, 
Additional citations: COSTA RICA: San José: H. Pittier 


7584 (E--photo, N--2 photos, S--photo, Z--2 photos). 


3a. AEGIPHILA ALBA Moldenke. 
Additional citations: ECUADOR: Los Rfos: Mexia 6656 (N-- 
photo of type, Z--photo of type). 


2la. AEGIPHILA AUSTRALIS Moldenke. = 
Additional citations: BRAZIL: Santa Catharina: Ule 152 
(N--photo of type, Z--photo of type). 


3. AEGIPHILA BOGOTENSIS (Spreng.) Moldenke. 

This binomial is incorrectly accredited to "(H.B.K.) 
Moldenke" by me in Phytologia 1: 188 (1937). 

Additional citations: COLOMBIA: Santander Norte: Schlim 


306 (Br). 


120. AEGIPHILA BRACTEOLOSA Moldenke. 
Additional citations: BRAZIL: Amazonas: Ducke 444 (N, Ww). 


98. AEGIPHILA CHRYSANTHA Hayek. 

Klug describes this species as a liana and collected it 
in anthesis in March and April, in forests at an altitude 
of 200 m. in Loreto, Peru. 

Additional citations: PERU: Loreto: Klug 2027 (S). 


10. AEGIPHILA COSTARICENSIS Moldenke. 

This binomial was erroneously cited by the Gray Herbari- 
um Card Index of New Species and Varieties, issue 142, to 
"ann. Mo. Bot. Gard. 33: 119. 1933", but this was later 
rectified on a correction card. 


eee ee eee 


—__ 


ee 


eee oe ee a 


— ee ee 


1938 Moldenke, Notes on Aegiphila 291 


A new synonym is Clerodendron Matudae Standl., Field Mus. 
Publ. Bot. 17: 206--207 (1937), of which the type was col- 
lected on Mt. Orando, Chiapas, Mexico, by Eizi Matuda (No. 
572) on December 16, 1936, deposited in the herbarium of the 
University of Michigan. This constitutes a notable extension 
in the range of the species. 

Additional citations: MEXICO: Chiapas: Matuda 572 (Mi—2, 
N--fragment & photo, S--photo, Z--photo). 


117. AEGIPHILA DEPPEANA Steud. 

The species, according to Dugand and Mina, is called 
"saiico de monte" in Colombia, where it has been collected at 
an altitude of 100--200 m. The flowers are described as 
"small, tubular, reddish". The Dugand & Mina collection has 
been erroneously distributed as A. mollis. 

Additional citations: COLOMBIA: Magdalena: Bertero s.fi. 
(N--photo, Z--photo); Department undetermined: Dugand & Mina 
950 (N). CULTIVATED: England: P. Miller 8 [Herb. Linnaeus 
G.810, S.9] (N--photo). 


124. AEGIPHILA ELATA Sw. 

The Moritz 973 and 1478, cited by me in Phytologia 1: 199 
(1937) as from an undetermined state in Venezuela, were act- 
ually collected in the State of Trujillo. The Kegel 173 and 
"Hostmann H.L.B. fee cited by Pulle in his Enum. 
Vasc. Pl. Surin. 403 (1906) are actually A. laevis. 

Additional citations: JAMAICA: P. Browne s.n. [Herb. Lin- 
naeus G.810, $.8] (N--photo). MEXICO: Veracruz: Ll. Williams 
9143 (N), 9271 (N), 9566 (N). 


61. AEGIPHILA FALCATA Donn. Sm. 
Additional citations: MEXICO: Chiapas: Matuda 666 (N). 


31. AEGIPHILA FERRUGINEA Hayek & Spruce. 

The species has been collected by Rimbach in forests, 
blooming in September. 

Additional citatione: ECUADOR: Chimborazo: Rimbach 616 
{[Mus. Yale School of Forestry 31,994] (N-~-2, S). 


63. AEGIPHILA FILIPES Mart. & Schau. 

Krukoff, on the label of the collection cited below, de- 
scribes the species as a shrub 18 feet tall, with a stem 
one inch in diameter. 

Additional citations: BRAZIL: Amazonas: Krukoff S042 (N). 


62. AEGIPHILA GLANDULIFERA Moldenke. 

Klug has collected the species in forests, at an altitude 
of 220 meters, in full anthesis in April. 

Additional citations: PERU: Loreto: Klug 3016 (S). 


292 PRET POG fk No. & 


62b. AEGIPHILA GLANDULIFERA var. PYRAMIDATA L. C. Rich. & 

Moldenke. 

The Moritz 364, cited by me in Phytologia 1: 204 (1937) 
as from an undesignated State of Venezuela, was actually 
collected in the State of Monagas. 

Additional citations: BRAZIL: Pard: L. C. Richard s.n. 
(E--photo of type, S--photo of type, W-photo of type). 


19. AEGIPHILA GRAVEOLENS Mart. & Schau. 
Additional citations: BRAZIL: S&o Paulo: Lund 796 [Mac- 
bride photos 7880] (N--photo of type, Z--photo of type). 


115a. AEGIPHILA HOEHNEI Moldenke. 

Additional citations: BRAZIL: Amazonas: Kuhlmann, Com. 
Rondon 2277 [Herb. Inst. Biol. S. Paulo 33,425] (E--photo of 
type, N--photo of type, S--photo of type, W--photo of type, 
Z--photo of type). 


42, AEGIPHILA INTEGRIFOLIA (Jacq.) Jacks. 

The Mexia 4203 and 4500 cited by me as this species in 
Brittonia 1: 339 (1934) and Phytologia 1: 228 (1937) seem to 
represent A. Sellowiana instead. 

Additional citations: BRITISH GUIANA: M. R. Schombur 
404, in part (Br). PERU: San Martin: Klug 3468 (S). 


44. AEGIPHILA INTERMEDIA Moldenke. 
Additional citations: BRAZIL: Amazonas: Ducke 136 (W). 


82. AEGIPHILA KILLIPII Moldenke. 
This specific name already occurs in some herbaria mis— 


spelled "Killippii". 


53. AEGIPHILA LAETA H.B.K. 

The Saer 602, cited by me in Phytologia 1: 229 (1937) as 
from an undetermined State of Venezuela, is actually from 
the State of Lard, and the Mocquerys 1018, cited by me on 
the same pags, was actually collected in the State of Zulia. 

Additional citations: COLOMBIA: Caldas: Bonpland 1664 (E 
-—-photo of type, S--photo of type, W--photo of type). 


78. AEGIPHILA LAEVIS (Aubl.) Gmel. 

The species has been described as a shrub or tree 6--20 
feet tall in the forests of Surinam. It is often confused 
with A, elata. The Kegel 173 and 687 and "Hostmann H.L.B. 


903 ,322--349" cited by Pulle in his Enum. Vasc. Pl. Surin. 

3 (1906) as A. elata are actually A. laevis. An additional 
synonym is Aegiphile laevis f. angustifolia Wullschl., in 
herb. 


1938 Moldenke, Notes on Aegiphila 293 


Additional citations: SURINAM: Kegel 173 (Gt), 687 (Gt); 
Wullschltigel 405 (Gt), 1983 (Gt). 


76. AEGIPHILA LAXIFLORA Benth. 

The Otto 1092 cited by me in Brittonia 1: 391 (1934) and 
in Phytologia 1: 231 (1937) as from an undetermined State of 
Venezuela, was actually collected in the State of Bolfvar. 


67. AEGIPHILA LONGIFOLIA Turcz. 
Additional citations: COLOMBIA: Santander Sur: Schlim 688 
( Br--isoty pe). 


71. AEGIPHILA MARTINICENSIS Jacq. 

The Otero 308, cited below, definitely represents the 
"“variabilis" form of the species -- almost all of the sever- 
al hundred calyxes on the specimen are deeply lobed!’ 

Additional citations: PORTO RICO: Otero 199 (Kr), 508 
(Kr). DOMINICA: G. P. Cooper 60 (N), 167 (N). MARTINIQUE: 
Stehlé 2127 (N). 


83. AEGIPHILA MOLLIS H.B.K. 

Elias 1458, erroneously distributed under this name, is 
(at least insofar as the New York specimen is concerned) a 
species of Phoradendron, while Dugand & Mina 950, also dist- 
ributed as A. mollis, is actually A. Deppeana. 

Additional citations: COLOMBIA: Cundinamarca: Bonpland 6. 
n. (E--photo, S--photo, W--photo). VENEZUELA: Amazonas: Bon- 
pland 9835 (E--photo, S--photo, W--photo). BOLIVIA: Santa 
Cruz: Steinbach 14,781 (S). 


15. AEGIPHILA MONSTROSA Moldenke. 
Additional citations: MEXICO: Oaxaca: Ll. Williams 9120 
(N). 


7a. AEGIPHILA MORTONI Moldenke. 
Additional citations: PERU: Ouzco: Cook & Gilbert 1234 (N 
--photo of type, Z--photo of type). 


69. AEGIPHILA PANAMENSIS Moldenke. 

Hayes describes the species as a shrub 12--15 feet tall, 
with light sulphur-colored flowers. 

Additional citations: COSTA RICA: Alajuela: Brenes 4030 
(N), 4345b (N); San José: Skutch 2680 (3). PANAMA: Canal 
ha Hayes s.n. (Panama, Dec. 7, 1862; Natal Herb. 10,208] 
Na). 


34. AEGIPHILA PARAGUARIENSIS Brigq. 
Additional citations: BRAZIL: S&o Paulo: Carvalho s.n. 
(Herb. Inst. Biol. S. Paulo 748] (W). 


294 | PHY TO b.06 LA No. 8 


64. AEGIPHILA PERUVIANA Turcz. 

Klug has collected the species at an altitude of 1100-- 
1600 meters in San Martin, Peru. 

Additional citations: PERU: San Martin: Klug 3511 (S). 


60. AEGIPHILA PLICATA Urb. 

The isotype of A. plicata, cited below, from the De Can- 
dolle Herbarium, was mounted on the same sheet there with an 
isotype of A. trifida Sw. 

Additional citations: JAMAICA: Bertero 2105 (N--photo of 
isotype, Z--photo of isotype). 


91. AEGIPHILA QUINDUENSIS (H.B.K.) Moldenke. 

This binomial appears mis-spelled "Aegiphita quinduensis" 
in Fedde, Repert. 42: 248 (1937). The reference given by me 
in Brittonia 1: 415 (1934) for Petitia tenuifolia should 
read "Willd. ex Schult. in Roem. & Schult. Syst. Veg. Mant. 
3: 50. 1827 [not P. tenuifolia Willd. ex Walp. Repert. 4: 
72, in syn. 1845)" and the reference for Petitia quinduensis 
should be dated "1818". 


44a. AEGIPHILA SALTICOLA Moldenke. 
Additional citations: BRAZIL: Pard: Mexia 5922 (S-- 
isotype). 


63a. AEGIPHILA SCANDENS Moldenke. 
Additional citations: BRAZIL: Acre Territory: Ule 9/2l 
(E--photo of type, S--photo of type, W--photo of type). 


108a. AEGIPHILA SCHIMPFFII Moldenke. 

Additional citations: ECUADOR: Province undetermined: 
Schimpff 1003 (E--photo of type, N--photos of type & of iso- 
type, Z--photos of type & of isotype). 


59. AEGIPHILA SELLOWIANA Cham. 

Mexia 4205 and 4500, cited by me under A. integrifolia in 
Brittonia 1: 339 (1934) and Phytologia 1: 228 (1937), seem, 
rather, to represent A. Sellowiana and they are therefore 
now cited here below. 

Additional citations: BRAZIL: Minas Geraes: Mexia 4203 
(B, Bm, Cb, D, E, G, I, N, P, S, W), 4500 (A, B, Bm, Cb, D, 
e¢, 1,-kr, WN. P, S, ¥). 


94. AEGIPHILA SMITHII Moldenke. 

Miss Mexia describes the species as a common woody vine, 
climbing up medium-sized trees, blooming in February, with 
white flowers, inhabiting cut-over woods at an altitude of 
about 110 m. 

Additional citations: PERU: Loreto: Mexia 6499 (S). 


1938 Moldenke, Notes on Aegiphile 295 


14. AEGIPHILA SORDIDA Moldenke. 
Additional citations: PERU: Loreto: Killip & Smith 27,193 


(W). 


123. AEGIPHILA SUFFLAVA Moldenke. 

Klug collected the species in a forest at an altitude of 
200 meters, blooming in March and April, and describes it as 
a liana with yellow flowers. 
‘oye citations: PERU: Loreto: Klug 2076 (S), 2574 

N). 


123ea. AEGIPHILA SUFFLAVA var. KLUGII Moldenke, var. nov. 

Haec varietds a forma typica speciei recedit per omnes 
partes densiore strigoso-pubescentibus (pilis elongatis val- 
de adpressis sufflavis) et calicibus usque ad 7 mm. longis 
et 5.5 mm. latis extus glandulis rotundatis ornatis. 

This variety differs from the typical form of the species 
in being more densely strigose-pulescent throughout, with 
longer and decidedly appressed yellowish hairs, and having 
larger calyxes (to 7 mm. long and 5.5 mm. wide), which are 
marked with several rounded glands near the apex outside, 
often a gland on every lobe. 

The type of this variety was collected by Guillermo Klug 
(No. 2511) in a forest, altitude 100 meters, at iiishuyacu, 
near Iquitos, Loreto, Peru, in February, 1932, deposited in 
the Britton Herbarium of the New York Botanical Garden, and 
is named in his honor. It is described as a liana. 

Citations: PERU: Loreto: Klug 2511 (N--type). 


56. AEGIPHILA TRIFIDA Sw. 

The isotype cited below, from the De Candolle Herbarium, 
was mounted on the same sheet there with an isotype of A. 
plicata Urb. 

Additional citations: JAMAICA: Swartz s.n. [Jamaica] (N-- 
photo of isotype, Z--photo of isotype). 


47. AEGIPHILA VERTICILLATA Vell. 

Additional citations: BRAZIL: Minas Geraes: Riedel 313 (L 
--3); Schwacke 9941 (Cb), 13,726 (Cb); Warming s.n. [Lagoa 
Sante] (Cp--3); Widgren 1222 (Br), s.n. [Minas Geraes] (Br, 
S); So Paulo: Burchell 4068 (K), 4105 (K); Campos Novaes 

(Vu); Gehrt s.n. [Herb. Inrt. Biol. S. Paulo 5489] (N); 
Glaziou 8184 (Op, P, Z--photo); Guillemin 515 (Z--photo); 
Lund 820 (Bm, Cp--2, Dc), sen. [Taubaté, November '353] (Dc); 
Martius 480 [Herb. Monac. 1045] (Ma), 8.n. (Herb. Monac. 
1046 & 1047] (Mu--2); Riedel & Luschnath 1453 (L--2); Saint- 
Hilaire GC’. 1016 (P--3); Schwacke 6594 (Cb); Sellow 2267 (B, 
Z—-2 photos), 5122 (Be~2, N—2 .photos, Z--2 photos); Usteri 
20 (B), 21 (B); Parané: Dusén 7236 (E, W), 14,871 (W), 


206 PHYTOLOGIA No. 8 


15,982 (B, E, N--2 photos, Z--2 photos), s.n. [Jaguariahyva, 
May 1914] (B, Cb, G, K); J&nsson 405a (A--photo, B--photo, D 
—photo, F--photo, K--photo, N—photo, P--photo, S=--photo, U 
--photo, V--photo, W--photo, X--photo, Z--photo); State un- 

determined: Raben 876 (Br); Sellow s.n. [Brasilia] (B, Br, K 
—3). PARAGUAY: Hassler 11,260 (Z--photo). 


45. AEGIPHILA VILLOSA (Aubl.) Gmel. 

An additional synonym is Manabea tomentosa Perrottet, in 
herb. The generic name Menabea is sometimes mis-spelled 
"Monobea", On page 343 of my monograph I stated that the 
lower surface of the leaf-blades is tomentose. This is an 
error of terminology. Actually the lower leaf-surfaces are 
villous with long straight hairs. The type specimen was col- 
lected near Cayenne, French Guiana, and is deposited in the 
herbarium of the British Museum (Natural History) in London. 

Additional citations: BRITISH GUIANA: Jenman 4068 (N, N-- 
photo, U, Z--2 photos), 6689 (U), 6869 (K, Z--photo). FRENCH 
GUIANA: Aublet s.n. [Cayenne] (Bm—-type); Benoist 1306 (P-- 
3); Collector undesignated 354 (Us), 359 (Us); Leblond 269 
(P); Leprieur s.n. [Guiana frangaise, 1838] (Le, P--2), s.n. 
[Guyane francaise, 1840] (Cb); J. Martin e.n. [Cayenne] (Bm, | 
P--2); Mélinon 128 (K, P), 145 (Cb, N--photo, P, Z--photo), 
372 (B~—4, Bm, K, N--2 photos, P, Z--3 photos), 439 (P); 
Perrottet s.n. [1820] (Cb--3); Poiteau s.n. (Dc, P, Z--2 
photos); L. CG. Richard s.n. [Guian. fr.] (N--photo, P--2, Z 
--photo); Sagot 472 (A--photo, B--photo, Bm, Cb--2, D-- 
photo, F--photo, G--photo, K, N=--photo, P--3, Ut, V, #- 
photo, Z--photo), s.n. [Cayenne, Mars 1859] (P); Von Rohr s. 
n. [Cayeme] (Bm); Wachenheim 175 (P). ILLUSTRATIONS: Aubl. 
Hist. Pl. Guian. 1: 61. 1775 (P); Lemarck, Illustr. pl. 70, 
ea2s 4791: (3). 


111. AEGIPHILA VILLOSISSIMA Moldenke. 

Additional citations: BRAZIL: Mattogrosso: Krukoff 1400 
(N--photo of type & of 2 isotypes, W#-photo of type, Z-- 
photo of type & of 2 isotypes). 


96. AEGIPHILA VITELLINIFLORA Klotzsch. 

Klug describes the species as a liana, with yellow flow 
ers, and collected it in forests at an altitude of 200 met- 
ers, blooming from March to July. An additional synonym is 
Aegiphila compacta Mart., in herb. The species is said to 
also inhabit thickets. It has been confused by some with A. 
filipes; Bowie & Cunningham confused it with A. foetida and 
Zuccarini confused it with A. martinicensis’ The type speci- 
men of A. cuspidata Mart. is deposited in the Munich herbar- 
ium. The "Hoehne s.n. [Inst. Biol. 20,562]" cited by me on 
page 4735 of my monograph is actually A. lanceolata; its be- 


1938 Moldenks, Notes on Aegiphila 297 


ing referred to A. vitelliniflora was through a typographic 
error. The "Instituto Biologico de Defesa Agricola e Animal, 
SHo Paulo" citations listed on page 474 will hereafter for 
convenience be referred to as "Herb. Inst. Biol. S. Paulo" 
The Nos. 20,099 and 20,562 cited in this list as A. vitell- 
iniflora are actually A. lanceolata. On page 423 the "Cult. 
Hort. Berol. s.n. (B-type, B-—isotype, B--cotype?)" should 
read as follows: we: "Cult. Hort. Berol. s.n. [Jun. 1831] (B); 
Lystler s.n. (Cult. Hort. Berol. Sept. 1839] (B-type & iso- 
type)". 


Additional citations: PERU: Loreto: Klug 2104 (A, B, E, 

G, K, Mi, S, W), 2204 (B, Ob, E, G, K, S, W). BRAZIL: Rio 
Grande do Norte: Schott 4917 (V--2); Pernambuco: G. Gardner 

aon. Say (Bm, Z--photo); Pickel 1084 (B, I); Bahia: Blan- 
chet 163 (Cb), 1294 (cb), 3269 (Bm, Br, Cb--3, F, P, V, X, Z 
--photo); Luschnath s.n. (Herb. Martius 1041; Herb. Monac. 
1014] (A--photo, Br, Dc, F--photo, G--photo, K, Le, Mu, N--2 
photos, Z--2 photos); Minas Geraes: Clausen s.n. (Cb); Warm 
ing 291 (Cp), a.n. [Lagoa Santa] (Bm, Cp--2, F, P); Matto—_ 
grosso: Gaudichaud 60 (P); Rio de Janeiro: Barboza s.n. 
[1846] (Br--2); Burchell 2625 (K, Le); susan 107 (Cb, 
De, P); Glaziou 807 (Br--2, C; Cp, P, Z—photo), Ch-=2, 
Cp, K, N--photo, P--2, Z--photo); Miers s.n. [M ote), BEB to Frei- 
chal, 1 May 1838] (Bm); Rudio s.n. (N--photo, Z-~photo); 
Schtich s.n. [Sebastianopolis] (A--photo, B--photo, D--photo, 
F--photo, G--photo, Mu, ibaa S--photo, V--3, W--photo, 
Z—-2 photos); Sellow (B, N--photo, Z--photo), 1020 (B), 

gen. (Z--2 photos); potter 15 (Cb); Weddell 578 (Cb, P); 
S&o Paulo: Riedel & Lund 0,39 (L); Rio Grande do Sul: Blan- 
chet 904 [Herb. lionac. ¢. 1015 a Br, Ob, wily State undet- 
Teen Bowie & Cunningham 336 (Bm--fruit), 496 (Bm); Herb. 

Imp. Vien. 152 (K); Mendonga 477 (B); Raben s.n. (Cp); 
Swainson s.n. (K). BOLIVIA: Santa Cruz: D'Orbigny 509 (Z-- 
photo), 554 esoneiols CULTIVATED: Germany: Collector un- 
designated | gen. [H.B. 30] (B); Cult. Hort. Berol. s.n. [Jun. 
1831} (B, N--photo, Z--2 photos); Herb. Zuccarini aon. 
(Herb. Monac. 1016] (Mu); Lystler a.n. [Cult. Hort. . Berol. 
Sept. 1839] (A--photo of isotype, B--photo of isotype, D—- 
photo of isotype, F--photo of isotype, N--photo of isotype, 
S--photo of isotype, W--photo of isotype, Z--photos of type 
& of isotype). LOCALITY OF COLLECTION UNDESIGNATED: Herb. 
Caes. Petrop. sen. (Herb. Martius] (Br). ILLUSTRATIONS: 
Martin f. 1, Il, & III (B); Martius, Fl. Bras. 9, tab. (B). 


The geographic distribution of the species and varieties of 
Aegiphilas 


OUBA: A. aurea [Havana] -- A. slata [Pinar del Rfo, Havana, 
Senta Clara, & Oriente] — A. martinicensis. 


298 PHY 7.0 062A No. 8 


CAYMAN ISLANDS: A. caymanensis — ~ Ae elata -- A. martinicen- 


sis. 
JAMAICA: A. elata -- A. foetida — A. martinicensis -—- A. 
martinicensis var. ~oligoneura = Se nervosa -- A. obtusa 


-- A. plicata -- A. punctata -- A. Swartziana -- - A. trif- 
ida — m~ Ae uniflora. 

HISPANIOLA: . A. elata [Hafti & Dominican Republic] — A. ner- 
~vosa [Hafti]. 

PORTO RICO, MARTINIQUE: A. elata — A. martinicensis = ie 
martinicensis var. oligoneura. 

ST. THOMAS, ST. JOHN, ST. OROIX, ST. EUSTACHE, ANTIGUA, GUA- 
DELOUPE, ST. LUCIA, GRENADA: A. martinicensis. 

ST. KITTS, MONTSERRAT, DOMINICA, ST. VINCENT: A. martinicen- 
sis -- A. martinicensis var. oligoneura. 

BARBADOS: A. barbadensis -- A. martinicensis. 

TOBAGO: A. obovata -- A. perplexa. 

TRINIDAD: A. elata —- A. integrifolia -- A. laxiflora -- A. 


macrantha — A. martinicensis -- A. obovata -- A. per- 
plexa. 


MEXICO: A. costaricensis [Chiapas] -- A. Deppeana [Tamauli- 
pas, Nayarit, Hidalgo, Veracruz, Oaxaca, & Smee ~~ he 
elata [Veracruz, Oaxaca, & Tabasco] -- A. falcata (Chiap- 
as] -- A. monstrosa [Oaxaca]. 

GUATEMALA: oe elata [Alta Verapaz & Izabal] -- A. falcata 
[Quezaltenango, Retalhuleu, Suchitepequez, & Escuintla] — 
A. fasciculata [Altea Verapaz] -- A. Hastingsiana -- A. 
laxicupulis icupulis [Chiquimula & Santa Rosa] — A. monstrosa a [El 
Petén, Alta Verapaz, & Izabal]. 

BRITISH HONDURAS: A. elata — A. monstrosa -- A. pauciflora. 

HONDURAS: A. elata [Sante Bérbara, Oortés, Yoro, & Atlantida] 
-- A. monstrosa [Cortés, Yoro, & Atl&ntida]. 

SALVALOR: be laxicupulis [Ahuachapén, Cuscatlan, & San Vicen- 


te] -- A. martinicensis var. oligoneura [La Paz]. 

NICARAGUA: mre ~ laxicupulis pustazetps | -- A. magnifica (Chin- 
andega & Chontales] —- paniculata [Segovia]. 

COSTA RICA: A. aculeifera fice. San José, & Cartago] -- 
A. anomala a {Alajuela & mers -—- A. costaricensis [Guane- 
caste & Funtarenas] -- A. Deppeana [Guanacaste & San José] 
-- A. elata (Puntarenas, | ee eae Limén, & San José] -- 
Ae falcata [Cartago] -- A. glandulifera [Alajuela] -- A. 
magnifica [Guanacaste & San José] —- A. odontophylla [Gua- 
nacaste & Heredia] -- A. panamensis [Alajuela, San José, & 
Cartago] -- A. Standleyi [San José] -- A. Valerii [Guane- 


seh . 
PANAMA: csphalophora ahaa Zone] -- A. costaricensis [Bo- 
cas ae Toro] -- A. Deppeana [Chiriquf, Veraguas, & Canal 


Zone] —- A. elata ~[Panemé] -- A. falcata [Bocas del Toro] 
-- A. filipes [Panamé] -- A. clandulifera [Canal Zone] — 
A. laeta [Taboga Island] -- - A. magnifica [Panamé] -- A. 


1938 Moldenke, Notes on Aegiphila 299 


martinicensis [Bocas del Toro] -- A. mollis [Canal Zone] 
—- Aw pananene?s [Bocas del Toro, Canal Zone, Colon, & 
= A. 


Panamaé A. paniculata [Bocas del Toro, Canal Zone, & 
Panamé] -- A. pendula [Canal Zone]. 


COLOMBIA: A. aculeifera [El Cauca] —- A. bogotensis [Santan- 
der Norte, Tolima, Cundinamarca, El Cauca, & Narifio] -- 
A. caucensis [Caldas] -- A. Deppeana [Magdalena, Bolfvar, 
& Cundinamarca] -- A. elata [Magdalena, Bolivar, El Valle, 
& Cundinamarca] -- A. filipes [Magdalena] -- A. glandu- 
lifera [Antioquia, Santander Sur, Chocd, & Tolima] -- A. 

landulifera var. pyramidata [Méta] -- A. Goudotiana 
Peers -- A. grandis [Tolima & Cundinamarca] — 
A. guianensis [Cundinamarca & Méta] -- A. hirsutissima 
[Bolfvar] -- A. integrifolia [Magdalena, Boyacaé, Méta, & 
El Cauca] -- A. Killipii [Santander Norte] -- A. laeta 
(Magdalena, Santander Sur, & Caldas] -- A. laevis [Antio- 
quia] -- A. Lehmannii [Boyacé, Chocd, & El Cauca] -- A. 
longifolia [Santander Sur] -- A. martinicensis [Antioquia 
& Méta] -- A. membranacea [Chocd & Métal] -- A. mollis 
(Magdalena, Atldéntico, Bolfvar, El Valle, Tolima, Cundin- 
amarca, & El Cauca] — A. mollis var. intermedia [Magde- 
lena & Bolfvar] -- A. montana [Huila] -—- A. novogranaten- 
sis [Antioquia & El Cauca] -- A. paniculata [Santander 
Sur] -- A. Pennellii [Tolima] -- A. puberulenta [Bolfvar] 
-- A. quinduensis [Tolima & Cundinamarca] -- A. racemosa 
[El Valle] -- A. reticulata [Cundinamarca] -- A. sessili- 
flora [Antioquia] -- A. sylvatica [Santander Sur] -- A. 
ternifolia [Cundinamarca] -- A. truncata [Cundinamarca]. 

VENEZUELA: A. elata (Trujillo, Carabobo, & Aragua] — A. 
Fendleri [Aragua] -- A. floribunda (Aragua] -- A. glandu- 
lifera var. pyramidata [Monagas] — A. guianensis [Amaz- 
ones] -- A. hirsutissima [Miranda] -- A. integrifolia 
[Zulia & Amazonas] -- A. laeta (Zulia, Lard, & Anzoateg- 
ui] -- A. laxiflora [Bolfvar] -- A. Lewisiana [Federal 
District] —- A. macrophylla -- A. martinicensis [Zulia & 
Mérida] — A. membranacea [Falcon] -- A. mollis [Lard, 
Carabobo, Aragua, Federal District, Miranda, Guarico, 
Sucre, Anzoategui, Monagas, Bolfvar, & Amazonas] -- A. 

arviflora [Bolfvar] -- A. pendula [Zulia] -- A. perplexa 
Delta Amacuro] -- A. quinduensis [Cerabobo & Federal 
District] -- A. racemosa [Zulia] -- A. ternifolia -- A. 
verrucosa [Aragua]. 

BRITISH GUIANA: A. bracteolosa -- A. elata -- A. Gleasonii 
-- A. guianensis -- A. integrifolia -- A. laevis -- A. 
laxiflora -- A. macrantha —- A. racemosa -- A. roraimen- 
sis -- A. villosa. 


SURINAM: A. elata -—- A. laevis -- A. membranacea -~ A. race- 
mosa. 


FRENCH GUIANA: A. Deppeana -- A. elata — A. glandulifera 


500 PHY b0:L:0 Gos No. & 


var. pyramidata —- Ae integrifolia -- A. laevis -- A. 


macrantha -- A. martinicensis -- Ae membrenacea -- xe Ta- 
cemosa -- A. villosa. 


ECUADOR: A. albe [Guayas & Los Rfos] -- A. bogotensis [Car- 
chi, Pichincha, & Los Rfos] — A. icsiatin Manabi & 
Guayae] -- A. ferruginea [Carchi, Imbabura, Pichincha, & 
Chimborazo] — A. glandulifera [Napo-Pastaza] -- A. glom 
erata [Menabij -- A. integrifolia [Napo-Pastaza] —- A. 
monticola [Pichincha & Chimborazo] -- A. Pavoniana [Guay- 
ac} — A. pendula [Oriente] — A. Rimbachii — A. Schim- 
prii. 

PERU: A. bracteolosa [Loreto] -- A. chrysantha [Loreto] -- 
Ae cordata [Loreto] -- A. cordifolia [Amazonas & Huanuco] 
-- A. cuneata [Loreto] -- A. elegans [Loreto & Junfn] — 

As fi. ilipes [Loreto] -- A. glabrate [Junin] -- A. glandu- 
lifera [Loreto] -- A. insignis [Ancachs] — A. " integri- 
folia [San Martin, Loreto, Huanuco, & Junfn] -- A. longi- 
petiolate eee cal -- A. membranacea [Loreto] -- As, Mort- 
oni [Cuzco] -- A. multiflora [Huanuco & Pufio] -- A. ovate 
[Junin] —- Ae eruviana [San Martin & Loreto] -- A. ul- 
cherrima [Junin] -- A. Smithii [San Mart{n, Loreto, & Ju- 
nin] —- A. sordide [Loreto] — A. sufflava [Loreto] — A. 
sutflava var. Klugii [Loreto] -- - A. triflora [Loreto] -- 
A. velutinosa [Cuzco] -- A. vitelliniflora [Loreto]. 

BRAZIL: A. australis [Santa Catharina] -- A. brachiata [Rio 
de Janeiro, Parané, Santa Catharina, & Rio Grande do Sul] . 
-- A. bracteolosa pages & Pard] -- A. brasiliensis 
[Rio de Janeiro] — A. Candelabrum [Mattogrosso] -- A. 

capitata [Sfo Paylo] -- A. casseliaeformis [S&o Paulo] — 
A. chrysantha [Pernambuco & Bahia] —- A. conturbata [Mar- 
anhfio] -- A. cordate [Acre Territory] -- A. coriacea -- 
A. pe finas Gerees, SHo Paulo, & Parana | —- A. cun- 
eata [Acre Territory] -- A. dentata [S&o Paulo] -- we l= 
egans [Amazonas] -- A. filipes 6s (Amazonas, Parad, & Acre 
Territory] -- A. fluminensis [Bahia, Rio de Janeiro, S&o 
eee & paréadT —= A. glandulifera [Amazonas & Parg] -- 

A. glandulifera var. “paratnsis [Pard] -- A. glandulifera 

var. pyreamidata [Pard] -- A. gloriosa [Behiey -- Aw 
Goeldiana [Para] -- A. graveolens [Bahia, Rio de Janeiro, 
& So Paulo] -- Ae uianensis [Paré] -- A. Hassleri [Par- 
ané] — A. Hoehnei [Amazonas] -- Ae integrifolia folia [Amazon- | 
as, Pard, Bahia, M Minas Geraes, Goyaz, fattogrosso, Rio de 

Janeiro, & So Paulo] -- A. intermedia [Anmazonas, Pard, & 
Marenhfio] —- A. laevis [Bahia, Mattogrosso, & Rio de Jan- | 
eiroj] -- A. lanata [Goyaz] — A. lanceolata [Pard, Minas : 
Geraes, Espirito Santo, Rio de ~Jamiro, & SKo Paulo] -- 
A. Lhotzkiana [Cearé, Parahybe, ee Geraes, Goy= 
az, Espirito Santo, Sho Paulo, & Parand] -——- A. Luschnathi 
[Bahia, Minas Geraes, Rio de Janeiro, Sfo Paulo, & Santa 


~~ —" ee eee 


1938 Moldenke, Notes on Aegiphila 301 


Catharina] -- A. macrantha [Paré] -- A. mediterranea [Rio 
de Janeiro & SHo Paulo] -- A. medullose [Rio de Janeiro] 
-- A. novofriburgensis [Rio de Janeiro] -- A. obducta 
[Amazones, Minas Geraes, Rio de Janeiro, So Paulo, Para- 
né, & Santa Catharina] -- A. paraguariensis [Minas Gere- 
es, Goyaz, Mattogrosso, Rio de Janeiro, Sho Paulo, & Par- 
ané] -- A. parviflora [Paré] -~ A. pernambucensis [Per- 
nambuco & Rio de Janeiro] -- A. racemosa [Amazonas, Pard, 
Pernambuco, Bahia, & Rio de Janeiro] -- A. Riedeliana 
[Bahiea, Minas Geraes, Rio de Janeiro, S&o Paulo, Sante 
Catharina, & Rio Grande do Sul] -- A. salticola [Parg] -- 
A. scandens [Acre Territory] -- A. Sellowians [Pard, Per- 
nambuco, Bahia, Minas Geraes, Goyaz, Mattogrosso, Espiri- 
to Santo, Rio de Janeiro, S&o Paulo, Parand, & Rio Grande 
do Sul] -- A. splendens [Minas Geraes] —- A. Spruceana 
[Amazonas] -- A. Surfaceana [Amazonas & Para] -- A. vert- 
icillata [Minas Geraes, Mattogrosso, Rio de Janeiro, Sfo 
Paulo, & Parandé] -- A. villosissima [Mattogrosso] — A. 
virgata [Rio de Janeiro] -- A. vitelliniflora [Pard, Rio 
Grande do Norte, Pernambuco, Bahia, Ninas Geraes, Matto- 
grosso, Rio de Janeiro, So Paulo, & Rio Grande do Sul]. 
BOLIVIA: A. boliviana [Sante Cruz] -- A. breviflora [El Be- 
ni] -- A. Buchtienii [La Paz] — A. chrysantha [Santa 
Cruz] -- A. elata [Santa Cruz] -- A. elegans [La Paz] -- 
A. elongata [La Paz] -- A. filipes [El Beni] -- A. Herz- 
ogii [Santa Cruz] —- A. hirsuta [La Paz] — A, integrifo- 
lia [El Beni, La Paz, & Santa Cruz] -- A. mollis [Santa 
Cruz] -- A. multiflora [El Beni & Cochabamba] -- A. ovata 
[La Paz] -- A. peruviena [El Beni] -- A. Sellowiana [E1 
Beni, La Paz, & Santa Cruz] —- A. setiformis [La Paz] — 
A. spicata [La Paz] -- A. vitelliniflora [Santa Cruz]. 
PARAGUAY: A. brachiata —- A. Candelabrum — A. Hassleri — 


A. lanceolata -- A. paraguariensis -- A. platyphylla -- 
A. verticillata -- A. vitelliniflora. 

URUGUAY: A. Hassleri. 

ARGENTINA: A. Hassleri [Misiones & Corrientes]. 

CULTIVATED: A. Deppeana [England] -- A. elata [Florida, Cu- 
ba, & Austria] -- A. integrifolia [Hefti] -- A. lanceole- 
ta [Brazil] -- A. Luschnathi [France] — A. martinicensis 
[England, France, Netherlands, Belgium, Germany, & Cey- 
lon] — A. martinicensis var. oligoneura [British Guiana 
& Belgium] -- A. mediterrenea [Cameroons] -- A. obducta 
[Brazil & France] -- A. obovata [British Guiana & Eng- 
land] -- A. Sellowiana [Brazil] -- A. vitelliniflora 
(Germany ]. 


An alphabetized list of citations (additional to those pub- 
lished in Brittonia 1: 472--477. 1934), including cor- 
rections: 


3502 PHY TO: 0 G:t-s No. 8 


Ackermann s.n. [1831] (Sellowiana); Acufia s.n. [Herb. Ro- 
ig 4232] (elata); Aitken 1082 (racemosa); Alexander, D. W., 


s.n. (Trin. Bot. Gard. Herb. 5022] (perplexa); Alexander, R. 
C., sen. [Jamaica, 1850] (foetida), s.n. [Moneague] (elata), 
a.n. [Pleasant Valley, Moneague, 10 Apr. 1850] (foetida), s. 
n. (elata); Allen, C., 416 (laeta); André K.1554 (mollis), 
K.1556 (mollis); Appun 1204 (roraimensis, type coll.); 
Archer 392 (sessiliflora, type coll.); Arechavaleta 43 (Has- 
sleri), B (Hassleri), s.n. (Hassleri); Aublet s.n. [Cayenne] 
(villosa, type coll.), s.n. (integrifolia, type coll. of 
Manabea arborescens), s.n. (laevis, type coll.); Aviles 988 
(cephalo hora). 
Badier 176 (martinicensis); Bailey & Bailey 575 (lenceo- 
lata), 662 (cephalophora); Baker, O. F., 204 (magnifica, 
type coll.); Balansa 2094 (lanceolata); Bang, M., 671 (in- 
tegrifolia), 878a, in part (setiformis, type coll.), 878a, 
in part (spicata, type coll.), 1732 (setiformis, type coll. 
of densiflora); Barboza s.n. [1846] (Luschnathi), s.n. 
[1846] (vitelliniflora); Barclay 632 (glomerata), 2498 (lae- 
ta); Bartlett, H. H., 11,941 (monstrosa), 13,011 (monstro- 
sa); Baudin s.n. (martinicensis); Bélanger 260 (martinicer- 
Be 290 (martinicensis), 5/1 (martinicensis); Benoist 740 
membranacea), 786 (membranacea), 78/7 (me mbranacea), 1306 
(villosa); Benzon 127 (martinicensis), 3713 (martinicensis); 
Bernoulli 748 (laxicupulis); Berro 1019 (Hassleri), 5087 
(Hassleri); Bertero 55, in part (elata), 2046 (martinicen- 
sis), 2104 (elata), 2105 (plicata, type coll.), s.n. [S. D.3 
Herb. Monac. 1019] (elata), s.n. [Majo 1831] (plicata), 
a.n. (Deppeana, type coll. of Berteriana), s.n. (elata); 
Berthoud-Coulon 196 (laevis), 550 (racemosa); Beyrich s.n. 
(Luschnathi); Biolley 7409 (falcata); Blanchet 46 (Lhotzki- 
ana), 79 (Lhotzkiana), 163 (vitelliniflora), 231 (graveo- 
lens), 354 (Lhotzkiana), 682 (fluminensis), 729 (Lhotzkia- 
na), 862 (laevis), 904 (vitelliniflora), 1064 (Lhotzkiana), 
1294 (vitelliniflora), 1398 (gloriosa, ee oases 
(laevis), 1605 (fluminensis), 1998 (gloriosa), 20 Lhotz- 
kiana), 3397 (Lhotzkiana), 3451 (graveolens), s.n. [1834] 
(fluminensis), 8.n. [1857] (Lhotzkiana), s.n. (Lhotzkiana), 
a.n. (racemosa), s.n. (Sellowiana); Boldingh 189 (martini- 
censis), 2 martinicensis), 386a (martinicensis), 410 


martinicensis), 8 martinicensis ); Bonpland 755 (Hass- 
leri), 9 inte rifolia), 983 (mollis, type coll. of salu- 
teria), 1664 (laeta, type coll.), 5864 (quinduensis), sen. 
(mollis); Boog s.n. (mediterranea); Bowie & Cunningham 336 
(vitelliniflora), 496 (vitelliniflora), s.n. [Road to Sta. 
Amaro, near Sta. Paulo] (obducta), s.n. (obducta); Brade 
7464 (Sellowiana), 10,542 (obducta); Brenes 105 (panamens- 
is), 105a (panamensis), 110 (panamensis), 180 (panamensis), 
301 (aculeifera), 3574 (panamensis), 4030 (panamensis), 


ct tate tn 


1938 Moldenke, Notes on Aegiphila 503 


4320 (panamensis), 4345b (panamensis), 4395 (panamensis), 
5709 (aculeifera), 6193 (elata), 6652 (anomala), 14,313 


(panamensis), 15,661 (anomala), s.n. [Herb. Instit. Physico- 
geogr. Nat. Costaric. 14, (glandulifera); British Guiana 
Bot. Gard. s.n. [May, 1905] (obovata); Britton & Cowell 594 
(martinicensis); Britton & Harris 10,726 (elata); Broedway, 
W. E., 142 (martinicensis), 1410 (martinicensis), 2584 (obo- 
vata), 2973 (obovata), 3157 (obovate), 3334 (elata), 4243 
(perplexa), 6349 (obovata), 6396 (obovata), 9064 (obovata), 
9128 (obovata), s.n. [Mount Parnassus, June, 1906] (martini- 
censis), s.n. [Oct. 31, 1905; Hort. Thenensis 1.4507] (mart- 
inicensis), s.n. [Year 1906?] (martinicensis), s.n. [Manzan- 
illa, Aug. 1918] (inteprifolia), s.n. [Trin. Bot. Gard. 
Herb. 7164] (integrifolia); Broadway, Mrs. ¥. E., s.n. [St. 
Ann's) (laxiflora, type coll. of trinitensis); Browne, P., 
s.n. [Herb. Linnaeus G.810, S.8] (elata, type coll. of 
Knoxia 2); "B.S." s.n. (martinicensis); Buchtien 1645 (el- 
ongata, type coll.), 1715 (hirsuta, type coll.), 1717 (Buch- 
tienii, type coll.), 2992 (multiflora), s.n. [XI.1910] (mul- 
tiflora); Bunbury 297 (obducta), 463 (obducta); Burchell 9 
(pernambucensis), 1985 (Riedeliana), 2581 (integrifolia), 
shi8. in part (integrifolia), 3418, in part (Sellowiana), 
3547 (capitata, type coll.), 4105 (verticillata), 4604 (Sel- 
lowiana), 8345 (paraguariensis), 9645 (lanceolata); Buswell 
s.n. (Gov't. Grounds, July 22, 1934] (elata). 

Caley a.n. [Jan. 1823] (martinicensis ); Campos Novaes 933 
(verticillata); Cardenas 16, special (breviflora, type 
coll.); Carleton 422 (monstrosa, type coll.); Carvalho s.n. 
(Herb. Inst. Biol. 748] (parazuariensis); Casaretto 926 (ob- 
ducta), 2022 (Lhotzkiana), 2624 (verticillata); Castelnau s. 
n. [Rio Ucayali] (integrifolia); Chaffanjon 245 (mollis); 
Chamberlain s.n. [25 Juli 1817; Herb. Monac. 1023] (flumin- 
ensis); Chrysler 1636 (elata); Clausen 8 (obducta), 53, in 
part (Luschnathi), 134 (novofriburgensis, type coll.), 352 
(Lhotzkiana), 630 (lanceolata), 652 (obducta), s.n. [Curvel- 
lo] (Lhotzkiena), s.n. [Aug.--April, 1840] (Lhotzkiana), s. 
n. [Aug.—-April, 1840] (obducta), s.n. [Aug.--April, 1840] 
(Sellowiana), s.n. [Aug.--April, 1840] (verticilletea), s.n. 
(Lhotzkiana), s.n. (vitelliniflora); Collector indig. Surin- 


ame 62 (laevis), 87 (racemosa); Collector undesignated 23 
(obducta), 285 (fluminensis), 354 (villosa), 359 (villosa), 


404 (paraguariensis), 426 (martinicensis), 3653 (Sellowiana 
on, co Portenschlag] (elata), s.n. [Cayerme] (laevis), 8. 
n. [Aug. 1824] (martinicensis), s.n. {"Bois de Calme"] (ner- 


vosa), 8.n. (H. B. 30] (vitelliniflora), s.n. (fluminensis), 
S.n. (integrifolia), s.n. (laevis), 8.n. (martinicensis), s. 
n. (racemosa); Cook & Collins 291 (martinicensis); Cook & 
Gilbert 1234 (Mortoni, type coll.), 1382 (velutinosa, type 
coll.); Cooper, G. P., 60 (martinicensis), 80 (martinicen- 


304 PHYTOLOGI-A No. 8 


sis), 167 (martinicensis); Crliger s.n. [17 Aug. '46] (integ- 
rifolia), sen. (laxiflora); Cult. Hort. Berol. s.n. (Jun. 
1831] (vitelliniflora); Cult. Hort. Schtnb. s.n. (elata); 
Cult. Palm Stove Kew 8.n. (martinicensis). 

Dahlgren & Sella 524 (racemosa); Damazio 1786 (obducta), 
1926 (Sellowiana); Dancer s.n. (elata); Dannouse s.n. (laxi- 


flora); Dawe 192 (bogotensis), 472 (glandulifera, type 
coll.); De la Ossa s.n. [1825] (martinicensis); Department 
of Forestry 2850 (racemosa); Desclussaux s.n. [Herb. Jussieu 
5035a] (elata); Desportes s.n. (Herb. Jussieu 5035b & 5041) 
(elata); Diels 942 (glandulifera); Distin s.n. (foetida); 
Dombey 251 (multiflora), s.n. [P&son Huare-hueri] (multi- 
flora), s.n. (integrifolia); Don, G., 8.n. (elata); D'Orbig- 
ny 1086 (mollis), 1094 (Sellowiana); Drake s.n. (racemosa); 
Duchassaing s.n. [Panama] (panamensis); Ducke 136 (intermed~ 
ia), 444 (bracteolosa), S44 (intermedia), 6739 (integrifol- 
ia), 7351 (bracteolosa), s.n. [Herb. Rio de Janeiro 18,949] 
macrantha, type coll.); Dugand 101 (mollis), 469 (mollis), 
639 (mollis); Dugand & Mina 950 (Deppeana); Dusén 9382 (Has- 
sleri), 10,541 (crenata, type coll.), s.n. [May 5, 19117] 
crenata), s.n. [Jaguariahyva, May 1914] (verticillata); 
Duss 303 (martinicensis), 2389, in part (martinicensis). 
Edwall s.n. [Herv. Geogr. e Geol. 4362; Herb. Inst. Biol. 
15,614] (dentata, type coll.); Eggers 501, in part (martini- 
censis), 5184b (elata), 6617, in part (martinicensis), 6617, 
in part (martinicensis var. oligoneura), 71535 (martinicens- 
is), sen. [Balao, 31/12/1891] (alba), a.n. [Decbr. 1882] 
(martinicensis); Ekman H.8971 (nervosa, type coll. of subop- 
posita), H.12,310 (elata), H.13,279 (elata); Elias 617 pub- 
erulenta, type coll.), 1102 (mollis); Estebom 7 (laevis). 
Fawcitt s.n. [May, 1888] (elata); Fendler 571 (perplexa), 
592 (perplexa), 841 (verrucosa), 844 (Lewisiana, type coll.), 
845 (floribunda, type coll.), 2032 (Fendleri, type coll.); - 
Fiebrig 5201 (Candelabrum), 5631 (paraguariensis), 6241 
(lanceolata, type coll.); Finlay s.n. trin. Bot. Gard. 
Herb. 2386] (obovata), s.n. [Trin. Bot. Gard. Herb. 2389] 
(obovata, type coll. of dubia), s.n. [Trin. Bot. Gard. Herb. 


2391] (laxiflora); Fleischmann 520 (integrifolia); Focke 396 
(elata); Forsstrtm s.n. (martinicensis); Forster s.n. ["Ind- 
ia"] (martinicensis); Forsyth s.n. (martinicensis); Fox 463 


(obducta); Frambach 123 (Sellowiana); Freyreiss 8.n. (coria- 
cea, type coll.), e.n. (Luschnathi); Funck & Schlim 81 (mar- 


tinicensis). 
Gabriel s.n. [1602] (laevis); Galeotti 1268 (Deppeana), 
7238 (elata); Gardner, G., 100 (mediterranea, type coll. of 


is 


cestrifolia), 3401 (paraguariensis), 5830 (obducta, type 
coll. of lanuginosa), 8.n. [1836] (vitelliniflora); Gaudi- 
chaud 60 (vitell:niflora), 110 (Luschnathi), 265 (obducta), 


468, in part (fluminensis), 468, in part (Luschnathi). 


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: PHY TOLOGIA 


Designed to expedite botanical publication 


Vou. 1 January, 1939 No. 9 


| 
| 
| 
CONTENTS 
| 


C. L. Lunpexr: Six New Trees and Shrubs from Tropical North 
OPEL OE SO aN ke Sate bao art AL a lat Nak al 305 


Published by H. A. Gleason and Harold N. Moldenke 


The New York Botanical Garden 
Bronx Park, New York, N. Y. 


Price of this number, 50 cents; per volume, $5.00 in advance. 


Vol. 1, number 8 , was issued May 17, 1938 


SIX NEW TREES AND SHRUBS FROM TROPICAL NORTH AMERICA (a) 


C. L. Lundell 


MAYTENUS GUATEMALENSIS Lundell, ap. nov. 

Frutex 1.3-metralis, 3 om. diam. Folia subcoriacea, ob- 
longo-elliptica vel elliptica, 3.8--7 cm. longa, 1.4--3 cm. 
lata, epice acuminata, acumine obtusiusculo, basi late oune- 
eta, obscure et minute crenulato-serruleta, nervis laterali- 
bus utrinque 7--9; petiolo 1.5--%3 mm. longo. Inflorescentiae 
fasciculatae, axillares; pedicellis ca. 5 mm. longis. Capsu- 
la@ monosperma, obovoidea, ca. 1! mm. longa. 

A glabrous shrub, 1.3 m high, about 3 cm. in diameter; 
branchlets slender, wiry, angled. Leaves subcoriaceous, ob- 
long-elliptic or elliptic, 3.8 to 7 om. long, 1.4 to 3 cm. 
wide, epex sbruptly acuminate, the acumen obtusish, base 
broadly cuneate, cbscurely and minutely crenulate-serrulate, 
costa slightly elevated on both surfaces, main lateral veins 
7 to 9 on each side, visible but faint on both surfaces; 
petioles canaliculate, 1.5 to 3 mm. long. Infructescence 
fasciculate in the axils; pedicels about 5 mm. long, some- 
times solitary (7). Persistent calyx S-dentate, the teeth 
broadly deltoid, sbout 0.5 mm. long, about 1.3 mm. wide. 
Capsules obovoid, about 11 mm. Long, 2-celled with 2 erect 
ovules in each cell, 1l-seeded. 

Type in the Herbarium of the University of Michigan, W. 
A. Schipp S-635, collected in swampy forest shade, at Camp 
35, British Honduras - Guatemala boundary survey, Toledo 
district, et alt. of about 750 m., June 10, 1934. 


MAYTENUS SCHIFPII Lundell, sp. nov. 

Arbor 8--10-metralis, 13--23 cm. diam. Folia subcoriacea, 
obovato-elliptica vel elliptica, 5--11.5 om. longa, 2.3--5.4 
cm. lata, apice abrupte acuminata, acumine obtuso, basi late 
cuneata, crenulato-serruleta, nervis lateralibus utrinque 
7--11; petiolo 6--S mm. longo. Infructescentiae fascicula- 
tae, axillares; pedicellis 4--6 mm. longie. Capsule obovoid- 
ea, 9--12 mm. longa. Semina 1--3; cotyledonibus suborbicu- 
laribus, ca. 7 mm. longis, basi biappendiculatis; radicula 
parva, ca. 0.5 mm. longa. 

A glabrous tree 8 to 10 m. high, 13 to 23 cm. in diam., 
with cream-colored wood (Schipp); branchlets slender, wiry, 
slightly angled. Leaves thinly subcoriaceous, obovate-ellip- 
tic or elliptic, 5 to 11.5 cm. long, 2.3 to 5.4 cm. wide, a- 
pex abruptly acuminate, the acumen obtuse, base broadly cun- 
eate, finely crenulate-serrulate, sometimes very obscurely 
so, costa elevated on both surfaces, main lateral veins 7 to 


305 


306 PHYTOLOGIA No. 9 


11 on each side, finely prominulous on both surfaces, vein- 
lets openly reticulate; petioles canaliculate, 6 to 9 m. 
long. Infructescence fasciculate in the axils; pedicels 4 to 
6 mm. long. Persistent calyx 5-dentate. Capsules bright or- 
enge (Schipp), obovoid, 9 to 12 mm. long, 2-celled, with 2 
erect ovules in each cell, 1l- to 3-seeded, eril red. Endo- 
sperm of seed abundant; cotyledons thin, equal, suborbicu- 
lar, inequilateral, about 7 mm. long, besring a minute ap- 
pendage at base on each side; redicle small, about 0.5 mm. 
longe 

Type in the Herbarium of the University of Michigan, W. 
A. Schipp 1014, collected in shade in swampy places, at 
Dixie » Punta Gorda, Toledo District, British Honduras, 
alt. about 65 m., Sept. 26, 1922. 

Additional specimens examined: BRITISH HONDURAS: Toledo 
District, Rio Grende, alt. 15 m., in swamp forest or on 
river banks, Schipp 1224, Nov. 18, 1933. 

M. Schippii is closely related to M. guatemalensis Lund- 
ell, but differs in being a tree with larger, usually obo- 
vate-elliptic rether than oblong-elliptic leaves, and with 
petioles more than twice as long. 


MAYTENUS TEXANA Lundell, sp. nov. 

Frutex. Ramuli minute puberuli. Folia crassa, pallida, 
oblongo-elliptica vel obovato-elliptica, 1.6--3.1 cm. lon- 
ga, 9--18 mm. lata, apice late obtusa vel rotundeta, minute 
apiculata, basi rotundeta, subintegra, enervia, costa basi 
prominula; petiolo crasso, 1--43 mm. longo, parce et minute 
puberulo. Flores dioeci, fasciculati. Pedicelli 2 mm. longi 
vel minores. Calyx quinquefidus, lobis rotundatis, ca. 0.65 
mm. longis. Petala 5, ovata, 1.2--1.5 mm. longa, lineati- 
punctata. Ovarium 37~ raro 4-loculare. Cvulum in loculis sol- 
itarium. Capsula obovoidea, ca. 12 mm. longa. Semina 1--3, 
erillate. 

A shrub, much-branched; branchlets usually short, rather 
thick, very minutely puberulent. Leaves thick, very pale, 
ashy gray when dry, oblong-elliptic or obovate-elliptic, 1.6 
to 3.1 cm. long, 9 to 18 mm. wide, spex broadly obtuse or 
rounded, usually minutely apiculate, base rounded, entire 
below, subentire above, usually with 2 or 3 minute teeth on 
each side, venation obscure, costa prominulous at base only; 
petioles stout, 1 to 3 mm. long, sparsely and minutely pub- 
erulent. Flowers dioecious, fasciculate in the leaf axils; 
pedicels less than 2 mm. long. Staminate flowers: Calyx S- 
lobed, the lobes rounded, about 0.65 mm. long, very minute- 
ly erose-ciliolate, punctate. Petals 5, ovate, 1.3 te 1.5 
mm. long, erose at apex, lineate-punctate. Filaments subula- 
te, about 0.6 mm. long. Anthers broadly cordate, rufous 
punctate dorsally. Pistillate flowers: Petals smaller, sbout 


1939 Lundell, Six New Trees & Shrubs 307 


1.2 mm. long, rufous-punctate. Stamens rudimentary. Disk 
large, flat. Ovary usually 3-celled, sometimes 4-celled, 
with one erect basal ovule in each cell. Capsules obovoid, 
about 12 mm. long, 3- or 4-celled, 1l- to 3-seeded; aril red; 
seed lineate, oblong-obovoid, about 5 mm. long; endosperm 
present; cotyledons oblong-elliptic, about 2.8 mm. long, 
rounded at apex and base, flat; radicle about 0.8 mn. + ORs: 
ecutish. 

Type in the Herbarium of the University of Michigan, 
Elzada U. Clover 986, collected in mesquite woods, between 
Los Fresnos and Port Isabel, Cameron Co., Texas, April 22, 
1953. 

Additional specimens examined: TEXAS: Cameron Co., Los 
Fresnos, in heavy brush, Clover 1731, Feb. 8, 1934; common 
in the lower Rio Grande valley. 

The oblong-elliptic or obovate-elliptic, short-petiolate 
leaves rounded at the base, and the smaller rufous-punctate 
flowers distinguish M. texana from M. phyllanthoides Benth., 
its closest relative. In the latter, the leaves are obovate, 
cuneate eat the base, larger, and have much longer petioles. 


BUGENIA TOLEDINENSIS Lundell, sp. nov. 

Arbor 15 m. alta, 30 cm. diam.; ramulis ut videtur novel- 
lis rufo-tomentosis, demum glabratis. Folia 5--6 mm. longe 
petiolata, subcoriacea, adulta glabra, oblonga, 9.5--12.5 
cm. longa, 2.8--4.1 cm. lata, apice obtuse acuminata, basi 
acuta. Pedunculi et pedicelli rufo-tomentosi, crassi, 2--5 
mme longi, axillares. Fructus globosus, ca. 2 cm. diame, 
rufo-tomentosus. 

A tree 15 m. high, 30 cm. in diam.; branchlets rather 
slender, apparently rufous-tomentose at first, glabrous with 
age. Leaves thin, subcoriaceous. Petioles stout, terete, 5 
to 6 mm. long. Leaf-blades narrowly oblong, 9.5 to 12.5 om. 
long, 2.8 to 4.1 om. wide, apex obtusely acuminate, base a- 
cute, glabrescent with age, costa slightly raised above, 
prominent beneath, reticulate-veined on both surfaces, main 
lateral veins 12 to 15 on each side, widely ascending, near- 
ly horizontal to midrib, anastomosing into submarginal 
veins, conspicuous on both surfaces. Pedicels solitary or 
several in a short raceme, axillary, the peduncles and pedi- 
cels stout, 2 to 5 mm. long, rufous-tomentose. Fruits glob- 
ose, about 2 cm. in diam., rufous-tomentose, vrowned by per- 
sistent rufous-tomentose calyx-lobes. 

Type in the Herbarium of the University of Michigan, W. 
A. Schipp S-644, collected on creek bank in forest shade at 
Camp 23, British Honduras - Guatemala boundary survey, Tole- 
do District, British Honduras, at alt. of about 600 m., Feb. 
16, 1934, 

Eugenia toledinensis apparently is related to E. xalapen- 


308 PHYTOLOGIA No. 9 
sis (HBK.) DC. 


EUGENIA VACANA Lundell, sp. nov. 

Arbor 5 cm. diam.; ramulis novellis rite Aiden teen dem- 
um glabratis. Folia subcoriacea, anguste elliptico-oblonga, 
2.5--4.5 cm. longa, 1--1.7 cm. lata, apice obtuse acuminata, 
basi cuneata, juventute parce rufo-tomentosa, meturitate 
glabra; petiolis 2--43.5 mm. longis. Flores breviter racemosi, 
ut videtur fasciculati. Pedicelli graciles, 4--8 mm. longi, 
parce rufo-tomentosi. Calycis lobi inaequales, ovati, 1--1.5 
mm. longi. Fetala orbicularia, 2.8--3.4 mm. longa, ciliata. 

A tree; diam. 5 cm.; branchlets slender, reddish, at 
first loosely rufous-tomentose, puberulent or glabrescent 
with age; internodes 1 to 1.8 cm. long. Leaves subcoria- 
ceous, nerrowly elliptic-oblong, 2.5 to 4.5 cm. long, 1 to 
1.7 cm. wide, apex acuminate, the acumen obtuse, base cune- 
ate, loosely rufous-tomentose on both surfaces at first, 
glabrescent early, costa impressed above, prominent beneath, 
the veins obsolete above, faint beneath; petioles canalicu- 
late, 2 to 3.5 mm. long. Flowers white, very shortly racenm- 
ose, appearing fasciculate. Pedicels slender, 4 to 8 m. 
long, loosely rufous-tomentose. Calyx pubescent, 4-lobed, 
the lobes ciliate, ovete, unequal, 1 to 1.5 mm. long, round- 
ed or broadly obtuse. Petals 4, orbicular, 2.8 to 3.4 mm. 
long, ciliate. 

Type in the Herbarium of the University of Michigan, 
Percy H. Gentle 2535, collected on hillside opposite Vaca, 
El Cayo District, British Honduras, April 30, 1938; vernacu- 
lar name "walk naked", 

Of the species in the Yucatan Peninsula, its closest ally 
appears to be E. bumelioides Standl., which has ovate or el- 
liptic broeder leaves with denser darker tomentum. The local 
name, alluding to the smooth brownish or reddish trunk, is 
applied to several other British Honduras eugenias. 


OSMANTHUS MEXICANA Lundell, sp. nov. 

Arbor 6--7 m. alta, 25 cm. diam. Folia glabra, coriacea, 
anguste lanceolata vel oblanceolata, 4--9 cm. longa, 1--2.4 
cm. lata, apice caudeto-acuminata, acumine anguste obtusius- 
culo, basi acuminata, nigripunctata, costa supra impressa; 
petiolo gracili, 1--1.8 cm. longo. Inflorescentise axillar- 
es, anguste paniculatae, 1.5-~-3 cm. longae. Calyois lobi 4. 
Corolla puberulenta, ca. 4 mm. longa, ad mediam partem co- 
alita, lobis late ovatis vel suborbicularibus, rotundatis. 
Ovarium glabrum, biloculare. Fructus ellipsoideus, ca. 15 
mm. longus, 8 mm. dian. 

A tree 6 to 7 m. high, and 25 cm. in diam.; branchlets e- 
rect, glabrous, slender, with short internodes. Leaves oppo- 
site, coriaceous, zlabrous, narrowly lanceolate or oblanceo- 


1939 Lundell, Six New Trees & Shrubs 309 


late, 4 to 9 cm. long, 1 to 2.4 cm. wide, apex caudate- 
acuminate, the acumen narrowly obtusish, base acuminate, de- 
current on the petiole, densely black-punctate, costa 
slightly impressed above, prominent beneath, main lateral 
veins 4 to 6 on each side, nearly obsolete; petioles slender, 
1 to 1.8 cm. long. Panicles of pistillate flowers axillary, 
narrow, small, 1.5 to 3 cm. long, each node bibracteate, 
sparsely and minutely puberulent, glabrescent. Flowers ses- 
sile or subsessile, the pedicels less than 1 mm. long. Calyx 
small, 4-lobed, the lobes triangular, about 0.6 mm. long, 
puberulent above, ciliolate. Corolla puberulent, densely so 
above, about 4 mm. long, 4-lobed to the middle, the Lobes 
broadly ovate or suborbiculer, rounded. Staminodes 2. Ovary 
glabrous, 2-celled, with 2 ovules in each cell. Style stout, 
about 1.5 mm. long. Stigma capitate, large. Fruits ellips- 
oid, about 13 mm. long, 8 mm. in diam. 

Type in the Herbarium of the University of Michigan, Eizi 
Matuda 2025, collected on Cerro Laguna, Mapastepec, Chiapas, 
Mexico, Jan. 1938. 

By its small narrow caudate-acuminate leaves, blackened 
when dry, costa impressed above, and large stigma, O. mexi- 
cana may be readily distinguished from the two other Ameri- 
oan species. 


Sn an ne ee | 


ADDITIONAL NOTES ON THE BRIOCAULACBAE -- I 


Harold N. Moldenke 


On December 1, 1937, I published a discussion of the &r- 
focaulacese of North America in North American Flora 19: 
17--50. The editorial policy of this flora, however, is 
such that one is in many cases not permitted to give com- 
plete synonymy, complete lists of published illustrations 
in the case of abundantly illustrated species, or any lists 
of examined material. Of these the most serious omission is 
that of the citation of herbarium material examined, for 
only by a consultation of such a list of cited material can 
future workers retrace the steps of a monographer and see 
actually on what he based his concept of any given genus, 
species, or variety. Lists of examined material are also 
invaluable in showing exactly what is known of the geo- 


510 PA FTOROGIA No. 9 


graphic distribution of any form, where gaps occur which 
might be filled in by more intensive collecting, what the 
basis is for the limits given, and in what herbaria critical 
specimens are deposited and where they can be studied for 
verification. To make up for these omissions the following 
notes are presented as a permanent record of the material 
examined in the preparation of the above-mentioned work. 
Additional material examined since its publication is also 
included here, together with the description of several new- 
ly discovered species and other miscellaneous notes. 

It should be noted that throughout the work in North Am- 
erican Flora the number of peduncles given in specific or 
varietal descriptions applies in each case to one individu- 
al plant only. This is worth emphasizing, for in many cases 
herbarium material has been mounted in such a way that sev- 
eral or many distinct plants are crowded so close together 
in one mass that the casual observer might be led to suppose 
that only one plant is represented and that the total numb- 
er of peduncles per plant is actually much larger than the 
number given in the description. The phrase “per plant" was 
originally included in all descriptions to emphasize this 
necessity on the part of the observer to determine how many 
plants he is actually looking at on a given specimen, but 
was omitted by the editors to conserve space. It may also be 
mentioned, in passing, that the citation of Literature ref- 
erences in this work is considerably at variance with that 
adopted in all my other publications; again, in order to 
comply’ with the established policies of North American Flora. 
In the present supplementary notes my own system, estab- 
lished in some 75 previous publications, is maintained. 

In the citation of specimens on the following pages, the 
following abbreviations of the names of herbaria are employ- 
ed: Be = Barnard College, New York City; C = Columbia Univ- 
ersity, New York City; De = Delzie Demaree Herbarium, Monti- 
cello, Arkansas; Es «= Estacion Experimental Agronomica, Hav- 
ana; F = Field Museum of Natural History, Chicago; Go = Bot~ 
aniska Trdédgard, Géteborg; I = Langlois Herbarium, Catholio 
University of America, Washington; It = Cornell University, 
Ithaca, N. Y.; K = Royal Botanic Gardens, Kew; Ki = EB. P. 
Killip Herbarium, Washington; M = Meisner Herbarium, New 
York Botanical Garden; Mi = University of Michigan, Ann Arb- 
or, Mich.; N = Britton Herbarium, New York Botanical Garden, 
New York City; P = Muséum Netional d'Histoire Naturelle, 
Paris; Q = Jardin Botanico, Madrid; T = Torrey Herbariun, 
New York Botanical Garden; Tc = Torrey Botanical Club Herb- 
arium, New York Botanical Garden; and Z = H. N. Moldenke 
Herbarium, Watchung, New Jersey. To the directors and cura- 
tore of these 18 herbaria the author extends his sincere 
thanks for their kindness and cooperation in permitting him 


1939 Moldenke, Notes on Eriocaulaceae 511 


to study their material of this group. All material so stud- 
ied -- 920 specimens and mounted photographs -~-- is herein 
cited. 


sc aeent L. Sp. Pl., ede 1, 87. 1753; Gen. Pl., ed. 5, 38. 
1754. 

Cespa Hill, Herb. Brit. 1: pl. 66 [some copies]. 1769; 
Nasmythia Huds. Fl. Ang., ed. 2, 414. 1778; Randalia Petiv. 
ex Desv., Ann. Sci. Nat. Paris, sér. 1, 13: 47, pl. 5, f. 2. 
1828; Symphachne P. Beauv. ex Desv., Ann. Sci. Nat. Paris, 
sér. 1, 13: 47, pl. 5, f. 3. 1828; Sphaerochloa P. Beauv. ex 
Desv., Ann. Soi. Nat. Paris, sér. 1, 13: 47. 1828 ("Sphoero- 
chloa", op. cit. pl. 5, f. 1. 1828]; Leucocephala Roxb. Fl. 
Ind. 3: 612. 1832; Electrosperma F. Muell., Trans. Phil Soc. 
Victor. 1: 23. 1855; Dichrolepis Welw. Apont. Phyt.-geogr. 
542. 1859; Lasiolepis Béck., Flora 56: 90. 1873. 

A genus of about 360 species, widely distributed in 
marshy places in tropical and subtropical regions, the 
greatest number in tropical America; numerous also in troni- 
cal Asia and Africa; one species in northwestern Europe an. 
northeastern North America. 

Excluded species: Eriocaulon leptodictyon A. Gray is cit- 
ed by Durand & Jackson, Ind. Kew. Suppl. 1: 158 (1902) to 
"S. Wats. in Proc. Am. Acad. 22 (1887) 420.--Mexic.", but 
this Watson reference plainly describes Bupatorium (not Eri- 


ocaulon!) leptodictyon. 


3. ERIOCAULON ARENICOLA Britton & Small ex Britton, Bull. 
Torrey Club 44: 31. 1917. 
This species inhabits white sand and has been collected 
in anthesis in February. 
Citations: CUBA: Isla de Pinos: Britton, Britton, & Wil- 


son 14,179 (N--type). 


31. ERIOCAULON BENTHAMI Kunth, Bnum. Pl. 3: 545. 1841. 

Eriocaulon Benthamii Kunth ex Ruhl. in Mngl. Pflanzen- 
reich 4°°: 48, sphalm. 1903. 

The Herb. Hooker specimen of Hartweg 258 at Kew is in- 
scribed "Zacatecas", but the other two specimens of this 
number are inscribed “Lagos” (in Jalisco). The species in- 
habits swamps and marshes, often growing directly in the 
water, and has been collected in anthesis in March and from 
June to August. 

Citations: MEXICO: Chihuahua: E. W. Nelson 6028 (N--2); 
Jalisco: Hartweg 258 (K--2 isotypes, N--isotype); Mexico: 


Hinton 627 (K), 3488 (K), 3638 (K, N), 4549 (K, N). 


1. ERIOCAULON BILOBATUM Morong, Bull. Torrey Club 19: 226. 
1892. 


312 PHY T OL @.G.1 A No. 9 


Eriocaulon Jaliscanum S. Wats., Proc. Amer. Acad. 26: 
157, in part (1891) is often cited as a synonym of this spe- 
cies, but Watson's name more properly is synonymous with E 
Schiedeanum. E. bilobatum inhabits wet hollows and wet 
places in rocky slopes, ascending to SOOO feet, and has been 
collected in anthesis in October. 

“wena MEXICO: Jalisco: Pringle 3855 (C--type), 6299 
(C). 


7. ERIOCAULON COMPRESSUM Lam. Encycl. Méth. 3: 276. 1789. 

Eriocaulon decangulare Walt. Fl. Car. 83. 1788 (not E. 
decangulare L., 1753]; Eriocaulon gnaphalodes Michx. Fl. 
Bor. Am. 2: 165. 1803; EKriocaulon cephalotes Michx. ex Poir. 
in Lam. Rncycl. Méth. Suppl. 3: 161. 1813; Sphaerochloa com- 
pressa (Lam.) P. Beauv. ex Desv., Ann. Sci. Nat. Paris, Bere 
1, 13: 47, pl. 5, f.- 1. 1828 ["Sphoerochloa compressa 
Beauv." in N. Am. Fl. 19: 22. 1938]; ?Briocaulon filiformis 
Raf. Atl. Journ. 121. 1832 (not E. filiformis Bong., Act. 
Petrop. Sci. Math. sér. 4, 1: 634. 1841]. 

The species inhabits moist sandy pinelands, cypress ponds 
and swamps, pine-barren swamps, bogs, flatwoods, millponds, 
swempy woods, moist pine-barrens, open or shaded muddy 
places, hammocks, sandhills, wet sandy soil, snady low 
ground in woods, borders of ponds, everglades, and even the 
water of canals, and has been collected in anthesis from 
January to August. 

Illustrations: Britton & Br. Ill. Fl., ed. 1, 1: f. WOO. 
1896; Ann. Rep. Vew Yersey State Mus. 1910: pl. 64, f. 2. 
1912; Britton & Br. Ill. Fl., ed. 2, 1: f. 1142. 1913. 
Citations: N&* JERSEY: Burlington Co.: Chrysler s.n. (13 

July, 1930] (N), sen. (Chatsworth, 3 June, 1931] (N); Com- 
mons s.n. [Atsion, June 22, 1882] (N); Eames s.n. [Chats- 
worth, VI.12.1894] (N); H. HR. Rusby son. {Bass River, ate 

’ 


1875) (Mi); Torrey & Cooper s.n. [Near Quaker Bridge] (N 
Torrey Botanical Club s.n. [Forked River, May 29--June 2, 
18596] (C); Ocean Co.: Kraemer s.n. [Tom's River, June 30, 
1891] (C); MacElwee 658 (N); H. N. hioldenke 10,577 (N); 
Rich s.n. [Tom's River, May 30, 1887] (C); Camden Co.: Ber- 
inger s.n. [Atco, June, 1895] (Mi); Atlantic Co.: Bassett 
8en. [Hammonton] (I, N--2); Beals s.n. (Hammonton Lake, 
July 1, 1922] (N); Gross 3118 (N), a.n. (N); Cape May Co.: 
C. S. Williamson s.n. (Bennet, June 16, 1907] (N); County 
undetermined: T. F. Allen s.n. [Pine barrens, June, 1871] 
(C); Herb. Columbia University s.n. [Pine barrens] (Cc); 
Herb. Torrey s.n. [Pine barrens] (T). MARYLAND: Baltimore 
Co.: Morong a.n. [Cantcnsville, Aug. 11, 1873] (Bc); Wicom- 
ico Co.: A. V. Smith 225 (I), 333 (1), sen. (May, 1936] 
(1). NORTH CAROLINA: Onslow Co.: House 5108 (N). SOUTH CAR- 
OLINA: Lexington Co.: Weatherby 6123 (N); Charleston Co.: 


ae 


1939 Moldenke, Notes on Eriocaulaceae 313 


Gibbes s.n. [lst March, 1839] (N). GEORGIA: Sumter Co.: R. 
M. Harper 2219 (N); Bryan Co.: R. M. Harper 2170 (N); Charl- 
ton Co.: J. S. Harper 394 (N); County undetermined: A. Gra 
een. [1840] (M). FLORIDA: Duval Co.: Curtiss 3017 (Bo, I); 
Hogg 8 (C); Lighthipe 172 (N); Clay Co.: Canby s.n. [Hiber- 
nia, March, 1869] (T); St. Johns Co.: M. C. Reynolds s.n. 
(St. Augustine, 1877] (C); Levy Co.: O'Neill 732 (I); Leke 
Co.: Nash (C, Mi); Underwood 1928 (C); Orange Co.: Huger 
aon. ("X.Q."] (N); B. J. Palmer 38,325 (N); Pasco Co.: 
O'Neill 1947 (1), 7780 (I); Hillsboro Co.: Leavenworth s.n. 
[Tampa Bay] (T); Polk Co.: McFarlin 4491 (1); Topping 2611 
(N); De Soto Co.: Small & Dewinkeler 9044a (N); Lee Co.: H. 
N. Moldenke 689a (N); Sheehan s.n. (Cat-tail Island, Feb. 
26, 1519] (N); C. Skottsberg s.n. [Buckingham, 19.5.1935] 
(Go); J. P. Standley 15 (N); Broward Co.: Small, Carter, & 
Small 3361 (N); Seminole Co.: O'Neill 6512 (I); Dade Co.: 
O'Neill 7778 (I); J. K. Small aes (N); Small & Carter 2991 
(N); Smell, Carter, & Small 3264 (N); Collier Co.: O'Neill 
I--2); County undetermined: Chapman s.n. [Florida 

T); Herb. Chapman s.n. [Florida] (C--2). ALABAMA: Mobile 
Co.: Gates, Fletcher, & Jewett Luh (Mi); Herb. Univ. Michi- 
gan sen. [Mobile, May 7, 1839] (Mi); Escambia Co.: C. F. 
Baker 1557 (N). LOUISIANA: Calcasieu Par.: &. J. Palmer 
7718 (N). UNITED STATES: State undetermined: Collector un- 
designated 272 (T); Walsh s.n. [Southern States] (T--2). 


7a. ERIOCAULON COMPRESSUM var. HARPERI Moldenke, N. Am. Fl. 
19: 23. 1937. 

This variety inhabits dry or moist pine-barrens, low or 
dry sandy pinelands, sandhills, wet sandy ditches, swamps, 
and shallow ponds, and has been collected in anthesis from 
February to July. 

Citations: GEORGIA: Sumter Co.: R. M. Harper 1395 (N); 
Charlton Cont fe Herper 383 (N). FLORIDA: Jackson Co.: R. 
M. Harper N--type); Washington Co.: Small & Wher 
11,695 Reg Co.: Biltmore Herb. 22968 (N); Leon 
Co.: Kurz s.n. (May 16, 1926] (N); Wakulla Co.: H. N. Mold- 
enke 1123 (N); De Soto Co.: Small & Dewinkeler 9044 (N); 
Lee Co.: H. N. Moldenke 689 (N). ALABAMA: Mobile Co.: Gates, 
Fletcher, & Jewett 18 Gay, 190 (Mi), 192 (Mi), s.n. [Mob- 
ile, May 15, 1839] (Mi). MISSISSIPPI: Harrison Co.: C. F. 
Baker 1558 (N); Langlois s.n. [Pasa Christian, March, 1880] 
(I); Tracy 5032 (Mi, N). LOUISIANA: St. Tammany Par.: Lang- 
lois s.n. [Near Mandeville, 1.V.1893] (I). 


2. ERIOCAULON CUBENSE Ruhl. in Fedde, Repert. 22: 29. 1925. 
This species inhabits alkali flats and white sand, and 
has been collected in anthesis in November. 
Citations: CUBA: Isla de Pinos: Ekman 12,065 (N-isotype). 


314 PHYTOLOGIA No. 9 


4, ERIOCAULON DECANGULARE L. Sp. Pl., ec. 1, 8&7. 1753. 

Eriocaulon statices Crantz, Inst. 1: 360. 1766; Eriocaul- 
on serotinum Walt. Fl. Car. 83. 1788; Randalia decangulare 

L.) P. Beauv. ex Desv., Ann. Sci. Nat. Paris, ser. 1, 13: 
47, pl. 5, f. 2. 1828 ("Randalia decangularis" according to 
Index Kewensis, Ruhland, and North American Flora]; Symph- 
achne xyrioides P. Beauv. ex Desv., Ann. Sci. Nat. Paris, 
ser. 1, 13: pl. 5, f. 3. 1828 ["Symphachne xyroides" accord- 
ing to North American Flora]; Randalia americana Petiv. ex 
Ruhl. in Engl. Pflanzenreich 43°: 33, in syn. 1903. 

The species inhabits wet or moist meadows, argillaceous 
or siliceous bogzy depressions, swemps, wet sandy soil, 
flatwoods, mountain bogs, savannas, wet or low pinelands, 
moist pine-barrens, pine-barrer swamps and ponds, wet or 
boggy places, sandy bogs, moist ditches, cypress swamps, 
ponds, and pond-margins, ascending to 2500 feet, and has 
been collected in anthesis from May to January. Pennell 9075 
has the heads binary on two very short stalks at the apex 
of the peduncle, while the Columbia University sheet of 
Herb. Chapman s.n. also has binary heads with many of the 
outer bractlets elongated into stiff leafy proliferations. 

Illustrations: Ann. Sci. Net. Paris, sér. 1, 13: pl. 5, 
f. 2 & 4. 1828; Lodd. Bot. Cab. 14: pl. 1310. 1828; Curtis, 
Bot. Mag. 59: pl. 3126. 1832; Britton & Br. Ill. Fl., ed. l, 
1: f. S01. 1896; Ann. N. Y. Acad. Soi. 17! : pl. 24, f. ls 
1906; G.-T..Stevens, Ill. Guide Fl. Pl. pl. 9, f. 5. 19103 
Ann. Rep. New Jersey State Mus. 1910: pl. 28, f. 2. 1912; 
Britton & Br. Ill. Fl., ed. 2, 1: f. 1143. 1913. 

Citations: NEW JERSEY: Monmouth Co.: D. C. Eaton s.n. 
{1860] (C); Burlington Co.: Chrysler sen. [W. of Speedwell, 
13 July, 1930] (N); Fogg 4602 (N), 4655 (N); Leggett Sen. 
(Quaker Bridge, July 8, 1864] (Tc); H. H. Rusby een. [Bass 
River, Aug. 1875] (Mi); Van Pelt s.n. (Chatsworth, Aug. 20, 
1907] (N); Ocean Co.: N. L. Britton sen. [Forked River, 
Aug. 17, 1889] (C); Clute 214 (N); Mackenzie 1050 (N), 16 
(Ny, 2361 (N), 4 (N), 4257 (N), 474. (N--2), 5482 (N); 
Poggenburg s.n. (Manchester, Sept. 19, '85] (Tc); Ruger s. 
ne Furialister] (Tc); Schrenk s.n. (Manchester, July 25, 
'77] (Tc); N. Taylor 2853 (N); F. Wilson s.n. (Opposite Is- 
land Heights, July 26, 1909] (N--3); Camden Co.: Bassett s. 
n. [Atco] (I), sen. [Atco, July 19, 19235] (N); Beringer s. 
n. [Atco, Aug. 1890] (Mi); Morong sen. [Atco, Sept. 3, 
1873] (Bc); Pennell 9075 (N), 9079 (N); Atlantic Co.: Grosa 

16 (N); Cumberland Co.: Dreisbach 3593 (Mi); Pennell 1927 

N); Cape May Co.: Parker s.n. [Opp. Sea Isle City, Aug. 
18, '82] (Bc); Ccunty undetermined: T. F. Allen s.n. [1870] 
(N); Collector undesignated s.n. [Pine barrens] (C); Com- 
mons sen. [Pine barrens, Oct. 1, 1872] (N); Herb. Torrey 8. 
n. [Pine barrens, June, 1816] (T); Leggett a.n. [Pine bar- 


DN a el oe 


1939 Moldenke, Notes on Eriocaulacese 15 


rens, Sept. 16--18, 1873] (Tc); A. V. Smith s.n. [(N. J. pine 
barrens, July 28, 1935] (I). DELAWARE: Sussex Co.: N. L. 
Britton 56 (M); Canby s.n. [Ellendale, July 24, 1893] (Mi); 
Commons sen. [dune 18, 1875] (Bc); O'Neill 9096 (1); Pennell 
12 (N). MARYLAND: Prince Yeorges Co.:S. F. Blake 10,665 
I); Tidestrom sen. [July 4, 1907] (I); Worcester Co.: H. H. 
Rusby 8.n. (Stockton, Aug. 1889] (C); County undetermined: 
Holm sen. (Surattsville, Aug. 6, 1900] (1). DISTRICT OF COL- 
UMBIA: Holm sen. (Holmead, July 22, 1888] (1). VIRGINIA: 
Prince George Co.: Fernald, Long, & Smart 6 (N). NORTH 
CAROLINA: Catawba Co.: Small & Heller 180 (C); Rowan Co.: A. 
A. Heller 180 (C, N), 181 (C), sen. (N); J. K. Small sen. 
[Dunn's Mtn., Aug. 18--27, 1894] (C); Dare Co.: Bartle & 
Pontius 503 (N); Buncombe Co.: Biltmore Herb. 3867a (1, N), 
38674 (N); Rugel sen. [Swanano, Jul. 1841 } tis, Cumberland 
Co.: Biltmore Herb. 3867f \(N); Brunswick Co.: Drushel 10,075 
(N); New Hanover Co.: Biltmore Herb. 3867b (N); Coville 202 
(N); County undetermined: Schweinitz sen. (T). SOUTH CAROLI- 
NA: Orangeburg Co.: Eggleston 502 (N). GEORGIA: Columbia 
Coe! a Oo an Bone reas Sumter Co.: R. 
M. Harper N); Lee Co.: Barle 3123 (N), s.n. (Leesburg, 
6720/1895] (N); McIntosh Co.: J. K. Small sen. [About suph by 
Jct., June 25--27, 1895] (C); Lowndes Co.: J. K. Small san. 
(Near Valdosta, June 6--12, 1895] (C); County undetermined: 
Boykin s.n. (T). FLORIDA: Holmes Co.: Drushel 10,095 (N); 
Franklin Co.: Biltmore Herb. 3867c (N); Leon Co.: Berg 5.n. 
{Near Tallahassee] (N); Duval Co.: Curtiss 3016 [June] (Bs, 
C), 3016 [July] (Bc), 5690 (N); Highlands Co.: McFarlin 7545 
(N); Marion Co.: O'Neill 7781 (I); Charlotte Co.: J. H. 
Simpson 396 (C); Lake Co.+ Nash 847 (C, Mi), 1722 (Mi); Or- 
ange Co.: O'Neill 7782 ["Count 13"] (1); Pasco Co.: O'Neill 
81 (I), i153 (I), 1815 (I); Hillsboro Co.: Leavenworth 9.n. 
(Tampa Bay] (T); Polk Co.: McFarlin 3418 (I); Brevard Co,: 
H. N. Moldenke 233 (N); Manatee Co.: Tracy 7587 (N); Lee 
Co.: A. S. Hitchcock 375 (N); Broward Co.: Small & Carter 
1029 (N); Dade Co.: H. N. Moldenke 5591 (N); County undeter- 
mined: Herb. Chapman aen. [Florida] (C--2),. ALABAMA: Mobile 
Co.: C. F. Baker a.n. (Mobile, 7/20/1897] (N); Bush 71 (N); 
Mackenzie 4051 (N); Fennell 4472 (N); Baldwin Co.: O. Blant- 
on 86 (I); Mohr a.n. [Oct. 4, 1894] (I); Tracy 8043 (N--2). 
MISSISSIPPI: Harrison Co.: C. F. Baker 780 (cy, sen. (Bil- 
oxi, 7-23-1897] (N); Lloyd & Tracy 319 (N); Tracy 3428 (N), 
6417 [Biloxi] (N), 6417 [Ocean Springs] (N--2). LOUISIANA: 
Natchitoches ap E. J. Palmer 7961 (N); Calcasieu re 
A. Allison 207 (N); St. Tammany Par.: Langlois sen. |9.Ik. 
1892] (I), *. (Mandeville, 15.VIII.1892} (I); Pennell 4144 
(N); Orleans Par.: T. Drummond 356 (N); Parish undetermined : 
Hale 456 (T), sen. (N). TEXAS: Smith Co.: Reverchon 2766 
(N); Waller Co.: EB. Hall 635 (N). UNITED STATES: State un- 


316 PHYTOLOGIA No. 9 
determined: M. A. Curtis s.n. (C). 


4a, ERIOCAULON DECANGULARE var. LATIFOLIUM Chapm. ex Molden- 
ke, N. Am. Fl. 19: 21. 1937. 
This variety inhabits sandy pinelands, and has been col- 
lected in anthesis in August. 
Citations: FLORIDA: Franklin Co.: Herb. Chapman s.n. 
[Florida] (C--type). ALABAMA: Mobile Co.: Pennell 4465 (N). 


6. ERIOCAULON DIOECUM Ruhl. in Fedde, Repert. 22: 29. 1925. 
This species has been collected in anthesis in March. 
Citations: CUBA: Pinar del Rf{o: Exman 12,807 (N--photo of 

type). 


19. ERIOCAULON ECHINOSPERMOIDEUM Ruhl. in Fedde, Repert. 22: 
31. 1925. 


16. ERIOCAULON ECHINOSPERMUM C. Wright ex Sauv., Anal. Acad. 
Ci. Habana 7: 716. 1871. 

This species inhabits borders of pineland lagoons, moist 
savannas, and shallow water at the edge of ponds, and has 
been collected in anthesis in January, September, and Novem- 
ber. 

