MATE) PARTOUT ETES RCIP PERLES Ag a aa sags VED, Sasa het ad. î aa Rae rade DENT die impari Deren Tr ia CARE degl id wtp a ae hess Besten A HP are 02051074 PS WI ! DE A OUEST PES Bear us DOTE CIONI FTT i PRE NTFITIO heeegine she POLE $, 040, Tete te” PEL ring bat bat feierte READ nto Apa giaro prac ytetenren ete ARCHE AIDANT ER LOTIR) EGLI vii arte i hat tt Sy Pye ta teense HANNAH S satana rato ERP mena ritirate anita se ES Teen ere CCR rin « Pye 4 385 len PRE mat CRIER fq th tata retire = DRE ; DEN Pe etere DRE i os 3 pa PENSO cana een Sha ata eta Etes MILITE RE HE FEU «spazza; sa va ste) DRE sarà ze E erin sg rara tates visiva mmie Sagre Sara anregt 5; ERBEN" LR RE hi E PO UN CRT AR NITRO REVUE SUISSE DE ZOOLOGIE REVUE SUISSE ZOOLOGIE ANNALES DE LA SOCIÉTÉ SUISSE DE ZOOLOGIE ET DU MUSEUM D'HISTOIRE NATURELLE DE LA VILLE DE GENEVE lal GENEVE 1998 ISSN 0035-418X TABLE DES MATIÈRES TOME 105 — 1998 Fascicule 1 WEBER, Claude. Catalogue révisé des types primaires de la collection ichtyo- logique du Muséum d’ histoire naturelle de la Ville de Genève (MHNG). . . . MARTI, Philippe & Jean WUEST. Répartition géographique et morphologie fione de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). ............. ROWELL, C. Hugh F. A revision of the genus Munatia Stàl, 1875 (Orthoptera, Caelifera, Romaleidae, Romaleinae). ................................ HUBER, Bernhard A. Report on some pholeid spiders collected in Guatemala and Hondurasi(Araneae-sPholcidae)i A E, OO I A seo LANG, Claude. Using a submarine to monitor the biological recovery of deep sediments in Lake Geneva (Switzerland). ............................ FRISCH, Johannes. A revision of some West Palaearctic species of Scopaeus Erichson (Coleoptera, Staphylinidae, Paederinae)...................... ANGELINI, Fernando & Luigi DE Marzo. Supplement to the knowledge of the Agathidiini of Taiwan (Coleoptera, Leiodidae)........................ PACE, Roberto. Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae). . . .. Fascicule 2 KLIMASZEWSKI, Jan & Stewart B. PECK. A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphylinidae, Aleo- CHAFIMAC) ie se ALA OL dedica DER left BAEHR, Martin. Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae)..................... VASILEVA, Gergana P., Boyko B. GEORGIEV & Todor GENOV. Redescription of Hymenolepis hoploporus Dollfus, 1951, with the erection of the new genus Dollfusilepisi(GestodasHymenolepididae) 4444444008004 eee ae Pages 3-14 15-23 25-48 49-80 81-88 89-124 125-138 139-220 221-260 261-318 319-329 VI TABLE DES MATIERES HUBER, Jean H. A new Cyprinodont species with a uniquely-colored female, Aphyosemion hera n. sp. (Cyprinodontiformes, Pisces), from northwestern DALENS, Henri. Endémisme pyrénéen: sur une nouvelle espèce épigée du genre Oritoniscus: O. rousseti n. sp. (Crustacea, Isopoda, Oniscidea)............ COMELLINI, André. Notes sur les Psélaphines néotropicaux (Coleoptera, Staphy- linidae, Pselaphinae) 10 - un nouveau genre et quatre nouvelles espèces de la CES MEN ES MR IE ANGELINI, Fernando & Luigi DE Marzo. Agathidiini from China, with description ofglAimewsspeciesi(Coleoptera Lelodidae) RE SCHAWALLER, Wolfgang. The genus Laena Latreille (Coleoptera: Tenebrionidae) inghhailandSwithidescupions'ofinewaspecies 5.20.46 45 RE PUTHZ, Volker. Die Gattung Stenus Latreille in Vietnam (Coleoptera, Staphy- indio) ec UN COTE PACE, Roberto. Aleocharinae della Cina: Parte II (Coleoptera, Staphylinidae)..... Fascicule 3 Pozzi, Stefano, Yves GONSETH & Ambros HANGGI. Evaluation de l’entretien des prairies sèches du plateau occidental suisse par le biais de leurs peuplements arachnologiquesi(ArachnidazAraneae) oe = EE ASSING, Volker. A revision of the Habrocerinae of the world. Supplement II (Colcoptera:#Staphiylimida)t OOO IE UHMANN, Gerhard. Beschreibung von vier neuen Arten der Gattung Derarimus (Coleoptera Anthieidae) aus Malaysia Beer EEA MARTENS, Jochen & Peter SCHWENDINGER. A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opi- lionesSfaniatores): u... 2328,04: i rs ae EROE TAN, Heok Hui & Maurice KOTTELAT. Redescription of Betta picta (Teleostei: Osphronemidae) and description of B. falx sp. n. from central Sumatra. ... . BARBALAT, Sylvie. Importance of forest structures on four beetle families (Col.: Buprestidae, Cerambycidae, Lucanidae and phytophagous Scarabaeidae) in theyAreuseiGorsesi(lNeuchaätel, Switzerland) 22 2 eee DE ANDRADE, Maria L. Fossil and extant species of Cylindromyrmex (Hymen- @ptera: Formieidae)..... re ose Age CH Se PACE, Roberto. Aleocharinae della Cina: Parte III (Coleoptera, Staphylinidae).... 331-338 339-343 345-350 351-373 375-382 383-394 395-463 465-485 487-492 493-497 499-555 557-568 569-580 581-664 665-732 TABLE DES MATIERES Fascicule 4 ZOOLOGIA ET BOTANICA ‘98, Geneva, 18-20 February 1998 (Joint meeting of the Swiss Society of Zoology and the Swiss Society of Botany)............. BAUD, François J. Nécrologie Louis de Roguin (1948 - 1998)................ MAHNERT, Volker & Renata DE ANDRADE. Description of a new troglophilous species of the genus Maxchernes Feio, 1960 (Pseudoscorpiones, Cherne- tidac) sromuBrazile(SaorRaulorState) EN AE EP EE ASSING, Volker. New species and records of Masuria Cameron from Nepal (Coleoptera StaphylinidacyAleochaninae) sw a. O een Te LOURENÇO, Wilson R. & Lionel Monop. Redescription of Compsobuthus rugo- sulus (Pocock, 1900) (Scorpiones, Buthidae) based on specimens from [PUSSIES 00 AL de eS CPO TRL Gree, CONE CORE I Re PE ZEIDAN-GEZE, Najla & Daniel BURCKHARDT. The jumping plant-lice of Lebanon (ElemipterasBsylloidea) o WUEST, Jean. Les organes producteurs de phéromones de quelques Hespérides (dFepidopreraszilesperidaestleSpetliMae) Rs ee KURBATOV, Sergei A. & Ivan LOBL. Nouvelles espèces asiatiques du genre Bryaxis et quelques données sur des espèces connues (Coleoptera: Staphy- [inidae#Esclaphim DE AE NITTO ANGELINI, Fernando & Jonathan COOTER. A new species of Stetholiodes Fall, 1910 (Coleoptera, Leiodidae, Agathidiini) from Taiwan................. MAHUNKA, Sändor. New data on Oribatids (Acari: Oribatida) from St. Lucia (Antlles)2(AcarologiealGenavensia III PAGÉS, Jean. Japygoidea (Diplura) du Sud-Est asiatique n° 8: Indonésie (Java, Bali), Singapour et Brunei - Dicellurata Genavensia XXIII -............. AZPELICUETA, Maria de las Mercedes & Atila Esteban GoszToNYI. Redescription of Diplomystes mesembrinus (Siluriformes, Diplomystidae).............. PACE, Roberto. Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). . . . VII Pages 733-765 767-770 771-775 777-787 789-796 797-812 813-822 823-833 835-837 839-877 879-899 901-910 911-982 Fi Lk ae NUE À té a n # nt ; ig gd ves le N i AA TO Me dom eee LES i di E 14 ehe, INDEX DES AUTEURS par ORDRE ALPHABÉTIQUE ANGELINI, Fernando & Jonathan COOTER. A new species of Stetholiodes Fall, 1910 (Coleoptera, Leiodidae, Agathidiini) from Taiwan. ............... ANGELINI, Fernando & Luigi De MARZO. Agathidiini from China, with description ofalZonewaspecics (Coleoptera, lee1odidac) meer ANGELINI, Fernando & Luigi DE Marzo. Supplement to the knowledge of the Agathidiniyotlaiwani(Coleopterayle1odidac) 5 Re, i nno ASSING, Volker. A revision of the Habrocerinae of the world. Supplement II (Coleoptera Sta piylii di) RETTA e ASSING, Volker. New species and records of Masuria Cameron from Nepal (@oleopteragStaphylinidaesAleocharinae)e 22. 2. ra: AZPELICUETA, Maria de las Mercedes & Atila Esteban GoszTonyı. Redescription of Diplomystes mesembrinus (Siluriformes, Diplomystidae). ............. BAEHR, Martin. Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae). .................... BARBALAT, Sylvie. Importance of forest structures on four beetle families (Col.: Buprestidae, Cerambycidae, Lucanidae and phytophagous Scarabaeidae) in thé AreuselGorsesi(Neuchate Switzerland) IRR BAUD, François J. Nécrologie Louis de Roguin (1948 - 1998)................ COMELLINI, André. Notes sur les Psélaphines néotropicaux (Coleoptera, Staphy- linidae, Pselaphinae) 10 - un nouveau genre et quatre nouvelles espèces de la (MDUITESIMELO PIASINI PERS TM ER NE A DALENS, Henri. Endémisme pyrénéen: sur une nouvelle espèce épigée du genre Oritoniscus: O. rousseti n. sp. (Crustacea, Isopoda, Oniscidea). ........... DE ANDRADE, Maria L. Fossil and extant species of Cylindromyrmex (Hymen- Opterawghk ONNICIGAS) AN EI Me a hee Ae EE ae Te FRISCH, Johannes. A revision of some West Palaearctic species of Scopaeus Erichson (Coleoptera, Staphylinidae, Paederinae). ..................... HUBER, Bernhard A. Report on some pholeid spiders collected in Guatemala and ondurasi¢Araneaes Pholcidae) EEE RR eee ee Huser, Jean H. A new Cyprinodont species with a uniquely-colored female, Aphyosemion hera n. sp. (Cyprinodontiformes, Pisces), from northwestern KLIMASZEWSKI, Jan & Stewart B. PECK. A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphylinidae, Aleo- CHALMAO) ERIN SIVIERO: ARIE Gaels eMC Rt te a Pages 835-837 351-373 125-138 487-492 777-787 901-910 261-318 569-580 767-770 345-350 339-343 581-664 89-124 49-80 331-338 221-260 x INDEX DES AUTEURS KURBATOV, Sergei A. & Ivan LOBL. Nouvelles espèces asiatiques du genre Bryaxis et quelques données sur des espèces connues (Coleoptera: Staphy- finidae sPselaphimae) En ore 20. A ee ee RE LANG, Claude. Using a submarine to monitor the biological recovery of deep sediments in Lake Geneva (Switzerland). ............................ LOURENCO, Wilson R. & Lionel Monop. Redescription of Compsobuthus rugo- sulus (Pocock, 1900) (Scorpiones, Buthidae) based on specimens from PARIS EAN ts Oe TIE ma eo ie RAC ALS G65 0.0 0°0 MARTI, Philippe & Jean WUEST. Répartition géographique et morphologie fione de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). ............. MARTENS, Jochen & Peter SCHWENDINGER. A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opi- lonessEanlatoresy)A e RL ATO OI eee MAHNERT, Volker & Renata DE ANDRADE. Description of a new troglophilous species of the genus Maxchernes Feio, 1960 (Pseudoscorpiones, Cherne- tidae) tromuBrazali(SaoyRauloyState)s 5 ee EE EE ee MAHUNKA, Sandor. New data on Oribatids (Acari: Oribatida) from St. Lucia (Antilles). (Acarologica Genavensia LXXXIX)........................ PACE, Roberto. Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae). . . .. PACE, Roberto. Aleocharinae della Cina: Parte II (Coleoptera, Staphylinidae). . . .. PACE, Roberto. Aleocharinae della Cina: Parte III (Coleoptera, Staphylinidae). . . . PACE, Roberto. Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). . . . PAGÉS, Jean. Japygoidea (Diplura) du Sud-Est asiatique n° 8: Indonésie (Java, Bali), Singapour et Brunei - Dicellurata Genavensia XXIII -............. Pozzi, Stefano, Yves GONSETH & Ambros HANGGI. Evaluation de l’entretien des prairies sèches du plateau occidental suisse par le biais de leurs peuplements arachnologiques (ArachnidaswAraneae) el 222 PUTHZ, Volker. Die Gattung Stenus Latreille in Vietnam (Coleoptera, Staphy- MTG AS) eae E SIRO CALI AI LOT ROWELL, C. Hugh F. A revision of the genus Munatia Stal, 1875 (Orthoptera, Caelifera, Romaleidae, Romaleinae). .. .:.................... onen SCHAWALLER, Wolfgang. The genus Laena Latreille (Coleoptera: Tenebrionidae) inthaland with descriptions Of MEW species. re eee TAN, Heok Hui & Maurice KOTTELAT. Redescription of Betta picta (Teleostei: Osphronemidae) and description of B. falx sp. n. from central Sumatra... . . UHMANN, Gerhard. Beschreibung von vier neuen Arten der Gattung Derarimus (EoleopteraAnthieidae)laus Malaysiar a2. 2 eee VASILEVA, Gergana P., Boyko B. GEORGIEV & Todor GENOV. Redescription of Hymenolepis hoploporus Dollfus, 1951, with the erection of the new genus Dolljusilepis(Gestoda Hymenolepididae) RSS ORE WEBER, Claude. Catalogue révisé des types primaires de la collection ichtyo- logique du Muséum d'histoire naturelle de la Ville de Genève (MHNG). . . WÜEST, Jean. Les organes producteurs de phéromones de quelques Hespérides (ÉEDIdOptErA HESpeIdaE MES PETINAC) re Pages 823-833 81-88 789-796 15-23 499-555 771-775 839-877 139-220 395-463 665-732 911-982 879-899 465-485 383-394 25-48 375-382 557-568 493-497 319-329 3-14 813-822 INDEX DES AUTEURS XI Pages ZEIDAN-GEZE, Najla & Daniel BURCKHARDT. The jumping plant-lice of Lebanon (FemipteraBsyIloidea) e RR I EE eee 797-812 ZOOLOGIA ET BOTANICA ‘98, Geneva, 18-20 February 1998 (Joint meeting of the Swiss Society of Zoology and the Swiss Society of Botany)............. 733-765 to, TOME IZZO] i Pre bene reno Sha entire pere: are TI È Ee A TP ul RUS NH ANNALES de la SOCIETE SUISSE DE ZOOLOGIE et du MUSEUM D'HISTOIRE NATURELLE de la Ville de Genève tome 105 fascicule 1 1998 5 A jal GENEVE MARS 1998 ISSN 0035 - 418 X È A REVUE SUISSE DE ZOOLOGIE REVUE SUISSE DE ZOOLOGIE TOME 105 — FASCICULE 1 Publication subventionnée par l'Académie suisse des Sciences naturelles et la Société suisse de Zoologie VOLKER MAHNERT Directeur du Muséum d'histoire naturelle de Genève FRANÇOIS BAUD È Conservateur au Muséum d'histoire naturelle de Genève CHARLES LIENHARD Chargé de recherche au Muséum d’histoire naturelle de Genève Comité de lecture Président: Ivan LOBL — Muséum de Genève Il est constitué en outre du président de la Société suisse de Zoologie, du directeur du ‘Muséum de Genève et de représentants des Instituts de zoologie des universités suisses. Les manuscrits sont soumis à des experts d’institutions suisses ou étrangères selon le sujet étudié. La préférence sera donnée aux travaux concernant les domaines suivants: biogéo- graphie, systématique, écologie, éthologie, morphologie et anatomie comparée, physiologie. Administration MUSÉUM D'HISTOIRE NATURELLE 1211 GENÈVE 6 Internet: http://www.geneva-city.ch:80/musinfo/mhng/publications/revues.htm PRIX DE L'ABONNEMENT: SUISSE Fr. 225.— UNION POSTALE Fr. 230.— (en francs suisses) Les demandes d'abonnement doivent étre adressées à la rédaction de la Revue suisse de Zoologie, Muséum d'histoire naturelle, C.P. 6434, CH-1211 Genève 6, Suisse ANNALES de la SOCIÉTÉ SUISSE DE ZOOLOGIE et du MUSEUM D'HISTOIRE NATURELLE de la Ville de Genève tome 105 fascicule 1 1998 REVUE SUISSE DE ZOOLOGIE g : kl GENEVE MARS 1998 ISSN 0035 - 418 X REVUE SUISSE DE ZOOLOGIE TOME 105 — FASCICULE 1 Publication subventionnée par l'Académie suisse des Sciences naturelles et la Société suisse de Zoologie VOLKER MAHNERT Directeur du Muséum d'histoire naturelle de Genève FRANÇOIS BAUD Conservateur au Muséum d'histoire naturelle de Genève CHARLES LIENHARD Chargé de recherche au Muséum d'histoire naturelle de Genève Comité de lecture Président: Ivan LOBL — Muséum de Genève Il est constitué en outre du président de la Société suisse de Zoologie, du directeur du Muséum de Genève et de représentants des Instituts de zoologie des universités suisses. Les manuscrits sont soumis à des experts d institutions suisses ou étrangères selon le sujet étudié. La préférence sera donnée aux travaux concernant les domaines suivants: biogéo- graphie, systématique, écologie, éthologie, morphologie et anatomie comparée, physiologie. Administration MUSÉUM D'HISTOIRE NATURELLE 1211 GENÈVE 6 Internet: http://www.geneva-city.ch:80/musinfo/mhng/publications/revues.htm PRIX DE L'ABONNEMENT: | SUISSE Fr. 225.— UNION POSTALE Fr. 230.— (en francs suisses) Les demandes d'abonnement doivent étre adressées | à la rédaction de la Revue suisse de Zoologie, Muséum d'histoire naturelle, C.P. 6434, CH-1211 Genève 6, Suisse REVUE SUISSE DE ZOOLOGIE 105 (1): 3-14; mars 1998 Catalogue révisé des types primaires de la collection ichtyologique du Muséum d'histoire naturelle de la Ville de Genève (MHNG) Claude WEBER Muséum d'histoire naturelle, case postale 6434, CH-1211 Genève 6, Suisse. Revised types catalogue of the ichthyological collection in the Natural History Museum, City of Geneva (MHNG).- The ichthyological collection of the MHNG includes primary type material of 130 species or subspecies. Complete type material (362 taxa), more detailed data and remarks concerning localities, collectors and origin of type specimens can be found under http:/www.ville-ge.ch/musinfo/mhng/erpi/cat2.html Key-words: Catalogue - types - Pisces - Hyperoartia - Elasmobranchii - Osteichthyes - Actinopterygii - Internet - MHNG INTRODUCTION Fondé en 1820, le Muséum d'histoire naturelle de la Ville de Genève (MHNG) s'installe en 1965 dans un bâtiment neuf qui offre alors les meilleures conditions pour la gestion des collections et les activités scientifiques. Dans ce contexte et conjointement à une augmentation significative des collaborateurs scientifiques, certaines structures sont remaniées: en 1971 est créé le département d'herpétologie et d'ichtyologie, issu de la division de celui des vertébrés. Les priorités muséologiques de ce nouveau département sont la mise en ordre des collections et la publication du catalogue des types de poissons, amphibiens et reptiles (MAHNERT 1976), qui ne compte alors que 26 espèces de poissons, holotypes et autres catégories comprises. Depuis 1965, le MHNG a développé son activité scientifique: collaborations régulières avec des spécialistes suisses ou étrangers, formation d'étudiants, organi- sation de campagnes de récoltes, souvent associées à des programmes d'inventaires biologiques ou à des études d'impact, politique d'acquisition et d'échanges orientés aussi sur la recherche, sont des éléments qui ont largement contribué à l'accroissement spectaculaire des collections scientifiques. Avec un total de 362, le nombre des espèces ou sous-espèces de poissons représenté par des types a plus que décuplé depuis 1976, et un nouveau catalogue s'imposait. Le présent travail consacré aux poissons sensu lato, constitue la deuxième partie de la révision du catalogue des types de MAHNERT (1976) après la publication du catalogue des types d'amphibiens et de reptiles par SCHATTI & PERRET (1997a). Manuscrit accepté le 15.07.1997 4 CLAUDE WEBER MÉTHODES La présente liste des types compte 130 taxa répertoriés par ordre systématique des ordres selon la classification d'ESCHMEYER (1990), et alphabétiques des noms de genres, espèces et sous espèces, selon la nomenclature originale. Elle ne mentionne que les types primaires (holotypes, lectotypes, néotypes et syntypes), sans autres indications que la catégorie de type, le numéro MHNG et le nombre de spécimens. Les espèces mentionnées par MAHNERT (1976) sont précédées d'un astérisque. De même que le catalogue des types d'amphibiens et de reptiles possède une version plus détaillée sur serveur électronique (SCHATTI & PERRET 1997b), cette liste succincte est publiée conjointement à la création d'un accès Internet au catalogue complet des types (y compris paratypes et paralectotypes), détruits ou disparus inclus, avec mention détaillée des localités, dates de capture et récolteurs. Des précisions concernant leur historique sont ajoutées en remarques. Ce catalogue, régulièrement mis à jour, est accessible à l'adresse: http://www.ville-ge.ch/musinfo/mhng/erpi/cat2.html PRINCIPALES ADDITIONS AU CATALOGUE DES TYPES DEPUIS 1976. L'apport le plus important provient de la fructueuse collaboration entretenue depuis 1971 avec Jacques Géry, spécialiste des poissons Characiformes. Sa partici- pation à l'étude de nombreuses récoltes, dont celles réalisées par le MHNG, et surtout le dépôt, en 1984, de sa collection personnelle, qui ne compte pas moins de 20.000 spécimens, ont apporté une quantité considérable de matériel type, représentant 205 espèces ou sous-espèces, dont 43 par des types primaires. Géry est auteur ou coauteur de 130 d'entre-elles, les 75 autres (décrites par Cope, Eigenmann, Myers, Schultz, etc.) proviennent d'échanges réalisés avec des instituts nord-américains. Plusieurs autres ichtyologistes ont contribué de l'extérieur à l'enrichissement de la collection des types: citons notamment Maurice Kottelat, qui a déposé des spé- cimens types de 21 espèces, dont 8 sont représentés par des types primaires, ainsi que Sven Kullander, qui étudie régulièrement les Cichlidés récoltés au cours des missions du MHNG, avec 13 espèces, dont 4 sont représentées par des types primaires. Enfin, depuis le précédent catalogue, les collaborateurs du MHNG ont décrit 25 espèces, dont 14 sont représentées par des types primaires. La révision des anciennes collections a mis en évidence quelques types ignorés et a surtout permis d'identifier du matériel type connu mais appartenant à des collections à problèmes (nomenclature confuse, mélanges de spécimens, etc.). Les types de Victor Fatio font partie de ces derniers. Ils sont maintenant revus, leur nomenclature est discutée en détail dans le travail de KOTTELAT (1997) et 16 taxa ont été retenus dans le catalogue. Outre celà, la consultation de diverses archives du Muséum a mis en lumière l'existence d'une petite collection de poissons envoyés en échange par Achilles Valenciennes. Nombre d'entre eux sont des spécimens historiques, et 6 syntypes d'espèces décrites dans l'Histoire naturelle des Poissons de CUVIER ET VALENCIENNES (1828-1849) ont été retrouvés. TYPES DE LA COLLECTION ICHTYOLOGIQUE DU MHNG CATALOGUE HYPEROARTIA Classe CEPHALASPIDOMORPHI Ordre Petromyzontiformes Petromyzon branchialis Linné, 1758 Neotype: MHNG 816.18 Petromyzon fluviatilis Linné, 1758 Neotype: MHNG 816.18 GNATHOSTOMATA Chondrichtyes Classe ELASMOBRANCHII Ordre Myliobatiformes *Pteroplatea binotata Lunel, 1879 Holotype: MHNG 1213.89 Osteichthyes Classe ACTINOPTERYGII Ordre Osteoglossiformes *Marcusenius cubangoensis Pellegrin, 1936 Syntype: MHNG 858.85 (1) Ordre Anguilliformes *Gymnomuræna brevicauda Regan, 1903 Holotype: MHNG 665.54 *Muræna grandimaculis Regan, 1903 Holotype: MHNG 665.39 Ordre Clupeiformes Alosa Finta var. lacustris Fatio,1890 Lectotype: MHNG 656.48 Ordre Cypriniformes Alborella maxima Fatio, 1882 Holotype: MHNG 815.80 6 CLAUDE WEBER Barbichthys laevis var. sumatranus Volz, 1904. Syntype: MHNG 683.23 (1) Barbus condei Mahnert & Géry 1982 Holotype: MHNG 1544.49 Barbus foerschi Kottelat, 1982 Holotype: MHNG 2058.98 Blicca intermedia Fatio, 1882 Holotype: MHNG 656.16 *Botia multifasciata Regan, 1905 Holotype: MHNG 677.98 *Brama saussurii Lunel, 1865 Holotype: MHNG 940.85 Ciprinus [sic] agonus Scopoli, 1786 Neotype: MHNG 656.48 Nemacheilus baenzigeri Kottelat, 1983 Holotype: MHNG 2081.32 Nemacheilus troglocataractus Kottelat & Géry, 1989 Holotype: MHNG 2407.54 Osteochilus pentalineatus Kottelat, 1982 Holotype: MHNG 2059.02 Pectenocypris korthausae Kottelat, 1982 Holotype: MHNG 2073.72 Phoxinellus libani Lortet, 1883 Syntypes: MHNG 611.24 (10) Rasbora hobelmani Kottelat, 1984 Holotype: MHNG 2160.46 Rhodeus syriacus Lortet, 1883 Syntypes: MHNG 611.22 (2) Ordre Characiformes Anostomus anostomus longus Géry, 1961 Holotype: MHNG 2197.04 Asiphonichthys condei Géry & Knôppel,1976 Holotype: MHNG 2229.05 TYPES DE LA COLLECTION ICHTYOLOGIQUE DU MHNG Astyanax validus Géry, Planquette & Le Bail, 1991 Holotype: MHNG 2435.77 Axelrodia lindeae Géry, 1973 Holotype: MHNG 2229.08 Brycinus derhami Géry & Mahnert, 1977 Holotype: MHNG 1183.06 Brycinus fwaensis Géry, 1995 Holotype: MHNG 2572.07 Chilobrycon deuterodon Géry & de Rham, 1981 Holotype: MHNG 2045.13 Creagrutus paraguayensis Mahnert & Géry, 1988 Holotype: MHNG 2386.01 Geisleria junki Géry, 1971 Holotype: MHNG 2229.07 Hemigrammus aereus Géry, 1959 Holotype: MHNG 2181.86 Hemigrammus guyanensis Géry, 1959 Holotype: MHNG 2181.23 Hemigrammus mahnerti Uj & Géry, 1989 Holotype: MHNG 2412.82 Hemigrammus micropterus boesemani Géry, 1959 Holotype: MHNG 2181.80 Hemigrammus unilineatus cayennensis Géry, 1959 Holotype: MHNG 2179.61 Hemiodopsis huraulti Géry, 1964 Holotype: MHNG 2151.14 Hemiodopsis vorderwinckleri Géry, 1964 Holotype: MHNG 2151.20 Hyphessobrycon arianae Uj & Géry, 1989 Holotype: MHNG 2412.79 Hyphessobrycon guarani Mahnert & Géry, 1987 Holotype: MHNG 2366.99 Hyphessobrycon procerus Mahnert & Géry, 1987 Holotype: MHNG 2385.68 8 CLAUDE WEBER Hyphessobrycon pytai Géry & Mahnert, 1993 Holotype: MHNG 2543.86 Hyphessobrycon simulans Géry, 1963 Holotype: MHNG 2171.19 Hyphessobrycon vilmae Géry, 1966 Holotype: MHNG 2229.04 Iguanodectes adujai Géry, 1970 Holotype: MHNG 2229.06 Iguanodectes geisleri Géry, 1970 Holotype: MHNG 2229.02 Jobertina eleotrioides Géry, 1960 Holotype: MHNG 2201.13 Laemolyta garmani macra Géry, 1974 Holotype: MHNG 2197.38 Laemolyta petiti Géry, 1964 Holotype: MHNG 2229.01 Leporinus lebaili Géry & Planquette, 1983 Holotype: MHNG 2152.48 Micralestes ambiguus Géry, 1995 Holotype: MHNG 2572.08 Microbrycen Cochui Ladiges, 1950 Syntype: MHNG 2187.75 (1) Microschemobrycon geisleri Géry, 1973 Holotype: MHNG 2229.09 Petitella georgiae Géry & Boutière, 1964 Holotype: MHNG 2150.28 Phenacogrammus bleheri Géry, 1995 Holotype: MHNG 2572.09 Phenacogrammus taeniatus Géry, 1996 Holotype: MHNG 2583.28 Piabarchus torrenticola Mahnert & Géry, 1988 Holotype: MHNG 2385.70 *Pseudochalceus longianalis Géry, 1972 Holotype: MHNG 1226.90 TYPES DE LA COLLECTION ICHTYOLOGIQUE DU MHNG Rhinopetita myersi Géry, 1964 Holotype: MHNG 2229.03 Steindachnerina varii Géry, Planquette & Le Bail, 1991 Holotype: MHNG 2435.76 Thayeria ifati Géry, 1959 Holotype: MHNG 2173.42 Tyttobrycon hamatus Géry, 1973 Holotype: MHNG 2172.30 Tyttobrycon xeruini Géry, 1973 Holotype: MHNG 2229.10 Ordre Siluriformes *Amphilius platychir var. cubangoensis Pellegrin, 1936 Syntype: MHNG 858.86 (1) Ancistrus pirareta Muller, 1989. Holotype: MHNG 2450.10 Ancistrus piriformis Muller, 1989 Holotype: MHNG 2450.11 Callomystax schmidti Volz, 1904 Syntype: MHNG 683.22 (1) Cetopsis candiru Spix et Agassiz, 1829 Syntype: MHNG 210.05 (1) Dysichthys quadriradiatus Mees, 1989 Holotype: MHNG 2157.21 Erethistes maesotensis Kottelat, 1983 Holotype: MHNG 2096.63 Farlowella platorhynchus Retzer & Page, 1997 Holotype: MHNG 2389.57 Hypostomus dlouhyi Weber, 1985 Holotype: MHNG 2229.43 Hypostomus latifrons Weber, 1986 Holotype: MHNG 2256.67 Hypostomus microstomus Weber, 1987 Holotype: MHNG 2367.90 10 CLAUDE WEBER Hypostomus piratatu Weber, 1986 Holotype: MHNG 2265.03 Lepthoplosternum altamazonicum Reis, 1997 Holotype: MHNG 2551.01 *Macrones argentivittatus Regan, 1905 Lectotype: MHNG 677.99 Macrones bimaculatus Volz, 1904 Syntype: MHNG 683.28 (1) Mystus misrai Anuradha, 1986 Holotype: MHNG 603.95 *Phreatobius cisternarum Geeldi, 1904 Syntypes?: MHNG 1213.97 (1) et MHNG 1505.91 (3) Ordre Gymnotiformes Gymnorhamphichthys hypostomus petiti Géry & Vu, 1964 Holotype: MHNG 2167.15 Ordre Salmoniformes Coregonus asperi dispar Fatio, 1885 Syntypes: MHNG 807.48 (3) Coregonus asperi maraenoides Fatio, 1885 Syntypes: MHNG 816.42 (2) Coregonus Balleus Fatio, 1885 Lectotype: MHNG 717.45 Coregonus candidus Goll, 1883 Neotype: MHNG 656.36 Coregonus crassirostris compactus Fatio, 1885 Syntypes: MHNG 715.93 (2) Coregonus exiguus albellus Fatio, 1890 Lectotype: MHNG 816.22 Coregonus fatioi Kottelat, 1997 Lectotype: MHNG 809.59 Coregonus lavaretus (Linné, 1758) Neotype: MHNG 2583.51 TYPES DE LA COLLECTION ICHTYOLOGIQUE DU MHNG tl] Coregonus nobilis Haack, 1882 Neotype: MHNG 656.56 Coregonus restrictus bondella Fatio, 1885 Lectotype: MHNG 656.36 Coregonus restrictus Niisslini Fatio, 1885 Syntypes: MHNG 715.94 (2) Coregonus Schinzii alpinus Fatio, 1885 Lectotype: MHNG 717.45 Coregonus Suidteri Fatio, 1885 Syntypes: MHNG 676.07 (1), MHNG 715.89 (1), MHNG 816.26 (1) Coregonus Wartmanni Alpinus Fatio, 1890 Nom indisponible (Kottelat 1997), voir Coregonus fatioi Kottelat, 1997. Coregonus Wartmanni dolosus Fatio, 1885 Lectotype: MHNG 656.53 Salmo lacustris var. excelsa Fatio, 1890 Syntype: MHNG 816.09 (2) Salmo lacustris var. meridionalis Fatio, 1890 Lectotype: MHNG 656.06 Salmo lacustris var. Rhenana Fatio, 1890 Syntype: MHNG 806.96 (1) Salvelinus salvelinus var. profundus Fuhrmann, 1903 Syntype: MHNG 809.61 (1) Trutta variabilis Lunel, 1874 Syntypes: MHNG 807.35 (1), MHNG 816.07 (1), MHNG 816.06 (1) Ordre Atheriniformes *Bedotia madagascariensis Regan, 1903 Holotype: MHNG 665.07 Ordre Cyprinodontiformes Rivulus elongatus Fels & de Rham, 1981 Holotype: MHNG 2079.63 Rivulus intermittens Fels & de Rham, 1981 Holotype: MHNG 2079.33 12 CLAUDE WEBER Rivulus iridescens Fels & de Rham, 1981 Holotype: MHNG 2079.57 Rivulus rectocaudatus Fels & de Rham, 1981 Holotype: MHNG 2079.10 Rivulus rubrolineatus Fels & de Rham, 1981 Holotype: MHNG 2079.46 Rivulus speciosus Fels & de Rham, 1981 Holotype: MHNG 2079.68 Ordre Syngnathiformes *Doryichthys multiannulatus Regan, 1903 Holotype: MHNG 665.51 *Penetopteryx tæniocephalus Lunel, 1881 Syntypes: MHNG 843.48 (3) Ordre Scorpaeniformes Apistus dracoena Cuvier, 1829 Syntype: MHNG 148.43 Ordre Perciformes Aequidens patricki Kullander, 1984 Holotype: MHNG 2163.93 Ambassis alta Cuvier, 1828 Syntype: MHNG 148.06. Ambassis Commersoni Cuvier, 1828 Syntype: MHNG 148.04 Apistogramma nijsseni Kullander, 1974 Holotype: MHNG 1595.82 Apogon rex mullorum, var. Americana Pictet, 1836 Syntype: MHNG 1060.89 (1) Bujurquina ortegai Kullander, 1986 Holotype: MHNG 2205.27 Chromis magdalenae Lortet, 1883 Syntypes: MHNG 611.21 (2) TYPES DE LA COLLECTION ICHTYOLOGIQUE DU MHNG 13 Chromis tiberiadis Lortet, 1883 Syntype: MHNG 611.19 (1) Cichlasoma pusillum Kullander, 1983 Holotype: MHNG 2131.94 Datnia virgata Valenciennes, 1831 Syntype: MHNG 148.22 *Eleotris pectoralis Regan, 1903 Holotype: MHNG 665.06 Gymnogeophagus setequedas Reis, Malabarba & Pavanelli, 1992 Holotype: MHNG 2518.19 *Heros octofasciatus Regan, 1903 Holotype: MHNG 665.55 Lates nobilis Cuvier, 1828 Syntype: MHNG 148.01 *Novacula temporalis Regan, 1905. Holotype: MHNG 678.02 *Percichthys altispinis Regan, 1905 Syntypes: MHNG 677.100 (2) Polynemus longifilis Cuvier, 1829 Syntype: MHNG 148.24 *Sciæna (Bairdiella) bedoti Regan, 1905 Syntype: MHNG 678.01 (1) Ordre Pleuronectiformes *Solea borbonica Regan, 1905 Holotype: MHNG 678.03 Ordre Tetraodontiformes *Balistes mauritianus Regan, 1903 Holotype: MHNG 665.43 REMERCIEMENTS Que soient remerciés ici tous ceux qui ont apporté aide et collaboration à ce travail, en particulier Sonia Muller, Nadia Zoppi, Jacques Géry, Pierre-Joseph Haymoz, Maurice Kottelat et Volker Mahnert. 14 CLAUDE WEBER RÉFÉRENCES CUVIER, G. L. C. & A. VALENCIENNES. 1829-1849. Histoire naturelle des Poissons. 24 vol. Levrault, Paris. ESCHMEYER, W. N. 1990. Catalog of the Genera of Recent Fishes. California Academy of Sciences, San Francisco. pp.697. KOTTELAT, M. 1997. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, Sect. Zool. 52 (suppl. 5). 1-271 MAHNERT, V. 1976. Catalogue des types de poissons, amphibiens et reptiles du Muséum d'histoire naturelle de Genève. Revue suisse de Zoologie 83 (2):471-496 SCHATTI, B & J.-L. PERRET. 1997a. Catalogue révisé des types d'amphibiens et de reptiles du Muséum d'histoire naturelle de Genève. Revue suisse de Zoologie 104 (2): 357-370. SCHATTI, B & J.-L. PERRET. 1997b. Catalogue révisé des types d'amphibiens et de reptiles du Muséum d'histoire naturelle de Genève. http://www.ville-ge.ch/musinfo/mhng/erpi/ catl.html REVUE SUISSE DE ZOOLOGIE 105 (1): 15-23; mars 1998 Répartition géographique et morphologie fine de Broelemanneuma gayi (Diplopoda: Craspedosomatidae) Philippe MARTI! & Jean WÜEST? | Departement de Zoologie et de Biologie animale, Station de Zoologie expérimentale, 154, rte de Malagnou, CH-1224 Chêne-Bougeries, Switzerland. 2Muséum d'histoire naturelle, 1, rte de Malagnou, case postale 6434, CH-1211 Genève 6, Switzerland. Geographical distribution and micromorphology of Broelemanneuma gayi (Diplopoda: Craspedosomatidae). - The distribution and ecology of the craspedosomatid diplopod Broelemanneuma gayi Demange, 1968 is discussed and morphological characters are illustrated by SEM. Previously unknown characters on fore legs in both sexes are described and illustrated. Key-words: Broelemanneuma gayi - Diplopoda - Craspedosomatidae - geographical distribution - secondary sexual characters. INTRODUCTION Lors d’explorations dans des grottes du bassin de Flaine, l’un de nous (P. M.) a procédé à la pose de pièges et nous avons eu la chance de récolter en abondance le Diplopode Broelemanneuma gayi Demange, 1968. Cela représente une nouvelle localité pour ce troglobie limité à la Haute-Savoie. Nous nous proposons de discuter sa répartition géographique et son écologie. D’autre part, l'examen de sa morphologie fine nous a permis de mettre en évidence un nouveau caractère sexuel secondaire présent chez les mâles ainsi que des peignes de soies modifiées sous les deux premières paires de pattes des deux sexes, que nous décrivons pour la première fois. Au sujet de ce Diplopode Craspedosomatidae, une remarque d’ordre nomen- clatural s'impose. L’orthographe du nom de genre est très variable suivant les sources: nous avons trouvé Brolemanneuma, Brohlemanneuma, Bròlemanneuma ou Broelemanneuma. D'ailleurs, le dédicataire de ce genre orthographiait son nom indifféremment Broelemann ou Brölemann avant 1920, et Brolemann, sans tréma ni «oe» après 1920. Si nous nous référons à la description originale de VERHOFF (1905), l’orthographe est Brölemanneuma, soit Broelemanneuma puisque le code de nomen- clature zoologique conseille de supprimer les accents dans les noms de taxa. Manuscrit accepté le 10.07.1997 16 PHILIPPE MARTI & JEAN WÜEST MATÉRIEL ET MÉTHODES Les pièges (gobelets en plastique remplis de bière) ont été posés dans le gouffre du Calumet (bassin de Flaine) entre -50 et -60m de la surface (couche de calcaire urgonien, Fig. 1), soit dans un éboulis et entouré de cailloux pour en faciliter l’accès aux animaux (piège A), soit contre la paroi d’une marmite remplie d’argile (piège B). La pose des pièges a été effectuée le 11 août 1996 et la récolte le 2 novembre , soit 83 jours plus tard. Pour notre étude en microscopie électronique à balayage, les exemplaires utilisés ont été déshydratés dans une série d’alcool, passés dans l’acétate de méthyle et séchés par la méthode du point critique dans du CO, liquide. Ils ont été métallisés par pulvérisation cathodique d’or et observés dans le MEB ZEISS 940A du Muséum de Genève. FIG. 1 Relevé topographique du début du gouffre du Calumet, indiquant l’emplacement des pièges (d’après FAVRE 1994, échelle en m). BROELEMANNEUMA GAYI, RÉPARTITION, CSS 17 RESULTATS RÉCOLTES Les récoltes ont été faites au Gouffre du Calumet, dans le bassin de Flaine par l’un de nous (P.M.). Le piège A, situé dans un éboulis peu stable de pierres de toutes tailles (Fig. 1) a donné un grand nombre (une cinquantaine) de Diplopodes, compre- nant des mâles et des femelles, ainsi qu’un Diptère Trichoceridae du genre Tricho- cera. Ce genre est typique des grottes, et l’espèce T. regelationis (L.) est trogloxène et fréquemment signalée de grottes d’ Europe centrale. Dans le piège B (marmite avec argile, Fig. 1), nous avons récolté un Diplopode ? de même qu’un Diptère Trichocera. Sur la base des gonopodes des mâles, ce Diplopode a pu être attribué à l’espèce Broelemanneuma gayi Demange 1968, selon DEMANGE (1981). La récolte a été révisée par J.-P. Mauriès du Muséum de Paris. Quelques exemplaires sont déposés dans les collections du MNHN de Paris. Le reste, soit 16 S, 5 d immatures et juvéniles, 25 9 et juvéniles, est déposé au MHN de Genève. Le Gouffre du Calumet constitue une localité nouvelle pour B. gayi (Fig. 2, point 1). Ce gouffre se trouve le long d’une faille qui part du col de Monthieu en direction de Flaine et son entrée se trouve à une altitude de 2150 m. Cette grotte est à considérer comme froide, comme toutes les cavités situées à plus de 1500m d’altitude. Il y a beaucoup de courants d’air en été. Par contre, pendant la saison hivernale, l’entrée est complètement obstruée par la neige et il n’y a donc plus de courants d’air. RÉPARTITION GÉOGRAPHIQUE DE B. gayi Cette espèce a été décrite de la grotte de la Diau, près d’ Annecy (DEMANGE 1968). Elle n’aurait plus été signalée depuis selon DEMANGE (1981), et ne figure pas dans les listes récapitulatives de GEOFFROY & MAURIES (1992); aucune référence concernant ce genre ne se trouve non plus dans le Zoological Record entre 1981 (où figure l’ouvrage de DEMANGE (1981)), et 1996. Pourtant, elle était déjà connue de la grotte de la Barme Froide (BOURNE 1975a), mais cette mention a paru dans un périodique plutòt confidentiel, sans préciser le nombre d’individus récoltés. Elle avait été précédée, dans la méme grotte, de la capture d’une femelle “Craspédosomide indéterminable” selon Demange (BOURNE 1973). Il se pourrait qu’il s’agisse de la même espèce étant donné la localisation et l’appartenance à la même famille. D’autre part, DEMANGE (1970) la signale de deux grottes de Haute-Savoie, la grotte du Vieux Taquin et le gouffre de la Tournette et MEYSSONNIER et al. (1987) y ajoutent le Gouffre Jean-Bernard, la grotte et la mine de fer de Sambuy. M. Mauriès nous a signalé encore quelques exemplaires des collections du Muséum de Paris provenant de la Grotte de Charrieu à Tournette et de la Grotte du Désert de Platé. Ces indications ne modifient pas la chorologie de l’espèce. La grotte de Charrieu fait partie du complexe de cavités du massif de la 18 PHILIPPE MARTI & JEAN WÜEST Tournette (point 3 de la Fig. 2). La grotte du Désert de Platé fait partie de l’ensemble de cavités de ce massif de lapiaz, comme la Barme Froide (point 4 de la Fig. 2). Nous complétons cette liste avec le Gouffre du Calumet dans le massif de Flaine (Fig. 2, point 1). Reculet A CF Genève A A Dents du Midi À , Tanin À ges +2 Fe È Recto Bl : Cluses Sixt A Filiere : © A R 55 Pointe Percée À A + 1 A A *4 Arve f Chamonix Annecy Sallanches A Y Thones Grandes Jorasses À Id A DÀ 7 Mont Blanc . Faverges\ g = N te) “6 Ja 4 Fic. 2 Carte des localités où le Diplopode Broelemanneuma gayi a été signalé: 1. Gouffre du Calumet, 1996 (938,3; 119,1; 2150m) (Massif du Plate, Flaine) - 2. Gouffre Jean-Bernard, 1982-1983 (943,41; 132,15; 1860m) (Samoéns) (ARIGNANO 1982, 1983; GEOFFROY 1983) - 3. Gouffre de la Tournette, 1969 (To I; 907,50; 100,56; 1760m) (Massif des Bornes) (DEMANGE 1970); Grotte de Charrieu, à Tournette, 28.11.71, coll. Deharveng, 1 6, 1 © (det. Mauriès). - 4. Grotte de la Barme Froide, (941,63; 118,88; 2060m) (Massif de la Barme Froide, vallon de Laouchet) (BOURNE 1973); Grotte du Désert de Plate, 12.7.64, coll. Guignard, 1 d (det. Mauriès) (Coll. MNHN Paris, inédit). - 5. Gouffre du Vieux Taquin, 1969 (1550m) (Mont Saxonnex) (DEMANGE 1970) - 6. Grotte de la Sambuy, 1984 (906,08; 84,51; 2020m) (Seythenex) (MEYSSONNIER 1984) - 6bis. Mine de fer de la Sambuy, 1986 (905,92; 84,84; 2045m) (Seythenex) (MEYSSONNIER et al. 1987) - 7. Grotte de la Diau, 1967 (905,80; 114,11; 962m) (Massif des Bornes, Mt Parmelan-Pertuis-Mt Terret) (DEMANGE 1968). BROELEMANNEUMA GAYI, RÉPARTITION, CSS 19 TABLEAU Distances (en km) entre les grottes où le Diplopode Broelemanneuma gayi a été signalé. Calumet Jean Tournette | Barme Vieux Bernard I Froide Taquin 907.5 941.63 919.01 100.56 118.88 123.19 2060 1550 Winther ee ee 602 tto "EEE RN RA 7. Diau MORPHOLOGIE FINE L’adaptation à une existence troglobie est manifeste du fait que l’espèce est totalement blanche et aveugle (Fig. 3). Dans l’ordre des Craspedosomatida, ce sont les paires de pattes 8 et 9 du mâle qui sont modifiées en organes copulateurs (Figs 6 et 7). L’espece est caractérisée par la forme des gonopodes 8 qui sont transformés en lames recouvertes de petits tubercules, absents chez les autres représentants du genre Broe- lemanneuma (Figs 6 et 7). La seconde paire de gonopodes (pattes 9) est relativement peu modifiée et présente l’allure de pattes raccourcies (Fig. 6). Les orifices génitaux mâles se présentent comme deux papilles sur les coxae de la seconde paire de pattes (Fig. 5). Il n'y a pas de pénis dans la famille des Craspedosomatidae. L’ orifice génital des femelles ne présente aucune ornementation spécifique pouvant aider à la déter- mination (Figs 3 et 4). En examinant en détail des specimens de B. gayi, nous avons découvert un nouveau caractère sexuel secondaire, qui concerne les pattes 3 à 7 du mâle. L’article tarsal terminal est dilaté à l’extrémité et pourvu, à sa face ventrale, d’une cinquantaine de soies modifiées (Figs 8 et 9). Portées par un court pédoncule, elles sont élargies en raquettes à l’extrémité. Placées sans ordre apparent, elles mesurent environ 12 u de hauteur. Le type de cuticule, sur les faces de la raquette, semble différer de celui du pédoncule, ce qui pourrait indiquer des possibilités d’adhérence de la zone élargie qui pourrait constituer une sorte de ventouse. Les exemplaires juvéniles ne présentent pas ce caractère, même si les gonopodes 8 sont déjà différenciés. Dans les deux sexes, les pattes 1 et 2 portent, sous l’article tarsal terminal, une rangée d’une vingtaine de soies modifiées en bâtonnets aplatis présentant une torsion et munies d’une épine latérale (Figs 10 et 11). Ces soies sont dirigées vers l’arrière et mesurent 30-50 u de longueur. Du côté antérieur de cette rangée, quelques soies modifiées sont placées plus ou moins en ligne. On en trouve également quelques-unes sous le premier article tarsal et sous le tibia; leur longueur atteint ici 100 u. Les soies bordant ces rangées sont légèrement modifiées: elles présentent une épine latérale et sont aussi tordues. Dès la BROELEMANNEUMA GAYI, RÉPARTITION, CSS DA troisième paire de pattes chez les femelles et sur les pattes postérieures aux gonopodes 8 et 9 chez le mâle, les soies sont droites et non modifiées. DISCUSSION Les organismes troglobies sont considérés comme peu mobiles une fois qu'ils se sont adaptés au milieu des grottes. L’éloignement des localités de captures (voir carte et tableau), leur localisation dans des massifs séparés par des vallées de basse altitude (vallée de l’Arve), font qu'il est difficile d'expliquer la répartition de B. gayi. Il n’est certainement pas limité à un seul système souterrain. Cependant, il semble bien que son aire géographique soit restreinte à la région de la Haute-Savoie. Mais la prospection d’autres grottes d’altitude dans les régions voisines (Valais, Savoie) pourrait apporter des éléments nouveaux à cette répartition, qui pourrait se révéler plus large, alpine par exemple. Ses préférences écologiques devront orienter les recherches vers des grottes d’altitude ou des grottes très froides, contenant des dépôts fins riches en matière organique permettant le régime géophage mis en évidence par Bourne (1975b). La localisation de cette espèce dans des endroits sans courants d’air (BOURNE 1975a) est sujette à discussion, puisque la grotte du Calumet présente de forts courants d’air. L'utilisation de pièges contenant de la bière pourrait cependant constituer un appât attirant les diplopodes hors des zones protégées des courants d’air. Cette espèce n’a jamais été décrite à l’état vivant. Les récoltes pouvant parfois être massives, il serait intéressant de la voir en place, pour savoir si elle existe sous forme de populations denses et groupées et si elle présente un éventuel comportement grégaire. La récolte d’un seul individu dans un des pièges et de près de 50 individus dans l’autre piège, distant de quelque 10m, pose la question de la mobilité de cette espèce. De plus, la récolte massive vient d’une zone de cailloux, alors que le piège ayant fourni un seul individu se trouvait dans une zone d’argile, substrat de prédi- lection pour cette espèce selon BOURNE (1975a, b), qui précise les conditions écolo- giques de ses lieux de capture ainsi: “sol argileux, température constance à 2,5°C et absence de courants d’air”. Ces conditions sont celles de la plupart des grottes dites d’altitude, ainsi que des localités mentionnées pour B. gayi, la grotte de la Diau (localité-type de la description de DEMANGE 1968) présentant ces mêmes conditions écologiques malgré une altitude nettement plus basse. Fics 3-11 3: Tête de B. gayi montrant le détail des pièces buccales et l’absence d’ocelles. On peut voir également l’orifice génital 9 (flèche). - 4: Orifice génital 9. - 5: Base des pattes 2 du d montrant les orifices génitaux mâles sous la forme de deux papilles. - 6: Région des gonopodes de B. gayi montrant les pattes 8 fortement modifiées et les pattes 9 de morphologie nettement plus normale. - 7: Gonopodes 8 de B. gayi en vue latérale montrant leur surface antérieure cou- verte de tubercules. Ces gonopodes sont particulièrement larges chez cette espèce. - 8: Extrémité d’une patte antérieure aux gonopodes (P6) chez le 4. L’article précédant la griffe terminale porte à sa face inférieure une série de sortes de petites ventouses. - 9: Détail d’une des ventouses terminales des pattes antérieures du d. - 10: Dernier article des pattes antérieures de la 2 (P2) montrant les deux rangées de soies modifiées. - 11: Poils modifiés de la patte 2 de la 9. Echelle = 400u (Fig. 3), 150u (Fig. 4), 40p (Fig. 5), 200u (Fig. 6), 90u (Fig. 7), 23,5u (Fig. 8), Su (Fig. 9), 115u (Fig. 10), 16u (Fig. 11). 22 PHILIPPE MARTI & JEAN WÜEST En ce qui concerne la morphologie fine de B. gayi, la présence de structures particulières sous les articles terminaux des pattes antérieures constituant un caractère sexuel secondaire n’a jamais été mentionnée ni figurée. Mâles et femelles portent sous les articles terminaux des pattes 1 et 2 des lignes de soies modifiées en bâtonnets tordus, semblables dans les deux sexes. Cette structure pourrait faire penser au peigne de nettoyage des pièces buccales signalé chez les Iulidae sur les tibias et tarses des pattes 1 et 2, formé d’épines ordonnées en ligne (VERHOEFF 1926). Par contre, les mâles présentent des renflements caractéristiques des tarses II des pattes 3 à 7, recouverts de soies élargies en raquettes ou en ventouses. Ces soies modifiées du mâle ressemblent aux soies qu’on trouve à l'extrémité des pattes de nombreux insectes et qui leur servent à adhérer sur des surfaces lisses. On peut penser à des structures (éventuellement associées à des glandes) assurant une bonne adhérence sur le corps d’un partenaire pendant l’accouplement ou a des récepteurs sensoriels, qui pourraient être également impliqués dans l’accouplement. L'examen d’autres représentants du genre Broelemanneuma ou de la famille des Craspedosomatidae devrait permettre de vérifier la présence de ces structures et de l’utiliser éventuellement dans la carac- térisation de ces taxa. REMERCIEMENTS Nous remercions M. Jean-Paul Mauriès, du Muséum national d’ Histoire natu- relle de Paris, qui a bien voulu examiner notre matériel, compléter nos informations et relire le manuscrit. La détermination du Diptère Trichoceridae est due à M. Jean-Paul Haenni, du Musée d’Histoire naturelle de Neuchâtel, auquel va toute notre gratitude. Notre reconnaissance va à M. Villy Aellen, directeur honoraire du Muséum d’histoire naturelle de Genève, et à M. Strinati, pour leurs conseils dans la rédaction de la présente note. Nous remercions M. Slim Chraiti pour le traitement informatique des figures | et 2. BIBLIOGRAPHIE ARIGNANO, D. 1982. Des bétes dans le Jean-Bernard, Samoéns, Haute-Savoie. Echo des Vulcains 42: 2-3. ARIGNANO, D. 1983. Premiers éléments sur la faune du Jean-Bernard. Echo des Vulcains 43: 3-5. BOURNE, J. D. 1973. Biospéléologie. Hypogées 30: 10. BOURNE, J. D. 1975a. Notes écologiques sur la grotte de la Barme Froide (alt. 2000m.) et sur les environs. Hypogées 36: 45-58. BOURNE, J. D. 1975b. Premières constatations sur quelques facteurs écologiques pour les diplopodes troglobies Brohlemanneuma gayi Dem. Actes 5° Congrès National Suisse de Spéléologie, Stalactite suppl. 9: 23-26. DEMANGE, J.-M. 1968. Un nouveau Myriapode de Haute-Savoie: Brölemanneuma gyai nov. sp. (Diplopoda: Craspedosomoidea: Craspedosomidae). Annales de Spéléologie 23: 189- 190. DEMANGE, J.-M. 1970. Sur une collection de Myriapodes de France (Savoie, Haute-Savoie, Ardèche) rassemblée par par M. L. Deharveng. Bulletin du Muséum national d Histoire naturelle, Paris, 2° serie, 42: 502-508. DEMANGE, J.-M. 1981. Les Mille-pattes. Ed. Boubée, Paris, 284 pp. BROELEMANNEUMA GAYI, RÉPARTITION, CSS 23 Favre, D. 1994. Camp d'été 93. Hypogees 61: 10-11. GEOFFROY, J.-J. 1983. Broelemanneuma gravi est-il le mille-pattes le plus profond du monde? Bulletin de liaison, Société de Biospéléologie 4: 25-26. GEOFFROY, J.-J. & J.-P. MAURIES. 1992. Diplopodes cavernicoles, édaphiques et souterrains de France. Données récentes, répartition des espèces nouvellement décrites et peu connues. Mémoires de Biospéléologie 19: 127-133. MEYSONNIER, M. 1984. Observations fortuites de chauves-souris. en 1983. S.C.V. Activités 45: 29-30. MEYSSONNIER, M., V. AELLEN & P. STRINATI. 1987. Faune souterraine du département de la Haute-Savoie. Emergences Spéléos Rhône-Alpes, N° spécial 1. 120 pp. VERHOEFF, K. W. 1905. Über «Ceratosoma» pectiniger Bròl. Zoologische Anzeiger 29: 223- 224. VERHOEFF, K. W. 1926. Diplopoda. Bronn’s Tierreich. Vol. 5, Fasc. 2, lère partie, 1071 pp. Leipzig. REVUE SUISSE DE ZOOLOGIE 105 (1): 25-48; mars 1998 A revision of the genus Munatia Stal, 1875 (Orthoptera, Caelifera, Romaleidae, Romaleinae). C. Hugh F. ROWELL Zoologisches Institut der Universitàt Basel, Rheinsprung 9, 4051 Basel, Switzerland. A revision of the genus Munatia Stàl, 1875 (Orthoptera, Caelifera, Romaleidae, Romaleinae). - The genus is redescribed. Keys to the genera of the Procolpini and to the species of Munatia are given. M. punctata Stàl and M. biolleyi Carl are both redescribed, and a female allotype of the former designated. M. decorata Carl is synonomized with M. punctata. Lectotypes of both M. decorata Carl and M. biolleyi Carl are designated. Biological data on both valid species are provided. The younger larvae of at least M. punctata are gregarious and provided with a visually striking pattern. Key-words: Orthoptera - Acridoidea - Romaleidae - Procolpini - taxo- nomy. INTRODUCTION The genus Munatia was created by STAL (1875) with the new species M. punctata as the type. The same author later (1878) contrasted Munatia with his (1873) genus Procolpia, using the shape of the tips of the elytra and the prominence of the medial spine of the hind knee as distinguishing characters. REHN (1955) revised the genus Procolpia; he rejected Stal's discriminating characters while clarifying the distinction between the two genera on the basis of others, but did not revise or redescribe Munatia. REHN & GRANT (1959) included Munatia (along with Procolpia Stal, Aeolacris Scudder, Prorhachis Scudder and Xomacris Rehn) in a tribe Pro- colpini of the Romaleinae (a somewhat reduced successor to GIGLIO-Tos's (1898) group Procolpiae), which probably corresponds to a real clade. Three further species of Munatia have been described since 1875. Munatia australis Bruner (1906) from Paraguay was synonomized with Procolpia minor Giglio-Tos, 1894 by REHN (1955). CARL (1916) erected two new species (M. biolleyi and M. decorata) on the basis of Costa Rican material, without discussing the characters of the genus. He effectively ignored Stal's punctata, noting only that it was too briefly described to provide a basis for comparison and in any case did not agree Manuscript accepted 10.07.1997. DE C. HUGH F. ROWELL with his biolleyi, and he did not examine the type specimen. Additionally, Procolpia inclarata (Walker) was treated by BRUNER (1907) in his text as P. emarginata (Serville) but figured as Munatia inclarata. Specimens of Munatia from Costa Rica (Tucurrique, Turrialba, Juan Vifias and Carrillo) were referred, in part erroneously, to punctata Stàl by REHN (1904) and BRUNER (1907), but since 1916 CARL's name decorata has mostly been used for Costa Rican material which does not agree with the description of M. biolleyi. REHN & GRANT (1959) examined and figured the male genitalia of Munatia decorata Carl but did not examine any other species of the genus. Several orthopterists have indicated to me that they consider it likely that all three current species of Munatia represent the same taxon. With access to plentiful material and much field experience of the Costa Rican species I here show that decorata Carl is indeed a synonym of punctata Stàl, but that biolleyi Carl is a valid species. In view of the inadequacy of previous descriptions, I also redescribe the genus and its two species and designate an allotype female of M. punctata Stàl and lectotypes of both M. decorata Carl and M. biolleyi Carl. Abbreviations of depositories: ANSP, Academy of Natural Sciences, Philadelphia, USA; INBio, Instituto Nacional de Biodiversidad, Santo Domingo de Heredia, Costa Rica; MNHNP, Muséum National d'histoire naturelle, Paris, France; MHNG, Muséum d'histoire naturelle, Geneva, Switzerland; NRS, Naturhistoriska Rijksmuseum, Stockholm, Sweden; RC, the author's collection; UCR, Museo de Entomologia, Universidad de Costa Rica, San José, Costa Rica; UMMZ, University of Michigan Museum of Zoology, Ann Arbor, USA. KEY TO GENERA OF PROCOLPINI: l Medial carina of pronotum absent, lateral carinae present and decorated with granular points. At least the more proximal spines of hind tibia conspicuously flattened and dorsoventrally produced at their base. Male Withtlarsepale Spots On'el VIT ON re en ORI Aeolacris Scudder - Medial carina of pronotum present, lateral carinae absent. ............... 2 2 Medial carina straight or simply arcuate, not incised by sulci. Elytra n'arrowAwithmo!costalilobe 8 TRO. SONATA e AREE Munatia Stàl - Medial’ carina incised by ‘sulci, costal lobe present. 2.1... Spree 3 3} Medial carina not produced dorsally to form large teeth. Lateral lobe not bearing a conspicuous lateral tubercle or spine. .......... Procolpia Stàl - Medial carina divided into teeth in prozona; lateral lobe bearing a lateral 'Spine’or tubércle} 1... 17... MR EN E AR 4 4. Medial carina anterior to first sulcus without prominent raised tooth. ateralimargins oftastieiumspined 45) 55455-5400" Prorhachis Scudder - Medial carina of pronotum with three large teeth in prozona, the first one anterior to the first sulcus. Lateral margins of fastigium simple. DIO Sera nea ea Xomacris Rehn REVISION OF MUNATIA 27 Munatia punctata O' IN Fic. 1 M. punctata. Male, lateral view. Intersegmental membranes at base of legs are shown in black for clarity, but are pale brown in life. Munatia Stal, 1875 STAL 1875: 28 Type species: M. punctata Stal, 1875. REHN 1905: 404; BRUNER 1907: 223; KIRBY 1910: 366; REHN 1955b: 37-39; REHN & GRANT 1959: 239; Uvarov & DIRSH 1961: 158; AMEDEGNATO 1974: 198. REDESCRIPTION OF GENUS Displays the characters delimiting the tribe Procolpini as defined by REHN & GRANT (1959). The most obvious of these are generally elongate shape, ensiform antennae, well developed rostrum, fully developed wings and tympana, smoothly elongated male subgenital plate, and the inequality between the external and internal row of the hind tibial spines, the latter being notably long and curved. Additionally characterised and distinguished from the remaining genera of the tribe as follows (see also Key to Genera above): Rostrum somewhat rounded in profile. Frontal ridge below medial ocellus absent (male) or obsolete (female) (vide Procolpia and Xomacris). Infra-ocular carinae weak or obsolete. Fastigium smooth-sided, without lateral processes (vide Prorhachis). Lateral carinae of pronotum absent (vide Aeolacris), medial carina well marked, in lateral view straight or forming a low curving crest, not incised by transverse sulci (vide Procolpia, Prorhachis, Xomacris), and not markedly higher in 28 C. HUGH F. ROWELL prozona (vide Xomacris and Prorhachis) than elsewhere; anterior margin of pronotum only weakly notched medially. Elytra long and slender with narrow rounded tips (also true of some species of the other genera), and with no proximal lobe on the costal margin. Wing dominated by the anal region; radial and medial areas strongly reduced, especially in the male, in which the remigium forms only a narrow elongate strip at the leading edge, somewhat or markedly longer than the anal area. Alar stridulatory apparatus very reduced or absent, transverse veins of the first anal area obsolete or incomplete and usually without denticles. Fenestration of the 2nd alar area absent (vide Aeolacris). Terminal medial tooth of metathoracic knee small in the adult (vide most other members of the tribe). Marginal spines of hind tibiae always simple and of circular cross-section, never laterally flattened and dorsoventrally produced at base (vide Aeolacris); 8-10 external and 8-9 internal spines on hind tibia, the bottom 3 and the upper 1-2 internal spines short, the remainder long and curved towards the animal's midline. Abdominal segments with well-marked medial carina. Male furcula simple, weak. Male supra-anal plate triangular, rounded at tip, simple, with a proxi- mal medial longitudinal furrow bordered by melanized edges. Male subgenital plate biolleyi 5 mm FIG. 2 Frontal view of head of males of Munatia. In biolleyi the eyes are more produced dorso- ventrally and the facial ridges more pronounced than in punctata. REVISION OF MUNATIA 29 twice as long as supra-anal plate (vide Procolpia, Prorhachis, Xomacris). Aedeagal valves with a weak subapical, latero-posteriorly directed process, and with weak transverse ridges on dorso-lateral surfaces of their tips (very similar to that of Xomacris (AMÉDÉGNATO & POULAIN 1986: Fig. 90)). Epiphallus with large pointed lophi. Distribution: Panama, Costa Rica, and Nicaragua. KEY TO SPECIES OF Munatia: 1) Males. Lateral lobe of pronotum brown, with 2 pairs of circular yellow or gold spots, not touching the ventral margin of lobe. Frons, genae, pronotum, tho- racic pleura and outer face of hind femur devoid of black tubercles. All longitudinal veins of elytron unbranched, except for the radius (Fig. 5); elytron with green or yellow margins along proximal regions of both leadinsfanditrailinszedgesun Secs ss RE punctata Stal Lateral lobe of pronotum not as above, yellow or green areas extend to ventral margin and are often fused. Frons, genae, pronotum, thoracic pleura and outer face of hind femur with numerous small black tubercles. Radius, media and cubitus veins of elytron branched (Fig. 5); elytron with green or yellow margin along proximal region of trailing edge OT 3.75 NG ONZE a ee biolleyi Carl 2) Females. Medial carina of pronotum low and straight, not at all arcuate (Fig 4A). Distal medial surface of subgenital plate smooth and convex (Fig. 8B). Black tubercles absent and venation of elytron unbranched, as in male. . punctata Stal Medial carina of pronotum raised and clearly arcuate (Fig 4C). Distal medial surface of subgenital plate bearing two minutely toothed ridges, separated by a deep medial groove (Fig. 8D). Black tubercles present and longitudinal elytral veins branched, as in male............. biolleyi Carl 1. Munatia punctata Stal, 1875 Munatia punctata Stal, 1875: 28. Holotype male, Chiriqui, Panama (Boucard), no date, Naturhistoriska Rijksmuseum, Stockholm (examined). PICTET & SAUSSURE 1887: 340.; REHN 1905: 405 (misidentification of M. biolleyi, as shown by figure of wing outlines); BRUNER 1907: 223 (in part misidentification of M. biolleyi, specimens examined). Allotype female. Female, here designated. Costa Rica, Centr. Am. (P. Biolley), ANS Philadelphia. No other data; bears additional labels "Munatia punctata Stal", "Hebard Collection", "M. decorata Carl", "Munatia punctata Stal 1875 det. C.H.F. Rowell, 96041". Munatia decorata Carl, 1916: 506, Fig. 12. Lectotype male, here designated (selected and labelled "Hololectotype" by C.S. CARBONELL 1966), Costa Rica (Prov. Cartago), Carrillo, 600 m; (Prov. Alajuela) Sarapiqui, Carablanco, 600 m (P. Biolley); paralectotype male (labelled "paratype" by C.S. CARBONELL 1966), no data, but considered by Carl to come from the other of the two localities indicated by Biolley on the label on lectotype male; both Muséum d'histoire naturelle, Geneva (examined). Syn. n. 30 C. HUGH F. ROWELL ia pera O 7 0 punctata EEE, A FT o) à 3 o | Fic. 3 Side view of head in Munatia. The rostrum is slightly longer and more pointed in punctata. In biolleyi there are also numerous small black tubercles on frons and genae (not shown). REVISION OF MUNATIA 31 biolleyi Fic. 4. Pronota of Munatia. In both sexes biolleyi is distinguished by having more numerous, smaller bosses along the anterior margin, the presence of a single raised pale tubercle laterally just anterior to the third sulcus (present but obscure in punctata), and numerous small black tubercles. Female biolleyi are additionally distinguished by the presence of a higher, arcuate crest, and males by a different distribution of pigment (dotted lines). 32 C. HUGH F. ROWELL REDESCRIPTION Stal's description of punctata is very brief and based on a poorly preserved specimen (see below), and is not illustrated. BRUNER (1907) did not figure the genus. Carl's description of decorata is largely limited to a comparison with biolleyi, and includes no figures apart from an outline of the wings. The female has not been described. Male (Fig. 1). Large (40 mm < body size < 50 mm). Integument matte and finely pitted, especially on head and thorax. Head. Fastigium triangular, rounded at tip, longer than vertex, forming a rostrum, dorsal surface convex. Frontal ridge (Fig. 2) very narrow dorsally, widening beneath antennal sockets, extending to the medial ocellus and there slightly grooved. Profile of frons (Fig. 3) concave, culminating in rostrum. Eyes globose, prominent, vertical dimension 1.3X horizontal dimension. Interocular space large, more than twice width of antennal scape. Antennae ensiform, longer than head and pronotum, 21 segments in flagellum. Thorax. Median carina of pronotum (Fig. 4) well developed, straight, not incised by the three transverse sulci. Anterior margin of pronotum shortly produced in midline, with a small medial notch. Anterior edges of pronotal lobes with 8-10 pairs of smooth raised bosses, anterior edge of prothoracic episternum with 2-3 such bosses. Posterior margin of pronotum produced posteriorly to a triangular point, forming a 70° angle. Prosternal process long, slender, vertical, tapering, rounded at tip. Mesosternum narrower than metasternum; metasternal interspace wider than long (Fig. 6C). Elytron (Fig. 5) long, narrow, with fine rounded tip, projecting well beyond hind knee. RI, CUI and CU2 unbranched. Wing long and narrow with 8 anal veins; stridulatory area of wing (area anterior to 2A) with very reduced transverse veinlets, entirely without denticles, presumably non-functional. Hind femora long, slender, exceeding length of abdomen, dorsal and ventral medial carinae slightly toothed, dorsal carina terminating in a minute apical spine; outer medial area of hind femur with reticulate pattern. Hind tibia with 8-10, usually 9, external spines and 9 internal spines. Abdominal segments with well-marked medial carina, produced in 9th seg- ment into a boss slightly overhanging supra-anal plate (Fig. 6A). Furcula present but weakly developed (Fig. 6A). Cerci (Figs. 6A, B) simple, short tapering, rounded at tips. Supra-anal plate (Fig. 6A) as in generic description. Subgenital plate elongate, subcylindrical, tapering, rounded at posterior tip, twice as long as supra-anal plate (Fig. 6A, B). Epiphallus (Fig. 7 A-C) bridge-shaped, with a medial dorsal protu- berance; lophi large, vertical, with outwardly directed tips, ancorae small. Lateral epi- phallic sclerites present. Cingulum (Fig. 7D-E) simple, saddle shaped, without anterior apodemes. Anterior apodemes of endophallus laterally flattened in form of two vertical concave plates, joined dorsally by a thin transparent chitinous plate (Fig. 7G, K). Aedeagal sclerites as in generic description. REVISION OF MUNATIA 33 punctata == o = CUT CU2 CUIB CUIA MP MA zen, C SC CUT CU2 R M MA MP Ri RS biolleyi | biolleyi = FIG. 5 A. Elytra and wings of female and male M. punctata and M. biolleyi. In punctata both the elytron and the wing are relatively narrower than in biolleyi, and this is associated with a reduction in the branching pattern of the longitudinal nerves of the elytron (especially in the male) and in the number of anal veins in the wing. Nomenclature of the venation and abbreviations after RAGGE (1955). 34 C. HUGH F. ROWELL Coloration. Antennae blackish brown. Head green; eyes brown; postocular stripe brown, extending also anteriorly around and under rostrum to form a brown horizontal band at eye level. Clypeus, labrum and mandibles brown. Maxillary and labial palps green. Pronotum brown, with a broad green medial stripe; two large yellow spots on pronotal lobe, one anterior and one posterior to the second transverse sulcus (Fig. 1, Fig. 3). Meso- and metathorax brown; meso- and metasterna each with a large yellow spot. Legs green, tibial spines tipped with black, ventral surfaces of tarsi black. Semilunar processes of hind knee dark brown. Abdomen brown, often with a horizontal yellow stripe along the ventral half of the abdominal tergites. Elytron brown, the marginal regions anterior to RS and posterior to 1A green. Wing pale yellow, with a broad smudged black border, widening anteriorly. FIG. 6 M. punctata. A, B, extremity of male abdomen, A, dorsal view, B, lateral view. C. Male thoracic sternal plate. REVISION OF MUNATIA 35) Female. Gigantic (70 mm < body size < 80 mm). Pronotum as in Key to Species and Fig. 4, with low, straight, medial carina. Ovipositor valves (Fig 8B, C) robust, long, straight, in side view slightly hooked distally, outer margins melanized but without teeth. Subgenital plate (Fig 8A) smooth and convex in its distal medial part. Spermatheca not examined. Coloration: all females seen to date have been plain green on head, thorax, wings and abdomen; no brown forms are known. Antennae, eyes, legs, hind knees, tarsal spines as in male. Measurements: see Table 1. The values for Stàl's Panamanian type (measured in this study) fall comfortably within the maximum and minimum values of modern Costa Rican specimens. The same is true of the values given by Carl for his M. decorata, given that his F = 17 mm is a misprint for 27 mm (specimen examined). Using the dimension P (length of the pronotum in the midline) as a reference point, females have relatively shorter hind femora and tarsi, a thinner antennal pedicel and slightly shorter elytra than the males. Larvae (Fig 9A). Larvae are laterally compressed with a prominent pronotal crest and a well developed spine on the hind knee. With successive moults these two characters reduce to the adult condition. Young larvae are dark brown, marked with gold or orange on the antennae and as a conspicuous patch above the tympanum, and more variably behind the eye and on the hind femora. Females have lost all orange markings by the third instar and are thereafter uniformly brown (or occasionally green) until the final moult. Males retain the supratympanal orange spot until the end of the third instar; in the fourth instar they adopt the adult coloration with the four additional pairs of golden spots on the pronotum and thoracic episterna. In the fourth, fifth and adult instars the location of the supratympanal spot is covered by the wings or wing rudiments and the pigmentation is absent. TAXONOMIC DISCUSSION Stal's type, the only specimen to date from Panama, is indistinguishable from the Costa Rican material. It was clearly a newly moulted adult, dried slowly under damp tropical conditions - the author has collected similarly poorly preserved speci- mens which show exactly the same facies of shrunken and wrinkled abdomen and a colour reversal of the characteristic spots on the thorax, which are now artifactually darker, rather than lighter, than their surroundings. This reversal is the origin of Stal's description "lateribus thoracis maculis quattuor nigricantibus notatis", which in turn apparently convinced Carl that his material was different from that of Stal ("elle (M. punctata) semble différer considérablement de M. biolleyi, notamment par la présence de 4 taches noiratres sur les cotés du thorax du male"). Stal had no female specimen, and there is no female specimen present in Carl's type series. For this reason an allotype female is designated. It is from the same collector and quite possibly the same locality as the lectotype of M. decorata. MATERIAL EXAMINED. Type material of punctata Stal and decorata Carl as indicated above. Additionally: 36 Munatia punctata Males Dimensions in millimetres: Hind femur (F) Rostrum-subgen. plate (L) Pronotum (midline) (P) Pronotum longest Interocular space (10) Antennal pedicel (width) Antenna (A) Antenna -> 11th. segment Hind tarsus Ist + 2nd segments Hind tarsus 3rd segment *Elytron length (E) Rostrum, tip to eye Ratios F/P L/P IO/P IO/pedicel Tarsus 3/ Tarsus 1+2 Tarsus 1+2+3/F Tarsus 1+2+3/P AIP SEP *E/L Females Dimensions in millimetres: Hind femur (F) Rostrum-subgen. plate (L) Pronotum (midline) (P) Pronotum longest Interocular space (10) Antennal pedicel (width) Antenna (A) Antenna -> | 1th. segment Hind tarsus Ist + 2nd segments Hind tarsus 3rd segment *Elytron length (E) Rostrum, tip to eye IO/P IO/pedicel Tarsus 3/ Tarsus 1+2 Tarsus 1+2+3/F Tarsus 1+2+3/P A/P *E/P *E/L C. HUGH F. ROWELL Mean 28.45 47.22 13.07 13.16 222 1.26 2923 19.31 3.62 3:95 47.21 3.64 S.D. 2235 TABLE 1. Dimensions of M. punctata Max Min B & D D D D PB BE BR BR $ & D D D D BR R R R Stal's Carl's type values ei CNT 41.00 45 MESSE 1893 1.90 1.20 1025 3.89 309) 44.50 50 357 * In many specimens the elytron tip is broken. Only the maximal values are meaningful. REVISION OF MUNATIA 27 Say O biolleyi Fic. 7 Male genitalia. A-G, M. punctata; H-M. M. biolleyi. A-C, H-J, epiphallus, dorsal, axial and lateral views; D, dorsal view of posterior region of complete phallic complex (area indicated by arrow in E); E. Complete phallic complex, lateral view; F-G, L-M, endophallus, lateral and dorsal views; K, diagrammatic transverse section through M at the point indicated, to show relationships of anterior apodemes of endophallus, the dorsal plate joining them, and the ejaculatory duct. Scale bar 1 mm, except for D, E and K, not to scale. 38 C. HUGH F. ROWELL COSTA RICA: Prov. Alajuela: Sarapiqui, Cariblanco, 700 m, April 20, 1977 (DeVriess P), INBio, no. CRIOO1 0130581, III instar female; August 28, 1981 (Simons Y), UCR, 1 male. Sarapiqui, Cinchona waterfall, 1470 m, August 24, 1983 (Rowell CHF), RC, no. 83407, 1 male. Sarapiqui, nr. Virgen del Socorro, 800-1000 m, June 21, 1980 (Rowell CHF, Rowell-Rahier M, Hyde C), RC, no. 80110, | male; June 22, 1980, no. 80132a, 1 male; 80132b & c, 2 larvae; nos. 80115a, 1 male, 80115b, 80115c, 2 larvae. Prov. Cartago: Orosi, September 9, 1982 (Marin F), UCR, 1 male. Orosi, Embalse El Llano, January 22, 1981 (Alvarado A), UCR, 1 male. Aquiares, nr. Turrialba, March 17, 1930 (Lankester CH), ANSP, 1 male. 13 km by rd. NW Turrialba (0.7 km NW Santa Cruz), site #131 (same locality as prevous one), October 1, 1961 (Hubbell TH, Cantrall I, Cohn T), UMMZ, 1 female, 4 II instar larvae. Santa Cruz, crossing of R. Aquiares & rd., 0.7 km NW of church, 1475 m (same locality as previous one), July 6, 1980 (Rowell CHF, Rowell-Rahier M, Hyde C), RC, nos. 80192a, 80192b, 2 larvae; July 10, 1980 (Rowell CHF Rowell-Rahier M Hyde C) RC, no. 80213, 1 male. Tapanti, Ref. Nac. Fauna Silv., Quebrada Segunda, 1250 m, March 1992 (Mora G), INBio, no. CRIOOO 741239, 1 male; no. CRIOO! 964248, fragmentary larva I or II, female; July 20, 1985 (Solis A), INBio, nos. CRIOOI 013059, CRIOOI 013060, CRIOOI 013056, CRI001 013062, CRIOO1 013057, 5 larvae HI female; no. CRIOO1 013408, 1 larva IV male. Prov. Guanacaste: Guanacaste (no other data), March 4, 1972 (Acevedo A), RC, no. 76001, 1 male. Sta. Cecilia, 9 km S, Est. Pitilla, 700 m, August 1988 (GNP Biodiversity Survey), INBio, no._ CRIOOI 014106, 1 male. Tierras Morenas, Bajo Los Cartagos, R. San Lorenzo, 1050 m,. April 1991 (Alvarado C), INBio, no. CRI000 463114, 1 male; April 1992 (Quesada F), INBio, no. CRI000 772687, 1 female. S.E. slope of Volcan Cacao, 1200 m, July 24, 1991 (Rowell CHF, Elsner N, Chavez C), RC, no. 91149, 1 larva. Volcan Cacao, Estac. Cacao, 1000-1400 m, SW side, April 1988 (Espinoza M), INBio, no. CRIOOO 036280, 1 larva III female; October 1989 (Blanco R, Chaves C), INBio, no. CRIOOO 097917, 1 male; December 1, 1989 (Blanco R, Chaves C), INBio, no. CRI000 204475, I male. Volcan Cacao, Estac. Mengo, 1100m, SW side February 1989 (GNP Biodiversity Survey), INBio, no. CRIOO! 014107, 1 male. Volcan Tenorio: nr. summit of rd. from Tierras Morenas to Bajo Los Cartagos, 1040 m, July 21, 1991 (Rowell CHF, Elsner N), RC, no. 91097, 1 larva II male. Prov. Puntarenas: Monteverde, Cerro Amigos, 1840 m, August 29, 1993 (Zumbado MA), INBio, no. CRIOO! 973681, 1 male. Monteverde, Est. La Casona, 1520 m, May 1991 (Obando N), INBio, no. CRIOO1 326073, 1 larva HI; March 1992 (Flores K), INBio, no. CRIOOO 788757, 1 larva. April 1992 (Flores K), INBio, no. CRIOOO 990325, 1 larva II female. Monteverde, San Luis, 1040 m, January 1993 (Fuentes Z), INBio, no. CRIOOI 371007, 1 male. Prov. S. José: 14 km N. of S. Isidro General on Pan-American Hwy, subtropical wet forest. July 19, 1961 (Futuyma D), UMMZ, 1 larva III male. La Hondura, 1300 m, May 18, 1929 (Valerio M), ANSP, 1 male. Bajo La Hondura. 