PHYTOLOGIA An international journal to expedite plant systematic, phytogeographical and ecological publication Vol. 87 December 2005 No. 3 D. S. SEIGLER and J. E. EBINGER, New combinations in the genus Vachellia (Fabaceae: Mimosoideae) from the New World......................139 M.H. MACROBERTS and B.R. MACROBERTS, Status and management of Eriocaulon koernickianum (Eriocaulaceae) in Texas ...............0.e.e eee es 179 B. L. TURNER, A Recension of The Mexican species of Roldana MM HCC INOCDE ire 20 8. n i cee wbvtian nS ude Velde enuinape te vppnb dwn «aae 204 _§S. MAHMOOD and M. ATHAR, Soil seed banks and dispersal patterns of Sea Rush, Juncus maritimus in spatially and temporally disturbed coastal EM Tea eats id, SA ede s cids Seid 5c enn cae ied aul e's ve iinn «smb onl s dee ny eden 250 B. L. TURNER, Stevia strotherana (Asteraceae), a newly proposed name for MRTUMEVITE, SPC VICA FISDIGUI A. 0. ois a4 so es aes ep ds neice detnveagevs enervadeesus sed 261 New Names and New combinations in volume 87(3)................20.eeee eee 262 Subscription Information & Information for Authors................-....s0+ 264 List of article reviewers for Phytologia volume 87.............-.:e:0eeecee eee 265 Publication dates for issues of Phytologia volume 87.....................00: 266 PHYTOLOGIA (ISSN 00319430) Phytologia, a journal of plant systematics, phytogeography and vegetation ecology, is published three times a year. Editor Justin K. Williams Department of Biological Sciences Sam Houston State University Huntsville, Tx 77341-2012 PHONE: (936) 294-1552 FAX (936) 294-3940 e-mail: bio_jkw@shsu.edu Associate Editor Billie L. Turner Plant Resources Center University of Texas Austin, Tx 78712-0471 Assistant Editor Steven D. Koether Department of Biological Sciences Sam Houston State University Huntsville, Tx 77341-2012 Subscription Phytologia Information & P.O. Box 2012 Scholarly Exchange Sam Houston State University Huntsville, TX 77341-2012 Copyright 2006 Phytologia. Printed by the Sam Houston Press. Publication date January 4, 2006. MISSOURI BOTANICAL FEB 16 2006 Phytologia (Dec 2005) 87(3) 139 GARDEN LIBRARY NEW COMBINATIONS IN THE GENUS VACHELLIA (FABACEAE: MIMOSOIDEAE) FROM THE NEW WORLD. David S. Seigler’ and John E. Ebinger’ ‘Department of Plant Biology, University of Illinois, Urbana, Illinois 61801, U.S.A. E-mail: seigler@life.uiuc.edu "Emeritus Professor of Botany, Eastern Illinois University, Charleston, Illinois,61920, U.S.A. E-mail: jebinger@eiu.edu ABSTRACT Based on the evidence that the genus Acacia s.l. is polyphyletic, morphological and genetic differences separating the subgenera of Acacia s.l., along with the change to a new conserved type for the genus necessitate the following new combinations: Vachellia acuifera (Benth.) Seigler & Ebinger, V. albicorticata (Burkart) Seigler & Ebinger, V. allenii (D. H. Janzen) Seigler & Ebinger, V. anagadensis (Britton) Seigler & Ebinger, V. aroma (Gillies ex Hook. & Arn.) Seigler & Ebinger, V. barahonensis (Urb. & Ekman) Seigler & Ebinger, V. belairioides (Urb.) Seigler & Ebinger, V. biaciculata (S. Watson) Seigler & Ebinger, V. bilimekii (J. Macbr.) Seigler & Ebinger, V. brandegeana (1. M. Johnst.) Seigler & Ebinger, V. bucheri (Marie- Victorin) Seigler & Ebinger, V. californica (Brandegee) Seigler & Ebinger, V. campechiana (Mill.) Seigler & Ebinger, V. campechiana forma houghii (Britton & Rose) Seigler & Ebinger, V. caurina (Barneby & Zanoni) Seigler & Ebinger, V. caven (Molina) Seigler & Ebinger, V. X cedilloi (Rico Arce) Seigler & Ebinger, V. chiapensis (Saff.) Seigler & Ebinger, V. choriophylla (Benth.) Seigler & Ebinger, V. collinsii (Saff.) Seigler & Ebinger, V. constricta (Benth.) Seigler & Ebinger, V. cookii (Saff.) Seigler & Ebinger, V. cornigera (L.) Seigler & Ebinger, V. cucuyo (Barneby & Zanoni) Seigler & Ebinger, V. curvifructa (Burkart) Seigler & Ebinger, V. daemon (Ekman & Urb.) Seigler & Ebinger, V. gentlei (Standley) Seigler & Ebinger, V. X gladiata (Saff.) Seigler & Ebinger, V. glandulifera (S. Watson) Seigler & Ebinger, V. globulifera (Saff.) Seigler & Ebinger, V. hindsii (Benth.) 140 Phytologia (Dec 2005) 87(3) Seigler & Ebinger, V. insulae-iacobi (L. Riley) Seigler & Ebinger, V. janzenii (Ebinger & Seigler) Seigler & Ebinger, V. macracantha (Humb. & Bonpl. ex Willd.) Seigler & Ebinger, V. mayana (Lundell) Seigler & Ebinger, V. melanoceras (Beurl.) Seigler & Ebinger, V. oviedoensis (R. Garcia & M. Mejia) Seigler & Ebinger, V. pacensis (Rudd & Carter) Seigler & Ebinger, V. pennatula (Schltdl. & Cham.) Seigler & Ebinger, V. polypyrigenes (Greenm.) Seigler & Ebinger, V. pringlei (Rose) Seigler & Ebinger, V. rigidula (Benth.) Seigler & Ebinger, V. roigi (Ledn) Seigler & Ebinger, V. rorudiana (Christophersen) Seigler & Ebinger, V. ruddiae (D. H. Janzen) Seigler & Ebinger, V. schaffneri (S. Watson) Seigler & Ebinger, V. schottii (Torr.) Seigler & Ebinger, V. sphaerocephala (Schltdl. & Cham.) Seigler & Ebinger, V. X standleyi (Saff.) Seigler & Ebinger, V. tortuosa (L.) Seigler & Ebinger, V. vernicosa (Britton & Rose) Seigler & Ebinger, V. villaregalis (McVaugh) Seigler & Ebinger, and V. zapaensis (Urb. & Ekman) Seigler & Ebinger. The following new combinations and new statuses are also necessary: V. aroma var. huarango (Ruiz & J. Macbr.) Seigler & Ebinger, V. bravoensis (Isely) Seigler & Ebinger, V. farnesiana var. minuta (M. E. Jones) Seigler & Ebinger, V. farnesiana var. pinetorum (F. J. Herm.) Seigler & Ebinger, V. guanacastensis (Clarke, Seigler, & Ebinger) Seigler & Ebinger, and V. pennatula var. parvicephala (Seigler & Ebinger) Seigler & Ebinger. Two other species that we recognize had previously been transferred to the genus by others: Vachellia astringens (Gillies in Hook. et Arn.) Speg. and V. farnesiana (L.) Wight & Arn. KEY WORDS Acacia sensu lato, Fabaceae, Mimosoideae, Vachellia. Recent morphological and genetic studies have shown that the genus Acacia s.|. is polyphyletic. Also, there has been an accumulation of data derived from molecular studies that has led to a better understanding of the probable relationships within the genus Acacia s.l., as well as the position of the genus within the Mimosoideae (Maslin 1988, Chappill and Maslin 1995, Clarke et al. 2000, Maslin et al. 2000, 2003, Miller and Bayer 2000, 2001, 2003, Luckow et al. 2003, Miller et al. 2003, Murphy et al. 2003). These studies suggest that the genus Acacia s.l. should be separated into as many as five Phytologia (Dec 2005) 87(3) 14] genera. These genera, for the most part, correspond to the four currently recognized infrageneric taxa of Acacia, namely the former subgenus Acacia, subgenus Aculeiferum sect. Aculeiferum and sect. Filicinae, and subgenus Phyllodineae, and a small group of North and Central America species related to Acacia coulteri Benth. The genus Acacia s.l., has recently been retypified as a result of endorsement of a proposal by Orchard and Maslin (2003); the type was changed from Acacia nilotica to Acacia penninervis (subgenus Phyllodineae) (McNeill, personal communication). The morphological and genetic differences separating the subgenera of Acacia s./., along with the change to a new type for the genus necessitate the following new combinations. The earliest valid name available for members of Acacia subgenus Acacia is Vachellia Wight & Arnott (1834). This name honors Reverend G. H. Vachell who collected extensively in China. Of these five groups, the former subgenus Acacia (series Gummiferae) consists of approximately 163 species, 60 species in the Americas, 73 species in Africa, 36 species in Asia (including ca. 15 that are also found in Africa), and 9 species in Australia (Maslin et al., 2003; Orchard and Maslin, 2003; http://www.worldwidewattle.com). Members of this subgenus can be separated from other members of the genus Acacia s./. by a combination of characters: the presence of stipular spines, the absence of prickles, the presence of an involucre on the peduncle, bipinnately compound leaves, colporate pollen apertures, pollen columellae, and the smooth pollen exine ornamentation. Descriptions of most New World species of this subgenus are available in publications completed by the authors and their students during the past 20 years (Clarke et al. 1989, 1990, 2006, Ebinger et al. 2000, 2002, Lee et al. 1989, Seigler and Ebinger 1988, 1995). Complete synonymies are provided below. In accord with the ICBN and an anonymous editor, we are using variety for all subspecific categories (Turner and Nesom, 2000) 142 Phytologia (Dec 2005) 87(3) VACHELLIA Wight & Arm., Prodr. fl. Ind. orient. 