Bot. Serials QK1 P87 P toe yh ht BA a Svea ay ae ays | ee t Th al ae ' 4 ” iy ’ j p wis te y ao we 1 ‘aie rt Wins i ; d ' ) 3} ' >» Z ‘eT ih “ ie y e i de) f Pa 4 - +] oF h i 4 vig at 1 we i {'oeey ‘ Ae ’ ~« ® We: : a | ¥ 5 a! bh ah a , v ¥ 4 rT 4 7 ? : } ‘ y i 49) 4 ie j 7 7 Pr i! 4 . f Did Me he | ae 4% av ike ae . ‘ } , ‘ ‘| iy / 4 yi aa wi } . ‘Ey ee ae ie ‘ ' dje wee i oy ED er mete ‘ot . Ri ys rey) » ‘ Mm i ‘ 1 Wa ‘he - a oe a) a si ay Jak @ 87 wry © hued iy a ae Saad | 7 2 ' ey ae 7. DA V UR esi Sy Oe e Pe) Be 4 a, ‘ i. 2 th Materials for a Flora of Central Yukon Territory National Museum of Natural Sciences Publications in Botany, No. 4 Published by the National Museums of Canada Staff editor Frances Smith Musée national des Sciences naturelles Publications de botanique, n° 4 Publié par les Musées nationaux du Canada Materials for a Flora of Central Yukon Territory A. E. Porsild ©Crown copyrights reserved National Museum of Natural Sciences National Museums of Canada Ottawa, Canada First quarter 1975 Catalogue No. NM95-9/4 Available by mail from the ; National Museums of Canada Marketing Services Ottawa, Ontario K1A OM8 P0987654321 Y798765 Printed in Canada ©Droits réservés au nom de la Couronne Musée national des Sciences naturelles Musées nationaux du Canada Ottawa, Canada Premier trimestre 1975 No de catalogue NM95-9/4 L’éditeur remplit les commandes postales adressées aux Musées nationaux du Canada Service de distribution Ottawa, Ontario K1A O0M8 T0987654321 A798765 Imprimé au Canada Contents List of Plates, vii Résumé, viii Summary, ix Preface, xi Annotated List of Selected Species, 1 Bibliography, 75 aur ~ ’ 2 nge sayy! Muncie of Moeturet Sc shige le Minho sateen): ; Le fai eS | ne —_ ase TAS es eS a Mi a. . iia. Comene Ofttawe, Cangce. xi sperensé vows een oe a re A vaste Sr ha iene Kin . oe fi ah | Win. bx Ba0ter4 = CWI Per tne aan ~~ Mmahertiptarpee NMGG-BI4 15 ae ns 4 r tatsiosge heioete? to hal) betatommnA _ af q ewe 6 oe - — — i ok ‘ : ; ty {! ¢) Wail i, ee yi Clatlag! ty! i) pn 7 oi os gc; mae wen @ BR i a ig i a a <> or! | ar: os , That ae hae sad Akira Ae M4) 4 rrmerstiny scwti A 5 Se Cre Ow io “owe Sara ave yi wa ie : ; a > _ 4 1 a7 | F a were Pi ® aids : Pietra? | i a7. eunne au Cétiag 9) + - fo I a ~a4 © e “4 ay 7 ow J : : i : y 7 ‘ J a ; . Pn ae * Fo j » 4 A : v ine fi | Pe ; = + > ae ~ ry a ioe ae es) < ° List of Plates 1 Calamagrostis Robertii n. sp., 6 2 Claytonia arctica Adams, 21 3 Melandrium apetalum (L.) Fenzl. ssp. ogilviense n. ssp. (flowering specimens), 22 4 Melandrium apetalum (L.) Fenzl. ssp. ogilviense n. ssp. (fruiting specimen), 24 5 Anemone multiceps (Greene) Standley, 28 6 Draba barbata Pohle, 32 7 Draba yukonensis n. sp., 36 8 Saxifraga rufopilosa (Hult.) n. stat. (flowering specimens), 42 9 Saxifraga rufopilosa (Hult.) n. stat. (fruiting specimen), 43 10 Potentilla Ledebouriana Porsild, 44 11 Oxytropis nigrescens (Pall.) Fisch. var. lonchopoda Barneby, 48 12 Antennaria Ellyae n. sp., 62 13 Taraxacum mackenziense n. sp., 72 vil Vili Resume L’inaccessible forteresse montagneuse située au nord de la Colom- bie-Britannique et au Yukon est demeurée longtemps l’une des moins connues des régions du Canada. II existe maintenant des cartes exactes, tracées a l’aide de photographies aériennes, de cer- taines parties de ce territoire, mais l’absence de chemins et de lieux d’atterrissage convenables, méme pour de petits avions, ont rendu trés difficiles l’exploration de cette région et l’étude de sa faune et de sa flore. . Depuis 1944, plusieurs realisations de voirie importantes ont per- mis d’atteindre certaines contrées montagneuses qui jusque-la n’étaient accessibles qu’aux expéditions munies de bétes de somme. Ces lieux se situent en partie au sud-est du Yukon, en bordure de la route Canol, et en partie le long des trongons de la route de |’Alaska traversant le méme territoire (voir la bibliographie). L’étude se rapporte principalement a la flore des monts Ogilvie devenus accessibles depuis la construction de la route Dempster, commencée en 1962 et qui réunira éventuellement le centre du Yukon avec Inuvik et le delta du Mackenzie. L’ensemble des renseignements compilés provient des collec- tions d’herbier faites principalement dans les monts Ogilvie. Parmi les 428 especes de plantes vasculaires que |l’auteur €numére ou décrit, 11 seraient des taxa nouveaux, 6 apparaitraient pour la pre- miére fois dans la flore du Canada, une serait une addition nouvelle a la flore de l’Amérique du Nord et au moins 38 se trouvent notées pour la premiére fois en provenance du Yukon. Summary The mountain fastnesses of northern British Columbia and Yukon Territory have long remained among the least Known parts of Can- ada. Accurate maps constructed from aerial photographs are now available for some of these parts, but the absence of roads and the scarcity of suitable landing places, even for small aircraft, have made ground exploration, including the study of plant and animal life, difficult. Since 1944, several major road projects have given access to mountainous regions in the Yukon hitherto accessible only by pack- horse trails. Among these regions are parts of southeastern Yukon adjacent to the Canol Road, as well as portions of the Yukon section of the Alaska Highway (see Bibliography for additional references). The present report deals mainly with the flora of the Ogilvie Moun- tains, now made accessible by the construction of the Dempster Highway, which was started in 1962, and when completed will con- nect central Yukon with Inuvik in the Mackenzie Delta. The bulk of information reported in the catalogue following has been obtained from collections of plants mainly from the Ogilvie Mountains. Among the 428 species of vascular plants listed or dis- cussed, 11 are believed to be ‘‘new” or heretofore unrecognized taxa; 6 are believed to be “‘new’”’ to the flora of Canada; one is ‘‘new”’ to the flora of North America; and no less than 38 are reported for the first time from the Yukon Territory. M 7 i a sabia en. nats hn, on, (VS, So _ " Oe’ ne ray i create iw soars ; Sitics re te org bows yl Yongec ty an a ae, i evad gioplorg tak ae inven, rn a. a | pod Ny aay ie sbieeoush ¢ ig be i dh Mauonisteie err) } | anil ha AHH Oe Bey: ao Lele eer h alae ad Pye GHhe ener (DH GE ior euIN de .o — alteotiy'ta UNE Bee ORT gia Geld Si af . nar te ise Cah Sa aVineke lees. | sta on WG sd eben won (eniat : wie RAw nsw rath ams feat tyaherart octet = wi yee Ain ney Nic oma =a) el rey 16 tether Me “GHODMio® matt beniaidg teed — oS a4 ck ie STROMA Sy et das: weet Gusltrts yeea ae é : F<. $. us tore os Ppt sue hu he et 90 men is pete notéee 7 SGur te pretties tote on prover anos du Yukon, ae ¥ 2 Preface When the Dempster Highway in the Yukon Territory is completed, it will provide access to what is probably the largest remaining botani- cally unexplored part of Canada.* Only the area adjacent to the southern part of the highway has been partly explored botanically, and the results reported in this paper indicate that the flora of the central and unglaciated Yukon Plateaut is rich and, phytogeographi- Cally, very interesting. Construction of the Dempster Highway was started in 1962. When completed, it will connect central Yukon with Inuvik on the east channel of the Mackenzie Delta, a distance of roughly 400 miles. The Dempster Highway branches off the main Dawson—Whitehorse Highway 26 miles east of Dawson. For approximately 44 miles it follows the wooded North Klondike River valley, climbing to about 4,000 feet in the North Fork Pass, which is on the watershed between the Yukon and Mackenzie drainages. From the pass there is an im- pressive view of some of the highest peaks of central-eastern Yukon Territory. Northeast of the pass is ‘Sheep Mountain” (elevation 6,650 feet), and beyond it the slightly lower “Herringbone Moun- tain”. Beyond the pass, the highway descends to approximately 3,500 feet into the valley of an unnamed minor tributary of the Black- stone River that joins the main Blackstone River near Chapman Lake at Mile 68. The floor of both valleys is unforested, but here and there, in sheltered places, solitary, stunted white spruce may be seen. The upper slopes and the summits of “Sheep” and “Herringbone” mountains, the lower slopes of mountains to the west of the valley, the gravelly river flats and river meadows, and the bogs, small lakes and ponds, provide an abundance of rich and varied plant habitats. In 1964 P.M. Youngman, zoologist with the National Museum of Canada, assisted by Gaston Tessier, made incidental collections of plants — chiefly those that serve as food for small mammals — in the vicinity of Miles 54—55 of the Dempster Highway. Among the plant material that they brought me, | discovered fragments of Draba sibirica (Porsild 1964a), which is regularly gathered by the singing vole (Microtus miurus). At that time, Draba sibirica was ‘‘new’”’ to the flora of North America, its nearest Known station being 1,600 miles to the west, on the Kolyma River of eastern Siberia. Subsequent and more ample collections by Messrs. Youngman and Tessier, and later by my brother, Mr. R.T. Porsild of Whitehorse, Yukon Territory, proved D. sibirica to be very common locally in the rich alpine and subalpine herbmats along the Dempster Highway between Miles 50 and 58. In preparation for his new Flora of Alaska and Neighboring Ter- ritories, Eric Hultén (1966:315) reported a brief visit to the Ogilvie Mountains of central Yukon Territory at approximately Miles 50-54 on the Dempster Highway. In 1967 he discussed at some length “changes in nomenclature or conception in the forthcoming ‘Flora’ as compared to the earlier ‘Flora’ [Flora of Alaska and Yukon, 1941-— 50], and also additional taxa discovered in the time between publica- tion of the two works” (Hultén 1967:1). However, he did not attempt to document all the new stations or changes that were to appear in the new or revised distribution maps of his new Flora. In regard to * For a history of botanical exploration of the Yukon Territory prior to 1944, see Porsild 1951a: 21-26. t Foraglacial history of central Yukon Territory, see Bostock 1966. xi xii Preface the Yukon Territory, some of the ‘‘new’”’ dots are presumably based on Hultén’s own, but otherwise unreported, collections in the Ogilvie Mountains. However, a great many range extensions, thus far not documented, are reported in the present paper by citation of speci- mens. The majority of these specimens were collected by Mr. R.T. Porsild, and are now deposited in the National Herbarium of the National Museum of Natural Sciences, Ottawa. In 1966, 1967, and 1968, R.T. Porsild, while under contract with the National Museums of Canada, collected vascular plants in central Yukon. In 1966 and 1968 he collected plants mainly in the parts of the Ogilvie Mountains then accessible from the Dempster Highway, of which the first 89 miles had been completed. Most of the summer of 1967 was spent in the Mayo district, where special attention was paid to the still largely unexplored alpine floras of Keno Hill and Mount Haldane (both somewhat isolated mountains, or mountain massifs, slightly over 6,000 feet high) and to the remarkably rich and also unexplored floras of adjacent ponds, lakes, and bogs. The collections made during those three summers total 1,871 numbers, most of them with abundant and excellent duplicate material. Of this total, 1,057 are from the Ogilvie Mountains adjacent to Miles 44-89 on the Dempster Highway, mainly from elevations of 3,500 feet at road level, and from 6,000 to 6,500 feet on mountains east and west of the highway. In addition to regular collections for the National Herbarium of Canada, Mr. R.T. Porsild brought back a number of photographs and numerous samples of seeds from which chromosome counts were obtained for 58 taxa, of which 21 were previously unknown (Mulligan and Porsild 1969, 1970). During the three summers, Mr. Porsild was accompanied by his wife, who assisted him in collecting and preparing the specimens, and undertook the responsibility of drying and caring for them. From 3 to 12 July 1970, | accompanied my brother to sites along the Dempster Highway and in the Mayo district, where he had made major collections in 1966, 1967, and 1968. Following these visits, a few days were spent examining the rather different flora of the lime- stone formation, which, beyond Mile 89, overlays the crystalline and volcanic rocks exposed along the southern and central parts of the Ogilvie Mountains. Collections made then, chiefly near Miles 110- 112, added a number of species not observed elsewhere along the Dempster Highway (Porsild 1972). The 428 taxa listed and discussed in the catalogue following in- clude some that are believed new to science, some that are new to the flora of Canada or to the Yukon Territory, as well as a large number that are rare or have not been reported previously from the central Yukon Plateau. The bulk of the information reported herein has been obtained from the rich collections of R.T. Porsild, made in the years 1966, 1967, and 1968. In addition, some important range extensions are reported, obtained from a few other collections submitted to me for determination, and now deposited in the National Herbarium of Canada. The more important of these, mainly from northern or central Yukon, were contributed by Dr. A.M. Pearson of the Cana- dian Wildlife Service, Mr. P.M. Youngman of the National Museums of Canada, and Mr. Gaston Tessier of the Canadian Wildlife Service, formerly of the National Museums of Canada. Preface Abbreviations following a collector’s field number or a herbarium accession number indicate the herbarium in which the specimen is deposited. CAN National Herbarium of Canada, Ottawa DAO Canada Department of Agriculture, Ottawa LCU Catholic University of America, Washington, D.C. NY New York Botanical Garden, New York Ss Botanical Museum, Stockholm, Sweden US U.S. National Herbarium, Washington, D.C. Where no collector’s name is given, the specimens cited are from the collections of Mr. R.T. Porsild of Whitehorse, Yukon Territory. Unless otherwise stated, all specimens cited in this paper are in the National Herbarium, National Museum of Natural Sciences, Ottawa (CAN). The following 11 taxa are believed to be new or heretofore un- recognized: Calamagrostis Robertii n. sp., Trisetum spicatum f. viviparum n. forma, Melandrium apetalum ssp. ogilviense n. ssp., Pulsatilla ludoviciana f. glabrata n. forma, Papaver alaskanum f. stenopetalum n. forma, Draba yukonensis n. sp., Lomatogonium rotatum ssp. tenuifolium f. albiflorum n. forma, Pedicularis lanata ssp. yukonensis n. ssp., Antennaria Ellyae n. sp., Erigeron grandi- florus ssp. arcticus n. ssp., and Taraxacum mackenziense n. sp. The following 6 taxa are believed to be new to the flora of Canada: Polygonum O’Neillii, Claytonia arctica, Papaver Walpolei, Taraxa- cum carneocoloratum, T. maurolepium, and T. mitratum. Draba barbata is new to the flora of North America. And, finally, the following 38 taxa are new to the flora of Yukon Territory: Potamogeton foliosus var. macellus, Scheuchzeria palus- tris, Calamagrostis purpurascens ssp. Maltei, Poa Porsildii, Carex atherostachya, C. chordorrhiza, C. Heleonastes ssp. Heleonastes, C. holostoma, C. lasiocarpa, C. laxa, C. misandra, C. norvegica, Juncus stygius ssp. americanus, Luzula groenlandica, Tofieldia glutinosa, Spergularia rubra, Stellaria Alsine, Caltha natans, Ranun- culus natans var. intertextus, R. septentrionalis, Braya glabella, Car- damine microphylla, Draba exalata, D. fladnizensis, D. Macounii, Smelowskia calycina var. media, S. calycina var. Porsildii, Medicago falcata, Oxytropis Jordalii, Epilobium arcticum, Myriophyllum verti- cillatum, Cicuta bulbifera, Phlox alaskensis, Antennaria densifolia, A. neoalaskana, Arnica diversifolia, Erigeron linearis and Aster pansus. xiii auretes fei paoniiay rraigo a tiaiasethih D i . : ' Ps pee p Aipwepiorect of Meng HAAS . Patt Wer Ole See oe : he Eee ee r ¥ . , o 4 ve ro E v a ry 1 in ies 7 , a math, a yy a Ae Oe we A ' i ora ¢ "yp i if d Pe POA Sb 7 shy ey, a : Me fds, fi ile Pigtine | MED AEP athe sh yl ays vies bie wf ee ay ON . ' i ae . 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Cea 8 1 4 ae oe wr er. \ rary Oras ye JDOR® MADGHBA cs Vienne, (om 'y rom 2 AA "4 4 - 6 + } Sah Fo « * j anti a! Tui is, wee Comite bv Lr, AML Pearce o1-48 Lae 4 ; 2. - » & Y a : me ais, , ns _ \ us ee Mon Whidite Ge:rvios: Mr. Fo M. Yaoutigiran of She Kehoe Mubeunms of Verada; and Mr. Ganian T oaaeol oO! the Garvecian Viigo Sato yinerty of the National Museunre of Canes . Annotated List of Selected Species POLYPODIACEAE Cryptogramma Stelleri (S.G. Gmel.) Prantl Ogilvie Mountains: In rock crevices by a lake west of Dempster Highway, Mile 58, Nos. 325 and 358; on moist shaded rock ledges, Miles 54-55, No. 1839; on moist shaded ledges west of Dempster Highway, Mile 100, No. 1711. A rare species in Yukon, where it has been collected twice before, near Dawson (Porsild 1951a). Cystopteris montana (Lam.) Bernh. Ogilvie Mountains: Dempster Highway, Mile 100, on moist rocky ledges under Alnus cris- pa, No. 1708; the present collection extends the Known northern range in Yukon by ap- proximately one hundred miles (Porsild 1966: map 1). Dryopteris Robertiana (Hoffm.) C.Chr. Stewart Plateau: Mayo, on shady rock ledges of canyon below water reservoir, elevation 1,900 ft, No. 1011; in rocky scree above high- way between Mayo and Elsa Mine, approxi- mately 63°45/N, 135°45/W, A.E. & R.T. Porsild, No. 23,562. Ogilvie Mountains: Dempster Highway, Mile 100, on rocky slope under Alnus crispa, Nos. 1706 and 1710. In Yukon otherwise known from a few early collections near Dawson. Woodsia alpina (Bolton) S.F. Gray Ogilvie Mountains: Cliffs in canyon east of Dempster Highway, Miles 50-54, No. 97; on dry, rocky ledges of mountain east of Mile 51, No. 1578 (Porsild 1951a). OPHIOGLOSSACEAE Botrychium Lunaria (L.) Sw. var. minganense (Vict.) Dole Stewart Plateau: Vicinity of Mayo on river ter- race with Potentilla anserina, No. 698 (Porsild 1966:11). EQUISETACEAE Equisetum hiemale L. var. californicum Milde Stewart Plateau: Vicinity of Mayo, along road to Moose Creek, No. 991. In Yukon otherwise known from Mount Gray near Whitehorse, No. 1326, and from Dawson (Porsild 1951a). Equisetum palustre L. Common in rich meadows along Dempster Highway, Miles 96-98, No. 1725; in damp, cal- careous tundra south of Dempster Highway, near Mile 110, approximately 66°40’N, 128° 00’/W, A.E. & R.T. Porsild, No. 23,588. The last station extends the known Yukon range of this species by about 100 miles. Equisetum pratense Ehrh. Old Crow River near mouth of Johnson Creek, elevation 800 ft, Youngman & Tessier No. 635; Mayo 63°35/N, 135°54’W. Common in open woods near public camp grounds, A.E. & R.T. Porsild, No. 23,580. Equisetum scirpoides Michx. Ogilvie Mountains: Dempster Highway, Mile 110, in damp tundra at foot of limestone hills, 66°40’/N, A.E. & R.T. Porsild, No. 23,589. The collection-closes a gap between the Dawson area and the Old Crow Flats. = Equisetum sylvaticum L. In southeastern Yukon | noted (Porsild 1951 a) that on Precambrian rocks along the Canol Road all E. sylvaticum observed had smooth branches and thus belonged to var. paucira- mosum Milde. In the present collection E. sylvaticum is represented by 4 numbers, all from the general vicinity of Mayo. In Nos. 510 and 910 from Minto Road, the internodes of the main stems and the branches are strongly scabrous and thus represent typical E. sylva- ticum, whereas in Nos. 499 and 818 from lake- shores and swampy flats north of Mayo and from Hunker Creek near Dawson, No. 1400, the internodes and branches are smooth and, accordingly, should be referred to var. pauci- ramosum. Equisetum variegatum Schleich. Ogilvie Mountains: Dempster Highway, Mile 110, in damp tundra at foot of limestone hills, 66°40/N, A.E. & R.T. Porsild, No. 23,590. The collection closes a gap in the known distribu- tion of the species between the Dawson area and the Arctic Coast. LYCOPODIACEAE Lycopodium clavatum L. var. monostachyon Grev. & Hook. Keno Summit, 63°58’N, 135°42’W, in open willow thickets, A.E. &R.T. Porsild, No. 23,881. Annotated List of Selected Species Lycopodium Selago L. Ogilvie Mountains: Dempster Highway, Mile 110, in damp tundra at foot of limestone hill, A.E. & R.T. Porsild, No. 23,591. SELAGINELLACEAE Selaginella selaginoides (L.) Link Ogilvie Mountains: Dempster Highway, Mile 83 by calcareous seepage, No. 1614. The present collection is the first in central north- western Yukon (Porsild 1966: map 4). Selaginella sibirica (Milde) Hieron. Stewart Plateau: Mount Haldane, among rock slabs in alpine tundra, elevation 4,300-—5,500 ft, No. 668. Ogilvie Mountains: Mountains east of Dempster Highway, Miles 50-54, elevation 5,000 ft, No. 41; dry lichen tundra on river ter- races along Miles 55-56, No. 151; gravelly river terraces and rockslides by Miles 84—86, Nos. 1565, 1637, and 1687 (Porsild 1966: map 5). PINACEAE Abies lasiocarpa (Hook.) Nutt. Stewart Plateau: South branch of McQuesten River north of Mount Haldane, elevation 2,000 ft, No. 475. On Keno Hill, 63°57’N, 135/13°W, the upper altitudinal limit of Abies lasiocarpa is well over 5,000 ft, where several trees with well-formed cones were observed. The sta- tions cited represent the Known northern limit for this cordilleran species (Porsild 1951a: 74-75). Picea glauca (Moench) Voss var. Porsildii Raup Stewart Plateau: Vicinity of Mayo. This is probably the common race of white spruce in Yukon, dominant in rich soil in lowland situa- tions. In the valley of the Mackenzie River and in central Alaska, the var. Porsildii is also the dominant spruce on rich alluvial soils. Picea mariana (Mill.) B.S.P. On Keno Summit, 63°58’N, 135°42’W, eleva- tion 6,000 ft, black spruce and alpine fir seed- lings were noted and collected well above the timberline and well above the upper limit of Picea glauca, A.E. & R.T. Porsild, No. 23,582. Pinus contorta Loud. var. latifolia Engelm. Stewart Plateau: Dawson Highway, near Mile 188, between Carmacks and Pelly Crossing, No. 1391; Minto Road, 13 miles North of Mayo, Nos. 543, 900, and 1253, all with well- formed cones. As far as | know, this is the most northerly stand of pines in Yukon. TYPHACEAE Typha latifolia L. Fruiting specimens from shallow ponds north- east of Mayo, about 63°55’/N, 135°28’W, ele- vation 1,600 ft, Nos. 452 and 1252. The col- lections bridge a major gap between Norman Wells on the Mackenzie River and the Alaska Highway in central eastern Alaska. SPARGANIACEAE Sparganium angustifolium Michx. Stewart Plateau: Vicinity of Mayo, Nos. 890, 1003, 1013, 1249, and A.E. & R.T. Porsild, No. 23,557. Ogilvie Mountains: Dempster High- way, Mile 72, No. 1763; Miles 90-93, No. 1590. In Yukon otherwise known from a few collec- tions along the Canol Road (Porsild 1951a). Sparganium hyperboreum Laest. Stewart Plateau: Vicinity of Mayo, Nos. 513 and 802; Halfway Lakes north of Mayo, No. 951. Ogilvie Mountains: River flats along Dempster Highway, Mile 55, in shallow ponds, No. 424; Bell River near Lapierre House on the west slope of Richardson Mountains, Youngman & Tessier, No. 603. The last is a new northern limit for Yukon (Porsild 1966: map 6). Sparganium multipedunculatum (Morong) Rydb. Southeastern Yukon: 10-12 miles south from Ross River on Watson Lake Road, 6-7 Sept. 1964. Ogilvie Mountains: Dempster Highway, Miles 90-91, in deep ponds with Potamogeton species and Hippuris vulgaris, No. 1750. ZOSTERACEAE Potamogeton foliosus Raf. var. macellus Fern. Stewart Plateau: Five Mile Lake north of Mayo, No. 788, with young, well-formed fruits on 12 July 1967; Halfway Lakes north of Mayo, fruiting specimens on 19 July 1967, No. 886; Gramineae south branch of McQuesten River, elevation 2,000 ft, in shallow pond, with young fruits on 20 Aug. 1967, No. 1159. Ogilvie Mountains: South end of Chapman Lake, No. 1764; lake along Dempster Highway, Mile 74, No. 1816; in deep floating bog with P. a/pinus ssp. tenui- folius, near Miles 90-91, No. 1748, the last with fully matured fruits on 2 Aug. 1968. New to the flora of Yukon; in northwestern North America otherwise known only from a few collections in central Alaska and in the upper Mackenzie Basin (Great Bear River and Great Slave Lake). Potamogeton Friesii Rupr. Stewart Plateau: In a small lake between the south branch of McQuesten River and Mount Haldane, No. 1148, fruiting specimens on 10 Aug. 1967. Ogilvie Mountains: Lake west of Dempster Highway, Mile 65, elevation 3,750 ft, Nos. 317 and 481; Mile 72, south end of Chapman Lake, No. 1761. In Yukon previous- ly Known from near Carcross and from Wat- son Lake (Porsild 1966: map 7). Potamogeton perfoliatus L. var. gracilis Fries Ogilvie Mountains: Lakes west of Dempster Highway Miles 58 ana 65, elevation 3,800 ft, Nos. 483 and 497, flowering on 16-17 Aug. 1966. In Yukon otherwise known from a few collections on the Ross River, on the Canol Road, and from Carcross. Potamogeton praelongus Wulf. South end of Chapman Lake, Dempster High- way, about Mile 72, in water over 7 ft deep, where it is not common, No. 1767. In Yukon otherwise known from a single collection from Kluane Lake near Burwash Landing, H.M. Raup, No. 12,390. Potamogeton pusillus L. Stewart Plateau: 4 miles north of Mayo in small lakes, with almost mature fruits on 13 July 1967, No. 814; fruiting specimens in shal- low, alkaline lake on 28 Aug. 1967, No. 1246. In Yukon previously known only from the Dawson region and from near Whitehorse (Porsild 1951 a). Potamogeton strictifolius A. Benn. var. rutiloides Fern. Stewart Plateau: In shallow ponds northeast of Mayo, No. 817, specimens with young fruits and winter buds already forming on 13 July 1967. In Yukon otherwise known from a single collection in the Pelly River valley (Porsild 1951a). Potamogeton vaginatus Turcz. Stewart Plateau: In alkaline lakes northeast of Mayo, Nos. 809 and 1247. Specimens with well-formed fruits on 17 July 1967. In Yukon otherwise known from stations along the Canol Road, where it was reported common (Porsild 1951 a). JUNCAGINACEAE Scheuchzeria palustris L. var. americana Fern. Stewart Plateau: Vicinity of Halfway Lakes north of Mayo, in wet sphagnum bog asso- ciated with Carex limosa, C. Heleonastes and Eriophorum russeolum; with fully matured fruits on 20 Aug. 1967, No. 1186, and on 1 Aug. 1970, No. 1973. In North America long Known only from sta- tions south of the 60th parallel, Scheuchzeria palustris has recently been reported from the upper Kuskokwim River in southwestern Alaska (Drury 1956) and from peat bogs southeast of Great Slave Lake in the Mac- kenzie District (Cody and Porsild 1968). The present collection is the first from Yukon, and bridges an 800-mile gap between its known stations in Alaska and the Mackenzie District. GRAMINEAE Agropyron boreale (Turcz.) Drobov ssp. boreale Ogilvie Mountains: Along Dempster Highway, Mile 57, in dry ravine, elevation 4,200 ft, No. 354; Miles 83-86, Blackstone River valley, dry slope at foot of canyon, No. 1621B. Agropyron boreale (Turcz.) Drobov ssp. hyperarcticum (Polunin) Melderis Ogilvie Mountains: Along Dempster Highway, sandy riverbank meadow, near Mile 67, Nos. 1792-93; Blackstone River valley, dry slope, near Miles 83-86, No. 1621A. Agropyron pauciflorum (Schwein.) Hitchc. ssp. novae-angliae (Scribn.) Melderis Bonnet Plume Lake, 64°20’N, 132°00’W, W.H. Butler & N. Olsen, No. 10, 11-27 July 1966. Our specimens are a good match for A. pauci- florum ssp. novae-angliae, except that the anthers are 3-4 mm long. Annotated List of Selected Species Agropyron sericeum Hitchc. A. macrourum (Turcz.) Drobov Gravelly river terraces near Mayo, 63°35’N, 135°55’W, No. 853; gravelly slopes near Half- way Lakes, 63°48’N, 135°47’W, 4 July 1970, A.E. & R.T. Porsild, No. 23,560. Ogilvie Moun- tains: On gravel bars along Dempster High- way, Mile 95, No. 1733. Agropyron spicatum (Pursh) Rydb. Stewart Plateau: Vicinity of Mayo, 63°35/N, 135°45’W, in canyon below water reservoir, on gravel slope at base of canyon wall, No. 1009. In Yukon otherwise known from Fort Selkirk and Dawson. Agropyron subsecundum (Link) Hitchc. Vicinity of Mayo, 63°35/N, 135°45’W, on grav- elly river terraces, No. 929. Agropyron violaceum (Hornem.) Lge. ssp. andinum (Scribn. & Sm.) Melderis Ogilvie Mountains: River flats along Dempster Highway, near Mile 57, No. 364. Agropyron violaceum (Hornem.) Lge. ssp. violaceum Ogilvie Mountains: Upper Blackstone River valley, Dempster Highway, Mile 8, 64°50’N, 138°12’-15’W, elevation 4,200 ft, on high banks facing river, No. 1113; same place but on dry slope at foot of canyon wall, No. 1621. Alopecurus aequalis Sobol. Vicinity of Mayo: 13 miles north of Mayo, in meadows near lake, elevation 2,000 ft, No. 906; 4 miles north of Mayo, 63°35’N, 135° 45’W, lakeshore meadows, No. 786. Ogilvie Mountains: Dempster Highway, Miles 54—56, on edge of partly summer-dry pond, Nos. 446 and 456; Miles 93-95, by wet margins of ponds, Nos. 1589 and 1737. Chromosome counts of 2n=14 were obtained from Nos. 456 and 906 (Mulligan and Porsild 1969). Alopecurus glaucus Less. Stewart Plateau: Keno Summit, in moist al- pine tundra between 5,500 and 6,000 ft eleva- tion, Nos. 975, 1054, and 1281; the last had matured seed on 22-23 Aug. 1967. Ogilvie Mountains: Dempster Highway, Miles 67-69, in dry river-flat meadows, Nos. 1594 and 1813 (Porsild 1951a, 1966). Apparently a rare plant in Alaska and Yukon that, unlike A. alpinus, appears to be restricted to alpine situations even when oc- curring close to the seacoast. A. alpinus, on the other hand, although not strictly speaking littoral, is very rarely found far from the sea- coast. Arctagrostis poaeoides Nash Ogilvie Mountains: Along Dempster Highway, Miles 54-55, in grassy places above flood- plain, 11 Aug. 1968. No. 1829. A rare and perhaps not too-well-understood member of a taxonomically difficult group; thus far Known only from about one dozen stations in central Yukon and Alaska. Calamagrostis kolymaensis Kom., Not. Syst. Herb. Hort. Bot. Petrop. 2:129 (1921). | reported this Siberian species (Porsild 1965a:81) from a number of stations in arctic Alaska, where it had been identified as C. neglecta var. borealis or as C. deschamp- sioides. Hultén (1967:11; 1968:107), on the authority of Tolmatchev (1964:64—-68), had noted that C. kolymaensis is a synonym for C. Holmii Lge. (Lange in Holm 1885: 16; Pl. 1, fig. 2; Pl. 2, figs. 9-16). | have not seen the type of C. Holmii, nor that of C. kolymaensis, but as noted (Porsild 1965a:81), the description of the latter by Rozhevits (in Komarov 1934:219) certainly fits the plant reported by me from Alaska. How- ever, it differs in several important points from Lange’s description of C. Holmii (Lange in Holm 1885), where it is stated: “foliis omni- bus angustissimis, complicatis v. involutus”, although the plant shown in Lange’s plate 1 actually has flat, rather than inrolled, leaves. However, a more important difference is shown in Lange’s plate 2, where figures 9-16 show details of the floral parts of C. Holmii. According to the explanation accompanying figures 9-13 (Lange in Holm 1885:68), the floret, glumes, lemma, and palea are said to be magnified six times, and, if so, the glumes of C. Holmii are 10 mm long and the floret only slightly shorter. In the North American plant reported by me as C. kolymaensis (Por- sild 1965a), and also in the specimens dis- tributed as C. Holmii Lge. ex. Herb. Inst. Bot. Acad. Sci. URSS, from the Taymyr Peninsula and the lower Lena River, the leaves are flat, and the glumes 3 to 4 mm long, or less than half of those of C. Holmii. Since my report (Porsild 1965a), | have ex- amined plants from the Canadian Arctic matching the description of C. kolymaensis. The plants were discovered in the following Gramineae locations: Prince Patrick Island, Mould Bay, 76°12’N and 119°25’W, S.D. MacDonald, No. 57: Melville Island, Sherard Bay, ca. 76°N and 108°W, M. Kuc, July 1971. Calamagrostis chordorrhiza Porsild is not synonymous with either C. Holmii or C. koly- maensis, as suggested by Hultén (1967, 1968), nor is it even closely related to them, differ- ing, for example, by its strongly rhizomatose habit, very scabrous leaves, and taller and more robust culms, whose upper third parts are distinctly scabrous, and by its longer ligules. Calamagrostis purpurascens R. Br. ssp. Maitei (Polunin) Porsild n. comb. C. purpurascens var. Maltei Polunin, Nat. Mus. Can. Bull. 92: 51 (1940). Yukon: Ogilvie Mountains, Dempster High- way in dry alpine herbmats near Miles 57-58, elevation about 5,000 ft, No. 400; on limestone ridges near Mile 81, elevation 4,200 ft, No. 1491. Alaska: Brooks Range, De Long Moun- tains, Feniak Lake, 68°17’N, 158°20’W, Hol- men and Martensson, No. 61-1032. Thus far, ssp. Maltei was known only from the type collection made in 1927 near Pond Inlet in northern Baffiii Island in the eastern Canadian Arctic (No. 118,833). The new collections from Alaska and the Yukon, although remote by a gap of 1,500 miles, match perfectly the Pond Inlet plant, and clearly indicate that ssp. Maltei is no trivial variation of C. purpurascens. Sub- species Maltei differs most strikingly from C. purpurascens by its horizontal and always dis- tinct rhizome and by its consistently shorter but broader and more open panicle, averag- ing 4-6 cm in length and about 2 cm in diam- eter, whereas in typical C. purpurascens the length-diameter ratio of the panicle is ap- proximately 8:1. In ssp. Maltei the glumes are glabrous, dark purplish and 5-7 mm long, compared with 4 mm long in C. purpurascens, and the strongly geniculate awn is well ex- serted and spreading; the anthers are 2.0- 2.2 mm long and fertile; the culms mostly solitary, 25-30 cm tall, smooth except just below the panicle; the leaves tend to be shorter than in C. purpurascens, mostly flat, 3-4 mm wide. Calamagrostis Robertii n. sp. Gramen perenne dense caespitosum; culmi in fasciculis erecti 30-40 cm alti, tenuibus, laeves, bi-nodosi; folia culmo maturo demidia breviora, lamina involutis attenuata; vaginae laeves, superior 8-10 cm longus; liguda usque ad 4 mm longa, truncata. Panicula laxiuscula, 3-7 cm longa et 1.0-2.0 cm lata, rami breves parce hispidae, inferioribus remotiusculis; spiculae 4.5-—5.0 mm longae, glumae subae- quales, purpureae vel fusco-purpureae, ad carinam parce scabrida; palea inferior pur- purea infra medium dorsum arista recta lema- ta haud superante; calli pilis copiosi, dimi- diam paleam sub-aequantes; antherae 1.5mm longae. C. Robertii differs strikingly from all mem- bers of the genus with which | have been able to compare it by its densely caespitose habit, smooth purplish culms and leaf-sheaths, the uppermost of extraordinary length, the tip of its blade reaching the base of the panicle. The type of C. Robertii came from the Ogilvie Mountains in northern Yukon from the gravelly river terraces near Mile 56 on Demp- ster Highway, 6 Aug. 1966, No. 441 (Type: CAN), Plate 1. Calamagrostis Robertii is named for Robert T. Porsild, of Whitehorse, Yukon, in recogni- tion of his very important contributions to our knowledge of the vascular flora of central Yukon. Deschampsia brevifolia R. Br. Stewart Plateau: Keno Summit, elevation 6,000 ft, gravelly places in damp alpine tun- dra, Nos. 520, 757, 963, 982, 1029, and 1232. Ogilvie Mountains: Tundra along Miles 81-86 on Dempster Highway, Nos. 309, 500, 1122, 1133, 1540, and 1655. Gjaerevoll’s conclusion (1958: 25) that