#3 PHYTOLOGIA An international journal to expedite botanical and phytoecological publication Vol. 63 August 1987 No. 3 CONTENTS FOOTE, M., The algae of New Jersey (U.S.A.). X Bacillariophyta (Diatoms). A. Fragilariales ............-+--- 137 FOOTE, M., The algae of New Jersey (U.S.A.). XI Bacillariophyta (Diatoms). B. Eunotiales and DI Cr ae oie we arenas cp ce oF ane nibie's Winn eels okie esse 142 FOOTE, M., The algae of New Jersey (U.S.A.). XII The occurrence of Cylindrotheca gracilis (Breb ex Kutz) Grun in the Hackensack River estuary ........---++++++++- 148 LOURTEIG, A., Nomenclatura plantarum americanarum. UT are elas (o's ees are Dd eid iw a oe dn, deren ewe tees 153 DEKKER, J., Mutant weeds of Iowa: fasciation in Taraxacum OPPICHIIE Sy wet ajt ek Se kiere We, slut 5 2s aes ie nyo wine omnes eo ae eee 155 PARFITT, B.D., Echinocereus nicholii (L. Benson) Parfitt, Stat. NOvs(CACHICEDE) i 65). . 6s ne ec ene eee noes 157 ST. JOHN, H., Diagnosis of Hawaiian species of Stenogyne (Labiatae): Hawaiian Plant Studies 139 .........---+++++++: 159 ST. JOHN, H., Diagnoses of Phyllostegia species (Labiatae): Hawaiian Plant Studies 140 .............2222 cece cece eee 172 ST. JOHN, H., Diagnoses of Hawaiian species of Touchardia (Urticaceae): Hawaiian Plant Studies 141...........-----+-- 183 ST. JOHN, H., Diagnoses of new species of Cladocarpa (Cucurbitaceae): Hawaiian Plant Studies 142 ...........---- 185 ST. JOHN, H., Diagnoses of new species of Sicyos (Cucurbitaceae): Hawaiian Plant Studies 143 ...........---- 187 SCHUSTER, R.M., Preliminary studies on Anthocerotae ....... 193 TURNER, B.L., Three new species of Koanophyllon (Asteraceae-Eupatorieae) from western Mexico .........---- 202 MOLDENKE, A.L., Book reviews ...........-220eeeeeeeeeees 207 PE » ees ease ae 'AUG 31 1087 NEy/ YORK BOTANICAL Garter. ay Published by Harold N. Moldenke and Alma L. Mo 590 Hemlock Avenue N.W. : Corvallis, Oregon 97330-3818 — U.S.A. Price of this number $3.00; for this volume $16.00 in advai after close of the volume; $5.00 extra to all foreign addcaniee dl dealers; 512 pages constitute a complete volume; claims f in the mail must be made immediately after receipt of the n number for free replacement; back volume prices apply if received after a volume is closed. THE ALGAE OF NEW JERSEY (U.S.A.) X. BACILLARIOPHYTA (DIATOMS). A. FRAGILARIALES MaryAnn Foote College of Mt. St. Vincent Riverdaie, New York 10471 This is the tenth paper in the series examining the distribution of aigae in the state of New Jersey. The genera are listed alphabetically and the collection cates chronologically within them. Further ecological data may be obtainea from the original sources. Publication was made possible by a Faculty Development Grant, College of Mt. St. Vincent. DIVISION BACILLARIOPHYTA DIATOMS Fragilariales Asterionella glacialis Costracane northern shore (8) Asterionella ralfsii var americana Egg Harbor Stream (9) Diatoma anceps (Ehr) Kirchn Buda Lake (1); New Brunswick (4) Diatoma hiemale (Roth) Heiberg Mullica Hill, Blackwood, Kirkwood, Marlton (1); Oyster Creek (7) Diatoma hiemale var mescdon (Ehr) Grun ‘Metlars Brook (5) Diatoma tenue var elongatum Lyngb Lake Hopatcong (10); Hackensack River (11) Diatoma vulgare Bory Mullica Hilll, Mariton, Drakesville deposit (1); New Brunswick (4); Metlars Brook (5) Diatoma vulgare var linearis VH Lake Hopatcong (10) Fragilaria bicapitata A. Mayer Egg Harbor Stream (9) Fragilaria brevistrata Grun Delaware/Raritan Canal (D/R Canal) (3); Lake Hopatcong (10); Hackensack River (11) 137 138 PHY TO.L'0 6 TA Vol. 63, No. Fragilaria brevistrata var inflata (Pant) Hust Lake Hopatcong (10) Fragilaria capucina Desm Camden, Kirkwood, Woodstown, Sharptown, Marlton, Hamburg, Andover, Culvers Gap (1); New Brunswick (4); Lake Hopatcong (10); Hackensack River (11) Fragilaria capucina var mesolepta Rabh Lake Hopatcong (10) Fragilaria constructa Ehr Egg Harbor Stream (9) Fragilaria constricta var trinodes Egg Harbor Stream (9) Fragilaria construens (Ehr)Grun Hamburg, Andover, Waterloo, Drakesville deposit (1); New Brunswick (4); Egg Harbor Stream (9); Hackensack River (11) Fragilaria construens var binodes (Ehr) Grun Absecon Pond (1); Lake Hopatcong (10) Fragilaria construens var pumile Grun Lake Hopatceng (10) Fragilaria construens var subsalina Hust Egg Harbor Stream (9) Fragilaria construens var venter (Ehr) Grun Lake Hopatcong (10) Fragilaria crotonensis Kitton D/R Canal (3); Barnegat Bay (6); northern shore (8); Lake Hopatcong (10); Hackensack River (11) Fragilaria cylindrus Grun Barnegat Bay (6) Fragilaria floridiana Hanna Oyster Creek (7); Egg Harbor Stream (9) Fragilaria lapponica Grun Metlars Brook (5) Fragilaria leptostauron (Ehr) Hust Lake Hopatcong (10); Hackensack River (11) ragilaria leptostauron var dubia (Grun) Hust Lake Hopatcong (10) 1987 Foote, New Jersey Diatoms 139 Fragilaria investiens (W Sm) A Cl Oyster Creek (7) Fragilaria nitzschoiaes Grun Lake Hopatcong (10) Fragilaria oceanica Cl Cape May Fragilaria pinnata Ehr northern shore (8); Lake Hopatcong (10); Hackensack River (11) Fragilaria stranguiata Egg Harbor Stream (9) Fragilaria vaucheriae (Kutz) Peters Lake Hopatcong (10); Hackensack River (11) Fragilaria virescens Raifs Springs ana brocks (1); D/R Canal (3); New Brunswick (4); Metlars Brook (5); Oyster Creek (7); Egg Harbor Stream (9); Lake Hopatcong (10) Meridion circulare (Grev) Ag Marlton, Woodstown, Sharptown, Blackwood, Kirkwood, Lake Como, Andover (1); New Brunswick (4); Metlars Brook (5); northern shore (8); Hackensack River (11) Meridion circular var constrictum (Ralfs) VH freshwater (1); Metlars Brook (5); Hackensack River (11) Opephora martyii Herib Oyster Creek (7); Lake Hopatcong (10); Hackensack River (11) Opephora marina (Greg) Petit northern shore (8) Opephora pacifica (Grun) Petit Oyster Creek (7) Synedra acus Kutz D/R Canal (3); New Brunswick (4); Lake Hopatcong (10); Hackensack River (11) Synedra amphicephala Kutz Lake Hopatcong (10) Synedra capitata Ehr Hewitts Pond, Drakesville deposit (1) Syneara delicatissima W Ga Lake Hopatcong (10); Hackensack River (11) 140 P By Y .T,.0).L. 0: Gel A Vol. 63, No. Synedra fasciculata (Ag) Kutz Barnegat Bay, Cape May marshes (1); New Brunswick (4); northern shore (8); Oyster Creek (7); Hackensack River (11) Synedra fulaens (Grev) W Sm Shark River, Manasquan River, Shrewsbury River, Barnegat Bay (1); Atlantic City (2) Synedra gaillonii (Bory) Ehr Oyster Creek (7) Synecra incisa 2 Metlars Brook (5); Hackensack River (11) Synedra parasitica (W Sm) Hust Culvers Gap (1); Lake Hopatcong (10); Hackensack River (11) Synecra parasitica var subconstricta (Grun) Hust Lake Hopatcong (10) Synecra pulchella Ralfs ex Kutz Lake Como, Andover, Marlton (1); Metiars Brook (5); Oyster Creek (7); northern shore (8); Lake Hopatcong (10) Synedra pulchella var saxonica (Kutz) Grun Marine, common along coast (1) Synedra radians kutz Mullica Hill, Marlton, Lake Hopatcong, Hamburg (1); D/R Canal (3); Hackensack River (11) Synedra rumpens Kutz Metlars Brook (5); Lake Hopatcong (10); Hackensack River (11) Synedra rumpens var familiaris (Kutz) Hust Metlars Brook (5) Synedra rumpens var fragilariodes Grun Metlars Brook (5); Hackensack River (11) Synedra rumpens var meneghiniana Grun Bag Harbor Stream (9 Synedra socia Wallace Metlars Broox (5) Synedra ulna (Nitz) Ehr state (1); D/R Canal (3); New Brunswick (4); Metlacs Brook (5); Lake Hopatcong (10); Hackensack River (11) Synedra ulna var chaseana Thomas Lake Hopatcong (10) 3 1987 Foote, New Jersey Diatoms 141 Synedra ulna var contracta Ostr Metlars Brook (5) Synedra ulna var ramesi (Herib) Hust Metlars Brook (5) Tabeliaria binalis (Ehr) Grun Egg Harbor Stream (S) Tabellaria fenestrata (Lyngb) cedar swamps (1); Pine Barrens (2); D/R Canal (3); Metlars Brook (5); Egg Harbor Stream (9) Tabellaria fiocculosa (Roth) Kutz state (1); Pine Barrens (2); Oyster Creek (7); Egg Harbor Stream (9); Lake Hopatcong (10); Hackensack River (11) REFERENCES l. Britton, W. 1889. Catalogue of Plants Found in New Jersey. John L. Murphy Pub Co. Trenton, N.J. 2s Boyer, C.S. 1927. Proceedings of Acad Natl Sci 7931-582 3. Renlund, R.W. 1953. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. 4. Keller, J.M. 1954. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. 5. Edgar, R. 1968. M.S. Thesis. Rutgers University. New Brunswick, N.J. ek Mountford, K.. 1969. M.S. Thesis. Rutgers University. New Brunswick, N.J. 7. Hein, M. 1977. M.S. Thesis. Rutgers University. New Brunswick, N.J.] 8. Olsen, P. and M. Cohn. 1979. Bull NJ Acad Sci 24:59-69 9. Patrick, R and N.A. Roberts. 1979. Algae of the Pine Barrens. IN: R.T.T. Forman, Editor. Pine Barrens: Ecosystem and Landscape. Academic Press. N.Y. 10. Beiisle, E. 1983. M.S. Thesis. Rutgers University. New Brunswick, N.J. ll. Foote, M.A. 1983. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. THE ALGAE OF NEW JERSEY (U.S.A.) XI. BACILLARIOPHYTA (DIATOMS). B. EUNOTIALES AND ACHNANTHALES MaryAnn Foote College of Mount St. Vincent Riverdale, New York 10471 This is the eleventh paper in the series examining the distribution of algae in the state of New Jersey. The genera are listed alphabetically and the collection dates chronologically within them. Further ecological data may be obtained from the original sources. Publication was made possible by a Faculty Development Grant, College of Mt. St. Vincent. DIVISION BACILLARIOPHYTA DIATOMS Eunotiales Eunotia arcus Ehr Kirkwood, Mullica Hill, Hammonton, Absecon (1); Oyster Creek (7); Lake Hopatcong (11) Eunotia arcus var bidens Grun Metlars Brook (5) Eunotia bactriana Ehr Tom's River (2) Eunotia bidentula W Gn May's Landing, Tom's River (2); Oyster Creek (7); Egg Harbor Stream (9) Eunotia carolina Patr Egg Harbor Stream (9) Eunotia cordillera Hohn & Hellerman Metiars Brook (5) Eunotia curvata (Kutz) Lagerst White Hall, Marlton, Lake Hopatcong, Kirkwood, Sharptown, Lake Como (1); New Brunswick (4); Egg Harbor Stream (9); Lake Hopatcong (11); Hackensack River (12) Eunotia denticulata var genuina A Cl Hammonton Pon 1 Eunotia diodan Ehr common in ponds and brooks (1) 142 1987 Foote, New Jersey Diatoms 143 Eunotia elegans Ostr Egg Harbor Stream (9) Eunotia exigua (Breb ex Kutz) Rabh Absecon, Hammonton, Toms River, Kirkwood, Mullica Hill (1); Oyster Creek (7); Egg Harbor Stream (9) Eunotia fallax A Cl Absecon Pond (1); Egg Harbor Stream (9) Eunotia flexuosa Breb ex Kutz Hammonton Pond (1); New Brunswick (2); Oyster Creek (7); Egg Harbor Stream (9) Eunotia formica Ehr Egg Harbor Stream (9); Lake Hopatcong (11) Eunotia glacialis Meist Kirkwood, Camden, Marlton, Hammonton, Mullica Hill (1) Eunotia gracilis var mayeri A Cl Cranberry bogs near Ancora (1) Eunotia incisa W Sm ex Greg Mays Landing, Pavonia (1); Egg Harbor Stream (9); Lake Hopatcong (11) Eunotia indica Grun Lake Hopatcong (11) Eunotia lapponica Grun ex A Cl Oyster Creek (7) Eunotia luna Ehr freshwater (2) Eunotia maior (W Sm) Rabh Camden, Lake Hopatcong, Culver's Gap (1) Eunotia microcephala Krasske ex Hust EBgg Harbor Stream (9) Eunotia monodon Ehr common in ponds anc meadow ditches (1); Egg Harbor Stream (9) Eunotia monodon var constricta Cl-Eul Egg Harbor Stream (9 Eunotia naegeli Migula Egg Harbor Stream (9) Eunotia pectinalis (O F Mulier) Rabh Hammonton, Sharptown, Culver's Gap (1); cedar swamps (2); 144 PHY TOL OG LA Vol. 63, No. New Brunswick (4); Metlars Brook (5); Oyster Creek (7); Egg Harbor Stream (9) Eunotia pectinalis var minor Rabh Metlars Brook (5); Oyster Creek (7); Egg Harbor Stream (9); Lake Hopatcong (11) Eunotia pectinalis var recta A Mayer ex Patr Bog Harbor Stream (9) Eunotia pectinalis var unduiata (Ralfs) Rabh Ancora, Mays Landing Hammonton, Absecon, Sharptown, Budd Lake, Lake Hopatcong (1); cedar swamps (2); Egg Harbor Stream (9); Lake Hopatcong (11) Eunotia pectinalis var ventricosa Grun Mays Landing Eunotia praerupta Ehr Mullica Hiil, Kirkwood, Hammonton, Absecon (1); Metlars Brook (5); Oyster Creek (7) Eunotia praerupta var bidens (Ehr) Grun Egg Harbor Stream (9) Eunotia pseucopectinalis Hust Oyster Creek Eunotia repens var arcuata (Naeg) A Cl Oyster Creek (7) Eunotia rostellata Hust ex Patr Metlars Brook (5); Oyster Creek (7) Eunotia septentrionales Ostr Egg Harbor Scream (9) Eunotia serra Ehr Ancora, Kirkwood, Toms River, Culver Gap (1); New Brunswick (4); Oyster Creek (7); Egg Harbor Stream (9); Hackensack River (12) Eunotia serra var diadema (Ehr) Patr Metlars Brook (5); Egg Harbor Stream (9) Eunotia soleirolii (Kutz) Rabh Moorestown Eunotia sudetica O Mull Egg Harbor Stream (9) Eunotia tautoniensis Hust ex Patr Egg Harbor Stream (9) 1987 Foote, New Jersey Diatoms 145 Eunotia tenella (Grun) Cl Metlars Brook (5); Egg Harbor Stream (9); Lake Hopatcong (11) Eunotia triodon Ehr Toms River, Hammonton, Ancover, Culvers Gap, Cape May Point (1) Eunotia vanheurckii Patr Bgg Harbor Stream (9) Eunotia vanheurckii var intermedia (Krass ex Hust) Patr Bgg Harbor Stream (S$) Eunotia zygodon Ehr Toms River (Z); Egg Harbor Stream (9) Peronia fibula (Breb ex Kutz) Ross Metlars Brook (5) Achnanthales Achnanthes biasolettiana (Kutz) Grun Manahawkin salt marsh (10) Achnanthes controversa Hust Oyster Creek Achnanthes exigua var constricta (Grun) Hust Lake Hopatcong (11) ‘Achnanthes fimbriata (Grun) Ross Oyster Creek (7) Achnanthes gibberula Grun New Brunswick (4) — Achnanthes orientalis Hust Bgg Harbor Stream (9) Achnanthes submarina Hust Manahawkin Salt marsh (10) Achnanthes stenera Hust northern shore (8) Cocconeis costata Greg northern shore (8) Cocconeis dirupta var flexella (Jan) Rabh rare in Manasquan River (1 Cocconeis disculoides Hust Oyster Creek (7) 146 Pema y TOLL O°e 1a Vol. 63, No. Cocconeis disculus (Schum) Cl Oyster Creek (7); Hackensack River (12) Cocconeis fluvitilis Wallace HackenSack River (lz) Cocconeis molesta (Kutz) Grun Culver's Gap (1) Cocconeis ornata Greg Atlantic City (2) Cocconeis pediculus Ehr abundant, Statewide (1); Delaware/Raritan Canal (3) Cocconeis placentula Ehr fresh water (1); Metlars Brook (5); Oyster Creek (7); Hackensack River (12) Cocconeis placentula var euglypta (Ehr) Cl Metlars Brook (5); Manahawkin salt marsh (10); Oyster Creek (7); Egg Harbor Stream (9) Cocconeis placentula var intermedia (Her & Per) Thist Oyster Creek (7) Cocconeis placentula var lineata (Ehr) Cl Dennis Creek (1); Metlars Brook (5); Lake Hopatcong (11); Hackensack River (12) Cocconeis pseudodiruptoides Foged Oyster Creek (7) Cocconeis quarnerensis (Grun) AS northern shore (8) Cocconeis scutellum Ehr Sait ana brackish water (1); northern shore (8); Oyster Creek (7) Cocconeis scutellum f parva Grun Oyster Creek (7); Manahawkin salt marsh (10) Cocconeis scutellum var speciosa (Greg) A Cl Oyster Creek (7) Cocconeis scutellum var stauroneiformis W Sm Oyster Creek (7); Manahawkin salt marsh (10) Rhoicosphenia curvata (Kutz) Grun ex Rabh Absecon, Pleasantville (1); New Brunswick (4); Metlars Brook (5); northern shore (8); Hackensack River (12) 1987 Foote, New Jersey Diatoms 147 REFERENCES 1. Britton, W. 1889. Catalogue of Plants Found in New Jersey. John L. Murphy Pub Co. Trenton, N.J. 2. Boyer, C.S. 1927. Proceedings of Acad Natl Sci 7921-582 3. Renlund, R.W. 1953. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. 4. Keller, J.M. 1954. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. 5. Edgar, R. 1968. M.S. Thesis. Rutgers University. New Brunswick, N.J. 6. Mountford, K. 1969. M.S. Thesis. Rutgers University. New Brunswick, N.J. 7. Hein, M. 1977. M.S. Thesis. Rutgers University. New Brunswick, N.J. ] 8. Olsen, P. and M. Cohn. 1979. Buil NJ Acad Sci 24:59-69 9. Patrick, R and N.A. Roberts. 1979. Algae of the Pine Barrens. IN: R.T.T. Forman, Editor. Pine Barrens: Ecosystem and Landscape. Academic Press. N.Y. 10. Sugihara, T. and C. Yearsley, J.B. Durand and N.P. Psuty. Compariscn of Natural and Altered Estuarine Systems: Analaysis. Center for Coastal and Environmentai Studies. Rutgers University. New Brunswick, N.J. ll. Belisle, E. 1983. M.S. Thesis. Rutgers University. New Brunswick, N.J. 12. Foote, M.A. 1983. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. THE ALGAE OF NEW JERSEY (U.S.A.) XII. THE OCCURRENZE OF CYLINDROTHECA GRACILIS (BREB EX KUTZ) GRUN IN THE HACKENSACK RIVER ESTUARY MaryAnn Foote College of Mount St. Vincent Riverdale, New York 10471 Abstract A 16—-month survey of a 15 km stretch of the upper Hackensack River estuary in New Jersey yielded 131 specimens of the pennate diatom Cylindrotheca gracilis (Breb ex Kutz) Grun. Correlation Seertictante indicate a strong positive correlation between C. gracilis and elevated anounts of silica, nitrates and water temperature. Introduction Cylindrotheca gracilis (Breb ex Kutz) Grun is a weakly Silicified and spindle shaped pennate2 diatom of the Family Nitzchiaceae. This species has been reported in Maryland, Ohio and Iowa (1) and Arizona (2) in the United States. Marine littoral samples fron Texas, Georgia, North Carolina, California and Lousiana and seawater samples for Massachusetts, Long Island Sound and Sandy Hook, New Jersey have yielded specimens of Cylindrotheca gracilis (3). It has been reported as a salt or brackish water form (4;5). Lowe (6) considers this taxon to be alkaliphilous, eutrophic, mesohalobic to halophilous, and euplanktonic or periphytic. Czarnecki and Blinn (2) found C. gracilis had its best development in the psammon with a preference for moderate to high conductivity, high alkalinity and periphytic habitats. However, relatively little information is available about the ecology of this species. Tne Hackensack River, lying approximately 10 km west of Manhattan Island, begins near Haverstraw, New York and extends south to Newark Bay, a linear distance of nearly 45 kn. A 16-month survery of the upper Hackensack River estuary in New Jersey yielded 131 specimens of this taxon, making it one of the 30 most abundant species collected. Cylindrotheca gracilis achieved its greatest abundance at a sample site 5.7 kn South of the reservoir dan (site #3) and was never encountered in the freshwater segment of the river (site $1). This paper provides more information concerning the ecology and distribution of Cylindrotheca gracilis. 