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Statement relating to the Society’s Property : December 31st, 1915. a5 & OW Balance in the Bankers’ hands, December 31st, 1915: » On Current Account ............6:. ce cee cee eee ees WAS 83) 7 », Deposit eA Aan aT sear Sse at Mayne ee RA 500 0 0 Balance in the Clerk’s hands, December 31st, 1915 719 9 Due from Messrs. Longmans & Co., on account of the Quarterly Journal, Vol. LX XI, etc. .... 364 «3 Arrears of Admission-Fees (estimated) ........ 50 8 O Arrears of Annual Contributions .......... Se oo e OnG (Estimated to produce £192 18s. Od.) Funded Property, at cost price :— £2500 India 8 per cent. Stock ............ 2623 19 0 £300 London, Brighton, & South Coast Rail- way 5 per cent. Consolidated Preference StOCK . 6.66 seek eee ee eee e erent teres 502 15 3 £2250 London & North-Western Railway 4 per cent. Preference Stock ............ 2898 10 6 £2800 London & South-Western Railway A per cent. Preference Stock ...........- 3607 7 6 £2072 Midland Railway 23 per cent. Per- petual Preference Stock. .........00000- 1850 19 £267 6s. 7d. Natal 3 per cent. Stock........ 250 0 £2000 Canada 33 per cent. Stock .......... 1982 11 [Norre.—The above amount does not include the value of the Libr and Stock of unsold Publications. The value of the Funded KOO Leet: a 635 18 4 468 12 9 £1104 61 £13716 2 9 ary, Furniture, Property of the Society, at the prices ruling at the close of business on December 3ist, 1915, amounted to £9,837 3s. 2d. | BEDFORD McNEILL, Treasurer. January 29th, 1916. x] PROCEEDINGS OF THE GEOLOGICAL SOcIETY. [ vol. Ixxii, AWARD OF THE WoniAston MEDAL. In handing the Wollaston Medal, awarded to Dr. ALEXANDER Perrovicn Karprysky, to M. Constantin Nazoxorr, Councillor of the Imperial Russian Embassy, the Prestpent addressed him as follows :— Councillor NaBoxorr,— The Council of the Geological Society has this year awarded the Wollaston Medal, its highest distinction, to Dr. Alexander P. Karpinsky, Honorary Director of the Geological Committee of Petrograd, which is responsible for the geological survey of the Russian Empire. Dr. Karpinsky’s activities have extended over a period of more than forty years, and so long ago as 1874 he made one of his most important discoveries, that of a marine formation in the Ural Mountains intermediate between the Carboniferous and the Permian Systems. This Artinskian Stage, as Dr. Karpinsky termed it, has now been traced in Russia almost from the Arctic Ocean to the Caspian Sea, besides being recognized in more remote regions, as in the Salt Range of India. Its interesting fauna has also been the subject of several important monographs, of which one of the most valuable is that on the Ammonoids, contributed by Dr. Karpinsky himself to the Imperial Academy of Sciences of Petrograd in 1889. Dr. Karpinsky has continued to take the deepest interest in the geological problems presented by the Urals, and has treated them with remarkable versatility from every point of view, whether tectonic, petrographical, or paleontological; but as official director of the surveys from 1885 to 1903 he also extended his researches to many other districts, and took a pro- minent part in the preparation of the beautiful geological maps which were issued during his period of active service. The useful Geological Map of Russia in Europe, which he edited in 1893, is especially well known. All Dr. Karpinsky’s work is characterized by the most painstaking thoroughness, of which I need only cite his two exhaustive memoirs on the Carboniferous ichthyolite, Helicoprion, as conspicuous examples. Those who have the privilege of his personal acquaintance recognize in him an un- assuming and enthusiastic student, still absorbed in following and part 1] ANNIVERSARY MEETING—MURCHISON MEDAL. xl aiding the progress of our science, and pre-eminently one whom - the Geological Society delights to honour. The Council will be glad if you will convey this medal to. Dr. Karpinsky as a token of its esteem and admiration, with an expression of its best wishes. Councillor NaBoxorr replied in the following words :— Mr. PREsIDENT,— Please accept my sincere thanks for the honour that you have done me in asking me to come here to-day and to convey to Dr. Karpinsky, with the expression of your good wishes, the Wollaston Medal which the Council of the Geological Society has awarded to him. I feel certain that this great distinction will be deeply appreciated by the recipient of the medal, as well as by the Russian Geological Committee as a high tribute to their Director. My distinguished friend, Dr. H. H. Hayden, Director of the Geological Survey of India, who crossed the Pamirs from India into Russian Turkestan a few months before the war, has often expressed to me the wish and hope that the highly interesting and valuable scientific researches, which have been carried out on both sides of the Pamirs by the British and Russian geologists, may be linked up and conducted on a basis of firmer and more complete unity and coordination. I venture to avail myself of this opportunity of expressing, on behalf of my countrymen, the same wish, and the confident hope that the ties of friendship which now unite Britain and Russia may extend from the fields of battle to the lofty peaks of science and enlightenment. Awarb oF THE Murcuison MEDAL. The PrestpEnt then handed the Murchison Medal, awarded to Dr. Ropert Kripsron, F.R.S., to Dr. F. L. Kircury, for trans- mission to the recipient, and addressed him as follows :— aa Dr. Krrcury,— The Council has awarded to Dr. Robert Kidston the Murchison Medal as a mark of its appreciation of his numerous and valuable contributions to our knowledge of fossil plants, especially those of the Carboniferous Period. For nearly forty years he has devoted xli PROCEEDINGS OF THE GEOLOGICAL socteTy. [ vol. lxxu, himself to an exhaustive and successful study of the external characters of the plant-remains associated with the various coal- seams ; and in this manner he has acquired an unrivalled knowledge of the distribution of the Carboniferous flora, which has proved of fundamental importance both to the geologist and to the practical miner. J] may mention, as examples of this work, his classic memoirs on the fossil plants of the Yorkshire and Staffordshire Coalfields and of Belgian Hainaut. During more recent years he has also extended his researches to various facts of structure and morphology which have a direct bearing on evolutionary problems. His memoir on the fructification of Newropteris heterophylla was the first description of the seed of a Pteridosperm in direct continuity with the frond; while his account of the microsporangia of the Pteridosperms first demonstrated the nature of the male organs in plants of this transitional group. His description of the internal structure of Szgillaria, and his remarkable series of memoirs, with the late Prof. Gwynne-Vaughan, on the evolution of the Osmundacez, must also be specially mentioned. While pursuing his researches he has continually recognized the importance of careful field-work, and has thus made a large and valuable collection of specimens, which has always been placed freely at the disposal of his fellow-paleobotanists. In transmitting this medal, please express our hope that he will treasure it not only as a token of our admiration, but also of our gratitude. Dr. Krrceuiy replied in the following words :— Mr. PrREesipENT,— It gives me great pleasure and satisfaction to be here to-day as Dr. Kidston’s representative, to accept on his behalf this valued award and to convey to you his thanks for the honour conferred on him by the Council of this Society. And I desire to thank you, Sir, in his name, for the kind words with which you have accompanied this presentation. It is gratifying to be the transmitter of the Murchison Medal to one who, a Scotsman himself, has laboured so long and so assiduously in elucidating the stratigraphical bearings of the Carboniferous Flora. Dr. Kidston, I feel sure, would have received this medal with enhanced pleasure, could he have listened to your graceful and appreciative references to his work. He asks me to express to you his great regret that he is unable to be here in person; and part 1] ANNIVERSARY MEELING—LYELL MEDAL. xl I may add that he is detained by responsible public duties, which have the first claim upon his time. | I have received a letter from Dr. Kidston which, with your permission, I will read :— ‘Will you please express to the President my sorrow at not being able to be present to thank the Society personally for the honour that they have done me in presenting me with the Murchison Medal, an honour which, it is needless for me to say, I very much value and appreciate. ‘The award of this Medal brings vividly to my memory that a number of years ago the Society awarded to me the Balance of the Proceeds of the Murchison Geological Fund, and I would like them to know that these proceeds were spent in the purchase of books dealing with Paleozoic Botany. It is only workers situated where not a single book on their special subject of study is obtainable for reference, who can fully appreciate the value of the help that I received from that award, and I hope that the books will eventually be placed where they will be of help to others. ‘I have now only to thank the Council of the Geological Society once more for its kind and encouraging recognition of my work.’ AWARD OF THE LYELL MEDAt. In presenting the Lyell Medal to Dr. Cuartes WinLiAM Anprews, F.R.S., the Prestpent addressed him as follows :— Dr. ANDREWS,— The Council has awarded to you the Lyell Medal as an acknow- Jedgment of the value of your numerous researches in Vertebrate Paleontology. Since your appointment to the Geological Depart- ment of the British Museum in 1892, you have made excellent use of the opportunities for research afforded by your official duties, and have made important contributions to our knowledge of fossil reptiles, birds, and mammals. You were soon attracted by the unique Leeds Collection of Oxfordian marine reptiles, and your studies of this collection eventually culminated in the two handsome volumes of the Deseriptive Catalogue, published by the Trustees of the British Museum (1910-13), which must always remain a standard work of reference on Ichthyopterygia, Sauropterygia, and Crocodilia. Your papers on the South American Stereornithes, on Rails from islands in the Southern Seas, and on Prophaéthon from the London Clay, are equally valuable contributions to our knowledge of extinct birds. Your researches on the fossil mam- mals of Egypt, many of them discovered by yourself, are still more noteworthy ; and your Descriptive Catalogue of the Tertiary xliv PROCEEDINGS OF THE GEOLOGICAL SOCIETY. | vol. xxii, Vertebrata of the Faytim (Egypt), published by the Trustees of the British Museum in 1906, began a new era in the history of mammalian life. Your demonstration of the stages in the evolution of the Proboscidea, and of the relationship between the Proboscidea and the Sirenia; your description and interpretation of the strange Eocene genus Arsinoitherium ; and your recognition of the early differentiation of the Hyracoids in Africa are especially funda- mental contributions to biological and geological science. I would further add that all your writings are characterized by remarkable thoroughness and insight into the meaning of the facts described. As your colleague in the British Museum during the whole period of your service, it gives me great pleasure to hand to you this medal, which the Council of the Geological Society could not have more worthily bestowed. Dr. ANDREWS rephed in the following words :— Mr. Presipent,— I wish to express my most sincere thanks to the Council of the Geological Society for the honour that it has done me in awarding to me the Lyell Medal, and to you, Sir, for the too flattering terms. in which you have made the presentation. I am_ particularly pleased to have received this medal from the hands of one with whom I have been associated for so many years. You will remember that, exactly twenty years ago, you yourself received this award from Dr. Henry Woodward, and that at the same time I received a moiety of the Balance of the Proceeds of the Lyell Geological Fund. If I have been able to accomplish something in Vertebrate Paleontology, it is mainly due to the fortunate environment in which I have found myself. An assistant in the British Museum possesses quite exceptional advantages, having free access to the great libraries and to the ever-increasing collections, and lastly, but by no means least, having many opportunities of making the personal acquaintance of workers interested in his subject. Having enjoyed these privileges, I feel that I have somewhat fallen short of what I ought to have accomplished; but, although it is just now uncertain what the future may have in store for us, I hope that I may still have opportunities of doing further work such as. will justify this award. part 1] ANNIVERSARY MEETING MURCHISON FUND. xlv AWARD FROM THE WoLLAston Donation Funp. The PrestpEnT then handed the Balance of the Proceeds of the Wollaston Donation Fund, awarded to W1ii1am BourkE WRIGHT, B.A., to Mr. G. W. Lampruen, F.R.S., for transmission to the recipient, addressing him as follows :— ! Mr. LamriueH,— The Balance of the Proceeds of the Wollaston Donation Fund is awarded to Mr. William Bourke Wright, in recognition of his contributions to Quaternary Geology. After completing his geological studies under Prof. J. Joly at Dublin University, Mr. Wright joined the Irish branch of the Geological Survey, and came under the influence of yourself when you were engaged in working out the glacial problems of the Dublin district. He took part in the revision of the memoirs and drift-maps of the Dublin, Belfast, and Cork districts, and shared with Mr. H. B. Maufe the discovery of a continuous raised-beach feature older than the Glacial Period. He also observed this pre-Glacial rock-shelf or beach in the West of Scotland, showing that a general subsidence allowed the sea to enter the valleys along the coasts of the British Isles, almost at the present sea-level, before they were occupied by the ic& After some experience both in Scotland and in England, Mr. Wright returned to Ireland, where, as Senior Geologist of the Trish Survey, he has since been successfully engaged on the glacial geology of the Kenmare and Killarney district. Much of his leisure has been devoted to the preparation of an important work on ‘The Quaternary Ice Age,’ in which he has made good use of his observations not only in the British Isles, but also in Scandinavia. Impressed by the value of Mr. Wright’s researches, the Council will be glad if you will transmit this award to him, with its best wishes for the progress of the work which he has so well begun. AWARD FROM THE Murcuison GronocicaL Funp. In handing the Balance of the Proceeds of the Murchison Geological Fund, awarded to Mr. Groraz Wanrer TyRRELL, E.G.S., to Dr. H. Lapworrn, Sec.G.S., for transmission to the recipient, the PRESIDENT addressed him in the following words :— xvi PROCEEDINGS OF THE GEOLOGICAL Society. | vol. lxxu, Dr. Larwortu,— The Balance of the Proceeds of the Murchison Fund has been awarded to Mr. G. W. Tyrrell in recognition of his contributions to the petrology of South-Western Scotland. His keen petrographic insight was first shown in his description of the quartz-dolerite sills of Kilsyth. His results of most general interest to geologists are those connected with the Paleozoic alkaline rocks ; for his investiga- tion of lugarite has added to petrology a peculiar rock-species and important evidence in favour of the differentiation of igneous rocks by the sinking of their heavier constituents. In several papers on the Auchineden Hills he has described their igneous rocks and their glacial and physical features; and, in his recent account of the ravine known as the Whangie, he has advanced conclusive evidence of its formation by earth-movements. As the Senior Assistant in the Geological Department of Glasgow University, and later also as Lecturer on Petrology there, he has done much towards the development of that school of geology. The Council hopes that this award may encourage and assist him in further research. AWARDS FROM THE LyELL Gronogican Funp. The PruestpEnr then presented a moiety of the Balance of the Proceeds of the Lyell Geological Fund to Mr. Marrin A. C. Hinton, addressing him as follows :— Mr. Hinton,— The Council has awarded to you a moiety of the Proceeds of the Lyell Fund in recognition of your researches on the British Pleistocene Mammalia, and as an incentive to further work of the same kind. Under circumstances frequently discouraging, you have for many years devoted yourself especially to the study of the Rodentia and the Insectivora, and have obtained a remarkable knowledge of the skeleton and teeth of certain groups which are most commonly met with among fossils. In this manner you have made discoveries with an important bearing on many problems of Pleistocene geology, which you have never failed to recognize. As one who has followed your work with great interest for several years, I have much pleasure in handing to you this award. part 1] ANNIVERSARY MEETING—LYELL FUND. xlvit The PrestpEnr presented the other moiety of the Balance of the Proceeds of the Lyell Geological Fund to Mr. Anrrep SanTER Kewnarp, F.G.S8., addressing him in the following words :— Mr. Kennarp,— It is particularly appropriate that the second moiety of the Proceeds of the Lyell Fund should be awarded to you, who have worked so long and so successfully with Mr. Hinton at problems of Pleistocene geology in the South of England. In the leisure of a busy life, you also have made yourself thoroughly acquainted with a group of fossils, the non-Marine Mollusca, which are of fundamental importance in classifying and interpreting the various deposits in which they occur. Both alone, and with Mr. B. B. Woodward, you have published many interesting notes and lists of such mollusca from Pleistocene and Holocene deposits in different parts of Britain. The Council desires to acknowledge the value of this work, and I have much pleasure in handing to you é tangible expacssion of its good wishes. xlviii PROCEEDINGS OF THE GEOLOGICAL socinTy. [ vol. ]xxil, THE ANNIVERSARY ADDRESS OF THE PRESIDENT, Arrnur Smita Woopwarp, LL.D., F.R.S. Amone those whom we have lost by death during the past year are two distinguished Foreign Members, Count Solms-Laubach and Dr. ©. R. Zeiller, who devoted themselves to Paleeobotany. For obituary notices of them I am indebted to Prof. A. C. Seward. ‘There are also two Foreign Correspondents, and several Fellows who have done great service to Geology. Hermann Graf zu Sorms-Lavpacnu died on November 24th, 1915, in his seventy-third year. He was elected a Foreign Member ot the Linnean Society in 1887, of the Royal Society in 1902, and of the Geological Society in 1906; in 1911 the Gold Medal of the Linnean Society was awarded to him, and with other foreign visitors he received an Honorary Degree at Cambridge on the occa- sion of the Darwin Celebration in 1909. Solms-Laubach belonged to ‘one of the most ancient German families, who were sovereigns in their own right down to the year 1806’ (‘ Nature,’ January 13th, 1916). He occupied the Chair of Botany at Gottingen, and succeeded the eminent botanist de Bary as Professor in the University of Strassburg. Though well known as the author of many important papers on recent plants, it is the distinguished part that he took in the reformation and revival of paleobotanical research which more particularly appeals to geologists. When he began to write his ‘ Hinleitung in die Paladontologie,’ he found it necessary to study the unrivalled collection of sections in the possession of Prof. W. Crawford Williamson, and for this purpose paid several visits to Manchester, which, as he says in a sym- pathetic biographical sketch of his friend published in ‘ Nature,’ September 5th, 1895, knitted closer the bonds of reverence and friendship between himself and his English colleague. The publication of Solms-Laubach’s book in 1887 led to a greater appreciation of the value of Wilhamson’s work, and his critical treatment of the widely-scattered literature enabled botanists to obtain a general view in truer perspective than had previously been possible of the significance of paleobotanical records. In the preface he wrote :—‘ Botany, which in former times generally treated Paleophytology in a very stepmotherly manner, now part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. xix finds it to be a subject of the highest interest to herself on account of the prominence at present assumed by the point of view of the theory of descent.’ The publication of an English translation in 1891 by the Clarendon Press conferred a boon upon many students to whom Count Solms’s involved and difficult style was a serious obstacle. The book contains a mass of valuable information, based in great measure on an actual examination of the specimens described; it is characterized by vigorous and discriminating criticism of the conclusions of previous writers, and the whole bears striking testimony to the author’s grasp of his subject, his exceptional power of handling details without losing sight of guiding principles, and his wonderful energy. Hor many years Solms-Laubach contributed to the ‘ Botanische Zeitung,’ and later to the ‘ Zeitschrift fiir Botanik,’ eritical summaries of recent work, which have played a con- spicuous part in keeping botanical readers informed of the more important results of paleobotanical enquiry. The great majority of Solms-Laubach’s published papers deal with petrified plants, and it was but rarely that he concerned himself with impressions. He made notable additions to our knowledge of the Paleozoic genera Medullosa, Protopitys, Sphenophyllum, and other types, and supplied fresh data of special interest from an evolutionary point of view. His researches into the structure of Lower Carboniferous and Upper Devonian plants yielded results of the greatest interest; he not only corrected the mistakes of earlier investigators, but presented for the first time an accurate picture, so far as the fragmentary nature of the material permitted, of the morphological characters of some of the oldest known plants. The discovery of several new generalized types gave emphasis to the view that the extinct Devonian genera represent highly-complex terms in a series extending back into ages far beyond those that have left any decipherable records. The account of his investigations on Stig- mariopsis and other plants in the quarries of St. Etienne threw new light on a subject that is still far from exhausted, and in a more recent paper he cleared up certain difficulties connected with the method of growth of the root-encircled stems of Psaronius. In collaboration with Capellini, Solms gave a systematic account of the splendid Cycadean stems from Northern Italy in the Bologna Museum, and, by his discovery of pollen-grains asso- ciated with a female flower, foreshadowed the later discoveries of Wieland, who had at his disposal the much more complete VOL. LXXIT. d 1 PROCEEDINGS OF THE GEOLOGICAL society. [vol. lxxu, American material. His work on the reproductive shoots of the English Cycadean stem Bennettites gibsonianus, origimally described by Carruthers, is especially noteworthy as an example of intensive study combined with rare morphological insight. Papers on the structure of some Conifers from the copper-bearmg Permian beds of Ilmenau and Frankenberg, and on Jurassic Gymnosperms collected in Franz-Josef Land by members of the Jackson-Harmsworth Expedition in 1894-96, are of special value as elucidating the structural features of genera previously known only as casts or impressions. Count Solms added greatly to our knowledge of recent and fossil calcareous Alge, not only in his text-book, but also in his monograph of the Acetabulariez published by the Linnean Society in 1895, and in other papers. His name will long be remembered with gratitude and respect by students of ancient plants; he raised the subject of Palzo- botany to a higher plane, and by his writings, as also in no small degree by his enthusiasm and infectious energy, he was the means of attracting many botanists to a branch of the science which had suffered neglect and had been discredited through its treatment at the hands of authors insufficiently equipped with a knowledge of recent plants. Solms-Laubach paid several visits to this country, where he was always a welcome guest. He was a man of considerable force of character, cultivated, and endowed with a sense of humour; a warm-hearted friend; and an in- vestigator of marked originality, whose work, more especially in the domain of Paleobotany, has had a wider influence than that of many men whose output was larger. PAR Cys CHARLES René ZEILLER died in Paris on November 27th, 1915, after a long and painful illness, at the age of 68. He was born at Nancy, his father being Ingénieur-en-Chef des Ponts et Chaussées, and his mother a great-granddaughter of the Lorraine artist Guibel, sculptor to King Stanislaus of Poland, Duke of Lorraine. He married in 1877 the sister of Léon Ollé-Laprune, the Catholic philosopher and a Member of the Institute, whose religious and philosophical views he shared. Prof. Zeiller was a Member of the Institute, Commander of the Legion of Honour, Inspector-General of Mines, and Professor at the Ecole Nationale Supérieure des Mines ; in 1905 he was elected a Foreign Member of the Linnean Society, and of the Geological Society in 1909. Zeiller rarely travelled in other countries than part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. li his own: he paid his first visit to England in 1909, when he attended the Darwin Celebration at Cambridge; and it is a satisfaction to learn from his daughter that, during his illness, he derived no little pleasure from the recollection of the days spent in this country. His dignified bearing and handsome face made him conspicuous among the distinguished foreigners upon whom the Chancellor conferred Honorary Degrees. Despite ex- acting official duties, Prof. Zeiller found time to make numerous substantial contributions to Paleobotany and, incidentally, to Geology. One of his earliest works, published in 1878 (the year in which he instituted the course of lectures on Paleobotany at the School of Mines), is an account of the plants of the French Coal Measures, the first of a series of admirable volumes on Upper Carboniferous and Permian floras. The two volumes on the botany of the Valenciennes Coalfield which appeared: in 1888 afford a good example of the author’s thoroughness of treatment and lucidity of style; they are not merely important from the point of view of the systematist and stratigraphical geologist, but the enlightened and philosophical treatment of the extinct types in their relation to recent plants gives them a high botanical value. Zeiller’s attitude was thoroughly scientific; he was always ready to consider criticisms and suggestions, and punctilious in his reference to the labours of colleagues: in him Solm-Laubach’s aphorism, ‘Bescheidenheit ist eine Zier,’ was conspicuously demonstrated, The description of the flora of Commentry and of the plants of Autun was shared with Renault. In the volumes on the Brive, Creusot, and Blanzy fossils, Zeiller made many additions of the first importance to our knowledge of several Paleozoic types, notably, in the Brive flora, as regards the main anatomical features of Psaronius. His work on the Coal Measures of Heraklia, in Asia Minor, and his description of Permian plants from Lodéve, contain much of great botanical interest. Without attempting to give a list of the types which he was the first to deseribe, or to enumerate the genera on which his investigations threw new light, one may refer to his memoir on the fructification of Sphenophyllum, published in 1898, as a remarkable example of the valuable results that can be obtained by a patient and skilful examination of unpromising material. Reference must also be made to the discovery of Paleozoic species closely allied to Selaginella, to his more recent researches into the anatomical characters of Lepidostrobus, and to his a2 ln PROCEEDINGS OF THE GEOLOGICAL SOCTETY. | vol. xxii ? discovery of new forms of Fern-like fructifications. Zeiller was the first to recognize the generic identity of the Indian genus Trizygia and the European Sphenophyllum; and his ingenious explanation of the peculiar features of the widespread Indian and Southern-Hemisphere genus Vertebraria is the best that has so far been suggested. One of the outstanding features of Zeiller’s work is the high standard of excellence of his descrip- tions of plants preserved as impressions: the comparatively few papers on petrified plants show that he was also thoroughly competent as an anatomist, but it is the high standard of his descriptive work, the determination to exhaust every method of attack, and his sanity of judgment and breadth of view that give a permanent value to his achievements. Zeiller’s accurate stratigraphical knowledge of the French coalfields enabled him to contribute in no small degree to the better appreciation of the value of plants as indices of geological age. He considerably extended our knowledge of the GJlossopteris flora in Brazil and South Africa, and described some new forms from the Lower Gondwana rocks of India; his paper on ‘ Les Provinces Botaniques a la Fin des Temps Primaires,’ in the ‘ Revue Générale des Sciences’ (1897), is a model of clear exposition and a highly-suggestive presentation of one of the most fascinating problems of geo- graphical distribution and plant-migration. Another contribution of especial interest from the same point of view is his critical examination of the Siberian plants referred by Schmalhausen to a Jurassic age, which led to their recognition as members of a Permian flora closely allied to those of Gondwanaland. Among many publications dealing with Mesozoic floras, the monograph of the Tongking Rheetic plants deserves special mention; it is one of the best works of the kind that we possess, not merely because of its excellent illustrations, but for the wealth of information which it contains and the masterly treatment of the rich material. Zeiller described collections of fossil plants from many parts of the world, from New Caledonia, Madagascar, China, Peru, North Africa, the Balkans, and elsewhere; his ‘Eléments de Paléobotanique,’ published in 1900, though of necessity greatly condensed, is a useful book for beginners as well as for experienced students. For several years Zeiller supplied comprehensive and critical summaries of recent palzobotanical literature to the ‘ Revue Générale de Botanique,’ and the thoroughness with which the laborious task was performed is characteristic of the man; he part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. lui was ever ready to help others, and unsparing of himself in the interests of the subject which he loved. It was a privilege to count Zeiller a friend; he was a man of strong affections, and deeply religious. While we admire his intellectual power and his whole-hearted devotion to the advancement of knowledge, it is his singularly-attractive personality and his lovable character that bulk largest in our recollection of the colleague whom we -have lost. To quote the words used by Zeiller in a biographical sketch of the Marquis of Saporta: he has left for all ‘le souvenir d’un maitre aussi aimé que respecté, en méme temps que dun des esprits les plus éminents dont ait a s’enorgueillir la paléontologie.’ pA Casa Micuren Férix Movrton, who was elected a Foreign Corre- spondent in 1899, was born on May 11th, 1845, and graduated at Brussels in 1867. He was appointed Conservator of the Royal Museum of Natural History at Brussels in 1872, and became Director of the Geological Survey of Belgium in 1897. Between 1875 and 1883 he contributed important papers on the Devonian formations of Belgium to the ‘ Bulletin’ of the Royal Academy of Brussels. He also edited the Memoirs on the Cretaceous and Tertiary formations of Belgium prepared for the Survey by A. Dumont. In 1880-81 he published his useful ‘ Géologie de la Belgique’ in two volumes, and during more recent years he wrote several papers on the Tertiary and Quaternary geology of that country. Epmonp Ricavx, who received an award from the Lyell Fund in 1883, and became a Foreign Correspondent in 1893, was an able amateur geologist residing at Boulogne. He made many important contributions to our knowledge of the geology of the Lower Boulonnais, which were eventually summarized in a memoir published by the Société Académique de Boulogne-sur-Mer in 1892. His acquaintance with the Jurassic rocks and fossils was especially profound, and his amiable services were always at the disposal of those who visited the district for geological work. He died in April 1915. Glacial Geology has lost a pioneer by the death of Prof. JAMES GEIKIE, which occurred at Edinburgh on March Ist, 1915. The younger brother of Sir Archibald Geikie, he was ‘born in liv PROCEEDINGS OF THE GEOLOGICAL soctEty. ([vol. Ixxu, Edinburgh on August 23rd, 1839, and was educated at the High School and University of that city. Having been naturally in- clined to geological studies from’ early youth, he joined the Geological Survey of Scotland in 1861, and became a District Surveyor in 1869. He was employed chiefly in the Lowlands and Southern Uplands of Scotland, often in districts of which the solid geology had already been examined, and his most important duty was to study, map, and describe the superficial deposits or ‘ drift.’ Such formations proved to have a special fascination for him, and he spent several vacations in investigating them in the Highlands, Outer Hebrides, and other areas which were beyond his official sphere. The peat-bogs especially soon attracted his notice, and seemed to him to indicate a succession of climatic changes during the period of their formation, which he described in his first important contribution to Pleistocene geology, ‘On the Buried Forests & Peat-mosses of Scotland, & the Changes of Climate which they indicate’ Trans. Roy. Soc. Edinb. vol. xxiv, 1867. He thus gradually arrived at the conclusion that the Pleistocene glacial period had not been continuous, but had been interrupted by several mild episodes or interglacial periods, and his results were eventually summarized in 1874 in his well-known volume on ‘The Great Ice Age & its Relation to the Antiquity of Man,’ of which new editions appeared in 1877 and 1894. This work was supplemented by another on ‘ Prehistoric Europe’ in 1881. In 1882 Dr. Geikie succeeded his brother as Murchison Professor of Geology in the University of Edinburgh. In that year he contributed an unportant memoir on the geology of the Ferde Islands to the Transactions of the Royal Society of Edinburgh, and expressed his views on ‘The Aims & Method of Geological Inquiry’ in his inaugural lecture at the University. He began educational work in earnest, devoting especial attention to the improvement of geographical teaching; and in 1884 he became one of the founders of the Royal Scottish Geographical Society, which fostered this work. From 1904 until 1910 he was President of that Society, and for many years he was honorary editor of its Magazine. His University students were provided for by his ‘Outlines of Geology,’ which passed through four editions (1886— 1903), and his ‘Structural & Field Geology for Students’ (three editions, 1905-12) ; while the relations of geology and geography were treated in a more general way in ‘ Fragments of Earth- Lore’ (1898), ‘Earth Sculpture, or the Origin of Land-forms’ part 1]. ANNIVERSARY ADDRESS OF THE PRESIDENT. lv (1898), and ‘ Mountains, their Origin, Growth, & Decay’ (1913). In the midst of all these new interests he continued to the end to pursue his enquiries into glacial geology, and an admirable brief summary of his latest conclusions was given in his Munro Lectures in 1913, published in book-form as ‘The Antiquity of Man in Europe’ in the following year. He retired from his Professor- ship in June, 1914. Prof. Geikie was an ideal teacher, both in the class-room and in the field, and gained from his numerous students the same deep esteem as that in which he was held by all who had the privilege of his friendship. He was elected a Fellow of the Geological Society in 1873, but had already contributed his first geological paper on the metamorphic Lower Silurian rocks of Carrick (Ayr- shire) to the Quarterly Journal in 1866. Our Murchison Medal was awarded to him in 1889, and he received the Brisbane Medal of the Royal Society of Edinburgh in the same year. He was elected a Fellow of the Royal Society of London in 1875, and was President of the Royal Society of Edinburgh at the time of his death. A list of his writings and a portrait appear in the ‘ Geo- logical Magazine,’ dec. 5, vol. x (1913) pp. 241-48 & pl. ix. RicuarD LYDEKKER, who was born in 1849 and died. on April 16th, 1915, was a Fellow of the Geological Society from 1883 to 1914, served on the Council, and became a Vice-President in 1894-97. Educated at Trimity College, Cambridge, he graduated in 1871, taking the second place in the first class of the Natural Science Tripos. In 1874 he was appointed to the Geological Survey of India, and proceeded to the exploration of the mountains of Kashmir, on which he wrote an important geological memoir. While there, his opportunities for sport gradually led him to take a special interest in the mammals and birds of India. and on returning to Calcutta he began to study the Indian Tertiary vertebrate fossils in the Survey collection. Planning a series of memoirs on these fossils for the ‘ Paleontologia Indica,’ he soon felt the necessity of ample materials for comparison. He accordingly retired from the Indian service, returned to England in 1882, and had the Calcutta collection sent in instalments to the British Museum, where he completed his work. Between 1885 and 1887 he prepared a Catalogue of the Fossil Mammals in the British Museum, in five parts. This was followed in 1888-90 by a similar Catalogue of the Fossil Reptilia & Amphibia, in four lvi PROCEEDINGS OF THE GEOLOGICAL soctery, [vol. lxxi, parts; and in 1891 by a Catalogue of the Fossil Birds, in one part. The whole series of ten small volumes did useful service to Verte- brate Paleontology at the time, and included the first systematic attempt to correlate European with American vertebrate fossils. In 1893 and again in 1894, lydekker visited the Argentine Republic at the invitation of Dr. Francisco P. Moreno, Director of the La Plata Museum. Here he studied the remarkable collection of mammalian remains from the Tertiary and post-Tertiary forma- tions of the Republic, besides a few dinosaurian bones from a Cretaceous deposit in Chubut. Some _ beautifully - illustrated memoirs in the ‘ Anales’ of the La Plata Museum were the result. By this time Lydekker had begun to realize the important bearing of recent discoveries of fossil mammals on the problems of geo- graphical distribution ; and in 1896 he published the most original and most philosophical of his works, ‘ The Geographical History of Mammals.’ During all these researches he contributed numerous preliminary notes to various serials, including the Quarterly Journal of the Geological Society. In 1889 Lydekker wrote the volume on vertebrates for the third edition of Prof. H. A. Nicholson’s ‘ Manual of Paleontology,’ and in 1891 he joined Prof. (afterwards Sir) William H. Flower in the authorship of a valuable ‘ Introduction to the Study of Mammals.’ From 1893 to 1896 he edited the ‘Royal Natural History,’ to which he himself contributed the sections on vertebrates. From 1896 until his death, much of his time was spent in arranging the public exhibition of Recent Mammals in the British Museum (Natural History), for which he was specially employed by the Trustees. His later studies therefore became more purely zoological, and he published several more or less popular volumes adapted to the needs of sportsmen. He also wrote a ‘Catalogue of the Ungulate Mammals’ and some Guide-books for the British Museum. His industry was phenomenal, but he sometimes bewailed his facility for literary composition which betrayed him into errors that on further reflection became at once apparent to him. In his ancestral home at Harpenden, though oppressed by sad domestic trouble, he continued absorbed in his favourite pursuits until the end, and his last volume for the British Museum was published posthumously. He had a large circle of devoted friends, whose admiration for his character increased the more closely they became associated with him. An excellent portrait of him appears as frontispiece to the fourth volume of the ‘Catalogue of the Ungulate Mammals in the British Museum.’ part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. lv Mr. Lydekker was elected a Fellow of the Royal Society in 1894, and the Geological Society awarded to him the Lyell Medal in 1902. CuarLes CaLtaway, who was a Fellow of the Geological Society from 1875 to 1906, and to whom the Murchison Medal was awarded in 1903, was a pioneer in the study of the British pre-Cambrian rocks, and also made valuable contributions to our knowledge of the Cambrian and Ordovician Systems. He was born at Bristol in 1838, and died at Cheltenham on September 29th, 1915. From 1876 until 1898 he resided at Wellington in Shrop- shire, and began his original researches in the area of the Wrekin. So long ago as 1874 he discovered an Upper Cambrian fauna in the Shineton Shales. A few years later he identified the under- lying greenish sandstone with the Hollybush Sandstone of Malvern. He was thus able to prove that the ancient masses of the Wrekin and the Longmynd represented a pre-Cambrian formation, which the termed Uriconian. Callaway next studied Anglesey, and came to the conclusion that the unfossiliferous metamorphic rocks of that island were also probably of pre-Cambrian age. He then visited the still more difficult region of the North-West High- lands of Scotland, and took an important share in the discussion of the great thrust-planes now recognized there. After so much experience of metamorphic rocks, Callaway began to consider the theory of dynamo-metamorphism, and applied it to the explanation both of the crystalline schists of the Malvern Hills and of certain old rocks in Galway, Donegal, and Wexford. He finally returned to the geology of Anglesey, and re-interpreted it in the ight of new knowledge, attempting to show that large masses of crystalline schists had been produced by intense changes in igneous rocks. This revised interpretation has been largely confirmed by the researches of Mr. Edward Greenly during more recent years, and will be fully dealt with in the forthcoming Geological Survey Memoir on the island. A portrait of Dr. Callaway and a list of his writings are published in: the ‘Geological Magazine,’ dec. 6, vol. 1 (1915) pp. 525-28 & pl. xvin. Arrnur VauGHaN was a brilliant exponent of the modern methods of stratigraphical geology, which he applied with success to the Lower Carboniferous formations. Born in London in 1868, Ivii PROCEEDINGS OF THE GEOLOGICAL socrEery. [vol. Ixxu, he graduated both at Cambridge and at London with high mathe- matical honours, and removed in 1891 to Clifton, where he resided until 1910. In the latter year he was appomted Lecturer in Geology in the University of Oxford, where he died on December 3rd, 1915. Beginning with the study of the Earth’s crust from the point of view of mathematical physics, he gradually became interested in all geological questions; and an intimate association with our late Fellow, Edward Wilson, definitely turned his attention to the stratigraphical distribution of fossils. After various preliminary researches, Vaughan proceeded to examine in detail the sections of Carboniferous Limestone in the Avon Gorge, and the results of his work, published in our Quarterly Journal for 1905, and in the Proceedings of the Bristol Naturalists’ Society for 1906, were not only a fundamental advance in our knowledge of the Carboniferous Limestone itself, but also a stimulating new departure in stratigraphical studies. Further researches in other areas in England, Wales, Iveland, and Belgium, enabled him to show how widely his results could be applied; and he gradually acquired so precise a knowledge of the mutations and variations of the Lower Carboniferous fossils, especially corals and brachiopods, that he was able to attempt correlations which otherwise would have been impossible. His last great work, a comparison of the Belgian with the British Carboniferous, was published im our Quarterly Journal so recently as September last. Vaughan had been in failing health for some years, and much of his investigation was carried on under difficulties, but he ever - retained his cheerful and buoyant spirit and unflagging zeal. He was an inspiring teacher and genial friend, and his untimely loss is mourned by all who knew him. He was elected a Fellow of the Geological Society in 1900, served on the Council from 1912 onwards, received the Wollaston Fund in 1907, and the Lyell Medal in 1910. A list of his writings and a portrait appear in the ‘ Geological Magazine,’ dee. 6, vol. iii (1916) pp. 92-96 & pl. v. OrviLtLE ADALBERT DeErBy, who died suddenly at Rio de Janeiro on November 27th, 1915. had devoted his life to the study of the geology of Brazil, and had done great service to our science. He was born at Kelloggsville (New York) on July 23rd, 1851, and proceeded in 1868 to Cornell University, where he came under the influence of Prof. Charles F. Hartt, who had accompanied the Agassiz Expedition to Brazil in 1865, and part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. lix had published his pioneer volume on the Geology of Brazil in 1870. After graduating in 1873, Dr. Derby became Assistant to Prof. Hartt, who had just been appointed Director of an Imperial Geological Commission of Brazil; and in the following year he went to Rio de Janeiro to act as Assistant in the con- templated survey. On the death of Hartt in 1878 the work of the Commission ceased, and in 1879 Derby was made chief of the geological section of the National Museum of Rio de Janeiro. The most important of his early memoirs on the Region of the Lower Amazons, the Cretaceous Basin of Bahia, and the Diamantiferous Region of the Parana, were published in the ‘Archivos’ of the Rio Museum in 1877 and 1878. In 1886 Derby left Rio and became Director of the Geographical and Geological Commission of the State of Sao Paulo, where he remained until 1905. He then undertook an examination of the diamond-bearing rocks of the State of Bahia, and shortly afterwards returned to Rio as Director of a new Geological and Mineralogical Survey of Brazil, which he carried on energetically until his lamented death. In 1887 and 1891 Derby contributed two papers on the nepheline-rocks of Brazil to the Quarterly Journal of the Geological Society. He was especially interested in petrology and in the occurrence of accessory minerals in rocks, and published many papers in the ‘American Journal of Science’ and in the ‘Journal of Geology,’ which are not only of scientific importance but of economic value. At the same time, he did not neglect any aspect of the science ; and one of his latest papers was devoted to an exhaustive study of the stem of the Permo- Carboniferous plant, Psaronius brasiliensis. Derby was an attractive personality, full of enthusiasm for his science, and always eager to welcome geologists who visited the land of his adoption. He gave special help and encouragement to Prof. J. C. Branner during his numerous researches in Brazil, and he similarly aided Dr. I. C. White in the preparation of his monumental volume on the coal-bearing rocks of Rio Grande do Sul. He was also associated with our Fellow, Mr. Joseph Mawson, when he was resident in Brazil and devoting attention to the geology of the country. In 1896, and again in 1907, I had the pleasure of experiencing his weleome both in Rio de Janeiro and in Sio Paulo, and learned to appreciate the difficulties under which he pursued his work with a strangely unsympathetic Government. He was elected a Fellow of the Geological Society Ix PROCEEDINGS OF THE GEOLOGICAL society. [ vol. lxxu, in 1884, and received an award from the Wollaston Fund in 1892. An excellent portrait of him appropriately appears as the frontispiece of the first edition of Prof. J. C. Branner’s ‘ Geologia Elementayr,’ published at Rio de Janeiro in 1907. Henry Hyarr Howenn was born on July 18th, 1834, and began his lifelong career on the Geological Survey of Great Britain in the autumn of 1850. As Assistant-Geologist, his first task was to accompany Beete Jukes in mapping the South Staffordshire coalfield; and for a few subsequent years he was occupied in the survey of the Midland counties. His most important memoir referred to the Warwickshire coalfield (1859). In 1855 Howell was transferred to Scotland, and devoted himself chiefly to the mapping of the Mid- and East-Lothian and Fifeshire coalfields. In 1857 he was promoted to the rank of Geologist, and he co- operated with (Sir) Archibald Geikie in producing their well-known volume on ‘The Geology of the Neighbourhood of Edinburgh,’ published in 1861. His work in association with Geikie and John Young laid the foundation of our present knowledge of the Carboniferous succession in Southern Scotland. In 1861 Howell returned to England, where he surveyed Jurassic areas in Northamptonshire, Bedfordshire, and Huntingdonshire. He then supervised the staff engaged in the survey of the North-Hastern counties, and was promoted to the rank of District Geologist in 1872. Ten years later he succeeded (Sir) Archibald Geikie as Director for Scotland, and eventually in 1888 became Director for Great Britain. He retired in 1899, after nearly half a century of disinterested public service, highly esteemed by all his colleagues. He had been elected a Fellow of the Geological Society in 1853. An account of the official work of Mr. Howell, with a portrait, appears in the ‘Geological Magazine,’ dec. 4, vol. iv (1899) pp. 483-37 & pl. xxi. He died in June 1915. Wiritam ANDERSON, who was born at Edinburgh in February 1860, and died at Sydney (New South Wales) on May 30th, 1915, was elected a Fellow of the Geological Society in 1899. After studying in the University of Edinburgh, he joined the Geological Survey of New South Wales in 1886, and wrote many valuable reports on the geology and mineral resources of that State, besides undertaking a series of researches on water-supply. In 1893 he retired from the New South Wales Survey, and, after a few years’ part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. Ixi service as Mining Specialist to the Geological Survey of India, he proceeded to South Africa. From 1899 to 1905 Anderson was Government Geologist of Natal, and he published three important reports on his work, with descriptions of the fossils by various specialists. After leaving South Africa, he returned to Scotland, but the climate proved too severe for his failing health, and he finally retired to Sydney. He was an excellent observer, full of enthusiasm for our science, and gained the esteem of all who knew him. South African geology loses one of its most active workers by . the premature death of Herpert Kynaston on June 28th, 1915. He was born on July 19th, 1868, at Durham, and educated at Eton and at King’s College, Cambridge. The Worts Travelling Fund was awarded to him in 1892, and he began his geological career by studying the Cretaceous Gosau Beds of Austria, of which he contributed an account to the Quarterly Journal of the Geological Seciety in 1895. From 1895 to 1902 he was engaged on the Geological Survey of Scotland, and in February 1903 he became Director of the Geological Survey of the Transvaal. Later, on the formation of the Union of South Africa, he assumed the direction of the newly-constituted Geological Survey under the Department of Mines. He carried on his Survey work with enthusiasm and success, and his annual reports were looked forward to no less by the student of pure geology than by the practical miner, whose needs in such a country are naturally paramount. Mr. Kynaston was of a retiring disposition and little known outside his depart- ment, in which he was universally esteemed ; but he took part in the proceedings of the Geological Society of South Africa, and was its President in 1908. He was elected a Fellow of the Geological Society in 1894. The Hon. Rogperr Marsnam (since 1893 Marsyam-Town- SHEND), who was born on November 15th, 1834, and died in April 1915, took a deep interest for many years in the study of stratigraphical geology, and in 1877 presented his valuable collection of British fossils to the British Museum. When in Brazil as Attaché at Rio de Janeiro, 1855-59, he obtained some Cretaceous fishes from Ceara, which he also gave to the British Museum. He was elected a Fellow of the Geological Society in 1859. Ix PROCEEDINGS OF THE GEOLOGICAL society. | vol. Ixxu, Sir SanprorpD Fremine, who was born at Kirkcaldy (Fife) in 1827, and died at Halifax (Nova Scotia) on July 22nd, 1915, became a Fellow of the Society in 1877. During the preliminary surveys and the early years of its construction, he was Engineer- in-Chief of the Canadian Pacific Railway. He was also associated with other railway enterprises in Canada; and his important public work was recognized by the conferment on him of the Knight Commandership of the Order of St. Michael & St. George in 1897. His interests were many and varied, and geological science, both theoretical and practical, was among them. Wiriiam Gryitits Apams, born on February 16th, 1836, was elected a Fellow of the Geological Society in 1865. He was a distinguished mathematician and physicist, and Professor of Natural Philosophy & Astronomy at King’s College, London, from 1865 to 1906. He became a Fellow of the Royal Society in 1872, and an account of his work is published in the Proe. Roy. Soc. ser. A, vol. xei (1915) pp. lxin, lsiv. He died in April 1915. Ricnarp AssHETon was a distinguished vertebrate embryo- logist, interested in all branches of biological and geological science. Born at Downham Hall (Laneashire) on December 23rd, 1563, he died at Grantchester, Cambridge, on October 24th, 1915. He had been elected a Fellow of the Geological Society in 1886, and of the Royal Society in 1914. Herpert Srantey Briony, who was elected a Fellow of this Society in 1911, died on June 6th, 1915, at the early age of 27. He was born in India, and educated at University College, London, graduating as B.Sc. at London University. He joined the Geo- logical Survey of India as Assistant-Superintendent in 1911, and was engaged chiefly in Burma and Kashmir. Forrescur Winiram Mizert, who was born in 1833, and died on February Sth, 1915, became a Fellow of the Geological Society in 1900. He was a well-known authority on the Foraminifera, and contributed an account of those found in the St. Erth Clay to the Palzontographical Society’s Monograph of the Foraminifera of the Crag. part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. lxill AnpDREW Duniop, who graduated as M.D. at Edinburgh Uni- versity in 1863, resided for 47 years in Jersey, and devoted much of his leisure to the study of the geology of that island. He contributed papers to the Quarterly Journal of the Geological Society on the Jersey Brick-clay and ou Jersey Raised Beaches in 1889 and 1893 respectively ; and so recently as November 1914 he communicated to the Society another paper on a raised beach on the southern coast of Jersey. He became a Fellow of the Society in 1874, and died at the age of 73 on December 30th, 1915. GEORGE HENRY HoLiinawortnH was an active member of the Manchester Geological Society, of which he was Treasurer for many years and President in 1903-1904. He was especially interested in the geology of coal-mining, but he also studied other geological questions in Lancashire, and in 1881 he contributed to the Quarterly Journal of the Geological Society a description of a peat-bed interstratified with Boulder-drift at Oldham. He was elected a Fellow in 1879, and died in April 1915. Brensamin Hongarr, who was born at Leeds in 1838, became a Fellow of the Geological Society in 1877. He devoted the leisure of his long life to a study of the geology and natural history of the district round Leeds, and made several contributions to the Transactions of the Leeds Geological Association. A portrait and a list of his writings are published in ‘ The Naturalist’ for April 1915 (pp. 145, 146). WiLLiaAM SIMPSON was an active member of the Yorkshire Geological Society and the Yorkshire Naturalists’ Union. He was especially interested in glacial geology, but also published notes on borings in the Millstone Grit at Halifax, where he resided until 1908. He was elected a Fellow of the Geological Society in 1893, and died at Catteral Hall, near Settle, in March 1915, aged 56. Joun Turner Horprack resided at Norwich, and for a long period took an active part in the public life of the city. He was a Member of the Museum Committee, and a valued supporter of the local scientific societies. In 1899-1900 he was President liv PROCEEDINGS OF THE GEOLOGICAL SOCIETY. |[ vol. Ixxu, of the Norfolk & Norwich Naturalists’ Society, and communi- cated to it in 1906 a paper on excavations in the Castle Mound. He was elected a Fellow of the Geological Society in 