Citations: CUBA: Pinar del Rf{o: ae Britton, & Gager 


7257 (N); Exman 18,128 (N); Shafer 11,705 (N); C. Wright 
3738 (T--isotype); Province undetermined: C. Wright een. (T). 


38. ERIOCAULON EHRENBERGIANUM Klotzsch ex Kérn. in Mart. Fl. 
Bras. 3 : 491. 1863. 

Eriocaulon Benthami Schlecht. ex Kérn. in Mart. Fl. Bras. 
3! 491, in syn. 1863 [not E. Benthami Kunth, 1841]; Erio- 
caulon Benthami Seem. apud Hemsl. Biol. Centr. Am. Bot. 3: 
443, in syn. (1885), Hook. f. & Jacks. Ind. Kew. 1: 877. 
1895 [not E. Benthami Kunth, 1841]; Eriocaulon microcephalum 
Hook. & Arn. apud Seem. Bot. Voy. Herald 221, in syn. (1855) 
and apud Hemsl. Biol. Centr. Am. Bot. 3: 443, in syn. 1885 
(not E. microcephalum H.B.K., 1816]; Eriocaulon anceps Sessé 
& Moc. Fl. Mex. 17. 1893 [not E. anceps Walt., 1788 J. 

Cotypes of E. Ehrenbergianum | are Schiede s.n. from swamps 
in the valley of Toluca near La Ventilla, Mexico, and kren- 
berg & Aschenborn 531, Berlandier 760, and Schiede sen. from 
marshy meadows in the neighborhood of Mineral del Monte and 
in the hills and other localities around Mexico City, Feder- 
al District. The first of these 4 cotypes is by inference 
designated as the logotype in N. Am. Fl. 19: 35 (1937). Co- 
types of E. anceps are Sessé, Mocino, Castillo, & Maldonado 
456 and 465, deposited in the Madrid herbarium. 

E. Ehrenbergianum inhabits springy bogs and open marshes 
at high elevations, ascending to 7500 feet, and has been 


t 


1939 Moldenke, Notes on Eriocaulaceae SLi 


collected in anthesis from July to October and December. 
Citations: MEXICO: Chihuahua: Townsend & Barber 117 (N-- 
2); Nayarit: J. N. Rose sen. [August, 1897] (N); Jalisco: 
Beechey 8.n. [Jalisco] (K); Collector undesignated s.n. 
Sept. 6, '92] (F); Edw. Palmer 44 (C, K); Hidalgo: Pringle 
8989 (N); Mexico: Schaffner 226 (C, Mi), 226a (N); Oaxaca: 
W. He. Camp 2265 (N); State undetermined: Sessé6, Mocifo, Cas- 
tillo, & Maldonado 456 (N--photo, Q, Z--photo), 465 (N-- 
ees Z--photo). GUATEMALA: Chimaltenango: Skutch 617 
Mi, N). 


26. ERIOCAULON EKMANNII Ruhl. in Fedde, Repert. 22: 30.1925. 

It is worth noting that Ruhland apparently intentionally 
spelled this specific name "Exmannii", for he did the same 
with Lachnocaulon Ekmannii. In the opinion of the present 
writer the original spelling should be retained. The species 
inhabits swamps and has been collected in anthesis in Octo- 
ber. , 

Citations: CUBA: Pinar del Rfo: Cuesta 374 (N); Ekman 


17,888 (N--isotype). 


20. ERIOCAULON FULIGINOSUM C. Wright ex Griseb. Cat. Pl. 
Cub. 226. 1866. 

Eriocaulon scirpoides Griseb. Cat. Pl. Cub. 226. 1866; 
Eriocaulon trichosepalum C. Wright ex Sauv., Anal. Acad. Ci. 
Habana 7: 715. 1871; Eriocaulon sphaerospermum C. Wright ex 
Sauv., Anal. Acad. Ci. Habana 7: 716. 1871. 

This species has been confused by some with &. Schiedea- 
num and specimens have been erroneously distributed under 
that name. It inhabits moist and pineland savannas, banks of 
lagoons, mud at brooksides or in the shade of grasses, pine- 
lands, and the edges of ponds, and has been collected in an- 
thesis from September to December. The type collection of BE. 
trichosepalum is C. Wright 3740 and of E, sphaerospermum is 
C. Wright 3739, while a cotype collection of E. scirpoides 
ie C. Wrignt 2238. 

Citations: CUBA: Pinar del Rfo: Britton, Britton, & Gager 
7060 (N); Ekman 17,864 (N), 18,127 (N); Shafer 10,663 (N); 
C. Wright 3239 (N--isotype, T--2 isotypes), 3740 (T); Isla 
de Pinos: Ekman 12,106a (N); Santa Clara: Combs 588 (N); 
Leén & Cazanas 5910 (N); Oriente: C. Wright 15 (N), 3238 
(N, T); Province undetermined: C. Wright 3739 (T). BRITISH 
HONDURAS: Gentle 993 (F, I, Mi--2, N); Pelly 73 (F). 


29. BRIOCAULON FUSIFORME Britton & Small in Britton, Bull. 
Torrey Club 44: 32. 1917. 
This species inhabits pinelands and has been collected in 
anthesis in February. 
Citations: CUBA: Isla de Pinos: Britton, Britton, & Wils- 


} 


318 PHYTOLOGIA No. 9 


on 14,951 (N--type). 


39. ERIOCAULON GUADALAJARENSE Ruhl. in Rngl. Pflanzenreich 
4°: 60. 1903. 
Thin species inhabits wet places and has been coLleeten 
in anthesis in November. 
Citations: MEXICO: Jalisco: Pringle 1734 (C--isotype, K-- 
isotype). 


27. ERIOCAULON HETEROPETALUM Ruhl. in Fedde, Repert. 22: 43. 
1925. 
This species inhabits moist white sand on the shores of 
lakes and has been collected in anthesis in September. 
Citations: CUBA: Pinar del Rf{o: Ekman 17,253 (N-~photo of 


type ) 6 


21. ERIOCAULON INSULARE Ruhl. in Fedde, Repert. 22: 32. 1925. 
This species inhabits wet grassy places, pinelands, moist 
white sand barrens, and the borders of lakes, and has been 
collected in anthesis from September to November. 
Citations: CUBA: Pinar del Rfo: Britton, Britton, & Gager 
6960 (N); Ekman 17,808 (N--isotype); Isle de Pinos: Ekman 


12 ,029a (N). 


9. ERIOCAULON KINLOCHII Moidenke, N. Am. Fl. 19: 23--24. 
1937. 
Citations: BRITISH HONDURAS: Kinloch 213 (F--type, N-- 
photo of type, Z--photo of type). 


25. ERIOCAULON KORNICKIANUM Van Heurck & Mill. Arg. in Van 
Heurck, Obs. Bot. 101. 1870. 

This specific name is often written "Koernickianum", but 
in the opinion of the present writer the original spelling 
should be retained. The species inhabits springy places on 
prairies and wet sandy ground, and has been collected in an- 
thesis in June and July. 

Citations: ARKANSAS: Benton Co.: F. be way gots [Silo- 
am Spring, June, 1885] (N); Logan Co.? px De). OKLA- 
HOMA: Pushtamaha Co.: E. J. Palmer 8720 


35. ERIOCAULON LACUSTRE Ruhl. in Fedde, Repert. 22: 53. 1925. 
This species inhabits the water of shallow lakes and has 
been collected in anthesis in October. 
Citations: CUBA: Pinar del Rfo: Ekman 17,877 (N--isotype). 


8. ERIOCAULON LINEARE Small, Fl. SE. U. S., ed. 1, 236 & 
1328. 1903. 
This species inhabits moist or wet pine-barrens, the 
shallow or miry mergins and wet sandy shores of ponds, shal- 


— 


1939 Moldenke, Notes on Eriocaulacese 319 


low pine-barren ponds, bogs, wet woods, and even still water 
to 2 feet deep, and has been collected in anthesis from 
April to June and September. 

Illustrations: Ann. N. Y. Acad. Sci. 17!': pl. 23 & 25, f. 
2. 1906; Small, Man. SE. Fl. 258. 1943. 

Citetions: NORTH CAROLINA: HRenderson Co.: Wherry s.n. 
[Flet Rock Ste., 5-30-1927] (N). GEORGIA: Montgomery Co.: R. 
M. Harper 2146 (N); Bulloch Co.: R. M. Harper 830 (N--type); 
Lowndes Co.: R. M. Harper 1608 (N). FLORIDA: Sante Rosa Co.:? 
R. M. Harper 85 (N); Walton Co.: Curtiss son. (Summer, 1885] 
(N--2); Re M. Harper 47 (N); Gulf Co.: Biltmore Herb. 3865e 
(N); Leon Co.: R. M. Harper 223 (N); Pasco Co.: O'Neill sen. 
(Fish Lake, Feb. 20, 1927] (I). ALABAMA: Baldwin Co.: R. Mi. 
Harper 22 (N). 


34, ERIOCAULON MELANOCEPHALUM Kunth, Enum. Pl. 3: 549. 141. 
Eriocasulon aquaticum Sagot ex Kérn. in Mart. Fl. Brac. 
31; 4O8, in syn. 1863; Lasiolepis aquatica Béck., Flora 56: 

91. 1873. 
Citations: CUBA: Criente: C. Wright 756 (N); Province ur - 
determined: C. Wright 3240 (N, T). 


34a. ERIOCAULON MELANOCEPHALUM ver. LONGIPES Griseb. Cat. 
Pl. Cub. 226. 1866. 

E. P. Killip, in a letter dated July 5, 1938, reports 
thet his No. 32,380 from southwestern Pinar del Rfo is 
clearly this variety. He comments, however: "I just can't 
reconcile myself to its being even a variety of a South Am- 
erican species. We have no material [in the U. S. National 
Herbarium] of the species from South America, but you might 
just meke a mental note that it is worth considering if the 
Cuban is not a full-fledged species". I may say, in res- 
ponse, that I, too, had my doubts about the Cuban "KE. mel- 
anocephalum" and ita variety longipes being conspecific 
with the South American E. melanocephalum. However, no 
authentic South American material of this species has been 
available to me for study and comparison and the original 
deacrintion yields no characters which would separate the 
two. Furthermore, some members of this family seem to have 
unusual distributions. For instance, Tonina fluviatilis and 
Paepalanthus Lamarckii are found both in Ouba and the Gvia- 
nas. More inteneive collecting in the West Indies and Cent- 
ral America may reveal all three species in the intervening 
area. 

Citations: CUBA: Pinar del Rfo: C. Wright 3241 (T-- 
isotype). 


33. ERIOCAULON MEXICANUM Moldenke, N. Am. Fl. 19: 33. 1937. 
This species inhabits springy places and has been col- 


320 PHYTOLOGIA No. 9 


lected in anthesis in July. 
Citations: MEXICO: Jalisco: Pringle 11,202 (N--type). 


37. ERIOCAULON MICROCEPHALUM H.B.K. Nov. Gen. & Sp. Pl. 1: 
255. 1816. 

Eriocaulon pusillum Willd. ex Kérn. in Mart. Fl. Bras. 3% 
492, in syn. 1863 (not E. pusillum R. Br., 1810]; Eriocaulon 
brachypus Van Heurck & Mull. Arg. in Vam Heurck, Obs. Bot. 
1: 96. 1870; Eriocaulon mexicanum Liebm., in herb. [not E. 
mexicanum Moldenke, 1937]. 

The species has been collected in wet meadows and gravel- 
ly soil around mountein springs, at altitudes of 9000 to 
10,000 feet, flowering in July, August, and October. 

Citations: CALIFORNIA: Kern Co.: Xantus de Vesey s.n. 
[Fort Tijon] (T). MEXICO: Mexico: Pringle 6144 (C, K), 7361 
4 ae 7. 13,228 (K); Rose & Painter [929 (N); Schaffner 
226b (C, Mi, N), 426 (N); Pueble: Liebmann s.n. [Chinantla] 
(K). COSTA RICA: San José: Standley & Valerio 43,637 (F), 
43,830 (F). ECUADOR: Carchi: Lehmann 567 (K); Pichincha: 
Jameson 206 (K); Province undetermined: André K.1737 (K). 


13. ERIOCAULON MINUTISSIMUM Ruhl. in Fedde, Repert. 22: 32. 
1925. ‘ 
The species inhabits moist places among grasses and has 
been collected in anthesis in November. 
Citations: CUBA: Pinar del Rfo: Ekman 17,948 (N-- 
isotype). 


24, ERIOCAULON MISERRIMUM Ruhl. in Fedde, Repert. 22: 30. 
1925. 
The species inhabits moist places along readsides and has 
been collected in anthesis in October. 
Citations: CUBA: Isla de Finos: Ekman 11,956 (N--isotype} 


5. ERIOCAULON OLIVACEUM Moldenke, N. Am. Fl. 19: 22. 1957. 
The species inhabits wet places and moist white sand of 

barrens and has been collected in anthesis in November. 
Citations: CUBA: Isla de Pinos: Ekman 12,029 (N--type}. 


28. ERIOCAULON OVOIDEUM Britton & Small in Britton, Bull. 
Torrey Club 44: 32. 1917. 
The species inhabits white sand areas and has been col- 
lected in anthesis in February. 
Citations: CUBA: Isla de Pinos: Britton, Britton, & Wils- 
on 14,220 (N--type). 


32, ERIOCAULON PALMERI Ruhl. in Engl. Pflanzenreich 4°°: 48. 


1903. 
The species has been collected in anthesis from April to 


1939 Moldenke, Notes on Eriocaulaceae 321 


November. 
Citations: MEXICO: Durango: Edw. Palmer 172 (N--isotype). 


30. ERIOCAULON PANAMENSE Moldenke, N. Am. Fl. 19: 31--32. 
1937. 
Citations: PANAMA: Chiriquf: Killip 3614 (F--type, It-- 
photo of type, Ki--isotype, N--photo of type, Z--photo of 
type); Woodson, Allen, & Seibert 1130 (N). 


18. ERIOCAULON PARKERI B. L. Robinson, Rhodora 5: 175. 1903. 

Illustrations: Britton & Br. Ill. Fl., ede 2, 1: fe 1141. 
1913; Marie-Vict. Fl. Laurent. f. 244. 1935. 

Cotypes ere T. P. James s.n. from the shore of the Dela- 
ware River near Cooper's Creek, N. J., and C. F. Parker san. 
from between high and low water merk} Camden, N. J. The for- 
mer is by inference regarded as the logotype in N. Am. Fl. 
19: 27. 1937, although if one of the two collections is to 
be so designated it should certainly be the Parker collect- 
ion, since the species if named after Parker and not after 
James, The species inhabits tidal mudflats, tidel pools in 
arrillaceous ledges, shallow water along river shores and 
banks, and other places mostly submerged at high tide, flow- 
ering from July to October. The leaves and peduncles are of- 
ten purplish-green throughout. 

Citations: MAINE: Penobscot Co.: Fernald & Long 13,165 
(I, Mi, N--2), 13,166 (N); Sagadahoc Co.: Fassett 157 (N); 
Fernald & Long, Plant. Exsicc. Gray. 174 (I, Mi, N--2), 
15,167 (N). CONNECTICUT: Fairfield Co.: Eames 8976 (N); New 
London Co.: R. W. Woodward s.n. [Old Lyme, Aug. 1, 1518] 
(N). NEW YORK: Greene Co.: House 25,128 (N); Columbia Co.: 
Muenscher & Curtis 5600 (N); Rockland Co.: Muenscher & Curt- 
is 5598 (N). PENNSYLVANIA: Philadelphia Co.: Conrad s.n. 
[Near Philadelphia] (T). NEW JERSEY: Mercer Co.: Mackenzie 

236 (N); Monmouth Co.: Carter s.n. (Belmar, July 12, 1910] 
OF Burlington Co.: B. Long 4767 (N); Mackenzie 5671 (N), 
5684 (N); N. Taylor 2558 ( . Ocean Co.: Mackenzie s.n. 
[Weat of Mantoloking, Sept. 21, 1912] (N); Camden Co.: Mack- 
enzie 7344 (N); Pennell (N); C. S. Williamson s.n. 
[Fish House, Oct. 2, 1905] (N), s.n. [Fish House, July 28, 
1906] (N); Atlantic Co.: B. Long 4721 (N); Cumberland Co.: 
B. Long s.n. (Millville, Oct. 7, 1909] (N). DELAWARE: Sus- 
sex Co.: N. L. Britton 24 (N). MARYLAND: Wicomico Co.: Cart- 
er son. (Salisbury, July 15, 1904] (N); County undetermibed: 
Carter a.n. [Gunpowder Sta., Auge 27, 1903] (N). VIRGINIA: 
Alexandria Co.: Dowell s.n. [Aug. 13, 1910] (N); New Kent 
Co.: Grimes 4145 (N). 


14, ERIOCAULON PINARENSE Ruhl. in Fedde, Repert. 22: 22. 
1925. 


322 PHYTOLOGIA NOR 


This species inhabits pinelands, mud, and the borders of 
small ponds, and has been collected in anthesis in February 
and March. 

Citations: CUBA: Pinar del Rf{o: Exman 18,769 (N--isotype); 
Isla de Pinos: Britton, Britton, & Wilson 15,008 (N). 


43, ERIOCAULON PRINGLEI S. Wats., Proc. Am. Acad. 23: 283, 
1888. 
The species is found in damp and wet places on plains, 
and has been collected in anthesis in October. 
Citations: MEXICO: Chihuahua: Pringle 2018 (C). 


11. ERIOCAUt.LON PSEUDOCOMPRESSUM Ruhl. in Urb. Symb. Ant. 1: 
492. 1900. 

Eriocaulon gnephalodes C. Wright apud Ruhl. in Urb. Symb. 
Ant. 1: 492, in syn. 1900 [not E. gnaphalodes Michx., 1803]. 

The species inhabits the Eleocharis belt at the edges of 
lakes and lagoons, and has been collected in anthesis in 
February. 

Citations: CUBA: Pinar del Rfo: Cuesta 371 (N); Exman 
11,221 (N); Province undetermined: C. Wright 3741 (N--iso- 
type, T--2 isotypes). 


17. ERIOCAULON RAVENELII Chapm. Fl. S. U. S., ed. 1, 503. 
1860 e 

The species inhabits wet places, low pinelands, hammocks, 
- and everglades, and has been collected in anthesis from Oct- 
ober to January. 

Citations: SOUTH CAROLINA: Berkeley Co.: Ravenel s.n. 
(St. Johns] (C--type). FLORIDA: Broward Co.: Small & Small 
4429 (N), 4447 (N); Dade Co.: A. A. Eaton sen. (Miami, Nov. 
& Dec. 1903] (N); Small & Carter 654 (N), 888 (N), 1148 (N), 
3067 (N), s.n. [Between Cocoanut Grove & Cutler] (N); Small 
& Nasn sen. [West of Miami, Nov. 1 & 9, 1901] (N). 


42, ERIOCAULON SCHIEDEANUM Kérn. in Mart. Fl. Bras. 3': 492. 
1863. 

Eriocaulon Jaliscanum S. Wats., Froc. Am. Acad. 26: 157 
(1891), of which the type is Pringle 2936, properly a 
straight synonym of this species, is sometimes cited “in 
part" as synonymous with E. bilobatum Morong. The specific 
name is usually lower-cased, but was originally published 
with a capital initial letter. Some specimens from British 
Honduras have been distributed under the name "E. Schiedea- 
num", but are actually &. fuliginosum, which see. 

The species inhabits swamps and wet hollows in rocky 
slopes, ascending to 5000 feet, and has been collected in 
anthesis in October and November. 

Citations: MEXICC: Jalisco: Barnes & Land 159 (F); 


1939 Moldenke, Notes on Eriocaulaceae 323 


Pringle 2 (C), 6146 (C), sen. [Near Guadalajara, Oct. 7, 
1890} (c). i 


36. ERIOCAULON SCHIPPII Standl. in Standl. & Record, Field 
Mus. Publ. Bot. 12: 90, hyponym (1936); Moldenke, N. 
Am. FL. 19: 34. 1937. 

The species inhabits the water of shellow pools in brack- 
ish swamps at low elevations, and has been collected in an- 
thesis in September. 

Citations: BRITISH HONDURAS: Schipp 647 (F--isotype, Mi-- 
isotype, N--type). 


15. ERIOCAULON SCLEROCEPHALUM Ruhl. in Fedde, Repert. 22: 
Ble! L926 

Cotypes of this species are Ekman 12191, 11975, 1190, 
and 10822 from moist places near the bay at Los Indios on 
white quartzite sand in mostly dried-out holes between dense 
tussocks of grass at Santa Bérbera, Isla de Pinos, and in 
sandy somewhat damp places at Herradure, Pinar del Rfo, 
Cuba. The species also inhabits pinelands, mud along brook- 
sides, and sand where water sifts through, and has been col- 
lected in anthesis in February, October, and November. 

Citations: CUBA: Isla de Pinos: Britton, Britton, & Wils- 


on as 48 (N); Ekman 11,975 (N--cotype), 11,990 (N--cotype), 
(N 


12,10 yy 


22. ERIOCAULON SEEMANNII Moldenke, N. Am. Fl. 19: 28--29. 
1937. 
Citations: PANAMA: Canali Zone: Seemann 295 (K--type, N-- 
photo of type, Z--photo of type); Panama: Woodson, Allen, & 
Seibert 1650 (N). 


10. ERIOCAULON SEPTANGULARE With. Veg. Brit. 784. 1776. 
?Cespa aquatica Hill, Herb. Brit. 1: pl. 66 [some cop- 
ies]. 1769; Nasmythia articulata Huds. Fl. Angl., ed. 2, 1: 
415. 1778; Eriocaulon pellucidum Michx. Fl. Bor. Am. 2: 166. 

1803; Eriocaulon pumilum Ref. Atl. Journ. 121. 1832; Erio- 
caulon Noveboracens Pluk. ex Hook. in Curtis, Bot. Mag. 59: 
pl. 3126, in syn. 1832; Nasmythia septanguleris (With.) 
Mart., Nov. Act. Physivo-med. Acad. Caes. Leopolc.-Carol. 
Neat. Cur. 17!': 58, pl. 2, f. 2, hyponym. 1835; Eriocaulon 
articulatum (Huds.) Morong, Bull. Torrey Club 18: 353. 
1891; Eriocaulon decangulare Hope apud Hook. f. & Jacks. 
Ind. Kew. 1: 878, in syn. 1895 (mot EB. decangulere L., 
1753]; Eriocaulon aquaticum (Hill) Druce, Pharm. Journ. ser. 
4, 29: 700. 1909 [not §. acguaticum Sagot, in syn., 1863]; 
Eriocaulon septangulare var. natans Hexamer & Meier, in herb. 
The 1762 date given by Ruhland for Nasmythia articulate 
is erroneous. This binomial was nov published until 1778, es 


724 PB Y, P.0 by OG) Fok No. 9 


has been pointed out by me in N. Am. Fl. The 1818 date given 
by Morong for E. septangulare is also erroneous (as has been 
pointed out by Fernald in Rhodora) and led to his faulty use 
of the specific name “articulatum", taken from Hudson's Na- 
amythia articuleta (1778). Eriocaulon septangulare was act- 
ually first published in 1776. 

The species inhabits tamarack and other bogs, swamps, 
ponds, brackish marshes, sandy and muddy tidal flats, quiet 
pools, muddy or mucky margins and banks of ponds and stream, 
stream deadwaters, shallow snady-bottomed ponds, low. sandy 
or drying mucky shores, gravelly soil, sandy shallows and 
shallow pools in peaty barrens, sandy shores of rivers, 
pond-holes in savannas, boggy soil at the margins of ponds, 
and in shallow or even rather deep quiet water. It has been 
collected in anthesis from July to September. 

Tllustrations: Hill, Herb. Brit. 1: pl. 66 [some cop- 
jes]. 1769; Philos. Trans. Lond. 59: 243, pl. 12. 1770; Sow- 
erby & Sm. Engl. Bot. 11: pl. 733. 1800; Curtis, Fl. Lond- 
ine, ed. 2, 4: pl. 52. 1819--1821; Nov. Act. Physico-med. 
Acad. Caes. Leopold.-Carol. Nat. Cur. 17’: pl. 2, f. 2. 
835; M. E. Jackson, Pict. Fl. f. 1387. 1840; Baxter, Brit. 
Bot., ed. 2, 6: pl. 465. 1843; Deakin, Florigr. Brit. 3: f. 
1457. 1847; Johnson & Sowerby, Brit. Wild Fl. f. 1303. 1858 
--1860; Trans. Linn. Soc. Lond. 22: pl. 68, f. t1--15. 1859; 
T. Moore, Field Bot. Comp. pl. 24. 1862; Benth. I11. Handb. 
Brite Fl. f. 1066. 1865; I-e Maout & Decaisne, Traité Gén. 
Bote, ed. 1, 597 & 598. 1868; Syme, Mngl. Bot. 10: pl. 1546. 
1870; Le Maout & Decaisne, Gen. Syst. Bot. 872. 1873; Pratt, 
Fl. Pl. Grasses, Sedges, & Ferns Gr. Brite, ed. 3: 5: pl. 
228. 1873; Le Maout & Decaisne, Traité Gén. Bot., ed. 2, 611 
& 612. 1876; Hogg & Johnson, Wild Fl. Gr. Brit. 11: pl. 894. 
1880; Baillon, Hist. Pl. 12: 398, f. 371. 1894; Britton & 
Br. Ill. Fl., ede 1, 1: 371, f. 899. 1896; Creevey, Flow. 
Field, Hill & Swamp 129. 1898; Rendle, Fl. Pl. 1: f. 122. 
1904; Praeger, Tour. Fl. W. Ireland pl. 6. 1909; Walton, 
Pract. Guide Wild Fl. & Fr. f. 32. 1909; Journ. Roy. Hort. 
Soce Lond. 36: 301, f. 107. 1910; Karst. & Schenck, Veg.- 
Bild. 8: pl. Jl. 1910; G. T. Stevens, Ill. Guide Fl. Pl. 
pl. 9, f. 9. 1910; Ann. Rep. New Jersey State Mus. 1910: pl. 
28, fe 1. 1912; Creevey, Harper's Guide Wild Fl. 45. 1912; 
Horwood, Plant Life Brit. Isles 3: 340. 1915; Britton & Br. 
Ill. Fl., ed. 2, 1: 454, f. 1140. 1913; Walton, Flower- 
Finder, ed. 1, 129. 1914; ed. 2, 129. 1916; House, Wild Fl. 
New York [Mem. N. Y. State Mis. 15:] 1: pl. 6, f. A. 1918; 
Fitch & Sm. Ill. Brit. Fl., ed. rev. 4, Iss. 2: f. 1082. 
1919; N. Taylor, Guide Wild Fl. f. 13. 1928; Pool, Flowers 
& Fl. Pl., ede 1, fe. 169. 1929; Marie-Vict. Fl. Laurent. f. 
244, 1935. | 

Citations: IRELAND: Groves & Groves sen. [8.VIII.1892] 


= 
« 


1939 Moldenke, Notes on Eriocaulaceae 325 


(I); Tidestrom 11,236 (I). SCOTLAND: Salmon sen. [circa 
1843] (I). ISLE OF SKYE: Greville a.n. (Mi). ST. adit Ar- 
séne 130 (N). SABLE ISLAND: Macoun 22,639 (N); St. John 1168 
(N). NEWFOUNDLAND: Fernald, Long, & Dunbar 26,459 (N); Fern- 
ald & Wiegand 5068 (N); Howe & Leng 856 (N), 1421 (N); Rob- 
inson & Schrenk 112 (C), sen. [12 Aug. 1894] (C). NOVA SCO- 
TIA: Cape Breton Co.: Howe & Lang 686 (N), 765 (N); Macoun 
sen. (North Sydney, July 14, 1884] (C); Nichole 1623 (Mi); . 
C. B. Robinson 396 (N); Richmond Co.: Rousseau 35,5/9 [Nat. 
Herb. Canada 130,088] (N); Guysborough Co.: C. A. Hemilton 
25,149 (N); Halifax Co.: Howe & Leng 1583 (N), 1605 (N); 
Digby Co.: Fernald & Long 20,594 ( ° Yarmouth Co.: Fernald, 
Bissell, Graves, Long, & Linder 20,597 (N). NEW BRUNSWICK: 
St. John Co.: Fowler sen. [St. John, Sept. 1873] (Mi). QUE- 
BEC: Rimouski Co.: C. S. Williamson 1106 (N); Temiscouata 
Co.: J. I. Northrop 184 (C); Portneuf Co.: Marie-Victorin 

8 Ne [Lacs du Laurentides, Aotit, 1912] (I); Montcalm Co.: 
Pennell 16,704 (N); Ottawa Co.: Marie-Victorin 15,737 (N); 
County undetermined: Anselm 435 (N). ONTARIO: Simcoe Co.: 
Britton, Britton, & Timmerman s.n. [Port Sandfield, Sept. l, 
1889] (C); St. Joseph Island: Harper & Harper s.n. [Aug. 3, 
'97] (I). MAINE: Penobscot Co.: Fernald 369 (C); F. L. Har- 
vey son. [Orono, 1884] (C); Waldo Co.: Fernald & Long 13,162 
OF Friesner 10,224 (N); Mount Desert Island: T. G. White 
sen. [Shores of Jordan's Pond] (C); County undetermined: 
Briggs s.n. (Chemo Stream, Aug. 1891] (N). NEW HAMPSHIRE: 
Coos Co.: Minns sen. (Crawford's, Aug. 1887] (C); Grafton 
Co.: Knight sen. [Echo Lake, 12.7.79] (N); Langdon sen. 
[Little Squaw Lake, July 27, 1894] (Mi); Merrimack Co.: 
Clinton sen. [Warner, August, 1935] ray Cheshire Co.: E. J. 
C. Gilbert s.n. [Keene, Sept. 1875] (N); Hillsboro Co.: Bat- 
chelder sen. (Merrimack, Aug. 4, 1917] (N); Sheehan s.n. 
(Goffstown, July 31, 1931] (1). VERMONT: Addison Co.: Brain-=- 
erd sen. [Lake Dmmore, Aug. 16, 1878] (Mi, N); Herb. Chap- 
man sen. [Lake Dmmore, 1859] (C); Orange Co.: Denslow s.n. 
[Newbury, July 31, 1923] (N). MASSACHUSETTS: Berkshire Co.: 
E. Davis sen. [Pittsfield] (T); Hampshire Co.: Collector un- 
designated sen. [Southampton Pond, 1829] (C); Hampden Co.: 
Seymour 268 (N); Essex Co.: L. T. Chamberlain s.n. [Wenham 
Pond, Auge 1899] (N); Morong s.n. (Hamilton, July 27, 1875] 
(Bc); Re He Rich sen. [July 26, 1896] (I); Middlesex Co.: H. 
H. Bartlett 313 (Mi); Morong s.n. [South Natick, Sept. 15, 
1881] (Bc); Norfolk Co.: 3S. F. Blake 4366 (I); Ruggles s.n. 
{Near Milton] (Mi); Plymouth Co.: Herb. Torrey Botanical 
Club sen. [Hanover] (Tc); Barnstable Co.: Hollick s.n. 
[Provincetown] (N); Dukes Co.: E. P. Bicknell 231 (N); Blod- 
gett s.n. [Martha's Vineyard, Aug. 19, '93] (N--2); Fo 

2508 (N); A. B. Northrop a.en. [Nashawena, Aug. 1901] (N), 8. 
n. [Nashawena, July--Aug. 1901] (N); Nantucket Co.: C. A. 


326 PHYTOLOGIA No. 9 


Davis a.en. (Nantucket, 14 Aug. 1911] (Mi); County undeterm- 
ined: C. F. Austin s.n. [East Mass., 1860] (C). CONNECTICUT: 
Litchfield Co.: Leggett s.n. [Canaan Pond) (Tc); Underwood 
3124 (C), sen. [West Goshen, Aug. 1890] (C); Middlesex Co.: 
Denslow s.n. (Killingworth, Sept. 5, 1902] (N); County un- 
determined: D. C. Eaton's.n. (T). NEW YORK: St. Lawrence Cos 
Phelps 296 (N); Oswego Co.: House 5822 (N); Hamilton Co.: By 
B. Lambert 8 (N); Madison Co.: House sen. [North Bay of On- 
eida Lake, July, 1901] (N), s.n. [North shore, Oneida Lake, 
Aug. 1903] (N); Warren Co.: W. Cooper son. [Lake Yeorge, 
Aug. 1817] (T); Wehmeyer s.n. [Hart Lake, Sept. &, 1930] 
(Mi); Washington Co.: A. Fitch s.n. [East Greenwich, 19.4. 
13) (Mi); Rensselaer Co.: H. H. Eaton s.n. [Sand Lake, 1829] 
(T); House 21,951 (N); Steuben Co.: Lucy 2237 (N); Sullivan 
Co.: P. Wilson aon. [Sand Pond, Aug. 13, 1918] (N), sen. 
(Lake Shandelee, Aug. 7, 1918] (N); Ulster Co.: P. Wilson s. 
n. [Lake Katrine, Aug. 18; 1916] (N); Dutchess Co.: Pennell 
= (N); Orange Co.: Lt. Bailey sen. [West Point] (T); Raup 
(N); Putnam Co.: Herb. Chapman sen. [Lake Mahopac, Aug. 
1848] (C); Westchester Co.: W. Cooper son. [Near Tarrytown, 
August, 1819] (T); Cushmen & Word s.n. [Shore of South Pond, 
Aug. 1910] (N); Leggett s.n. [Lake Mohegan, Sept. 6, 1868] 
(C);-Muenscher & Curtis 5611 -(N); Pennell 8400 (N); Queens 
Co.: Bisky s.n. [Lakeville] (C); Nassau Co.: EB. P. Bicknell 
230 (N), sen. [Rockville Center, July 25, '03] (N); Clute & 
Wilson sen. (Hempstead, July 20--21, 1899] (N); Ferguson 
2483 (N), sen. [Meadow Brook, 8-26-20] (N), sen. [Hempstead, 
7-25-19] (N); P. Wilson an. (Merrick, Sept. 11, 1915] (N); 
Suffolk Co.: Clute 116 (N); Ferguson 302b (N), 1869 (N), 
2296 1/2 (N), 7755~(N). PENNSYLVANIA: Luzerne Co.: J. Ke 
Smell s.n. [Lily Lake, Aug. 15, 1889] (N); Pike Co.: Nash s. 
ne (Twin Lakes, July 30, 1909] (N). NEW JERSEY: Sussex Co.: 
N. L. Britton sen. [Morris Pond, Sept. 1%, 1887] (C); Mack- 
enzie 4417 (N--2), 7277 (N); W. de W. Miiler 1334 (N); Nash 
sen. [July 26, 1909] (N); J. K. Small son. [Budd's Lake, 
August 12--14, 1890] (I); Morris Co.: Collector undesignated 
s.n. [Schooley's Mtn., August, 1820] (C); Mackenzie 796 (N), 
1021 (N); Tuckerman s.n. [Mountein Lakes] (T); Ocean Co.: 
Gleason, Smith, & Alexander 173 (N); Hexamer & Meier s.n. 
[Manchester pond, Aug. 25] (N); Mackenzie 46 (N--2), 4850 
(N), S277 (N), sen. (Waretown, Sept. 1922] (N); Camden Co.: 
Crawford & Bliss sen. [Atco, July 10, 1927] (N); O'Neill 
8150 ( J; Atlantic Co.: Beals s.n. [Hammonton Lake] (N); 
Gross 3017 (N); Pennell 8194 (N); County undetermined: Brin- 
ton s.n. [Pine barrens, Sept. 1892] (C). MARYLAND: Harford 
Co.: C. S. Williamson s.n. [Havre-de-Grace, Sept. 1, 1906] 
(N); Wicomico Co.: Canby sen. (Salisbury, Sept. 1887] (C); 
Worcester Co.: H. H. Rusby sen. [Stockton, Aug. 1889] (C). 
INDIANA: Steuben Co.: C. C. Deam 1290 (N), 20,896 (N), 


a 


: 
: 
: 
: 


1939 Moldenke, Notes on Eriocaulaceae 327 


20 (N); Whitley Co.: C. C. Deam son. [Round Lake, 9-1- 
97] i). MICHIGAN: Isle Royale Co.: C. A. Brown 3462 (Mi), 

26 (Mi); Foote son. (July 19, 1868] (Mi); McFerlin 2381 
ro Univ. of Michigan party s.n. (July, 1868] (Mi), sen. 
[1868] (Mi); Houghton Co.: C. A. Davis a.n. (Sept. 1905] 
(Mi); Iron Co.: C. A. Davis sen. [Crystal Falls, Aug. 1905] 
(Mi); Marquette Co.: C. K. Dodge 468 (N), a.n. [Aug. 30, 
1916] (Mi); Duckworth a.n. (Little Lake, Auge 4, 1906] (Mi); 
Metoalf 2172 (N); Alger Co.: Fernald & Pease 3220 (Mi); 
Chippewa Co.: C. K. Dodge s.n. (Aug. 30, 1914] (Mi); F. J. 
Hermann 7146 (Mi, N); N. A. Wood s.n. [Whitefish Point, 
1912] (Mi); Cheboygan Co.: Ehlers 112 (Mi), 1109 (Mi), 1686 
(Mi), 5153 (Mi); Brlanson 395 (Mi); Gleason & Gleason 202 
(N); Miner sen. [Aug. 11, 1931] (I); Presque Isle Co.: 
Enlers 5440 (Mi), 6150 (Mi); F. J. Hermann 7012 (Mi, N); Me- 
gon Co.: Chaney 67 (N); Gratiot Co.: O. A. Davis son. (Alma, 
Aug. 13, '95] (ii--3), sen. [Alma, Aug. '95] (Mi); Case Coe: 
Gates 858 (Mi); Herb. Univ. of Michigan sen. [Aug. 25, 1838] 
(Mi). WISCONSIN: Burnett Co.: Fassett 7/11 (N); Calumet Co.:. 
Hasse sen. [Lake Hilbert, Aug. 18h] (N); County undetermin- 
ed: Hasse sen. [Northern Wisconsin, July, 1882] (N); Hough- 
ton sen. [Lac des Isles, Aug. 5, 1831] (Mi). MINNESOTA: St. 
Louis Co.: Butters sen. petra: Aug. 24, 1919] (N); 
Cook Co.: Butters & Buell N); Crow Wing Co.: Sandber 
sen. [Aug. 1891] (N); tae Co.: B.C. Taylor s.n. (Linn 
Lake, Aug. 1892] (C), sen. [Center City, Aug. 1892] (Mi). 
LOCALITY OF COLLECTION UNDESIGNATED: G., L. Ames s.n. (Mt. 
Luber, W., Sept. 1857] (Mi); Collector undesignated s.n. 
(C); G. W. Wright aon. [1876] (To). 


23. ERIOCAULON SIGMOIDEUM C. Wright ex Sauv., Anal. Acad. 
Ci. Habana 8: 48. 1871. 
Citations: CUBA: Province undetermined: C. Wright 3737 
(T--isotype). 


40, ERIOCAULON TEPICANUM Moldenke, N. Am. Fl. 19: 36. 1937. 
This species has been collected in anthesis in January 
and February. 
Citations: MEXICO: Nayarit: Edw. Palmer 2029 (N--type). 


12. ERIOCAULON TEXENSE Kérn., Linnaea 27: 594. 1856. 
This species inhabits swamps and the wet soil of moist 
pinelands, and has been collected in anthesis in May. 
Citations: TEXAS: Smith Co.: Reverchon 4 (N); County 
undetermined: T. Drummond I1.409 (T--isotype). 


41. ERIOCAULON WILLIAMSII Moldenke, N. Am. Fl. 19: 36. 1937. 
This species inhabits the clay of dried-up ponds, and has 
been collected in anthesis in February and March. 


328 PHYTOLOGIA No. 9 


Citations: BRITISH HONDURAS: W. C. Meyer 134 (F). PANAMA: 
Coclé: R. S. Williams 299 (N--type). 


LACHNOCAULON Kunth, Mum. Pl. 3: 497. 1841. 
A genus of about 10 species, all North American. 


10, LACHNOCAULON ANCEPS (Walt.) Morong, Bull. Torrey Club 
18: 360. 1891. 

Eriocaulon anceps Walt. Fl. Car. 83. 1788; Eriocaulon 
villosum Michx. Fl. Bor. Am. 2: 166. 1803; ?Briocaulon pub- 
igerum Bong., Mém. Acad. St.-Pétersb., sér. 6, 1: 628, pl. 

2. 1831; Lachnocaulon Michauxii Kunth, Enum. Pl. 3: 497. 
1841. 

The species inhabits sandy and wet places, low or moist 
sandy pinelands, flatwoods, ditches, marshes, pinewood 
swamps, wet peaty margins of pine woods, sphagnum-magnolia 
swamps, low grassy pine-barrens, and argillaceous or sili- 
ceous biggy depressions and prairies, and has been collect- 
ed in anthesis from May to October. Some material of L. 
minus has been confused with this species in the past and 
distributed under the name of L. Michauxii. 

The Beyrich specimen cited below is of special interest. 
It was originally identified and labelled as "Eriocaulon 
villosum Michx.", but in pencil has been added "L. Beyrioch- 
ianum Kérn. non villosum". It does not, however, agree 
with the description of L. Beyrichianum and so may not actu- 
ally represent the type collection of that species. 

Illustrations: Mém. Acad. St.-Pétersab., sér. 6, 1: pl. 
42, 18351; Britton & Br. Ill. Fl., ed. 1, 1: 373, f. 905. 
1896; Engl. Pflanzenreich 43°: 241, f. 36. 1903; Britton & 
Beni tiie Fle, eds 2; 1: 456, 2. 1145. 1913. 

Citations: VIRGINIA: Isle of Wight Co.: Fernald & Long 
6121 (N); James City Co.: Grimes 3761 (N); Prince George 
Co.: Fernald, Long, & Smart 5698 (N). NORTH CAROLINA: Bladen 
Co.: Biltmore Herb. 2755 (N); Oosting 3555 (N); New Hanover 
Co.: M. EB. Hyams sen. (Wilmington, July, 1879] (Bc). SOUTH 
CAROLINA: Kershaw Co.: House 2685 (N); Florence Co.: E. 3B. 
Bartram 2277 (N); Ravenel .s.n. [Florence, 1879] (C, Mi); 
Aiken Co.? Ravenel s.n. [Aiken, 1870] (C); Dorchester Co.: 
Gibbes s.n. (Summerville, May 25, 1855] (N). GEORGIA: Rich- 
mond Co.: Outhbert 141 (N), a.n. [Augusta, May, 1881] (N); 
Sumter Co.: R. M. Harper 443 (N); Hmanuel Co.: R. M. Harper 
804 (N); Worth Co.: Pollard & Maxon 562 (N); County undeter- 
mined: Boykin 12 (T); Le Conte sen. (T). FLORIDA: Alachua 
Co.: O'Neill 633 (I); Nassau Co.: Drushel 10,135 (N); Duval 
Co.: Curtiss 3021 (Bc, C, I); Keeler sen. [Vicinity of May- 
port & Jacksonville] (C); Lighthipe 452 (N); Clay Co.: Canby 
gen. (Hibernia, 1869] (N); Lake Co.: A. S. Hitchcock sen. 
(Eustis, June--July, 1894] (1); Nash 1942 (C, Mi); Okeecho- 


1939 Moldenke, Notes on Eriocsaulacease 329 


bee Co.: O'Neill "Count 181" (I); Orange Co.: Huger s.n. 
("x.C.I."] (N); O'Neill "Count 100" (1);Pasco Co.: O'Neill 
786 (I), 7783 (1), 7787 (I); Hillsboro Co.: Britton & Wilson 
19 (N); Folk Co.: MoFarlin 6381 (I); Manatee Co.: Tracy (586 
N); Seminole Co.: C. Skottaberg s.n. (Sanford, 14.5.1935] 
i)3 yoniee eae? cele gens Lie West] (T); County un- 
etermined: ciltmore Herb. 27 > Herb. Chapman s.n. 
{Florida] (C); Herb. Columbia Univ. a.n. [Fla.] (C--2); 
Herb. Le Roy sen. [Fla.] (C); O'Neill son. (Prairies, Oct. 
28, 1929] (1). ALABAMA: De Kalb Co.: Ruth 133 (N); Mobile 
Co.: C. F. Baker 841 (C); Bush 107 (N); Dukes 9 (N); Gates, 
Fletcher, & Jewett 188 (Mi); Jewett s.n. (Mobile, May 9, 
1839] (Mi--2); backenzie 4059 (N); County undetermined: 
Buckley sen. [June] (T). MISSISSIPPI: Harrison Co.: Lloyd & 
Tracy 320 (N); Jackson Co.: Tracy 5031 (Mi, N). LOUISIANA: 
St. Tammany Par.: Langlois s.n. (Mandeville to Covington, 
April, 1879] (I); Orleans Par.: Ingalls sen. (T); Parish un- 
determined: Hale 160 (T). TEXAS: Hardin Co.: Whitehouse 581 
(N). CUBA: Isla de Pinos: Ekman 12,410 (N). LOCALITY OF COL- 
LECTION UNDZSIGNATED: Beyrich s.n. [Amer. bor.] (Mi). 


7. LACHNOCAULON BEYRICHIANUM Sporleder ex Korn., Linnaes 27: 
567. 1856. 

This species inhabits low or rather dry pine-barrens, the 
borders of swamps, sandy shores, and springy places, and has 
been collected in anthesis from March to June and August. 
The binomial is often inaccurately accredited to Kornicke. 
See under L. anceps for comments about 4 Beyrich specimen in 
the herbarium of the University of Michigan. 

Citations: GEORGIA: Charlton Co.: R. M. Harper 1491 (N). 
FLORIDA: St. Johns Co.: M. C. Reynolds s.n. (St. Augustine, 
Mar.--June, 1875] (C, Mi). 


LACHNOCAULON CUBENSE Ruhl. in Fedde, Repert. 22: 34.1925. 
The species inhabits 3andy savannas, and has been coll- 
ected in anthesis in August. 

aan CUBA: Santa Clara: Ek:ian 17,118 (N--photo of 
type). 


z 


se 


6. LACHNOCAULON DIGYNUM Kérn., Linnaea 27: 570. 1856. 

Lachnocaulon diandrum Van Heurok & Mijll.-Arg. in Van 
Heurok, Cbs. Bot. 1: 108. 1870; Lachnocaulon ee Benth. & 
Hook. f'. apud Ruhl. in Rngl. Pflanzenreich 430; 242, in syn. 
1903 [not L. anzeps (Walt.) Morong, 1891]. 

The species inhabits moist sandy pinelands, and nas been 
collected in anthesis in September. 

Citations: ALABAMA: Mobile Co.: Pennell 4474 (N). MISS- 
ISSIPPI: Harrison Co.: Lloyd & Tracy 318 (N). 


330 PHYTOLOGIA No. 9 


8. LACHNOCAULON ECILIATUM Small, Fl. SE. U. 8., ede 1, 235 & 
1328. 1903. 

The species inhabits sandy shores of small lakes and mar- 
gins of ponds, and has been collected in anthesis in Janue- 
ry, August, and September. 

Citations: FLORIDA: Walton Co.: Curtiss 3022 (Bo--isotype, 
C--type, I--isotype, N--isotype); Putnam Co.: R. M. Harper 7 
(N); Lake Co.: Biltmore Herb. 15,001d (N). 


2. LACHNOCAULON EKMANNII Ruhl. in Fedde, Repert. 22: 34, 
1925. 

The species inhabits wet sandy places, sandy banks of 
lakes, and white sand in pinelands, and has been collected 
in anthesis in November and December. It is worth pointing 
out that Ruhland apparently spelled this specific name "Ex- 
mennii” intentionally, for he spelled it thus also in the 
case of Eriocaulon Exmannii. In the opinion of the present 
writer, the original spelling should be retained. The spec- 
ies has in the past been confused with the continental L. 
Engleri and specimens distributed under that name. 

Citations: CUBA: Pinar del Rfo: Ekman 18,122 (N--2, N-- 
fragment & photo); Shafer 11,011 (N--2). 


1. LACHNOCAULON ENGLERI Ruhl. in Engl. Pflanzenreich 4°°: 
241. 1903. 

Eriocaulon maritimum Chapm., in herb.; Lachnocaulon mar- 
itimum Torr., in herb.; Lachnocaulon glabrum Chapm., in 
herb. [not L. glabrum Kérn., 1856]. 

The species inhabits damp sandy and springy places, 
ditches, and lake shores, and has been collected in anthes- 
is in January and July. Vivipary is quite conspicuous on 
some heads of O'Neill 7785a. The species has been confused 
by some with L. glabrum, while the Cuban material distribut- 
ed under this name is ectually L. Exmannii. 

Citetions: FLORIDA: Washington Co.: Chapman sen. (St. An- 
drews Bay, 1838] (C); Herb. Le Roy sen. [St. Andrews Bay] 
(N--2); Putnam Co.: R. M. Harper 8 (N); Lake Co.: Nash 1184 
(C--isotype, Mi--isotype); Pasco Co.: O'Neill 5a (I); Or- 
ange Co.: O'Neill s.n. [Lake Ola, July 2, 1929] (1); County 
undetermined: Chapman sen. [Florida] (T); Collector undesig- 
nated a.n. [Florida] (N). 


3, LACHNOCAULON FLORIDANUM Small, Fl. SE. U. S., ed. 1, 235 
& 1328. 1903. 

The species inhabits low sandy places, and has been col- 
lected in anthesis in June. The specific name is often 
written with a capital initial letter. 

Citations: FLORIDA: Lake Co.: Nash 1981 (C--type, N-- 
isotype). 


1939 Moldenke, Notes on Eriocaulacese 351 


4, LACHNOCAULON GLABRUM Kérn., Linnaea 27: 568. 1856. 

The species inhabits low pinelands, damp woods, prairies, 
palmetto hammocks, flatwood ponds, and wet sandy places, and 
has been collected in anthesis in Merch, May, July, Auguat, 
October, and November. 

Citations: FLORIDA: Lee Co.: A. S. Hitchcock 374 (N); J. 
K. Small 8339 (N--2); J. P. Standley 33 (N); Broward Co.: 
Small & Carter 1024 (N); Small & Wilson 1608 (N), 1783 (N); 
Dade Co.: Small & Carter 690 (N); Small, Mosier, & Small 
6892 (N), 6912 (N); Small & Wilson 1613 (N). 


9. LACHNOCAULON MINUS (Chapm.) Small, Fl. SE. U. S., ed. l, 
235. 1903. 

Lachnocaulon Michauxii var. minor Chapm. Fl. S. U. S., 
ed. 3, 531. 1897; Lachnocaulon minor (Chapm.) Smali, F. SE. 
U. S., ed. 1, 1328, sphalm. 1903; Lachnocaulon Michauxii 
minus Chapm. ex Small, Man. SE. Fl. 257, sphalm. 1933. 

The species inhabits moist soil or cultivated ground, low 
pine-barrens, pinelands, and the area above high-water mark 
on the sandy shores and margins of ponds and lakes, and has 
been collected in anthesis in March, from May to September, 
arid in November. It has been confused by some in the past 
with L, Beyrichianum, L. glebrum, and L. Michauxii [#=L. an- 
ceps], and with Paepalanthus flavidulus [=Syngonanthus flav- 
idulus] and the Brazilian P. pilulifer Kérn. and herbarium 
material has been distributed under those names. 

Citations: NORTH CAROLINA: Halifaz Co.: C. S. Williamson 
sen. [Weldon] (N); New Hanover Co.: Canby sen. (Prope Wilm 
ington, Oct. 1867] (Mi); C. S. Williamson sen. [Wilmington] 
(N). GEORGIA: Lowndes Co.: R. M. Harper 1607 (N). FLORIDA: 
Walton Co.: Curtiss 5911 (N); Liberty Co.: Herb. Chapman s. 
Ne (Bristol ] (N--type ; Leon Co.: Berg son. (Near Tallahass- 
ee] (N); Clay Co.: Canby a.n. (Hibernia, March, 1869] (N); 
Lake Co.: R. M. Harper 42 (N); A. S. Hitchcock sen. (Eustis, 
June--July, 1894] (N); Nash 148 (C, Mi), 1295 (C, Mi), 1855 
ay Volusia Co.: Curtiss 6894 (N); Pasco Co.: O'Neill 7785 
(I); Broward Co.: Small & Carter 1037 (N). 3 


PAEPALANTHUS Mart., Nov. Act. Physico-med. Acad. Caes. 
Leopold.-Carol. Nat. Cur. 17': 13. 1835. 

Dupatya Vell. Fl. Flum. 35. 1825 [nom. rejic.]; Stephano- 
phyllum Guill. in Deless. Icon. Sel. 3: 61, pl. 96, in obs. 
1837; Cladocaulon G. Gardn. in Hook. Icon. Pl. pl. 528. 
1844; Limnoxeranthemm Salzm. ex Steud. Syn. Pl. Cyp. 2: 
281, in syn. 1855; Lasiolepis Béck., Flora 41: 90, in part. 
1873. 

Sections III and IV of Eriocaulon as considered in Steud. 
Syn. Pl. Cyp. 2: 273 (1855) also belong here. It is most un- 
fortunate that the present International Rules force us to 


332 PHYTOLOGIA No. 9 


use the generic name Paepalanthus instead of the name 
Dupatya, which was published validly 10 years earlier! 

A genus of about 450 species of tropical America, 
most numerous in Brazil. 


5. PAEPALANTHUS ALSINOIDES C, Wright ex Sauv., Anal. Acad. 
Ci. Habana 8: 49, 1871. 

This binomial is accredited by Britton to Wright and Sau- 
valle and the synonymous cheironym, Dupatya alsinoides 
(Wright & Sauv.) Britton, occurs on widely distributed spec- 
imens. The species inhabits gravelly pinelands and dry sandy 
pine woods, and has been collected in anthesis in September 
and December. 

Citations: CUBA: Pinar del Rfo: Britton, Britton, & Gager 
7O90 (N), 7130 (N); Ekman 17,918 (N); ©. Wright 3743 (N-- 
pres T--isotype); Province undetermined: C. Wright s.n. 
(T e 


5a. PABPALANTHUS ALSINOIDES var. MINIMUS Jennings, Ann. Car- 

The variety inhabits white sand or gravelly soil, sandy 
pinelands and pine-barrens, and hillsides, and has been col- 
lected in anthesis in February, March, May, and October. 

Illustrations: Ann. Carnegie Mus. 11: pl. 17, f. E--H. 
1917. 

Citations: CUBA: Pinar del Rfo: Haman 11,034 (N); Shafer 
10,682 (N--2); C. Wright 3743a (N); Isla de Pinos: Britton 
= : = ? 
Britton, & Wilson 14,144 iar Ekman 11,965 (N), 12,100 (N); 
Jennings 387 (N--isotype). 


3, PAEPALANTHUS CHIAPENSIS Moldenke, N. Am. Fl. 19: 39. 1937 
The species inhabits wet localities and even shallow wat- 
er of ponds, and has been collected in anthesis in May. 
er MEXICO: Chiapas: Purpus 179 (N), 10,565 (N-- 
type). 


1. PAEPALANTHUS COSTARICENSIS Moldenke, N. Am. Fl. 19: 38. 
1937-6 
The species inhabits sphagnum bogs at high elevations, 
ascending to 6400 feet, and has been collected in anthesis 
in December. 
Citations: COSTA RICA: San José: P. Oo. Standley 42,726 
(F--type, N--photo of type, Z--photo of type). 


9. PAEPALANTHUS DOMINGZNSIS Ruhl. in Urb. Symb. Ant. 1: 485. 
1900, 
Cotypes are Eggers 2216 and 2216b from among rocks at 
Valle Nuevo, altitude 2270 m., Dominican Republic. The spe- 
cies also inhabits grassy places at high elevations, and has 


\ 
. 
} 


a = 


1939 Moldenke, Notes on Eriocaulaceae 533 


been collected in anthesis in June and July. 
Citetions: HISPANIOLA: Dominican Republic: Fuertes 1748 
(N); Tirckheim 3422 (N). eee wal 


7. PASPALANTHUS GENTLEI Noldenke, N. Am. Fl. 19: 40-41. 1937. 
The species has been collected in anthesis in “ecember. 

It apparently grown in company with P. Lamarckii, for both 

species are intimately mixed on many sheets of the type col- 


lection. The P. Lamarckii materiel is here regarded as 


Gentle 992a. 
Citations: BRITISH HONDURAS: H. H. Bartlett 11,874 (Mi); 
Gentle (F--isotype, I--isotype, Mi--2 isotypes, N-- 


type); O'Neill 8547 (I, Mi). 


6. PAEPALANTHUS LAMARCKII Kunth, Mum. Pl. 3: 506. 1841. 

Sriocaulon fasciculatum Lam. Rncycl. Méth. 3: 276. 1789 
(not E. fasciculatum Rottb., 1778]; Paepalanthus Ottonis 
Klotzech in Schomb. Reise in Br. Guian. 3: 1115. 1848; Erio- 
caulon Lamarckii (Kunth) Steud. Syn. Pl. Oyp. 276. 1855; 
Lasiolepis pilosa Bock., Flora 56: 9. 1873; Dupatya Lamarck- 
ii (Kunth) Kuntze, Rev. Gen. Pl. 2: 746. 1891. 

The species apparently grows in company with P. Gentlei 
and Syngonanthus Bartlettii, for it is intimately mixed with 
these species on sheets of Gentle 992 and H. H. Bartlett 
11,263 _. The latter collection also has several other cont- 
aminations, mostly xyridaceous, cyperaceous, and juncaceous. 
It inhabits roadsides, pinelands, swampy places, and savan- 
nas near sea-level and is said to be “rare” in British Hon- 
duras. It has been collected in anthesis in November and De- 
cember and February. It occurs rather abundantly in Venezue- 
la and the Guianas. 

Illustrations: Lam. Encycl. Méth. Ill. pl. 50, f. 3. 1791. 

Citations: BRITISH HONDURAS: H. H. Bartlett i1,263 (Mi-- 
2); Gentle peas (Mi, N); Schipp S-130 (F, N). PANAMA: Coclé: 
H. Pittier 4932 (N). CUBA: Pinar del Rfo: Exmen 18,121 (N-- 
2); Isla de Pinos: Ekman 12,015 (Mi, N); Province undeternm- 
a C. Wright 3742 (T--2). TRINIDAD: W. E. Broadway 9310 
(I). 


8. PAEPALANTHUS MELLII Moldenke, N. Am. Fl. 19: 41. 1937. 
The species has been collected in anthesis in November, 
Citations: MEXICO: Veracruz: C. D. Mell sen. (Minatitlan, 

Nov. 28, 1928] (N--type). 


13. PAEPALANTHUS MONTANUS (Britton) Moldenke, Revista Sud- 
ame Bot. 4: 17% 1947. 
Dupatya montana Britton, Bull. Torrey Club 44: 33. 1917. 
The species inhabits compact red iron ore, open places a- 
long trailsides, banks of streams, and pine woods at high 


elevations, ascending to 5000 feet, and is said to be scarce 
and local. It has been collected in anthesis in November, 
December, and February. 

Citations: CUBA: Oriente: Ekman = (N); Shafer 4104 (N 


--3), 4473 (N--type & 2 isotypes), 8045 5 (N), 8251 poe 


11. PAEPALANTHUS PUNGENS Griseb. Cat. Pl. Cub. 224, 1866, 
Dupatya pungens (Griseb.) Britton, Bull. Torrey Club 44: 
33. 1917.~ 
The species is found along the coast, ascending to 2300 
feet, and has been collected in anthesis in May and July. 
Citations: CUBA: Oriente: Exman 2341 (N), 5709 (N). 


334 PHYTOLOGIA. No. 9 


I 
4, PAEPALANTHUS RETUSUS C. Wright ex Sauv., Anal. Acad. Ci. 
Habana 8: 50. 1871. ; 
The species inhabits sandy pine woods and pinelands and 
is said to be “rare”. It has been collected in anthesis in 
May. 
Citations: CUBA: Piner del Rfo: Ekman 12,806 (N); Prov- 
ince undetermined: C, Wright 3744 (T--isotype). 
7 
} 


12. PABPALANTHUS RIPARIUS Moldenke, N. Am. Fl. 19: 42--43, 
1937. 

The species inhabits moist banks bordering streams, as- 
cending to 2700 feet, and has been collected in anthesis in 
February. 

Citations: CUBA: Oriente: Shafer 4106 (N--type). 


2. PAEPALANTHUS SESLERIOIDES Griseb. Cat. Pl. Cub. 224.1866. 

Dupatya seslerioides (Griseb.) Britton, in herb. 

The species inhabits dry white quartz sand, sandy pine- 
lands and savannas, plains, pine-barrens, and the edge of 
clay banks of rocky rivers, and has been collected in anthe- 
sis in February, March, May, September, and December. 

ure sae CUBA: Pinar del Rfo: Britton, Britton, & Jeger 


O (N), LA IN), 7251 (N); Exman 11,033 (N), 17,810 (N); 

n & Roca nee O (N); Shafer 10,882 ss, 10,956 (N); C. 
ce 2a T--2 isotypes); Tele de Pinos: Britton, Britton 
& Wils 225 (N); Britton & Wilson 14,319 (N), 15,689 
Gi)s Jeminge 3 a 338 (N); Oriente: Shafer 4093 (N); C. Wright 
8 


10. PAEPALANTHUS TUERCKHEIMII Ruhl. in Urb. Symb. Ant. 7: 
173. 1912. 
The species inhabits pine woods, ascending to 6800 feet, 
and has been collected in anthesis in September. 
Citations: HISPANIOLA: Dominican Republic: Chardon 28 
(N); Tiirckheim 3327 (N--isotype). 


1939 Moldenke, Notes on Eriocaulaceae 335 


SYNGONANTHUS Ruhl. in Urb. Symb. Ant. 1: 487. 1900. 

The following subgenera of Paepalanthus as considered in 
earlier works, belong here: Thyrsanocephalus Korn. in Mart. 
Fl. Bras. 3!: 429, 1863; Bulepis Bong., Mem. Acad. Sr.- 
Pétersb. Sci. Math., sér. 6, 1: 618 & 635. 1841; Andras pid- 
opsis Kérn., op. cit. 439; Psilocephalus Kérn., op. cit. 
451, in part; and Carphocephalus Kérn., op. cit. 465. 

A genus of about 160 species, mostly of tropical America; 
a few in tropical Africa. 


2. SYNGONANTHUS ANDROSACEUS (Griseb.) Ruhl. in Urb. Symb. 
Ant. 1: 488. 1900. 

Paepalanthus androsaceus Griseb. Cat. Pl. Cub. 225. 1866; 
Paepalanthus androsaceus var. flavescens Griseb. Cat. Pl. 
Cub. 225. 1866. 

The species inhabits moist sand, sandy pastures and pine- 
lands, wet sandy savannas, roadsides, and the margins of 
railroad tracks, and has been collected in anthesis in Janu- 
ary and April. The type collection of Paepalanthus androsac- 
eus var. flavescens is C. Wright 3236. 

Citations: CUBA: Pinar del Rfo: Ekman 10,792 (N); Leén 
15,946 (N); Shafer 11,784 (N); Van Hermann 570 (N); C. 
fright 749 (N), 3235 (T--isotype), 3236 a. 


7. SYNGONANTHUS BARTLETTII Moldenke, sp. nov. 

Herba parva; foliis caespitosis 3--12 mm. longis valde 
recurvatis angustissimis; pedunculis solitariis vel paucis 
pilosis, pilis longis valde patentibus; bracteolis hyalinis. 

Very small herb, 2--7 cm. tall; leaves caespitose, lin- 
ear, dark-olivaceous, conspicuously recurved and appressed 
to the ground, 3--12 mm. long, about 0.5 mm. wide, blunt at 
apex, glabrate or very obscurely puberulent on both surf- 
aces; sheaths closely appressed, conspicuously surpassing 
the leaves, conspicuously pilose with long wide-spreading 
(hirsute) hairs, sharply acute or acuminate at apex; pedun- 
cles solitary or few, very slender, obscurely sulcate, 
slightly twisted, conspicuously pilose throughout with elon- 
gated hirsute wide-spreading hairs like the sheaths; heads 
solitary, 2--5 mm. wide; involucral bractlets hyaline, com- 
pletely colorless throughout, elliptic-lanceolate, sharply 
acute or acuminate at apex, glabrous; staminate florets: 
pedicels about 0.5 mm. long, very densely long-tomentose at 
base; sepals 3, elliptic-obovate, 1--1.125 mm. long, free, 
hyaline, acute at apex, ocuneate-narrowed at base; petals 
very hyaline, closely connate, forming a very narrow tube; 
pistillate florets: pedicels about 0.5 mm. long; sepals 
free, lanceolate-ovate, about 1.5 mm. long, acuminate at a- 
pex, not ciliate on the margins; petals linear-oblanceolate, 
very narrow, about half as wide as the sepals, connate by 


5336 PHY PB Od eerie No. 9 


their margins. 

The type of this species was collected by Dr. Harley Har- 
ris Bartlett (No. 11,670) -- in whose honor it is named -- 
on a flat wet upland at Mountain Pine Ridge, El Cayo Distr- 
ict, British Honduras, on February 22, 1931, and is deposit- 
ec in the herbarium of the University of Michigan. The spec- 
ies also inhabits pine ridges and has been collected in an- 
thesis in Jenuary and February. It apparently grows in comp- 
any with Paepalanthus Lamarckii, since H. H. Rartlett 11,263 
originally contsined a mixture of these two species, the 
new species being here regerded as No. 11,26%a. 

BRITISH HONDURAS: H. H. Bartlett 11,263a (Mi--2, N), 
11,670 (Mi--type & isotype, N--fragment of isotype). 


10. SYNGONANThHUS CAULESCENS (Poir.) Ruhl. in Engl. Pflanzen- 
reich 43°: 267. 1903. 

Eriocaulon caulescens Poir. in Lam. Encycl. Méth. Suppl. 
3: 162. 18135; Eriocaulon splendens Bong., Mém. Acad. St.- 
Pétersb. Sci. Math., ser. 6, 1: 633, pl. 66. 1831; Paepalan- 
thus caulescens (Poir.) Kunth, Enun. Pl. 3: 537. 1841; Faep- 
alanthus surinamensis Miq., Linnaea 19: 126. 1847; Paepalan- 
thus procerus Klotzsch in Schomb. Reise in Br. Guian. 3: 
1115. 1848; Eriocaulon surinamense Miq. ex Steud. Syn. Pl. 
Cyp. 2: 275. 1855; Eriocaulon geraénse Steud. Syn. Pl. Cyp. 
2: 276. 1855; Eriocaulon simillimum Steud. Syn. Pl. Cyp. 2: 
277. 1855; Eriocaulon subuncinatum Steud. Syn. Pl. Cyp. 2: 
277. 1855; Paepalanthus splendens (Bong.) Mart. ex Kérn. in 
Mart. Fl. Bras. 3!: 466. 1863; Dupatye caulescens (Foir.) 
Kuntze, Rev. Gen. Pl. 2: 745. 1891. 

The species ascends to over 4300 feet and has been col- 
lected in anthesis in February. 

Citations: COSTA RICA: Province undetermined: H. Pittier 
sen. [Herb. Instit. Physico-geogr. Nat. Costaric. 11,043 
(N). 


1. SYNGONANTHUS FLAVIDULUS (Michx. ) Ruhl. in Engl. Pflenzen- 
reich 43°: 256. 1903. 
Eriocaulon flavidulum Michx. Fl. Bor. Am. 2: 166. 1803; 


?7Eriocaulon spathaceum Raf. Atl. Journ. 121. 1832; Faepalan- 
thus flevidulus (Michx.) Kunth, Enum. Pi. 3: 532. 1841; Pae- 
alanthus flavidus Kunth ex A. Gray, Man. Bot. North. U. 5., 
ed. 2, 489, sphalm. 1856; Eriocaulon flavidum Michx. ex A. 
Gray, Man. Bot. North. U. S., ed. 2, 589, in syn., sphalm. 
1856; Dupatya flavidula (Michx.) Kuntze, Rev. Gen. Pl. 2: 
745. 1891. 
The species inhabits low and wet sandy pinelands, margins 

of sandhill ponds, dogs, wet scrub-land, prairies, low and 
sandy pine-barrens, wet sandy ditches, moist sandy soil, send 


the borders of ponds. 


ie _ 
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PHY TOLOGIA 


Designed to expedite botanical publication 


Vou. 1 November, 1939 No. 10 


CONTENTS 


C. L. Lunpetn: New or Otherwise Noteworthy Plants from Mexico 
| Wad Ceninal America Oey Sle i ee Me espe etter e PF 


+H. A. Gueason: Four Central American Melastomes.....++++++ 340 
i: N. Moxpenxe: Additional Notes on the Eriocaulaceae--II..... S43 


H. N. Mowpene: Additional Notes on the Genus Aegiphila--V.... 364 


- Published by H. A. Gleason and Harold N. Moldenke > 


js The New York Botanical Garden 
Ay ue hadi Fare Sev York, N. Y. 


2 


\ Hi : ‘Veo. 4 ‘nomber % was ined January 31, 1939 


ut Bit re . ' aah 
Tale 


7 


mm (ay : 
NEW OR OTHERWISE NOTEWORTHY PLANTS FROM MEXICO AND CENTRAL 
AMERICA (a) 


C. L. Lundell 


PIPER MAYANUM Lundell, sp. nov. 

Arbor parva, ca. 4m. alta, 6.5 cm. diam., ramulis et 
petiolis villoso-tomentosis; folia chartacea, vetiolo 3--5.5 
cm. longo, alato; lamina chartacea, oblongo-ovata, 23--50 
cm. longa, 12.5--24.5 cm. lata, apice abrupte acuminata, ac- 
umine obtusa, basi inaequilateraliter auriculata sinu later- 
ali, supra glabra, punctata, subtus minute villosa, pinnati- 
nervi, venis circiter 7 vel 9, subtus villoso-tomentosis; 
spicae usque ad 43.5 om. longee, 0.7 cm. crassae, pedunculo 
usque ad 10.5 cm. longo. -~ BRITISH HONDURAS: Stann Creek 
District, Middlesex, in high ridge forest on hill top, June 
12, 1939, Percy H. Gentle 2866 (type in University of Michi- 
gan Herbarium). -- P. mayanum closely resembles P. hians 
Trel. of Costa Rica, a species with much coarser pubescence, 
subobovate-oblong leaves, open basal leaf sinus, and spikes 
only 18 cm. long. In P. mayanum the leaf sinus is closed by 
the strongly overlapping lobes. 


TREMA STRIGILLOSA Lundell, sp. nov. 

Arbor 8-metralis, 12.5 cm. diam., erecta; ramulis puberu- 
lis et parce hirsutis; folia petiolata, petiolo 7.5--10 mn. 
longo, canaliculato; stipulae ovato-lanceolatae, ca. 4 mm. 
longae, acuminatae; lamina lanceolata vel oblongo-lanceolata, 
7-5--10 cm. longa, 2--43.1 cm. lata, apice caudato-acuminata, 
basi late rotundata, subaequali, inconspicue crenulato- 
serrulata, costa et venis supra impressis, subtus elevatis, 
supra scabrida, subtus minute strigillosa; cymae usque ad 
2 cme longae, puberulae; calyx breviter stipitatus; sepala 
puberula, inaequalia, lanceolata, 1--2 mm. longa, ciliata; 
overium glabrum; stigmate rufo-villoso, lobis divaricatis; 
fructus ignotus. -- BRITISH HONDURAS: Stann Creek District, 
Middlesex, alt. about 60 m., in secondary forest; flowers 
cream colored, faintly perfumed; Nov. 11, 1929, W. A. Schipp 
439 (type in the University of Michigan Herbarium). -- The 
narrowly lanceolate unequal sepals 1 to 2 mm. long and the 
leaf blades rounded at base and minutely strigillose distin- 
guish T. strigillosa. T. floridana and T. micrantha, the 
other two species of the peninsula, have ovate equal sepals 
about 1 mm. long, leaf blades cordate or subcordate at base, 
and under surface of leaf with spreading pubescence. 


PHYLLANTHUS ANTILLANUS (Adr. Jussieu) Muell. Arg. var. HYPO- 
557 


338 PHY TOLOGIA No. 10 


MALACUS (Standl.) Lundell, comb. nov. 
Phyllanthus nobilis (L. f.) Muell. Arg. var. hypomalacus 
Standl., Carmegie Inst. Wash. Publ. 461: 68. 1935. 


HIPPOCRATEA AUSTIN-SMITHII Lundell, sp. nov. 

Frutex scandens; ramuli subteretes, patentes, hornotini 
viridescentes, glabri; folia glabra, membranacea, utrinque 
viridia, petiolsta, petiolo supra sulcato, 4--8 mm. longo, 
gracili; lemina lenceolata, 6--10.5 cm. longa, 2--3.8 cm. 
leta, apice acuminata, acumine obtusiusculo, basi cuneata, 
subintegra, venis lateralibus, utroque latere 4--6, utrin- 
que conspicue prominulo-reticulata; cynae axillares, glab- 
rae, pauciflorae, 1.5--2.5 cm. longae (pedunculo incluso), 
pedicellis ultimis usque ad 2 mm. longis; flores ad 5 mn. 
lati; calyx stipitatua; sepala 5, depresso-ovata vel rotund- 
ata, ciliolata, usque ad O.7 mm. longa; petala 5, late 
ovato-elliptica, ca. 2 mm. longa, glsbra, subintegra; discus 
carnosus, annularis vel subcupularis; stamina 35; carpidia 
desunt. -- COSTA RICA: Province Alajuela, Canton San Carlos, 
Villa Guesada, alt. 850 m., in rain forest, a woody vine 
climbing 5 m., Feb. 28, 1939, Austin Smith H-1725 (type in 
the University of Michigan Herbarium). 


HIPPOCRATEA LANCIFOLIA Lundell, sp. nov. 

Frutex alte scandens, ramulis glabris, atrorubris, novel- 
lis quadrangularibus; folia glabra, petiolata, petiolo 2.5-- 
3.5 mm. longo, canaliculato; lemina subcoriacea, lanceolata 
vel anguste elliptico-oblonga, 5.5--7.3 cm. longa, 1./--3-3 
cm. lata, apice obtuse acuminsta vel obtusa, basi acutiuscu- 
la, repanda, venis lateralibus utroque latere 4--6, venulis 
remote reticulatis; infructescentiae glabrae; carpidia 3, 
complanata, ovata vel elliptica, 3--4.8 cm. longa, 2--2.5 cm. 
lata, apice late obtusa, basi rotundata, crasse coriacea, 
nervosa. -- BRITISH HONDURAS: Stann Creek District, Sittee 
River, along river bank in shade, March 20, 1930, We. Aw 
Schipp 715 (type in the University of Michigan Herbarium). 
-- The ovate or elliptic strongly veined capsules and the 
small subcoriaceous openly reticulated leaves characterize 
H. lancifolia, a relative of H. celestroides H.B.i. 


DOLIOCARPUS CORIACEUS (Mart. & Zucc.) Gilg. 

Pinzona coriacea Mart. & Zucc. -- BRITISH HONDURAS: Stann 
Creek District, Middlesex, a woody vine in high ridge on 
hillside, July 2, 1939, Percy H. Gentle 2892, fruits. -- Ap- 
parently this species has been known previously only from 
equatorial Brazil. 


EUGENIA COCQUERICOTENSIS Lundell, Bull. Torrey Bot. Club 64: 
554. 1937. 


a 


at “a= 


a See oe 


1939 j Lundell, New or Noteworthy Plants 339 


BRITISH HONDURAS: El Cayo District, on hill top ovnosite 
Vaca, a tree 8 in. diam., flowers white, April 30, 1948, 
Percy H. Gentle 2533; vernacular name "female boy Job". -- 
From the type (Lundell 4090) in the University of Michigan 
Herbarium, this collection differs aligntly in having pedi- 
cels up to 2.5 mm. long and somewhat larger, less conspicu- 
ously veined leaves. 


EUGENIA GENTLEI Lundell, Carnegie Inst. Wasn. Publ. 478: 
216. 1937. 

BRITISH HONDURAS: Belize District, one mile north of Gra- 
cie Rock on the Sibun River, in pine ridge, Sept. 21, 19%, 
Hugh O'Neill 8761. Stann Creek District, Melinda Pine Ridge, 
in pine lands, a small tree, flowers white, Jan. 25, 1937, 
Percy H. Gentle 1887. -- From these additional collections 
the originsl description is amplified as follows: 

Flowers sessile, solitary or geminate, borne on older 
branchlets; calyx glabrous, limb 4-lobed, the lobes obl ong 
or ovate-oblong, 2 to 3.5 mm. long, rounded, ciliate, sparse- 
ly and rather finely glandular-punctate; petals obovate, 5 
to 7 mm. long, 4 to 4.5 mm. wide, rounded, long-ciliate, 
glabrous otherwise; stamens numerous; style glabrous; ovary 
2-celled; mature fruits unknown. 


ASPIDOSPERMA MATUDAI Lundell, sp. nov. 

Arbor 20-metralis, 60 cm. diam.; ramulis parce lepidotis; 
folia alterna, firme membranacea, glabra, longe petiolata, 
petiolo 3--3.8 cm. longo; lamina oblonga, 12--17.5 cm. lon- 
ga, 4--6 cm. lata, apice acuta vel obtusa, rary acuminata, 
basi late cuneata, utrinque prominenter costata, multiveno- 
sa; inflorescentiae 5.5--10 cm. longae (pedunculo incluso), 
longe pedunculatae, pedunculis lepidotis, ramulis et pedi- 
cellis adpresse albi-tomentulosis; pedicellis 4 mm. longis 
vel minoribus; calyx albi-tomentulosus, lobis lanceolatis, 
ca. 2.6 mm. longis; corolla ca. 7 mm. longa, extus glabra, 
tubo urceolato, ca. 4 mm. longo, intus sub apice puberulo, 
lobis subuleatis, ca. 3 mm. longis; ovarium glabrum. -- MEX- 
ICO: Chiapas, Escuintla, January, 1938, Eizi Matuda 2030 
(type in the University of Michigan Herbarium); vernacular 
name “chichi", -- A, Matudai differs notably from the other 
Mexican and Central American species in its large thin 
leaves with remarkably long slender petioles. It apvears to 
be related closely to A. cruentum ioodson, which is known 
only from fruiting material. 


MARKEA CAMPANULATA (Donn. Sm.) Lundell, comb. nov. 
Merinthopodium campanulatum Donn. Sm., Bot. az. 47: 257. 
1909. 


340 PHYTOLOGIA No. 10 


MARKEA INTERNEXA (Blake) Lundell, comb. nov. 
Merinthopodium internexum Blake, Contrib. Gray Herb. 52: 
85. 1917. 


MARKEA UNIFLORA Lundell, sp. nov. 

Frutex scandens; ramulis angulatis, rugulosis, hirsutis, 
crassiusculis; folia alterna, petiolata, petiolo sulcato, 
3--15 mm. longo; lamina chartacea vel subcoriacea, obovata 
vel elliptica, 4--13 cm. longa, 1.7--6.8 cm. lata, apice ob- 
tusiuscula vel abrupte breviterque subacuminata, basi obtus- 
juscula in petiolum attenuata, integerrima, subconcolor, 
supra glabra, subtus parce hirsute, venis lateralibus 5--8- 
jugis, supra impressis, subtus prominentibus; flores solit- 
arii; pedicelli graciles, parce hirsuti, 4.5--7 cm. longi; 
sepala 5, ovato-lanceolata, 3.7--4.2 cm. longa, basi usque 
ad 1.8 cm. lata, obscure trinervia, glabra; corolla usque ad 
7.5 cm. longa, parte tubiformi ca. 2 cm. longa, parte cam- 
paniformi ca. 3.5 cm. longa, lobis 5, ovatis, ca. 2 cm. lon- 
gis, rotundatis; filamenta ad basin villosa; antherae ex- 
sertae, ca. 1.4 cm. longae; stylus ca. 6 cm. longus; ovarium 
triloculare, multiovulatum, glabrum. -- MEXICO: Chiapas, 
Volcan de Tacana, alt. 2800 m., March 27, 1939, Bizi Matuda 
2816 (type in the University of Michigan Herberium). -- M. 
uniflora is a close relative of M. campanulata (Donn. Sm.) 
Lundell, & poorly known species described from incomplete 
material. The flowers of M. uniflora, solitary and apparent- 
ly terminal, ere considerably smaller than those described 


for M. campanulata. 


FOUR CENTRAL AMERICAN MELASTOMES 


H. A. Gleason 


CENTRADENIA GRANDIFOLIA var. BREVISEPALA Gleason, var. nov. 

Folia basi uno latere anguste cuneata altero late cunea- 
ta, secus venas albo-vittata. Sepala brevie truncata marg- 
ine minute ciliate. 

Type, Stuart 15, collected at Finca Pansamala, Dept. Al- 
taverapaz, Guatemala, in the herbarium of the University of 
Michigan. In the typical element of the species, .the sepals 
are ovate-triangular and 1.5-~-2 mm. long. 


1939 Gleason, Central American Melastomes 341 


LEANDRA LEPIDOTA Gleason, sp. nov. 

Frutex, caulibus junioribus, petiolis, venis, inflores- 
centia et floribus canescenti-lepidotis. Folia magna oblon- 
go-obovata S-pli-nervia. Hypanthium 10-sulcatum. Calycis 
limbus subhyalinus ad anthesin in lobos irregulares fissus; 
dentes exteriores lanceolati erecti. Petala lanceolata acu- 
ta. Antherae isomorphae semi-obovatae 2-loculares, poris 2 
dehiscentes; connectivum in appendicem lanceolatam product-. 
UM. 

Branching shrub 4 m. high, the younger stems, petioles, 
principal leaf-veins, inflorescence and flowers densely 
gray-lepidote with lanceolate scales 0.2 mm. long. Petioles 
stout, 1--3 cm. long. Blades unequal in each pair, obovate- 
oblong, 18--32 cm. long, 9--15 cm. wide, abruptly short- 
acuminate to an obtuse and mucronulate tip, obscurely re- 
pand, broadly cuneate at base, 5-pli-nerved, the innermost 
primaries alternate, 3--7 cm. from the base. Veins plane a- 
bove, elevated beneath; secondaries nearly straight, 4--6 
mm. apart; tertiaries prominulous beneath, obsolete above. 
Inflorescence terminal, the straight axis bearing several 
pairs of short lateral branches 5--8 mm. long; flowers 5- 
merous, sessile in terminal glomerules, subtended by lan- 
ceolate bracts 1.5--2 mm. long. Hypanthium subglobose, 
thick-walled, 2.8 mm. long to the torus, deeply but roundly 
10-sulcate. Calyx-tube erect, 0.6 mm. long; calyx-limb very 
thin and diaphanous, about 1.1 mm. long, sparsely stellete, 
splitting at anthesis into irregular triengular lobes; ex- 
terior teeth erect, lanceolate, acute, 1.6 mm. long. Fetals 
erect, 4.5 mm. long, more or less involute distally, the 
actual epex incurved, 0.2 mm. long, the exterior tooth er- 
ect, O.4 mm. long. Filaments erect, glabrous, flat, 2.4 mm. 
long. Anthers radially flattened, 1.6 mm. long; connective 
prolonged into a basal eppendage O.5 mm. long, lanceolete 
in the epipetalous series, obscurely 3-lobed in the episep- 
a@lous. Ovary wholly inferior, S-celled, its subconic summit 
glebrous; style glabrous, slender, 3 mm. long; stigma trun- 
cate. 

In rain forest at Ville Quesada, Prov. Alajuela, Costa 
Rica, alt. 240 meters, Austin Smith H-1735 (type in the 
Britton Herbarium eat The New York Botanical Garden). Relat- 
ed to L. lasiopetala Cogn., also of Costa Rica, which is 
glandular-pubescent rather than lepidote. 


MICONIA PANAMENSIS Gleason, sp. nov. 

Sect. Bumiconia: Rami juniores minutissime furfuracei 
demum glabrescentes. Petioli crassi 4 mm. longi. Laminee 
ovato-lanceolatae, usque ad 16 cm. longae 6 cm. lateae, lon- 
ge acuminatae, inferne rotundatae ad basin cordulatam, 3- 
nerviae, supra glebrae subtus ad venas majores minutiss ime 


342 EA. YT OL OG aA No. 10 


stellatae. Inflorescentia pyramidelis, remis gracilibus, in- 
ferioribus elongatis; bracteolae triangulares minimae. Flor- 
es sessiles S-meri. Hypanthium campanulatum, ad torum 2 mm. 
longum, stellato-furfuraceum. Calycis tubus 0.4 mm. longus 
suberectus; sepala late triangulsria 0.6 mm. longa; dentes 
exteriores conici vix prominuli. Fetala alba elliptica equi- 
latera integra 2.6 mm. longa. Stemina dimorpha. Filementa 
gracilia glabri 3.5 vel 2.7 mm. longa. Antherae lineares 
subrectae 2-loculares poro ventrali-terminali dehiscentes, 
%,4--3.7 vel 2.6--2.8 mm. longae; connectivum in ser. ext. 
0.8 mm. infra thecas productum, in lobos erosos 2 laterales 
deflexos 1 dorsalem divisum, in ser. int. 0.6 mm. productum, 
in lobos parvos integros 2 laterales 1 dorsalem productun. 
Ovarium 3-loculare, semi-inferum, ovulis in quoque loculo 
paucis adscendentibus; stylus glaber 6 mm. longus; stigma 
capitellatum. 

Type, iledel 6, from Maccaw Hill, Frov. Bocas del Toro, 
Panama, in the Britton Herbarium. Related to a small group 
of South American species, of which M. Chamissois Naud. is 
best known, in which the connective of the outer series of 
stamens is expanded into a large cordate structure. Within 
this group, our species most resembles the Peruvian M. 
longiracemosa Gl., which (as also M. Chamissois) has pli- 
nerved leaves narrowed to the base. 


BLAKEA CALYCOSA Gleason, sp. nov. 

Frutex epiphyticus, ramis supremis obscure furfuraceis. 
Petioli crassi, 18--4O mm. longi tenuiter furfuracei. Lamin- 
ae membranaceae ellipticae, usque ad 24 cm. longae 9 cm. 1a- 
tae, breviter caudato-acuminetae, basi rotundsatae, subpelta- 
tae, 3-nerviae jugo submarginali neglecto, supra glabrae, 
subtus ad venas furfuraceae; venae secundariae alternatim 
validae debiles. Flores 6-meri, pedicellis 5 mm. longis. 
Bracteae exteriores subpatulae rotundae, extus pubescentes, 
11 mm. longae 14 mm. latae, 4 mm. connatae, 3-nerviae, intra 
glebrae; interiores basi hypanthium amplectentes glabrae, 5 
mm. connatae, superne patulae foliaceae, limbo lunato medio 
5--6 mm. longo. Hypanthium hemisphaericum 5 mm. altum 8 mn. 
latum glabrum. Sepala horizontaliter patula ovato-oblonga, a 
toro 6--7 mm. longa, glabra, apice rotundata appendiculata, 
appendice foliacea valde reflexa pubescente 7 mm. longa. 
Petala rosea srathulata obtusa basi late unguiculata, 22 
mie longa, multinervia, utrinque verrucosa, latere uno spar- 
se villoso-ciliata, altero minutissime reflexo-ciliata. Fil- 
amenta lata recta 5 mm. longa. Antherae late ellipticae ob- 
tusae, 6 mm. longae, poris 2 terminalibus, connectivo in 
carinam dorsalem elevato, basi in calcarem late conicum ob- 
tusum 2 mm. longum producto. Ovarium inferum 6-loculere, 
summo depressum circa styli basin elevatum. Stylus 14 mm. 


} 
q 
; 
F 


OO ee a 
‘- ‘ 


1939 Moldenke, Notes on Eriocaulaceae 343 


longus infra medium glanduloso-hirsutulus; stigma punctifor- 
me. 

Type, Allen 1788, from the vicinity of #1 Valle, Prov. de 
Coclé, Panama, alt. 600--1000 m., in the Britton Herbarium. 
In the foliaceous reflexed appendages of the sepals it is 
quite unlike any other known species of Blakea. 


ADDITIONAL NOTES ON THE ERIOCAULACKAE -- II 


Harold N. Moldenke 


The following notes constitute a continuation of the 
notes published by me in Phytologia 1: 309--336. 


1. SYNGONANTHUS FLAVIDULUS (Michx.) Ruhl. 

The species has been collected in anthesis from February 
to June and in September. Curiously, Ruhland placed the name 
Eriocaulon flavidulum Michx. in two places: (1) under Syngo- 
nanthus flavidulus, for which it is the name-bringing syno- 
nym, and also (2) as a valid species of Eriocaulon [imgl. 
Pflanzenreich 439: 43 & 256. 1903]. For the latter he gives 
as distribution "along banks of streams from Pennsylvania to 
the Carolinas". Just what he meant by this latter species is 
doubtful. 

Illustrations: Britton & Br. Ill. Fl., ed. 1, 1: f. We. 
1896; ed. 2, 1: 455, f. 1144. 1913; Small, Man. SE. Fl. 257. 
1933. 