148 1987 Foote, New Jersey Cylindrotheca 146 Sample Sites The section of the river investigated covered an area from the Oradell Reservoir Dam to just above Overpeck Creek. Site $1 was northernmost and site #5 is most bayward. Site #2 received run-off from a parking lot and also drainage from a duckpond. A bridge at the point where Coles Brook and French Creek enter the river was site #3 and the footbridje at Fairleigh Dickinson University, site #4. Site #5 was a bridge in the city of Hackensack, slightly upstream from an oil depot. Materials and Methods Grab samples of whole plankton were taken along a 15 km segnent of the Hackensack River from July 1980 to October 1931. Chemical analyses of the samples were done using a Hach model DR-EL Portable Water Engineer's Laboratory Kit as outlined in the Hacn Methods Manual (7). Salinity was assessed using the Mohr method of titration with silver nitrate (3). Temperature was determined in situ with a mercury thermometer in the field. Samples were prepared by the incineration technique (9) and cleaned material was mounted in the high refractive medium, Hyrax, as described by Foote (10). The slides were exanined with a Bausch and Lomb stereoscopic microscope equipped with 100X aprochronatic oil immersion objective and 10X occulars. Counts of 250-500 individuals were done for each collection. Niche breadth values were calculated using the formula of McIntire and Overton (11) and correlation coefficient with physiochemical data were computed. Results and Discussion Tadle 1 lists the mean values for some physio-chemical parameters of the Hackensack River, the number of frustules of Cylindrotheca gracilis encountered at each site during the cqurse of the Study and the niche breadth value, Bi. The correlation coefficient values are given in Table 3. A review of the sparse literature concerning this taxon indicates that there is only one specific mention of the diatom being frequent in fresh water but that most researchers have classified it as a frequent estuarin2 form (1). An extensive review of the diatom flora lists over the last century for the state of New Jersey indicate that C. racilis has never been reported for this area. The data presented in Table 1 indicate that C. gracilis was found in a Slightly, rather than strongly mesohaline brackish (3-10 pet) environment. Comparing station $3, where this species achieved its greatest abundance, with the next lower estuarine site, #4, one notices in Table I that there is little difference in temperature and alkalinity but greater differences in the values of salinity, sulfate, magnesium, total hardness, calcium and chloride. This appears to Support the 150 Pury ' O20 6.278 Vol. 63, No. conclusions of Christensen and Reimer (1) that this diatom should not be considered to be a salt water form ani that conductivity and concentrations of sulfate, majnesiun and calciun, or some combination of factors, may be more critical to this taxon's growth and distribution than chloride or sodium ions. When the niche breadth values are calculated for seasons, rather than sanple sites, it is found that C. gracilis grew best in the autumn and winter (5 October 1930 to 22 February 1981). The average values for physio-chemical parameters in the estuarine portion of the river (sites #2-#5) for this period are given in Table 2. The data in this table are somewhat misleading, however, because the river was frozen from sites #2-#4 throughout much of January and this prevented collecting in the upper and middle estuary. The values given in Table 2, therefore, are probably slightly elevated, based primarily on lower estuarine collections. Correlation coefficients listed in Table 3 show a strong positive correlation between Cylindrotheca gracilis and elevated anounts of silica, nitrate anid water temperature at site $3, the site where most of the specimens were collected. It appears that C. gracilis is a species capable of withstanding mesohaline brackish water but it is not a true marine species. Cylindrotheca gracilis also appears to be able to withstand high concentrations of chloride, sulfate, alkalinity, calcium, magnesium ani total hardness. Acknowledgements I very gratefully acknowledge the critical review of these data and manuscript given by Charles W. Reimer. Computer programming was afforded by Michael Levandowsky, Pace University. The publication was made possible by a Faculty Development Grant, The College of Mt. St. Vincent. 1987 Foote, New Jersey Cylindrotheca TABLE 1. Mean chemical and physical parameter values for the Hackensack River estuary and collection data. Values are mg/l unless noted otherwise. STATION 1 2 3 4 5 5 distance fron dan, km 0.07 B52 511 als 1058 ~ 24.6 salinity, ppt 0.11 6 2-OF P42 555 azo temperature, C 16.2 15-9507 .4.0 1628. 17 csmer20s4 alkalinity 74 93 103, =i21 128 144 chloride 4) 947 1152 2122 2961 4153 nardness 121 399 594 831 1133 1498 magnesiun 29 233 493 547 826 1154 calciun 95 152 192 242 274 345 sulfate 238 2 7526S 354 458 nunber of cells 9 3i3. 49 42 3 5 niche breadth value ——) | Sa2 6.5 7.6 3 1.9 TABLE 2. Mean chenical and physical values for the Hackensack River estuary from 5 October 1980 to 22 February 1931. salinity 5.12 pot temperature 5 hs fal alkalinity 120 mg/1 chloride 1139 mg/1 hardness 1160 mg/1 magnesiun §53 m/1 calciun 295 mg/1 sul fate 382 mg/l 151 152 PHY T:0.L:0 G.I A Vol. 63, No. TASLE 3. Correlation coefficients for Cylindrotheca gracilis and 7 hydrographic variables. STATION NUM3ER PARAMETER 2 3 4 5 5 phosphate Pay) 0.02 “0.95 9.05 0.12 sulfate 0.091) 90502 0.03 0.095 > =0508 Silica 0.14 0.64 0.05 0.02) 9 :=0812 dissolved oxygen 091s 9502 0.03 0.004 -0.04 salinity aed 0.02 0.03 '0.004 =O512 nitrate 9.14 9.72 -0.05 0.02 -0.09 water temperature a3 0.52 0.25 0.005 -0.03 References l. Christensen, C.L. and Reimer, C.W. 1958. Iowa Acad Sci 75: 35-41 2. Czarnecki, D.B. and Blinn, D.W. 1939. Diatoms of Lower Lake Powell and Vicinity. Bibl Phycologia 28:1-119 3. Reimann, B.E.F and Lewin, J.C. 1954. Jrl of Royal Micro Soc 33:283-95 4. Cholnoky, B.J. 1953. Die Oxologie der Diatomeen in Binnengewassern. J. Kramer. Lehre. 599 p. 5S. Hustedt, F. 1955. Kieselalgen (Diatomeen). W. Keller and Company. Stuttgart. 70 p. 6. Lowe, R.L. 1974. Environnental requirenents and pollution tolerances of freshwater diatoms. U.S. Government Printing Office. Washington, D.C. 334 p. 7. Hach Chenical Company. 1976. Methods Manual. 12th Edition. Ames, Iowa. 118 p. 8. Strickland, J.D.H. and T.R. Parsons. 1958. Bulletin 157 Fisheries Res Board Canada. 311 p. oS Foote, M.A. 19389. Phytologia 45:95-93 10, - 1982. J. Microscopy 128:199-201 ll. McIntire, C.D. and Werton, W.S. 1971. Ecology 52:758-77 3 NOMENCLATURA PLANTARUM AMERICANARUM V. LAURACEAE Alicia Lourteig Muséum National d’Histoire Naturelle, Paris. PLUMIER describiéd el género Borbonia,dedicado al Prfncipe de Borbon quien adem&és de otras virtudes posefa e] amor a la BotAnica. el cual fué adoptado por Linné d&ndole el nombre de Laurus. Bajo ese nombre descri- bié varias especies, otros autores lo imitaron y antes de un siglo vistos los cearacteres dispares de las especies inclufdas, se separaron varios grupos en diversos géneros. Plumier ilustr6 y describi6 tres especies, pero en su Catalogo de Gé- neros, al describir Borbonia, s6lo cit6 dos (p.4): Borbonia fructu oblongo nigro, calyce coccineo que corresponde al MSS 6: Ic. 98 Borbonia fructu globoso niyro calyce © viride rubente que corresponde al MSS 6: Ic.99 la otra es: Borbonia fructu ex auro virescente, calyce coraliino y corresponde al MSS 63 Ic. 97 Burmann (Edit.) Plant. Americanarum, fasc. 3. 1756 en p&g. 50 cita el primer prot6logo de Plumier, luego las obras de Linné que no correspon- den a ninguna especie de Plumier, da una descripci6n que es una mezcla de especies (de América del Norte y de Antillas) y publica la l4mina 60 en le cual los detalles A = D copiados del Cat8logo de Géneros de Plumier no corresponden a la planta ilustrada por el h&bito; en ésta las hoje presen- tan 5 nervadures secundarias, como en el Ic6n original de Plumier ( 4 6 5) pero, quiz4s por fantas{a del grabador aceptada por Burmann, agrega unas raras nerveduras terciarias que no existen y falsean el icén. Esto fué ob- servado por Urban (Rep. Sp. Nov.Beihiefte 5: 40,0BS.1). Linné en su Systema,1759 y en Species Plant.ed. 2. cita para Laurus Borbonia el primer prot6logo de Plumier y la l&mina 60 de Burmann, que no corresponde. Lamarck, Encyc. p. 450 bajo Laurus Borbonia L. distingue la especie antillana de Plumier como var. f para el prot6logo de Plumier pero cita también el Icdén 60 de’ Burmann que corresponde a otro ( MSS 6: 98). En p. 451 cita Laurus globosus Aublet y tipifica ese binomio (cuyo protélogo fu€é citado por Aublet) con el Ic6n MSS 6: 99 que es el que corresponde, reproducido por Burmann en lémina 60 (s6lo el h&bito con fructificacio~ nes) excluyendo los detalles A - D. Esta especie es de las Islas Antillas La descripcién de Lamarck es la traducci6n franceega de la latins de Plu— mier inédite. Nees, 1836, p. 316, coloca este binomio en la sinonimia de Nectandra Banguinea Roland ex Rotboell (1778), citando adem&s, Lamarck y fereoon, acompafiado al final por un 7? y reafirma la relaci6n con el protélogo y el Ic6n MSS Plumier. Al mismo tiempo sinonimiza Laurus Borbonia var.§ que es otra especie. Expone sus dudas en p. 320 Adnotatio. 153 154 PHY -TOnLs0rGst A Vol. 63, No. 3 Meissner in Candolle, 1864, ignora el binomio de Aublet y describe Nec- tandra antillana, citando especimenes de las Antillas. Mez, 1889, p. 415, hace la combinaci6én Nectandra globosa (Aublet) Mez, citando el bas6nimo y la obra de Lamarck, pero curiosamente todo el mate- rial as{ como la sinonimie son extra-antillanos, correspondiento a una es— pecie guayanesa. A pesar de citar Lamarck parece que no lo ha lafdo. En p. 425 vuelve a citar el binomio en Lamarck pero como sinénimo, esta vez, de N. antillana Meissn. Una de las inconsistencias de la obre de Mez. Urban, 1920, acepta N.antillana pero, como dije m4s arriba, sefials los errores de Burmann en cuanto al uso de los MSS de Plumier y sinonimiza con ? Laurus globosa Lamarck, etc. Rohwer, 1985, fiel a "ez, no ha percibido el] problema de N. globosa. Cita un "halotype BM n.v." inexistente, puesto que Aublet ni vi6 ni cité material alguno, y Lamarck fué quien tipific6 el binomio con la ilustra— cié6n del protélogo citado por Aublet. Este autor s6lo puso el protélogo de Plumier en nomenclature binomiel (V. Phytologia, 53 (4): 245-246. 