1: 272. 1834. — TYPE: Vachellia farnesiana (L.) Wight & Arn., based on Mimosa farnesiana L.. Sp. Pl. 521. 1753. VACHELLIA ACUIFERA (Benth.) Seigler & Ebinger, comb. nov. Basionym: Acacia acuifera Benth., London J. Bot. 1: 496. 1842. Bahamia acuifera (Benth.) Britton & Rose, N. Amer. Fl. 23: 86. 1928. TYPE: BAHAMAS. W. Swainson s.n. (holotype: K). Pithecellobium micrantha Benth., Trans. Linn. Soc. London 30: 589. 1875. Cojoba micrantha (Benth.) Britton & Rose, N. Amer. FI. 23: 30. 1928. — TYPE: DOMINICAN REPUBLIC. Santo Domingo, 1852, R. Schomburgk 149 (holotype: B, destroyed; isotypes: F fragment and photo, P, NY fragment). NOTE: For additional information, see Rico-Arce 1991 and Barneby 1992). VACHELLIA ALBICORTICATA (Burkart) Seigler & Ebinger, comb. nov. Basionym: Acacia albicorticata Burkart, Darwiniana 7: 504. 1947. - TYPE: ARGENTINA. SALTA: Tartagal, alt. 500 m, 29 Oct 1924, R. Schreiter 3355 (holotype: SI; isotype: ele): VACHELLIA ALLENII (D. H. Janzen) Seigler & Ebinger, comb. nov. Basonym: Acacia allenii D.H. Janzen, Smithsonian Contr. Bot. 13: 53. 1974. - TYPE: COSTA RICA. PUNTARENAS: Osa Peninsula, bank of tributary of Rio Agua Buena, 3.5 mi. SW of Rincon, 9 Mar 1967, D. H. Janzen 1769 (holotype: US; isotypes: F, GH, K, MEXU, MICH, MO, NY, UC). VACHELLIA ANEGADENSIS (Britton) Seigler & Ebinger, comb. nov. Basionym: Acacia anegadensis Britton, Mem. New York Bot. Gard. 6: 572. 1916. Fishlockia anegadensis (Britton) Britton & Rose, N. Amer. Fl. 23: 91. 1928. — TYPE: BRITISH VIRGIN ISLANDS. Anegada, rocky plains, The Settlement, 19- 20 Feb 1913, N. L. Britton & W. C. Fishlock 990 (holotype: NY; isotypes: F, GH, NY, US). Phytologia (Dec 2005) 87(3) 143 VACHELLIA AROMA (Gillies ex Hook. & Arn.) Seigler & Ebinger, comb. nov. Basionym: Acacia aroma Gillies ex Hook. & Arn., Bot. Misc. 3: 206. 1833. Acacia lutea (Mill.) Britton var. aroma (Gillies ex Hook. & Arn.) Kuntze, Revis. gen. pl. 3(2): 47. 1898. Vachellia lutea (Mill.) Speg. var. aroma (Gillies ex Hook. & Arn.) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 307. 1924. Vachellia lutea (Mill.) Speg. f. aroma (Gillies ex Hook. & Arn.) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 313. 1924.—- TYPE: ARGENTINA. a little way south of San Juan, Dec 1823, J. Gillies s.n. (holotype: K). VACHELLIA AROMA (Gillies ex Hook. & Arn.) Seigler & Ebinger var. AROMA Acacia moniliformis Griseb., Abh. Konigl. Ges. Wiss. Gottingen 19: 136. 1874. Acacia lutea (Mill.) Britton var. moniliformis (Griseb.) Kuntze, Rev. gen. pl. 3(2): 47. 1898 (as monoliformis). Vachellia lutea (Mill.) Speg. var. moniliformis (Griseb.) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 303. 1924. Vachellia lutea (Mill.) Speg. f. moniliformis (Griseb.) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 312. 1924.- TYPE: ARGENTINA. TUCUMAN: La Cruz, 20-24 Apr 1872, T. G. Lorentz 190 (holotype: GOET). Acacia michelii Rusby, Mem. Torrey Bot. Club 6: 28. 1896. — TYPE: BOLIVIA. vic. Cochabamba, 1891, M. Bang 1095 (holotype: K; isotype: NY). Acacia aroma Gillies ex Hook. & Arn. var. cochlearis Griseb., Abh. Konigl. Ges. Wiss. G6ttingen 24: 122. L879. c= TYPE: ARGENTINA. CORDOBA: north of San Javier, 18 Mar 1876, T. G. Hieronymus 558 (holotype: GOET). Vachellia lutea (Mill.) Speg. f. oocephala Speg., Bol. Acad. Nac. Ci. Cordoba 26: 312. 1924.—TYPE: none cited. Vachellia lutea (Mill.) Speg. f. thlipsacantha Speg., Bol. Acad. Nac. Ci. Cordoba 26: 312. 1924.— TYPE: none cited. 144 Phytologia (Dec 2005) 87(3) Vachellia lutea (Mill.) Speg. var. aroma (Gillies ex Hook. & Arn.) Speg. f. leptocarpa Speg., Bol. Acad. Nac. Ci. Cordoba 26: 305. 1924.— TYPE: none cited. Vachellia lutea (Mill.) Speg. var. aroma (Gillies ex Hook. & Arn.) Speg. f. pachycarpa Speg., Bol. Acad. Nac. Ci. Cordoba 26: 307. 1924.— TYPE: none cited. VACHELLIA AROMA (Gillies ex Hook. & Arn.) Seigler & Ebinger var. HUARANGO (Ruiz ex J. Macbr.) Seigler and Ebinger, comb. nov. Basionym: Acacia huarango Ruiz ex J. Macbr., Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 90. 1930. Acacia aroma Gillies ex. Hook. & Arn. var. huarango (Ruiz ex J. Macbr.) Ebinger, Seigler and Clarke, Syst. Bot. 25: 602. 2000. — TYPE: PERU. HUANUCO: near Huanuco, low spreading shrub, Ruiz & Pavon s.n. [lectotype, designated by Macbride (1943): F]. VACHELLIA ASTRINGENS (Gillies) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 278. 1924. Prosopis astringens Gillies in Hook. & Arn. Bot. Misc. 3: 204. 1833. Acacia atramentaria Benth., London J. Bot. 1: 392. 1842. Acacia farnesiana y atramentaria (Benth.) Kuntze, Revis. gen. pl. 3(2): 47. 1898. (based on same type). — TYPE: ARGENTINA. Mendosa and San Juan, J. Gillies s.n. (holotype: GL; isotype: K). NOTE: Not Acacia astringens A. Cunn. ex G. Don (1832), a species with phyllodes native to Australia and a synonym of Acacia falciformis DC. (Maslin, 2001a). Not Acacia adstringens Mart. in Spix and Martius (1828), an unarmed species of legume from Brazil that is not an Acacia. Acacia prosopoma O. Schnyd., Anales Soc. Ci. Argent. 3: 152. 1877. — TYPE: none cited, from Argentina, Mendoza, Cordoba. VACHELLIA BARAHONENSIS (Urb. & Ekman) Seigler & Ebinger, comb. nov. Basionym: Acacia barahonensis Urb. & Ekman in Urb Ark» Bot: 22a .(8): 292 19283) — “DYPE: DOMINICAN REPUBLIC. BARAHONA: between Coueve-en- Phytologia (Dec 2005) 87(3) 145 Haut and Trujin, quaternary calcareous soil, 22 Sep 1926, E. L. Ekman H7043 (holotype: B destroyed; isotype: S). VACHELLIA BELAIRIOIDES (Urb.) Seigler & Ebinger, comb. nov. Basionym: Acacia belairioides Urb., Symb. antill. 9: 439. 1928. Feracacia belairioides (Urb.) Britton and Leon, N. Amer. Fl. 23: 87. 1928. — TYPE: CUBA. HOLGUIN: Holguin, in carrascales (serpentine barrens) toward El Paraiso, 26 Aug 1916, E. L. Ekman 7589 (holotype: S; isotypes: MT, NY). VACHELLIA BIACICULATA (S. Watson) Seigler & Ebinger, comb. nov. Basionym: Acacia biaciculata S. Watson, Proc. Amer. Acad. Arts 21: 452. 1886. Acaciopsis biaciculata (S. Watson) Britton & Rose, N. Amer. Fl. 23: 96. 1928. — TYPE: MEXICO. CHIHUAHUA: on sandy plains near Chihuahua, 20 Aug 1885, C. G. Pringle 662 (holotype: US; isotypes: F, NY, VI): VACHELLIA BILIMEKII (J. Macbr.) Seigler & Ebinger, comb. nov. Basionym: Acacia bilimekii J. Macbr., Contr. Gray Herb. 59: 6. 1919. Acaciopsis bilimekii (J. Macbr.) Britton & Rose, N. Amer. Fl. 23: 94. 1928. — TYPE: MEXICO. MORELOS: Cuernavaca, 10 Jan 1866, D. Bilimek 128 (holotype: GH). Acacia ambigua Rose, Contr. U. S. Natl. Herb. 8: 31. 