-- PHYTOLOGIA An international journal to expedite plant systematic, phytogeographical and ecological publication De get peepee genplan enti ne nnn rn Vol. 80 February 1996 No. 2 CONTENTS _, ESPARZA SANDOVAL, S. & Y. HERRERA ARRIETA, Revisién de Bouteloua barbata Lagasca (Poaceae: Eragrostideae)........5....2....2000.6.. 73 _~ TURNER, B.L., A new Species of Psacalium (Asteraceae, Senecioneae) from Oaxaca, WMéxiaky Atonccn cen. ethene Sartor ay ope. Cia eh Dba 92 ~ TURNER, B.L., Two new species of Senecio (Asteraceae) from México. ....95 » TURNER, B.L., Two new cliff-dwelling species of Pinaropappus (Asteraceae, Lactuceae) from Combine ie Miot i tceeces r att oH Reet a od ates : 100 » PATTERSON, T.F., A new species of Conoclinium (Asteraceae, Eupatorieae) rOM PAGKICO, o's posi tno haere coe abt ana ede ester ests Benes each thane 104 ~~ SIQUEIROS. DELGADO, -M.E., New Leguminosae records from ‘A puascalientes; MEXICO: 0 i5c5 55 sits yoo pew Kans adnwbenaiaeaue sis aweieos tok! 108 _»° TURNER, B.L., The genus Liabwm (Asteraceae, Liabeae) in the Dominican Republic and Plait a en Pik codiin sak oe ioded pesee ie dc ucb ems wan pens tad ey dp 115 .- TURNER, B.L., A new species of Gentiana (Gentianaceae) from Nuevo Leon, MIGRNOOS Sid's facies dun sehk faves ne pie ca pues RaWeetis Ih By wee tas Sun's yee eave once pets 118 ~ WORTHINGTON, R.D., Comments on plant species added to the flora of Texas from El Paso County with more additions. ..............6.c.e cece ee 121 » TURNER, B.L., A new species of Ageratina (Asteraceae, Eupatorieae) from Jalisco, Mexican ty vie Re Gnenbert Cyc ear 128 “ TURNER, B-L., Taxonomic study of the yt iret Sag arizonicus-C. sonoranus (Asteraceae, ‘Heliantheae) ODINIIRX WN seg ena Ra eae ee tay teas Ty 133 CHANTARANOTHAL, P., Pouteria ealnccenals (Supolaceas), new to Thailand. HR Cats cect h a vo en Fak ae RUE aA AU a ame a nana ies spare Bees ec 140 Corrections and adqinOnss 2 Fis eici ices cope porate nowk hh opeativ cakeaseoniaioneds Le 142 Phviclogia Memos lO Available oso ises GIS aS eee 144 CE ee ee eee eee ee ee eee eee ee eee ee Published by Michael J. Warnock 185 Westridge Drive’ Huntsville, Texas 77340 USA. PHYTOLOGIA is printed on acid free paper. LIBRARY NOV 6 - 1996 NEW YORK BOTANICAL GARDEN PHYTOLOGIA (ISSN 00319430) is published monthly with two volumes per year by 185. Westridge Drive, Huntsville, TX 77340. Second Class postage paid at Huntsville, TX. © domestic individual | Michael J. Warnock, 1996 by. PHYTOLOGIA. = Annual subscription (12 issues): $40.00. Annual domestic institutional issues): $44.00. Foreign and/or airmail postage . extra. issue and back issues volume 72°to present: 72); $3.00, add $.75 per copy postage and handling US [$1.50 per copy foreign]). issue sales ‘by volume: $17.00 per volume 42-71 (not all available as complete volumes); $21.00 per volume 72-present; add $3.00 per volume postage “US ($6.00 per volume foreign). POSTMASTER: Send address changes to’ Phytologia, _ 185 midis.) subscription (12 Single copy sales: current Drive, Huntsville, TX .77340-8916: $4.00; back issues (previous to volume a Phytologia (February 1996) 80(2):73-91 REVISION DE BOUTELOUA BARBATA LAGASCA (POACEAE: ERAGROSTIDEAE) Sandra Esparza Sandoval Centro Basico, Universidad Aut6noma de Aguascalientes, Avenida Universidad s/n, Aguascalientes, Ags., 20100 MEXICO & Y olanda Herrera Ameta CIIDIR Unidad Durango, Instituto Politécnico Nacional, Apartado Postal 57, Durango, Dgo., C.P. 34000 MEXICO Becana de la COFAA RESUMEN Se revisan los taxa infraespecificos de Bouteloua barbata, se analizan los caracteres anat6micos en la epidermis abaxial de laminas foliares y en secciones transversales de lamina foliar, vaina y tallo de estos taxa, con el fin de observar las variaciones infraespecificas. Se reconsideran los caracteres morfolégicos que delimitan los taxa y se plantea una clave para la separacion de variedades. PALABRAS CLAVE: Bouteloua barbata, Poaceae, anatomia, taxonomia ABSTRACT The infraspecific taxa of Bouteloua barbata are revised; anatomical features from their leaf abaxial epidermis and transverse sections of leaves, sheaths and culms were analyzed in order to observe the infraspecific anatomical vanation. Morphological characters were reconsidered and a key was proposed to separate the vaneties of this species. KEY WORDS: Bouteloua barbata, Poaceae, anatomy, taxonomy 73 74 PHYTOLOGIA February 1996 volume 80(2):73-91 INTRODUCCION El género Bouteloua Lagasca se distribuye solo en el continente Amencano, especialmente en México, donde 34 de las aproximadamente 40 especies reconocidas se distribuyen a todo lo largo del pafs. Cuatro especies se distribuyen en Norte y Sud América y solamente B. megapotamica Spreng. se restringe a Sud América. Bouteloua curtipendula (Michx.) Torr. ha sido introducida a las islas del Hawaii como pasto forrajero por su polimorfismo y agresividad naturales. Ha sido reconocido como uno de los mds importantes géneros de las gramineas, componente de las praderas y planicies del sur de Norte América. No solo debido a que sus especies son elementos significativos de las asociaciones naturales de pastizales, sino que también son las de mayor calidad forrajera en las dreas de pastoreo. Las especies mas ampliamente distribufdas y econédmicamente importantes son B. curtipendula (bandenilla o triguillo), B. gracilis (H.B.K.) Gnffiths (navajita), B. eriopoda (Torr.) Torr. (navajita negra), y B. hirsuta Lag. (navajita peluda). Las principales contribuciones al conocimiento taxonémico del género son los estudios realizados por. Griffiths (1912), Hitchcock (1913, 1935), Gould (1979), Reeder y Reeder (1980), y Beetle y col. (1987). Dentro del género Bouteloua \a especie B. barbata Lag. ha sufrido varios cambios y reconsideraciones para delimitar sus variedades. Se le reconocen tres variedades: B. barbata var. barbata, B. barbata var. rothrockii, y B. barbata var. sonorae, cuya distribucién va desde el S de Colorado y Utah hacia el sur hasta Guerrero en México. Creciendo en suelos arenosos y dreas de disturbio cercanas al nivel del mar hasta aproximadamente 2550 m (en el caso de la variedad tipica). Bouteloua barbata Lagasca, es posiblemente una de las especies vegetales que mas se le ha prestado atencién ya que ha sido descrita por un numero considerable de autores, habiendo recibido diversos nombres genénico-especificos, desde el mds antiguo 3. barbata, en Varied. Ci. 2(4):141. 1805.; a las denominaciones posteriores (como se resefia en Gould, 1979): Actinochloa barbata (Lag.) Roemer & Schultes, Syst. Veg. 2:420. 1817.; Eutriana barbata (Lag.) Kunth, en Rev. Gram. 1:96. 1829.; Chondrosium polystachyum Bentham, en Bot. Voy. Sulph. 56. 1844.,; Chondrosium subscorpioides C. Mueller, en Bot. Zeitung 14:347. 1856.; Bouteloua pumila Buckley, en Proc. Acad. Nat. Sci. Philadelphia 93. 1862.; Erucaria tetrastachya Cerv., en Naturaliza 1:349. 1870.; Chondrosium exile Fournier, en Mex. Pl. Gram. 2:137. 1886.; Chondrosium microstachyum Fournier, en Mex. Pl. Gram. 2:138. 1886.; Bouteloua microstachya (Fourn.) L.H. Dewey, en Contr. U.S. Natl. Herb. 2:531. 1894.; Bouteloua arenosa Vasey, en U.S.D.A. Div. Bot. Bull. 12(1): pl. 34. 1890.; Bouteloua micrantha Scribner & Memll, en U.S.D.A. Div. Agrostol. Circ. 32:8. 1901, Bouteloua barbata Lag. var. rothrockii (Vasey) Gould, en Ann. Missouri Bot. Gard. 66:403. 1979. Fué inicialmente descrita como B. polystachya major Vasey, en Wheeler, Rep. U.S. Surv. 100th Merid. 6:287. 1878.; postenormente reconocida como B. rothrockii Vasey, en Grasses U.S. 33. 1883.; como B. micrantha Scribner & Mermill, en U.S. Dept. Agr. Div. Agrost. Circ. 32:8. 1901.; y mds tarde como B. barbata var. major (Vasey) Beetle, y col., en SARH-COTECOCA II:63. 1987. Esparza Sandoval & Herrera Armeta: Bouteloua barbata 13 Bouteloua barbaia Lag. var. sonorae (Griffiths) Gould, en Ann. Missoun Bot. Gard. 66:403. 1979. Fué promero descrita como Bouteloua sonorae Griffiths, en Contr. U.S. Natl. Herb. 14:389. 