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- - THE TWENTY-SECOND
es:
ANNUAL REPORT
AND
pesTRact OF PROCEEDINGS
OF THE
BRIGHTON AND SUSSEX
NATURAL HISTORY SOCIETY,
ADOPTED AT A MEETING HELD
THURSDAY, SEPTEMBER 9th, 1875.
Price ONE SHILLING AND SIXPENCE. °
*
BRIGHTON :
_ PRINTED BY D. O'CONNOR, “GUARDIAN” OFFICE, 34, NORTH STREET.
1875.
THE TWENTY-SECOND
ANNUAL REPORT
AND
fresTRactT OF PROCEED! NGS
OF THE
BRIGHTON AND SUSSEX
NATURAL HISTORY SOCIETY,
ADOPTED AT A MEETING HELD
THURSDAY, SHPTEMBER OQth, 1875.
Price ONE SHILLING AND SIXPENCE.
BRIGHTON :
PRINTED BY D. O'CONNOR, ‘‘GUARDIAN” OFFICE, 34, NORTH STREET.
1875.
£,
President:
MR. J. DENNANT.
Wice-Presidents :
MR. HOLLIS, DR. HALL,
MR. BIGGE, MR. T. GLAISYER,
DR. HALLIFAX, MR. T. H. HENNAH,
MR. SIMONDS, MR. F. MERRIFIELD,
MR. PENLEY, MR. HASELWOOD,
MR. GWATKIN, MR. T. B. HORNE,
MR. W. E. C. NOURSE, MR. ALDERMAN COX.
SIR J. CORDY BURROWS,
Treaster :
MR. THOMAS GLAISYER,
12, North Street.
Bommittes :
MR. BENJAMIN LOMAX, MR. G. D. SAWYER,
MR. C. P. SMITH, MR. E. H. WILLETT,
MR. DENNET, MR J. WILLS.
Honorary Secretaries:
MR. T. W. WONFOR, MR. J. C. ONIONS,
38, Buckingham Place. 56, Middle Street.
Honorary Librarian and Burator:
MR. ROBERT GLAISYER,
The Dispensary, Queen’s Road.
At the Twenty-second Annual Meeting of the BricHToN anp
Sussex Naturat History Society, held in the Curator’s Room,
Free Library and Museum, Church Street, September 9th, 1875.
It was RESOLVED,—
That the Report, Abstract of Proceedings, and Treasurer’s
Account, now brought in, be received, adopted, and entered on the
minutes, and printed for circulation as usual.
That the cordial thanks of this Meeting be given to the Honorary
Secretaries, Honorary Treasurer, and Honorary Librarian, for their
labours in preparing the same.
That the following gentlemen be elected as Officers of the Society
for the ensuing year :—President: Mr. Dennant; Treasurer: Mr.
Thomas Glaisyer; Committee: Mr. Benjamin Lomax, Mr. C. P.
Smith, Mr, Dennet, Mr G. D. Sawyer, Mr Ernest H. Willett, and Mr.
J. Wills. Honorary Secretaries: Mr. T. W. Wonfor and Mr. J. C.
Onions; Honorary Librarian and Curator, Mr. R. Glaisyer.
That the sincere thanks of this Meeting be given to the Vice-
Presidents, Treasurer, Committee, Honorary Secretaries, and
Honorary Librarian for their services during the past year.
SPECIAL BUSINESS.
That the following words be inserted in Rule 14 after the word
“evening”: “Except in the months of July and August, during
‘which no meetings will be held;” and that in Rules 4 and 16 the
word “October” be substituted for the word “ September.”
That the Committee appoint sub-Committees to undertake the
several branches indicated in Mr. Wonfor’s paper, read at the
Ordinary Meeting on the 12th of August last.
(Signed) J. EK. HASELWOOD,
Chairman.
It was ALso REsoLveD,—
That the warmest thanks of this Meeting be presented to Mr.
Ald, Cox for his able conduct as President during the past year.
REPORT.
In presenting the Twenty-second Annual Report your Committee
have the pleasure of recording the continued prosperity of the
Society, which has considerably increased in number during the
past year. They regret the loss which the Society has sustained
by the death of two of its Members, viz., Mr. J. Howell and Mr.
G. Scott, the latter one of the Vice-Presidents and Honorary
Librarian, and who had for many years taken an active part in
promoting the prosperity of the Society, and rendered valuable aid
in obtaining the accommodation for the Society at the Free Library
and Museum.
The state of the finances is very satisfactory. The sum of
42 10s. remains in the hands of the Treasurer, after expending
£29 4s. 2d. in the purchase of new books and periodicals.
The following books have been presented to the Society
during the year:— Eleventh Annual Report of the Belfast
Naturalists’ Field Club and Guide to Belfast (from the Society) ;
Report and Proceedings of Eastbourne Natural History Society
(from the Society) ; Journal of the Quekett Club (from the Society) ;
~
[3 |
Annual Report and Proceedings of Maidstone and Mid-Kent
Natural History, Microscopical, and Philosophical Society for 1875
(from the Society); Report of Lewes and East Sussex Natural
History Society (from the Society); Quarterly Reports of Sub-
Wealden Exploration (from the Secretary) ; Remarkable Forms of
Animal Life on the Norwegian Coast, by G. O. Sars (from the
Author) ; Sur les Brachiopodes Tertiares de Belgique, by T. David-
son, F.R.S., F.G.S. (from the Author); on the Trimerellidz, by T.
Davidson, F.R.S., F.G.S., and W. King (from the Authors) ;
Geological Magazine (from Mr. H. Willett); The Glaciation of
the Southern Part of the Lake District, and the Presidential
Address before the Keswick Literary and Scientific Society, by J.
Clifton Ward, F.G.S. (from the Author) ; On the Cephalopoda Bed
and Oolite Sands at Dorset and Somerset, by J. Buckman, F.G.S.
(from the Author) ; Flora of Eastbourne, by F. C. S. Roper, ris.
F.G.S. (from the Author) ; Proceedings of Geologists’ Association
(from the Society). From the Smithsonian Institution, Washington,
U.S.A.: Report of the Society for 1873; Daily Bulletin of
Weather Reports for October and November, 1872; Birds of the
North West; Geological and Geographical Survey of Colorado ;
Geological Survey of Missouri, with Atlas ; List of Elevations West
of the Mississippi River; Cretaceous Flora of the Territories ;
Catalogue of the Publications of the United States Geological
Survey of the Territories.
The following books have been purchased during the year :—
Medical Botany, Woodville ; Missionary Travels and Researches,
Livingstone ; Last Journals of Livingstone, by H. Waller, F.G:S. ;
Contributions to Solar Physics, Lockyer; Text Book of Botany,
Li a
Sachs ; Corals and Coral Islands, J. D. Dana; Log Book of a
Fisherman and Geologist, F. Buckland; Insectivorous Plants,
Darwin ; Memoirs of Sir R. I. Murchison, Geikie; Climate and
Time, Croll; Our Summer Migrants, Harting ; Harvesting Ants
and Trap-door Spiders, Appendix, Moggeridge ; Mineralogy and
Appendix, Dana; Cave Hunting, Boyd Dawkins; On the Microscope,
Carpenter ; Scottish Cryptogamic Flora, Greville ; Episodes of
Insect Life ; Dictionary of Science, Brande and Cox; Heart of
Africa, Schweinfurth.
Periodicals: Entomologist; Entomologists’ Monthly Magazine ;
Geological Magazine; Grevillea; Linnean Society, Transactions
of; Monthly Microscopical Journal; Popular Science Review ;
Quarterly Journal of Microscopical Science ; Quarterly Journal of
Science ; Science Gossip ; Zoologist.
The number of volumes in the Library is 858, exclusive of
pamphlets and unbound current periodicals, and your Committee
are pleased to learn that the circulation of books continues to
increase.
A numbered Catalogue of the books belonging to the Society
can be consulted at all times at the Library counter of the Free
Library and Museum, and Members can obtain the Society’s books
on loan on application to the Assistant-Librarian during the hours
the Institution is open.
Your Committee are pleased to hear that the public have
made great use of the Society’s books during the past year, and that
~ the books have not received any damage at the hands of the
public.
ae
During the year the Society’s Microscopical Cabinet, which
now contains 257 slides, has received donations of slides from the
following gentlemen: Mr. T. Curteis and Mr. T. W. Wonfor.
The Conversazione held in the month of February, 25th,’ at
the Royal Pavilion, was very successful, both in the number of
those who were present and in the variety and character of the
entertainment. A detailed account will be found in the pro-
ceedings.
Scarcely any response having been made to the circular
inviting assistance in verifying the fauna and flora of Sussex, as
suggested in Mr. Wonfor’s paper of April roth, 1873, that gentleman
read, on August 12th, this year, a second paper, in which he
suggested a scheme by which the Society might accomplish that
and other objects, and which your Committee hope will meet with
your approval.
The thanks of the Society are due to those gentlemen who
have read papers, exhibited objects, or presented slides for the
cabinet, photographs and drawings for the Society’s album, or
specimens.
The Society is to be congratulated on the increased attendance
at its meetings, the far greater accommodation afforded at the Free
Library and Museum, and the growing interest taken by the
Members in the welfare of the Society.
The Field Excursions since the last report have been as
follows :—September, 1874, Cissbury ; October, Falmer; May, 1875,
[9]
Balcombe ; June, West Grinstead (postponed in consequence of
weather) ; July, Newhaven ; August, West Grinstead.
The Annual Excursion took place on the znd August to
Battle Abbey, and the Sub-Wealden Exploration at Netherfield.
In concluding their Report, your Committee beg to request
the Members to endeavour to promote the prosperity of the
Society by bringing its merits under the notice of their friends, by
contributions of works of Natural History to the Library, of
photographs or drawings of objects of Natural History for the
: album, and of slides for the Microscopical Cabinet, and particularly
Sh
by Reading Papers during the ensuing year.
mootmact ‘OF PROCEEDINGS:
18'74.-5.
SEPTEMBER toth.
ANNUAL MEETING.
The retiring President, Mr. J. H. HASELWooD, before vacating
the chair, said that as it was officially his “last dying speech and
confession,” he asked for their sympathy. First of all, he thanked
very cordially the whole of the Members for the loyalty they had
on all occasions manifested towards him, and for their attendance
at the various meetings. During the year they had shifted from
their old place of meeting into more comfortable quarters, and
this, probably, had attracted more to their meetings.
Future years, he hoped, would find the number still increasing,
for there could be no doubt that it was much more instructive and
enjoyable when the attendances were large than when small.
With regard to the Committee, he was glad to be able to say
that he had never been associated with a body of men between
whom there existed a more kindly feeling. And this was equally
the case with the Secretaries, from whom he had received the
greatest assistance, and who worked together most amicably, their
departments, the scientific and the business, never clashing the one
with the other, or causing any jealousies to arise between them.
“Thee
[il]
While speaking of the Secretaries, there was one point to which he
should like to allude—the statement which was sometimes made
(though it had never been made to him) as to the important and
continuous part played by Mr. Wonfor in the affairs of the Society.
It was said there was too much of Mr. Wonfor; and so, perhaps,
there was in one sense. But this was the fault of the Members
themselves; if they came forward and did their duty better than
they had yet done, there would be no occasion for him to act as a
stop-gap so often as he had to.
Knowing Mr. Wonfor, he was in a position to say that no man
had less desire to monopolise the time of the meetings than he; he
would only be too glad if the other Members came forward and took
their share in the discussions, without falling back so much upon
him. His position, however, was one which must always be
recognised, for the greater portion of the burden of the Society
fell upon his shoulders.
Those Members who, during his period of office, had con-
tributed papers, he now took the opportunity of thanking; and he
was pleased to notice that their efforts had not been confined to the
comparatively small sphere embraced by the Society, but that,
through the medium of the Press, they had been made known to
the public, and had, he believed, in many cases, been extensively
read.
With the Microscopical Members he was, perhaps, more at
home than the others, and he would here hint to them the advisa-
bility of coming out with their instruments more than they did at
the ordinary meetings. To some, perhaps, it might appear that
the results obtained from the Society were not so great as they
might be. But it should always be remembered that the majority
of the Members, being engaged in professions or business, had not
the necessary time at their command to devote to original
research; all they were able to do was to make themselves in some
measure acquainted with the labours of those who devoted the
whole of their time to the advancement of particular studies. And
in doing this they felt all the better, the habits which they thus
acquired proving to be most useful in other walks of life; they
were led to be studious, careful, and exact, and the possession of
these qualities could not but be productive of good results,
[12]
But, it might be said, what had he for them in the shape
of suggestions for the future. He had two suggestions. One was
that papers should occasionally be read, dealing with the elementary
aspect of the subjects of which they treat. Many of them, he was
sure, oftentimes failed to appreciate some of the papers as much as
they would like for the want of elementary knowledge respecting
the particular questions referred to.
The other suggestion was that papers should also be read
occasionally explaining the cant scientific terms now coming into
use; such, for instance as “ The conservation of energy,” “The
correlation of physical forces,” “ Natural selection,” “The survival
of the fittest,’ and ‘ Evolution.” Very frequently these terms
were used as bugbears, for they were often used by persons in a
very random and careless manner, showing that they were really
ignorant of their true meaning. Let him, however, not be mis-
understood ; he did not wish to make these subjects of controversy
—that would be unwise, and would sure to be productive of evil.
On the other hand, papers explaining these terms, and showing in
what sense they were used and understood by the master minds of
the day would, he was quite convinced, be useful to them. Though
the sphere of the society was small, he believed it had been, and
would still continue to be, very useful to the town, helping to elevate
some, at least, of its inhabitants into something higher than they
would have been but for its influence.
It was with great pleasure that he vacated the chair in favour
of such a man as Alderman Cox—one of the ex-Mayors of
Brighton—who, he was sure, would preside over their gatherings
in a manner which he had never been able to attain to.
The President, Mr. Alderman Cox, in taking the chair, said it
was not usual, he was told, for the incoming President to make any
remarks on first taking the chair; but he would take this oppor-
tunity of thanking them for the honour they had done him, and of
saying that if he succeeded in performing the duties of his office as
well as the President who had just retired from the cbair, he should
consider that he had performed them well.
Mr. M. PENtey said he had been asked to propose that a warm
vote of thanks be given to Mr. Haselwood for his able conduct as
ie
[ 13 ]
President during the past year. He did so with the greatest
possible pleasure. He had not been able to attend many of the
meetings, but those that he had, had shown him how well qualified
the ex-President was for the post he had held.
Mr. G. Scorr seconded, observing that he had regarded Mr.
Haselwood’s performance of his duties with the utmost admiration ;
but this was only another instance of the manner in which the
Society got through its labours under the presidency of the most
varied men.
The meeting then resolved itself into an ordinary meeting for
the exhibition of specimens.
Mr. G. Scorr showed some interesting specimens which were
about to be added to the Museum, including a March ichneumon
(Egypt), an Alpine marmot (North Europe), a Coati mondi (South
America), &c.; several rude flint implements which had recently been
discovered in a new district—Pyecombe; and a ray from one of the
fins of the gigantic sunfish recently caught by Mr. Lawler, of the
Brighton Aquarium, off the Sussex Coast. Mr. Scott also exhibited
arough drawing of the fish, and explained its general characteristics.
The fish was supposed to have been the largest caught in British
waters. It weighed 12 cwt. Mr. Scott also showed a spine of the
fish, measuring about 2% feet long.
Mr. F. E. Sawyer showed a specimen of the arundo arenaria,
obtained from the dunes in Holland, where it binds together the
loose sand. Mr. Dennet made some remarks upon the subject,
in the course of which he suggested the use of the pinus maritimus
as a shrub suitable for sea-side gardening.
Mr. C. Cayton showed a good specimen of the jaws and
puddle of an ichthyosawrus, brought from the blue lias formation
at Street, Somersetshire.
Mr. Wonror exhibited some Indian mosquitoes, as well as a
small case vf moths,—the Vapourer O. Antiqua,—the inmates of
which had formed the subjects of a sevies of experiments on the
attraction furnished to the males by the females of the species,
A long discussion followed this exhibition, the question being
as to what particular sense enabled the males to discover the
females when long distances intervened between them.
[ 4 ]
SEPTEMBER 24TH.
MICROSCOPICAL MEETING.—“ POND LIFE.”
Mr. T. W. Wonror announced the receipt from Mr. Curties,
of High Holborn, London,—who is an honorary member,—of
a photograph of a group of thirty-five separate specimens
of British microscopical animals and plants, collected from a
pond at Leytonshire, near London, and drawn from life by Mr.
H. C. Richter. The objects, which had been very beautifully
grouped, were inspected with much interest; and the diagram,
with the accompanying key, rendered the photographs more
valuable. In the absence of Mr. Lomax, who had suggested the
subject of the evening, but who was unfortunately prevented
attending.
The President, Mr. Alderman Cox, asked Mr. Wonfor to in-
troduce the subject for the evening.
Mr. Wonror remarked that he had lately experienced the
truth of an observation made at the last meeting that, although
they might propose to show living objects, the difficulty would be
to get them. Only the evening before he had some hundreds of
specimens of the volvox globator, and he had purposed exhibiting
the rolling, turning, volvox; but he was unable to do so as they had
now disappeared.
A few bottles of specimens from ponds at Lewes and Furze
hill, would, however, be submitted for examination ; and he had also
managed to secure some larve of the English gnat,—in other
words, of a mosquito,—which would doubtless be looked upon with
interest, remembering the discussion which took place at a previous
meeting. The specimens of pond life would be likewise interest-
ing, because of the anxiety manifested at times to know whether
or not certain water was drinkable. Now, the mere fact of finding
certain forms of animal life in water would not necessarily mili-
tate against it for drinking purposes; but if he found in stagnant
or well water certain other forms of animal life, he should begin to
inquire what it was that had filtrated into it.
[ 15 ]
There was before them a sample of water, taken from a well,
to which, for drinking purposes, he should greatly object. There
were in it forms of life of the kind invariably found in water into
which some organic substance, or, in other words, sewage matter,
was filtrating. That being so, and if sewage matter were there,
he should not be surprised if, at some time or another, infectious
disease broke out in the neighbourhood in which the water was
consumed.
Certain forms of animal life were always to be found where
the water was contaminated with sewage matter; in such cases
there was always danger; and, consequently, important lessons
were always to be drawn from examining the forms which the
water contained. In reference to the water supplied by the Cor-
poration of Brighton, there were circumstances connected with it
which rendered it virtually impossible for it to contain organic life.
It was brought up at such a depth from the chalk that, if examined
microscopically when taken from the main, not a trace of organic
life could be observed in it.
The PRESIDENT: Yes; and do not forget the fact that all the
reservoirs have been covered over.
Mr. Wonror: And, besides, it remained such a short time
in the reservoirs after it had been brought up from the chalk.
Examination had proved that the Brighton water was invariably
free from any trace of organic impurity.
The PRESIDENT: Then you prefer the constant service to
intermittent ?
Mr. WonFor replied that he had the constant service at his
own house; and, in continuing, said if all could have their drinking
water drawn direct from the main they would avoid the risk of
having animal life introduced, by water standing in a cistern exposed
more or less to the atmosphere. There were, however, certain
forms of life in pure water which rendered it drinkable. For
instance, the American weed, anacharis, having been thrown into
water at Ditchling, never before drinkable, that water afterwards
became pure and palatable, and was eagerly sought by those
persons living in the neighbourhood.
[ 16 ]
Dr. HaLLirax said it was acknowledged as a general principle
that the growth of vegetables purified the water; and this was
found to be the case even in tanks or globes, where the purity of
either fresh or sea water was preserved by the presence of weeds.
The rapidity of growth of the anacharis weed was then dilated
upon.
Mr. C. F. DENNET enquired whether there was no mineral
substance pumped up with the Brighton water ?
Mr. Wonror replied that there was a quantity of chalk in
solution—he believed about fifteen grains to the gallon.
Mr. GLAISYER said the proportion was about seven grains to
the gallon, as supplied by the Corporation.
Mr. Wonror observed that boiling would to a great extent get
rid of the chalk; or, if the Brighton Corporation would adopt the
plan which used to be a patent, but which was now public property,
the quantity would be reduced to about three grains, the water
would lose what was commonly called its hardness, there would be
a wonderful saving to the ratepayers in soap, and to the house-
holders in connection with their boilers and kitchen ranges.
Mr. DenNET handed in a newspaper paragraph which stated
that Dr. Letheby had come to the conclusion that moderately hard
water was safer and more healthy for drinking than soft water.
Mr. Wonror remarked that, as a matter of fact, absolutely
pure water was not good for drinking.
The meeting then resolved itself into a conversazione for the
exhibition and inspection of the various objects of pond life
which had been brought by Dr. Hallifax, Mr. Wonfor, Mr. J. E.
Haselwood, Mr. T. Glaisyer, Mr. R. Glaisyer, and Mr. G. Nash.
Among the most noticeable were the rotifera exhibited by Dr.
Hallifax.
Mr. Wonror specially drew the attention of the meeting to a
specimen of the Daphnia which was covered with Vorticellae—small
bell-like animals.
The larve of the English gnat, and specimens of impure well
water containing Bacteria, were also shown.
Lea
OcroBER 8th.
ORDINARY MEETING.—MR. BENJAMIN LOMAX ON
“FACTS IN THE CHEMISTRY OF PLANTS.”
Although the chemical changes which the life history of every
plant involved were discussed in most scientific works, and the
number of organic compounds directly obtained from a vegetable
source was recognised by all conversant with our manufactures,
there remained an amount of chemical effect wrought on surround-
ing nature by the agency of plants which altogether escaped
popular notice, and was apt to be under estimated even by the
naturalist. He must leave to those who had better opportunities
than himself the task of adding to their knowledge on this point,
contenting himself with a reference to some facts with which all
were acquainted, but few thought of in this connection, and
trusting to the after discussion for further information. The
evolution of oxygen by plants had been so exhaustively considered
in botanical works that he should merely draw attention to the
amount of work it actually represented. Every human being gave
off, on an average, 154lbs. weight of pure carbon per annum.
Assuming the population of the world at 20,000,000,000, the
wonderful amount of 137 millions and a half of tons of carbon
was expired from human lungs alone in one year. Add to this
the carbon evolved from animals, large, small, and microscopic,
the products of combustion, and the decomposition of organic
matter, and some idea might be formed of the work which plants
had to perform; and which they did so well that the amount of
carbonic acid in the atmosphere was seldom, if ever, perceptibly
increased. i
The deposition of silica on the exterior surface of the culm in
the cereals, sedges, and bamboo, might be, perhaps, regarded as
necessary for the stability of the plant, but the geology of certain
districts must be considerably modified by the change this forma-
tion indicated. Silica was usually found in a coarse, crystalline
Cc
[ 18 J
form, even the typical grain of sand having a very perceptible size
and shape, whereas the siliceous coating of a wheaten straw was
deposited in sufficient quantity to turn the edge of a knife, with
a uniformity and a fineness of division resembling that produced
by the electrotype process. Whether this silica was extracted
from sand, as indicated by the constant presence of reeds and
sedges on the sea shore, or separated from the soluble silicates
everywhere present in arable soil, a curious chemical process must
have taken place, which there was no means of tracing, and the
question was not simplified but rather increased by the fact that
the deposit was not always a powder. Frequently it was crystalline
or needle-shaped, sometimes laminated, as at the nodes of bamboo,
from whence it was collected in considerable quantity under the
name of tabasheer.
Then, again, the importance of the change might be estimated
by the scale on which depositien took place. The horsetails and
reeds yielded in their ashes 97 per cent. of pure silica, the grasses
nearly as much. The diatoms were almost made of silica, and
when it was remembered that the deposit remained unchanged by
burning or natural decomposition, it would readily be conceived
that in this way also plants performed a most important work in
nature. The silica so deposited was puve, and the purification of
salts was one of the most remarkable functions of plants. In the
laboratory, purification was generally performed by crystallising
from a solution, and it was always assumed that a crystal posses-
sing the proper form and colour was a pure specimen of the salt
under examination.
In like manner plants produced various salts in purity, as
those knew who were dicussing the beautiful crystals called
raphides, found in the cells of almost all plants, but especially in
the liliaceee and kindred orders, with the crystals on the leaves of
the deutzia, &c. In this pure form were found the carbonate,
oxalate, phosphate, and sulphate of calcium, and those who weve
curious in the various shapes which crystals of the same compound
assumed, might find fresh field for investigation in the fact that all
the varieties of form common to the bodies mentioned were found
in different plants, but that the same species always presented the
same form and composition of crystal. Whether raphides were
{ 19 ]
useful to plants or not, there could be little doubt that the continual
separation, on so large a scale, of salts in their purest form, must
have a very marked influence on the economy of nature.
Another strange function of plants was the suspension of
odours, disagreeable or otherwise. A peculiarity which, in a less
degree, was shared by other members of the goosefoot family, gave
its specific name to the chenopodiwm vulvaria. Its leaves, when
rubbed, imparted the disgusting odour of putrid salt fish, and
superficial observers had regarded this plant as a contrivance for
creating and diffusing a stench. But this was hardly the case. The
stench was, he believed, antecedent to the plant. The chenopodiwm
was found on the sea shore amongst decaying seaweed, or in heaps
of rubbish, whence nasty smells might reasonably be expected to
issue. The odour, which might otherwise have disgusted or
poisoned the passer by, remained unnoticed when absorbed by the
leaves, and it was not the fault of the goosefoot if we chose to
handle and smell our fingers afterwards. During the exhibition of
the local flora at the Brighton meeting of the British Association,
one of these plants was kept alive for several days, and afterwards
died in the same spot, and he could bear witness that no offensive
smell was ever perceived, except by those who touched. The whole
question of plant odours was an interesting one, as there seemed
no rule or uniformity in them. Thus the Viola odorata and
Viola hirsuta, the Reseda odorata and Reseda lutea, plants so
similar as frequently to be mistaken for each other, differ essen-
tially in this particular. The common Diplotaxis had a flower
scented like the field convolvulus, while its leaves were simply
disgusting. The Silene inflata had the taste and smell of young peas,
while the Poterium sanguisorba smelt of cucumbers, and the Lychnis
vespertina had a sweet odour in the twilight, and none by day.
Another peculiarity in plant economy was the power of
emitting light. A common plant of Tasmania which had the form
of a lettuce, and the pale, leathery appearance like that of a fungus,
or liverwort, glowed at night with a phosphoric luminosity from
every part of its surface. The children cut its leaves into fantastic
forms and hung them in dark rooms. Though this plant was very
abundant, he was unacquainted with its generic name, and did not
know whether to assign it to the Phenogamia or Cryptogamia.
[ 20 ]
The light was bluish, like that of the English glowworm. The Aus-
tralian glowworm shone with a copper coloured radiance. Similar
phenomena had been regarded by Linnzus and others, as indica-
ting an absorption, and subsequent radiation of diffused light, he
might, without committing myself to this theory, point out a
direction in which there seemed room for profitable investigation.
There could be no doubt that light was largely utilized in vegetable
economy. In the absence of light the decomposition of carbonic
acid ceased. The juices of celery and endive changed their
character, and colour was no longer produced. It appeared to him
that we were justified in supposing that these functions were
entirely discharged by light itself, and that the office of the plant
was to expose the tissues under favourable circumstances. If this
were so, an investigation of those conditions ought to enable us to
make light do the same for us. As light produced all shades of
colour in plants, a naturally coloured photograph was a possible
production, and might be ultimately obtained, and as the character
of certain juices was entirely changed by light, we might find out
from plants the secret by which organic compounds, similar in
composition, but different in properties and appearance, might be
interchanged. It would be useless to refer to the countless organic
compounds produced directly by plant organism; to starch,
dextrine, mannite, gum, &c., &c. It was sufficient to remark that
these substances were produced without any visible mechanism,
and under circumstances apparently exactly similar, to show that
in the simplest plant was locked up a secret of chemical manipula-
tion to which at present we had not the slightest clue—but it was
within the scope of his present object to point out as a great
portion of plant work the conversion of gases into solid matter.
Those who had grown mustard and cress upon a piece of wet
flannel knew that a certain quantity of mould was produced. So,
on a larger scale, every plant took its substance from the
atmosphere and laid it finally on the soil, building up daily what
would be cainozoic strata when we ourselves were fossils. The whole
of our linen and canvas was thus woven out of thin air, for the flax
plant, after the thread fibres had been removed from it, presented
the same analysis as the soil from which it was taken.
At the request of the CuarrMAN (Mr. Haselwood), a vote of
thanks was awarded to Mr, Lomax for his instructive paper.
f a J
Mr. Wonror remarked that, although they could not well
separate chemical from physiological or life action, cases cropped
up in which chemical action was seen to be produced without,
apparently, the intervention of life. There was another curious
fact, namely, that while on the one hand there was such a variety
of colour produced by the action of light, the brightest sea weed
was found in the deepest water, and in the darkest caves.
Mr. Dowsett added, the bright sea weed could not be grown
in aquaria, unless the light reached it through a tinted medium.
The CHAIRMAN (Mr. Haselwood) thought this would form an
argument to show tbat colour was not entirely due to light.
Mr. DowseEtt mentioned another strange circumstance; that
lemon, turpentine, and bergamot were chemically the same, yet
emitted different odours. What produced the odours was unknown.
Mr. Dennant directed attention to one practical outcome,
namely, the importance of instructing children in the elements of
science. If, he said, the poor children of large towns knew ever so
little of the importance of maintaining a balance between animal
and vegetable life, their health would be much better than it was.
If they would foster and nurture plants in their neighbourhood,
they would be much happier and stronger than they were without
any kind of foliage.
OcToBER’ 22nd.
MICROSCOPICAL MEETING. — “SECTIONS AND.
SECTION CUTTERS,” BY DR. HALLIFAX,
The CHarrmMan (Mr. J. E. Haselwood) read a letter from Mr.
T. W. Wonfor, who had announced the subject, apologising for his
absence through illness, and called upon Dr, Hallifax, who had
consented to fill up the gap.
Dr. Hatuirax said he much regretted the absence of Mr.
Wonfor, who would have enlightened them upon the subject much
more than he could. His object would be to give them a practical
[ 22 ]
illustration—practical as far as possible—of cuttings for micro-
scopes, and of all the instruments which they had at command, as
well as the mode in which they were employed.
The first and most valuable cutting apparatus which he should
bring before them was the instrument invented by Richard Beck,
which possessed a very fine screw, having 43 turns to the inch, so
that one revolution would lift it a 43rd part of an inch, and a fraction
of a turn would obviously diminish the distance accordingly. The
large head of the screw was so devised that it was divided into
25 parts, so that one was enabled to make a cutting by this
machine of a thousandth part of an inch theoretically, and 500
parts of an inch practically. The plate, too, was remarkably true.
The instrument was invented for the purpose of cutting
wood and sections of vegetable substances of a certain degree of
hardness with a chisel. If, however, the subject were soft, a razor
must be used, and for specimens of this nature he could not say
Beck’s invention was so valuable as it was for hard substances.
He had fixed upon the surface of the machinea piece of plate glass,
—the truest that he could procure,—to avoid friction and to allow
the instrument used for cutting to pass over it more readily, an
idea which would be found to be as correct in practice as it
was in theory. There was another machine sold by Baker, of
Holborn. The operation of the screw was the same, but it was not
finished in the same neat and exact manner, so that it was
impossible to get the same refinement in the sections produced by
it. Mr. Haselwood, he believed, had also brought one of the most
recent inventions, which had the advantage of a refrigerating
apparatus.
Mr. HaseELWwoop explained a section cutting machine known
as Professor Rutherford’s. This was provided with a refrigerator,
which, being filled with ice and salt, solidified the wax or other
substance in which the object cut was “ mounted.” The screw was
a very fine one and was furnished with a spring, which would fix it
at any part of its turnings. There was a difficulty of obtaining a
proper section of very soft substances, such as animal tissue, but
he believed the Germans were able to cut a section clean through a
man’s brain, a very difficult operation.
{ 23 ]
Dr. HALLIFAX said he believed the Germans possessed the
secret of the meaus of obtaining such a section, which they did not
promulgate to the world. The Germans cut such sections for sale,
and did it on a very large scale.
Dr. HALLIFAX next showed the method of preparing soft objects
for cutting, illustrating his remarks by producing a dead wasp,
ulready attached to the cylinder by a mixture of gum and creosote,
which he recommended as a preservative of like specimens, as the
creosote remedied the brittleness of the gum. Wrapping a strip
of paper round the cylinder so that the top edge of the paper was
about on a level with the wasp, he poured melted white wax into this
yeadily made capsule, completely submerging the body of the
insect. When cooled, the wax held the body firmly, a cutting of
the soft body being thus safely made. He also showed another
similar apparatus, the “well” of which, however, was surrounded
by a glass plate, the smoothness of which obviated, as far as
possible, friction with the cutting instrument, which he advised
should be moistened with methylated spirit. He mentioned that
another mode of cutting was to place the object in the hand,
and there was, too, a knife with two blades for cutting purposes,
having a screw by which the two blades were brought together.
Mr. Hasetwoop pointed out that there was a newer invention
—a knife with three blades, so that two sections, showing the con-
tiguity of the parts, might be cut at the same time.
Dr. Hatirrax then said that, in order to procure an accurate
specimen of a minute section, it was necessary to have one side of
the razor ground perfectly flat so that it might come in direct
opposition with the object, and not scoop the object and thus
destroy the specimen.
Mr. Hasetwoop observed that it had been recommended
that the ls army razor should be used, on account of its thin
blade.
It was suggested by a member that the Plantagenet razor was
very useful for such a purpose.
Mr. Hasetwoop further informed the meeting that, with the
use of a new medical instrument, and the ether spray, medical
[ 24 J
men were enabled to take from the body pieces of diseased skin,
so that they might study the various forms of skin disease.
Dr. HALLIrax next exhibited a saucer arranged to facilitate
the placing of cuttings on slides. Very thin sections, if placed
in water, and the slides placed under them and then raised, rarely
retained the desired position. To obviate this he had a small hole
drilled in the bottom of the saucer, which was tightly fitted with a
slip of wood. After the object had been floated into its right
position, the plug was removed, and the water, running gently
away, left the section in its proper place. The drilled hole should
not, however, be too large, otherwise a current would be formed
and destroy the effect of its operation.
At the conclusion of the meeting, the Chairman announced
that the next microscopical meeting was the last of the series this
year, and it was hoped that they would have a microscopical
conversazione, and that members would bring with them the
choicest specimens of their collection.
NOVEMBER 12th.
ORDINARY MEETING.-—“ ON RECENT EXCAVATIONS
AT CISSBURY,” BY MR. ERNEST WILLETT.
Mr. Wonfor had informed them that excavations had been made
at Cissbury during the past summer: indeed, several of the members
had themselves visited the spot; and, on the understanding that a
more detailed account would not be unacceptable to the Brighton
and Sussex Natural History Society, he had arranged a few notes
on the subject into the form of a paper, for the defects and short-
comings of which he must ask their indulgence. Had the site of
the work been other than at Cissbury, it would have been desirable
to have preceded the paper with a description of the locality and
situation of the earthwork, but the spot must be so familiar to
every Sussex man who was interested in ethnology, and the camp
had been so frequently described that, in the absence of fresh
information bearing on the subject, it would be inexpedient to
recapitulate minutely the accounts given by the various archwo-
logical authorities who had treated on the subject. The etymology
of the name had, too, been a wide field for speculation. Camden
in the 16th century, Cartwright, Turner, M. A. Lower in this, had
discussed the question at length, but the result of their several and
combined efforts was, he was afraid, eminently unsatisfactory.
They all attributed it to the “bury ” (or fort) of a Saxon General,
of whose occupation of the camp, or presence within its walls, from
internally derived evidence, we had no proof whatever. He could
not accept the theory, in the absence of all traces possible of
Saxon tenancy; it might be supposed that had the hill been
occupied by a military chief (sufficiently powerful and well known
to give his name to the place) some relics would have been found
in the numerous excavations that had been carried on in and about
Cissbury, of so highly a civilized people as the Anglo-Saxons.
This was the case with the Romans, who, though they were by far
more advanced in the culture of arts and manufactures and
manners of living than our German forefathers, were but tempo-
rarily established in this country, of them we had sufficient
evidence in bones, pottery, coins, &c., to believe that, if indeed they
were not the actual engineers of the Camp, they selected it as an
important post, and held it for some time. With this brief intro-
duction he would proceed at once to the subject more properly
before them, hoping, though far from being sanguine, that future
excavations might reveal a Saxon burial ground, or that a deriva-
tion supported by more patent external evidence might be found
by some one in an unlooked for source. He proposed to divide
his paper into three parts :—
I. To give a description of the pit, the galleries connected
with it, and the material with which they were filled in.
II. To state the reason for which it appeared the pit and
galleries were made, and to describe the various manufactured
articles found, their occurrence and probable uses.
III. To compare the workings with those of similar character
in other parts of the country, and to shortly consider the question
as to who the people were who originated them.
It was difficult, even if it were desirable, to prevent these three
[ 26 ]
questions overlapping and running one into the other, as they were
so intimately connected one with the other.
The pit was rather at the south extremity of the series of
basin-shaped hollows which occupied the south-west corner of the
camp. He had no especial reason for selecting this one in par-
ticular, except that it appeared not to have been previously
disturbed ; it was of an average size, and the depression from the
natural slope of the hill was very slight, only about 16 degrees.
The result of the excavation showed:—First, surface soil,
similar to that in all on this side of the camp, containing chipped
flints, flakes, and broken implements, land and oyster shells,
numerous water-worn pebbles, a few pieces of bone, and fragments
of a coarse pottery. Below this, to a total depth of five feet, came
a layer of small chalk, rubble, and loam, of a yellowish colour, con-
taining a few flint implements, one or two fragments of bone, and
a deposit of charcoal surrounded by calcined chalk. This stratum
seemed to extend beyond the area of the shaft in north, south, and
westerly directions, and was the layer from which Col. Lane Fox,
in his extensive excavations in 1872, obtained the greater number
of the flint implements on which he founded his able treatise, read
before the Society of Antiquaries. Dead land shells also occurred
in this layer, of which by far the greater portion were Cyclostoma
Elegans ; but specimens of Helix nemoralis, and one of the rare
shell H. obvoluta (now almost extinct in Sussex), were also found.
At five feet (the depth at which the solid chalk at the edge of the
pit commenced) the filling-in entirely altered, being composed of a
moist red earth, full of flints worked and unworked, numberless
chippings, a fragment or two of stags’ horns, and several patches
of charcoal. This feruginous deposit was much thicker in the
centre than at the sides; it seemed much trodden and worn, the
implements were mostly huddled together at the centre, but
unworked or merely broken flints occurred all through the layer.
At 10 feet it was replaced by large blocks of pure chalk (in
some cases roughly squared) loosely thrown in, the interstices not
filled up by looser or smaller material. No change was observable
for about three feet, but here, viz., at 15 feet, came a repetition of
the red earth above mentioned, with the same characteristics, but
not so thick as the one above, two feet being the greatest depth at
[ a7 J
the centre and thinning out to six inches towards the circum-
ference, and this did not extend to the sides by 18in. The
remainder of the debris, ‘.e., from 16 feet to the bottom, was pure
chalk, but it differed from the layer above in that the blocks,
though loosely thrown in, had their interstices filled with small
chalk, rubble and loam. Throughout this layer were distributed
the implements, which would hereafter be more fully described,
and with which he concluded the original sinking was done. They
consisted for the most part of the broken pieces of the antlers of the
red deer, showing more or less marks of usage and manufacture,
- four scapulz (shoulder blades) of bos longifrons, and one of the
common pig, one or two flint implements, and a few broken flints.
The thickness of the last deposit made up a total of 20ft. from the
ground, and which was tabulated as follows :—
Surface soil 2ft., chalk rubble 3ft., red earth, &c., 5ft., chalk
blocks 3ft., red earth, &c., 3ft., chalk blocks 4ft., total 20ft. Ata depth
of 17£t. from the surface in the north-west corner he, not altogether
unexpectedly, came upon the mouth of a cave. He said not
altogether unexpectedly, because he had previously seen similar
caves in another pit in the camp, though on a much smaller scale.
And as the workmen proceeded, one by one were displayed to
view the series of eight caves, which constituted the chief interest
in their work. These caves, as could be seen by the ground plan,
yan laterally in all directions, and were filled before the excavation
to within from 1ft. to 2ft. from the top with loose blocks of
chalk partly converted into a kind of stalagmite by being
cemented together by carbonate of lime, derived from the percola-
tion of rainwater through the chalk above; it was exceedingly.
hard, and required a considerable blow to detach the pieces which
jt was necessary to remove, but this hardness was not so observable
at the entrances, and from the mouth, running in a few feet, there
was distributed some red earth, containing a few pieces of bone
and an implement or two. The caverns were of an irregular
height, owing to the yoof giving way more in some places than
in others, but the variation was from three to five feet; the width
of each was undetermined, as in some instances the separation
from one another was merely effected by a barrier of this stalag-
mite, and had it been considered safe to remove this they would
probably have found that, with the exception of a block of chalk
[ 28 ]
left here and there for support to the roof, the whole series was, in
fact, very nearly one large cave, with several openings. This was
not the case in every instance, and where they were enabled to
measure, from side to side, the width of the actual chamber was
about 4ft.
Cave No. 1 was explored to a length of 19ft., at which the solid
chalk was not reached, for the wall there was still composed of
blocks of stalagmite, and on placing a lighted candle to a chink
the effect of a strong draught was observed. Now, on the surface,
at a distance of 20ft., there was an indication of another shaft,
from which he thought it was but natural to conclude that they
communicated one with another. A similar case occurred in No.
6 cave, but there an unimportant difference was observed, for in
this instance they did clear the pit to the solid wall, and a squared
aperture of a few inches was found. If this communicated with
another open sbaft (as it was reasonable to suppose was the case),
it was somewhat strange that it was so small as to preclude the
interchange of anything except words or very little articles
between people stationed on either side. Part of a layer of undis-
turbed flint, of an exceedingly fine quality, lay at the bottom of
the galleries, and several detached pieces occurred all through the
filling up of the caves. Another layer of an inferior quality was to
be seen in the west side of the shaft, at a depth of 10ft., and two
vertical veins cut through the pit, one running N.W. and $.E., the
other nearly due E. and W.; the latter dipped to the south, at an
angle of 63 degrees.
Commencing the second division of the subject, various had
been the suggestions as to the purpose for which these galleries
had been made. A very natural theory, and which seemed to
present itself first to the mind was, that they were dwelling places,
or at least caves of refuge, if not for the whole year, at least for
seasons when the winter storms would render the rude wattle huts
but a poor shelter against the inclement weather; or when
harassed by war, the tribe in possession might have a safe retreat
for the women and children. He did not say that this might not
have been a secondary purpose to which they might have been
applied, but he deemed it highly- improbable primarily that for
either of these purposes they would have gone so deep; and
secondly that, had they done so, that a people even so barbaric as
these inhabitants must have been, should not have left some traces
in pottery, charcoal, remains of bones, of a continued ‘habitation.
Another suggestion was that they were storage barns for grain,
and what valuables they became possessed of, but if so why should
the shafts he filled up again so carefully 2 And how was the entire
absence of anything of the sort accounted for? ara:
| 49 J
January 14th, 1875.
ORDINARY MEETING.—* ON WINGLESS BIRDS.”—
MR. T. W. WONFOR.
