PLANNING FOR WILDLIFE IN THE UNITED STATES PART IX OF THE SUPPLEMENTARY REPORT OF THE LAND PLANNING COMMITTEE TO THE NATIONAL RESOURCES BOARD For aale by the Superintendent of Documants, WashlnSton, D. C. - - Price 10 centa (paper cover) SXJPPLEMENTARY REPORT OF THE LAND PLANNING COMMITTEE • TO THE NATIONAL RESOURCES BOARD Published in eleven separate parts Part I. General Conditions and Tendencies Influencing the Nation's Land Requirements Part II. Agricultural Exports in Relation to Land Policy Part III. Agricultural Land Requirements and Available Resources Part IV. Land Available for Agriculture Through Reclamation Part V. The Problem of Soil Erosion Part VI. Maladjustments in Land Use Part VII. Certain Aspects of Land Problems and Government Land Policies Part VIII. Forest Land Resources, Reqmrements, Problems, and PoUcy Part IX. Planning for Wildhfe in the United States Part X. Indian Land Tenure, Economic Status, and Population Trends Part XI. Recreational Use of Land in the United States From the collection of the i— m Prelinger 'ibrary p fj San Francisco, California 2008 PLANNING FOR WILDLIFE IN THE UNITED STATES PART IX OF THE REPORT ON LAND PLANNING THIS PART WAS PREPARED BV THE BUREAU OF BIOLOGICAL SURVEY. DEPARTMENT OF AGRICULTURE AND THE FOREST SERVICE, DEPARTMENT OF AGRICULTURE FOR THE LAND PLANNING COMMITTEE OF THE NATIONAL RESOURCES BOARD The National Resources Board assumes no responsibility for the views and opinions expressed herein UNITED STATES GOVERNMENT PRINTING OFFICE WASHINGTON : 1935 PREFACE On November 2S, 1934, the National Resources Board submitted to the President, in accordance with an Executive order, its report on National Planning and Public Works in Relation to Natural Resources and Includino; Land Use and Water Resources. Part II of that report was the Report of the Land Planning Committee. In the course of preparing part II of the above report a large volume of basic data and infor- mation was collected which could not then be included. The publication of the present report is for the purpose of making such data and infonnation available to interested persons and organizations. The present land report has been organized into 1 1 parts according to subject matter and the contributmg agencies. These 11 parts are made available as 11 separate publications. Organization and publication on this basis was done because many persons and agencies are interested only in certain parts of the present report, and the necessity of purchasing the whole report in order to obtain the desired part or parts is thereby eliminated. The present land report, when conceived as a whole, does not purport to be a complete work on the subject of land utilization, or of its related problems and pro- posed lines of action ; neither is it designed to be a thor- oughly integrated piece of work. The prunary aim here has been to set forth the facts, analyses, and v,he recommended lines of action as developed by each of the various contributing governmental bureaus, divi- sions, sections, or individuals, on the problems with which each of such agencies or persons is concerned. The points of view are, therefore, those of the con- tributmg agencies or individuals themselves. The Land Planning Committee presents the report as information, but assumes no responsibility for the opinions expressed in it. This report was prepared under the direction of Dr. L. C. Gray, director of the Land Section of the National Resources Board, aided by John B. Bennett, who served as administrative assistant and as secre- tary to the Land Planning Committee. Editing and preparation of the report for publication were under the direction of Mr. H. H. Erdmann, agricultural economist of the Land Section, National Resources Board. Authorship by agencies and individuals is acknowl- edged in their respective contributions. The follow- ing governmental agencies have contributed to the whole report: The Geological Survey, the Division of Grazing Control, the Office of Indian Affairs, the National Park Service, and the Bureau of Reclamation, in the United States Department of the Interior; and the Bureau of Agricultural Engineering, the Biological Survej', the Bureau of Chemistrv' and Soils, the Forest Service, the Soil Conservation Service, the Weather Bureau, the Divisions of Land Economics, of Farm Management and Costs, and of Farm Fmance in the Bureau of Agricultural Economics, and the Land Policy Section, the Production Planning Section, the Import-Export Section, and the Agricultural-Indus- trial Relations Section of the Division of Program Planning of the Agricidtural Adjustment Administra- tion in the United States Department of Agriculture. Credit also is due to the State agricultural expermicnt stations and extension services, State planning boards, commissions, and other State organizations and indi- viduals for aid in preparation of several sections of the report. Land Planning Committee M. L. Wilson, Chairman. Oscar Chapman. W. G. Mendenhall. H. H. Bennett. mordecai ezekiel. Jacob Baker. Charles W. Eliot, 2d. L. C. Gray, Director. LAND REQUIREMENTS AND POLICIES WITH RESPECT TO WILDLIFE Contents Page Preface iii Section I. Requirements for Wildlife Areas 1 Economic importance of wildlife — Social and recreational importance — Need of wildlife management — Possibilities and limitations in associating wildlife with other uses of land — Public areas employed as special- ized refuges — Public areas designated as refuges but devoted to other major uses — Public areas needed for specialized refuges — -Public areas in other uses that should be designated as wildlife refuges — Private lands employed as a habitat for wildlife. Section II. Policies With Respect to Wildlife 12 Areas now in public ownership devoted to wildlife — Programs of acquisition; suggestions for modifica- tion— Integration of Federal and State policies — Encouraging game production on private lands. Section III. Wildlife Management in National Forests 15 Encouraging game management on public land in other use — Economic and social values of wildlife areas — Wildlife and other land uses — Areas suitable only for refuges — Areas specially adapted to wildlife — • Areas available for wildlife — Needed additions to forests — Privately owned land as a hibitat for wildlife. SECTION I REQUIREMENTS FOR WILDLIFE AREAS* Economic Importance of Wildlife Wildlife is as essentially a product of land (or land and water) as is timber, agricultural crops, or domestic livestock. A multitude of kinds, bearing importantly upon human welfare, are especially adapted to various kinds of habitat, from dense forest and open range to cultivated fields, water, and marsh. It follows, there- fore, that wildlife cannot be ignored in any compre- hensive, well-considered plan for land utilization. Mammals, birds, fishes, and other wild vertebrates con- stitute a major national resource that is becoming more generally appreciated, as evidenced by increasing interest in every phase of nature study. Many of the various forms, fur bearers and fishes especially, are of direct economic value. An example is the income derived from the sale of hunting, trapping, and fishing licenses; the furs of course have a great commercial value also, and in some cases the meat supphes a highly prized form of food. That the meat and fur value of wild mammals and birds is considerable is indicated by an estimated total for 1 year of more than $190,000,000 for the whole country. Capitahzed on a low-percentage basis the total value of wildlife would far exceed $1,000,000,000. Other economic values are receipts from the sale of hunting and fishing equipment, including guns, ammunition, fishing tackle, and clothing, expenditures of sportsmen for transporta- tion, board, hire of guides, and for hunting and fishing privileges on private lands. To the foregoing may be added the output of the fur trade, including the fur manufacturing industry, the annual expenditures for the maintenance of the numerous and widely distributed hunting and fishing clubs; also the general expenses of tourists and others attracted primarily by an abundance of wildlife. It is these various wildlife values that go to make up the estimated grand total of at least $1,000,- 000,000 annually. Last but not least should be mentioned the incal- culable value of wildlife, chiefly birds, as destroyers of the insects that prey on agricultural crops and forest trees. Social and Recreational Importance The social values also, including the recreational and educational advantages arising from an abundance of wildlife in general, are more intangible and therefore more difficult to appraise than the economic values, but thev arc none the less real. * Contributed by the Bureau of Biological Survey. In Europe from time immemorial hunting has been a sport restricted mainly to the nobility or the wealthy classes, who usually assume direct owTiership of all game ranging on their large estates. In America, on the other hand, the traditional concept is that owner- ship of most game, whether on privately owned or public land, is vested in the State or the Federal Gov- ernment, and that hunting, subject to restrictive laws and regulations, including those in regard to trespass, is for all who wish to indulge in it. The relative free- dom of the American system is due to the fact that for the early settlers the pursuit and killing of game and fur bearers was a vital necessity in providing meat for food and furs for clothing. The skilled hunter became an expert marksman who defended his home and gave an excellent account of himself in early military cam- paigns. The role of the hunter, important first in gaining a livelihood, led naturallj^ through successive generations to the development of a love of the chase as sport. The joy of the chase provides wholesome recreation of a kind that can be obtained in no other way, and one that perhaps only a hunter can fully appreciate. The many difi'erent kinds of hunting for large and small game and the several kinds of fishing afford the variety adapted to the need and inclination of the various classes of our growing population. The pursuit of large game, especially, which often requires persistent effort as well as skill, tends to bring out quaUties contributing to the virility of the race. A considerable number to whom the taking of animal life is repugnant, satisfy a natural urge for contact wath primitive nature by stalking wildlife to obtain photographs at short range, or to make detailed studies of the life habits of the several species. A report of the Senate Committee on Conservation of Wildhfe Resources (S. Kept. 1329, 71st Cong.), estimates that there was during the decade ending in 1930 a 400-percent increase in the number of people who enjoy the pastimes of hunting and fishing. It estimates the number of licensed hunters in the United States in 1929 at 7,000,000 and the total of all hunters and fishermen at probably 13,000,000. The current trend toward shorter hours and fewer working days should add tremendoush- to the number of men who seek such out-door recreation. Hunting and fishing as a sport attract men to the forest and mountains, the lakes, and the streams, where for a brief period they may escape the artificial life to which by modern conditions most are confined. 1 Land Planning Report From such an outing they return to their usual duties improved mentally and physically, with a broadened outlook and a quickened appreciation of natural re- sources that should contribute immeasurably to the wealth, comfort, and well-being of the people. Need of Wildlife Management Conservation Alone Not the Solution: Recognition of wildlife as a national asset of major importance brings into relief the problem of management. Much has been written on the conservation of wildlife by authors representing many shades of opinion, but compara- tively little indicating a broad, clear concept of its corollary, wildlife management. The emphasis on conservation has doubtless been due to the necessity of building up a militant opposition to destructive forces leading toward the extermination of so many species. In many directions this danger is not yet past, but conservation should be recognized in its true rela- tion as a means to an end and not an end in itself. The overabundance of game in one region may present a more pressing and difficult problem than would its absence or scarcity in another. Game management consists largely in the regulation of numbers in accord- ance with food supply and cover, with due regard for other local interests. At the time of the discover}^ of North America, its •\vildlife, probably unequaled elsewhere in diversity' and abundance, ranged in everj' kind of habitat throughout the length and breadth of the continent. Great herds of grass-feeding animals — buflFalo, ante- lope, elk, and mountain sheep — occupied the open plains or high mountains of the West. Moose and caribou had an extensive range in the northern forests, and bears were numerous wherever conditions were suitable. Fur bearers in great variety, including the beaver, otter, wolverine, marten, mink, fox, raccoon, and muskrat, were widely distributed and soon became the basis of an extensive and profitable pioneer indus- try. Upland game birds, as wild turkeys, passenger pigeons in amazing numbers, and the various kinds of grouse and prairie chickens, occupied great areas adapted to their divergent needs. Waterfowl in count- less millions, especially ducks, geese, and swans, nested over an enormous territory but particularly in the vast prairie regions of the Middle West, the Northwest, and areas extending into Canada and Alaska ; and in concen- trated niA-riads migrated southward to winter. Many writers attest to their almost incredible abundance. For example, Grinnell records that in Eastern and Hogg Bay, Chesapeake Bay, one might see redlieads rafted in bodies miles in extent, probably not less than 50,000 ducks in a mass.' Incidentally the redhead is new so reduced in nuinbei*s that only special protection will prevent its extermination. Geese also were in such ' Grinnell, G. B. American Duck Shooting. New York, 1901. pp. 485-86. abundance that, especially in California, they became destructive to winter wheat on a large scale. Previous to the arrival of the settlers, the scattered Indian population, hunting mainly with bows and arrows, and without effective tools, was negligible in its effect upon the game. Predatory animals — wolves, mountain lions, bobcats, and, in the West, coyotes — took their toll, but there was ample food for all. Essentials of WihUife Management: A re\'iew of the general field, and contrast with past abimdance, shows that wildlife has nearly every^vhere been sadly neg- lected as a national resoiu-ce or grossly mismanaged. The essentials of wildlife management are relatively simple but exceedingly difficult to put into effect, owing to public apathy, sheer ignorance or misunder- standing, selfishness, misguided sentiment, and politics. First of all, only a well-informed public opinion can overcome the inertia and prejudice that tend to paralyze constructive effort. The management of game, and of wildlife as a whole, calls for professional skill of the highest