Citations: NORTH or SOUTH CAROLINA: County undetermined: 
Michaux s.n. (N--photo of type, P--type). SOUTH CAROLINA: 
Florence Co.: Ravenel s.n. [Florence, 1879] (Mi, T); Beau- 
fort Co.: J. H. Mellichamp sen. [Bluffton] (Be). GEORGIA: 
Charlton Co.: F. Harper sen. [Floyd's Island] (N); J. S. 
Harper 161 (N); Emanuel Co.: R. M. Harper 803 (N). FLORIDA: 
Walton Co.: Mohr sen. [De Funiac Springs, May 12, 1892] (C); 
Franklin Co.: Biltmore Herb. 3162a (N); Leon Co.: Berg sen. 
{Near Tallahassee] (N); Jefferson Co.: Lighthipe s.n. [Mont- 
icello, March, 1891] (N); Alachua Co.: S. J. Knight 2 (C); 


O'Neill 611 (I), 652 (I); Duval Co.: Curtiss 41 (Cc); T. 
Hoge 8 (C); Lighthipe 173 (N); Clay Co.: W. M. Canby s.n. 
[Hibernia, March, 1869] (N); C. Skottsberg s.n. [Below Jack- 
sonville & Atlantic Beach] es Johns Co.? M. C. Rey- 
nolds sen. (Mar.--July, 1875] (C); Lake Co.: A. S. Hitchcock 
aon. [Mustis, June--July, 1894] (I); Nash 14 y (C, Mi), 1944 
(C); Volusia Co.: Lugren sen. [Volusia] (NN); Orange Co.: F. 
S. Blanton 6491 (1); Huger sen. [Winter Park, April, 1900] | 


344 PHYTOLOGIA No. 10 


(N); BE. J. Palmer 38,324 (N); Pasco Co.: O'Neill 7788 (1), 
7875 (I), sen. [Fish Lake, Feb. 20, 1926] (I), sen. [Big Cy- 
press Swamp, April, 1927] (1); Hillsboro Co.: Leavenworth s. 
n. [Tampa Bay] (T); Pinellas Co.: Tracy 6642 (N=-2), 7588 ae 
(N); Polk Co.: Topping 2609 (N); Brevard Co.: Edw. Palmer 
580 (Bc); Manatee Co.: Garber sen. [Manatee, Apr. 1876] (C); 
Tracy 6643 (N); Lee Co.: H. N. Moldenke 688 (N), 940 (N); 
Sheehan s.n. [Boakum Place, March 6, 1919] (N); J. P. Stand- 
ley 10 (N); County undetermined: Chapman 182 (T), s.n. (C, T 
--2), ALABAMA: Mobile Co.: Pennell 4509 (ii). LOCALITY OF 
COLLECTION UNDETERMINED: Gibbes s.n. [Sept. 1833] (N). 


8. SYNGONANTHUS HONDURENSIS Moldenke, sp. nov. 

Herba parva; foliis caespitosis patentibus 1--1.4 cm. 
longis lineeribus; pedunculis 4 vel 5 adpresso-pubescenti- 
bus, vaginis quam foliis brevioribus dense breviterque pub- 
escentibus; brecteolis ad mediam olivaceis, unque ad 2.5 mm. 
longis, glsbris. 

Small herb, 6--7 cm. tall; Leaves caespitose, linear, ol- 
ivaceous, pétent, not noticeably recurved nor appressed, l-- 
1.6 cm. long, about 1 mm. wide, rather blunt at apex, glab- 
rate on both surfaces; sheaths closely appressed, shorter 
than the leaves, densely short-pubescent, acuminate at apex; 
neduncles 4 or 5 per plant, very slender, obscurely sulcate, 
slightly twisted, conspicuously white-pilose throughout with 
usually closely appressed hairs or sometimes also a few 
spreading ones toward the apex; heads solitary, 2--6 mm. 
wide; involucral bractlets elliptic, membranous, olivaceous 
in a median band from apex to base, gray and scarious at the 
margins, 1.6--2.5 mm. long, about 0.875 mm. wide, regularly 
nerrowed from the middle to the sharply acute or scuminate 
apex, glabrous, not ciliate; staminate florets: pedicels a- 
bout 1.25 mm. long, glabrous; sepals 3, free, membranous, 
with an olivaceous median band, elliptic, about 1.125 mn. 
long, sharply acute at apex, glabrous, not ciliate; petals 
united to form a tube about as long as the sepals, olivace- 
ous throughout to about the middle, olivaceous only along 
the midrib above, glabrous; pistillate florets: pedicels a- 
bout 1 mm. long, densely long-villous at base with white 
hairs longer than the pedicel; sepals 3, free, membranous, 
with an olivaceous median band, lanceolate, about 1.875 mm. 
long, sharply acute at anex, long-ciliate along the margins, 
glabrous; petals loosely connate along the margins, narrow- 
oblong, slightly shorter than the sepals, densely long- 
villous on the outer surface with wnite ascending hairs. 

The type of this species was collected by Rev. Hugh 
O'Neill (No. 8543) in a trovical pineland, 3 miles west of 
Boomtown, British Honduras, on August 23, 1936, and is dep- 
osited in the herbarium of the University of Michigan. It 


; 
: 
4 
P 


(N); C. Wright 3237 (N--Laotype, ja; wnt an, 


1939 Moldenke, Notes on Eriocaulaceae 345 


was mixed with No. 8543a, Syngonanthus Oneillii, in the ori 
inal collection. — 

BRITISH HONDURAS: O'Neill 8543 (I--isotype, Mi--type, N-- 
isotype). 


4, SYNGONANTHUS INSULARIS Moldenke, N. Am. Fl. 19: 45, 1937. 
The species inhabits white sand and has been collected in 
anthesis in February. . 
Citations: CUBA: Isla de Pinos: Britton, Britton, & Wils- 
on 14,162 (N--type); Ekman 12 12,095 (Mi). *e 


3. SYNGONANTHUS LAGOPODIOIDES (Griseb.) Ruhl. in Urb. Symb. 
Ant. 1: 489. 1900. 

Paepalanthus lagopodioides Griseb. Cat. Pl. Cub. 225. 
1866; Paepalanthus lagospodioides Griseb. ex Sauv. Fl. Cub. 
165, sphalm. 1871. 

The species inhabits sandy and moist pinelands, plains, 
moist savannas, streamsides, moist or wet sand along reail- 
road tracks, white sand, or wet black soil, and hes been 
collected in anthesis from December to March, May, August, 
and September. 

Citations: CUBA: Pinar del Rf{o: Britton, Britton, srle, 
+s Gager 6488 (N); erties, Britton, & Sa er 7058 (N), 7254 

Ekman 10,823 (Mi, N), veo 11,061 (Mi, 11,145 (i) te = 
Ay Roca a 6930 (N); Shafer 10,866 866 (N--2), Stina), 11,782 


8 Isla 
- Pinos! Britton, Britton, & Wilson 14,21 (N), i yee (N), 
15,767 (N), 15,78 15,789 ( (N); Ekman 1 11,973 (N), 12,071 


9. SYNGONANTHUS LUNDELLIANUS Moldenke, sp. nov. 

Herbe parva; foliis caespitosis 1--1.5 cm. longis patent- 
ibus linearibus; pedunculis pluribus pubescentibus, pilis 
hirsuto-patentibus, vaginis quam foliis longioribus, laxe 
longeque pilosis; bracteolis plusminus olivaceis, usque ad 8 
mm. longis, villosis. 

Small herb, 6.5--8.5 cm. tall; leaves caespitose, linear, 
olivaceous, 1--1.5 cm. long, patent, not noticeably recurved 
nor appressed, rather blunt at apex, appressed-puberulent on 
both surfaces; sheaths closely appressed, conspicuously sur- 
passing the leaves, loosely long-pilose, acute or subacumin- 
ate at apex; peduncles numerous, very slender, few-sulcate, 
streight (not twisted), conspicuously pilose throughout with 
loosely spreéding (hirsute) elongated hairs; heads solitary, 
2--15 mm. wide; involucral bractlets linear-oblong, often e- 
longate to 8 mm. long (4), the outer ones much shorter or 
all much shorter, the short ones olivaceous in a median band 
from base to apex, the elongated ones olivaceous in several 
parallel bands from base to apex, acute at apex, conspicu- 
ously villous on the outer surface with long loose hirsute 


346 PHYTOLOGIA No. 10 


capitate hairs, glabrous within; staminate florets: pedicels 
about 0.75 mm. long, densely long-villous-tomentose at base 
with white hairs longer than the pedicel; sepals 3, free, 
hyaline, elliptic, about 1 mm. long, acuminate at both ends, 
glabrous, not ciliate; petals united into a narrow nyaline 
glabrous tube almost as long as the sepals; pistillate flor- 
ets: pedicels about 0.5 mm. long, densely lonz-villous with 
white hairs longer than the pedicel; sepals 3, free, nyal- 
ine, elliptic, about 1.625 mm. long, regularly narrowed to 
the acute or acuminate apex, glabrous, not ciliate; petals 
narrow-oblong and strap-shaped, shorter than tne sepals, hy- 
aline, anpressed-villous on the back, loosely connate by 
their margins. 

The type of this species was collected by ev. Hugh 
O'Neill (No. 8546) under Quercus oleoides var. australia 
Trelease, Broken Ridge, 8 miles northeast of Boomtown, Brit- 
ish Honduras, on September 17, 1936, and is deposited in the 
herbarium of the University of Michigan. Vivipary is so pro- 
nounced on the Catholic University isotype that young plants 
are actually in flower in the inflorescence-heads of the old 
plants! The species is named in honor of my good friend and 
esteemed colleague, Dr. Cyrus Longworth Lundell, who is do- 
ing such noteworthy and important work on the natural hist- 
ory of the Yucetén Peninsula. 

BRITISH HONDURAS: O'Neill 8546 (I--isotype, Mi--type, N-- 
isotype). 


6. SYNGONANTHUS ONEILLII Moldenke, sp. nov. 

Herba parva; foliis caespitosis 1--2.5 cm. longis plus- 
minus recurvatis; pedunculis pluribus glabris vel subglab- 
ris; bracteolis olivaceis. 

Smell herb, 6--10 cm. tall; leaves caespitose, linear, 
olivaceous, spreading and more or less recurved, 1--2.5 cm. 
long, O.5--1l mm. wide, abruptly acute at apex, glabrate on 
both surfaces or somewhat loosely pilose; sheaths closely 
appressed, equaling or shorter then the leaves, very sparse- 
ly and loosely pilose or glabrate, sherply acute or acumin- 
ate at apex; peduncles numerous, very slender, sulcate, 
twisted, glabrous or subglabrous throughout; heads solitary, 
3--5 mm. wide; involucrel bractlets oblanceolate-elliptic, 
rather light-olivaceous or browmish throughout, lighter a- 
long the margins, very abruptly acute or obtuse at apex, 
glabrous; staminate florets: pedicels about 0.875 mm. long, 
glabrous except for a few long hirsute white hairs at the 
base surpassing the pedicel; sepals 3, 1--1.125 mm. long, 
oblanceolate, about 0.3575 mm. wide at the widest part, glab- 
rous, pleinly cellular, hyaline, short-acuminate or sinuate- 
tridentate at apex, cuneate-nearrowed to the base, free; 
petals united at the middle, free at apex and base, slightly 


1939 Moldenke, Notes on Eriocuulaceae 347 


shorter than the sepals, olivaceous in a median band from a- 
pex to base, incurved and pilose at the very apex on the 
outer surface with a few black hairs; pistillate florets: 
pedicels about 0.75 mm. long, long-hirsute at base; sepals 
about 1.25 mm. long, lanceolate-ovate, acute st apex, long- 
ciliate on the marzins toward the apex; petals narrow- 
oblong, connate by their margins. 

The type of this species was collected by my good friend, 
Rev. Hugh O'Neill (No. 8548) -- in whose honor it is named 
-- in & trovical pineland, Boomtown, British Honduras, on 
September 14, 1936, and is deposited in the herbarium of the 
University of Michigan. : 

BRITISH HONDURAS: O'Neill 8543e (I, Mi), 8548 (I--iso- 
type, Mi--type). 


The new species from British Honduras, described above, 
may be distinguished by means of the following artificial 
key: 

1. Peduncles glabrous or practically 30.........5. Oneillii. 

la. Peduncles conspicuously pubescent. 

2. Involucral bractlets hysline, completely colorless..... 
S. Bartlettii. 
2a. Involucral bractlets more or less olivaceous. 

3. Sheaths shorter than the leaves, densely short- 
pubescent; pubescence on peduncles mostly appressed; 
brectlets to 2.5 mm. long, glabrous...S. hondurensis. 

Ja. Sheaths lonzer than the leaves, loosely lLong-pilose; 

pubescence on peduncles spreading; bractlets to 8 
mm. Long, VillOUS...ceeeeeeeveeeeeeede Lundellianus. 


_ 


5. SYNGONANTHUS WILSONII Moldenke, N. Am. Fl. 19: 45. 1937. 
The species inhabits white sand and has been collected in 
anthesis in Merch. 
Citations: CUBA: Isla de Pinos: Britton, Britton, & Wils- 


on 15,789a (N--type). 


TONINA Aubl. Hist. Pl. Guisn. Fr. 2: 856. 1775. 
Hyphydra Schreb. Gen. 666. 1791. 
A monotypic genus of tropical portions of the New World. 


1. TONINA FLUVIATILIS Aubl. Hist. Fl. Guian. Fr. 2: 857, pl. 
530. 1775. 

Hyphydre amplexicaulis Vahl, Symb. Bot. 3: 99. 1794 [ac- 
credited to Schreber by Ruhlend!]; Eriocaulon amplexicaule 
(Vahl) Rottb. Descr. Fl. Surinam. 7, pl. 1, f. 1. 1798. 

The species inhabits quiet water of marshes, swamps, 
ponds, and streams, and has been collected in anthesis in 
every month of the year. 

Illustrations: Aubl. Hist. PL. Guian. Fr. 4: pl. 330. 


348 PHYT:? O'L°O GA No. 10 


1775; Acta Lit. Univ. Hafn. 1: pl. 1, f. 15 Lam. Encycl. 
Méth. Ill. pl. 772, f. 1. 1798; Rottb. Descr. Pl. Surinam. 
pl. 1, f. 1. 1798; Ann. Sci. Nat. Paris, sér. 1, 13: pl. 5, 
f. 4. 1828; Nov. Act. Physico-med. Acad. Caes. Leopold.- 
Carol. Nat. Cur. 17’: pl. 4, f. 2. 1835; Schnitzlein, Icono- 
gre 1: pl. 46, f. 13--19. 1845; Lindl. Veg. Kingd. 122, f. 
82. 1846; Mart. Fl. Bras. 3): pl. 38, f. 1. 1863; Rngl. & 
Prantl, Nat. Pflanzenfam. 2%: 24, f. 14. 1888; Bot. Tidsskr. 
18: pl. 20, f. A. 1893; Baill. Hist. Pl. 12: 399, f. 372-- 
374. 1894; Engl. Pflanzenreich 4°°: 239, f. 35. 1903. 
Citations: MEXICO: Veracruz: Orcutt 3285 (F). BRITISH 
HONDURAS: Schipp 693 (Mi, N). COSTA RICA: San José: Skutch 
2465 (Mi, N). CUBA: Pinar del Rfo: Acuna 10,753 (Es); Ekman 
11,120 (N); Province undetermined: C. Wright 3242 (N, T). 


Geographic distribution of the species and varieties of 
Eriocauleceae thus far cited in these notes: 


ISLE OCF SKYE, IRELAND, & SCOTLAND: Eriocaulon septangulare. 

SAINT PIERRE, SABLi& ISLAND, & NEWFOUNDLAND: Eriocaulon sept- 
angulare. 

NOVA SCOTIA: Eriocaulon septangulare [Cape Breton, Richmond, 
Guysborough, Halifax, Digby, « Yarmouth]. 

NEW BRUNSWICK: Zriocaulon septangulare [St. John]. 

QUEBEC: Eriocaulon septangulare |Rimouski, Temiscouata, 
Fortneuf, Montcalm, & Ottawa]. 

ONTARIO: Eriocaulon septangulare [Simcoe & St. Joseph Is- 
land]. 

MAINE: Eriocaulon Parkeri [Penobscot &« Sagadahoc] -- E. sep- 
tangulare [Penobscot, Waldo, & Mount Desert Island]. 

VERMONT: Eriocaulon septangulare [Addison & Orange]. 

NEW HAMPSHIRE: Eriocaulon septangulare [Coos, Grafton, Mer- 
rimack, Cheshire, & aii Teborel. 

MASSACHUSETTS: Eriocaulon septangulare (Berkshire, Hamp- 
shire, Hampden, Essex, Middlesex, Norfolk, Flymouth, 
Barnstable, Dukes, & Nantucket]. 

CONNECTICUT: Eriocaulon Parkeri [Fairfield & New London] -- 
E. septangulare [Litchfield & Middlesex]. 

NEW YORK: Eriocaulon Parkeri [Greene, Columbia, & Rockland] 
-- E. septangulaere (St. Lawrence, Oswego, Hamilton, Madi- 
son, Warren, Washington, Rensselaer, Steuben, Sullivan, 
Ulster, Dutchess, Orange, Putnam, Westchester, Queens, 
Nassau, & Suffolk]. 

PENNSYLVANIA: Eriocaulon Parkeri [Philadelphia] -- E. sept- 
angulare [Luzerne & Pike]. 

NEW JERSEY: Eriocaulon compressum [Burlington, Ocean, Cam- 
den, Atlantic, & Cape May] -- E. decangulare [Monmouth, 
Burlington, Ocean, Camden, Atlantic, Cumberland, & Cape 
May] -- E. Parkeri [Mercer, Monmouth, Burlington, Ocean, 


Se 


1939 Moldenke, Notes on Eriocaulaceae 349 


Camden, Atlantic, & Cumberland] -- E. septangulare [Sus- 
sex, Morris, Ocean, Camden, & Atlantic]. 

DELAWARE: Eriocaulon decangulere [Sussex] -- E. Parkeri 
[Sussex]. 

MARYLAND: Eriocaulon compressum [Baltimore & Wicomico] -- E. 
decangulare [Prince Georges &« Worcester] -- E, Parkeri 
[Wicomico] -- &. septangulare [Harford, Wicomico, & Wor- 
cester ]. . 

DISTRICT OF COLUMBIA: Eriocaulon decangulare. 

VIRGINIA: Eriocaulon decangulare [Prince George] -- E. Park- 
eri [Alexandria & New Kent] -- Lachnoceaulon anceps [Isle 
of Wight, James City, & Prince George]. 

NORTH CAROLINA: Eriocaulon compressum [Onslow] -- E. decang- 
ulare (Catawba, Rowan, Dare, Buncombe, Cumberland, Bruns- 
wick, & New Hanover] -- E. lineare [Henderson] -- Lach- 
nocaulon anceps [Bladen & New Hanover] -- L. minus [Heli- 
fax & New Hanover]. 

SOUTH CAROLINA: Eriocaulon compressum [Lexington & Charles- 
ton] -- E. decangulare [Orangeburg] -- E. Ravenelii 
[Berkeley] -- Lachnocaulon anceps [Kershaw, Florence, Ai- 
ken, & Dorchester] -- Syngonanthus flavidulus {Florence « 
Beaufort ]. 

GEORGIA: Eriocaulon compressum [Sumter, Bryan, & Charlton] 
-- E. compressum var. Harperi [Sumter & Charlton] -- E. 
decangulare [Columbia, Sumter, Lee, McIntosh, « Lowndes] 
-- E. lineare [Montgomery, Bulloch, & Lowndes] -- Lachno- 
caulon anceps [Richmond, Sumter, Emanuel, & Worth] -- L. 
Beyrichianum [Charlton] -- L. minus [Lowndes] -- Syngo- 
nanthus flavidulus [Emanuel & Charlton]. 

FLORIDA: Eriocaulon compressum [Duval, Clay, St. Johns, Le- 
vy, Lake, Orange, Pasco, Hillsboro, Polk, De Soto, Lee, 
Broward, Seminole, Dade, & Collier] -- E. compressum var. 
Harperi [Jackson, Washington, Franklin, Leon, Wakulla, De 
Soto, & Lee] -- EB. decangulare [Holmes, Franklin, Leon, 
Duval, Marion, Highlands, Charlotte, Lake, Orange, Pasco, 
iilleboro, Polk, Brevard, Nanatee, Lee, Broward, & Dade] 
-- E. decangulare var. latifolium [Franklin] -- 5. line- 
are [Santa Rosa, Walton, Gulf, Leon, & Pasco] -- E. Rav- 
enelii [Broward & Dade] -- Lachnocaulon anceps [Alachua, 
Nassau, Duval, Clay, Lake, Okeechobee, Orange, Pasco, 
Hillsboro, Polk, Manatee, Seminole, & Monroe] -- L. Bey- 
richianum [St. Johns] -- L. eciliatum [Walton, Putnam, & 
Lake] -- L. Rngleri [Washington, Putnam, Lake, Pasco, & 
Orange] -- L. floridanum [Lake] -- L. glabrum [Lee, Brow- 
ard, & Dade] -- L. minus [Walton, Liberty, Leon, Clay, 
Lake, Volusia, Pasco, & Broward] -- Syngonanthus flavidu- 
lus [Walton, Franklin, Leon, Jefferson, Alachua, Duval, 
Clay, St. Johns, Lake, Volusia, Orange, Pasco, Hillsboro, 
Pinellas, Polk, Brevard, Manatee, & Lee]. 


350 PHYTOLOGIA No. 10 


ALABAMA: Eriocaulon compressum [Mobile & Escambia] -- E. 
compressum ver. Harperi [Mobile] -- E. decangulare [Mob- 
ile & Baldwin] -- &. decangulare var. latifolium [Mobile] 
-- E. lineare [Baldwin] -- Lachnocaulon anceps [De Kalb & 
Mobile] -- L. digynum [Mobile] -- Syngonanthus flavidulus 
[Mobile]. 

MISSISSIPFI: Eriocaulon compressum var. Harperi [Herrison] 
-- &, decangulare (Herrison] -- Lechnocaulon anceps [Har- 
rison & Jackson] -- L. digynum [Harrison]. 

INDIANA: Eriocaulon septangulare [Steuben & Whitley]. 

MICHIGAN: Eriocaulon septangulare [Isle Royale, Houghton, 
Iron, Marquette, Alger, Chippewa, Cheboygan, Presque 
Isle, Mason, Gratiot, &« Cass]. 

WISCONSIN: Eriocaulon septangulare [Burnett & Calumet]. 

MINNESOTA: Eriocaulon septangulare [St. Louis, Cook, Crow 
Wing, & Chisago]. 

OKLAHOMA: Eriocaulon Kérnickianum [Pushtamaha]. 

ARKANSAS: Eriocaulon Kérnickianum [Benton & Logan]. 

LOUISIANA: Eriocaulon compressum [Calcasieu] -- E. compress- 
um var. Harperi (St. Tammany ] -- E, decengulare ({Natchi- 
toches, Calcesieu, St. Tammany, & Orleans] -- Lachnocaul- 
on anceps [St. Tammany & Orleans]. 

TEXAS: Briocaulon decangulsre {Smith & Waller] -- E. texense 
[Smith] -- Lachnocaulon anceps [Hardin]. 

CALIFORNIA: Eriocaulon microcephalum [Kern]. 

MEXICO: Eriocaulon Benthami (Chihuahua, Zacarecas, Jalisco, . 
& Mexico] -- E. bilobatum [Jalisco] -- E. Ehrenbergianum 
(Chihuahua, Neyarit, Jalisco, Hidalgo, Mexico, Federal 
District, & Oaxaca] -- E. guadalajarense (Jalisco] -- E. 
mexicanum [Jalisco] -- E. microcephalum [Mexico & Puebla] 
-- E. Palmeri [Durango] -- E. Pringlei [Chihuahua] -- 5. 
Schiedeanum [Jalisco] -- E. tepicanum [Nayarit] -- Faep- 
alanthus chiapensis [Chiapas] -- P. iellii [Veracruz] -- 
Tonina fluviatilis [Veracruz]. 

BRITISH HONDURAS: Eriocaulon fuliginosum -- & Kinlochii -- 
E. Schippii -- & Williamsii -~ Paepalanthus Gentlei -- 
P, Lamarckii -- Synzonanthus Bartlettii -- S. hondurensis 
-- S. Lundellianus -- S. Oneillii -- Tonina fluviatilis. 

GUATEMALA: Eriocaulon Ehnrenbergianum {Chimaltenango]. 

COSTA RICA: Eriocaulon microcephalum (San José] -- Paepalan- 
thus costaricensis [San José] -- Syngonanthus caulescens 
-- Tonina fluviatilis [San José]. 

PANAMA: Erioceulon panamense [Chiriqu{] -- E. Seemannii 
(Canal Zone & Panama] -- & iilliameii [Coclé] -- Peepal- 
anthus Lamarckii [Coclé]. 

CUBA: Hriocaulon arenicola [Isla de Pinos] -- E. cubense 
[Isla de Pinos] -- E. dioecum [Pinur del Rfo] -- E. ech- 
inospermoideum [Santa Clara] -- E. echinospermum (Pinar 
del Rfo] -- &. Exmannii [Pinar del Rio] -- & fuliginosum 


ee eee ee 


1939 Moldenke, Notes on Eriocaulaceae 551 


{Pinar del Rfo, Isla de Pinos, Santa Clara, & Oriente] -- 
E. fusiforme [Isla de Pinos] -- E. heteropetalum [Pinar 
del Rfo] -- E. inaulare [Pinar del Rfo & Isla de Pinos] 
-- E. lacustre [Pinar del Rfo] -~ E, melanocephalum [Ori- 


ente] -- E, melanocephalum var. longipes [Piner del Rfo] 
-- &. minutissimum [Pinar del Rfo] -- E. miserrimum [Isla 
de Pinos] -- E. olivaceum [Isla de Pinos] -- E, ovoideum 


(Isla de Pinos] -- E. pinarense [Pinar del Rf{o & Isla de 
Pinos] -- &. pseudocompressum [Pinar del Rfo] -- &. scle- 
rocephslum [Isla de Pinos] -- E. sigmoideum -- Lachnocau- 
lon anceps [Isla de Pinos] -- L. cubense [Santa Clara] -- 
L. Ekmannii [Pinar del Rfo] -- Paepalanthus alsinoides 
[Pinar del Rio] -- P. alsinoides var. minimus [Piner del 
R{o & Isla de Pinos] -- P. Lamarckii [Pinar del Rfo & Is- 
la de Pinos] -- P. montanus [Oriente] -- P. pungens [Ori- 
ente] -- P. retusus [Piner del Rfo] -- P. riparius [Ori- 
ente] -- P, seslerioides [Pinar del Rfo, Isla de Pinos, & 
Oriente] -- Syngonanthus androsaceus [Pinar del Rfo] -- 
S. insulsris [Isle de Pinos] -- S. lagopodioides {Pinar 
del Rfo & Isla de Pinos] -- S. Wilsonii [Isla de Pinos] 
-- Tonina fluvietilis [Pinar del Rfo]. 
HISPANIOLA: Paepalanthus domingensis [Dominican Republic] -- 
P. Tuerckheimii [Dominican Republic]. 
TRINIDAD: Paepalanthus Lamarckii. 
HCUADOR: Eriocaulon microcephalum (Carchi & Pichincha]. 


An alphabetized list of citations of herbarium material in 
the foregoing notes: 


Acuna 10,753 (Tonina fluviatilis); Allen, T. se S.ne 
[Pine barrens, June, 1871] (Eriocaulon compressum), s.n. 


[1870] (Briocaulon decangulare); Allison 207 (Eriocaulon 
deca Lere); Ames, Gs L., sen. (Mt. Luber, W., Sept. 1857] 
(Eriocaulon septangulare); André K.1747 (Eriocaulon micro- 
cephalum); Anselm 435 (Eriocaulon septangulare); Arsene 130 
(2riocaulon septangulare); Austin sen. [Hast Mass., 1860] 


(Zriocaulon septanculare). 

Bailey, Lt., sen. (West Point] (Zriocaulon septangulare); 
Baker, “tthe 0 aortoceulon decangulare), Ot gugomnceul- 
on anceps), 1557 (Eriocaulon com ressum), 1558 (Eriocaulon 
compressum var. Har eri), @en. Liobile, 7/20/1897] (Brio- 
caulon decangulare), sen. [Biloxi, 7-25-1897] (Eriocaulon 
decangulare); Barnes & Land 159 (Eriocaulon Schiedeanum ); 
Bartlett, H. H., 313 (Eriocailon septangulere), 11,263 (Pae- 


elanthus Lamarckii), 11,263a (Syngonanthus Bartlettii), 
11,670 (Syngonanthus Bartlettii, type coll.), 11,874 (Paep- 
alanthus Gentlei); Bartley & Pontius 503 (Eriocaulon decang- 
ulare); Bartram 2277 (Lachnocaulon anceps); Bassett s.n. 

Hammonton] (Eriocaulon compressum), sen. [Atco] (Hriocaulon 


352 PHYTOLOGIA No. 10 


decangulare), s.n. [Atco, July 19, 1923] (Eriocaulon decang- 
a ieene Batchelder s.n. (Merrimack, Aug. 4, 1917] (Eriocaul- 


on septangulare); Beals s.n. [Hammonton Lake, July 1, 1922] 
(Zriocaulon compressum), s.n. [Hammonton Lake] (Eriocaulon 
septangulare); Beechey s.n. [Jalisco] (Eriocaulon hrenberg- 
eae Berg s.n. [Near Tallahassee] (Hriocaulon decangula- 
re), sen. [Near Tallanassee] (Lachnocaulon minus), sen. 
[Near Tallahassee] (Syngonanthus flavidulus); Beringer sen. 
[Atco, June, 1895] (Hriocaulon compressum), aen. [Atco, Aug. 
1890] (Eriocavlon decangulare); Berlandier 760 (Zriocaulon 
Bhrenbergianum, cotype coll.); Beyrich s.n. [Amer. bor. ] 
(Lachnocaulon anceps); Bicknell 230 (Hriocaulon septengula- 
re), 231 (Eriocaulon septanculare), s.n. [Rockville Center, 
July 25, '03] (Eriocaulon septangulare); Biltmore Herb. 
2296a (Eriocaulon compressum var. Harperi), 2755 (Lachno- 
caulon anceps), 2755a (Lachnocaulon anceps), 3162a (Syngon- 
anthus flavidulus), 3865a (Eriocaulon lineare), 3867a (Hrio- 
caulon decangulare), 3867b (Eriocaulon decangulare), 3867c 
(Eriocaulon decangulare), 3867d (Eriocaulon decangulere), 
3867f (Briocaulon decangulare), 15,001d (Lachnocaulon ecili- 
atum); Bisky sen. [Lakeville] (Hriocaulon septan ulare); 
Blake, S. F., 4366 (Eriocaulon septangulare), 10,665 (srio- 
caulon decangulare); Blanton, F. S., 6491 (Syngonanthus 
flavidulus); Blanton, 0., 86 (Eriocaulon decangulare); 
Blodgett s.n. [Martha's Vineyard, Aug. 19, '93] (Eriocaulon 
septangulare), 8eNe ikey West] (Lachnocaulon anceps); Boykin 
12 (Lachnocaulon anceps), sen. (Eriocsulon decangulare); 
Breinerd s.n. [Lake Dunmore, Aug. 16, 1878] (Hriocaulon sep- 
tangulare); Briggs sen. [Chemo Stream, Aug. 1891] (Eriocaul- 
on septangulare); Brinton a.n. [Pine barrens, Sept. 1892] 
(Eriocaulon septangulare); Britton, N. L., 24 (Eriocaulon 
Parkeri), 56 (Eriocaulon decengulare), sen. {Forked River, 
Aur. 17, 1889] (Erioceulon decangulare), sen. {Morris Pond, 
Sept. 13, 1887] (Eriocaulon septangulare); Britton, Britton, 
Earle, & Gager 6488 (Syngonanthus legopodioides); Britton, 
Britton, & Gager 6960 pees insulare), 6980 (Paepalan- 
thus seslerioides), 7058 (Syngonanthus lagopodioides), 7060 
Eriocaulon fuliginosum), 7090 (Paepalanthus alsinoides), 


71350 (Paepalanthus alsinoides), 7131 (Paepalanthus sesleri- 


oides), 7251 (Paepalanthus seslerioides), 72 Syngonanthus 


lagopodioides), 7257 (#riocaulon echinospermum); Britton, 
Britton, « Timmerman sen. [Port Sandfield, Sept. 1, 1887] 
(Brioceulon septangulare); Britton, Britton, & Wilson 14,144 
(Paepealanthus alsinoides var. minimms), 14,162 (Syngonanthus 
insularis, type coll.), 14,179 (Briocaulon arenicola, type 
coll.), 14,217 (Syngonanthus lugo octosdes), 14,220 (Erio- 
caulon ovoideum, type coll.), 1-205 ngonanthus lagopodi- 
oides), 14,225 (Paepalanthus pt Sl pe 14,948 (Erio- 
caulon sclerocephalum), 14,951 (Eriocaulon fusiforme, type 


1939 Molcenke, Notes on Eriocaulaceae 353 


coll.), 15,008 (Eriocaulon pinarense), 15,787 (Syngonanthus 
lagopodioides), 15,789 (Syngonanthua lLagopodioides), 15,789a 
(Syngonanthus Wilsonii, type coll.}; Britton & Wilson 19 
(Lachnocaulon anceps), 14,319 (Paepalanthus seslerioides), 
15,689 (Paepalanthus seslerioides); Broadway, W. &., 9310 
(Paepalanthus Lamarckii); Brown, C. A., 3462 (Eriocaulon 
septangulure), 3626 (Erioceaulon septangulare); Buckley s.n. 
a (Lachnocaulon anceps) ; Bush 71 (Eriocaulon decangul- 
are), 107 (Lachnocaulon anceps); Butters s.n. [Clear Lake, 
Auge 24, 1919] (Zriocaulon septangulare); Butters & Buell 
474 (@riocaulon septangulere). 

Camp 2265 (riocaulon Ehrenbergianum); Canby sen. [Hiber- 
nia, Merch, 1869] (&riocaulon compressum), sen. [Ellendale, 
July 24, 1893] (Eriocaulon decangulare), s.n. (Salisbury, 
Sept. 1887] (Ericcaulon septangulare), s.n. (Hibernia, 1869] 
(Lachnocaulon anceps), sen. [Prope Wilmington, Oct. 1867] 
(Lachnocaulon minus), sen. (Hibernia, March, 1869] (Lachno- 
caulon minus), s.n. [Hibernia, March, 1869] (Syngonanthus 
flavidulus); Carter sen. (Gunpowder Sta., Aug. 27, 1903] 

Eriocaulon Parkeri), s.n. (Salisbury, July 15, 1904] (Erio- 
caulon Parkeri), sen. [Belmar, July 12, 1910] (Briocaulon 
Parkeri); Chamberlain s.n. [Wenham Pond, Aug. 1899] (Erio- 


caulon septangulare); Chaney 67 (Hriocaulon septangulere); 
Chapman 182 (Syngonanthus flevidulus), sen. [Florida] (Erio- 
caulon compressum), Bene [Florida] (Lachnocaulon Engleri), 
Sen. (St. Andrew's Bay, 1838] (Lechnocaulon Engleri), 3 oNe 
(Syngonanthus flevidulus); Chardon 28 (Paepalanthus Tuerck- 
heimii); pes 8.ne 113 July, eget Eriocaulon compres- 
sum), sen. (Chatsworth, 3 June, 1931] (Zriocaulon compress- 
un), sen. [W. of Speedwell, 13 July, 1930] (Eriocaulon dec- 
angulere); Clinton s.n. [Warner, August, 1935] (Briocaulon 
septangulare); Clute 116 (Hriocaulon septangulare), 214 (Er- 
{ocaulon atari Clute & Wilson s.n. [Hempstead, July 
20--21, 1899] (Eriocaulon septangulare); Collector undesig- 
nated 272 (Hriocaulon Soninanaiaye sen. [1832] (Hriocaulon 
decan ulare), 8 Ne {Fine barrens] (Eriocaulon decan ulare), 
8 oNe ose 6, '92] (Hriocaulon Ehrenbergianum), sen. 
{Schooley's Mtn., August, 1820] (riocaulon septangulare), 
sen. (Southampton Pond, 1829] (Eriocaulon septan ulere), 8. 
n. [Florida] (Lachnocaulon Engleri), s.n. (Eriocaulon sept- 
angulare) ; Columbia University s.n. [Pine barrens ] (Erio- 
caulon compressum); Combs 588 (Eriocaulon fuliginosum); 
Commona sen. [Atsion, June 22, 1882] (Eriocaulon compress- 
um), s.n. [Pine barrens, Oct. 1, 1872] (Eriocaulon decangu- 
lare), sen. [June 18, 1875] (Hriocaulon decangulare); Con- 
rad sen. [Near Philadelphia] (Eriocaulon FParkeri); Cooper, 
W., Ben. [Lake George, Aug. 1817] (Sriocaulon septangulere), 
s.n. [Near Tarrytown, August, 1819] (Eriocaulon septangu- 
lare); Coville 202 (Eriocaulon decangulare); Crawford & 


354 PHYTOLOGIA No. 10 


Bliss sen. [Atco, July 10, 1927] (friocaulon septangulare); 
Cuesta 371 (Eriocaulon pseudocompressum), 374 (Hriocaulon 
Emannii); Curtis, M. A., Sen. (Hriocaulon decangulare); 
Curtiss 3016 [June] (Eriocaulon decangulare), 3016 [July] 
(Eriocaulon decan ulare), 3017 (Eriocaulon compressum), 3021 
(Lachnocaulon anceps), 3022 (Lachnocaulon eciliatum, type 
coll.), 4140 (Syngonanthus flavidulus), 5690 (Eriocaulon de- 
cangulere), 5911 (Lachnocaulon minus), 6894 (Lachnocaul on 
minus), s.n. [Summer, 1885] (Hriocaulon lineare); Cushman & 
Word s.n. [Shore of South Pond, Aug. 1910] (Eriocaulon sept- 
angulare); Cuthbert 141 (Lachnocaulon anceps), s.n. (August, 
May, 1881] (Lechnocaulon anceps). 

Deam, C. C., 1290 (Eriocaulon septangulare), 20,896 (Bri- 


ocaulon septangulare), 20,952 (Eriocaulon septangulare), s. 
ne [Round Lake, 9-1-97] (Eriocaulon senkedoal ice 'e Davis, C. 


A., sen. (Alma, Aug. 13, '95] (Eriocaulon septangulare), 8. 
n. (Alma, Aug. '95] (Hriocaulon septan ~ ems y sen. (Crystal 
Falls, Aug. 1905] (Eriocaulon septangulare), s.n. (Sept. 
1905] (Briocaulon septangulare), sen. [Nantucket, 14 Aug. 
1911] (Hriocaulon septangulare); Davis, E., s.n. [Pitts- 
field] (Eriocaulon septangulare); Denslow s.n. {Killing- 
worth, Sept. 5, 1902] (Eriocaulon septangulare), s.n. [New- 
bury, July 31, 1923] (Eriocaulon septangulare); Dodge 468 
(Sriocaulon septangulare), s.n. [Aug. 30, 1914] (Eriocaulon 
septangulare), sen. LAug. 30, 1916] (Eriocaulon septangule- 
ar Dowell sen. [Aug. 13, 1910] (Hriocaulon Parkeri); 
Dreisbach 3593 (Zriocaulon decangulare); Drummond, T., 356 
(Eriocaulon decan ulare), II.409 (Hriocaulon texense, type 
colle); Drushel 10,075 (Briocaulon decanrulare), 10,095 (Er- 
iocaulon decangulare), 10,135 (Lachnocaulon anceps); Duck- 
worth sen. [Little Lake, Aug. 4, 1906] (Eriocaulon septangu- 
lare); Dukes 9 (Lachnocaulon anceps). 
Eames 8976 (Eriocaulon here x 8ene (Chatsworth, VI.12. 
1894] (Eriocaulon compressum); Earle 3123 (Eriocaulon decan- 
gulere), s.n. [Leesburg, 6/20/1895] (Eriocaulon decangulare} 
Eaton, A. A., sen. (Miami, Nov. & Dec. 1903] (Eriocaulon 
Ravenelii); Eaton, D. C., aen. [1860] (Hriocaulon decangula- 
re), 8en. (Eriocaulon septangulare); Eaton, H. H., sen. 
(Sand Lake, 1829] (Eriocaulon septangulare); Eggers 2216 
(Paepalanthus domingensis, cotype coll.), 2216b (Paepalanth- 
us domingensis, cotype coll.); Eggleston 5027 (Eriocaulon 
decanpulere); Ehlers 112 (Eriocaulon septangulare), 1109 
(Eriocaulon septangulare), 1686 (Eriocaulon septangulare), 
5155 (Briocaulon septangulare), 5440 (Eriocaulon septangule- 
re), 6150 (Eriocaulon septangulare); Ehrenberg & Aschenborn 


531 (Hriocaulon Ehrenbergianum, cotype coll.); Ekman 2341 


(Paepalanthus pungens), 3522 (Faepalanthus montanus), 5709 
($ fe) 


(Paepalanthus pungens), 10,792 (Syngonanthus androsaceus), 
10,822 (#riocaulon sclerocephalum, cotype coll.), 10,823 


1939 Moldenke, Notes on HZriocaulaceae 5 


(Syngonanthus lagopodioides), 11,033 (Paepalanthus sesleri- 
oides), 11,034 (Paepalanthus alsinoides var. minimus), 11061 
(Synzonanthus Lagopodioides), 11,120 (Tonina fluviatilis), 
11,145 (Syngonanthus lagopodioides), 11,221 (#riocaulon 
seudocompressum), 11,956 (Hriocaulon miserrinum, type 
coll.), 11,965 (Paepalanthua alsinoides var. minimus), 
11,973 (Syngonanthus lagopodioides), 11,975 (Eriocaulon 
aclerocephalum, cotype coll.), 11,990 (Zriocaulon sclero- 
cephalum, cotyve coll.), 12,015 (Paepalanthus Lamarckii), 
12,029 (Eriocaulon olivaceum, type coll.), 12,029a (ario- 
caulon insulare), 12,065 (Eriocaulon cubense, type coll.), 
12,071 (s onanthus lagopodioides), 12,095 (Syngonanthus 
insularis), 12,100 (Paepalanthus alsinoides var. minimus), 
12,106 (riocaulon sclerocephalum), 12,106a (Zriocaulon ful- 
iginosum), 12,191 (Eriocaulon sclerocephalum, cotype coll.), 


12,410 (Lachnocaulon anceps), 12,806 (Paepalanthus retusus), 
12,807 (Hriocaulon dioecum, type coll.), 17,118 (Lachnocaul- 


on cubense, type colle), 17,253 (riocaulon hetero etalun, 
type coll.), 17,808 (Hriocaulon insulare, type coll.), 
17,810 (Paepalanthus seslerioides), 17,864 (Eriocaulon ful- 


iginosum), 17,877 (Eriocaulon lacustre, type coll.), 17,888 
(Eriocaulon Exmannii, type coll.), 17,918 (Paepalanthus als- 
inoides), 17,948 (Sriocaulon minutissinum, type coll.), 

18,121 (Paepalanthus Lamarckii), 18,127 (ariocaulon fuligin- 


osum), 18,128 (Eriocaulon echinospermum), 18,132 (Lachno- 
caulon Exmannii), 18,769 (Eriocaulon pinarense, type coll.); 
Erlanson, C. 0., 4595 (Eriocaulon septangulare). 

Fassett 157 (Eriocaulon Parkeri), 7711 (Hriocaulon sept- 
angulare); Ferguson 302b (Eriocaulon septanpulare), 1869 
(Eriocaulon septangulare), 2296 1/2 (Eriocaulon septangule- 
re), 2483 (Eriocaulon septangulare), 7755 (HEriocaulon sept- 
angulare), sen. [Hempstead, 7-25-19] (Zriocaulon septangul- 
ey sen. [Meadow Brook, 8-26-20] (Eriocaulon septanguls- 
re); Fernald 369 (riocaulon septangulare); Fernald, Biss- 
ell, Graves, Long, & Linder 20,597 (Eriocaulon septangule- 
re); Fernald & Long 6121 (Lachnocaulon anceps), 13,162 (#r- 
iocaulon septangulare), 15,165 (Hriocaulon Parkeri), 13,166 
(Eriocaulon Parkeri), 15,167 (Eriocaulon Parkeri), 20,594 


(Eriocaulon septangulare), Plant. Exsicc. Gray. 174 (Erio- 
caulon Rikon Ye Fernald, Long, & Dunbar 26,459 (sriocaulon 


septanrulare); Fernald, Long, & Smart 5698 (Lachnocaulon 
ance ay 6790 (Eriocaulon decangulare); Fernald & Pease 
3220 (Eriocaulon septangulere); Fernald & Wiegand 5068 (Er- 
iocaulon septanculare); Fitch, ar Sen. (East Paarpestin 
19.4.13] (Eriocaulon septangulare); Fogg 2508 (Eriocaulon 
septangulare), Wr (Zriocaulon decangulare), er (Erio- 
caulon decangulare); Foote sen. [July 19, 1868] (Eriocaulon 
etc alarey; Fowler sen. (St. John, Sept. 1873] (Erio- 
caulon septangulare); Friesner 10,224 (Zriocaulon septangu- 


356 PHYTOLOGIA No. 10 


lare); Fuertes 1748 (Paepalanthus domingensis). 

Garber sen. [Manatee, Apr. 1876] (Syngonanthus flavidu- 
lus); Gates 858 (riocaulon agate Gates, Fletcher, 
& Jewett 188 (Lachnocaulon anceps), 189 (Hriocaulon compres- 
sum var. Harperi), 190 (Eriocaulon compressum var. Har eri), 
191 (Eriocaulon compressum), 192 (Hriocaulon compressum var. 
Harperi), sen. [Mobile, May 15, 1839] (ariocaulon compressum 
ver. Harperi); Gentle 992 (Paepalanthus Gentlei, type 
coll.), 992a (Paepalanthus Lamarckii), 993 (Eriocaulon ful- 
iginosum); Gibbes s.n. [lst March, 1839] (Eriocaulon comp- 
ressum), sen. (Summerville, May 25, 1855] (Lachnocaulon an- 
ceps), sen. [Sept. 1833] (Syngonanthus flavidulus); Gilbert, 
E. Je Ce, Sense (Keene, Sept. 1875] (Eriocaulon septangulare)} 
Gleason & Gleason 202 (Hriocaulon septangulare); Gleason, 
Smith, & Alexander 173 (Hriocaulon septangulare); Gray, A., 
sene [1840] (Eriocaulon compressum); Greville s.n. (Eriocau- 
lon septangulare); Grimes 3761 (Lachnocaulon enceps), 4135 
(Eriocaulon Parkeri); Gross 3016 (Eriocaulon decangulare), 


3017 (a@riocaulon septangulare), 3118 (Eriocaulon compress- 
um), Sen. (Eriocaulon compressum) ; Groves & Groves 8.n. (8. 


VIII.1892] (Eriocaulon septangulare). 

Hale 160 (Lachnocaulon anceps), 456 (Eriocaulon decangul- 
are), 5en. (Eriocaulon decangulare); Hall, E., 635 (Hrio- 
caulon decangulare); Hamilton, C. A., 25,149 (Eriocaulon 
septan aiagey: Harper, F., 383 (Eriocaulon compressum var. 
Harperi), sen. [Floyd's Island] (Syngonanthus flavidulus); 
Harper, J. S., 161 (Syngonenthus flavidulus), 394 (ariocaul- 
on compressum) ; Harper, R. M., 7 (Lachnocaulon eciliatum), 8 
(Lachnocaulon Engleri), 22 (Zriocaulon lineare), 42 (Lachno- 
caulon minus), 47 (riocaulon lineare), 79 (Eriocaulon comp- 
ressum var. Harperi, type coll.), 85 (Sriocaulon lineare), 
22% (Eriocaulon lineare), 443 (Lachnocaulon anceps), 44 
(Briocaulon decangulare), 803 (Syngonanthus flavidulus), 804 
(Lachnocaulon anceps), 830 (Eriocaulon lineare, type coll.), 
1395 (Eriocaulon compressum var. Harperi), 1491 (Lachnocaul- 
on Be; ichianum), 1607 (Lachnocaulon minus), 1608 (Hriocaul- 
on lineare), 2146 (driocaulon lineare), 2170 (Hriocaulon 
compressum), 2219 (Eriocaulon compressum); Harper & Harper 
Sen. LAug. 3, '97] (Eriocaulon septangulare); Hartweg 258 
(Eriocaulon Benthami, type colle); Harvey, F. L., sen. [Sil- 
oam Spring, June, 1885] (Eriocaulon Kérnickianum), sen. [Or- 


ono, 1884] (Eriocaulon septangulere); Hasse s.n. (Northern 
Wisconsin, July, 1882] (Eriocaulon septangulare), sen. [Lake 
Hilbert, Aug. '84] (Eriocaulon septangulare); Heller, A. A., 
180 (Zriocaulon decangulere), 181 (Eriocaulon decangulare), 
sen. (Eriocaulon decangulare); Herb. Chapman s.n. [Florida] 
(Eriocaulon compressum), 8.n. [Florida] (driocaulon decangu- 
lere), sen. (Florida] (Briocaulon decangulare var. latifoli- 
um, type coll.), s.n. [Lake Mahopac, Aug. 1848] (Eriocaulon 


{<a 


1939 Moldenke, Wotes on Hriocaulacese 357 


septanzulere), s.n. [Lake Dunmore, 1859] (Eriocaulon septan- 
pulare), sen. [Florida] (Lachnocaulon anceps), a.n. ([Brist- 


ol] (Lachnocaulon minus); Herb. Columbia Univ. s.n. [Fla.] 
(Lachnocaulon anceps); Herb. Le Roy aen. [Fla.] (Lachnocaul- 
on anceps), Sen. LSt. Andrew's Bay] (Lachnocaulon Engleri); 
Herb. Torrey s.n. [Fine barrens] (Hriocaulon compressum), 8. 
n. {Pine barrens, June, 1818] (Eriocaulon decangulsre); 
Eerb. Torrey Botanical Club s.n. [Hanover] (Hriocaulon sept- 
angulere); Herb. Univ. Michigan s.n. [Mobile, May 7, 1849 

Eriocaulon compressum), a.n. [Aug. 25, 1838] (Hriocaulon 
septangulare); Hermann, F. J., 7012 (Hriocaulon septangule- 
aay: 7146 (Eriocaulon septangulare); Hexamer & Neier sen. 
[Manchester pond, Aug. 25] (Er iocaulon septangulare); Hinton 
627 (Hriocaulon Benthami), 3488 (Eriocaulon Benthami), 3638 
(Eriocaulon Bentham), 4549 (Hriocaulon Benthami); Hitchcock, 
A. S., 574 (Lachnocaulon glubrum), 375 (riocaulon decangu- 
lare), sen. [Eustis, June--July, 1894] (Lachnocaulon anc- 
eps), sen. [Hustis, June--July, 1894] (Lachnocaulon minus), 
Sen. [Kustis, June--July, 1894] (Syngonanthus flavidulus); 
Hogg 8, in part (Eriocaulon compressum), 8, in part (Syngon- 
anthus flevidulus); Hollick sen. [Provincetown] (Eriocaulon 
septangulere); Holm a.n. [Holmead, July 22, 1888] (Hriocaul- 
on decangulare), sen. (Surattsville, Aug. 6, 1900] (Erio- 
caulon decangulare); Houghton sen. [Lac des Isles, Aug. 5, 
1831] (Hriocaulon sevtangulare); House 2685 (Lachnoceulon 
anceps), 5108 (Hriocaulon compressum), 5822 (Hriocaulon sep- 
tangulsre), 21,951 (Briocaulon septangulare), 25,128 (Erio- 
caulon Parkeri), s.n. [North Bay of Oneida Lake, July, 1901] 
(Eriocaulon septangulare), sen. (North shore, Cneida Lake, 
Aug. 1903] (riocaulon septan ulere); Howe & Lang 686 (Erio- 
caulon septangulare), 765 (Erioceulon septangulare), 856 
(Eriocaulon septangulare), 1421 (Hriocaulon septangulere), 
1583 (Eriocaulon septangulare), 1603 (Eriocsulon septangula- 
re); Huger, A. M., sen. ("x.a."] (Eriocaulon comvesbin’. 
sene ["X.C.1I."] (Lachnocaulon anceps), sen. [Winter Park, A- 
pril, 1900] (Syngonanthus flavidulus); Hyams, M. E., sen. 
(Wilmington, July, 1879] (Lechnoceulon anceps). 

Ingalls s.n. (Lachnocaulon anceps). 

James s.n. [Delaware Bixer near Bets ma (Zrio- 
caulon Parkeri, cotype coll.); Jameson 206 (Eriocaulon mic~ 
rocephalum); Jennings 338 (Paepalanthus seslerioides), 387 
(Paepalanthus alsinoides var. minimus, type colle); Jewett 
Bene (Mobile, lay 9, 1839] (Lachnocaulon anceps). 

Keeler net {Vicinity of aise & Jacksonville] (Lachno- 
caulon anceps); Killip 3614 (Briocaulon panamense, type 
colle), 32,380 (Eriocaulon melanocephalum); Kinloch 213 (Er- 
iocaulon Kinlochii, type coll.); Knight 2 (Syngonanthus 
flavidulus), sen. [Echo Lake, 12.7.79] (Eriocaulon septangu- 
lare); Kraemer sen. [Tom's River, June 30, 1891] (Hriocaulon 


358 PHYTOLOGIA No. 10 


compressum); Kurz sen. [lay 16, 1926] (Hriocaulon compressum 
ver. Harperi). 
Lambert, 3. B., 8 (Eriocaulon septangulare); Langdon s.n. 
{Little Squaw Lake, July 27, 1894] Cet cunt septangulare); 
Langlois sen. [Pass Christian, Merch, 1880] (Eriocaulon com 
pressum var. Herperi), s.n. [Near Mandeville, i.V.1893] (Er- 
jocaulon compressum var. Harperi), s.n. [Nandeville, 15. 
VI11I.1892] (Eriocaulon decangulare), s.n. [9.1X.1892] (Erio- 
caulon decangulere), Sen. [Mandeville to Covington, April, 
1879] (Lachnocaulon anceps); Leavenworth sen. [Tampa Bay ] 
(Erioceulon compressum), sen. [Tampa Bay] (Zrioceulon decan- 
gulere), sen. LTampa Bay] (Syngonanthus flevidulus); Le Con- 
te s.n. (Lachnocaulon Be Bons Legzett s.n. [Quaker Bridge, 
July 8, 1864] (Hriocaulon decangulare), sen. [Fine barrens, 
Sept. 16--18, 1873] (Eriocaulon decangulare), s.n. [Canaan 
Pond] (Eriocaulon septangulare), s.n. LLake Mohegan, Sept. 
6, 1868] (riocaulon septangulare); Lehmann 567 (Eriocaulon 


microcephalum) ; Leén 15,946 (Syngonanthus androsaceus); Leon 
& Cazanas 5910 (Eriocaulon fuliginosum); Leén & Roca 6930 
(Syngonanthus lago odioides), 6970 (Paepalanthus seslerioid- 
es); Liebmann s.n. [Chinantla] (Eriocaulon microcephalum); 
Lighthipe 172 (Eriocaulon compressum), 173 speccagienes 
flavidulus), 452 (Lachnocaulon anceps), sen. [Monticello, 
March, 1891] (Syngonanthus flavidulus); Lloyd & Tracy 318 
di ) 


(Lachnocaulon num), 319 (Hriocaulon decangulare), 320 
(Lachnocaulon anceps); Long, B., 4721 (Er iocaulon Parkeri), 
4767 (Hrioceulon Parkeri), sen. [Killville, Oct. 7, 1909] 
(Eriocaulon Parkeri); Lucy 2237 (Eriocaulon septangulere); 
Lugren s.n. [Volusia] (Syngonanthus flavidulus). 

MacElwee 658 (Zriocaulon compressum); Wackenzie 796 (Hri- 
ocaulon septangulare), 1021 (Hriocaulon septangulare), 1050 
(Eriocaulon decangulere), 1638 (Eriocaulon decangulare), 
2361 (Eriocaulon decan ulare), 3694 (Eriocaulon decangula- 
re), 3846 (Eriocaulon sepeanculere), 4051 eee decan- 
gulere), 4059 (Lachnocaulon anceps), 4257 (Briocaulon decan- 
sulere), 4417 (Eriocaulon septengulare), 4741 (Eriocaulon 
decangulere), 4850 (Eriocaulon septengulare), 5277 (Er io- 
caulon septangulare), 5482 (Eriocaulon decangulare), 5671 
(Erioceulon Parkeri), 5684 (Eriocaulon Parkeri), 7236 (Erio- 
caulon Parkeri), 7277 (Eriocaulon septangulare), 7344 (Hrio- 
caulon Parkeri), a.n. [West of Mantoloking, Sept. 21, 1912] 

Hriocaulon Parkeri), sen. [Waretown, Sept. 1922] (Hriocaul- 
on septangulare); Macoun 22,639 (Eriocaulon septangulare), 
gen. [North Sydney, July 14, 1883] (Eriocaulon septangulere 
Merie-Victorin 15,737 (Eriocaulon septangulare), sen. [Lacs 
du Laurentides, Aout, 1912] (Eriocaulon septangulare); Mc- 
Farlin 2381 (Eriocaulon septangulare), 3418 (Eriocaulon dec- 
angulare), 4491 (Eriocaulon compressum), 6381 (Lachnocaulon 
anceps), 7545 (Eriocaulon decangulare); Well, C. D., sen. 


1939 Moldenke, Notes on Eriocaulaceae 359 


(Minatitlan, Nov. 28, 1928] (Paepalanthus Mellii, type 
coll.); Mellichamp sen. [Bluffton] (Synzonanthus flavidul- 


us); Metcalf 2172 (Eriocaulon septangulare); Meyer, . C., 
134 (Eriocaulon Williamsii); Michaux sen. (Syngonanthus 
flavidulus, type coll.); Miller, W. de W., 1534 (riocaulon 
septangulare); Miner s.n. (Aug. 11, 1931] (@riocaulon sept- 
anpulare); Minns s.n. (Crawford's, Aug. 1887] (Briocaulon 
septangulare); Mohr s.n. [Oct. 4, 1894] (@riocaulon decengu- 
lare), 3en. [De Funiac Springs, May 12, 1892] (Syngonanthus 
flavidulus); Moldenke, H. N., 233 (Sriocaulon decangulare), 
688 (Syngonanthus flavidulus), 689 (Hriocaulon compressum 
var. Harperi), 689a (Eriocaulon compressum), 940 (Syngonan- 
thus flavidulus), 1123 (Eriocaulon compressum ver. Harperi), 


5591 (Zriocaulon deca ulare), 10,577 (Briocaulon compress- 
um) ; raw sen. [Cantonsville, Aug. 11, 1873] (Hriocaulon 


compressum), sen. [Atco, Sept. 3, 1873] (Hriocaulon decangu- 
lare), s.n-e (Hamilton, July 27, 1875] (Eriocaulon septangu- 
lare), s.n. [South Natick, Sept. 15, 1881] (riocaulon sept- 
angulare); Muenscher & Curtis 5598 (Hriocaulon Parkeri), 
5600 (Hriocaulon Parkeri), 5611 (Hriocaulon septangulare). 
Nash 92 (Hriocaulon co ressum), 143 (Syngonanthua flav~ 
idulus), 148 (Lachnocaulon minus ), 847 (Briocaulon decangu- 
lare), 1184 (Lachnocaulon Engleri, type colle), 1295 (Lach- 
nocaulon minus), 1722 (Hriocaulon decangulare), 1855 (Lach- 
nocaulon minus), 1942 (Lachnocaulon anceps), 1944 (Syngonan- 


thus flavisalus), 1981 (Lachnocaulon floridanum, type 


colle), Sen. Luly 26, 1909] (Hriocaulon septangulare), s.n. 
{Twin Lakes, July 30, 1909] (Hriocaulon septan ulare); Nat. 
Herb. Canada 130,088 (Eriocaulon septangulare); Nelson, &, 
W., 6028 (Hriocaulon Benthami); Nichols 1623 (#riocaulon 
septangulare); Northrop, A. 3., sen. {Nashawena, July--Aug. 
1901 ] Pio a rcvtune septangulare), s.n. [Nashawena, Aug. 1901] 
(Zriocaulon septan i Northrop, J. I., 184 (Hriocaulon 
septangulare). 

O'Neill 81 (Eriocaulon decangulare), 611 (Syngonanthus 
flavidulus), 633 (Lachnocaulon anceps), 652 ( onanthus 
flavidulus), 722 (Hriocaulon compressum), feos Lachnocaulon 
anceps), 1153 (Eriocaulon decangulare), 1815 (Zriocaulon de- 
cangulare), 1947 (Br iocaulon compressum), 6512 (Eriocaulon 
compressum), 7778 (Hriocaulon compressum), 7779 (ir iocaulon 
compressum), 7780 (@riocaulon compressum), 7781 (Erioceulon 
decangulare), 7782 (Hriocaulon decangulare), 7783 (Lachno- 
caulon anceps), 7785 (Lachnocaulon minus), 7785a (Lachno- 


caulon Engleri), 7787 (Lachnocaulon anceps), 7788 (Syngonan- 
thus flavidulus), 7875 (Syngonanthus flavidulus), 8190 (Eri- 
ocaulon septangulere), sols (Syngonsnthus hondurensis, type 
coll.), seh se © onanthus Oneillii), 8546 (Syngonanthus 

Lundellianus, type coll.), 8547 (Paepalanthus Gentlei), 8548 
(Syngonanthus Oneillii, type coll.), 8549 (Syngonenthus 


360 Pa‘E ‘FOL OhoEd & No. 10 


Oneillii), 9096 (Hriocaulon decangulare), "Count 13" (Erio- 
caulon decan ulare), "Count 100" (Lachnocaulon anceps), 
*Count 181" iiiabowstens anceps), sen. [Fish Lake, Feb. 20, 
1927] (Eriocaulon lineare), aen. [Prairies, Oct. 28, 1929] 
(Lachnocaulon ance a), sen. [Lake Ola, July 2, 1929] (Lach- 
nocaulon Engleri), sen. (Fish Lake, Feb. 20, 1926] (Syngon- 
anthus flavidulus), sen. [Big Cypress Swamp, April, 1927] 
(Syngonanthus flavidulus); Oosting 3555 (Lachnocaulon an- 
cevs); Orcutt 3285 (Tonina fluviatilis). 

Palmer, Edw., 44 (Eriocaulon Ehrenbergianum), 172 (Erio- 
caulon Palmeri, type coll.), 580 (Syngonanthus flavidulus), 
2029 (Hriocaulon te icanunm, type coll.) Palmer, EB. J., 7718 
(Eriocaulon compressum), 7981 (Eriocaulon decangulare), 8320 
(Eriocaulon Kérnickianum), 38,324 (Syngonanthus flevidulus), 
38,325 (Eriocaulon compreasum); Parker, C. F., sen. LOpp. 
Sea Isle City, Aug. 18, '82] (Eriocaulon decangulare), sen. 
(Camden] (Eriocaulon Parkeri, cotype coll.); Pelly 73 (Erio- 
caulon fuliginosum); Pennell 1927 (Eriocaulon decangulere), 


Gi44 (Eriocaulon decangulare), 4465 (Eriocaulon decangulare 
var. latifolium), 4472 (Eriocaulon decangulare), 4474 (Lack. 


nocaulon digynum), 4509 (Syngonanthus flavidulus), 8194 (Er- 
iocaulon septangulare), 8400 (Eriocaulon septangulare), 8431 
(Eriocaulon septangulare), 9075 (Eriocaulon decangulare), 
9079 (EZriocaulon decungulare), 9898 (Eriocaulon Parkeri), 
12,903 (Eriocaulon decangulare), 16,704 (Eriocaulon septang- 
ulare); Phelps 296 (Eriocaulon septangulare); Pittier, H., 
4932 (Paepalanthus Lamarckii), sen. [Herb. Instit. Physico- 


geogr. Nat. Costaric. 11,043] (Syngonanthus caulescens); 
Pogzenburg sen. [Manchester, Sept. 19, '85] (Eriocaulon dec- 
angulare); Pollard & Maxon 562 (Lachnocaulon anceps); 
Pringle 1734 (Eriocaulon guadalajarense, type coll.), 2018 
(Hriocaulon Pringlei), 2936 (Eriocaulon Schiedeanum; type 
coll. of E. Jaliscanum), 3855 (Hriocaulon bilobatum, type 
coll.), 6144 (Hriocaulon microcephalum), 6146 (Hriocaulon 


Schiedeanum), 6299 (Eriocaulon bilobatum), 7361 [Eriocaulon 


microcephalum), 8989 (iriocaulon hrenbergianum), 11,202 
(Eriocaulon mexicanum, type coll.), 13,228 (Eriocaulon mic- 


rocephalum), a.n. (Near Guadelajara, Oct. 7, 1890] (Erio- 
caulon Schiedeanum); Purpus 179 (Faepalanthus chia ensis), 
10,565 (Paepalanthus chiapensis, type colle); Pyle 754 
(Eriocaulon Kérnickianum). 

Raup 7748 (Eriocaulon septangulare); Ravenel s.n. [St. 
Johns] (Eriocaulon Ravenelii, type coll.), sen. [Aiken 1870] 
(Lachnocaulon anceps), sen. (Florence, 1879] (Lachnocaulon 
anceps), gen. (Florence, 1879] (Syngonanthus flavidulus); 
Reverchon 2766 (Hriocaulon decangulare), 4359 (Eriocaulon 
texense); Reynolds, M, C., Sen. etc Augustine, 1877] (Erio- 
caulon compressum), s.n. [St. Augustine, lMar.--June, 1875 
(Lachnocaulon Beyrichianum), s.n. [Mar.--July, 1875] (Syngo- 


1939 | Moldenke, Notes on Zriocaulsceae 361 


nanthus flevidulus); Rich, 2. H., sen. [Ton's River, May 30, 


1887] (Sriocaulon compressum), sen. [July 26, 1896] (Brio- 


caulon septanzulare); Hobinson, C. B., 396 (Eriocsulon sept- 
angulare); Robinson & Schrenk 112 (Eriocaulon septangulare), 
S.Ne 


12 Aug. 1894] (Brioceulon septangulere); Kose, J. ie, 
8.n. [Auguat, 1897] (trioceulon Ehrenberrianum); Rose & 
Painter 7929 (Eriocaulon microcephalum); Rousseau 35,579 
(Eriocaulon septansulere); Rugel s.n. [Swanano, Jul. 1841] 
(Eriocaulon decenpulare); Ruger a.en. (Manchester] (driocaul- 
on decen ulere); Ruggles sen. (Near Milton] (Hrioceulon sep- 
tanpulare); Rusby, H. H., sen. [Bass River, sug. 1875] (Br i- 
ocaulon compressum), s.n. [Bass River, Aug. 1875] (#riocaul- 
on decangulare), sen. [Stockton, Aug. 1889] (Hrioceulon dec- 
angulsre), s.n. (Stockton, Aug. 1889] (riocaulon septengul- 
are); Ruth 133 (Lachnocaulon anceps). 

Salmon sen. [circa 1843] (Hrioceulon septanculare); Sand- 
berg sen. (Aug. 1891] (Eriocaulon septan PoE Scheffner, 
We, 226 (Hriocaulon ghrenbergianum), 2268 (Eriocaulon Zhren- 
bergianum), 226b (Eriocaulon microcephalum), 426 (Eriocsulon 
microcephalum); Schiede s.n. [Minerel del Monte] (Erioceulon 
Shrenbergianum, cotype coll.), sen. [Toluce] (Zriocaulon Eh- 
renbergianum, cotype coll.); Schipp 647 (Eriocaulon Schipp- 
ii, type coll.), 693 (Tonina fluviatilis), 5.130 (Paepalan- 
thus Lamarckii); Schrenk s.n. [Manchester, July 25, '77] 
(Briocaulon decangulare); Schweinitz sen. (Hriocaulon decan- 
gulure); Seemann 295 (Eriocaulon Seemannii, type coll.); 
Sessé, liocino, Castillo, & Maldonado 456 (Eriocaulon Ehren- 
bergianum; cotype coll. of E, anceps Sessé & lMoc.), 465 (ir- 
ioceulon Ehrenbergianum; cotype coll. of & anceps Sessé & 
Moc. )3 Seymour 268 (Eriocaulon septangulare); Shafer 4093 
(Paepalenthus seslerioides), 4104 (Paepalanthus montanus), 
4106 (Paepalanthus riparius, type colle), 447% (Paepalanthus 
montunus, type coll.), 8045 (Faepalanthus montanus), 8251 
(Paepalanthus montanus), 10,663 (Sriocuulon fuliginosum), 
10,682 (Paepalanthus alsinoides var. minimus), 10,866 (Syn- 

onanthus lagopodioides), 10,882 (Paepalanthus seslerioid- 
es), 10,956 (Pac alanthus seslerioides), 10,995 (Syngonanth- 
us tries, 11,011 (Lachnocaulon Ekmannii), 11,705 
(Er iocaul on echinos ermum), 11,782 (Ss ngonanthus lagopodi- 
oides), 11,784 (Syngonanthus androsaceus); Sheehan sen. 
Cat-tail Island, Feb. 26, 1919] (Hriocaulon compressum), s. 
n. [Goffstown, July 31, 1931] (Hriocaulon septangulere), 5. 
n. (Boakum Place, March 6, 1919] (Syngonanthus flavidulus); 
Simpson, J. H., 396 (Hriocaulon decangulare); Skottsberg, C., 
sen. (Buckingham, 19.5.1935] (Eriocaulon compressum), 8.n. 
(Sanford, 14.5.1935] (Lachnoceulon anceps), sen. [Below 
Jacksonville & Atlantic Beach] (Syngonanthus flavidulus); 
Skutch 617 (Eriocaulon Ehrenbergianum), 2465 (Tonina fluvi- 


atilis); Small, J. K., 7386 (driocaulon com ressum), 8339 


362 PYoT OLOGIA No. 10 


(Lachnocaulon glsbrum), s.n. [Dunn's Ntn., Aug. 18--27, 
1894] (Eriocaulon decangulere), Sen. (Near Valparaiso, June 
6--12, 1895] (Eriocaulon decangulare), s.n. [About Darien 
Jet., June 25--27, 1895] (Eriocaulon decangulere), s.n. [Li- 
ly Lake, Aug. 15, 1889] (Eriocaulon septangulare), sen. 
(Budd's Lake, August 12--14, 1890] (Hriocaulon septangula- 
re); Small & Carter 654 (Eriocaulon Ravenelii), 690 (Lachno- 
ceulon glabrum), 888 (Eriocaulon Ravenelii), 1024 (Lachno- 
ceulon glabrum), 1029 (Hriocaulon decangulure), 1037 (Lach- 
noceulon minus), 1148 (Eriocaulon Ravenelii), 2991 (H#riocau- 
lon compressum), 3067 (Erioceulon Havenelii), sen. [Between 
Cocoanut Grove & Cutler] (Hriocaulon Ravenelii); Small & 
Heller 180 (driocaulon decanpulare); Small, Carter, & Small 
2264 (rioceulon compressum), 3361 (Hriocaulon compressum); 
Small & Dewinkeler 9044 (Er iocaulon compressum var. Harper- 
i), 944a (Eriocaulon compressum); Small, Mosier, & Small 
6892 (Lachnocaulon glabrum), 6912 (Lachnocaulon glabrum); 
Small & Nash sen. [West of Miami, Nov. 1 & 9, 1901] (Erio- 
caulon Ravenelii); Small & Small 4429 (#riocaulon Ravenel- 
ay 4447 (Briocaulon Ravenelii); Small & Wherry 11,695 
(Erioceulon compressum var. Harperi); Small & i/ilson 1608 
(Lachnocaulon glabrum), 1614 (Lachnocaulon glabrum), 1783 
(Lachnocaulon glabrum); Smith, A. V., 325 (driocaulon comp- 
ressum), 333 (Eriocaulon compressum), sen. [liay, 1956] (Er- 
iocaulon compressum), sen. [N. J. pine barrens, July 28, 
1935] (Hriocaulon decangulare); Standley, J. F., 10 (Syngo- 
nanthus flavidulus), 15 (Eriocaulon compreasum), 33 (Lach- 
nocaulon slabrum); Standley, P. C., 42,326 (Paepalanthus 
costaricensis, type coll.); Standley & Valerio 43,637 (Eri- 
ocaulon microcephalum), 45,830 (Eriocaulon microcephelum); 
St. John 1168 (Eriocaulon septanpulare). 

Taylor, B. C., aen. [Center City, Aug. 1892] (Hriocaulon 
septangulare), a.n. [Linn Lake, Aug. 1892] (Hriocaulon sep- 
tangulere); Taylor, N., 2558 (Zriocaulon Parkeri), 2853 (Er- 
jocaulon decangulare); Tidestrom 11,236 (Eriocaulon septang- 
ulare), 8en. (July 4, 1907] (EZriocaulon decangulare); Topp- 
ing 2609 (Syngonanthus flavidulus), 2611 (riocaulon comp- 
ressum); Torrey Jotenical Club sen. [Forked River, May 29-- 
June 2, 1896] (riocaulon conmpressum); Torrey & Cooper sen. 
{Near Quaker Bridge] (Hriocaulon compressum); Townsend & 
Barber 117 (Zriocaulon Ehrenbergianum); Tracy 3428 (#rio- 
caulon decangulere), 5031 (Lachnocaulon anceps), 5032 (Erio- 
caulon compressum var. Harperi), 6417 [Biloxi] (Zriocaulon 
decangulare), 6417 [Ocean Springs] (riocaulon decangulare), 
6642 “agent flevidulus), 6643 (Syngonanthus flavidu- 
lus), 7586 (Lachnocaulon anceps), 7587 (Hriocaulon decangu- 
tare} 7588 (Syngonanthus flavidulus), S043 (Hriocaulon dec- 
angulare); Tuckerman sen. (Mountain Lakes] (Hriocaulon sept- 
angulare); Turckheim 3327 (Paepalanthua Tuerckheimii, type- 


1949 Moldenke, Notes on Eriocaulaceae 363 


coll.), 3422 (Paepalanthus domingensis). 
Underwood, L. '., 1928 (Hriocaulon compressum), 3124 (zr- 


( 
iocaulon septangulere), s.n. [West Goshen, Aug. 1890] (Hrio- 
caulon septangulare); Univ. of Michigan party s.n. (July, 
1868] (Hriocaulon septangulere), sen. [1868] (Eriocaulon 


septangulare), 

Yan Hermann 570 (Syngonanthus androsaceus); Van Pelt aen. 
(Chatsworth, Aug. 20, 1907] (Hriocaulon decanzulare). 

Walsh sen. [Southern States] (Eriocaulon com ressum); 
Weatherby, C. A., 6123 (briocaulon compressum) ; Wehmeyer 3. 
n. (Hart Lake, Sept. 8, 1930] (Erioceulon septangulere); 
iherry sen. [Flat Rock Sta., 5-30-1927] (Hriocaulon line- 
are); White, T. &., s.n. (Shores of Jordan's Pond] (Erio- 


caulon septangulare); “hitehouse 581 (Lachnocaulon anceps); 
Williams, R. S., 299 (Hrioceulon Williamsii, type patieae 
Williamson, C. S., 1106 (Hriocaulon septangulare), sen. 
(Bennet, June 16, 1907] (Eriocaulon compressum), a.n. [Fish 
House, Oct. 2, 1905] (Eriocaulon Parkeri), sen. (Fish House, 
July 28, 1906] (@riocaulon Parkeri), s.ne [Havre-de-Grace, 
Sept. 1, 1906] (Hriocaulon septangulare), s.n. [Weldon] 
(Lachnocaulon minus), s.n. [Wilmington] (Lachnocaulon min- 
us); tlilson, P., sen. [Opposite Island Heights, July 26, 
1909] (Eriocaulon decangulare), sen. [iferrick, Sept. ll, 
1915] (Eriocaulon septangulare), sen. [Lake Katrine, Auge 
18, 1916] (Eriocaulon septangulare), sen. [Lake Shandelee, 
Auge 7, 1918] (Eriocaulon septangulare), sen. {Sand Pond, 
Aug. 13, 1918] (Hriocaulon septangulare); Wood, N. A., Sen. 
(Whitefish Point, 1912] (Eriocaulon septanpulare); Woodson, 
Allen, & Seibert 1130 (Hriocaulon panamense), 1650 (@rio- 
caulon Seemannii); woodward s.n. [Old Lyme, Aug. 1, 1918] 
(Eriocaulon Parkeri); Wright, C., 748 (Paepalanthus sesleri- 
oides), 749 (Syngonenthus androsaceus), 753 (Hrioceulon ful- 
iginosum), 756 (rioceulon melanocephalum), 3229 (Eriocaulon 
fuliginosum, type colle), 4234 (Paepalanthus seslerioides, 
type coll.), 3235 (Syngonanthus androsaceus, type coll.), 
3236 (Syngonenthus androsaceus), 3257 (Syngonanthus lezopod- 
foides, type coll.), 3238, in part (Hriocaulon fuliginosum; 
cotype coll. of E, scirpoides), 3238, in part (Syngonanthus 
lagopodioides), 3240 (Hrioceulon melanocephalum), 3241 (Eri- 


ocaulon melanocephalum var. longipes, type coll.), 3242 
(Tonina fluvistilis), 3737 (Eriocaulon sigmoideum, type 


coll.), 3728 (Hriocaulon echinospermum, type coll.), 3738 
(Eriocaulon fuliginosum; type coll. of EB. aphaerospermum), 
3740 (Briocaulon fuliginosum; type coll. of E, trichosepal- 
um), 3741 (Hriocaulon pseudocompressum, type coll.), 5710 
(Paepalanthus Lamarckii), 3743 (Peepalanthus alsinoides, 
type coll.), 3743a (Paepalanthus alsinoides var. minimus), 
3744 (Paepalanthus retusus, type coll.), sen. (Eriocaulon 
echinospermum), s.n. (Paepalanthus alsinoides); Wright, G. 


364 PHYTOLOGIA No. 10 


¥., sen. [1876] (eriocaulon septangulare), 
Xantus de Vesey s.n. [Fort Tijon] (Zriocaulon microcepha- 
lum). 


_- -—-— = =& —«— =—§ =—|—= =§ =& =—§ = =~ = = s&s = = =e =e = = 


ADDITIONAL NOTES ON THE GENUS AEGIPHILA -- V 


Harold N. Moldenke 


The following notes constitute a continuation of those 
published in Phytologia 1: 182--208, 222--240, and 248--272 
(1937) and 289--304 (1938). 

An alphabetized list of citations [additional to those 
published in Brittonia 1: 472--477 (1934) and Phytologia 1: 
301--304 (1938) ]: 


Gaudichaud 630 (Luschnathi), 1801 (brachiata), sen. 
[1830] (Sellowiana); Geay s.n. [Guaritico] (mollis); Gehrt 
sen. [Herb. Instit. Biol. S. Paulo 30,081] (graveolens); 
Gentle 263 (monstrosa), 404 (monstrosa), 948 (monstrosa); 


Glaziou 11,333 (lanceolata; type coll. of A. ferruginea 
Glaz.), 21,917 (lanata, type coll.), A (inteerifolia); Glea- 
son 237 (Gleasonii, type coll.); Glocker 578b (Lhotzkiana), 
2166 (Lhotzkiana); Goeldi 6999 (racemosa), 7002 (racemosa), 
8166 (Goeldiana, type coll.); Goudot 2 (mollis), sen. [Bogo- 
ta] (bogotensis), s.n. (elata), s.n. [Pandi] (Goudotiena, 
type coll.), sen. [Bogota] (grandis), s.n. [Portachuela, 
Quindiu] (grandis, type coll.); Gouin sen. [1867] (Deppea- 
na); Greg s.n. (obovata); Grisebach s.n. (martinicensis ); 
Grosourdy Cat. 13 s.n. (parviflora), sen. [1864] (Hastingsi- 
ana, type coll.); Guilding s.n. (martinicensis); Guillemin 
54 (Luschnathi), 630 (Luschnathi), 975 (obducta). 

Hahn sen. [Medelli] (Deppeana), s.n. [Fotrero] (elata); 
Hallier f. C.240 (martinicensis); Handro sen. (Herb. Instit. 
Biol. S. Paulo 43,523] (obducta); Harris, W¥., 5533 (uniflor- 
a, type coll.), 8166 (martinicensis var. oligoneura, type 
coll.), 8996 (obtusa, type coll.), sen. [7.41.95] (elata); 
Hart s.n. (elata); Hassler 1674 (Hassleri, cotype coll.), 
1921 (psraguariensis, cotype coll.), 3193 (Hassleri, cotype 
coll.), 4271 (Hassleri, cotype coll.), 4498 (paraguarien- 
sis, cotype coll.), 5056 (paraguariensis, cotype coll.), 
6766 (paraguariensis, cotype coll.), 6780 (Hassleri, cotype 
coll.), 6941 (paraguariensis, cotype coll.), 8056 (platy- 

hylla, type pore ae 8120 (Candelabrun, type coll.), 8632 
Maasiers). 10,852 (para ariensis); Haught 1629 (glandulif- 
era); Hayes 269 (glandulifera), s.n. [Dec. 7] (glandulifer- 


1939 Moldenke, Notes on Aegiphila 365 


a), sen. [Fanama, Dec. 7, 1862; Natal Herb. 10,208] (panem- 
ensis); Heinrichs 499 (pendula); Henschen sen. [Herb. Reg- 
nell. 1.184, 1868] (Sellowiana); Herb. Adeanson 227 (integri- 
folia), s.n. (mertinicensis); Herb. Baillon sen. (bozoten- 
sis), sen. (mollis), s.n. (ternifolia); Herb. Bernhardi s.n. 
(Lhotzkiana); Herb. Boas 1113 (fluminensis); Herb. Caes. Pe- 
trop. sen. (Herb. Martius] (vitelliniflora); Herb. Colsmann 
Sen. (martinicensis); Herb. De Candolle 829 (inteprifolia), 
869 (Sellowiana), 876 (Sellowiana), 892 (Lhotzkiana, type 
coll.); Herb. Gen. ius. Para. 2270 (intermedia, type coll.); 
Herb. Hooker s.n. (mertinicensis); Herb. Hornemann a.n. [Ex 
Ind. occid.?] (martinicensis); Herb. Imp. Vien. 149 (splend- 
ens, type coll.), 152 (vitelliniflora), 155 (Lhotzkiana), 
1643 (lenceolata); Herb. Instit. Biol. S. Paulo 748 (parag- 
uariensis), 5489 (verticillata), 8179 (Sellowiana), 15,é1h 
(dentata, type coll.), 20,089 (pernambucensis), 20,099 (lan- 
ceolata), 20,562 (lanceolata), 28,666 (Sellowiana), 40,081 
(graveolens), 31,636 (Sellowiana), 33,425 (Hoehnei, type 
coll.), 33,523 (obducta); Herb. Instit. Fhysico-geogr. Nate 
Costaric. 9167 (contaricensis, type coll.), 13,629 (magnifi- 
ce), 14,484 (glandulifera); Herb. A. L. Jussieu 5034a (mart- 
inicensis), 5034b (martinicensis), 5045a (elata), 5035b (el- 
ata), 5036 (integrifolia), 5039 (racemosa), 5041 (elata), 8. 
ne (martinicensis); Herb. Lamarck 261 (nervosa), sen. [Guad- 
eloupe] (martinicensis); Herb. Liebmann s.n. (martinicens- 
is); Herb. Link s.n. (leevis); Herb. Linnaeus G.810, S.8& 
(elata), G.810, S.9 (Deppeana); Herb. Martius 379 (pareguar- 
fensis), 1C41 (vitelliniflora; type coll. of A. cuspidata), 
sen. Herb. Monac. 1450] (Sellowiana), sen. (fluminensis), 
sen. (vitelliniflora); Herb. Monac. 1000 (racemosa), 1002 
(inteprifolia), 1003 (bracteolosa), 1004 (bracteolosa; co- 
type coll. of A. arborescens var. longiflora), 1005 (bracte- 
vom toes) coll. i A, arborescens var. lon flora), 1006 
bracteolosa), 1007 (De eana), 1008 (recemosa), 1009 (race- 
mosa), 1011 (racemosa), 1C12 (racemosa), 1014 (vitelliniflo- 
ra; type coll. of A. cus idata), 1015 (vitelliniflors), 1016 
(vitelliniflora), 1017 (elata), 1018 (elata), 1019 (elste), 
1020 (filipes, cotype coll.), 1021 (filipes), 1022 (filipes), 
1023 (fluminensis), 1024 (fluminensis), 1026 (greveolens), 
1028 (Lhotzkiana), 1029 (Lhotzkiana), 1030 (Lhotzkians), 
1031 (paragueriensis), 1032 (Lhotzkiana), 1033 (parviflora), 
1034 (Lhotzkiana), 1035 (martinicensis), 1036 (martinicens- 
is), 1C37 (mertinicensis), 1038 (martinicensis), 1039 (mart- 
inicensias), 1040 (mediterranea), 1042 (Sellowiana), 1043 
(Sellowiana), 1045 (verticillata), 1046 (verticilleta), 1047 
(verticilleta), 1091 (graveolens), 1188 (membranacea), 1189 
(laevis), 1447 (Lhotzkiana), 1448 (martinicensis), 1449 
(martinicensis), 1450 (Sellowiana), 1657 (verticilleta), 
1658 (Sellowiana), 1689 (filipes, cotype colle), 1692 (grev- 


366 PY FOG O°G 2a No. 10 


eolens), 1723 (lLaxicupulis), 1724 (falcata), 1725 (Sellowia- 
ne), 1726 ajgert lis , 1727 (setiformis; type coll. of ae 
densiflora), 1843 (perplexa, type coll.), 3706 (glomereta), 
3822 eR a Gadetdin 3831 (integrifolia), 3833 (elate), 
4033 (Hassleri), 4094 (costaricensis), 4297 (fasciculata, 
type coll.), 4298 (fasciculata, type coll.), 4305 (obovata); 
Herb. Mus. Goeldi 2270 (intermedia, type coll.); Herb. Mus. 
Paris sen. (longifolia); Herb. Pavon sen. [Huayaquil] (Pav- 
oniana), sen. ("Peruvia"] (Pavoniana, type coll.), sen. 
(Deppeana); Herb. Persoon s.n. (mertinicensis); Herb. Pierre 
sen. (ferruginea); Herb. Puerar s.n. [1824] (martinicensis); 
Herb. Regnell 1.184 [1868] (Sellowiana); Herb. Rio de Janei- 
ro 19,949 (mecrentha, type coll.), 22,947 (obducta); Herb. 
Roig 4232 (elata); Herb. Spruner a.n. (Lhotzkiana); Herb. U- 
niv. Christian. sen. (martinicensis); Herb. Vahl s.n. (mart- 
inicensis); Herb. Ventenat s.n. (elata), s.n. (Luschnathi); 
Herb. ‘Jilldenow 2834 (mollis; type coll. of A. salutaris); 
Herb. Zuccearini s.n. [Herb. Monac. 1034] (Lhotzkiana), sen. 
{Herb. Monac. 1016] (vitelliniflora), s.n. (Herb. lonac. 
1043] (Sellowiana); Herminier s.n. (martinicensis); Herv. 
Com. Geogr. e Geol. S. Paulo 4362 (dentata, type coll.); 
Herzog 1369 (Herzogii, type coll.); Heyde 191 (laxicupulis, 
type coll.); Hitchcock, A. S., sen. (1843] (elata), sn. 
(Grand Cayman, 1-17-'91] (caymanensis, type coll.), sen. 
{Grand Cayman, 1-17-'91] (elata); Hoehne, F. C., pl. viv. 
192 (Sellowiana), pl. viv. 442 (obducta), sen. [Herb. Inst. 
Biol. S. Paulo 33,523] (obducta), Com. Rondon 1287 (laevis), 
1341 (integrifolia), 4301 (Candelabrum), 5122 (integrifol- 
ia), 6109 (verticillata); Hooker, W., s.n. [18435] (elata); 
Hort. Boyton s.n. [Ay Bs Lambert, 1807; Lord Seaforth, West 
Indies] (obovata, type coll.), sen. [A. B. Lambert, 1807; 
Lord Seaforth, iiest Indies] (martinicensis; type coll. of A. 
diffuse), s.n. [A. 8. Lambert, 1808; Lord Seaforth, West In- 
dies] (martinicensis); Hort. Kew. sen. {1785] (martinicens- 
4s); Hort. Liverpool sen. (martinicensis); Hort. Monac. s.n. 
[Herb. Monac. 1028] (martinicensis); Hort. Thenensis 1.4507 
(martinicensis); llostmann 89 (membranacea, type coll.), 543 
(laevis), 721 (laevis), s.n. (laevis); Hostmann & Keppler 
89, in part (membranacea); Houllet sen. (obducta); Huber 809 
(intermedia), 4253 (integrifolia); Huggins sen. [Trin. Bot. 
Gard. Herb. 2385] (integrifolia); Humboldt & Bonpland s.n. 
{El Moral, Quindiu] (quinduensis, type coll.), s.n. [Woods 
of the Orinoco] (mollis, type coll.). 