1903. — TYPE: MEXICO. PUEBLA: on the hills above Matamoras, where it is common, 26 Jun 1899, J. N. Rose & W. Hough 4698 (holotype: US). NOTE: Not Acacia ambigua Hoffannsegg (1826), which is based on a sterile specimen from Senegal, the identity of which is unknown (Ross 1979). Not Acacia ambigua Vogel (1836), a species from the Caribbean with prickles and lacking stipular spines, which is a synonym of Acacia vogeliana Steud. (Britton and Rose 1928). Acacia sericocarpa Rose, Contr. U. S. Natl. Herb. 8: 300. 1905. — TYPE: MEXICO. PUEBLA: on the hills above Matamoras, where it is common, 26 Jun 1899, J. N. Rose & W. Hough 4698 146 Phytologia (Dec 2005) 87(3) (holotype: US). NOTE: Not Acacia sericocarpa Fitzgerald (1904), a phyllodinous acacia species from Western Ausralia (Maslin, 2001b). VACHELLIA BRANDEGEANA (I. M. Johnst.) Seigler & Ebinger, comb. nov. Basionym: Acacia brandegeana I. M. Johnst., Contr. Gray Herb. 75: 27. 1925. Acaciopsis brandegeana (1. M. Johnst.) Britton & Rose, N. Amer. Fl. 23: 94. 1928.— TYPE: MEXICO. BAJA CALIFORNIA SUR: Agua Verde Bay, 26 May 1921, 7. M. Johnston 3881 (holotype GH; isotypes: A, F, MO, NY, UC, US). Pithecollobium acuminatum M. E. Jones, Contr. W. Bot. 18: 38. 1933. — TYPE: MEXICO. BAJA CALIFORNIA SUR: above Primiera Aqua near Loreto, 20 Oct 1930, M. E. Jones 27255 (holotype: RSA). VACHELLIA BRAVOENSIS (lsely) Seigler & Ebinger, comb. et stat. nov. Basionym: Acacia schaffneri (S. Watson) F. J. Herm. var. bravoensis Isely, Sida 3: 383. 1969. - TYPE: UNITED STATES. TEXAS: San Patricio Co., 7 miles S of Taft in clay loam soil, 29 Mar 1950, F. B. Jones 100 (holotype: SMU). VACHELLIA BUCHERI (Marie-Victorin) Seigler & Ebinger, comb. nov. Basionym: Acacia bucheri Marie-Victorin, Contr. Inst. Bot. Univ. Montréal 49: 57. 1945. —- TYPE: CUBA. ORIENTE: Moa, sur la plateforme littorale serpentine-limonitique, 27-31 May 1943, F. Marie-Victorin & Bro. Clément 21804 [holotype: MT, F photo; isotypes: F, HAC (Alvarez and Oliver 1991), MO, NY]. VACHELLIA CALIFORNICA (Brandegee) Seigler & Ebinger, comb. nov. Basionym: Acacia californica Brandegee, Proc. Calif. Acad. Sci. II. 3: 221. 1892. Acacia pringlei Rose subsp. californica (Brandegee) Lee, Seigler & Ebinger, Syst. Bot. 14: 99. 1989. Acaciopsis californica (Brandegee) Britton & Rose, N. Amer) Fly 2327096 ;,))1928.. 0 TYPE 2 MEXICO, sBAJA CALIFORNIA SUR: La Palma, 31 Mar 1892, 7. S. Brandegee s.n. [lectotype, designated by Lee et al. (1989): UC]. Phytologia (Dec 2005) 87(3) 147 Acacia sonorensis Rose, Contr. U. S. Nat. Herb. 8:31. 1903. Acaciopsis sonorensis (Rose) Britton & Rose, N. Amer. Fl. 23: 95. 1928.— TYPE: MEXICO. SONORA: near Guaymas, 5-11 Jun 1897, J. N. Rose 1247 (holotype: US). VACHELLIA CAMPECHIANA (Mill.) Seigler & Ebinger, comb. nov. Basonym: Mimosa campeachiana Mill. Gard. dict. ed 8, no. 20. 1768. Poponax campechiana (Mill.) Britton & Rose, N. Amer. Fl. 23: 90. 1928. Acacia cymbispina Sprague & Riley in Riley, Kew Bull. 1923: 394. 1923 (based on the same type). Poponax cymbispina (Sprague & Riley) Britton & Rose, N. Amer. Fl. 23: 90. 1928. Acacia milleriana Standl. Jour. Arnold Arb. 11: 29. 1928. (Based on the same type). (Rudd 1966, Seigler & Ebinger 1988). — TYPE: MEXICO. VERACRUZ: Veracruz, 1731, W. Houstoun s.n. (holotype: BM). NOTE: Not Acacia campechiana Schenck (1913), which is considered a synonym of the ant-acacia species Acacia cornigera (L.) Willd. by Seigler and Ebinger (1995). Note: We have used the spelling Vachellia campechiana. VACHELLIA CAMPECHIANA (Mill.) Seigler & Ebinger f. CAMPECHIANA Acacia cochliacantha Humb. & Bonpl. ex Willd., Sp. Pl. 4:1080. 1806. Mimosa cochliacantha (Humb. & Bonpl. ex Willd.) Poir. in Lam. Dict. Suppl. 1: 78. 1810. - TYPE: MEXICO. without exact locality [erroneously given as Guayaquil, Ecuador (Rudd 1966)], Humboldt & Bonpland s.n. (holotype: B destroyed, F photo, MO photo). Acacia cymbacantha Zucc. ex Benth., Trans. Linn. Soc. London 30: 501. 1875.—TYPE: MEXICO. W. Karwinski s.n. (holotype: M). NOTE: Acacia cymbacantha Zucc. ex Benth. is listed in synonomy in the original description of Acacia cochliacantha with the note “may be the same species with very much larger spines, at least 1 in. long and 1/2 in. broad”. 148 Phytologia (Dec 2005) 87(3) Poponax attenuata Britton & Rose, N. Amer. Fl. 23: 90. 1928. — TYPE: MEXICO. PUEBLA: Rio de Santa Lucia, in the vicinity of San Luis Tultitlanapa, near Oaxaca, Aug 1908, C. A. Purpus 3050 (holotype: NY). VACHELLIA CAMPECHIANA (Mill.) Seigler & Ebinger f. HOUGHII (Britton & Rose) Seigler & Ebinger, comb. nov. Basionym: Poponax houghii Britton & Rose, N. Amer. Fl. 23: 90. 1928. Acacia cochliacantha Humb. & Bonpl. ex Willd. f. houghii (Britton & Rose) Seigler & Ebinger, Syst. Bot. 13: 14. 1988. — TYPE: MEXICO. OAXACA: Tomellin Canon, 23 Jun 1899, J. N. Rose & W. Hough 4669 (holotype: NY; isotype US). VACHELLIA CAURINA (Barneby & Zanoni) Seigler & Ebinger, comb. nov. Basionym: Acacia caurina Barneby & Zanoni, Moscosoa 5: 20. 1989. — TYPE: HAITI. NORD-OUEST. sprawling shrub on rocks, arid, steep mountain slope west of bay, vicinity of Port a l'Ecu, 15-17 Mar 1929, E. C. Leonard & G. M. Leonard 13863 (holotype: US; isotypes: A, GH, NY). VACHELLIA CAVEN (Molina) Seigler & Ebinger, comb. nov. Basionym:'Mimosa caven Molina, Sag. stor. nat. Chili, ed 1, 174. 1782. Mimosa cavenia Molina, Eassai hist. nat. Chili, 338. 1789. ‘Acacia caven (Molina) Molina, Sag. stor. nat. Chili, 2nd. ed., 163, 299. 1810. Acacia cavenia (Molina) Hook. & Arn., Misc. Bot. 3: 206. 1833. Acacia farnesiana (L.) Willd. B cavenia (Molina) Kuntze, Revis. gen. pl. 3(2): 47. 1898. Acacia_farnesiana (L.) Willd. var. cavenia (Hook. & Arn.) Arechav., Anales Mus. Nac. Montevideo 1: 436. 1901.\!WVachellia farnesiana (L.) Wight & Arn. f. cavenia (Molina) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 297. 1923. ‘Acacia farnesiana (L.) Willd. f. cavenia (Hook. & Arn.) E. C. Clos., Bol. Minist. Agric. Nac. (Argentina) 28: 455. 1930.— TYPE: CHILE. Raneagna, Oct 1828, C. G. Bertero s.n. [lectotype, designated by Aronson (1992): SG0]. NOTE: We interpret the specific epithet cavenia to be a misspelling. Phytologia (Dec 2005) 87(3) 149 tAcacia adenopa Hook. & Arn., Bot. Misc. 3: 206. 1833. — TYPE: Islands of the Uruguay, J. Tweedie s.n. (holotype: K). ‘Acacia aromatica Poepp., Mart., Fl. Bras. 15: 395. 1879. — TYPE: CHILE. Valparaiso, FE. Poeppig 177 [lectotype, designated by Ebinger et al. (2000): W]. /Acacia farnesiana (L.) Willd. var. brachicarpa Kuntze, Revis. gen. pl. 1: 156. 1891.—TYPE: none cited, from Argentina, Cordoba. ‘Acacia farnesiana (L.) Willd. y heterocarpa Kuntze, Revis. gen. pl.. 3(2): 47. 1898. TYPE: ARGENTINA. CORDOBA: Dec 1891, O. Kuntze s.n. [lectotype, designated by Ebinger et al. (2000): MO]. “Acacia caven (Molina) Molina var. dehiscens Burkart ex Ciald., Darwiniana 25:76. 1984.- TYPE: ARGENTINA. CORDOBA: Ascochinga, 22 Sep 1936, E. G. Nicora 962 (holotype: SI). Acacia caven (Molina) Molina var. microcarpa (Speg.) Burkart ex Ciald., Darwiniana 25: 77. 1984. -Vachellia farnesiana (L.) Wight & Arn. f. microcarpa Speg., Bol. Acad. Nac. Ci. Cordoba 26: 301. 1924. - TYPE: ARGENTINA. FORMOSA: Depto. Patino, Fortin Soledad, A. Krapovickas 1283 [lectotype, designated by Cialdella (1984): SI; isotype: LIL]. \Acacia caven (Molina) Molina var. stenocarpa (Speg.) Burkart ex Ciald., Darwiniana 25: 78. 1984. ‘Vachellia farnesiana (L.) Wight & Arn. f. stenocarpa Speg., Bol. Acad. Nac. Ci. Cordoba 26: 301. 1924. TYPE: ARGENTINA. MISIONES: Depto. Candelaria, Santa Ana, A. Burkart 14734 [lectotype, designated by Cialdella (1984): SI]. “Acacia caven (Molina) Molina var. sphaerocarpa Burkart ex Aronson, Ann. Missouri Bot. Gard. 79: 964. 