The idea ordinarily associated in the mind with the word
“bird” was that of a creature possessing the power of flight and,
as a matter of consequence, provided with wings. In the north of
England “bird” or “brid” was a term used of any flying thing, so
that a moth, butterfly, or beetle would be equally included under
the term “bird.” Now as there were wingless or brevipennate
insects, the question might arise, are there wingless birds ? in the
usual application of the term. We know that the expressions
“wingless” and “brevipennate” had been applied to some
members of the family Aves, yet still, if we inquired into the
character of those birds which had been so designated we found
that, though we could not in all cases hold that they were flying
creatures, yet no one member of the great family of Aves was
absolutely deprived of wings. They might be merely rudimentary,
as in the case of the Apterix, or changed into flappers or paddles
as seen in the Penguins, but still they were present and corresponded
with the anterior pair of limbs found in all members of the family
possessing the power of flight.
THE APTERIX AUSTRALIS.
It was related that a few years since the skin of a bird brought
home by someone from New Zealand was given to a taxidermist to
set up, and he, taking into consideration the shortness of the wing
and absence of tail, and assuming that it was a penguin, stuffed it
in a sitting posture, with the head and neck arranged after the
manner of the penguins. This bird proved to be the Apterix
Australis, a native of New Zealand, and, for a bird, exhibiting
peculiar features. In addition to an almost want of wing, it pre-
sented a strange appearance, the plumage consisting of long flat
slender lanceolate feathers, each furnished with a soft shining
silken down, for the basal third of their length, and then narrow-
ing rapidly towards the extremity, where they presented the
appearance of single shafts, with hair-like webs on each side. The
G
[ 50 |
quill portion was very small and short, and overlapped by the
down, when the feathers were removed from the skin. Above and
below the eye, at the base of the beak, and on the forehead were
groups or pencils of long whisker-like hairs. As the kiwi, as the
natives called it, was nocturnal in its habits and lived among the
fern beds, boring imto the ground and seeking shelter in deep
excavations, these hairs might serve the same purpose as the
whiskers of the cat ; and supplement the eye, which, unlike that of
the other nocturnal birds, was small. So small was the organ of
vision, while the cornea was very convex, and the olfactory system
so much larger than in other birds, that Professor Owen remarked,
“The nocturnal habits of this bird, combined with the necessity for
a highly developed organ of smell, which chiefly compensates for
the low condition of the organ of vision, produce the most singular
modifications which the skull presents, so that it may be said that
those cavities which in other birds are devoted to the lodgment of
eyes are in the apterix almost exclusively devoted to the nose.” The
nostrils externally were very narrow, very small, avd set on each
side of the tip of the long curved beak, which at this point was
somewhat swollen. The internal olfactory apparatus and the
pituitory surface, on which the olfactory nerve freely ramified, was
complex and extensive. ° Like all the Cursores the legs were very
strong and muscular, the tarsi short and stout, the toes, four in
number, without intervening webs, the three anterior strong and
armed with powerful claws, the hinder one short and terminated
by a sort of spur, with which it was said to defend itself very
vigorously by striking very rapidly, and with great force. The
eggs, for so small a bird, were of great size, and showed the
absurdity of judging of the size of an animal from the size of the
egg. The eggs weighed on an average 143 ounces, while the adult
bird weighed 60 ounces; the proportion being about 1-4th, while
in the common fowl it was 1-48th, and in the case of the ostrich
1-100th; with some animals, such as the alligator and crocodile it
was much less.
GIGANTIC BIRDS.
There are, or, if extinct, were, birds of far more gigantic
proportions, viz., the dinornis and notornis of New Zealand, and
ABpiornis of Madagascar. The Dinornis giganteus stood from 14ft. to
16ft. high. Two portions of bone in the Brighton Museum would quite
[ 51 ]
lead, were other evidence wanting, to the belief that the bird was
not only of gigantic dimensions, but possessed of immense power
of stride. In connection with this bird, it is said that some years
ago a traveller returning from New Zealand, brought with him
a few inches of bone. This was before the days of the gold
discovery; the traveller had heard of the gigantic birds of New
Zealand, and also of Professor Owen, to whom the piece of bone
was presented. It was big enough in diameter to have been the
bone of an ox, but upon examination Professor Owen noticed a
peculiar cancellated structure only found in the bones of birds. A
bird to possess such a bone must have been considerably larger than
an ostrich, He therefore set to work to construct an outline of
the bird. The figure, when produced, caused no small amount of
disbelief and, as some thought, shewed hasty conclusions on Owen’s
part. But, causing enquiries to be made in New Zealand, some
disjointed bones turned up; these were packed up and sent in a
trunk to London; when put together they formed the incomplete
skeleton in the Museum of the College of Surgeons. From 14ft. to
16ft. appeared to be the height assigned to the dinornis giganteus,
the (tibia) legbone measured 2ft. 10in., and the legs to the root of
the tail 6ft. An egg found in the voleanic sand by Mr Walter
Mantel, was so large that he said his hat would serve as an egg-
cup for it. What became of this egg was not known. _ If, as was
asserted by some, the bird was still existing, we might in time
obtain a living specimen, or if the capture of it alive was impossible,
at least the skin and bones.
THE NOTORNIS MANTELLI.
Another brevipennate bird of the same country, the Notornis
Mantelli, was captured alive in the year 1849 by some seal hunters.
“They were ashore in one of the coves of Dusky Bay, the S.W.
extremity of Middle Island, when, observing the footprints of a
large and strange bird in the snow, they followed the trail and at
length came in sight of the birditself. Afteralong chase, in which
their dogs were very much distressed, they came up with and caught
it alive, in a gully behind Resolution Island. They kept it alive on
board their schooner for some days, and then killed and skinned it,
roasting and eating the flesh, which they pronounced delicious.
The skin was procured by Mr W. Mantell, and sent to England.
This bird stood about two feet high, the beak was short and strong,
[ 52]
wings very short and rounded, plumage feeble, legs and feet more
adapted for the land than those of the ordinary rails; plumage
rich purple on neck, breast, and abdomen, back and wings decked
with green and gold, tail scanty and white beneath; beak and legs,
when the bird was alive, bright scarlet.
AN ENORMOUS EGG.
In the Museum was the cast of an egg of enormous size. This
was in reality a fac-simile of an egg of the Hpiornis Maximus (a
supposed extinct bird of Madagascar), in the possession of Mr Dawson
Rowley, of Brighton, who had published a pamphlet upon it. Some
idea of its comparative size might be gathered when it was stated that
it would take 148 fowl’s eggs to equal it. The idea respecting this
egg was that it was laid by a bird of far larger size than the Dinornis,
and, if not still existing in the unexplored parts of Madagascar, that
it had not been extinct more than a couple of centuries. Professor
Owen, speaking of the eggs of this bird, in a paper read before the
Zoological Society in 1852, entitled “Notes on the egg and young
of the Apterix, and on the casts of the eggs and certain bones of
the Aipiornis,” thought it very unscientific to estimate the size of
a bird from the size of its egg, and proceeded to show, first that the
egg of the Apterix might have led to a supposition of its having
been laid by a much larger bird; and secondly, that from a
comparison of the bones, the Mpiornis did not equal in size the
Dinornis Giganteus.
BREVIPENNATE BIRDS.
Little more than 200 years ago there was on the islands of
Mauritius, Bourbon, and Rodiguez sundry brevipennate birds, the
Dodo, Solitaire, &c., in great abundance, and, till recently, only a
head, a couple of feet or so, and a few bones, with some paintings,
were all that remained to tell us of a very interesting group of
brevipennate birds. Much interesting matter respecting these
would be found in a work in their Library, “The Dodo and its
kindred.” The living specimen of the Dodo exhibited in London in
1639 passed into the hands of Tradescant, and, when his museum
was presented to the University of Oxford by Ashmole, it contained
a perfect stuffed dodo. On January 8th, 1755, by an order of the
Vice-Chancellor and his co-trustees, it was ordered to be burnt, the
bead and foot alone escaping destruction. Excavations made in
fF 53 J
1865 in the Mauritius by M. de Bissy, for the purpose of utilising
the soil of a marsh for manure, !ed to the discovery of various bones,
including those of the deer and tortoise. Mr Clark, who had long
had an opinion respecting the possibility of finding bones of the
dodo, told M. de Bissy his views. This led toa systematic ex-
ploration, and the discovery of many bones of that bird, and now,
in the British Museum, might be seen an almost perfect skeleton.
AFRICAN OSTRICHES,
Leaving the region of the comparatively unknown, we came to
well-known examples of wingless or brevipennate birds, all belonging
to the true Cursores; these were the ostriches, the emeu, and the
cassowaries. The best known species was the ostrich, Struthio
Camelus, an inhabitant of the African Continent. This bird, which
had been celebrated since the most remote antiquity, and a dish of
whose brains was an epicurean dish in Old Rome, measured from
six to eight feet in height; its feet consisted of only two toes; the
head and neck were nearly naked, the general plumage very lax, the
quill feathers of the wings and tail remarkable for the length of
their barbs, which, though furnished with barbules, were com-
pletely separated from each other, and formed the well-known
ostrich feathers of commerce. The ostriches lived together in large
flocks, feeding upon grass, grain, wild melons, &c., and, like the
gallinaceous birds, which they resembled in their food, had an
enormous crop and a strong gizard. Ina state of nature it picked
up and swallowed small pebbles; but in confinement it had
swallowed brickbats, knives, old shoes, scraps of wood, tenpenny
nails, bits of iron, and feathers; one went to the length of
swallowing in succession the whole of a brood of young
ducks—whether impelled by normal hunger, a morbid appetite,
or sheer mischief, was an open question. Another tried
to swallow its blanket. The voracity of the ostrich formerly
gave rise to the belief that it fed on iron. The African
ostrich was polygamous and gregarious. The female scratched
a hole in the sand, in which she lay ten or twelve eggs in
an upright position. The male and female both sat upon the eggs
during the night, and this sitting, supplemented by the heat of the
sun, hatched those in the middle of the nest, the outer ones, when
the centre eggs were hard and the young birds nearly hatched,
being quite fit for food; the eggs weighed upon an average 3lbs.,
i ome
and were regarded as great delicacies. Though equal in weight to
twenty-four hen’s eggs, one was not thought enough for a meal,
and in one instance two men finished five eggs in the course of an
afternoon. The approved method of cooking was to place the egg
upright on the fire, and break a hole in the top, through which a
forked stick was forced. This was made to rotate by rubbing with
the hands, and so beat up the contents while cooking.
AMERICAN OSTRICHES.
The American Ostriches contained two species. Rhea
Americana and R. Darwenii, and were scarcely more than half the
size of the African species, from which they also differed in having
the head and neck covered with feathers, and the feet furnished
with three toes. The feathers of the wing and tail, though
elongated, possessed none of the beauty of the African ostrich, and
were only employed in the manufacture of light dusting brooms.
They were very abundant in the large plains of America. The food
consisted mainly of grasses, roots, and other vegetable substances ;
but they would occasionally eat animal food, being known to come
down to the mud banks of the rivers for the purpose of eating the
little fish that had been stranded in the shallows. Darwin, who had
frequent opportunities of observing these birds, had given an
excellent account of their habits. He says:—‘They take the
water readily, and swim across broad and rapid rivers, and even
from island to island in the bays. They swim slowly, with the
greater part of the body immersed, and the neck extended a little
forwards. On two occasions I saw some ostriches swimming across
the Santa Ciruz river, where it was 400 yards wide and the stream
rapid.” It was polygamous; the male bird prepared the nest,
collected the eggs, which were frequently laid by the females at
random on the ground, and performed all the duties of incubation.
Darwin said four or five females had been known to lay in the same
nest, and the male, when sitting, lay so close that he himself nearly
rode over one. At thie time they were very fierce, and had been
known to attack a man on horseback, trying to kick and leap on
him.
THE AUSTRALIAN EMEU.
The Emeu of Australia, Dromaius Nove Hollandie was nearly
as large as the African ostrich, measuring from 5 to 7ft, in height,
[ 3 J
It had three toes on each foot, and these were each furnished with
nearly equal claws. The head and neck were covered with feathers,
the throat being bare, the plumage of the body, closely resembling
long hairs, hung down on each side of the body, from a central line
or parting. These birds, at one time abundant in Australia, were
now becoming extinct, for natives and Europeans were fast thinning
them, the former eating the eggs and hunting down the emeus for
food, but not allowing boys or women to partake of it, the flesh
being reserved for warriors and counsellors. Europeans and settlers
ran it with dogs, trained on purpose, for food, sport, and also for a
valuable oil, of which as much as six or seven quarts were yielded
by asingle emeu. This oil was of a light yellow colour, was used
as an embrocation for bruises or strains, and not readily congealing
or becoming glutinous, was also useful for oiling the locks of fire-
arms. They were monogamous, the male performing the office of
incubation; the nest was made by scooping out a shallow hole in
the ground, in some scrubby spot, and in this depression a variable
number of eggs was laid. Dr. Bennett remarked that “there is
_ always an odd number, some nests having been discovered with
nine, others with eleven, and others with thirteen.” These eggs
were nearly as large as those of the ostrich, but of a dark green
colour, and the young, when first hatched, were elegantly striped
with black and grey. In defending itself it did not kick forward
like the ostvich, but sideways and backwards like a cow.
THE CASSOWARY AND THE MOORUK.
The Cassowaries, of which there were at least two—the
Cassowary proper, Caswarius Guleatus, and the Mooruk, Casuarius
Benettii—were natives of the Eastern Archipelago. The former,
standing 5ft. high, was distinguished by the possession of a peculiar
horny crest or helmet upon the head, by the wings being furnished,
instead of feathers, with about five cylindrical stalks, destitute of
barbs, and by the large size of the claw on the inner toe. The head
and neck were naked and wattled, and of a bright red, variegated
with blue. The rest of the body, which was very stout, was clothed
with long glossy black pendant feathers, more closely resembling
hair than those of the emeu. It fed upon herbs, fruit, and seeds,
and, like the ostrich, swallowed hard substances. The eggs were of
a greenish tint. The eye was fierce and resolute, and the character
[ 56 ]
of the bird was tetchy, and it was apt to take offence without any
apparent provocation. Scarlet cloth excited its ire, and it had a
great antipathy to ragged and dirty persons. The height of the
mooruk was 3ft. to the top of the back, and oft. when standing
erect. The colour was rufous, mixed with black on the back and
hinder portions of the body, and raven black about the neck and
breast; the loose wavy skin of the neck was coloured with irridiscent
tints of bluish purple, pink, and an occassional shade of green;
the feet and legs were large and strong, of a pale ash colour, and
exhibited a peculiarity in the extreme length of the claw of the
inner toe of each foot, it being nearly three times the length of the
claws of the other toes. Instead of the helmet-like protuberance
of the cassowary, it had a horny plate resembling mother-of-pearl
darkened with black lead.
THE PENGUINS.
Another set of birds, if not wingless, should also be mentioned,
these were the Penguins, in which birds the wings were reduced to
a rudimentary character, were destitute of quills, and were covered
with a scaly skin forming flat fin-like paddles, the scales being
rudimentary feathers. In the water, which appeared their natural
element, they used them in swimming and diving. On shore they
used the paddles as anterior legs. From the backward position of
their feet the Penguins could only stand in a very upright
attitude, in which position they might be seen in countless numbers
arranged in as compact a manner and in as regular ranks as a
regiment of soldiers, and classed with the greatest order, the
moulting birds in one place, the young ones in another, the sitting
hens by themselves; the clean birds in another place, &. So
strictly did birds in a similar condition congregate that should a
bird in a moulting state intrude amongst those which were clean,
it was immediately ejected from among them.
THE PLUMAGE OF WINGLESS BIRDS.
Apart from what might be called the absence of wings or
rather the presence of merely rudimentary wings, wingless birds,
leaving possibly the penguin and dodo out of the category, were
distinguished from other birds by certain marked qualities. In all
of them the plumage differed from that of those possessing the
power of flight, the barbs of the feathers being always separate,
L 57 J
and the whole covering approaching very nearly the character of
ths hair of animals. The bones, too, were almost destitute of the
air cells which gave so much lightness to the skeletons of ordinary
birds, and assimilated more closely to mammalian bone. From
the mere rudimentary character of the wings there was an almost
absence of pectoral muscles, and the sternum was reduced to a
simple convex shield without any trace of the keel which in other
birds gave attachment to the powerful pectoral muscles. To
compensate for this deficiency, the great size and muscularity of
the legs rendered the pace of these birds in running very swift. The
pelvis was of a large size, and the two sides of the arch united at
the pubis; this was not the case in other birds. The anterior toes
were strong, either two or three in number, and terminated by
strong nails. The hinder toe, except in the genus Apterix, where
it was rudimentary, was entirely wanting. So strong were the
legs and muscles in the ostrich, that it would knock over a hyena
with a stroke, or rip up a dog with its claw. At bay it had
knocked down, and trampled upon the hunter who had approached
it incautiously.
THE ABODE OF BREVIPENNATE BIRDS
It must have been noticed that all the brevipennate cursores
were confined either to islands or two Southern continents, and
were of great size. How could this be accounted for; and was
there anything to explain the fact? First, as regarded the islands,
it seemed geologically evident that they, comparatively recently in
geological time, formed a part of the great Asio-African Continent
and that, when the separation took place, the ancestors of all these
gigantic and brevipennate birds were shut up in the islands
together with smaller birds, with the same lax plumage and feeble
powers of flight, those only of large size and swiftness of foot escaping
from their natural enemies shut up with them. That a ravenous
and active enemy in a small island would soon cause the extinction
of a small and feeble brevipennate bird was seen by what happened
in the Samoan group of islands. There was a Dodolet, the
— didunculus strigirostris, a pet with the natives, and a connecting
link between the true pigeons and the Dodo. European vessels
touching at these islands left behind them rats, which increased
and multiplied until, like Dick Whittington’s so-called cat of
H
[ 58 J
fable, but not the ship of reality, they introduced the domestic
cat; it not only kept down the rats, but destroyed the didunculus,
which became, as was believed, extinct, until a few months ago
living specimens were found and wrongly described in the journals
as the true Dodo.
THE SILK FOWLS.
Absence of wings and hairiness of plumage was seen under
domestication in what were known as Silk Fowls. The origin was
said to have been from an ordinary pair of Chinese fowls. A single
bird thus clothed was hatched from an egg laid by the hen, the
cause, though determining the variation, being unknown. By a
careful and long-continued selection of the offspring of this fowl
showing the most complete tendency to develope the peculiarity of
feathers, the breed of silky cochins was at length established. It
was well-known to pigeon fanciers that similar laxity of plumage
existed among them. Given therefore a tendency to laxity of
plumage, and but moderate powers of flight, natural selection would
explain on Darwin’s hypothesis, how, as the powers of flight
diminished, only the larger and more powerful would survive, while
the smaller and feebler would easily fall a prey to their natural
enemies, while the use of the legs and disuse of the wings would
also in time produce the swift and strong cursores like the ostrich,
emeu, Xe.
A vote of thanks to Mr Wonfor for his paper was proposed by
the President, Mr Alderman Cox. He thought it was very
fortunate for them that they could have such a stop-gap as they
had. Being personally perfectly ignorant of the subject treated,
the meeting, he said, would understand what a great temptation it
was for him to say much about it. There was a tendency in
human nature to say most about that of which they knew least,
but he would refrain from indulging in this natural propensity by.
merely proposing a vote of thanks in return for the instructive
paper read.—This proposition was heartily responded to.—A_ short
conversation was then engaged in, the Chairman, Mr Moore, Mr
Hennah, Mr Onions, Mr Dowsett, Mr C. W. Wallis, and one or
two others taking an active part in it.
To the Chairman the question suggested itself, whether the
rudimentary organs of brevipennate birds were in progress of
[ 59 J
development into something else, or had been wings and became
reduced through disuse, or had been always stunted.
Darwin’s belief in the second alternative was alluded to by Mr
Wonfor; but in opposition to this, Mr Moore gave the result of his
practical experience in New Zealand. There be had found that
brevipennate birds existed principally on soil roots, and consequently
had-no use for wings.
Mr Dowserr would not allow that there were so great
modifications in nature as Darwin suggested, although he admitted
that there were great modifications, the result of human agency.
Why were the birds mentioned not specially created in a form
adapted to the spheres they occupied ?
Another difficulty in believing Darwin’s theory was referred to
by the Chairman ; this was the density of the bone in the
brevipennate birds, in contrast with the animals themselves.
To meet Darwin half-way, Mr Wonfor suggested that, instead
of admitting that all forms of birds had been developed from a few
types, some acquiring greater powers of flight, and some ceasing to
fly altogether, they might concede that brevipennate birds had all
developed from one type, of which, perhaps, the ostrich or dinornis
was the original.
January 28th.
MICROSCOPICAL MEETING.—“ ROCK SECTIONS.”
The subject was to have been introduced by another gentleman,
but was in his absence introduced by Mr, T. W. Wonfor, the hon.
sec., who, since the last meeting of the society, had been elected
Curator of the Free Library, Museum, and Picture Gallery
After referring to the barrénness of scientific literature upon
the subject, Mr Wonfor proceeded to give a brief but comprehensive
sketch of the rock systems, constituting the crust of the earth, and
explained the characteristics of the three groups, the igneous, the
aqueous, and the organic. Dealing with the latter first, he showed
how at the present time rocks were being built upin the beds of
r 60 ]
the Atlantic, the Pacific, and elsewhere, the component parts of
which were similar to our chalk cliffs, mountain limestone, or coal
formations. The first two were made up of minute organisms, and,
taking into consideration the time necessary to form these rocks,
the age of the earth must be pushed much further back than where
it had been placed by many geologists.
The aqueous rocks bore distinct evidences of having been
formed under water; and in the igneous rocks, more particularly
in granites, minerals were found; while many rocks supposed to
be igneous contained water, glass, stone, and gas cavities.
One stumbling block to independent Microscopical research
amongst rocks was that so much aid was required from the
lapidary’s wheel, and the preparing of specimens for microscopic
inspection thus became an expensive amusement.
The meeting then became a conversazione, when about six
dozen sections, supplied by Mr. Wonfor, were exhibited, by different
gentlemen present amongst the principal being granite, porphyry,
felspar, jasper, obsidian, basalt, pitchstone, quartz, agate, lime-
stone, oolites, chalk, coal, and lignite.
FEBRUARY 14th.
ORDINARY MEETING.—MR. T. DAVIDSON, F.R.S.,
E.G.S., (ON “WHAT IS A. BRACHIOPOD?”
Having been requested by some members of the Brighton
Natural History Society to tell them what a Brachiopod is, he would
endeavour to do so in as brief a manner as so extensive a subject
would admit. All were aware'that it was very often much easier to
put a question than to obtain an entirely satisfactory answer, and
he was consequently sorry to have to begin his few observations on
a very extensive group of organisms by stating that zoologists and
comparative anatomists had not yet entirely agreed as to the exact
position it should occupy among the invertebrate animals. The
first species belonging to the class were imperfectly and quaintly
described and figured by Prince Fabio Columna, as far back as
1606, and for many years were supposed to be referable to the genus
f ory
Anomia of the Lamelli branchiata; but, as was judiciously observed
hy Edward Forbes, “a close examination shows that there is no
relationship between them, but only a resemblance through formal
analogy.” It was during the present century that the class itself
had been worked out and understood, and this had been achieved
only after the most lengthened and persevering researches.
NAME.
The name Brachiopod (an arm, a foot) was proposed for the
class by Cuvier in 1805, and by Dumeril in 1809, and had since been
very generally adopted. Professor King, perhaps rightly, objected
to the term on the grounds that it was a misnomer, for the two
variously curved and cirrated brachial appendages, improperly
designated as arms or feet, were subsequently found not to subserve
the function of locomotive organs. Blainville, in 1824, proposed as
a substitute for Cuvier’s name that of Palliobranchiata (a mantle,
gills), on account of the respiratory system being combined with
the mantle on which the vascular ramifications are distributed.
SHELL.
Before alluding to the position the Brachiopod should occupy
amongst the invertebrata, he might, at once, observe that the
animal was protected by a shell composed of two distinct valves;
and that these valves were always, except in cases of malformation,
equal sided, but not equivalved. The shell was likewise most varied
and beautiful in its endless shapes and variations; in some species
it was semi-transparent, and glassy, in others massive; generally
the shell was from a quarter of an inch to about fonr inches in size,
but in certain species was nearly a foot in breadth by something
less in length, as was the case with Productus Giganteus. The
valves were often very unequal in their respective thickness, as
might be seen in Productus Llangollensis, Davidsonia Verneuilii,
&e., and while the space allotted to the animal was very great in a
number of species it was extremely small in many others. The
outer surface of many of the species presented likewise the most
exquisite sculpture, heightened by brilliant shades, stripes, or spots
of green, red, yellow, and bluish black. The valves had been
distinguished by various names, but those of dorsal and ventral were
in more general use. The ventral one was usually the Jarger; in
[ 62 |
many genera, as Terebratula and Rhynchonella, had a prominent beak
with a circular or otherwise shaped perforation or foramen at, or
near, its extremity, partly completed by one or two plates termed a
deltidium ; through the foramen passed a bundle of muscular fibres
termed a peduncle, and by which the animal was,in many species,
attached to sub-marine objects during at least a portion of its
existence. It was, however, certain, from the admirable researches
of Professor Morse, that the embroyo of some, if not of every
species, swan most actively in every direction and turned abruptly
about; but that, in the fourth stage of its development, it became
attached, the peduncle widening at its end into a sucking disk.
Other species showed no indication of ever having been attached,
while some, that had been fixed by means of a peduncle during a
part of their existence became attached at a more advanced stage
of life, the opening for the peduncle becoming gradually cicatrized.
Lastly, certain forms appeared to have adhered to submarine
objects by a larger or smaller portion of the surface of their ventral
valve, or by spines (Strophalosia) during their entire life. In
external shape the valves were essentially symmetrical, which was
not the case with the Lamellibranchiata, so much so, that certain
Brachiopod shells received the name of Lampides from some early
naturalists, but while such might bear an obscure kind of
resemblance to an antique Etruscan lamp, by far the larger number
in no way resembled one. The valves were also either articulated
by means of two curved teeth developed from the margin of the
larger valve and fitting into sockets in the corresponding part of
the smaller one, or they were unarticulated and kept in place or in
juxtaposition by the means of muscles especially provided for that
purpose.
SHELL STRUCTURE.
The structure of the shell had been shown by Dr Carpenter,
Professor King, and others to be generally distinct from that
of the Lamellibranchiata or Gasteropoda. In some families,
according to Professor King, it consisted of three divisions,
the innermost and middle divisions, which constitute the entire
thickness of the valve, were calcareous, with a prismatic or fibrous
structure ; while the outer division consisted of a very thin
membrane. The innermost and intermediate divisions in some
families were traversed by minute tubular canals, which passed
{ 63 ]
from one surface to the other, for the most part in a vertical
direction, and at tolerably regular intervals; but, just before
terminating near the outer surface of the epidermis, their tubular
orifices suddenly became dilated, the lower half of the canal was
often considerably smaller in diameter than the upper half. The
‘canals were occupied by coecal processes proceeding from the
mantle or fleshy covering of the animal. Their functional nature
was, according to Dr. Carpenter, branchial in their office, and
subservient to respiration; but,as observed by Professor King, the
outer epidermis, which covered their expanded termination, would
seemingly prevent any communication between the surrounding sea
water and the mantle, so that it might be questioned whether they
were at all connected with the respiratory function. In certain
genera, such as Lingula, there were no canals, the shell being found
to consist of flattened prisms of considerable length, and which lay
parallel to one another with great regularity, and at a very acute
angle. The shell substance was also either almost entirely
composed of a horny or of a calcareous substance.
INTERIOR OF. VALVES.
Having described the exterior of the shell, he must add a few
words with respect to the interior. On the imner surface of both
valves several well defined muscular impressions were observable;
these varied considerably in position and shape in different species.
They formed either indentations of greater or lesser size and depth,
or occurred as variously shaped projections. In the Trimerellide
some of the muscles were attached to a massive or vaulted platform
situated in the medio—longitudinal region of the posterior half or
umbonal portion of both valves. In addition to these there existed
in the interior of the dorsal valve, in some genera a variously
modified thin calcified ribbon-shaped lamina or skeleton for the
support mainly of the brachial or labial appendages, and so varied
and constant were these in shape to certain species that this
laminal apophysis had served as one of the chief characters
in the creation of both recent and extinct genera. In certain forms
the lamina was more or less developed, and so bent as to assume the
shape of a loop. The loop was in some genera attached only to the
hinge-plate, as in Terebratula, Waldheimia, &c., in others likewise
so to a central longitudinal plate or septum. In certain families
the apophysis- presented the form of two spiral processes, which
[ 64 J
nearly filled the interior of the shell, the ends of the spires being
either directed outwardly towards the cardinal angles (Spirifer)
or placed horizontally with their apices directed inwards and
towards the centre of the concave surface of the same valve, which
they almost filled, the inner side of the spires were pressed together
and flattened with thin terminations close to each other, near the
centre of the bottom of the shell (Atrypa). In the Rhynchonellide
again it assumed the shape of two short slender curved lamine,
while in many genera, and even families, Productide, &c., there
existed no calcified support for the branchial appendages.
SOFT PARTS OF ANIMAL.
He must very briefly allude to the soft parts of the animal,
but it would require much more time than could be bestowed to
his short lecture, to do anything like justice to so difficult and
complicated a subject. He was, however, truly happy in being
able to inform the society that this important enquiry had been
most ably and successfully elaborated during the last twenty
years, and by some of the most distinguisned anatomists of the
period. To such men as Owen, Huxley, Hancock, Gratiolet, Vogt,
Morse, and others, we were indebted for an extensive series of the
most elaborate and accurate dissections and observations, which
had defined to a very considerable extent, what were the true
characters of the Brachiopod, while some important researches
elaborated by Steenstrup, Lacage-Duthiers, Morse, Dr. Fritz
Miller, Oscar Schmidt, McCrade, Kowalevsky, and others, had
thrown much additional light upon the embiyology, or early stages,
of the groups. Some differences of opinion, it was true, had been,
and were still, entertained with respect to the exact function to be
attributed to certain parts of the animal; but on all essential
questions there was a pretty general agreement. Before describing
the soft parts of the animal, it might be as well to mention that
Brachiopoda had been divided into two great groups by Bronn,
which he termed Apygia and Plewropygia, considering them to be
inadmissible on certain grounds. Professor King substituted the
name Clistenterata for the first group, on account of its including
animals destitute of an anal aperture; and the term Tretenterata
for the second, as it embraced animals provided with this organ.
The former division contained species which had their valves
L 65 |
articulated as Terebratula, Spirifer, Rhynchonella, the latter com-
prised species with non-articulated valves as Lingula discina, ec.
Some very important modifications in the animal were connected
with these two divisions, especially in what related to the muscular
system. According to Morse, the Brachiopod were reproduced by
eggs, generally kidney-shaped and irregular, which were discharged
from the anterior margin of the shell, and dropped just beyond the
pallial membrane, hanging in clusters from the sete. Some
uncertainty had prevailed as to whether there was a male and
femule individual; Lacage-Duthiers and Morse stated that the sexes
were separate and described them as such in Thecidium and Tere-
bratulina, and the French zoologist went so far as to suggest that
a difference was even observable in the shell. Professor Morse
described the embryo of Terebratulina with great minuteness during
its six stages of development. It was divided into two, three, or
four lubes clothed with vibratile cilia; and before becoming
attached, swam or whirled head foremost by means of vibratile
cilia which covered the body.
MANTLE.
Both valves were lined by the delicate membrane, termed the
“pallium,” or mantle; it secreted the shell, and was generally
fringed with horny bristles, or sete. The mantle was composed of
an outer and inner layer, between which were situated blood
channels or lacunes. These arterial trunks and veins corstitute the
vascular system. On certain parts of the surface of the mantle
occurred a vast number of microscopic, flattened, calcareous,
denticulated plates, or spicule, destined, no doubt, to stiffen the
portions that contained them. The digestive organs, viscera, as well
as the muscles, took up only a small place in the contiguity of the
beak, and were separated from the general cavity by a strong
membrane, in the centre of which the mouth is located. The nervous
system consisted of a principal ganglion, of no great size.
BRACHIAL APPENDAGES.
He must next briefly describe those beautiful and singular
organs, eminently characteristic of the Brachiopoda, termed arms,
or more correctly labial, on account of each member being a
prolongation of the lateral portion of the lips or margin of the
mouth; the lamellibranchs, or conchifera, had analagous
[ 66 |
appendages, but very much less developed. They assumed different
shapes in different genera, and were supported, or otherwise, by the
more or less complicated calcified skeleton, already described.
These two long, bent, or spirally convoluted organs, occupied the
larger portion of the cavity of the shell, and were mainly composed
of a membranous tube, fringed on one side with long fiexible cirri.
The brachial appendages occupied, therefore, nearly the entire
pallial cavity, but were not capable of being protruded in those
families and genera in which they were folded upon themselves,
or supported by a calcareous skeleton. In Rbyncbonella,
where the elongated spiral arms were but slightly supported
at their origin by two short projecting calcareous processes,
they could, at the will of the animal, be unrolled, and
protruded to some distance beyond the margins of the shell, and,
when forcibly stretched out, were said to be more than four times
the length of tlie shell, and to support some 3.000 cirri. It must,
however, remain for ever uncertain whether, in those extinct
genera, such as Spirifer and Atrypa, where the spirally coiled fleshy
arms were supported throughout their entire length by a calcified
skeleton, the animal could protude its brachial appendages beyond
the margin of the valves. In some families, Rhynchonellide,
Productide, the arms are spiral and separate, in others, Lingulide,
only at their extremities, and it was almost certain that these
beautiful organs by means of their cirri and the cilia, with which
they were doubtless furnished, were not only instrumental in carry-
ing floating nutrimental particles or minute organisms to the
mouth, but were subservient to the functions of respiration.
MOUTH: HEARTS.
The mouth conducted by a narrow cesophagus to a simple
stomach, which was surrounded by a large granulated liver.
Owen’s “ hearts” had been found to be oviducts, while the true
heart would consist of a pyriform vesicle, appended to the dorsal
surface of the stomach.
MUSCLES.
He must next speak of the muscular system, and as the num-
ber and position of these muscles differed materially in the two
great divisions into which the Brachiopoda had beeu separated, and
to some extent also in the different genera of which each division
[ mm
was composed, it might be desirable to treat the subject under two
different heads in the Clistenterata, of which the genus Terebratula
might be taken as an example, five or six pairs of muscles were
stated by Hancock and others to be connected with the opening
and closing of the valves, or with their attachment to, or movements
upon the peduncle. First of all, the adductors, or occlusors, consisted
of two muscles, which, bifurcating near the centre of the shell
eavity, produced a large quadruple impression on the internal sur-
face of the small valve, and a single divided one towards the centre
of the large or ventral valve. The function of this pair of muscles
was the closing of the valves. Dr. Gratiolet who had likewise de-
scribed with great minuteness, the muscles of the Brachiopoda, in-
formed us that those which close and open the valves were the only
ones known to Pallas, but that he defined their positions and
functions clearly. The same was done by Blainville and Quenstedt.
The next two pairs of muscles had been termed divaricators, or
cardinal muscles, and had for function the opening of the valves.
The divaricators proper were stated by Hancock to arise from the
ventral valve, one on each side, a little in advance of and close to
the occlusors, and after rapidly diminishing in size became attached
to the cardinal process, a space or prominence between the sockets
in the dorsal valve. The accessory divaricators were, according to
the same authority, a pair of small muscles which had their ends
attached to the ventral valve, one on each side of the median line,
a little behind the united basis of the occlusors, and again to the
extreme point of the cardinal process. Two pairs of muscles
appeared to be connected with the peduncle and its limited move-
ments had been minutely described by Hancock as having one of
their extremities attached to this organ. The dorsal adjustors
were attached to the ventral surface of the peduncle and again in-
serted into the hinge plate in the smaller valve. The ventral
adjustors were considered to pass from the inner extremity of the
peduncle and to become attached by one pair of their extremities
to the ventral valve, one on each side of, and a little behind, the
expanded base of the divaricators. The function of these muscles
was not only that of erecting the shell, but also to attach the peduncle
to the shell, and thus effect the steadying of it upon the.peduncle.
Such was the general arrangement of the shell muscles in the divi-
sion comprising the articulated Brachiopoda, allowance being made
t
f 68 ]
for certain unimportant modifications observable in the shells com-
posing the different families and genera thereof. In the Tretenterata,
of which Lingula might be quoted as an example, the myology was
much more complicated, and anatomists had differed considerably
in their respective views concerning the function of some of the
muscles. It would not, however, be possible to discuss the opinions
of Hancock, Vogt, Gratiolet, and others, but he would refer to
and adopt those recently advocated by Professoy King, as
they seemed to carry with them a greater degree of plausibility.
Of shell or valvular muscles, Professor King, made out five pairs
and an odd one. Three pairs were laterals having their members
limited to the sides of the shell; one pair were transmedians, each
member passing across the middle to reverse side of the shell, one
pair had its members confined to the central region; while the odd
muscle occupied the umbonal cavity, and he assumed that the
central and umbonal muscles effected.the direct opening and closing
of the shell ; the laterals enabled the valves to move forward and
backward on each other, while the transmedians allowed the similar
extremities (the rostral) of the valves to turn from each other to the
right or the left on an axis subcentrically situated, that was in the
medio transversed region of the dorsal valve. It was longa matter
in discussion whether the animal could displace its valves sideways
when about to open its shell, but this matter had been set at rest by
Professors Semper and Morse, who actually observed the animal
perform the operation; they mention that it was never done
suddenly or by jerks, as the valves were at first always pushed to
one side several times and back again on each other, at the same
time opening gradually in the transverse direction, till they rested
opposite to each other and widely apart. Those who had not seen the
animal in life, or who did not believe in the possibility of the valves
crossing each other with a slight obliquity, would not consent to
appropriating any of its muscles to that purpose, and consequently
attributed to all the lateral muscles the simple function of keeping
the valves in an oppusite position, or holding them adjusted. They
had not only the observations of Semper and Morse, but the
anatomical investigations of King, to confirm the sliding action or
divarication of the valve of Lingula. In the clistenteratu, where no
such sliding action of the valves was necessary or possible, no
muscles for such an object were required; consequently none took
rise from the lateral portions of the valves as in Lingula, but, in an
[ 69 |
extinct group, the Trimerellidw, which seemed to be somewhat
intermediate in character between the Tretenterata and Clisten-
teruta, certain scars which had been found, appeared to have been
produced by rudimentary lateral muscles, but it was doubtful
(considering the shells are furnished with teeth, though but rudely
developed) that such muscles enabled the valves, as in Lingula,
to move forward and backward upon each other. There were
muscles connected with other portions of the animal, such as the
parietal muscles so strongly defined in the Tretenterata, and dis-
tinctive peculiarities of the Peduncle, &c., but which the limited
. time at his disposal would not allow him to describe.
AFFINITIES OF THE BRACHIOPODA,
It was, however, time that he should say a few words in
connection with the systematic position of the Brachiopoda. As
they were aware, the invertebrata had been grouped into five sub-
kingdoms, namely, the Protozoa, Celenterata, Annuloida, Annulosa,
and Mollusca; and that for many years the Brachiopods had been
considered to constitute a separate class among the mollusca, a view
still maintained by several distinguished naturalists. Subsequently,
the class was removed from the Mollusca proper, and placed along
with the Polyzoa and Tunicata into a division to which the name
Molluscoidea was given. Again, within the last few years, Professors
Kowalevsky and Morse, in several very able memoirs, had strongly
urged that the Brachiopods should be considered to constitute a
separate group among the Annelids, and it must be admitted that
the American and Russian zoologists have clearly shown that
several portions of the animal of the Brachiopod and Amphetrite
presented important characters in common; but, at the sam® time,
as was observed to him by Professor Verrill, almost any invertebrate
group may be anneledelized by overrating certain points of their
affinities; nor could one cast aside many true molluscons characters
presented by the Brachiopod, and, as was justly observed by
Stoliczka, there cannot be, be thought, much doubt as to the true
molluscous character ‘of the Brachiopod and their proper classifica-
tion between the Anomiide of the Pélecypoda, and Saccopoda, and
_ the arm bearing section of the Ciliopoda. The subject in connection
with the systematic position of the Brachiopoda must, therefore, be
left, for the present, as an open question, although he was still in
Lm
favour of the opinion that the class should be considered to
constitute a distinct group among the Molluscoidea.
RANGES OF DEPTH.
All Brachiopoda were inhabitants of the sea, and had been met
with at depths ranging from less than a foot to that of 2,600 fathoms,
between Kerpules Island and Melbourne. (Her Majesty’s Challenger
expedition). Some Lingule lived partly buried in mud; while Pro-
fessor Morse described a species of the genus which he found in
vast numbers “in a sand shoal at low water mark buried just
below the surface of the sand. The peduncle, six times the length
of the shell, was partly encased in a sand tube differing in no
respect from the sand tubes of neighbouring annelids. He
observed (likewise) that Lingula pyramidala had the power of
moving over the sand by the sliding motion of the two valves, using
at the same time the fringes of sete, which swung promptly back
and forth like a galley of oars, leaving a peculiar track in the sand.
In the motion of the sete he noticed the impulse commencing from
behind and running forward. The larger number of species lived,
however, at depths from five to three or four hundred fathoms,
some apparently in the free condition, others attached by their
peduncle to various marine objects, and very often to coral reefs.
Discina had been found attached to stone at low water, and been
dredged from depths ranging from five to nearly 2,000 fathoms
(Discina atlantica). Mr. Jeffreys did not believe that the habitat
of any invertebrate animal was affected by bathymetrical conditions.
Jukes got any number of Waldheimia flavescens while boating in
Australia among the reefs, they were merely washed by the tide,
and he gathered them with his hand like limpets on the shore.
Certain minute forms were found by Mr Jeffreys fixed to seaweeds,
while others adhered by a larger or smaller portion of one of their
valves to submarine rocks, stones, corals, &e.
_ CLASSIFICATION.
It would not be possible on so brief a paper to enter at any
length on the subject relating to the classification of the Brachiopoda.
This important matter would be found fully discussed up to the
-year 1853 in the general introduction to his great work on “ British
Fossil Brachiopoda.” He then published his own views, and which
[a ]
were subsequently very generally adopted both by British and
foreign palzontologists, but he did not omit to impress upon bis
readers that we were not then, nor were we even yet, quite in a
condition to prepare a really complete, permanent, or wholly
satisfactory classification of the numerous species composing the
class. In 1853, he divided the Brachiopoda into eight families,
comprising twenty-four genera and twenty-two sub-genera, but
during the years that had elapsed since 1853 up to the present time
some seventy-two more genera or sub-genera had been proposed,
and of which he published a list in the Sussea Daily News newspaper
for the 20th of August. 1872. Much consideration on his part had
been devoted to the subject, but he felt that in order to place the
known genera and species into their respective families, or into new
ones that would have to be created, much more information would
have to be acquired. The subject was immense, when one had to
grapple with between five and six thousand described species,
varieties, or synonyms. J. Bigsby gave a list of upwards of three
thousand species alone from the paleozoic rocks, and it must likewise
be borne in mind that many of the extinct genera had as yet been
but imperfectly elaborated. The material in hand was, however, so
great that, with time, palzontologists would be able to lay before
the public a complete history of a class, which, as would be shortly
shown, had played an important part in the great life-system of
our globe, from its dawn to the present time. It would be necessary
though, to admit the two great divisions Tretenterata and
Clistenterata into any scheme of classification, although it was
impossible to say whether or not all the extinct genera were
provided or otherwise with an anal aperture. The Tretenterata
comprised the families Lingulide, Discinide, Craniade, and (?)