900 m, May 1971 (Echeverria L), UCR, | male; August 9, 1978 (K. Paulsen), UCR, 1 male. Carrillo, June 1903, ANSP, 1 male, 4 male larvae, 1 female larva (Hebard Collection); August 1903, ANSP, I male; August-October 1903, ANSP, 1 male. Parque Nacional Braulio-Carrillo, La Montura, 1100 m, April 26, 1980 (DeVriess P), RC, nos. 80282a, 80282b, 2 larvae. Pozo Azul de Pirris, 325-550 ft (98-167 m) May-June 1903, ANSP, 1 larva. Distribution. M. punctata is a characteristic and sometimes common species of lower montane rain forest in Costa Rica, replacing M. biolleyi as one ascends from the lowlands. Virtually all records come from between 600 and 1600 m altitude. The only certain record from a lower altitude is a single larva from Pozo Azul de Pirris, 98-167 m, in 1903. The type of punctata from Chiriqui Province (Pacific slope, abutting the REVISION OF MUNATIA 39 FIG. 8 Female genitalia. A-C, M. punctata; D-G, M. biolleyi. À, D, ventral view of subgenital plate and ventral ovipositor valves; B, E, dorsal view of supra-anal plate and dorsal ovipositor valves; C, F & G, lateral view. 40 C. HUGH F. ROWELL Costa Rican border) is the only recorded example from Panama. It is not so far recorded north of Costa Rica, and is absent from Astacio-Cabrera's (1975) com- pilation of species from southern Nicaragua. Distribution map, Fig. 10. NATURAL HISTORY Males are active and conspicuous with their shining yellow spots, females sluggish, cryptically coloured and more rarely collected. Larvae are recorded from April to October; adults are first seen in July, are common from August to November, and occur as isolated individuals through January and March. These data are compa- tible with a one-year generation time, the eggs being laid in the (North Temperate Zone) Autumn and hatching with the onset of the Spring rains in March/April. The larvae early aggregate into groups. These groups can contain more than 100 individuals of mixed ages, which suggests that the progeny of more than one egg pod may coalesce. They are visually attracted to each other, collecting by a proffered mirror, as in the related romaleine Chromacris, or the African pyrgomorphid genus Phymateus (ROWELL, unpublished data and 1967). The early larvae are reddish brown in colour and conspicuously marked with two bright yellow patches. They are strongly compressed laterally, hold the hind legs flexed high over their backs, knees together and feet off the substrate, and often lie on their sides - and so look at first sight like anything but grasshoppers. These groups of young larvae bear a striking superficial resemblence to those of a similarly coloured and equally gregarious redu- vid bug which occurs in the same environment, and although no experimental data are available, it seems likely that there is a mimetic relationship between the two. Later instars too tend to stay in groups at first but occur as solitary individuals by the fifth instar - presumably (as in Phymateus) dispersal is caused by loss of visual contact due to the more cryptic coloration. M. punctata is moderately polyphagous, eating the leaves of a variety of trees, shrubs and herbs, including Hyptis (Lamiaceae), Lantana (Verbenaceae), Croton and Alchornia (Euphorbiaceae) and Clibadium and Vernonia (Asteraceae). Most larval groups have been found on Lantana or Clibadium. All of these plants are strongly odorous or otherwise chemically defended. It has not been seen to feed on monocots, either in the wild or when offered them in captivity. It also refuses many dicoty- ledenous leaves, including those of species of Solanum (Solanaceae), Hibiscus (Mal- vaceae), Phenax (Urticaceae), Conyza and Erechtites (Asteraceae). Munatia biolleyi Carl, 1916 Munatia biolleyi Carl, 1916: 504, lectotype male, here designated (selected and labelled "hololectotype" by C.S. Carbonell, 1966), no collection data: paralectotype male, Costa Rica (Prov. Cartago), Carrillo, herbes aux soleil (P. Biolley); paralectotype female (selected and labelled "allolectotype" by C.S. Carbonell, 1966), Costa Rica (label in Biolley's hand writing), no other collection data; all Muséum d'histoire naturelle, Geneva (examined). It is highly probable that all of Carl's type series were collected in Carrillo by Biolley. REHN 1905: 405 (as M. punctata, but figured outline of wing indicates actually biolleyi); HEBARD 1924a: 100. REVISION OF MUNATIA 4] REDESCRIPTION Male. Very similar to M. punctata, from which it differs a) as specified in the Key to Species above and b) as follows: Frontal ridge (Fig. 2) shorter than in punctata, not reaching the medial ocellus. Preocular ridges present, frons in general with more sculpturing than in punctata. Eyes longer in vertical dimension (1.45 X the horizontal dimension). Rostrum (Fig. 3) more rounded in profile. Pronotum (Fig. 4) bears only 4-6 pairs of bosses on the anterior margins of the lateral lobes, and these are smaller than in punctata. The prothoracic episternum 1s rounded, with no decoration on anterior margin. A prominent white tubercle present anterior to the third transverse sulcus on the side of the pronotum (also present in punctata, but there much smaller and darker). Both elytron and wing (Fig. 5) broader than in punctata; CUI of the elytron is branched, and the wing has 10 anal veins. Stridulatory area of wing slightly better developed than in punctata, but probably still nonfunctional. Hind tibia with 8-9, usually 8, external and internal spines. There are no consistent differences in the internal or external genitalia of the two species (Figs. 6, 7). Coloration. The dorsal medial stripe running along fastigium, vertex, and pronotum, and continued along the folded anal region of the elytron, can be pale brown, yellow, or green, and is narrower than in punctata - in the latter species it is apparently always green. The same difference in range of coloration applies to the legs. Pattern on pronotal lobes as in Key to Species. The leading edge of the elytron is invariably brown, lacking the anterior green stripe of punctata. Female. Differs from female of punctata as described in Key to Species, namely in the clearly arcuate crest of the pronotum (Fig. 4) and the medially grooved extremity of the subgenital plate (Fig 8D). Ovipositor valves (Fig. SE-G) shorter and in dorsal view more divergent than in punctata, otherwise similar in form. Sperma- theca not examined. Coloration. Unlike punctata, the females of biolleyi can be either brown or green in general coloration, usually the former. The trailing margin of the wing (i.e. the anal area) is usually yellow or a lighter brown, thus forming a dorsal pale strip when the wings are folded. The fastigium and vertex are also pale in brown forms, but there is no medial pale stripe on the pronotum as in the males. Measurements: see Table 2. Carl's values fall within the range of measure- ments made in this study, allowing for the fact that his F = 29 mm (male) is an over- estimate (specimen examined). The same sexual differences in relative proportions are seen as in punctata. Comparing the two species, using ratios relative to the dimension P as a reference point, punctata has relatively longer hind femora and elytra than biolleyi. Larvae (Fig 9B.) The morphological changes occuring in the larvae are similar to those described for punctata. Only the first instar however has orange markings, all others are cryptically coloured. MATERIAL EXAMINED. Type material of biolleyi Carl as indicated above. Additionally: 42 Munatia biolleyi Males: Dimensions in millimetres: Hind femur (F) Rostrum-subgen. plate (L) Pronotum (midline) (P) Pronotum longest Interocular space (10) Antennal pedicel (width) Antenna (A) Hind tarsus 1st + 2nd segments Hind tarsus 3rd segment Elytron length (E) Rostrum, tip to eye IO/P IO/pedicel Tarsus 3/ Tarsus 1+2 Tarsus 1+2+3/F Tarsus 1+2+3/P A/P E/P E/L Females: Dimensions in millimetres: Hind femur (F) Rostrum-subgen. plate (L) Pronotum (midline) (P) Pronotum longest Interocular space (IO) Antennal pedicel (width) Antenna (A) Hind tarsus Ist + 2nd segments Hind tarsus 3rd segment Elytron length (E) Rostrum, tip to eye IO/P IO/pedicel Tarsus 3/ Tarsus 1+2 Tarsus 1+2+3/F Tarsus 1+2+3/P A/P E/P E/L Mean C. HUGH F. ROWELL SD: TABLE 2. Dimensions of M. biolleyi OO Un \O OO 0 0 0 \O 46 35 20 60 REVISION OF MUNATIA 43 PANAMA Prov. Bocas del Toro: Bocas del Toro, July 1-10, 1908 (Robinson W), ANSP, 1 male. CosTa RICA Prov. Cartago: 2 mi SE Turrialba (grounds of Inst. Interamer. de Sci. Agricolas), site #129, September 30, 1961 (Hubbell T, Cantrall I, Cohn T), UMMZ, 2 larvae IH, I male; October 3, 1961 (Hubbell T, Cantrall I, Cohn T), UMMZ, 2 females,1 male. Juan Vinas, March (no year given) (Bruner L), ANSP, 1 female. Prov. Guanacaste: Cerro El Hacha, 300 m, 12 km SE La Cruz, May 1988 (Espinoza M), INBio, no. CRIOOO 094299, | female. Sta. Cecilia, 9 km S, Est. Pitilla, 700 m, May 1988 (GNP Biodiversity Survey), INBio, no. CRIOOO 121179, 1 larva V female; July 1988, no. CRIOO1 014100, I larva V male; no. CRI001 014089, 1 male; no. CRIOOO 129732, 1 male; no. CRIOOO 129731, 1 male; no. CRIOOO 129446, 1 male; no. CRIOOO 088000, 1 male; no. CRIOOO 129797, 1 female; no. CRIOOO 088007, 1 female; no. CRIOOI 014121, 1 larva; August 1988, no. CRIOOI 014105, 1 larva III f.; no. CRIOO1 014090, 1 male; no. CRIOOI 014117, 1 female; September 1988, no. CRIOO1 013295, 1 male; November 1988, no. CRIOO1 014113, 1 larva V female; no. CRIOOO 136257, 1 male; April 6, 1989, no. CRIO00 091743, 1 female; June 1989, no. CRIOOO 011236, 1 male; September 1989, no. CRIO00 035769, 1 male; September 1989 (Moraga C, Rios P), INBio, no. CRIO00 046485, 1 male; May 1990 (II Curso Parataxon.), INBio, no. CRIOO1 147678, 1 larva IV male, no. CRIOOO 241688, 1 larva V male.; no. CRIOOO 293438, 1 female; August 16, 1991 (Moraga C), INBio, no. CRIO00 409548, 1 female; July 21, 1993 (Rios P), INBio, no. CRIOO! 767206, 1 male; December 10, 1993 (Moraga C), INBio, no. CRIOO1 948084, 1 female. Prov. Heredia: Parque Nacional Braulio-Carrillo, Est. Magsasay, June 1990 (Alvarado C), INBio, no. CRI000 272954, 1 female; Puerto Viejo, Finca La Selva. 40 m, September 3, 1975 (Rowell CHE), RC, nos. 75005 & 75006 | male, 1 female, in cop.; August 13, 1976 (Walz S, Rowell CHF), RC , nos. 76003 & 76004, 1 male, 1 female, in cop.; September 12, 1979 (Rowell CHF, Rowell-Rahier M), RC, no. 79246, | male; September 13 1979 (Rowell CHF, Rowell-Rahier M), RC, no. 79247, 1 male; July 10, 1980 (Braker HE), RC, no. 80018, 1 female; May 3, 1982 (Marquis B), ANSP, no. 82-106, 1 female; June 1982 (Braker HE), ANSP, no. 82-137, 82-138, 2 males; July 12, 1982 (Braker HE), ANSP, no. 82-140, 1 female; July 20, 1982 (Braker HE), ANSP, no. 82-143, 1 female; September 2, 1983 (Rowell CHF), RC, nos. 83447a & 83447b, 2 males; September 1, 1991 (Waltz S), RC, no. 75004, 1 male. Prov. Limon: 5 km N. of Suretka, trail to Rio Uatsi, 200-220 m, September 20, 1983 (Rowell CHF), RC, nos. 83060a & 83060b, 1 male, 1 female, in cop.; Amubri, 70 m, August 23, 1992 (Gallardo G), INBio, no. CRI000 734414, 1 male; no. CRIO00 734404, 1 female; May 14, 1994 (Gallardo G), INBio, no. CRI001 871114, 1 male; no. CRIOO1 871113, 1 male; Barra del Colorado, R.N.F.S., Rio Sardinas, 10 m, April 10, 1994 (Araya F), INBio, no. CRIOO1 848425, 1 larva: June 1994 (Araya F), INBio, no. CRIO0I 847939, 1 female; Parque Nacional Tortuguero, Est. Cuatro Esquinas, 0 m asl, June 1990 (Chavarria U), INBio, no. CRI000 272555, 1 female; June 1990 (Quesada E), INBio, no. CRI000 462818, 1 male; July 1990 (Chavarria U), INBio, no. CRI000 244702, 1 female; June 1990 (Chavarria U), INBio, no. CRI000 272558, 1 male. Rio Segundo affl. Rio. Banano, 500 m, April 27, 1985 (Solis A), INBio, no. CRIOO! 013052 to 013055, 4 larvae. Rio Toro Amarillo, 7 km. W. of Guäpiles, late second growth tropical wet forest, August 21, 1964 (Hubbell SP), UMMZ, 1 male, 1 female; Rio Toro Amarillo, 10 km N of Guapiles, S of Quebrada Grande on trail to S. Valentino, 650 m, September 10, 1993 (Rowell CHF), RC, no. 93207, 1 male, 1 female, in cop.; Valle de la Estrella, Res. Biol. Hitoy Cerere, Est. Miramar, 500 m, July 1993 (Carballo G), INBio, no. CRIOOI 955534, 1 male; Est. Hitoy Cerere, 100 m, July 1991 (Carballo G), INBio, no. CRIOOO 585772, 1 larva V female; November 16, 1991 (Carballo G), INBio, no. CRIO00 524119, 1 female. Distribution. M. biolleyi is apparently confined to lowland rainforest of the Caribbean slope of northern Panama, Costa Rica, and southern Nicaragua (Nicaragua: 44 C. HUGH F. ROWELL punctata larvae Fic. 9. Larvae of Munatia. A. M. punctata. B. M. biolleyi. Conventions applying to both dia- grams: stippled areas, orange or gold: hatched areas, green or yellow, filled areas, black; open areas, dark brown. Scale 10 mm throughout, except for instar II larvae, where the scale is 5 mm. Both species show progressive reduction with age in the curvature of the pronotal crest and the REVISION OF MUNATIA O 77: relative length of the spine on the hind knee, and similar changes in general coloration. They differ from each other throughout the larval instars in the more angular rostrum and pronotum of punctata, in the detail of colour patterning, especially on the antennae of young larvae and the pronota of older males, and in the presence in fourth and later instars of biolleyi of a conspicuous white tubercle in the centre of the pronotal lobe. 46 C. HUGH F. ROWELL Zelaya, El Recreo (forêt), 30 m, October 1984 (Amedegnato C, Poulain S), MNHNP, specimen not examined). It occurs between sea level and about 700 m altitude. A specimen of Bruner's bears the locality label "Juan Vifias", which town lies at 1165 m. However, the surrounding countryside is precipitous, and even in Bruner's day sui- table forest habitat was probably restricted by agriculture to the bottom of the valley of the Rio Reventazön at less than 800 m. At least the modern records from "Turrialba" (nominally at 630 m) also certainly refer to the gorge of the Reventazon, here at about 500 m. Distribution map: see Fig. 10. NATURAL HISTORY The natural history of the adults of M. biolleyi is similar to that described for M. punctata. Males are active, females sluggish and cryptic. They are usually found in secondary vegetation in light-gaps caused by tree-falls or path construction. One female was captured on the pendant root of an Aroid epiphyte at 25 m up a large Dipteryx tree in closed forest (D. Perry, pers. comm., 1978 - specimen examined), so females too must either fly or at least climb up into trees. They are however certainly not typically arboricole, as is said for the related genus Xomacris (AMEDEGNATO & POULAIN 1986). Adults have been recorded from March to November, and copulations seen in August and September. Given the poor collecting intensity in December and January and the paucity of larval records, these data do not indicate whether the species breeds seaso- nally or all the through the year. The natural history of the larvae is not known. In particular it is not known whether they display the same visually based gregarious behaviour as larvae of M. punctata, but their more cryptic coloration makes this per- haps improbable. M. biolleyi is polyphagous on dicotyledons, like M. punctata. In the wild it has been seen eating Neurolaena (Asteraceae) and Alchornia and Plukenetia (Euphor- biaceae). In captivity it has accepted species of Convolvulaceae, Amaranthaceae, Rubiaceae, and the (monocotyledenous) Marantaceae, but refused to eat other monocotyledons (various Araceae, Arecaceae, Bromeliaceae, Cyclanthaceae, Helico- niaceae, Poaceae), ferns, or the dicotyledons Aspidospermum (Apocyanaceae), Ana- xagoria (Annonaceae), Cecropia (Moraceae), Solanum (Solanaceae), and various Melostomataceae (feeding trial records in part from H.E. Braker, pers. comm.). It is occasionally parasitized internally by the larvae of Tachinid flies. ACKNOWLEDGEMENTS I am grateful to the following colleagues for loan of material and for data from their collections: Drs. D. Azuma (Philadelphia), B. Hauser (Geneva), T. Kronstedt (Stockholm), M. O'Brien (Ann Arbor), and A. Solis (Santo Domingo de Heredia). I also thank Prof. C.S. Carbonell (Montevideo) for discussion and for photographs of various type specimens, Dr. C. Amédégnato (Paris) for the data on her Nicaraguan specimen, and Dr. D. Quintero (Panama City) for information on Panamanian locality names. Some field collecting was supported in part by a grant from the National Science Foundation (USA). REVISION OF MUNATIA 84 83 Munatia biolleyi 10 | 9 Munatia punctata D 85 84 Fic. 10 83 47 11 Distribution map of Munatia. The type locality of M. punctata in N. Panama is not known more precisely than the province (Chiriqui); the corresponding symbol (an open square) is here placed arbitrarily between Cerro Punto and Boquete. A further locality (not shown) for M. biolleyi is known in E. Nicaragua at 12°10' N, 84°19' W. The map includes sight records of the present author not listed in "Material examined". All localities for biolleyi are in the Caribbean lowlands or in the floor of river valleys draining into them. The localities for punctata are all in the highlands near the Pacific/Caribbean watershed, with one exception (Pozo Azul de Pirris in the Pacific lowlands). 48 C. HUGH F. ROWELL REFERENCES AMÉDÉGNATO, C. 1974. 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An analysis of the tribes of the Romaleinae with special reference to their internal genitalia (Orthoptera; Acrididae). Transactions of the American Entomological Society 85: 233-271. ROWELL, C. H. F. 1967. Experiments on aggregations of Phymateus purpurascens (Orthoptera, Pyrgomorphinae). Journal of Zoology, London 152, 179-193. STAL, C. 1873. Orthoptera nova descripsit. Kongliga Vetenskaps-akademiens Förhandlingar, Stockholm 30(4): 39-53. STAL, C. 1875. Observations orthopterologiques (1). Bihang till Kongliga Svenska Vetenskaps- akademiens Handlingar 3: 1-43. STAL, C. 1878. Systema acridiodeorum. Essai d'une systematisation des acridoidées. Bihang till Kongliga Svenska Vetenskaps-akademiens Handlingar 5(4): 1-100. Uvarov, B. P. & V. M. DirsH 1961. The diagnostic characters, scope and geographical distri- bution of the subfamily Romaleinae (Orthoptera: Acrididae). Proceedings of the Royal Entomological Society, London (B) 30: 153-160. REVUE SUISSE DE ZOOLOGIE 105 (1): 49-80; mars 1998 Report on some pholcid spiders collected in Guatemala and Honduras (Araneae, Pholcidae) Bernhard A. HUBER* Escuela de Biologia, Universidad de Costa Rica, Ciudad Universitaria, Costa Rica Report on some pholcid spiders collected in Guatemala and Honduras (Araneae, Pholcidae). - Descriptions and redescriptions of nine pholcid species from Guatemala and Honduras are given. The following species are new: Modisimus pana sp.n., Mod. ixobel sp.n., and Metagonia asintal sp.n. from Guatemala, Modisimus lancetilla sp.n. and ‘Coryssocnemis’ tigra sp.n. from Honduras. For two species the males are newly described: Meta- gonia blanda Gertsch, 1973 and Mer. belize Gertsch, 1986. Two species are redescribed: “Coryssocnemis’ furcula Cambridge, 1902 and Modisimus cornutus Kraus, 1955. Key-words: Pholcidae - neotropics - Guatemala - Honduras. INTRODUCTION Pholcids are apparently among the most common and diverse spider families in the neotropics (Huber 1997 b). About 350 species are presently known from the New World, but this is probably only a small fraction of the actual number. The present paper reports on part of the pholcids collected by the author in Guatemala, Honduras and Nicaragua during a 4-week trip in September and October 1996. The fact that about 35 species were collected in this short time demonstrates the abundance of pholcids in these countries, for which only about 15 species were previously recorded. This paper presents a subjective selection of the most ‘inte- resting’ species. The criteria for “interesting” were quite arbitrary, such as extraor- dinary morphological character states (e.g. Modisimus pana: pedipalps; “Coryssocne- mis’ tigra: epigynum, male bulb; Metagonia belize, Met. lancetilla, and Modisimus ixobel: male chelicerae); unusual variation in genitalia (‘Coryssocnemis’ furcula: epi- gynum); the first case of stridulation in the genus Metagonia (M. asintal); the surprisingly wide distribution of a troglophile species (Metagonia blanda). Modisimus cornutus Kraus, which was previously only known from the island of Utila, is reported from the mainland. For two species (Metagonia belize, Met. blanda), the *Present address: Department of Entomology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024. Manuscript accepted 30.07.1997 50 BERNHARD A. HUBER males are newly described. The first is new for Guatemala, the second is new for Honduras. The known range of ‘Coryssocnemis’ furcula (new for Honduras) is widely extended. ‘Coryssocnemis’ furcula is redescribed under the original name although it is almost certainly not congeneric with the type species of the genus (C. callaica Simon, 1893 from Venezuela; see Huber in press b for discussion of the genus). This is also true for ‘C.’ tigra n.sp. which may or may not be congeneric with °C.’ furcula. Only future revisions and phylogenetic analyses may justify the creation of a new genus (or new genera) for these species. MATERIAL AND METHODS Previously described species were borrowed from the following institutions: American Museum of Natural History, New York (AMNH), Natural History Mu- seum, London (BMNH), Senckenbergmuseum Frankfurt (SMF). Holotypes and paratypes of new species as well as vouchers are deposited in the AMNH. Further types and vouchers are deposited in the Muséum d’histoire naturelle, Genève (MHNG). The other material is provisionally deposited in the author’s collection. Descriptions follow the style currently used tor pholcid spiders (for discussion of style see Huber in press a). Drawings were made with a compound microscope with camera lucida and later completed with a dissecting microscope. Measurements (all in mm) were taken with ocular micrometers in a compound or a dissecting micro- scope. Averages (arithmetic means) are given for N25. Prosoma length was defined as the distance between frontal face of eye region and posterior border of carapace medially, but it varies widely with the angle at which the prosoma is viewed. “Cara- pace” is referred to as the dorsal part of the prosoma. The most accurate indicators of size are probably prosoma width and tibia length. Total size is simply the sum of prosoma length and opisthosoma length, regardless of the petiolus, and is given as an approximate indication of overall size. The tibia index (“tibind”) is the length of the tibia divided by its width at the middle, and is thus a measure of the ‘slenderness’ of the legs. In the diagnoses, species with an average total length of >3 mm are defined as “large”, those smaller than 2.5 mm are “small”. Diagnoses of the genera are not given, since there are recent discussions of each of the treated genera (Modisimus Simon, 1893: Huber in press a; Coryssocnemis Simon, 1893: Huber in press b: Metagonia Simon, 1893: Huber 1997a). DESCRIPTIONS AND REDESCRIPTIONS Modisimus pana sp.n. Figs 1-10 MATERIAL EXAMINED: Male holotype and one female paratype from creek near Panajachel, Dept. Sololä, Guatemala, elev. about 1700 m, 17 Sept. 1996 (B. A. Huber), in AMNH. | male and 1 female paratypes, same collection data, in MHNG. 2 males, 1 female, 1 juv., same collection data, in authors collection. Etymology: ‘Pana’ is the vulgar name of Panajachel, the type locality. PHOLCIDS FROM GUATEMALA AND HONDURAS 5] Diagnosis: Small to medium sized Modisimus with high eye turret (Fig. 1), characterized by the male genitalia (bulbal apophyses; cymbium with bent procursus and additional dorsal apophysis - Figs 3-4, 7-8), the male chelicerae with their characteristically modified hairs (Figs 5-6), and the female epigynum and internal genitalia (Figs 9-10). The species is similar to M. palenque Gertsch, 1977 (I have seen the male holotype of this species), but easily distinguished by details in the characters mentioned. Distribution: Known only from above material from type locality. DESCRIPTION Male: Prosoma dorsally pale ochre, with brown mark medially (Fig. 2) and brown clypeus, eye turret brown posteriorly, sternum medially pale ochre, laterally slightly darker. Opisthosoma dorsally grayish ochre with large black spots and few small white spots (Figs 1-2), ventrally lighter, with brownish genital plate and brown spot between genital plate and spinnerets. Legs yellow-brown, with dark rings on femora (distally) and tibiae (proximally and distally). Six eyes on high eye turret (the holotype and one other male completely lack the AMEs, but two males have minute AMEs, cf. Fig. 2). Chelicerae with high number of modified hairs anteriorly (Figs 5- 6). Pedipalps distinctive, as shown in Figs 3-4. Measurements of male holotype: Total length: 2.4; prosoma width: 0. OSE length: 0.8; opisthosoma length: 1.6; legs: I 2 3 4 fem 6.1 4.2 3.4 4.1 pat 0.4 0.4 0.4 0.4 tib 5.9 3.9 3.1 3.6 met 10.1 6.4 4.6 SI tar 1.4 1.0 0.8 0.9 total 23.9 15.9 12 14.5 tibind 69 42 36 42 Female: Colors as in male, minute AMES present in two individuals, absent in the third. Epigynum simple brown plate (Fig. 9). Internal genitalia as in Fig. 10. Measurements of a female paratype: Total length: 2.6; prosoma width: 0.79; length: 0.8; opisthosoma length: 1.6; legs: I 2 3 4 em 3.8 27 2.2 27 pat 03 03 03 03 tib 38 2.4 19 2.2 me 6.0 36 2.8 33 ar 13 0.8 0.7 0.7 total 15.2 9.8 7.9 9.2 Are 52 33 30 30 52 BERNHARD A. HUBER Fics 1-4 Modisimus pana sp.n. 1, Male, lateral view. 2, Male, dorsal view. 3, Left pedipalp, prolateral view. 4, Left pedipalp, retrolateral view. Scales: (1,2) 1 mm, (3,4) 0.3 mm. PHOLCIDS FROM GUATEMALA AND HONDURAS 53 Fics 5-10 Modisimus pana sp.n. 5, Male chelicerae, frontal view. 6, Modified hairs on male chelicerae. 7, Right procursus, retrolateral view. 8, Left bulb, retrolatero-ventral view. 9, Epigynum, ventral view. 10, Epigynum, dorsal view. Scales: (5,7-10) 0.2 mm, (6) 0.01 mm. Variation: All males except the holotype are quite pale, which is usual for recently molted spiders. Tibia 1 length in other material: males: 5.7, 6.2, 6.4; females: 3.3, 3.5. Habitat: The spiders were found in small webs with a domed sheet of silk, about 50 cm above the ground, between leaves in the low vegetation. ) 54 BERNHARD A. HUBER Modisimus ixobel sp.n. Figs 11-20 MATERIAL EXAMINED: Male holotype and 2 female paratypes from forest near Finca Ixobel, near Poptun, Dept. Petén, Guatemala, 22 Sept. 1996 (B. A. Huber), in AMNH. 2 female paratypes, same collection data, in MHNG. 5 females, 3 juvs, same collection data, in author’s collection. Etymology: Named for Finca Ixobel, which 1s close to the type locality. Diagnosis: Small Modisimus with high eye turret (Fig. 11), distinguished from other species of the genus by the male genitalia (bulb with dorsal apophyses; simple procursus; palpal femur with distal bulge - Figs 13-16), the male chelicerae with their characteristically modified hairs in an uncommon position (Figs 17-18), and the female epigynum and internal genitalia (Figs 19-20). Distribution: Known only from above material from type locality. DESCRIPTION Male: Prosoma dorsally pale ochre yellow, with brown mark medially and on eye turret (Fig. 12), clypeus and sternum ochre yellow. Opisthosoma dorsally greenish gray with large black spots and few small white spots (Figs 11-12), ventrally lighter, with brown- ish genital plate and brown stripe between genital plate and spinnerets. Legs ochre yellow, without rings. Six eyes on high eye turret. Chelicerae with five strong modified hairs on outer distal margin (Figs 17-18). Pedipalps distinctive, as shown in Figs 13-14. Measurements of male holotype: Total length: 1.9; prosoma width: 0.79; length: 0.7; opisthosoma length: 1.2; legs: | 2 3 4 fem 6.2 4.3 By) 4.1 pat 0.4 0.4 0.3 0.4 tib 6.2 4.1 3.1 3.6 met 10.9 6.3 4.7 Doll tar 1.4 1.0 0.8 0.9 total DIA 16.1 12.1 14.7 tibind 82 56 44 49 Female: Colors as in male, but legs with dark rings on femora (distally) and tibiae (proximally and distally). Epigynum simple brown plate (Fig. 19). Internal genitalia as in Fig. 20. Measurements of a female paratype: total length: 2.5; prosoma width: 0.92; length: 0.8; opisthosoma length: 1.7; legs: l 2 3 4 fem 6.2 4.4 3.6 4.4 pat 0.4 0.4 0.4 0.4 tib 6.2 4.1 3.3 3.6 met 10.6 65 4.9 5.9 tar 1,7 112 0.9 0.9 total 25.1 16.6 Sel 1522 tibind 70 46 38 40 PHOLCIDS FROM GUATEMALA AND HONDURAS 55 LPS SES eyz Fics 11-14 Modisimus ixobel sp.n. 11, Male, lateral view. 12, Male, dorsal view. 13, Right pedipalp, retro- lateral view. 14, Right pedipalp, prolateral view. Scales: (11,12) 1 mm, (13,14) 0.3 mm. 56 BERNHARD A. HUBER Fics 15-20 Modisimus ixobel sp.n. 15, Right bulb, retrolateral view. 16, Right procursus, prolateral view. 17, Male chelicerae, frontal view. 18, Modified hairs on male chelicerae. 19, Epigynum, ventral view. 20, Epigynum, dorsal view. Scales: (15,17) 0.2 mm, (16) 0.1 mm, (18) 0.03 mm, (19-20) 0.3 mm. Variation: Tibia | in 7 other females: 5.4-6.6; x=6.0. Habitat: The spiders were found in small webs with a domed sheet of silk, close to the ground, mostly in dark sheltered places. Modisimus cornutus Kraus, 1955 Figs 21-31 Modisimus cornutus KRAUS 1955: 13-14; pl. 1, figs 19-21. MATERIAL EXAMINED: Male holotype and 9 female paratypes from Isla Utila, Honduras, 26 Sept. 1951 (Peters), SMF 8685-8687 and 8710. 7 males, 5 females, 3 juvs from Jardin Botanico Lancetilla, near Tela, Dept. Atlantida, Honduras, elev. about 50 m, 28 Sept. 1996 (B. A. Huber), one pair in AMNH, one pair in MHNG, others in author’s collection. PHOLCIDS FROM GUATEMALA AND HONDURAS 57 Diagnosis: Small Modisimus with high eye turret (Fig. 21), distinguished from other species of the genus by the male genitalia (bulb with a pair of dorsal processes; cymbium with dorsal apophysis and procursus with translucent dorsal spine - Figs 23- 24, 26, 28), the male chelicerae with a patch of simple spines (Fig. 27), and the female epigynum and internal genitalia (Figs 29-31). Note: The diagnosis of the original description refers to three cylindrical processes on the pedipalpal tarsus. However, two of these processes do not originate from the tarsus (cymbium) but from the bulb (Fig. 26). Distribution: Known only from above material from two localities in northern Honduras. REDESCRIPTION Male (Lancetilla): Prosoma ochre, only eye turret darker. Opisthosoma dorsally greenish gray with large black spots and few small white spots (Figs 21-22), ventrally lighter, with light brownish genital plate and short black stripe between genital plate and another brown spot near spinnerets. Legs ochre brown, without rings. Six eyes on high eye turret. Chelicerae with a patch of about 10 modified hairs on each side (Fig. 27). Pedipalps distinctive, as shown in Figs 23-24. The male holotype is much paler, and the patches of modified hairs on the chelicerae are located slightly more proximally. Measurements of a male from Lancetilla: Total length: 2.3; prosoma width: 0.89; length: 0.8; opisthosoma length: 1.5; legs: | 2 3 4 fem 5.4 3.6 2.9 3.6 pat 0.4 0.4 0.4 0.4 tib IH 3.6 2.9 35) met 9.7 ID 4.2 Sl tar 1.4 0.9 0.8 0.8 total 22.6 14.0 2 34 tibind 75 49 41 46 Female (Lancetilla): Colors as in male, but with broad dark median stripe dorsally on prosoma, legs also without rings. Epigynum brown, slightly protruding, with posterior sclerotized plate (Figs 29-30). Internal genitalia as in Fig. 31. Most paratypes are very pale, but in several the dark spots on the opisthosoma are still discernible. The epigynum looks identical in side view, but in ventral view the median arch on the frontal plate (arrow in Fig. 30) is not visible in some of the females. Measurements of a female from Lancetilla: Total length: 2.0; prosoma width: 0.79; length: 0.7; opisthosoma length: 1.3; legs: 58 BERNHARD A. HUBER Fics 21-24 Modisimus cornutus Kraus. 21, Male, lateral view. 22, Male, dorsal view. 23, Left pedipalp, prolateral view. 24, Left pedipalp, retrolateral view. Scales: (21,22) 1 mm, (23,24) 0.3 mm. PHOLCIDS FROM GUATEMALA AND HONDURAS 59 Fics 25-31 Modisimus cornutus Kraus. 25, Left palpal trochanter and femur, prolateral view. 26, Left bulb, retrolateral view. 27, Male chelicerae, frontal view. 28, Left procursus, prolateral view. 29, Epigynum, lateral view, frontal side on the right. 30, Epigynum, ventral view; arrow points to arch that is not visible in some females. 31, Epigynum, dorsal view. Scales: (25-27, 29-31) 0.2 mm, (28) 0.1 mm. I 2 3 4 fem 3.6 24 1.9 2.5 pat 0.3 0.3 0.3 0.3 tib 3.6 22 1.8 22 met 5.7 3.3 2.6 3.2 tar 1.2 0.8 0.7 0.7 total 14.4 9.0 7.3 8.9 tibind 977 35 29 25 60 BERNHARD A. HUBER Variation: Tibia 1 in other material: Lancetilla: 6 males: 4.8-5.4; x=5.2: 4 females: Sooo oa Ons Ut Mimale 7.0: > temales: 2579) 1°"); Habitat: In Lancetilla the spiders were found in small webs with a domed sheet of silk, close to the ground, mostly in dark sheltered places, along a dried brook-bed. ‘Coryssocnemis’ tigra sp.n. Figs 32-40 MATERIAL EXAMINED: Male holotype and female paratype from Parque Nacional La Tigra, (about 10 km NE Tegucigalpa), Dept. Francisco Morazan, Honduras, elev. about 1800-1900 m, 2 Oct. 1996 (B. A. Huber), in AMNH. 1 female paratype, same collection data, in MHNG. 2 females and 4 juvs, same collection data, in author’s collection. Etymology: Named for type locality. Diagnosis: Medium sized to large dark pholcid with eight eyes on slightly elevated ocular area. Male and female genitalia highly distinctive (bulb with a pair of large ventral processes; simple, slender procursus; epigynum with a pair of long posterior processes - Figs 39-40). Male chelicerae unmodified. Female with paired stridulatory apparatus between prosoma and opisthosoma. Distribution: Known only from above material from type locality. DESCRIPTION Male: Prosoma dorsally pale ochre with large dark brown area medially (Fig. 33). Ocular area and clypeus also brown. Opisthosoma dorsally greenish gray with large black spots and few small white spots (Figs 32-33), ventrally lighter, with brownish genital plate. Legs ochre brown, with dark rings on femora (distally) and tibiae (only proximally). Eight eyes on slightly elevated ocular area (Fig. 32). Chelicerae unmo- dified. Pedipalps distinctive, as shown in Figs 34-35. Measurements of male holotype: Total length: 2.9; prosoma width: 1.43; length: 1.3; opisthosoma length: 1.6; legs: I 2 3 4 fem 6.2 SO 4.4 5.2 pat 0.6 0.6 0.5 0.5 tib 6.5 4.9 4.1 4.9 met 8.4 6.2 IS) 6.2 tar 1.9 1185) 13 13 total 23.6 18.4 15.6 18.1 tibind 42 30 DI 34 Female: Colors as in male. The structure of the external genitalia is unique among pholcids (Fig. 39). Internal genitalia as shown in Fig. 40. Stridulatory organ con- sisting of a pair of humps on the rear side of the prosoma dorsally, and a corres- ponding pair of sclerotized plates on the opisthosoma (arrow in Fig. 39). Measurements of a female paratype: total length: 3.3; prosoma width: 1.24; length: 1.2; opisthosoma length: 2.1; legs: PHOLCIDS FROM GUATEMALA AND HONDURAS 61 Sio zz, 717 7 Fics 32-35 ‘Coryssocnemis’ tigra sp.n. 32, Male, lateral view. 33, Male, dorsal view. 34, Right pedipalp, retrolateral view. 35, Right pedipalp, prolateral view. Scales: (32-33) 1 mm, (34-35) 0.5 mm. 62 BERNHARD A. HUBER Fics 36-40 ‘Coryssocnemis’ tigra sp.n. 36, Right bulb, retrolateral view. 37, Right procursus tip, retro- lateral view. 38, Right procursus tip, prolateral view. 39, Female opisthosoma, ventral view; arrow points to one of the two sclerotized plates which are part of the ‘stridulatory apparatus’. 40, Epigynum, dorsal view. Scales: (36, 39-40) 0.5 mm, (37-38) 0.1 mm. PHOLCIDS FROM GUATEMALA AND HONDURAS 63 2 3 4 fem 4.8 3.8 3.0 3.9 pat 0.5 0.5 04 04 tib 49 3.4 27 3.5 met 62 44 NG 4.6 can 1.7 13 12 12 me: 18.1 13.4 10.9 13.6 Gbind 39 27 22 28 Variation: In one female the rings on the legs are very dark, and both are preceded and followed by light rings, giving the legs a very vivid pattern. Tibia 1 in 3 other females: 4.8, 5.1, 5.1. Habitat: The spiders were found in essentially the same type of web and in the same habitat as ‘C.’ furcula (see below). ‘Coryssocnemis’ furcula Cambridge, 1902 Figs 41-52 Coryssocnemis furcula CAMBRIDGE 1902: 371; pl. 35, figs 8, 8a-b. Kraus 1955: 14; pl. 2, figs 22-23. TYPE MATERIAL: Female holotype from “Tecpam in the Los Altos region” (Tecpän Guatemala, Dept. Chimaltenango), Guatemala, elev. about 2300 m, no date (Stoll), in BMNH, examined. OTHER MATERIAL EXAMINED: HONDURAS: 4 males, 3 females, 5 juvs from Parque Nacional La Tigra, (about 10 km NE Tegucigalpa), Dept. Francisco Morazän, elev. about 1800-1900 m, 2 Oct. 1996 (B. A. Huber), one pair in AMNH, one pair in MHNG, others in authors collection. GUATEMALA: 2 males, 4 females from near Panajachel (this is only about 20 km from the type locality), Dept. Solola, elev. about 1700 m, 17 Sept. 1996 (B. A. Huber), in authors collection. 3 females from near Zunil (about 7 km SE Quetzaltenango), Dept. Quetzaltenango, elev. about 2150 m, 18 Sept. 1996 (B. A. Huber), in author’s collection. EL SALVADOR: | male, 4 females from Finca San Jorge near Santa Ana, Dept. Santa Ana, elev. about 1000 m, 25 April 1951 (O. Kraus), SMF 8545-8546. Diagnosis: Very large dark pholcid with strong legs, characterized by the male genitalia (bulb with rounded prolateral process; procursus slender with distal spine - Figs 43-46), the male chelicerae with a pair of strong pointed apophyses (Fig. 47), and the female epigynum and internal genitalia (Figs 48-51). Distribution: The species has been found in mountainous regions of Guatemala, El Salvador, and Honduras (Fig. 52). REDESCRIPTION Male: Prosoma dorsally pale ochre gray with characteristic pattern of dark brown spots (Fig. 42). Clypeus and sternum light brown. Opisthosoma dorsally dark greenish gray with many black and (smaller) white spots (Fig. 41), ventrally lighter, with brown genital plate and black spinnerets. Legs brown, femora with two dark rings distally, patellae dark, tibiae with dark rings proximally and distally, distal ring followed by light ring. Eight eyes on slightly elevated ocular area (Fig. 41). Cheli- cerae with a pair of large anterior apophyses (Figs 41, 47). Pedipalps distinctive, as 64 | \ BERNHARD A. HUBER Hare (A A < “a Fics 41-44 ‘Coryssocnemis’ furcula Cambridge. 41, Male, lateral view. 42, Male prosoma, dorsal view. 43, Left pedipalp, prolateral view. 44, Left pedipalp, retrolateral view. Scales: (41-42) 2 mm, (43-44) | mm. PHOLCIDS FROM GUATEMALA AND HONDURAS 65 YELLE PHT Fics 45-51 ‘Coryssocnemis’ furcula Cambridge. 45. Left genital bulb, ventral view. 46, Left procursus, prolateral view. 47, Male chelicerae, frontal view. 48, Epigynum, female from La Tigra, ventral view. 49, Epigynum, female from La Tigra, lateral view, frontal side on the right. 50, Epigynum, female from La Tigra, dorsal view. 51, Epigynum, female from Panajachel, lateral view. Scales: (45-47) 0.5 mm, (48-51) 1 mm. 66 BERNHARD A. HUBER 'Coryssocnemis' furcula e Metagonia blanda EIGS92 Distribution of ‘Coryssocnemis’ furcula Cambridge and Metagonia blanda Gertsch. shown in Figs 43-44. The palpal femur bears distally, on the ventral side, a small apophysis. It is not clear whether this is a homologue of the ‘pup-apophysis’ that characterizes the Modisimus-group sensu Huber (in press b). Measurements of a male from La Tigra: Total length: 7.7: prosoma width: 3.19; length: 2.8: opisthosoma length: 4.9; legs: | 2 3 4 fem 12.0 9.7 8.1 10.4 pat 1.4 1.4 3 152 tib 12.0 8.6 6.8 9.0 met [SA les 9.0 11.8 tar 52 32 2.4 2.6 total 45.7 34.2 27.6 35.0 tibind 28 23 18 24 Female (La Tigra). Colors as in male, but with large brown epigynum (Figs 48-49). Internal genitalia relatively small in relation to the large epigynum (10250) Measurements of a female from La Tigra: Total length: 7.4; prosoma width: 2.61; length: 2.3; opisthosoma length: 5.1; legs: PHOLCIDS FROM GUATEMALA AND HONDURAS 67 | 2 3 4 fem 8.8 Tail 59 8.0 pat 12 1.1 1.0 1.0 tib 93 6.5 4.9 6.8 met 10.9 8.0 6.4 8.6 tar 4.1 2.6 2.0 210 total 34.3 2573 20.2 26.4 tibind 24 18 14 19 Variation: While the male genitalia are almost identical in all the studied specimens, there is substantial variation in the female epigynum. Females from Honduras (La Tigra) have relatively small frontal apophyses (Fig. 49), while those from Guatemala (Tecpan, Panajachel, Zunil), have much larger apophyses (Fig. 51; Cambridge 1902: figs 8, 8b). Females from El Salvador (Santa Ana) are intermediate, though closer to those from Guatemala. Tibia 1 in other material: La Tigra: 3 males: 10.0, 11.3, 12.6; 2 females: 7.0, 9.3. Panajachel: 2 males: 11.3, 12.0; 4 females: 9.9, 10.1, 10.1, 10.4; Zunil: 2 females: 7.4, 9.3. Santa Ana: 1 male: 12.8; 4 females: 10.6, 10.7, 11.2, 11.3. Habitat: The spiders were found in sheet webs close to the ground, mostly in dark sheltered places, along creeks (Panajachel, Zunil) or footpaths (La Tigra). When distrubed the spiders fled into a funnel that led into the substrate, much like agelenids, but with the difference that the funnel was the continuation of the underside of the sheetweb. Metagonia lancetilla sp.n. Figs 53-62 MATERIAL EXAMINED: Male holotype and female paratype from Jardin Botanico Lancetilla, near Tela, Dept. Atlantida, Honduras, elev. about 50 m, 28 Sept. 1996 (B. A. Huber), in AMNH. 1 female paratype, same collection data, in MHNG. Etymology: Named for type locality. Diagnosis: Small epigean Metagonia, distinguished from other species of the genus by male and female genitalia (procursus with long ventral hinged process; epigynum with posterior process - Figs 54-55, 59-62), and male chelicerae with a pair of apophyses (Figs 56, 58). Distribution: Known only from above material from type locality. DESCRIPTION Male: Entire body pale ochre yellow (Fig. 53), only tibia-metatarsus joints dark brown, patellae only slightly darkened. Pedipalps as shown in Figs 54-55, procursus with long hinged process (arrow in fig. 59). Palpal femur with ventral apophysis distally (Fig. 57). Chelicerae with a pair of large distinctive apophyses frontally (Figs 56, 58), which seem to carry about four deeply inserted modified hairs each. Clypeus unmodified. Measurements of male holotype: Total length: 1.7; prosoma width: 0.67; length: 0.6; opisthosoma length: 1.1; legs: 68 BERNHARD A. HUBER Fics 53-55 Metagonia lancetilla sp.n. 53, Male, dorsal view. 54, Left pedipalp, prolateral view. 55, Left pedipalp, retrolateral view. Scales: (53) 0.5 mm, (54-55) 0.3 mm. 1 2 3 4 fem 4.5 2.9 2.0 3.0 pat 0.3 0.3 0.3 0.3 tib 4.5 DT] 127 DIS met Tot 4.3 2.5 3.8 tar 1.3 0.9 0.6 0.6 total 18.3 11.1 7.1 10.2 tibind 71 43 30 40 Female: Colors as in male. Epigynum of same color, of distinctive shape (Figs 60- 61). Internal genitalia as in Fig. 62. Measurements of a female paratype: total length: 1.7; prosoma width: 0.57; length: 0.6; opisthosoma length: 1.1; legs: PHOLCIDS FROM GUATEMALA AND HONDURAS 69 Fics 56-62 Metagonia lancetilla sp.n. 56, Male chelicerae, frontal view. 57, Male palpal femur, lateral view. 58, Apophysis on left male chelicera. 59, Left procursus, prolateral view: arrow points to ‘hinged process’. 60, Epigynum, lateral view, frontal side on the right 61, Epigynum, ventral view. 62, Epigynum, dorsal view. Scales: (56, 59, 62) 0.2 mm, (57) 0.1 mm, (58) 0.03 mm, (60- 61) 0.3 mm. I 2 3 4 fem 3.6 255 1.7 2.8 pat 0.3 0.3 0.2 0.3 tib 33 2.1 1.4 DD. met 5.8 3.3 2.0 3.3 tar 1.2 0.7 0.5 0.6 total 14.2 8.9 5.8 9.2 tibind 2 35 25 39 70 BERNHARD A. HUBER Tibia 1 in other female: 3.5. Habitat: Underside of large leaves. Metagonia asintal sp.n. Figs 63-73 MATERIAL EXAMINED: Male holotype, 1 male and 2 female paratypes from near El Asintal (about 8 km NW Retalhuleu), Dept. Retalhuleu, Guatemala, elev. about 300 m, 19 Sept. 1996 (B. A. Huber), in AMNH. 2 male and 2 female paratypes, same collection data, in MHNG. 3 males, 4 females, 3 juvs, same collection data, in author’s collection. Etymology: Named for type locality. Diagnosis: Small epigean Metagonia, distinguished from other species of the genus by male and female genitalia (details of procursus; large flap directed anteriorly on epigynum - Figs 65-67, 70-73), and male chelicerae with only a pair of tiny modified hairs on each side (Figs 68-69). Distribution: Known only from above material from type locality. DESCRIPTION Male: Prosoma pale ochre yellow, with dark spots at ocular area and on rear side of prosoma dorsally (Figs 63-64). Sternum whitish. Legs also pale ochre yellow, with brown patellae and dark brown tibia-metatarsus joints. Opisthosoma grayish ochre, with or without spots dorsally. Pedipalps as shown in Figs 65-67. The chelicerae appear unmodified in the dissecting microscope, but they have two tiny club shaped hairs on each side, and stridulatory ridges laterally (Figs 68-69). The scrapers of the stridulatory organs are club shaped hairs on the prolateral sides of the pedipalpal femora (Figs 65-66). Clypeus unmodified. Measurements of male holotype: Total length: 2.5; prosoma width: 0.78; length: 0.8; opisthosoma length: 1.7; legs: | 2) 3 4 fem al 2 DAS 32 pat 0.3 0.3 0.3 0.3 tib 5.1 3.0 1.9 2.8 met 8.6 4.5 2.8 4.1 tar 1.4 0.9 0.7 0.7 total 20.5 11.9 8.0 11.1 tibind 65 43 30 40 Female: Colors as in male, but spots on prosoma less dark. Epigynum pale, with distinctive lobe directed forwards (Figs 71-72). Internal genitalia as in Fig. 73. Measurements of a female paratype: total length: 2.4; prosoma width: 0.70; length: 0.8; opisthosoma length: 1.6; legs: PHOLCIDS FROM GUATEMALA AND HONDURAS il NENNEN WAAL \ | \ 4] FıGs 63-67 Metagonia asintal sp.n. 63, Male, lateral view. 64, Male, dorsal view. 65, Left pedipalp, prolateral view. 66, Modified hair (scraper) on palpal femur. 67, Left pedipalp, retrolateral view. Scales: (63-64) 1 mm. (65,67) 0.3 mm, (66) 0.01 mm. 72 BERNHARD A. HUBER Fics 68-73 Metagonia asintal sp.n. 68, Male chelicerae, frontal view. 69, Modified hairs on male chelicerae. 70, Left procursus, prolateral view. 71, Epigynum, ventral view. 72, Epigynum, lateral view, frontal side on the right. 73, Epigynum, dorsal view. Scales: (68, 70) 0.1 mm, (69) 0.01 mm, (71-73) 0.2 mm. Il 2 3 4 fem 4.1 2.8 17 2.9 pat 0.3 0.3 0.3 0.3 tib 3.9 2.3 1.4 25 met 6.0 3.6 DID; 3.4 tar 1.2 0.8 0.6 0.7 total 155 9.8 6.2 9.6 tibind 56 37 22 33 PHOLCIDS FROM GUATEMALA AND HONDURAS 73 Variation: The pattern on the opisthosoma is highly variable: some specimens have no spots at all, others have only a pair of dark spots medially (as in Fig. 64), others have only many white spots. In some males the spots on the prosoma are less dark. Tibia 1 in other material: 5 males: 4.3-5.4, x=4.7; 8 females: 3.4-3.9, x=3.6. Habitat: Underside of large leaves. Metagonia blanda Gertsch, 1973 Figs 52, 74-82 Metagonia blanda GERTSCH 1973: 152; figs 20-22. GERTSCH 1986: 57. MATERIAL EXAMINED: GUATEMALA: Female holotype from Gruta de Silvino, Dept. Izabal, 34 km SW Puerto Barrios, 20-22 August 1969 (S. & J. Peck), AMNH. 12 males, 9 females from cave near Finca Ixobel, near Poptun, Dept. Petén, 22 Sept. 1996 (B. A. Huber), 2 males, 2 females in AMNH, 2 males, 2 females in MHNG, others in author’s collection. 1 female from Cueva Seamay, Finca Senahu, Senahu, Dept. Alta Verapaz, 24-26 Aug. 1969 (S. & J. Peck), AMNH. 3 females from Cueva de la Coche, 1.5 mi W Livingston, Dept. Izabal, 21 Aug. 1969 (S. & J. Peck), AMNH. 2 females, 3 juvs from Cueva Lanquin, Lanquin, Dept. Alta Verapaz, 28. Aug. 1969 (S. & J. Peck), AMNH. HONDURAS: 5 males, 4 females from cave at Quebrada Sesesmil, near Copan, Dept. Copan, elev. about 600 m, 26 Sept. 1996 (B. A. Huber & O. A. Cardona), in author’s collection. Diagnosis: Small troglophile Metagonia, distinguished from other species of the genus by male and female genitalia (procursus with long ventral hinged process; epigynum with paired posterior lobe - Figs 76-78, 81-82) and the male chelicerae with usually three modified hairs distally on each side (Figs 79-80). Distribution: This species is now known from six caves in Guatemala and Honduras. For being a troglophile pholcid, the range is surprisingly wide (Fig. 52; cf. troglophile species of the genus Anopsicus in Mexico which are usually confined to just one single cave - Gertsch 1982). DESCRIPTION Male: Prosoma pale ochre yellow, with darker spot on rear side of prosoma dorsally (Fig. 75). Sternum whitish. Legs also pale ochre yellow, with slightly darker patellae and tibia-metatarsus joints. Opisthosoma whitish, without. spots (Figs 74-75). Pedipalps as shown in Figs 76-77. The chelicerae appear unmodified in the dissecting microscope, but they have a group of usually three modified hairs on each side near the bases of the fangs (Figs 79-80). Clypeus with small unpaired projection (Fig. 75). Measurements of a male from Poptun: Total length: 2.5; prosoma width: 0.92; length: 0.8; opisthosoma length: 1.7; legs: I 2 3 4 om 5.4 36 6. 30 pat 0.4 0.4 0.4 0.4 tib 57 316 2.3 33 He 9.1 5.5 35 49 (an 13 0.9 0.7 0.7 total 21.9 14.0 9.5 13.0 tibind 72 46 29 4l 74 BERNHARD A. HUBER EER E N || NC Fics 74-77 Metagonia blanda Gertsch. 74, Male, lateral view. 75, Male, dorsal view. 76, Left pedipalp, prolateral view. 77, Left pedipalp, retrolateral view. Scales: (74-75) 1 mm, (76-77) 0.3 mm. PHOLCIDS FROM GUATEMALA AND HONDURAS 75 FIGs 78-82 Metagonia blanda Gertsch. 78, Left procursus, prolateral view. 79, Male chelicerae, frontal view. 80, Modified hairs on male chelicerae. 81, Epigynum, ventral view. 82, Epigynum, lateral view, frontal side on the right. Scales: (78-79) 0.2 mm, (80) 0.01 mm, (81-82) 0.3 mm. REDESCRIPTION Female: Colors as in male. Epigynum pale, of distinctive shape (Figs 81-82). Measurements of female holotype: total length: 2.9; prosoma width: 1.05; length: 1.0; opisthosoma length: 1.9; legs: 76 BERNHARD A. HUBER I 2 3 4 fem Sif 4.1 3.0 4.5 pat 0.4 0.4 0.4 0.4 tib 5.9 3.8 2.6 3.9 met 9.7 5.9 3.8 5.4 tar 1.4 1.0 0.7 0.7 total 23.1 152) 10.5 14.9 tibind 62 40 27 41 Measurements of a female from Poptun: total length: 2.4; prosoma width: 0.89; length: 0.8; opisthosoma length: 1.6; legs: 1 2 3 4 fem 5.0 3.6 2.6 3.8 pat 0.4 0.4 0.4 0.4 tib 5.0 3.3 DD 3.3 met 8.0 4.9 379 4.8 tar 1.4 0.9 0.7 0.7 3 total = 198 dr on m : 92 e. 13,0 tibind 53 35 23 35 Variation: Tibia 1 in other material: Poptun: 10 males: 5.3-5.9, x=5.7; 8 females: 4.8- 5.4, X=5.0. Copan: 5 males: 5.0-5.6, X=5.2; 4 females: 4.1, 4.4, 4.8, 4.8. Habitat: Both in Poptun and in Copan, the spiders were only found within the caves, several meters beyond the entrance, although in Poptun the cave was surrounded by a dense humid forest. In the caves they lived in small crevices and shelters were they apparently built simple attached webs. When disturbed the spiders swiftly ran away over the rock surface. Metagonia belize Gertsch, 1986 Figs 83-92 Metagonia belize GERTSCH 1986: 55; figs 38-39. MATERIAL EXAMINED: BELIZE: Female holotype from unnamed fissure near Mountain Cow Cave, Cayo District, May 1977 (L. McNatt), in AMNH. GUATEMALA, Dept. Petén: 6 males, 6 females from forest near Finca Ixobel, near Poptun, 22 Sept. 1996 (B. A. Huber), 2 males, 2 females in AMNH, 2 males, 2 females in MHNG, others in author’s collection. 5 males, 6 females from Parque Nacional Cerro Cahui, elev. about 150 m, 21 Sept. 1996 (B. A. Huber), in author’s collection. 7 males, 2 females from Tikal, elev. about 150 m, 21 Sept. 1996 (B. A. Huber), in author’s collection. Diagnosis: Small epigean Metagonia, distinguished from congeners by details of male and female genitalia (Figs 85-86, 89-92), and especially the male chelicerae with a unique row of modified hairs on each side (Figs 87-88). Note: In the original description (Gertsch 1986) the species is characterized as troglophile. The new records show that it is a typical epigean species instead (also the original record was not from within a cave!). Distribution: Known from type locality and the three new localities above. PHOLCIDS FROM GUATEMALA AND HONDURAS Tal DESCRIPTION Male: Prosoma and legs pale ochre yellow, without dark marks (Figs 83-84), patellae slightly darker, tibia-metatarsus joints dark. Opisthosoma without spots. Pedipalps as shown in Figs 85-86. Palpal femur with ventral hump (Figs 85-86). Chelicerae with highly distinctive row of modified hairs on each side and a blunt projection more proximally (Figs 87-88). Clypeus unmodified. Measurements of a male from Poptun: Total length: 2.4; prosoma width: 0.73; length: 0.8; opisthosoma length: 1.6; legs: 1 2 3 4 fem I 3.6 2.4 3.6 pat 0.4 0.4 0.3 0.4 tib 5.4 3.5 2.0 39 met 9.0 5.3 3.0 4.6 tar 1.5 0.9 0.7 0.7 total DM) 13.7 8.4 125 tibind 74 52 32 48 REDESCRIPTION Female: Colors as in male. Epigynum pale, with distinctive black knob (Figs 90-91). Internal genitalia as in Fig. 92. Measurements of female holotype: prosoma width: 0.75; length: 0.7; (opistho- soma damaged); leg 3: fem: 2.1, pat: 0.3, tib: 1.7, met: 2.5, tar: 0.6 (other legs missing). Measurements of a female from Poptun: total length: 2.5; prosoma width: 0.73: length: 0.8; opisthosoma length: 1.7; legs: 1 DI 3 4 fem 4.3 3] DD 3.2 pat 0.4 0.4 0.3 0.3 tib 4.3 2.8 1.8 2.8 met 7.0 4.3 2.6 4.1 tar 1.3 0.9 0.6 0.7 total 17.3 11.5 TS 11.1 tibind 61 44 29 40 Variation: Tibia | in other material: Poptun: 5 males 4.9-5.4, x=5.2; 5 females: 3.8- 4.3, x=4.1. Cerro Cahui: 5 males: 4.5-5.4, X=4.9; 6 females: 3.6-4.0, x=3.8. Tikal: 6 males: 4.4-5.5, x=5.0; 2 females: 4.1, 4.2. Habitat: Underside of large leaves. 78 BERNHARD A. HUBER ped B127079-0.81,EEW1R131.222ET0.5-0.557M 1465,67 4227812212 POR OOK OI SMOSH OSs 08 M7 2 0) CS" MATERIAL EXAMINED. Holotype d, Israel: Galilee, Hula, 25.04.1982, Besuchet, Lob] (MHNG). Paratypes. 26 , 49, same data as holotype (JFCG, MHNG). COMMENTS. Scopaeus minimus and S. palaestinus may be distinguished from other species by the characteristic shape of the aedeagus, of which the dorsal lobe is right- angled bent ventrally in the basal portion and divided into two long spines, which are pointing proximally toward the phallobase, and by the spermatheca having a distinctly elongated chamber and a sclerotized ductus inserted distally. Both species are placed in the S. minimus group, which is distributed in Central Europe, South-East Europe, Anatolia and in Middle East. Scopaeus ryei Wollaston (Figs 80-82, 89, 97, 100, 103, 109) Scopaeus ryei Wollaston, 1872: 34. Lectotype d, England, Devon, Slapton Ley, Wollaston (BMNH); here designated (examined). Scopaeus (Hyposcopaeus) ryei; COIFFAIT 1968: 419. Scopaeus (Stilpon) baudrimonti Coiffait, 1952: 6. Holotype ©, France, Hautes-Pyrénées, Pragnères, 16.04.1945, Tempère (MNHN); examined. Syn. n. Scopaeus (Polyodontus) jarrigei Coiffait, 1953: 267. Holotype d , France, Indre, Chateauroux, 1939, Coiffait (MNHN); examined; synonymised by COIFFAIT 1968: 419. Scopaeus (Hyposcopaeus) jarrigei; COIFFAIT 1960: 285. Scopaeus (Polyodontus) minimus forcipis Ochs, 1954: 65. Holotype ¢, France, Alpes Mari- times, Pré du Lac, 03.12.1947, Ochs (MHNG); examined; synonymised by COIFFAIT 1968: 419. DESCRIPTION. Length 2.5-2.9 mm; forebody 1.3-1.5 mm. Body yellowish-brown to brown, disc of elytra sometimes slightly darker. Abdomen light brown to dark brown, appendages light yellowish-brown. Surface dull, head and pronotum with dense reti- culation, puncturation very fine and dense, notably coarser on elytra. Tempora slightly enlarged, head with strongly rounded hind angels and straight posterior mar- gin. Eyes very small, 0.40-0.47 of temporal length. Pronotum relatively slender, 0.23 up to 0.30 times longer than wide. Elytral lateral length up to 1.2 times as long as pronotal length, sutural length about a tenth shorter than latter. Metathoracic wings entire. Protarsomeres 1-4 conspicuously slender, in both sexes slightly wider than long. Mesotibia moderately enlarged, about 5-6 times as long as wide. Laterotergite 9 102 JOHANNES FRISCH (fig. 97) with strong dorsal tooth. Tergite 10 (fig. 100) and valve (fig. 103) relatively slender. Posterior margin of male sternite 8 (fig. 89) with very shallow, arcuate emargination in distal 1/14 and smooth posterio-median area surrounded by median pointing setae. Aedeagus (figs 80-82) large and robust with bloated phallobase. Apical lobes distinctly shortened and slender, less than half as long as dorsal lobe, and pointing ventrally. Dorsal lobe deeply divided into two processes with basal portions bent ventrally, and distal halves widened and pointing longitudinally. Lateral lobes prominent and also orientated ventrally, each with a group of numerous, long setae. Ventral endophallic process in distal half deeply divided into two spines, which are right-angled bent distally in lateral view. A long flagellar spine reaching apex of apical lobes. Spermatheca as in fig. 109. RATIOS. HLW 1.13-1.2; PLW 1.23-1.3; HPW 1.09-1.2; HPL 1.02-1.09; PSL 0.95- 1.14; PLL 0.79-0.93; ELW 1.13-1.24; ET 0.4-0.47; MT 5.0-6.1; A 2.2, 1.2, 1.3, 1.2, Ip LOOSE O85 O25 OOS los 1D AUP WES) x MATERIAL EXAMINED (380 specimens). Austria: Burgenland (MKCH, NHMW, SMTD); Carinthia (MHNG); Tyrol (BKCB, TLMF, JFCG, MKCH, MSCB, NHMW); Upper Austria (NHMW); Vienna (NHMW); Vorarlberg (NHMW). Bosnia-Hercegovina (NHMW, SMTD, VACH). Czech Republic: Jihomoravsky Kraj (HNHM, NHMW, NMPC); Severomoravsky Kraj (NHMW). England: lectotype 4, Devon, Slapton Ley (BMNH). France: holotype d of S. jarrigei, Indre, Chateauroux (MNHN); holotype d of S. minimus forcipis, Alpes Maritimes, Pré du Lac (MHNG); holotype ® of S. baudrimonti, Hautes-Pyrénées, Pragnères (MNHN); Bouches-du-Rhône (MHNG); Gers (ISNB); Haute-Saône (MHNG); Rhône (NHMW); Var (DEIC, MHNG); Vaucluse (HNHM); Vienne (NHMW). Germany: Bavaria (ZMHB, ZSMC); Hesse (VACH). Hungary: Györ-Moson-Sopron (MHNG). Italy: Emilia Romagna (ZMHB); Friuli-Venezia Giulia (MKCH); Lombardia (NHMW); Piedmont (MCSN); Trentino-Alto Adige (TLMF, JFCG, MHNG, MKCH, MSCB, NHMW, SMTD). Poland: Bielsko-Biala (ZMHB). Romania: Caras-Sevérin (HNHM). Slovakia: Stredoslovensky Kray (NHMW, SMTD). Spain: Aragon (NHMW). Switzerland: Bern (JFCG); Fribourg (MHNG); Genève (MHNG); Valais (TLMF). DISTRIBUTION. Scopaeus ryei is an European species distributed from South England (Devon, type locality) throughout France, Switzerland, Central and South Germany and Austria southwards to North-East Spain (Aragon), the Abruzzese Mountains (BINAGHI 1935) and Bosnia-Hercegovina. Easternmost data are from South Poland, Slovakia and West Romania. The record from Denmark (JOHANSEN 1914), overtaken by PALM (1963) and SILFVERBERG (1992), is doubtful. HABITAT. Scopaeus ryei is a thermo-hygrophilous inhabitant of damp, sandy margins of rivers, streams or lakes with gravel and poor vegetation (BOHAC 1985; SCHATZ 1996). It occurs also in secundary biotopes such as gravel pits (PEEZ & KAHLEN 1977). British authors (EDMONDs 1931; HYMAN & PARSON 1994) recorded the species from coastal shingle. COMMENTS. Scopaeus ryei was first mistaken for S. minimus by NEWBERRY (1914), followed by BINAGHI (1935), who described the aedeagus, and subsequent authors (e.g. BOHAC 1985; CoIFFAIT 1968, 1984; LOHSE 1964; LOHSE & LUCHT 1989). Thus, the data in HORION (1965) on S. minimus refer rather to S. ryei. FOWLER (1888) hold S. ryei for S. rubidus, restricted to South-West Europe, and HEYDEN et al. (1906) WEST PALAEARCTIC SPECIES OF SCOPAEUS 103 synonymised it with S. minutus. EDMONDS (1931) revalidated S. ryei. The confusion grew when OCHS (1954) described S. minimus forcipis based on a male of which the apical lobes of the aedeagus are bent out of shape and running longitudinally along the phallobase. CoIFFAIT (1968) synonymised this name with S. ryei, but later (COIFFAIT 1984) distinguished falsely S. rvei with longer apical lobes, which in fact are bent out of shape, and S. minimus with undamaged aedeagus. Obviously, COIFFAIT (1984) illustrated the damaged aedeagus of S. minimus forcipis, which he falsely identified as that of S. ryei. BOHAC (1985) and LOHSE & LUCHT (1989) followed COIFFAIT (1984). See also comments under S. minimus. Scopaeus brevicuspis Binaghi (Figs 83-85, 90, 104, 110) Scopaeus (Polyodontus) brevicuspis Binaghi, 1935: 102. Holotype 6, Italy, Sardinia, Cagliari, 03.06.1901, Dodero (MCSN); examined. Scopaeus (Hyposcopaeus) brevicuspis; COIFFAIT 1960: 285. DESCRIPTION. Length 2.5-2.9 mm; forebody 1.4-1.6 mm. Body brown, pronotum sometimes slightly lighter, elytra, except hind margin and posterior half of suture, darker, abdomen blackish. Appendages brown. Forebody notably shining, punctu- ration relatively fine and dense, reticulation indistinct. Head with slightly enlarged tempora, strongly rounded hind angles and straight posterior margin. Eyes about as long as tempora. Elytra relatively long, lateral length exceeding pronotal length by a tenth up to a quarter, sutural length slightly exceeding latter, or up to a tenth shorter. Metathoracic wings entire. Protarsomeres 1-4 in both sexes twice as wide as long. Mesotibia relatively slender. Antennomeres slender, not wider than long, except slightly transverse segment 10. Laterotergite 9 as in S. littoralis (fig. 96), with strong dorsal tooth. Valve (fig. 104) relatively slender. Sternite 8 in male (fig. 90) with triangular emargination in distal sixth with lateral margins moderately convex. Aedeagus (figs 83-85) with apical lobes broad and distinctly shortened, only a third as long as dorsal lobe, truncate in lateral view, each bearing a lateral group of long setae. Dorsal lobe deeply divided, almost right-angled bent ventrally, gradually widened toward apex and truncate apically, ventro-proximal angle bearing a short dent. Lateral lobes short, truncate, pointing ventrally, each with a group of long setae. Ventral endophallic process hook-shaped, but evenly arcuate longitudinally, slender and car- ved at apex in ventral view. Endophallic flagellum extended far over dorsal lobe. Process of spermatheca (fig. 110) slender and bent at apex, chamber triangular. Ratios. HLW 1.12-1.2; PLW 1.16-1.28; HPW 1.04-1.11; HPL 1.0-1.05; PSL 0.93- (PIG RIE OSEO SEEN ITA 25) El 048-0572 Mills. 3-5.97A 2.1, ed eS. eS. PS2 HO SalO = 1050295 16-9 V (00168 MATERIAL EXAMINED (46 specimens). Algeria: Annaba (MHNG). France: Corse-du-Sud (DEIC, HNHM); Haute-Corse (DEIC, MHNG). Italy: holotype &, Sardinia, Cagliari (MCSN); para- types 16, 19, Sicily, Pachino (MCSN); Sardinia MHNG, NHMW, SMTD); Sicily (NHMW). DISTRIBUTION. Scopaeus brevicuspis occurs in the West Mediterranean region. Examined material is from Corsica, Sardinia, Sicily and Algeria. According to BINAGHI (1935), it occurs also in Tunisia. The record of S. minimus from Corsica and Sardinia (PORTA 1926) refers obviously to S. brevicuspis. 104 JOHANNES FRISCH Scopaeus littoralis Ochs (Figs 86-88, 91, 96, 105, 111) Scopaeus (Polyodontus) littoralis Ochs, 1958: 276. Holotype d, France, Var, St. Aygulf, 11.1957, Ochs (MHNG); examined. Scopaeus (Hyposcopaeus) littoralis; COIFFAIT 1960: 285. DESCRIPTION. Length 2.4-2.9 mm; forebody 1.3-1.4 mm. Body light brown to brown, abdomen and elytra, except humeral callus, hind margin and posterior half of suture, dark brown or blackish, appendages light brown. Forebody shining, lacking reticu- lation, puncturation clear. Head and pronotum relatively slender. Head with slightly enlarged tempora, strongly rounded hind angles and straight posterior margin. Eyes half as long as tempora or somewhat shorter. Elytral lateral length exceeding pronotal length by a fifth, sutural length as pronotal length or up to a tenth shorter. Mesothoracic wings entire. Protarsomeres 1-4 slender, in both sexes almost twice as wide as long. Mesotibia notably enlarged, about five times longer than wide. Dorsal margin of laterotergite 9 (fig. 96) with strong dorsal tooth, valve relatively slender (fig. 105). Sternite 8 in male (fig. 91) as in S. brevicuspis (fig. 90), but triangular emargination with proximal angle more obtuse. Aedeagus (figs 86-88) with short, slender apical lobes, which are pointing ventrally, truncate at apex and bent toward each other in ventral view. Dorsal lobe projecting far over apical lobes, deeply divided into two ventrally curved processes by a nearly semicircular incision. Processes each at apex enlarged into a right-angled distal projection and a long proximal hook. Endophallic flagellum extended far over dorsal lobe. Lateral lobes strongly reduced, each bearing a group of short setae, which is pointing ventrally. Phallobase with an additional, median group of few setae. Ventral endophallic process reduced to two short portions. Spermatheca (fig. 111) small, with strongly curved, slender portions. Ratios. HLW 1.14-1.22; PLW 1.24-1.29; HPW 1.07-1.18; HPL 1.0-1.12; PSL 1.0- [ell PIEIL0:79-0:86; ELW 115-123; ET 0.45-0.5; MT 4554 AVS MEZZA SIE LOIERO O00 ONO" 1720 12°979,©),6.8: MATERIAL EXAMINED (16 specimens). France: holotype d and paratypes 4d, 89, Var, St. Aygulf (MHNG): Gard (MHNG); Var (JFCG, MHNG). DISTRIBUTION. The distribution pattern of Scopaeus littoralis is unknown. The presumed West Mediterranean species is confirmed only from Var and Gard in South France. COMMENTS. Scopaeus ryei, S. brevicuspis and S. littoralis share characteristic external and aedeagal features and are combined as S. ryei group, which occurs in Central Europe, Western Europe and in the western Mediterranean region. Members of the S. ryei group may be distinguished by the aedeagus having reduced, ventrally bent apical lobes and a remarkable dorsal lobe, which is deeply divided and of which processes are also pointing in ventral direction. The aedeagus is furthermore featured by ventrally orientated lateral lobes with long setae and by a long endophallic flagellum, which is projecting far over dorsal lobe. The species of the S. ryei group are also linked in having slender protarsomeres, which are less widened than in most Scopaeus species, and share a slender head with slightly widened tempora and strongly rounded posterior angles. Judging from external and aedeagal features, the West Mediterranean S. portai Luze, which was redescribed in FRISCH (1997b), is included as well. The species is distinguished by the lobes of the aedeagus, which are not bent ventrally. WEST PALAEARCTIC SPECIES OF SCOPAEUS 105 Scopaeus hercegovinensis sp. n. (Figs 112-115) DESCRIPTION. Length 3.2 mm; forebody 1.9 mm. Body brown, elytra in apical half gradually lighter, abdomen blackish, appendages light brown. Puncturation fine and distinct, reticulation indistinct. Head broad, almost as long as wide, with slightly enlarged tempora and straight posterior margin. Eyes relatively large, notably exceeding half length of tempora. Elytral lateral length exceeding pronotal length by almost a fifth, elytral suture as long as pronotum. Metathoracic wings entire. Protarsomeres 1-4 more than twice as wide as long. Mesotibia slender. Apical fourth of male sternite $ (fig. 115) with triangular emargination. Aedeagus with characters as in S. heinzi subgroup (FRISCH 1994). Apical lobes gradually widening toward apex, with truncate apices and slightly concave ventral margins, bearing minute setae in proximal half. Apical lobes very slender with parallel outer margins and shortly arcuate apices in dorsal view, widening toward apex in ventral view. Dorsal lobe relatively broad and parallel, reaching apex of apical lobes and bearing a long spine bent ventrally in a right angle. Ventral endophallic flagellum longitudinal. RAROSMAE WEP OGC PENA 1221 APN SI IT HIP] 40 PSET.07PEE 0:81 JEEWT23; ROD EAP OMS MES 220 EDR OO OO 0870917 MATERIAL EXAMINED. Holotype d, Bosnia-Hercegovina: Jablanica (SMTD). COMMENTS. The aedeagal features of S. hercegovinensis fit those of the S. heinzi subgroup as defined by FRISCH (1994). Scopaeus hercegovinensis is very close to S. haemusensis Frisch (figs 116, 117) from Bulgaria and S. graecus Frisch from Greece and Dalmatia (FRISCH 1994), but is unique in having a one-spined dorsal lobe. Scopaeus graecus may be easily distinguished by the three-spined dorsal lobe, and S. haemusensis may be distinguished in having a two-spined dorsal lobe and strongly convex ventral margins of the apical lobes. ACKNOWLEDGEMENTS I thank the following colleagues who generously lend specimens from the respective institutes: K. Adlbauer, IHUG; M. Baehr, ZSMC; N. Berti, MNHN; M. Brancucci, NHMB; J.D. Brendell, BMNH; E. De Boise, BMNH; J. Clary, MHNL; M. Daccordi, MLZT; R. Danielsson, MZLU; D. Drugmand, D. Haghebaert, ISNB; O. Jäger, SMTD; J. Jelinek, NMPC; M. Kahlen, TLMF; I. Löbl, MHNG; R. Poggi, MCSN; H. Schillhammer, NHMW,; G. Szél, HNHM; M. Uhlig, ZMHB and L. Zerche, DEIC. Specimens from private collections were provided by V. Assing, Hannover, C. Brandstetter, Bürs, V. Gollkovski, Berlin, A. Kapp, Bürs, C. Morkel, Butzbach, M. Schiilke, Berlin and H. Terlutter, Osnabriick. A. G. Kirejtshuk, Zoological Museum of the Academy of Sciences, St. Petersburg, provided the type specimen of Xantholinus breviventer from the Charkow University collection, Ukraine. In particular I am indebted to L. Herman, New York, I. Löbl, Geneva, and V. Puthz, Schlitz, for rviewing earlier versions of the manuskript and many helpful comments. V. Wolters, University of Gießen, kindly provided microscopes and material for my work. The present paper is part of a project toward the degree of Ph. D. at the University of Gießen, funded by the Studienstiftung des Deutschen Volkes, Bonn-Bad Godesberg. 106 JOHANNES FRISCH Fics 1-3. Scopaeus gracilis, 3, Greece, Peloponnese: aedeagus in 1) lateral, 2) ventral, 3) dorsal view. Scale bar = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 107 Fics 4-6. Scopaeus flavofasciatus, & holotype: aedeagus in 4) lateral, 5) ventral, 6) dorsal view. Scale bar = 0.1 mm. JOHANNES FRISCH 108 Fics 7-9. Scopaeus gracilis, 8 lectotype of S. trossulus: aedeagus in 7) lateral, 8) ventral, 9) dorsal view. Scale bar = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 109 Fics 10-12. Scopaeus siculus, $ lectotype: aedeagus in 10) lateral, 11) ventral, 12) dorsal view. Scale bar = 0.1 mm. 110 JOHANNES FRISCH Fics 13-23. Scopaeus siculus, 2 paralectotype: 13) laterotergite 9, 21) valve, 23) tergite 10. d lectotype: 18) sternite 8. Scopaeus flavofasciatus, 9 paratype: 14) laterotergite 9, 20) valve, 22) tergite 10. d holotype: 16) sternite 8. Scopaeus gracilis, 2 paralectotype of S. trossulus: 15) laterotergite 9, 19) valve. Scopaeus gracilis, Greece, Peloponnese: 17) 4 sternite 8. Figs 13-15, 19-23: scale bar a), figs 16-18: scale bar b), scale bars = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 111 Fics 24-29. Spermatheca: 24) Scopaeus flavofasciatus, 9 paratype. 25) Scopaeus siculus, 9 paralectotype. 26) Scopaeus gracilis, ?, Germany, Hesse. 27), 28) Scopaeus gracilis, 9, Teneriffa. 29) Scopaeus gracilis, ? paralectotype of S. trossulus. Scale bar = 0.1 mm. Fics 67-73. Spermatheca: 67), 68), 69) Scopaeus micropterus, ®, Italy, Emilia. 70) Scopaeus alaschiacus, 9 paratype. 71) Scopaeus minutoides, ®, Turkey, Antalya. 72) Scopaeus championi, 9, Austria, Lech river. 73) Scopaeus gladifer, 2 paratype. Scale bar = 0.1 mm. 112 JOHANNES FRISCH Fics 30-36. Scopaeus micropterus, à , Italy, Urbino: aedeagus in 30) lateral, 31) ventral, 32) dor- sal view. Scopaeus championi, 3, Austria, Lech river: aedeagus in 33) lateral view. d, Bosnia- Hercegovina, Sarajevo: aedeagus in 34) lateral, 35) ventral, 36) dorsal view. Scale bar = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 113 Fics 37-42. Scopaeus alaschiacus, 3 holotype: aedeagus in 37) lateral, 38) ventral, 39) dorsal view. Scopaeus gladifer, 3 holotype: aedeagus in 40) lateral, 41) ventral, 42) dorsal view. Scale bar = 0.1 mm. 114 JOHANNES FRISCH Fics 43-51. Scopaeus minutoides, $ holotype: aedeagus in 43) lateral view. d, Turkey, Istanbul: aedeagus in 44) lateral, 45) ventral, 46) dorsal view. d sternite 8: 47) Scopaeus micropterus, Italy, Urbino. 48) Scopaeus championi, Bosnia-Hercegovina, Sarajevo. 49) Scopaeus alaschiacus, holotype. 50) Scopaeus minutoides, holotype. 51) Scopaeus gladifer, holotype. Figs 43-46: scale bar a), figs 47-51: scale bar b), scale bars = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 115 Fics 52-66. Scopaeus micropterus, ®, Italy, Emilia: 52) laterotergite 9, 57) tergite 10, 62) valve. Scopaeus championi, 9, Austria, Lech river: 53) laterotergite 9, 58) tergite 10, 63) valve. Scopaeus alaschiacus, 9 paratype: 54) laterotergite 9, 59) tergite 10, 64) valve. Scopaeus gladifer, 2 paratype: 55) laterotergite 9, 61) tergite 10, 65) valve. Scopaeus minutoides, ®, Turkey, Istanbul: 56) laterotergite 9, 60) tergite 10, 66) valve. Scale bar = 0.1 mm. 116 JOHANNES FRISCH Fics 74-79. Scopaeus minimus, 6 Hungary, Neusiedler See: aedeagus in 74) lateral, 75) ventral, 76) dorsal view. Scopaeus palaestinus, 3 holotype: aedeagus in 77) lateral, 78) ventral, 79) dorsal view. Scale bar = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 117 Fics 80-85. Scopaeus ryei, 3, Austria: aedeagus in 80) lateral, 81) ventral, 82) dorsal view. Scopaeus brevicuspis, dè paratype: aedeagus in 83) lateral, 84) ventral, 85) dorsal view. Scale bar = 0.1 mm. 118 JOHANNES FRISCH Fics 86-93. Scopaeus littoralis, $ holotype: aedeagus in 86) lateral, 87) ventral, 88) dorsal view. d sternite 8: 89) Scopaeus ryei, Austria. 90) Scopaeus brevicuspis, paratype. 91) Scopaeus littoralis, holotype. 92) Scopaeus palaestinus, holotype. 93) Scopaeus minimus, Hungary, Neusiedler See. Figs 86-88: scale bar a), figs 89-93: scale bar b), scale bars = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 119 Fics 94-105. Scopaeus minimus, 9, Hungary, Neusiedler See: 94) laterotergite 9, 98) tergite 10, 101) valve. Scopaeus palaestinus, 9, paratype: 95) laterotergite 9, 99) tergite 10, 102) valve. Scopaeus littoralis, 2 paratype: 96) laterotergite 9, 105) valve. Scopaeus brevicuspis, 2 paratype: 104) valve. Scopaeus ryei, 2, Austria: 97) laterotergite 9, 100) tergite 10, 103) valve. Scale bar = 0.1 mm. 120 JOHANNES FRISCH Fics 106-111. Spermatheca: 106) Scopaeus minimus, 9, Hungary, Neusiedler See. 107), 108) Scopaeus palaestinus, 2 paratype. 108) Scopaeus ryei, 9, Austria. 110) Scopaeus brevicuspis, 2 paratype. 111) Scopaeus littoralis, 2 paratype. Scale bar = 0.1 mm. WEST PALAEARCTIC SPECIES OF SCOPAEUS 121 amb Fics 112-117. Scopaeus hercegovinensis sp. n., 6 holotype: aedeagus in 112) lateral, 113) ventral, 114) dorsal view, 115) sternite 8. Scopaeus haemusensis, & holotype: aedeagus in 116) lateral, 117) ventral view. Figs 112-114, 116-117: scale bar a), fig. 115: scale bar b), scale bars = 0.1 mm. 122 JOHANNES FRISCH REFERENCES BERNHAUER, M. & K. SCHUBERT 1910. Staphylinidae 1. /n: Junk, W. & S. Schenkling (eds), Coleopterorum Catalogus 19. Berlin, 86 pp. BINAGHI, G. 1935. Studio sul genere Scopaeus Erich. (Coleopt.: Staphylin.). Memorie della Socièta Entomologica Italiana 14: 84-115. BoHAc, J. 1985. Review of the subfamily Paederinae (Coleoptera, Staphylinidae) in Czecho- slovakia. Part II. 