nester, N. H. a DR. ARNOTT. THE POPULAE SCIENCE MONTHLY. CONDUCTED BY E. L. YOU MANS, VOL. X. NOVEMBER, 1876, TO APRIL, 1877. NEW YORK: D. APPLETON AND COMPANY, 54 9 & 551 BROADWAY. . 1877. Copyright by D. APPLETON AND COMPANY, 1877. lO + lk THE POPULAR SCIENCE MONTHLY. NOVEMBER, 1876. WHAT AMERICAN ZOOLOGISTS HAVE DONE FOR EVOLUTION.1 Br Professor EDWAED S. MOKSE. I. IT would be pleasant indeed if only a lecture or an essay were ex- pected from the presiding officer of the Section ; but an address implies a great deal more, and the giver of it is not only expected to be entertaining, where perhaps he never entertained before, but in- structive upon grounds upon which, perchance, he has made but par- tial survey. Among the many questions of sustaining interest, a number of subjects intrude themselves. A general review of the work accomplished since the last meeting of the Association would seem an appropriate subject for discourse. Yet beyond my special studies I feel quite incompetent to scan so broad a field. In this year of Centennial reviews, one might naturally fall into an attempt to sketch the growth of science and the work accomplished within the last hundred years, but that would not only be too vast a field, but would on the whole be unprofitable, since time-boundaries, like the surveyor's lines bordering a State, have no definite existence in Na- ture. The natural boundaries of oceans and sierras do indeed isolate and impress peculiarities of thought and action upon man, as upon the creatures below him, and for this reason we may with propriety ex- amine the work of our nation in any line of investigation. Never be- fore has the study of animals been raised to so high a dignity as at present. While chemistry could point to its triumphs in the arts, and geology to the revelations of hidden wealth in the rocks, zoology was for the most part a mere adjunct to geology, or a means to thwart the 1 An address delivered at the meeting of the American Association for the Advance- ment of Science. Read at Buffalo, New York, August, 1876. By Edward S. Morse, Vice-President Biological Section. roL. x. — 1 2 THE POPULAR SCIENCE MONTHLY. ravages of insects. Now, however, it is the pivot on which the doc- trine of man's origin hinges. The worlds themselves are too old to study, though tbe spectroscope reveals the existence of celestial pro- toplasm as their physical basis. The rocks are too rigid and the time too immense to come within tbe compass of our minds, but the living facts of evolution are with us to-day in these graceful forms and their constant changes, while the records more or less preserved in past times give us a clew to things hinted at in the earlier changes of present ex- isting forms. It seems, therefore, at the present time, that a review of the work accomplished by American students for the doctrines of natural selection might be acceptable for several reasons, and first among them might be mentioned the fact that thus far no general re- view of the kind has been made; and, secondly, that with few excep- tions all the general works upon the subject are from English or Ger- man sources, and filled with the results of work done there oftentimes to the exclusion of work done elsewhere. The oft-repeated examples in support of the derivative theory belong to Europe. The public are fa- miliar with these facts only, and come naturally to believe that these examples alone exist, and from their remoteness do not carry the weight of equally or perhaps more suggestive facts which lie concealed in the technical publication's of our own societies. A review of the work accomplished by American students bearing upon the doctrine of descent must of necessity be brief. Even a review of a moiety of the work is beyond the limits of an address of this nature. And for obvious reasons I must needs here restrict it to one branch of biology, namely, zoology. For material, the scientific publications of the coun- try have been scanned, and an attempt has been made to bring to- gether the more prominent facts bearing upon natural selection. In this review the zoological science of the country presents itself in two distinct periods : The first period, embracing as to time-limits the greatest portion, may be recognized as embracing the lowest stages of the science ; it included among others a class of men who busied them- selves in taking an inventory of the animals of the. country, an im- portant and necessary work to be compared to that of the hewers and diggers who first settle a new country, but in their work demanding no deep knowledge or breadth of view. And so the work to be done in tabulating the animals has more often been done by specialists who neither knew nor cared to know the facts lying beyond the limits of their studies ; a work often prompted by the same spirit that one sees among children in the collection of birds'-eggs and postage-stamps. The workers in this class were compared by Agassiz to those who make the brick and shape the stone for the edifice, an indispensable work, but with it was raised not the edifice but an almost insuperable barrier against the acceptance of views more in accordance with rea- son and common-sense. So thoroughly interwoven with this work were certain conceptions believed to be infallible, that overpowering AMERICAN ZOOLO GIS TS AND EVOL UTION. 3 indeed has been the argument to render as coadjutors the very men who so thoroughly opposed Darwin at the outset. It seems unneces- sary to point out the mode of work adopted by the class above de- scribed. Their honor involved as soon as their name had been at- tached to a supposed new species, and any deviation from the type oftentimes persistently overlooked, what wonder, when every local variety received a new name and that name stamped upon a supposed valid creation — what wonder, I repeat, that whole groups of animals have been so thoroughly scourged by such work that few have the courage to engage in the task of revision ? Emerson's reflections on the science of England in 1847 would apply with far more propriety to our country even at a much later date, where in his words " one hermit finds this fact and another finds that, and lives and dies ignorant of its value." With the noble ex- amples of Dana, Wyman, Leidy, and Burnett, before them, they did not profit. In fact, the labors of these honored men, and early in the century Lesueur and others, gave the country its largest claim to recognition abroad. The second period dates from the advent of Agassiz in this country. With his presence a gradual but entire change took place. He rendered the study a dignity rather than a pastime, No longer were the triflers to fling their loose work before the academies unrebuked. The protests he uttered in this Association were the means of elevating the tone of the communications. In fact, nothing indicates the poverty of our attainments in zoology more than an examination of the volumes preceding Agassiz's presence and the succeeding volumes. With his honest repudiation of all that was bad, he frightened away the lighter chaff, and there was but little solid work left to take its place. Agassiz made men, and his example, and the methods of work taught by him, spread to other parts of the country. He brought the American student into intimate ac- quaintance with the classical work of European naturalists. In his public lectures the names of Cuvier, Von Baer, Leuckart, and others, became familiar. The public caught the enthusiasm of this great teacher, and money was lavishly given by the citizens and the State in aid of his scientific undertakings. Agassiz's earnest protest against evolution checked the too hasty acceptance of this theory among American students. But even the weight of his powerful opposition could not long retard the gradual spread of Darwin's views ; and now his own students,- last to yield, have, with hardly an exception, adopted the general view of derivation as opposed to that of special creation. The results of his protest have been beneficial in one sense. They have prompted the seeking of proofs in this country, and now our students are prepared to show the results of their work in evidence of the laws of progressive development, and it is mainly this work that I wish to review. So much is claimed for birthplace that, in the way of history, it may not be amiss to call attention to 4 THE POPULAR SCIENCE MONTHLY. the fact that the first clear premonition of the theory of natural selec- tion came from this country. William Charles Wells, born in this country, at Charleston, South Carolina, in 1757, in a paper read before the Royal Society, in 1813, first substantially originated the theory to account for the black skin of the negro. He limits his application to races of men and certain peculiarities of color, correlated with an immunity from certain dis- eases ; in proof of it he cites domesticated animals, and the selection by man in precisely the same line of argument urged by Darwin. In the preface to the last edition to the " Origin of Species," Darwin refers to Wells's essay as entitled to the credit of containing the ear- liest known recognition of the principle. Dr. Wells first shows that varieties among men as among animals are always occurring, and having cited the way in which man selects certain qualities among domesticated animals and thus secures different breeds, -calls atten- tion to the well-known fact that the black as well as the white races are differently affected by certain diseases of the countries which they inhabit. He finds a coincidence between the immunity from certain diseases and the black color of the skin, though why this is so he does not attempt to explain. He thinks that, through the suc- cessive survival of dark skins, the dark variety of the human race has become fixed. Referring to the man's selective action regarding domesticated animals, he says : " But what is here done by art seems to be done with equal efficacy, though more slowly, by Nature, in the formation of varieties of mankind fitted for the country which they inhabit." These sentences have such a Darwinian sound that, when we remember they were dragged from obscurity by Mr. Darwin him- self, we can share in what a recent writer ' happily calls " Mr. Dar- win's evident delight at discovering that some one else had said his good things before him, or has been on the verge of uttering them." As early as 1843, Prof. Haldeman2 discussed some of the arguments brought forward by the opponents of the Lamarckian theory, and offered certain views in favor of the transmutation of species. While he does not hint at the laws of natural selection, he recognizes fully the value of varieties and their persistency and ultimate divergence. He says, "Although we may not be able artificially to produce a change beyond a given point, it would be a hasty inference to suppose that a physical agent acting gradually for ages could not carry the variation a step or two farther, so that instead of the original one we will say four varieties, they might amount to six, the sixth being suf- ficiently unlike the earlier ones to induce a naturalist to consider it distinct." In the year 1850, Dr. Joseph Leidy, in a paper on entophyta in living animals, wrote as follows: " The essential conditions of life are 1 Gray's " Darwiniana," p. 284. 2 Journal of the Boston Society of Natural History, vol. iv., p. 368. AMERICAN ZOOLOGISTS AND EVOLUTION. 5 five in number, namely: a germ, nutritive matter, air, water, heat, the four latter undoubtedly existing in the interior of all animals." 1 Dr. Leidy affirms his belief that very slight modifications of these essen- tial conditions of life were sufficient to produce the vast variety of liv- ing beings upon the globe. The theory of derivation based upon the principles of natural selection demands the following admissions : that species vary, that peculiarities are transmitted or inherited, that a greater number of individuals perish than survive, and that the physi- cal features of the earth are now and have been constantly changing, and that precisely the same conditions never recur. These are ad- mitted facts. Now comes the theoretical part of natural selection, namely, that the varieties which survive are those which are more in harmony with the environments of the time. These propositions, with minor ones, form the theory of Darwin. Lamarck and others had recognized the gradual enhancement of varieties into species, but had not struck the key-note of natural selection, though Wells in the beginning of the century had clearly recognized it in a pertinent example. If we look impartially at these propositions, we need no demonstration to prove the inheritance of characters the most minute, and even the perpetuation of the most subtile features. On general principles, too, the proposition, that those individuals best adapted to their surroundings survive, need only be stated to be accepted by a reasonable mind. In truth, to deny it would be to deny, as Alphonse de Candolle says, that a round stone would roll down-hill faster and farther than a flat one. Indeed, this eminent botanist affirms that natural selection is neither a theory nor an hy- pothesis, but the explanation of a necessary fact. The constant physi- cal changes in the past and present condition of the world are incon- trovertibly established. It seems, then, that the prime question re- solves itself into whether each species as a whole has something inherent which prompts it to vary irrespective of its environments, or whether a correlation can be established between the variation of spe- cies and certain physical conditions inducing these variations, and here let me add that of all groups of animals from species through genera to higher divisions, that group of individuals recognized as a species has the most tangible existence. And, as a proof of this, there need only be mentioned the fact that many naturalists, while regarding species as clearly distinct, have on the other hand looked upon classification as an artificial method to facilitate the study, and hence the innumerable schemes and the successive interpolation of sub- classes, sub-orders, sub-families, and sub-genera, which simply circum- scribed smaller proofs than had before been recognized. The rapid multiplication of some of these groups has already formed a serious obstacle to the study of systematic zoology. What would good Dr. Mitchell have said if he could have foreseen 1 " Proceedings of the Philosophical Academy," vol. iii., p. 7. 6 THE POPULAR SCIENCE MONTHLY. the generic lists of to-day ! In an article on the " Proteus of Lake Erie," he expressed his aversion to multiplying names in zoology, and lamented the tendency. He protested as follows, fifty years ago : " By some, these innovations have been so wantonly introduced, as almost to threaten in the end the erection of every species into a dis- tinct genus." l Though these words were undoubtedly aimed at Rafi- nesque, they were none the less prophetic. Whatever may be said of the existence in nature, of other groups, there can be no question that species have the most definite existence, and it would seem then that nothing more need be proved for the theory of descent as opposed to the theory of special creation, than the establishment of the fact that species assume the characters of new species, or disap- pear altogether with a change of surroundings. As examples might be cited, the transplanting of Alpine seeds to warmer regions below, and an accompanying change of the plant into another species before known in the warmer region, or, more remarkable still, the change of a species of Crustacean which lives in salt water, to another species with a partial freshening of the water, and this freshening slowly persisted in, the form changing into another genus, and in so doing losing: one of its segments. In the first case we see the effect ol temperature, and in the second case the physical influence of salt anfl water in different proportions. Now, these and hundreds of similar examples can be incontestably proved. Even the prolonged existence of the form of some animals, like Lingula, may be referred to an inherent vitality which enables them to survive changes that caused the death of thousands of others. In an early discussion of Darwin's theory,2 Prof. Agassiz cited the persistence of Lingula as fatal to the theory, and Prof. William B. Pogers replied that the vital characters of some animals would enable them to survive above others. Ten years later, I had an opportunity of studying living Lingula on the coast of North Carolina, and brought specimens home alive in a small jar of water, and kept them in a common bowl for six months without the slightest care. Their power of surviving under changed conditions — their vitality, in other words — seems incredible.3 (For further details, see reference below.) It has for a long time been suspected that the species of Jfollusca, described in such profusion in this country, would be redubed when the slightest attention to their habits had been made. Dr. James Lewis4 long ago observed that a certain species of fresh-water mussel, described as Alasmodonta truncata, is only the truncate form of another species, A. marginata. From a careful study of the condi- 1 American Journal of Science and Arts, vol. vii., 1829. s " Proceedings of the Boston Society of Natural History," vol. vii., p. 231, December 15, I860. 