Trimerellide, Bach of these families was composed of several
genera. The Clistenterata would include the families Terebratulide,
Rhyenchonellide, Spiriferide, Strophomenide and Productide. By
far the larger number of described genera and species would have to
be located in this great division of the Brachiopoda.
DISTRIBUTION IN TIME.
Assuming that they were all acquainted with the geological
divisions into which the crust of this earth had been grouped, he
might at once observe that, as was justly remarked by Barrande,
[ 72 ]
the Trilobites and Brachiopoda occupied the most important place
in the primordial fauna of our globe, and if we excluded the proble-
matical Hozoon canadense from the animal creation. as some
naturalists had done, we should find the Brachiopoda making their
first appearance in the oldest known fossiliferous deposits, for Mr.
Hicks bad obtained undoubted examples of Lingula or Lingulella,
(L. primeva) from the very base of the whole Cambrian series of St.
Davids. They were less numerous during the Permian and Triassic
periods, while they again became abundant, although comparatively
much less numerous, during the Jurassic and Cretaceous times.
In the Tertiary period they had materially decreased in number, and
were represented at the present time by about one hundred species.
But what wonderful changes had been operating during the in-
calculable number of ages in which the creation (?) and extinction
of a large number of genera, and thousands of species, had taken
place, some few only of the first created genera, such as Lingula,
Discina, and Crania, have fought their way and struggled for
existence through the entire sequence of geological times. Many
were destined to a comparatively ephemeral existence, while others
had a greater or lesser prolongation of reproduction. These
remarks led him to give some extracts from a letter which he
received from Darwin as far back as the 26th of April, 1861. In
that letter, that eminent and admirable observer writes, “I do not
know whether you have read my ‘ Origin of Species.’ In that book
I have made the remark, which I apprehend will be universally
admitted, that, as a whole, the fauna of any formation is inter-
mediate in character between that of the formation above and
below. But several really good judges have remarked to me
how desirable it would be that this should be exemplified
and worked out in some detail, and with some single group of
beings. Now, every one will admit that no one in the worid could
do this better than you with Brachiopods. The result migbt turn
out very unfavourable to the views which I hold ; if so, so much the
better for those who are opposed to me. But I am inclined to
suspect that, on the whole, it would be favourable to the notion of
descent with modification. I can hardly doubt that many curious
points would occur to anyone thoroughly instructed in the subject,
who could consider a group of beings under this point of view of
descent with modification. All those forms which have come down
» Sha Rat AN ae
lL 7 |
from an ancient period very slightly modified ought, I think, to be
omitted; and those forms alone considered which have undergone
considerable change at each successive epoch. My fear is whether
the Brachiopods have changed enough. The absolute amount of
difference of the forms in such groups at the opposite extremes of
times ought to be considered; and how far the early forms are
intermediate in character between those which appeared much
later in time. The antiquity of a group is not really diminished,
as some seem to think, because it has transmitted to the present
day closely allied forms. Another point is, how far the succession
of each genus is unbroken from the first time il appeared to its
extinction, with due allowance made for formations poor in fossils.
T cannot but think that an important essay (far more important
than a hundred literary reviews), wight be written by one like
yourself, and without very great labour,’ &c. In several subse-
quently written letters, Darwin reiterated bis suggestion. Hecould
assure them that he had not neglected a request coming from so
eminent a quarter; but he was bound to state that he had found
the subject beset with so many apparently inexplicable difficulties,
that year after year had passed away without his being able to
trace the descent with modification among the Brachiopoda which
the Darwinian Doctrine required. The imperfection (one due, he
believed to our slight acquaintance with the subject) in the geolo-
gical record could not in many cases be doubted, but we had no
right to make capital out of unknown data, we must therefore deal
with facts as we found them, and see how far they would bear upon
the subject under examination. It was quite true that strata at
great distances could not be positively asserted to be, strictly speak-
ing, absolutely contemporaneous, although they might contain the
same animal remains. It was very probable that some species
might have emigrated from the sea bottom on which they originally
lived to some more favourable locality, and become to some extent
modified. No one could seriously doubt that life had continued to
be represented under one form or another ever since it was first
brought into existence. He ‘consequently could not agree with
Deshayes and others who believed in a total extinction of the
animal creation at certain specified periods. Notwithstanding the
theoretical doctrine that had been promulgated with respect to the
origin of species, we were still, and would probably for ever remain,
in the dark, or within the region of suppositions with respect to so
K
[ & J
important a question. In his admirable address to the Belfast
meeting of the British Association, Tyndall observes, “If you ask
me whether there exists the least evidence to prove that any one
form of life can be developed out of matter, without demonstratable
antecedent life, my reply is that evidence, considered perfectly con-
clusive by many, has been adduced, . . . . . but there is in
the true man of science a wish stronger than the wish to have his
beliefs upheld—namely, the wish to have them true ;” and He adds
further on, ‘‘ that these observers will frankly udmit their inability
to point to any satisfactory experimental proof that life can be
developed save from demonstratable antecedent life,” and that the
whole process of evolution is the manifestation of a power absolutely
inscrutable to the intellect of man.
Darwin’s tempting and beautiful theory of descent with modifi-
cation bore a charm that appeared to be almost irresistible, and he
would be the last person to assert that it might not represent the
actual mode of specific development. It wasafar more exalted con-
ception than the idea of constant independent creations, but we were
stopped by a number of questions that seemed to plunge the concep-
tion into a maze of inexplicable, nay mysterious, difficulties; nor had
Darwin, as far as he was aware, said how he supposed the first pri-
mordial form to have been introduced. The theory was at best, as
far as we could at present perceive, with our imperfect state of know-
ledge, but balf the truth, being well enough in many cases between
species and species; for it was evident that many so-termed species
might be nothing more than modifications produced by descent. It»
applied likewise to accidental variations, as between closely allied
genera, yet there was much more than this with respect to which the
Theory seemed insufficient. The strange geological persistency of
certain types such as of Lingula, Discina, Nautilus, &c., seemed also
to bar the, at present, thorough acceptance of such a theory of
general descent with modification,
What did the Brachiopoda tell us upon this question. Taking
the present state of our knowledge as a guide, but admitting at
the same time that any day our conclusions and inductions might
require to be modified from fresh discoveries, let them ascertain
whether they revealed anything to support the Darwinian ideas.
They found that by far the larger number of genera made their
first appearance during the Paleozoic period, and, since then, had
[ % |
been decreasing in number to the present period. They might
leave out of the question the species, for they varied so little that
it was often very difficult to trace really good distinctive characters
between them, it was different with genera as they were, or
should be, founded on much greater and permanent distinctions.
Thus, for example, the family of Spiriferide included genera .
characterised by calcified spiral lamina for the support of the
brachial appendages, and however varied they might be they always
retained the distinctive characters of the group from their first ap-
pearance to their extinction. The Brachiopodist laboured under the
difficulties of not being able to determine what are the simplest, or
which are the highest, families into which either of the two groups
of his favonrite class was divided, so far, then, he was unable to
point out any evidence favouring progressive development in it,
But, confining himself to species he had often before him great
varietal changes, so much so, as to make it difficult for him to
define the species; and it led him to the belief that such groups
were not of independent origin, as was universally thought before
Darwin published his great work on the origin of species. But in
this respect the Brachiopoda revealed nothing more than other
groups of the organic kingdom. All the Productide had their
permanent and constant characters, and so on. It would appear
that the earliest forms among the Brachiopoda are referable to the
division Vretenterata, which included the genera lingulella, lingula,
discina, and obolella. Of these, lingula and discina only had lived
on with but slight modifications in external shape, during the
entire sequence of geological time, and they were still represented
by several species, but in rocks, somewhat later in age, the
Menevian group, or lower lingula flags, there occurs a species of
orthis, the first representative, as far as they were aware, of the
division Clistenterata, but there was not the slightest evidence
that orthis was ever derived from any of the four genera of
Tretenterata above named. Since that period both divisions
continued to be represented, without ever showing a tendency to
pass one into the other. Now, although certain genera, such as
Terebratala, Spirifer, Rynchonella, Crania, Discina, and Lingula,
had enjoyed a very considerable geological existence, and had
varied very little since their first appearance, and might be looked
upon as among the earliest. Brachiopoda, there were genera such
as Stringocephalus, Uncites, Porambonites, Koninchina, &c., which
[ 76 J
made their appearance very suddenly and without any warning,
and disappeared in a similarly abrupt manner, having enjoyed a
comparatively short existence. They were all possessed of such
marked and distinct internal characters that we could not trace
between them and associated or synchronous genera any evidence
of the result of descent with modification, and no imperfections
in geological record would come to our aid to clear away the
difficulty. Therefore, although far from denying the possibility or
probability of the correctness of the Darwinian Theory, we could
not conscientiously affirm that the Brachiopoda, as far as we were
at present acquainted with them, would be of much service in
proving a theory, which, however delightful in its conception, had
still to be made certain by positive and indisputable evidence, and it
was a subject worthy of the continued and serious attention of every
well-informed man of science, for the sublime Creator of the
universe had bestowed on man a thinking mind, therefore, all that
was discovered was legitimate. Science had also this advantage
over theology, that it was continually on the advance, and ever
ready to correct its errors when fresh light or new discoveries make
such necessary. The importance of the study of the Brachiopoda
was very great, when they remembered that they were among the
they first well authenticated indications of life in this world, and that
had continued to be very extensively represented up to the present
time. They were also very abundant and characteristic fossils, by
which rocks at great distances could be identified, and without its
being even necessary for the Palwontologists to visit the district
from whence they were derived, and as we became more intimately
acquainted with their characters, their interest and usefulness was
augmented. They were, as Mantell would have termed them, sure
medals of the creation, with the date of their appearance firmly
stamped upon them, and their distinctive characters so legibly
impressed as to defy misinterpretation.
Mr. Davidson’ resumed his seat amid loud and continued
applause. The paper was illustrated by beautifully designed and
executed diagrams, by Mr Davidson himself, and specimens
illustrating the Brachiopoda were exhibited. A hearty vote of
thanks to Mr Davidson, for the trouble he had taken in the prepara-
tion of the paper read, was proposed by the Chairman, Mr Alderman
Cox, He thought that a more than usually hearty vote bad been
wi
f @]
well-merited. Asking pardon for digressing from the subject
before them, he broached a tender matter by expressing how deeply
he regretted that, for a time, Mr Davidson’s services had heen lost
to the Museum). He, however, hoped that Brighton would still be
favoured with the services of a gentleman whose name was known
wherever the name of geology or science itself was recognised. The
motion was unanimously carried.
The deep regret experienced at the loss of Mr Davidson’s
services to the Museum was about to be further commented upon
by Mr Alderman Mayall, but the Chairman interposed by remarking
that he thought they had better not go into that subject, although
he had been allowed to revert toit. Myr Alderman Mayall then
paid a tribute to the great care with which Mr. Davidson had
investigated science, and asked that gentleman whether, in any of
the dredging expeditions, extinct genera had heen re-discovered.
Mr. Davipson replied that he had had several letters from
the Challenger Expedition, and had seen all the forms discovered
in the course of the other expeditions—he, in fact, had been asked
to describe the Brachiopoda obtained by the Challenger Expedition,
having done so for various expeditions—and not a single extinct
form had been found. In reply to the Chairman, Mr Davidson
further stated that he had 40,000 specimens, from every part of the
world, and fully four-fifths of the known species.
Mr. Wonror observed that the geologic story as told by the
Brachiopoda did not bear out the theory of evolution. Besides the
fact that when the genus disappeared it never reappeared, one could
hardly imagine that Tretenterata with a mouth and anal opening
degenerated into the Clistenterata with a mouth but no anal
opening. In the latter, one aperture had to perform the double
office of mouth and anus, and in the former these functions
were performed by two apertures—an evidently higher type than
the other.
Mr HAsELWoop reminded the meeting that this theory of
evolution was only a working hypothesis, and one of the best which
they had. He also took this opportunity of remarking that they
could not over-estimate the value of such a gentleman as Mr.
Davidson. If the world was to progress, it was by such devotion
Reo
as his, and by that alone; and when a gentleman of his means and
capacity would set apart his valuable life to a seemingly out-of-the-
way subject like this, and proclaim such facts as those to which he
has given utterance that night, no man was more worthy of the
praise and honour of his fellow men.
In answer to Mr Panxuurst, it was pointed out that the
Brachiopoda served a purpose in the animal economy. They con-
stituted food for other animals, and were even eaten by some in
America. Several other gentlemen engaged in the conversation.
At its close, the specimens exhibited by Mr Davidson were ex-
amined with interest.
FEBRARY 25th.
THE ANNUAL SOIREE.
The 4th Annual Soiree was held at the Royal Pavilion, the
whole suite of rooms being engaged for the occasion, and proved
none too large for the gathering of members and their friends,
which numbered between 700 and 800.
The Banqueting Room, two Drawing Rooms, and Saloon
were devoted to a variety of interesting objects, while the Music
Room was converted into a dark room in which short quarter of an
hour lectures, illustrated by the magic lantern, were given at
intervals during the evening.
The exhibition of objects of interest was not only more varied
than on former occasions, but in many cases unique. Inthe North
Drawing Room, the Sub-Wealden Exploration was brought pro-
minently under notice, Mr. Ernest Willett here presiding over cases
in which were shown a number of the latest cores extracted from
the bore hole, as also those which were brought to the surface just
prior to that unfortunate breakdown, with the particulars of which
the public had been already informed through the reports of Mr,
Henry Willett, F.G.S., the Honorary Secretary to the undertaking.
During the evening, the cores were eagerly and studiously scanned.
Close by the Sub-Wealden cores, Mr. Ernest Willett exhibited the
bones and flints which he found during his excavations at the
;
"
[ 79 ]
Cissbury pits, and respecting which he read a paper before the
members of the society.
A fine collection of shells, some of them beautifully marked,
was exhibited by Miss Glaisyer.
In the Saloon, the cases of stuffed birds lent by Mr. H. T. Booth
at once attracted attention. With respect to these, it was, as Mr.
Wonfor remarked in the course of the evening, difficult to tell
where “ Nature ended and Art began.” While the natural features
were well preserved, Art came in to aid in the accurate represen-
tation of the surroundings. The birds exhibited formed part of a
magnificent collection which Mr. Booth has formed. They bad all
been shot by himself and are, eventually, to be place in a permanent
building, which has been erected especially for their reception, on a
piece of ground abutting on the Dyke road. Every bird in the
collection is shown with something of its natural surroundings at
the time of the capture. This has been done by sketching the spot
and afterwards reproducing it by means of modelling. The
mounting is entirely artificial; but is, nevertheless, wonderfully
realistic, and this even in the most minute detail. Regard being
had merely to those cases shown in the Saloon, it might be said
truly that ornithology never found a more artistic and complete
exemplification. Whether one looked upon the owls taken at.
Chiltington, the moor hens obtained on Loch Glyn, in Ross-shire,
or the specimens of the cuckoo, the wagtail, and other members of
the feathery family taken in the vicinity of Brighton, there were
still to be seen the botanical productions peculiar to each, while the
geologist, would find it difficult to discover a defect in the character of
the strata in each case represented. Mr. Booth has made a special
study of the subject; and, no expense being spared, he has, by dint
of great labour, succeeded in getting together the unique collection
of which he is the fortunate possessor. A specimen of the great
African Albatross, together with a pair of splendid lyre-birds,
supplemented Mr. Booth’s collection. These latter were exhibited
in tbe Drawing Boom by the President of the Society (Alderman
A. H. Cox, J.P.)
In the South Drawing Room Mr F. E. Sawyer had a large
collection of meteorological instruments, weather charts, and other
appliances for the study of the atmosphere and its effects.
Apparatus for the analysis of milk, and the detection of adultera-
[ 80 ]
tions in that necessary article, as well as specimens of its affinities,
were shown by Mr. E. Moore, amongst these being some of the
artificial milk used for feeding infants during the siege of Paris.
Close by was placed the tank of the Aquarium Company. Special
arrangements as to lighting, &c., had been made for the exhibition
of its contents; and Mr. Lawler certainly had no unattractive
collection. Here was a specimen of the sea anemone—a double
dianthus—with two mouths, two sets of tentacles, but only one
body! A similar “freak of Nature” bad also happened in regard
to a baby trout, hatched in the Aquarium. Coming from one ova,
it had a veritable Siamese twin appearance; and, as it was carefully
held up to view in a glass bottle, its two heads presented a very
peculiar appearance as they converged to the centre and terminated
ina single tail. A number of tiny salmon were also shown, and
the sea horses in the tank, together with anemones,. tube worms,
zoophytes, madrepores, all gracefully exposed among rock-work and
sea-weed, furnished a noteworthy contribution.
Electrical experiments were provided by Mr J. P. Capon, who
had a powerful battery at work, and the kaleidograph, lent by Mr.
G. Nash, was a never-ceasing source of fascination. Amongst the
more general collections were skeletons of turtles, one of which
before death weighed 80lbs, exhibited by Mr Mutton; sections of
telegraphic cables, including portions of the deep sea cables of 1865
and 1866; curious opium pipe from China, from the Grammar
School Museum; a pipe fish exhibited by Mr W. Saunders, and
other curiosities.
In the Banqueting Room Mr. Dowsett exhibited three fine
cases of beetles; Messrs Wallis and Clayton exhibited an interesting
case in which were an iron Crucifix found in the ruins of Lewes
Priory ; a pipe of the time of Cromwell, antique silver sugar tongs,
spur of the time of Henry VI., dug up in the Queen’s road,
Brighton, mummy beads, mummy charms, a diminutive mummy
crocodile, fossils found in the },)ue lias of Somersetshire, lent by
Mr. C. E. Clayton, including paddles and jaws of the ichthyosaurus.
A model of the link motion of a locomotive, lent by Mr. Mutton, a
capital collection of eggs, and a portion of the jaw of the sperm
whale, some six feet long, with most formidable teeth, from the
Brighton Grammar School Museum, were also in this room. Mr.
Hubert Saunders was a large contributor, and exhibited a dangura,
L sl J
or Himalayan wood axe, and a kookeree, or knife used by the
Ghoorkas, and Arab slippers. Photographs of the natives of
Steamer Point, Aden, and of the Point itself, calabash covers from
Central Africa, flint implements from Cissbury, &c.
Spectroscopes, &e., were exhibited by Messrs. T. Rowley and
Son, as well as a photorama and cuminoscope. Telegraphic
and other apparatus were shown by Mr. Volk. ‘Tree felling
instruments in green stone, used by the natives of New Guinea,
were exhibited by Mr H. Willett. The most interesting table
in the room was that of Mr C. F. Dennet, who has devoted
himself, and with the most encouraging and gratifying results, to
the discovery of substitutes for cotton, silk, &c., and materials for
the manufacture of rope, paper, and other things. He was led to
this by the cotton famine in the last decade and by the reward
offered by the Government; the result being that several growths
which have, up to the present time, been regarded as all but useless
have been utilized and have been found to yield well. Specimens
of China grass, New Zealand flax, and other plants, in various
stages of preparation were shown, and the fact that they are now
finding a ready sale in the market is a satisfactory proof of the
success of the experiment. Hitherto the chief difficulty in dealing
with these products has been the want of a machine which should
properly separate the fibre, and such a machine Mr Dennet has just
managed to secure.
Microscopes, spectroscopes, galvanic batteries, meteorological
and other scientific instruments, were scattered throughout the
rooms.
Amongst those who presided over microscopes were Dr. Corfe,
Dr. Tuthill Massy, Dr. Hallifax, Mr. Haselwood, Mr. W. H. Smith,
Mr. Savage, Mr. Puttick, Mr. Wonfor, Mr. Ryall (Eastbourne), Mr.
Wonfor, ‘jun., Mr. J. Ridge, jun., Mr. G. D. Sawyer, Mr. C.
Hamilton, Mr. T. Glaisyer, Mr Gwatkin, Mr. H. Saunders, jun., Mr.
Lawler, Rev. W. J. Payne, Mr. B. Lomax, Mr. Mills, Mr. H. Lee
(Croydon), Mr. F. E. Sawyer, Mr. Moginie, Quekett, and Mr. T.
Curties (Royal Microscopical Society); the latter gentleman,—an
honorary member,—in addition to supplying a large collection of
tropical microscopical insects, also providing some two hundred
_ mounted objects for the use of members. The special object shown
[ 82 ]
by Mr. Wonfor was a newly-hatched salmon, and by Mr. Lee the
head of the well-known insect, the “ Daddy Long-legs,” beautifully
mounted in glycerine.
Galvanic batteries and electrical] apparatus were exhibited by
Mr. Benjamin Lomax, Mr. J. Capon, Mr. R. Glaisyer, and T. Cooper.
At half-past eight, the PRESIDENT, Mr Alderman Uox, delivered
an inaugural address in the Banqueting Room. It was a rule of
the Natural History Society that, at the annual soirée, the President
and other members should deliver short addresses. For the sake
of the assembly, as well as his own, he was glad to say that ten
minutes was the usual time allotted. He recollected the distinguished
ability of many of the gentlemen who had preceded him in the chair
which he had now the honour of occupying; and he had always
felt it to be somewhat presumptuous in his venturing to accept an
office which was pressed upon him by those who had too favourable
an opinion of his attainments. He knew that their society had
amongst its members nearly all the scientific men in Brighton.
He also knew that on these occasions they were proud of the
company of many young persons and ladies who had not made
natural history their particular pursuit. It was to those he wished
to address his few remarks; for he desired to create in their
minds an intelligent interest in the pursuits of the society and the
specimens of natural history in the museum. He was not going to
trouble his young friends with any technical phrases, but perhaps
in one of the rooms they might have observed lumps of granite.
These were called primary rocks; they were crystaline in their
character, broke like pieces of lump sugar, and when broken showed
no signs of either vegetable or animal life. They were most
durable in their character; but, like everything in creation, they
were subject to change. Jersey was built upon the primary rocks,
and at places might be seen large masses which had become
displaced, and nearer the seashore smaller pieces. As the tide rose
and fell, ebbed and flowed, those pieces were rubbed the one
against the other, and fine sand was formed. After the lapse of
ages, this sand became a sedimentary rock; they were stratified ;
‘they broke into layers, and did not break, like the primary rock,
into crystaline masses.
Some of their pavements were composed of this species of stone,
and most valuable it was for building purposes. He did not allude
{ 83 J
to the Caithness stone, some specimens of which might be seen
near the Post Office. That was mud consolidated by the lapse of
ages, and in which had lived and died myriads of creatures, the oil
from whose bodies gave it that greasy appearance so observable.
Coming toa more recent formation, it would be seen, when passing
through Clayton Tunnel towards London, they came upon a large
expanse of comparatively flatcountry. This was an estuary of thesea,
in which a mighty river once poured its flood and deposited its mud.
This extended over a large district which existed where the English
Channel now was, and could be distinctly seen on the opposite coast.
After passing through the wealden district they came to the north
down, composed, as were the south downs, of chalk. Unquestion-
ably, the north and south downs were connected, and covered the
space where the wealden clay now existed. This chalk extended to
London and beyond; and as all chalk was formed at the bottom of
the sea, it followed that where London now stood the sea once
ebbed and flowed. The hills surrounding Brighton were of chalk,
and the London and Lewes roads were the valley through which
rivers ran. Within their own time they might see some of these
changes. Thirty years ago he visited Offham chalkpits to collect
specimens. Barges for the conveyance of the burnt chalk or
lime were brought to the foot of the hill. Last summer he was
there; and the barges could not approach within half a mile of the
same place; that which was covered by water had become dry land.
Above the chalk the London clay was found. That formation
cropped up in their own neighbourhood, and in it had been found
many plants now grown in tropical regions. He had omitted any
reference to the intermediate strata. Did time allow, he would call
their attention to the carboniferous, in which coal was found—the
vegetable remains of a past time. Buckland said that, in the coal
mines of Bohemia, the most elaborate imitations of living foliage
upon the painted ceilings of Italian palaces, bore no comparison
with the beauteous profusion of extinct vegetable forms with which
the galleries of these instructive coal mines were overhung. The
roof was covered as with a canopy of gorgeous tapestry, enriched
with festoons of most graceful foliage, flung in wild irregular pro-
profusion over every portion of its surface. The effect was
heightened by the contrast of the coal black colour of these
vegetables with the light groundwork of the rock to which they
were attached. The spectator felt himself transported, as if by
[ 84 J
enchantment, into the forests of another world; he beheld
trees, of forms and characters now unknown upon the surface
of the ‘earth, presented to his senses almost in the beauty
of their primeval life; their scaly stems and _ bending
branches, with their delicate apparatus of foliage, were all spread
forth before him, little impaired by a lapse of countless ages, and
bearing faithful records of extinct systems of vegetation which
began and terminated in times of which those relics were the
infallible historians. He had attempted, he knew most feebly, to
describe those physical changes ; he would now briefly remind them
that in accord with those changes different grades of life appeared.
The earlier formations showed but a low order of either animal or
vegetable life, and it was only in the later geological epochs that
they saw the higher order of life, with its higher development and
its more exquisite sensibility to the joys and pains of existence.
Some of their kind friends had brought their microscopes, and they
would show them that the lower order of life had its organization
perfect for the circumstances in which it was placed, and the
various clusters of contrivances showed the power and wisdom of
the Great Creator. The microscope would also show that there
were other worlds beside our own, for it was well said by Channing
that, “whilst the telescope revealed a world in every star, the
microscope revealed a world in every leaf,’ and so minute were some
of the little creatures formed, that no less than 41 millions were
found in a cubic inch of tripoli, or the polishing slate. Millions
and millions of years had elapsed to produce those changes, but how
manifest through all those countless ages were shown the power
and wisdom of God in the works of His creation? One more word
and he had done. Let them not neglect to see the spectroscope.
They would see every gas and liquid examined by it showed its own
lines of colour and coloured liquids, exactly alike to the naked eye,
had a marked difference when subjected to its influence. His task
was done, and he thanked them for the attention with which they
» had listened to him; and if he could induce one person to join the
Society and feel an interest in the contents of the Museum, he
should be more than satisfied. He could assure them they would
meet with enjoyments unalloyed with regrets; and he believed they
would feel as he felt, under deep obligations to those who had
established the Society, and that their warmest thanks were due to
their excellent Hon. Secretaries, Mr. Onions and Mr. Wonfor, who
fF 85 |
had been most active in preparing this intellectual treat for their
enjoyment.
Mr. Wonror then gave a brief epitome of the contents of the
rooms, the object being to direct the attention of visitors to
particular places for particular features. Referring to the tank of
specimens lent by the Aquarium Company, he said that, although
some people out of Brighton sneered at the Aquarium, and said it
did nothing to aid science, those in Brighton knew the Company
‘did everything in their power to aid the progress of science, and,
perhaps, he might, before many months were over, tell them what
he thought the Aquarium had done for the cause of science.
An adjournment was now made to the Music Room, where,
before a large audience,
Mr BE. A. Panxuurst lectured on “The Forms of Water,” his
remarks being illustrated by a magic lantern. He proposed to
consider, for a few moments, water under the three forms in which
it presented itself to us, viz., as a gas, a liquid, and a solid, and he
would commence with the form best known to us—the liquid. Here
they had (referring to a slide exhibited on the screen) a view of a
clear, pellucid lake lying in its solitary beauty among the Sierras
of California: it was the mirror lake of the world—renowned
Yasemite Valley. How marvellously clear it was, how truly it
deserved its name, for not a feature of those hills was in the mirror
slighted. Butzit was not of its beauty that he wished to speak to
them so much as its power. It was strength in repose; it was
power potential, not active. Follow the stream that ran from it
down the valley. Here the giant was enclosed, and in this view of
the Lower Falls they beheld a picture of the work which it had
done and was still doing. For thousands of years it had been
eating away the hills, and in the photograph of the Yasemite
Valley itself—the stream running 6,000 feet beneath the peaks on
either side—they saw what a memorial it had left behind it of the
work it had done. Or if that example of the force of water drew
too much upon their imagination, let them stand on the brink of
Niagara, and gaze at the swirling, surging torrent hurl itself into
the abyss below. The world scarcely furnished a more impressive
spectacle. Whence did these lakes and rivers have their source ?
The answer was, in the rain that falls on the earth. The rain came
-
[ 86 ]
from the clouds; and the clouds, whence did they arise? These
masses of vapour were creatures of the earth, born of the land, the
rivers, and seas. They could not follow them in the passage from
earth to sky, for steam was invisible. By means of - heat
water became an invisible gas. It issued thus from the tea-
kettle, or the chimney, or the lccomotive engine. But, at a
short space from its source, it was condensed by the colder
air, and they had the dust of water as Tyndall called
it, which was somewhat incorrectly termed “steam.” It
was the heat of the sun which caused the water of the earth to
pass into this invisible gaseous form. By reason of its lightness
it rose through the heavier air till it condensed into clouds.
The canopy of clouds that so often hung above might not be very
high. It was his privilege once, during an ascent of Mont Blane, to
be above the clouds, and to see the sun rise on the other side of
them. Even there, the ubiquitous photographer had been. But
the scene, beautiful as it was, could not give them all the glories of
the reality. A sea of vapour threw its fleecy waves around the
rocks at his feet. Far as the eye could reach it stretched its
flocculent billows, and the great black peaks of the surrounding
mountains were set as islands in its midst. The sun, gradually
climbing above the horizon, flooded by degrees the depths of its
ghost-like waves with mystic light, and edged their crests with
silver. But this mass of vapour, if at a greater elevation, would not
fall in rain, but would be congealed to snow.
They had seen lately those white flakes of frozen vapour
coming down in multitudes. How many of them dreamed,
as they fell around, of the beauty of their geometry? Here
were examples of some of the “snow flowers;” they would
observe that each one had six petals, and though the forms
were so various, the number was invariable. In Brighton
the snow did not last long, for the heat soon converted it
into liquid, and as such it found its way into the sea. But on the
summits of the Alps it was not so; there the cold was too great.
Not all the snow that fell was melted, only a portion. Were it to
be piled up from year to year, in a century the Alps would have
nearly 4,000 feet of snow on them. But they would see, by the view
of Monte Rosa, that the great stormy peaks still cropped up above
them. What became of it then? It slid down the sides of the
{ 87 ]
mountains and filled up the valleys. But in doing that a great
change was effected init. They knew that, after walking in the
snow for some time, the bottom of their boots was covered, not
with snow, but with an icy mass. The pressure on their heels had
generally hardened a frozen lump there. They learned the
important fact that pressure could convert snow into ice. It was
soin the Alps. The vast superincumbent masses of snow pressed
that beneath it into the narrow valley, and so formed what was
called a glacier. It was somewhat difficult to describe a glacier,
but they might imagine a huge river winding among the hills, its
surface rugged with the vast waves which a stormy wind had raised
on it, and then imagine that, in a moment, the river was frozen.
The crests of the waves were ridges of ice. The depressions
between them ran down for hundreds of feet. Such was the great
ice river called a glacier. Amongst researches of scientific men
during recent years, few had more interest than those of Agassiz,
Forbes, and recently Tyndall, on the subject of glaciers. To the
works of Pro. Tyndall on the “ Forms of Water,” from which he
had borrowed the title of his lecture, he would refer those wishing
to know what there was to be said on snow and ice. One would
scarcely think such a mass of rigid ice moved, but it had been
proved that it moved just as a river, faster in the middle than at the
sides, faster at the point towards which it bent than that from
which it turned. In fact, a glacier did everything that a river did,
and they had often had melancholy provf of their movement.
Glaciers were the sources of rivers; the Rhine, the Rhone, the
Oregon, and others had their sources in them. The hot summer
melted them as they came down into the narrow valleys. The
water trickled through the cracks, plunged into their crevasses in
hundreds of little cataracts, and at last found a vent at the bottom
of the valley. But the Alps were not the only mountains where
there were glaciers—the stupendous peaks of the Himalayas were
surrounded with them. In the midst of their inaccessible
peaks could be discerned the glittering of the great ice rivers.
How they dug out the valleys and polished the sides of the
mountains between which they moved. How they carried
down to their termination the huge boulders which had fallen
from the mountains upon their surface during their journey,
and, when they melted, left them stranded on the plains. The
hills and valleys of Scotland and Wales still bore marks
[ 88 ]
of the great glaciers which had passed over and along them, and
the “scars of Cumberland and Westmoreland still showed where
the irresistible ice plain had passed along.
In conclusion, he wished to say a word or two more, parti-
cularly with regard to ice. Were it not so common, how
wonderful it would be to see clear liquid water turned to
brittle, crystaline, solid ice. But in England, Nature was not
seen exerting her crystalising power in all its grandeur. If
they wanted the poetry of frost they must go to Russell, Lovell, and
to Emerson, and, looking at such forms, they could well understand
the beauty that had inspired those American poets. But how was
ice builtup? The skaters, hurrying over the glossy surface, little .
thought of the wondrous forms of which every portion was built
up. Still true to her plan, Nature never varied from the hexegon.
Why that necessity so often constrained her, Science could
not answer. Often during the frost preceding Christmas
he had wished for the aid of the photographer to render permanent
the evanescent forms on the windows. ‘Those delicate graceful
forms seemed to rival Nature’s beauty of leaf and flower. They
could think for a moment of the wonderfully complex forces of
which they were built up, of the power, at once impelling and
restraining, that had drawn such a curve, as with a master’s hand,
but they could know nothing of Nature’s method in her artistic
movements. Perchance she would keep the secret always hidden,
or perhaps some day Science would throw open the door, and they
would see the manipulation of the artist, but at present they could
only admire that which was at once nature and art in one,
Mr. T. W. Wonror lectured on “The Life History of an
Insect,” also illustrated by magic lantern views.
Strictly speaking, an insect consisted of three separate and
distinct parts, head, thorax, and abdomen, and possessed six legs
and one or two pairs of wings. The stages through which it passed
were described as egg, larva, chrysalis, and perfect insect; there
were exceptions to this general law, but generally the female laid her
eggs, either in or on the food plant or animal of the future caterpillar
or grub; in some cases she had an instrument by means of which
she pierced the body or substance in which they were placed, and a
wonderful instinct came in play, when, through the stem of a plant,
L 89 |
she was able to deposit her egg in the body of some creature within
the stem. The eggs, and the number laid by an insect, varied froma
few score to some hundreds, were curiously deposited in the majority
of cases on food differing essentially from that of the mother; she
might be honey loving, while the grub was carnivorous; she
might eat nothing during her brief existence, while the
caterpillar would only eat certain plants. She must also guard
against accidents, such as the fall of the leaf and glue, in the
autumn, the eggs from which the larva would escape in the spring,
to the stem of their food plant. All this she did, and much more
besides, with what we, for want of a better name, called unerring
instinct. ' Some insects, like the flesh-fly retained the eggs, which
hatched within themselves, and laid the maggot ready to commence
its work as a scavenger ; others, like the blackbeetle, carried them
in horny cases till the young fry were ready to escape.
The eggs themselves, often very minute, were objects of great
beauty and variety, and though so small, far exceeded in marking
and ornamentation any eggs belonging to any other order of
animals. The shell consisted of three layers, an outer one, tough
and raised, a middle one of non-nucleated cells, and an inner irri-
discent one. Internally, they consisted of a yolk membrane, the
yolk substance, and the germinal vesicle. There was one part of
the egg called the micropyle, or little gate, and some affirmed the
larva eat its way out at this point, but his experience showed that
at that part of the egg to which its head was nearest, the caterpillar
escaped and made its first meal, in many cases off its egg shell.
The eggs of many bird parasites were furnished with a lid which
lifted, and in the case of some, especially the Mallee bird and
black Indian peacock, when moisture fell upon the egg, certain
leaf-like appendages folded back and protected the cover.
The next stage was the Jarva, very voracious, very destruc-
tive, and constantly changing his skin; this stage presented
great variety of markings and colours, sometimes so closely like
the food that inexperienced eyes could not detect them. Some
larvee were covered with hairs and spines, and some of these hairs
possessed the property of inflicting great pain, for their sharp
points penetrated the skin and set up a most irritating annoyance.
Some larve like those of the gnat, mosquito, or whirligig beetle,
M
[ 90 ]
lived in the water, and there, by eating decaying animal and
vegetable matter, kept our streams pure.
When the caterpillar or grub had stored up sufficient fatty
matter, it prepared to enter the next stage of its existence as a
chrysalis. Somespun cocoons, others formed retiring rooms in the
barks of trees so deftly as to defy the scrutiny of an ordinary
eye; others hung down head foremost, or lashed themselves by
silken threads to twigs and branches ; while some buried themselves
in the ground, until the time arrived to burst their bonds and
appear as the perfect insect.
Here, according to the absence or presence of wings, their
number and their nature, insects were classified. There was one
point which often startled, viz., that the great bulk of the insect world
did not in the perfect state, grow. As regarded the perfect insect
there was much of interest,—the compound eye, the antenne,
organs of some sort, but of what naturalists were not agreed, stings,
lancets, tongues,—each and all would take more time than was at
his disposal. Marvellous were their powers of finding out the food
for their young, and most marvellous was the attractive power
possessed by the females of some creatures of drawing up numbers
of the opposite sex from long distances.
A third lecture from Mr. B. Lomax, on “Spectrum Analysis,”
(illustrated by Browning’s Electric Spectroscope) was to have been
given; but, owing to a want of power in the battery which was to
have illuminated the electric lamp, Mr. Lomax had to abandon the
experimental part of his contribution to the entertainment, and
confine himself to a few remarks on the nature of the study.
The rooms throughout were pleasingly embellished with a
variety of rare plants and shrubs, lent by Messrs. Balchin and Nell ;
and, during the evening, light refreshments were dispensed by Mr.
Booth to the company at a counter in the Banqueting Room.
Marcu 11th.
ORDINARY MEETING.
MR. C. F. DENNET ON “VEGETABLE FIBRES.”
It was so much the fashion for the theologian and man of
science to refer back to pre-historic times, men, and things, as a
[ 1 ]
starting point for evidence and precedent of the verity of the
subject on which he spoke, that he feared if he did otherwise he
might be thought out of fashion, presumptuous, and lacking in
reverence for the wisdom of the past. He would, therefore, refer
back to the earliest of time-servers and workers as a fit reference
for a textile fibrous lecture. He found in an edition of the Bible
— Translated according to the ‘ Ebrew and Greeke,’and conferred
with the best translations in divers languages, with most profitable
Annotations upon all hard places, and other things of great im-
portance” (Barker, Printer to the Queen, 1599—276 years ago—
Genesis chap. iii., v. 7,)—it is written, “ Then the eyes of them both
were opened, and they knew they were naked, and they sewed figge
tree leaves together, and made themselves breeches.” This was, he
thought, about the first intimation on record of the utilization of
vegetable fibres. This edition was known amongst old bookworms
as the “ Breeches Bible.” So much by the way of preface to the
paper, which, by request, he was about to read on the anniversary-
day of the first London daily paper, 1702. On the specimens of
vegetable textile fibrons materials now before them—which
attracted so much attention at the recent soirée of the society,
he should try to make the subject to which he had devoted years
of attention, much money, and in which he had found good instrue-
tion and delight, agreeable to them in so far as the limited time at
command for preparation and his feeble talents would permit his
giving them some details respecting their origin, from the rude to
the elaborate product of nature, and he might add, art; shewing
their value and advantage to commerce, manufactures, and man-
kind—developing a new industry—and which had become a source
of wealth as well as a benefit to the whole human family.
RHEA, OR RAMIE.
What was it? A textile fibrous plant, within the last few
years introduced anew from British India to the European world,
and now grown extensively in the southern and western Pacific
States of America. The samples before them showed the whole
process progressively.—1. From the root of the plant. 2. Stalk or
stem. 3. The epidermis'and fibre freed of the wood, gum, resinous,
and glutinous matter. 4, The fibre separated and left in its purity
and strength. Finally—bleached, carded, roved, spun into thread
[ 92 ]
and yarn—woven into goods, dyed in a variety of colours. They
would also observe some specimens of Kurachee hemp, American
black hemp, flax, and jute, which had been treated in the same
way; and some hemp, which he styled cottonised, for spinning, like
cotton. This was much used as a substitute for cotton during the
horrible American war.
China-grass, now well known,in commerce, was a member of the
urtica family. Urtica nivea of Wildenow (differing widely from the
common stinging nettle, wrtiea wrens), the Callooee hemp, Kalmoi
or Ramie of Sumatra, though the fibre was very similar, and forall
practical purposes was identical, was yet a distinct species of wrtica,
obtained from the urtica utilis, urtica tenacissima of Dr. Roxburgh,
and was now making the tour of the world. Thanks to the active
age in which we lived, and the inventive genius of man, we should
soon hear of some further rapid strides in the improved treatment
of cotton and this wonderful fibre.
A French journalist, in 1869, claimed the credit of introducing
to European notice L’Ortie de la Chine, or the White Nettle of
China, for one Father Voisin, a French missionary to the Chinese
Empire, as the first person who made discovery of its practical
utility, and presented it to the attention of the Societé d’acclimata-
tion of Paris, 1844-5. It might be well to correct this impression.
It was known that the wrtica nivea (Boehmeria nivea) had been
utilised for centuries in the Indian and Chinese Empires, where it
was employed for its great strength and beauty—strength for fish-
ing nets; utility and beauty for wearing apparel. ‘“ Kankhura”
or “ Ramy,” of the Islands of Malay Peninsula, to which was given
the name of urtica tenacissima, was introduced in 1803, from
Bencoolen to Caleutta, where it was cultivated at the Botanical
Garden by Dr. Roxburgh, and a considerable quantity of this
species was brought to Great Britain by Captain Joseph Cotton,
The Society for the Encouragement of Arts, Manufactures, and
Commerce, awarded a silver medal to Captain Cotton for its intro-
duction to their notice. The fibre was tested and pronounced of
practical value, but fell into disuse for the want of proper appliances
for manipulating and preparing it for manufacturing purposes.
Cotton, too, at that time was plentiful and cheap, and the cost of
importing the fibre from Jndia told against it.
[ 93 J
The late American war caused a famine in the cotton industrial
worlds, and a want of that staple article, and all other texile fibrous
materials, for the spinner and paper maker, either to be worked
alone oras an admixture with wool, cotton, silk, hemp, flax, &c.,
&c., caused public attention to be turned to anything and everything
which could be produced and made available for either of the uses
mentioned. The fibrous substance now known in commerce are :—
Rhea or ramie, or China grass, Japanese grass, Jute, New Zealand
flax, Bamboo, Aloe, Yucca, Espartero or Spanish grass, Algerian,
Egyptian diss grass, the various palms and canes, the agave or sisal
hemp, oat straw, wheat straw, broom corn, corn-husks, hop vine, &e.,
without saying anything of the utilisation of wood pulp (for paper),
which came prominently into notice, and commanded high prices.
