National Museums National Museum of Canada of Natural Sciences Ottawa 1975 Publications in Biological Oceanography, No. 8 A New Species of Centrolophid Fish from Easter Island and Rapa Iti Island in the South Pacific Don E. McAllister and John E. Randall . OF at Oe | ‘¥,— 5 | ~~ FU ITS Publications d’Océanographie biologique, n°8 Musées nationaux Musée national des du Canada Sciences naturelles A New Species of Centrolophid Fish from Easter Island and Rapa Iti Island in the South Pacific National Museum of Natural Sciences Publications in Biological Oceanography, No. 8 Published by the National Museums of Canada Staff Editor Bonnie Livingstone Musée national des Sciences naturelles Publications d’Océanographie biologique, n°8 Publié par les Musées nationaux du Canada A New Species of Centrolophid Fish from Easter Island and Rapa Iti Island in the South Pacific Don E. McAllister and John E. Randall ©Crown copyrights reserved National Museum of Natural Sciences National Museums of Canada Ottawa, Canada First quarter 1975 Catalogue No. NM95-7/8 Available by mail from the National Museum of Natural Sciences Ottawa, Ontario K1A 0M8 P0987654321 Y798765 Printed in Canada ©Droits réservés au nom de la Couronne Musée national des Sciences naturelles Musées nationaux du Canada Ottawa, Canada Premiére trimestre 1975 N° de catalogue NM95-7/8 L’editeur remplit les commandes postales adressées aux Musée national des Sciences naturelles Ottawa, Ontario K1A O0M8 T0987654321 A798765 Imprimé au Canada Contents List of Figures, vi Summary, vii Résumé, vii Resumen, vii Biographical Note, viii Acknowledgements, ix. Introduction, 1 Taxonomy, 1 Discussion, 4 References Cited, 6 vi List of Figures 1 Lateral, dorsal and frontal views of the holotype of Schedophilus labyrinthicus n.sp., 1 2 Schedophilus labyrinthicus n.sp., 2 3 Schedophilus ovalis (Valenciennes), 4 Summary Four specimens from the marine shore waters of Easter Island (Isla de Pascua) and Rapa Iti Island in the South Pacific Ocean are referred to the genus Schedophilus and described as a new species, Schedophilus labyrinthicus. The holotype is deposited in the National Museum of Natural Sciences, Ottawa. Résumé Quatre spécimens provenant des eaux cétiéres de I’ile de Paques (Isla de Pascua) et de l’ile de Rapa Iti, dans le sud de l’océan Pacifique, et se rattachant au genre Schedophilus ont servi a décrire une nouvelle espece, Schedophilus labyrinthicus. L’holotype est déposé au Musée national des Sciences naturelles, a Ottawa. Resumen Se describen cuatro ejemplares procedentes de las aguas maritimas de la Isla de Pascua y de la Isla de Rapa Iti, en el Sur del Océano Pacifico en el género Schedophilus, como una especie nueva Schedophilus labyrinthicus. El holotypo esta en el Museo Nacional de Ciencias Naturales, Ottawa. vii viii Biographical Note Don E. McAllister Don E. McAllister, Curator of Fishes at the National Museum of Natural Sciences, Ottawa, since 1958, is also a lecturer at the University of Ottawa. He was born in Victoria, B.C. and received his Ph.D. from the University of British Columbia in 1964. His studies have taken him to such places as Newfoundland, Quebec, the Northwest Territories, British Columbia, and Japan. He has published over 65 scientific papers, including studies on Arctic fishes, sculpins, smelts, fishes found in archaeological sites, light organs in fishes, the classification of bony fishes, and a popular guide, The Freshwater Sport Fishes of Canada. His professional memberships include the Canadian Society of Wildlife and Fishery Biologists and the American Society of Ichthyologists and Herpetologists. His hobbies include photography, skin diving, cross- country skiing and bonsai. John E. Randall John E. Randall has been an ichthyologist with the Bernice P. Bishop