1900, and died on November Ist, 1915, aged 67. Henry Roreg, who was born on February 15th, 1839, and died on March 2nd, 1915, became a Fellow of the Society in 1890. He was a distinguished waterworks engineer, and constructed important works for the water-supply of many English towns. In 1905 he visited and reported on the water-supply of British Hast Africa. Ricuarpd Kerr, who was elected a Fellow of the Society in 1882, was engaged for many years in educational work at Folkestone, and subsequently became well known as a popular exponent of science. He died at Mitcham (Surrey) on May 21st, 1915. WitiraMm Hurron WixitaMs, who was born at Clifton, Bristol, in 1875, and became a Fellow of the Society in 1905, was a mining engineer who had done much work in Korea, Southern Manchuria, and India. He helda commission as Captain in the East Surrey Regiment, and was killed in action in France in May 1915. I have also to record the death on December 17th, 1915, of Wiri1am Ruprerr Jones, who was Assistant in the Geological Society’s Library from 1872 until 1918. He was born in 1855, the eldest son of Prof. T. Rupert Jones, F.R.S., and inherited his father’s retentive memory, which enabled him to give most valuable help to the Fellows using the Library. From its beginning until 1912 he prepared the slips for the Society’s annual Record of Geological Literature. part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. Ixv ‘Tue Use or tur Higner VERTEBRATES IN SE RAPHICAL GEOLOGY. JA a Tie study of fossil fishes, to which I referred last year, seems to show that each of the successive dominant groups is sharply distinguished from its immediate predecessor by some fundamental character. marking an advance towards the extreme adaptation for locomotion in water, which was ultimately attained in the Cretaceous Period. It isalso evident that various members of each of these successive groups soon became specialized for every possible mode of lite in the circumstances of the time. Fishes of the same general outward appearance and habit have thus originated repeatedly from progressively higher groups; similar adaptations have recurred with only minor differences ; and nearly the same changes, though perhaps with increasing intensity, have always marked the approach to racial old age. These phenomena are, indeed, so remarkable, that the question arises as to whether animals of apparently the same family, genus, or species may not originate more than once from separate series of ancestors. We may even hesitate further in deciding whether or no the really fundamental advances in life at successive periods have occurred more than once in the faunas of which they are respec- tively characteristic. The study of fishes, however, is scarcely sufficient to solve these problems, because the large majority of the fossils are marine, the animals would spread rapidly and widely, and the limits of the seas in which they lived are never clearly recognizable. The higher vertebrates, which inhabited the land, seem to be a much more hopeful source of necessary facts; for the land has always been subdivided into well-defined ar eas, aint by seas, mountains, and deserts. Animals in these sasrerel areas must often have deve- loped independently for long periods; alterations in the barriers ean be detected by the geologist when he studies the migrations and mingling of faunas ; while, as researches progress, the varying geography of successive periods may be more or less successfully restored. Like the fishes, the terrestrial vertebrates have advanced by successive fundamental steps towards perfection in powers of locomotion, and during this progress have at each stage diverged into various analogous specializations. They have, indeed, advanced one step farther by the final increase in the relative size and Ole DNL e Ixvi PROCEEDINGS OF THE GEOLOGICAL SocrIETY. | vol. lxxii, efficiency of the brain, which culminated in. Man. If, in these circumstances, the same type of animal has originated more than once, it should be possible, in some cases at least, to discover the phenomenon and determine its limitations. Although the enquiries involved are almost entirely biological, it is especially important for the geologist to take note of them, because students of shells from the modern standpoint are unanimous in recognizing what they term homcomorphy. They continually find ammonites, gastropods, and brachiopods, for example, which are essentially identical when full-grown but differ completely in their early stages. When the fossils happen to be perfect, these early stages are, of course, preserved and distin- guishable ; and if they reproduce, even only approximately, the ancestral condition in each case, they certainly suggest that the same type of shell may have arisen from more than one source. If this be so, shells from different horizons and widely-separated localities need very careful scrutiny before they can be used for correlating geological formations ; for, even if nearly similar, they may represent animals that have no really close affinity, and any. diagnostic features that they show may be inconspicuous points which have generally been overlooked as insignificant. For a clearer view of general principles, we therefore turn with ex- pectaney to vertebrate skeletons, which have much more numerous and tangible characters, and approach senility in more varied ways. Even among vertebrates it is by no means easy in every case to interpret the evidence that most concerns the geologist, and I need only refer to the supposed thylacines (Sparassodonts) and the horned tortoises (JMolania) of the Argentine Tertiary, which have often been quoted as specially strong proofs of the former existence of an Antarctic continent uniting the Australian and South American regions. So far as can be determined from their fossil remains, the Sparassodonts of the Santa Cruz Beds of Patagonia differ only in: the slightest particulars from the Pleistocene and existing thylacines of Australia, and they have been placed in the same family by American paleontologists. It is, however, to be noted that the thylacines and Sparassodonts are essentially identical with the primitive Creodonta, which are known to have ranged over all the continental land of the Northern Hemisphere at the beginning of the Tertiary Era. They only differ from these early mammals in certain senile specializations which might be expected in any long-lived group ; part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. Ixvii and, as these specializations are not altogether the same in the two cases, it is probable that they have arisen independently. Again, Miolania is a senile heavily-armoured form of the Pleurodiran Tortoises, which had a universal distribution in the late Secondary and early Tertiary Eras. The arrangement of the bony bosses on the skull is not absolutely the same in the species from Australia and Patagonia, and these indubitable marks of racial old-age may therefore have been acquired by separate groups’ in different regions. In other words, the essential identity of these land-animals does not prove a former direct connexion between Australia and South America; they may be merely survivors of cosmopolitan races at the two extremes of their former range, with certain inevitable marks of senility. In making comparisons, indeed, it is not enough to distinguish the fundamental and merely adaptive characters of animals; it is also essential to note separately those characters which depend on the early, mature, or senile position of the particular animals in the evolving series to which they belong. In the present state of our knowledge there seems to be only one case in which we begin to have materials for forming a judg- ment as to whether fundamental advances—the ‘ expression points’ of Cope—occur more than once. I refer to the acquisition of warm blood and its correlatives or consequences: which started the eareer of the mammals. The only reptiles that ever made a close approach to mammals in their skeleton existed during the Permian and Triassic Periods, and they were precisely intermediate between the Paleozoic Amphibia and the Mesozoic Mammalia. It is, therefore, presumably in Permo-Triassic times that mammals arose. These intermediate or-Theromorph reptiles, however, were spread widely over many lands. Their remains were first found in the Karoo formation of South Africa, were afterwards recognized in India, Scotland, Northern Rusgia, and Central Kurope, and also proved to occur in great numbers in the Permian of North America. Traces of them have also been met with in Southern Brazil. The African, Asiatic,and Huropean remains belong to animals so closely sunilar, that it is probable that they lived on a continuous land- area; but the American forms, although in the beginning not unlike, soon began to evolve into several groups which are almost or completely unknown in the Old World. The American Thero- morphs, therefore, probably flourished in isolation. Large collec- tions from both regions have been studied during recent years by e2 Ixvil PROCEEDINGS OF THE GEOLOGICAL SocIETY. [ vol. Ixxu, Dr. R. Broom, Dr. E. C. Case, and others, and a critical summary of the results was published by Dr. Case last summer.! It now appears that all the specializations in the American groups were in the direction of higher reptiles; while all those in the South African groups made a progressively closer approach to mammals, and as nearly as possible culminated in typically mammalian skeletons. Hence, although we have evidence of two possible sources of mammals, only one appears to have produced them. Secondly, consider a case of less fundamental character, the origin of the Monkeys or lower Anthropoidea. It is generally admitted that they arose from the next lower grade, that of the Lemurs or Lemuroidea, which were almost universal in their distribution over the great continents at the beginning of the Tertiary Era. It is also suspected (though the paleontological evidence is still very scanty) that the two markedly distinct groups of Monkeys, the Platyrhines in America and the Catarhines in the Old World, were derived independently from the Lemurs in these two separate continental areas. If this be so, a zoologist’s ‘sub- order’ has arisen by two parallel, though readily distinguishable, developments from an earlier and lower ‘sub-order.’ Now, the surviving Lemurs of the Asiatic and African regions and Mada- gascar have lived on from early Tertiary times practically un- changed ; but there is one isolated area, the island of Madagascar, where some families during the later Tertiary periods attained remarkable specialization. Unfortunately, we only know the Pleistocene and Holocene members of these families that are discovered in the caverns and swamps of Madagascar; but they represent the culminating genera, and suffice to indicate what happened. In this isolated region the highly-specialized late Tertiary Lemurs curiously resembled in some characters both the Platyrhine and the Catarhine Monkeys. ‘The first portion of skull of Nesopithecus, in fact, appeared to so experienced an observer as Dr. Forsyth Major so monkey-like that he originally placed it among the Monkeys, in a new family intermediate between the South American Cebidee and the Old-World Cercopithecide. I+ is now clear, however, from the researches of Dr. G. Grandidier and Dr. H. F. Standing, that none of these animals really passed beyond the typical Lemur-grade. They eventually grew large, 1 #. C. Case, ‘The Permo-Carboniferous Red Beds of North America & their Vertebrate Fauna’ Publication 207 (1915) of the Carnegie Institution of Washington, p. 121. a part 1} ANNIVERSARY ADDRESS OF THE PRESIDENT. ]xix some of them (Megaladapis) almost rivalling donkeys in size; they also diverged for various modes of life, a few of them even entering the proper sphere of hoofed animals. Removed from the stress of competition with other mammals such as swarmed on the continents, they flourished in luxurious ease ; and we are tempted to speculate whether it was not the strenuous competition in the crowded continental forests of America and Africa (or Asia) that made ‘the difference, and led to the increase of brain-power which is the specially distinctive characteristic of Monkeys. If this be so, the lower race passed into the higher race only where the struggle for existence was keenest. Why the higher race never advanced further in America, while it began at once to give rise to the man-like Apes in the Old World, is still an unsolved problem. The case of the Anthropoidea suggests that a widely-distributed group of early Tertiary mammals experienced in three separate regions an essentially-similar initial impulse to become a higher group, evolved on parallel though not identical lines, and ended at ‘three different stages in its onward progress. It is, therefore, interesting to consider the geological history of some of the families of Ungulata which are already sutticiently well known by a succes- sion of fossils to reveal the main episodes in their career. This history is especially instructive, because all the surviving groups originated in the Northern Hemisphere on two or three large continental areas which were sometimes united, at other times separate; and the fossils are beginning to show how and when the various connexions and isolations occurred. Northern Africa, probably with part of Southern Asia, is one of the areas on which Ungulates made an independent start; Europe, with North-Central Asia, is another ; while North America is the third. The Rhinoceroses are a well-marked family, of which numerous fragmentary fossil remains can easily be recognized and compared. They seem to have arisen at the beginning of the Eocene Period from some small generalized Perissodactyl which had a slender snout and a regular close series of 22 low-crowned teeth in each jaw; and, as ages passed, the nasal bones became enlarged and thickened to support a dermal horn, the front teeth tended to disappear as the lips became more prehensile, while the molars and premolars increased in effectiveness for grinding hard and dry herbage. As in other Perissodactyls, the premolars were at first comparatively simple, but afterwards gradually approached the xx PROCEEDINGS OF THE GEOLOGICAL sociEry. |[vol. lxxu, molars in complexity ; while the successive representatives of the family showed a progressive increase in bodily bulk. ‘These changes took place almost simultaneously both in the Old World and in North America, though in the latter region the whole family died out in early Pliocene times as soon as a rudimentary nasal horn was beginning to appear. After the earliest stage, however, the European series at least can readily be distinguished from the American series by the premolar teeth, and there must have been independent evolution in the two regions. According to Prof. O. Abel, the oldest known recognizable member of the family is the small Prohyracodon from the Middle Eocene of Transylvania ; and both this and ‘the later Mpiaceratherium, from the Lower Oligocene of Northern Italy, agree with the earliest North American Rhinoceroses in having the foremost premolar more nearly like a molar than is the hindmost premolar. They thus show the first step in the complication of the premolars from front to back, which, according to Prof. H. F. Osborn, is the direction of this complication during geological time in all the American Rhinoceroses. On the same account, they differ from all the known Rhinoceroses which followed them in Europe: for in these the complication of the premolars always began with the hindmost, and proceeded forwards. Not only can two distinct progressive groups be thus recognized in two distant regions, but more than one line of development can also be traced within each. The fossils, indeed, seem to justify the conclusion that the Rhinoceroses arose from a common stock, but evolved in several different ways and at varying rates towards the same goal of specialization. It is curious that extreme specialization in the family was reached only in the Old World, and still more curious that the large-horned woolly rhinoceros (2h. antiquitatis), which lived in the Arctic Regions with the mammoth, never accompanied that animal to North America. Equally interesting is the exceptional case of high specialization in the Old-World Pleistocene Hlasmo- therjum, in which the extreme deepening, enlargement, and complication of the molar and premolar teeth seem to have been correlated with the reduced development of the epidermal horn. The large bony boss on the frontals of this animal was probably covered only by a very thin capping of horn. At least, it is difficult to suggest any other interpretation for such a horn, which was found isolated a few years ago by the Prussian Geological part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. Ixx1 Survey, in a superficial deposit in the Luckau district south of Berhn. A detailed study of the tooth-pattern in the successive groups of extinct Horses, seems to reveal the same phenomenon that 1s observable in the Rhinoceroses—the approach to nearly-identical extreme specialization by several separate lineages in approximately the same time. ‘The earliest Eocene forerunners of the Horses are essentially similar in Europe and in North America, and some of the later forms also exhibit many resemblances which may be due to migrations from one region to the other; but the most ex- haustive modern researches leave no doubt that the gradual reduction of the foot to a single toe and the deepening of the teeth for effective grinding of hard food, with various correlated specializations, occurred in several distinct groups of Horses in different localities. The climax was reached during the Pliocene Period, both in Europe and Asia and in North America; while the greatest diversity of form appeared in the latter region and among the immigrants to South America, where all Horses died out before historic times. Even in families of more restricted geographical range, the same unswerving tendency towards a fixed goal is very clearly recognizable. Among the Camels, for instance, which seem to have accomplished the whole of their evolution in part of North America, the characteristic cushioned foot appears to have been produced more than once. All the small early ancestors of the Camels had feet like those of deer or gazelles, with pointed toes ; and Prof. W. B. Scott has observed that even the Lower Miocene genera, which were already differentiated into two groups, still exhibited the same primitive feature. All the later Miocene Camels of both these groups, however, possess the irregularly- nodular ungual phalanges which indicate the presence of the cushions or pads. It is thus evident that two progressive series independently acquired one and the same structure. A study of the extinct representatives of several other familiar Ungulata has led to the discovery of many facts which tend to confirm the results just mentioned. In every case, either the feet become perfectly adapted for rapid locomotion over hard ground, or the teeth acquire more grinding power by deepening, folding, and the frequent infilling of the hollows with cement; or both these progressive adaptations take place at the same time. When, however, this evolution first started in the Eocene Period, the / Ixxii PROCEEDINGS OF THE GEOLOGICAL soctety. | vol. Ixxu, | common ancestors of the groups that were thus destined to flourish were accompanied by other generalized Ungulates which began to acquire running feet without any of the requisite changes in the arrangement of the wrist- and ankle-bones, and often showed a deepening or other complication of the grinding teeth without any infilling of the resultant hollows by cement. These animals with ‘inadaptive modifications’ (as Kovalevsky termed them) were soon handicapped in the race with their better-endowed contemporaries, and all became extinct before they had advanced far. In South America, on the other hand, which was an isolated centre of mammalian development during the greater part of the Tertiary Era, all the early Ungulates began to advance in the ‘inadaptive ’ manner, without any more efficient contemporaries and competitors. Here we are, therefore, able to follow to the end | a course of evolution which was abruptly stopped in the Northern | Hemisphere. It resulted in less variety than is observable among ordinary Ungulates, but some of the later genera are strange mimics of the Rhinoceroses and Horses in outward shape. At least one of the bulky three-toed forms acquired a rhinoceros-like horn: and some of the small one-toed forms exhibited a greater reduction of the lateral toes even than in the horse. The molar teeth were also sometimes invested with a little cement, but they never became such effective grinders as those of the Horses and Elephants. The latest of these South American animals, indeed, soon disappeared when a land-connexion in the Pliocene Period allowed the animals of North America to invade the Southern Continent and compete with them. None of the South American Ungulates developed the sym- metrical pair of toes such as characterizes so large a proportion of the northern forms, but their single enlarged toe, which became the centre of symmetry, was always the third, as in the ordinary tapirs, rhinoceroses, and horses. It is, therefore, interesting to note that among the Australian Marsupials, which have inde- pendently developed a hind foot for rapid locomotion or leaping on hard ground, the largest toe is not the third, but the fourth. So far, unfortunately, we lack nearly all the ancestors of these animals; but available evidence seems to show that they are descended from tree-dwellers in which the first digit had become opposable. When they began to live habitually on the ground and the foot became modified accordingly, this digit dwindled to a projecting rudiment (as preserved in Diprotodon), and the axis of part 1 | ANNIVERSARY ADDRESS OF THE PRESIDENT. Ixxi the foot was displaced outwards from the third to the fourth toe. In other words, an essentially ungulate foot was produced as usual, only disturbed by an initial twist. From these and many similar facts in mammalian evolution, we may therefore conclude that parallel development or ‘homco- morphy’ undoubtedly occurs. Some of the changes are adaptations to the same altering modes of life, perhaps also to the same progressive modifications in the environment; among which may be enumerated the increasing efficiency of the teeth for grinding and of the feet for running. Other changes are the inevitable marks of racial maturity or old age, such as increase of size, the development of excrescences, and the reduction in number of the teeth. While, however, both these series of changes always take place in approximately the same order and may be used to mark the successive periods of Tertiary time by a paleontologist who is accustomed to deal with such evidence, there is still no reason to suppose that identical animals have arisen from more than one source. Distinct families or genera which may be difficult to separate on superficial examination, are readily recognized in most eases when studied in detail with our present knowledge of general principles. It is, indeed, usually possible to distinguish between traits of heredity that are fundamental, and those that are merely adaptive or depend on racial antiquity. The new palzontology is thus as useful in relation to world-problems, as was the old paleontology in determining the relative ages of the rocks in the restricted Huropean area to which it was first applied. The adaptive characters just mentioned change with much rapidity, and allow a tolerably detailed subdivision of the series of strata which contain the fossils exhibiting them. It is therefore interesting to notice that some other characters which can scarcely be correlated with utility or efficiency, persist without change for comparatively long periods and are of little value for stratigraphical geology. Huxley long ago directed attention to the problem of ‘persistent types,’ as he termed them, and most of them are still as inexplicable as they were when first considered. Only in the ease of certain fishes have I suggested that advance was stopped by the appearance of various modifications or specializations in the wrong order. It may be that single peculiar characters persist when they cannot be affected by Natural Selection. One of the most striking examples was described by Egerton in the Society’s Journal for 1871 in a Chimeroid fish from the Lower Lias of Ixxiv PROCEEDINGS OF THE GEOLOGICAL society. {[vol. lxxi, Lyme Regis, which he named Ischyodus orthorhinus (now known to be Prognathodus). The prolongation of the snout in this early Jurassic fish is of precisely the same remarkable shape as that in the existing Chimeroid Callorhynchus. Notwithstanding the many changes which have occurred in Chimeroid fishes since that remote period, including a total replacement of the genera, the pattern of snout in one group at least has persisted. Again, the existing Lamnid sharks are characterized by a curious dwarfing of the third or fourth tooth, or both, on each side of the upper jaw. A study of associated sets of teeth of some of the earliest- known members of the family from the Chalk, proves that this feature was already established even in Cretaceous times: not only in the genera which have survived, but also in the extinct Coraz. Finally, in the Society’s Quarterly Journal for 1910, I pointed out that in the Lower Jurassic carnivorous dinosaurian, Megalosaurus, three or four pairs of the teeth in front of the lower jaw are diminutive, perhaps functionless. Last year, through Mr. W. E. Cutler, the British Museum received from the Upper Cretaceous of Alberta (Canada) the jaws of a gigantic Megalosaurian, evidently of another genus, in which exactly the same reduction of the teeth at the mandibular symphysis oceurs. Through a wide range of time and space, therefore, this small and apparently insignificant feature persisted, and even passed from one genus to another. The strength of heredity is, in fact, one of the most remarkable phenomena with which a paleontologist is repeatedly impressed. Even when one great group gives rise to another, the later type often seems to be handicapped at first by the inheritance of some characters which are no longer congruous. The large majority of the Permian and Triassic Reptiles, for example, although true land-animals, still retained the large head and short neck which were well adapted to the aquatic habits of their amphibian and piscine ancestors. This was first strikingly shown by Seeley’s restoration of the South African Aeirognathus, afterwards by Boulenger’s sketch of the Scottish Velerpeton, and has been emphasized more recently by the numerous restorations of Permian reptiles from North America by Williston, Case, and others. Similarly, when Reptiles passed into Mammals, the relatively-large tail of the gliding or swimming animal was no longer needed; but it persisted in nearly all Mammals at least through the first half of the Eocene Period, and even at the . part 1] ANNIVERSARY ADDRESS OF THE PRESIDENT. xxv present day the abdominal region gradually tapers into the thick root of the large tail in such lowly types as the Marsupial Thylacinus and the Kdentate Orycteropus. Archeopteryx suggests that the persistence of the reptilian tail was also a handicap to the earliest birds. It may be that the same conservative tendency is the origin of various other apparently incompatible structures, for which adaptive explanations have been sought in vain. For instance, it has been surmised that the great bony brow-ridges of apes like the gorilla are needed to resist the strain produced by the working of the powerful jaws; but the same explanation will not apply to the nearly similar forehead of Neanderthal Man, in which the jaws are no heavier than those of many modern men. This must be a case either of direct inheritance or (as at present seems more probable) of curious reversion in a higher group to a tendency characterizing an ancestral lower group. Many similar instances might be cited. The paleontologist is, indeed, now continually tempted to trespass on the domain of the speculative philosopher. He has had to abandon the old vague methods of comparative anatomy by which the solid foundations of his science were first laid. He now sorts out characters into several categories before beginning to compare them, and arrives at his interpretations in accordance with certain general principles which I have tried to illustrate. If, as Sir Jethro Teall has said, ‘the state of advancement of a science must be measured, not by the number of facts collected but by the number of facts co-ordinated,’ Paleeontology is well in the forefront of progress. Its devotees may be wrong in some of their broader speculations, but during recent years they have rarely met with new facts which could not be reconciled with the general scheme of things developing in their minds. The value of its results for the purposes of geology must thus be constantly increasing. Leaving high philosophy, it now only remains for me, in conclusion, to thank the Fellows of the Geological Society for the great honour which they conferred upon me two years ago when they elected me to be their President. I also wish to express my appreciation of the kind help and forbearance of the Officers, Council, Fellows, and Permanent Staff, which have made my term Ixxvi PROCEEDINGS OF THE GEOLOGICAL socrnty. [vol. lxxu, of service to the Society one of the most pleasant and memorable episodes of my life. My own special studies are so completely on the borderland of our science, that I fear I must sometimes have wearied you with matters that seem almost extraneous; but it is one of the greatest charms of geology that it needs co-operation with the whole circle of the sciences to further its aims. You have now turned from paleontology to petrology, and I have much pleasure in resigning the Presidential Chair to an old friend, an accomplished Fellow who has indeed delved deeply into the mineral structure of the Earth. part 11 PROCEEDINGS OF THE GEOLOGICAL SOCIETY. xxvii February 23rd, 1916. Dr. ALFRED Harxker, F.R.S., President, in the Chair. William David Purdy, 7 Christchurch Terrace, Chelsea, S.W., and George James Roberts, Noyna, Avenue Rise, Bushey (Hertfordshire), were elected Fellows of the Society. The List of Donations to the Library was read. The following communication was read :— ‘On the Origin of some River-Gorges in Cornwall and Devon.’ By Henry Dewey, F.G.S. (Communicated by permission of the Director of H.M. Geological Survey.) Lantern-slides were exhibited in illustration of the above paper. A nodule surrounded by a layer of cone-in-cone structure, without any trace of calcite, from Lower Paleozoic rocks near Machynlleth (Montgomeryshire), was exhibited by Dr. A. Smith Woodward, In pldiaera« Vele(Crise March Sth, 1916. Dr. Atrrep Harker, F.R.S., President, in the Chair. Wilfrid Baker, Royal Grammar School, Worcester; and Arthur G. Pomeroy, M.A., B.Eng., 21 Orange Street, Haymarket, S.W., were elected Fellows of the Society. The List of Donations to the Library was read. The PresrpEn’ referred with regret to the death, on March 3rd, of Prof. Jonn Westey Jupp, C.B., LL.D., F.R.S., Past President of the Society. He spoke of the value of Prof. Judd’s contri- butions to geological science, and of his eminence as a teacher of the science, and stated that the Society was well represented at the funeral. ' Dr. AuBREY Srrawan, F.R.S., Director of H.M. Geological Survey, exhibited and described briefly a set of specimens from the Western Front, illustrating the character of the rocks in which trenches, tunnels, etc. are being dug. They included specimens from the Cretaceous and Tertiary formations showing remarkable similarity in characters to the contemporaneous forma- tions in Britain. 4 Ixxvil PROCEEDINGS OF THE GEOLOGICAL socrery. [vol. lxxu, The following communication was read :-— “On some Insects from the British Coal Measures.’ By Herbert Bolton, M.Sc., F.R.S.E., F.G.8., Reader in Paleontology in the University of Bristol. Lantern-slides and specimens were exhibited in illustration of the above paper. March 22nd, 1916. Dr. AurreD Harxker, F.R.S., President, in the Chair. Lieut. Kenneth Neville Moss, B.Sc., Royal Engineers, Fieldgate, Walsall, was elected a Fellow of the Society. The List of Donations to the Library was read. Dr. A. SmirH Woopwarp, F.R.S., V.P.G.S., exhibited speci- mens of the problematical ichthyolite, Celorhynchus, from an Eocene deposit in the Ombialla district (Southern Nigeria), and discussed the nature of this fossil. Microscope-sections of the well-preserved Nigerian specimens confirmed W. C. Williamson’s determination that Calorhynchus is an essentially dermal struc- ture. A similar section of part of the rostrum of the teleostean fish Blochius, from the Upper Eocene of Monte Bolca, near Verona, showed an almost identical structure. The precise nature of this rostrum remained to be determined, but there could be no doubt that the so-called ‘ Celorhynehus’ is the corresponding part, either of Blochius or of an alhed genus. The followimg communication was read :— ‘The Pseudo-Tachylyte of Parijs (Orange Free State) and’ its Relation to “Trap-Shotten Gneiss”” and “ Flinty Crush-Rock.”’ By S. James Shand, D.Se., F.G.S., Professor of Geology in the Victoria College, Stellenbosch (8.A.). April 5th, 1916. Dr. ALFRED Harker, F.R.S., President, in the Chair. The List of Donations to the Library was read. The PRESIDENT announced that the Council had awarded the ' part 1] PROCEEDINGS OF THE GEOLOGICAL SOCIETY. Ixxix Proceeds of the Daniel-Pidgeon Fund for the present year to Joun Kaye Cuariteswortn, M.Sc., Ph.D., F.G.S., who proposes to conduct researches in connexion with the Glaciation of Donegal. The following communication was read :— ‘The Picrite-Teschenite Sill of Lugar (Ayrshire) and its Differ- entiation.” By George Walter Tyrrell, A.R.C.Sc., F.G.S. Lantern-slides were exhibited in illustration of the above paper. A Bronze Bust of Sir Charles Lyell, presented to the Society by Mrs. J. W. Judd, through Prof. W. W. Watts, was also exhibited. May 10th, 1916. Dr. AnFRED Harker, F.R.S., President, in the Chair. Keir Arthur Campbell, B.A., Trinity College, Cambridge ; Ednyfed Wynne Hughes, M.Sec., The Grange, Lesness Park, Belvedere ; and Christopher Luke Waite, Newlands, Pontefract Road, Castleford, were elected Fellows of the Society. The List of Donations to the Library was read. The followmg communications were read :— 1. ‘ Carboniferous Fossils from Siam.’ By F. R. Cowper Reed, MEAS se.D. E.G-S: 2. ‘The Lurgecombe Mill Lamprophyre and its Inclusions.’ By Herbert Gladstone Smith, B.Sc., F.G.S. Carboniferous fossils from Siam were exhibited by F. R. Cowper Reed, M.A., Sc.D., F.G.S., in illustration of his paper. Lantern-slides, rock-specimens, and microscope-sections were exhibited by H. G. Sinith, B.Sc., F.G.S., in illustration of his paper. Sapphire-bearing xenoliths, from a Tertiary sill in the Island of Mull, were exhibited on behalf of H.M. Geological Survey. ~ May 24th, 1916. Dr. AufFrep Harker, F.R.S., President, in the Chair. The List of Donations to the Library was read. Dr. A. Smrru Woopwaxp, F.R.S., V.P.G-S., exhibited De- vonian fish-remains from Australia and the Antarctic Ixxx PROCEEDINGS OF THE GEOLOGICAL SOCIETY. | vol. Ixxu, Regions, and discussed our present knowledge of the Devonian fish-fauna of the Southern Hemisphere. So far as is known, there are no strange elements in this fauna, and the remains Hiccomerad closely resemble those met within the Northern Hemisphere. Even the rocks are very similar to those containing the corresponding fossils in the Northern Hemisphere. There is, as yet. no satisfactory evidence of the basal Devonian fish-fauna such as occurs in the Downtonian of England and Scotland; but both Lower and Upper Devonian forms occur in Victoria and New South Wales (Australia ). A Coccostean related to Phlyctenaspis trom Gippsland (Victoria ), and another related to Macropetalichthys from Goodra Vale (New South Wales), may be regarded as Lower or Middle Devonian ; typical plates of Bothr iolepis from the Harvey Range (New South Wales) indicate an Upper Devonian fauna. The fish-remains obtained by the ‘ Discovery’ Expedition in Granite Harbour (Antarctica) comprise Bothriolepis, another Ostracoderm related to Byssacanthus, Acanthodian scales, Selachian dermal tubercles, a Coceostean, scales of Osteolepidie, and scales of a very small Paloniscid. They must be regarded as Upper Devonian. Dr. Smith Woodward expressed his indebtedness to the Govern- ment Geologist of New South Wales for the loan of the Australian specimens exhibited. Mr. R. Butten Newton exhibited some so-called Orbitoidal Limestones from Dutch New Guinea, the microscopical structures of which were shown by lantern-illustrations. The specimens were collected by Dr. A. F. R. Wollaston during his expedition to that country in 1912-13, on the snow-line of Mount Carstensz at a height of 14,200 feet, this mountain forming the highest elevation mee New Guinea, with an altitude of rear 16,000 feet. The foraminiferal organisms determined in this material included five species of Lepidocyclina (sumatrensis, martine, neodispansa, murrayana, and cf. tnsule-natalis), Amphi- stegina, Carpenteria, Cycloclypeus (ct. orbitoideus), ete.; the marine alga or nullipore, Lithothamnium, was also largely repre- sented. This assemblage compares favourably with that which characterizes rocks of similar age in other Pacific regions, such as Christmas Island (Indian Ocean), Formosa, the Philippines, Borneo, Celebes, Sumatra, Nias, Timor, and Australia, besides indicating a Miocene origin. It was pointed out that the genus Orbitoides of A. VOrbigny had been restricted by Schlumberger (relying on the researches of Giimbel, Verbeek, and others) “to species having rhomboidal equatorial chambers and belonging only to Cretaceous times ; species furnished with rectangular chambers, and recognized as Or thophr agmina of Munier- Chains were limited to the Hlpcene and Oligocene formations ; while Giimbel’s genus Lepidocyclina, with rounded or hexagonal chambers, included species of Miocene and later age. As the result of a study of species from Borneo and the Philippines, Prof. Douvillé had proposed to divide Lepidocyclina part 1] PROCEEDINGS OF THE GEOLOGICAL SOCIETY. Ixxx1 into two sections—Hulepidina and Nephrolepidina: the first including forms of generally large size, recognized as Aquitanian ; the second for those of small dimensions, regarded as Burdigalian— these geological divisions representing the oldest stages of the Miocene System. This distinction, however, was not applicable to the New Guinea limestones nor to corresponding rocks from Christmas Island, as both large and small species occurred in association ; it was, therefore, suggested that the age of the New Guinea material might be referable to the later part of the Aquitanian. Several writers have already written on rather similar limestones from various parts of New Guinea, although we are indebted to Dr. K. Martin for the first announcement in connexion therewith: he reported the discovery in 1881 of Lepidocycline organisms from rocks found in the northern and south-western districts of the country (Geelvink Bay, islands of Kei, Aru, etc.), which he attributed to the older Miocene. ‘The same author also referred to the occurrence of similar organisms in Mount Wil- helmina, obtained by Dr. Lorentz, beneath which the Alveolina Limestone was identified, proving the existence of Hocene rocks. The Cycloclypeus remains, which are of frequent occurrence in the present material, bear a strong resemblance to Prof. Douvillé’s new genus and species from the Miocene of Borneo, known as Spiroclypeus orbitoideus. This genus was stated to have all the characters of Cyeloclypeus, but differs from it in the possession of superficial chamberlets in the shelly layers of the central region. Recent investigations, carried out by the speaker, had proved the presence of this character (hitherto unrecognized) in recent forms of Cycloclypeus from Funafuti; hence the retention of Spiro- clypeus now appeared to be unnecessary. A full report on this New Guinea collection by the speaker had been lately published, entitled: ‘Notes on some Organic Limestones, &c. collected by the Wollaston Expedition in Dutch New Guinea,’—forming No. 20 of a series of reports, and it was to be included in vol. ii of those reports. _ By permission of the Secretary of State for the Colonies, Mr. J. F. N. Green exhibited a manuscript report by Mr. J. H. Haglesome, C.M.G., on the Udi Colliery in Nigeria, containing a number of photographs and plans, and made some observations on the economic importance of this development in Equatorial Africa. There were placed with it for comparison two photo- graphs of the outcrop of manganese-ore in the Gold Coast, also discovered by Mr. A. E. Kitson, F.G.S. VOL. LXXEt. f lxxxil PROCEEDINGS OF THE GEOLOGICAL society. {[vol. lxxu, June 7th, 1916. Dr. AuFRED HarxkeEr, F.R.S., President, in the Chair. The List of Donations to the Library was read. Dr, F. L. Kircuty, M.A., F.G-.8., exhibited a representative set of Mesozoic fossils obtained from deep borings and pit-sinkings in Kent. The specimens were selected from the collections of the Geological Survey, by permission of the Director. The life of the successive zones present in the various sections was illustrated by the arrangement of the specimens in sequence, the series comprising a time-range from the Lower Lias up to the top of the Lower Greensand. The section revealed in the Brabourne Boring is remarkable for the range of formations found there in superposition, and may be regarded as the type-section for the study of the hidden Mesozoic succession in Kent. Specimens were exhibited from that locality and from the shafts at Dover, but these were supplemented by materials obtained more recently from various borings situated east of a line drawn from Folkestone to Canterbury. The speaker described the principal characters of the faunas* as developed in this area, and made incidental references to the nature and distribution of some of the associated rock-types. At some horizons, the molluscan assemblage assumes a particular aspect, by reason of the preponderance of species fitted for life amidst the special conditions of deposition. On the other hand, there are evolutionary phases which recur repeatedly with considerable uni- formity, and seem to arise independently of immediate surrounding conditions. Such are illustrated by the degenerative changes shown to occur in many ammonites, and by some of the forms repeatedly assumed by the members of separate series among Mesozoic oysters. The evidence of ammonites, so important for the purpose of zonal determination, is frequently forthcoming at these localities in Kent, even in borings of narrow diameter; but it is often necessary to rely entirely upon the aid afforded by the more abundant bivalves. Many of these, although belonging to un- described species, are found to have a limited vertical range, and by their distr ibution thr oughout this area, as well as farther afield, prove of much service in these correlation-studies. Specimens of many undescribed species have come to light, as well as others which are known from their occurrence in Continental localities, though not previously recorded in this country. A small series of Jurassic Cephalopoda from Kachpur (Russia), collected by the late G. ¥. Harris, F.G.S., at the time when the International Geological Congress met at Petrograd (1897), was exhibited by James Francis, F.G.S. part 1] PROCEEDINGS OF THE GEOLOGICAL SOCIETY. Ixxxil June 28th, 1916. Dr, ALFRED Harker, F.R.S., President, in the Chair. Herbert Theodore Mayo, B.A., care of Port Master, Rawal Pindi (India) ; and William Guy de Gruchy Warren, Capel Issa, Manordilo (Carmarthenshire), were elected Fellows of the Society. The List of Donations to the Library was read. The following communications were read :— . ‘Ona New Species of Hdestus from the Upper Carboniferous of Yorkshire. By A. Smith Woodward, LL.D., F.R.S., V.P.G-S. With a Geological Appendix by John Pringle, EGS. Paine Tertiary Volcanic Rocks of Mozambique.’ By Arthur Holmes, D.I.C., B.Se., A.R.C.Se., F.G.S. Dr. A. Srranan, F.R.S., exhibited cores from borings in Kent, showing pebbles of coal embedded in Coal- Measure sandstones. With the coal-pebbles occurred a few partly-rounded fragments of chert, and in one of these radiolaria had been identified by Dr. G. J. Hinde. The chert resembled that which had been described from Lower Carboniferous rocks elsewhere. Its occurrence suggested that the sequence of strata had been similar in South Wales and Kent, and, taken in con- nexion with the piping of the limestone-surface at Ebbsfleet and the absence of Millstone Grit in Kent, tended to confirm the view that there is unconformity between the Coal Measures and the Carboniferous Limestone in that county. Mr. F. P. Mennetr exhibited a geological sketch- map of the northern margin of Dartmoor. He said that the central part of Devon was to a great extent a terra incognita ; but, as regarded the fringe of altered Carbon- iferous rocks along the northern border of the Dartmoor granite, he had been led, in the course of observations originally concerned with the petrology alone, to the’ conclusion that it might prove possible to establish a definite order of succession. This was rendered feasible by the occurrence of some well-characterized bands of rock, especially limestones and tuffs, which were exposed in every good river-section. It was true that almost everywhere overfolds, sometimes accompanied by thrusts, were to be de- tected, and tended to make the observer somewhat doubtful of his ground. Nevertheless, it seemed impossible to escape the conclusion that, as one approached the granite from the north, continuously older rocks were met with, and the extremely Ixxxiv PROCEEDINGS OF THE GEOLOGICAL society. [ vol. lxxil, continuous character of some of the beds seemed to show that, despite all minor disturbances, the general sequence could be trusted. The comparison of the different lines of section leading up to Dartmoor showed them to be strikingly similar. The granite was, moreover, intruded all along at precisely the same horizon, and its direct offshoots never reached into the lower of the two important bands of limestone, but were confined to the altered shales at the bottom of the series, which afforded, where fresh, good examples of andalusite-hornfels. The series, which extends from south of Sourton to Drewsteignton, and perhaps right round to Doddiscombeleigh, appears clearly older than the shales which have been so carefully searched for fossils in the Exeter region by Mr. F. J. Collins. These last are considered to be of Pendleside age, and nowhere contain any traces of limestone. The probability is thus indicated that the distinctly calcareous series under consideration may represent part of the Carboniferous Limestone. It may be noted that, although a number of bands of epidiorite representing intrusions of dolerite occur roughly parallel to the strike of the sediments, the contemporaneous rocks are never of such basic character. The main band of tuff stretches from Lake, near Bridestowe, to beyond Sticklepath, and, of the numerous well-preserved rock-fragments that it contains, most are of rhyo- litic or trachytie character, with some which represent aitered andesites. Lantern-slides and specimens of Hdestws were exhibited by A. Smith Woodward, LL.D., F.R.S., V.P.G.S., in illustration of his paper. Lantern-slides, microscope-slides, and rock-specimens were ex- hibited by Arthur Holmes, D.I.C., B.Sc., A.R.C.Se., F.G.S., in illustration of his paper. THE QUARTERLY JOURNAL OF THE GEOLOGICAL SOCIETY OF LONDON, Vou. LXXII FoR 1916. 1. On a New Species of Epvesrus from the Uppnr Carpon- IFEROUS of YorKsuIRE. By Artuur Smira Woopwarp, LL.D., F.R.S., V.P.G.8. With a GrotogicaL APPENDIX by Joun Prinexe, F.G.S. (Read June 28th, 1916.) [Puate I. ] THe remarkable Upper Paleozoic fossil Hdestvs has already been proved to represent a row of symphysial teeth of an Elasmobranch fish, but it has hitherto been found only once in direct association with portions of jaws.! A second specimen, still more instructive, has now been obtained by the Geological Survey from the upper part of the Millstone Grit at Brockholes, near Huddersfield, and Iam indebted to Dr. Aubrey Strahan and Dr. F. L. Kitchin for the opportunity of studying it. The circumstances of its discovery are described in the Appendix by Mr. John Pringle. The new fossil, shown of two-thirds the natural size in Pl. I, fig. 1, displays a single example of Hdestus, with a detached dental crown and another fragment of the same form, near the tapering ends of a symmetrical pair of cartilages (¢) which evidently repre- sent a jaw. Whether they are upper or lower is uncertain, on 1 O. P. Hay, ‘ On an Important Specimen of Hdestus ; with Description of a New Species, Hdestus mirus’ Proc. U.S. Nat. Mus. vol. xlii (1912) pp. 31-38 & pls. i-ii. Q. J. G.S. No. 285. B 2 DR. A. SMITH WOODWARD ON [vol. Ixxu, account of the shortness of the portions preserved; but, as the anterior ends suddenly begin to taper and eventually become very slender, they are probably the ~pterygo-quadrates of the upper jaw. The cartilage is well calcified in very small tesserz, and, as shown both by the portions of jaws themselves and by remains in front of the fossil, the calcification penetrates more deeply than is usual in recent Elasmobranchs. The best-preserved outer surface of the cartilage, on the side of the specimen not shown in the figure, is shghtly marked with scattered fine pittings, such as have already been deseribed in Hdestus mirus.} The row of fused symphysial teeth (s) of the Hdestus type is bilaterally symmetrical, and arched almost in the form of a semi- circle. The eight teeth of which it is composed do not increase much in size backwards, and’ the depth of each crown slightly exceeds twice that of its root. The crown is laterally compressed and triangular in shape, much deeper than wide, with the sharp anterior and posterior edges nearly straight, and the postero-inferior angles much produced into slender, pointed extensions, which clasp the next following tooth about as far as the posterior edge of its crown. The superticial gano-dentine is smooth, but the base of the crown is impressed irregularly with a few large vertical plications or flutings, which are deepest in the anterior half and gradually disappear in the posterior extension. There is also a faint longi- tudinal median ridge on part of this extension. The anterior and posterior edges of the crown are coarsely serrated, the serrations (Pl. I, fig. 7) being about 30-in number on each edge, bluntly rounded (not crenulated), and those in the apical portion inclined upwards. In most of the teeth, however, the serrations are much worn by an apparently single row of opposing teeth. They are especially worn in the foremost tooth, irregularly on its anterior edge, most deeply in the Jower half of its posterior edge. The wear is nearly similar in the second, third, and fourth teeth; while in the fifth and sixth teeth a large worn hollow (w) is conspicuous in the upper half of the posterior edge. The apical portion of the seventh tooth is unfortunately lost, but the eighth tooth 1s complete and unworn, displaying all the serrations as previously described. The separate tooth (¢) behind is crushed at the base, and thus was probably not completely developed ; but the crown exhibits well its smooth face and unworn serrated edges. It may be either a ninth tooth of the series displayed or one of the opposing dentition, which is otherwise represented only by a broken fragment of one tooth in the edge of the shaly matrix of the fossil. The root of the fused symphysial teeth is well seen in front, where it has been extricated from the matrix, but it becomes vague behind where the teeth were in process of formation. In side view (Pl. I, fig. 1) the limits of the successive components are only just distinguishable; but in lower view (Pl. I, fig. 2) 1 OQ. P. Hay, Proc. U.S. Nat. Mus. vol. xlii (1912) p. 32. Ee eS eel cee ee part 1] A NEW SPECIES OF EDESTUS. 3 the divisions between the roots of the three anterior teeth are well marked by lines of calcite. In each tooth the root is much shorter than is usual in Hdestws, scarcely extending backwards beyond the hinder production of the base of the crown. As shown by the foremost tooth (figs. 2 & 3), the anterior margin of the root slopes downwards and backwards, and is compressed to an edge nearly as sharp as that of the crown. Its truncated lower face (fig. 2) is excavated into a triangular hollow, so that the lower face of the arch of clasping teeth is impressed by a wide longi- tudinal groove, of which the shape and depth are well seen in cross- section (fig.4). Sections prove that both root and crown are solid, consisting of the usual vascular dentine of rather open texture. Irregularly scattered over the shale below the arch of Hdestus are the more or less broken remains of comparatively small Orodont teeth (0), of the form commonly described as Campodus or Agassiz- odus. Some of these teeth are much extended laterally, witha low central cusp (Pl. I, fig. 8); while others appear to have had little lateral extension, but a relatively-large and elevated central cusp (figs. 9 &10). The longitudinal ridge on the summit of the crown and all the vertical buttresses on its outer and inner face are sharp and simply serrated; the superficial gano-dentine is otherwise smooth. ‘The serrated buttress of the central cusp is especially conspicuous on its outer face; but both this and the lateral but- tresses, which are widely spaced in from one to four pairs, are much more prominent on the outer than on the inner face. The com- pressed root is of very open texture, and seems to be inclined slightly inwards. The fossil itself affords no definite proof that the Orodont teeth thus described belong to the same jaw as the Mdestus, and their very small size seems at first to make their connexion improbable. Teeth of the same type, however, have already been found in association with Hdestus mirus1; and one jaw of Campodus has been described, in which a single arched series of high-crowned symphysial teeth is relatively enormous.? Moreover, it will be noticed that in the new fossil from Yorkshire there is no difficulty in regarding the teeth of the HMdestus type as an extreme modifi- cation of those of the Campodus type which oceur with them. In the new Hdestus the central cusp of Campodus has become exces- sively enlarged and laterally compressed, while the lateral extensions of the tooth are reduced and attenuated, and their sharp crest is represented merely by a faint longitudinal ridge. The irregular basal plications or flutings in the Hdestus are the last remnants of the anterior (outer) lateral buttresses in Campodus. 'The first approach, indeed, towards this extreme modification occurs in the symphysial teeth of the typical jaw of Campodus (Agassizodus) itself, as shown in the accompanying text-figure (p. 4). Here the 1 ©. P. Hay, Proc. U.S. Nat. Mus. vol. xlii (1912) p. 36. 2 ©. R. Hastman, ‘On the Nature of Hdestus & Related Forms’ Bull, Mus. Comp. Zool. Harvard Coll. vol. xxxix (1902) pp. 55-77 & pls. i-iv. B2 4, DR. A. SMITH WOODWARD ON [ vol. Ixxii, central cusp is much enlarged and elevated, and begins to be some- what laterally compressed, with sharp anterior and posterior (outer and inner) edges; but the lateral extensions of the tooth, although attenuated, still retain the characteristic anterior (outer) buttresses, the apical ridge throughout their length, and the typical root (8). In the new specimen of Hdestus the same enlargement of the central cusp with reduction of the lateral extensions has progressed so far that the central feature predominates immensely, while the diminutive lateral parts curve backwards along the modified root to clasp the next successional tooth (c). In typical Hdestus the tooth is almost entirely a laterally-compressed central cusp, while the clasping root is excessively enlarged (»). Diagrammatic sketches of half of a lateral tooth of Campodus variabilis, wpper view (A), and-symphysial teeth of Campodus variabilis (B), Edestus newton1 (c), avd Kdestus minor (»), left side view, to show progressive modification. A. B. Gc ID. [m = longitudinal ridge. } There seems thus to be no doubt that the small teeth of the Campodus type occurring with the new fossil really belong to it, and a question arises as to how the species represented by this jaw shall be named. Ca ampodus (1844) is an older generic term than Edestus (1856), but it is evident that the teeth to which its definition applies belong to more than one genus: for the sym- physial teeth of Cantos variabilis, as made ‘known by Eastman,! must be regarded as generically dienes from those of the specimen now described, ile the former must have been arranged as a single row only i in one jaw, and as a paired row in the opposing jaw; whereas in the Yorkshire teeth the surfaces of wear show that there must have been an unpaired median row in each jaw. As. the latter arrangement has already been proved by Hay to charac- terise Hdestus, “and as the symphysial teeth in the new fossil merely differ from those of the typical Hdestus in the relativ ely- small size and extension of the root, 1 propose to regard it as 1 Bull. Mus. Comp. Zool. Harvard Coll. vol. xxxix (1902) p. 58. Quart. Journ. Geo. Soc. Vor. LXXIl, Pt. I. Kqsaq'oj\09 ‘asouwag ‘Aaou ‘ds ‘INOLMA4N Siees = de {9p ‘PUeEMPOoN ‘WD part 1] A-NEW SPECIES OF EDESTUS. 