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Zur Morphologie der weiblichen Terminalia einiger Staphylinidenarten (Cole- optera). Verhandlungen des 11. SIEEC Gotha 1986: 227-237. WOLLASTON, T. V. 1864. Catalogue of the Coleopterous Insects of the Canaries in the Collec- tion of the British Museum. London, 661 pp. WOLLASTON, T. V. 1872. Scopaeus rvei n. sp. The Entomologist’s Monthly Magazine 9: 34-35. REVUE SUISSE DE ZOOLOGIE 105 (1): 125-138; mars 1998 Supplement to the knowledge of the Agathidiini of Taiwan (Coleoptera, Leiodidae) Fernando ANGELINI* & Luigi DE MARZO** * S.S.7 per Latiano, Km. 0.500, 1-72021 Francavilla Fontana (Brindisi), Italy ** Dipartimento di Biologia, Difesa e Biotecnologie agro-forestali, Università della Basilicata, Via Nazario Sauro 85, I-85100 Potenza, Italy Supplement to the knowledge of the Agathidiini of Taiwan (Coleop- tera, Leiodidae). - New records and/or descriptions are given for 32 species of Agathidiini from Taiwan, collected by Dr. Ales Smetana. New species are: Agathidium (A.) subnitidum n.sp., Agathidium (A.) deli- catulum n.sp., Agathidium (A.) alesi n.sp., Agathidium (A.) pictum n.sp. New records for Taiwan are: Cyrtoplastus seriepunctatus (Bris.), Agathi- dium (Neoceble) kyotoense Ang. & Dmz, A. (Neoceble) confusum Bris., A. (A.) mequignoni Roubal. Spermatheca is figured for the first time of: Agathidium (A.) honestum Ang. & Dmz., A. (A.) meifengense Ang. & Dmz., A. (A.) egregium Ang. & Dmz. Key-words: Leiodidae - Agathidiini - Taiwan - new species - new records. INTRODUCTION This new contribution increases the number of the species of Agathidiini known from Taiwan to 60. It is based on the study of 32 species in 430 specimens of Cyrtoplastus and Agathidium, collected in 18 localities by Dr. Ales Smetana, and includes: (a) the descriptions of 4 new species: Agathidium (A.) subnitidum n.sp., Aga- thidium (A.) delicatulum n.sp., Agathidium (A.) alesi n.sp., Agathidium (A.) pictum n.sp.; (b) 4 new records for Taiwan: Cyrtoplastus seriepunctatus (Bris.), Agathidium (Neoceble) kyotoense Ang. & Dmz, A. (Neoceble) confusum Bris., A. (A.) mequignoni Roubal; (c) some female characters of 3 species known previously only from male specimens. The specimens are deposited in Geneva Museum (MHNG), Taichung Museum (NMNT) and F. Angelini's private collection (AC). Manuscript accepted 11.07.1997 126 FERNANDO ANGELINI & LUIGI DE MARZO We are indebted to Dr. Ales Smetana (Ottawa) for the opportunity of studying his very interesting material and to Dr. Ivan Löbl for comments on the manuscript of this paper. Cyrtoplastus Reitter, 1884 Cyrtoplastus smetanai Ang. & Dmz. Cyrtoplastus smetanai Angelini & De Marzo, 1995: 184. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 & and 3 © (MHNG, NMNT, AC). Discussion: These specimens agree in all characters with the types. Body length: 2.3- 2.5 mm. Distribution: Taiwan. Cyrtoplastus seriepunctatus (Brisout) Agathidium seriepunctatus Brisout in Grenier, 1867: 174. Cyrtoplastus seriepunctatus, GANGLBAUER 1899: 239; ANGELINI & DE Marzo 1988: 65: HOSHINA 1996: 204. MATERIAL: Taiwan, Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 1 ex. (AC). Discussion: This specimen differs from the European material in the shape of the aedeagal apex; in dorsal view. Distribution: Europe, Siberia, Mongolia, Japan, Taiwan. New record for Taiwan. Agathidium Panzer, 1797 Subg. Cyphoceble Thomson, 1859 Agathidium (Cyphoceble) yushanicum Ang. & Dmz. Agathidium (Cyphoceble) yushanicum Angelini & De Marzo, 1995: 186. MATERIAL: Taiwan, Nantou Hsien, Nenkaoshan, 1.5 Km SW Tenchi Hut, 2830 m, 6.V.92, 2 exx. (MHNG, AC). Distribution: Taiwan. Agathidium (Cyphoceble) geniculatum Ang. & Dmz. Agathidium (Cyphoceble) geniculatum Angelini & De Marzo, 1995: 191. MATERIAL: Taiwan, Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 21.1V.92, 1 G and 1 2 (AC); Taichung Hsien, Anmashan, 2225 m, 11.V.92, 1 2 (NMNT); Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 6 and 1 2 (MHNG); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2400 m, 20.VII.93, 1 4 (MHNG). Distribution: Taiwan. Subg. Neoceble Gozis, 1886 nigripenne group Agathidium (Neoceble) kyotoense Ang. & Dmz. Agathidium (Neoceble) kyotoense Angelini & De Marzo, 1988: 79; ANGELINI 1995: 136. SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 197, MATERIAL: Taiwan, Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 1 4 (AC): Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.VII.93, 1 5 (MHNG); Nantou Hsien, Meifeng, 2130 m, 10.VII.93, 1 2 (NMNT). Discussion: These specimens agree with the types from Japan in all characters, except one which has the dorsum black. Distribution: Japan (Honshu), Taiwan. New record for Taiwan. Agathidium (Neoceble) nigrocastaneum Ang. & Dmz. Agathidium (Neoceble) nigrocastaneum Angelini & De Marzo, 1995: 192. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 d and 2 9 (MHNG); Taichung Hsien, Anmashan, 2120 m, 13.V.92, 1 8 and 1 9 (AC); Taichung Hsien, Anmashan, 2225 m, 14.V.92, 1 ex. (NMNT). Discussion: These specimens agree fully with the types. Body length: 2.5-2.7 mm. Distribution: Taiwan. Agathidium (Neoceble) confusum Brisout Agathidium (Neoceble) confusum Brisout in GRENIER 1863: 9; HLISNIKOVSKY 1964: 90; ANGELINI 1995: 179. MATERIAL: Taiwan, Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.V11.93, 1 9 (MHNG); Kaohsiung Hsien, Peinantashan trail, 1950 m, 8.V11.93, 1 2 (AC). Discussion: These specimens agree in all characters with European material. Distribution: Europe, Caucasus, Siberia, Mongolia, Japan, Taiwan. New record for Taiwan. canariense group Agathidium (Neoceble) rufomarginatum Ang. & Dmz. Agathidium (Neoceble) rufomarginatum Angelini & De Marzo, 1995: 197. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, trail at 1500 m, 1.V.92,5 6 (MHNG, NMNT): Taichung Hsien, Anmashan, 2225 m, 11.V.92, 1 4 (MHNG); Taichung Hsien, Anmashan, creek, 2185 m, 12.V.92, 1 2 (MHNG); Taichung Hsien, Anmashan, 2120 m, 13.V.92, 1 8 (MHNG); Nantou Hsien, Meifeng, 2130 m, 10.VII.93, 2 d and 2 2 (MHNG, AC). Discussion: Agathidium rufomarginatum Ang. & Dmz. was described on the basıs of 1 male and 1 female. The additional specimens agree with the types in all characters. Body length: 2.1-2.5 mm. Distribution: Taiwan. Subg. Macroceble Angelini, 1992 Agathidium (Macroceble) oblitum Ang. & Dmz. Agathidium (Macroceble) oblitum Angelini & De Marzo, 1995: 198. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2225 m, 11.V.92, 2 2 (MHNG, AC). Distribution: Taiwan. Subg. Agathidium Panzer, 1797 madurense group 128 FERNANDO ANGELINI & LUIGI DE MARZO + Fics 1-4 Male metafemora of: 1, Agathidium subnitidum n.sp.; 2, A. delicatulum n.sp.; 3, A. alesi n.sp.; 4, A. pictum n.sp. Agathidium (Agathidium) subnitidum n.sp. task S75 17 Length 3.3-3.5 mm (holotype d: 3.4 mm). Dorsum either black or dark reddish-brown; venter reddish-brown; antennae uniformly testaceous; legs reddish- brown. Microreticulation almost absent: traceable on pronotum and elytra; punc- turation fine and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus deeply emarginate; clypeal line absent; eyes flattened. Antennal segment 3 1.8 times as long as 2 and as long as 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation absent; punctures very small, super- ficial, hardly distinct, separated from each other by 1-10 times their diameter. Pronotum: 1.3 times as broad as head, slightly broader than long (W/L = 1.33) and very convex (W/H = 1.31); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation almost absent: only traceable; punctures as large as those on head but sparser, separated from each other by 2-15 times their diameter. Holotype: length 1.20 mm, width 1.60 mm, height 1.22 mm. Elytra: Slightly narrower than pronotum, as broad as long and moderately convex (W/H = 1.8); lateral outline with very weak humeral angle. Microreticulation almost absent, only traceable; punctures as large as those on head, separated from each other by 5-15 times their diameter. Holotype: length 1.45 mm, width 1.50 mm, height 0.83 mm. Metathoracic wings absent. Meso- and metasternum: median carina absent, lateral lines absent, femoral lines complete. Legs: Male metafemora enlarged distally (fig. 1). Tarsal formula: d 5-5-4, 9 5-4-4. Male copulatory organ (figs 5-7): Aedeagus stout, with ring-like proximal part, lateral margins gently convergent towards a rounded apex, ventral piece shallowly emarginate. Parameres sinuate, broadened apically. SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 129 we Fics 5-10 Male copulatory organ (lateral view, dorsal and ventral view of apex) of: 5-7, Agathidium subnitidum n.sp.; 8-10, A. delicatulum n.sp. Scale: 1 division = 0.1 mm. 130 FERNANDO ANGELINI & LUIGI DE MARZO Spermatheca (fig. 17): Basal and ventral parts slender, differing in length. HOLOTYPE d : Taiwan, Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2135 m, 30.1V.92 (MHNG). PARATYPES: as holotype, 1 2 (MHNG), 1 d (NMNT), 1 G and I 2 (AC). Discussion: Agathidium subnitidum n.sp. is closely related to Agathidium tardum Ang. & Dmz.; it differs from the latter in the darker colour of the dorsum, the colour of the antenna, the ratio of the pronotum/head width, the absence of the metathoracic wings and the characters of the meso- and metasternum. Distribution: Taiwan. Agathidium (Agathidium) delicatulum n.sp. Figs 2, 8-10, 18 Length 2.9-3.8 mm (holotype d: 3.1 mm). Dorsum either black or dark reddish-brown; venter dark reddish-brown, mesosternum testaceous; antennae uni- formly testaceous; legs reddish-brown. Microreticulation absent from dorsum; punc- turation fine and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus moderately emarginate; clypeal line absent; eyes flattened. Antennal segment 3 1.5 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures very small, superficial, hardly distinct, separated from each other by 1-10 times their diameter. Pronotum: 1.3 times as broad as head, slightly broader than long (W/L = 1.31) and very convex (W/H = 1.45): anterior margin sharply curved; lateral outline broadly rounded. Punctures as large as those on head. Holotype: length 1.10 mm, width 1.45 mm, height 1.00 mm. Elytra: Moderately narrower than pronotum, as broad as long and moderately convex (W/H = 1.8); lateral outline with very weak humeral angle. Punctures as large as those on head. Holotype: length 1.40 mm, width 1.35 mm, height 0.75 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines complete. Legs: Male metafemora enlarged distally (fig. 2). Tarsal formula: & 5-5-4, 9 5-4-4. Male copulatory organ (figs 8-10): Aedeagus slender, with ring-like proximal part, lateral margins sinuously convergent towards a subacute apex, ventral piece deeply emarginate. Parameres slender, gently narrowing towards apex. Spermatheca (fig. 18): Basal part pear-shaped; apical part short. HOLOTYPE dé: Taiwan, Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93 (MANG). PARATYPES: as holotype, | d and 1 2 (MHNG), | 6 and 1 2 (AC); same but 2390-2490 m, 5.VII.93, 1 6 and 3 2(MHNG), 1 d and 1 £ (NMNT), 1 6 and 1 2 (AC); Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 22.VII.93, 1 4 and 3 2 (MHNG). Discussion: Agathidium delicatulum n.sp. is closely related to Agathidium alesi n.sp. and A. distinguendum Ang. & Dmz.; it differs from them in the shape of the male metafemora and ratio of the 3rd/2nd antennal segments. The three species can be dis- tinguished by the male copulatory organ. Distribution: Taiwan. SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 131 Agathidium (Agathidium) alesi n.sp. Figs 3, 11-13, 19 Length 3.0-4.1 mm (holotype d: 4.1 mm). Dorsum black, mesosternum red- dish-brown, metasternum darker; antennae uniformly testaceous; legs reddish-brown. Microreticulation absent from dorsum; punctures small and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus deeply emarginate; clypeal line absent; eyes flattened. Antennal segment 3 1.3 times as long as 2 and longer than segments 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures very small and superficial, separated from each other by 2-10 times their diameter. Pronotum: 1.4 times as broad as head, slightly broader than long (W/L = 1.25) and very convex (W/H = 1.45); anterior margin sharply curved; lateral outline broadly rounded. Punctures as large as those on head. Holotype: length 1.40 mm, width 1.75 mm, height 1.20 mm. Elytra: Moderately narrower than pronotum, slightly longer than broad (W/L = 0.8) and moderately convex (W/H = 1.68); lateral outline with very weak humeral angle. Punctures very small, hardly distinct, separated from each other by 2-15 times their diameter. Holotype: length 2.00 mm, width 1.60 mm, height 0.95 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines incomplete; a small tubercle between the metacoxae. Legs: Male metafemora with a pronounced tooth at the posterior margin (fig. 5) slarsaliformula: d).5-5-4, 95-44 Male copulatory organ (figs 11-13): Aedeagus very slender, with a hook-like proximal part, lateral margins sinuously converging to rounded apex, ventral piece shallowly emarginate. Parameres slender, abruptly reflexed near apex. Spermatheca (fig. 19): Basal and ventral parts slender, differing in length. HOLOTYPE d: Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5. VII.93 (MHNG). PARATYPES: as holotype, 5 2 (MHNG), 1 d and 1 2 (NMNT), 3 d and 2 9 (AC); same but 2500 m, 4.VII.93, 4 4 (MHNG); Nantou Hsien, Nenkaoshan Tenchi Hut, 2895 m, 7.V.92, 1 6 (MHNG). Discussion: See under A. delicatulum n.sp. Agathidium alesi n.sp. is most closely related to A. distinguendum Ang. & Dmz.; it differs in the pronotal shape and the aedeagal apex. Distribution: Taiwan. Agathidium (Agathidium) ? distinguendum Ang. & Dmz. Agathidium (s.str.) distinguendum Angelini & De Marzo, 1995: 202. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2125 m, 27.IV.92, 1 2 (MHNG). Discussion: Our identification of this female is reported as doubtful because of the difficulties in distinguishing the species of the madurense group without examination of the male copulatory organ. Distribution: Taiwan. 132 FERNANDO ANGELINI & LUIGI DE MARZO 11 14 LU ASS | 15 16 Fics 11-16 Male copulatory organ (lateral view, dorsal and ventral view of apex) of: 11-13, Agathidium alesi n.sp.; 14-16, A. pictum n.sp. Scale: 1 division = 0.1 mm. SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 133 Agathidium (Agathidium) familiare Ang. & Dmz. Agathidium (s.str.) familiare Angelini & De Marzo, 1995: 203. MATERIAL: Taiwan, Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2650 m, 21.1V.92, 1 3 and 2 2 (MHNG); Taichung Hsien, Anmashan, 2225 m, 11.V.92, 1 4 (MHNG); Taichung Hsien, Anmashan, 2230 m, 12.V.92, 2 8 and 1 2 (MHNG, AC); Taichung Hsien, Anmashan, 2225 m, 14.V.92, 1 d and 1 2 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 3 d and 4 2 (MHNG, AC); Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.VII.93, 3 6 and 8 2 (MHNG, NMNT); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2400 m, 20.VII.93, 1 d and 2 2 (MHNG, AC). Distribution: Taiwan. seminulum group Agathidium (Agathidium) mequignoni Roubal Agathidium (s.str.) mequignoni Roubal, 1911: 49; HLISNIKOVSKY 1964: 221; ANGELIN 1995: 278. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2150 m, 13.V.92, 1 & (AC). Discussion: This specimen agrees in all characters with the type material from Cau- casus. Distribution: Caucasus, Turkey, North Iran, Kirgizstan, Siberia, Korea, Taiwan. New record for Taiwan. laevigatum group Agathidium (Agathidium) honestum Ang. & Dmz. Fig. 20 Agathidium (s.str.) honestum Angelini & De Marzo, 1995: 208. MATERIAL: Taiwan, Kaohsiung Hsien, Kuanshan, Kunanoshing Hut, 3020 m, 18.1V.92, 1 6 (MHNG); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2130 m, 27.IV.92, 2 d and 2 © (MHNG, NMNT); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2125 m, 27.1V.92, 3 4 and 4 ? (MHNG, NMNT, AC); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2135 m, 30.IV.92, 3 ¢ and 1 2 (MHNG, AC); Kaohsiung Hsien, Peinantashan trail, ridge at 2800 m, 3.VII.93, 1 G (AC); Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 1 4 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.VII.93, 2 & (AC). Discussion: Agathidium honestum Ang. & Dmz. was described on the basis of 3 males. The additional specimens agree in all characters with the types. Female tarsal formula: 5-4-4. Spermatheca as in fig. 20. Body length: 2.6-2.9 mm. Distribution: Taiwan. Agathidium (Agathidium) lucidum Ang. & Dmz. Agathidium (s.str.) lucidum Angelini & De Marzo, 1995: 208. MATERIAL: Taiwan, Nantou Hsien, Nenkaoshan trail, Yuenhal Hut, 2350 m, 4.V.92, 1 d and 2 2 (MHNG); Nantou Hsien, Nenkaoshan Tenchi Hut, 2900 m, 5.V.92, 3 6 and I 9 (MHNG); Nantou Hsien, Nenkaoshan, 2.5 Km SW Tenchi Hut, 2720 m, 6.V.92, 11 6 and 6 © (MHNG, NMNT, AC); Ilan Hsien, Taipingshan, 1950 m, 13.VII.93, 1 & (MHNG). Distribution: Taiwan. 134 FERNANDO ANGELINI & LUIGI DE MARZO 17 20 21 da Fics 17-22 Spermatheca of: 17, Agathidium subnitidum n.sp.; 18, A. delicatulum n.sp.; 19, A. alesi n.sp.; 20, A. honestum Ang. & Dmz.; 21, A. meifengense Ang. & Dmz.; 22, A. egregium Ang. & Dmz. Scale: | division = 0.1 mm. Agathidium (Agathidium) tenebroides Ang. & Dmz. Agathidium (s.str.) tenebroides Angelini & De Marzo, 1995: 210. MATERIAL: Taiwan, Kaohsiung Hsien, Kuanshan, Kunanoshing Hut, 3020 m, 18.IV.92, 1 4 (MHNG); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2400 m, 20.1V.92, 2 à and 1 £ (MHNG); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2650 m, 21.1V.92, 4 d and 10 2 (MHNG, NMNT, AC); Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 21.1V.92, 1 & (MHNG); Kaohsiung Hsien, Peinantashan trail, 2400 m, 4. VII.93, 1 d (AC); Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 2 & and 3 2 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.VII.93, 9 d and 6 2 (MHNG, NMNT, AC); Kaohsiung Hsien, Peinantashan trail, 2080 m, 6.VII.93, 1 3 and 1 2 (MHNG); Kaoh- siung Hsien, Peinantashan trail, 2020 m, 7.VII.93, 2 4 (MHNG); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2400 m, 20.VII.93, 1 4 (MHNG); Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 22.V11.93, 3 & (MHNG). Distribution: Taiwan. Agathidium (Agathidium) meifengense Ang. & Dmz. eve, ZI Agathidium (s.str.) meifengense Angelini & De Marzo, 1995: 213. MATERIAL: Taiwan, Taichung Hsien, Anmashan, creek, 2185 m, 12.V.92, 1 d and 2 © (MHNG, NMNT); Taichung Hsien, Anmashan, 2230 m, 12.V.92, 3 8 and 4 9? (MHNG, AC); Taichung Hsien, Anmashan, 2120 m, 13.V.92, 1 d (MHNG). SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 13 Un Discussion: Agathidium meifengense Ang. & Dmz. was described on the basis of 4 males. Female tarsal formula: 5-4-4. Spermatheca as in fig. 21. These specimens agree in all characters with the types. Body length: 3.9-4.2 mm. Distribution: Taiwan. Agathidium (Agathidium) kaohsiungense Ang. & Dmz. Agathidium (s.str.) kaohsiungense Angelini & De Marzo, 1995: 217. MATERIAL: Taiwan, Kaohsiung Hsien, Peinantashan trail, 2080 m, 6.VII.93, 2 & and 2 ? (MHNG, AC); Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 22.VII.93, 2 8 and 2 2 (MHNG, NMNT, AC). Distribution: Taiwan. Agathidium (Agathidium) fuliginosum Ang. & Dmz. Agathidium (s.str.) fuliginosum Angelini & De Marzo, 1995: 221. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, ridge at 1288-2910 m, 28.IV.92, 4 6 (MHNG, NMNT, AC). Discussion: Agathidium fuliginosum Ang. & Dmz. was described on the basis of 1 male. Female remains unknown. The additional specimens agree in all characters with the type. Body length: 3.4-3.7 mm. Distribution: Taiwan. Agathidium (Agathidium) alpestre Ang. & Dmz. Agathidium (s.str.) alpestre Angelini & De Marzo, 1995: 223. MATERIAL: Taiwan, Nantou Hsien, Nenkaoshan, 2.5 Km SW Tenchi Hut, 2720 m, 6.V.92, 2 & and | 2 (MHNG, NMNT); Nantou Hsien, Nenkaoshan Tenchi Hut, 2895 m, 7.V.92, 1 8 and 1 2 (MHNG); Nantou Hsien, Nenkaoshan trail, 2050-2150 m, 8.V.92, 3 & and 2 2 (MHNG, AC). Discussion: Apart from some specimens being lighter coloured, this new material agrees in all characters with the types. Body length: 3.0-3.3 mm. Distribution: Taiwan. Agathidium (Agathidium) comptum Ang. & Dmz. Agathidium (s.str.) comptum Angelini & De Marzo, 1995: 227. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, trail at 1500 m, 1.V.92, 1 & (MHNG). Further two specimens are doubtfully referred to the same species: Kaohsiung Hsien, Kuanshan, Kunanoshing Hut, 3020 m, 18.1V.92, 1 4 (AC): Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 21.1V.92, 1 4 (MHNG). Discussion: The two doubtful specimens differ slightly from the types in the shape of the aedeagus. Distribution: Taiwan. Agathidium (Agathidium) egregium Ang. & Dmz. Fig. 22 Agathidium (s.str.) egregium Angelini & De Marzo, 1995: 228. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2225 m, 11.V.92, 1 & (MHNG): Taichung Hsien, Anmashan, creek, 2185 m, 12.V.92, 2 d and 2 2 (MHNG, NMNT, AC); 136 FERNANDO ANGELINI & LUIGI DE MARZO Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 6 and 1 9 (AC); Taichung Hsien, Anmashan, 2225 m, 14.V.92, 1 & (MHNG); Ilan Hsien, Taipingshan, 1950 m, 13.VIL.93, 2 & (MHNG). Discussion: Agathidium egregium Ang. & Dmz. was described on the basis of 1 male. The additional specimens agree in all characters with the types, except for the colour of some of them, which are either darker or lighter dorsally. Female tarsal formula: 5- 4-4. Spermatheca as in fig. 22. Body length: 2.3-2.6 mm. Distribution: Taiwan. Agathidium (Agathidium) anmashanense Ang. & Dmz. Agathidium (s.str.) anmashanense Angelini & De Marzo, 1995: 228. MATERIAL: Taiwan, Nantou Hsien, Nenkaoshan trail, Yuenhal Hut, 2350 m, 4.V.92, 1 3 and 1 2 (MHNG); Nantou Hsien, Nenkaoshan Tenchi Hut, 2900 m, 5.V.92, 3 d and 6 9 (MHNG, AC); Nantou Hsien, Nenkaoshan, 2.5 Km SW Tenchi Hut, 2720 m, 6.V.92, 8 6 and 5 © (MHNG, AC); Nantou Hsien, Nenkaoshan, 1.5 Km SW Tenchi Hut, 2830 m, 6.V.92, 4 & and 3 2 (MHNG, AC); Nantou Hsien, Nenkaoshan trail, 2050-2150 m, 8.V.92,5 d and 3 © (MHNG, NMNT). Distribution: Taiwan. Agathidium (Agathidium) subalpinum Ang. & Dmz. Agathidium (s.str.) subalpinum Angelini & De Marzo, 1995: 230. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, trail at 1500 m, 1.V.92, 6 © (MHNG, NMNT, AC). Distribution: Taiwan. Agathidium (Agathidium) exoletum Ang. & Dmz. Agathidium (s.str.) exoletum Angelini & De Marzo, 1995: 236. MATERIAL: Taiwan, Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2130 m, 27.1V.92, 2 4 (MHNG); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2125 m, 27.1V.92, 1 4 (MHNG); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2135 m, 30.1V.92, 1 ¢ and 1 2 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 4 & and 6 © (MHNG, NMNT, AC). Discussion: The additional specimens agree in all characters with the types, except for the lighter colour of some of them. Body length: 3.2-3.5 mm. Distribution: Taiwan. dentatum group Agathidium (Agathidium) pictum n.sp. Figs 4, 14-16 Length 4.0 mm (holotype d and paratype). Dorsum black, venter reddish- brown, meso- and metasternum lighter; antennae uniformly testaceous; legs reddish- brown. Microreticulation absent on entire dorsum; punctures very small on head and pronotum, absent from elytra. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus deeply emarginate; clypeal line absent; eyes flattened. Antennal segment 3 1.6 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th SUPPLEMENT OF THE AGATHIDIINI OF TAIWAN 137 and 10th antennal segments. Punctures very small and superficial, not very distinct, separated from each other by 4-6 times their diameter. Pronotum: 1.3 times as broad as head, slightly broader than long (W/L = 1.2) and very convex (W/H = 1.28); anterior margin sharply curved; lateral outline broadly rounded. Punctures hardly distinct, separated from each other by 1-10 times their diameter. Holotype: length 1.50 mm, width 1.80 mm, height 1.40 mm. Elytra: Moderately narrower than pronotum, as broad as long and moderately convex (W/H = 1.7); lateral outline with very weak humeral angle. Puncturation, except for few very small punctures, absent. Holotype: length 1.65 mm, width 1.70 mm, height 1.00 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines incomplete; a small tubercle between metacoxae. Legs: Male metafemora with a pronounced tooth at posterior margin (fig. 4). Tarsal formula: d 5-5-4, 2 not known. Male copulatory organ (figs 14-16): Aedeagus very slender, with hook-like proximal part, lateral margins converging to small subacute tip, ventral piece shallowly emarginate. Parameres slender, bent up at apex. HOLOTYPE d : Taiwan, Taichung Hsien, Anmashan, 2225 m, 11.V.92 (MHNG). PARATYPE: as holotype, 1 d (AC). Discussion: A. pictum n.sp. is closely related to A. umbratum Ang. & Dmz. and A. fenestratum Ang. & Dmz.; it differs in the shape of the male hind femora and the ratio of the 3rd/2nd antennal segments. The aedeagus is very similar in these species but the shape of the male hind femora is characteristic. Distribution: Taiwan. Agathidium (Agathidium) splendidulum Ang. & Dmz. Agathidium (s.str.) splendidulum Angelini & De Marzo, 1995: 245. MATERIAL: Taiwan, Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2650 m, 21.1V.92, 8 G and 12 2 (MHNG, NMNT, AC); Pingtung Hsien, Peitawhushan, Kuai-Ku Hut, 2130 m, 27.1V.92, 1 4 (MHNG); Pingtung Hsien, Peitawhushan, ridge at 1288-2910 m, 28.1V.92, 3 G and 1 2 (MHNG); Pingtung Hsien, Peitawhushan, above Kuai-Ku Hut, 2680 m, 29.IV.92, 4 8 and 3 2 (MHNG); Nantou Hsien, Nenkaoshan Tenchi Hut, 2900 m, 5.V.92, 1 4 and 1 2 (MHNG); Nantou Hsien, Nenkaoshan, 2.5 Km SW Tenchi Hut, 2720 m, 6.V.92, 2 d (MHNG); Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 d (MHNG); Taichung Hsien, Anmashan, 2225 m, 14.V.92, 1 6 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2400 m, 4.VIL.93, 3 4 and 2 2 (MHNG); Kaohsiung Hsien, Peinantashan trail, 2500 m, 4.VII.93, 7 & and 10 2 (MHNG, AC); Kaohsiung Hsien, Peinantashan trail, 2390-2490 m, 5.VII.93, 13 & and 14 2 (MHNG, NMNT, AC); Nantou Hsien, Meifeng, 2130 m, 10.VII.93, 1 & (MHNG); Kaohsiung Hsien, Kuanshan, trail at Kaunshanchi Riv., 2400 m, 20.VII.93, 1 é (MHNG); Kaohsiung Hsien, Kuanshan, trail above Kaunshanchi Riv., 2550 m, 22.VII.93, 8 & and 2 2 (MHNG, NMNT, AC). Discussion: Agathidium splendidulum Ang. & Dmz. was described on the basis of 4 males. The additional specimens are variable in puncturation and colour. Body length: 3.5-3.9 mm. Distribution: Taiwan. Agathidium (Agathidium) melanocephalum Ang. & Dmz. Agathidium (s.str.) melanocephalum Angelini & De Marzo, 1995: 246. 138 FERNANDO ANGELINI & LUIGI DE MARZO MATERIAL: Taiwan, Nantou Hsien, Nenkaoshan trail, 2050-2150 m, 8.V.92, 7 & and 12 ® (MHNG, NMNT, AC). Distribution: Taiwan. Subg. Microceble Ang. & Dmz., 1986 grouvellei group Agathidium (Microceble) klapperichi Ang. & Dmz. Agathidium (Microceble) klapperichi Angelini & De Marzo, 1984 (1985): 41 (s.str.); 1986: 454; 1995: 248. MATERIAL: Taiwan, Taichung Hsien, Anmashan, 2230 m, 12.V.92, 1 8 (MHNG). Distribution: Taiwan. REFERENCES ANGELINI F., 1995. Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie del Museo Regionale di Storia naturale di Torino 18: 1-484. ANGELINI F. & L. DE Marzo, 1985. Descrizione di 13 specie nuove di Anisotomini dell'Asia sudorientale. Entomologica, Bari, 19: 23-49 (1984). ANGELINI F. & L. DE Marzo, 1986. Agathidium from India and Malaya: expedition of Geneva Natural History Museum (Coleoptera, Leiodidae). Revue suisse de Zoologie 93: 423- 455. ANGELINI F. & L. DE Marzo, 1990. Anisotomini del Giappone (Coleoptera, Leiodidae). Entomologica, Bari, 23: 47-122 (1988). ANGELINI F. & L. DE Marzo, 1995. Agathidiini from Taiwan collected by Dr. A. Smetana (Coleoptera, Leiodidae, Agathidiini). Revue suisse de Zoologie 102 (1): 175-255. BRISOUT DE BARNEVILLE CH., 1863. Matériaux pour servis à la faune des Coléoptères francais. I. In Grenier A., Paris: 1-135. GANGLBAUER L., 1899. Die Käfer von Mitteleuropa. III, Wien: 202-251 (Liodini). HLISNIKOVSKY J., 1964. Monographische Bearbeitung der Gattung Agathidium Panzer. Acta Entomologica Musei Nationalis Pragae, suppl. 5: 1-255. HOSHINA H., 1996. A taxonomic study on the genus Cyrtoplastus of Japan. Japanese Journal of systematic Entomology 2 (2): 201-206. Roubal J., 1911. Koleopterologické vysledky mé cesty na Kavkaz v cervenci r. 1911. Casopis Ceske Spolecnoski Entomologicke 8: 48-50. REVUE SUISSE DE ZOOLOGIE 105 (1): 139-220; mars 1998 Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae) Roberto PACE Via Vittorio Veneto, 13. 37032 Monteforte d’Alpone (Verona), Italia. Aleocharinae from China: Part I (Coleoptera, Staphylinidae). - In this paper, one hundred known species listed, 52 new species for China are recorded and 65 species are described as new for science. These new species, collected in very recent years by A. Smetana, G. de Rougemont and others, belong to the following tribes: Masuriini (1 species), Hypo- cyphtini (1), Myllaenini (7), Termitohospitini (1), Gyrophaenini (18), Homalotini (27), Bolitocharini (6), Sahlbergini (1) and Eusteniamorphini (3). Four genera are described as new, assigned to following tribes: Homalotini (Taraktomora n. gen., near Silusa), Bolitocharini (Omologlusa n. gen., Neoleptusa; Methistemistiba n. gen., near Leptusa) and Sahlbergini (Derougemontius n. gen., near Loeblius). Eight new combination and five new synonymies are proposed. The major diagnostic caracters are illus- trated. The species of Aleocharinae taken in China again reflect the mixture of Palaearctic and Oriental elements. Key-words: Coleoptera - Staphylinidae - Aleocharinae - Taxonomy - China. INTRODUZIONE Il catalogo delle specie della sottofamiglia Aleocharinae della Cina finora comprendeva solamente 147 specie. Data la vastità e la varietà del territorio cinese, ciò non poteva che essere il segno tangibile che ricerche in questo campo sono ancora agli inizi. Grazie a recentissime ricerche effettuate dal Dr. Ales Smetana del “Centre for Land and Biological Resources Research” di Ottawa, svolte su territori montuosi, mai prima entomologicamente esplorati e per merito del collega stafilinidologo Guillaume de Rougemont di Londra, che ha applicato tecniche di ricerca nuove, il catalogo delle specie di Aleocharinae della Cina si è ora arricchito ulteriormente di 56 specie nuove per la Cina e di 336 nuove per la scienza, con 14 nuovi generi. Con il presente lavoro e i successivi della serie, oggi il catalogo delle Aleocharinae della Cina comprende 552 specie, includendo anche le prime 7 nuove specie del genere Leptusa Kr. della Cina continentale, da me descritte in un lavoro precedente (PACE 1997), e le specie di Taiwan. (139° Contributo alla conoscenza delle Aleocharinae) Manoscritto accettato il 25.09.1997 140 ROBERTO PACE AI materiale raccolto da A. Smetana e da G. de Rougemont, ho aggiunto per lo studio tassonomico quello raccolto da Jonathan Cooter, noto studioso di Liodidae di Hereford (Gran Bretagna), da Garry Ades, da Graham Reels, dal Dr. Jeng-Tze Yang della “National Chung Hsing University” di Taiwan e dal Prof. Shugiang Li di Stuttgart (Germania). Gli holotypi delle nuove specie sono conservati nel “Muséum d’ Histoire naturelle” di Ginevra (MHNG). ELENCO DELLE SPECIE (escluse le nuove) PRONOMAEINI Pronomaea thaxteri Bernhauer, 1915 Pronomaea thaxteri Bernhauer, 1915: 148; CAMERON 1939: 31; PACE 1986: 141. 3 es., China, Zhejiang, Tianmushan, 29.1V.1993, 2.1X.1994; 13 es., China, Yunnan, Xishuang- banna, Chayanhe, 24.1.1993, Jing Hong, 11.1993, tutti de Rougemont leg. Distribuita in India, Sumatra, Malaysia, Bali, Sabah, Filippine. Nuova per la Cina. LEUCOCRASPEDINI Leucocraspedum scorpio (Blackburn, 1895) Barronica scorpio Blackburn, 1895: 203 Leucocraspedum scorpio; CAMERON 1939: 7 | d, China, Yunnan, Xishuangbanna, Chayanhe, 24.1.1993, de Rougemont leg. Specie distribuita in Sri Lanka, India, Giava, Singapore, Filippine e Australia. Nuova per la Cina. MYLLAENINI Myllaena lombokensis Pace, 1896 Myllaena lombokensis Pace, 1986: 166 8 es., China, Beijing, Yingtaogou, III.1994, 1 d, China, Zhejiang, Tianmushan, 29.IV.1993, tutti de Rougemont leg. Specie nuova per la Cina. Myllaena ledouxi Pace, 1988 Myllaena ledouxi Pace, 1988: 405 Myllaena xianbeorum Pace, 1993: 80, syn. n. 1 del 2, Hong Kong, XII.1995-1.1996, de Rougemont leg. Specie diffusa in Nepal e Cina: Sichuan, Yunnan. Nuova per Hong Kong. GYROPHAENINI Gyrophaena (s. str.) cicatricosa Motschulsky, 1858 Gyrophaena cicatricosa Motschulsky, 1858: 231 Gyrophaena (s. str.) cicatricosa: CAMERON 1939: 89 ALEOCHARINAE DELLA CINA 141 1 d el ©, Hong Kong, X11.1955-1.1996; 33 es., Hong Kong, Tai Po, III.1996, from fungi tutti de Rougemont leg. Distribuita in Sri Lanka, India e Sumatra. Nuova per la Cina. Gyrophaena (s. str.) thoracica Cameron, 1939 Gyrophaena (s. str.) thoracica Cameron, 1939: 112 5 es., China, Sichuan, Gongga Shan, above camp 2, 2800 m, 25.VIII.1994; 1 2, idem, above camp 3, 3050 m, 22.VII.1994, tutti Smetana leg. Specie dell’ India settentrionale, nuova per la Cina. Gyrophaena (Acanthophaena) appendiculata Motschulsky, 1858 Gyrophaena appendiculata Motschulsky, 1858: 228 Gyrophaena (Acanthophaena) appendiculata; CAMERON 1939: 60 1 4, Hong Kong, Kadoorie Farm, VII.1996, flight interception trap, de Rougemont leg. Specie distribuita in India, Malaisia e Filippine. Nuova per la Cina. HOMALOTINI Silusa aliena Bernhauer, 1916 Silusa aliena Bernhauer, 1916: 33 7 es., China, Sichuan, Gongga Shan, above camp 2 and 3, 2800-3050 m, 22-26.V11.1994, Smetana leg.; 6 es., China, Dali, 9.11.1993, de Rougemont leg. Specie diffusa solo in Cina. Silusa bodemeyeri (Eppelsheim, 1883) Leptusa bodemeyeri Eppelsheim, 1883: 252 Silusa bodemeyeri: Pace 1989: 24 9 es., Hong Kong, Tai Po, III-IV.1996, de Rougemont leg. Specie distribuita in Romania. Nuova per la Cina. Nota. - Non esistono signi- ficative differenze nella forma dei pezzi copulatori del sacco interno dell’edeago del tipo di bodemeyeri da me esaminato a confronto di quelli dell’edeago degli esemplari di Hong Kong. Probabile diffusione attraverso il commercio di legname. Coenonica javana Bernhauer, 1914 Coenonica javana Bernhauer, 1914: 104 Neosilusa hongkongensis Pace, 1993: 80, syn. n. l es., China, Yunnan, Xishuangbanna, Jing Hong, 11.1993; 1 es., Hong Kong, XII.1995-1.1996, flight interception trap, tutti de Rougemont leg. Specie diffusa a Giava e Hong Kong. Nuova per la Cina continentale. Coenonica ming Pace, 1993 Coenonica ming Pace, 1993: 80 4 es., China, Yunnan, Kunming, 1.11.1993, de Rougemont leg. Specie finora nota solo dello Yunnan. Coenonica impressicollis (Motschulsky, 1858) Phloeopora impressicollis Motschulsky, 1858: 257 142 ROBERTO PACE Coenonica impressicollis; CAMERON 1939: 161 1 ©, China, Yunnan, Ruili, 4.11.1993, de Rougemont leg. Specie finora nota della sola India. Nuova per la Cina. Stenomastax cribrum (Fauvel, 1878) Thectura cribrum Fauvel, 1879: 297 Stenomastax cribrum; CAMERON 1939: 171 31 es., Hong Kong, X11.1995-1.1996-IV.1996, flight interception trap, de Rougemont leg. Specie distribuita in India, Singapore, Filippine e Nuova Guinea. Nuova per la Cina. Stenomastax tuberculicollis (Kraatz, 1859) Homalota tuberculicollis Kraatz, 1859: 33 Stenomastax tuberculicollis; CAMERON 1939: 177 1 d, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. Specie diffusa in Sri Lanka e a Singapore. Nuova per la Cina. Stenomastax nigrescens (Fauvel, 1905) Homalota nigrescens Fauvel, 1905: 147 Stenomastax nigrescens, CAMERON 1939: 170 6 es., China, Zhejiang, Tianmushan, 29.1V.1993; 23. es., China, Yunnan, Xishuangbanna, Jing Hong, II. 1993, 2 4 d e 2 2 © Hong Kong, Kadoorie Farm, 19-31.V.1996, IV.1996, VIII.1996, tutti de Rougemont leg. Specie diffusa in India, a Sumatra, Singapore, Malaysia, Giava e Sabah. Nuova per la Cina. Stenomastax nepalensis (Pace, 1982) Neomalota nepalensis Pace, 1982: 89 Stenomastax nepalensis: Pace 1989a: 487 1 d, China, Yunnan, Ruili, 4.11.1993, de Rougemont leg. Specie finora nota solo per il Nepal. Nuova per la Cina. Neosilusa moultoni Cameron, 1920 Neosilusa moultoni Cameron, 1920: 233 Plagiusa moultoni: Bernhauer & Scheerpeltz 1926: 540 Neosilusa moultoni: Pace 1992: 235 1 d, China, Jiangsu, Nanjing, 17.VIII.1994, de Rougemont leg. Specie nota di Singapore e della Thailandia. Nuova per la Cina. Neosilusa ceylonica (Kraatz, 1857) Stenusa ceylonica Kraatz, 1857: 8 Plagiusa ceylonica: Camero, 1939: 167 Neosilusa ceylonica: Pace 1984: 15; Pace 1993: 71 1 2, China, Beijing, Panshan, 8.V.1993, 18 es., China, Zhejiang, Tianmushang, 2.IX.1994, 29.IV.1993, 2 8 d, China, Zhejiang, Hangzhou, 27.1V.1995; 1 es., China, Yunnan, Sanchahe: elephant reserve, 24.1.1993, de Rougemont leg.; 1 es., China, all Jiangsu Prov., Nanjing Zijinshan, 8.V.1996, J. Cooter leg.; 1 9, Hong Kong, Kadoorie Farm, flight interception trap, VII.1992, G. Ades leg.; 1 es., Hong Kong, XII.1995-1.1996; 17 es., Hong Kong, Tai Po, N. T., ALEOCHARINAE DELLA CINA 143 flight interception trap, III.1996, IV.1996, V-VI.1996, IX.1996; 1 es., Hong Kong, Kadoorie Farm, vegetable refuse, f. i. t., VI.1996, tutti de Rougemont leg. Specie distribuita dalle Mascarene, all’ India, Malesia, Cina e Giappone. DIESTOTINI Diestota testacea (Kraatz, 1859) Bolitochara testacea Kraatz, 1859: 7 Diestota testacea; FAUVEL 1905: 86; CAMERON 1939: 164; PACE 1984: 15 9 es., China, Yunnan, Xishuangbanna, Jing Hang & Mengdien, I-II.1993; 4 es., China, Yunnan, Ruili & Sanchahe (elephant reserve), 24.1.1993, 4.11.1993; 1 4 e 1 ®, Hong Kong, 25.VIIL. 1994, de Rougemont leg.; 25 es., Hong Kong, flight interception trap, XII.1995-1.1996; 23 es., Hong Kong, Tai Po, N.T., vegetable refuse, III-V.1996, IV-V.1996. IX.1996, all de Rougemont leg. Specie distribuita dalle Mascarene alla Regione Orientale. BOLITOCHARINI Pseudatheta franzi Pace, 1992 Pseudatheta franzi Pace, 1992: 240 10 es., Hong Kong, Tai Po, from fungi, III.1996; 1 4, Hong Kong, Tai Po, III.1996; 1 d, Hong Kong, N.T., IV.1996; 1 ¢, Hong Kong, Kadoorie Farm, flight interception trap, VI.1996, all de Rougemont leg. Specie nota della Thailandia, nuova per la Cina. Pseudatheta pulchra Pace, 1992 Pseudatheta pulchra Pace, 1992: 240 1 2, Hong Kong, Chinese University, Auricularia fungus, 1.IX.1996, de Rougemont leg. Specie nota della Thailandia, nuova per la Cina. Pseudatheta meorum Pace, 1992 Pseudatheta meorum Pace, 1992: 240 3 es., China Yunnan, Xishuangbanna, Cheyanhe F.P., 24. I. 1993, de Rougemont leg. Specie della Thailandia, nuova per la Cina. AUTALIINI Autalia rivularis (Gravenhorst, 1802) Aleochara rivularis Gravenhorst, 1802: 73 Autalia rivularis; MANNERHEIM 1831: 501; BERNHAUER & SCHEERPELTZ 1926: 569 1 d, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, Smetana leg. Specie diffusa in Europa settentrionale e centrale e Caucaso. Nuova per la Cina. FALAGRIINI Cordalia vestita (Boheman, 1858) Falagria vestita Boheman, 1858: 25 144 ROBERTO PACE Cordalia vestita; CAMERON 1939: 236; PACE 1993: 71 2 dd e 1 &, China, Jiangsu, Nanjing, 17.VII.1994; 10 es., China, Yunnan, Ruili, Dali; Kunming & Sanchahe (elephant reserve), 24.1.1993, 1-4-9.11.1993, all de Rougemont leg.; | es., Hong Kong, N.T., Shek Kong, V.1990, G. Reels leg.; 19 es., Hong Kong, N.T., vegetable refuse, flight interception trap. XII.1995-I.1996, IV-V-IX.1996, de Rougemont leg. Specie diffusa in Cina, India e Indonesia. Falagria (s. str.) caesa Erichson, 1839 Falagria caesa Erichson, 1839: 295 Staphylinus sulcatus Paykull, 1789: 32 (nec Staphylinus sulcatus O. F. Müller, 1776: 97) Falagria sulcata (Paykull, 1789) auct.; PACE 1993: 71 58 es., China, Beijing, B.N.U. Campus from bird’s nesting box, 7.VII.1993, at light, V- VI.1993, flight interception trap 10.VI-10-VN.1993; 14 es., China, Beijing, Yingtaogou, III.1994; 7 es., China, Xishan, [X.1992; 5 es., China, Shaanxi, Nanwutai & Xian, 16.X.1993, 17.1X.1995; 9 es., China, Xinjiang, Turfan & Baiyanggou, 2.X.1993, 10.X.1993; 14 es., China, Henan, Luoyang, 18.X.1993; 73 es., China, Hebei, Changde & Beidaihe, 29.V.1993, 3.X.1993, tutti de Rougemont leg.; 1 es., China, Gansu, Yonghai, ca. 20 Km SW Yuzhong, 2700-2800 m, 9.VIII.1994, Smetana leg.; 2 es., China, Gansu, 120 Km S Lanzhou Guanghe Xian Mai Jia, 2300 m, 8.VII.1994, Smetana leg. Specie diffusa in tutta l Europa, nel Nordafrica, in Cina e in Giappone. Falagria (Myrmecocephalus) dimidiata Motschulsky, 1858 Falagria dimidiata Motschulsky, 1858: 260 Falagria (Stenagria) dimidiata; CAMERON 1939: 250 1 ©, Hong Kong, vegetable refuse, V.1996, de Rougemont leg. Specie diffusa nello Sri Lanka e in India. Nuova per la Cina. Falagria (Myrmecocephalus) innocua Pace, 1984 Falagria (Stenagria) innocua Pace, 1984: 435 Falagria (Myrmecocephalus) innocua; PACE 1993: 72 2 es., China, Yunnan, Kunming, 1.11.1993; 2 es., China, Zhejiang, Tianmushan, 29.1V.1993; 1 2, China, Chengde, 3.X.1993, tutti de Rougemont leg. Specie finora nota solo della Cina. Falagria (Myrmecocephalus) semilucens Cameron, 1950 Falagria (Stenagria) semilucens Cameron, 1950: 106, nec Falagria (Stenagria) semi- lucens Coiffait, 1984: 154 Falagria (Myrmecocephalus) semilucens; PACE 1993: 72 9 es., China, Yunnan, Ruili & Xishuangbanna: Jing Hong, 11.1993, 4.11.1993; 84 es., Hong Kong, Kadoorie Farm, Tai Po, flight interception traps, vegetable refuse, 25.VIII.1995, XII.1995-1.1996, V.1996, IX.1996, tutti de Rougemont leg. Specie diffusa nella Penisola Malese e in Cina. Falagria (Myrmecocephalus) coiffaiti nom. n. per Falagria (Stenagria) semilucens Coiffait, 1984: 154, nec Falagria (Stenagria) semilucens Cameron, 1950: 106. Falagria (Myrmecocephalus) pallipennis Cameron, 1939 Falagria (Stenagria) pallipennis Cameron, 1939: 253 Falagria (Stenagria) innocua Pace, 1984: 435, syn. n. ALEOCHARINAE DELLA CINA 145 Falagria (Stenagria) innocua pagana Pace, 1984: 435, syn. n. 3 es., China, Beijing, Panshan, 8.V.1993; 26 es., China, Yunnan, Ruili, Xishuangbanna: Meng- dien, Jing Hong, 26.1.1993, 9.11.1993; 68 es., Hong Kong, Kadoorie Farm, flight interception trap, VIII.1992, V.1996, 19-31.V.1996, tutti de Rougemont leg. Diffusione della specie: India, Thailandia e Cina. Falagria (Myrmecocephalus) chang Pace, 1993 Falagria (Myrmecocephalus) chang Pace, 1993: 86 1 3, China, Yunnan, Ruili, 4.11.1993; 1 ©, China, Shanxi: Cuihua Shan, 11.VII.1994, tutti de Rougemont leg. Specie diffusa solo in Cina. Falagria (Myrmecocephalus) tsin Pace, 1993 Falagria (Myrmecocephalus) tsin Pace, 1993: 84 1 ©, China, Zhejiang, Tianmushan, 2.IX.1994, de Rougemont leg. Specie diffusa solo in Cina. Falagria (Myrmecocephalus) ficta Pace, 1992 Falagria (Myrmecocephalus) ficta Pace, 1992: 245 34 es., China, Yunnan, Xishuangbanna: Chayanhe F.P. 6 Mengdian, 24.1.1993, 26.1.1993, de Rougemont leg. Specie diffusa in Thailandia. Nuova per la Cina. Falagria (Leptagria) densipennis Cameron, 1939 Falagria (Anaulacaspis) densipennis Cameron, 1939: 256; Pace 1984a: 428 12 es., China, Yunnan, Ruili, Sanchahe (elephant reserve) & Xishuangbanna: Jing Hong, 24.1. 1993, 11.1993, de Rougemont leg. Specie diffusa in India e in Birmania. Falagria (Leptagria) assamensis Pace, 1985 Falagria (Anaulacaspis) assamensis Pace, 1985: 160 (nom. nov. per F. latesulcata Cameron, 1939 dell’ Assam, nec F. latesulcata Cameron, 1939 di Giava) 7 es., China, Yunnan, Xishuangbanna: Mengdian, 26.1.1993, de Rougemont leg. Specie nota dell’ Assam, è nuova per la Cina. Gnypeta (s. str.) modesta Bernhauer, 1915 Gnypeta (s. str. modesta Bernhauer, 1915a: 239; PACE 1984a: 441 2 es. China, Zhejiang, Tianmushan, 29.1V.1993; 13 es., China, Yunnan, Xishuang- banna: Jing Hong, II.1993; 17 es., Hong Kong, vegetable refuse, flight interception trap, XII.1995-1.1996, V. 1996, IX.1996, tutti de Rougemont leg. Specie diffusa a Sumatra, Birmania, Thailandia e Cina. Gnypeta (s. str.) yaoana Pace, 1992 Gnypeta (s. str.) yaoana Pace, 1992: 247 1 2, China, Yunnan, Xishuangbanna: Mengdian, 26.1.1993, de Rougemont leg. Specie della Thailandia, nuova per la Cina. 146 ROBERTO PACE ATHETINI Nehemitropia jiniana Pace, 1993: Nehemitropia jiniana Pace, 1993: 90 1 4, China, Yunnan, Xishuangbanna: Mengdian, 26.1.1993, de Rougemont leg. Specie finora nota solo della Cina. Nehemitropia lividipennis (Mannerheim, 1831) Oxypoda lividipennis Mannerheim, 1831: 484 Nehemitropia sordida (Marsham, 1802), auct. 9 es., China, Beijing, Xishan, B.N.U., V-VI.1993, 10.VI-10.VII.1993, IX.1993, flight inter- ception trap; | d, Zhejiang: Tianmushang, 2.IX.1994; 2 es., China, Henan, Luoyang, 18.X. 1993; 1 d, China, Hebei, Beidaihe, 29.V.1993; 13 es., China, Hebei, Changde, 3.X.1993; 1 ©, China, Yunnan, Dali, 9.11.1993; 2 es., China, Xinjiang, Turfan, II-X.1993, tutti de Rougemont leg.; 4 es., China, Gansu, Yonghai, ca. 20 Km SW Yuzhong, 2700-2800 m, 9.VIII.1994, Smetana leg. x Specie subcosmopolita, assente nella regione intertropicale. Atheta (Philhygra) ingenua Pace, 1993 Atheta (Notothecta) ingenua Pace, 1993: 108 14 es., China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. Specie nota solo della Cina. Atheta (Philhygra) palustris (Kiesenwetter, 1844) Homalota palustris Kiesenwetter, 1844: 318 Atheta (Philhygra) palustris; BERNHAUER & SCHEERPELTZ 1926: 629 10 es., China, Shanxi, Wutaishan, 4-5.VI.1993, de Rougemont leg. Specie diffusa nella Regione Paleartica, nuova segnalazione per la Cina. Atheta (Philhygra) pseudoelongatula Bernhauer, 1907 Atheta (Metaxya) pseudo-elongatula (sic!) Bernhauer, 1907: 411 Atheta (Philhygra) pseudoelongatula; SAWADA 1977: 182 1 d, China, Beijing, Xishan, IX.1993; 13 es., China, Beijing, B.N.U., 10.VI-10.VI1.1993, flight interception trap; 2 es., China, Beijing, Yingtaogou, III.1994, tutti de Rougemont leg. Specie diffusa in Giappone, nuova per la Cina. Atheta (Sipalatheta) ciu Pace, 1993 Atheta (Sipalatheta) ciu Pace, 1993: 92 18 es. Hong Kong, Tai Po, N.T., XII.1995-1.1996, ITI-IV.1996, de Rougemont leg. Specie nota solo della Cina. Atheta (Coprothassa) coriaria (Kraatz, 1859) Homalota coriaria Kraatz, 1859: 282 Atheta (s. str.) coriaria; CAMERON 1939: 344 Atheta (Xenota) coriaria; PACE 1984: 263; PACE 1990: 92 16 es., China, Beijing, Xishan & B.N.U., IX.1992, 10.VI-10.VII.1993, flight inter- ception trap; 1 2, China, Yunnan, Dali, 9.11.1993; 1 ©, China, Chengde, 3.X.1993; 2 22, Hong Kong, N.T., IV e IX. 1996, tutti de Rougemont leg. Specie cosmopolita. ALEOCHARINAE DELLA CINA 147 NOTA SINONIMICA. - In base alla forma della spermateca il sottogenere Xenota Mulsant & Rey, 1874 è syn. n. di Coprothassa Thomson, 1859. Atheta (Coprothassa) dilutipennis Motschulsky, 1858 Homalota dilutipennis Motschulsky, 1858: 252 Atheta (s. str.) dilutipennis; CAMERON 1939: 351 11 es., Hong Kong, Tai Po & N.T., HI-IV-IX.1996, de Rougemont leg. Specie diffusa nella regione intertropicale orientale, con infiltrazioni nella zona temperata settentrionale. Nella regione intertropicale occidentale (Africa e Americhe) vive un’altra specie confusa finora con dilutipennis avendo il maschio la medesima caratteristica spina mediana posteriore al sesto urosterno libero. Atheta (Coprothassa) melanaria (Mannerheim, 1831) Homalota melanaria Mannerheim, 1831: 484 Atheta (Coprothassa) melanaria; BERNHAUER & SCHEERPELTZ 1926: 670 Atheta (Acrotona) melanaria; BRUNDIN 1952: 102 1 ©, China, Gansu Mts., 25 Km E Xiahe, 2805-2925 m, 3.VIII.1994, Smetana leg.; 3 2 9, China, Yunnan, Xishuangbanna, Mengdian, 26.1.1993; 16 es., China, Shanxi, Wutaishan, 4- 5.V1.1993, tutti de Rougemont leg. Specie diffusa in Europa, in Transcaucasia e nell’ Altai. Nuova per la Cina. Atheta (Acrotona) annuliventris (Kraatz, 1859) Homalota annuliventris Kraatz, 1859: 40 Atheta (Acrotona) annuliventris; CAMERON 1939: 408 4 es., China, Jiangsu Prov., Nanjing Zijinshan, 8.V.1996, J. Cooter leg.; 23 es., Hong Kong, Tai Po, Chinese University (in Auricularia fungus), N.T., Kadoorie Farm, VI.1991, IX-X.1991, V. 1994, G. Ades leg., III-V-IX.1996, de Rougemont leg. Specie diffusa in India, a Singapore e Taiwan. Nuova per la Cina continentale. Atheta (Acrotona) birmana Pace, 1984 Atheta (Acrotona) birmana Pace, 1984: 445; PACE 1991: 116 53 es., China, Yunnan: Xishuangbanna, Mengdian, Ruili, Sanchahe (elephant reserve), Cha- yanhe, Jing Hong, 26.1.1993, 24.1.1993, 11.1993, 4.11.1993; 2 9 9, China, Zhejiang, Tian- mushan, 29.1V.1993, tutti de Rougemont leg.; 2 © ©, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, Smetana leg.; 138 es., Hong Kong, Kadoorie Farm, Tai Po, flight interception trap, 19-31-V.1996, VII.1996, de Rougemont leg. Specie diffusa in Nepal e in Thailandia. Nuova per la Cina. Atheta (Acrotona) fungi (Gravenhorst, 1806) Aleochara fungi Gravenhorst, 1806: 157 Atheta (Acrotona) fungi; CAMERON 1939: 402; PACE 1993: 73 16 es., China, Beijing, Dong Ling Shan, 1900 m, leaf litter Quercus-Alnus, 1.VII.1993, de Rougemont leg.; 6 es., China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994; 1 2, China, Gansu Mts., 25 Km E Xiahe, 2805-2975 m, 3.VIII.1994; 23 es., China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, tutti Smetana leg.; 1 9, Hong Kong, Kadoorie Farm, 19-31-V.1996, de Rougemont leg. Specie diffusa nella regione paleartica. Prima segnalazione per la Cina. In base alla forma della spermateca il sottogenere Mocyta Muls & Rey, 1874 è syn. n. di Acrotona Thomson, 1859. 148 ROBERTO PACE Atheta (Acrotona) vicaria (Kraatz, 1859) Homalota vicaria Kraatz, 1859: 38 Homalota inornata Kraatz, 1859: 39 Atheta (Acrotona) vicaria; CAMERON 1939: 396; PACE 1987: 434 Atheta (Acrotona) taedia Cameron, 1933: 215; SAWADA 1977: 198 Atheta (Acrotona) pseudoparens Cameron, 1933: 215; SAWADA 1977: 138 Atheta (Acrotona) vicaria immixta Pace, 1085: 177; PACE 1991: 116, syn. n. Atheta (Acrotona) cariei Pace, 1984: 263, syn. n. 1 3d, China, Beijing, Xishan, IX.1992; 5 es., China, Yunnan, Ruili ca. 700 m, Xishuangbanna: Mengdian, 26.1.1993, 3.11.1993; 4 es., China, Zhejiang, Tianmushan, 2.IX.1994; 162 es., Hong Kong, Kadoorie Farm, N.T., flight interception trap, IV-V.1996, 19-31-V.1996, VI.1996, tutti de Rougemont leg. Specie diffusa dalle Mascarene allo Sri Lanka, all’ India, Nepal, e Giappone. Atheta (Acrotona) neglecta Cameron, 1933 Atheta (Colpodota) neglecta Cameron, 1933: 215; Sawada 1977: 194 9, China, Beijing, Songshan, 15.1V.1993; 1 ©, China, Zhejiang, Tienmushan, 29.1V.1993; , China, Hebei, Chengde, 3.X.1993, tutti de Rougemont leg. Specie diffusa in Giappone. Nuova per la Cina. 2 I +O +O Atheta (Acrotona) paedida (Erichson, 1840) Homalota paedida Erichson, 1840: 917 Atheta (Acrotona) paedida; PACE 1984: 265 1 ®, Hong Kong, Tai Moshan, 600 m, cow dung; 1 4, Hong Kong, Tai Mo Shan, 800 m, forest floor litter, 20. VI.1996, tutti de Rougemont leg. Specie diffusa nel Madagascar, nelle Mascarene, alle Andamane, in India, in Cina, in Malesia e nelle Filippine. Atheta (Acrotona) probans Pace, 1984 Atheta (Acrotona) probans Pace, 1984a: 445; PACE 1993: 74 4 es., China, Yunnan, Sanchahe (elephant reserve) 24.1.1993; 2 es., China, Yunnan, Xishuang- banna: Chayanhe F.P. & Jing Hong, 24.1.1993, II. 1993, tutti de Rougemont leg. Specie diffusa in Thailandia e in Cina. Atheta (Acrotona) siamensis Pace, 1984 Atheta (Acrotona) siamensis Pace, 1984a: 443 11 es., China; Yunnan, Xishuangbanna: Mengdian, 26.1.1993; 8 es., China, Yunnan, Ruili, 4.11.1993, tutti de Rougemont leg. Specie finora nota solo della Thailandia. Nuova per la Cina. Atheta (Acrotona) subclientula Cameron, 1939 Atheta (Acrotona) subclientula Cameron, 1939: 405 27 es., China, Beijing, Xiaolongmen, 1.VII.1993, de Rougemont leg. Specie diffusa nell’ India settentrionale. Nuova per la Cina. ALEOCHARINAE DELLA CINA 149 Atheta (Acrotona) suspiciosa (Motschulsky, 1858) Homalota suspiciosa Motschulsky, 1858: 90 Atheta (Acrotona) suspiciosa, CAMERON 1939: 397; PACE 1993: 73 26 es., China, Shaanxi, Nanwutai, 17.1X.1995, de Rougemont leg. Specie diffusa nello Sri Lanka, in India e in Cina Atheta (Bessobia) occulta (Frichson, 1839) Homalota occulta Erichson, 1839: 317 Atheta (Bessobia) occulta; BERNHAUER & SCHEERPELTZ 1926: 626 Atheta (Bessobia) erichsoni Bernhauer, 1907: 397; Yost & SAWADA 1976: 80 1 3, China, Beijing, Panshan, 8.V.1993; 1 6 e 4 2 2, China, Beijing, Ying Taogou, III.1993, tutti de Rougemont leg. Specie diffusa dall’ Europa settentrionale e centrale alla Siberia e al Giappone. Nuova per la Cina. Atheta (Microdota) amicula (Stephens, 1832) Aleochara amicula Stephens, 1832: 132 Atheta (Microdota) amicula; BERNHAUER & SCHEERPELTZ 1926: 631 1 ©, China, Beijing, Songshan, 15.1V.1993, de Rougemont leg. Specie a diffusione paleartica, ma con infiltrazioni nella regione tropicale. L’amicula citata da CAMERON (1939: 326) è un insieme di cinque specie: A. amiculoides Cameron, A. kathmanduensis Pace, A. gahanensis Pace, A. arniensis Pace e A. notatella Pace. Atheta (Microdota) denticauda Bernhauer, 1907 Atheta (Metaxya) dentiventris Bernhauer, 1907: 412 Atheta (Microdota) denticauda Bernhauer, 1907: 401; SAWADA 1977: 173 2 es., China, Beijing, Yingtaogou, III.1993, de Rougemont leg. Specie finora nota solo del Giappone. Nuova per la Cina. Atheta (Microdota) mon Pace, 1992 Atheta (Microdota) mon Pace, 1992: 251 16 es., Hong Kong, Kadoorie Farm, flight interception trap, XII.1995-1.1996, 19-31.V.1996, de Rougemont leg. Specie diffusa in Thailandia, nuova per la Cina. Atheta (Microdota) subcrenulata Bernhauer, 1907 Atheta (Microdota) subcrenulata Bernhauer, 1907: 403 Atheta (Amidobia) subcrenulata; SAWADA 1974: 166 1 3, China, Beijing, flight interception trap, 10.VI-10.VII.1993; 2 8 &, China, Zhejiang, Tian- mushan, 29.IV.1993, tutti de Rougemont leg. Specie diffusa in Thailandia, nuova per la Cina. Atheta (Microdota) subcrenulata Bernhauer, 1907 Atheta (Microdota) subcrenulata Bernhauer, 1907: 403 Atheta (Amidobia) subcrenulata; SAWADA 1974: 166 150 ROBERTO PACE 1 3, China, Beijing, flight interception trap, 10.VI-10.VII. 1993; 2 G 4, China, Zhejiang, Tianmushan, 29.1V.1993, tutti de Rougemont leg. Specie giapponese, ora nota anche della Cina. Atheta (Microdota) vagans Bernhauer, 1907 Atheta (Microdota) vagans Bernhauer, 1907: 404 Atheta (Amidobia) vagans; SAWADA 1974: 152 107 es., Hong Kong, Kadoorie Farm, Tai Po, flight interception trap, XII.1995-I.1996; ITI-IV- V-VIII.1996, de Rougemont leg. Specie diffusa nello Sri Lanka, a Taiwan e in Giappone. Nuova per la Cina. Atheta (Datomicra) lewisiana Cameron, 1933 Atheta (Datomicra) lewisiana Cameron, 1933: 214 Atheta (Datostiba) lewisiana; SAWADA, 1976: 19 3 99, China Beijing, Xiaolongmen, B.N.U. & Songshan, at light, 15.1V.1993, V-VI.1993, 1.VII.1993; 8 es., China, Yunnan, Dali, 9.11.1993; 8 es., China, Zhejiang, Tianmushan & Hangzhou,, 29.1V.1993, 27.1V.1995; 18 es., China, Shanxi, Wutaishan Shaanxi, Nanwutai, 4- 5.VI.1993, 17.IX.1995; 1 2, China, Jiangsu, Nanjing, 17.VIII.1994; 2 2 ©, Hong Kong, NT. vegetable refuse, IV-V.1996, tutti de Rougemont leg. Specie diffusa in Cina, Giappone e Giava. Atheta (Datomicra) poroshirica Sawada, 1978 Atheta (Datostiba) poroshirica Sawada, 1978: 243 6 es., China, Beijing, Xiaolongmen, 1100-1500 m, 1.VHI.1993; 1 2, China, Shaanxi, Nanwutai, 17.IX.1995, tutti de Rougemont leg. Specie diffusa in Giappone. Nuova per la Cina. Atheta (Datomicra) sordiduloides Cameron, 1939 Atheta (Datomicra) sordiduloides Cameron, 1939: 386 1 8, China, Shaanxi, Nanwutai, 17.IX.1995, de Rougemont leg.; 1 3, China, Gansu, 120 Km S Lanzhou, Guanghe Xian Maijia, 2300 m, 8.VII.1994, Smetana leg. Specie diffusa dal Kashmir all India. Nuova per la Cina. Atheta (Datomicra) subsericans Cameron, 1939 Atheta (s. str.) subsericans Cameron, 1939: 355 1 d, China, Sichuan, Gongga Shan, above camp 2, 2800 m, 26.VII.1994, Smetana leg. Specie dell’ India settentrionale. Nuova per la Cina. Atheta (Dimetrota) furtiva Cameron, 1939 Atheta (Dimetrota) furtiva Cameron, 1939: 378 1 4, China, Yunnan, Ruili, ca. 700 m, 3.11.1993; 2 2 2, China, Yunnan, Dali, 9.11.1993, tutti de Rougemont leg. Specie diffusa dall’ India settentrionale alla Cina. Atheta (Dimetrota) guizhouensis Pace, 1993 Atheta (Dimetrota) guizhouensis Pace, 1993: 100 1 es., China, Zhejiang, Tianmushan, 29.1V.1993: 30 es., China: Yunnan, Xishuangbanna: ALEOCHARINAE DELLA CINA 151 Mengdian & Dali, 26.1.1993, 9.11.1993; 4 ® 9, China, Shaanxi, Nanwutai & Cuihuashan, 11.VIIL.1994, 17.1X.1995, tutti de Rougemont leg. Specie nota solo in Cina. Atheta (Dimetrota) cinnamoptera Thomson, 1856 Atheta cinnamoptera Thomson, 1856: 105 Atheta (Dimetrota) cinnamoptera; BERNHAUER & SCHEERPELTZ 1926: 662 53 es., China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, Sme- tana leg. Specie diffusa nell Europa settentrionale e centrale. Nuova per la Cina. Atheta (s. str.) atramentaria (Gyllenhal, 1810) Aleochara atramentaria Gyllenhal, 1810: 408 Atheta atramentaria, THOMSON 1861: 92 Homalota transfuga Sharp, 1874: 13 Atheta (Athera) transfuga; YOSn & SAWADA 1976: 69 Atheta (Dimetrota) sublaevana Cameron, 1939: 379; PACE 1991: 122; PACE 1993: 75 Atheta (Dimetrota) atramentaria; PACE 1984: 447 Atheta (Notothecta) kunmingensis Pace, 1993: 106, syn. n. Atheta (s. str.) atramentaria; PACE 1991: 122 1 2, China, Beijing, Yingtaogou, 900 m, III.1993; 13 es., China, Yunnan, Ruili & Dali, 4. 11.1993, 9.11.1993, tutti de Rougemont leg. Specie subcosmopolita: regione paleartica e regione etiopica. Atheta (s. str.) euryptera (Stephens, 1832) Aleochara euryptera Stephens, 1832: 132 Atheta (s. str.) euryptera; BERNHAUER & SCHEERPELTZ 1926: 642 13 es., China, Zhejiang, Hangzhou, 27.1V.1995, de Rougemont leg. Specie diffusa nella Regione Olartica. Prima segnalazione per la Cina. Atheta (Notothecta) pseudocoriaria Cameron, 1939 Atheta (s. str.) pseudocoriaria Cameron, 1938: 345 Atheta (Xenota) pseudocoriaria; PACE 1986: 154 39 es., Hong Kong, Kadoorie Farm, Tai Po, N.T., malaise trap, vegetable refuse, V.1992, VI.1993 G. Ades leg., IU-IV-V-VII.1996, de Rougemont leg. Specie diffusa in India, Nepal e Giava. Nuova per la Cina. Atheta (Notothecta) reitteriana Bernhauer, 1939 Atheta (Acrotona) reitteriana Bernhauer, 1939: 109 Atheta (Notothecta) reitteriana; Yosu & SAWADA 1976: 44; PACE 1993: 75 Atheta (s. str.) cameroni Pace, 1987: 403; Pace 1993: 75 5 es., China, Beijing, Xishan & Songshan, IX.1992, 15.1V.1993; 35 es. China, Yunnan, Dali, Ruili & Xishuangbanna: Mengdien, 26.1.1993, 4.11.1993, 9.11.1993; 4 es., China, Yunnan, Kunming, 1.11.1993; 9 es., China, Zhejiang, Tianmushan, 29.1V.1993; 8 es., China, Shaanxi, Nanwutai, 17.1X.1995; 16 es., Hong Kong, Tai Po & N.T., flight interception traps, XII.1995- 1.1996, III-IV.1996, tutti de Rougemont leg. Specie presente in Cina, Giappone, Taiwan, Nepal, India settentrionale e Bir- mania. 152 ROBERTO PACE Geostiba (Indatheta) rougemonti Pace, 1993 Geostiba (Indatheta) rougemonti Pace, 1993: 90 9 es., China, Gansu, Xilong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, Smetana leg.; 1 6, China, Yunnan, Kunming, 1.11.1993; 1 2, China, Zhejiang, Tianmushan, 29.1V.1993; 1 2, China, Zhejiang, Lin An County, 1000 m, W. Tianmu Shan N. R., 18.V.1996, tutti de Rougemont leg. Specie endemica della Cina. Pelioptera opaca Kraatz, 1857 Pelioptera opaca Kraatz, 1857; CAMERON 1939: 418 1 2, China, Beijing, Yingtaogou, HI.1993; 3 es., China, Yunnan, Xishuangbanna, Jing Hong, 11.1993; 1 8, Kadoorie Farm, flight interception trap, 19-31.V.1996, tutti de Rougemont leg. Specie diffusa dallo Sri Lanka all India e a Singapore. Nuova per la Cina. Pelioptera testaceipennis (Motschulsky, 1858) Homalota testaceipennis Motschulsky, 1858: 251 Homalota pelioptera Kraatz, 1859: 30; SAWADA 1980: 51 Atheta (Dimetrota) testaceipennis; CAMERON 1939: 377 Pelioptera pelioptera; CAMERON 1939: 414 Pelioptera testaceipennis; SAWADA 1980: 51; PACE 1984a: 428 1 ®, China, Zhejiang, Tianmushan, 2.IX.1994; 91 es., Hong Kong, Kadoorie Farm, Tai Po, flight interception trap, XII.1995-1.1996, III-IV-V.1996, 19-31.V.1996, tutti de Rougemont leg. Specie presente nello Sri Lanka, in Nepal, India, Birmania, Sabah e Giappone. Nuova per la Cina. Pelioptera micans Kraatz, 1858 Pelioptera micans Kraatz, 1857: 56; CAMERON 1939: 415 I 3 e 1 2, Hong Kong, Kadoorie Farm, flight interception trap, XII.1995-1.1996, 19-31.V. 1996, de Rougemont leg. Specie presente in Sri Lanka, India e Singapore. Nuova per la Cina. Gastropaga (Rougemontia) siamensis (Pace, 1984) Rougemontia siamensis Pace, 1984: 450 Gastropaga (Rougemontia) siamensis; PACE 1993: 76 1 3, China, Beijing, Xishan, IX.1992, de Rougemont leg. Specie presente in Thailandia e Cina. THAMIARAEINI Thamiaraea insigniventris Fauvel, 1878 Thamiaraea insigniventris Fauvel, 1878: 299 1 d, Hong Kong, N.T., IV.1996, de Rougemont leg. Specie presente in Sri Lanka, Singapore, Sumatra, Celebes, Sabah, Nuova Guinea e Filippine. Schistogenia crenicollis Kraatz, 1857 Schistogenia crenicollis Kraatz, 1857: 40; CAMERON 1939: 424; PACE 1993: 76 ALEOCHARINAE DELLA CINA 153 6 es., China, Zhejiang, Tianmushan, 2.1X.1994: 2 es., China, Yunnan, Xishuangbanna, Jing Hong, 11.1993; 6 es., Hong Kong, N.T., IX.1996, tutti de Rougemont leg. Specie diffusa in Sri Lanka, India, Malesia, Indonesia e Cina. Mimacrotona orousseti Pace, 1990 Mimacrotona orousseti Pace, 1990a: 167 18 es., Hong Kong, Kadoorie Farm, Tai Po, N.T., VII-VIH-IX.1996, de Rougemont leg. Specie del Nepal, nuova per la Cina. MYRMEDONIINI Amaurodera yaoana Pace, 1992 Amaurodera yaoana Pace, 1992: 257 6 es., China, Yunnan, Xishuangbanna, Chayanhe F.P., 24.1.1993, de Rougemont leg. Specie della Thailandia, nuova per la Cina. Zyras (s. str.) songanus Pace, 1993 Zyras (s. str.) songanus Pace, 1993: 114 I d, China, Shanxi, Wutaishan, 4-5.V1.1993, de Rougemont leg. Specie presente solo in Cina. Lomechusa minor Reitter, 1887 Lomechusa minor Reitter, 1887: 210; SCHILOW 1981: 219 1 ©, China, Gansu, Pass btw Hezuo & Amgog, 3300 m, 12.VII.1994, Smetana leg., 1 es., Xinjiang, Nanshan, ca. 40 Km S.W. Uromgi, VIIL1981, de Rougemont leg. Specie diffusa solo in Cina (SCHILOW 1981). OXYPODINI Chilopora longitarsis (Erichson, 1839) Calodera longitarsis Erichson, 1839: 698 Chilopora longitarsis; KRAATZ 1858: 147 1 2, China, Shanxi, Wutaishan, 4-5.VI.1993, de Rougemont leg. Specie diffusa nella Regione Paleartica occidentale. Nuova per la Cina. Phloeopora teres (Gravenhorst, 1802) Aleochara teres Gravenhorst, 1802: 79 Phloeopora teres; BERNHAUER & SCHEERPELTZ 1926: 720 4 es., China, Hebei, Chengde, 3.X.1993, de Rougemont leg. Specie paleartica occidentale, nuova per la Cina. Amarochara (s. str.) umbrosa (Erichson, 1839) Calodera umbrosa Erichson, 1839: 304 Amarochara umbrosa; THOMSON 1860: 300 ld e2 2 2, China, Hebei Prov., Yongnian, 6.X.1995, Shugiang Li leg. Specie diffusa nella Regione Paleartica occidentale e nella Regione Neartica. Nuova per la Cina. 154 ROBERTO PACE Apimela lineola (Kraatz, 1859) Oxypoda lineola Kraatz, 1859: 27 Pseudomeotica lineola; CAMERON 1939: 582; PACE 1986: 164; PACE 1992: 123. 46 es., Hong Kong, Kadoorie Farm, Tai Po, N.T., HI-IV-V-VII-IX.1996, de Rougemont leg. Specie diffusa dallo Sri Lanka, all’ India, Singapore e Ball. Apimela consors Pace, 1992 Apimela consors Pace, 1992a: 281 24 es., Hong Kong, Kadoorie Farm, Tai Po, 19-31.V.1996, de Rougemont leg. Specie nota del Nepal, nuova per la Cina. Ocalea himalayica Cameron, 1939 Ocalea himalayica Cameron, 1939: 578 2 99, China, Zhejiang Prov., Anji County, ca. 500 m, Long Wang Shan N.R., 12.V. 1996, J. Cooter leg. Specie finora nota solo dell India settentrionale. Nuova per la Cina. Oxypoda (Podoxya) shuteae Pace, 1993 Oxypoda (Podoxya) shuteae Pace, 1993a: 172 1 ©, China, Yunnan, Xishuangbanna, Chayanche, F.P., 24.1.1993, de Rougemont leg. Specie diffusa in India settentrionale, ora presente anche in Cina. Oxypoda (Podoxya) subsericea Cameron, 1939 Oxypoda (Podoxya) subsericea Cameron, 1939: 603 5 es., China, Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. Specie dell’ India settentrionale, nuova per la Cina. Oxypoda (Paroxypoda) morosa Cameron, 1939 Oxypoda (Paroxypoda) morosa Cameron, 1939: 597 1d e2 99, China, Beijing, Dong Ling Shan, 1900 m, leaf litter Quercus-Alnus, 1.VII.1993, de Rougemont leg. La specie era nota solo dell’India settentrionale per il solo holotypus. Nuova per la Cina. Oxypoda (Sphenoma) connexa Cameron, 1939 Oxypoda (Sphenoma) connexa Cameron, 1939: 615; PACE 1992: 268 1 2, China, Sichuan, Gongga Shan, above camp 2, 2750 m, 24.VII.1994, Smetana leg.; 1 2, China, Yunnan, Dali, 9.11.1993, de Rougemont leg. Specie particolarmente diffusa in India settentrionale e in Nepal. Nuova per la Cina. HOPLANDRIINI Pseudoplandria beesoni Cameron, 1939 Pseudoplandria beesoni, Cameron, 1939: 672 1 2, China, Yunnan, Ruili, 4.11.1993, de Rougemont leg. Specie diffusa nell India settentrionale, nuova per la Cina. ALEOCHARINAE DELLA CINA [55 Pseudoplandria osellaiana Pace, 1984 Pseudoplandria osellaiana Pace, 1984b: 488 2 2 2, China, Zhejiang, Lin’an County, W. Tianmu Shan N.R., 16-22.V.1996, J. Cooter Leg.; 5 es., Hong Kong, Kadoorie Farm, VIII.1996, de Rougemont leg. Specie della Thailandia, nuova per la Cina. Pseudoplandria bohaci Pace, 1992 Pseudoplandria bohaci Pace, 1992b: 127 1 2, Hong Kong, IV.1996, de Rougemont leg. Specie del Vietnam, nuova per la Cina. ALEOCHARINI Aleochara (Coprochara) bipustulata (Linnaeus, 1761) Staphylinus bipustulatus Linnaeus, 1761: 232 Aleochara (Coprochara) bipustulata; CAMERON 1939: 650 1d e1 ®, China, Beijing, Panshan & Xiaolongmen, 8.V.1993, 1.VII.1993, de Rougemont leg. Specie a diffusione olartica e sudafricana. Aleochara (Calochara) formosanorum Pace, 1993 Aleochara (Calochara) formosanorum Pace, 1993a: 178 1 4, Hong Kong, XII.1995-1.1996, flight interception trap, de Rougemont leg. Specie diffusa a Taiwan, nuova per la Cina continentale. Aleochara (Xenochara) puberula Klug, 1833 Aleochara puberula Klug, 1833: 139 Aleochara (Xenochara) puberula; KLIMASZEWSKI & JANSEN 1993: 72 1 2, China, Yunnan, Xishuangbanna, Mengdien, 26.1.1993; 9 es., Hong Kong, Tai Mo Shan, 600 m, 20.VI.1996, cow dung, tutti de Rougemont leg. Specie cosmopolita infatti le sue larve sono predatrici di larve di Musca domestica, Stomoxys calcitrans, ecc. Aleochara (s. str.) nigra Kraatz, 1859 Aleochara nigra Kraatz, 1859: 13 Aleochara (s. str.) nigra; CAMERON 1939: 626 1 d, Hong Kong, VIII.1991, G. Ades leg. Specie diffusa dallo Sri Lanka all’ India, alla Birmania, alla Penisola Malese e a Sumatra. Nuova per la Cina. DESCRIZIONI MASURIINI Masuria (Oncosomechusa) chinensis sp. n. Figg. 1-4 Tipi. Holotypus d, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VII.1994, A. Smetana leg. (MHNG). Paratypi: 38 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,70 mm. Corpo lucido e bruno con elitre nero-brune e addome nero con estremità distale bruna; antenne brune con i tre antennomeri basali 156 ROBERTO PACE rossicci; zampe rossicce. La punteggiatura del capo e del pronoto è composta di punti grandi e superficiali. Le elitre e l'addome sono coperti da tubercoletti svaniti. L’avancorpo è privo di microscultura reticolare, l’addome è coperto di distinta reticolazione. Edeago figg. 2-3, spermateca fig. 4. COMPARAZIONI. Due sono le specie note del sottogenere Oncosomechusa Pace, 1982a: besucheti Pace, 1982a e tashigaonensis Pace, 1989b, entrambe del Nepal, note solo su esemplari femmine. La nuova specie è ben distinta da entrambe per avere il pronoto meno trasverso, elitre meno ridotte e bulbo prossimale della spermateca più sviluppato del distale e non molto meno sviluppato come nelle due specie del Nepal. HYPOCYPHTINI Cypha yunnanensis sp. n. Figg. 5-7 Tipi. Holotypus ¢, China, Yunnan, Ruili, ca. 700 m, de Rougemont leg. (MHNG). Paratypi: 2 es., stessa provenienza. DESCRIZIONE. Lunghezza 1,4 mm. Corpo lucidissimo, molto convesso e bruno scuro, con estremità addominale rossiccia; antenne e zampe giallo-rossicce. Su tutto il corpo non vi è traccia di reticolazione. La punteggiatura del capo è estremamente fine e rada; quella del pronoto e delle elitre è estremamente superficiale e molto fine. Sui tre uroterghi basali vi è una fila di punti un po’ allungati. Edeago figg. 6-7. COMPARAZIONI. In base alla forma dell’edeago, la nuova specie mostra affinità con C. besuchetiella (Pace, 1985a), comb. n. (“olim” Hypocyphtus besuchetiellus Pace, 1985a: 81), dell’ India. La nuova specie differisce da besuchetiella per avere gli occhi meno ridotti, l'addome più ristretto all'indietro e per l’edeago meno sviluppato, con un ciuffo di setole del sacco interno, assente in besuchetiella, qui sostituito da due lame. Nora. Il riconoscimento che il genere Hypocyphtus Gyllenhal, 1827 è sinonimo del genere Cypha Leach, 1819, comporta la necessità di stabilire le seguenti nuove combinazioni: Cypha helvetiorum (Pace, 1985a), comb. n., “olim” Hypocyphtus helvetiorum Pace, 1985a: 81. Cypha loebliella (Pace, 1985a), comb. n., “olim” Hypocyphtus loebliellus Pace, 1985a: 81. Cypha nepalensis (Pace, 1985a), comb. n., “olim” Hypocyphtus nepalensis Pace, 1985a: 81. Cypha puer (Pace, 1985a), comb. n., “olim” Hypocyphtus puer Pace, 1985a: 84. Cypha pusilla (Pace, 1984a), comb. n., “olim” Hypocyphtus pusillus Pace, 1985a: 84. Cypha hystrix (Pace, 1985a), comb. n., “olim” Hypocyphtus hystrix Pace, 1985a: 84. Cypha senilis (Pace, 1985a), comb. n., “olim” Hypocyphtus senilis Pace, 1985a: 84. MYLLAENINI Myllaena adesi sp. n. Figg. 8-11 Tipi. Holotypus 2, Hong Kong, N.T., Kadoorie Agricultural Research Centre, malaise trap, V.1992, G. Ades leg. (MHNG). 1577 ALEOCHARINAE DELLA CINA 01 mm Olmm FIGG. 1-7 Habitus, edeago in visione laterale e ventrale e spermateca. 1-4: Masuria (Oncosomechusa) chinensis sp. n.; 5-7: Cypha yunnanensis sp. n. 158 ROBERTO PACE Paratypi: 13 es., Hong Kong, N.T., IV.1996; 2 d 4, idem IX.1996; 20 es., Hong Kong, Tai Mo Shan, 600-800 m, forest floor litter, cow dung, 20.VI.1996, tutti de Rougemont leg. DESCRIZIONE. Lunghezza 2,7 mm. Corpo lucido e bruno-rossiccio con elitre brune tranne la base e con addome rossiccio; antenne giallo-rossicce con gli antennomeri basali 2, 3 e 4 rossicci; zampe gialle. L'intero corpo è coperto di fitta pubescenza aderente d’aspetto sericeo. Edeago figg. 9-10, spermateca fig. 11. COMPARAZIONI. In base alla forma dell’edeago e soprattutto della spermateca, la nuova specie si presenta affine a M. himalayca Cameron, 1939, dell’India setten- trionale. Ne è nettamente differente perché l'apice dell’edeago di himalayca è acu- tissimo e non a punta ogivale come nella nuova specie e la spermateca ha bulbo distale molto più sviluppato e volto verso il lato destro, mentre in himalayca il bulbo distale è chiaramente poco sviluppato e non volto verso il lato destro. ETIMOLOGIA. Specie dedicata al suo primo raccoglitore Garry Ades, zoologo collega di Guillaume de Rougemont. Myllaena kunmingensis sp. n. Figg. 12-15 Tipi. Holotypus 4, China, Yunnan, Kunming, 1.11.1993, de Rougemont leg. (MHNG). Paratypi: 1 9 e 1 es. senza addome, stessa provenienza. DESCRIZIONE. Lunghezza 2,7 mm. Corpo lucido e bruno con estremità addominale distale rossiccia; antenne brune con antennomero basale giallo sporco; zampe gialle. L'intero corpo è coperto di pubescenza sericea aderente. Edeago figg. 12-13, sper- mateca fig. 14. COMPARAZIONI. Specie appartenente al gruppo di M. lateritia Kraatz, 1859, dello Sri Lanka, per la forma della spermateca, ma più affine a M. lombokensis Pace, 1986, di Lombok, per la presenza di appendici laterali dell’introflessione del bulbo distale della spermateca, appendici assenti ai lati dell’introflessione del bulbo distale della spermateca di lateritia. La parte prossimale della spermateca della nuova specie descrive due strette spire, mentre quella della spermateca di /ombokensis descrive tre spire. Inoltre l’apice dell’edeago della nuova specie è uncinato, in visione laterale, mentre non lo è in /ombokensis. Myllaena chinoculata sp. n. Figg. 16-19 Tipi. Holotypus 4, China, Zhejiang, Tianmushan, 2.IX.1993, de Rougemont leg. (MHNG). Paratypi: 2 © 9, stessa provenienza. DESCRIZIONE. Lunghezza 1,9 mm. Corpo debolmente lucido e giallo-rossiccio con addome rossiccio con apice giallo-rossiccio; antenne e zampe gialle. Tutto il corpo è coperto di fitta pubescenza sericea aderente. Il quinto urotergo libero non presenta margine posteriore bianco. Edeago figg. 17-18, spermateca fig. 19. COMPARAZIONI. La nuova specie, per la forma della spermateca, appartiene al gruppo di M. lateritia Kraatz, 1859, dello Sri Lanka, ma questa specie ha occhi molto più sviluppati, pronoto molto ampio ed elitre lunghe quanto il pronoto e non nettamente ALEOCHARINAE DELLA CINA 159 01 mm 005 mm 11 Olmm Fico. 8-15 Habitus, edeago in visione laterale e ventrale e spermateca. 8-11: Myllaena adesi sp. n.; 12-15: Myllaena kunmingensis sp. n. 160 ROBERTO PACE più corte del pronoto, come nella nuova specie. Inoltre la spermateca della nuova specie ha bulbo distale più lungo che largo, mentre in lateritia detto bulbo è più largo che lungo. Myllaena tianmushanensis sp. n. Figg. 20-23 Tipi. Holotypus ¢, China, Zhejiang, Tianmushan, 2.IX.1994, de Rougemont leg. (MHNG). Paratypi: 16 es., stessa provenienza; 5 es., China, Zhejiang Prov., Lin’an County, 1000 m, W. Tianmu Shan N.R., 18.V.1996, J. Cooter leg. DESCRIZIONE. Lunghezza 2,8 mm. Corpo debolmente lucido, con capo e addome bruni, pronoto ed elitre giallo-bruni e antenne bruno-rossicce con i due antennomeri basali e l’undicesimo gialli; zampe rossicce. L'intero corpo è coperto di pubescenza sericea. Edeago figg. 20-21, spermateca fig. 22. COMPARAZIONI. Sia per la forma dell’edeago, che per quella della spermateca, la nuova specie mostra sicure affinità con M. yunnanensis Pace, 1993, pure della Cina. La nuova specie presenta una debole bozza ventrale dell’edeago, mentre in yun- nanensis tale bozza è molto saliente; l'apice dell’edeago, in visione ventrale, nella nuova specie è arcuato, mentre in yunnanensis è a forma di corta punta di lancia. Myllaena speciosa sp. n. Figg. 24-27 Tipi. Holotypus d, China, Yunnan, Dali, 9.11.1993, de Rougemont leg. (MHNG). Paratypi: 36 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido, molto convesso e bruno, con estre- mità addominale rossiccia; antenne brune, con i tre antennomeri basali e l'undicesimo rossicci; zampe rossicce. L'intero corpo è coperto di pubescenza sericea. Edeago figg. 25-26, spermateca fig. 27. COMPARAZIONI. Una specie che presenta come nella nuova specie la spermateca con un numero di spire superiore a dieci è M. ming Pace, 1993, pure della Cina. Tuttavia essa ha bulbo distale della stessa spermateca, all’apice, per nulla ristretto e volto al lato destro, come nella nuova specie, ma con bulbo distale subsferico. Inoltre l’undi- cesimo antennomero è bruno in ming e rossiccio nella nuova specie. Myllaena salamannai sp. n. Figg. 28-29 Tipi. Holotypus ©, China, Zhejiang, Tianmushan, 2.IX.1994, de Rougemont leg. (MHNG). Paratypi: 3 9 ©, stessa provenienza. DESCRIZIONE. Lunghezza 3,5 mm. Corpo debolmente lucido e nero-bruno, con pro- noto ed estremità addominale bruno-rossicci e con elitre bruno-rossicce scure; antenne brune con i due antennomeri basali rossicci e con apice dell’undicesimo di un giallo sporco; zampe giallo-rossicce. Tutto il corpo è coperto di pubescenza sericea. Spermateca fig. 29. COMPARAZIONI. Per la taglia corporea e per la forma della spermateca, la nuova specie è affine a M. ming Pace, 1993, pure della Cina. Tuttavia il colore del pronoto è bruno- ALEOCHARINAE DELLA CINA 161 Figc. 16-23 Habitus, edeago in visione laterale e ventrale e spermateca. 