Museum, Honolulu, Hawaii, since 1966. He was born in Los Angeles in 1924, did his undergraduate work at the University of California, Los Angeles, and received his Ph.D. at the University of Hawaii in 1955. A Yale University-Bishop Museum fellowship enabled him to spend more than a year in ecological research on fishes in Tahiti. From 1957 to 1965, he worked for the University of Miami, spending from 1963 to 1965 directing ’a marine biological survey of St. John, Virgin Islands. From 1961 to 1965, he was a professor of biology at the University of Puerto Rico; from 1962 he also served as Director of the University’s Institute of Marine Biology. He has published over 130 articles, chiefly on the systematics and biology of tropical marine fishes. Acknowledgements The authors are grateful to Dr. lan A. Efford of the University of British Columbia for making the specimens from the Canadian Expedition to Easter Island available for study. Dr. Per Pethon of the Universitet i Oslo kindly provided counts and measurements on the holotype of Seriolella christopherseni. Staff of the British Museum (Natural History) assisted Dr. Don E. McAllister during his visit to examine specimens. Dr. M. Stehmann, Institut fur Seefischerei, Hamburg, generously made his and Dr. W. Lenz’s manuscript available prior to publication. Drs. Frederick H. Berry and William D. Anderson, Jr., of Charleston, South Carolina, permitted reference to their undescribed new species from the Pacific coast of Costa Rica. Dr. Richard L. Haedrich of Woods Hole Oceanographic Institution and C. G. Gruchy of the National Museum of Natural Sciences, Ottawa, criticized the manuscript. Charles H. Doug- las made the drawing. T. A. Willock, J. Aniskowicz and the late W. H. Van Vliet took X rays of the specimens. The National Geographic nga ag ate financial support for the collections made by Dr. John . Randall. Opee eceins ups et to mots Ane @ of whatetg ew erortiun-arft Pvoens PMC OM. Oe tert. arntapacnds quan sot iid Plate an! Jo notiotiee! @G . aweanhecelane tidalatan |. Pea oled pase RAM aH aEoY DEE Ghwen- dekiyom, Bothhroles - eqenplet sewn mvCOUM He qelsc HP siege odeats ina ainneg Poe oe ee Sa ane: .Chbetrteas yr: VMN AaY BIT ET 8 goureearh ater ts rmernet oS aM UNG PAID, 8 HOG, ads An ine Mranrtis Cok haan FeO AARNE), 205 Were... dine att Snecma raat attr aelenysayen | vodhacsbou edt abagermetan Bettrnend Baile ee ert: less fea og sat Me vb aioilt, rob eal olen: rodanoo antiaes ai CARBUM Aa l) OFk hay vue? 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ALL i af Pv ! lee 4 oo avtorm =, _Oe 7 t : 7 . 5 a ges Aytg sree 1a rr nig PD VINVor any rowel wy 1995 A fae Linivertdiy-Bienay Muaesuih (¢!! yp Ehoned 7 ONG TREND The 5 ee ¢a>' « ' 7 Ys ry @ 4) 4" 3 st . ~ 9 1a Ap ue 6 Temes) ML y iy ouewT 7 ‘ d roe 965 if - ; ‘ A aoe Dec &@ VOT) Pa vey ( St wu (lars A i » *O8 ) ' ' Uri ef a Poors * Som (pez | ’ ats (UAn C) (Aare < , ’ ’ Ny Vie Seo “teia ‘ i™ ria - a ' » 7 ee : ' e ve . st } Min ad ATS if re LJ 7 od Eins t ; ' e : _ a iy fal rs Rographicat Mote | Introduction The centrolophid genus Schedophilus Cocco, 1839 (sensu Haedrich and Horn 1972; Stehmann and Lenz 1972) is known from the waters of the North and South Atlantic, South Africa, the southern Indian Ocean, southeast Asia, Australia, and New Zealand (Haedrich and Horn 1972). This paper reports on the first specimens of an undescribed species of Schedophilus from Easter Island (Isla de Pas- cua), a Chilean territory, and Rapa Iti Island, a French territory, in the South Pacific Ocean. Haedrich (1967), Haedrich and Horn (1969, 1972) and Stehmann and Lenz (1972) treat the systematics of Schedophilus. Drs. lan A. Efford and John A. Mathias col- lected two specimens of Schedophilus on the 1964-65 Canadian Medical Expedition to Eas- ter Island. One specimen, the