1983). De esta situaci6n confusa, resulta evidente que Nectandra globosa(Au- blet) Mez no es aplicable al material de la regién quayanesa citado por Mez, sino al de Antillas, lo que corresponderfa a Nectandra antillana Meissner. El material citado, de Gusyana tendr& que llamarse quiz4s, se — gGn alguno de los sinénimos citados por Mez u otro binomio prioritario. La nomenclatura es como sigue: —_ ee MSS 6: 99, reppoducido por Burmann, l.c. excl. detalles A —- D. Laurus globosa Aublet, Fl. Guiana Frang. 1: 364. 1775. Lamarck, Encyc. Mé- thod.3: 451, 1792. Persea globosa (Aublet) Sprengel, Systema 2: 269. 1825. Nectandra globosa (Aublet) Mez, Bot. Jahrb. 5: 415. 1889 excl. descr., si- nén. y material. Rohwer, Mitteil. Inst. Allgem. Bot. Hamburg 20: 51. 1985. Sinénimos: Nectandra antillana Meissner in Candolle, Prodromus 15 (1): 153. 1664, Mez, loc. 425. Urban, Symb. Antill. 6:233. 1920 et 1l.c. Adams, Flowe- ring Pl. Jamaica 253, 1972, Fournet, Fl. Illustrée Phanér. Guadeloupe et Martinique 492 - 493. 1978. Rohwer, 1.c. 52 = 53 (in syn.). MUTANT WEEDS OF IOWA: FASCIATION IN Taraxacum officinale Jack Dekker, Agronomy Department, Iowa State University, Ames, IA 50010 and Robin G. Dekker, RR #1, Madrid, Iowa 50156 Fasciation in plants causes a wide variety of flattened, coalesced and malformed morphological changes in vascular plants. These include a widening or broadening of shoot axis parts, enlargements of plant parts and a fusing of vegetative and reproductive parts. These changes can affect single plant parts or the entire plant. Fasciation is common in many families of vascular plants (White, 1945, 1948). This trait is often inherited but can occur with environmental, or exogenous chemical, stimuli (Gorter, 1965; Scheibe and Wohrmann-Hillmann, 1957; and White, 1948). The purpose of this report is to describe the recent finding (May, 1987) of fasciation in dandelion (Taraxacum officinale). Ten to 15 plants were observed in an abandoned pasture in Douglas Township, Boone Co., Iowa. The plants were localized in an area about 30 m by 20 m. They were the only fasciated members of about 20,000 - 30,000 plants in a local community with, primarily, Kentucky bluegrass (Poa praetense). The fasciation took two forms, both confined to the reproductive portions of the plant. Plant tissue in all cases appeared healthy with normal coloration. Of multiple peduncle stalks, the central peduncles were fused together to form a broad (1-2 cm) flower stalk. Only a single, central, fasciated peduncle appeared in a single plant. Individual stems were visible in the fused stalk, often 2-4 peduncles appeared together. The second evidence of fasciation was in multiple flower heads. Flower heads (2-4) were fused at their base and formed a longitudinal organ. Flower color and seed production appeared normal. All other plant parts appeared normal: leaves and taproots. Seed and asexual organs were collected for further studies. This report represents the first of a series of reports on naturally occurring mutants of the Iowa flora. Literature Cited Gorter, C. J. 1965. Origin of fasciation. In Handbuch der Pflanzenphysiologie, W. Ruhland, ed., Pp. 330-351. Vol. 15. Springer, New York. Scheibe, A. and B. Wohrmann-Hillmann. 1957. Gene-controlled developmental processes in the stem structure of pea. Z. Bot. 45:97-121. 155 156 PHYTOL 0 G1A Vol. 63, No. 3 The biology of fasciation (and its relation White, 0. E. 1945. Herid. 36:11-22. to abnormal growth). J. White, 0. E. 1948. Fasciation. Bot. Rev. 14:319-358. ECHINOCEREUS NICHOLII (L. BENSON) PARFITT, STAT. NOV. (CACTACEAE) Bruce D. Parfitt Department of Botany, Arizona State University, Tempe, AZ 85287 For the past 43 years the yellow-spined southern Arizona hedgehog with the pale pink flowers has been known as a variety of Echinocereus engelmannii. Recent studies have shown it to be a genetically isolated, morphologically distinct species. Echinocereus nicholii (L. Benson) Parfitt, stat. nov. Basionym: Echinocereus engelmannii (Parry ex Engelm.) Lemaire var. nicholii L. Benson. 1944. Proc. Calif. Acad. Sci. ser. 4, 25:258. Type: Arizona, Pima County, Silver Bell Mountains. 28 March 1941. L. Benson 19729 (ARIZ 24989). During an ongoing survey of chromosome numbers in the Cactaceae of the western United States, E. nicholii was found to be diploid, 2n=22 (Appendix 1), whereas all varieties of E. engelmannii for which chromosome numbers are known are tetraploid, 2n=44 (Parfitt 1978; Pinkava & McLeod 1971; Pinkava & Parfitt 1982; Pinkava et al. 1977, 1985, unpubl.). This difference in ploidy level represents more than a diagnostic character; it represents a reproductive barrier between the two taxa. Any hybridization that may occur between them would result in a sterile triploid (2n=3x=33), effectively blocking the flow of genes between the parent taxa. The discovery that E. nicholii is diploid led to a closer examination of the morphological differences between it and the tetraploids. Although they are similar in size and in the presence of a downward-pointing flattened spine, E. nicholii may be readily separated from E. engelmannii by a greater distance between the areoles on each rib, flowers pale pink instead of medium magenta, basal portion of the floral cup green instead of colored, and smaller seeds with large, distinct papillae instead of low, coalescent papillae. ACKNOWLEDGEMENTS I wish to thank Wendy C. Hodgson, Allan D. Zimmerman (both DES) and Donald J. Pinkava (ASU) for their contributions to the study. LITERATURE CITED Parfitt, B.D. 1978. Cactaceae, in A.Lbve, ed., IOPB chromosome number reports. LIX. Taxon 27:54. 157 158 PHY 2TOLO0¢ TA Vol. 63, No. 3 Pinkava, D.J. & M.G.McLeod. 1971. Chromosome numbers in some cacti of western North America. Brittonia 23(2):171-176. , L.A.McGill & T.Reeves. 1977. Chromosome numbers in some cacti of western North America. III. Bull. Torrey Bot. Club 194(2) 3195-119. & B.D.Parfitt. 1982. Chromosome number in some cacti of western North America. IV. Bull. Torrey Bot. Club 199:121-128. , M.A. Baker, B.D.Parfitt, M.W.Mohlenbrock & R.D.Worthington. 1985. Chromosome numbers in some cacti of western North America. V. Syst. Bot. 19(4) :471-483. Appendix 1. Documentation for chromosome number determinations in Echinocereus nicholii: n=11 AZ, Pima County: TYPE LOCALITY, SW side of Silver Bell Mountains, elev. 2689 ft., Parfitt 3579 and 3573 with A. Zimmerman (ASU). All determinations are from meiotic anther material prepared according to the methods of Pinkava & McLeod (1971). DIAGNOSIS OF HAWAIIAN SPECIES OF STENOGYNE (LABIATAE) Hawaiian Plant Studies 139 Harold St. John Bishop Museum, Box 19000A, Honolulu, Hawaii, 96817, USA. A lengthy study by the author of Hawaiian Stenogyne has produced a large manuscript. Since this is not ready to publish, it is logical to print the diagnoses of the new species and varieties, in order to make them available. The types, unless otherwise located, are in the Bishop Museum, Honolulu. Stenogyne acuta sp. nov. Ramulis puberulis, petioles 5-8 mm longis, laminis 4.5-6 cm longis ellipticis, calycibus 8.5-10.2 mm longis, corollis 21 mm longis pilosis, loba infera 4.5 mm longa, ovata. Typus: Hawaii I., Volcano, F. R. Warshauer 1,938. S. adpressa sp. nov. Ramulis pilosulis, petioles 10-17 mm longis pilosis, laminis 2-4 cm longisanguste ellipticis infra pilosulis, nodis 18-22-floriferis, calycibus 10-12 mm longis. Typus: Hawaid 2., Cu N. Forbes 1183.0... 1n part. S. adscendens sp. nov. Ramulis hirsutulis, petiolis 15-30 mm longis hir- sutulis, laminis 4-6.5 mm longis ellipticisinfra in nervis hirsutis, nodis 2-floriferis, calycibus 5 mm longis, corollis 12 mm longis puberulis, loba infera 1 mm longa ovata. Typus: Hawaii I., L. W. Cuddihy 893. S. alakaiensis sp. nov. Ramulis pilosulis, petiolis 6-13 mm longis pilosulis, laminis 3.5-6.5 cm longis lanceolatis, nodis 6- (4)- floriferis, calycibus 9 mm longis puberulis, corollis 26 mm longis puberulis, loba infera 5 mm longa ovata. Typus: Kauai I., R. Gustafson 1,061. Sy alba isp nove Ramulis pilosulis, petiolis 5-8 mm longis pilosulis, laminis 3.5-4.7 cm longis ellipticis infra pilosulis, nodis 4-6-floriferis, calycibus 9-10 mm longis pilosulis, corollis 15 mm longis pilosulis, loba infera 1.6 mm longa ovata. Typus: Hawaii I., W. Hillebrand 397. S. albimontis sp. nov. Ramulis glabris, petiolis 8-15 mm longis ciliatis, laminis 1.5-4.3 cm longis ovatis, glabris, nodis 6-flor- iferis, calycibus 13 mm longis puberulis, corollis 19 mm longis puberulis, loba infera 3.3 mm longa suborbiculari. Typus: Hawaii I., C. N. Forbes 469.H. S. alternans sp. nov. Ramulis cum valleculis alternatis pilosulis, pet- iolis 5-8 mm longis pilosis, laminis 22-43 mm longis ovatis\infra nervis pilosulis, nodis 6-floriferis, cal- ycibus 6-8 mm longis, corollis 17 mm longis, loba infers 159 160 PH Yr O L*0 Gir A Vol. 63, No. 3 3 mm longa, ovata. Typus: Hawaii I., H. McEldowney 19. S. ambiobtuta sp. nov. Ramulis pilosulis, petiolis 12-19 mm longis pilosulis, laminis 5-6 cm longis ellipticis infra pilosulis, nodis 6-floriferis, calycibus 10-11 mm longis pilosulis, cor- ollis 18 mm longis pilosulis. Typus: Hawaii I., S. Anderson 489. S. angularis sp. nov. Ramulis hirsutis, petiolis 6-20 mm longis hirsutis, laminis 18-40 mm longis ovatis hirsutis, nodis 6-florif- eris, calycibus 11-12.5 mm longis hirsutis. Typus: Hawaii I.,EF. R. Warshauer 1,996. S. bracteosa