Scientific men and mechanical geniuses taxed their brains and skill
to produce appliances, machinery, or a process to speedily manipu-
late and utilise the best, as well as meanest, of nature’s products,
and to make them available for the use of man :—at first the want
of knowledge of the chemistry of the plant; second, the use and
abuse in the employment of chemicals of the right sort, and in pro-
portionate quantities to a given quantity of material in cleansing
and bleaching, caused much provocation, discouragement, and loss,
Fibres are sensitive, and have an affinity for that whichis bad, like iron,
4 which, in course of treatment, takes to sulphur. It had recently
been truthfully stated that cotton, “if it is saturated with chromic
acid, or with permanganate of potash, and subsequently washed,
although the fibre presents no apparent change, it is found to be
seriously weakened when treated with any alkali.” They had seen
tons of the best flax and hemp injured by unskilled men. Whilst
a resident in France, he learned that the public documents of 1830
in the archives of the French Government were going to destruction,
all arising from the improper use of poor material and the more
imprudent use of chemicals. Mr. Plimsoll might find that the occa-
sion of the loss of more than one good vessel per annum was from
the use of cheap canvas, used-up duck and cordage, the fibre of which
had been partially destroyed by acids. He had to return to his
subject. Success crowned the efforts of some enterprising spirits
—men of great enterprise—whom he could name, and the result
had proved that vegetables fibres had been levelled up from the
lust immensely, and now held their own as staple articles of
é
[ 9% ]
commerce in the great markets of the world, were quoted day by
day, like cotton, flax, hemp, or any of the essentials of life.
For instance, Espartero or Spanish grass, which in 1850 was
little known, and considered of doubtful character, a drug in the
market at 50s. the ton, now commanded eleven or twelve guineas,
if to be had at all. Two hundred thousand tons would probably be
a low estimate for the importation and consumption of “Sparta
Grass” for paper making in 1873-4, On the New World side of
the ocean, the ordinary broom-corn (of carpet brooms) which used
to be plentiful at three to five guineas per ton, was now contracted
for in advance of its growth, at £12 to £15.
If they ran their eye down the Mincing lane produce sales of
almost any day, they would find 1,000, 2,000, or 3,000 bales “sold
on the spot” at prices ranging from £15, £17, £20. Some marks
and brands of jute, the product of good growers, excelling in
quaiity, like Major Hallett’s cereals, commanded premium rates in
advance of arrival. There was no end to the uses to which this
useful fibre was applicable. He had seen thousands and thousands
of bales landed at Boulogne from London to go inland to the
French beehives of industry.
New Zealand flax, Phormiwm Tenax had not made the headway
that several other fibres had. It grew abundantly in all the
districts, and in one place, wild in the woods; the Colony being so
far distant and in a primitive state, they had not given the atten-
tion to it they ought. To get proper machines to manipulate and
render it supple and the readier for manufacturing purposes, Dr.
Featherston, the Agent-General for New Zealand, in London, had
recently furnished his confreres with a quantity, and it was hoped
they would soon have a happy surprise for them, in machines which
would do the work as it ought to be done, and thereby revive a
demand for and use of a very superior fibre for many purposes.
He was sorry to tell them, and it might be no news, that with
so much of good alloted us, also came much of evil; good and evil
ought not to go hand in hand together, but so it was; there were
adulterations in textile manufactured goods, and the officials at
Somerset House ought to have an eye to the adulterators as well as
upon the grocer, baker, and other tradesmen. It required longer
and nicer calculation to produce refined shoddy goods for wearing
[ % J
apparel to cover our bodies than to make wooden nutmegs to tickle
our palates. Look at the glowing announcement (samples
handed round) setting forth “as a well-known fabric, being all wool
weft, and well woven, the shades dyed to suit the prevailing fashion
__ at the leading watering places, and being ingrain colours, they are
perfectly fast.” A chief recommendation seemed to be they were
5 perfectly fast colours! Perhaps the vendor thought his goods
adapted to this watering place on that account? They now had
cheap enough articles. But how good? Paper made of the
meanest old worn out, washed out materials charged with kaoline
or pipe clay and sizing to give it weight. Newspapers for a half-
penny so rotten they would hardly hold together during one
reading and scarcely bear a short mail route. Trousers made of
shoddy and refuse jute yarns—“ guaranteed all pure wool” for 13s,
Japanese silks, full length and width, twelve yards for 12s. 6d.
Silk-rep, “ warranted all pure silk and wool,’—only demi-rep, well
dyed shoddy and jute. Again— Your attention is invited to our
warranted all pure silk stuffs at 1s. 6d. per yard””—which was all
stuff !—puzzling both seller and buyer to show where the silk came
_ in; and in weft, warp, or woof, in most of the very attractive fast-
coloured fashionable goods now filling the shop windows in every
large city or town in the kingdom or on the continent. He believed
_ the continentals had much to answer for; the neighbouring streets
of Bowbells were filled with German and French manufacturers,
agents taking orders by samples, competing with the British
manufacturer—all making haste to be rich.
7
ay eT es
é
THE TWENTY-THIRD
ANNUAL REPORT
"ABSTRACT OF PROCEEDINGS,
» OF THE
| BRIGHTON AND SUSSEX
NATURAL HISTORY SOCIETY,
ON THURSDAY, OCTOBER 12tu, 1876,
PRICE ONE SHILLING AND SIXPENCE.’
- BRIGHTON :
‘PRINTED BY FLEET AND BISHOP, “HERALD” OFFICE.
1876,
‘
resident
MR. G. D. SAWYER.
Vice-Hresidents :
‘MR. HOLLIS, DR. HALL,
MR. BIGGE, MR. T. GLAISYER,
. HALLIFAX, MR. F. MERRIFIELD,
SIMONDS, MR. HASELWOOD,
. PENLEY, MR. T. B. HORNE,
. GWATKIN, MR. ALDERMAN COX,
. W. E. C. NOURSE, MR, J. DENNANT.
Greasurer :
MR. THOMAS GLAISYER,
12, North Street.
; Committee :
. BENJAMIN LOMAX, MR. J. WILLS, |
. DENNET, MR. ALDERMAN MAYALL,
E. H. WILLETT, MR. A. DOWSETT.
; Honorary Secretaries :
[R. T. W. WONFOR, MR. J. C. ONIONS,
38, Buckingham Place. 56, Middle Street.
Honorary Librarian and @urator:
MR. ROBERT GLAISYER,
The Dispensary, Queen’s Road.
At the Twenty-third Annual Meeting of the BRIGHTON AND
SUSSEX NATURAL HrisTory SOCIETY, held in the Curator’s Room,
Free Library and Museum, Church-street, October 12th, 1876,
IT WAS RESOLVED,-—
That the Report, Abstract of Proceedings, and Treasurer’s Account,
now brought in, be received, adopted, and entered on the minutes, and
printed for circulation as usual.
That the cordial thanks of this Meeting be given to the Honorary
Secretaries, Honorary Treasurer, and Honorary Librarian, for their
labours in preparing the same.
That the following gentlemen be elected as Officers of the Society
for the ensuing year :—President: Mr. G. D. Sawyer ; Treasurer :
Mr. Thomas Glaisyer; Committee: Mr. Benjamin Lomax, Mr.
Dennet, Mr. Ernest H. Willett, Mr. J. Wills, Mr. Alderman Mayall,
and Mr. A. Dowsett. Honorary Secretaries: Mr. T. W. Wonfor and
Mr. J. C. Onions; Honorary Librarian and Curator, Mr. R. Glaisyer.
That the sincere thanks of this Meeting be given to the Vice-
Presidents, Treasurer, Committee, Honorary Secretaries, and
Honorary Librarian, for their services during the past year.
(Signed) JOHN DENNANT,
Chairman.
IT WAS ALSO RESOLVED,—
That the warmest thanks of this Meeting be presented to Mr. J.
Dennant for his able conduct as President during the past year.
REPORT.
In presenting the Twenty-third Annual Report your Committee
have the pleasure of again recording the continued prosperity of
the Society. They deeply regret the loss which the Society has
sustained by the death of two of its Vice-Presidents, viz., Mr. T-
_ Bi. Hennah and Sir J. Cordy Burrows ; the former was one of the
~ earliest Members of the Society, and had always taken a very
active part in promoting its prosperity, and in the diffusion
of scientific knowledge, particularly in those branches of science
a
which depend upon the use of the microscope.
The state of the finances continues to be satisfactory ; the sum
of £4 18s. 4d. remains in the hands of the Treasurer, after
; expending 428 16s. 3d. in the purchase of new kooks and
. periodicals. The number of members has slightly increased during
3 the year. It is also gratifying to the Committee to report that
the Society is becoming much more widely known ; the number of
% similar Societies in correspondence with it, both in this country
and America, has considerably increased.
~
Presentations of the following Books have been received by.
6
the Granitic, Granitoid, and Associated Metamorphic Rocks,
three pamphlets (by J. C. Ward) ; On the Outline of the Flora of
Sussex (by W. B. Hemsley); On Turf Carvings (by Dr. J.
Stevens) ; A Lecture on the Geology of Croydon (by J. Morris) ;
two pamphlets on the Chalk Cliffs of Eastbourne, and of Dorset
and Devon, by W. Whitaker (from their Authors) ; Observations on
the Genus Unio, 7 vols. (by Dr. J. Lea); Materieux pour la
Paleontologie Suisse, by T. J. Pictet, 2 vols. (from Mr. T. David-
son); The Human Race, by L. Figuier (from Mr. C. Paine) ;
Annals of Philosophy, by Thomson and Phillips, 28 vols. (from Mr.
J. H. Glaisyer) ; Two Addresses before the Geological Society of
London, by J. Evans ; Quarterly Journal of Geological Society for
1871; Free Trade (by Cobden Club); Reforme Economique
(1875, 1876); The Coinage of the Ancient Britons (by J. Evans) ;On
Siliceo-fibrous Sponges (by J. S. Bowerbank) ; Geological Record for
1874 (by W. Whitaker) ; On Landslips, by Conybeare and Dawson
(from Mr H. Willett) ; Report of Smithsonian Institution for 1874 ;
on the Zapus Hudsonius and White Tailed Ptarmigan (by Dr. E.
Coues); 2nd Vol. Crustaceous Vertebrata (by E. D. Cope);
Geological Survey of Colorado for 1874 (by F. V. Hayden) ;
Invertebrate Palaeontology (by F. B. Meek); Geometrid Moths,
by A. S. Packard (from F. V. Hayden, Geologist in charge,
Washington, D.C., U.S.A.; Nebraska (by E. A. Curley) and a
Map of the Line (from the Union Pacific Railway Company,
U.S.A.); Memoirs of Peabody Academy of Science; Sixth Annual
Report of ditto ; Check Lists of Ferns of North America, 2 vols. ;
American Naturalists (from Peabody Academy of Science, Salem,
Mass., U.S.A.) ; Annual Reports for 1873, 1874, 1875, of Belfast
Naturalists’ Field Club; Annual Report for 1876 of North
7
Staffordshire Naturalists’ Field Club ; Annual Report for 1875-6
of Chester Society of Natural Science ; Four Parts of Transactions
for 1875-6 of Watford Natural History Society ; Annual Report
and Parts of Journal of Quekett Microscopical Club; Annual
Report for 1875 and Five Numbers of Proceedings of Geologists’
Association ; Two Parts of Proceedings of Natural History Society
of Glasgow ; Annual Reports (10, 11, and 12) of Lewes and East
Sussex Natural History Society ; Annual Reports for 1875-6 of
Eastbourne Natural History Society; Annual Report for 1875
and other Papers of Leeds N aturalists’ Club ; Report for 1875 of
Brighton Free Library and Museum ; Annual Report for 1875-6
of Chichester and West Sussex Natural History Society Four-
teenth Quarterly Report and other Papers of Sub-Wealden Explo-
ration ; Report of Cardiff Naturalists’ Society (from the Secretaries
_ of the Societies).
The following Books have been purchased during the year,
viz. :—Mental Physiology (by Dr. W. B. Carpenter) ; The Octopus
(by H. Lee) ; History of Ferns (by J. Smith) ; Vol. t., Parts 1, 2, 3,
Botany, and Vol. 1., Parts 1, 2, 3, Zoology, and Index to Vols.
26 to 30 of Linnean Society ; Jummoo and Cashmir Districts
(by F. Drew); Geology for Students (by A. H. Green) ;
Animal Parasites (by Van Benenden); Contributions to
Molecular Physics (by J. Tyndall); Climbing Plants (by C.
Darwin) ; Fermentation (by P. Schiitzenberger) ; Notes on Col-
lecting and Preserving Objects (by J. E. Taylor) ; Geology of
England and Wales (by H. B. Woodward) ; The Moon (by E.
Neison); Geographical Distribution of Animals (by A. R. Wallace).
The Geology of Surrey, Petrifactions, Catalogue of Organic
Remains of Sussex; Osteology of the Iguanodon and Hylaeo-
saurus; Fossils; Reptiles of South East of England (by G. Mantell).
The number of the Pooks is now 917, exclusive of unbound
current Periodicals and Pamphlets. The Committee are pleased
to report that the use of books, both by Members and by the
Public in the Reading Room of the Free Library, continues to
increase.
As a new Catalogue was required, the system of arrangement
has been altered, the classification being by subjects, with a list of
the Authors at the end, which plan it is considered will render the
reference to works on a particular subject more convenient.
Members will be able to obtain the new Catalogues on application.
A donation of seven slides, illustrating different varieties of
starch and arrowroot, has been presented to the Society’s Micro-
scopical Cabinet, by Mr. W. H. Smith, and also six slides of
Polycystina, by Mr. Haselwood.
The Conversazione, held on the 29th February, at the Royal
Pavilion, was very successful, a detailed account of it will be found
in the proceedings.
The thanks of the Society are due to those gentlemen who
have read Papers, exhibited objects, or presented specimens or
slides for the cabinet, or photographs or drawings for the Society’s
album.
The Field Excursions since the last Report have been as
follows :—September, 1875, Hayward’s Heath ; October, Worthing ;
4 9
May, 1876, Balcombe ; June, Isfield ; July, Lancing ; August,
i “Steyning ; September, Berwick.
The Annual Excursion took place on the 6th of July to
Petworth.
In concluding their Report, your Committee beg to request’
i. _ the Members to endeavour to promote the prosperity of the Society
4 by bringing its merits under the notice of their friends, by con-
a tributions of works of Natural History to the Library of photo-
# graphs or drawings of objects of Natural History for the album,
and of slides for the Microscopical Cabinet, and particularly by
4 reading ‘Papers during the ensuing year.
ABSTRACT OF PROCEEDINGS.
1875-6.
SEPTEMBER 9QTH.
ANNUAL MEETING.--MR. C. POTTER ON “THE
SO-CALLED FOREST BEDS.”
Mr. DENNANT, the President, returned thanks for his election,
and expressed a hope that the progress of the Society during the year
‘just begun would be very real, and that the efforts of the Members
would end in even greater usefulness than hitherto. He also spoke
briefly in favour of very short papers by Members who felt some
diffidence in reading long ones at their ordinary meetings; and
expressed the pleasure he felt in being associated on the Committee
with their two able Hon. Secretaries, than whom they could not find
better were they to search the country through.
Mr. POTTER explained that what he should say would refer
principally to the Cheshire shore, not so much because the beds were
confined to that coast, as because they were seen there to greater
advantage than in any other part of the country. At one time he
believed that the whole of the forestal remains so largely exposed on
the Cheshire shore, and to be found in corresponding beds in nearly
the whole of the marsh lands and river valleys of Great Britain,
Ireland, and in many parts of the North of Europe, germinated, grew,
and decayed, where they were now found; but having accidentally
come upon a small unresinous fir in an undecayed state, underlying
the bole of a large.oak, both of which were firmly embedded in their
surrounding matrix, a doubt at once arose in his mind whether his
II
preconceived opinions were correct. Subsequently, he ascertained that
seldom were two of these trunks to be found lying parailel to each
other, but upon and crossing each other, often at right angles. He
also observed that the roots of the standing butts were as likely to
spread out above these prostrate trunks as below them, and further,
that the standing butts which were embedded in the lower part were
not more than as one to ten of the trunks.
As previous observers appeared now to have noticed these facts,
he determined to examine into and satisfy himself on the subject.
These investigations caused him to change his original opinion ; he
concluded that the whole of the forestal remains found in these beds
drifted from elsewhere. Under any circumstances, the tree-trunks
and boughs must have been covered up where they were now found
by the natural growth and deposition of ee fresh water plants: of
which these beds were formed.
Was it possible to account for these different conditions of sound-
ness, if the fact were excluded that one must have been more decayed
than the other previous to embedding? Strong evidence that the
trees exposed on the surface had not grown and been broken down
where they were now found was afforded by the fact, that of the
hundreds of trees exposed, it was almost impossible to find a prostrate
trunk in such a position near a standing butt as would suggest the
idea that they might have formed one entire tree. Nevertheless, the
smallest roots and branches of these trees were, with their bark and
epidermis, in the most perfect condition preserved in these beds ; and
it should be borne in mind by those who argued that these trees had
lived and died where they were found, that such portions as had
grown above the earth, including trunks, branches, &c., would lie
exposed to the decaying influence of air, moisture, and insects for an
indefinite period of time before they could, from the natural growth
of vegetation from below, and the annual fall of the leaves, &c., from
above, become covered and buried by a vegetable soil.
The tree butts had sunk through water to where they were found,
and the materials of which the beds were made up was gradually
formed by deposition. To his mind one fact was in itself conclusive,
that these trees could not have grown where they were now seen,
The peats being stratified and very finely laminated would give proof
or evidence of the intrusion of a foreign body, and he hesitated not to
assert that it would be impossible for a root in its first stage of growth
12
as a delicate filament to expand with the growth of the tree until it
had attained a diameter of eighteen inches or more and not dispense
with or in any way crush the laminz or stratification above or below.
He believed that it was impossible for a tree, under any circumstances,
to fall where it had grown, and there remain in a comparatively sound
state, whilst a soil derived from the ordinary growth and decay of
vegetation accumulated over it.
SEPTEMBER 23RD.
MICROSCOPICAL MEETING.—“ POND LIFE.”
The President, Mr. J. DENNANT, in the absence of any paper on
the subject of the evening, “ Pond Life,” invited those present who had
brought objects of interest to give an account of them.
Mr. T. W. WONFOR said he had purposed obtaining some
specimens from Lewes, but the weather had been so unfavourable that
he had not been able to do so. He had, however, procured froma
pond on the Furze Hill—which generally proved a fertile source of
supply—what proved, on microscopical examination, to be one of a
very interesting class of creatures, /i/usoria, so-called from the forms
of life found in various infusions. All /zfusorta were at one time sup-
posed to be animals, and Ehrenburg, the great German microscopist,
had taken a red spot which he called the “eye dot” as a mark of their
being so. Many had since been proved to be the early stages of
plants. This particular specimen was known by the name of exg/ena,
and the “ eye dot” was plainly to be seen. Its shape was cylindrical,
and the green nuclei might be observed circulating in its body. Its
form changed as it moved about, and one that he noticed appeared to
have absorbed some particle of some other animal for food. There
was not much doubt that it was a true though very low type of
an animal. The same -pond furnished plenty of cyclops, larve of
gnats, and many vegetable forms.
Mr. T. GLAISYER stated that he had received from a friend an
object picked up from the surface of the sea, some 300 miles from
Singapore, which he had ascertained to be a portion of a vegetable
structure,
!
13
Mr. HASELWOOD observed that he had brought some forms of
life from Hayward’s Heath. From the same place he had carried
away some newts, but they had unfortunately died.
The Meeting then resolved itself into a microscopical one, at
which were exhibited, in addition to the objects already mentioned by
the Members, the following :—
Mr. DENNANT, the President, diatoms and desmids from the
Shoreham marshes ;
Mr. HASELWOOD, a cadis worm and some daphnia ;
Mr. PUTTICK, cyclops, rotiferze, and minute plants ;
Mr, R. GLAISYER, water fleas and wheel bearing animalcuze.
OcToBER 14th.
ORDINARY MEETING.—MR. T. WwW. WONFOR
ON “MANNA.”
“Tn the morning the dew lay round about the host, And when
the dew that lay was gone up, behold, upon the face of the wilderness
there lay a small round thing as small as the hoar frost on the ground.
And when the children of Israel saw it, they said one to another, It
is Manna ; for they wist not what it was.” Over this word manna,
_ and what it was, there had been great differences of opinion. He
_ did not purpose entering into any question of a controversial character,
nor yet to raise any doubt respecting the miraculous production of .
food for the Israelites during their wanderings in the desert ; but to
indicate first what natural productions had at different times been
_ considered to approach in resemblance the nearest to the Scripture
food, and then to show that even now a substance was found, a
_hatural growth, which corresponded with the description given in the
Mosaic account.
But first there was a difference of opinion in regard to the term
Manna; some asserted that, as the word man means ‘‘ what is it?” it
derived its name from the exclamation of surprise uttered when the
_ substance was first seen. It has been urged against this derivation
that the use of the term Manna in the New Testament as a substance
14
militated against this derivation ; but this seemed of little account, for
though a mere exclamation in the first instance, by usage it became a
term applied to a material substance. Others derived the name from
Manah, a gift, others from Minah, to prepare, and others from Munahon,
“provision for a journey.” It seemed to him that the first was the
simplest and most natural derivation, and agreed with the evident
surprise with which the Israelites first saw the (to them) new food.
Those who had written about Manna divided themselves, as it
were, into two groups, viz., those who took it literally to be a substance,
which fell like dew from heaven, z.2., the atmosphere, and those who
considered it a vegetable production or a plant itself’ Among those
who affirmed that the Manna of Holy Writ was a species of con-
densed dew or honey, they had Salmasius, who said that the Manna
of the Israelites was only a species of honey, identical in its nature
with wild honey, which supplied food to St. John in the wilderness
and that the miracle did not consist in the production of any new
substance, but in the abundance and in the regularity in which it was
dispensed for the maintenance of so vast a host.
The idea of Salmasius was sustained by Atlian in his work, De
Natura Animalium, where he described a natural phenomenon in
India. ‘In India, and particularly in the country of the Prasii
(who extended through the richest part of India, from the Ganges
to the Punjaub), it rains liquid honey, which, falling on the
grass and leaves of reed, produces wonderfully rich pasture for
sheep and oxen ; the cattle are driven by the herdsmen to the
spots where they know quantities of this sweet dew have fallen.
The animals enjoy a rich banquet on these pastures, and furnish very
sweet milk. There is no necessity to mix it with honey as the Greeks
do.” Athenzus, quoting from Amyntas, who wrote an Indian
itinerary, says, Amyntas, in his first book, speaking of the honey from
the atmosphere, writes thus : “ They collect it with the leaves, making
it into the form of a Syrian cake ; some make it into the form of a ball ;
and when they wish to enjoy it, breaking off a portion, they melt it in
wooden cups called tabetiz, and after they have passed it through a
sieve, drink it. It is much like diluted honey, though somewhat
sweeter.”
In Italy, especially in Calabria, during periods of excessive heat,
drops of a honey-like substance fall to the ground, and are called by
the inhabitants manna. This substance, of a sweetish glutinous
15
flavour, suddenly appearing on the leaves of plants, was asserted to
stop their growth and act injuriously on the plants. Such leaves were
spoken of as “ Foglie ammanate” (leaves affected by manna). They
used a similar term of grapes which have acquired a peculiar flavour
when covered with the same substance as “ Vino ammanato.”
He would take next the vegetable productions described as
Manna. On this latter subject there were many writers, but one of
them, Buckhardt, in “ Notes on the Bedouins and Waliabys,” when
speaking of Wady-el-Shiekh, to the north of Mt. Serbel, says, “ In
many parts it was thickly overgrown with the tamarisk or farfa. It is
from this ¢avfa that the manna is obtained ; and it is very strange that
the fact should have remained unknown in Europe till M. Seetzin
mentioned it in a brief notice of his tour to Sinai, published in the
Mines del Orient. This substance is called by the Arabs Mann, and
accurately resembles the description of the Manna given in Scripture.
In the month of June it drops from the thorns of the tamarisk upon
the fallen twigs, leaves, and thorns, which always cover the ground
beneath the tree in the natural.state ; the Manna is collected before
sunrise, when it is coagulated, but it dissolves as soon as the sun shines
upon it. The Arabs clear away the leaves, dirt, &c., which adhere to
it, boil it, strain it through a coarse piece of cloth, and put it into
leathern skins ; in this way they preserve it till the following year, and
use it, as they do honey, to pour over their unleavened bread, or to dip
their bread into. I could not learn that they ever made it into cakes
or loaves. The Manna is found only in years when copious rains
have fallen ; sometimes it is not produced at all. I saw none of it
among the Arabs, but I obtained a piece of last year’s produce at the
Convent, where, having been kept in the cool shade and moderate
temperature of that place, it had become quite solid, and formed a
small cake ; it became soft when kept some time in the hand, if placed
in the sun for five minutes ; but when restored to a cool place it
became solid again in a quarter of an hour. In the season at which
the Arabs gather it, it never acquires that degree of hardness which
allows of it being pounded, as the Israelites are said to have done
(Numb. xi. 8.) The colour is dirty yellow, and the piece which I saw
was still mixed with the tamarisk leaves ; its taste is agreeable, some-
what aromatic, and as sweet as honey.”
This ann, or manna, which was said to drop from the Tamarix
mannifera, was said by some not to exude from the tree, but to be
16
formed by an insect which abounded on the tamarisk. Others affirmed
that an insect, a coccus, punctured the tree, that the juice exuded and
dropped on the ground as described, and that the dissolving after sun-
rise was simply the evaporating of the fluid part of the exudation,
There was another substance known by the name of manna, and com-
monly employed in medicine. This was merely a sweet condensed
juice of certain plants, but especially obtained from an ash tree,
fraxinius rotundifolia, a tree indigenous to Italy and the south of
Europe, and found growing abundantly without culture in Calabria,
where collecting manna was a regular trade, commenced about the end
of July.
The gatherers of manna made a horizontal cut in the trunk of the
tree, and on the following day a second cut was made, into which the
point of amaple leaf was fixed, while the stalk part was placed in the
first slit, thus forming a kind of cup to receive the exuding juice.
Sometimes they applied thin straws to the incisions, or pieces of twig,
on which the manna ran as it exuded, and so formed tubular pieces,
which were called Manna in Cannoli, and fetched a higher price than
that collected on leaves. The season for collecting manna was over by
the end of August. At one time, manna from Syria was in request,
but the Calabrian was thought better. In Mexico, a manna was
obtained which seemed to take the place of cheese, while in other
places a manna was eaten much in the same way as honey. There
seemed to be some relationship between these so-called mannas and
some of the sugars obtained by making incisions in the trunks of trees,
notably in the case of sugar maple. It was a well-known fact that the
juices of many trees and plants when made to exude, in a somewhat
similar way, did on the evaporation of the watery particles produce
substances akin to but differing from manna in their qualities and
properties.
In the year 1849, Mr. Giles Munby read a paper before the British
Association at Birmingham, on the “ Botanical Productions of the
Kingdom of Algiers,’ in which he called attention to a lichen,
L. Esculentus or Lecanoru esculenta, as it was now named, which he
considered agreed more nearly than any other substance hitherto dis-
covered with the description of the manna on which the Israelites fed
during their wanderings in the desert. He mentioned that this lichen,
which was found on the sands of the desert, sprung up during the night
much in the same way as mushrooms, That during an expedition to
17
the south of Constantine, the French soldiers made bread of it and
cooked it in various ways, actually subsisting on it for several days. In
. Science Gossip for 1872, at page 60, was given a description of this
: curious plant, specimens of which had been presented to the Museum
| by the gentleman, T. B. W., who furnished the material for that article
; and articles in subsequent numbers, 1872, p. 186; 1873, p. 118 ; and
; 1875, p- 146. From these it appeared Pallas figured and described
. this plant in 1776, and said “it occurs in the very driest limestone of
7 the Tartarian desert, scarcely distinguishable from small stones.
except by the expert.”
The specimens before them were collected at Reboud Djelfa, in
the desert of the Great Atlas Chain, and were about one-fifth the size
of those figured by Pallas. Berkeley mentioned that, lying loose on
the ground without any attachment, it was easily rolled along by the
wind, and sometimes piled together in layers several inches thick, and
that at times being carried up by whirlwinds, it was showered down on
the ground. One such shower fell at Erzeroun during a time of great
scarcity, and afforded very opportune relief to the inhabitants.
Lindley, in his Vegetable Botany, speaking of Z. Esculentus, says “ it
sometimes appears in immense quantities in Persia, Armenia, and
_ Tartary, where they are devoured by the natives, who fancy that they.
must fall from heaven, not knowing how to account for the prodigious
numbers of the plants, of the origin of which they are ignorant.”
Parrot says that “in some districts of Persia they cover the ground to
the depth of five or six inches,” and Eversman, who had an opportunity
_ of studying it, was convinced that even in its earliest stages, the plant
had not the slightest attachment to a grain of sand.
Another fact in connection with Z. Esculenta and L. Affinis was,
that, instead of being produced at uncertain intervals and only during
a few months, it was produced during the whole year. It appeared
_ from an article by Dr. Ruke, reprinted in the Pharmaceutical Journal
for September, 1860, that Z. Esciwlenfa was presented to the Academy
of Sciences in 1828. The lichen was described as being of a fawn
colour, granulated, composed of broken crusts, which had fallen in the
neighbourhood of Mount Ararat, and which a Russian Generai of the
; Persian army had given to M. Thénard, who presented it to the
Academy. It seemed this lichen dried up during the summer in the
_ Mountains, and was transported by the winds to great distances ; this
18
led the inhabitants to say the grain had fallen from heaven. Sucha
shower fell in 1845, in the Crimea, and covered the ground to the
depth of three or four inches, and was used as food by the inhabitants
for several days.
Wherever this lichen occurred in any quantity men and cattle eat
it. In Asia Minor also it soinetimes formed beds several inches thick,
on which not only the sheep were nourished, but a species of bread
was made from it, and consumed by the poor, who regarded it as true
manna sent from Heaven. There was one fact in connection with
lichens generally, all contained nitrogeneous matters and starch, and
many kinds were used as food.
It was often argued, and especially by a writer in the Dictionary
of the Bible, that the manna of Scripture was wholly miraculous, and
not in any respect a product of Nature. All the plants mentioned
did produce the so-called manna for only a few months : but these
lichens might be gathered all the year round. The lichens again were
produced in large quantities, but not, as he states, at the rate of
15,000,000lbs. a week, which quantity might, by Divine agency,
assuming a lichen to have been the manna of Scripture, have been
produced by increasing the growth without depriving the plants of
their character as natural products.
Another difficulty which had puzzled many was the fact that the
manna collected on the sixth day “did not stink” on the Sabbath,
though that collected on other days and kept over did; but it seemed
the injunction, “ Bake that ye will bake zo-day, and seethe that ye will
seethe,” showed how by the process of cooking decomposition was
arrested. He could quite understand that this very lichen, gathered
in large quantities while the dew was on it, and heaped up damp and
moist, would decompose, ferment, and deteriorate. The fact that the
Israelites were supplied with manna for forty years, that a double
quantity fellon the sixth day, and that it ceased when their wandering
ended, placed its production among the miracles of Divine ageney
without taking away from it the character of being a natural pro-
duction, created only when first seen by the children of Israel, and
no longer existing as a part of the scheme of the life of the globe.
Of all the substances hitherto assumed to be the manna of
Scripture, this Z. Esculenta seemed to approach the nearest, in its
being produced all the year round, being round and hard, able to be
19
pounded and converted into bread, appearing in large quantities, as if
from Heaven, while all the others were found only at certain seasons,
and did not in so many respects correspond with that described in
Holy Writ.
The President, Mr. J. DENNANT, proposed a vote of thanks for
the paper, and, in inviting discussion, said that neither religion nor
politics were allowed to be discussed.
Mr. G. D. SAWYER said that thouch religion was not to be dis-
cussed, the Scriptures had been appealed to, and he should like to ask
how, if they accepted the fact that the manna could not be found on
the seventh day, they could still believe that it was a natural produc-
tion. To his mind it was a convincing proof, if they accepted the
fact that it was not found on the seventh day, that it was not a natural
production.
Mr. WONFOR replied by asking Mr Sawyer if he ever went out to
gather mushrooms, because if he had, he must have noticed, that
where none could be found one morning, there would be abundance
the next.
Mr. SAWYER : Every seventh day ?
Mr. WoNFOR said he did not intend to lead to questions of that
kind. It would be equally a miracle to produce a natural substance
in more than its natural abundance, and for longer than its natural
period, as to create an entirely new substance. In the chapter that
contained the account of the manna, was also an account of that
wonderful production of quails. That was one reason why, with all
deference to the opinions of others, and with the most reverent
feeling, he suggested the possibility of a substance already in existence
being produced in double quantity.
Mr. G. D. SAWYER mentioned it with no feeling, but as in a
scientific discussion. He could not take it to bea natural substance
if it could not be found.
Mr. WonFror: It was only recorded that they went out one
Sabbath day and could not find it. It did not say that they ever
“went out again and could not find it.
Mr. C. F. DENNET expressed dies great pleasure he had felt in
: listening to the paper, especially after the proceedings of the week at
the Social Science Congress, It was an agreeable purging of their
20
minds of the intellectual stuff that had been crammed into their
mouths ; and he felt specially delighted, and thanked Mr. Wonfor for
the information he had afforded on the subject.
Mr. DowsETT pointed out that the manna previously found could
not have been the manna eaten by the children of Israel, inasmuch
as it was not nutritious ; and was also slightly purgative.
NOVEMBER 4TH.
ORDINARY MEETING.—DR. HALLIFAX ON “THE
NERVOUS SYSTEM.”
In introducing to the Members for their discussion “The Nervous
System and its Functions,” he would remark that, although the subject
might appear almost a professional one, it was, to his mind, one which
was of great importance to every student of scientific matters.
The knowledge of early physiologists had been very vague and
indefinite with regard to the nervous system, and it had only been
within the last hundred years that any great advancement had been
made on the general observations of those who had in earlier ages
bestowed their attention to the system of nerves with which all the
members of the animal kingdom were endowed, or that the knowledge
that had been gained had been placed before the public in the manner
which it merited.
To Dr. Bell, among others, who had thrown much light on the
functions of the nerves, the greatest praise must be bestowed. But,
though he and other physiologists who had since devoted their time
and talents to the study of this particular branch of physiology had
laid such great results before the scientific students of modern times,
there were yet many problems to be solved, and much to interest the
general observer. The nerves which proceeded from the cerebro-
spinal axis, as it was named, in all the higher forms of animals formed
the wonderful apparatus which by a rare development presented us
with the glorious phenomena of the great minds of a Shakespeare, a
Bacon, or a Newton. ;
2I
The spine was the seat of the motive power of the many passions
which we possessed, some good enough to obtain the approval of
Heaven, and others bad enough to have their origin in “another
place.” These passions, impulses, and emotions were all put in motion
and guided by the action of the nerve trunk, which consisted of a
number of fibres running parallel with each other and the ganglia,
_ or knots which the fibres and nerve cells formed at particular points
in the cerebro-spinal axis. These gazg//a were united in one superior
ganglion, which in the higher animals was situated immediately
beneath the brain, which surmounted it. The nerve cells were called
globular, but, as a matter of fact, the prolongations proceeding from
them made the form stellate,
The ganglia were connected with each vertebra by two distinct
and separate fibres, one of which was named the sentient and the
other the motive nerve. These had two separate, although similar,
functions, and in many respects bore an analogy to the phenomena
presented by the electric telegraph. As in the telegraph, two wires
were required for the efficient performance of the work, so two fibres
were required to sustain the nervous force in its integrity. And as
the telegraphic apparatus required stimulating by a relay of electric
_ batteries, so was the amount of nervous energy supplemented by the
ganglia or knots which occurred at intervals throughout the system.
The simpler the animal in construction, the more simple the
nervous system. For instance, in the case of the ascidians a single
_ ganglion only was required to perform the offices required, and this
single ganglion was typical of the more elaborate nervous system of
the higher animals.
The motions produced by these nerves were quite involuntary, and
must in their consideration be kept quite distinct from the actions in
which the will was brought to bear; the ejection of matter from the
interior of one of the ascidians being precisely the same as the ejection
__ of a crumb or other foreign matter from the throat of a human being,
- for the latter could not help the violent action which naturally took
Place to free the body from the pain and danger attending the lodg-
ment of such foreign matter in the throat. .
In the case of the articulata, the number of geng/iz corresponded
with that of the articulations, so that each segment of an insect was
_ provided with its own ganglion. In the higher animals the ganglia
22
were united at the base of the brain, and thus the whole of the nervous
system was placed in direct connection and communion with the seat
of the intellectual and mental powers. «
At the PRESIDENT’S suggestion, Dr. HALLIFAX consented to
continue the subject at the next ordinary meeting.
NOVEMBER 25TH.
MICROSCOPICAL MEETING.—MR. T. W. WONFOR ON
“ POLYCYSTINA.”
Almost every microscopist possessed a slide or slides marked
“ Barbadoes Earth,” or “ Polycystina,” and many only regarded them
as beautiful and exquisite objects, without, possibly, considering what
they were or what the nature of the organism, whether animal or
vegetable, which originated these beautiful crystal-like forms of almost
every conceivable shape.
If we turned to the manuals on the microscope, we found that
they were the minute siliceous shells of a class of animals, whose living
part or substance consisted of a brownish coloured “ sarcode,” some-
what resembling that met with in the protean animal, the amceba,
“4 little particle,’ as it had been said, “ of apparently homogeneous
jelly, changing itself into a greater variety of forms than the fabled
Proteus, laying hold of its food without members, swallowing it without
a mouth, digesting it without a stomach, appropriating its nutritious
material without absorbent vessels or a circulating system, feeling (if
it has any power to do so) without nerves ; and not only this, but in
many instances forming shelly coverings of a symmetry and complexity
not surpassed by those of any testaceous animals.”
That was actually what the Polycystina accomplished, being mere
lumps of savcode. They spun, or wove, or, at least, formed for them-
selves coverings resembling the most costly and delicate filigree-work,
rivalling in beauty Chinese carvings in ivory, or ancient Peruvian or
modern Indian or Maltese silver filigree, out of a substance less work-
able or ductile, but infinitely more beautifu', than ivory or silver--
—
23
namely, silex, extracted, in some way at present unexplained, either
from their food or the element in which they lived—the sea. These
exquisite habitations were perforated to allow for the extrusion of
the pseudo-podia, as they were called, to be sent out in all directions
to collect whatever was needed for nutriment or continuing the build-
ing up of their habitations.
They were not confined, as was once thought, to fossil forms and
periods, but were to be dredged up living in the Mediterranean, the
North Sea—notably off Cuxhaven—the Pacific, and the Atlantic ; yet,
though the greatest care had been used in drawing them up through
the water, the mere friction or pressure generally caused the pseudo-
podia to shrink and collapse. According to Dr. Carpenter, the sarvode
substance did not always fill the shell, often occupying only the upper
part, and showed a tendency toa regular division into four lobes.
Some of the shells were spheres with smaller spheres like Chinese
balls within them ; these spheres were often connected by radiating
rods, sometimes prolonged into very delicate spines. Some of the
globes had lesser globes attached to them, with ovoid or conical pro-
longations ; others were star-shaped, with the spaces between the rays
filled in with a most beautiful network.
Although they were almost as widely distributed and quite as
varied in their form as the foraminifera, yet they must in past times
geologically have been more numerous, as we found whole rocks com-
posed of their shells, mingled with the siliceous valves of diatoms ;
thus the chalks and marls of Sicily, Greece, and Oran, in Africa, and
the diatomaceous deposits, many feet thick, in Bermuda and Richmond
(Virginia), contained them in great variety ; but the great source of our
supply was from Cambridge and Chimborazo, in the island of Barba-
does, where immense masses of a chalk-like rock were found, composed
almost entirely of the siliceous shells of Polycystina. A supply of this
rock containing most lovely forms, had been sent by their kind friend
and member, Mr. T. Curties, of Holborn, for distribution that evening,
and he had also sent a couple of slides prepared by Topping and Cole,
_ from the same deposit, with a hope that some of the members might
be incited to emulation, and produce slides as good.
Those who wished to prepare the material would find an account
a how best to manipulate it by Mr. Furlong, in the first volume of the
___ “ Miscroscopical Society's Transactions” (new series), or in “ Davis
24
on Microscopic mountings and preparation.” They should be mounted
dry, or in balsam, and viewed as opaque or transparent objects, and
with the spot lens or paraboloid. There was a method of giving them
an enamel-like opacity by burning them on platinum foil, but whatever
method of mounting or illumination was adopted, all who had scen the
Polycystina under the microscope must allow they were very beautiful
and delicately sculptured objects, while the question how the creatures
whose shelis they were, constructed such charming habitations,
remained among the, at present, unsolved problems of Natural History.
The Meeting then became a Conversazione, when some exquisite
and beautiful specimens of Polycystina were exhibited under the
microscope.
DECEMBER 9th.
ORDINARY MEETING.—DR. HALLIFAX ON “ THE
NERVOUS SYSTEM AND ITS FUNCTIONS.”
( Continuation).
Dr. HALLIFAX said, at the last meeting he endeavoured to trace the
different gradations of the nervous system, as they appeared through-
out the series of animals, beginning at the lowest ; and his object was
to show that, as they ascended, the complexity of the nervous system
kept pace with the complexity of the animal organization, and was
not only developed on the introduction of a new organ, but even on
the exaltation of an organ already existing. The insect tribes
approached nearer than any other of the series to the mammalian ;
and, by way of illustration, he would select one of the social insects, a
bee or a wasp, as denoting in its action the exercise of an intelligent
principle. In the higher forms of insects was found the cerebro-spinal
axis, the great central organ, consisting of the spinal cord and the
cerebrum. In man the cord was of great length, proceeding from the
cerebrum down to the lower portion of the frame, sending off, on each
side, throughout the system, thirty-one pairs of spinal nerves. Each
25
two of these nerves were contained in one sheath, although fulfilling
entirely different functions; one at the posterior ministering to the
purposes of sensation, and the other, at the anterior, ministering in
response to the contraction of the corresponding muscles.
Next, they approached that important doctrine which was called
the ‘excito-motor ”—the sensory nerve conveying the impression from
without to the ganglionic matter, which impression was responded to
by a mysterious action in the motor, or those nerves the filaments of
which supplied the muscular system that holds throughout. If they
examined the ventral cord of the higher insects,—so called from being
found in the ventral region,—two cords could be distinctly traced ;
and these were knotted into ganglionic little cells. Going on to the
thorax of insects, to which were fixed the legs and wings, they all knew
the remarkable activity of those limbs,—they found the ganglia agegre-
gating together to provide due energy for them. Advancing: to the
cephalic ganglion, there was a still further enlargement and evolution °
of this nervous matter, it being the neighbourhood of that wonderful
organ, the insect’s eye. It was under the guidance of visual impres-
sions that almost all the movements of insects were performed and
controlled, and thus they would observe how the proportion was kept
up between the ganglionic centre and the organ it supplied,
With regard to this double cord in the insects, it was remarkable
that, in the embryonic stage of the human form, the spinal cord had
also two distinct lines, so that, in the former, they had an element of
what was the commencement of the human spinal cord; while the
sensorium of the human brain corresponded with the cephalic ganglion
of the insect. In insects, the reflex action was automatic, all the
operations going on by virtue of the creature’s structure, without the
interposition of an intelligent principle, the majority of the actions of
the social insects being thus explained.