holotype, lacks half the caudal fin; the other, a paratype, has two deep cuts in the nape. Aside from frayed fins and some lost scales, these two specimens are in satisfactory condition. Two other speci- mens in good condition, one from Easter Island, the other from Rapa Iti Island, were obtained by Dr. John E. Randall. Taxonomy ORDER PERCIFORMES SUBORDER STROMATEOIDEI! FAMILY CENTROLOPHIDAE GENUS Schedophilus Cocco, 1839 Schedophilus labyrinthicus n.sp., Figure 1 Holotype NMC67-501: 502 mm in standard length; from an unascertained locality near Easter Island; obtained from islanders by lan A. Efford and Jack A. Mathias, 22 December 1964; deposited in the ichthyological collection, National Museum of Natural Sciences, Ottawa. Paratypes BC65-411: 477 mm in standard length; from Hanga Pico, Easter Island; collected by lan A. Efford and Jack A. Mathias; deposited in the ichthyological collection of the Institute of Ani- mal Resource Ecology, University of British Columbia, Vancouver. BPBM 6642: 525 mm in standard length; from Easter Island, in about 170 fathoms; obtained from fisherman by John E. Randall, 11 Feb- ruary 1969; deposited in the Bernice P. Bishop Museum, Honolulu, Hawaii. Drawing by C. H. Douglas Figure 1 Lateral, dorsal and frontal views of the holotype of Schedophilus labyrinthicus n. sp. Caudal fin resto- ration from University of British Columbia paratype. The upper two rays of the pectoral are broken. Figure 2 Schedophilus labyrinthicus n. sp. Top: holotype. Island specimen 12253. Note larger, more ovate eyes Centre: University of British Columbia Easter Island of the two bottom specimens. (Photographs by T. A. paratype 65-411 and Bishop Museum Easter Island Willock and Don E. McAllister) paratype 7323. Bottom: Bishop Museum Rapa Iti Taxonomy Other material BPBM 12253: 512 mm in standard length; from Rapa Iti Island; collected by Alfred Make, March-August 1971: deposited in the Bernice P. Bishop Museum, Honolulu, Hawaii. Diagnosis Orbit 20-26 per cent of head length; snout 7-8 per cent and snout to anus 53-54 per cent of standard length; gill rakers comprising .68-.83 of longest gill filament; dorsal spines VII-VIII; dorsal rays 26-29; anal rays 18-19; vertebrae (abdominal + caudal) 10+15; pyloric caeca 4-8; digitiform; scales slightly deciduous; dor- sal spines becoming longer posteriorly and graduating into the soft dorsal rays (last spine about 2/3 length of first soft ray); 2 free inter- neurals. Description In counts and proportions below, the holotype is mentioned first, the University of British Colum- bia paratype next, and the Bishop Museum paratype 6642 and specimen 12253 next. Dorsal spines and rays, VII,27, Vil,27, VIII, 26, VII,29; anal, 111,19, I1,19, II,18*, 111,18; ventral, 1,5, 1,5, 1,5, 1,5; pectoral, i21, i22, i21, i20; pored lateral line'scales, ca. 96, ca. 92, ca. 91, ca. 96; scales above the lateral line, 20, 21, 21, 20; scales below the lateral line, 38, —, 37, 37; branchiostegals, 7, 7, 7, 7; gill rakers, 7+1+16, 7+14+17, 7+1+18, 7+1+16; verte- brae (abdominal + caudal), 10+15, 10+15, 10+ 15, 10+15 (including urostyle). Head short, snout blunt and almost vertical, head length 3.8, 3.6, 3.4, 3.5 in standard length. Jaws short, extending to just under front edge of orbit, but with gape not reaching orbit. Upper jaw lacking frenum, slightly protractile, with su- pramaxillary. Angle of jaws about 35° to hori- zontal axis in holotype. Single row of small teeth in each jaw; palate and floor of mouth toothless. Well-developed flexible teeth on pharyngeal bones above and below. Orbit slightly or very oval, with longest axis vertical; horizontal diameter 4.7, 5.0, 4.1, 3.9, in head length. Preoperculum denticulate at and above angle, jutting posteriorly up to 1/2 orbit diameter. Interoperculum and lower suboperculum