3 Ibid., vol. xv., p. 315. 4 Ibid., vol. v., p. 121. AMERICAN ZOOLOGISTS AND EVOLUTION. 7 tions surrounding the first form in the Mohawk River, he had reason to believe that the rapid currents which pass over it bear along sub- stances that, coming in contact with the exposed edges of the shell, break them down, thus retarding the growth of the shell at this point, and the animal concentrates its growth -powers to the repairs of the broken portion. The same gentleman also shows that the so-called species Lymnana elodes, catascopium, and marffinata, " are modifica- tions of one type or species, influenced by locality and temperature varying the method of development." ' A. G. Wetherby 2 calls attention to the variation in form of a group of fresh - water snails, found in the greatest abundance in certain streams of Tennessee and North Alabama. In showing the varied influences they are subjected to he cites the rapid currents of the channels, and the greater liability of the snails being torn from the rocks. He shows that they are exposed in various ways to the effects of these currents, with all their changing impetus of high and low water — exposed also to privation of food from the scouring sand re- moving the conferva?, upon which they subsist, from the rocks. He takes into account temperature, chemical action, and the like, and says, " No greater vicissitude can be imagined than this growth in an unstable element." Coincident with these diverse conditions he finds an enormous variety of forms, and frankly acknowledges that many of those described as distinct species must be reduced to synonyms. George W. Ti-yon, in his large work on the American Melanians, published by the Smithsonian Institution, having finished his manu- script in 1865, says, under date of 1873, when the work was finally published, "A more enlarged acquaintance with fresh-water shells con- vinces me that a much greater reduction of the number of species than I have attempted must eventually be made." If we now look upon the definition of a species, as given by a gen- tleman foremost in the ranks as a describer of species, we find it formu- lated as follows : A species represents " a primary established law, stamped with a persistent form (a type) pertaining solely to itself, with the power of successively reproducing the same form, and none other ;" and this gentleman has not hesitated to base these " primary organic laws" upon the evidence of a single specimen, and in some cases even the fragments of one have offered him a sufficient induce- ment ! But it has been argued by some that a wide variation may be the case "with many species. Prof. Agassiz,3 at a meeting of the American Academy, reiterated his opinion that what are called varieties by naturalists do not in reality exist as such. He found a great abun- dance of diverging forms in Echinoderms, which, without acquaintance 1 " Proceedings of the Boston Society of Natural History," vol. v., pp. 121-128. 2 Proceedings of Cincinnati Society of Natural Science, No. 1, June, 1876. 8 " Proceedings of the American Academy," vol. v., p. 72. 8 THE POPULAR SCIENCE MONTHLY. with connecting ones, would be deemed distinct species, but he found they all passed insensibly into each other. Prof. Parsons suggested that more extended observations might connect received species by intermediate forms, no less than so-called varieties ; and Prof*. Gray remarked that the intermediate forms, con- necting by whatsoever numerous gradations the strongly divergent forms with that assumed as a type of a species, so far from disproving existence of varieties, would seem to furnish the best possible proof that these were varieties. Without the intermediate forms they would, it was said, be taken for species; their discovery reduced them to va- rieties, between which (according to the ordinary view), intermediate states were to be expected. Recognizing, then, the existence of varieties, and of varieties suffi- ciently pronounced to have led careful naturalists to regard them as distinct species, what shall we say when it is found that these marked forms are correlated with certain physical conditions, many of which have originated within comparatively recent times ? Dr. J. G. Cooper,1 after a careful study of the California land snails, ascertained that " species, sub-species, and varieties, living in cool, damp situa- tions, become more highly developed (but not always larger) than the others ; the shell assuming a more compact (imperforate) form, and losing those indications of immaturity referred to, viz., sharp, deli- cate sculpture, bristles, and angular periphery. These characteristics, however, remain more or less permanently for indefinite periods, and give that fixedness to the various forms, even when living under the same conditions, which enables us to retain them as sub-species differ- ing from varieties in permanency, and from races in not inhabiting distinct regions." It may be added that Stearns, Bland, and Binney, have likewise observed the same peculiar variations associated with aridity. In a broader field, and compassing different classes, Prof. Spencer F. Baird, Mr. J. A. Allen, and Mr. Robert Ridgway, have severally shown that marked and specific changes are seen in birds and mam- mals corresponding to differences in their surroundings. Prof. Baird, in a paper entitled " The Distribution and Migration of North Amer- ican Birds," 2 has shown that birds in high altitudes and those bred at the North are larger than those born South and at low altitudes ; that Western birds of the same species have longer tails than eastern exam- ples, and that the bill increases in size in those birds occurring in Florida as compared with those found north of that State, and that on the Pacific coast the birds are darker in color than those found in the interior. Mr. J. A. Allen3 has made a more special study of this matter, and 1 "Proceedings of the California Academy of Natural Science," vol. v., p. 128. 2 American Journal of Science and Arts, vol. xli., January and March, 1866. 3 "Proceedings of the Boston Society of Natural History," vol. xv., p. 156. AMERICAN ZOOLOGISTS AND EVOLUTION. 9 his work ranks among the most important contributions to this sci- ence. Mr. Allen finds that there are marked geographical variations in mammals and. birds. He shows that northern mammals of the same species are more thickly and softly furred, and that toward the south the peripheral parts, such as the ears and feet, are more devel- oped. The same law holds good in birds, a diminution in size being observed toward the south, and the individuals being darker in color. As one goes south he meets with the same species of birds, whose bodies are shorter, but whose beak, tail, and claws, are longer. On the Plains, also, he found the birds with plainer tints, while south- ward the colors became more intense. On drawing up a table indi- cating the regions of lighter varieties, and comparing it with a chart of mean annual rainfall, Mr. Allen found the lighter forms occurred in dry regions, and the dark forms in relatively humid regions. To sum up : Mr. Allen finds in latitudinal variation climatic influences affecting color as well as altering the size of bill, claw, and tail, while longitudinal variation usually affects color alone. He states that these laws are now so well known that a species may be predicted to assume a given color if under certain specific climatic conditions. Mr. Robert Ridgway ' has in a similar way called attention to the relation between color and geographical distribution in birds as ex- hibited in melanism and hyperchromatism, and has shown that red areas " spread " or enlarge their field in proportion as we trace cer- tain species to the Pacific coast, and that in the same proportion yel- low often intensifies in tint. The results of these investigations can be easily understood. Nearly if not quite one hundred and fifty species of birds, which were recognized as distinct, are at once reduced to varieties, though less than twelve years ago they were looked upon as good species, with which no external influence had anything to do. Nearly if not quite a fifth of the number of species of birds have been reduced by the investigations of Baird, Allen, Coues, and Ridgway. The mammals, through the same study of geographical variation, will have been reduced at least one-fourth. Already Mr. Allen2 has studied the geographical variation of the squirrels, and the result is that a reduction has been made of one-half the number of species before recognized. Prof. Baird, in his monograph of North American squirrels, reduced the number from twenty-four, as acknowledged by Audubon and Bachman, to ten well-established species and two doubtful varieties. Allen, with still greater advantage in the shape of a mass of material from the Western surveys, l-educed the ten species to five species, with seven geographical varieties. 1 American Journal of Science and Arts, vol, iv., December, 1872, p. 454, and vol. v., p. 39. 2 " Proceedings of the Boston Society of Natural History," vol. xiv., p. 276. io THE POPULAR SCIENCE MONTHLY. Should it be urged that the present tendency toward reducing species be taken as an evidence that species had not before been properly defined, then it offers a stronger argument still in favor of the fact that species are even more variable than had before been supposed, leaving the greater possibility of larger numbers of these ultimately surviving. Again, the assumption that the limitation of specific variation had not been properly indicated, shows how repre- hensible has been the work of some of those who have burdened our literature with their bad species. The fact is, the work has in a measure been justifiable, and is not' to be wholly condemned. The workers in this line have followed the teachings of their masters. A group of individuals removed from an allied group of individuals by an extra dot or darker shade, perpetuating their kind from generation to generation, marked with persistent characters, and in every way coming up to the standard recognized as specific, had the right to be judged as such. It is only when a whole series of forms are collected, and climatic influences are seen to affect these in the same way that they affect other groups of species even in different classes, that the mere influence of moisture and temperature is shown to be the sole cause of many of these supposed specific characters. Dr. A. S. Packard, in his remarkable monograph of a group of moths, the Phalcenidce, published under the auspices of the Hayden Survey, finds that with some species there are changes analogous to those pointed out by Baird and Allen ; and while he does not find enough to establish a law, yet to his mind enough is seen " to illus- trate how far climatic variation goes as a factor in producing primary differences in fauna? within the same zones of temperature," and he admits that varietal and even specific differences may arise from these climatic causes alone. Dr. Packard, in the same work, under the head of " Origin of Genera and Species," says, " The number of so- called species tends to be reduced as our specimens and information increase." The genera also " are as artificial creations as species and varieties. The work of the systematic biologist often amounts to but little more than putting Nature in a strait-jacket." An application of the influence of temperature is here proper, as explaining, on a rational ground, the persistence of peculiar arctic forms of animals and plants on the summits of Mount Washington and other high peaks. With a knowledge of glacial phenomena, we are capable of judging the condition of things which must, of neces- sity, have existed directly after the recedence of the great ice-sheet : its southern border slowly retreating, and, with the encroachment of the warmer zone, the arctic forms dying out, or surviving under changed conditions ; but, in high plateaus and mountains, local glaciers flourished for a while, and at their bases arctic forms flour- ished, and, lingering too long, were ultimately cut off by the retreat AMERICAN ZOOLOGISTS AND EVOLUTION. n of the main field. This interpretation of arctic forms on high peaks, though attended to by several American naturalists, is not new. Os- wald Heer, in discussing the origin of certain animals and plants, coincides with De Candolle that Alpine plants are relics, as it were, of a glacial epoch. Prof. Gray ' had also independently arrived at the same conclusions, based on a comparison of the plants of Eastern North America and Japan. In the position he maintained regarding the derivation of species from preexisting ones, he stood far in ad- vance of his brother naturalists in this country, for this was before Darwin's great work had appeared, and before Heer had developed the host of fossil plants from the arctic zone. Mr. S. I. Smith, in speaking of mountain faunae, points out the gradual encroachment of glaciers, and the drawing down of northern forms ; and, as the gla- ciers retreated, these forms were caught, "the mountain-summits being left as aerial islands." Dr. Packard and Mr. Scudder have severally called attention to the same thing. Prof. A. R. Grote has more fully dealt with the subject in a paper read before this Association, and in a graphic way shows that the "former existence of a long and widely-spread winter of years is offered in evidence through the frail brown OEneis butterflies, that live on the top of the mountains within the temperate zone." I have been thus explicit, in order to contrast these more rational views with those formerly entertained by eminent naturalists, whose minds were imbued at the time with the idea of special creation. Mr. Samuel H. Scudder2 read before the Boston Society of Natural History an ac- count of distinct zones of life on high mountains, as illustrated in the insect-life of Mount Washing-ton. He called attention to certain insects which he supposed peculiar to the summit, and not found farther north, though showing a remarkable correspondence to certain arctic forms. Prof. Wyman asked whether all the facts might not be ac- counted for on the theory of migration northward after a glacial epoch, and Prof. Rogers suggested that the facts might be accounted for on the migratory theory if we added thereto the supposition of subsequent variation induced by isolation. Yet these views were persistently opposed by the other naturalists present. The mass of evidence already contributed, as to the extraordinary variation in color, markings, and size of species coinciding with their physical sur- roundings, though perhaps trivial in itself, becomes important when the proofs are grouped together, and all bear upon the theory of derivation. So slight a thins; as change of food is found to influence certain animals even to a degree usually regarded specific. The late Dr. B. D. Walsh 3 discovered some very curious features among in- 1 "Memoirs of the American Academy," vol. vi., pp. 37*7-458 (1859). 3 " Proceedings of the Boston Society of Natural History," vol. ix., p. 230. 3 " On Phytophagic Varieties and Phytophagic Species," " Proceedings of the Ento- mological Society of Philadelphia," vol. iii., p. 403. 