order. The wildlife administrator should com- bine broad technical knowledge and the instincts of a natiu-aUst with capacity for dealing with hard, factual realities. Such a combination of qualities is not easily obtained, and it is rare where game officials are ap- pointed merely as a reward for political party services. Occasionally a political appointee will develop some capacity as a game manager, but by the time he has become familiar with the intricacies of the work his services are apt to be terminated. His political suc- cessor may be sincere, but untrained, and the splendid game assets suffer for lack of the attention that can be given only by those who would make wildlife manage- ment a career. In order to function properly, game commissions should be composed of well-informed indi- viduals, free from political pressure, and must have authority to deal promptly and effectivel}' with rapidly changing conditions as the\ arise. The adequate management of big game on national forests ^ is urgentlj" needed and presents especial diffi- culty owing to conflicting views with regard to juris- diction. Game is a product of the forest, and too abundant game on national forests may be destructive to forest reproduction and may seriously interfere with the proper regulation there of the grazing of domestic stock. Proper game management calls for the regula- tion of niunbers, ordinarily by himting under a limited license plan, the game supply to be maintained with due regard to range-carrying capacity and other local interests. The game should be fostered and made to yield an annual crop that will be harvested in an orderly way adapted to local conditions. Some States may have what is regarded as compe- tent game administration, but it is obviously impossible 3 See also contribution by the Forest Service on wildlife management on national forests. Sec. Ill of part IX. Wildlife in the United States for them, subject as they are to changing policies and political expediency, to equal the efficiency of the wild- life service that can be extended to national forests by the Federal Government under plans coordinating all forest uses. Possibilities and Limitations in Associating Wildlife with Other Uses of Land Wildlife in a multiplicity of forms has a more or less direct bearing upon land uses nearly everywhere. Certain kinds may be highly beneficial, others neutral, or some maj' seriously interfere with economic utiliza- tion of lands. Crows, for example, are beneficial in consuming grubs or cutworms that injure agricultural crops, but they destroy many eggs of wild ducks, the breeding of which constitutes a highly important form of marshland utilization. And these differing crow activities may be carried on over closely adjoining lands. Competitors with Wildlife for Land: With the appear- ance of the settlers, the clearing of the forests for farms and town sites, the occupation of grasslands for agri- cultural purposes, or the grazing of domestic stock, the general aspect of much of the former domain of wildlife was transformed. As the tide of civilization and gen- eral land occupation moved westward across the con- tinent the greater part of the plains animals either were wiped out or resorted to the forests, which are the great reservoirs of so much of our remaining upland game. Though deplorable, the displacement of much of the game was inevitable. The final passing of the buft'alo on open ranges could not have been prevented, for great herds of buft'alo have no place, as game, under modern conditions. Inevitable also was the passing of most of the antelope, originallj' more numerous perhaps than the buft'alo. The intensive settlement of the vast prairie areas of the Middle West and north into Canada, and the devel- opment of one of the greatest grain-producing regions in the world, led directly to the virtual elimination of large sections of the best breeding ranges of our most important waterfowl. The surface absorption of mois- ture by the cultivation of land, and the deliberate drain- age of so many of the originallj' innumerable ponds, lakes, and marshes, have wiped out most of the local waterfowl breeding grounds. Another serious result has been the lowering of underground water levels, shown in the failure of wells, the drying up of springs that should feed streams, and the general desiccation of the country. Meanwhile many of the lands drained have proved to be of little agricultural value, and during periods of drought wind erosion has strongly tended to complete the picture of desolation. Restoration Measures Attempted: Efforts are now being made to restore some of the 84,000,000 acres that have been drained in the country as a whole; and 144090 — 36 2 yet drainage engineers may in many localities be con- tinuing the destructive policy of the past. All drainage projects of the Reclamation Service should be scrutinized with a view to determining their effect upon the native wildlife and other water uses and values, and these should be balanced against their prospective value for the purposes already planned. Pressure on wildlife by human occupation of the land, therefore, together with almost unrestricted killing by hunters, inroads by predatory animals, and other inci- dental factors, has led to the disappearance of game of many kinds or to its reduction to comparatively small numbers over most of its former range. In the Eastern States the larger predatory animals virtually disap- peared many years ago along with the larger game, but the destruction of game by the mountain lions, wolves, coyotes, and bobcats that still remain in the Western States continues on a scale that is not generally realized. The greatest predatorj"^ animal after all, however, is man. Control must be exercised over the too indiscrim- inate use of modern firearms by the increasing millions of hunters, if a sufficient breeding stock of game is to be maintained. This does not mean that hunting under proper conditions should be discouraged, but rather that its implications should be realized and that it should be carried on and regulated as a game-crop harvesting operation. Although the general outlook for forms of wildlife hunted as game may seem discouraging, conservation progress is being made and much imiy yet be done through the properly coordinated use of land, not only to save the remnants, but greatly to increase numbers and restore game to vast sections from which it has been eliminated. To deal intelligenth' with the problem requires an appreciation of the requirements of the game and of all the factors that maj' result in decreases or increases in numbers of wildlife forms. With these requirements clearly in mind wildlife values should be measured against land values for other uses everywhere. Forest Wildlife: The fact that the forests are the great reservoirs of most of our upland game leads logically to a consideration of the relation of forestry to game con- servation with a view to the best use of forest lands.* Since the maximmn production of timber tends to con- flict with the maximum production of game, the rela- tive economic and social values and all the factors involved should be carefully weighed in connection with general forest numagement. Forest reproduction is so unsatisfactory in many places, from the forester's stand- point, that planting has to be resorted to. In such proj- ects of forest restoration the probable trend in future comparative values of timber and game should be antici- pated as far as possible. These comparative values, difficult to measure, will vary with manj' local condi- tions. The rapid substitution of other more durable • See also contribution by the Forest Service on wildlife managemeDt on national forests. (Sec. HI of part IX.) Land Planning Report materials for the wood that was formerly used in con- struction of all kinds, and the slow growth of mer- chantable timber suggests that in planting operations the potential values of wildlife as well as of timber values should be given consideration. Large game animals, as well as beavers, porcupines, rabbits, and other smaller forms, are apt to become too abundant and destructive to the forest. In places the competition between unregulated game and domestic stock for forage has become acute and the range per- mancntlj- impaired. Serious problems thus tend to arise. Wilderness Preservation and Restoration: Conditions even appro.ximating the primeval wilderness can hardly be maintained anywhere unless the areas available are extensive. Wilderness, or natural, areas typical of the principal kinds of land surface and landscape aspect, includuig humid and arid, forested and unforested, should be preserved in the various parts of the country for values that cannot readily be measured in dollars and cents. The fauna is dependent upon the flora, and typical samples of both should be perpetuated. Such a plan contemplates, as examples, the setting aside for posteritj" of suitable tracts of such divergent character as the great forest of the Olympic Peninsula of Washington, the Everglades of Florida, and the giant cactus desert of southern Arizona. The areas should to well-rounded units and include, as far as possible, both summer and winter range for game, and should function as wildhfe preserves of the highest type. In such areas modifying human influences should be limited to such emergency measures as fire and flood control and the reduction of excessive num- bers of large game and predatory animals. Suitably distributed wilderness or natural areas should be maintained in the national forests; several in the larger national parks might be so designated, and others might be carved from the unreserved public domain. It is probable that few typical areas of the long-grass and short-grass prairies that played so prominent a part in our national development remain unmodified, but some might be restored to an approximation of their original condition. As such, and supporting suitable wildlife, they would be of surpassing interest to posterity. Some suitable prau'ie areas would probably have to be acquired by purchase. The present limited system of refuges. Federal and State, designed for the perpetuation of particular species, as the buffalo, should be extended to provide more amply for other animals, including the antelope, valley elk of Cahfornia, the peccary, and others. Human capacity to transform the land surface and to render it imfit for habitation by wildhfe, especially game, has been clearly demonstrated. That a reversal of the destructive process can bring about an amazing restoration of game has also been amply attested. It foUows then, that in manipulation of the environment, game should be given due consideration as a natural resource in all plans for land utilization. Game must be subject to appropriate control measures, but it will often be found highl.y profitable to encourage its pro- duction along with other land uses. Relations of Big Game, Predators, and Livestock: Large game animals, as deer, antelope, and elk, often compete directly with domestic stock for forage. To maintain the food-producing capacity of a given area may require a reduction in the numbers of either or both. Adjustment of the conflicting interests of game and the grazing of domestic stock, and the relation of both to predatory animals, is a compUcated problem on the national forest and the public domain. Or- ganized predatory animal-control operations are carried on by the Federal Government, largely on national forest areas and the pubhc domain but also on State and private holdings in cooperation with the States. The direct objective is the protection of the domestic stock for which grazing fees on the national forest are collected by the Forest Service. The work is financed jointly by the Federal Government, which through the Biological Survey is charged with the direction; and the States or stock associations, which furnish the larger share of the funds required. This seems an equitable arrangement, as the States are heavily interested in the stock industry both on and oft' the forest, while the Government is the principal landowner. But the same national forests are the principal reservoirs of our larger game, especially deer, and a heavy toU is nor- mally taken every year by mountain Uons, coyotes, and bobcats. \Vhile mountain hons are the more notorious deer slayers, each year kiUing great numbers of deer in sections where abundant, the inroads of coyotes, especially on fawns and the smaller game species, are usually more serious, owing to the greater number of these wild members of the dog family. Predatory animal control, undertaken primarih' in the interest of domestic stock, now incidentaUy extends similar protection to game, which only a few years ago became threatened with the actual danger of local extinc- tion. On many of the domestic stock ranges of the West, notably in the national forests of central and south- eastern Arizona, deer, antelope, and elk have become overabundant and threaten the forage supply for both stock and game. This maladjustment is due partly to the creation of game refuges that are far too large, and partly to lack of adequate provision for the harvesting of the game crop. In such places, where hunters are unable to utilize the game, projects of predatory animal repression should be pressed only as necessary to prevent serious injury to local economic interests. Buffalo and Other Big Game: The herds of wild and semidomesticated buffalo that range in an isolated part of Yellowstone National Park and the larger herds in Canada, together with numerous groups under fence, Wildlife in the United States assure the perpetuation of a species once threatened with extermination. Deer, moose, and black bears now tend to increase where accorded protection, especially in some of the national parks. A few cari- bou still persist in continental United States in an area closed to hunting in northern Minnesota. Mountain Sheep, or Bighorns: Formerly widel}- dis- persed in the more rugged sections of the West, mountain sheep, or bighorns, still occur in places in the higher mountains in the national forests and national parks, and in the desert ranges of the Southwest, usualh^ in small bands. As a rule these show Uttle or no increase, although under the laws of most States no killing is permitted. In many sections local extinction was due to unrestricted huntmg, especially for meat to supply mining camps. There seems to be good reason to believe that scabies and perhaps other diseases introduced by domestic sheep have killed manj' mountain sheep; and predatory animals have taken their toll. In some parts of the public domain little utilized by man the causes that led to the elimination of mountain sheep appear no longer to be operative. Mines have been worked out and abandoned. Domes- tic sheep are now dipped and kept more generally free from scabies, and under proper control may be ex- cluded from areas especiallj' adapted to their wild relatives. In some of the rugged, desert mountains, especially of California, Nevada, Arizona, and New Mexico, there is sufficient forage, but ranges cannot be used bj' domestic stock of any kind except possibly at certain seasons, owing to