finely denticulate. Lower jaw, preoperculum, cheek, snout, and top of head posterior to the nape, with lateral line network opening to surface through numerous tiny pores. Gill rakers slen-. der, toothed on dorsal edge; length of longest is * An X ray shows one ray missing, so the anal count is given as III,18, rather than the apparent III,17. .75, .68, .83, .75 of longest gill filament. Cheek, interoperculum, and operculum with scales. Spinous dorsal begins 1/2 orbit diameter in front of end of operculum; spines graduated, lower than rayed dorsal but continuous with it. Rayed dorsal ends in front of end of rayed anal. Pectoral long, falcate, almost reaching to above vent. Pelvics shorter than pectorals, attached to abdomen by a short membrane and folding into shallow groove, origin about 4 mm posterior of a vertical through the pectoral origin. Anal and dorsal spines stiff, stout, first spine buried in skin. Caudal peduncle slender. Caudal fin well- forked. Vent in a shallow groove, its centre about 28 mm anterior to origin of anal fin, slightly behind midpoint of body in the holotype. Scales moderate sized, cycloid, extending onto base of fins, cheek, operculum and isthmus, not highly deciduous. In the holotype and the Bishop Museum 7323 (Figure 2) the lateral line follows the dorsal profile and straightens out horizontally on the urosome above the posterior quarter of the anal fin; due to twisting of the urosome the latter feature cannot be deter- mined on the U.B.C. paratype. Pyloric caeca large, digitiform, 8, 6, 4, 7. All specimens have two free interneurals before the dorsal fin. Pre- served colour dark brown dorsally, yellowish below. Fins without dark markings. Buccal and branchial cavities light. Peritoneum light with. fine speckling. The eye coloration is visible only in the Bishop Museum 7323, where its iris is golden, appearing as a complete ring undivided by a bar. Etymology The specific name /abyrinthicus is derived from the Greek AaBupuv Gos, or maze, in reference to the labyrinthine complex of the cephalic lateral- line sensory system. Discussion Stehmann and Lenz (1972) have rechar- acterized the Schedophilus ovalis species- group as having two free predorsal interneurals as opposed to three in the others of the genus and in the related genera Seriolella and Hyperoglyphe. We follow these authors in the generic assignment of the Easter and Rapa Iti specimens to Schedophilus. The Easter and Rapa Iti specimens can be immediately distinguished from several other species in the genus by their fewer vertebrae. Schedophilus huttoni (Waite, 1910) from New Zealand and Australia has 12+18-19, S. griseolineatus (Norman, 1937) from off Argen- tina has 10+ 16 (and numerous pyloric caeca), and S. maculatus Gunther, 1860, from the Southern Ocean has 12+17 (counts from Haedrich and Horn 1972), whereas the Easter and Rapa Iti specimens have only 10+ 15 ver- tebrae. S. medusophagus Cocco, 1839, from the Mediterranean and North Atlantic is sharply distinguished from our specimens by posses- sion of 44-50 dorsal elements (spines and rays) as opposed to 34-36, S. pemarco (Poll, 1959) by its 12-14 as opposed to 16-18 gill rakers on the lower gill arch and its lower dorsal and anal ray counts, S. haedrichi Chirichigno, 1973, by its fewer dorsal rays, 22-23, as op- posed to 26-29, fewer anal rays, 15-16, as opposed to 18-19, and its longer pre-anal and head lengths (Chirichigno 1973). Our species differs from an undescribed new species of the genus from the Pacific coast of Costa Rica by its fewer dorsal elements IV,22 as opposed to VI-VII,26-29, fewer anal elements Ill,15 as opposed to IIl,18-19, and 3 free interneurals as opposed to 2 (F. H. Berry and W. D. Ander- son, Jr., MS). Of the currently accepted forms in the genus, this leaves only the