5 representing an extremely generalized species of KHdestus. It ditfers from all known forms, not only in the shape and proportions ot the root, but also in the shape of the crown and the significant flutings and faint lateral ridge at its base. It represents, in fact, a very distinct species, which may be appropriately named Hdestus newtont, atter Mr. H. T. Newton, F.R.S., who first recorded the occurrence of Hdestus in British Carboniferous rocks.! EXPLANATION OF .PLATE I. Edestus newtoni, sp. nov.; anterior ends of cartilages of jaw (c), median symphysial dentition (s), a detached symphysial tooth (¢), and small seattered Orodont teeth (0).— Millstone Grit; Brockholes, near Huddersfield, Yorkshire. (Museum of Practical Geology, London.) [ Figs. 1-6 are two-thirds of the natural size, figs. 7-10 are twice the natural size. | Fig. 1. The whole fossil. w=worn surface on edge of tooth. 2. Anterior end of symphysial dentition, lower view. 3. Front view of the same. 4, Vertical transverse section of root of third tooth of the same. Figs. 5 & 6. Transverse sections of second and eighth teeth of the same. Fig. 7. Serratfons of the border of the eighth tooth. 8. Left half of an elongated Orodont tooth (Campodus), front view. 9. Elevated Orodont tooth (Campodus), front view; and (9a) upper view. 10. Elevated Orodont tooth (Campodus), back view. APppENDIX.—NoreE on the WeELL-SECTION at Rock Mints and the Fauna associated with Epresrus. By JoHN PRINGLE, E.G.S. THe well at Rock Mills (Messrs. Joseph Sykes & Co.) is situated in the mill-grounds in the valley of the River Holme at Brock- holes, about 4 miles south of Huddersfield. According to the Geological Survey-map (Quarter-Sheet 88 S.H.), the alluvium in the valley at this pomt rests on shales which underlie the Rough Rock, the uppermost member of the Millstone Grit. The shales appear to vary considerably in thickness in this district. Green ? states that 100 feet of shales are present between the Rough Rock and Grit A about Holmfirth, which lies 2 miles south of Brockholes; while on Holestone Moor, west of Huddersfield, he estimates that they are 200 feet thick between the same horizons. Spencer? has shown that in the district embraced in the next sheet to the north (Quarter-Sheet 88 N.E.) the shales contain a marine fauna; and the discovery of Hdestws should stimulate the efforts of the local geologists to further work on this horizon. 1H. T. Newton, ‘On the Occurrence of Edestus in the Coal-Measures of Britain’ Q. J. G.S. vol. lx (1904) pp. 1-8 & pl. i. 2 A. H. Green & others, ‘The Geology of the Yorkshire Coalfield’ Mem. Geol. Surv. 1878, p. 59. 3 J. Spencer, ‘ Additional Notes on the Millstone Grit of the Parish of Halifax’ Trans. Manchester Geol. Soc. vol. xiii (1873-74) p. 109. or) A NEW SPECIES OF EDESTUS. [vol. lxxii, During the excavation of the well no record of the various strata passed through was made by the sinkers; but it appears, from notes supplied by Mr. J. R. Simpson, of Bale Honley, that the rocks were mainly shales, with some thin beds of sandstone. At the depth of 120 feet the specimen of Hdestus was obtained by Mr. H. H. Freer in a bed of rock—rather friable shale,—which also contained numerous fragments of marine shells. Among the specimens the following have been identified :—Postdoniella levis (Brown), Gastrioceras sp., Glyphioceras reticulatum (Phill.), and Orthoceras cf. aciculare Brown. At the depth of 142 feet was a dark-grey flagey micaceous shale, crowded with shells which appear identical with JJodiola transversa Hind. ‘This shale formed the roof of a 83-inch coal which overlies a hard grey sand- stone. The coal-seam probably corresponds to the thin coal which lies on the top of Grit A in the Huddersfield district. Water was reached at the depth of 163 feet, and the sinking was stopped after being carried down a few feet lower. Our thanks are due to Mr. J. R. Simpson, who brought the discovery of the specimen of Edestus to our notice, and for his kindness in placing his specimens of the associated fauna at the disposal of the Geological Survey. The specimen was presented to the Museum of Practical Geology by Mr. E. Crowther, Managing Director of Messrs. Joseph Sykes & Co. Discussion Dr. A. SprauvAN expressed his appreciation of the skill with which the fossil had been developed at the Natural History Museum. He wished also to take the opportunity of acknow- ledging the obligation under which geologists had been laid by My. EH. Cramtinge in placing this unique specimen in a National Museum, where it will be studied by specialists from all parts of the world. Mr. J. Prinaie remarked that it was seldom that so interest- ing a specimen came before the Society, and he congratulated the Fellows on having had the opportunity of hearing Dr. Smith Wood- ward’s lucid and illuminating account of this Eee known genus. He referred briefly to the eeolocical position of the specimen and to the associated fauna. He had had the opportunity of studying exainples of marine shells obtained from the shales which yielded the Edestus. These comprised species of Glyphiocer as, Gastrio- ceras, Orthoceras, and Posidoniella. Mr. EK. T. Nrewron asked Dr. Smith Woodward whether he’ could explain how it was that, while in Hdestws the base of the tooth seemed to be extended backwards from the crown, in Helicoprion it was directed forwards. Dr. A. Smirn Woopwarp agreed with the last speaker that it was difficult to explain the difference in the direction of extension of the tooth in Hdestus as compared with Helicoprion. The two forms seemed to have had a distinct origin. part1] FOSSILIFEROUS LIMESTONE FROM THE NORTH SEA. 7 2. On a FosstiifeRous Limestone from the NortH Sea.t By Ricnarp Burien Newton, F.G.S. (Read June 23rd, 1915.) [Puate II.] THRouGH the good services of Mr. R. W. Thomson, of the Fishery Office, Aberdeen, the British Museum has been placed in possession of two blocks of limestone, one of considerable size, crowded with the remains of marine shells, which had been obtained from the bed of the North Sea by the steam-trawler, the ‘ Procyon,’ com- manded by Captain Wood, some 80 miles east of Orkney or 100 miles north-east $ north of Buchan Ness. In order to acquire ton ther information as to this unique occur- rence, | communicated with Dr. A. W. Gibb, the Curator of the Geological Museum of Marischal College, Aberdeen, who kindly sent me some interesting particulars on the subject. He was familiar with the rock, having had a sample forwarded to him by Dr. Bowman, the scientific investigator on the North Sea Fisheries’ boat, the ‘Goldseeker, who from considerable experience had gained some knowledge of the physical characters of the floor of the North Sea. Dr. Bowman informed Dr. Gibb ‘that at the spot where it [the limestone] occurs, there is what seems to be a gorge or submerged channel of some kind, with much deeper water than in the adjoining sea, and he thinks that there is a considerable mass of the rock in all probability [in situ ?],as trawlers report that they frequently break their gear upon it.’ Although found beneath the sea, this limestone nowhere displays any unusual abrasion, its aspect being entirely that of a rock which might have been obtained from an ordinary quarry or land exposure. There is nothing in its appearance to suggest transportation by glacial agencies, and it can only be/surmised that deep down in the North Sea a considerable development of the rock may actually occur 7 situ. So far as can be ascertained at present, no similar limestone is known in either England or Scotland. The so-called ‘Crag Strata’ of Aberdeenshire, described by Jamieson as occur- ring beneath the Boulder Clay and consisting of stratified sand and gravel, have yielded marine mollusca of a northern type, mixed with a few Red Crag species, which Searles Wood regarded as of Red Crag age ?; although, according to Clement Reds 3 such beds resemble the ‘Bridlington Crag’ deposits of Yorkshire, and would therefore be of Pleistocene horizon. Mr. Reid further stated that there is no Pliocene in Scotland, the Red Crag shells of the glacial beds of Aberdeenshire having been probably derived from strata of that age lying beneath the North Sea. A very 1 Communicated by permission of the Trustees of the British Museum. 2 Q.J.G.S. vol. xvi (1860) p. 373. 3 ¢The Pliocene Deposits of Britain’ Mem. Geol. Surv. 1890, p. 8 MR. R. B. NEWTON ON A FOSSILIFEROUS _ [vol. Ixxu, similar molluscan fauna to that occurring in the Scottish deposits has been found in the rocks of Iceland which Gwyn Jeffreys con- sidered as Post-Tertiary and Searles Wood as ‘not later than Middle Red Crag.’ ! Reference may also be made to Jamieson’s account of some semi-fossil shells of Arctic character, dredged by Robert Dawson from glacial beds underneath the sea, off the Aberdeenshire coast, from 3 to 8 miles away from land, at a depth varying from 30 to 45 fathoms. Similar shells had even been brought up by fishermen’s lines at a distance of 30 miles from the coast, and none of the species were thought to occur alive in that district.? Besides this, we have Alfred Tylor’s account? of fossil marine Arctic shells dredged off the Shetland Islands by Gwyn Jeffreys in about 90 fathoms of water—the species being stated to occur fossil in Sweden and still living in extreme Arctic seas. These records of fairly late fossiliferous deposits beneath the sea are of great interest, but scarcely assist us in understanding the history of the present rock. This North Sea limestone is of a dark-greyish colour, and is built up almost entirely of shell-remains belonging chiefly to the Pelecypoda. Microscopical tests have proved it to be highly siliceous, thus accounting for its great hardness and tenacity, which have frequently militated against complete development of some of the organisms, especially when the attempt was made to expose in- ternal @haracters: otherwise, speaking generally, the fossils are well preserved, and many of their finer external markings are perfectly displayed. It has been possible to determine 23 different forms of mollusea, ten being Gastropoda, while the remainder belong to the Pelecypoda, a new species being included in that group. If the material had been more dismantled, some additional forms might have been available for study, although in that case the larger block would have diminished in value as a museum specimen. The shells are essentially of southern character, although it is interesting to mention that one of the commonest forms is the large Cyprina islandica, which is so well known as existing im boreal seas and around our British coasts, but has never been recorded from the Mediterranean: its origin, however, is to be found in the Vindobonian stage of the Miocene Period. I have mentioned this species incidentally, because it frequently occurs in the Scottish and Icelandic deposits previously mentioned, which I consider of much later age than the present limestone. Occasional small brown masses of woody structure* occur in Q.J.G.S. vol. xli (1885) p. 96. Ibid. vol. xxi (1865) p. 200. Ibid. vol. xxv (1869) p. 8. The occurrence of drift-wood in the Coralline Crag near Sudbourn Church was mentioned by Clement Reid (- The Pliocene Deposits of Britain’ Mem. Geol. Surv. 1890, p. 28) ; but, so far as can be ascertained, no such material has been systematically described. em Ww Mw part 1] LIMESTONE FROM THE NORTH SEA. 9 contact with the limestone, but of very much softer character, which from a preliminary examination appear to be of a coniferous nature, although requiring further study before a more accurate deter- mination can be attempted. Out of the 23 determined molluses of this limestone 18 (or nearly 80 per cent.) trace their origin from the Vindobonian stage of the Miocene Period, 10 (or about 40 per cent.) may be regarded as extinct, whereas 12 (or 50 per cent.) are found living in recent seas. The extinct species include :— Streptochetus sexcostatus. Ficus | Pyrula] simplex. Ringiculella ventricosa. Yoldia oblongoides. Levicardim fragile. Sinodia tertiaria, sp. nov. Tellina benedent. Panopea menardi. Arcoperna sericea. Nucula levigata. The existing species are found chiefly in British and Mediter- yanean seas, although Spiswla ovalis and Cyprina islandica belong to more boreal waters, and are never found so far south as the Mediterranean. ‘This list embraces :— Nucula nucleus. Dentilucina borealis. Thyasira flexuosa. Cyprina islandica. TIsocardia humana (= cor). Spisula ovalis. Ranella gigantea. Turritella communis. Aporrhais pespelicant. Naticina alder. Actzxon tornatilis. < Scaphander lignarius. The majority of the species recognized are fairly evenly repre- sented in both the Coralline and the Red Crag deposits, 15 occurring in the former and about 12 in the latter. Among them the genus Yoldia, belonging at the present day entirely to Arctic seas, is well distributed through the limestone, specimens having been identified as Wood’s oblongoides, which is known from the Red Crag, Norwich Crag, and later deposits of East Angha, although never yet recorded from the Coralline Crag. Another species, however, of this genus, but not occurring in the present limestone, is Y. semistriata, which is restricted to Coralline Crag beds, and therefore forms an illustration of the Arctic genus Yoldia having been associated with a more southern fauna than prevailed during Red Crag or Jater geological times as represented in England. Further support is given to the southern aspect of the fauna by the occurrence in the limestone of a new Dosinitorm shell, Sznodza tertiaria, which presents affinities that are only to be found in the Indian Ocean and regions of Southern Asia. The comparatively large number of extinet species is greatly in favour of the North- Sea rock being of older horizon than the Red Crag, as among them are Arcoperna sericea, Tellina benedeni and Panopea menardi (= gentilis), which are not known of later age in this country than the Coralline Crag, although Zellina benedeni occurs as well in the Lenham Beds and Panopea menardi in the Boxstones. In con- nexion with the antiquity of this fauna, mention should also be made of the occurrence of Streptochetus sexcostatus and Ficus simplex, 10 MR. R. B. NEWION ON A FOSSILIFEROUS __ [vol. Ixxu, Streptochetus sexcostatus,a Fusiform shell, belongs to the Upper Miocene deposits of Northern Germany and Belgium (Anversian Beds), which are variously regarded as Messinian, Mio-Pliocene, — or the Sarmatian-Pontian Series ; in Holland the species has been recorded as of Vindobonian age. ‘his is its first acknowledgment as a British fossil, although remains of it are found in the Lenham Beds, a fact, however, published since the reading of this paper.! Ficus | Pyrula| simplex occurs only in the Messinian Beds of Northern Germany, never having been previously determined from British rocks. In addition to these species, Gottsche® has recorded from the same horizon and country, Scaphander lignarius, Arcoperna sericea, Levicardium fragile, and Dentilucina borealis, all of which form part of the fauna of the North-Sea rock. On the assumption, therefore, that the limestone belongs to some part of the Crag system, there appears to be ample evidence that it is of older age than the Red Crag, on account both of the total absence of Arctic species, and of the comparatively large number of extinct forms which have been recognized in it. It is more likely, therefore, to be of Coralline Crag age: for, in addition to the fact that the fauna agrees in the main with the molluscan fauna of that period, there are some lithological features which may be worthy of mention. According to the constitution of the Coralline Crag deposits of East Anglia, as explained by Prestwich,? the lower beds of the series contain ‘irregular seams of shelly mestone’; in more recent years Mr. F. W. Harmer has also referred to a similar formation in the same deposits as ‘tabular-layers of limestone, very hard, and difficult to penetrate,’ which were discovered in a boring at. Gedgrave Hall.“ I have not seen specimens of these limestones, although it would be desirable to compare them with the present rock. It does seem possible, however, that this North-Sea material may represent the lower portion of the Coralline Crag, and has, therefore, no connexion with Red Crag deposits, which are through- out of a sandy nature and furnish no evidence of intercalated calcareous beds. Considering the large number of extinct molluscan species fur- nished by the limestone, namely, 10 out of a total of 23 (or about 40 per cent. ), as against Searles Wood’s computation of the Coralline Crag mollusca made in 1874, that out of 391 species 142 were not now living® (thus representing about 36 per cent. of extinct forms), one may infer that, if the North-Sea rock really belongs to 1 R. B. Newton, ‘ On the Conchological Features of the Lenham Sandstones of Kent & their Stratigraphical Importance’ Journal of Conchology, vol. xv (1916) p. 74. 2 Verhandl. Ver. Naturwissensch. Unter. Hamburg, 1876—78, pp. 182-85. 3 Q. J.G.S. vol. xxvii (1871) pp. 123, 125. 4 Ibid. vol. liv (1898) p. 336. 5 See Clement Reid, ‘The Pliocene Deposits of Britain’ Mem. Geol. Surv. 1890, p. 38. part 1| LIMESTONE FROM THE NORTH SEA. eh the Coralline Crag, that formation is probably of earlier age than it is usually supposed to be at the present day; and it is interesting in this connexion to note that, in his earlier researches on the Crag mollusca, Searles Wood was of opinion that the Coralline Crag deposits represented the remains of the Miocene Period in Hngland—a view, however, which he afterwards relinquished ! in favour of regarding them as early Pliocene. Clement Reid placed them in his ‘Older Phlocene” group in association with the St. Erth Beds, the Lenham Beds, and the Boxstones,? whereas Mr. F. W. Harmer regards them as the base of the ‘ Newer Pliocene’ Series and younger than the Lenham and Boxstone Beds, which he restricts to the ‘Older Pliocene’ division as represented in this country.? It should be also stated that long before Reid and Harmer published their views on this subject, Mayer-Eymar had estab- lished the group-name ‘ Messinian’ for the later Miocene deposits of Europe, including in it the Lower Crag because a great number of its species were found in the Helvetian stage of the Miocene.* From my own studies of these mollusca, and especially those of the Lenham sandstones, I believe that no great disparity of age separates the so-called ‘ Older Plocene’ deposits. The faunas may differ somewhat in detail according to the special environments which governed their existence; but they all exhibit a strong southern character, and in other ways appear to resemble more the later Miocene life of Europe than that of the succeeding period. Judging entirely, therefore, from paleontological evidence, and apart from any physical considerations, I am inclined to regard this limestone as of Coralline Crag age. List of Molluscan Species. GASTROPODA. Family Lampusip 2. RaNELLA GrGanrEA Lamarck. (PI. I, fig. 1.) Ranella gigantea Lamarck, ‘ Hist. Nat. Anim. sans Vert.’ 1822, vol. vii, p. 160. Ranella reticularis Hoernes, Abhandl. K.IK. Geol. Reichsanst. 1853, vol. iii, p. 211 & pl. xxi, figs. 1-2. Remarks.——A small example, represented by a cavity in the limestone which, with the aid of a wax squeeze, yields the sculp- ture characters of this species. Distribution.—Vindobonian (Austria); Plaisancian (France and Italy) ; Recent (Mediterranean ). 1 Monogr. Pal. Soc. 1848, p. v (Introduction) ; ibid. 1857, pp. 301, 302. 2 «The Pliocene Deposits of Britain’ Mem. Geol. Surv. 1890, p. 2. 3 * Geology in the Field’ Jubilee Vol. Proc. Geol. Assoc. 1909, p. 90. 4 * Catal. Syst. Foss. Tert. Mus. Zurich’ pt. 2 (1867) p. 13. \ Ses eh LOSE RS *‘(Uesleg 03 1eqTRIGQLD) UBOUBIIOJIpPSP, puR YS 4 x *purlaoy —ULIUBIMOITpPa PT PUB FSI x SCH ESC “OGLE T pure uvauvitezyIpeyy ‘snttes “UVOUIE}IpPaT, PUB YSTLIG 4 See “UBAUBILOJ IPI x M/A) ere) uy) 2 Ale 0 in 99 PB ee | ee | es S| = Sh ai eal ‘SVEQ INDOaYy 8 4 Pi" : ie “SBI OULT[BIOD | Sped WH 4S | | | x é ae else al Se ce | = | a ao | oF lo} s z eae w| 2 @ ee “UBIySal(y | x tas > 2 8 Qs. i=] ss a= ism {oe) pe Bos mie ao Sse) = nai {9°} lex} ae iY) j=] *(UBLUOIIOT, =) UBIUOpery | x x x x x x =| oe Lei & Bele 0g a5] » of] & EB) = =, 2 | = a an (a2) (2, < (q>} ot Ste pe) =} | | ‘VAS HIMON AHL NOW ANOLSHWIT SHOUAMITISSO HY Ha —— NI GNQOd VOSOTIOP-, AHL FO MOVES GNV AW, NI NOWOAATELSt qd ‘uvlueqinbd y | *(Aqtemog *D ep *f) psooniquan pijanorbury (Soq10,7) 20a) U20290 AT *(snauury) iunorjadsad sipy.t.cody ‘ds w1oydouary OSSIY S2wmuU OD Y))A2RVMANT, (yortseg) wards snoyy *(qorsog) sngpysoavas srjayooj.dai4g yoinmury vagunhrh nppownay | “VadOdOULSV) —_—<—<—<—$<$___ ——_—__—_+ —_—__ ‘Seloodg pusw Brlouey “UGIUBLIOFTpPoa TAL jou arial aca [serpy pay Ur eatyvatieq | [Seip pay ur eargearreq | ‘vISW UIeyyNOG HO seas 0} poejoraqysea snes y UBOUBLIOFI pa] PUB YS WRIUBII} -Ipey| JoU—[vo10g pue YSyIAg “UBOTIOULY 910 NT pue “uvouvtnoqrpayy “YS “UBOTIOULY Y910 NT pue “Uvouvireqtpayy “YStyTg “Aup quesoad otf} 48 S¥OQ DIJOTV 0} SSUOTEG snuUer) ‘pueluesry Jou “UvIAvUIpUBOG pus ‘uvauRtojipeyy “Sty URAUBILOIpaT, pues YySsTyTIG “UBIUBLIOII pap, pus sty eenfelaie (Aq.tamog a) $7)900 pynsidgy eget sekvysa(] wpuwueu vadoung ‘diopuegse (A : wy ISA Wapauag vur7197, mers “Aou “ds ‘y2.1029.107 wrypoUng (109 =) (snenuIry) vuvwuny vypuvo0sT (sneuuIy) vorpwnjsr vuudhD * (1ypo01g) aprbouf wnrpimar0daT “ (nsevIMoy) vsonwayl v.vusvhy, 7, |" (sheuUry) s27nav0g vutonzyuaq “(P0OM “A “S) Saprohuojgo vipj0 x Aq19Mog “f vpnbr0H) DynonAT vogunt (sneuUUly) snaponw vynanjT Jrseeee (UtoIg) veorLes vusadoouy Vdqaod ANN TAd *(sneuury) smApuby sapunydvogy (SheUUIyT) s27270U.102 UWODI0F 14 MR. R. B. NEWTON ON A FOSSILIFEROUS _ [vol. Ixxu, Family Fustp#. STREPTOCHETUS SEXcostatTusS (Beyrich). (PI. II, figs. 2 & 3.) Fusus sexcostatus Beyrich, Zeitschr. Deutsch. Geol. Gesellsch. 1856, vol. viii, p. 73 & pl. xxiv, fig. 2 Streptochetus sexcostatus Cossmann, ‘ Essais de Paléoconchologie Comparée’ 1901, pt. iv, p. 31. Remarks.—This species, I believe, is represented in the English Crag deposits by Wood’s Fusus waeli,) a form that I am unable to recognize as the Belgian shell which was originally described by Nyst from the Oligocene (Rupelian) rocks of that country. A perfect comparison, however, is impossible, as Wood’s type- specimens do not appear to be in the British Museum. It is worthy of notice that Wood directed attention to an artistic error in his figure 106, which exhibited denticulations on the inner surface of the labrum, a character which was neither present in the English shell nor in S. sexcostatus from the German Miocene. Distribution.—Vindobonian (Holland) ; Messinian (North Germany); Anversian (Belgium) ; Lenham Beds (Britain). Family Frcrp 2. Ficts sIMpLEx (Beyrich). IL, figs. 4 & 5.) Pyrula simplex Beyrich, Zeitschr. Deutsch. Geol. Gesellsch. 1854, vol. vi, p. 777 & pl. xviii (xv), fig. 3 Ficula simplex Gottsche, a Ver. Naturwissensch. Unter. Hamburg, 1878, vol. 11, p. 182. Remarks.—Bolten’s Ficus of 1798 antedates both Lamarck’s Pyrula and Swainson’s Ficula, all of which are founded on the same Linnean tvpe of Bulla jficus. Distribution.—Messinian (North-Western Germany). Family TURRITELLID &. "TURRITELLA COMMUNIS Risso. Turritella communis Risso, ‘ Histoire Naturelle de Europe Méridionale’ 1826, vol. iv, p. 106 & pl. iv, fig. 37. Turritella terebra J. de C. ‘Sowerby, “Mineral Conchology’ 1827, vol. vi, p. 126 & pl. dlxv, fig. 3, now Linnzeus. Distribution.—Vindobonian (France); Coralline Crag to Post-Phocene (Britain) ; Recent (Mediterranean and British Seas). Family XENOPHORID#. XENOPHORA sp, (PI. IT, fig. 6.) Distribution.—This genus is unknown in the East Anglian Crag deposits, although occurring in the Lenham Beds of Kent. 1 Mon. Pal. Soc., Crag Mollusca, Suppl. 2 (1879) p. 9. & pl. i, figs. 10 a-10c. part 1] LIMESTONE FROM THE NORTH SEA. 15 It belongs more particularly to the Mio-Pliocene strata of Italy, Belgium, Holland, and Austria, besides surviving in recent seas (Mediterranean and Pacific). Family APORRHAID#. APORRHAIS PESPELICANT (Linnezeus). Strombus pespelicant Linneus, ‘Systema Nature’ 10th ed. 1758, p. 742. Distribution.—Vindobonian (Austria, Italy); Plaisancian and Astian (Italy, France); Anversian and Sealdisian (Bel- gium); Lenham Beds (Britain); Coralline Crag to Post-Plocene (Britain); Recent (Mediterranean and British Seas). Family Natterp a. NATICINA ALDERI (EH. Forbes). (Pl. II, figs. 7 & 8.)- 5 WNatica alderi HK. Forbes, ‘ Malacologia Monensis’ 1888, p. 31 & pl. ii, figs. 6-7. Natica (Naticina) alderi Dollfus, C. R. Assoc. Frane. Av. Sci. (Cherbourg, 1905) 1906, p. 368. Distribution.—Vindobonian (Holland) ; Messinian (Ger- many); Redonian (France); Coralline and Norwich Crags to Post-Pliocene ( Britain) ; Recent (Mediterranean and British Seas). Family Rine@rcuLip ». RINGICULELLA VENTRICOSA (J.deC.Sowerby). (PI.IL, figs. 9 & 10.) Auricula ventricosa J. de’ C. Sowerby, ‘Mineral Conchology’ 1824, vol. v, p. 99 & pl. ececlxy, fig. 1 Ringiculella auriculata var. ventricosa, Sacco, ‘ Moll. Terz. Piemonte’ 1892, pt. 12, p. 25; ibid. 1904, pt. 30, p. 110 & pl. xxiv, figs. 25-26. Remarks.—The shorter, broader, and more inflated volutions, as mentioned by 8. V. Wood, are sufficient to separate this species from Brocchi’s &. buccinea, which belongs to the Italian Tertiaries and existing seas. ‘The genus is well known in the Mediterranean. Distribution.——Vindobonian to Astian (Italy); Scaldisian (Belgium); Coralline Crag to Norwich Crag (Britain ). Family ACT HONID A. ACTON TORNATILIS (Linnzus). Bulla tornatilis Linneus, ‘Systema Nature’ 10th ed. 1758, p. 728. Acteéon striatus J. de C. Sowerby, ‘Mineral Conchology’ 1824, vol. v, p. 87 & pl. ccecelx, fig. 2. Acteon tornatilis S. V. Wood, Monogr. Pal. Soc. (Crag Mollusca) 1848, p. 170 & pl. xix, fig. 5 _ Distribution.— Vindobonian (Holland); Plaisancian and Astian (Italy); Anversian and Scaldisian (Belgium); Lenham Beds (Britain) ; Coralline Crag to Post-Pliocene (Britain) ; Recent (Mediterranean and British Seas). 16 MR. R. B. NEWTON ON A FOSSILIFEROUS [Vol. lxxii, Family SCAPHANDRID#. SCAPHANDER LIGNARIUS (Linneeus). Bulla lignaria Linneus, ‘Systema Nature,’ 10th ed. 1758, p. 727. Scaphander lignarius Montfort, * Conchyl. Syst.’ 1810, vol. 11, pp. 334-35. Distribution.—Vindobonian (Austria, Holland, Italy); Redo- nian (France); Plaisancian and Astian (Italy); Bolderian to Scaldisian (Belgium); Lenham Beds (Britain); Coralline and Red Crags (Britain) ; Recent (Mediterranean and British Seas). PELECYPODA. Family Myrrnip. ARCOPERNA SERICEA (Bronn). Modiola sericea Bronn, ‘TItaliens Tertiar-Gebilde’ 1831, p. 112; Philippi, ‘Enum. Moll. Sicilia’ 1836, vol. i, p. 71 & pl. v, fig. 14. Arcoperna sericea Sacco, ‘ Moll. Terz. Piemonte’ 1898, pt. 25, p. 43 & pl. xii, figs. 8-9. Distribution.—Vindobonian (Austria, Italy); Messinian (North Germany); Anversian to Sealdisian (Belgium) ; Coralline Crag (Britain); Astian (Italy). Family NucunipZ. Nvctura nucievs (Linneus). Arca nucleus Linneus, ‘Systema Nature’ 10th ed. 1758, p. 695; ibid. (Gmelin) 1790, 13th ed. vol. i, pt. 6, p. 8314. ~ Distribution.—Vindobonian (Austria); Redonian (France) ; Anversian to Scaldisian (Belgium); Coralline Crag to Post- Phocene (Britain) ; Recent (Mediterranean and British Seas, ete. ). Noucuta tavieata J. Sowerby. (PI. II, fig. 11.) Nucula levigata J. Sowerby, ‘Mineral Conchology’ vol. ii, 1818, p. 207 & pl. excii, figs. 1-2. Remarks.—This large and tumid Nuculoid shell is of frequent occurrence in the idnestiorae. and in general features resembles Nucula cobboldie of J. Sowerby, bub is without the divaricate sculpture of that species. Distribution.—Vindobonian (Holland); Redonian (North- Western France); Anversian to Scaldisian (Belgium) ; Coralline and Red Crags (Britain) ; Astian (Holland). Family NucunanNipH. YoLpIA OBLONGOIDES (S. V. Wood). (PI. II, fig. 12.) Nucula oblongoides 8. V. Wood, Mag. Nat. Hist. (Charlesworth), 1840, n. s. vol. iv, p. 297 & pl. xiv, fig. 4. Leda myalis 8. V. Wood, Monogr. Pal. Soc. (Crag Mollusca) 1851, p. 90 & pl. x, fig. 17, non Couthouy. part 1] LIMESTONE FROM THE NORTH SEA. Le Remarks.—This species is of fairly abundant occurrence, and shows relationship both to Y. hyperborea Torell and tu Y. limatula Say, of modern seas (boreal), from which it chiefly differs in the possession of thicker and more convex valves. Distribution.—Red Crag to Post-Plocene (Britain). Family Locryip a. DENTILUCINA BOREALIS (Linneus). Venus borealis Linneus, ‘Systema Nature’ 12th ed. 1767, vol.i, pt.2, p. 1134. Dentilucina borealis Sacco, ‘Moll. Terz. Piemonte’ 1901, pt. 29, p. 80 & pl. xviul, figs. 23-26. Distribution.