lack of water, which, however, could be made available at moderate cost by impound- ing the run-off at favorable sites. Many such areas, now scarcely utilized for any purpose, could undoubtedlj' be stocked and developed as excellent mountain sheep ranges. Owing to the habit of twinning, mountain sheep have a high potential rate of increase, a rate actually attainable under favorable conditions, as shown for a time on the National Bison Range, in Montana. A system of well-administered Federal mountain sheep refuges would encourage State conser- vation efforts and should in a few years afford a surplus for stocldng many ranges on which limited himting of one of the finest game animals in the world could be permitted. Antelope: The antelope, regarded by manj- as the most beautiful and characteristically American of our game mammals, have decreased rapidly in most places during recent years. Though they still occur in parts of most of the Western States, many of the smaller groups, unfortunatelj', are in sections where unfavor- able local conditions make their extinction practically certain. Antelope have held on with remarkable tenacity, but they are not likelj' to survive except on areas where they are accorded special protection. They are reasonably safe in the national forests and national parks, but much of their natural range is on the public domain or on lands to which adequate pro- tection or management has not extended. In some sections, however, as in parts of Arizona, California, Nevada, Oregon, and Wyoming, protective measures have favored substantial increases. On the Coconino National Forest and adjoining territory in Arizona, where extinction seemed imminent a few years ago, antelope have now increased to thousands. In the competition between antelope and domestic stock, the low-growing food plants are cropped with killing effect, and the normal forage supply is so diminished that the antelope are forced to browse on junipers and other unpalatable trees as high as thej- can reach, leaving them completely defoliated to a sharp line that is always indicative of seriously destructive overutiliza- tion on stock or game ranges. Such forage curtail- ment, increasing general surface erosion, results in rapid, permanent deterioration of the range and starva- tion on a large scale. Here the mounting numbers of the antelope, on areas closed to hunting, hke those of other game in many parts of the West, have been coincident with the control of predator}^ animals, mainly coyotes, instituted primarilj' in the interest of domestic stock production. The general public, and even many who regard themselves as conservationists, often uninformed and accustomed to think of the antelope as a vanishing species, may oppose that feature of game manage- ment that involves the IdiUng of surplus animals. This case is an example, of which manj- could be cited, of the necessity for clearer general understanding of the requirements of game and the urgent need for its consistent management in well-considered plans for the multiple use of land. W}-oming contains far more surviving antelope than any other State, and, as in some other places, mounting numbers on the public domain constitute local problems pressing for solution. Federal refuges established primarilj- for antelope in suitable sections of the public domain in several Western States would also extend needed protection to mule deer and sage hens. On other parts of the public domain the grazing of domestic stock should be controlled in accordance with the reasonable needs of game as well as with a view to watershed protection and the prevention of soil erosion. Deer: Owing to large numbers and wide distribution the deer are by far the most important of the big game of the United States. They constitute a national resource capable of great development. Although now extinct over large areas formerly occupied, four species of deer still divide territor}' or overlap in geographic Land Planning Report range. In order of importance these are, first, the white-tailed, or Virginia, deer of the Eastern and Northern States, extending west and invading the general territory of the mide deer in the northern Rocky Mountain region. Second in importance is the mule deer, commonly called the blacktail, of the more arid West, from the Rocky Mountains west to the Cascades, reaching the Pacific coast in southern Cali- fornia and extending southward into Mexico. Third, the true blacktail of the Pacific coast region, the close relative of the mule deer, occurs from California north to southeastern Alaska. And, fourth, the Arizona whitetail of the mountains of southern Arizona, south- ern New Mexico, and western Texas, has a main range extending far south along the Sierra Madre of Mexico. As these four members of the deer family are quite distinct and also differ considerably in the character of terrain occupied, their relations to one another should be clearly understood and considered in connection with plans for land utilization. Elk-: The elk, one of the largest and most majestic members of the deer family, had a range measured originally by the width of the continent, from Penn- sylvania to California. Unfortunately so large an animal can be destructive to agricultural interests, and like the buffalo, its displacement in densely settled com- munities was inevitable. Thousands have remained, however, in the wilder parts of the Rocky Mountain region, chiefly in and around Yellowstone National Park, and in the mountains nearer the Pacific coast in Oregon and Washington. These elk are of special interest, as they constitute the only really large herds of big game remaining in the United States, exclusive of Alaska, but while comparatively numerous they are mere remnants of the former great herds. In the West, the elk were accustomed to summer largely in the high mountains and to migrate to winter- ing areas on the lower mountain slopes and the sur- rounding plains, where there was less snow and abun- dant food. Occupation of the grasslands at the lower levels for agricultural purposes or for grazing stock forced many of the elk to winter in the higher and more inaccessible mountains, where they suffer pitifully, many dying from starvation and disease in hard winters. The result has been the elk problem about which so much has been written. The preservation of the elk is a land utilization problem of major importance. Elk of a northern group are fed in winter in Yellowstone National Park by the National Park Service. Those of a more southern group winter in the Snake River drainage, in Jackson Hole, Wyo. Here the Winter Elk refuge, embracing 4,500 acres of meadow land, maintained by the Bureau of Biological Survey, together with supplemental feed provided by the Wyoming Game Commission, has repeatedly prevented a final disaster. Several thou- sand elk frequently congregate on the feeding ground, where they crowd close about the wagons from which the hay is distributed, and the spectacle thus presented is one long to be remembered by the fortunate visitor to the place. But such large concentrations are unde- sirable, and the feeding facilities are inadequate. In a solution of the problem of winter range and forage for these elk, about 12,000 acres of additional land are urgently needed. Authorization for the acquisition of this land has now been obtained, and actual acquisition is under way. A bill providing for the acquisition of the lands selected has been before Congress for several years. Elk are hardy animals, adapted to a fairly wide range of conditions. Owing to their large size they are more destructive to agricultural interests than the smaller deer, and for this reason they should not be introduced in the vicinity of farming communities. In some of the wilder and more mountainous parts of the West, however, the range of the elk might be extended. Under proper management, surplus elk should afford not only rare sport but an ample supply of excellent meat for the fortunate hunter. As hunting is prohibited in national parks, the provision of open hunting grounds for elk should be given careful con- sideration in connection with any plans for park extension. Management oj Beavers and Other Fur Animals: The original range of the beaver included practically the entire country. This fine fur bearer is at home along streams in suitable places from the lower Rio Grande and the lower Colorado River on the Mexican boundary to near timber line on the highest mountains. It is easily trapped and was eliminated or reduced to small numbers in many localities, where it should be restored and colonies increased as a major asset. Like many other kinds of wdldlife, beavers require expert manage- ment. The location of their dams may in some places seriously injure other important human interests. Roads or fields may be flooded, or valuable timber killed. Fortunately beavers can readily be trapped and moved alive and uninjured to sites where their activities will do no harm. At low elevations streams desired for trout fishing may become too sluggish and warm owing to the impounding of water by beavers, but cold, swiftly flowing streams at the liigher elevations are more likelj' to be improved for these fish, as the dams tend to equalize the stream flow and prevent erosion. In many forested sections food is abundant and general conditions are still suitable for beavers. Under proper management the value of the annual fur crop from this source alone could be made to reach high figures. Other fur bearers, as the muskrat, marten, mink, and raccoon, should be more adequately managed, with a view to developing neglected wildlife assets. Forest Game Restoration: Along with the clearing of forests incident to settlement in the eastern United Wildlife in the United States States, the game, notablj' the deer that played an important part in pioneer development, disappeared or were greatly reduced in numbers as they were hunted and killed without stint. With the decline of the deer, the larger predatorj- animals that prej-ed upon them — mountain lions and wolves — became extinct nearl}' throughout the East. Much of the forested land, however, was too moun- tainous or rocky for agricultural purposes. The large, mature timber was gradually cut for use in construc- tion of manj' kinds. The logging operations, taking about all the merchantable timber, were extended successiveh' from area to area nearly throughout the forested regions of the East, continuing over a long period down to the present. In the original forest, the older trees, largely hemlock, white pine, and mature hardwoods, heavily shaded the ground. The result of the shading was a thin stand of small trees, berry- producing shrubs and other vegetation, and a limited supply of tender browse for deer and fruit for bears and other wildlife. The removal of the forest canopj" through lumbering operations, however, brought a great change. The dense new growth springing up afforded far more food and cover for game. The larger predatory animals were eliminated as has been men- tioned. In marginal or submarginal territorj^, such as in many sections of the Ozark Mountain section of southern Missouri and northern Arkansas, and the Appalachian Mountain region in half a dozen States, now virtually barren of game, the forest setting has thus been prepared for the restoration of deer, bears, rabbits, grouse, and wild turkeys on a scale far exceed- ing the game populations of the same areas before the coming of the white man. If such areas were restocked and properly administered, the annual game and fish crop that could be taken would go far to reheve the general poverty so prevalent there at the present time. The phenomenal success of forest-game restoration in cut-over areas in Pennsylvania, New York, Michigan, and other States where large predatory animals are now absent or scarce is indicative of what could be done in similar sections in many other States. Conditions in the West differ materially from those in the East, as the western forests have been cut over only in part, domestic stock grazing is a more important factor to be reckoned with, and large predator}^ animals, especially coyotes, are still numerous. The remarkable increases in game, especially deer and antelope, coinci- dent with predatory animal control undertaken pri- marih- to protect domestic animals, have already- been mentioned. In spite of these differences it is evident that the general principles bearing upon the relation of game to the use of forested lands in the West are the same as in the East. Wildlife Refuge Needs: The importance of establish- ing more refuges, Federal, State, numicipal, and private, as a part of a comprehensive program of wildlife man- agement, is becoming more generally recognized; but these must be suitably located and properly adminis- tered to achieve their highest usefulness. Wildlife refuges of one kind or another are under the jurisdiction of several departments of the Federal Gov- ernment, their varied situations being largely due to expediency, as the control of wildlife tends to run con- currentl.y with that of the land (or water) it inhabits. The national parks under the Interior Department and refuges on the national forests under the Forest Service, Department of Agriculture, have alread}' been men- tioned. Over 100 refuges, mainly for birds, but includ- ing a few big game ranges, under the Bureau of Bio- logical Survey, Department of Agriculture, are far flung in general distribution — from Puerto Rico and the coastal islands of the United States, to the Aleutian Archipelago in Alaska and islands of the Hawaiian group. These Federal refuges are administered with special emphasis on the interests of all the wildlife found on them. The protection of marine mammals, notably the northern fur seal, which breeds on the Pribilof Islands, and the sea otter, is intrusted to the Bureau of Fisheries, Department of Commerce. Specialized Refuge Needs for Upland Game: The ideal upland game refuge is a protected area that will favor the rapid increase of game within its borders and from which the surplus may readily spread to suitable sur- rounding territory that may be open to hunting. It need not necessarily be very large, but alone or in con- junction with neighboring refuge units should provide j^ear-long range, ample forage, water, and cover for resident game. This is the type of refuge that has succeeded so admirably in building up the game supply in the State forests of Pennsylvania and other Eastern States, and on Federal and State refuges on the national forests, mainly in the West. ^Mierever game thrives, a surplus representing a game crop may reasonably be expected. It can be removed by hunters, consumed by predatory animals, or allowed to die of starvation or disease. Game management, therefore, calls not only for adequate protection but the prompt and well- planned disposal of all surplus. The national parks serve as great game reservations in which no shooting by sportsmen is allowed. Game Refuge Needs on Foreats: Additional refuges of suitable size and distribution should be created for forest game in the forested areas, but are even more urgently needed for restoring mountain sheep, antelope, mule deer, peccaries, sage hens, and other denizens of the more arid parts of the public domain. The un- wieldj' surplus of deer, antelope, and elk that has already resulted from overprotection in certain places should not be taken to mean that the protective prin- ciple is wrong, or that additional refuges are not de- sirable, but should be considered as examples of game mismanagement. 