Imray 85 (martinicensis), 420 (martinicensis); Isert 87 
(mertinicensis ). 

Jacquin s.n. (martinicensis, type coll.); Jard. Bot. Par-— 
is s.n. [de l'isle de Cuba] (martinicensis); Jard. Malmaison 
s.n. (martinicensis); Jelski s.n. [Cayenne] (membranacea); 
Jenman 5948 (integrifolia), 6689 (villosa), 6715 (racemosa); 


1959 . Moldenke, Notes on Aegiphila 367 


Jobert 630 (bracteolosa); Julio 454 (multiflora). 

Kappler 543 (laevis), sen. (laevis); Karsten a.n. [1847] 
(verrucosa), a.n. (Colonia Tovar, 1848] (verrucosa: type 
coll. of Briickea srandifolia), a.n. (floribunda); segel 173 
' (laevis), 687 (laevis); Kenoyer 607 (cephalophora, type 

coll.); Kerber 305 (Deppeana); Killip & Smith 14,729 (panic- 
ulata, type coll.), 14,849 (sylvatica, type coll.), 19,706 
bogotensis), 20,870 (Killipii, type coll.), 25,503 (glabra- 
ta, type coll.), 26,520 (ovata, type coll.), 26,882 (fili- 
pea), 26,957 (Smithii, type coll.), 27,055 (elegans, type 
coll.), 27,439 (sufflava, type coll.), 27,795 (sordida), 
28,386 (cuneata, type coll.), 28,858 (triflora, type coll.), 
30,661 (glandulifera var. paraénsis, type coll.); llug 2511 
(aufflava var. Klugii, type coll.), 2574 (sufflava), 3016 
(glandulifera), 3468 (integrifolia), 3511 (peruviana), 3894 
(Smithii); Kohaut s.n.-(martinicensis); Koscinsky 225 (Sel- 
lowiana); Kraus 273 (martinicensis); Krebs sen. (martinicen- 
sis); Krukoff 1400 (villosissima, type coll.), 5060 (bracte- 
olosa), 5125 (filipes), 5923 (glandulifera var. paraénsis), 
8041 (filipes), 8042 (filipes), 8290 (glandulifera), 8701 
(elegans ); Yuhlmann, Com. Rondon 2277 (Hoehnei, type coll.); 
‘untze 8.n. (chrysantha). 

Labroy 9 (integrifolia); Lambert s.n. (martinicensis ); 
Lang & Persaud 253 (bracteolosa, type coll.); Langsdorff s. 
n. ([Mandiocca; Riedel} (Sellowiana); Lawrance 548 (integri- 
folia); Leblond 269 (villosa), 283 (martinicensis), 362 
(laevis); Ledru s.n. (martinicensis); Lee s.n. (Dec. 2, 
1887] (martinicensis); Legit Anonymus ante ann. 1840 s.n. 
(martinicensis); Lehmann 4066 (novogranatensis, type coll.), 
6692 (mollis; type coll. of Cornutia velutina), 8524 (acule- 
ifera, type coll.), 3.T.1117 (Lehmannii, type coll.); Lepri- 
eur sen. {Guiana frangaise, 1838] (villosa), s.n. [Guiana 
frangaise, 1840] (villosa), s.n. (laevis); Lhotaky s.n. 
{Herb. De Candolle 892] (Lhotzkiana, type coll.), aen. (flu- 
minensis); Liebmann 11,957 (Deppeana); Lillo 10,448 (medi- 
terranea); Lindberg 496 (verticillata); Lindley 222 (paragu- 
ariensis), 222b (paragusriensis); Lockhart s.n. [All hills 
that have been lately cultd.] (laxiflora), sen. [Bahia] 
(Lhotzkiana); Lofgren 245 (paragueriensis); Luetzelburg 
12,345 (Sellowiana), 22,296 (bracteolosa), 26,190 (Lhotzkia- 
na); Lund 796 (graveolens, type coll.), 820 (verticillata), 
sen. [Taubaté, November '33] (verticillata), s.n. (Lhotzkia- 
na); Lundell, C. L., 1492 (monstrosa); Luschnath sen. [Lagoa 
Sacaremo] (Luschnathi, cotype coll.), sen. [Brasilia] (Lus- 
chnethi, cotype coll.), sen. (Herb. Martius 1041; Herb. Mon- 
ac. 1014) (vitelliniflora; type coll. of A. cuspidata), an. 
[Brasilia] (fluminensie), sen. [Capocabona] (fluminensis), 
sen. (fluminensis); Lystler sen. (Cult. Hort. 3erol., Sept. 


1839] (vitelliniflora, type coll.). 


368 PHY T0 LOG 24 No. 10 


Macbride photos 7879 (Luschnathi, photo of cotype), 7880 
(graveolens, photo of type), 20,350 (filipes, photo of co- 
type), 22,775 (odontophylla), 24,614 (Lhotzkiana), 24,619 
(peruviana, photo of isotype); Malme 898 (Riedeliana), 951 
(Riedeliana), 2473a (paraguariensis); March 902 (trifida), 
972 (elata), 1411 (foetida), 1461 (elata); Martin, J., sen. 
[Rudge] (laevis), s.n. [Cayenne] (villosa), s.n. (laevis); 
Martius 0,39 (racemosa), 120 (graveolens), 166 (mediterran- 
ea), 448 (Sellowiana), 480 (verticillata), 1040 (Luschnathi, 
cotype coll.), 1112 (fluminensis), 1934 (graveolens), 2222 
(Lhotzkiana), 2724 (racemosa), s.n. [Herb. Monac. 1000 & 
1009] (racemosa), sen. [Forto dos Miranhao, Decbr.; Herb. 
Monac. 1004] (bracteolosa; cotype coll. of A. arborescens 
var. lonziflora), s.n. (Barra de Rio Negro, Octbr.; Herb. 
Yonac. 1005] (bracteolosa; cotype coll. of A. arborescens 
var. longiflora), 8.n. L[Prov. Rio Negro; Herb. Monac. 1006] 
(bracteolosa), sen. [Sylvia secus Amazon, Paré; Herb. Monac. 
1020 & 1689; Macbride photos 20,350] (filipes, cotype coll.), 
a.n. [Frov. Paraénsis; Herb. Monac. 1021 & 1022] (filipes), 
sen. [Herb. Monac. 1026] (graveolens), s.n. {Herb. Monac. 
1031] (paraguariensis), s.n. (R. Negro; Herb. Monac. 1011] 
(racemosa), sen. [Herb. Monac. 1046 & 1047] (verticillata), 
Sene (Herb. Monac. 1030] (Lhotzkiana); Masson s.n. (elata); 
Mathews, A., s.n. ([Chacapoyas] (cordifolia); Matuda 666 
(falcata); Mayerhoff s.n. [1859] (integrifolia); McFadyen s. 
n. (elata), sen. (trifida); Melin s.n. [Iquitos] (Smithii); 
Mélinon 83 (membranacea), 111 (membranacea), 330 (macranth- 
a), 436 (membranacea), 439 (villosa), 486 (membranacea), s. 
n. (in 1864] (membranacea), s.n. [1845] (laevis), sen. 
(1877] (macrantha); Mendonca 727 (paraguariensis); Mexia 
5922 (salticola, type coll.), 6499 (Smithii), 6656 (alba, 
type coll.), 7173 (integrifolia), 7187 (inte rifolia), 7289 
(interprifolia), 7446 (ferruginea); Miers 3096 (mediterran- 
ea), 3191 (fluminensis), 3712 (Luschnathi), s.n. (Laranjei- 
ra] (fluminensis), s.n. [Tejuco] (fluminensis), s.n. [Barra 


de Iguassu] (graveolens), sen. {Arraras, Jan. 7, 1838] 
(Riedeliana), sen. [Organ Mount., 1828] (Sellowiana), a.n. 
(Imbuby, Organ Mt., Dec. 1837] (Sellowiana), 8.n. [ilage to 
Freichal, 1 May 1838] (vitelliniflora), s.n. (Organ Mount. ] 
(obducta); Mikan sen. (Luschnathi); Miller, P., 8 (Deppea- 
na); Mocquerys 910 (martinicensis), 1018 (laeta); Moricand 
@en. (Lhotzkiana); Moritz 365 (mollis), 364 (glandulifera 
var. amidata), 897 (verrucosa, type coll.), 973 (elata), 
1910 TORT Mosén 2005 (Riedeliana); Moss sen. [1919] 
(glandulifera var. paraénsis); Miller, Fr., s.n. [Schwacke 
1465] (brachiata); Mus. Bot. Berol. sen. ["Bras.?"] (martin- 
icensis; type coll. of A, straminea); Mus. Yale School of 
Forestry 22,820 (monticola, type coll.), 28,648 (Rimbachii, 


type coll.), 31,994 (ferruginea).— 


CES aie 
pegtiegs 


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— 


> H Y T OLOGIA 
Designed to expedite botanical publication 


September, 1940 


CONTENTS 


be Ht: i Glasicn ai Harold N. Moldenke he tte — 
The I New York Botanical Garden ere Sand Re eRe 
Bronx | Park, New York, N. Y. ike i ete aed 


, 50 os cents; A volume, $5.00 in advance 


RIDKRAL 
NEW YOR 
BOTANIC, 

GARDEN 


ADDITIONS TO THE FLORA OF MEXICO AND CENTRAL AMERICA (a) 


O. Le Lundell 


PRUNUS MATUDAI Lundell, Bpe NOVe 

Arbor glaberrima, 7 m. alta, 17 cm. diam. Petioli 5--7.5 
mm. longi. Lamina basi 1--5 glandulosa, integra, chartacea, 
lanceolata, 5--8.5 cm. longa, 1.7--3.3 cm. lata, acuminata, 
basi rotundata. Racemi axillares solitarii, usque ad 5 om. 
longi; pedicelli usque ad 6 mm. longi. Sepala 5 vel 6, obtu- 
se triangularia, ca. 0.5 mm. longa. Petala 5 vel 6, usque ad 
2 mm. longa, 2-2 mm. lata. Stamina 2.5--3 mm. longa, anther- 
ae 0.5 mm. Stigma ca. 0.7 mm. diam. 

A glabrous tree, 7 m. high, 17 cm. diam.; branchlets 
slender, striate. Petioles slender, canaliculate, 5 to 7.5 
mm. long. Leaf blades entire, chartaceous, entirely glab- 
rous, lanceolate, 5 to 8.5 cm. long, 1.7 to 3.3 om. wide, 
acuminate, base rounded, costa and veins slightly impressed 
above, the veins scarcely discernible, costa elevated be- 
neath, the veins fine and inconspicuous, bearing at or near 
the base 1 to 5 glands about midway between costa and marg- 
in. Racemes axillary, solitary, up to 5 cm. long, entirely 
glabrous, drying reddish-black, pedunculate, bearing up to 
27 flowers, these not crowded; pedicels slender, up to 6 mm. 
long. Hypanthium turbinate, about 2.3 mm. long, 2.5 m. in 
diameter. Sepals 5 or 6, minute, obtusely triangular, about 
0.5 mm. long. Petals 5 or 6, suborbicular, up to 2 mm. long, 
2-2 mn. wide, subentire or remotely erose. Stamens 2.5 to 3 
mme long, slender; anthers about 0.5 mm. long. Stigma about 
O.7 mm. diam. 

Type in the University of Michigan Herbarium, Eizi Matuda 
2453, collected on Volcan de Tacana, Chiapas, Mexico, Aug. 
1938, alt. 1000--2000 m. 

P. Matudai is related to P. erythroxylon Koehne, P. axit- 
liana Standl., and P. Lundelliana Standl. 


CALLIANDRA CHIAPENSIS (Britt. & Rose) Lundell, comb. nov. 
Anneslia chiapensis Britt. & Rose, N. Amer. Fl. 23: 61. 
1928. 


ZANTHOXYLUM SONORENSE Lundell, sp. nov. , 

Frutex. Folia hirtella, imparipinnata, 2.5--5 cm. longa; 
foliola 5--9, oblanceolata, obovata, vel oblanceolato- 
oblonga, 0.4--1.8 om. longa, 0.25--0.6 om. lata, apice obtu- 
sa vel rotundata et emarginata, basi cuneata, crenulata. In- 
fructescentia racemosa, usque ad 1.5 om. longa. Sepala 4. 
Follioula 1 vel 2, sagen 3--3.5 mm. diam., 4--4.5 mm. 

9 


370 PHYTOLOGIA No. 11 


longa, stipitata. 

A shrub armed with short nearly straight slender prick- 
les; twigs slender, hirtellous. Leaves hirtellous, odd- 
pinnate, 2.5 to 5 cm. long, with prominently winged petiole 
and rachis; leaflets 5 to 9, sessile or nearly so, callosed 
at base, oblanceolate, obovate, or oblanceolate-oblong, 0.4 
to 1.8 om. long, 0.25 to 0.6 em. wide, apex obtuse or round- 
ed and minutely emarginate, base cuneate, margin crenulate, 
glandular, the glands chiefly below the crenatures, slightly 
paler beneath, hirtellous on both surfaces but sparingly so 
above. Inflorescence lateral, axillary, short racemose, up 
to 1.5 cm. long, the rachis hirtellous. Persistent sepals 4, 
minute, puberulent, ovate-deltoid, about 0.45 mm. long. Foll- 
icles 1 or 2, stipitate, the stipe 3 to 5.5 mm. long, the 
follicles ellipsoid, 3% to 4.5 mm. in diam., 4 to 4.5 m. 
long, glandular; seeds subglobose, about 2.5 mm. in diam., 
black, shining. 

Type in the University of Michigan Herbarium, Forrest 
Shreve 6727, collected 16 miles NE of Ures, on the road to 
Baviacora, Sonora, Mexico, Sept. 21, 1934, alt. about 900 m. 

Z. sonorense is very closely related to Z. Fagara (L.) 
Sargent, and probably represents a local desert segregate. 
It may be distinguished by the smaller hirtellous leaflets, 
puberulent sepals, and coarser pubescence throughout. The 
collection was distributed as a species of Bursera. 


TRAGIA POTOSINA Lundell, sp. nov. 

Volubilis; caules graciles. Fetiolus 2.3--4.5 cm. longus, 
gracilis; limbus oblongus vel ovato-oblongus, 8--12.5 om. 
longus, 3.0--7.3 cm. latus, acuminatus, basi cordatus, cren- 
ato-dentatus, basi 3- vel S-nervius; stipulae ca. 5 mm. lon- 
gee. Inflorescentiae eglandulosae, pedunculatae, pauciflor- 
ae, basi florem 2 unicum gerentes, ceterum é. Sepala o 5, 
stamina 4. Sepala f 6, subaequalia. Capsula parva. Semina 
cae 3 mm. diam., globosa. 

A vine; stems slender, rather sparingly hairy. Petioles 
slender, 2.3 to 4.5 cm. long, pilose. Leaf blades oblong or 
ovate-oblong, 8 to 12.5 cm. long, 3.5 to 7.3 cme wide, rath- 
er abruptly short-acuminate, base shallowly cordate and 3- 
or 4—veined, rather coarsely crenate-dentate, sparsely hisp- 
id above, hirtellous beneath with coarser hairs along veins. 
Inflorescence opposite the leaves, hirtellous, less than 2.5 
cm. long, with long peduncle, each bearing one pistillate 
flower with staminate flowers above. Bracts of pistillate 
flowers 3-parted, those of staminate flowers simple, linear- 
lanceolate, about 2 mm. long, sparingly hispid. Pedicels of 
pistillate flowers about 1 mm. long, those of staminate 
flowers subequal, erticulate slightly above base. Sepals of 
staminate flowers 3, ovate-elliptic, about 1.2 mm. long, 


POW Sale Se 


1940 Lundell, Flora of Mexico & Central America 371 


hispidulous outside. Stamens 4; filaments thick, apparently 
glandular, subequaling anthers. Sepals of pistillate flowers 
6, linear-lanceolate, up to 3 m. long, sparingly hispid, 
not glandular. Cvary hispid and glandular-hairy. Style con- 
nate to above the middle, obscurely rugulose. Capsule less 
than 1 cm. in diam. Seed globose, about 3 mm. in diam. 

Type in the University of Michigan Herbarium, C. L. Lund- 
ell end Ametia A. Lundell 7200, collected in second growth 
on hillside, at Tamazunchale, San luis Potosi, Mexico, July 
153, 1957, alt. 200 m. 

T. potosina is referable to the section Butragia, and 
it has 1 affinity with T. cordata Michx. The oblong leaves, e- 
glandular small inflorescences, long peduncles, short pedi- 
cels, and small capsules and seeds are distinguishing char- 
acteristics. 


DAVILLA MATUDAI Lundell, sp. nov. 

Frutex scandens. Rami scabriusculi. Folia petiolata, pet- 
iolo 7--25 mm. longo, parce strigoso, chartacea, haud scab- 
riuscula, elliptica vel obovato-elliptica, 6.5--12.5 cm. 
longa, 3.1--7.8 om. lata, integra, apice rotundata vel raro 
apiculata, basi subcuneata, costa et vena parce substrigosa, 
venis lateralibus 9--14. Inflorescentia paniculata, parva, 
dense puberula, pauciflora. Pedicelli usque ad 7 mm. longi. 
Sepala ciliolata, minutissime scabriuscula, suborbicularia, 
3 exteriora 2.4--3.5 m. longa, 2 interiora usque ad 8 mm. 
longa. Petala 5, obovata. Stamina usque ad 5 mm. longa. 


_Ovaria 1 vel 2, glebra. 


A woody vine. Stems at first zigzag, slender, brown and 
puberulent, with age straight, dark red and finely scabrous. 
Petioles sparingly strigose, narrowly winged, usually 7 to 
12 mm. long, sometimes up to 25 mm. long. Leaf blades chart- 
aceous, smooth, elliptic or obovate-elliptic, 6.5 to 12.5 
em. long, 3.1 to 7.8 cm. wide, entire, apex rounded or rare- 
ly apiculate, bese subcuneate and decurrent, sparingly hairy 
along costa above, and beneath along costa and main lateral 
veins, the hairs usually subappressed, costa and veins plane 
or slightly impressed above, prominent beneath, the main 
lateral veins 9 to 14, reticulation fine. Flowers in small 
panicles, the panicle branches densely puberulent, rather 
crowded. Pedicels slender, puberulent, up to 7 mm. long. Se- 
pals microscopically scabrous, ciliolate, not pubescent, 
suborbicular, the smaller 3 from 2.4 to 3.5 mm. long, the 
inner 2 up to 8 mm. long, the innermost with strongly re- 
curved margin. Petals 5, obovate, apparently emarginate. 
Stamens up to 5 mm. long, the filaments enlarged above, the 
anthers ovate-elliptic. Ovaries 1 or 2, glabrous, l-celled, 
2-ovulate; style elongate, 3 to 3.5 mm. long; stigma large, 
discoid. Fruits unknown. 


372 PHYTOLOGIA No. 1l 


Type in the University of Michigan Herbarium, Eizi Matuda 
3647A, collected in forest, Javalinero, near Palenque, Chia- 
pas, Mexico, July 6--9, 1939. 

According to description, the affinity of D. Matudai is 
with the Brazilian D. multiflora St. Hil. 


GILIBERTIA MATUDAI Lundell, spe nov. 

Arbor parva, 5 me alta, glabra. Petioli 0.8--5 om. longi. 
Lamina membranacea, oblanceolato-oblonga vel oblonga, 10-=-21 
em. longa, 4--9 cm. lata, apice obtuse acuminata vel late 
obtusa, basi cuneata, prominente sinuato-dentata. Umbellae 
racemosae, terminales. 

A small tree, 5 m. high, glabrous throughout. Petioles 
slender, striate, inconspicuously canaliculate, 0.8 to 5 cme 
long. Leaf blades membranaceous, oblanceolate-oblong or ob- 
long, 10 to 21 cm. long, 4 to 9 om. wide, apex obtusely 
short-acuminate or bluntly obtuse, base broadly cuneate, ob- 
tuse to acute, margin prominently sinuate-dentate with re- 
mote coarse obtuse or rounded teeth, pinnately veined, costa 
nearly plane above, elevated beneath, veins and veinlets 
prominulous beneath, reticulate on both surfaces. Umbels ra- 
cemose, terminal, 6 to 9 in an inflorescence; the inflores- 
cences less than 3.5 cm. long, short-pedunculate; bractlets 
fimbrillate; stalks of the umbels up to 1.5 om. long, bi- 
bracteolate near the middle. Pedicels slender, about 5 mm. 
long. Flowers S- or 6-merous. Calyx tube about 2.3 mm. long, 
5- or 6-denticulate, the teeth deltoid, acutely apiculate. 
Petals thick, ovate-deltoid, 1.7 to 2 mm. long, acute. Anth- 
ers about 1 mm. long, equaling filaments. 

Type in the University of Michigan Herbarium, Eizi Matuda 
3152, collected in advanced forest at Reforma, near Balan- 
can, Tabasco, Mexico, May 22--26, 1939. 

Although G. Matudai obviously belongs to the complex of 
G. arborea (L.) March., it may be readily distinguished by 
its coarsely sinuate-dentate leaves. 


ADDITIONAL NOTES ON THE GENUS AEGIFPHILA -- VI 


Harold N. Moldenke 


Ys 


The following notes constitute a continuation of those 


1940 Moldenke, Notes on Aegiphila 373 


published in Phytologia 1: 182--208, 222--240, and 248--272 
(1937), 289--304 (1938), and 364--368 (1939). The conclusion 
of the alphabetized list of citations [additional to those 
published in Brittonia 1: 472--477 (1934) and Phytologia 1: 
301--304 (1938) and 364--368 (1939)] follows: 


Mutis 305 (mollis, cotype coll. of A. Mutisii), 423 (mol- 
lis, cotype coll. of A. Mutisii), 782 (quinduensis), 857 
(reticulata, type colle), 985 (reticulata, type coll.), 2332 


(bogotensis), 3660 (mollis, cotype coll. of A. Mutisii), 
5191 (truncata, type colle), sene (bogotensis, cotype coll.). 


Natal Herb. 10,208 (panemensis); Nelson, E. W., 4245 
(Deppeana, cotype coll. of A. pacifica), 4254 (Deppeana, co- 
type coll. of A. pacifica); Newman sen. (conturbata, type 
coll.); Nicholls 94 (martinicensis); Niederlein 1705 (Hass- 
leri); Nyst sen. (Hassleri). 

Orcutt 3057 (Deppeana), 3418 (Deppeana), 5622 (trifida); 
Grsted 11,178 (panamensis), 11,180 (De eana), sen. [1846] 
(martinicensis); Otero 78 (martinicensis), 199 (martinicens- 
is), 308 (martinicensis). 

Pabst 435 (australis); Paul 153 (panamensis); Pavon sen. 
{"Mexique"] (Pavoniana); Peckolt 422 (Sellowiena); Pennell 
3185 (Pennellii, type coll.), 7097 (bogotensis); Pennell, 
Killip, & Hazen 8667 (caucensis, type coll.); Perrottet s.n. 
[1820] (villosa), sene [1819] (laevis), sen. [1820] (laev- 
is), sen. [18 Juin 1824] (martinicensis), sen. [Juillet 
1841] (martinicensis), sen. [Jardin des Fl. de Paris, 1818] 
(martinicensis); Picard sen. (martinicensis); Pickel 526 
(pernambucensis), 3030 (pernambucensis), 3042 (pernambucen- 
sis, type colle), 3642 (racemosa); Pittier, H., 288 (odonto- 
hylla, type coll.), 6782 (elata), 6815 (panamensis, type 
colle), 7584 (aculeifera), 8257 (hirsutissima, type coll.), 
8806 (quinduensis), 8806a (elata), 10,645 (pendula), sen. 
(Herb. Instit. Physioo-geogr. Nat. Costaric. 16,711] (anoma- 
la, type coll.), sen. (Herb. Instit. Physico-geogr. Nat. 
Costaric. 16,034] (costaricensis), sen. (Herb. Instit. 
Physico-geogr. Nat. Costaric. 12,017] (elata); Pittier & 
Tonduz 6782 (elata), sen. (Herb. Instit. Physico-geogr. Nat. 
Costaric. 9167] (costaricensis, type coll.); Plée 172 (mart- 
inicensis), sen. (elata); Poeppig 2158 (cordata, type coll.), 
2314 (chrysantha, logotype aah: Bene [Collares] (macran- 
tha); Fohl 1022 (splendens, type coll.), 4392 (Riedeliana, 
cotype coll.), sen. (Herb. Imp. Vien. 151] (mediterranea), 
gene (Herb. Imp. Vien.] (Lhotzkiana), sen. (Luschnathi); 
Poiteau een. (laevis), sen. (villosa); Ponthieu sen. [Ind. 
occid.] (martinicensis); Potter 5154 (laeta); Puiggari 3200 
(obducta), 3225 (obducta); Purdie sene [Port Royal Mtns., 
Aug. 1843] (trifida), een. (Mt. Diablo, April, 1844] (foeti- 
da), een. (Summit of Quindiu] (bogotensis), sen. (bogotens- 


3574 PHYTOLOGIA No. 11 


is), sen. (elata); Purpus 6982 (falcata), 7521 (falcata). 

Quentin 17 (martinicensis). 

Raben 749 (Luschnathi), 842 (casseliaeformis, type coll.), 
876 (verticillata), sen. (Iuschnathi), sen. (vitelliniflo- 
ra); Raimondi 578 (filipes), 981 (filipes); Ramage sen. [21 
Jan. 1888] (pernambucensis), s,n. [April 24, 1888] (martini- 
censis), sen. [Auge 22, 1888] (martinicensis), sen. [Sept. 
23, 1888] (martinicensis), sen. (martinicensis); Ravn sen. 
(martinicensis); Read sen. (martinicensis); Regnell 156 
(luschnathi), 1.184 [1845] (Sellowiana), 1.184 [1866] (Sell- 
owiena), 1.184 [1877] (Sellowiana), I.310 [1845] (verticill- 
ata), 1.310 [1862] (verticillata), 1.310 [1866] (verticille- 
ta), 1I.310x [1865] (Ihotzkiana), 1.310x [1867] (Lhotzkiana), 
11.310 [1856] (verticillata), II.310 [1874] (verticillata); 


Rehder sen. (elata); Richard, Le C., sen. (glandulifera var. 


ramidata, type coll.), sene (laevis), sene [S.-D.] (nervo- 
sa), sen. (martinicensis); Ridley, H. N., sen. (martinicen- 
gis); Riedel & Luschnath 1812 (Sellowiana); Riedlé 202 (mar- 
tinicensis), sen. [Herb. A. Le Jussieu 5034a] (martinicens- 
is), sen. (martinicensis); Riley, L. A. M., 64 (laxiflora); 


Rimbach 118 (monticola, type coll.), 234 (Rimbachii, type 


colle), 466 (bogotensis), 616 (ferruginea); Robert, A., 434 
(verticillata); Rodriguez sen. [Lillo 10,448] (mediterran- 
ea); Roig 1203 aatet: Rudio sen. (vitelliniflora); Ruiz 


187 (multiflora), 188 (cordifolia, type coll.); Ruiz & Pavon 
Sen. [Mina, Panatahue] (cordifolia, type colle), sene [Pent- 


ahua & Chichao] (integrifolia, type coll. of Callicarpa 

lobiflora), sen. [Peruvia et Chili] (integrifolia), sen. 
fimacc cases & Pallao] (multiflora, type colle), sen. (mul- 
tiflora); Rusby, He He, 2472 (filipes, type coll. of A. ob- 
ee 2619 (multiflora, type coll. of Clerodendron 
bolivianum); Rusby & Pennell 701 (montana, type coll.); Rus- 
by & Squires 316 (perplexa, type coll.); Butten-Pekelharing 
40 (racemosa); Ryan sene (Deppeana), sen. (integrifolia), 6. 
ne (martinicensis). 

Saer 602 (laeta); Sagot 473, in part (racemosa), 473, in 
part (glandulifera var. pyramidata), sen. [Boura, 1858] 
(laevis), sen. [Cayenne, Fevrier 1859] (laevis), sen. [Cay- 
enne, Mars 1859] (villosa), sen. [Karovany, 1854--58] (race- 
mosa), sen. (membranacea); Sagra 360 (elata); Saint-Hilaire 
2229 (Sellowiana), 2229 ter (Sellowiana), A.361 (fluminen- 
sis), A*.663 (fluminensis), A!.665 (fluminensis), C.50 (med- 
ullosa, type colle), C'.1016 (verticillata), C#.1721 bis 

Luschnathi), C2.1774 (obducta), sen. Litabura] (Sellowia- 
na), Sen. (Lhotzkiana); Salzmann 432 (Lhotzkiana), sen. 
11831] (Unotzkiana), sen. (Lhotzkiana); Sartorius sen. (Dep- 
peana); Schiede 1165 (Deppeana, type ooll.), sen. [Estero, 
Jan. 29] (Deppeana); Schimpff 252 (monticola), 267 (bogoten- 
sis), 279 (chrysantha), 1003 (Schimpfii, type coll.); Schipp 


Me ate 


1940 Moldenke, Notes on Aegiphila 375 


1083 (monstrosa); Schlim 688 (longifolia, type coll.); 
Schomburgk, M. Re, 404, in part (guianensis, type coll.), 
404, in part (integrifolia), 772 errr type coll.); 
Schott 4918 (Luschnathi); Schiich sen. [Sebastianopolis] 
(vitelliniflora), sen. (Sellowiana); Schunke 443 (pulcherri- 
ma, type colle); Schwacke 1465 (brachiata), 1939 (obducta), 

35 (bracteolosa), 5381 (fluminensis), 6594 (verticillata), 
9556 (obducta), 9941 (verticillata), 11,264 (Sellowiana), 
12,966 (Riedeliana), 13,098 (obducta), 13,726 (verticille- 
ta); Seemann 87 (magnifica), 335 (glandulifera); Seitz 61 
(obovata), 92 (obovata); Sellow 9 vcsenlanst, 2189 (cgrave- 
olens), 5122 (verticillata, type coll. of A. tomentosa), s. 
Ne (Brasilia ] (Ftuminanata), BeNe (vitelliniflora), BeNe 
[flowers] (Sellowiana, cotype coll.), sene [fruit] (Sellowi- 
ana, cotype coll.), sen. (brachiata), sen. (laevis), sen. 
(ihotzkiana), sen. (obducta), sen. (paraguariensis); Sessé, 
Mocino, Castillo, & Maldonado 603 (Deppeana), 1074 (Deppea- 
na); Sinclair sen. [Salango Isl.] (glomerata, type coll.); 
Sintenis 6492 (martinicensis), sen. [L.89] (martinicensis), 
e.ne [1895] (martinicensis); Skinner sen. (monstrosa); 
Skutch 2680 (panamensis); Smith, G. W., 93 (martinicensis); 
Smith, H. H., 329 (mollis var. intermedia, type colle), 330 
(laeta, type coll. of A. stricta); Smith, J. D., 2111 (fale- 
ata, type coll.); Snethlage 109 (racemosa); Sodiro 22 (ferr- 
u on), 125 (ferruginea); Soubirou sen. [Prés Cayenne] 
(laevis), sen. (laevis); Spruce 342 (parviflora), 589 (par- 
viflora, type coll.), 691 (integrifolia), 1013 (bracteolos- 
a), 1283 (bracteolosa), 2296 (Spruceana, type colle), 3113, 
in part (guienensis), 3113, in part (integrifolia), 4275 
(peruviana, type coll.), 5473 (ferruginea, type coll.), 9473 
{probably an error for 5473], sen. [Aug. 1858] (ferruginea), 
sen. [In vicinibus Sentarem] (integrifolia); Standley, P.C., 
37,570 (Standleyi, type coll.); Standley & Valerio 44,597 
(costaricensis), 44,606 (costaricensis); Stehlé 1247 (mart- 
iniaensis), 2127 (martinicensis); penlaser 3186 (bolivia- 
na), 5066 (boliviana, cotype coll.), 6437 (boliviana, cotype 
colle), 14,781 (mollis); Stephan sen. [1843] (paraguarien- 
sis); Stevenson, D., 5 (pauciflora, type coll.); Stork 2230 
(aculeifera), 2290 (anomala); Suringar sen. [Puerto Cabello ] 
(elata); Swabey sene [Trin. Bot. Gard. Herb. 12,232] (integ- 
rifolia); Swainson sen. (vitelliniflora), sen. (pernembucen- 
is), Sen. (mediterranea); Swartz sen. [Jamaica] (Swartzian 
type coll.), sen. [Jamaica] (trifida, type coll.), sen. 
(Jamaica] (foetida, type coll.), sen. (glata, type coll.), 
sen. (nervosa, type coll.), sen st pO 

Tafalla sen. [Chicoplaya] (insignis, type colle); Talbot 

sene (laevis); ‘ate 162 (Surfaceana, type coll.); “errasson 
62 (martinicensis); Tesamann 5963 (bracteolosa); Thompson, 
W. Je, 6493 (elata), 8012 (elata); Tonduz 929%3b (falcata), 


376 PHYTOLOGIA No. ll 


13,629 (magnifica), sen. (Herb. Instit. Physico-geogr. Nat. 
Costaric. 13,629] (magnifica), sen. [Herb. Instit. Physico- 
geogr. Nat. Costario. 6782] (elata); Triana 375 (integrifol- 
ia), 2083, in part (Lehmannii), 3712 (grandis), 3713, in 


part (glandulifera var. pyramidata), 3713, in part (guianen- 
sis), 3713, in part (elata), 3713, in part (Lehmannii), 
3713, in part (membranacea), 3713, in part (mollis), 3743 
(bogotensis), sen. (grandis); Trin. Bot. Gard. Herb. 1899 
(macrantha), 2389 (obovata, type coll. of A. dubia); Troll 
1704 (integrifolia); Tirckheim 4013 (fasciculata, type 
coll.), 7961 (elata). 


Ule 335 (Lhotzkiana), 1520 (australis, type coll.), 4436 
(Luschnathi), 4854 (mediterranea), 8293 (filipes), 9718, in 
part (elegans), 9718, in part (ovata), 9720 (cordata), 9721 
(scandens, type coll.), 9723 (filipes), 9859 (cuneata), sen. 
(obducta); United States Exploring ed. [Wilkes] sen. [Rio 
de Janeiro] (brasiliensis, type coll.), sene [Rio de Janei- 
5 (fluminensis); Usteri 20 (verticillata), 21 (verticilla- 
ta). 
~~ Vehl sen. (1796; Herb. A. L. Jussieu 5036] (integrifol- 
ia); Valerio, J., 148 (Valerii, type coll.); Versuchsenstalt 
Kamerun 270 (mediterranea); Vogl 1177 (mollis), 1179 (mol- 
lis), 1180 (mollis), 1181 (mollis), 1183 (mollis); Von Rohr 
97 (Deppeana), sen. [Cayenne] (villosa), sen. (Deppeana), s. 
ne (elata), sen. [Cayenne] (laevis); Von Seneloh 139 (flun- 
inensis). 

Waby 72 (martinicensis); Wachenheim 175 (villosa), 188 
(membranecea), 484 (macrantha), 489 (macrantha); Wagner sen. 
(Herb. Monac. 1007] (Deppeana); Warming 71 (barbadensis), 
101 (barbadensis, type coll.), 291 (vitelliniflora), Sen. 
(Lagoa Santa] (verticillata), sen. [Lagoa Santa] (Sellowia- 
na), sen. [Juli 1866] (fluminensis), sen. [Ad Rio de Janei- 
ro] (fluminensis), son. (Sellowiana); Weberbauer 4667 (lon- 

ipetiolata, type coll.); Weddell 40 (fluminensis), 95 
(riaianats), 578 (vitelliniflora); "West Indies, Nov. 17- 
17" (martinicensis vare oligoneura); Widgren 1177 (Sellowi- 
ana), 1222 (verticillata), 1223 (mediterranea), II.117 
(Sellowiana), 11.1176 1/2 (Sellowiana), sen. (mediterranea); 
Wiles sen. (elata); Williams, Ll., 5054 (Smithii, type 
coll.), 9120 (monstrosa), 9145 (elata), 9271 (elata), 9566 
(elata); Williems, R. 0., sen. [Trin. Bot. Gard. Herb. 
11,115] (obovate), sen. [Trine Bot. Gard. Hert. 12,026] 
(obovata), sen. (Trin. Bot. Gard. Herb. 12,180] (obovata); 
Williems, Freeman, & Cheesman s.n. [Trin. Bot. Gard. Herb. 
11,329] Antegrifolis); Wilson, N., 224 (elata); Winkler, 
H., 628 (mediterranea); Wolle sen. (Swartziana); Wright, W., 
sen. (elata), sen. (foetida), sen. (martinicensis); Wull- 
schlégel 405 (laevis), 4235 (martinicensis), 424 (martinicen- 


sis), 974 (elata), 1364 (elata), 1983 (laevis). 


— 


1940 Moldenke, Notes on Aegiphila 377 
Yuncker 4749 (elata). 


Three additional generic synonyms are Egiphylla Jacq. ex 
Moldenke, Prelim. Alph. List Inyalid Names 25, in syn. 
(1940), Stigmatococca Mart. ex Moldenke, Prelim. Alph. List 
Invalid Nemes qo, in syn. (1940), and Echiphylla Jacq., in 
herb. Gerth van Wijk in his "A Dictionary of Plant Nemes", 
page 30 (1911), records two additional common names for men- 
bers of this genus: "Ziegenbaumchen" and "geitenboompje". 
The Ducke sen. [Herb. Rio de Janeiro 35,661] distributed as 
a species of Aegiphila by the collector, is actually Rudgea 
Krukovii Standl. in the Rubiaceae. Standley in Field Mus. 
Publ. Bot. 10: 334 (1931) uses the name “aegiphiles" as a 
common name for members of this group, although "goatwoods" 
would certainly be preferable. His description of the group 
as being "shrubs or small trees with entire leaves", with 
"exserted stamens", "a corolla with spreading limb", and 
fruit which "is a fleshy drupe", is not true for the genus 
as a whole. Many species are low bushes, vines, or tall 
trees; most species have two types of flowers, one type 
with included stamens; the corolla is often infundibular; 
and the fruit is often dry and not at all fleshy. The gen- 
eric description in Field Mus. Publ. Bot. 18: 993 (1938) is 
more accurate, but still is misleading in many respects. If 
generic descriptions in floras of circumscribed regions are 
drawn up from only the species represented in that region 
and are meant to apply only to those species (which may well 
be exceptional or even aberrant ones! ), the author ought to 
plainly state this fact. Unless such a qualifying statement 
is given, it is natural for other workers to regard such 
generic descriptions as being actually generic descriptions 
and not just compilations of the characters of the few spec- 
ies of the genus inhabiting that particular area. 

Since the bibliographic report given in Mhytologia 1: 289 
(1938) was drawn up, ten more papers referring to the genus 
have been reviewed and 4 more contributors added. It is 
worth noting that the generic name Amerina, cited by me in 
Brittonia 1: 250 and 280 (1934), should be accredited to 
"P, DC." The Gentle 3242 and 3246 distributed as a species 
of sonia by Lundell, are actually Dermatocalyx parvi- 
florus Mrst. in the Scrophulariacese. 

The herbarium abbreviations herein employed are those ex- 
plained in my original monogreph of the genus in Brittonia 
1: 249--250 (1934) and in previous supplements in Phytolog- 
fa. Additional abbreviations herein used for the first time 
are "Ar" = United States National Arboretum, Washington, D. 
0.3 "Dp" = DePauw University, Greencastle, Indiana; "Ha" = 
Colegio de La Salle, Vedado, Havana, Ouba; "Jo" = J. Quatre- 
casas herbarium, Instituto Botanico, Bogota, Colombia; "Lu"= 


378 PHYTOLOGIA No. 11 


Botanisk Museum, University of Lund, Lund, Sweden; and "Po"= 
Pomona College, Claremont, California. 


49a. AEGIPHILA ACULEIFERA Moldenke. 

The species is described by A. Snith from living material 
as a tree 4--10 m. tall, its base to 42 om. in diameter, its 
bark light-browm, with "reticulation" or distantly spaced 
large raised "dots"; terminal twigs with "lepidote scales"; 
cambium—layer pale-green; leaves rather flaccid or membran- 
ous and basally stiffened, glabrous, slightly shining and 
"polished" above, the venation "rather reticulated on under 
side"; inflorescence mostly terminal; peduncles and pedicels 
bright-green or gray-brown, the former with "lepidote scales; 
flower-buds clavate, opening campanulate, the small ones 
gray-brown; calyx bright- or pale-green, "pointed", short- 
pubescent; petals “opening flat", "indifferently 4 or 5", 
pure white; stamens short, shorter than the style, included; 
style white, elongate, longer than the stamens, exserted; 
immature fruit hard and green, round, flat-topped. Standley 
in Field Muse Publ. Bot. 18: 994 (1938) describes the leaves 
as obovate-oblong, the cymes mostly as long as the petioles 
or shorter, densely stellate-furfuraceous, the calyx some~ 
what bilabiate, its lobes 2 mm. long, and the corolla-tube 8 
mm. long, with its lobes 6 m. long. He comments on the fact 
thet the recurved prickles which cover the branchlets and 
peduncles of the type collection are not in evidence in the 
Costa Rican material. They are, however, very numerous and 
conspicuous on the peduncles and inflorescence-branches of 
the Sneidern collection cited below, on which they are also 
found to some extent on the tips of the twigs and on the 
petioles. 

The species has been collected at altitudes of 1500 m. in 
Colombia and 1500-2200 m. in Costa Rica, and in anthesis al- 
so in April and August. A. Smith reports the style as "bi- 
furceted", but actually it is the stigma which is bifid. He 
states that the species inhabits the subtropical zone in 
Costa Rica, found at the edges of forests in semi-shade, in 
wet mouldy soil mixed with loam. Sneidern reports it as an 
inhabitant of the primeval forest. 

Additional citations: COSTA RICA: Alajuela: Skutch 3255 
(N); A. Smith (N--3), 1037 (N--3). COLOMBIA: El Cauca? 
Sneidern 1661 (S). 


42a. AEGIPHILA AMAZONICA Moldenke, Geogr. Distrib. 24, nom. 
nud. (1939), spe nov. 

Frutex vel arbor; ramulis argute tetragonis dense ad- 
presso-strigosis; petiolis dense flavido-strigosis; laminis 
submembranaceis ellipticis vel obovatis acutis vel breviter 
acuminatis, integris, ad badim cuneato-attenuatis, subtus 


1940 Moldenke, Notes on Aegiphila 379 


densiuscule breviterque pubescentibus vel velutinis; inflor- 
escentiis axillaribus cymosis dense multifloris. 

Shrub or small tree to 5 m. tall; brenchlets and twigs 
medium-stout, very sharply tetragonal, often pronouncedly 
flettened, densely appressed=pubescent with short and strig- 
ose or felt-like grayish or yellowish pubescence; pith 
large; nodes not annulate, often somewhat flattened; buds 
densely yellow-velutinous or -villous; principal internodes 
1.5--10 cm. long, sometimes abbreviated to 3 m.; leaves de- 
cussate-opposite; petioles rather slender, 0.7--3 cm. long, 
or very greatly abbreviated on young twigs, densely appress- 
ed-pubescent with short an yellowish strigose hairs; blades 
submembranous or thin-chartaceous, dark-green and brunnes-~ 
cent (in drying) above, lighter beneath, elliptic or obov- 
ate, 3.5--18 cm. long, 2--6 om. wide, acute or short-acumin- 
ate at apex, entire, cuneately narrowed and long-attenuate 
into the petiole at base, finely puberulent above when 
young, glabrescent when mature, rather densely short-pubes- 
cent with grayish or yellowish hairs beneath, subvelutinous 
when immature, the hairs on the lower surface bulbous-based; 
midrib rather stout, flat above, very large and prominent 
beneath; secondaries slender, 8--14 per side, arcuate- 
ascending, flat or subprominulous above, prominulous be- 
neath, arcuately joined in many loops near the margins be- 
neath; inflorescence axillary, opposite, sparse, cymose; 
cymes solitary in each axil, densely many-flowered, 3--6.5 
cme long, 2--5 cm. wide, many times dichotomous, its branch- 
es similar to the peduncles in color, texture, and pubes- 
cence; peduncles slender, 1.5-—4 om. long, densely short- 
pubescent with yellowish or grayish strigose hairs; pedicels 
slender, elongate, 3--5 mm. long, densely strigose-pubes- 
cent; bracts and bractlets absent; prophylla minute, hidden 
by the pubescence; calyx infundibular, about 5.8 mm. long, 
1.2 mm. wide to above the ovary, about 3 mm. wide at apex, 
lightly appressed-puberulent or short-strigose on the outer 
surface and often marked with scattered glandular disks a- 
bove, its rim deeply 4~lobed, the lobes equal, regular, a- 
bout 1.9 mm. long and 1.4 mm. wide, acute at apex; corolla 
white, hypocrateriform, its tube rather slender, about 6.3 
mm. long, 0.9 mm. wide at base, abruptly ampliate to 5 mn. 
at apex, glabrous, the limb spreading, 4-parted, its lobes 
ovate-lingulate, about 3 mm. long and 1.5 mm. wide at base, 
obtuse at apex; stamens 4, inserted about 0.5 mm. below the 
mouth of the corolla~tube, long-exserted; filaments filiforn, 
5--6 mm. long, glabrous; anthers oblong, about 1 m. long 
and 0.5 m. wide, dorsifixed just below the middle, bilobed 
at base; pistil slightly exserted; style capillary, about 5 
mn. long, glabrous; stigma bifid, its branches about 1 mm. 
long; ovary subglobose, about 0.8 mm. long and wide, glab- 


380 PHYTOLOGIA No. 11 


rous e 

The type of this species was collected by Joa&o Geraldo 
Kuhlmann [Herb. Rio de Janeiro 22,546] in a secondary forest 
at Manés, Amazonas, Brazil, on March 17, 1924, and is depos- 
ited in the Britton Herbarium at the New York Botanical Gar- 
den. The species is closely related to A. integrifolia. 

Citations: BRAZIL: Amazonas: J. Ge Kuhlmann sen. (Herb. 
Rio de Janeiro 3386] (N), sen. (Herb. Rio de Janeiro 22,546] 
(N--type); Para! Ducke s.n. (Herb. Rio de Janeiro 14,281; 
Herb. Amaze Mus. Para. 4738] (N), s.n. [Herb. Rio de Janeiro 
18,952] (N). 


6. AEGIPHILA ANOMALA Pittier. 

Standley in Field Mus. Publ. Bot. 18: 994 (1938) gives 
the following additional characters: a shrub or small tree 
of 4.5 m., the branches densely ochraceous-puberulent; 
leaves short-petiolate, the blades oblanceolate or oblong- 
obovate, mostly 10--20 cm. long, acute or abruptly short- 
acuminate, long-attenuate to the base, densely and minutely 
puberulent on both surfaces; cymes small and dense, 5 cm. 
long or less, the branches densely puberulent; calyx densely 
puberulent, 3--5S-lobate; corolla salver-form; fruit globose, 
about 1 cm. in diameter. He quotes Austin Snith as stating 
that the species grows at altitudes of 1700--1950 m., on 
clay ridges in open forests; a tree 11--15 m. tall, of open 
growth, the trunk 45--60 cm. in diameter, the bark browm, 
cork-like in structure, well-sutured, thick; cambium-layer 
cream—ysllow; leaves soft and rather lax, light-green, vel- 
vety to the touch; flower-buds buffy-~yellow; and the corolla 
pure white. The Austin Smith material, siad by Standley to 
represent this species and collected about Zarcero, has not 
been seen as yet by the present writer. 


70. AEGIPHILA BARBADENSIS Moldenke. 
Additional citations: BARBADOS: Warming 101 (Mi--photo of 


type). 


3. AEGIPHILA BOGOTENSIS (Spreng. ) Moldenke. 

The binomial "Amerina tomentosa", cited by me in Britton- 
ia 1: 283 (1934), should be accredited to "(H.B.K.) P. DC." 
The Schimpff 252 material in the Berlin ( 2 sheets), Deless- 
ert, and Britton herbaria, cited in Phytologia 1: 188 (1937) 
as this species, is actually A. monticola. 

Additional citations: COLOMBIA: Department undetermined : 
Mutis 5774 (W). 


80. AEGIFHILA BOLIVIANA Moldenke. 
This binomial is erroneous written "Aegiphila bolivana" 
in Brittonia 1: 391 (1934). The species has been collected 


1940 Moldenke, Notes on Aegiphila 381 


at 450 m. altitude, in enthesis in September, and has been 
confused with A. martinicensis. 

Additional citations: BOLIVIA: Santa Cruz: Steinbach 2799 
(N), 3116 (N), 5066 (Mi--photo of cotype). 


23. AEGIPHILA BRACHIATA Vell. 

An additional synonym is Aegiphila Glazioviana Taub. ex 
Moldenke, Prelim. Alph. List Invalid Names 2, in syn. (1940). 
The species is listed as A. triantha by Herter in Revist. 
Sudam. Bot. 4: 185 (1937). 


120. AEGIPHILA BRACTEOLOSA Moldenke. 

The flowers of this species are described as white. It 
has been collected in anthesis in March, 

Additional citations: BRAZIL: Amazonas: Ducke 444 (S); 
Spruce sen. [In vicinibus Barra] (Lu). 


99. AEGIPHILA CANDELABRUM Brigq. 
The species has been collected in anthesis in November. 
Additional citations: PARAGUAY: Hassler 7974 (F). 


40. AEGIPHILA CAPITATA Moldenke. 
Additional citations: BRAZIL: Sao Paulo: Burchell 3547 
(Mi--photo of isotype). 


98. AEGIPHILA CHRYSANTHA Hayek. 

An additional synonym is Aegiphila lutea Poepp. ex Mold- 
enke, Prelim. Alph. List Invalid Names 2, in syne (1940). 
The specimen cited below is noteworth in possessing a binary 
leaf -- one with two apexes! 

Additional citations: ECUADOR: Manabi: Zggers 14,838 (Lu) 


112. AEGIPHILA CORDATA Poepp. 
Additional citations: PERU: Loreto: Poeppig 2158 (Mi-- 
photo of type). 


112a. AEGIPHILA CORDATA var. COLOMBIANA Moldenke, Geogr. 
Distrib. 18, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit foliis dist- 
incte petiolatis; laminis ad basim acutis, ad apicem longe 
acuminatis; cymis parvioris minus densis, distincte stipita- 
tis non valde bracteolatis; floris albis. 

This variety differs from the typical form of the species 
in its leaves being acute at base, long-acuminate at apex, 
distinctly petiolate, the hairs on the branchlets shorter, 
the cymes smaller, less dense, distinctly stipitate, not 
plainly bracteolate, and the flowers: white. 

The type of this variety was collected by Oscar Haught 
(no. 1885) in the vicinity of Barranca Bermeja, alt. 100-- 


382 PHYTOLOGIA No. 11 


500 m., Magdalena valley, between Sogamoso and Colorado Riv- 
er, on June 16, 1936, and is deposited in the Britton Herb-— 
arium at the New York Botanical Garden. It is described as 
an abundant scrambler in second-growth forests, about 10 m. 
tall, with rather conspicuous white flowers. 

Citations: COLOMBIA: Santander Sur: Haught 1885 (N--type). 


113. AEGIPHILA CORDIFOLIA (Rufz & Pav.) Moldenke. 

The species has been confused with and distributed as A. 
Mutisii H.B.K. 

Additional citations: PERU: Huanuco: Macbride 3922 (N); 
Department undetermined: Ruiz & Pavon sen. [Mina, Panatahua] 
(Mi--isotype). . 


10. AEGIPHILA COSTARICENSIS Moldenke. 

A tree to 8 m. tall, the trunk to 15 cm. in diameter at 
breast height; branches glabrous; leaf-blades attenuete to 
the base; corolla white. 

The Tonduz and the Pittier & Tonduz specimens cited by me 
from Bocas del Toro, Panama, in Brittonia 1: 295 (1934) and 
Phytologia 1: 196 (1937), were actually collected in Carta- 
go, Costa Rica, according to a letter from Dr. Henri Pittier 
quoted under A. falcata in these supplementary notes, and 
the numbers are in each case Herb. Instit. Physico-geogr. 
Nat. Tostaric. numbers. This means that the species is as 
yet unknown from Panama. Standley in Field Mus. Fubl. Bot. 
18: 994--995 (1938) cites the type collection as "Pittier & 
Tonduz 9167". He gives the species' distribution in Costa 
Rica as "Pacific tierra caliente and probably also in the 
Atlantic". It ascends to 2400 feet in Guatemala, blooming in 
December. The Skutch collection cited below was originally 
distributed as "Vitex sp." 

Additional citations: MEXICO: Chiapas: Matuda 2101 (Mi, 
N). GUATEMALA: Quezaltenango: Skutch 2012 (N, W). 


35. AEGIPHILA CRENATA Moldenke. 
Additional citations: BRAZIL: Paranaé: Dusén 16,238 (Iu). 


9. AEGIFPHILA CUNEATA Moldenke. 
Additional citations: PERU: Loreto: Killip & Smith 28,386 
(Mi--photo of isotype). 


4). AEGIPHILA DENTATA Moldenke. 

The species is listed by Hoehne in his "Resenha Histori- 
ca para a Commemoragaéo do Vigesimo Anniversario da Seccao de 
Botanica e Agronomia annexa ao Instituto Biologico de Sao 
Paulo", pages 153 and 157 (1937). 


117. AEGIPHILA DEPPEANA Steud. 


; 
i 
; 


1940 Moldenke, Notes on Aegiphila 383 


The synonymous binomial, Aegiphila brachiata, cited by me 
in Brittonia 1: 450 (1934), should more properly be accred- 
ited to "Schlecht. & Cham." instead of "Cham. & Schlecht." 
The Dugand & Mina collection cited by me in Phytologia 1: 
291 (1938) as from en undetermined department of Colombia, 
was actually collected in the Panama Canal Zone, although 
the labels read "Juan Mina, Colombia". These collectors des- 
cribe the flowers as "small, tubular, reddish". Standley in 
Field Mus. Publ. Bot. 18: 995 (1938) states that the species 
is sometimes subscandent, with short-petiolate leaves, whose 
blades are cuneate at the base. 

Additional citations: PANAMA: Canal Zone: Dugand & Mina 
950 [Mus. Yale School of Forestry 32,382] (N, Y). COLOMBIA: 
Atlantico: Elias 1621 (N). 


27a. AEGIPHILA DUCKEI Moldenke, Geogr. Distrib. 24, nom. 
nud. (1939), sp. nov. 

Arbor; ramulis obtuse tetragonis densissime adpresso- 
pubescentibus glabrescentibus; petiolis abbreviatis strigo- 
sis plusminus merginatis; laminis tenuiter chartaceis obova- 
tis abrupte breviterque acuminetis integris, ad basim cunea- 
tis, supra plusminus pulverulentis, subtus minutissime pub- 
erulentibus; inflorescentiis axillaribus cymosis multiflor- 
is. 

Small tree; branchlets medium-slender, obtusely tetragon- 
al, very densely appressed=pubescent when young with yellow- 
ish felt-like strigose hairs, glabrescent in age; nodes not 
ampliate, not annulate; leaf-scars on older branchlets large 
and corky, semicircular; principal internodes 1--3 cm. long; 
leaves decussate-opposite; petioles rather slender, abbrevi- 
ated, 5--15 mm. long, flattened above, densely strigose- 
pubescent beneath, less densely so above, more or less marg- 
ined; blades thin-chartaceous, dark-green above, lighter be- 
neath, obovate, 7.5--13 cm. long, 3.5--6 cm. wide, abruptly 
short-acuminate at apex (the acuminetion itself sharp- 
pointed or rounded), entire, cuneate at base and prolonged 
into the margined petiole, more or less pulverulent above 
(sometimes with very tiny obscure hairs interspersed), very 
minutely puberulent beneath with tiny scattered appressed 
hairs; midrib rather stoutish, flat above, very large and 
prominent beneath; secondaries slender, 8--12 per side, ar- 
cuate-ascending, flat or subprominulous above, prominulous 
beneath, obscurely joined at the margins; inflorescence ax- 
illary, cymose; cymes solitary in the uppermost axils, oppo- 
site, 6--11.5 om. long, 2.5--3.5 om. wide, several times di- 
chotomous, many-flowered; peduncles very slender, elongate, 
3--6.4 cm. long, densely strigose-pubescent with whitish or 
yellowish appressed pubescence; pedicels slender, 1--45 mn. 
long, densely velutinous with short appressed heirs; bracts 


384 PHYTOLOGIA No. 11 


rether large, a pair at each principal cyme-furcation, stip- 
itate, oblanceolate, 1--2 cm. long, 2--3 mm. wide, lightly 
pulverulent-puberulent; bractlets numerous, conspicuous, 
spatulate, 4--8 mm. long, 1--1.8 mm. wide; calyx infundibu- 
lar, about 6.3 mm. long, about 1.7 mm. wide to above the ov- 
ary and 53 mm. wide at apex, densely strigose-pubescent, its 
rim truncete, 4-apiculate; corolla infundibular, white, its 
tube narrow-cylindric, about 10 mm. long and 1 mm. wide at 
base, abruptly ampliate to 4 m. at apex, densely pulveru- 
lent-puberulent outside (especially above the calyx), its 
limb 4-parted, its lobes regular, erect, ovate-lingulate, a- 
bout 4.9 mm. long and 2.4 mm. wide, subacute or obtuse at a- 
pex, densely pulverulent-puberulent outside; stamens 4, in- 
serted about 3 mm. below the mouth of the corolla-tube, 
long-exserted; filaments flattened, about 10 mm. long, glab- 
rous; anthers broadly oblong, about 1.7 mm. long and wide, 
dorsifixed below the middle, bilobed at base; pistil includ- 
ed; style capillary, about 5.4 mm. long, glabrous; stigma 
deeply bifid, its branches about 3.4 mm. long. 

The type of this species was collected by my esteemed 
friend and colleague, Dr. Adolfo Ducke [Herb. Rio de Janei- 
ro 35,662] -- in whose honor it is respectfully named -- in 
@ non-inundated forest at Igarapé Macacuny, near Cucuhy, on 
the Rio Negro, Amazonas, Brazil, on September 20, 1935, and 
is deposited in the Britton Herbarium at the New York Botan- 
ical Garden. It is apparently related to A. Lehmannii of 

Colombia. 
Citations: BRAZIL: Amazonas: Ducke sen. [Herb. Rio de 
Janeiro 35,662] (N--type). 


124. AEGIPHILA ELATA Sw. 

Stendley in Field Mus. Publ. Bot. 10: 334 (1931) and 18: 
995 (1938) and various collectors on their labels have added 
-a few more items to the composite description of the spec- 
ies: It is sometimes a medium-sized shrub of bushy habit of 
growth or a subscandent shrub; stem to 4 inches in diameter; 
branches sometimes long and trailing; leaf-blades varying to 
broadly elliptic, abruptly short-acuminate at apex, obtuse 
at base, glabrous or nearly so on both surfaces or appressed 
pilose beneath along the veins or almost glebrous; cymes of- 
ten forming a rather large and open panicle; corolla varying 
from cream-colored to yellowish-white, pale~yellow, or 
greenish-yellow; fruit bright-orange or deep-yellow, to 1 
em. in diameter, often borne in large clusters 4--6 inches 
long. It inhabits brushy slopes, forests, moist soil at the 
margins of swamps, wet thickets, high ridges, sides of 
creeks, savannas, advanced forests, wooded swamps, and 
coastal thicketse Gentle found it in "acahual" areas and 
Standley in “ierra caliente". It has been collected at alti- 


: 


a le ee ns 


1940 Moldenke, Notes on Aegiphila 385 


tudes of from 30 to 1000 m., in fruit in January, March, 
June to August, and October. It is called "tall aegiphila" 
by Lindley. Schipp describes the species as "common" at some 
localities in British Honduras and "occasional" in others. 
It is oultivated in the Plant Introduction Garden at Miami, 
Florida, source of seed unknowm. It is recorded by Yuncker 
in Field Mus. Publ. Bot. 9: 329 (1940). 

The Kegel 687 cited by Pulle in his Rnum. Vase. Pl. Sur- 
in. 403 (1906) is actually A. laevis. The Steinbach 3259 
cited by me in Brittonia 1: 466 (1934) and Phytologia 1: 199 
(1937) with a question-mark, proves on closer examination to 
be A. Herzogii. Therefore, A. elata is not now known to occ~ 
ur in Bolivia and should be removed from my lists of species 
know from that country. The binomial, Aegiphila macrophylla 
H.BeKe, cited by me in Brittonia 1: 462 (19354) as a synonym, 
was published in 1818, not in "1817" as stated on that page, 
and the "Aegiphila macrophylla Rich." on the same page 
should more accurately be accredited to "A. Rich." An addit- 
ional synonym is Aegiphila macrophtla HeBeK. ex Moldenke, 
Prelim. Alph. List Invalid Names 2, in syn. (1940). 

Additional citations: CUBA: Senta Clara: lama 493 (Ha); 
Oriente: Acuna 9866 (Es); Leén 10,872 (Ha), 12,199 (Ha); 
Leén, Clément, & Roca 10,431 (Ha); Lean & Seifritz 18,185 

N). MEXICO: Tabasco: Matuda 3031 (N), 3081 (N), 3406 (N). 
HONDURAS: Atlaéntida: Yuncker 4749 (Dp); Yuncker, Koepper, & 
Wagner S2tt (Dp, Mi, N). BRITISH HONDURAS: Gentle 2633 (N-- 
2), 2643 (N), 3047 (N). CULTIVATED: Florida: Dade Co.: Pope- 
noe 32 Corm-3.) 


121. AEGIPHILA ELEGANS Moldenke. 
aig citations: BRAZIL:-Amazonas: Krukoff 8701 
Kr )e 


101. AEGIPHILA ELONGATA Moldenke. 
Additional citations: BOLIVIA: La Paz: Buchtien 1645 (Mi 
--photo of type). 


21lb. AEGIPHILA EXIGUIFLORA Moldenke, Geogr. Distrib. 24, 
nom. nud. (1939), sp. nov. 

Arbor; ramulis obtuse vel subacute tetragonis dense furf- 
uraceo-farinaceis vel tomentosis; petiolis valde abbreviatis 
vel subobsoletis; laminis firme chartaceis late ellipticis 
vel oblanceolatis vel ovatis acutis vel brevissime acuminat- 
is integris, ad basim-longe cuneatis vel abrupte acuminatis, 
supra adpresso-pubescentibus vel pulverulentis et nigro- 
punctatis, subtus dense vel sparse brunneo-tomentosis; in- 
florescentiis axillaribus abbreviatis subcapitatis pauci- 
floris. 

Small tree with corky bark; branches obtusely tetragonal, 


386 PHYTOLOGIA No. 1l 


grey, somewhat flattened and ampliate at the nodes, with 
small white pith, pulverulent or glabrescent; branchlets 
more slender, obtusely or subacutely tetragonal, densely 
furfuraceous-ferinaceous or tomentose with yellowish or 
brown pubescence; leef=-scars rather large and corky; nodes 
not annulate; principal internodes 1.2--35 cm. long; leaves 
decussate-opposite; petioles very much abbreviated or subob- 
solete, 1--4 mm. long, stout, margined, densely furfuraceous 
pubescent or tomentose with brown hairs, flat above; blades 
firmly chartaceous, uniformly dark-green on both surfaces, 
shiny above, broadly elliptic or oblanceolate, varying to o- 
vate, 4.5--13 om. long, 2.4--6.8 cm. wide, acute or very 
shortly acuminate at apex, entire, varying from long-cuneate 
to shortly and very abruptly acuminate at base, varying from 
appressed-pubescent with short yellowish strigose hairs a- 
bove when very immature to merely pulverulent and black- 
punctate when mature, sparsely or densely brownish-tomentose 
beneath; midrib stoutish, sharply prominulous above, round- 
ed-prominent beneath; secondaries very slender, 6--13 per 
side, arcuate-ascending, prominulous on both surfaces, not 
plainly joined at the margins; veinlet reticulation coarse, 
the larger portions prominulous on both surfaces; inflores- 
cence axillary; cymes abbreviated, subcapitate, 1--2 cm. 
long, 8--15 mm. wide, several-flowered, rather dense, solit- 
ary in the upper axils on young branchlets, opposite; pedun- 
cles very slender or subfiliform, to 1 cm. long, sometimes 
subobsolete, furfuraceous; pedicels filiform, 1--4 mm. long; 
calyx cempanulate, about 3.4 mm. long, about 0.9 mm. wide to 
apex of the ovary, ampliate to 3.1 mn. at the rim, densely 
furfuraceous~puberulent outside, its rim truncate, minutely 
4-apiculate; corolla small, its tube narrow-cylindric, about 
5 mm. long, about 0.7 mm. wide at base and ampliate to 1.7 
mm. at apex, its limb not seen; stamens 4, inserted about 
1.4 mm. below the mouth of the corolla-tube; filaments fili- 
form, glabrous; style capillary, glabrous; ovary tetragonal, 
about 0.7 mm. long and wide, 4~lobed and umbilicate at apex, 
glabrous; fruiting-calyx greatly enlarged and indurated, 4-- 
5 mm. long, 6--8 mm. wide, pulverulent-puberulent, its rim 
truncate, irregularly and shallowly incised. 

The type of this species was collected by Adolfo Ducke 
(Herb. Rio de Janeiro 18,951] on a high campo, Campos do 
Jutahy, between Almeirim and Prainha, Pardé, Brazil, in flow- 
er in September, 1923, and in fruit on April 13, 1923, and 
is deposited ‘in the Britton Herbarium at the New York Botan- 
ical Garden. 

Citations: BRAZIL: Parad: Ducke sen. [Herb. Rio de Janeiro 
18,951] (N--type). 


1940 Moldenke, Notes on Aegiphila 387 


Standley in Field Mus. Publ. Bot. 18: 995 (1938) adds the 
following items to the composite description of the species: 
Shrub; branches glabrous; leaves opposite, their blades oft- 
en somewhat falcate, glabrous on both surfaces; cymes axill- 
ary and forming large, terminal, thyrsoid panicles; pedicels 
53 mm-e long or less; calyx acutish at base, puberulent; cor- 
Olla pale-yellow, its lobes 6 mm. long. He records it as in- 
habiting the "Atlantic tierra caliente". The species has 
been collected at altitudes of 1100--2050 feet. The author- 
ity name is often abbreviated "J. D. Gn." 

The H. Pittier 8643 and the Tonduz 8627, 9292, 9293, and 

b cited by me in Brittonia 1: 364 and in Phytologia 1: 
Ae (1937) as from Boeas del Toro, Panama, were actually 
collected in Cartago, Costa Rica, according to a letter re- 
ceived by me from Dr. Pittier on February 28, 1939. Dr. Pit- 
tier states definitely that the new boundary between the two 
countries did not change the status of the localities in 
which he and Tonduz collected. All these localitie are still 
in Costa Rica. He states also that the numbers of these col- 
lections and of the ones cited as "H. Pittier 11,244" and 
"13,216" are actually Herb. Instit. TPhysiso=georrs Bi Nat. Cos= 
taric. numbers and should be so cited. They are not collect- 
ors’ numbers. 
Additional citations: GUATEMALA: Retalhuleu: J. D. Smith 


1479 (MMi--photo). 


15. AEGIPHILA FASCICULATA Donn. Sm. 
This binomial is sometimes erroneously accredited to H.B. 
Kes 


31. AEGIPHILA FERRUGINEA Hayek & Spruce. 

Collectors have furnished the following additional items 
for the composite description of the species: a wide-spread- 
ing tree or tree-like shrub, 3--6 m. tall, "said to grow to 
a large tree, with wood excellent for cabinet work" Qu exia); 
leef-blades dark dullish-green above, greenish-brown tomen- 
tose beneath; calyx light-green; corolla white within, 
greenish-white outside; fruit green and hard. It has been 
collected at altitudes of 2700--4300 m., in flower and fruit 
in June, sometimes found along roadsides. A vernacular name 
is "valso". 

Additional citations: ECUADOR: Carchi: Mexia 7446 (Ar); 
Pichincha: Firmin 632 (F, Mi--photo); Penland & Summers 939 
(N); Spruce 5473 (lu--isotype); Chimborazo: Rimbaoh 616 
(Mus. Yale School of Forestry 31,994] (N). 


63. AEGIPHILA FILIPES Mart. & Schau. 
Ducke records the following additional notes about this 
species, saying it is a small tree, with aromatic leaves and 


388 PHYTOLOGIA No. 11 


white flowers, blooming in December. 

Additional citations: =a Amazonas: Krukoff 8041 
(Kr), 8042 (Kr); Spruce 1761 (lu); Paré: Ducke sen. [Herb. 
Rio de Janeiro 22,550] (N). 


28. AEGIPHILA FLUMINENSIS Vell. 
Ducke describes the species as a "scandent shrub". 
Additional citations: BRAZIL: Rio de Janeiro: Ducke sen. 
[Herb. Rio de Janeiro 601] (N); State undetermined: Wied- _ 
Neuwied s.n. [Brasilia] (lu). 


68. AEGIFHILA GLABRATA Moldenke. 
Additional citations: PERU: Junin: Killip & Smith 25,503 
(Mi--photo of type). 


62. AEGIPHILA GLANDULIFERA Moldenke. 

Standley in Field Mus. Publ. Bot. 18: 996 (1938) adds the 
following items to the composite description of this species: 
inflorescence-branches glabrous or puberulent; calyx almost 
2 mm. long; corolla yellow or pale-yellow, its tube almost 8 
mme long, its lobes 4.5 mm. long. It has been collected in 
anthesis in September. 

Additional citations: PANAMA: Canal Zone: P. White 255 
(N). COLOMBIA: Santander Sur: Dawe 472 (I--photo of isotype, 
Mi--photo of type). 


62a. AEGIPHILA GLANDULIFERA var. PARAENSIS Moldenke. 
Kuhlmann describes this variety as a shrub, with ochrace- 
ous flowers, growing in secondary forests, blooming in Octo- 
bere 
Additional citations: BRAZIL: Acre Territory: J. G Kuhl- 
mann sen. (Herb. Rio de Janeiro 22,545] (N). 


32. AEGIPHILA GOELDIANA Huber & Moldenke. 

Seandent shrub; branchlets very stout, apparently glab- 
rous, but uniformly white pulverulent-dotted under the hand- 
lens, the outer bark very papery and soon peeling off; prin- 
cipal internodes to 7 cm. long; blades varying to suborbicu- 
lar, 6--9.5 cme long, 6--8 cms wide; inflorescence red. 

The type of this remarkable species was collected by, 
Goeldi at Peixeboi on the railroad between Belem do Para and 
Braganca on March 14, 1907. The number "8166" is actually an 
herbarium number instead of being a collector's number as 
implied by me in Brittonia 1: 523 (1934) and Phytologia 1: 
204 (1937). 

Additional citations: BRAZIL: Paré: Goeldi sen. (Herb. 
Amaz. Mus. Para. 8166; Herb. Rio de Janeiro 35,664] (N-- 
isotype). 


1940 Moldenke, Notes on Aegiphila 389 


19. AEGIPHILA GRAVEOLENS Mart. & Schau. 

Contrary to what is stated in Brittonia 1: 305 (1934), 
this species is based not on one collection (Lund 796), but 
also on several collections of Martius, Riedel, and Sellow 
-- unless the presence of only the Lund specimen in the De 
Candolle Herbarium at “eneva is to be taken as indicating 
that it is the type specimen. Unless this view is taken, the 
Martius 120 and sen. and Lund ue specimens cited in Phyto- 
logia 1: 206 (1937) and 1: 292 (1938) and probably some of 
the Riedel and Sellow specimens cited there and in Brittonia 
1: 305 (1934) are actually cotypes. 


43, AEGIPHILA GUIANENSIS Moldenke. 
Additional citations: BRITISH GUIANA: M. Re Schomburgk 
4o4, in part (Mi--photo of type). 


22. AEGIFHILA HASSLERI Briq. 

THe species is listed as “Aegiphila trientha" by Herter 
in Beith. Bot. Centralbl. 59: 275 (1939), but Latzina in 
Trab. Inst. Bot. y Farm. Buenos Aires 54: 112 (1935) lists 
it correctly. Rodriguez describes the flowers of the spec- 
ies as white. 

The "Benara umbraticola Arech." mentioned by Jorge Cheb- 
ateroff in Revist. Sudam. Bot. 5: 166, 167, & 170 (1938) and 
figured in fig. 2 on page 167, is actually Aegiphile Hass- 
leri, but does not agree with the description bt Arechavale- 
ta [Anal. Mus. Montev. 2: 280 (1899) and Arech., Fl. Urug. 
2: 108 (1903)] in many important respects, so probably is 
merely a case of misidentification. The true Banara umbrat- 
fcola is a member of the Flacourtiaceae. The Chebataroff 
specimen cited below was originally distributed as “Banara 
umbraticola" by Herter. Aegiphila Hassleri ascends to 250 m. 
in Uruguay, collected in fruit in February. 

Additional citations: PARAGUAY: Balansa 2085 (lu). ARGEN- 
TINA: Misiones: D. Rodriguez 566 (N), sene (Herb. Mus. Arg- 
ent. Cienc. Nat. 23,983] (N). URUGUAY: Chebataroff s.n. 
(Herb. Herter 1885a and 99,551] (K). 


95. AEGIPHILA HERZOGII Moldenke. 

The Steinbach 3259 (B, F, Z--photo) cited by me in Britt- 
onia 1: 466 (1934) and Phytologia 1: 199 (1937) as A. elata, 
is actually A. Herzogii. The species ascends to 450 m. and 
is described as a shrub 3--4 m. tall, with yellowish flow- 
ers, blooming in February. 

Additional citations: BOLIVIA: Santa Cruz: Herzog 1369 
(Mi--photo of isotype); Steinbach 3259 (N). 


86. AEGIPHILA HIRSUTISSIMA Moldenke. 
Killip and Garcfa state that the species is a shrub, in- 


390 PHYTOLOGIA No. 11 


habiting dense forests at altitudes of SO--100 m., with pen- 
dent fruiting-branches, a green calyx, and yellow fruit in 
February. It is listed by Pittier in his "Suplemento a las 
Plantas Usuales de Venezuela", page 54 (1939). 

Additional citations: COLOMBIA: Chocé: Killip & Garcfa 
53,563 (N--fragment & photo, W, Z--photo). 


115a. AEGIPHILA HOEHNEI Moldenke. 
The species is listed by Hoehne in his "Resenha Historica 
Secc. Bot. Inst. Biol. S. Paulo", pages 153 and 157 (1937). 


42. AEGIPHILA INTEGRIFOLIA (Jacq.) Jacks. 

Collectors describe the species as a small tree or sub- 
scandent shrub, with greenish-white buds, inhabiting second- 
growth forests, dense forests, cut-over woods, and the bord- 
ers of forests. It has been collected in anthesis also in 
July, and at altitudes of 70--100 m. in Colombia and 400-- 
650 m. in Ecuador. The vernacular name "bois de golette" is 
recorded by H. L. Gerth van Wijk in his "A Dictionary of 
Plant Names" (1911 & 1916) and the species is listed by Pit- 
tier in his "Suplemento a las Plantas Usuales de Venezuela", 
page 54 (1939) and by Martyn in his "Index of the Phanerog- 
amae in the Jenman Herbarium", page 461, mss. (1937). 

Additional citations: COLOMBIA: Chocd: Killip 35,495 (N). 
ECUADOR: Napo~Pastaza: Mexia 7173 (Ar), 7187 (Ar), 7289 
(Ar). BRAZIL: Amazonas: Ducke sen. [Herb. Rio de Janeiro 
35,663] (N); Poeppig 1615 (Mi--photo); Para: Spruce 311 
nie BOLIVIA: La Paz: M. Bang 584 (lu); Buchtien 719 (lu-- 
2). 


44, AEGIPHILA INTERMEDIA Moldenke. 

Ducke describes the species as a small tree, with white 
flowers, blooming in January. 

Additional citations: BRAZIL: Amazonas: Ducke 136 (N, 8). 


82. AEGIPHILA KILLIPII Moldenke. 
Additional citations: COLOMBIA: Santander Norte: Killip & 
Smith 20,870 (Mi--photo of type). 


53. AEGIPHILA LAETA H.BeKe 

Dugand describes the species as a rather large shrub, 4 
(sometimes 6) meters tall, inhabiting forests at altitudes 
of 100--200 m.; the fruits orange when unripe, bright red 
when ripe, fruiting in October. He records the vernacular 
name "San Juan de la Verdad". Haught describes the flowers 
as pale-yellow, in January. White describes his specimen as 
a plant 4 feet tall, with "green" and faintly fragrant flow- 
ers, in September. The Pennell 3689 (G, N) cited by me in 
Brittonia 1: 382 (1934) as A. martinicensis, is actually A. 


ite bess VEC eo 


1940 Moldenke, Notes on Aegiphila 391 


laeta and thus adds an Antioquia, Colombia, record for the 
species. 

Additional citations: PANAMA: Canal Zone: Gene White 175 
(N). COLOMBIA: Atlaéntico: Dugand G. 1149 (F); Qundinemarca: 
Haught 2148 (N). 


78. AEGIPHILA LAEVIS (Aubl.) Gmel. 

An additional synonym is Aegiphila sarmentosa L. C. Rich. 
ex Moldenke, Prelim. Alph. List Invalid Names 3, in syn. 
(1940). On the same page of this work the binomial "Aegiph- 
ila levis", mentioned in Phytologia 1: 229 (1937), is ace- 
redited to Vahl and to "(Aubl.) Gmel.", and the binomial 
"Aegiphila levis" is accredited to Vahl. The vernacular name 
"manprasara” is recorded for the species. 

Additional citations: FRENCH GUIANA: Aublet sen. (Mi-- 
photo of isotype). 


47. AEGIPHILA LANATA Moldenke. 
Additional citations: BRAZIL: Goyaz: Glaziou 21,917 (Mi-- 
photo of isotype). 


97. AEGIFHILA LANCEOLATA Moldenke. 

The Balansa 2094 cited by Briquet in Ann. Conserv. Jard. 
Bot. Genev. 7--8: 318 (1904) as A. cuspidata, is actually A. 
lanceolata. 


79. AEGIPHILA LAXICUPULIS Moldenke. 

The misspelling of this binomial, "Aegiphila laxicupula", 
is recorded in Moldenke, Prelim. Alph. List Invalid Names 2, 
in syn. (1940). The species has been collected at altitudes 
of 2600--3500 feet in Guatemala. It is described by Skutch 
as a small tree, with cream-colored flowers, inhabiting 
second-growth thickets. It has been confused with A. martin- 
icensis. The Choussy 12 (W) cited by me in Brittonia 1: 383 

1934) as A. martinicensis var. oligoneura, is actually A. 

lexicupulis, and therefore adds a La Paz, Salvador, record 
for this species. 

Additional citations: GUATEMALA: Quezaltenango: Skutch 
1280 (N). 


76. AEGIPHILA LAXIFLORA Benth. 
Additional citations: BRITISH GUIANA: M. R. Schomburgk 
Wl2 (Mi--photo of type). 


27. AEGIPHILA LEHMANNII Moldenke. 

Sneidern has collected this species in a primeval forest, 
at an altitude of 800 m., in El Cauca, Colombia, blooming in 
_ March. His specimen has the leaf-blades rather more densely 
_ strigillose above and more obovate in shape than the previ- 


392 PHY F 0) L:056) 5 2 No. 11 


ously cited material. 
Additional citations: COLOMBIA: El Cauca: Sneidern 1660 


(s). 


33. AEGIPHILA LHOTZKIANA Cham. 

Additional citations: BRAZIL: Cearaé: Freire Allemao s.n. 
(Herb. Rio de Janeiro 31,757] (N); Minas Geraes: Regnell 
I.310xa (Br--photo); Parané: Dusen 1028a (I--photo, Mi-- 
photo), sen. [Herb. Rio de Janeiro 31,756] CN) 


93. AEGIPHILA LONGIPETIOLATA Moldenke. 
Additional citations: PERU: Loreto: Weberbauer 4667 (I-- 
photo of type, Mi--photo of type). 


Ducke describes the species as scandent,’ with white odor- 
ous flowers, inhabiting ground that is not inundated period- 
ically (terra firma). Sandwith calls it a bush-rope of the 
green-heart forest, with creamy-white corollas, blooming and 
fruiting in August. The first fruits to be collected of this 
species are found on the sheet of Sandwith 1192 in the Kew 
herbarium. In Moldenke, Prelim. Alph. List Invalid Nemes 19, 
in gyn (1940), the binomial "Clerodendron capitatum 
Klotzsch" is recorded as a synonym. The number "22,549" cit- 
ed as a collection number of Ducke in Brittonis 1: 320 (1934 
and Phytologia 1: 234 (1937), is actually an Herb. Rio de 
Jeneiro number and not a collector's number. 


129. AEGIPHILA MACROPHYLLA H.B.K. 
This binomial was actually first published in 1818, hot 
in "1817" as cited by me in Brittonia 1: 470 (1934). 


73- AEGIFPHILA MAGNIFICA Moldenke. 

Standley in Field Mus. Publ. Bot. 18: 996 (1938) and on 
labels gives some additional characters for the composite 
description of this species: a shrub to 10 feet tall, some- 
times scandent; branches densely puberulent; leaves short- 
petiolate; cymes axillary and terminal, forming a thyrsoid- 
pyramidal terminal panicle, whose branches are puberulent; 
fruit somewhat tetragonal, orange; inhabiting damp thick- 
ets, rare at altitudes of 330--600 m., fruiting in January. 
The label on one of the Pomona specizens of C. F. Baker 204 
indicates that Baker's number 658 is the same collection, 
or, at least, was recognized by the collector as being the 
very same species. 

Additional citations: GUATEMALA: Suchitepéquez: P. C. 
Stendley 62,196 (N). NICARAGUA: Chinandega: C. F. Baker 204 

Po--2 isotypes). 


1940 Moldenke, Notes on Aegiphila 593 


71. AEGIPHILA MARTINICENSIS Jacq. 

H. L. Gerth von Wijk in his "A Dictionary of Plant Names" 
(1911 & 1916) records the common names "bois de bouc", "bois 
cabril", "bois de cabril", "bois de fer", and "westindisches 
Eisenholz". The additional synonym, Aegiphila pyremidata L., 
is recorded in Moldenke, Prelim. Alph. List Invalid Names 3, 
in syn. (1940). Pittier records the species in his "Suple- 
mento a las Plantas Usuales de Venezuela", page 54 (1939). 
The Pennell 3689 (G, N) cited by me in Brittonia 1: 382 
(1934) proves actually to be A. laeta. The Otero 199 and 308 
recorded from the Krukoff herbarium in Phytologia 1: 293 
(1938) have now been officially transferred to the Britton 
Herbarium. The binomial, Aegiphila macrophylla H.B.Ke, cit- 
ed in Brittonia 1: 377 (19 as having been p@blished in 
"¥817" actually was not published until 1818. 

Standley in Field Mus. Publ. Bot. 18: 996 (1938) gives a 
few added characters for the composite description of the 
species: shrub or small tree; leaves short-petiolate; blades 
thin, mostly 10--20 cm. long, acuminate or narrowly long- 
acuminate at apex, glabrous or sparsely and inconspicuously 
puberulent; cymes rather lax, forming a thyrsoid terminal 
panicle, whose branches are puberulent; corolla-lobes 3--6 
mme long. He records it from the Changuinola Valley in Pana- 
ma and states that it grows "doubtless elsewhere in the At- 
lantic tierra caliente". Box in an as yet unpublished Flora 
of Antigua describes it as an “undershrub in wooded valleys 
and ravines in the S.W. district. Infrequent and local." He 
cites Box 833 and 834, not yet seen by me. Holdridge des- 
cribes it as a sparingly branched shrub 8 feet tall, with 
orange fruit in February. The specific name is sometimes 
written with a capital initial letter. It has been collected 
in anthesis in November. 

Additional citations: PORTO RICO: Holdridge 3 (N); Otero 
270 (N), 696 (N). TRINIDAD: Sieber, Fl. Trinit. 85 (Iu). 
LOCALITY OF COLLECTION UNDESIGNATED: Swartz sen. (lu). 


7la. AEGIPHILA MARTINICENSIS var. OLIGONEURA (Urb.) Moldenke. 

The Choussy 12 (W) cited by me in Brittonia 1: 383 (1934) 
as this variety proves actually to be A. laxicupulis. This, 
then, removes the only supposed record of A. martinicensis 
vare oligoneura from Salvador. 


38. AEGIPHILA MEDITERRANEA Vell. 
Additional citations: BRAZIL: Rio de Janeiro: Ule sen. 


(Herb. Rio de Janeiro 31,558] (N). 


12. AEGIPHILA MEDULLOSA Moldenke. 
Additional citations: BRAZIL: Rio de Janeiro: Saint- 
Hilaire C.50 (Mi--photo of type). 


394 PHYTOLOGIA No. 11 


83. AEGIPHILA MOLLIS H.B.K. 

Davidson describes the species as an "herb", doubtless an 
error in observation. The flowers are said to be greenish- 
yellow and fragrant, soon falling, blooming in April and 
May, fruiting in January. It has been collected in clumps on 
open "llanos" and is said to be abundant on limestone soil 
and on savannas. Haught describes the leaves as "soft", 
probably meaning that they are soft-pubescent. The species 
is recorded by Pittier in his "Suplemento a las Plantas Usu- 
ales de Venezuela", page 54 (1939). It is said to be a shrub 
1--5 m. tall by most collectors, growing at altitudes of 100 
--1250 m. The Hartweg 13559* cited below is a different col- 
lection from 4artweg 1559. The latter is Petrea rugosa 
H.B.K. ’ 
Additional citations: PANAMA: Chiriqu{: Devidson 638 (F). 
COLOMBIA: Magdalena: Haught 2288 (N); El Valle: Garcia 
Barriga 6449a (W); Cundinamarca: Hartweg 1359* (Iu); Meta: 
Cuatrecasas 43539 (Jce--2); El Vichada: Haught 27935 (N). VEN- 
EZUELA: Sucre: Funck 643 (Iu). BOLIVIA: Santa Cruz: Stein- 
bach 5168 (N). p 


13. AEGIFHILA MONSTROSA Moldenke. 

Standley gives numerous additional characters for the 
composite description of this species: shrub or tree; 
branches pale, obtusely 4~angled, exceedingly brittle and 
easily broken from the plant with little effort; wood pink- 
ish-gray, very light and soft, medium-textured; leaves very 
large, long-petiolate, the blades usually broadly elliptic, 
acute or acuminate at Apex, narrowed to the base, nearly 
glabrous; flowers small, pure white, in small clusters in 
the axils of the leaves and below the leaves at the nodes of 
the naked branches, rather numerous and handsome, suggesting 
those of coffee at a short distance; fruit globose, about 8 
mm. in diameter. Schipp describes it as a tree to 10 m. 
tall, with a stem-diameter of 8 inches and sweetly perfumed 
flowers. Lundell refers to it as a "bush" in Guatemala. It 
has been collected in pine regions of humid areas in Hondur- 
as, in anthesis in May, August, November, and December, and 
in fruit in February and April. It has been collected at al- 
titudes of from 50 to 4500 feet. It is recorded in Fedde, 
Bot. Jehresber. 59™: 416 (1939) and by lundell in Carnegie 
Inst. Wash. Publ. 478: 47 & 75 (1937). 

Additional citations: GUATEMALA: Alta Verapaz: H. V. 
Johnson 520 (La). HONDURAS: Yoro: Von Hagen & Von Hagen 1020 
(N); Atlantida: P. ©. Standley 53,176 (I--photo); Cortés: 
Carleton 422 (Mi--photo of type). BRITISH HONDURAS: Gentle 


948 (La); Schipp 1083 (F). 
32a. AEGIPHILA MONTICOLA Moldenke. 


—— a ee 


—_-. ee eee. =— 


1940 Moldenke, Notes on Aegiphila 3595 


The Schimpff 252 (B--2, Ob, N--fragment) cited as A. bog= 
otensis in Phytologia 1: 188 (1937) is actuelly A. monticola 
instead. 

Additional citations! ECUADOR: Chimborazo: Rimbach 118 
(Mus. Yale School of Forestry 22,820] (Mi--isotype). $52 


11. AEGIFHILA MULTIFLORA Rufz & Pav. 

Steinbach states that the entire inflorescence is violet- 
red, the filaments white, and the anthers gray or cinnamon- 
colored. 
atin citations: BOLIVIA: Cochabamba: Steinbach 5809 

F, N). 