1912. Los resultados que aquf se reportan fueron generados como parte del proyecto general “Estudio biosistematico de las gram{fneas de importancia forrajera en México”, que se lleva a cabo en el CIIDIR Unidad Durango, en colaboraci6n con el personal del Herbario de la Universidad Aut6noma de A guascalientes. MATERIALES Y METODOS Se utilizaron ejemplares de Bouteloua barbaia tanto de matenal fresco colectado especialmente para este estudio, como de ejemplares de herbano obtenidos en préstamo. Los ejemplares revisados para complementar el estudio provinieron de los siguientes herbarios mexicanos: ANSM, CIIDIR, ENCB, HUAA, IEB, IBUG, MEXU, SLPM, y UAG; asf como del US, de EUA. La metodologia que se empleo en el estudio morfoldgico es aquella propuesta por Lott y Chiang (1990), que se utliza regularmente en los estudios floristicos, para llevar cabo las descnipciones de los taxa y la clave de separacién de vanedades. Para el estudio anat6mico se utilizaron muestras colectadas en fresco (Cuadro 1), fijadas en Formalina-Ac. acético-Alcohol (FAA), seleccionando el tercio medio de l4minas foliares maduras, los tallos de la parte central cercana al nudo, la vaina que los cubre en la porcién superior en cada ejemplar en fresco. Los cortes fijos en FAA fueron sometidas a deshidratacién en series de etanol siguiendo la metodologia propuesta por Berlyn y Miksche (1976). Se hicieron cortes de 10 a 12 micras de espesor y se tifieron con safranina al 1% durante 45 min, luego con verde de malaquita al 0.2% cubriendo la laminilla con este colorante y lavandola casi inmediatamente. Las muestras se suavizaron con una solucion 0.1 glycerina-etanol- agua antes de ser examinadas con un microscopio de contraste de fases y a un aumento de 40x. Las muestras tomadas de ejemplares de herbano fueron rehidratados en agua caliente y ablandados con KOH al 5%, y los cortes transversales de tallo, vaina y lamina foliar fueron hechos en fresco con una navaja de rasurar, sin tincidn. Las descripciones anat6micas se llevardn a cabo siguiendo el sistema propuesto por Ellis (1976). De forma complementana se llev6 a cabo la descnpcién anatomica de epidermis con base en el método estandarizado de Ellis (1979). DESCRIPCIONES ANATOMICAS Lamina foliar en secci6n transversal. Ldmina en corte transversal generalmente muy abierta, en forma de V, con angulos menores a 30°, brazos rectos y simétnicos, de 0.1 a 0.15 mm de espesor; margenes 76 PHYeT OLOGTA February 1996 volume 80(2):73-91 profundamente ondulados, 1/4 o mds del grosor de la lamina en ambas superficies. Hv primarios (HvI°) y Hv secundanos (HvII°) presentes, dispuestos centralmente en la lamina, subiguales, de forma circular a tnangular-ovados en la var. sonorae, 3 a S{- 7) HvI°. HyvII° intercalados en pares entre los HvI°, (3 HvII° entre cada HvI° en la var. rothrockii). Hvi° con una vaina compuesta por 10 a 11 células Kranz (cK), interrumpidas abaxialmente por una viga de esclerénquima; una vaina interna de mestoma que rodea al haz vascular; células del floema esclerosadas; xilema con 2 vasos metaxilematicos adyacentes al floema y con | a 2 vasos protoxilematicos (1 en la var. rothrockii), de paredes engrosado-angulosas. HyvII° ininterrumpidos, con una vaina de 8 a 10 cK y una vaina interna de mestoma que rodea al xilema y floema. Esclerénquima presente a manera de vigas angostas adaxial y abaxialmente en los HvI° y HvII°, y en los margenes estrechos de proyeccién ligeramente puntiaguda. Células del clorénquima en una sola hilera, rodeando cada Hv, y separadas por | 0 2 hileras de células del parénquima esponjoso. Células buliformes en forma de abanico presentes en la unién adaxial de los Hvs. Epidermis adaxial y abaxial unicelular, adaxialmente con tricomas unicelulares cortos de base no bulbosa, unicuspidales a bicuspidales en la var. sonorae; papilas unicelulares, pequenas, redondeadas a ovadas, infladas. Abaxialmente con micropelos bicelulares y con ganchos cortos en las vars. rothrockii y sonorae (Figura 1). Tallo en seccion transversal. Corte transversal obloide, con bordes ondulados a lisos en la var. sonorae, sin pubescencia 0 con pubescencia en la var. rothrockii. Amillo de esclerénquima interrumpido por haces vasculares secundanos (HvII°) penféricos; HvII° con una vaina de clorénquima radial constnfido en su porcién media por una viga de esclerénquima subepidérmico, seguida de una vaina de 5 células Kranz (cK), semicirculares; 9 a 10 HvII° dispuestos en las crestas, en forma alterna con los HvI° internos. Anillo de esclerénquima multiestratificado, ocupando aproximadamente 2/5 partes de la superficie total. HvI° intenores en un solo nivel del circulo, bordeados en Su parte supenor por el anillo de esclerénquima; adicionalmente la var. rothrockii presenta un anillo de HvI° mucho mds pequefios, alternos a los grandes e inmersos hasta la mitad de su superficie en el anillo de esclerénquima, mientras que la var. sonorae presenta 2 HvI° inmersos en el parénquima central del tallo; HvI° ovalados con 3 vasos metaxilemdticos (2 laterales externos, grandes y i central interno, pequeno). Parénquima continuo hasta el centro, aproximadamente 3/5 partes de la superficie total (Figura 2). Seccién transversal de estolén en Bouteloa barbata Lag. var. sonorae (Griffiths) Gould. Corte transversal obloide con bordes lisos, sin pubescencia. Anillo de parénquima subepidérmico ocupando aproximadamente 1/4 parte de la superficie total, interrumpido por HvII° penféncos. HvII° con una vaina de clorénquima radial Esparza Sandoval & Herrera Ameta: Bouteloua barbata 77 constrifiida en su porcién media por una viga de esclerénquima subepidérmico, en seguida una vaina de 5 cK, colocadas en un semicirculo, rodeando el HvII°; HvII° alternos a los 90 10 HvI° del primer anillo. Anillo de esclerénquima adyacente al parénquima subepidérmico, ocupando 1/4 de la superficie total, interrumpido por los HvI°‘s que estan inmersos casi totalmente en la banda de esclerénquima, ovalados, alternos con los HvI° de un segundo anillo interno, con 2 a 4 vasos metaxilemdticos; HvI° del segundo anillo semejantes a los HvI° del pnmer anillo, inmersos en el parénquima central. Parénquima central ocupa aproximadamente la 1/2 de la superficie total (Figura 2). CUADRO 1. Ejemplares utilizados en las descnipciones anat6émicas Bouteloua barbata Lag. var. barbata Aguascalientes: M. de la Cerda 3225, 3273, 3807, 3829, 4168, 4203 (HUAA); M. Siqueiros 2478 (HUAA); Plan Lerma 1117 (HUAA); J.R. Reeder & C.G. Reeder 1325 (MEXU). Durango: S. Gonzdlez y S. Acevedo 2508 (CIIDIR); Y. Herrera 319 (CIIDIR); Y. Herrera 1347, 1350(CIIDIR-HUAA). Hidalgo: R. Hdz. Magara 6677 (ENCB, MEXU); J.R. Reeder, C.G. Reeder, & L.N. Goodding 1621 (ENCB); /. Diaz, J. Vilchis, y A. Diaz 350 (MEXU). San Luis Potosi: J. Rzedowski 4211; J.L. Jiménez s/n (SLPM); Rodarte s/n (SLPM); A. Gdmez-Glz. s/n; F. Miranda 8918 (MEXU); C.D. Orcutt 5437 (MEXU); Schaffner 1020 (MEXU). Sinaloa: F.W. Gould 12116(ANSM). Sonora: Y. Herrera y M. Siqueiros 1322, 1326, 1328, 1330, 1332 (CIIDIR-HUAA); R.S. Felger 85-949 (IEB); R.S. Felger 14971 (IEB,ENCB, MEXU). Bouteloua barbata Lag. var. rothrockii (Vasey) Gould Sinaloa: Y. Herrera y M. Siqueiros 1313, 1316 (CIIDIR-HUAA); R. McVaugh 1439 (IEB); H.H. LeRoy 8772 (ENCB); H.S. Gentry 14338 (MEXU). Sonora: W.R. Anderson 12547 (MEXU). Bouteloua barbata Lag. var. sonorae (Griffiths) Gould Sinaloa: Y. Herrera y M. Siqueiros 1309 (CIIDIR-HUAA). Vaina foliar en secci6én transversal. Vaina en corte transversal en forma de U, borde abaxial ondulado. Epidermis sin tricomas en la var. rothrockii, con tricomas unicelulares cortos de base no bulbosa, 78 PHYTOLOGIA _ February 1996 volume 80(2):73-91 unicuspidales y papilas unicelulares, pequefias ovadas de pared del gada, intercalados a lo largo de la epidermis abaxial y adaxial en la var. sonorae y solo en la epidermis abaxial en la variedad tfpica. Esclerénquima subepidérmico continuo en el borde abaxial a lo largo de la vaina hasta los mdrgenes, a discontinuo por los haces vasculares (Hv) en la var. rothrockii. Hv dispuestos abaxialmente en un mismo nivel. HvI° y HvII° presentes, intercalados 1:1 en la var. barbata, 1:2-3 en la var. rothrocki, y 1:3 en la var. sonorae, HvI°‘s obloides a semicirculares, con una vaina de clorénquima radial seguida de una vaina de 3 a 4 células Kranz (cK) a cada lado (2 en la var. rothrockii), interrumpidas adaxialmente por parénquima y abaxialmente por una viga de esclerénquima; con una vaina de mestoma interna que rodea el floema y xilema; con 2 vasos metaxilematicos, 1 a cada lado de la porcidn superior (3 vasos en la var. sonorae, el tercero central). HvII° semicirculares a obloides en la var. sonorae; con una vaina de clorénquima radial seguida de una vaina de 2 cK a cada lado en la var. rothrockii, de 3 cK a cada lado en la var. tipica y de 8 cK en la var. sonorae; interrumpidas adaxialmente por parénquima y abaxialmente por una viga de esclerénquima, en las vars. barbata y rothrockii, a ininterrumpidas en la var. sonorae (Figura 3). Epidermis foliar en vista abaxial. Epidermis vista abaxialmente (envés), con zona costal e intercostal presentes. Zona intercostal constituida por bandas estomdticas (Be), bandas de células intercostales largas (Bil), con células intercostales cortas y micropelos (Bilcm), y con ganchos cortos (Bilg). En algunas células intercostales se observan papilas. Arreglo de Ja distnbucién entre los diferentes tipos de bandas, partiendo de la uni6n costal 1(Bilg) -1(Be) -1(Bil) -1(Bilcm) -1(Bil) -1(Be) -1(Bilg). Células intercostales largas 3 veces o mds largas que anchas, rectangulares, paredes moderadamente engrosadas, con ondulaciones profundas en forma de U en las vars. barbata y rothrockii a ligeramente onduladas en la var. sonorae. En Bilg, células intercostales largas con ganchos cortos intercalados irmegularmente entre ellas. En Bilcm, cada célula intercostal larga separada por una célula intercostal corta (1 a 2 en la var. sonorae). En Bilc, células intercostales cortas, igual de altas que células intercostales largas pero muy estrechas, no silificadas. Micropelos bicelulares, emergiendo de la base de las células intercostales cortas, no formando Angulo con la base; célula basal ligeramente mas larga que distal, mds larga que ancha, de base bulbosa; célula distal algo mds corta que basal, inflada con dpice ahusado. Ganchos presentes en las vars. rothrockii y sonorae, de base igual a ligeramente mds