1992.— TYPE: ARGENTINA. CORRIENTES: ca. 27° 27'S, 58° 46'W, 60 m. "alrededores de la ciudad de Corrientes, antiguo camino a Matadero, 50 m de la ruta, 150 Phytologia (Dec 2005) 87(3) 17 Feb 1989, S. G.Tressens & A. Radovancich 3539 (holotype: K; isotype: CTES). ‘Acacia caven (Molina) Molina var. macrocarpa Aronson, Ann. Missouri Bot. Gard. 79: 965. 1992. - TYPE: ARGENTINA. SALTA: Salta, Chicoana, El Carril, 19 Oct 1948, A. Burkart 17577 (holotype: SI). ‘Vachellia farnesiana (L.) Wight & Arn. f. brachypoda Speg., Bol. Acad. Nac. Ci. Cordoba 26: 301. 1924.— TYPE: not cited, from Argentina, Buenos Aires Prov., San Fernanda, Gualeguaychu, Ibicuy y chaco santefecino. VACHELLIA X CEDILLOI (L. Rico Arce) Seigler & Ebinger, comb. nov. Basionym:/Acacia x cedilloi L. Rico Arce, Acta Bot. Mex. 26: 7. 1994.—- TYPE: MEXICO. QUINTANA ROO: km 6.5 de la carretera Vigia Chico-F. Carrillo Puerto, selva mediana subperennifolia, alt. 8 m, 9 Apr 1986, R. Villanueva 714 (holotype: MEXU: isotypes: CIQRO, XAL). VACHELLIA CHIAPENSIS (Saff.) Seigler & Ebinger, comb. nov. Basionym: :Acacia chiapensis Saff., J. Wash. Acad. Sci. 5: 356. 1915. — TYPE: MEXICO. CHIAPAS: near San Fernando, between Tuxtla and Chicoasén, 12 Jan 1907, G. N. Collins 164 (holotype: US, F photo). WACHELLIA CHORIOPHYLLA (Benth.) Seigler & Ebinger, comb. nov. Basionym: ‘Acacia choriophylla Benth., London J. Bot. 1: 495. 1842.-Lucaya choriophylla (Benth.) Britton & Rose, N. Amer. Fl. 23: 87. 1928.— TYPE: BAHAMAS: W. Swainson s.n. (holotype: K). WACHELLIA COLLINSII (Saff.) Seigler & Ebinger, comb. nov. Basionym: “Acacia collinsii Saff., Science N.S. 31: 677. 1910. \Myrmecodendron collinsii (Saff.) Britton & Rose, N. Amer. FI. 23: 92. 1928. — TYPE: MEXICO. CHIAPAS: between Chicoasén and San Fernandino, 1,005 ft., 13 Jan 1907, G. N. Phytologia (Dec 2005) 87(3) 15a Collins & C. B. Doyle 180 (holotype: US, F photo). NOTE: Safford (1910) listed G. N. Collins as the only collector, and cited 14 Jan. 1907 as the date collected. On the label of the type specimen G. N. Collins and C. B. Doyle are listed as the collectors and 13 Jan. 1907 is given as the date of the collection. J/Acacia glutea Ram. Goyena, Fl. Nicaragiiense 1: 382. 1909.— TYPE: not cited. NOTE: Although the original description of Ramirez Goyena is in Spanish and no type is cited (Rico Arce, 2001), this species was described before either of these was required. Because this name is not used by most workers, is not supported by type material, nor is a designated herbarium extant, in order to contribute to nomenclatural stability, we have made a proposal to conserve the name of Safford (1910), which is widely used. If accepted, the name for this taxon must become Vachellia collinsii (Saff.) Seigler & Ebinger. VAcacia costaricensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 361. 1913. Whyrmecodendron costaricensis (Schenck) Britton & Rose, Nee Amen MFl.. “232993700 1928. =, 8 TYPE: wCOSTASRICA: ALAJUELA: Alajuela, alt. 900 m, 1896, J. D. Smith 6488 [lectotype, designated by Seigler and Ebinger (1995): S (B, destroyed); isotypes: BM, GH, K, MO]. VAcacia panamensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 362. 1913.—TYPE: PANAMA. ALAJUELA: A. Koch s.n. (holotype: B, destroyed). VYAcacia yucatanensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 361. 1913. TYPE: MEXICO. YUCATAN, 1895, G. F. Gaumer 353 [lectotype, designated by Seigler and Ebinger (1995): US (B, destroyed); isotypes: BM, GH, F, MO]. Wadia nelsonii Saff., J. Wash. Acad. Sci. 4: 363. 1914. — TYPE: MEXICO. GUERRERO: Acapulco, sea level, 30 Apr 1903, E. W. Nelson 7024 (holotype: US; isotypes: F, GH). 152 Phytologia (Dec 2005) 87(3) “Acacia penonomensis Saff., J. Wash. Acad. Sci. 4: 363. 1914. — TYPE: PANAMA. COCLE: Penonomé, 50-1,000 ft., 23 Feb-22 Mar 1908, R. S. Williams 113 (holotype: NY, F photo, US fragment and photo; isotype: US). VACHELLIA CONSTRICTA (Benth.) Seigler & Ebinger, comb. nov. Basionym: Acacia constricta Benth. in Gray, Pl. Wright. 1: 66. 1852. -Acaciopsis constricta (Benth.) Britton & Rose, N. Amer. Fl. 23: 96. 1928. - TYPE: MEXICO. CHIHUAHUA: May-Oct 1849, C. Wright 162 (holotype: K; isotypes: GH, NY, US). “Acacia constricta Benth. var. paucispina Wooton & Standl., Bull. Torrey Bot. Club 36: 105. 1909.“Acaciopsis constricta Benth. var. paucispina (Wooton & Stand.) Moldenke, Revista Sudamer. Bot. 4: 15. 1937. — TYPE: UNITED STATES. NEW MEXICO: Sierra Co., Animas Creek in the Black Range, alt. 1500 ft., 13 Jul 1904, O. B. Metcalfe 1123 (holotype: US). “Eee arcuata M. Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10: 310. 1843.— TYPE: MEXICO. OAXACA: se trouve sur le flanc des montagnes calcaires et cactiferes, a l’est de Tehuacan de las Grandas, vers 6,000 pieds. Fl. jaunes, H. Galeotti 3222 (holotype: BR; isotypes: G, K). (Rudd 1976-1982). NOTE: Not Acacia arcuata Sieber ex Sprengel (1826), a taxon with phyllodes that is a synonym of Acacia melanoxylon R. Br. (Cowan and Maslin, 2001). In response to the recent conservation of the type for Acacia with an Australian species, the necessity to transfer species of the former subgenus Acacia to the genus Vachellia, the fact that the name Acacia constricta Benth. has been used by most workers, because this name is widely employed in floras of both Mexico and the United States, and in order to contribute to nomenclatural stability, we have made a proposal to conserve the name of Bentham (1852). If accepted, the name for this taxon must become Vachellia constricta (Benth.) Seigler & Ebinger. Phytologia (Dec 2005) 87(3) 153 VACHELLIA COOKII (Saff.) Seigler & Ebinger, comb. nov. Basionym: “Acacia cookii Saff., Science N. S. 31: 677. 1910. ‘Myrmecodendron cookii (Saff.) Britton & Rose, N. Amer. Fl. 23: 93. 1928. TYPE: GUATEMALA. ALTA VERAPAZ: Finca Trece Aguas, near Secanguim, alt. 300 m, 8 Mar 1907, G. P. Goll 102 (holotype: US). SAcacia bucerophora B. L. Rob., Proc. Amer. Acad. Arts 49: 502. 1913. - TYPE: BELIZE. Belize, about Toledo, 4 mi. from the coast and 5 miles from Punta Gorda, sea level, 29 Mar 1907, M. E. Peck 632 (holotype: GH, F fragment and photo, US fragment and photo; isotypes: F, K). VACHELLIA CORNIGERA (L.) Seigler & Ebinger, comb. nov. Basionym: “imosa cornigera L., Sp. Pl. 520. 1753. Acacia cornigera (L.) Willd., Sp. Pl. 4: 1080. 1806. Tauroceras cornigerum (L.) Britton & Rose, N. Amer. Fl. 23: 86. 1928. — TYPE: from a cultivated plant grown in the garden of George Clifford, between Haarlem and Leyden, Holland, collected by Linnaeus (No. 4) and bearing his label "Mimosa cornigera" presumably grown from Mexican seed (Rudd 1964). (holotype: BM, US fragment and photo). ‘Acacia spadicigera Schltdl. & Cham., Linnaea 5: 594. 1830. “Tauroceras spadicigerum (Schltdl. & Cham.) Britton & Rose, N. Amer. Fl. 23: 85. 1928.— TYPE: MEXICO. VERACRUZ: near La Laguna Verde, Mar 1928, C. Schiede & F. Deppe 685 [lectotype, designated by Seigler and Ebinger (1995): US fragment, (B, destroyed)]. ci ee campecheana Schenck, Repert. Spec. Nov. Regni Veg. 12: 361. 1913.— TYPE: MEXICO. CAMPECHE: von Chrismar s.n. (holotype: B, destroyed). /Acacia cubensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 360. 1913. - TYPE: CUBA. N coast, 21 Apr 1863, C. Wright 2402 [lectotype, designated by Seigler and Ebinger (1995): US, (B, 154 Phytologia (Dec 2005) 87(3) destroyed); isotypes: G, GOET, HAL, JE, K, MO, US]. NOTE: A note on one herbarium sheet (JE) indicates that the seeds came from Martinique. “Acacia interjecta Schenck, Repert. Spec. Nov. Regni Veg. 12: 361. 