119. AEGIPHILA OBDUCTA Vell. 

Additional synonyms are Aegiphila fontex Schwacke ex Mol- 
denke, Prelim. Alph. List Invalid Nemes 2, in syn. (1940) 
and Aegiphila salicifolia Sellow ex Moldenke, Prelim. Alph. 
List Invalid Names 3, in syne (1940). Ducke describes the 
species as a small tree, with white leaves (meaning leaves 
white beneath?), blooming in June. Riedel 452 was mididenti- 
fied and originally distributed as a species of Buddleia. 

Additional citations: BRAZIL: Minas Geraes: Riedel 452 
(W); Rio de Janeiro: Ducke sen. [Herbe Rio de Janeiro 
22,551] (N); Herb. Rio de Janeiro 31,720 (N); Parana: Jonss- 
on 379a (lu); Santa Catharina: Schwacke sen. [Herb. Rio de 
Janeiro 31,777] (N). ode 


52. AEGIPHILA ODONTOPHYLLA Donn. Sn. 

Standley in Field Mus. Publ. Bot. 18: 996--997 (1938) 
adds a number of characters to the composite description of 
this species: small tree, to 10 m. tall; trunk to 30 cm. in 
diameter; bark grey or grayish-brown, roughened; leaves 
short-petiolate, opposite, the blades elliptic-oblong, more 
or less granulose-tomentose; cymes all axillary, dense, 
densely tomentose; corolla creamy-white; fruit pale-yellow 
when ripe. He states that the type was collected on the 
southern (not northern) slope of Volcan de Barba and that 
the species is endemic to the wet forests of the central 
mountains at altitudes of 2000--2400 m. 


69. AEGIPHILA PANAMENSIS Moldenke. 

Standley in Field Mus. Publ. Bot. 19: 997 (1938) and var- 
ious collectors have added the following items to the comp~ 
osite description of this species: slender shrub or small 
tree to 8 m. tall, or scandent; branches densely puberulent; 
leaves short-petiolate, the blades thin, blackening in dry- 
ing, caudate-acuminate at apex, rather densely puberulent on 
both surfaces; cymes small, dense or lax, forming a terminal 
thyrsoid panicle, whose branches are densely puberulent; ca- 


396 PHYTOLOGIA No. 11 


lyx almost 3 mm. long; corolla pale-yellowish, cream-colored 
or white, its tube 5--7.5 mm. long, its lobes 4 mn. long; 
fruit subglobose, 1 cm. long, truncate at apex. The species 
has been collected at an altitude of 40 m. by Allen and in 
anthesis in October. It is said by Standley to inhabit the 
"thickets of the Atlantic tierra caliente....at 1,100 meters 
or lesse" The Skutch 2680 and Matuda 2115 cited below may 
possibly be A. paniculata instead of the present species, 
since without the fruit the two species are difficult to 
distinguish. A. panamensis is recorded in Fedde, Bot. Jahr- 
esber. 59%: 416 (1959). 

Additional citations: MEXICO: Chiapas: Matuda 2115 (Mi, 
N). COSTA RICA: San José: Skutch 2680 (Mi). PANAMA: Coclé: 
Woodson, Allen, & Seibert 1756 (N); Canal Zone: P. H. Allen 
2016 (N); Darien: P. H. Allen 906 (N). 


74. AEGIPHILA PANICULATA Moldenke. 

Standley in Field Mus. Publ. Bot. 18: 997 (1938) adds the 
following supplementary characters to the composite descrip- 
tion of the species: shrub or tree; branches glabrous or 
puberulent; leaves short-petiolate, the blades thin, brun- 
nescent in drying, acuminate or acute at base; inflorescen- 
ce-branches glabrous or puberulent; fruit truncate at apex. 
“eo says that it inhabits the Changuinola Valley "and doubt- 
less elsewhere in the Atlantic tierra caliente". He remarks 
"The species is close to A. panamensis and perhaps should be 
united with it." The Skutch 2680 and Matuda 2115 cited above 
under A. panamensis may possibly represent A. paniculata. 
ie “bags is recorded by Fedde in Bot. Jahresber. 592%: 416 

1939)-« 


34. AEGIPHILA PARAGUARIENSIS Briq. 

The Malme 2473 (S--2, W) cited by me as A. Sellowiana in 
Brittonia 1: Ss! (1934) and in Phytologia 1: 266 (1937) is 
actually A. paraguariensis. 

Additional citations: BRAZIL: Mattogrosso: Malme 2473 
(Lu); Parané: Dusén 10,472 (Iu). 


72. AEGIFHILA PENDULA Moldenke. 
This species is recorded by Pittier in his "Suplemento a 
las Plantas Usuales de Venezuela", page 54 (1939). 


35a. AEGIPHILA PERNAMBUCENSIS Moldenke. 

The species has been collected in anthesis in November. 
In Phytologia 1: 258 (1937) four isotypes are cited from the 
Britton Herbarium. This is an error -- there are only two i- 
sotypes there. 

Additional citations: BRAZIL: Parahyba: Zenaide 31 [Herb. 
Inst. Biol. S. Paulo 36,703] (Sp). 


: 


1940 Moldenke, Notes on Aegiphila 397 


75. AEGIPHILA PERPLEXA Moldenke. 

The species is described by Sandwith as having dirty yel- 
lowish-white flowers, blooming in October. 

Additional citations: TOBAGO: Sandwith 1801 (N). 


64, AEGIPHILA PERUVIANA Turcz. 
Additional citations: PERU: San Martin: Spruce 4275 (Lu-- 
isotype); Ll. Williams 6836 (Mi--photo). 


88. AEGIPHILA PUBERULENTA Moldenke. 
Additional citations: COLOMBIA: Bolfvar: Elias 617 (Mi-- 
photo of type). 


91. AEGIPHILA QUINDUENSIS (H.B.K.) Moldenke. 

Williams and Delgado add the following characters to the 
composite description of this species: slender tree,.to 11 
me tall; crown spreading or the branches erect or upright; 
trunk round, slightly bent, to 30 cm. in diameter, unbranch- 
ed for 4 or 5m. from the base; inhabiting transition for- 
ests, blooming in January. The species is listed by Pittier 
in his "Suplemento a las Plantas Usuales de Venezuela", page 
54 (1939). 

Additional citations: VENEZUELA: Carabobo: Karsten sen. 
[Puerto Cabello] (Mi--photo); Aragua: Delgado 115 (W); Li. 
Williams 10,251 (W). 


115. AEGIFPHILA RACEMOSA Vell. 

The species is described by Haught as a shrub or small 
tree, 2--4 m. tall, with light-yellow flowers. It has been 
collected at 250 m. altitude, in anthesis in February and 
October, and is listed by Martyn in his "Index to the Fhan- 
erogamae of the Jenman Herbarium", page 461, mss. (1937). 

Additionel citations: COLOMBIA: Boyacé: Haught 2612 (N). 
BRITISH GUIANA: De la Cruz 3627 (Mi--photo). BRAZIL: Para: 
Guedes sen. (Herb. Rio de Janeiro 31,581] (N). 


24, AEGIPHILA RIEDELIANA Schau. ; 
Additional citations: BRAZIL: Bahia: Pohl 4392 (I--photo 

of cotype); Santa Catharina: Ule 1537 (Br--photo, Mi-- 

photo). ILLUSTRATIONS: Pohl, Icon. Plant. Brasil. 333 (V). 


14a. AEGIPHILA RIMBACHII Moldenke. 
Additional citations: ECUADOR: Los Rios: Rimbach 234 
(Mus. Yale School of Forestry 28,648] (F--isotype). 


Ae, AEGIPHILA SALTICOLA Moldenke. 
Small tree; principal internodes 1.5--5.5 cm. long; bla- 

des 3.8--7 cm. wide, very sparsely scattered-pilose or gla- 

brescent above; cymes to 7 cm. long and 5 cme wide, very 


398 PHYTOLOGIA No. 11 


densely subvillous with flavescent hairs throughout; pedunc- 
les 2--4 cm. long; pedicels filiform, 1--2 mm. long, densely 
subvillous-pubescent; calyx elongate-campanulate, thin and 
translucent, 5.8--6 mm. long, about 3.4 mm. wide at apex, l 
mm, wide toward base, densely villous with appressed or sub- 
spreading flavescent hairs, its rim 4-toothed, the teeth 
triangular-ovate, about 1.5 mm. long and 1.7 mm. wide at 
-base, acute; corolla white, hypocrateriform, its tube nar- 
row-cylindric, about 5 mm. long, glabrous, its limb 4~-part- 
ed, the lobes elliptic-lingulate, venose, about 3 mm. long 
and 1.7 mm. wide, subacute at apex, glabrous on both surf- 
aces; stamens 4, inserted about 1.2 mm. below the mouth of 
the corolla-tube, long-exserted; filaments filiform, 8.8--9 
mm. long, flattened, glabrous; anthers oblong, about 1.2 mm. 
long and 0.5 mm. wide, dorsifixed near the base; pistil in- 
cluded or short-exserted; style capillary, about 5.1 mm. 
long, glabrous; stigma bifid, its branches about 1 mm. long. 

Additional citations: BRAZIL: Amazonas: Ducke sen. [Herb. 
Rio de Janeiro 25,593] (N). 


39. AEGIPHILA SELLOWIANA Cham. 

The Malme 2473 (S--2, W) cited by me in Brittonia 1: 334 
(1934) and in Phytologia 1: 266 (1937) is actually A. parag- 
uariensis. The Mexia 4500 cited in Phytologia 1: 294 (1938 
as from the Krukoff Herbarium has since been transferred to 
the Britton Herbarium. The trinomial, Aeziphila tomentosa 
var. silvestris Regnell, cited as a synonym of A. verticill- 
ata in Brittonia 1: 329 (1934), is actually synonymous with 
A. Sellowiana [see Moldenke, Prelim. Alph. List Invalid 
Names 4. 1940]. Another new synonym is Aegiphila integerrima 
(Jacq. ) Jacks. ex Moldenke, Prelim. Alph. List Invalid Names 
2, in syn. (1940). The two Herb. Rio de Janeiro specimens 
cited below show beautifully the remarkable difference in 
the size of the male- and the female-predominant flowers of 
this species -- no. 31,585 is the female type and has very 
small flowers, while no. 31,521 is the male type and has 
very large flowers. 

Additional citations: BRAZIL: Minas Geraes: Regnell 1.184 
[1845] (Lu); Rio de Janeiro: Freire sene [Herb. Rio de Jan- 
eiro 31,521] (N); Sdo Paulo: R. D. Gongalves een. [Herb. 
Inst. Biol. S. Paulo 39,285] (Sp); Herb. Rio de Janeiro 31,- 
585 (N); Parané: Dusén 15,855 (Iu); Rio Grande do Sul: Malme 


799 (Iu). 


16. AEGIPHILA SESSILIFLORA Moldenke. 
Additional citations: COLOMBIA: Antioquia: Archer 392 (Mi 
--photo of type). 


104, AEGIPHILA SSTIFORMIS Rusby. 


1940 Moldenke, Notes on Aegiphila 399 


Additional citations: BOLIVIA: La Paz: M. Bang 878a, in 
part (I--photo of type), 1732 (Mi--photo). _ 


27b. AEGIPHILA SKUTCHII Moldenke, Geogr. Distrib. 13 & 15, 
nome nud. (1939), spe nov. 

Arbor; foliis oppositis; laminis anguste ellipticis acum- 
inatis integris, ad basim acutis vel subacuminatis, supra 
minutissime puberulis, subtus subglabratis densissime gland- 
ulosis; inflorescentiis axillaribus; cymis pedunculatis; 
calyce campanulato truncato; corollae lobis 4 obovatis. 

Tree, to 16 m. tall; trunk to 47 om. in diameter; 
branchlets rather slender, obtusely tetragonal, gray, minu- 
tely but not densely appressed-puberulent, glabrescent in 
age; principal internodes 1--2./7 cm. long; leaves decussate- 
opposite, numerous; petioles slender, 2--3 cm. long, dense- 
ly and minutely appressed-puberulent, canaliculate above; 
leaf-blades thin-chartaceous, rather uniformly dark-green on 
both surfaces, narrowly elliptic, 8--16 om. long, 1.5--4.8 
cm. wide, acuminate at apex, entire, acute or subacuminate 
at base, very minutely and sparsely puberulent above, more 
densely so along the midrib and secondaries, subglabrate be- 
neath but very densely glandulose with small black puncti- 
form and discoid glands; midrib slender, subprominulent and 
often lighter-colored above, prominent and often lighter- 
colored beneath; secondaries slender, 7--9 per side, rather 
irregular and with numerous tertiaries intervening between 
them, arcuate-ascending, flat or subprominulous above, prom 
inulous beneath, often lighter in color than the lamina on 
both surfaces; veinlet reticulation rather sparse and ob- 
scure on both surfaces; inflorescence axillary, abundant to- 
ward the apex of the branchlets; cymes 4--8 cm. long, 2.5--4 
ems wide, brachiate, many-flowered; peduncles rather slend- 
er, 2--4.5 cm. long, divaricate-ascending, densely appress- 
ed-puberulent; pedicels very slender, 1--1.5 mm. long, minu- 
tely puberulent; bractlets and prophylla linear, 1--2.5 mm. 
long, cinereous-puberulent; calyx campanulate, about 3 mM « 
long and wide, very sparsely and minutely puberulent, its 
rim truncate and entire or slightly repand; corolla hypo- 
crateriform, its tube cylindric, about 5 mm. long, slightly 
ampliate above, its limb 4-parted, the lobes obovate-lingul- 
ate, about 5 mm. long, 3--4 mm. wide, rounded at apex, glab- 
rous on both surfaces; stamens 4, inserted about 1.7 mm. be- 
low the mouth of the corolla-tube, long-exserted; filaments 
ita about 10 mm. long, glabrous; anthers oblong, about 

1.7 mm. long and 0.7 mm. wide; pistil included; style capil- 
lary, about 3.7 mm. long, glabrous; stigma bifid, its bran- 
ches about 2 mm. long; immature fruiting-calyx accrescent, 
very minutely puberulent or glabrescent, its rim irregularly 
split e 


400 PHYTOLOGIA No. 1l 


The type of this very distinctive species was collected 
by Alexander Frank Skutch (no. 1551) -- in whose honor it is 
named -- at Finca Moca, altitude 4700 feet, Suchitepéquez, 
Guatemala, on October 29, 1934, and is deposited in the 
Britton Herbarium st the New York Botanical Garden. The col- 
lector notes that the type tree was 50 feet tall, with a 
trunk-diameter of 20 inches at breast height. Immature fruit 
has been collected in August. 

Citations: MEXICO: Chiapas: Matuda 1653 (Mi--2, N--2, N-- 
fragment). GUATEMALA: Suchitepéquez: Skutch 1551 (N--type). 


94. AEGIFPHILA SMITHII Moldenke. 
Additional citations: PERU: Loreto: Klug 1460 (I--photo, 
Mi--photo ). 


92. AEGIPHILA SPRUCEANA Moldenke. 
Additional citations: BRAZIL: Amazonas: Spruce 2296 (Lu-- 
isotype, N--isotype). 


26. AEGIFHILA STANDLEYI Moldenke. 

Standley in Field Mus. Publ. Bot. 18: 997 (1938) gives 
the following additional characters for the composite des- 
cription of this species: branches glabrate; leaves oppos- 
ite, short-petiolate, the blades attenuate to the base; 
cymes all axillarys, equaling or shorter than the petioles, 
short-pedunculate; calyx 3.5--4 mm. long. 


84, AEGIFHILA SURFACEANA Moldenke. 

Kuhlmann describes this species as subscandent, with 
ochraceous-green flowers, inhabiting the secondary forest, 
blooming in April. It is recorded by Fedde in Bot. Jahres- 
ber. 592: 416 (1939). 

Additional citations: BRAZIL: Para: Dehlgren & Sella 173 
(Mi--photo); Herb. Amaz. Mus. Para. 10,751 Herb. Rio de 
Janeiro 35,660] (N); J. G. Kuhlmann sen. [Herb. Rio de Jan- 
eiro 22,548] (N). 


55. AEGIPHILA SWARTZIANA Urb. 
Additional citations: JAMAICA: Swartz sen. [Jemaica] (I-- 
photo of type). 


1. AEGIPHILA TERNIFOLIA (H.B.K.) Moldenke. 
Additional citations: COLOMBIA: Qindinamarca: Goudot sene 
(Near Bogota] (Mi--photo). 


56. AEGIPHILA TRIFIDA Sw. 

Harris describes the species as a straggling shrub 8 feet 
tall. : 

Additional citations: JAMAICA: W. Harris 6273 (Mi--photo) 


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 ~PHYTOLOGIA 


Designed to expedite botanical publication 


Vor. 1 October, 1940 No. 12 
mY est 
wi 
Fi 


Bic | CONTENTS 


‘ 
7 
PB 


ui, Published iy H. A. Gleissn and Harold N. Moldenke 
The New York Botanical Garden _ 
Bronx Park, New York, N. Y. 


a. Wr eee 


as 


NEW SPECIES OF CROTON FROM THE YUCATAN PENINSULA (a) 


Ooi tes Lundell 


' CROTON AGUILARI Lundell, sp. nov. 


Frutex, stellato-tomentosa. Petioli usque ad 7.5 om. lon- 


- gi, apice glandulosi. Lamina chartacea, denticulata, late o- 


vata, apice caudato-acuminata, basi cordata, supra subglab- 
ra, subtus albido-tomentosa. Inflorescentiae bisexuales. 
Flores 9 sessiles; calyce usque ad 3.5 mm. longo, lobis tri- 
angulari-acuminatis; stylis ca. 6 mm. longis. Flores & pedi- 
cellati, glomerulati; calyce ca. 53.5 mm. longo; staminibus 
36 vel 37. Capsula parce stellato-tomentosa, cae 7 mm. long. 

A shrub with whitish to tawny tomentum of stellate hairs. 
Branchlets thick, with large pith, sulcate, covered at first 
with tawny tomentum, at length glabrescent. Petioles stout, 
tawny tomentose, up to 7.5 om. long, with 4 conspicuous 
glands at apex on under surface. Leaf-blades thin-chartace- 
ous, irregularly denticulate or subserrulate, the teeth 
gland-tipped, broadly ovate, up to 25 om. long, 15.5 cm. 
wide, apex caudate-scuminate, base cordate, covered beneath 
with whitish tomentum of stellate hairs, sparsely stellate 
hairy above except along the veins, costa and primary veins 
elevated beneath, slightly impressed above, base 3~ or 5- 
nerved. Racemes terminal, bisexual, up to 38 ome long, the 
rachis and flowers tawny tomentose, the staminate flowers 
glomerulate. Pistillate flowers sessile; calyx up to 3.5 mn. 
long, deeply S-lobed, the lobes strongly unequal, triangu- 
lar-acuminate, subsetaceous, 1 to 43 mm. long; petals reduced 
to leciniae; ovary tammy tomentose with stellate hairs; 
styles sparsely hairy, about 6 mm. long, each once branched. 
Staminate flowers with slender pedicels up to 6.5 mm. long; 
calyx about 3.5 mm. long, deeply S-lobed, the lobes ovate; 
petals 5, oblong-elliptic, about 3.5 mm. long, minutely pap- 
illate, long-pilose within at base; stamens 36 or 37, fila- 
ments glabrous; receptacle pilose. Capsules sessile, about 7 
mme long, sparsely tawny tomentose. 

Type in the University of Michigan Herbarium, Mercedes 


Aguilar H. 463, collected in secondary forest (acahual), 


near La Libertad, Department of Petén, Guatemala, January 
30, 1935; vernacular name "caret". 

Mr. Aguilar reports that it has a bitter sap which flows 
copiously. The species is allied to C. asteroides Lundell, 
and belongs to the complex of C. xalapensis H.B.K. 


CROTON AMELIAE Lundell, sp. nov. 
Frutex, ramulis sail" hata Petioli usque ad 3 
1 


402 ba: Bane sky a ae Pha (ig te oe No. 12 


cme longi. Lamine membranacea, integra, ovata, apice acumin- 
ata, basi anguste cordata, subtus albido-tomentosa. Inflor- 
escentiae bisexuales. Floresg pedicellati; calyce usque ad 
11 mm. longo, lobis 5, oblong-lanceolatis, stipitato-glandu- 
losis. Flores & pedicellati; calyce 2.6 mm. longo; petelis 
5, oblongis; staminibus 16. 

Arborescent shrub, about 2 m. high, with whitish tomentum 
of stellate hairs. Branchlets erect, slender, white toment- 
ellous and sparsely hirsute with stellate hairs. Stipules 
eristate, up to 1.3 ome long. Fetioles slender, white tomen- 
tellous, up to 3 cm. long. Leaf-blades membranaceous, en- 
tire, ovate, 3 to 6.8 om. long, 1.5 to 3.7 cm. wide, apex a- 
cuminate, base cordate with a closed sinus, stellate puberu- 
lent and green above, persistently white tomentellous be-_ 
neath, costa elevated beneath, slightly impressed ebove, 5- 
Or 7-veined at base. Racemes terminal, bisexual, up to 7.5 
ome long, white tomentellous and sparsely hirsute. Pistill- 
ate flowers with stellate-hirsute pedicels up to 7 mm. long; 
calyx up to 11 mm. long, deeply S-lobed, the lobes oblong- 
lanceolate, stellate-hirsute,-the lower two-thirds bearing 
stalked prominent glands along the margins outside, the 
lobes tomentose within; ovary stellate hairy; styles united 
at base, digitately lobed. Staminate flowers with pedicels 
up to 5 mm. long; calyx 2.6 mm. long, the lobes lanceolate; 
petals 5, oblong, about 2.3 mm. long; stamens 16, filaments 
pilose below middle; receptacle pilose. 


uous forest along Kaua road, east of Chichen Itza, Yucatan, 
Mexico, June 8, 1938. 


CROTON ASTEROIDES Lundell, sp. nov. 

Arbor, stellato-tomentosa. Petioli 1--5 cm. longi, apice 
glandulosi. Lamina chartacea, minute denticulata, ovata vel 
lanceolato-ovata, apice caudato-acuminata, basi subcordata 
vel rotundata, supra subglabra, subtus albido-tomentosa. In- 
florescentiae bisexuales. Flores ? sessiles; calyce ca. 2.8 
mm. longo, lobis 5, triangulari-acuminatis, subsetaceis; 
stylis ca. 5 mm. longis. Flores & pedicellati; calyce ca. 
3.2 mm. longo; staminibus 30 vel 31. Capsula stellato-tomen- 
tosa, ca. 6.5 mm. longa. 

A tree, 12.5 om. in diam., with whitish tomentum of stel- 
late hairs. Branchlets rather slender, somewhat flattened 
and ‘angled. Petioles slender, 1 to 5 om. long, with 2 or 4 
conspicuous sessile or subsessile yellow glands at apex on 
under surface. Leaf-blades chartaceous, minutely denticulate 
with glandular teeth, ovate or lanceolate~ovate, 5.5 to 13 
eme long, 2.8 to 665 cme wide, apex caudate-acuminate, base 
shallowly cordate or rounded, white tomentose beneath with 


1940 Iundell, New Species of Croton 403 


soft stellate hairs, green and very sparsely stellate hairy 
above except along the veins, costa and primary veins eleva- 
ted beneath, fine and with age slightly impressed on upper 
surface, base 3- or S-nerved. Racemes terminal, bisexual, up 
to 13.5 cm. long, the rachis and flowers white tomentose 
with soft stellate hairs. Fistillate flowers sessile; calyx 
about 2.8 mm. long, star-like, deeply S-lobed, the lobes 
narrowly triangular-acuminate, subsetaceous; ovary white to- 
mentellous with stellate hairs; styles sparsely hairy, about 
5S mme long, each once branched. Staminate flowers with slen- 
der pedicels up to 7 mm. long; calyx about 3.2 mn. long, 
deeply S-lobed, the lobes ovate; petals 5, oblong-elliptic 
or obovate-oblong, 4.5 mm. long, minutely papillate, long- 
pilose within at base; stamens 50 or 31, filaments glabrous; 
receptacle pilose. Capsules sessile, about 6.5 mm. long, 
white tomentose with short stellate hairs. 

Type in the University of Michigan Herbarium, Peroy H. 
Gentle 2218, collected on hillsides at Vaca, El Cayo Dist- 
rict, British Honduras, February 22, 1938. 

The calyx of the pistillate flowers is asteroid in app- 
earance, whence the specific name. 


CROTON FLAVOGLANDULOSUS Lundell, sp. nov. 

Frutex. Petioli 3--7 cm. longi. Lamina chartacea, integ- 
ra, anguste lanceolata vel lineari-lanceolata, apice attenu- 
ata, acuta vel acuminata, basi obtusa, supra stellato- 
hirtella, subtus albido-tomentosa, biglandulosa. Inflores- 
centiae bisexuales. Flores @ pedicellati; calyce 3--3.5 mm. 
longo, lobis 5, oblongis. Flores & pedicellati; calyce ca. 3 
mme longo, lobis 5; staminibus 10 vel 11. Capsula albido- 
tomentosa, cae 5 mm. longa. 

A shrub. Branchlets slender, wiry, tawny, densely stell- 
ate-hirtellous. Petioles rather stout, canaliculate, 4 to 7 
mm. long. Leaf-blades chartaceous, entire, narrowly lanceol- 
ate or linear-lanceolate, 2.3 to 5 cme long, 0.6 to 1.4 om. 
wide, apex attenuate, acute or acuminate, base obtuse, upper 
surface covered with small stellate hairs, beneath with 
whitish tomentum of appressed stellate hairs, pinnately 
veined, the veins evident beneath, scarcely discernible a- 
bove, costa slightly impressed above, prominent on lower 
surface, with a pair of stipitate yellow glands at base be- 
neath. Racemes terminal, bisexual, up to 5.5 cm. long, tom- 
entellous with whitish stellate hairs. Pistillate flowers 
with stout pedicels about 1.8 mm. long; calyx 3 to 3.5 m. 
long, deeply S-lobed, the lobes oblong, up to 3 mm. long, 
1.3 mm. wide, entire, obtuse, accrescent; petal vestiges 
threadlike; ovary stellate-tomentellous; styles hairy, each 
once branched. Staminate flowers with slender pedicels up 
to 3 mm. long; calyx about 3 mm. long, deeply S-lobed, the 


404 PHYTOLOGIA No. 12 


lobes narrowly ovate, up to 2.5 mm. long; petals 5, oblong- 
elliptic, about 3 mm. long, 1-35 mm. wide, long-pilose at 
base; stamens 10 or 11, filaments glabrous, up to 3 m. 
long, anthers 1 mm. long; receptacle pilose, Capsule white 
tomentellous with stellate hairs, about 5 m. long. 

Type in the University of Michigan Herbarium, Eizi Matuda 
3042, collected in savanna at Achotal near Balancan, Tabas- 
co, Mexico, May 9-14, 1939. 


CROTON ICCHE Lundell, sp. nov. 

Arbor parva, lepidota. Petioli 4--10 m. longi. Lamina 
chartacea, subintegra, eglandulosa, lanceolata vel oblongo- 
lanceolata, apice obtuse acuminata, basi obtusa vel rotunda- 
ta. Inflorescentiae usque ad 4.5 om. longae. Flores 9 pedi- 
cellati; calyce ca. 2.8 mm. longo, lobis 5, ovatis. Flores 
pedicellati; staminibus 9--1l. Capsule ca. 7 mm. longa, lep- 
idota, haud tuberculata. 

A small tree or arborescent shrub up to 5 m. high and 5 
cme in diam., lepidote throughout; branchlets slender, ferr- 
uginous-lepidote, angled and somewhat compressed, terete 
with age. Petioles slender, canaliculate, 4 to 10 mm. long. 
Leaf-bledes thinly chartaceous, subentire, eglandular, lanc- 
eolate or oblong-lanceolate, 3.5 to 9.5 cm. long, 1.2 to 3.8 
cm. wide, apex obtusely acuminate, base obtuse or rounded, 
often subcuneate, green and sparsely lepidote above, at 
first densely lepidote beneath, glabrescent with age, costa 
plane or slightly raised on upper surface, prominent be- 
neath, pinnetely veined, the veins fine and inconspicuous. 
Racemes axillary and terminal, staminate or bisexual, up to 
4.5 om. long, ferruginous-lepidote. Pistillate flowers with 
short stout pedicels about 2.5 mm. long; calyx about 2.8 mm. 
long, deeply 5-lobed, the lobes ovate, tomentose within; 
petals 5, oblanceolate or spatulate, clawed, about 3 mm. 
long, lepidote outside, densely short villous-tomentose 
within; overy lepidote; styles punctate, branched 2 or 3 
times. Stammate flowers with slender pedicels about 2 mm. 
long; calyx about 2.8 mm. long, deeply S-lobed; petals 5, 
elliptic, about 3 mm. long, clawed; stamens 10 or ll, fila- 
ments sparsely pilose; receptacle pilose. Capsules ferrugin- 
ous-lepidote, smooth, not tuberculate, about 7 mm. long; 
fruiting pedicels and calyx not accrescent. 

Type in the University of Michigan Herbarium, C. L. Lund- 
on llerida road between Piste and Yokdzonoot, Yucatan, Mexi- 
co, July 11, 1938. In second growth nearFisté, Lundell and 
Lundell 7547, June 14, 1938. In advanced forest east of 
Coba, Quintana Roo, Mexico, Llundell and Lundell 7719, June 
30, 1938; vernacular name "icche". 

In Maya, ic is chili end che is wood or tree. The leaves 


1940 Lundell, New Species of Croton 405 


and wood of the tree have an odor suggestive of the chili 
pepper, whence the Maya name “icche". C. Icche is a member 
of the complex which includes C. glabellus Le, C. perobtusus 
Lundell, and C. pseudoglabellus Lundell. 


CROTON ITZAEUS Lundell, sp. nov. 

Frutex, stellato-tomentosa. Fetioli usque ad 1 cm. longi. 
Lemina chartacea, serrato-dentata, lanceolato-oblonga, apice 
acuta vel obtusa, basi obtusa vel rotundata. Inflorescentiae 
bisexuales, usque ad 3.5 ome longae. Flores ¢ pedicellati; 
calyce usque ad 6 mm. longo. Flores 6 pedicellati; stamini- 
bus 11--13. 

Shrub, 1 m. high, intricately branched, stellate-tomento- 
se. Branchlets slender, at first tomentose, the stellate 
hairs with central ray elongated sharply. Petioles up to 1 
em. long. Leaf-blades chartaceous, lanceolate-oblong or ov- 
ate-lanceolate, 1.5 to 5 cme. long, 0.8 to 2.2 cm. wide, apex 
usually acute, sometimes bluntly obtuse, base obtuse or 
rounded, yellow-green above and densely covered with soft 
stalked stellate hairs, with whitish tomentum of stellate 
hairs beneath, margin irregularly serrate-dentate, usually 
with small short-stalked glands between the serratures and 
at base of blade, pinnately veined, the costa and veins con- 
spicuous beneath. Racemes terminal, bisexual, up to 3.5 cm. 
long, with whitish tomentum of stellate hairs. Pistillate 
flovers with stout pedicels about 1 mm. long; calyx up to 6 
me long, S-lobed, the lobes unequal, oblong or obovate- 
elliptic; petal vestiges minute, resembling staminodes; ov- 
ary stellate-tomentose; styles 3, eech once branched. Stam- 
inate flowers with pedicels up to 3 mm. long; calyx 2.8 mn. 
long, lobes ovate-oblong; petals 5, oblanceolate, up to 4.5 
mm. long, ciliate, the hairs below middle elongated; stamens 
11 to 13, filements glabrous; receptacle pilose. 


tan, Mexico, July 22, 1938. 


CROTON PEROBTUSUS Lundell, sp. nov. 

Arbor, lepidota. Fetioli 0.5--1.8 om. longi. Lamina sub- 
coriacea, integra, oblonga vel lanceolato-oblonga, utrinque 
obtusa. Inflorescentiae bisexuales, usque ad 5 cm. longae. 
Flores ¢ subsessiles; calyce fere ad basim partito, lobis 5, 
ovatis, 2.2--2.5 mm. longis. Flores S pedicellati; staminib- 
us 1l vel 12. Fedicelli fructiferi 2--3 mm. longi. Capsula 
haud tuberculata, lepidota, ca. 6.5 mm. longa. 

A tree up to 18 m. high and 40 om. in diame, silvery lep- 
idote throughout, Branchlets stout, angled. Petioles stout, 
0.5 to 1.8 cm. long, shallowly canaliculate, eglandular at 


406 PHYTOLOGIA Noe 12 


apexe Loer-blades subcoriaceous, entire, oblong or lanceola- 
te-oblong, 7 to 14.5 cm. long, 3.3 to 5.3 cm. wide, apex and 
base usually bluntly obtuse, apex sometimes obtusely subac- 
uminate, sparingly lepidote above at first, glabrous with 
age, densely lepidote beneath, costa slightly elevated ab- 
ove, prominent beneath, pinnately veined, the primary veins 
prominulous on both surfaces. Racemes subspicate, bisexual, 
axillary, up to 5 cm. long. Pistillate flowers subsessile, 
the pedicels less than 1 mm. long; calyx 5-lobed nearly to 
the base, the lobes ovate, 2.2 to 2.5 mm. long, entire, with 
finely lanate margins; petals 5, obovate or elliptic, up to 
53 mme long, short-clawed, margins long-ciliate-lanate; ovary 
lepidote; styles 3. Staminate flowers short-pedicellate, 
calyx and oorolla as in pistillate flowers; stamens 11 or 12, 
filaments 35 mm. long, anthers 1 mm. long; receptacle pilose. 
Fruiting pedicels stout, usually 2 to 3 mm. long, rarely up 
to 4 mm. long. Capsules smooth or nearly so, lepidote, about 
bed Mle long. 

Type in the University of Michigan Herbarium, Eizi Matuda 
3327 (flowers), collected in advanced forest et La Palma, 
near Balancan, Tabasco, Mexico, June 1--6, 1939. At Reforma, 
near Balancan, Matuda 3160 (fruits), May 22--26, 1939. 


CROTON PETENENSIS Lundell, sp. nov. 

Arbor, stellato-lepidota. Petioli 1--3.5 cm. longi, apice 
biglandulosi. Lamina membranacea, crenato-dentata, elliptico 
-oblonga vel obovato-elliptica, apice rotundata, emarginata, 
vel obtuse subacuminata, basi obtusiuscula, utrinque parce 
stellato-lepidota. Infructescentiae usque ad 11.5 cm. long- 
ae. Capsula parce stellato-lepidota, ca. 9 mm. longa.. 

A tree 8 m. high, stellate-lepidote, the scales silvery. 
Brenchlets rather slender, at first densely stellate-lepid- 
ote. Fetioles very slender, canaliculete, 1 to 3.5 cm. long, 
beering at apex on upper side two prominent subsessile 
glands. Leaf-blades thin-membranaceous, crenate-dentate, the 
teeth gland-tipped, elliptic-oblong or obovate-clliptic, 3 
to 11 cm. long, 1.4 to 4.7 cm. wide, apex rounded, emargina- 
te or obtusely subacuminate, base obtusish, at first sparse- 
ly stellate-lepidote on both surfaces, glabrescent above 
with age, base 3-nerved, costa elevated beneath, veins prom- 
inulous. Pistillate flowers unknown. Staminate flowers in 
bud only, pedicellate, stellate-lepidote; calyx-lobes 5, 
petals 5, the petals pilose; stamens 16, filaments glabrous; — 
receptacle pilose. Infructescence terminal, up to 11-5 cm. 
long; fruiting pedicels stout, recurved, 3 to 4 mm. long. 
Capsules rather sparsely stellate-lepidote, about 9 mm. lom 

Type in the University of Michigan Herbarium, C. L. Lund- 
ell 4128, collected in secondary forest along Aguada Tigre-- 
Yaxha road, Department of Petén, Guatemala, June 17, 1933. 


1940 Lundell, New Species of Croton 407 


CROTON PSEUDOGLABELLUS Lundell, sp. nov. 

Arbor, lepidota. Petioli 2--4.5 mm. longi. Lamina charta- 
cea, integra, eglandulosa, ovata, apice obtusa vel acutius- 
cula, basi rotundata, lepidota. Inflorescentiae bisexuales. 
Flores Y subsessiles; calyce cae. 2.8 mm. longo, lobis 5, ov- 
atis vel lanceolatis. Flores & subsessiles; steminibus 9-- 
ll. 

A tree 6 m. high, lepidote; branchlets erect, rather 
stout, angled. Petioles 2 to 4.5 mm. long. “eaf-blades 
chartaceous, entire, ovate, 2.2 to 5.5 cm. long, 1.2 to 3.1 
ome wide, apex obtuse or acutish, base rounded, at first 
densely lepidote on both surfaces, glabrescent above with 
age, costa prominent beneath, less conspicuous above, pinn- 
ately veined, the veins obscure. Racemes axillary, bisexual, 
crowded, up to 2.3 cm. long, ferruginous-lepidote. Pistill- 
ate flowers with pedicels about 1.6 mm. long; calyx about 
2.8 mm. long, deeply S-lobed, the lobes ovate or lanceolate; 
petals 5, spatulate, about 2.5 mm. long, short-clawed, 
villous-ciliate; ovary ferruginous-lepidote; styles 3, 
branched 3 times. Staminate flowers with pedicels about 1 
mme long; stamens 9 to ll, filaments sparsely pilose below 
middle. 

Type in the University of Michigan Herbarium, G. F. Gaum- 
er 1407, collected at Lake Chichankanab, Quintana Roo, Mexi- 
CO. 

The small ovate leaves and compact inflorescences distin- 
guish this species from C. glabellus L. and its allies. 


CROTON SUTUP Lundell, sp. nov. 

Frutex, stellato-tomentosa. Petioli 0.5--1.8 cm. longi. 
Lamina chartacea, integra, eglandulosa, oblanceolata vel ob- 
longo-elliptica, apice acuta vel acuminate, basi obtusa. In- 
florescentiae bisexuales. Flores 9 pedicellati; calyce 5.5 
mms longo, lobis lineari-oblongis. Flores & pedicellati; 
staminibus ll. . 

A shrub, about 2 m. high, with soft whitish or brownish 
tomentum of stellate hairs. Branchlets slender, brownish to- 
mentose with age. Stipules aristate. Fetioles slender, 0.5 
to 1.8 cm. long. Leaf-blades chartaceous, entire, oblanceol- 
ate or oblong-elliptic, 4 to 8 om. long, 1.2 to 3.8 om. 
wide, apex acute or acuminate, base obtuse, green and spar- 
sely hairy above, whitish tomentose beneath, costa and veins 
impressed above, prominent beneath, pinnately veined with 
base 4-nerved. Racemes terminal, bisexual, up to 8.5 cm. 
long, whitish tomentose. Pistillate flowers with stout pedi- 
cels about 2 mm. long, sharply accrescent; calyx 5.5 mm. 
long, accrescent, S-lobed, the lobes linear-oblong; styles 
3, digitately lobed. Staminate flowers with slender pedicels 
about 4 mm. long; calyx 2.5 mm. long, lobes ovate~oblong; 


408 PHYTOLOGIA No. 12 


petals 5, oblanceolate, about 2.8 mm. long, ciliate, the 
basal hairs elongated; stamens 11, filaments sparsely pilo- 
se below middle. Immature capsules stellate-tomentose. 

Type in the University of Michigan Herbarium, C. L. lund- 
ell and Amelia A. lundell 7363, collected in low second 
growth near Piste, Yucatan, Mexico, June 4, 1938; vernacular 
name “sutup". 


CROTON TABASCENSIS Lundell, sp. nov. 

Arbor, stellato-hirsuta vel tomentosa. Fetioli usque ad 
1.8 cme longi. Lamina chartacea, ovata vel oblongo-ovata, a- 
pice acuta vel subacuninata, basi subcordata vel rotundata, 
glabrata, margine minute denticulata. Inflorescentiae bisex- 
uales. Flores > pedicellati; calyce ca. 6.5 mm. longo, lobis 
5, ovatis. Flores & pedicellati; calyce ca. 4 mm. longo, lo- 
bis 5; staminibus 15 vel 16. Capsula stellato-hirsuta. 

A tree 7 m. high and 25 cm. in diam., stellate hairy 
throughout. Branches stout; branchlets slender, densely 
short-hirtellous with appressed stellate hairs, the central 
ray elongated sharply. Petioles slender, up to 1.8 cm. long, 
shallowly canaliculate. Leaf-blades chartaceous, ovate or 
oblong-ovate, 2.5 to 7 cme long, 1.4 to 3.4 cm. wide, apex 
acute or short-acuminate, base shallowly cordate or rounded, 
minutely denticulate, eglandular, with scattered appressed 
stellate hairs on both surfaces, glabrate with age, palmate- 
ly veined, basal veins 5 or 7, costa and veins promimlous 
and whitish beneath, inconspicuous and slightly impressed a- 
bove. Cymes terminal, bisexual, up to 9 cm. long, rachis and 
flowers stellate-tomentose. Pistillate flowers with pedicels 
3 to 6 mm. long; calyx about 6.5 mm. long, deeply S-lobed, 
the lobes ovate, about 5 mm. long, 4 mm. wide, irregularly 
toothed below middle, acutish; ovary stellate-tomentose; 
styles stellate hairy, twice branched. Staminate flowers 
with pedicels up to 3 mm. long; calyx 4 mm. long, the lobes 
ovate, entire, ebout 3 mm. long, 2 mm. wide; petals 5, nar=- 
rowly oblong-elliptic, about 3.5 mm. long, 2 mm. wide, pil- 
ose; stamens 15 or 16, filaments pilose below middle, 3 to 4 
mme long; receptacle pilose. Capsule densely stellate-hirs- 
ute, about 7 mm. long (hamatuea) 

Type in the University of Michigan Herbarium, Eizi Matuda 
3145, collected at Reforma, near Balancan, Tabasco, Niexico, 
May 22--26, 1939. 

Probebly related to C. Soliman Schlecht. & Cham. 


CROTON YUCATANENSIS Lundell, ep. nove - 

Frutex, stellato-lepidota. Petioli usque ad 3.5 cm. lon- 
gi. Lamina membranacea, subintegra, lanceolata vel ovata, e- 
glandulosa, apice acuminata, ecuta vel obtusa, basi subcord- 
ata vel rotundata, subtus albido-lepidota. Inflorescentiae 


1940 lundell, New Species of Croton 409 


bisexuales, ad 21 om. longae. Flores $ pedicellati; calyce 4 
mme longo, lobis late ovatis. Flores & pedicellati; stamini- 
bus 13--18. Capsula ca. 5 mm. longa. 

Arborescent shrub, up to 2 me. high, with whitish scale- 
like stellate hairs. Branchlets slender. Stipules reniforn, 
at first small and long-aristulate-cuspidate, becoming foli- 
aceouse Petioles slender, up to 3.5 cm. long. Leaf-blades 
membranaceous, Obscurely and minutely denticulate, appearing 
entire, lanceolate or ovate, 2 to 10 cm. long, 1.3 to 5 cme 
wide, apex acuminate, acute or obtuse, base subcordate or 
rounded, green and very sparsely lepidote. above, densely 
white lepidote beneath, base 5= or 7-nerved, costa and veins 
conspicuous beneath, slightly impressed above. Racemes very 
slender, terminal, bisexual, up to 21 cme long. Pistillate 
flowers with pedicels about 2.5 mm. long, acorescent; calyx 
4 mm. long, thin, sharply accrescent, 5-lobed to middle, the 
lobes broadly ovate, acutish, becoming rounded, ruffled with 
age; petal vestiges pilose; ovary white atellate-lepidote; 
styles branched 3 or 4 times. Staminate flowers with slender 
pedicels about 5 mm. long; calyx 3 mm. long, lobes ovate; 
petels linear, variable in number; perfect stamens 13 to 18, 
filaments pilose. Capsules about 5 mm. long. 

Type in the University of Michigan Herbarium, C. L. Lund- 
ell and Amelia A. lundell 7400, collected in second growth 
at kilometer 77, “erida-Chichen Itza road, Yucatan, Mexico,. 
June 3, 1938. In low second growth bordering Secred Cenote 
at Chichen Itza, Lundell and lundell 7524, June 12, 1938. 

Related to C. Watsonii Standl. 


NOVELTIES IN THE AVICENNIACEAE AND VERBENACEAE 


Harold N. Moldenke 
r te) 


AVICENNIA BALANOFHORA Stapf & Moldenke ex Moldenke, Geogr. 
Distrib. 34, nom. nud. (1939), sp. nov. 

Arbor; ramulis gracillimis subteretibus dense griseo- 
farinaceis glabrescentibus; nodis annulatis articulatis; 
petiolis dense griseo- vel flavido-farinaceis; leminis firme 
chartaceis vel subcoriaceis lanceolatis vel anguste ellipti- 
cis acutis vel longe acuminatis vel caudatis integris, ad 
basin attenuatis vel longe acuminatis, supra obscure pulver- 


410 PHYTOLOGIA No. 12 


ulentis vel glabris, subtus densissime incano- vel flavido- 
farinaceis; inflorescentiis axillaribus terminalibusque, 
capitatis solitariis vel geminatis; fructibus oblongis ad a- 
picem et basin rotundatis, ad apicem apiculatis. 

Small tree; branchlets and twigs very slender, subterete, 
densely grayish-farinaceous, becoming glabrous in age, ob- 
scurely or not at all lenticellate; nodes swollen, annulate, 
articulate; principal internodes 2.8--6.5 cm. long; leaves 
decussate-opposite; petioles rather slender, 0.8--2 cm. 
long, densely farinaceous with gray or yellowish hairs, 
flettened above, wrinkled longitudinally beneath in drying, 
slightly ampliate at base; blades firmly chartaceous or sub- 
coriaceous, dark-green and rather shiny above, incanous or 
flavidous beneath, lanceolate or narrowly elliptic, 2.5--9.3 
em. long, 6--24 mm. wide, varying from acute (on young 
leaves) to long-acuminate or caudate (on mature leaves) at 
apex, entire, attenuate or long-acuminate into the petiole 
at base, very obscurely fine-pulverulent or glabrate above, 
very densely incanous- or yellowish-farinaceous beneath; 
midrib slender, slightly prominulous above and usually can- 
aliculate to the apex or to 1/2 or 3/4 the distance from the 
base, rounded-prominulous beneath; secondaries very slender, 
ascending, slightly prominulous on both surfaces, arcuately 
joined in meny loops at the margins, mostly obscure beneath; 
vein and veinlet reticulation sparse, the larger portions 
subprominulous on both surfaces, mostly obscure or hidden 
beneath; inflorescence axillary and terminal, the axillary 
ones solitary or paired, capitate, 1.4--5 cm. long, 5-19 
mm. wide, several-flowered, the terminal one often 3-branch- 
ed et apex and there conspicuously bracteate; bracts, when 
present, foliaceous, very narrowly elliptic, short-stipitate 
or subsessile, 17--25 mm. long, 4--6 mm. wide, densely far- 
inaceous with incanous or yellowish furf on both surfaces, 
acute at apex, attenuate or acuminate at base; fruiting- 
calyx practically unchanged, densely short-pubescent, its 
segments and the similar bractlets end prophylla mostly 
brown-margined, closely appressed to the base of the fruit; 
fruit oblong, 6--9 mm. long, 4--4.5 mm. wide, rounded at 
both ends, very densely puberulent with grayish-flavidous 
hairs, apiculate at (usually) the exact center above. 

The type of this species was collected by Ferdinand von 
Mueller along the Brisbane River, Queensland, Australia, and 
is deposited in the herbarium of the Royal Botanic Gardens 
at Kew. It is known also from Keppels Islands, Queensland. 


AVICENNIA ALBA var. LATIFOLIA Moldenke, Geogr. Distrib. 34, 
nome nud. (1939), var. nov. 
Haec varietas a forma typica speciei recedit laminis fol- 
iorum meturis late ellipticis usque ad 6.5 cm. latis. 


CS ee at be ee 
pee ne at ee 1 


1940 Moldenke, Novelties AY) 


This variety differs from the typical form of the species 
in its mature leef-blades being broadly elliptic, to 6.5 cm. 
wide. 

The type was collected by Noerkas (no. 58) on the Van 
Vuuren Expedition at Cape Wadjo on Celebes Island, and is 
deposited in the Rijksherbarium at Leiden. 


AVICENNIA MARINA var. ACUTISSIMA Stapf & Moldenke ex Molden- 
ke, Geogr. Distrib. 32, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit laminis fol- 
iorum ad apicem argute acutis vel acuminatis; petiolis brev- 
issimis; tomentis ramulorum pedunculorum elbis. 

This veriety differs from the typical form of the species 
chiefly in its decidedly sharp-acute or acuminate leaf-apex. 
The branches, branchlets, twigs, peduncles, brectlets, pro- 
phylla, and fruits are also more plainly and regularly white 
~farinaceouse The petioles are very uniformly extremely 
short, 1--6 mm. long. The fruit is not beaked. 

The type was collected by R. Ke Phide in the creek near 
Boriwle Station of the B. Be. & C. I. Re Re, Salsette Konkan, 
Bombay, India, on April 4, 1904, and is deposited in the 
herbarium of the Royal Botanic Gardens at Kew. The veriety 
is known thus far only from Bombay and Sind. 


AVICENNIA MARINA ver. ANOMALA Moldenke, Geogr. Distrib. 455, 
nome nud. (1939), var. nove 

Haec varietas e forma typica speciei recedit inflorescen- 
tiis attenuatis, floribus geminatis oppositis vel trregular- 
iter dispositis. 

This variety differs from the typical form of the species 
in its attenuete inflorescences with scattered or opposite 
pairs of flowers. 

The type was collected by De Henne and Carl Wilhelmi on 
Low Ieiend, Tasmania, and is deposited in the Bernhardi Her- 
barium at the Botanisches Museum in Berlin. 


BURROUGHSIA HMoldenke, gene nove 

Plantee suffrutescentes; caulibus sublignosis multiramo- 
sis brevibus; remis numerosis gracilibus plerumque fascicu- 
latis; foliis simplicibus oppositis vel ternatis sessilibus 
plerumque lanceolatis; inflorescentiis axillaribus spicatis 
vel capitatis; bracteolis ovatis vel ovato-lanceolatis dist- 
inctis acutis; calyce bipartito persistente; corolla hypo- 
crateriformi, limbo bilabiato 5-partito; staminibus 4 didyn- 
amis; antheris ventralibus longe appendiculatis; stigmate 
Obliquo discoideo. 

Somewhat woody perennials or shrubby, much branched; 
stems short, somewhat ligneous, covered with a light-colored 
cracked cortex; branches numerous, slender, flexuous, ascen- 


Ayo PHYTOLOGIA Noe 12 


ding, often fascicled; leaves simple, opposite or ternate, 
sessile, small, usuelly lanceolate in outline, pinnatifid or 
crenate, often plicate and prominently veined beneath; in- 
florescence axillary, spicate or capitate; peduncles solit- 
ary, 2--4 times as long as the subtending leaves; heads hem- 
ispheric or oblong and few-flowered, or spikes oblong and 
dense; bractlets ovate or ovate-lanceolate, distinct, acute, 
ciliate, pubescent; calyx much exceeding the bractlets, 2- 
parted, persistent, united nearly to the top and truncete or 
minutely 2-toothed; corolla purplish or white, much exsert- 
ed, hypocrateriform, its tube cylindric, its limb bilabiate, 
the upper lip 2-lobed, the lower 3-cleft, all the lobes 
crenate or crisped, the ventral lobe somewhat larger than 
the others; stamens 4, didyneamous; filaments longer than the 
enthers, the ventral ones with slender erect appendages 
springing from the connective and exceeding the anthers in 
length; anther-cells somewhat separate and oblique; pistil 
very short; stigma strictly simple, oblique, disk-like; ov- 
ules one in each cell. 

This remarkable genus of the Verbenaceae combines the 5- 
parted corollas and appendeged anthers of Verbena and the 2- 
cleft celyx and entire oblique stigmas of Lippia. It is nam- 
ed in honor of John Burroughs (April 135, 1837--March 29, 
1921), distinguished American naturalist, philosopher, and 


writer. Type species, B. appendiculata. 


BURROUGHSIA APPENDICULATA (Robinson & Greenm.) Moldenke 
Pests <M) kt Twe4d- Nenies? 30, tigpanyas CSA 
combe nOvVe 


Lippia appendiculata Robinson & Greenm., Préc. Am. Acad. 
29: 390. 89h. 


BURROUGHSIA FASTIGIATA (T. S. Prandege) Moldenke, Suppl. 
List Common Names 6, nome nud. (1940). Prnlim. Alph. 
List Invalid Names 30, hyponym (1940), comb. nov. 
Lippie fastigiata T. S. Prandeg., Proc. Calif. Acad., 
sere 2» 2: 196. 1889. 


CITHAREXYLUM AFFINE var. GLANDULIFERUM Moldenke, Geogr. 
Distrib. 13, nom. nud. (1939), var. nov. 

Faec varietas a forma typica speciei recedit laminis 
foliorum late ovatis, usque ad 12 cm. longis et 6.5 cme la-~ 
tis, irregulariter arguto-dentatis, subtus puberulentibus et 
glanduliferis; glandulis paucis discoideis. 

This variety differs from the typical form of the species 
in its broadly ovate leaf-blades, which are to 12 cm. long 
and 6.5 cm. wide, irregularly sharp-dentate along the marg- 
ins when mature, puberulent throughout on the lower surface, 
and glandulose with several brown disk-like glands along the 


r 
* . - a 
eee ee eee 


1940 Moldenke, Novelties 413 


midrib and at the base beneath. The sharply tetragonal and 
glabrous branches, branchlets, and twigs are many~striate 
and abundantly impressed glandular-punctate (!). 

The type of this variety was collected by George B. Hint- 
on (no. ) in an oak woods at Yperricones, Tamasceltepec, 
Mexico, Mexico, on February 7, 1935, and is deposited in the 
herbarium of the Royal Botanic Gardens at Kew. 


CITHAREXYLUM GHIESBREGHTII Moldenke, spe nov. 

Frutex; ramulis obtuse tetragonis sparse strigillosis 
vel glabratis nitidulis; petiolis gracilibus obscure spars- 
issimeque pulverulento-puberulis vel glabratis; leminis lan- 
ceolatis vel ellipticis attenusto-subacuminatis integris, ad 
basin acutis, utrinque glabratis vel sparse punctatis; in- 
florescentiis axillaribus confertis racemiformibus erectis 
vel nutentibus. 

Shrub; branchlets slender, buff or brownish, rather 
shiny, striate in drying, obtusely tetragonal, sparsely 
strigillose or glabrate, somewhat ampliate at the nodes; 
nodes annulate, mostly oblique; principal internodes 5--6 
em.e long; leaves decussate-opposite; petioles slender, 1.5-- 
2e2 cme long, obscurely and very sparsely pulverulent-puber- 
ulent or glebrate, not noticeably ampliate at base, flatten- 
ed above, usually somewhat canaliculeate-margined toward the 
apex; blades thin-chartaceous, rather uniformly dark-green 
on both surfaces, lanceolate or elliptic, 16--21 cm. long, 
4,1--5.5 ome wide, attenuate to a subacuminete point at the 
apex (or obtuse and emarginate on stunted leaves), entire, 
acute at base, glabrate and sparsely punctate on both surf- 
eces; midrib slender, flattened above, prominent beneath; 
secondaries very slender, 10--135 per side, arcuate-ascend- 
ing, flattened or subprominulous above, sharply prominulous 
beneath, not conspicuously anastomosing at the margins; vein 
and veinlet reticulation very abundant, prominulous through- 
out above, plane beneath; inflorescence axillary, usually 
crowded at the tips of the branchlets, racemiform, 9.5--22 
cme long, about 1 cm. wide throughout, densely many-flower- 
ed, erect or nutant; peduncles slender, to 4.5 cm. long, us- 
ually more abbreviated, sparsely strigillose or glabrate; 
pedicels very slender, about 1 mm. long, sparsely strigillo- 
se or glabrate, spreading; prophylla equaling the pedicels, 
sessile, sharp-pointed; calyx campanulate, about 2 mm. long 
and wide, glabrous except for the puberulent rim; corolla 
about 5 mm. long, its limb about 5 mm. in diameter. 

The type of this species was collected by Auguste Boni- 
face Ghiesbreght -- in whose honor it is named -- in the 
state of Yaxaca, “exico, in 1842, and is deposited in the 
herbarium of the Muséum National d'Histoire Naturelle at 
Paris. 


Ad Pen YS 0: LOG. TA No. 12 


CITHAREXYLUM HIDALGENSE Moldenke, Geogr. Distrib. 13, nome 
nude (1939), sp. nov. 

Frutex vel arbor; remis ramulisque obtuse tetragonis 
breviter hirsutis vel furfuraceis glabrescentibus; sarmentis 
gracilibus nigrescentibus sparse breviterque hirsutis; peti- 
olis gracilibus hirsutis; foliis membranaceis oblongo- 
ellipticis vel subobovatis acutis vel breviter acuminetis 
integris vel ad apicem asymmetrico-angulatis, ad basin acut- 
is vel subacuminatis glanduliferis, supra hirsuto-puhescent- 
ibus vel in senectute dense puberulis, subtus dense hirsut- 
is; inflorescentiis axilleribus vel terminelibus nutantibus 
simplicibus multifloris. 

Shrub or tree, to 7 m. tall; brenches and branchlets med- 
ium, gray, obtusely tetragonal, short-hirsute or furfurace- 
ous at the apex, soon becoming glabrescent, lenticellate; 
twigs slender, nigrescent in drying, sparsely short-hirsute; 
nodes obscurely annulate; principal internodes 1--3.5 cm. 
long; leaves decussate-opposite; leaf=scars large, borne on 
extremely large and massive, divergent-ascending, corky 
sterigmata to 5 mm. long; petioles slender, 1.5--3.3 cm. 
long, more or less hirsute; blades membranous, brunneous or 
nigrescent above in drying, lighter beneath, oblong-elliptic 
or elliptic to subobovate, 5.2--11 cm. long, 2--/7./ cm. wide, 
acute or short-acuminate at apex, asymmetrically eangulate 
near the apex or entire, acute or subacuminate and often 
more or less asymmetrical at the base, usually bearing 1 or 
2 black glands at the very base, shortly hirsute-pubescent 
above, becoming merely densely puberulent in age, densely 
hirsute (especially along the larger venation) beneath; mid- 
rib slender, not prominent on either surface; secondaries 
very slender, 5--8 pairs, arcuste-ascending, not prominent 
on either surface, conspicuous beneath only because of their 
more or less distichous pubescence; vein and veinlet reticu- 
lation obscure or indiscernible; racemes axillary or termin- 
al, nutant, 8--16 cms long, about 2 cm. wide in anthesis, 
simple, many-flowered; peduncles and rachis slender, fiore or 
less hirsute-pubescent, brown, the former 5--7 mm. long; 
pedicels filiform, about 2.5 mm. long, short-pubescent; 
bracts and bractlets none; prophylle linear-setaceous, about 
1 ma. long; fruiting-calyx light, cupuliform, about 4 mm. 
long and 8 mm. wide, S-ribbed, minutely puberulent or glab- 
rescent, its rim shallowly S-angulate or subentire; fruit 
oblong, about 10 mm. long and 7 m. wide, fleshy, glebrous, 
shiny, wrinkled and blackening in drying. 

The type of this species was collected by Cyrus Guernsey 
Pringle (no. 8969) below the Trinided Iron Works, at en alt- 
itude of 5000 feet, Hidalgo, Mexico, on April 22, 1904, and 
is deposited in the Britton Herbarium at the New York Botan- 
ical Garden. It is closely related to C. Sessaei D. Don and 


i i iit lO 


1940 Moldenke, Novelties 415 
is known also from Zacatecas, Veracruz, and Puebla. 


CITHAREXYLUM MOCINI var. LONGIBRACTEOLATUM Moldenke, vare 
NOVe 

Haec varietas a forma typica speciei recedit bracteolis 
elongatis linearibus vel spathulatis 4--25 mm. longis. 

This variety differs from the typical form of the species 
in having the bractlets of its flowering racemes linear or 
narrowly spatulate, 4--10 mm. long, elongating in fruiting 
racemes to 25 mm. and then even more conspicuous, persist- 
ente 

The type of this variety was collected by Zizi Matuda 
(no. S.181) on Mount Pasitar, Chiapas, Mexico, on December 
28, 1936, and is deposited in the Britton Herbarium at the 
New York Botanical Garden. 


CITHAREXYLUM SHREVEI Moldenke, Geogr. Distrib. 14, nom. nud. 
(1939), sp. nov. 

Frutex; remis ramulisque gracilibus griseis subteretibus; 
sarmentis gracillimis obtuse tetragonis minute brevissime- 
que albido-puberulis; foliis oppositis; petiolis gracillimis 
vel obsoletis; leminis chartaceis anguste oblongis vel ell- 
ipticis acutis vel obtusis integris revolutis, ad basin acu- 
tis vel subcuneatis, utrinque strigilloso-scabrellis. 

Shrub; branches and branchlets slender, gray, subterete; 
twigs extremely slender and tenuous, white of very light 
gray, minutely puberulent with very short albidous hairs, 
obtusely tetragonal; nodes not annulate; principal inter- 
nodes 0.4--2.5 cm. long; leaves decussate-opposite; petioles 
very slender and less than 1 mm. long or obsolete; blades 
chertaceous, grey-green on both surfaces, narrowly oblong or 
elliptic, 0.4 to 2.2 cm. long, 2.5--5.5 mm. wide, bluntly a- 
cute or obtuse st apex, entire and usually more or less rev- 
Olute along the margins, acute or subcuneate at base, strig- 
illose-roughened on both surfaces with minute albidous 
bulbous-based hairs; midrib very slender, very slightly pro- 
minulous beneath; secondaries about 3 per side, very slend- 
er, usually indiscernible above and obscure beneath; vein 
and veinlet reticulation indiscernible; inflorescence and 
fruit not seen. 

The type of this species was collected by Daniel Trembly 
MacDougal and Forrest Shreve (no. 59) at Picu Pass, Sonora, 
Mexico, on November 19, 1923, and is deposited in the United 
States National Herbarium at Washington. It is named in 
honor of my good friend, Dr. Forrest Shreve, of the Carnegie 
Institution's Desert Laboratory at Tucson, Arizona, who has 
contributed so much to our knowledge of the flora of the 
southwestern United States and Mexican deserts. 


416 Pa FT ULOsSers No. 12 


ALOYSIA BARBATA (T. S. Brandeg.) Moldenke, Prelim. Alph. 
List Invalid Names 30 & 31, hyponym (1940), comb. nov. 
Lippia barbata T. S. Brandeg., Proc. Calif. Acad. Sei., 
sere 25 2: 196. 1889. 


CLERODENDRUM AFRICANUM Moldenke, spe nov. 

Frutex vel arbor; remis ramulisque gracillimis submargin-~ 
ato-tetragonis glabris; foliis oppositis; petiolis glabris; 
laminis membranaceis anguste oblongo-ellipticis acutis, ad 
basin acutis vel attenuatis, utrinque pulverulentis vel 
glabris, subtus punctatis; inflorescentiis axillaribus; cym- 
ulis solitariis unifloris. 

Tree or shrub; branches and branchlets very slender, 
brunnescent, submargined-tetragonal, glabrous; leaves oppos- 
ite; petioles 1--4 mm. long, brunnescent, glabrous; blades 
membranous, dark-green (nigrescent in drying) above, paler 
beneath, narrowly oblong-elliptic, 1.5--5.5 cm. long, 5.5-- 
14 mm. wide, acute at apex, acute or attenuate at base, pul- 
verulent or glabrous on both surfaces, punctate beneath; 
midrib very slender, flet or subimpressed above, flat or 
very slightly subprominulous beneath, nigrescent; secondar= 
ies few, very slender, 2--5 per side, irregularly arcuate- 
ascending, plainly anastomosing near the margins, obscure or 
subimpressed above, flat and nigrescent beneath; vein and 
veinlet reticulation indiscernible on either surface; in- 
florescence axillary; cymules single, opposite, about equal- 
ing the subtending leaves, 1-flowered; corolla purple. 

The type of this species was collected by J. Wylie in the 
Greytown District, Natal, Union of South Africa, in Decemb- 
er, 1932, and is deposited in the Natal Government Herbarium 
at Durban. 


CLERODENDRUI£ LIGUSTRINUM var. NICARAGUENSE Moldenke, Alph. 
List Common Nemes 16 & 28, nom. nud. (1939); Geogr. 
Distrib. 16 & 37, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit foliis et 
cymis axillaribus plerumque ternatis et laminis foliorum 
subtus constanter puberulentibus. 

This variety differs from the typical form of the species 
in its leaves and axillary cymes being generally ternate and 
its leaf-blades being regularly puberulent beneath. 

The type of this variety was collected by D. Chaves (no. 
227) et Managua, Nicaragua, on July 26, 1926, and is depos- 
ited in the United States National Herbarium at Washinzton. 


CLERODENDRUM PITTIERI Moldenke ex Standl., Field Mus. Publ. 
Bot. 18: 1003, without Latin diag. (1938); Geogr. Dist- 
rib. 15, 17, 19, 20, & 22, nom. nud. (1939), sp. nov. 

Frutex vel fruticulus; ramis sarmentosis rigidis; ramulis 


1940 Moldenke, Novelties 417 


sarmentisque subgracilibus rigidis minute puberulis vel sub- 
glabrescentibus; internodiis abbreviatis; foliis oppositis 
vel ternatis; petiolis gracillimis 1--5 mm. longis pulveru- 
lento-puberulis vel glabratis; laminis tenuiter chartaceis 
anguste ellipticis, ad apicem obtuse rotundatis, integris, 
ad basin acutis vel subcuneatis, utrinque glabris, subtus 
dense punctatis; inflorescentiis axillaribus, cymis abbrevi- 
atis 1--3-floris; pedunculis gracilibus minutissime puberul- 
is; pedicellis subglebratis. 

Bush or shrub; branches twiggy, stiff; branchlets and 
twigs rather slender, light-gray, minutely puberulent with 
greatly appressed hairs, becoming subglabrate in age, the 
twigs often short, stiff, and sharp-pointed; nodes not annu- 
late, marked with 2 or 3 aculeate leaf-scars; principal in- 
ternodes much abbreviated, 3--30 mm. long; leaves decussate- 
Opposite or ternate; petioles very slender, 1--5 mm.-long, 
very obscurely pulverulent-puberulent or glabrate; blades 
thin-chartaceous, rather dark-green on both surfaces, nar~ 
rowly elliptic, 6--24 mm. long, 3.5--10 mm. wide, rounded or 
obtuse at apex, entire, acute or subcuneate at base, glab- 
‘rous on both surfaces but densely punctate beneath; midrib 
very slender, almost indiscernible or subimpressed above, 
prominulous beneath; secondaries very slender, 2--4 per 
side, indiscernible above, obscure beneath; vein and veinlet 
reticulation indiscernible; inflorescence exillary; cymes 
abbreviated, solitary, opposite, in the upper axils, 1--3- 
flowered, to 5-5 om. long (including the mature flower), 
lax; peduncles slénder, 8--14 mm. long, very minutely and 
obscurely puberylent with appressed hairs; pedicels slender, 
5--8 mm. long, subgleabrate; bracts and bractlets none; pro- 
phylla linear-subulate, few, 1--2 mm. long; calyx about 3 
mme long, minutely puberulent or glabrate, truncate; corolla 
white, about 2 cm. long, with a long and slender tube. 

The type of this ee ae was collected by Henri Frangois 
Pittier de Fabrega (no. 4965) -- in whose honor it is named 
-- along the outskirts of the tidal belt and in savannas 
near sea-level at Aguadulce, Coclé, Panama, on December 5, 
1911, and is deposited in the United States National Herbar- 
ium et Washington. The species is also known from Guatemala, 
Colombia, and Venezuela. 


CLERODENDRUM SCHEFFLERI var. ELLIPTICUM Moldenke, var. nov. 

Haec verietas a forma typica speciei recedit gracillimis 
sparse strigillosis; leminis tenuiter membranaceis late ell- 
ipticis acutis, ad basin acuminatis, utrinque sparsissime 
puberulento-strigillosis vel glabratis. 

This variety differs from the typical form of the species 
in the following characters: leaf~scars large, corky, eleva- 
ted, with flaring margins; leaves opposite; petioles very 


418 PHYTOLOGIA No. 12 


slender, 5--1l mu. long, sparsely strigillose; blades thin- 
membranous, darkgreen or brunnescent above, lighter beneath, 
broadly elliptic, 2.8--8 cm. long, 1.7--4.7 cm. wide, acute 
at apex, acuminete at base, very sparsely puberulent-strig- 
illose or glabrate on both surfaces; calyx parted half way 
or farther. 

The type of this varisty was collected by Maurice Smeth- 
urst 3vans (no. 544a) at Berea, Durban, Natal, Union of 
South Africa,.in October, 1894, at an elevetion of 5000-6000 
feet, and is deposited in the Natal Government Herbarium at 
Durban. It is described as a bush with violet flowers. 


CLERODENDRUM SUFFRUTICOSUM var. NATALZENSE Moldenke, var.nov. 

Haec varietas a forma typica speciei recedit foliis oppo- 
sitis; leminis ellipticis vel subobovatis acuminatis vel 
subcaudatis, ad basin acuminatis, marginibus densiuscule ir- 
regulariterque inciso-dentatis (dentibus saepe incurvatis 
vel recurvetis), utrinque leviter adpresso-pubescentibus. 

This variety differs from the typical form of the species 
in its leaves being opposite; retioles 3--7 mm. long, dense- 
ly appressed-pubescent; blades membranous, elliptic or sub- 
obovate, dark-green above, lighter beneath, 2--4.5 cm. long, 
1--2 cm. wide, acuminate at base and apex or subcaudate at 
apex, the mergins rather deeply end irregularly incised with 
antrorse teeth which are often incurved or recurved at the 
tip, finely appressed-pubescent on both surfaces, more dens- 
ely so beneath; calyx split half way to the base. 

The type of this variety was collected by John Medley 
Wood (no. 657) at Inanda, Natal, Union of South Africa, in 
September, 1880, and is deposited in the Natal Government 
Herbarium at Durban. The collector states that four segments 
of the corolla are white end the fifth is violet. The vari- 
ety is also known from Trensvaal and Zululand. 


GMELINA SPECIOSA Moldenke, sp. nove 

Arbor; ramulis gracilibus lenticellatis densissime puber- 
ulis; foliis oppositis; petiolis gracilibus densissime pub- 
erulis; laminis late ovatis vel deltoideis acutis vel brevi- 
ter acuminatis integris, ad basin truncatis vel acutis vel 
breviter acuminatis, supra glabratis nitidulis, subtus dens- 
issime adpresso-puberulis; inflorescentiis terminalibus pan- 
iculatis; calyce glanduloso. 

Tree, to 8 m. tall; branchlets slender, prominently lent- 
icellate, very densely puberulent with yellowish-brown pub- 
erulence, becoming less conspicuously so in age; leaves dec- 
ussate-opposite; petioles slender, 2--5 cm. long, very den- 
sely puberulent; blades broadly ovate or deltoid, dark-green 
above, grey beneath, 6--20 cm. long, 4.3--11.5 cm. wide, ac- 
ute or short-acuminate at apex, entire, varying from trunc- 


1940 Moldenke, Novelties 419 


ate to abruptly acute or short-acuminate at base, glabrete 
and rather shiny above, extremely densely appressed-puberul- 
ent with gray puberulence beneath; inflorescence terminal, 
paniculate; flowers very large; calyx distinctly glandulose, 
the glands small, numerous, basal. 

The type of this species was collected by Benedict Balen- 
sa (no. 3806) in forests in the valley of Lankok, Mont-Bavi, 
Tonkin, French Indo-china, on June 24, 1887, end is deposit- 
ed in the herbarium of the Royal Botenic Gardens at Kew. 


GMELINA TONKINENSIS Moldenke, sp. nov. 

Frutex subspinosus; ramulis gracillimis dense breviterque 
pubescentibus vel dense puberulis; foliis oppositis; petio- 
lis grecillimis dense breviterque pubescentibus; leminis o- 
vatis acutis vel breviter acuminetis integris, ad basin cor- 
detis vel truncatis vel abrupte acutis, supre sparsissime 
strigilloso-pubescentibus glabrescentibus, subtus dense 
breviterque pubescentibus et puberulis; inflorescentiis ter- 
minalibus racemiformibus; calyce distincte glanduloso. 

Somewhat spiny shrub; branchlets very slender, the young- 
er parts densely short-pubescent with fulvous hairs, the 
older parts merely densely puberulent, sparsely but promin- 
ently lenticellate; leaves decussate-opposite; petioles very 
slender, 0.8--6 cm. long, densely short~pubescent with fulv- 
ous hairs like the young twigs; blades thin-chartaceous or 
membranous, ovate, 3./--13 cm. long, 2-.5--9.1 cm. wide, a- 
cute or short—acuminate at epex, entire, varying from cord- 
ate or truncate to abruptly acute at base, very sparsely 
strigillose-pubescent (especially along the larger venation) 
above, becoming glabrous, densely short-pubescent beneath 
over the lower puberulence; inflorescence terminal racemi- 
form; calyx small, with distinctive large glands eat its apex 

The type of this species was collected by Benedict Balan- 
sa (no. 3807) -long the sides of the road leading from Tu- 
chep to the rocks of Notre Dame, Tonkin, French Indo-china, 
in May or June, 1887, and is deposited in the herberium of 
the Jardin Botanique Principal at Leningrad. 


LANTANA PRUTILLA Moldenke, sp. nov. 

Frutex; ramis griseis obtuse tetragonis srarsissime 
strigilloso-pubescentibus; ramulis sarmentisque pergracili- 
bus elongatis brunneis sparse strigillosis; foliis. opposit- 
is; petiolis gracilibus sparse albido-strigillosis; laminis 
membranaceis ovetis acutis vel subacuminatis, ad basin rot- 
undatis vel abrupte acuminatis, regulariter serratis, supra 
' sparse strigillosis, subtus dense breviterque pubescentibus; 
inflorescentiis axillaribus; capitulis numerosis. 

Shrub, to 1.5 m. tall; branches woody, gray, obtusely 
tetragonal, very sparsely strigillose-pubescent with scat- 


400 PHYTOLOGIA No. 12 


tered white hairs; brenchlets and twigs very slender, elong- 
ate, brownish, submargined on the obtuse angles, sparsely 
strigillose like the branches; leaves decussate-opposite; 
petioles slender, 3--5 mm. long, sparsely whitish-strigill- 
ose; blades membranous, bright-green above, somewhat lighter 
beneath, ovate, 1.8--6 cm. long, 1--3 cm. wide, acute or sub 
acuminate at apex, rounded at base or abruptly acuminate and 
prolonged into the petiole, uniformly serrate from the wid- 
est part to the epex with rather appressed acute or bluntish 
teeth, sparsely strigillose above, densely short-pubescent 
beneath; inflorescence axillary; heads solitary in the axils 
and opposite, numerous, 1--2 cm. long, to 1 cm. wide, dense- 
ly many-flowered; corolla-tube about 4 mm. long, its limb a- 
bout 3 mm. in diameter. 

The type of this species was collected by George B. Hint- 
on (no. 6455) in I. R. F. Cutzemala, at Pungarabato, Coyuca, 
Guerrero, Mexico, on August 17, 1934, and is deposited in 
the herbarium of the Royal Botanic Gardens at Kew. The spec- 
ies is also known from-Michoacén, Morelos, Hidalgo, Chiapas, 
and the state of México. 


LANTANA GROSSESERRATA Moldenke, sp. nove 

Frutex; ramulis acute tetragonis sparsiuscule adpresso~ 
strigosis; foliis oppositis; petiolis brevissimis substrigo- 
s0-pubescentibus; laminis chartaceis ellipticis ad basin et 
apicem acuminetis, supra mediam grosse serratis, supra spar- 
sissime brevissimeque pilosulis, subtus dense rebinoso- 
punctatis in venatio sparsissime brevissimeque pilosulis; 
inflorescentiis axillaribus longe pedunculatis cepitatis; 
bracteolis perspicuis numerosis elongatis anguste ellipti- 
cise 

Straggling shrub, to 2.5 m. tall; branchlets slender, a- 
cutely tetragonal, rather sparsely or densely appressed- 
strigose with yellowish hairs; nodes rather-obscurely annul-~ 
ete; principal internodes 5--9 cm. long; leaves decussate- 
opposite; petioles very much abbreviated, 5--7 mm. long, 
densely short-pubescent with substrigose hairs; blades char- 
taceous, slightly lighter beneath, elliptic, 7--12.5 cm. 
long, 2.8--4.8 cm. wide, attenuate-acuminate at base and a- 
pex, coarsely and rather irregularly serrate-dentate from a- 
bout the middle to the apex with sharply acute broadly tri- 
angular and divergent teeth, very sparsely and minutely 
pilosulous above, densely resinous-punctate beneath and also 
very sparsely short-pilosulous on the venation beneath; mid- 
rib slender, prominulous above, prominent beneath; secondar- 
ies slender, 5 or 6 per side, arcuate-ascending, mostly 
plane above, prominent beneath, not plainly anastomosing; 
veinlet reticulation obscure or indiscernible above, plane 
and not very conspicuous beneath; inflorescence axillary, 


OE ee F 


1940 Moldenke, Novelties 421 


long-pedunculate, capitate; peduncles slender, 6.5--7.5 cm. 
long, rather sparsely strigose; bractlets numerous, conspic- 
uous, narrowly elliptic, elongate, 6--20 mm. long, 2--4 mm. 
wide, densely resinous-punctate, venose, appressed-strigill- 
ose On the venation, acute at apex; corolla white, its tube 
about 6 mm. long, its limb about 2 mm. wide; drupes black. 

The type of this distinct species was collected by Alex- 
ander Frank Skutch (no. 4128) in a bushy clearing, at an al- 
titude of 730 m., in the vicinity of El General, San José, 
Coste Rica, in February, 1939, and is deposited in the Brit- 
ton Herbarium at the New York Botanical Garden. 


LANTANA MEARNSII Moldenke, sp. nov. 

Frutex; remis gracilibus obtuse tetragonis densiuscule 
hirsutulo-pubescentibus, in senectute scabridis; foliis 
ternatis; petiolis plusminus marginatis dense breviterque 
subadpresso-pubescentibus; laminis lanceolatis vel ellipti- 
cis acutis regulariter crenatis, ad basin acutis vel acumin- 
atis, supra dense puberulis scabris saepe bullatis, subtus 
densissime canescento-puberulis. 

Shrub; stems slender, obtusely tetragonal, rather densely 
hirsutulous-pubescent, scabridous in age; branches similar, 
more slender, more densely hirsutulous-pubescent; nodes not 
annulate; principal internodes 1.5--7.5 cme long; leaves 
ternate; petioles 3--8 cm. long, more or less margined, den- 
sely short-pubescent with mostly subappressed hairs; blades 
thin-chartaceous, dark-green above (brunnescent in drying), 
much lighter beneath, lanceolate or elliptic, 2.5--/.5 cm. 
long, 1.65--3-5 cm. wide, acute at avex, acute or acuminate 
at base, regularly crenulete with blunt antrorse teeth ex- 
cept at the very base, densely puberulent and scabrous ab- 
ove, often more or less bullate, very densely canescent-- 
puberulent; midrib, secondaries, and veinlet reticulation 
impressed above, often deeply so, prominulous beneath; in- 
florescence spicate, subcapitate when young, later elongat- 
ing, opposite or ternate, long-pedunculate; peduncles slend- 
er, 1.5--8.5 cm. long, densely puberulent; spikes 1.5--2 cm. 
wide, elongated to 1.5 ome in age; outer bractlets large and 
foliaceous, broadly elliptic or ovate, 7--14 mm. long, 4--9 
mme wide, acute, entire, venose, densely puberulent; corolla 
about 1 cm. long, its limb ebout 5 mm. in diameter. 

The type of this species was collected by Edgar Alexander 
Mearns on the Smithsonian Institution Expedition under the 
direction of Colonel Theodore Roosevelt in the vicinity of 
Fort Hall, at an altitude of about 1200 m., Kenya, between 
September 10 and 12, 1909, and is’ deposited in the Britton 
Herbarium at the New York Botanical Garden. The species has 
been widely confused in literature and herbaria with the 
South African L. salvifolia Jacq. and the tropical American 


400 PHYTOLOGIA No. 12 
L. trifolia Le, from both of which it is quite distinct. 


LANTANA NOTHA Moldenke, Spe NOVe 

Frutex; ramulis dense albido-puberulis et plusminus dense 
albido-hirsutis inermibus vel minute sparsissimeque aculeol- 
atis; petiolis pergracilibus dense puberulis et hirsutis; 
leminis chartaceis ovatis acutis grosse serratis, ad basin 
subtruncetis vel abrupte acutis, utrinque dense cinereo- 
pubescentibus; inflorescentiis axillaribus capitatis; bract- 
eolis lanceolatis vel oblongis dense cinereo-puberulis. 

Shrub; branchlets slender, obtusely tetragonal, densely 
white-puberulent end also more or less densely white-hirsute 
with hairs 2 or 3 times as long, more densely so toward the 
apex and on.younger parts, unarmed or armed with a very few 
minute prickles; nodes usually annulate with a band of dens- 
er hirsute hairs; principal internodes 4--7.5 cm. long; 
leaves decussate-opposite; petioles very slender, 3--12 m. 
long, very densely white-puberulent and hirsute; blades thin 
-chartaceous, uniform'y gray-green on both surfaces, ovate, 
1.6--5 em. long, 1--2 cm. wide, acute et apex, coarsely ser- 
rate from base to apex with broadly triangular subacute 
teeth, subtruncate at base (or abruptly acute when imnat- 
ure), somewhat prolonged into the petiole at the middle, 
densely cinereous-pubescent on both surfaces, the hair sub- 
atrigose above; midrib slender, subimpressed and more dense- 
ly pubescent above, prominulous beneath; secondaries very 
slender, 4--6 per side, short, ascending, not much arcuate, 
subimpressed and not pubescent above, prominulous beneath; 
veinlet reticulation abundant, the lerger portions subim- 
pressed and not pubescent above and subprominulous beneath; 
inflorescence axillary, cavitate, surpassing the subtending 
leaves; peduncles slender, 3--/7 cm. long, grayish-puberulent 
and also more or less sparsely white-hirsute; heads 1.2--2.35 
cme wide, many-flowered; bractlets lanceolate or oblong, 5-- 
6 mm. long, 1--2 mm. wide, blunt or subacute at apex, dense- 
ly cinereous-puberulent; corolle-tube 7--8 mm. long, its 
limb about 5 mm. wide. 

The type of this species was collected by Joseph Nelson 
Rose, Paul Carpenter Standley, and Paul George Russell (no. 
13573) along the river in the vicinity of Fuerete, Sinaloa, 
Mexico, on March 27, 1910, and is deposited in the Britton 
Herbarium et the New York Botanical Garden. 


LANTANA SCABRIFOLIA Moldenke, sp. nov. 

Herba sublignosa; caulibus obtuse tetragonis, in juventu- 
te adpresso-strigosis, in senectute minute asperulis vel 
glebratis; foliis oppositis breviter petiolatis; laminis 
firme chertaceis ovatis obtusis regulariter crenulatis, ad 
basin rotundatis vel subtruncatis, supra scaberrimis, sub- 


1940 Moldenke, Novelties 4o3 


tus dense puberulis; inflorescentiis axillaribus longe ped- 
unculatis; spicis subcapitatis oblongis dense multifloris 
valde bracteolatis. 

Probably a rather woody herb; stems apparently not much 
branched or simple, obtusely tetragonal, at first densely 
appressed-strigose, later minutely asperulous, finally glab- 
rate; nodes annulate; principal internodes 4.5--7 cm. long; 
leaves opposite, usually with several smaller ones on great- 
ly abbreviated twigs in their axils; petioles very slender, 
5--7 mm. long, sparsely puberulent and asperulous, canalicu- 
late above; blades firmly thick-chartaceous, rather uniform- 
ly gray-green on both surfaces, ovate, 2--7.5 cm. long, 1.4 
-3./ cme wide, obtuse at apex, regularly crenulate from apex 
to base, rounded or subtruncate at base, very scabrous ab- 
ove, densely puberulent and slightly scabridous beneath; 
midrib, secondaries, and veinlet reticulation impressed a- 
bove and prominent beneath; inflorescences axillary, paired 
in each axil near the tip of the stem; peduncles very slend- 
er, 2.5--4.5 cm. long, densely appressed-puberulent, ascend- 
ing; spikes subcapitate, becoming oblong and to about 1 cm. 
long, 8--10 mm. wide, densely many-flowered; bractlets lan- 
ceolate-ovete, conspicuous, the lowermost ones to 6 mm. long 
and 2 mme wide, long-attenuate to the apex, densely puberul- 
ent; flowers equaling or barely exceeding the bractlets; 
corolla-limb about 2 mm. in diameter. 

The type of this species was collected by Edgar Alexender 
Mearns (no. 267) on the Smithsonian African Expedition under 
the direction of Colonel Theodore Roosevelt in the vicinity 
of Nairobi on the Uganda Railway, at an altitude of about 
2000 m., Kenya, between May 26 and June 1, 1909, and is dep- 
osited in the Britton Herbarium at the New York Botanical 
Gardene The species has hitherto been confused with L. salv- 
ifolia Jacq., of South Africa, which has much smsller leaves 
and larger flowers. 


LIPPIA CONTROVERSA Moldenke, sp. nov. 

Frutex; ramulis gracilibus sparse hirsutis glabrescentib- 
us; foliis crasse chartaceis ovatis vel ovato-ellipticis a- 
cutis vel breviter acuminetis regulariter serrulatis, ad 
basin rotundetis, supra bulboso-pilosis scaberrimis, subtus 
sparsissime hirsutis; inflorescentiis axillaribus, peduncul- 
is solitariis vel geminatis simplicibus; capitulis valde 
brecteatis. 

Shrub, to 2 m. tall; branchlets slender, stramineous, ob- 
tusely tetragonal, sparsely hirsute, becoming glabrate in 
age; nodes flattened, annulate; principal internodes 5--9 
em. long; leaves decussate-opposite; petioles rather stout- 
ish, short, 5--10 ma. long, sparsely hirsute, more or less 
_margined; bledes thick-chartaceous, uniformly gray-zreen on 


kok PHYTOLOGIA No. 12 


both surfaces, ovate or ovate-elliptic, 3.5--7 cme long, 
1.5--4.5 cme wide, acute or short-acuminate at apex, regu- 
larly serrulate from base to apex, rounded at base and more 
or less attenuate into the petiole at the center, acute when 
immature, densely and harshly pilose above with bulbous- 
based hairs and very rough-scabrous, very sparsely hirsute 
or almost subglabrous beneath; midrib slender, plane or im- 
pressed above, prominent beneath; secondaries slender, 7 or 
8 per side, arcuate-ascending, mostly impressed above, prom- 
inent beneath; veinlet reticulation abundant, mostly impres- 
sed above, plane or the larger portions subprominulous be- 
neath; inflorescence axillary, simple, capitate; peduncles 
very slender, one or two from each of the upper leaf-axils, 
2.5--4 om. long, densely short-pubescent or puberulent with 
glandular hairs; heads 1.5--2.5 cme wide; bracts very large 
and conspicuous, numerous, ovate, the lowermost about 10 
mn. long and 6 mm. wide, acute, densely puberulent; corolla 
cream-colored or pale greenish-yellowe 

The type of this species was collected by Paul Carpenter 
Standley (no. 58067) on a brushy slope near Las Lajas, Saca- 
tepequez, at an altitude of about 1200 m., Guatemala, on 
November 28, 1938, and is deposited in the Britton Herbarium 
at the New York Botanical Garden. The species is also known 
from Escuintla. 


LIPPIA COSTARICENSIS Moldenke, sp. nove 

Arbor; ramulis acute tetragonis saepe submarginsatis plus- 
minus substrigoso-pilosis; foliis oppositis; petiolis brevi- 
bus plusminus adpresso-strigosis; laminis chartaceis anguste 
ellipticis serrulatis, ad basin et apicem attenuatis acutis 
vel acuminatis, supre minute strigillosis glabrescentibus, 
subtus minute puberulis; inflorescentiis axillaribus psnicu- 
latis. 

Tree, to 18 m. tall; branchlets stoutish or rather slend- 
er, acutely tetragonal, often submargined, more or less sub- 
strigose-pilose; nodes plainly annulate, often conspicuously 
80; principal internodes 2--4.5 cm. long; leaves decussate- 
opposite; petioles slender, 1--2 cm. long, usually short, 
more or less appressed-strigose; bledes chartaceous, rather 
uniformly green on both surfaces, narrowly elliptic, 8--18 
cme long, 1.5--5 cm. wide, attenuate and acute or acuminste 
at base and apex, regularly serrulate from base to apex with 
_ very fine rounded and appressed teeth, often subrevolute a- 
long the margins, minutely strigillose above, glabrescent 
end merely punctate in age, minutely puberulent beneath, 
often only on the venation; midrib rather stout, plene a- 
bove, prominent beneath; secondaries slender, 8--10 per 
side, plane ebove, prominent beneath; veinlet reticulation 
abundant, plane (often obscure) above, the larger portions 


——————— 


1940 Moldenke, Novelties 425 


subprominulous beneath; inflorescence axillary, paniculate, 
with 1--3 whorls of capitate branches subtended by more or 
less foliaceous bracts; heads 5--10 mm. wide, conspicuously 
bracteolate; bractlets large, ovate, stramineous, the lower- 
most ebout 6 mm. long and 4 mm. wide, minutely puberulent; 
corolla yellow. 