sp. nov. Ramulis pilosulis, petiolis 6-11 mm longis ciliatis, laminis 17-27 mm longis ovatis, glabris, nodis 6-flor- iferis, calycibus 5.5=6.5 mm longis glabris, corollis 22-23 mm longis pilosulis, loba infera 3.3 mm longa ovata. Typus: Hawaii I., J. Davis 946. S. calaminthoides A. Gray, var. kulaniensis, var. nov. Calycibus 8-9 mm longidpuberulis, nodis 8-10-flori- feris, ramulis pilosulis, petiolis pilosulis. Typus: Hawaad 1 4,)\H. St.gohn (25,320. S. calvata sp. nov. Ramulis in lateribus pilosulis, petiolis 3-7 mm longis ciliatis, laminis 1.9-3.9 cm longis ovatis infra in nervis pilosis, nodis 6-floriferis, calycibus 6.5-7 mm longis pilosis, corollis 15.5 mm longis pil- osis, loba infera 3 mm longa ovata. Typus: Hawaii lI., G. Clarke 109. S. (Glliata sp. nev. Ramulis pilosulis, petiolis 3-7 mm longis cil- iatis, laminis 10-13 mm longis elliptici-ovatis infra pilosulis, nodis 2-floriferis, calycibus 8-8.5 mm longis puberulis, corollis 22 mm longis puberulis, loba infera 3.5 mm diametri suborbiculari. Typus: Hawaii I., Hosts Jon 26,962. Ss. conscandens sp. nov. Ramulis in angulis pilosis, petiolis 3-7 mm longis ciliatis, laminis 5-10 mm longis ovatis ciliatis,.nodis 2-floriferis, calycibus 7.5 mm longis ciliolatis, corollis 18-19 mm longis pilosulis, loba infera 3 mm longa ovata. Typus: Hawaii I., K. M. Nagata 1,689. S. coriacea sp. nov. Ramulis glabris, petiolis 6-8 mm longis pilosulis, laminis 28-37 mm longis ellipticis glabris, nodis 3-4-floriferis, calycibus 11-12.5 mm longis puberulis, corollis 13 mm longis. Typus: Hawaii I., C. Wakida 326. S. Davisae sp. nov. Ramulis pilosulis, petiolis 6-9 mm longis ciliatis, laminis 18-27 mm longis ovatis glabris, nodis 4-6-flor- iferis, calyeibus 6.5-7 mm longis glabris, corollis 1987 St. John, Hawaiian Stenogyne 161 23 mm longidpilosis, loba infera 5 mm longa. Typus: Hawaii Tr, J. Dava's “490. S. decumbens sp. nov. Ramulis puberulis, petiolis 3-8 mm longis ciliatis, laminis 1.3-2.7 cm longidellipticis glabris, nodis 6-flor- iferis, calycibus 7-8 mm longis pilosulis, corollis 12-14 mm longis puberulis, loba infera 3.3 mm longa elliptica. Typus: Hawaii I., H. McEldowney 20. S. deltoidea sp. nov. Ramulis hirsutulis, petiolis 7-13 mm longisciliatis, laminis 2-3.8 cm longis ovatis, glabris, nodis 6-florif- eris, calycibus 9-10 mm longis; corollis 35 mm longis pil- osulis, loba infera 5 mm longa elliptica. Typus: Hawaii Io, CoilNashOnbes O76O—H - S. diantha sp. nov. Ramulis pilosulis, petiolis 5-9 mm longis ciliatis, laminis 4.5-5.8 cm longis ellipticis glabris, nodis 2-floriferis, calycibus 9-10 mm longis pilosulis, cor- ollis 20 mm longis pilosulis. Typus: Hawaii I., F. R. Warshauer 1,938. S. Douglasii sp. nov. Internodis glabris, petiolis 8-18 mm longis ciliatis, laminis 1.8-6.2 cm longis ellipticis infra\in nervis pilosulis, nodis 10-14-floriferis, calycibus 7.5 mm longis puberulis, corollis 16 mm longis puberulis, loba infera 2 mm diametro suborbiculari. Typus: Sandwich Is. D. Douglas 22 (K). S. earina sp. nov. Internodis glabris, petiolis 2-10 mm longis glabris, laminis 25-40 mm longis eliptici-lanceolatis glabris, nodis 3-6-floriferis, calycibus 8.5-9 mm longis puber- ulis, corollis 17-18 mm longis puberulis, loba infera 1.8 mm longa ovata. Typus: Hawaii I., C. Wakida 325. S. elliptica sp. nov. Ramulis hispidulis, petiolis 2-4 mm longis ciliatis, laminis 2.2-3.1 cm longis ovatis infra hispidulis, nodis 2-6-floriferis, calycibus 10-12 mm longis his- pidis, corollis 22-28 mm longis hispidulis, loba infera 3-4.5 mm longa ovata vel suborbiculari. Typus: Sandwich Is., D. Nelson (BM). S. elliptilobata sp. nov. Nodis 6-floriferis, calycibus 16-17 mm longis pilosulis, corollis 40 mm longis pilosulis, loba in- fera 5.5 mm longa oblate orbiculari. Typus: Hawaii I., J. Jacobi 360. Steitlavaisp., nov. Ramulis in valleculis alternatis pilosulis, ramis lateralibus cum petiolis 6-12 mm longis ciliatis, laminis 2.5-5 cm longis ellipticis infra in nervis pilos- ulis, nodis 6-floriferis, calycibus 11-12 mm longis pilosulis, corollis 19-20 mm longis pilosulis, loba infera 1.5 mm longa ovata. Typus: Hawaii I. J. Davis 555. 162 PH) AP O10 GORA Vol. 63, Nos S. Forbesii (Sherff) comb. nov. S. purpurea H. Mann, var. Forbesii Sherff, Am. Journ. Bot... 215) 701;,°:1934.. S. f£ruticosa, sp... nov. Ramulis pilosulis, petiolis 6-7 mm longis ciliatis, laminis 16-26 mm longis lancei-ovatis infra in nervis pil- osis, nodis 6-4-floriferis, calycibus 8.5-9 mm longis pilosulis, corollis 15 mm longis pilosulis, loba infera 3 mm longa ovata. Typus: Hawaii I., O. & I. Degener 34,639. © (G). S. fusca sp. nov. Internodis glabris, petiolis 10-21 mm longis glabris vel puberulis, laminis 4-7 cm longiselliptici-lanceolatis glabris, nodis 6-floriferis, calycibus 6-8 mm longis, sub- glabris, corollis 15 mm longis puberulis, loba infera 2.6 mm longa ovata. Typus: Hawaii I., F. R. Warshauer 3,131. S. gracilis (Sherff) comb. nov. S. macrantha Benth., var. gracilis Sherff, Bot. Gaz. 96: 140, 1934. S. hamakuaensis sp. nov. Ramulis glabris, petiolis 10-22 mm longis ciliatis, laminis 4-6 cm longis eee ae in nervis hirsut- is, nodis 8-6-floriferis, calycibus 8-%.5 mm longis pub- erulis, corollis 26-28 mm longis puberulis, loba infera 7 mm longa ovata. Typus: Hawaii I., W. Hillebrand 344A (K). F S. hawaiiensis sp. nov. Ramulis glabris, petiolis 9-17 mm longis laminis 4-5.5 cm longis ovatis infra glabris, nodis 6-floriferis, caycibus 13 mm longis puberulis, corollis 32 mm longis pilosis, loba infera 6 mm longa semiorbiculari. Typus: Hawaii I., J. Remy 377 (P). S. hemidasys sp. nov. Lateribus alternatis ramularum pilosis, petiolis 2-3 mm longis ciliatis, laminis 5-11 cm longis ovatis, nodis 2-floriferis, calycibus 6-6.6 mm longis pilosulis, corollis 12-13 mm longis pilosulis, loba infera 2 mm longa ovata. Typus: Hawaii I., S. Anderson 439. S. hiloensis sp. nov. Ramulis pilosis, petiolis 7-12 mm longis pilosulis, laminis 2.5-3.7 cm longis ovatis infra puberulis, nodis 10-8-floriferis, calycibus 6-7 mm longis, corollis 34 mm longis puberulis, loba infera 4.5 mm longa reniformi. Typus: Hawaii I., J. Davis 670. S. Hosakae spl nov. Herbis glabris, petiolis 8-15 mm longis, laminis 3-5 cm longis, nodis 2-floriferis, calycibus 7.5-8 mm longis, corollis 15-18 mm longis puberulis, loba infera 2.5 mm longa elliptica. Typus: Hawaii I., E. Y¥. Hosaka Pee 9 a 1987 St. John, Hawaiian Stenogyne 163 S. huluhuluensis sp. nov. Ramulis pilosis, petiolis 2-4 mm longis pilosulis, lam- inis 3-7 mm longis ovatis infra pilosulis, nodis 2-flor- iferis, calycibus 7.5-8 mm longis pilosulis, corollis 20-21 mm longis, loba infera 3 mm longa ovata. Typus: Hawaii I., L. W. Cuddihy 912. S. humululaensis sp. nov. Internodis glabris, petiolis 5-25 mm longis, laminis 2-8.5 cm longis ellipticis infra glabris, nodis6-4-flor- iferis, calycibus 7.5-9 mm longis ciliatis. Typus: Hawaii I., F. R. Warshauer 3,125. S. integra sp. nov. Ramulis glabris, petiolis 0.5-2 mm longis, laminis 2-3.5 cm longis elliptis glabris, nodis 2Ufloriferis, calycibus 11-12 mm longis puberulis, corollis 20 mm longis pilosulis, Typus: Maui I., C..N, Forbes 2,437.M. S. kaalae Wawra, var. Obatae var. nov. Ramulis glabris, petiolis ciliatis, laminis in basi rotundatis, nodis 2-floriferis, calycibus 13 mm longis puberulis. Typus: Oahu I., J. Obata. S. kaalae Wawra, var. Pearsallii var. nov. Ramulis glabris, calycibus 9 mm longis glabris. Typus: Oahu I., G. Pearsall. S. Kamehamehae Wawra, forma albiflora (Sherff) comb. nov. S. Kamehamehae Wawra, var. albiflora Sherff, BoterGcaze. 9620 139),, 1934. S. kauaiensis sp. nov. Ramulis pilosis, petiolis 5-9 mm longis pilosis, laminis 4-5.8 cm longis ovatis infra in nervis pil- osulis, nodis 6-8-floriferis, calycibus 10-12 mm longis puberulis, corollis 40 mm longis pilosulis, loba infera 7 mm longa ovata. Typus: Hawaii I., J. Davis 662. S. kilaueae sp. nov. Ramulis pilosulis, petiolis 8-10 mm longis pilosulis, laminis 3.5-5 cm longis ellipticis infra in nervis pilosulis, calycibus 6-7 mm longis pilosulis, corollis 17 mm longis pilosulis, loba infera 2 mm longa ovata. Typus: Hawaii I., O. Degener 5,456. S. kipahuluensis sp. nov. Ramulis pilosis, petiolis 10-23 mm longis pilosis, laminis 2.2-5 cm longis ovatis infra in nervis pilosis, calycibus 12 mm longis puberulis, corollis 5.7 cm longis puberulis, loba infera 7 mm longa semiorbiculari. Typus: Maui I., C._ A. Russell 545, S. konaensis sp. nov. Ramulis pilosis, petiolis 8-16 mm longis pilosis, laminis 23-40 mm longis ovatis infra pilosulis, nodis 6-4-floriferis, calycibus 7-9 mm longis pilosis, cor- ollis 28-30 mm longis pilosis, loba infera 4-4.5 mm longa ovata. Typus: Hawaii I., F. R. Warshauer 2,011. 