D. caespitosa ssp. orientalis “is identical with D. brevifolia’”’ is puzzling. D. caespitosa ssp. orientalis is a littoral and strictly oceanic race with an open panicle, whereas D. brevifolia is arctic-alpine, and its panicle is always dense and contracted. Chromosome counts made from No. 309 gave 2n=26 (Mulligan and Porsild 1969), which agrees with counts reported from Greenland (Jorgensen, Sgrensen, and Westergaard 1958). When it was reported from MacMillan Pass in the Mackenzie Mountains (Porsild 1951a), this taxon was otherwise known only from the Canadian Arctic Archipelago and from north- ern East and West Greenland. Gjaerevoll (1958) reported it from the White Mountains and the Alaska Range of central Alaska. | have also seen it from the St. Elias Mountains, D. Murray, No. 1435, from the Brooks Range, Plate 1 iZ_ ry “4, Zi a”, Le Calamagrostis Robertii n. sp. (Type) Approximately 2 natural size Gramineae and from elsewhere along the Arctic Slope of northern Alaska (Porsild 1964a: map 27). In 1970 | noted it as very common in alpine tun- dra along the Dempster Highway between Miles 80 and 110. According to Tolmatchev (1964), D. brevifolia is also wide-ranging in arctic parts of the USSR, from Nova Zembla to the Lena River. In the Canadian Arctic D. brevifolia is a dwarf species rarely more than 15 cm tall; in the mountains of Yukon and Alaska it is more high-grown, often with culms 30-35 cm high, and always restricted to truly alpine situations. Elymus innovatus Beal Ogilvie Mountains: Calcareous, gravelly river flats along Dempster Highway, Miles 81-83, Nos. 1559 and 1606; Mile 110, A.E. & R.T. Porsild, Nos. 23,592-93. A strongly calcico- lous species, apparently common on the Peel Plateau. Festuca baffinensis Polunin Ogilvie Mountains: Mountain east of Miles 50-54 on Dempster Highway, on dry, rocky exposure, No. 505. The collection bridges a gap between southwestern Yukon and the Richardson Mountains east of the Mackenzie Delta (Porsild 1955: 84-86; 1964b: map 54). Festuca brachyphylla Schultes Ogilvie Mountains: Mountains east of Miles 50-57 on Dempster Highway, Nos. 378, 395, and 505A. The collection bridges a gap be- tween southwestern Yukon and the Arctic Coast. In the first two numbers the glumes are of very unequal length, and although most lemmas are glabrous, scattered short hairs are present on some. In these characters, as well as in general appearance, these two numbers match F. hyperborea Holmen, de- scribed from northeastern Greenland and in recent years also recognized from the north- ernmost islands of the eastern Canadian Arc- tic; however, in Nos. 378 and 395 the leaves are distinctly tapered toward the apex, not equally broad to the apex asin F. hyperborea. The anthers in Nos. 378 and 395 are less than 1 mm long, and therefore our plant cannot be referred to F. ovina ssp. alaskana Holmen, described from the eastern Endicott Moun- tains of northern Alaska and reported from the Richardson Mountains of northeastern Yukon and northwestern Mackenzie District (Porsild 1966: 13-14). Festuca rubra L. ssp. Richardsonii (Hook.) Hult. F. rubra var. arenaria auct. non E. Fries Ogilvie Mountains: Sandy river meadow along Dempster Highway, Mile 67, Nos. 1696 and 1795; Mile 82, No. 1139. Festuca rubra L. ssp. rubra Ogilvie Mountains: Sandy river meadows along Dempster Highway, Mile 67, Nos. 1794 and 1800; limestone hill west of Mile 82, ele- vation 5,200 ft, No. 1104; gravel flats along upper Blackstone River, Nos. 1116 and 1123 (Porsild 1966: map 11). Festuca saximontana Rydb. Stewart Plateau: Gravelly slopes north of Mayo, elevation 1,600 ft, No. 800; sandy banks of creek along road to Moose Creek, eleva- tion 3,000 ft, No. 987. A cordilleran species that reaches central interior Alaska through southern and central Yukon. Festuca vivipara (L.) Sm. Stewart Plateau: Keno Hill, open gravelly patch on herbmat slope, elevation 4,500 ft, No. 955. In Yukon otherwise known from two collections in the St. Elias Mountains. For a discussion of this and the three preceding taxa, see Porsild 1966: maps 11 and 12. Glyceria borealis (Nash) Batch. Stewart Plateau: 4 miles north of Mayo on gravelly edge of small lake, elevation 1,900 ft, No. 947. In Yukon: otherwise known only from single stations near Dawson and from Little Atlin Lake (Raup, No. 11,371). Glyceria grandis S. Wats. Ogilvie Mountains: In shallow slough west of Dempster Highway near Mile 95, No. 1743. This station extends, by about 100 miles, the known northern limit in the Yukon for the species. Glyceria pulchella (Nash) Schum. Stewart Plateau: In wet meadow bordering Five Mile Lake north of Mayo, No. 815 (Por- sild 1966: map 13). Hierochloe odorata (L.) Wahlenb. Ogilvie Mountains: River flats along Demp- ster Highway, Mile 57, in dry heath, No. 231; Blackstone River valley, near Miles 82-86, on gravelly river flats, Nos. 1127 and 1622. The Annotated List of Selected Species last two collections fill a gap in the known distribution of the species in central Yukon. Phippsia algida R. Br. Ogilvie Mountains: River flats along Demp- ster Highway, Mile 57, elevation 2,500 ft, on dry gravel, No. 131, with mature seeds on 9 July 1966. A chromosome count (unpub- lished) agrees with 2n=28 reported from Greenland (Bocher, Holmen, and Jakobsen 1968); Dempster Highway, Mile 58 by over- flow from cold spring, No. 495. In Yukon P. algida has previously been reported from mountains south of Kluane Lake (Porsild 1966). Poa annua L. Minto Lake Road, Miles 9 and 13, in damp gravel road, Nos. 911 and 1295. Common and spreading rapidly along new roads and new settlements. In Yukon perhaps mainly bien- nial. No. 1295 had mature seeds on 26 Aug. 1969, and provided a chromosome count of 2n= 28 (Mulligan and Porsild 1969). Poa lanata Scribn. & Merr. Mountain summit east of Dempster Highway, Miles 57-58, elevation about 6,000 ft, No. 379. An alpine and perhaps often overlooked species; the present collection is the first from central Yukon and represents a slight northward extension of its known Yukon range. Poa paucispicula Scribn. & Merr. Keno Summit: 63°58/N, 135°42’W, common in rather moist alpine heath and herbmats from 4,500 to 6,000 ft elevation, Nos. 759, 968, and 1028; “Highlet Gulch’’, tributary to Minto Creek, 63°45’N, 136°10’W, elevation 4,000 ft, Nos. 841 and 842. The collections are the first from central Yukon (Porsild 1951a: 91). Poa Porsildii Gjaerevoll, Kgl. Nor. Vidensk. Selsk. Forh. 29 (16): 73-76 (1956). P. vaseyochloa Hult., Ark. Bot., ser. 2, vol. 7 (1): 16 (1967) non Scribn. Ogilvie Mountains: Granitic ridge east of Mile 57 on Dempster Highway, elevation 6,000 ft, No. 375; in turfy alpine tundra near Mile 81, No. 1499. Stewart Plateau: Keno Summit, south slope, elevation just under 6,000 ft, in alpine heath near snowbed, Nos. 1023, 1031, and 1223; same place, large tussock of stami- nate plant, A.E. & R.T. Porsild, No. 23,566, and nearby pistillate plant, No. 23,567. The description of P. Porsildii was based on over-mature specimens collected in Mac- Millan Pass near the Yukon—Mackenzie bor- der, in September, when the first snow had already fallen (Type: Porsild & Breitung, No. 11,185)..With the better and more abundant material now available, not only from the Mackenzie Mountains (Cody and Porsild 1968), but also from the Ogilvie Mountains and from the Stewart Plateau where, more- over, | was able to examine live material in July 1970, some emendations can now be added to the original description. The most important of these, perhaps, is that Poa Por- sildii appears to be completely dioecious. The type cited above consists of two tufts, each containing 4 and 5 culms; both tufts were part of a large tussock. All florets are entirely staminate. This is also the case with No. 375 from Ogilvie Mountains, of which the collector prepared no less than nine duplicate sheets, all from one tussock. In the Yukon, in 1970, | examined hundreds of tussocks of P. Por- sildii; all proved either male or female and were fully functional; in none did | find bi- sexual florets. The culms are 25 to 30 cm tall in the type, but in the large material that | examined some culms measured 35 cm. In the description the leaves are said to be narrow; in life the blade may be 2 mm wide, but is often so tightly rolled as to appear narrower. The ligule, de- scribed as being ‘1.5-2.0 mm long, ovate- transversely cut and laciniate’ (Gjaerevoll 1956: 73-76) is rarely quite that long. The spreading, or even drooping, lower panicle branches noted and well illustrated by Gjaerevoll (1956: figs. 1 and 2) are excel- lent field characters that are readily observed even from a distance. Hultén’s categorical statement (1967: 16) that “Poa Porsildii proved [sic!] to be identical with this species [P. vaseyochloa Scribn.]”’ is difficult to understand. The latter, accord- ing to Hitchcock, Cronquist, and Ownbey (1969:667), is identical to P. Leibergii Scribn. described earlier; a still earlier synonym of P. vaseyochloa is P. pulchella Vasey, not P. pulchella Salisb. Vasey (1882:32) described his Poa pulchella as ‘“‘a handsome dwarf Poa, related to P. Jaxa but distinct...”, and A.S. Hitchcock (1951) wrote about P. vaseyochloa, ‘“.. small dense soft lax tufts; culms erect, Cyperaceae 10 to 20 cm tall... panicle ovate, 2 to 4 cm long, few-flowered, open, the slender branches spreading bearing 1 or 2 spike- lets...’. These comments apply to the plant illustrated in Hitchcock, Cronquist, and Own- bey (1969) and to four sheets of P. vaseyo- chloa from Klicktitat County, Washington, de- posited in the National Herbarium of Canada. Among these sheets are two from the original Suksdorf collections, but they are not at all the robust Yukon bluegrass, which is twice as tall, strictly dioecious, has drooping panicle branches bearing 2 to 5 spikelets, normally forms huge tussocks, and is clearly an alpine endemic of unglaciated central Mackenzie Mountains, Northwest Territories, Yukon, and Alaska. P. Leibergii Scribn. (P. vaseyochloa) is known only “from hot interior sagebrush plains and foothills of Oregon and Washing- ton” (Vasey 1882:32), and is separated from the Yukon plant by a gap of 1,400 miles. Poa Williamsii Nash Stewart Plateau: Keno Hill, in gravelly creek bed, No. 977. Ogilvie Mountains: River flats along Dempster Highway, Miles 55-56, on gravel ridges, elevation 2,500 ft, No. 273. The collections are the first from central Yukon. (Porsild 1951a, 1966). Puccinellia interior Sgr. Mayo District near Five Mile Lake, where it was common along gravelly lakeshore, A.E. & R.T. Porsild, No. 23,554. The present col- lection represents a new northern limit for the Yukon, where it was first reported from the vicinity of Mackintosh, south of Kluane Lake (Porsild 1966). Trisetum spicatum (L.) Richt. f. viviparum n. forma: spiculis viviparis Ogilvie Mountains: Gravelly river bank along Mile 82 on Dempster Highway, No. 1142 (Type: CAN). The specimen consists of a single tuft in which one panicle is viviparous while the remaining three appear normal, al- though they are probably sterile. Typical T. spicatum were noted as common on Keno Summit, No. 1282, and in the Ogilvie Moun- tains along Dempster Highway. A chromo- some count from seeds of No. 308 from river flats, Miles 57-58 on Dempster Highway, gave 2n = 28 (Mulligan and Porsild 1969). CYPERACEAE Carex aenea Fern. Vicinity of Minto Bridge, 63°41’N, 135°51’W, in wet meadows, No. 885. The collection bridges a gap between southeastern Yukon and the Dawson area. Carex athrostachya Olney Vicinity of Mayo, 63°37’N, 135°55’W, on shores of small bog pond, with well-formed heads on 12 May 1967, No. 799. New to the flora of Yukon, the nearest stations are near Fairbanks, Anchorage, and southeastern Alaska. Carex atratiformis Britt. sso. Raymondii (Calder) Porsild Vicinity of Mayo, 63°35’N, 135°45’W, bog on gravelly river terrace, No. 690. The collection bridges a gap between southeastern Yukon and the Dawson area (Porsild 1966: 17-18). Carex atrofusca Schk. Ogilvie Mountains: Dempster Highway, Mile 72 on wet shores of Chapman Lake, No. 1776; Mile 110 in moist, calcareous tundra, A.E. & R.T. Porsild, No. 23,599. The present collec- tion bridges a gap between mountains of southwestern Yukon and the Arctic Coast. Carex atrosquama Mack. Stewart Plateau: Mount Haldane, along creek in Open spruce woods, elevation 3,500 ft, No. 626. The collection bridges a gap between mountains of southeastern Yukon and those of central Alaska. (Porsild 1966: map 22). Carex bicolor All. Ogilvie Mountains: River flat along Dempster Highway, Mile 57, on turfy edge of dry bog, No. 311. In Yukon otherwise known from a few collections in the southeastern and south- western parts of the territory (Porsild 1951a, 1966). Carex bonanzensis Britt. Stewart Plateau: Vicinity of Mayo in meadows along Five Mile Lake, No. 816. Ogilvie Moun- tains: Dempster Highway in wet meadows near Mile 95, No. 1738. In view of its reported Asiatic range, from Kanin Peninsula and the Ural Mountains to Anadyr in eastern Siberia (Tolmatchev 1966), it is strange that in North America C. bonanzensis appears to be en- tirely absent in the Bering Sea region, and is Annotated List of Selected Species rare and localized from central Yukon and Alaska and from western Mackenzie District north to the Mackenzie Delta. Carex brunnescens (Pers.) Poir. ssp. alaskana Kalela Ogilvie Mountains: Dempster Highway, open gravelly river terraces, Miles 68-69, No. 1592; Mile 81, No. 1549. The present collection bridges a gap in the known range of ssp. alaskana between southeastern Yukon and central Alaska (Kalela 1965). Carex capillaris L. s. /at. Three easily distinguished “‘races” of C. ca- pillaris occur in central Yukon: A. Terminal spike pistillate or sometimes with a few staminate flowers at the base; basal leaves flat, always fresh-green, and the mature flowering stems usually 10-20 cm tall. C. capillaris ssp. robustior (Lge. ex Drej.) Bocher (C. Bocheriana Love et al., but not C. Krausei Boeckl.). Ogilvie Mountains: River flats along Dempster Highway, Mile 57, on clay spots in tundra, No. 226. AA. Terminal spike always staminate; basal leaves grey-green. B. Mature flowering stems rarely over 10 cm tall. Common in alpine tundra. Arctic-alpine. C. capillaris L. s. str. BB. Mature flowering stems 20-30 cm tall. Common in woodland meadows. Bo- real forest. C. capillaris L. var. elon- gata Olney (C. capillaris L. ssp. chloro- stachys (Stev.) Love et al.). Carex capitata L. Ogilvie Mountains: Peaty bogs along Demp- ster Highway, Miles 68-69, No. 1593; calcar- eous tundra along Dempster Highway, Mile 81, No. 1462. The second collection is slightly beyond the previously known northern limit for Yukon. A decided calciphile of western boreal-alpine rather than arctic range, in the southwestern Hudson Bay region barely over- lapping that of the eastern and amphi-Atlantic C. arctogena Sm. Carex chordorrhiza Ehrh. Ogilvie Mountains: River flats along Dempster Highway, Mile 56, in wet sohagnum moss by shallow pond, No. 419; swampy lakeshore, near Mile 75, No. 1781; upper Blackstone River near Mile 82 on Dempster Highway, in moist tundra, No. 1137. These collections, 10 which are the first from Yukon, bridge a major gap in the known range of C. chordorrhiza between the Mackenzie District and central Alaska. Carex concinna R. Br. Vicinity of Mayo, 63°35’N, 135°45’W, gravelly river terrace, No. 441; edge of shallow pond, No. 483. Ogilvie Mountains: Wooded calcar- eous Cliffs, Dempster Highway, Mile 83, ele- vation 3,800 ft, No. 1510. The last collection is a new northern limit for the species in Yukon. Carex consimilis Holm Mayo District: Keno Summit, 63°58/N, 135°42’ W, elevation 5,500-6,000 ft, alpine tundra, No. 1057; same place, in wet sphagnum bog, No. 1227. Ogilvie Mountains: Along Dempster Highway in alpine herbmats, Miles 51-52, No. 204; alpine tundra, elevation 4,500 ft, Miles 57-58, No. 260; peaty hummocks in open spruce woods, Mile 83, No. 1615; moist tundra flat at foot of limestone mountain, Mile 110, A.E. & R.T. Porsild, No. 23,601. For general range and alleged relation to C. Bigelowii, see discussion of C. /ugens. Carex eleusinoides Turcz. C. kokrinensis Porsild Keno Summit, 63°58’N, 135°42’W, elevation 5,500—6,000 ft, in wet places below seepage, A.E. & R.T. Porsild, No. 23,583. New to central Yukon. Carex glacialis Mack. Ogilvie Mountains: Limestone hills west of Miles 82-83, on Dempster Highway in dry, al- pine heath where it was associated with Tha- lictrum alpinum, Nos. 1103 and 1520, the last with culms 20 cm high; slopes of poorly vege- tated limestone hills west of Mile 110, asso- ciated with Kobresia myosuroides and Dryas sylvatica, A.E. & R.T. Porsild, No. 23,602. Apparently a rare species in Alaska and Yukon, where it has been collected only a few times, mostly in high-alpine situations and probably always in soil rich in lime (Porsild 1951a, 1966). Carex gynocrates Wormskj. Mayo District: Vicinity of Minto Bridge, 63°41’N, 135°51’W, Nos. 530-31. The local abundance of this species, for example in southwestern Yukon and along the Canol Road, suggests that itis, perhaps, more abun- dant and wide-ranging in Yukon than would Cyperaceae appear from the relatively few collections on hand. Carex Heleonastes Ehrh. ssp. Heleonastes Stewart Plateau: Vicinity of Halfway Lakes, elevation 2,300 ft, in a wet peat bog, No. 1188, where it was associated with Scheuchzeria palustris and Carex limosa. The collection is the first from Yukon and constitutes a major extension of the range of this taxon known for its “spotty” distribution, which may be ac- counted for to some extent by its preference for very boggy places difficult to approach. Its nearest known station is Liard Hot Springs in northern British Columbia (Porsild and Crum 1961), except for a specimen from Rink Rapid on the Yukon River near Carmacks, er- roneously reported as C. neurochlaena Holm (C. Heleonastes ssp. neurochlaena (Holm) Bocher). Carex holostoma Drej. Ogilvie Mountains: In wet tundra, approxi- mately 64°50’N near Mile 82 on Dempster Highway, with fully ripe fruits on 5 Aug. 1967, No. 1129. New to the flora of the Yukon, and in Alaska known only from three widely dis- junct stations, two of them on the shores of Bering Strait, the third on the northern slope of the Wrangell Mountains. Carex lasiocarpa Ehrh. Stewart Plateau: Lower northern and eastern slopes of Mount Haldane, elevation 2,000 ft, No. 1161, on wet edge of bog where it grew with Equisetum limosum. New to the flora of Yukon, the present station bridges a gap be- tween eastern central Alaska and the Mac- Kenzie River valley (Porsild 1951a: 116). Carex laxa Wahlenb. Vicinity of Mayo, 63°48’N, 135°47’W, on swampy flat near Halfway Lakes, where it was associated with Drosera anglica and Utricu- laria minor, No. 887; south branch of Mc- Questen River, in a wet bog with Drosera anglica and Tofieldia glutinosa, elevation 2,000 ft, Nos. 1154 and 1207. Since Hultén’s report (1942) of C. /axa as new to North America, based on collections from Richardson Highway in central Alaska (J.P. Anderson, No. 2712B), a second station was added from the Mackenzie Delta (Porsild 1951a). The three collections cited above are the first reports from Yukon, and they close a gap between central Alaska and northwest- ern Mackenzie District. Carex leptalea Wahlenb. Stewart Plateau: Vicinity of Halfway Lakes north of Mayo in a wet bog, No. 1189. Since first reported from Yukon on the Canol Road (Porsild 1951a), a few more stations have been added in the central and southern parts. Carex limosa L. Stewart Plateau: Vicinity of Halfway Lakes north of Mayo, in a sphagnum bog with C. Heleonastes, Nos. 480 and 502; vicinity of Minto Bridge, on floating bog, No. 535; north- eastern slope of Mount Haldane, in a bog with C. livida and Scirpus hudsonianus, No. 1162. Carex livida (Wahlenb.) Willd. Ogilvie Mountains: Dempster Highway, cal- careous river-flat bogs, Miles 53-55, Nos. 428, 429, and 1827; Mile 75, No. 1782. In Yukon otherwise known from a single collec- tion near Watson Lake. Carex lugens Holm, Amer. J. Sci. 10: 269, figs. A-D (1900). Ogilvie Mountains: River flats along Dempster Highway, Miles 57-58, common in not-too-dry meadows between gravel ridges, elevation 4,200 ft, No. 297; common in wet tundra along Dempster Highway, Mile 110, A.E. & R.T. Porsild, No. 23,603; Hungry Lake, 65°40’N, 136°00’W, elevation 800 ft, in sparsely wooded boggy places along stream, Youngman & Tes- sier, No. 691. In Yukon previously known only from the Dawson area. Although Carex /ugens and C. consimilis are of similar habit, they are, nevertheless, taxonomically as well as ecologically distinct; both are endemic in unglaciated Alaska, cen- tral Yukon, and northwestern Mackenzie Dis- trict. C. Jugens commonly grows in damp, mossy places by small streams, whereas C. consimilis prefers deep, peaty soil in spruce bogs (muskegs). Both are tussock-forming, and in both the primary roots are character- istically velvety because of a dense cover of pale, shaggy “hairs” (? mycorrhiza) that are approximately 1 mm long. In C. /ugens the slender culms are from 50-100 cm tall or taller, and the long, flat leaves are rarely over 2 mm wide; the more or less sessile, nearly contiguous pistillate spikes, 3 (2) in number, are commonly 1-2 cm long, but rarely over 3 mm in diameter. In C. consimilis the culms 11 Annotated List of Selected Species are coarser, but rarely over 25 cm tall, the leaves 3-4 mm wide, and the 3 (rarely 4) pis- tillate spikes commonly 1.0 or rarely 1.5 cm long and 5 mm wide. Carex consimilis was described as being stoloniferous, but the type from Klondike Indian Divide, Yukon Territory, J. Macoun, No. 53,878 (CAN 25,199), is caespitose and entirely lacking stolons. Mackenzie (1935:401) listed C. yukonensis Britt., C. consimilis Holm, and C. cyclocarpa Holm (the types of all three from the Dawson area of central Yukon) as synonyms of C. lugens, of which the type came from Kasilof on the north shore of Kenai Peninsula, Alas- ka. Holm’s figure (1900) agrees perfectly with a duplicate of the type, W.H. Evans, No. 725 (CAN 25,178), and shows two erect, well- separated pistillate spikes, the lower distinct- ly pedicellate, though in the text Holm stated, “spikes 3-4, contiguous’”’. It is of particular interest that Holm’s illus- tration of C. Jugens shows the hairy roots, a feature that, as far as | am aware, has not been described by anyone else. The hairs are ‘best developed in plants growing in peaty soil, and are therefore always present in C. consimilis, but are absent or weakly devel- oped in C. lugens when growing in wet, mar- shy places or in mineral soil. Hultén (1942: 334; 1968: 248) placed C. con- similis in synonymy under C. Bigelowii Torr., perhaps influenced by Holm’s statement (1900) that the plant is ‘“‘stoloniferous’” and “the perigynium ... sharply denticulate along the upper margins’. Neither of these state- ments actually applies to the type. But even so, C. consimilis is certainly quite distinct from C. Bigelowii, which has cord-like and scaly rhizomes terminating in leafy shoots and is not a muskeg plant at all. Moreover, C. Bigelowii is an amphi-Atlantic rather than a circumpolar species, not known in North America west of longitude 125°W, nor, ac- cording to Tolmatchev (1966), in Eurasia east of Murmansk. Carex membranacea Hook. Ogilvie Mountains: Dempster Highway, Mile 110, common in moist tundra below limestone hills, A.E. & R.T. Porsild, No. 23,604. The col- lection closes a gap between southwestern Yukon and the Arctic Coast. 12 Carex microglochin Wahlenb. Ogilvie Mountains: Wet places along Demp- ster Highway, Miles 53-54, Nos. 426 and 1826. In the Yukon otherwise known from the south- eastern and southwestern parts. Carex microptera Mack. Stewart Plateau: In open poplar-willow forest along road to Moose Creek, No. 986. Ogilvie Mountains: Along Mile 67 on Dempster High- way on gravelly river flat, No. 1791. When first reported from Yukon (Porsild 1966), the habitat, which was near a clearing made for a pipeline near Pine Creek in south- eastern Yukon, suggested a recent introduc- tion. The environment of these additional Yukon stations leaves no question that C. microptera is native to Yukon. Carex misandra R. Br. Ogilvie Mountains: River-flat tundra along Dempster Highway, Mile 52, Youngman & Tessier, No. 376; along Miles 57-58 on ridge along creek, Nos. 240, 295, and 374; in tundra bogs near Mile 81, No. 1476; Mile 110, A.E. & R.T. Porsild, No. 23,605; also known from the St. Elias Mountains near Steele Glacier, D.F. & B.M. Murray, No. 1424. C. misandra is an arctic species, new to the Yukon but probably also occurring along the Arctic Coast and on the west slopes of the Richardson and Mackenzie mountains. Carex nardina E. Fries var. atriceps Kuk. Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 50-54 and 57-58, Nos. 386 and 509; on limestone hills west of Mile 82, No. 1081. Appears to be rare in Yukon, where it has been reported only a few times. Carex norvegica Retz. Ogilvie Mountains: River flats along Dempster Highway, near Mile 58, elevation 2,800 ft, in damp treeless tundra, approximately 65°N, 138°W, No. 294. Densely tufted culms 3-4 dm tall, slender but stiff, about twice as long as the flat or slightly involute leaves, 2-3 mm wide, and basal bladeless sheaths, dark purple and shiny. Spikelets 3, approximate, the scales black with a faint median nerve, slightly nar- rower and shorter than the distinctly dark- brown tight-fitting perigynia, 1.8-2.2 mm long, distinctly broadest above the middle and abruptly tapering to a distinct beak. Cyperaceae The discovery of this low-arctic ‘‘amphi- Atlantic” taxon in unglaciated northern cen- tral Yukon, 1,200 miles west of its nearest North American station, in Keewatin near the 100th meridian, is significant in view of the report of C. norvegica ssp. conicorostrata Kalela (1944) from the Chukotsk Peninsula in easternmost Siberia. Typical C. media R. Br., with pale-green elliptic perigynia, 3.0—3.3 mm long, were col- lected in open willow corridors on nearby river flats, Nos. 296 and 422. , Carex paupercula Michx. Stewart Plateau: Halfway Lakes in sphagnum bogs, No. 882. In Yukon otherwise known from the vicinity of Dawson and from a few stations along the Canol Road (Porsild 1951a). Carex petricosa Dew. Ogilvie Mountains: In dry, alpine heath on limestone mountain, elevation 5,200 ft, west of Dempster Highway, Miles 82-83, Nos. 1100, 1477, 1512, and 1527; common locally on cal- careous slopes and floodplains near Mile 110, A.E. & R.T. Porsild, Nos. 23,606-07. The known range of this still rare and little- collected species, until recently recorded only from mountains of Alberta near the type locality at approximately 53°N, has lately been extended north through and beyond the Mackenzie District to Victoria Island in the western Canadian Arctic Archipelago, south- eastern and central Yukon (Porsild 1951a, 1955, 1966), and Alaska. Its known southern limit is in Banff National Park, Alberta, at ap- proximately 51°N. Carex rariflora (Wahlenb.) Sm. Ogilvie Mountains: River flats along Dempster Highway, Miles 55-56, No. 274; in swampy tundra along upper Blackstone River from Mile 72 to 82 on Dempster Highway, Nos. 1136, 1770, and 1773. An arctic species ap- parently rare in Yukon, where it is probably restricted to alpine situations. Carex rostrata Stokes Vicinity of Mayo: Marshes along Five Mile Lake, No. 819. Ogilvie Mountains: Swampy river flats along Dempster Highway, Miles 57 and 95, Nos. 312 and 1744. No. 1744 is a very stout specimen, over 1 m tall, in which the 3 terminal spikes are staminate and linear, 6 cm long, and the lower 3 androgynous, the lowermost 9 cm long, with lateral branches or fascicles in the lower third. Carex tenuiflora Wahlenb. Ogilvie Mountains: River flats and bogs along Dempster Highway, Mile 56, Nos. 418 and 420; Mile 84, in damp meadows, No. 1695. Stewart Plateau: In wet swamp near Halfway Lakes, No. 1190. In Yukon thus far collected only a few times. Carex vaginata Tausch Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 54—57, in sedge and wil- low meadow, Nos. 250, 397, and 1824; Mile 110, floodplain, A.E. & R.T. Porsild, No. 23,608. Stewart Plateau: Vicinity of Minto Bridge in wet places in open spruce woods, No. 878. Carex viridula Michx. Carmacks Road, Mile 71 near Twin Lakes, Nos. 1864 and 1908. In Yukon otherwise Known only from Whitehorse and Watson Lake. Carex Williamsii Britt. Ogilvie Mountains: River flats along Dempster Highway, Mile 53, No. 413A. In Yukon other- wise known only from the type locality, which is near Dawson, and from one station in the Kluane Lake area. Eleocharis acicularis (L.) Roem. & Schultes Not uncommon in soft, muddy places along quiet lake and pond shores. Halfway Lakes north of Mayo, 63°48/N, 135°47’W, No. 1187; Mayo Lake Road 13 miles north of Mayo, No. 1181; in pond on lower slope of Mount Haldane, 63°50/N, 135°50’W, No. 1156; lake west of Dempster Highway, Mile 58, No. 330; shallow pond, Dempster Highway, Mile 84, No. 1692. Eleocharis palustris (L.) Roem. & Schultes Shallow pond shores north of Mayo, 63°37’N, 135°55’W, Nos. 528 and 778; Dempster High- way, boggy places at Miles 55-56, No. 275, and Miles 96-98, No. 1716. The last collection is a northward extension of the known Yukon range. Eleocharis pauciflora (Lightf.) Link var. Suksdorfiana (Beauv.) Svens. Twin Lakes, approximately 61°40’N, 136°00’ W, on moist, sandy lakeshore, No. 1905; McLean Lake near Whitehorse on marly lake- 13 Annotated List of Selected Species shore, No. 1955. (Porsild 1951a: 102; 1966: - map. 31). Eriophorum angustifolium Honck. Common in swampy places on river flats along Dempster Highway, Miles 57 and 84-88, Nos. 293 and 1690. The last collection is about 100 miles beyond the previously known north- ern limit for this species in Yukon. Eriophorum callitrix Cham. Mayo District, Keno Summit, 63°58’N, 135°42’W, No. 760. Ogilvie Mountains: Along Dempster Highway, Mile 81, elevation 4,200 ft, calcareous spots in alpine tundra, No. 1496; Mile 110, in moist calcareous spots in tundra, A.E. & R.T. Porsild, No. 23,595. In Yukon pre- viously reported only from near Pine Creek in southwestern Yukon (Porsild 1966). Eriophorum russeolum E. Fries var. albidum Nyl. Ogilvie Mountains: River flats and swampy lakeshore along Dempster Highway, Miles 50-51, No. 501; Mile 57, No. 314; Mile 72, No. 1772; Mile 82, No. 1128. In Yukon otherwise known from the southeastern part along the Canol Road (Porsild 1951a). Eriophorum Scheuchzeri Hoppe Ogilvie Mountains: River flats along Dempster Highway, Mile 58, No. 304. Probably common in central and northern Yukon, although thus far not actually collected north of the Ogilvie Mountains. Eriophorum triste (T. Fries) Hadac & Love Mayo District: Keno Summit, common in moist mineral spots of alpine tundra, Nos. 720, 761, and 1055. Ogilvie Mountains: Moun- tain east of Dempster Highway, Miles 57-58, elevation 5,800 ft, No. 269. Eriophorum vaginatum L. ssp. vaginatum Ogilvie Mountains: Peat bog along Dempster Highway, Mile 82, 64°50’N, No. 1135; Peel River basin, Hungry Lake, 65°50’N, 136°00’W, Youngman & Tessier, No. 692. Kobresia hyperborea Porsild Ogilvie Mountains: In turfy alpine tundra along Dempster Highway, Mile 82, No. 362. In Yukon otherwise known only from a few col- lections in St. Elias Mountains and on the Arctic Coast. 14 Kobresia simpliciuscula Mack. Ogilvie Mountains: Calcareous river flats along Dempster Highway, Miles 72-75, Nos. 433, 1774-75, 1785, and 1825; Dempster High- way, Mile 110, A.E. & R.T. Porsild, Nos. 23,595-98. The series partly closes a gap in the Yukon range of this species, between southwestern and southeastern Yukon and the Arctic Coast (Porsild 1951a). Scirpus caespitosus L. ssp. austriacus (Pall.) Aschers. & Graebn. Common in peat bogs everywhere; one col- lection, Dempster Highway, Mile 85, No. 1553, extends the known Yukon range by approxi- mately 60 miles. Scirpus hudsonianus (Michx.) Fern. Stewart Plateau: Vicinity of Mayo, Minto Bridge, No. 542; Halfway Lakes, No. 861; north slope of Mount Haldane in wet peat bog with Carex livida var. Grayana, C. limosa, and Dro- sera anglica, No. 1163. Scirpus Rollandii Fern. Boggy shores of marly pond near Twin Lakes, approximately 61°40’N, 136°00’W, No. 1878. This is the second collection from Yukon, and it was found about 190 miles northeast of the earlier station near Mackintosh in southwest- ern Yukon (Porsild 1966: 22, map 32). Scirpus validus M. Vahl Dempster Highway, Miles 91-93, in wet places by pond margins, Nos. 1587 and 1752, speci- mens with fully ripened achenes on 16 July and 2 Aug. 1968. Vicinity of Mayo, Nos. 672 and 1243, the last with mature achenes on 28 Aug. 1967. LEMNACEAE Lemna minor L. Stewart Plateau: In ponds northeast of Mayo, Nos. 454 and 1199. In Yukon thus far collected a few times in the Dawson area and in the southwest (Porsild 1966: map 33). Lemna trisulca L. Vicinity of Mayo in a shallow lake, No. 1200. Ogilvie Mountains: In ponds west of Dempster Highway, Mile 65, Nos. 321 and 485; Mile 82, south end of Chapman Lake, No. 1758. The last station bridges a gap between central and northern Yukon and the Mackenzie Delta. Juncaceae JUNCACEAE Juncus balticus Willd. var. alaskanus (Hult.) Porsild Ogilvie Mountains: Very common in wet places on river flats along Dempster Highway, Miles 57, 81-82, Nos. 291, 1131, and 1537. The last station is slightly beyond the previously known northern limit for Yukon. Juncus biglumis L. Ogilvie Mountains: River flats along Dempster Highway, Mile 53, elevation 4,200 ft, No. 435; mountains east of Miles 50-54, elevation 5,500 ft, No. 475. Stewart Plateau: Keno Sum- mit, on wet frost boils with Claytonia tube- rosa, Nos. 961, 1024, and 1230. An arctic species that in Yukon is known only from a few collections on high granitic mountains (Porsild 1966). Juncus bufonius L. Vicinity of Mayo, 63°35’/N, 135°45’W, on wet, gravelly beaches of small lakes, 4 miles north of town, No. 948. Ogilvie Mountains: Demp- ster Highway, Miles 96-98, by the edge of summer-dry ponds in river meadow, No. 1720. The last is a new northern limit for Yukon. Juncus Drummondii E. Mey. -Stewart Plateau: ‘‘Highlet Gulch’, 63°45’/N, 136°10’W, in gravelly creek beds, elevation 4,000 ft, Nos. 838 and 843; vicinity of Mayo, 63°35’N, among willows in floodplain, No. 1174; Keno Summit, 63°58’N, elevation 6,000 ft, No. 1283. An alpine cordilleran species, in Yukon thus far not reported from north of the Stewart Plateau, although even there it is restricted to alpine meadows and snowbeds. Juncus filiformis L. Ogilvie Mountains: Dempster Highway, Mile 96, in damp floodplain meadows of the Black- stone River valley, No. 1717. The collection is a slight extension of the known northern limit for Yukon. Juncus filiformis is circumboreal and, throughout its range, is confined to non- calcareous lakeshores and fens; in the Mac- kenzie District it has not been collected north of the Precambrian Shield. Juncus stygius L. ssp. americanus (Buch.) Hult. Stewart Plateau: Vicinity of Halfway Lakes, north of Mayo, in wet sphagnum bog with Drosera anglica, Tofieldia glutinosa, and Carex livida ssp. americana, Nos. 888, 1218, 1241, and 1969; south branch of McQuesten River in wet bog near Elsa, No. 1167. New to the flora of Yukon. Its nearest stations are in the Alaska Range and on Great Slave Lake in the upper Mackenzie Basin. Juncus triglumis L. Ogilvie Mountains: Common in river flats and in moist spots in tundra along Dempster High- way, Miles 53 and 58, Nos. 432 and 490; south end of Chapman Lake, Mile 72, No. 1771; moist riverbank meadows, Mile 81, Nos. 1469 and 1777; Miles 85-86, damp creek bank, Nos. 1550 and 1648; Mile 110, moist flood- plain tundra, A.E. & R.T. Porsild, No. 23,609. Mature capsules average 6 mm in length and are well exserted beyond the tips of the perianth; the seeds, including tails, are 2 mm long or only slightly shorter than the measure- ment given by Fernald (1924: 201-03) for J. triglumis, but are distinctly longer than the measurement reported for J. albescens (Lge.) Fern. In No. 1771 some culms are 29 cm tall. Luzula arctica Blytt ssp. arctica Stewart Plateau: Keno Summit, elevation 5,500-6,000 ft, in alpine tundra, Nos. 1059 and 1229; east fork of Vancouver Creek; Mc- Questen area, approximately 64°N, 136°W, J.D. Campbell, No. 576. In Yukon otherwise known from two collections from the Arctic Coast and from the Pelly Mountains along the Canol Road (Porsild 1951a, 1966). Luzula arctica Blytt ssp. latifolia (Kjellm.) n. comb. L. arcuata f. latifolia Kjellm., Vega Exped. Vetensk. lakttag. 