The type of this species was collected by Alexander Frank 
Skutch (no. 2292) at an altitude of 915 m., in the vicinity 
of El General, San José, Costa Rica, in December, 1935, and 
is deposited in the Britton Herbarium at the New York Botan- 
ical Garden. The species is also known from Alajuela. 


LIPPIA CURTISIANA Moldenke, sp. nov. 

Frutex; ramis ramulisque gracilibus dense breviterque 
pubescentibus, pilis sordidis; foliis oppositis vel ternat- 
is; petiolis gracilibus brevibus dense breviterque pubes- 
centibus; laminis chartaceis saepe falcatis ovatis acutis 
regulariter serrulatis, ad basin abrupte acutis, supra stri- 
gosis scebris bullatis, subtus breviter pubescentibus; in- 
florescentiis axillaribus capitatis simplicibus ebbreviatis 
confertis; bracteolis maturis reniformibus venosis magnis 
densis. 

Shrub; branches and branchlets slender, grey, obtusely 
tetragonal, densely short-pubescent with sordid appressed 
hairs, glabrescent in age; nodes not annulate; principal in- 
ternodes 0.5--3 cm. long; leaves decussate-opposite or tern- 
ate; petioles slender, 4-7 mm. long, densely short-pubes- 
cent with appressed sordid hairs; blades chartaceous, often 
falcate and conduplicate in drying, rather uniformly gray- 
ish-green on both surfaces or slightly lighter beneath, o- 
vate, 1.5--5.5 cm. long, 0.2--3.4 cm. wide, acute at apex, 
regulerly serrulate with small blunt appressed or spreading 
teeth from almost the base to the apex and usually slightly 
revolute, abruptly acute or rounded at base, strigose and 
scabrous-bullate above, short-pubescent beneath (especially 
along the venation); midrib very slender, subimpressed a- 
bove, prominulous beneath; secondaries very slender, 4--8 
per side, ascending, not much arcuate, impressed above, pro- 
minulous beneath; vein and veinlet reticulation abundant, 
more or less impressed above and often subprominulous be- 
neath; inflorescence axillary, capitate, simple, abbreviat- 
ed; peduncles very slender, usually 3 per axil, O./--2 cm. 
long, densely appressed-pubescent with sordid-gray hairs; 
heads to 9 mm. wide in anthesis, to 13 mm. wide in fruit; 
brectlets ovate, the lowermost to 4 m. long and 3 mm. wide, 
densely appressed=puberulent, in fruit greatly expanded and 
reniform, venose, to 9 mm. wide, stramineous. 

The type of this species was collected by Edward Palmer 
(no. 479) at the city of Durango and vicinity, at an alti- 


426 PHY TOLO.GI-A No. 12 


tude of 6207 feet, Durango, Mexico, between April and Novem- 
ber, 1896, and is deposited in the Britton Herbarium at the 
New York Botanical Garden. The species is named in honor and 
grateful appreciation of Dr. Carleton Clarence Ourtis, for 
many years professor of botany at Columbia University, auth- 
or of valuable texts, inspiring teacher, and valued friend 
and adviser of hundreds of the present generation of Ameri- 
can botanists, who will always honor and revere his memory. 


LIPPIA DURANGENSIS Moldenke, sp. nove 

Fruticulus; ramulis gracilibus dense adpresso-pubescent- 
ibus; foliis oppositis vel ternatis vel quaternatis; petiol- 
is gracillimis dense puberulis et resinoso-glandulosis; lam- 
inis tenuiter chartaceis ellipticis saepe subfalcatis acutis 
reguleriter serratis, ad basin acutis vel acuminatis, supra 
scaberrimis, subtus densiuscule puberulis et resinoso- 
punctatis; inflorescentiis axillaribus capitatis confertis 
simplicibus. 

Shrub; branchlets slender, obtusely tetragonal, brownish, 
densely appressei-pubescent; nodes annulate; principal in- 
ternodes 3--7 cm. long; leaves opposite, ternate, or wnorled 
in 4's; petioles very slender, 5--7 mm. long, densely puber- 
ulent end resinous-glandular, less so in age; blades thin- 
chertaceous, dark-green above, much lighter beneath, ellip- 
tic, often falcate and more or less conduplicate in press- 
ing, 4--6 cme long, 1.3--3 cm. wide, acute at apex, regular- 
ly serrate from almost the base to the apex with rather 
small rounded and often appressed teeth, acute or acuminate 
at base, very rough-scabrous above, rather densely puberu- 
lent and resinous-punctate beneath; midrib slender, plane or 
subimpressed above, prominulous beneath; secondaries very 
slender, 5--8 per side, arcuate-ascending, subimpressed a- 
bove, promimulous beneath; veinlet reticulation abundant, 
subimpressed above, the larger portions prominulous beneath; 
inflorescence axillary, capitate, simple, crowded; peduncles 
very slender, 1--5.5 cme long, 2--5 or more in each axil, 
more or less densely short-pubescent with substrigose whit- 
ish hairs, often also resinous-punctate; heads 6--9 mm. in 
diameter; bracts large and conspicuous, ovate, very plainly 
venose with a conspicuous reticulum, the lowest about 5 mm. 
long and wide, acute at apex, puberulent-margined; corolla 
greenish-yellow. 

The type of this species was collected by Francis Whit- 
tier Pennell (no. 18189) on a rocky andesitic slope, at an 
altitude of 1900--2000 m., at Chupaderos, north of Durango, 
in the Sierra Madre Occidental, Durango, Mexico, on August 
24, 1934, and is deposited in the United States National 
Herbarium at Washington. The beautifully venose bracts are 
quite characteristic. 


1940 Moldenke, Novelties 4o7 


LIPPIA INDICA Moldenke, sp. nov. 

Herba perennis; ceulibus ramulisque gracilibus obtuse 
tetragonis dense hirsutulis vel breviter pubescentibus vel 
incano-puberulentis; nodis annulatis; foliis oppositis; pet- 
iolis gracilibus albido-hirsutis; laminis chertaceis ovatis 
obtusis serratis, supra bullatis densiuscule strigosis, sub- 
tus dense tomentulosis; inflorescentiis axillaribus spicat- 
is; spicis elongatis. 

Perennial herb; stems and branches slender, obtusely tet- 
ragonal, densely hirsutulous or short-pubescent, becoming 
more sparsely so or merely puberulent in age, the hairs 
whitish; nodes annulate; leaves decussate, often with fas- 
cicles of small ones on greatly abbreviated twigs in their 
axils; petioles slender, 3--15 mm. long, very densely or 
sparsely hirsute with whitish hairs; blades chartaceous, 
grey-green, ovate, 0.8--7.5 om. long, 0.6--3.5 cm. wide, ob- 
tuse at apex, regularly serrate from base to apex with ex- 
tremely small and blunt somewhat appressed teeth, very bull- 
ate and rether densely strigose with appressed whitish heirs 
above, densely tomentulose beneath; inflorescence axillary, 
clustered toward the tips of the branches, spicate; spikes 
elongate, 2--5.5 cm. long, 1--45 in each upper axil, erect or 
ascending, the floriferous part 0.4--2.6 cm. long, uniformly 
5--6 mm. wide throughout; peduncles very slender, 1.5--2.6 
cme long, densely short-pubescent or puberulent with whitish 
hairs, often binary. 

The type of this species was collected by James Sykes 
Gamble (no. 17895) at Sigte Ghat, at an altitude of 3000 
feet, Nilgiri district, Madras, India, in August, 1886, and 
is deposited in the herbarium of the Royal Botanic Gardens 
at Kew. The species is also known from Coimbatore. 


LIPPIA JALISCANA Moldenke, spe nov. 

Frutex vel arbor; ramis ramulisque gracilibus brunneis 
tetragonis breviter pilosis velde scabris; foliis oppositis; 
petiolis gracilibus brevibus breviter pilosis scabris; lam- 
inis coriaceis lenceolatis acutis reguleariter serrulatis, ad 
basin acutis vel acuminatis, supra bullatis scaberrimis, 
subtus puberulis dense glanduloso-punctatis; inflorescentiis 
axillaribus paniculatis vel raro simplicibus capitatis. 

Shrub or small tree; branches and branchlets slender, 
brown, tetragonal, shortly appressed-pilose, decidedly 
scabrous, especially on the margins; nodes annulate; princi- 
pal internodes 1.5--6 cm. long; leaves decussate-opposite; 
petioles very slender, 4--17 mm. long, short-pilose and sca- 
brous; blades coriaceous, rather uniformly green on both 
surfaces or slightly lighter beneath, lanceolate, 4-11 cm. 
long, 1.7--3.2 cm. wide, acute at apex, regularly serrulate 
from almost the base to the apex with blunt more or less ap- 


408 PHYTOLOGIA No. 12 


pressed revolute teeth, acute or acuminate at base, bullate 
and very scabrous above, puberulent and densely glandular- 
punctate beneath; midrib slender, plane or subimpressed a- 
bove, prominent beneath; secondaries very slender, 8--10 per 
side, arcuete-ascending, impressed above, sharply prominent 
beneath; vein and veinlet reticulation abundant, impressed 
above, sharply prominent beneath; inflorescence axillary, 
mostly paniculate with the heeds borne in secondary umbels 
of 3--6, rarely a few simple ones included; peduncles very 
slender, 2--4 cm. long, densely short-pubescent with spread- 
ing brownish hairs; secondary peduncles 1--1.5 cm. long; 
heads 6--10 mm. wide in anthesis, densely many-flowered; 
bractlets ovate, about 3 mm. long and wide, sharply acute at 
apex, densely puberulent; corolla greenish-yellow, exceeding 
the bractlets. 

The type of this species was collected by Ynes Mexia (no. 
1636) in an oak-clad open forest on steep slopes on the 
trail from Real Alto to San Sebastian, at an altitude of 
2000 m., in the Sierra Madre Occidental, Jalisco, Mexico, on 
Paheverny 3, 1927, and is deposited in the Britton Herbarium 
at the New York Botanical Garden. 


LIPPIA TRANSVALENSIS (Kuntze) Moldenke, comb. nove 
Camara salviaefoline transvalensis Kuntze, Rev. Gen. Pl. 
3, part 2: 250. 1898. 


LIPPIA WHYTEI Moldenke, sp. nov. 

Herba; remis obtuse tetragonis brunneis dense breviterque 
pubescentibus vel puberulis; nodis valde annulatis; foliis 
oppositis breviter petiolatis; leminis lanceolatis acutis 
vel breviter acuminatis, ad basin acutis vel acuminatis, 
crenulatis, supra bulletis sceberrimis, subtus tomentellis; 
inflorescentiis axillaribus sessilibus vel subsessilibus ge- 
minatis; spicis subcapitatis dense multifloris. 

Herb; stems and branches slender, obtusely tetragonal, 
brownish, the younger parts densely short-pubescent with 
sordid hairs, the older parts puberulent with minute brown- 
ish hairs; nodes plainly ennulate; principal internodes 1.5 
--4.5 cme long; leaves opposite; petioles 2--5 mm. long, 
alate-margined, sparsely short-pubescent; blades firmly 
chartaceous, dark-green above, lighter beneath, lanceolate 
or elliptic, 2.5--4 cm. long, 8--18 mm. wide, acute or 
short-acuminate at apex, regularly crenulate from base to a~ 
pex, acute or acuminate at base, bullate and very scabrous 
above, densely tomentellous with sordid-grayish hairs be- 
neath; midrib, secondaries, and veinlet reticulation deeply 
impressed above, only the larger portions (midrib and sec- 
ondaries) prominulous beneath; inflorescence axillary, abun- 
dent toward the tips of the branches, a pair in each axil; 


1940 Moldenke, Novelties 409 


spikes sessile or subsessile, subcapitate, to about 12 mm. 
long and 10 m. wide, very densely many-flowered; bractlets 
ovate, about 2 mm. long, sharply acute or short-acuminate, 
strigose with short sericeous hairs; flowers very small, 
hardly exceeding the bractlets. 

The type of this species was collected by Alexander Whyte 
in northern Nyasaland, and is deposited in the Britton Herb- 
arium at the “ew York Botanical Garden. It has hitherto been 
confused with Lantana salvifolia Jacq., of South Africa, 
with long-pedunculate heads and fleshy fruits. 


PRIVA PEDICELLATA Moldenke, Geogr. Distrib. 32, nom. nud. 
(1939), sp. nov. 

Herba; caulibus ramisque gracilibus argute tetragonis 
breviter pubescento-strigillosis vel puberulis; foliis opp- 
Ositis; petiolis gracillimis sparsiuscule breviterque pubes- 
cento-strigillosis; laminis submembranaceis ovatis acutis 
vel obtusis serratis, ad basin subtruncatis vel subcordatis 
vel abrupte acutis, utrinque sparsiuscule strigillosis vel 
albido-strigosis; inflorescentiis terminalibus racemiformi- 
bus erectis laxe multifloris; pedicellis gracillimis elonga- 
tis. 

Herb, to 5 dm. tall; stems and branches slender, sharply 
acute, shortly pubescent-strigillose, becoming sparsely so 
or merely puberulent in age, the older parts gray, the 
younger ones turning brown in drying; nodes annulate on 
young parts, indistinctly so or not annulate on older parts, 
not swollen; principal internodes 1.3--7.5 cm. long; leaves 
decussate-opposite; petioles very slender, 5--11 mm. long, 
rather sparsely short-pubescent or strigillose, flattened a- 
bove, convex beneath, not noticeably ampliate at base; 
blades submembranous, rather uniformly bright-green on both 
surfaces or slightly lighter beneath, ovate, 2--3.8 cm. 
long, 1--2.5 cm. wide, acute or obtuse at apex (in outline), 
varying from subtruncate or subcordate to abruptly acute at 
base, usually somewhat prolonged into the apex of the peti- 
ole at the center, rather coarsely serrate from near the 
base to the apex with broad and acute antrorse teeth, rather 
sparsely strigillose or strigose with distant appressed non- 
bulbous hairs on both surfaces; midrib very slender, plane 
above, slightly subprominulous beneath; secondaries very 
slender, 3 or 4 per side, ascending, not much arcuate, plane 
on both surfaces, not conspicuously anastomosing; vein and 
veinlet reticulation very delicate, obscure or indiscernible 
above, plane beneath; inflorescence terminal, racemiform, 7 
--17 cm. long, 0.5--2 cm. wide in anthesis, loosely many- 
flowered, erect; peduncles (2.5--3.8 cm. long) and rachis 
exactly similar to the adjacent branches or stems in color, 
shape, texture, and pubescence; pedicels very slender, elon- 


430 | PHYTOLOGIA No. 12 


gate, 5--7 mm. long in anthesis, rather sparsely short- 
pubescent or puberulent; prophylla about 2 mm. long, lanceo- 
late-ovate, attenuate-acuminate at apex, sessile, puberul- 
ent; fruit not seen. 

The type of this remarkable species was collected by A. 
E. English somewhere in Burma and is deposited in the B. H. 
Lace Herbarium, purchased in 1918 by the Royal Botanic Gard- 
en at Edinburgh. 


SPARTOTHAMNELLA TEUCRIIFLORA (F. Muell.) Moldenke, Geogr. 
Distrih. 34, nom. nud. (1939); Prelim. Alph. List In- 
valid Name 40, hyponym (1940), comb. nov. 

Spartothamus teucriiflorus F. Muell. in Wing, South. Se. 

Record 2: 55. 1882. 


STACHYTARPHETA FRANTZII var. PATENTIFLORA Moldenke, var.nov. 

Haec varietas a forma typica speciei recedit calycibus 
rhachidem non arcte adpressis et pilis ubique sparsioris. 

This variety differs from the typical form of the species 
in having its rachis more slender and often slightly flexu- 
ous, the bractlets and calyxes not closely appressed to and 
sunken in the rachis, but more or less spreading before, 
during, and after anthesis, and the pubescence throughout 
(especially on the rachis, bractlets, and calyx) more 
sparse. 

The type of this variety was collected by Adolfe Tonduz 
(no. 626) in broken ground? near Guatemala City, Guatemala, 
at an altitude of 1400 m., in July, 1921, and is deposited 
in the Britton Herbarium at the New York Botanical Garden. 


STACHYTARFHETA GUATEMALENSIS var. LUNDELLIANA Moldenke, var. 
nove 

Haec varietas a forma typica speciei recedit ramulis fol- 
iisque spicisque plusminus adpresso~pilosis et laminis foli- 
orum majoribus crassiore serratis. 

This variety differs from the typical form of the species 
in having the branchlets, petioles, leaf-blades, peduncles, 
rachis, bractlets, and calyxes more or less appressed-pilose 
with very short and sparse whitish hairs, and in having the 
leaf-blades larger, to 8 cm. long and 4 cm. wide, more att- 
enueate and sharply acute or subacuminate at apex, and much 
more coarsely serrate with broadly triangular and spreading 
sharply acute or apiculate teeth. 

The type of this variety was collected by Eizi Matuda 
(no. 1762) at Finca Fuarez, Chiapas, Mexico, on August 12, 
1937, and is deposited in the Britton Herbarium at the New 
York Botanical Garden. It is named in honor of my esteemed 
friend and colleague, Dr. Cyrus Longworth Lundell of the U- 
niversity of Michigan, who is doing such important and note- 


1940 Moldenke, Novelties 43) 
worthy work on the flora of Mexico and Central America. 


STACHYTARPHETA HINTONI Moldenke, sp. nov. 

Herba perennis (7); ramis gracilibus dense breviterque 
pubescentibus vel puberulis subglabrescentibus; foliis oppo- 
sitis; petiolis gracilibus alatis dense vel sparse breviter- 
que pubescentibus; laminis membranaceis ellipticis acutis 
vel acuminatis serratis, ad basin longiuscule acuminatis, 
utrinque sparse strigillosis; rhachide valde excavato pubes- 
cente vel puberulo. 

Perennial herb (?); stems and branches slender, buff- 
colored, rather densely short-pubescent when young, becoming 
merely puberulent or even subglabrate in age; leaves decuss- 
ate-opposite; petioles slender, 8--10 mm. long, alate- 
margined, densely or sparsely short-pubescent; blades memb- 
ranous, very brittle and:fragile in drying, elliptic, 5.5-- 
9.5 cm. long, 3--5 cm. wide, acute or acuminate at apex, 
rather long-acuminate at base and prolonged into the petiole 
at the center, regularly or irregularly serrate (except on 
the basal and apical ecuminations) with coarse acute antror- 
se somewhat divergent teeth, sparsely strigillose on the 
lamina above with short appressed antrorse whitish hairs, 
similarly strigillose on the venation beneath; inflorescence 
terminal, spicate, 12--27 ome long, densely many-flowered, 
nutant before anthesis, erect in fruit, the flowers closely 
appressed and imbricate, sunken into the deeply excavated 
short=-pubescent or puberulent rachis; bractlets lanceolate- 
ovate, 8--10 mm. long, to 2.5 mm. wide at base, sharply att- 
enuate at apex, sparsely puberulent. 

The type of this species was collected by George B. Hint- 
on (no. 8236) -- in whose honor it is named -- in a shaded 
barranca at Luvianos, Temascaltepec, Mexico, Mexico, on Sep- 
tember 29, 1935, and is deposited in the herbarium of the 
Royal Botanic Gardens at Kew. 


STACHYTARPHETA INCANA Moldenke, Prelim. Alph. List Invalid 
Names 46, hyponym (1940), sp. nov. 

Fruticulus; ramulis gracilibus tetragonis dense incano- 
strizgillosis; foliis oppositis sessilibus; laminis tenuiter 
chartaceis ovato-ellipticis vel ellipticis acutis vel suba- 
cuminatis arrute serratis, ad basin longe caudatis, supra 
sparsiuscule strigillosis, subtus densissime puberulento- 
strigillosis vel subvelutinis; rhachide valde excavato den- 
sissime incano-strigilloso. 

Shrubby; branchlets rather slender, gray-brown, rather 
plainly tetragonal, densely incanous-strigillose with very 
short antrorsely appressed hairs; nodes not distinctly ann- 
ulate; principal internodes 1.5--5 cm. long; leaves decuss- 
‘ate-opposite, regularly disposed to the base of the spikes, 


432 PHYTOLOGIA No. 12 


sessile; blades very thin-chartaceous, dark-green on both 
surfaces, brunnescent in drying, ovate-elliptic or elliptic, 
4,5--8.5 com. long, 1.7--3-7 cm. wide, sharply acute or suba- 
cuminate at apex, long-caudate at base, sharply serrate with 
antrorse teeth except on the basal prolongation and et the 
very apex, rather sparsely strigillose above, very densely 
puberulent-strigillose beneath (subvelutinous on the larger 
venation); midrib slender, plane above, subprominulous be- 
neath; secondaries slender, 3--5 per side, ercuate-ascend- 
ing; spikes terminal, 17--19 cm. long, about 5 mm. wide 
throughout exclusive of the corollas; peduncles 10--12 mm. 
long, incanous-strigillose; rachis very densely incanous- 
strigillose, deeply excavated; bractlets broadly ovate, ab- 
out 5 mm. long and 35 mm. wide at base, abruptly acuminate 
on the upper 1/3, incanous-strigillose and ciliate, but not 
as densely strigillose as the rachis. 

The type of this species was collected by Martin Sesse, 
José Mariano Mocifio, Juan Diego del Castillo, and José Mald- 
onado (no. 137) somewhere in Mexico between 1787 and 1804, 
and is deposited in the herbarium of the Jardin Botsnico at 
Madrid. The species is known otherwise only from Guerrero. 


STACHYTARPHETA INCANA var. ANGUSTIBRACTEATA Moldenke, Pre- 
lim. Alph. Lis Invalid Names 44, hyponym (1940), var. 
noVe 

Haec varietas a forma typica speciei recedit spicis elon- 
gatis angustioribus et bracteolis lanceoletis, ad basim 1 
mme latis, ad apicem longissime acuminato-ceudatis. 

This variety differs from the typical form of the species 
in its somewhat more slender and more elongated spikes, 
which are 28--30 cm. long, and its lanceolate bractlets, on- 
ly 1 mm. wide at base end very long-acuminate-caudate at the 
aPeXe 

The type was collected by Sessé, Mocino, Castillo, & Mal- 
donado (no. 136) somewhere in Mexico between 1787 and 1804, 
and is deposited in the herbarium of the Jardin Botanico at 
Madrid. The variety is otherwise known from San luis Fotosf, 
Neyerit, Sinaloa, Veracruz, and Oaxaca. 


VERBENA LITORALIS var. ALBIFLORA Moldenke, var. nov. 

Haec varietas a forma typica speciei recedit corollas 
albis. 

This variety differs from the typical form of the species 
in its white corollas. 

The type was collected by George B. Hinton (no. 13,965) 
in a llano at an altitude of 1000 m., Coalcoman, district of 
Coalcoman, Michoacan, Mexico, on July 20, 1939, and is dep- 
osited in the herbarium of the University of California at 
Los Angeles. 


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PHY TOLOGIA 


Designed to expedite botanical publication 


Vor. 1 October, 1940 No. 13 
CONTENTS 

L. Crowat: A New Series of Croton FON COLE DIA iin nee BNW 433 

H.N. Motpvenxe: Additional Verbenaceous Novelties. ....e.00e- 433 


Published by H. A. Gleason and Harold N. Moldenke 
The New York Botanical Garden 
Bronx Park, New York, N. Y. 


Price of this number, 25 cents; per volume, $5.00 in advance 


wa 


A NEW SPECIES OF CROTON FROM COLOMBIA 


Leon Croizat 


CROTON CUPREATUS Croiz., sp. nov. 

Planta lignosa, frutex vel arbor, innovationibus furfura- 
ceo-lepidotis, pube nulla. Foliis ovato-ellipticis, apice 
brevius acuminatis, basi levissime cordatis ad rotundatis, 
margine integris, supra dissite subargenteo-lepidotis, caet- 
erum olivaceis, subtus lepidibus confertioribus nitore plum- 
beo, vel cupreo-rubiginosis, costa venisque rubiginosis, 
venis optime penninerviis ad 10-jugis, vix anastomosatis 8-- 
9 cm. longis, 3--4 cm. latis; petiolo lepidoto, ad 6 om. 
longo, glandulis 2 bacilliformibus apice dilatatis insigni- 
to. Cymis ad 15 om. longis, utriusque sexus, floribus totis 
capitulatis, capitulis inferioribus 2-sexualibus, superiori- 
bus more generis masculis: flore foemineo pedicello crassi- 
usculo ad 8 m. longo fulto, confertissime furfuraceo- 
lepidoto, calyce ad 4 mm. lato, lepidoto, lobis triangulari- 
bus ad 2 mm. longis, petalis ligulatis vel ligulato- 
truncatis, vel ligulato-bicornutis, margine tanto puberulis; 
ovario toto aureo-lepidoto ca. 4 mm. magno, stylis 3, crass- 


_iusculis, apice bifidis, pro genere brevibus, nec ultra 1 


mm. longis, patentibus neque convolutis: flore masculo pedi- 
cello tenuiori ad.5 mm. longo fulto, ca. 5 mm. lato, pallide 
lepidoto, petalis integerrimis, glabris, ovatis ad 1.5 mm. 
longis, lobis triangularibus ca. 1.5--2 mm. longis; stamini- 
bus ad 12. 

Typus: COLOMBIA: sine loco, Quatrecasas 8667 in herb. 
Arnold Arboretum. 

C. cupreatus is a very distinct species, with a type of 
indument reminiscent of that of C. punctatus Jacqe, but al- 
together unlike this plant. I can not suggest at present the 
name of another American endemic which it approaches. It was 
collected and sent by Dr. J. Quatrecasas in August, 1940, 
without indication of precise locality. 


ADDITIONAL VERBENACEOUS NOVELTIES 
Harold N. Moldenke 
STACHYTARPHRBTA INDICA f. MONSTROSA Moldenke, Prelim. List 


Invalid Nemes 7, hg or (1940), f. nov. 
oy) 


43h PHYTOLOGIA No. 13 


Haec forma a forma typica speciei recedit spicis multo- 
ramosis et floribus abortis. 

This form differs from the typical form of the species in 
having its spikes modified into usually flowerless many- 
branched panicles, the individual branches of the panicle e- 
rect or ascending, very slender, 1--15 cm. long, issuing 
from the axils of the normal bractlets in the deep excava- 
tions of the normal rachis, or else even the normal rachis 
absent and replaced by a very slender one similar to the 
branches; bractlets numerous and often closely imbricate on 
the branches, subtending greatly aborted often long-pedicel- 
late flowers or no flowers at all. 

The type of this form was collected by George Thomson in 
Mysore or the Carnatic region of Madras, India, and is dep- 
osited in the herbarium of the Jardin Botanique Principal at 
Leningrad. This obviously abnormal and monstrous form has 
been found repeatedly in India and, more recently, in Sumat- 
rae Wallich regarded it as an abnormal form of Bouchea hyd- 
erobacensis Walp. [=Svensonia hyderobadensis (Walp. ) Molden- 
ke] and others have thought that it may represent a bigener- 


ic cross between Stachyterpheta indica (L.) Vahl and Svens- 
onia hyderobadensis (cfr. Moldenke in Fedde, Repert. qi: 


132. 1936]. 


LANTANA LANGLASSEI Moldenke, sp. nov. 

Frutex; ramulis gracillimis acutiuscule tetragonis sparse 
stricillosis; foliis oppositis; petiolis gracillimis sparse 
strigillosis; laminis submembransceis lanceolatis attenuatis 
subacuminatis, ad basin abrupte acutis vel breviter acumina- 
tis, regulariter serratis, supra sparse albido-strigillosis, 
subtus glabratis densiuscule nigro-punctatis. 

Low shrub; branchlets extremely slender, rather acutely 
tetragonal, sparsely strigillose with appressed albidous 
heirs; leaves decussate-opposite; petioles very slender, 3-- 
5 mm. long, sparsely strigillose; blades submembranous, lan- 
ceolate, 4--8 cm. long, 1.8--3.5 cm. wide, attenuate to a 
subacuminete point at the apex, abruptly acute or short- 
aouminate at base, regularly serrate from the widest part to 
the apex with greatly appressed bluntish teeth, sparsely al- 
bidous-strigillose above, glabrate but rather densely black- 
punctate beneath; inflorescence axillary, 1--1.8 cm. long; 
peduncles very slender, 5--12 mm. long, strigose; heads a- 
bout 5 mm. long and 13 m. in diameter; flowers white. 

The type of this species was collected by Bugene Langlas- 
86 (no. 286) -- in whose honor it is nemed -- at Pantla, at 
an altitude of about 5O m., Guerrero, Mexico, on August 18, 
1898, and is deposited in the Delessert Herbarium of the 
Conservatoire et Jardin Botaniques at Geneva. It is closely 
related to L. hispida H.BeK. 


1940 Moldenke, Additional Novelties 435 


STACHYTARPHETA LUNDELLAE Moldenke, sp. nov. 

Herba perennis, ad basin lignosa; ramis gracilibus obtuse 
tetragonis dense breviterque hirsutulo-pubescentibus; foliis 
oppositis; petiolis alatis dense patento-pubescentibus; lam- 
inis firme chartaceis obovato-subspathulatis, ad apicem rot- 
undatis vel obtusis, ad basin longe acuminatis, regulariter 
serratis, supra dense substrigoso-pubescentibus, subtus den- 
sissime velutinosis; inflorescentiis terminalibus spicatis 
ubique dense puberulis gracilibus. 

Perennial herb, woody at base; stems light-gray, glabres- 
cent; branches slender, obtusely tetragonal, densely short- 
pubescent with spreading hirsutulous sordid hairs; nodes not 
plainly annulate; principal internodes 3--8 cm. long; leaves 
decussate-opposite; petioles not very distinct, winged, 5-- 
10 mm. long, densely hirsutulous-pubescent; blades firmly 
chartaceous, uniformly gray-green on both surfaces, obovate- 
spatulate, 4--8 om. long, 1.8--35 cm. wide, obtuse or rounded 
at apex, regularly serrate from the apex to the base of the 
expanded portion, very long-acuminate at base and indisting- 
uishably attenuate into the petiole, densely substrigose- 
pubescent above, very densely velutinous beneath; midrib 
slender, plane above, prominent. beneath; seconderies slend- 
er, 4--6 per side, ascending, not much arcuate, plane above, 
prominulous beneath; veinlet reticulation indiscernible ab- 
ove, usually obscure beneath; inflorescence spicate, termin- 
al, 17--23 cm. long, densely many-flowered, densely puberu- 
lent throughout; rachis slender, deeply excavated, densely 
puberulent; bractlets lanceolate, 7--8 mm. long, long-atten- 
uate at apex, uniformly puberulent throughout; corolla cer- 
ise-red, purplish in the throat, its tube about 10 mm. long, 
its limb 6--7 mm. in dismeter. 

The type of this showy species was collected by Cyrus 
Longworth Lundell and Amelia A. Lundell (no. 7412) -- in 
whose honor it is named -- along a roadside at Chichen 
Itza, Yucatan, Mexico, in June or July, 1938, and is deposi- 
ted in the Britton Herbarium at the New York Botanical Gard- 
en. The species is related to S. purpurea Greenm., from 
which it may be distinguished at once by its larger bract- 
lets, which are uniformly puberulent, instead of being mere- 
ly ciliate-margined. 


STACHYTARPHETA MEXICANA Moldenke, sp. nov. 

Frutex; ramis obtuse tetragonis vel subteretibus dense 
albido-pubescentibus; foliis oppositis; petiolis gracilibus 
marginatis dense albido-pubescentibus, ad basin amplexicaul- 
ibus; laminis membranaceis rhomboideo-ellipticis vel rare 
lanceolato-ovatis acutis vel acuminatis serratis, ad basin 
longe cuneatis, supra scabrellis et sparse pilosis, subtus 
canescento-pubescentibus; spicis dense canescento-pubescent- 


436 PHYTOLOGIA No. 13 


ibus; rhachide leviter excavato. 

Shrub, about 1 m. tall; branches obtusely 4-angled or 
subterete, brown, densely white-pubescent with short and 
more or less spreading hairs; upper internodes 5.5--/+5 cm. 
long; modes more or less annulate; leaves decussate-opposi- 
te; petioles slender, 1--1.5 cm. long, not distinct, margin- 
ed, clasping at base, densely white-pubescent; blades memb~- 
ranous, lighter beneath, rhomboid-elliptic or rarely lanceo- 
late-ovate, 7--14 cm. long, 2--5.5 cm. wide, acute or acum- 
inate at apex, long-cuneate at base into the winged petiole, 
coarsely serrate along the margins except on the lower 1/3 
or 1/2, scabrellous and sparsely pilose above, canescent- 
pubescent beneath with very short spreading hairs; midrib 
slender, mostly flat and not prominent on either surface; 
secondaries very slender, 4--6 per side, ascending at an a- 
cute angle, mostly not very arcuate, plane above, very ob- 
scurely prominulous beneath; spikes terminal, 15--25 cm. 
long, slender, densely canescent-pubescent throughout with 
short spreading hairs, densely flowered, the flowers over- 
lapping except at the base of the spike, closely appressed 
into shallow excavations in the rachis, the spikes florifer- 
ous to the base or with a peduncle 1--2 cm. long; bractlets 
large and conspicuous, ovete-lanceolate, 11--15-mm. long, a- 
bout 2 mm. wide at base, long-acuminate and subulate at a- 
pex, densely short-pubescent; corolla blue. 

The type of this species was collected by George B. Hint- 
on (no. 4504) in an oak woods at luvianos, district of Tem- 
ascaltepec, Mexico, Mexico, on August 10, 1933, and is dep- 
osited in the herbarium of the Royal Botanic Gardens at Kew. 


DURANTA REPENS var. CANESCENS Moldenke, vare nove. 

Haec varietas a forma typica speciei recedit rhachidibus 
pedicellisque celycibusque densissime incano-strigosis et 
calycibus plerumque longioribus angustioribusque. 

This variety differs from the typical form of the species 
in having the rachis of its racemes and its calyxes very 
densely white-strigose, the pedicels very densely short- 
pubescent with canescent spreading hairs, and the calyx in 
anthesis somewhat longer and narrower, usually 4--4.5 mm. 
long and 1--1.5 mm. wide. 

The type was collected by Ellsworth Paine Killip and Alb- 
ert Charles Smith (no. 14990) at an altitude of 1000--1500 
m. on the northern slope of the Mesa de los Santos, Santand- 
er Sur, Colombia, between December 11 and 15, 1926, and is 
deposited. in the herbarium of the Field Museum of Natural 
History at Chicago. It is known also from Margarita Island 
and Lara, Venezuela. The corolla is purple, white at center. 


STACHYTARPHETA MUTABILIS var. VIOLACEA Moldenke, var. nov. 


1940 Moldenke, Additional Novelties 437 


Haec varietas a forma typica speciei recedit spicis 5 m. 
in diametro et corollis violaceis vel caeruleis vel purpur- 
eis. 

This variety differs from the typical form of the species 
in its spikes being more slender, averaging about 5 mm. in 
diameter (excluding the corollas) instead of 10 mm., and the 
corollas being blue, violet, or purple in color. 

The type was collected by Alexander Frank Skutch (no. 
4024) in a clearing, as an escape from cultivation, at an 
altitude of 1055 m., in the vicinity of El General, San Jo- 
sé, Costa Rica, in February, 1939, and is deposited in the 
Britton Herbarium at the New York Botanical Garden. The var- 
iety is known also from Cartago and from Guerrero, liexico. 
It is said to be a shrub, sometimes 7 or 8 feet tall, used a 
great deal in Costa Rice as a fence and in gardens, where it 
is s favorite of hummingbirds. 


STACHYTARFHETA TABASCANA Moldenke, sp. nov. 

Fruticulus; ramulis gracillimis glaberrimis pernitidis 
valde tetragonis; foliis oppositis; petiolis indistinctis 
marginetis glaberrimis; laminis submembranaceis ellipticis 
vel oblanceolatis abrupte acutis serratis, ad basin longe a- 
cuminatis, utrinque glaberrimis; spicis gracillimis ebbrevi- 
atis ubique glaberrimis. 

Shrubby; branches and branchlets very slender, light-gray 
or almost white, conspicuously tetragonal, often with marg- 
ined engles on the larger parts, perfectly glabrous, shiny; 
nodes annulate; principal internodes 1--5.5 cm. long; leaves 
decussate-cpposite; petioles indistinct, perhaps about 5 m. 
long, winged-margined, glabrous; blades submembranous, rath- 
er uniformly dark-green on both surfaces or slightly lighter 
beneath, elliptic or oblanceolate, 4--8 om. long, 1.4--% cm. 
wide, abruptly acute at apex, regularly and often somewhat 
coarsely serrate from almost the base of the expanded port- 
ion to the apex with rather broadly triangular sharply acute 
or apiculate somewhat spreading teeth, long-ecuminate and 
attenuate into the petiole at base, perfectly glabrous on 
both surfaces; midrib slender, plane above, prominulous be- 
neath; seconderies very slender, 4 or 5 per side, ercuate- 
ascending, short, usually obscure above, subprominulous be- 
neath; vein end veinlet reticulation obscure or indiscernib- 
le on both surfaces; inflorescence terminal, spicate; spikes 
abbreviated, about 12 cm. long, very slender; rachis glab- 
rous, excavated; bractlets lanceolate, 53--3.5 mm. long, att- 
enuate at apex, glabrous; flowers very small. 

The type of this species was collected by Eizi Metuda 
(no. 3218) at Reforma, Balancan, Tabasco, Mexico, between 
May 22 and 26, 1939, and is deposited in the Britton Herbar- 
ium at the New York Botanical Gerden. 


438 PHYTOLOGIA No. 13 


STACHYTARFHETA VELUTINA Moldenke, sp. nov. 

Frutex; ramulis obtuse tetragonis dense brevissimeque 
pubescentibus; foliis oppositis vel approximatis sessilibus; 
laminis chartaceis anguste ellipticis acutis vel acuminatis 
irregulariter serratis, ad basin cuneatis, supra densiuscule 
piloso-pubescentibus, subtus densissime velutinis; spicis 
vercrassis dense multifloris ubique dense hirtello-pubescen- 
tibus; rhachide valde excavato. 

Shrub, about 2 m. tall; branchlets very obtusely tetrag- 
onal, gray-brown, very slightly 4-margined, densely pubes- 
cent with very short gray-brown hairs; principal internodes 
2--4 cm. long; nodes not annulate; leaves decussate-opposite 
or approximate, the members of a pair sometimes separated by 
intervals of 2--15 mm.; petioles obsolete; blades chartace- 
ous, dark-green above, somewhet lighter beneath, narrow- 
elliptic, 5.5--11 cm. long, 1.5--3.5 cm. wide, acute or ac- 
uminate at apex, irregularly serrate along the margins ex- 
cept at the bese, cuneete at base, rather abundantly pilose- 
pubescent above with spreading hairs of various lengths, 
very densely velutinous beneath; spikes terminal, 20--32 cm. 
long, very stout and heavy, densely flowered almost to the 
base, the flowers closely appressed in deep excavations in 
the incrassate rachis, almost overlapping, densely hirtell- 
ous-pubescent throughout; bractlets lanceolate-ovate, 7--10 
mm. long, about 2 mm. wide eat base, sharply acute or acumin- 
ete et apex, divaricate in anthesis, later appressed; style 
and stigmas persistent long after the corollas have dropped 
off. 

The type of this species was collected by George B. Hint- 
on (no. 1801) by the river, at an altitude of about 1080 m., 
Vigas, district of Temascaltepec, Mexico, Mexico, on Septem- 
ber 22, 1932, and is deposited in the herbarium of the Royal 
Botanic Gardens at Kew. The species is also known from Mich- 
oacane 


VERBENA HARBISONII Moldenke, spe nove 

Herba; ramis acutiuscule tetragonis dense canescento- 
hirtellis; nodis valde annulatis; foliis oppositis; petiolis 
alatis dense canescento-hirtellis; laminis chartaceis prof- 
unde 3-partitis hirsutulo-pubescentibus, lobis lanceolatis 
argute acutis subrevolutis; inflorescentiis spicatis longe 
pedunculetis; spicis oblongis densissime multifloris. 

Herb; stems erect, slender, rather acutely tetragonal, 
ridged, rather densely hirsutulous-pubescent with short, di- 
vergent, canescent hair; branches similar but usually more 
densely canescent-hirsutulous, ascending, short; nodes on 
stems usually elongate, 4--8 cm. long, more abbreviated on 
branches, plainly annulate; leaves decussate-opposite; peti- 
oles 3--10 m. long, winged-margined, densely hirsutulous- 


1940 Moldenke, Additional Novelties 439 


pubescent with canescent heir; blades chartaceous, uniformly 
green on both surfaces, 2--5.5 cm. long, 2--5 om. wide, the 
larger ones deeply 3-parted (practically to the base), the 
divisions lanceolate in outline, deeply and irregularly pin- 
netisect, all the lobes sharply acute at apex, the mergins 
slightly revolute, the lateral divisions widely divergent, 
the uppermost leaves much reduced, with linear divisions, 
all hirsutulous-pubescent above and on the venation beneath; 
venation slightly impressed above, prominent beneath; in- 
florescence spicate, rather long-pedunculate, terminating 
the upper branchlets; peduncles slender, acutely tetragonal, 
densely canescent-hirsutulous, 2--5.5 cm. long; spikes ob- 
long, 1--2.5 cme long, about 1.5 cm. wide, very densely 
many-flowered; bractlets lanceolate, equaling the calyx, 5-- 
6 mm. long, about 1 mm. wide at base, long-attenuate to a 
setaceous apex, densely hirsutulous with short hairs; calyx 
membranous, cylindric, its tube about 5 mm. long, 0.9 mm. 
wide at base and 2.2 mm. at apex, densely short-pubescent 
with spreading white hairs, its rim 5-lobed, the lobes about 
2.5 mm. long, setulose, densely pubescent; corolla hypocrat- 
eriform, glabrous outside, its tube about 6.8 m. long, its 
limb about 5 mm. in diameter, the lobes small, about 2 m. 
long and wide, rounded and bifid at apex; stamens 4, didyna- 
mous, inserted about 2.4 and 4.4 mm. below the mouth of the 
corolla-tube, included; filaments filiform, very short, 0.2- 
-0.4 mm. long; anthers about 0.9 mm. long and 0.7 mm. wide, 
une ppendaged; style capillary, about 4.9 mm. long, glabrous; 
ovary oblong, about 1.4 mm. long and 0.7 mm. wide, glabrous. 

The type of this species was collected by Charles F. Har- 
bison -- in whose honor it is named -~- at Agua del Refugio, 
Baja California, ‘iexico, on April 1, 1935, and is deposited 
in the herbarium of the Sen Diego Society of Natural History 
at San Diego, California. The species is quite distinct from 
211 known species of this region. Its most obvious charact- 
ers are its long-pedunculete densely-flowered oblong spikes, 
elongate setaceous brectlets, and deeply 3-parted leaves 
whose lobes are all pleinly acute at the apex. 


VERBENA HINTONI Moldenke, sp. nov. 

Herba; ramis gracillimis acute tetragonis plusminus hirs- 
utulis; nodis annulatis; foliis oppositis; petiolis breviss- 
imis vel obsoletis; laminis lanceolatis vel ellipticis, ad 
apicem acutis, ad basin cuneatis, marginibus crasse pauci- 
dentatis vel 3-laciniatis vel sublobatis utrinque plusminus 
hirsutulis; inflorescentiis spicatis elongatis gracilibus. 

Herb, to 1 m. tall; stems and branches very slender, 
brownish, acutely tetragonal, more or less short-hirsutul- 
ous; nodes annulate; principal internodes 1.5--6 cm. long; 
leaves opposite, the upper ones sessile or subsessile; peti- 


440 PHYTOLOGIA No. 13 


oles very short and obscure or obsolete; blades thin-charta- 
cesous, uniformly green on both surfaces, narrowly lanceolate 
or elliptic in outline, the upper ones 1.5--2 cm. long, 2-- 
11 mm..wide, acute at apex, cuneate at base, coarsely few- 
dentate along the margins or 3-leciniate or 3-lobed below 
the middle, more or less short-hirsutulous on both surfaces, 
especially on the venation beneath; midrib and the l or 2 
secondaries very slender, impressed above, prominulous be- 
neath; veinlet reticulation indiscernible on both surfaces; 
inflorescence axillary and terminal, spicate, elongate, to 
23 om. long, many-flowered, the flowers densely imbricate 
before and during anthesis, loosely scattered in fruit; ra- 
chis very slender, sparsely pilosulous; peduncles obsolete 
or extremely short; bractlets lanceolate, 1.5--2 mm. long, 
acuminete, ciliate; calyx 2.5--45 mm. long, strigillose; 
corolla light-purple, about 7 mm. long, its limb about 4 m. 
in diameter. 

The type of this species was collected by George B. Hint- 
on (no. 11991) -- in whose honor it is named -- on a grassy 
bank at Zitacuaro-Bosque, district of Zitacuaro, Michoacan, 
Mexico, on June 28, 1938, and is deposited in the Britton 
Herbarium at the New York Botanical Garden. 


VITEX QUINATA var. WITTROCKIANA Moldenke, Geogr. Distrib. 
40, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit ramulis sar- 
mentisque pedunculisque rhachideque pedicellisque petiolis- 
que calyceque ubique glabris; inflorescentiis parvioribus 
saepe non-brachiatis; cymis longe stipitatis divaricato- 
furcetis, ramvlis longis gracillimis; pedicellis elongatis 
filiformibus. 

This variety differs from the typical form of the species 
in its smaller inflorescences, which are only 10--12 cm. 
long, less thyrsoid, often not branched, with few-flowered 
and very lax cymes which are long-stipitate and divaricate- 
furcate with long and very slender branches, and elongate 
filiform pedicels 3--5 mm. long; the branchlets, twigs, ped- 
uncles, rachis, cyme=branches, pedicels, petioles, and calyx 
glabrous throughout; and the flowers larger. 

The type was collected by Benedict Balansa (no. 3815) in 
cultivation at Tonkin, French Indo=china, in May, 1887, and 
is deposited in the herbarium of the Royal Botanic Gardens 
at Kew, It is also known from cultivation in Hongkong. 


ALOYSIA DUSENII Moldenke, sp. nov. 

Frutex; ramulis gracillimis leviter adpresso-puberulis; 
foliis oppositis; petiolis gracillimis; leminis chartaceis - 
anguste ellipticis acutis vel subapiculatis intezris vel 
vaucidentatis, ad basin acutis, utrinque levissime adpresso- 


1940 Moldenke, Additional Novelties Ady 


puberulis, in senectute supra pustulato-asprellis scabris; 
inflorescentiis spicatis; spicis dense multifloris nutentib- 
US. 

Bushy; branchlets very slender, finely appressed-puberul- 
ent, twigry; leaves decussate-opposite; petioles very slend- 
er, 1--3 ma. long, finely appressed-puberulent; blades thin- 
chartaceous, narrow-elliptic, 1.8--5 cm. long, 7--14 m. 
wide, acute and often subapiculate at apex, entire of with 1 
Or 2 small teeth on each margin near the apex, acute at 
base, very finely appressed-puberulent on both surfaces, be- 
coming pustulate-asprellous and roughened above in age; in- 
florescence snicate; spikes to 9 cm. long, 6--9 mm. wide, 
densely meny-flowered, nutant; peduncles elongate, very 
slender, to 4 cm. long, densely appressed-puberulent; calyx 
about 2 mm. long, hirsute; corolla about 5 mn. long, its 
limb about 4 mm. in diameter. 

The type of this species was collected by Per Karl Hjal- 
mar Dusén (no. 1050a) on a shrubby campo at Tamandaré, Pare- 
na, Brazil, on October 4, 1914, and is deposited in the her- 
berium of the Botanisches Museum at Berlin. 


ALOYSIA VIRGATA var. ELLIPTICA (Briq.) Moldenke, comb. nov. 
Lippia virgata var. elliptica Briq., Ann. Conserv. Jard. 
Bote Genev. 7--8 7304, 1904. 


CITHAREXYLUM KOBUSKIANUM Moldenke, Geogr. Distrib. 23, nom. 
nud. (1939), sp. nov. 

Arbor; ramis mediocriter gracilibus glabris; hornotinis 
gracillimis mimute puberulis; nodis smplietis; cicatricibus 
elevatis suberosis; foliis oppositis; petiolis gracillimis 
minute puberulis; laminis chartaceis oblongis vel oblongo- 
ellipticis acutis brevissime apiculatis integris, ad basin 
acutis vel breviter acuminatis, supra sparsissime puberul- 
is, subtus dense puberulis; inflorescentiis axillaribus rac- 
eniformibus; calyxe puberulo. 

Tree; branches medium-slender, grayish, glabrous; young- 
est twigs very slender, brownish, minutely puberulent; nodes 
somewhat ampliste; principal internodes 1--4 cm. long; older 
leaf-scars lerge, prominent, corky; leaves opposite; petiol- 
es very slender, 2--5 mm. long, minutely puberulent; blades 
chartaceous, uniformly dull-green on both surfeces, oblong 
or oblong-elliptic, 2.5--5 cme long, 1.-5--2.2 cm. wide, acm 
ute and very shortly apiculate at apex, entire, acutely or 
shortly acuminate at base, very sparsely puberulent above, 
densely puberulent beneath; midrib slender, subimpressed and 
densely puberulous above, prominulous beneath; secondaries 
slender, 7--10 per side, slightly arcuate-ascending, obscure 
above, slightly prominulous beneath; veinlet reticulation 
indiscernible above, very slightly promimlous beneath; in- 


Ao PHYTOLCGIA No. 13 


florescence axillary, racemiform, to 6.5 cm. long, many- 
flowered; rachis slender, puberulent; fruiting-calyx very 
light and herbaceous, puberulent outside; fruiting-pedicels 
about 4% mm. long, brownish-puberulent; fruit subglobose or 
oblong, about 5 mm. long end 4 mm. wide, fleshy, glabrous. 

The type of this species was collected by Jelski (A. Rai- 
mondi 6303) at Chota, Chichayro, Cajamarca, Feru, on Septem- 
ber 10, 1877, and is deposited in the herbarium of the Bota- 
nisches Museum at Berlin. The species is named in honor of 
my good friend and colleague, Dr. Clarence Emmeren Kobuski, 
of the Arnold Arboretum, who has produced an excellent mono- 
graph of the genus Priva. 


CITHAREXYLUM PERNAMBUCENSE Moldenke, Alph. List Common Names 
26, nome nud. (1939); Geogr. Distrib. 25, nom. nud. 
(1939), sp. nov. 

Arbor; ramis griseis glebretis; ramulis subteretibus min- 
ute puberulentibus glabrescentibus; foliis oppositis vel 
suboppositis; petiolis gracillimis minute puberulentibus; 
foliis tenuiter chartaceis subovatis vel oblongis vel obova- 
tis acutis vel obtusis intesris, ad basin ecuminatis, supra 
minutissime obscureque puberulis vel strigillosis glabres- 
centibus, subtus dense velutino-pubescentibus; inflorescent- 
iis terminelibus axillaribusque erectis multifloris, florib- 
us pseudo-subsecundis. 

Tree; branches gray, glabrate; branchlets brown, subtere- 
te, minutely puberulent, glebrescent in age; nodes not annu- 
late; principal internodes 2.7--5.4 cme long; leaves decuss- 
ate-opposite or subopposite; petioles very slender, 9--15 mm 
long, brown, minutely puberulent, conspicuously canaliculate 
ebove; blades thin-chartaceous, rather shiny above, subov- 
ate or oblong to obovate, 8.8--12 cm. long, 5.3--7 cm. wide, 
acute or obtuse at apex, entire, acuminate at base end bear- 
ing two elongate glands elong the midrib beneath, very minu- 
tely and. obscurely puberulent or strigillose above, glabres- 
cent in age, densely velutinous-pubescent beneath; midrib 
slender, practically flat above, prominent beneath; second- 
aries slender, 5--7 per side, arcuate-ascending, not plainly 
anastomosing, prominulous beneath; racemes terminel and ax- 
illary, erect, to 24 cm. long, about 2.5 cm. wide, many- 
flowered, the flowers pseudo~subsecund; peduncles 1.5--6.5 
cme long, slender, densely puberulent; rachis slender, dens- 
ely puberulent; pedicels to 1 mm. long, densely puberulent; 
calyx about 5 mm. long and 4 m. wide, puberulent; corolla- 
tube 15--17 mm. long, its limb about 10 mm. in diameter. 

The type of this species was collected by my good friend, 
Don Bento Pickel (no. 1501) at Caruarti, Tapera, Fernambuco, 
Brazil, in Januery, 1928, end is deposited in the herbarium 
of the Botanisches Museum at Berlin. 


1940 Moldenke, Additional Novelties 443 


CITHAREXYLU!! RIMBACHII Moldenke, spe nov. 

Arbor; ramis remulisque percrassis fistulosis valde tet- 
regonis pulverulento-puberulis; nodis valde annuletis; peti- 
olis percreassis puberulis; leminis magnis coriaceis ovatis 
acutis integris, ad basin subtruncetis, supra glabris, sub- 
tus dense adpresso tomentosis; inflorescentiis exillaribus 
spicatis dense multifloris. 

“iddle-sized forest tree, with broad crown and thick ra- 
ther smooth bark exfoliating in longitudinal strips; branch- 
es and branchlets very stout and robust, hollow, brown, de- 
cidedly tetrezgonal, much flattened st the nodes, pulverul- 
ent-puberulent, usually decidedly corky-margined helow the 
nodes; nodes annulate, the annulation deeply U-shaped (at 
least on one side); leaves opposite; petioles very stout, 
3.5--4 om. long, flettened above, ridged in drying, ampliate 
at base, puberulent; blades very large, coriaceous, oval or 
ovate, 25--30 cm. long, 15--19 cm. wide, dark-green above, 
somewhet lighter beneath, acute at apex, entire, subtruncete 
et base (rarely ebruptly acute), glabrous above, densely ap- 
pressed-tonentose beneath with yellowish many-brenched 
hairs; midrib very stout, slightly impressed above, very 
prominent end merely puberulent beneath; seconderies strong, 
11--16 per side, arcuate-ascending, very slightly impressed 
above, very prominent beneath with the pubescence wearing 
off from the most elevated portions; vein and veinlet retic- 
ulation abundant, only the coarsest portions prominent be- 
neath, only the finest portions visible ebove (!); inflores- 
cence axillary, spicate; spikes opposite, solitary, 13--21 
cme long, cebout 2 cm. wide in anthesis, densely many-flower- 
ed; peduncles stout, 2.5--3 cm. long, densely furfureceous- 
tomentose with cinereous hairs, engulate in drying; rechis 
stout (stouter than the peduncle), more densely cinereous- 
tomentose with longer hairs than the peduncles; bracts and 
brectlets none; prophylla conspicuous (especially when the 
calyxes have dropped off), reflexed, 1.5--2 mm. long, dense- 
ly tomentose, linear-subulate; pedicels none; celyx in anth- 
esis very large and heavy, more or less infleted, 5.5--7 mm. 
long, tubuler or broadly cupuliform or urceolate, densely 
short-tomentose, often 5—-costate with darker or less pubes- 
cent lines, its rim 5-toothed, the teeth broadly triangular, 
about 1 mm. long, acute; immeture fruiting-calyx decidedly 
urceolate, almost enclosing the fruit, very decidedly in- 
fleted, with the 5 dark ribs very conspicuous; immeture 
fruit globose. 

The type of this species was collected by Dr. August Rim- 
bach (no. 374) -- in whose honor it is named -- in the for- 
est region between the Chimbo river and the village of Bals- 
apempa in the Western Cordillera, «t an altitude of 2500 m., 
Ecuador, in December, 1934, and is deposited in the herbar- 


4h PHY FO L@6 FAs No. 13 
ium of the Naturhistoriska Riksmuseet at Stockholm. 


CLERODENDRUM ACULEATUM var. GUIANENSE Moldenke, Geogr. Dist- 
rib. 22, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit margine cal- 
ycis tantum breviter denteto, dentibus triengulari-ovatis 
O.7--1 mu. longis acutis nec ecuminetis nec caudetis nec 
patento-reflexis. 

This variety differs from the tynical form of the species 
in its celyx-rim being only short-toothed, the teeth triang- 
ular-ovate, O.7--1 mm. long, acute, not at ell acuminate or 
caudate or spreeding-reflexed. 

The type was collected by Jean Baptiste Leblond (no. 282) 
in French Guiane in 1792, and is deposited in the Delessert 
Herberium of the Conservatoire Botenique at Geneva. 


CLERODENDRUM DUCKEI Moldenke, Geogr. Distrib. 26, nom. nud. 
(1939), Spe NOVe 

Frutex; remulis gracilibus obtuse tetregonis pvlusminus 
puberulis vel subglabrescentibus; sarmentis gracilibus brev- 
ibus dense flavo-tomentosis; foliis ternatis; petiolis grac- 
illimis 5--8 mm. longis dense breviteraue pubescentibus; 
laminis ellipticis vel elliptico-oveatis acutis-vel breviter 
acuminatis integris, ad basin acutis vel acuminetis, supra 
pulverulento-puberulis vel glabrescentibus, subtus plusminus 
pulverulento-puberulis et punctatis; inflorescentiis axill- 
aribus terminealibusaue abbreviatis confertis, cymis pauci- 
floris; pedunculis pedicellisque gracillimis dense pubescen- 
tibus. 

Shrub; branchlets slender, obtusely tetragonal, grayish, 
more or less puberulent or becoming subglabrate in age; 
twigs slender, short, subterete or obscurely tetragonal, 
densely pubescent with tomentose yellowish hairs; nodes not 
annulate, marked with 3 prominent and divaricate corky ster- 


igmata 1--2 mm. long; principal internodes much abbreviated, © 


5--32 mm. long or much less; leaves ternete (or the lowermo- 
st on « twig only approximate); petioles very slender, 5--8 
mm. long, densely short-pubescent; blades thin-chartaceous, 
dark-green (nigrescent in drying) above, lighter (brunnes- 
cent in drying) beneath, elliptic or elliptic-ovate, 2--5.6 
cme long, 1--3.2 cm. wide, acute or short-acuminate at apex, 
entire, acute or acuminate at base, pulverulent-puberulent 
above, becoming glabrate, more or less pulverulent-puberul- 
ent and punctate beneath; midrib very slender, subimpressed 
above, prominulous beneath; secondaries very slender, 5--7 
oer side, arcuste-ascending, often short and not very arcu- 
ate, obscure or indiscernible above, flet or very slightly 
prominulous beneath; vein and veinlet reticulation sparse, 
indiscernible above, very inconspicuous or obscure beneath; 


ee ———-_--- 


1940 Moldenke, Additional Novelties 44S 


inflorescence axillary in the uppermost axils and terminal, 
ebbreviated, congested, the cymes few-flowered, to 2.5 cm. 
long; peduncles very slender, 5--8 mm. long, densely pubes- 
cent; sympodia and inflorescence-branches obsolete or much 
abbreviated and densely pubescent; pedicels very slender, 1- 
2 mm. long, densely pubescent; bracts 1 or more pairs, sub- 
tending the larger cymes of the terminal inflorescence, ses- 
sile, oblenceotate-spatulate or oblong, densely yellowish- 
or cinereous-pubescent, 3--8 mn. long; bractlets and vrophy- 
lla linear-subulate or setaceous, 1--2 mm. long, densely 
pubescent; calyx about 3 mn. long and 2 mm. wide, appressed- 
puberulent; corolla-tube ebout 15 mm. long, its limb 9--10 
mne in diameter. 

The type of this species was collected by Adolfo Ducke 
(no. 9924) -- in whose honor it is nemed -- at Igarapé do 
Zreré, ‘‘onte Alegre, Farad, Brazil, on December 14, 1908, and 
is deposited in the Delessert Herbarium of the Conservatoire 
Botanique at Geneva. 


CLERCDENDRUM EKMANI Moldenke, Geogr. Distrib. 29, nom. nud. 
(1939); Prelim. Alph. List Invalid Nemes 32, h:ponym 
(1940), sp. nov. 

Fruticulus; ramulis gracilibus obtuse tetragonis densiss- 
ime villoso-tomentosis; petiolis gracilibus dense hirsutis; 
leminis ovatis vel ovato-ellipticis obtusis vel rotundatis 
regulariter serratis, ad basin subcordatis vel rotundetis 
vel acutis, utrinoue dense villosis; inflorescentiis axill- 
aribus cymosis; corollis valde elongatis. 

Low bush; branchlets slender, obtusely tetragonal, very 
densely villous-tomentose with long. albidous hairs; nodes 
not annulate; principal internodes 1-~3 cm. long; leaf-scars 
not conspicuous; leaves decussate-opposite; petioles slend- 
er, 5--17 mm. long, densely hirsute with long whitish hairs, 
flettened above, not noticeably annulate ut base; blades 
membranous, rather uniformly dark-green on both surfaces, o- 
vate or ovate-elliptic, 3.3--5.8 om. long, 1.5--4 om. wide, 
obtuse or rounded at apex (in outline), varying from subcor- 
date to rounded or acute at base, usuclly sonewheat prolonged 
into the petiole, regularly serrate from base to apex with 
acute or bluntish antrorse teeth of uniforz size throughout, 
densely villous on both surfaces with whitish hairs 1--1.5 
mie long, especially dense on younz leaves and on the larger 
venetion beneath; midrib slender, plene above, prominulous — 
beneath; secondaries very slender, 4--6 per side, ercuate- 
ascending, 10stly plane on both surfaces; vein and veinlet 
reticulation obscure or indiscernible on both surfaces, 
hidden by the dense pubescence; inflorescence axillary, cyn- 
ose; peduncles very slender, about 1 cm. long, hirsute like 
the petioles; corolla-tube to 9 cm. long. 


446 PHYTOLOGIA No. 13 


The type of this remarkable species was collected by H. 
Quiroga at San Ignacio, !isiones, Argentina, on Cetober 2l, 
1913, and is deposited in the Delessert Nerbarium of the 
Conservatoire Botanique at Geneva. It is named in honor of 
Zrik Leonard Elman, who slso collected it and who considered 
it to represent a new genus of Verbenecese. 


CLERODENDRUM MARGARITENSE Moldenke, Geogr. Distrib. 20, nom. 
nud. (1939), sp. nov. 

Frutex; ramulis sarmentiscque subgracilibus obtuse tetrag- 
onis brunneis dense brevitercue pubescentibus vel tantum 
puberulis; internodiis abbreviatis; foliis ternetis; petio- 
lis gracillimis dense breviterque pubescentibus; leminis 
chertaceis ellipticis acutis vel brevissime subacuninatis 
integris (plerumque plusminus revolutis), ad basin acutis 
vel subecuminatis, supra densiuscule subtus densissime brev- 
iterque pubescentibus et punctatis, pilis flevidis; inflor- 
escentiis axilleribus, ad epicem sarmentorum confertis; cyn- 
is abbreviatis ternatis laxiuscule multifloris ubiaque dens- 
issime breviteraue pubescentibus; pvedunculis pedicellisque 
grecilibus. 

Shrub; brenchlets and twigs rather slender, obtisely tet- 
razonal, brownish, densely short-pubescent or merely puberu- 
lent in ege; younger nodes seeningly ennulate through the 
confluence or persistent pnetiole-base margins, older nodes 
not ennulete; principal internodes 1--21 mm. long, usually 
extremely abbrevicted on voung twigs or even subobsolete; 
leaves ternate or ternete-anproximate; petioles very slend- 
er, 1--/7 mn. long, densely snort-pubescent, its base persis- 
ting as e stout corky «nd non-aculeate spur-like projection 
l--2 mn. long after the blade is shed, in whose axil may of- 
ten be found a bluntish or sharp spine 3--44 mm. long, which 
is the lowest pert of the peduncle left when the remcinder 
broke off; blades chertsceous, derk grey-green scbove, bright 
-green or yellow-zreen beneath, elliptic, 1--4.5 cm. long, 6 
--12 mm. wide, acute or very shortly subecuminate at anex, 
entire (and often more or less revolute in drying) slong the 
margins, acute or subacuminate at base, rether densely short 
-pubescent above, much more densely so beneath with yellow- 
ish brown hairs end densely punctate; midrib very slender, 
impressed above, prominulous beneath; secondaries very slen- 
der, 3--7 on each side, ercuate-ascending, obscure or indis- 
cernible above, slightly prominulous or obscure beneath; 
vein and veinlet reticulation very delicate, impressed or 
obscure above, not at all prominulous and often even obscure 
beneath; inflorescence exillary, congested at the termine- 
tions of the twigs so as to appear terminal; cymes abbrevia- 
ted, ternate, solitary, 1.5--5 cm. long, 0.6--2 cm. wide, 
rather loosely meny-flowered, very densely short-pubescent 


1940 Moldenke, Additional Novelties AAT 


(like the lower leaf-surfaces) throughout; peduncles slender 
and 6--10 mm. long, pubescent; pedicels slender, 2--4 mn. 
long, pubescent; bractlets linear-subulate, 2--5 mm. long,. 
densely pubescent; prophylla minute, setaceous; calyx about 
3 mme long and wide, appressed short-pubescent; corolla-tube 
7--10 mm. long, its limb ebout 7 mm. in diameter. 

The type of this species was collected by John Robert 
Johnston (no. 82) en route from El Valle to Asuncion, “arg- 
arita Island, Venezuela, in August, 1903, and is deposited 
in the United States National Herbarium at Washington. It 
has hitherto been confused with the continental C. molle 
H.B.K. 


CLERODENDRUM RUSBYI Moldenke, Geogr. Distrib. 26, nom. nud. 
(1939), spe nov. 

Frutex vel arbor; ramulis gracilibus obtuse tetragonis; 
nodis non annuletis; petiolis gracillimis glabris, in ster- 
igmatibus magnis prominentibus suberosis non aculeiferis or- 
ientibus; leminis membraneceis ellipticis ecutis vel brevit- 
er acuminatis intesris, ad basin acutis, ut videtur gleb- 
ris; inflorescentiis axillaribus 3-floris laxis; pedunculo 
vedicellisaue -racilibus glabratis; calyce campenulato cras- 
so glabro, margine S-apiculato vel breviter 5-dentato; cor- 
olla hypocrateriforni slebra, tubo elongato gracillimo. 

Shrub or tree; brenchlets slender, obtusely tetregonal, 
more or less 4-sulcete, grey; nodes not annulate; leaves de- 
cussate-opposite; petioles very slender, about 5 m. long, 
glabrous, borne on very large and prominent corky sterigmata 
about 1 mm. long and 2.2 mm. in diameter, not aculeiferous; 
blades membranous, elliptic, dark-green end glabrous above 
(lower surface not es ebout 5.4 em. long and 2.7 cme 
wide, acute or very snort-acuminete at apex, entire, acute 
at base; midrib very slender; seconderies very slender, ab- 
out 6 per side, ascending, not much ercuate, very slightly 
prominulous above; inflorescence axillary, 4-flowered, oppo- 
site in the uppermost axils, lax; peduncle slender, 1.7--2 
cme long, glabrate; pedicels slender, S--7 mm. long, glab- 
rous; calyx campenulate, heavy, 3.5--5 mm. long, 4--6 mm. 
wide, glebrous, the rim S-epiculate or shortly 5-toothed; 
corolle hypocreteriform, glabrous, its tube very slender, 
2=--2.7 cm. long, 1 mn. wide for 3/4 its length, ampliate to 
53 mme below the limb, the limb spreading or reflexed, 5- 
parted; stamens long-exserted; filaments filiform, exserted 
2--2.5 cm. from the corolla-tube, glabrous; anthers oblong, 
about 2 mm. long and 1 mm. wide, 2-celled. 

The type of this species was collected by Henry Hurd Rus- 
by (no. 2572) -- in whose honor it is named -- at the falls 
of the ‘‘adeira River, Brazil, in Cctober, 1886, and is dep- 
osited in the Columbie University herberium at the New York 


Ae PH: ¥“8-0-5 0-6 Tk No. 13 
Botenical Garden. 


CLERODENDRUM TESSMANNI Moldenke, Geogr. Distrib. 23, nom. 
nud. (1939), spe nov. 

Frutex vel arbor; ramulis tetragonis crassis subglabris 
vel ad nodos complanatos vlusminus strigillosis; internodiis 
elongetis; foliis oppositis vel saepe anproximatis; petiolis 
crassiusculis vel crassis minute pulverulentis vel glabrat- 
is, ad basin in juventute corrugatis in senectute valde in- 
cressetis; leminis chartaceis ellipticis breviter acuminatis 
integris, ad basin cuneatis, utrinaque glabris; inflorescent- 
iis axilleribus vel supra-axillaribus; cymis 2--/-floris; 
redunculis crassis stremineis vel purpureis glabratis; pedi- 
cellis crassis slabratis. 

Shrub or tree; branchlets obtusely or ecutely tetregonal, 
buff-colored, stout, often very much decussate-flattened and 
broadened at esch node, subglabrate and shiny or more or 
less strigillose et end below the nodes and around the leaf 
sears; nodes not annulete; principal internodes elongate, 3- 
6.5 om. long; leaves decussate-opposite or often approximate 
with the 2 members of e pair separated by as much as 1.5 cms 
petioles stovtish or very stout when mature, 7--20 mm. long, 
minutely pulverulent or glabrate, more or less corrugated at 
base when immature, much incrassate at base when mature, of- 
ten deeply canaliculate above; blades chartaceous, rather 
uniformly bright-green on both surfaces, elliptic, 7--30.5 
cme long, 2./--8 cm. wide, short-acuminate at apex, entire, 
cuneete et base, glabrous on both surfaces; midrib stout, 
flat above, very prominent beneath; secondaries very slend- 
er, 8--1l per side, ercuste-ascending, flat or subimpressed 
ebove, sharvly prominulous beneath; vein and veinlet reticu- 
lation very slender, mostly obscure above, very slightly 
prominulous beneath; inflorescence axillary or supra-axill- 
ary; cymes solitary, ascending, opposite, 8--15 cm. long, 2- 
6 em. wide, 2--7-flonered; peduncles stout, 4--9 cm. long, 
stramineous or purplish, glabrate; pedicels stout, 5--l12 m. 
long, glebrate, erect or ascending, not divaricate; bracts 
none; bractlets few, narrowly spatulate, to 1.2 cm. long,’ . 
slebrate, often curvate, stipitete; prophylle .linear-subula- 
te, to 5 mm. long; calyx thick-textured, cempanulate-elonga- 
te, sbout 10 mm. long, 8 ma. wide at apex; corolla-tube ab- 
out 22 mn. long, its limb about 25 mm. in diameter. 

The type of this species was collected by Giinther Tess- 
mann (no. 3244) -- in whose honor it is named -- in the rain 
forest of the Ucayali from 10° S. to its mouth, Loreto, Peru 
in 1923, end is Deposited in the Delessert Herbarium of the 
Conservetoire Botenicue at Geneva. The species is also known 
from Ayacucho. 


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PHYTOLOGIA 


iveare to ‘Papedite botanical publication 


_ November, 1940 


CONTENTS 


ee Noteworshy spermatophytes from Mexico and Central 
bas Ameritas 449 


THe 
- 


_N. i Novelties among the American Verbenaceae..... 453 


5 Vol. bse 


Sts ; 
a, 


% 


NOTEWORTHY SPERMATOPHYTES FROM MEXICO & CENTRAL AMERICA (a) 


Ce be Lund ell 


CROTON CHICHENENSIS Lundell, spe nov. 

Frutex, albido-tomentosus. Petioli 0.6--2 cm. longi. Lam- 
ina chartacea, minute serrulato-denticulata, lanceolato- 
oblonga vel ovato-lanceolata, apice acuta vel acuminata, 
mucronulata, basi subcordata, supra stellato-pilosa, subtus 
elbido-tomentosa. Inflorescentiae unisexuales. Flores 9 ses- 
siles vel subsessiles; calyx 2--2.5 mm. longo, lobis triang- 
uleribus.: Flores & pedicellati; calyce 3 mm. longo; stamini- 
bus 18. Capsula stellato-tomentosa. 

A shrub, 1 m. high, with amber colored sap, whitish or 
pale tawny tomentose with stellate hairs. Branchlets slend- 
er, angled. Stipules inconspicuous, subulate, up to 3 mms 
long. Petioles 0.6 to 2 cm. long. Leaf blades chartaceous, 
minutely serrulate-denticulate, lanceolate-oblong or ovete- 
lanceolate, 3% to 8.5 cm. long, 1.1 to 3 om. wide, apex acute 
or acuminate, mucronulate, base subcordate, finely stellate- 
pubescent above, whitish tomentose beneath, pinnately vein- 
ed, costa and veins slightly impressed above, conspicuous 
beneath. Racemes unisexual, terminal, tomentose, the stamin- 
ate up to 13 cme long, the pistillate spicate, congested, up 
to 3 cm. long. Pistillete flowers sessile or subsessile; 
calyx 2 to 2.5 mm. long, S-lobed, the lobes trianguler or 
trisngular-lanceolate, glabrous within; petals sometimes de- 
veloped and resembling calyx lobes, sometimes vestigial; ov- 
ary stellate-tomentose; styles once-branched, practically 
glabrous on inner surfaces. Staminate flowers with pedicels 
up to 4 mm. long; calyx 3 mm. long, lobes ovate, glabrous 
' within; petals oblanceolate-oblong, about 3 mm. long, cili- 
ate, the hairs greatly elongated below middle, practically 
glabrous otherwise; stamens 18, filaments glabrous; recep- 
tacle pilose. Capsules tomentose. 


ico, May 30, 1938. 
Allied to C. Cortesianus H.B.K. 


CROTON PSEUDONIVEUS Lundell, sp. nove 
Arbor lepidotea. Petioli 0.7--2.3 cm. longi. Lamina memb- 
ranacea, repanda, ovata, apice subacuminata, obtusa, -basi 
rotundata vel subtruncata. Inflorescentiae bisexuales, fasc~ 
iculatae, usque ad 1.2 cme longae. Flores — pedicellati; 
calyce ca. 3 mm. longo, eebig vente. Flores 6 pedicellati; 
9 


450 PY EOL Oo -£ No. 14 


staminibus 10 vel ll. 

Tree, 6 me high, lepidote throughout. Branchlets slender. 
Petioles 0.7 to 2.3 cm. long, slender. Leaf blades membrana- 
ceous, repand, ovate, 5 to 8.5 cme long, 3.2 to 7 cm. wide, 
apex obtuse, base rounded or subtruncate, 3- or 5-veined, 
rather sparsely lepidote on both surfaces, the scales silv- 
ery with red center. Racemes bisexual, fasciculate in the 
leaf axils, crowded, very short, up to 1.2 cm. long, bracts 
villous-ciliate. Pistillate flowers with pedicels about 2 
mm.e long; calyx about 3 mm. long, lobes ovate; petals 5, 
broadly obovate, clawed, 3 mm. long, villous-ciliate; ovary 
lepidote; styles 3, branched 3 or 4 times. Staminate flowers 
with pedicels 1.5 mm. long; stamens 10 or 11, filaments 
glabrous. 

Type in the University of Michigan Herbarium, Ynes Mexia 
921, collected in thorny coast jungle, alt. 5 m., et Los 
Lebrados, Sinaloa, Mexico, Oct. 14, 1926; vernacular names 
"vera blanca" and "mueltilla". 

Closely related to C. niveus Jacq. with which it has been 
confused. 


CROTON TACANENSIS Lundell, sp. nov. 

Arbuscula videtur, farinoso-stellato-tomentosa. Petioli 
3--8 cm. longi. Lamina membranacea, subserrulata, ovata, ap- 
ice acuminata, basi subtruncata, emarginata. Inflcrescentiae 
bisexuales. Flores 9 pedicellati; calyce 4 mm. longo, lobis 
lenceolato-oblongis. Flores S pedicellati; staminibus 13--16. 

Apparently a small tree, with loose farinose yellowish 
tomentum of stellate heirs, the central ray of hairs elonga- 
ted and acicular on branchlets, vetioles, and inflorescence. 
Petioles 3 to 8 cm. long, with a pair of stalked glands at 
apex beneath. Leaf blades membranaceous, ovate, 7 to 15 cme 
long, 3.5 to 8 cm. wide, apex acuminate, base subtruncate, 
shallowly emarginate, the tomentum dense and persistent be- 
neath, at length nearly glabrous above, farinose, margin 
subserrulate, base 3-veined. Racemes terminal, up to 20 cm. 
long, lower bracts subtending both staminate and pistillate 
flowers. Pistillete flowers with pedicels up to 5-5 mn. 
long; calyx 3 mm. long, deeply S-lobed, lobes lanceolate- 
oblong; ovary stellate-hispid; styles bifid to base. Stamin- 
ate flowers with pedicels up to 3 mm. long; calyx 2-5 mm. 
long; petals 5, oblanceolate, about 2.5 mme long, villous; 
stamens 13 to 16, filaments villous below middle, receptacle 
villous. 

Type in the University of Michigen Herbarium, Eizi Matuda 
2943, collected at an altitude of 2100 m. on the north side 
of Volcan de Tacana, Chiapas, Mexico, on April 2, 1939. 

The species is a member of the complex of Croton gossypi- 
ifolius Vahl. 


1940 Lundell, Noteworthy spermatophytes 451 


CROTON TENUICAUDATUS Lundell, sp. nov. 

Arbor, 18 m. alta, lepidota et hirtella. Fetioli usque ad 
4.3 cme longi. Lamina membranacea, integra, ovato-oblonga 
vel elliptico-oblonga, 10--16.5 cm. longa, 4.8--8 cm. lata, 
eapice abrupte et obtuse caudato-acuminata, basi late rotund- 
ata. Inflorescentiae bisexuales, usque ad 16 cm. longae. 
Flores 2 pedicellati, pedicello 4--7 m. longo; celyce 3 mme 
longo, lobis 5, ovatis; petalis 5; stylis 3. Flores & pedi- 
cellati, pedicello 3--4 mm. longo; staminibus 12 vel 13. 

A tree 18 m. high, densely ferruginous-lepidote and short 
hirtellous. Branchlets rather stout, angled. Petioles slend- 
er, up to 4.3 om. long. Leef blades membrenaceous, entire, 
ovate-oblong or elliptic-oblong, 10 to 16.5 ome long, 4.8 to 
8 cme wide, apex abruptly and obtusely caudate-acuminate, 
base broadly rounded, densely lepidote at first on both sur- 
faces, glabrescent with age, eglandular, costa elevated be- 
neath, pinnately veined, primary veins 8 or 9 pairs, rather 
conspicuous beneath. Racemes axillary and terminal, bisexu- 
al, up to 16 cm. long. Pistilleate flowers with pedicels 4 to 
7 mm. long; calyx 3 mm. long, S-lobed, lobes ovate; petals 
5, broadly spatulate, long-clawed, up to 5.5 mm. long, lepi- 
dote without, pilose within, villous-ciliate; ovary lepid- 
ote; styles 3, each branched 4 times. Staminate flowers with 
pedicels 3 to 4 mm. long; calyx 3.2 mm. long, lobes ovate; 
petals 5, up to 4 m. long; stamens 12 or 13, filaments 
sparsely pilose, up to 5 mm. long; receptacle pilose. 

Type in the University of Michigan Herbarium, Alexander 
F. Skutch 2575, collected in open woods, vicinity of El Gen- 
eral, Prov. San José, Costa Rica, alt. 825 m., Feb. 1936. 

Referable to the complex of C. glabellus L. 


PERROTTETIA COSTARICENSIS Lundell, nom. nov. 
Perrottetia racemosa Standl., Field Mus. Bot. 18: 633. 
1937, not FP. racemosa Loes. 


PERROTTETIA SESSILIFLORA Lundell, sp. nov. 

Arbor, ramulis glabris. Petioli 6--10 mm. longi. Lamina 
integra, chartacea, lenceolato-oblonga, apice caudato- 
acuminata, basi rotundata et acutiuscula. Inflorescentiae 4 
paniculateae, flevo-puberulae. Flores sessiles vel subsess- 
iles, puberuli, ca. 1 mm. longi. 

Tree, 8 m. high, branchlets slender, glabrous. Petioles 
6 to 10 mm. long. Leaf blades entire, lanceolate-oblong, 
10.5 to 16 cm. long, 3.6 to 5.8 om. wide, apex caudate- 
acuminate, base rounded and acutish, thinly chartaceous, 
barbate in the axils beneath, otherwise entirely glabrous, 
primary veins 7 to 10 on’ each side, the costa and veins 
conspicuous beneath. Inflorescence paniculate, axillary, 
the panicles much-branched, subsessile, up to 7e5 cme long, 


452 PHYTOLOGIA No. 14 


9 cme wide, densely yellowish-puberulent. Pistillate flowers 
minute, about 1 mm. long, greenish-yellow, crowded, sessile 
or subsessile, puberulent. Calyx lobes about 0.4 mn. long, 
acutish. Petals triangular, acute, about 0.6 mm. long, glab- 
rous inside. Stamens shorter than petals, inserted on margin 
of thin shellow disk. Ovary free, glabrous, depressed-glob- 
ose, 2-celled, with 2 erect ovules in each cell; style 
slightly longer then ovary; stigma shallowly bifid. Stamin- 
ate flowers and fruits unknown. 

Type in the University of Michigan Herbarium, Alexander 
F. Skutch 2325, collected in vicinity of El General, Prov. 
San Jose, Costa Rica, alt. 915 m., December 1955. Duplicate 
in United States National Herbsrium (no. 1642337). 

This remerkably distinct species has entire leaves and 
sessile or subsessile flowers. 


GUNNERA KILLIPIANA Lundell, sp. nov. 

Herba perennis. Folia adpresse hirsuta, scabrida, corda- 
to-rotundata, lobata et dentata. Inflorescentiae adpresse 
hirsutee, spicae usque ad 1.8 cm. longae, bracteee fimbriat- 
ae, usoue ad 1.8 cm. longae. Fructus ovoideus. 

Coarse perennial herb, scabrid. Leeves long-stalked, 
large, cordate-rounded, palmately lobed, dentate, the teeth 
acute or acuminate, the longest ones up to © mm. long, both 
surfaces scabrid, the upper at first densely hirsute with 
short stout appressed brownish hairs, the under surface hir- 
sute only along the veins, palmately veined, the veins prom- 
inent beneath. Inflorescence densely subappressed hirsute, 
the fertile pert 25 cm. long. Flowers in compact short- 
stalked spikes up to 1.8 cm. long, each spike subtended by a 
subequal bract; calyx lobes up to 1.2 mm. long, with long 
filiform tip, the filiform part deciduous early; petals 
sparsely hirsute, deciduous early. Infructescence long- 
stalked, the fertile part up to 75 cm. long, the spikes up 
to 9 cm. long, hirsute; bract et base of spike lanceolate, 
oblong or oblanceolate, up to 1.8 om. long, sppressed- 
hirsute, fimbriate. Fruits (immature) glabrous, sessile, ov- 
oid, about 2 mm. long including persistent part of calyx 
lobese 

Type in the University of Michigan Herbarium, Sizi Matuda 
2763, collected at alt. of 1400 m. on Volcan de Tacena, Chi- 
epas, Mexico, March 17--23, 1959. 

Mr. E. P. Killip was first to recognize the plant as a 
distinct species, and it is with pleesure that I dedicate 
it to him. Gunnera Killipiana is near to G. insignis 
(Oerst.) A. DC. CGunnera mexicena Brandeg., the only other 
Mexican species, is know to the writer from the brief orig- 
inal description only. It apparently differs from G. Killip- 
iana amply in its leaf form being attenuate at the base 


1940 Lundell, Noteworthy spermatophytes 453 


rather then deeply cordate. 


NOVELTIES AMONG THE AMERICAN VERBENACEAE 


Harold N. kioldenke 


Continued studies in the herbarium of the New York Botan- 
ical Garden and elsewhere heve brought to light « good many 
as yet undescribed novelties in the Verbeneceae, some of 
which are presented herewith. More complete and detailed 
descriptions, as well as discussion of relationship end cit- 
ation of specimens examined, will be found in the author's 
forthcoming generic monographs. 


LIPPIA GEHRTII Moldenke, sp. nov. 

Frutex; ramis ramulisque gracilibus dense albido-hispid- 
is; nodis plusminus ennulatis; foliis oppositis; petiolis 
elato-marginatis dense albido-hispidis; laminis chertaceis 
late ellipticis vel subovatis, ad apicem rotundetis, ad 
basim truncatis vel subtruncetis, refzuleriter serreto- 
dentatis utrinque albido-hirsutis. 

Shrub; branches and brenchlets slender, obtusely tetrag- 
_onal, densely hispid with harsn white divergent hair 2.5--3 
mm. long; nodes hidden, but apparently more or less annule- 
te; principal internodes 0.9--% cm. long; leaves decussate- 
opposite; petioles slender, 3--7 mm. long, canaliculate a- 
bove, more or less wing-margined, densely white-hispid like 
the branchlets; blades chartaceous, rether uniformly derk- 
green on both surfaces, broadly elliptic or subovate, 1--4 
cme long, 0.7--3.2 cm. wide, rounded at apex, regularly 
serrate-dentate from base to apex with rounded or subacute, 
broad, short teeth, truncete or subtruncete ct base (very- 
ing to acuminate and prolonged into the petiole on young 
twigs), abundantly hirsute on both surfaces with long white 
hair similar to that on the branchlets; midrib slender, im- 
pressed above, prominent beneath; secondaries slender, ye) 
per side, ercuate-ascending, impressed above, prominulous 
beneath; veinlet reticulation abundant, impressed above, 
prominulous beneath; inflorescence borne in terminal open 
panicles, consisting of 2 or 3 (or more) decussate pairs of 
stipitate heads and a sessile terminal one, the stipitate 


454 Po YO LO EGsas No. 14 


ones ebout 1.5--2.5 em. long, subtended by e pair of foliac- 
eous bracts which are similar to the smaller leaves in ell 
respects or somewhat obovate, 6--15 mm. long and 5--10 mm. 
wide (or lerger), short-stipitate, densely hispid on both 
surfaces; axillary heads also often present in the uppermost 
leaf-axils beneath the penicle; inflorescence-branches dens=- 
ely white-hispid like the branchlets; heads oblong, 1.2--1./ 
em.e long, 1--1.5 cm. wide, densely many-flowered; bractlets 
large and conspicuous throughout the head, ovate, sessile, 
very thin-textured, 5--7 mm. long, 4 m. wide at base, sharp 
and acute at apex, softly pubescent with multicellular whit- 
ish somewhat bulbous-based hairs which are dense on the mar- 
gins and more scattered on the body of the bractlets; corol- 
la about 8 mm. long, its limb about 5 mm. in diameter. 

The type of this species was collected by Guilherme Gehrt 
-- in whose honor it is nemed -- at Itirapina, Sao Paulo, 
Brazil (Herb. Instit. Biol. S. Paulo 8313] on April 29, 
1923, and is deposited in the Britton Herbarium at the New 
York Botanical Garden. 


LIPPIA TEPICANA Moldenke, sp. nov. 

Frutex vel arbor; ramis ramulisque subgrecilibus saepe 
suberoso-marginatis dense resinoso-punctatis et griseo- 
puberulis; nodis annulatis; foliis oppositis; petiolis dense 
velutinis vel strigosis dense resinoso-granularibus; leminis 
crasse chartaceis, supra scaberrimis bullatis, ellipticis 
vel lanceolatis, ad apicem acutis, ad basim acuminetis, reg- 
ulariter serrulatis, supra bulboso-pilosis, subtus dense 
breviteraque pubescentibus vel tomentellis. 