16-19: Myllaena chinoculata sp. n.; 20-23: Myllaena tianmushanensis sp. n. 162 ROBERTO PACE FIGG. 24-29 Habitus, edeago in visione laterale e ventrale e spermateca. 24-27: Myllaena speciosa sp. n.; 28-29: Myllaena salamannai sp. n. ALEOCHARINAE DELLA CINA 163 rossiccio nella nuova specie e bruno pece in ming. Ma è soprattutto la forma del bulbo distale e dell’introflessione apicale di esso della spermateca che permette di distinguere nettamente le due specie; introflessione lunghissima nella nuova specie, corta in ming; bulbo distale tronco-conico nella nuova specie e oviforme in ming. ETIMOLOGIA. Specie dedicata al Prof. Giovanni Salamanna dell’ Università di Genova, come piccolo segno di riconoscenza per il prezioso lavoro per i soci, come direttore delle pubblicazioni della Società Entomologica Italiana. Myllaena tianmumontis sp. n. Figg. 30-32 Tipo. Holotypus d, China, Zhejiang Prov., Lin’an County, 1000 m, W. Tianmu Shan N.R., 18.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido e giallo-bruno, con estremita addo- minale giallo-rossiccia; antenne giallo-rossicce; zampe gialle. Una uniforme pubes- cenza sericea copre l’intera superficie del corpo. Edeago figg. 31-32. COMPARAZIONI. In base alla forma dell’edeago, la nuova specie si presenta affine a M. lateritia Kraatz, 1859, dello Sri Lanka, ma la nuova specie, oltre ad avere tale organo strettamente incavato al lato ventrale (largamente incavato in /ateritia), ha l’apice dell’edeago stesso, in visione ventrale, a punta di lancia (a punta smussata in late- ritia). Fico. 30-32 Habitus ed edeago in visiona laterale e ventrale. 30-32: Myllaena tianmumontis sp. n. 164 ROBERTO PACE TERMITOHOSPITINI Sinophilus rougemonti sp. n. Figg. 33-36 Tipi. Holotypus ©, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, 19-31.V.1996, de Rougemont leg. (MHNG). Paratypi: 4 es., Hong Kong, Tai Po, flight interception, V-VII-IX.1996, de Rougemont leg. DEscRIZIONE. Lunghezza 2,6 mm. Corpo lucidissimo con capo bruno-rossiccio, pro- noto bruno-rossiccio con margini giallo-rossicci, elitre brune con base rossiccia e addome giallo-rossiccio con margine posteriore degli uriti e quinto urite libero, bruni; antenne giallo-rossicce con antennomero basale giallo e 1 successivi 2°, 3° e 4° bruno- rossicci; zampe giallo-rossicce. L’avancorpo è privo di reticolazione; la reticolazione dell’addome è poco distinta. La punteggiatura del capo è estremamente svanita. Il pronoto presenta solo alcuni punti isolati. I tubercoletti che coprono le elitre e l’addome sono svaniti. Il sesto urotergo libero sia del maschio che della femmina è profondamente inciso al margine posteriore. Edeago figg. 34-35, spermateca fig. 35. COMPARAZIONI. La nuova specie appartiene a un genere descritto di recente (KISTNER 1985) per una nuova specie della Cina: S. xiai Kistner, 1985, della provincia dello Zhejiang. La nuova specie se ne distingue perché ha il pronoto trasversalmente im- presso, le elitre più sviluppate in lunghezza, l’introflessione apicale del bulbo distale della spermateca stessa, nettamente più profonda e la parte prossimale della sper- mateca descrive 4 sinuosità invece di due come in xiai. ETIMOLOGIA. Specie dedicata al collega stafilinidologo Guillaume de Rougemont di Londra, che l’ha raccolta. GYROPHAENINI Brachida kadooriorum sp. n. Figg. 37-38 Tipo. Holotypus ©, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, VIII.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,0 mm. Corpo lucido, molto convesso e rossiccio con elitre oscurate di bruno alla parte posteriore; antenne rossicce con 1 tre antennomeri basali gialli; zampe gialle. La punteggiatura del capo è fine e svanita, quella del pronoto è così fine da essere quasi indistinta. I tubercoletti che coprono le elitre sono super- ficiali, quelli dell’addome sono allungati e distinti. Due fossette svanite stanno sul capo. Il pronoto non presenta punti grandi isolati. Non vi è presenza di reticolazione su tutto il corpo. Spermateca fig. 38. COMPARAZIONI. Specie simile a B. crassiuscula (Kraatz, 1859), dello Sri Lanka, per il colore del corpo; ma gli occhi sono più sporgenti, il quarto antennomero è nettamente trasverso (Più lungo che largo in crassiuscula) e per la spermateca a pareti sottili (spesse in crassiuscula) con introflessione apicale del bulbo distale presente (assente in crassiuscula). ETIMOLOGIA. Specie dedicata ai fratelli Kadoorie, grandi filantropi di Hong Kong che hanno fondato il “Kadoorie Agricultural Research Centre” dell’ Università di Hong ALEOCHARINAE DELLA CINA 165 Kong, luogo dove molte ricerche entomologiche sono state compiute da G. de Rougemont e dove è stata raccolta la nuova specie di Brachida. Brachida hongkongensis sp. n. Figg. 39-40 Tipo. Holotypus 2, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, VIII.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,1 mm. Corpo lucido, molto convesso e giallo-rossiccio con la meta posteriore delle elitre bruno-rossiccia; antenne rossicce con 1 tre antenno- meri basali gialli; zampe giallo-rossicce. La punteggiatura del capo è finissima, quasi indistinta, quella del pronoto è assente. Tubercoletti fini e superficiali coprono le elitre, quelli dell’addome sono allungati e netti. Non è presente reticolazione sul corpo. Due punti isolati stanno sul capo e sul pronoto. Spermateca fig. 39. COMPARAZIONI. Specie distinta dalla precedente B. kadooriorum sp. n., per avere la pubescenza al lati del corpo meno lunga, per la presenza di due punti isolati sul pronoto (assenti in kadooriorum) e soprattutto per la forma del bulbo distale della spermateca: ovale trasverso e senza introflessione apicale nella nuova specie, sub- sferico e con introflessione apicale in kadooriorum. L'assenza di lunga appendice sulla spermateca distingue ulteriormente la nuova specie da kadooriorum e da crassi- uscula (Kraatz, 1859), dello Sri Lanka. Brachida solifuga sp. n. Fig. 41 Tipo. Holotypus ®, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, VIII.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido, molto convesso e interamente giallo- rossiccio, comprese antenne e zampe. La punteggiatura del capo è indistinta. I tuber- coletti che coprono il pronoto sono assai svaniti, quelli delle elitre sono salienti. I tre uroterghi basali sono coperti di fitti tubercoletti salienti, tranne che alla base e nel solco basale, dove sono assenti tubercoletti e punteggiatura; i restanti uroterghi sono coperti di tubercoletti svaniti. Purtroppo all’atto dell'esame microscopico, la sperma- teca non era presente nella cavità addominale. COMPARAZIONI. Per il colore del corpo interamente giallo-rossiccio, la nuova specie è da avvicinare tassonomicamente a B. flava Cameron, 1939, del Bengala. Tuttavia questa specie ha taglia inferiore (2,0 mm) e non possiede i caratteri unici osservabili nella nuova specie, quali due file di tre punti sulla fascia longitudinale mediana del pronoto e tubercoletti salienti sui tre uroterghi basali, totalmente assenti sulla fascia basale di ciascun urotergo. Neobrachida jiangsuensis sp. n. Figg. 42-45 Tipo. Holotypus d, China, Jiangsu Prov., Nanjing Zijinshan, 8.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 1,4 mm. Corpo lucido e bruno con elitre e addome bruno- rossicci; antenne rossicce con antennomeri basali 1°, 2° e 3° gialli e l’undicesimo 166 ROBERTO PACE \ \ TRUMAN RN ENNA 944 URS v . v SE PLIS i vv use Fico. 33-40 Habitus, edeago in visione laterale e ventrale e spermateca. 33-36: Sinophilus rougemonti sp. n.: 37-38: Brachida kadooriorum sp. n.; 39-40: Brachida hongkongensis sp. n. ALEOCHARINAE DELLA CINA 167 bruno; zampe gialle. Il capo è privo di punteggiatura; il pronoto ha radi punti svaniti. Le elitre sono coperte di tubercoletti svaniti e di reticolazione trasversa distinta. L’addome, coperto di reticolazione molto superficiale, mostra tubercoletti allungati tra cui alcuni sono più salienti. Edeago figg. 43-44, sesto urotergo libero del maschio fig. 45. COMPARAZIONI. Il genere Neobrachida Cameron, 1920a, comprende la sola specie castanea Cameron, 1920a, dello Sri Lanka. L'esame dell’holotypus mi ha permesso di osservare che esso è una femmina. La nuova specie differisce da castanea per il pronoto e le elitre molto sparsamente punteggiate, per il pronoto molto meno largo delle elitre (elitre appena più larghe del pronoto in castanea) e per l'assenza di scul- tura squamiforme sui tre uroterghi basali, come si osserva in castanea. Neobrachida punctum sp. n. Figg. 46-48 Tipo. Holotypus 2, China, Zhejiang Prov., Lin’an County, 350 m, W. Tianmu Shan N.R., 16-22.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 1,4 mm. Corpo lucido e bruno-rossiccio con capo e uriti liberi terzo e quarti bruni; antenne giallo-brune con i tre antennomeri basali gialli e l’undicesimo bruno; zampe gialle. La reticolazione della superficie del capo è molto superficiale, quella sul resto del corpo è svanita. La punteggiatura del capo è poco distinta. I tubercoletti del pronoto e delle elitre sono spuerficiali, quelli dell’addome sono distinti. Spermateca fig. 47. COMPARAZIONI. La nuova specie per avere il pronoto e le elitre fittamente pubescenti, sembra più affine a N. castanea Cameron, 1920a, dello Sri Lanka, che a N. jiang- suensis sp. n. sopra descritta, che ha pubescenza del pronoto a delle elitre molto rada. La nuova specie ha taglia corporea molto minore di quella di castanea: 1,4 mm invece di 2,0 mm. La spermateca della nuova specie ha bulbo distale poco trasverso e senza introflessione apicale, mentre castanea ha il bulbo distale della spermateca ovale molto trasverso e con un’introflessione apicale larghissima e profonda fino quasi a raggiungere l’asse longitudinale del bulbo stesso. Gyrophaena (Phaenogyra) lividoides sp. n. Figg. 49-52 Tripi. Holotypus d, Hong Kong, Tai Po, VII.1996, de Rougemont leg. (MHNG). Paratypi: 2 4 4, Kadoorie Agricultural Research Centre, VHI.1996; 3 es., Hong Kong, Chinese University, in Auricularia fungus, 1.IX.1996; 1 3, Hong Kong, N.T., IX.1996, tutti de Rougemont leg. DESCRIZIONE. Lunghezza 1,3 mm. Corpo lucido con capo e pronoto bruno-rossicci, elitre brune con base e lati esterni bruno-rossicci, addome giallo-bruno; antenne brune con i tre antennomeri basali gialli; zampe gialle. La punteggiatura del capo è distinta, quella del pronoto assai svanita e quella delle elitre superficiale. Il capo e il pronoto non presentano microsultura reticolare; le elitre mostrano una reticolazione svanita e l'addome una reticolazione a maglie poligonali irregolari distinte. Il quinto urotergo libero del maschio ha un tubercolo mediano triangolare piatto. Edeago figg. 50-51, sesto urotergo libero del maschio fig. 52. 168 ROBERTO PACE RAR HS ma EN KAHN Figc. 41-52 Habitus, edeago in visione laterale e ventrale, sesto urotergo libero del maschio o della femmina e spermateca. 41: Brachida solifuga sp. n., 42-45: Neobrachida jiangsuensis sp. n.; 46-48: Neobrachida punctum sp. n.; 49-52: Gyrophaena (Phaenogyra) lividoides sp. n. ALEOCHARINAE DELLA CINA 169 COMPARAZIONI. La nuova specie ha caratteri dell’edeago e del sesto urotergo libero del maschio tali da permettere di avvicinarla tassonomicamente a G. livida Mot- schulsky, 1858, dello Sri Lanka. Infatti entrambe le specie hanno il pezzo copulatore dell’edeago lungamente sporgente dall’orifizio apicale e il sesto urotergo libero del maschio ha un lobo mediano in entrambe le specie. La differenze tuttavia sono nume- rose e alcune appariscenti. La nuova specie ha un'appendice ventrale dell’edeago presso la “crista apicalis”, assente in livida e la porzione preapicale dell’apice dell’edeago, in visione laterale, è larga nella nuova specie e stretta in livida. Il lobo mediano del margine posteriore del sesto urotergo libero del maschio della nuova specie ha a ciascun lato una debole prominenza, mentre in livida a ciascun lato di detto lobo mediano vi è una lunga spina a base larga. Gyrophaena (Phaenogyra) cooteri sp. n. Figg. 53-57 Tipi. Holotypus d, China, Jiangsu Prov., Nanjing Zijinshan, 8.V.1996, J. Cooter leg. (MHNG). Paratypi: 3 es., stessa provenienza. DESCRIZIONE. Lunghezza 1,4 mm. Corpo lucido e bruno; antenne gialle con antenno- meri 9 e 10 di un giallo sporco; zampe gialle. La reticolazione del capo, del pronoto e dell'addome è svanita, quella delle elitre è distinta. La punteggiatura del capo è poco distinta. I tubercoletti che coprono la superficie del pronoto sono molto superficiali, quelli delle elitre e dell'addome sono distinti. Edeago figg. 54-55, spermateca fig. 56, sesto urotergo libero del maschio fig. 57. COMPARAZIONI. La nuova specie per la forma dell’edeago e del margine posteriore del sesto urotergo libero del maschio, mostra affinità tassonomiche con G. obscurella Cameron, 1939, dell’ India settentrionale. Ma le differenze morfologiche tra le due specie sono molteplici. Le più vistose sono: la parte preapicale ventrale dell’edeago della nuova specie, in visione ventrale, è sensibilmente allargata, mentre è a lati rigo- rosamente paralleli tra loro in obscurella. Il margine posteriore del sesto urotergo libero del maschio della nuova specie mostra due corti dentini mediani inquadrati a ciascun lato da un largo lobo, mentre in obscurella i denti mediani sono piuttosto lunghi e a ciascun lato esiste una lunga spina a base stretta. ETIMOLOGIA. Specie dedicata al suo raccoglitore Jonathan Cooter di Hereford (Gran Bretagna), noto studioso di Liodidae. Gyrophaena (Phaenogyra) cristifera sp. n. Figg. 58-61 Tipo. Holotypus d, China, Yunnan, Xishuangbanna, 20.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Corpo lucido e giallo-rossiccio con elitre e addome di un giallo sporco; antenne giallo-rossicce con i due antennomeri basali gialli; zampe gialle. La reticolazione del capo e delle elitre è distinta, quella del pronoto e dell’addome è assai svanita. La punteggiatura del capo è superficiale e assente sulla fascia longitudinale mediana: si notano due impressioni circolari sulla fronte. I tuber- coletti che coprono la superdicie del pronoto sono poco distinti: distinti sono al 170 ROBERTO PACE Fico. 53-61 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 53:57: Gyrophaena (Phaenogyra) cooteri sp. n.; 58-61: Gyrophaena (Phaenogyra) cristifera sp. n. ALEOCHARINAE DELLA CINA 171 contrario i tubercoletti della superficie delle elitre. Edeago figg. 59-60, sesto urotergo libero del maschio fig. 61. COMPARAZIONI. La nuova specie ha il margine posteriore del sesto urotergo libero del maschio evidentemente di struttura simile a quello di G. obcurella Cameron, 1939, dell’ India: due denti mediani sporgenti all'indietro, inquadrati da due lunghe spine late- rali. La taglia corporea delle due specie però è molto differente: 1,1 in obscurella e 1,8 nella nuova specie. La struttura dell’edeago è ancor più differente nelle due specie: l’unica corta appendice ventrale dell’edeago di obscurella, nella nuova specie è sosti- tuita da due lunghe appendici ricurve, che insieme all’apice dell’edeago simulano una cresta. Gyrophaena (s. str.) gonggana sp. n. Figg. 62-64 Tipo. Holotypus d, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VIL. 1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e rossiccio scuro, con lati del pronoto e addome, tranne il quarto urite libero che è bruno, rossicci; antenne giallo-brune con i tre antennomeri basali gialli; zampe giallo-rossicce. La punteggiatura del capo è composta di punti enormi e assenti sulla linea mediana; quella del pronoto è composta da punti medi e grandi e quella delle elitre è composta di punti irregolarmente distri- buiti e distinti. Il capo e le elitre non presentano reticolazione, il pronoto e l’addome hanno reticolazione molto svanita. Il sesto urotergo libero del maschio mostra un tubercolo mediano posteriore compresso. Edeago figg. 63-64. COMPARAZIONI. Dato che l’edeago è molto espanso lateralmente nella regione pre- apicale, la nuova specie mostra affinità tassonomiche sia con G. multiplex Pace, 1989a, che a G. sitalaiana Pace, 1989a, entrambe del Nepal. Ne è chiaramente dis- tinta per la corta appendice apicale dell’edeago, assente in multiplex e lunghissima in sitalaiana, e per numerosi altri caratteri minori. Gyrophaena (s. str.) facilis sp. n. Figg. 65-69 Tipi. Holotypus 4, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII.1994, A. Smetana leg. (MHNG). Paratypi: 4 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,2 mm. Corpo lucido e bruno con lati del pronoto gialli; antenne brune con i tre antennomeri basali di un giallo sporco; zampe gialle. La reti- colazione del capo e delle elitre è svanita, quella dell'addome è netta e quella del pronoto assente. La punteggiatura del capo è svanita, quella del pronoto distinta, ma assente ai lati e quella delle elitre è poco distinta. Il quinto urotergo libero del maschio non mostra alcuna reticolazione. Edeago figg. 66-67, spermateca fig. 68, sesto uro- tergo libero del maschio fig. 69. COMPARAZIONI. La nuova specie sicuramente è tassonomicamente molto vicina a G. difficilis Cameron, 1939, dell’India settentrionale. Se ne differenzia, tra l’altro, per il pronoto nettamente meno trasverso, per l’appendice preapicale dell’edeago, corta e robusta, e non lunghissima come in difficilis, e per la taglia corporea maggiore: 2,2 mm invece di 1,6 mm come in difficilis. 722 ROBERTO PACE Fico. 62-69 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 62-64: Gyrophaena (s. str.) gonggana sp. n.; 65-69: Gyrophaena (s. str.) facilis sp. n. ALEOCHARINAE DELLA CINA 173 Gyrophaena (s. str.) chinensis sp. n. Figg. 70-74 Tipi. Holotypus d, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.V111.1994, A. Smetana leg. (MHNG). Paratypi: 7 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,6 mm. Corpo lucido e bruno (lievemente immaturo), con lati del pronoto, base ed estremità addominale bruno-rossicci; antenne rossicce con 1 tre antennomeri basali gialli; zampe gialle. L’avancorpo è privo di reticolazione, l’addome presenta una reticolazione svanita. La punteggiatura del capo e del pronoto è netta e profonda, quella delle elitre è distinta. Gli uroterghi basali presentano tubercoletti svaniti. Il quinto urotergo libero del maschio presenta un largo tubercolo mediano compresso. Edeago figg. 71-72, spermateca d fig. 73, sesto urotergo libero del maschio fig. 74. COMPARAZIONI. La nuova specie mostra lontane affinità tassonomiche con G. quadri- fida Cameron, 1939, del Kashmir e del Nepal, se si osserva la forma dell’edeago. Però l’appendice apicale dell’edeago è più larga e meno lunga nella nuova specie e 1 pezzi copulatori dell’edeago della nuova specie sono molto più lunghi. Inoltre il sesto uro- tergo libero del maschio della nuova specie ha i denti mediani molto corti e non lun- ghissimi come in quadrifida. Gyrophaena (s. str.) vidua sp. n. Figg. 75-76 Tipi. Holotypus 9, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VII.1994, A. Smetana leg. (MHNG). Paratypi: 2 2 9, stessa provenienza. DESCRIZIONE. Lunghezza 2,7 mm. Corpo lucido e bruno-rossiccio; antenne bruno- rossicce con i tre antennomeri basali e l'undicesimo gialli; zampe gialle. La reticola- zione del capo è estremamente svanita, quella del pronoto è distinta, quella delle elitre è assente e quella dell'addome è netta. La punteggiatura del capo e del pronoto è profonda e quella delle elitre è irregolare e distinta. Spermateca fig. 75. COMPARAZIONI. La spermateca della nuova specie ha forma molto simile a quella di G. densicollis Cameron, 1939, dell’India settentrionale e del Nepal. Tuttavia la sper- mateca della nuova specie è molto più sviluppata, con bulbo distale ovale più tras- verso. Inoltre il pronoto della nuova specie presenta punteggiatura irregolarmente distribuita e profonda e non uniformemente distribuita come quella di densicollis. Gyrophaena (s. str.) beijingensis sp. n. Figg. 77-80 Tipo. Holotypus 4, China, Beijing, Xiaolongmen, 1100-1500 m, 1.VII.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Corpo lucido e giallo-rossiccio sporco con capo bruno; antenne giallo-brune con i tre antennomeri basali gialli; zampe gialle. La reti- colazione del capo è estremamente svanita, quella del pronoto è svanita, quella delle elitre è distinta e quella dell’addome è quasi netta. La punteggiatura del capo e del pronoto è profonda, quella delle elitre è meno netta di quella del pronoto, ben distinta. Edeago figg. 78-79, sesto urotergo libero del maschio fig. 80. 174 ROBERTO PACE Imm Fico. 70-76 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 70-74: Gyrophaena (s. str.) chinensis sp. n.; 75-76: Gyrophaena (s. str.) vidua sp. n. ALEOCHARINAE DELLA CINA 175 COMPARAZIONI. La nuova specie, per la forma dell’edeago, è avvicinabile tassono- micamente sia a G. thoracica Cameron, 1939, dell India settentrionale e del Nepal, che a G. elegans Pace, 1989, del Nepal. Tuttavia la nuova specie mostra l’edeago con struttura più simile a elegans. G. beijingensis sp. n. si distingue da elegans per l’inca- vatura ventrale dell’edeago molto stretta (ampia in elegans), per il pezzo copulatore del sacco interno terminante in un flagello lunghissimo (senza flagello in elegans), e per le due lunghe spine al margine posteriore del sesto urotergo libero del maschio (due lobi in elegans). Gyrophaena (s. str.) vexillifera sp. n. Figg. 81-85 Tipi. Holotypus d, Hong Kong, Tai Po, flight interception, V.1996, de Rougemont leg. (MHNG). ) Paratypus: 1 ©, stessa provenienza. DESCRIZIONE. Lunghezza 1,7 mm. Corpo lucido e bruno con pronoto e base delle elitre giallo-bruni e con addome giallo-rossiccio con quarto urite rossiccio; antenne rossicce con i tre antennomeri basali gialli; zampe gialle. La reticolazione del capo è distinta sul disco e svanita ai lati, quella del pronoto è netta sulla fascia mediana e assente ai lati, sulle elitre è svanita e sull’addome è svanita sui due uroterghi basali: sui tre seguenti la reticolazione è distinta e netta alla base di ciascuno. La punteggia- tura del capo è svanita. I tubercoletti che stanno sul pronoto sono molto salienti e quelli delle elitre sono distinti. Edeago figg. 82-83, sesto urotergo libero del maschio fig. 85. COMPARAZIONI. Per l’habitus e per la forma dell’edeago, la nuova specie mostra alcune affinità tassonomiche con G. densicollis Cameron, 1939, dell’India setten- trionale e del Nepal. Il pezzo copulatore molto sporgente dall’orifizio apicale dell’e- deago di densicollis, non è così dilatato alla metà apicale quanto quello della nuova specie e la debole bozza preapicale ventrale dell’edeago della nuova specie, in densi- collis è diventata una lunga appendice stretta. Il sesto urotergo libero del maschio di densicollis ha solo due dentini marginali mediani, mentre nella nuova specie i denti sono come da fig. 85. Gyrophaena (s. str.) penetrans sp. n. Figg. 86-88 Tipo. Holotypus 2, Hong Kong, de Rougemont leg. (MHNG). DESCRIZIONE. Lungh. 1.8 mm. Corpo lucido e giallo (un po’ immaturo), con capo ed elitre giallo-bruni; antenne brune con i tre antennomeri basali gialli e i seguenti fino all’ottavo rossicci; zampe gialle. La reticolazione del capo e del pronoto è molto sva- nita, quella delle elitre è distinta e quella dell’addome è superficiale. La punteggiatura del capo e del pronoto è netta e profonda, quella delle elitre è superficiale. Spermateca fig. 87, sesto urotergo libero della femmina fig. 88. COMPARAZIONI. In base alla forma della spermateca, la nuova specie è probabilmente tassonomicamente vicina a G. quadrifida Cameron, 1939, del Kashmir e del Nepal. Se ne distingue per avere il bulbo distale della spermateca il doppio più sviluppato, 176 ROBERTO PACE F1GG. 77-85 Habitus, edeago in visione laterale e ventrale, sesto urotergo libero del maschio e spermateca. 77-80: Gyrophaena (s. str.) beijingensis sp. n: 81-85: Gyrophaena (s. str.) vexillifera sp. n. ALEOCHARINAE DELLA CINA 1077 nonostante la taglia corporea sia minore nella nuova specie (1,8 mm) e maggiore in quadrifida (2,2 mm). Inoltre il pronoto della nuova specie è più trasverso, le elitre meno larghe rispetto al pronoto e la reticolazione della superficie delle elitre della nuova specie è distinta, mentre quella delle elitre di quadrifida è estremamente sva- nita. Gyrophaena (s. str.) herebi sp. n. Figg. 89-91 Tipo. Holotypus 2, Hong Kong, Tai Po, V.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,5 mm. Corpo lucido e nero-bruno con addome rossiccio avente bruni i lati degli uriti liberi terzo, quarto e quinto; antenne di un giallo sporco con i tre antennomeri terminali bruni; zampe giallo-rossicce con tibie medie e poste- riori brune a metà. Spermateca fig. 90, sesto urotergo libero della femmina fig. 91. COMPARAZIONI. Specie che per la taglia corporea ridotta e per il bulbo distale della spermateca, è probabilmente affine a G. immatura Kraatz, 1859, dello Sri Lanka. Tuttavia il bulbo distale della spermateca della nuova specie è il doppio più sviluppato rispetto quello di immatura e il colore del corpo è nero-bruno con addome rossiccio nella nuova specie e giallo rossiccio con elitre bruno-rossicce in immatura. Gyrophaena (s. str.) paula sp. n. Figg. 92-94 Tipi. Holotypus 9, China, Yunnan, Xishuangbanna, Chayanhe F.S., 24.1.1993, de Rougemont leg. (MHNG). Paratypus: 1 ©, stessa provenienza. DESCRIZIONE. Lunghezza 1,5 mm. Corpo lucido e nero-bruno con pronoto bruno e addome giallo sporco avente gli uriti liberi quarto e quinto giallo-bruni; antenne brune con i due antennomeri basali di un giallo sporco; zampe gialle. L'intero corpo non mostra alcuna reticolazione. La punteggiatura del capo è composta di punti grandi allineati sul disco e di punti medi sul resto della superficie. La punteggiatura del pro- noto è sparsa e distinta e quella delle elitre è netta e meno sparsa di quella del pro- noto. Spermateca fig. 93, sesto urotergo libero della femmina fig. 94. COMPARAZIONI. La nuova specie per la taglia corporea, per l’habitus e per la forma della spermateca, è simile a G. livida Motschulsky, 1858, dello Sri Lanka. Però è spe- cie ben distinta per avere le elitre evidentemente più corte rispetto al pronoto, gli occhi più sporgenti, l’addome non reticolato (addome distintamente reticolato in livida) e per la spermateca con bulbo distale più sviluppato, con appendice inserita presso il bulbo prossimale, larga e corta (stretta e lunga in livida). Encephalus chinensis sp. n. Figg. 95-98 Tipo. Holotypus d, China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII. 1994, A. Smetana leg. (MHNG). DescRIZIONE. Lunghezza 2,4 mm. Corpo lucido, molto convesso con addome scafoide e nero con margine posteriore degli uroterghi rossiccio; antenne brune; zampe ros- sicce con femori bruni. La reticolazione del capo e del pronoto è assente; quella delle 178 ROBERTO PACE ss. Imm Fico. 86-94 Habitus, spermateca e sesto urotergo libero della femmina. 86-88: Gyrophaena (s. str.) penetrans sp. n.; 89-91: Gyrophaena (s. str.) herebi sp. n.; 92-94: Gyrophaena (s. str.) paula sp. n. ALEOCHARINAE DELLA CINA 179 elitre è svanita e quella dell’addome è netta. La punteggiatura del capo e dell’addome è netta e profonda, quella delle elitre è distinta e irregolarmente distribuita, assente ai lati. Edeago figg. 96-97, sesto urotergo libero del maschio fig. 98. COMPARAZIONI. Per la forma dell’edeago, la nuova specie si presenta simile a E. hima- laviensis Pace, 1987, del Nepal. E. chinensis sp. n. si distingue da questa specie per la punteggiatura del corpo più rada, per quella delle elitre per nulla grossolana come in himalayiensis, per 1 due denti mediani al margine posteriore del sesto urotergo libero del maschio per nulla prolungati come in himalayiensis, ma corti. L’edeago della nuova specie è meno ricurvo al lato ventrale e il pezzo copulatore del sacco interno dello stesso organo è robusto e sinuato all’apice nella nuova specie e sottile e retto all'apice in himalayiensis. HOMALOTINI Silusa (s. str.) chinensis sp. n. Figg. 99-102 Tipi. Holotypus 4, China, Zhejiang, Tienmushan, 29.1V.1993, de Rougemont leg. (MHNG). Paratypi: 4 es. stessa provenienza. DESCRIZIONE. Lunghezza 3,2 mm. Corpo lucido; capo bruno, pronoto bruno-rossiccio, con margini laterali e posteriore rossicci, elitre di un giallo sporco con 1 lati esterni bruni, addome bruno con margine posteriore degli uroterghi e lati degli uroterghi liberi primo, secondo e terzo, rossicci; antenne bruno-rossicce; zampe gialle. La reti- colazione del capo è assente, quella sul resto del corpo è estremamente svanita. La punteggiatura del capo e del pronoto è molto svanita, quella delle elitre è netta, ma assente lungo il margine posteriore dove si osservano tubercoletti salienti. Edeago figg. 100-101, spermateca fig. 102. COMPARAZIONI. La nuova specie è distinta da S. indica Cameron, 1939, dell’ India, per gli occhi ridotti, le tempie molto più lunghe e per le elitre più sviluppate. E° pure distinta da S. aliena Bernhauer, 1916, della Cina, per avere i pezzi copulatori del sacco interno dell’edeago lungamente sporgenti dall’orifizio apicale dell’edeago stesso (appena sporgenti in aliena). Silusa (s. str.) smetanai sp. n. Figg. 103-104 Tipo. Holotypus 9, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII. 1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e bruno con margine posteriore dei tre uroterghi basali rossiccio come l’estremità addominale; antenne bruno-rossicce con 1 due antennomeri basali rossicci; zampe rossicce. Su tutto il corpo non vi è presenza di reticolazione. La punteggiatura ombelicata del capo è netta e assente sul disco. Il pronoto e le elitre sono coperti da distinti e fini tubercoletti. Spermateca fig. 104. COMPARAZIONI. La nuova specie è ben distinta dalla precedente S. chinensis sp. n., per l'assenza di impressione mediana basale del pronoto e per l’introflessione apicale del bulbo distale della spermateca, larghissimo e non strettissimo come in chinensis. La 180 ROBERTO PACE Fico. 95-102 Habitus, edeago in visione laterale e ventrale, sesto urotergo libero del maschio e spermateca. 95-98: Encephalus chinensis sp. n.; 99-102: Silusa (s. str.) chinensis sp. n. ALEOCHARINAE DELLA CINA 181 nuova specie è pure distinta da S. indica Cameron, 1939, dell’India, per avere tra l’altro, il quarto antennomero trasverso e non lievemente più lungo che largo, come si osserva in indica. Silusa (s. str.) cooteri sp. n. Figg. 105-108 Tipi. Holotypus d, China, Zhejiang Prov., Lin’an County, 1000 m, W. Tanmu Shan N.R., 18.V.1996, J. Cooter leg. (MHNG). Paratypus: 1 9, stessa provenienza. DESCRIZIONE. Lunghezza 2,6 mm. Corpo lucido e bruno; antenne e zampe bruno- rossicce. Su tutto il corpo non vi é traccia di reticolazione. La punteggiatura del capo è netta, quella delle elitre è distinta e a raspa. I tubercoletti che coprono la superficie del pronoto sono fini, quelli dell’addome sono distinti. Edeago figg. 106-107, sper- mateca fig. 108. COMPARAZIONI. La nuova specie è distinta da S. aliena Bernhauer, 1916, della Cina, per avere taglia minore, edeago di un terzo minore e pezzi copulatori del sacco interno meno robusti, di cui uno profondamente incavato presso l’apice (non incavato in aliena). ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore Jonathan Cooter di Here- ford, noto studioso di Liodidae. Taraktomora gen. n. Figg. 109-111 DIAGNOsI. Genere assai affine al genere Silusa Erichson, 1837, per avere la ligula intera e palpi labiali allungati; distinta per la ligula nettamente più lunga del primo articolo dei palpi labiali e con un rigonfiamento basale su cui stanno quattro pori sen- sitivi e per la spermateca che presenta bulbo distale non sferico come si riscontra nelle varie specie note di Silusa. E’ mia opinione che questo nuovo genere si colloca tassonomicamente in posizione intermedia tra il genere Silusa Er. e il genere Leptusa Kr. che pure presenta specie a ligula intera e molto lunga. Inoltre il tipo di spermateca del nuovo genere è associabile a quello di moltissime specie di Leptusa. Tuttavia i palpi labiali sono eccessivamente lunghi nel nuovo genere per attribuire la specie al genere Leptusa. DESCRIZIONE. Corpo convesso; lobo interno ed esterno dei palpi mascellari come in Silusa; palpi labiali di 2 articoli, meno lunghi che in Silusa; ligula nettamente più lunga del primo articolo dei palpi labiali, con rigonfiamento basale su cui stanno quattro pori sensitivi; paraglosse nulle, mento come in Silusa (fig. 111); 11 antenno- meri; occhi molto sviluppati; pronoto molto trasverso, in avanti più ristretto che all’indietro; elitre a lati paralleli; processo mesosternale ad angolo acuto, sicché le mesocoxe sono tra loro contigue, come in Silusa e Leptusa; formula tarsale 4-4-5; primo tarsomero posteriore appena più lungo del seguente; bulbo distale della spermateca subreniforme. ETIMOLOGIA. Il nome del nuovo genere significa “Confinante e conturbante”, ciò a significare che Taraktomora gen. n. si pone in posizione tassonomica tra Silusa e 182 ROBERTO PACE Fico. 103-110 Habitus, spermateca ed edeago in visione laterale e ventrale. 103-104: Silusa (s. str.) smetanai sp. n.; 105-108: Silusa (s. str.) cooteri sp. n.; 109-110: Taraktomora orientis gen. n., Sp. n. ALEOCHARINAE DELLA CINA 183 Leptusa e che per questo può sconvolgere l’ordine stabilito in tassonomia, dato che può essere considerato “l’anello di congiunzione” tra i generi Silusa e Leptusa. GENERE GRAMMATICALE. Taraktomora gen. n. è di genere femminile. TYPUS GENERIS. Taraktomora orientis sp. n. Taraktomora orientis sp. n. Figg. 109-111 Tipo. Holotypus 9, Hong Kong, Kadoorie Agricultural Research Centre, VIII.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,9 mm. Corpo lucido; capo bruno-rossiccio, pronoto giallo- rossiccio, elitre brune con base e sutura giallo-brune, addome giallo-rossiccio con uriti liberi quarto e metà basale del quinto bruni; antenne bruno-rossicce con i tre anten- nomeri basali e l'undicesimo giallo-rossicci; zampe gialle. La reticolazione del capo è assente, quella sul resto del corpo è estremamente svanita. La punteggiatura ombeli- cata del capo è fitta e assai svanita, quella del pronoto è superficiale e quella delle elitre è distinta. Spermateca fig. 110, labio con palpo labiale fig. 111. Coenonica absurda sp. n. Figg. 112-116 Tipi. Holotypus ¢, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, VIII.1996, de Rougemont leg. (MHNG). Paratypi: 1 4 e 1 2, Hong Kong, Chinese University, in Auricularia fungus, 1.IX.1996; 1 4, Hong Kong, N.T., IX.1996; 1 ©, Hong Kong, Tai Po, VII.1996, tutti de Rougemont leg. DESCRIZIONE. Lunghezza 2,7 mm. Avancorpo debolmente lucido, addome lucido. Corpo bruno con addome bruno-rossiccio; antenne brune con 1 tre antennomeri basali e l’undicesimo bruno-rossicci; zampe gialle. La reticolazione del capo è vigorosa, quella del pronoto è svanita; assente è sul resto del corpo. La fronte è coperta da punteggiatura ombelicata distinta su un fondo a reticolazione svanita, sul resto della superficie del capo la punteggiatura è cancellata dalla vigorosa reticolazione. Il pronoto presenta ai lati una netta rugosità, sul resto della superficie sta una punteggiatura distinta; la fossetta posteriore è profonda. Le elitre sono coperte di tubercoletti molto salienti. Edeago figg. 112-113, spermateca fig. 114, sesto urotergo libero del maschio fig. 115. COMPARAZIONI. La nuova specie mostra un carattere che si riscontra anche in C. angusticollis Cameron, 1920: un tubercolo mediano posteriore sul quinto urotergo libero del maschio. Nonostante la presenza di esso, per i caratteri dell’edeago e della spermateca, le differenze tra le due specie sono marcate. L’edeago della nuova specie ha forma unica nell’insieme delle specie note, così è per la spermateca. Coenonica longwangensis sp. n. Figg. 117-120 Tipi. Holotypus d, China, Zhejiang Prov., Anji County, ca. 400 m, Long Wang Shan N.R., 13-14.V.1996, J. Cooter leg. (MHNG). Paratypus: 1 ©, China, Zhejiang Prov., Lin’an County, 1000 m, W. Tianmu Shan N.R., 18.V.1996, J. Cooter leg. DESCRIZIONE. Lunghezza 2,1 mm. capo e pronoto opachi, elitre debolmente lucide, addome lucido; corpo bruno con addome bruno-rossiccio; antenne brune con i due 184 ROBERTO PACE FIGG. 111-120 Labio con palpo labiale, edeago in visione laterale e ventrale, spermateca, sesto urotergo libero del maschio e habitus. 111: Tataktomora orientis gen. n., sp. n.; 112-116: Coenonica absurda sp. n.; 117-120: Coenonica longwangensis sp. n. ALEOCHARINAE DELLA CINA 185 antennomeri basali bruno-rossicci; zampe gialle. Il capo e il pronoto sono coperti da fittissima punteggiatura, tanto da dare un aspetto rugoso alla superficie, tranne tra le antenne dove non vi è punteggiatura e la superficie è lucida; il pronoto ha una debole depressione mediana posteriore; i tubercoletti che coprono le elitre sono fittissimi e confluenti tra loro, sì da formare linee logitudinali interrotte. Gli uroterghi presentano tubercoletti salienti e reticolazione molto svanita. Edeago figg. 117-118, spermateca fig. 109: COMPARAZIONI. C. longwangensis Sp. n. è più affine a C. ming Pace, 1993, della Cina, che a C. javana Bernhauer, 1914, di Giava, Hong Kong, Malesia e Filippine. Ciò in base alla forma dell’edeago e dell’habitus. C. javana ha pronoto poco trasverso ed edeago appiattito al lato ventrale, mentre la nuova specie e C. ming hanno pronoto nettamente trasverso. La nuova specie differisce da ming per avere gli occhi più sviluppati, sicché le tempie sono più corte, il pronoto sinuato davanti agli angoli pos- teriori, l’edeago meno sviluppato, con incavatura ventrale meno ampia e con “crista apicalis” appena distinta (ben distinta in ming) e differente forma dei pezzi copulatori del sacco interno dell’edeago stesso. Coenonica angularis sp. n. Figg. 121-122 Tipi. Holotypus 9, China, Zhejiang Prov., Lin’an County, 350 m, W. Tianmu Shan N.R., 16-22.V.1996, J. Cooter leg. (MHNG). Paratypus: 1 ©, stessa provenienza. DESCRIZIONE. Lunghezza 3,0 mm. Avancorpo debolmente lucido, addome lucido. Corpo rossiccio con capo e uriti liberi quarto e quinto bruni; antenne brune con 1 tre antennomeri basali e l’apice dell’undicesimo giallo-bruni; zampe giallo-rossicce. La reticolazione del capo è netta, quella del pronoto è molto svanita, quella delle elitre e dell’addome è assente: solo nel fondo dei solchi trasversi basali degli uroterghi la reti- colazione è netta. La punteggiatura del capo è netta e fitta, assente in avanti, quella del pronoto è indistinta, tranne dei robusti punti basali. Le elitre mostrano una pun- teggiatura netta e profonda solo sui due terzi anteriori, sul terzo posteriore la punteggiatura è fine e svanita. Spermateca fig. 122. COMPARAZIONI. Specie simile a C. exuta Pace, 1984, del Nepal, sia per l’habitus, che per la forma della spermateca. Entrambe sono avvicinabili tassonomicamente a C. puncticollis Kraatz, 1857, specie a larga distribuzione geografica. La nuova specie ha spermateca di dimensione troppo riddotta, rispetto quella di puncticollis e il sesto uro- tergo libero della femmina è privo di denti marginali, per essere considerata specie molto prossima. La nuova specie è invece molto più affine a exuta, da cui si distingue per la taglia corporea maggiore (3,0 mm invece di 2,4 mm), per il pronoto privo di punteggiatura (pronoto coperto di tubercoletti salienti in exuta), per le elitre più lunghe, rispetto al pronoto, per il fondo punteggiato dei solchi trasversi basali degli uroterghi (non punteggiato in exuta) e per il bulbo prossimale della spermateca netta- mente conformato, anche se poco sviluppato (pressoché nullo in exuta). Coenonica parens sp. n. Figg. 123-125 Tipo. Holotypus d, China, Zhejiang, Tienmushan, 29.1V.1993, de Rougemont leg. (MHNG). 186 ROBERTO PACE DESCRIZIONE. Lungh. 2,1 mm. Capo e pronoto opachi resto del corpo lucido. Capo, pronoto ed elitre, tranne la loro sutura, nero-bruni, fascia suturale delle elitre bruno- rossiccia, addome bruno-rossiccio con gli uriti liberi quarto e quinto bruni; antenne nere con antennomero basale nero-bruno; zampe giallo-rossicce. Sul capo e sul pro- noto i tubercoletti della superficie sono così contigui tra loro e così salienti, sì da dare un aspetto rugoso alla superficie. La punteggiatura delle elitre è distinta, posta su un fondo non reticolato come quello degli uroterghi. Edeago figg. 123-124. COMPARAZIONI. Specie del gruppo di C. ming Pace, 1993, della Cina, più affine a questa specie che a C. longwangensis sp. n. sopra descritta, per avere gli occhi lunghi quanto le tempie (e non più lunghi come in /ongwangensis), per avere il pronoto non sinuato davanti agli angoli posteriori e per avere l’edeago ampiamente ricurvo al lato ventrale. La nuova specie differisce da C. ming per avere l’edeago ancor più ampia- mente ricurvo al lato ventrale e il pezzo copulatore del sacco interno dell’edeago stesso, non dilatato e ricurvo all’ apice. Coenonica semimutata sp. n. Figg. 126-129 Tipo. Holotypus dé, China, Yunnan, Xishuangbanna, Sanchaha, elephant res., 24.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,3 mm. Avancorpo opaco, addome lucido. Corpo bruno- rossiccio con capo e lati esterni delle elitre bruni; addome giallo-rossiccio con gli uriti liberi quarto e quinto rossicci; antenne brune con 1 due antennomeri basali e l’apice dell’undicesimo rossicci; zampe giallo-rossicce. Il capo e il pronoto sono coperti di contigui e grossolani tubercoli che danno un aspetto rugoso alla superficie. La pun- teggiatura delle elitre è fittissima, a punti contigui fra loro e netti. Tubercoletti salienti stanno sulla superficie degli uroterghi. che non è reticolata. Edeago figg. 127-120, sesto urotergo libero del maschio fig. 129. COMPARAZIONI. La nuova specie è sicuramente molto affine a C. mutata Pace, 1984, del Nepal, se si osserva l’habitus e la struttura generale dell’edeago. Se ne distingue per l’edeago più esile, con “crista apicalis” molto più lunga. Il profilo ventrale dell’edeago è bisinuato e dentellato nella nuova specie (arcuato e senza dentini in mutata), l’apice è meno acuto e i pezzi copulatori del sacco interno sono più distesi e meno robusti, con tubulo mediano distinto (assente in mutata). Inoltre il margine posteriore del sesto urotergo libero del maschio, presenta due dentini accostati fra loro e altri denti come da fig. 129, mentre C. mutata mostra 8 denti uguali regolarmente distanziati fra loro, inquadrati da due lunghe spine laterali a base stretta (spine corte e a base larga nella nuova specie). Coenonica truncata sp. n. Figg. 130-134 Tipi. Holotypus d, China, Yunnan, Xishuangbanna, Mengdian, 26.1.1993, de Rouge- mont leg. (MHNG). Paratypi: 20 es., stessa provenienza. DESCRIZIONE. Lunghezza 3,7 mm. Capo opaco tranne il disco che è lucido, resto del corpo lucido. Capo nero, pronoto bruno, elitre di un giallo sporco, addome giallo- ALEOCHARINAE DELLA CINA 187 FIGG. 121-129 Habitus, spermateca, edeago in visione laterale e ventrale e sesto urotergo libero del maschio. 121-122: Coenonica angularis sp. n.; 123-125: Coenonica parens sp. n.; 126-129: Coenonica semimutata Sp. n. 188 ROBERTO PACE rossiccio con uriti liberi quarto e la metà basale del quinto rossicci; antenne brune con i due antennomeri basali e la base del terzo giallo-rossicci; zampe gialle. Solo sul disco del capo la punteggiatura è robusta e profonda: sul resto della superficie la punteggiatura è indistinta data la scabrosità della superficie stessa. La punteggiatura del pronoto è netta e più densa in avanti sulla linea mediana, posta su un fondo non reticolato. La punteggiatura delle elitre è svanita. Elitre e addome sono privi di reti- colazione. Edeago figg. 131-132, spermateca fig. 133; sesto urotergo libero del maschio fig. 134. COMPARAZIONI. La nuova specie, per l’habitus e per il reniforme bulbo distale della spermateca, è sicuramente affine a C. pendleburyi Cameron, 1936, della Penisola Malese. Se ne distingue per la taglia corporea maggiore (2,9 mm in pendleburyi), per gli occhi più sviluppati, per il quarto antennomero nettamente trasverso (lungo quanto largo in pendleburyi), per avere il pronoto nettamente punteggiato, con punti adden- sati in avanti a metà (pronoto coperto di tubercoli superficiali in pendleburyi) e per la spermateca meno sviluppata (nonostante la taglia corporea maggiore), con bulbo distale reniforme regolare (e non ristretto verso l'apice come in pendleburyi, il cui holotypus unico è una femmina). Coenonica aliena sp. n. Figg. 135-136 Tipo. Holotypus 9, China, Yunnan, Dali, 9.11.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 3,3 mm. Capo e pronoto debolmente opachi, resto del corpo lucido. Capo nero-bruno, pronoto bruno con margini laterali e posteriore bruno-ros- sicci, elitre giallo-brune, compresa la sutura con terzo posteriore bruno, addome rossiccio con uriti liberi quarto e quinto bruni; antenne brune con antennomero basale rossiccio; zampe di un giallo sporco con tarsi e tibie anteriori e medie bruni. La reti- colazione del capo è netta e fine, quella del pronoto e dell’addome è assente e quella delle elitre è svanita. La punteggiatura del capo è netta e profonda solo sul disco: sul resto della superficie è svanita.La punteggiatura del pronoto è netta e fitta e presenta alcuni punti grossolani alla base, quella delle elitre è irregolarmente distribuita e fitta. Spermateca fig. 136. COMPARAZIONI. Per l’habitus, per il sistema di reticolazione del capo, per il pronoto molto trasverso e per la forma della spermateca, la nuova specie è probabilmente affine a C. varicornis (Kraatz, 1859), dello Sri Lanka. Se ne distingue per la taglia corporea maggiore (1,9 mm in varicornis), per le elitre coperte di punteggiatura irre- golarmente distribuita, invece di tubercoletti fitti come in varicornis e per il bulbo distale della spermateca tre volte maggiore e di forma non sferica (a sfera in vari- cornis). Coenonica zhejiangensis sp. n. Figg. 137-140 Tipi. Holotypus 4, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. (MHNG). Paratypi: 5 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,9 mm. Corpo lucido e bruno con margine posteriore degli uroterghi basali e l’estremità distale dell’addome rossicci; antenne bruno-rossicce; ALEOCHARINAE DELLA CINA 189 Imm 136 Fico. 130-136 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 130-134: Coenonica truncata sp. n.; 135-136: Coenonica aliena sp. n. 190 ROBERTO PACE zampe rossicce. La reticolazione del capo è vigorosa, tranne che sul disco dove è distinta, quella sul resto del corpo è assente. La punteggiatura del capo è ombelicata e netta sul disco, svanita ai lati; quella delle elitre è distinta. Il pronoto presenta superficie coperta di tubercoletti salienti, molto salienti in avanti e ai lati dove sono anche più fitti. Edeago figg. 138-139, spermateca fig. 140. COMPARAZIONI. Specie simile a C. ming Pace, 1993, della Cina. Ne è distinta per il pronoto e le elitre meno trasversi, per i punteggiati solchi trasversi basali degli uroterghi (senza punti in ming) e per l’edeago più ampiamente e meno profondamente ricurvo al lato ventrale, con apice, in visione ventrale, sinuato lateralmente (non sinuato in ming), con pezzo copulatore del sacco interno sottile (e non dilatato e ricurvo come in ming). Coenonica tianmushanensis sp. n. Figg. 141-144 Tipi. Holotypus ®, China, Zhejiang, Tianmushan, 29.1.1993, de Rougemont leg. (MHNG). Paratypi: 1 9, stessa provenienza; 1 d, China, Zhejiang, Lin’an County, 1000 m, W. Tianmu Shan N.R., 18.V.1996, J. Cooter leg. DESCRIZIONE. Lunghezza 2,4 mm. Corpe lucido e giallo-rossiccio con capo nero- bruno; antenne brune con 1 tre antennomeri basali e l’undicesimo, tranne la sua base, rossicci; zampe giallo-rossicce. La punteggiatura del capo in avanti è fine, sul disco è netta e profonda, all’indietro e ai lati è svanita, confusa nella vigorosa reticolazione che sul disco è assai svanita. Tubercoletti molto salienti coprono la superficie del pro- noto e delle elitre, su un fondo distintamente reticolato sul pronoto e a reticolazione svanita sulle elitre. Sull’addome non vi è reticolazione, tranne nel fondo dei solchi trasversi basali. Edeago figg. 141-142, spermateca fig. 143. COMPARAZIONI. Specie affine a C. varicornis (Kraatz, 1959), dello Sri Lanka, per l’habitus simile, per la struttura dell’edeago e della spermateca. E° chiaramente distinta da essa, per la taglia corporea maggiore (1,9 mm in varicornis), per il bulbo basale dell’edeago poco sviluppato (molto sviluppato in varicornis), per la “crista apicalis” dell’edeago stesso saliente (stretta e corta in varicornis) e per il maggiore sviluppo del bulbo distale della spermateca. Coenonica arcusifera sp. n. Figg. 145-148 Tipi. Holotypus dé, China, Yunnan, Xishuangbanna, Mengdien, 26.1.1993, de Rouge- mont leg. (MHNG). Paratypus: | S, China, Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e bruno con margini del pronoto ed elitre rossicci, addome bruno-rossiccio con margini posteriori degli uriti basali ros- sicci e uriti liberi quarto e quinto bruni; antenne brune con i tre antennomeri basali rossicci e con l’apice dell’undicesimo bruno-rossiccio; zampe rossicce. La pun- teggiatura del capo è composta di punti fitti tra loro contigui, più netti sul disco che sul resto della superficie cefalica dove gradualmente svaniscono fino a scomparire là dove la reticolazione è vigorosa, mentre sul disco la reticolazione è distinta. La ALEOCHARINAE DELLA CINA 191 Ol mm 142 141 Fico. 137-144 Habitus, edeago in visione laterale e ventrale e spermateca. 137-140: Coenonica zhejiangensis sp. n.; 141-144: Coenonica tianmushanensis sp. n. 192 ROBERTO PACE punteggiatura del pronoto è fitta, profonda e robusta, assente agli appuntiti angoli pos- teriori, posta su un fondo a reticolazione estremamente svanita. La punteggiatura delle elitre è netta, profonda, irregolarmente distribuita e assente presso gli angoli poste- riori: è posta su un fondo distintamente reticolato. La stria suturale delle elitre si allontana dalla sutura nella metà posteriore. Gli uroterghi non presentano reticola- zione e sono coperti di rada punteggiatura netta. Edeago figg. 146-147, sesto urotergo libero del maschio fig. 148. COMPARAZIONI. L’habitus della nuova specie è simile a quello di C. sharpi (Fauvel, 1901), del Giappone: solo le impressioni del pronoto sono più brevi nella nuova specie e le elitre hanno punteggiatura meno fitta e sono più lunghe, rispetto al pronoto e più brevi in sharpi. Purtroppo la serie tipica di sharpi è costituita da sole femmine, per cui non sono possibili comparazioni dell’ edeago. Coenonica yunnanensis sp. n. Figg. 149-150 Tipi. Holotypus 9, China, Yunnan, Xishuangbanna, Chayanhe F.P., 24.1.1993, de Rougemont leg. (MHNG). Paratypi: 3 2 2, stessa provenienza. DESCRIZIONE. Lunghezza 2,7 mm. Corpo lucido. Capo e pronoto neri, elitre nero- brune con base giallo-bruna, addome bruno con margine posteriore dei tre uriti basali rossiccio; antenne nero-brune con base ed estremità dei due antennomeri basali ros- sicce; zampe brune con tarsi giallo-rossicci. Su tutto il corpo non vi è traccia di reticolazione. La punteggiatura del capo è ombelicata, profonda e robusta. Il pronoto è coperto di tubercoli allungati e salienti che sono assenti lungo il margine posteriore. Le elitre presentano tubercoli molto salienti, più fitti e grossolani alla base e ai lati. Spermateca fig. 149. COMPARAZIONI. La nuova specie è affine a C. drescheri Cameron, 1939a, di Giava, sia per la robusta punteggiatura del capo e per la granulosità delle elitre, sia per la presenza di una sporgenza mediana all’orlo del solco basale degli uriti liberi terzo e quarto, sia per la forma globulare della spermateca. La nuova specie è però chiara- mente distinta da drescheri perché ha il pronoto coperto di tubercoli allungati salienti, mentre in drescheri il pronoto è coperto di punteggiatura svanita. Inoltre la sporgenza mediana dell’orlo dei solchi basali degli uriti liberi terzo e quarto è molto lunga (bre- vissima in drescheri) e la spermateca è di un terzo più sviluppata di quella di drescheri, nonostante la taglia corporea sia uguale nelle due specie. Stenomastax chinensis sp. n. Figg. 151-154 Tipi. Holotypus ¢, China, Yunnan, Xishuangbanna, Chayanhe F.P., 24.1.1993, de Rougemont leg. (MHNG). Paratypi: 8 es., stessa provenienza; | 4, Yunnan, Mengdian, 26.1.1993; 1 dé, China, Zhejiang, Tianmushan, 29.1V.1993; 2 dd e 2 9 9, China, Xishuangbanna, Jing Hong, 11.1993, tutti de Rougemont leg. DESCRIZIONE. Lunghezza 2,0 mm. Corpo debolmente lucido e bruno con elitre giallo brune aventi base e lati esterni bruni e con i due uriti basali e il margine posteriore del ALEOCHARINAE DELLA CINA 193 A dr } i Ka FiGG. 145-150 Habitus, edeago in visione laterale e ventrale, sesto urotergo libero del maschio e spermateca. 145-148: Coenonica arcusifera sp. n.; 149-150: Coenonica yunnanensis Sp. n. 194 ROBERTO PACE terzo urite libero, rossicci; antenne brune con antennomero basale giallo-bruno; zampe gialle. Sull’avancorpo indistinta è la reticolazione che sull’addome è estre- mamente svanita. La punteggiatura del capo è fittissima e distinta. Il pronoto e le elitre sono coperti di tubercoletti fittissimi e distinti. Edeago figg. 152-153, sper- mateca fig. 154. COMPARAZIONI. L’habitus della nuova specie è pressoché identico a quello di S. vari- ventris (Kraatz, 1859), largamente diffuso dalle Mascarene alla Nuova Guinea. L’ede- ago e la spermateca sono simili. Le differenze sono le seguenti: l’apice dell’edeago della nuova specie è più sottile se visto al lato ventrale e i pezzi copulatori del sacco interno sono molto sporgenti dall’orifizio apicale dell’edeago e descrivono un’ampia spira all’esterno. La parte prossimale della spermateca descrive una spira nella nuova specie e molte spire in variventris. Stenomastax raptoria sp. n. Figg. 155-158 Tipi. Holotypus d, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. (MHNG). Paratypi: | 9, stessa provenienza; 8 es., China, Yunnan, Xishuangbanna, Jinghong, 11.1993, de Rougemont leg. DESCRIZIONE. Lunghezza 2,0 mm. Corpo debolmente lucido, compresso e bruno con i due uriti basali e l'apice addominale brunoerossicci; antenne brune con i due anten- nomeri basali bruno-rossicci; zampe giallo-rossicce. La punteggiatura del capo è fittissima, composta di punti tra loro contigui e svaniti. La reticolazione del pronoto è netta, quella delle elitre e dell'addome è distinta. Il pronoto non presenta distinti tubercoletti o punteggiatura. I tubercolletti che coprono le elitre sono fini e distinti. I quattro uroterghi basali mostrano tubercoletti ben salienti, il quinto urotergo libero è coperto di tubercoletti svaniti. Edeago figg. 155-156, spermateca fig. 157. COMPARAZIONI. Per la forma dell’edeago, la nuova specie si mostra affine a S. cele- bensis Pace, 1986, di Celebes. Si distingue per avere l’apice dell’edeago molto più lungo e, se visto ventralmente, protratto a lati paralleli e non a lati convergenti, come in celebensis. Inoltre 1 pezzi copulatori del sacco interno sono nettamente più lunghi e con un gancio apicale, mentre in celebensis tale gancio è assente. Stenomastax diogenes sp. n. Figg. 159-162 Tipo. Holotypus ¢, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,0 mm. Capo e pronoto debolmente opachi, resto del corpo lucido. Capo e pronoto neri, elitre giallo-brune con lati esterni e base bruni, estremità addominale bruno-rossiccia; antenne nere con antennomero basale bruno; zampe rossicce. La reticolazione del capo e del pronoto è quasi vigorosa, quella delle elitre è svanita, quella dei tre uroterghi basali è estremamente svanita, quella del quarto urotergo libero è superficiale e quella del quinto è distinta. La punteggiatura del capo è ombelicata e distinta sul disco: sul resto della superficie è confusa nella reticola- zione quasi vigorosa. Il pronoto non presenta distinta punteggiatura, nè tubercoletti. I 195 ALEOCHARINAE DELLA CINA 01 mm 4 A 01 mm 197. 156 155 Figc. 151-158 Habitus, edeago in visione laterale e ventrale e spermateca. 151-154: Stenomastax chinensis Sp. n.; 155-158: Stenomastax raptoria sp. n. 196 ROBERTO PACE tubercoletti che coprono le elitre sono superficiali. Edeato figg. 160-161, sesto uro- tergo libero dal maschio fig. 162. COMPARAZIONI. Specie molto affine a S. platygaster (Kraatz, 1859), largamente distri- buito nella Regione Orientale. E’ distinta da esso per avere il pronoto più trasverso, il sesto urotergo libero del maschio assai poco incavato al margine posteriore (profon- damente incavato in platygaster) e per l’apice dell’edeago poco protratto, senza debole gancio apicale e, in visione ventrale, con apice tronco (e non con apice con debole gancio apicale e con apice non tronco come in platygaster). Stenomastax yunnanensis sp. n. Figg. 163-164 Tipi. Holotypus 2, China, Yunnan, Ruili, ca. 700 m, de Rougemont leg. (MHNG). Paratypus: | ©, stessa provenienza. DESCRIZIONE. Lunghezza 2,1 mm. Corpo debolmente lucido, un po’ depresso e bruno con capo e uriti liberi terzo, quarto e base del quinto neri; antenne brune con anten- nomero basale bruno-rossiccio; zampe gialle. La reticolazione del corpo è da poco distinta ad assente. La punteggiatura del capo è distinta e fittissima, quella del pronoto e delle elitre è assente per la presenza di una superficie d’aspetto reticolato. I tubercoletti degli uroterghi sono ben salienti. Spermateca fig. 163. COMPARAZIONI. In base alla forma della spermateca, la nuova specie è affine a S. vari- ventris (Kraatz, 1859) largamente diffuso dalle Mascarene alla Nuova Guinea. Ne è distinta per avere il bulbo distale della spermateca più sviluppato, con docce interne molto lunghe (corte in variventris) e per la parte prossimale della spermateca stessa descrivente due spire e non rettilinea, con stretta spira terminale come in variventris. Stenomastax pulcher sp. n. Figg. 165-168 Tipo. Holotypus 4, China, Yunnan, Xishuangbanna Mengdian, 26.1.1990, de Rouge- mont leg. (MHNG). DESCRIZIONE. Lunghezza 1,9 mm. Corpo lucido e rossiccio con capo e quarto urite libero bruni, elitre gialle con i tre quarti posteriori sfumati di bruno; antenne brune con i tre antennomeri basali giallo-rossicci; zampe gialle. La reticolazione del capo è netta, quella del pronoto è nettissima e quella delle elitre e del pronoto è svanita. La punteggiatura del capo è fitta e svanita, quella del pronoto è assente. Distinti tuber- coletti coprono la superficie delle elitre e dell'addome. Edeago figg. 165-166, sesto urotergo libero del maschio fig. 167. COMPARAZIONI. Specie molto affine a S. distincta (Pace, 1982), comb. n., “olim” Anomognathus distinctus Pace, 1982: 89, del Nepal. S. pulcher sp. n. si distingue per avere l’edeago meno profondamente ricurvo al lato ventrale e il pezzo copulatore del sacco interno ricurvo e non rettilineo con un granulo preapicale dorsale come in distincta. I denti laterali del margine posteriore del sesto urotergo libero del maschio sono lunghi e stretti nella nuova specie, di conseguenza l’incavatura contigua è molto profonda, mentre in distincta i denti laterali del corrispondente urotergo, sono corti e a base assai larga, sicché l’incavatura contigua è poco profonda. ALEOCHARINAE DELLA CINA 197 CHAR ET E ALI RITA) rege = 4 per Imm Imm Ficc. 159-168 Habitus, edeago in visione laterale e ventrale, sesto urotergo libero del maschio e spermateca. 159-162: Stenomastax diogenes sp. n.; 163-164: Stenomastax yunnanensis Sp. n.; 165-168: Stenomastax pulcher sp. n. 198 ROBERTO PACE Stenomastax serrula sp. n. Figg. 169-171 Tipo. Holotypus ¢, China, Yunnan, Xishuangbanna, 20.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lungh. 2,0 mm. Avancorpo debolmente lucido, addome lucido. Corpo nero con estremità addominale bruna; antenne nere con i tre antennomeri basali bruni; zampe di un giallo sporco con femori giallo-bruni. La reticolazione del capo è quasi vigorosa, quella del pronoto è vigorosa, quella delle elitre è svanita e quella dell'addome è distinta. La punteggiatura del capo è netta solo sulla metà basale. Il pronoto non mostra punteggiatura, nè tubercoletti. Le elitre sono coperte di tuber- coletti distinti. Edeago figg. 170-171. COMPARAZIONI. La nuova specie presenta l'apice dell’edeago seghettato al lato ventrale. Un simile carattere, ma molto obliterato, si riscontra anche in S. platygaster (Kraatz, 1859), dello Sri Lanka, pertanto l'affinità tra le due specie, anche per altri caratteri affini qui non elencati per ragione di brevità, è pressoché sicura. La sinuosità ventrale dell’edeago della nuova specie non si osserva nell’edeago di platygaster e il tubulo mediano interno dell’edeago è ricurvo e robusto nella nuova specie e rettilineo ed esile in platygaster. Stenomastax contermina sp. n. Figg. 172-173 Tipo. Holotypus 2, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,9 mm. Avancorpo debolmente lucido, addome lucido. Capo bruno, pronoto bruno-rossiccio, elitre giallo-brune, addome giallo-rossiccio con gli uriti liberi terzo, quarto e base del quinto bruni; antenne brune con i tre anten- nomeri basali rossicci; zampe gialle. La reticolazione del capo e del pronoto è netta, quella delle elitre è distinta. La punteggiatura del capo è molto svanita e confusa. Il pronoto e le elitre mostrano tubercoletti poco distinti. Spermateca fig. 173. COMPARAZIONI. Per la presenza di un largo solco mediano del pronoto e per la forma della spermateca, la nuova specie si pone tassonomicamente vicina a S. tuberculicollis (Kraatz, 1859), diffusa dallo Sri Lanka al Borneo e presente anche in Cina. Il bulbo distale della spermateca è subsferico nelle due specie, però nella nuova specie il bulbo prossimale dello stesso organo è nullo, mentre in tuberculicollis è ben conformato e terminante con una corta spira. Stenomastax kadooriorum sp. n. Figg. 174-178 Tıpı. Holotypus é, Hong Kong, Kadoorie Farm, V.1996, de Rougemont leg. (MHNG). Paratypi: | 4, Hong Kong, XII.1995-1.1996, de Rougemont leg.; 1 2, Hong Kong, Tai Po, VII.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 1,8 mm. Avancorpo debolmente lucido, addome lucido. Corpo giallo-rossiccio con capo ed elitre, tranne la base, rossicci, quarto urite libero bruno; antenne brune con i tre antennomeri basali giallo-rossicci e metà apicale dell’undicesimo rossiccia; zampe gialle. La reticolazione del capo è svanita, visibile solo ai lati esterni, quella del pronoto è netta e quella delle elitre e dell’addome 199 ALEOCHARINAE DELLA CINA mm 01 mm 4 Fısc. 169-178 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 169-171: Stenomastax serrula sp. n., 172-173: Stenomastax contermina sp. n.; 174-178: Steno- mastax kadooriorum Sp. n. 200 ROBERTO PACE assente. La punteggiatura del capo è distinta e fitta. Il pronoto presenta tubercoletti confusi nella reticolazione. I tubercoletti che coprono la superficie delle elitre sono svaniti. Edeago figg. 175-176, spermateca fig. 177, sesto urotergo libero del maschio fig. 178. COMPARAZIONI. Per la forma dell’edeago molto simile e per la presenza di 4 denti e 2 spine al margine posteriore del sesto urotergo libero del maschio, la nuova specie si colloca tassonomicamente vicino a S. densissima Cameron, 1941, delle Filippine (di cui è noto solo l’holotypus maschio). Enormi differenze si notano nella forma e dimensione dei pezzi copulatori del sacco interno dell’edeago. S. densissima mostra un robustissimo pezzo copulatore semicircolare nel sacco interno dell’edeago, accompagnato da tre spine diafane, mentre la nuova specie mostra vari pezzi copulatori tra loro separati. I 4 denti al margine posteriore del sesto urotergo libero del maschio della nuova specie sono tra loro ugualmente distanziati, mentre in densissima sono raggruppati due a due e a metà del margine vi è un largo intervallo. Inoltre le spine laterali dello stesso urotergo non superano la lunghezza dei denti nella nuova specie, mentre in densissima li superano di molto. ETIMOLOGIA. Specie dedicata ai fratelli Kadoorie, grandi filantropi di Hong Kong per aver fondato il “Kadoorie Agricultural Research Centre” dell’Università di Hong Kong. Placusa (Calpusa) yunnanicola sp. n. Figg. 179-183 Tipi. Holotypus d, China, Yunnan, Xishuangbanna Jinghong, 11.1993, de Rougemont leg (MHNG). Paratypi: 1 ©, stessa provenienza; 1 d e 2 £ ©, Hong Kong, XII.1995-I.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 1,9 mm. Corpo lucido e bruno; antenne brune con i due antennomeri basali e la base del terzo gialli; zampe gialle. La reticolazione del capo e dell’addome è svanita, quella del pronoto è molto superficiale e quella delle elitre è distinta. La punteggiatura del capo è svanita. Il pronoto e le elitre sono coperti di tubercoletti distinti. Edeago figg. 180-181, spermateca fig. 182, sesto urotergo libero della femmina fig. 183. COMPARAZIONI. Specie, a una superficiale osservazione, apparentemente identica a P. furcifera Pace, 1986 di Bali. E° invece ben distinta per lo stretto apice delle due appendici ventrali dell’edeago, per la spermateca più robusta (con parte prossimale della spermateca nettamente sottile in furcifera) e per i sottili e lunghi denti del mar- gine posteriore del sesto urotergo libero della femmina e del maschio. Placusa (Calpusa) sculpticollis sp. n. Figg. 184-186 Tipo. Holotypus 2, China Yunnan, Xishuangbanna, Sanchahe, elephant res., 24.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,2 mm. Corpo lucido e bruno con margine posteriore degli uroterghi basali giallo-brunni; antenne brune con i due antennomeri basali e la base del terzo gialli; zampe giallo-brune con femori bruno-rossicci. La reticolazione del ALEOCHARINAE DELLA CINA 201 Figc. 179-186 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero della femmina. 179-183: Placusa (Calpusa) yunnanicola sp. n.; 184-186: Placusa (Calpusa) sculpticollis sp. n. 202 ROBERTO PACE capo, delle elitre e dell’addome è distinta, quella del pronoto è svanita. La punteggia- tura del capo è svanita. Tubercoletti distinti stanno sul pronoto e sulle elitre. Sper- mateca fig. 185, sesto urotergo libero della femmina fig. 186. COMPARAZIONI. La nuova specie si distingue da P. yunnanicola sp. n. sopra descritta perché ha pronoto con due deboli fossette longitudinali posteriori, invece di un debole solco mediano posteriore come in yunnanicola. La spermateca della nuova specie è dello stesso tipo di yunnanicola, ma la sua parte prosimale è distintamente più lunga e termina ad angolo, invece di essere arrotondata. Ma è la forma del margine posteriore del sesto urotergo libero della femmina della nuova specie che distingue nettamente i due taxa, figg. 183 e 186. Placusa (s. str.) montium sp. n. Figg. 187-188 Tipo. Holotypus ©, China, Gansu Mts., 25 Km E Xiahe, 2805-2925 m, 3.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 2,6 mm. Corpo debolmente lucido e nero pece; antenne brune con 1 due antennomeri basali bruno-rossicci; zampe bruno-rossicce con femori bruni. Avancorpo coperto di fitti tubercoletti molto salienti. L’addome mostra una fitta pubescenza d’aspetto sericeo. Spermateca fig. 188. COMPARAZIONI. Specie affine a P. acuminata Kraatz, 1859, dello Sri Lanka, distinta per avere il quarto antennomero nettamente trasverso (e non lungo quanto largo come in acuminata), per gli occhi meno ridotti e soprattutto per avere la spermateca più sottile e più lunga, tanto che la sua parte prossimale descrive una spira (spermateca a forma della lettera S rovesciata e robusta, senza spira prossimale in acuminata). Linoglossa (Axinocolya) chinensis sp. n. Figg. 189-190 Tipo. Holotypus 9, China, Sichuan, Gongga Shan, above capo 2, 2800 m, 25.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,7 mm. Corpo lucido e bruno scuro con addome bruno; antenne brune; zampe rossicce. Il capo presenta un profondo solco discale a superficie non reticolata, ai lati la reticolazione è vigorosa; tubercoletti svaniti coprono la super- ficie. Il pronoto è coperto di tubercoli robusti e salienti sulla fascia mediana e radi e fini ai lati; esiste una profonda fossetta posteriore. Le elitre sono coperte di tubercoli allugati molto salienti, soprattutto verso i lati esterni; la loro superficie è priva di reti- colazione, come quella dell’addome. Spermateca fig. 190. COMPARAZIONI. La nuova specie è affine a L. smetanai Pace, 1989, del Nepal, ciò in base all’habitus simile e per la struttura della spermateca. Le differenze più evidenti sono le seguenti: il pronoto della nuova specie non è marcatamente sinuato davanti agli angoli posteriori come in smetanai, inoltre tubercoli grossolani sono addensati sulla fascia longitudinale mediana del pronoto della nuova specie, mentre in smetanai i tubercoli sono uniformemente distribuiti sul pronoto. E’ tuttavia la spermateca che presenta i caratteri differenziali più marcati: il bulbo distale della spermateca della nuova specie è molto più lungo e appena reniforme, mentre in smetanai è nettamente reniforme, con lato destro profondamente angolare verso | interno. ALEOCHARINAE DELLA CINA 203 BOLITOCHARINI Omologlusa gen. n. Figg. 191-195 DIAGNOSI. Genere che mostra alcuni caratteri riscontrabili anche nel genere Neo- leptusa Cameron, 1939, dell’ India, Nepal e delle Filippine. E’ chiaramente distinto per la ligula assai larga e appena incavata al margine anteriore, per il lobo esterno delle mascelle molto corto e, soprattutto, per il tipo di spermateca nuovo per i generi della tribù. La sua forma, se non si osservasse la formula tarsale 4-4-5, indurrebbe a porre senza difficoltà il nuovo genere nella tribù Athetini, in cui tale tipo di spermateca è frequente. DESCRIZIONE. Capo e pronoto piuttosto stretti rispetto alle larghe elitre; collo largo; antenne corte; palpi mascellari di 4 articoli: il quarto è ipertrofico rispetto il secondo che è corto, fig. 195; lobo interno delle mascelle robusto, con una fila di 8 spine mar- ginali interne assai lunghe; lobo esterno molto corto, sicché il ciuffo di setole apicali è molto sviluppato in lunghezza: palpi labiali di 3 articoli: i primi due sono corti e larghi, fig. 192; ligula larghissima e intera, con margine anteriore appena incavato; paraglosse nulle; margine anteriore del mento rettilineo, fig. 193; processo meso- sternale a punta acuta, sicché le mesocoxe sono contigue tra loro; formula tarsale 4-4- 5; pro-mesotibie con due lunghe setole esterne, metatibie lungamente ciliate esterna- mente; primo tarsomero posteriore corto; spermateca trisinuata: è presente l’intro- flessione apicale del bulbo distale, fig. 194. Typus GENERIS: Omologlusa rougemonti sp. n. ETIMOLOGIA. Il nome del nuovo genere significa “Colei che è ammessa concor- demente”. GENERE GRAMMATICALE. Il nuovo genere è femminile. Omologlusa rougemonti sp. n. Figg. 191-195 Tipo. Holotypus ®, China, Beijing, Xiaolongmen, 1100-1500 m, 1.VII.1993, de Rougemont leg. (MHNG). DescRIZIONE. Lunghezza 2,0 mm. Corpo lucido e rossiccio con addome bruno-ros- siccio; antenne brune con i due antennomeri basali e l'apice dell’undicesimo bruno-ros- sicci; zampe gialle. La reticolazione del disco del capo è netta, quella dei lati e della parte posteriore è svanita, quella del pronoto e delle elitre è netta e quella dell’adome è distinta. La punteggiatura del capo è fine e distinta, come quella del pronoto. Tuber- coletti poco distinto stanno sulla superficie delle elitre. Spermateca fig. 194. Phymatura chinensis sp. n. Figg. 196-200 Tipi. Holotypus d, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII.1994, A. Smetana leg. (MHNG). PARATYPI: 2 © ©, stessa provenienza. DESCRIZIONE. Lunghezza 4,0 mm. Corpo debolmente lucido. Capo nero-bruno, pro- noto di un rossiccio sporco, elitre brune con omeri rossicci, addome di un rossiccio ROBERTO PACE 01 mm Fico. 187-193 Habitus, spermateca, labio con palpo labiale e mento. 187-188: Placusa (s. str.) montium sp. n.; 189-190: Linoglossa (Axinocolya) chinensis sp. n.; 191-193: Omologlusa rougemonti gen. n., sp. n. ALEOCHARINAE DELLA CINA PORTE Un PE 1 Vf 199 E = 200 Ficc. 194-200 Spermateca, mascella con palpo mascellare, habitus, edeago in visione laterale e ventrale e sesto urotergo libero del maschio. 194-195: Omologlusa rougemonti gen. n., sp. n.; 196-200: Phymatura chinensis sp. n. 206 ROBERTO PACE sporco con uriti liberi quarto e base del quinto bruni; antenne rossicce con 1 tre anten- nomeri basali giallo-rossicci; zampe rossicce. La reticolazione del capo è fine e svanita, quella del pronoto e delle elitre è distinta e quella dell’addome assente. I tubercoletti che coprono il capo sono salienti, quelli del pronoto e delle elitre sono più fitti di quelli del capo e superficiali. L’addome è coperto di fitta pubescenza. Edeago figg. 197-198, spermateca fig. 199, sesto urotergo libero del maschio fig. 200. COMPARAZIONI. In base alla forma dell’edeago e della spermateca, la nuova specie è vicina tassonomicamente a P. picta Cameron, 1939, dell'India. Se ne distingue age- volmente perché ha taglia corporea maggiore (4,0 mm, invece di 2,7 mm, in base a mia misurazione di un esemplare maschio da me scelto come lectotypus. Cameron dà solo la misura di 3,5 mm). Inoltre l’edeago della nuova specie è poco arcuato al lato ventrale e ha un robusto pezzo copulatore (fortemente arcuato e con sottile pezzo copulatore in picta) e per la spermateca che nella nuova specie non ha bulbo distale così dilatato e con profonda e larga introflessione apicale, quanto quello della spermateca di picta. Phymatura smetanai sp. n. Figg. 201-205 Tipi. Holotypus d, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII.1994, A. Smetana leg. (MHNG). Paratypus: | ©, stessa provenienza. DESCRIZIONE. Lunghezza 3,8 mm. Corpo lucido. Capo nero-bruno, pronoto di un ros- siccio sporco; elitre dello stesso colore, con il quarto posteriore bruno, addome rossiccio con una fascia mediana bruna sugli uroterghi basali e con gli uroterghi liberi quarto e base del quinto pure bruni; antenne rossicce con i tre antennomeri basali giallo-rossicci; zampe rossicce. La reticolazione del capo e del pronoto è distinta, quella delle elitre è svanita e quella dell’addome è molto superficiale. I tuberceletti che coprono il capo sono salienti, quelli del pronoto sono molto salienti e quelli dell’addome sono svaniti. La punteggiatura delle elitre è fitta e distinta. Edeago figg. 202-203, spermateca fig. 204. COMPARAZIONI. Poiché il margine posteriore del sesto urotergo libero del maschio della nuova specie è privo di denti mediani, P. smetanai sp. n. potrebbe essere affine a P. intermedia Cameron, 1939, dell’India. Tuttavia la taglia corporea della nuova specie è maggiore (3,8 mm, invece di 2,7-3,1 mm), gli uriti liberi quinto e sesto sono privi di tubercolo mediano e i dentini del margine posteriore del quinto urotergo libero del maschio, sono meno lunghi e a ciascun lato di essi un dente è minuscolo e non supera in lunghezza i denti contigui (li supera di molto in intermedia). ETIMOLOGIA. Specie dedicata al suo raccoglitore, il Dr. Ales Smetana del “Centre for Land and Biological Resources Research” di Ottawa e insigne stafilinidologo. Phymatura gonggaensis sp. n. Figg. 206-210 Tipi. Holotypus d, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII.1994, A. Smetana leg (MHNG). Paratypi: 1 d e 2 2 9, stessa provenienza. nee ALEOCHARINAE DELLA CINA 207 vali QUARONI ANR vun i SORA i Imm III Um) ] lon 201 205 | vy iyi 210 207 208 Ficc. 201-210 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 201-205: Phymatura smetanai sp. n.; 206-210: Phymatura gonggaensis sp. n. 208 ROBERTO PACE DESCRIZIONE. Lunghezza 3,6 mm. Corpo debolmente lucido. Capo nero-bruno, resto del corpo bruno con margine posteriore del quinto urotergo libero e l’estremità addo- minale rossicci; antenne bruno-rossicce con i quattro antennomeri basali giallo- rossicci; zampe rossicce con femori bruni. La reticolazione del capo è molto svanita, quella del pronoto è superficiale e quella delle elitre è distinta. I tubercoletti della superficie del capo e dell'addome sono distinti, quelli del pronoto sono molto salienti. La punteggiatura delle elitre è svanita. Edeago figg. 207-208, sesto urotergo libero del maschio fig. 209, spermateca fig. 210. COMPARAZIONI. Una specie di Phymatura che presenta una quindicina di corti dentini al margine posteriore del sesto urotergo libero del maschio, non è stata finora ancora osservata. D'altronde anche la spermateca ha una conformazione tale che si discosta dal tipo di spermateca usuale in Phymatura: il bulbo distale assume forma di berretto frigio e il bulbo prossimale è chiaramente ben conformato (poco distinto in altre specie). Pseudatheta cooteri sp. n. Figg. 211-212 Tipo. Holotypus 9, China, Jiangsu Prov., Nanjing Zijinshan, 8.