374 (1883). L. arctica var. latifolia Buchenau, Pflanzenr. IV, 36, Heft 25: 69 (1906). L. tundricola Gorodk., Herb. Inst. Bot. Acad. Sci. URSS (1935) nomen nudum; Vassiljev, Acad. Nauk SSSR Bot. Inst. Bot. Mater. Herb. 15: 40 (1953). L. beeringiana Gjaerevoll, Kgl. Nor. Vidensk. Selsk. Skr., no. 5: 63 (1958). Ogilvie Mountains: Mountain east of Miles 50-54 on Dempster Highway, in rocky saddle, elevation 5,500 ft, No. 507; 20 miles south of Chapman Lake, 64°35’N, 138°20’W, Young- man & Tessier, No. 493 (Porsild 1966: map 38). Gjaerevoll (1958) in his discussion of L. beeringiana noted that “this species has 15 Annotated List of Selected Species caused much confusion’, which undoubtedly is correct, but why this should be is not so obvious. To anyone familiar with the two subspecies of L. arctica, it should be obvious that they are, at most, geographical races; L. arctica ssp. arctica is circumpolar and truly arctic, whereas ssp. /atifolia is amphi-Berin- gian and apparently oceanic, with relatively few alpine inland stations; its eastward range, as is the case with the majority of amphi- Beringian species, does not extend beyond the Mackenzie Delta (Porsild 1951a: 129-30; 1966: map 38). Although the differences between ssp. arc- tica and ssp. /atifolia are constant and easily observed, they are by no means fundamental. Both are densely caespitose; the fully devel- oped leaves in each one (Porsild 1951a) are rarely over 4 mm wide. Lid (1963) stated that in Scandinavia the leaves of L. arctica are 3-5 mm wide, while for Greenland, Bocher, Holmen, and Jakobsen (1968) reported leaves 4.0-4.5 mm wide. In typical L. arctica Hultén (1942, 1968) reported the leaves to be 2-3 mm wide, whereas in L. tundricola (L. arctica ssp. latifolia) they are said to be 4-8 mm wide; the last figure is clearly a misprint, for leaves even 5 mm wide are indeed rare. Vassiljev (1953) noted that the leaf margins in his L. tundricola bear long, thin hairs, which is cor- rect, but they are soon lost and can rarely be seen except in young leaves; such caducous hairs are also found in ssp. arctica, whicn differs from ssp. /atifolia mainly by its dark- coloured, more compact and rarely branched inflorescence that, like the upper part of the stem as well as the upper half of the leaves, is consistently darker than in ssp. /atifolia. In the details of flower and fruit | have been unable to detect any consistent differences in the two subspecies. Whether to consider ssp. /atifolia a geo- graphical race or a good species is merely a matter of convention. For Hultén to consider ssp. /atifolia a good species is certainly in- consistent with his treatment (1968) of L. mul- tiflora, in which he deals with four subspecies (two of them with varieties). Of these varie- ties, ssp. multiflora var. Kjellmanii is as dif- ferent from L. multiflora ssp. comosa as L. arctica is from L. tundricola (L. arctica ssp. latifolia). Luzula arcuata Wahlenb. Ogilvie Mountains: Canyon Creek, Mile 48 on Dempster Highway, No. 454. Stewart Plateau: 16 Keno Summit, in alpine heath, Nos. 1026 and 1030. The present collections are the first from central Yukon. This species was pre- viously reported as common on mountains along the Canol Road (Porsild 1951a). Luzula groenlandica Bocher, Medd. Grgnland 147 (7):18, pls. 1-3 (1950). Ogilvie Mountains: Dempster Highway near Mile 57, on dry, gravelly river terraces, eleva- tion 4,200 ft, No. 230; mountain east of Miles 50-54, in saddle between rock towers, eleva- tion 5,500 ft, No. 506; boggy lakeshores near south end of Chapman Lake, Mile 62, No. 1769; moist floodplain below limestone hills near Mile 110, elevation about 3,500 ft, A.E. & R.T. Porsild, No. 23,610. The material is very uniform, and complete- ly agrees with L. groenlandica from Green- land and also from eastern arctic Canada, where it was reported from Labrador and from several places on the west coast of Hudson Bay (Cody and Porsild 1968: 266). Ripe seeds are present in all three collections cited above, and measure 1.0-1.2 mm in length; the anthers are 0.5 mm long, and the perianth leaves about 2.0 mm long. A chromo- some count of 2n = 24 was obtained from root tips grown from seeds of No. 1769, which was erroneously reported as L. multiflora (Retz.) Lej. (Mulligan and Porsild 1970). L. groenlandica is a tundra species, wide- ranging from central West Greenland across subarctic North America, from northern Lab- rador—Ungava, the shores of Hudson and James bays, and northern Keewatin and Mac- kenzie districts, and appears to be truly alpine in the Yukon and Alaska. Throughout this range it was formerly treated as L. sudetica, L. multiflora, or perhaps more often as L. mul- tiflora ssp. frigida var. arctica; in the last two taxa the perianth leaves are longer, and the chromosome count is 2n = 36 (Bocher, Hol- men, and Jakobsen 1968). L. groenlandica is “new” to Yukon and Alaska. Luzula multiflora (Retz.) Lej. ssp. frigida (Buch.) Krecz. Southwestern Yukon, Haines Road near Mile 85, C.H.D. Clarke, No. 551; mountain above Vulcan Creek, south of Kluane Lake, by mar- gin of alpine pool, Schofield & Crum, No. 8212; vicinity of Ptarmigan Heart, approximately 61°46-49’N, 138°32-37’W, in alpine tundra and muskeg, Raup, Drury & Raup, Nos. 12,698, 13,599, and 13,681; southeastern Yukon, Liliaceae Canol Road, Mile 245, south fork of MacMillan River, A.E. Porsild & Breitung, No. 11,412; central Yukon, Midnight Dome near Dawson, 64°04’N, 139°42’W, No. 1415. In the last num- ber, collected on 21 June 1968, the heads are still close together and may thus be referred to the var. contracta Sam. Apparently rare or little-collected in Yukon, where it was previously reported only from the Canol Road near the Yukon—Mackenzie divide (Porsild 1951a). Luzula rufescens Fisch. Stewart Plateau: Vicinity of Mayo on gravelly river terraces, No. 507A; Midnight Dome near Dawson, open wooded slopes, No. 1417. Ogil- vie Mountains: In willow thickets on flood- plain near Dempster Highway, Mile 67, No. 1810. An amphi-Beringian species, in Yukon previously known from the vicinity of Dawson. Beyond Yukon Territory, it has recently been collected in the Mackenzie Delta and in north- eastern British Columbia. Luzula spicata (L.) DC. Stewart Plateau: Keno Summit, in open al- pine tundra, elevation 6,000 ft, No. 1042; Mount Haldane, gravelly slope of main cirque, elevation 5,500 ft, No. 590. The present col- lections apparently extend the known north- ern limit for L. spicata in the Yukon. In eastern North America the range of L. spicata follows the familiar pattern of a low-arctic amphi- Atlantic species, extending from southern and central East and West Greenland, south- ern Baffin Island, and Labrador—Ungava to the east coast of Hudson Bay, south to New- foundiand, Gaspé, and the high mountains of New England and northern New York. The range of the western population of L. Spicata extends from the Mackenzie Moun- tains over the Alaska Range and the Pacific Coast of Alaska to Unalaska Island, and southward mainly along the Continental Di- vide to Colorado and Arizona, with disjunct populations on Vancouver Island and in the southern coastal Mountains of British Colum- bia. To the west of Bering Strait, L. spicata appears to be absent across Siberia until we reach the Ural Mountains. From the 1,500-mile wide “gap” between the eastern and western populations, we know of a few isolated stations, each disjunct by several hundred miles from the main areas: Great Bear Lake, A.E. Porsild & R.T. Porsild, Nos. 3652 and 5224, and Nisutlin Lake in northern Manitoba, Baldwin, Nos. 2211 and 2292. It would thus appear that the eastern population is amphi-Atlantic and low- arctic-alpine, whereas the western is cordil- leran North Pacific and not arctic. (Porsild 1964b: map 110). LILIACEAE Smilacina trifolia (L.) Desf. Although R.T. Porsild thoroughly explored a great many sphagnum bogs during three seasons of collecting, he was unable to add S. trifolia to his long list of range extensions. A member of the flora of the eastern boreal forest, this species barely enters south- eastern Yukon, where it has been collected once near the east end of Watson Lake, ap- proximately 60°05’N, 128°43’W, Raup & Cor- rell, No. 11,058. Apparently, it is also absent from Alaska but, in common with some other woodland species, reappears in forested parts of eastern Siberia. Tofieldia coccinea Richards. Ogilvie Mountains: In dry herbmats on moun- tains east of Dempster Highway, Miles 57-58, elevation 4,500 ft, No. 256; near lake west of Mile 57, No. 332; limestone hills west of Miles 81-85, in alpine tundra, Nos. 1099, 1489, and 1564. Noted as very common on limestone hills near Miles 110-13. Tofieldia glutinosa (Michx.) Pers. Stewart Plateau: South branch of McQuesten River in a bog west of Elsa, No. 1164; in bogs near Halfway Lakes, Nos. 505A, 862, 1242, and 1296. In Yukon otherwise known from Watson Lake, Raup & Correll, No. 11,018. This is the first documented report from Yukon. Tofieldia pusilla (Michx.) Pers. In interior northwestern America there ap- pears to be a strong discrepancy between the stature of this plant and the epithet by which it is now recognized. Thus, in the National Herbarium of Canada there are scores of sheets with individual specimens having scapes well over 20 cm tall, and quite com- monly up to 25 and 30 cm tall, bearing ra- cemes composed of up to 62 flowers. Some of the above were found in the following loca- tions. Yukon Territory: Stewart Plateau, vicin- ity of Mayo, Nos. 797, 898, and 1291; Alaska Highway near White River, J.P. Anderson, No. 9278. Alaska: Wiseman, J.P. Anderson, No. 17 Annotated List of Selected Species 5836. Mackenzie District: Mackenzie Moun- tains, Brintnell Lake, Raup & Soper, No. 9625. Elsewhere within most of its northern range, T. pusilla is very rarely over 15 cm tall, and in the Arctic and Subarctic it is quite often shorter, with short racemes often no more than 1 cm long, and rarely with more than 11 flowers, often no more than 5-6. Zygadenus elegans Pursh Noted as common, mainly on calcareous soils, north in the Ogilvie Mountains at least to Mile 81 on the Dempster Highway, No. 1501. ORCHIDACEAE Calypso bulbosa (L.) Oakes Stewart Plateau: Vicinity of Mayo in rich birch-spruce woods, Nos. 451A, 699, and 1016. In Yukon otherwise known from Daw- son and Johnsons Crossing, Mile 835 on the Alaska Highway (Porsild and Crum 1961). Cypripedium passerinum Richards. Minto Road north of Mayo, approximately 63°41-—43’N, 135°51-52’W, elevation 2,000 ft, in dry meadow among willows, No. 897; in dry meadow above Minto Bridge, No. 891. Ogilvie Mountains: Blackstone River valley, Demp- ster Highway, Mile 83, calcareous turfy open- ings in spruce woods, Nos. 1524 and 1618. Goodyera repens (L.) R. Br. Stewart Plateau: Vicinity of Mayo in mossy spruce woods with Calypso bulbosa, Nos. 474, 700, 855, and 1193. In Yukon otherwise known from Kluane Lake, White River and the Canol Road (Porsild 1951a). Habenaria obtusata (Pursh) Torr. The most common representative of the genus in central Yukon. Ogilvie Mountains: Common. in mossy spruce woods along the Dempster Highway, Nos. 1611 and 1627. Spiranthes Romanzoffiana Cham. & Schlecht. Apparently common in certain types of peaty bogs of the Mayo District. Ogilvie Mountains: Along Dempster Highway, Miles 91-93, No. 1586. A slight northward extension of its Known northern range in Yukon. 18 SALICACEAE Salix alaxensis (Anderss.) Cov. var. longistylis (Rydb.) Schneid. Vicinity of Minto Bridge, 63°41’N, 135°51/W, in thickets by small lake, No. 536. Ogilvie Mountains: Dempster Highway near Mile 110, at base of limestone hills north of road, A.E. & R.T. Porsild, No. 23,613. In Yukon not pre- viously reported from north of the Dawson area. Salix arctica Pall. Mount Haldane, 63°51’N, 135°40’W, rocky ledges above cirque, elevation 4,000 ft, No. 611; Keno Hill, 63°58’N, 135°42’W, on south- western slope, elevation 5,500 ft, No. 737. Ogilvie Mountains: Dempster Highway on mountains east of Miles 50-58, elevation 6,000 ft, Nos. 149 and 266; Mile 110, moist floodplain below limestone hills, A.E. & R.T. Porsild, No. 23,611. The present series closes a gap in the known range of this species be- tween the Dawson area and the Arctic Coast. Salix Barclayi Anderss. Ogilvie Mountains: Dempster Highway, Mile 52, elevation 4,560 ft, Youngman & Tessier, No. 364; Halfway Lakes north of Mayo, 63° 48’N, 135°47’W, bushes 5 feet tall by edge of pond, A.E. & R.T. Porsild, No. 23,564. In Yukon otherwise known only from the Canol Road (Porsild 1951a) and from a few collections in the extreme southwestern part of Yukon. Salix Barrattiana Hook. Ogilvie Mountains: On snowbed slopes by al- pine brook on mountain east of Miles 50-54 on Dempster Highway, elevation 4,500 ft, No. 168; moist subalpine slope east of Mile 51, No. 357; same place, Youngman & Tessier, No. 366; damp hollows in floodplain tundra at foot of limestone hills near Mile 110, A.E. & R.T. Porsild, No. 23,612. A cordilleran species that reaches mountains in central and northern Yukon and Alaska. Salix Bebbiana Sarg. S. depressa L. ssp. rostrata (Anderss.) Hiit. A boreal forest species, common in central Yukon north to the Ogilvie Mountains, reap- pearing again in the Porcupine River basin, where it has been collected 13 miles south- east of Lapierre House, elevation 2,300 ft, Youngman & Tessier, No. 588. Along the Mackenzie River valley, S. Bebbiana is com- Polygonaceae mon north to the Mackenzie Delta, where it has been collected near Aklavik (A.E. Lind- sey, No. 657). Salix candida Fliigge Stewart Plateau: Vicinity of Mayo, on floating margin of marly bog, No. 655. A member of the eastern boreal forest element, barely reaching Yukon and central Alaska (Porsild 1966: map 46). Salix fuscescens Anderss. S. arbutifolia auct. non Pall. Ogilvie Mountains: In sphagnum bog on river flat along Dempster Highway, Mile 54, No. 16. Also from Bell River near Lapierre House in the Porcupine River basin of northern Yukon, Youngman & Tessier, No. 608. In Yukon other- wise known only from the Arctic Coast (Por- sild 1943: 27). Salix glauca L. ssp. glabrescens (Anderss.) Hult. Ogilvie Mountains: Dempster Highway near Mile 110, erect shrub 3-4 ft high by a brook, A.E. & R.T. Porsild, No. 23,614. In Yukon the var. glabrescens was not previously collected north of latitude 64°N.. Salix planifolia Pursh McQuesten area, head of Bennett Creek, about 63°43’N, 136-38°00’W, elevation 3,800 ft, J.D. Campbell, No. 316; mouth of Bennett Creek, bushes 10 ft tall, idem, No. 332; Bell River, southwest of Lapierre House, approxi- mately 67°40’N, 136°30’W, Youngman & Tes- sier, No. 530. An eastern boreal-subarctic species, still common in the southwestern parts of the Mackenzie District and also reported from southeastern and southwestern Yukon (Por- sild 1951a, 1966), but until now not reported from central and northern Yukon. Salix Scouleriana Barr. McQuesten area, Bear Creek, approximately 63°40’N, 136°40’W, in open muskeg, J.D. Campbell, No. 296; old burn on south-facing hillside, idem, No. 820. Ogilvie Mountains: Foot of limestone hills along Dempster High- way near Mile 110, A.E. & R.T. Porsild, No. 23,614. A north-cordilleran woodland species, in Yukon not previously reported from north of Dawson. MY RICACEAE Myrica Gale L. Stewart Plateau: Swampy flats north of Mayo, No. 503A, flowering specimens with young leaves on 15 June 1967; edge of bog north- east of Mayo, No. 656, and at Mayo Lake, No. 1180. A rare and little-collected species in Yukon, where it appears to be restricted to the Dawson-—Mayo districts. SANTALACEAE Geocaulon lividum (Richards.) Fern. Mount Haldane, 63°51’N, 135°50’/W, in open spruce forest, elevation 3,500 ft, No. 621. Ogilvie Mountains: Dempster Highway, Mile 83, in open spruce forest, elevation 3,800 ft, No. 1511. A boreal forest species, common in mossy spruce forest, mainly on well-drained river flats. POLYGONACEAE Koenigia islandica L. Ogilvie Mountains: River flats along Dempster Highway, Miles 57-58, flowering specimens on 18 July 1966, No. 229, and with flowers and young fruits on 7 Aug. 1966, No. 445; Mile 47, on wet gravel in creek bed with almost mature fruits on 14 Aug. 1968, No. 1851. Ap- parently a rare, or no doubt often overlooked, plant in Yukon, where it is otherwise known only from two widely disjunct stations along the Canol Road and on Bennett Lake (Porsild 1951a). Oxyria digyna L. Ogilvie Mountains: Common locally along Dempster Highway, Mile 47, No. 1844, fruiting on 14 Aug. 1968. A chromosome count of 2n= 14 confirmed earlier reports from Green- land (Bocher, Holmen, and Jakobsen 1968). In central Yukon found mainly in places where snow or overflow ice persists until late into the summer. Polygonum alaskanum (Small) Wight Stewart Plateau: Keno Hill, in moist ravines, elevation 4,500-—5,500 ft, Nos. 999 and 1290. Ogilvie Mountains: River flats along Dempster Highway, Mile 63, No. 347. Peel Plateau: “La- pierre River’ and Richardson Mountains, Youngman & Tessier, Nos. 516 and 586; Old Crow River, /idem, No. 641. 19 Annotated List of Selected Species In the collections cited above, the leaves are glabrous except for the sparsely ciliate margins, and may thus be designated var. glabrescens Hult. (Porsild 1966: map 52). Polygonum caurianum Robins. Stewart Plateau: South branch of McQuesten River, in disturbed soil near Elsa, No. 1150. Apparently rare in Yukon, where it has been collected only a few times (Porsild 1951a). Polygonum O’Neillii Brenckle P. pennsylvanicum L. ssp. Oneillii (Brenckle) Hult. Stewart Plateau: Vicinity of Mayo on wet, gravelly lakeshore, Nos. 949 and 1007, in be- ginning anthesis in the last week of July; same place with ripe achenes, No. 1309, 19 Aug. 1967, A.E. & R.T. Porsild, No. 23,558. In both collections the leaves are narrowly lan- ceolate, 6 to 7 times longer than broad, to- mentose in youth, glabrate in age. The speci- mens closely match a collection from Nak- nek, Alaska Peninsula, E. Lepage, No. 24,111, determined by Brenckle, except that the peduncles of our specimens are totally gla- brous, whereas in the Lepage specimen a few minute, stipitate glands are present (Hultén 1967: 46). The collections are the first from Canada, otherwise known only from southern and southwestern Alaska. Rumex arcticus Trautv. Very common in damp, turfy meadows and river flats along Dempster Highway, ascend- ing to at least 3,000 ft elevation, Nos. 155, 850, 1141, 1642, 1806, and A.E. & R.T. Porsild, No. 23,616, the last at Mile 110 on the Blackstone River. A chromosome count of 2n=40 was obtained from No. 155 (Mulligan and Porsild 1969). Rumex occidentalis S. Wats. Vicinity of Mayo, 63°31’N, 135°54’W, willow flat north of town, No. 1146; Hungry Lake, 65°40’N, 136°00’W, in mossy, wet places along stream, Youngman & Tessier, No. 697. Rumex triangulivalvis (Danser) Rechn. fil. Dawson, on old gold dredge tailings, No. 544, with mature fruit on 19 Aug. 1968; same place, M.O. Malte, No. 282; Teslin Lake, M.P. Porsild, No. 774; Alaska Highway, Mile 1022, on dry roadside, Schofield & Crum, No. 8256. A chromosome count of 2n= 20 was obtained from No. 544 (Mulligan and Porsild 1969). 20 PORTULACACEAE Claytonia arctica Adams Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 50-54, in moist gravelly places, elevation 5,800 ft, fruiting specimens on 15 Aug. 1966, No. 447 (Plate 2). A chromo- some count from seedlings grown in Ottawa gave 2n=10 (Mulligan and Porsild 1969). The collection is the first from Canada, and is an interesting extension of the range of a spe- cies that is also rare and little-collected in Alaska, where, moreover (for reasons | shall document below), it has generally been con- fused with C. sarmentosa C.A. Mey. The following Alaska specimens are in the National Herbarium of Canada, Ottawa: Brooks Range, Peters Lake, 69°20’N, 145° 00’W, common in wet niggerhead tundra, Raymond D. & Mildred Wood, No. 341, and colour photographs Nos. 469-1961; ‘‘Lake Schrader’, approximately same latitude and longitude, elevation 3,000 ft, iidem, No. 1878; Anaktuvuk Pass, 68°17’N, 151°25’W, L.A. Spetzman, No. 784 (the last two distributed as C. sarmentosa); Cape Lisburne, Seeman, summer of 1850, photograph of original speci- men in Herb. Kew. Although Hultén (1944) and others accept- ed Seeman’s report of Claytonia sarmentosa C.A. Mey. from Cape Lisburne on the north- west coast of Alaska (Seeman 1852:27), | was puzzled, because Seeman’s fine plate (1852: pl. 5) purported to represent C. sarmentosa, but is in fact C. arctica Adams, a little-known species in North America reported by Hultén (1937, 1968) only from the westernmost islands of the Aleutian chain, and in Asia known from northern Kamchatka, the Com- mander Islands, and from widely disjunct stations across arctic Siberia. The description of C. sarmentosa in See- man (1852) is copied verbatim from C.A. Meyer (1829:137), and clearly does not apply to the plant Seeman illustrated (1852: pl. 5). At Kewin 1954, | examined the well-preserved Seeman specimens from Cape Lisburne, and at once confirmed that Seeman had indeed collected C. arctica Adams, rather than C. sar- mentosa, but had failed to recognize it as C. arctica. Two of the 12 specimens on the sheet preserved at Kew show plants with taproots, so well described by Adams (1817:94): “‘ra- dice fusiformi longa carnosa, caudice brevi turbinato simplisissimo haud stolonifera...’’; in the remainder of the Seeman specimens, Caryophyllaceae Plate 2 Claytonia arctica Adams Approximately 73 natural size the taproots were lost during the collecting, but the thickened lower part of the stem is still there, and shows the place of attachment. In none of the 12 specimens are runners pres- ent, such as those formed in C. sarmentosa. The label on the sheet reads: ‘3339. N.W. America, Seeman. 1850’. Across the upper right corner W.J. Hooker had written ‘“Clay- tonia sarmentosa C.A. Mey.”’. In 1957 | was unable to discover the type of C. arctica Adams in the Komarov Herbarium in Leningrad, where it is said to be preserved. However, some other specimens more or less contemporary with the type, labelled C. arc- tica, agreed well with the plant illustrated by Seeman (1852: pl. 5), and also agreed with the photograph of the Kamchatkan C. arctica (Hultén 1928: pl. 2). On the other hand, some recent collections, for example, those from Wrangel Island, 8 Aug. 1938, Gorodkov, are depauperate and condensed, with a short and stouter taproot, and shorter basal leaves forming a rosette at the head of the taproot. Rydberg (1932:302) cited Claytonia sar- mentosa, Seem., not C. sarmentosa C.A. Mey., under C. arctica Adams, thereby showing that he recognized Seeman’s illustration (1852: pl. 5) as acorrect representation of C. arctica Adams; however, he failed to point out that the plant described by Seeman (1852) is C. sarmentosa C.A. Mey. Rydberg’s opening statement, “. . . fleshy taproot 5-10 cm thick”’ is obviously a misprint and should have been “5-10 mm”. The Siberian range of C. arctica Adams, according to Tolmatchev (1966: map 67), is from the Lena Delta to the head of the Gulf of Anadyr, with a disjunct population in Wrangel Island. Incidentally, C. arctica Adams was described as being one-flowered, with pale- yellow petals (dilute flavis). Hultén (1928: pl. 2) shows one Kamchatkan plant with a single- flowered scape, and a second in which one flower is fully expanded and two are still un- opened; the petals are said to be white. In the Yukon and Alaska C. arctica usually has more than one flower, and the petals are pale pink with purple veins. Claytonia tuberosa Pall. Stewart Plateau: Keno Hill in alpine tundra, elevation 5,500-6,000 ft, Nos. 753-54 and 1255, A.E. & R.T. Porsild No. 23,571; on rich herbmat slopes, No. 998. In stony and gravelly places on Keno Sum- mit, the corms of C. tuberosa were frequently noted close to the surface when rarely more than one flowering stem is produced. This condition is obviously due to frost heaving, for when they grow in more sheltered places with ample snow cover in winter, the corms are invariably well below the surface, and normally produce several flowering stems, but never “‘long, filiform, very brittle runners’, as reported, but not illustrated, by Hultén (1968: 405). C. tuberosa is of amphi-Beringian range, but in Alaska and Yukon it appears to be rare and always of local and disjunct oc- currence. In Yukon it was first reported from the McArthur Group, of which Keno Hill is the northern spur (Porsild 1951a; 1966: 63). CARYOPHYLLACEAE Arenaria humifusa Wahlenb. A. longipedunculata Hult., Bot. Notis. 119 (2): 313 (1966). Stewart Plateau: Vicinity of Minto Bridge, 63°41’N, 135°51’W, very sparse and local on open gravel patches on wooded slope, eleva- tion 2,000 ft, No. 928. Apparently a rare spe- cies in Yukon and Alaska. In interior Alaska, the Yukon, and the Mac- kenzie District of the Northwest Territories, A. humifusa tends to be of tall growth, with the fruiting peduncles raised well above the plant mat, which may be firm or loosely tufted, 21 Plate 3 Melandrium apetalum (L.) Fenzl. ssp. ogilviense n. ssp. Flowering specimens. Approximately % natural size 22 Caryophyllaceae with long, creeping branches. Hultén (1966) considered this taller plant distinct from A. humifusa, but an examination of a representa- tive material of A. humifusa from Alaska, the Yukon, and the Mackenzie District, as well as from isolated mountain stations in Alberta and British Columbia, failed to disclose any consistent characters whereby long-pedun- cled plants can be distinguished from low- grown A. humifusa, which, furthermore, is found coexisting with A. “/ongipedunculata”’ throughout its range. That this is also the case in the eastern North American Arctic was noted by Polunin (1940: 198), who wrote: “Thus, in most southern specimens the pedi- cels are elongated sufficiently to carry the capsule clear of the tuft of leaves that in the North almost envelops the bud and some- times even the capsule.” It would appear then that the elongation of the peduncle in A. humi- fusa is merely the result of a favourable mi- croclimate or local edaphic conditions. A. cylindrocarpa Fern. (Fernald 1914), from Newfoundland and Gaspé, Quebec, with free- ly branching stems and fruiting peduncles up to 3 cm long, which Fernald himself later re- duced, differs no more, at least in appear- ance, from A. humifugsa Wahlenb. than does A. longipedunculata Hult. Cerastium vulgatum L. Vicinity of Mayo, 63°35’N, 135°54’W, on grav- elly river terraces, where obviously fully es- tablished and flowering abundantly on 14 June 1967, No. 506. Numerous last season’s fully formed but now empty capsules are still attached. In Yukon the species was first col- lected near Dawson in 1899 by R.S. Williams and J.B. Tarleton (Britton and Rydberg 1901). Melandrium affine (J. Vahl) Hartm. Ogilvie Mountains: Dempster Highway along Miles 44-46, in subalpine herbmat, elevation 3,800 ft, No. 1436, with almost ripe seeds on 25 June 1968; Miles 83-86 on rocky ledges in canyon of Blackstone River, No. 1632. In Yukon previously reported from a few col- lections along the Canol Road and along the Arctic Coast (Porsild 1951a). Melandrium apetalum (L.) Fenzl. ssp. arcticum Hult. Keno Summit, 63°58’N, 135°42’W, on solifluc- tion lobes, elevation 5,500-6,000 ft, No. 1074. Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 50—54, in alpine herbmat, No. 183; near summit of same mountain, ele- vation 6,500 ft, No. 1569; gravelly river flat near Mile 57, No. 222; northern slope of mountain east of Miles 54-57, No. 389. The above series is rather uniform and con- sists of rather small tufted plants, with slen- der stems rarely over 20 cm tall; the single flower is nodding in anthesis, but stiffly erect when in seed. A chromosome count based on No. 1078, erroneously published as M. Tay- lorae, gave 2n=24 (Mulligan and Porsild 1970). Melandrium apetalum (L.) Fenzl. ssp. attenuatum (Farr) Hara, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6:42 (1952). Keno Summit, 63°58’N, 135°42’W, elevation 5,500-6,000 ft, on gravelly slide above creek beds, No. 710. Ogilvie Mountains: Along Dempster Highway near Miles 83-86, on dry ledges and slopes below canyon wall, No. 1631. As | have previously stated (Porsild 1966: 27-28, map 59), there are now good and suffi- cient reasons for considering this taxon a dis- tinct cordilleran race, or perhaps even a spe- cies, related to, but constantly distinct from, M. apetalum, the range of which rarely, if ever, overlaps that of ssp. attenuatum. Melandrium apetalum (L.) Fenzl. ssp. ogilviense n. ssp. Caules 40-60 cm longi, erecti, pilis crispulis sparsis vestiti; folia anguste-oblanceolata 5-6 cm longa et 3-5 mm lata, uninervia, ad margi- nem ciliata. Flos unicus vel plerumque 2 (3), per anthesin nutans, postea erectus, pedun- culo ad 10-20 cm longo; calyx 1.5 cm longus, fructifer inflatus, obscuro-striatus; corolla purpurea calyce sublongior; semine renifor- mi, 2.0-2.4 mm lato, ala lata inflata, 2n= 48. Ogilvie Mountains: Dempster Highway, Mile 81, calcareous river flat along Black- stone River, elevation 2,300 ft, flowering on 3 July 1968, No. 1545 (Plate 3); same place, with mature fruits, 19 July 1968, No. 1612A (Type: CAN), Plate 4; Blackstone River valley, Miles 83-85, on calcareous talus slope, fruit- ing on 23 July 1968, No. 1630; on river flat, No. 1557; Dempster Highway, Mile 115, in damp, calcareous tundra flat, No. 1975. Chro- mosome count on root tips from No. 1612 gave 2n = 48 (Mulligan and Porsild 1970, where reported as M. apetalum). The ssp. ogilviense differs from M. apeta- lum ssp. apetalum by its stems up to 60 cm 23 Plate 4 Melandrium apetalum (L.) Fenzl. ssp. ogilviense n. ssp. (Type). Fruiting specimen. Approximately 24 natural size 24 Caryophyllaceae tall, by the number of flowers, 2-3, rarely one, and by its higher chromosome count. The seeds are similar to those of ssp. apeta/um, being 2.0-2.4 mm in diameter, including the broad, more or less inflated margin. The ssp. ogilviense is common locally on calcareous soils in the Ogilvie Mountains between Miles 81-150, but not seen elsewhere, and is ap- parently endemic in central-northern Yukon. Melandrium Taylorae (Robins.) Tolm. Vicinity of Mayo, 63°35’/N, 135°45/W, on grav- elly river terraces, No. 469; Mount Haldane, 63°51’N, 135°40’W, floor and bottom slopes of main cirque, elevation 4,300 ft, in slide- rock, No. 581; Midnight Dome near Dawson, 64°04’/N, 138°15’/W, in open woods near sum- mit, No. 1423. Ogilvie Mountains: Blackstone River valley, gravelly river flats, along Demp- ster Highway, Miles 81-82, Nos. 1121, 1472, and 1531. A chromosome count from root tips grown from seed from No. 1121 gave 2n=48 (Mulligan and Porsild 1969). The material is rather uniform, with few to many slender flowering stems, up to 40 cm tall, from a rather weak taproot. Nos. 469, 581, 1423, and 1472 are in anthesis when the 3-5 flowers are congested on short peduncles that elongate as the seeds ripen and, at ma- turity, may be 3-10 cm long. The mature cap- sules are 6-8 mm long, and the seeds about 1 mm in diameter, with a narrow but distinct wing. Although the peduncles are not actual- ly glandular, the seeds are often seen to ad- here to the moniliform, transversely walled hairs covering the peduncles. In Yukon other- wise known from a few collections from the Dawson area. In habit M. Tay/orae resembles M. triflorum (R. Br.) J. Vahl of northern Greenland and the eastern Canadian Arctic, and also the west- ern M. taimyrense Tolm., but differs from them by its reddish-brown seeds that are flat and have a narrow but distinct wing, whereas in M. triflorum and M. taimyrense the seeds are almost black, angular, or prismatic, and entirely wingless (Porsild 1943). Minuartia arctica (Stev.) Aschers. & Graebn. Stewart Plateau: Mount Haldane, elevation 5,000 ft, on slide-rock, No. 667; Keno Summit, forming large cushions in alpine tundra, ele- vation 6,000 ft, Nos. 538 and 704. A chromo- some count of 2n=ca. 80 was obtained from No. 704 (Mulligan and Porsild 1969). Ogilvie Mountains: River flats along Dempster High- way, Mile 56, No. 53; mountain east of Demp- ster Highway, elevation 5,000-6,000 ft, Nos. 146 and 404; Mile 82, in calcareous alpine tundra, No. 1495; moist floodplain tundra at foot of limestone hills, Mile 110, A.E. & R.T. Porsild, Nos. 23,617 and 23,619. A calci- philous alpine species common locally in Yukon, but collected only a few times north to the Arctic Coast (Porsild 1951 a). Minuartia biflora (L.) Schinz & Thell. Arenaria sajanensis Willd. Mayo District: Keno Hill, 63°58’N, 135°42’/W, in alpine herbmats near summit, elevation 4,500-6,000 ft, Nos. 954, 1027, and 1256. Ogil- vie Mountains: On river flats along Dempster Highway, Mile 51, elevation 4,200 ft, No. 123. Minuartia dawsonensis (Britt.) Mattf. Minuartia stricta sensu Hult., Fl. Alaska & Yukon (1944), pro max. pte., not M. stricta (Sw.) Hiern., nor Arenaria stricta Michx., FI. Bor.-Am. 1:274 (1803). Stewart Plateau: Minto Bridge, damp mead- ows by lakeshores, No. 529; gravelly river banks, Nos. 922 and 940. Hunker Creek near Dawson, on gravelly creek bank, No. 1399. Ogilvie Mountains: Dempster Highway, in damp, sandy, and gravelly places, near Miles 83-86, Nos. 1558 and 1624; Miles 96-98, No. 1712. A lowland species of quite different dis- tribution and ecology than the smaller and arctic-alpine A. uliginosa Schleich. (Minuartia stricta (Sw.) Hiern.), with which it is some- times confused. For a detailed description of differences and habit, see Porsild 1951a: 166. Gjaerevoll (1958: map 8) is mistaken in be- lieving A. dawsonensis is endemic in central Alaska and Yukon. It is actually a wide- ranging eastern species, with a western limit in central Alaska (Fernald in Gray 1950: 620). Minuartia obtusiloba (Rydb.) House Minuartia yukonensis Hult., Ark. Bot. ser. 2, vol. 7 (1):52 (1967). Dawson area: Dominion Creek, 63°37’N, 138°42’W, in damp alpine tundra, No. 1407; Midnight Dome, 64°04’N, 139°42’/W, in open woods near summit, No. 1422. Mayo district: Moose Creek, 63°37’N, 136°25’W, sandy creek banks, No. 990. Ogilvie Mountains: Dempster Highway, Mile 84, in moist herb- mats, No. 1672; Richardson Mountains: 13 miles northeast of Lapierre House, Young- man & Tessier, No. 591; Firth River 68°50’N, 140°33’W, iidem, No. 154. 