12 THE POPULAR SCIENCE MONTHLY. sects connected with a change of food. First, he established the fact that insects accustomed to one kind of plant could acquire a taste for another kind, and he has shown that in thus changing the food of the insect a change took place in the appearance of either the larva, pupa, or imago, and sometimes all three stages were affected. Dr. Fitch had observed that changing an insect's larva from the leaf to the fruit affected the appearance of the larva. It would be impossible to give even an abstract of Dr. Walsh's remarkable essay. It may be said, however, that his investigations led him irresistibly to the conclusion that the present species have been derived from preex- isting ones, and in numberless cases he is capable of showing the successive stages from the dawn of a plant-eating variety, where the changes are slightly seen in the larva only, to the plant-eating species in which profound changes are seen in the larva, pupa, and imago. The minor factors of natural selection, such as protective coloring and mimicry, have been variously illustrated by Mr. R. E. C. Stearns, Dr. Kneeland, Prof. Cope, Dr. Charles C. Abbott, and others. In a special paper on " The Adaptive Coloration of Mollusca," ' I have en- deavored to show not only a wide-spread application of this feature to mollusks, and especially those exposed by the tide, but in some cases a mimicry of inanimate objects, as the accumulation of clay or grains of sand upon the shell. Wallace's theory of birds'-nests finds interesting confirmations in the observations of Dr. Abbott, who made a special study of a large number of robins'-nests, and found the widest variation among them. He studied also the nests of the Baltimore oriole, where, according to the theory of Wallace, a concealing nest should be made, the bird being exceedingly bright-colored. He found that, away from the habitations of man, the orioles built concealing nests ; but in villages and cities, on the other hand, where they were in no special danger from predatory hawks, the nests were built comparatively open, so that the bird within was not concealed.8 The differences in the habits of animals of the same species are noticed in different parts of the country, and such facts militate against the idea that certain unerring ways were implanted in them at the outset. Indeed, such facts go to show that these various creat- ures not only become adapted to their surroundings, but that individ- ual peculiarities manifest themselves. The observations of Dr. Cones, Mr. Allen, and Mr. Martin Trippe, go to prove that certain birds change their habits in a marked degree. In their behavior, too, cer- tain birds, which are wild and suspicious in New England, are com- paratively tame in the West. In their resting-places they show wide individual variation. 1 " Proceedings of the Boston Society* of Natural History," vol. xiv., p. 141, 5 Popular Science Monthly, vol. vi., p. 481. AMERICAN ZOOLOGISTS AND EVOLUTION. 13 Prof. A. E. Verrill,1 on the supposed eastern migration of the cliff- swallow, traces historically its first appearance in various places in the East, and is inclined to the opinion that as the country became settled by Europeans the birds left their native haunts for barns and houses, and increased in number to a greater extent than before on account of the protection invariably furnished by man. Rev. Samuel Lockwood2 records a curious case of the Baltimore oriole acquiring a taste for the honey-sacs of bees, tearing off the heads of those insects, and, having secured the honey-sacs, rejecting the rest of the body. Prof. Wyman3 observes a curious case in Florida of a colt and a number of pigs and cows thrusting their heads under water and feed- ing on the river-grass, in some cases remaining with their heads im- mersed for half a minute. Hon. A. II. Morgan 4 observes the widest difference in the habits of the same species of beaver iu the Lake Superior region and in the Missouri, constructing their dams and ways differently, and meeting the varied conditions, not by a blind instinct, but by a definite intelli- gence manifested for definite purposes. All of these facts, simple in themselves, yet together go to prove that animals do vary in their habits, and with a persistent change in habits arises the minute and almost insensible pressure to swerve and modify the animal. So much does the influence of season, with its accompanying pe- culiarities of food, temperature, humidity, and the like, affect certain animals developing coincidently with its different phases, that it is instructive to note that in certain species of insects two or three dif- ferent forms occur. Thus Mr. Edwards 6 has in an elaborate way worked up the history of a polymorphic butterfly [Ephiclides ajctx), showing that there are three forms heretofore regarded as distinct species, which are only varieties of one and the same species, but ap- pearing at different times of the year, and consequently confronted by different influences as to temperature, moisture, food, and the like. These forms are known under the names of Walshii, Telemonides, and JIarcellus, and both sexes are equally affected. The first form mentioned represents the early spring type, Telemonides the late spring type, and Marcellus the summer and autumn type {see also Mr. Scudder's paper 6). If these influences affect species, we should expect to see the greatest variety of forms in a country possessing the widest diversity of conditions. Some suggestive paths of investigation have been pointed out by 1 " Proceedings of the Boston Society of Natural History," vol. ix., p. 276. a American Naturalist, vol. vi., p. 721. 3 Ibid., vol. viii., p. 237. 4 " " The American Beaver and his Works." 6 " Butterflies of North America," part ix. 6 American Naturalist, vol. viii., p. 257. 14 THE POPULAR SCIENCE MONTHLY. Prof. N. S. Shaler ! on the connection between the development of the life and the physical conditions of the several continents, showing first that the greatest amount of shore-line in proportion to the internal areas indicates a greater diversity of surface within. Another proposition he attempts to establish : that in propor- tion to the shortness of the shore-lines, or, in other words, to the want of variety in their surfaces, will be the diversity of animal life in the continent. He then proceeds from Darwin's standpoint, and follows out many curious and instructive lines of thought regard- ing increased amount of influences in diversified surfaces — a level plain having the same conditions throughout, but a mountainous re- gion having for each one thousand feet of elevation a new condition of things, in the form of streams, winds, humidity, and the like. In areas of simple outline and unvarying surfaces wTe do, in fact, have a less diversity of forms. Recognizing the mutation of continents through past geologic ages, we again see the accompanying physical changes in not only modify- ing forms, but in selecting them afterward by succeeding changes. The widely-diversified nature of the facts bearing on the doc- trine of natural selection baffles all attempts at a systematic classifica- tion of them. Of such a nature are many of the valuable communi- cations of Prof. Wilder. At the meeting of this Association 2 he has, among other matters, confirmed in a young lion the discovery of Prof. Flowers that, in the young dog and probably in other carnivora as well, the scapho-lunar bone has at the outset three centres of ossification, and that these really represent the radiate intermedium and centrale of the typical carpus. By study of a foetal manatee, Prof. Wilder is able to de- termine its affinities, and to point out the probable retrograde meta- morphosis of some ancient ungulate animal, and that the mana- tee is widely removed from the whales Avith which it has been associated. Mr. William K. Brooks has published a very remarkable paper on certain free swimming tunicates, the /Salpa, giving for the first time a clear and comprehensive history of certain ohscure points, and has at the same time applied the principles of natural selection theoreti- cally in showing the origin of salpa from sessile tunicates, and mak- ing clear the peculiar modification of parts which accompany these changes. In the field of entomology some capital work has been done, both practical and theoretical. Prof. Riley's demonstration of the yucca-moth is unique in its way. Dr. Engelmann has discovered that the yucca depends upon insects for fertilization ; and Prof. Riley, by patient study, not only 1 " Proceedings of the American Academy," vol. viii., p. 349. 2 " Proceedings of the American Acc.demy of Arts and Sciences," vol. xxii., p. 301. AMERICAN ZOOLOGISTS AND EVOLUTION. 15 discovered the moth which fertilizes the flower, but finds an anoma- lous change in the maxillary palpi of the insect, by means of wbich the moth collects bundles of pollen, which it inserts into the stigmatic tube, and during this peculiar act deposits her eggs in the young fruit. Prof. Riley has reasons to believe that this is the only insect engaged in the fertilization of this plant. A mutual dependence is here met with of extreme interest. The yucca unfertilized forms no fruit, and the larva of the moth consequently perishes. Prof. Augustus K, Grote, in an examination of butterflies, finds successive gradation in their structures, and shows that as these or- gans " become less serviceable to the insect they become more rigid and in position more elevated above the head in the butterfly, while in the moth they are more whip-like and directed forward." While protesting against the separations which have been made in the order based upon the antenna?, he directs attention " to the real differences in antennal structure between the butterflies and moths, while show- ing that the antenna? are modified by desuetude in the higher and former group." Prof. Grote,1 in dealing with a family of moths, the Noctuidce, calls attention to the unequal value of Acronycta, and is forced to admit that these differences become clear through the theory of evolution. He says : " Where in Acronycta there is a gen- eral prevailing uniformity in the appearance in a single group of spe- cies and generally broad distinctions between the larval forms, it is a not unreasonable conclusion that these larval differences are gradu- ally evolved by a natural protective law, which intensifies their char- acters in the direction in which they are serviceable to the continu- ance of the species." Those who have believed in types as fixed laws, rigidly impressed at the outset of life, are those also who have recognized in the cells of a honey-bee, as well as in the arrangement of leaves about the axis of a plant, a perfect mathematical adjustment of parts, which were stamped at the beginning, and have so continued to exist with- out deviation. For nearly two hundred years it has been believed that the instinct of a bee guided it to shape a cell which of all other forms should use the least amount of material. A theory having been established as to the constant shape of a bee's cell, namely, that it was an hexagonal prism with trihedral bases, each face of the base being a rhomb with certain definite angles, a mathematician wTas given the problem to construct similar cells, and to determine the best possible form with the use of the least amount of material. The coincidence between theory and observation and experiment was so remarkable as to settle apparently for all time the question as to the perfectly-implanted instinct of the bee with its unconscious power of accurate work. Prof. Jeffries Wyman,2 to whose memoir I am indebted 1 " Proceedings of the Buffalo Society of Natural Science," vol. i., p. 130. 2 " Proceedings of the American Academy," vol. vii., p. 68. 16 THE POPULAR SCIENCE MONTHLY. . for the above facts, has by an ingenious study of the cells of bees shown, first, that, a cell of this perfection is rarely if ever attained. Further- more, that, while the honey-cells " are built unequivocally in accord- ance with the hexagonal type, they exhibit a range of variation which almost defies description ; " that the worker-bees, from incorrect alignment and other causes, build cells, the measurement of which shows the widest limit of variation ; that the drone-cells are liable to substantially the same variations, while the transition-cells, namely, those in which drones and worker-cells are combined in the same piece of comb, are extremely irregular. As the drone-cells are one- fifth larger than worker-cells, " a transition cannot be made without some disturbance in the regularity of the structure." And Prof. Wyinan states distinctly that the bees do not have any systematic method of making the change, adding that " the cell of the bee has not that strict conformity to geometrical accuracy claimed for it," and the assertion, like that of Lord Brougham, that there is in the cell of the bee " perfect agreement between theory and observation, in view of the analogies of Nature, is far more likely to be wrong than right, and his assertion in the case before us is certainly wrong." Prof. AYyman closes his essay by saying that " much error would have been avoided if those who have discussed the structure of the bee's cell had adopted the plan followed by Mr. Darwin, and studied the habits of the cell-making insects comparatively, beginning with the cells of the humble-bee, following with those of the wasps and hornets, then with those of the Mexican bees, and finally with those of the common hive-bee ; in this way they would have found that, while there is a constant approach to the perfect form, they would at the same time have been prepared for the fact that even in the cell of the hive-bee perfection is not reached. The isolated study of any- thing in Nature is a fruitful source of error." The remarkable ingenuity, so characteristic of Prof. Wyman's ex- periments, is fully shown in this memoir. He made plaster-casts of the comb, and then sawed transverse sections, and by slightly heat- ing the plaster the wax was melted and absorbed, leaving the deli- cate interspaces representing the partitions. From these sections electrotypes were taken, and thus the veritable figures were used to illustrate the absolute structure of the comb. The results of these brilliant researches were published in the " Proceedings of the Amer- ican Academy of Sciences." \To be continued.] THE EARLY HIS TOBY OF FIRE. i7 THE EAKLY HISTORY OF FIRE.1 Br Professor N. JOLY, OF THE FACULTY OF SCIENCES, TOULOUSE. FIRE, the common source of heat, of light, and of life, and the active principle of a multitude of industries, and of metallurgi- cal industry in particular, is unquestionably one of the greatest con- quests achieved by man over Nature. The discovery of fire was more than a benefit ; it was, in fact, a giant stride on the road to civilization. With fire arose sociability, the family, the sacred joys of the domestic hearth, all industries, all arts, together with the wonders they have produced, and still pro- duce from day to day. Hence w'e can readily understand how it is that fire has ever been and still is, among many nations, the object of a special worship (priests of Baal, Ghebers, Hindoo Brahmans, Roman vestals, priestesses of the sun in Peru, etc.) ; and that it has often fig- ured in the relioious or funereal rites of nations most remote from one another, both in time and space, as the Chaldees, Hebrews, Greeks, Romans, Peruvians, Mexicans, etc. But how and when was this great discovery made, in the absence of which we can hardly conceive of the possibility of human arts or even of human existence ? Did man, as we are told in the myths of India and Greece, steal fire from heav- en ; or did he, as other legends affirm, take advantage of spontaneous forest-fires, arising from the violent rubbing together of dry branches under the action of the wind ; or, finally, was man so ingenious, even from the beginning, as to devise one of those simple and practical contrivances by means of which certain savage and half-civilized tribes in our own time obtain the fire they need for their daily uses ? However far back we may trace man's history, we find him always in possession of fire. The story of Prometheus getting fire from Olympus is nothing but the Vedic myth which tells of the god Agni, or heavenly fire (Latin, ignis), as squatting in a hiding-place whence he is compelled by Matarichvan to come forth in order to be com- municated to Manu, the first man, or to Bhrigu (the shining one), the father of the sacerdotal family of the same name. The very name of Prometheus is of purely Vedic origin, and calls to mind the process employed by the ancient Brahmans in getting the sacred fire. For this they used a spindle called rnatha or pramatha, the prefix pra adding the idea of taking by force to the signification of the root matha ; this latter is from the verb mathndmi, or manthdmi — " to bring out by friction." Prometheus, therefore, is the one who dis- covered fire, brings it forth from is hiding-place, steals it and gives it 1 Translated from the French by J. Fitzgerald, A. M. vol. x. — 2 18 THE POPULAR SCIENCE MONTHLY. to mankind. From Pramantka, or Pramathyus — " lie who hollows out by rubbing," "he who steals fire" — the transition is easy and natural, and there is only one step from the Indian Pramathyus to the Prome- theus of the Greeks, who stole the heavenly fire to light the spark of life, the soul, in the clay-formed man. The spindle or pramantha had wound round it a cord of hemp mixed with cow-hair, and with this cord the priest of Brahma gave it an alternating rotary motion from right to left and from left to right. In rotating the spindle, one end of it rested in a depression made at the intersection-point of two crossed pieces of wood, the ends of which were bent to a right angle, and firmly secured with four bronze nails, thus preventing them from moving. The entire appa- ratus was called sioastika.1 The father of the sacred fire was named Twastri, i. e., the divine carpenter who made the sioastika and the pramantka, the mutual rubbing of which together produced the divine babe Agni. Its mother was named Maya. Agni took the name of Akta (i. e., anointed, christos) after the priest had poured on its head the soma, and on its body the purified butter of the sacrifice. In his interesting work on the " Origin of Fire," Adalbert Kuhn gives to the r-M and to this other like sign, r^H the name of arani, and both of them he regards as the religious symbols, par excellence, of our old Aryan ancestors — the symbols of sexual reproduction. This fire-myth occurs also in the Zendavesta, or sacred book of the Persians, and in the Vedic hymns of the Hindoos, under a two- fold form, both material and metaphysical. But the authors of these hymns bear witness that this same myth was, long before their time, symbolized in a great national religion, the founder of which, Rhibu, is no other than Orpheus. This tradition, common to Greeks, Hin- doos, and Persians, carries us back to those ancient times when the as yet undiscovered branches of this stock wandered upon the banks of the Oxus. In his " Researches into the Early History of Mankind," Tylor gives interesting details about the discovery of fire, and the various modes of obtaining it in every age. The primitive method of all would seem, according to him, to have consisted in rubbing together two pieces of dry wood, but this process was perfected in the course of time. Thus, friction is produced by means of a stick which is made 1 It is well worthy of note that the swastika, !