corta que los estomas, con la punta encorvada, corta, intercalados de manera discontinua en las células intercostales largas, adyacentes a la zona costal. Papilas redondeadas, circulares (en forma de media luna en la vaniedad tfpica), grandes, las mayores 1/2 de la anchura vertical de las células intercostales largas, infladas, pared engrosada, comunmente elongadas, con el dpice c6ncavo, distnbuidas en al menos del 50% de las células intercostales largas, distalmente, una papila por célula. Estomas elipsoides en forma de domo bajo ovoide; células subsidianas anexas alargadas y algo constrefiidas en su parte media. Células interestomales presentes en bandas estomaticas, intercaladas entre los estomas, rectangulares a romboides en la var. sonorae, 2 a 3 veces mds largas que anchas, de paredes ligeramente engrosadas, con ondulaciones profundas en forma de U a ligeramente onduladas en la var. sonorae, con extremos céncavos. Zona costal Esparza Sandoval & Herrera Armeta: Bouteloua barbata 79 constituida por bandas de células costales largas (Bcl) y bandas de células costales cortas y de siflice (Bccs); células costales largas, tan largas como las intercostales largas pero 3 veces menos anchas que éstas; células costales cortas rectangulares a cuadradas, de paredes sinuosas, alternando | célula costal corta con | célula de sflice (1 a 2 en la var. sonorae); cuerpos de sflice en forma de silla de montar, desiguales en longitud a las células cortas, de paredes lisas, de bordes ligeramente redondeados. Células de corcho raras a escasas en la var. sonorae, presentes en menos del 25% de la superficie (Figura 4). TRATAMIENTO SISTEMATICO Bouteloua barbata Lagasca, Varied. Ci. 2(4):141. 1805. Planta anual o perenne, tallos decumbentes y abiertos 0 erguidos y amacollados, a veces formando estolones hasta de 5O cm de largo, muy vanable en aspecto y tamanio, dependiendo de su habitat y grado de desarrollo. Raices fibrosas por lo regular delicadas. Tallo generalmente ramificado en el segundo nudo. Hoyas persistentes, principalmente basales. Vainas por lo general cortas, con mechones de pelos largos en cada lado del collar. Ligula siempre presente en forma de un anillo de pelos o membrana corta, de 0.5 mm de longitud. Ldminas foliares de 0.5 a 7 (-13) cm de longitud y de 1 a 3 (-4) mm de ancho, acuminadas en el dpice, a menudo escabroso- pilosas a esparcido estrigosas en la superficie adaxial. © Inflorescencia de espigas purpureas, persistentes, cortamente pediceladas, no ramificadas, pectinadas, ligeramente curveadas (navajitas), con (2-) 4 a 8 (-12) espigas por tallo, de 1 a 2.5 an de longitud, de 1 a 5 mm de ancho, con (20-) 25 a 40 (-55) espiguillas por espiga, sobre un raquis plano, glabro a diminuto escabroso, a veces presenta pelos de base papilar. Espiguillas de 2.5 a 4 mm longitud, incluyendo las anstas cortas, por lo general con dos rudimentos armba de la flor perfecta. Glumas glabras, desiguales, ovado-lanceoladas, acuminadas o ligeramente emarginadas y mucronadas; pnmera gluma de | a 1.5 mm de longitud, por lo general hialina; segunda gluma de 2.5 mm de longitud, por lo general purpurea, en ocasiones presenta pelos de base papilar. Raquilla con un mechén de pelos abajo del rudimento aristado. Lema de la flor perfecta de 2.5a3 mm de longitud, densamente pubescente al menos en los margenes, tnlobada, tnanstada, aristas de 0.5 a 3 mm de longitud, vilosa en la cara externa. Pdlea de 2 a 2.8 mm de longitud, biaristada, vilosa en la cara externa. Flores rudimentarias 2, rudimento inferior con |6bulos redondeados, tnaristados, anstas casi tan largas como aquellas de la lema, con un anillo de pelos en el estipite corto, de 1 mm de longitud; rudimento superior pequeno, sin aristas, en forma de abanico. Anteras rojizas a anaranjadas. Candépsis obovada, apiculada en la base, de casi | mm de longitud. Habitat: Matorrales de zonas andas y semidndas, bosque de encino, pastizales y zonas de disturbio, selva baja caducifolia en suelo gravoso y/o arenoso, costas y playas arenosas. Se le encuentra desde altitudes cercanas al nivel del mar hasta los 2550 m (solo la var. barbata). 80 PHY TOLOGIA February 1996 volume 80(2):73-91 6 Gewese: ate ) ome Wen aoe & comens ¢ Goes ote | oom memetemenees 9 ewesns © sae Retomes ipaes prereset me eT Log = betes Anatomie 2 tere the 62 Bovtetous betes aore 0 owners aes Foes Sew © are @ comen ¢ miewe eee | eaten 9 Oaee ees » ete Rees (oats pe bp Me en ee Arete €: wre thy 6o Peventees barton Log ver Sv eced (vessy} Ova Figura 1. EE A EEN APOE 81 Bouteloua barbata Esparza Sandoval & Herrera Armieta: nomen on wot oe Cnn teat a emcees (Very) Const Figura 2. February 1996 volume 80(2):73-91 PHY TOLOGTA eeeeened » Caerenmees 2 Et 8 it & yeney (ase, ED ae Or) NT PTY 2 Um STA SD STREET UE as Or) RE SY 6 apy oe 6p ery Figura 3. Bouteloua barbata Esparza Sandoval & Herrera Ameta: he ee ee ee ee ee ey eet eet fe of ee ee eae ee - Ro eNO, OS, eoetee ke Q FEL OVNOC Q ous ome, Rom cwlo me} “ONO: ao, x Aes. () s ey : i = ; —— <= ——- moan pe — a aie ‘i oe i Ww OE) (.) ® =) a=) on 2 : i Sra = 3 een : © Rete a's ze Cit ot ome PP Py ‘oe qebondaoadagcd pe ik em - c ; Forte | | BREPRRRTEI| alo a 2c f Wie QO} GO : Lupinus aif. leptocarpus Benth. Macroptilum atropurpureus (DC.) Urban. aS 4 Fr. Lotus repens (Don.) Stand]. & Steyerm Medicago lupulina L. 112 PHY TOL@GIA February 1996 volume 80(2): 108-114 TABLE 1. (cont.). en eee eS Se eer eee FERRE [SUBFAMILY PAPILIONOIDEAE (cont) _[|——sSsSsd SSS | Melilotus alba.Dest. 8 | ee Phaseolus grayanus Woot. & Standl. a aes ae ee eee eee [SUBEAMILY CAESALPINOIDEAR 7 | es ee Chamaecrista nictitans (L.) Moench. var. jaliscensis hem arte (Greenm.) Irwin & Barneb 3 le Chamaecrista serpens Greene var. wrightii (A. Gray) Pers Irwin & Barneb 13 le,2a Hoffmannseggia glauca (Ort.) Eifert. Pee tt ..... dle ee Distnbution (municipios of the state of Aguascal Aguascalientes, 5) Asientos, 3. Calvillo, 4. Cosio, 5. Jestis Maria, 6. Pabellén, 7. Rincédn de Romos, 8. San José de Gracia, 9. Tepezala. Vegetation: 1. Scrub: a-thorny, b-subthorny, c-inerme, d-subinerme, e-subtropical; 2. Forest of: a-Quercus, b-Pinus, c-Quercus-Pinus, d-Pinus-Quercus, e-Juniperus, 3. Grassland; 4. Disturbed vegetation; 5. Agniculture; 6. Riparian vegetation. On the other hand, most of the species belonging to the subfamily Mimosoideae which were reported as new records, are characteristic trees from the tropical scrub. This type of vegetation is being decreased day by day because of conversion of the area for agriculture (citns and guava plantations). Acacia acailensis and Calliandra grandiflora are some of the scarce trees, only found in conserved areas of this region. The same situation exists with Conzattia multiflora (subfamily Caesalpinoideae) an uncommon tree, known only from its site collection. Nevertheless, other trees are a dominant part of the landscape in this type of vegetation. Here we can mention Leucaena esculenta, Lysiloma acapulcense, Lysiloma microphyllum, and Albizia plurijuga. Mimosa zygophylla is located only in Tepezala and Asientos, over xero- calcareous soils common in the north of the country and in these municipios of Aguascalientes. In the same region, Senna bauhinioides and Hoffmannseggia glauca (subfamily Caesalpinoideae) were collected. Siqueiros Delgado: Leguminosae in A guascalientes 113 In conclusion, it is possible to mention that Aguascalientes is probably not offering new discoveries to science, given its geographic situation. In fact, few botanic expeditions had occurred specifically on its mountainous region, making some slopes interesting for botanists at present. ACKNOWLEDGMENTS I acknowledge the species determinations of Desmodium and Lupinus by Dr. R. McVaugh; Phaseolus, Macroptilium, and Cologania by A. Delgado; Calliandra, Zapoteca, and Erythrina by Dr. H. Hernandez; Crotalaria by C. Soto; Ramireziella by H. Ochoterena; and Tephrosia by O. Telles. Special thanks are given to Y. Herrera- Amieta for reviewing and translating to English this manuscript. I also thank the facilities received in the consultation of MEXU specimens. REFERENCES Anénimo. 198la. Las Leguminosas del Estado de México. Gobierno del Estado de México. Direccién de Recursos Naturales. México, D.F., México Anoénimo. 1981b. Sinteis Geogrdfica de Aguascalientes. Secretarfa de Programacion y Presupuesto. México, D.F., México 98 pp. Barneby, R.C. 1964. Atlas of North American Astragalus. Vol. 13 Part | and II. Memoirs of the New Botanical Garden. Bronx, New York. Barneby, R.C. 1977. Dalea Imagines. Vol. 27. Memoirs of the New York Botanical Garden. Bronx, New York. Bolio, A.E., J.M. de la Puente, R. Moncallo, & S. Villa 1970. Estudio de la Vegetaci6n Forestal del Estado de Aguascalientes. Inventario Nacional Forestal, México. Correll, D.S. & M.C. Johnston. 1970. Manual of the Vascular Plants of Texas. Texas Research Foundation. Renner, Texas. 1881 pp. de la Cerda, L.M. & M.E. Siqueiros. 1985. Estudio Ecolédgico y Floristico del Estado de Aguascalientes. Programa de Investigaciones Bioldégicas. Centro Basico, U.A.A. Aguascalientes, Ags. México. 69 pp. Garcia, R.G. 1989. Plantas Medicinales de San José de Gracia U.A.A. Aguascalientes, Ags. México. 