—Vindobonian (Holland, Italy, Austria); Re- donian (North-Western France); Messinian (North Germany) ; Anversian, Diestian, and Scaldisian (Belgium); Plaisancian and Astian (Italy, France) ; St. Erth Beds to Post-Pliocene (Britain) ; Recent (Mediterranean and British Seas, etc.). Family THyastRipZ. THYASIRA FLEXUOSA (Montagu). Venus sinuosa Donovan, ‘Nat. Hist. British Shells’ 1801, pl. xli, fig. 2, non Pennant, 1777. Tellina flexuosa Montagu, * Test. Britannica’ 1803, pt. 1, p. 72. Thyasira flecuosa (Leach MS.), Lamarck, ‘ Histoire Naturelle des Animaux sans Vertébres’ 1818, vol. v, p. 492. Distribution.—Redonian (North-Western France); Anversian to Scaldisian (Belgium); Coralline Crag, Chillesford Beds, and Post-Pliocene (Britain); Recent (Mediterranean and British Seas, etc.). Family Carpiip2. L&VICARDIUM FRAGILE (Brocchi). Cardium fragile Brocchi, ‘Conchiologia Fossile Subapennina’ 1814, vol. iu, p. 505 & pl. xiii, fig. 4. Levicardium norvegicum Sacco, ‘ Moll. Terz. Piemonte’ 1899, pt. 27, p. 51 & pl. xi, figs. 41-42. Distribution.—Vindobonian (Austria); Messinian (North- Western Germany); Plaisancian and Astian (Italy). Family Cyprinip#. CyprINA ISLANDICA (Linneus em. Miiller). Venus islandica Linneus, ‘Systema Nature ’ 1767, 12th ed. vol.i, pt. 2, p. 1131 (without reference to a type form); Miiller, ‘ Zool. Danice Prod.’ 1776, p. 246 (no. 2977). Cyprina islandica Nyst, ‘Coquilles & Polypiers Foss. Tert. Belg.’ Mém. Cour. Acad. Roy. Belg. 1843-43, vol. xvii, p. 146, pl. ix, fig. 1 & pl. xi, fig. 1. Q.J.G.S. No. 285. C 18 MR. R. B. NEWTON ON A FOSSILIFEROUS _ [vol. lxxu, Remarks.—The valves of this species are of abundant occur- rence, while some of the adult examples measure from 80 to 85 millimetres in length and height. Distribution.—Vindobonian (Holland); Anversian to Scal- disian (Belgium); Plaisancian and Astian (Italy); Boxstones, Lenham Beds, and Coralline Crag to Post-Pliocene (Britain) ; Recent (British and Boreal seas, non Mediterranean ). IsocaARDIA HUMANA (Linneus). (PI. II, fig. 13.) Cardium humanum Linneus, ‘Systema Nature’ 1758,-10th ed. p. 682. Chama cor Linneus, ‘Systema Nature ’ 1767, 12th ed. vol. i, pt. 2, p. 1187. Tsocardia cor Lamarck, ‘ Hist. Nat. Anim. sans Vert.’ 1819, vol. vi, pt.1, p. 31. Tsocardia humana Dall, Trans. Wagner Free Inst. Sci. Philadelphia (Tertiary Fauna of Florida), 1900, vol. iii, pt. 5, p. 1064, Distribution.—Tortonian (Italy, Holland); Plaisancian and Astian (Italy); Diestian and Scaldisian (Belgium) ; ; Boxstones, Lenham Beds, Coralline Crag, and derivative in Red Crag (Britain); Recent (Mediterranean and British Seas). Family Dosrn1ip 2. SINODIA TERTIARTA, sp. nov. (PI. I, figs. 14-16.) Deseription (Right valve).—Shell solid, ineequilateral, sub- orbicular, height in excess of length, well arched ; posterior region deep, sloping, marginal curvature extensive; dorso-anterior margin oblique, straight, long; ventral margin oradually ascending to unite with the expanded and rounded edge of the lower anterior side ; lunuloid region superficial, large, ovately- elongate, obscurely circumscribed ; hinge area massive, deep, bearing two divergent, thick, robust, cardinal teeth, also a third and smaller cardinal in front, of more or less laminate appearance ; ligamental furrow wide, long, and prominent; pallial sinus triangulate, moderately wide at the base, apex obtuse; internal margin smooth ; surface covered with well-marked, nearly equidistant, elevated growth-lines and numerous finer concentric strize occupying the intervening spaces, the whole crossed by obscure radial striations. Dimensions in millimetres.—Largest example, right valve. Length = 50; height = 53; diameter = 16. Remarks.—The more striking features of this shell include the anterior obliquity, the long and superficial lunuloid area, the nearly equidistant growth-lines, and the extensive curvature of the pos- terior margin. It appears to be allied to Chemnitz’s Venus excisa rather than to the other forms referred to this genus, which are of more distinctly trigonal contour. The shell is fairly common in the North-Sea limestone, although it is difficult to obtain specimens with perfect internal characters; one example, however, does exhibit a partial internal cast of a left valve, showing the presence of a well-marked, obtusely-pointed, and triangulate pallial sinus. It is proposed to name this shell Scnodia tertiaria. part 1] LIMESTONE FROM THE NORTH SEA. — 19 It is interesting to find among these fossils a new Dosiniform shell that can be referred to Jukes-Browne’s S’nodia,! which was established on the type of Dosinia trigona of Reeve, and also includes the Venus eaxcisa of Chemnitz. Sinodia is chiefly distinguishable from the true Dosinia by reason of its oblique and straight dorso-anterior margin, followed by an expanded anterior side. Jt is known only in the living state, being restricted to the waters of the Indian Ocean, particularly the coasts of Southern Asia. Family 'TELLINID ®. TELLINA BENEDENT Nyst & Westendorp. (PI. II, fig. 17.) Tellina zonaria Nyst, “Recherch. Cog. Foss. Anvers’ 1835, p. 4, now Basterot. Tellina benedeni Nyst & Westendorp, Bull. Acad. R. Sci. Bruxelles, 1839, vol. vi, pt. 2, pl. 11, fig. 5 bis & pl. iii, fig. 5, p. 399. =. fallax Beyrich, a manuscript name. Remarks.—This is by far the most abundant molluse in the limestone, and well agrees with the typical form from the Belgian Upper Tertiaries. Examples are chiefly preserved as natural casts which exhibit well-marked muscular and other impressions, as also the extensive pallial sinus with its elevated and obtuse summit. Distribution.—Anversian to Scaldisian (Belgium); Middle Pliocene (Holland); Lenham Beds and Red Crag—derivative (Britain). Family Saxicavip 2. PaNoPHA MENARDI Deshayes. Panopea faujasii Basterot, Mém. Soc. Hist. Nat. Paris, 1825, vol. ii, pt. 1, p. 95, non Ménard de la Groye. Panopea menardii Deshayes, ‘ Dict. Class. Hist. Nat.’ 1828, vol. xiii, p. 22. Panopea gentilis J. de C. Sowerby, ‘Mineral Conchology’ 1840, vol. vii, p- 1 & pl. dex, fig. 1. Distribution.—Vindobonian (France, Poland, Austria, Switzer- land, Italy, Holland) ; Anversian and Scaldisian (Belgium) ; Boxstones, Coralline Crag, and Red Crag—probably derivative ( Britain). Family Macrrip#. _ SPISULA OVALIS (J. Sowerby). Mactra ovalis & dubia J. Sowerby, ‘ Mineral Conchology * 1817, vol. 11, p. 136 & pl. clx, figs. 2-5. Distribution.— Vindobonian (Switzerland); Redonian (North- Western France); Middle Phocene (Holland); Scaldisian (Bel- gium); Red Crag to Post-Phocene (Britain) ; Recent (British Seas, non Mediterranean ). 1 Proe. Malacol. Soc. London, vol x (1912) pp. 100-104. c 2 20 MR. R. B. NEWTON ON A FOSSILIFEROUS [vol. lxxui, PostscRIPt. [Since the reading of this paper two memoirs have been published on some Miocene limestones from the North-Sea basin (Denmark and Germany) containing mollusean remains, several species of which are identical with those found in the North-Sea Limestone. In a Middle Miocene boulder from Esbjerg (Denmark), EH. M. Norregaard! has reported the occurrence, among other shell- remains, of Arcoperna sericea, Tellina benedeni, Naticina alderi, Streptochetus sexcostatus, and Acte@on tornatilis; while Karl Gripp 2 enumerates the same species as occurring in some older Miocene deposits of Denmark and Germany, with the following additional forms :—TIsocardia humana (=cor), Thyasira flexuosa, Dentilucina borealis, Levicardium fragile, Cyprina cf. islandica, Tellina benedeni (=fallax), Panopea menardi, Ficus simplex, and Ringiculella ventricosa, all of which are found in the North- Sea Limestone. Again, attention may be directed to a recently published account of the Lenham Sandstone Mollusca,? which have been referred to a late Miocene age, the following species contained therein indicating affinities with the fauna of the North-Sea Lime- stone :—Aporrhais pespelicani, Streptochetus sexcostatus, Acteon tornatilis, Scaphander lignarius, Yoldia oblongoides, Isocardia humana, Dentilucina borealis, Tellina benedeni, and Panopea menardt, R. B. N. March 27th, 1917. } EXPLANATION OF PLATE II. [All the figures are of the natural size, except where otherwise stated. | Fig. 1. Ranella gigantea Lamarck. (See p. 11.) Prepared from a wax model taken from a limestone cavity. . Streptochetus sexcostatus (Beyrich). (See p. 14.) . The same, showing magnified sculpture. . Ficus simplex (Beyrich). (See p. 14.) . The same, showing details of sculpture, magnified. . Xenophora sp. (See p. 14.) Fragmentary specimen exhibitirg adherent cavities on the whorl. . Naticina alderi (KH. Forbes). (See p. 15.) . The same. . Ringiculella ventricosa (J. de C. Sowerby). (See p. 15.) Front. view, X 3. 10. The same, showing sculpture lines, magnified. 11. Nucula levigata J. Sowerby. (See p. 16.) 12. Yoldia oblongoides (S. V. Wood). (See p. 16.) 13. Isocardia humana (Linneus). (See p. 18.) BD oe 0 LO 65 OO ~T 1 « Mellem-Mioczene Blokke fra Esbjerg’ Danmarks Geol. Underség. ser. 4, vol. i, No. 5 (1916) pls. i-iii, pp. 58. , 2 See, for example, J. Joly, Phil. Mag. vol. xxiv (1912) p. 694; and A. Holmes, Sci. Progress, no. 33 (1914) p. 15. 2 Foid. p. 17. 3 Geol. Mag. ser. 6, vol. i (1915) pp. 68-69. Silbid. py lulls > Ibid. p. 67. part 3] VOLCANIC ROCKS OF MOZAMBIQUE. 27 there were a considerable relief of the pressure appropriate to such depths. In this connexion, the boundary-fault along which the lavas are aligned is of particular interest, for it seems to mark a zone along which pressure actually was relieved to an extent and depth sufficient to promote fusion. The boundary-fault system is probably a consequence of isostatic readjustment between the uplifted mainland and the sunken region of Mozambique Channel. Long denudation of the crystal- line plateau should, on the hypothesis of isostasy, naturally lead to its uplift, and that such a movement has taken place is demon- strated by the presence of a coastal belt of Cretaceous and Tertiary sediments, and by the existence of still more recent raised beaches at various levels along the coast (see fig. 5, p. 280). More- over, the heavy denudation of the crystalline rocks implies the removal of an important section of the ‘radioactive layer.’ This process therefore leads, like uplift, to the cooling and contraction of the lithosphere, and thus additional tension-stresses are developed that can only find relief in fissures and faults. Doubtless the actual formation of fissures was helped by regional movements of « tensional character, associated in a complementary way with the compressive movements of Tertiary mountain-building elsewhere. In conclusion, I wish to place on record the unfailing courtesy and hospitahty with which we were everywhere received by the _ Portuguese officials in Mozambique, who were always ready to help us in every way that lay in their power. For permission to publish the geological observations made during the expedition, I owe my thanks to the Directors of the Memba Minerals Ltd. ; in particular, I have to thank them for placing at my disposal a large collection of rock-specimens, including those collected in the Sanhuti-River district, and many of those collected by the staff of the Company during the 1910 season. From my friends and former colleagues Messrs. E. J. Starey, E. J. Wayland, and D. A. Wray, I have received notes and sketches referring to the districts in which they worked, and I desire to express my indebtedness to them for help thus afforded. In connexion with the investigation of the rocks in the laboratory, I wish to thank Prof. W. W. Watts for granting every possible facility for work at the Imperial College of Science & Technology; Prof. the Hon. R. J. Strutt for continuing to lend me his apparatus for determining radium in rocks and minerals; Prof. C. G. Cullis and Dr. J. W. Evans for examining many of my sections, and offering valuable suggestions ; and Dr. G. T. Prior and Lieut. W. Campbell-Smith for looking through my sections, and allowing me to compare the Mozambique lavas with specimens of similar rocks in the Natural History Museum from British East Africa, Abyssinia, Nyasaland, and other African localities. In making the analyses published in this aper I received considerable help from Dr. H. F. Harwood, Lieut. J. L. Harris, M.C., the late Lieut. G. Kirby, Lieut. A. McIntyre, and Mr. HE. Spencer, and for their assistance in an Q. J. G.8. No. 287. x 278 DR. A. HOLMES ON THE TERTIARY [ vol. Ixxii, arduous task I am particularly grateful. Finally, I owe my thanks to Mr. G. 8. Sweeting, who kindly cooperated with me in the preparation of the photomicrographs reproduced in Plates XX & XXI. EXPLANATION OF PLATES XX & XXI. PLATE XX. Fig. 1. Sélvsbergite (153) found between Miali and the Sanhuti River, Mozam- bique. Chief minerals: anorthoclase, cossyrite, and soda-pyroxenes. x 20. (See p. 233.) 2, Aigirine-trachyte (152) north of Sokoto Hill. Chief minerals : anortho- clase, cossyrite, and egirine. X 20. (See p. 235.) 3. Phonolite (150) near the Saahuti River. Chief minerals: anorthoclase, nepheline, natrolite, soda-pyroxenes, and amphiboles. The dark phenocryst is cossyrite surrounded by a green chlorite-like mineral. x 20. (See p. 236.) - 4. Basalt with ophitic patches (156) near the Sanhuti River. Chief minerals: labradorite and augite, with some palagonite in the ground- mass. xX 20. (See p. 241.) 5. Picrite-basalt (155) near the Sanhuti Ree. Chief minerals: purple augite and olivine; labradorite, purple augite, and giass in the ground-mass. xX 20. (See p. 244.) . Micro-vesicular basalt (128) from a sill at the western base of Sokoto Hill. Chief minerals: labradorite and augite, in a base of dark-brown glass. The vesicles are occupied by Chalcedony and chlorite. x 20. er) PuatTEe XXI. Fig. 1. Basalt (141) from the dyke at Mochelia. Chief minerals: labra- dorite and augite in a base of dark-brown glass. X36. (See p. 246.) . Amyedaloidal sandstone (112) from contact with the Mochelia dyke (text-fie. 5). Grains of quartz, orthoclase, and microcline are present in the sandstone, cemented by calcite which is replaced near the dyke hy opal. Inthe steam-holes opal, chalcedony,and quartz are present. x 12. (See p. 249.) : . Amygdaloidal basalt (124) south of the Monapo River. Chief minerals : labradorite and augite in a base of dark-brown glass. x 20. (See p. 250.) 4. Amygdale in amygdaloidal basalt (136) east of Murimatigri Quartel. Tilustrating the replacement of heulandite (dark areas) by chal- cedony. xX 20. (See p. 252.) 5. Hornblende-andesite (126) from a dyke penetrating amygdaloid south of the Monapo River. Chief minerals: andesine, hornblende, and biotite, in a ground-mass consisting of the same minerals, together with quartz and orthoclase. x 20... (See p. 256.) 6. Pyroxene-andesite (144) found near the Monapo River. Chief minerals : andesite, augite, and hypersthene, in a ground-mass of hyalopilitie texture. x 20. (See p. 259.) bo ee) _ Discussion. My. F. P. Mernnetr said that he had listened to the paper with the greater interest, in that he had spent more than ten years ip a neighbouring part of Africa. He was quite in accord with [November 23rd, 1917. | Quart. Journ. Geor. Soc. Vor. LXXII, Pr. XX. ik ae ie or at ARAL 7 AH. & 6.S.S,, Photomicro. Bemrose, Collo., Derby VOLCANIC ROCKS oF THe DISTRICT o— MOZAMBIQUE. Quart. Journ. Geot. Soc. Vor. LXXIl, Pi. XXI. AH. & G.S.S.,Photomicre. Bemrose, Collo., Derby VOLCANIC ROCKS oF tHe DISTRICT oF MOZAMBIQUE. ail if Jans part 3] VOLCANIC ROCKS OF MOZAMBIQUE. 279 the Author, that alkaline and non-alkaline rocks were to be found along the African coast in intimate association. It was not clear, however, from what had been said, on what principle the Author separated the rocks that he put into his alkaline group from the others with which he had dealt, and it was not easy to see why they should be treated apart. It was interesting to note that. ‘although alkaline voleanic rocks were so often Sonndicn the coastal regions, there were no occurrences known in the inland districts of this part of Africa. Dr. J. W. Evans congratulated the Author on his presentation of the results of several years’ work on some very interesting rocks. He had not confined himself to simple descriptions of structures and minerals, but had endeavoured to ascertain what light the facts disclosed threw on the evolution of igneous rocks. especially those rich in alkalies. The speaker enquired whether there was any evidence that the alkali-rocks were later than those of normal types in this area. Mr. A. E. Kirsoy congratulated the Author on the paper. He asked whether there was any general linear arrangement of the dykes, and if so, whether the ‘lines were parallel w with the coast. He said that on both sides of Africa, wherever late volcanic rocks occurred, it seemed that they had been extruded along lines of fracture roughly parallel with the coast, or in Peon -zones inland, and that on the Gold Coast volcanic necks and plugs occurred in several places along the Volta River; but he had not found similar evidence in that colony outside of zones of fracture. The AvTHoR, in reply, thanked the Fellows for their kind reception of his paper, and stated that most of the points raised in the discussion were Exily dealt with in the paper itself. Answering Mr. Mennell~he said that the division of the lavas into—40_ series _we< based on analyses, varlation-diagrams, the composition of the amygdale-minerals, and in part on the sequence. so far as this was known. In reply to Dr. Evans, he was sorry to say that there was no definite field-evidence from which the relations of the two series could be deduced. Referring to Mr. Kitson’s remarks, he observed that, as in many other of the African coast- lands, the main faults were parallel to the prevailing trend of the coast, and most of the basaltic dykes followed the same direction. ADMISSION AND PRIVILEGES OF FELLOWS OF THE GEOLOGICAL SOCIETY OF LONDON. Every Candidate for admission as a Fellow must be proposed by three or more Fellows, who must sign a Certificate in hisfavour. 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Bonney, F.R.S., Sc.D. Crown 4to. lds. net. Contains the results of further visits to the Lipari Islands, Vesuvius (after the outburst in 1906), and to Etna, together with photographs taken after the great eruptions of the Soufriére in St. Vincent and Mont Pelé in Martinique, which occurred in 1902. Dr. Anderson again visited these islands in 1908 and obtained illustrations to show the return of vegetation and other changes. The present volume gives photographs of some Mexican volcanoes, taken in 1908, and of others from Guatemala, New Zealand, Matavanu, in the Samoan Islands, and Kilauea. It concludes with a selection from the photographs obtained by Dr. Anderson on his last journey—that to Java, Krakatau, and the Philippine Islands—so as to form with its predecessor an exceptionally large collection of Volcanic Studies from all parts of the globe. JOHN MURRAY, Albemarle Street, LONDON, W. 1. CONTENTS. PAPERS READ. Page 9. Dr. W. R. Jones on the Secondary Stanniferous Deposits of the Kinta District (Plates ADL OKV) 522 a caus cee areas eee eae so arnch s eteewis seu ataee te 165 10. Prof. S. J. Shand on the Pseudotachylyte of Parijs, Orange Free State (Plates 2X ViTSXDRe) ences oe Ee eedocle eh ier tare ac eae gates. ete ieee meee 198 11. Dr. A. Holmes on the Tertiary Volcanic Rocks of the District of Mozam- bigue (Plates Kexe & XN) I Sere nie se esac oupty aaa eer sie ee ee ae OemeecirmEs 222 [The Editor of the Quarterly Journal is directed to make it known to the Public that the Authors alone are responsible for the facts and opinions contained in their respective Papers.| *,.* The Council request that all communications intended for publication by the Society shall be clearly and legibly written on one side of the paper only, with proper references, and in all respects in fit condition for being at once placed in the Printer’s hands. Unless this is done, it will be in the discretion of the Officers to return the communication to the Author for revision. The Library at the Apartments of the Society is open every Week Day from Ten o'clock until Five, except on Saturdays (when the hours are from 10 to 1), and except during the fortnight commencing on the first Monday in September, when the Library is closed for the purpose of cleaning. Vol. LXXII. No. 288. Parr 4. THE QUARTERLY JOURNAL OF THE GEOLOGICAL SOCIETY. EDITED BY THE PERMANENT SECRETARY. [With Three Plates, illustrating Dr. Stanley Smith’s Paper. | DrcEMBER 31st, 1917. LONDON : LONGMANS, GREEN, AND CO. PARIS :—CHARLES KLINCKSIECK, 11 RUE DE LILLE. SOLD AL8O AT THE APARTMENTS OF THE SOCIETY. Price Five Shillings. RRR RII I IPD IRIN GLI LL IIS II IEEE DO OOO PRADEEP ODE LIST OF THE OFFICERS AND COUNCIL OF THE GEOLOGICAL SOCIETY OF LONDON. Blected February 16th, Ue President. Alfred Harker, M.A., LL.D., F.R.S. Gice-Prestvents. R. Mountford Deeley, M.Inst.C.H. Edwin Tulley Newton, F.R.S. William Johnson Sollas, M.A., LL.D., Se.D., F.R.S. Arthur Smith Woodward, LL.D., F.R.S., F.L.S. Secretaries. Herbert Henry Thomas, M.A., Sc.D. Foreign Secretary. | Herbert Lapworth, D.Sc., M.Inst.C.E. Creasurer. Sir Archibald Geikie, O.M., K.C.B., D.C.L., | James Vincent Hlsden, D.Sc. LL.D., Se.D., F.R.S. COUNCIL. Charles William Andrews, D.Sc., F.R.S. Prof. John Cadman, C.M.G.,D.Sc., M.Inst. C.E. Prof. Charles Gilbert Cullis, D.Sc. Arthur Morley Davies, D.Sc., A.R.C.Sc. R. Mountford Deeley, M.Inst.C.H. James Vincent Elsden, D.Sc. Prof. Edmund Johnston Garwood, M.A., Se.D., F.R.S. Sir Archibald Geikie, O.M., K.C.B., D.C.L., LL.D., Se.D., F.R.S. Walcot Gibson, D.Sc. Alfred Harker, M.A., LL.D., F.B.S. Finlay Lorimer Kitchin, M.A., Ph.D. George William Tamplugh, F.R.S. Herbert Lapworth, D.Sc., M.Inat.C.E. Prof. John Edward Marr, M.A., Se.D., E.R.S. Edwin Tulley Newton, F.E.S. Richard Dixon Oldham, F.R.S. Robert Heron Rastall, M.A. Prof. Thomas Franklin Sibly, D.Sc. Prof. William Johnson Sollas, M.A.,UL.D., Se.D., F.R.S. Sir Jethro J. Harris Teall, M.A., D.Sc., LL.D., F.R.S. Herbert Henry Thomas, M.A., Se.D, Samuel Hazzledine Warren. Arthur Smith Woodward, LL.D., F.R.S., E.L.S. Permanent Secretary. L. L. Belinfante, M.Sc. Librarian. C. P. Chatwin. Clerk. M. St. John Hope. Assistant tn Library. Arthur Greig. STANDING PUBLICATION COMMITTEE. Dr. A. Harker, President. Dr. H. Lapworth. Dr. Herbert H. ieee Secretaries. Dr.\F. A. Bather Prof. J. Cadman. Prof. C. G. Cullis. Dr. J. V_ Hisden. Dz. W. Gibson. Dr. F. L. Kitchin. Mr. G. W. Lamplugh. Mr. HE. T. Newton. Mr. R. D. Oldham. Prof. T. F. Sibly. Prof. W. J. Sollas. Sir Jethro Teall. Dr. A. Smith Woodward ORDINARY MEETINGS OF THE GEOLOGICAL SOCIETY TO BE HELD AT BURLINGTON HOUSE. Szssron 1917-1918. 1918. Wednesday, January .. 9*—23* ss February (Anniversary, Friday, February 15th) . wee eee eee see neneee Se a 6*—20* [Business will commence at 5.30 p.m. precisely. | The asterisks denote the dates on which the Counoil will meet. part 3] VOLCANIC ROCKS OF MOZAMBIQUE. 279 the Author, that alkaline and non-alkaline rocks were to be found along the African coast in intimate association. It was not clear, however, from what had been said, on what principle the Author separated the rocks that he put into his alkaline group from the others with which he had dealt, and it was not easy to see why they should he treated apart. It was interesting to note that, although alkaline voleanic rocks were so often found in the coastal regions, there were no occurrences known in the inland districts of this part of Africa. Dr. J. W. Evans congratulated the Author on his presentation of the results of several years’ work on some very interesting rocks. He had not confined himself to simple descriptions of structures and minerals, but had endeavoured to ascertain what light the facts disclosed threw on the evolution of igneous rocks, especially those rich in alkalies. The speaker enquired whether there was any evidence that the alkali-rocks were later than those of normal types in this area. : Mr. A. E. Krrson congratulated the Author on the paper. He asked whether there was any general linear arrangement of the dykes, and if so, whether the lines were parallel with the coast. He said that on both sides of Africa, wherever late voleanic rocks occurred, it seemed that they had been extruded along lines of fracture roughly parallel with the coast, or in fracture-zones inland, and that on the Gold Coast voleanic necks and plugs occurred in several places along the Volta River; but he had not found similar evidence in that colony outside of zones of fracture. The AvurHor, in reply, thanked the Fellows for their kind reception of his paper, and stated that most of the points raised in the discussion were fully dealt with in the paper itself. Answering Mr. Mennell, he said that the division of the lavas into two series was based on analyses, variation-diagrams, the composition of the amygdale-minerals, and in part on the sequence, so far as this was known. In reply to Dr. Evans, he was sorry to say that there was no definite tield-evidence from which the relations of the two series could be deduced. Referring to Mr. Kitson’s remarks, he observed that, as in many other of the African coast- lands, the main faults were parallel to the prevailing trend of the coast, and most of the basaltic dykes followed the same direction. Q. J. G.S. No. 288, ———— 280 DR. STANLEY SMITH ON AULINA ROTIFORMIS, (vol. xxii, ~ 12. Avrivna ROTIFORMIS, gen. et sp. nov., PHILLIPSASTR.EA HENNAHT (Lonsdale), and OrronasTR#£.4, gen. nov. By SrantEy Suira, B.A., D.Sc., F.G.S. (Read November 8th, 1916.) [Puatres XXII-XXIV.] 7 CONTENTS. Page dBRL bai ndoleuneinlorabymandesnhocwaucsbcooatepetooceddiccanost aabane oncanb sd yoDe so a5. 280 Il. Definition of certain Terms employed ...... ........:::.:ecee eden 281 III. The Astreeiform Corallum ........ ee Sete A ohh ge cane ae 281 IN EI ACUUC SECRETE cane aun catnabaosphbdnnaboncanesscaracunenwevescagas0tpoan. 284 Ai, HISTO Is e313 Oi “soonSesanaco sdesnd =JoSec ono nnbar> sods cedisbancuococbasue 290 VI. Orionastred, GON. NOV. .....6c.csceccscee cece etc e ete eet ene tee neneet ees 294 VII. Notes on Cyathophyllum regium and on Koninckophyllum sp. 304 “ I. IyrRopvucTIoN. PHILLIPSASTRZA is a genus of Devonian Corals possessing - certain well-marked characters. The genus Ovionastrea has been established to include certain Carboniferous species very closely related to Lithostrotion, but which have been regarded by many writers as congeneric with those of Phillipsastrea. O. phillipst (McCoy) is commonly known as Ph. radiata (Martin). Aulina is a new genus, found at the horizon of the Millstone (Grit. One species only has been recognized: this has been confused hitherto with O. phillipsi, and hence recorded as Ph. radiata. I vegard Phillipsastrea as the ancestor of Aulina, but do not consider that these are related to Orionastrea. All three genera, however, are colonial in habit, and possess a similar type of corallum: namely, that in which the individual corallites have lost their epitheca, and consequently are united by their dissepiments—a type of colony which may be termed ‘astreiform.’! The foregoing statements explain my reasons for including the history of the name Phillipsastrea and the description of its genotype in a communication primarily concerned with Car- boniferous genera. I am deeply indebted to the late Prof. T. MecKenny Hughes, to Dr. A. Smith Woodward, Dr.’ F. L. Kitchin, Dr. W. G. Lee, and Mr. Peter MacNair for the loan of material (including various type-specimens) preserved in the Sedgwick Museum, 1 The term is frequently used by Edwards & Haime in their well-known Monograph, as also by other writers, to indicate the type of corallum in which the septa of the corallites are confluent, and does not in any way imply rela- tionship to the Astreide. I here employ the word in a slightly more extended sense, so as to include any Madreporarian colony (Rugose or Aporose) in which the corallites are united as above described. : , io y part 4] PAILLIPSASTR 4A HENNAHI, AND ORIONASTR4A. 281 Cambridge, the British Museum (Natural History), the Museum of Practical Geology, the collection of the Geological Survey of Scotland, and the Glasgow Museum & Art Gallery; and to Mx. Charles Edmonds, Prof. E. J. Garwood, and Prof. T. F. Sibly for use of the specimens that they had collected. My thanks and acknowledgment are also cordially extended to Dr. F. A. Bather, Mr. W. D. Lang, and Mr. C. D. Sherborn for their ever-ready advice and kindly assistance. IL. DEFINITION OF CERTAIN TERMS EMPLOYED. On account of the unfortunate lack of uniformity in the nomenclature of Corals, it seems advisable, before proceeding farther, to define clearly certain terms here employed. I denote the skeleton of a single individual, whether solitary or member of a colony, by the term ‘corallite,’ and use the term ‘corallum’ to mean the skeleton of the whole colony. In the ease of solitary or ‘simple’ corals the terms are synonymous. I divide the Rugose corallite into two regions: (1) the intrathecal region, which is built up of tabule or of tissue ontogenetically derived from tabule, and (2) the surrounding extrathecal tissue, which is built up of dissepiments. The junction of the tabular with the dissepimental tissue constitutes an annular wall—the theca. The corallite is usually clothed with an outer mural investment—the epitheca, the nature and develop- ment of which I have discussed in a previous publication.! The terminology applied to the septa will be found in a paper by Mr. R. G. Carruthers,? or in my own paper ‘On the Genus Aulophyllum. ® IIL. Tue Astr#1rorm CoraLLumM. Among the Rugose colonial corals there are certain genera, or certain species within a genus, in which the corallites have lost their epitheca, and are united by their dissepimental tissue.