_L1 , of India occurs very frequently in two forms, viz., !ZLJ and w o "P, on the earthen-ware disks found in such great numbers I 1— L'-T1 by Dr. Schliemann among the ruins of ancient Ilium. From this it would seem to follow that the Trojans were of Aryan origin. As for the analogies, or even direct resemblances, between certain ceremonies to the worship of Agni and certain rites of the Catholic worship, they, too, may be explained, at least to some extent, by community of origin : Agni, as Akta, would be Christ ; Maya, the Virgin Mary; Twastri, Saint Joseph. THE EARLY HISTORY OF FIRE. 19 to slide rapidly to and fro upon a piece of dry, soft wood laid upon the ground (in Tahiti, the Sandwich Islands, New Zealand, Timor, etc.). This process Tylor denominates the stick-and-groove (Fig. 1), but the fire-drill (Figs. 2 and 4) is more generally used. In its simplest form, the fire-drill consists of a stick, one extremity of which is inserted in Fig. 1.— The Stick-and-Groove. (Tylor.) Fig. 2.— The Fiee-Drill. (Tylor.) a hole bored in a piece of dry wood, while the stick itself is twirled between the hands and pressed downward {see Fig. 2). This instrument occurs not only in Australia, Sumatra, the Caro- line Islands, and Kamtchatka, but also in China, South Africa, and North and South America. It was employed by the ancient Mexicans, and is still in use among the Yenadis of Southern India, and the Veddas of Ceylon (Fig. 3). ^l^^T^^^^ Fig. 3.— Ancient Mexican Fire-Drill. (Tylor.) It is still further modified by causing the stick to whirl by means of a thong wound round it, the ends of which are pulled in opposite directions alternately. This is the instrument described in the Ve- das, and it is still employed by the Brahmans of our own day for lighting the sacred fire. For, as Tylor well observes, we very often see fire obtained for use in religious rites by the ancient processes, 20 THE POPULAR SCIENCE MONTHLY rather than by the readier means discovered in later times. Thus, when the vestals permitted the sacred fire to go out, it was rekindled by means of the sun's rays, concentrated by a lens. A similar method was employed by the ancient priests of Peru in kindling the sacrificial fire. Fig. 4— Fike-Dkill of the Oauchos. (Tylor.) An instrument resembling that employed by the Brahmans of India is to this day in use among the Esquimaux and the Aleuts (Fig. G). It consists of a rod, one end of which fits into a mouth-piece, and the other into a hole in a piece of dry wood. The rod is twirled by means of a thong wound twice around it, and pulled to the right and left alternately by the hands. Slight modifications occur in the form of the fire-drill, and various instruments have been devised to serve the same purpose. For in- stance, there are the bow-drill and the pump-drill, which latter is used both for obtaining fire and for boring holes in wood, stone, and metal (Figs. 5 and V). Fig. 5.— Bow-Drill, used bt the Sioux. (Tylor.) Of other means of procuring fire we will simply mention, in passing, the striking together of flints, or flint and steel, or iron pyrites; strik- ing together two pieces of bamboo (this method peculiar to China) ; compressing air in a tube of ivory or of wood (a process adopted by the Malays, etc.). THE EARLY HISTORY OF FIRE. 21 The dried parenchyma of the Boletus igniarius, frayed cedar-bark, dry leaves, carbonized vegetal fibres, and the like, are the combustible materials commonly employed to receive the spark produced by friction or by concussion. Is there or has there ever been a people absolutely ignorant of the means of producing fire ? Many authors answer this question affirm- atively. Thus we are told that the natives of Tasmania, though ac- quainted with fire and making use of it, nevertheless are ignorant of the means of producing it. Hence it is the special duty of their women to carry fire-brands that burn day and night, and by the light of which the tribe make their way through the woods. If the torches or brands go out, it may be necessary to make a long journey in order to have them lighted again from the fire kept up by another tribe. Nearly every family, too, carries about a cone of banksia, which burns slowly like amadou. Fig. 6.— Esquimau Thong-Drill. (Tjlor.) Fig. 7.— Pump-Drill. (Tylor.) That the Australians are not so ignorant of the uses of fire as they are said to be, is shown by a legend current among them about the origin of fire. This legend we copy from Wilson, who, in his work, " Prehistoric Man," devotes a highly-interesting chapter to the ques- tion we are considering : " A long, long time ago, a little bandicoot ' was the sole owner of a fire-brand, which he cherished with the great- est jealousy. So selfish was he in the use of his prize that he obsti- nately refused to share it with the other animals. So they held a general council, where it was decided that the fire must be obtained from the bandicoot either by force or strategy. The hawk and the pigeon were deputed to carry out thts resolution ; and, after trying to induce the fire-owner to share its blessings with his neighbors, the pigeon, seizing, as he thought, an unguarded moment, made a dash to obtain the prize. The bandicoot saw that affairs had come to a crisis, 1 A small, sharp-nosed animal, not unlike the Guinea-pig. 22 THE POPULAR SCIENCE MONTHLY and, in desperation, threw the fire toward the river, there to quench it forever. But, fortunately for the black man, the sharp-eyed hawk was hovering near, and, seeing the fire fall into the water, with a stroke of his wine: he knocked the brand far over the stream into the long, dry grass of the opposite bank, and the flames spread over the face of the country. The black man then felt the fire, and said it was good." ' Did prehistoric man possess fire ? If we are to believe the Abbe Bourgeois, man was in possession of fire since Miocene times. This assertion rests upon the discovery in the sands of the Orleanais of a fragment of artificial paste 2 mixed with charcoal, and lying in the midst of mastodon and dinotherium bones. It also rests, but not so firmly, upon the discovery by the same savant of cracked flints in the neighborhood of Thenay, not far from the banks of the lake of Beauce. These flints appear to bear plain traces of the action of fire ; but may not these be due to lightning ? If not, where are the ashes, where the charcoal which naturally would accompany the flints if they had been really brought under the action of fire ? Then, where is the fire- place ? Hence, the Abbe Bourgeois's deduction is not an impossible one, though in my opinion it is by no means demonstrated. • But, though the discovery of fire in Miocene times may be ques- tioned, it cannot be denied that in the earliest Quaternary times this element was known to man. Several fireplaces, ashes, charcoal, bones, either entire or partly calcined, fragments of coarse pottery black- ened by smoke, and similar objects, have been found in caverns be- longing to the epoch of the Cave Bear, and of the Reindeer and the Polished Stone age. These things prove that the men who inhabited the caves commonly enough cooked their food, thus making it more readily digestible. With the aid of fire, prehistoric man cremated his dead, hollowed out his pirogue, and saved from too rapid destruction the lower extrem- ity of the piles on which he built his lake-dwellings. And not only did the troglodyte and the lacustrian know how to cook their food and warm their habitations, but they also were acquainted with vari- ous methods of lighting them during the darkness of night. There have been found in the Lake of Geneva carbonized sticks of resinous wood, which, in all probability, once were employed for this latter purpose. Just as the Esquimaux now light their snow huts by means of lamps fed with the oil of the porpoise or the whale, so did the Danes of the kitchen-middens use, for illuminating purposes, a wick made of moss, one end of which was introduced into the stomach of a great penguin (Alca impennis) filled with fat. The use of flint, quartz, and iron pyrites, in the Lacustrian period, for procuring fire by striking these substances against one another, is 1 Vol. i., p. 139. 2 Paste, the mineral substance in which other minerals are imbedded. — Webster. PHYSICAL SCIENCE IN ENGLISH SCHOOLS. 23 proved by the discovery, in the lakes of Switzerland (at Meilen, Moos- seedorf, Wangeu, and Robenhausen), of bits of tinder prepared from the Boletus igniarius. And, if we accept the views of Messrs. Ed. Lartet and Christy with regard to certain blocks of granite, hol- lowed out in the centre, which have been found in the caves of Peri- gord, these blocks would appear to have been intended for the pur- pose of procuring fire by rapidly rotating a wooden rod in the central cavity, as is done by the priests of Brahma. And, indeed, how could it happen that fire should have been un- known even in the earliest periods of Quaternary time, considering the chances of fire being struck from these flints, whether in the workshop or in battle, and of the sparks falling upon combustible materials — for instance, dry leaves ? This explanation we hold to be simpler and more natural than the other, which refers the discovery of fire to the spontaneous conflagrations of forests, or to the friction of dry branches of trees. Fire, we repeat with profound conviction, must have been very early known to man, for we cannot conceive of his living without it. And hence, " who can picture the joy, the gladness, the radiant ec- stasy of that one of our unknown forefathers who first triumphantly exhibited to his astonished tribesmen the smoking brand from which he had succeeded in causing flame to burst forth ? " ' Thus we have seen that fire gave rise to nearly all the arts, or at least promoted their development. Metallurgy, architecture, ceramic arts, agriculture, navigation, commerce, industry, all are quickened by its vivifying flame. It has played, and still does play, an impor- tant part in the religious ceremonies and the funereal rites of nations, both savage and civilized. But then, in turn, as though by a law of Fate, evil accompanies the good : fire destroys with greater rapidity than it produces by forging those formidable engines, those imple- ments of death, by which in the twinkling of an eye the flower of nations is cut down on the battle-field. — Revue Scientifiqtie. ♦ «» PHYSICAL SCIENCE IN ENGLISH SCHOOLS. AT a meeting of the British Association five years ago, the subject of science-teaching in our higher schools excited unusual inter- est. Not only were papers read and followed by enthusiastic discus- sion, but a committee was privately formed, including more than twenty leaders of the Association, all of whom undertook to combine in pressing the claims of science on our head-masters, and in offer- ing counsel as to systems and methods, apparatus, and expenditure. 1 Albert Reville, Revue des Deux Mondes, tome xl. 24 THE POPULAR SCIENCE MONTHLY. Technical difficulties prevented the formal nomination of the com- mittee in that year ; and before the next meeting came round the Science Commission was in full work, and the ground was covered. Five years have passed; the commission has reported ; and the Brit- ish Association, if it deals at all Avith the problem that lies at the root of our scientific progress, will have to face the fact that only ten endowed schools in England give as much as four hours a week to the study of science ; in other words, that, in spite of ten years of talk, the eclat of a Royal Commission, a complete consensus of scientific authority, and the loud demands of less educated but not less keen- sighted public opinion, the organization and practical working of science in our higher schools has scarcely advanced a step since the Schools Inquiry Commission reported in 1868. Are the causes of this strange paralysis discoverable, and are they capable of present remedy ? We believe that they are notorious, and that it is in the power of the British Association at the present mo- ment to overrule them. It is therefore in the hope of rekindling a productive enthusiasm at a critical moment in the history of our science-teaching that we appeal with all the earnestness of which we are capable to the leaders of the great parliament, whose session will have opened before this goes to press. The first obstacle to be understood and reckoned with is the amazing confusion in the minds of unscientific leaders of opinion as to the very nature of education. An ex-Lord Chancellor gives away prizes to a school, declares in stately terms that Greek and Latin must always form the backbone of high intellectual training, and that the sciences can only be tolerated as a sort of ornament or capital to this great central vertebral column. On the following day an ex-Chancel- lor of the Exchequer gives away prizes at another school, assures the boys that modern scientific teaching is their being's end and aim, and envies them by comparison with himself, who at Winchester and Ox- ford basked only in the " clarum antiques lucis jubar." 1 In all such public utterances chaos reigns supreme. Men take side with one or other branch of mental discipline, unconscious of the Nemesis which waits on the divorce of literature from science, or of science from liter- ature, forgetful of the fundamental truths that all minds require gen- eral training up to a certain point, and that the period at which special education should supervene is the problem which awaits solution. The hostility of the clergy ranks high among the difficulties we have to recognize. To the gi-eat public schools this is matter of in- difference; but the vigorous head-master of a young and rising coun- ty school, who attempts, being himself a clergyman, to make real science compulsory in his school, is rattened by the vulgar heresy- hunters, who swarm in every diocese. The hint and shrug in society, the whisper at clerical conferences, the warning to parents attracted 1 The bright radiance of ancient light. PHYSICAL SCIENCE IN ENGLISH SCHOOLS. 