106 pp. Hernandez, H. 1982. Female sterility in Erythrina montana. Erythnna Simposium IV. Allertonia 3(1). Hernandez, H. 1986. Zapoteca: A new genus of Neotropical Mimosoideae. Ann. Missouri Bot. Gard. 73:755-862. Hernandez, H. 1989. Systematics of Zapoteca (Leguminosae). Ann. Missoun Bot. Gard. 76:78 1-862. Isley, D. 1973. Leguminosae of the United States: I. Subfamily Mimosoideae. Memoirs of the New York Botanical Garden Vol. 23(1): 1-152. Isley, D. 1975. Leguminosae of the United States: II]. Subfamily Caesalpinoideae. Memoirs of the New Y ork Botanical Garden Vol. 2{2):I-VIII, 1-168. 114 PHYTOLOGIA February 1996 volume 80(2): 108-114 Isley, D. 1981. Leguminosae of the United States: III. Subfamily Papilionoideae. Tribes Sophoreae, Podalyrieae, Loteae. Memoirs of the New York Botanical Garden. Vol. 25(3): 1-264. Leén Cazares, JM. 1970. Estudio Ecolédgico de los Pastizales del Estado de Aguascalientes. Tesis Doctoral, UNAM. México, México. Lott, A. & F. Chiang. 1986. Manual de Herbario, Consejo nacional de la Flora de México. la. Edicién. México, D.F., México McVaugh, R. 1987. Flora Novo Galiciana. Vol. 5, Leguminosae. The University of Michigan Press, Ann Arbor, Michigan. 786 pp. Rzedowski, J. 1972. Notas sobre la Flora del NE del Estado de Aguascalientes. Escuela Nacional de Ciencias Bioldégicas, México. 19:31-43. Rzedowski, J. 1979. Flora Fanerogdmica del Valle de México. Vol. I. Editonal Continental. México. México. 403 pp. Rzedowski, J. 1988. Analisis de la distribucién geogrdfica del complejo Prosopis (Leguminosae, Mimosoideae) en Norte Aménca. Acta Botanica de México. 3:7- 19. Standley, P.C. 1920. Trees and Shrubs of Mexico. Contr. U.S. Nat. Herb. 23:1- 1721: Standley, P.C. & J.A. Steyermark. 1946. Flora of Guatemala. Fieldiana: Botany. Vol. 24. Part V. Chicago Natural History Museum. Vines, R.A. 1960. Trees, Shrubs and Woody Vines of the Southwest. University of Texas Press. Austin, Texas. 1033 pp. Phytologia (February 1996) 80(2):115-117. THE GENUS L/ABUM (ASTERACEAE, LIABEAE) IN THE DOMINICAN REPUBLIC AND HAITI B.L. Turner Department of Botany, University of Texas, Austin, Texas 78713 U.S.A. ABSTRACT Liabum in the Dominican Republic and Haiti is treated as having but a single variable species, L. poiteaui (Cass.) Urb. The seven other names of Liabum proposed for this region are treated as synonyms. KEY WORDS: Asteraceae, Liabeae, Liabum, Haiti, Dominican Republic, systematics LIABUM Adanson Perennial herbs or shrublets. Leaves simple, mostly bicolored, opposite, usually connate, often forming a nodal disk; petioles usually winged; blades broadly ovate to elliptical, 3-nervate from or near the base, the margins serrate. Heads campanulate, one to numerous in a rather open or congested capitulescence. Involucres campanulate to broadly turbinate, 5-6 senate, the bracts ovate (outermost series) to lanceolate, graduate, persistent. Receptacle with bristly chaff or strongly alveolate. Ray florets 20-120 or more, usually in 2 or more senes, the ligules linear and yellow. Disk florets 10-80+, the corollas yellow. Achenes small, prismatic, ca. 10-nbbed, hispidulous, the inner pappus of 15-40 white or tawny bnistles. Base chromosome number, x=9. Type: Liabum brownei Cass. [= L. umbellatum (L.) Sch.-Bip.] According to Robinson (1983), a genus of perhaps 37 species, most of these found in South America. A single highly vanable species occurs in the Dominican Republic and Haiti. Robinson (1983) recognized six species as occurring in this region, but all of these appear to be but forms of Liabwm poiteaut. 115 116 PHY TOL @GGTA February 1996 volume 80(2): 115-117 LIABUM POITEAUI (Cass.) Urb., Ark. Bot. 23A:87. 1931. BASIONYM: Andromachia poiteaui Cass., Bull. Soc. Philom. 184. 1817. TYPE: HAITI. “in via ad Barriere Couchant,” Poiteaui s.n. (P, not examined). Liabum domingense Rydb., Fl. N. Amer. 34:290. 1927. TYPE) DOMINICAN REPUBLIC: near Constanza, Santa Domingo, 1400 m, Tuerckheim 3113 (HOLOTYPE. NY'!): Liabum subacaule Rydb., Fl. N. Amer. 34:290. 1927. TYPE: HAITI: Between Petit Borgne and Mount Casse, Nash 502 (HOLOTYPE: NY'!). Liabum barahonense Urban, Arkiv. Bot. 23A:85. 1931. TYPE: DOMINICAN REPUBLIC: prov. Barahona near Paradis, 150 m, Tuerckheim 2785 (Isotype: NY!). Liabum oblanceolatum Urb. & Ekman, Arkiv. Bot. 23A:89. 1931. TYPE: DOMINICAN REPUBLIC: Santo Domingo, Prov. de la Vega, Cordillero Central, “in scopulosis umbrosis Valle Nuevo ad nvulum”, 2400 m, Ekman H. 13827 (HOLOTYPE: S, not examined). Liabum ovatifolium Urb., Arkiv. Bot. 23A:89. 1931. TYPE: DOMINICAN REPUBLIC: Santo Domingo, Prov. Espaillat, near Moca, “in colonia Jamao locos saxosis”, 900 m, Ekman H.12578 (HOLOTYPE: S, not examined). Liabum polycephalum Urb. & Ekman, Arkiv. Bot. 23A:88. 1931. TYPE HAITI: Massif de la Hotte, near Torbec above Lie-Mare-Proux, 900 m, Ekman H. 5346 (Isotype: NY!). Liabum selleanum Urb., Arkiv. Bot. 23A:86. 1931. TYPE: HAITI: “Massif de la Selle in decivibus Mome Cabaio”, 1950 m, Ekman H. 1548 (Isotype: NY'!). After examination of a large suite of specimens of Liabum from Haiti and the Dominican Republic (GBSD,NY ,LL,TEX), I conclude that there is but a single highly variable species on the island concerned, this being Liabwn poiteaui, the earliest available name for the aggregate. Urban, in his recognition of that species (among many others!), noted that Rydberg’s acceptance of L. cubense Rydb. as occurring in Haiti was a misidentification of forms referable to L. poiteaui. | concur with this assessment. Finally, I should note that a broader species concept than that espoused here might show that Liabum umbellatum (L.) Sch.-Bip. will encompass L. poiteaui, but my cursory examination of a wide range of Caribbean material suggests otherwise. This research was occasioned by my early efforts to “work up” the genus Liabwn for the Dominican Republic. This has now been superseded by a more ambitious study over a broader region by others. ACKNOWLEDGMENTS I am grateful to my colleague Ted Delevoryas and Justin Williams for reviewing the paper. Tumer. Liabum on Hispanola 117 REFERENCES Robinson, H. 1983. A generic review of the tribe Liabeae (Asteraceae). Smithsonian Contr. Bot. 54: 1-66. Rydberg, P.A. 1927. Liabeae, in N. Amer. Fl. 34:289-301. Phytologia (February 1996) 80(2):118-120. A NEW SPECIES OF GENTIANA (GENTIANACEAE) FROM NUEVO LEON, MEXICO B.L. Turner Department of Botany, University of Texas, Austin, Texas 78713 U.S.A. ABSTRACT A new species of Gentiana, G. hintoniorum B.L. Turner, from Nuevo Leé6n, México, is described and illustrated. It belongs to the sect. Pneumonanthe and relates to Gentiana laevigata, the latter known only from the Mexican states of Chiapas and Oaxaca. KEY WORDS: Gentianaceae, Gentiana, México, Nuevo Leon, systematics GENTIANA HINTONIORUM B.L. Tumer, spec. nov. Figure 1. TYPE: MEXICO. Nuevo Leén: Mpio. Aramberm, Cerro Viejo, 2435 m, pine-oak forest, 19 Nov 1993, Hinton et al. 23961 (HOLOTYPE: TEX!). Similis G. laevigatae Mart. & Gal. sed tubis calycum fississimis (vice non fissorum) et corollis parvioribus, ca. 2 cm longis (vice 2.5-4.0 cm longis). Perennial herbs ca. 25 cm high. Stems reddish, minutely hispidulous, the vestiture 0.1 mm high or less. Leaves opposite throughout, gradually reduced upwards. Midstem leaves linear-lanceolate to subfalcate, entire, 2-3 cm long, 0.3-0.4 cm wide, uninervate, pubescent like the stem, especially along the margins and midnb. Flowers arranged in terminal spike-like racemes 3-13 cm long, the pedicels 4-12 mm long. Bracts at base of calyx 2, linear-lanceolate, 6-10 mm long, ca. 0.5 mm wide. Calyces 8-12 mm long, tubes 5-7 mm long with a lateral sinus 3-4 mm deep; lobes 5- 6, linear-lanceolate, 2-4 mm long. Corollas blue, ca. 2 cm long; lobes 5, broadly ovate, apiculate, between these, 1-2 appendages 2-3 mm long. Stamens 5, ca. 18 mm long, glabrous; anthers yellow, ca. 2 mm long, basally attached, or nearly so. Capsules (immature) ca. 18 mm long, glabrous; beak ca. 1.5 mm long; stigmas two, ca. 0.8 mm long, sharply recurved. This species belongs to the sect. Pneuwmonanthe of Gentiana and, except for its deeply cleft calyx, would key to or near G. laevigata in Pringle’s (1977) treatment of the Mexican species of this section. The latter, so far as known, occurs only in the Mexican states of Chiapas and Oaxaca. 118 Tumer. New Gentiana from Nuevo Leén Figure 1. Gentiana hintoniorum B.L. Turner, from holotype. 