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Corpo lucido e giallo-bruno con pronoto bruno- rossiccio ed estremità addominale rossiccia; antenne nero-brune con i due antenno- meri basali gialli; zampe giallo-rossicce. La reticolazione del capo è svanita, quella del pronoto e delle elitre è distinta e quella dell'addome è molto superficiale. La pun- teggiatura del capo è assai svanita. Il resto del corpo è coperto di tubercoletti distin- tamente salienti. Spermateca fig. 212. COMPARAZIONI. Per la forma simile della spermateca, la nuova specie è forse affine a P. elegans Cameron, 1920, di Singapore e dell'India. Se ne distingue per avere gli occhi più sviluppati, sicché le tempie sono molto più corte degli occhi (e non lunghe quanto gli occhi come in elegans); per il pronoto meno tresverso, con rapporto lar- ghezza/lunghezza pari a 1,27 (1,35 in elegans) e soprattutto per la maggiore dimen- sione della spermateca, con bulbo distale lievemente conico (e non reniforme come in elegans), con microscultura reticolare sulla parete interna della parte mediana e del bulbo prossimale della spermateca (senza microscultura reticolare alla spermateca di elegans). ETIMOLOGIA. Specie dedicata al suo raccoglitore Jonathan Cooter di Hereford (Gran Bretagna), noto studioso di Liodidae. Methistemistiba gen. n. Figg. 213-216 Diagnosi. Corpo simile a Placusa; ma per i palpi labiali di tre articoli si colloca nella tribù Bolitocharini. Tra i generi della tribù Bolitocharini, solo il genere Leptusa Kr. presenta ligula intera, come quella del nuovo genere, tuttavia in Leptusa è molto stretta e molto lunga. Inoltre il tipo di spermateca del nuovo genere non si è mai osservato nel genere Leptusa, a cui perciò non appartiene. ALEOCHARINAE DELLA CINA 209 DESCRIZIONE. Capo e pronoto appena più stretti delle elitre; addome a lati appena divergenti all’indietro; tempie marginate: palpi mascellari di 4 articoli: il secondo è poco più stretto del terzo; lobo esterno delle mascelle lungo, con corte setole apicali; lobo interno delle mascelle stretto, con 5 corte e robuste spine al margine interno; palpi labiali di 3 articoli: il secondo è molto lungo, fig. 215; ligula intera, appuntita e molto larga; paraglosse nulle; mento a margine anteriore pressoché retto, fig. 216; processo mesosternale appuntito; mesocoxe contigue tra loro; formula tarsale 4-4-5; primo tarsomero posteriore corto; tibie con due setole erette esterne; spermateca a forma della lettera Z, robusta e con enorme introflessione apicale del bulbo distale, fig. 214. TYPUS GENERIS. Methistemistiba zhejiangensis sp. n. ETIMOLOGIA. Il nome del nuovo genere significa “Orma che si colloca altrove”. GENERE GRAMMATICALE. Methistemistiba gen. n. è di genere femminile. Methistemistiba zhejiangensis sp. n. Figg. 213-216 Tipo. Holotypus 2, China, Zhejiang Prov., Lin’an County, 350 m, W. Tianmu Shan N.R., 16-22.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 2,0 mm. Corpo lucido e rossiccio con una bruna fascia che non tocca i margini basale e posteriore, sugli uriti liberi terzo, quarto e quinto; antenne e zampe giallo-rossicce. La reticolazione del capo è molto svanita, quella del pronoto e dell'addome è distinta e quella delle elitre è netta. Le maglie di retico- lazione sui quattro uriti basali sono molto trasverse: il quinto urite ha tali maglie trasverse solo ai lati esterni. I tubercoletti della superficie del capo sono quasi indis- tinti, quelli del pronoto e delle elitre sono assai svaniti. Spermateca fig. 214. SAHLBERGIINI Derougemontius gen. n. (vedi Addenda) Figg. 217-224 DIAGNOSI. Per avere il collo stretto, le mandibole senza dente interno, antennomeri compatti e formula tarsale 4-5-5 il nuovo genere può essere incluso nella tribù Sahlbergiini Kistner, 1993, anche se la base dell’addome non è molto ristretta, né le zampe sono molto lunghe. Per questi due ultimi caratteri e per la particolare forma del capo, del pronoto e delle elitre sarebbe necessaria istituire una nuova tribù. Il nuovo genere ha dei caratteri simili a quelli del genere Loeblius Pace, 1985b, come la forma delle antenne e la struttura della spermateca. DESCRIZIONE. Capo con spogenze acute laterali, fig. 218; collo stretto; antenne di 11 antennomeri, fig. 222; tempie non marginate; palpi mascellari di 4 articoli: il secondo e il terzo sono simili tra loro; lobo esterno delle mascelle più lungo dell’interno che mostra lunghe spine al margine interno; palpi labiali di 3 articoli: il primo è più lungo del secondo, fig. 220; ligula larghissima, divisa in due lembi larghi; paraglosse nulle; labbro superiore, fig. 221; prosterno crestato sulla linea mediana anteriore e lieve- mente su quella posteriore, fig. 223; pronoto semicircolare, con una larga impressione 210 ROBERTO PACE mediana e una a ciascun lato; processo mesosternale molto prolungato fra le meso- coxe che sono fra loro lievemente separate; elitre con depressione laterale limitata da uno spigolo saliente; formula tarsale 4, 5, 5; primo tarsomero posteriore appena allungato, fig. 224; spermateca a struttura bisinuata, con distinta introflessione apicale del bulbo distale, fig. 219. TYPUS GENERIS. Derougemontius mirabilis sp. n. ETIMOLOGIA. Genere dedicato al suo raccoglitore Guillaume de Rougemont di Londra, noto studioso di Staphylinidae. GENERE GRAMMATICALE. Derougemontius gen. n. è di genere maschile. Derougemontius mirabilis sp. n. (vedi Addenda) Figg. 217-224 Tipi. Holotypus 9, Hong Kong, Tai Po, flight interception trap, V.1996, de Rougemont leg. (MHNG). Paratypus: 1 9, Hong Kong, Kadoorie, Agricultural Research Centre, flight inter- ception trap, VI.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 2,0 mm. Avancorpo debolmente opaco, addome debolmente lucido. Corpo bruno con pronoto avente le parti concave laterali rossicce, con elitre giallo-brune e con addome giallo-rossiccio; antenne brune con i due antennomeri basali giallo-rossicci e 1 successivi antennomeri fino al sesto rossicci; zampe giallo- rossicce. Il capo è coperto di tubercoletti tra loro contigui che danno un aspetto cesellato alla superficie e presenta una profonda impressione a V sulla fronte. Il pro- noto è coperto di tubercoletti tra loro contigui che danno un aspetto rugoso alla super- ficie e mostra un largo e profondo solco mediano, limitato a ciascun lato da una profunda concavità. Le elitre hanno superficie d’aspetto crivellato dato che la pun- teggiatura è composta di punti tra loro contigui; lateralmente esse mostrano una profonda depressione limitata all’esterno da una saliente carena. Il fondo del solco trasverso basale del primo urotergo libero mostra una fila di setole. Tutto l'addome è coperto di corte e aderenti setoline. Spermateca fig. 219. EUSTENIAMORPHINI Eusteniamorpha zhejiangensis sp. n. Figg. 225-226 Tipi. Holotypus 9, China, Zhejiang, Tianmushan, 2.1X.1994, de Rougemont leg. (MHNG). Paratypus: | ®, stessa provenienza. DESCRIZIONE. Lunghezza 1,9 mm. Avancorpo debolmente lucido, addome lucido. Corpo rossiccio con capo ed elitre bruno-rossicci; antenne rossicce; zampe giallo- rossicce. Il capo e il pronoto sono coperti di tubercoletti molto salienti che danno l’aspetto di una superficie di raspa. Il solco mediano del pronoto è distinto: a ciascun lato di esso verso la base del pronoto una fossetta è profonda. La punteggiatura delle elitre è irregolarmente distribuita ed è composta di punti grandi e di punti fini poco visibili per il fondo nettamente reticolato a maglie ampie. I tubercoletti che coprono gli uriti sono superficiali. Spermateca fig. 226. ALEOCHARINAE DELLA CINA ‘o we 212 01 mm 1mm 214 Fico. 211-217 Habitus, spermateca, labio con palpo labiale e mento. 211-212: Pseudatheta cooteri sp. n.: 213- 216: Methistemistiba zhejiangensis gen. n., sp. n.; 217: Derougemontius mirabilis gen. n. sp. n. (vedi Addenda). 212 ROBERTO PACE 3 005 mm 220 0! mm I \ Olmm A \ \ Lada FIGG. 218-223 Capo in visione ventrale, spermateca, labio con palpo labiale, labbro superiore, antenna e prosterno con protibia. 218-223: Derougemontius mirabilis gen. n. sp. n., (vedi Addenda). 213 ALEOCHARINAE DELLA CINA 01 mm 01mm 226 | FicG. 224-226 Meso-metasterno con meso-metatibia, habitus e spermateca. 224: Derougemontius mirabilis gen. n., Sp. n.; 225-226: Eusteniamorpha zhejiangensis sp. n. 214 ROBERTO PACE COMPARAZIONI. La nuova specie è tassonomicamente affine a E. livida Bernhauer, 1928, delle Filippine, ciò in base alla forma simile della spermateca. Ma la nuova specie ha il bulbo distale della spermateca molto lungo, con microscultura interna evidente e introflessione apicale poco sviluppata, mentre livida ha bulbo distale corto, con poco distinta microscultura interna e con introflessione apicale molto profonda. Inoltre il bulbo prossimale della spermateca è più sviluppato in livida, che nella nuova specie. Anche esternamente si notano differenze: gli occhi della nuova specie sono molto più sviluppati, mentre in livida sono ridotti, sicché le tempie sono molto più lunghe degli occhi. Il pronoto della nuova specie è lievemente trasverso, mentre in livida è lievemente più lungo che largo. Le elitre della nuova specie mostrano pun- teggiatura composta di punti grandi e irregolarmente distribuiti, mentre in livida la superficie delle elitre è coperta da distinti tubercoli. Eusteniamorpha ruiliensis sp. n. Figg. 227-230 Tipi. Holotypus d, China, Yunnan, Ruili, ca. 700 m, 3.1.1993, de Rougemont leg. (MHNG). Paratypi: 5 9 9, stessa provenienza. Descrizione. Lunghezza 2,1 mm. Capo e pronoto molto opachi, resto del corpo lucido. Capo bruno, pronoto bruno-rossiccio con larga fascia rossiccia al margine posteriore, elitre brune con base bruno-rossiccia, addome rossiccio con uriti liberi quarto e quinto bruni; antenne bruno-rossicce con i due antennomeri basali rossicci; zampe rossicce. Il capo è coperto da tubercoletti tra loro contigui che conferiscono alla superficie un aspetto aspro. La medesima superficie sta sul pronoto, ma è estesa solo fino all’altezza della profonda fossetta mediana posteriore: qui la superficie è liscia e lucida, con una fila di punti a ciascun lato. Debole è il solco mediano del pro- noto. La punteggiatura delle elitre è profonda e irregolarmente distribuita, su un fondo non reticolato. L’urotergo basale presenta una fossetta mediana basale attraversata da una carena. Tutti gli uroterghi sono coperti di punteggiatura fine su un fondo non reticolato. Edeago figg. 228-229, spermateca fig. 230. COMPARAZIONI. La nuova specie presenta grande taglia corporea e pronoto fortemente ristretto all'indietro, pertanto mostra affinità con E. monstrosicollis Bernhauer, 1928, delle Filippine, specie nota per il solo holotypus femmina. E. ruiliensis sp. n. si distingue da essa perché non ha un solco mediano e due fossette basali del pronoto come in monstrosicollis, ma un corto e superficiale solco mediano e una fossetta mediana posteriore. La carena mediana del primo urotergo libero della nuova specie, non raggiunge il margine posteriore dell’urotergo stesso, mentre in monstrosicollis lo raggiunge. E’ tuttavia la forma delle spermateca che permette una netta distinzione delle due specie. La nuova specie ha bulbo distale della spermateca molto lungo, mentre in monstrosicollis è cortissimo. Eusteniamorpha chinensis sp. n. Figg. 231-234 Tipi. Holotypus d, China, Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. (MHNG). Paratypi: 2 © 9, stessa provenienza, ma anche in data 4.11.1993. ALEOCHARINAE DELLA CINA 215 227 A FIGG. 227-234 Habitus, edeago in visione laterale e ventrale e spermateca. 227-230: Eusteniamorpha ruiliensis sp. n.; 231-234: Eusteniamorpha chinensis sp. n. 216 ROBERTO PACE DESCRIZIONE. Lunghezza 1,7 mm. Corpo lucido. Capo bruno. pronoto bruno-rossiccio, elitre brune con base bruno-rossiccia, addome rossiccio con uriti liberi terzo, quarto e quinto bruni; antenne rossicce; zampe giallo-rossicce. La punteggiatura del capo è fitta e profonda, assente su una fascia longitudinale mediana. La punteggiatura del pronoto è pure profonda e fitta, ma assente lungo i lati esterni dove la superficie appare lucidissima. Il solco mediano del pronoto posteriormente si divide in due rami lievemente scostati tra loro. A ciascun lato di detto solco sta una profonda fossetta basale. Le elitre sono coperte di robusti tubercoli molto salienti che scompaiono lungo una fascia del margine posteriore. I tubercoletti dell’addome sono svaniti, tranne che sulla metà posteriore del quarto urite libero dove i tubercoli sono salienti e sul quinto dove sono molto salienti, come sul sesto. Edeago figg. 232-233, spermateca fig. 234. COMPARAZIONI. La spermateca della nuova specie ha parte distale che descrive nume- rose spire che si fondono tra loro distalmente. Un tale tipo di spermateca hanno due specie: E. aspera (Fauvel, 1905), di Giava (nota sul solo holotypus femmina) ed £. philippina Bernhauer, 1928, delle Filippine. Tuttavia, mentre la parte distale della spermateca di aspera descrive soltanto 7 spire, di cui le ultime 3 sono tra loro fuse, e la parte prossimale della spermateca di philippina descrive 4 spire libere e un numero imprecisabile di spire fuse tra loro, tanto da formare una parte del bulbo semplice- mente stretto e lungo, la nuova specie ha la parte distale della spermateca che descrive 4 spire libere e circa 10 spire fusa tra loro. Per questi caratteri e per la cortissima appendice ventrale dell’edeago (molto lunga invece in philippina), la nuova specie si distingue nettamente dalle due citate specie e da altre che hanno caratteri differenziali in numero maggiore. ADDENDA Il presente lavoro era già stato consegnato per la pubblicazione, quando verso la fine del 1997 è sato pubblicato il seguente lavoro: KISTNER, D.H., WEISSFLOG, A., ROSCISZEWSKI & MASCHWITZ, U. 1997. New Species, New Genera and New Records of Myrmecophyls Associated with Arma Ants (Aenictus sp.) with the Description of a New Subtribe of Staphylinidae (Coleoptera; Formicidae: Aenictinae). Sociobiology 29: 123-221. In questo lavoro è descritto il nuovo genere Dentaphila Kistner. Esso è lo stesso genere da me descritto nel presente lavoro con il nome di Derougemontius. Pertanto va stabilita la seguente sinonimia: Dentaphila Kistner, 1997: 182 Derougemontius Pace, 1998, hoc opus, syn. n. La nuova specie descritta nel presente lavoro resta invece valida per l'assenza di spine alla base del collo, per il capo meno lungo, per l’assenza di due fossette anteriori del pronoto e per altri caratteri. Essa pertanto va chamata Dentaphila mira- bilis (Pace, hoc opus). ALEOCHARINAE DELLA CINA 217 RINGRAZIAMENTI Rivolgo i miei più sentiti ringraziamenti a quanti mi hanno generosamente affidato in studio le Aleocharinae della Cina oggetto del presente lavoro e frutto di recenti raccolte: il Dr. Ales Smetana di Ottawa, i colleghi Guillaume de Rougemont di Londra, Jonathan Cooter di Hereford (Gran Bretagna), Garry Ades, Graham Reels, il Dr. Jeng-Tze Yang del “National Chung Hsing University” di Taiwan e il Dr. Shugiang Li di Stuttgard (Germania). Per il prestito di tipi e di materiale di confronto, rigrazio molto il Dr. A. F. Newton del “Field Museum of Natural History” di Chicago, il Dr. P. M. Hammond del “Natural History Museum” di Londra, il Dr. L. Baert dell’“Institut Royal des Sciences Naturelles” di Bruxelles, il Dr. H. Schönmann del “Naturhistorisches Museum” di Vienna e il Dr. L. Zerche del D.E.I. di Ebers- walde. BIBLIOGRAFIA BERNHAUER, M. 1907. Zur Staphyliniden-Fauna von Japan. Verhandlungen der zoologisch- botanischen Gesellschaft in Wien 57: 371-414. BERNHAUER, M. 1914. Neue Staphyliniden der Indo-malayischen Fauna. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien 64: 76-109. BERNHAUER, M. 1915. Neue Staphyliniden der indomalayischen Fauna, insbesonders der Sunda-Insel Borneo. Verhandlungen der zoologisch-botanischen Gesellschaft in Wien 65: 134-158. BERNHAUER, M. 1915a. Neue Staphyliniden aus Java und Sumatra. Tijdschrift voor Ento- mologie 58: 213-243. BERNHAUER, M. 1916. Kurzflügel aus dem deutschen Schutzgebiete Kiautschau und Cina. Archiv für Naturgeschichte 81: 27-34. BERNHAUER, M. 1928. Die Staphyliniden der Philippinen. Verhandlungen der zoologisch- botanischen Gesellschaft in Wien 78: 29-44. BERNHAUER, M. 1943. Neuheiten der paläarktischen Staphylinidenfauna. Mitteilungen der Miinchner entomologischen Gesellschaft 33: 169-188. BERNHAUER, M. & O. SCHEERPELTZ. 1926. Staphylinidae 6: 499-988. Pars 82. In: W. Junk (ed.): Coleopterorum Catalogus. Berlin. BLACKBURN, T. 1895. Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia 19: 201-258. BOHEMAN, C.H. 1858. Coleoptera. Species novae discripsit, pp. 1-112. In: Kongliga Svenska Fregatten Eugenies Resa Omkring Jorden unter Bafäl af C.A. Virgin, Aren 1851-1853. Andra Delen, Zoologici 1, Insecta. Stockholm. BRUNDIN, L. 1952. Acrotona-Studien (Gattung Atheta, Col. Styphylinidae). Entomologisk Tidskrift 73: 93-145. CAMERON, M. 1920. New Species of Staphylinidae from Singapore. Transactions of the Ento- mological Society of London 1920: 212-284. CAMERON, M. 1920a. New Species of Staphylinidae from Ceylon. Part II. Entomologist’s Monthly Magazine 56: 49-53. CAMERON, M. 1933. Staphylinidae of Japan. Entomologist’s Monthly Magazine 69: 208-219. CAMERON, M. 1936. New species of Staphylinidae (Col.) from the Malay Peninsula. Journal of the Fed. Malay State Museum 18: 40-53. CAMERON, M. 1939. The Fauna of British India, including Ceylon and Burma. Coeloptera, Staphylinidae. Vol. IV, 410 pp. London. 218 ROBERTO PACE CAMERON, M. 1939a. Fauna Javanica. The Staphylinidae collected by Mr. C.F. Drescher. Tijdschrift voor Entomologie 82: 1-29. COIFFAIT, H. 1984. Staphylinides (Col.) de la Région Himalayenne et de l’Inde. II. Tachy- porinae, Oxytelinae et Aleocharinae. Entomologia Basilensia 9: 116-157. EPPELSHEIM, E. 1883. Neue Staphyliniden der österreichischen-ungarischen Monarchie und der angrenzenden Ländern. Wiener Entomologische Zeitschrift 2: 251-255. ERICHSON, G.F. 1839. Genera et species staphylinorum, insectorum coleopterorum familiae 1, pp. 400. Berlin. ERICHSON, G.F. 1940. Genera et species staphylinorum, insectorum coleopterorum familiae 2: 401-954, Berlin. FAUVEL, A. 1901. Liste des Staphylinidae du Japon central recueillis par M. le Dr. Harmand. Bulletin du Muséum National d'Histoire Naturelle, Paris 1901:62-66. FAUVEL, A. 1905. Staphylinidae exotiques nouveaux (3e partie). Revue d’Entomologie (Caen) 24: 113-147. GRAVENHORST, J. L.C. 1802. Coleoptera microptera Brunsvicensia nec non exoticorum quot- quot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. 206 pp. Brunsvigae. KISENWETTER, H. von. 1844. Die Staphylinidenfauna von Leipzig s Umgegend. Stettiner Ento- mologische Zeitung 5: 307-320; 340-356. KISTNER, D.H. 1985. A new genus and species of termitophilous Aleocharinae from Mainland China associated with Coptotermes formosanus and its zoogeographical significance (Coleoptera: Staphylinidae). Sociobiology 10: 93-104. KISTNER, D.H. 1993. Cladistic analisis, taxonomic restructuring and revision of the Old World genera formely classified as Dorylomimini with comments on their evolution and behavior (Coleoptera: Staphylinidae). Sociobiology 22: 151-383. KLIMASZEWSKI, J. & R.E. JANSEN. 1993. Systemastics, biology and distribution of Aleochara Gravenhorst from Southern Africa. Part I: Subgenus Xenochara Mulsant & Rey (Coleoptera: Staphylinidae). Annals of the Transvaal Museum 36: 53-107. KRAATZ, G. 1857. Genera Aleocharinorum illustrata. Linnaea Entomologica 11: 1-43. KRAATZ, G. 1859. Die Staphyliniden-Fauna von Ostindien, insbesonders der Insel Ceylan. Archiv für Naturgeschichte 25: 1-196. MANNERHEIM, C.G. 1831. Précis d’un nouvel arrangement de la Famille des Brachélytres de l’ordre des Insectes Coléoptères. Mémoires de l’Académie Impériale des Sciences de St- Pétersbourg 1: 415-501. MOTSCHULSKY, V. DE. 1858. Enumération des nouvelles espèces de Coléoptères rapportées de ses voyages. Bulletin de la Société impériale des Naturalistes de Moscou 3: 204-264. MULSANT, M.E. & C. REY. 1874. Tribu des Brévipennes: Familles des Aléochariens (suite): Sixième branche: Aléocharaires. Annales de la Société Linnéen de Lyon 20: 285-447. PACE, R. 1982. Aleocharinae del Nepal e dell’India settentrionale raccolte dal Prof. Herbert Franz I: Bolitocharini. Bollettino della Società entomologica italiana 114: 4-7. PACE, R. 1982a. Ocosomechusa besucheti n. gen., n. sp. dell’India ed Emmelostiba besucheti n. gen., n. sp. del Libano. Revue suisse de Zoologie 89: 443-450. PACE, R. 1984. Aleocharinae delle Mascarene, Parte I et II. Revue suisse de Zoologie 91: 3-36; 249-280. PACE, R. 1984a. Aleocharinae della Thailandia e della Birmania riportate de G. de Rougemont. Bollettino del Museo civico di Storia naturale di Verona 11: 427-468. PACE, R. 1984b. Aleocharinae del’ Asia Sudorientale raccolte dal Dr. Osella. Bollettino del Museo civico di Storia naturale di Verona 11: 481-491. PACE, R. 1985. Aleocharinae del’ Himalaya raccolte da Guillaume de Rougemont. Bollettino del Museo civico di Storia naturale di Verona 12: 165-191. PACE, R. 1985a. Hypocyphtini dell’ India, Ceylon e Nepal del Museo di Ginevra (Coleoptera, Staphylinidae). Revue suisse de Zoologie 92: 77-87. ALEOCHARINAE DELLA CINA 219 PACE, R. 1985b. Uno straordinario mirmecofilo: Loeblius nepalensis gen. et sp. nov. (Cole- optera, Staphylinidae). Revue suisse de Zoologie 92: 291-296. PACE, R. 1986. Aleocharinae dell’ Asia Sudorientale raccolte da G. de Rougemont. Bolletino del Museo civico de Storia naturale di Verona 13: 139-237. PACE, R. 1987. Staphylinidae dell Himalaya Nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). Courier Forschungs-Institut Senckenberg 93: 383-441. PACE, R. 1988 (1987). Aleocharinae dell’ Himalaya raccolte da Marc Tronquet e Georges Ledoux. Bollettino del Museo civico di Storia naturale di Verona 14: 403-419. PACE, R. 1989. Monographia del genere Leptusa Kraatz. Memorie del Museo civico di Storia naturale di Verona Is. A Biologia 8: 1-307 pp. PACE, R. 1989a. Aleocharinae nepalesi del Museo di Ginevra Parte I (Coleoptera, Staphy- linidae). Revue suisse de Zoologie 96: 483-539. PACE, R. 1989b. Aleocharinae nepalesi del Museo di Ginevra Parte II. Revisione del genere Masuria Cameron (Coleoptera, Staphylinidae). Revue suisse de Zoologie 96: 713-727. PACE, R. 1990. Aleocharinae delle Filippine. 82° Contributo alla conoscenza delle Aleo- charinae. /n: Berti, N. (ed.): Miscellanées sur les Staphylins. Mémoires du Museum National d'Histoire naturelle (A) 147: 57-113. PACE, R. 1990a. Aleocharinae del Nepal. 101° Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). In: Berti, N. (ed.): Miscellanées sur les Staphylins. Mémoires du Muséum National d'Histoire Naturelle (A) 147: 155-169. PACE, R. 1991. Aleocharinae nepalesi del Museo di Ginevra Parte IV: Autaliini et Athetini (Coleoptera, Staphylinidae). Revue suisse de Zoologie 98: 107-158. PACE, R. 1992 (1989). Aleocharinae della Thailandia. Bollettino del Museo civico di Storia naturale di Verona 16: 227-268. PACE, R. 1992a. Aleocharinae nepalesi del Museo di Ginevra Parte VII (conclusione): Oxypodini e Aleocharini (Coleoptera, Staphylinidae). Revue suisse de Zoologie 99: 263-342. PACE, R. 1992b. Aleocharinae del Vietnam (Coleoptera, Staphylinidae). Nouvelle Revue d’Entomologie (N.S.) 9: 119-129. PACE, R. 1993 (1990). Aleocharinae della Cina. Bollettino del Museo civico di Storia naturale di Verona 17: 127-180. PACE, R. 1993a (1990). Nuove Aleocharinae Orientali. Bollettino del Museo civico di Storia naturale di Verona 17: 127-180. PACE, R. 1997. Specie del genere Leptusa in Cina. Monografia del genere Leptusa Kraatz: Supplemento VII (Coleoptera, Staphylinidae). Revue suisse de Zoologie 104: 751-760. PAYKULL, G. VON. 1789. Monographia Staphylinorum Sveciae. 82 pp. Upsaliae. REITTER, E. 1887. Insecta in itinere CI. N. Przewalskii in Asia Centrali novissime lecta, Pars 6, Clavicornia, Lamellicornia et Serricornia. Horae Societatis Entomologicae Rossicae (St. Petersburg) 21: 201-234. SAWADA, K. 1974. Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphy- linidae) I: Amidobia. Contributions of the Biological Laboratory of the Kyoto Uni- versity 24: 145-186. SAWADA, K. 1977. Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphy- linidae) III: Japanese Species described by the previous Authors. Contributions of the Biological Laboratory of the Kyoto University 25: 172-222. SAWADA, K. 1978. Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphy- linidae) IV: Three new species from Japan. Contributions of the Biological Laboratory of the Kyoto University 25: 241-268. SCHILLOW, W.F. 1981. Die Lomechusa-Arten der Sowjetunion und angrenzender Gebiete. Reichenbachia 36: 213-223. 220 ROBERTO PACE STEPHENS, J.F. 1832. Illustrations of British Entomology. Mandibulata. Vol. V. 448 pp. + 4 pls. London. THOMSON, C.G. 1856. Nagra nya arter af Insekt-slaegtet Homalota. Ofversigt af Kungliga Svenska Vetenskaps-Akademiens Forhandlingar 1856: 91-107. THOMSON, C.G. 1860. Skandinaviens Coleoptera, synoptiskt bearbetade. II. Tom. 304 pp. Lund. Yosı, R. & K. SAWADA. 1976. Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae) II: Diagnostic characters of Genera and Subgenera with description of representative Species. Contributions of the Biological Laboratory of the Kyoto University 25: 11-140. REVUE SUISSE DE ZOOLOGIE Tome 105 — Fascicule 1 WEBER, Claude. Catalogue révisé des types primaires de la collection ichtyologique du Muséum d’histoire naturelle de la Ville de Genève NEIN Os das are) E cul le open MARTI, Philippe & Jean WUEST. Répartition géographique et morphologie fine de Broelemanneuma gayi (Diplopoda: Craspedosomatidae). . . . ROWELL, C. Hugh F. A revision of the genus Munatia Stàl, 1875 (Ortho- perd CaeliferasRomaleidae Romaleinae) Ei HUBER, Bernhard A. Report on some pholcid spiders collected in Guate- malavand) Honduras (Araneae, Pholcidae). ag 2.2.2008 LANG, Claude. Using a submarine to monitor the biological recovery of deepisedimentsin Eake Geneva (Switzerland) a wm ee FRISCH, Johannes. A revision of some West Palaearctic species of Sco- paeus Erichson (Coleoptera, Staphylinidae, Paederinae)........... ANGELINI, Fernando & Luigi DE MARZO. Supplement to the knowledge of the Agathidiini of Taiwan (Coleoptera, Leiodidae).............. PACE, Roberto. Aleocharinae della Cina: Parte I (Coleoptera, Staphyli- RICA) Es se sd ARR RE TS ET A PAP Re ca Pages 3-14 15-23 25-48 49-80 81-88 89-124 125-138 139-220 REVUE SUISSE DE ZOOLOGIE Volume 105 — Number | WEBER, Claude. Revised types catalogue of the ichthyological collection in the Natural History Museum, City of Geneva (MHNG)............ MARTI, Philippe & Jean WUEST. Geographical distribution and micromor- phology of Broelemanneuma gayi (Diplopoda: Craspedosomatidae). ROWELL, C. Hugh F. A revision of the genus Munatia Stàl, 1875 (Ortho- pleray Caclitera RomalerdaewRomaleinae)a. 52444 62 ren HUBER, Bernhard A. Report on some pholcid spiders collected in Guate- mala anditonduras (Araneae; Pholcidae) EP as Wakes eee LANG, Claude. Using a submarine to monitor the biological recovery of deep sedimentsun Wake'Geneva(Switzerland) OR ET = ee FRISCH, Johannes. A revision of some West Palaearctic species of Sco- paeus Erichson (Coleoptera, Staphylinidae, Paederinae)........... ANGELINI, Fernando & Luigi DE MARZO. Supplement to the knowledge of the Agathidiini of Taiwan (Coleoptera, Leiodidae).............. PACE, Roberto. Aleocharinae from China: Part I (Coleoptera, Staphyli- MICAS) MAMAN EN. RATA URSS PRISER ES LOR Indexed in CURRENT CONTENTS, SCIENCE CITATION INDEX PUBLICATIONS DU MUSÉUM D'HISTOIRE NATURELLE DE GENÈVE CATALOGUE DES INVERTÉBRÉS DE LA SUISSE Fisc S AR CODINES park. PENARD To cee osm ca n ee Fr. 12.— 2 EBREI OPODES pat lh SRINGEEIN gr ne 12.— BEE NIRAIGNEES pa dR EIESSERI ee i ee ee Sa eee 42.— AS SISOR@DES Mat J ZEART OT AR TO I 8.— SMESEUDOSCORPRIONSIpatsRADE ESSERI ar 2 OI 5.50 CAUNEUSOIRESIParESANDREGE SRI AO IAN IT 18.— 7. OLIGOCHETES par E. PIGUET et K. BRETSCHER : . -. ............"...... "tn 18:— Sa CORERODES pam MEI HIEBAUDE rae EEE PRE O ERP De Tee 18.— ISEOENMONSIP ABIREHD ENTFES SERIES Re ECC PEN ER ER ECM EME O RE 11.— OMS CORPIONSIPAMRADENEESSERDO ER II CE RCE CE CENTRO CT ETC ET 3.50 RIÉRSROMATEURS'PAdE = EAWEBEREUGAMONTEN sys PRE EE ti a DE CARODES Dani CAR EU NE oh. n 11.— IBMPACANTHOCERHADES parE SANDRELLI TT TTT TP 11.— PNG ASTÉROMNRICHES par G MONTE © se) OO: I eee ee 18.— ISTENMPENIBODES Dar TR CARE te re RO N IE 12.— 16. 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Les manuscrits sont soumis à des experts d’ institutions suisses ou étrangères selon le sujet étudié. La préférence sera donnée aux travaux concernant les domaines suivants: biogéo- graphie, systématique, écologie, éthologie, morphologie et anatomie comparée, physiologie. Administration MUSÉUM D'HISTOIRE NATURELLE 1211 GENÈVE 6 Internet: http://www.geneva-city.ch:80/musinfo/mhng/publications/revues.htm PRIX DE L'ABONNEMENT: SUISSE Fr. 225.— UNION POSTALE Fr. 230.— (en francs suisses) Les demandes d'abonnement doivent étre adressées à la rédaction de la Revue suisse de Zoologie, Muséum d'histoire naturelle, C.P. 6434, CH-1211 Genève 6, Suisse REVUE SUISSE DE ZOOLOGIE 105 (2): 221-260; juin 1998 A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphylinidae, Aleocharinae) Jan KLIMASZEWSKI! & Stewart B. PECK? 1 BC Research Inc., 3650 Wesbrook Mall, Vancouver, BC Canada V6S 2L2; 2 Carleton University, Department of Biology, 1125 Colonel By Drive, Ottawa, ON Canada K1S 5B6 A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphylinidae, Aleocharinae). - The Galapagos aleocharine rove beetles are reviewed. Fifteen species (3 new), in 9 genera and 7 tribes are recognised in our collections: Athetini: Atheta (Acrotona) pseudoclaudiensis sp. n. (endemic), A. (Atheta) coriaria (Kraatz) (intro- duced, new record), A. lurida (Erichson) (native), A. dichroa (Gravenhorst) (native), A. galapagoensis Pace (endemic); Myllaenini: Myllaena leleupi Pace (endemic); Rothium littoralis sp. n. (probably endemic); Hypo- cyphtini (=Oligotini): Oligota (Holobus) chrysopyga Kraatz (introduced, new record); Homalotini (=Bolitocharini): Diesota (Apheloglossa) fran- ziana (Pace) (status uncertain, possibly endemic), D. (Apheloglossa) leleupi Pace (probably endemic), Phanerota tridentata sp. n. (probably introduced), Thecturota franzi Pace (probably endemic); Falagriini: Myr- mecocephalus concinnus (Erichson) (introduced, new record); Placusini: Euvira scalesia sp. n. (endemic, new record) and Oxypodini: Feluva franzi Pace (probably endemic). The tribes Falagriini, Hypocyphtini and Placu- sini are reported from the islands for the first time. Diesota galapagosensis Pace (1985a), and Rothium ashlocki Ahn & Ashe (1996), previously reported from Galapagos, are not confirmed in our material. All but the unconfirmed species are described/redescribed and their diagnostic features illustrated. Data on bionomics and species affiliations are briefly discussed. Keys for identification are provided. Most naturally occurring species occur in the arid vegetation zone. The 17 known species seem to represent at least 17 separate colonization events. Key-words: Coleoptera - Staphylinidae - Aleocharinae - Galapagos Islands - native species - endemic species - island insects. Manuscript accepted 15.01.1998 222 JAN KLIMASZEWSKI & STEWART B. PECK INTRODUCTION The Galapagos Islands of Ecuador (Map. 1) lie 800-1000 km west of the coast of Ecuador, and span 304 km east to west and 341 km northwest to southeast. There are 45 named islands, islets, or rocks. The total land area is approximately 7856 km. The largest island is Isabela (4670 km?), and the second largest island is Santa Cruz (904 km2). The next five islands in order of size are Ferdinandina (635 km?), Santiago (572 km?), San Cristöbal (552 km?), Floreana (171 km”), and Marchena (130 km?). The islands are oceanic and volcanic in origin. They lie at the edge of the Central Pacific dry zone with two seasons: rainy and warm from about January to May, and dry and cool from about May to December (PECK & KUKALOVA-PECK 1990). Scientific interest in the fauna and flora of the Galapagos Archipelago was first generated after Charles Darwin and H.M.S. Beagle visited the islands in 1835. The islands are now considered as a model system for estimating the dynamics of biotic dispersal to, evolutionary differentiation in, and ecological structuring of oceanic islands (PECK & KUKALOVA-PECK 1990). Darwin collected a total of 29 beetle species, which were described in 1845 by Waterhouse (VAN DyKE, 1953). The general beetle fauna was reviewed by PECK & KUKALOVA-PECK (1990), and the entire insect fauna was summarized by PECK (1996). There are now 418 known beetle species in 238 genera belonging to 59 families of Coleoptera (PECK 1996). The vegetation and moisture conditions in the islands change seasonally and from sea level to the top of the volcanic mountains, two of which reach an elevation of about 1700 m. The following seven biotic zones are reported for the large main islands (PECK & KUKALOVA-PECK 1990, PECK 1991, 1996): littoral zone, arid zone, transition zone, humid forest zone, evergreen shrub zone, fern-sedge or “pampa” zone, and an agricultural zone. The littoral zone: a narrow belt of salt tolerant vege- tation (mangroves, succulents) extending from the beach line to some 10 m or more inland. The arid zone: the largest zone of microphyllous and xerophytic vegetation (Jasminocereus and Optuntia cacti, seasonally deciduous trees of Acacia, Bursera, and Prosopis) extending up to 80-120 m altitude on the southerly face of the islands and up to 200-300 m on the leeward side of the larger islands: The transition zone: a relatively more moist zone consisting of some evergreen plants and numerous tree species, including the genera Pisonia, Pisidium and Piscidia. The humid forest (or Scalesia) zone: a mesophylous, mainly evergreen forest often with a rich under- growth, extending from 180 to 550 m with average rainfall 1040 mm. The forest often contains endemic trees of Scalesia pedunculata, other Scalesia species and Zantho- xylum. The evergreen shrub (or Miconia) zone: a zone above the humid forest, especially on Santa Cruz, Santiago, and San Cristobal, extending from about 450 to 625 m (average rainfall 1694 mm), and often dominated by the endemic Miconia robinsoniana a mesophyllous evergreen shrub often forming a low closed canopy with a dense growth of epiphytic mosses and liverworts. A fern-sedge (or pampa) zone: a zone extending from 550 to 650 m to 1000 m or more on the mountains on higher islands, lacking native woody vegetation, sometimes with abundant Preridium bracken fern. The agricultural zone: a human-influenced zone dominated by intro- GALAPAGOS ALEOCHARINE ROVE BEETLES 223 duced plants in the altered vegetation of former transition, humid forest and evergreen shrub zones. In 1981 COIFFAIT reviewed the rove beetles from the Galapagos, with new collections from N. & L. Leleup. The latest review of the Galapagos aleocharine rove beetles was that of PACE (1985a), based on material collected by Prof. H. Franz in 1975. He recorded 9 aleocharine species, of which 6 were described as new to science, all belonging to four tribes: Bolitocharini (sensu Pace), Callicerini (=Athe- tini), Myllaenini, and Oxypodini. His studies were a well illustrated contribution but were unfortunately based on a small sample size and very restricted collecting area and time period. The results reported here are based on very large sample sizes and extensive collecting over 5 expeditions, temporally extending from early January to July, totalling 15 months of field work. We here present new records for 15 species of aleocharines in 9 genera (Atheta, Diesota, Euvira, Feluva, Myllaena, Oligota, Phane- rota, Rothium, Thecturota) in 7 tribes (Athetini, Falagriini, Homalotini, Hypocyphtini, Myllaenini, Oxypodini, Placusini). Six species in four genera, Atheta, Euvira, Oligota, Phanerota, and three tribes, Falagriini, Hypocyphtini, Placusini, are recorded from Galapagos for the first time (see Table 1). Knowledge of the South American aleocharines, including Colombia, Ecuador, Peru and the West Indies, despite several recent publications (AHN & ASHE 1996, PACE 1982, 1983a, b, c, 1984, 1985a, b, 1986, 1987a, b, c, 1988, 1990a, b) is still fragmentary. This is the major predicament in assessing the origin and phylogenetic affiliations of Galapagos aleocharine beetles, and Galapagos insects in general. However, our preliminary assessment indicates 11 species to be potentially endemic, 2 to be native (to South America), and 4 probably represent very recent accidental introductions by humans (Atheta coriaria, Myrmeco- cephalus cingulatus, Oligota chrysopyga, Phanerota tridentata) (see Table 1 for details). The species of the genera Diesota, Euvira, Feluva, Myllaena, Phanerota, Rothium and Thecturota, are ultimately of South American origin, while Oligota is cosmopolitan and the Atheta are either endemic (A. galapagoensis, A. pseudo- claudiensis), or cosmopolitan (A. coriaria), or of South American origin (A. dichroa, A. lurida). The genus Diesota is particularly species rich on the South American mainland where it has undergone a massive radiation (PACE 1985a, 1986). No endemism at the generic level in Aleocharinae was found, which indicates fairly recent colonizations. As indicated by KUSCHEL (1963) the arthropod fauna of the archipelago is young, but he placed its origin as far back as the Eocene or part of the Oligocene. Present geological understanding places the origin of the present islands to be about 3.5 million years at most (PECK 1996). The Galapagos Islands are famous for Darwin’s Finches, a classic group showing within-archipelagic species formation and adaptive radiation. One might expect other groups of Galapagos organisms to provide similar exemples. Do we see this in the aleocharine staphylinids? Of the 10 genera present, only three have more than one endemic species: Rothium (2 species), Diesota (3 species) and Atheta (2 species). In no case is it clear 294 JAN KLIMASZEWSKI & STEWART B. PECK that these represent sister species decendant from a single common ancestor and originating from Within the Archipelago (only D. galapagoensis has not been seen by us, but the other two Diesota are more closely comparable to different continental species). Thus, the pattern seems to be that each of the 17 known species originated from at least 17 separate colonization events (in the distant past (Pleistocene) for the endemic species, in Holocene times or more recently for the native species, and in Historical times for the introduced species. This lack of within-archipelago species multiplication is similar to the pattern found by BORKENT (1991), in which the 11 known species of Galapagos ceratopogonid flies represent 10 or 11 separate colo- nization events, and is the norm in Galäpago insects (PECK 1996). Table 2 shows the distribution of our samples of aleocharine species in the different vegetation zones. The highest species diversity is equivalent in the arid, transition and humid forest zones with 10 species each, and 9 species in the evergreen shrub zone. Thus, the species diversity is relatively uniform through these 4 vege- tation zones. Excluding introduced species yields a diversity which decreases with increased elevation: arid (9), transition (8), humid forest (7), littoral and evergreen shrub (6), and pampa (4). Six of nine non-introduced species occurring in transition to evergreen shrub zones are able to persist after these are converted to agriculture. MATERIAL AND METHODS Approximately 7321 specimens, collected by S. B. Peck and associates during 5 expeditions to the Galapagos (1985-1996) were examined in this study. The majority of specimens were collected in Flight Interception Traps (FIT), Malaise traps, carrion baited pitfall traps, by sifting litter, and by general collecting. Many specimens were dissected to confirm an identity based on genitalic structures. Genitalia and terminalia were dehydrated in absolute ethanol, washed in xylene, and embedded in Canada balsam on small microslides attached to the pins with the specimens. The habitus illustrations were made using a drawing tube attached to a WILD M3Z stereomicroscope. Genital structures were studied using an image processing system consisting of a compound microscope (JENAMED 2), digital camera (JAVELIN) and an IBM computer with an image processing programe (COMPUTER EYES/1024, Digital Vision). The images were printed in Microsoft Word and used for tracing. All drawings were inked on V-C® tracing film. The terminology used in this paper follows that of ASHE (1984), HOEBEKE (1985), KLIMASZEWSKI (1982a, b), and SEEVERS (1978). The following main taxo- nomic sources were consulted in the course of this study: AHN & ASHE 1996, ASHE (1984), BERNHAUER (1907, 1908, 1941), BERNHAUER & SCHEERPELTZ (1926), COIFFAIT (1981), ERICHSON (1840), FAIRMAIRE & GERMAIN (1861), FAUVEL (1865), HOEBEKE (1985), KLIMASZEWSKI (1982a, b), KUSCHEL (1963), LOHSE et al. (1990), PACE (1982, 1983a, b, c, 1984, 1985a, b, 1986, 1987a, b, c, 1988, 1990a, b), SCHUBERT (1849), SOLIER (1849), and VAN DYKE (1953). GALAPAGOS ALEOCHARINE ROVE BEETLES 225 CONVENTIONS Je Localities and specimen data: Localities are grouped under individual island names, which form the Galäpagos Archipelago (see Map 1). We use the conventional Ecuadorian (not English) names for the indicated islands. Di Specimen Repository abbreviations: DEI Deutsches Entomologisches Institut, Ebersvalde, Germany; (L. Zerche) HFC H. Franz collection, Modling, Austria HUB Museum für Naturkunde, Humboldt Universität, Berlin, Germany; (M. Uhlig) IRSNB Institut Royal des Sciences Naturelles de Belgique, Bruxelles, Belgium; D. Drugmand JKC J. Klimaszewski collection, BC Research Inc., Vancouver, Canada RPC R. Pace collection, Verona, Italy SBPC S. B. Peck collection, Carleton University, Ottawa, Canada SEM Snow Entomological Museum, University of Kansas, Lawrence, U.S.A. Surplus specimens from SBPC will be deposited first in the collections of the Muséum d’Histoire Naturelle, Geneva, Switzerland; and then in Canadian National Collection, Agriculture Canada, Ottawa; Charles Darwin Research Station, Santa Cruz, Galapagos; and the Catholic University, Quito, Ecuador. 3% Citation of information on primary type labels: Text of each label is enclosed within double quotes (“”’), a forward slash (/) with a space on each side separates each line, and information enclosed by brackets ([]) provides further details about the specimen associated with the label. 4, Terms: Frank & McCoy (1990) proposed definitions and a classification of commonly used terms such as “introduced” and “endemic” as applied to species occurring in an area under discussion. They recommend the use of “precinctive” rather than “endemic” when referring to species confined to an area, because of prior and continuing use of “endemic” in a somewhat different sense in epidemiology. However, we prefer to continue to use “endemic” in its traditional biogeographic sense. It is a well under- stood zoological term, is commonly used in entomological literature, and is unlikely to be confused with its use in epidemiology. We also use the term “native” for those species which have probably arrived through natural dispersal (from Mexico or Central and South America) and “introduced” for those accidentally brought in by human activity. We know of no intentional introductions of insects into the Galapagos Islands, for bio-control or other purposes. While our use of terms may not satisfy everybody, it is consistent with the established literature on Galapagos organisms and ideas of their origins and distribution. 226 JAN KLIMASZEWSKI & STEWART B. PECK RESULTS SYSTEMATICS Order Coleoptera Family Staphylinidae Subfamily Aleocharinae KEY TO TRIBES OF ALEOCHARINAE RECORDED FROM GALAPAGOS 7(6). Antenna 10-segmented with distinct 4-segmented club (Fig. 3), tarsal iormula: AA ATX. aisi. 2. } Hypocyphtini (=Oligotini) (Oligota p. Antenna 11-segmented without distinct club, tarsal formula not as above . . . Harsallfonmula 4525; 25.14.75 2 suerte de NE Antennal segments 8-11 bead-shaped, pronotum trapezoidal, widest at apex and narrowest at base (Fig. 2), wings absent, labial palpi extre- melycloneate (Rill) esta re Re Myllaenini (Rothium p. Antennal segments 8-11 never bead-shaped, pronotum differently sha- ped, widest at base or at middle, wings present, labial palpi moderately clongate az ee Guten. oe anale siae ot CRC IEEE RE Head with clearly visible slender neck, pronotum with median sulcus, body ant-like (Fig. 14), legs extremely elongate Re LEE ER Falagriini (Myrmecocephalus p. Head without clearly visible slender neck, pronotum without median sulcus, body not ant-like (Figs 9-13), legs moderately elongate cf MA ei ine bietet en o Athetini (Atheta p. Marsa OMM ASIA RR ca IRR TOS Oxypodini (Feluva p. Marsal formula:4-4-5) 2 #5, ue. Le peo EE Body elongate, fusiform (Fig. 1), with evenly dense, short and adhering pubescence directed posteriorly, abdomen with lateral bristles, labial palpuiextremelyielongate Per re oe Myllaenini (Myllaena p. Body narrowly elongate but not fusiform, pubescence not as above, abdomen without or with less pronounced lateral setae, labial palpi short(exception: Diesota) 13m noie RIE Head approximately quadrate, genae angular posteriorly, neck distinct and narrow, one half as wide as head (Fig. 8) ...... Placusini (Euvira p. Head not as above, neck not visible from above (Figs 4-7) Homalotini (=Bolitocharini) (Diesota, Phanerota, Thecturota p. 230, 231, I. TRIBE Myllaenini Ganglbauer, 1895 229) DPA) 235) 232) (for details on characteristics and phylogenies see AHN & ASHE 1996, KLIMASZEWSKI 1982a, 1992) GALAPAGOS ALEOCHARINE ROVE BEETLES 291 Diagnosis. Tarsal formula 4-4-5 or 4-5-5; body fusiform (Myllaena, Fig. 1) or subparallel (Rothium, Fig. 2); abdomen with protruding prominent setae; labial palpi exceptionally elongate (3-segmented in Rothium, Fig. 18); maxillary palps 4- segmented, last segment minute; tergite 10 deeply bifid (Myllaena) or entire (Rothium). Hydrophilous species with Rothium confined to the seashores of the Pacific coast of Mexico, Ecuador, Peru, and the Galapagos (Ahn & Ashe 1996). Inclusion of Rothium in Myllaenini (Ahn & Ashe 1996) requires redefinition of the tribe. 1. Myllaena leleupi Pace, 1985a (Figs 1, 16) PACE 1985a: 450 (habitus and spermatheca illustrations). Holotype (@): “I. Isabela Sud., a 4 km de la côte, tamisage, XI-1964, N. Leleup leg.” (IRSNB). Diagnosis. Body length 1.8 mm; fusiform (Fig. 1), narrow; uniformly dark brown; slightly glossy; punctation minute and dense; pubescence greyish, very short, dense, and adhering to body; antennal segments 6-10 slightly elongate; head wide at base and produced anteriorly, temples shorter than diameter of eye; pronotum transverse, straight basally and apically, arcuate at sides; elytra transverse; abdomen elongate, gradually tapering posteriorly with some protruding setae (Fig. 1); legs slim. é Unknown. 9 Spermatheca with spherical capsule and short, sinuate duct which is coiled posteriorly (Fig. 16). Tergite and sternite 8 elongate and truncate apically. Bionomics. Our single new female was collected from Buttonwood mangrove (Conocarpus) litter soil washing in the littoral zone. Collecting period: May and November (Pace 1985a). Distribution. Pace (1985a) recorded this species for the first time from Isabela. We confirm his record. Endemic. Material examined (new). One female: Isabela, 2 km W Villamil (SBPC). Remarks. Body form similar to A. cuneata Notman (SE USA), but spermatheca distinct. For details on M. cuneata and other Nearctic species see Klimaszewski (1982a). Rothium Moore & Legner 1977 (Fig. 2) Type species: Rothium sonorensis Moore & Legner We here provide characteristics for this genus because of its unsettled taxonomic position. For discussion see AHN & ASHE (1996). Diagnosis. Tarsal formula 4-4-5, or 4-5-5. Body linear, superficially Leptusa- like (Fig. 2), finely pubescent with macrosetae on pronotum, elytra and abdomen; punctation fine and dense; head with frontal suture; antenna with 3 basal segments strongly elongate, first longest, second and third slightly shorter than first and of equal length, 4-8 elongate and progressively shortening apically, 9-11 bead-shaped; maxillary palpus 4-segmented, basal and apical segments minute, second and third elongate; lacinia elongate, narrow, with teeth in apical fourth; galea narrower than 228 JAN KLIMASZEWSKI & STEWART B. PECK lacinia, and with surface smooth; labial palpi longer than mentum, with a few micro- setae; mentum trapezoidal with anterior angles produced anteriad; labrum transverse, narrow; mesocoxae widely separated, mesocoxal process wide, rounded apically and nearly reaching mesocoxal base; tarsal claws large, sickle-shaped; female tergite and sternite 8 elongate, truncate apically. Members of Rothium are known from the sea- shores of Mexico (Acapulco, Guerrero, Sonora, Sinaloa), Peru (Paita), and Ecuador (Galapagos Islands; continental: Punta Galera, Salinas) (AHN & ASHE 1996). 2. Rothium ashlocki Ahn & Ashe, 1996 Ahn & Ahe, 1996:247 (illustrations of habitus, mouthparts, genitalia). Holotype: Galapagos, Santa Cruz Is., Academy Bay, 15.02.1964, on rocks low tide level, P.D. Ashlock (SEM). Diagnosis (based on original description). Body length 2.4 mm. Linear in shape, antennae, elytra and legs light brown, and head, pronotum and abdomen dark reddish- brown; head almost as long as wide; pronotum about 0.58 times as long as wide; elytral length to pronotum length ratio 1.17. For details and illustrations of genitalia see Ahn & Ashe (1996). See also further under Remarks of À. littoralis. 3. Rothium littoralis sp. n. (Figs 2, 17, 18) Diagnosis. Body length 4.5 mm. Linear in shape (Fig. 2), uniformly reddish- brown, appendages slightly lighter; pubescence short and dense; punctation dense, large and sligthly coarse on abdomen; head triangular, temples rounded posteriorly, as long as diameter of eye, pubescence directed anteriad and obliquely anterolaterad; pronotum overlapping elytra at base, trapezoidal in form, narrowest at base (Fig. 2), pubescence radiating from middle line anteriorly, laterally and posteriorly; elytra short, pubescence directed straight posteriad; abdomen subparallel, tergites 1-5 with shallow basal impressions; legs moderately elongate with strong claws. ¢ Unknown. ? Spermatheca with capsule small, approximately hemispherical, and slightly broader than diameter of duct, duct sinuate and coiled posteriorly (Fig. 17). Tergite and sternite 8 elongate, truncate apically, densely pubescent, pubescence long, protruding setae in 4 rows. Bionomics. The unique female was collected in March from a pitfall trap baited with sea lion dung in the littoral zone. Distribution. Rothium littoralis is known only from Floreana Island, Galapagos. Endemic. Material examined. Holotype (9 ): “ECU.[ador], Galapagos / Floreana, Black Beach [from loberia (Sea lion colony) about 2 km S of Black Beach] / 23-27.111.89, littoral / sea lion dung tp. [=trap] / B.J. Sinclair, 89-159 “ (SBPC). Remarks. The genus Rothium was originally described by MOORE & LEGNER (1977) based on the new species R. sonorensis from Sonora, Mexico. They placed this species in the tribe Bolitocharini (=Homalotini). AHN & ASHE (1996) revised this genus, recognizing 5 species, 3 of which were newly described, and transferred it to GALAPAGOS ALEOCHARINE ROVE BEETLES 229 the tribe Myllaenini. They believe that Rothium is closely related to members of the tribes Myllaenini and Pronomaenini because they have a similar gland opening on tergite 7, antero-lateral angles of mentum produced apically, similarly formed lacinia and galea, ligula short and entire, and elongate labial palpi (for details see AHN & ASHE 1996). Rothium is superficial similar to Polypea Fauvel (Diglottini), known only from Aru Islands, New Guinea (KLIMASZEWSKI 1982b). Both genera share the following characteristics: similar body form, similarly shaped maxillae with 4-segmented maxillary palpus, narrow lacinia and galea, elongate glossae with few microsetae, mentum with two latero-apical projections, and mesocoxae widely separated. Rothium is distinct, however, by having 4-5-5-segmented tarsi, labial palpi longer than mentum, mental apical margin between lateral projections straight, lacinia with apical teeth of a different shape (triangular), galea narrower and glabrous, and female tergite 8 not dentate apically. Rothium littorallis is closely related to R. ashlocki described from Santa Cruz, Galapagos by AHN & ASHE (1996). It is readily distinguishable from R. ashlocki by its twice larger body size, uniform color, more slender lacinia and galea, and the spermatheca with contracted coils of posterior duct (Fig. 17). We think these differences are too great for them to be descended from a common ancestor in the Galapagos. They then represent separate colonizations from the continent. Etymology. The specific name is derived from the adjective littoral, in allusion to the habitat where this species was found and to which it may be restricted. II. TRIBE Hypocyphtini Laporte, 1835 (=Oligotini) LAPORTE 1835:135; NEWTON & THAYER 1992 (nomenclature). Diagnosis. Tarsi 4-4-4-segmented; antenna 10-segmented with distinct club (Fig. 3); hind coxae with lamella over base of femur; body minute. Species of Oligota feed on mites. 4. Oligota (Holobus) chrysopyga Kraatz, 1859 (Figs 3, 19-21) KRAATZ 1859: 25; PACE 1984: 9; FRANK (1972) (illustrations of habitus and median lobe of aedeagus in last two references) Diagnosis. Body length 0.8-1.0 mm; subovate (Fig. 3), widest at elytra; robust: rust-brown or dark brown with light brown / reddish tip of abdomen, legs and base of antennae; moderately glossy; punctation fine; pubescence short and dense on fore- body; scale-like microsculpture present on abdomen (Fig. 3); antenna with two basal segments enlarged, segments 3-6 slightly elongate, 7-10 transverse and forming club; head partially or completely concealed by pronotum, eyes large, pubescence directed anteriad and obliquely antero-laterad; pronotum strongly transverse and strongly convex, hypomera not visible in lateral view, pubescence directed straight and obli- quely posteriad; elytra transverse, pubescence directed straight posteriad; abdomen subparallel basally and tapering apically. 4 Median lobe of aedeagus with small 230 JAN KLIMASZEWSKI & STEWART B. PECK round bulbous (Figs 20, 21) and extremely elongate tubus which is produced ventrally at apex (Fig. 20). Tergite and sternite 8 transverse and truncate apically. 2 Sperma- theca with spherical capsule and narrow, short and approximately L-shaped duct (Fig. 19). Tergite 8 strongly transverse, truncate apically, slightly pointed medially; sternite 8 transverse, and moderately strong pointed apically. Bionomics. Adults have been collected widely from arid zone, thornscrub, transition forest, agricultural zone, guava thicket, Scalesia forest, Miconia forest etc. Some specimens were found in frass under bark, and on Fomes fungi. Collecting methods: Malaise traps, Flight interception Traps, general collecting. Altitudes: 20-570 m above sea level. Collecting period: February to July. Species of this genus are known to feed on mites (Seevers 1978). Distribution. A cosmopolitan species recorded from Africa, Madagascar, Canary Is., Mascarene Is., India, Jamaica, New Caledonia, Sechelles, and Sri Lanka (FRANK 1972, PACE 1984). We report this species for the first time from Galapagos: Isabela, San Cristébal, St. Cruz. Probably a recent accidental introduction by humans. Material examined. (56 specimens, 4 4,7 2, 45 sex undetermined) (SBPC, JKC): Isabela: Tagus Cove (SBPC); 23 km NW Villamil, Jabonocillo forest (SBPC). San Cristobal: 1 km E Progressso (SBPC). St. Cruz: Academy Bay (SBPC); Charles Darwin Research Station (SBPC); Los Gemelos (SBPC, JKC); 2, 4, 21 km N Bellavista (SBPC, JKC); 1 km N Puntudo (SBPC); 7.2 km N St. Rosa (SBPC); Tortoise Reserve (SBPC). III. TRIBE Homalotini Heer, 1839 (=Bolitocharini, Gyrophaenini) HEER 1839: 305; ASHE 1984a, 1991, 1992 (phylogenetic relationships); NEWTON & THAYER 1992 (nomenclature). Diagnosis. Tarsal formula 4-4-5; body elongate, subparallel, or short and compact (morphologically diversified, Figs 4-7); ligula elongate, entire, bifid apically or divided. ASHE (1992) defined this tribe by the following features: presence of more or less developed denticles in the molar region of the ventral (condylar) side of the mandible; narrowing of the distance between the medial setae of the prementum so that the setal insertions are close or contiguous; and narrowing of the medial pseudopores. For details and subtribal classification of Homalotini see ASHE (1992). KEY TO GALAPAGOS SPECIES le Body minutes length 10-13 mm, 2 - Body moderately large; length 1:8-2.5:. mm... 222.222. RSA 3 2(1). Body short and wide, oval in outline, strongly glossy (Fig. 4) ne Bern nant ef see tete o et peat ad Phanerota tridentata sp. n. - Body linear, narrow, moderately glossy (Fig. 5)...... Thecturota franzi Pace 3(1). Antennal segments 5-10 at most twice as wide as long; pubescence shortsibody lengthwles=2-0immi (Fics) Reese rer Diesota franziana (Pace) = Antennal segments 5-10 approximately three times as wide as long; pubescence long; body length 2.1-2.5 mm (Fig. 7) ..... Diesota leleupi (Pace) GALAPAGOS ALEOCHARINE ROVE BEETLES 23] Subtribe Gyrophaenina 5. Phanerota tridentata sp. n. (Fig. 4, 22) Diagnosis. Body length 1.3 mm; shortly-oval, compact and strongly glossy: forebody uniformly dark-brown with abdomen slightly lighter, legs yellowish, antenna brownish apically and yellowish basally; antenna with basal 3 segments elongate, 4 as wide as long, 5-10 transverse,, nearly twice as wide as long; punc- tation and setation throughout body very sparse; head with large eyes, some 12 moderately large and scattered setigerous punctures in each half, frons flattened; pronotum narrowly oval, strongly transverse, setigerous punctures large and scattered, distributed in 4 rows and on disc margin; elytra transverse, with scattered setigerous pores, pores slightly elevated; abdomen sparsely pubescent, basal 3 terga impressed at base, integument with transversely meshed microsculpture. 4 Median lobe of aede- agus with tubus strongly produced ventrally (Fig. 22). Tergite 8 conically produced medially and with two narrow lateral projections. Sternite 8 slightly transverse and rounded apically. 2 Unknown. Bionomics. Collected in May from guava thicket in agricultural zone using Flight Interception Trap. Altitude: 360 m above sea level. For life history and habits of some species see ASHE (1981, 1982). Distribution. Known only from St. Cruz. Probably introduced, because it 1s known only from the disturbed agricultural area. Material examined. Holotype (3): “ECU.[ador], St. Cruz / 2 km N Bellavista / 14, V.85, 85- / 159, 360 m” “S. & J. Peck, guava /thicket, FIT, Agric. / area” (SBPC). Remarks. For details on the genus see ASHE (1984b, 1986). Ph. tridentata may be readily separated by the distinctive shape of male tergite 8 with three apical pro- jections. No close relative was established. Ph. brunnessa Ashe from Florida also bears three (but smaller) projections on male tergite 8 (see ASHE 1986). PACE (1987a, b, 1990a) reviewed and illustrated some Latin American Gyrophaenina. We think the species belongs to Phanerata, and not Gyrophaena, because the latter does not have a stongly bent median lobe of the aedeagus in such a manner, and male sternite 8 has 3 strong projections, which are not typical in Gyrophaena (see ASHE 1986). Etymology. The specific name tridentata, having three teeth, relates to the three dentate (toothed) apex of male tergite 8 of this species. Subtribe Homalotina 6. Thecturota franzi Pace, 1985a (Figs 5, 23-25) PACE 1985a: 450 (habitus, spermatheca and median lobe of aedeagus illustrations). Holotype (d ): “Santiago Gebirge [mountains], V- VI. 1975, H. Franz leg. “(HFC). Diagnosis. Body length 1.0-1.3 mm; narrowly subparallel (Fig. 5); uniformly rust-brown, dark-brown or nearly black, occasionally with paler legs and basal antennal segments; moderately glossy; punctation fine, slightly asperate particularly on pronotum, dense on forebody and sparse on abdomen; antennal segments 4-10 232 JAN KLIMASZEWSKI & STEWART B. PECK strongly transverse, up to 3 times as wide as long (Fig. 5); head slightly wider than pronotum, with gena long (longer than diameter of eye), subparallel and rounded posteriorly, dorsum flattened, pubescence directed laterad from midline of disc (Fig. 5); pronotum narrowly trapezoidal dorsally, pubescence directed horizontally laterad from median line (Fig. 5); elytra elongate with pubescence directed obliquely latero-posteriad (Fig. 5); abdomen widening apically at 5/6 of its length and narrowed at apex, 4 basal tergites impressed basally (Fig. 5); legs short. d Median lobe of aedeagus with moderately large ovoid bulbous and narrowly elongate tubus which has sinuate venter in lateral view (Figs 23, 24). Tergite 8 transverse, truncate apically; sternite 8 transverse, rounded apically and slightly pointing. 2 Spermatheca with capsule narrowly spherical, duct short and curved posteriorly (Fig. 25). Tergite 8 transverse, truncate apically; sternite 8 transverse, broadly rounded apically. Bionomics. Adults have been collected from: arid zone, Bursera forest, transition zone forest (of Bursera, Trema, Zanthoxylon), agriculture zone, rose-apple thicket, in forest litter (Tournefortia litter), guave/fern litter, litter in grietas (lava cracks), epiphyte and rotten wood litter, frass under bark, cow, horse and tortoise dung, Scalesia and Miconia forest, and pampas. Altitudes: 10-700 m. Collecting methods: sifting litter and dung, carrion traps, Flight Interception Traps, and UV light traps. Distribution. PACE (1985a) recorded this species from Santiago, and St. Cruz. Both records are confirmed here. New island records: Fernandina, Floreana, Isabela, Marchena, Pinta, Pinzon, and San Cristobal. Material examined. (282 specimens, d, 2, sex undetermined) (SBPC, JKC): Fernandina: 5, 10 km NE Cabo Hammond (SBPC, JKK). Floreana: 6 km E Black Beach (SBPC); Cerro Pajas (SBPC). Isabela: Alcedo (east crater) (SBPC); Cerro Azul (SBPC, JKC); Sierra Negra (SBPC, JKC); 2 km NE Tagus Cove (SBPC, JKC); 13 km NW Villamil, Jabonicillo forest (SBPC). Marchena: Pta. Espejo (SBPC); SW Playa (SBPC). Pinta: trans. zone forest (SBPC, JKC). Pinzön: SE slope (SBPC). San Cristöbal: El Junco (base & rim) (SBPC); 1 km E Progresso (SBPC, JKC); 3-5 km E Wreck Bay (SBPC). Santiago: Aguacate Camp and vicinity (SBPC, JKC); Santiago Camp (SBPC); 8 km SE Playa Espumilla (SBPC). St. Cruz: 2 km N Bellavista (SBPC); El Granillo (SBPC); 1 km NE Media Luna (SBPC, JKC); Puntudo (SBPC, JKC); 7 km N Puerto Ayora (SBPC, JKC); 7, 7.2, 13 km N St. Rosa (SBPC, JKC); Tortoise Reserve (SBPC). Subtribe Silusina 7. Diesota (Apheloglossa) franziana (Pace, 1985a) (Figs 6, 27-30) Parasilusa franzi PACE 1985a: 451 (illustrations of habitus and median lobe of aedeagus). Diesota (Apheloglossa) franziana (PACE 1986: 422). Holotype (9): “S. ta [=Sta.] Cruz, Wald uber [forest above] S. ta [=Sta.] Rosa, V-VI.1975, H. Franz “(HFC). Diagnosis. Body length 1.8-2.0 mm; narrowly subparallel; flattened (Fig. 6); color variable, either approximately uniformly dark brown to black, sometimes with reddish-brown legs and abdomen, or reddish brown with dark brown head, or reddish- brown with dark brown head, pronotum, elytra, and posterior abdomen; antenna uniformly rust-brown, black, or rust-brown basally and black apically; moderately GALAPAGOS ALEOCHARINE ROVE BEETLES 233 glossy, less so on forebody; punctation fine, dense and asperate on forebody; microsculpture present; pubescence short; antennal segments 5-10 transverse, at most about twice as wide as long (Fig. 6); head as wide as long, aproximately subequal in width to pronotum, abruptly produced in front of eyes, rounded posteriorly, temples shorter than diameter of eye, pubescence directed anteriad medially and laterad on either side from midline (Fig. 6); pronotum transverse, rectangular in shape, and with small mediobasal impression, pubescence directed laterad and obliquely laterolaterad (Fig. 6); elytra as long as wide or slightly transverse, insignificantly wider than pronotum and abdomen, pubescence directed approximately straight posteriad (Fig. 6); abdomen subparallel, 4 basal tergites shallowly impressed basally, pu- bescence directed straight posteriad (Fig. 6). 4 Median lobe of aedeagus with ovoid bulbous and moderately narrow, tapering apically tubus (Fig. 30), venter of tubus approximately straight laterally (Fig. 29), internal sac with inconspicuous structures. Tergite 8 transverse with 6 apical teeth, 2 lateral ones being longer (Fig. 28). Sternite 8 transverse and truncate apically with numerous apical bristles. 2 (new record). Spermatheca with long (variable in length) frequently and regularly coiled duct and narrow, elongate capsule (Fig. 27). Tergite and sternite 8 transverse, and truncate apically. Bionomics. Adults have been collected from: arid zone, thornscrub, Opuntia forest, transition zone, agricultural zone, avocado grove, pastures, Bursera, Scalesia, Miconia forest, in leaf litter, fruit litter, soil and rotten cactus, and in carrion. Collecting methods: sifting, Malaise traps, Flight Interception Traps (FIT), carrion and banana baited traps. Altitudes: 10-620 m above sea level. Collecting period: February to August. Distribution. PACE (1985a) recorded this species from St. Cruz (Sta Rosa), and we confirm his record here. New island records: Floreana, Isabela, Marchena, Pinzon, Santiago, San Cristébal. Status uncertain (possibly endemic). Material examined. (153 specimens, 18 6, 18 ©, 117 sex undetermined) (SBPC, JKC): Floreana: 6 km E Black Beach (SBPC). Isabela: Sto. Tomas; 1/2 and 3 km W Sto. Tomas; Corazon Verde; 13 km NW Villamil (SBPC, JKC). Marchena: SW Playa (SBPC). Pinzon: SE slope (SBPC). Santiago: Aguacate Camp (SBPC, JKC). San Cristébal: 5 km E Wreck Bay (SBPC). St. Cruz: Academy Bay (SBPC); Bellavista and vicinity (SBPC, JKC); Charles Darwin Research Station (SBPC); Horneman Farm (SBPC, JKC); Los Gemelos; 7.2, 15 and 31 km N Sta Rosa (SBPC); Media Luna (SBPC); Puerto Ayora and vicinity (SBPC). Remarks. PACE (1985a) affiliated D. franziana with three Latin American species described by ERICHSON (1840): D. laesicollis (Er.), from Brazil; D. flavipennis (Er.), from the Antilles; and D. melanura (Er.), from Puerto Rico. We have examined the original type material of the latter three species and found consistent differences in body size, color, proportion of pronotum and genitalia between D. franziana and D. flavipennis and D. melanura. However, we were not able to find important differences in external morphology, color and the shape of spermathecae between D. franziana (reddish-brown form) and D. laesicollis. Should further studies including comparison of males of the two species (D. laesicollis male unknown) prove that there are no substantial differences, the status of D. franziana should be reconsidered. Dark brown 234 JAN KLIMASZEWSKI & STEWART B. PECK or black specimens in our material with slightly narrower pronota than those of the reddish-brown form are considered to be color variations of D. franziana because they retain the same genitalic structures. However, they may represent sibling species which can not be discriminated using morphological methods. 8. Diesota (Apheloglossa) leleupi (Pace) (Figs 7, 31-34) Parasilusa leleupi PACE 1985a, PACE 1986 (as Diesota Subg. Apheloglossa). Holotype (9 ): I. Isabela Sud, à 4 km de la côte, tamisage, XI-1964, récolté en forêt humide, N. Leleup leg. (IRSNB). Diagnosis. Readily distinguishable from D. franziana and the mainland Diesota species by the extremely transverse antennal segments 7-10 (approximately 3 times as wide as long, Fig. 7). Body length 2.1-2.5 mm; narrowly oval, abdomen subparallel (Fig. 7); uniformly reddish-brown (all but one) or dark brown (one specimen), with darker, brown to greybrown, apical portions of antennae (segments 5-11); strongly glossy; punctation fine and sparse on forebody and large and coarse on abdomen, especially in tergal impressions; pubescence moderately long to long on abdomen; microsculpture not apparent; head nearly as wide as pronotum, abruptly produced in front of eyes, widely rounded behind, temples shorter than diameter of eye, pu- bescence directed inward medially and lateroanteriad elsewhere (Fig. 7); pronotum strongly transverse, slightly narrower than elytra, approximately rectangular in shape, with U-shaped basal impression, pubescence directed anteriad on middle line of disc and laterally on sides (Fig. 7); elytra slightly transverse, pubescence directed straight- and slightly obliquely posteriad (Fig. 7); abdomen with 4 basal tergites impressed basally, impressions with large and coarse punctures, punctures outside impressions asperate (Fig. 7). d (new record). Median lobe of aedeagus with bulbous oval in outline and tubus wide at base and gradually narrowed apically (Fig. 33), laterally tubus slightly sinuate subapically and produced ventrally (Fig. 32). Tergite 8 trans- verse, with 6 apical narrowly elongate teeth of approximately equal length. Sternite 8 transverse, and widely rounded apically. 2 Spermatheca with extremely long regu- larly and irregularly twisted duct, capsule narrow and elongate (Fig. 31). Tergite 8 strongly transverse, truncate apically. Sternite 8 slightly transverse, widely rounded apically with slightly emarginate apex. Bionomics. Adults have been collected from: littoral zone, under Sesuvium and Heliotropium debris on upper beach, cormorant nest debis, and from moist litter in rock crevices. Altitudes: 1-2 m above sea level. Collecting methods: sifting debris, yellow pan trap, Flight Interception Trap (FIT). Apparently mostly a littoral zone species. Although the Leleup holotype record is reported to be from “humid forest”, the actual locality is in the arid zone in a low spot where groundwater, under tidal influence, presents a local zone of moisture. Distribution. Recorded by PACE (1985a) from Isabela I. New island record: St. Cruz. Endemic. Material examined. (23 specimens, 5 4,5 2, 13 sex undetermined) (SBPC, JKC). St. Cruz: Charles Darwin Research Station, 29.1.89, 89-3, S. Peck (SBPC, JKC); Charles GALAPAGOS ALEOCHARINE ROVE BEETLES 235 Darwin Research Station, littoral zone, 19.1-9.11.89, yellow pan trap; B. J. Sinclair (SBPC); Tortuga Bay, 18.V.85, 85-167, S. & J. Peck (SBPC), labelled as previous one except: 23.V.85, back beach Heliotropium currassavicum lit. [ter]? (SBPC). Isabela: Bahia Urvina, 23.V.92, S. Peck 92-200 (SBPC, JKC); Villamil 1 km W, 2-15.111.89, 89-88, Peck & Sinclair (SBPC). Remarks. This species resemblances D. patagonica (Scheerpeltz) but has more strongly transverse antennal segments 7-10, and has a differently shaped spermatheca. It is readily distinguishable from all South American species known to us by having antennal segments 7-10 extremely transverse, approximately three times wider than long. For illustrations of some South American Diesota see PACE (1985b, 1986, 1987a, b, 1990a). 9. Diesota galapagosensis (Pace), 1985a Parasilusa galapagosensis PACE 1985a: 452; PACE 1986: 422 (as Diesota). Holotype (9): Isla Marchena, S. Seite, 5/6. 1975, H. Franz leg. (HFC). Diagnosis (based on the original description). Body length 2 mm, narrowly oval, abdomen subparallel; head brown, pronotum and abdomen reddish except for brown abdominal segments 4 and 5, antennae with two basal segments reddish and remain- der brown. Habitus and male genitalia illustrated by PACE (1985a). Remarks. Unfortunately we were not able to borrow the holotype specimen for study. IV. TRIBE Placusini Mulsant & Rey, 1871 MULSANT & REY 1871: 102; ASHE 1989, 1991, 1992 (definition, phylogenetic relationships): NEWTON & THEAEYER 1992 (nomenclature) Diagnosis. Tarsal formula 4-4-5; head with strongly angular posterior genae and with distinct narrow neck (Euvira only, Fig. 8); labium rounded medially with small a-sensillum, epipharynx with longitudinal medial field of small pores flanked on either side with row of large scales, mandibles with dorsal ‘velvety patch’ modi- fied to transverse rows of large teeth, labium with short and widely rounded ligula (ASHE 1991). For larval characteristics see ASHE (1991). 