8 Land Planning Report Waterfowl Refuge Area N'eeds: While a sufficient breeding stock of most kinds of upland game, and even of some of the more important fur bearers, seems within relatively easy reach, the plight of out migratory water- fowl is far more critical. Human encroacliment, the primary cause of the great reduction of waterfowl — ducks, geese, and swans — that formerly nested in the Prairie States of the Middle West, and the Prairie Provinces of Canada, and migrated southward in uncounted millions, has already been described. To this primary cause should be added overshooting by an ever-increasing army of hunters. The combined unfavorable influences have led to the diminution of waterfowl at a rapidly accelerating rate, and, unless they can be modified, they must lead to the early extermination of one species after another. In order to check the depletion of breeding stocks of waterfowl common to the United States and Canada, especially the ducks and geese. Federal regulations are annually promulgated under the Migratory Bird Treaty with Great Britain. State laws and regulations tend to con- form with those of the Federal Government; but restrictive regulations alone will not suffice to save the birds. An appraisal of the relative value of many prairie areas before and after drainage brings into relief the vital importance of putting into effect a wildlife restoration program by the Federal and State Govern- ments in cooperation with all other agencies interested. A system of Federal refuge areas, covering especially ihe Middle West and the great migration fiyway through the Mississippi Valley, is planned to take the form of inviolate refuges, on which the birds can breed unmolested, and on which also they will have feeding and resting grounds. Wildlife Management on Privately Oumed Lands: Meanwhile the cooperation of private landowners should be enhsted, with a view to restricting as far as practicable the grazing of domestic stock and other destructive practices about prairie ponds and in marsh areas of critical importance to breeding waterfowl. On Farms: The relation of small game, as rabbits and squirrels, the fur bearers, and the various species of upland game birds, to private holdings, and especially farmland utilization, is of major importance owing to the vast extent of the land involved. The cultivation of land brings radical changes in environmental condi- tions, and these may be either beneficial or harmful, depending on the varying needs of the different classes of wildlife. Farm operations may provide an abun- dance of food and shelter at certain seasons and leave the game and fur bearers bereft of these prime neces- sities at others. The attitude of the farmer toward the hunter has an important bearing in this connection. Farm game, the fur bearers, and the fishes should be regarded as having a potential crop value similar to that of any other product of the soil. Since a farm must be managed for profit, there should be recognition of the fact that the farmer has a proprietary interest in the game attached to liis land, and that he is justly entitled to a monetary return, proportionate to his efforts in its behalf. He should be encouraged to expect a profit tluough the sale of shooting or trapping rights, under regulations fixed by the State. The rela- tion of local game, fur bearers, or fishes to other farm crops of the region should be clearly understood. Rabbits, for example, may be too numerous and injurious to field crops or to horticidture in a given locaUty, while a larger number would be harmless or even highly desirable in another place. The food and cover required for wildhfe may be provided by less intensive cultivation of land, in conformity with current human needs. In the Prairie States many hedgerows and bordering thickets, with a higlily beneficial wind- break value, and incidentally affording game food and shelter, were uprooted and displaced by barbed wire in order to make the land yield a Httle more grain in war time. The result has been increased wind erosion, soil exhaustion, and game elimination. Obviously, and in conformity with the shelter belt idea, such hedgerows should be restored for their beneficial effect in the general use of the land. The strip cultivation of land, employed to prevent wind erosion and as a crop reduction measure, coidd also be made the means of increasing farm game. On Areas Generally: Wildlife, in its multiplicity of forms adapted to every sort of environment, should be accorded its proper place in the use of land everywhere. It should be recognized as a rich endowment, one to be wisely managed and used, and then passed on unim- paired to future generations. Public Areas Employed As Specialized Refuges Administered by the Bureau of Biological Survey: Federal areas devoted exclusively to wUdUfe protection under the administration of the Bureau of Biological Survey, United States Department of Agriculture, include more than 900,000 acres of surveyed lands in the United States, besides unsurveyed islands in Alaska, Hawaii, and Puerto Rico. A Ust is given in table I and acreages by States in table II. Most of these refuges are specifically for the protection of birds, but several big-game preserves, on which birds also are protected, are included. Administered by the Bureau of Fisheries: The Pribilof Islands, Alaska, of 49,000 acres, are administered by the Bureau of Fisheries, Department of Commerce, primarily for the conservation and utiHzation of the fur seals. Wildlife in the United States 9 Table I. — -Specialized wildlife refuges under administration by the Bureau of Biological Survey, showing the year established, the acreage of the surveyed lands, and the chief species for the protection of which each tras established Table I. — Specialized wildlife refuges under administration by the Bureau of Biological Survey, showing the year established, the acreage of the surveyed lands, and the chief species for the protection of which each was established — Continued Year state and designation estab- lished Acres Chief species protected Alabama: Petit Bois Island 1903 956 Laughing gulls, least terns, black skimmers, Louisiana herons, brown Alaska: pelicans. Alaska Railway Musk- 1927 4.160 Muskrats, beavers. rat and Beaver Ref- Aleutian Islands 1913 (') Puffins, auklets. murres. gulls, ducks, geese, ptarmigan, sea otters, red and blue foxes: on Uniraak Island cari- bou, big-brown bears. Bering Sea (St. Mat- 1909 (') Puffins, auklets, kittiwakes, glaucous thew and Hall Is- gulls, sandpipers, snow buntings. lands). Bogoslof 1909 (') Puffins, auklets. murres. gulls, sea Chamisso Island 1912 (') lions. Horned puffins. Pallas murres. Pacific kittiwakes. glaucous gulls. Curry Bird, Game, 1927 8,960 Grouse, ptarmigan, black bears, foxes. and Fish Refuge. lynx, ermines, fishers. Forrester Island 1912 (■) Puffins, auklets. guillemots, murres. gulls, petrels, cormorants. Hazy Island 1912 (') Puffins, auklets, guillemots, murres. gulls, cormorants. Nunivak Island 1929 (') Waterfowl, ptarmigan, reindeer, cari- bou, musk oxen, foxes, minks. 1909 (0 gulls, petrels, cormorants. 1932 1909 8,290 (') Tuxedni Pacific kittiwakes, glaucous gulls, Arkansas: eider ducks. Big Lake 1915 8,937 Ducks. California: Farallon - . . 1908 91 Puffins, auklets, guillemots, murres. Delaware: gulls, cormorants. Killcohook (see also 1934 586 Ducks, geese, muskrats. New Jersey). Florida: 1925 12 Brown pelicans. Caloosahatchee 1920 1929 40 236 Herons. Chinsegut Hill 1932 2,033 birds. 1906 90 Louisiana and little blue herons. 1908 3,321 Key West 1908 2,030 Cormorants, brown pelicans, man-o'- war birds, white ibises, herons. Matanzas.- 1927 267 Black skimmers, clapper rails. Matlacha Pass 1908 1908 1905 10 1 36 Cormorants, brown pelicans, berons. Laughing gulls, terns, skimmers, cor- morants, herons, sandpipers. Pelican Island 1903 15 Brown pelicans. Pine Island 1903 1931 31 36, 862 Brown pelicans, herons. St. Marks Gulls, ducks, geese, sandpipers and Georgia: other shore birds. Blackbeard Island 1926 4.659 Herons, ducks, clapper rails, quail, chachalacas, white-tailed deer, rac- coons, opossums. Savannah River (see 1927 424 Ducks, herons, rails, coots, shore birds. also South Caro- lina). Wolf Island. 1930 538 Waterfowl, shore birds. Hawaii: Hawaiian Islands 1909 W Terns, albatrosses, petrels, shearwaters, boobies, man-o'-war birds, Laysan teal, rails, finches. Johnston Island 1926 (0 Sooty and noddy terns, wedge-tailed shearwaters, petrels, boobies, man- Illinois: o'-war birds. Upper Mississippi 1925 19, 759 Ducks, gee.se, shore birds, upland game River Wildlife and birds, muskrats. minks, beavers. Fish Refuge (see also foxes, raccoons, fishes, mollusks. Iowa. Minnesota, and Wisconsin). Iowa: Upper Mississippi 1925 30, 616 Do. River \\ ildlife and Fish Refuge (see also Illinois, Minne.sota, and Wisconsin). Louisiana: Breton Island . 1904 955 Laughing gulls, royal and Cabot's terns, skimmers, herons, willels. F-ast Timhnlipr 1907 337 Gulls, roval terns, skimmers, brown pelicans, herons, clapper rails. Shell Keys 1907 77 Royal terns, brown pelicans. Tern Islands 1904 1,000 Laughing gulls, royal. Cabot's and Maine: Forster's terns, brown pelicans. 1933 12 Maryland: 1932 8,241 Michigan: birds, muskrats. Huron Islands.. 1905 83 Herring gulls, ducks. Slsklwlt Islands 1905 9 Do. State and designation Minnesota: Mille Lacs Upper Mississippi River Wildlife and Fish Refuge (see also Iowa, Illinois, and Wisconsin). Montana: Benton Lake National Bison Range.. Nebraska: Crescent Lake Niobrara Nevada: .\naho Island Charles Sheldon Railroad Valley New Jersey: Killcohook (see also Delaware). North Carolina: Lake Mattamuskeet... Swanquarter.. North Dakota: Chase Lake... Long Lake... Stump Lake. SuUysHill... Oklahoma: Salt Plains. Wichita Mountains Wildlife Refuge.-' Oregon : Qoat Island Lake Malheur .-. Malheur (Blitzen Val- ley). Three .\rch Rocks Puerto Rico: Culebra Desecbeo Island... South Carolina: Cape Remain Year estab- lished Savannah River also Georgia). Utah: Bear River (see Locomotive Springs. .. Washington: Columbia River Copalis Rock Dungeness Spit Ediz Hook Flattery Rocks Smith Island Quillayute Needles Wisconsin: Gravel Island Green Bay Upper Mississippi River Wildlife and Fish Refuge (see also Iowa. Illinois, and Minnesota). Wyoming: Bamforth Lake Elk Retuge Flat Creek , Button Lake , 1915 1925 1929 1909 1931 1912 1913 1931 1934 1934 1932 1908 1931 1905 1914 1930 1905 1935 1908 1935 1909 1912 1928 1931 1926 1907 1915 1915 1907 1914 1907 1913 1912 1925 1932 1912 1922 1932 Acres 23.288 12.235 18. .521 40. 782 16.681 248 30. 321 135, 184 856 49. 925 15. 493 2.960 8,795 27 994 19. 453 61.500 21 95, 155 (■) (') 2,559 64,216 1.031 83 125 65 117 27 2 64.749 2.163 4.558 Chief species protected 40 ,708 Galls, ducks, geese. Ducks, geese, shore birds, upland game birds, muskrats, minks, beavers, foxes, raccoons, fishes, mollusks. Ducks, geese, shore birds. Ducks, grouse, pheasants, buffalo, deer, mountain sheep, elk. Terns, ducks, geese, coots, sandpipers and other shore birds. Prairie chickens, sharp-tailed grouse quail, hufialo, elk. antelope, beavers Gulls, cormorants, white pelicans. Sage grouse, antelope. Ducks, geese, ibises, avocets. shori birds. Ducks, geese, muskrats. Ducks, geese, swans, shore birds, muskrats. Ducks, geese, brant, swans, sandpipers and other shore birds. Gulls, white pelicans, ducks, shore birds, grouse. Gulls, terns, ducks, coots, and willets, sandpipers and other shore birds. Grebes, gulls, terns, ducks, Wilson's phalaropes. Goldeneye and wood ducks, geese, pheasants, buflalo, elk, white-tailed deer. Franklin's gulls, black terns, waterfowl, bitterns, rails, shore birds. Ducks, geese, wild turkeys, buflalo, elk, deer, antelope, Texas longhorns. California murres. gulls, puffins, geese. Grebes, gulls, terns, cormorants, peli- cans, ducks, gee.se, swans, herons. Grebes, gulls, terns, cormorants, peli- cans, ducks, geese, swans, ibises, herons, sandhill cranes, sage grouse, deer, antelope, beavers. Puffins, guillemots, murres, gulls, fork- tailed petrels, cormorants. Gulls, royal terns, Bahama ducks, herons, coots, ground doves. Terns, boobies, man-o'-war birds, oyster catchers. Ducks, curlews, egrets, herons, pelicans ibises, sanderlings. dowitchers. oyster catchers, sandpipers and other shore birds, sea turtles, diamond -.back terrapin. Ducks, herons, rails, coots, shore birds. Ducks, geese, coots, shore birds, peli- cans, ibises, pheasants, beavers, muskrats. Ducks, coots, curlews, avocets, sand- pipers and other shore birds. Gulls, ducks, geese, blue herons. Puffins, miu-res, glaucous and western gulls, petrels, cormorants. Grebes, loons, gulls, ducks. Pigeon guillemots, California murres, cormorants. Tufted puffins, pigeon guillemots, California murres. Western grebes, pigeon guillemots, California murres, cormorants, ducks. Grebes, auklets, gulls, cormorants. Herring gulls. Do. Ducks, geese, shore birds, upland game birds, muskrats. minks, raccoons. b^vers, (axes, Qsbes, mollusks. Ducks, geese, sandpipers and other shore birds. Ducks, geese, sage grouse, elk (in winter) . Ducks, geese, elk (in winter). Ducks, geese, and sandpipers and other shore birds. 'Unsurveyed areas. ' Jurisdiction transferred from Forest Service, 193S. 10 Land Planning Report Table II. — Acreage by Stales of surveyed lands administered as uildtife refuges by the Bureau of Biological Survey, Department of Agriculture State Acres Remarks 9a6 21,410 8,937 91 686 44,984 5,621 Alaska Figures for Alaska do not include the acreage of a number of unsurveyed islands. Florida Illinois . 19. 759 30.616 2,369 12 8,241 92 23,289 30, 756 57.463 165,753 856 65. 418 12. 776 80, 953 159,909 Maryland - . North Dakota Including Wichita Mountains Wild- life Refuge. Unsurveved islands. South Carolina . 56, 620 7.000 65, 247 630 64,778 8.469 936, 591 Utah Wyoming - Total surveyed acreage... Public Areas Designated as Refuges but Devoted to Other Major Uses Administered by the Biological Survey on Reclamation Reservoirs: In 11 States the Biological Survey adminis- ters wildlife refuges, mainly for birds, on areas the pri- mary use of most of which is as reservoirs under projects developed by the Bureau of Reclamation of the Interior Department (tables III and IV). As there is usually wide fluctuation of water levels, however, their value for wildlife is limited. Administered by Other Federal Agencies: Publicly owned areas administered by several Federal agencies have been devoted primarily to various major uses and designated as wildlife refuges, as shown in table V. State Game Refuges: The list in table VI wiU indicate the approximate acreage nominally in State game refuges. Some areas are State owned or leased, and Table III.