119 120 Piyel) OW OIG IVA February 1996 volume 80(2): 118-120 It is a pleasure to name this apparently localized endemic for the Hinton family, who continue to garner a remarkable array of endemic plants from northeastern México. ACKNOWLEDGMENTS I am grateful to Gayle Tumer for the Latin diagnosis, and to her and Ted Delevoryas for reviewing the paper. Maria Thompson provided the illustration. LITERATURE CITED Pringle, J.S. 1977. Taxonomy and distnbution of Gentiana (Gentianaceae) in México. Sida 7:174-217. Phytologia (February 1996) 80(2):121-127. COMMENTS ON PLANT SPECIES ADDED TO THE FLORA OF TEXAS FROM EL PASO COUNTY WITH MORE ADDITIONS Richard D. Worthington Floristic Inventories of the Southwest Program, P. O. Box 13331, El Paso, Texas 79913 U.S.A. ABSTRACT Piant taxa recently added to the flora of Texas from El Paso County are listed and those for which little was published other than presence are discussed. New additions to the flora of Texas are Malacothrix sonorae Davis & Raven and Spartium junceum L. Problematic locality data for records of Mimulus rubellus A. Gray and Physalis acutifolia (Miers) Sandwith are discussed and new records presented. KEY WORDS: Texas, El Paso County, floristics In the mid-1970’s, an intense flonstic survey was started for the Franklin Mountains, E]) Paso County, Texas, and surrounding areas. In the course of that survey, a number of species were added to the known flora of Texas (Correll & Johnston 1970) and others thought to be a part of the flora were documented for the first ime. Some discoveries have been reported elsewhere and include the following (family acronym and reference given): Telosiphonia brachysiphon (Torr.) Henr. APO Hennckson 1996 Baccharis sarothroides A. Gray CMP Worthington 1990a Filago californica Nutt. CMP Worthington 1990a Prenanthella exigua (A. Gray) Rydb. CMP Worthington 1990b Stylocline micropoides A. Gray CMP Worthington 1990a, 1990b Arabis perennans S. Wats. CRU Rollins 1993a, 1993b Brassica tournefortii Gouan. CRU Lemke & Worthington 1991; Rollins 1993a, 1993b Chorispora tenella (Pallas) DC. CRU Lipscomb 1984 Diplotaxis tenuifolia (L.) DC. CRU Spellenberg, ef al. 1986 Lepidium latifolium CRU Worthington 1990a Sisymbrium orientale L. CRU Rollins 1993a, 1993b 122 PHYTOLOGIA February 1996 —_ volume 80(2):121-127 Streptanthus carinatus Wright subsp. arizonicus (S. Wats.) Kruckeberg, Rodman, & CRU Kruckeberg, Rodman, & Worthington Worthington 1982 Mirabilis commata (Small) Standl. NYC Tumer 1993 Collections of the species added to the Texas flora and of those precisely documented for the first tme have been deposited at UTEP and one or more of the following herbaria: TEX, SRSC, NY, and UCR. Johnston (1990) has noted most of these, mentioning them in his update, and Worthington (1989) has included most of them in his El Paso County checklist. However, details of the nature of the occurrences in Texas have not yet been reported for many of them. In this report the collection localities and other information about the species are presented and other species are reported for the first time. APOCYNACEAE TELOSIPHONIA BRACHYSIPHON (Torr.) Henr. This species was recently found in the Franklin Mountains on a granite rock outcrop in a canyon bottom 1.1 air mi. NE from the top of Anthony’s Nose (near 31° 58’ 10” N, 106° 29’ W), 4800 ft., 19 Jul 1995, Worthington 25068 (Henrickson 1996). This collection represents a significant range extension from extreme southwestern New Mexico. This population is restricted to crevices in an outcrop of granite rock measuring no more than 50 m square. ASTERACEAE HYMENOTRRIX WISLIZENII A. Gray Tumer (1962) cited an 1881 Vasey collection from the county that lacked specific locality data. Correll & Johnston (1970) state “not definately known to occur in Texas .. . to be expected in the E] Paso area.” A significant population occurs on the east side of the Franklin Mountains at 4300 ft. elevation on igneous rock denved soils (generally near 31° 54’ 15” N, 106° 27’ 25” W), 11 Jun 1978, Worthington 2953, 08 Jul 1978, Worthington 306]. Another population was located at the Three Sisters Hills at 4100 ft. elevation on a mixed igneous and limestone alluvial substrate (near 31° 52’ N, 106° 33’ 30” W), 15 Sep 1988, Worthington 174006. Worthington: Flora of El Paso County 123 MALACOTHRIX SONORAE Davis & Raven This species was collected in the Franklin Mountains 1.1 air mi. ENE from the top of North Franklin Mountain (near 31° 54’ 13” N, 106° 28’ 30” W), S000 ft., ina steep-walled granite rock canyon, Worthington 24701. This is a significant range extension to Texas from the nearest known population in the Tres Hermanas Mountains, Luna County, New Mexico (Spellenberg ef al. 1986). BRASSICACEAE ARABIS PERENNANS S. Wats. This species of Arabis is now known from a number of localities on the east side of the Franklin Mountains mostly from igneous rock substrates (granite and rhyolite) or from mixed igneous and limestone alluvium at elevations of 4600-5500 ft. Specific localities are: 1.6 mi. WNW ject. Trans-Mountain Rd. with Gateway North-South (31° 54’ 17" N, 106° 28’ W), 11 Mar 1979, Worthington 3980; 1.1 air mi. NNE from the top of Anthony’s Nose (31° 58’ 26” N, 106° 29’ 28” W), 4 Mar 1979, Worthington 3919; 1.5 air mi. NNE from the top of Anthony’s Nose (31° 58’ 45” N, 106° 29’ 25” W), 4 Mar 1979, Worthington 3920; Mundy’s Spring (31° 55’ 10” N, 106° 29’ 15” W), 10 May 1981, Worthington 7036; 0.1 mi. E of McKelligon Canyon Theatre parking lot, 19 Mar 1978, Worthington 2287. SISYMBRIUM ORIENTALE L. This species has become established on disturbed sites on the west side of El Paso. It has been found in the spring on scraped lots, roadsides, and dumped dirt at 4000- 4100 ft. elev. (El Paso at the comer of Ressler and Escondido, Mar 1980, Worthington s.n.; lot on the NW side of El Paso, 9 May 1985, Worthington 13101, Paso at the Three Sisters Hills, 19 Apr 1992, Worthington 20561). CRASSULACEAE SEDUM COCKERELLII Bnitt. Uhl (1972) reports that the species occurs in the Davis Mountains in Madera Canyon at 5800 ft. and gives a chromosome count of n= 15. One collection made by the author from above Limpia Creek east of Mt. Livermore was sent to the late Robert J. Clausen as a live specimen and was confirmed to be this species. Presumably, the species has been collected many tmes in the Davis Mountains, but not prepared in a 124 PHY TOLOGTA February 1996 volume 80(2): 121-127 way that makes determination certain. One collection from the Franklin Mountains is also this species (0.5 air mi. NE from top of North Franklin Mountain, 6600 ft., N- facing metamorphic rock cliff, 13 Oct 1984, Worthington 12748). FABACEAE LOTUS HUMISTRATUS Greene This winter annual occurs on the east side of the Franklin Mountains on granite derived soils (0.9 air mi. WNW jct. Trans-Mountain Rd. with Gateway North/South (31° 54’ 13” N, 106° 27’ 22” W), 4300 ft. elev., Worthington 4182. SPARTIUM JUNCEUM L. This ornamental shrub, native to Europe, is now widely cultivated in El Paso County and probably elsewhere in Texas. It commonly escapes from cultivation and becomes established on vacant lots, roadsides, and arroyos. One voucher collection is from the 5400 block of North Mesa in El] Paso, where it was found growing on a scraped lot (Worthington 19254, 8 May 1991). PORTULACACEAE TALINUM LONGIPES Woot. & Standl. This species has been collected several times at high elevations (6250-6400 ft.) on North Franklin Mountain (0.7 mi. NW from the top of North Franklin Mtn., 10 Sep 1978, Worthington 3464; 0.5 mi. NE of the top of North Franklin Mtn., 27 Aug 1988, Worthington 17079). SCROPHULARIACEAE MIMULUS RUBELLUS A. Gray Correll & Johnston (1970) include this species in the flora of Texas stating that it occurs in “moist and wet places in extreme west Texas.” The basis for the inclusion appears to be the mentioning of the collectuon made by Thurber (# /35) in the “Hueco Mountains” in the onginal descnption (Torrey 1858). Unfortunately, the data on the Thurber collection do not give a specific locality or even mention the state in which it was collected. It is most likely that Thurber made this collection on igneous substrate in the Texas part of the Hueco Mountains, but the possibility exists that it was Worthington: Flora of El Paso County 125 collected in New Mexico. Two recent collections of this taxon in the Franklin Mountains of El Paso County, document this species as part of the Texas flora. The first collecton was from 0.7 air mi. NNW from the top of North Franklin Mountain on fine talus at 6000 ft., (31° 54’ 45” N, 106° 29’ 50” W), 13 May 1995, Worthington 24760. The second was found 0.7 mi. SE from the top of North Franklin Mountain (near 31° 53’ SO” N, 106° 29’ W), about 5900 ft., also on finer grain igneous talus, Worthington 24829. SOLANACEAE PHYSALIS ACUTIFOLIA (Miers) Sandwith Gray (1875) described Physalis wrightii A. Gray, now considered a synonym of P. acutifolia, from a collection made by Charles Wright (# 1602), allegedly from “prairies along the San Pedro River, southwestern Texas.” Waterfall (1958) in his monograph of Physalis, accepted the data on the label of the type as correct, but found no additional Texas collections, and cited only “recent” collections from California and Anizona. Correll & Johnston (1970) followed Waterfall, noting that the species was known from Texas, but only from the one early collection. Apparently the label on the type collection is in error due to the fact that Wnght collected along two San Pedro Rivers. Early in his trip, Wnght collected along the Devil’s River in Texas, at that tme known as the San Pedro River (Hartmann 1992). Late in his trip, he traveled to the San Pedro River in Anzona, where he actually collected the type of Physalis wrightii S of Benson. Additional evidence to this error is the high Wnght collection number, the later years on the label (1851-1852), and the mention of “prairie” which is absent from the Devil’s River in Texas. The species has been