Schedophilus ovalis com- plex. Stehmann and Lenz (1972) synonymized S. christopherseni (Sivertsen 1946) from Tris- tan da Cunha, and suggested the probable synonymy of S. velaini (Sauvage 1879) of St. Paul Island in the southernmost Indian Ocean, and the Easter and Rapa Iti islands material with S. ovalis (Valenciennes, in Cuvier and Valenciennes 1833), which was originally de- scribed from the Mediterranean and extended by later authors to the eastern and central Atlantic. However, noting certain patterns in the data, we place a different interpretation on them. The 12 specimens of S. ovalis in the Mediterranean, central and eastern Atlantic in the Northern Hemisphere for which we have data (BMNH 1953.11.1.529, Figure 3; BMNH 1955.19-19.317; BMNH 1925.1.8.3; an un- numbered BMNH Madeiran specimen; plus data in Stehmann and Lenz 1972; Steindachner 1868; Cuvier and Valenciennes 1833) have 21-23 anal rays and 5-6(7) gill rakers on the upper arch, whereas all specimens known from the Southern Hemisphere (St. Paul, Tristan da Cunha, Easter, and Rapa Iti islands) have 18-20 anal rays and 6-7 gill rakers on the upper arch. Further, the Northern Hemisphere specimens are deeper, the depth comprising .34-.42 of standard length as opposed to .29-.35 for the Southern Hemisphere speci- mens and the caudal peduncle depth being .098-.121 as opposed to .078-.090 of the stan- dard length. The northern specimens range from 144 to 452 mm in standard length, whereas the southern ones range from 477 to 525 mm in standard length. It is possible, there- fore, that the proportional differences may be Figure 3 Schedophilus ovalis (Valenciennes), BMNH 1953.11. 1.529, from Funchal market, Madeira. Note the deeper body and caudal peduncle. (Photograph by Don E. McAllister) Discussion attributable to negative allometric growth, al- though it seems odd that the northern ones should all be smaller than the southern ones. The meristic differences remain, however. Stehmann and Lenz (1972) suggest that, since the meristic variation in the better-known Seriolella punctata (45 specimens) equals or exceeds that amongst the various Schedophilus ovalis samples (Sensu /ato, 16 specimens), the latter complex should be re- garded as conspecific. Although the logic is not impeccable, the argument is persuasive. To apply the limits of variation of one genus to another, even though the fishes are closely related, may not always be justified. More im- portantly, it overlooks the geographical pattern in the data, northern populations with 21-23 anal rays, and southern with 18-20. Chi-square tests, with and without Yates correction, show the Northern Hemisphere specimens to be significantly different from the Southern Hemi- sphere specimens at the p = <.001 level. We conclude, therefore, that the name Schedophilus ovalis (Valenciennes) should be applied only to those populations of Sche- dophilus in the warmer waters of the central and eastern Atlantic Ocean and the Mediterra- nean Sea that have 10+15 vertebrae and 21-23 anal rays. 7 This leaves us with three allopatric popula- tions in the Southern Hemisphere whose status must be discussed: the Tristan da Cunha S. christopherseni, the St. Paul S. velaini, and the Easter and Rapa Iti islands populations. The standard lengths of the samples are 505 mm, 550 mm, and 477-525 mm respectively. Unlike the Easter and Rapa Iti specimens, both S. christopherseni and S. velaini have 20 anal rays instead of 18-19 and an orbit to snout ratio of 1.14-1.16 instead of .70-1.00. Further, S. christopherseni has 6 dorsal spines instead of 7 or 8, only 85 pored lateral-line scales instead of 91-96, and rounded instead of pointed tips to the caudal fin lobes. S. velaini has a crescentic anal fin, with the first soft ray twice the length of the last as opposed to 1.5 