25 by the school against " atheistic tendencies," keep down his numbers and wear out his energies, till his enterprise becomes a warning in- stead of an example to his admirers at other schools. In a neighbor- hood of rural squires and clergy, untempered by a large town's neigh- borhood, and unchecked by any man of education and intelligence holding sovereignty by virtue of superior rank and wealth, a school which treads doggedly in the ancient paths, and is flavored with gen- tle " High-Church tendencies," will certainly succeed even in second- rate hands, while a school which under superior chieftainship asserts the claims of science, and whose theology is therefore suspect, will as certainly long struggle for existence, if it does not finally succumb. The head-masters, with no inveterate intention, but by the force of circumstance's, are potent allies upon the side of nescience. Their position is peculiar. Enlightened, able, high-minded, and most labo- rious, to speak of them with disrespect would be to forfeit claim to a heai-insr. But of their whole number not more than two or three know anything at all of science ; they have gained honors and supremacy by proficiency in other subjects ; to teach well these sub- jects which they know, forms their happiness and satisfies their sense of duty ; and they feel natural dismay at the proposal to force upon them new and untried work which they have not knowledge to super- vise, and which must displace whole departments of classical study. Bifurcation they do not mind, for they hope that the dunces will be drafted into the modern school, and the clever boys retained upon the classical side ; but the momentous recommendation of the Royal Commission that six hours a week of science-teaching should be given to every boy in every school has taken away their breath ; it was only once alluded to at the last head-masters' meeting, and then with something between a protest and a sneer. They are too clear-sighted not to see that the demand for science-teaching is real, and too liberal not readily to accede to it, if some central authority, which they respect, at once puts pressure on them, and tenders such assistance and advice as they can trust. But, until these two things are done, they will pursue a policy of inaction. Nor is there any hope that this reluctance of head-masters will be stimulated by exuberant energy on the part of governing bodies. The instances in which these pet creations of the Endowed Schools Commission have appeared before the public hitherto make it evi- dent that absolute inactivity is the service they are best calculated to render to the cause of education ; but their probable devotion to science may be guessed from an incident reported in our columns some months ago, where a body of trustees, composed of country gentlemen of local mark, having to arrange a competitive examination under a scheme of the Charity Commission, adopted the machinery of the University Leaving Examination, but inserted a distinct pro- viso that no scientific subject recognized by the university regula- 26 THE POPULAR SCIENCE MONTHLY. tions should under any circumstances be taken up by the candidates, either as an alternative or a positive branch of work. Will the universities help or impede the spread of school science- teaching ? The universities adhere at present to their fatal principle that only one-sided knowledge shall find favor within their walls. A boy who knows nothing but classics, nothing but mathematics, noth- ing but science, may easily win a scholarship ; a boy who knows all three must seek distinction elsewhere ; and this rule shapes inevita- bly the teaching of the schools. The science scholarships at Oxford, of which we hear so much, fall mainly to three distinguished schools : two so large and wealthy that they can overpower most competitors by their expenditure on staff and apparatus ; the third planted in Oxford, with access to the university museum and laboratory, and with a pick of teachers'from the men of whom examiners are made ; and these schools insure success in science by abandoning other sub- jects almost or altogether in the case of the candidates they send up. No school which should carry out the recommendations of the com- missioners by giving six hours a week to science, and the rest of its time to literature and mathematics ; no school which should realize its function as bound to develop young minds by strengthening in fair proportion all their faculties of imagination, reason, memory, and observation — could offer boys for any sort of scholarship under the present university system with the faintest chance of success. What these institutions are powerless to avert or helpless to bring about is, we repeat, within the scope of the British Association to effect. All institutions, political or educational, will bow to a strong- ly-formed committee of scientific men, formally commissioned by the Association and speaking with authority, delegated as well as per- sonal, on scientific subjects. Let such a committee be revived as died on paper in 1871, including the acknowledged leaders of pure science, and weighted with the names of such educationalists as have shown themselves zealous for science-teaching. Let their functions be — 1. To communicate with the head-masters and governing bod- ies, calling attention to the recommendations of the Duke of Devon- shire's commission, asking how far and how soon each school is pre- pared to carry these out, and tendering advice, should it be desired, on any details as to selection and sequence of subjects, teachers, text- books, outlay. 2. Let them appeal to the universities, to which many of them belong, as to the bearing of science scholarships and fellow- ships upon school-teaching, and the extent to which such influence may be modified or ameliorated in that rearrangement of college funds which next session will probably be commenced. 3. Let them be instructed to watch the action of Government in any proposal made either in pursuance of Lord Salisbury's bill, or as giving effect to the Duke of Devonshire's commission, and let them be known to hold a brief for school science in reference to all such legislation. A NATURE OF THE INVERTEBRATE BRAIN. 27 single meeting of such a committee before the Association separates would settle a basis of action and compress itself into a working sub- committee. The time for papers and discussions is past ; they have done their work. What the schools and the head-masters want is authoritative guidance — the guidance not only of a blue-book, but of a living leadership, central, commanding, and accessible, to which they may look with confidence, and bow without loss of prestige. The precision of its dicta will clear up public confusion ; its abil- ity, conscientiousness, and pojmlarity, will overawe the clergy ; schools and universities will listen respectfully to suggestions echoed by their own best men ; and the three great departments of intellectual culture, equal in credit, appliances, and teaching power, will bring out all the faculties and elicit the special aptitudes of every English boy. " Hinc omne principium, hue refer exitum." l — Nature. -+*+- NATURE OF THE INVERTEBRATE BRAIN. By Professor H. CHAELTON BASTIAN. II. IT now remains for us to consider the disposition of the nervous sys- tem in some of the principal types of the sub-kingdom Mollusca. These are animals wholly different in kind from those we have just been considering, mostly aquatic, and all of them devoid of hollow, articulated, locomotor appendages. Their organs of vege- tative life attain a disproportionate development. On the other hand, what are termed the " organs of relation " present a wide range of variation, as may be imagined from the fact that while some of the simplest representatives of the Mollusca consist of mere motionless sacs or bags, containing organs of digestion, respiration, circulation, and generation, its more complex forms are active predatory creatures, endowed with remarkable and varied powers of locomotion, and with sense-organs as keen and as highly developed as those of insects. The lower type is represented by the motionless ascidian, and the higher by the active and highly-endowed cuttle-fish. Omitting any reference to the Polyzoa, we may turn our attention first of all to the Tunicata, of which the solitary ascidians may be taken as the type. They are marine animals, possessing no powers of locomotion, and having no head. The current of sea-water, serving for respiratory purposes, and, at the same time, containing food-parti- cles, enters a large branchial chamber, through an open, funnel-like projection of the investing tunic of the animal, the orifice of which 1 With this begin, to this refer the end. 28 THE POPULAR SCIENCE MONTHLY. is guarded by sensitive tentacula and a sphincter muscle. The mouth is situated at the bottom of this branchial sac, down the side of which minute particles of food are swept by ciliary action, so as to be brought within the simple commencement of the oesophagus. The effete sea-water passes through the walls of this branchial cavity into a general body-chamber, in which the viscera are contained. This cavity is bounded externally by a muscular expansion, lining the outer cellulose tunic. By the periodical contraction of this muscular sac, the water which enters it, together with food-residues and ova, is ex- pelled through another funnel-like opening, adjacent to and very similar to that by which it gains entry to the branchial chamber. Although these ascidians have a definite alimentary canal, a circu- latory system, and respiratory organs, together with a distinct genital apparatus, their life of relation with the external world is of the sim- plest description. They are stationary creatures, and have no pre- hensile organs, food being brought to the commencement of their alimentary canal by ciliary action. In correspondence with such a simple mode of life, we might ex- pect to find a very rudimentary nervous system, and this expectation is fully realized. The Tunicata possess a single small nervous gan- glion lying between the bases of the two funnels through which water is taken in and discharged. This ganglion receives branches from the tentacula guarding the orifice of the oral funnel, and possibly from the branchial chamber, while it gives off outgoing, filaments to the various parts of the muscular sac, and perhaps to the alimentary canal, and some of the other internal organs. In some of the solitary Tunicata a rudimentary visual function is presumed to exist. At all events, pigment-spots are situated on, or in very close relation with, the solitary ganglion. This single body seems to serve for the per- formance, in a rudimentary manner, of the various functions dis- charged by at least two pairs of ganglia in a large number of higher Mollusca, viz., those known as the cerebral and the parieto-splanchnic or branchial. The brachiopods are among the oldest and most wide-spread of the forms of life in the fossil state, and the geographical distribution of their living representatives at the present day is also very wide. Like the Tunicata, they are headless organisms, and lead a sedentary existence, attached either by a pedicle or by one division of their bivalve shells. The mouth is unprovided with any appendages for grasping food — nutritive particles being brought to it by means of ciliary currents. Numerous muscles exist which connect the valves of the shell to one another, and with the inclosed animal. And, though the visceral organization of the brachiopods is somewhat complex, no definite sense-organs have yet been detected in any of them. In the nervous system of these sedentary animals, there is, therefore, nothing answering to a brain as it is ordinarily constituted, NATURE OF THE INVERTEBRATE BRAIN. 29 though ganglia exist around the oesophagus which must receive afferent impressions of some kind, and from which branches proceed to the various muscles and viscera of the body. Such low sensory endowments as are presented by the Brachiopoda would be wholly incompatible with that degree of visceral complexity of organization which they possess, had it not been for the fact that they lead such a passive existence 'in respect to quest of food. They do not go in search of it at all — they remain securely anchored while food is brought to the entrance of their alimentary canal by means of cilia. • The absence of sense-organs and of a brain is, indeed, only compatible with a ^wtm-vegetative existence such as this. The lamellibranchs, or ordinary headless bivalve 3Iollusca, also include some representatives — such as the oyster and its allies — which lead a sedentary life after the fashion of the Mollusca already men- tioned. The valves of the shell in these lamellibranchs are lateral, instead of being dorsal and ventral as among the branchiopods. The shell is, however, closed by a single adductor muscle, and it is opened, when this relaxes, by means of an elastic hinge. The mouth of the oyster is surrounded by four labial appendages, whose functions are not very definitely known. It presents no other appendages of any kind in the neighborhood of the mouth, and, as in the two types of Mollusca already described, the food which it swal- lows is brought to the entrance of its oesophagus by means of ciliary currents. This well-known animal has a large and important nervous ganglion (Fig. 8, b) situated posteriorly, and close to the great ad ductor muscle. It gives off' branches to this muscle, to each half of the mantle, to the gills (c, c), and it sends forward two long parallel branches (d, c7), which serve to connect it with a much smaller an- terior ganglion (a, a) situated on each side of the mouth. These an- terior or labial ganglia are joined by a commissure arching over the mouth, and also by a more slender thread beneath the mouth, from which filaments (e) are given off to the stomach. These latter filaments may be considered to have a function similar to that of the stomato- gastric nerves in insects. The anterior ganglia receive nerves (/) from the labial processes, probably for the most part afferent in function. At all events, these processes have no distinct muscular structure. Other lamellibranchs possess a remarkable muscular appendage known as the foot, w*hich is in relation with an additional single or double nervous ganglion, and is used in various ways as an organ of locomotion. The animals possessing this organ are also provided with a second adductor muscle for closing their shells. Speaking of the various uses of the foot among bivalves, Prof. Owen says : " To some which rise to the surface of the water it acts, by its expansion, as a float ; to others it serves by its bent form as an instrument to drag them along the sands ; to a third family it is a burrowing organ ; to many it aids in the execution of short leaps." 3° THE POPULAR SCIENCE MONTHLY. These bivalves possessing a foot present three pairs of ganglia in- stead of two — the anterior or oval, the posterior or branchial, and the inferior or pedal. It occasionally happens, however, that the ganglia of the posterior or of the inferior pair become approximated or even fused into one. The fusion of the posterior pair takes place, as in the oyster, when the branchiae from which they receive nerves come close together posteriorly. On the other hand, in those mollusks in which the branchiae are farther apart, the two ganglia remain separate, and are connected only by a short commissure, as in the mussel (Fig. 9, b). Fig. 8. — Nervous System of the Oyster. Fig. 9. — Nervous System of the Common Mcssel. The separate existence or fusion of the inferior or pedal ganglia depends upon the size and shape of the foot. The nerves in relation with these ganglia are distributed almost wholly to this organ and its retractor muscles. Where the foot is broad the ganglia remain sep- arate, and are merely connected by a commissure. But where the foot is small and narrow, as in the mussel, the two ganglia become fused into one (Fig. §,p). Some of the special senses are unquestionably represented among these headless Mollusca, though the distribution of the different or- gans is very peculiar. Thus in Pecten, Pinna, Spondulus, the oyster, and many others, very distinct and often pedunculated ocelli are dis- tributed over both margins of the pallium or mantle. These vary in number from forty to two hundred or more, and are in connection with distinct branches of the circurupallial nerves. In the razor-fish, NATURE OF THE INVERTEBRATE BRAIN. 