25 Annotated List of Selected Species Hultén (1943), with some reservations, fol- lowed Gray (1895) and Fernald (1919), who identified the plant then known from ungla- ciated central Yukon and Alaska with Arena- ria laricifolia L. (Minuartia Jaricifolia (L.) Schinz & Thell.), observing that Gray and Fernald also had wondered if the North American plant really could be conspecific with that of central European mountains. Without offering new evidence, Hultén (1967), on geographical rather than morphological grounds, proposed Minuartia yukonensis n. sp. for the Yukon and Alaska plant that he had formerly called Arenaria /aricifolia. As the type, he selected Calder & Billiard, No. 3359, from near the mouth of Klondike River (near Dawson City). It is not clear, however, if Hutén intended to include under M. yukon- ensis the plant that he referred (1943:682) to M. laricifolia from “‘south of the pleistocene glaciation from Washington and Montana to Idaho and Wyoming”, which, according to Hitchcock and Cronquist (1964), is A. obtusi- loba (Rydb.) Fern. Unfortunately, Hultén’s diagnosis of M. yukonensis is so general that it fits equally well not only the central Euro- pean plant, from which he intended to sepa- rate M. yukonensis, but also the plant of Washington, Montana, and Wyoming, which, incidentally, is also found in the Canadian Rocky Mountains. A. obtusiloba, in the Rocky Mountains as well as in the Yukon and Alas- ka, is quite variable in general habit as well as in regard to degree and type of pubes- cence. In moist herbmats it tends to be loose- ly matted, whereas on slide-rock or in ex- posed situations it is caespitose, often ‘‘an- chored”’ by a thick, subligneous taproot. Hultén’s illustration of M. obtusiloba (1968: 430) was drawn from a condensed plant that had probably grown in a dry, exposed habitat. In the Yukon as well as in the Rocky Moun- tains, specimens that match Hultén’s illustra- tions of either Minuartia obtusiloba or M. yukonensis can be found on the same moun- tain, but on different exposure or elevation (Hultén 1968:432) . A. obtusiloba was also reported from the east branch of the Mackenzie Delta (Porsild 1943:30). Several published chromosome counts of the European Minuartia (Arenaria) laricifolia report 2n= 26, which is also the number ob- tained for A. obtusiloba from Boulder, Colo- rado (G.A. Mulligan, verbal communication). 26 Minuartia Rossii (R. Br.) Graebn. ssp. elegans (Cham. & Schlecht.) Maguire Ogilvie Mountains: River flats along Dempster Highway, Mile 57, No. 224, with young fruits on 18 July 1966; on hummocks in alpine cal- careous tundra, Miles 81-83, Nos. 1498 and 1523; calcareous, stony floodplain, Mile 110, A.E. & R.T. Porsild, No. 20,618. The present collections close a gap between southwest- ern Yukon (Porsild 1966) and the Arctic Coast. Silene acaulis L. var. subacaulescens (Williams) Fern. Ogilvie Mountains: Along Dempster Highway near Mile 44, in turfy places by brook, eleva- tion 3,800 ft, No. 1440; Mount Haldane, 63° 51’N, 135°50’W, northern slope on rocky ledges of cirque, No. 570. Typical S. acaulis were collected in the Ogilvie Mountains near Dempster Highway, Mile 53, in alpine herb- mats, elevation 4,200 ft, No. 113. The present reports are the first for central Yukon, where both taxa are apparently rare and alpine (Porsild 1951a: 170-71). Spergularia rubra (L.) Presl Minto Road, 13 miles north of Mayo, in grav- elly place by creek, elevation 2,000 ft, Nos. 512 and 646. A chromosome count of 2n=36 was obtained from No. 646 (Mulligan and Porsild 1969); “Highlet Gulch’, a tributary to Minto Creek, 63°45/N, 136°10’W, in gravelly places by creek, elevation 4,000 ft, No. 837. An introduced weedy species, but apparently well established in the Mayo mining area. The present is the first report of this species from the Yukon Territory. Stellaria Alsine Grimm Minto Road, 13 miles north of Mayo, along banks of small creek near road, elevation 2,000 ft, where it forms dense clumps with stems 30 cm tall; the plants were fully with- ered, with masses of ripe seeds on 26 Aug. 1967, No. 1239, 2n=52 (G.A. Mulligan and A.E. Porsild, unpublished). An introduced, but locally well-established, weedy species new to the flora of Yukon. Stellaria crassifolia Ehrh. Vicinity of Mayo, 63°35/N, 135°45’W, in moss by small lake in the shade of birch and wil- lows, No. 1196. Ogilvie Mountains: Dempster Highway, Mile 95, in wet sand by a pond, No. Ranunculaceae 1734. The last number bridges a gap between the Dawson area and the Arctic Coast. NYMPHAEACEAE Nuphar polysepalum Engelm. Stewart Plateau: Vicinity of Mayo, in small lake north of town, in full flower on 13 July 1967, No. 808; small lake west of Elsa, No. 1166, with fully ripened seeds on 11 Aug. 1967. Ogilvie Mountains: Dempster Highway in lake near Miles 74-75, Nos. 315, 1786, and 1815; Gravel Lake on Stewart River, 60 miles south of Dawson, No. 187. RANUNCULACEAE Anemone multiceps (Greene) Standley Hultén’s statement (1967:56; 1968:466) that Anemone multiceps is merely “a low-grown arctic form of A. Drummondii”’ is very difficult to understand. As confirmed also by Boraiah and Heimburger (1964:919), “The Pulsatilla- like appearance of its flowers with deep red styles and blue anthers [and filaments], make the species morphologically distinct [from A. Drummondii], as does its tufted habit so well characterized by the’specific name given it by Greene, as against the less compact habit of A. Drummondii.” In the Richardson and British mountains of northern Yukon and in the mountains of northern and northwestern Alaska, A. multi- ceps and A. Drummondii may grow together, but in no instance have | noted specimens that in any way could be considered interme- diate. By its dwarfed and compact habit and dark purplish-blue flowers up to 4 cm broad, itis a strikingly beautiful species. In A. Drum- mondii the flowering stems are commonly twice as tall as those of A. multiceps, the ex- panded flowers usually 2 cm and very rarely 3 cm in diameter, the sepals creamy white or pale bluish dorsally, and the anthers and fila- ments always creamy white, as are the styles. A. multiceps is apparently endemic in cal- Careous areas in the mountains of northern Yukon and Alaska, and thus far has not been reported east of the 140th meridian. A. Drum- mondii is a herbmat species, and in the Cana- dian Rocky Mountains often ascends far above the timberline, where it may be dwarf- ish of habit but never even remotely simulates A. multiceps. A. multiceps is from Cape Thompson, Alas- ka (Plate 5). Caltha natans Pall. Ogilvie Mountains: In wet, swampy places along Dempster Highway, Miles 90-93, Nos. 1584 and 1746, the last with fully ripened fruits on 2 Aug. 1968, which provided a chro- mosome count of 2n = 32 (Mulligan and Por- sild 1970). Caltha palustris L. ssp. arctica (R. Br.) Hult. Ogilvie Mountains: Common in ponds and marshes along Dempster Highway, Miles 55— 60, No. 2. Delphinium glaucum S. Wats. Vicinity of Mayo, 63°35’N, 135°45’W, boggy depression in river terrace, No. 857. Ogilvie Mountains: Dempster Highway, Mile 57, on river terraces, No. 370; Mile 86, on wooded slope along Blackstone River, No. 1666; Mile 110 among willows in wet floodplain, A.E. & R.T. Porsild, No. 23,624. These collections bridge a gap in the distribution of D. glaucum between central Yukon and northern Richard- son Mountains. Pulsatilla ludoviciana (Nutt.) Heller Mount Nansen, growing with P. ludoviciana f. glabrata, No. 1385; Mayo district, 63°35/N, 135°55’W, on gravelly river terraces, eleva- tion 1,600 ft, No. 449, with fully mature fruits on 31 May 1967. Pulsatilla ludoviciana (Nutt.) Heller f. glabrata n. forma Planta omnina glabra, sepalis albis. A single, totally glabrous and white-flow- ered specimen, growing among typical P. ludoviciana, the flowers and several leaves apparently normal and functional. Mount Nansen, 62°06’/N, 137°17’W, 40 miles west of Carmacks, top of scree, No. 1386 (Type: CAN). Ranunculus abortivus L. Stewart Plateau: Minto Road, 13 miles north of Mayo, shores of small lakes and creeks, No. 509A. In Yukon otherwise known from the Canol Road, where | believed it to be fully indigenous (Porsild 1951a). Ranunculus aquatilis L. var. eradicatus Laest. Stewart Plateau: Crystal Lake northwest of Keno Hill, in water 0.5 m deep, No. 1237; in small lake on the northeastern slope of Mount Haldane, No. 1147. Ogilvie Mountains: Shal- low pool in river flat along Dempster High- 27 Plate 5 Photograph by Raymond D. Wood) ( Anemone multiceps (Greene) Standley tely % natural size ima Approx 28 Ranunculaceae way, Miles 51-58, Nos. 302 and 438 (Porsild 1966: 29). Ranunculus aquatilis L. var. subrigidus (Drew) Breitung R. circinatus Sibth. var. subrigidus (Drew) Benson Stewart Plateau: Minto Road, 13 miles north of Mayo, in shallow creek, with ripe seeds on 13 Aug. 1967, No. 1184, 2n = 32 (Mulligan and Porsild 1969). Ogilvie Mountains: Lake west of Dempster Highway, Mile 58, No. 328; Mile 82, Chapman Lake in water 2 m deep, flowering on 5 Aug. 1968, No. 1757 (Porsild 1966). " Ranunculus Eschscholtzii Schlecht. Stewart Plateau: Keno Summit, on moist herb- mat slopes, elevation 5,500—6,000 ft, Nos. 522, 723, 777, 1039, 1043, and 1044; Mount Hal- dane, 63°51’N, 135°50’W, snowbed slopes in open Abies forest, elevation 4,000 ft, No. 605; alpine slopes to 5,500 ft, No. 596. Ogilvie Mountains: West of Dempster Highway, Mile 51, No. 194; Mile 58, near snowflush, Nos. 331 and 491, where it grew associated with A. ni- valis. No. 723 from Keno Summit and No. 331 from the Ogilvie Mountains represent a pe- culiar peloric form in which all, or at least some, of the stamens are petaloid and deeply notched, or even 3-4-fid. R. Eschscholtzii is the most common representative of the genus in alpine herbmats of central Yukon. Ranunculus gelidus Karel. & Kiril. Ogilvie Mountains: Mountains east of Miles 57-58 on Dempster Highway, on slide-rock, elevation 6,000 ft, No. 257, 2n = 16 (Mulli- gan and Porsild 1969). See also Porsild 1966: map 65. Ranunculus Gmelini DC. Minto Bridge, 63°41’N, 135°51’W, in shallow ponds with Potamogeton and Myriophyllum, No. 881; Halfway Lakes north of Mayo, 63°48’ N, 135°47’W, in shallow pond, No. 640. Ogil- vie Mountains: Dempster Highway, in ponds along Miles 83-86, No. 1629. Ranunculus hyperboreus Rottb. Ogilvie Mountains: River flats along Dempster Highway, Miles 54—56, on wet clay by edge of pond, elevation 2,500 ft, No. 11; flowering on 27 June 1966, Nos. 442 and 443, with flowers and ripe fruits on 6 Aug.; Keele Lake area, 63°31’N, 130°28’W, W.H. Butler, No. 19. A chromosome count of 2n = 32 was obtained from No. 442 (Mulligan and Porsild 1969). Ranunculus lapponicus L. Vicinity of Mayo, 63°35’/N, 135°45’W, in damp, mossy willow thicket, No. 630; upper Rosebud Creek, 63°17’N, 138°25’W, J.D. Campbell, No. 101; Hungry Lake, 65°40’N, 136°00’W, Youngman & Tessier, No. 699. Ranunculus natans C.A. Mey. var. intertextus (Greene) Benson Vicinity of Minto Bridge, 63°41/N, 135°51’W, elevation 1,900-2,000 ft, in swampy pools, No. 547. The specimens collected on 21 June 1967 have flowers and young fruits. By its larger size, distinctly reniform leaves, some up to 1.5 cm wide, and by its globose fruiting heads of 40 or more achenes, R. na- tans var. intertextus is at once distinguished from R. hyperboreus Rottb. New to the flora of Yukon, and separated by a gap of 1,000 miles from the nearest known station of R. natans in the Alberta Rocky Mountains. Ranunculus nivalis L. Ogilvie Mountains: Rather common in alpine tundra in mountains east of the Dempster Highway, Miles 50-58, Nos. 94, 342, 343, 408, and 465. Stewart Plateau: Keno Summit, in al- pine tundra, elevation 6,000 ft, No. 721; Mount Haldane, in damp moss, elevation 4,000 ft, No. 572. An arctic species in central and southern Yukon, restricted to alpine habitats. Ranunculus occidentalis Nutt. var. brevistylis Greene Since first reported from Yukon (Porsild 1951a: 181), this taxon has been collected twice in southwestern Yukon: Alsek River val- ley, north side of Lowell Glacier, 60°11’/N, 137°19’W, A.M. Pearson, No. 67-162A; Bul- lion Creek, 60°58’N, 138°36’W, elevation 4,000 ft, idem, No. 70-9. Ranunculus Purshii Richards. Stewart Plateau: Along road to Moose Creek, 63°37’N, 136°25’W, in shallow water, No. 984. An eastern boreal forest species, in Yukon thus far reported a few times in the southwest (Porsild 1966: map 66). Of similar habit as R. Gmelini, which, however, is a smaller and an amphi-Beringian plant with a chromosome count of 2n = 32 or 2n = 16, whereas that of 29 Annotated List of Selected Species the eastern boreal forest species, R. Purshii, is 2n = 64 (Mulligan and Porsild 1969). Ranunculus pygmaeus Wahlenb. Stewart Plateau: Keno Hill, in wet gravel, ele- vation 5,500 ft, No. 962. Mayo district: From Black Ram to Hotspring Creeks, H. Bostock, No. 209. Ogilvie Mountains: Alpine herbmats and snowflushes on mountain east of Demp- ster Highway, Miles 50-54, elevation 4,800 ft, No. 164; mountain north of McQuesten River, R.L. Christie, No. 43; Mount Decoeli in south- western Yukon, by alpine streamlet, Scho- field & Crum, Nos. 8056 and 8075. In Yukon otherwise known from the Canol Road and from the Arctic Coast (Porsild 1951a). Ranunculus reptans L. Ogilvie Mountains: Dempster Highway, Mile 54, in small summer-dry pond, No. 457, asso- ciated with R. hyperboreus, No. 443. In Yukon previousy known only north to Mayo. Ranunculus sceleratus L. ssp. multifidus (Nutt.) Hult. Stewart Plateau: Vicinity of Mayo, lakeshore northeast of town, No. 674; in meadow near Five Mile Lake, No. 780; riverbank under shading willows, No. 1216, 2n = 64 (Mulligan and Porsild 1969). Ogilvie Mountains: Demp- ster Highway, Mile 81, in wet gravel, No. 1529; Carmacks Road, 71 miles north of White- horse, approximately 61°40’N, wet, calcar- eous lakeshore, Nos. 1860 and 1870. Ranunculus septentrionalis Poir. Stewart Plateau: Vicinity of Mayo, 63°35/N, 135°45’/W, edge of slough by mouth of Mayo River, No. 791. An eastern species of marshy woodland situations of which only a few wide- ly disjunct western stations are known: East of Fort Liard in the southwestern corner of the Mackenzie District (Jeffrey 1961); Nordegg, in southwestern Alberta, M.O. Malte, No. 1438. New to the flora of Yukon. Thalictrum alpinum L. Ogilvie Mountains: Mountain east of Demp- ster Highway, Mile 54, elevation 5,000 ft, No. 165; subalpine herbmats on limestone, Miles 81-85, Nos. 1110, 1467, and 1561; very com- mon on lower slopes of limestone hills near Mile 110, A.E. & R.T. Porsild, No. 23,623; Bon- net Plume Lake, 64°20’N, 132°00’W, Butler and Olsen, No. 44. 30 Thalictrum sparsiflorum Turcz. var. Richardsonii (A. Gray) Boivin Vicinity of Mayo, Nos. 696, 923, and 1205. Ogil- vie Mountains: In willow thickets along Demp- ster Highway, Mile 95, No. 1746. The last is its known northern limit in Yukon. A chromo- some count of 2n=14 was obtained from No. 1205 (Mulligan and Porsild 1969). FUMARIACEAE Corydalis pauciflora (Steph.) Pers. Apparently common in alpine herbmats on “Keno Hill Dome”, Nos. 764 and 1077, on Mount Haldane, No. 600, as well as-in the Ogilvie Mountains, north to about Mile 65 on Dempster Highway, Nos. 63 and 129; the last is the var. albiflora Porsild, described from the Canol Road in southeastern Yukon (Por- sild 1951a). PAPAVERACEAE Papaver alaskanum Hult. f. stenopetalum n. forma Petalis anguste-oblanceolatis, 2.0 cm longis, 0.5 cm latis differt. Ogilvie Mountains: River flat along Demp- ster Highway, Nos. 19A and 19B (Type: CAN), from a colony of tall plants, differing from typical P. alaskanum by their narrowly oblan- ceolate petals 2.0 cm long and 0.4-0.5 cm wide. While in most plants in the colony the flowers appeared sexually normal, the an- thers in some had been converted into pecu- liar leathery scales, 4 mm long and 2 mm wide, dorsally covered with stiff black bristles similar to those normally found on a Papaver capsule, and fringed by a thin, yellowish- green ruffle or pleat along the edge. In none of these peloric flowers were there normal stamens, whereas the yet immature ovary and stigma appeared to have developed normally, though they were obviously nonfunctional. A chromosome count obtained from seed grown from a nearby and quite normal colony of P. alaskanum, No. 365, gave 2n = 42 (Mul- ligan and Porsild 1969), which is the count commonly obtained from the Papaver radica- tum type from Alaska and the Yukon, whereas in P. Macounii the count was 2n = 28 (John- son and Packer 1968). In No. 365 the capsules were unusually large, averaging 2.2 cm in length, tapering gradually from the stigma towards the peduncle. The petals were not narrower than in typical P. alaskanum. Cruciferae Papaver Keelei Porsild, Nat. Mus. Can. Bull. 101:20 (1945); ibid., 121:185; Pl. 16, figs. 4-7 (1951) not P. Macounii sensu Hult. Fl. Alaska & Neighboring Territories. 491 (1968). Stewart Plateau: Keno Summit, elevation 6,000 ft, in moist, alpine tundra below scree, Nos. 519, 1260, and 1262. Dempster Highway: Miles 50-54, No. 214; alpine herbmats east of Miles 50-58, Nos. 384 and 474; Mile 110, on moist tundra at the base of limestone hills, flowering on 10-11 July 1970, A.E. & R.T. Por- sild, Nos. 23,626-27. | The type of P. Keelei came from the Plains of Abraham near the crest of the Mackenzie Mountains at Mile 82 east of the Canol Road, where it was common on limestone rubble at 6,000 ft elevation (A.E. Porsild & Breitung, No. 11,782). Since 1945 it has been collected a number of times in the Mackenzie and Rich- ardson mountains, as well as in the moun- tains of central and northern Yukon and Alaska, and everywhere it appears to favour calcareous soil. In this preference and in its non-caespitose habit and narrow capsule (well illustrated in the plate cited above), it differs markedly from most native poppies reported from northwestern North America, and perhaps most of.all from P. Macounii, to which Hultén (1968) refers it. A chromosome count from root tips grown from seeds of No. 519 from Keno Hill gave 2n = 70, whereas that of P. Macounii is reported to be 2n = 28 (L6ve and Freedman 1956). Papaver Walpolei Porsild, Rhodora 41:231; Pl. 552, figs. 4-10 (1939). Ogilvie Mountains: In moist, calcareous seep- age at base of limestone hill south of Demp- ster Highway, Mile 110, A.E. & R.T. Porsild, No. 23,625. New to the flora of Canada (Por- sild 1972: 236). Our specimens, past flowering on 11 June 1970, are a perfect match for this dwarf spe- cies, until now Known only from the Seward Peninsula, Alaska, where, since its discovery in 1926, it has been collected a number of times near Nome, at Teller, and is also known from East Cape, Siberia. P. Walpolei is the smallest of the North American poppies, with scapes rarely more than 15 cm tall; the leaves are totally gla- brous, shallowly 3-lobed, or subentire, and the flowers are rarely more than 2 cm in diameter. CRUCIFERAE Arabis Holboellii Hornem. var. retrofracta (Grah.) Rydb. Vicinity of Mayo, 63°35’N, 135°45’/W, gravelly river terraces, Nos. 456 and 934. Arabis lyrata L. ssp. kamchatica (Fisch.) Hult. Mayo District: Keno Hill, in moist, open grav- elly places ascending to 6,000 ft, Nos. 749, 847, and 1275. Ogilvie Mountains: Along Dempster Highway, Miles 40-47, No. 1452; Mile 56, No. 412; Mile 63, No. 44; Miles 67 and 95, Nos. 1788 and 1739. A chromosome count from No. 1275 gave 2n = 32 (Mulligan and Porsild 1969). Nos. 1739 and 1788 also gave 2n = 32 (Mulligan and Porsild 1970). Hultén (1967: 65) reported having collected Arabis arenicola (Richards.) Gelert in north- ern Alaska, “‘at Umiat, Meade R. village, Col- ville R. Delta, and at Ogotoruk Creek near Cape Thompson’, and he later listed and illustrated what he considered to be A. areni- cola (1968:544). On the strength of his dis- covery, he also accepted (and mapped) an earlier report of that species from Golofnin Bay on the south shore of Seward Peninsula, Alaska (Gray and Watson 1895: 137). How- ever, he apparently overlooked the fact that Hopkins (1937: 78), following an examination of the Golofnin Bay specimen preserved in the Gray Herbarium, had reported it to be “‘in the early flowering stage” and “to belong to some other genus of the Cruciferae, most probably to Parrya”’. | have not seen Hultén’s alleged specimens of the Alaskan “A. arenicola’’, but, judging from the illustration (Hultén 1968:544) and from specimens and photographs (obtained in 1961 at Meade River village by the late Ray- mond D. Wood when together with Hultén), the plant in question is not A. arenicola but A. lyrata ssp. kamchatica (Fisch.) Hult. Arabis arenicola is always perennial, es- sentially glabrous, its leaves toothed-entire, somewhat fleshy and shiny in life; the siliques are greenish purple, flat, 2.0-2.5 cm long and never less than 2 mm wide, and the seeds are distinctly in two rows. It is endemic in the eastern North American Arctic, where it is known from East Greeniand, west across sub- arctic Canada, to slightly beyond longitude 110°W (Porsild 1964b: map 197). It is well il- lustrated by Gelert (1898: figs. 1A:c and d), by Porsild (1964b: fig. 40a) and by Bocher, Holmen, and Jakobsen (1968: fig. 22a). 31 Plate 6 Draba barbata Pohle Approximately 1% natural size 32 Cruciferae In Arabis lyrata ssp. kamchatica at least some of the leaves are pinnatifid-lyrate, sparsely pubescent, not fleshy, or greenish blue, or shiny; the siliques are linear, round in cross-section, 2—4 cm long, but never more than 1 mm in diameter, and the seeds are always in a single row. A. lyrata var. glabra (DC.) Hopkins may superficially resemble A. arenicola, but it is normally biennial, its siliques are always nar- rower, and the seeds are always in a single row. Braya glabella Richards. Ogilvie Mountains: Moist spots on lower slope of limestone hill west of Dempster High- way, Mile 110, A.E. & R.T. Porsild, Nos. 23,629 and 23,630; same place in moist calcareous sand by brook, jidem, No. 23,628. In all these places we found plants in anthesis, with short flowering heads becoming much elongated in fruit. Our specimens are almost entirely glabrous; some almost mature siliques meas- ured 1 cm in length, with slender styles 2 mm long. New to the flora of Yukon. Braya Richardsonii (Rydb.) Fern. B. humilis (C.A. Mey.)Robins. ssp. Richard- sonii (Rydb.) Hult. Ogilvie Mountains: Calcareous gravel bars along Blackstone River, elevation 3,800 ft, Dempster Highway, Mile 83, Nos. 1514 and 1701. A chromosome count of 2n = 42 was obtained from seeds of No. 1701 (Mulligan and Porsild 1970); limestone hills near Mile 110, A.E. & R.T. Porsild, Nos. 23,631-32. Cardamine digitata Richards. Ogilvie Mountains: Wet, calcareous flats along Blackstone River near Dempster High- way, Mile 81, Nos. 1464 and 1536. A new southern limit in Yukon for this western arc- tic-alpine species. As shown by the writer (Porsild 1943: 41; 1951a: 190) and by Rollins (1952: 260), C. di- gitata Richards. is the valid name for this species, and C. hyperborea Schulz as well as C. Richardsonii Hult. are therefore super- fluous. Hultén (1967: 62) apparently accepted this view but, nevertheless, he later (1968: 516) used C. hyperborea. Cardamine microphylla Adams C. Blaisdellii Eastw. “Sam Lake’’, Old Crow Flats, Yukon, 68°30/N, 138°05’W, dry area at top of hill, elevation 2,500 ft, A.M. Pearson, No. 71-21; Mount Sedgwick, approximately 68°58/N, 139°00’W, in moist area along creek, idem, No. 70-26E. An amphi-Beringian species, shown in Hul- tén’s distribution map (1968: 516) from the north slope of Sadlerochit Mountains, and re- cently reported from the Richardson Moun- tains (Porsild and Cody 1968). In the Bering Sea area the leaflets in C. microphylla are distinctly 3-lobed; in the present collection and in Hultén’s figure (1968) they are elliptic- oval and entire-margined. New to the flora of Yukon. Cardamine purpurea Cham. & Schlecht. Ogilvie Mountains: Alpine herbmats east of Dempster Highway, Miles 50-54, No. 466; river flats along Miles 56-57, Nos. 52 and 57; calcareous tundra, elevation 4,200 ft, near Miles 81-82, Nos. 1080 and 1485; calcareous, stony floodplain tundra, near Mile 110, A.E. & R.T. Porsild, Nos. 23,634—-35. Mount Nansen, 62°06’N, 137°17’W, 40 miles west of Car- macks, in wet moss by alpine brook, No. 1364. Draba barbata Pohle sensu Busch, FI. Sib. et Orient. 25:304-05 (1913), but not Schulz in Engler, Pflanzenr. IV, 105, Heft 89:101 (1927), at least insofar as references to North Amer- ican plants. Ogilvie Mountains: Vegetated stripes on dry limestone scree west of Dempster Highway, Mile 110, elevation 4,000 ft, A.E. & R.T. Por- sild, No. 23,633 (Plate 6). New to North Amer- ica. Otherwise Known only from easternmost Siberia. Dense caespitosa. Rhizoma ramosum. Caules rosulis foliorum vetustis diu persistent- ibus obtecti. Folia dense imbricata, integer- rima, liguliformi-linearia, obtusa, rigida, cari- nata, 7-12 mm Ig. ca. 1 mm It; * margine pilis ad 2mm longis et rigidis ciliata, supra et sub- tus pilis mollioribus densissimis simplicibus et ramosis canescentia. Scapi aphylli, cum racemo 1-4 cm lg., densissime pilis simpli- cibus et ramosis hirsuti. Racemus 6-12-florus, pedunculis densissime hirsutis, floriferis 1.5— 9 mm Ig. Sepala oblongo-ovata, dense pilis * Clearly a misprint. In Flora of the USSR, vol. 8:386, the leaves of D. barbata are said to be 1.5 mm broad. 33 Annotated List of Selected Species mollibus simplicibus et ramosis, pubescentia, 1.75-2 mm lg. Petala obovata v. oblongo- obovata, obtusa v. retusa, 3.5-4.5 mm lg., 2-2.5 It. Siliculae ovatae, glabrae v. hirsutae, stylo ca. 1.25 mm Ig.” Our specimens were in fruit, but matched rather closely the above description, except in their leaves, 2.0-2.5 mm broad (see foot- note, p. 33); they are quite unlike the Rocky Mountain plant cited by Schulz, which is D. Paysonii MacBr. var. Treleasii, and do not re- semble Gelert’s illustration (1898: 294-95, fig. 4), which Schulz (1927) had intended as a representation of D. barbata. Gelert, how- ever, thought it was D. glacialis Adams var. Y Hook., but, except for the remains of a pre- vious year’s silique, Gelert’s figure 4 almost certainly shows the flowering state of D. oli- gosperma Hook. In our plant the almost ma- ture siliques are lanceolate, 5-6 mm long and 2 mm wide, which agrees with the measure- ment given for D. barbata in Flora of the USSR, vol. 8:386; the upper surface of the leaves is loosely covered by stalked, 2-4- forked hairs, and the underside is glabrous and prominently keeled. Draba borealis DC. Stewart Plateau: Keno Hill, in gravelly places near summit, elevation 5,500-6,000 ft, Nos. 979, 1032, and 1276. Ogilvie Mountains: Dempster Highway, Miles 81-83, on gravelly hummocks in river valley, No. 1600. In Yukon otherwise Known from half a dozen stations in the southern part. Hultén’s map (1968: 534) shows a type of distribution typical of oceanic amphi-Berin- gian species. Mulligan (1970: 1434), on the other hand, considered D. borealis an arc- tic-montane species, and on his map (1970: map 3) indicated that it is wide-ranging in the Canadian Rocky Mountains, and that beyond Great Bear Lake it reaches the southern islands of the Canadian Arctic Archipelago. * Quoted from Busch (1913: 304-05), the above is presumably the diagnosis originally proposed by Pohle (1914:467), and is at variance at a great many points with that given by Schulz (1927). Schulz had seen no Siberian material and had identified D. barbata with the cordilleran plant now recognized as D. Paysonii Macbr., which does not even resemble D. barbata Pohle. The illustration in Hultén (1968: 529) closely matches flowering speci- mens of D. barbata in the Leningrad Herbarium. 34 Draba cinerea Adams Ogilvie Mountains: On dry limestone ledges west of Mile 82 on Dempster Highway, eleva- tion 4,200 ft, No. 1082. Also on Mount Gray northeast of Whitehorse, on limestone cliffs, elevation 5,000 ft, No. 1353A. Draba crassifolia Grah. Keno Hill, 63°58’N, 135°42’W, in herbmat slopes between 4,500 and 5,500 ft elevation, No. 953. Ogilvie Mountains: River flats along Dempster Highway, Miles 65-66, in damp sand, No. 72; mountain east of Miles 50-54, elevation 4,800-—5,000 ft, Nos. 167A and 476; Mile 47, No. 1852. In Yukon otherwise known from a single station in the Pelly Mountains (Porsild 1951a), and from mountains in south- western parts. Draba exalata El. Ekm., Svensk Bot. Tidskr. 25 (4):489; PI. 5, fig. 3 (1931). Mount Gray northeast of Whitehorse, on lime- stone cliffs near summit, elevation 5,000 ft, No. 1353A. The two specimens in the collec- tion are young plants, flowering for the first time, each with a single, rooted rosette. A very rare and little-collected species, long Known only from the type locality near Nome, Alaska, with caespitose, long-mar- cescent, and crowded basal leaves, softly stellate-pubescent on both sides, the under- side with a prominent midrib; scapes short- pubescent, 5-7 cm tall; petals pale yellow, siliques ovate, 4-5 mm long, thinly pubescent, style 0.5 mm long. Draba exalata is “new” to the Yukon, but was reported by Cody and Porsild (1968: 268) from several stations in the southern Mac- kenzie Mountains, Northwest Territories. In- cidentally, Hultén’s illustration (1968: 524) purporting to represent D. exalata, does not at all resemble the type illustrated by Ekman (1931: Pl. 5, fig. 3). Draba fladnizensis Wulf. Stewart Plateau: Keno Summit, in open gravel patches above continuous vegetation, eleva- tion 6,000 ft, No. 1069; same place on rocky ledges, No. 708. Ogilvie Mountains: On rocky ledges along Dempster Highway, Mile 86, Nos. 1645-46. These collections are the first from Yukon Territory. Draba glabella Pursh Ogilvie Mountains: River flats along Dempster Highway, Mile 57, Nos. 227 and 228; rocky Cruciferae ledges in canyon near Mile 86, No. 1644; on herbmat slopes near Mile 84, No. 1677. In all collections the siliques are very narrow, aver- aging from 2.0—2.5 mm wide, broadest below the middle, and from 10-18 mm long, on peduncles one-third to one-half the length of the silique: the valves are glabrous or finely hirtellous, and the style is slender and only 0.5 mm long. Draba lactea Adams Ogilvie Mountains: In alpine herbmats on mountain east of Miles 50-54 on Dempster Highway, No. 470. Apparently rare in central Yukon (Porsild 1966).. Draba lanceolata Royle Keno Hill, 63°58’N, 135°42’W, on dry rocky southwestern slope, No. 739. Ogilvie Moun- tains: On gravelly river flats along Dempster Highway, Miles 57-58, Nos. 288 and 427; on dry, calcareous river flats, Mile 81, elevation 4,200 ft, No. 1490. Also from Mount Gray northeast of Whitehorse, on limestone cliffs, No. 1331. Draba longipes Raup Keno Hill: Very common on rather moist herb- mat slopes and occasional on moist gravel near the summit, elevation 6,065 ft, Nos. 528, 705-07, 743, 974, 1022, 1070, and 1277-78; flowering from early July. Ogilvie Mountains: Alpine herbmats on mountain east of Demp- ster Highway, Miles 57-58, Nos. 401-02 and 494; Miles 44-46, Nos. 1441-42; Mile 83, No. 1515. Old Crow Flats, 68°N, 138°W, elevation 2,500 ft, A.M. Pearson, No. 70-21. As predicted (Porsild 1966:33) Draba long- ipes has now become recognized as one of the most common members of the genus in moist, alpine herbmats throughout Yukon. Draba Macounii Schulz Ogilvie Mountains: Very sparse at 5,800—6,500 ft on wet, gravelly northern slope of mountain east of Dempster Highway, Miles 51 and 54, Nos. 471 and 1579. The latter collection con- sists of 6 specimens, each with a short, com- pact branched caudex and a stout taproot; the leaves are oblong, about 5-7 mm long, loosely imbricated, nerveless, long-ciliate and glabrous on both surfaces; the compact flowering heads on 7 July were barely raised above the leaf-cushion; remains of last year’s fruiting scapes, 5-6 mm long; sepals oval, glabrous or with a few simple hairs; petals pale yellow, obovate, the young siliques gla- brous with a short, coronate style. No. 471 was collected on 15 Aug. 1966, which was obviously a more favourable season, because the scapes in that collection are 1.5-—2.0 cm long, each with 3-5 lanceolate siliques, 5-7 mm long. Both collections closely match the follow- ing: the type of D. Macounii from Kicking Horse Lake, Rocky Mountains, Alberta, J. Macoun, 20 July 1885, CAN No. 61,572; a large series of D. Macounii in my collection from the Canadian Rockies; four collections from the vicinity of Summit Pass, British Columbia, 58°31’N, 124°34’W, H.M. Raup & D.S. Correll, Nos. 10,660, 10,684, 10,688 and 10,692, dis- tributed as Draba macrocarpa Adams; and H.M. Raup & E.C. Abbe, No. 3970 from Mount Selwyn, British Columbia, 56°01’N, 123°39’W, distributed as D. alpina. Draba Macounii is new to the flora of Yukon, but was recently reported from mountains of the Mackenzie District (Cody and Porsild 1968). Draba nivalis Liljebl. Common in the Ogilvie Mountains in alpine tundra and on gravelly river terraces along Dempster Highway, Miles 44-69, Nos. 40, 42, 382, 464, 1443, 1577, and 1598. Draba oligosperma Hook. Mount Gray northeast of Whitehorse on lime- stone cliffs and in alpine tundra, elevation 3,500-5,000 ft, No. 1336. This number is a single but exceptionally large specimen, measuring about 40 cm in diameter, rising from a central taproot, the crown of which is repeatedly divided into hundreds of densely massed flowering and fruiting branches that average 15 cm in length, each bearing from 5-10 ovate siliques, on peduncles 2 to 3 times the length of the silique. Draba oligosperma is now known from a number of stations in southern Yukon, all on calcareous rocks. For its general range, see Porsild 1966: map 79. Draba Palanderiana Kjellm. D. caesia sensu Hult., Fl. Alaska & Yukon 5:849 (1945) non Adams (see Porsild 1966:35, map 80). Stewart Plateau: Mount Haldane, rocky ledges on the north face, elevation 4,000 ft, No. 562; on south face, on open gravel-slide, elevation 5,500 ft, No. 669. Ogilvie Mountains: 35 Plate 7 Draba yukonensis n. sp. (Type) Approximately natural size 36 Cruciferae On rock exposure on mountain east of Demp- ster Highway, Miles 50-54, elevation 5,000 ft, No. 145; in herbmats on north side of moun- tain ridge east of Miles 57-58, No. 382; Mount Nansen, 62°06’N, 137°17’W, common on dry ridges, No. 1369. Also on Mount Sedgwick, 68°58’N, 139°01’W, A.M. Pearson, No. 70-22B. In life the petals of D. Palanderiana are creamy white at first, and only the claws are yellow, but as the season advances, the petals turn a deeper yellow that increases in density when drying, whereas in the morpho- logically similar D. nivalis the petals are milky white and do not darken when drying. In D. Palanderiana ‘the siliques are lanceo- late and about twice as long as broad; in D. nivalis they are narrower and 4-5 times longer than broad. Draba sibirica L. D. ogilviensis Hult., Bot. Notis. 