7 It was the same in the human brain ; but here they had the
_ Superadded organ, the cerebrum, the material instrument of thought
and will; and the degree of intelligence and self control corresponded
__ to the degree of development of that organ. It consisted of the
_ medullary matter, which was invested on its outer surface with a layer
_ of ganglionic cells, called cineritious, the medullary matter itself being
_ composed merely of nerve fibres, connected with the cortical substance
and the sensorium, From the sensgorium radiated fibres going to the
26
convolutions of the brain, these fibres being in two sets ; one carrying
an impression upwards to the cortical substance, the force returning by
the other to the motor part of the sensorium. The cineritious layer
was made up of vascular members by which the great part of the
circulating fluid was conveyed to the cortical substance, and the whole
action of the brain was dependent upon that supply.
As showing the importance of this, it might be mentioned, it had
been established that, although the brain was estimated at a 45th part,
by weight, of the entire body, these arteries, conveying blood to the
cranium for the purposes of the brain, were actually one-fifth of the
whole circulating mass ; and although there were four principal vessels
conveying arterial blood into the cranium, encircling it and branching
off to supply the whole external surface of the brain, they were so
powerful that one was sufficient to keep up the circulation in a subdued
degree.
With regard to the cerebellum, it was little understood ; but the
theory had been propounded that it controlled the reproductive system.
The whole brain was divided into two hemispheres, united by a band,
in order to give them a harmonious co-operation ; and it was a curious
fact that, in some instances, where this connection was greatly deficient,
the person lacked foresight, and was utterly unable to act upon
experience. Before resuming his seat, Dr. Hallifax said the subject he
had endeavoured to deal with was a great and complex one. He had
given a brief outline of what he had thought worthy their consideration,
and he recommended those who sought further information to study
Dr. Carpenter's book on “ Mental Physiology.”
The President, Mr. J. DENNANT, conveyed the best thanks of
the meeting to Dr. Hallifax for his interesting lecture, remarking that
no Member of the Society was so competent to deal with the physiology
of the nervous system as the eminent doctor who had addressed
them.
A discussion followed, in which the ex-President, Alderman Cox,
Mr. J. E. HASELWOOD, Mr. WONFoR, the PRESIDENT, and Mr.
DENNET took part. The interest of it centred in the automatic
action or otherwise of the nervous system, as laid down by Dr. Hallifax,
and opinions varied on the point ; Alderman Cox holding that the
different functions developed a certain mental activity which produced
27
a certain effect on the muscles; while the President expressed his
opinion that problems were often worked out by the brain automatically
and without any mental exertion.
JANUARY 13th.
ORDINARY MEETING.—MR. C. P. SMITH ON “ BEES.”
The bee was an insect consisting of a head, thorax, and abdomen,
united by cylinders which were at times so reduced as to make it
wonderful how the minute nervous cord passing through it was able to
control all the functions of the organs of sensation and motion. The
head was the most important section, and carried the antennz, the
compound eyes, the simple eyes, and the organs of the mouth. The
antennz, or feelers, were variously constructed, being sometimes
straight and at other times curved, or like a string of beads. In a
wild bee the integument of the antennz was of a hexagonal structure.
The antennz had been supposed to be organs of smell and also of
hearing, but this was by no means certain; they were, however, in
constant motion, and were frequently protruded into a flower before the
insect entered, for the purpose, apparently, of ascertaining its fitness.
The fact of a special apparatus for the purpose of cleaning them
would also show that Nature had attached great importance to them.
On each side of the head were the large compound eyes which varied
in the various genera, and consisted of a number of small hexagonal
facets, each one having its own branch of the optic nerve. The use
of these compound eyes was supposed to be the examination of near
objects. On the crown of the head were the three stemata, or
simple eyes, which were supposed to be used for long vision,
as when covered up by a black varnish Réaumur found that the bee
invariably flew straight up, and was lost. The organs of the mouth,
called trophi, were very variable in development in the different
genera, and, when complete, consist of a labrum, or upper lip, a
4 epipharynx, or gullet, which formed the mouth, the labrum, or lower
lip, and an organ called the tongue. Besides these single organs the
_ organs in pairs were the mandibles, the maxille, maxillary palpi, the
labial palpi, and the paraglosse. The labrum was of a horny or
28
leathery consistence, and closed over the organs below it, being
covered when at rest by the mandibles. The gullet was placed close
under the labrum, and was closed by the valve above, beneath which
was a small triangular appendage, which received the honey or food.
The labrum had two joints, one of which carried the maxilla, or lower
jaws, which in a state of rest folded together and formed a sheath for
the lingual apparatus. The tongue was not, as was formerly thought,
tubular, but flat, and when at repose was broader than thick. It could
be lengthened and shortened with great rapidity, thus pushing the
syrup up to the gullet. Réaumur proved that the food always passes
over the surface of the tongue, and not through the aperture which was
thought to exist in it. The thorax was divided into three parts, to the
front of which the front legs were attached. On the tibia of the fore-
legs in all bees was a velum, so named from its resemblance to a little
sail, opposite which, at the base of the palmz, was a deep incision
called the strigilis or currycomb, from the pecten or comb of short stiff
hair which fringed it. This apparatus was for the purpose of cleaning
the antennee, which were drawn through the hollow incision while
being pressed by the velum against the currycomb. In the boring bees
the mandibles do the cutting and excavating work, while the forelegs or
hands were used to clear away the rubbish thus formed ; but by the artizan
becthe forelegs were used like trowels to work up the soft clay into cells.
The bee had the power of locking the wings together in flight by
means of a row of hooks on the inferior wing which caught in a ridge
or ledge in the upper wing, and thus gave the insect a much greater
power of beating the air. On the hinder portion of the thorax were
placed the hind legs, the shank of which in the social bee was dilated,
for the purpose of carrying pollen. The abdomen was often elegantly
coloured, and consisted of six imbricated segments in the female and.
seven in the male. In the artizan bee pollen was conveyed on the
abdomen instead of the hind legs, perhaps because of the narrow
entrances to their nests.
Passing on to the consideration of the honey bee, in particular,
they found the male or drone was distinguished by its more cylindrical
shape, its larger size, and by being more densely hairy all over.
The large compound eyes met at the top of the head and covered
all the side of the face. The abdomen consisted of seven segments,
being thus distinguished from the queen, which had six seg-
ments. ‘The structure of the drone, as of all male bees, incapacitated
29
him from any share in the work of the hive. So far as had, been
ascertained its sole use was to impregnate the young queens, and to
accomplish this end, Nature had been lavish in the production of drones,
about 1000 being produced in each hive during a summer, although, as
seldom more than two swarms departed, but three drones were required
for that purpose. The large eyes of the drone no doubt assisted him in
finding the queen during her nuptial flight, while the greater sweep of the
hinder wings enabled him to rise above her. The large number of
drones was no doubt required by the circumstance of fertilization taking
place high up in the air, and by the great importance of a speedy
return of the queen to the hive, as in the air she was in great fear of
being snapped up by birds, and thus in some cases the entire hive
must perish. The fact of the drones not adding to but diminishing
the honey had induced bee keepers as long ago as Aristotle to use
traps for their exclusion and destruction. The working bees, however,
in the latter end of July, suddenly made up their minds to stop the
consumption of honey by driving out the drones and preventing their
return, so that they perished from want. At the same time, the workers
dragged out and stung to death all drone grubs, being apparently filled
with the utmost fury. During a wet summer, when honey could not
be obtained, the drones were driven out earlier. The queen bee was
the mother of the hive, and was easily distinguished from the other
bees by her size, being much longer and the abdomen being much more
spindle-shaped. Her chief function was to lay eggs ; and this she did
both day and night. The manner in which the eggs were fertilized
was a great mystery, until Dzierzon showed.that there was communica-
ting with the oviduct, through which the eggs passed, a spermatic
reservoir. The fertilization of the queen took place on the wing, and
it appeared probable that it only took place once in life. Two or
three days after impregnation she began to lay ; and if not permitted
to leave the hive she lay all the same, though she only produced
drones. This had been proved by Siebold who kept some virgin
queens caged and found they produced drones only. From observa-
tions and experiments it seemed probable that the queen only laid two
kinds of eggs, viz. : worker and drone, the former being capable by
£ appropriate feeding and management of being developed into queens.
- The queen only laid eggs, leaving the workers to look after them and
_ attend to the young. The other peculiarities of the queen were the
-want of a honey bag, the inability to secret wax, the convex thigh
30
which prevented the gathering of pollen, and the blunt sting that could
not pierce the human hand. It was, therefore, dependent on the
workers for food, which was continually presented to her by at-
tendants who, like true courtiers, never turned their backs on her. The
neuter, or worker, was principally distinguished by its pollen-collecting
apparatus. Though called a neuter, it was in reality an undeveloped
female, and occasionally laid eggs, as previously noticed. Perhaps
the most important organ of the worker was the marvellously compli-
cated muscular tongue with which the honey of flowers was not sucked,
but lapped up, and passed along the grooved surface to the mouth, and
then to the honey bag. The tongue of the hive-bee was twice folded
in repose. The honey was collected from a great number of flowers,
of which the lime, heather, and white clover, were perhaps the princi-
pal. The proboscis of the hive-bee was not long enough to reach the
nectary of the red clover, which was visited by the humble-bee. Asa
great deal of honey was needed to make wax for forming the comb,
means had been adopted to empty the combs and return them to the
hive. Combs were built most rapidly at night, the bees working during
the day at honey collecting. The peculiar structure of the working.
bee could not but be admired in conjunction with its work of cell
building. Their mandibles were endentate and like spoons, and were
so used for strengthening and plastering their work, while the brushes
on the posterior feet and the little auricle enabled the hind legs to
gather up the wax as it was formed, and pass it on to the mouth to be
masticated and moulded by the mandibles. The want of spurs onthe
hind legs (in which the hive-bee differed from the humble-bee), gave a
greater freedom and play to the comb of short stiff bristles at the ex-
treme edge of the shank. Propolis, an adhesive substance, gathered from
the buds of poplar, hollyhock, willow, and other trees, was gathered
and used to strengthen the defences of the hive. Propolis was with
difficulty kneaded into a ball and brought home in the corbiculz, but
it dried so quickly that it was sometimes with difficulty torn by the
bees from the legs of the collector. This substance was indispensable
for filling up cracks and cementing the hive to its floor boards, which
was always done for the sake of keeping out intruders. The eggs of
the bee were of a long oval shape, and being covered with a glutinous
matter stuck to the bottom of the cell. In three or four days they
hatched a little white worm which had fifteen segments, each, except
the head and four terminal ones, supplied with a pair of spiracles. It
31
had no feet, but small knobs existed on each side of the segments,
giving it a slight power of motion. This grub was very voracious, and
was sedulously supplied with food by its nurses. On this food it
throve until in four days it was fully grown and filled the cell. The
worker then sealed over the cell, and the grub spun a cocoon of silk in
which it grew to maturity, which is in twenty-one days. It then burst
the covering of the cell and flew out. The cells for the queens were
four or five in number, and they hung with their mouths downward.
They were four times as large as the others, and much stronger. The
queen grubs were so largely supplied with food, that they were in a
thick bed of jelly, some of which usually remained after hatching.
They arrived at maturity one-third earlier than the others, in conse-
quence of the difference of their food, which also prolonged the life of
the queen to two or three years instead of a few months. The young
queen was jealously guarded by the workers from the queen mother,
and at its nuptial flight the old queen generally left with other bees,
who gorged themselves with honey before departing to seek a new
hive.
Mr. H. Goss asked if a difference in the temperature of the cell
of the queen bee had anything to do with the development of a worker
- into a queen.
Mr, C. P. SMITH did not think it to be the case, and in answer
to further questions from Mr. Goss, he said he did not know that the
vertical position of the cell had anything to do with it; or if the food
given to the queen was different to that given to the workers.
Mr. Goss said he had seen it stated that a queen laid from
70,000 to 80,000 eggs.
Mr. SMITH added that a bee only lived seven or eight weeks
during the summer.
DR. INGLE called attention to the shape of the cells, and said it
had been a mathematical problem as to whether the equilateral
triangle, the hexagonal, or the square shape would waste the least
space, and it had been found that the bee adopted ¢#e one—the
hexagonal.
32
THE DEATH OF Mr. HENNAH.
The CHAIRMAN said it was a painful duty for him to bring before
their notice the death of Mr. Thomas Hennah, one of the Vice-
Presidents. Those who had been members of the Society for some
years would realise the real loss that had fallen upon them. He had
ably sustained the office of President, was at all times a wise
counsellor, and certainly exhibited a mind of no ordinary culture.
He did not think he would be wrong if he said that Mr. Hennah was
one of their best microscopists. He was kind and genial in his
manner, willing at all times to impart to younger members the
knowledge which they sought. He had made himself so completely a
helper in the work of the Society that his loss would be felt by all.
There were very few to take the places of those friends they had lost
lately, and he hoped that when their loss was made known to the
public, men of scientific attainments would consider it their duty to
think of the Society. In conclusion, the Chairman moved the following
resolution :—
“The Brighton and Sussex Natural History Society, deeply
lamenting the loss it has sustained in the death of Mr. T. H. Hennah,
desires to convey to Mrs. Hennah and her family a sincere expression
of sympathy with them in their sad bereavement, and to express a hope
that it may be some solace to them to know that her late husband was
held in the highest esteem and respect by his co-workers in the Society,
who cannot fail to miss in no ordinary way a Member of such con-
spicuous ability and genial character.”
The funeral would take place on the following day. He regretted that
his health would not permit him to be present, but he hoped that as
many Members as could would be present to show their respect to
their old friend.
Mr. WONFOR seconded the resolution, as a personal friend of Mr.
Hennah and his family for more than eighteen years. During the
time he had known them he had always found Mr. Hennah a
man of great kindness and courtesy. His death to him was a great
shock ; and he felt that in losing him they had lost a great support to
the Society. It was he who established the microscopical meetings
which had proved so interesting ; and not only had he done this, but,
. 33
while his health would permit it, he helped to keep up a microscopical
club. Not only was his death a great loss to this Society, but also to
the town at large. There was scarcely a gentleman in the town who
worked with his microscope but was indebted for advice, assistance,
and instruction to Mr. Hennah, who had been always willing and
ready, even though suffering from physical debility, to give information
to the mere tyro equally with the advanced student. All who came
in contact with him could bear witness to his affable and gentlemanly
bearing, and the genial way in which he conveyed information to
others.
JANUARY 27TH.
MICROSCOPICAL MEETING.—“THE ANATOMY OF
THE BEE,” IN ILLUSTRATION OF MR. C. P.
SMITH’S PAPER ON THE BEE, READ AT THE
JANUARY ORDINARY MEETING.
The PRESIDENT (Mr. J. Dennant) said there was no paper to be
read, as Mr, Smith’s paper at the last meeting had almost exhausted
the subject, but as there was neither time noropportunity on that occa-
_ sion to examine the several parts of the bee described by Mr. Smith,
an opportunity would be afforded them that evening of so doing. He
4 believed Mr. Smith and other gentlemen had brought down a number
_ of objects of a very interesting character.
Mr. T. W. WONFOR remarked that he had since the last meet-
_ ing examined the curious bodies in the specimen of American honey
~ shown by Mr. C. F. Dennet, and found they were crystals of honey
sugar. It would be seen by comparing them with crystals of cane
qj ‘sugar how much they differed from it in their mode of crystallization.
‘He found also plenty of pollen grains in the honey, but whether they
were of the orange, on the flowers of which the bees were said to feed,
he had not had the opportunity of verifying. He had also examined
34
the Zsoezes exhibited at the last meeting, and had found both macro-
spores and microspores.
Mr. C. P. SMITH, in reply to the President, said he had brought
down for exhibition the general anatomy of the hive bee, mounted on
one slide, the ovaries of the queen bee, a very uncommon object, in
which the eggs were shown, as well as the sting, the tongues of the
hive and other bees, and the sting of wasp.
Mr. WONFOR said he had the tongues of the humble bee, the
honey bee, and the wasp for comparison, the stings of the wasp and
bee, the latter showing the two serrated stings protruded from the
sheath, the heads of the hive bee and wasp, parasites of bee, and
crystals of honey and sugar.
The meeting then became a conversazione, when the above-
mentioned and other objects illustrating the anatomy of the bee were
exhibited by the President, Messrs. Haselwood, C. P. Smith, E.
Glaisyer, R. Glaisyer, and Wonfor.
FEBRUARY IOTH.
ORDINARY MEETING.—MR. BENJAMIN LOMAX “ON
THE BRANCHING OF TREES.”
The subject of the paper had always possessed the greatest
interest for him, principally, he thought, because he seemed to have it
all to himself. Until the time of Lindley, physiologists appeared to
look upon a tree as a structure composed entirely of a root and
flowers, and though Le Maout, Sachs, and others had lately bestowed
much attention on Taxophylly, the growth of wood, and the develop-
ment of branches from buds, that symmetrical arrangement by which
each tree acquired its marked and unmistakeable character and out-
line, preserved to the finest veining of its leaves, had been
entirely neglected by the botanists, and only remarked on, from an
artistic point of view, by Ruskin, in his “ Modern Painters.”
That the form of a tree was regulated by general laws, independent
:
35
of its minute structure, would seem probable to every naturalist ; and
the Society would be prepared to look with favour upon an explana-
tion which referred it to the composition of two combining or opposing
forces, as in Mechanics and Electricity, even though the way should
not seem clear for pointing out the exact nature of the forces them-
selves. Such an explanation he had to submit. It would account for
every peculiarity in the form of branches and leaves, considered as
such, but it was so entirely independent of the well-known laws which
regulated the formation of wood rings, &c., that it seemed impossible
at present to harmonise the two. If his position were true, they would
ultimately be harmonised. If false, the facts at least were worthy of
_ notice; in any case he claimed for his scheme the consideration due
to lengthened investigations under very favourable circumstances.
For fifteen years he had bestowed pretty constant attention 'to this
subject. He had made some hundreds of measurements in England,
_ Australia, and Peru, on trees of all sizes, from the lilac and alder to the
_ Eucalypti. For this purpose, he had used, successively, a lady’s
_ Measuring tape, a surveyor’s chain, and a theodolite or sextant,
a and on one occasion had allowed himself to be hoisted 120 feet in the
air by a rope drawn over with the help of a bow and arrow, but never
had he found a single instance of departure from the principles
described.
7
4
3
te It was not at every period of a tree’s life that it possessed a
perfect form. During what might be called its childhood the branches
were not yet visible ; in its advanced maturity the true branches were
intermixed with subsidiary limbs, forming no part of the original plan,
and produced by the development of what would have been mere
wigs but for accidents to which he should hereafter refer; while old
age showed us a confused and tangled mass of branches, broken and
‘repaired, or bent in distorted forms to avoid collision with others, or to
ce advantage of some accidental condition. But there was a
e in a tree’s history which might be likened to our own useful
sapling nor the unintelligible luxuriance of the maturer growth,
when the laws to which he had referred might be easily recognised
d fully traced. He had spoken of branches as normal and subsidiary
accidental, and it was necessary that he should justify such a
By way of doing so, he would ask those who had seen
36
trunk or Jarger limbs at the point where a branch left them. He had
had much to do with sawmills, and had seen many such sections cut,
and often one tree, falling beneath his axe, had lighted in the fork of
another and split it to the ground. The appearance then presented was
that presented by the forked twig which a schoolboy split to make his
bird trap.
The wood from bark to bark formed one continued strip of
white, from which the white wood of the branch turned off without
mark of junction, just as rivers were indicated ina map. These are
normal branches. Sometimes, however, a very different appearance
presented itself—a branch was apparently zuser/ed into the trunk,
penetrating only through the outer rings, and surrounded by a kind of
socket. Such limbs had commenced their growth after the branches
above them, and, though produced late in life, had outstripped their
predecessors, partly because nearer the roots, and partly because they
had been deriving their nourishment from the outer and softer layers
of the cambium region, where alone the sap descended in any con-
siderable quantity. It was only of the normal limbs that he should
speak for the present.
A well formed tree, then, consisted of a trunk or trunks, ascending
to a certain distance and there throwing off one or more limbs. These
limbs might be large or small in comparison with the parent stem,
might be opposite or slightly separated, and might turn off at any
angle not exceeding go°, Where the limb turned off the stem was
deflected in the opposite direction. That deflection might be very
slight, but it could always be traced and sometimes was so great as to
render it impossible to tell which of the two or move divisions was to
be regarded as the continuation of the parent stem. The original
pattern thus given was afterwards adhered to, and throughout the
whole structure each branch with its branchlets repeated the same
form of subdivision, so that it might be traced to the minutest ramifi-
cations, and could be seen in the skeleton of the leaves. If, then, we
rightly understood one forking, we understood all, and he proposed to
consider one minutely. There were several points requiring attention,
which might be enumerated as follows :—
1. The size of the limb (i.e., its sectional area),
2. The size of the stem above the joint.
3. The angle at which the limb branches off.
4, The amount of deflection in the stem.
5, The relative length of the previous and succeeding internodes.
37
If his view were correct, all these were connected together by one
common law, so that if one element were given the rest could be
deduced. The required data were the amounts and proportion of two
forces whose nature he did not yet attempt to define, the one tending
to lengthen the stem and its offshoots, the other tending to separate
the wood fibres laterally, the two energies holding much the same
relation to each other as the centrifugal force and that of gravity did
to each other in the planetary system. If the first force predominated,
the tree would be tall and narrow; if the latter, it would be wide-
spreading. The ratio of the two gave a constant fraction, which
might be called the “ co-efficient of ramification,” and from which the
whole outline of the tree might perhaps be calculated, just as a curve
might be traced from its equation.
The size of the first limb was doubtless determined by some in-
herent law, as trees of the same species showed a striking uniformity
in this respect, but he was not able even to imagine its nature. He
knew, however, that the proportion of the limb to the stem once fixed
was preserved throughout, and that every shoot bore the same propor-
tion to its parent as the first branch to the primiiive stem. The size
of the barrel above the limb was far easier of determination. Its
sectional area, added to that of the limb, always gave the sectional
area of the trunk from which they sprung, and this was equally true
where the division was into three or more branches. Thus, if from a
trunk 37in. in circumference two limbs, measuring respectively 18in.
and 12in. round, forked off, the resulting stem would measure 3oin. in
¥ circumference, whether the two branches were at one point or separate,
_ or, in other words, the sum of the squares of 18, 12, and 30 equalled
_ the square of 37, nearly. This law, which held good, without ex-
ception, in every tree and every twig, settled a most important
_ question, showing us that limbs were not off-shoots or lateral growths,
but actual divisions of the stem from which they sprung; and this
being so, we were led to infer that the whole of the leaf stalks on a
tree, if united in one bundle, would make up the thickness of the main
trunk, a conclusion as startling as it was inevitable.
This result might be illustrated by binding together, say, a hun-
' twigs, afterwards dividing them into ten, and these successively
nto fives, twos, and ones. The effect would be, that a fair model of
t tree would be produced, in which the ultimate divisions Donne
38
across, a very different spectacle was produced from that seen in the
cross section of a forest tree. In the latter we saw a series of cor.-
centric rings traversed by radiating lines, known as “ medullary rays ;”
in the former, a collection of circles, which, under extreme com-
pression, might assume the hexagonal form. But this dissimilarity
vanished on closer inspection and a more careful consideration of the
manner in which trees grow. If we surrounded one twig by a circle
of others, and subjected the whole to uniform pressure, we should find
that the outer twigs, if sufficiently compressible, would form one con-
tinued ring, wherein each separate piece had the wedge-form produced
by intersecting radii. Ring after ring might be thus formed, and the
whole section of an aged tree satisfactorily imitated.
It would be noticed that in the fancied model each medullary
wedge was connected with one twig and one only ; and it might be
asked whether he considered such to be the order of actual tree-
growth? To this he answered that his illustration was but an illustra-
tion, and was intended to prove nothing ; but the supposition was quite
possible, though he knew nothing whatever concerning its truth. The
angle of ramification was very uniform in trees of the same species,
and was readily explained on the principle to which he wished to
draw their attention.
Every student of dynamics knew that if two forces acting on or
from one point be graphically represented by two lines of equivalent
length and direction, the diagonal of the parallelogram so indicated
would give the length and direction of their resultant, and wice versd.
So if we knew the longitudinal and lateral forces at any given fork, we
should be able to trace the angle at which each division would leave
the point of separation. As these data were not forthcoming, another
means of investigation must be found. It was manifest that a force
tending to separate any pair of fibres must vary in total amount
directly as the number of fibres present ; and as these together made
up a limb, it followed that the amount of pressure tending to cause
any two limbs to diverge from the original direction would be in direct
proportion to the sectional area ot the limbs themselves. On the
other hand, there was frida facie reason to suppose that which he
hoped to establish by another line of reasoning : that the tendency to
longitudinal extension was at its greatest at the commencement, and
decreased according to the amount of work done, or according to the
distance from the root. On such a supposition we readily deduced a
39
result which agreed exactly with observed fact, namely, that when
two branches of equal size separated, they would have exactly the
. same angle of divergence ; that where one exceeded the other, the
larger would be nearer to, and the smaller farther from, the line of
original direction; and that where the limb was very small in com-
parison with the stem, as in some of our coniferz, the angle of rami-
fication would be indefinitely near to a right angle, and the stem
_ practically perpendicular,
They next came to the question of internodes ; and here they
might with profit refer to the herbaceous plants, which for the most
part diminished them with great regularity, but in addition suppressed
them altogether when any large draught was made on the vital
reserve. Thus we found the lower leaves alternate, the higher
opposite, and those surrounding the ovary verticillate. Such a rule
obtained in forest trees; the internodes ordinarily decreasing so
regularly that the proportion between the length and thickness of any
internode was that of any other in the same tree ; but this regularity
disappeared, and whole internodes were suppressed where any
abnormal necessity required the tree, for the preservation of its
balance, to push out extra branches, or extend branches beyond their
intended length. Without wearying by details that had been very
pleasant to him, he might say that he had never seen a case of ramifi-
cation which could not be explained on the assumption of two com-
_ bining forces, and he did not expect to find one.
Between this humble attempt and the time when each tree should
i have its equation, whence its outline and taxophylly might be pre-
_ dicated, there was a great gulf fixed. If ever such a supposition could
Bie reconciled with the annual succession of rings, it must be by the
_ application of laws not yet enunciated, and many generations might
pass before the connection could be held possible, but till that connec-
tion were established, the likeness of ramification to the most familiar
_ effects of mechanical composition must be held as one of the most
remarkable coincidences in Nature.
He would, in conclusion, remark that any gentleman wishing to
_ pursue the same line of observation must prepare himself for difficulties,
4 The effect of gravity, the reflection of heat from walls and banks, the
presence of water in hot climates, and the alternate loss and recovery
as of balance, supplied a series of complications which must be patiently
40
eliminated before the true law of ramification could be traced; but if
any one, after such reasonable deductions, should fail to find every-
where the same unvyarying proportions mentioned, he must have an
experience widely differing from his own.
The PRESIDENT (Mr. J. ennant), on behalf of those present,
heartily thanked Mr. Lomax for his paper, short, but interesting and
valuable, being original.
The discussion on the paper was opened by Mr. WONFOR pointing
out that the subject had been treated from such a novel aspect, that
difficulty would be experienced in discussing it, and thanking Mr.
Lomax for suggesting ideas which might be considered in the coming
spring.
Mr. G. D. SAWYER and Mr. HASELWOOD followed, the latter
observing that there was a strong tendency to account for biological
facts by mechanical laws. This was not new to them—they had that
theory promulgated before, though not in so practical a form as that
night. But he understood that the gentleman who asserted mechanical
laws as an explanation of biological facts went further than Mr. Lomax
had gone, and, therefore, he advised those present to be cautious in
adopting mechanical laws as accounting for those things, as such an
adoption might carry them further than the application of those laws
to mere vegetable productions. He also asked if the laws laid down
held good to any degree in reference to the downward progress of the
tree as well as the upward—to the ramifications of the roots as to those
of the branches ?
Mr. SAWYER gave instances of the continual variation of the
growth of trees.
Mr. LoMAX replied to Mr. Haselwood that he looked upon
the forces which he had mentioned in his paper as forces of
vitality, which had to give way to the ordinary mechanical laws, and
that he had not studied the downward progress of the tree.
Mr. C. F. DENNET also took part in the discussion.
CURIOUS ARCHITECTURAL SPECIMEN.
Mr. T. W. WONFOR, one of the Hon. Secretaries, brought forward
a large stone block, which, he pointed out, had on one side ecclesiastical
carving common to the thirteenth century, and, on the other, civil
=
4I
carving belonging to the seventeenth century. The block was taken
from a house of Mr. H. Gorringe, at Kingston-on-Sea, while some
alterations were being made, and, being of a rather interesting
character, was forwarded to the Brighton Museum.
IMPOSITION,
Mr. C. F. DENNET submitted specimens of imitations of silk, the
cheapness of which, had they been silk, attracted his attention. The
fabric, however, was almost jute alone, so manufactured as to appear
silk to the untutored eye, and to deceive the touch of an experienced
person.
FEBRUARY 25TH.
MICROSCOPICAL MEETING.—MR. T. W. WONFOR
ON “THE SCALES OF BUTTERFLIES.”
Among insects, scales were not confined to any one group, though
they were found on every member of one division, the Lepidoptera, the
scale-winged, as their name implied, but in connection with microsco-
pical work the scales obtained from certain insects had been, and still
were, favourites with microscopists, very deservedly, because, through
differences of opinion as to the markings on sundry scales, together
with an attempt to resolve those markings, we owed the great improve-
ments made in objectives since the achromatic microscope had been
an instrument of research, and not a mere optical toy,
As all knew, certain scales, such as those from the gnat, Zepisma,
Podura, or Lepidocyrtus, and three or four butterflies, viz. a blue and
a white of English origin, and a couple of gorgeously coloured
_ foreigners, had been employed, along with sundry siliceous valves of
plants named diatoms, as /es¢ objects, and our objectives had been
_ considered up to or below the mark accordingly as they were able or
~ not to resolve certain figured markings, seen by objectives of particular
aperture and magnifying power. Again, to enable these objectives or
others of different aperture and power to resolve the same or other
markings, various adjuncts to the microscope had been devised, such
42
as condensers, prisms, &c. And though many, alas! too many,
microscopists in this country had spent nearly all their time in trying
to see exactly the same things that other men had, or said that they
had, seen, yet within the last few years a very considerable addition
had been made to our knowledge of many physiological facts, as well
as the resolution of diatom and scale markings ; and though we might
not accept all Dr. Royston Piggott’s conclusions, yet had he not
ventilated the subject through his “ vouleaux of beads” and “ green
peas,” we might have gone on plodding in the same steps as those
before us, and accepted one view of an object as a sufficient des¢ of the
power of an objective.
Whatever view was taken of the markings of scales, there seemed
no manner of doubt that the scales of insects were nothing more nor
less than modified hairs, of greater or less thickness, more or less
flattened or cylindrical, according to circumstances and their position
on the body, legs, or wings of the animal on which found. If then we
regarded all scales as only modified hairs, and considered that hazrs
were composed of cells, we might see our way out of the difficulties
into which the learned amongst microscopists had led us, and we
might also understand an wnuder and upper surface, two lamina,
striated surfaces, ribs, and sundry other puzzling terms which had
come into existence when speaking of scales.
Those who had written or spoken about scales and their markings -
within the last few years, were Dr. Royston Piggott, and those who
inclined to his deaded theories on the one hand, among whom must be
included Mr Slack, one of the Secretaries of the Royal Microscopical
Society, and on the other Colonel Woodward, Dr. Maddox, Dr.
Anthony, and Mr. McIntyre. It was with the latter group he felt he
must ally himself, because much, if not all, of the so-called structure
described by Dr. Piggott he believed was purely optical and not
structural.
Primarily, the scales of insects, and specially the /epidopiera
were more or less flattened hairs of a cylindrical or tube-like shape,
inserted by a pedicle, differing in character according to circumstances,
into the wing membrane, on both sides of which they were arranged
in symmetrical rows, like the tiles on the roof of a house. The
majority of those on the wing, or rather the flat part of the wing, were
43
flat, or nearly so ; interspersed among them were others quite hair-like
in their character, and in some cases, to be attended to presently, were
some which were balloon-like in shape and, he believed, were capable
of being inflated like a balloon. He should mention that from six to
eight different kinds, or rather shapes, of scales were found on each
butterfly or moth, and when it was borne in mind how many thousands
of moths and butterflies were known to science, it might be imagined
that, though the varieties were not commensurate in number with the
varieties of the insects themselves, yet the varieties in shape and
markings were very numerous. Anyone devoting a little time and
attention to scales would be well rewarded for his pains, and would
learn far more by comparison than he could possibly glean by study-
ing one or two special scales.
Among other points he would find that the colour of insects was
not always due to what to the eye appeared the colour on the wings ;
a striking illustration of this was seen in the case of the common orange
tip, the green colour on the back of the wings proved on examination —
to be produced by an intermingling of black and yellow scales. Some
certainly owed their colour to pigments, but in a great many it was due
to the incidence of light, for if, while under examination under the
microscope, the stage were rotated, as great a change would be seen
in colour as with some objects under polarised light. Seen as trans-
parent objects, brown seemed the prevailing hue.
Again, if scales were taken from living insects or from those
recently killed, and gently squeezed by pressure on a thin covering
glass, greenish oily particles would be seen to ooze from the pedicle,
leading to the idea that in the living insect there was something like
the circulation of a fluid from the wing membrane. It was certain
they would not require any fluid to repair waste or to accelerate growth,
because they were of full size the instant'the insect escaped from the
chrysalis. There was no growth proper in the scales, any more than
in the wings. The instant the moth or butterfly emerged from the pupa
case, it came forth full-grown, the membranes expanded laterally and
longitudinally, and by so doing the scales of the full size and zz situ
were simply drawn wider and farther apart, but of growth there was
none. This could be seen by cutting open the pupa case just before
the emergence, or taking the wing of a recently emerged insect. This
must be done at once, because in some cases, within ten minutes of
44
the emerging the wings were of full size. There was, therefore, no re-
plenishing of waste nor reparation of parts inthe scales of lepidoptera,
as in the hairs of the mammalia.
Nor could we trace in all scales the cuticular layer, cortical sub-
stance, and medullary substance of such hairs ; we seemed to obtain
a cortical substance, with or without a cuticular layer, and sundry rib-
like strengtheners, which had given rise to the diversity of opinion as
regarded structural markings. There was another curious fact some-
tiraes noticed—viz., that there appeared to be a power on the part of
the insect to raise the rows of the scales as well as to inflate some
scales. He had often found the scales on the wing of a butterfly
caught and killed in flight, not lying flat to the wing membrane, but
raised on it at a more or less acute angle, as though in flight the insect
possessed the power of raising the rows of scales. This might, if we
imagined the power of inflation of the scales as well, give greater
buoyancy to a butterfly or moth in flight. He had every reason to
believe this to be true of some scales ; if so, it would give to the wing
membrane with its nervures a power not accorded to it in Manuals on
Entomology.
It was now twelve years ago he started one afternoon with a
medical friend to look for a certain fern, said to grow about twelve
miles from Brighton. They got out of a train at the Hassock’s Gate
Station, and diverged a little from their course to see whether Veottia
Sprralis (autumn lady’s tresses), the orchis which Darwin watched the
bees fertilizing, by carrying the pollen masses from one flower to
another, was in flower. While searching for this plant, some blue
butterflies got up. His friend and he managed to catch several. This
fact was strongly impressed on his mind, because, after walking nearly
a mile, they retraced their steps to search for a stethoscope, which, in
the eagerness of the chase, had dropped from his friend’s hat when
catching a “blue.”
On the way they both talked over the question of why they had
not been able to find “ battledore” scales on blue butterflies. Was it
they were only on one kind, for though they had both searched for
them on the part described in the Micrographic Dictionary, p. 564,
under Polyommatus, they had neither been able to find them. The
words there run, “ The scales upon the under surface of the wings of
45
P. argiolus and P. argus have been proposed as test objects. They
are of two kinds—one resembling in structure the ordinary scales of
insects, the other of a battledore form.”
What his friend did with his blues he did not know, he (Mr. W) care-
fully removed the scales from the under surface of one of the blues and
found not a single battledore. Having placed a wing upperside down-
wards on a glass slide to enable him to search over the wing, he found,
after a futile and long continued search, that a considerable number of
scales detached from the upper surface of the wing were upon the
glass slide ; focussing down to these he found battledores in plenty,
but not exactly like those figured in the book. He then realised this
fact, that battledores were to be found on the upper surface of the
wings of Polyommatus, or, as it was now called, Lycena alexi zs, differ-
ing in form from those figured as obtained from Z. argtolus. This led
him to try whether he could find “ battledore” scales on other blues
than Z. alexis, the common blue, or Z. argiolus, the azure blue, for to
those unacquainted with such differences among “blues” he should
mention we had, in England, nine different species of “ blues,” such as
the Common, Azure, Clifden, Mazarine, Chalk- hill, Silver-studded,
“large” and “little” blue, and rare “ long tailed” blue.
Through the kindness of an entomological’ friend he secured
examples of nearly all, and, strangely enough, found battledore scales
on the upper surface of both wings in some cases, and not in others,
It turned out that the insects on which he had been unable to find
battledores were females, while those on which he had found them
were males, This caused him to make a critical examination of the
blue family generally, and he then discovered these two facts—
the battledores were found on the upper side of the wings of the males
ondy, and in rows beneath the ordinary scales, and at their intervals of
overlapping. He should mention that, among the blues, the males
only were of a deep blue colour ; the females generally were of a
brownish hue, with a few blue scales, and this might have led to the
assertion that battledore scales were found on the “blues.” It was
true some females adorned themselves with so great a profusion that
they simulated the garb of the males, much in the same way as the
erstwhile “weaker sex” among humanity, since the cry of woman’s
rights had been raised, assumed the get-up and garments of the
“lords of creation,” but on no female, however highly coloured, had
he been able to find “ battledores.”
46
Since then, he had had the opportunity of examining many foreign
and tropical blues, and in each case, even when the female assimilated
closely in colour to the male, only on the male, and invariably on the
upper surface and in rows beneath the ordinary scales, had he found
battledores. As might be expected, these battledores differed in size
and shape—the blade was longer or broader in some than others
relatively, the top was more or less rounded or squared, while the
length of the handle or pedicle varied in length, so that, when two
species of blues resembled each other very closely in their markings,
the shape and size of the battledore might be used as a ready means
of settling species. There were species among the Lyczenidz in which
neither males nor females were “blue” or “bluish” in colour, and,
curiously, among these no single example of male with “ battledores”
had been found. Mr. Watson, who was working on “plumules ” at
the same time he was carrying out his investigations, had had
opportunities of examining some hundreds of species of “ blues” and
“brown” among the Lycznidz, and had never met with “ battledores ”
on any but the “ blue” proper.
He would turn next to another characteristic scale, the “ tasseled,”
described in the “ Micrographic Dictionary” under the head of Pontia,
p. 571, thus—“ The form and structure of certain scales existing upon
the uuder side of the male is curious.” Here, too, as he should show,
the writer had made a slight mistake, instead of wuder they should
read upper, for it was there, and there only, they would find scales
similar to those figured in the “‘ Micrographic Dictionary.” As might
be supposed, having got an inkling from the “ blues,” respecting the
situation of “battledores,” he was not long before he searched the
whites. The first to come under examination were the “large” and
“small” cabbage white Pon?tia, or rather now Preris brassice, and
P. rape, both of which gave on the upper side the characteristic
scales called “ tasseled,” or, as some preferred, “ plumules.” These
scales differed essentially from the “ battledores ” in shape, ornamenta-
tion, markings, and pedicle ; some were long and very slender through-
out, and gradually tapering to a point, others seemed cut short, while
others comparatively broad at the basal end, suddenly narrowed and
terminated either as though cut short or fined off to a slender point ;
all, whatever were their comparative breadth and length had a cup-like
or ball-and-socket termination to the pedicle. At the apex of each
scale was a tasseled fringe of great beauty, from which circumstance
_——
47
they had been called ‘‘tasseled” scales. All our English whites,
together with the orange-tip, possessed these peculiar scales, and if
the great family of Pieridz, which included a vast number of con-
tinental and tropical forms, were examined, they would find the males
invariably possessed a tasseled scale, either of the form of our English
whites and orange-tip, or a modification of these, both as regards the
terminal fringe and the pedicle. He had had opportunities of examin-
ing a good many, but Mr, Watson had enumerated upwards of 200
English and foreign Pieridz examined by him, on which, in every case,
the males, and the males alone, possessed the characteristic scale.
As might be supposed, there were great diversities of size and
form, and which, from what had been said respecting the English
Pieridze, might doubtless be very useful in determining species. One
curious confirmation of the value of these characteristic scales Mr.
Watson drew his attention to in 1868, viz., that two hitherto-believed
different species of Pzerzs turned out to be the male and female of the
same, the one having, and the other being without, the characteristic
scales. They were arranged in rows behind the other scales, as in the
“blues,” but many being long and hair-like, they appeared only as
hairs 27 situ.
The next English family to be noticed was that named Hifparchia:
the common meadow brown A, Fanira had a scale brush-like, taper-
ing like the large white, but differing from it in the markings in the
ribbon-like portion, as well as having a pedicle similar to that possessed
by the “blues.” Investigating this family, he found the same story
told throughout—a distinctive scale, ribbon-like, and tasseled on every
male of the family, but never a scale of such a character on ‘the
females.
In each of the families described there was generally a difference
of wing-markings between the males and females ; occasionally they
_ resembled each other, and some few insects had been found with the
_ wings on one side with the male markings, and on the other side with the
female markings. It would be very curious to find out if, with this
_ assumption of marking, they also on the male marked wings ‘had this
undoubted sex distinction.
_____ There was only one other family of English butterflies possessing,
as far as he had been able to make out, a characteristic scale, and
_ that was the Argynnide or Fritillaries, or, at least, those among them
48
the underside of whose wings was marked by metallic spots. The
male scales were of a very decided character, differing essentially
from those of the Pzerid@, on the one hand, and from the Aipparchie,
which they somewhat resembled, on the other. Some were ofa very
long, narrow, and ribbon-like form, with the tassel at the apex, while
others were shorter and broader. Some, too, like the Hipparchia,
were nearly opaque, except at the apex, while others had the ribbon-
like portion opaque for one-half its length. The position of these
scales on the wing differed from that of any others described, for,
instead of being placed in rows beneath the ordinary scales, they were
situated on the nervures or black veins of the upper surface, and had
mingled with them in some species very peculiar Indian club-shaped
scales or hairs. Mr. Watson, whose opportunities of examination had
been far greater and more extensive than his own, had found
“plumules,” as he calls them, on thirty genera of butterflies, or nearly
600 species. In every case they were found on the male insects
alone.
He consequently drew the inference, a very reasonable one,
that battledores, tasseled scales, or plumules, wherever found, were
characteristic of sex, and that of the male sex. What purpose they
served in the insect economy was not yet clear. Their paucity or
abundance on individuals could not be, as some had suggested, marks
of greater or less virility, because, as had been seen, scales did not
grow with the age of the individual, nor did they find more scales on
one freshly-emerged butterfly than another of the same species. They
might render the males more buoyant on the wing ; but here they were
met by this difficulty, well-known to field entomologists, the females
were more rapid in their movements, in most cases, than the males.