* The septa of adjacent corallites in the ‘astreiform’ colony tend to become confluent; but all stages of this development, from that in which it is incipient to that of perfect confluence, are to be found. In some cases, on the other hand, the. septa of one corallite do not extend to those of another, and leave an inter- vening space to the sole occupation of the dissepiments. Astreiform colonies make their appearance in widely divergent stocks and at different periods of time: being, it would seem, the ultimate terms in a progressive deve- lopment along a well-defined line. The steps towards this end are :— 1Q. J. G.S. vol. lxxi (1915) pp. 228-29. 2 Ann. & Mag. Nat. Hist. ser. 7, vol. xviii (1906) pp. 356-63. 3 Q. J. G.S. vol. lxix (1913) pp. 60-62. 4 An epithecal growth external to the whole corallum may, nevertheless, be retained in the astreeiform colony. Nee 282 DR. STANLEY SMITH ON 4UL/NA ROTIFORMIS, (vol. lxxu, 1. The simple or solitary coral. 2. The fasciculate colony, in which the corallites are not contiguous or do not press on one another, and are therefore cylindrical in form, as (for example) Lithostrotion martini. 3. The basaltiform colony, in which the corallites are contiguous, and are prismatic through mutual pressure. Although the corallum is massive, the corallites are defined by an epitheca, as (for example) Lithostrotion basaltiforme. And, finally, 4, The astreiform colony: for instance, Orionastrea. The examples quoted are from a single lineage. The number of genera represented by the astreeiform type of colony among the Rugosa does not appear to be numerous at an period, although the individuals may be extremely abundant.! Strombodes is fairly common in the Upper Silurian ; Phillips- astrea is very plentiful in the Upper Devonian; while, among the British Carboniferous Rugosa, the only examples of astreeiform genera and species that have come under my notice are Awlina and Orionastrea, Cyathophyllum regiwm Phillips, and a species of Koninckophyllam (or Diphyphyllum). All these forms are from the uppermost part of the Carboniferous Limestone Series—D, and higher beds. Certain Characters assumed by the Astizeiform Corallum in the Rugosa, and Factors that lave produced them. The coralla of Ovionastrea and of Aulina exhibit certain interesting characters—namely, a flattened form, an extensive development of extrathecal tissue, and epithecal growth external to the colony, even in cases where there is no trace of epitheca within it. (i) The form of the corallum is correlated with the mode of growth of the corallites. These may arise at any point on the surface of the corallum, but the region of most active growth is the margin. The peripheral corallites at first grow outwards, away from the centre of. the colony, and then bend upwards into a more ver teal position, as is shown in fig. 1 (p. 288). The extreme case of this form of colony is to be found, not in the Rugose but among the ‘tabulate’ corals—for example, Heliolites, where an almost horizontal base is often developed. 1 This type of corallum, while exceptional in the Rugosa, is characteristic of the massive Aporose corals—in fact, exceptions are rare.—but it must be remembered that the epitheecal covering to the skeleton (always present in Rugose simple corals) is often lacking in the case of Aporose simple corals. The mezandriform colony (due to repeated but incomplete fission so common among the Aporose stocks) is unknown in the Rugosa, and necessarily so, on the assumption that Rugose colonies increase by gemmation only, but never by fission. (See my notes on the subject in Q. J. G.S. vol. Ixxi, 1915, p. 233, and W. D. Lang, ‘Homceomorphy in Fossil Corals’ Proce. Geol. Assoc. vol. xxviii, 1917, p. 90.) : part 4] . PHILLIPSASTR.4:4 HENNAHI, AND ORIONASTR_EA. 283 Fig. 1.— Longitudinal section,of Orionastrea placenta (MeCoy), holotype of Sarcinula placenta, showing the corallite in its earliest stage. x 5 diameters. W. Tams photo. (ii) The development of dissepimental tissue can be shown to be progressively greater, as we follow the passage of dendroid and fasciculate into basaltiform, and these into astreeiform colonies. The case is not merely one of phylogenetic change, but may be observed to take place wherever Fig. 2.—Group of corallites two or more corallites of a den- of Lithostrotion martini. droid-fasciculate colony approach x 2 diameters. each other to the extent of con- tiguity. This is illustrated by the accompanying figure of a group of corallites of Lithostrotion martini Edwards & Haime. The phenomenon may be accoun- ted for, by supposing that two neighbouring polyps mutually ex- tend one. towards the other until they are in contact. This results in a stretching of the area of dis- sepiment-secreting tissue where the two touch, and in a_ consequent widening of the dissepimental area of the skeleton when secreted. (ii) The external epitheca, which stretches round the corallum as a continuous surface, usually displays pronounced rugosity. It differs neither in its origin nor in character from the epitheca that clothes a simple coral or an individual corallite ; yet its presence is often emphasized in descriptions of corals (especially if the corallum in question is discoid in form), and 284: DR. SLANLEY SMITH ON AULINA ROTIFOKMIS, (vol. lxxil, it is frequently defined as the ‘basal epitheca.’ It represents — merely the epitheca of the outer side of the peripheral corallites. * IV. Pururipsasrrma VOrbigny. Summary of Research. William Lonsdale (1840), in Sedgwick & Murchison’s ‘ Physical Structure of Devonshire, & on the Subdivisions & Geological Relations of its older Stratified Deposits, etc.’ Trans. Geol. Soe. ser. 2, vol. v, pt. 3, p. 697, pl. lvili, figs. 3, 34, & 346, described, under the name of ‘ Astrea (Stderastrea de Blainville) hennahir (sp. n.),’! a type of colonial coral since proved very character- istic of the Upper Devonian. The specimen (or specimens) upon which the species was established was stated to have been found at Barton, north-west of St. Marychurch, and to have been in the collection of Daniel Sharpe.? This collection is now in the Museum of Practical Geology. The specimen represented by fig. 3 can be recognized with absolute certainty in No. 6185; but the source of fig. Ba 1s. uncertain: it was probably another fossil. Edwards & Haime assumed such to be the case, although Lonsdale made no mention of there being two specimens. John Phillips (1841), ‘ Figures & Descriptions of the Paleozoic Fossils of Cornwall. Devon, & West Somerset’ p. 12, pl. vi, figs. 16aa, 16/36, 16), & pl. vu, fig. 15 pb, ustrated Bh hennahi by a drawing (fig. 16a a) clearly intended to represent the weathered surface of Lonsdale’s actual specimen, and by two figures—16 8b (enlarged) & 16 ¢ (natural size)—ot a polished specimen in his own possession. His description of the species does not differ materially from that given by Lonsdale-—in fact, Phillips appended to it the following statement :— ‘The above description is almost verbatim from Mr. Lonsdale. I have added the words in brackets, from a beautiful polished specimen in my possession (fig. 16c¢), which may be distinct.’ Phillips’s specimen unquestionably belongs to the same genus as Lonsdale’s; but the figure is not sufficiently exact to enable one to pronounce a certain opinion as to the species. Pl. vu, fig. 15 p, is that of a specimen which Phillips found in the Carboniferous Limestone of Flintshire, and figured under the impression that it was allied to, if not identical with, A. hennaht Lonsdale. As a matter of fact, the species bears no relationship at all to the Devonian coral, but is the form that I here deseribe as Ov7on- astrea phillipst (McCoy). Untortunately, this figure has been the cause of much subsequent confusion. It must be borne in ? J.B. Lamarck, ‘Systéme des Animaux sans Vertébres’ 1801, p. 371. ? Daniel Sharpe’s Collection was presented to the Geological Society by Henry Sharpe, his brother, in 1856, and passed into the possession of the Museum of Practical Geology in 1911, when the Society divided its collection between that institution and the British Museum—the British material going to the former and the foreign to the latter. e part 4] PHILLIPSASTR4#A HENNAHJ, AND ORIONASTR 4A. 285 mind, in connexion with the controversy which followed, that Phillips did not state that it was ‘A. hennahii, but, in his own words, a ‘specimen from Mountain Limestone much allied to Astrea hennahii. HK. A. Roemer (1843), ‘Die Versteinerungen des Hartzge- birges’ p. 5, pl. ii, figs. 13 a-136, & pl. iii, fig. 1, contributed to the literature of the genus a description and figures of two species allied to ‘Astrea hennahii, both from the Devonian ot the neighbourhood of Griind in the Harz, namely ‘ Astrea hennahii’ Lonsdale? and ‘ Astrea parallela N.’ :— 4 Astrea hennahii (pl. ii, figs. 13a & 1386).—The figure indicates a form differing from the type-specimen in regard to the smaller size of the corallites and the apparent presence of an epitheca. It suggests Phillipsastrea pentagona Goldtuss (Acervularia penta- gona Goldfuss auctt.). Astrea parallela (pl. iti, fig. 1).—In the figure of this species the intrathecal regions are very small and widely separated, and the septa are markedly confluent. Rcemer identified this species with Phillips’s polished specimen, also with the coral from North Wales, for which he assumed a Devonian age. Alcide d’Orbigny (1849), ‘Note sur des Polypiers Fossiles’ p- 12, introduced the generic name ‘ Phillipsastrea, and he defined the genus in the following words :— ‘G. Phillipsastrea, @’Orb., 1847.1 Le dessin qu’en a donné M. Phillips pourrait faire croire que ce sont des calices petits, 4 cloisons rayonnantes, entourées-de cloisons costales communes. On en connait deux espéces de Vétage déyonien. Ex.: Astrea parallela et hennahit, Phillips.’ In the ‘Prodrome de Paléontologie’ published the followmg year (1850), on pp. 106-107, he tabulated under various generic names most of the species that I have previously mentioned, namely :— : Inthostrotion hennahii (Lonsdale, pl. lviii, fig. 3). Actinocyathus hennahii (Phillips, pl. vi, fig. 16 a). Phillipsastrea parallela (Roemer, pl. iii, fig. 1). Phillipsastrea hennahii (Phillips, pl..vi, fig. 16 3, and pl. vii, fig. 15 D). Furthermore, he re-defined the genus Phillipsastrea thus :— ‘Ce sont des Siderastrea dont la columelle, au lieu d’étre styliforme saillante, est large et divisée en cloisons rayonnantes, comme chez les Columnastrea.’ H. Milne Edwards & J. Haime, in 1850, in the Introduction to their monograph on ‘British Fossil Corals’ pp. lxx—lxxi, defined the genus Phillipsastrea, and quoted as the type-specimen ‘ Astrea hennahi’ Lonsdale. Later, however (1851, ‘Mon. Polypiers Fossiles’ pp. 417 & 421; and again in 1852 & 1853, ‘ British Fossil Corals’ pp. 203 & 240), they restricted the name Phillipsastrea to a genus represented by Phillips’s Carboniferous species, which they identified with ’ Martin’s species Hrismatolithus radiatus (see p. 296), and quoted 1 This date is untenable, since the tract was not published until October 10th, 1849, fide C. D. Sherborn. 286 DR. STANLEY SMITH ON AULINA ROTIFORMIS, [vol. lxxii, “A. hennahii Lonsdale’ fig. 3 (excluding fig. 3 a1) as the type of “a new genus—Smithia.. Phillipsastrea, they stated, possessed a columella, and Smthza did not. While acknowledgment is due to the service rendered by these ‘authors in separating the Devonian and Carboniferous forms, the legitimacy of their use of the name Phillipsastrea must be challenged: because, although the figure of the Carboniferous species was quoted by d’Orbigny, he clearly intended the name Phillipsastrea to be applied to the Devonian species, since he framed his definition of the genus upon the figures and descriptions of those corals. It is quite certain that, in quoting fig. 15D, he did so under the impression that the species was identical with that represented by figs. 1666 & ec. It is also true that he mentions the presence of a columella as one of the generic characters of Phillipsastrea; but, here again, he was misled by the figures that he quoted, and interpreted as a coral structure the interstitial calcite which filled the intrathecal region and stood out from the weathered surface as a boss. The name Phillipsastrea must, therefore, be retained for the Devonian genus, and a new one given to the Carboniferous species. The actual genotype of Phillipsastrea is the species to which Phillips’s specimen be- longed, but there is nothing in the figures to warrant the assumption that it was not specifically identical with the type- specimen of Astrea hennahi—it may, or may not, have been. In either case the two specimens belong to the same genus, and, even supposing that they represented different species, Phillips’s specimen is lost, and we are at liberty to closeé’a neogenotype-— besides, in so doing, we rectify an anomaly. ~~ Edwards & Haime also described under the name Acervularia 2 (‘Polypiers Fossiles’ 1851, pp. 416, 421, and ‘ British Fossil Corals’ 1853, pp. 236-41) certain species of Devonian corals, which differed from Phillipsastrea (Smithia of these authors) only in the presence of a thin but distinct epitheca. \ T agree with Frech in regarding the two as congeneric—in fact, careful examination of material shows that certain species of the one are identical with certain ‘species’ of the other, and that the development (or, rather, loss) of the epitheca is one of degree only. Indeed, in a single corallum, epitheca may be present in one region and absent in another. Frederick McCoy (1851), ‘ British Paleozoic Fossils’ p. 72, 1 They founded upon this figure a new species, Smithia pengellyi, and __ thereby definitely fixed for us the type of A. hennaht. ~ 2 The genus Acervularia was founded by Schweigger, ‘Handbuch der Naturgeschichte’ &c., 1820, p. 418, upon the figure and description of a Silurian coral from Gothland by Henry Fougt, ‘ Corallia Baltica’ in ‘ Ameeni- tates Academic’ pp. 92-98, pl. iv, fig. ix and No. 2. Fig. ix bears a strong superficial resemblance to weathered specimens of the so-called Devonian Acervularia, but it is not at all likely that these are congeneric with the coral figured by Fougt. No. 2 (also fig. viii, referred to as a variety of the same ‘ madrepora composita’) undoubtedly represents Acervularia luxurians. —— part 4] PHILLIPSASTR.£A HENNAHI, AND ORIONASTR EA. 287 assigned Astrea hennahi of Lonsdale to Dana’s genus drachno- phyllum. Fritz Frech (1885), ‘Die Korallenfauna des Oberdevons in Deutschland’ Zeitschr. Deutsch. Geol. Gesellsch. vol. xxxvu, pp- 44 et segg., has anticipated me in merging Smethia and the Devonian species of ‘Acervularia’; he also questioned the right of including Phillipsastrea radiata in the same genus. Rudolph Schifer (1889), ‘On Phellipsastrea WOrb., with special reference to Phillipsastrea radiata 8. Woodward sp. & Phillipsastrea tuberosa McCoy sp.’ Geol. Mag. dec. 3, vol. vi, 1889, pp. 398-409, pl. xii, reviewed the literature of Phillipsastrea, and redescribed the species of Orvzonasti¢a mentioned in the title of his paper. He examined McCoy’s types at Cambridge and specimens of Orionastrea in the British Museum, and came to the conclusion that these forms did not possess a columella, and therefore the basis upon which Edwards & Haime separated Phillipsastrea and Smithia had no real existence. I must modify Schifer’s assertion, that the Carboniferous species do not possess a columella,! by the statement that a columella may be present or absent; although in most cases it is present, the columella is admittedly merely the dilated prolongation of the counter-septum, but in this origin it does not differ from the columella in other Rugose genera. Even if Schiafer had been correct in his statement concerning the absence of a columella in Orionastrea, there still remains the very striking character of Phillipsastrea that is not developed in the former: namely, the peculiar septal dilations already described. Several other paleontologists have discussed the genera Phil- lipsastrea and Smithia; but, in every case, they have contented themselves with the question of the identity of the two ‘ genera,’ and have neglected to inquire into the legitimate application of the names. In fairness to these authors, it must be stated that they decided that the genera were identical, and consequently the other question did not concern them. Generic Characters. The corallum is composite and massive; the corallites are united by their dissepimental tissue, or are separated by a thin epitheca only : in the former case the septa are often confluent. In the 1 Tt should be noted that some of Schiafer’s own figures (5, 6, & 7) show the columella, and that, although in figs. 1 & 2 no columella is represented, yet in the actual specimens (R. 541 & 56740) the columella is prominent. Schafer quoted Kunth (Zeitschr. Deutsch. Geol, Gesellsch. vol. xxii, pp. 30-87, pl. i, figs. 4a—4.d) to support his contention that there was no fundamental difference between the Devonian and the Carboniferous species of Phillipsastrea; but, as a matter of fact, Kunth’s figure of Ph. hennahi bears very little resemblance to theactual type, and may be dismissed from the argument. The major septa are shown in these figures to unite in the centre of the corallite, and so form a ‘columellarian tubercle’; moreover, they do not’ appear to dilate at the theca. 288 DR. STANLEY SMITH ON AULINA ROTIFORMIS, { vol. lxxu, mature corallite the septa assume 2 radial symmetry, although the bilateral symmetry of its early growth-stages is sometimes traceable in the adult state through the presence of a cardinal septum shorter than the rest, and is indicated in some species. by a prolonged counter-septum. Both the major and the minor septa dilate at the theca, and the latter terminate there ; but the major septa, in an attenuated form, advance into the intrathecal region, and frequently dilate again at their axial edge. The central part of the corallite is occupied solely by the tabule. Confluence and dilatation of septa are characters comparatively rare in the Rugosa, but of very frequent occurrence in the Aporosa. Genotype: Phillipsastrea (Astrea) hennahi (Lonsdale). Type-specimen of Ph. hennahi: No. 6185, Museum of Practical Geology. The type is possibly the holotype, but is much more probably one of two syntypes. Edwards & Haime assumed the Jatter to- be the case, and chose it as the lectotype. Description of Type-Specimen of Phillipsastrea hennuhi (Lonsdale). (Pl. XXII, figs. 1—4.) No. 6185 & Sections 28348-28349, Museum of Practical Geology (‘ Daniel Sharpe Collection’—part of the ‘ Geological ee Collection’). For literature and history, see p. 284. The specimen is a weathered and rounded block measuring some 10 cm. along its greatest length and about 4 cm. through its thickest part. There are two plane polished surfaces, transverse and longitudinal respectively to the direction of growth. The material is translucent and of a grey colour, except near the surface, where it is bleached and opaque. Superficially, it is iron-stained. Weathering has reacted more vigorously upon the coral tissue than upon the interstitial calcite, and so has engraved the coral structures into the stone. The calcite occupying the interseptal spaces stands out from the surface, and counterfeits actual septa, while that filling up the intrathecal region projects as a central prominence. External Characters. None present. Internal Characters. (Pl. X X11, figs. 2-4.) Transverse section.—Epitheca almost but not entirely absent, weak traces being detectable here and there. Confluence on the part of the septa is not highly developed: the septa may be confluent; but, for the greater part, those of one corallite abut against those of another, or intermingle in a confused network. The thee are of uniform size, measuring 3 mm. in diameter; part 4] PHILLIPSASTR_£A HENNAHI, AND ORIONASTR_EA. 289 but the extrathecal tissue is very unevenly developed, so that the intrathecal regions may lie closely together or widely apart. There are twenty-six septa present im the majority of the corallites, and the dilation of these at the theca extends for a distance of about 1mm. ‘The dilation at the axial edge is but feebly developed in this specimen. Longitudinal section.—The dissepiments are small and crowded ; the tabule are closely set. Shallow concave tabule occupy the central part of the intrathecal region, and these are supplemented near the theca by smaller proximally-inclined plates. The figure should be compared with that of Aulina, p-292. The peripheral plates do not appear to be present in all species of the genus. Notes upon Lonsdale’s Figures and Statements. Fig. 3 combines a view of the weathered surface and the ibveranalle: -polished face of specimen No. 6185. The features displayed have been most accurately drawn,! and the portion included in the figure can be precisely defined by a line. It is evident that Lonsdale (and others after him) mistook the etched- out interstitial calcite of the surface for actual coral-tissue. Fig. 36 shows, on an enlarged scale, the longitudinal section through a few septa as seen in the polished surface. It supplies no useful information. Fig. 3a illustrates a transverse section, but, as has been pre- viously mentioned, this figure has probably been supplied by another specimen. The corallites are represented as being larger, and the septa as being more numerous and more perfectly confluent, than those in the specimen figured in fig. 8 a—the type. I have searched through the Daniel Sharpe Collection, and, among those examined, the specimen with which the figure most closely agrees is No. 6192, a thin polished slab of irregularly pentagonal shape. In this specimen the thece are larger (5 mm. in diameter) and the septa are more numerous (about 36) than in the type-specimen ; and, furthermore, the dilation of the septa at the theca is less localized and consequently not so obvious. The characters displayed agree with Lonsdale’s statements that there are 36 septa (or rays, as he termed them), ‘ unequal in length and breadth, and of a crenulated structure.’ The last remark is of some importance in identifying the specimen: it refers to a feature not untrequently found in mineralized corals, and is strongly shown in the present case—a feature due to the obli- teration or partial obliteration of the dissepiments, except where these meet the septa. Despite his assertion that the number of septa is 36, Lonsdale only shows about 30 in this figure. The origin of his fig. 8@ must, nevertheless, remain uncertain. 1 The drawing is reversed on the plate, so that the right side in the original is the left in the figure—a common occurrence in plates of the period. 290 DR. STANLEY SMITH ON AULINA ROTIFORMIS, [vol. xxii, V. AULINA, gen. nov. Family ? Phillipsastreide. | The structure of Awlina is in most respects similar to that of Phillipsastrea, but it appears to carry to a further stage of development the septal characters peculiar to the latter genus. Generic Characters. The corallum is massive, and the corallites are united by their extrathecal tissue. All the septa dilate at the theca, and those of the major cycle again dilate at their axial edges, in such a manner as to fuse together and so to form a cylindrical wall or tube within the theca (see fig. 4, p. 292). Stability of character is a feature strongly maintained. Genotype: -dulina rotiformis 8. Smith. Type-specimen of 4. rotiformis. The holotype has been cut in two: one half is in the British Museum (R. 17497), and the other in the Sedgwick Museum, Cambridge. AULINA ROTIFORMIS, Sp. Nov. 1910. Phillipsastrea radiata (Martin), S. Smith, Trans. Nat. Hist. Soc. Northumberland, &c. n. s. vol. iii, pt. 3, pp. 629-30. a 1912. Phillipsastrea radiata (Martin), E. J. Garwood, Q. J. G.S. vol. Ixviii, pp. 542-43. : 1916. Aulina rotiformis S. Smith, Abs. Proc. Geol. Soc. No. 995, pp. 2, 3. Aulina rotiformis was recorded as Phillipsastrea radiata from South Northumberland by myself and from Hurdorthwaite Moor (North Yorkshire), by Prof. E. J. Garwood, in the publications above cited. No description of the form was given in either paper. I remarked that the form was plentiful in and charae- teristic of the Fell Top Limestone around Harlow Hill, while Prof. Garwood stated that it was somewhat abundant in the Botany Beds on Hurdorthwaite Moor and that he had not found it ¢ sttw at any other horizon.! iExternal Characters. The corallum is massive, and is typically depressed in form. The corallites are not defined by epitheca ; consequently, the only epithecal development present is that external to the whole colony. The upper or distal surface of the corallum is generally flattened and fairly uniform. The thecze measure about 1:5 mm. in diameter; they are near together and regularly arranged, as compared with those of Orionastrea, lying 1 to 3 mm. apart.® 1 He recorded the finding of an angular and unworn specimen of Phillips- astrea radiata resting upon the Tyne Bottom Limestone in High Cup Gill and of a cast of the same from the Drift of Ravenstonedale. The High-Cup Gill specimen was correctly identified, and therefore not Aulina. but Orion- astra. * In one of the specimens collected from the Botany Beds the calices are larger and proportionately more widely separated, and the surface more mammiferous, than in the Harlow-Hill material. part 4] PHILLIPSASTR£ZA HENNA, AND ORIONASTR AA, 291 The general surface of the corallum (as will be noticed in the text-figure) rises towards the thece, so as to form mound-like borders to the calicular depressions, suggesting the idea of minute Fig. 3.—Distal surface of the type-specimen of Aulina rotiformis, sp. nov. x 2 diameters. A. H. Harrow photo. [Fell Top Limestone, Harlow Hill (Northumberland). One half is in the British Museum (Natural History), R. 17497, and the other half in the Sedgwick Museum, Cambridge. | craters. Within the calice the tubular wall terminates as an elevated ring, forming a hollow axis against which the major septa abut like the spokes of a wheel. Internal Characters. (PI. XXII, figs. 6-11, & text-fig. 4.) The most distinctive feature is the tube traversing the tabular tissue. Within this tube the tabule are horizontally disposed, and are remarkably flat; externally to it, they take the form of cones perforated by this axial tube. The dissepiments are very small and regularly formed. All the septa dilate at the theca, and there the minor series abruptly terminate. In these respects Awlina is similar to Phillipsastrea. The major septa, in an attenuated form, advance towards the tube, to which their axial edges appear to be fused. Actually, the septa dilate along their axial edges in such a way as to present in cross-section the letter T, and form the tube by the mutual fusion of these peculiar dilations into a perfect wall. Confluence of the septa is well developed. Transverse section.—In this section the septa display the 292 DR. STANLEY SMITH ON AULINA ROTIFORMIS, [ vol. Ixxii, characters above mentioned, and the tube appears as a polygonal ring. The number of septa in each cycle is usually eleven; very rarely does it exceed Fig. 4.