- — Acreage by States of Reclartwtion Service reservoir areas administered as bird refuges by the Biological Survey State Acres Remarks Arizona 333, 807 88,763 25,540 67,870 5,107 360,513 91.908 94, 249 13,680 14,080 1,120 34,949 Including Boulder Canyon project. Idaho Nebraska . New Mexico Oregon. - Utah Wyoming _ .. Total acreage 1,131,586 others are Federal lands, cliiefly in national forests, on which the States exercise jurisdiction over the game. Owing to frequent changes in status the figures are incomplete and subject to constant correction. Public Areas Needed for Specialized Refuges Additional Federal lands needed for the purpose of specialized wildlife refuges fall into two main divisions, as follows: For Upland Game: Parts of the unreserved public domain, aggregating 21,000,000 acres needed for up- land game, especially such big game mammals as mountain sheep, antelope, and mule deer. For Migratory Birds: Marsh and water areas, aggre- gating 17,000,000 acres, needed for the restoration and conservation of waterfowl, mainly ducks, geese, and swans, in accordance with a national plan, in further- ance of the Migratory Bird Treaty protecting birds that pass the year between Canada and the United States. Table IV. — Wildlife refuges administered by the Bureau of Bio- logical Survey on areas primarily devoted to other uses — chiefly on reservoirs for irrigation projects of the Bureau of Reclamation State and designation Year estab- lished .icres Chief species protected Arizona; Boulder Canyon (see also Nevada) Salt River . _ - 1933 1909 1911 1908 1930 1928 1909 1909 1929 1921 1921 1912 1909 1916 1933 1931 1909 1909 1909 1908 1927 1928 1909 1909 1909 1928 312,687 21, 120 33, 840 45, 560 9,363 12.300 13. 240 56,954 2,022 2.540 3.155 3,199 5,107 346, 443 14, 070 18, 680 73,228 2,677 81, 619 1,813 8,140 13,680 14,080 1,120 34,949 Waterfowl, mammals. Cormorants, white pelicans, ducks, California; Clear Lake geese. Gulls, cormorants, white pelicans. Klamath Lake (see also Oregon) geese, herons, ducks. Gulls, ducks, geese, coots, shore birds. TuIeLake- birds. Idaho; Deer Flat Ducks, geese, pheasants. Minidoka Montana; white pelicans, ducks, coots, herons, avocets, sage grouse. bitterns, phalaropes, avocets, sand- pipers, yellowlegs, plovers, sharp- tailed grouse, pheasants. Pablo Do. Pishkun Gulls, ducks, geese, swans. WUlow Creek Nebraska; North Platte... Ducks, geese. Ducks, geese, swans, shore birds. Nevada; Boulder Canyon (see also Arizona) Fallon. Waterfowl, mammals. New Mexico; rants. Rio Qrande Grebes, cormorants, ducks, geese. Oregon; Cold Springs shore birds. Ducks, geese, swans, herons, sharp- Klamath Lake (see also California). McKay Creek Upper Klamath South Dakota; Belle Fourche Utah; Strawberry Valley. . . Washington; tailed grouse. Gulls, ducks, geese, coots, shore birds. Ducks, geese. Do. Ducks, geese, curlews, prairie chickens, pheasants. Ducks, sage grouse. Wyoming: Pathfinder tailed grouse, Hungarian partridges Ducks, geese. Wildlife in the United States 11 Table V. — Areas administered as wildlife refuges by other departments incidental to their primary use Table VI. — Areas nominally in Stale game refuges, 1931 Alaska California Georgia Do Kentucky Louisiana Maryland.- Michigan Mississippi Montana _.. New Mexico North Carolina Ohio Pennsylvania South Dakota Tennessee Wyoming Total acreage Acres 612,000 129 6,562 150 110 5,640 40 2,680 1,324 1,281 45,423 677 57 2,530 5, 548 4,299 56, 132 644, 582 Department administering Commerce. Do. Interior. Commerce. Interior. Commerce. Interior. Do. Do. Do. Do. Do. Do. Do. Do. Do. Do. Public Areas In Other Uses That Should Be Designated as Wildlife Refuges For a discussion of this subject see contributions by the Forest Service and the National Park Service. Private Lands Employed as a Habitat for Wildlife Nearly all privately owned lands bearing vegetation are, or may be, employed as a habitat for some form of wildlife. In many States no shooting is allowed on certain areas, and these, therefore, function as wildlife State Acres State Acres 100,000 2,499,670 35,495 2,399,215 2,601,960 18, 026 12,788 2,472,713 26,525 245 640 Arizona. California.. North Carolina 124,894 9,684 20,928 Connecticut Ohio Florida 2,731,662 14,000 3,479,120 3,360 26,120 8,951 7,955 63,000 329,863 244,212 40,269 9,530 71,560 3,084,756 Oregon 2, 936, 720 Georgia Pennsylvania: Primary 77,836 Au.\iliary 27,635 Idaho Illinois 105 471 17,447 Kansas. .. _- South Carolina 67 423 190, 933 1,560,520 Maine Texas 3, 075, 905 Utah 2,119,000 12,088 16,700 Washington 2,619.040 Mississippi . . 120 547 58, 213 2,419,336 72, 622 3,425,112 9,830 46,385 Montana 4, 461 370 Total acreage Nevada 44,016 458 refuges. Marsh areas, fence rows, and tree- or shrub- bordered stream or ditch banks should be utilized for wildlife production. Recommendations by tlie Biologi- cal Survey for improving the farm environment for wildlife have been set forth in a recent publication of the Department of Agriculture. ° • Grange, W. B., and McAtee, W. L. Improving the Farm E.nvirosment for Wildlife. U. S. Dept. Agr. Farmers' Bui. 1719, 62 pp.. iUus. 1934. SECTION II POLICIES WITH RESPECT TO WILDLIFE' Areas Now in Public Ownership Devoted to Wildlife Federal Ownership: Among Federal lands on which wildlife is protected are national parks, national forests, certain national monuments, and national wildlife refuges established specifically for game birds and mammals, fur bearers, and other valuable and interest- ing forms of animal hfe. For all these there should be formulated definite policies wdth respect to wildlife. Specialized: There are now under Federal owner- ship and administered by the Bureau of Biological Survey, of the Department of Agriculture, approxi- mately 1,000,000 acres of surveyed lands in the United States, specifically devoted to wildlife protection, besides unsurveyed islands of some extent designated for the purpose in Alaska, Hawaii, and Puerto Rico. Most of these areas are breeding and feeding grounds for birds, including some for sea birds and migratory waterfowl, but a few big game and fur animal refuges also are included. Table II gives the areas by States of the refuges administered by the Biological Survey. In table I is given a list of these refuges and the chief species for the protection of winch each was established, and in tables III and IV are treated the wildlife refuges administered by the Biological Survey on areas pri- marily devoted to other uses, cliiefly on reservoirs for irrigation projects under the Bureau of Reclamation. In addition to these areas it is imperative that water- fowl habitat to the extent of approximately 17,000,000 acres be obtained for the conservation of migratory game and other birds protected under treaty with Great Britain. These land and water areas for the waterfowl should be in two classes — breeding grounds and feeding and resting grounds. Migratory water- fowl are in a particularly percarious condition and, unless positive action in their behalf is taken without delay, there is little hope of saving some of the native species now in dire straits. These waterfowl areas should be grouped along the major flyways in such way that the birds will find some sanctuary where they can feed and rest in safety, and where they can breed without human interference or molestation by predators. This will involve in some cases setting aside Federal lands, but more frequently lands now privately owned should be purchased and placed imder Federal administration. > Contributed by the Bureau of Biological Survey. 12 For upland game birds and game mammals, approxi- mately 21,000,000 acres are needed, especially for such big-game animals as mountain sheep and antelope, for which no protection is provided on the national forests except on a few game refuges. The largest antelope herds are now on pubhc domain and private holdings, a fact that must be taken into consideration in making provision for the future of the herds of this most interesting North American mammal. Moun- tain sheep are in a precarious condition, and certain of the desert ranges, particularly in Nevada and Arizona, should be set aside and developed for the use of these unique animals. For the use of sage hens, prairie chickens, and sharp-tailed grouse, areas on the public domain need also to be designated as wildlife refuges. Primarily Devoted to Other Major Uses But Designated for Wildlife Sanctuaries: In the majority of cases national forests and national parks provide adequate summer range for the deer, elk, and other big game animals that are found therein. Wliere there is any deficiency in ability to care for these herds, it is usually from lack of winter forage. This is a pressing problem from the wildlife standpoint, and the lower lying lands along the boundaries of both the forests and the parks should be designated as winter refuges, and forage should be reserved for these big-game animals. For a full discussion of this subject, see contributions by the Forest Service and National Park Service. State Ownership: While the total acreage in many of the State game refuges is impressive, httle of the land is owned by the States involved. A few States, notably Pennsylvania, New York, and others in the East, have made some progress toward actually acquiring land for use as game refuges. In most of the country, however, such areas are either in Federal or private ownership but have been designated as wildlife refuges by legis- lative authorization or by regulations under State game commissions. Many, however, are merely paper refuges, with no control of factors detrimental to the valuable wUdlife they purport to foster. This condi- tion wiU probably continue to exist until some method can be worked out of removing State game administra- tion from political control. It is almost a.xiomatic now that whenever a man begins to get some acquaint- ance with the problems of his State and with methods of handUng them, he is tlirown out for pohtical reasons, to be succeeded aU too frequentl}^ by one who is entireh' ignorant of the problems he faces. Until this defect is Wildlife in the United States 13 corrected, additional State refuges are not so important in tlie wildlife conservation scheme as woidd be the better handUng of national lands and the development of a consistent Nation-wide policy in game management on Federal lands. There should be a national wildhfe program administered and coordinated by a single Federal wildlife agency. Units Owned by Local Government: Units owned by other local governments are usually too small to be entirely satisfactory as game management units or refuges. Provisions should be made for the exchange or purchase of many local areas, to consolidate them with major holdings of either State or Federal organiza- tions. In a few cases, such as at Lake Merritt, in Oakland, CaUf., a small refuge has been biult up into a public attraction of the first order. There is room for many more such well administered local units through- out the country. At present many cities and villages are interested in such a progi'am, and every encouragement should be offered them to continue such interest. Programs of Acquisition ; Suggestions for Modification The present program of acquisition of migrator.y-bird refuges by the Federal Government has as its objective two major purposes. The predominant purpose is the acquisition of all possible areas wliich by reason of their present natural environment will attract migratory birds in the nesting season, or which through develop- ment or by restoration to primeval conditions may be made attractive to nesting waterfowl. The second objective contemplates the acquisition of more or less extensive resting and feeding areas in the flight lanes used by migratory birds spring and fall, in order that they may be provided with conveniently situated sanc- tuaries f urnisliing water, food, and cover in an unmolested env-ironment, protected from both man and predators. At present, as pointed out by Sal.yer,- the major efforts are being concentrated on the breeding-ground phase of this migratory-bird-refuge restoration program, but not entirely to the exclusion of efforts for providing resting and feeding grounds. The size of refuges to be so acqidred should as a rule be not less than 10,000 acres, but single units where the en\'ironmental elements are favorable might comprise 50,000 acres or more. An ideal sj'stcm of refuges woidd contemplate a series of major projects approximately 300 miles apart in each one of the four major waterfowl flyways described by Lincoln," that extend from the Canadian border to the southern Umits of the United States, and in proximitj' ' Salyer. J. C, U. A Program ok Waterfowl Restoration. U. S. Dept. Agr. Circ. 339, U p.. 1934. ' Lincoln, F. C. The Waterfowl Fltways or XORTn America. U. S. Dept. Agr. Circ. 342, 12 p. illus., 1935. to most, if not all, of these might well be created groups of refuges smaller in size that would be beneficial in the wildhfe restoration and conservation program. These are the ideal objectives. Their accomplish- ment is largely contingent upon the availabihty of adequate funds with wliich to carry them out, but in some parts of the United States, notably where the natural environmental conditions are most attractive to nesting migratory birds, there should be a radical change in point of view regarding the drainage of lands and the ill-ad viscil use of water through waste and futile reclamation projects. Without some such change in point of \'iew, it will not be possible to accomplish fully the objectives of the Biological Survej^ in the matter of an adequate system of refuges. The critical situa- tion of the waterfowl in 1934, following a series of un- favorable 3'ears, has been elsewhere presented bj- Bell and Preble,* their findings having been based on inten- sive investigations by members of the Biological Sur- vey and cooperating organizations and individual sportsmen and other conservationists. Integration of Federal And State Policies Federal and State Governments should work in close cooperation in deahng with game problems and as far as possible any causes of friction should here- moved and questions of jm-isdiction settled. The national forests are the natural reservoirs of most of our large game, especially deer. The Forest Service is charged with the general care of these forests, but the game pohcy, or lack of pohcy, of some States in which forests are located may hamper cooperation or completely nullify well rounded forest management, which should encompass all forest resources — game as well as timber. For example, on many western na- tional forests the grazing of domestic stock is an impor- tant industry and may constitute the major use of the land. In places, deer, elk, and other kinds of big game in uncontrolled numbers seriously compete with domestic stock for food, and overbrowsing by both classes of animals may be seriously destructive to forest reproduction. In addition, such intensive use results in general watershed denudation, and this greatly accelerates destructive erosion. Obviously the urgent need is the regulation of numbers of both stock and game in accordance with the food supply, in such way that none of the conflicting interests will be sacrificed. Such regulation is a dillicult and complex problem at best, and calls for a comprehensive and sustained plan of wildlife management. An adequate solution of the i)rol)k'ni is virtually impossible where dependent upon ])olicies changing with State politics. < Bell, W B., and I'reble, E. A. Status of Waterfowl in 1934. U. S. Dept. Agr. Misc. Publ. 210, IS p., illus. 