119 (2):315 (1966). Ogilvie Mountains: Dempster Highway, Miles 40-46, No. 1460; mountain east of Mile 50, Nos. 9, 62, 141, 167, 463, 1460, and 1580. Very common in herbmats and in moist, open wil- low thickets of the valley, ascending to 6,500 ft; flowering and fruiting abundantly; often associated with D. longipes. The first report of D. sibirica from North America (Porsild 1964a) was based on flower- ing fragments found among other plants regu- larly “harvested” by the collared lemming in the Ogilvie Mountains, and were brought to me for identification in 1963 by Messrs. P.M. Youngman and G. Tessier of the National Museum of Canada. With an abundant and well-preserved series of the Yukon plant be- fore me, as well as with the large series re- ported from the central Mackenzie Moun- tains, Northwest Territories (Cody and Por- sild 1968), | compared these with other speci- mens of D. sibirica representative of its pecul- iarly discontinuous range from central East Greenland, the Ural Mountains, Waigatsch Island, and the adjacent mainland of arctic Russia, including the west Caucasus Moun- tains, the lower Lena River of eastern Siberia, the Chukotsk Peninsula, and East Cape (Por- sild 1967). The variations observed proved no greater than those noted in D. sibirica ob- tained at different elevations and exposures between elevation 3,800-—6,500 ft on a single mountain in the Ogilvie Mountains. * D. ogil- viensis Hult., proposed for the Yukon plant, is therefore superfluous and must be rejected. Draba yukonensis n. sp. Herba parva hibernanti-biennis. Caules fili- formes, simplices vel ramosi, fructifer usque ad 18-20 cm alti, 1-3-phylli, pilis patentibus plerisque floccoso-stellatis vestitis. Folia ba- salia rosulata, lanceolata, acuta, integra, 8-10 mm longa, nervo medio ad basin manifeste, utrinque pilis rigidis, simplices vel rarius bi- vel trifurcatis vestitis; folia caulinaria 1-3, ovata, 5-8 mm longa, sessilia. Racemus flo- riger non vidi, fructifer jam initio laxiusculus, dein valde elongatus et laxus, 5-20-florus. Pedicelli 1-3 mm longi. Siliquae teretiusculae, erectopatentes, 3-5 mm longae, 2 mm latae, nunquam compressae. Valvae cum pilis bre- vissimis furcatis sive stellatis hirtae. Stylus 0.3-0.4 mm longus coronatus. Semina ellip- soidea brunnea, 0.8 mm longa. Southwestern Yukon: Vicinity of Mackin- tosh, Mile 1022 of the Alaska ‘Highway, in dry area among grasses, 29 June 1957, W.B. Schofield and H.A. Crum, fruiting specimens, No. 7319 (Type: CAN), Plate 7; approximately in the same place (approximate latitude 60° 47'N, and longitude 137°38’W) on open stony ridges on ancient beach, H.M. and L.G. Raup, No. 11,938, 26 June 1944. By its biennial habit, by its tiny basal ro- settes of leaves covered on both sides by pre- dominantly simple, relatively long and rather stiff hairs, by its erect, slender, simple or branched, elongating fruiting stems, bearing 1-3 tiny, sessile leaves, by its tiny, short-pedi- celled, terete siliques covered by short, soft stellate hairs, and by its very short style, Draba yukonensis differs markedly from Draba praealta Greene, as well as from other members of section Leucodraba. |n both col- lections cited, mature seeds are present; at the base of some of the lowermost siliques, wilted but distinctly white petals still adhere. Erysimum Pallassii (Pursh) Fern. Ogilvie Mountains: Dempster Highway near Miles 44—46, elevation 4,200 ft, on gravel ter- races above floodplain, No. 1439; Mile 55, No. 55, with almost mature fruits on 25 June 1966, n= 14 (Mulligan and Porsild 1969). A similar conclusion was reached by Tolmat- chev (1939: 446-47). 37 Annotated List of Selected Species Eutrema Edwardsii R. Br. Ogilvie Mountains: Mountain east of Demp- ster Highway, Mile 58, elevation 5,800 ft, No. 251. Stewart Plateau: Keno Summit, elevation 5,500-6,000 ft, Nos. 534, 775, 1021, and 1265. The specimens are all unusually robust, and in No. 1265 one last year’s fruiting stem has two lateral branches; this is unique for this taxon, which normally has one simple floral axis, as confirmed by the examination of the 180-odd sheets of Eutrema Edwardsii in the National Herbarium of Canada. Lesquerella Calderi Mulligan & Porsild, Can. J. Bot. 47(1):215-16, pl. 1 (1969). Ogilvie Mountains: Slopes of limestone hills west of Dempster Highway, Mile 82, elevation 3,800 ft, No. 1095; Miles 83-85, Nos. 1509, 1556, and 1609; rocky slopes of limestone hills southwest of Dempster Highway, Mile 110, A.E. & R.T. Porsild, Nos. 23,637-38, flowering and fruiting specimens on 10-11 July 1970; same place, on moist, calcareous floodplain-tundra, jidem, No. 23,639. The type of L. Calderi came from the Ogilvie Moun- tains, 66°02’/N, 138°44’W, Calder & Gillet, No. 26,016 (DAO). L. Calderi differs from the rather similar L. arctica by consistently longer and broader petals and larger siliques; L. arctica is dodecaploid, whereas L. Calderi is tetraploid, both with the base number X =5 (Mulligan and Porsild 1970). Rorippa barbareaefolia (DC.) Kitag. Ogilvie Mountains: Dempster Highway, Miles 96-98, in rich meadows along the Blackstone River, No. 1722, with mature seeds on 30 July 1968, 2n=16 (Mulligan and Porsild 1969). An amphi-Beringian species of spotty and rather local range, suggesting distribution by man (Porsild 1939:233). Until now, known in Canada only from the Dawson area and main- ly in places disturbed by placer mining. R. barbareaefolia differs from the somewhat similar R. palustris by its pear-shaped sili- ques, opening by four rather than two valves. Smelowskia borealis (Greene) Drury & Rollins Ogilvie Mountains: Very sparse and local in slide-rock on the face of limestone mountain southwest of Dempster Highway near Mile 110; the specimen consists of this year’s ro- settes and last season’s elongated inflores- cence, A.E. & R.T. Porsild, No. 23,636. The above station is approximately 100 miles 38 southeast of the type locality at Runt Creek on the Alaska—Yukon boundary, latitude 66° 10’N. Smelowskia calycina (Steph.) C.A. Mey. var. media Rollins & Drury Mount Sedgwick, approximately 68°58/N, 139°01’W, on dry slope at top of ridge, A.M. Pearson, No. 70-27D. Until now, this variety was known only from the east end of the Brooks Range, Alaska, and from the east slope of the Richardson and Mackenzie mountains, but it had not been reported from Yukon Territory. Smelowskia calycina (Steph.) C.A. Mey. var. Porsildii Drury & Rollins, Rhodora 54:105 (1952). S. Porsildii (Drury & Rollins) Yurtsev, Not. Syst. Rast. 6:309 (1970). Ogilvie Mountains: On slide-rock on mountain east of Dempster Highway, Miles 57-58, ele- vation 6,000 ft, Nos. 255 and 383. The var. Porsildii is distinguished from other varieties of S. calycina by the blade of its basal leaves being entire or sometimes with 2-3 shallow apical notches, linear to narrowly spatulate in outline and shorter than its slender petiole. Thus far, the variety is known only from a few stations in the Brooks Range of northern Alaska and from the type locality in the Kok- rines Hills, 65°20’N, 154°30/W, A.E. & R.T. Porsild, No. 741. The Ogilvie stations listed above are the first for Canada. DROSERACEAE Drosera anglica Huds. Stewart Plateau: Halfway Lakes north of Mayo, in bogs, Nos. 500, 641, 863, 1219, and 1254; McQuesten River valley west of Elsa, No. 1165. Common locally in wet bogs near Mayo. In Yukon otherwise known from a sin- gle collection from the base of Mount Shel- don in eastern Yukon (Porsild 1951a). Drosera rotundifolia L. Stewart Plateau: Vicinity of Halfway Lakes north of Mayo, in a bog, No. 864; near Minto Bridge, on floating quagmire, No. 541. In Yu- kon otherwise known from two stations along the Canol Road and from the Dawson district (Porsild 1951a). Saxifragaceae SAXIFRAGACEAE Boykinia Richardsonii (Hook.) Gray Saxifraga Richardsonii Hook., Fl. Bor.-Am. 1:247 (1832). Mount Sedgwick, 68°58/N, 139°01’W, in moist places by alpine brook, A.M. Pearson, No. 70-23. Hooker (1832:247), on Richardson’s au- thority, gave the range of his new and strik- ingly handsome species as “Arctic Sea- shore between Mackenzie and Coppermine Rivers’. With our present knowledge of the range of this very conspicuous and distinctly alpine species, the above statement certainly is puzzling. At any rate, | and all subsequent collectors have looked in vain for Boykinia Richardsonii along the Arctic Coast as well as inland, east of the Mackenzie River. | can only assume that Richardson’s statement, as quoted by Hooker, is in error, and that the specimens seen by Hooker were actually col- lected on the Arctic Coast of Yukon or north- ern Alaska by members of the Franklin Ex- peditions Western party travelling by boat west to Point Barrow, and that the specimens later got mixed up with Richardson’s plant collections from eas? of the Mackenzie River. Boykinia Richardsonii, endemic in Alaska and northwestern Yukon, is apparently not uncommon in mountains of northern, central, and western Alaska, where it is restricted to alpine herbmats with ample snow cover in winter. It has been collected a few times in mountains of northern and central Yukon, and it will likely also turn up on the east slope of the Richardson Mountains. Chrysosplenium Wrightii Franch. & Sav. Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 50-54, in shaded ravine between rocks, elevation 5,500-—5,800 ft, Nos. 148, 469, and 1568. In Yukon otherwise known from Kluane Lake and Donjek River. A chro- mosome count of 2n= 24 was obtained from No. 469 (Mulligan and Porsild 1969); also re- ported recently from a number of stations on the east slope of the Richardson Mountains, Mackenzie District (Cody and Porsild 1968). Ribes glandulosum Grauer Stewart Plateau: Minto Bridge, edge of spruce woods, with mature fruits on 17 July 1967, No. 875; vicinity of Halfway Lakes north of Mayo, No. 1217, the latter with sprawling stems freely rooting from the nodes. An east- ern boreal forest species, in Yukon otherwise known from a few stations along the Canol Road (Porsild 1951a). Ribes hudsonianum Richards. Vicinity of Mayo, 63°35’N, 135°55’W, wooded riverbanks, No. 472. Ogilvie Mountains: In willow thickets west of Dempster Highway, Mile 95, No. 1740. The last is a slight exten- sion of the known northern limit for Yukon. Ribes triste Pall. Ogilvie Mountains: Base of rock scree, Demp- ster Highway, Mile 86, No. 1638. An extension of the Known northern limit for Yukon. Saxifraga adscendens L. ssp. oregonensis (Raf.) Bacigalupi Ogilvie Mountains: River flats along Demp- ster Highway, Mile 57, on wet gravel, No. 220, n=11 (Mulligan and Porsild 1969). See also Porsild 1966: map 87. Saxifraga aizoides L. Ogilvie Mountains: Calcareous river flats along Dempster Highway, Miles 81-85, Nos. 1555 and 1601, the latter with flowers and young fruits on 18 July 1968; in calcareous gravel by cold brook west of Mile 110, A.E. & R.T. Porsild, No. 23,642. A pronounced calci- cole. The collections represent a slight ex- tension of the known western limit of this amphi-Atlantic species, only recently report- ed from Yukon (Porsild 1951 a). Saxifraga caespitosa L. ssp. monticola (Small) Porsild Ogilvie Mountains: Rocky ledges, elevation 5,500-5,800 ft, on mountain east of Demp- ster Highway, Miles 50-54, Nos. 182, 472, and 1570; mountain east of Miles 57-58, No. 410; limestone hills west of Mile 82, elevation 5,200 ft, No. 1094. A well-marked eastern cordilleran race that extends, mainly on calcareous moun- tains, from the Mackenzie and Richardson mountains to southwestern and central Yu- kon. West of the 141st meridian known froma single collection from ‘‘Gunsight Mountain”, Mile 120, on Glenn Highway, approximately 61°51’N, 147°31’/W, L. Spetzman, No. 4373. See also Porsild 1966:39-—40; pl. 6B; map 88, on which the stations cited above should be shown. 39 Annotated List of Selected Species Saxifraga davurica Willd. ssp. grandipetala (Engler & Irmsch.) Hult. Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 50-58, elevation 5,500-— 6,000 ft, Nos. 184, 385, and 467; river flats along Miles 51-52, No. 205; damp, rocky places on mountain east of Mile 51, No. 1573; Stewart Piateau, Keno Summit, in damp al- pine tundra, elevation 6,000 ft, Nos. 513, 725, and 1261 (Porsild 1966: map 89). Saxifraga flagellaris Willd. ssp. flagellaris Ogilvie Mountains: River flat along Dempster Highway, Miles 51-52, Nos. 122 and 212; mountain east of Miles 50-54, elevation 6,000 ft, No. 25 (Porsild 1954, 1966). Saxifraga hieracifolia Waldst. & Kit. Ogilvie Mountains: Moist alpine tundra and herbmats on mountain east of Dempster High- way, Miles 50-54, Nos. 27, 89, and 480; on gravel ridge along river near Miles 57-58, No. 300; Mile 67, No. 1807; in damp river meadows near Miles 82-86, Nos. 1118 and 1657, the latter with stout flowering scapes 6 dm tall. Stewart Plateau: Keno Hill, 63°58’N, 135°40’W, alpine tundra, elevation about 6,000 ft, Nos. 518 and 996. Common in turfy places of alpine tundra. Saxifraga Hirculus L. Ogilvie Mountains: Common in calcareous tundra and herbmats, along Dempster High- way, Miles 51-58, Nos. 132, 213, and 488; Mile 67, in mossy bog on river flat, No. 1803; Mile 83, by seepage in open woods, No. 1613; Mile 110, wet, calcareous bog, A.E. & R.T. Por- sild, No. 23,642. Stewart Plateau: Keno Hill, in herbmats near summit, elevation 5,500-— 6,000 ft, Nos. 533, 709, and 956. The speci- mens are best placed with the arctic and cir- cumpolar var. propinqua (R. Br.) Simm., al- though in some the petals are up to 15 mm long; in No. 1613 the slender flowering stems are over 2 dm tall, with 7-9 sessile cauline leaves. Very common in damp, mossy bogs and in damp, alpine herbmats. Saxifraga punctata L. ssp. Nelsoniana (D. Don) Hult. Ogilvie Mountains: In wet gravel by alpine seepage, along Dempster Highway, Miles 44— 46, elevation 4,200 ft, No. 1446; mountain east of Miles 50-54, in alpine herbmats, elevation 3,800 ft, No. 95; same mountain, elevation 5,000 ft, No. 28; Mile 86, on rocky ledges of a 40 canyon, No. 1636. A chromosome count of 2n = ca. 84 was obtained from S. punctata L., No. 95 (Mulligan and Porsild 1969). The range of ssp. Nelsoniana is amphi- Beringian, extending along the North Slope of Alaska and Yukon, east only to the eastern slope of the Richardson Mountains, west of the Mackenzie Delta and south across north- ern Yukon to the Ogilvie Mountains, where it meets and slightly overlaps that of ssp. Por- sildii. Subspecies Porsildii is apparently an inland or continental race, consisting of two discrete populations of which the western is confined to the Mackenzie Mountains and the Stewart Plateau of central Yukon, and the eastern to the northeastern Mackenzie and western Keewatin districts. The two popula- tions are separated by a forested gap nearly 1,000 miles wide. Saxifraga punctata L. ssp. Porsildii Calder & Savile Mount Haldane, 63°51’N, 135°50’W, elevation 4,300 ft, in gravel by a creek, No. 588; in mossy openings under spruce, elevation 3,500 ft, No. 622; Keno Hill, 63°58’N, 135°42’W, wet spots in rich herbmat slope, elevation 4,500—5,500 ft, Nos. 745 and 965; on rock ledges, elevation 5,500-6,000 ft, No. 1225. Ogilvie Mountains: Wet rocky ledges in canyon along Dempster Highway, Mile 47, No. 1850; on snowbed slope, No. 1853; on river flats along Dempster Highway, Mile 56, No. 192. The ssp. Porsildii is distinguished by its lower stature, essentially glabrous leaves, purplish peduncles, sepals, and follicles, and by its terminal flower often, if not always, 3- or even 5-follicled. Its range, as shown above, is remarkably disjunct from that of ssp. Nel- soniana. Saxifraga radiata Small Stewart Plateau: Mount Haldane, 63°51’N, 135°50’W, elevation 4,300 ft, in wet moss by alpine creek, No. 602; Keno Hill, 63°58’N, 135°42’W, herbmat slopes and ledges be- tween elevation 4,500 and 6,000 ft, Nos. 744, 1025, 1224, and 1280. A chromosome count of n = 13 was obtained for No. 1224 (Mulligan and Porsild 1969). Ogilvie Mountains: river flats and moist rocky ledges along Dempster Highway, Miles 44-63, Nos. 43, 191, 215, 1438, and 1843. Apparently quite common locally in moist alpine herbmats from the central Yukon Plateau northwards to the Arctic Coast. Saxifragaceae Saxifraga reflexa Hook. Stewart Plateau: Mount Haldane, 63°51’N, 135°50’W, on gravelly slope, elevation 5,500 ft, No. 582; Keno Hill, 63°58’N, 135°46’W, on rocky ledge near summit, elevation 5,500 ft, No. 1269. Ogilvie Mountains: Dempster High- way, rocky slopes of mountain east of Miles 50-54, Nos. 20 and 93; mountain ridge east of Miles 57-58, No. 388; river flat near Mile 67, No. 1804; rocky ledges near Mile 86, No. 1647. Not uncommon on dry alpine slopes and river flats. | Saxifraga rivularis L. var. flexuosa Engler & Irmsch. . Stewart Plateau: Keno Hill, common above timberline in wet, gravelly places and on rocky ledges to elevation 6,065 ft, Nos. 529, 766, 1257, and 1258. A chromosome count from Nos. 1257 and 1258 gave n = 13 (Mul- ligan and Porsild 1969). Ogilvie Mountains: damp, rocky ledges near Mile 47 on Demp- ster Highway, No. 1849, in wet gravel by cold spring, near Mile 58, No. 496. As noted by Hultén (1944), typical S. rivu- laris in Alaska and Yukon are largely re- stricted to the Arctic Coast, whereas in the interior most S. rivuleris belong with the taller and more slender var. flexuosa, in which the flowers are smaller, the lowermost on long and strictly erect peduncles. Saxifraga rufopilosa (Hult.) n. stat. S. nivalis var. rufopilosa Hult., Ark. Bot. ser. 2, vol. 7 (1): 69 (1967). S. (?) eriophora Porsild, Can. Field-Natur. 79 (2): 82 (1965) non S. eriophora S. Wats. An examination of the type of S. nivalis var. rufopilosa Hult. found on Steese Highway, mountain west of Mastodon Dome (central Alaska, approximately 65°35’N, 145°00’W), 25-26 July 1964, E. Hultén (S), and the large series listed below, but not seen by Hultén, suggests that this taxon, although undoubted- ly distinct, is in several characters more closely related to S. tenuis (Wahlenb.) Sm. than to S. nivalis L. In the flowering state S. rufopilosa super- ficially resembles S. tenuis by its subcapitate inflorescence, but as the fruits mature, its flowering axes and their branches elongate, so that the inflorescence becomes racemose- paniculate (see Plate 8), and from 4 to 6 cm long, but never corymbose as in S. tenuis, S. rufidula J.M. Macoun, or S. eriophora S. Wats. In all parts S. rufopilosa is a smaller and more delicate plant than S. nivalis in which, moreover, the petals are always white and the mature capsules average 6 mm in length; whereas in S. rufopilosa the petals are pink or pale purplish, and the capsules about 3 mm long. In S. rufopilosa as well as in S. tenuis, the underside of the leaves is loosely matted by long, crinkly, rust-coloured hairs, whereas in S. nivalis some rust-coloured hairs may be present on the upper as well as the lower leaf-surface. S. rufopilosa appears to be endemic in un- glaciated alpine parts of central Alaska and southern and central Yukon Territory. Alaska: Eagle Creek, near Eagle Summit on the Steese Highway, Mile 105, 65°22’/N, 145°20’W, elevation 3,500-4,000 ft, on north- facing rock stripes and talus, 30 July and 7 Aug. 1963, Robert W. Weeden, fruiting spec- imens, No. 283,877 (Plate 9); same place, 28 June 1966, idem, No. 303,792; Canada: Mountain slopes northeast of Ptarmigan Heart, approximately 61°49’N, 138°35’W, 16 July 1948, H.M. Raup, W.H. Drury, and K.A. Raup, No. 13,740; St. Elias Mountains, Bear Creek near Mount Archibald, Mile 1028, Alas- ka Highway, elevation 4,150 ft, A.M. Pearson, No. 1967-297A; Dawson Range, 62°35/N, 138°20’W, 27 June 1941, Hugh Bostock, No. 256. The numbers that follow were collected by R.T. Porsild. Mount Nansen, 40 miles west of Carmacks, 62°06’N, 137°17’W, on moist spots in alpine tundra, 16-17 June 1968, No. 1372. Ogilvie Mountains: “Sheep Mountain’ east of Dempster Highway, Miles 50-54, eleva- tion 4,500-—5,000 ft, in alpine herbmats, No. 32, Plate 8, flowering specimens on 24 June 1966; same place, in damp, gravelly places near summit, elevation 6,500 ft, with flowers and young fruits on 7 July 1968, No. 1574. Saxifraga serphyllifolia Pursh Stewart Plateau: Keno Summit, in half shade on mossy rock ledges, elevation 6,000 ft, Nos. 527, 773, and 1076. Ogilvie Mountains: Sum- mit of mountain east of Miles 50-54, on Demp- ster Highway, elevation 6,500 ft, Nos. 176, 461, and 1575; mountain east of Miles 57-58, No. 381 (Porsild 1966: map 91). 41 Plate 8 Saxifraga rufopilosa (Hult.) n. stat. Flowering specimens. Approximately 1% natural size 42 Rosaceae Plate 9 Saxifraga rufopilosa (Hult.) n. stat. Fruiting specimen Approximately 2 natural size ROSACEAE Amelanchier alnifolia (Nutt.) Nutt. Vicinity of Mayo, 63°35’N, 135°45’W, where it appears to be restricted to river terraces; No. 462, flowering on 7 June 1967, and No. 1178 with fully ripened fruits on 10 Aug. Through- out its Yukon—Alaska range, restricted to the main river systems. Chamaerhodos erecta (L.) Bunge ssp. Nuttallii (T. & G.) Hult. Stewart Plateau: Vicinity of Mayo, on gravelly river terrace, Nos. 464 and 553; flowering specimens 28 cm tall on 7 and 22 June 1967, fruiting on 7 Aug. 1967, No. 1307, 2n = 14 (Mulligan and Porsild 1969). A western plains and foothill species, reaching southwestern and central Yukon. The present collection ex- tends its known northward range by about 100 miles. Dryas alaskensis Porsild * On mountain east of Miles 57-58 on Dempster Highway, ascending to 6,000 ft, where still flowering on 31 July 1966, No. 393; Mile 110, stony floodplain below limestone hill, A.E. & R.T. Porsild, No. 23,649; Richardson Moun- tains, 17 miles northeast of Lapierre House, approximately 67°43’N, 136°20’W, Youngman & Tessier, No. 595. The last is a new northern limit for this species in Yukon. Dryas alaskensis is perhaps the most com- mon member of the genus in central Yukon, particularly abundant on gravelly floodplains and slopes rich in lime. Dryas crenulata Juz. Mount Gray northeast of Whitehorse, com- mon on limestone cliffs and in alpine tundra between 3,500 and 5,000 ft elevation, Nos. 1334 and 1349. Ogilvie Mountains: In rich herbmat slopes along Dempster Highway, Mile 84, No. 1679, with fully ripened achenes on 26 July 1968, 2n = 18 (Mulligan and Por- sild 1970). See also Porsild 1966. Dryas integrifolia M. Vahl Ogilvie Mountains: Limestone hills west of Dempster Highway, Mile 110, elevation 2,500 ft, A.E. & R.T. Porsild, No. 23,645. Apparently rare in the interior of Yukon and central Alas- ka, where it is restricted to alpine situations. Dryas octopetala L. s. str. Mount Nansen, 62°06/N, 137°17’W, in alpine lichen heath, No. 1366; summit of Coldspring Mountain, 63°02’N, 137°13’W, J.D. Campbell, No. 125. See Porsild 1966 for stations in south- western Yukon. Dryas punctata Juz. Ogilvie Mountains: Limestone mountain west of Dempster Highway, Mile 110, A.E. & R.T. Porsild, No. 23,648; Bonnet Plume Lake, 64°20/N, 132°00’W, W.H. Butler & N. Olsen, No. 57. The Ogilvie station bridges a gap in the known Yukon range of this taxon between southeastern Yukon and the Arctic Coast. * For a monographic treatment of the North American members of the genus Dryas, see Porsild 1947. Hultén’s somewhat different treatment (1968: 629) is influenced by ‘‘the fact that hybrid swarms seemingly are formed between all taxa—with the exception of D. Drummondii, which is distinct.” 43 Plate 10 Potentilla Ledebouriana Porsild Approximately % natural size ae Rosaceae Dryas sylvatica (Hult.) Porsild Ogilvie Mountains: River flats along Dempster Highway, Mile 56, No. 54; in alpine tundra, elevation 5,000 ft, Miles 54-55, No. 1830; a dominant species on sparsely wooded lime- stone gravel-terraces near Mile 83, No. 1517; limestone hills west of Mile 110, A.E. & R.T. Porsild, Nos. 23,646-47; McQuesten area, lower forks of Vancouver Creek, J.D. Camp- bell, No. 590; Parker Creek, idem, No. 782. Geum glaciale Adams Mount Sedgwick, approximately 68°58’N, 139°01’/W, in moist area along creek, flower- ing 5 July 1970, A:M. Pearson, No. 70-26B. The collection bridges a gap between the Richardson Mountains west of the Macken- zie River and the eastern Brooks Range of Alaska. Geum macrophyllum Willd. ssp. perincisum (Rydb.) Hult. Vicinity of Mayo, 63°35’N, 135°45’W, on grav- elly river terrace, No. 688; in open woods along river, No. 789. Ogilvie Mountains: Dempster Highway, Miles 96-98, in rich flood- plain meadow, No. 1721. The last station is a new northern limit for this taxon in Yukon. Geum Rossii (R. Br.) Ser. Stewart Plateau: Mount Haldane, rocky slope, elevation 4,000 ft, flowering on 23 June 1967, No. 566. Keno Hill: in alpine herbmats, eleva- tion 5,000 ft, Nos. 535, 994, and 1259; and in open heath on summit, elevation 6,000 ft, No. 1259, 2n = 70 (Mulligan and Porsild 1969). Ogilvie Mountains: River flats along Dempster Highway, Mile 52, flowering specimens 45 cm tall on 25 June, with last year’s fruiting axis, No. 58; mountains east of Miles 57-58, eleva- tion 6,000 ft, No. 403. “Sam Lake” in the British Mountains, approximately 68°30’N, 138°05’W, elevation 2,500 ft, A.M. Pearson, No. 70-21. The last is the northernmost sta- tion known for this species in Yukon. See Porsild 1966 for general distribution. Potentilla biflora Willd. Ogilvie Mountains: River flats along Demp- ster Highway, Mile 57, elevation 2,500 ft, No. 350; mountain east of Miles 50-54, elevation 5,500-6,500 ft, Nos. 142, 185, 468, and 1838 (Porsild 1966: map 95); Mount Sedgwick, ap- proximately 68°58’/N, 139°01’W, flowering on 5 July 1970, A.M. Pearson, No. 70-25A. Potentilla hyparctica Malte Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 57-58, on shady cliffs of north-facing slopes, No. 394; alpine rock ledges of mountain east of Mile 51, No. 1572 (Porsild 1966). Potentilla Ledebouriana Porsild P. Vahliana sensu Hult., Fl. Alaska & Neigh- boring Territories. 612 (1968) non Lehm. Stewart Plateau: Keno Summit, 63°58’N, 135°42’W, gravelly ledge, elevation 6,000 ft, No. 718; Mount Haldane, 63°51’N, 135°50’W, rock scree, elevation 4,000 ft, No. 567; Gale- na Hill, 63°53’N, 135°24’W, on rocky ledge, No. 1212. Ogilvie Mountains: Mountain east of Miles 50-54 on Dempster Highway, dry rocky ledges, elevation 5,000 ft, No. 144; river terraces near Mile 57, elevation 2,400 ft, No. 139; gravelly ridges near Mile 74, No. 1821, 2n = 28 (Mulligan and Porsild 1970). The plants listed above are typical of the area under consideration. More condensed plants were found in ex- posed situations. Stewart Plateau: Keno Sum- mit, elevation 6,000 ft, exposed rocky places, No. 1266, 2n= 28 (Mulligan and Porsild 1969); on gravel scree, elevation 5,500 ft, No. 730; Mount Haldane, alpine gravel slope, elevation 5,500 ft, No. 682. Ogilvie Mountains: Demp- ster Highway, mountain east of Miles 50-54, elevation 5,000 ft, No. 38; limestone hills west of Mile 82, elevation 5,200 ft, on rock tower, No. 1093 (Plate 10). In my work (Porsild 1955: 152-55, pl. 21) | have endeavoured to show that true P. Vah- liana Lehm., described from West Greenland, are endemic in West Greenland, the Cana- dian Arctic Archipelago, the Arctic Coast westward to longitude 130°W, the northern Hudson Bay region, and northern Keewatin, in contrast to P. Ledebouriana, which is a western species, distributed from eastern Siberia along mountain ranges in Alaska and the Yukon eastward to the Richardson Moun- tains west of the Mackenzie Delta, and south in the Rocky Mountains to Oregon and Colo- rado. | further noted that in exposed and high-alpine situations in the Yukon and Alaska, dwarfed and condensed forms of P. Ledebouriana undeniably may resemble P. Vahliana. However, on phytogeographical grounds alone, it seems very doubtful that true P. Vahliana should occur in North Amer- ica west of the Mackenzie River valley, as maintained by Hultén (1946, 1967, 1968). 45 Annotated List of Selected Species To test the hypothesis of condensed growth forms in an area where P. Vahliana definitely does not occur, | studied P. Ledebouriana in the Canadian Rocky Mountains, and thereby confirmed that here, also, condensed growth form is directly related to exposure. | illus- trated four plants (Porsild 1955: pl. 21) col- lected on the same mountain in Banff National Park, Alberta, approximately at the same ele- vation, but in different situations in regard to snow cover and exposure to wind. The centre plant (Porsild 1955: Pl. 21, fig. 3) is one of many cylindrical branches of a large, hemi- spherical cushion; the lower part of the branch is covered by the persistent leaf bases blackened by age and moisture. The remain- ing three specimens illustrated are parts of plants of less exposed habitats; in these the caudices are short, and the old leaf-sheaths have remained reddish brown and, as might be expected, the white, silky pubescence of the leaves is less dense than in the plants from exposed situations. An even more extreme example of con- densed P. Ledebouriana is shown in Plate 10. In life this dense and compact cushion con- sisted of a large number of branches, each terminated by a fascicle of leaves, some bare- ly 1 cm long, and the strongest branches ter- minated by a solitary flower on a peduncle 3 cm long. The abundant material of P. Ledebouriana collected in the Ogilvie Mountains by R.T. Porsild between 1966 and 1968 shows that condensed plants, resembling P. Vahliana in habit, occur there on exposed ridges, where- as taller plants of more matted growth are found in more sheltered situations, especially those that are assured ample snow cover in winter. Chromosome counts from Yukon plants representing both life-forms gave 2n=28 (Mulligan and Porsild 1969). For P. Vahliana from West Greenland, Jgrgensen Sgrensen, and Westergaard (1958) reported 2n=42, and Dansereau and Steiner (1956) reported 2n=48 and 2n=56 for plants from Baffin Island. Potentilla nivea L. ssp. Hookeriana (Lehm.) Hiit. Vicinity of Mayo, 63°35/’N, 135°45’W, on gravelly river terraces, Nos. 442 and 493. Potentilla nivea L. ssp. nivea Ogilvie Mountains: Dry limestone ledges along Dempster Highway, Mile 86, No. 1658. 46 Potentilla pennsylvanica L. var. strigosa Pursh Stewart Plateau: Vicinity of Mayo, on gravel- ly river terraces, Nos. 458, 523, and 658. A grassland and foothill species, in the north- west confined to the major interior river valleys. Sibbaldia procumbens L. Ogilvie Mountains: Canyon Creek near Mile 48, on Dempster Highway, Nos. 448 and 1841; mountains east of Miles 50-54, No. 87. A chromosome count of 2n=14 (unpublished), which agrees with the number reported from Greenland (Bocher, Holmen, and Jakobsen 1968), was obtained from No. 1841. Stewart Plateau: Mount Haldane, gravelly snowbed areas in cirque, elevation 5,500 ft, No. 583; Keno Hill, in herobmats bordering snowbeds, elevation 5,500-—6,000 ft, Nos. 983 and 1040. In Yukon otherwise known from mountains along the Canol Road (Porsild 1951a) and from south of Kluane Lake. LEGUMINOSAE Astragalus aboriginorum Richards. Ogilvie Mountains: West slope of mountains east of Dempster Highway, Miles 50-57, No. 83, flowering on 23 June 1966, and Nos. 246 and 392 with fully formed pods on 20 and 31 July 1966. The last two numbers represent a slight northward extension of the known range of the species in Yukon. Astragalus alpinus L. South branch of McQuesten River along east slope of Mount Haldane, No. 1204, 2n=32 (Mulligan and Porsild 1969); vicinity of Minto Bridge, 63°41’N, 135°51’W, common on river meadows, No. 649. Ogilvie Mountains: Com- mon along the Dempster Highway north to at least Mile 81, Nos. 1463 and 1538. Astragalus umbellatus Bunge Stewart Plateau: Northern slope of Mount Haldane, 63°51’N, 135°50’W, elevation 4,000 ft, No. 568; on west slope below timberline, flowering on 25 June 1967, No. 617; Keno Hill, 63°58’N, 135°42/W, in rich herbmat slopes, elevation 5,500 ft, No. 776; in rich alpine tun- dra, elevation 6,000 ft, No. 1284. Ogilvie Moun- tains: River-flat meadow along Dempster Highway, Mile 57, No. 138, 2n=16 (Mulligan and Porsild 1969); in rich herbmats by Mile 83, No. 1533. Common in alpine herbmats. Leguminosae Astragalus yukonis Jones Ogilvie Mountains: On calcareous riverbanks among willows, Dempster Highway, Mile 82, No. 1126. A new northern limit for Yukon. Hedysarum Mackenzii Richards. Vicinity of Mayo, 63°35’N, 135°45’W, among willows, No. 1010A. Ogilvie Mountains: In al- pine heath and on herbmat slopes of lime- stone mountains west of Miles 81-82 on Dempster Highway, Nos. 1088 and 1502; low- er slopes of limestone hills north and south of Mile 110, A.E. & R.T. Porsild, Nos. 23,651—52. The last station cited bridges a gap in the known distribution of the species between Dawson and the Arctic Coast of Yukon. Medicago falcata L. Stewart Plateau: Vicinity of Mayo among wil- lows near the airport, No. 1203. An apparent- ly well-established roadside weed on 21 Aug. 1967, with flowers and almost mature seeds. The collection is the first from Yukon Ter- ritory. Melilotus officinalis (L.) Lam. Stewart Plateau: Vicinity of Mayo, near road to airport, No. 1175,-on 10 Aug. 1967, with flowers and mature seeds. Noted in 1970 as a well-established roadside weed, spreading into nearby clearings. Oxytropis deflexa (Pall.) DC. var. foliolosa (Hook.) Barneby Stewart Plateau: Keno Hill, elevation 3,800 ft, No. 510. Ogilvie Mountains: Gravelly river flats along Dempster Highway, Mile 81, No. 1466. In Yukon otherwise known from the Canol Road (Porsild 1951a) and from south of Kluane Lake. Oxytropis deflexa (Pall.) DC. var. sericea or & GC, Stewart Plateau: Vicinity of Minto Bridge, 63°41’N, 135°51’W, gravelly riverbanks, No. 921; vicinity of Mayo, 63°35’/N, 135°45’W, in open woods, No. 945. Ogilvie Mountains: River flats along Dempster Highway, Miles 57-58, on gravel ridge, No. 298. Oxytropis glutinosa Porsild Stewart Plateau: Keno Hill, elevation 5,500 ft, in gravel scree, No. 732. Ogilvie Mountains: Mountain east of Dempster Highway, Miles 50-54, elevation 5,000-6,000 ft, Nos. 33, 82, 181, 732, and 1834. A chromosome count of 2n=16 was obtained for No. 1834 (Mulligan and Porsild 1970). See also Porsild 1966. Per- haps the most common alpine member of the genus in Yukon. Oxytropis Huddelsonii Porsild, Nat. Mus. Can. Bull. 121: 242; Pl. 17, fig. 5 (1951). Stewart Plateau: Keno Summit, in alpine tun- dra, elevation 6,000 ft, Nos. 532, 771, and 1268. No. 1268 provided a chromosome count of 2n=16 (Mulligan and Porsild 1969), and had mature legumes on 11 Aug. 1966. A caespitose-pulvinate dwarf species form- ing low, flat cushions, rarely more than 12 cm in diameter, from a strong multicipital tap- root; the branches are cylindrical, their lower part densely covered by persistent, charta- ceous, white stipules that are glabrous on both sides, but copiously white-ciliate along the margins. The leaves are short, 2-3 cm long, and the rachis purplish grey, its 7-9 leaf- lets (not 15-19 as stated by Hultén 1968: 660) * sparsely covered by stiff white hairs, their margins strongly involute and always pink. Flowers 1-2, small, dark purple, the standard about 7 mm long, calyx short-hirsute with brown hairs, its short and triangular lobes one-third as long as the tube. Legumes ses- sile, almost glabrous, 2.5 cm long and 0.8-0.9 cm wide, ovoid and somewhat inflated when mature, on very slender and weak purplish- grey peduncles. In habit and by its ovoid and more or less inflated but not stipitate legumes, O. Huddel- sonii resembles O. podocarpa, and some early reports of O. podocarpa from northern British Columbia and southwestern Yukon (Britton and Rydberg, 1901) undoubtedly were based on immature specimens of O. Huddel- sonii. In O. podocarpa the legumes are broad- er and always distinctly stipitate within the Calyx; its nearest Known station is in Jasper National Park, Alberta. O. Huddelsonii is apparently another en- demic of southeastern and central Yukon and central-eastern Alaska; the type came from the Golden Horn Mountain, south of White- horse (Porsild 1966: 46, map 104). * Obviously a typographical error because the correct number is clearly shown in his illustration of O. Huddelsonii. 47 Plate 11 Oxytropis nigrescens (Pall.) Fisch. var. lonchopoda Barneby Approximately natural size 48 Leguminosae Oxytropis Jordalii Porsild, Can. Field-Natur. 