1913. — TYPE: Engler 3870a [lectotype, designated by Seigler and Ebinger (1995): JE, (B, destroyed)]. NOTE: From material growing in the Singapore and Kew Botanical Gardens (Janzen 1974). ‘Acacia nicoyensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 360. 1913. — TYPE: COSTA RICA. GUANACASTE: | shore of the Gulf of Nicoya, sea level, Feb 1900, A. Tonduz 13538 [lectotype, designated by Seigler and Ebinger (1995): US, (B, destroyed), isotypes: BM, GH, K, NY, US]. /Acacia rossiana Schenck, Repert. Spec. Nov. Regni Veg. 12: 361. 1913. — TYPE: MEXICO. VERACRUZ: Santa Lucrezia, Isthmus of Tehuantepec, 8 Oct 1906, H. Ross 91/8 [lectotype, designated by Seigler and Ebinger (1995): M, US photo]. Acacia furcella Saff., J. Wash. Acad. Sci. 4: 359. 1914. — TYPE: MEXICO. VERACRUZ: shore of Lake Catemaco, southern Veracruz, alt. 1,000 ft., 26 Apr 1894, E. W. Nelson 427 (holotype: US). Acacia hernandezii Saff., J. Wash. Acad. Sci. 4: 358. 1914. — TYPE: MEXICO. SAN LUIS POTOSI: vicinity of Rascon, 19-22 Jul. 1905, E. Palmer 699 (holotype: US; isotypes: F, GH, NY). Acacia turgida Saff., in W. M. Wheeler, Bull. Mus. Comp. Zoology Harvard Coll. 90: plate 45, 1942.— TYPE: (holotype: plate 45 in Wheeler 1942). “Acacia cornigera (L.) Willd. var. americana DC. Prodr. 2: 460. 1825. — TYPE: none cited. Phytologia (Dec 2005) 87(3) ie) A ACHELLIA CUCUYO (Barneby & Zanoni) Seigler & Ebinger, comb. nov. Basionym: Acacia cucuyo Barneby & Zanoni, Moscosoa 5: 14. 1989. —- TYPE: DOMINICAN REPUBLIC. AZUA: Sierra Martin Garcia, Loma del Copey, approximately 3 km (by air) SW of Barrero, 18° 18'N, 70° 56'W, alt. 780 m, 23 Feb 1987, T. Zanoni, J. Pimentel, & R. Garcia 38370 (holotype: JBSD; isotypes: FLAS, FTG, GH, K, MO, NY, S, TEX). VACHELLIA CURVIFRUCTA (Burkart) Seigler & Ebinger, comb. nov. Basionym: “Acacia curvifructa Burkart, Las Leguminosas Argentinas, 2nd ed., 97, 541, 1952. —- TYPE: PARAGUAY. CHACO: Puerto Casado, Dec 1916, T. Rojas 2138 (holotype: SI). VACHELLIA DAEMON (Ekman & Urb.) Seigler & Ebinger, comb. nov. Basionym: Acacia daemon Ekman & Urb., Symb. antill. 9: 438. 1928, #eracacia daemon (Ekman & Urb.) Britton & Rose N. Amer. Fl. 23: 86. 1928. —- TYPE: CUBA. MATANZAS: prope Ceiba Mocha in cuabales ad Canasi versus locis valde sterilibus, | Mar 1924, E. L. Ekman 18591 (holotype: S; isotypes: A, F, G, HAC photo, ILL, K, NY, US). VACHELLIA FARNESIANA (L.) Wight & Arm., Prodr. fl. Ind. orient. 272. 1834. Mimosa farnesiana L., Sp. pl. 521. 1753. Acacia farnesiana (L.) Willd., Sp. pl. 4: 1083. 1806. Poponax -farnesiana (L.) Raf., Sylva tellur. 118. 1838.— TYPE: Aldinus, Exactissima descriptio rariorum plantarum Romae in Horto Franesiano: 2-7, 1625, which provides, under the name Acacia Indica Farnesiana, a detailed description and two illustrations of a plant in cultivation in the garden of Cardinal Farnese in Rome [lectotype, designated by Ross (1975)]. VACHELLIA FARNESIANA (L.) Wight & Arn. var. FARNESIANA Acacia pedunculata Willd., Sp. pl. 4: 1084. 1806. Mimosa pedunculata (Willd.) Poir., in Lam. Encycl. Suppl. 1:81. 1810. \Acacia farnesiana (L. ) Willd. f. pedunculata (Willd.) Kuntze, 156 Phytologia (Dec 2005) 87(3) Revis. gen. pl. 3(2): 47. 1898.— TYPE: JAVA: (holotype: B- Willd.). vAcacia acicularis Humb. & Bonpl. ex Willd., Enum. pl. 1056. 1809. — TYPE: in America meridionali (holotype: B-Willd.). “Acacia edulis Humb. & Bonpl. ex Willd., Enum. pl. 1056. 1809. — TYPE: in America meridionali (holotype: B-Willd.). “Acacia ferox M. Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10: 314. 1843. — TYPE: MEXICO. OAXACA: se trouve abondamment dans les plaines d'Oaxaca, de Tlacolula et d'Etla, de 5,000 pieds, Fl. jaunes, H. Galeotti 3223 (holotype: BR; isotype: K). NOTE: For more information, see Rudd 1976-1982. /Acacia lenticellata F. Muell., J. Proc. Linn. Soc., Bot. 3: 147. 1859. VAcacia farnesiana (L.) Willd. var. lenticellata (F. Muell.) F. M. Bailey, Compr. Queensland PI. 164. 191352, TYPE: AUSTRALIA. NORTHERN TERRITORIES: McArthur River, F. Mueller 43 (Pedley 1979). Acacia smallii Isely, Sida 3: 384. 1969.— TYPE: UNITED STATES. LOUISIANA: Lafourche Parish, along Bayou La Fourche near Cut-off, 16 Apr 1931 and Aug 1931, J. K. Small & E. J. Alexander s.n. (holotype: NY; isotypes: US, WIS). Farnesia odora Gasp., Descr. nuov. gen. Leg. 1836. — TYPE: none cited [lectotype, designated here, plate of Farnesia odora in Gasparrini (1836)]. Poponax venosa Britton in Britton and Killip, Ann. New York Acad. Sci. 35: 139. 1936. — TYPE: COLOMBIA. ANTIOQUIA: vicinity of Medellin, 20 Aug 1927, R. A. Toro 478 (holotype: NY). Vachellia densiflora Alexander ex Small, Man. s. e. fl. 655, 1505. 1933.Acacia densiflora (Alexander ex Small) Cory, Rhodora 38: Phytologia (Dec 2005) 87(3) Py 406. 1936. Acacia minuta (M. E. Jones) R. M. Beauch. subsp. densiflora (Alexander ex Small) R. M. Beauch., Phytologia 46: 7. 1980. - TYPE: UNITED STATES. LOUISIANA: Lafourche Parish, along Bayou La Fourche near Cut-off, 16 Apr 1931 and Aug 1931, J. K. Small & E. J. Alexander s.n. (holotype: NY; isotypes: US, WIS). VACHELLIA FARNESIANA (L.) Wight & Arn. var. MINUTA (M. E. Jones) Seigler & Ebinger, comb. et stat. nov. Basionym: Pithecellobium minutum M. E. Jones, Contr. West Bot. 18: 38. 1933. Acacia minuta (M. E. Jones) R. M. Beauch., Phytologia 46: 5. 1980. Acacia farnesiana (L.) Willd. subsp. minuta (M. E. Jones) Ebinger & Seigler, Southw. Naturalist 47: 90. 2002. — TYPE: MEXICO. BAJA CALIFORNIA: Cacachilla Mts., 2 Oct 1930, M. E. Jones 27265 (holotype: POM-RSA). VACHELLIA FARNESIANA (L.) Wight & Arn. var. PINETORUM (F. J. Herm.) Seigler & Ebinger, comb. et stat. nov. Basionym: Acacia pinetorum F. J. Herm., J. Wash. Acad. Sci. 38: 237. 1948. Acacia farnesiana subsp. pinetorum (F. J. Herm.) Ebinger and Seigler, Southw. Naturalist 47: 90. 2002. —- TYPE: UNITED STATES. FLORIDA: Dade Co., hammocks, Long Key (Everglades), 18-26 Jan 1909, J. K. Small & J. J. Carter 2975; in pinelands, Long Key, 6-7 May 1903, J. K. Small & P. Wilson 1778 (syntypes: NY). Vachellia insularis J. K. Small, Man. s. e. fl. 655, 1505. 1933. — TYPE: UNITED STATES. FLORIDA: Monroe Co., pinelands, Big Pine Key, 21 Mar 1915, J. K. Small & C. A. Mosier 6018; pinelands, Big Pine Key, 27 Feb 1911, J. K. Small, J. J. Carter & G. K. .Small 3549 (syntypes: NY). NOTE: Not Acacia insularis A. Richard (1845), an herbaceous legume lacking stipular spines and prickles that is a synonym of Neptunia pubescens Benth. (Bassler 1998). Vachellia peninsularis J. K. Small, Man. s. e. fl. 654, 1505. 1933. — TYPE: UNITED STATES. FLORIDA: Dade Co., hammocks, 158 Phytologia (Dec 2005) 87(3) Long Key (Everglades), 18-26 Jan 1909, J. K. Small & J. J. Carter 2975; in pinelands, Long Key, 6-7 May 1903, J. K. Small & P. Wilson 1778 (syntypes: NY). NOTE: Not Senegalia peninsularis Britton & Rose (1928), a species from Baja California, Mexico that has prickles and lacks stipular spines, nor Acacia peninsularis (Britton & Rose) Standley (1936). VACHELLIA GENTLEI (Standl.) Seigler & Ebinger, comb. nov. Basionym: Acacia gentlei Standl., Publ. Field Mus. Nat. Hist., Bot.:Ser.) 222 77s 1940.c=> TYPE: “ BELIZES PGenile 185 (holotype: F). VACHELLIA X GLADIATA (Saff.) Seigler & Ebinger. comb. nov. Basionym: Acacia x gladiata Saff., J. Wash. Acad. Sci. 5: 359. 1915. Myrmecodendron gladiatum (Saff.) Britton & Rose, N. Amer. Fl. 23: 92. 1928. = TYPE: MEXICO. SINALOA: vicinity of Rosario, 1849, J. Gregg 1135 (holotype: MO, F photo, NY photo, US fragment and photo). VACHELLIA GLANDULIFERA (S. Watson) Seigler & Ebinger, comb. nov. Basionym: Acacia glandulifera S. Watson, Proc. Amer. Acad. Arts 25: 147. 