10. Euvira scalesiai sp. n. (Figs 8, 26) Diagnosis. Body length 1.7-1.9 mm; uniformly dark-brown with reddish-brown antennae (except apex), maxillary palpi and legs; glossy; punctation sparse and coarse; integument with reticulate microsculpture present; pubescence sparse, moderately long; antennal segments 5-10 strongly transverse, at least twice as wide as long (Fig. 8); head with long subparallel temples, angular posteriorly, with distinct neck approximately half as wide as base of head, pubescence directed medially (Fig. 8); pronotum transverse, about 1.4 times as wide as long, sides widely rounded, base sinuate laterally, posterior angles distinct and slightly pointed posteriad, pubes- cence directed approximately straight posteriorly (Fig. 8); elytra elongate, pubescence directed straight posteriorly (Fig. 8); abdomen subparallel, widely rounded apically, 4 basal tergites deeply impressed basally (Fig. 8). d Unknown. 9 Spermatheca with 236 JAN KLIMASZEWSKI & STEWART B. PECK capsule elongate and globular, as in Fig. 26. Tergite 8 strongly transverse, truncate apically; sternite 8 approximately as wide as long, apical margin arcuate. Bionomics. Adults (all females) have been collected only from forests of Scalesia pedunculata. Collecting methods: vegetation sweeping and Malaise traps. Altitudes: 550-620 m above sea level. Collecting period: April, May. Most specimens were taken by sweeping low grass-herbaceous vegetation in the humid forest highland zone. Ashe & Kistner (1989), recorded larvae and adults of Euvira diazbatresae Ashe from the nests of communal pierid butterfly Eucheria socialis in Mexico. It remains to be seen if other species of the genus exhibit similar behaviour. Distribution. Known only from St. Cruz Island, Galapagos. Endemic. Material examined. (12 9) (SBPC, JKC): Holotype (9 ): “Galap.[agos], Santa Cruz / Los Gemelos, 620 m / 17.V.91, J. Heraty / Scalesia zone H91/38” (SBPC); Paratypes (9 9): 3 labelled as holotype (SBPC); “Galap. [agos], Santa Cruz / 3 km N Santa Rosa, 600 m / Scalesia H91/011” (SBPC) 1; “Galap., Santa Cruz, Los Gemelos, 620 m / 1.V.91, J. Heraty / Scalesia H91/012” (SBPC, JKC) 4; “Galap. St. Cruz / 1.7 km N Sta. Rosa, 1-30.V1.91, 550 m /Scalesia forest malaise / S. Peck, 91-233” (SBPC) 2; St. Cruz, 7.2 km N Sta. Rosa (SBPC) 1. Etymology. Name of this species derived from the generic name Scalesia pedunculata, a tree common to the humid forest zone of some islands of the Galapagos. Remarks. We were able to examine the type series (7 specimens, HUB) of Euvira fervidula Erichson from Columbia. The specimens are generally similar to our spe- cimens of E. scalesia in body form and size but have the forebody orange instead of uniformly brown and have a slightly differently shaped spermatheca. V. TRIBE Athetini Casey, 1910 (=Callicerini) Diagnosis. Tarsi 4-5-5-segmented, mesocoxae usually narrowly separated, intercoxal process highly variable, male tergite eight usually modified and bearing teeth (Figs 51, 53, 54), median lobe of aedeagus with an oval compressor plate and in front with a transverse sclerotised strip called the ‘athetine bridge’ (SEEVERS 1978). Maxillary palpi 4-segmented, labial palpi usually 3-segmented. KEY TO SPECIES I. Body fusiform (Figs 9, 12), length 1.0-3.0 mm (average 2.0 mm); pronotum slightly transverse, approximately trapezoidal in outline with lateral margins strongly converging apically; convex; pubescence directed straight or antero-laterad, laterad or obliquely posteriad, forming straight lines radiating from midline of disc (Figs 9, 12).......... 2 - Body linear (Figs 10, 11, 13), length 2.1-3.1 mm (average 2.6 mm); pronotum strongly transverse, approximately rectangular in outline with lateral margins slightly converging apically: flattened; pubescence directed anteriad, laterad or lateroposteriad, forming arched lines from midline of disc,(Figs 10) 11; 131. I Hr Er RT Re 3 GALAPAGOS ALEOCHARINE ROVE BEETLES 237 2(1). Body length 1.3-2.0 mm; moderately glossy; pronotal punctation asperate; pronotal pubescence directed anterolaterad, laterad and pos- teriad from midline of disc (Fig. 12); antennal segments 8-10 appro- ximately twice as wide as long (Fig. 12); median lobe of aedeagus and spermatheca as in Figs 41, 42, 44. . Atheta (Acrotona) pseudoclaudiensis sp. n. - Body length 1.8-3.0 mm; strongly glossy; pronotal punctation not asperate; pronotal pubescence directed straight and obliquely posteriad (Fig. 9); antennal segments 8-10 at most 1.5 times as wide as long (Fig. 9); median lobe of aedeagus and spermatheca as in Figs 35, 36, 43 So dhs “oro RT SATA URI at Atheta galapagoensis Pace 3(1). Antennal segments 7-10 strongly transverse, about twice as wide as long (Fig. 11); body strongly glossy; pronotum strongly transverse, nearly twice as wide as long (Fig. 11); median lobe of aedeagus and spermathecalas nEies 4749 nern. Atheta (s. str.) coriaria (Kraatz) - Antennal segments 7-10 moderately transverse (Figs 10, 13), about 1.5 times as wide as long; body moderately glossy; pronotum moderately transverse, at most 1.4 as wide as long; genitalia different ............... 4 4(3). Eyes large, diameter much longer than temples (Fig. 13); temples short and abruptly converging posteriorly (Fig. 13); median lobe of aedeagus and spermatheca distinct (Figs 39, 40, 45) ..... Atheta dichroa (Gravenhorst) - Eyes moderate in size, diameter shorter or at most as long as temples (Fig. 10); temples long, subparallel anteriorly (near eyes), and gradually converging posteriorly (Fig. 10); median lobe of aedeagus and sper- mathecasasyineEies327,38,.40. 2. 2.20 m es Atheta lurida (Erichson) 11. Atheta galapagoensis Pace, 1985a (Figs 9, 35, 36, 43, 52) PACE 1985a: 452 (habitus, aedeagus and spermatheca illustrations). Holotype (4): “Isabela I., Cerro Azul 5/6-1975, H. Franz leg.” (HFC). Diagnosis. Body length 1.8-3.0 mm; narrowly oval (Fig. 9), from uniformly dark brown or nearly black to light brown with paler pronotum, elytra, abdominal apex and appendages; strongly glossy; punctation moderately dense on pronotum and elytra and sparser elsewhere; pubescence short, moderately dense on forebody, and sparse on abdomen; meshed microsculpture present on forebody; antennal segments 5-10 slightly transverse, each segment at most 1.5 wider than long (Fig. 9); head approximately quadrate, temples widely round, eyes large, longer than temples (Fig. 9); pronotum much wider then head, transverse, strongly convex, pubescence straight and directed slightly obliquely posteriad (Fig. 9); elytra transverse with pubescence directed posteriad (Fig. 9); abdomen widely arcuate laterally and gradually narrowing apically, four basal tergites with deep basal impressions (Fig. 9). & Median lobe of aedeagus with moderately large, ovoid bulbous and narrowly elongate tubus (Figs 35, 36), internal sac with two subapical structures (Fig. 36). Tergite $ truncate apically with arcuate apex, apical margin entire (Fig. 52), sternite 8 truncate and slightly 238 JAN KLIMASZEWSKI & STEWART B. PECK pointed apically. 2 Spermatheca with enlarged, elongate capsule, and sinuate pos- terior duct (Fig. 43). Tergite 8 truncate apically, sternite 8 strongly pointed medially. Bionomics. Adults have been collected from littoral zone, arid zone, transition zone and pampa zone, leaf litter, deep soil litter, soil washing under Croton and Sesuvium (littoral zone), shrub litter, lava flow edge. Cryptocarpus and Manchineel litter, cow and horse dung, sea lion dung, epiphytes, dead wood (e.g., rotted avocado), frass under bark, under bark of dead Manchineel, rotted logs, guava/mosse litter, lake edge litter, mangrove litter, Trema/Zanthoxylum litter, Zanthoxylum/lichen litter, moss forest litter, fern/moss and fern/sedge litter, treefern litter, rotting Opuntia trunks, cave litter, in moss forest, Miconia and Scalesia forests, grass and Bursera forest, Pisonia forest, Zanthosylum/Pisonia forest, coffee plantation. Collecting methods: Malaise traps, Flight Interception Traps (FIT), general sweeping, sifting leaf litter, dung baited traps, deep soil traps, soil washing, and UV traps. Altitudes: from 1 m to 1700 m above sea level. Collecting period: January to July. Distribution. Originally recorded by Pace (1985) from Isabela, Pinta, Pinzon, Santiago, San Cristobal, and St. Cruz Islands. Our data confirm all of Pace’s records. New island records: Bartolomé, Espanola, Fernandina, Floreana, Marchena, Plaza Sur, Rabida, and St. Fe. Material examined. (3523 specimens: 14 4,3 9, 3506 sex undetermined) (SBPC, JKC). Bartolomé: (SBPC). Española: Bahia Manzanillo (SBPC). Gardner at Española: arid zone (SBPC). Fernandina: Crater Rim (SBPC); 8, 10, 11 km NE Cabo Hammond (SBPC, JKC). Floreana: 3, 5, 6, 8 km E Black Beach (SBPC); Pta. Cormorant (SBPC); Cerro Pajas (SBPC); Finca Cruz (SBPC). Isabela: 7-10 km NE P. Bravo (SBPC); Cerro Azul and vicinity, 7 km NE caleta Iguana (SBPC); 11 km SW Playa (SBPC); Sierra Negra (SBPC, JKC); 4, 8 km NW Sto. Tomas (SBPC, JKC); Tagus Cove and vicinity (SBPC, JKC); 4,12 and 2 km NW Villamil (SBPC); Volcan Alcedo and vicinity (SBPC, JKC). Marchena: Pta. Espejo (SBPC); SW Playa (SBPC); Pinta: Playa Ibbetson (SBPC, JKC); transition zone forest (SBPC); Zanthoxylum-lichen forest (SPPC). Pinzon: SE slope (SBPC). Plaza Sur: S Plazas (SBPC). Rabida: Red Beach (SBPC). Santiago: Central Camp (SBPC); Aguacate Camp and vicinity (SBPC); 9 km SE Playa Espumilla (SBPC); Playa Espumilla (SBPC); 1 km E Progresso (SBPC). San Cristobal: Baquerizo and vicinity (SBPC); El Junco and vicinity (SBPC); Poza Colorada (SBPC); 2-12 km SE Wreck Bay (SBPC). St. Fé: litoral zone (SBPC, JKC). St. Cruz: 2, 3 km W Bellavista (SBPC); Cerro Crocker and vicinity (SBPC); 2 km E Charles Darwin Research Station (SBPC, JKC); Cueva Tres Entradas (SBPC); Los Gemelos and vicinity (SBPC, JKC); Media Luna and vicinity (SBPC); Pta. Roca fuerte (SBPC); Pto. Ayora and vicinity, Tortuga Bay (SBPC); Puntudo (SBPC, JKC); 1.7, 3, 7.2, 10, 13, 15 km N Sta. Rosa (SBPGE, JKC). Remarks. According to PACE (1985a) A. galapagoensis belongs to the species group of A. propinqua (Erichson, 1840) which is a distinct species and should not be considered as a synonym of A. dichroa (Gravenhorst, 1802). A. propinqua is known from the Antilles (BERNHAUER & SCHEERPELTZ, 1926). 12. Atheta lurida Erichson (Figs 10, 37, 38, 46, 53) Atheta lurida Erichson, 1840, PACE 1985a. Type: see “material examined”. to black with rust-brown elytra, apex of abdomen, legs and 2-3 basal antennal GALAPAGOS ALEOCHARINE ROVE BEETLES 239 segments, antennal segments 4-11 mat black; moderately glossy; punctation fine and moderately dense, slightly asperate on forebody; meshed microsculpture clearly visible on forebody; pubescence moderately dense on pronotum and abdomen and sparse elsewhere; body sides with strong bristles; antennal segments 5-10 slightly transverse, at most 1.5 times wider than long (Fig. 10); head slightly elongate with pubescence directed medially and anteriad, temples as long as eye diameter, arcuate and narrowed posteriorly (Fig. 10); pronotum transverse, subequal in width to elytra, pubescence at midline of disc directed anteriad and laterally on the sides (Fig. 10): elytra transverse with pubescence directed straight posteriad or obliquely postero- laterad (Fig. 10); abdomen sparsely pubescent with 3 basal terga bearing basal impressions (Fig. 10). ¢ Median lobe of aedeagus with moderately large bulbous, tubus wide basally and tapering apically, internal sac with two subapical and median structures (Figs 37, 38); tergite 8 bearing two wide median and two narrow lateral teeth apically (Fig. 53); sternite 8 widely arcuate apically. 2 Spermatheca with elongate capsule and slightly posteriorly arched duct (Fig. 46); tergite 8 slightly emarginate apically; sternite 8 rounded posteriorly. Bionomics. Adults were collected from carrion in a shady ravine. Collecting methods: carrion baited traps. Altitudes: 400 m above sea level. Collecting period: May. Distribution. Atheta lurida was originally described from Brazil (ERICHSON 1840). PAcE (1985a) recorded this species from Galapagos: Santiago Gebirge, St. Cruz. We record it here for the first time from Fernandina. Because of its remote collecting site on pristine Fernandina Island, we interpret this as a natural dispersal, and the species to be native. It is interesting that we had no other collections of this species. Material examined. (4 specimens, 2 4,3 ©) (SBPC, HUB, JKC). Type: “lurida Er.[ichson] / Brasil Reich”, “6994”, “Type”, “Zool. Mus. Berlin” (HUB) 1 @, studied. Non type material: Fernandina: 10 km NE, Cabo Hammond, 400 m, 6-10.V.91, S. & J. Peck (SBPC, KE): 13. Atheta (Atheta) coriaria (Kraatz, 1856) (Figs 11, 47-49, 51) Homalota coriaria KRAATZ 1856, FRANK 1980, MUONA 1984. Aedeagus and spermatheca illustrated by HANSEN (1954), and LOHSE (1974). Syntypes: “Germany: Berlin (DEI) 6, sex undetermined, Leipzig “(DEI) 1 sex undetermined (Geadike 1981). Diagnosis. Body length 2.1-2.8 mm; narrowly subparallel (Fig. 11), dark brown to rust brown, usually with lighter pronotum, elytra, base and apex of abdomen, legs and base of antenna (segments 1-4); strongly glossy; punctation fine, slightly asperate, and dense on forebody; fine, meshed microsculpture clearly visible on head and pronotum; pubescence short and moderately dense except on abdomen; body sides with some bristles; antennal segments 5-10 transverse, at most almost twice as wide as long (Fig. 11); head approximately as long as wide, with temples arcuate and narrowing posteriorly, shorter than diameter of eye, pubescence directed anteriad (Fig. 11); pronotum transverse, pubescence directed anteriad along midline, and 240 JAN KLIMASZEWSKI & STEWART B. PECK lateroposteriad on sides of disc (Fig. 11); elytra slightly transverse, pubescence directed straight posteriad (Fig. 11); abdomen sparsely pubescent, strongly glossy and with 3 basal tergites strongly impressed basaly (Fig. 11). d Median lobe of aedeagus with large bulbous, and narrow and short triangularly dorsally shaped tubus (Fig. 48), internal sac with complex and strongly sclerotised structures (Figs 47, 48); tergite 8 slightly emarginate apically with two large lateral teeth and several medial teeth (Fig. 51); sternite 8 rounded apically. ? Spermatheca with horizontally subdivided capsule, and narrow and short posterior duct (Fig. 49): tergite 8 broadly arcuate apically; sternite $ truncate apically. Bionomics. Adults have been collected from transition forest, Scalesia forest, in Miconia forest, pampa zone, agricultural zone, litter under coffee, forest litter, rotting oranges, cow dung, tortoise dung and guava thicket. Collecting methods: sifting organic litter, Flight Interception Traps (FIT). Altitudes: 200 to 1000 m above sea level. Collecting period: February to May. In Europe A. coriaria is often found on mushrooms (Polyporaceae), and in compost (HANSEN 1954, LOHSE 1974). Larva described by Ashe (1984). Distribution. Europe, North America (California, Florida) (Frank 1980, Lohse 1974, Muona 1984). New island records: Floreana, Isabela, San Cristébal, and St. Cruz. Material examined. (82 specimens, 10 4,3 2, 69 sex undetermined) (SBPC, JKC). Floreana: 5, 6 km E Blackbeach (SBPC, JKC). Isabela: Sierra Negra (SBPC; JKC). San Cristobal: 1 km W Progresso (SBPC, JKC). St. Cruz: Bellavista (SBPC); Cueva Tres Entradas . (SBPC); Los Gemelos (SBPC); Sta. Rosa (SBPC). Remarks. An introduced European species recorded for the first time from the Galapagos Islands. Well established. Superficially similar to smaller A. clientula Erichson from South America but has differently formed genitalia. 14. Atheta (Acrotona) pseudoclaudiensis sp. n. (Figs 12, 41, 42, 44, 50) Diagnosis. Body length 1.3-2.0 mm; fusiform (Fig. 12), uniformly dark brown to black, often with legs slightly paler; moderately glossy; punctation dense and asperate on forebody and sparse on abdomen; microsculpture scarsely visible; pubescence short and dense except for abdomen; sides, especially on abdomen, with bristles; antennal segments 4-10 transverse and 7-10 approximately twice as wide as long (Fig. 12); head with temples about as long as diameter of eye, arcuate and gradually narrowing posteriorly, pubescence directed anteriad and anterolaterad (Fig. 12): pronotum transverse, pubescence directed laterad and posterolaterad from midline of dise (Fig. 12); abdomen strongly narrowing apically and pointed (Fig. 12), three basal tergites with basal impressions. Meso- and metatibia with strong bristle in basal half. d Median lobe of aedeagus with narrowly elongate bulbous and narrow subparallel apically truncate tubus (Fig. 42); internal sac with fine structures (Figs 41, 42). Tergite 8 truncate apically and with slightly emarginate base (Fig. 50). Sternite 8 widely arcuate apically and with straight base. 2 Spermatheca with small spherical GALAPAGOS ALEOCHARINE ROVE BEETLES 241 capsule and sinuate coiled posteriorly duct (Fig. 44). tergite and sternite 8 similar to those of male. Bionomics. Adults have been collected predominantly from tortoise dung, some from horse and cow dung, carrion, cormorant nest debris, in arid zone shrub and succulent litter, lagoon edge, tidal meadow, arid zone beach forest, Bursera forest, and Pisonia forest, transition forest, shrub forest, guava thicket, pampas. Collecting methods: shifting litter and dung, in dung and beer baited pitfall traps, Flight Interception Traps (FIT). Altitudes: 2 m to 1100 m above sea level. Collecting period: February to July. Distribution. New island records: Fernandina, Floreana, Isabela, Marchena, Pinta, San Cristébal, and St. Cruz. It is probably an endemic species. Material examined. 1234 specimens, 5 d, 11 2) (SBPC, JKC). Holotype (2): “Ecu.[ador], Galap.[agos], St. Cruz/Sta. Rosa 180 m/7.02.89, 89-34”, “S. Peck, Tortoise/Res.[erve], trans.[ition] for.[est]/tortoise dung” (SBPC). All remaining specimens listed bellow are considered paratypes. Fernandina: 5-10 km NE Cabo Hammond (JKC, SBPC). Floreana: 3-8 km E Black Beach, Cerro Pajas, Finca Cruz (SBPC). Isabela: Alcedo (Volcan), E crater, NE slope, 14 km NE Playa (JKC, SBPC); Bahia Urvina (SBPC); Bellavista, 21 km N (SBPC); Cerro Azul and vicinity (SBPC); Los Gemelos, 31 km N Sta. Rosa (JKC, SBPC); Sierra Negra (SBPC); St. Rosa, 15 km N (KJKC, SBPC); Tagus Cove, 2-10 km NE (JKC, SbPC); 13 km NW Villamil, Jabonocillo forest (SBPC). Marchena: Pta. Espejo (SBPC); SW Playa (SBPC). Pinta: Playa Ibbetson (SBPC). Plaza Sur: (SbPC). San Cristobal: El Junco (SBPC). St. Cruz: Bella Vista (SBPC); Charles Darwin Research Station (SBPC); Puntudo (SBPC); Sta. Rosa, 3 km N St. Cruz (SBPC); Tortoise Reserve (SBPC). Remarks. This species is externally similar to A. (A.) neolutea Pace from Brazil (PACE 1990a), A. (A.) praemeditata Pace from Argentina, A. (A.) hoyoana Scheerpeltz from Argentina and Chile (ScHEERPELTZ 1972, PACE 1987a, b), and A. (A.) magellanica Pace from Chile (PACE 1987b). It is distinct, however, in the shape of the spermatheca which bears a small spherical capsule with a characteristic invagination (Fig. 44), and two posterior coils and the shape and structures of the median lobe of the edeagus. This species has a strong affiliation with the mainland South American fauna of the genus. For illustrations of genitalia of the mainland species see PACE (1987a, b, 1990a). The genitalia of A. pseudoclaudiensis superficially resemblance those of the European A. alterrima (Grav.), and A. pusilla Brundin, however A. alterrima is on average twice larger and A. pusilla has differently shaped antennae (segments elon- gate) and median lobe of the aedoeagus. For illustrations and description of European species see BRUNDIN (1952). Atheta pseudoclaudiensis has a spermatheca very similar to that of A. claudiensis Pace (1990a) described from a unique female specimen collected in Brazil (Espirito Santo, Laranja de Terra, HUB). A. claudiensis however is smaller and lighter in color and the male of the species remains unknown. 15. Atheta dichroa (Gravenhorst) (Figs 13, 39, 40, 45, 54) Aleochara dichroa Gravenhorst, 1802; BLACKWELDER 1943; Moore & LEGNER 1975 (as Brundinia), PACE 1985a (as Atheta). Type material listed under ‘material examined”. Diagnosis. This species has two distinct color forms; dark and light, with the light being predominant. Specimens of the two forms are sympatric in distribution. 242 JAN KLIMASZEWSKI & STEWART B. PECK Dark form: uniformly brownish-black to blackish, or with elytra, tip of abdomen and 3 basal antennal segments, and tarsi or entire legs, usually paler and brownish; moderately glossy. Light form: dark-brown with light-brown elytra, tip of abdomen, legs and 2-3 basal antennal segments, head usually the darkest; moderately to strongly glossy. Body length 2.1-3.1 mm; subparallel (Fig. 13), punctation fine and moderately dense on forebody, slightly asperate especially on elytra; pubescence moderately dense on forebody and sparse on abdomen; body sides with bristles; meshed microsculpture clearly visible on forebody; antenna with segments 2-3 glossy and 4- 11 mat, segments 6-10 slightly transverse, less than 1.5 as wide as long (Fig. 13); head approximately round dorsally, flattened medially, with temples shorter than diameter of eye, and strongly narrowing posteriorly, pubescence directed medially and anteriorly (Fig. 13); pronotum nearly as wide as elytra at base, pubescence directed anteriorly on midline of disc and lateroposteriad laterally (Fig. 13); elytra transverse, pubescence directed posteriad (Fig. 13); abdomen subparallel and pointed apically (Fig. 13). d Aedeagus with moderately large, approximately oval bulbous (Fig. 40); tubus elongate, subparallel medially, and sharply pointed apically (Figs 39, 40); in lateral view venter of tubus straight, narrow apically and pointed ventrally (Fig. 39); internal sac with two well defined dark structures in bulbous and two pairs of inconspicuous structures in tubus (Fig. 40); tergite 8 with apical margin bearing two lateral sharp and two median rounded teeth (Fig. 54); sternite 8 rounded apically. ? Spermatheca with arched capsule bearing small conical projection posteriorly, and spherical ending of duct (Fig. 45); tergite 8 truncate apically; sternite 8 arcuate apically. Bionomics. Adults have been collected predominantly from carrion (e.g., dead tortoise), tortoise-, horse-, and cow dung, rotting Opuntia, forest litter, roseapple litter, cormorant nest debris, in littoral zone (e.g., lagoon edge), arid zone, agricultural zone, Bursera forest, Scalesia forest, Zonthoxylon-lichen humid forest, Miconia forest, mixed forest, shrub forest, open forest, guava thicket, /nga pods, pampas etc. Collecting methods: carrion traps, dung traps, light traps, Flight Interception Traps (FIT), Malaise traps, bottle traps, and general collecting techniques. Altitudes: 2 m to 1100 m above sea level. Collecting period: February to July. This is the most common aleocharine species on the Galapagos Islands. Distribution. The original series of this species consists of 6 specimens from South America and West Indies, St. Thomas (HUB), and subsequent records are from Grenada, St. John, St. Thomas, St. Vincent, Tortola; Bolivia and Galapagos (St. Cruz) (BLACKWELDER 1943, PACE 1985a). We judge it to be native. Atheta dichroa is also known to occur in North America (BLACKWELDER 1943, MOORE & LEGNER 1975). New island records: Baltra, Fernandina, Floreana, Isabela, Marchena, Pinta, Santiago, San Cristobal. Material examined. (1906 specimens, 41 d, 18 9, 1847 sex undetermined) (SBPC, JKC). Syntypes: “dichroa / Gr.[avenhorst] / Am.[erica] spt [=septemptrionalis], Zimm.[er- mann]”, “5423”, “Typus” (HUB) I d; “Americ.[a] sept.[emptrionalis] / Zimmermann / Nr. 5423”, “Typus” (HUB) 2 3, 1 2; “6994”, “Type” (HUB) 1 9; “St. Thomas / Moritz / 5424” (HUB) 1 ©. Non type material: Baltra: Arid zone (SBPC). Fernandina: 5-10 km NE Cabo GALAPAGOS ALEOCHARINE ROVE BEETLES 243 Hammond (SBPC); near summitt (SBPC). Floreana: 3-8 km E Black Beach (SBPC); Cerro Pajas (SBPC). Isabela: Alcedo (E crater rim, NE slope) (SBPC); Bahia Urvina (JKC, SBPC); Cerro Azul, 2-9 km NE (SBPC); Finca Cruz (SBPC); Sierra Negra (JKC, SBPC); Sto. Thomas and vicinity (SBPC); Tagus Cove, 2-9 km NE (JKC, SBPC); Villamil and 12 km NW Villamil (SBPC). Marchena: Pta. Espejo (SBPC); SW Playa (SBPC). Pinta: littoral zone, Playa Ibbetson (SBPC), Zanthoxylon-lichen forest (SBPC). Santiago: Aguacate Camp (SBPC); Central Camp (SBPC). San Cristobal: 2 km NE and 3 km E Baquerizo; 5 km E Wreck Bay (SBPC); Casetta (SBPC); El Junco and vicinity (JKC, SBPC); Progresso, 1-5 km E (SBPC). St. Cruz: Academy Bay (SBPC); Bellavista (SBPC); 3 km W, 2-4 km N Bellavista (JKC, SBPC); Devine Farm (SBPC); Charles Darwin Research Station; Pto. Ayora (SBPC); Horneman Farm (SBPC); Media Luna and vicinity (SBPC); Los Gemelos; 31 km N Sta. Rosa (JKC, SBPC); 7.2-10 km N Sta. Rosa; Tortoise Reserve (JKC, SBPC). Remarks. We consider the two color forms as belonging to the same species because they occur sympatrically, do not differ in external morphology, and have a similar shape of the median lobe of aedeagus and spermatheca. Atheta dichroa is externally very similar to A. propinqua (Er.) which is known from Brazil, but has a differently shaped spermatheca and should be considered as a different species. The female type of the latter species was examined and is deposited in HUM. VI. TRIBE Falagriini Mulsant & Rey, 1873 (First record of this tribe from Galapagos) Diagnosis. Tarsi 4-5-5-segmented, body ant-like (Fig. 14); head with distinctly constricted neck which is no more than one-third as wide as head (Fig. 14); pronotum narrow at base, no more than three-fourths maximum width, usually with median sulcus (Fig. 14): mesothoracic peritremes enlarged and sclerotised and almost always present behind procoxae; mesocoxal cavities moderately separated; maxillary palpus 4-segmented, labial palpus 3-segmented; legs long and slender. 16. Myrmecocephalus concinnus (Erichson) (Figs 14, 55-57) ERICHSON (1840) (Falagria). HOEBEKE 1985 (illustrations); PACE (1990a). Lectotype: “Brasil, Germ.[any], Hist. Coll. Nr. 5288 / Typus, Zool. Mus. Berlin” (HUB). Diagnosis. Body length 2.2-2.6 mm; ant-like (Fig. 14), with long and slender legs; brown to rust-brown, with rufous basal and apical portions of antennae, tibiae, tarsi, base and tip of abdomen and elytra, sometimes elytra appears rufous, mottled with dark brown; glossy; punctation fine and moderately dense on forebody; pu- bescence short, except slightly longer on abdomen; antennal segments 1-7 elongate and 8-10 as wide as long or slightly transverse (Fig. 14); head arcuately truncate at base, slightly wider than thorax (Fig. 14); pronotum elongate, convex, strongly converging at base, hind margin reflexed, deeply sulcate along median line (Fig. 14): scutellum partially carinate at base; elytra much wider than prothorax, with pronounced shoulders; abdomen narrowed at base and apex, 3 basal terga deeply impressed basally, first impression with some punctures. d Median lobe of aedeagus with large, widely oval bulbous, and short and apically truncate tubus (Figs 56, 57; internal sac with two sclerites in bulbous (Fig. 57). Tergite 8 truncate and without 244 JAN KLIMASZEWSKI & STEWART B. PECK comb of denticles on apical margin; sternite 8 rounded apically. 2 Spermatheca S- shaped (Fig. 55). Tergite and sternite 8 truncate apically. Bionomics. Adults have been collected from Miconia and Scalesia forest, and a guava filled ravine. Collecting methods: Flight Interception Traps (FIT). Altitudes: 370-550 m above sea level. Collecting period: February and March. Distribution. A cosmopolitan species originally described from Brazil (ERICHSON 1840), and recorded from Bolivia (PACE 1990b), and across the United States (HOEBEKE 1985). We report this species for the first time from Galapagos: San Cristobal, St. Cruz. We judge it to be introduced. Material examined. (19 specimens, 4 6,2 9, 13 unsexed (SBPG JK©) San Cristébal: 1 km E Progresso (SBPC) I sex undetermined. St. Cruz: Media Luna (SBPC, JKC); Los Gemelos (SBPC); Puntudo (SBPC). VII. TRIBE Oxypodini Thomson, 1859 Diagnosis. Tarsi 5-5-5-segmented, mesocoxae narrowly to moderately widely separated and set in margined acetabula, intercoxal process slender, median lobe of aedeagus with elongate compressor plate and without a transverse strip called the ‘athetine bridge’. Maxillary palpi 3 or 4-segmented, labial palpi 3-segmented. Genus Feluva Blackwelder, 1952 Type species: Feluva varicolor (Fauvel) (Brachyglossa) 17. Feluva franzi Pace, 1985a (Figs 15, 58-60) PACE 1985a: 454. Habitus and spermatheca illustrated by PACE (1985a). Holotype (¢ ): Isabela I., Cerro Azul, 5/5-1975, H. Franz leg. (RPC) Diagnosis. Body length 1.9-2.1 mm; narrowly subparallel (Fig. 15), dark brown or nearly black to pale brown with rust tinge, usually with darker antennal segments 5-11, head, pronotum and abdomen except for the apex; moderately glossy; punc- tation dense especially on head and pronotum; forebody with granulate surface; antennal segments 5-10 strongly transverse, each at least twice wider than long (Fig. 15); head with long (approximately as long as diameter of eye) subparallel temples, abruptly constricted posteriorly into narrow neck (Fig. 15); pronotum equal in width to head, slightly narrowed posteriorly, with pubescence directed horizontally from the median line of disc (Fig. 15); abdomen slightly widening posteriorly, three basal tergites with deep basal impressions (Fig. 15). 4 Median lobe of aedeagus with enlarged oval bulbous, and short triangular tubus in dorsal view, internal sac struc- tures inconspicuous (Figs 58, 59). Tergite 8 transverse, truncate apically, with several protruding setae, sternite 8 transverse, broadly rounded apically with several pro- truding setae. 2 Spermatheca with slightly elongate spherical capsule and approxi- mately two posterior coils (Fig. 60), tergite and sternite 8 elongate, truncate posteriorly, with protruding long setae. i ne ae ri GALAPAGOS ALEOCHARINE ROVE BEETLES 245 Bionomics. Adults have been collected from beach zone forests, arid zone, agriculture Zone, transition zone, and Scalesia and Miconia forests. Collecting methods: Malaise traps, Flight Interception Traps (FIT), carrion trap, and beating forest branches. Altitudes: 10 to 650 m above sea level. Collectintg period: January to May. Distribution. First recorded from Galapagos (Isabela, Marchena) by Pace (1985). Isabela and Marchena records are here confirmed. New island records: Floreana, Pinta, Rabida, St. Cruz. An endemic species. Material examined. (26 specimens: d, 2, sex undetermined) (SBPC, JKC): Floreana: 3 and 8 km E Black Beach (SBPC). Isabela: 4 km NW Villamil (SBPC); Cerro Azul, 2 km E Caleta Iguana (SBPC): Marchena: Pta. Espejo (SBPC); SW Playa (SBPC). Pinta: Playa Ibbetson (SBPC). Rabida: NE Coast, Palo Santo forest (SBPC). St. Cruz: Bellavista (SBPC, JKC); Charles Darwin Research Station, arid zone (SBPC); Los Gemelos (SBPC, JKC); Puntudo (SBPC, JKC); agricultural zone (SBPC); 1-7.2 km N. St. Rosa (SBPC). ACNOWLEDGMENTS We thank the curators of the institutions cited in section 2 for information and the loan of specimens under their care: especially D. Drugmand (IRSNB), R. Pace (RPC), M. Uhlig (HUC). We also appreciate the assistance of Roberto Pace who provided valuable information and line illustrations of genitalia of two Atheta species from South America. This study was partly supported by Natural Sciences and Engineering Research Council of Canada operating grants to S. B. Peck, and by grant number 5563-95 of the National Geographic Society. Field sampling during the 15 months spent in the field in the Galapagos Islands was by permit from the Galapagos National Park (F. Cepeda, Arturo Izurieta and E. Cruz, Superintendents, Department of Forestry, Ministry of Agriculture, Republic of Ecuador). Fieldwork was facilitated by logistical support from the Charles Darwin Research Station (CDRS), Isla Santa Cruz (G. Reck, D. Evans, C. Blanton, Directors). Field sampling was helped by J. K. Peck, J. Cook, J. Heraty, B. Sinclair, B. Landry, C. Vogel, S. Abedrabbo, Elvia Moraima Inca, Maria-Teresa Lasso, T. Finston, Eduardo Vilema and Lazaro Abelo Roque. TAME helped with reduced airfares. Contribution No. 537 of the Charles Darwin Research Station. 246 JAN KLIMASZEWSKI & STEWART B. PECK TABLE | Checklist of species Distribution Taxon Original Status Description (New Galapagos records with asterisk) Myllaenini Ganglbauer, 1885 confirmed 1. Myllaena leleupi Pace, 1985 Isabela confirmed probably endemic 2. Rothium ashlocki Ahn & Ashe, 1996 Sta. Cruz unconfirmed probably endemic 3. Rothium littoralis sp. n. Floreana probably endemic Hypocyphtini Laporte, 1835 new record 4. Oligota (Holobus) Kraatz, 1859 Africa, Madagascar, cosmopolitan, chrysopyga Mascarene Is., India, introduced New Caledonia, Sechelles, Sri Lanka; Isabela*, San Cristöbal*, Sta. Cruz* Homalotini Heer, 1839 confirmed 5. Phanerota sp. n. Sta. Cruz* probably introduced tridentata 6. Thecturota franzi Pace, 1985a Fernandina*, well established, Floreana*, Isabela*, Marchena*, Pinta*, Pinzön*, San Cristöbal*, probably endemic Santiago, Sta. Cruz 7. Diesota Pace, 1985a well established, Floreana*, Isabela*, (Apheloglossa Marchena*, status uncertain, franziana Pinzön*, Santiago”, possibly endemic San Cristébal*, Sta. Cruz 8. Diesota Pace Isabela, Sta. Cruz* well established, (Apheloglossa) probably endemic leleupi 9. Diesota Pace Marchena unconfirmed, (Apheloglossa) possibly endemic galapagosensis Placusini Mulsant & Rey, 1871 new record GALAPAGOS ALEOCHARINE ROVE BEETLES 247 10. Euvira scalesia sp. n. Sta. Cruz* probably endemic Athetini Casey, 1910 confirmed and new pen e) e dit ARE PU TOD enge records 11. Atheta Pace, 1985a Bartolomé*, well established, galapagoensis Espanola*, probably endemic Fernandina*, Floreana*, Isabela, Marchena*, Pinta, Pinzon, Plaza Sur*, Räbida*, Santiago, San Cristobal, Sta. Cruz, St. Fé* 12. Atheta lurida Erichson, 1840 Brazil; Santiago, established, Fernandina* native 13. Atheta (s. str.) Kraatz, 1856 Europe, North cosmopolitan, coriaria America; Floreana*, established, Isabela*, San introduced Cristobal, Sta. Cruz* 14. Atheta sp. n. Fernandina*, well established, (Acrotona) Floreana*, Isabela*, endemic pseudoclaudiensis Marchena*, Pinta*, Plaza Sur*, San Cristöbal*, Sta. Cruz* 15. Atheta dichroa Gravenhorst, 1802 North America, established, native South America (Bolivia), West Indies (St. Thomas); Baltra*?, Fernandina*, Floreana*, Isabela*, Marchena*, Pinta*, Santiago*, San Cristöba*, Sta. Cruz Falagriini Mulsant & Rey, new record 1873 16. Erichson, 1840 North America, established, Myrmecocephalus Bolivia, Brasil; San introduced concinnus Cristobal, Sta. Cruz* Oxypodini Thomson, 1859 confirmed 17. Feluva franzi Pace, 1985a Floreana*, Isabela, well established, Marchena, Pinta*, probably endemic Rabida*, Sta. Cruz* 248 JAN KLIMASZEWSKI & STEWART B. PECK TABLE 2 Aleocharine species distribution (including earlier literature records) in biotic zones on the Galapagos Islands and a total species diversity per zone as depicted in Table 1. (X) indicates presence; column one indicates species name, and its probable status. Columns 2-8 biotic zones. Last row contains information on species diversity per biotic zone (Litt.=littoral zone; Ar.=arid zone; Tr.=tranzition zone; HF.=humid forest; Es.=evergreen shrub; Pa.=pampa zone; Agr.=agricultural zone). Species and its probable status Litt. Ar. Dr HF. ES: Pa. Agr. Myllaena leleupi X (endemic) Rothium ashlocki X (endemic) Rothium littoralis X (endemic) Oligota chrysopyga X x X X X (introduced) Phanerota tridentata x (introduced) Thecturota franzi x X X X xX X (endemic) Diesota franziana X X X X X (probably endemic) Diesota leleupi X xX (endemic) Diesota galapagosensis x (endemic) Euvira scalesia X x (endemic) Atheta galapagoensis x X X x x x X (endemic) Atheta lurida xX xX (native) Atheta coriaria xX x x X X (introduced) Atheta pseudoclaudiensis x X x x x x X (endemic) Atheta dichroa xX xX X xX x x x (native) Myrmecocephalus concinnus x X (introduced) Feluva franzi x X X X X (endemic) Total Species diversity in bn sd O TOO 5 9) different zones Endemic and native species 6 og 7 6 4 6 | diversity in different zones . GALAPAGOS ALEOCHARINE ROVE BEETLES 249 Map | Map of the Galapagos Archipelago (numbers on contour lines indicate elevation in hundreds of meters). o DARWIN MARCHENA © @ GENOVESA SANTIAGO Seymour Daphne, O Br i \ Pinzon@ FERNANDINA SAN CRISTOBAL SANTA CRUZ ce Santa Fé ISABELA o, Gardner FLOREANA CÀ) ESPANOLA 10 20 30 40 50 KM 250 JA. JAN KLIMASZEWSKI & STEWART B. PECK \/ \ GALAPAGOS ALEOCHARINE ROVE BEETLES 251 77777 AAA zZ D) SICH == \ ER (ft N \ | \ fi Ò 2 Veh IG, Pipe PP ob, if pene aroreceay 7 VÀ D in) JAN KLIMASZEWSKI & STEWART B. PECK JAN KLIMASZEWSKI & STEWART B. PECK 254 GALAPAGOS ALEOCHARINE ROVE BEETLES 255 256 JAN KLIMASZEWSKI & STEWART B. PECK 58 59 60 Fics 1-8 Schematic habitus illustrations (pubescence shown only on right side of head, pronotum and abdomen; legs not shown). 1. Myllaena leleupi (length 1.8 mm); 2. Rothium littoralis (length 4.5 mm); 3. Oligota chrysopyga (length 0.9 mm); 4. Phanerota tridentata (length 1.3 mm); 5. Thecturota franzi (length 1.1 mm); 6. Diesota franziana (length 1.9 mm); 7. Diesota leleupi (length 2.2 mm); 8. Euvira scalesia (Length 1.8 mm). Fics 9-15 Schematic habitus illustrations (pubescence shown only on right side of head, pronotum and abdomen; legs not shown). 9. Atheta galapagoensis (length 2.2 mm); 10. Afheta lurida (length 2.8 mm); 11. Atheta coriaria (length 2.7 mm); 12. Atheta claudiensis (length 1.9 mm); 13. Atheta dichroa (length 2.8 mm); 14. Myrmecocephalus concinnus (Erichson) (length 2.4 mm); 15. Feluva franzi (length 2.0 mm). Fics 16-26 Genital structures and labial palpi of selected species. 16. Spermatheca of Myllaena leleupi; 17. spermatheca of Rothium littoralis; 18. labial palpi or glossae of À. littoralis; 19-21. Oligota chrysopyga: 19. spermatheca; 20. median lobe of aedeagus in lateral view; 21. median lobe of aedeagus in dorsal view; 22. Phanerota tridentata, median lobe of aedeagus in lateral view; 23- 25. Thecturota franzi: 23. median lobe of aedeagus in dorsal view; 24. median lobe of aedeagus in lateral view; 25. spermatheca; 26. spermatheca of Euvira scalesia. Fics 27-34 Genital structures and tergites of selected species. 27-30. Diesota franziana: 27. spermatheca; 28. apical portion of male tergite 8; 29. median lobe of aedeagus in lateral view; 30. median lobe of aedeagus in dorsal view; 31-34. Diesota leleupi: 31. spermatheca; 32. median lobe of aedeagus in lateral view; 33. median lobe of aedeagus in dorsal view; 34. apical portion of male tergite 8. GALAPAGOS ALEOCHARINE ROVE BEETLES 257 Fics 35-46 Genital structures of selected species. 35, 36, 43, Atheta galapagoensis: 35. median lobe of aedeagus in lateral view; 36. median lobe of aedeagus in dorsal view; 43. spermatheca; 37, 38, 46. Atheta lurida: 37. median lobe of aedeagus in lateral view; 38. median lobe of aedeagus in dorsal view: 46. spermatheca. 39, 40, 45. Atheta dichroa: 39. median lobe of aedeagus in lateral view; 40. median lobe of aedeagus in dorsal view; 45. spermatheca; 41, 42, 44. Atheta pseudoclaudiensis: 41. median lobe of aedeagus in lateral view; 42. median lobe of aedeagus in dorsal view; 44. spermatheca. Fics 47-57 Genital structures and tergites of selected species. 47-49, 51. Atheta coriaria: 47. median lobe of aedeagus in lateral view; 48. median lobe of aedeagus in dorsal view; 49. spermatheca. 51- 54. apical portion of male tergite 8:50. Atheta pseudoclaudiensis; 51. Atheta coriaria: 52. Atheta galapagoensis; 53: Atheta lurida; 54. Atheta dichroa. 55-57. Myrmecocephalus cingulatus: 55. spermatheca; 56. median lobe of aedeagus in lateral view; 57. median lobe of aedeagus in dorsal view. Fics 58-60 Feluva franzi: 58. median lobe of aedoeagus in dorsal view; 59. median lobe of aedeagus in lateral view; 60. spermatheca. REFERENCES AHN K.-J. & J.S. AsHE. 1966. 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Berlingska Bok- tryckeriet, Lund. 290 pp. VAN Dyke, E.C. 1953. The Coleoptera of the Galapagos Islands. California Academy of Sciences, Occasional Papers, 22: 181 pp. REVUE SUISSE DE ZOOLOGIE 105 (2): 261-318; juin 1998 Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae) Martin BAEHR Zoologische Staatssammlung, Münchhausenstr. 21, D-81247 München, Germany. Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae). - The genus Gunvorita Landin is revised and the following 13 new species from Nepal and northeastern India are described and illustrated: Gunvorita angusticeps, G. besucheti, G. depressipennis, G. hamifera, G. inermis, G. laeviceps, G. minor, G. nepalensis, G. ovaliceps, G. punctipennis, G. schawalleri, G. smetanai, and G. uncinata. The male genitalia of G. indica Darlington are described and figured. A key to all 16 known species is provided. Scenario's of the possible evolution and biogeographic history of the Oriental-Australian Leleupidiini are presented, even though the phylo- genetic relations of and within the three genera occurring in the Oriental region are not yet settled. Striking gaps in distribution of Leleupidiini in the Oriental region and some phylogenetic evidence indicate at least two, or even more probably, three independent colonization events of Leleu- pidiini in southern Asia: namely the genus Paraleleupidia to southern India, the ancestral stock of Colasidia to Malaysia, and the ancestral stock of Gunvorita to the pre-Himalayan area of Nepal to northeastern India. But another scenario is also conceivable with colonization of the ancestral stock of Colasidia and Gunvorita to Malaysia and with later spreading of Colasidia to the southeast and Gunvorita to the northwest. The immigration is believed to have occurred by drifting on the Indian plate (Paraleleupidia) and on terranes of the so-called "Sundaland" (Cola- sidia and Gunvorita). Consequently, the Oriental-Australian Leleupidiini are derived from ancestors that were formerly distributed in the part of Gondwanaland situated east to Africa. Therefore, Leleupidiini most pro- bably are of pre-Cretacious age, and they are a further example of rather recent immigration of an old Gondwanan element into the Australian region from the north. Key-words: Coleoptera - Carabidae - Zuphiinae - Leleupidiini - Gunvorita - Taxonomy - Oriental Region - Biogeography. Manuscript accepted 27.01.1998 262 MARTIN BAEHR INTRODUCTION As a second part of a general review of the Oriental Leleupidiini the genus Gunvorita is treated. Specimens of this genus, like Oriental Leleupidiini in general, until now were very rare in collections, though became increasingly numerous in the last few years, which is certainly due to more intense collecting and to specialized sampling methods. However, Leleupidiini are yet unrecorded from large parts of the Oriental region. Apparently, they either concentrate in few regions, or have been only sampled in these areas, because other regions have not been adequately worked until now. The history of the discovery of the Oriental Leleupidiini was briefly recorded in the first part of this review (BAEHR 1997b). Including the species of the genus Colasidia described in that paper and the new species of Gunvorita described in the present paper 49 species of Leleupidiini are now known from the Oriental and Australian Regions. Most Oriental-Australian species have been included in the genus Colasidia Basilewsky, but except for three species from South India that were arranged within the genus Paraleleupidia Basilewsky, subgenus Megaleleupidia Mateu, so far only three species were described as belonging to the genus Gunvorita Landin. Although the present classification of Oriental Leleupidiini is not really satisfactory, I think this generic concept should be maintained for the present. Hence those species are included in the genus Gunvorita that have a markedly elongate head with generally small eyes, a rather elongate, highly convex prothorax with inconspicuous lateral margins, possess rather distinct circular impressions medially of the eyes, though lack - at least on head and prothorax - the rather coarse puncturation present in the genus Colasidia. The crotchet on the lower surface of the apex of the aedeagus is highly characteristic in many species of Gunvorita, though a group of species lacks this character without showing striking differences in any characters of the external morphology. Therefore, characteristics of the aedeagus at present are not as useful for characterization of the genus Gunvorita as one might believe. There is some reason to believe that male genitalic characters generally can be used for better recognition of the genera, but at present male genitalia were known of little more than half of the species of Colasidia, and of no species at all of the genus Paraleleupidia. Moreover, the known aedeagi of Colasidia are very diverse and thus far do not give a clear picture. Hence, for the present, in Colasidia as well, male genitalia have been only used for distinction of species. By courtesy of Dr I. Löbl (Muséum d'histoire naturelle, Genève - MHNG) I received samples of the genus Gunvorita (and other Leleupidiini - see BAEHR 1997b) for identification that had been collected during the last 25 years by staff of the museum in different countries of southern Asia. When the present paper was almost ready for being printed, I received a further sample of specimens of the genus Gunvorita by courtesy of Dr W. Schawaller (Staatliches Museum fiir Naturkunde - SMNS) that were collected by Prof. J. Martens during his frequent expeditions to THE GENUS GUNVORITA 26 W Nepal, but also partly in the course of J. Martens' joint trips together with W. Schawaller. I decided to include this sample in the paper because it contains alto- gether 7 Gunvorita species of which 5 are new again. Hence, this paper covers 13 new species, and the three species known hitherto are reviewed briefly and in parts redescribed. Altogether, 76 specimens of Gunvorita were examined for this paper. MEASUREMENTS Measurements have been made under a stereo microscope using an ocular micrometer. Length has been measured from tip of labrum to apex of elytra, therefore, measurements may slightly differ from those of other authors. Length of head is measured from anterior border of clypeus to anterior border of "neck". The ratio length of orbit/length of eye is likewise measured to anterior border of "neck". CHARACTERS Best characters for differentiation of the species of the genus Gunvorita are in the structure of the male aedeagus that is available in all species of this genus. Fairly useful characters are also provided by shape of head, pronotum, and elytra (expressed in a number of measurements and ratios), size of eyes and of appendages of head, degree and shape of puncturation of upper surface, and pilosity. In other respects the species are rather similar. DEPOSITION OF TYPES The holotypes of the new species are deposited at the Muséum d'histoire naturelle, Genève (MHNG) and at Staatliches Museum fiir Naturkunde, Stuttgart (SMNH), some paratypes are deposited in the working collection of the author (CBM) at the Zoologische Staatssammlung, Miinchen. The types of the three yet described species are in Naturhistoriska Riksmuseet, Stockholm (NHRS), The Natural History Museum, London (BMNH), and Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt/M. (SMF). Genus Gunvorita Landin Gunvorita Landin, 1955: 467, fig. 90; DARLINGTON 1968: 208, fig. 1; MATEU 1981: 721, figs 4, 7; PERRAULT 1982: 76; CASALE 1985: 41, figs 1, 2; BAEHR 1988: 115; BAEHR 1990: 16; BAEHR 1991: 194. Type species: Gunvorita elegans Landin, 1955 In this genus species with the following characteristics are combined: head always posteriorly somewhat widened, shape of head either pentagonal or more or less triagonal; eyes small; clypeal suture with a pair of deep grooves; frons with a pair of circular or horseshoe-shaped grooves; pronotum narrow and dorsally convex, with inconspicuous lateral margin; elytra depressed, posteriorly conspiuously widened, 264 MARTIN BAEHR markedly triangular; puncturation on head and pronotum fine, on elytra fine to moderately coarse; pilosity elongate, hirsute, rather dense; male aedeagus almost always rather elongate, internal sac usually with two large sclerotized areas furnished with very elongate sclerotized teeth; female stylomere 2 always elongate, little curved, with elongate, acute apex, two elongate ventral ensiform setae, one elongate dorsal ensiform seta, a well developed nematiform seta, and stylomere 2 with 1-2 well visible nematiform seta(e) at median rim. It is yet unsettled, whether the combination of these character states is suffi- cient to distinguish Gunvorita from Oriental Colasidia or even from African Leleu- pidia. In certain of these character states species of Gunvorita remember (supposedly) primitive species of Colasidia from Malaysia (see Baehr 1997 b). Although the species are rather similar in external morphology, the aedeagi, in particular their apices, are highly characteristic in all species. Hence examination of male genitalia is indispensible, but species distinction by use of male genitalia is reasonably easy. KEY TO THE SPECIES OF THE GENUS Gunvorita LANDIN Because some of the published figures of the yet described species are of great value for identification, though are not included in the present paper, in the key the numbers of the respective figures have been added under the following chiffres: L55: Landin 1955; D68: Darlington 1968; M81: Mateu 1981; C85: Casale 1985. It should be mentioned, however, that the figure of the aedeagus ascribed to G. indica Darlington by Mateu (1981, fig. 8) actually does not show the aedeagus of this species. Identification of females may be difficult, unless they are associated with males. Although many species are very similar in certain external characters, I give an additional key using only external characters and ratios that may be useful for identification of females. Nevertheless, whenever females can be associated with males d genitalia should be examined. le Owen VEDRA oh High womens FAs oe ae RE 083 400 0:5 5 00.0 2 5 0 = < 2 > D - DO Lun As we tétanie oid ee create die dated E LIRE ne 18 DI Apex of aedeagus with crotchet at lower side, doubtful species under both couplets (MSI fig. 7; C85 fig. 2; figs. 1-9)... 22: RRSSAA 3 = Apex of aedeagus without crotchet at lower side (figs 10-14) ............ 13 3), Head posteriorly markedly widened, rather triagonal (L55 fig. 90; M81 fig. 4; C85 fig. 1; figs 37-40) and smaller species, length <4.5 mm and apical third of aedeagus distinctly bent up (M81 fig. 7; C85 fig. 2; figs Le mans be wt deci asd ASSE + - Head posteriorly widely rounded off (Figs 41-47); usually larger species, length mostly >4.5 mm; apical third of aedeagus not distinctly bent up (Figs 3-9); if in doubt of the latter, species > 5 mm long .......... 7 THE GENUS GUNVORITA 265 Eyes smaller in comparison to orbits (C85 fig. 1; fig. 38); aedeagus with remarkably slender apical part, apex semicircular, with minute denticle on lower side (C85 fig. 2). Eastern Nepal .......... martensi Casale Eyes larger in comparison to orbits (LSS fig. 90; figs 37, 39, 40): aedeagus with less slender apical part, apex not semicircular, with distmecterotchet((MiSdohisem/ tics IAD) cx 45% sett ee 5 Head markedly widened posteriorly, strongly triagonal (Fig. 40); lower surface of aedeagus with characteristic edge in middle, internal sac with short sclerotized area (Fig. 2). Northeastern India............ inermis sp. n. Head less widened posteriorly, less triagonal (M81 fig. 4; figs 37, 39); lower surface of aedeagus without edge in middle, internal sac with clonpatescleronzed'area (MSKRE TUST atone TEE UC 6 Aedeagus elongate, apex more delicate, apical third suddenly turned upwards, upper margin of apex not markedly upturned (Fig. 1); punc- turation of head very fine and sparse. Northeastern India . .... laeviceps Sp. n. Aedeagus shorter, apex shorter, stouter, less distinctly turned upwards, upper margin of apex distinctly upturned (M81 fig. 7); puncturation of head coarser and denser. Eastern Nepal, Sikkim, northeastern India A TC mn ebl'anatoiai elegans Landin Crotchet situated immediately at apex of aedeagus (figs 3-7); eyes slightly smaller, orbit >3.5 x as long as eye (Figs 41-45)................. 8 Crotchet considerably removed from apex of aedeagus (Figs 8, 9); eyes slightly larger, orbit <3.5 x as long as eye (Figs 46, 47)................ 12 Aedeagus markedly short and stout, with very elongate crotchet (Fig. 7); size small, length <4.3 mm; for further discrimination of 2 9 from © © of hamifera sp. n.: eyes smaller, orbit >3.75 x as long as eye (Fig. 45) and elytra slightly narrower, ratio 1/w <1.96 (Fig. 29). Eastern Nepal pre eel Shee od eet neon ne Manet weet schawalleri sp. n. Aedeagus longer and narrower, with shorter crotchet (Figs 3-6); size larger, usually >4.5 mm, except for small specimens of hamifera sp. n.; for further discrimination of 9 9 of hamifera sp. n. from 99% of schawalleri sp. n.: length >4.3 mm and eyes larger, orbit <3.75 x as long as eye (fig. 44) and elytra slightly wider, ratio 1/w >1.96 (Fig. 28) ..... 9 Head elongate, posteriorly markedly oval-shaped, ratio /w >1.6 (Figs 41, 42); apex of aedeagus somewhat upturned (Figs 3, 4). Northeastern India... 10 Head shorter, posteriorly shortly rounded, ratio I/w <1.6 (Figs 43, 44); ape xiOlaedeaeus barelyuptunned (igs) S TON RE Il Larger species, length 5.65 mm; head longer and narrower, ratio 1/w 1.78 (Fig. 41); prothoraxwider, ratio I/w 1.23 (Fig. 57); aedeagus longer and mioredelicate (Ei) Megha. RA indica Darlington Smaller species, length 5.05 mm; head shorter and wider, ratio 1/w 1.61 (Fig. 42); prothorax narrower, ratio I/w 1.31 (Fig. 58); aedeagus shorter didistoutera(Bio post trento re a ovaliceps sp. n. 266 MARTIN BAEHR Head longer and narrower with smaller eyes, ratio /w of head 1.51-1.60 (Fig. 43); elytra shorter, ratio /w 1.38-1.44, wider in relation to pro- thorax (Fig. 27): aedeagus with shorter crotchet and with one elongate, coiled, denticulate sclerite (Fig. 5). Central Nepal . . ..... nepalensis sp. n. Head shorter and wider with larger eyes, ratio I/w of head 1.42-1.47 (Fig. 44); elytra longer, ratio I/w 1.53-1.55, narrower in relation to prothorax (Fig. 28); aedeagus with longer crotchet and with two narrow, denticulate sclerites (cao) ME astem Nepales te RR RA TE hamifera sp. n. Larger species, length c. 5 mm; prothorax longer and narrower, ratio 1/w 1.21-1.27 (Fig. 62); elytra longer, ratio 1/w 1.55-1.60, >2.05 x as wide as prothorax (Fig. 30); apex of aedeagus remarkably curved up, upturned apex and crotchet form a very oblique line (Fig. 8). Central Nepal oe ba AR tri at fiir AT BD aA SOE SE EEE DIN RME smetanai sp. n. Smaller species, length <4.7 mm; prothorax shorter and wider, ratio 1/w 1.16-1.19 (Fig. 63); elytra shorter, ratio I/w 1.51-1.52, <1.90 x as wide as prothorax (Fig. 31); apex of aedeagus barely curved up, upturned apex and crotchet form a distinct angle (Fig. 9). Eastern Nepal...... uncinata sp. n. Aedeagus with remarkably slender apical part, apex semicircular, with minute denticle on lower side (C85 fig. 2). Eastern Nepal..... martensi Casale Aedeagus without any denticle on lower side of apex, apical part not remarkably slender (Pigs 10-14) eae 2. ee he Eee 14 Apex of aedeagus distinctly curved down, wide or slightly club-shaped as seen from below; genital ring rather wide (Figs 13, 14)................ IS Apex of aedeagus straight of faintly curved up, narrow and acute as seen from below; genital ring narrow and rather triangular (Figs 10-12) ......... 16 Head shorter and wider, posteriorly shortly rounded, eyes rather large, orbit <2.7 x as long as eye (Fig. 35); aedeagus shorter and stouter, apex slightly club-shaped as seen from below (Fig. 13). Northeast India EVEN IO II AIM ORI RT cca ew ot) a cy depressipennis sp. n. Head longer and narrower, posteriorly markedly ovalish, eyes small, orbit c. 3.75 x as long as eye (Fig. 36); aedeagus longer and narrower, apex wide as seen from below (Fig. 14). Nepal . .......... angusticeps Sp. n. Large species, length >4.8 mm; head posteriorly markedly ovalish (Fig. 48); lower surface of aedeagus gently bisinuate and denticulate sclerite in internal sac elongate (Fig. 10). Northeastern India, Darjeeling area SE hat SII, CELA MU DIL POE COR OOO SPO besucheti sp. n. Smaller species, length <4.0 mm; head posteriorly shortly rounded (Fig. 49) or markedly triangular (Fig. 50); lower surface of aedeagus either almost straight (Fig. 11) or denticulate sclerite in internal sac short (Fig. 12). Northeastern India, Khasi Hills, or Eastern Nepal............. Sl Head posteriorly shortly rounded (Fig. 49); prothorax longer and narrower, ratio I/w 1.38 (Fig. 65); lower surface of aedeagus straight (Fig!) sNortheastem India, Khasithhllss. ee eee eee minor Sp. n. 19: DD D DS? 24. THE GENUS GUNVORITA 267 Head posteriorly triangular (Fig. 50); prothorax shorter and wider, ratio I/w 1.23 (Fig. 66); lower surface of aedeagus bisinuate (Fig. 12). Eastern ING PAR non ORTO IRE ARTEN: punctipennis Sp. n. Posterior part of head elongate, markedly oval-shaped, head rather pentagonal, comparatively narrow and elongate, ratio /w >1.58 (Figs 41, 42, 48, 52); always rather large species, length >4.7 mm.............. 19 Posterior part of head either shortly rounded (Figs 43-47, 49, 51) or distinctly triagonal (L55 fig. 90; M81 fig. 4; C85 fig. 1; figs 37-40, 50), head not pentagonal, usually shorter and wider, ratio 1/w <1.60, com- monly far less; mostly smaller species, length rarely up to5.0mm......... 22 Very large species, length 5.65 mm; prothorax comparatively short and wide, ratio 1/w 1.23 (Fig. 57), ratio width pronotum/head 1.25. Northeast I ARS IRE IRE UNE Lo OR A O indica Darlington Smaller species, length <5.3 mm; prothorax longer and narrower, ratio I/w >1.27 (Figs 58, 64, 68), ratio width pronotum/head <1.18 (Figs 26, SANS CRE RTE On Sa Laka ES AL tete RE MO 20 Eyes comparatively large, orbit <3.25 x as long as eye (Fig. 48). Norineasternilindia gt Sree NE ate et tn Ra RR. besucheti sp. n. Eyesisinallertonbite>3: 75 xas longtasreye (E1ss'42452) i eae ae an. 21 Head longer and narrower, ratio 1/w 1.77 (Fig. 52); pronotum longer and narrower, ratio I/w 1.38, with comparatively wide base, <2 x as wide as elymallles On 68) Nepal ar wire. cere Na cherie angusticeps Sp. n. Head shorter and wider, ratio I/w 1.61 (Fig. 42); pronotum shorter and wider, ratio /w 1.31, with comparatively narrower base, >2 x as wide as elyirar(Ees\2 6458) Northeasterm India vie eos hors ovaliceps sp. n. Posterior part of head considerably widened, head markedly triagonal (LSS figs. 90; M81 fig. 4; C85 fig. 1; figs 37-40, 50); small species, lenethtaliwaysi<4:StmMmMIN® fe ER BR Er N ee IO). 02 23 Posterior part of head less widened, more or less rounded off, head not markedly triagonal (Figs 43-47, 49, 51); commonly larger species . ........ 27 Head rather elongate, ratio I/w 1.53; eyes comparatively small, orbit >3.5 XgasMlonptasteyei(C8Sthg gl ho738)aEast Nepal an MEET martensi Casale Head shorter, ratio I/w <1.47; eyes slightly larger, orbit <3.2 x as long as eye(E3533127904MS1 ne lies 37539. 40,50)8 rare MO IE 24 Eyes comparatively large, orbit slightly >2.5 x as long as eye (Fig. 50): whole surface, in particular elytra very coarsely punctate (Fig. 34). East ING pale ers osc A A RATS ER IN RE ee det punctipennis sp. n. Eyes smaller, orbit >3 x as long as eye (L55 fig. 90; MSI fig. 4; figs 37, SOA0)Surracclesscoarselipunctate (tie 2, >> DA) each RO Head longer and narrower, ratio I/w >1.42 (Figs 39, 40); prothorax longer and narrower, ratio I/w >1.23 (Figs 55, 56); generally smaller SPECIES Mem Ot A SON EE wet ee III 26 268 26 30. Go LD MARTIN BAEHR Head shorter and wider, ratio I/w <1.36 (Fig. 37); prothorax shorter and wider, ratio /w >1.17 (Fig. 53); generally larger species, length >4.25 mm. Eastern Nepal, Sikkim, northeastern India .............. elegans Landin Slightly larger species, length >4 mm; leytra slightly shorter, ratio 1/w <1.51 (Fig. 23); puncturation on head very sparse and fine (Fig. 39). Northeastermilndiaiss riens La es eee laeviceps sp. n. Slightly smaller species, length <3.9 mm; elytra slightly longer, ratio 1/w >1.52 (Fig. 24): puncturation on head denser and coarser (Fig. 40). Nontheasternindiaztze: pere hesitate inermis sp. n. Eyes larger, orbit <2.75 x as long as eye (Figs 49, 51). Northeastern India . . . 28 Eyes smaller, orbit >3.33 x as long as eye (Figs 43-47). Nepal ............ 29 Larger species, length >4.5 mm; pronotum slightly shorter and wider, ratio I/w <1.32 (Fig. 67); elytra markedly depressed, apical margin distinctly oblique and redressed towards suture (Fig. 35). . depressipennis sp. n. Smaller species, length 4 mm; pronotum slightly longer and narrower, ratio I/w 1.38 (Fig. 65); elytra less depressed, apical margin transverse, noteredressedstowardsisutune (Fiss33) 2 ap Sent oe PRE ERP minor Sp. n. Head posteriorly with rather elongate curvature, fairly oval-shaped (Fig. 43); elytra shorter, ratio I/w <1.44, perceptibly >2 x as wide as prothorax (io2#MaCEentraliNepalia:- 2324.23 eee ER TE nepalensis Sp. n. Head posteriorly with rather short curvature, less oval-shaped (Figs 44- 47); elytra longer, ratio 1/w >1.49, barely >2 x as wide as prothorax or narrowem(Figs; 2823) ekki: LOUE voleva dae EEE 30 Larger species, length >4.95 mm; head longer and narrower, posteriorly less widened, ratio I/w >1.55 (Fig. 46); elytra slightly longer and narrower rationl/w155) (Figs30)s Central Nepal ar ee smetanai sp. n. Smaller species, length <4.7 mm; head shorter and wider, posteriorly more widened, ratio /w <1.53 (Figs 44, 45, 47); elytra slightly shorter and wider, ratio I/w <1.55 (Figs 28, 29, 31). Eastern Nepal............... 31 Generally smaller species, length <4.3 mm; eyes generally smaller, orbit MOTS On AS eye (ie rdS) Bet EC I EEE schawalleri sp. n. Generally larger species, length >4.3 mm; eyes generally larger, orbit HS xaslong.asteye (Figsi44;'47)i... tea ate RI a 32 Head slightly shorter, posteriorly more widened, ratio 1/w <1.47; eyes slightly smaller, orbit >3.5 x as long as eye (Fig. 44); prothorax slightly longer and narrower, ratio /w >1.20 (Fig. 60); elytra slightly longer, ratio I/w >1.53, c. 2 x as wide as prothorax (Fig. 28).......... hamifera sp. n. Head slightly longer, posteriorly less widened, ratio I/w >1.50; eyes slightly larger, orbit <3.5 x as long as eye (Fig. 47); prothorax slightly shorter and wider, ratio I/w <1.19 (Fig. 63); elytra slightly shorter, ratio I/w <1.52, perceptibly <2 x as wide as prothorax (Fig. 31)..... uncinata sp. n. THE GENUS GUNVORITA 269 Gunvorita elegans Landin, 1955 Figs 21, 37, 53 Landin, 1955: 467, fig. 90; DARLINGTON 1968: 210; MATEU 1981, 721, fig. 7; CASALE 1985, 41. Note: The male genitalia of this species were figured by MATEU (1981) who examined the holotype. Therefore, there is no need to redescribe the male genitalia of this species. For the benefit of the user, however, measurements and ratios of some specimens alluded to this species by examination of the male genitalia are added. SUPPLEMENTARY DESCRIPTION: Measurements: Length: 4.25-4.45 mm; width: 1.5-1.55 mm. Ratios. Length/width of head: 1.35-1.36; length orbit/eye: 2.97-3.02; length/width of pronotum: 1.16-1.17; width widest part/base of pronotum: 1.76; width pronotum/head: 1.10-1.11; length/width of elytra: 1.46-1.48; width elytra/pronotum: 1.91. The aedeagi of both specimens mentioned below exactly match the figure of the holotype given by Mateu (1981, fig. 7). This species is now known from eastern Nepal, Sikkim, and northeastern India. New records: 1 d, Nepal Expeditionen Jochen Martens, 255 Ilam Distr., zw. Mai Pokhari, Mai Majuwa und Gitang Khola 2100-1800 m Kulturland 26 Aug 83 Martens & Daams leg. (SMNS); 1 6, Nepal Expeditionen Jochen Martens, 254a Ilam Distr. Mai Pokhari 2150-2250 m 23-25 Aug 1983 Berlese Martens & Daams leg (CBM). Gunvorita martensi Casale, 1985 Figs 22, 38, 54 Casale, 1985: 42. Type material: Holotype: 185, Nepal, Ilam Dist. zwischen Mai Pokhari und Ilam, 1330 m, quelliger Hang, Kulturland, Martens & Ausobsky | Apr. 80/ Holotypus d Gunvorita martensi n. sp. 1984 A. Casale Det. (SMF). Note. The measurements in Casale’s original description (CASALE 1985) do not agree with measurements made taken from the holotype which is actually considerably smaller than noted by Casale. Apart from that discrepancy, the description is detailed and reliable, and the male aedeagus is correctly figured. Therefore, the male genitalia are not redescribed. Diagnosis: Rather small species with elongate, distinctly triangular head, distinguished by odd shaped aedeagus with two characteristic edges on lower surface, elongate, rather straight though upturned apical part and a hemispherical apex that bears a tiny hook at the lower surface. SUPPLEMENTARY DESCRIPTION: Measurements: Length: 4.2 mm; width: 1.4 mm. Ratios. Length/width of head: 1.53; length orbit/eye: 3.67; length/width of pronotum: 1.29; width widest part/base of pronotum: 1.74; width pronotum/head: 1.09; length/width of elytra: 1.51; width elytra/pronotum: 2.05. This species is thus far known from the type locality in eastern Nepal only. 270 MARTIN BAEHR Gunvorita laeviceps sp. n. Figs 17159932333 Type material: Holotype: d India W. Bengal, Darjeeling distr. Sevoke 200 m. 7.X.78, Besuchet Löbl (MHNG). (MHNG). Paratypes: 1 9, same data (MHNG); 2 2 ©, India W. Bengal, Darjeeling distr. Kalimpong-Algarah 1400 m, 8.X.78 Besuchet Löbl (CBM, MHNG): 1 ©, India W. Bengal, Darjeeling distr. Singla 300 m, 17.X.78 Besuchet Löbl (MHNG). Diagnosis: Rather small species with posteriorly markedly triagonal head, further distinguished by shape of aedeagus with rather elongate, markedly upturned apical part and a very small crotchet situated shortly in front of apex. DESCRIPTION: Measurements: Length: 4.0-4.25 mm; width: 1.4-1.45 mm. Ratios. Length/width of head: 1.42-1.45; length orbit/eye: 2.98-3.20; length/width of pronotum: 1.23-1.27; width widest part/base of pronotum: 1.70-1.81; width pronotum/head: 1.10- 1.15; length/width of elytra: 1.50-1.51; width elytra/pronotum: 2.05-2.08. Colour: Piceous, labrum, palpı, legs, and antennae yellowish. Head: Comparatively short, posteriorly considerably widened, markedly triangular, widest near base, orbit posteriorly shortly rounded off. Upper surface gently convex. Eyes small, depressed, length c. 1/3 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a shallow, circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but compara- tively short. Antenna moderately elongate, slightly surpassing middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere <2/3 as long as Ist, <1.5 x as long as 2nd antennomere. Surface glossy,with traces of microreticulation only on clypeus. Puncturation sparse, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both supraorbital setae elongate, though not much longer than pilosity, posterior supraorbital seta situated far behind eye at base of head. Pronotum: Moderately elongate, rather cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, basal angles slightly projecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta shorter, situated right on basal angle. Surface without microreticulation, glossy, with moderately sparse, coarse puncturation. Distance between punctures c. as wide as ne THE GENUS GUNVORITA 271 FIG. 1 Gunvorita laeviceps sp. n. 3 genitalia: Aedeagus (left side), shape of apex (from below), left paramere, genital ring. Scale: 0.25 mm. diameter of punctures. Pilosity moderately sparse, elongate, hirsute, irregularly incli- ned, though rather erect. Elytra: Fairly elongate, triangular, laterally slightly curved, widest in posterior third, upper surface moderately convex, at base distinctly raised. Shoulders narrow, oblique, not projecting. Apex wide, faintly sinuate, slightly oblique, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation rather sparse, coarse, rather irregular, in apical and lateral parts punctures slightly finer. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, glossy. Pilosity rather sparse, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring narrow, elongate, ovalish-triangular, with moderate apical plate, almost symmetric. Aedeagus elongate, with elongate, suddenly upturned apical third that bears a very small crotchet shortly in front of apex. Apex very narrow seen from below. Lower surface almost straight in basal two thirds, then suddenly curved, then again straight. Internal sac in middle with large, coiled, markedly dentate sclerite, and with another dentate sclerite on top. For parameres see fig. 1, left paramere rather elongate, at apex evenly rounded. Right paramere unknown. Female genitalia: Stylomere 2 narrow and elongate with acute apex, with 2 elongate ventral ensiform setae the lower one being much shorter, one elongate 272 MARTIN BAEHR dorsal ensiform seta, and a nematiform seta situated in a large groove in middle between base and apex. Apex of stylomere 1 medially with 1 nematiform seta. Variation: The specimen from Singla somewhat varies in having a shorter and wider prothorax and in coarser puncturation on head. Etymology: The name refers to the smooth surface of the head. Distribution: Northeastern India, Darjeeling region. Collecting circumstances: Collected by seving of leaf litter at low to median altitudes. Gunvorita inermis sp. n. Figs 2, 16, 24, 40, 56 Type material: Holotype: d, India W. Bengal, Darjeeling distr. Mahanadi 1200 m, 6.X.78 Besuchet-Löbl (MHNG). Paratypes: 11 84,7 9 9, same data (CBM; MHNG); 1 6, 2 © 2, same locality, 19.X.78 (MHNG, ZSM). Diagnosis: Small species with posteriorly markedly triagonal head, further distinguished by shape of aedeagus with a characteristic edge in middle of lower surface, moderately elongate, slightly upturned apical part and a rather small crotchet sıtuated shortly at the very apex. DESCRIPTION: Measurements: Length: 3.6-3.9 mm; width: 1.2-1.3 mm. Ratios. Length/width of head: 1.44-1.47; length orbit/eye: 3.05-3.20; length(/width of pro- notum: 1.24-1.28; width widest part/base of pronotum: 1.71-1.76; width pronotum/ head: 1.05-1.11; length/width of elytra: 1.52-1.56; width elytra/pronotum: 1.93-2.02. Colour: More or less dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae yellowish. H ead: Comparatively short, posteriorly considerably widened, triangular, widest near base, orbit posteriorly shortly rounded off. Upper surface gently convex. Eyes small, laterally slightly produced, length slightly <1/3 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a shallow, ovalish to even rather horseshoe-shaped, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large, comparatively elongate. Antenna modera- tely elongate, slightly surpassing middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere distinctly <2/3 as long as Ist, c 1.3 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Punc- turation sparse, fine and superficial, distance between punctures about 3-4 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both THE GENUS GUNVORITA Te Fic. 2 Gunvorita inermis sp. n. 3 genitalia: Aedeagus (left side), shape of apex (from below), left and right parameres, genital ring. Scale: 0.25 mm. supraorbital setae elongate, though not much longer than pilosity, posterior supraorbital seta situated far behind eye at base of head. Pronotum: Moderately elongate, rather cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin convex in anterior half, then almost straight, sinuate just in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, basal angles slightly pro- jecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta slightly shorter, situated right on basal angle. Surface without microreticulation, glossy, with fairly dense, coarse puncturation. Distance between punctures slightly less wide than diameter of punctures. Pilosity moderately sparse, elongate, hirsute, 1rregu- larly inclined, though rather erect. Elytra: Fairly elongate, triangular, laterally slightly curved, widest in posterior third, upper surface moderately convex, at base slightly raised, odd intervals, especially the 5th, slightly raised in basal half. Shoulders moderately narrow, oblique, not projecting. Apex wide, absolutely straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation rather sparse, coarse, rather irregular, in apical and lateral parts punctures slightly finer. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very 274 MARTIN BAEHR difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and | pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, glossy. Pilosity rather sparse, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring narrow, elongate, ovalish-triangular, with narrow, elongate apical plate, slightly asymmetric. Aedeagus fairly elongate, with moderately elongate, gently upturned apical part that bears a small crotchet at the very apex. Apex narrow and acute seen from below. Lower surface with a characteristic edge about in middle, then gently bisinuate-oblique. Internal sac in middle with large though rather short, coiled, dentate sclerite. For parameres see fig. 2, both parameres rather elongate. Female genitalia: Stylomere 2 narrow and elongate with acute apex, with 2 elongate ventral ensiform setae the lower one being much shorter, one elongate dorsal ensiform seta, and a nematiform seta situated in a large groove in apical two fifth. Apex of stylomere | medially apparently with | nematiform seta. Variation: Little variation noted. Etymology: The name refers to the small apical hook at the aedeagus. Distribution: Northeastern India, Darjeeling region. Known only from type locality. Collecting circumstances: Collected by sieving of leaf litter at median altitude. Gunvorita indica Darlington, 1968 Figs 3; 25,415 ou Darlington, 1968: 208, fig. 1; MATEU 1981: 722, figs 5, 8 (erroneously!); CASALE 1985: 42. Type material: Holotype: 4, Holotype/ India: Darjeeling. Ghoom. 