164 P'H ¥ TO) L0G TA Vol. 63, Noo S. kukuiensis sp. nov. Internodis glabris, petiolis 1.5-2.5 mm longis hir- sutis, laminis 2.2-2.5 cm longis ellipticis glabris, nodis 2-floriferis, calycibus 12 mm longishirsutulis, corollis 22 mm longis pilosulis, loba infera 3.3 mm longa rotun- date oblonga. Typus: Maui I., R. Hobdy 2,231. S. kulaensis sp. nov. Ramulis hirsutis, petiolis 1-2 mm longis hirsutis, laminis 4-7 mm 4-7 mm longis ellipticis hirsutis, nodis 2-floriferis, calycibus 4.3-5 mm longis pilosulis, corollis 16 mm longis pilosis, loba infera 3 mm wet! deltoidea. Typus: Maui I., R. Hobdy 2,231. S. lactea sp. nov. Ramulis hirsutulis, petiolis 30 mm longis hirsutis, laminis 5.2-8.5 cm longis ovatis infra puberulis, nodis 6-floriferis, calycibus 16-19 mm longis pilosis, corollis 36 mm longis pilosulis, loba infera 6 mm longa orbiculari. Typus: Hawaii I, J. D.. Jacobi 360). S. laculata sp. nov. Ramulis pilosulis, petiolis 10-25 mm longis ciliatis, laminis 25-55 mm longis ovatis infra pilosulis, nodis 6-4-floriferis, calycibus 6-8 mm longis pilosulis, cor- ollis 23-25 mm longis pilosulis, loba infera 10 mm longa Ovata. Typus: Hawaii I., F. R. Warshauer 1,857. S. laevis sp. nov. Ramulis glabris, petiolis 7-22 mm longis ciliatis, laminis 5-10.5 cm longis lanceolatis infra midnervo sub- hirsuto, nodis 6-floriferis, calycibus 8-9 mm longis pil- osulis, corollis 21 mm longis pilosulis, loba infera 2.5 mm longa subovata. Typus: Hawaii I., H. St. John 18,453. S. lanceolata sp. nov. Ramulis puberulis, petiolis 2-5 mm longis ciliatis, laminis 25-27 mm longis lanceolatis glabris, nodis 2-4- floriferis, calycibus 7-6 mm longis puberulis, corollis 16 mm longis pilosulis, loba infera 2 mm longa suborbic- ulari. Typus: Hawaii I., F. R. Warshauer 3,153. ; latisepala (Sherff) comb. nov. S. kaalae Wawra, var. latisepala Sherff,Occas. Papers B. P.. Bishop Mus. 20: 16, fig.) 7.23055 S. leptophylla (Sherff) comb. nov. S. purpurea H. Mann, var. leptophylla Sherff, Bot. Gaz. 96: 142, 1934). S. leptophylla (Sherf£) St... wohn, var. retrorsa, var. nov. Petiolis retrorse pilosulis, laminis infra in nervis pilosulis, calycibus 10-11 mm longis puberulis, labia supera corollae 7 mm longa. Typus: Kauai I., J. F. Rock 5,744. S. lehuensis sp. nov. Internodis in linea unica hirsutulis, petiolis 2-4 mm longis hirsutis, laminis 2-3 cm longis ovatis 1987 St. John, Hawaiian Stenogyne 165 subglabris, nodis 2-floriferis, calycibus 11 mm longis hirsutulis, corollis 22 mm longis hirsutis, loba infera 4 mm longa cordata. Typus: Maui I., R. Hobdy 517. S. linearis sp. nov. Ramulis hirsutis, petiolis 4-8 mm longis ciliatis, laminis 4.5-10 cm longis lanceolatis glabris, nodis 6-floriferis, calycibus 10.5-11 mm longis hirsutulis, corollis 16 mm longis hirsutis, loba infera 2.5 mm longa suborbiculari. Typus: Kauai I., W. Takeuchi 2,100. S. longiloba sp. nov. Ramulis hirsutulis, petiolis 5-12 mm longis cil- iatis, laminis 3-9.7 cm longis glabris, nodis 2-flor-. iferis, calycibus 15-17 mm longis hirsutulis, corollis 10 mm longis hirsutulis, loba infera 2.5 mm longa ovata. Typus: Hawaii I., J. D. Jacobi 1,104. S. lutea sp. nov. Ramulis glabris, petiolis 10-17 mm longis glabris, laminis 2.5-4.5 cm longis glabris, nodis 4-6-floriferis, calycibus 9-10.5 mm longis glabris, corollis 17 mm longis pilosis. Typus: Hawaii I., C. N. Forbes 469.H. S. mamanecola sp. nov. Internodis glabris, petiolis 6-12 mm longis ciliatis, laminis 15-25 mm longis ovatis glabris, nodis 6-floriferis, calycibus 6-7 mm longis pilosulis, corollis 22 mm longis pilosulis, loba infera 3 mm longa ovata. Typus: Hawaii I., S. Anderson 440. S. Meeboldii (Sherff) comb. nov. S. angustifolia A. Gray, var. Meeboldii Sherff, Bull Boab bLshoprMus.. 36, 7/2), E1qenz2on loss S. mollissima (Skottsb.) comb. nov. S. rugosa Benth., var. mollis Sherff, f. mollissima Skottsb., Bot. Notis. 1943: 368, 1943. S. montana sp. nov. Internodis glabris, petiolis 4-9 mm longis cil- iatis, laminis 7-18 mm longis ellipticis glabris, nodis 2-floriferis, calycibus 7-8 mm longis glabris, corollis 16 mm longis pilosulis, loba infera 4 mm longa ellip- tica.. Typus: Hawaii I., H. McEldowney 1. S. montideae sp. nov. Ramulis glabris, petiolis 12-25 mm longis hirsut- ulis, laminis 4-6.5 cm longis lanceolatis infra in nervis pilosulis, nodis 2-4-floriferis, calyxibus 7-10 mm longis palosulis:... Typus:) Hawaii. 1., Bec. 1955) J. (Fs Roek. S. multiflora sp. nov. Ramis glabris, petiolis 5-13 mm. longis ciliatis, lam- inis 3.5-5.3 cm longis elliptis glabris, nodis 10-12- floriferis, calycibus 6-10 mm longis glabris, corollis 18 mm longis puberulis, loba infera 3 mm longa subor- biculari. Typus: Hawaii I.,-C. N. Forbes 334.H. 166 PHYTO L,O1G IeA Vol. 63, Nowes S. oxyodonta (Sherff) comb.- nov. S. calaminthoides A. Gray, var. oxyodonta Sherff, Any iWournie Bot. 21°2 700, V9 sa. S. pedunculata sp. nov. Ramulis glabris, petiolis 10-26 mm longis glabris, laminis 5-8.6 cm longis glabris\lancei-ovatis, nodis 8-20-floriferis, calycibus 9-13 mm longis puberulis. Typus: Hawaii I., Wailuku R., L. W. Bryan. S. pilosa sp, nov. Ramulis pilosis, petiolis 7-15 mm longis pilosis, laminis 4.3-7.5 cm longis ovatis infra in nervis pilosu- lis, no@is 8-floriferis, calycibus 9 mm longis pilosis. Typus: Hawaii I., 0. Degener 5,393. S. pubicostata sp. nov. Ramulis hirsutulis, petiolis 4-9 mm longis puberulis, laminis 3.5-6.2 cm longis lanceolatis infra glabris, nodis 2-floriferis, calycibus 6.5-7 mm longis hirsutulis, corollis 15-16 mm longis pilosulis, loba infera 2.8 mm longa ovata. Typus: Hawaii I. F. R. Warshauer 2,000. S. recta sp. nov. Ramulis pilosis, petiolis 7-10 mm longis pilosis, laminis 2/3-3/8 cm longis ovatis infra in nervis pil- osis, nodis 6-floriferis, calycibus 9-10 mm longis sub- glabris, corollis 19 mm longis, loba infera 4 mm longa ovata. Typus: Maui I., P_.K .Higashino 9,129. S. Remyi (Sherff) comb. nov. S. scrophularioides Benth., var. Remyi Sherff, Bot. Gaz. 962.740, 1934. S. repens sp. nov. Ramulis setosis, petiolis 6-;4 mm longis cil- iatis, laminis 1.3-2.7 cm longis ovatis glabris, nodis 6-floriferis, calycibus 5.5-6 mm longis subglabris, corollis 19-21 mm longis pilosulis, loba infera 4.5-5 mm longa elliptica. Typus: Hawaii I., J. F. Rock 8,314 (A). S. retrorsa sp. nov. Ramu;is pilosis, petiolis 7-9 mm longis pilosis, laminis 15-25 mm longis ovatis infra in nervis pilosulis, nodis 6-floriferis, calycibus 7-8.5 mm longis pilos- ulis, corollis 23-25 mm longis pilosulis, loba infera 4.5 mm longa ovata. Typus: Maui I., F. R. Warshauer 2,543. S. rhuakos sp. nov. Ramulis pilosulis, petiolis 5-14 mm longis glabris, laminis 17-34 mm longis ovatis infra in nervis pilosulis, nodis 2-floriferis, calycibus 8-10 mm longis pilosulis, corollis 8.5 mm longis pilosulis, loba infera 3 mm longa ovata. Typus: Hawaii I. U. S. Exped., sheet 57,505. 1987 St. John, Hawaiian Stenogyne 167 S. nahukuensis sp. nov. Ramulis pilosulis, petiolis 7-12 mm longis ciliatis, laminis 23-40 mm longis ovatis infra in nervis hirsutulis, nodis 6-floriferis, calycibus 6-9 mm longis puberulis, corollis 22-26 mm longis puberulis, loba infera 3.5 mm longa ovata. Typus: Maui I., F. R. Warshauer 2,530. S. nigra sp. now. Ramulis hirsutis, petiolis 6-10 mm longis ciliatis, laminis 20-30 mm longis ovatis glabris, nodis 8-14-flor- iferis, calycibus 5-6 mm longis glabris, corollis 17-18 mm longis pilosulis, loba infera 3 mm longa suborbic- ulari. Typus: Hawaii I., J. Davis 404. S. novalimontis sp. nov. Ramulis hirsutulis, petiolis 7-14 mm longis ciliatis, laminis 2-3.7 cm lonagis ovatis infra puberulis, nodis 6-floriferis, calycibus 6-7 mm longis otaan is. Typus: Hawaii I., F. R. Warshauer 1,415. Sic oblonga (Sherff) comb. nov. S. rotundifolia A. Gray, var. oblonga _ Sherff, Am. JOURN. BOtl 2s OL 934: S. ohiascandens sp. nov. Internodis glabris, petiolis 7-17 mm longis glabris, laminis 11-32 mm longis ovatis glabris, nodis 2-4-florif- eris, calycibus 5-6 mm longis glabris, corollis 14 mm longis, loba infera 1.5 mm longa suborbiculari. Typus: Hawaii I., N. Balakrishnan 931. S. olaaensis sp. nov. Ramulis pilosulis, petiolis 10-23 mm longis cil- iatis, laminis 3-8 cm longis ovatis infra in nervis pilosulis, nodis 6floriferis, calycibus 7-8 mm longis pilosulis. Typus: Hawaii I., N. Balakrishnan 773. S. olowaluensis sp. nov. Ramulis glabris, petiolis 2-5 mm longis glabris, laminis 1.7-4.8 cm longis ovatis glabris, nodis 2-6-floriferis, calycibus 6-8 mm longis puberulis, corollis 19 mm longis pilosulis, loba infera 4 mm longa suborbiculari. Typus: Maui I., C. N. Forbes 2,436.M. S. oonoides sp. nov. Ramulis pilosulis, petiolis 8-12 mm longis pil- osis, laminis 3-5.5 cm longis ovatis infra in nervis puberulis, nodis 3-floriferis, calycibus 8-9 mm longis pilosulis, corollis 23-25 mm longis pilosulis, loba ° infera 4-4.5 mm in diametro suborbiculari. Typus: Hawaii I., Kona, F. R. Warshauer. S. ovata sp. nov. Ramulis pilosulis, petiolis 10-17 mm longis cil- iatis, laminis 3-4.3 cm longis ellipticis infra pub- erulis, nodis 6-floriferis, calycibus 10-13 mm longis puberulis, corollis 35 mm longis pilosulis. Typus: Hawaii I., O. & I. Degener 27,557. 168 Puy Similar to the A. punctatus crispulus phenotype in the cris- pate and freely lamellate thalli; distinct from A. punctatus and vir- tually all other taxa of Aspiromitus in the sharp distinction between a spinose-areolate external spore face and plane, only obsoletely sculptured inner spore faces. Type. North Carolina: Toxaway R., Transylvania Co. (RMS 45231). Known again only from above Jocassee, Estatoe Cr., South Carolina (RMS). This species is described and illustrated in detail in the final volume of The Hepaticae and Anthocerotae of North America; here only the Latin diagnosis is given, since material of the species (labelled Anthoceros appalachianus) by now has been widely distributed during the last three decades. 200 BHY TO L0G LA Vol. 63, No. 3 I know of no member of Aspiromitus in which the strongly spi- nose-foveolate external spore face is contrasted more strongly to the inner (proximal) spore faces. Acknowledgement: I thank Dr. Hannah Croasdale for generously preparing the Latin diagnoses. 