—Diayram combining a transverse twelve or is it less and a longitudinal section of Aulina than ten. The theca rotiformis. x about 18 diameters. is emphasized by a very slight secondary thickening of the dis- sepiments which form it. The diameter of the theca is about 1°5 mm., and the diameter of the tube about 0°5 mm. ‘The dilation of the septa at the theca does not extend far in a peripheral direction (as indicated in the figure ). _ Longitudinal section (Pl. XXII, fig. 8)—The tube is represented by two vertical walls limiting the horizontal tabule ; the tabule external to these walls are seen as inclined plates. Ontogeny. In the earliest growth-stages observed in duw/zna the septa then present meet in the centre of the corallites (Pl. XXII, figs. 9-11). Later, these leave the centre and form the initial stage of the tube, while new members are being added to their number. Although, by the time that it reaches the adult stage, the form displays a radial symmetry, the insertion of septa follows the general rule appertaining to the Rugosa as a class. A cardinal fossula (in some cases very conspicuously, but in others less pro- minently displayed) and a distinct pinnate symmetry may be observed in the immature stage. Phylogeny. The morphology of Awlina clearly points to its derivation from Phillipsastrea: the axial tube in the former and the dilated edges of the septa in the latter are probable stages in a continuous line of development. The interval of time, however, separating the epochs in which the two genera lived respectively is consider- able—practically the whole of the Lower Carboniferous Period, and no intermediate forms have as yet been recorded from the beds of that age. Horizon and locality.—Awlina occurs abundantly in the Fell Top Limestone in the vicinity of Harlow Hill,! where the ! Harlow Hill lies on the Neweastle and Carlisle Road, 11 miles west of the former city (Quarter Sheet 105 N.W. New Series 14). : : part 4] PAILLIPSASTR.ZA HENNAHI, AND ORIONASTRAA. 293 outcrop of that limestone is exposed by a chain of old quarries extending from the south side of the village to Stob Hill, near Cheesburn Grange, some 2 miles in a north-easterly direction. In that locality the Fell Top Limestone lies somewhere about 1000 feet above the Great Limestone, and is between 20 and 30 feet thick (I have not been able, however, to expose the actual base nor find the exact top). The lower portion consists of well- bedded crystalline limestone; the middle is much more argillaceous, and is divided up by bands and partings of shale; but the upper part, again, is more massive and more purely calcareous. The fauna does not materially differ from that of the lime- stones between it-and the Great Limestone, except in a few (but noteworthy) particulars —namely, the presence of Aulina and the abundance of Lithostrotion ct. junceum (but approaching L. irregulare), which oceurs as small stunted colonies of tortuous growth-habit. These colonies form in places a band at the top of the lower and more massive portion, and appear to have been buried as they lived by the overlying silt (now calcareous shale). Dibunophyllum near y Vaughan (small, conical and often twisted forms) is plentiful, and Lithostrotion portlocki is common. The most striking feature of the brachiopod-fauna is the great abundance of Productus latissimus. Fenestella is present in large quantity. The other locality in which Aulina has been found is Hurdor- thwaite Moor, from a thin series of limestones and shales inter- calated in the Millstone Grit, to which series Prof. E. J. Garwood has given the name of Botany Beds,! after the farm north of their outcrops. Prof. Garwood? defines the stratigraphical position and geo- graphical situation of the beds, and describes their character in the following passages :— ‘The beds lie some distance [200 feet or so] above the base of this series [the Millstone Grit], and occupy a tract extending for 2 miles in a general east-and-west direction on Hurdorthwaite Moor, south of Botany Farm. They are repeated by a strike-fault with a southward upthrow, which causes an extension of the outcrop round the south of the moor, and affords additional exposures. ‘The beds are well seen in several old quarries at Scoletree, Howgill Head, and Greenhill. They consist of a few feet of compact crystallized lime- stone at the base, which is overlain by some 10 to 15 feet of impure calcareous and ferruginous shales’ [and I may add—especially towards the top—lime- stone-bands]. The sequence suggests at once the Fell Top Limestone, and the resemblance is still further strengthened by the characteristic appearance of the pale crumbly shale packed with Fenestella. Faunistically also the two series bear a striking similarity, since, in addition to Aulzna, the Botany Beds have yielded the same 1 Botany Farm lies 23} miles west-south-west of Romaldkirk, and about half a mile north of the Reservoir (Quarter Sheet 102 S.E. New Series 31). 2 Q. J. G.S. vol. Ixviti (1912) p. 542. . 294: DR. STANLEY SMITH ON AULINA ROTIFORMIS, [ vol. \xxii, stunted and twisted forms of Lithostrotion junceum, the same dwarfed and conical forms of Débunophyllum, and also Litho- strotion portlocki.| 'The list of fossils from these beds published by Prof. Garwood? is very similar to that of the Fell Top Limestone which | previously published.? Thus, in their agreement in faungl contents, lithological cha- racter, and stratigraphical position there is strong evidence for correlating the Botany Beds with the Fell Top Limestone.* This correlation being accepted, the bed represents the highest known limestone of standard conditions ® occurring at a horizon occupied in most places by arenaceous deposits (‘ Millstone Grit’) or argil- laceous beds (‘ Pendleside Series’). Assuming that subsequent investigation does not discover Awlina in beds lower than those which it is known to occupy, I suggest that the horizon in which it occurs be removed from the Dibunophyllum Zone to con- stitute a new zone—the Auwlina Zone, which, in accordance with Vaughan’s system, would be indicated by the initial letter A: a definite limit would thus be given to the Dibunophyllum Zone, which at present tends to extend indefinitely upwards. The Postdonomya Zone® in Gower and elsewhere is approximately equivalent to the Aulina Zone, but represents a shallow-water phase, and, appearing earlier in time in some localities than in others, must also represent beds belonging to ‘ D,’ as well as ‘A.’ The question as to whether the Avonian Stage should include the Aulina Zone, or be restricted so as to end with the Dibwno- phyllum Zone, is worthy of future consideration, but cannot be discussed here. VI. ORIONASTRHA, gen. Nov. Family : LirHostRorronvTip az. Species.— O. phillipsi (McCoy). Genotype. O. placenta (McCoy). O, ensifer (Edwards & Haime). The species grouped together under the name Ovzonastrea are closely allied to the species of Lithostrotion, and appear to . 1 Although I paid a visit to these beds in the summer of 1915, in company with Mr. C. T. Trechmann and Dr. D. Woolacott, and found Awlina and other forms, the principal source of information for the above facts was the material collected y Prof. Garwood, which he kindly allowed me to examine. 2 Q. J. G. S. vol. Ixvili (1912) p. 542. 3 Trans. Nat. Hist. Soc. Northumberland, n. s. vol. iii, pp. 630-31. 4 The identity of the two limestones was suggested long ago by J. G. Goodchild, but he does not appear to have ever published these views— fide Prof. G. A. Lebour. ® Standard conditions as opposed to a phase: that is, any interruption in a sequence of deposits formed under conditions which have been fixed for the purpose of zoning. See A. Vaughan, Proc. Bristol Nat. Soe. ser. 4, vol. i (1906) p. 79. ° B. E. L. Dixon & A. Vaughan, Q. J. G.S. vol. lxvii (1911) p. 494. i i ert alee Pe + gal a part 4] PHILLIPSASTR4ZA HENNAHI, AND ORIONASTR ZA. 295 represent the phylogerontic (or senile) stage in the history~of that stock. The separation of Lithostrotion ensifer from the other species of that genus and its inclusion here may encounter reasonable objection, since the differences that it exhibits are cer- tainly not great; but, as 1t possesses (although in a less-developed condition) those features which distinguish the genotype of Orionastrea, I believe that the step is justifiable. Generic Characters. The corallum is composite and massive, and the corallites are either defined by a thin epitheca, or, in the more typical forms, by no epitheca at all; in this latter case the corallites are united by the dissepimental tissue, and the septa are confluent. Both major and minor septa are well developed, and a columella is present (except in O. placenta). The characters of Orionastrea are essentially those of Litho- strotion, but in a modified form, and their structures are more unstable and variable. Even in the same corallum the difference between the corallites is often very marked, as in Pl. XXIV, fig. 2. From a large quantity of material examined three distinct types of Orionastrea can be readily isolated, represented respectively by the three species here described; but a large proportion of the specimens falls between two species rather than within one of them, and might, therefore, be ascribed to the one species equally as well as to the other. To O. ensifer specific rank can with little hesitation be accorded, and it is easily distinguished from O. phillipsi and O. placenta. Furthermore, it represents almost exclusively the genus in the Bristol area, whereas it is verv rare in North Wales, where O. phillipsi is so abundant and where O. placenta also occurs. I have considerable doubt as to whether, in the strict zoological sense, it is correct to regard O. phillipst and O. placenta as separate species: both occur together, and the difference may be nothing more than that between individual colonies of the same species. Since, however, in the absence of sufficient evidence this question cannot be settled, and since it may prove useful to have separate names for the two types, 1t seems permissible and even desirable to recognize MeCoy’s species rather than keep them merged under Martin’s name ‘ radiata.’ Genotype: O. phillipsi (McCoy). Type-specimen of O. phillipsi: No. 2138.a, Sedgwick Museum, Cainbridge. (Lectotype chosen from two syntypes.) Summary of Research. The earliest figures of Orionastrea are those of a highly siliceous specimen from Winster (Derbyshire), published -;by William Martin in 1809, ‘ Petrificata Derbiensia’ pl. xvii, figs. 2 & 3. Q. J.G.S. No. 288. Z 296 “DR. STANLEY SMITH ON AULINA ROTIFORMIS, (vol. xxi, Martin described the specimen under the name ‘ H7ismatolithus tubiporites (radiatus1)’, and stated that the coral was built up of ‘straight tubes connected by transverse dissepiments or partitions ... embedded in a calcareous cellular mass.’ From his fig. 2 these ‘tubes’ can be readily recognized as cherty casts of the intrathecal region of corallites, and the ‘transverse dissepiments ’ as silicified layers in the extrathecal tissue. The confluent character of the septa is mentioned by Martin, and is well shown in the figure. John Fleming (1828), in ‘A History of British Animals ’ p- 529, waeonan Martin’s species as ‘ Tubipor a radiata. This citation appears to have been overlooked hitherto, and those paleontologists who refuse to acknowledge Martin’s claim to the authorship of the species which he named, on the grounds that his terminology did not conform to the laws of Linnzan nomen- clature, credit the trivial name ‘radiata’ to Samuel Woodward, ‘Synoptical Table of British Organic Remains’ p. 5 (1830). John Phillips’s ‘ Figures & Descriptions of the Paleozoic Fossils of Cornwall, Deron, & West Somerset’ (1841) contained the figure (pl. vii, fig. 15 p) of the coral from the Carboniferous Lime- stone of Flintshire which Phillips believed to be allied to ‘ Astrea hennahii’ (see p. 284), but which actually belonged to the same genus as Martin’s ‘ Hrismatolithus tubiporites (radiatus). ‘The form is common, and very characteristic of the Lower Carboniterous of North Wales. Frederick MeCoy, in 1549, ‘On some New Genera & Species of Palzonoic Corals & Foraminifera’ Ann. & Mag. Nat. Hist. ser. 2, vol. iui, pp. 124-25, and in 1852, ‘A Systematic Description of the British Paleozoic Fossils in the Geological Museum of the University of Cambridge’ p. 110, pl. 118, figs. 8 & 9, described, under the generic name Sar cinula,2 three ‘species’ of Orion- astred, namely, S. tuberosa, S. placenta, and S. phillipsi. he last he rightly identified with the coral from North Wales figured by a Phillips. Brief notes upon the ieee specimens will be found on pp. 299-800. H. Milne Edwards & J. Haime, ‘Monographie des Polypiers Fossiles’ 1851, pp. 448 & 449 and * A Monograph of the British Fossil Corals’ 1852, pp. 203-204, pl. xxxvu, fig. 2, merged S. phillipsi and S. placenta, and identified these with Martin’s coral, but retained as a separate species S. tuberosa. Now that IT have cut tie type-specimens, I consider it more desirable to recognize S. phillips: and S. placenta as distinct species, and to merge S. twberosa® with the former. Moreover, on grounds 1 The name ‘radiatus’ has not been retained, for reasons subsequently explained. 2 J. B. Lamarck, ‘ Histoire Naturelle des Animaux sans Vertébres’ vol. ii (1816) p. 222 (for two recent forms, one of which he confuses with a Silurian fossil). 3 H.M. Edwards & J. Haime, ‘ Polypiers Fossiles des Terrains Paléozoiques’ p. 447, include an American species, P. vernewili in the same genus; but, arguing from the species so named in the British Museum, I do not consider the form congeneric with the British species. part 4] PHILLIPSASTRLA HENNAHI, AND ORIONASTR-EA. 297 already adduced (pp. 285 e¢ seqq.), these authors restricted the use of A. dOrbigny’s. generic name Phillipsastrea to these species—by which name they have since been known. In the same works (pp. 442-43 and p. 193 respectively) Edwards & Haime described a new species of Lithostrotion (L. ensifer), which I propose to include in the genus Orionastrea, although acknowledging its equal claim for inclusion in either genus: it constitutes, in fact, the link connecting Orionastrea phillipsi with the typical forms of Lithostrotion. James Thomson, in 1883, ‘On the Development & Generic Relation of the Corals of the Carboniferous System of Scotland ’ Proce. Phil. Soc. Glasgow, vol. xiv, pp. 394-96, pl. iv, figs. 1, 1 a, 16, Le, & 2, recorded Orionastrea (as ‘Phillipsastrea’) from Blackridge (Linlithgowshire), and thus widened its known area of distribution to include Scotland. Moreover, he demonstrated its instability of character by enumerating the varieties that he ob- tained from a very limited horizon at that locality, which he stated was the only one in Scotland where the genus had been found. He specified four varieties, distinguished by the following characters :— Ist variety. Columella absent, tabule regular. 2nd do. do. do. do. irregular. 3rd do. do. present. 4th do. do. do. mammilliform surface (=S. twherosa (MeCoy)). He illustrated his description by figures of : (a) a form in which the counter-septum alone invades the intrathecal region, and constitutes a slender and inconstant columella ; (b) a form in which several septa unite in the centre of the intrathecal region. I consider it probable that his varieties 1 & 2 are O. placenta, and that 3 & 4 are O. phillipst. Rudolph Schafer’s (1889) contribution to the literature, ‘On Phillipsastrea dOrb., with especial reference to Phillipsastrea radiata 8. Woodward sp. and Phillipsastrea tuberosa_ McCoy sp. Geol. Mag. dec. 3, vol. vi, pp. 898-409, pl. xii, has already been discussed (p. 287); wherefore here I merely desire to draw attention to the text-figures on p. 403, representing calices over the floors of which a number of septa have straggled: the figures are intended to demonstrate the absence of a columella. I admit the accuracy of the figures; but unfortu- nately they illustrate selected calices, while others of the same specimens in which the columella can be detected have been overlooked. Schafer followed Edwards & Haime in merging McCoy’s species S. phillipsi and S. placenta, and retaining his S. tuberosa. A. Vaughan (1903), ‘ Notes on the Corals & Brachiopods obtained from the Avon Section & preserved in the Stoddart Collection’ Proc. Bristol Nat, Soc. n. s. vol. x, pp. 109-10, included 72 -~ 298 DR. STANLEY SMITH ON AULINA ROTIFORMIS, [ vol. Ixxti, remarks on Lithostrotion ensifer Edwards & Haime; and in 1905, in ‘The Paleontological Sequence in the Carboniferous. Limestone of the Bristol Area’ Q. J. G. S. vol. xi, p, 199, he stated the exact paleontological horizon at which the species. occurs—namely, the upper part of the Dibunophyllum Zone: that is, Subzone of Lonsdaleia floriformis (D,). ORIONASTRMHA PHILLIPSI (McCoy). 21809. Hrismatolithus tubiporites (radiatus) W. Martin, ‘ Petrificata Derbiensia” pl. xviii, figs. 2 & 3. [O. phillipsi or O. placenta. | 21828. Tubipora radiata (Martin), J. Fieming, ‘A History of British Animals’ p. 529. [O. phillipsi or O. placenta. | ? 1880. Tbonor radiata (Martin), S. Woodward, ‘A Synoptical Table of British Organic Remains’ p. 5. [O. phillipsi or O. placenta. | 1841. ‘Specimen from the Mountain Limestone much allied to Astrea hennahii Lonsdale’ J. Phillips, ‘ Paleozoic Fossils, &c.’ p. 12 & pl. vii, fig. 15 D. 1849. Sarcinula phillipsi FE. McCoy, Ann. & Mag. Nat. Hist. ser. 2, vol. ui. p. 125. Sarcinula tuberosa. Id. ibid. p. 124. 1850. Phillipsastrea hennahii Lonsdale, partim A. @Orbigny, ‘Predrome de Paléontologie’ vol. 1, p. 107. 1851. Phillipsastrea radiata (Martin), H. Milue Wdwards & J. Haime, ‘ Polypiers Fossiles des Terrains Paléozoiques’ p. 448. Phillipsastrea tuberosa (McCoy). Id. ibid. p. 449. 1851. Sarcinula phillipsi KF. McCoy, ‘ British Paleozoic Fossils’ p. 110. Sarcinula tuberosa. Id. ibid. p. 110, pl. iB, figs. 8 & 8a. 1852. Phillipsastrea radiata (Martin), H. Milne Edwards & J. Haime, ‘ Mono- graph of the British Fossil Corals’ Pal. Soc. p. 203, pl. xxxvil, figs. 2 & 2 a. Phillipsastrea tuberosa (McCoy). Id. ibid. p. 204. 1888. Phillipsastrea radiata (Martin), J. Thomson, Proc. Phil. Soc. Glasgow, vol. xiv, p. 394, pl. iv. figs. 1, la, 16, & 2. 1885. Phillipsastrea radiata (Martin), F. Frech, Zeitschr. Deutsch. Geol. Gesellsch. vol. xxxvii, p. 48. Phillipsastrea tuberosa (McCoy). Id. ibid. p. 48. . 1889. Phillipsastrea radiata Martin, partim R. Schafer, Geol. Mag. dec. 3, vol. vi, p. 401, pl. xii, figs. 1, 3, 4, 7, 8, & 9, text-tigs. 1-3 (p. 403). Phillipsastrea tuberosa (McCoy). Id. ibid. p. 407, text-figs. 4-6 (p. 403). Phillipsastrea radiata (Martin) includes, in all cases quoted, Sarcinula phillipsi and S. placenta. External Characters. (Pl. XXIII, figs. 1, 3, & 4; Pl. XXIV, tig. 2.) The corallum is typically depressed, and the distal surface is usually flat. Nevertheless, the species occurs in more tumular masses, and may display a mamiillate surface (see Pl. XXIII, fig. 4). “The dissepimental tissue, which constitutes the greater part of the skeleton, is often unevenly developed ; consequently, the cali- cular depressions may be near together, or widely separate: these depressions are not infrequently “pounded by a sharply-elevated border (Pl. XXIII, fig. 1). A columella is present, and is often prominent. Hpitheca clothes the lower surface of the corallum. ENNAHI, AND ORIONASTR.EA. 299 (Pl. XXIII, figs. 2 & 5; XIV, fic. 1.) septa are confluent; the inequality ber number of the major septa is the latter advance far into the r major septa may extend to wh is prolonged into the centre to form a conspicuous columella: en becomes isolated from the rest of (Many of these features are is In section. ) s from 30 to 40 in all, 15 to ough varying in size, measure Transverse seétion between the minor and @ ‘not marked, since on intrathecal region; bu the columella. The & of the corallite, and the the swollen axial portion of the septum in the ephebic stage. displayed equally well ingthe alice The number of septa pres 20 in each cycle, and the theee on the average 2°5 mm. in @ Longitudinal section tabulze are conical in form, the columella is persisten dissepiments are fine. The © absence of the epithecal boun to the corallites, and perhaps to a small extent the less uniform disposition of the tabule, distinguishes the species from those of Lithostrotion. 2 to more typical examples. t to much variation: a less with O. placenta, and less epta merges the members of The foregoing description applies on The characters enumerated are subj well-developed axis leads to eom perfect confluence on the part this species with those of O {[ Sarcinula phillipse Notes upon the Type-Speceimens | berosa McCoy in the McCoy (genotype) and 8. 7 Sedgwick Museum, Cambr: 1. SARCINULA PHILLIPSI: Specimen Merionethshire. (PI. The species was described, bu figured, by McCoy. ‘There are two syntypes mounted on the same tablet, and of these I have chosen the first (213 a, Pl. XXIII, fig. 1) as the lectotype. ~ Photographs of both specimens are reproduced in Pl. XXIII, which illustrate sufficiently well the external characters to obviate the necessity of detailed description. The two types of calices ~ present in the specimens shoulc be observed, namely, those in which the theca is marked by a raised border (fig. 1) and those in which the extrathecal region simply rounds off at the intra- thecal depression (fig. 3, and certain calices in fig. 1). In the ‘ease of 213 (fig. 3) the intrathecal depressions are filled with matrix. The internal characters of the lectotype agree with the general description, and need not here be repeated. ‘The number of septa present ranges from 32 to 40. & 2136; trom Corwen, 3.) é 300 _ DR. SPANLEY SMITH ON AULINA ROTIFORMIS, [vol, xxu, 2. SARCINULA TUBEROSA: Specimen 212; found in Derbyshire, and presented by W. Hopkins. (Pl. XXIII, figs. 4 & 5.) The holotype is a silicified specimen, perforated by several large drusy cavities. The coral structures are nevertheless well pre- ‘served, and the interstices are but incompletely filled with mineral deposit. The distal surface, which is in a fair state of preservation, displays in a somewhat exaggerated form a mam- millate development. The thecz are ‘slightly larger than usually observed, and the columella is not very prominently shown in the transverse section (Pl. XXIII, fig. 5). The number of septa present is smaller than is generally the case, being only about 30: otherwise the characters of this form are in agreement with those of O. phillipsi. Its one pronounced and distinguishing feature—the mamimilli- form calices—may be found associated in a single corallum with calices of the types illustrated as typical of O. phillipsi. It seems therefore inadvisable to regard the form, simply because of its external characters, as a separate and distinct species. McCoy illustrated (from a fragment of the type) what he con- ceived to be the internal characters of the species (‘ British Paleo- zoic Fossils’ pl. ins, fig. 8a), but he omitted the columella, and showed the tabulz as concave—an idea probably due to the tangential nature of the sections, which missed the former, and incompletely displayed the character of the latter. ORIONASTREA PLACENTA (McCoy). 1849. Sarcinula placenta K. McCoy, Aun. & Mag. Nat. Hist. ser. 2, vol. iii, p. 124. 1851. Sarcinula placenta F. McCoy, ‘ British Paleozoic Fossils’ p. 110 & pl. 118, figs. 9, 9a, 9b. See also synonymy and notes on O. phillipsi. O. placenta differs from O. phillipsi solely in the absence of a columella, and consequently in the form of the tabule, which, deprived of their axial support, are concave instead of being conical. The major and the minor septa are almost equal in length, the former advancing very little farther beyond the theca than do the latter. O. placenta bears the same relationship to O. phillipsi as the non-columellate forms of Lithostrotion (Diphyphyllum and Stylastrea)! do to the columellate forms. At certain horizons (for example, at the base of the Great Limestone around Chollerford, South Northumberland) ‘ Diphy- phyllum’ not infrequently occurs to the exclusion of Lithostrotion locally. It is conceivable that similarly O. placenta may subse- quently be found, to the exclusion of O. phillips?, at some particular place or level. 1 See W. Lonsdale, in R. I. Murchison’s ‘Geology of Russia, &c.’ vol. i (1845) pp. 619 & 624: Diphyphyllwm refers to the fasciculate, and Stylastrea to the massive, variety. part 4] PHILLIPSASTR ZA HENNAHI, AND ORIONASTRZA, . 301 Notes upon the Holotype of the Species in the Sedg- wick Museum, Cambridge, No. 211, presented to the Collection by W. Hopkins. (PI XXIII, figs. 6 & 7.) The type-specimen of Sarcinula placenta McCoy is part of a depressed corallum measuring some 6 by 4 cm., and is about 2 cm. thick. Both the upper and the lower surfaces are re- markably flat. For the greater part, the coral interstices are free from mineral deposit, and consequently the material is very friable. Along certain planes, however, it is highly silicified, bemg then converted into thin bands of chert. Superficially, the fossil is stained a deep red. The theecze measure about 2 mm. in diameter, and are fairly regularly spaced, the interval between them averaging 4 to 5 mm. None of the septa invade the intrathecal region, and there is no columella. Im consequence of the absence of a columella, the tabulz are concave, but are somewhat irregular in habit. In this specimen it may be observed that the thecz are some- what smaller than in the average examples, and the number of septa correspondingly less—about 25. Attention must be drawn to the striking resemblance of this type to Martin’s figure of Hrismatolithus radiatus. In mode of preservation it would seem identical with the specimen illustrated in ‘ Petrificata Derbiensia.’ Unfortunately, since we know nothing of the intrathecal character of Martin’s coral, it is impossible to identify McCoy’s type with Martin’s figure, although the pre- sumption is that they are the same. ORIONASTRHA ENSIFER (Hdwards & Haime). 1851. Lithostrotion ensifer H. Milne Edwards & J. Haime, ‘ Polypiers Fossiles | des Terrains Paléozoiques’ p. 442. 1852. Lithostrotion ensifer H. Milne Edwards & J. Haime, ‘ Monograph of the British Fossil Corals’ Pal. Soc. p. 193 & pl. xxxvil, figs. 2, 2a. 1889. Phillipsastrea radiata (Martin), partim R. Schafer, Geol. Mag. dec. 3, vol. vi, pp. 401-407 & pl. x1, figs. 2, 5, ?6. 1903. Lithostrotion ensifer Kdwards & Haime, A. Vaughan, Proc. oe Nat. Soc. n. s. vol. x, p. 109. 1905. Lithostrotion ensifer Edwards & Haime, A. Vaughan, Q.J.G.S. ‘Fak Ixi, p. 199. The specimens upon which Edwards & Haime established the species came from Bristol, and were preserved in the British Museum. The identification of these is not possible; nevertheless, there are several specimens which were in that collection at the time when these authors were engaged upon their researches. The figures and descriptions given by them leave no room for doubt as to the identity of the species. They drew attention to its resemblance ue ‘ Phillipsastrea’ (that is, Orionastrea), and remarked that ‘in this fossil the columella is more prominent than in any other species of the same genus, and the walls much thinner.’ I cannot quite agree with the statement concerning the columella, although 302 DR. STANLEY SMITH ON AULINA ROTIFORMIs, ETC. { vol. 1xxu. J admit that the prominence of this axis is a characteristic feature of the species. The material examined by Schiafer included specimens of Orionastrea ensifer from Bristol, as well as of O. phillipsi and O. placenta from North Wales and Derbyshire. The Bristol specimen (No. 56740, British Museum), which he figured as ‘P. radiata, could be referred almost oe as well to O. phillipst as to O. ensifer; but I consider it to belong to the latter, rather than to the former species. O. ensifer is considerably less differentiated from the species of Lithostrotion than is the genotype O. phillipsi, and constitutes, as previously suggested, the passage-form between Lithostrotion and Orionastrea. Characters not described here may be taken to be in agreement with those of the genotype. External Characters. (Pl. XXIV, fig. 5.) The corallum has the same general form and characters as O. phillipst, except in regard to the distal surface; in this it differs very shghtly from Lithostrotion basaltiforme. The indi- vidual calices are clearly defined, and each is divisible into an intrathecal depression and an extrathecal platform. 'The latter is broad and sloping, and meets that of the contiguous corallites in a sharp ridge. In Lithostrotion this ridge is surmounted by a distinct wall of epitheca, and it is in the absence or weak development of this that O. ensifer is distinguished from the massive forms of Litho- strotion. Calices of the two genera are figured in juxtaposition for the purpose of comparison (Pl. XXIV, figs. 5 & 6). Internal Characters. (Pl. XXIV, figs. 3 & 4.) A thin epitheca may divide the corallites; but, if present at all, it is usually reduced to a palisading of isolated rods which in transverse section present a line of dots. The septa never attain a confluent condition. .The number of septa present is a fairly constant character: the number, including both series, closely approximates to 86. The major septa extend farther into the intra- thecal region than those of O. phillips generally do, and are much more uniform in their length. The minor septa, on the other hand, are but feebly developed. The columella is persistent and stoutly built, but this axis is not found swollen to the extent sometimes observed in O. phillipsi. 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