26.V.1931. Dr M. Cameron. B.M.1931-452./ Gen. et sp. nov. Place provisionally after Agastus/ Holotype Gunvorita indica Darl. (BMNH). Note: The figure given by MATEU (1981, fig. 8) of the very strange male genitalia ascribed to G. indica was taken from a specimen from Nepal that was not available for reeximantion. Unfortunately, Mateu failed to examine and compare this specimen (and additional specimens) with the holotype. It is not easily understood how Mateu came to a reliable identification using the raw sketch of the habitus of G. indica and the fairly poor description given by DARLINGTON (1968, fig. 1). Examination of the holotype of G. indica revealed a completely different aedeagus bearing a crotchet right at the apex. Hence, the specimen sketched by Mateu must represent another, yet un- described species. Diagnosis: Very large species with elongate, posteriorly rounded, not triangular though characteristically oblique head, further distinguished by elongate aedeagus with elongate, rather straight apical part and a small crotchet situated imme- diately at apex. SUPPLEMENTARY DESCRIPTION: Measurements: Length: 5.65 mm; width: 1.8 mm. Ratios. Length/width of head: 1.78; length orbit/eye: 3.93; length/width of pronotum: 1.23; width widest THE GENUS GUNVORITA D SI Un Fic. 3 Gunvorita indica Darlington. 5 genitalia. For legend see fig. 2. part/base of pronotum: 1.78; width pronotum/head: 1.25; length/width of elytra: 1.54; width elytra/pronotum: 2.02. Male genitalia: Genital ring narrow and elongate, almost parallel, with deep apex, rather symmetric. Aedeagus elongate, with elongate, almost straight, barely upturned apical part that bears a small crotchet at the very tip. Apex moderately wide seen from below. Lower surface gently convex. Internal sac in middle with large, coiled, dentate sclerite. For parameres see fig. 3, left paramere rather elongate, right paramere short. This species is thus far known only from the type locality in northeastern India. Gunvorita ovaliceps sp. n. Figs 4, 26, 42, 58 Type material: Holotype: d, India W. Bengal, Darjeeling dist. 13 km N. Ghoom 1500 m. 15.X.78, Besuchet Löbl (MHNG). Diagnosis: Large species with rounded, posteriorly markedly pentagonal head, further distinguished by shape of aedeagus with moderately elongate, fairly upturned apical part and a small crotchet situated at the very apex. DESCRIPTION: Measurements: Length: 5.05 mm; width: 1.6 mm. Ratios. Length/width of head: 1.60; lengh orbit/eye: 3.87; length/width of pronotum: 1.31; width widest part/ base of pronotum: 1.75; width pronotum/head: 1.14; length/width of elytra: 1.59; width elytra/pronotum: 1.94. 276 MARTIN BAEHR FIG. 4 Gunvorita ovaliceps sp. n. $ genitalia. For legend see fig. 2. Colour: Piceous, elytra slightly lighter than fore body, suture narrowly reddish. Labrum, palpi, legs, and antennae yellowish. Head: Elongate, oval-shaped, posteriorly markedly widened, rather distinctly pentagonal, widest in posterior third, orbit posteriorly rather oblique, rounded off. Upper surface gently convex. Eyes very small, laterally slightly produced, length about 1/4 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a rather deep, circular, slightly oblique groove on either side that posteriorly in middle combine to a more shallow triangular groove. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, trian- gular tooth. Labium anteriorly concave. Maxillary palpus rather elongate, apex obli- quely cut. Terminal segment of labial palpus large and very elongate. Antenna comparatively elongate, slightly surpassing middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere <2/3 as long as Ist, c. 1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Punc- turation very sparse especially on frons and vertex, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both supraorbital setae broken, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Rather elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly THE GENUS GUNVORITA 27/1) convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, basal angles slightly projecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior setae shorter, situated right on basal angles. Surface without microreticulation, glossy, with modera- tely sparse, rather coarse puncturation. Distance between punctures c. as wide as dia- meter of punctures, only near median line punctures somewhat finer and sparser. Pilosity moderately sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, odd intervals in anterior half slightly raised. Shoulders narrow, oblique, not projecting. Apex wide, faintly concave, slightly oblique, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation rather sparse, coarse, rather irregular, in apical and lateral parts punctures finer and somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and | pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, rather glossy. Pilosity rather sparse, fairly elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring rather narrow, ovalish-triangular, without distinct apical plate, slightly asymmetric. Aedeagus rather short, with short, distinctly upturned apical part that bears a fairly small crotchet at the very apex. Apex narrow as seen from below. Lower surface almost straight. Internal sac in middle with large, very elongate, coiled, markedly dentate sclerite. For parameres see fig. 4, left paramere triagonal and rather elongate. Female genitalia: Unknown. Variation: Unknown. Etymology: The name refers to the markedly ovalish shape of head. Distribution: Northeastern India, Darjeeling region. Known only from type locality. Collecting circumstances: Collected by sieving of leaf litter at median altitude. Gunvorita nepalensis sp. n. Figs 5, 17, 27, 43, 59 Type material: Holotype: 4, Nepal (Pr. Bagmati) Phulchauki nr. Kathmandu, 2300 m, 10.V.81, I. Lobl, 66 (MHNG). Paratypes: 10 dd,3 99, same data (CBM, MHNG); 2 6d, Nepal 424 Lalitpur Distr. Phulchoki Mt. 1800-2000 m 25.1V.1995 Martens & Schawaller (SMNS). Diagnosis: Large species with rounded, not triangular head, distinguished by shape of aedeagus with elongate, rather straight apical part and a small crotchet situated immediately at apex. 278 MARTIN BAEHR DESCRIPTION: Measurements: Length: 4.55-5.0 mm; width: 1.65-1.8 mm. Ratios. Length/width of head: 1.51-1.61; length orbit/eye: 3.73-3.95; length/width of pronotum: 1.20-1.26; width widest part/base of pronotum: 1.70-1.74; width pronotum/head: 1.14- 1.21; length/width of elytra: 1.38-1.44; width elytra/pronotum: 2.03-2.14. Colour: More or less dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae light reddish. Head: Elongate, oval-shaped, posteriorly widened, though not triangular, widest in posterior third, orbit posteriorly widely rounded off, rather oblique. Upper surface gently convex. Eyes small, depressed, length slightly >1/4 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but comparatively short. Antenna comparatively elongate, surpassing middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere <2/3 as long as Ist, c. 1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation sparse, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae elon- gate, though not much longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Rather elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin strongly convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, posterior angles slightly projecting, obtuse. Lateral margin inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta shorter, situated right on basal angle. Surface without microreticulation, glossy, with Sparse, very fine puncturation. Distance between punctures c. 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, odd intervals in anterior half slightly raised. Shoulders narrow, oblique, not projecting. Apex wide, almost straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly dense, rather coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse punc- THE GENUS GUNVORITA 279 FIG. 5 Gunvorita nepalensis Sp. n. d genitalia. For legend see fig. 2. turation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and | pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, rather glossy. Pilosity dense, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring wide, ovalish, widened to apex, rather symmetric. Aedeagus fairly elongate, with elongate, almost straight, barely upturned apical part that bears a small crotchet at the very tip. Apex narrow seen from below. Lower surface gently bisinuate. Internal sac in middle with large, coiled, strongly dentate sclerite. For parameres see fig. 5, left paramere rather elongate. Female genitalia: Stylomere 2 narrow and elongate with acute apex, with 2 elongate ventral ensiform setae, one elongate dorsal ensiform seta, and a nematiform seta situated in a large groove in middle between base and apex. Apex of stylomere | medially with 2-3 nematiform setae. Variation: Little variation noted apart from minor differences in relative shape of head, pronotum, and elytra. Etymology: The name refers to the range of this species. Distribution: Central Nepal. Known only from a small area around type locality. Collecting circumstances: Collected by sieving ground litter at median to fairly high altitudes. 280 MARTIN BAEHR Gunvorita hamifera sp. n. Figs 6, 18, 28, 44, 60 Type material: Holotype: 4, Nepal Expeditionen Jochen Martens, 412 Sankhua Sabha Distr., Arun Valley betw. Mure and Murure, mixed broad-leaved forest, 2050-2150 m, 9- 17 June 88 Martens & Schawaller leg. (SMNS). Paratypes: 246,1 ®, same data (CBM, SMNS); | ©, Nepal Expeditionen Jochen Martens, 414 Sankhua Sabha Distr., Arun Valley, Chichila, 1900-2000 m, Quercus forest, bushes near village, 10-20 June 88 Martens & Schawaller leg. (SMNS); 1 2, Nepal Expeditionen Jochen Martens, 297 Terhathum Distr., Tinjura Dara, 2450-2850 m, artenreicher Laubmischwald, 17 Sep 83 (Martens & Daams 1. (SMNS). Diagnosis: Medium sized species with rounded, not triangular head and small eyes, further distinguished by rather elongate aedeagus with elongate, rather straight apical part and a large crotchet situated immediately at apex. DES CRIPTION: Measurements: Length:4.3-4.65 mm; width: 1.45-1.55 mm. Ratios. Length/width of head: 1.42-1.47; length orbit/eye: 3.55-3.78; length/width of pronotum: 1.20-1.27; width widest part/base of pronotum: 1.69-1.77; lwidth pronotum/head: 1.05- 1.12; length/width of elytra: 1.53-1.55; width elytra/pronotum: 1.95-2.02. Colour: More or less dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae light reddish. Head: Elongate, oval-shaped, posteriorly widened, though not triangular, widest in posterior third, orbit posteriorly widely rounded off, fairly oblique. Upper surface gently convex. Eyes small, depressed, laterally little protruded, length slightly >1/4 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly pro- jecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, in middle with a large, shallow, about v-shaped groove the acute part of which is directed posteriad. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Men- tum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large, compara- tively elongate. Antenna comparatively elongate, almost attaining base of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere barely longer than 2/3 as long as Ist, c. 1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation sparse, very fine and superficial, dis- tance between punctures about 5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae elongate, perceptibly longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Rather elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin strongly convex in anterior two thirds, sinuate in front of pos- terior angles. Apex fairly wide, rather excised, anterior angles obtuse, slightly pro- THE GENUS GUNVORITA 281 FIG. 6 Gunvorita hamifera sp. n. 3 genitalia. For legend see fig. 2. jecting. Base narrow, laterally oblique but not excised, posterior angles slightly pro- jecting, obtuse. Lateral margin distinct though without distinct border line, marginal channel absent. Median line fine, barely impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta shorter, situated right on basal angle. Surface without microreticulation, glossy, with fairly sparse and fine puncturation. Distance between punctures c. 2-3 x as wide as diameter of punctures. Pilosity moderately sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, in most specimens odd intervals in anterior half slightly raised. Shoulders narrow, oblique, not projecting. Apex wide, almost straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly dense, moderately coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and | pore at apex of 3rd stria. Setae when present very elongate. Surface without microreticulation, glossy. Pilosity dense, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring moderately wide, ovalish, narrowed to apex, fairly symmetric. Aedeagus fairly elongate, with elongate, almost straight, barely upturned apical part that bears a large, very acute crotchet at the very tip. Apex moderately wide seen from below. Lower surface gently convex. Internal sac in middle 282 MARTIN BAEHR with two large, somewhat coiled, strongly dentate sclerites, one situated at left side near bottom, the other running from top of right side anteriad to bottom. For parameres see fig. 6, right paramere rather elongate. Female genitalia: Stylomere 2 narrow and elongate with acute apex, with 2 elongate ventral ensiform setae, one elongate dorsal ensiform seta, and a nematiform seta situated in a large groove in middle between base and apex. Apex of stylomere | medially with 1-2 nematiform setae. Variation: Little variation noted apart from minor differences in relative shape of head, pronotum, and elytra. Etymology: The name refers to the hamiform apex of the aedeagus. Distribution: Eastern Nepal, known from a restricted area around type locality. Central Nepal. Collecting circumstances: Collected by sieving ground litter in mixed broad-leaved forest, Quercus forest, and rich, mixed broad-leaved forest at rather high altitudes. Gunvorita schawalleri sp. n. Figs-/, 29; 45, 61 Type material: Holotype: d, Nepal Expeditionen Jochen Martens, 412 Sankhua Sabha Distr., Arun Valley betw. Mure and Murure, mixed broad-leaved forest, 2050-2150 m, 9- 17 June 88, J. Martens & W. Schawaller leg. (SMNS). Paratypes: 6 dd, same data (CBM, SMNS); 1 d, Nepal Expeditionen Jochen Martens, 414 Sankhua Sabha Distr., Arun Valley, Chichila, 1900-2000 m, Quercus forest, bushes near village, 10-20 June 88, J. Martens & W. Schawaller leg. (SMNS). Diagnosis: Rather small species with rounded, not triangular head and small eyes, easily distinguished by the very short and compact aedeagus bearing a large crotchet immediately at apex. DESCRIPTION: Measurements: Length: 4.1-4.3 mm: width: 1.4-1.45 mm. Ratios. Length/width of head: 1.45-1.49; length orbit/eye: 3.77-3.85; length/width of pronotum: 1.18-1.22; width widest part/base of pronotum: 1.75-1.76; width pronotum/head: 1.05- 1.11; length/width of elytra: 1.49-1.55; width elytra/pronotum: 1.90-1.96. Colour: More or less dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae reddish. Head: Elongate, oval-shaped, posteriorly widened, though not triangular, widest in posterior third, orbit posteriorly widely rounded off, rather oblique. Upper surface gently convex. Eyes small, depressed, laterally little protruded, length slightly >1/4 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly pro- jecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular THE GENUS GUNVORITA 283 Rio Gunvorita schawalleri sp. n. 3 genitalia. For legend see fig. 2. tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but comparatively short. Antenna rather short, attaining middle of pronotum. Median antennomeres as long as wide, 3rd antennomere <2/3 as long as Ist, <1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation sparse, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae elongate, perceptibly longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Moderately elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin strongly convex in anterior two thirds, slightly sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, posterior angles slightly projecting, obtuse. Lateral margin without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta slightly shorter, situated right on basal angle. Surface without microreticulation, glossy, with rather sparse and fine puncturation. Distance between punctures c. 3 x as wide as diameter of punctures. Pilosity rather sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Moderately elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, odd intervals in anterior half faintly raised. Shoulders comparatively wide, oblique, not projecting. Apex wide, almost straight, 284 MARTIN BAEHR transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly dense, rather coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and 1 pore at apex of 3rd stria. Setae when present very elongate. Surface without microreticulation, glossy. Pilosity fairly dense, elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring wide, ovalish, strongly narrowed to apex, rather symmetric. Aedeagus markedly short and compact, with short and wide, almost straight, slightly upturned apical part that bears a very large, acute crotchet at the very tip. Apex very wide seen from below. Lower surface gently convex. Internal sac in middle with two large, coiled, strongly dentate sclerites, one situated on left side near the top, the other near bottom on the right side. For parameres see fig. 7, left paramere rather elongate, right short. Female genitalia: Unknown. Variation: Little variation noted apart from minor differences in relative shape of head, pronotum, and elytra. Etymology: The name is a patronym in honour of W. Schawaller. Distribution: Eastern Nepal, known from a small area around type locality. Collecting circumstances: Collected by sieving ground litter in mixed broad-leaved forest and Quercus forest at fairly high altitudes. Gunvorita smetanai sp. n. Figs 8, 30, 46, 62 Type material: Holotype: d, Nepal (Pr. Bagmati) Pokhare NE Barahbise 2800 m, 2.V.81, I. Löbl & A. Smetana (MHNG). Paratype: 1 d, same data (CBM). Diagnosis: Large species with rounded, not triangular head, distinguished by shape of aedeagus with moderately elongate, markedly upturned apical part and a large crotchet situated far from apex. DESCRIPTION: Measurements: Length: 4.95-5.0 mm; width: 1.65 mm. Ratios. Length/width of head: 1.55-1.59; length orbit/eye: 3.55-3.57; length/width of pronotum: 1.21-1.27; width widest part/base of pronotum: 1.66-1.68; width pronotum/head: 1.15- 1.18; length/width of elytra: 1.55-1.60; width elytra/pronotum: 2.05-2.06. Colour: More or less dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae yellowish. Head: Elongate, oval-shaped, posteriorly widened, though not triangular, widest in posterior third, orbit posteriorly widely rounded off. Upper surface gently convex. Eyes small, depressed, length considerably >1/4 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral THE GENUS GUNVORITA 285 Fig. 8 Gunvorita smetanai sp. n. d genitalia. For legend see fig. 2. [=] [=] (©) angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a shallow, circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but comparatively short. Antenna comparatively elongate, reaching middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere <2/3 as long as Ist, c. 1.5 x as long as 2nd antennomere. Surface glossy, with traces of micro- reticulation only on clypeus. Puncturation sparse, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both supraorbital setae elongate, though not much longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Rather elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, basal angles slightly projecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior mar- ginal seta elongate, situated at anterior third of pronotum, posterior seta shorter, situated 286 MARTIN BAEHR right on basal angle. Surface without microreticulation, glossy, with moderately sparse, fine puncturation. Distance between punctures c. 3-4 x as wide as diameter of punctures. Pilosity moderately sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, odd intervals in anterior half slightly raised. Shoulders narrow, oblique, not projecting. Apex wide, almost straight, slightly oblique, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly dense, rather coarse, rather irregular, in apical and lateral parts punctures finer and somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and | pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, rather glossy. Pilosity dense, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring narrow, ovalish, with large apical plate, slightly asymmetric. Aedeagus rather short and stout, with stout, moderately elongate, markedly upturned apical part that bears a large crotchet far behind apex. Apex modera- tely wide seen from below. Lower surface gently convex. Internal sac in middle with large, coiled, strongly dentate sclerite. For parameres see fig. 8, left paramere rather elongate. Female genitalia: Unknown. Variation: Due to small number of available specimens, little variation noted. Etymology: The name is a patronym of Dr A. Smetana, collector of this species. Distribution: Central Nepal. Known only from type locality. Collecting circumstances: Collected by sieving of leaf litter at rather high altitudes. Gunvorita uncinata sp. n. Figs 9, 31, 47, 63 Type material: Holotype: 4, Nepal Expeditionen Jochen Martens, 352 Taplejung Distr. above Yamputhin, left bank of Kabeli Khola, bushes, open forest, 1800-2000 m, 27-29 Apr 1988, J. Martens & W. Schawaller leg. (SMNS). Paratype: 1 6, Nepal Expeditionen Jochen Martens, 351 Taplejung Distr., Yamputhin, cultural land, open forest, 1650-1800 m, 26 Apr.-1 May 1988, J. Martens & W. Schawaller leg. (CBM). Diagnosis: Medium sized to fairly large species with rounded, not triangular head and rather small eyes, distinguished by the rather elongate aedeagus with elongate, somewhat spoon-shaped apical part that bears a small crotchet far remo- ved from apex. DESCRIPTION: Measurements: Length: 4.5-4.7 mm; width: 1.55-1.6 mm. Ratios. Length/width of head: 1.50-1.53 length orbit/eye: 3.38-3.43; length/width of pronotum: THE GENUS GUNVORITA 287 Fic. 9 Gunvorita uncinata sp. n. 3 genitalia. For legend see fig. 2. 1.16-1.19; width widest part/base of pronotum: 1.68-1.73; width pronotum/head: 1.16- 1.18; length/width of elytra: 1.51-1.52; width elytra/pronotum: 1.82-1.88. Colour: Rather dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae light reddish. Head: Elongate, oval-shaped, posteriorly widened, though not triangular, widest in posterior third, orbit posteriorly widely rounded off. Upper surface gently convex. Eyes rather small, slightly convex, somewhat protruded laterally, length slightly <1/3 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) dis- tinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a deep, circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but comparatively elongate. Antenna comparatively short, barely surpassing middle of pronotum. Median antenno- meres c. as long as wide, 3rd antennomere <2/3 as long as Ist, c. 1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Punc- turation sparse, very fine and superficial, distance between punctures about 4 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae elongate, perceptibly longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature. 288 MARTIN BAEHR Pronotum: Comparatively short, rather cordiform, perceptibly longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin strongly convex in anterior two thirds, sinuate in front of posterior angles. Apex wide, considerably excised, anterior angles obtuse, slightly projecting. Base rather narrow, laterally oblique but not excised, posterior angles barely projecting, obtuse, prebasal part of lateral margin almost parallel. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta slightly shorter, situated right on basal angle. Surface without microreticulation, glossy, with moderately dense, fairly coarse puncturation. Distance between punctures c. 1-2 x as wide as diameter of punctures. Pilosity moderately dense, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Moderately elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, odd intervals in anterior half slightly raised. Shoulders fairly narrow, oblique, not projecting. Apex wide, almost straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly dense, coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse punc- turation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation,glossy. Pilosita dense, rather elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring moderately wide, rather parallel, narrowed to apex, rather symmetric, basal part elongate. Aedeagus fairly elongate, with elongate, almost straight, barely upturned apical part that bears a medium-sized crotchet far removed from the tip. Apex rather wide though tapering seen from below. Lower surface gently bisinuate. Internal sac in middle with elongate, coiled, strongly dentate sclerite. For parameres see fig. 9, both parameres rather elongate. Female genitalia: Unknown. Variation: Little variation noted apart from minor differences in relative shape of head, pronotum, and elytra. Etymology: The name refers to the uncinate apex of the aedeagus. Distribution: Eastern Nepal, known from a small area around type locality. Collecting circumstances: Collected by sieving ground litter in bushes and open forest, partly on cultivated land, at median to fairly high altitudes. Gunvorita besucheti sp. n. Figs 10, 19, 32, 48, 64 Type material: Holotype: d, India W. Bengal, Darjeeling dist. Algarah 1800 m. 8.X.78, C. Besuchet & I. Löbl (MHNG). Paratypes: 2 99, same data (CBM, MHNG); 1 3, India W. Bengal, Darjeeling dist. Algarah-Labha 1900 m. 11.X.78, C. Besuchet & I. Lôbl (MHNG). THE GENUS GUNVORITA 289 Fig. 10 Gunvorita besucheti sp. n. 3 genitalia. For legend see fig. 2. Diagnosis: Large species with posteriorly rather pentagonal head, further distinguished by shape of the very elongate aedeagus with slightly upturned apical part and without a crotchet. DESCRIPTION: Measurements: Length: 4.8-5.3 mm; width: 1.55-1.65 mm. Ratios. Length/width of head: 1.58-1.70; length orbit/eye: 3.11-3.25; length/width of pronotum: 1.27-1.35; width widest part/base of pronotum: 1.63-1.70; width pronotum/head: 1.11- 1.18; length/width of elytra: 1.53-1.60; width elytra/pronotum: 2.02-2.10. Colour: Dark piceous to almost blackish, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae yellowish, basal antennomere and femora reddish. Head: Elongate, oval-shaped, posteriorly markedly widened, rather distinctly pentagonal, widest in posterior third, orbit posteriorly rather oblique, rounded off. Upper surface gently convex. Eyes small, laterally slightly produced, length slightly <1/3 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly pro- jecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a rather shallow, circular to somwehat triangular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Men- tum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus remarkably stout, widened to apex, apex very obliquely cut. Terminal segment of labial palpus large and very elongate. Antenna comparatively elongate, slightly surpassing 290 MARTIN BAEHR middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere barely more than half as long as Ist, barely longer than 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation very sparse especially on frons and vertex, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures, middle of frons and vertex almost impunctate. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both supra- orbital setae very elongate, posterior supraorbital seta situated far behind eye at beginning of basal curvature. Pronotum: Rather elongate, fairly cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, rather deeply excised, anterior angles obtuse, markedly projecting. Base narrow, laterally oblique but not excised, basal angles slightly projecting, obtuse, Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta slightly shorter, situated right on basal angle. Surface without microreticulation, glossy, with moderately sparse, rather coarse puncturation. Distance between punctures c. as wide as diameter of punctures, only near median line punctures somewhat finer and sparser. Pilosity moderately sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, barely triangular, laterally weakly curved, widest in posterior third, upper surface depressed, odd intervals in middle slightly raised. Shoul- ders wide, oblique, not projecting. Apex wide, straight, slightly oblique, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation sparse, remar- kably irregular, in basal half coarse, becoming very fine and sparse behind middle. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 2 postmedian, 4-5 apical pores, and | pore at apex of 3rd stria. Setae very elon- gate. Surface without microreticulation, glossy. Pilosity rather sparse, elongate, hirsute, irregular, inclined posteriorly, though rather erect. Male genitalia: Genital ring rather narrow, elongate, ovalish-triangular, with small, asymmetrically curved apical plate. Aedeagus very elongate, with small, gently upturned apex that lacks a crotchet. Apex narrow and acute seen from below. Lower surface gently bisinuate. Internal sac in middle with large, elongate, coiled, dentate sclerite. For parameres see fig. 10, both parameres rather short. Female genitalia: Stylomere 2 narrow and elongate with narrow, acute apex, with 2 elongate ventral ensiform setae the lower one being much shorter, one elongate dorsal ensiform seta, and a nematiform seta situated in a large groove in middle between base and apex. Apex of stylomere | medially with 1 nematiform seta. Variation. Some variation noted in relative width of head, prothorax, and elytra, and in distinctness of puncturation of surface of head. Etymology: The name is a patronym of Dr C. Besuchet, collector of this species. THE GENUS GUNVORITA 29] Distribution: Northeastern India, Darjeeling region. Known only from the Algarah area. Collecting circumstances: Collected by sieving of leaf litter at median altitude. Gunvorita minor sp. n. Figs 11, 33, 49, 65 Type material: Holotype: d, India Meghalaya, Khasi Hills 27.X.78 Mawsynram- Balal 1000 m, C. Besuchet & I. Löbl, 33B (MHNG). Diagnosis: Rather small species with posteriorly widened though neither pentagonal nor triagonal head, further distinguished by shape of the elongate, at lower surface almost straight aedeagus with faintly upturned apex that lacks a crotchet. DESCRIPTION: Measurements: Length: 4.0 mm; width: 1.32 mm. Ratios. Length/width of head: 1.49; length orbit/eye: 2.63: length/width of pronotum: 1.38; width widest part/ base of pronotum: 1.63; width pronotum/head: 1.06; length/width of elytra: 1.56; width elytra/pronotum: 1.97. Colour: Piceous, head slightly darker than rest of surface. Labrum, palpi, legs, and antennae yellowish. Head: Rather elongate, oval-shaped, posteriorly slightly widened, though neither triagonal nor pentagonal, widest in posterior third, orbit posteriorly widely rounded off. Upper surface gently convex. Eyes comparatively large, though rather depressed, length slightly <2/5 of orbit length to beginning of curvature. Clypeus anteriorly rather convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a rather shallow, rather irregular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus stout, apex obliquely cut. Terminal segment of labial palpus large and rather elongate. Antenna comparatively short, barely surpassing middle of pronotum. Median antennomeres c. as long as wide, 3rd antennomere slightly <2/3 as long as Ist, slightly >1.33 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation very sparse especially on frons and vertex, moderately fine, distance between punctures c. 4-5 x as wide as diameter of punctures, laterally puncturation denser, distance slightly less. Pilosity sparse, elongate, hirsute, erect, inclined anteriorly. Both supraorbital setae very elongate, posterior supraorbital seta situated far behind eye behind beginning of basal curvature. Pronotum: Fairly elongate, gently cordiform, considerably longer than wide, barely wider than head, widest in anterior third. Upper surface convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, gently excised, anterior angles obtuse, slightly projecting. Base narrow, laterally 292 MARTIN BAEHR Fic. 11 Gunvorita minor sp. n. d genitalia. For legend see fig. 2. oblique but not excised, basal angles slightly projecting, obtuse. Lateral margin rather inconspicuous, slightly raised, marginal channel narrow, almost absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated slightly behind anterior third of pronotum, both posterior setae broken, situated right on basal angles. Surface without microreticulation, glossy, with rather sparse, coarse puncturation. Distance between punctures slightly smaller than diameter of punctures. Pilosity rather sparse, moderately elongate, hirsute, irregularly inclined. Elytra: Rather elongate, narrowly triangular, laterally weakly curved, widest in posterior third, upper surface fairly depressed. Shoulders rather wide, oblique, not projecting. Apex wide, straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation rather sparse, coarse, becoming finer and sparser towards apex. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, appa- rently consisting of 8 basal, 2 postmedian, 5-6 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, glossy. Pilosity rather sparse, moderately elongate, hirsute, irregular, inclined posteriorly, rather depressed. Male genitalia: Genital ring elongate, moderately wide, rather triangular, without distinct apical plate, gently asymmetric. Aedeagus elongate, with narrow, rather elongate, very faintly upturned apex that lacks a crotchet. Apex narrow and acute seen from below. Lower surface almost straight. Internal sac in middle with large, elongate, coiled, dentate sclerite. For parameres see fig. 11, both parameres rather short. Female genitalia: Unknown. THE GENUS GUNVORITA 293 Variation: Unknown. Etymology: The name refers to the lesser size in comparison with the foregoing species from the same area. Distribution: Northeastern India, Khasi Hills. Known only from type locality. Collecting circumstances: Collected by sieving of leaf litter at median altitude. Gunvorita punctipennis sp. n. Figs 12, 34, 50, 66 Type material: Holotype: d, Nepal Expeditionen Jochen Martens, 308 Ilam Distr., Sanishare 5 km N, feet of Siwalik Mts, 270-300 m, mixed Shorea forest, 3.-5. April 1988, J. Martens & W. Schawaller leg. (SMNS). Diagnosis: Small species with rather triangular head, distinguished by elongate aedeagus with elongate, rather straight apical part without a crotchet, gently bisinuate lower surface, and slightly upturned apex. DESCRIPTION: Measurements: Length: 3.8 mm; width: 1.3 mm. Ratios. Length/width of head: 1.46; length orbit/eye: 2.64; length/width of pronotum: 1.23; width widest part/base of pronotum: 1.73; width pronotum/head: 1.13; length/width of elytra: 1.54; width elytra/pronotum: 1.94. Colour: Rather dark piceous, pronotum and elytral faintly lighter, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae yellowish. Head: Comparatively short, posteriorly widened, perceptibly triangular, widest in posterior quarter, orbit posteriorly very shortly and widely rounded off, rather straight. Upper surface gently convex. Eyes rather small, slightly convex, laterally slightly protruded, length slightly <2/5 of orbit length to beginning of curvature. Clypeus anteriorly slightly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a circular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large, fairly elongate. Antenna comparatively short, attaining middle of pronotum. Median antennomeres as long as wide, 3rd antennomere <2/3 as long as Ist, <1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation sparse, very fine and superficial, distance between punctures about 4-5 x as wide as diameter of punctures. Pilosity sparse, elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae very elongate, much longer than pilosity, posterior supraorbital seta situated very far behind eye at beginning of basal curvature. Pronotum: Moderately elongate, cordiform, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. 294 MARTIN BAEHR Fig. 12 Gunvorita punctipennis sp. n. d genitalia. For legend see fig. 2. Lateral margin strongly convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, distinctly excised, anterior angles obtuse, slightly projecting. Base narrow, laterally oblique but not excised, posterior angles slightly projecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Ante- rior marginal seta situated at anterior third of pronotum, both setae broken, posterior seta rather short, situated right on basal angle. Surface without microreticulation, glossy, with moderately sparse, coarse, even somewhat vermiculose puncturation. Dis- tance between punctures c. 1-2 x as wide as diameter of punctures. Pilosity rather sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, all intervals in anterior half rather distinctly raised. Shoulders comparatively wide, oblique, not projecting. Apex wide, almost straight, transverse, not redressed to suture. Striae irregularly marked by rows of punctures, puncturation fairly sparse, though very coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly reco- gnizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, glossy. Pilosity moderately dense, rather elongate, hirsute, irregular, inclined poste- riorly, fairly depressed. THE GENUS GUNVORITA 295 Male genitalia: Genital ring narrow, elongate, fairly ovalish, narrowed to apex, rather symmetric, with elongate and apically acute basal part. Aedeagus elon- gate, with elongate, almost straight, faintly upturned apical part, without a crotchet at apex. Apex narrow and acute seen from below. Lower surface gently bisinuate. Internal sac in middle with large, though short, coiled, strongly dentate sclerite. For parameres see fig. 12, left paramere rather elongate, right paramere short. Female genitalia: Unknown. Variation: Unknown. Etymology: The name refers to the markedly punctate elytra of this species. Distribution: Eastern Nepal. Known only from type locality. Collecting circumstances: Collected by sieving ground litter in Shorea forest of low altitude. Gunvorita depressipennis sp. n. Figs 18 2. 02SE51E07 Type material: Holotype: d, India Meghalaya Khasi Hills Nongpoh 700 m. 5.X1.78 C. Besuchet & I. Löbl (MHNG). Paratypes: 2 99, same data (CBM, MHNG). Diagnosis: Medium-sized species with posteriorly widened though neither pentagonal nor triagonal head, further distinguished by shape of the fairly short, at lower surface markedly bisinuate aedeagus with distinctly downcurved apex that lacks a crotchet. DESCRIPTION: Measurements: Length: 4.5-4.6 mm; width: 1.45-1.55 mm. Ratios. Length/width of head: 1.49-1.54; length orbit/eye: 2.47-2.70; length/width of pronotum: 1.26-1.32; width widest part/base of pronotum: 1.56-1.60; width pronotum/head: 1.08- 1.14; length/width of elytra: 1.49-1.56; width elytra/pronotum: 1.94-2.0. Colour: Piceous, labrum, palpi, legs, and antennae yellowish. Head: Elongate, oval-shaped, posteriorly slightly widened, though neither triagonal nor pentagonal, widest in posterior third, orbit posteriorly widely rounded off. Upper surface gently convex. Eyes comparatively large, though rather depressed, length slightly <2/5 of orbit length to beginning of curvature. Clypeus anteriorly rather convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side two hairs. Clypeal suture posterio-laterally with a large, deep groove each side. Frons convex, between eyes with a rather shallow, rather irregular, slightly oblique groove on either side. Labrum anteriorly barely excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Men- tum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus fairly stout, apex obliquely cut. Terminal segment of labial palpus large and rather elongate. Antenna comparatively elongate, slightly surpassing middle of pronotum. Median antennomeres slightly longer than wide, 3rd antennomere c. 2/3 as long as Ist, slightly <1.5 x as long as 2nd antennomere. Surface glossy, with traces of micro- 296 MARTIN BAEHR Fig. 13 Gunvorita depressipennis sp. n. d genitalia. For legend see fig. 2. reticulation only on clypeus. Puncturation very sparse especially on frons and vertex, moderately fine, distance between punctures c. 4 x as wide as diameter of punctures, laterally distance slightly less. Pilosity sparse, elongate, hirsute, erect, inclined anterior- ly. Both supraorbital setae very elongate, posterior supraorbital seta situated far behind eye behind beginning of basal curvature. Pronotum: Rather elongate, narrow, gently cordiform, considerably longer than wide, slightly wider than head, widest in anterior third. Upper surface moderately convex, basally highly convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, rather deeply, v-shaped excised, anterior angles obtuse, markedly projecting. Base narrow, laterally oblique but not excised, basal angles slightly projecting, obtuse. Lateral margin rather conspicuous, slightly upturned, marginal channel narrow though marked. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta slightly shorter, situated right on basal angle. Surface without microreticulation, glossy, with rather sparse, coarse puncturation. Distance between punctures considerably smaller than diameter of punctures. Pilosity rather sparse, moderately elongate, hirsute, irregularly inclined. Elytra: Elongate, narrowly triangular, laterally weakly curved, widest in posterior fifth or quarter, upper surface markedly depressed. Shoulders wide, oblique, faintly projecting. Apex wide, straight, markedly oblique, distinctly redressed to suture. Striae irregularly marked by rows of punctures, puncturation sparse, in apical third irregular, very coarse, becoming finer and sparser behind middle. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores THE GENUS GUNVORITA 297 very difficult to detect when setae broken, apparently consisting of 8 basal, 2 post- median, 4 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, glossy. Pilosity rather sparse, moderately elongate, hirsute, irregular, inclined posteriorly, rather depressed. Male genitalia: Genital ring moderately wide and elongate, ovalish, with large apical plate, gently asymmetric. Aedeagus rather short, with small, distinctly downcurved apex that lacks a crotchet. Apex rather narrow, slightly club-shaped seen from below. Lower surface markedly bisinuate. Internal sac in middle with large, elongate, coiled, dentate sclerite, in basal part with another, complexly folded, non- dentate sclerite. For parameres see fig. 13, both parameres rather short. Female genitalia: Stylomere 2 narrow and elongate with acute apex, with 2 elongate ventral ensiform setae the lower one being much shorter, one elongate dorsal ensiform seta, and a nematiform seta situated in a large groove in apical two fifth. Apex of stylomere 1 medially apparently with 1 nematiform seta. Variation: Little variation noted. Etymology: The name refers to the depressed surface of elytra. Distribution: Northeastern India, Khasi Hills. Known only from type locality. Collecting circumstances: Collected by sieving of leaf litter at rather low altitude. Gunvorita angusticeps sp. n. Figs 14, 36, 52, 68 Type material: Holotype: d, Nepal 426 Kaski Dist. above Pothana 2000 m, 27.- 29.1V.1995, Martens & Schawaller (SMNS). Diagnosis: Medium sized species with characteristic elongate, oval- shaped, not triangular head, further distinguished by very sparse pilosity on head and pronotum and by aedeagus with elongate, rather straight, apically wide apical part that does not bear a crotchet but is distinctly curved down. DESCRIPTION: Measurements: Length: 4.7 mm; width: 1.5 mm. Ratios. Length/width of head: 1.77; length orbit/eye: 3.78; length/width of pronotum: 1.38; width widest part/base of pronotum: 1.58; width pronotum/head: 1.09; length/width of elytra: 1.56; width elytra/pronotum: 2.10. Colour: Rather dark piceous, suture of elytra narrowly reddish. Labrum, palpi, legs, and antennae light reddish. Head: Very elongate, characteristically oval-shaped, posteriorly widened, widest in front of posterior third, orbit posteriorly very widely rounded off, very elongately oblique, by no means triangular. Upper surface gently convex. Eyes small, depressed, length slightly >1/4 of orbit length to beginning of curvature. Clypeus posteriorly markedly convex, surface in middle convex, uneven, lateral angles (above base of antenna) distinctly projecting. Clypeal seta far removed from apex, at apex on either side apparently a single hair. Clypeal suture posterio-laterally with a deep groove 298 MARTIN BAEHR Fig. 14 Gunvorita angusticeps Sp. n. d genitalia. For legend see fig. 2. each side. Frons convex, between eyes with a circular, slightly oblique groove on either side. Labrum anteriorly slightly excised, 6-setose, inner 4 setae slightly shorter, lateral margin rather sparsely pilose. Mandibles short, at apex sharply incurved. Mentum with short and wide, triangular tooth. Labium anteriorly concave. Maxillary palpus rather short, apex obliquely cut. Terminal segment of labial palpus very large but compara- tively short. Antenna comparatively short, barely attaining middle of pronotum. Median antennomeres as long as wide, 3rd antennomere <2/3 as long as Ist, <1.5 x as long as 2nd antennomere. Surface glossy, with traces of microreticulation only on clypeus. Puncturation extremely sparse and fine, superficial, distance between punctures about 6-8 x as wide as diameter of punctures. Pilosity very sparse, very elongate, hirsute, rather erect, inclined anteriorly. Both supraorbital setae elongate, though not much longer than pilosity, posterior supraorbital seta situated far behind eye at beginning of basal curvature, though due to elongate basal part of head far more anteriorly than in other species. Pronotum: Elongate, narrow, less cordiform than in other species, considerably longer than wide, distinctly wider than head, widest in anterior third. Upper surface markedly convex. Lateral margin convex in anterior two thirds, sinuate in front of posterior angles. Apex fairly wide, slightly excised, anterior angles obtuse, slightly projecting. Base comparatively wide, laterally oblique but not excised, pos- terior angles slightly projecting, obtuse. Lateral margin rather inconspicuous, without distinct border line, marginal channel absent. Median line fine, not impressed. Prebasal grooves almost absent. Anterior marginal seta elongate, situated at anterior third of pronotum, posterior seta shorter, situated right on basal angle. Surface without THE GENUS GUNVORITA 299 microreticulation, glossy, with very sparse and fine puncturation. Distance between punctures c. 5-6 x as wide as diameter of punctures. Pilosity very sparse, elongate, hirsute, irregularly inclined, though rather erect. Elytra: Rather elongate, triangular, laterally slightly curved, widest in posterior third, upper surface depressed, intervals barely raised. Shoulders compara- tively wide, oblique, not projecting. Apex wide, almost straight, transverse, not re- dressed to suture. Striae irregularly marked by rows of punctures, puncturation rather sparse, coarse, rather irregular, in apical and lateral parts punctures somewhat rasp-like. Fixed setae in third interval hardly recognizable within the coarse puncturation. Series of marginal pores very difficult to detect when setae broken, apparently consisting of 8 basal, 3 postmedian, 6 apical pores, and 1 pore at apex of 3rd stria. Setae very elongate. Surface without microreticulation, rather glossy. Pilosity fairly sparse, elongate, hirsute, irregular, inclined posteriorly, fairly depressed. Male genitalia: Genital ring wide, ovalish, slightly widened to apex, fairly symmetric. Aedeagus fairly elongate, with elongate, almost straight apical part that is distinctly downcurved though lacks an apical crotchet. Apex wide seen from below. Lower surface bisinuate. Internal sac in middle with elongate, coiled, strongly dentate sclerite. For parameres see fig. 14, left paramere fairly large and short, right paramere rather elongate. Female genitalia: Unknown. Variation: Unknown. Etymology: The name refers to the conspicuously narrow head of this species. Distribution: Eastern Nepal. Known only from type locality. Collecting circumstances: Collected by sieving ground litter in fairly high altitude. Y (| \ a, | | 15 16 17 18 19 20 Fics 15-20 2 stylomere 2 and base of stylomere 1. 15. Gunvorita laeviceps sp. n.; 16. G. inermis sp. n. 17. G. nepalensis sp. n.; 18. G. hamifera sp. n.; 19. G. besucheti sp. n.; 20. G. depressipennis sp. n. 300 MARTIN BAEHR APPENDIX Because measurements and rations are rather useful in species differentiation, the used ratios for all Gunvorita species are compiled in the following table. TAB. | Measurements and ratios of the species of the genus Gunvorita. L. Length (in mm). I/w h. Length/width of head. I o/e. Length of orbit/eye. I/w pr. Length/width of pronotum. w d/b. Width widest diameter/base of pronotum. w pr/h. Width of pronotum/head. I/w el. Length/width of elytra. w el/pr. Width of elytra/pronotum. HT. Holotype. Species L /w h l o/e I/w pr G. angusticeps 4.7 hadi) 3.78 1.38 G. besucheti 4.8-5.3 1.58-1.70 3.11-3.25 1.27-1.35 G. depressipennis 4.5-4.6 1.49-1.54 2.47-2.70 1.26-1.32 G. elegans 4.25-4.45 1.35-1.36 2.97-3.02 1.16-1.17 G. hamifera 4.3-4.65 1.42-1.47 3.55-3.78 1.20-1.27 G. indica HT 5.65 1.78 3.93 1.23 G. inermis 3.6-3.9 1.44-1.47 3.05-3.20 1.24-1.28 G. laeviceps 4.0-4.25 1.42-1.45 2.98-3.20 (12341027 G. martensi HT 4.2 1.53 3.67 1.29 G. minor 4.0 1.49 2.63 1.38 G. nepalensis 4.55-5.0 1.51-1.60 3.73-3.95 1.20-1.26 G. ovaliceps 5105 1.61 3.87 Lesa G. punctipennis 3.8 1.46 2.64 1923 G. schawalleri 4.1-4.3 1.45-1.49 3.77-3.85 1.18-1.22 G. smetanai 4.95-5.0 ESS IRSO 333.371 1.21-1.27 G. uncinata 4.5-4.7 1.50-1.53 3.38-3.43 1.16-1.19 w d/b w pr/h I/w el w el/pr G. angusticeps 1.58 1.09 1.56 2.10 G. besucheti 1.63-1.70 1.11-1.18 1.53-1.60 2.02-2.10 G. depressipennis 1.56-1.60 1.08-1.14 1.49-1.56 1.94-2.00 G. elegans 1.76 1.10-1.11 1.46-1.48 1.91 G. hamifera 1.69-1.77 1.05-1.12 1.53-1.59 1.95-2.02 G. indica HT 1.78 1625 1.54 2.02 G. inermis 1.71-1.76 1.05-1.11 1.52-1.56 1.93-2.02 G. laeviceps 1.70-1.81 1.10-1.15 1.50-1.51 2.05-2.08 G. martensi HT 1.74 1.09 RSI 2.05 G. minor 1.63 1.06 1.56 1.97 G. nepalensis 1.70-1.74 1.14-1.21 1.38-1.44 2.03-2.14 G. ovaliceps 975 1.14 159 1.94 G. punctipennis 173 113 1.54 1.94 G. schawalleri 1.75-1.76 1.05-1.11 1.49-1.55 1.90-1.96 G. smetanai 1.66-1.68 1.15-1.18 1.55-1.60 2.05-2.06 G. uncinata 1.68-1.73 1.16-1.18 SS 1.82-1.88 BIOGEOGRAPHIC REMARKS Even when the actual distribution of the Oriental-Australian Leleupidiini is by no means adequately recorded, certain striking distribution patterns can be recognized: The genus Gunvorita is apparently confined to the pre-Himalayan areas of Nepal, THE GENUS GUNVORITA 301 Sikkim, and adjacent northern India; the genus Paraleleupidia, subgenus Megaleleu- pidia is indigenous to a rather restricted montane area in South India; and the genus Colasidia is distributed from Malaysia through Sumatra and northern Borneo to Papua New Guinea and northeastern Australia (Fig. 69, tab. 2). But this extensive range of the Oriental Leleupidiini is by no means continuous, though bears large and striking gaps, from where until now no records are available: Ceylon (Sri Lanka), central India, Burma, Thailand, Indochina, southern China, Java, the Indonesian part of Borneo, the Philippines, the Moluccas including Celebes (Sulawesi), and Western New Guinea (Irian Jaya). In some areas (e. g. southern Borneo, Java, western New Guinea) Leleu- pidiini do presumably occur and I think they will be detected when these areas are more appropriately sampled. In other countries, however, such as main parts of Burma, Thailand, Indochina, southern China, perhaps also the Philippines, Leleupidiini perhaps do not occur. The reason for this prediction is the supposed biogeographical history of the subfamily in the Oriental region. TAB. 2 Current distribution of the Oriental- Australian Leleupidiini. = Number of species Country Gunvorita Colasidia Paraleleupidia Nepal 10 Sikkim l Northeast India 8 South India 3 Malaysia Sumatra Sarawak (northern Borneo) Sabah (northern Borneo) Borneo altogether (only northern part) Papua New Guinea Australia (northern Queensland) — | © D Un -J © 00 Total number of species 16 30 3 Leleupidiini occur outside of the Oriental-Australian regions only in central and eastern Africa (where they have been discovered and where they are still most numerous) and they are thus an Old World Tropical faunal element. At the first glance Leleupidiini could be supposed to belong to the so-called Old Gondwanan faunal element which was always adapted to subtropical to tropical conditions. Remnants of this element are still present in subtropical and tropical parts of South America, Africa, and Australia. But this faunal element is usually well represented in large parts of Australia, whereas Leleupidiini in Australia have a very restricted range in the northeast and presumably immigrated rather recently from the north, probably via New Guinea. Where, therefore, the Oriental- Australian Leleupidiini came from? 302 MARTIN BAEHR Apparently, the species of Paraleleupidia of southern India are more closely related to their African counterparts of the same genus, than to either the Asiatic Gunvorita or Colasidia. This may be evidence of their independent biogeographical history and of their different place of origin. The different biogeographical history, on the other hand, may explain the apparent gaps in the present distribution of Leleupidiini in the Oriental region. For explanation of these questions a short summary of the present ideas about the geological history of the southern continents is required that refers mainly to RIDD (1968), MCKENNA (1973), SHIELDS (1979, 1983), POWELL et al. (1981), OWEN (1983), KEAST (1983), AUDLEY-CHARLES (1987), TRUSWELL et al. (1987), SCOTESE et al. (1988), VEEVERS ef al. (1991), BURRETT ef al. (1991), DALY et al. 1991, CRANSTON & NAUMAN (1991), and DE BOER (1995). According to present knowledge the first breakup of the southern landmass (Gondwanaland) in a western and eastern part began in late Jurassic (c. 155 mio b.p.), when a deep sea trench formed between present South America + Africa in the west and present Antarctica + India + Australia in the east. This seaway which became later the Indian Ocean spread southwards and subsequently India began its northwards mo- vement. At the same time separation of South America and Africa began from the north through development of a narrow seaway, the later South Atlantic, and subsequently Africa drifted likewise northwards under a certain amount of rotation. Also separation of Australia had commenced in the late Jurassic by the development of a deep seaway in a west to east direction between Antarctica and the western part of Australia. Com- plete segregation of Australia from Antarctica, however, occurred in early Tertiary, when the seaway between the Indian and Pacific Oceans was completed. Since its separation from Antarctica, Australia drifted steadily northwards, until it came in contact with the south Asiatic insular belt in mid Miocene (c. 15 mio b.p.). There is some reason to believe that southern Asia (excluding India) is not a compact block but is composed from rather small components of different origin and geological history, because certain parts (terranes) of Gondwanaland that formerly had been located east of Africa drifted likewise northwards and eventually came in contact with the northern continent Laurasia or with those parts of it that now build central Asia. Especially the terranes of the so called “Sundaland” (present Malaysia, southern Thailand and Burma, and the Greater Sunda Islands, most probably even northeastern India and Nepal) were originally situated between later Africa and Australia and drifted away from Gondwanaland during late Jurassic or early Cretaceous. Unfortunately (see below) it is still unsettled, whether these terranes always formed a more or less conti- nuous landmass, whether they were always separated and arrived at different times and in different places at the Laurasian continent. It is further unknown, to which extent they were submerged during their drifting history. Certainly the mentioned terranes arrived at their present position prior to the time when Australia came in contact with southern Asia — this 1s likewise demonstrated by geological and biogeographical evidence. However, during late Tertiary as well as during most of Pleistocene Sundaland was more or less isolated from the mainland of Asia and this is true even for those components of Sundaland that are now part of the THE GENUS GUNVORITA 303 Asian mainland, e.g. Malaysia, southwestern Thailand, and perhaps even northeastern India, Sikkim, and Nepal. On the basis of the sketched geological and geographical evidence and of the present distribution scheme, the following scenario of the biogeographical history of the Oriental-Australian Leleupidiini is suggested: The Oriental Leleupidiini are not monophyletic, because at least the species of Paraleleupidia are more closely related to African Leleupidiini than to the remaining Oriental genera, and they arrived at their present position in the Oriental region by independent drift on the Indian plate. This would also explain the apparent distributional gap of Leleupidiini in central India. The genus Colasidia apparently developed in that part of Gondwanaland that was originally located east of Africa, and later the original stock of Colasidia drifted on terranes of the later Sundaland to the north, to meet southern Asia where it subse- quently diversified. Unfortunately the phylogenetic relations of and within the genus Colasidia are still uncertain due to limited material and too little knowledge of the male genitalia, and to the lack of a general generic revision of the Leleupidiini as a whole (see BAEHR 1997b). However, in the following I dare a very preliminary attempt to fix the supposed plesiomorphic state for some external characters. Although this does not implicitly mean that apomorphic states could not have evolved convergently, there is some reason to believe that they are synapomorphic states so that the more derived species would build up a monophyletic group. I regard the following character states as plesiomorphic (mainly by outgroup comparison with other leleupidiine genera): e large size e rather elongate body shape e triangular shape of elytra e depressed dorsal surface e fine and dense puncturation of surface e dense and diffuse puncturation of elytra e dense, hirsute pilosity e large eyes e rounded or ovalish head the posterior part of which is not strikingly widened e elongate antenna e simple male aedeagus without striking features at apex and within internal sac. If these character polarizations are correct, then the most primitive members of the genus Colasidia have been thus far observed on the Malayan Peninsula (in parti- cular C. oviceps, C. depressa, C. rougemonti, C. lagadiga, C. malavica, C. triangularis, and C. attenuata in about that sequence), but not elsewhere. Nevertheless, even on the Malayan Peninsula some rather advanced species exist. All species hitherto recorded from Borneo, New Guinea, and Australia, and most species from Sumatra, on the other hand, are in most respects apomorphic and perhaps form a monophyletic unit, and a single species from northernmost Sumatra only (C. denticollis) seems less advanced and occupies a somewhat intermediate position. 304 MARTIN BAEHR Fic. 69 Current distribution of the Oriental Leleupidiini. Genus Colasidia: — — — — — ; genus Gunvorita: ; genus Paraleleupidia: - - - - - - - ; Even if the opinion expressed by MORVAN (1994) that “there is no reason to divide the genus Leleupidia” — which means that Coladisia (and perhaps also Gun- vorita) should be simply included in Leleupidia — should prove right for the presumably primitive stock of Colasidia, probably this does not apply to the group of advanced species mentioned above. Perhaps, this species-group could be simply separated from Colasidia to accomodate the mentioned relationships. However, this should be left to a future revisor who must have a reasonable knowledge of the African and Oriental Leleupidiini. Contrary to the genus Colasidia, the genus Gunvorita forms a rather homo- genous unit, likewise regarding external morphological characters as characters of the d and ® genitalia. That is the reason why phylogenetic relations within the genus are not easily settled. Even when a character evolution comparable to that in Colasidia is postulated, the distribution of character states throughout the species is confusing. To give an example: assumed that absence of the crotchet at the apex of the aedeagus, posteriorly rounded or ovalish head, and large eyes are plesiomorphic cha- racter states, these cannot be reasonably correlated, because of the five known species that lack the crotchet angusticeps has very small eyes, and punctipennis has a markedly triangular head. Hence, we must assume convergent origin in many of the characters mentioned, though without having any opportunity to decide between synapomorpy of characters or convergent origin. This heterobathmic character distribution is evident also in several other character states. Hence, it ıs likewise impossible to correlate the phylogenetic relations of the species with their distribution pattern. Actually, although some apparently very THE GENUS GUNVORITA 305 Fic 70. Supposed migration routes of the Oriental Leleupidiini. Different scenario’s of the biogeographic history of genus Gunvorita are described in figs 70 a and b (see text). Immigration routes into the Oriental region: > ; migration routes within the Oriental and Australian regions: — — — — — > . Abbreviations: C: Colasidia; G: Gunvorita; CG: common ancestral stock of Colasidia + Gunvorita; P: Paraleleupidia. closely related species possess adjacent or even overlapping ranges, other — likewise related — species may have quite different distributions. As a consequence, at present it is impossible to fix the most primitive species of Gunvorita and nothing can be said about evolution and historical biogeography within the genus. 306 MARTIN BAEHR HENNIG (1966) expressed and BRUNDIN (1966) firstly adopted the opinion that plesiomorphy of a taxon and place or origin are commonly correlated, and that the most apomorphic taxa are therefore usually found at the margins of the range of the supra- specific taxon, no difference whether speciation is believed to have been caused only by vicariance or also by some migration. If this opinion is true (and likely it is true at least for flightless, endogeous insects of low vagility as Leleupidiini are), then the genus Colasidia should have evolved on the northern part of the drifting Sundaland that firstly came into contact with the Asian mainland and which today forms the Malayan Peninsula. Although this area reached the mainland first, members of the genus Colasidia apparently did not cross the western and northern boundaries of the former Sundaland, even though these northernmost parts where later firmly attached to the Asian mainland. Or, at least, if some did so, there are no remnants left in these areas, presumably caused by extinction. However, stocks of Colasidia did extend over the whole area of Sunda- land (whether it was a continuous landmass or an insular belt), they diversified on Suma- tra and Borneo, crossed the eastern boundaries of Sundaland and the Wallace line into the Australian region (and plate), and colonized New Guinea and northeastern Australia, where some taxonomic radiation occurred BAEHR 1987, 1991, DARLINGTON 1971). Certainly, species of Colasidia and Leleupidiini in general ought to be very poor dispersers, because they are flightless — perhaps already since a long time — and because they live semi-endogeous. Then, why they where able to disperse so widely to the south and southeast in the course of which dispersal they certainly did some island hopping and crossed rather important waterbodies, but apparently were not able to cross the borders of the former Sundaland to the north and west to colonize the whole of tropical South Asia? Or, if they did so, why they were extinct in the northern part of its areal, but not in the southern part? Unfortunately the phylogenetic relations of Colasidia and Gunvorita are still unsettled, although there is some reason to believe that Gunvorita in certain respects of external and genitalic morphology is more advanced than the apparently most primitive members of Colasidia, though less advanced than the highly evolved stock of Colasidia that ranges mainly over Sumatra, Borneo, and the Papuan-Australian region. With respect to the unsettled phylogenetic relations, therefore, two biogeographic scenario’ s are conceivable: e either Gunvorita evolved independently from Coladisia, perhaps from a common ancestor, which implies that Colasidia is a monophyletic unit; e or Gunvorita is more closely related to certain of the more advanced members of Colasidia, and then Colasidia would be paraphyletic. In the first scenario Gunvorita and Colasidia could have reached their present ranges by independent drift on different terranes of the drifting Sundaland that attached separately and at different times to mainland Asia. If that would prove true, then colonization of the Oriental region by Leleupidiini occurred threefold and presumably also at three different times (Fig. 70a). This scenario would also explain the apparent gap between the present ranges of Colasidia and Gunvorita in Burma and western Thailand. THE GENUS GUNVORITA 307 In the second scenario the common ancestor of Colasidia and Gunvorita arrived on the northernmost part of drifting Sundaland (present Malayan Peninsula) at its pre- sent position. After the fusion of this part with the Asian mainland some evolution took place and the ancestral stock of Gunvorita reached its present northern range by range spreading through Burma, but further evolution of Gunvorita likewise occurred in its present pre-Himalayan range (Fig. 70b). In this scenario, however, the apparent distri- bution gap between both genera can be only explained by extinction of the required intermediate Burmese populations — all under the provisio that the apparent distribution gap of Leleupidiini in Burma really exists and is not caused by unsatisfactory exploration. Unfortunately, a final decision is not yet possible in view of the present unsuffi- cient knowledge of geological history, phylogenetic relations, and chorology of the species. In both scenario’s, however, New Guinea and Australia would have been colo- nized rather recently from the north, which implicates for Australia that old Gondwanan faunal elements in the Australian Region may have been indigenous for long periods (as most of those elements have been), but also may be very recent immigrants from the north. In any case, Leleupidiini must be a very old group that evolved before the final breakup of the Gondwanan continent — at least before the final separation of Africa, India, and the terranes of “Sundaland”. As a consequence, the origin of Leleupidiini should be pre-Cretaceous. This general scenario proposed for Leleupidiini finds some support by a rather similar scenario of the biogeographical history in thegenus Cryptocephalomorpha of the carabid subfamily Pseudomorphinae (BAEHR 1997). In this genus, the apparently most primitive species occurs in the northern part of South Africa (Natal), more advanced species live in Malaysia, southern Thailand, Sumatra and Java, and the most advanced species groups occur either on Borneo and the Philippines, either in New Guinea, New Britain, and northernmost Australia, respectively. Apparently this genus likewise originated in Africa, the original stocks of the Asian species presumably arrived drifting on terranes of “Sundaland”, rather little evolved species persist in the area of former “Sundaland”, and the most advanced species transgressed the boundaries of this area and colonized the Australian region (and plate), where today the most highly evolved species live. Similarly to the genus Colasidia, Cryptocephalomorpha did not spread much to the north and east on the Asian mainland, but on the mainland is restricted to the southern parts of Malaysia and Thailand. Therefore, in Cryptocephalo- morpha presumably the same — still unknown — mechanisms prevented further sprea- ding on the mainland as they did in Colasidia. As a conclusion, the biogeographical pattern of drifting on terranes from the African component of Gondwanaland to mainland Asia, subsequent range spreading and evolution in the South Asian insular belt, and later colonization of the Australian region may prove to have been a regular and even fairly common event during the colonization of the Asian-Australian regions. 308 MARTIN BAEHR Fics 21-24 Entire view. 21. Gunvorita elegans Landin; 22. G. martensi Casale; 23. G. laeviceps sp. n.; 24. G. inermis sp. n. Lengths: 4.45 mm: 4.2 mm: 4.0 mm; 3.6 mm. THE GENUS GUNVORITA 309 Fics 25-28 Entire view. 25. Gunvorita indica Darlington; 26. G. ovaliceps sp. n.; 27. G. nepalensis sp. n.; 28. G. hamifera sp. n. Lengths: 5.65 mm; 5.05 mm; 4.75 mm; 4.35 mm. 310 MARTIN BAEHR Fics 29-32 Entire view. 29. Gunvorita schawalleri sp. n.; 30. G. smetanai sp. n.; 31. G. uncinata sp. n.; 32. G. besucheti sp. n. Lengths: 4.15 mm; 4.95 mm; 4.5 mm; 4.8 mm. THE GENUS GUNVORITA 311 Fics 33-36 Entire view. 33. Gunvorita minor sp. n.; 34. G. punctipennis sp. n.; 35. G. depressipennis sp. n.: 36. G. angusticeps sp. n. Lengths: 4.0 mm; 3.8 mm; 4.5 mm; 4.7 mm. 312 MARTIN BAEHR Fics 37-44 Head. 37. Gunvorita elegans Landin; 38. G. martensi Casale; 39. G. laeviceps sp. n.; G. inermis sp. n.; 41. G. indica Darlington; 42. G. ovaliceps Sp. n.; 43. G. nepalensis sp. n.; 44. G. hamifera sp. n. All figures to scale. THE GENUS GUNVORITA 313 Fics 45-52 Head. 45. Gunvorita schawalleri sp. n.; 46. G. smetanai sp. n.; 47. G. uncinata sp. n.: 48. G. besucheti sp. n.; 49. G. minor sp. n.; 50. G. punctipennis sp. n.; 51. G. depressipennis sp. n.; 52. G. angusticeps sp. n. All figures to scale. 314 MARTIN BAEHR Fics 53-60 Prothorax. 53. Gunvorita elegans Landin; 54. G. martensi Casale; 55. G. laeviceps sp. n.; 56. G. inermis sp. n.; 57. G. indica Darlington; 58. G. ovaliceps sp. n.; 59. G. nepalensis sp. n.; 60. G. hamifera sp. n. All figures to scale. THE GENUS GUNVORITA 315 FIGs 61-68 Prothorax. 61. Gunvorita schawalleri sp. n.; 62. G. smetanai sp. n.; 63. G. uncinata sp. n.; 64. G. besucheti sp. n.; 65. G. minor sp. n.; 66. G. punctipennis sp. n.; 67. G. depressipennis sp. n.; 68. G. angusticeps sp. n. All figures to scale. 316 ACKNOWLEDGEMENTS MARTIN BAEHR My thanks are due to Dr I. Lòbl, Genève, for kindly submitting the bulk of the mentioned material, Dr W. Schawaller, Stuttgart, for providing material collected by Prof. J. Martens and by Martens and himself in Nepal. I also thank Mr S. Hine, London and Dr D. Kovac, Frankfurt/M. for kindly loaning the types of Gunvorita indica and G. martensi. I am further indebted to Prof. Dr G.E. Ball, Edmonton, for his critical reading of a first draft of the biogeographic section of the paper and for some very interesting and helful comments. ALPHABETIC CHECKLIST OF THE LELEUPIDIINI FROM THE ORIENTAL AND AUSTRALIAN REGIONS Colasidia angusticollis Baehr, 1988 Colasidia atra Baehr, 1997 Colasidia attenuata Baehr, 1997 Colasidia borneensis Baehr, 1997 Colasidia brevicornis Baehr, 1988 Colasidia burckhardti Baehr, 1997 Colasidia convexior Baehr, 1993 Colasidia denticollis Baehr, 1997 Colasidia depressa Berhr, 1997 Colasidia gerardi Perrault, 1982 Colasidia globiceps Baehr, 1991 Colasidia helvetorum Baehr, 1997 Colasidia kokodae Baehr, 1991 Colasidia lagadiga (Morvan, 1994) Colasidia laticeps Baehr, 1997 Colasidia loebli Baehr, 1997 Colasidia lustrans Baehr, 1991 Colasidia macrops Baehr, 1990 Colasidia madang Darlington, 1971 Colasidia malayica Basilewsky, 1954 Colasidia mateui Baehr, 1997 Coladisia monteithi Baehr, 1987 Colasidia oviceps Baehr, 1997 Colasidia papua Darlington, 1971 Colasidia pumila Baehr, 1990 Colasidia riedeli Baehr 1990 Colasidia rougemonti (Morvan, 1994) Colasidia similis Baehr 1997 Colasidia taylori Baehr 1988 Coladisia triangularis Baehr, 1997 Gunvorita angusticeps Sp. n. Gunvorita besucheti sp. n. Gunvorita depressipennis sp. n. Gunvorita elegans Landin, 1955 Gunvorita hamifera sp. n. Gunvorita indica Darlington, 1971 Gunvorita inermis Sp. n. Gunvorita laeviceps sp. n. Gunvorita martensi Casale, 1985 Gunvorita minor sp. n. Sarawak (Borneo) Sarawak (Borneo) Malaysia Sabah (Borneo) Sarawak (Borneo) Sabah (Borneo) Sumatra Sumatra Malaysia Sabah (Borneo) Sumatra Sumatra Papua New Guinea Malaysia Sabah (Borneo) Malaysia Sumatra Sarawak (Borneo) Papua New Guinea Malaysia Sabah (Borneo) Queensland (Australia) Malaysia Papua New Guinea Sarawak (Borneo) Sarawak (Borneo) Malaysia Sumatra Sarawak (Borneo) Malaysia East Nepal Northeast India Northeast India East Nepal, Sikkim, Northeast India East Nepal Northeast India Northeast India Northeast India East Nepal Northeast India THE GENUS GUNVORITA 317 Gunvorita nepalensis sp. n. Central Nepal Gunvorita ovaliceps sp. n. Northeast India Gunvorita punctipennis Sp. n. East Nepal Gunvorita schawalleri sp. n. East Nepal Gunvorita smetanai Sp. n. Central Nepal Gunvorita uncinata Sp. n. East Nepal Paraleleupidia besucheti Mateu, 1981 South India Paraleleupidia linearis Baehr, 1990 South India Paraleleupidia loebli Mateu, 1981 South India REFERENCES AUDLEY-CHARLES, M. G. 1987. Dispersal of Gondwanaland: relevance to evolution of the Angiosperms, 5-25. In: Biogeographical evolution of the Malay Archipelago (Whitmore, T.C., ed.). Clarendon Press, Oxford. BAEHR, M. 1987. Revision of the Australian Zuphiinae 2. Colasidia monteithi sp. nov. from North Queensland, first record of the tribe Leleupidiini in Australia (Insecta: Coleoptera: Carabidae). Memoirs of the Quensland Museum 25: 135-140. BAEHR, M. 1988. Three new Leleupidiini from Sarawak (Coleoptera, Carabidae, Zuphiinae). Mitteilungen der Miinchner Entomologischen Gesellschaft 78: 115-123. BAEHR, M. 1990. Four new species of Leleupidiini from the Oriental Region (Coleoptera, Cara- bidae, Zuphiinae). 