times, or less, the length of the last soft ray in the Easter and Rapa Iti material. S. christopherseni and S. velaini appear morphologically similar, and both in- habit oceanic islands near the northern limit of drift ice and within the seasonal oscillation of the Antiboreal Convergence (Ekman 1953). Easter and Rapa Iti islands, on the other hand, are 900 and 1400 miles respectively beyond the northern limit of drift ice and over 600 miles north of the Antiboreal Convergence. The Eas- ter and Rapa Iti populations, therefore, appear morphologically and ecologically distinct from S. velaini and S. christopherseni and are here recognized as a new species, Schedophilus labyrinthicus. The four specimens of Schedophilus labyrin- thicus are fairly homogeneous with respect to meristic and most mensural characteristics, but the Bishop Museum paratype from Easter Is- land does differ in having a larger orbit (4.1 instead of 4.9-5.0 in head) and a larger head (3.4 instead of 3.6-3.8 in standard length). The specimen from Rapa Iti Island differs most in having the snout less declivous, 29 instead of 26-27 soft dorsal rays and the orbit diameter 3.9 instead of 4.1-—5.0 in head; it is most like the Bishop Museum Easter Island paratype. Be- cause of these differences we do not give the Rapa Iti specimen paratype status. We were struck by the variation in shape as well as by the size of the orbits. The left orbit is small and circular in the holotype and in the U.B.C. paratype, large and subcircular in the Bishop Museum Easter Island specimen, and large and ovate in the Rapa Iti specimen. However, the contralateral orbit of the Bishop Museum Easter Island specimen is ovate, that of the Rapa Iti specimen is large and oval, yielding orbit into head proportions for left and right orbits of 4.1 and 3.8 for the Bishop Museum Easter Island specimen and 3.9 and 3.4 for the Rapa Iti specimens (based on maximum dimension). Because of the bilateral variation in individual specimens, we do not wish at the moment to over-emphasize the differences from specimen to specimen, although, in the future, additional material may reinforce the significance of such differences. We also noted variation in orbit shape of Schedophilus ovalis (sensu Stricto). The left orbit of BMNH 1953.11.1.529 was small and circular while that of BMNH 1955.19-19.317 was large and oval (both from Madeira). An ophichthid eel about 295 mm in total length was found in the body cavity between the gut and gas bladder of the Rapa Iti specimen. The eel was bent in a U-shaped position, and appeared to be enclosed in a membrane. As its coloration was normal, it would appear to have burrowed out of the stomach before digestion began. Walters (1955) discussed other exam- ples of ophichthid eels found in the coelomic cavity of predacious fishes; he called them pseudo-parasites. The Rapan name for Sche- dophilus is ‘‘paipai’. References Cited Chirichigno, Norma (1973). Nuevas especies de peces de los generos Mustelus (Fam. Triakidae), Raja (Fam. Rajidae) y Schedophilus (Fam. Centrolophidae). Inst. Mar Peru (Callao) Informe (42):1-40, 8 figs. Cuvier, G., and A. Valenciennes (1833). Histoire naturelle des poissons. Vol. 9. F. G. Sevrault, Paris. 379 pp. Ekman, Sven (1953). Zoogeography of the sea (Transl. from Swedish by Elizabeth Palmer.) Sidgwick and Jackson. London. 417 pp. Guichenot, Alphonse (1848-49). Fauna Chilena: Peces. Pages 137-372, in Claude Gay, Historia fisica y politica de Chile. Tom. 2, Repitilia and Pisces. Chile en el Museuo de historia natural de Santiago, 1844-71. Paris. Haedrich, Richard L. (1967). The stromateoid fishes: systematics and a classification. Mus. Comp. Zool. Bull. 135(2):31-139, 56 figs. Haedrich, Richard L., and Michael H. Horn (1969). A key to the stromateoid fishes. (Unpublished manuscript.) Woods Hole Oceanographic Institution, Ref. No. 69-70. 