31 the cockle, Venus, and other bivalves possessing those prolongations of the mantle known as siphon-tubes, the eyes are situated either at the base or on the tips of the numerous small tentacles distributed around the orifices of these tubes, which in those of them living in the sand are often the only parts appearing above the surface. The margins of the mantle are also garnished by a number of short though apparently very sensitive tentacles, in which the creature's most spe- cialized sense of touch seems to reside. Some of these tactile append- ages, as well as some of the ocelli, send their nerves to the posterior or parieto-splanchnic ganglia, while those situated on the anterior borders of the mantle communicate with the anterior or oral ganglia. The latter ganglia also receive filaments from the so-called labial ap- pendages, whose function is uncertain, though it has been suggested that they may be organs of taste or smell. Lastly, in close relation with the pedal ganglia or ganglion, there are two minute saccules (Fig. 9, s), to which an auditory function is usually ascribed. Thus we find among these headless mollusks a distribution of spe- cially impressible parts or sensory organs, such as cannot be paralleled among any other animals. The sense of touch and the sense of sight seem to be more especially in relation with the great posterior gan- glia. These sensory functions are, however, to a minor extent shared by the oral ganglia, which are also in relation with parts that may possibly be organs of taste or smell. On the other hand, auditory impressions are invariably brought into relation with the inferior or pedal ganglia. In these headless mollusks, therefore, the functions pertaining to the brain in other animals are distributed in a very re- markable manner, and the anterior ganglia cannot in them be proper- ly regarded as representing such an organ. The viscera in these lamellibranchs are also in relation with the three pairs of ganglia, and not exclusively with any one of them. Filaments to the intestinal canal and the liver are usually given off from the commissures between the anterior and the posterior ganglia; the genital organs are in connection with filaments coming from the commissures between the anterior and the inferior or pedal ganglia ; while the branchiae are in relation with the ganglia at the posterior part of the body.1 There is another interesting class of mollusks — the Pteropoda — which,, in respect of powers of locomotion and the possession of a dis- tinct head, may, if for no other reasons, be said to lead us on from the comparatively sluggish bivalve Mullmca to the gasteropods and the cephalopods, all of which are distinguished by definite and wide- reaching powers of locomotion, and by the possession of a distinct head carrying sense-organs, and a more or less developed brain. 1 In speaking of the nervous system of the lamellibranchs, I have not alluded to cer- tain small accessory ganglia which exist in some of them in relation with peculiar special- ly developed contractile structures. 3z THE POPULAR SCIENCE MONTHLY. The possession, by many members of this class, of two fin-like mus- cular expansions attached to the side of the head induced Cuvier to give them the name Pteropoda. Prof. Owen says : "All the species of Pteropoda are of small size ; they float in the open sea, often at great distances from any shore, and serve, with the Accdep/ice, to people the remote tracts of the ocean. In the latitudes suitable to their well-being, the little Pteropoda swarm in incredible numbers, so as to discolor the surface of the sea for leagues ; and the Clio and the Limacina constitute, in the northern seas, the principal article of food of the great whales." Some of the least highly-organized members of this class, such as the Hyalaceos, are provided with a bivalve shell, and cannot be said to possess a head. They have a simple commencement of the ali- mentary canal at the anterior extremity of the body ; but since this anterior extremity has no tactile appendages and no eyes, and inas- much as it also contains no cerebral ganglia, it can have no claim to be considered as a head. Their chief nervous centre consists of a flat, somewhat quadrate, sub-oesophageal ganglion, to the anterior angles of which is attached a nervous commissure which extends upward so as to encircle the gullet, though there are no ganglia either on or at the sides of this tube in the usual situation occupied by cere- bral ganglia. In other pteropods devoid of a shell, we meet with a higher organ- ization. Thus in Clio there is a distinct head bearing sensory ap- pendages in the form of two tentacula and two eyes, and containing in its interior a brain. This brain is represented by two connected super-oesophageal ganglia, which are in relation, by means of nerves, with the cephalic sensory organs, and in connection with the sub-ceso- phageal commissure are the two pedal and two branchial ganglia. The two pairs of ganglia exist separately in Clio and its allies, though they are combined into one quadrate mass in Hyalea. In this latter there are two acoustic vesicles in contact with the anterior part of the great ganglion, while in Clio similar vesicles are in connection with the anterior pair of sub-cesophageal ganglia — that is, with the pair which corresponds with the pedal ganglia of the common bivalve mollusks. Gasteropods constitute a class of organisms which, in point of numbers, can only be compared with the still more numerously repre- sented class Of insects. Their name is derived from the fact that these animals crawl by means of a large muscular expansion stretched out beneath the viscera. The locomotion of the members of this class may be said to be, in the main, dependent upon their own individual efforts, so that, in this respect, they differ widely from the pteropods, whose locomotions are brought about by wiuds driving them along the surface of the water on which they float. Some gasteropods are terrestrial, air-breathing animals, though by NATURE OF THE INVERTEBRATE BRAIN. 33 far the greater number are aquatic, and breathe by means of gills. But being all of them, as Prof. Owen says, "endowed with power to attain, subdue, and devour organic matter, dead and living," we find their nervous system not only better developed, more complex and concentrated, but also in relation with more highly-evolved organs of special sense and exploration. It offers considerable variations in its general arrangement, especially as regards relative positions of gan- glia, though these modifications are, to a great extent, referable to differences in the outward configuration of the body. Some of the differences in external form which are to be met with among gasteropods are well illustrated by the limpet or the chiton, as compared with the snail. Here differences in form coexist with dif- ferences in habit, so that we almost necessarily meet with notable variations in the disposition of the principal parts of the nervous system. In the limpet we find that the two small cerebral ganglia (Fig. 10, a) are widely separated from one another, and lie at the side of the cesophagus. Each receives a rather large nerve from one of the ten- tacles, and a smaller optic nerve. A commissure connects these cere- bral ganglia above the cesophagus with one another, while each of them is also in relation by means of two descending commissures with a series of four connected ganglia forming a transversely-ar- ranged row beneath the cesophagus. Of these the two median gan- glia (JB) correspond with the pedal, while the two external ( C) cor- respond with the branchial ganglia, though they are here separated from one another by an immensely wide interval. Fig. 10.— Nervous System of the Common Limpet. Fig. 11. — Nervous System of Chiton marmoralus. However small and undeveloped t-he duplex brain of the limpet may be, this organ exists in an even more rudimentary state in some other gasteropods. Thus, in the chiton, which is a close ally of the limpet, and about the most simply organized of all the gasteropods, there are neither tentacles nor eyes, and, as a consequence of this, VOL. X. — 3 34 THE POPULAR SCIENCE MONTHLY. there are (Fig. 11) no supra-oesophageal ganglia. There is nothing, in fact, to which the term brain can be appropriately applied. But, if we turn now to the much more active snail, we find the ner- vous system existing in a more developed and concentrated form. There is (Fig. 12, I) a large ganglionic mass situated over the oesopha- gus, each half of which receives a considerable bundle of nerve-fibres (f) from the eye (b) of the smaller side, which is situated at the tip of the larger tentacle. It also receives another bundle of nerves (k) Fia 12.— Head and Nekvous System of the Common Snail. from the small tentacle on each side, which has in all probability a tactile function. The auditory vesicles are here in a new position. They are in immediate relation with the posterior aspect of these ganglia constituting the brain, though in other gasteropods »they are, as in bivalve Mollusca, found to be connected with the pedal ganglia. That gasteropods are endowed with a rudimentary sense of smell is now generally admitted by naturalists, though hitherto they have been unable to locate this endowment in any particular organ or sur- face-region. The brain of the snail is connected, by means of a triple cord or commissure on each side of the cesophagus, with a still longer double ganglionic mass (m). This latter body, situated beneath the oesopha- gus, represents the pair of pedal and the pair of branchial ganglia of the bivalve Mollusca. Here nerves are received from the integument and given off to the muscles of the foot, while they are also received and given off from the respiratory and other organs. In the nautilus and some other representatives of the next class, Cephalopoda, the nervous system attains a development only slightly in advance of that met with among the highest gasteropods, though in the active and predaceous cuttle-fish, and in its near ally, the octo- pus, we find the nervous system presenting the highest development to be met with araons; the sub-kino-dom Mollusca. NATURE OF THE INVERTEBRATE BRAIN. 35 One of the most striking characteristics of the principal nerve- centres of the cuttle-fish is the fact of the existence of a very large optic ganglion (Fig. 13, 2), in "connection with a well-developed eye, on each side. Each optic lobe, according to Lockhart Clarke, is " as large as the rest of the cephalic ganglia on both sides taken together." From each of these lobes an optic peduncle passes inward to join a supra-oesophageal ganglionic mass, which bears on its surface a large bilobed ganglion (1), thought by Clarke to be homologous with the cerebral lobes of fishes. It is connected, by means of two short cords, Fig. 13.— Nervous Ststem of the Common Cuttle-fish (Sepia officinalis). with a much smaller bilobed ganglion, known as the pharyngeal (7). This double ganglion receives nerves from what are presumed to be the organs of taste and smell, and gives off nerves to the tongue and powerful parrot-like jaws with which the creature is provided. The supra-oesophageal mass is connected, by cords at the sides of the oesophagus, with a very large ganglion lying beneath it (4), which is partially divided into an anterior and a posterior division. The anterior division is in relation, by means of large nerves (6), with the feet and tentacles. A commissure also unites it with the pharyngeal ganglion, so that the tentacles and arms are thus able to be brought 3 6 THE POPULAR SCIENCE MONTHLY. into correlated action witb. the jaws. The posterior portion of the sub-oesophageal mass receives nerves from, and also gives off nerves (14) to, the branchiae and other viscera, as well as to the mantle (13, 13). The auditory organs and their nerves are also connected with this branchial and pallial ganglion. These organs are lodged in the sub- stance of the cartilaginous framework investing the nerve-ganglia — a structure which seems to answer to a rudimentary skull. The roots of the auditory nerves are probably principally in relation with the pallial portion of the branchio-pallial ganglion. The locomotions of these creatures are largely brought about by contractions of the pal- lial chamber, though these contractions of the mantle are also subser- vient to the respiratory function. The share which the branchio-pallial ganglia take in bringing about and regulating the movements of the cuttle-fish would seem to explain the connection of the auditory nerves with them rather than with the homologues of the pedal ganglion, with which the auditory saccules are in relation in most other mollusks. But, whatever may be the precise explanation of the different connections of the auditory nerves in the cuttle-fish tribe, the fact remains that their connections are still away from the brain proper. They are, as in most other Mollusca and in those insects in which auditory organs are known to occur, in intimate relation with one of the principal motor centres. This survey of some of the principal forms of the invertebrate brain, brief though it has been, should have sufficed to call attention to the following important facts and inferences : 1. That sedentary animals, though they may possess a nervous system, are often headless, and then have nothing answering to a brain. 2. That where a brain does exist, it is invariably a double organ. Its two halves may be widely separated from one another, though at other times they are fused into a single mass. 3. That the component or elementary parts of the brain in these lower animals are ganglia in connection with some of those special impressible parts or sense-organs, by means of which the animal is brought into harmony with its environment or medium. 4. That the sensory ganglia, which as an aggregate constitute the brain of invertebrate animals, are connected with one another both on the same and on opposite sides of the body, either by continuous growth or by means of commissures. 5. The size of the brain as a whole, or of its several parts, is strictly regulated by the development of the animal's special sense- organs. This is so, because, the more these impressible surfaces be- come elaborated and attuned to help in discriminating between nu- merous different external impressions, the larger are the ganglionic masses with which their nerves ai*e in relation. 6. Of the several sense-organs and sensory ganglia whose activity NATURE OF THE INVERTEBRATE BRAIN. 37 lies at the root of the intellectual and instinctive life (such as it is) of invertebrate animals, some are much more important than others. Two are notable for their greater proportional development, viz., tactile organs and visual organs. The former are soon outstripped in importance by the latter. The visual sense, indeed, and its related nerve-ganglia, attain an altogether exceptional development in the higher insects and mollusks. V. The sense of taste and that of smell are developed to a much lower extent. It is even difficult to point to distinct organs or im- pressible surfaces as certainly devoted to the reception of impressions of this kind. 8. The sense of hearing is also developed to a very slight extent. No distinct sense-organ of this kind has been discovered, except in a few insects and in members of the sub-kingdom Mollusca It is, however, of no small interest to find that, where these organs do exist, the nerves issuing from them are not in direct relation with the brain, but are immediately connected with one of the principal locomotor nerve-centres of the body. 9. The associated ganglia representing the single or double brain are, in animals possessing a head, the centres in which all impressions from sense-organs, save those last mentioned (the auditory), are re- flected on to appropriate groups of muscles. This "reflection occurs either at once or after the stimulus has passed through other ganglia, whence it is passed along nerves to those groups or combinations of muscles whose simultaneous or successive contractions give rise to the organism's reply to such impressions. It may be easily under- stood, therefore, that in all such animals perfection of sense-organs, size of brain, and power of executing varied muscular movements, are intimately related to one another. 10. But a fairly parallel correlation also becomes established be- tween these various developments and that of the internal organs. An increasing visceral complexity is gradually attained. Such in- creased visceral complexity carries with it the necessity for a further development of nervous communications. The several internal or- gans have to be brought into more perfect relation with the sensori- motor nervous system, and also with one another, for all joint actions in which two or more of them may be concerned. 11. In invertebrate animals the visceral system of nerves has, when compared with the rest of the nervous system, a greater pro- portional development than among vertebrate animals. Its importance among the Invertebrata is not dwarfed by the enormous development of the brain and spinal cord, which gradually declares itself among the Vertebrata. 