4 Footnotes’ t Subfam. Dendrocerotoideae Schust., subfam. n. Subfamilia a An- therocotoidis differens quod (a) sporophyta sine stomatibus; (b) elateres spirales; (c) omnis locellus antheridialis 1 vel 1(2) an- theridia continet. Type: Dendroceros Nees. Aspiromitus subg. Folioceros (Bharadw.) Schust., status nov. Basi- onym: Folioceros Bharadw., Geophytology 1(1):9, 1971. Type: Aspiro- mitus assamicus (Bharadw.) Schust., comb. n. [Basionym: Folioceros assamicus Bharadw., ibid. 1(1):9, 1971]. Megaceros subg. Nothoceros Schust., subg. n. Subgenus a subg. Megacerote differens quod thalli raro ad sparse furcati, segmentis linearibus ad ligulata; a Dendrocerote differens quod sporae tempore liberationis unicellulares. Type: Megaceros endiviaefolius Steph. (M. endiviifolius) of South America; M. giganteus (L. & L.) Steph. of New Zealand also belongs here. 4 Apoceros Schust., subg. n. Subgenus a subg. Dendrocerote differ- ens: (a) costa cavernosa aut lacunosa; (b) alae thalli lacunosae aut perforatae, lacunis magnitudine variantibus. Type: D. cavernosus Hasegawa (1980, p. 306, fig. 11). 5 Aspiromitus appalachianus Schust., sp. n. Species A. punctato crispulo phenotypo similis quod thalli crispati et libere lamellati; distincta ab A. punctato et fere omnibus aliis taxis Asperomiti quod superfacies externa sporae spinoso-areolata bene distincta e super- ficiebus sporarum internis planis et modo obsolete sculptis. References Frye, T. C. & L. Clark. 1937-47. Hepaticae of North America. Univ. Wash. Publ. Biol. 6(1-5):1-1018. . Haessel, G. G. de Menendez. 1963. Estudio de las Anthocerotales y Marchantiales de la Argentina. Opera Lilloana 7:1-297. Hasegawa, J. 1980. Taxonomical studies on Asian Anthocerotae II. Some Asian species of Dendroceros. J. Hattori Bot. Lab. 47: 287-309. 1981. Dendroceros borbonicus. Steph. A new record for Sri Lanka. Misc. Bryol. et Lichen. 9(2):39-41. 1987 Schuster, Studies on Anthocerotae 201 1982. Note on Samoan species of Dendroceros. Ibid. 9:97-100. 1983. Taxonomical studies on Asian Anthocerotae. III. Asian species of Megaceros. J. Hattori Bot. Lab. 54:227-240. 1984. Ibid. IV. A revision of the genera Anthoceros, Phaeo- ceros and Folioceros in Japan. _Ibid. 57:241-272. Jones, E. 1958. An annotated list of British hepatics. Trans. Brit. Bryol. Soc. 3(3):353-374. Mueller, K. 1951-58. Die Lebermoose Europas, in Rabenhorst's Krypt- ogamen-Flora, 3rd ed., 6:1-1365, figs. 1-514. Proskauer, J. 1958. Nachtrag zur Familie Anthocerotaceae, in K. Mueller, Die Lebermoose Europas, Rabenhorst's Kryptogamen-Flora, 3rd ed., 7:1303-1319, figs. 508-14. 1960. Studies on Anthocerotales VI. On spiral thickenings in the columella and its bearing on phylogeny. Phytomorphology 10: 1-19, figs. 1-36. Renzaglia, K. S. & Z. B. Carothers. 1986, Ultrastructural studies of Spermatogenesis in the Anthocerotales. IV. The blepharoplast and mid-stage spermatid of Notothylas. J. Hattori Bot. Lab. 60:97- 104. Schuster, R. M. Boreal Hepaticae, a manual of the liverworts of Minne- sota and adjacent regions. Amer. Midl. Nat. 49:i-v, 257-684. 1966. The Hepaticae and Anthocerotae of North America. Vol. I. Pp. i-xvii, 1-802, figs. 1-83. Columbia Univ. Press, N. Y. Stephani, F. 1909. Dendroceros, eine Gattung der Lebermoose. Sitzb. Naturw. Gesell. Leipzig 35(1908):11-20. THREE NEW SPECIES OF KOANOPHYLLON (ASTERACEAE-EUPATORIEAE) FROM WESTERN MEXICO B. L. Turner Department of Botany, University of Texas, Austin TX 78713 A forthcoming treatment of the Asteraceae of Mexico (Turner & Nesom, in prep.) necessitates description of the following novelties. While described within Koanophyllon this need not mean I favor acceptance of the generic name, as already noted (Turner, 1983). KOANOPEYLLON SINALOENSIS B. Turner, sp. nov. EK. xi affinis sed foliis pubescentibus, capitulis parvioribus, et acheniis hispidis differt. Shrub to ca 2m high. Stems brown, puberulent, the internodes mostly 3-7 cm long; leaves opposite throughout, 11-17 cm long, 3-7 cm wide; petioles 2.5-5.0 cm long, puberulent; blades thin, broadly ovate to somewhat deltoid, the flanges rounded, 3-nerved from the base, moderately pubescent above and below, especially along the veins, atomiferous-glandular, the apices long attenuate, the margins crenulo-serratulate; heads 4-5 mm high, numerous in terminal or axillary cymose-panicles, the ultimate peduncles mostly 3-6 mm long; involucres 2-seriate, 2.2-2.7 mm long; bracts 12-13, linear-lanceolate; receptacle glabrous; florets 13-19 per head; corollas reportedly white or pale lavender, glabrous or a few atomiferous glands present, ca 2.5 mm long, the tube ca 1 mm long gradually flaring into the throat; achenes ca 1.5 mm long, brown, markedly hispid; pappus of 20-30 white bristles, 2.0-2.5 mm long. TYPE: MEXICO. SINALOA: Mcpio. Badiraguato, 3 mi N of Los Ornos along road to Ocurahui, Sierra Suratato, 6500 ft, 3 Nov 1969, D.. E. Breedlove & E. &. Kawahara 16906 (holotype MICH). Additional Specimens Examined: CHIHUAHUA: Mpio. Ocampo, area of Cascada de Basaseachic at the confluence of Rio Basaseachic and Rio Durazo, ca 2 mi S of Basaseachic (28°02'N x 107955'wW), 2100-2200 m, 17-20 Oct 1986, G. Nesom & J. Vorobik 5566 (TEX, MEXU). SINALOA: Mcpio. Sinaloa y Vela, 5 mi NW of Los Ornos along road to Mocorito, 5800 ft, 6 Nov 1969, Breedlove & Kawahara 17091 (MICH). The species is superficially similar to Koanophyllon ij B. Turner (1983), a species of northeastern Mexico with nearly glabrous leaves, larger heads and eciliate, glandular, achenes. - 202 1987 Turner, New Species of Mexican Koanophyllon 203 KOBNOPHYLLON RZEDOWSKII B. Turner, sp. nov. affinis sed foliis parvioribus tenuioribus valde pubescentibus petiolis longioribus et acheniis hispidis differt. Shrub to 1.5 m high; stems densely tan-hirsutulous; leaves opposite, 5-8 cm long, 2.0-3.5 cm wide; petioles 0.5-1.0 cm long; blades broadly ovate, obtuse or rounded at base, 3-nervate, moderately pubescent above, both hirsute and atomiferous-glandular beneath, the margins denticulate to nearly entire; heads numerous, terminal and axillary in corymbose panicles; involucre 4-5 mm high, 3-4 seriate, graduate, the bracts 2-5 mm long, puberulent, narrowly lanceolate, the apices sharply acute; receptacle plane, glabrous; florets 30-34 per head; corollas whitish, glabrous, 3.0-3.5 mm long, the lobes ca 0.5 mm long; anther appendages thin, somewhat flattened, ca 0.25 mm long, 0.20 mm wide, rounded at the apices; achenes ca 2.5 mm long, brown, the ribs 5, yellow, hispidulous, carpopod well-marked, annular; pappus of 20-25 ciliate, tawny, bristles ca 3 mm long. TYPE: MEXICO. SAN LUIS POTOSI: Sierra de San Miguelito, canon arriba de Terrero, 2250 m, 8 Sep 1954, G. C. Rzedowski 4324 (holotype MICH). The holotype was determined by its collector to be Eypatorium aASDGs 2c which is a synonym of E, collinum, as noted by McVaugh 1984). King and Robinson treat this latter taxon within their concept of the genus Chromolaena. Indeed, K, rzedowskii much resembles E, collinum but differs in numerous details (nonurceolate, persistent,involucral bracts, etc.). Nevertheless, it is likely that E, gollinym is better positioned within the Keanophyllon grouping than within Chromolaena itself, as indicated by Whittemore (1987). Keanephyilon rzedowskii is closely related to K xichardsonii B. Turner, which differs in having larger, thiner, nearly glabrous leaves, with much longer petioles, glandular instead of hispid achenes, etc. It is also close to K. xeyrobinsonii of Nuevo Leon. All of these are in turn related to K, longifolia (B. L. Rob.) King & H. Rob. and K, gracicaulis (Sch-Bip. ex B. L. Rob.) King & H. Rob which appear to make up a fairly close-knit species group in KOANOPHYLLON GUERREROANA B. Turner, sp. nov., Fig. 1 a i affinis sed foliis grandibus ellipticis coriaceis et corymbis axillaribus differt. Said to be a "vine"; stems white; leaves opposite, thick and leathery, 15-20 cm long, 4.5-6.0 cm wide, minutely pubescent below 204 PHYTOLOGIA Vol. 63, No. 3 to glabrate; petioles 5 mm long; blades elliptical, prominently 3- nerved from 10-15 mm above the base; heads ca 9 mm high, 10-15,in short axillary clusters, the ultimate peduncles 1-6 mm long; involucres 2-3 seriate, ca 6 mm long; bracts linear-lanceolate, dark, without well-defined nerves, puberulent throughout, the apices acute; florets 18-20 per head; corollas 4-5 mm long, glabrous, the lobes atomiferous-glandular; achenes (immature) ca 2.5 mm long, pubescent; pappus of ca 50 bristles, 4-6 mm long. TYPE: MEXICO. GUERRERO: District Coyuca, Santa Barbara, 6 Apr 1935, G. B, Hinton et al, 7592 (holotype GH). The type and only known collection is depicted in Fig.1. It is very closely related to K. albicaulis (Klatt) King & H. Rob. but is so strikingly differt as to leaf shape, size, and texture, capitulescence and habit (described as a “vine” on the collectors’ label) that I dare describe it as new. At least the collection stands out from among the hundreds of specimens of this plant which I have examined at GH, LL, TEX, UC and US. Nevertheless, it is possible that the collection is from an aberrant, late-flowering individual taken from older parts of the plant, thus perhaps the peculiar capitulescence and thicker, larger leaves. While Koanophyllon albicaulis occurs along the Pacific Coast of Mexico from Chiapas to Sinaloa, I have not previously seen specimens from the states of Guerrero or Oaxaca. Nor had McVaugh (1984), to judge from his account of the taxon for Flora- Novogaleciana. B. L. Robinson recognized the material from northwestern Mexico as Eupatorium albicaule var. laxius B. L. Rob., which was distinguished from the var. albicaule by its loose capitulescence and linear-attenuate involucral bracts. He subsequently elevated the variety to specific rank as i SAASREL B. L. Rob. Neither McVaugh (1984) nor King and Robinson 1971) recognize the varietal taxon, although its treatment as a regional variant of E albicaule would seem to have merit. In any case the present species has involucral characters of var. laxius. The type locality (Santa Barbara) is on the flood plain of the Rio Balsas ("habitat: IRF Balsas” is typed on the label), which is located by Hinton and Rzedowski (1975) at ca 18°16'N = 100°31'W at an elevation of ca 800 m, well above the 300 m limit which McVaugh notes for B, albicaule. ACKNOWLEDGEMENTS I am grateful to GH and MICH for the loan of material and to Dr. Guy Nesom for the Latin diagnoses. Ms. Katy Bear produced the Figure. 