They seemed rather to be the analogues of the beard in man, the
mane in the lion, the comb in the cock, or the more brilliant
plumage of some birds, and, possibly, to insect eyes rendered their
possessors more attractive than the duller-coloured and non-plumuled
sisters of their species.
There were many debated points of structure he scarcely felt
justified in touching on, because he had rather dealt with the scales as
a means of differentialising species or determining sex, and not as tests
for objectives.
49
In obtaining the scales for examination, he had found the best way
to examine a wing was to lay it on a clean slide, then place another
clean slide or covering glass upon it and gently press. | Upon remov-
ing the upper slide or covering glass, plenty of scales would be
obtained in their relative positions. The covering glass could be
mounted, or if a ring of cement was run round the slide, a cover added,
and, when dry, a finishing coat were put on, the slide was ready for the
cabinet. He feared he had been too prolix in his remarks, and
not added any new fact to the store of the members of the Society ;
but if he were able to enlist one to pursue the study of scales of insects,
he should have done his quota towards the use of the microscope as a
means of research, for there were many paths open, besides the beaten
tracks, by which not only, if that was needed as a spur, fame might be
won by the discovery of new facts in physiology, but much pleasure
and interest might be derived by prying into the secrets of Nature
hidden from the unaided eye.
The paper was illustrated by chalk drawings on a blackboard.
A discussion followed, in which Mr, HASELWOOD, Alderman Cox,
Mr. DENNET, and others took part.
The meeting then resolved itself into a conversazione; when
a number of slides, showing the different scales and the wings of
butterflies and moths, were examined under the microscope.
FEBRUARY 29TH.
ANNUAL SOIREE,
The Fifth Annual Soirée was held at the Royal Pavilion with
_ great success, some six hundred members and friends being present.
___ Flowers and plants were judiciously arranged about the rooms,
and in the Corridor the staircases at each end were banked with ever-
greens, whilst the fire-places were filled in and the mantel-pieces
cked with floral adornments, Messrs. Balchin and Nell carrying out
S portion of the arrangements.
5°
In the North and South Drawing Rooms, the Saloon, and
Banqueting Room (where also light refreshments were supplied by Mr.
Booth), various objects of interest were set out for inspection, the
numerous microscopes, and other scientific instruments belonging to
the members of the Society, very appropriately engaging the chief
share of attention. Amongst those exhibiting microscopes were Mr.
S. Aylen, Mr. W. Ardley, Mr. J. Capon, Mr. T. Cooper, Mr. J.
Dennant, Mr. J. Gwatkin, Mr. R. Glaisyer, Mr. E. Glaisyer, Mr. J. E.
Haselwood, Mr. B. Lomax, Mr. Mills, Mr. Moore, Mr. W. Mitchell,
Mr, Moginie (Quekett Club), Mr. G. Nash, Mr. Pankhurst, Mr. W.
Puttick, Mr. H. Saunders, Mr. G. D. Sawyer, Mr. Welch, Mr. T. W.
Wonfor, Mr. T. W. C. Wonfor, Dr. Hallifax, and Dr. Tuthill Massy.
A very conspicuous centre of attraction was a huge case, three yards
long by two yards high, containing a pair of splendid mute swans,
with cygnets, which occupied one side of the Saloon. These noble
birds, which came from the Norfolk Broads, were lent by Mr. E. T.
Booth, the owner of the Dyke-road Museum, and excited much ad-
miration. They were mounted and stuffed by Mr. Savile, in the
short space of one week. The artificial foliage was remarkably
natural, and here and there a close observer could discern snails,
caterpillars, &c., modelled to the life. Arctic birds were exhibited by
Mr. Pratt ; Miss Glaisyer contributing others, and the same lady had
a very fine case of shells on view, some of them being exquisitely
beautiful. A number of “ Eskimo” articles, from Disco Island, where
the Arctic Expedition touched, were shown by Mr. H. Willett, as well
as a number of photographs relative to the search for the North-West
Passage, by Alderman Cox. A curious Indian bow for slinging stones,
an assortment of woods used in manufacture, and a strange weapon,
of ibex horn, from Somali, were shown by Mr. W. Saunders. Consider-
able interest was also manifested in a series of illustrations by Dr.
Corfe, of the hands and feet in animal life, whilst those fond of geology
found scope for investigation in the drawings of, and cores from, the
Sub-Wealden borings (which had just been completed for the
Scientific Exhibition at South Kensington), lent by Mr. H. Willett.
Mr. E. Moore, Corporation Analyst, had a very interesting
assortment of various descriptions of butter and cheese from all parts
of the world; his table being much admired for the very tasteful
manner in which the wares were set forth, Ivy, trailing leaves, moss,
butterflies, and other insects, were skilfully used to adorn the palatable-
OE
EWE EE LUIS ig tt be tos
51
looking ornamental pats of butter, and doubtless in this regard many
ladies got a hint which will not be thrown away. One of the specimens
deserved special mention,—that sent by Miss Hadlow, of Uckfield.
The young lady milked the cow, made the butter, and manufactured
the very pretty basket in which it appeared at the Soirée ; and, what
with tasteful adjuncts in the way of flowers and ferns, she carried off
the first prize in the minds of the numerous critical inspectors. Mr.
H. Bull, of Hurst, also sent a miniature hamper of butter, which looked
all that could be desired.
Superb collections of English and Foreign beetles, were contri-
buted by Mr. Dowsett ; butterflies by Mr. Colling ; and Indian butter-
flies and moths by Mr. Wills, occupied other tables, whilst no one could
fail to appreciate the beautiful skeleton leaves prepared by Miss
Gwatkin, and in the arrangement of which the greatest taste and skill
were manifest. Mr. C. Bellingham’s fine assortment of casts of medals
and cameos attracted considerable attention ; and another great object
of interest was a beautiful shell basket shown by Mr. Cox, that gentle-
man also exhibiting some splendid specimens of the lyre bird.
Electrical instruments of the most approved character were lent
by Mr. Whiting, of the Post office, and many of the company amused
themselves by forwarding messages to each other; whilst electricity
__under another aspect was seen in its application to vacuum tubes, pro-
ducing some beautifully tinted fireworks ; Mr. J. Capon being the
demonstrator. Mr. C. J. Rowsell had his combination graphoscope
and other fine instruments on view, and various other apparatus and
objects of interest, including some excellent photographs shown by Mr.
D. B. Friend, were placed here and there about the. various rooms.
During the evening, the following addresses were delivered in the
Music Room. The first was by the President.
Mr. J. DENNANT said it is usual on these occasions for the Presi-
_ dent-to offer a few informal observations preparatory to the work of
_ the evening. These, I think, are fitting occasions when we may fairly
bring before your notice the work of our Society. I may say, to com-
mence, that we are ina most flourishing condition. Financially, we
are extremely healthy, and the number of our Members is something
over 200. We have an excellent and valuable library of over 900
_ volumes of standard scientific works, and I am glad to say that the
’ library is now used by our townspeople without let or hindrance.
52
During the last year we have hada series of papers of a very valuable
character read before the Society. I propose to mention them, that
you may see the class of subjects which come under our
attention. The first to which I will allude is one delivered last
year by our very eminent townsman, Mr. Davidson, F.R.S. He was
generous enough to make known, through our Society, his original
researches in geology, and he gave us the gist of his observations on
brachiopods, in a paper entitled “What is a Brachiopod?” Mr.
Dennet followed with a paper on “ Vegetable Fibres,” and Mr. F. E.
Sawyer read another on ‘‘ The Birds and Mammals of Sussex.” Our
good and untiring friend Mr. Wonfor discoursed “On the Brighton
Aquarium, and what it has done for Science.” Then came Mr. B.
Lomax with a paper on “The Minor Diseases of Plants,” and Mr.
Pankhurst with one on “ The Ores of Iron.” Next we had a paper
from Mr. Potter “ On the so-called Forest Beds.” Mr. Wonfor, who
always comes in when any of our friends fail us, gave a second paper
on “ Manna ;” and we had a valuable paper read by Dr. Hallifax on
“ The Nervous System and its functions.” This was followed by Mr.
C. P. Smith’s paper “On Bees,” which proved very interesting, and
still another from Mr. B. Lomax on “The Branching of Trees.”
These papers have been so amply reported, that it would be un-
necessary and unwise to detain you further with them, except to
remark that the majority of these papers were considerably in advance
of ordinary merit, and, indeed, would have done credit to any of the
learned Societies of this country. Most of them evinced no inconsi-
derable amount of original research. I think that man has missed a
very important element of happiness in common every day life who
has not cultivated his mind and provided for himself an intelligent and
rational pursuit, such as the study of literature, science, or art. You
who are working hard in your profession or business, pause before you
allow your life to slip away without the study of some subject which will
elevate the taste and lead you out of yourself for the time being, and
provide you with a mental tonic which will brace your mind with fresh
energies for your ordinary pursuits (hear, hear). It isa matter of great
importance that young people should have a knowledge of natural
history or natural science generally. It is a matter for congratulation
that those estimable persons who have undertaken the charge of
educating the young are awaking to a knowledge of this fact. Messrs.
Macmillan have, I notice, recently issued an admirable series of science
53
schoolbooks and science p ~ >
"superiority over the equilibrium theory, is still but an imperfect
‘representation of the real circumstances of the actual ocean.
__ Accordingly Laplace again attacked the problem from another side.
‘The law of the rise and fall of the sea may be broadly
stated as follows. If we suppose a circle traced on a dock
wall, so that the lower extremity of its vertical diameter may coincide
with low water mark, and the upper extremity with high water mark,
and if we divide its circumference between these points into six equal
_ parts, representing each one hour, then the tide will, as it rises, just cover
_ one of these divisions each hour, and similarly uncover one for each s
hour that it falls. A rule founded on. this is in practical use among a
seamen for finding the alteration produced by the tide in the soundings: fs,
_ marked on the chart. This motion of the water is what is termed a ss
simple harmonic motion. ee
_ ~ Let us suppose that, as the minute hand of a clock travels round E:
the face, a point moves along the vertical diameter, so as always to e “ig
‘keep on the same level as the extremity of the hand. This point then a
will have a simple harmonic motion. Let us suppose the hand and oe
‘the point to start from the figure VI. of the dial: at first, then, the —~__ >
ipward motion of the point will be very slow ; as the hand gets nearer _
to IX. the point will move faster, attaining its greatest speed when the —_— é
hand is at IX. ; its speed will then decrease as the hand goes towards 5
ee, Thus we see that the velocity of a simple harmonic motion is
= te circle on the desk wall in equal lines, its vertical rise in the same
_ time will vary greatly.
se ei!s ator just as the apeaivte One: in space of a man on hos? a
m otion relative to the earth, and of the earth’s motion in space, so . :
May conceive a motion compounded of several simple harmonic
otions.
_ Owing to the varying distances and positions of the sun and moon,
tide-producing forces which they exert are always varying in
ntensity ; for example, the varying distance of the moon from the
produces one variation in her tide-producing force, her varying
_ distance from the equator produces another. Now, Laplace was led
0 conclude, from his mathematical results, that each such periodical
iation in the tide-producing forces causes a corresponding variation
102
in the height of the tide, and that the height of the tide at any port is
the sum of these separate oscillations. This conclusion—almost the
only practical result of the dynamical theory—may be stated thus :
the height of the water at any place is the sum of the heights due to
several simple harmonic motions, whose periods depend on the
motions of the sun and moon. F
Each of these simple harmonic motions may be termed a tidal
constituent, or, for brevity, a tide. To render Clearer this complex
dynamical action, Laplace conceived each of these partial tides as
produced by a corresponding imaginary star moving uniformly in the
plane of the equator.
The periods of these tides being known, their other data are
determined from actual observations by the application of a profound
analysis.
A Committee of the British Association, under the presidency of
Sir William Thompson, is at present engaged in reducing series of
tidal observations made in different parts of the world. One of its
first reports was made at the meeting held in Brighton.
The larger a sea is, the more perceptible must be the phenomena
of the tides. In a fluid mass the impressions which each molecule
receives communicate themselves to the whole mass ; and thus it is
that the action of the sun, which is insensible on an isolated molecule,
produces on the ocean such remarkable effects. The Black Sea and
the Caspian have no tides, and the greatest range of the tide in the
Mediterranean is not more than four feet.
But a sea which is too small for the production of perceptible
tides by the sun and moon, may have derivative tides ; that is to say,
undulations which are excited primarily by the disturbing effect of the
luni-solar tide-wave, and subsequently propagated by mechanical
action among “the particles of water. The tides in the English
Channel are of this nature.
It appears a universal rule throughout the English Channel, that
at any great distance from either shore the tide-current runs up the
Channel nearly three hours after high water, and down the Channel
nearly three hours after low water; and that on the English side of
the Channel, especially opposite the entrances of bays, the direction of
the current revolves in 12 hrs. 20 min. through all the points of the
“—~
a ee a
=)
5 SAS I ne hn aia Oa een i
Sree bee Vere eae
103 4
compass, in the same direction as the hands of a watch, the observer’s a
face being directed up the Channel ; and that on the French side they 45.
. _ turn in the opposite direction. Thus, at some distance from shore, As?
we see that the time of high water does not coincide with that of slack. ie
This is well-known to sailors, who term the tide-current which flows P es
_ up Channel for three hours after high water tide and half-tide. ? ?
Nevertheless, they have often noted down the time of slack water as a
that of high water, and thus caused no small confusion and trouble in naa.
the discussion of their observations. then called Hermaphrodite ; an abnormal state of organism to which et.
os he would allude more particularly afterwards. hk,
_ On the subject of single or equivocal generation he had ‘but little - ‘es
4 ‘to say. By means of it only ie very lowest order of organism was
Drirred 3 in that of a particular species of louse, which originated upon
ae and collected in great numbers at particular spots.
— Certain species of acari were also developed in the same
.. z Professor Burmeister says :—“ The second kind of propagation, 5
mere ha at by shoots, has not yet been observed in insects ; it is also perfectly
{ itradictory to the idea of creatures so highly organised as they are. »
4 Some observations, however, seem to confirm the possible development of
wiki 4 .
*y f insects from germens or eggs laid by an unimpregnated female.”
it was of not unfrequent occurrence. This mode of propagation by :
hens constantly and regularly took place in certain genera. In
‘it occurred only occasionally. As a regular mode it was ascribed
wf)
T16
nation, according to De Geer and Bonnet, produced living female
young ones. ‘This spontaneous generation was repeated to the tenth
generation. After this a generation of males and females again made
their appearance.
On this subject he could not do better than quote Professor Von
Siebold, whose work on the subject of Parthenogenesis had been trans-
lated by Mr. W. S. Dallas. Von Siebold says: “It is well-known
that in the Aphides, a sexual generation represented by separate
males and females, is followed by a series of generations only in-
cluding a single form, which proceed from each other in manifold
repetition, without any previous copulation, until after about seven to
eleven such generations a generation of males and females again
makes its appearance.
“ Steenstrup regarded these forms of Aphides, which are capable
of reproduction without the influence of the male generative organs,
and which had been previously looked upon as virgin female Aphides,
as Nurses, and consequently as those members of an animal species
subjected to an alteration of generations which are capable of pro-
ducing young in the asexual state, Those Aphides which bring forth
living young, without a preliminary copulation, are in reality quite
different in their organization from the female Aphides, which lay eggs
capable of development after the act of copulation.
“In the viviparous Aphides those organs, specially from which
the living young are produced, have quite a different form and
organization from the sexual organs of the oviparous Aphides ;
so that in opposition to the ovaries, the products of which (eggs) only
become capable of development by the action of the male semen, we
may with perfect justice indicate the organs as germ stocks, which
are capable of producing young of themselves without the influence
of male fertilizing organs.
“‘ These nurselike viviparous Aphides therefore, which, instead of
ovaries, bear germ stocks in their interior, are also destitute of the
seminal receptacle which occurs universally in the female of insects
and plays an important part in the fecundation of eggs.”
The terms “germ stocks” and “ germ body” had been used to
distinguish the reproductive organs of the viviparous asexual Aphides
from the eggs and ovaries of the oviparous female Aphides.
Thus much on the subject of spontaneous generation of germ
y
4
ae peri . Se aus
117
mode of spontaneous generation by true virgin females. This mode
_ of reproduction had been called by the older naturalists “ Lucina sine
concubita,” and is now generally known as Parthenogenesis.
: Burmeister says “that unimpregnated females lay eggs may be
_ observed in the Bombycidee if, some days after their escape from the
io - pupa case, they be impaled and allowed to die slowly. The females
er the ine hadl ser the same ; but never the butterflies, according
a "This, of course, was known to be the case from the personal obser-
vation of all who had bred any of these insects. It must not be
supposed that these unfertilized ova as a rule produced larve. It was
indeed only very rarely that young were disclosed. As a rule these
eggs were sterile, but the exception to this rule was known as
Parthenogenesis or the generation from ova laid by unimpregnated
or virgin females. Y
according to the observations of such unquestionable authorities as
_ De Geer, Réamur, Bonnet, Schiffer, Miiller, Pallas, Ochsenheimer,
eRe
Roesel, and others, to be of not unfrequent occurrence.
In the Lepidopterous Genus Psyche, not only the Entomologists
ore named had observed it, but many now living could furnish
stimony in support of their statements. He would cite instances
is ir which this mode of generation had been observed.
pice €
Naturalists in 1701 a memoir, relating that he took a brown pupa,
which had spun itself up in a black currant bush, and preserved it
om. it. At the end of July a moth of a yellowish white colour
me caped from it (the moth was not more particularly described). ‘The
moth in a few days laid a great number of eggs and then died. In
pril of the following year, Albrecht again looked after the glass, and
‘ / ee 9 Geet ae Cen, Oe) 0 ee ae nl at
y oe eee eS WA, Te SS re ae re bas SP ea
dies by asexual individuals. He would next allude to the other —
_ The oldest communication on the subject was due to J.T. ‘ a,
brecht, of Hildesheim, who wrote to the Leopoldine Academy of.
ur nder a glass in his summer house to see what moth would be evolved
astonished at finding young black caterpillars in it instead of the
7
118
Querct-folia, related by Bernouilli as having been observed by Pro-
fessor Basler. _ Bernouilli himself observed an instance in Cceruleo-
cephala.
Treviranus observed an instance of the same spontaneous
development in S. Ligustri, Suckow in G. Pzz, and Dr. Nordman in
S. Populi.
Shirach stated that a queen bee would sometimes lay fertile eggs
without copulation with the drone, and that the females produced by
such eggs would again lay productive eggs without having copulated.
Professor Von Siebold had confirmed this statement of Partheno-
genesis in the honey-bee ; and, although originally very sceptical of
the occurrence of this phenomenon in the cases before mentioned,
subsequent observations convinced him that Parthenogenesis con-
stantly occurred in.some of the Tineidz, viz., in the Genus Taleeporia
or, more properly, Solenobia. The two species of Solenobia in which
he observed Parthenogenesis constantly to occur were S. Lichenella
and S. Tviguetrelia. The two species of sac bearers just mentioned
were not, however, the only representatives of true Parthenogenesis ;
an equally striking example of the virgin reproduction of a female
insect was presented by Psyche Helix.
Von Siebold had further established the constant occurrence of
Parthenogenesis in the case of Bomdbya Mori (the silkworm moth).
One more instance he would cite before closing this part of his
subject. He alluded to the Genus Cynips (or the Gall Flies). Of this
_genus some twenty eight species were known, which according to
Hartig were all destitute of males. Hartig was said to have inspected
9,000 or 10,000 of Cy#ips divisa and 3,090 or 4,000 of Cynips Soli
and found no single male amongst them. He even collected C. jolt
for eight years and never obtained anything but females, and he
observed these female Cynipidz proceed to the deposition of their eggs
immediately after their issuing from the galls. A male cynips had,
however, been subsequently discovered. Mr. Frederick Smith, of the
British Museum, the well-known Hymenopterist, writing in the May
number of the Entomologists’ Monthly Magazine for 1869, says,
“through the kindness of Mr. Darwin I have received both sexes of a
species of Cynips ; they were bred from the black oak by Mr. Benjamin
Walsh, the American Hymenopterist. The gall from which the male
and female were obtained is larger than the bullet gall of the oak, so
ea ee
o be an 119
“common in England, being two sitiched or more in diameter. According
to Mr. Walsh’s observations the males are only obtained from those
galls which develop flies early in the season, two months before the
is great autumnal brood appears, the latter all being invariably of the
3 female sex, Following up this we may hope this year to obtain males
of Cynips Lignicola.”
**
}
s
ob
a
__- It will be seen from the instances cited that Parthenogenesis cer-
tainly occurred more generally in the insect world than the few at
_ present discovered examples led us to suppose. Von Siebold was of
4 opinion that it occurs in accordance with determinate laws which had
_ hitherto entirely eluded our observation.
~.
>
In Nature definite objects were probably obtained by it which we
could only comprehend when we should have learnt to know the life
and actions of insects in general more exactly than was at present the
case.
E. . A Before commencing the subject of sexual generation, a few obser-
og _ vations were necessary on Hermaphroditism among insects. It was no
doubt well known that perfect Hermaphrodites amongst animals were
_ found only in the tape worm, many Annelida (for example, the leech
and the earth worm), and the majority of the Mollusca. In insects,
according to present observations, Hermaphroditism was but one-
_ sided, the left side generally exhibiting female forms and organs and
the opposite side male organs.
this subject were made known by Schiffer ; then Scopoli described an
meet s
_-Esyer described a specimen of 7. Crategi, in which the right side
BD Quercus. Capieux saw an Hermaphrodite of .S. Carfind, the left
= wing of which was male, but the right with the rest of the body
female.
: _ Ochsenheimer made a complete list of the Hermaphrodites which,
—
a
_ possessed,
‘He divided all Hermaphrodites into two groups, viz., into perfect,
w |
_ According to Professor Burmeister, the earliest observations on
stance. Communications of this kind then became more numerous.
was ‘male and the left side female. Then Hetthuger a similar ore of —
Pp to his time, had been described, or which he had himself seen or
’ in which one side was perfectly female and the other perfectly male, “4
d into imperfect Hermaphrodites, where the habit of one sex pre- ©
a ’
ay
eh a
ht
Pe
ey eee
iis tae oe
Le
Lp ie ry ES
iP Waa tte es
‘
"te
Rte, ae
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a Wl mw
any
ae > a
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6
PRES) Fe
yg bebe
120
vailed throughout the entire insect, and the forms of the other were,
perceptible in solitary parts. From this list of observed Hermaphro-
dites, made by Ochsenheimer, it appeared that in fourteen out of the
twenty-three instances there cited of true Hermaphrodites, the right
side was male and the left female.
Curiously enough, in the only specimen of a true Hermaphrodite
which he ever saw alive, and which was now in his cabinet, the right
side was female and the left side male, in exception to the general rule.
This Hermaphrodite was a specimen of Gonepteryx Rhamni, and the
greenish white colour of the female (right) wings contrasted very
strongly with the bright sulphur yellow colour of the male (left) wings.
Among the imperfect Hermaphrodites the majority, viz., six out
of eleven in Ochsenheimer’s list, were females, and the minority male
with female characters.
Scopoli seemed to have been of opinion that Hermaphrodites
sufficed to themselves, but subsequent writers on the subject appeared
to think this not to be the case.
It appeared that many of these so-called Hermaphrodites were
really nothing more than monstrosities, because on anatomical in-
spection they were found not to possess perfect female organs in
conjunction with perfect male organs, which must exist to constitute
true natural Hermaphroditism.
In conclusion he would allude to the third and regular method of
generation, which was called sexual. The methods of generation
previously alluded to being of course quite exceptional and abnormal.
Asa rule ail insects were of separate sexes and required the admixture
of both sexes in order.to be fruitful.
The union of the sexes by copulation was followed by the forma~
tion of the egg and the development of the embryo.
The differences of the sexes in size, form, and colour, were very
- striking in most species.
First as to size and form. It might be stated asa rule that the
body of the female was always thicker, larger, and generally more
convex. On the other hand the male was generally slenderer, smaller,
more delicately formed, and provided with longer legs. These
characteristics were particularly noticeable amongst the Coleoptera.
Another point worthy of notice was that the females of certain
a>
Tat
“species were invariably apterous, the males never so. In the Lepidop-
4 tera the females of the two species of the genus Orgyia amongst the
- Bombycidze were invariably apterous. So were the Psychide.
Amongst the Geometrz were the following genera the females of which
were entirely apterous, Phigalia and Myssia, and the following in
which the females were either entirely apterous or possessed wings only
i. _ Partially developed, Hybernia (including five species), Anisopteryx
- and Cheimatobia (two species).
ah,
_ Another peculiarity of sex was that the males of many of the
3 Peeibyces and Geometre had doubly pectinated antenna, whereas
_ those of the females were much less strongly pectinated, or were
. _ merely simple and setiform.
“ag
_ In the Genus Blatta, of the order Orthoptera, the females were
partially apterous.
- The differences of colour in the sexes were also very remarkable.
a rule (to which, however, there were some striking exceptions) the
colours of the male were brighter, more beautiful, and more glittering
_ than those of the female. In the Genus Lyczna, or Poly6mmatus of
the e Lepidoptera, the males were (with two exceptions out of 10 species)
Be beans shades of blue, whereas the females, with two exceptions
(in Argiolus and Arion), were of a dull brown. In Iris, as was well
own, the wings of the male were shot with splendid purple, but the
male was of a dull brown.
Biri One of the most notable exceptions to this rule occurred in Thecla
; Quercus, the males of which were dull purple throughout, whereas the
- fore ‘wings-of the female were enlivened by a patch of brilliant
_ The butterflies copulated about noon in the brightest sunshine.
The crepuscular and nocturnal insects copulated at those times.
copulation the male frequently died from exhaustion.
ms of the eggs lying on the tubes of her ovaries.* -After the de-
elopment of the egg was completed, the period for depositing it
OS: The impregnation of the female produced the development of the :
*
*,
it
122
immature egg. When the egg had been laid a distinct life, that of
the embryo, commenced in it. Externally the egg appeared to consist
of a horny shell, which got hard after exposure to the air. Beneath
the external membrane lay a second and more delicate one, which
formed the case of the fluid contained in the egg. The fluid was the
yolk, which was stated chemically to consist of albumen, animal glue,
yellow fat, and sulphate and phosphate of natron. According to
Suckon’s observations, a small dark spot was formed in the centre of
the originally tolerably clear yolk, which was the sign of the
commencement of the embryo. After a short time the em-
bryo appeared as a half moon shaped body, at the end of
which the head was already perceived. The embryo swam in
a bright green but clear fluid, and was enclosed by two other
membranes besides the shell. Michelotti’s experiments (quoted by
Burmeister) upon the eggs of Z. Dispar and L. Mari had proved that
the eggs during their development decomposed air, viz, imbibed
oxygen and gave out carbonic acid, but only in a temperature of from
15° to 20° (Reaumur). As the embryo increased in size, its external
formation could be seen through the egg shell. Burmeister stated
that the sexual organs might be observed during the last few days of
the embryo period and presented themselves in both sexes as small
knobs.
The commencement of the nervous system consisted of two
extremely delicate, scarcely perceptible filaments, into which the
nervous matter by degrees accumulated. The muscular layers beneath
the skin were also indicated, and particularly the head, with its
mandibles, the legs, and the anal horn.
The matured embryo, in its convoluted position, might imme-
diately before its development into the larval state be seen through
the thin egg shell. After these evolutions the young larva bored its
way through the egg shell, immediately commencing its first and most
important occupation, eating. The first meal was very frequently the
egg shell from which the larva had just emerged.
The President (Mr. J. DENNANT), on behalf of the members of
the Society, thanked Mr. Goss for his paper, and hoped it would not
be the last they would hear from him.
Mr. T. W. WoONFOR mentioned that, in 1869, some female
bullet galls which had been reared in the British Museum were
ua
*
Ar
rs
“
123
Mr. J. E. HASELWOOD said it had been noticed that the aphides
g the summer produced only females for ten generations, but
a Sania. To test this, some had been kept in warm atmo-
- sphere and supplied with plenty of nutrition, the result being that
“g ‘s ould not be completed in one as well as two. It was said that the
aries began to develop before impregnation, and when they reached
2 certain stage, if not impregnated, they decayed.
rave
_ Mr. WoNFOR said there was another point with regard to the
bides. There was a miscroscope slide commonly marked “leaf
sect,” i in consequence of it being covered with leaf-like appendages.
\ hat was merely one form of the maple leaf insect, and which also
ould be found on the sycamore. These peculiar females produced
ze species, one capable of producing their kind exactly alike, and
ae e other incapable of producing its kind.
- ger nination was the rule when the organism was not so developed ;
p> but when it became more complex bi-sexual re-production became
j ‘the tule, the latter having a greater tendency to sustain the form and
Ze give a longer duration of life than the other.
“iia
“A NEW SPINNING JENNY,”
a
is
. Mr. HASELWoopD did not see why all the necessary conditions
7
'
: sen thy
= It was remarked that Darwin had pointed out that uni-sexual
Mr. F. C. DENNET brought before the notice of the meeting a
Machine for spinning cotton that had been invented in America, ~ .
cing it with a statement of the futility of the attempts made i in this ae
NE >
wr Whe
ae te SS
Wn gh iwi eee tke
ee Ob at BO ee
1s
se ne eA
a ae
Dae Pe ae
hy
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» ni
hoa a
“y
peur
i
”
ed
it
a4"
t
‘
A
124
country to check the “ adulteration” of cotton. He claimed for the
“new machine” that it would not only effectually stop any tampering
~ with cotton, but would save half the usual waste, make stronger and
better threads, do away with a great deal of the present costly
machinery, and dispense with a great deal of labour. The machine
was worked on the ground where the cotton was grown, the seed
cotton being spun directly it was culled, so that it could not be
adulterated in any way. He produced some specimens of the cotton
for examination.
The CHAIRMAN observed that no doubt it was a clever invention,
but it was likely to damage the English and Scotch cotton trade.
Mr. F. C. DENNET did not believe it would have that effect, but
would be more likely to improve it, since it would send into the market
a really dond fide article ; a thread far better in every way than that
which could be made from bale cotton.
MAY 25TH, 1876.
MICROSCOPICAL MEETING.—MR. T. W. WONFOR
ON “THE SO-CALLED FUNGUS FOOT OF
INDIA.”
That fungi were able to induce disease was an accepted theory.
Their spores, being very minute, are able to penetrate the pores and
openings on the cuticle of animals and plants, and, having once found a
nidus, to thrive at the expense of the host into which they had insinu-
ated themselves. Nature, though, had fixed a barrier between the
sporule and the organism on which it might settle, and that barrier
was healthy vitality. The vitality might be impaired by disease or
old age, but when this did not exist, it had not, at present, been shown
that the organisms would necessarily succumb to the attack of the
fungus. It had been specially noticed that the less vitalized portions
of plants and animals were subject to and became the prey of fungus
growth. ‘This might especially be seen in the leaves of plants in the
autumn, and on the epidermal tissues, such as the wing covers, and
.
|
}
|
4
.
125
the articular plates of flies and insects, and the branchial plates of
_ fishes ; but though such was the case it remained to be proved that the
animals attacked were in a healthy condition at the time, or whether
the fungus did not the rather follow than produce the disease. In
other words, it was a moot point whether the bodies of insects were
ever attacked’ by fungus growth while the animals were in perfect
sg health. This was a very important matter in considering the nature of
the influence exerted by fumgi in the production and maintenance of
disease.
That the entire bodies of flies, beetles, wasps, bees, moths, and
other insects, when affected by fungus, were found, when examined
after death, permeated in every direction by mycelium, would bea very
significant fact, if we could determine beyond a doubt whether the
; tissues were not diseased before the advent of the fungus.
Since, then, it was by a large number an accepted opinion that
at that a very peculiar form of disease, met with in India, and believed
atthe time to attack omly the feet of some of the natives who went
about with bare feet, should, from a variety of causes, be attributed
to a fungus, the more especially as certain bodies found in amputated
"y _ Tins or in the matter discharged from limbs attacked, resembled and
_ were thought to be the resting spores of a fungus. Moreover, micro-
_ scopic examination and the planting these black particles so as to
enable them to, as was believed, grow, led men of the highest eminence
‘as mycologists to come to the conclusion that the cause of the disease
_ was a fungus, which they believed they had been able to rear from
particles obtained from undoubted cases of the so-called “ Fungus
Foot, ” or “ Madura Foot,” of India.
oe The first person to describe this disease as of vegetable origin
was Dr. H. Vandyke Carter, in 1860; the Rev. M. J. Berkeley
lescribed and figured the fungus in No. X. of the “ Intellectual
bserver,” vol. ii, November, 1862; while Mr, H. J. Carter, F.R.S.,
_ described it in 1862 in the “Linnzan Society’s Journal” and the
igh “6 ‘Annals of Natural History,” and in 1874 Dr. Carter published a
Bonoeraph on “‘ Mycetoma,” or Fungus Disease of India. All these
_and other publications went to prove that a fungus was the cause of
= disease. Dr. Carter had ample opportunities of examining speci-
ye nen. of amputated feet, and matter discharged from affected feet in
fungi induced and did not follow disease, it was not to be wondered
126
India, and separated the disease into two forms—one the d/ack and
the other the Za/e variety ; in addition, there appeared to be an inter-
mediate stage, characterised by the presence of pink granules,
The part affected was more or less distorted, and had numerous
openings communicating with channels of various sizes. The materials
which escaped from these openings contained brownish-black granules
like grains of gunpowder, in one variety, and roe-shaped ones in the
pale variety. The alteration of the tissues and bones caused a greasy
or waxy degeneration to the parts affected. Dr. Carter considered
these black particles the resting spores of a fungus, and the pale an
advanced stage of the disease due to a change—seemingly a degenera-
tion—of the darkened masses.
A series of experiments and observations with the black particles,
when mixed with cotton soil, placed on rice paste or moist ground rice,
both in India and in England, by Dr. Carter and Mr. Berkeley, led
to the conclusion that the cause of the disease was a fungus, to which
the name Chionyphe Carteri was given. The theory of the introduc-
tion was that spores becoming attached to the bare feet of the natives
germinated and sent in through the pores of the skin their mycelial
threads, hence the disease ; but the pink mould or fungus, of which
the black particles were thought to be the resting spores, had been ob-
served to grow without any connection with the black particles, these
black particles being the only substance associated with the malady in
which the existence of fungoid elements had been definitely established.
These particles too were on every occasion found to be unchanged.
Moreover, the pink mould grew just as luxuriantly in connection with
preparations which had been preserved in spirit as in connection with
specimens of the morbid tissues which had not been subjected to the
influence of any preservative fluid.
It was at this stage of the enquiry that Drs. T. B. Lewis and D.
D. Cunningham, who had been appointed special assistants to the
Sanitary Commissioner with the Government of India to conduct an-
enquiry into cholera, were deputed to investigate special diseases, as
well as continuing their cholera enquiry, turned their attention to that
form of disease known as “ Madura Foot” or “Fungus Foot” of
India.
The materials they had for examination were entire preparations
of the «pJer and /ower limbs —for both hands and feet, it turns out,
127
os:
234 are ‘attacked by this peculiar disease—as well as numerous smaller
preparations. They not only closely examined the condition of the
3 _ tissues and the nature of the morbid materials present in the various
q _ forms of the disease, but morbid tissues and products in other diseases
affecting similar parts of the body; they also made various attempts
at cultivating the morbid products as well as the study of the resultant
& _ organisms and the effects of re-agents on them and other vegetable
growths.
‘Inthe pale variety they found the bones were reduced to mere
Pisces of soft fat, the muscular and tendinous structures were well
preserved, but the most careful microscropic examination of all the
_ tissues and materials failed to afford the faintest evidence of the
q _ presence of any fungal or fungoid bodies, or of anything save degenera-
tion of the normal tissues. In a very early stage of this disease,
preservation of the muscles and tendons, less fatty matter, and the
"presence of the dark grains like gunpowder mentioned before. Here,
Bie no trace of fungal or fungoid substances was found in any part of
_ the tissues. Not only were both the pale and dark products subjected
to every possible test with re-agents and failed, save in some vague
2 points of form, to present anything which could suggest their vegetable
ieee Parasitic origin ; but in various cultivation experiments no fungus
: "growth was produced in any material in which the supposed spores
s were planted that was not also produced in some of the same materials
_ matter evidence any tendency to germinate or become altered in any
_ Similar experiments with the pale failed to give positive results ;
in fact, the fungus Chionyphe Carteri is unhesitatingly pronounced not
to cause the disease, and, moreover, that it cannot be developed from
»elements contained in the morbid products.
“In short, the “ Fungus Foot” is a misnomer, and does not owe its
toafungus. This is very important, because up to these ob-
tions, the opinion that the “ Madura Foot” was produced by
s has formed the basis for generalizations as to the supposed
of other diseases.
a in which nothing was planted. Never in any experiment did the black
128
The full reports, with the experiments and subjects operated upon,
will be found in the form of an admirable article in the t1th annual
report of the Sanitary Commissioner with the Government of India,
from the pens of Drs. Lewis and Cunningham, of which he had made
considerable use in the brief exposition of the so-called “ Fungus Foot”
of India.
Various engraved plates were used to illustrate the subject ; and
Mr. Wonfor made considerable quotation from the official report to
which he called attention in his paper. Mr. C. P. M. SMITH, Mr, H. E.
HASELWOOD, the President (Mr. J. DENNANT), and others, took part
in a short discussion which followed the reading. The point specially
discussed was which of these theories was correct—that germs in the
atmosphere attacked highly vitalised organisms, or only organisms in
a low state of vitality or debilitated by disease. At the close, speci-
mens of fungi, supposed to be the cause of disease in plants and
animals, were exhibited under the microscope.
Mr. WONFOR also submitted for examination the hard, earthy
cocoon of a beetle. It had been dug up by Mr. Brown, of Trafalgar-
street, in his garden on the Round Hill Estate, and it had been brought
under the notice of the Society through Mr. Haselwood. The cocoon
contained a larva ; and, as at one end it was partly broken, the insect
could be seen at work, It was placed on a table with the opening to
the light, when the grub was seen to be moving its head about, and
apparently endeavouring to close the opening by taking from the out-
side small particles of earth. This went on for a day and on the
following morning the aperture was found completely closed with a
small piece of flint, obtained from the exterior of the cocoon. The_
question whether or not the extra expenditure of energy and fatty
matter in repairing the cocoon would prevent the full development of
the insect,—-as was often the case where caterpillars had been com-
pelled from accident to make a second cocoon,—awaited solution in
regard to this particular case. The President said the point was so
interesting that he would suggest its reconsideration at another meet-
ing of the Society.
Beret oe ane Leth She ee tote aa eS
129 +
JUNE 18TH.
a ORDINARY MEETING.—MR. F. E. SAWYER, F.M.S.,
At ON “THE EROSION OF THE SUSSEX COAST,
. -WITH SPECIAL REFERENCE TO GREAT
STORMS WHICH HAVE VISITED tHe COUNTY.”
The present shape of the coast line of Sussex was mainly owing
to the two ranges of hills which traversed the County, and in a lesser
degree to geological causes and to the rivers. One of these—the
“South Downs, which in the western part of the County were about 10
Be Head ; the other—the North Downs, which, passing through Petworth,
4 Peeminated 3 in Fairlight Cliffs. The district situated to the south of the
. South Downs chiefly embraced the Tertiary formations, and it was this
osed of chalk had suffered less denudation than the alluvial plains to
_ its west, and thus the bay had been gradually formed. Another,
2 ‘though smaller bay, extended from Beachy Head to Fairlight Cliffs,
and this owed its origin to similar causes.
- At one time considerable estuaries of the English Channel are
aid to have extended to Lewes and to Bramber, and it was somewhat
difficult to conjecture why these have ceased to exist. In the time of
d probably, therefore, a considerable quantity of water made its exit
* hrough the valleys of the Adur and the Ouse. When, however, the
= forests were cut down the rainfall decreased, and consequently less
3 ‘water flowed down the valleys, and a gradual silting up took place.
He should surmise that this had taken place at the latest by the ca
9, if not much earlier. Hayley, in his “ History of Chichester,”
4 ‘states that in the time of the Romans the Lavant flowed entirely round
oat he City of Chichester, whereas it now flowed round two sides only,
os and that it was probable that the bed of the river was, at an early
: period, diverted by an earthquake shock. He (Mr. Sawyer) was further
inclined to think that since the time of the Romans the whole of
the coast of Sussex west of Beachy Head had been slightly raised
__ by an earthquake shock.
_ There were no very early records of storms on this coast. Tegg’s
miles distant from the sea, terminated in the bold headland of Beachy —
district which had suffered most from the sea. A long but shallow bay ~
sxtended from Selsea Bill to Beachy Head. The latter being com-.
‘the Romans the Weald was covered with large forests and lagoons, —
sh hy
Soe SAN hae SOR S's os
4
nls
Sis 43
eA 2 ae
»
130
Chronology mentions a “storm on the coast of Kent, Sussex, and
Hampshire, in 566 ;” and a “ Terrible hurricane, at Chichester” was
recorded in 995; but no particulars were known respecting them. The
earliest storm of which any definite account existed occurred on
October Ist, 1250, and was thus described in a manuscript formerly in
the possession of Mr. Thomas Martin, of Winchelsea :—
“The moon being in her prime, the sea passed its accustomed
bounds, flowing twice without ebb, and made so horrible a noise that
it was heard a great distance inland, not without the astonishment of
the oldest man that heard it. Besides this, at dark night, the sea
seemed to be alight and to burn, and the waves to beat with one
another insomuch that it was past the mariners’ skill to save their
ships ; and to omit others, at a place called Huckebourn (Eastbourne)
three noble and famous ships were swallowed up by the violent rising
of the waves and were drowned.” *
Holinshed’s account was somewhat similar :-—
“ On the first day of October (1250) the moon, upon the change,
appearing exceedingly red and swelled, began to show tokens of the
great tempest of wind that followed, which was so huge and mightie,
both by land and sea, that the like had not been lightlie known, and
seldom, or rather never, heard of by men then aliue. The sea, forced
contrarie to his natural course, flowed twice without ebbing, yielding
such a rooring that the same was heard (not without great wonder) a
farre distance from the shore. Moreover, the same sea appeared in
the night to burn, as it had been on fire, and the waves to strive and
fight togither after a maruellous sort, so that the mariners could not
deuise how to save their ships where they laie at anchor, by no cunning
or shift which they could deuise. At Hertbourne, three tall ships
perished without recouerie, besides other smaller vessels. At Win-
chelsey, besides other hurte that was doone, in bridges, mills, breakes,
and banks, there were 300 ae and churches drowned with the oe
rising of the water course.’
This storm diverted the course of the river Rother, which now
entered the sea near Rye, but before the storm had its exit to the east
* Horsfield’s History of Sussex,
131
“4 Gor 1 Dungeness, 12 miles eastward of the present mouth. Camden,
= after treating of Hastings, says ;—
d . jm Hence the shore retires backwards and is followed inwards,
_ being full of windings and creeks, within which stands Winchelsea,
Pi built i in the time of King Edward I., when a more ancient town of the
i same name was quite swallowed up by the raging of the tempestuous
_ ocean, in the year 1250, at which time the face of the earth, both here
and i in the adjoining coast, was much altered.”