65 (2):77-78, pl. 1 (1951). Ogilvie Mountains: Limestone hills west of Miles 81-82, on Dempster Highway, elevation 4,200 ft, where it is common in turfy, alpine tundra, flowering on 23-28 June 1968, No. 1486; with mature fruits on 4 Aug. 1967, No. 1083; stony lower slope of limestone hill south of Mile 110, flowering on 10 July 1970, A.E. & R.T. Porsild, Nos. 23,655—-56; moist, stony floodplain, same place, /idem, No. 23,654. Hultén (1967: 79) reduced O. Jordalii to a subspecies of O. campestris, “closely related to O. campestris ssp. sordida (Willd.) Hartm.”’. To me, the alleged “close” relationship to O. sordida Willd. or, for that matter, to any other species of Oxytropis is not obvious. In this view, | am supported by Gjaerevoll (1963: 79), who collected large series of O. Jordalii in the White Mountains of central Alaska, and who considered it “a very distinct species’, not suggestive of O. campestris var. varians, as suggested by Wiggins and Thomas (1962). Thus far, O. Jordalii was known only from the eastern Brooks Range of northern Alaska and from the central Mackenzie Mountains, Northwest Territories, and is therefore new to Yukon Territory. Oxytropis Maydelliana Trautv. ssp. Maydelliana Stewart Plateau: Keno Summit, on gravel patches, elevation 6,000 ft, Nos. 523, 769, and 1065. Ogilvie Mountains: Mountain east of Dempster Highway, Mile 51, elevation 5,000 ft, No. 36; in rich alpine tundra near Mile 81, elevation 4,200 ft, No. 1500; stony lower slope of limestone hill north of Mile 110, A.E. & R.T. Porsild, No. 23,653. For a discussion of the two races of O. Maydelliana in North America see Porsild 1966, where O. Maydelliana ssp. melanocephala (Hook.) Porsild is proposed for the eastern race, and where map 105 shows the distinct ranges of the two races. Oxytropis nigrescens (Pall.) Fisch. var. lonchopoda Barneby, Leafl. West. Bot. 10:23 (1963). O. pygmaea sensu Porsild, Nat. Mus. Can. Bull. 121: 243-44 (1951); ibid., 162: 47 (1966) non (Pall.) Fern. Ogilvie Mountains: Mountain ridge 52 miles northeast of Dawson, elevation 5,200 ft, Youngman & Tessier, No. 355; mountains east of Miles 50-54 on Dempster Highway, eleva- tion 5,000-6,000 ft, in herbmats and on rocky ledges, Nos. 35 and 177, flowering on 24 June 1966. O. nigrescens var. lonchopoda is other- wise known only from the type locality: Cathe- dral Rocks in the Ogilvie Mountains, 66°02’N, 138°44’W, J.A. Calder & J.M. Gillett, DAO, No. 9395 (Type). Also from 138 miles north of Watson Lake and 5 miles east of Little Hyland River, elevation 6,000 ft, Youngman & Tessier, No. 418; Ogilvie Mountains, Yukon Territory, approximately 65°37’N, 138°56’W, J.A. Calder and J.M. Gillett, No. 25962, DAO 116523 (Plate TH). Oxytropis Scammaniana Hult. Ogilvie Mountains: In alpine situations along Dempster Highway, Mile 52, flowering on 15 July 1963, Youngman & Tessier. No. 379; al- pine herbmat slopes, elevation 6,000 ft, east of Miles 57-58, flowering on 31 July 1966, No. 405; Mount Nansen, 62°06/N, 137°17’W, in alpine Dryas-lichen heath, No. 1365. O. Scammaniana, in common with O. Hud- delsonii, O. Jordalii, and O. sheldonensis, were described from mountains of ungla- ciated central Alaska and Yukon; all except O. Huddelsonii have now also been reported from the Mackenzie Mountains, Northwest Territories. The type locality of O. Scamma- niana is Eagle Summit near Circle Springs on Steese Highway. It is a very handsome spe- cies that commonly forms extensive mats in alpine Dryas-lichen heath. Characteristic and conspicuous are its elongated and freely branched caudices that are covered by the im- bricated, glabrous, and prominently 3-nerved stipules, the uppermost with rounded, pale- green and free tips that turn straw-coloured in age. The slender leaf rachides and the flowering peduncles are characteristically purplish grey, as in O. Huddelsonii. \ts bright- purple flowers are very showy, usually in clusters of 3; the calyx is coal-black from dense pubescence, its teeth narrowly trian- gular or almost linear, two-thirds as long as the tube; the subsessile legumes are densely black hirsute, rarely over 15 mm long, and, when mature, conspicuously elevated above the plant mat by elongation of the scape. Oxytropis splendens Doudgl. Vicinity of Halfway Lakes north of Mayo, 63°48’N, 135°47’W, on high, gravelly river terraces, Nos. 527 and 657. A dry grassland and foothill species. A slight northward ex- tension of the Yukon range for this species. 49 Annotated List of Selected Species GERANIACEAE Geranium Bicknellii Britt. Stewart Plateau: Vicinity of Minto Bridge ina river meadow, No. 647, with flowers and well- formed fruits on 30 June 1967. In Yukon other- wise known from Dawson, Teslin Lake, and Whitehorse (Porsild 1951a). The sporadic occurrence of G. Bicknellii in the Canadian northwest strongly suggests a fairly recent introduction by man. CALLITRICHACEAE Callitriche hermaphroditica L. Stewart Plateau: Crystal Lake north of Keno Hill, in shallow water, No. 1234; north branch of McQuesten River, in small lakes with Ra- nunculus aquatilis, No. 1158. Ogilvie Moun- tains: Lake west of Dempster Highway, Mile 58, in shallow water, Nos. 326, 484, and 499. In Yukon otherwise known from Whitehorse and from two stations on the Canol Road (Porsild 1951 a). Callitriche verna L. emend. Lonnr. Vicinity of Mayo: In small lakes 4 miles north of town, elevation 1,900 ft, No. 787; Minto Bridge in shallow pond, No. 874. Ogilvie Mountains: In shallow ponds along Dempster Highway, Miles 57-58, Nos. 301 and 437; and Miles 90-91 in ponds on river flats, Nos. 1753 and 1755; Bell River near Lapierre House, Youngman & Tessier, No. 622. The last is a new northern limit for Yukon. VIOLACEAE Viola epipsila Ledeb. ssp. repens (Turcz.) Becker Apparently common in moist, grassy places along the Dempster Highway where it was the only member of the genus noted, and where it may ascend above the timberline, Nos. 74, 108, 279, and 1823. V. epipsila is the only North American member of the genus to range beyond the Arctic Circle almost to the shores of the Arctic Ocean. Mount Sedg- wick, 68°58’N, 139°01’W, A.M. Pearson, No. 70-24A. 50 ELAEAGNACEAE Shepherdia canadensis (L.) Nutt. Apparently common on wooded river ter- races of the Ogilvie Mountains, along Demp- ster Highway, Miles 54-82, Nos. 237, 489, 1633, and 1831. ONAGRACEAE Epilobium anagallidifolium Lam. Stewart Plateau: Keno Hill, common in damp moss by alpine brooks, elevation 5,500-—6,000 ft, Nos. 972, 1018, 1036, 1053, and 1221-22. Ogilvie Mountains: In wet moss by creek along Dempster Highway, Mile 47, No. 1845, which is a new northern limit for this species in Yukon; it has been collected, however, in the Richardson Mountains west of the lower Mackenzie River and, no doubt, also occurs in mountains of northern Yukon. Epilobium arcticum Sam. Ogilvie Mountains: In alpine herbmats east of Dempster Highway, Mile 54, elevation 6,000 ft, No. 473. A truly arctic species of amphi- Atlantic range, in North America thus far known from central East and West Greenland, the islands of the Canadian Arctic Archipel- ago, including Axel Heiberg Island west to Banks Island, to the northern Hudson Bay shores, with one disjunct station in the Rich- ardson Mountains west of the Mackenzie Delta. The present collection is the first from Yukon. Our specimens are 6-8 cm high, with stiffly erect capsules containing almost mature seeds on 15 Aug., when the short overwinter- ing offsets were fully formed. E. davuricum grew on forested river flats in the valley below. In E. arcticum the leaves are oblong- oblanceolate, 2-4 times longer than wide, whereas in E. davuricum they are linear to narrowly lanceolate, 10-13 times longer than wide. For a discussion of the status of E. arcticum see Porsild 1955: 165. Epilobium davuricum Fisch. Ogilvie Mountains: On forested river flats along Dempster Highway, Miles 57-58, eleva- tion 2,500 ft, on gravel bars, specimens 40 cm tall, with flowers and mature fruits on 23 July 1966, No. 306. Umbelliferae Epilobium glandulosum Lehm. var. adenocaulon (Hausskn.) Fern. Vicinity of Mayo, 63°35’N, 135°55’W, along shores of small lake, Nos.1202, 803, and 1014. A chromosome count of 2n= 36 was obtained from No. 1202 (Mulligan and Porsild 1969). Ogilvie Mountains: In rich meadow west of Dempster Highway, Miles 96-98, No. 1729, which represents a new northern limit for Yukon. In Yukon our plant was reported from the southern parts of the Canol Road (Porsild 1951a) as E. glandulosum var. perplexans (Trel.) Fern., which, according to Fernald (in Gray 1950), is a synonym for E. ciliatum Raf. Epilobium Hornemannii Reichenb. s. /at. Stewart Plateau: Mount Haldane in wet gravel in cirque, elevation 4,500 ft, No. 589; “‘Highlet Gulch’, tributary to Minto Creek, Nos. 824, 825, and 826. Ogilvie Mountains: River flat along Dempster Highway, Miles 51-52, No. 218. Southwestern Yukon: Environs of Mack- intosh south of Kluane Lake, Schofield & Crum, Nos. 7774, 7777, 8021, and 8129. E. Hornemannii and the closely related E. lactiflorum are alpine but not arctic species, with bicentric and similar ranges in North America. Thus, the range of the eastern popu- lation of E. Hornemannii extends from central East and West Greenland over Labrador- Ungava, Newfoundland, to the Gulf of St. Lawrence and the foot of James Bay. Its western population, separated by a gap of nearly 1,500 miles of woodland, grassland, or tundra, extends from Seward Peninsula in western Alaska, through the Canadian Cor- dillera, south to Colorado. Thus far, it has not been reported from the Northwest Terri- tories, but will likely turn up in the mountains west of the Mackenzie River. Epilobium lactiflorum Hausskn. Keno Hill, 63°58’N, 135°42’W, elevation 5,500 ft, on rich herbmat slope, No. 973. In Yukon previously reported from the Mayo district and from several stations along the Canol Road and Macmillan Pass (Porsild 1951a). Southwestern Yukon: Known from the vicinity of Mackintosh, Schofield & Crum, No. 7711; from St. Elias Mountains, Dezadeash River valley, on snow patch slope at west end of Kathleen Lake, elevation 4,200 ft, A.M. Pear- son, No. 67-221A. HALORAGACEAE Myriophyllum exalbescens Fern. Ogilvie Mountains: Lake west of Dempster Highway, Mile 58, elevation 3,100 ft, in shal- low water, Nos. 320, 482, and 493. In the sum- mer of 1966 only sterile specimens were seen, and on 17 Aug. some had well-formed winter buds; south end of Chapman Lake, near Mile 72, No. 1759, in beginning anthesis on 5 Aug. 1968. Stewart Plateau: Minto Bridge, in shal- low pond, No. 879; small lake near Elsa, No. 1160, fruiting specimens on 11 Aug. 1967 (Porsild 1951a, 1966). Myriophyllum verticillatum L. var. pectinatum Wallr. Stewart Plateau: In shallow ponds north of Mayo, Nos. 805 and 1245, the last with well- developed spike of well-formed but still im- mature fruits on 28 Aug. 1967. The collections are the first from Yukon, and bridge a wide gap between central Alaska and the Macken- zie River valley (Cody and Porsild 1968). UMBELLIFERAE Cicuta bulbifera L. Stewart Plateau: In shallow water or wet mud by lake northeast of Mayo, Nos. 1250, 1857, and 1974. New to the flora of Yukon, and a new western limit in Canada, where its pre- viously Known westernmost station was at Norman Wells on the Mackenzie River. Cicuta mackenzieana Raup Stewart Plateau: In wet meadow 4 miles north of Mayo, Nos. 783 and 1244, associated with C. bulbifera. The present collections close a gap between the Dawson district and Shel- don Lake on the Canol Road. Conioselinum cnidiifolium (Turcz.) Porsild Stewart Plateau: Edge of woods northeast of Mayo, No. 673; gravelly riverbank near Daw- son Highway near Mile 210, between Car- macks and Pelly Crossing, No. 1390. A com- mon species of alluvial lakeshores and river- banks, but thus far not reported from the mountains separating the Yukon and lower Mackenzie drainages. 51 Annotated List of Selected Species PYROLACEAE Pyrola secunda L. The ssp. secunda and ssp. obtusata are wide- ranging across Canada, but as noted (Porsild 1951a:261), ssp. secunda is a woodland plant, whereas ssp. obtusata, always easily distin- guished by its smaller and pale yellowish- green leaves (1.5 cm long and 1.0 cm wide) and few-flowered racemes, is arctic-alpine, ranging far beyond the treeline, and even reaching southern Banks and Victoriaislands. In Yukon ssp. secunda has not been reported north of latitude 65°. ERICACEAE Cassiope tetragona (L.) D. Donssp. saximontana (Small) Porsild The cordilleran ssp. saximontana differs from the circumpolar ssp. tetragona by its consis- tently smaller flowers and much shorter pe- duncles; its corollas are about 5 mm long, against 7-8 mm in ssp. tetragona; in ssp. tetragona the flowers are commonly raised well above the tip of the branch on a peduncle up to 2.cm long, whereas in ssp. saximontana the flowers are clustered near the tip of the branches on very short peduncles that scarcely elongate in the post-floral period. Since ssp. saximontana was first reported from the southern half of the Canol Road in southeastern Yukon (Porsild 1951a), it has been collected, as predicted, in the St. Elias Mountains and in southwestern Yukon: West of Kluane River in alpine tundra, elevation 4,800 ft, L.A. Spetzman, No. 292; on Mount Bratnober in alpine tundra, elevation 5,000 ft, idem, No. 293; near terminus of Dusty Gla- cier, 60°23’/N, 138°12’W, elevation 4,400 ft, A.M. Pearson, No. 230. In southeastern Yu- kon, as noted (Porsild 1951a), ssp. saximon- tana was found mainly on snowflushes in ravines, and was not seen to form extensive heath. Only on Mount Sheldon were both races seen, although never together, nor was any sign of intergradation noted. The ssp. saximontana, in common with several other cordilleran taxa, barely enters Yukon in mountains of the south and south- west, but it has not been seen in mountains of northern or central Yukon, nor in the northern Mackenzie or Richardson mountains. How- ever, according to Gjaerevoll (1967:15) and Hultén (1968:725), its range extends over a2 southwestern Yukon, westward to longitude 150° in the Alaska Range. Loiseleuria procumbens (L.) Desv. This species is strongly acidophilous, and its spotty distribution in Yukon and elsewhere within its general range is, no doubt, largely controlled by its preference for acid rocks. In Yukon it is known from granitic parts in the south, but is apparently lacking in the central and largely calcareous plateau, reap- pearing again in the northern parts, where it was recently collected on the south slope of the British Mountains: ‘Sam Lake’, 68°30/N, 138°05’W, elevation 2,500 ft, A.M. Pearson, No. 70-20. Rhododendron lapponicum (L.) Wahlenb. Stewart Plateau: Mount Haldane, among Abies lasiocarpa scrub, elevation 5,500 ft, No. 663. Ogilvie Mountains: River flat along Dempster Highway, Mile 53, No. 112, and noted in alpine tundra near Mile 115. Vaccinium uliginosum L. Mount Haldane, 63°51’N, 135°50’W, on the northern slope ascending to the timberline, approximate elevation 4,000 ft, where the var. alpinum L. forms a low scrub, No. 576; above the timberline var. alpinum is replaced by var. microphyllum Lge., which grows here as a low, flat shrub in open lichen heath, No. 661 (Porsild 1951a:268). The var. alpinum was noted as common in muskeg and open spruce bogs along Dempster Highway, Miles 51—52, No. 203, and was still noted occasionally as far as Mile 115. DIAPENSIACEAE Diapensia obovata (F. Schm.) Nakai The soil preference and the Yukon range of this species are rather similar to that of Loiseleuria procumbens, although thus far Diapensia obovata has not been reported in Yukon from north of latitude 64°. It is, how- ever, known from the east slope of the Richardson Mountains (Porsild 1943:43), and will, no doubt, also be found in the mountains of northern Yukon. The westernmost station known for D. lapponica is at the head of Bathurst Inlet (longitude 106°W) on the Arctic Coast of Canada. Gentianaceae PRIMULACEAE Androsace Chamaejasme Host. Mount Nansen, 62°06’N, 137°17’/W, in dry, al- pine-lichen heath, No. 1369A. Ogilvie Moun- tains: dry, calcareous tundra along Dempster Highway, Miles 81-82, No. 1474; rocky ledges and screes west of Mile 100, var. arctica R. Knuth, Nos. 1704 and 1856, 2n= 20 (Mulligan and Porsild 1970). These all belong with the common northwestern representative of A. Chamaejasme, variously distinguished as var. arctica R. Knuth or as ssp. Lehmanniana (Spreng.) Hult.; its leaves are narrowly lan- ceolate, 6-8 mm long, flat and densely villous from long, simple hairs, at least on the lower surface and along the margins; the expanded corollas are 5-8 mm in diameter, usually creamy white, often turning pink in drying; the scapes are 5-12 cm tall, elongating in post-anthesis, as are the flowering peduncles. In the northwest, as well as in the Canadian Rocky Mountains, Androsace Chamaejasme is commonly confined to well-drained calcar- eous, stony, or gravelly soils. A smaller and more delicate plant was col- lected south of Dempster Highway, Mile 110, on the lower slope of limestone hill, No. 1111, A.E. & R.T. Porsild, No. 23,658. It differs rather strikingly from the series cited above by its smaller size and above all, by its linear- oblong, prominently keeled leaves, 3-4 mm long, crisply short-villous on both sides, by its glabrate scapes 2-5 cm tall, and by its umbels, usually 2-3-flowered, subtended by 2 prominently saccate bracts. Calyx short- villous, about 3 mm long, corolla 3-4 mm in diameter, pink. Douglasia Gormanii Constance Mount Nansen, 62°06/N, 137°17’W, among lichens in alpine heath, No. 1359; Mount Hal- dane, 63°51’N, 135°50’W, in slide-rock, eleva- tion 5,500 ft, No. 666; rocky patches in Cas- siope heath, elevation 4,000 ft, No. 561. Ogil- vie Mountains: Mountain east of Dempster Highway, Mile 51, approximately 64°00’N, 138°20’W, not uncommon in damp, gravelly places and on “frost boils” in alpine tundra, elevation 6,500 ft, Nos. 1576 and 1582. New northern limit for Yukon Territory. In the flowering state D. arctica and D. Gormanii are not always readily distinguish- able, but as noted (Porsild 1951a), the latter can nearly always be recognized by its more oulvinate-caespitose habit, by its short, branched hairs on the margins and the upper surfaces of the leaves, and above all by its short, non-elongating fruiting peduncle that is rarely more than1cm long. In D. arctica the peduncles are 2-5 cm long, and the leaves are glabrous above (Porsild 1966: map 117). Primula mistassinica Michx. Bonnet Plume Lake, approximately 64°20/N, 132°00’W, W.H. Butler & N. Olsen, No. 79, 11-27 July 1966. The first, and the only other, Yukon specimen of P. mistassinica known to me is from the Frances River, approximately 61°N, 129°W, collected by Geo. M. Dawson in 1887. Primula sibirica Jacq. Braeburn Lake, 61°27’N, 135°46’W, on peaty lakeshore, No. 1920. The North American dis- tribution of P. sibirica is peculiar; an eastern population is known from southwestern Yu- kon and adjacent Alaska, and a western population is apparently confined to the east- ern and western shores of Bering Sea and Bering Strait. Thus far, no specimens have been reported from the intervening 500-mile gap. Trientalis europaea L. ssp. arctica (Fisch.) Hult. Stewart Plateau: Vicinity of Mayo, in rich spruce-birch woods, No. 697. The collection bridges a gap between the Dawson area and the upper Canol Road (Porsild 1966: map 120). GENTIANACEAE Gentiana algida Pall. Ogilvie Mountains: Open, damp places among tall willows along Dempster Highway, Mile 58, No. 272; Mile 67, No. 1814 and Mile 82, elevation 4,200 ft, No. 1143. The last sta- tion is a new northern limit for Yukon. Gentiana prostrata Haenke Keno Hill, 63°58/N, 135°42’W, elevation 5,500 ft, in rich herbmat with G. arctophila, No. 952A. Ogilvie Mountains: River flats along Dempster Highway, Mile 57, elevation 2,500 ft, No. 156; near Mile 85, openings in willow thickets, No. 1554. The last is a slight north- ward extension of the known range for Yukon, where thus far it has been collected a few times along the Canol Road, in the Kluane Lake region, and near Dawson. 53 Annotated List of Selected Species Lomatogonium rotatum (L.) E. Fries ssp. tenuifolium (Griseb.) Porsild Stewart Plateau: Carmacks Road, Mile 71, near Twin Lakes, in a wet meadow, No. 1865. Ogilvie Mountains: In moist tundra near Miles 82-86 on Dempster Highway, Nos. 1089, 1620, 1626, and 1115; No. 1115 is a rather striking white-flowered form: f. albiflorum n. forma, corollis albis (Type: CAN). As noted (Porsild 1955:174), Lomatogo- nium rotatum in North America is composed of two distinct geographical races, of which ssp. rotatum is strictly littoral,“ of lower stat- ure, and with fewer and larger flowers. The ssp. tenuifolium (Griseb.) Porsild is a non- littoral inland race, distinguished by taller and more fastigate growth, narrower leaves and calyx lobes, and by more numerous but smaller flowers. The latter is wide-ranging on interior floodplains from central Alaska, Yu- kon, and the Mackenzie River valley, south through foothills and interior valleys of the Rocky Mountain region to Colorado. Menyanthes trifoliata L. Ogilvie Mountains: Common in ponds along Blackstone River near Mile 75 on Dempster Highway, Nos. 186 and 1780; Old Crow River near mouth of Johnson Creek, 67°35/N, 139° 50’/W, elevation 800 ft, Youngman & Tessier, No. 662. The last is a new northern limit for Yukon Territory. POLEMONIACEAE Collomia linearis Nutt. Vicinity of Minto Bridge, 63°41’N, 135°51/W, in a meadow above the bridge, flowering on 18 July 1967, No. 892. Ogilvie Mountains: Dempster Highway, in rich meadow near Miles 96-98, apparently well established, with fully ripe fruits on 30 July 1968, No. 1718. The Genus Phlox in Alaska and the Yukon Hooker (1838:73) noted that specimens of what he considered to be Phlox sibirica from Kotzebue Sound, Alaska, collected during Capt. Beechey’s Voyage, “‘differ in no respect whatever from the numerous Siberian and Dahurian ones in my herbarium’. Until re- cently, students of the flora of Alaska accept- * A single, truly inland station is known from the east shore of Great Bear Lake, Northwest Terri- tories, where | noted ssp. rotatum by a cold mineral spring (A. E. & R. T. Porsild, No. 5183). 54 ed this view, although Hultén (1948:1319) re- corded that ‘Alaska specimens have shorter and broader leaves than most of the Siberian specimens seen’, but this, he added, “‘is natural, as the Alaskan specimens are of the northernmost within the area of the species”. The American range then known to Hultén (1948: map 998) included the American shores of Bering Strait from just north of the Yukon Delta north to Cape Lisburne, and, in the interior, four stations on the south slope of the Brooks Range, the nearest disjunct by some 200 miles from the coastal population. In recent years P. “‘sibirica’’ has been col- lected in Alaska on calcareous, volcanic rocks wherever they outcrop: from Norton Sound north to Cape Beaufort, in the central Brooks Range, and on calcareous shale or limestone in the British, Richardson, and Ogilvie mountains of Yukon Territory. The large series of “sibirica’’ specimens that | have examined from Alaska and from the Yukon River are very uniform, but although they are obviously related to, they cannot satisfactorily be placed with, P. sibirica L. |In- cidentally, P. sibirica L. is the only Old World member of this otherwise North American genus. Although its ancestors may have origi- nated in North America, its present range ex- tends from the southern Ural Mountains, east- ward along the northern rim of the central Asiatic highlands to the upper Kolyma River in eastern Siberia, and it is thus separated from its closest North American relative, P. alaskensis Jordal, by a gap of 1,500 miles. In addition to P. “sibirica’’ and P. alasken- sis, two other species of Phlox have been re- ported from our area. These are P. Richard- sonii Hook. and P. Hoodii Richards. The first, although reduced to P. sibirica ssp. Richard- sonii Hult. (Hultén 1967), is in my opinion abundantly distinct and actually endemic in a small area in the Canadian Arctic, with a few disjunct and alpine stations in unglaciat- ed mountains of northern Yukon and central Alaska. P. Hoodii is neither alpine nor arctic, but is a foothill species of dry, gravelly river- banks and terraces, wide-ranging in these situations in the Canadian Prairie Provinces and in the northern and central United States. As noted (Porsild 1955:175; 1966:51—53), all Phlox Hoodii reported from Alaska and the Yukon are merely extreme and depauperate forms of P. Richardsonil. Polemoniaceae The following key should be helpful in dis- tinguishing northwestern members of the genus Phlox. A. Loosely caespitose or matted; leaves op- posite, connate, their thin, flat blades 1-3 cm long and about 2 mm wide, spaced by internodes more or less glabrous, 1-2 cm long; leaf margins villous from coarse, flat, crinkly hairs, more or less septate; flowers terminal, 1-3, on glandular or vil- lous peduncles from 0.5-3.0 cm long; co- rolla lilac or pink, the limb 1.5-2.5 cm in diameter. B. Leaves 1.0-1.5 cm long; flowers soli- tary, not scented and not fading in dry- ing, on very short, villous, but not con- spicuously glandular peduncles; calyx lobes villous; limb of corolla 1.5 cm in diameter or less; locules one-ovuled. Endemic in unglaciated, calcareous mountains of western and northwest- ern Alaska, eastward to northern Mac- kenzie Mountains, Northwest Terri- tories. P. alaskensis Jordal (=P. sibi- rica, of Amer. auctt. non L.). BB. Leaves 1.5-3.0 cm long; flowers com- monly more than one, on peduncles up to 3 cm long, densely short-glandu- lar, but usually not villous; calyx lobes very narrow, glabrous or thinly villous; locules 2-ovuled; limb of corolla about 2.0 cm in diameter. Mountains of cen- tral and eastern Siberia east to Kolyma River; not Known in North America. P. sibitica L.s.str; AA. Densely caespitose, from a strong, sub- ligneous taproot: leaves imbricated, mar- cescent for many years, with strongly revolute margins, commonly less than 1.0 cm long, cobwebby beneath; flowers soli- tary and terminal, the limb of the corolla less than 1.5 cm in diameter. B. Leaves awl-shaped, silvery grey; flowers small, not scented, the corolla white, 7-8 mm in diameter. Prairie and foothill species from southern Alberta, east to Manitoba, southward through foothills to Montana, Idaho, Wyoming, Nebraska, and Oregon. P. Hoodii Richards. BB. Leaves distinctly apiculate, flowers scented, corolla pink or lilac, drying white, 1.2-1.4 cm in diameter. Arctic- alpine. Endemic in western islands of the Canadian Arctic Archipelago, south to the Arctic Coast of the west- ern Mackenzie District and moun- tains of unglaciated western Yukon and central Alaska. P. Richardsonii Hook. Phlox alaskensis Jordal, Rhodora 54:38 (1952). P. sibirica auctt. non L. P. borealis Wherry, Morris Arboretum Monogr. S217 (1955): P. Richardsonii ssp. alaskensis (Jordal) Wher- ry, Baileya 4:98 (1956). P. sibirica ssp. borealis (Wherry) Shetler, Rhodora 65:220 (1963). Ogilvie Mountains: Mountain east of Miles 50-54 along the Dempster Highway, rocky ex- posures in alpine herbmats, elevation 4,800 ft, No. 169; same place on gravelly scree, ele- vation 6,000 ft, flowering on 24 June 1966, No. 21, and No. 1840 with ripe seeds and late flowers on 12 Aug. 1968; a chromosome count of 2n=14 was obtained from No. 1840 (Mul- ligan and Porsild 1970); limestone hills west of Miles 81-85, on rock stripes in alpine heath, elevation 5,200 ft, with ripe fruits on 4 Aug. 1967, Nos. 1106, 1521, 1566, and 1779; a chromosome count of 2n= 14 was obtained from No. 1779 (Mulligan and Porsild 1970); lower slope of limestone hills south of Mile 110, A.E. & R.T. Porsild, No. 23,659. In addition to the above, the following specimens of P. alaskensis from Yukon Ter- ritory and Mackenzie District are in the Na- tional Herbarium of Canada: Yukon Territory: Along the 141st meridian, approximately 67°N, south of Porcupine River, F.H. Nelles, No. 31; Peel River basin, 10 July, idem, No. 105; C. Camsell, No. 67,891; Firth River, 68°50/N, 140°33’W, Youngman & Tessier, No. 142; Mount Sedgwick, 68°58’N, 139°01’W, A.M. Pearson, No. 70-27B. Mackenzie Dis- trict: Eastern slope of Richardson Mountains west of Mackenzie Delta, approximately 68°N, 136°W, A.E. Porsild, No. 6712, reported as P. Richardsonii (Porsild 1948). Phlox Richardsonii Hook. Ogilvie Mountains: Mountain east of Demp- ster Highway, Miles 57-58, No. 264. Stewart Plateau: South of Mayo, 63°N, elevation 4,800 ft, Hugh Bostock, No. 223. In addition, the following Yukon collections of P. Richard- sonii are in the National Herbarium of Cana- da: “Selkirk Trail’? near Aishihik, J.B. Tyrrell, No. 19,443; same place, flowering on 15 June 1960, Mrs. J. Lammers; Alaska Highway be- 95 Annotated List of Selected Species tween Alaska boundary and Champagne, Yukon Territory, Lloyd A. Spetzman, No. 317; St. Elias Mountains, moraines along Steele Glacier, elevation 6,000 ft, D.F. & B. Murray, No. 1368; west of Rocky Mountains between Fort Yukon and Lapierre House, McT[avish]; Red Mountain, Yukon Territory, 61°0O0/N, 133°46’W, W.H. Gorman, No. 974; “Selkirk Trail’ near Aishihik, Yukon Territory, J.B. Tyrrell, 6 Sept. 1898. The last two numbers were cited as P. Hoodii by Hultén (1948:1318). Polemonium occidentale Greene, Pitt. 2:75 (1890). Polemonium: Brand in Pflanzenr. IV, 250, Heft 27:33 (1907). P. coeruleum ssp. occidentale (Greene) J.F. Davidson, Univ. Calif. Publ. Bot. 23:225 (1950). P. acutiflorum Willd. ssp. occidentale (Greene) Hult., Fl. Alaska & Yukon 8:1323 (1948). Ogilvie Mountains: Dempster Highway, Mile 23, on gravelly floor of abandoned rock quar- ry, flowering on 23 June 1968, No. 1430, with ripe seeds on 2 Aug., No. 1755. These, as well as the series of P. occidentale from central Yukon that | cited (Porsild 1951a), are insepa- rable from P. occidentale from Alberta, British Columbia, Washington, Montana, Wyoming, and Nevada. Polemonium occidentale is a cordilleran species, commonly up to 1 m tall, with an elongating inflorescence of 20 or more flow- ers, the petals not ciliate. P. acutiflorum is rarely more than half as tall; the inflorescence (cf. illustration in Hultén 1968:767) is short, often branching, not elongating in fruit, rare- ly with more than half a dozen flowers, the petals distinctly ciliate. Its range is typically amphi-Beringian, and in the Canadian Rocky Mountains it has not been collected south of latitude 55°N. Its chromosome numbers are 2n=18 (Mulligan and Porsild 1970), as are those of P. pulcherrimum and P. acutiflorum (Mulligan and Porsild 1969). The fact that the northern population of P. occidentale is dis- junct from that of the Rocky Mountains by a gap of more than 1,200 miles, and that in cen- tral Yukon and Alaska it is, to my knowledge, restricted to placer mining areas strongly suggests that P. occidentale is a recent intro- duction in this area, dating only as far back as the early days of gold mining, when horses were widely used in the mining camps, and when each year thousands of tons of hay were imported from Pacific ports of Canada 56 and the United States. In Alberta P. occiden- tale is commonly seen by roadsides or as a ‘“‘weed”’ in mountain pastures and hayfields. HYDROPHYLLACEAE Phacelia mollis McBride Ogilvie Mountains: River flat along Dempster Highway, Mile 57, growing in dry, gravelly places, with flowers and fully ripe seeds on 18 July 1966, No. 221; in a similar place near Mile 67, No. 1796, 2n=22 (Mulligan and Por- sild 1970). Avery rare plant, thus far collected a number of times in the Dawson Range of southwestern Yukon and in central Alaska. BORAGINACEAE Eritrichium aretioides (Cham. & Schlecht.) DC. Ogilvie Mountains: Limestone mountains west of Miles 81-83 on Dempster Highway, approx- imately 65°00’N, 138°15-20’W, dominant lo- cally in dry, calcareous tundra, elevation ap- proximately 4,200 ft, Nos. 1098, 1475, and 1619; in similar places on Dempster Highway south of Mile 115, A.E. & R.T. Porsild, Nos. 23,661-62. Common locally in northern Yu- kon, but perhaps always restricted to shallow, turfy places on limestone; as yet not reported from the Richardson Mountains in the North- west Territories (Porsild 1966). Eritrichium splendens Kearney A very rare plant described in 1902 from Baird Mountain in northwestern Alaska and long known only from a few other stations in moun- tains of northern and central Alaska; it was reported recently from the Richardson Moun- tains, Northwest Territories (Cody and Porsild 1968), and will likely also turn up on the west slope of the Richardson Mountains and else- where in northern Yukon Territory. LABIATAE Dracocephalum parviflorum Nutt. Stewart Plateau: Vicinity of Mayo on gravel banks near wood clearing, tall flowering specimens on 22 June 1967, No. 558; Hunker Creek near Dawson, 63°57’N, on gravelly creek bank, No. 1394; Midnight Dome near Dawson, 64°04’N, in open woods, No. 1418. | reported the species (Porsild 1951a) from a number of stations along the newly con- structed Canol Road in southwestern Yukon, Scrophulariaceae where its occurrence suggested recently es- tablished colonies in disturbed soil. If this taxon is not native to Yukon, it is now at least well established in many places. Mentha arvensis L. Braeburn Lake near Mile 54 on Whitehorse-— Carmacks Road, approximately 61°27’N, 135°48’W, on moist, peaty lakeshore, flower- ing on 15 July 1969. Scutellaria epilobiifolia A. Ham. ) Stewart Plateau: Vicinity of Mayo, by small lake 4 miles north of town, flowering on 13 July 1967, No. 806; Five Mile Lake with ma- ture fruits on 26 Aug. 1967, No. 1297. In Yukon thus far Known only from the vicinity of Mayo, where first reported by J.P. Anderson (1944). SCROPHULARIACEAE Castilleja hyperborea Pennell Mount Haldane, 63°51’N, 135°50’W, open al- pine heath, elevation 5,500 ft, Nos. 603 and 683; Keno Hiil, 63°58’N, 135°42’W, in alpine herbmats, elevation 5,500-—6,000 ft, Nos. 537, 980, and 1220; Galena Hill, in alpine heath, No. 1169. Ogilvie Mountains: Alpine herbmats east of Miles 57-58 on Dempster Highway, elevation 6,000 ft, Nos. 254 and 346; limestone hills west of Miles 81-82 in alpine heath, ele- vation 4,200-5,200 ft, Nos. 1108 and 1504; common near Mile 110, A.E. & R.T. Porsild, No. 23,662. Common locally and perhaps al- ways above timberline. Castilleja pallida (L.) Spreng. ssp. caudata Pennell Ogilvie Mountains: Common lowland species of riverbank meadows and among willows along Dempster Highway, Miles 57-58, Nos. 135 and 407, and Miles 81-82, Nos. 1134, 1541, and 1650. Castilleja villosissima Pennell Southwestern Yukon: St. Elias Mountains in Alsek River valley near “Beachview Creek Pass’, elevation 4,500 ft, near the type lo- cality, A.M. Pearson, No. 67-182A; Burwash Landing, in a dry meadow near the airstrip, A.E. Porsild, No. 23,528. Castilleja villosis- sima is perhaps most closely related to C. hyperborea and C. yukonis, and thus far is Known only from a few collections, all in southwestern Yukon. Castilleja yukonis Pennell Ogilvie Mountains: Calcareous floodplain on gravel bars in Blackstone River valley near Miles 81-83, Dempster Highway, No. 1604. Less common than C. hyperborea, and per- haps restricted to well-drained calcareous soils. Euphrasia subarctica Raup Mayo, along footpath to an abandoned Indian cemetery, No. 933. My map (Porsild 1966: map 126), which gives