46 pp., 7 figs. (1972). A key to the stromateoid fishes. (Unpublished manuscript.) Rev. ed. Woods Hole Oceanographic Institution. 46 pp. 7 figs. Miranda Ribeiro, Alipio de (1915-18). Fauna Brasiliense: Peixes. Tomo V. Elev- therobranchios, Aspirophoros, Physoclist. Archivos de Museu Nacional de Rio de Janeiro, vol. 21. 227 Pp. Regan, C. Tate (1902). A revision of the fishes of the family Stromateidae. Ann. Mag. Natur. Hist. Ser. 7, vol. 10: 115-31, 194-207. Rendahl, Hialmar (1921a). The fishes of the Juan Fernandez Islands. Pages 49-58 in C. Skottsberg, ed., The Natural History of Juan Fernandez and Easter Islands, vol. 3, pt. 1. Almgrist and Wiksell, Uppsala. (1921b). The fishes of Easter Island. Pages 59-68 in C. Skottsberg, ed., The Natural History of Juan Fer- nandez and Easter Islands, vol. 3, pt. 1. Almgrist and Wiksell, Uppsala. Ringuelet, Raul A., and Raul H. Aramburu (1960). Peces marinos de la Republica Argentina. Agro (Buenos Aires) 2(5):1-141, 72 figs. Sauvage, H. E. £ (1879). Mémoire sur la faune ichtyologique de I'lle Saint-Paul. Arch. Zool. Exp. Gen., Notes Rev. 8:1-46, pl. 1-3. Sivertsen, Erling (1945). Fishes of Tristan da Cunha, with remarks on age and growth based on scale readings. Result. Nor. Sci. Exped. Tristan Cunha 1937-38, no. 12:1-44, 8 pl., 17 figs. Stehmann, M., and W. Lenz (1972). Ergebnisse der Forschungsreisen des FFS “Walther Herwig” nach Sidamerika: XXVI. Syste- matik und Verbreitung der Artengruppe—Seriolella punctata (Schneider, 1801), S. porosa Guichenot, 1848, S. dobula (Gunther, 1869)—sowie taxo- nomische Bemerkungen zu Hyperglyphe Gunther, 1859 und Schedophilus Cocco, 1839 (Osteichthyes, Stromateoidei, Centrolophidae). Arch. Fischereiwiss. 23(3):179-201, 8 figs. Walters, Viadimir (1955). Snake-eels as pseudoparasites of fishes. Copeia 1955(2):146-47. Table 1 Measurements of Schedophilus labyrinthicus n. sp., in millimetres/hundredths of standard length Holotype Paratype Paratype Easter Is. Easter ls. Easter Is. Rapa Iti Is. NMC67-501 BC65-411 BPBM6642 BPBM12253 Standard length 502/100 477/100 525/100 512/100 Greatest body depth 170/34 158/33 162/31 150/29 Head length 132/26 131/28 155/30 148/29 Snout to centre of anus 269/54 251/53 277/53 277/54 Pelvic fin to anal origin 156/31 119/254 151/29 141/28 Horizontal orbit diameter 28/6 26/6 38/7 38/7 Snout length 37/7 37/8 39/8 38/7 Upper-jaw length 45/9 43/9 52/10 49/10 Pectoral-fin length 108+/22+° 114+/24+» 136/26 128/25 Pelvic-fin length Se/t2 65/14 66/12 64/12 Caudal-peduncle depth 39/8 42/9 46/9 46/9 Caudal-peduncle length 89/18 86/18 83/16 75/15 Caudal-fin iength - 112/23 - - Longest gill raker 17/3 14/3 19/4 17/3 Longest gill filament 22/4 20/4 22/4 23/4 @ Foreshortened by contraction following cutting of nape muscles. > Tips of pectorals of both specimens broken off. Errata The last sentence on page 4, column 1, should read: An undescribed new species of the genus from the Pacific coast of Costa Rica differs from our species by its fewer dorsal elements IV,22 as opposed to VI-VII,26-29, fewer anal elements Ill,15 as opposed to IIl,18-19, and 3 free inter- neurals as opposed to 2 (F. H. Berry and W. D. Anderson, Jr., MS). Where it appears on pages 2 and 3, BPBM 7323 should read: BPBM 6642. 7 late i. WV : 1 a 5 ia i) j ' ‘ 7 2 oe ft } } 7 4 *~ \ i A 7 ‘ 4 P ) a “ 7 ' eer g i TS BTaS fy Wo ye 20) A ete He , | vie ' ¥ Wi, he On er iy yates 6 regen weve ty an Vea { wy - bi dies y ' 7 Leena LN pllihating ty Ancisny toy ; ; an) we eon aT: an POT 4 * » ; j Wie? n ‘ ‘ shag a , je yt ‘ St WY Pik ‘ 4 ‘ ' , ioe peaely , wis ee | ae a Tete : Niky = ‘ - - = war A ; . 1 +’ % as) 7 : A : eos , ‘ , PS me os et VMS emaahe ’ 5» Mee! a] - . te : 7s ° Z P 7 om . 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