4 - Shortly after this storm, and before the town of Winchelsea had
recovered from the injuries it had received, another storm occurred
__ which was described ina Latin memorandum found by Jeakes amongst
__ the Rye records, and thus translated :—
“ Be it remembered, that in the year of our Lord, 1287, in the
ven of St. Agath, the Virgin, was the town of Winchelsea drowned
oe and all the lands between Climesden and the Vocher of Hithe. The
same year was such plenty of corn throughout all the countries of
_ England, Scotland, and Wales, that a quarter of wheat was sold for
_ two ee a
In the year 1292 Pope Nicholas thé Fourth granted the tenths
f all the ecclesiastical benefices in England to Edward I. for six
for defraying the cost of an expedition to the Holy Land, and a
; mplete valuation was accordingly made. About the year 1340 the
= 2 ninths of wool, &c., were granted to Edward III. A careful enquiry
yas made in each parish, and the returns afforded a complete record
f what changes had occurred since Pope Nicholas’s Taxation. These
nquiries were called “ Nonarum Inguisitiones.” From this source we
it that between the years 1292 and 1340 no less than 5,500 acres
in 1 this County were submerged by the sea! The None return for
ie 340 for Brighton (made 1341) says :—
e* Ba “Tr dicunt qd ix ps pdea no respondet nc attinge potest ad taxam
ecclie pdce p eo qd xl acr’ tre submis sunt p mare imppetuu que valuer
nu x1.s,”
whi ich might be thus translated :—
“The same (the jurors) say that the 9th part aforesaid cannot
er nor attain to the taxation of the church aforesaid, for that 40
of land are submerged by the sea for ever, which were worth
r annum 40s.”
ae’
= The return for Hove stated that 150 acres of land were submerged
.
iy
sth
7
*
mee
, a
Ligeia amp A ities yon Sot as Eee aaa
he
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132
in that parish worth ro marks a year, and at Aldrington and Middle-
ton 40 acres were inundated. Early in the 14th century (Dallaway)
in his “History of Chichester,” says not many years before 1345.
Pagham Harbour was formed by a sudden irruption of the sea by which
2,700 acres of land were devastated. Inthe year 1432 the inhabitants
of Shoreham petitioned the King for a reduction of their taxes, “ for-
asmuch as by the encroachment of the sea, &c., the number of in-
habitants was reduced from 500 to 36.” “A sudden incursion of the
sea at Rye, in which the water rose 8 or 9 feet in men’s houses at mid-
night” was recorded in 1570. Elliott’s manuscript says :—
“In Elizabeth’s reign the sea in a storm broke through the beach
bank at Bishopstone and formed what is now called the Old Harbour
which was in use till the Newhaven one was made a safer exit. This
may have been the same storm as deepened so materially Rye
Harbour.”
This was the second instance of the course of a Sussex river being
diverted by a storm. Formerly, it might, however, be mentioned, the
mouth of the river Adur shifted from year to year, going gradually
farther eastward towards Brighton, and then back again towards
Shoreham. The following interesting account of a storm on Novem-
ber Ist, 1597, was from the Hastings Corporation Records. The pier
there was destroyed by the sea, and in 1595 the work of re-building it
was begun, but all the works were washed away and a second attempt
made, which, it would be seen, also failed.
* This woorke was with singular industrie and arte brought above
the full, and by all Hollantyde, 1597, were nere finished, viz., XXX foote
high, and C foote at least long, bewtyful to behold, huge, invincible,
and unremovable in the judgment of all the beholders, amounting to a
great charge, whereunto the whole shire and div’s beholders were con-
tributoryes of benevolence, besides the townes great expences. But
behold when men were most secure, and thought the woorke to be per-
petual, on All Saints’ daie, 1597, appeared the mighty force of God,
who with the finger of His hand, at one great and exceeding high
spring tyde, with a south-est wynd, overthrow this huge woorke in less
than a hower to the great terror and adasmt of all beholders, to the
great discredit of the lyke woorke hereafter with the country, and to
he manifest undoing of the town.”
In the following century Brighton suffered severely from the en-
he.
croachments of the sea. Godwin’s rental of the Manor, made in 1665,
“stated that previous to that year no less than 22 tenements under the
_ cliffs were destroyed by the sea, and between 1645 and 1655 frequent
grants of land were made to tenants who had suffered from the inundation,
and in one case a tenant lost eight acres. In 1665 there remained, how-
ever, 113 tenements under the cliff. The next storm which visited the
county was that known as “the Great Storm,” which occurred on
November 27th, 1703. It was one of the most disastrous storms ever
recorded, and indeed some writers had stated that it was the most
Spee storm since the Deluge! Defoe who published a book on
espe storm, says :—
“At Shoram the Market House, an Antient and very strong
ilding, was blown flat to the Ground, and the town shattered.
_ Brighthelmston, being an old built and poor tho’ populous Town, was
_ “The Bishop of Bath and Wells and his lady were killed at
ells, Somersetshire, by a wall, which was blown down, and ‘ the
hop of London’s sister, Lady Penelope Nicholas, was killed in like
nner at Horsley in Sussex, and Sir John Nicholas, her husband,
_ The same book contains the following curious and quaint apie
t — the effects of the storm in a letter to Defoe :—
ar From Medhurst in Sussex, the following Letter is a short
account of the loss of the Lord Montacute there which is extraordinary
« one in and about our Town. I praise God we came off indifferent
e e of them fell on part of the Great Hall, which did considerable
age ; and the Church Steeple of Osborne, half a mile from us, was
hn Mills, a oil pace Tiled Barn.
“Your humble servant,
eviously hurt.’ "—Milner's Gallery of Nature. -
sreat, tho’ abridged in the letter :-— Sir, I received a letter from you,
herein you desire me to give you an account of what damage was —
; the greatest damage we received, was the untiling of houses, and _
ax down at the same time ; and fa Lord had above 500 trees torn >
“Medhurst, Jan. 18, 1703-4. : “JOHN PRINKE.” ~ :
Aart CS Mohs
em Th Mees ORE
s
\! ‘
'd
PROT rae
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134
The following was the record of the storm in the Brighton Town-
book :— ;
* Memorand.—November 27th, 1703, there was a very great and
remarkable tempest, which begun after midnight, and continued in its
violence till about eight in the morning, being Saturday. Many houses
__ in town were damnified, two windmills in the east blown over, several
of the church leads turned up, and several vessells belonging to the
town were shipwracked, to the great impoverishment of the place.”
The rain which accompanied the storm appeared to have been
very salt, as would be seen by the following description from the
Philosophical Transactions :—
“ Part of a letter from John Fuller, of Sussex, Esq., concerning
a strange effect of the late great storm in that County.
‘ December 6, 1703.
‘We live Ten Miles off the Sea in a direct line, and yet can
scarce perswade the Country People that the Sea water was blown
thus far, or that during the Tempest the Rain was salt, for all the
Twiggs of the Trees the day after were white, and very salt, as I am
informed almost by every body, tho’ I did not taste them enough
«myself, nor observe it, and that not only upon this Hill where we live
facing the Sea, but in all other places within 14 or 15 miles of the Sea,
as well as in the Valleys between which and the Sea are several very
high Hills, as on the Hills themselves.’”
No doubt Mr. Fuller was unable to make the people believe that
the rain was not salt when they could taste it, and indeed salt rain was
not so very unusual, for it had been observed on several occasions.
Another account of this salt rain was also given in the Philosophical
Transactions, by a clergyman at Lewes :—
“A Physician, travelling soon after the Storm to Ticehyrst, about
20 miles from Lewes, and as far from the Sea, as he rode he pluckt
some tops of Hedges, and chawing, found them Salt. Some Ladies of
Lewes hearing this tasted some Grapes that were still on the Vines,
and they also had the same relish. The Grass on the Downs in his
Parish was so Salt that the Sheep in the morning would not feed until
hunger compelled them, and afterwards drank like Fishes, as the —
‘Shepherds report. This he attributes to Saline Particles driven from
the Sea.”
iagrs
The Gestation of the undercliff town of Brighton, which was
_ Brighthelmstone ” shews : —
> ‘s “ Another dreadful storm reached this town about one o’clock in
the morning, but raged not with its greatest fury till after three, and
then continued with unremitting violence till the hour of eight. It
_ destroyed a great many houses in the town, and blew off most of the
2 church leads. Every habitation under the Cliff was utterly demolished,
and its very site concealed from the owner’s knowledge ee a
ound of beach.”
a . He might then remark that there was a very prevalent idea that
the town of Brighton formerly stood about as far from the shore as the
_ Chain Pier Head was. From a careful study of the plans, Court rolls,
_ &c., he was convinced that this was an erroneous idea, and he thought
that within the historic period the coast line was not more than 200 feet
e - from the present line. Another exploded notion was that the Block
. House stood in the centre of the town. Horsfield stated that in 1734
ee the Gun Garden adjoined the Block House.” | The Burrell MSS.
3 ecorded i in January, 1749—“ By reason of extraordinary high tides the
i 1 the farm lands called Salts, and did considerable damage to the
land adjacent.” On January 30th, 1775, a portion of the Battery at
_ the end of East Street was washed away, and on November 3oth, 1786,
a egan to take steps for sevens further encroachments of the
‘sea. This was the first time in the history of the County that we found
a etod of an attempt to keep back the sea. “ Magna Brittannia”
(put plished i in 1738), says that at a moderate computation the damage
-sust: 1 ined by the town was £40,000. It was found that the tendency
of the tide was to roll the beach from west to east, and it was deter-
mi ed to erect wooden barriers (called groynes) stretching from the
to low water mark. The origin of the term groyne was very
an.
reat Seal, issued briefs by which collections were made in the
Bermenced by this storm, was completed by another storm on August
ie ih SAYS
et oe mn Via
Pre
sea broke in at Brighthelmstone, washed away part of the Block House |
are. The inhabitants accordingly, being too poor to provide the ©
ek Ft
1ey for erecting groynes, in 1722, by virtue of Letters Patent under —
may
Pn Vs,
w-
> A
a
os.
re hi
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i, roe
ae Fy
ASG
e
136
various churches and chapels throughout the County for this object.
. About £1,700 was raised, and, aided by local contributions, the town
was protected as far as the Steine.
During the present century much has been done in the way of
groyning, and the coast of Brighton, instead of being washed away,
was advancing farther out to sea, and the concrete groyne opposite the
Clarendon Hotel appeared to have had a very great share in the work
of accumulating the beach to the west of it, which had rendered it
possible to widen the King’s Road once or twice. The effect of the
groynes was somewhat peculiar ; on their western side a great heap
of beach was formed, while on their eastern side the beach was from
10 to 15 feet lower, and on this side the sea rushed up with great
violence. This was specially noticeable in the heavy gale of March
12th, 1876. It might be considered whether this could not be obviated
by making the groyne V-shaped or by putting posts or buttresses on
the eastern side, which would break the force of the waves.
At the present time there seemed to be two weak places on the
coast, viz., at Lancing and beyond Pevensey. He had an impression
that the cause of the special attacks of the sea at these places was
very similar to that of the sea on the eastern sides of groynes. The
neighbouring promontories of Worthing Point and Beachy Head
respectively acted very similarly to the groynes and collected beach on
their western sides, while the sea rushed up on their - eastern
sides.
One of the most destructive storms during the present century
occurred on November 22nd, 1824. A brisk wind sprang up at night
from S.W., and increased all night. Towards daylight of 23rd it
became extremely calm for a short time but increased with the tide. ~
High water was at one p.m., when the gale was tremendous, and it ~
continued without abatement till midnight. At Brighton the platform
of the Chain Pier was washed away and forty-houses between Royal —
Crescent and Kemp Town were washed down. The Arunand the Ouse ©
overflowed their banks. At Littlehampton the sea broke over the —
beach and inundated the fields; the damage between there and Pag-
ham was estimated at £15,000. At Pagham and Sidlesham great
damage—loss £30,000. At Bosham the mud wall, which cost £20,000, —
was washed away, and at Emsworth the tide was three inches higher j
than was ever known. In conclusion, he must apologise for the —
137
a The PRESIDENT (Mr. Dennant), on behalf of the members,
Beyeepked Mr. Sawyer for his admirable paper, and expressed a hope
“that it would lead to an interesting and profitable discussion.
aa Mr. E. WILLETT pointed out that the tendency of eanhquakee
& - was rather to uplift than to sink the coast, but that it was pretty clear
_ from observations which his father was making, and of which he could
P _ hot say more at present, that the coast line of the county was gradually
and very considerably sinking. Allusion was also made to the present
theory of groyning, which he (Mr. Willett) thought was wrong. By
__ adopting it, there was as much wash on the lee side of a groyne as
gt was beach on the windward. If the groynes were made to lean
a little to the south-west, kept very low, built up as the beach accumu-
F inted, and constructed in zig-zag fashion, a natural barrier to the sea
_ would’soon be secured. These were ideas of his father, and had been
~~ successfully enforced elsewhere.
_ Mr. Wonror remarked that it was believed the whole of the
southern coast of England was sinking, and that the time would come
when Brighton and the adjacent dow-lying parts would return to the
5 "sea. This prospect was so alarming to some people that one gentle-
m an whom he knew had determined not to purchase a house on the
_ Brighton Cliff, being convinced that although he should not witness
_the disappearance of the coast-line, his grandchildren might.
_ Encouragement for such a conviction was offered in the fact that with-
‘im the memory of some old people there were two roads from Brighton
to Rottingdean south of the present. The silting up of rivers and
rbours in this part of the country was also referred to; and it was
dicted that cither greater local energy than was at present shown in
se ose
Now at (Eningen the Carnivorous Coleoptera are to the
total number. of species of that order, in the proportion of
1: 4:62, and to the Herbivorous species of that order, in the pro-
portion of 1 : 3°62; that is, they are rarer in this locality than in
Europe at the present day, although they are much more
numerous than they are at the present day in tropical countries,
especially in South America.
The fact that the Carnivorous species are in such a decided
minority to the total number of species of the order compared
with the proportion existing between the two classes at the
present day, stamps the fossil fauna of (Eningen with even a more
southerly character than one would have expected from the
flora. From this it may be safely assumed that the climate of
this part of Europe during the Upper Miocene period approached
much nearer to that of the Tropics than at the present day ; and
this assumption is materially strengthened by a consideration of
the present Geographical distribution of many of the genera and
-species then inhabiting Giningen.
93
According to Dr. Heer and M. Oustalet the majority of
the Ciningen species belong to genera distributed at the present
«lay over the Old and New World, and these genera are said to
contain nearly two-thirds of the total number of species dis-
covered in these strata. Heer estimates the proportion which
these genera bear to the total number of genera to be 114 to
180. Amongst the other genera, according to M. Oustalet, two
axe not found at the present day out of Africa, two are specially
American. Some are met with both in Africa, Asia, and
America, particularly in the last named country.
A great number of the species of this period are allied to
existing American species. Seventeen genera still inhabit the
Old World ; seven of them, as well as those which are exclusively
European, are still represented at the present day in the
Mediterranean fauna. On the subject of the insect fauna of this
locality Sir Charles Lyell observes:—“Few of the genera of
Insects are extinct, but many of them imply a Geographical
distribution widely different from that now obtaining in the
_ same part of the world.”
On the whole, Dr. Heer found that the results furnished by
an examination of the Insect fauna of (Eningen agreed with those
derived from a study of the flora, although the character of the
flora is more strictly American and more. tropical than that of the
fauna, and induced him to believe that the summer climate of
4Eningen was not tropical, but that the winter climate was
extremely mild.
} To this period (Upper Miocene) Dr. Heer refers certain
strata at Parschliig in Styria, Austria, from which he enumerates
fourteen species of insects, viz :-—
Coleoptera 2 Ait re 7 species
Orthoptera wad wat ea OA.
Neuroptera Tis es,
Hymenoptera Sashes
Diptera Dk ibs
el
re
94
The classical deposits of Senigallia, in the north-east of
Italy, also belong to the Miocene period. These deposits, so:
famous for the quantity of animal and vegetable remains
detected therein, contain, says Signor Mas:alongo, numerous
impressions of insects. It was from these strata that Signor
Procaccini-Ricci obtained a considerable number of species,
which afterwards formed part of the collection of Signor Giuseppe
Scarabelli di Imola.
Signor Massalongo describes two fossil larve of genus.
Libellula, which he states that he found by chance in splitting
open some slabs of stone from this locality. He has named one
species Libellula Eurynome and the other species Libellula Doris.
With this exception I have failed to meet with the description of
any fossil insects from Italian strata of this period,
MippLe MI0cENE.
To this period are referred the Marls of Radaboj in Croatia
which have been productive of a great quantity of insect remains.
Professor Unger was the first author who published (in 1839)
on the fossil insects of this locality a Memoir of any importance,
This was followed, in 1843, by a paper by Von Charpentier (in the
Acta, Acad. Leop. Carrol). Since then Professor Heer has-
thoroughly studied the fossil insect fauna of Radaboj, in which
he has recognised a great number of tropical forms including some:
gigantic Termites.
The insects from this locality comprise the following :—
Coleoptera Se oe oe 42 species.
Orthoptera ey ae ae 13" ste
Neuroptera = LS ies 20 sg
Hymenoptera... = = 83\> 35
Lepidoptera ke ees AM a Soe
Diptera... Le eee 3 3 ,
Hemiptera 4 is — 2 55
Orthoptera 4 43 —2 ,;
Neuroptera 2 2, —2 ,,
Quite recently Mr. Scudder has published a description of
two species of Orthopterous insects from the Rocky Mountain
Tertiaries. The first species Mr. Scudder has described as
Homeogamia ventriosus and the latter as Labidura tertiaria.
Since the date of the last mentioned paper, Mr. Scudder has
published detailed descriptions of 31 species of Coleoptera, most of
which were included in those before referred to as having been
found by Professor Denton and Mr. Richardson, and a few others
of which were found by Mr. Mead or by members of the United
States Geol. Survey.
“Tn this paper,” said Mr. Scudder, “are made known the
first fossil Coleoptera from the Tertiaries of the United States ;
Pe ene ts
109
indeed, if we except some doubtful remains in the red sandstone
of the Connecticut Valley, the first distinctively American
Coleoptera from any formation. Two beetles have been figured
by Heer from the Miocene of Northern Greenland, and these are
all that have yet been described from the New World.”
These valuable contributions to our knowledge of the fossil
insects of the American Tertiaries were alluded to by Professor
Westwood in his anniversary address to: the Entomological
Society of London on the 7th February last, who also stated that
Mr. Scudder had published “the descriptions of several fossil
species of Thripside from the North American Tertiaries,
including two new genera, Lithadothrips and Paleothrips.
Mr. Scudder has very recently described several species of
insects obtained by Mr. G. M. Dawson from tertiary beds at
Quesnel, British Columbia. In addition to fragmentary indeter-
minate remains, there are some 24 species distributed among the
undermentioned orders, as follows :—
Coleoptera... — ... as x 1 species.
Neuroptera td pre mA Koy 33
Hymenoptera... ae doe ds 95
Hemiptera... Red 3 ear 1 es
Diptera... an Riad se ENS!
24
AMBER.
Before closing this paper I must say a few words about
amber and the insects which have been detected in it.
Nowhere have fossil insects been found in greater numbers
or in a more beautiful state of preservation than in this “resinous
and bitumenous substance.” By most geologists amber is con-
sidered as antediluvian, but its origin has been the subject of
much dispute. Some ancient writers attributed it to ants, and some
considered it to be of mineral origin, It is now considered by all
authorities to be of vegetable origin, and to have been formed
from the resin which distilled from certain species of fir trees of
the Tertiary epoch, whose fossilized trunks are said to form the
lignite in which it is frequently obtained. Dr. Berendt and
110
M. Goeppert are said to have discovered proofs that the Prussian i
Amber is the product of a fir tree, the species of which is now
extinct. In some cases the leaves of the Conifere have been
found in amber, and although, according to Dr. Berendt, they
appear to differ from all known species, he thinks himself
justified in ascribing them to the genus Pinus.
On this subject the Rev. F. W. Hope, F.R.S., says, “In a
letter lately received from Dr. Berendt he informs me that the
anotomico-microscopical examination of the wood places it beyond
a doubt that the amber was a Pinus; but of what species cannot
be now known, and that Pinus Balsamea approaches nearest to it
in appearance.”
The period during which this species of fir tree existed is
supposed to have been about the commencement of the Tertiary
epoch. The lignites or Brown Coal formed by it are superior to
the Chalk and inferior to the Tertiary and diluvial gravels and
sands of Germany.
Amber is said to be occasionally thrown on the beach of the
East Coasts of England along with masses of jet, and if not torn
from the bed of the sea may be washed from the Baltic, where
there are regular mines of it. It is also abundant on the shores
of Sicily and the Adriatic Sea and also in the beds of lignite or
Brown Coal in Germany, and is occasionally found in Spain and
France. Dr. Berendt states that he is of opinion “that the
geographical focus of the amber wood was at the bottom of the
Baltic.”
The insects at present detected in amber appear to relate to
species extra European, many of them belonging to tropical
climes, while some approach South America and Indian forms,
They do not appear related to any existing species, and are there-
fore probably extinct, and this seems to be the opinion of Germar
Jussieu, De Jean, Hope, Dr. Leach and others, On this
subject the Rev. F. W. Hope says, “The major part of the
insects discovered in amber exhibit a close resemblance to
111
existing species, and can be satisfactorily classed under published
genera. That any of those which are found in amber are identical
with existing species I do not believe, for out of many hundred
specimens, nay, I may say thousands, which have fallen under my
notice, none have yet induced me to change my opinion that they
are otherwise than of the Tertiary period.”
The number of species of the different orders of insects dis-
‘covered in amber, and according to the list at the end of Mr.
Hope’s paper on “ Succinic Insects” and the investigations of
Sendelius, Berendt, Germar, Bronn, Gravenborst, Ehrenberg,
Pictet, and other writers on the subject, is as follows :—
which includes :—
species from Amber, Waa
Mr. Hope also gives a list of
Of the Coleoptera, about 160 species.
,, Hemoptera, as 11 iy
;, | Neuroptera, - 10 3
», Hymenoptera, ;, 25 34
5, Orthoptera, 35 12 3
» Hemiptera, 4, 15 Ri
,, Lepidoptera, ;, 10 by
», Diptera, i. 22,
species from Copal and Auimé,
Coleoptera 9 species.
Hymenoptera 2 A
Hemiptera 1»
12
In his “Fauna der Vorwelt,” Dr. Giebel enumerates 318
Coleoptera 106 species.
Hymenoptera 14 oe
Lepidoptera 3 a
Diptera 141 .
Neuroptera 28 FE
Orthoptera 4 o
Hemiptera 22 35
318
One of the best accounts of the history and origin of amber
is given by Professor Zaddach of Kénigsberg, an abstract of which
in English, appeared in the Quarterly Journal of Science, for April,
1868; and since then the Rey.
P. B. Brodie has written an
112
admirable paper on the subject of amber and the fossil which its
contains, which was read at the March meeting of the Warwick-
shire Naturalists’ Field Club, in 1871.
The lecture was attentively listened to, and heartily
applauded at the close, and, at the request of the PRESIDENT, a
cordial vote of thanks was awarded to Mr. Goss for his paper.
The very patient attention the meeting had given to it proved
that the research which the lecturer had evidently made was
thoroughly appreciated, and they could not but be surprised
at what he had revealed.—Mr. T. W. Wonror (one of the hon.
secretaries) also expressed his gratification at many of the
matters which had been brought to light by Mr. Goss. Mr.
Goss having announced that the lecture he had given would be the
first of a series of three, which he intended to publish as soon as
they were delivered, Mr. B. LomMAx expressed his opinion that
the series would constitute a most important geological volume.
The conversation was continued by Mr. Dennet and Mr.
Dowsett.
MARCH 22ND.
MICROSCOPICAL MEETING.
This being a general evening objects of interest were shown
under microscopes by Mr. G. D. Sawyer, the President, Mr. T.
W. Wonfor, one of the hon. secs., Mr. S. Aylen, and Mr, A.
D. Michael. Mr. Sawyer exhibited, among other things, a
section of stellate tissue, eggs and larve of moth, and elytron
of the diamond beetle ; Mr. Wonfor, larve of the ant lion, a
lobster insect found in melon pits, and two species of the gnat ;
Mr. Aylen, a shaving of the horn of the rhinoceros ; and Mr.
Michael, coralines in fluid, with extended tentacles, and diatoms
in chain. At a meeting of the Committee of the Society, held
previous to the microscopical meeting, it was decided to offer a
prize of five guineas for the best botanical collection in the
113
following orders, made by residents in Brighton, Hove, and
Preston :—Ranunculacee, Papaveracee, Crucifere, Violacex,
Polygalacez, and Caryophyllacee.
APRIL 12TH.
ORDINARY MEETING.—DR. CORFE ON THE STRUC-
TURE AND DINETICAL CURRENTS OF SPONGE-
LETS AND FIBRILS OF ROOTS.
The object of his communication to the Society was to give
publicity to a series of microscopical appearances on the structure:
and dinetical currents of spongelets and fibrils of roots, which,
whilst they confirm the researches of eminent phytologists on
cyclosis in leaves, stems, and hairs, appear to have escaped
attention in this field of life-germs.
If it is asked, “‘ What inscrutable power presides over the
intricate machinery of vegetable and animal life, of elaboration,.
digestion, and assimilation, whether it be in the tiny weed we
tread upon or the colossal form of an elephant?” we reply
unhesitatingly, electrical force—a force which man has recently
enlisted in his service and has so utilised that he conveys his
thoughts in a few seconds to his fellow-men through a space as wide-
apart as the Arctic and Antarctic seas. He would remind them
that the surface of the globe and the bowels of the earth afford
two mighty currents of this force; the one is moving from us on
the eastern side and is at right angles to the magnetic meridian,
or parallel to the equator, and corresponds to the negative pole of
an electro-magnetic battery; the other, on the Western side, is
moving ‘owards us, and answers to the positive pole, and is.
parallel to the straight line which joins the poles. Every
substance above, upon, and below the earth’s surface contributes
its quota to these currents, some so highly that they are styled
“Paramagnetic,” by Farady; others, being feeble, he called
“ Diamagnetic,” and thus the bosom of our globe is one stupendous.
114
-and infinite reservoir of electrical fluid, oxygen in gases, iron in
metals standing highest in the former, hydrogen and potassium
in the latter.
In contemplating the whole cycle of animal life, they were
at once met by wheels within wheels ; thus they had the vascular
cycle within the skull of a mammal; again, that within the liver ;
and another within the kidney—each one differing from and
independent of the other.
Without further preface he would direct their attention to the
dominant cycle and mainspring of the whole fabric—the venous
blood, ere it reaches the pulmonary heart. What was it they
found?
H u s w
Year.| -25 AS As AS
he ile A hee
o 75 OD Ts O é
co) r= ae ra) FS o) a
1857 | 47
1858 | 79
1859 | 91
1860 | 87*
1861 | 123
1862 | 93 60 53
1863 | 97 90 60
1864 | 59 91 69 32
1865 | 92 81 76 39
1866 | 88 110 86 75
1867 | 76 95 87 61
1868 | 98) |121) 106) 82)
1869 | 111 124 113 74
1870 | 106 111 98 81
1871 | 91 89 114 77
1872 | 71 $92)117 | 105/101 ;+ 102) 96/81
1873 | 83 100 78 74
1874 | 92 98} — |104 81
1875 | 90 92 97 80
1876 | 86) 93 103 84
* Supposed ears imperfect from wet season.
oe ___
Total
Contents of the grains.
original earof each variety 47 60 53° 32 =192
at starting.
Average contents of
best ear annually of
each variety 92 105 102 $1 =380
throughout last 9 years J
of series.
The average productive powers of each kind of wheat is here
clearly doubled.
In conclusion I will briefly explain the practical bearing of these
results. The bushel of ordinary wheat containing about 700,000
grains, by the present practice of sowing two bushels of wheat,
nearly one and a half millions of grains are deposited upon an
acre of land. The number of ears produced upon an acre
never equals this, Whether we plant single grains one foot
135
apart each way, or nine inches apart each way (if we plant
them sufficiently early), or whether we sow one bushel, two
bushels, or three bushels per acre, the number of ears produced per
acre is practically the same. In June, 1874, I was on a visit to a
friend farming some 4,000 acres in the Vale of Eversham, who had
two plots of my wheat—the one planted with single grains twelve
inches apart every way (but not early enough), the other with
grains nine inches apart every way. The ears were magnificent,
but everyone talked of it as the “thin” wheat. He was much
astonished upon my telling him that there were as many ears per
acre on the plot planted nine inches apart (the other plot was not
planted soon enough for the greater distance) as he had on any of
his magnificent fields of wheat drilled with one bushel, or with two
bushels, or with three bushels, per acre, if he could show me any
such. He, his son, and I forthwith proceeded to test the matter,
with the following result, which astonished him even more than
the apparent boldness of my original assertion :—
Ears per
1s In. In. square yard.
Single grains planted (too late) et 12x12 gave... 229
” ” ” 9x9 ahi dO
Drilled October 11 ... 1 bush. per acre ue Pater Salleh
” ” pak, ” 2” 9 22 ee 283
“* Coles’ Field” drilled
end of October 2 ,, Bn Ss (ae GS
‘*Dears” drilled
NOV: Soc. -cecusay eis y 2 jo: se 269
4 | 1080
AVEFage:.s ctaseadius 270
or six less per yard than the number produced by single grains
nine inches apart each way. And this does not fully represent
the whole truth, for many of the stems of the thick sown wheat,
credited each with an ear, would fail to produce one at harvest,
while every one of the stems counted upon the single-grain Bie
would produce a fully developed one.
The Agricultural Returns now furnish us with pretty correct
‘information as to the acreage of wheat grown in the United
136
Kingdom, but until within the last few years we have been with-
out this knowledge. Nevertheless, statements have been for more
than a century past confidently made upon this point, and are
often now repeated by those who are considered authorities upon
the subject. The only foundation upon which such statements
rest are an assumed consumption of so many bushels per head, the
returns as to population, the deduction of the known imports, and
the division of the remainder by the number of acres grown in the
United Kingdom—which number of acres was, until the quite
new institution of Agricultural Returns itself, absolutely un-
known.
What value can be attached to such estimates, which are, in
fact, mere guesses? It is sufficient for my present purpose to take
40 bushels per acre as an excellent and thoroughly good crop.
This gives us 28 millions of grains as the produce of one million
ears per acre, or 28 grains only per acre! And this, for the
purposes of my present contention, is putting the case as strongly:
as possible against myself. For if the average crop be taken (as
supposed) at 30 bushels per acre, the number of grains produced
becomes reduced to 21 millions, or only 21 per ear; while if 1?
millions of ears per acre be claimed, the produce per ear can even
upon a crop of 40 bushels amount to only the same. Butif, while
maintaining the number of ears per acre, we double their contents,
this is doubling the crop, and would save annually the sending
out of the country a sum equal to the interest upon the National
Debt ! Beyond the increased contents of the ears produced in fully-
developed plants, the increased size of the grains alone adds 40 or
50 per cent. to the crop. Thus a bushel of fully-developed
pedigree wheat contains 460,000, a bushel of ordinary wheat
700,000 grains. Theretore, in two crops consisting ot precisely
the same number of grains, that from fully-developed plants would
be 70 bushels against 46 bushels, or 2 quarters against 6 quarts
per acre. So in barley the crop would be from this cause alone 7
quarters against 5 quarters per acre.
As this has been in fact a history of my labour in working
137
out this subject, I should be most ungrateful not to acknowledge
the generous support and encouragement given me from the first
by the Press, throughout my own and foreign countries, as also
by several kind friends, and by none more than the late Mr.
Merrifield, when such support and encouragement were especially
welcome ; but I shared, what is, I suppose, the fate of all pioneers,
meeting generally with obstinate prejudice and opposition for
many years, and that, too, from those who should have been the
very first to give me their countenance. However, I went
steadily on upon the magna est veritas principle, and I am happy
to say this has pretty well all died away, and I now reap a
sufficient reward in finding my system officially adopted by the
Governments of other countries and by agriculturists generally,
abroad as well as at home. I thank you for having so patiently
listened to a paper which has grown under my hands much longer,
and, I fear, more tedious, than I had at the outset intended it to
be.
The PresmpENT, Mr. G. D. Sawyer, on behalf of the
meeting, heartily thanked Major Hallett for his paper, one of
the best with which the Society had been favoured for a long
period.—A long and interesting conversation followed.
The PRESIDENT opened it with a few remarks expressive
of the results of his observation of wheat producing, which led
Major Hallett to state that Jethero Tull said that 90 per cent. of
_a square yard of very thickly sown wheat would be missing at
harvest, and to point out that the development on the stems
ceased as soon as the roots became impeded by crowding—as
soon as, in Darwinian phraseology, there was a struggle for
existence.
In reply to Dr. CorFe, the Major remarked that in all his
experiments he had employed no manure, his object being to
ascertain, so far as he could, the relative value of each grain, an
object whose attainment would be defeated if any substance was
used which might tend to the superiority of one growth over
another, by an unequal distribution of that substance.—Mr. C. F.
138
DENNET highly complimented Major Hallett upon the resolution,
courage, and perseverance, which he had exhibited in carrying out
his experiments.
Mr. W. SAUNDERS made a few remarks which induced the
Major to affirm that there was nothing in his system which should
prevent quantity going hand-in-hand with quality. Neither, said
he, in answer to a question from the President, was much time or
labour expended in following out his system as compared with
that usually spent.
Mr. J. HAMBLIN and Mr. J. E. HASELWOOD were among the
other interrogators, and, in reply to the latter, Major HALLETT
assured the meeting that but for selection the inherent power of
the wheat would belost—theavoidance of over-crowding in planting
was insufficient to retain it.
May 247TH.
MICROSCOPICAL MEETING.
The PresipENtT (Mr. G. D. Sawyer) expressed his pleasure
at hearing that one of the hon. secretaries (Mr. T. W. Wonfor)
was still improving in health, and was expected to be enabled to
get out of doors ina few days. He also stated that Mr. John
Mayall, jun., F.R.M.S., had found such difficulty in getting a lens
which was necessary for the elucidation of the subject of the paper
he had intended reading that night, on “The Microscope : its
Modern Development,” that he was compelled to withold that
paper till their next microscopical meeting, which would be held
on the fourth Thursday in June. Under these circumstances the
meeting would resolve itself into a conversazioné, and would
devote itself to the examination of objects under the microscope,
provided by Mr. J. E. Haselwood, Mr. R. Glaisyer, the librarian,
and himself.
139
JUNE 14TH.
MR. H. GOSS, F.L&., F.GS., &., ON “THE INSECT
FAUNA OF THE SECONDARY OR MESOZOIC
PERIOD, AND THE BRITISH AND FOREIGN FOR-
MATIONS IN WHICH THEIR REMAINS HAVE BEEN
DETECTED.”
I do not propose, in this paper, to make any preliminary
observations on the importance of an acquaintance with fossil
entomology, or on the valuable conclusions bearing upon the
geological conditions of the earth in former ages, which may be
arrived at from a study of insect remains. Such observations
as I had to offer on this portion of the subject have already been
made, in the introduction to my first* paper, an acquaintance with
which I shall presume on the part of all hearers and readers of
this, the second, paper of the series.
In the secondary rocks of this country, insects in a fossil
state are well represented. For our knowledge of them we
are indebted mainly to the Rev. P. B. Brodie, M.A., F.G.S.,
who, as I before stated in my first paper, is the author
of the only book on fossil insects which has as yet been
published in this country. In the determination of a large
number of the insects described and figured in this book, as well
as for some very important introductory observations therein, Mr.
Brodie had the great advantage of the assistance of Professor
Westwood, M.A., F.L.S. We are indebted to Mr. Brodie not only
for the very interesting and valuable work before mentioned, but
also for numerous important papers and notices on fossil insects
‘which have, from time to time, appeared in the “ Proceedings of
the Geological Society,” the “Quarterly Journal of the Geological
Society,” the “Annals and Magazine of Natural History,” the
“Proceedings of the Warwickshire Naturalists’ Field Club,” and
elsewhere. Amongst the names of other contributors to our know-
* See paper on ‘‘ The Insect Fauna of the Recent and Tertiary Periods
&c.,” read before the Society on the 8th March, 1877,
140
ledge of the fossil insects of the British formations of this period,
T must not omit to mention those of Professor Buckman, Messrs.
W. R. and H. Binfield, Professor Morris, F.G.S., Dr. Mantell,
F.R.S., Professor Edward Forbes, F.R.S., Mr. W. R. Brodie, Mr.
C. Wilcox, the Rev. O. Fisher, Captain Woodley, the Rev. Dr.
Buckland, F.R.S., the Rev. J. H. Austen, the Rev. E. F. Witts,
Mr. H. E. Strickland, F.G.S., Mr. Arthur Gardiner Butler, F.L.S.,
Mr. S. H. Scudder, Mr. E. T. Higgins, &c.
The British formations of this epoch, in which insect
remains have as yet been discovered in the greatest number,
are the upper series of the Upper Oolite (ie, the Purbecks)
and the Lias. The only other formations of the period
in which they appear to have been yet discovered in this
country are the Hastings sands, and some other subordinate
members of the Wealden system, the Kimmeridge clay of the
Upper Oolite, the Oxford clay of the Middle Oolite, the Forest
Marble, Great Oolite, and Stonesfield Slate of the Lower Oolite ;
and the Rheetic series between the Lias and the Trias. As yet no
remains have been recorded from the British upper cretaceous
system, the upper members of the lower cretaceous, or Neocomian
system, the Portland stone and sand, coral rag, Kelloway rock,
Cornbrash, Fuller’s earth and inferior Oolite, or from the Trias.
It is needless to say that one must not infer that because no insects
have as yet been discovered in these strata, none existed at the
different geological epochs of their deposition. Either they have
been overlooked, or their absence may be accounted for by the fact
that as these formations are of marine origin, the circumstances
attending their deposition were unfavourable to the preservation
of the insects of the period, and that, consequently, we have few or
no traces of them. Although, as I have observed in my first paper,
the presence of insects in marine formations may in some cases
be accounted for by supposing them to have been driven by the
wind into the sea when in its neighbourhood, or when attempting
to cross it, yet, as a rule, where their remains have been detected
in any abundance in decidedly marine formations, this may generally
141
be considered as satisfactory evidence that land was not far distant
at the time they became embedded ; and the abundance of other
terrestial remains, usually accompanying these fossils in marine
formations, leaves but little doubt upon the subject. One notable
exception to this rule is the continental formation known as the
**Solenhofen Slate” of Bavaria (belonging to the Upper Oolite),
in which the remains of insects are said to be mixed up, almost
exclusively, with the débris of the marine animals and plants.
Numerous lacustrine deposits have been discovered in the middle
of the marine formations, constituting the greater part of the
Jurassic system, which are supposed to indicate the situation of
ancient islands. If this supposition be correct, the presence of the
remains of terrestrial animals and plants in marine formations is
easily accounted for.
Before proceeding to review the various strata of the period
in which fossil insects have been detected, I think it may not
be out of place to call attention to the striking difference in the
state of preservation of the insects from English formations to
this period, with that of those from one of the most important of
the foreign formations (i.¢., the Solenhofen Slate). On the
subject of the deposition of the Solenhofen insects, I cannot do
better than quote Dr. Hagen,* who observes, “on comparing the
insects of Solenhofen and Eichstadt with those of England, there
appears, in the first place, a difference which may possibly admit
of interesting inferences ; the insects of the Bavarian strata are
almost universally preserved entire; wings, legs, head and
antenne are in their proper places; most of the Libellule have
their wings expanded. He who has noticed on the sandy shores
of the Baltic how depositions of insects are now taking place, will
admit that the insects of the Solenhofen strata were already dead
when deposited.” The insects would be, as now, driven by the
wind into the sea, thrown on the shore dead or dying, and there
*See a paper read by Dr. Hagen before the Geological Section of the
British Association at Manchester, September, 1861, and see the
Entomologists’ Annual for 1861.
142
gradually covered with sand by the rippling waves. That this
process took place gradually and slowly in the Solenhofen strata,
is evident also from another circumstance, for we frequently find
the cavities of insects, the head, thorax, and body filled up with
regular crystals of calcareous spar. Hence the pressure of the
stratum overlying the insects must have been very slight, when
such delicate parts as the abdominal segments of a dragon fly
could oppose resistance for a sufficient length of time to admit of
the formation of crystals. Naturally there do occur, here and
there, in the Solenhofen strata, impressions of insects obtained in
a different way, which admit of the idea of a very heavy pressure
from the superincumbent strata; yet these specimens are scarce,
and form only a small proportion of the entire number.” Now
the state of preservation of fossil insects from English strata of
this period is, as Dr. Hagen goes on to observe, in strong
contrast to that of the Solenhofen insects. The fossil insects
from English strata are seldom entire, and their remains generally
consist of single wings, or parts of wings. The imperfect con-
dition of the English insects may be accounted for by supposing
that their deposition did not take place gradually and slowly, like
that of the Solenhofen insects, but only after they had for a long
time been tossed about by storms or other commotions, or
remained for years soddening in water. It may seem almost
incredible that such fragile things as the wings of insects should
have been preserved to us at all, except under extremely favour-
able circumstances ; but, according to Dr. Hagen, the wings of
insects are almost indestructible in water, and he states that he
“has kept the wings of dragon flys in water for years without
observing the slightest change in their texture.” On the
Continent of Europe, so far as present researches enable us to
judge, the secondary rocks do not, as a rule, appear to contain
fossil insects in any abundance ; but to this rule the Solenhofen
slate of Bavaria, and the Lias of Schambelen in the Swiss Alps,
form a striking exception. For our knowledge of the fossil
insects of the Solenhofen slate we are indebted chiefly to
a hl
—— =
143 5
Professor Germar,* Count Munster, and Dr. Hagen. To Dr.
Hagen we are especially indebted for making us acquainted with
the numerous gigantic Newroptera obtained from this formation.
To Professor Oswald Heer we owe our knowledge of the fossil
insects from the Lias of the Swiss Alps. This indefatigable
savant, in comparison with whose labours in this particular branch
of science those of other paleontologists sink into comparative
insignificance, has obtained from Schambelen about 2,000
specimens of insects, comprising 143 species. A most interesting
account of these insects and of the strata in which they have
been found is given by Dr. Heer in his Urwelt der Schweiz,” an
English translation of which by Mr. W. S. Dallas, F.L.S., edited
(with some additions) by Mr. J. Heywood, F.R.S., has recently
been- published. With the exception of one insect from the
Swiss carboniferous system and five from the Trias, these insects
from Schambelen are the most ancient in Switzerland, and
consequently possess great interest. Among the names of other
students of, and writers on, the fossil insects of the continental
formations of this period, must be mentioned Schmiedel, Schroter,
Esperm, Van der Linden, Van Buch, Von Meyer, Dr. Geinitz,
Marcel de Serres, Professor Bronn, Professor Pictet, Dr. Giebel,
and Herr Weyenberg. _I will now review, as briefly as possible,
in the descending order of geological succession, the various strata
of this period, in which insect remains have been detected, and the
several orders and species to which such remains have been
referred. The English strata and their remains will be treated of
before proceeding to those of the Continent.
BRITISH STRATA.
FLOWER CRETACEOUS OR NEOCOMIAN.