Shrub or tree; branches and branchlets rather slender, a- 
cutely or obtusely tetragonal, sometimes corky-margined, 
densely puberulent with grayish eppressed hairs, less so in 
age, densely resinous-punctate; nodes ennulate; principal 
internodes 0.5--3.5 cme long; leaves decussate-opposite; 
petioles slender, 0.5--1.5 cm. long, varying from densely 
velutinous when young to merely short-pubescent or strigose 
and densely resinous-granular; bledes firmly thick-chartac- 
eous, uniformly gray-green on both surfaces or somewhat 
lighter beneath, very rough-scabrous and bullate ebove, ell- 
iptic or lanceolate, 2.5--9 cme long, 1--5 cme wide, acute 
at apex, regularly serrulate from almost the base to the a- 
pex with small acute antrorse teeth with subrevolute marg- 
ins, acuminate at base, pilose above with harsh bulbous- 
based whitish hairs, densely short-pubescent or tomentellous 
beneath (velutinous-tomentose when young); midrib slender, 
mostly impressed above (or appearing as though raised by 
being more densely pilose), rounded-prominent beneath; sec- 
onderies slender, 5--9 ver side, arcuate-ascending, impress- 
ed above, prominent beneath; veinlet reticulation abundant, 


Ve ete 


ee 


1940 ny Moldenke, Novelties 455 


impressed above, sharply prominent beneath; inflorescence 
axillary, peniculate, one panicle and sometimes also 1 or 2 
single-headed peduncles issuing from each axil or sometimes 
only several single-headed peduncles; the simple infloresc- 
ences 1--3 ome long, the penicles to 9 cme long; peduncles 
and inflorescence-branches very slender, 1--4.5 em. long, 
' densely puberulent, short-pubescent, or strigose with ciner- 
eous hair; penicles with 1--3 whorls of capitate branches; 
heads hemispheric during anthesis, oblong in fruit, densely 
many~flowered, 3--14 mm. long, 5--10 mm. wide; bircetibhe 
large, conspicuous, imbricate in many ranks, persistent, dry 
and scarious in fruit, reniform, 3--5 mm. long, 4--6 m. 
wide, densely puberulent on both surfaces during anthesis, 
glabrate and prominently venose in fruit; receptacle 3--7 
mu. longe 
oe type of this species was collected by Edward Palmer 

(no. 69), at Tepic, Nayarit, Mexico, between January 5 and 
ame oe 6, 1892, and is deposited in the Britton Herbarium 
et the New York Botanical Garden. 


STACHYTARPHETA CALDERONII Moldenke, Suppl. List Vern. Names 
23, nom. nud. (1940), sp. nov. 

Herba; remis crassiusculis subglabratis vel sparse albido 
-hirsutis; nodis hirsuto-annulatis; foliis oppositis; petio- 
lis alato-marginatis sparse hirsutis; laminis membranaceis 
ellipticis, supra scaberrimis, ad eapicem acutis, ad basim 
longe acuminetis, crassiuscule serratis utrinque sparse 
hirsutis. 

Herb; stems rather stoutish, obtusely tetragonal, subgla- 
brous or besring a few scattered, long, white, hirsute hairs 
at the nodes and when young; nodes annulate, mostly marked , 
by a circumferential band of long, white, more or less div- 
ergent, hirsute hairs 1--2 mm. long; leaves decussate- 
opposite; petioles slender, 5--10 mm. long, wing-margined, 
sparsely hirsute with widely scattered white hairs like 
those at the nodes; blades membranous, rather uniformly dark 
Breen on both surfaces, very scabrous above, elliptic, 5--9 
ems long, 1.9--2.6 cm. wide, acute at apex, long-acuminate 
at base and prolonged into the petiole-wings, rather coarse- 
ly serrate from below the middle to the apex with sharply a- 
cute often irregular teeth, sparsely hirsute on the lerger 
venation on both surfaces with scattered white hairs like 
those on the petioles and very sparingly so on the lemina 
itself or hirsutulous throughout on the upper surface, den- 
sely pustulate-punctate above; midrib slender, flat above, 
prominulous beneath; secondaries very slender, 6--8 per 
side, ercuate-ascending, flat above, flat or slightly prom- 
inulous beneath, joined in many loops near the margins; 
veinlet reticulation abundant, obscure or indiscernible a- 


456 PHY PO LO afta No. 14 


bove, flet beneath; spikes terminal, 11.5--25 cme long, very 
densely many-flowered, erect; flowers imbricate to the base; 
peduncles obsolete; rachis deeply excavated, stout, 3--4 mm. 
wide, glabrous or sparsely hirsute at the very base; bract- 
lets lanceolate, 6--7 mm. long, to 2 mm. wide, long-acunin- 
ate at apex, closely appressed, glabrous, the mergins entire 
and not scarious; corolla 8--10 mm. long, its limb about 5 
mme in diameter, glabrous. : 

The type of this species was collected by Salvador 
Salderén y Arana (no. 989) -- in whose honor it is named -- 
near Chalchuapa, Salvador, in 1922 and is deposited in the 
United States Nationel Herbarium at Washington. 


x STACHYTARPHETA HYBRIDA ifoldenke, hybre nov. 

erba perennis multibrachiatis; caulis ramisque gracili- 
bus sparse albido-hirsutulis glabrescentibus; nodis indist- 
incte annulatis; foliis oppositis plerumque fasciculatis; 
vetiolis obsoletis vel brevibus et late alatis; laminis ten- 
uiter chartaceis ellipticis, ad apicem acutis vel rotundat- 
is, ad basim abrupte cuneatis et in petiolum attenuetis, 
refuleriter serratis utrinque sparse albido-hirsutulis. 

Probably a hybrid between S. jamaicensis (L.) Vahl end. S. 
strigosa Vahl. *erennial herb, abundantly branched from the 
base; stems and branches slender, acutely or obtusely tet- 
ragonal, scattered-pilose with long, divergent, whitish, 
hirsutulous hairs, glabrescent in age; nodes indistinctly 
annulate; principal internodes 0.8--4 cm. long; leaves dec~ 
ussate~opposite, usually beering 2 or more smaller ones on 
greatly abbreviated twigs in their axils; petioles obsolete 
or, if present, broadly winged and scarcely distinguishable 
from the long-acuminate base of the blade; blades thin- 
chertaceous, uniformly colored on both surfaces, often brun- 
nescent in drying, the expanded portion elliptic, 1--45.5 cm. 
long, 0.5--2.1 cm. wide, acute at apex (or rounded in out- 
line), regularly serrate with short antrorse teeth, abruptly 
cuneate at base and attenuate into the long petiole-like 
acumination, which is to 2 cm. long, sparsely pilose on both 
surfaces with scattered whitish hirsutulous hairs, more 
densely so on the midrib and secondaries; midrib slender, 
flat on both surfaces or slightly prominulous beneath; sec- 
ondaries 4--6 per side, very slender, flat on both surfaces 
or slichtly prominulous beneath;. veinlet reticulation indis- 
cernible on both surfaces; spikes terminal, 15--24 cm. long, 
rather slender, many-flowered, the flowers imbricate but not 
especially dense; peduncles abbreviated, 1--2 cme long, 
sparsely pilose-hirsutulous; rachis about 3 mm. wide, deeply 
excavated, very sparsely scattered-pilose at base, glabres- 
cent above; brectlets lanceolate, 5--8.5 mme long, to 2 mm. 
wide, regularly triangular-attenuate from about the middle 


li he at i Ri ee 


r 
| 
4 
; 
| 


1940 Moldenke, Novelties 457 


to the filiform apex, the upper margins subscarious and usu- 
ally several-denticulete, very minutely and obscurely puber- 
ulous on the back and margins. 

The type of this hybrid was collected by Erik Leonard 
dxman (no. He11,978) in e pineland at an altitude of about 
1100 m., Sierra de Ocoa, prov. de Azua, San José de Ocoa, 
near Bejucal, Dominican Republic, on “iarch 18, 1929, and is 
deposited in the United States National Herbarium at Wash-- 
ington. 


STACHYTARPHETA MUTABILIS var. MAXONI Moldenke, var. nov. 

Haec varietas a forma typica speciei recedit spicis 
tenuioribus; ramis petiolisque sparsiore pubescentibus; lam- 
inis foliorum supra sparse pilosis, subtus dense adpresso- 
puberulis. 

This variety differs from the typical form of the species 
in its more slender spikes (4--5 mm. in diameter, exclusive 
of the corollas) and its less densely pubescent branches, 
petioles, and leaf-blades -- the upper leaf=surface being 
merely scattered-pilose, the lower surface merely densely 
puberulent with very short and appressed hair. The leaves 
are also larger,to 13 cm. long and 6 cm. wide, abruptly 
long-acuminete at base into the winged petiole. 

The type of this variety was collected by William Rand- 
olph Maxon (no. 7742) -- in whose honor it is named -- on an 
open bank neer shore of lake, et an altitude of about 300 m. 
at Laguna de Masaya, Nicaragua, on July 6, 1923, and is dep- 
osited in the United States National Herbarium at Washington. 


STACHYTARPHETA QUIROSANA Moldenke, spe nove 

Fruticulus; ramis gracilibus sparsiuscule subadpresso- 
pilosis; nodis annuletis; foliis ternatis fascieulatis; pet- 
iolis subobsoletis vel usque ad 5 mm. longis dense breviter- 
que pubescentibus; laminis tenuiter chartaceis ellipticis 
vel obovetis, ad apicem acutis, ad basim cuneato-acuminetis, 
regulariter serratis, supra sparsiuscule adpresso-pilosis 
non scabris, subtus dense breviterque pubescentibus. — 

Shrubby; stems slender, obtusely tetragonal (or S-engled) 
and rather sparsely pilose with short, white, subappressed, 
antrorse hairs; nodes annulate; principal internodes 3.5-- 
9.5 cm. long; leaves ternate, usuelly with several additio- 
nal small ones on greatly abbreviated twigs in their axils; 
petioles mostly obsolete on mature leaves, or slender, to 5 
mae long, and densely short-pubescent on small leaves; 
blades thin-chartaceous, dark-green above, slightly lighter 
beneath, elliptic or obovate, 2.5--7.5 cme long, 1--3.2 cm. 
wide, acute at apex, regularly serrate-dentate with broadly 
trianguler mucronulate teeth from about the middle or below 
the middle to the apex, cuneately nerrowed to the long- 


458 PHYTOLOGIA . No. 14 


acuminate base, rather sparsely scattered-pilose above with 
appressed antrorse whitish hair, not scabrous, densely 
short-pubescent with sordid eppressed hairs beneath; midrib 
slender, flat above, prominent beneath; secondaries very 
slender, 5 or 6 per side, flat and usually obscure above, 
arcuate-ascending, prominulous beneath; veinlet reticulation 
delicate, indiscernible above, flat and often obscure be- 
neath; spikes terminal, solitary, 16--50 cm. long, stout or 
slender, 5--10 mm. in diameter (exclusive of the corollas), 
densely many-flowered, the flowers imbricate above, barely 
so or separate toward the bese, somewhat loosely divergent 
during anthesis; rachis slender or stout, 2--5 mm. in diam- 
eter, densely appressed-pilose when young, less so in age, 
deeply excavated; brectlets lanceolate, ¢--7 mm. long, 1-- 
1.3 mm. wide, long-ettenuate to a filiform apex, sparsely 
pilose; corolla hypocrateriform, its tuba about 2 cm. long, 
its limb about 1--1.5 cm. in diameter. 

The type of this species was collected by August Weber- 
bauer (no. 6343) above Curilcds, in the valley of the 
Quiros, alt. 1700--2000 m., prov. Ayavaca, Piura, Peru, in 
May, 1912, and is deposited in the herbarium of the Field 
Museum of Naturel History eat Chicago. 


STACHYTARPHETA RORAIMENSIS Moldenke, spe nove 

Frutex (7); laminis foliorum chartaceis lenceolatis, ad 
apicem acutis, ad basim longe acuminetis, regulariter serra- 
tis subrevolutis, supra sparse pilosulis et minute pustule- 
to-scabris, subtus densiuscule puberulis; bracteolis lanceo- 
letis longe caudatis minute puberulis vel glabretis. 

Shrubby (7); leaf-blades chartaceous, dark-green above, 
somewhat lighter beneath, lanceolate, about 10 cm. long and 
53.5 cm. wide, acute et apex, regularly and abundantly serr- 
ate from below the widest point to the apex with antrorse 
rounded teeth about 1--2 mm. long and subrevolute-mergined, 
gradually narrowed to the long-ecuminate base, the basal 
acuminetion entire, sparsely pilosulous above and minutely 
pustulate, scabrous, rather densely puberulent (especially 
on the venation) beneath with short sordid hair; midrib 
slender, subimpressed above, prominent beneath; secondaries 
very slender, about 7 per side, arcuate-ascending, impress- 
ed above, prominulous beneath, not directly joined at the 


margins; veinlet reticuletion abundant, the larger portions | 


subimpressed above and prominulous beneath; spikes stout, 
about 1 cm. wide (exclusive of the corollas), very densely 
many-flowered with closely eppressed and imbricate flowers; 
rachis slender, cbout 2.5 ma. in diameter, glebrate; bract- 
lets lanceolate, about 1 cms long and 1 mm. wide at the 
widest point, long-caudate to a filiform apex, very minutely 
puberulent or glabrate. 


OX 


Bcd 


ee 


1940 Moldenke, Novelties 459 


The type of this species was collected by Moritz Richard 
Schomburgk on Mount Roraime, British Guiana, in 1842 or 1843 
and is deposited in the herbarium of the Royal Botanic Gard- 
ens at Kew, ea fragment in the herbarium of the Field Museum 
of Natural History et Chicego. 


VERBENA HALEI f. ROSEIFLORA (Benke) Moldenke, comb. nov. 
Verbene officinelis f. roseiflora Benke, Rhodora 45: 45, 


1933-6 


VERBENA NIVEA Moldenke, spe nov. 

Herba nane procumbens; ramis radiato-patentibus crecill- 
imis densiuscule cinereo-puberulis glabrescentibus; hornot- 
inis densissime breviterque cinereo-pubescentibus; nodis ob- 
scure annulatis; foliis oppositis numerosis fasciculetis; 
petiolis anruste slato-marginatis densiuscule vel sparse 
cinereo-strigosis; leminis charteceis ovatis utrinque dense 
cinereo-strigosis 3-lobatis vel 3-partitis, lobis 2- vel 3- 
lobuletis, lobulis obtusis revolutis. 

Low procumbent herb, branched from the base; branches 
spreading in radial fashion, very slender, sometimes elmost 
filiform, obtusely subtetrezonal, rather densely vcuberulent 


with very short svreading cinereous heir, more sparsely so 


or zlebrescent in age, the youngest branchlets very densely 
short-pubescent with spreading cinereous hairs; nodes faint- 
ly annulate; principal internodes 0.5--2.5 cm. long, mostly 
greatly abbreviated; leaves decussate-onposite, numerous, 
usually with severel much reduced ones in their axils; peti- 
oles slender, 1--/i m. long, flattened, deeply cenaliculete 
above, nerrovly winged-“ergined, rether densely or’ sparsely 
strigose with appressed whitish hairs; blades chartaceous, 
uniformly grey-green on both surfaces, ovate in outline, 4-- 
& mm. long and wide, densely strizgose with appressed whitish 
entrorse hair on both surfaces, 5-lobed or -perted elmost to 
the base, each division often again 2- or 35-lobed, the lobes 
all obtuse at epex and revolute-margined; the very slender 
midrib anc secondaries impressed above, slightly prominulous 
beneath; veinlet reticuletion indiscernible on both surf- 
eces; inflorescence terninel, capitate; heads small, meny- 
flowered, dense, subsessile or very short-pedunculete; cclyx 
tubuler, 3--/: mm. long, often purplish, densely short- 
pubescent with whitish rather spreading hair; corolla white, 
its tube cebout 5 mm. long, its limb about 2.5--5 mn. in di- 
ameter. 

The type of this species was collected by Santiago Vent- 
uri (no. 10,014) along the highway to San Antonio, Rosario 
de Lerma, elt. 3000 m., Salta, Argentina, in Yecember, 1929, 
and is deposited in the Britton “erbarium ct the New York 
Botcnical Gardene 


460 P.H YY 7.0¢L .0-G:iE A No. 14 


CLERCDENDRUM TERNIFOCLIUM var. MEXIAE Moldenke, Geogr. Dist- 
rib. 22, nom. nud. (1939), Vere NOVe 

Haec varietas a forme typica speciei recedit lobis caly- 
cis late triengulato-ovetis ad apicem acutis. 

This variety differs from the typical form of the species 
in the calyx-lobes being broadly tricengular-ovate and merely 
acute at apex, instead of long-ettenuate and acuminate. 

The type was collected by Ynes Mexia (no. 6757) -- in 
whose honor it is named -- on an overflow flet near sea lev- 
el between Guayaquil end Salinas, Guayas, Ecuedor, between 
December 10 and 14, 1934, and is deposited in the United 
States National Herbarium at Washington. 


DURANTA PSEUDOREPENS Moldenke, sp. nove 

Frutex; remulis pergracilibus tetragonis marginetis ad- 
rresso-puberulis; foliis oppositis; petiolis gracilibus min- 
ute puberulis; laminis subccriaceis ellipticis vel ovetis, 
ed epicem rotundatis, ad basim breviter acuminatis, serratis 
supre glabrescentibus, subtus minutissime puberulis; inflor- 
escentiis terminalibus exilleribusque dense puberulis. 

Shrub; branchlets very slender, tetragonal, margined, ap- 
pressed-puberulent; nodes not annulate; principal internodes 
1.5--4.5 cm. long; leaves ovposite or subopposite; petioles 
very slender, 5--6 mm. long, minutely puberulent; blades 
subcoriaceous, derk-sreen above, somewhat lighter beneath, 
elliptic or ovate, 2.8--4.5 cm. long, 1.6--3 cm. wide, roun- 
ded at apex, short-acuminate at base, serrate from the wid- 
est part to the apex, glabrescent above, very minutely pub- 
erulent beneath, revolute, the mergins rolled in egainst the 
top of the petiole et base; midrib slender, subimpressed a- 
bove, prominent beneath; secondaries slender, 3 or 4 per 
side, subimpressed above, sharply prominent beneath, arcuate 
-ascending; veinlet reticulation sparse, the larger perts 
subimpressed ebove and prominulous beneath; inflorescence 
axillary end terminal, racemiform, the axillary ones two per 
node, ebbreviated, 2--4 cm. long, rether few-flowered, the 
terminal one about 7.5 cm. long; rachis densely short- 
pubescent; pedicels slender,l--2 mm. long, densely short- 
pubescent; foliaceous bractlets often present, about 5 mm. 
long and 2.5 mm. wide; prophylla setaceous, 2 mm. long, 
short-pubescent; calyx tubular, 6--7 mm. long, 3--4 mn. 
wide, eppressed-puberulent, 5S-cpiculate; corolla-tube curv- 
ate, 1--1.3 cm. long. 

The tyre of this species was collected by Richard Spruce 
(no. 6042) in the Andes of Ecuador, between 1857 and 1859, 
and is deposited in the Britton “erbarium et the “ew York 
Botanical Garden. 


DURANTA HITCHCOCKII Moéldenke, sp. nov. 


1940 Moldenke, Novelties 461 


Frutex vel arbor; ramulis rracilibus valde spinosis dense 
adpresso-pubescentibus, pilis cinereis subtomentellis, in 
senectute dense puberulis, acute vel obtuse tetregonis; 
spinis oppositis vel ternatis; foliis oppositis vel approxi- 
matis vel ternatis coriaceis breviter petiolatis; laminis 
ellipticis vel obovatis, ad apicem rotundatis minute apicu- 
latis, integris revolutis, ad basim cuneatis, supra minute 
pilosulis vel glabratis, subtus plusminus dense puberulis 
glanduloso-punctatis; inflorescentiis axillaribus terminali- 
busque abbreviatis. 

Shrub or small tree, to 4 m. tall; branches and branch- 
lets slender, acutely or obtusely tetragonal, conspicuously 
spinose, densely appressed-pubescent with cinereous tangled 
hair, in age merely densely puberulent, opposite or ternate; 
spines opposite or ternate, slender, 6--20 mm. long, un- 
branched, very sharp and stiff; nodes not ennulate; princip- 
al internodes 1--2.5 cm. long; leaves numerous, opposite, 
approximate, or ternate; petioles very slender, 1--5 mm. 
long, densely appressed-pubescent; blades coriaceous or sub- 
coriaceous, slightly lighter beneath, elliptic or obovate, 
1--2.5 cm. long, 5--13 mm. wide, mostly rounded end minutely 
apiculate at apex, sometimes subacute, entire and slightly 
revolute along the mergins, cuneate at base and usually pro- 
longed into the petiole, minutely pilosulous sbove or glab- 
rate in age, more or less densely puberulent and glandular- 
punctate beneath; midrib slender, impressed above, prominent 
beneath; secondaries very slender, 2 or 3 per side, ascend- 
ing, not much arcuate, obscure or subimpressed above, prom- 
inulous beneath; veinlet reticulation indiscernible on eith- 
er surface; inflorescence axillary and terminal, racemiforn, 
abbreviated, few-flowered; calyx tubular, about 5 mm. long 
and 2 mme wide, densely short-pubescent; corolla-tube about 
12 mm. long,. its limb about 10 mm. wide, sweet-scented, 
pale-blue or lilac. 

The type of this species was collected by this distingu- 
ished American botanist, Albert Spear Hitchcock (no. 20821) 
-- in whose honor it is named -- between Otavalo and Malch- 
ingui, Imbabura or Pichincha, ecuador, at an altitude of 
2400--35000 me, on August 12, 1923, and is deposited in the 
Britton Herbarium at the New York Botanical Garden. It is 
also known from Chimborazo and Bolfvar. 


DURANTA PENLANDI Moldenke, spe nov. 

Frutex spinosus; ramis gracilibus debilibus; ramulis ac- 
ute tetragonis marginetis minutissime puberulis glabrescent- 
ibus ut videtur inermibus; foliis verticillatis; petiolis 
brevibus subglabretis; laminis subcoriaceis ellipticis ab- 
rupte acutis supra mediam serratis subrevolutis, infra med- 
jam integris valde revolutis, ad basim breviter acuminatis, 


462 PHYTCLOGIA No. 14 


utrinque glebris; inflorescentiis numerosis racemiformibus 
ubique minutissime puberulis glabrescentibus. 

Spiny shrub; branches slender, weak, to 4 m. long; 
branchlets slender, acutely tetragonal, margined, very minu- 
tely puberulent, soon glabrescent, apparently unarmed; nodes 
not annulate; principal internodes 3--4 cm. long; leaves 
whorled in 4's; petioles very slender and short, 2--4 m. 
long, subglebrate; blades coriaceous, dark-green and lust- 
rous ebove, slightly lighter beneath, elliptic, 1.9--3.8 cm. 
long, 1.2--2.1 cm. wide, abruptly acute at apex, short- 
acuminete at base, uniformly serrulate from about the middle 
to the apex with small appressed sharp-pointed teeth and 
subrevolute, entire and pronouncedly revolute from about the 
middle to the base, the two margins rolled in egainst the 
top of the petiole in charecteristic fashion, glabrous on 
both surfaces; midrib slender, subimpressed above, sharply 
prominent beneath; secondaries slender, 4-j-7 per side, sub- 
impressed above, sharply acute beneath; veinlet reticulation 
very sperse, obscure or indiscernible on both surfaces; in- 
florescences axillary, racemiform, 4 per node, 3.5--9 cm. 
long, rather loosely many-flowered, the flowers subsecund ; 
rachis and peduncles sharply tetragonal, very minutely pub- 
erulent, glabrescent; pedicels 2--4 mm. long, very minutely 
puberulent or glabrescent; prophylla minute, setaceous, 1--2 
mm. long; calyx tubular, 6--7 mm. long, ebout 4mm. wide, 
apiculate, very minutely puberulent, bluish-purple and 
green; corolla pale blue-lilac, its tube curvate, about 1 
em. long; fruit tough, juicy, dull-yellow, splitting the 
mature calyx. 

The type of this species was collected by Charles William 
Theodore Penland and Robert H. Summers (no. 958) in the vic- 
inity of Tambillo, at an altitude of 2800 m., Pichincha, Ec- 
uedor, on June 26, 1939, and is deposited in the Britton 
Herbarium at the New York Botanical Garden. It is named in 
honor of its distinguished senior collector. 


LANTANA GLAZIOVII Moldenke, sp. nove 

Nerba perennis nana, ad basim lignosa; remis erectis per- 
sracilibus, simplicibus densissime lanato-pubescentibus; 
foliis oppositis paucis sessilibus; laminis ellipticis vel 
subrotundis, ad basim et apicem acutis vel obtusis, subin- 
tegris vel serrulatis, utrinque dense lenato-pubescentibus. 

Very dwarf perennial herb, woody at base, to about 12 cm. 
tell, gnerled at base; stems several or numerous, erect, 
very slender, simple, very densely lanate-pubescent with 
white or cinereous hairs; leaves decussate-opposite, few, 
sessile; blades thin-chartaceous, elliptic or subrotund, 4-- 
14 mm. long, 3--6 mm. wide, acute or obtuse at both ends, 
varying from subentire to uniformly serrate with small blunt 


: 
: 


1940 Moldenke, Novelties 463 


teeth from almost the base to the apex, densely lanate- 
pubescent with albidous hairs on both surfaces; inflorescen- 
ce axillary; capitulae abundant in ell the upper leef-axils, 
solitary, opposite; peduncles very slender, 1--3 cm. long, 
ascending, densely hirsutulous-pubescent with whitish hairs; 
corolla-tube about 7 mm. long, its limb about 3 mm. wide. 
The type of this species was collected by Auguste Frang- 
ois Marie Glaziou (no. 21,892) -- in whose honor it is 
named -~- on the central pleteau of Goyaz, Brazil, in .1894 or 
1695, and is deposited in the Delessert Herbarium et the 
Conservatoire Botsnique et Geneva. 


TANTANA HAUGHTII Moldenke, sp. nove 

Frutex; ramis gracilibus tetragonis dense adpresso- 
puberulis glabrescentibus; foliis oppositis; setiolis grac- 
ilibus dense breviterque albido-pubescentibus; leminis firme 
chartaceis lanceolatis obtusis regulariter serrulatis, ad 
basim acutis vel acuminetis, supra bullatis dense adpresso- 
strigosis, subtus densissime velutino-tonentellis; inflores- 
centiis axillesribus capitatis ubique dense canescento- 
puberulis vel breviter pubescentibus; bracteolis megnis late 
ellipticis vel ovetis. 

Shrub; branchlets slender, obtusely or acutely tetragon- 
al, densely eppressed-puberulent, glabrescent in age; nodes 
not ennulete; leef-scars lerge, corky, divergent; principal 
internodes 1--7.5 cme long; leaves decussate~-opposite, often 
with several much smaller ones borne on much abbrevieted 
twigs in their axils; petioles very slender, 4--8 mm. long, 
densely short-pubescent with whitish heirs; bledes firmly 
chertaceous, dark-green above, grey-green beneath, lanceola- 
te or elliptic-ovate, 1--3.5 one long, O./--2 cme wide, ob- 
tuse et epex, reculerly serrulate, sacute or acuminate at 
base, bullate and densely appressed-strigose above, very 
densely velutinous-tomentellous benesth with sordid or whit- 
ish hairs; midrib slender, impressed above; prominent be- 
neeth; secondaries slender, 4--6 per side, ascending, not 
much. arcuate, deeply impressed above, slightly prominulous 
beneath; veinlet reticulation abundant, deeply impressed 4- 
bove, prominulous or hidden by the tomentum beneath; inflor- 
escence axillary, capitate; peduncles very slender, 2 per 
node, 3--6 om. long, densely appressed-pubescent or albidous 
-puberulent; heeds densely many-flowered, 1--1.5 om. in di- 
ameter; bractlets lerge, conspicuous, broadly elliptic or 
ovete, to 9 mn. long and 4 mm. wide, acute at apex, densely 
canescent-puberulent or substrigose; corolle white, its ‘ube 
about 7 mm. long, its limb cbout 3 mm. wide. 

The type of this species was collected by Oscer Hauzht 
(no. 139) -- in whose honor it is named -- in the Amotepe 
Sills, Piura, Feru, in November, 1927, and is deposited in 


464 PHYTOLOGIA No. 14 
the Britton Herbarium at the New York Botanical Garden. 


LANTANA PAVONII Moldenke, sp. nov. 

Frutex; ramulis sarmentisque gracillimis sparsissime hir- 
sutis; foliis oppositis; petiolis grecilibus alatis sparse 
hirsutis; laminis chartaceis ellipticis vel subrotundis acu- 
tis serruletis, ad basim acuminetis, supra sparsiuscule hir- 
sutulis (in senectute scabris bulletis), subtus dense hirsu- 
tis. 

Low shrub (7); brenchlets and twigs very slender, very 
sparsely hirsute with scattered hairs; leaves decussate- 
opposite; petioles slender, 2--5 mm. long, hirsute with sca- 
ttered hairs, alate-mergined; blades chartaceous, elliptic 
or subrotund, 1.2--3 em. long, 7--15 mm. wide, acute (or ob- 
tuse) at apex, rether densely serrate from the widest part 
(or below) to the apex with very small rounded teeth, acum- 
inate at base, rather sparsely hirsutulous above, becoming 
very rough and bullate in age, more densely hirsute (especi- 
ally along the venation) beneath; inflorescence axillary, 
2.5-=3.5 cme long, solitary, opposite, ascending; peduncles 
very slender, 2--2.8 cm. long, hirsutulous like the twigs, 
capitulae to & mm. long and 19 mm. wide. 

The type of this perplexing species was collected by José 
Antonio Pavon -- in whose honor it is named -- somewhere in 
Peru, and is deposited in the Delessert Herbarium at the 
Conservatoire Botanique at Geneva. It was named "Lantana 
scabra Pavon, ne spe" by the collector. 


LANTANA RUBELLA Moldenke, sp. nove 

Frutex; ramulis cracilibus tetragonis marginatis; sarmen- 
tis pergracillimis numerosis abbreviatis dense breviterque 
pubescentibus; foliis oppositis; petiolis gracillimis brevi- 
bus adpresso-puberulis; laminis submembranaceis ellipticis 
obtusis supra mediam serruletis, ad basim acutis, supra min- 
utissime puberulis vel glabrescentibus, subtus puberulis 
dense glanduloso-punctatis; inflorescentiis axillaribus sub- 
spicatis; brecteolis oblenceolatis; calycibus dense hirsutis 

Zrect shrub, to 2.5 m. tall; branchlets slender, obtusely 
or subacutely tetragonal, margined (the margins coming loose 
and splitting off in me densely short-pubescent with 
brownish hairs, glebrescent in age; twigs very slender, num- 
erous, abbreviated, densely short-pubescent; nodes annulate; 
principal internodes on twigs 3--12 mm. long, on branchlets 
to 5.5 cme long; leaves decussate-opposite, numerous; peti- 
oles very slender, short, 1--5 mm. long, appressed=puberul- 
ent with dark brownish hairs; blades submembranous, dark- 
green above, much lighter beneath, elliptic, 1--2e5 cm. 
long, 3--10 mm. wide, obtuse at apex, serrulate from the 
widest part to the apex with subacute teeth, acute at base, 


————— = - 


ee 


1940 Moldenke, Novelties 465 — 


very minutely puberulent or glabrescent above, puberulent 
and densely glendular-punctate beneatn; midrib very slender, 
ususlly subimpressed above, prominulous beneath; secondaries 
very slender, usually subimpressed above, 3 or 4 per side, 
subprominulous beneath; veinlet reticulation obscure or in- 
discernible above, plane beneath; inflorescence axillary, 
subspicate, few-flowered; peduncles very slender, 5--15 mm. 
long, appressed-pubescent; bractlets oblanceolate, about 5. 
mm. long and 1.5 mm. wide, rounded or truncate at apex, app- 
ressed-puberulent; calyx densely spreading-hirsute and res- 
inous-punctate; corolla deep-pink, its tube about 5 mm. 
long, its limb about 2 mn. wide. > 

The type of this species was collected by Ynes Mexia (no. 
5845) in a shallow moist valley at Diamantina, on the upper 
slope of the Serra do Rio Grande, at an altitude of about 
1290 m., Minas Geraes, Brazil, on May 15, 19351, and is dep- 
Osited in the Britton Herbarium at the New York Botenical 
Garden. 


LANTANA TOMASII Moldenke, sp. nov. 

Fruticulus; ramis ramulisque mediocriter gracilibus ob- 
tuse tetragonis dense puberulis; foliis oppositis; petiolis 
grecilibus dense tomentellis; lsminis chartaceis vel sub- 
membranaceis ovetis acutis vel subacuminatis, regulariter 
serratis, ad basim ecutis vel rotundatis, supra scabro- 
pilosis subbulletis, subtus dense tomentellis; inflorescen- 
tiis axillaribus capitetis; bracteolis magnis oblongo-ovatis 
dense breviterque pubescentibus. 

Shrubby, about 80 cm. tall; branches and branchlets med- 
ium-slender, obtusely tetragonal, unarmed, densely puberul- 
ent or tomentellous; nodes annulate; principal internodes 1 
--9 cm. long; leaves decussate-opposite; petioles slender, 
about 1 cm. long, densely tcementellous; bledes chartaceous 
or submembranous, ovate, 3--1l cm. long, 2--6 cm. wide, ac- 
ute or subacuminate at apex, regularly serrate from apex al- 
most to the base, rounded at base or acute when immeture, 
scabrous-pilose and subbullate above, densely tomentellous 
beneath; the slender midrib, secondaries, and veinlet retic- 
ulation impressed above and prominulous beneath; inflores- 
cence axillary, capitate, densely many-flowered, about 1.5 
eme long end wide; bractlets large and conspicuous, the low- 
ermost oblong-ovate, ebout 1 cm. long and 6 or 7 mm. wide, 
densely short-pubescent, obtuse at apex; corollas about 9 or 
10 mm. long, purple. 

The type of this species was collected by Brother Tomas 
(no. 579) -- in whose honor it is nemed -- at Béelo, Antio- 
quia, Colombia, on June 14, 1959, and is deposited in the 
United States National Herbarium et Yashington. It is obvi- 
ously closely releted to Lentane boyacana Moldenke, 


66 PHYTOLOGIA No. 14 


LANTANA V&LUTINOIDES Moldenke, sp. nov. 

Frutex; ramulis gracilibus acute tetragonis densiuscule 
pubescentibus; foliis ternatis; petiolis gracilibus dense 
pubescentibus; laminis chartaceis lanceolato-ovatis, ad api- 
cem acutis vel breviter acuminatis, ad basim rotundatis, 
regulariter serratis, supra dense pubescentibus, subtus den- 
sissime velutino-tomentosis; inflorescentiis axillaribus 
perabbreviatis spicatis; bracteolis ovatis dense breviterque 
pubescentibus. 

Shrub; branchlets slender, rether acutely tetragonal, 
rather densely short-pubescent; nodes not annulate; princip- 
al internodes 2.5--3-5 cm. long; leaves ternate; petioles 
slender, about 5 mm. long, densely short-pubescent; blades 
chartaceous, dark-green above, very much lighter beneath, 
lanceolate-ovate, 3--4.5 cme long, 1.5--2 cm. wide, ecute or 
short-acuminate at apex, regularly serrate from base to apex 
with small rounded teeth, rounded at base, densely short- 
pubescent with eppressed hairs ebove, very densely velutin- 
ous-tomentose with sordid hairs beneath; midrib slender, im- 
pressed above, prominulous beneath; seconderies very slend- 
er, impressed ebove, prominuloys beneath, 4--6 per side, ar- 
cuate-ascending; veinlet reticulation very abundant, impres- 
sed above, prominulous beneath and plainly visible through 
the tomentum; inflorescence axillery, much shorter than the 
subtending leaves, exceeding the petioles, 1.5--2.4 em. 
long, cepitete-spicate, densely many-flowered; peduncles 
slender, 5--9 mm. long, densely short-pubescent; bractlets 
conspicuous, ovete, about 5 mm. long and 3 mm. wide, densely 
short-pubescent with cinereous hairs, sharply acute et apex. 

The type of this species was collected by H. lmederwaldt 
(Herb. Instit. Biol. S. Paulo 15,669] eat Ipiranga, Sao 
Paulo, Brazil, in March, 1912, and is deposited in the Brit- 
ton Herbarium at the New York Botanical Garden. 


LANTANA VELUTINOIDES var. BRUCHII Moldenke, ver. nove 

Haec varietas a forma typica speciei recedit foliis in- 
florescentiisque oppositis, pedunculis usque ad 4.5 cm. lon- 
gis, spicis usque ad 3 em. longis, et bracteolis usque ad 8 
vel 9 mm. longis et leatis. 

This variety differs from the typical form of the species 
in having its leaves and inflorescences opposite, peduncles 
to 4.5 cm. long, spikes to 3 cm. long, and bractlets to 8--9 
mae long and wide. 

The type of this variety was collected by C. Bruch (no. 

2831) -- in whose honor it is named -- at Unquillo, Cordoba, 
Argentina, in 1926, end is deposited in the Britton Herbar- 
ium at the New York Botanical Garden. 


LIPPIA ASPERRIMA var. ROTUNDATA Moldenke, vare nove 


————— 


—=—S 


1940 Moldenke, Novelties 467 


Haec varietas a forma typica speciei recedit leminis fol- 
iorum ad epicem obtusis vel rotundatis. 

This variety differs from the typical form of the species 
in having its leaf-blades obtuse or rounded et the apex. The 
petioles are 1--4 mm. long, mergined; leaf-blades elliptic, 
1.8--3.7 cme long, 1.2--2.4 cm. wide, regularly serrate al- 
most to the base with blunt teeth, ecute at base and prolon- 
ged into the short petiole; venation decidedly impressed ab- 
ove, Only the midrib and seconderies slightly prominulous 
beneath; peduncles 5--3.5 cm. long; branchlets densely 
glandular-pubescent with short derk hairs and also copiously 
hirsute with white srreading hairs twice as long, less den- 
sely so in age. The hairs on the upper leaf-surface cppar= 
ently become bulbous-besed in age, first along the margins, 
finelly over the entire lamina. The upper leaf=-surfece is 
scabrous at all times. 

The type of this variety was collected by Frederico Carl- 
os Hoehne et Fonte Grossa, Paren&, Brazil, on November l, 
1928, and is deposited in the herberium of the Instituto Bi- 
Ologico de Yefesa Agricola e Animal et Sao Paulo [no. 23268] 


LIPPIA HOSHNEI Moldenke ex Hoehne, Resen. Hist. Comm. Viges. 
Anniv. Secc. Bot. 153 & 161, hyponym (1937), sp. nov. 

Fruticulus; remulis sarmentisque grecilibus ubique dense 
puberulis; foliis oppositis sessilibus amplexicaulibus; lam- 
inis suborbiculeribus rotundetis, ad basim cordetis, refu- 
laritcr serratis vel sinuatis, supra nitidulis minute puber- 
ulis, subtus densiuscule pubescentibus; inflorescentiis ax- 
illaribus terminalibusque paniculatis; brecteis mernis ovat- 
is sessilibus acutis intezgris vel serratis; capitulis densis 
breviter pedunculstis confertis. 

Shrubby; branchlets and twigs slender, ascending, buff- 
colored, densely puberulent throughout with very short erect 
brovmish hairs; nodes ennulate; leaves déecussate-opposite, 
sessile, clasping at base; bledes suborbicular, 3--6 cm. 
long and wide, rounded (in outline) at apex, cordate at 
base, refzularly serrate from base to apex with appressed and 
bluntish teeth or sinuate below, the very apex often marked 
with ea sharply «cute or acuminate tooth, minutely puberulent 
and rather shiny above, rather densely spreading-pubescent 
on the whole venetion beneath; midrib, seconderies, and vein 
and veinlet reticulation subimpressed above, very prominent 
and conspicuous beneath in a beautiful reticulum; inflores- 
cence axillery and terminal, clustered in a dense panicle at 
the tips of the branches, the inflorescence-branches erect 
or ascending, conspicuously brecteate with large ovate sess- 
ile and entire or serrate acute bracts; cepitulee dense, 
short-pedunculate, crowded. 

The type of this handsome species was collected by fred- 


468 Pon EP’ Och. oO: Gara No. 14 


erico Carlos Hoehne [Com. Rondon 2177] -- in whose honor it 
is named -- at Trés Jacts, Mattogrosso, Brazil, in 1908, and 
is deposited in the herbarium of the Instituto Biologico de 
Defese Azricola e Animal et Sao Paulo. 


LIPPIA MATTOGROSSENSIS Moldenke, sp. nov. 

Herba (?) perennis; remulis grecilibus obtuse tetragonis 
dense hirsutis; foliis oppositis; vetiolis sracilibus dense 
hirsutis; laminis tenuiter chartaceis ovatis vel elliptico- 
ovatis acutis serratis, ead basim acutis vel rotundatis, sup- 
ra hirsutis, subtus dense hirsuto-tomentosis; inflorescent- 
iis axillaribus capitetis confertis; capitulis densis. 

Herb (?) or shrub; branchlets slender, obtusely tetragon- 
al, brown, densely hirsute with long hite hairs (1--1.3 mm. 
long); rather twiggy; principal iriternodes 3--4.5 om. long; 
leaves decussate-opposite; petioles slender, 4--10 mm. long, 
densely hirsute like the branchlets; blades thin-chartace- 
ous, dark-green, lighter beneath, elliptic-ovate or ovate, 
2--5.5 om. long, 1.2--2.6 cm. wide, acute, serrate from base 
to apex with small blunt teeth (6--9 per cm. of margin), ac- 
ute or rounded at base, hirsute above with hairs precisely 
like those on the branchlets, twigs, and petioles, densely 
hirsute-tomentose beneath with somewhat shorter tangled 
whitish or cinereous hairs and also densely granuler-pulver- 
ulent and resinous; midrib slender, flat or obscurely subim- 
pressed above, prominulous beneath; secondaries slender, 6-- 
10 per side, usuelly irregular, arcuate-ascending, slightly 
subimpressed above, prominulous beneath; vein and veinlet 
reticulation abundant, rether obscurely subprominulous ab- 
ove, prominulous and dark beneath; inflorescence axillary, 
capitate, numerous, 4i--6 in each upper leaf-axil (rarely re- 
duced to 1 per axil in the lowermost axils of the twig 
peduncles very slender, 5--16 mm. long, densely long-hirsute 
like the twigs; heads dense, 4--10 mm. long, 7--9 ma. wide. 

The type of this species was collected by Frederico Carl- 
os Hoehne [Com. Rondon 470] at Coxipd da Ponte, lattogrosso, 
Brazil, in March, 1911, and is deposited in the herberium of 
the Instituto Biologico de Defesa Agricola e Animal at Sao 
Paulo. 


LIPPIA iiICROMERA var. PALUDICOLA Moldenke, var. nove 

Haec verietas a forma typica speciei recedit ramulis sar- 
mentisque tantum pulverulento-punctatis (non patento-pubes- 
centibus), foliis angustis integris tantum pulverulento- 
punctatis (non breviter pubescentibus vel puberulis). 

This variety differs from the typicel form of the species 
in having its branchlets and twigs merely densely pulveru- 
lent-punctate, instead of spreading short-pubescent, and in 
having its petioles and the very narrow and entire leaf- 


SS ny 


1940 Moldenke, Novelties 469 


blades also merely: pulverulent-punctate on both surfaces, 
instead of short-pubescent or puberulent. 

The type was collected by Jacques Semuel Blanchet (no. 
2872) in swamps at Ilhabina, Bahie (7), Brazil, in 1839, end 
is deposited in the Meisner Herbarium at the New York Botan- 
icel Garden. 


PETREA DUCKEI Moldenke, sp. nov. 

Frutex scandens; ramis gracilibus minute puberulis glab= 
rescentibus; foliis oppositis; petiolis crassis minute pub- 
erulis; laminis firme charteceis ellipticis acutis vel brev- 
iter acuminatis integris, ad basim acutis vel obtusis, ut- 
rinque minutissime puberulis glabrescentibus scabrellis, in- 
florescentiis axillcribus recemiformibus; lobis calycis per- 
magnis, in fructu late ellipticis. 

Scandent shrub; branches slender, grayish, minutely pub- 
erulent. glabrescent in age, sozewhat lenticellate; nodes 
not annulate; principal internodes 1.2--11.5 cm. long; 
leaves decussate-onposite; petioles stout, °--12 mn. long, 
minutely puberulent, flettened above; blades firmly cherta- 
ceous, stiff, rether uniformly gray-zreen on both surfaces, 
somewhat shiny ebove, elliptic, 10.5--16 om. long, 4.3--8 
cm. wide, abruptly acute or short-acuminate at apex (the 
very point often obtuse), entire, acute or rounded eat base, 
very minutely puberulent and scabrellous on both surfaces, 
glabrescent and merely punctate in age, the immature blades 
very thin-membrenous and nigrescent in drying; midrib rather 
stout at base, rapidly diminishing in size as the apex is 
epproached, prominent on both surfaces; secondaries very 
slender, 9--15 per side, prominulous above, sharply promin- 
ent beneath; vein and veinlet reticulation abundant, prom- 
inulous on both surfaces, the tertiaries sharply prominent 
beneath; inflorescence axillary, racemiform, 14--25 em. 
long, rether loosely many-flowered; rachis slender, minutely 
puberulent; pedicels slender, 2--5 mn. long, elongate to 9 
mae in fruit, minutely puberulent; calyx subtended by 1--5 
foliaceous prophylla, which sre thin-membrenous, elliptic, 
venose, 5--6 mm. long, 3--4 mm. wide, sharply acute or at- 
tenuete-acuminete at apex; fruiting-calyx indurated, its 
tube 6--7 mm. long, 5--7 mm. wide at apex, very minutely 
puberulent, its lobes greatly enlarged, broadly elliptic, to 
about 13 mm. long end 12 mm. wide, pinnately venose, very 
minutely and obscurely puberulent, abruptly acute or obtuse 
at apex. 

The type of this species was collected by Adolfo Ducke 
{Herb. Jard. Bot. Rio de Janeiro 22,542] -- in whose honor 
it is named -- on inundated shores at Parand de Anevilhana, 
on the lower Rio Negro, Amazonas, Brazil, on July 24, 1939, 
and is deposited in the Britton “erbarium et the-New York 


470 PH YoD- 9 1. 6:6:07h No. 14 


Botenical Garden. The species is obviously closely related 
to P. insignis Schau. 


STACHYTARPHETA AUSTRALIS Moldenke, Frelim. 4lph. List Invalid 
ames 42, hyponym (1940), sp. nov. 

Fruticulus; ramis gracilibus obtuse tetrazsonis zlabres- 
centibus; remulis gracilibus p!usminus breviter pubescentib- 
us; foliis oppositis; petiolis zracilibus adpressopubescent- 
ibus; leminis cherteceis ellipticis acutis arsute serratis, 
ad basim cuneato-attenuatis, supra dense puberulis scabrell- 
is, subtus breviter adpresso-pubescentibus; spicis dense 
multifloris ubique plusminus pilosis, floribus in rhachidem 
non velde immersis. 

Shrubby; branches slender, rather obtusely tetragonal, 
glabrous or subglabrate in age, light-grey; branchlets very 
slender, browmish, more or less densely short-pubescent; 
nodes not ennulate; principal internodes 1--5.5 cm. long; 
leaves decussate-opposite, small; petioles very slender, 5-- 
8 mm. long, avpressed-pubescent; bledes chartaceous, brunn- 
escent eb>ve in drying, somewhat lighter beneath, elliptic, 
1.5--4.5 om. long, 1--2 cm. wide, acute at apex, regularly 
serrate from almost the base to the apex with sharply acute 
small antrorse teeth, cuneate-ettenuate at base and prolong- 
ed into the petiole, densely puberulent and scabrellous ab- 
ove, appressed-pubescent beneath with short hair often more 
dense on the larger venation; midrib slender, plane or im- 
pressed above, prominulous beneath; secondaries very slend- 
er, 4+-6 per side, arcuate-ascendinz, plene or impressed ab- 
ove, prominulous beneath; veinlet reticulation indiscernible 
above, usually obscure beneath; inflorescence spicate, term- 
inel, to about 20 cm. long, densely many-floered; rachis 
very slender, more or less densely whitish-pilose; bractlets 
lanceolate, about 5 mm. long, scarious~margined, long- 
attenuate, more or less pilosulous-ciliate; flowers not 
deeply imbedded in the rachis. 

The type of this species was collected by Svnreiter 
(Herb. lus. Argent. Cience lat. 26/1347] et Tartagal, in the 
devartment of Oran, Salta, Argentina, on September 29, 1925, 
and is deposited in the Britton Herbarium at the New York 
Botanical Garden. The species is common in southern South 
America and is known from many localities in Brezil, Paragu- 
ay, Uruguay, and Argentina, as well. es in Hawaii and Java. 


STACHYTARPHETA AZUREA Moldenke, sp. nove 

Fruticulus (?); foliis epproximatis ellipticis maar 
us, ad apicem rotundetis, ac basim cuneatis serratis subrev- 
_ olutis tlebratis vel minute puberulis; inflorescentiis tern- 
inalibus elongetis spicetis longissime pedunculatis densiss- 
ime multifloris. 


1940 itoldenke, Novelties 471 


Shrubby (7); stems and branches not seen; uppernost leav- 
es approximate, the members of the pair about 3 em. apart, 
sessile, broadly elliptic, firmly ch: rtaceous, 9--10 cm. 
long, 4--5 cm. wide, rounded at apex, cuneate to the clasp= 
ing base, regulerly serrate from below the middle to the a- 
pex with rether bluntish teeth and subrevolute, ftabrous or 
minutely puberulent on both surfeces; inflorescence termin- 
el, spicete, elongate, long-pedunculate; peduncles stout, a- 
bout 1° cm. long, glubrate or minutely puberulent; florifer- 
ous portion of the spike rather stout, 25--29 cm. long or 
longer, sbout 1 cme in diemeter, very densely many-flovered, 
the flowers closely overlapping; rechis excavated; bractlets 
lanceolate, about 1 cm. long, lonz-eristate at apex, gleb- 
rate or minutely vuberulent; corollas blue. 

The type of this species was collected by Frederico Carl- 
os Hoehne [Commissao Rondon no. 692] at Forto Isperidiéo, 
uettogrosso, Brazil, in November, 1908, end is deposited in 
the herbarium of the Instituto Biologico de Defesa Agricola 
e Animal at Sado Faulo. 


STACHYTARPHETA BOLDINGHII Moldenke, sp. nove 

Fruticulus; ramvlis subteretibus obscure puberulis; fol- 
iis oppositis sessilibus; laminis ellipticis, ad apicem ro- 
tundetis, ad basim cunectis, merginibus reguleriter serrat- 
is, supre glabratis, subtus minute puberulis et dense punct- 
atis; inflorescentiis terminalibus spicatis elonzctis dens- 
issime multifloris; floribus imbricatis; rheachide profunde 
excavato; brecteolis enguste ellivticis longe aristulatis 
glabris. 

Shrubby; branchlets subterete, obscurely puberulent; 
nodes not <nnulete; leaves opposite or subopposite, sessile; 
blades chartaceous, somewhat lighter benesth, elliptic, the 
uppermost ones 1.5-2.5 cm. long, O./--1.5 cm. wide, rounded 
at apex, cuneate at base, regulerly serrate from the widest 
part to the apex with acute or bluntish teeth, minutely 
strigillose or subslebrate above, minutely puberulent and 
densely punctulate beneath; midrib and the 2 or 5 pairs of 
secondaries very slender, subimpressed above, prominulous 
benesth; veinlet reticulation obscure or indiscernible on 
both surfaces; inflorescence terminal, spicete, elonrate, to 
37 cm. long, very densely many-flowered, the flowers closely 
imbricate end deeply sunken in the rachis; rechis rether 
stout, ebout 3 mm. wide, glabrous, deeply excavated during 
and after anthesis; bractlets lerge, abundant, conspicuous, 
nerrow-elliptic, cbout 7 mm. long end 1.5 mm. wide, long- 
aristulete ct apex, scarious in a wide bend elong the marg- 
ins, glabrous; corolla-tube about 1 cm. long. 

‘ne type of this species was collected by Isa&e Boldingh 
(no. €352) -- in whose honor it is named -- on the island of 


472 PH ¥'T: 0:2: O:G-Eh No. 14 


Arube in 1909 or 1910 and is deposited in the Britton Herb- 
erium et the New York Botanical Garden. 


STACHYTARFHETA ELATIOR var. JENMANI Moldenke, Suppl. List 
Common Names 7, nom. nud. (1940), var. nov. 

Haec varietas a forma typica speciei recedit foliis ut- 
rinque plusminus hirsutis. 

This variety differs from the ty7ical form of the species 
in having both surfeces of its leaf-blades more or less 
short-hirsute, sometimes rather densely so, instead of glab-= 
rouse The stems and branches are sometimes also more or less 
hirsute. 

The type was collected by George Samuel Jenman (no. 5542) 
-- in whose honor it is named -- at Nuinatta on the Rupununi 
River, British Guiana, in October, 1889, and is deposited in 
the Columbia University Herbarium at the New York Botanical 
Garden. It is knovn also from Guarigo, Venezuela, and re- 
minds one of S. Calderonii “‘oldenke from Central America and 
S. pycnodonta Urb. from the West Indies. The corolles ere 
described as purple. A vernaculer neme is “esponjilla". 


STACHYTARFHETA HAXIMILIANI ver. CILIARIS iloldenke, var. nove 
Haec varietas a forma typica speciei recedit rnechide 
celycibusque spersissime pilosis et bracteolis tantum spar- 

siuscule cilictis. 

This variety differs from the typical form of the species 
in having the rechis and calyxes very sparsely pilose with 
short and widely scettered obscure wiitish hairs and the 
bractlets glabrous except for the sparsely and irregularly 
ciliate margins. 

The type was collected by Lyman Bradford Smith (no. 1420) 
along a peth on the brook trail between Faineiras and Jardim 
Botanico, et an altitude of 100--400 m., Rio de Janeiro, Fe- 
deral District, Brazil, on December 4, 1928, end is deposit- 
ed in the Britton Herberium at tne New York Botanical Garden 


STACHYTARPHETA MEXIAE Moldenke, spe nove 

Fruticulus; ramis ramulisaque obtuse tetregonis crassiusc- 
ulis densissime sordido-hirsutulis; foliis oppositis; petio- 
lis brevibus densissime sordido-hirsutulis; leminis chartac- 
eis ovetis ecutis reguleriter serratis, ad basim rotundatis, 
supra dense velutinoso-pubescentibus, subtus densissime sor- 
dido- vel canescento-tomentosis; spicis abbrevietis termin- 
elibus densissime multifloris ubique densissime albido- 
hirsutulis; brecteolis enruste lanceolatis elongatis. 

Shrubby, to 3.3 m. tall, with loosely spreading branches; 
brenches and branchlets rather stoutish, obtusely tetragon- 
al, very censely nirsutulous with sordid-gray or brownish 
heirs; nodes not ennulete; principel internodes 3--7 cm. 


1940 Moldenke, Novelties 473 


long; leaves opposite; petioles short, 5--9 mm. long, very 
densely sordid-hirsutulous; blades chartaceous, derk-creen 
ebove, greyish beneath, ovate, 2--5 cm. long, 1.5--3 em. 
wide, acute at apex, regularly serrate from base to apex 
with rather small acute teeth, rounded at base or often 
slightly prolonged into the petiole at the middle, densely 
velutinous-pubescent above with more or less appressed and 
forward-pointing hairs, very densely sordid- or whitish- 
tomentose beneath; midrib slender, plene above, prominulous 
beneath; secondaries slender, plane above, 4--7 per side, 
escending, slishtly arcuate, prominulous beneath; veinlet 
reticulation indiscernible above, plain beneath; inflores- 
cence terminal, spicate; spikes sessile, short, 5--9 cm. 
long, stout (about 1.5 cm. wide), very densely many-flower- 
ed; rachis very densely canescent-hirsutulous; brectlets 
dense, cons™icuous, very narrowly lanceolate, elongate, 10-- 
15 mm. long, densely canescent-hirsutulous, cttenuete at a- 
pex; corolla brignt-blue, about 2 cm. long. 

The type of this species was collected by Ynes “exia (no. 
56824) -- in whose honor it is named -- on damp ground in the 
edge of a tangle on an overgrowm slope at “hristais, Diamen- 
tinea, at ean altitude of 116C m., Mines Geraes, Brazil, on 
May 13, 1931, and is deposited in the Britton Herbarium at 
the New York Botanical Garden. 


STACHYTARFHETA PERUVIANA Moldenke, sp. nove 

Herba perennis; ramulis gracilibus glebratis nitidis; 
foliis membranaceis ovetis acutis serratis, ad basim longe 
acuminetis, utrinque slabris nitidis; spicis terminalibus e- 
longetis persracilibus ubiaque glabris. 

Perennial herb, to about 1 m. tall; branches very slend- 
er, glabrous, shiny, obtusely tetregonal, stremineous; nodes 
not annulate; principal internodes 5.5--11.5 cm. long; 
leaves decussate-opposite; petioles slender, indistinct, 
winged, «bout 1 cm. long, zlebrous; blades thin-membranous, 
uniformly bright~-freen and shiny on both surfaces, ovate, & 
--10 cm. long, 4--5.5 come wide, acute «et apex, rather uni- 
formly and coarsely serrate from almost the base of the ex- 
panded nortion to the apex with broadly triercular subacute 
and minutely epiculete teeth, very long-acuminate «t base 
and indistinzuishably attenuate into the petiole, cslabrous 
on both surfaces; midrib slender, plane ebove, flattened and 
herdly prominulous beneath; secondaries very slender, 5--7 
per side, ascending, not much arcuate, plane on both surf- 
aces; veinlet reticulation sparse, obscure above, plene be- 
neath; inflorescence terminal, spicate; peduncles very abb- 
reviated or obsolete, glabrous; rachis very slender, gleb- 
rous, 23--230 cm. long; brectlets lanceolate, about 4 m. 
long, shorter then the calyx, long-cttenuate et apex, scari- 


47h PHYTCOLOGIA No. 14 


ous-mergined near the base, glabrous; fruiting-calyx rather 
deeply impressed in the excavated rechis; corolla pinkish- 
blue, peler at the center within, its tube about 7 mm. long, 
extremely slender, its limb about 4 mm. wide. 

The type of this distinctive species wes collected by 
Elisworth Peine Killip and Albert Charles Smith (no. 22853), 
at the edge of a dense forest along the beach, et an altit=- 
ude of 400 m., near Kimpitiriki, in the Rfo Apurimac valley, 
Ayecucho, Peru, on May 10, 1929, end is deposited in the 
herbarium of the Field Museum at Chicago. 


STACHYTARFHETA SPRUCEI Moldenke, sp. nov. 

Fruticulus; reamulis acute tetragonis puberulis; foliis 
oppositis; petiolis gracilibus merginatis caneliculatis pub- 
erulis; leminis tenuiter -chartaceis brunnescentibus oveto- 
ellipticis acutis vel brevissime acuminatis regulariter ser- 
ruletis, ad basim acuminetis, supra sparse strigillosis 
scabrellis, subtus densissime brevitercue pubescentibus vel 
velutinosis; inflorescentiis terminslibus spicatis breviter 
pedunculatis; spicis densissime multifloris elongetis ubique 
dense puberulis; brscteolis anguste lanceoletis, ad eapicem 
longe setulosis.e 

Shrubby; branchlets acutely tetragonal, densely puberul- 
ent; nodes obscurely annulete with a longer band of pubes- 
cence; leaves opposite; netioles slender, 1.2--1-5 cme long, 
margined, deeply canaliculate above, densely puberulent; 
blades thin-chartaceous, brunnescent in drvinc, ovate-ellip- 
tic, +4.5--10 cm. long, 2--5 om. wide, acute or very snort- — 
acuminate at apex, regulerly serrulate from below the middle 
to the apex with very inuch appressed apiculate teeth, acum- 
inate at base, spersely strigillose and scabrellous above, 
very densely short-pubescent or ve'’utinous beneath; midrib 
slender, plane above, pvrominulous beneath; secondaries slen- 
der, 5--3 per side, arcuate-ascending, plane above, vrominu- 
lous above; veinlet reticulation obscure or indiscernible a= 
bove, plain beneath; inflorescence terminal, spicate; pedun- 
cles short, about 2.5 cm. long, densely puberulent; rachis 
medium-slender, not excavated, densely puberulent; florifer- 
ous portion of the spikes very densely many-flovered, to 22 
em. long, about 1 cm. wide, the flowers densely imbricete; 
bractlets nerrow-lanceolate, about 10 ma. long, long-setul- 
ose at apex, densely puberulent, ciliolate along the merg- 
ins; corolla-tube ebout 1.5 cm. long. 

The type of this species wes collected by Richard Spruce 
(no. 651) near Maypures, on the Orinoco River, Vichada, 
Colombia, in June, 1854, and is deposited in the Britton 
Herberium at the ew York Botcnical Garden. The species is 
named in honor of its distinruished and famous originel 
collector, whose name is so indelibly associated with the 


ee ee ae 


1940 Moldenke, Novelties 475 
netural history of South America. 


STACHYTARPHETA STRAIINEA Moldenke, sp. nov. 

Subligneous herb; stems more or less tetre7onal, fletten- 
ed at the nodes, rather densely puberulent, glabrescent in 
age; nodes not annulate; branches similar, slender, densely 
puberulent; principal internodes 3.5--6.5 cm. long; leaves 
decussate-opposite; petioles 0.6--2 cm. long, broadly alate, 
sparsely pilosulous-puberulent; blades thin-chartaceous, o- 
vate-elliptic or ovate, 35--5 cme long, 1.0--3 cm. wide, acu- 
te at apex, abruptly acuminete into the petiole at base, 
regularly serrate except et the very base with broadly tri- 
angular apiculete teeth, very minutely puberulent on both 
surfaces with obscure and scattered hairs on the lemina, 
more densely so on the lerger venation; midrib slender, 
plane above, very slightly prominulous beneath; secondaries 
slender, 3--6 per side, ascending, plane on both surfaces, 
ercuately joined near the mergins; veinlet reticulation ob- 
scure or plane on both surfaces; spikes terminal, 8--26 cm. 
long, slender, many-flowered, erect; peduncles very short, 
usually about 1 cm. long, canescent-puberulent; rechis slen- 
der, excavated, densely or sparsely canescent-puberulent ; 
flowers overlapping, closely appressed; bractlets lanceo- 
late, about 5 mae long, about 0.75 mm. wide at base, long- 
ettenuate to the setaceous apex, glabrous, usually stremin- 
eous, scarious-mergined on the lowest 1/3; calyx 4--5 m. 
long, stramineous, glabrous; corolla small, 8--10 mm. long. 

Herba sublignosa; remis densiuscule puberulis clabres- 
centibus; netiolis late alatis sparse pilosulo-puberulis; 
laminis ovetis vel ovato-ellipticis acutis, ad basim abrup- 
te acuminetis serretis utrinoue minute puberulis; brecteolis 
lanceolatis setaceo-attenuatis slabris stremineis. 

The type of this species was collected by José Quatrecas- 
as (no. 9627) somewhera in Colombie and is deposited in the 
Britton Herbarium at the New York Botanical Garden. The spe- 
cies is apparently closely related to the common and widely 
distributed S. cayennensis (L. ©. Rich.) Veht, from which 
its larger bractlets quickly distinguish it. 


STACHYTARPHETA SUBULATA Moldenke, spe nOve 

Herbe (2); ramulis grecillimis glebris; foliis oppositis 
Oblongis, ad apicem obtusis vel subacutis, ed basim longe 
cuneato~attenuatis, supra basim regulariter serratis, utrin- 
que glabris; petiolis obscuris vel alatis; inflorescentiis 
terminelibus spicatis ¢grecillimis laxe multifloris ubique 
glabris. 

Perennial herb (?); branchlets very slender, obtusely 
tetragonal or subterete, glabrate; nodes a.nulate; principal 
internodes ebout 5 cm. long; leaves decussate-opposite, usu- 


476 PHYTCLOGIA No. 14 


ally with some smaller ones on e very much abbreviated twig 
in their exils; petioles indistinct, merging into the leaf- 
bese, broadly alate, glebrate; blades membranous, oblong, 
rether uniformly zreen on both surfaces, 5--7 cme long, 1--2 
cme wide, obtuse or subacute et apex, long-cuneate and att- 
emiate at base, regularly serrate along the margins except- 
on the basal acumination, gl<ebrate on both surfaces; inflor- 
escence terminal, spicate, sessile, elongate, very slender, 
laxly meny-flovered; rachis very slender, shallowly excavat- 
ed, clzebrate; flowers barely imbricate; bractlets linear, a- 
bout 5 mm. lonz, subulete-setaceous at apex, glabrate. 
Tne type of th his species wes collected by Jacques Semuel 

Blanchet (no. 31392) in Bahia, Brazil, in or before 1840, 
and is dep s0sited in the herbarium of the Field Museum at 

Chicezo. 7 was identified as S. dichotoma (Rufz & Fav. ) 
Vehl end cited under tniis name by Scheuer in De Candolle's 
"Frodromus". S. dichotoma, however, is synonymous with S. 
ccyennensis and DSlanchet's plant is not that species. 


STACHYTARFEZTA TRINITENSIS Moldenke, spe nove 

Herba perennis; ramis fracilibus obtuse tetragonis densi- 
uscule adpresso-puberulis; foliis oppositis griseo-viridib- 
us; vetiolis indistinctis alatis; leminis ellipticis abrupte 
acutis vel obtusis serratis, ad basim longe attenuato- 
aciminatis, supra sparsissime albido-pilosis, subtus in ven- 
is majoribus puberulo-pilosis; spicis terminalibus elongatis 
mediocriter gracilibus; rhechide plusminus adpresso-piloso 
glebrescente; brecteolis megnis late ellipticis vel oblence- 
Oletis ciliatis, ad apicem scarioso-marginatis. 

Perennial herb; brenches slender, obtusely tetragonal, 
rether densely aprressed-puberulent with whitish hairs, more 
densely so at the apex and on youngest parts; nodes not ann- 
ulate; principal internodes 3--6 cm. long; leaves decussate- 
oprosite, usually with e fascicle of smaller ones on very 
much abbreviated twigs in their axils; petioles not dist- 
inct, « few mm. long, winged and merging into the leaf-base, 
puberulent; blades thin-chartaceous or membranous, uniformly 
crey-creen on both surfaces, elliptic, 3.5--8.5 cm. long, 
1./--3./7 cme wide, acute or obtuse at apex, rather uniformly 
serrate from almost the base of the expanded portion to the 
anex, lonz-acuminete at base and attenuate into the petiole, 
very sparsely white-pilose above, puberulent-pilose beneath 
especially along the larger venation; midrib slender, plane 
ebove, subprominulous beneeth; secondaries very slender, ab- 
out 5 per side, ercuate-ascending, plane above, very slight- 
ly prominulous beneath; veinlet reticulation sparse, indis- 
cernible ebove, mostly obscure beneath; inflorescence term- 
inal, spicate, elongate; peduncles greatly abbreviated, ab- 
out 2 cme long, densely eppressed-pubescent with short white 


ce 


es oe ee 


1940 Moldenke, Novelties 477 


hairs; rachis medium-slender, deeply excaveted in age, more 
or less cppressed-pilose, more densely so toward the base, 
glebrescent in age; bractlets lerzge, broadly elliptic or ob- 
lenceolate, 7--9 mm. long, 2--3 mm. wide, long-aristate- 
acuminate eat apex, zlebrous except for the ciliate margins, 
scarious and more or less lacerete along the margins from 
the widest pert to the base of the terminal aristula. 

The type of this species was collected by ‘alter Zlias 
Broadway et Tabacuite, Trinidad, on Sentember 20, 1918, end 
is deposited in the Britton Herbarium at the New York Botan- 
ical Gerden. The species is also lmown from Vlargerita Is- 
lend, Venezuela. 


STACHYTARPHETA WEBSRBAUERI lMoldenke, spe nove 

_ Fruticulus; ramis ramulisque acutissime tetragonis subal- 
atis densissime puberulis; nodis annuletis; foliis oppositis 
subsessilibus; leminis ellipticis acutis vel subecutis regu- 
leriter serruletis, ad basim longe acuminatis, utrinaue 
dense tomentellis; inflorescentiis terminalibus spicctis el- 
Ongetis densissime multifloris ubique breviter pubescentibus; 
rhachide excavato; bracteolis oblongo-elongatis. 

Shrubby; brenches and branchlets very sharply tetragonal, 
mergined or even subalate, very densely puberulent; nodes 
annulete; principal internodes 1--/+ em. long; leaves oppos- 
ite, sessile or subsessile; blades chartaceous, uniformly 
green On both surfeces or slightly lighter beneath, ellipt- 
ic, 2.2--4 om. long, 1--2 cm. wide, ccute or subacute at a- 
pex, long-acuminate at base, recularly serrulate except on 
the basal petiole-like ecumination, densely tomentellous on 
both surfaces; iiidrib slender, subimpressed above, prominul- 
ous beneath; secondcries slender, 5--7 per side, subimpress- 
ed above, prominu'ous beneath; veinlet reticuletion abund- 
ant, usuelly subimnressed above; inflorescence terminal, 
spicate; reduncles 0.5--2 cm. long, sharvly tetraronal or 
subelate, densely short-pubescent; rechis dee»ly excavated 
during and after anthesis, densely short-pubescent; florif- 
erous portion of srike 7--25 cme long, very densely many- 
flowered, €--10 mm. wide, the flowers closely imbricate; 
brectlets consnicuous, oblong-elongate, ubout 1 cm. long, 
densely short-pubescent, subulcte ct apex; corolla-about 2 
cm. long. 

The type of this species was collected by Ausust Weber- 
peuer (no. 6178) -- in whose honor it is named -- at an 
altitude of 1200 m., at Tabaconas on the “iaranon river, Caj- 
amarca, Feru, between 1909 and 1914, end is deposited in the 
Britton Eerbarium et the New York Botanical Garden. 


VERBSIA BALLSII ‘‘olcenke, cpe nov. 
Nerke csecpitosa, «ed basim lignose multo-ramosa; ramis 


478 PHYTOLOGIA Noe 14 


dense pubescentibus; foliis oppositis sessilibus; ‘taminis 
charteceis cuneato-obovatis trilobatis (lobis seepe trilobu- 
letis) utrinque dense pubescentibus; inflorescentiis termin- 
elibus magnis cepitatis; capitulis globosis densissime mult- 
ifloris; corollis rubellis; brscteolis linearibus elongatis 
dense pubescentibus. 

Dense meny-stemmed plants, woody at base, forming domed 
tufts to lm. ecross; stems simple or very spersely branch- 
ed, 15--22 cm. long, densely pubescent throughout; nodes en- 
nulete; principe! internodes 1--3 om. long; leaves opposite, 
sessile; bledes cnartaceous, cuneate-obovate, 1--35 cm. long; 
1--2.4 cm. wide, rounded in outline at apex and distinctly 
3-lobec (each lobe often ezein 3-lobed, oblong, blunt), 
long-cuneate to the base, densely short-pubescent with 
brownish or grsyisn often gland-tipped hairs on both surf- 
aces, less densely so in ege and then often with the pubes- 
cence beneath limited to the larger venation; midrib and 
secondaries impressed above, prominulous beneath; veinlet 
reticulation prectically indiscernible on either surface; 
inflorescence terminel, epparently compound, but forming an 
extremely lerge and dense slobuler many-flowered head 5.5--S 
cme in diameter; peduncles obsolete or very short; brectlets 
numerous, linear, about 1 cm. long, very densely spreading- 
pubescent; celyx tubular, about © mm. long, densely spread- 
ing-pubescent with whitish gland-tipped hairs; corolle 
creamy-pink, sweet-scented, its tube about 1 cm. long, its 
limb about 6 mm. in diameter. 

The type of this showy and most distinctive species was 
collected by Zdward K. Balls (no. 6036) -- in whose honor it 
is named -- in the Chorru valley, neer Tilcara, et an altit- 
ude of 12,800 feet, Jujuy, Argentine, on February 13, 1959, 
end is deposited in tre United States National Herbarium at 
Yashington. The collector states that it grows among rocks 
on dry sunny exposures in shaley formations well above the 
moisture line and thet its leaves ere gray, softly hairy, 
loosely clothing the stems. 


VERBENA CABRERAE lMoldenke, spe nove 

Herba; ceulibus ramisaue srecilibus obtuse tetregonis ad- 
presso-pilosis; foliis oppositis; petiolis alatis albido- 
strigosis; leminis tripartitis, partibus laciniatis vel 
pinnetifidis, leciniis acutis, utrinque strigoso-pilosis; 
inflorescentiis terminalibus spicatis, spicis abbreviatis 
congestis. 

Herb; stems and brenches slender, obtusely tetragonal, 
rether sparsely appressed-pilose with whitish hairs, more 
densely so toward the apex and on younger parts; nodes eannu- 
late; principal internodes 2--4 cm. long; leaves decussate- 
opposite, usually with several smaller ones on very much ab- 


1940 Moldenke, Novelties 479 


breviated twigs in their axils; petioles slender, 5--10 mn. 
long, winged, white-strigose; blades very thin-chartaceous, 
uniformly green on both surfaces, ovate in outline, 4--6 cm. 
long and wide, deeply 3-parted to the base, the divisions 
laciniate or pinnatifid, the lobes/oblong and acute (often 
sharply so), strigose-pilose with appressed whitish hairs on 
both surfaces; midrib and secondaries plane or subimpressed 
above, prominulous beneath; veinlet reticulation indiscern-- 
ible on both surfaces; inflorescence terminal; peduncles 
very slender, appressed-strigose, 2.5--4.5 cm. long; spikes 
abbreviated, congested, many-flowered; calyx narrow-tubular, 
about 9 mm. long and 1 mm. wide, densely white-strigose; 
corolla-tube about 12 mm. long, its limb 7--9 mm. wide,blue. 

The type of this species was collected by Angel L. Cabre- 
ra (no. 4199) -- in whose honor it is named -- at Quebrada 
del Rio Caraparf, department of Oran, Salta, Argentina, on 
July 15, 1937, and is deposited in the Britton Herbarium at 
the New York Botanical Garden. It is known also from Tucum- 
én and is a most distinctive species. 


VERBENA OCCIDENTALIS Moldenke, sp. nov. 

Rerba; ramis simplicibus erectis dense hirsuto-pubescent- 
ibus, pilis griseis glanduliferis; foliis oppositis fascicu- 
latis; petiolis obsoletis vel brevissimis et alatis; laminis 
chertaceis 3-partitis profunde pinnatifidis utrinque dense 
griseo-hirsutulis, marginibus subrevolutis. 

Herb; stems simple, erect, slender, tetragonal, densely 
hirsute-pubescent with sordid-grayish gland-tinped hair; 
principal internodes 1.5-~3 cm. long; leaves decussate- 
opposite, usually with a few smaller ones fascicled in their 
axils; petioles obsolete or very short and winged; blades 
chartaceous, uniformly gray-green on both surfaces, 3-perted 
almost to the winged petiole-like base, each division again 
deeply pinnetifid, densely hirsutulous-pubescent on both 
surfaces with grayish hair like that on the stems, sometimes 
substrigose above, the margins slightly revolute; midrib and 
seconderies very slender, impressed above, prominulous be- 
neath; veinlet reticulation indiscernible on both surfaces; 
spike terminel, single, simple, unbranched, 7--8 or more cm. 
long, very densely many-flowered; peduncles slender, about 
325 cme long, densely hirsute-pubescent like the stems; flo- 
wers very densely imbricate (except the lowest 2 or 3, which 
may be erranged in ea whorl somewhat separated from the rest 
of the nici Or more separated so as to be barely imbricate 
and somewhat divergent in fruit; bractlets large, linear or 
narrow-lenceolate, 5--8 mm. long, less than 1 mm. wide, 
densely hirsutulous with gland-tipped hairs like the pedunc- 
les; calyx tubular, about & mm. long and 2 mm. wide, S-cost- 
ate, very thin-textured between the ribs, densely hirsutul- 


480 PHYTOLOGIA Yo. 14 


ous with spreading whitish gland-tipped hairs, its rim 5- 
apiculate; corolla hypocrateriform, its tube ebout 1 cm. 
longe 

The type of this species was collected by Andrew Mathews 
(no. 198) somewhere in Peru in 1834 and is deposited in the 
Meisner Herbarium et the New York Botanical Garden 


VERBENA REINECKII Moldenke, spe nov. 

Herba; caulibus ramisque gracilibus plusminus hirsutulis 
et puberulis; foliis oppositis breviter petioletis; laminis 
ovatis vel ovato-ellipticis profunde laciniatis vel lobatis 
(lobis obtusis latis), ad apicem subacutis, ad basim in pet- 
jolum cuneato-attenuatis, supra sparse scabro-pilosis, sub- 
tus in venetio hirsutulis; inflorescentiis axillaribus term- 
inelibusque subcepitatis hirsutulis. 

Herb; stems end branches slender, more or less hirsutul- 
ous end puberulent, less so in age; principal internodes 1-- 
5 em. long; leaves opposite; petioles very slender, to 5 mm. 
long end sparsely hirsutulous, or obsolete; blades chartace- 
ous, rether uniformly bright-green on both surfaces, ovate 
or ovate-elliptic, 0.5--2.5 cm. long, 0.3--2 cm. wide, sub- 
acute at apex, cuneate-ettenuate into the petiole at base, 
deeply leciniate or lobate with broad obtuse lobes, sparsely 
scabrous-pilose with bulbous-based hairs above, hirsutulous 
along the venation end margins beneath; midrib and the 2 or 
3 secondaries very slender, impressed above, sharply promin- 
ulous beneath; veinlet reticulation very srarse, usually ob- 
scure; inflorescence axillary and terminal, subcapitate- 
spicate, 8--15-flowered; peduncles elender, hirsutulous, 1-- 
2.5 cme long; bractlets much shorter than the calyx, lanceo- 
late, about 4 mm. long and 1 mm. wide, hirsutulous, ettenua- 
te at apex; calyx tubular, about 9 mm. long, hirsutulous, 
the hairs sometimes gland-tipped; corolla-tube ebout 1.5 om. 
long, its limb to 1 cm. in diameter. 

The type of this species was collected by Sduard Martin 
Reineck -- in whose honor it is named -- and Josef Czermak 
(no. 21) on a sunny hill near Moinhos do Vento, Forto Aleg- 
re, Rio Grande do Sul, Brazil, in November, 1897, and is 
deposited in the herbarium of Pomona College at Claremont, 
California. The species has been hitherto confused with 
with "Verbena chemaedryfolia Juss.", which, however, is con- 
specific with V. peruviana (L.) Britton and is an entirely 
different plant with brilliant scarlet flowers and quite 
different foliar characters. 


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| a 
 PHYTOLOGIA 
Designed to expedite botanical publication 
Vor. 1 January, 1941 No. 15 
CONTENTS 
C, L. Lunpett: Three new trees from Tabasco and Campethe..... g8I 


H. N. Moxpenxe: New South American Verbenaceae.....+++++ GBF 
CGR IAD E ONUTRE ODE gh) «ial 5, -os¥ ages wm 0 6 ae bie. Will O Wt ell in pwede We 491 


PR RE ROMP R UPS So tah ety Kak Apa la aus ching eS abso aw Skee ee 512 


Published by H. A. Gleason and Harold N. Moldenke 
. The New York Botanical Garden 
Bronx Park, New York, N. Y. 


Price of this number, 50 cents; per volume, $5.00 in advance 


Vol. 1, No. 14, was issued November 25, 1940 


Mwy tt’ 


THREE NEW TREES FROM TABASCO AND CAMPECHE (a) 


C. Le Lundell 


PITHECOLOBIUM CAMPECHENSE Lundell, sp. nov. 

Arbor. Stipulae subulatae, usque ad 3 mm. longae. Petioli 
usque ad 2.4 cme longi. Pinnae l-jugae. Foliola l-juga, ob- 
longa, chartacea, basi barbata, apice rotundata vel emargin- 
ata. Inflorescentiae spicatae, adpresse puberulae; bracteae 
minutae. Calyx 1--1.3 mm. longus. Corolla 4 mm. longa. 

A tree 7 to 10 m. high, 30 to 60 cm. in diam., branchlets 
rather slender, glabrous, striate. Stipular spines minute, 
stout, less than 3 mme longe Petioles up to 24 cme long, 
often less than 1 cme long, glabrous, slender, with a disc- 
oid apical gland. Pinnae 1 pair. Leaflets l-pair, obliquely 
oblong, 2 to 4.3 cm. long, up to 1.9 cm. wide, chartaceous, 
barbate at base beneath, entirely glabrous otherwise, apex 
rounded or emarginate, both surfaces reticulate-veined. 
Spikes up to 4 cm. long, borne in panicles, puberulent; 
bractlets minute, less than half the length of calyx, pers- 
istent. Tlowers sessile. Calyx appressed-puberulent, 1 to 
1.3 mm. long. Corolla appressed-puberulent, 4 mm. long. 
Stamen-tube shortly exserted. Ovary subsessile, covered with 
appressed hairs. 

MEXICO: Campeche, Palizada, in "swamp side forest", July 
25--28, 1939, Eizi Matuda 3866 (type in the University of 
Michigan Herbarium); vernacular name "tinta". 

From P. lanceolatum (He & B.) Benth., to which it is al- 
lied, Pe " campechense mpechense differs in having minute stipuler 
spines, | much shorter flower spikes, and smaller calyx and 
corolla. The fruits are not known. 


EUGENIA BALANCANENSIS Lundell, sp. nov. 

Arbor parva, glabrae Folia petiolata, chartacea, ovato- 
lanceolata, apice attenuata, obtuse acuminata, basi acuta. 
Inflorescentiae racemosae, axillares, usque ad 1-7 cme long- 
ae. Pedicelli usque ad 6 mm. longi. Petala oblonga. 

Tree, 4 m. high, 15 cm. diam., glabrous throughout ex- 
cepting the sparsely puberulent buds. 3ranchlets slender, 
compressed at first, glandular-punctate. Petioles short, 
canaliculate, 3 to 6 mm. long. Leaf-blades thin, chartace- 
ous, punctate, concolorous or slightly paler beneath, ovate- 
lanceolate, 4 to 9.5 cm. long, 1.3 to 3-5 cme wide, apex at- 
tenuate, obtusely acuminate, base acute, costa nearly plane 
above, inconspicuously elevated beneath, the main lateral 
veins 6 to 10 on each side, anastomosing into a submarginal 
vein, prominulous beneath. Inflorescence racemose, the rac- 

481 


482 PHYTOLOGIA No. 15 


emes axillary, solitary or fasciculate, up to 1.7 cme long 
with a short rachis; the rachis and pedicels glandular- 
punctate. Pedicels slender, up to 6 mm. long. Salyx small, 
the tube subglobose, less than 1 mm. long, 4-lobed, the 
lobes rounded, unequal, the larger up to 1.5 mm. long, minu- 
tely ciliolate. Petals oblong, up to 3.2 mm. long, conspicu- 
ously glandular-punctate. 

MEXICO: Tabasco, San Isidro near 3alancan, in advanced 
forest, June 7--11, 1939, Eizi Matuda 3361 (type in the Uni- 
versity of Michigan Herbarium). 


EUGENIA TABASCENSIS Lundell, spe nov. 

Arbor parva, ramulis albido-sericeis. Folia petiolata, 
chartacea, anguste elliptico-oblonga, apice caudato-acumin- 
ata, basi cuneata, sericea vel subglabra. Inflorescentiae 
racemosae, axillares, fasciculatae, usque ad 4.5 cm. longae. 
Pedicelli usque ad 6-5 mm. longi. Fetala obovata vel ellip-— 
tica, 2.6--3.-4 ma. longa, ciliolata. 

A small tree. Branchlets slender, whitish-sericeous, at 
first compressed and sulcate at the nodes, reddish. Leaves 
attenuate at the base into short, slender, canaliculate, 
sparingly sericeous petioles up to 5 mm. long. Leaf-blades 
chartaceous, narrowly elliptic-oblong, 3 to 7.5 cm. long, 1 
to 3 cme wide, apex caudate-acuminate, the cusp obtuse, base 
cuneate, punctate, sparingly sericeous on both surfaces at 
first, at length with only a few hairs persisting along the 
costa, the midvein impressed above, elevated beneath, the 
lateral veins numerous, approximate, prominulous below, less 
evident above, reticulate, mercing into a prominulous sub- 
marginal vein. Inflorescence densely sericeous, racemose, 
the racemes axillary, usually fasciculate, up to 4.5 cm. 
long, sessile or pedunculate, the rachis slender. Pedicels 
slender, up to 6.5 mm. long. Calyx whitish-sericeous, 4- 
lobed, the lobes sparingly puberulent on both surfaces, 1 to 
1.3 ma. long. Petals obovate or elliptic, 2.6 to 3.4 mm. 
long, ciliolate, glabrous otherwise, punctate; receptacle 
puberulent. 

MEXICO: Tabasco, Boca Cerro on the Usumacinta River above 
Tenosique, on limestone, July 1--5, 1939, Eizi Matuda 3561 
(type in the University of Michigan Herbarium). 

The species is related to Eugenia yucatanensis Standl. 
and E. cocquericotensis Lundell. 


ee a 


NEW SOUTH AMERICAN VERBENACEAE 


Harold Ne Moldenke 


DURANTA REPENS vare MICROPHYLLA (Desf.) Moldenke, comb. nov. 
Durante microphylla Desf., Cat. Hort. Paris, ede 3, 392.. 
1829. 


JUNELLIA STRAGULOIDES Moldenke, spe nove 

Fruticulus nanus caespitosus; caulis abbreviatis lignosis 
ramis gracilibus abbreviatis puberulis; internodiis valde 
abbreviatis; foliis oppositis rigidis tripartitis amplexi- 
caulibus, lobis patentibus puberulis argute acutis integris; 
inflorescentiis unifloris; corollis rubellis. 

Diminutive alpine shrub, forming small rosettes, pressed 
together into dense wood-hard cushions to 1 me in diameter, 
usually flattened, sometimes mounding slightly in the cent- 
er; stems abbreviated, woody, prostrate; branches slender, 
abbreviated, prostrate or decumbent, puberulent; internodes 
greatly abbreviated; leaves opposite, rigid, uniformly 
yellow-green on both surfaces, tri-partite, clasping at 
base, the divisions oblong, 2—5 mm. long, sharply acute at 
apex, divergent, puberulent, entire, usually obdscurely vein- 
ed; inflorescence reduced to solitary axillary flowers, 
borne in great abundance; calyx cupuliform, about 2 mm 
long, white-puberulent or strigillose; corolla pale-pink, 
its tube 4-—-5 mn. long, its limb about 4 mm. in diameter. 

The type of this species was collected by Edward Ke Balls 
(noe 6000) in screen-like open patches among upland grasses 
on rolling non-calcareous soil in valleys above San Gregor- 
io, Tilcara, at an altitude of 14,000 feet, Jujuy, Argent- 
ina, on February 11, 1939, and is deposited in the United 
States National Herbarium at Washington. 


VERBENA PARAGUARIENSIS Moldenke, spe. nove 

Herba; caulibus simplicibus erectis albido-strisosis; 
foliis oppositis sessilibus ad caulem arcte adpressis; lam- 
inis lanceoletis attenuatis acutis integris subrevolutis, ad 
basin rotundatis vel obtusis, utrinque strigosis; inflores- 
centiis terminalibus spicatis multifloris, floribus alternis 
imbricatis. 

Herb, to 6 dm tall; stems usually simple and erect, 
strigose with closely appressed whitish antrorse hairs; 
leaves decussate-opposite, numerous, appressed to the stems, 
sessile; blades lanceolate, 6--23 mm. long, 1--3 mm. wide, 
attenuate above to the rather sharply acute apex, rounded or 
obtuse at base, entire and usually subrevolute along the 


483 


484 ra. -T:O tL: U's Bak Noe 15 


margins, strigose on both surfaces like the stems; inflores—- 
cence terminal, spicate, terminating the stems, solitary 
10--20 cme long, about 1 cm. wide throughout, many-flowered, 
the flowers’alternate, imbricate, somewhat spreading; rachis 
slender, more or less flexuous, strigose like the stems; 
bractlets 2--4 mm. long, closely appressed to the calyx, 
strigose. 

The type of this peculier species was collected by Teod- 
oro Rojas (Hassler 9751) on high plateau and declevities, 
Sierra de Amambay, Paraguay, in December, 1907, and is dep- 
osited in the Delessert Herbarium of the Conservatoire 3ot- 
anique at Genevae 


VITEX APPUNI Moldenke, Geogr. Distrib. 21 % 26, nom. nud. 

(1939), sp. nove 

Frutex vel arbor; ramlis sarmentisque obtuse tetragonis 
vel subteretibus medullosis dense breviterque pubescentibus 
subglabrescentibus; foliis 3- vel 5-foliolatis, plerumque 3- 
Poliolatis; petiolis gracillimis dense breviterque pubescen- 
tibus; foliolis sessilibus submembranaceis ellipticis vel 
sudobovato-ellipticis, ad basin et apicem acuminatis, inte g- 
ris vel crasse gerratis, supra dense puberulis, subtus den- 
sissime pubescentibus vel! tomentellis; inflorescentiis ax- 
illaribus capitatis dense m:ltifloris. 