1934. 14 Land Planning Report Encouraging Game Production On Private Lands Studies of local needs should be made and food and cover plants developed where required. In many places the restoration of game that has become extinct is necessary. One measure usually needed in all places is to extend more adequate protection to the wildlife already present. Educational programs should be developed, includ- ing the use of motion pictures, to acquaint landowners with practical means of producing game and with profitable ways of harvesting the crop. The attitude of the farmer especially has an important bearing in this connection. Farmers should be shown that rab- bits, quail, pheasants, and other game, as well as fur- bearing animals and fishes, have a potential crop value similar in kind to that of any other product of their acres. They should be encouraged to expect a profit through the sale of shooting or trapping rights, com- mensurate with their efforts to produce game, under regulations fixed by the State. Planned utilization of land for wildlife development, for game management, and for human enjoyment of large natiu-al wilderness areas is deserving of the inter- est and active support of all concerned — the land economist, the sportsman and conservationist, and the farmer. Tlie policies in such a program should be such as not to sacrifice human interests, but rather to advance them tlu-ough better realization of the importance of the wildlife resources and of their func- tion in human economy. The policies should not give wildlife priority rights everywhere, to the exclusion of other enterprise. They should, however, recognize clearly that wildlife is entitled to its share of land and water and to have adequate areas set aside solely for its use and benefit. The time when the best uses are being sought for submarginal private holdings is appropriate for making plans to retm-n a vast acreage to its original and best use — as habitat for wildlife. The need is now so pressing that it deserves a major place in the national land utilization program. SECTION III WILDLIFE MANAGEMENT IN NATIONAL FORESTS' Encouraging Game Management On Public Land in Other Use Wildlife in National Forests: The production and management of wildlife is an important objective in the coordinated management of the national forests to produce the liighest economic, scientific, aesthetic, and recreational values. Management contemplates the production of the largest wildlife population con- sistent with: (1) Use and needs of other resources, and (2) permanent food supply of animals, birds, and fish. Wildhfe includes all species of game animals (large and small), game birds, fur-bearers (both predatory and herbivorous), nongame birds, and fish. Of all wildhfe, big game animals — deer, elk, bear, mountain sheep, and others — are by far the most spectacular. On some national forests, however, small game animals and birds — squirrel, rabbit, grouse, and turkey- — are at present far more abundant, and consequently of greater importance than big game. Although the national forests are the habitat of comparatively large proportions of the total numbers of many of the important wildlife species- — particu- larly in the Western States^ — distribution of wildhfe on the forests is by no means uniform. Some areas are overstocked, others could support larger popula- tions. Each locality presents its own peculiar prob- lems of correlation of uses and interrelation of wildhfe species. Scientific Management of Game on Federal and State Refuges: Despite inadequate basic knowledge, lack of sufficient trained personnel, and a question as to the legal control over wildhfe, the Forest Service aided by the Biological Survey has made a start in scientific management. On Federal game refuges, notablj" the Kaibab in Arizona and the Pisgali in North Carolina, the regulation of game populations within reasonable bounds, by means of controlled hunting, is being practiced. The 283 State game refuges, comprising 21,000,000 acres on national forests, provide sanctuary and breeding grounds. Forest officers cooperate with the States in the enforcement of game laws. Fire protection alone is of extreme value in the develop- ment and maintenance of adecjuate wildlife popula- tions. Stream surveys in cooperation with the Bureau of Fisheries and the improvements of habitat in ' Contributed by C. E. Rachrord and L S. Qroas o( the Forest Service, Depart- iiiCDt of Agriculture. accordance with prescribed plans are also underway. Wliile wildhfe ranges have long been studied by local forest officers and preliminary management plans developed, this work is being carried further in co- operation with the Biological Survey, whose trained personnel brings to it the expert advice so badly needed. Essentials of Management Program: Two tlungs need to be done: (1) Bring abovit adequate stocking of all land and water areas on national forests with suitable species; and (2) initiate satisfactory man- agement. A number of obstacles must be overcome in the consummation of this program. Of greatest present concern is the inadequacy of the game laws of most States. Existing game laws, almost without excep- tion, have been directed toward preservation rather than sustained production of wildlife. Results vary with character of the laws and their enforcement. Biological relationships are not changed by legal enact- ments and frequently operate to nullifj' them. Mod- ern hunting, fishing, and trapping laws, based upon control of numbers of game lulled by locahties, are essential to rational management on a sustained production basis. Western national forests at present include only about 40 percent of the winter range needed to support big-game populations large enough to utihze available summer range. Despite all possible adjustments of range use between wildlife and domestic livestock, this shortage of winter game range constitutes a serious limitation to increased wildlife populations. Control of disease and predators constitute fields wherein increasing activity is needed in research into basic principles and practical methods of application. Law enforcement is, and probably always will be, an important part of wildlife management. Close coop- eration with State game officials, encouragement to the States to paj^ greater attention to this phase, and, where necessary, increase in Federal enforcement personnel are indicated. Cooperation of Forest Service and States. — In order to redeem its responsibilitj- in wildhfe management, the Forest Service will continue to extend full cooperation to all States willing to join in constructive efforts for sound development and use of tlus resource. In any case where it is not practicable to proceed on this basis, the Forest Service should assume full authority and responsibihty for wildhfe management in the 15 16 Land Planning Report national forests. The former course is much to be preferred. Steps necessary to its accomplishment are: 1. Educational courses in universities and colleges to develop personnel trained in scientific and practical wildUfe management as related to forestry. 2. Provision for the employment of an adequate, qualified personnel, both Federal and State. 3. Clear-cut definition of the responsibility for wild- life management on Federal land by Forest Service and State officials. 4. Complete revision of State game laws, based on biological facts rather than pohtical consideration and vesting broad general powers in a nonpartisan board or commission. 5. Careful survey of wdldhfe resources, both publicly and privately owned, by competent personnel; devel- opment of wildhfe management plans. 6. Adequate appropriations for the employment of personnel necessary to the application of management plans and the extension of research. Economic and Social Values of Wildlife Areas Value, Eitent, and Conditions oj Wildlife on Forested Lands : The forested lands of the United States provide the largest part of habitat for most of that remaining wildlife which is important for food, fur, hunting, and aesthetic purposes. In 1929 the number of hunters and fishermen in the United States was placed, by the Senate Committee on Conservation of Wildlife Re- sources, at 13,000,000, an increase, it is estmiated, of 400 percent in a single decade. The total positive national value of all wildlife is estimated (by the Bio- logical Survey) at more than $1,000,000,000 annually. Wildlife has decreased, and is stUl decreasing, on much of the forest land of our country as a whole. - Reasons are obvious: The enormous increase in num- bers of hunters, disease, deterioration, or destruction by fire (and otherwise) of forest cover and forage, lack of management on at least four-fifths of the more than 600,000,000 acres now classed as forest or potential forest land. Increase of Game Animals in the National Forests: Contrasted with the general situation the number of game animals on 167,000,000 acres of national forests presents a difl'erent picture. Estimates, based upon observations made by hundreds of forest officers who spend a large part of their time on the game ranges, show an increase of 100 percent between 1921 and 1933. (See chart showing trends of big game populations 1921-33.) Double Value of Wildlife: Three-fifths of the esti- mated total wildlife value is classed as economic, the remainder recreational. Economic values include serv- ' "Wildlife Conservation" S. Eept. No. 1329, 71st Cong., 3d sess. ices of birds, rodents, and other mammals in destruction of insect pests, as well as returns in the form of meat and fur from hunting, fishing, and trapping. Measurable social or recreational values consist of expenditures for licenses, transportation, and equip- ment by hunters and tourists. The large intangible benefits derived by more than 31,000,000 people who annually visit the national forests cannot be ade- quately estimated in cash values. Hunting, fishing, and nature study attract large numbers of these people. Contribution of Game to Busiiiess Activity: That those who go in search of game contribute in many ways to business activity throughout the country, is shown in the total annual returns of about $158,000,000 for hunters' expenditures and $254,000,000 of tourist ex- penditures credited to wildlife attraction. Hunters' expenditures include equipment, arms, and ammunition purchases and, in addition, transportation, lodging, food, guide, and other personal expenses. Tourist expenditures are concerned with all of these except arms and ammunition purchases. Examples of the value of national forest wildlife may be cited. During the 1933 regulated hunting season in the Kaibab National Forest in Arizona, 932 hunters spent more than $43,000 incident to their sport which resulted in bagging 859 deer. The average hunter spent $47, 40 percent of which represented subsistence, 25 percent transportation, 15 percent license fees, and 20 percent other expenditm-es. On the more accessible Sierra National Forest, the records for 1932 show that 1,400 deer were killed by 6,145 hunters, whose average expenditures approximated $23 each. The meat ob- tained by hunting and fisliing is a welcome addition to the larder of many local residents, and highlj^ prized by many of the hunters and fishermen who travel con- siderable distances to enjoy their favorite sport. Trap- ping fur-bearers provides a cash return for many a family, and, as wildlife management becomes more intensive, will no doubt assume greater importance in connection with the conscious effort of the Forest Serv- ice to supply more work for the dependent population on each national forest, by developing all resources so as to secure maximum yields. Recent studies by the Forest Service show that there are within, and near the national forests, more than 5,000 farms and ranches primarily operated for recre- ational purposes. A considerable portion of the fiveh- hood of the owners and operators of these ranches is derived from conducting hunting and fishing parties on the national forests. An example of the magnitude of these undertakings is obtained from the 1929 report of the Dude Ranchers' Association. Fifty-one ranches comprising property valued at approximately $6,250,000 showed receipts in 1929 of nearly a milhon and a half dollars. Wildlife in the United States 17 t9ZJ I60 1922 /92J /92<- •J /92S /926 /927 f928 /929 /930 /93/ r932 k 10 1933 /60 /4-0 Qrtginot Estimates Corrected by More Complete Data for R-a , 1923. 1925, 1926 and R-9 , 1929 'fO 8.084^ 115,197- FIGURE I- INCREASES AND DECREASES IN BIG-GAME ON THE NATIONAL FORESTS 1921-33 Increasing OjJjiort unity fur Wildlife Observation: With increasing intensity of wildlife management on the na- tional forests, much greater ojiiiortuiiities will also be created for natm-e study and wildlife observation. A large proportion of the millions of annual visitors are primarily concerned with getting back to nature for a short period. A chance to see — and perhajis photo- graph— a feeding deer, a band of elk, a drumming grouse, or a beaver at work adds zest to the adventure and greath' increases the satisfaction derived from the trip. Wildlife and Other Land Uses Interrelation: Wildlife is directly concerned with practically all other forest-land uses. The full pos- sibilities of such cot)rdination, while apparently great, can be determined onh- by a most careful study of the biological relationships between the flora and fauna of each locality. It is desired, however, to set forth some of the significant developments in the interrelation of wildlife with other uses of the national forests. In general it can be stated that those things neces- sary for the protection of the forest from fire, as well 18 Land Planning Report as the application of such measures as are necessary to keep forest land productive, contribute to the welfare of wildUfe. The requu'ements of game for various types of cover are directly related to silvicultural methods used in cutting, slash disposal, and reforestation. Cutting operations break the forest canopy, and provide food and cover by encom'aging the growth of low herbaceous and shrubby vegetation, and succulent sprouts highly relished by various wildlife species. On the other hand, game populations so increased may be expected to diminish gradually as the forests mature and the food supplj' diminishes. Management must seek a proper balance based on the relative importance of each resource in the locaUty concerned. Modifications of management plans for both timber and wildlife will often be possible. For example, it may be practicable on intensively managed areas to so locate timber cuttings that at all times there wiU be sufficient feed and shelter, within the cruising range of the wildlife species concerned, to maintain relatively large populations. Silvicultural Practices and Wildlife: Silvicultural practices on eastern national forests are designed to aid in the production of wildlife as well as timber. This is