the Alaska—Yukon dis- tribution, should include a dot for Mayo. Lagotis Stelleri (Cham. & Schlecht.) Rupr. Ogilvie Mountains: Dempster Highway near Mile 110, common on moist, stony floodplain below limestone hills, A.E. & R.T. Porsild, Nos. 23,663-64. The collection closes a gap be- tween central Yukon and the Arctic Coast. Mimulus guttatus DC. In Yukon long known only from an early col- lection from Whitehorse Rapids, John Ma- coun, 1902 (Porsild 1951a:282). We now have confirmatory collections, both from the White- horse area: 4 miles south on “Fish Lake Road’, Micky Lammers, 5 Aug. 1963; Mcin- tyre Creek below water reservoir, No. 1929. Pedicularis arctica R. Br. Keno Summit, 63°58’N, 135°42’W, elevation 6,000 ft, No. 1047. Ogilvie Mountains: Com- mon in boggy tundra along Dempster High- way, Miles 44-110, Nos. 267, 280, 1448, and A.E. & R.T. Porsild, No. 23,665. P. arctica is wide-ranging across the Cana- dian Arctic Archipelago to northwestern Greenland, and is also not uncommon in al- pine situations west of the Mackenzie River valley, as well as in mountains of the Yukon and Alaska. In continental Northwest Ter- ritories, on the other hand, it is Known only from a single station on the Thelon River, 64°16/N, 103°35’W, G.B. Rossback, No. 6504. Pedicularis lanata Cham. & Schlecht. ssp. yukonensis n. ssp. Differt a P. lanata* galea tenuiter lanata. Yukon Territory: Dawson Range, Mount Nansen, 62°06’N, 137°17’W, 40 miles west of Carmacks, in damp, alpine tundra, flowering on 16-17 June 1968, No. 1377 (Type: CAN); Dawson Range 62°40/N, 138°00’W, elevation * P. Kanei Durand, according to Hultén 1967. S57 Annotated List of Selected Species 5,000 ft, Hugh Bostock, No. 273; approximate- ly the same place, elevation 5,500 ft, 19 July 1933, idem, No. 11. Kluane Lake: Burwash Landing, above timberline at 6,000-—7,000 ft, 3 July 1943, C.H.D. Clarke, No. 273; Burwash Creek, 61°30/N, 139°17’W, in damp meadow, elevation 4,500 ft, A.M. Pearson, No. 70-1; summit of “Sugden Pass’, 12 miles southwest of Haines Junction, elevation 4,900 ft, idem, No. 261; vicinity of Steele Glacier, moist lower slopes, David F. & Barbara M. Murray, No. 1444; Yukon River (probably near the Alaska border), Wm. Ogilvie, July 1887. Alaska: Di- vide Mountain, approximately 64°05’N, 141° 00’W, in tundra, 29 July 1967, Lloyd Spetzman. A few collections from Yukon Territory of P. lanata, with thinly hirsute galeas, have long been known to me. Because these speci- mens differed from typical P. Janata in no other respects, | did not consider the pres- ence of a few hairs on the galea of taxonomic significance. However, alerted by Hultén’s straight transfer of the Siberian P. Adamsii Hult. to P. Kanei ssp. Adamsii (1967:121), which he later described as “similar to ssp. Kanei, but galea pubescent and the lip ciliate in the margin” (Hultén 1968:827), | carefully examined all material of P. Janata in the Na- tional Herbarium of Canada, in addition to some material from other herbaria. The ex- amination showed that among the several hundred collections, eight from alpine situa- tions in central and southeastern Yukon and one from adjacent mountains of central Alas- ka could be distinguished from typical P. fanata by their hirsute galeas and ciliate lips. The fact that all nine collections came from alpine situations within a narrowly restricted area, subject only to local glaciation, sug- gests that the hairiness of the galea and the ciliate lip peculiar to these plants was per- haps of some taxonomic significance, and, moreover, suggests great age and long geo- graphical isolation. In the National Herbarium of Canada there are three flowering specimens labelled P. Adamsii Hult., all from the lower Lena River in central Siberia, distributed by Inst. Bot. Acad. Sci. URSS. The collectors are: V. Scha- murin, 1955; T. Polsova & B. Jurtzev, 1956; and L. Malyshev, 1964. These specimens match exactly the description of P. Adamsii Hult. (Hultén 1930:117), and differ from P. lanata Cham. & Schlecht., as well as from ssp. yukonensis, by their spikes (up to 5 cm wide) of very large flowers, in which the corolla is 58 3cm long, and the galea strongly falcate, dis- tinctly toothed below the apex. The plant illustrated by Hultén as P. Kanei ssp. Adamsii (1968:827) is smaller, but in habit and in the strongly falcate galeas close- ly matches the Lena River specimens cited above. In view of the discrepancies cited, the rela- tion of P: Adamsii Hult. to P. lanata (P. Kanei) is not very obvious, and it is puzzling that in his key Hultén (1968:816) places P. Kanei ssp. Adamsii under the heading: ‘Galea lacking in teeth in the apex’. At any rate, it can be defi- nitely stated that P. Janata ssp. yukonensis is not synonymous with P. Adamsii Hult., and that it appears to be endemic in a small and isolated alpine area in southwestern Yukon and adjacent parts of Alaska. Pedicularis lapponica L. Ogilvie Mountains: Dempster Highway, rare and local in rich alpine heath and herbmats, Miles 44—46, elevation 4,200 ft, No. 1459, and on river flat near Mile 63, No. 13; Old Crow River flats near “Sam Lake’’, A.M. Pearson, No. 70-19A. A subarctic, circumpolar tundra species, in Yukon and Alaska very rare and local south of latitude 67°N. Pedicularis Oederi Retz. Ogilvie Mountains: Rare and local in wet al- pine tundra rich in lime, near Mile 81 on Dempster Highway, elevation 4,200 ft, No. 1486 (Porsild 1966:54). Pedicularis sudetica Willd. Very common in wet tundra and in boggy places along the Dempster Highway, where it is the most common member of the genus. Ogilvie Mountains: Miles 44-110, Nos. 12, 241, 340, 504, 548, 748, 1046, 1120, 1408, and 1505. Very variable regarding shape and de- gree of elongation of the inflorescence, as well as in the shape and cut of cauline bracts and basal leaves. Pedicularis verticillata L. Ogilvie Mountains: Rare and local in moist tundra along Dempster Highway, Miles 81-82, Nos. 1532, 1132A, and 1132B. No. 1132A was with fully ripe fruits, and No. 1132B was in full flower. Both were collected in the same meadow on 5 Aug. 1967, but 1132B was in an area where overflow ice had remained late; near Mile 110 on calcareous floodplain, A.E. & R.T. Porsild, No. 23,667. Lentibulariaceae Rhinanthus groenlandicus Chab. Lake Laberge, in meadows along Richthofen Creek, 61°06/N, 135°24’W, No. 1953. In Yukon otherwise known from a few collections from near Whitehorse and from the Haines Road (Porsild 1951a:286). Synthyris borealis Pennell Ogilvie Mountains: Common locally in stony alpine herbmats on granitic mountains east of Dempster Highway, Miles 50-58, elevation 6,000-6,500 ft, Nos. 263, 178, 179, 411, and 1571. A chromosome count of 2n = 24 was ob- tained from No. 178 (Mulligan and Porsild 1969). See also Porsild 1966. Veronica alpina L. As noted (Porsild 1951a:283), var. a/terniflora and var. unalaschcensis are not so distinct in the Yukon as in eastern North America, and they are certainly not “so strongly marked geographic varieties’ as suggested by Fer- nald (1939:447). The first five numbers cited below were collected in alpine situations, and are best treated as var. unalaschcensis. Mount Haldane, 63°51’N, 135°50’W, on snowflushes in main cirque, elevation 5,500 ft, No. 584; Keno Summit, 63°58’N, 135°42’W, in alpine herbmats, elevation 6,000 ft, No. 1056. Ogilvie Mountains: Snowbeds along Dempster Highway, Miles 50-56, elevation 4,200-4,500 ft, Nos. 281 and 460; on late snowbed in canyon, near Mile 47, No. 1855, 2n=18 (Mulligan and Porsild 1970). One collection, No. 830, from tributary to Minto Creek, 63°45’N, 136°10’W, on mossy riverbank among willows, by its strongly elon- gated fruiting head of mostly alternate cap- sules on pedicels 4-9 mm long, and by its opposite leaves drying green, suggests the var. alterniflora Fern. OROBANCHACEAE Boschniakia rossica (Cham. & Schlect.) Fedtsch. Ogilvie Mountains: Dempster Highway, Mile 100, in mossy, open spruce woods on lime- stone hill, elevation 3,000 ft, parasitic mainly on the roots of Picea glauca and Alnus crispa, with mature seeds on 30 July 1968, No. 1714. LENTIBULARIACEAE Pinguicula villosa L. Vicinity of Halfway Lakes north of Mayo, 63°48’N, 135°47’W, elevation 2,300 ft, Nos. 498 and 645 .Ogilvie Mountains: On sphagnum hummocks in black spruce bog near Mile 55 on Dempster Highway, No. 153; in similar bog with Oxycoccus microcarpus near Mile 83, No. 1518; sphagnum bog at foot of limestone hills north of Mile 110, A.E. & R.T. Porsild, No. 23,668. Pinguicula vulgaris L. Ogilvie Mountains: Limestone hills west of Dempster Highway, elevation 5,200 ft, Miles 81-83, Nos. 1096, 1567, and 1601; lower slope of hills south of Mile 110, elevation about 3,000 ft, where very common on calcareous gravel, A.E. & R.T. Porsild, No. 23,669, asso- ciated with Dryas alaskensis. | noted (Porsild 1951a) that in 1944 Pingui- cula vulgaris was common on the eastern portion of the Canol Road, but | had not seen it west of the Yukon—Mackenzie border, nor had it then been reported from elsewhere in the Yukon. Pinguicula vulgaris is a pronounced calci- phile, which accounts for its uneven distribu- tion throughout its range. In addition to the Yukon stations cited above, it is now known from the following stations: Twin Lakes, ap- proximately 61°50’N, 136°00’/W, in boggy places by lakeshore, No. 1890; vicinity of Burwash Landing in southwestern Yukon, Raup et al., Nos. 12,280, 12,460, 12,488, and 13,925. Arctic Slope: Firth River, June Mc- Neish, CAN No. 236,763; Firth Lake 68°50/N, 140°33’W, Youngman & Tessier, No. 162, and, finally, Hultén (1967) reported it from a lake in the Mackenzie Mountains, 64°47’/N, 133° 31’/W. Utricularia intermedia Hayne Stewart Plateau: In a shallow pond northeast of Mayo, in post-floral state on 1 July 1967, No. 653. In Yukon otherwise known from one collection near Dawson and another on the Alaska Highway near the Alaskan border, J.P. Anderson, No. 1210. Utricularia minor L. Stewart Plateau: Vicinity of Halfway Lakes north of Mayo, flowering on 17 July 1967, No. 859. Ogilvie Mountains: River flats along Dempster Highway, Mile 55, in shallow ponds 59 Annotated List of Selected Species in a bog, Nos. 10 and 423, both sterile, the latter with winter buds forming at the tip of the branches on 3 Aug. 1966. In Yukon other- wise known from single collections near Dawson. Utricularia vulgaris L. var. americana A. Gray Vicinity of Mayo, 63°35’N, 135°45’W, common in small lakes, elevation 1,600—1,700 ft, Nos. 677, 810, 860, and 1201, the last with nearly ripe fruits on 21 Aug. 1967, the remainder with flowers on 1 July 1967. Ogilvie Mountains: In lake near Mile 75, on Dempster Highway, No. 1784; west of Miles 90-91, No. 1751; Nos. 1784 and 1751 sterile on 2 and 21 Aug. 1968. In Yukon this is the only member of the genus that is known to flower regularly, and in ex- ceptionally warm summers it may even ripen seeds. PLANTAGINACEAE Plantago septata Morris P. canescens sensu Hult., Fl. Alaska & Yukon 9 (1949) and FI. Alaska & Neighboring Terri- tories (1968) non Adams. Vicinity of Mayo, 63°35/N, 135°55’W, on river terraces, Nos. 457 and 1206. A chromosome count of 2n=12 was obtained from No. 1206 (Mulligan and Porsild 1969); Stewart Cros- sing, on gravelly river terraces, No. 1392. Described from the Yukon River valley, 62°03’/N, 135°40’W, P. septata appears to be- a lowland species of calcareous prairie and river terraces. It is endemic in northwestern North America, reaching the Arctic Coast of the Mackenzie District and the southwestern islands of the Canadian Arctic Archipelago, but in central Alaska not extending west of longitude 150°. In the eastern Rocky Moun- tain foothills known to me south to Jasper Park, Alberta, and according to Moss (1959: 222), extends to southwestern Alberta, but was reported from Montana by Rydberg (1922). RUBIACEAE Galium trifidum L. ssp. trifidum Vicinity of Mayo, 63°35’N, 135°45’W, marshy lakeshores, Nos. 654, 821, 1008, and 1196A. A chromosome count of n=12 was obtained from No. 1196A (Mulligan and Porsild 1969). Ogilvie Mountains: Swampy islands in lake west of Dempster Highway, Miles 90-93, No. 1585, with mature fruits on 13 July 1967. Bell 60 River, 67°18/N, 137°50’W, Youngman & Tes- sier, No. 626. The last is its northernmost station known from the Yukon. ADOXACEAE Adoxa Moschatelina L. Stewart Plateau, Mount Haldane, elevation 4,000 ft, No. 577. Ogilvie Mountains: River flats along Dempster Highway, Mile 56 on peaty ridges, flowering on 25 June 1966, No. 96; common locally in rich, loamy soil under willows along Miles 44-47, Nos. 1433 and 1848, the latter with mature fruits and 14 Aug. 1968. For the northwestern North American range of Adoxa, see Porsild 1966: map 133. VALERIANACEAE Valeriana sitchensis Bong. Mount Nansen, 62°06’N, 137°17’W, by alpine brook, No. 1382; Midnight Dome near Daw- son, 64°04’/N, 139°42’W, in moist woods, No. 1419; Keno Hill, 63°58’N, 135°42’W, rich herb- mat at 5,500 ft, No. 752; Bonnet Plume Lake, 64°20’/N, 132°00’W, Butler & Olsen, No. 89. Ogilvie Mountains: 52 miles northeast of Daw- son, Youngman & Tessier, No. 347; Dempster Highway near Mile 83, No. 1663. Valeriana sitchensis is a cordilleran-Pacific Coast species; in western Yukon its range overlaps that of the amphi-Beringian V. capi- tata Pall. (Porsild 1966: maps 134 and 135). CAMPANULACEAE Campanula aurita Greene Ogilvie Mountains: Limestone hills along west side of Blackstone River valley near Dempster Highway, Miles 81-83, Nos. 1086, 1607, and 1702, where it is common on cal- careous floodplains and river terraces; very common on limestone scree south of Demp- ster Highway, Mile 110, A.E. & R.T. Porsild, No. 23,670. A pronounced calciphile, endemic but not uncommon locally in Alaska, Yukon, and central Mackenzie Mountains, Northwest Territories. Campanula lasiocarpa Cham. Very common mainly in alpine situations in southern and central Yukon, but thus far not reported from northern Yukon or from ad- jacent parts of the Mackenzie District. The following stations are from slightly beyond the previously known northern limit for the Compositae species in Yukon. Ogilvie Mountains: Along Dempster Highway, Miles 50-54, Nos. 110, 163, and 172; moist alpine slopes and herb- mats near Miles 83-84, Nos. 361 and 1671. Campanula uniflora L. Ogilvie Mountains: River flats along Demp- ster Highway, Miles 51-52, Nos. 114 and 210; in alpine herbmat, Miles 50-54, No. 147; turfy, calcareous tundra and herbmat slopes near Miles 81-86, Nos. 1084, 1482, 1488, and 1652. In Alaska and Yukon C. uniflora appears to be consistently taller than in eastern Canada; thus, among the last four numbers cited there are fruiting specimens over 30 cm tall. COMPOSITAE Achillea nigrescens (E. Mey.) Rydb. A. borealis Bong. Ogilvie Mountains: Dempster Highway, Miles 81-86, Nos. 1140, 1470, and 1625. Very com- mon everywhere on dry slopes and gravelly floodplains. As noted (Porsild 1951a), itis one of the first native species to become estab- lished on disturbed soil along roadsides. Achillea sibirica Ledeb. Vicinity of Mayo, 63°35/N, 135°45’W, Nos. 687 and 858; Minto Bridge, Nos. 873 and 950. Ogilvie Mountains: Common in rich meadow on alluvial river flat west of Miles 96-98 on Dempster Highway, No. 1726. Although less common and of more re- stricted range than A. nigrescens, A. sibirica likewise tends to become weedy and to readi- ly invade disturbed soil along highways, river- banks, and townsites. The Genus Antennaria In 1950 ! noted (Porsild 1950a:1512) that the number of species of Antennaria in Alaska was small, but that their numbers and com- plexity increased rapidly as one entered east- ern and southeastern Yukon and British Co- lumbia. In the Old World the genus is repre- sented by only six species, or, according to Flora of the USSR, vol. 25, nine species. The flora of eastern and southeastern Yukon is clearly an extension of the cordilleran flora and, like it, is rich in species of this essen- tially North American genus. In the 20 years that have elapsed, a great deal of new Antennaria material from Alaska and the Yukon has come to me. In addition, during seven seasons of fieldwork in the northern Rocky Mountains, | became well acquainted with the rich Antennaria flora there. Contrary to the views expressed by Hultén (1967), | now find that this new mate- rial, as well as the wider experience gained in the Rocky Mountains, by and large confirms rather than weakens the conclusions ex- pressed in my 1950 monograph (Porsild 1950b) of the northwestern members of the genus. It seems fairly obvious that certain species are wide-ranging arctic-alpines, and are therefore restricted to alpine situations in central Alaska and the Yukon; some are low- land species belonging with what | have called the eastern boreal forest element of the northwestern flora, of which few members reach into central Yukon, and very few are remnants of a once more widespread prairie or open grassland flora. It seems obvious, too, that the genus Antennaria is essentially North American, and that it is unrealistic to attempt to relate North American members of the genus to those of the Old World. - Section Alpinae Bracts of the involucre (phyllaries) with pale greenish-brown to olivaceous or dark-brown tips, usually acuminate and erose. Antennaria angustata Greene Ogilvie Mountains: Dempster Highway, Mile 56, on grassy lower slopes of “Sheep Moun- tain’, A.E. & R.T. Porsild, No. 23,671. An east- ern arctic-alpine species, in Yukon collected only once before, on Firth River near the Arc- tic Coast, E.H. McEwen, No. 206, 6 Aug. 1953. Antennaria compacta Malte Mount Nansen, 62°06’N, 137°17’W, 40 miles west of Carmacks, in alpine lichen heath near summit, No. 1376. Densely caespitose dwarf species, wide-ranging from northern Green- land across northern islands of the Canadian Arctic Archipelago to mountains of central and northern Yukon and Alaska. Antennaria densifolia Porsild Ogilvie Mountains: Limestone hills west of Mile 82, on Dempster Highway, rare in alpine heath, No. 1109; on solifluction lobes of cal- careous slopes, No. 1481; lower slopes of limestone hills south of Mile 110, A.E. & R.T. Porsild, No. 23,672; on moist calcareous gravel at base of same hill, idem, No. 23,673. In all the collections cited, male and female plants are represented; at anthesis the male 61 Plate 12 Yi, ‘? « Antennaria Ellyae n. sp. (Type) Approximately % natural size 62 Compositae flowers are quite showy because of their ex- panded pinkish-yellow corollas. The collec- tions are the first from the Yukon. By its caes- pitose habit, short, crowded, and sessile branches, and small dioecious heads, Anten- naria densifolia is one of the most easily recognized members of the genus. Thus far, A. densifolia was known only from a dozen or so collections from limestone hills of the east slope of the central Mackenzie Moun- tains, between latitudes 60°N and 64°N. Antennaria Ekmaniana Porsild Keno Summit, 63°58’N, 135°42’W, elevation 6,000 ft, in gravel spots on alpine tundra, No. 712. Ogilvie Mountains: Gravelly solifluction lobes on mountain east of Miles 44-46, Demp- ster Highway, elevation 4,200 ft, No. 1444; mountain east of Miles 57-58, elevation 6,500 ft, No. 372; in alpine herbmats near elevation 5,000 ft, No. 399. Arctic species wide-ranging from Green- land to northern Alaska and south, almost to latitude 60°N in high mountains of Yukon. Antennaria Ellyae n. sp. Humifusa, stolonibus bene evolutis, 3-6 cm longis, sublignosis, foliis valde confertis per- brevibus, cuneato-obovatis vel late oblanceo- latis-obtusis nec mucronatis, 4-6 mm longis, 3-4 mm latis, utringue dense coacto-tomen- tosis. Caules floriferi 8-17 cm alti, coacto- tomentosis, foliis 5-11 linearibus ca. 1.0 cm longis, 1-2 mm latis. Folia inferiora mere acuta, superiora apice anguste scarioso- apiculata. Calathiae femineae 3-7, cymam elongatam formantes, pedunculis inferioribus 7.5-3.0 cm longis. Involucrum 3 mm altum, bracteis bi-vel triseriatis sub-aequalibus; ex- terioribus basi atro-brunneis. Stylus bene exsertus. Achaenia laevia. Planta mascula ignota. Ogilvie Mountains: Gravelly river flats near Dempster Highway, Miles 81-83, elevation 3,500 ft, fruiting on 26 July 1968, No. 1700 (Type: CAN), Plate 12; same place, No. 1605, 19 July 1968; rocky ledges on Dempster High- way, Mile 148, No. 2016. By its very short, broad, and densely to- mentose leaves, A. E/lyae resembles A. dens,- folia, from which it is, however, at once dis- tinguished by its spreading, leafy, stolon-like branches and by its flowering stems, twice as tall and elongated, cymose inflorescence, strongly contrasting with the head-like, few- flowered inflorescences of A. densifolia, in which the male plant is apparently as com- mon as the pistillate. On the limestone hills of the Ogilvie Moun- tains between Miles 80-110 A. Ellyae was noted on calcareous river terraces near Miles 81-83, whereas A. densifolia invariably grew in more alpine situations. Antennaria Ellyae is named for Mrs. Elly Porsild of Whitehorse, Yukon Territory, who, during the summers of 1966, 1967, and 1968, accompanied and greatly aided her husband in the collection and preservation of vascular plants in the wilderness of central Yukon for the National Herbarium of Canada. Antennaria monocephala DC. Mount Haldane, 63°51’N, 135°50’W, above elevation 5,000 ft, Nos. 575, 660, and 670; Keno Summit, elevation 6,000 ft, 63°58’N, 135°42’W, No. 719B; Mount Nansen, 62°06’N, 137°17’W, alpine snowbeds, No. 1362. Ogilvie Mountains: Alpine situations along Dempster Highway, Miles 44—45, No. 1447; Miles 50-54, Nos. 80, 96, 143, 208, and 209; A.E. & R.T. Porsild, No. 23,671. Perhaps the most common member of the genus above the timberline, where it is ubiq- uitous on moist, neutral, and well-leached snowbed soils. In most populations the male plant is as common as the female. Antennaria neoalaskana Porsild Long known only from the type locality in the Richardson Mountains west of the Mackenzie Delta, A. neoalaskana has recently turned up elsewhere in the northern Richardson Moun- tains: 68°08’N, 136°46’W, J.D.H. Lambert, No. 26; 68°20/N, 136°30’W, elevation 3,500 ft, idem, No. 23. In each of the two collections that | examined, one plant is male and the other female; both came from the east of the Yukon—Mackenzie border. Two other collec- tions, from a few miles to the east and just in- side the Mackenzie District, are Lambert & Morrison, Nos. 65-07-0604 and 65-08-1102; both are female plants. A fifth collection, from Mount Sedgwick in the British Mountains, 68°58’N, 139°01’W, A.M. Pearson, No. 70-27C, consists of two staminate plants. | reported this species (Porsild 1950a:1523) from the Sadlerochit River on the northern slope of Brooks Range (Spetzman, No. 1100). | have since examined three additional Alas- kan collections: Mount Fairplay, 63°40/N, . 142°05’W, No. 1945; Alaska Range, Mile 14 63 Annotated List of Selected Species on Denalji Highway, Spetzman, No. 4051; and Ketchumstuk Mountain, idem, No. 5165. Antennaria pedunculata Porsild This rather striking species was described from the Rose River in southeastern Yukon (Porsild 19505). It is of stoloniferous habit and grows on gravelly river terraces, where it forms mats of up to 1 min diameter. In com- mon with A. sto/onifera, its achenes are very hispid, but it differs from that species by its extraordinary, long flowering peduncles and by the very prominent, flat, brown, scarious tips of the cauline leaves. Only the female plant was seen and collected. | now have a specimen that can only be the male plant of A. pedunculata. It came from Babbage River, due east of “Little Trout Lake’, Yukon Territory, approximately 69° 00/N, 138°13’W, Lambert & Morrison, 16 July 1965, No. 65-07-1603, CAN No. 308,595. Antennaria philonipha Porsild Stewart Plateau: Keno Hill, in alpine herb- mats, elevation 5,500-6,065 ft, Nos. 524 and 742. An alpine-arctic snowbed species, in Yukon previously reported from mountains of southeastern Yukon and from the Stewart Plateau (Porsild 1950b). The species is also known from the Mackenzie Delta and the east slope of the Richardson Mountains. It will un- doubtedly also be discovered in mountains of northern Yukon. Antennaria stolonifera Porsild Keno Summit, 63°58/N, 135°42’W, on gravel- ly solifluction stripes near elevation 6,000 ft, Nos. 719A and 1045. The specimens are a good match for A. stolonifera from south- eastern Yukon. In addition it has been col- lected in southwestern Yukon, A.M. Pearson, No. 67-217A, and was reported by Gjaerevoll (1967) from the Alaska Range (Porsild 1951 a). Section Dioicae The inner phyllaries papery, white, straw- coloured, or pink, and usually with spatulate- ligulate tips; in our area only the female plants are known. Antennaria alborosea Porsild Stewart Plateau: ‘‘Highlet Gulch’, tributary to Minto Creek, in gravel among willows, eleva- tion 4,000 ft, No. 846; vicinity of Halfway Lakes north of Mayo, edge of abandoned gravel pit, No. 883; Minto Road, 13 miles north 64 of Mayo, No. 909. Ogilvie Mountains: Along Dempster Highway, Miles 96-98, No. 1719 (Porsild 1966: map 137). A rather attractive and well-marked spe- cies, often pioneering on non-alpine, well- drained sites that are not too dry, such as river terraces or gravelly floodplains, where, because of its stoloniferous habit, it often forms mats up to 1 m in diameter. Charac- teristic of A. al/borosea are the slender flower- ing stems, up to 30-35 cm tall and very leafy, that are often bent or S-shaped because of the weight of the rather large, pinkish, and dense flowering head. Antennaria incarnata Porsild Vicinity of Mayo, 63°35/N, 135°45/’W, on grav- elly river terraces, elevation 1,600 ft, No. 694. Described from the Pelly Mountains in south- eastern Yukon, A. incarnata is also known from a few collections in southwestern Yu- kon and from Great Bear Lake, Northwest Territories. It is a rather handsome species of matted growth. The flowering heads are rather dense, and their outer bracts are pale pink. Antennaria isolepis Greene Stewart Plateau: Keno Summit, in gravelly openings in alpine tundra, elevation 6,000 ft, No. 1034. Ogilvie Mountains: River flats along Dempster Highway, Mile 57, Nos. 133 and 234; Canyon Creek east of Mile 48, No. 450. Subarctic North America from northern La- brador to central Alaska, northward to or slightly beyond the limit of trees. Antennaria nitida Greene Vicinity of Mayo, 63°35’N, 135°45’W; com- mon on gravelly river terraces, Nos. 694A, 930, and 1182; Midnight Dome near Dawson, No. 1410A; Mount Gray northeast of White- horse, in alpine heath, No. 1345. A western prairie and grassland species, reaching southern Mackenzie District and southern half of Yukon. Antennaria rosea (Eaton) Greene Minto Road north of Mayo, 63°41’N, 135° 51’/W, open aspen woods, Nos. 650 and 913; Midnight Dome near Dawson, 64°04’N, 139° 42’W, on rocky ledge, No. 1410. Wide-ranging western species, reaching southern Macken- zie District and southern half of Yukon, with a disjunct population in southern James Bay. Compositae Antennaria subviscosa Fern. Vicinity of Mayo, 63°35’/N, 135°45’W, common on gravelly river terraces, Nos. 488, 490, and 522. The series match perfectly the plant re- ported from southeastern Yukon (Porsild 1951a), and, like it, differ from A. subviscosa in Gaspé, Quebec, mainly in the smooth rather than papillose achenes. Since 1951, the 1500-mile gap between Gaspé and the nearest western station of A. subviscosa has been bridged by collections from the north shore of Lake Superior (Porsild 1966: map 141). Arnica alpina (L.) Olin ssp. attenuata (Greene) Maguire Ogilvie Mountains: Alpine calcareous tundra along Dempster Highway, Mile 81, Nos. 1471, 1478, and 1503; river flats and terraces, Mile 84, Nos. 1680 and 1688; lower slope of lime- stone hills south of Mile 110, A.E. & R.T. Por- sild, Nos. 23,675—76. This is apparently by far the most common Yukon Arnica, and it is rarely absent on well-drained calcareous soils of the central Yukon Plateau. Arnica diversifolia Greene St. Elias Mountains in southwestern Yukon, in valley south of Serpentine Creek, eleva- tion 4,000 ft, A.M. Pearson, No. 67-337. New to the flora of Yukon. This species has been reported from Alas- ka by Maguire (1943:475), without, however, exact locality. In the National Herbarium of Canada there is a fine specimen of A. diversi- folia from the Alaska Range: Tangle Lake, ap- proximately 63°N, 146°W, O. Gjaerevoll, No. 1315, reported by him as A. mollis (1967:48). Arnica frigida C.A. Mey. A. louiseana Farr. ssp. frigida (Meyer ex Iljin) Maguire Keno Hill, 63°58’N, 135°42’W, alpine tundra, elevation 5,500 ft, No. 729. Ogilvie Mountains: In alpine tundra on river flat along Dempster Highway, Miles 51-52, No. 198. Apparently rare and alpine in central Yukon (Porsild 1966: map 144). Arnica Lessingii Greene Mount Haldane, 63°51’N, 135°50’W, edge of snowflush in main cirque, elevation 5,500 ft, No. 595; Keno Summit, 63°58’N, 135°42’W, in alpine tundra, elevation 5,500-6,000 ft, Nos. 733 and 1287. Ogilvie Mountains: Alpine tun- dra, elevation 4,200 ft, along Dempster High- way, Mile 51, No. 195; Miles 55-56, No. 5; alpine tundra on limestone hill south of Mile 110, A.E. & R.T. Porsild, No. 23,674. A chro- mosome count based on No. 1287 gave 2n=57 (Mulligan and Porsild 1969). Arnica mollis Hook. This species was first reported from Yukon (Porsild 1951a:329) from the upper Rose River in southeastern Yukon; a second and more recent collection from southwestern Yukon came from Serpentine Creek, A.M. Pearson, No. 67-338A. Hultén (1967:141) stated that a duplicate in Stockholm of the Rose River specimen cited above (A. mollis, A.E. Porsild and Breitung, No. 10,583) was misidentified, and is actually A. amplexicaulis. A re-examination of the specimen in Ottawa confirmed, however, that itis a perfectly good A. mollis, and that, fur- thermore, the determination was verified in 1947 by Dr. Bassett Maguire. Artemisia alaskana Rydb. Ogilvie Mountains: Gravel terrace on moun- tain east of Dempster Highway, Miles 57-58, elevation 4,800 ft, No. 265. This is a slight northward extension of the Yukon range shown by Porsild (1966: map 145). Artemisia arctica Less. Stewart Plateau: ‘‘Keno Hill Dome’, com- mon in alpine herbmats, Nos. 971, 995, and 1273. A chromosome count of 2n= 18 was ob- tained from No. 1273 (Mulligan and Porsild 1969). Ogilvie Mountains: Dempster Highway, common on gravelly river flats, Mile 57, No. 369; Dempster Highway, Mile 86, No. 1651; rocky tundra on limestone hills south of Mile 110, A.E. & R.T. Porsild, No. 23,677. Richard- son Mountains, 13 miles northeast of Lapierre House, Youngman & Tessier, No. 599. Com- mon everywhere in moist alpine herbmats. Aster pansus (Blake) Cronq. A. commutatus sensu Hult., Fl. Alaska & Neighboring Territories. 