collected twice recently in E] Paso County (Three Sisters Hills near 31° 52’ N, 106° 33’ 30” W, 4100 ft., arroyo of mixed igneous and limestone alluvium, 4 Sep 1988, Worthington 17196, 0.3 mi. W of the Borderland Bridge across the Rio Grande at 31° 53’ 15” N, 106° 36’ W, 3760 ft., 27 Aug 1978, irrigation ditch along field, Worthington 3239). West of E] Paso this species is often an abundant weed in old fields. Nesom (pers. comm.) searched the collection at TEX and found one sheet from Cameron County that appears referable to the species, but is worthy of a closer look (Runyon 4243, 11 Jul 1943). ZY GOPHY LLACEAE ZYGOPHYLLUM FABAGO L. Johnston (1990) references D.M. Porter (pers. comm., 1980) to the occurrence of this species as an adventive weed in E] Paso County. Hatch, ef al. (1990) state its 126 PHY OLOGWA February 1996 volume 80(2): 121-127 occurrence in the Trans-Pecos is doubtful. The species occurs along the Rio Grande levee road 0.7 road miles WNW of the Vinton Road bridge (SW of Anthony) in the extreme western tip of Texas and El Paso County (Worthington 4733, 4 Jul 1979; Worthington 5889, 4 May 1980). ACKNOWLEDGMENTS I wish to express my thanks to the following individuals who either reviewed duplicate collections sent to them or made determinations: R.W. Kiger, G. Nesom, A.M. Powell, R. Rollins, B.L. Turner, and the late R.J. Clausen. LITERATURE CITED Correll, D.S. & M.C. Johnston. 1970. Manual of the Vascular Plants of Texas. Texas Research Foundation, Renner, Texas. Gray, A. 1875. Synopsis of North American species of Physalis. Proc. Amer. Acad. Arts 10:62-68. Hartmann, C.P. 1992. Charles Wright, botanizer of the boundary. Password 37:55- 70, 171-187. ; Hatch, S.L., K.N. Gandhi, & L.E. Brown. 1990. Checklist of the vascular plants of Texas. Texas Agricultural Expenment Station Publ. MP-1655, College Station, Texas. Henrickson, J. 1966. Studies in Macrosiphonia (Apocynaceae): generic recognition of Telosiphonia. Aliso 14(3):179-195. Johnston, M.C. 1990. The Vascular Plants of Texas. A List Up-dating the Manual of the Vascular Plants of Texas. 2nd Ed. Published by the author, Austin, Texas. Kruckeberg, A.R., J.E. Rodman, & R.D. Worthington. 1982. Natural hybndization between Streptanthus arizonicus and S. carinatus (Cruciferae). Syst. Bot. 7(3):291-299. Lemke, D. & R.D. Worthington. 1991. Brassica and Rapistrum (Brassicaceae) in Texas. Southw. Naturalist 36(2): 194-197. Lipscomb, B. 1984. New additions or otherwise noteworthy plants of Texas. Sida 10(4):326-327. Rollins, R.C. 1993a. The Cruciferae of Continental North America. Stanford University Press, Stanford, California. Rollins, R.C. 1993b. Brassicaceae (Cruciferae): mustard family. Jn: Hickman, J.C., (Ed.). The Jepson Manual. Higher Plants of California. University of California Press, Berkeley, California. pp. 392-448. Spellenberg, R, R. Worthington, P. Knight, & R. Fletcher. 1986. Additions to the flora of New Mexico. Sida 11(4):455-470. Torrey, J. 1858. Botany of the Boundary. Jn: Emory, W.H. United States and Mexican Boundary Survey. U.S. Government Printing Office, Washington, D.C. pp. 29-270. Turner, B.L. 1962. Taxonomy of Hymenothrix (Helenieae, Compositae). Bnittonia 14: 101-120. Worthington: Flora of El Paso County 127 Turner, B.L. 1993. The Texas species of Mirabilis (Nyctaginaceae). Phytologia 75:432-451. Uhl, C.H. 1972. Intraspecific vanation in chromosomes of Sedum in the southwestern United States. Rhodora 74:301-320. Waterfall, U.T. 1958. A taxonomic study of the genus Physalis in North America north of Mexico. Rhodora 60: 152-173. Worthington, R.D. 1989. An annotated checklist of the native and naturalized flora of El Paso County, Texas. El Paso Southwest Botanical Miscellany No. 1:1-56. Worthington, R.D. 1990a. Additions to the flora of Texas from El Paso County. Sida 14(1):135-137. Worthington, R.D. 1990b. Effects of El Paso pollutants on the lichen, moss, and winter annual flora on andesite rock formations. Jn: Ganster, P. & H. Walter, eds. Environmental Hazards and Bioresource Management in the United States- Mexico Borderlands. Los Angeles: UCLA Latin Amencan Center Publications, Los Angeles, California. pp. 105-115. Phytologia (February 1996) 80(2):128-132. A NEW SPECIES OF AGERATINA (ASTERACEAE, EUPATORIEAE) FROM JALISCO, MEXICO B.L. Turner Department of Botany, University of Texas, Austin, Texas 78713 U.S.A. ABSTRACT A new species, Ageratina jocotepecana B.L. Tumer, is described and illustrated from Jalisco, México. It belongs to the A. mairetiana complex and is distinguished by its very large broadly deltoid leaves which are subcordate at the base. A map showing the distnbution of the four closely related members of this complex in Jalisco is provided. KEY WORDS: Asteraceae, Eupatorieae, Ageratina, México, systematics Routine identification of Mexican Asteraceae has revealed the following novelty. It belongs to the Ageratina mairetiana (DC.) King & H. Rob. species complex as conceived by Turner (1987) a group characterized by its biseriate pappus, the outer bristles much shorter than the inner. AGERATINA JOCOTEPECANA B.L. Tumer, spec. nov. Figure 1. TYPE: MEXICO. Jalisco: Mpio. Jocotepec, Cerro Viejo, “antes de llegar a la cima - (Bola del Viejo) subiendo por Zapoutan - de Hidalgo.” ca. 2650 m, 5 Mar 1989, M. Chazaro B. et al. 5871 (HOLOTYPE: TEX!). Similis A. lasioneurae (Hook. & Arn.) King & H. Rob. sed foliis late deltatis cum basibus subcordatis, costs infra, dense glandulifens- pubescentibus ubique, et achenibus cum pilis plerumque effusis (vice acheniorum atomiferorum-glanduliferorum). Suffly erect perennial herbs 1.6-2.0 m high. Stems submaculate to tannish, densely glandular-hirsute just below the capitulescence, glabrate with age. Leaves opposite throughout, mostly 10-20 cm long, 6-10 cm wide, not much reduced 128 Tumer: New Ageratina from Jalisco Figure 1. Ageratina jocotepecana, from holotype. 130 PHY FOLOGCTA February 1996 volume 80(2): 128-132 HO 'y OW 10I°26 W 23°25'N 23°25'N + + DUR ZRE SAN Figure 2. Distribution of Ageratina mairetiana complex in Jalisco (excluding A. cerifera): A. mairetiana var. mairetiana (closed circles); A. m. var. elucens (open circles); intermediate to var. mairetiana and var. elucens (half circle); A. lasioneura (open triangles); A. jocotepecana (open squares). Based upon matenal at LL, TEX. Tumer. New Ageratina from Jalisco 131 upwards; petioles 3-6 cm long; blades deltoid to subcordate, 5-7 palmately nervate from or near the base, the margins irregularly dentate, glandular-pilose beneath, especially along the major veins. Heads arranged 5-10 in axillary bracteate clusters, the ultimate peduncles 5-15 mm long. Involucres campanulate; bracts 2-seriate, subequal, 8-12 mm long, glandular-pubescent with short hairs, the apices acute to acuminate. Florets 50-80 per head (estimated); corollas white, 5-6 mm long, glabrous or nearly so; tube ca. 3 mm long; lobes ca. 0.8 mm long, acute, glabrous (rarely 1 or 2 short hairs). Achenes cylindric, ca. 4mm long, moderately hispid throughout; pappus of 30-40 white bristles about 7 mm long, below these an outer series of fewer, shorter, bristles 1-3 mm long. ADDITIONAL SPECIMENS EXAMINED: MEXICO. _ Jalisco: Mpio. Jocotepec, “Vereda por la cresta a la Bola del Viejo”, oak woodlands, 2100-2850 m, 5 Feb 1987, Machuca N. 5449 (WISC); Mpio. Tecolotlan, Sierra de Quila, Cerro Huehuenton, ladera SE, ca. 2300 m, 4 Feb 1990, Guerrero N. 654 (TEX). The very broad deltoid blades, subcordate at the base, having lower surfaces with midveins densely glandular-pubescent, readily distinguishes this taxon from Ageratina lasioneura. In addition, it can be distinguished from the latter by the densely glandular-pubescent branches of its capitulescence (vs. eglandular and tomentose). Nearly similar leaf forms to those of A. jocotepecana occur elsewhere in Jalisco (e.g., Mpio. Venustiana Carranza, Villa C. 668 [TEX]), but otherwise such plants have all of the characters of A. lasioneura. It should also be noted that both A. jocotepecana and A. /asioneura occur on Cerro Huehuenton, flower at about the same time (e.g., Guerrero N. 656 [TEX]), but intergrades between them have not been detected. Two of the above cited specimens (TEX), including the type, were included in my broad concept of Ageratina lasioneura (Tumer 1987). I stll maintain Eupatorium chapalense S. Wats. as synonymous with A. Jasioneura, although the former approaches A. jocotepecana in having broadly ovate leaves and pubescent achenes. It differs from A. jocotepecana, however, in possessing eglandular pubescence and leaves with blades merely obtuse or rounded at the base. McVaugh (1984) suggested that such plants might prove distinct; if recognized, the A. mairetiana complex (excluding A. cerifera [McVaugh] H. Robins.) in Jalisco would include at least four named taxa (A. mairetiana var. mairetiana, A. mairetiana var. elucens, A. lasioneura, and A. jocotepecana) as shown in Figure 2. ACKNOWLEDGMENTS I am grateful to Gayle Tumer for the Latin diagnosis, and to her and Ted Delevoryas for reviewing the manuscript. Maria Thompson provided the illustration. 