given definite apphcation on the AUeghenj^ Forest in Pennsylvania. Of the timber trees there, beech is the most important producer of wildlife food, beechnuts being utilized bj^ deer, bear, squirrels, and grouse. Beech, although a valuable tunber tree, has a decided tendency to develop a wide-spreading "wolf tree" habit in open stands. It is silviculturally desu-able, therefore, to remove defective limby beech on cut-over areas, so that the next stand may be composed of tall, straight, well-formed stems. Following a study of the situation it was decided, however, that it was desirable to reserve an average of four large crowned beech per acre, incident to silvicultural unprovement of cut-ov^er areas. This policy has been put into practice as a definite measure for coordinating wdldlife with timber production. Serviceberry, a nontimber species, is definitely protected on the Allegheny, because of its value as a producer of wildlife food. The George Washington National Forest in Virginia furnishes another example of the effect of silvicultural measures on wildlife. In 100,000 acres, of the Massa- nutten Working Circle 1,249 acres of the best sites were subjected to timber stand improvement treatment by the Civilian Conservation Corps in 1933. An average of 37 stems were cut per acre, 70 percent of which were 10 inches or less in diameter. The average acre supported 253 stems after treatment, including such producers of wildUfe food as hickory, oaks, cucumber, dogwood, black gum, and sumac. The area treated on this working circle is only slightly more than 1 per cent of the total, and the treatment is expected to promote better production of timber without inter- ference with wildlife. As more fundamental data regarding intricate local biological relationships become available, even greater coordination should be possible. Improvement of Game Conditions an Objective of Forest Planting: The importance of wildlife require- ments is also recognized in the national forest planting program. Improvement of game conditions is listed ^ as one of the objectives in planting. It is the policj' to plant first those areas which may be expected to produce the most valuable timber crops; that is, the most productive sites. It is recognized, however, that exten- sive areas of unbroken coniferous forest — either virgin stands or maturing plantations — afford ample cover, but little feed for many wildlife species. It is there- fore necessary to make definite provision, whenever possible, for supplies of fruits or browse, within or near coniferous plantations. Natural growth may some- times be utilized for this purpose. The Canaan Moun- tain planting project of the Monongahela National Forest in West Virginia may be cited. This area of 21,000 acres originally supported dense stands of spruce, hemlock, and hardwoods. Logging and repeated fires wrought colossal changes. Since the tract has come into national forest ownership, protec- tion from fire has resulted in a new growth of hard- woods, with some admixtm-e of conifers, over most of the area. On about 2,500 acres of the most severe burn, however, it has been necessary to reestablish the forest by plantmg. Red spruce, the native species, was the most logical choice, and was the chief species used. Small quantities of black cherry, a wildlife food pro- ducer, were planted in mixture with the spruce. Of greater importance to wildlife, however, is the natural growth of mountain ash, viburnum, holly, and other hardwood species. During the establishment of plantations on this project, most of this native hard- wood growth was undisturbed. As the plantations mature, it is anticipated that adequate wildlife cover and food will be suppUed by the resulting coniferous hardwood mixture. The planting problem on the Ozark National Forest in Arkansas, on which the predominating type is hard- woods with more or less short-leaf pine in mixture, consists of restocking abandoned old fields scattered throughout the forest. Short-leaf pine is used almost exclusively for planting the old fields. The result will be comparatively small blocks of almost pure conifers, breaking the hardwood cover. Many of these old fields, however, support a scattered stand of persim- mon, sassafras, black gum, and other hardwood pro- > p. 129-S, National Forest -Manual. Wildlife in the United States 19 ducers of \vildlife food. These hardwoods, as in the case of those in the Monongahela Forest, may be expected to develop in mixture with the coniferous plantations. The plantations should eventually pro- vide both shelter and subsistence for wildlife. Some wildlife species, notably rabbits and deer, maj' become inimical to the estabUshment and growth of plantations. In the national forests of the Lake States, rabbits have increased to the point where brush areas cannot be successfully planted. Planting must either be linoited to more open areas (where rab- bit damage is much less severe) or measures under- taken to reduce the rabbit population. Modification of Wildlife Alanagement: Wildlife management plans must sometimes be modified to accord with plans for timber management on specific areas. Relatively inaccessible portions of many na- tional forests must necessarily be handled under extensive forest management, involving less frequent cuttings wliich cannot be confined to small areas scattered tlirough the entire unit. Under such condi- tions, wildhfe management must be directed toward smaller populations, changes in relative number of species, or a fluctuation in populations with character of cover on the various portions of the unit. Wildlife and Domestic Livestock- on Forest Range: Probably in no other phase of forest-land management is the interrelation of wildlife production and other forest uses so apparent as in the use of forest range by wildhfe and domestic hvestock. Increase of wildlife on the national forests, requiring in some instances adjustments between game and domestic stock, have sharply focused the attention of foresters and game specialists upon the need for care- fully planned and coordinated use of forest grazing areas. The outstanding example is on tiie Kaibab Plateau where an area now comprising about 727,000 acres was set aside in 1906 as a Federal game preserve, with deer the principal game species. A measm'e of pro- tection was afforded by the operations of the United States Biological Survey up to 1923 in destroying predators, and conditions were made favorable for the deer in other ways. As a result, deer increased until the forage producing capacity of the area was insuffi- cient not only for the deer and domestic stock, but even for the deer alone. Notwithstanding a reduc- tion of domestic livestocks, to a point where competi- tion with deer was at a minimum, the continued in- crease in deer resulted in great tlamage to the more valuable forage plants and timber reproduction. The productive capacity had been reduced to a degree which would sustain not more than 5 to 10 percent of the game which it was capable of supporting under proper conditions of normal use. Action has been under way for several 5'ears to remedy the situation, and the excessive deer population has been reduced. Restoration of properly balanced wildhfe and vegeta- tive conditions presents intricate problems of biological relationships and management. Increasing knowledge of forage requirements of game, as difl'erentiated from the recjuirements of domestic hvestock, offers excellent opportunities for determining a well-balanced relation between numbers of game and hvestock. Satisfactory determination of permanent ratios requires additional research and, in tlic final analysis, should afford the means of obtain- ing the best development and use of the wildhfe resource in its proper relation to timber and other lines of pro- duction. Proi'iding a Proiier Balance Between Wildlife Species: The proper balance between the various classes of wildlife is also important. Experience in many places has amply demonstrated that the status of the natural wildlife population may be disturbed bj- changes of food and cover conditions that favor one or more species over others, or protection by man of certain species from natural enemies. The ultimate goal of land use is the mdespread application of principles that will result continuously in proper balanced pro- duction of timber, other forest products, and a variety of wildlife. Competition between ^\•ildlife species may at times be a vital factor. In the southwestern part of the Sacramento Mountains in New Mexico, on the Lincoln National Forest, wild turkey are very scarce. They have, in fact, nearly- disappeared from that part of the mountains. On this particular range, deer have in- creased on private holdings and adjacent national forest land. Use of the range bj^ deer and domestic stock has so decreased the available quantities of the more palatable shrubs that the deer are forced to supplement their usual food with acorns and juniper berries. This use of mast by deer, together with reduction of other turkey food tlu^ough overgrazing, destroys the winter forage supply for turkey. In the north end of the Sacramento Mountains, where deer are not plentiful, there are great numbers of turkeys. The Sitgrcaves National Forest, in Arizona, provides another instance of the need for careful balance between wildlife species. Elk have increased on the simuner range here to such an extent that they are kiUing out the wallows in high mountain meadows. This, together wnth overgrazing of yuccas and other plants utilized bj' elk along the canj-on walls, and trampling of the soil, is resulting in erosion. The delicate relationships between a constant supply of clear water, the develop- ment of aquatic life, and sliaded pools, so necessaiy to provide proper habitat for trout in the streams flowing from this range, are upset by erosion. Floods 20 Land Planning Report scour the streams, wash out plant Hfe, destroy trout shelter and breeding places, and materially reduce the capacity of the streams to produce trout. This serves to illustrate a little-considered relationsliip between big game animals and fish. Winter Range Jor Game Animals: Game animals have free access to every acre of the national forests which is suited to them, including the 82,000,000 acres of range used by domestic livestock. In stocking these ranges with domestic livestock, it is the object to provide amply for the needs of game during that period of the year that game is dependent upon forest range for feed. Emphasis should be laid upon that last statement. Generally speaking, the national forests would support during the summer season many times the present numbers of game animals. In the Western States, however, most of the best winter ranges are outside the national forests, generally on private lands on which little if any consideration in given to the welfare of game. This presents one of the most important prob- lems to those interested in big game. Obviously, any expectation that a larger number of game animals can be maintained than is represented by the capacity of the winter range is based upon a slender reed of hope which fails to recognize realities. To be more specific, it is estimated that national forest ranges in Colorado could support during the summer months 300 percent more big game animals than are now present, if protection and mangement were provided on some 10,000,000 acres of winter range outside the national forests. In this case, as in many others, winter range is the governing or limit- ing factor in game production. The part which winter range plays in game production might be illustrated by many other specific examples. The Targhee National Forest, in Wyoming, however, is sufficient to show the relationship between sheep and big game animals. This forest supports more than 200,000 head of domestic sheep and a small number of game animals, yet inspec- tion by grazing experts mdicates that its summer range is conservatively used by sheep, and that there is an enormous quantity of food available for game animals. The area is deficient, however, in winter feed for game. Cold and severe winters force the game animals to the lower areas on private lands outside the forest, where they are subjected to shortage of feed and other perils. The exclusion of sheep from the national forest would fail to relieve a condition of tliis kind. Grazing of sheep, particularly on western ranges, may conflict with ground nestmg game birds, such as grouse, sage hen, and turkey. On national forest ranges, however, the nesting habits of game birds have been taken into consideration in the estabhshment of opening dates for the sheep grazing season. This date has been set sufficiently late in the season so that the eggs have hatched and the chicks have left the nest before sheep are admitted to the range. Management of the domestic animals is so regulated as to avoid a concentration of grazing on any particular area. This method greatly reduces the actual conflict and damage to nests which occurs on areas of heavy concentration of domestic livestock, such as lambing grounds, drive- ways, and bedding grounds. Control oj Predatory Animals: Predatory annual con- trol is an important factor in wildlife management. This is emphasized by reports for 1932 from field officers, which show that for every deer killed by hunters on the national forests, one and a half deer were killed by predatory animals. This raises the question as to whether game is raised to afford sport or to feed pred- ators. It seems reasonable to assume that the pred- ator may play an important part in assisting in the regulation of numbers of game anmials to the capacity of the range, or may be responsible for a serious decline in the herds. That sheep grazing is not an important limiting factor in game production can be illustrated by the situation in the eastern and southern regions. These regions have more than 5,000,000 acres of the best and most natural game range. A total of only 2,600 sheep are grazed on 8 national forests, yet there is a very small game population per acre. About one-fifth of the big game animals on the forests in the Eastern United States are on a Federal game refuge (the Pisgah Refuge), where full protection is accorded under the authority of the Forest Service. But wildlife on areas outside the game refuge is subject to all the perils of a poor regulatory system, including illegal killing and destruction by dogs. Fishing Benefited by Forest Management: Fishing in the 60,000 miles of national forest trout streams is also benefited by scientific forest and range manage- ment. These measures tend to conserve water sup- plies and maintain continuous stream flow, so essential to sustain production of satisfactory quantities of game fish. Fur Bearers in National Forests: The production of sustained wildlife yields on the national forests involves not only game animals, game birds, and fish, but also fur bearers. Among the latter are such carnivores as the mountain lion, lynx, fox, and mink. Careful study of local biological relationships between fur bearers and both game species and domestic livestock is one of the pre-requisites to the formulation of wildhfe-manage- ment plans. Damage