860 (1968), in part only, not as to illustration and not (Torr. & Gray) A. Gray. A. elegantulus sensu Hult., ibid., non Porsild, Nat. Mus. Can. Bull. 121: 297-98; PI. 28, figs. 1 and 2 (1951). Richthofen Creek on west shore of Lake La- berge, growing in dense masses on wet clay banks subject to spring flooding, 3 Aug. 1969, No. 1947; Mile 985 on Alaska Highway east of 65 Annotated List of Selected Species Haines Junction, 60°48’N, 136°45’W, Calder & Kukkonen, No. 28,250 (DAO). Stems one to several, clustered, from a short, erect rootstock; the entire plant glaucous-green and rough-hairy from short, stiff hairs; leaves small and narrow, very numerous; inflorescence racemose, rather dense, with numerous spreading branches, one- to several-headed; phyllaries in 3 series, of unequal length, prominently ciliate, their tips spreading. Aster pansus is a wide-ranging western grassland species, until now known in Cana- da only from western Alberta; however, it is also common locally on banks of the Macken- zie River, between latitudes 62°N and 64°N; opposite Simpson, Raup & Soper, No. 9964; willow thickets and sand banks west of Fort Simpson, jidem, No. 9866; shelving river banks below Fort Good Hope, Wynne- Edwards, Nos. 8580 and 8616; same place, A.E. Porsild, No. 16,743. The series cited above are all in the National Herbarium of Canada, and were tentatively referred to Aster falcatus by Raup (1947); A. falcatus, however, as now understood, has a horizon- tally soreading rhizome, whereas the rhizome of A. pansus is short and erect, as is the case with all the specimens cited above. Hultén’s illustration (1968:860) of what he considers Aster commutatus was obviously drawn from an isotype of A. elegantulus Por- sild (A.E. Porsild & Breitung, No. 10,703, from Whitehorse) in the Botanical Museum, Stock- holm. Hultén noted (1967:129) that A. elegan- tulus ‘‘agrees well with A. commutatus (T. & G.) A. Gray’’, and also cited two “other” spec- imens said to be A. commutatus. The first, Calder & Kukkonen, No. 28,250 (cited above) is definitely A. pansus; the second, Calder & Kukkonen, No. 28,087, from Von Wilczek Lakes near Minto on the Yukon River, 62° 42/N, 136°41’W (both in DAO), definitely is not A. pansus but is a perfect match for A. elegantulus, but not for A. commutatus, as represented in the National Herbarium of Canada from grasslands of Saskatchewan and Alberta. In A. elegantulus the stems are slender, 30- 40 cm tall, one or several together from a short, compact base. The entire plant is fresh green, not nearly so leafy as A. pansus, and the stems and leaves are sparingly downy- pubescent. The inflorescence is open, race- mose-paniculate, the very slender branches 66 3-6 cm long, each bearing one large terminal head and 1-3 smaller lateral heads. Chrysanthemum integrifolium Richards. Ogilvie Mountains: Common in alpine tundra on limestone hills, ascending to at least 5,000 ft, along Dempster Highway, Miles 81-83, Nos. 1097, 1480, 1526, and 1617. An arctic- alpine and obligate limestone plant, also noted as common in rocky tundra near Mile 110. In Yukon thus far known only from a few stations. Crepis nana Richards. Ogilvie Mountains: Moist sandbars and river flats along Dempster Highway, Miles 56-59, No. 126; near Mile 67, No. 1809, and in similar places near Mile 110, A.E. & R.T. Porsild, No. 23,679. Erigeron compositus Pursh Because of its preference for sandy, non- acid soils, the distribution of E. compositus in Yukon, as elsewhere, is spotty. It was not seen in the southern and central Ogilvie Mountains, but has been collected in the Por- cupine River basin. No. 550, on dredge tailings near Dawson, differs from typical E. compositus by its caes- pitose habit and long, narrowly cuneate leaves, terminating in 3 shallow lobes and by its pale-purple or pink pappus. It may be a hybrid: E. purpuratus X EE. compositus. Achenes apparently well formed, but appear abortive. Similar, but fertile, plants are known to me from the Alberta Rocky Mountains: Bow River Pass, on shaly slope, elevation 7,200 ft, A.E. Porsild & Breitung, No. 14,938; Bow Pass above Peyto Lake, A.E. Porsild, No. 18,136; Brazeau National Forest, alpine ridges in moist sand, elevation 7,100 ft, idem, No. 20,791. In No. 20,791 the pappus is white as are the ligules. In habit all look strikingly different from E. compositus. The Rocky Mountain plant is clearly E. trifidus Hook. (Hooker 1834:17, pl. 120). Erigeron elatus Greene Ogilvie Mountains: Moist tundra along Demp- ster Highway, Mile 82, elevation 4,200 ft, No. 1117; Bell River near Lapierre House, Young- man & Tessier, No. 264. New northern limit for Yukon. Compositae Erigeron glabellus Nutt. ssp. pubescens (Hook.) Cronq. Gravelly hillsides, “Steamboat Slough’, Yu- kon Territory, M.W. Gorman, No. 108; near mouth of Lewis River, 62°N, 18 Aug. 1887, G. Dawson, CAN No. 11,143; flats along Rink Rapid, Yukon River, 19 July 1902, John Ma- coun, CAN No. 78,994; McQuesten area, river flat one mile east of ‘‘Pelly Farm’, J.D. Camp- bell, Nos. 59 and 78, flowering on 14 June 1948; seven miles upstream from junction of Pelly and Yukon rivers, Mickey Lammers, July 1965, CAN No. 289,652; Minto Road, Mile 13 north of Mayo, lake banks, No. 1251; vici- nity of Mayo, 63°35/N, 135°45’W, 4 miles east of Stewart Crossing, No. 1389. The Yukon plant is aclose match for the plant of western Canada and adjacent United States. In Yukon it is well established along highways and townsites, near existing and former gold- mining camps, and was most likely intro- duced with shipments of baled hay in the early years of gold mining. Erigeron grandiflorus Hook. ssp. arcticus n. ssp. Differt a ssp. grandiflorus caulis 15-25 cm altis, foliis radicalibus oblanceolatis anguste petiolatis usque ad 5-6 cm longis; capitulus 2.5-3.5 cm latus; pedunculus infra capitalum phyllariesque glandibus destitutis. | have noted (Porsild 1955) that the plant reported as E. grandiflorus from Alaska, the Yukon, the Arctic Coast of the Mackenzie District, and the western islands of the Cana- dian Arctic Archipelago differs consistently from E. grandiflorus Hook. of the Alberta Rocky Mountains (Hooker 1834:18, pl. 123), where it was long known only from the type locality, at approximate latitude 53°30’N. The Rocky Mountain plant is well illustrated in Hultén (1968:865), where the figure was drawn from a Rocky Mountain specimen from southern Alberta (A.E. Porsild & Lid, No. 19,314). The arctic plant for which ssp. arc- ticus is now proposed was tentatively de- scribed and illustrated (Porsild 1955:181-82; Pl. 22, figs. 1 and 2). The type (CAN) is A.E. Porsild, No. 17,342, Victoria Island, southwest coast, vicinity of Holman Island trading post, 8 Aug. 1949. In the Yukon E. grandiflorus ssp. arcticus is known from mountains in the south and southwest and from the Arctic Coast. The fol- lowing are unreported. Ogilvie Mountains: Mountain east of Dempster Highway, Miles 57-58, in grassy ravine, No. 252. Richardson Mountains: McDougall Pass, Horn Lake, ele- vation 2,100 ft, Youngman & Tessier, No. 75. A gap of about 1,000 miles separates the areas occupied by ssp. grandiflorus and ssp. arcticus. Erigeron humilis Grah. E. unalaschkensis (DC.) Vierh. Keno Summit, elevation 5,500-6,000 ft, 63° 58’N, 135°42’W, very common in rich alpine herbmats and snowbed slopes, Nos. 717, 747, and 981. Ogilvie Mountains: Mountains east of Dempster Highway, Miles 50-57, Nos. 90 and 249; damp limestone ledges, Mile 83, elevation 3,800 ft, No. 1516. Erigeron hyperboreus Greene E. alaskanus Cronq., Brittonia 6(2):228 (1947). E. grandiflorus sensu Porsild, Rhodora 41 :292 (1939) non Hook. Long known only from the type locality on Porcupine River in northeastern Alaska, E. hyperboreus is now known from a number of stations along the Brooks Range of northern Alaska to the Seward Peninsula and to Pas- tolik, north of the Yukon Delta. Thus far it has been collected in the Yukon only twice, at 65°18’N, 141°00’W (D.D. Cairnes, No. 83,056), and on the lower Firth River south of Herschel Island (June McNeish, CAN No. 236,767). In habit E. hyperboreus is somewhat similar to E. yukonensis Rydb., described from Daw- son, but differs, among other aspects, by the hairs on the involucrum and especially by the hairs on the tip of the phyllaries having pur- plish-black cross-walls. In E. yukonensis and in E. grandiflorus the cross-walls are clear and colourless. Erigeron linearis (Hook.) Piper Diplopappus linearis Hook. Southwestern Yukon: Mountain slope on east side of ‘Big Arm’’, Kluane Lake, approximate- ly 61°33/N, 138°40’W, 20 Aug. 1944, H.M. & L.G. Raup, No. 12,918, which is a first record for Yukon. The occurrence in southwestern Yukon of E. linearis, disjunct by more than 1,200 miles from the northernmost part of its main range in southwestern British Columbia and Wash- ington, is most unexpected. Erigeron lonchophylius Hook. Haines Road, subalpine slopes south of Deza- deash River, A.E. & R.T. Porsild, No. 22,321; 67 Annotated List of Selected Species McQuesten area, Bear Creek, J.D. Campbell, No. 286; vicinity of Minto Bridge, 63°40’N, 135°51’W, No. 648. Ogilvie Mountains: River flats along Dempster Highway, Mile 57, No. 219; moist tundra and gravelly places along Miles 81-83, No. 1603. The last is a new north- ern limit for Yukon. Erigeron peregrinus (Pursh) Greene ssp. callianthemus (Greene) Cronq. Tincup Lake in southwestern Yukon, approxi- mately 61°50’/N, 139°40’W, on a rockslide above the lake, No. 1986. A cordilleran race, shown from one station in southern-central Alaska and from another in southwestern Yukon on the Alaska High- way near the Alaska border (Hultén 1968:869). Erigeron purpuratus Greene Ogilvie Mountains: Solifluction stripes in al- pine tundra, elevation 4,200 ft, east of Demp- ster Highway, Miles 44-46, Nos. 1456-58; gravelly slopes of mountain east of Miles 50-51, elevation 4,200-4,400 ft, No. 1583; gravel ridges along river, Mile 56, Nos. 49 and 278; sandy river flat near Mile 67, No. 1802. Common locally and forming loose mats from a central taproot from the crown of which issue slender, spreading, and leafy shoots up to 10 cm long; the leaves oblanceo- late, 1-2 cm long, the blade 4-5 mm wide, entire or very rarely shallowly toothed near the apex, gradually tapering into a slender petiole; scapes very slender, 3-5 cm long; heads solitary, small, about 1.5 cm in diame- ter; involucrum viscid under the dense cover of tangled, flat hairs; ligules varying from pure white to pale purple; pappus pink; leaves and scapes viscid under the villous spreading hairs. A chromosome count of 2n=18 was obtained from No. 1802 (Mulligan and Por- sild 1970). An alpine species endemic in mountains of Alaska, southwestern Yukon, and northern- most British Columbia, closely related to E. pallens Crong. (Cronquist 1947:240). The type of E. pallens came from Mount MacDonald near Glacier, British Columbia. This species is common locally on alpine slide-rock slopes in Banff National Park, Alberta (A.E. Porsild & Breitung, Nos. 13,671, 14,018, 14,188, 14,252— 53, 14,577, and 15,941). It differs from E. pur- puratus mainly by the colour of its pappus and ligules, which are yellowish white instead of pale purple. 68 Erigeron yukonensis Rydb., Bull. N.Y. Bot. Gard. 2(6):185 (1901); Crongq., Brittonia 6(2): 227 (1947); Hult., Fl. Alaska & Yukon 10:1510 (1950) pro max. pte. E. glabellus Nutt. ssp. pubescens (Hook.) Cronq. var. yukonensis (Rydb.) Hult., Ark. Bot., ser. 2, vol. 7(1):132 (1967). E. grandiflorus Porsild, Sargentia 4(1):68 (1943) non Hook. In addition to the Yukon specimens cited by Cronquist (1947), E. yukonensis is now also known from the Ogilvie Mountains: Dempster Highway, Mile 83, on calcareous, stony tun- dra, No. 1507; northern Northwest Territories from Dolomite Lake east of upper Mackenzie Delta, 68°39’N, 133°62’W, Dorothy Swales, No. 522; Arctic Coast, Nicholson Island, ap- proximately 70°N, 129°W, A.E. & R.T. Porsild, No. 2950; Tree River, Coronation Gulf, R.E. Miller, No. 165. A somewhat critical species that has been confused with, or misidentified as, Aster yukonensis, Erigeron grandiflorus, and E. hyperboreus. |n general habit it does resem- ble E. hyperboreus, which, however, appears to be restricted to Alaska. Furthermore, as Cronquist pointed out, in E. hyperboreus the hairs on the involucre have black or dark- purple cross-walls, whereas in E. yukonensis and E. grandiflorus the cross-walls are clear white. |! am unable to follow Hultén in reducing E. yukonensis to a variety of the biennial E. glabellus ssp. pubescens. In his monograph, Cronquist (1947) even places E. glabellus ssp. pubescens in a different subsection of the genus. Gnaphalium uliginosum L. Stewart Plateau: Minto Road, 13 miles north of Mayo, in dry gravel, Nos. 915, 1183, and 1292. An introduced annual, in Yukon col- lected at Mayo as early as 1944 (J.P. Ander- son, No. 9740), and now apparently well established and spreading. Hieracium gracile Hook. Mount Haldane, 63°51’N, 135°30’W, near road in bottom of main cirque, elevation 4,300 ft, No. 681; Galena Hill near Calumet and Elsa mines, openings in willow thicket, No. 1172; “Highlet Gulch’, tributary to Minto Creek, No. 851; Keno Summit, herbmat slopes, ele- vation 5,500 ft, No. 1038. Not seen in the Ogil- vie Mountains. Compositae Saussurea angustifolia DC. var. angustifolia Ogilvie Mountains: Limestone hills west of Dempster Highway, Miles 81-82, 64°58’N, ele- vation 4,200 ft, in turfy places, Nos. 1087, 1130, 1468, and 1546. No. 1130 provided a chromo- some count of 2n=52 (Mulligan and Porsild 1969), and No. 1546 was 45 cm tall; moist, stony floodplain below limestone hills south of Mile 110, A.E. & R.T. Porsild, No. 23,680. Saussurea angustifolia DC. var. yukonensis Porsild, Nat. Mus. Can. Bull. 121:337 (1951). Keno Summit, 63°58’N, 135°42’W, herbmat slopes, elevation 5,500 ft, No. 992; in moist alpine heath, elevation 5,500-6,000 ft, No. 1228. Ogilvie Mountains: Damp river meadow along Dempster Highway, Mile 67, No. 1811. The var. yukonensis differs from typical S. angustifolia mainly by its dwarf habit; the leaves are usually broader and often with sinuate-dentate margins, but they are never viscid, and the underside may be glabrous or floccose as in S. angustifolia. | therefore see no reason for placing the var. yukonensis under S. viscida Hult., as was done by Hultén (1950, 1968). Senecio conterminus Greenm. Ogilvie Mountains: On gravelly slope of moun- tain east of Mile 57 on Dempster Highway, No. 247; river flats near Mile 60, No. 73. S. conterminus and S. resedifolius both have glabrous achenes, but in the latter the stems, and often other parts, are strongly tinged with purple; S. conterminus is never purplish- tinged, but is grey-green and distinctly floc- cose-tomentose, especially in the axils of the leaves, but often also on the petioles and flowering stems. A southern population of Alberta and British Columbia is isolated from that of the Yukon and Alaska by a gap of 1,200 miles (Porsild 1966:60, map 155). Senecio indecorus Greene S. pauciflorus Pursh var. fallax sensu Hult., FI. Alaska & Neighboring Territories. 930 (1968). Minto Road, 13 miles north of Mayo, No. 902. A peculiar variation, with slender stems, 6 dm tall, branching irregularly, some from below the middle, with only discoid heads. Senecio Kjellmanii Porsild, Rhodora 41:299 (1939). Cineria frigida Richards. f. tomentosa Kjellm., Vega Exped. Vetensk. lakttag. 2:13, pl. 1 (1883) non Senecio tomentosus Michx. S. atropurpureus (Ledeb.) Fedtsch. ssp. to- mentosus (Kjellm.) Hult., Ark. Bot., ser. 2, vol. 7(1):141 (1967). The holotype of this species came from St. Lawrence Island in Bering Strait and is a very striking plant, well illustrated in Kjellman (1883: pl. 1); it differs strikingly from S. atro- purpureus (Ledeb.) Fedtsch. by its dense, brownish-yellow and long-woolly indument that completely covers the upper half of the stem, the uppermost cauline leaves, and the phyllaries of the large, solitary head from 2-4 cm in diameter and 2-3 cm high; the blade of the basal leaves, unlike that of S. atropur- pureus, is broadly lanceolate and coarsely dentate, and the rootstock is very stout, ascending-erect, and rarely branched. In 1939 | had seen only a few specimens of S. Kjel/manii from Alaska and northern Yukon that fully matched the type. Numerous addi- tional collections are now Known from north- ern Alaska and the Yukon, and from alpine situations in central Alaska, southwestern and central Yukon, and the Richardson Moun- tains, Northwest Territories. Arctic Slope, Alaska: 69°20’N, 145°00’W, Spetzman, Nos. 618 and 1479. Central Alaska: Eagle Summit 105 miles north of Fairbanks, Galen Smith, No. 2359; Ketchumstuk Mountain west of Chicken, Spetzman, No. 5169; Porcupine Dome, elevation 3,450 ft, Gjaerevoll, No. 1146. Southwestern Yukon: “High Pass” north of “Bridge Creek’, 61°35’N, 138°50’W, H.M. Raup, et al., No. 13,849; Amphitheatre Moun- tain 16 miles west of Burwash Landing, above timberline, elevation 7,000 ft, C.H.D. Clarke, No. 284; same area, Gladstone Creek, Ruby Range, James A. Neilson, No. 1289; Burwash Creek, 61°30’N, 139°17’W, alpine meadow, elevation 4,500 ft, A.M. Pearson, No. 70-5. Central Yukon: Ogilvie Mountains east of Miles 57-58 on Dempster Highway, No. 406. Northern Yukon: British Mountains, Mount Sedgwick, 68°58’N, 139°01’W, A.M. Pearson, No. 70-24B. Mackenzie District: Richardson Mountains west of Mackenzie Delta, A.E. Por- sild, No. 6722. The above series differs strikingly from typ- ical S. atropurpureus (Ledeb.) Fedtsch., but throughout the range of S. Kjel/manii, speci- mens do occur that resemble that species in general habit, but lack the dense, yellowish- brown indument. In view of the considerable polymorphy within this plastic group, the taxonomic status of S. Kjellmanii may be un- certain; however, the proposed change to 69 Annotated List of Selected Species ssp. tomentosus n. stat. (Kjellm.) Hult. is ille- gal because of S. tomentosus Michx. of the coastal plains of the eastern United States. Senecio Lindstroemii (Ostf.) Porsild S. fuscatus sensu Hult., Fl. Alaska & Neigh- boring Territories. 927 (1968) non (Jord. & Fourr.) Hayek. See Porsild 1966:60-—61 for a discussion of identity and distribution of the North American plant. Ogilvie Mountains: On mountain east of Dempster Highway, Miles 51—52, on turfy al- pine slopes, elevation 4,500-—6,500 ft, Nos. 37, 216, and 1581; same place, Youngman & Tes- sier, No. 350. Apparently common locally, but in Yukon thus far reported only from the Arc- tic Coast and the St. Elias Mountains. Senecio resedifolius Less. McQuesten area, east slope of White Moun- tain, elevation 4,800 ft, on talus slope, J.D. Campbell, No. 190; south peak of “Little S. Klondike Syenite Mt.”, elevation 6,100 ft, idem, No. 699. Ogilvie Mountains: East of Dempster Highway, Miles 57-58, elevation 4,200-6,000 ft, No. 245; dry, calcareous tun- dra, near Mile 81, No. 1483; limestone hill west of Mile 82 on rocky ledge, No. 1102; damp tundra below limestone hill south of Mile 110, A.E. & R.T. Porsild, No. 23,681. A dwarf species usually with stems 10-15 cm tall, but in No. 245 several are as tall as 25 cm. Readily distinguished from S. conter- minus by its deep purplish-tinged phyllaries, glabrous stems, and basal leaves. Senecio triangularis Hook. Stewart Plateau: Mount Haldane, near snow- beds along creek, elevation 5,500 ft, No. 680. “Highlet Gulch’, tributary to Minto Creek, in damp places by stream, No. 827. Ogilvie Mountains: River flats along Dempster High- way, Miles 51-52, No. 277; on snowflushes west of Mile 58, elevation 3,800 ft, No. 486. A cordilleran—Pacific Coast species of alpine snowbed habitats, probably always growing on soils derived from neutral or acidic rocks. The present collection extends its known Yukon range beyond the Stewart Plateau. As yet not reported from mountains of central Alaska, although it is apparently common in alpine situations along the Pacific Coast of Alaska as far as Kodiak Island. 70 Senecio yukonensis Porsild Keno Hill, 63°58’N, 135°42’W, common in al- pine meadows and on snowbed slopes, eleva- tion 5,500-6,000 ft, Nos. 539, 740, 1075, and 1272; same place, A.E. & R.T. Porsild, Nos. 23,576-77; Mount Haldane, bottom of main cirque, elevation 5,500 ft, No. 686. Ogilvie Mountains: Damp alpine meadows near Dempster Highway, Miles 55-56, No. 3; among willows near Mile 81, No. 1548. Apparently common in damp alpine meadows of central and northern Yukon and Alaska, but thus far not reported from the St. Elias Mountains. Taraxacum alaskanum Rydb. Dwarf species, in habit similar to T. sibiricum Dahlst., from which, however, itis at once dis- tinguished by its runcinate-pinnatifid leaves in which the lobes are opposite and entire- margined, the terminal lobe spear-shaped and larger than the lateral. In T. sibiricum the leaves are narrowly pinnatifid, and the lobes, and often also the rachis, have one or more small but sharp teeth. Arctic-alpine: Central Yukon: Canol Road, Miles 10-14, elevation 4,500-6,000 ft, A.E. Por- sild & Breitung, No. 11,018; Rose River Pass, elevation 4,000-6,000 ft, iidem, No. 10,938; upper Rose River, iidem, Nos. 10,529 and 11,945. Southwestern Yukon: Vicinity of Mac- kintosh, Mile 1022 on the Alaska Highway, Schofield & Crum, No. 8070. General Range: Amphi-Beringian; east along the Arctic Coast of Alaska to 125°W; south in the Richardson and Mackenzie mountains, in British Columbia slightly be- yond 60°N, with a disjunct station in 49°04’N, 120°12’W, but apparently absent in Alberta. Taraxacum carneocoloratum A. Nels., Amer. J. Bot. 32:290 (1945). Ogilvie Mountains: Common locally on grav- elly riverbanks along Dempster Highway, Miles 55-66, elevation 4,200 ft, flowering on 25 June 1966, No. 50; gravelly river flats and meadows, Miles 67-69, Nos. 1597, 1461, and 1789. A chromosome count from seedlings grown from No. 1789 gave 2n=32 (Mulligan and Porsild 1970). An alpine species, until now Known only from the type locality in Mount McKinley Na- tional Park, Alaska. It differs strikingly not only from other members of the section Arc- tica, but from all arctic and boreal species of the genus by its ligules than are deep pink or flesh-coloured in life, turning dark purple in Compositae drying. In our specimens the fruiting scapes are up to 16 cm tall. The involucral bracts are in three distinct rows, the first and second broadly ovate, with a pale-green centre and much paler margins; those of the third and upper row are dark green, gradually tapering to a truncate and scarcely dilated point. The body of the ripe achene is straw-coloured, about 5 mm long, broadest in the middle and spiny in the upper half. New to the flora of Canada. Taraxacum dumetorum Greene In Yukon Territory thus far collected only a few times in mountains of the southern parts (Porsild 1951a:338).-. Taraxacum lacerum Greene Apparently common in open woods, on river- banks and lakeshores, and lately also in dis- turbed soil along roads; north to the Arctic Coast. Taraxacum mackenziense n. sp. Planta mediocris ca. 10-30 cm alta; folia oblanceolata vel lingulata, vix 1.0 cm lIata, brevissime dentata vel integra; petiolis an- gustis, anguste-alatis, sat purpureis; involu- crum mediocre vel sat parvum, 10-18 mm longum, vulgo sat laete viridi; squamae in- volucri exteriores erecto-patentes, ovato- lanceolatae, anguste albido-marginatae, sub apice corniculus usque ad 2 mm longis mu- nitae, interiores corniculus minoribus; ligulae marginales albis, extus dilute roseo-purpu- reae ornatae; antherae parce polliniferae; achenium olivaceo-stramineum, 4 mm lon- gum, superne valde muricato-tuberculatum, rostro ca. 8 mm longo. By its narrow or subentire leaves, small heads with light-green involucrum, and white ligules that are faintly purplish on the outside, T. mackenziense differs from all other Taraxa- ca known to me. Thus far it, has been col- lected only a few times and only on the east branch of the Mackenzie Delta, by dolomite quarry near Inuvik, with mature achenes on 7 July 1965, Dorothy E. Swales, CAN No. 291,036 (Type: CAN), Plate 13; Richards Island 69°32’/N, 133°45’W, on a sandy beach, G.W. Scotter, No. 10,129 (CAN No. 300,244); Mackenzie River, Bear Rock, 64°55/N, 125° 42’W, A.E. & R.T. Porsild, No. 3399 (CAN No. 111,694). Although thus far known only from the lower Mackenzie River and the Mackenzie Delta, 7. mackenziense should be looked for in the Yukon Territory, in damp, calcareous habitats. Taraxacum maurolepium Hagl., Svensk Bot. Tidskr. 43(1):111, figs. 2 and 5C (1949). Ogilvie Mountains: River flats along Dempster Highway, Mile 57, elevation 4,200 ft, No. 242. Our specimens are with young, almost mature fruits, 5 mm long and slate-grey on 19 July 1966, and match the description closely. The type is from Umiat on the Colville River in northern Alaska, E. Lepage, No. 23,784, a duplicate of which | have examined (CAN No. 204,344). Judging from the pale-green colour of the elongated leaves, up to 25 cmin length, the type specimen must have grown in partly shaded areas, among grasses or wil- lows. In our No. 242, which grew in the open, the leaves are one-third as long, dark green and of firmer texture, and incidentally are a close match for Spetzman, No. 1379, from Umiat, Alaska, cited by Hultén (1950). New to the flora of Canada. Taraxacum mitratum Hagl., Svensk Bot. Tidskr. 40(4):350, figs. 13 and 14 (1946). Ogilvie Mountains: River flats along Demp- ster Highway, Miles 55-56, elevation 4,200 ft, No. 128. The large material represented in No. 128 is very uniform and agrees rather closely with the description and illustrations of T. mitra- tum from the Katmai region, Alaska Penin- sula, of which the type is A.E. Miller (US No. 1,072,536), as well as with specimens from Hope, J.P. Anderson, No. 6596, determined by Haglund. Our specimens were collected on 5 July 1966; among the two dozen specimens, some heads were already in the post-floral state, while several had almost mature achenes. In No. 128 the somewhat shrivelled ligules are now creamy white; the involucral bracts are in three distinct series, all uniformly dark green, with prominent appendages; those of the first row are ovate and almost as broad as they are long; the achenes are 5 mm long, and spiny nearly to the base. The collection is the first from Canada. Taraxacum multesinum Hagl., Svensk Bot. Tidskr. 43(2):313, fig. 11 (1951). Known to Haglund only from the type: Alaska, Moose Pass on Kenai Peninsula, along rail- road, J.P. Anderson, No. 6490 (S and CAN). 71 Plate 13 Taraxacum mackenziense n. sp. (Type) Approximately natural size "2 Compositae Another specimen that closely matches the type was collected in southwestern Yukon, in the vicinity of Mackintosh, elevation 4,000 ft, near Mile 1022 on the Alaska Highway, Scho- field & Crum, No. 8156. In the Yukon speci- men, mature achenes are reddish yellow, spiny in the upper part, the body about 3.0 mm long or about half as long as the beak. Taraxacum pellianum Porsild, Can. Field- Natur. 64:44 (1950); Porsild, Nat. Mus. Can. Bull 121:339, pl. 38 (1951). Ogilvie Mountains: In stony, alpine herbmats on mountain east of Dempster Highway, Miles 50-54, elevation 6,000 ft, No. 29. Our speci- mens perfectly match-the type, which came from the Pelly Mountains in southeastern Yukon. T. pellianum is now also known from southwestern Yukon (Raup, No. 13,078) and from the lower Mackenzie River and the Mac- kenzie Delta (Porsild and Cody 1968). Taraxacum phymatocarpum J. Vahl In Yukon Territory long Known only from Herschel Island on the Arctic Coast. An un- reported collection from McDougall Pass in the Richardson Mountains, Northwest Terri- tories, suggests that 7. phymatocarpum may also turn up in the mountains of northern Yukon Territory. Taraxacum sibiricum Dahlst. T. alaskanum sensu Porsild, Nat. Mus. Can. Bull. 121:338 (1951) pro max. pte. Central Yukon: Stewart Plateau, Keno Hill, alpine herbmats, elevation 6,065 ft, No. 516; same place, on south-facing rich herbmat slope, elevation 5,500 ft, Nos. 768 and 996. Ogilvie Mountains: River flats along Dempster Highway, Mile 57, elevation 4,200 ft, No. 105; on snowflushes of mountain east of Miles 50-54, elevation 4,500 ft, No. 170; Canyon Creek, near Mile 48, No. 449. Arctic-alpine dwarf species of similar habit and habitats as 7. alaskanum, from which, however, it is easily distinguished by the dif- ferent cut of its leaves (see T. alaskanum). Long known only from the type locality at the mouth of the Lena River in central Siberia, T. sibiricum is now known to be wide-ranging in North America from the Seward Peninsula, Alaska, east to the Mackenzie Mountains, Northwest Territories, and south, again with large gaps, to mountains of interior British Columbia; in Alberta south to latitude 49°N. 73 ins we ‘ Ps { 1 Ih herd YT >| : een Ata ot 1 ia we ee i 2arresing , I i Lah eoveqiial = " i ei es A yy 5 ot Ge. te) se = * 7 ' mad} WH ‘ ™ i? ten Ais ‘ 7 fri r 7 a Hy | .. a cr py OL Pores, ’ Spee “oT eg hi ah set er ariehtyer of al Duditel OF ra AAA Ae Bibliography Adams, M. F. (1817). Descriptiones plantarum minus cognitarum Sibiriae praesertim orientalis quas in itinere ann. 1805 et 1806 observavit. Moscou Soc. Nat. Mem. 5:89-116. Anderson, J. P. (1944). Flora of Alaska and adjacent parts of Canada. lowa State University Press, Ames, lowa. Bocher, Tyge W., Kjeld Holmen, and Knud Jakobsen (1968). The flora of Greenland, Transl. By T. T. Elkington and M. C. Lewis, Haase, Copenhagen. Boraiah, G. and Margaret Heimburger (1964). Cytotaxonomic studies on New World Ane- mone (Section Eriocephalus) with woody root- stocks. Can. J. Bot. 42 (7): 891-922. Bostock, H. S. (1966). Notes on glaciation in central Yukon Terri- tory. Geol. Surv. Can. Pap. 65-36. Britton, N. L., and P. A. Rydberg (1901). An enumeration of the flowering plants col- lected by R. S. Williams and by J. B. Tarleton. Bull. N.Y. Bot. Gard. 2 (6): 149-87. Busch, N. A. (1913-31). Flora Sibiriae et Orientis extremi 25: 304-05. Akad. Nauk Bot. Muzei, Leningrad. Cody, W. J. (1971). A phytogeographic study of the floras of the continental Northwest Territories and Yukon. Natur. Can. 98 (2): 145-58. Cody, W. J., and A. E. Porsild (1968). Additions to the flora of continental North- west Territories, Canada. Can. Field-Natur. 82 (4): 263-75. Cronquist, Arthur (1947). Revision of the North American species of Erigeron, north of Mexico. Brittonia 6 (2): 121-300. Dansereau, P. M., and E. E. Steiner (1956). Studies in Potentillae of high latitudes and altitudes. 2, Central Baffin Island populations. Bull. Torrey Bot. Club 83 (2): 113-35. Drury, W. H. (1956). Bog flats and physiographic processes in the upper Kuskokwim River region, Alaska. Con- trib. Gray Herb., no. 178. Ekman, Elizabeth (1931). Contribution to the Draba flora of Green- land. Svensk Bot. Tidskr. 25 (4): 465-94. Fernald, M. L. (1914). The North American representative of Arenaria ciliata. Rhodora 16: 43-44. (1919). The American representatives of Arenaria sajanensis. Rhodora 21: 12-17. (1924). Juncus Triglumis and its American repre- sentative. Rhodora 26: 201-03. (1939). New species, varieties and transfers. Rho- dora 41: 423-61. Gelert, Otto (1898). Notes on arctic plants. Bot. Tidsskr. 21 (3): 287-318. Gjaerevoll, Olav (1956). Poa porsildii, a new species from Alaska- Yukon. Kgl. Nor. Vidensk. Selsk. Forh. 29 (16): 72-75. (1958). Botanical investigations in central Alaska, especially in the White Mountains, Pt. 1, Pterido- phytes and Monocotyledones. Kg!. Nor. Vidensk. Selsk. Skr., no. 5. (1963). Botanical investigations in central Alaska, especially in White Mts. Pt. 2, Dicotyledones Sali- caceae—Umbelliferae. Kgl. Nor. Vidensk. Selsk. Skr., no. 4. (1967). Botanical investigations in central Alaska, especially in the White Mts. Pt. 3, Synpetalae. Kgl. Nor. Vidensk. Selsk. Skr., no. 10. Gray, Asa (1950). Manual of botany. 8th (centennial) ed. Rev. and enl. by M. L. Fernald. American Book Com- pany, New York. Gray, Asa, and Sereno Watson (1878-95). Synoptical flora of North America. Con- tinued and ed. by B. L. Robinson. 2 vols. in 3. American Book Company, New York. Greene, L. H., and J. A. Roddick (1962). Dawson, Larsen Creek, and Nash Creek map areas, Yukon Territory. Geol. Surv. Can. Pap. 62-7. Hitchcock, A. S. (1950 [i.e.] 1951). Manual of grasses of the United States. 2nd ed., rev. by Agnes Chase. U.S. Govern- ment Printing Office, Washington, D.C. Hitchcock, C. Leo, and Arthur Cronquist (1964). Salicaceae to Saxifragaceae. Pt. 2 of C. Leo Hitchcock, Arthur Cronquist, Marion Ownbey, and J. W. Thompson, Vascular plants of the Pacific northwest, University of Washington Press, Seattle. Hitchcock, C. Leo, Arthur Cronquist, and Marion Ownbey (1969). Vascular Cryptogams, Gymnosperms, and Monocotyledons. Pt. 1 of C. Leo Hitchcock, Arthur Cronquist, Marion Ownbey, and J. W. Thompson, Vascular plants of the Pacific northwest. University of Washington Press, Seattle. 15 Bibliography Holm, H. T. (1885). Novaia-Zemlia’s Vegetation, saerlig dens Phanerogamer. Pages 5-59, pls. 1-2 in C. F. Lutken, ed., Dijmhnatogtets zoologisk-botaniske Udbytte. Copenhagen Zoological Museum, Copenhagen. Hooker, W. J. (1829-40). Flora Boreali-Americana. 2 vols. H. G. Bohn, London. Hopkins, Milton (1937). Arabis in eastern and central North Ameri- ca. Rhodora 39: 63-186. Hultén, Eric (1928). Flora of Kamtchatka and the adjacent islands. 2, Dicotyledoneae, Salicaceae, Cruciferae. Kgl. Svenska Vetensk. Hand. 5 (2). (1930). Flora of Kamtchatka and the adjacent islands. 4, Dicotyledoneae, Pyrolaceae, Composi- tae. Kgl. Svenska Vetensk. Hand. 8 (2). (1937). Flora of the Aleutian Islands. Bokforlags Aktiebolaget Thule, Stockholm. (1941-50). Flora of Alaska and Yukon. Pts. 1-10. Lunds Univ. Arsskr. N.F., Avd. 2, 37(1)—46(1). (1966). New species of Arenaria and Draba from Alaska and Yukon. Bot. Notis. 119 (2): 313-16. (1967). Comments on the flora of Alaska and Yukon. Ark. Bot., ser. 2, vol. 7 (1). (1968). Flora of Alaska and neighboring territories. Stanford University Press, Stanford, Calif. Jeffrey, W. W. (1961). Notes on plant occurrence along lower Liard River, Northwest Territories. Nat. Mus. Can. Bull. 171: 32-115. Johnson, A. W., and J. G. Packer (1968). Chromosome numbers in the flora of Ogo- toruk Creek, N.W. Alaska. Bot. Notis. 121: 403-56. Jorgensen, C. A., T. Sorensen, and M. Westergaard (1958). The flowering plants of Greenland: a taxo- nomical and cytological survey. Biol. Skr. Kgl. Dan. Vidensk. Selsk. 9 (4). Kalela, Aarno (1944). Systematiche u. Pflanzengeographische Studien an der Carex—Subsektion Alpinae. Ann. Bot. Fenn. 19 (3). (1965). Uber die Kollektivart Carex brunnescens (Pers.) Poir. Ann. Bot. Fenn. 2 (2): 173-218. Kjellman, F. J. (1883.) Fanerogamfloran pa St. Lawrence-6n. Pages 1-23 in N.A.E. Nordenskidld, Vega-expedi- tionens vetenskapliga iakttagelser, 1883, Bd. 2. Komarov, V. L., ed. (1934). Flora SSSR. Vol. 2, ed. by R. Yu. Rozhevits and B. K. Shishkin. Izdatel’stvo Akademii Nauk SSSR. 76 Lid, Johannes (1963). Norsk og Svensk Flora. Det Norske Samla- get, Oslo. Love, Doris, and N. J. Freedman (1956). A plant collection from S.W. Yukon. Bot. Notis. 109 (2): 153-211. Mackenzie, K. K. (1935). North American flora. Vol. 18, pt. 7. New York Botanical Garden, New York. Maguire, Bassett (1943). A monograph of the genus Arnica. Brittonia 4 (3): 386-510. Meyer, C. A. (1829). Novae plantarum species (Claytonia sar- mentosa, C. stolonifera, Ribes tubiflorum). Moscou Soc. Nat. Nouv. Mem. 7, n.s. 1: 135-46. Moss, E. H. (1959). Flora of Alberta. University of Toronto Press. Mulligan, G. A. (1970). Cytotaxonomic studies of Draba glabella and its close allies in Canada and Alaska. Can. J. Bot. 48 (8): 1431-37. Mulligan, G. A., and A. E. Porsild (1969). Chromosome numbers of some plants from the unglaciated central Yukon Plateau, Canada. Can. J. Bot. 47 (5): 655-62. (1970). Chromosome number reports. Pages 111- 12 in A. Love, ed., IOPB chromosome number re- ports, no. 25. Taxon 19 (1). Pohle, R. (1914). Bull. Jard. Bot. Petersb. 14:467. Polunin, Nicholas (1940). Botany of the Canadian Eastern Arctic. Pt. 1, Pteridophyta and Spermatophyta. Nat. Mus. Can. Bull. 92. Porsild, A. E. (1939). Contributions to the flora of Alaska. Rho- dora 41: 141-83; 199-254; 262-301. (1943). Materials for a flora of the continental Northwest Territories of Canada. Sargentia, no. 4: 1-79. (1945): The alpine flora of the east slope of Mac- kenzie Mountains, Northwest Territories. Nat. Mus. Can. Bull. 101. (1947). The genus Dryas in North America. Can. Field-Natur. 61 (6): 175-92. (1950a). The genus Antennaria. Pages 1511-39 in Eric Hultén, Flora of Alaska and Yukon, pt. 10. Lunds Univ. Arsskr. N.F., Avd. 2, 46 (1). (1950b). The genus Antennaria in northwestern Canada. Can. Field-Natur. 64 (1): 1-25. Bibliography (1951a). Botany of southeastern Yukon adjacent to the Canol Road. Nat. Mus. Can. Bull. 121. (1951b). Two new Oxytropis from arctic Alaska and Yukon. Can. Field-Natur. 65 (2): 76-79. (1954). The North American races of Saxifraga flagellaris Willd. Bot. Tidsskr. 51: 292-99. (1955). The vascular plants of the western Cana- dian Arctic Archipelago. Nat. Mus. Can. Bull. 135. (1961). The vascular flora of an alpine valley in the Mackenzie Mountains, N.W.T. Nat. Mus. Can. Bull. 171: 116-30. (1964a). Potentilla stipularis L. and Draba sibirica (Pall.) new to North America. Can. Field-Natur. 78 (2): 92-97. (1964b). Illustrated flora of the Canadian Arctic Archipelago. 2nd rev. ed. Nat. Mus. Can. Bull. 146. (1965a). Some new or critical vascular plants of Alaska and Yukon. Can. Field-Natur. 79 (2): 79-90. (1965b). The genus Antennaria in eastern arctic or subarctic North America. Bot. Tidsskr. 61 (1-2): 22-55. (1966). Contributions to the flora of southwestern Yukon Territory. Nat. Mus. Can. Bull. 216: 1-86. (1967). Draba sibirica (Pall.) Thell. in North Ameri- ca. Can. Field-Natur. 81 (3): 165-68. (1972). The vascular flora of limestone hills of the northern extension of the Ogilvie Mountains, Yukon Territory. Arctic 25 (3): 233-36. Porsild, A. E., and W. J. Cody (1968). Checklist of the vascular plants of the con- tinental Northwest Territories, Canada. Can. Dep. Agr. Plant Res. Inst., Ottawa. Porsild, A. E., and Howard Crum (1961). The vascular flora of Liard Hotsprings, B.C., with notes on some bryophytes. Nat. Mus. Can. Bull. 171: 131-97. Raup, Hugh M. (1947). The botany of southwestern Mackenzie. Sargentia, no. 6: 1-275. Rollins, R. C. (1952). Cardamine digitata Richardson (Crucife- rae). Rhodora 54: 260. Rydberg, P. A. (1922). Flora of the Rocky Mountains and adjacent plains. 2nd ed. The author, New York. (1932). North American flora. Vol. 21, pt. 4. Arctic Inst. N. Amer. Spec. Publ. 4. University of Toronto Press. Schulz, O. E. (1927). Cruciferae-Draba et Erophila. Pflanzenr. IV, 105, Heft 89. Seeman, B. C. (1852). The botany of the voyage of H.M.S. Herald ... 1845-51. L. Reeve, London. Tolmatchev, A. I., ed. (1964, 1966). The arctic flora of the USSR. Pts. 2 and 3. Izd-vo Nauka, Leningrad [in Russian]. Vasey, G. (1882). Some new grasses. Bot. Gaz. 7 (3): 32-33. Vassiljev, V. N. (1953). Luzula DC. Acad. Nauk SSSR Bot. Inst. Bot. Mater. Gerb. 15: 39-44. Wiggins, Ira L., and John Hunter Thomas (1962). A flora of the Alaskan Arctic Slope. Arctic Inst. N. Amer. Spec. Publ. 4. i ii ys : ty Ls "1 3 ure Fide a) ~ \ eed “ a ” a be 4 ult poly cary 7 Ae a ij Sil wedi DEG 104 Rabi ld ah ka a Mit ; Tay wn ; r ? he? if Wa mn cen; her ule ri Pe : i » “ fF F ¥ ‘ : . . \ 4 bi) bs vis 4 ee ban \ by aot . eal wheel ihc. eeind ma bait sik ve ‘sitter ANCOR a Le 9 nal | hy Me a y j ‘ | nie oii: dai Ow Me te We be ir i te Pligtt, 1 4 & q ‘ 4 g aaF ; re ii by 7 (eam - | | ak : (Pee hr to eiriald wus - P ; my bid 4euPA Agi oO¢ Seabei 9G A)T. (Sh 3 Bie an vi, ’ i | ' v oat (oi Y ye wy, ee antl } ) 7 i. ; ‘ rs , 3 i ; 5 a’? ; pe " rl , : a i] ~ si Ret nite & , i | i ' ‘ ) iv " ' fi at Ee | } ) (es | 7 7) ¥ 1 4 ~ Aa 4 / a ‘ . ¥ i? ‘ | - { vig | “Ay / ‘ ; wi | ny 9 mm Fide 4 _ . \ 4 : Sul 5 ae Teel. | b 2 a ’ y Tee yutne, | ) F 7a ofe “ac j i ' Por a VS Ee as wal 4 %) 2 if * ies? y tS ae } a ¥ \ of ' ap SS @ | = \ ] 7 * 4 4 J y+ Le § nS Ai | it we us! % WwW u | f » . 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