12. Impressions emanating from the viscera and stimulating the organism to movements of various kinds, whether in pursuit of food or of a mate, would, therefore, have a proportionally greater impor- 3 8 THE POPULAR SCIENCE MONTHLY. tance as constituting part of the ordinary mental life of invertebrate animals. Movements thus initiated will be found to afford a basis for the development of many so-called instinctive acts. -♦-*♦- PKENATAL AND INFANTILE CULTURE.1 By Dr. E. SEGUIN. TO educate children for themselves is rare in Europe, and is considered rather quixotic. The youth of the people are mer- chantable commodities, soon to be credited to the party which puts its stamp upon them. Therefore, when they are worth having, they are picked up as eagerly as nuggets. Priests pretend to teach them to think, yet care only to impose upon them a belief which implies obedience to their craft ; Kaisers claim their direction, not to elevate them, but to put them among their droves of subjects ; bourgeois and manufacturers give them a minimum of instruction, just sufficient to insure their working dependence, and to qualify their own sons to be fed at the public expense ; while the working-men themselves — de- moralized by such examples — put their apprentices at menial employ- ment, and cheat them out of their rightful technical training. From this standpoint we consider European children as in four groups : those who receive no education ; those who do not receive the education they need ; those who receive an education which dis- qualifies them for work ; and those whose education prepares them for work. From another point of view we saw that the European children enter the school younger, are trained longer, and are ad- vanced further, than the Americans. As a consequence of this last contrast, we shall have less to say about the primary and' grammar schools, and more about the infantile and the professional. We will leave the other consequences to issue naturally from observation. 1. The Cradle. — At the Vienna Exposition there was a pavilion de V enfant (infant pavilion), a room replete with the necessaries of the nursery — and also with its superfluities — intended altogether to represent the unbounded wishes of a mother for her baby's comfort and happiness. This palace of luxurious nursing ought, in the esti- mation of the writer, to have been accompanied by a little manual of what is necessary to protect and to prepare life before nativity, in re- lation to what may be called foetal education. During this first period the feelings come mainly through reflex impressions from the mother, a process which not only lays the foun- dation of health and vitality, but which forms the deeper strata of 1 Extracts from the Report of the Commissioners of the United States to the Inters national Exhibition held at Vienna, 1S73. PRENATAL AND INFANTILE CULTURE. 39 the moral dispositions and of the so-called innate ideas. The man- agers of the world " from behind the screens " know this, for it is at this time that they impose on plebeian women pilgrimages and ec- static "nove7ias"1 and keep those of a higher class under more strin- gent impressions. Here, in Vienna, for instance, from the time of the Emperor Charles V. till quite recently, when an heir to the throne was expected, the empress was given in charge of a special director, who would regulate all her actions and surroundings, in view of com- mencing the course of submissive education of the contingent mon- arch, as early as the first evolution from the yolk-substance of the human egg during embryogenesis. Similar influence is now claimed for an object diametrically opposed to the degeneracy thus arrived at in the house of Hapsburg. It can be attained by counsels printed either in book-form or on scrolls, as are the sentences of the Koran. But, whatever may be the form given to this magna charta of the rights of the unborn, let it be found precisely where these rights ought to be kept most sacred, in the nursery ; where their enforce- ment would protect the mother and elevate her function, at the same time that it would insure her fruit against the decay resulting from wrong prenatal impressions. We know that a cold contact with the mother makes the foetus fly to the antipode of its narrow berth ; that a rude shcck may destroy it, or originate life-long infirmities ; that, the emotion of fear in the mother is terror or fits within ; that harsh words vibrate as sensibly in the liquor of the amnion as in the fluid of the labyrinth of the ear. For instance, when a mother has lulled her home-sorrows with strains of soothing music, her child, too often an idiot, shows wonderful mu- sical proclivities amid the wreck of all the other faculties of his mind. For thirty-five years the writer has furnished his share of the facts, which abound in modern books on physiology, in support of this doctrine. It is useless to give here the illustrations detailed in the report ; but experienced physicians will testify that, when their hands receive a new-comer, they plainly read upon his features the dominant feel- ings and emotions of its mother during that intra-uterine education whose imprints trace the channel of future sympathies and abilities. Therefore, if it is noble work to educate or to cure the insane, the idiot, the hemiplegic, the epileptic, and the choreic, how much higher is the work of preventing these degeneracies in the incipient being, by averting those commotions which storm him in the holy region in- tended for a terrestrial paradise during the period of evolution ! To teach him reverence toward the bearer of his race, to instruct her in the sacredness of bland and serene feelings during the Godlike cre- ative process, is educating two generations at once — this is the high- est education of the nursery. 1 A nine days' season of prayer. 4o THE POPULAR SCIENCE MONTHLY. From this, the true cradle of mankind, let us look at that made for the baby. There was no end of cradles in the pavilion de Pen/ant ; and we may find more philosophy in them than the upholsterer in- tended to put there. Therein the infant will at first but continue his ovum-life; and for this the cradle must be fitted. Let us see. The head is bent, the extremities are drawn up, and the body shaped like a crescent. This attitude gives to the muscles the greatest relaxation, and to the cartilages, which cap the bones, the position most favor- able to nutrition and growth. Generally, the baby rests on the right side, to free the heart from pressure and to facilitate its movements. In this mode of reclining, the left hemi-cerebrum will contain more blood than the right, which is compressed by the pillow. Attitudes concordant with the sleepy habits of the first months, and the activity of the mind during this long sleepiness, indicate the future prepon- derance of the mental operations of the left over the right side of the brain, the approaching superior nutrition and dexterity of the right over the left hand, and later even the causation of paralysis on the left. For the present, and for some time yet, the infant will live mainly in his sleep ; during which, more than when awake, he will be seen angry, smiling, or thinking, even in his well-defined dreams. How important it is, then, that the cradle be formed in accord- ance with these natural indications ! A transitory abode between the basin and the bed, it should be a warm, soft, yet supporting recipient, ampler than the former, better defined in its shape than the latter, with curves less short than circles, and more varied than ovals. An egg, vertically split, would make two such cradles, or nests, suited either for child or bird. But as soon as the nursling awakes to the world, and wants to be introduced to everything, his couch must be enlarged and enlivened, and must look more and more like a school and play-room. Other- wise it becomes a prison, whence, Tantalus-like, baby looks at his surroundings. Here is his first lesson of practical sociability. To see, and not be able to reach, to perceive images, with no possibility of seizing the objects, renders him impatient, fretful, or unconcerned, and opens an era of exaction upon others, or of diffidence of himself, or of indiffex-ence for any attainment, which unavoidably ends in im- morality or incapacity, or in both. Viewed from this standpoint, these cradles, so varied, so elegant, so easy to keep clean, and to carry from the light of the window by day to the recess of the alcove at night — the best being of French and Austrian manufacture — are yet very imperfect in their bearing on education. Let us mark some of their shortcomings. Little ones have an instinctive horror of isolation. Whoever studies them knows that when they awake they look not, at first, with staring eyes, but with searching hands ; they seek not for sights, but for contacts. This love of contact, whence results the primary PRENATAL AND INFANTILE CULTURE. 4i education of the most general sense, the touch, is ill-satisfied with the uniformity of the materials at hand, as exemplified at Vienna or Paris. (In November last I saw a similar exhibition, a pavilion de V enfant, in the Champs Elysees, but it was no improvement on that of the Prater.) In this respect, the child of poor people fares better, having the opportunity of amusing himself for hours in experiencing the rude or soft, warm or cold, contacts of his miscellaneous surroundings ; whereas the hand of the offspring of the rich finds all around the sameness of smooth tissues, which awake in his mind no curiosity ; he calls for some one to amuse him, gets first angry, then indifferent, and does not improve the main and surest sense of knowledge, his touch. But soon other senses are awakened : audition — of which here- after— and vision, for the enjoyment of which the cradle becomes a kind of theatre. For a mother must be very destitute or despondent who does not try to enliven it with some bright things laid on, or flapping above. One may benevolently smile at the extravagancies of colors and patterns intended to express this feeling, but these ex- aggerations must also give a serious warning. Physiologically viewed, this is a grave matter. The form of the cradle demands fitness ; its ornamentation requires a more extended knowledge. When planning it, a mother must remember that the fixity of the eye upon any object — particularly upon a bright one, and more so if that object is situated upward and sideways from the ordinary range of vision — and through the eye the fixedness of the mind while the body is in a state of repose, constitute a concurrence of conditions eminently favorable to the production of hypnotism, and its terrible sequels, strabismus and convulsions. Hypnotism, which, when unsuspected, is not controlled, is often mistaken for tranquil happiness or natural sleep. Psychologically viewed, the decoration of the cradle is of equal moment. To surround an infant with highly wrought or colored figures, often grotesque, or at least untrue to Nature, may, by day, attract more attention than his faculties of perception can safely bestow ; hence fatigue of the brain, or worse ; but it will, by night, evoke other than the perceptive and rational powers, for, when the lights and shadows of dusk alter all the forms and deepen every color, the faculty of imprinting images being led astray, it photo- graphs distorted imprints from confused, often moving, sometimes rustling, ornaments. In this way the mind is made the subject of hal- lucinations, which it accepts as objective, without inquiring into their causes, till it comes to the fatal credo quia absurdum (I believe, be- cause it is absurd). The seeds of most of the insanities are sown at or before this time. These were the first impressions that forced themselves upon my 42 THE POPULAR SCIENCE MONTHLY. mind in the pavilion de P enfant. Here is, in a few words, a resume of them : Paucity of the material upon which the inexperienced yet inquisitive baby can exercise, with interest and profit, his sense of touch ; profusion, bad taste, and dangerous disposition of the objects which speak to the eye, if not always with the intention, at least with the almost uniform result, of giving wrong or dangerous im- pressions. Attention was next called to what had been done, and to what had been left undone, for the cultivation or the satisfaction of the other senses of the infant. But here it was soon perceived that our inqui- ries went beyond the sphere of what was exhibited. There were plenty of Farina's and Rimmel's volatilities, some of Alexander's, De- bain's, or Smith's sighing accordeons, but no means of cheering and educating the nascent yet already inquisitive senses. Further exami- nation showed that the perfumes were there as an attenuation and the music as a distraction, and both intended for other senses than the infant's. From these and other omissions it was concluded that nursery arrangements are as yet intended rather for the mother's and nurse's comfort than for the baby's improvement. 2.' The Ckeche. — This pavilion de V enfant ought to contain at least ene model crhche. Creche is the French name of the public nursery where working- women leave their little ones in the morning, and whence they bring them home at night. The cr&che I Horrid necessity ! Beginning of the communistic inclined plane upon which those who pay and do not receive rents slide with a fearful rapidity; yet a kind institution for those already fallen into the gulf. Since, therefore, creches must be, their latest improvements should have been represented at the Vienna Exhibition next to the appliances of the most luxurious nursing. There could have been tested the action of colors, of light, and its various attributes, on the organ of vision ; the influence of varied sounds, of harmonies and melodies on the virgin audi- tion, the mind, and the sympathetic centres ; the power of primary perceptions to awaken first ideas, to impel to determinations of the will, and to raise or calm the various passions ; the effects of diet upon those passions ; the effect of modification of food and digestion ; the influence of rest and sleep on the body's temperature, on the pulse and respiration ; the influence of the artificial, the moist, or the dry heat of the nursery on the too precocious development of the nervous centres, and, subsequently, on the prevalence of chronic or acute meningitis, diphtheria, and croup ; besides many other problems whose solution depends on the early study of phenomena wdiich can be found in the creche as surely as the flower in the bud. There, bet- ter than anywhere else, they may be studied with profit to all parties. Let us bear in mind that the rich man can never flatter himself that he does a gratuitous charity, since from its poor recipient comes many PROFESSOR HUXLEY'S LECTURES. 43 times its worth in useful experience, directly benefiting the would-be benefactor. We do not overlook the fact that many mothers, particu- larly among those both educated and fruitful, pay the closest atten- tion to these questions, and become expert therein, but, as they lack the means of record and transmission of their observations, their ex- perience dies, so to speak, with each generation. Hence the nursing of babies continues to be a work of devotion, but does not become the coordinated and progressive art it ought to be in well-organized creches open to criticism in public exhibitions. Thus in Vienna, at least, an opportunity was lost. -*♦*- PKOFESSOK HUXLEY'S LECTURES.1 THE THREE HYPOTHESES OF THE HISTORY OF NATURE. WE live and form part of a system of things of immense diver- sity and perplexity, which we call Nature, and it is a matter of the deepest interest to all of us that we should form just concep- tions of the constitution of that system and of its past history. With relation to this universe, man is, in extent, little nipre than a mathe- matical point ; in duration, but a fleeting shadow. He is a reed shaken in the winds of force ; but, as Pascal long ago remarked, although a reed, he is a thinking reed, and, in virtue of that wonderful capacity of thought, he has a power of framing to himself a symbolic concep- tion of the universe, which, although doubtless highly imperfect, and although wholly inadequate as a picture of that great whole, is yet sufficient to serve him as a guide-book in his practical affairs. It has taken Ions; asres of accumulated and often fruitless labor to enable man to look steadily at the. shifting scenes, phantasmagoria of Na- ture, to notice what is fixed among her fluctuations, and what is regu- lar among her apparent irregularities ; and it is only comparatively lately, within the last few centuries, that there has emerged the con- ception of a pervading order and a definite course of things, which we term the course of Nature. But out of this contemplation of Nature, and out of man's thought concerning her, there has in these later times arisen that conception of the constancy of Nature to which I have referred, and which at length has become the guiding conception of modern thought. It has ceased to be almost conceivable to any person who is familiar with 1 The first of three lectures on " The Direct Evidence of Evolution," delivered at Chickering Hall, New York, September 18th. From the report of the New York Tribune, carefully revised by Prof. Huxley. 