must always be weighed against value, and an effort made to secure maximum benefits. The beaver is an excellent example of a nonpredatorv fur bearer. Here again, compHcated relationships exist. Wildlife in the United States 21 TOTAL FOR UNITED STATES 133,086.262 ACRES 'includes SoLtth DakoTc: NebrasKa and Oklahoma Areas) 16.610 E J - 916 3° wwsawild Life Winter Ranges (Summer ond/brWmter use in Arizona and New Mexico) Area used by Cattle , Available Area used by Sheep \ for Area not used by either! vvild Life CatTle or Sheep Numerals on mop are percents ^^j Existing National Forests MiLCS FIGURE 2 - LOCATION OF GAME REFUGES AND WINTER RANGES NEEDED IN CONNECTION WITH NATIONAL FORESTS. AND PERCENTAGES OF NATIONAL FOREST LANDS USED BY CATTLE AND SHEEP 22 Land Planning Report Studies on a number of national forests show that beaver are of great value in watershed control. Beaver dams create small ponds and lakes, impounding and conserving the water. An instance in which this pro- duced measurable economic value was noted on the Gunnison Forest in Colorado. In 1924, the water supply for the Minnesota Reservoir ran low and suffici- ent water was not available for late irrigation. Four- teen large beaver ponds on the streams above were tapped, filling the reservoir. This supply, which had been stored by the beaver, practically saved the fruit crop on every orchard dependent upon this irrigation project for water. The fruit growers valued this emergency water supply at $15,000, an average of more than $1,000 for each of the dams tapped. Control and Regidation of the Beaver Necessary: Beaver are responsible, however, for considerable damage to some irrigation projects, by diverting the flow from ditches and flooding fields. Damage also occurs by the flooding of roads, trails, and railroad embankments. Under such conditions, careful control is necessary. Forage conditions may be improved by beaver dams which check the flow of rapid streams, spreading out the water and resulting in greater production of willows and succulent grasses. The beaver population must be regulated, however, since natural increase in numbers frequently results in shortage of aspen and other beaver feed. Cases are on record where shortage of food has resulted in wholesale migration of beaver to other watersheds. In other cases, unregulated trapping of beaver has also resulted in virtual elimination of the species on certain streams. The result of either forced migration or destruction by man are the same. The dams are washed out, erosion occurs, stream flow tends to revert to alternating floods and low stages. The solution to such problems involves maintenance of the beaver population on each stream at such a level that the maximum benefits will accrue without undue dam- age to other uses, and within the limits of available feed. This may be accomplished by regulated trapping. Beaver dams on cold, rapidly flowing streams usually improve fishing conditions by providing deep back- waters, suitable spawning grounds, and increased feed. On the contrary, the results of beaver dams on sluggish streams may be detrimental to game fish. For example, water in the dams may reach a higher temperature than can be tolerated by the fish, or decaying vegetable matter may create an undesirable habitat. Use of Water Areas: That the protection and propa- gation of wildlife species can be properly coordinated with other land uses is further indicated in the relation- ship of the use of water areas by waterfowl to the use by recreationists and domestic livestock. On some of the lakes on national forest areas where feed conditions are suitable to resting and breeding grounds for migra- tory birds, the shore line generally used for nesting purposes is also occupied by fishermen. Notable in- stances have occurred where, due to increase in recrea- tional use, migratory birds have been deprived of the nesting grounds along the shores of the lakes. This situation may be easily corrected by the prohibition of fishing on those portions of the lake shore best adapted to waterfowl use, and the posting of such areas against trespass. The same situation applies where many of these lakes afi'ord the only available watering place for domestic livestock. If the livestock is allowed to concentrate around the shores, the lakes are no longer valuable for wildlife purposes except as resting grounds. If, how- ever, the livestock is definitely restricted to watering at certain portions of the lake, the needs of both are met. Conclusions: The foregoing discussion and examples of the possibilities and limitations of the association and production of wildlife with other land uses point to a number of conclusions. 1. Wildlife production may be definitely coordinated with all other land uses on the national forests. 2. Many biological relationships, basic to complete coordination, remain to be investigated. 3. Coordination must be applied locally. Dominant use may vary within regions, national forests, drainages, or smaller units. 4. Sustained production of wildlife implies regulated use, necessary to control populations within available food supplies, and, in varying degree, within other land use requirements. 5. Adjustments between uses, involving priority ratings, must be expressed in integrated plans for management of wildlife and other national forest resources. Areas Suitable Only for Refuges Since the refuge idea must be fitted into the proper place in the management plan as a whole, permanent dedication of any area to a wildlife refuge has been found undesirable on the national forests, except as a means of perpetuating an almost extinct species or where the area will serve no other useful purpose. A refuge may provide excellent cover for breeding or other purposes when established, but, because of changing biological relationships, may not be satisfac- tory for that purpose on a permanent basis. The system of management must be sufficiently flexible to permit the closing of areas to hunting and fishing, and the establishment of other restrictive measures when and where conditions justify such action. It will probably always be necessary to reserve certain areas to meet the purpose contemplated by the refuge idea, but in the absence of complete management plans for Wildlife in the United States 23 iill areas there are no data available to indicate the percentage of land which will be primarily devoted to that purpose. Within the national forests, tliere are approximately 8,000,000 acres of land classed as "barren" on which some forms of wildlife are found. Tliis area, so far as can be seen, will serve no other purpose than the pro- duction of wildlife. ^Miile its classification would indicate lack of vegetative cover, this is not entirely true since within the cracks and crevices and along the narrow ledges on the faces of almost perpendicular cliffs, and on other inaccessible areas there is ample food for the species of wildlife inhabiting such localities, such as mountain goats, mountain sheep, and ptarmigan. In addition, there are within the national forests some 10,000,000 acres of "brush land", much of which will be permanently useful only for watershed protec- tion and wildlife production. Areas Specially Adapted to Wildlife The areas of private land and public domain listed in table II, as needed for additions to national forests in regions 1 to 10, inclusive, are vital to wildlife manage- ment on tlie western national forests. They are an integral part of the winter range for many of the big game herds which depend upon national forest ranges for subsistence during tlie summer months. Hea\T' snows and severe cold drive elk and deer from the high sunmier ranges to the lower countrj-, where climatic conditions are more favorable and food may be obtained during the winter. The capacity of the national forests to produce sustained crops of big game animals is very definitely limited b}^ the availabilitj^ of winter range. Under present conditions of ownership and use, that portion of winter range in private ownership (6,000,000 acres) and in the pubhc domain (14,000,000 acres), within and adjacent to national forests, is not capable of supplying forage adequate to maintain wildlife popu- lations as large as can be supported by national forest summer ranges. These areas, totaling 20,000,000 acres, should be added to the national forests, and other uses, chiefiy grazing by domestic livestock, should be sufficiently curtailed to allow the big game herds to increase to a size comparable with the carrying capacity of the liigh national forest summer ranges. Areas Available for Wildlife All national forest land is now and may in the future be employed for the production of some form of \\-ildIife in conjunction with other uses. The following tabu- lation shows the extent of these areas, by national forest regions. Table I.- -Exlenl of iialional forest area which is available for wildlife use National forest region Net area of national for- ests and pur- chase units on June 30, 1934 I Net area game refuges on national forests ' Federal State Acres 22,791,449 19,383,134 19, 932, 106 29, 183, 676 19.3.12,839 23,121,116 2, 554, 586 5, 512, 882 4,073.985 21.342,300 Acres Acres 2, 563, 700 2. Rocky Mountain-- 195. 158 837.115 26.'776" 4.769.517 2. 224. 397 4. InUTiiiuiinlaiii , - -- 5. f-'alifornia .-, 6. North Pacific 5. 348. 396 2.019,870 3, 111,440 90.278 S. Sonthern.- - 9. North Central- --- - 249. 132 2.671 2, 697, 225 251.750 864, 193 10. Alaska Total 167.248,073 4,002,071 21, 243, ,541 1 Includes land approved for purchase under the Weeks and Clarke-McNary laws. ' Included in net area national forests and purchase units. The 167,000,000 acres of national forest land, located in 33 States, Alaska, and Puerto Rico, is all available for wildlife. All of this land, except the 25,000,000 acres of Federal and State game refuges, is open for hunting and fishing, subject to the laws of the several States. Hunting is also permitted on some of the Fed- eral refuges, under special regulations. Almost 3,000,000 acres have been closed to use by domestic livestock, in the interest of wildlife production. These areas, however, may be used for other purposes, such as timber production and recreation. This illus- trates the policy of correlated land uses, a prmciplc which can be applied in safeguarding the wildlife re- source. Strictlj^ speaking, therefore, there are no areas in the national forests devoted exclusively to wildlife, but future studies may reveal the necessity for dedi- cating areas to only one use. Needed Additions to Forests The Forest Service believes that, in the interests of the production and management of wildlife, three classes of additions to the national forests are essential: (1) Completion of the purchase program in the East, within purchase units approved by the National Forest Reservation Commission; (2) purchase of certain es- sential areas of ^vinter game range within and adjacent to the national forests; and (3) transfer of certain areas of winter game range within the public domain to adjacent national forests. These recommended ad- ditions are shown in table II. Of the 34,000,000 acres needed for additions to the national forests, all but 14,000,000 acres of pubhc do- main will, if acquired, be devoted to wildlife in associa- tion with other uses. The pubhc-domain lands are urgently needed for speciahzed whiter range use by big- game animals summering on the national forests. These lands, if added to the national forests, will be so administered that the winter range for big-game animals, chiefly elk and deer, will be greatly enlarged 24 Land Planning Report Table II. — Needed additions to the national forests for production and management of wildlife Table III. -Privately owned lands within national forest and purchase unit boundaries National forest region Areas needed to complete purchase program on approved purchase units Areas of winter game range needed for ad- ditions to national forests Total needed additions to national forests Private land Public domain Acres Acres 471,893 3, 482, 535 Acres 123, 732 5.925,460 1, 367, 000 4, 125. 329 1,219,093 1,431,000 Acres 595,625 2 Rocky Mountain 9, 407, 995 1,367,000 4, IntArmnnntnin 868,491 451, 380 385,800 4, 993, 820 1, 670, 473 6. Northwestern 1,816,800 3.042,924 6,504,702 4,115,433 3, 042, 924 6, 504. 702 9 North Central 407, 080 4, 522, 513 10. Alaska Total 13,663,059 6,067,179 14, 191, 614 33,921,852 and improved. (See map sho^ving winter range and game-refuge areas, fig. 2.) Privately Owned Land As a Habitat for Wildlife Two classes of land are of interest to the Forest Service in connection with the discussion of the use of private lands for the production of wHdhfe: (1) Pri- vately owned lands intermingled ■with national forest lands, and (2) privately owned lands intermingled with the public domain. Table II hsts the areas in class (1). This 37,000,000 acres of privately owned land witliin the exterior boundaries of national forests and pur- chase units includes the 19,000,000 acres listed in columns 1 and 2 of table no. II as land needed for additions to the national forests. The remaining 18,000,000 acres will largely remain in private owner- ship. These lands are, by and large, of similar value and importance in wildlife production as are adjacent national forest lands. That is, on these lands wildlife may be coordinated with other private land uses, such as grazing, agriculture, timber production, mining, Lands within Lands within national national forest and forest and National forest region purchase unit boundaries, National forest region purchase unit boundaries. in private in private ownership as ownership as of Junes, 1934 of June 3, 1934. Acres Acres 1. Northern 3,768,837 7. Eastern . 4,112,912 2. Rocky Mountain 1,831,473 8. Southern 8. 485, 279 3. Southwestern 2, 085, 575 9. North Central 6. 546. 515 1, 594, 189 4, 857, 503 3,972,889 10. Alaska 54, 633 6 Northwestern Total 37, 309, 805 and recreation. The degree to which these inter- mingled private lands will produce wildlife is de- pendent upon the emphasis placed by the owners on this use as compared with other uses, and upon the degree of control or wildhfe management exercised thereon. Within western game ranges, but outside national forest boundaries, there is an estimated total of 55,- 000,000 acres of land, of which 13,000,000 acres is public domain on which the wildlife use should be considered of high priority. An additional 8,000,000 acres of public domain is less important for wildlife usages, but necessarj'' to a full coordination of uses both inside and outside national forests. The situation with respect to tliis 55,000,000 acres of intermingled private and pubhc domain lands is quite analogous, in at least one respect, to the 167,000,000 acres of national forests, with 37,000,000 acres of inter- mingled private holdings. The 55,000,000 acres is largely suited to the production of wildhfe, in coordina- tion with other land uses. A high degree of integration of uses, fully coordinated and sustained long-time programs, and definite and decisive execution of plans are necessary to bring about complete development and utiUzation of the wildlife resources on these lands. o