1987 Turner, New species of Mexican Koanophyllon 205 LITERATURE CITED Hinton, J. and J. Rzedowski. 1975. George B. Hinton, explorado or Botanico en el Sudoeste de Mexico. Ann. Escuela Nac. Cien. Biol. 21: 1-114. King, R. and H. Robinson. 1971. Studies in the Eupatorieae (Asteraceae) LXIV. The genus Koanophy}lon. Phytologia 22: 147-152. McVaugh, R. 1984. Euypatorium, in Flora Novo-Galiciana 12: 340- 424. Turner, B. L. 1983. Two new species of Koangphyllon (Asteraceae- Eupatorieae) from northeastern Mexico. Phytologia 52: 495- 498. Whittemore, A. 1987. Systematic study of Critonia (Asteraceae- Eupatorieae). Doctoral Thesis, University of Texas, Austin. 206 PHYTOLOGIA Vol. 63, No. 3 Fig. |. Koanophyllon querreroana, holotype. BOOK REVIEWS Alma L. Moldenke "McGRAW-HILL ENCYCLOPEDIA OF SCIENCE AND TECHNOLOGY, VOLUMES 1-20" Sixth Edition, edited by Sybil P. Parker and staff, 12700 pp., 15000+ illustrations, 69 unnumbered color plates. McGraw-Hill Book Company, New York, N.Y. 10011. 1987. $1,600.00. Readers can safely believe all of the "high class" publicity that the McGraw-Hill publishers have put forth about this new edition excellently produced, considerably enlarged and enriched by modernization of content and by new illustrative materials. Just as the previous editions were outstanding in their time, so this new one excels the others and also excels similar presently available works of other publishers. In this new 6th edition there are 1,900 new and 2,000 revised illustrations, 3,500 distinguished experts authoring contributions, measurements given throughout in both the U.S. Customary System and the International System, an Analytic Index of 150,000 entries and an extensive Topical Index, bibliographies given with signed articles, and articles with larger explanations than just the dictionary style so that interrela- tionships between ideas and processes can be better integrated and more broadly understood in the 7,/00 articles. The format and printing make for easier reading than most encyclopedias and the interesting content and style made for "run on" reading. University, school, laboratory, public and many home libraries need this new edition. Congratulations are due to several -- the authors and compilers, the artists and lay-out staff, the publishers and the fortunate owners and/or readers to be. "THE SEAWEED HANDBOOK - An Illustrated Guide to Seaweeds from North Carolina to the Arctic" by Thomas F. Lee, v + 217 pp., 81 black/white line drawings, 17 halftones and 1 map. Dover Publications Inc., Mineola, New York 11501. 1986. $5.95 paperbound. This is a most helpful and interesting book for the naturalist-scuba diver, -glass bottom-boat viewer, -swimmer, -diver, or even -beachcomber who wants to learn about these under saltwater simply structured plants. They are helpfully keyed, well described and effectively illustrated. There is fascinating information about seaweeds generally and more identifying details about 78 species found along the eastern coast of North America. This book is an unabridged, but author-corrected replication of the 1977 first edition from the Mariner's Press of Boston. 207 208 PHY, 1, 0:50) Ga A Vol. 63, No. 3 "ROOT DEVELOPMENT AND FUNCTION" edited by P.J. Gregory, J.V. Lake and D.A. Rose, xiiit+ 206 pp., 70 black/white figures incl. 36 photo. + 11 tables, Cambridge University Press, Cambridge and London, U. K. and New York, N. Y., 1987. $34.50. "This volume is based on the review papers. presented at the Annual Meeting of the Environmental Physiology Group of the Society for Experimental Biology ... with the Association of Applied Biologists and the British Ecological Society" 1985. The xeroxed typing is neat but hard on the eyes. Besides a well organized prologue there are 10 technical papers on such topics as: cellular organization of roots and responses to the physical environment, root tissue functions in nutrient transport, competition between root systems in natural communities, and goals for future research. "LA SELVA NUBLADA DE RANCHO GRANDE PARQUE NACIONAL ‘HENRI PITTIER': El Ambiente Fisico, Ecologia Vegetal y Anatomia Vegetal" edited by Otto Huber, 281 pp., 172 black/white fig. incl. 47 photo., 6 maps,4 pp. insert of 9 color photo. + 124 pollen grain photos. Fondo Editorial Acta Cientifica Venezolana, Caracas, Venezuela. 1986. Paperbound. This book is a very fine study of the beautifully unique cloud forest of el Rancho Grande named in honor of a wonderful botanist, ardent advocate of education and conservation, and an admired personal friend, Dr. Henri Pittier, who showed us much of this fascinating area two score years ago. Well aware of its unique plant, animal, atmosphere and other special features, he was even then advocating its being designated as a national park. There are 10 carefully prepared detailed papers by 10 authors with 4 by the editor who too is an excellent botanist and ecologist - on the geography, physiology, climate, soils and ecological interpretation of vascular plant adaptations in this misted area. The whole text is rich in illustrations and bibliography and is well documented with the careful research results of the other authors. "THE JOHN BURROUGHS REVIEW - Sesquicentennial Issue 1837-1987, Number I ii + 72 pp., 4 black/white photo., John Burroughs Assn., American Museum of Natural History, New York, N. Y. 12561. 1987. $5.00 paperback. The sesquicentennial is not for this journal but honors the birth on April 3, 1837 of this wonderful naturalist-philosopher- teacher-writer whom it was our privilege to know. There are articles about the Wake-Robin Club, the ties with Vassar College across the Hudson River, the journals, the Washington, D.C. years, and his books. Ah, "Wake-robin"! Inasmuch as we do no editing, papers accepted for publication must be submitted in exactly the form that the author wants to have them published. They will then be photographed and printed by photo-offset in exactly the form as submitted except that we will add page numbers and running-heads. Typescripts should be prepared single-spaced on clean white heavy bond smooth and opaque paper. Elite type is probably the most space-economical. Typescript text must not exceed a rectangle 5*/s inches wide (horizontal) by 8° inches high (vertical), not including the running-head and page number. The title of the paper should be typed in all uppercase (capital) letters with 2 blank lines above the title and one beneath; then the name of the author in ordinary upper- and lower-case letters, along with his address (if so desired); followed by 2 blank lines; then the first line of text. It is usually best to leave a blank line between paragraphs. All scientific plant and animal names and group names should be typed either in italic type (if available) or underscored. Any corrections in the text made by the author must be complete and neat as they will be photographed as they are. The finished typescript as submitted by the author will be reduced from the 85s x 55% inch size as submitted to 6% x 4 inches by the printer. It is therefore advisable to place a centimeter or millimeter scale on all text figures and plates included. Use a new heavily inked black typewriter ribbon and be sure to clean the type on the typewriter after each several pages of typing. Cost of publication at present is $12.00 US per page, with no subsequent rebates, but this rate may vary depending on inflation and costs, so it is best to inquire as to current rates. The page charges are due with the typescript and no paper will be published before payment is received in full. Each author will receive gratis a proportionate share of the printed copies remaining after paid subscrip- tions are filled, but if separates (reprints or offprints) are desired, these will be charged extra in accord with the current rate for offprints provided by the printer. The cost of all such separates ordered must also be paid for in advance at the time the typescript is sent. No orders for separates will be accepted later, nor can additions or corrections be accepted. Authors are asked to indicate in light pencil on the reverse side of each page of their typescript the page number so that no mistakes in sequence occur. All manuscripts accepted will be published in the next issue, so that the size of the numbers may vary greatly. A volume will contain 512 pages. The plan insures prompt publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain limitations of — size and proportion. An extra charge will be made for halftones, depending on their size, as fixed by the engraver. Articles dealing with research in all lines of botany and plant ecology, in any reasonable length, biographical sketches, and critical reviews and summaries of literature will be considered for publication.