44 THE POPULAR SCIENCE MONTHLY. the facts upon which that conception is based, that chance should have any place in the universe, or tbat events should follow anything but the natural order of cause and effect. We have come to look upon the present as the child of the past and as the parent of the future; and, as we have excluded chance from any share or part in the order of things, so in the present order of Nature men have come to neglect, even as a possibility, the notion of any interference with that order. And, whatever may be men's speculative notions upon those points, it is quite certain that every intelligent person guides his life and risks his fortune upon the belief that the order of Nature is constant, and the relation of cause to effect unchanged. In fact, there is no belief which we entertain which has so com- plete a logical basis as that to which I have just referred. It tacitly underlies every process of reasoning ; it is the foundation of every act of the will. It is based upon the broadest induction, and it is verified by the most constant, regular, and universal of deductive processes. But we must recollect that any human belief, however broad its basis, however defensible it may seem, is, after all, only a probable belief, and tbat our broadest generalizations are simply statements of the highest degrees of probability. Though we are quite clear about the constancy of Nature at the present time, and in the present order of things, it by no means follows necessarily that we are justified in expanding this generalization into the past, and in denying absolutely tbat there may have been a time when Nature did not follow a fixed order, when the relations of cause and effect were not definite, and when external agencies did not intervene in the gen- eral course of Nature. Cautious men will admit that such a change in the order of Nature may have been possible, just as a very candid thinker may admit that there may be a world in which two and two do not make four, and in which two straight lines do not inclose a space. In fact, this question with which I have to deal in the three lectures I shall have the honor of delivering before you, this question as to the past order of Nature, is essentially an historical question, and it is one that must be dealt with in the same way as any other histori- cal problem. I will, if you please, in the first place, state to you what are the views which have been entertained respecting the order of Nature in the past, and then I will consider what evidence is in our possession bearing upon these views, and by what light of criticism that evidence is to be interpreted. So far as I know, there are only three hypotheses which ever have been entertained, or which well can be entertained, respecting the past history of Nature. Upon the first of these the assumption is, that the order of Nature which now obtains has always obtained ; in other words, that the present course of Nature, the present order of things, has existed from all eternity. The second hypothesis is, that the present state of PROFESSOR HUXLEY'S LECTURES. 45 things has had only a limited duration, and that at some period in the past the state of things which we now know (substantially, though not, of course, in all its details) arose and came into existence without any precedent condition from which it could have naturally proceeded. The third hypothesis also assumes that the present order of Nature has had but a limited duration, but it supposes that the present order of things proceeded by a natural process from an antecedent order, and that from another antecedent order, and so on ; and on this hy- pothesis the attempt to fix any limit at which we could assign the commencement of this series of changes is given up. I am very anxious that you shall realize what these three hypotheses actually mean ; that is to say; what they involve, if you can imagine a spec- tator to have been present during the period to which they refer. On the first hypothesis, however far back in time you place that specta- tor, he would have seen a world essentially, though not perhaps in all its details, similar to that which now exists. The animals which ex- isted would be the ancestors of those which now exist, and like them ; the plants in like manner would be such as we have now; and the supposition is that, at however distant a period of time you place your observer, he would still find mountains, plains, and waters, with ani- mal and vegetable products flourishing upon them and sporting in them just as he finds now. That view has been held. It was a favor- ite fancy of antiquity, and has survived toward the present day. It is worthy of remark that it is an hypothesis which is not inconsistent with what geologists are familiar with as the doctrine of Uniformita- rianism. That doctrine was held by Hutton, and in his earlier days by Lyell. For Hutton was struck with the demonstration of astrono- mers that the perturbations of the planetary bodies, however great they may be, yet sooner or later right themselves, and that the solar system contained within itself a self-adjusting power by which these aberrations were all brought back to an equilibrium. Hutton imagined that something of the same kind may go on in the earth, although no one recognized more clearly than he the fact that the dry land is being constantly washed down by rain and rivers and deposited in the sea, and that thus in a certain length of time, greater or shorter, the inequalities of the earth's surface must be leveled, and its high lands brought down to the sea. Then, taking into account the internal forces of Nature, by which upheavals of the sea- bottom give rise to new land, he thought that these operations might naturally compensate each other, and thus, for any assignable time, the general features of the earth might remain what they are. And, inasmuch as there need be no limit under these circumstances to the propagation of animals and plants, it is clear that the logical develop- ment of this idea might lead to the conception of the eternity of the world. Not that I mean to say that either Hutton or Lyell held this conception — assuredly not ; they would have been the first to repu- 46 THE POPULAR SCIENCE MONTHLY. diate it. But, by the arguments they used, it might have been pos- sible to justify this hypothesis. The second hypothesis is that to which I have referred as the hypothesis which supposes that the present order of things had at some no very remote time a sudden origin, so that the world, such as it now is, arose. That is the doctrine which you will find stated most fully and clearly in the immortal poem of John Milton, the English " Divina Commedia," "Paradise Lost." I believe it is largely to the influence of that remarkable work, combined with daily teachings to which we have all listened in our childhood, that this hypothesis owes its general wide diffusion as one of the current beliefs of English- speaking people. If you turn to the seventh book of " Paradise Lost " you will find there stated the hypothesis to which I refer, which is briefly this: That this visible universe of ours made its appearance at no great distance of time from the present day, and that the parts of which it is composed made their appearance in a certain definite order in the space of six natural days, in such a manner that in the first of these days light appeared; in the second, the firmament or sky separated the water above from the water beneath it ; on the third day the waters drew away from the dry land, and upon it a varied vegetable life similar to that which now exists made its ap- pearance ; that the fourth day was devoted to the apparition of the sun, the stars, the moon, and the planets ; that on the fifth day aquatic animals originated within the waters; that on the sixth day the earth gave rise to our four-footed terrestrial creatures, and to all varieties of terrestrial animals except birds, which had appeared on the pre- ceding day ; and, finally, that man appeared npon the earth, and the work of fashioning the universe was finished. John Milton, as I have said, leaves no doubt whatever as to how, in his judgment, these marvelous processes occurred. I doubt not that his immortal poem is familiar to all of you, but I should like to recall one passage to your minds, in order that I may be justified in what I have said re- garding the perfectly concrete, definite conception which Milton had respecting the origin of the animal world. He says : " The sixth, and of creation last, arose With evening harps and matin ; when God said, ' Let the earth bring forth soul living in her kind, Cattle and creeping things, and beast of the earth, Each in their kind.' The earth obeyed, and straight Opening her fertile womb, teemed at a birth Innumerous living creatures, perfect forms, Limbed and full-grown ; out of the ground uprose, As from his lair, the wild beast, where he wons In forest wild, in thicket, brake, or den ; Among the trees in pairs they rose, and walked ; The cattle in the fields and meadows green ; Those rare and solitary, these in flocks PROFESSOR HUXLEY'S LECTURES. 47 Pasturing at once, and in broad hoards upsprung. The grassy clods now calved ; now half appears The tawny lion, pawing to get free His hinder parts, -then springs, as broke from bonds, And rampant shakes his brinded mane ; the ounce, The libbard, and the tiger, as the mole Kising, the crumbled earth above them threw In hillocks ; the swift stag from underground Bore up his branching head; scarce from his mould Behemoth, biggest born of earth, upheaved His vastness ; fleeced the flocks and bleating rose As plants ; ambiguous between sea and land, The river-horse and scaly crocodile. At once came forth whatever creeps the ground, Insect or worm." There is no doubt as to the meaning of this statement, or as to what a man of Milton's genius expected would have been actually visible to one who could witness the process of the origination of liv- ino; things. The third hypothesis, or the hypothesis of evolution, supposes that at any given period in the past we should meet with a state of things more or less similar to the present, but less similar in proportion as we go back in time ; that the physical form of the earth could be traced back in this way to a condition in which its parts were sepa- rated, as little more than a nebulous cloud making part of a whole in which wTe should find the sun and the other planetary bodies also re- solved ; and that, if we traced back the animal world and the vege- table woi'ld, we should find preceding what now exist animals and plants not identical with them, but like them, only increasing their differences as Ave go back in time, and at the same time becoming simpler and simpler, until finally we should arrive at that gelatinous mass which, so far as our present knowledge goes, is the common foundation of all life. The hypothesis of evolution supposes that in all this vast progres- sion there would be no breach of continuity, no point at which we could say " This is a natural process," and " This is not a natural pro- cess," but that the whole might be compared to that wonderful series of changes which may be seen going on every day under our eyes, in virtue of which there arises, out of that semifluid, homogeneous sub- stance which we call an egg, the complicated organization of one of the higher animals. That, in a few words, is what is meant by the hypothesis of evolution. I have already suggested that in dealing with these three hypoth- eses, in endeavoring to form a judgment as to which of them is the more worthy of belief; or whether none is worthy of belief — and our condition of mind should be that suspension of judgment which is so difficult to all but trained minds — we should be indifferent to all a 48 THE POPULAR SCIENCE MONTHLY. priori considerations. The question is a question of fact, historical fact. The universe has come into existence somehow or other, and the question is, whether it came into existence in one fashion, or whether it came into existence in another ; and, as an essential preliminary to our further discussion, permit me to say two or three words as to the nature of historical evidence, and the kinds of historical evidence. The evidence as to the occurrence of any fact in past time is of one or two kinds, which, for convenience' sake, I will speak of on the one hand as testimonial evidence, and on the other as circumstantial evidence. By testimonial evidence I mean human testimony ; and by circum- stantial evidence I mean evidence which is not human testimony. Let ine illustrate by a familiar figure what I mean by these two kinds of evidence, and what is to be said respecting their value: Suppose that a man tells you that he saw a person strike another and kill him ; that is testimonial evidence of the fact of murder. But it is possible to have circumstantial evidence of the fact of murder; that is to say, you may find a man dying with a wound upon his head having exactly the form and character of the wound which is made by an axe, and, with due care to take surrounding circumstances into ac- count, you may conclude with the utmost certainty that the man has been murdered — is dying in consequence of the violence inflicted by that implement. We are very much in the habit of considering cir- cumstantial evidence as of less value than testimonial evidence, and it may be in many cases, where the circumstances are not perfectly clear and perfectly intelligible, that it is a dangerous and uncertain kind of evidence ; but it must not be forgotten that in many cases it is quite as good as testimonial evidence, and that not unfrequently it is a great deal better than testimonial evidence. For example, take the case to which I referred just now. The circumstantial evidence is better and more convincing than the testimonial evidence, for it is impossible, under the circumstances that I have mentioned, to suppose that the man had met his death from any cause but the violent blow of an axe wielded by another man. The circumstantial evidence in favor of a murder having been committed, in that case, is as complete and as convincing as evidence can be. It is evidence which is open to no doubt and no falsification. But the testimony of the witness is open to multitudinous doubts. He may have been mistaken. He may have been actuated by malice. It has constantly happened that even an accurate man has declared a thing has happened in this, that, or the other way, when a careful analysis of the circumstantial evidence has shown that it did not happen in that way, but in some other way. Now we must turn to our three hypotheses. Let me first direct your attention to what is to be said about the hypothesis of the eter- nity of this state of things in which we now are. What will first strike you is, that that is an hypothesis which, whether true or false, is not capable of verification by evidence ; for, in order to secure testi- PROFESSOR HUXLEY'S LECTURES. 49 mony to an eternity of duration, you must have an eternity of wit- nesses or an infinity of circumstances, and neither of these is attain- able. It is utterly impossible that such evidence should be carried beyond a certain point of time, and all that could be said at most should be that there was nothing to contradict the hypothesis. But when you look, not to the testimonial evidence — which might not be good for much in this case — but to the circumstantial evidence, then you find that this hypothesis is absolutely incompatible with that cir- cumstantial evidence, and the evidence is of so plain and so simple a character that it is impossible in any way to escape from the conclu- sions which it forces upon us. IDEAL SECTION OP THE CRUST OF THE EARTH. -—Post-tertiary and Recent. — Pliocene. — Miocene. — Eocene. Cretaceous. •—Jurassic or Oolitic. — Triassic (New Red Sandstone). •— Permian. .... Carboniferous. — Devonian or Old Red Sandstone. — Silurian. — Cambrian. — Huronian. — Laurentian. B J m1 -> -.a • W ft. - t . ^^^mmm °IgSoI°r°I c .'£?■*« T *' : "