132 IP \St ae Gi ONL, OMG ALIN February 1996 volume 80(2): 128-132 LITERATURE CITED McVaugh, R. 1984. Eupatorium, in Flora Novo-Galiciana 12:240-424. University of Michigan Press, Ann Arbor, Michigan. Tumer, B.L. 1987. Study of the Ageratina mairetiana complex (Asteraceae- Eupatorieae). Phytologia 63:417-427. Phytologia (February 1996) 80(2):133-139. TAXONOMIC STUDY OF THE COREOCARPUS ARIZONICUS- C. SONORANUS (ASTERACEAE, HELIANTHEAE) COMPLEX B.L. Turner Department of Botany, University of Texas, Austin, Texas 78713 U.S.A. ABSTRACT Coreocarpus arizonicus and Coreocarpus sonoranus are closely related, relatively common elements of the Sonoran Desert of northwestern México. Both are quite variable and relatively few characters serve to distinguish between them. Study of a wide range of specimens from the area concerned suggests that within C. arizonicus four intergrading morphogeographical infraspecific elements can be identified: var. arizonicus, var. pubescens, var. filiformis, and var. sanpedroensis (E.B. Smith) B.L. Turner, comb. & stat. nov. Within C. sonoranus two infraspecific taxa are recognized: var. sonoranus and var. ibranus B.L. Turner, var. nov. The latter is known only from Sierra Libre, Sonora. A key to the taxa concerned is provided, along with distribution maps, and comments upon their regional vanability. KEY WORDS: Asteraceae, Heliantheae, Coreocarpus, México, Anzona, systematics Coreocarpus is a relatively small genus largely confined to the Sonoran Desert regions of northwestern México. Smith (1989) recognized Coreocarpus as having nine species, but two of these were subsequently transferred to the genus Bidens by Melchert (Melchert & Turner 1990); even with these two removals the genus appears paraphyletic, for at least two additional species retained in Coreocarpus, C. congregatus (S.F. Blake) E.B. Smith and C. insularis (Brandegee) E.B. Smith, would appear to have their relationships elsewhere. At least one would place in Coreocarpus only four species: C. dissectus (Benth.) S.F. Blake, C. parthenioides Benth., C. arizonicus, and C. sonoranus. All of these have monomorphic involucral bracts, excepi for an occasional 1 or 2 much reduced bractlets at the base of the involucre. This has been amply discussed by Smith. Those taxa removed from the genus by Melchert & Turner (1990) have a double involucre, as in Coreopsis or Bidens. Turner (1991) described an additional species of Coreocarpus from near Mexico City, (C. ixtapanus B.L. Turner) which he took to be a sister species of C. congregatus, noung 133 134 PHYTOLOGIA February 1996 volume 80(2): 133-139 Figure 1. Distribution of Coreocarpus arizonicus - C. sonoranus complex: C. arizonicus var. arizonicus (open circles), C. a. var. pubescens (closed tnangles); Cra: var. filifolius (open triangles); C. a. var. sanpedroensis (closed square); C. sonoranus var. sonoranus (closed circles); C. Ss. var. libranus (open square). Turmer. Coreocarpus arizonicus/sonoranus complex 135 that both of these were questionably placed in Coreocarpus. Smith (1991) promptly sunk C. ixtapanus into synonymy with his enlarged concept of C. insularis. But the question remains: do the taxa concerned belong to Coreocarpus, phyletically speaking? The impetus for the present paper has been occasioned by difficulues with two species of Coreocarpus, C. arizonicus and C. sonoranus. The two are largely distinguished by a syndrome of characters, but best identified by leaf shape, as noted below, and even the latter is subject to considerable variation upon occasion. This has been compounded by much infraspecific variation in both taxa, so much so that I refer to these two taxa in this paper as the C. arizonicus - C. sonoranus complex. The following key will distinguish among members of the Coreocarpus arizonicus - C. sonoranus complex, as recognized here. 1. Leaves with their ulumate divisions variously ovate to deltoid, not at all linear 2. Foliage glabrous or nearly so; lateral divisions of the leaf with sinuses extending to the mid-lines or nearly so; widespread but not in Sierra Libre...... Hera ois's detest dens vetoes ocdaue tence seaouascceeeucereuiessubonee 2a. var. sonoranus 2. Foliage moderately to densely pubescent throughout; lateral divisions of the leaf with sinuses not extending to mid-lines; Sierra Libre, Sonora (ca. 78 km N of RRUEAWINIAS) sich sioves cp eon tsi couuiigerbe sdsmehaunae assess gtcewe sage same 2b. var. libranus 1. Leaves with their ultmate divisions mostly linear to linear-lanceolate; rays mostly WeUow. Occasionally ‘WHItE...00bioee. cen esgeeaecantonssascnereacesceees 1. C. arizonicus 3. Involucres mostly 3-5 mm high; achenes with reduced corky marginal enations; Iptansam PEGIOy, Sins ur. sites. edhe abn. ion ooh ste eee om ol ld. var. sanpedroensis 3. Involucres mostly 5-6 mm high; achenes with well-developed corky wings and/or marginal enations; not on Isla San Pedro. 4. Foliage moderately and evenly pubescent throughout; pappus bnistles MHOSTV PIOSEMl ai acer. ve teseat sence otancateeteee sce cet lb. var. pubescens 4. Foliage glabrous or nearly so; pappus bristles mostly absent, rarely present. 5. Ultimate divisions of leaves relatively short, mostly 2-3(-4) cm long; southern Anzona and closely adjacent Sonora, MEXxico. .................. Be Sn re oO LOAN we acled cede apnerietta sowed awash cl taeda ya la. var. arizonica 5. Ultimate divisions of leaves relatively long and slender, mostly 3-6 cm long; southern Sonora, México and closely adjacent Sinaloa. ............ PEO Re sede oles de Coke Whales SNORE Sones LSS eae haan see else lc. var. filiformis 1. COREOCARPUS ARIZONICUS (A. Gray) S.F. Blake, Proc. Amer. Acad. Arts 49:344. 1913. As noted in the above key four varieties are recognized within this taxon, as follows: 136 PHY TOLGGIA February 1996 volume 80({2): 133-139 la. COREOCARPUS ARIZONICUS (A. Gray) S.F. Blake var. ARIZONICUS My concept of this taxon is essentially the same as that rendered by Smith (1989), with the exclusion of Coreocarpus arizonicus var. filiformis and C. arizonicus var. macrophyllus, which I would include in my concept of C. arizonicus var. filiformis, the latter distinguished by its mostly larger leaves with longer, more linear, divisions, as suggested by its epithet. Collections of var. arizonicus are nearly always glabrous or nearly so in the U.S.A., but in northeastern Sonora var. arizonicus grades into var. pubescens. Indeed, a case could readily be made for the recognition of but a single widespread highly vanable species without infraspecific categones, but this would deny the morphogeographical patterns portrayed in Figure 1. lb. COREOCARPUS ARIZONICUS (A. Gray) S.F. Blake var. PUBESCENS (B.L. Rob.) Fern., Proc. Amer. Acad. Arts 49:344, 1913. My concept of this taxon is about the same as that of Smith (1989). It is only weakly differentiated from var. arizonicus, intermediates between these not uncommon in areas of contiguity, as noted in the above. lc. COREOCARPUS ARIZONICUS (A. Gray) S.F. Blake var. FILIFORMIS (A. Gray) S.F. Blake, Proc. Amer. Acad. Arts 49:344. 1913. Leptosyne arizonicus var. filiformis Greenm. (1904). TYPE: MEXICO. Sinaloa: 80 km NE Choix (LECTOTY PE [selected by Smith 1989]). Coreocarpus arizonicus (A. Gray) S.F. Blake var. macrophyllus Sherff (1935). TYPE: MEXICO. Chihuahua: “southwestern Chihuahua”. Smith (1989) included this taxon in his broad concept of Coreocarpus arizonicus var. arizonicus, but I think it to be as distinctive, if not more so, than var. pubescens, which he maintained. It is seemingly equally close to C. sanpedroensis, hence my reduction of the latter, as follows: ld. COREOCARPUS' ARIZONICUS (A. Gray) S.F. Blake var. SANPEDROENSIS (E.B. Smith) B.L. Turner, comb. & stat. nov. BASIONYM: Coreocarpus sanpedroensis E.B. Smith, Amer. J. Bot. 72:262. 1985. This is a very weakly differentiated infraspecific element of Coreocarpus arizonicus and were it not confined to the Isla de San Pedro, likely not to have been recognized. Smith (1989) distinguished it from C. arizonicus by its achenes which are said to have margins which are corky and involute, dissected into separate teeth, albeit “highly reduced to nearly obsolete in C. sanpedroensis.” My examination of numerous achenes of var. arizonicus and var. sanpedroensis suggest that the differences between the two are mostly quantitative: var. sanpedroensis, in general, Turner: Coreocarpus arizonicus/sonoranus complex inv has somewhat smaller heads with somewhat smaller achenes having somewhat less well-developed marginal enations, otherwise they are scarcely different. 2. COREOCARPUS SONORANUS Sherff, Bot. Gaz. (Crawfordsville) 97:604. 