1890. Poponax glandulifera (S. Watson) Britton & Rose, N. Amer. Fl. 23: 88. 1928. — TYPE: MEXICO. COAHUILA: limestone hills, Carneros Pass, 7 Sep 1889, C. G. Pringle 2861 (holotype: GH; isotypes: F, NY, VT). NOTE: Not Acacia glandulifera Schinz (1900) which is considered a synonym of Acacia nebrownii Burtt Davy, a species with stipular spines from Africa (Ross 1979). VACHELLIA GLOBULIFERA (Saff.) Seigler & Ebinger, comb. nov. Basionym: Acacia globulifera Saff., J. Wash. Acad. Sci. 4: 360. 1914. Myrmecodendron globulifera (Saff.) Britton & Rose, N. Amer. Fl. 23: 93. 1928 — TYPE: MEXICO. YUCATAN: at the port of Silam (Tzilam), N coast of Yucatan, Apr 1895, G. F. Gaumer 1909 (holotype: F, US fragment and photo; isotypes: GH, MO, NY, US). Phytologia (Dec 2005) 87(3) 159 Acacia donnelliana Saff., J. Wash. Acad. Sci. 4: 361. 1914. Myrmecodendron donnelliana (Saff.) Britton & Rose, N. Amer. FI. 23: 93. 1928 - TYPE: HONDURAS. SANTA BARBARA: San Pedro de Sula, Cortés, alt. 600 ft., Mar 1888, C. Thieme 5216 (holotype: US). VACHELLIA GUANACASTENSIS (Clarke, Seigler & Ebinger) Seigler & Ebinger, comb. et stat. nov. Basionym: Acacia farnesiana (L.) Willd. var. guanacastensis Clarke, Seigler & Ebinger, Syst. Bot. 14: 562. 1989. Acacia guanacastensis (Clarke, Seigler & Ebinger) Ebinger & Seigler, Syst. Bot. 25: 610. 2000.— TYPE: COSTA RICA. GUANACASTE: Great Swamp, Comelco, near Bagaces, 11 May 1976, D. H. Janzen 10362 (holotype: MO; isotype: ILL). VACHELLIA HINDSII (Benth.) Seigler & Ebinger, comb. nov. Basionym: Acacia hindsii Benth., London J. Bot. 1: 504. 1842. Myrmecodendron hindsii (Benth.) Britton & Rose, N. Amer. FI. Z23209%. ) 1928 — "TYPE? ~ MEXICO): ) JALISCO:sshore sof Manzanilla Bay, sea level, 1841, R. B. Hinds 248 (holotype: K, F photo). Acacia bursaria Schenck, Repert. Spec. Nov. Regni Veg. 12: 363. 1913. — TYPE: GUATEMALA. AMATITLAN: Laguna Amatitlan, alt. 3,900 ft., Feb 1890, J. D. Smith 2304 [lectotype, designated by Seigler and Ebinger (1995): US (B, destroyed); isotypes: GH, K]. Acacia sinaloensis Saff., J. Wash. Acad. Sci. 4: 365. 1914. — TYPE: MEXICO. SINALOA: vicinity of Villa Union, growing about a pond, 2 Apr 1910, J. N. Rose, P. C. Standley & P. G. Russell 13972 (holotype: US; isotype: NY). Acacia tepicana Saff., J. Wash. Acad. Sci. 4: 366. 1914. — TYPE: MEXICO. NAYARIT: thickets, vicinity of Acoponeta, Tepic, alt. 30 m, 10 Apr 1910, J. N. Rose, P. C. Standley & P. G. Russell 14357 (holotype: US, F photo; isotype: NY). 160 Phytologia (Dec 2005) 87(3) VACHELLIA INSULAE-IACOBI (L. Riley) Seigler & Ebinger, comb. nov. Basionym: Acacia insulae-iacobi L. Riley, Bull. Misc. Inform. Kew 1925: 220. 1925. - TYPE: ECUADOR. GALAPAGOS: San Salvador (James Island), James Bay, in ravine near seashore, 27 Jul 1924, C. Penny & L. A. M. Riley 391 (holotype: K). VACHELLIA JANZENII (Ebinger & Seigler) Seigler & Ebinger, comb. nov. Basionym: Acacia janzenii Ebinger & Seigler, Southw. Naturalist 32: 245. 1987. — TYPE: MEXICO. TABASCO: 9.8 miles W of Lazaro Cardenas on hwy. 180, 17 Jun 1966, D. H. Janzen 515 (holotype: MO; isotypes: BM, CAS, F, GH, MICH, MO, UC, US). VACHELLIA MACRACANTHA (Humb. & Bonpl. ex Willd.) Seigler & Ebinger, comb. nov. Basionym: Acacia macracantha Humb. & Bonpl. ex Willd. Sp. pl. 4: 1080. 1806. Poponax macracantha (Humb. & Bonpl. ex Willd.) Killip in Little, Caribbean Forester 9: 241. 1948. Mimosa macracantha (Humb. & Bonpl. ex Willd.) Poir. in Lam., Encycl. Suppl. 1: 78. 1810. — TYPE: MEXICO. without exact locality, Humboldt & Bonpland s.n. (holotype: B-Willd., GH photo; isotypes: NY, P). NOTE: In response to the recent conservation of the type for Acacia with an Australian species, the necessity to transfer species of the former subgenus Acacia to the genus Vachellia, the fact that the name Acacia macracantha Humb. & Bonpl. ex Willd. has been used for this widespread species by most workers, because this name is widely employed in floras of both Latin America and the United States, and in order to contribute to nomenclatural stability, we have made a proposal to conserve the name of Bentham (1852). If accepted, the name for this taxon must become Vachellia macracantha (Humb. & Bonpl. ex Willd.) Seigler & Ebinger. Vachellia lutea (Mill.) Speg., Bol. Acad. Nac. Ci. Cordoba 26: 301. 1923. Mimosa lutea Mill., Gard. dict. ed. 8, 1768. Acacia lutea (Mill.) Britton, Bull. Torrey Bot. Club 16: 327. 1889. Poponax Phytologia (Dec 2005) 87(3) 161 lutea (Mill.) Britton & Rose, N. Amer. Fl. 23: 90. 1928.— TYPE: JAMAICA. 1731, W. Houstoun s.n. (holotype: BM; isotype: K). NOTE: Not Acacia lutea Leavenworth (1824) which is a synonym of Neptunea lutea (Leavenw.) Benth. (Correll and Johnston 1970). Acacia flexuosa Humb. & Bonpl. ex Willd., Sp. pl. 4: 1082. 1806. Mimosa flexuosa (Humb. & Bonpl. ex Willd.) Poir. in Lam., Encycl. Suppl. 1: 79. 1810. Poponax flexuosa (Humb. & Bonpl. ex Willd.) Britton & Rose in Britton & Killip, Ann. New York Acad. Sci. 35: 139. 1936.— TYPE: VENEZUELA. CUMANA: Humboldt & Bonpland 81 (holotype: B-Willd.; isotypes: HAL, P). Acacia flexuosa Humb. & Bonpl. ex Willd. 8 ambigua DC., Prodr. 2: 4635 1825:.— TYPE: GUADELOUPE. «C.-GBeriero: s.n: (holotype: G-DC). Acacia flexuosa Humb. & Bonpl. ex Willd. 8 /asiocarpa Griseb., Abh. KO6nigl. Ges. Wiss. G6ttingen 7: 211. 1857. —- TYPE: none cited, from Guadeloupe. Acacia obtusa Humb. & Bonpl. ex Willd., Sp. pl. 4: 1087. 1806. Mimosa obtusa (Humb. & Bonpl. ex Willd.) Poir. in Lam., Encycl. Suppl. 1: 82. 1810. - TYPE: VENEZUELA. ORINOCO: Humboldt & Bonpland s.n. (holotype: P-H.B.K.; isotype: B- Willd.). Acacia punctata Humb. & Bonpl. ex Willd., Sp. pl. 4: 1084. 1806. — TYPE: in America meridionali, Humboldt & Bonpland 1162 (holotype: B-Willd.). Acacia humboldtii Desv. in W. Ham., Prodr. Pl. Ind. Occid. 60. 1825. — TYPE: none cited, from Guyana. Acacia macracanthoides Bertero ex DC., Prod. 2: 463. 1825. Poponax macracanthoides (Bertero ex DC.) Britton & Rose, N. Amer. Fl. 23: 89. 1928.— TYPE: JAMAICA. C.G.Bertero s.n. (holotype: G-DC). 162 Phytologia (Dec 2005) 87(3) Acacia subinermis Bertero ex DC., Prodr. 2: 463. 1825. — TYPE: JAMAICA. C. G. Bertero s.n. (holotype; G-DC, MO photo). Acacia microcephala Macfad., Fl. Jamaica 1: 316. 1837. - TYPE: JAMAICA. at bridge beyond the second milestone on the windward road, J. Macfadyen s.n. (holotype: K). NOTE: Not Acacia microcephala Richard (1845) which has prickles and is considered a synonym of Acacia tenuifolia (L.) Willd. by Howard (1988). Acacia platyacantha Schlitdl., Linnaea 12: 565. 1838. — TYPE: MEXICO. HIDALGO: Mineral del Monte, C. A. Ehrenberg s.n. (holotype: B destroyed). Acacia pellacantha Vogel, Meyen, Nov. Actorum Acad. Caes. Leop.- Carol. Nat. Cur. 19. Suppl. 1: 45. 1843.— TYPE: PERU. Lima, Dombey, F. J. F. Meyen & Cuming, n. 1013. (holotype: not seen). Acacia macracantha Humb. & Bonpl. ex Willd. var. glabrescens Griseb., Fl. Brit. W. I. 222. 1860.— TYPE: JAMAICA. in dry plains, Mar, J. Macfadyen s.n. (holotype: K). Acacia macracantha Humb. & Bonpl. ex Willd. var. glabra Kitanov, Ann. Univ. Sofia Fac. Biol. 66(2): 31. 1974. —- TYPE: CUBA. ORIENTE: sur de Guantanamo en Novaliches, 30 Jun 1936, Hno. Leon s.n.; Cuba, Prov. Oriente, Santiago de Cuba-Caney, bordes del mar en Arroyo de la Costa, 9 Sep 1951, M. Lopez Figueiras s.n. (syntypes: SV). Mimosa atomaria Poir. in Lam., Encyle. Suppl. 1:81. 1810. - TYPE: none cited. Poponax cowellii Britton & Rose, N. Amer. Fl. 23: 89. 