In the Lower Cretaceous or Neocomian system remains of
insects are extremely rare and confined, so far as present in-
vestigations enable us to judge, to the very lowest members of
Epa RE) © fT oe
* Ueber die Versteinerten: Insecten des Juraschiefers von: Solenhofen,
1837. f
144
the series. The first remains recorded from these strata were
discovered in the neighbourhood of Hastings by Messrs. W. R. and
H. Binfield. They consisted of a few minute elytra and fragments
of Neuropterous wings, and were obtained in the courses of iron-'
stone exposed near low water mark, at a place called the
“ Govers,” near St. Leonards-on-Sea. Above the course of the
ironstone, in a bed of dark coloured shale, many elytra were.
found, as well as traces of wings of Coleoptera, Neuroptera, and
Diptera. A few fragments of Coleopterous elytra have also been
discovered in the Wealden marlstone, from between Tonbridge and
Maidstone, and are mentioned by Dr. Mantell. In the proceedings
of the Geological Society for 1854, Professor Westwood alludes to
the discovery, by Professor Edward Forbes, of some traces of fossil
insects in the Hastings series of the Isle of Wight, and of a few
doubtful specimens, by Mr. W. R. Brodie, in the Wealden series
of Punfield Bay Swanage.
Upper OOLITE.
PURBECK BEps.
In Mr. P. B. Brodie’s valuable and interesting work on
‘The Fossil Insects of the Secondary Rocks of England,” these
beds are referred to the Wealden formation; but modern
geologists are of opinion that in consequence of the organic
remains discovered therein, the Purbeck series has a close affinity
to the Oolitic group. It may, therefore, be considered the newest
or uppermost member of that formation, instead of the oldest or
lowest member of the Wealden formation. The “ Purbeck beds”
are of freshwater origin, and are so named from the Peninsula
of Purbeck in Dorsetshire, in the cliffs of which they were first
studied. These beds are of but limited geographical extent in
Europe, but they are of great geological importance, on account of
the succession of * three distinct sets of fossil remains which
they contain, these again differing from the fossils of the
Dorsetshire Purbecks, by Professor Edward Forbes, British Association,
Edinburgh, 1850.
145
overlying Hastings sands and Weald clay. This series is best
seen at Durdlestone Bay near Swanage, at Lulworth Cove, and
the neighbouring bays between Weymouth and Swanage. The
Purbeck beds were divided by Professor Edward Forbes into the
upper, middle, and lower. It is from the middle and lower
divisions that the greatest quantity of insect remains have been’
‘recorded, These remains have been discovered principally by the
Rev. O. Fisher, Mr. W. R. Brodie, Captain Woodley, Mr. C.
Wilcox, and especially by the Rev. P. B. Brodie, of whom
Professor Westwood observes that, “from his attention having
been especially directed to this branch of the subject, he has been
highly successful in detecting minute fragments of insect remains
in small slabs of stone, which would to a less educated eye have
been passed over as destitute of any traces of ancient animal
life.” *In the “Quarterly Journal of the Geological Society ” for
1846, Mr. Brodie recorded the discovery of a few imperfect’
remains of insects in Purbeck strata, in the neighbourhood of
Swindon, Wilts. A large collection of insect remains, consisting
of one hundred and eighteent small slabs, containing many
specimens, was made by Mr. Brodie from the middle Purbeck of
Durdlestone Bay, Dorsetshire. These remains included elytra,
bodies, and wings referable to Coleoptera, Orthoptera, and Diptera,
and portions of wings of Newropterous insects of doubtful family.
A number of specimens of fossil insects were obtained by the
Rev. O. Fisher from the Ridgway Quarries near Dorchester,
belonging to the lower Purbecks. A large collection of fossil
insects was also obtained by Mr. Wilcox from the Purbeck beds
near Swanage; the collection consisted of some 60 slabs, each
containing a number of remains. Professor Westwood says these
remains included a very extensive series of elytra of Coleoptera,
including Buprestide and Harpalde, also fragments of the wings
of Libellulide, Blattide, Tipulide, &c. A small collection of
* Quarterly Journal of the Geological Society,” 1846, vol. IIL,.
pp. 53-54.
+ ‘Proceedings of the Geological Society,” 1854.
146
insect remains, twenty-two in number, was made by Mr. W. R.
Brodie, of Swanage. Professor Westwood states that these
remains included Elateridw, Helopide, Curculionide, and the wing
of a gigantic Ant, which, in its perfect state, must have measured
at least two inches across the expanded wings ; it is most nearly
allied to some of the exotic forms of which Myrmica is the type
in temperate regions. ‘The discovery of such an insect,” says
Professor Westwood, “is of the highest importance in respect to
the question of the geographical range of the insects embedded in
the lower Purbeck series.” The largest collection of insect
remains from the lower Purbecks was made by the Rev.
P. B. Brodie. “It consisted,” says Professor Westwood, “of
three hundred and fifty small slabs of stone of various sizes.
Upon many of them only a single fragment of an insect occurs ;
but upon a considerable number, the remains are very numerous,
the fragments being crowded together, and often lying one upon
another.” These remains, of course, included quantities of elytra
of Coleoptera, and also wings and wing covers of Newroptera and
Orthoptera, and two wings supposed to be Lepidopterous. The very
fragmentary nature of these insect remains rendered the identifica-
tion of the majority of them a matter of impossibility, but from
Professor Westwood’s paper in the proceedings of the Geological
Society for 1854, and the list of species in Dr. Giebel’s* “Fauna
der Vorwelt” some eighty-six species seem to have been
«determined, viz. :—
Coleoptera... ae ae 3 45 species.
Orthoptera re ie a 1D er
Neuroptera si we! Res 10_=C*=»“;
Hemiptera sae “ee ses 1 A
Diptera... ome ae ae ae
Hymenoptera... by: se Dil sy,
(supposed) Lepidoptera 2 »
‘The two supposed species of Lepidoptera were respectively named,
by Professor Westwood, Cylloniwm Boisdwvatianum, and C.
* Fauna der Vorwelt, pp. 187, 393, and 426.
147
Hewitsonianum. Both species are included by Dr. Giebel in his.
list. On the subject of these supposed remains of Lepidoptera
‘Mr. Scudder observes : “I have not been able to find, even with
Mr. Brodie’s help, the first specimens referred to; but an
examination of the original of the latter proved that while it is
unquestionably an insect it cannot be referred to the Lepidoptera.
As the figure of the first species clearly resembles, in this
particular, the one I have seen, I am forced to the conclusion
that neither of these wings are Lepidopterous. Plainly, the only
reason why a new generic name was appended to these forms, was.
that their remains were too fragmentary to afford the slightest
guess as to what modern genus they might be referred.” All
geologists are, of course, aware that the formation known as the
“ Purbeck Beds” occurs not only in the peninsula of Purbeck in
Dorsetshire, but also in some parts of Wiltshire and Buckingham-
_ shire, and elsewhere. This formation in the vale of Wardour in
Wiltshire, consists, according to Mr. P. B. Brodie, of a series of
clays, white limestones, grits, sandstones, coarse blue limestones,
and fine white slaty limestones. In this last-mentioned limestone,
insect remains have been detected in such abundande, that it has
“been called, both here and in Dorsetshire, the “Insect Limestone.”
‘It is described as being of a somewhat coarse texture, often white
at the edges, and of a blue colour towards the centre, where it
becomes fine grained and harder, and then joins into a thin white
‘slaty limestone, very fine grained and having a laminated structure.
It was chiefly from these strata in the vale of Wardour in Wilts,.
and the vale of Aylesbury in Bucks, that the large collections of
fossil insects were obtained which furnished materials for Brodie’s
“History of the Fossil Insects in the Secondary Rocks of
England,” before referred to. Out of some two hundred and
forty specimens, or parts of specimens, from these formations,.
‘seventy-four * were figured by Professor Westwood in the work
last mentioned. The very fragmentary and imperfect condition
* Introductory observations to Brodie’s ‘‘ Fossil Insects,” by Professor
J. O. Westwood, M.A., F:L.S., &e.
148
-of a great quantity of the insect remains discovered in these
formations rendered it impossible, even for so skilled an entomolo-
gist as Professor Westwood, to determine, in many cases, the
species, or even the orders, to which they belonged. A great
many have, however, been identified, and, according to the lists
given by Mr. Brodie in his “ History of the Fossil Insects in the
Secondary Rocks,” and by Dr. Gliebel in his “ Fauna der Vorwelt”
(Insekten and Spinnen), the following orders are represented by
some sixty-four species, distributed amongst them in the
following proportions, viz. :—
Coleoptera... Ee oe xe 18 species.
Orthoptera ace 38 She 10°
Neuroptera ae Lae ee Oi or
Hemiptera ae 4 re 14 ~=~«C«z
Diptera... oe ac Ed 1s 3
64
The Coleoptera appear to have been very well represented,
-and to have included Buprestide, Carabide, Curculionidae.
Chrysomelide, Elateride, Cantharide, Tenebrionide, and Halophoride,
Professor Westwood also detected amongst the insect remains
from this formation Cercopide, Cimicide, Tipulide, Simulium,
Blatta, Aphis, Acheta, Of the species which have been determined
a large proportion are said to be herbivorous, and as the remains
-of plants are stated to be abundant in this formation, we have
reason for believing that the flora of the period was luxuriant.
Some of the species detected are certainly not herbivorous, but
the number of species which are vegetable feeders implies some
_abundance of carnivorous or predaceous ones. On the subject of
the remains from the Dorsetshire Purbecks, Professor Westwood
observes, “‘ With the exception of the dragon flies, of which there
_are as many as 34 fragments of single wings (from, which however,
it is impossible to affirm either a moderate or tropical climate)
and of a large Ant wing, it is worthy of remark that the whole of
these remains, not fewer than 700 or 800, are those of minute
insects, not more than a fourth or a third of an inch in length.”
Further on inthe same paper hesays, “If we take intoconsideration
149
the small and even minute size of the great majority of the insects,
and, indeed, of the whole of the Coleoptera which have been passed
under review, the idea that we have before us the wreck of an
insect fauna of a temperate region is at once raised ; for, although
it would be rash to assert that a mass of the existing tropical
insects might not be accumulated in which a large quantity of
minute beetles and flies would not be present, yet I cannot conceive
any process either arising from currents of water or chemical disso-
lution of insect matter, which would carry off or destroy the many
gigantic forms of insect life always occurring in the tropics. The
fossils before us show abundant evidence of the presence of numbers
of lignivorous species such as Elateride and Buprestide ; but we
nowhere find amongst them traces of the great Lamellicorn and
Longicorn Beetles. Herbivorous insects also occur in considerable
numbers, but we do not meet with the gigantic Grasshoppers and
Locusts of tropical climates.” Professor Westwood goes on to
observe that the question of by what means such masses of insect
remains could have been brought together, as are found in the
slabs of stone from this formation, is one for Geologists to decide.
Entomologists, however, are aware that sudden inundations, or
the rapid rising of rivers, are sure to bring with them vast numbers
of insects, which are carried away by currents in vast numbers, and
congregated together in masses. It appears, according to Professor
Westwood, that with the exception of the gigantic winged ants
and the wings of monster dragon-flies, that there is a very general
conformity between the insects from the Dorsetshire Purbecks and
those of Wilts and Bucks. There would, however, appear to have
been a great difference in the manner of disposition of the strata
-of the two districts, as evidenced by the remarkable contrast
presented by the state of preservation of the insect fauna of the
Wiltshire Purbecks, and that of the insect fauna of the Dorset
Purbecks. In the former, numbers of specimens are in a fair state
of preservation, while in the Jatter, the remains consist of almost
always of fragments of wings, elytra, or bodies. It appears from
_a careful examination of the fossil insect remains from these for-
150
mations, that the insects of this age are, with a few exceptions,
closely allied to forms now in existence, a result, upon the whole,
as Mr. Brodie observes, “ quite the reverse of that which we are
led to infer from the rest of the fossils of the Secondary Rocks.
Thus while we have in the Purbecks many strange and extinct
races among other divisions of the animal kingdom, the insects
which accompany them are more nearly related to existing genera,
and present, upon the whole, a decidedly European character ;
indeed, the greater part must have been of the inhabitants of a
temperate climate, although some few were adapted to a much
higher temperature. This is the more remarkable because the
colossal saurians, palms, and tree ferns in these strata,
evidently belonged to a hot country. It is true that these
are chiefly confined to the upper division ofthis deposit”
(i.e., the Wealden proper); “but many of the fossils in the
lower or Purbeck beds are equally conclusive in the matter,
as the remains of the Megalosaurus, crocodiles, turtles, and
cycas clearly prove.”
KIMMERIDGE CLAY.
Below the well known Portland stone, so extensively
quarried for building purposes, is a dense bed of sand called the
Portland sand, immediately below which is the Kimmeridge clay.
The Kimmeridge clay is said by Lyell to consist, in great part, of
a bituminous shale, sometimes forming an impure coal several
hundred feet in thickness. At Ringstead, in Dorsetshire,
according to Mr. Brodie, this clay is traversed by a bed of sandy
laminated stone about two feet thick, and this is succeeded by
thick strata of dark coloured shale and clay, containing large
blocks of Septaria, in one of which Mr. Brodie discovered a
striated elytron of a small beetle. With this exception, I find no
record of the discovery of any insect remains from this part of
the Upper Oolite system.
MIDDLE OOLITE.
OxForD CLAY.
The Oxford clay consists principally of laminated beds of
151
blue clay, including in some places, subordinate courses of lime-
stuuc culled Kelloway rock. The only trace of insect remains
from this division of the Oolitic group is the supposed larva of
an insect, found by Mr. J. C. Pearce, near Christian Malford,
which Professor Westwood thought might possibly be referred to
a Libellula. As the Oxford clay is supposed to have been
deposited in a deep sea, remains of insects are hardly to be
expected in it.
LOWER OOLITE.
FOREsT MARBLE.
The forest marble is found immediately below the cornbrash,
and belongs to the upper series of the Lower Oolite. It is said
to be composed chiefly of thin fissile and slaty oolitic limestones,
divided by clays, sometimes by sand and grit. The only record
of the discovery of fossil insects from these strata is made by Mr
P. B. Brodie, who states that in some of the quarries in this
series, at Farleigh, near Bath, he found a few elytra of small
beetles. They are undescribed.
GREAT OOLITE.
In the list of fossil insects at the end of Mr. P. B. Brodie’s
paper on “The Distribution and Correlation of Fossil Insects,”
&c., I find a record of elytra of Coleoptera from the Great Oolite
formation, in the eastern moorlands of Yorkshire. The discovery
of these remains is also alluded to by Dr. Mantell in the “ Medals
of Creation;” by Dr. Buckland in the “Bridgwater Treatise ;”
by Mr. R. C. Taylor, in the III. Vol. of “Loudon’s Magazine of
Natural History ;” and by Messrs. Young and Bird, in their
“Geology of Yorkshire.” In a note at page 379 of the
“ Geological Proceedings for 1854,” reference is made by Professor
Westwood to a fossil insect having been met with by Professor
Morris in the Great Oolite of Lincolnshire.
STONESFIELD SLATE.
The Stonesfield slate lies at the base of the Great Oolite, and
is found chiefly in Oxfordshire and in the Cotteswold Hills,
Gloucestershire, though there are, I believe, similar beds at Ketter-
\ 152
ing, Northamptonshire, and near Stamford, in Lincolnshire. Tt is
described by Lyell as “a slightly oolitic shelly limestone, forming
large lenticular masses, embeded in sand only six feet thick, but
very rich in organic remains.” Mr. Brodie says the (Stonesfield)
slate beds are largely developed in the north east of Gloucester-
shire, and have been traced from Wooton-under-edge to the neigh-
bourhood of Burford. From thence they may be followed in a
more northerly direction, and are well exhibited near Bourton on
the Water, Upper Slaughter, Stow-on-the-Wold ; and on the west
at Eyeford and Maunton to their farthest limit in this direction
at Sevenhampton, six miles east of Cheltenham. The remains of
insects in this formation appear to be almost confined to a few
families of Coleopiera, and they are generally in a very fragmentary
condition, and consequently most difficult to determine. In the
second volume of the “Geological Proceedings,” Dr. Buckland has
described the wing of a large Neuropterous insect from the
Stonesfield slate, which has been named Hemerobioides Giganteus.
In the list at the end of Mr. Brodie’s paper on “The Correlation and
Distribution of Fossil Insects ” before mentioned, another specimen
of a Newropterous insect is mentioned (Libellula Westwoodii) besides
two large wings, referred to the Libellule, which were found at
Eyeford on the Cotswolds by Mr. Brodie. The remains of
Coleoptera from this slate have been referred by Brodie and West-
wood to some seven families, including Buprestide, Curculionidae,
Coccinellida, Prioniide, Blapside, and Pimeliide. By far the most
interesting insect fossil from the Stonefield state is the wing of a
large insect, which Mr. Butler, Professor Westwood, and Mr.
Bates believe to be Lepidopterous. In the proceedings of the
Entomological Society of London, for 1872, we find the
following notice of its exhibition at a meeting of that Society.
“Mr. Butler exhibited a remarkably perfect impression of the
wing of a fossil buttertly in the Stonesfield slate. It appeared to
be most nearly allied to the now existing South American genus
Caligo. Mr. Butler subsequently named the species to which he
referred this fossil Paleontina Oolitica, and described and figured
153
it in his “Lepidoptera Exotica.” I quote his description—
_ “Though a British insect, this species belongs to a group so
completely tropical that I do not hesitate to describe and figure
it in the present work ; its nearest allies are the genera Calligo,
Dasyophthalma, and Brassolis, all three essentially tropical
American genera, LP. Oolitica is especially interesting as being the
oldest fossil butterfly yet discovered ; the most ancient previously
known to science having been found in the Cretaceous series
(white sandstone of Aix la Chapelle)” (this is an error of Mr.
Butler’s ; the insect was found at Aix in Provence, in strata be-
longing to the Upper Eocene period) “whilst the bulk of the
known species are from the lower Miocene beds of Croatia ; it is
also interesting as belonging to the highest family of butterflies
and to a sub-family intermediate in character between two others,
: ) viz., the Satyride and Nymphali, whilst the more recently dis-
covered fossils are referable, with one exception, to the two latter
groups. The nervures appear to have been impregnated with
iron, which will partly account for their well defined condition.”
Mr. Scudder, who states that he carefully examined the
original fossil, considers Mr. Butler to be in error in referring
this wing to a Lepidopterous insect, and is of opinion that it should
be considered Homopterous rather than Lepidopterous, and allied to
Cicada. Mr. Scudder’s opinion is presumedly formed from a
careful study of the neuration of the wing and a comparison of
the neurations of Lepidoptera with those of Cicada. Mr.
Scudder’s views on the subject are given at length in his “ Fossil
Butterflies,” pages 90-95, but time will not permit me to quote
them. Mr. Butler has himself directed my attention to the
neuration of this wing, and also to that of some of the Cicade in
the British Museum Collection, and, from a comparison of the
wing with these insects, I am inclined to agree with Mr. Butler,
who in his view is supported by Professor Westwood and Mr.
Bates. It will be observed that Mr. Butler refers this wing to a
butterfly of a tropical species, which is quite consistent with the
‘general character of the large Neuropterous wings and Bupresti-
154
dean elytra of the Stonefield slate, which is stated by Professor
Westwood to be “ decidedly tropical.” Probably Mr. Scudder is
somewhat prejudiced by the opinion which has, according to M.
Preudhomme de Borre, always prevailed amongst Continental
Naturalists, viz., that the order Lepidoptera was the last created,
and that none existed prior to the Tertiary epoch. I believe that
the same opinion is also held by such English entomologists as
have given the subject their attention. Sir John Lubbock, F.R.S.,
in his “ Origin and Metamorphoses of Insects,” observes, ‘ Well
characterized Orthoptera and Neuwroptera occur as early as the
Devonian strata ; Coleoptera in the coal measures ; Hymenoptera,
Hemiptera and Diptera in the Jurassic; Lepidoptera, on the
contrary, not until the Tertiary.” M. Oustalet appears to form an
exception to Continental Naturalists, in his view of the probable
date of the apparition of the Lepidoptera on the Geological
Horizon. At page 26 of his “Recherches sur les insectes fossiles,”
he gives the Upper Oolite (Epoque de Purbeck) as the period
during which this order first appeared. Since the date of the
discovery of P. Oolitica, another fossil butterfly is stated to have
been discovered in the Stonesfield slate of Oxford. This second
specimen is said to be in the possession of Mr. Parker, of Oxford,
but has not yet, I believe, been figured or described.
Lias AND RHETICS.
The Lias, including therein the Rheetic series, is in this
country the lowest formation of the secondary rocks in which
fossil insects have been detected. It is of great extent in this
country and generally rich in remains. It consists of Argillaceous
limestones, shell, marls, and clays, and is in England divided into
three formations, viz., the Upper, Middle, and Lower, and is for
the most part of marine origin. Sir Charles Lyell states that
some members of the series, especially in the lowest part of it,
have an estuarine character, and must have been formed within
the influence of rivers. In some sections of this formation insect
remains have been detected in such abundance that the beds
containing them have, as in the Purbeck strata, been called the
155
“Insect Limestones.” The remains of insects have been found
chiefly in the lower division of this formation in Gloucestershire,
Worcestershire, Warwickshire, Somersetshire, and on the borders
of Monmouthshire, and a few in Yorkshire. They are generally
in a much more fragmentary condition than those from the
Purbecks, and, although abundant, are less common than in the
latter formation. Mr. Brodie states that he first discovered these
interesting fossils in the immediate neighbourhood of Gloucester.
He also states that some of the beds of limestone in the lowest
division of the Lias, in the vale of Gloucester, abound in insects,
and that beautiful specimens of insect remains, chiefly elytra and
wings, have been found in the Upper Lias at Dumbleton and
Alderton. In this locality (Dumbleton), which is about twelve
miles north-east of Cheltenham, Mr. Brodie obtained from the
Upper Lias shales one nearly perfect Newropterous insect, of which
Professor Westwood says: “It possesses an arrangement of the:
wing veins differing from that of any English species, and also
from any foreign species known to me, but it comes nearest to the
small Libellule forming the genus Diplaz. In the Upper and
Middle portions of the Lower Lias, which are extensively
_ developed in the neighbourhood of Gloucester and Cheltenham,
traces of insects are said to be exceedingly scarce, but at Wainlode
Cliff, on the banks of the Severn, near Gloucester, the insect
_ limestone has produced, according to Mr. Brodie, remains of
_ several genera of Coleoptera, which are not very rare ;, and a few
wings not unlike those of the genus Tipula. With these were
found abdomens of some insects, and larve, apparently of the
-gnat tribe. In insect limestone, to the south-west of Combe Hill,
not far from the last mentioned locality, Mr. Brodie obtained a
great number and variety of insect remains, consisting chiefly of
the elytra of Coleoptera, and a few imperfect, but large, wings of
Tibellule.
At Apperley, near Wainlode Cliff, the remains of insects
have been found in plenty, and many small slabs, three or four
inches square, exhibiting several elytra and. wings, and. a few
156
small beetles. From the insect limestone near the village of
Hasfield, Gloucester, various elytra of Coleoptera and a few
beetles have been obtained. The same formation in the neigh-
bourhood of Forthampton, near Tewkesbury, has also furnished
insect remains similar to those found in the localities before
mentioned. At Strensham, two miles from Dafford Station, and
nine from Evesham, from a bed of insect limestone at the bottom
of a large quarry, numerous remains of insects have been obtained.
Amongst them was found part of the abdomen of a gigantic
species of Libellula, which Mr. Brodie named Libellula Hopei. In
the neighbourhood of Evesham the insect limsetone has produced
numerous remains of insects, the wings and elytra of many of
which are said to be beautifully preserved. In the lower division
of the Lias, in this neighbourhood, Mr. H. E. Strickland
discovered small elytra of Coleoptera and portions of the wings of
Libellula. ‘In the County of Warwickshire,” says Mr. Brodie,
“there is a large extension of the lower Lias in the neighbourhood
of Bidford and Binton. The quarries thereabouts afford some of
the finest sections in this part of the series. The insects are not
confined to one or two comparatively thin seams of limestone, but
occur in several distinct beds, the whole being developed to a
much greater extent than in any part of Gloucestershire hitherto
examined.” From one quarry near Bidford Mr. Brodie obtained
a small species of the family Gryllid~, which he named Gryllus
Bucklandi, in honour of Professor Buckland. In some of the
strata in this neighbourhood (Bidford) the wings of Libellula were
obtained not uncommonly, particularly at a place called the Nook,
where a beautiful specimen was found, which has been described
and figured by Mr. Strickland. Mr. E. T. Higgins obtained from
the Lower Lias or the Rheetics, in the southern parts of Gloucester-
shire and the adjoining county of Somersetshire, in the neighbour-
hood of Bristol, numerous remains of insects. From Aust, near
Bristol, and from Sudbury on the Monmouthshire side of the
Severn, about three miles from Chepstow, the insect limestone and
the landscape stone have afforded a quantity of remains. “In some
i i ae ie
157
slabs the insects were found embedded together in masses, In
one slab Mr. Higgins is said to have detected as many as thirty
small beetles. From the frequency of such delicate creatures as
insects in the “Landscape Stone,” and in another band of
limestone only a few feet higher, some of which are said to be
beautifully preserved, and could not have been long subject to the
action of the waves, Mr. Brodie supposes that this part of the
Lias may have been formed in an estuary, which received the
waters of some neighbouring coasts, and which brought down
the remains of insects and plants. The Coleoptera appear to have
been abundant in the Lias, for out of some three hundred
specimens of insects, or parts of specimens, from this formation,
examined by Professor Westwood, more than one-third were:
referable to this order, and comprised the families Buprestidae,
Elateride, Curculionide, Chrysomelide, Carabide, Telephoride, &c.
Most of the species appear to have been very minute, “never:
equalling,” observes Mr. Westwood, “in size those from the
Stonefield Slate.”
d The other orders represented in this formation are the
Orthogtera, the Neuroptera, the Hemiptera, and the Diptera ;
the minute Diptera and Trichoptera of the Purbecks are said by
Westwood to be absent. The remains of Orthoptera include
Gryllide and Blattide, the Hemiptera include Cicada and Cimez, and
the Neuroptera, Libellula, Agrion, Orthophlebia, Hemerobius, Eshna,
Chauliodes, and Ephemera. Among these various families we have
terrestial and fluviatile genera, which include omniverous,,
_ herbaceous, and predaceous species. A great number of the
families and genera found in the Lias are common both to it and
the Purbecks. The following Lias insects, however, have not, I
believe, been found in the Purbecks, viz, Gyrinus, Laccophilus,.
Berosus, Melolontha, Telephoride,; Gryllide, Agrion, Hemerobius,
Ephemera and Asilus. Although, as a rule, the remains of insects.
from this formation are very imperfect’ and fragmentary, the
detached wings of many Neuropterous insects are said to be
preserved in the greatest perfection, and have the nervures of the:
158
wings beautifully defined, and retaining the original colour, or
rather the spots. The size of the insects, judging from the
remains, appears to have been usually small and indicative, if
taken alone, of a temperate climate. Mr. Brodie observes that,.
“with some rare exceptions (viz.), the gigantic Libellulide,
Termitide, and larger Chanliodes, these insects seem to have
belonged to a temperate climate, and thus they offer, as in
the Purbeck series, a remarkable distinction in the contemporary
forms of animal life, which, both in the Purbecks and Lias,
are chiefly referable to extinct genera, and were most probably
natives of warm latitudes. In one respect, however, they differ
from the Purbeck insects, since they are less closely allied to
European forms, and more nearly to those which now inhabit
North America.” It may be observed that all the numerous
remains of insects from this formation have, with the exception
of a few specimens from the upper division, been obtained from
the lowest division of the Lias, or from the Rheetic series, between
the Lias and the Trias. These remains are very numerous, but
the majority of them were in such a fragmentary condition that
it has been impossible, even for those who have devoted their
especial attention to the subject, to make out the species to which
they belonged. About fifty-six species, however, are stated to
have been determined, which are distributed amongst five orders,
as follows, viz. :—
Coleoptera... Pe es es 29 species.
Meuroptera re Sos ae IZ Aces
Orthoptera ope oes es ae
Hemiptera... ee ae an OY My
(supposed) Diptera... > Feetane
No traces of eee or i ee have been met with,,
and the remains which have been referred to the Diptera are very
doubtful. In concluding this part of my paper, I would observe
that whilst fully appreciating the value of the learning, the
industry, and the research, of the Geologist, which have been the
means of bringing to light these interesting relics of a former
world, the value of the special knowledge and experience of the
Entomologist, must not be forgotten ; and it should be borne in
.
;
159
mind that the learning and skill of Professor Westwood, an
Entomologist of world-wide reputation, was, undoubtedly,
of the greatest assistance in the determination of the species, or:
of the orders, to which they belonged, of many of the specimens
obtained from the Purbecks and the Lias.
FOREIGN STRATA.
: EUROPE.
The Rocks of this period on the Continent, with theexception
of the Solenhofen slate, and the Lias of the Swiss Alps, have not,
as before stated, as yet produced any quantity of insect remains.
CRETACEOUS.
The Cretaceous system has as yet produced but very few fossil
insects. Professor Pictet alludes to some indistinct impression of
insects in the Schists of Glaris. Dr. Geinitz has called attention
to the remains of perforated wood in the upper and lower
greensand, of Saxony, which appear to him to be evidence of the
existence of Longicorn beetles during this epoch. He has referred
the insects, which he believed to have made these perforations, to
the Cerambycide. Professor Pictet further says that M.
Desmoulins found certain elytra of Coleoptera in the chalk marlof
the mountain of St Catherine, near Rouen.
Upper OOLITE.
SOLENHOFEN SLATE.
The most celebrated formation of this period from which
fossil insects have been obtained in any number, is that of the
well known lithographic stone of Solenhofen in Bavaria (before
referred to), belonging to one of the upper divisions of the Oolite.
Of this formation Sir Charles Lyell observes that, « although the
‘number of éestacea in this slate is small and the plants few, and
those all marine, Count Munster had determined no less than 237
species of fossils, including 26 species of insects. These insects,
among which is a Libellula, or dragon fly, must have been blown
out to sea, probably from the same land to which the flying
lizards and other contemporaneous reptiles resorted.” The
oldest account of fossil insects from this formation is found in
160
Schroter’s Lexicon, published in 1779. In a subsequent work
Schroter figured a Lepidopterous insect, which Germar named
Sphinz Schretert. In 1783 Esper refers to the discovery of a
species of Gryllus in the Solenhofen Slate. In 1826 Van der
Linden described a Neuropterous insect from Solenhofen as
Cschna Antiqua. Reference to fossil insects from this locality is
also made by M. Marcel de Serres in 1829, and Bronn, in 1835,
enumerates two genera of Coleoptera, four of Neuroptera, two of
Hymenoptera, and one of Lepidoptera, from the same locality. In
1839 Professor Germar published in the ‘“‘ Nova Acta” of the
Leopoddine Academy, descriptions of 25 species from Solenhofen,
including Coleoptera, Orthoptera, Hymenoptera, Hemiptera, Lepidop-
tera, and Diptera. Several species from this locality were also
described (in 1814) by Count Munster, including a Lepidopterous
insect, which he referred to the Tineina. In his “ Fauna der
Vorwelt” (Insekten and Spinnen), published in 1856, Dr.
Giebel enumerates 26 species from this locality, including :—
Coleoptera... as “ts sa 2 species.
Orthoptera mn Wes we ae
Neuroptera ah ses = = 12a
Hymenoptera... os <> i ane
Hemiptera... ‘ ay aE 4 Toms
Lepidoptera ey; Se ee oe ds
Diptera sag Re eas DA ee
26
Dr. Hagen, who has paid especial attention to the Solenhofen
Neuroptera, describes 24 species in the tenth volume of Meyer’s
Palontographica, published in 1862, and also gives a list of 37
species in the Royal collection in the Academy of Munich. In
Vol. XV. of Meyer’s Paleontographica, published in 1865, Dr.
Hagen described and figured eight further species of Neuwroptera
from the Solenhofen formation. Of the 37 species of Newroptera
in the Munich collection, 27 are said by Dr. Hagan to be dragon
flies, some of which belong to extinct genera, but others to such
_as now live in America and Australia. Dr. Hagan states that
many of the fossil insects from this formation, preserved in the
Munich collection, and the collection of Dr. Crantz, of Bonn, are
ae
161
in-a splendid state of preservation. One-third of the entire
collection are Libellulw, another third consists of Orthoptera (in-
cluding some gigantic Locustw) and Hemiptera (especially givantic
species of Belostoma, Pygolumpas, and Nvpa), and -the remaining
third consists of Coleoptera, Hymenoptera, and Diptera. Out of
450 specimens of fossil insects in the Munich collection, 150 are
Neuroptera, and out of these 136 belong to the Odonata. Dr.
Hagen says that of those not Odonata, six only (comprising four
- species) belong to the Newropfera as restricted by Erichson, viz.,
one species of Corydalis, one Chrysopa,a large Apochrysa,and a
beautiful Nymphes. The last two genera, which do not seem
very remote from Chrysopa, are now found only in the Southern
Hemisphere. Nymples is almost exclusively an Australian genus.
The Qdonata, continues Dr. Hagen, comprise the following, viz.:—
Libellulina ... ss a oe 4 species.
A’schnina ... oP ae es Dee
Gomphina ... be rer pi! TERE
Calopterygina Hes ae Se |i aie
Agrioniana ... a eS
The Gomphide are said to belong principally to species which
come near the genera Petalia, Phenes, and Petalura, of which a
few species occur at the present day in Chili and Australia.. The
fossil Gomphina of Solenhofen are generally very large, and some
are truly gigantic, 4 inches in length, with an expanse of wings of
74 inches. Of the Calopterygina, the group Heterophelbia contains
some of the most gigantic Newroptera, two of them belong to the
largest known Odonata, having an expanse of wing of 73-8 inches,
and bodies 34-4 inches in length. According to M. Oustalet, a
number of specimens from Solenhofen, preserved in the Museum
Tyler at Haarlem, have recently been described by Herr Weyen-
bergh, jun., who has distinguished 74 species, as follows :—
Of the Coleoptera 2 ... 26 species.
», Diptera... a hie er tee
», Orthoptera ve a TORS hs
s, Neuroptera ae A. Ao ge
» Lepidoptera ee B. | ae
74
From a study of the fossil insects from this formation and a com-
162
parison of them with those of England, Dr. Hagen has drawn
two conclusions, viz. :—Firstly, That the two fauna are extremely
closely allied, and that possibly some species occur in both
formations. Secondly, That the fauna of the English and
Bavarian strata are not only quite distinct from the existing
fauna, but also from those of Aix, of the Rhenish peat deposit,
Brown Coal of the Rhine, of CEningen and Radoboj, and from
that of Amber, differing not only in species but in genera.
LIAS.
From the lower marls of the Lias, at Schambelen, in the
Canton of Aagau, in Switzerland, about 2,000 specimens of fossil
insects, comprising 143 species, have been obtained by Professor
Heer. With the exception of one insect from the CarboniferouS
system, and five from the Trias, these are the most ancient
insects of Switzerland. Dr. Heer considers that Schambelen is
the only locality on the Continent in which so large a number of
primeval insects have been preserved to our time. The 143
species are distributed as follows :—
Coleoptera... a. - ... 116 species.
Orthoptera re Vi mah
Neuroptera Bes Lh Bi 95
Hemiptera iF ay <- 1
Hymenoptera... 3 as reas
143
No Lepidoptera or Diptera have as yet been discovered, and
the Hymenoptera are represented only by a single small wing.
The abundance of the Coleoptera may be due to the circumstance
of their hard elytra having been better adapted for preservation,
than the soft wings of most other insects ; but as the delicate
membranous wings of the Termites have been preserved, there
can be but little doubt that if the Diptera and Lepidoptera had
existed at this period, some traces of them would have been dis-
covered. In this deposit the order Coleoptera is represented by
more than four times as many species as all the other orders put
together. The species are referable to some sixteen families,
Of
oe
a
.
EE
Wn Be a *
163
including Carabide, Cryptophagide, Byrrhide, Hytrophilide,
Buprestide, Elateride, Rhyncophora, &c. Certain orders are
entirely unrepresented, including the Longicornes, Coccinelle
(Ladybirds), the Xylophagi, Melasomata and Brachelytra. The
Lamellicornes and Chrysomelinw are poorly represented. The
presence, in large numbers, of those beetles whose larve are wood
feeders, and who in their perfect state are usually found on or
under the bark of trees, is evidence of the existence of woods or
forests during this period. The presence of several genera of
fungus beetles implies the existence of fungi, but as yet none have
been discovered. Of the Byrrhida four species have been found
in this formation; three of them, Byrrhidium, Arenatum, B.
Morio, and B. Troglodytes, are among the most abundant insects
at Schambelen, and as they are all moss feeders, Professor Heer
supposes that the ground and trunks of the trees were more or
less covered with mosses, although no remains of these plants
have been preserved. The leaf-eating beetles are represented by,
amongst other species, two of the family Chrysomelide. The
presence of a small Coprophagous beetle (Aphodiites protogeus ),
so closely resembling the Aphodii which now live on the excre-
ments of cattle, would lead to the presumption of the existence
at that time of mammalia in Switzerland. All the before-named
beetles are herbivarous species. Of the carnivorous beetles we
have 29 species, including Telephoride and Carabide. The water
beetles are represented by some 20 species. The Orthoptera com-
prise three species of cockroaches, three grasshoppers, and one
ear-wig; and the Neuwroptera are represented by 6 species of
Termites (or white ants) and one dragon fly (Libellulide ), Gischna
Hageni, which is larger than any living species. It may be safely
inferred from the richness of the insect fauna that the land must
have been of considerable extent. The Aquatic insects,
especially, afford evidence of this fact. Their numbers are so
great as to imply the presence of a river or large lake, the exist-
‘ence of which in a small island is improbable. Many of the
insects are of large size, but the majority are small, some even
164
smaller than their smallest living relatives of the present day, as
is also the case with the insects of the English Lias. When the
conditions of climate are favourable to a luxuriant growth of
plants and insects, the larger forms of the insect class are
developed, as in tropical America and Asia, but side by side with
these large species thousands of minute species are to be found.
It may be inferred from the general smallness of the Liassic
insects, both from Continental and English strata, that insect life
was not at this period so luxuriant, or capable of producing such
large forms as the tropical world of the present day. The
numerical proportions of the families furnish evidence of a warm
climate. The Buprestide are very numerously represented. Of
the Termites eight tropical forms have been discovered, and the
cockroaches appear, says Heer, more closely allied to those now
inhabiting the Torrid Zone than to those of Switzerland at the
present day.
The only other record which I have been able to find of the
occurrence of insects in this formation on the Continent, is
contained in the second volume of bBronn’s * “Lethea
Geognostica.” At page 210 of that work it is stated that Count
Miinster detected some wings of insects in the Lias of Bayreuth.
Trias, oR Upper NEW RED SANDSTONE.
The Trias or Triple Group is separable on the Continent
into three distinct formations, called the ‘“Keuper,” the
“Muschelkalk,” and the “Bunter Sandstein.” The “ Muschelkalk,”
or Calcaire Coquillier of French geologists, is wanting in this
country. As has before been stated, no traces of insects have as
yet been found in any part of this formation in the United
Kingdom, and until recently none had been recorded from strata
_of this period on the Continent. M. Oustalet, in the first part of
_his “ Recherches sur les insectes fossiles,” published in 1871, says
“« Les insectes fossiles thanquent jusqu’a présent dans le terrain
* Jahrb, 1835, s. 332 and Lethea Geognostica, vol. i., p. 210.
165
pénéen et dans le Trias.” Since this was written, however, five
specimens appear to have been discovered in this formation. In
Mr. Heywood’s edition of Heer’s “Urwelt der Schweiz”
(Primeval Switzerland), it is stated that “in the Keuper Marls of
the Rutihard, Professor Heer has vainly sought for insects, but in
the Black Shales of Vadutz he has found two species of beetles,
viz. :—Glaphyroptera Pterophylli and Curculionites prodronus.”
Furtker on in the same work it is stated, “‘ In the Bunter Sanstein
of Trelitz and Salzmunde three more have been found, Legnophora
Girardi, Chauliodites, Picteti, and Chauliodites Zinckeni.”
AMERICA.
TRIAS.
In the Geological Magazine for May, 1868, Mr. Scudder
mentions one Colcopterous insect from the Trias of the North
American Continent. ‘This is, at present, the only fossil insect
recorded from the secondary rocks of America. With the
“Bunter Sandstein” we arrive at the lowest stratum of the
secondary period, and I am consequently able to bring this paper
to a conciusion. Before this paper is published, lists’ of the
families, genera, and species of the insects detected in the
secondary formations will be added to it, by reference to which
the fullest information respecting the ‘Insect Fauna of the
Secondary Period ® will be obtained,
The PRESIDENT, Mr. G. D. Sawyer, said he was sure it
would be the pleasure of the Meeting, as it was his, to pass
a vote of thanks to their essayist for the very exhaustive
paper he had placed before them. He was hopeful Mr. Goss
would not misunderstand a remark he made on the reading
of the previous paper, so as to think he (the Chairman)
was impatient for him to draw deductions when he had not
yet examined the evidence sufficiently to arrive at his deductions.
166
He took it that those who were close observers would be
of the very greatest importance to them in the investiga-
tion of scientific matters, and that if they would give them the
results of their close investigations, the time would come when
by a comparison of those investigations and of the evidence thus
obtained, they would be able to draw deductions which might be
still more interesting than the facts which had been put before
them that night. He himself had been greatly interesting in
what they had heard. It had not been to him at alldry, but he
had been a little disappointed at the barrenness of the upper
cretaceous white chalk period which we enjoyed so much round
here, that we could not do anything to illustrate the subject
spoken about; but he did not despair that the remains of
insects would be found in that upper cretaceous period for
Great Britain. It would have struck them, as it did him,
that the investigation of the subject of insect remains threw
a light upon other things that they found in very low
periods, such as the lower lias; they could tell what must have
been the surrounding state of things as to the woods, mosses, and
fungi, and so on by finding that insects which must have existed
in those days were obtained, and that they remained even after
the objects on which they had fed had dissappeared entirely.
They could never tell in investigating scientific matters what
light would be thrown on what they were about ; there was very
great encouragement, therefore, to persevere in close observation,
and the record of observations carefully taken was of very great
importance; and that seemed to have been the forte of their
worthy essayist. On their behalf, therefore, he begged to tender
him their hearty thanks for the excellent paper which he had
read,
Mr. Farr asked the lecturer if he had ever examined any of
the deep chalk pits in this neighbourhood.
Mr. Goss observed that the chalk had not produced any-
thing, and he thought it had been pretty well worked. He
; 167
believed a geologist would say there was hardly any strata which
was better worked for fossils of one sort or another than chalk.
The CHAIRMAN thought Mr. Farr would remember that the
objects discovered in the chalk were marine.
Mr. Goss said it was quite exceptional to find insect remains
in marine formations at all. They might be blown into the sea,
or by a strong river washed out in currents into the sea, but as a
rule they did not expect to find them in marine deposits. One of
the most important things which they drew from the study of
insect remains was what the climate of a former period was, and
they might arrive at some knowledge on the subject by consider-
ing the proportion existing between the carnivorous beetles and
the vegetable beetles, because it was well known at the present
day there was a certain ratio existing between those two groups,
which varied according as they come nearer to the equator or
departed further from it. The nearer they came to the equator
the'vegetable beetle got much commoner, and the carnivorous
much more rare.