1936. Two infraspecific categories are reorganized under this species, as follows: 1. Foliage glabrous; sinuses of leaves extending to or near their mid-lines; pappus awns mostly absent, sometimes present; widespread in western Sonora............. NE Pe oA Ne Rae A ccies ve ose sence vevennedtcr ciate ns nates rigs be 2a. var. sonoranus 1. Foliage markedly pubescent; sinus of leaves not extending to their midlines; pappus awns mostly present; Sierra Libre, Sonora................2.....20055 2b. var. libranus 2a. COREOCARPUS SONORANUS Sherff var. SONORANUS Coreocarpus johnstonii Sherff (1936) Coreocarpus shrevei Sherff var. latilobus Sherff (1935) Except for the inclusion of maternal referable to my newly described var. libanus (cf. below), this taxon is aptly described by Smith (1989). He notes that Coreocarpus johnstonii and C. sonoranus were described at the same time, but he selected the latter as the most desirable name since he considered C. johnstonii to be typified by an ecotypic (coastal) vanant. As can be noted in Figure 1, Coreocarpus sonoranus is relatively widespread but, so far as known does not co-occur with C. arvizonicus. Nevertheless, it is quite variable, both as regards ray color (mostly white, sometimes yellow), and pappus bristles (mostly absent, sometimes present). It is most readily distinguished from C. arizonicus by its less linear-dissected leaves, as indicated in my key. Nevertheless, occasional plants with young or immature secondary leaflets may superficially resemble C. arizonicus, and it is possible that the two taxa have exchanged genes in the distant past, this perhaps accounting for the occurrence of white and yellow rays in both taxa. 2b. COREOCARPUS SONORANUS Sherff var. LIBRANUS B.L. Turner, var. nov. TYPE: MEXICO. Sonora: Mpio. Hermansillo, Sierra Libre, Canyon E of Restaurante La Pintata, Cerro Bola, ca. 600 m, 3 Jan 1984, R.K. & Tom Van Devender 84-36 (HOLOTYPE: TEX!; Isotype: ARIZ!). Differt a C. sonorano Sherff habendo folia minus dissecta, moderate pubescentia, cum sinibus vadosioribus, et achenia plerumque papposa. Suffruticose perennial herbs 30-50 cm high. Leaves broadly ovate in outline, mostly tripartite-dissected, moderately pubescent, those at midstem mostly 3-6 cm long, 3-4 cm wide; petioles 1.0-2.5 cm long; sinuses of the blade not exiending to mid-lines. Heads 1-4, mostly terminal, the ultimate peduncles 1-3 cm long. Involucres pubescent, 5-6 mm high. Ray florets ca. 5; ligules white, 4-8 mm long. 138 PHY MOLOIGIVA February 1996 volume 80(2): 133-139 Disk florets 20-40, the corollas yellow. Achenes mostly 3.5-4.0 mm long, the margins beset with 6-9 corky enations; pappus of 2 persistent awns 1-2 mm long, sometimes absent. ADDITIONAL SPECIMENS EXAMINED: MEXICO. Sonora (all from Sierra Libre): Mpio. Hermansillo, Cerro Bola, 600 m, 18 Apr 1993, Burquez 93-030 (LL); near small reservoir in Cafiada Las Chivas, 0.1 km SE of cave with paintings, slopes of volcanic rock, 320 m, 16 Nov 1984, Burgess 6723 (ARIZ); La Pintada Canyon, 9 Nov 1986, Smith 3972 (TEX); La Pintada Canyon, 7 and 9 Feb, 1978, Van Devender s.n. (ARIZ); same locality 1 Jan 1982, Van Devender s.n. (ARIZ); same locality, 1 Jan 1983, Van Devender s.n. (ARIZ). This novelty was called to my attention (over protest!) by Dr. Tom Van Devender, who perceived its distinction from the more typical elements of Coreocarpus sonoranus. Smith (by annotation, 1986, and publication, 1984) identified most of the specimens cited above as belonging to the latter, but did note on annotations of both the holotype and isotype that the plants were “near Coreocarpus sonoranus Sherff, but achenes aristate & plant very pubescent!!” Indeed, C. sonoranus var. libranus, in my opinion, is as distinct, if not more so, from var. sonoranus as is C. arizonicus vat. sanpedroensis from C. arizonicus var. arizonicus, hence an additional reason for my reduction of C. sanpedroensis to varietal status here. Var. libranus receives its name from the Sierra Libre, to which it is apparently endemic. So far as known, var. sonoranus has not been collected in this massif. Yetman & Buirquez (1996) have presented an interesting account of Sierra Libre (100- 1100 m elevation). ACKNOWLEDGMENTS I am grateful to Tom Van Devender of the Desert Museum, Tucson, Anzona for his taxonomic proddings which prompted this study, and to the following herbaria for the loan of specimens: ARIZ, ASU, LL, TEX. These acquisitions were all annotated and served to construct Figure 1. I am grateful to Gayle Turmer for the Latin diagnosis, and to her and Ted Delevoryas for reviewing the paper. LITERATURE CITED Melchert, T. & B.L. Turner. 1990. New species names and combinations in Mexican Bidens (Asteraceae - Coreopsidinae). Phytologia 69:20-25. Smith, E.B. 1989. A biosystematic study and revision of the genus Coreocarpus (Compositae). Syst. Bot. 14:448-472. Smith, E.B. 1991. Coreocarpus ixtapanus B. Tumer (Asteraceae - Heliantheae - Coreopsidinae) a synonym of C. congregatus (S.F. Blake) E.B. Smith. Phytologia 71:29-30. Turmer: Coreocarpus arizonicus/sonoranus complex 139 Turner, B.L. 1991. A new species of Coreocarpus (Asteraceae - Coreopsideae) from México State, México. Phytologia 70:42-43. Yetman, D.A. & A. Burquez. 1996. A tale of two species: speculation on the introduction of Pachycereus pringlei in the Sierra Libre, Sonora, Mexico by Homo sapiens. Desert Plants 1966:23-32. Phytologia (February 1996) 80(2):140-141. POUTERIA MALACCENSIS (SAPOTACEAE), NEW TO THAILAND Pranom Chantaranothai Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, THAILAND ABSTRACT Pouteria malaccensis (C.B. Clarke) Baehni, previously known from Malaysia and Sumatra through to New Guinea, is reported from southern Thailand. KEY WORDS: Sapotaceae, Pouteria, Thailand My study of the Sapotaceae for the Flora of Thailand has revealed the occurrence of Pouteria malaccensis (C.B. Clarke) Baehni in Thailand, where it has not hitherto been reported. Both of the collections were made in Klong Na Ka Wildlife Sanctuary, Ranong province, as follows: C. Niyomdham 1244, 5 Nov. 1986 (E,L) and C. Niyomdham, R. Kubat, & W. Aajchomphoo 1441, 18 Mar. 1987 (BKF,E,L). Pouteria malaccensis has been collected in relative abundance in Peninsular Malaysia, Sumatra, Riau, Lingga, Sulawesi, and New Guinea (Herrmann-Erlee & Royen 1957; Ng 1972). The nearest known collections to the Ranong locality have been made about 600 kilometers to the south in Pulau Penang, Peninsular Malaysia. ACKNOWLEDGMENTS I thank Drs. D. Simpson (K) and M. Newman (E) for their reviews and comments. I also thank the curators of the following herbana for loans of their material: BKF, E, and L. Thanks to P. Beedle for providing the publication costs. 140 Chantaranothai: Pouteria malaccensis, new to Thailand 141 LITERATURE CITED Herrmann-Erlee, M.P.M. & P. van Royen. 1957. Revision of the Sapotaceae of the Malaysian area in a wider sense 9. Pouteria Aublet. Blumea 8(2):452-509. Ng, F.S.P. 1972. Sapotaceae. Jn: Whitmore, Tree Flora of Malaya. 1:388-439. Phytologia (February 1996) 80(2): 142-143. CORRECTIONS AND ADDITIONS Volume 78, issue 3, page 297, line 31, substitute herbacea for herbaceous. Volume 78, issue 5, page 358, line 24, substitute Careya for Carea. Volume 78, issue 5, page 358, line 28, substitute azedarach for azadirach. Volume 78, issue 5, page 358, line 30, substitute Acacia nilotica (L.) Willd. ex Del. for Acacia arabica Willd. Volume 78, issue 5, page 358, lines 31 and 32, substitute Sammania saman (Jacq.) Merr. for Enterolobium saman Prain. Volume 78, issue 5, page 358, line 33, substitute Taub. for Tanb. Volume 78, issue 5, page 359, line 1, substitute benghalensis for bengalensis. Volume 78, issue 5, page 359, line 3, substitute globulus for globulus. Volume 78, issue 5, page 359, line 6, substitute Ziziphus trinervia for Zyzyphus trinervi. Volume 78, issue 5, page 359, line 15, substitute disperma for disperina. Volume 78, issue 5, page 359, add to Acknowledgments: We are gratefully acknowledged to the Department of Science & Technology, Govt. of India, New Delhi, under the scheme of promotion of scientific interest in youth for financial assistance. We also express thanks to the Botanical Survey of India, Calcutta for the authentication of species. Volume 79, issue 1, page 13, first sentence under Hedyotis butterwickiae, should read “This species was first described by Terrell in 1979.” Volume 79, issue 1, page 16, the author provided additional information for this map after the paper had gone to press. The corrected map is shown on the page following this one. Note the hatching representing the range of Hedyotis nigricans var. austrotexana. Volume 79, issue 2, page 126, line 26, substitute Triodanis for Trifolium. Volume 79, issue 2, page 127, line 10, substitute Zornia for Zornea. Volume 79, issue 2, page 130, line 19, substitute Zornia for Zornea. 142 143 Corrections and additions TT : SET veneneis ue OX anaes amore. =a eS eos aqme seeecaten @ TBE TET ORR LNG iggse ree Rots SOU PT were AC a etre SSS beth aS as gles); var. papillacea (open Distribution of Hedyotis nigricans in U.S.A.: var. nigricans (closed circles), var. austrotexana (diagonals); var. floridana (closed trian circles); var. pulvinata (open tnanglcs). Figure 1. NOW AVAILABLE, PHYTOLOGIA MEMOIRS, Volume 10 THE COMPS OF MEXICO, Volume 6 TAGETEAE AND ANTHEMIDEAE B.L. TURNER 1996, 8 1/4 x 10 3/4 in., 97 pp., $29.95, ISBN 0-9642085- 1-2. The first of an anticipated ten volume series on the Asteraceae of México, this volume includes the tribes Tageteae and Anthemideae. Later volumes will include the remainder of the 2700+ species of Asteraceae known from México. The Mexican representatives of the family are particularly rich in the tribes Helenieae, Heliantheae, and Eupatorieae. 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