1928. Acacia cowellii (Britton & Rose) Leon in Leon & Alain, Contr. Ocas. Mus. Hist: Nat::Colecio “De Ma Salle) 9:58), 1950.5 aR: CUBA. ORIENTE: tree 5 m with spreading branches, hillside Phytologia (Dec 2005) 87(3) 163 thicket, vicinity of Santiago, 10-25 Mar 1912, N. L. Britton, E. G. Britton & J. F. Cowell 12593 (holotype: NY). Poponax canescens Britton ex Britton & Killip, Ann. New York Acad. Sci. 35: 139. 1936. Acacia canescens (Britton ex Britton & Killip) Garcia-Barriga & Forero-Gonzales, Cat. Il. PI. Cundinamarca Si oa POGSS ge) AEMERE: COLOMBIA. CUNDINAMARCA: Anapoima, alt. 400 m, 1851-1857, J. Triana 4439 (holotype: NY). NOTE: Not Acacia canescens Martens & Galeotti (1843), a member of subgenus Aculeiferum. VACHELLIA MAYANA (Lundell) Seigler & Ebinger, comb. nov. Basionym: Acacia mayana Lundell, Publ. Carnegie Inst. Wash. 478: 210. 1937.— TYPE: GUATEMALA. EL PETEN: near San Diego on the Rio Pasion, 10 Apr 1935, M. Aguilar H. 495 (holotype: MICH, isotypes: GH, NY, US). VACHELLIA MELANOCERAS (Beurl.) Seigler & Ebinger, comb. nov. Basionym: Acacia melanoceras Beurl., Kongl. Svenska Vetensk. Acad. Handl. 1854: 123. 1856. Myrmecodendron melanoceras (Beurl.) Britton & Rose, N. Amer. Fl. 23: 93. 1928. — TYPE: PANAMA. COLON: Portobello, sea level, Apr 1826, J. G. Billberg 289 [lectotype, designated by Seigler and Ebinger (1995): S, F photo, MO photo]. Acacia multiglandulosa Schenck, Repert. Spec. Nov. Regni Veg. 12: 362. 1913. - TYPE: PANAMA. COLON: Portobello, JZ. G Billberg 1825 [lectotype, designated by Seigler and Ebinger (1995): US photo (B, destroyed)]. VACHELLIA OVIEDOENSIS (R. Garcia & M. Mejia) Seigler & Ebinger, comb. nov. Basionym: Acacia oviedoensis R. Garcia & M. Mejia, Moscosoa 11: 7. 2000. — TYPE: DOMINICAN REPUBLIC. PEDERNALES: 22 km al W de Oviedo, El Papayo, lado Sur de la Carretera Oviedo-Pedernales. Bosque seco sobre roca caliza con: Guaiacum, Senna, Opuntia, Harrisia and Reynosia. 17°51'N, 71° 31'W, elev. 140 m, 13 May 1997, R. 164 Phytologia (Dec 2005) 87(3) Garcia & R. Pujols 6543 (holotype: JBSD; isotypes: B, F, MAPR, MO, NY, S). VACHELLIA PACENSIS (Rudd & A. M. Carter) Seigler & Ebinger, comb. nov. Basionym: Acacia pacensis Rudd & A. M. Carter, Madrofio 30: 177. 1983. — TYPE: MEXICO. BAJA CALIFORNIA SUR: La Paz, 1 Nov 1890, 7. S. Brandegee 190 (holotype: UC). VACHELLIA PENNATULA (Schltdl. & Cham.) Seigler & Ebinger, comb. nov. Basionym: /nga pennatula Schltdl. & Cham., Linnaea 5: 593. 1830. Acacia pennatula (Schltdl. & Cham.) Benth., London J. Bot. 1: 390. 1842. Poponax pennatula (Schltdl. & Cham.) Britton & Rose, N. Amer. Fl. 23: 88. 1928. — TYPE: MEXICO. VERACRUZ: municipio de Xico, region of Jalapa, Hacienda de la Laguna, Aug. 1828, C. Schiede & F. Deppe s.n. (lectotype: HAL). NOTE: The sheet containing the type material is a mixture from Jalapa and from near Hacienda de la Laguna. Schlechtendal & Chamisso (1830) mention both in the original description and both are Acacia pennatula. VACHELLIA PENNATULA (Schltdl. & Cham.) Seigler & Ebinger var. PENNATULA Acacia lanata M. Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10: 315. 1843. - TYPE: MEXICO. OAXACA: Misteca Alta, Apr 1840, a 7-8,000 pieds, H. Galeotti 3231 (holotype: BR; isotypes: G, P). (Rudd 1976-1982). Pithecollobium minutissimum M. E. Jones, Contr. W. Bot. 18: 38. 1933. — TYPE: MEXICO. JALISCO: Guadalajara, La Barranca, 25 Nov 1930, M. E. Jones 27266a (holotype: RSA-POM). VACHELLIA PENNATULA (Schltdl. & Cham.) Seigler & Ebinger var. PARVICEPHALA (Seigler & Ebinger) Seigler & Ebinger. comb. et stat. nov. Basionym: Acacia pennatula (Schltdl. & Cham.) Benth. subsp. parvicephala Seigler and Ebinger, Syst. Bot. Phytologia (Dec 2005) 87(3) 165 13: 12. 1988.— TYPE: MEXICO. YUCATAN: Chichén Itza, in second growth, 6 Jun 1938, C. L. Lundell & Lundell 7418 (holotype: F; isotypes: LL, NY, TEX, US). VACHELLIA POLYPYRIGENES (Greenm.) Seigler & Ebinger, comb. nov. Basionym: Acacia polypyrigenes Greenm. in Combs, Trans. Acad. Sci. St. Louis 7: 419 and plate 33. 1897. Poponax polypyrigenes (Greenm.) Britton & Rose, N. Amer. Fl. 23: 89. 1928. TYPE: CUBA. SANTA CLARA: District of Cienfuegos, not common, in open woods near long beach, near the entrance of the Bay of Cienfuegos, Faro Villa Nueva, E., 18 Sep 1895, R. Combs 602 (holotype: GH; isotypes: F, MO, NY, US). Acacia cupeyensis Leon in Leon and Alain, Contr. Ocas. Mus. Hist. Nat) Colegio "De La Salle” 9: 8) 1950: —-4PYPE> CUBA. ORIENTE: Puerto Padre, playa de El Cupey, 24 Feb 1930, M. Curbelo 512 [holotype: HAC - Herb. J. Roig 5075 (Bassler 1998); isotype: NY]. Acacia curbeloi Leon in Leon and Alain, Contr. Ocas. Mus. Hist. Nat. €olesio "De La ‘Salle? 0241. 195ie—/ TYPE: “CUBA. ORIENTE: Puerto Padre, playa de El Cupey, 1929, M. Curbelo 60 [lectotype: HAC - Herb. J. Roig 5075 (Bassler 1998)]. VACHELLIA PRINGLEI (Rose) Seigler & Ebinger, comb. nov. Basionym: Acacia pringlei Rose, Contr. U. S. Natl. Herb. 3: 316. 1895. Acaciopsis pringlei (Rose) Britton & Rose N. Amer. FI. 23: 95. 1928.— TYPE: MEXICO. OAXACA: Tomellin Canyon, alt. 3000 ft. a tree 20-30 ft. high, 22 Dec 1894, C. G. Pringle 6113 (holotype: US; isotypes: CM, F, MO, UC). Acacia unijuga Rose, Contr. U. S. Natl. Herb. 8:32. 1903. Acaciopsis unijuga (Rose) Britton & Rose, N. Amer. Fl. 28: 95. 1928. — TYPE: MEXICO. SAN LUIS POTOSI: Las Palmas, lowlands, 9 Mar 1899, C. G. Pringle 6989 (holotype: US; isotypes: CM, F, LE “MO;UG); 166 Phytologia (Dec 2005) 87(3) Acacia conzattii Standl., Contr. U. S. Natl. Herb. 20: 186. 1919. Acaciopsis conzattii (Standl.) Britton & Rose, N. Amer. Fl. 28: 95. 1928. TYPE: MEXICO. OAXACA: Estacién Almaloyas, alt. 700 m, Mar 1907, C. Conzatti 1756 (holotype: US). Acaciopsis sesquijuga Britton & Rose, N. Amer. Fl. 28: 95. 1928. Acacia sesquijuga (Britton & Rose) Standl., Field Mus. Nat. Hist. Bot. Ser. 3: 277. 1930.— TYPE: MEXICO. YUCATAN: Silam, G. F. Gaumer 1307 (holotype: MO). VACHELLIA RIGIDULA (Benth.) Seigler & Ebinger, comb. nov. Basionym: Acacia rigidula Benth., London J. Bot. 1: 504. 1842. Acaciopsis rigidula (Benth.) Britton & Rose, N. Amer. Fl. 23: 94. 1928. — TYPE: UNITED STATES. TEXAS: 7. Drummond 161 (holotype: K). VACHELLIA ROIGII (Leon) Seigler & Ebinger, comb. nov. Basionym: Acacia roigii Leon in Leon and Alain, Contr. Ocas. Mus. Hist. Nat. Colegio "De La Salle" 9: 7. 1950. — TYPE: CUBA. LAS TUNAS: Puerto Padre, yanales de El Cupey, 28 Feb 1930, M. Curbelo 290 [holotype: HAC -Herb. J. Roig 5077 (Bassler 1998); isotype: HAC (Alvarez and Oliver 1991)]. VACHELLIA RORUDIANA (Christophersen) Seigler & Ebinger, comb. nov. Basionym: Acacia rorudiana Christophersen, Nyt Mag. Naturvidensk. 70: 76. 1931. — TYPE: ECUADOR. GALAPAGOS: Santa Cruz Island (Indefatigable), Academy Bay, the dry zone, Dec 1926, B. Rorud 12 (holotype: O). Acacia tortuosa (L.) Willd. B glabrior Hook. f., Trans. Linn. Soc. London 20: 229. 1847. - TYPE: ECUADOR. GALAPAGOS: San Salvador (James Island), C. Darwin s.n. (holotype: K). VACHELLIA RUDDIAE (D. H. Janzen) Seigler & Ebinger, comb. nov. Basionym: Acacia ruddiae D. H. Janzen, Smithsonian Contr. Bot. 13: 34. 1974. TYPE: COSTA RICA. LIMON: ferry landing, Moin River, Moin, 13 Feb 1965, D. H. Janzen 828 Phytologia (Dec 2005) 87(3) 167 (holotype: US; isotypes: CAS, F, GH, MEXU, MICH, MO, NY, UC). VACHELLIA SCHAFFNERI (S. Watson) Seigler & Ebinger, comb. nov. Basionym: Pithecellobium schafjneri S. Watson, Proc. Amer. Acad. Arts 17: 352. 1882. Samanea schaffneri (S. Watson) J. Macbr., Contr. Gray Herb. 59: 2. 1919. Popanax schaffneri (S. Watson) Britton & Rose, N. Amer. Fl. 23: 89. 1928. Acacia schaffneri (S. Watson) F. J. Herm., J. Wash. Acad. Sci. 38: 236. 