Shrub or tree; branchlets and twigs medium or slender, 
obtusely tetragonal or subterete, medullose, gray—brown, the 
youngest shoots densely short-pubescent with cinereous pub- 
escence, the older ones sparsely puberulent or subglabrate; 
leaf-scars on older wood rather large and semi-circular, us-— 
ually with elevated margins; nodes not annulate; principal 
internodes 1.5--6 cme long or more abbreviated on short ax- 
illary twigs; leaves decussate-opposite, 3— or 5-foliolate, 
mostly 3-foliolate; petioles very slender, 1.2--6 cm. long; 
convex beneath, flattened above, densely short—pubescent 
like the young twigs; leaflets, when 3, subequal in size or 
the lateral ones slightly smaller, when 5, the 2 lowermost 
often mich reduced. all sessile or the central ones only 
subsessile; leaflet-blades submembranous, rather dark-green 
above, much lighter beneath, the central one elliptic or 
subobovate-elliptic, 3--9 cm. long, 1.5--3 cm. wide, acumin- 
ate at apex and base (the lateral ones usually acute at 
base), entire or coarsely and irregularly serrate with rath- 
er few blunt teeth, densely puberulent above, becoming uch 
more sparsely so in age, very densely short-pubescent or 
tomentellous beneath; midrib very slender, subimpressed ab-— 
ove, prominulous beneath; secondaries very slender, 9--11 
per side, flat (and often inconspicuous) or subimpressed a- 
bove, prominulous beneath, arcuate-ascending, not plainly 
anastonosing; vein and veinlet reticulation iostly obscure 


1941 Moldenke, South American Verbenaceae 485 


on both surfaces; inflorescence axillary, cymose, capitate, 
3—--7 cme long, 1--2.5 cme wide, densely many-flowered; ped- 
uncles slender, 2-—5.e5 cme long, more sparsely short-pubes- 
cent than the young twigs, flattened; pedicels very slender, 
1--3 mm. long; bracts none; bractlets few, linear, 2-—4 mm. 
long, densely pubescent; prophylla minute, setaceous, 1 mm. 
long or less, densely pubescent. 

The type of this species was collected by Carl Friedrich 
Appun (no. 1885) -- in whose honor it is named -- probably 
on or near Mount Roraima, British Guiana, and is deposited 
in the herbarium of the Royal Botanic Gardens at Kew. The 
species is known also from Amazonas, Brazile 


VITEX FROESII Moldenke, sp. nov. 

Arbor; ramulis glabrescentibus; hornotinis crassis dens- 
issime villoso-tomentosis, pilis fulvis; petiolis crassis 
densissime fulvo-villosis; foliolis 4 sessilibus subcoria- 
ceis late ellipticis vel obovatis, ad apicem rotundatis vel 
subcuspidatis integris utrinque dense villoso-velutinis; 
inflorescentiis glomeratis sessilibus bracteosis densissime 
longeque villosis. 

Tree, to 15 me tall; trunk to 18 cm. in diameter; 
branchlets medium-stout, gray, obtusely tetragonal, glab- 
rescent; young twigs stout, obtusely tetragonal, very den- 
sely villous-tomentose with long fulvous hairs; buds dense- 
ly long-villous; nodes somewhat enlarged and ampliate, not 
annulate; principal internodes 2.5--8 cme long; leaves de- 
cussate-opposite, trifoliolate; petioles stout, 4.5--11 cm. 
long, very densely villous-tomentose with fulvous hair sim- 
ilar to that on the twigs; petiolules absent; leaflets 3, 
sessile, subcoriaceous, broadly elliptic or obovate, light- 
er beneath, the central one 8=--18.5 cme long, 5--12e3 cme 
wide, rounded or very slightly cuspidate at apex, entire, 
slightly undulate or sinuate, cuneate and symmetrical at 
base, densely villous-velutinous on both surfaces, more 
conspicuously so beneath, with flavescent hairs; midrib 
stoutish, prominulous (except toward the apex) above, very 
prominent beneath; secondaries slender, 10--15 per side, 
ascending, not much arcuate (except toward the margins), 
close together, joined in many loops at the margins, very 
slightly impressed above, prominent beneath; veinlet retic- 
ulation abundant, slightly ispressed above, prominulous be- 
neath; lateral leaflets similar, practically the same size 
as the central one or slightly smaller, but usually asymm 
etric at base or asymmetric throughout, often rounded or 
subtruncate at base, or the outer half broadly rounded and 
the inner half cuneate at base; inflorescence glomerate, 
sessile, bracteose, very densely long-villous with fulvous 
hairs; bracts numerous, lanceolate, 1.8--2.e3 cme long end 


486 PAY 1:03.40 GA Noe 15 


2--5 mme wide at base, sessile, densely villous on the outer 
surface, concave from within, an opposite pair closely app-— 
ressed to the base of the calyx and alternate with its lips; 
calyx tubular, about 1.8 cm. long and 5 mm. wide, distinctly 
2-lipped, one lip 2-lobed at apex, the other 3-lobed, the 
lobes 1-—-4 mm. long, slender, triangular~acuminate, the sin- 
uses between the lips about 5 mm. deep, in fruit greatly en- 
larged and splitting to the base, the 2 halves broadly ov- 
ate-elliptic, to 2.5 cme long and 1.7 cme wide, the lobes 
5--7 mme long; corolla and fruit not seene 

The type of this most remarkable and distinctive species 
was collected by Re Frées (no. 11,660) -- in whose honor it 
is named —- on high land in a high forest, terra firma along 
the Rio Pindare, Rapoza Moncg&o, Maranhfo, Brazil, on Decemb-— 
er 15, 1937, and is deposited in the Britton Herbarium at 
the New York Botanical Garden. 


VITEX GUIANENSIS Moldenke, Alph. List Common Names 14, nom. 
nud. (1939); Geogr. Distrib. 21, nome nud. (1939), spe 
nove 
Arbor alta; ramulis gracilibus obtuse tetragonis dense 

breviterque rufescento-tomentosis; foliis 5-foliolatis; pet- 

iolis gracillimis dense rufescento-tomentosis; foliolis 
stipitatis ellipticis acutis vel breviter acuminatis integ- 
ris vel obscure repando-undulatis, ed basin acutis vel brev- 
iter acuminatis, supra sparse ferrugineo-hirsutulis, subtus 
dense hirsutulis; inflorescentiis axillaribus abbreviatise 
Tree, to 90 feet tall, slightly buttressed, inclined to 
be fluted, with pale-gray thin papery bark; branchlets rath- 
er slender, obtusely tetragonal, densely tomentose with ruf- 
escent rather short hairs, the tomentum wearing off in age; 
nodes annulate, flattened; principal internodes 2--6 cm. 
long; leaf-scars rether large and prominent; leaves decidu- 
ous, strongly scented, 5-foliolate; petioles very slender, 
4--10 (--18.5 1) cme long, densely rufescent-tomentose (in 
age stout and glabrous ? -- one old petiole, without leaf- 
lets, is mounted on an isotype, but may not belong to this 
species); leaflets thin-chartaceous or submembranous, dark- 
green above, slightly lighter beneath, unequal, the 2 lower- 
most mostly considerably smaller than thé rest, short-petio- 
lulate; petiolules very slender, 1.5--4 mm. long, densely 

rufescent-tomentose; central leaflet-blades elliptic, 8.6-- 

15-5 cme long, 301--6.8 cm. wide, acute or short-acuminate 

at apex, entire or obscurely repand-undulate, ecute or 

short-acuminate at base, sparsely hirsutulous above with 

scattered ferruginous hairs, very densely so along the mid- 

rib and lerger veins, somewhat more densely hirsutulous be- 

neath, especially along the larger venation; midrib slender, 

flat above, prominent beneath; secondaries very slender, 10- 


oo 


1941 Moldenke, South American Verbenaceae 487 


15 per side, arcuate-ascending, mostly flat and rather ob- 
scure above, prominulous beneath, anastomosing in many loops 
(indistinctly) at the margins; veinlet reticulatio indis- 
cernible above, the lerger portions slightly prominulous be- 
neath; inflorescence axillary; cymes abbreviated, much 
shorter than the petioles; peduncles, pedicels, end young 
shoots covered with a brown felt of hairs; flowers in erect 
umbelloid cymes; calyx pale-green, erecto-patent; corolla 
bilabiate, the upper lip composed of 2 white reflexed lobes, 
the lower lip composed of 3 blue-purple lobes, with a patch 
of pale-yellow hairs at the base of the central lobe, re- 
flexed; corolla-tube blue-striate; filaments up to 1/4 long- 
er than the corolla-tube, flushed with purple; enthers blue. 

The type of this species was collected in baniaballi 
(myrtaceous) bush on the top of an Ironstone Hill where 
there was no uniform soil covering, but only soil pockets, 
British Guiana (British Guiana Forestry Department 2543), on 
October 28, 1937, and is deposited in the herbarium of the 
Royal Botanic Gardens at Kewe The native Arawak name is 
"hakiaballi". 


VITEX KRUKOVII Moldenke, Geogr. Distribe 27, nome nud. 

(1939), sp. nove 

Arbor; ramilis compresso-tetragonis sparsissime pilosis; 
sarmentis densissime et valde conspicue villoso-tomentosis, 
pilis fulvis; foliis 5-foliolatis; petiolis gracilibus dens- 
issime villoso-tomentosis; foliolis stipitatis late ellipti- 
cis vel subobovatis breviter acuminatis integris, ad basin 
acuminatis vel cuneatis, supra in costa villosis, subtus in 
venis majoribus fulvo-villosis; inflorescentiis axillaribus 
capitatis 2- vel 3-floribus. 

Tree, about 50 feet tall; branchlets dark, flattened- 
tetragonal, very sparsely pilose with scattered hairs; twigs 
slender, extremely densely and conspicuously villous- 
tomentose with long fulvous hairs; leaves decusaate-oppos— 
ite, 5-foliolate; petioles slender, 6.5--10 cm. long, 
rounded beneath, flat above, very densely villous-tomentose 
with fulvous hairs, slightly ampliate at apex; leaflets 
mostly symmetrical, the central noticeably larger than the 
two lowest and borne on longer petiolules, young leaves with 
all the petiolules obscure; petiolules 4--11 mm. long, very 
densely villous-tomentose with fulvous hairs; central leaf- 
let-blades broadly elliptic or slightly obovate, 14--19 cm. 
long, 6-5--9 cme wide, short-acuminate at apex, entire, ac- 
uminate or cuneate at base, villous along the midrib above 
and along the midrib and secondaries beneath, with long ful- 
vous hairs, often also very sparsely so on the veinlet- 
reticulation beneath, basal leaflets similar but smaller, 
6--9.5 cme long, 2e8--4.5 cme wide; midrib somewhat impress- 


488 PHYTOLOGIA No. 15 


ed above, very prominent beneath; secondaries 10—-15 on each 
side, close together, rather straight, abruptly arcuate- 
joined near the margins; inflorescence axillary, solitary, 
opposite, cymose; cymes capitate, apparently 2- or 3-flower+ 
ed, 9--9.5 cm. long; peduncles slender, erect, divergent, 
7--8 cme long, densely villous-tomentose with long fulvous 
hairs (1—2 mm. long, like on the twigs and petioles), appa- 
rently bearing a few bractlets at the apex; fruiting—pedi- 
cels obsolete; fruiting-calyx cupuliform, large, about 9 mm. 
long and 12 mm. wide, densely strigose-villous with closely 
appressed rather long flavescent hairs, its rim deeply 5- 


lobed, the lobes triengular-ovate, long, rather narrow, ac- 


ute, erect; fruit ellipsoid or rounded, about 14 mm. long 
and wide, extremely densely and beautifully villous with > 
long (2--3 mm.) fulvous or brown glistening hairs, all for- 
ward=pointing and appressed. 

The type of this handsome species was collected by Boris 
Alexander Krukoff (no. 6279) -- in whose honor it is named 
-- on "varzea" land near Tres Casas, Municipality Humayta, 
Amazonas, Brazil, between September 14 and October 11, 1934, 
and is deposited in the Britton Herbarium at the New York 
Botanical Garden. 


VITEX MARTII Moldenke, Geogr. Distrib. 27, nom. nud. (1939), 

Sp. NOVe 

Arbor; ramis ramulisque gracilibus obtuse tetragonis 
dense puberulis glabrescentibus; sarmentis gracillimis ab- 
breviatis densissime fulvo- vel ferrugineo-puberulis; foliis 
3--5-foliolatis; petiolis gracillimis dense breviterque 
fulvo- vel flavescento-pubescentibus; foliolis sessilibus 
vel subsessilibus chartaceis ellipticis argute acutis integ- 
ris, ad basin acutis, supra leviter puberulis, subtus dens- 
iuscule puberulis; inflorescentiis axillaribus capitatis 
dense multifloris. 

Tree; branches and branchlets slender, grayish, obtusely 
tetragonal, somewhat swollen at the nodes, densely puberu- 
lent with extremely fine matted grayish puberulence, becom- 
ing subglabrate in age, not noticeably lenticellate; twigs 
very slender, mostly abbreviated, very densely puberulent 
with fulvous or ferruginous hairs; nodes not annulate; 
principal internodes 1--2.8 cme long or on young twigs ex- 
tremely abbreviated (to 1 mm.); leaf-scars large, corky, 
prominent; leaves decussate-opposite, 3--5-foliolate; petio- 
les very slender, 0.6--2.8 cm. long (at anthesis), densely 
short-pubescent with flavescent or fulvous hairs, flattened 
above, not noticeably ampliate at base; leaflets sessile or 
subsessile (at anthesis); leaflet-blades (at anthesis) char- 
taceous, rather uniformly bright-green on both surfaces or 
somewhat lighter beneath, the terminal one elliptic, 1.5-- 


1941 Moldenke, South American Verbenacese 489 


305 cme long, 6--11.5 mm. wide, sharply acute at apex, en- 
tire, acute at base, finely puberulent above, somewhat more 
densely so beneath, mostly conduplicate and reflexed; midrib 
very slender, plane above, slightly prominulous beneath; 
secondaries very slender, about 10 per side, ascending, 
short, not much arcuate; vein and veinlet reticulation very 
fine, indiscernible above, plane beneath; inflorescence ax- 
illary, capitate, 2--5 cm. long, 6--15 mm. wide, densely 
many-flowered, not brachiate, appearing before or with the 
leaves; peduncles very slender, 1.6--4 cm. long, densely 
short=-pubescent or puberulent with flavescent or fulvous 
hairs. 

The type of this species was collected by Carl Friedrich 
Philipp von Martius in the woods of the Catingas along the 
Rio Sao Francisco Salgado, Minas Geraes, Brazil, and is dep- 
osited in the herbarium of the Botanisches Museum at Munich. 
The species is known also from Bahia. 


VITEX POLYGAMA var. BAKERI Moldenke, Alphe List Common Names 

20, nome nud. (1939); Geogr. Distrib. 27 & 40, nom. nud. 
(1939), vare nov. 

Haec varietas a forma typica speciei recedit foliolis 5 
sessilibus, ad basin longe acuminatis vel attenuatis; pilis 
ubique sericeo-velutinis et ferrugineis vel hirsutis. 

This variety differs from the typical form of the species 
in its five leaflets being sessile and long-acuminate or at- 
tenuate at base, and the pubescence throughout being usually 
more silky-velutinous and ferruginous or hirsute. 

The type was collected by Charles Fuller Baker (no. 281) 
-- in whose honor it is named -- in the vicinity of Pard, 
Perd, Brazil, on February 20, 1908, and is deposited in the 
herbarium of the Botanisch Museum at Utrecht. The variety is 
also known from Maranh&o and from cultivation. 


VITEX POLYGAMA vare DUSENII Moldenke, Geogr. Distrib. 27, 
nome nude (1939); Prelim. List Invalid Names 51, hyponym 
(1940), vare nov. 

Haec varietas a forma typica speciei recedit bracteolis 
foliaceis, cymis dense congestis, lobis calycis longissimis 
foliaceis bracteoloideis. 

This variety differs from the typical form of the species 
in its extremely long and foliaceous calyx-lobes, which 
greatly resemble the foliaceous bractlets which surround 
them in the dense and congested cymes. 

The type was collected by Per Karl Hjalmar Dusén (no. 
16,111) at the margins of primeval woods in the littoral 

. 4 
region at Jacarehy, Parand, Brazil, on December 13, 1914, 
end is deposited in the herbarium of the Naturhistoriska 
Riksmseet at Stockholm. The variety is known also from Rio 


490 PHYTOLOGIA Noe 15 
de Janeiro. 


VITEX POLYGAMA var. GLAZIOVII Moldenke, Geogr. Distrib. 27, 
nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit lobis caly- 
cis angustissimis elongatis sublinearibus vel oblongis 
bracteoloideis, fructibus valde carnosis, endocarpio cent— 
rale parvo. 

This variety differs from the typical form of the species 
in its very fleshy fruits and small central endocarp, the 
fleshy portion being almost 2 cme long and wide and the 
stony endocarp only about 11 mm. long and 6 mm. wide. The 
celyx-lobes are also very narrow and elongate, sublinear or 
oblong, 5--10 mm. long, resembling the bractlets. 

The type was collected by Auguste Frangois Marie Glaziou 
(noe 5959) -- in whose honor it is named -- at Restinga de 
Mau&, in the Organ Momtains, Rio de Janeiro, Brazil, be- 
tween T1861 and 1887, and is deposited in the herbarium of 
the Royal Botanic Gardens at Kew. 


VITEX POLYGAMA vare WARMINGII Moldenke, Geogre Distrib. 27, 
nome nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit foliis sub 
anthesin valde immaturis cymis saequantibus, pilis conspicue 
flavescentibus vel ferrugineis et velutino-sericeis. 

This variety differs from the typical form of the species 
in its leaves being very immature at the time of flowering 
and equaled in length by the cymes, and the pubescence 
throughout being very conspicuously flavescent or ferrugin- 
ous and velutinous-silky, especially on the leaves, where it 
is very lustrous. 

The — was collected by Johannes Eugenius Bulow Yarming 
(noe 23/967) -- in whose honor it is named -- at Lagoa Sant- 
a, Minas Geraes, Brazil, and is deposited in the herbarium 
of the Universitetets Botanisk Museum at Copenhagen. 


VITEX SCHOMBURGKIANA var. GRANDIFLORA Moldenke, Geogr. Dist- 

rib. 27, nom. nud. (1939), var. nov. 

Haec varietas a forma typica speciei recedit floribus ma- 
joribus et cymis brachiatis. 

This variety differs from the typical form of the species 
in its large flowers (tne corolla-tube 7--8 or more mm 
long, its limb about 10 mm. wide) and brachiate cymes, the 
cyme—branches often to 2 cme long and very divergent. 

The type of this variety was collected by Gustav Wallis 
(noe 64) on the Rio Branco, Amazonas, Brazil, and is depos- 
ited in the herbarium of the Botanisches Museum at Berline 


INDEX TO 


Abuta, 275, 278, 281 

Abuta concolor, 278 

Abuta oblongifolia, 276 
Abuta parvifolia, 276 

Abuta Selloana, 278 
Acanthaceae, 250 
Acrodiclidium, 177 
Acrodiclidium mexicanum, 181 
Acrodiclidium misantlae, 180 
Adenodesma, 33, 3 

Adiptera submontana, 215 
Aegiphila, 182, 183, 191,222, 


VOLUME ONE 


500, 376 

Aegiphila breviflora, 
190, 191, 301 

Aegiphila Buchtienii, 192,301, 
503 | 

Aerziphila Candelabrum, 183, 
192, 300, 301, 304, 364,366, 
381 

Aeziphila capitate, 193, 300, 
381 

Aegiphila casseliaeformis, 193 
500, 374 


— 


a 


183, 


226, 229, 250, 289, 297,377 Aepiphila caucensis, 193, 299, 


Aegiphila aculeifera, 184,222 
254, 290, 298, 299, 302, 
3055 567s 3135s 575; 378 

Aegiphile aequinoctialis, 260 

Aegiphile alba, 185, 290,300, 
304, 368 

Aegiphila amazonica, 378 

Aegiphila enomala, 183, 186, 
222, 298, 505, 575, 375, 
380 

Aegiphila arborea, 206 

Aegiphila arborescens, 227 

Aegiphila arborescens var. 
lonziflora, 189, 227, 365, 
368 

Aeziphila austrelis, 187, 222 
290, 300, 373, 376 

Aeziphile barbadensis, 188, 
298, 376, 380 

Aegiphila bogotensis, 183, 
188, 249, 290, 299, 300, 
304, 364, 365, 367, 373, 
374, 376, 380, 395 

Aergiphila boliviana, 188, 301 
380 


Aegiphila brachieta, 189, 206 
300, 301, 364, 368, 375, 
381, 3835 

Aegiphile bracteolosa, 183, 
189, 191, 192, 227, 290, 
299, 300, 304, 365, 367, 
368, 375,. 381 

Aegiphile brasiliensis, 190, 


ps ie, 

Aepiphila caymanensis, 193,298 
366 

Aegiphila cephalophora, 193, 
298, 502, 367 

Aegiphile cestrifolia, 238 

Aegiphila chrysantha, 194,290, 
500, 301, "367, 373532 574; BOL 

Aegiphilea compacta, 296 

Aegiphila conturbata, 194, 300, 
bl ie, 

Aegiphila cordata, 182, 183, 
195, ‘226, 500, 9755. 5165-008 

Aegiphile cordata var. colomb- 
iana, 381 

Aegiphila cordifolia, 195, 223 
300, 368, 374, 382 


Aegiphila coriacea, 195, 300, 
30h 


Aeziphile costaricensis, 196, 
290, 298, 365, 366, 3733;3195; 
382. 6 

Aegiphila crenata, 196, 223, 
258, 300, 304, 382 

Aegiphila cuneata, 196, 271, 
300, 367, 376, 382 

Aegiphila cuspidata, 296, 365, 
367, 391 


Aeziphila densiflora, 302, 366 

Aeziphila dentata, 197, 300, 
304, 365, 366, 382 

Aepiphile Deppeana, 182, 193, 
184, 197, 223, 291, 293,298, 


491 


492 PHYTOLOG IA 


299; 301, 304, 364, 365, 
366, 367, 368, 373, 374, 
3755. 376, .382 

Aegiphila diffusa, 366 

Aegiphila dubia, 252, 253, 
304, 376 

Aegiphila Duckei, 483 

Aegiphila elata, 184, 198, 
223, 260, 291, 297, 298, 
299, 301, 302, 303, 304, 
364, 365, 366, 368, 373, 
514 3159 5165377 384, 
385, 389 


Aegiphila el : 8, 199, 223, 


e gan 
500, 301, 367, 376, 385 


Aegiphila elongata, 200, 301, 


303, 385 


Aegiphila exiguiflora, 385 


Aegiphila falcata, 200, 291, 


298, 302, 466, 368, 374, 
375; 386 


Aegiphila fasciculata, 200, 
223, 240, 298, 366, 376,387 


Aegiphile Fendleri, 200, 299, 
2 


Aegiphila ferruginea, 183, 
201, 223, 291, 300, 364, 
366, 368, 374, 375, 387 


Aegiphila wa tHE: 201, 225; 


255, 291, 296, 299, 300, 
301, 365, 367, 368, 374, 
376, 387 

Aegiphila floribunda, 201, 
299, 304, 367 


Aegiphila fluminensis, 202, 


233, 300, 302, 303, 304, 
365, 367, 368, 374, 375; 
376, 388 


Aegiphila foetida, 202, 260, 
356, 298, 302, 304, 368, 


373, 375, 376 
Aegiphila fontex, 395 
Aegiphila furia, 254 
Aegiphila fusca, 254 
Aegiphila 

567, 388 


Aegiphila glandifera, 232 


Aegiphila glandulifera, 203, 


232, 291, 298, 299, 500, 


labrata, 203, 300, 


NOe 15 


303, 304, 364, 365, 367, 
375, 388 


‘Aegiphile glandulifera var. 


paraénsis, 203, 300, 367, 
368, 388 
Aegiphila glandulifera var. 
pyremidata, 203, 260, 292, 
299, 300, 368, 374, 376 
Aegiphila Glazioviana, 381 


Aegiphila Gleasonii, 204, 299, 
Seb wae : 


Aegiphila glomerata, 204, 300, 


5302, 3566, 375 | 
Aegiphila gloriosa, 204, 300, 
402 


Aegiphila Goeldiana, 204, 300, 
364, 388 

Aegiphila Goudotiana, 205, 224 
299, 364 

Aegiphila grandis, 205, 206, 
224, 249, 299, 364, 376 

Aegiphila graveolens, 183,205, 
292, 300, 302, 364, 365,367, 
368, 375, 389 

Aegiphila puianensis, 206,226, 
299, 300, 375, 376, 389 

Aegiphila guyanensis, 205, 206 

Aegiphila Hassleri, 183, 189, 
206, 300, 501, 502, 304,364, 
366, 373, 389 

Aegiphila Hastingsiana, 207, 
298 

Aegiphila Herzogii, 183, 224, 
301, 366, 385, 389 

Aegiphila hirsuta, 224, 301, 
503 

Aegiphila hirsutissima, 224, 
299, 373, 389 

Aegiphila Hoehnei, 224, 292, 
300, 365, 367, 390 

Aegiphila insipnis, 226, 300, 
375 

Aegiphila integrifolia, 189, 
226, 227, 255, 263, 292, 
294, 298, 299, 300, 301, 
302, 303, 304, 364, 365, 
366, 367, 368, 374, 375, 
376, 380, 390 

Aegiphila integerrima, 398 


1941 


Aegiphila intermedia, 195, 
228, 292, 300, 50h, 365; 
366, 390 


Aegiphila Killipii, 183, 228, 


292, 299, 367, 390 
Aegiphila laeta, 228, 229, 
269, 292, 299, 302, 368, 
373, 374, 375, 390, 393 
Aegiphila laeviflora, 251 
Aegivhila laevis, 228, 229, 
292, 299, 500, 302, 303, 
304, 365, 366, 367, 368, 
BID» BT4s 3755-576, 3855 
391 
Aegiphila laevis f. angusti- 
folia, 292 


eee lanata, 2430, 300, 
564, 3591 


Aepgiphila lanceolata, 
296, 297, 300, 501, 
303, 304, 364, 365, 

Aegiphila lanuginosa, 

Aegiphila laxicupulis, 231, 
298, 502, 366, 391, 393 

Aegiphila laxiflora, 251, 
293, 298, 299, 303; 304, 
367, 374, 375, 391 

Aegiphila Lehmannii, 231, 
299, 367, 376, 384, 391 

Aegiphila Lewisiana, 241, 
299, 304 

Aegiphila Lhotzkiana, 232, 
300, 302, 364, 365, 
367, 368, 373, 374, 
376, 392 

Aesiphila longifolia, 
295, 299, 566, 374, 

Aegiphila longipetiolata, - 
233, 300, 376, 392 

Aegiphila Luschnathi, 183, 
233, 300, 301, 302, 304, 
BON, 506, 567, 360, 5733 
374, 375, 376 

Aegiphila lutea, 381 

Aegiphila macrantha, 234, 
298, 299, 300, 301, 304, 
366, 368, 373, 376, 392 

Aegiphila macrophila, 385 

Aegiphila macrophylla, 234, 


Index 


_ Aegiphila mollis var. 
dia, 240, 299, 375 


495 


299, 385, 392, 392 
Aegiphila magnifica, 234, 298, 
502; 565,. 37535 576, 592 
Aegiphila martinicensis, 183, 
225, 23555-2573 295, 2963 
297; 299, 300, 301, 
302, 303, 304, 364, 365, 
366, 368, 373, 374, - 
3155 381, 590, 391,395 
Aegiphila martinicensis var. 
Oligoneura, 235, 237, 298, 
301, 304, 364, 376, 391, 393 
Aegiphila mediterranea, 238, 
5301, 502, 304, 365, 367, 
368, 373, 374, 375, 376, 393 


Aegiphila medullosa, 238, 301, 
37h, 395 


Aegiphila membranacea, 238, 
299, 300, 302, 365, 366, 
368, 374, 376 

Aegiphilea minima, 245 

Aegiphila mollis, 183, 2539, 
291, 295, 299, 501, 302, 
303, 304, 364, 365, 366, 
367, 568, 573, 315, 376; 359% 


interme- 


Aegiphila monstrosa, 223, 240, 
293, 298, 302; 364, 367, 
375 394 

Aegiphila montana, 248, 299 

Aegiphila monticola, 248, 300, 
368, 374, 380, 394, 395 

Aegiphila Mortoni, 249, 293, 
400 

Aegiphila multiflora, 250, 
300, 301, 303, 304, 367, 
374, 395 

Aegiphila Mutisii, 239, 575; 
382 

Aegiphila nervosa, 251, 298, 
303, 304, 365, 374, 375 

Aegiphila nigrescens, 254 


Aegiphila novofriburgensis, 
251, - 501 


Aegiphila novogranatensis, 
185, 251, 299, 367 

Aegiphila obducta, 252, 501, 
302, 303, 304, 364, 365, 


Lod 


366, 368, 373, 374, 375, 
376, 395 

Aegiphila obovata, 18%, 252, 
253, 298, 301, 303, 304, 
364, 366, 375, 376 


Aepiphila obtusa, 253, 298, 
36h 

Aegiphila odontophylla, 185, 
55h, 298, 368, 373, 395 

Aegiphila 


Orbignyana, 266 

Aegiphila ovata, 25h, 300; 
301, 367, 376 

Aegiphila pacifica, 473 

Aegiphila panamensis, 254, 
293, 298, 299, 3502, 503; 
304, 365, 373, 375, 395, 
396 

Aegiphila paniculata, 254, 
298, 299, 367; 39 

Aegiphila paraguariensis, 
226, 255, 293, 3501, 303, 
304, 364, 365, 367, 368, 
375, 396, 398 

Aegiphila parviflora, 255, 

299, 301, 365, 375 
Aegiphila pauciflora, 255, 
375 

Aegiphila Pavoniana, 256, 
300, 366, 373 

Aegiphila pendula, 256, 299, 
300, 365, 373, 396 

Aegiphila Pennellii, 257; 
299, 373 

Aegiphila pernambucensis, 
257, 201, 909, 565, 517 
374, 375, 396 

Aegiphila perplexa, 258, 298, 
299, 302, 304, 366, 374,396 

Aegiphila peruviana, 259, 294 
300, 301, 367, 368, 397 

Aegiphila platyphylla, 259, 
301, 3 

Aegiphile plicata, 183, 259, 
294, 295, 298, 502 


Aegiphila puberulenta, 259, 
299, 304, 397 

Aegiphila pubescens, 202 

Aegiphila pulcherrima, 259, 
300, 375 


PHYTOLOGIA 


Noe 15 


Aegiphila punctata, 298 

Aegiphila amidata, 204, 393 

Aegiphila quinduensis, 260, 
294, 299, 302, 366, 375, 397 

Aegiphila racemosa, 226, 260, 
299, 300, 301, 302, 305, 
304, 364, 365, 368, 373, 
374, 375, 397 

Aegiphila reticulata, 251, 
261, 299, 373 - 

Aegiphila Riedeliana, 261, 
301, 503, 368, 373, 375, 397 

Aegiphila Rimbachii, 261, 300, 
368, 374, 397 

Aegiphila roraimensis, 263, 
299, 302 

Aegiphila salicifolia, 5% 

Aegiphila salticola, 263, 294, 


301, 368, 497 


Aegiphila salutaris, 239, 302, 
a 


Aegiphila saluteris var. lut- 
ea, 249 

Aegiphilea scandens, 264, 294, 
301, 376 


Aegiphila Schimpffii, 265, 
59h, 300, 374 


Aegiphila Sellowiana, 227, 
266, 292, 294, 301, 302, 
304, 364, 365, 366, 367, 
368, 373, 374, 375, 376, 
396, 398 

Aegiphila sessiliflora, 267, 
299, 302, 398 

Aegiphila setiformis, 267; 
301, 302, 366, 398 

Aegiphila Skutchii, 399 

Aegiphila Smithii, 183, 268, 
29h, 300, 367, 368, 400 

Aegiphila solanifolia, 189 

Aegiphila sordida,. 268; 295; 
300, 467 

Aegiphila spicata, 267, 268, 
301, 302 

Aegiphila splendens, 268, 
365, 373 

Aegiphila Spruceana, 268, 
301, 375, 400 

Aegiphila Standleyi, 268, 298,_ 


1941 


375, 400 
Aegiphila straminea, 368 
Aegiphila stricta, 229, 269, 
Fe 


Aegiphila subopposita, 304 
Aegiphile sufflava, 269, 295, 


300, 367 
Aegiphila sufflava var. 
Klugii, 295, 300, 367 
Aegiphila Surfaceana, 269, 
301, 375, 400 3 
Aegiphila Swartziana, 269, 
298, 375, 376, 400 
Aegiphila sylvatica, 252, 
253, 269, 299, 367 a 
Aegiphila ternifolia, 183, 
270, 299, 365, 400 
Aegiphila tetragona, 205 
Aegiphila tomentosa, 3/75 
Aegiphila tomentosa var. 
silvestris, 398 
Aegiphila triantha, 381, 389 
Aegiphila trifida, 270, 294, 
ste 298, 368, 3573, 375; 
00 
sees triflora, 270, 300, 
367 
Aegiphila 
Aegiphila 
373 


A ae 
=F a 


Aegiphila 
376 

Aegiphila velutinosa, 272, 
300, 503 

Aegiphila verrucosa, 272, 
299, 304, 367, 368 

Aegiphila verticillata, 272, 
295, 301, 303, 304, 365, 
366, 367, 368, 374, 375, 
376, 398 

Aegiphila vestita, 232 

Aegiphila villosa, 232, 296, 
299, 300, 302, 303, 366, 
367, 368, 373, 374, 376 

Aegiphila villosissima, 183, 
336, 296, 301, 367 

Aegiphile virgata, 198, 301 


trinitensis, 303 
truncata, 271, 299, 


uniflora, 271, 298, 


Valerii, 272, 298, 


Index 


495 


Aegiphile vitelliniflora, 183, 
230, 260, 296, 297, 300, 
301, 302, 303, 304, 365, 366 
367, 368, 374, 375, 376 

Aegiphylla elata, 198. 

Aepuephila, 182 

Aegyphilla, 182 

Aiouea guyanensis, 115 


Aiouea rubra, 115 

Aiouea scandens, 115 

Alchornea megalostylis, 63 

Allomaieta prandiflora, 37 

Aloysia barbata, 41 

Aloysia Dusenii, 440 

Aloysia ligustrina var. parag-— 
uariensis, 167 

Aloysia ligustrina var. Schul- 
zii, 95 

Aloysia macrostachya, 95 


Aloysia pulchra, 95 


Aloysia scorodonioides var. 
detonsa, 95 
Aloysia scorodonioides var. 


Mathewsii, 95 
Aloysia spathulata, 95 
Aloysia urugueyensis, 167 
Aloysia virgata ver. elliptic- 
a, ry) 


Aloysia virgata var. laxa, 95 
Alseis, 101, 172 


Alseis floribunda, 137 

Alseis latifolia, 137 

Alseis leiantha, 96 

Alseis Mutisii, 95 

Alsine asensis, 53 

Ambiyarrhena, 43, 44, 47, 48 

Ambrina ambrosioides var. an- 
thelminticun, 274 

Amerina, 249, 377 

Amerina ternifolia, 270 

Amerina tomentosa, 188, 380 

Andraspidopsis, 335 

Andromeda, 8% 

Aniba Brittonii, 116 

Aniba citrifolia, 118 

Aniba elliptica, 115 

Aniba flexuosa, 117 

Aniba juruensis, 116 

Aniba Krukovii, 117 


496 


Aniba Muelleriana, 117 
Anneslia chiapensis, 369 
Anomospermum, 281 

Anona, 8% 

Anthodiscus montanus, 29 
Anthodiscus obovatus, 29 
Anthodiscus peruanus, 29 
Anthodiscus trifoliatus, 29 
Aphelandra sciophila, 82 
Aphelandra priematica, 82 
Apoleya, 143 

Apoleya leiocarpa, 143 
Apoleya molaris, 143 


Apuleia, 143 
Apuleia leiocerpa, 143 


Apuleia molaris, 143 
_ Apuleja, 143 

Apuleja praecox, 143 

Arbuscula, 46 

Arbutus, 83 


Arctostaphylos, 83 
Arcythophyllum flavescens, 79 
Arcythophyllum nodosum, 78 
Aspidosperma cruentum, 349 
Aspidosperma Matudai, 439 
Autodiplosis Iheringi, 226 
Avicennia alba var. latifol- 
ia, 410 
Avicennia balanophora, 409 
Avicennia marina var. scutis- 
sima, 411 
Avicennia marina vare snom- 
ala, 411 
Avicennia nitida var. trini- 
tensis, 9 
Avicennia Tonduzii, 273 
Avicenniaceae, 171 
Balsamita major vare 
oides, 27 
Banara umbraticola, 389 
Barnhertia, 111 
Bartsia altissima, 75 
Bartsia Guggenheimiana, 76 
Bartsia hispida, 76 
Bartsia mutica, 76 
Bauhinia, 212, 214 
Bauhinia Calderonii, 214 
Bauhinia congesta, 214 
Bauhinia gigas, 213, 214 


tanacet— 


PHY T O-BsOs6 Tk 


Bauhinia 
Bauhinia 


Beuhinia 


Begonia sanguinea, 


Noe 15 


214 
214 
214 
68 


Jermyana, 
monantha, 


Purpusii, 


Begonia subrectangula, 67 
Begonia unilateralia, 68 


Belucia, 


3 


Benthamidia Nuttallii, 167 


Berberis 
Berberis 
Berberis 
Berberis 
Berberis 


Berberis 


Berberis 


Berberis 
Berberis 


aquifolium, 91, 4 
californica, 93 
dictyota, 93 
Fremontii, 93 
nervosa, 90 
Nevinii, 93 

innata, 90 

umila, 92 
repens, 92, 94 


Bernoullia, 110 
Bernoullia swietenioides, 109 
Besleria, 151 


Besleria 
Besleria 
Besleria 
Besleria 


Besleria 


Besleria 
Besleria 


Blakea, 


chiapensis, 151 
insolita, 153 
laxiflora, 151, 153 
lutea, 153 

montana, 152 
saxicola, 151 
verecunda, 151, 152, 


arborea, 108 
excelsa, 107, 108 
Orellana, 108 


platycerpa, 108 
sphaerocarpa, 108 


urucurana, 


108 
2 


Blakea calycosa, 342 
Blakea truncata, 46 


Bouchea, 
Bouches. 
Bouchea 
Bouchea 


18 


boliviana, 18 
hyderobadensis, 434 


incisa, 18 


Bouchea integrifolia, 18 


Bouchea 


Krookii, 18 


latifolia, 18 
longipetala, 18 


Bouchea namaquana, 18 


Schlechteri, 18 
sessilifolia, 18 
Wilmsii, 18 


1941 Index 4907 
Bourreria ovata, 182 Cassia submontana, 215 
Boussingaultia, 245 Cassia villosissima, 215 
Brachyotum barbiferum, 69 Cassia xanthophylla, 215 


Briickea prandifolia, 367 
Buddleia, 149, 197, 232, 395 
Buddleia Skutchii, 148, 149 
Burroughsia, 411 
Burroughsia appendiculata,412 
Burroughsia fastigiata, qi2 
Bursera, 370 
Butea frondosa, 161 
Caesalpiniaceae, 23 
Caespa aquatica, 423 
Caiophora macrophylla, 67 
Calceolaria cryptantha, 77 
Calceolaria lechioides, 77 
Calceoleria ribesiaefolia, 78 
Calliandra chiapensis, 369 
Callicarpa, 191, 192, 197,223 
Callicarpa americana, 242 
Oallicarpa ampla, 234 
Callicarpa globiflora, 227, 
37 
Calluna, 83 
Calyptranthes, 218 
Calyptranthes belizensis, 218 
Camara solviaefolia p trans— 
valensis, 428 
Caopia parvifolia, 65 
Carphocephalus, 435 
Casparea Calderonii, 214 
Casparea congesta, 214 


Casparea Jermyana, 214 
Sasparea monantha, 214 


Casparea Purpusii, 214 
Casselia ilicifolia, 97 
Cassia, 21 

Brittoniana, 214 
cobanensis, 214 


comayapuana, 214 
Doylei, 214 


fenixensis, 214 
logirostrata, 214 
Mayena, 215 
monserratensis, 215 
Pinoi, 215 

Seleri, 215 
simulans, 215 
stenocarpoides, 215 


: 


Cassia yucatana, 215 
Sastela, 18 


Castela galapagei, 8 
Castelaria, 18 — 
Castelaria galapagei, 8 
Jatesbaea, 245 
Savendishia, 131 
Cavendishia amalfiensis, 210 
Cavendishia compacta, 211 
Cavendishia confertiflora, 210 
Cavendishia crassifolia, 211 
Savendishia Mexiae, 210 
Cavendishia nitida, 131 
Cavendishia obtusa, 131 
Celosia argentea var. linear- 
is, 5 


Centradenia grandifolia vare 
brevisepala, 340 
Cerastium brevicarpicum, 54 
Ceratostema, 130 
Ceratostema colombianum, 130 
Ceretostema coronarium, 130 
Ceratostema parvifolium, 131 
Ceratostema Rimbachii, 130 
Cerdana Gerasacanthus, 16 


Sespa, 311 
Cestrum, 205, 235 


Chaetolepis nana, 153 
Chamaecrista chiapensis, 215 
Chamaecrista comayaguana, 214 
Chameecrista fenixensis, 214 
Chamaecrista Pinoi, 215 
Chamaecrista Seleri, 215 
Chamaecrista stenocarpoides, 
215 
Chamaecrista Tonduzii, 215 
Chamaecrista villosissima, 
215 
Chamaecrista catana, 215 
Chamaefistula chiapensis, 214 
Chanekia, 177, 178 
Chanekia campechiana, 177,178 
Chenekia caudata, 178 
Chanekia coriacea, 179 
Chanekia mexicana, 178, 181 
Cheanekia misantlae, 178, 180 


498 


Chanekia Peckii, 178 
Chascanum, 18 
Shascanum incisum, 18 
Chascanum inteprifolium, 18 
Chascenum Krookii, 18 
Chascanum longipetalum, 18 
Shascanum latifolium, 18 
Chascanun namaquanum, 18 
Shascanum reriflorum, 167 
Chascanum Schlechteri, 18 
Chascanum sessilifolium, 18 
Chascanum Vilmsii, 18 
Chenooodium anthelminticun, 
27 , 
Shiococca alba, 283 
Shitonia, 143 
Chitonia macrophylla, 143 
Chlaenaceae, 83, 84 
Chomelia, 270 
Chondodendron, 281 
Shondodendron filipendulun, 
280 
Chusquea lanceolata, 145 
Chyvtroma, 125 
Chytroma apiculata, 124 
Chytroma rorida, 124 
Sissanpelos, 281 
Cissampelos fasciculata, 274 
Cissus rhombifolia var. glab-— 
rescens, 8 
Citharexylum, 101, 172 
Citharexylum affine var. 
landuliferum, 412 
Citharexylum fruticosum var. 
subserratum, 17 
Citharexylum fruticosum vare 
villosum, 17 


Citherexylum Ghiesbreghtii, 
413 


Citharexylum hidalgense, 414 
Citharexylum integerrimun, 17 
Citharexylum Kobuskianum, 441 
Citharexylum linearifoliun,9/ 
Citharexylum Mocini var. lon- 
gibracteolatun, R15 
Citharexylum montevidense, 17 
Citharexylum myrianthum var. 
rigidum, 17 


Citharexylum pernambucense, 


Pon T T Ore OrG kek 


Noe 15 


Aho 
Citharexylum Rimbachii, 443 
Citharexylum rigidum, 17 
Citharexylum Shrevei, 415 
Citharexvlum subserratum, 17 
Citharexylum villosum var. in- 
tegerrimum, 17 
Cladocaulon, 331 
Slerodendron bolivisnum, 250, 
374, 375 
Clerodendron Matudae, 291 
Clerodendrum, 184, 197, 231, 
238, 250 
Clerodendrun aculeatum var. 


puianense, 
Clerodendrunm africanum, 416 


Clerodendrum capitatum, 234 

Slerodendrum Duckei, Th 

Clerodendrum Ekmani, 445 

Clerodendrum japonicum var. 
album, 167 

Clerodendrum Kaempferi vare 
album, 167 

Clerodendrum lisustrinum var. 


nicaraguense, 41 


Clerodendrum margaritense, 


Clerodendrum Pittieri, 415 
Clerodendrum Rusbyi, 447 
Clerodendrum Scheffleri var. 
ellipticum, 417 
Clerodendrum suffruticosum 
var. natalense, 418 
Clerodendrun ternifolium 
var. Mexiae, 
Clerodendrum Tessmanni, 448 
Clethra glaberrima, 219, 220 
Clethra Matudai, 220 
Clethra parvifolia, 220 
Clethra suaveolens, 220 
Clethra vulcanicola, 220 
Clidemia, 17 
Clidemia ampla, 46 
Clidemia ferox, 175 
Clidemia longisetosa, 176 
Clidemia ostrina, 45, 46 
Clitoria, 141 
Clusia, 212, 218 
Clusia Lundellii, 218 


1941 


Clusia suborbicularis, 218 

Coccoloba escuintlensis, 213 

Cocculeae, 280 

Cocculus, 280, 281 

Cocculus Filipendula, 

Somnelina reflexa, 50 

Cordia, 234 

Cordia Gerasacanthus, 

Cordia pubescens, 199 

Cornutia, 234 

Sornutis odorata var. 
cens, 2 

Cornutia pubescens, 182 

Sornutia velutina, 239, 367 

Cornus Nuttallii, 167 

Costus longifolius, 50 

Costus Tatei, 51 

Couratari macrosperma, 125 

Souratari pulchra, 125 

Cremanium, 31 

Cremanium serrulatum, 144 

Croton Aguilari, 4o1 

Croton Ameliae, 401 

Groton asteroides, 401, 402 

Groton Sascarilla, 167 

Croton chichenensis, . 449 

Croton Cortesianus, 449 

Croton cupreatus, 433 

Groton flavoglandulosus, 403 

Croton glabellus, 405, 407, 


451 
ossypiifolius, 450 
icche, 404 


itzaeus, 405 
Lamarckianus, 167 
niveus, 450 
erobtusus, 405 
etenensis, 406 


pseudoglabellus, 405, 


seudoniveus, 449 
punctatus, 433 
Soliman, 408 
sutup, 407 
tabascensis, 408 
tacanensis, 450 
tenuicaudatus, 451 
Watsonii, 409 


xalapensis, 401 


280 


16 


‘calves- 


<2 


Croton 
Croton 


Index 


499 


Croton yucatanensis, 408 
Cymosae, 191 

Cyphostyla, 37 

Cyphostyla hirsuta, 37 
Cyphostyla strigosa, 56, 37 
Cyphostyla villosa, aT 
Dalea dispar, 147 

Davilla Matudai, 371, 372 
Davilla multiflora, 372 
Decaraphe Hostmanni, 143 


Dendrostigma, 29 
Dendrostigma hystricina, 40 


Dendrostylis, 40 
Dentatae, 191 


Dermatocal perviflorus, 200, 
225, 240, 377 
Dialyanthera acuminata, 114 


Dialyanthera olacoides, 113, 
rik 


Dichrolepis, 311 
Diclidanthera, 111 


Diclidanthera octandra, 110, 
pe 
Diclidantheracese, 111 
Diffusa, 148 
Dioclea coriacea, 7 
Dioclea densiflora, 7 
Dioclea eee 
Dioclea pulchra, 
Dioclea rufescens, 56 
Diospyros Tectona, 155 
Diplochaeta, 143 
Diplochaeta Hostmanni, 14% 
Diplochaeta leucocephala, 143 
Diplochaeta serrulata, 14% 
Diplochaeta serrulata var. 
latifolia, 143 
Dipteracanthus Schauerianus, 
82 


Disciphania, 283 

Disciphania Killipii, 274 
Disporum Cahnae, 169 

Disporum Hookeri, 168 
Disporum lanucinosum, 168, 169 
Disporun maculatum, 168, 169 


Disporum oregsanum, 168 


Disporum pervifolium, 168 
Disporum Schaffneri, 167, 169 


Disporum Smithii, 168 


56 


Endiandrae 


— 


500 2 HY. Posh 


Disporum trachyandrum, 168 
Disporum trachycarpym, 168 


OGIA Noe 15 


Eriocaulon articulatum, 323,324 
Eriocaulon Benthami, 311, 316, 


Dissilaria baloahioides, 161 350, 356, 357, 359 


Disterigma, 130 
Doliocarpus coriaceus, 348 
Drimys, 83 

Drymariea glandulosa, 55 
Drymaria Ladewii, 54 
Drymaria pauciflora, 55 
Drymaria stricta, 55 


Duranta Hitchcockii, 460 
Duranta Penlandi, 461 
Duranta Plumieri, 74 


Duranta pseudorepens, 460 
Duranta recurvistachys, 73 


Duranta repens vare canes- 
436 


cens, 
Durenta repens vare micro- 


phylla, 483 
Dupatya, 331, 4352 
Dupatya alsinoides, ASO 
Dupatya caulescens, 336 
Dupatya flavidule, 336 
Dupatya Lamarckii, 453 
Dupatya montana, 433 
Dupatya pungens, 334 
Dupatya seslerioides, 334 
Echiphylle, 377 
Echites microcalyx, 13 
Egiphila, 182 
Egiphylla, 377 


Egiphylla macrophylla, 234 
Ehretia montevidensis, 17 
Elaeocarpaceae, 122 
Electrosperma, 311 
Eleocharis, 322 
Elephantopus, 40 
Elephantopus erectus, 40 
Elephantopus hirtiflorus, 40 
leuca, 161 
Endlicheria formosa, 118 
Endlicherie glaberrima, 118 
Eriocaulon, 311, 331, 343 
Eriocaulon amplexicaulis, 347 
Eriocaulon anceps, 316, 328, 
261 
Eriocaulon aqueticum, 319,323 
Eriocaulon arenicole, 311, 


350, 252 


‘Eriocaulon 


411 
ll; 


Eriocaulon Benthamii, 
Eriocaulon bilobatum, 

312, 322, 350, 360 
Eriocaulon brachypus, 420 
Eriocaulon caulescens, 336 


Eriocaulon cephalotes, 312 


Eriocaulon conpressum, 312, 
348, 3h9, 5503 551; De, 
353, 354, 356, 357, 358, 
359, 360, 361, 362, 363 

Eriocaulon compressum vare 
Harperi, 313, 349, 350, 351; 
552, 3565. 3585 35950 562 

Eriocaulon cubense, 313 

Eriocaulon decanpulare, 312, 
314, 323, 348, 349, 350, 
3515 352 5.:3535 2am 
356, 357, 358, 359, 360, 
361, 362, 363 

Eriocaulon decangulare vare 
latifolium, 316, 349, 350, 
356, 360 

Eriocaulon dioecum, 316, 350, 
355 

Eriocaulon echinospermoideun, 
316, 350 

Eriocaulon echinospermum, 316, 
350, 352, 355, 361, 363 

Eriocaulon Ehrenbergianun, 
316, 350, 352, 353, 354, 
360, 361, 362 

Eriocaulon Ekmannii, 317, 330, 
350, 354, 355 

Eriocaulon fasciculatum, 333 

Eriocaulon filiformis, 312 

Eriocaulon flavidulum, 336,343 

Eriocaulon Plevidum, 436 

Eriocaulon fuliginosum, 317, 
322, 350, 352, 3535 355; 
356, 258, 360, 361, 363 

Eriocaulon fusiforme, 317, 
Pls 352 

eraénse, 336 

Erioceulon pnaphalodes, 312, 
422 


Eriocaulon guadelajarense, 518, 


1941 
350, 360 


Eriocaulon heteropetalum, 318, . 


oe 

Eriocaulon insulare, 318, 
F015 D2S5: 59D 

Eriocaulon Jaliscanum, 312, 
360 

Erioceulon Koernickianum, 318 

Eriocaulon Kornickisanum, 318, 


350, 356, 3 
Erioceaulon Kinlochii, 318, 
350, 357 


Eriocaulon lacustre, 318, 
351, 355 

Eriocaulon Lamarckii, 333 

Eriocaulon leptodictyon, 311 

Eriocaulon lineare, 318, 


Index 


Eriocaulon 
Eriocaulon pusillum, 320 


501 


350, 360 


Eriocaulon pseudocompressum, 
522, 351,. 354, 355s. 565 


Eriocaulon pubigerum, 328 
umilum, 323 


Eriocaulon Ravenelii, 322, 
9, 354, 360, 362 . 

Eriocaulon Schiedeanum, 312, 
5175 322, 590, F5l,: 560 

Eriocaulon Schippii, 323, 
350, 361 

Eriocaulon ecirpoides, 317, 
565 


Erioceaulon sclerocephalun, 
Jee) SOL toes 5h, 255 


349, Eriocaulon Seemannii, 323, 


350, 352, 354, 356, 360,363 350, 361, 365 


Eriocaulon maritimum, 330 

Eriocaulon melanocephalun, 
519, 351, 357, 3563 

Eriocaulon melanocephalum 
vere longipes, 319, 351,363 

Eriocaulon mexicanum, 319, 
320, 350, 360 

Eriocaulon microcephalum, 316 
320, 350, 351, 357, 358, 
360, 361, 362, 364 


Eriocaulon minutissimum, 320, 
ek we OF, 

Eriocaulon miserrimum, 320, 
Pole 2D 

Eriocaulon Noveboracens, 323 

Eriocaulon olivaceum, 320, 
20) Pe op, 

Eriocaulon ovoideum, 320, 
351, 552 

Eriocaulon panamense, 321, 


350, 357, 362 

Eriocaulon Palmeri, 320, 350, 
y) 

Eriocaulon Parkeri, 321, 348, 
BND, B52s. 355, 55%). 355, 
356, 357, 358, 359, 360, 
362, 363 

Eriocaulon pellucidum, 323 

Eriocaulon pinerense, 3521, 
351, 353, 555 

Eriocaulon Pringlei, 322, 


Eriocaulon septanpulare, 324, 
348, 349, 350, 351, 352, 353 
354, 3555. 356,557, 3585 
359, 360, 361, 362, 363, 364 

Eriocaulon septanpulare var. 
natans, 323 

Eriocaulon serotinum, 314 

Eriocaulon sigmoideum, 327, 


351, 363 
Eriocaulon simillimum, 336 
Eriocaulon spathaceum, 336 


Eriocaulon sphaerospermum, 


317, 363 


Eriocaulon splendens, 336 
Eriocsulon statices, 314 
Eriocaulon subuncinatum, 336 
Eriocsulon tepicanum, 327, 


350, 360 
Eriocaulon texense, 327, 350, 
354, 360 


Eriocsulon trichosepalum, 317, 
ee ee 


363 


Erioceulon villosum, 328 


Eriocaulon Williamsii, 327, 


350, 359, 363 

Epacridaceae, 83 

Epacris, 83 

Eremolaena Humblotiana, 84, 
86, 88 

Eremolaena rotundifolia, 84, 
86, 88 


502 


Erica, &3 

Ericaceae, 83 
EBriocaulaceae, 309, 348 
Erythrina, 286, 287 
Erythrina Eggersii, 289 
Erythrina Folkersii, 286 
Erythrine horrida, 289 
Erythrina neglecta, 287 
Erythrina rubrinervia, 287,289 
Erythroxylon, 212 
Erythroxylon belizense, 215 


Erythroxylon rufum, 216 
Eschweilera apiculata, 124 


Eschweilera Fendleriana, 22 
Eschweilera Krukovii, 124 
Eschweilerea obtecta, 125 
Eschweilera rorida, 124, 125 
Eschweilere sessilis, 21 
Eubeslerie, 152, 153 
Eueschweilera, 124, 125 
Eugenia balancanensis, 481 
Eugenia belizensis, 218 
Eugenie bumelioides, 408 
Eugenia cocquericotensis, 438, 
Eugenia Gentlei, 439 

Eugenia Matudai, 218, 219 
Eugenia tabascensis, 482 
Eugenia toledinensis, 407 
Eugenia vacana, 408 


Eugenia xalapensis, 307, 308 
Euhenriettella, 31 


Eulepis, 445 

Eumiconia, 32, 48 
Eupatorium leptodictyon, 311 
Eutragia, 371 

Euweigeltia, 12 


Evea pectinata, 80 
Evea radiata, 79 


Exothea, 242 

Exothea copalillo, 242, 243 
Exothea diphylla, 242 
Exothea paniculata, 242 
Feramea occidentalis, 235 
Filipendula, 281 

Fischeria columbiana, 14 
Fiecheria viridis, 13 
Flacourtiaceae, 112, 389 
Flindersia Oxleyana, 161 


Fourcroya, &3 


> 


PvE TT Orb Oe 1A 


NOe 15 


Galium charoides, 80 
Galium pseudoaparine, 80 
Gentiana Tatei, /2 
Geranium Palmeri, 148 
Geranium pulchrum, 147, 148 
Gesneriaceae, 151 

Ghinia juncea, 169 

Ghinia spicata, 169 
Gilibertia arborea, 372 
Gilibertia Matudai, 372 
Glossocentrum, 32 

Gmelina speciosa, 418 
Gmelina tonkinensis, 419 
Godoya magnifica, 27 
Godoya splendida, 28 
Godoya sulcata, 28 
Goethalsia, 112 

Goethalsia isthmica, 112 
Graffenrieda, 41 
Graffenrieda colombiana, 41 
Graffenrieda emarginata, 41 
Gaultheria martaensis, 71 
Guatteria setosa, 55 


Guignardia prominens, 245 
Gunnera insignis, L52 


Gunnera Killipiana, 452 
Gunnera mexicana, 452 
Guttiferae, 122 

Gynandropsis brachycarpa, 6 
Gynandropsis hirsuta, 5 
Hebenstreitia rariflora, 167 
Hedeoma adscendens, 7 
Helenium decurrens, 169 
Helenium denticulatum, 169 
Hemipersea, 119 

Henriettella, 36 

Henriettella fascicularis, 46 
Henriettella fissanthera, 35 
Henriettella Goudotiana, 31 
Henriettella Lawrancei, 31 
Henriettella loretensis, 46 
Henriettella sylvestris, 108 
Henriettella tovarensis, 109 
Henriettella verrucosa, 31,109 
Hippocratea Austin-Smithii, 438 
Hippocratea celastroides, 438 
Hivpocratea lancifolia, 438 
Hobsonia Ackermanni, 235 


Hormocalyx, 17 


1941 


Hormocalyx hirsutus, 174 

Hortia, 25 

Hortia brasiliana, 25 

Hortia colombiana, 25 

Hufelandia, 119 

Humulus japonicus var. varie- 
atus, 169 

Humulus scandens, 169 

Humulus scandens var. varieg— 
atus, 169 

Hyperbaena, 276, 278, 281 


Hyperbaena domingensis, 274, 
Bf, 2f 

Hyperbaena graciliflora, 275 

Hyperbaena Hassleri, 276 

Hyperbaena oblongifolia, 276 

Hyperbaena ovalifolia, 276 

Hyperbaena solimoesana, 277, 
278 

Hypericum, 141 

Hyphydra, "3h7 


Hyphydra amplexicaulis, 347 
Ilex, 212 
Ilex belizensis, 217 
Tlex guianensis, Oerear 
Tlex panamensis, 217 
Tlex utilis, 8 
ee 147, 148 
Inga Lawranceana, 23 


Ipomoea cardiosepala, 14 
Ipomoea denticulata, 14 


Ipomoea Nil, 15 

Isandrina xanthophylla, 215 
Ixora fasciculata, 270 
Jacquinia, 83 

Jacquinia Sonzattii, 247 


Jatus 8s. Gaju jati, 156 
Juanulloa bicolor, 47, 38 


Juanulloa speciosa, 38 
Jucunda, - 4}, 4% 


Jugastrum obtectum, 125 


Juncus, 83 


Junellia straguloides, 483 
Jussiaea, 83 


Lachnaiocarpae, 156 

Lachnocaulon, 328 

Lachnocaulon anceps, 328, 
329, 351; » 350, 551, 
5595. 3545. 555 91556,) 550, 


Index 


503 


358, 359, 360, 361, 362, 363 
Lachnocaulon Beyrichianum, 328, 


329, 331, 349, 356, 360 


Lachnocaulon cubense, 329, 


351, 255 


Lachnocaulon diandrum, 329 


Lachnocaulon digynum, 329, 
DOD; Fee D 
Lachnocaulon eciliatum, 350, 


2h9, 352, 354, 356 
Lachnocaulon Ekmannii, 317, 

330, 351, 355, 361 
Lachnocaulon Engleri, 4330, 

349, 353, 356, 357, 359, 360 


Lachnocaulon floridanum, 4530, 


h9, 359 
Lachnocaulon glabrum, 440, 


351, JAD, Dis Se 
Lachnocaulon maritimum, 330 
Lachnocaulon Michauxii, 328, 

yp. 

Lachnocaulon Michauxii minus, 

531 
Lachnocaulon Michauxii var. 

minor, 331 
Lachnocaulon minor, 331 
Lachnocaulon minus, 328, 331, 


319, 352, 355, 354, 356, 

357, 359, 362, 3563 
Lacistema pedicellatum, 138 
Lacistema Pittieri, 138 
Laestadia prominens, Ae go & 
Lagerstroemia cicrOcarpes 161 


Langsdorffia hypogaea, 5 
Lantana aristata var.e angusti- 


folia, 170 
Lantana aristata var. pluripe- 
dunculata, 170 


Lantana boyacana, 465 


Lantana 
Lantana 


Chamissonis, 17 
cordobensis, 279 
Lantana frutilla, hig 
Lantana Glaziovii, 462 
Lantana prosseserrata, 420 
Lantana Haughtii, L63 
Lantana hispida, 434 
Lantana Junelliana, 279 


Lantana Langlassei, 434 
Lantana Mearnsii, 421 


504 


notha, 422 
Pavonii, 464 
rubella, 464 
salvifolia, 423, 429 
scabra, 
scabrifolia, 422 
spicata, 279 
tenuifolia, 74 
tiliaefolia, 74 
Tomasii, 
Lantana trifolia, 422 
Lantana velutinoides, 466 
Lantana velutinoides var. 
Bruchii, 466 
Lasiolepis, 311, 431 
Lasiolepis aquatica, 319 
Lasiolepis pilosa, 433 
Lauraceae, 115, 177 
Leandra, 48 
Leandra cuspidata, 48 
Leandra lasiopetala, 341 
Leandra lepidota, 341 


Lecythidaceae, 123 
Lecythis hians, 123 
Lecythis paraensis, 124 
Lecythis usitata, 124 
Ledum, 8% 

Leiocarpae, 156 
Leitgebia, 39 

Leitgebia ianensis, 39 


Lantana 
Lentana 
Lantana 
Lantana 
Lantana 
Lantana 
Lentana 
Lantana 
Lantana 
Lantane. 


PHY T Owe Ds 


No. 15 


Lippia costaricensis, 424 
Lippia Curtisiana, Hos 


Lippia durangensis, 426 


Lippia fastigiata, 412 
Lippia Gehrtii, 453 


Lippia Grisebachiana, 279 

Lippia Hoehnei, 467 

Lippie indica, 427 

Lippia jaliscana, 427 

Lippia lantenifolia, 279 

Lippia lantenifolia var. cren-= 
ata, 279 

Lippia ligustrina var. parag= 
uariensis, 167 

Lippia ligustrina var. 
Schulzii, 95 

Lippia mattogrossensis, 468 

Lippia micromera var. paludic- 
ola, 468 

Lippia nodiflora var. normalis 
fe sericea, 98 


Lippia nodiflora var. pusilla, 
170 
Lippia pulchra, 95 


Lippia scorodonioides var. de- 
tonsa, 95 

Lippia scorodonioides var. 
Mathewsii, 95 

Lippia spathulata, 9 

Lippia tepicana, 454 


Leitgebie Imthurniana, 39, 40 Lippia transvalensis, 428 


Leptolaena miltiflora, 384, 
86, 8&8 

Leptolobium leiocarpum, 14% 

Leptopoda denticulata, 169 

Leucopogon, &3 

Leucostomae, 147 

Limmoxeranthemum, 331 

Lippia, 412 

Lippia affinis, 167 

Lippia appendiculata, 412 

Lippia aristata f. pluriped- 
unculata, 170 


Lippia aristata ver. anpusti- 
folia, 170 


Lippie asperrima var. rotund- 
ata, Le 


Lippia canescens, 98 
Lippia controversa, 423 


Lippia urticoides vare laxa,95 
Lippia virgata ver. elliptica, 
iy 


Lippia Whytei, 428 


Lippia Wrightii var. macro- 
stachya, 95 


Lobatae, 191 

Lonchocarpus, 140 
Lonchocarpus denudatus, 140 
Lonchocarpue pallidus, 140 
Lonicera alba, 28% 

Lonicera canadensis, 283 
Lonicera ciliata, 283 
Loranthaceae, 113 

Lozania, 138 

Lozenia bipinneta, 138, 139 
Lozania Klugii, 138 

Lozania Mutisiana, 138 


1941 


Lozania pedicellata, 138 
Lozania Pittieri, 138 


Lupinus Buchtienii, 56 
Lupinus paniculatus, 58 
Lupinus prostratus, 5/7 
Lupinus soratensis, 58 
Lupinus Tatei, 57 

Luzula, 83 

Lycianthes novogranatensis,16 

Macleania, 131 

Macleania hirtiflora, 132 

Macleania mollis, 132 

Macleania nitida, 131 

Macleania rupestris, 131, 132 

Mahonia, 89 

Mahonia aquifolium, 89, 91,92 

dictyota, 90, 91, 9% 

fascicularis, 90 

fremontii, 90, 93 
lumosa, 90 

nérvosa, 89, 90 

nevinii, 90, 9% 
innata, 89, 90 


Mahonia 


Mahonia piperiana, 89, 91 
Mahonia pumila, 89, 90, 92 


repens, 89, 90, 92 

sonnei, 89, 92 

Maieta, 175 

Manabea, 296 

Manabea arborescens, 227 

Meanabea tomentosa, 296 

Markea campanulata, 339 

Merkea internexa, 340 

Markea uniflora, 340 

Martia, 141 

Martiodendron, 141 

Martiodendron elatum, 141,142 

Martiodendron excelsum, 141, 
142 

Martiodendron macrocarpon,142 

Martiodendron parvifoliun, 
141, 142 

Martiusia, 141 

Martiusia elata, 141 

Martiusia excelsa, 141 

Martiusia parvifolia, 141 

Mascapnia, 19 

Mascapnia dissimilis, 19 


Mascagnia nervosa, 19 


Index 505 


Matisia 
Matisia 


alchornaefolia, 27 
bicolor, 27 
Castano, 27 
Matisia cordata, 27 
Matisia Cornu-copiae, 27 
Matisia Dowdingii, 27 
Matisia plandifera, 27 
Matisia lasiocalyx, 27 
Matisia longiflora, 26 
Matisia obliquifolia, 27 
Matisia oblongifolia, 27 
Matisia ochrocalyx, 27 
Matisia paraensis, 2/7 
Mayna, 30 
Maytenus puatemalensis, 405, 
30 


Maytenus phyllanthoides, 307 
Maytenus Schippii, 305, 306 
Maytenus texana, 306, 307 
Meibomia adhaerens, 58 
Meibomia adpressa, 58 
Meibomia nummlaria, 59 
Meibomia simplicifolia, 40 
Meibomia tetrasperma, 50 
Melastomaceae Miconieae, 174 
Melastones, q Z 174 
Meliosma idiopoda, 243 
WYeliosma Matudai, 243 
Meliosma oaxacana, 243 
Mendoncia Lindavii, 17 
Menispermaceae, 274, 280, 281 
Menziesia, 83 
Merinthopodium campanulatun, 
539 
Merinthopodium internexum, 340 
Mesogynixa, 19 


Mespilodaphne, 120 
Miconia, 35, 48, 143, 144 


Miconia axinaeoides, 34, 35 
Miconia Chamissois, 342 


Miconia chrysophylla, 32 
Miconia Cookii, 43 

Miconia difficilis, 44, 45 
Miconia elongata, 

Miconia eugenioides, 48 
Miconia fiesa, 43 

Miconia fulva, 32 

Miconia galactantha, 144 


Miconia pratissima, 43 


506 PHYTOL 


Miconia Kraenzlinii, 32, 43 
Miconia leucocevhala, 144 
Miconia longiracemosa, 342 
Miconia macrophylla, 143, 144 
Miconia megalantha, 33, 34,35 
Miconia megaphylla, 43 
Miconia mesmeana, 

Miconia monzoniensis, 44 


Miconia mutisella, 47 
Miconia pachydonta, 41, 43 


Miconia panamensis, 341 
Miconia platyhedra, 144 
Miconia plena, 31 

Miconiea prasina, 144 
Miconia robusta, 144 
Miconia robustissima, 144 
Miconia serrulata, 143, 144 
Miconia tetrasperma, 48 
Miconia theaezans, 32 
Miconia tschudyoides, 144 
Miconia tubulosa, 32 
Mimosa Lawranceana, 23 
Mimosaceae, 23 

Misanteca, 177 

Misanteca campechiana, 178 
Misanteca Peckii, 178 
Mollinedia Krukovii, 114 
Mollinedia latifolia, 115 
Mollinedia Rusbyana, 115 
Mollinedia Selloi, 114 
Monimiaceae, 114, 126 
Monobea, 296 

Moschoxylum, 106 
Mucuna, 7 

Mucune. Holtoni, 7 

Mucuna urens, 7 
Myriaspora, 176 
Myriocarpa Tatei, 52 
Myristicaceae, 113 
Myrmidone, 176 

Myrtus Metudai, 246, 247 
Myrtus montena, 247 
Nasmythia, 311 

Nasmythia articulata, 323,324 


Nasmythia septangularis, 423 
Nautea, 154 
Nectandra lineatifolia, 120 


Nectandra superba, 120 
Nemochloa Beyrichii, 81 


OGIA No. 15 
elegans, 82 
Gaudichaudii, 81 
macrantha, 81 
Martiana, 81 


Nemochloa 
Nemochloa 
Nemochloa 
Nemochloa 


Nemochloa milleprana, 81 
Nemochloa sparsiflora, 82 
Nemochloa Urvillei, 82 
Neocastela, 18 

Nomochloa Orbigniana, 81 
Nomochlosa stricta, 81 


Nomochloa tenuiflora, 82 
Nomochloa turbinata, 82 
Ocotea campechiana, 178 
Ocotea lanata, 120 
Ocotea tenella, 119 
Odontadenia, 13 


Odontocarya, 280, 281 


Odontocarya filipendula, 280 
Odostemon aquifolium, 91 


Odostemon dictyota, 93 
Odostemon fascicularis, 90 
Odostemon fremontii, 93 
Odostemon nervosus, 90 
Odostemon nevinii, 93 
Odostemon nutkanus, 91 
Odostemon pumilus, 92 
Odostemon repens, 92 
Oldfieldia africana, 160 
Orchidaceae, 87 
Osmanthus mexicana, 308, 309 
Oxalis, 62 
Oxalis densissima, 61, 62 
Oxalis Tatei, 62 
Oxalis virgata, 62 
Oxymeris cuspidata, 48 
Pachylobium, 7 
Paepalanthus, 331, 332, 335 
Paepalanthus alsinoides, 452; 
351, 352, 355, 563 
Paepalanthus alsinoides vare 
minimis, 332, 351, 352, 355; 
357, 361, 363 
Paepalanthus androsaceus, 435 
Paepalanthus androsaceus vare 
flavescens, 335 
Paepalanthus caulescens, 336 
Paepalanthus chiapensis, 332, 
350, 3 


Paepalanthus costaricensis, 


1941 


332, 350, 362 
Paepalanthus domingensis, 


D525 751, B54, 356, 563 


Paepalanthus flavidulus, 331, 
536 


Paepalanthus flavidus, 336 

Paepalanthus Gentlei, 433, 
550, 351, 356 

Paepalanthus lagopodioides, 


Paepalanthus lagospodioides, 


eee eas Lamarckii, 319, 
52D 350, 351, 3535 355; 
356, 360, 361, 363 

Paepalanthus Mellii, 433, 
350, 359 


Paepalanthus montanus, 333, 
351, 361 


Paepalanthus Ottonis, 433 

Paepalanthus pilulifer, 3531 

Paepalanthus procerus, 336 

Paepalanthus pungens, 334, 
Ils 3D 

Paepalanthus retusus, 334, 
Dl, 255, 365 

Paepalanthus riparius, 334, 
5515. 361 

ae alanthus seslerioides, 

» Dl, oe? 20795! D0's 


<8" 361, 363 
Paepalanthus splendens, 336 


Paepalanthus surinamensis, 
336 
Paepalanthus Tuerckheimii, 
53h, 351, 353, 362 
Paniculatae, 191 
Parabesleria, 151 
Parathesis £ rsiana, ll 
Parathesis rubella, 11 
Parathesis serrulata, 11 
Parsonsia satureioides, 69 
Passiflora bauhinifolia, 67 
Passiflora Tatei, 66, 67 
Paullinia Tatei, 64 
Peiranisia Doylei, 214 


Peiranisia longirostrata, 214 


Peiranisia simulans, 215 
Periploca, 83 


Index 


507 
Pernettya, 71 
Pernettya densa, 70 
Pernettya Pentlandii, 71 
Pernettya schizosti » (0 


Perrottetia costaricensis, 451 

Perrottetia racemosa, 45] 

Perrottetia sessiliflora, 451 

Persea, 119 

Persea cuneata, 119 

Persea durifolia, 119 

Persea inaequalis, 118 

Petitia quinduensis, 294 

Petitia tenuifolia, 294 

Petraeeae, 101, 172 

Petrea Duckei, 469 

Petrea insignis, 470 

Petrea rugosa, 394 

Petrea Schomburgkiana, 17 

Phenax ballotaefolius, 5% 

Phenax bullatus, 52 

Phenax flsvifolius, 53 

Phoradendron, 293 

Phragmites, 170 

Phragmites communis, 170 

Phrsapmites communis vare vari- 
egatus, 170 

Phragmites maximus, 170 

Phragmites maximus var. varie- 

atus, 170 

Phyla nodiflora vare canes- 
cens, 98 

Phyla nodiflora var. 
170 

Phyla nodiflora vare sericea, 
98 

Phyllanthus antillanus vare 
hypomalacus, 337, 3438 


Phyllanthus nobilis ver. hypo- 
malecus, 438 


Picramie atudai, 241, 242 

Pilea pracilipes, 146 

Pilea quercifolie, 146 

Pinpuicula ae 99 

Pinguicule pumila, 99 

Pinpuicule See vare Pus- 
wellii, 98 

Pinzona coriacea, 438 

Piper h hians, 437 

Piper Mayanum, 337 


usilla, 


508 PHYTOLOGIA No. 15 


Pisonia, 235 Pseudaegiphila breviflora, 95, 
Pithecolobium campechense,481 — 189, 190 
Pithecolobium lenceolatum,481 Pseudobesleria, 151, 153 


Plenckia, 28h Psilocephalus, 345 

Plenckia bahiensis, 286 Psittacanthus biternatus, 20 
Plenckia integerrima, 284 Psittacanthus dilatatus, 20 
Plenckie microcarpa, 285, 286 Psittacanthus leptanthus, 113 
Plenckia populnea, 286 Psittacanthus si hon, 113 
Pleurostachys, @1 Psychotria, 235 


Pleurostachys Beyrichii, 61 Pterocarpus Marsupium, 161 
Pleurostachys elegans, 82 Pyrethrum Balsamita tanacet- 
Pleurostachys foliosa, 81 oides, 274 

Pleurostachys praminifolia,81 Pyxidanthus, 47 

Pleurostachys Gaudichaudii,81 Quercus oleoides var. sustral- 
Pleurostachys latifolia, &2 is, 34 


Pleurostachys Luetzelburgiena Quercus prinus, 147 


82 Raimondia quinduensis vare 
Pleurostachys macrantha, @1 latifolia, 5 


Pleurostachys Martiana, 81 Randalia, 311 

Pleurostachys millegrana, 81 Randalia americana, 314 
Pleurostachys Orbigniana, 81 Randalia decanpulare, 314 
Pleurostachys panicoides, 81 Randalia decangularis, 314 
Pleurostachys puberula, 82 Rapanea Rapanea denticulata, 72 
Pleurostachys spersiflora, &2 Rapanea latifolia, 72 
Pleurostachys stricta, 82 Recordia, 99, 101, 171, 172; 
Pleurostachys tenuiflora, 82 174 

Pleurostachys Urvillii, 82 Recordia boliviana, 99, 105, 
Pleurothyrium bifidum, 121 fae 417 

Pleurothyrium Krukovii, 121 Rhododendron, 83 
Pleurothyrium nobile, 120,121 Rhodolaena Bakeriana, 84, 86, 


Pleurothyriun parviflorun, &8 

121, 122 Rhodolaena parviflora, &, 
Pleurothyrium Poeppigii, 121 86, 88 
Podocarpus, 212 Rhynchanthere paludicola, 136 
Podocarpus Matudai, 212 Rhynchospora stricta, @1 
Polygalaceae, 111 Rhynchospora Urvillei, 82 
Polymeris, 17 Roraimanthus, 39 


Polypompholyx leciniata, 138 Roreimanthus Imthurnienus, 40 
Polvypompholyx longeciliata, Roucheria humiriifolia, 8 


138 Rubiaceae, 205, 254, 377 
Priva, 442 Ruellia prismatica, 82 
Priva pedicellata, 429 Ruellia quadrangularis, &2 
Prunus axitliana, 469 Ruellia Schaueriana, 8&2 
Prunus erythroxylon, 369 Ruellia solitaria, 82 
Prunus Lundelliana, 369 Rutidanthera, 28 
Prunus Matudai, 369 Ruyschia, 246 
Psanmisia nitida, 131 Ruyschia clusiaefolia, rigs 
Psammisia rupestris, 131 Ruyschia enervia, 245, 2 


Pseudeerziphila, 192 Ruyschia mexicana, 245 


1941 Index 509 


Ryania, 106 Siparuna niceraguensis, 127 
Ryania acuminata, 106 Sloanea Garckeana, 122 
Ryania Candollei, 106 Sloanea reticulata, 122 
Ryania canescens, 106 Solanaceae, 205, 250 

Ryania chocoensis, 106 Solanum atropurpureum, 75 
Ryania dentata, 106 Solanum tenuispinum, 75 
Ryania Mansoana, 106 Solenophora insienis, 150 
Ryania Riedeliana, 106 Solenophora Pirena, 149, 150 
Ryania Sagotiana, 106 Somphoxylon, oe 

Ryania sauricida, 106 Souroubea, 246 

Ryania speciosa, 106 Souroubea exauriculeta, 245 
Ryania stipularis, 106 Souroubea triandra, 2 244, 245 


Rynchospora, 81 Spartothamnella te teucriiflora, 
Rynchospora Beyrichii, 81 430 


Rynchospora fluminensis, 81 Spartothamus teucriiflorus, 
Rynchospora foliosa, 81 430 
Rynchospora Gaudichaudii, 81 Sphaerochloa, 311 


Rynchospora graminifolia, 81 Sphaerochloa com ressa, 412 
Rynchospora Martiana, 81 Sphoerochloa, 311 
Rynchospora millegrana, 81 Sphoerochloa compressa, 312 
Rynchospora Orbigniana, 81 Spigelia polystachya, 138 
Rynchospora panicoides, 81 Stachyterpheta australis, 470 


Rynchospora puberula, 82 Stechytarphets azurea, 470 
Rynchospora scalaris, 82 Stachytarpheta Boldinghii, 471 
Rynchospora sparsiflora, 82 Stachytearpheta Calderonii, 
Rynchospora tenuiflora, 82 455, 472 

Saccolenia dimorpha, 37 Stachiter hete cayennensis, 
Sagraea, 45 155 


Sarcolaena codonochlamys, 84, RE or dichotoma, 476 
86, 88 Stachyterpheta elatior var. 
Sarcolaena multiflora, 83, Jenmani, 472 


84, 86, 88 Stechytarpheta Frantzii var. 
Sarcophysa, 38 patentiflora, 430 
Schizolaena cauliflora, 88 Stachytarpheta aterezeneis 
Sciadotenia, 281 var. Lundelliana, 


Scrophulariaceae, 200, 377 Stachyterpheta Hin atic 45) 
Securidaca Corytholobium, 21 Stachytarpheta hybrida, 456 


Securidaca cristata, 20 Stachytarpheta incana, 431 
Senegalia - eed 24 Stachytarpheta incana vare 
Serjania lyrata angustibracteata, 442 
Sideroxylon jintadat. 221 Stachytarpheta indica, 434 
Siparuna, 222 Stachytarpheta indica f. mon- 
Siparuna Archeri, 127 strosa, 43% 


Siparuna episciaefolia, 127 Stachytarpheta jamaicensis, 
Siparuna janensis, 182 


5 
Siparune hemisphaerica, 126, Stachyterpheta Lundellae, 435 


127 Stachytarphete Maximiliani 
Siparune hispida, 128 vere ciliaris, 472 


Siparuna metensis, 128 Stachytarpheta Mexiae, 472 


PHYTOL 
435 


510 


Stachytarpheta mexicana, 
Stachytarpheta miniacea, 


Stachytarpheta mutabilis 
Maxoni x eid 

Stachytarpheta mtabilis 
violacea, 436 


Stachytarpheta peruviana, 473 


Stachytarpheta purpurea, 435 
Stachytarpheta pycnodonta,472 


vere 


vare 


OGIA 
335, 347, 350, 351 


Noe 15 


170. Syngonanthus caulescens, 336, 


350, 360 

Syngonanthus flavidulus, 331, 
336, 343, 349, 350, 352, 
353, 354, 356, 357, 358, 359 
360, 361, 362 

Syngonanthus hondurensis, 344, 


Ts 350, 359 


Stachytarpheta quirosana, 457 Syngonenthus insularis, 345, 


Stachyterpheta roraimensis, 
58 


Stachytarpheta Sprucei, 474 
Stachytarpheta straminea, 475 
Stachytarphete strigosa, 456 


Stechytarpheta subulata, 475 
Stachytarpheta tabascana, 437 
Stachytarpheta trinitensis, 


Z 
Stachytarpheta velutina, 438 


Stachytarpheta Weberbaueri, 
T7 


Stephidium, 175 
Stellaria cuspidata, 54 


Stephanophyllum, 331 
Stigmatococca, 205, 477 
Stigmatococca solanacea, 205 
Stizolobium, 7 

Stizolobium Holtoni, 7 
Stizolobium pruriens, 7 
Stylogyne, 12 

Styracacese, 111 

Svensonia hyderobadensis, 434 
Swietenia, 109, 126 
Swieteniae, 109 


Symphachne, 311 


Symphachne xyrioides, 314 
Symphachne xyroides, 314 


Symphoricarpos, 283 
Symphoricarpos albus, 283 


s hyllanthus Donnell- 
Smithii, 26 


Symphyllanthus plicatus, 26 

Synandropus, 281 

Syngonanthus, 335 

Synponanthus androsaceus, 
335, 251, 354, 358, 361, 
362, 363 


Syngonenthus Bartlettii, 433, 


351, 352, 355 
Syngonanthus lagopodioides, 
345, 351, 352, 353, 355, 

358, 361, 363 

Ss onanthus Lundellianus, 
345, 247, 350 

Syngonanthus Oneilii, 346, 
347, 350, 359, 360 

Syngonanthus ‘Jilsonii, 347, 
351, 353 

Talisia diphylla, 242 

Tamonea, 32 

Tamonea juncea, 169 

Tamonea spicata, 169 

Tamonopsis spicata, 279 

Tapirira, 212 

Tapirira macrophylla, 216 

Tapirira mexicana, 216 

Tassadia apocynella, 15 

Tassania recurva, 15 

Tectona, 154, 155, 162 

Tectona grandis, 155, 156, 162 

Tectona grandis f. abludens, 
156 

Tectona Hamiltoniana, 155, 159 

Tectona philippinensis, 155, 
164 

Tectona ternifolia, 155 

Tectona Theka, 156 

Tectonia, 163 

Tektona, 163 

Theka, 154, 155 

Theka grandis, 156 

Theka ternifolia, 155 

Thibaudia albiflora, 209 

Thibeudia Archeri, 209 

Thibaudia costaricensis, 209, 
210 


Thibaudia paniculata, 209 


1941 Index 511 


Thibaudia parvifolia, 131 Ungulipetalum, 279 


Thyrsanocephalus, 555 Ungulipetalum filipendulum, 280 
Tibouchina asperifolia, 136 Utricularia longeciliata, 138 
Tibouchina brachyphylla, 136 Vacciniaceae, 126, 130, 209 


Tibouchina citrina, 1455 Vaccinium, 83 

Tibouchina erythrophylle, 133 Vaccinium album, 283 

Tibouchina latifolia, 134 Vaccinium Tatei, 71 

Tibouchina nana, 1353 Valerianodes boliviana, 18 . 

Tibouchina nitida, 131 Varronia puianensis, 16 

Tibouchina Oerstedii, 133 Verbena, Ep 

Tibouchins Oerstedii var. Verbena Ballsii, 477 
subsessiliflora, 133 Verbena bonariensis, 171 


Tibouchina oligantha, 134 Verbena Cabrerae, 478 
Tibouchina paludicola, 136 Verbena chamaedryfolia, 480 


Tibouchina rupestris, 131 Verbena Halei f. roseiflora, 
Tibouchina saxosa, 135 459 

Tibouchina venosa, 134, 135 Verbena Harbisonii, 438 
Tinosporeae, A PEO Verbena Hintoni, 439 

Tococa, 175 Verbena litoralis var. slbi- 
Tonina, 347 flora, 432 


Tonine fluviatilis, 319, 347, Verbena nivea, 459 
~ 350, 351, 355, 260, 361,363 Verbena occidentalis, 479 


Torresia cearensis, 136 Verbena officinalis f. rosei- 
Tovomita acuminata, 122 flora, 459 

Tovomita Krukovii, 122 Verbena paraguariensis, 483 
Tovomita macrophylla, 123 Verbena patagonica, 171 
Tovomita micrantha, 123 Verbena peruviana, 480 
Tovomita Riedeliana, 122 Verbena Reineckii, 480 
Tovomita rubella, 123 Verbenaceae, 155, 157, 171, 
Tovomita Sprucean a;,°.123 4i2, 446, 453 

Tragia cordata, 371 Verbenoideae, 101, 172 
Tragia potosina, 370, 371 Vitis aestivalis, 10 

Trema floridana, 43/ Vitis caribaea, 10 

Trema micrantha, 337 Vitis cordifolia, 10 

Trema strigillosa, 437 Vitis novogranatensis, 9 
Trichilia Cipo, 107 Vitis tiliaefolia, 10 


Trichilia Krukovii, 106, 107 yitex Appi, 484 
Trigonia | crassiflora, 129 Vitex cymosa, 17 


Trigonia laevis, 129 Vitex divaricata, 103 
Trigonia membranacea, 128 Vitex erythrocarpa, 232 
Trigonia microcarpa, 129 Vitex Froesii, 485 

Trigonia virens, 130 Vitex puianensis, 486 
Trigoniaceae, 126, 128 Vitex Klugii, 101 

Triumfetta althaeoides, 11 Vitex Krukovii, 487 

Triumfetta boyacana, 10 Vitex littoralis, 161 
Triumfetta mollissima, 11 Vitex lucida, 102 

Triumfetta semitriloba, 11 Vitex Martii, 488 

Techudya robusta, 144 Vitex polygama var. Bakeri,489 


Typha, 83 Vitex polygama var. Dusenii, 


512 PHYTOLOGIA No. 15 


489 Waltheria americana, 65 
Vitex polygama var. Glaziovii Waltheria Ladewi, 

490 Weigeltia bogotensis, 12 
Vitex polygama var. Warmingii Weigeltia Lawrancei, 12 

490 Xerochlamys Boyeriana, 84, 86, 
Vitex quinata var. Wittrocki- 88 

~ dana, 440 Xylolaena Richardii,84,86,88 
Vitex Rusbyi, 103 Xylosteon, 283 
Vitex woe ee vare Xylosteon album, 283 

grandiflora, 490 Xylosteon canadense, 283 
Vitex triflora var. quinque- JZanthoxylum Fagara, 370 

foliolata, 104 Zanthoxylum sonorense, 369, 370 
=e. umbrosa, 103 Zenkera, 143 


Vogelocassia ‘cassia cobanensis, 214 Zygandra, 19 


Index to authors 


Abrams, Le R., 89 

Britton, Ne Le; 23 

Croizat, Le, 433 

pipet He Ao, 1; 255 359;-40, 41,106, 2125-153,_ 141 ee 
0 

Killip, E. Pe, 23, 140 

Krukoff, Be Aes 286 

gate Ce Le, 177, 212, 241, 284, 305, 337, 369, 401, 449, 
81 

Moldenke, H. Ne, 1, 5, 19, 95, 154, 167, 171, 182, 222, 248, 

273, 289, 3509, 343, 364, 372, 409, 433, 453, 483 

Morton, Co Ve; 19, 145, 251 ; 

Rusby, He Hes 49 

Smith, Ae Co, 20, 113, 126, 209 

Smith, Ce Ce; 83 

Smith, Le Be, 81, 138 

Walker, Ee He, 165 


are) eee 


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