1948. TYPE: MEXICO. SAN LUIS POTOSI: in the mountains about San Luis Potosi, C. C. Parry & E. Palmer 219 [lectotype, designated by Rzedowski (1963): GH]. Acacia subtortuosa Shafer in Britton & Shafer, N. Amer. Trees, 524. 1908. — TYPE: MEXICO. DURANGO: near the city of Durango, Apr - Nov 1896, E. Palmer 11 (holotype: NY; isotypes: F, MO, UC). VACHELLIA SCHOTTII (Torr.) Seigler & Ebinger. comb. nov. Basionym: Acacia schottii Torr. in Emory, Rep. U. S. Mex. Bound. Surv. 2: 62. 1858. Acaciopsis schottii (Torr.) Britton & Rose, N. Amer. Fl. 23: 96. 1928. - TYPE: UNITED STATES. TEXAS: Brewster Co., near the canon of San Carlos, at the Comanche Crossing of the Rio Grande, Sep, C. C. Parry 330 (holotype: NY; isotype: GH). VACHELLIA SPHAEROCEPHALA (Schltdl. & Cham.) Seigler & Ebinger, comb. nov. Basionym: Acacia sphaerocephala Schltdl. & Cham., Linnaea 5: 594. 1830. — TYPE: MEXICO. VERACRUZ: Actopan, sea level, Mar 1829. C. Schiede & F. Deppe 684 [lectotype, designated by Seigler and Ebinger (1995): HAL (B, destroyed), US fragment and photo). Acacia veracruzensis Schenck, Repert. Spec. Nov. Regni Veg. 12: 362. 1913.— TYPE: MEXICO. VERACRUZ: sand dunes south of Veracruz, sea level, 13 Oct 1908, culta in hort. bot. Darmstadt., H. Schenck 916 [lectotype, designated by Seigler and Ebinger 168 Phytologia (Dec 2005) 87(3) (1995): M, (B, destroyed), US fragment and photo; isotypes: F, HAL]. Acacia dolichocephala Saff., J. Wash. Acad. Sci. 5: 355. 1915. — TYPE: MEXICO. VERACRUZ: along the shore north of the city of Veracruz, 24 Jan 1906, J. M. Greenman 87 (holotype: F; isotypes: GH, NY, US). VACHELLIA X STANDLEYI (Saff.) Seigler & Ebinger, comb. nov. Basionym: Acacia x standleyi Saff., J. Wash. Acad. Sci. 4: 367. 1914. Myrmecodendron standleyi (Saff.) Britton & Rose, N. Amer. Fl. 23: 92. 1928. — TYPE: MEXICO. NAYARIT: along the river in the vicinity of Acaponeta, Territory of Tepic, W Mexico, 11 Apr 1910, J. N. Rose, P. C. Standley & P. G. Russell 14374 (holotype: US, F photo). Acacia x hirtipes Saff., J. Wash. Acad. Sci. 4: 367. 1914. Myrmecodendron hirtipes (Saff.) Britton & Rose, N. Amer. FI. 23: 92. 1918. - TYPE: GUATEMALA. SANTA ROSA: along the Rio de las Cafias, 3,000 ft., Apr 1892, Hevde & Lux 3299b (holotype: US, NY photo). Myrmecodendron oaxacanum Britton & Rose, N. Amer. Fl. 23: 92. 1928. - TYPE: MEXICO. OAXACA: Fonameca, alt. 110 m, 3 Jul 1925, E. Makrinius 488 (holotype: US). VACHELLIA TORTUOSA (L.) Seigler & Ebinger, comb. nov. Basionym: Mimosa tortuosa L., Syst. Nat., 10th ed., 2: 1312. 1759. Acacia tortuosa (L.) Willd. Sp. Pl. 4: 1083. 1806. Poponax tortuosa (L.) Raf., Sylva tellur. 118. 1838. — TYPE: JAMAICA. without exact locality, Patrick Brown(e) s.n. (holotype: LINN, microfiche). Acacia virescens DC., Cat. pl. horti monsp. 74. 1813. — TYPE: probably South American, the species was described from a plant grown in the Botanical Garden in Montpellier. The seeds for this Phytologia (Dec 2005) 87(3) 169 plant were obtained from the Botanical Garden in Madrid. (holotype: G-DC, microfiche). Acacia leucacantha Bertero, Spreng., Syst. veg. 3: 144. 1826. — TYPE: JAMAICA. C. G. Bertero s.n. [lectotype, designated by Ebinger et al. (2000): MO]. Acacia seifriziana Leon in Leon and Alain. Contr. Ocas. Mus. Hist. Nat: ‘Colegio: “De. LaSalle?) -92'8:' | 1950.) = TYPE: ‘CUBA. ORIENTE: low forest near Maisi, Jul 1938, Leon and Seifriz 18351 (holotype: HAC). NOTE: We have not been able to examine the type, however, based on the description, it appears referable to Acacia tortuosa, a conclusion also advanced by Bassler (1998). Mimosa salinarum Rohr ex Benth., Trans. Linn. Soc. London 30: 501. 1875.— TYPE: none cited. VACHELLIA VERNICOSA (Britton & Rose) Seigler & Ebinger, comb. nov. Basionym: Acaciopsis vernicosa Britton & Rose, N. Amer. Fl. 23: 96. 1928. Acacia vernicosa Standl. , Cont. U. S. Natl. Herb. 20: 187. 1919 (nom. Illeg.). Acacia constricta Benth. var. vernicosa L. Benson, Amer. J. Bot. 30: 238. 1943. Acacia neovernicosa Isely, Sida 3:380. 1969. (based on the same type). — TYPE: MEXICO. CHIHUAHUA: in the vicinity of Santa Rosalia, alt. 1200 m, 13-15 Jun 1908, E. J. Palmer 385 (holotype: US; isotypes: GH, MO, NY). NOTE: Not Acacia vernicosa Fitzgerald (1904), an Australian species with phyllodes, presently considered a synonym of Acacia incrassata Hook. (Maslin 2001c). VACHELLIA VILLAREGALIS (McVaugh) Seigler & Ebinger, comb. nov. Basionym: Acacia villaregalis McVaugh, Flora Novo-Galiciana 5: 141. 1983. - TYPE: MEXICO. JALISCO: Cima Cerro de Tequila, bosque de Quercus spp. y Pinus spp.., alt. 2850 m, 17 Mar 1974, L. M. Villarreal 6171 (holotype: MICH). 170 Phytologia (Dec 2005) 87(3) VACHELLIA ZAPATENSIS (Urb. & Ekman) Seigler & Ebinger, comb. nov. Basionym: Acacia zapatensis Urb. & Ekman, Ark. Bot.. 22a(8): 30. 1928.— TYPE: CUBA. MATANZAS: Zapata Peninsula, in shrub forest towards Bahia de Cochinos, in open places on limestone, 31 Jan 1924, E. L. Ekman 18358 (holotype: B, destroyed; isotype: NY). EXCLUDED NAMES The following species have been considered to belong to former subgenus Acacia by some authors: Acacia amentacea DC., Prod. 2: 455. 1825., based on a Sessé and Mocifo drawing that is ambiguous (Lee et al. 1989). No type specimen has been found and the quality of the drawing does not permit conclusive identification of the plant depicted. Acacia haematomma Bertero in DC., Prod. 2: 456. 1825. Examination of the type specimen indicates that this is a Calliandra species (Britton & Rose, 1928, Barneby, 1998). Acacia peruviana Willd., Enum. pl. 1056. 1809. The type specimen at P is not an Acacia as it has a single pair of pinnae and lacks stipular spines and prickles. The following names appear on specimens, but were never validly published, to our knowledge: Acacia /eoni Marie-Victorin. -- Name appearing on the label of type specimens of Acacia bucheri. Vacheliopsis cubana -- Name on the label of the isotype of Acacia cupeyensis (M.Curbelo 512) attributed to Britton & Wilson. Phytologia (Dec 2005) 87(3) 171 ACKNOWLEDGEMENTS The authors thank several colleagues for advice concerning questions of nomenclature and general taxonomic advice. Among these are: J. Lee Crane, Bruce R. Maslin, Mario Sousa, Paul Wilson, Gordon Tucker, and James Zarucchi. The views and conclusions (and errors) in this manuscript are our own and do not necessarily reflect any of their judgements. We wish to acknowledge support by the National Science Foundation (NSF DEB 04-15803) and by the American Philosophical Society. LITERATURE CITED Alvarez, A. C. and P.H. Oliver. 1991. Catalogo de las Colleciones Historicos y de Tipos del Herbario de la Academia de Ciencias de Cuba. Editorial Academia, La Habana, Cuba. Aronson, J. 1992. Evolutionary biology of Acacia caven (Leguminosae, Mimosoideae): infraspecific variation in fruit and seed character. Annals Missouri Botical Garden 79: 958-968. Barneby, R.C. 1992. Centennial beans: A miscellany of American Fabales. Brittonia 44: 224-239. Bassler, M. 1998. Flora de la Republica de Cuba. Fasc. 2. Mimosaceae. Koeltz Scientific Books. Koenigstein, Germany. Britton, N. L. and J.N. Rose. 1928. Mimosaceae. North American Flora. 23: 1-194. Chappill, J.A. and B.R. Maslin. 1995. A phylogenetic assessment of the tribe Acacieae. pp. 77-99. In M. D. Crisp and J. J. Doyle (eds.). Advances in Legume Systematics 7: Phylogeny. Royal Botanic Garden, Kew, London. Cialdella, A. M. 1984. El genero Acacia (Leguminosae) en la Argentina. Darwiniana 25: 59-111. 172 Phytologia (Dec 2005) 87(3) Clarke, H.D., S.R. Downie, and D.S. Seigler. 2000. 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