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REVUE SUISSE DE ZOOLOGIE 105 (2): 319-329; juin 1998 Redescription of Hymenolepis hoploporus Dollfus, 1951, with the erection of the new genus Dollfusilepis (Cestoda, Hymenolepididae) Gergana P. VASILEVA, Boyko B. GEORGIEV & Todor GENOV Central Laboratory of General Ecology, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria: e-mail: bbg@bbgcict.acad.bg Redescription of Hymenolepis hoploporus Dollfus, 1951, with the erec- tion of the new genus Dollfusilepis (Cestoda, Hymenolepididae). - The monotypic genus Dollfusilepis is erected for Hymenolepis hoploporus Dollfus, 1951, until now known only from Podiceps cristatus (Aves, Podicipedidae) in Morocco. D. hoploporus comb. n. is recorded from the same host species in Bulgaria and Switzerland (new geographical records). The cestodes identified by JOYEUX & BAER (1950) as Hymenolepis capillaris are recognized as D. hoploporus. The species is redescribed based on specimens from Bulgaria, Switzerland and syntypes from Morocco. Among the genera of the family Hymenolepididae, the new genus Dollfusilepis is unique by the peculiar structure of the genital apparatus presented by a protrusible poral part of the cirrus-sac armed with a crown of refractive, baton-like spines. Key-words: Dollfusilepis gen. n. - Dollfusilepis hoploporus comb. n. - Hymenolepididae - Cestoda - Podiceps cristatus - Bulgaria - Switzerland - Morocco. INTRODUCTION Hymenolepis hoploporus Dollfus, 1951 was described from the intestine of Podiceps cristatus (L.) in Morocco. Up to now, there were no other records of this species. Recently, it was found in three specimens of P. cristatus during faunistic studies on cestodes from grebes from the Bulgarian coast of the Black Sea. The material consisted only of strobilar fragments and, therefore, for the purposes of more reliable identification, they were compared with types from the collection of the Muséum National d’ Histoire Naturelle, Paris. On the other hand, in the course of taxonomic revision of hymenolepidid cestodes from Palaearctic grebes, we reexamined specimens of JOYEUX & BAER (1950) from the collection of the Muséum d’histoire naturelle in Geneva. The cestodes identified as Hymenolepis capillaris (Rudolphi, 1810) were found to closely resemble H. hoploporus. Manuscript accepted 06.10.1997 320 GERGANA P. VASILEVA, BOYKO B. GEORGIEV & TODOR GENOV The aim of the present paper 1s to redescribe Hymenolepis hoploporus on the basis of specimens from Bulgaria, Morocco and Switzerland and to clarify its generic allocation. On this basis, Dollfusilepis gen. n. is proposed for it. MATERIALS AND METHODS Specimens of Dollfusilepis hoploporus were collected from the small intestine of three birds of Podiceps cristatus. The grebes were captured at the village of Krapec, on northern part of the Bulgarian Black Sea coast. Cestodes were isolated from intestines alive, relaxed in tap water, fixed in 10% formalin solution and preserved in 70% ethanol. Specimens were stained in iron acetocarmine, dehydrated in alcoholic series, cleared in eugenol and mounted in Canada balsam. The material examined from Bulgaria consists of about 30 fragments of strobila in different stages of development, stained and mounted in Canada balsam (total 10 slides); scoleces were not found. Voucher specimens (2 slides) are deposited in the Muséum d’histoire naturelle, Geneva, Nos 23,629 INVE and 23,630 INVE. Comparative material: the following syntypes and voucher specimens were studied: — from the collection R.-Ph. Dollfus, Muséum National d’Histoire Naturelle, Paris, syntypes, from intestine of Podiceps cristatus, 16 October 1926, Rabat Sale, Morocco (2 slides, whole mounts): 1 slide containing stained fragments of strobila; 1 slide containing a scolex. — from the collection of the Muséum d’histoire naturelle, Geneva, Nos 089/035-037, specimens identified as Hymenolepis capillaris (Rudolphi, 1810) and mentioned by JOYEUX & BAER (1950), from intestine of Podiceps cristatus, Neuchâtel (3 slides, whole mounts): 1 slide containing a stained single specimen and several additional fragments of strobila; 1 slide containing stained strobilar fragments: 1 slide containing scolex. The measurements of the cirrus-sac, the external seminal vesicle and the seminal receptacle were taken from mature proglottides. The metrical and meristic data are given as the range, the mean in parentheses and the number of measurements or counts taken (n). The measurements are given in micrometers unless otherwise stated. DESCRIPTIONS Dollfusilepis gen. n. Strobila protandrous, slender. Scolex round. Rostellar apparatus musculo- glandular. Rostellum armed with single crown of ten hooks. Each rostellar hook consisting of aploparaksoid refractive particle and epiphyseal thickening of handle. Suckers round, unarmed. Proglottides craspedote, wider than long. Inner longitudinal muscle bundles numerous. Genital pores unilateral. Genital ducts dorsal to osmo- regulatory canals. Testes three, arranged in triangle, usually one of them porally to female primordia. External seminal vesicle oval or elliptical. Internal seminal vesicle large, elliptical. Cirrus-sac elongate, reaching mid-line of proglottis, often extending DOLLFUSILEPIS GEN. N. 321 to antiporal osmoregulatory canals. Poral end of cirrus-sac protrusible, provided with basal crown of baton-like refractive spines. Cirrus narrow, unarmed. Accessory sac and stylet lacking. Female glands disposed antiporally. Ovary with three compact lobes. Vitellarium oval, compact. Seminal receptacle voluminous, elliptical. Vagina with funnel-shaped, thick-walled, muscular copulatory part, surrounded with cellular sleeve and thin, tubular conductive part. Uterus sac-like, situated anteriorly and dorsally to female glands. Specific parasites of Podiceps cristatus (Podicipedidae). Palaearctic. Type-species: Hymenolepis hoploporus Dollfus, 1951. Dollfusilepis hoploporus (Dollfus, 1951) comb. n. (Figs 1-16) Hymenolepis (Weinlandia) hoploporus DOLLFUS, 1951 Dubininolepis hoploporus (Dollfus, 1951) YAMAGUTI 1959 Variolepis hoploporus (Dollfus, 1951) SCHMIDT 1986 Hymenoleopis capillaris (Rudolphi, 1810), JOYEUX & BAER (1950). DESCRIPTION OF SPECIMENS FROM BULGARIA (Figs 1-6: for some measurements see Table I): Strobila slender, band-like, with maximum width at post-mature pro- glottides. Proglottides (Figs 1-3) craspedote, always wider than long. Inner longitu- dinal muscle bundles more than 40. Genital pores unilateral, situated at about middle of lateral proglottis margin. Genital atrium (Figs 4-5) deep, funnel-shaped, thick- walled; divides into two separate canals: male and female. Atrium surrounded with intensely stained cells; when the cirrus is evaginated, atrium forming short genital papilla. Ventral and dorsal osmoregulatory canals without transverse anastomoses. Diameter of ventral osmoregulatory canals 18-36 (26, n=10), diameter of dorsal osmoregulatory canals 3-5 (5, n=10). Genital ducts dorsal to osmoregulatory canals. Strobila protandrous. Testes (Fig. 1) three, compact, oval, situated in triangle, one poral, two antiporal to vitellarium. External seminal vesicle elliptical or oval, situated dorsally to female glands, near to antiporal osmoregulatory canals. Cirrus-sac (Figs 1-2) thin-walled, highly elongate, crossing mid-line of proglottis and often extending to antiporal osmoregulatory canals. Intensely stained cells surrounding ductus ejaculatorius. Internal seminal vesicle very long, fills up almost 2/3 of cirrus- sac. Poral end of cirrus-sac (Figs 4-6) protrusible, cylindrical when everted, forming wide ductus when withdrawn; provided with basal crown of baton-like refractive spines; number of spines 30-35 (34, n=7), their length 6-7 (7, n=5). Evaginated cirrus (Fig. 6) thin, almost cylindrical, unarmed. Female genital organs (Fig. 2) disposed antiporally. Ovary consisting of three compact lobes. Vitellarium compact, elliptical or oval, situated postero-ventrally to ovary. Seminal receptacle (Fig. 2) voluminous, transversely elongate, situated ven- trally to cirrus-sac and dorsally to ovary. Vagina (Figs 4-5) opens on protrusible base of female atrial canal, often forming tiny papilla; vaginal orifice thick-walled, infundi- bular, with diameter 8-10 (8, n=10). Position of female atrial canal and vaginal orifice 322 GERGANA P. VASILEVA, BOYKO B. GEORGIEV & TODOR GENOV TABLE I Metrical and meristic data for Dollfusilepis hoploporus (Dollfus) comb. n. Locality Bulgaria Morocco Morocco Neuchâtel Source Present study Dollfus Present study Present study (1951) Range Mean n Range Range Mean n Range Mean n Strobila: length (mm) = Sii 70 40 — | 26.0 | width (mm) - - = I 0.6 SET] 0.3 ei Scolex: length — — = - - — = 188 Sp width - — = 155 170 = al 142 seg 4 Suckers: diameter _ - = - - - = 44-52 49 4 Rostellum: length - — = - - — = 62 = oath width - — = - - - = 39 — | Rostellar sheath: length - - = - - — = 104 Set] width - - - - - - = 64 | Rostellar hooks: total length - - = 20-21 20 21 220554 20-21 21 4 length of blade - - = - 10-11 11 4 10-11 1055 Testes: diameter 26-34 28.207 305,25 30523136, 533710 31-39 3720 Cirrus-sac: length 180-232 203 20 170 160-186 173 10 216-238 229 20 width 13-23 205520 20 18-232 2222210 23-31 27720 Ext. seminal vesicle: length 31-64 41 20 - ANC ST IO 62-129 65 15 width 21-39 26 20 - 31-46 36 10 44-59 3315 Vitellarium: length 36-41 380215 - 57-77 66 10 41-46 45 10 width 18-23 20815 - 41-46 43 10 28-34 Sik 10) Seminal receptacle: length 9022 12100615 - 90 — | 129-162 147 10 width 31-41 33.0215 - 34 ha 39-52 43 10 to male pore not constant, usually posterior, often anterior or lateral. Copulatory part of vagina fusiform, thick-walled, muscular, 18-23 (20, n=10) long, surrounded by cellular sleeve. Conductive part tubular, slender, straight or slightly convoluted. Developing uterus (Fig. 3) sac-like, transversely elongate, situated dorsally to female glands and osmoregulatory canals; passing beyond osmoregulatory canals. Proglottides with fully-developed uterus and ripe eggs not available. DOLLFUSILEPIS GEN. N. 328 Fics 1-6 Dollfusilepis hoploporus (Dollfus, 1951) gen. n., comb. n., specimens from Bulgaria: 1-2, mature proglottides; 3, proglottis with developing uterus; 4-5, terminal genital ducts; 6, evaginated cirrus. Scale-bars: 1-3 = 100 um; 4 = 50 um; 5-6 = 25 um. 324 GERGANA P. VASILEVA, BOYKO B. GEORGIEV & TODOR GENOV OBSERVATIONS ON SYNTYPES (Figs 7-11; for some measurements see Table 1): Most of the current observations are in agreement with the original description (DOLLFUS 1951) and, therefore, an entire redescription of the syntypes will not be given. The few details presented below extend or correct the original data. The position of the scolex in the slide does not permit us to characterize the structure of the suckers, the rostellum and the rostellar sheath. Rostellar hooks in number 10, with aploparaksoid refractive particle (Fig. 7) with length 16 (n=4); epiphyseal thickening of handle present; distance between handle-tip (of whole hook) and guard-tip 14 (n=4): distance between blade-tip and guard-tip 6-7 (7, n=4). Strobila fragmented. Testes (Fig. 9) arranged in triangle. External seminal vesicle oval. Cirrus-sac (Figs 9-10) crossing mid-line of proglottis; ductus ejaculatorius (Fig. 8) surrounded by intensely stained cells. Crown of baton-like, refractive spines on poral protrusible part of cirrus-sac (Fig. 8). Internal seminal vesicle very large, occupying almost 2/3 of cirrus-sac. Evaginated cirrus (Fig. 8) thin, almost cylindrical, unarmed. Ovary (Fig. 10) disposed antiporally, consisting of three compact lobes. Vitellarium well-visible, oval or elliptical, compact, postovarian. Seminal receptacle very large, situated postero-ventrally to cirrus-sac, dorsally to ovary. Young uterus (Fig. 10) sac-like, with poorly visible walls, situated anteriorly and dorsally to ovary. Developing uterus transversely elongate, crossing osmoregulatory canals dorsally. Proglottides with fully developed uterus and eggs not available. OBSERVATION ON SPECIMENS FROM SWITZERLAND (Figs 12-16; for some measurements see Table I): Few details, mainly referring to the morphology of the scolex (lacking in the Bulgarian material and in poor condition in the syntypes), are presented below. Scolex (Fig. 12) round, with maximum width at level of suckers. Suckers round, unarmed. Rostellum ovoid, with well-developed musculature; intensely stained cells situated in it. Rostellar sheath with weakly-developed musculature of walls, usually passing beyond posterior margin of suckers. Intensely stained cells present in rostellar sheath. Rostellar hooks (Fig. 13) with epiphyseal thickenings of handle; length of aploparaksoid refractive part 16-17 (n=2); distance between handle-tip (of whole hook) and guard-tip 14 (n=2); distance between blade-tip and guard-tip 8 (n=2). Diameter of ventral osmoregulatory canals 8-18 (12, n=10), diameter of dorsal osmoregulatory canals 3-8 (5, n=10). Genital atrium (Fig. 16) deep, antiporally divided into two separate canals. Cirrus-sac (Figs 14-15) thin-walled, highly elongate, crossing mid-line of proglottis and often extending to antiporal osmoregulatory canals. Poral part of cirrus-sac (Fig. 16) protrusible, armed with crown of baton-like spines. Cirrus thin, unarmed. Female genital glands disposed antiporally. Copulatory part of vagina (Fig. 16) with strong circular musculature. Proglottides with fully- developed uterus and ripe eggs not available. COMMENTS: Initially, these cestodes were recorded from Podiceps cristatus from the Lake of Neuchatel and identified as Hymenolepis capillaris (Rudolphi, 1810) by JOYEUX & BAER (1950). Their conclusion was based mainly on the morphology of the DOLLFUSILEPIS GEN. N. 805 i Fics 7-11 Dollfusilepis hoploporus (Dollfus, 1951) gen. n., comb. n., syntype specimens: 7, rostellar hooks; 8, evaginated cirrus; 9, young mature proglottis; 10, mature proglottis with early stage of uterine development; 11, terminal genital ducts. Scale-bars: 7-8 = 20 um; 9 = 150 um; 10 = 100 um; 11 = 50 pm. 326 GERGANA P. VASILEVA, BOYKO B. GEORGIEV & TODOR GENOV 16 UE Fics 12-16 Dollfusilepis hoploporus (Dollfus, 1951) gen. n., comb. n., specimens from Switzerland: 12, scolex; 13, rostellar hooks; 14, mature proglottis; 15, proglottis with developing uterus; 16, terminal genital ducts. Scale-bars: 12, 14 = 100 um; 13 = 20 um; 15 = 150 um; 16 = 50 um. DOLLFUSILEPIS GEN. N. 524] rostellar hooks as described and figured by KRABBE (1869) (PI. VII, Fig. 179). JOYEUX & BAER (1950) presented brief data about the morphology of the proglottides but did not describe the structure of the copulatory apparatus. By courtesy of Dr B. Neuhaus, we had the opportunity to reexamine the type-material of H. capillaris from the collection of the Naturkunde Museum, Berlin; its redescription, together with redes- criptions of other species of the genus Confluaria Ablasov, will be subject of another publication (Vasileva, in preparation). In spite of the similar shape of the rostellar hooks, the morphology of the genital system of C. capillaris is very different from that of the cestodes from the Lake of Neuchâtel. As seen from the above description, this material should be referred to D. hoploporus. DISCUSSION The comparison of the present results about the morphology of Hymenolepis hoploporus with the hymenolepidid genera as characterized by CZAPLINSKI & VAUCHER (1994) reveals that this species is unique in the peculiar morphology of its male copulatory apparatus, which possesses a protrusible poral part of the cirrus-sac armed with a crown of baton-like spines. The morphology of H. hoploporus fits to a group of avian hymenolepidid genera characterized by three testes, a rostellum armed with a single crown of 10 hooks and lacking an accessory sac and a cirrus-stylet. According to CZAPLINSKI (in CZAPLINSKI & VAUCHER 1994), this group includes Nadejdolepis Spasskii & Spasskaya, 1954, Microsomacanthus Lopez-Neyra, 1942, Matiarensis Dixit & Capoor, 1986. Parafimbriaria Voge & Read, 1954, Confluaria Ablasov in Spasskaya, 1966 and Wardium Mayhew, 1925. Hymenolepis hoploporus cannot be included in any of these genera. Its rostellar hooks are with an aploparaksoid refractive particle and possess well differentiated epiphyseal thickening of the handle whilst the species of the genera Nadejdolepis and Microsomacanthus have nitidoid and diorchoid hooks, respectively. The species belonging to Wardium also have 10 aploparaksoid hooks but without epihpyseal thickenings (CZAPLINSKI, in CZAPLINSKI & VAUCHER 1994). The genus Matiarensis is characterized by the presence of 4 accessory apical suckers. The species of Parafimbriaria lack external segmentation of the strobila. The species of Confluaria, especially C. capillaris, most closely resemble H. hoploporus. They are also parasites of grebes and also have ten aploparaksoid hooks with epiphyseal thickenings, a tri-lobed ovary and a sac-like uterus. However, they are characterized by a long, cylindrical or claviform cirrus, armed usually with different types of spines: sparse, rose-thorn shaped spines at distal part of cirrus and dense, minute, needle-shaped spines at its basal part. H. hoploporus has a different structure of the copulatory apparatus. The genital atrium of the species of Confluaria is simple, whilst in H. hoploporus it is thick-walled and antiporally divided into two separate canals, male and female. The cirrus-sac of Confluaria spp. rarely crosses the mid-line of the proglottis, whilst that of H. hoploporus is very long, often extending to the antiporal osmoregulatory canals. These morphological peculiarities, in addition to 328 GERGANA P. VASILEVA, BOYKO B. GEORGIEV & TODOR GENOV the armed protrusible poral part of the cirrus-sac, are not observed in species of Confluaria. Furthermore, H. hoploporus possesses numerous inner longitudinal muscle bundles; in contrast, Confluaria Spp. are characterized with only 8 bundles (CZAPLINSKI, in CZAPLINSKI & VAUCHER 1994). In addition, the genera Microsomacanthus and Wardium as defined by CZAPLINSKI (in CZAPLINSKI & VAUCHER 1994) seem to be heterogeneous groups. Therefore, the new genus has also to be distinguished from some of the genera believed to be their synonyms, especially those parasitizing aquatic birds and possessing, to some extent, similar morphological characters. The genus Anserilepis Spasskii & Tolkacheva, 1965 (type-species Hymenolepis barrowensis Schiller, 1952 from Anseriformes in Alaska and Siberia), as characterized by SPASSKI & TOLKACHEVA (1965), has 10 diorchoid hooks, female gonads situated antiporally to testes, and a cirrus armed with spines of three different shapes. Dubininolepis Spasskii & Spasskaya, 1954 (type-species Hymenolepis fuhrmanni Skrjabin and Matevosyan, 1942 from Gaviiformes and Podicipediformes in North America) is characterized by 8 inner longitudinal muscle bundles, a highly elongate rostellum, a deep rostellar pouch and rostellar hooks without epiphyseal thickenings (SPASSKAYA 1966). The species of Echinatrium Spasskii & Yurpalova, 1965 (type-species E. skriabini Spasskii & Yurpalova, 1965 from Anseriformes in Chukotka) possess 8 inner longitudinal muscle bundles and very deep genital atria (occupying almost one- fifth of the proglottis width) with spinose bases (SPASSKAYA 1966; TOLKACHEVA 1971). The genus Laricanthus Spasskii, 1963 (type-species L. lateralis (Mayhew, 1925) from Laridae) is characterized by 10 diorchoid rostellar hooks, a fan-shaped, multilobate ovary, a pyriform thick-walled cirrus-sac and highly muscular vagina (SPASSKAYA 1966). Oshmarinolepis Spasskii & Spasskaya, 1954 (type-species O. microcephala (Rudolphi, 1919) from Ardeidae) has testes situated in both median and lateral fields of the proglottis, 8 inner longitudinal muscle bundles and a strong, spherical genital atrium which may form a prominent genital papilla (SPASSKAYA 1966). Chelacanthus Yamaguti, 1959 (type-species C. parviceps (von Linstow, 1872) from birds of the tribe Mergini, Anatidae) possesses an armed vagina and a rosette- shaped ovary (YAMAGUTI 1959). Decacanthus Yamaguti, 1959 (type-species D. arcticus (Schiller, 1955) from Anseriformes in Alaska) has a rather short cirrus-sac which does not reach the median line of the proglottis, an ovary with “numerous digitiform lobules” and a vagina forming a characteristic loop (YAMAGUTI 1959). Lobatolepis Yamaguti, 1959 (type-species L. lobulata (Mayhew, 1925) from Podi- cipedidae in North America) has a rostellum provided with knob-like enlargement with 8-11 deep marginal lobes, each lobe bearing a rose-thorn rostellar hook, a rather short cirrus-sac and a transversely elongate ovary (YAMAGUTI 1959). None of the characters mentioned occurs in H. hoploporus and it cannot be placed, therefore, in any of the above discussed genera. CZAPLINSKI (1967) redescribed Wardoides nyrocae (Yamaguti, 1935) from Cygnus olor (Gmelin) in Poland. This species is characterized by a thin cirrus with an unarmed base. According to CZAPLINSKI (1967), “the bottom of the genital atrium is armed with very small hair-like hooks; when the genital atrium is protruded... has the DOLLFUSILEPIS GEN. N. 329 shape of a surrounded hat...” (see Fig. 3 of CZAPLINSKI 1967). This structure of the genital atrium, mentioned also by YAMAGUTI (1935) in the original description of this species, is similar to the male copulatory apparatus of H. hoploporus. However, the armed, protrusible “hat-like” papilla of W. nyrocae is formed by the bottom of the genital atrium; in contrast, in H. hoploporus the poral part of the cirrus-sac is armed and forms a similar protrusible papilla. In the original description of H. hoploporus, DoLLFUS (1951) also regarded baton-like spines as an armament of the genital atrium. On the basis of the above comparisons, the new monotypic genus Dollfusilepis is proposed for H. hoploporus. Up to now, D. hoploporus has been recorded only from P. cristatus in three regions of the Palaearctic. Therefore, it can be supposed to be a specific parasite of this grebe species. ACKNOWLEDGEMENTS We are grateful to Dr M.-C. Durette-Desset and Mrs R. Tcheprakoff (Paris), Dr B. Neuhaus (Berlin) and Dr C. Vaucher (Geneva) for the loan of specimens. Dr C. Vaucher (Geneva) and Dr R. A. Bray (London) kindly suggested some impro- vements of the manuscript. This investigation was completed with the financial support of the National Scientific Research Foundation of the Republic of Bulgaria, Grant B-404/1994. REFERENCES CZAPLINSKI, B. 1967. The anatomy of Wardoides nyrocae (Yamaguti, 1935) Spassky, 1962 (Cestoda) from Cygnus olor (Gm.). Acta Parasitologica Polonica 15: 113-121. CZAPLINSKI, B. & C. VAUCHER 1994. The family Hymenolepididae Ariola, 1899. In: Keys to the cestode parasites of vertebrates (L.F. Khalil, A. Jones & R.A. Bray, eds). CAB International, Wallingford (UK): 595-663 pp. DOLLFUS R. PH. 1951. Quelques trematodes, cestodes et acanthocephales. Miscellanea helmin- thologica maroccana. Archives de l’Institut Pasteur de Maroc 4: 104-229. JOYEUX, CH. & J.G. BAER 1950. Sur quelques espèces nouvelles ou peu connues du genre Hymenolepis Weinland, 1858. Bulletin de la Société neuchâteloise des Sciences natu- relles 73: 51-70. KRABBE, H. 1869. Bidrag til Kundskab om Fuglenes Baendelorme. Kjobenhavn Danske vidensk. Selskab. Skr., 5. Baekhe, naturvidenskabelig og mathematisk Afd. 8: 249-363. SPASSKAYA, L.P. 1966. [Cestodes of birds in the USSR. Hymenolepididae.| ‘Nauka’, Moscow, 698 pp. (in Russian). SPASSKII, A.A. & L.M. TOLKACHEVA 1965. [Anserilepis nov. gen. (Cyclophyllidea, Hyme- nolepididae), a new cestode genus from Anseriformes.] Trudy Gel’mintologischeskoy Laboratorii, Akademiya Nauk SSSR 15: 151-155 (in Russian). TOLKACHEVA, L.M. 1971. [New cestode species, Echinatrium clanguli nov. sp. and Micro- somacanthus strictophallus nov. sp. (Hymenolepididae) from Anseriformes.] Jn: Sbornik Rabot po Gel’mintologii Posvyashchen 90-letiyu so Dnya Rozhdeniya Aka- demika K.I. Skryabina. ‘Kolos’, Moscow: 406-410 pp. (in Russian). YAMAGUTI, S. 1935. Studies on the helminth fauna of Japan. Part 6. Cestodes of birds, I. Japanese Journal of Zoology 6: 183-232. YAMAGUTI, S. 1959. Systema Helminthum. Vol. II. The cestodes of vertebrates. Interscience Publishers, New York: 860 pp. à REN x É ANT sà i ie ser ters nr on LA n RA = REVUE SUISSE DE ZOOLOGIE 105 (2): 331-338; juin 1998 A new Cyprinodont species with a uniquely-colored female, Aphyosemion hera n. sp. (Cyprinodontiformes, Pisces), from northwestern Gabon Jean H. HUBER Laboratoire d'Ichtyologie, Museum national d'Histoire naturelle, 43, rue Cuvier, 75231 Paris Cedex 05, France. A new Cyprinodont species with a uniquely-colored female, Aphyo- semion hera n. sp. (Cyprinodontiformes, Pisces), from northwestern Gabon.- A new species of Aphyosemion is described from a single locality about 45 km northeast from Lambaréné, within the primary forest of northwestern Gabon. Together with some distinctive characters (the opposite insertion of dorsal fin to anal fin, a non mood-dependant broad dark longitudinal band), present in both sexes, the new species exhibits, in the female, a bright apple yellow to gold yellow coloration on belly and strong red pigments on sides and fins. On the contrary, females of the genus, encompassing over 120 species, are usually drab light brown with few markings. The systematic position is discussed: it is rather obscure since it does not fall into one of the eleven subgenera, as presently diagnosed. Key-words: Pisces - Cyprinodont - Aphyosemion - new species - N.-W. Gabon. INTRODUCTION The prospection of the species of the West African genus Aphyosemion can be today regarded as satisfactory, numerous expeditions, mainly by enthusiastic aqua- rists, having been organised throughout its range since the late sixties. For example, in Gabon, a small equatorial country of 268 000 square kilometers, more than 210 localities for Aphyosemion have been registered (HUBER 1996). The main obstacle, then, to discover a new species is the inaccessibility of large regions, like the northern Du-Chaillu mountains in central Gabon. Another factor is obviously hazard and this is what happens in the present case: the concerned area has been prospected several times, including by Radda and ourselves in 1976 and only A. gabunense Radda, 1975 was collected. The new species has been discovered by two Swiss experienced killi-hobbyists (i.e. aquarists specialized in oviparous Cyprinodonts), Mr Hermann Romer and Manuscript accepted 26.01.1998 332 JEAN H. HUBER Mr René Krumenacker in July 1996: the fish were rare in their biotope. Further, because of the raised interest, it has been recollected at the same place by two Americans, Mr Peter Tirbak and Mr Vladimir Derugin and one German, Mr Andreas Kliesch, also keen aquarists, in February 1997, even in smaller numbers (only 3 fishes about 15 mm long: Kliesch, comm. pers.). However, the fish were not found despite strong efforts by a third expedition in August 1997 at the same location and in the surroundings. DESCRIPTION OF Aphyosemion hera n. sp. (male, fig. 1 and female, fig. 2: not preserved topotypes). Material: Holotype: MHNG 2590.64, a male of 31.4 mm S.L. and 37.5 mm T.L., R. Krumenacker and H. Romer, coll. July 27, 1996. Paratypes: MHNG 2590.65, a female of 29.2 mm S.L. and 35.4 mm T.L.; MNHN 1997-184, 2 males, MNHN 1997-185, 4 females, USNM 347463, 3 specimens and BMNH 1998.1.21: 1-2-3, 3 specimens, all collected with the holotype; all maintained a few months in aquarium, except the USNM and the BMNH material which is from the F1 aquarium generation. TYPE LOCALITY: Gabon, 45 km northeast of Lambaréné (starting point: the bridge over the Ogooué river in the city) on the road to Bifoun, near Benguié, lower Ogooué basin. Geographical coordinates in degrees and hundredths: 0.47S, 10.32E. At this point, the Ogooué river flows less than 10 km away to the east; the Mbiné river, one of its tributary is about the same distance away to the west. DIAGNOSIS: Aphyosemion hera is a medium-sized species with a strong sexual dimorphism and dichromatism: the female is not subdued, as usual in the genus, but well specifically colored. The new species is besides diagnosed by the opposite insertion of the vertical fins (D/A= +0.4, average), by the asymmetrical pattern of the vertical and caudal fins in male, by the rare, in non annual species, trilobate caudal shape in male and by the deep anal fin in female; in addition, the longitudinal dark mid-band of the young male and the female at all stages is permanent and does not depend on mood, like the one seen in some other species, especially those of the subgenus Kathetys Huber, 1977. Methods of measurements and counting have been detailed in HUBER (1992). Morphomeristic data of the first quoted eight types, the holotype first in bold (after confirmation by radiophotographs): sex= male, female, male, male, female, female, female, female. D= 11, 11,12, 115 11, 11, 12) 1) (mean 11.25; SD 0 PB) APM 12, 12, 12, 11 (mean: 11.88: S.D.: 0:33). D/A= +1, +2, +1, 0, CII 4120" (mean +0.62; S'D.: 0.86). LL= 2842; 2844; 2942: 2744; 28+3: 2844: 2843: 29734 (mean: 28.12+3.12; S.D.: 0.6). Predorsal scales= 14, 15, 13, 13, 14, 13, 14, 14 (mean: 13.75; S.D.: 0.7). Transversal scales (TRAV.)= 9, 9, 10, 10, 10, 9, 10, 9 (mean: 9.6; S.D: 0.5). Circumpeduncular scales (CIR)= 15, 14, 16, 16, 17, 16, 17, 16 (mean: 15.9; S.D.: 0.9). S.L: (An mm)= 31-4; 29.2; 36.7; 28.0; 25:4; 23:83 22.45 204 eles (A0 S.E)= 119, 121, 125, 129, 118, 121, 120; 125) (mean 1223257 DS) (predorsal length)= 62; 64, 60, 61, 63, 65, 64, 69 (mean: 63.7; S.D.: 2.8). P.A. A NEW CYPRINODONT FROM GABON 333 (preanalifiensth) #64 Gk 59% Ole tOls 635962566) (mean 602722NS DEA 0) PAVE (preventralelenth)— 05 49 50N 53552, 52. 52,55 (mean 513725 D 19) Herehe at Anal level= 22, 19, 21, 21, 19, 21, 19, 22 (mean: 20.4; S.D.: 1.2). Height at peduncle level 287 Sinise 14513, A1 135 13) (mean 13-478.D!: 0%) Headilensth= 275285 25: Oe DO) San I MEAN 272$ D 122) Interorbitar= 6.17, WAS Sele 4S (Gean: 4 ES DER) Bye diameter 9) 95 859" 759358, 9i(@means 6:44.57) 2) O27) Snout= 7, 6, 6, 8, 6, 6, 6, 7 (mean: 6.4; S.D.: 0.6). The caudal fin of the dominant male is strongly trilobate, with short streamers on upper and lower tips; the dorsal and, less so, the anal fins bear short streamers; the female is having an unusual deep anal fin and a somewhat pointed dorsal. The D/A deviation has been checked on the radiophotographies of 6 additional specimens with the following results: +1, -1, -1, +1, 0, +1; for the total 14 specimens, the mean value is +0.43 and the standard deviation, 0.80. Vertebrae (abdominal+caudal), on 8 specimens= 13415, 12+15, 11+15, 12+15, 1115, 12+15, 12+15, 12+16 (mean: 27.0; S.D.: 0.71). The hypural plate is fully divided ın the middle, an unusual situation. The frontal scalation is of the G-type, but less regular than usual, one female being F-type. The frontal neuromast pattern is open, like in all Aphyosemion from Gabon: the channels are unusually wide, not protected by fleshy lobes. A few ctenoid scales are available on the old male sides. Colour in life: male, little pigmented on a blue green metallic background; the "shield" pattern (red lines, longitudinal below lip, oblique on operculum) is not conspicuous; only a few red blotches on the upper part of sides and near the basis of the dorsal, ventral and anal fins can be noticed; the middle of the dorsal, pectoral and caudal fins are flamed with red along rays; in addition, two submarginal red bands, the lower being wider, occur on the caudal but not on the dorsal fin; light blue margins are seen on dorsal, pectoral, and caudal fins, but not on ventral and anal fins, which are yellow-green overall, except their basis. Female, strongly pigmented with red spots over upper sides and with red flames on all fins; the dorsal fin and the upper part of the caudal fin is black margined, like the lower lip and the area below the eye: besides, a broad conspicuous dark band is present (also in the juvenile male) from behind the eye (in prolongement of the lower lip black line) until the tail; below this band, the entire belly is colored with a contrasting apple to gold yellow, like all paired and unpaired fins. Colour in alcool: male, with around 20 light big spots on upper sides over a dark brown background; the lower mid-sides are less dark; head, dark with two darker lines on lower lip and somewhat below the first one; dark shield, well marked; dark rays on vertical fins, except a light broad margin at lower caudal and a light edge at its upper part; the dorsal fin bears a streamer and reaches, like anal, the caudal peduncle level. Female, with a broad dark longitudinal band from snout until the caudal peduncle where it ends larger by a blotch; a second dark thiner line, parallel to the lower lip, goes beyond the rear part of eye, underlying it; above the broad band, the region of upper sides is mottled with grey markings, while below it, it is unmarked 334 JEAN H. HUBER and yellow; all fins, except pectorals, irregularly and discontinuously flamed with black, especially on mid-caudal following the band; in both sexes, a dark thin line on back from the A-scale to the dorsal fin insertion. Ecological data (Romer, pers. comm., June 14, 1997): the biotope is similar to that of other Aphyosemion species: a shady primary forest creek ("marigot"); the water was, in summer 1996 i.e. during the long dry season, very shallow, 1-2 m wide and 20 cm deep, with lots of dead leaves; the water was clear, of low conductivity, slightly acid, very similar to rain water as usual for Aphyosemion (a single measure- ment: conductivity= 20uS; pH= 6); at 2 p.m., the water temperature was 21°C. A dozen of specimens only could be caught, whereas the other sympatric species, A. gabunense was much more abundant. In the same locality, TIRBAK et al. (1997) were also able to collect two other Cyprinodonts: Epiplatys sexfasciatus Gill, 1862 and E. singa (Boulenger, 1899), the typical fauna of the area between Bifoun and Lambaréné. Aquarium experience (Romer, pers. comm.): a typical non annual species which prefers dark parts of the tank, furnished with lots of plants; fairly easy to breed, despite unbalanced sex-ratios (first generation in favor of males; second in favor of females); the pair spawns on perlon mops with the standard Z-type position of Aphyo- semion; no aggressivity displayed; incubation time: 20 days at room temperature; first food: Artemia nauplii; growth: relatively slow. Sexual differentiation may appear at 5 months and first breeding at 7. Grell (pers. comm.) reports that the sexual maturity is reached at 7 months and the adult size, at 12 months. Derivatio nominis: hera (the Greek Goddess), an invariable noun in apposition, the name refers to the beauty of the female, probably the most beautiful in Aphyosemion, if not in Cyprinodonts, but this is subjective. DISCUSSION The main morphomeristic characters and the colour pattern of the new species places it undoubtedly within the genus Aphyosemion among the tropical West African Cyprinodonts: average dorsal and anal fin basis (D<16; A<16), dorsal and anal insertion, not too far from each other (D/A< +8), lyre-shaped caudal fin, red pigments on sides and fins, and notably on head (the "shield"), no dark vertical bars; its low meristics (D= 11.3, A= 11.9, D/A= +0.4, on average) can only relate it to two taxa (HUBER 1996: average data): the subgenus Chromaphyosemion Radda, 1971 (D= 11.8; A= 13.6; D/A= 2.4) and the related genus Diapteron Huber and Seegers, 1977 (D= 10.7; A= 11.3; D/A= -2.3). However the new species is distinguished from the Chromaphyosemion components by the absence of the two longitudinal mood- dependant dark bands on sides of both sexes, by the shape of the vertical fins (pointed without long filaments, not trapezoid), by the shape of the caudal fin (without long filaments). It is distinguished from the Diapteron components by the shape of the vertical fins (pointed, not rounded), by the larger size (1 cm larger) and by the com- pletely different colour pattern of male (standard red pigments on blue background, versus the reverse). A NEW CYPRINODONT FROM GABON 335 No other Aphyosemion species combines a low anal fin count and super- imposed vertical fins. The colour pattern of the adult male of A. hera, so important in Aphyosemion speciation (the female chooses the conspecific male among sympatric species!) and systematics, reminds that of the A. gabunense superspecies from the same region. This superspecies encompasses three isomorphic allopatric valid species, separated by their colour patterns and by their caryotypes: A. gabunense (a blue symetrical phase), A. marginatum Radda & Huber, 1977 (a yellow symetrical phase), A. boehmi Radda & Huber, 1977 (a yellow phase, with an asymetrical pattern in the caudal fin). All show, unlike A. hera, strong and regular series of spots on male sides and inner fins, plus a broad symmetrical red margin on dorsal and anal fins; dominant males exhibit long filaments on caudal fin upper and lower streamers; females are grey brown, without any conspicuous dark band (RADDA 1975). The colour pattern of the adult male of A. hera and its body and fin shapes remind also another species with two subspecies (probably valid species): A. pascheni pascheni (Ahl, 1928) and A. p. festivum Amiet, 1987 from the Kribi area in south- western Cameroun, i.e. over 350 km from our locality, with no other population in- between. The festivum male pattern agrees especially with that of hera: red flamed colour pattern of the inner caudal fin, distinctive patterns at dorsal and anal fins (a rare feature in Aphyosemion), however, like the nominal subspecies, the male is heavily pigmented with red longitudinal lines and bears a red submargin at anal; and the female is gray brown with few red and yellow markings and without the charac- teristic longitudinal dark band. Indeed, the five just quoted taxa share with A. hera paucity in biotope and very isolated location: all are known only from their type localities and eventually one or two more places (HUBER 1981; AMIET 1987); on the other hand, caryotype studies (ScHEEL 1990) suggest that pascheni is related to the calliurum superspecies (with australe and ahli), whereas gabunense and its allied are related to the striatum super- species: these two superspecies are similar in fin shape, but not in body depth; they share a large part of their distribution (in Ecuatorial Guinea, Gabon, Congo), although the former is more restricted to near the coast; and they are reported sympatrically in a number of localities (in that case,it appears that the former chooses more open parts of the biotope). Finally, the position of A. hera remains unclear with our present morphological and field knowledge: no direct relationship can seemingly be derived from isomorphic features and the new species appears to be a distinctive morphospecies, a rare case in the genus; an attractive relationship exists with A. pascheni, but the position of the dorsal fin is very different, more advanced and there are several reported cases of colour convergence in Aphyosemion between two species belonging to two different phylogenies; at last, a putative relationship with the sympatric gabunense cannot be rejected, although related species are extremely rarely found in the same biotope in Aphyosemion and in Cyprinodonts in general. JEAN H. HUBER FIGS 1-2 1: male topotype Aphyosemion hera in aquarium. 2: female topotype Aphyosemion hera in aquarium. Photos Wolfgang Grell. A NEW CYPRINODONT FROM GABON DAT Fic. 3 The biotope of A. hera, A. gabunense, E. sexfasciatus, E. singa, a shady creek crossing the road. Photo René Krumenacker. CONCLUSION Low meristics and superimposed vertical fins, a colorful female, the presence of a black longitudinal band in the juvenile male and in the female, characterize Aphyosemion hera which lies apart in the phylogeny of its large genus: is it a primitive species linked with the species occuring in the same region? Is it a relict species, like pascheni, of a formerly well distributed group which suffered considerable extinction? Is it an offshoot of a species living in the nearby yet unknown northern Du Chaillu mountains? It is expected that the DNA techniques will bring clues: no doubt that they will help to sort out the puzzle of speciation that is seen in the genus, especially in primary forest equatorial hilly regions, such as that of A. hera! 338 JEAN H. HUBER It is hoped too that the nice beauties of this species and notably of the female will be appealing to killi-hobbyists for new collections, so that new places are discovered and this so different species better known. ACKNOWLEDGEMENTS I am indebted to MM. H. Romer and R. Krumenacker (Winterthur, Switzer- land) and to Mr P. Tirbak and Mr V. Derugin (Palo Alto, California) and Mr A. Kliesch (Gelsenkirchen, Germany) for having donated preserved material from their aquarium strains or information on their collections and on the ecology, to Mr Wolfgang Grell (Erlach, Germany) for the permission of using his nice photographs and complement information on breeding, to Mr Wolfgang Eberl (Schorndorf, Germany) for having raised our attention towards these fishes and for his permanent support, and to Mr Patrice Pruvost (Paris, M.N.H.N.) who kindly worked out several times the difficult radiophotographs. The manuscript has benefited from positive contributions by Prof. Jean-Louis Amiet (now retired in the South of France), who has discovered A. festivum, by Prof J. Daget (M.N.H.N, Paris, France) and by Univ.-Prof. Dr Alfred C. Radda (Wien, Austria), who has rediscovered A. pascheni, and by anonymous reviewers. They all receive my sincere gratitude. BIBLIOGRAPHY AMIET, J.L. 1987. Faune du Cameroun. Le Genre Aphyosemion Myers. Sciences Naturelles. Compiègne: 262pp., 76 pls. HUBER, J.H. 1978. Contribution à la Connaissance des Cyprinodontidés de l'Afrique Occi- dentale: Caractères taxonomiques et Tentative de Regroupement des Espèces du Genre Aphyosemion. Revue francaise d'Aquariologie, S: 1-29, 9 figs., 30 photos, 6 maps. HUBER, J.H. 1981. A Review of the Cyprinodont Fauna of the Coastal Plain in Rio Muni, Gabon, Congo, Cabinda & Zaïre, with taxonomic Shifts in Aphyosemion, Epiplatys & West African Procatopodins. British Killifish Association Publication (Separatum): 46 pp.. 27 figs., 16 photos. HUBER, J.H. 1992. Review of Rivulus. Ecobiogeography - Relationships. Cybium Supplément , Société Francaise d'Ichtyologie Publication, Paris: 586 pp., 40 pls., 85 figs, 8 tabs, 13 maps. HUBER, J.H. 1996. Killi-Data 1996. Updated checklist of taxonomic names, collecting localities & bibliographic references of oviparous Cyprinodont Fishes (Atherinomorpha); in french, english, & german. Cybium, Société francaise d'Ichtyologie, Ed., Paris: 400 pp., 20 maps. Leviton, A.E., R.H. G1BBS JR., E. HEAL, & C.E. Dawson. 1985. Standards in herpetology and ichthyology: Part I. Standard symbolic codes for institutional resource collections in herpetology and ichthyology. Copeia 1985: 802-832. RADDA, A.C. 1975. Contribution to the Knowledge of the Cyprinodonts of Gabon with the Description of four new species & one new subspecies of the Genus Aphyosemion Myers. British Killifish Association Publication (Separatum): 20 pp., 12 pls, tab. SCHEEL, J.J. 1990. Atlas of the Killifishes of the Old World. Tropical Fish Hobbyist Publi- cation, Neptune City, 448 pp., figs. REVUE SUISSE DE ZOOLOGIE 105 (2): 339-343; juin 1998 Endémisme pyrénéen: sur une nouvelle espèce épigée du genre Oritoniscus: O. rousseti n. sp. (Crustacea, Isopoda, Oniscidea) Henri DALENS Laboratoire d’Ecologie des Invertébrés terrestres Université Paul Sabatier 118, route de Narbonne, F-31062 Toulouse Cedex / France Pyrenean endemism: O. rousseti a new epigean species (Crustacea, Isopoda, Oniscidea). - The new species here described has a very restric- ted distribution and is closely related to the species O. baroussensis and O. aurensis which have been recently described. Key-words: Isopoda - Oniscidea - Oritoniscus - Morphology - Endemism. INTRODUCTION La recherche de zones de contact entre certaines des espèces ayant fait l’objet de précédentes publications (DALENS et al. 1996; 1997) a révélé, dans une toute petite vallée pyrénéenne transversale, celle du ruisseau de Beyrède, située au sud des Baronnies, une espèce nouvelle, laquelle fait l’objet de la présente note. Oritoniscus rousseti sp. n. (Figs 1-12) Matériel examiné: Holotype (MNHN-IS 5069): 1 3 provenant de la station type au Cap de la Pène sur la route forestière de Bignec, Cne de Beyrède-Jumet (Htes- Pyrénées), alt.: 940 m, UTM31: 283400/4759450, 29.V.1997, Dalens & Rousset réc. dans des suintements sur schistes. Paratypes (pour partie au Muséum d’histoire naturelle de Genève: 5 34,5 2 2; pour partie dans la collection de l’auteur): 44 8 & trouvés dans la même station avec 1 d appartenant à l’espece O. simplex et 89 ® ®; ruisseau du Mounet, route du col de Beyrede. Cne de Beyrède-Jumet (Htes-Pyrénées), alt. 850 m, UTM31: 284550/4759600; 2 d à, 2 9 2, 29.V.1997, Dalens & Rousset réc.; Cne de Beyrède-Jumet (Htes-Pyrénées), alt. 1350 m, UTM31: 283400/4758000, 1 3, 14.IX.1997, Bedos & Deharveng, dans la litière de hétraie. DESCRIPTION: Holotype de 6 mm ce qui est la taille la plus grande observée pour les mâles jusqu'ici. Coloration brun-acajou clair, assez uniforme, mais laissant apparaître en clair des insertions musculaires et une bande à la limite tergite-pleurépimère. Péréiopodes avec un réseau pigmentaire diffus, mais pléopodes non pigmentés. Al (fig. 1) de 3 articles portant à son apex jusqu’à 12 aesthétascs disposés en peigne et flanqués sur le bord externe d’une soie courte et trapue. Flagelle antennaire (fig. 2) formé de 6 pseudo-articles, le second nettement plus long que les autres, portant une plage de 7 longs aesthétascs flanqués de 2 autres plus courts et étagés (fig. 3). Les Manuscrit accepté 09.12.1997 340 HENRI DALENS ORISTONISCUS ROUSSETI N. Sp. 341 caractères sexuels secondaires mâles affectent le péréionite I et les péréiopodes V, VI et VII. Le péréionite I présente une fossette médiane parfaitement circulaire et discer- nable à l’œil nu sur les plus gros individus de sexe mâle (figs 4 & 5). Péréiopodes V (figs 7 & 8) avec la soie sterno-basale du méros légèrement hypertrophiée mais recti- ligne. Péréiopode VI avec la soie sterno-distale de l’ischion portée par un tubercule basal (figs 9 & 10). Enfin au niveau des péréionites VII, l’angle sterno-distal du basis porte une petite touffe de soies. Les pléopodes 1 du mâle (fig. 11) présentent des exopodites à bord externe fortement concave différenciant très nettement une pointe latero-externe distale, tandis que les endopodites se terminent par une pointe courte et fine qui à fort grossissement se révèle creusée d’une gouttière (fig. 12). Discussion: La fossette présente sur le péréionite I rappelle beaucoup au premier abord celle d’O. baroussensis, elle s’en différencie cependant par sa taille qui est plus réduite et par le degré et le type de différenciation des soies. Chez O. baroussensis les soies qui ceinturent la fossette sont à base élargie et nettement distinctes de celle du reste du tergite; chez O. rousseti, elles sont certes concentriques mais elles ne sont en rien différentes de celles du reste du tergite. Chez O. baroussensis, le fond de la dépression est tapissé de soies ou de formations squameuses qui forment un tapis plus ou moins plat; chez O. rousseti, seule la zone centrale présente des soies hyper- trophiées qui forment comme un còne avec apparemment une sorte de puits central (fig. 6). Par ailleurs par rapport à O. baroussensis la zone tergale portant la fossette est beaucoup moins bombée et les zones latérales du bord postérieur du péréionite I sont nettement moins sinuées. Pour ce qui est des péréiopodes V et VI, ils ne per- mettent nullement de différencier cette nouvelle espèce d’O. aurensis ou d’O. baroussensis. En ce qui concerne le péréiopode VII, la touffe de soies sterno-distale du basis est peut-être un peu plus développée que chez O. aurensis et O. baroussensis où elle se trouve être très discrète au point de passer souvent inaperçue. Quant aux pléopodes 1 du mâle ils ne présentent pas non plus une conformation qui puisse permettre de les différencier de façon certaine des deux autres espèces: O. aurensis et O. baroussensis; seule l'extrémité de l’endopodite 1 examinée à fort grossissement se révèle quelque peu différente de celle des deux autres espèces. AFFINITÉS: La nouvelle espèce semble manifestement très proche des espèces O. aurensis et O. baroussensis et partage avec elles les caractères suivants: une fossette circulaire, une soie mérale du P.V peu différenciée et rectiligne, la soie distale de l’ischion VI du mâle portée par un tubercule et des pléopodes 1 du mâle très semblables. Seul le degré de différenciation de la fossette permet de les discriminer sans risque d’erreur, car si le schéma de base de la fossette reste le même pour les Fics 1-6 Oritoniscus rousseti N. sp. 1: Al droite; 2: flagelle antennaire; 3: detail des aesthétascs du 2e article flagellaire; 4: péréionite I avec fossette en vue dorsale; 5: péréionite I avec fossette en vue latérale gauche; 6: détail de la zone centrale de la fossette du péréionite I. (clichés H. Dalens avec Hitachi S 450). n Z m — < A 2 Z. mM a0) ORISTONISCUS ROUSSETI n. Sp. 343 3 espèces, il existe des différences nettes au niveau des composantes de cette fossette, différences qui sont déjà marquées chez l’immature. Il paraît donc tout à fait logique d’en faire 3 espèces distinctes mais très proches et dérivant manifestement d’une souche commune. RÉPARTITION: Cette espèce paraît jusqu'ici localisée dans la vallée de la Beyrède. Encore doit-on noter que dans la partie haute de la vallée, elle paraît supplantée par O. flavus. Peut-étre cependant y subsiste-t-elle par places et avec des peuplements épars et peu denses. En effet, après réexamen des échantillons, il ne fait aucun doute que 3 individus trouvés le 8 octobre 1996 dans une station de la vallée du Gripp et classés alors comme O. aurensis ou O. baroussensis atypiques, sont en fait des O. rousseti. Un prélèvement très important fait en mai 1997 dans la même station n’a toutefois par permis d’y retrouver cette espèce, mais seulement des O. flavus. Derivatio nominis: Je dédie cette espèce à mon collègue et ami André Rousset, tout récemment disparu, avec qui au cours de nombreuses sorties dans les Pyrénées, nous avons fait progresser les connaissances sur les formes épigées du genre Oritoniscus. Egalement co-découvreur de cette dernière espèce, la maladie ne lui a pas permis d’en faire l’étude. REMERCIEMENTS: Ce travail s’inscrit et est financé dans le cadre du projet européen CEE n° EVSV-CT94-0435 «High Endemism areas, Endemic biota and the conser- vation of Biodiversity in Western Europe». REFERENCES DALENS, H., A. ROUSSET & D. FOURNIER. 1996. Les formes épigées du genre Oritoniscus (Crustacea, Isopoda, Oniscidea). I. Le complexe Oritoniscus flavus. Revue suisse de Zoologie 103 (3): 623-641. DALENS, H., A. ROUSSET & D. FOURNIER. 1997. Les espèces épigées du genre Oritoniscus (Crustacea, Isopoda, Oniscidea). II. Le complexe Oritoniscus bonadonai-pyrenaeus- remyi. Revue suisse de Zoologie 104 (3): 1-23. Fics 7-12 Oritoniscus rousseti n. sp. 7: méros du péréiopode V avec soie basale hypertrophiée; 8: détail de la soie basale du méros V; 8: ischion du péréiopode VI avec soie sterno-distale sur tubercule basal: 10: détail de la soie sterno-distale de l’ischion VI; 11: première paire de pléopodes mâles: 12: détail de l’extrémité de l’endopodite 1 droit. (clichés H. Dalens avec Hitachi S 450). ery — up LEUR SMISE iu REVUE SUISSE DE ZOOLOGIE 105 (2): 345-350; juin 1998 Notes sur les Psélaphines néotropicaux (Coleoptera, Staphylinidae, Pselaphinae) 10 - un nouveau genre et quatre nouvelles espèces de la tribu des Metopiasini André COMELLINI Muséum d'histoire naturelle, Case postale 6434, CH- 1211 Genève 6 Notes on Neotropical pselaphines (Coleptera, Staphylinidae, Psela- phinae) 10 - A new genus and four new species of the tribe Meto- piasini. The genus Chandleria is described to accomodate five species, all described and/or redescribed: Metopias elegans Sharp, C. angusta sp.n., C. biguttula sp. n., C. spinosa sp. n. from Panama, and C. colombiana sp. n. from Colombia. Key-words: Coleoptera - Staphylinidae - Pselaphinae - Metopiasini - taxo- nomy - Neotropics. INTRODUCTION La tribu des Metopiasini est constituée de six genres confinés à la région néotropicale. Elle est caractérisée par la forme de la tête et par les antennes coudées, avec un scape particulièrement long (SHARP 1877, RAFFRAY 1908, PARK 1942). Parmi les Metopiasini étudiés, Metopias elegans Sharp et quatre espèces nouvelles, provenant de Panama et de Colombie, forment un groupe homogène, isolé des autres Metopiasini. Le genre Chandleria gen. n. est établi et ces espèces sont décrites ci-dessous. Les dessins des mêmes structures sont à la même échelle. Les abbréviations suivantes ont été utilisées: BMNH = British Museum of Natural History, London: CNCI = Canadian National Collection of Insects, Ottawa; DENH = University of New Hampshire Insect Collection, Durham; FMNH = Field Museum of Natural History, Chicago; MHNG = Muséum d'histoire naturelle, Genève. SYSTÉMATIQUE Chandleria gen. n. Espèce type: Chandleria angusta sp.n. Ce genre est en premier lieu caractérisé par les antennes (fig. 1). L'article 3 de celles-ci est bien plus grand que les articles 2 et 4; le tégument de l'antenne est brillant, finement granulé-ponctué. Les yeux sont gros. Les élytres réunis sont plus longs que larges avec deux fossettes basales sur chacun. Un sillon longitudinal part de chaque fossette; il est profond en avant et s'efface vers l'arrière; le sillon externe est moins marqué que l'interne. Le dessous de l'abdomen, particulièrement les sternites 6 Manuscrit accepté 14.10.1997 ANDRE COMELLINI 346 PSÉLAPHINES NEOTROPICAUX NOUVEAUX 347 et 7, forme chez les mâles une structure très caractéristique, différente pour chaque espèce. La pilosité des pattes est dirigée vers l'extrémité de celles-ci. Etymologie: Genre dédié au Dr. Donald S. Chandler de l'Université du New Hampshire, Durham, USA; genre grammatical: féminin. Chandleria angusta sp. n. Tête (fig. 2) avec une large dépression entre les yeux. Tégument brilllant, très finement et éparsément ponctué. Pilosité fine, claire et longue, plus dense sur les côtés, dirigée vers l'apex. Pronotum nettement plus long que large avec les côtés arrondis; sa plus grande largeur au milieu. Un large et profond sillon transversal au tiers postérieur. Tégument lisse et brillant, très éparsément et finement ponctué. Pilosité longue et claire, recourbée, plus courte et plus dense sur les côtés. Elytres avec les sillons longitudinaux effacés avant la moitié antérieur. Angles huméraux marqués en crête arrondie. Tégument brillant, finement et assez densément ponctué-granuleux. Pilosité claire, longue et fine, dirigée vers l'arrière, plus dense sur les côtés et à l'apex. Dessous de l'abdomen chez le mâle: fig. 7 Pattes à pilosité claire et assez longue. Tégument brillant, très finement ponctué-granuleux. Longueur: 2,50 à 2,75 mm. Edéage: fig. 12 et 13; longueur 0,31 - 0,32 mm. d, holotype: Panama, Province de Bocas del Toro, Miramar, 9°N, 82°15’W, V. 16.80, H. Wolda, UV It (DENH). 18 d, paratypes, même provenance, différentes dates de capture (DENH et MHNG). Chandleria biguttula sp. n. Tète (fig. 3) avec les deux bosses à la base du lobe frontal très en relief, en forme de goutte. Tégument brillant finement et densément ponctué. Pilosité claire et longue. Pilosité des antennes longue et assez abondante, dirigée vers l'apex. Pronotum nettement plus long que large, avec les côtés arrondis, brusquement plus étroits au tiers postérieur à la hauteur du large sillon transversal. Tégument brillant, plus lisse et plus finement ponctué que celui de la téte. Pilosité claire, longue et fine, plus abondante, plus épaisse et plus courte sur les côtés. Elytres avec les sillons longitudinaux s'effaçant déjà au quart antérieur. Angles huméraux arrondis, peu marqués. Tégument brillant, finement ponctué-strié, plus ou moins densément par place. Pilosité claire, fine et longue, plus dense sur les côtés et en arrière. Fics 1-11 1, Chandleria sp., antenne. Dessus de la tête: 2, Chandleria angusta sp. n.; 3, C. biguttula sp. n.; 4, C. colombiana sp. n.; 5, C. elegans (Sharp); 6, C. spinosa sp. n. Derniers sternites: 7, C. angusta sp. n.; 8, C. biguttula sp. n.; 9, C. colombiana sp. n.; 10, C. elegans (Sharp); 11, C. spinosa sp. n. 348 ANDRÉ COMELLINI PSÉLAPHINES NEOTROPICAUX NOUVEAUX 349 Dessous de l'abdomen chez le mâle: fig. 8. Pattes à pilosité claire et assez longue. Tégument brillant, très finement ponctué-granuleux. Longueur: 2,50 à 2,75 mm Edéage: fig. 14 et 15; longueur 0,35 - 0,36 mm. $6, holotype: Panama, Province de Bocas del Toro, Miramar, 9°N, 82°15°W, 24. VII. 1979, H. Wolda, UV It (DENH); 19 paratypes ®, même provenance, différentes dates de capture (DENH et MHNG). Chandleria colombiana sp. n. Téte (fig. 4) avec une large dépression entre les yeux, limitée en arrière par une créte granuleuse. Pilosité claire, assez longue et dense, plus rare sur le dessus. Pilosité des antennes également assez longue et dense, claire, dirigée vers l'apex. Pronotum à peine un plus large que long avec un fort sillon transversal au tiers basal et les côtés nettement applatis. Tégument brillant, lisse, avec une fine ponctuation éparse. Pilosité claire, longue et très fine, plus dense sur les còtés. Elytres avec les sillons longitudinaux s'effaçant déjà au quart antérieur. Angles huméraux peu marqués, arrondis. Tégument brillant, assez densément ponctué. Pilosité claire, assez longue et fine, plus dense et plus épaisse vers l'arrière. Dessous de l’abdomen chez le mâle: fig. 9 Pattes à pilosité assez longue, claire et fine. Tégument brillant, lisse et ponctué. Longueur: 2,1 mm. Edéage: fig. 16; longueur 0,25 mm. $d, holotype: Colombie, Anchicaya, 1000’, VII.22-27.1970, J. M. Campbell, Malaise trap (CNCI). Chandleria elegans (Sharp), comb. n. Metopias elegans Sharp, 1887 Tête (fig. 5) avec un bourrelet transversal à l'arrière, échancré au milieu, cou- vert de grosses granulations; ce bourrelet est précédé d'une dépression assez profonde. Tegument brillant, irrégulièrement granulé-ponctué, plus fortement sur les côtés de la tête. Pilosité assez longue, claire, plus fine et plus éparse sur le dessus, plus dense et plus épaisse sur les côtés. Pilosité des antennes assez longue, fine et dense, dirigée vers l'apex. Pronotum plus long que large, cordiforme, partagé au tiers postérieur, sur toute sa largeur, par un large sillon en circonflexe transversal. Ponctuation du tégument très fine, irrégulière et éparse, granulée par places. Pilosité longue et assez dense, plus éparse sur le dessus. Fics 12-19 Edéages: Chandleria angusta sp. n., 12, profil et 13, face; C. biguttata sp. n., 14, profil et 15, face; C. colombiana sp. n., 16, face; C. elegans (Sharp), 17, profil et 18, face; C. spinosa sp. n., 19, profil. 350 ANDRÉ COMELLINI Elytres avec les sillons longitudinaux très atténués vers le milieu s'effaçant complètement ensuite. Angles huméraux très obtus, arrondis. Tégument brillant, très finement ponctué-granulé. Pilosité assez dense, courbe et longue sur les côtés et en arrière, dirigée vers l'apex. Dessous de l'abdomen chez le mâle: fig. 10 Pattes à pilosité courte et assez dense. Tégument peu brillant, finement et assez densément ponctué. Longueur: 2,4 mm. Edéage: fig. 17 et 18; longueur 0,32 mm. d, type : Chiriqui (Volcan), Panama (BMNH) Chandleria spinosa sp. n. Tête (fig. 6) avec une assez forte dépression en arrière. Tégument finement ponctué-strié. Pilosité courte, claire, plus épaisse sur les còtés du lobe frontal et sur les tempes. Pilosité des antennes courte et claire, dirigée vers l'apex. Pronotum aussi long que large, avec les côtés régulièrement convexes; sa plus grande largeur un peu en arrière du milieu; au quart postérieur un large sillon transversal Tégument brillant, plus fortement ponctué-réticulé que la téte. Pilosité courte, plus dense sur les côtés. Elytres avec les sillons longitudinaux s'effaçant déjà au tiers antérieur. Angles huméraux arrondis. Tégument brillant, très finement granuleux-réticulé. Pilosité claire et courte, assez épaisse, plus dense sur les côtés. Dessous de l'abdomen chez le mâle: fig. 11 Pattes avec à l'apex des fémurs I une série de petites épines sur le dessus de la partie épaisse. Pilosité courte et assez épaisse. Ponctuation du tégument peu brillant, très fine, éparse et granuleux. Longueur: 2,6 mm. Edéage: fig. 19; longueur 0,30 mm. d, holotype: Volcan Chiriqui, Panama, 1930, A. Bierig Collin (FMNH). REMERCIEMENTS Madame E. De Boise, London et messieurs D.S. Chandler, Durham, A.F. Newton, Chicago et J.M. Campbell, Ottawa m'ont communiqués le matériel étudié. Monsieur G. Roth, Genève, a mis au net les dessins. BIBLIOGRAPHIE PARK, O. 1942. A study in Neotropical Pselaphidae. /n Northwestern University Studies in the Biological Sciences and Medicine Number 1. Norwestern University, Evanston & Chicago, x + 403 pp, 21 pls. RAFFRAY, A. 1908 Coleoptera. Fam. Pselaphidae, 487 pp, 9 pls. /n Genera Insectorum Wytsmann, P. (ed.) . Fascicule 64. Rome. SHARP, D. 1887. Fam. Pselaphidae, pp. 1-46, pl. 1. Jn Biologia Centrali-Americana. Insecta. Coleoptera, vol. 2 (1). Taylor & Francis, London. REVUE SUISSE DE ZOOLOGIE 105 (2): 351-373; juin 1998 Agathidiini from China, with description of 14 new species (Coleoptera, Leiodidae) Fernando ANGELINI* & Luigi DE MARZO** * S.S.7 per Latiano, Km. 0.500, I-72021 Francavilla Fontana (Brindisi), Italy ** Dipartimento di Biologia, Difesa e Biotecnologie agro-forestali, Università della Basilicata, Via Nazario Sauro 85, I-85100 Potenza, Italy Agathidiini from China, with description of 14 new species (Cole- optera, Leiodidae). - Records and/or descriptions are given for 18 species of Agathidiini from China, deposited in the Geneva Museum. New species are: Agathidium (Neoceble) sichuanicum n.sp. (Sichuan), A. (A.) vagum n.sp. (Guangxi), A. (A.) occultum n.sp. (Yunnan), A. (A.) gonggaense n.sp. (Sichuan), A. (A.) rufescens n.sp. (Sichuan), A. (A.) procerum n.sp. (Sichuan), A. (A.) celatum n.sp. (Hubei), A. (A.) huaense n.sp. (Shaanxi), A. (A.) brunneipenne n.sp. (Hubei), A. (A.) inerme n.sp. (Hubei), A. (A.) lugubre n.sp. (Guangxi), A. (A.) indubium n.sp. (Sichuan), A. (Microceble) corticinum n.sp. (Yunnan), A. (Microceble) solutum n.sp. (Guangxi). New records for China are: A. (Microceble) venustum Ang. & Dmz. (Guangxi), A. (Microceble) manasicum Ang. & Dmz. (Guangxi). Key-words: Leiodidae - Agathidiini - China - new species - new records. INTRODUCTION A large amount of new data on the Agathidiini fauna of China results from a study of the following material deposited in the Geneva Museum: - 16 species (87 specimens), from 5 localities in Sichuan, Guangxi, Hubei, Shaanxi and Yunnan, leg. S. Kurbatov; - 2 species (8 specimens), from 1 locality in Sichuan, leg. A. Smetana. The new data concern: - 14 new species: Agathidium (Neoceble) sichuanicum n.sp. (Sichuan), A. (A.) vagum n.sp. (Guangxi), A. (A.) occultum n.sp. (Yunnan), A. (A.) gonggaense n.sp. (Sichuan), A. (A.) rufescens n.sp. (Sichuan), A. (A.) procerum n.sp. (Sichuan), A. (A.) celatum n.sp. (Hubei), A. (A.) huaense n.sp. (Shaanxi), A. (A.) brunneipenne n.sp. (Hubei), A. (A.) inerme n.sp. (Hubei), A. (A.) lugubre n.sp. (Guangxi), A. (A.) indubium n.sp. (Sichuan), A. (Microceble) corticinum n.sp. (Yunnan), A. (Microceble) solutum n.sp. (Guangxi). Manuscript accepted 11.07.1997 392. FERNANDO ANGELINI & LUIGI DE MARZO - 2 new records for China: Agathidium (Microceble) venustum Ang. & Dmz. (Guangxi), A. (Microceble) manasicum Ang. & Dmz. (Guangxi). The specimens are now deposited either in the Geneva Museum (MHNG) or in Angelini's collection (AC). We are indebted to Dr. Ivan Lòbl for making the material avalaible for study and comments on the earlier version of the manuscript. Anisotoma Panzer, 1797 Anisotoma becvari Ang. & Svec Anisotoma becvari Angelini & Svec, 1994: 5. Material: China, West Hubei, Shennongjia Nat. Res., 2000-2200 m, 3-8.VI.1995, leg. S. Kurbatov, 2 exx. (MHNG), 1 ex. (AC). DISTRIBUTION: China (Yunnan and Hubei). New record for Hubei. Agathidium Panzer, 1797 Subg. Neoceble Gozis, 1886 marginatum group Agathidium (Neoceble) sichuanicum n.sp. Figs 1-5 Length 2.7-3.1 mm (holotype d: 2.95 mm). Dorsum reddish-brown; venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish- brown. Microreticulation superficial on entire dorsum; puncturation fine and sparse on head and pronotum, absent from elytra. Sutural striae absent. Head: Widest just behind eyes: temple 1/4 as long as the eye; anterior-lateral margins not raised; clypeus slightly emarginate; clypeal line very superficial; eyes prominent (fig. 1). Antennal segment 3 1.3 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter with 1 vesicle in 9th and 10th antennal segments. Microreticulation very superficial; punctures very small, superficial, separated from each other by 1-6 times their diameter; some extremely small punctures interposed. Pronotum: 1.45 times as broad as head, moderately broader than long (W/L = 1.7), moderately convex (W/H = 1.61); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation more distinct than on head, uniform; punctures as large as those on head, sparser, separated from each other by 5-15 times their diameter. Holotype: length 0.85 mm, width 1.45 mm, height 0.90 mm. Elytra: Moderately narrower than pronotum, nearly as long as broad (W/L = 0.96), moderately convex (W/H = 1.77); lateral outline with sharp humeral angle. Microreticulation very superficial; puncturation absent, except for some very small and poorly distinct punctures. Holotype: length 1.40 mm, width 1.35 mm, height 0.76 mm. Metathoracic wings absent. Meso- and metasternum: median carina weak, lateral lines complete, femoral lines absent. Legs. Tarsal formula: & 5-5-4, 2 4-4-4. AGATHIDIINI FROM CHINA 353 = um == = Fics 1-5 Head, male copulatory organ (lateral view, dorsal and ventral view of apex) and spermatheca of Agathidium sichuanicum n.sp. Scale: 1 division = 0.1 mm. Male copulatory organ (figs 2-4): Aedeagus comparatively stout, with proximal part simple and lateral margins gently converging towards a rounded tip. Ventral piece absent. Parameres slender, slightly enlarged at apex. Spermatheca (fig. 5): S-shaped, enlarged at mid-length. HOLOTYPE d : China, Sichuan, Wolong Nat. Res., 1700 m, 17.V.1994, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 3 2 (MHNG), 1 d and 1 £ (AC). Discussion: Agathidium sichuanicum n.sp. is the only Chinese species of the marginatum group; therefore, it is easy to distinguish from the other species of the subg. Neoceble, as it lacks sutural striae and exhibits microreticulate dorsum. Distribution: China (Sichuan). 354 FERNANDO ANGELINI & LUIGI DE MARZO Subg. Agathidium Panzer, 1797 madurense group Agathidium (Agathidium) vagum n.sp. Figs 6, 7, 10-12 Length 3.4 mm (holotype d and paratype). Dorsum reddish-brown; venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish- brown. Microreticulation absent from entire dorsum; puncturation fine and sparse on head and pronotum, absent from elytra. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus slightly emarginate; clypeal line absent; eyes prominent (fig. 6). Antennal segment 3 1.4 times as long as 2 and as long as 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures small, superficial, separated from each other by 2-8 times their diameter. Pronotum: 1.6 times as broad as head, moderately broader than long (W/L = 1.45), very convex (W/H = 1.45); anterior margin sharply curved; lateral outline broadly rounded. Punctures as large as those on head, sparser, separated from each other by 8-15 times their diameter. Holotype: length 1.10 mm, width 1.40 mm, height 1.10 mm. Po ae Fics 6-9 Head and male metafemora of: 6-7, Agathidium vagum n.sp.; 8-9, A. occultum n.sp. AGATHIDIINI FROM CHINA 355 Elytra: Slightly narrower than pronotum, slightly longer than broad (W/L = 0.96), moderately convex (W/H = 1.82); lateral outline with very weak humeral angle. Puncturation almost absent: only some very small, sparse punctures present. Holotype: length 1.55 mm, width 1.50 mm, height 0.82 mm. Metathoracic wings present. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines incomplete; a small tubercle between the metacoxae. Legs: Male hind femora broadened distally (fig. 7). Tarsal formula: 4 5-5-4, 2 not known. Male copulatory organ (figs 10-12): Aedeagus slender, with hook-like proximal part, lateral margins sinuate and converging towards a subacute tip, deeply bifid ventral piece. Parameres slender, gently narrowing towards apex. HOLOTYPE d: China, Guangxi, 15 Km North Longsheng, 1000 m, 15-22.V1.95, leg. S. Kurbatov (MHNG). PARATYPE: as holotype but 20.VI.95, 1 & (AC). Discussion: Agathidium vagum n.sp. is closely related to A. occultum n.sp. and A. becvari Ang. & Svec, from which it differs in the colour of the antennae, the ratio of the 3rd/2nd antennal segment and the shape of the male hind femora. Distribution: China (Guangxi). Agathidium (Agathidium) occultum n.sp. Figs 8, 9, 13-16 Length 3.0-3.2 mm (holotype d: 3.1 mm). Dorsum reddish-brown; venter reddish-brown, mesosternum lighter; antennae darker at segments 9-10; legs reddish- brown. Microreticulation almost absent, traceable on elytra; puncturation fine and Sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus slightly emarginate; clypeal line absent; eyes prominent (fig. 8). Antennal segment 3 1.7 times as long as 2 and as long as 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures small, superficial, separated from each other by 1-6 times their diameter. Pronotum: 1.57 times as broad as head, moderately broader than long (W/L = 1.5), moderately convex (W/H = 1.65); anterior margin slightly curved; lateral outline broadly rounded. Punctures larger and more distinctly impressed than on head, separated from each other by 4-6 times their diameter. Holotype: length 1.10 mm, width 1.65 mm, height 1.00 mm. Elytra: Moderately narrower than pronotum, slightly broader than long (W/L = 1.1), slightly convex (W/H = 1.93); lateral outline with very weak humeral angle. Microreticulation almost absent, only traceable; punctures smaller and less well impressed than on head, separated from each other by 4-15 times their diameter. Holotype: length 1.40 mm, width 1.55 mm, height 0.80 mm. Metathoracic wings present. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines incomplete; a small tubercle between the metacoxae. Legs: Male hind femora broadened distally (fig. 9). Tarsal formula: 8 5-5-4, ? 5-4-4. 356 FERNANDO ANGELINI & LUIGI DE MARZO 12 16 Fics 10-16 Male copulatory organ (lateral view, dorsal and ventral view of apex) of: 10-12, Agathidium vagum n.sp.; 13-15, A. occultum n.sp. Spermatheca of: 16, A. occultum n.sp. Scale: 1 division = 0.1 mm. Male copulatory organ (figs 13-15): Aedeagus slender, with hook-like proximal part, lateral margins sinuate and converging towards a largely rounded apex, deeply bifid ventral piece. Parameres slender, abruptly enlarged at apex. Spermatheca (fig. 16): Basal part pear-shaped; apical part short. AGATHIDIINI FROM CHINA 357 HOLOTYPE d: China, South Yunnan, Mengyang Nat. Res., 9.IX.94, 500 m, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 2 2 (MHNG), 1 2 (AC); as holotype but 8.IX.94, 1 4 (AC). Discussion: See under A. vagum n.sp. Distribution: China (South Yunnan). seminulum group Agathidium (Agathidium) uliginosum Ang. & Svec Agathidium (s.str.) uliginosum Angelini & Svec, 1994: 17. Material: China, Sichuan, Gongga Shan, above Camp 3, 3050 m, 22.VII.94, leg. A. Smetana, 3 exx. (MHNG), 2 exx. (AC). Distribution: China (Yunnan and Sichuan). laevigatum group Agathidium (Agathidium) gonggaense n.sp. Figs 17, 18, 25-27 Length 4.2-4.6 mm (holotype d: 4.5 mm). Dorsum black; venter reddish- brown, mesosternum lighter; antennae uniformly testaceous; legs reddish-brown. Entire dorsum with superficial microreticulation and very small punctures. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus moderately emarginate; clypeal line absent; eyes flattened (fig. 17). Antennal segment 3 2.1 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation superficial, uniform; punctures very small, superficial, hardly evident, separated from each other by 3-10 times their diameter. Pronotum: 1.4 times as broad as head, slightly broader than long (W/L = 1.35), very convex (W/H = 1.45); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation less well impressed than on head; punctures as large as those on head, separated from each other by 5-6 times their diameter. Holotype: length 1.55 mm, width 2.10 mm, height 1.45 mm. Elytra: Slightly narrower than pronotum, as broad as long, slightly convex (W/H = 1.9); lateral outline with very weak humeral angle. Microreticulation more distinctly impressed than on pronotum; punctures as large as those on head, separated from each other by 10-15 times their diameter. Holotype: length 2.00 mm, width 2.00 mm, height 1.05 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines complete; a small tubercle between the metacoxae. Legs: Male hind femora with pronounced tooth at posterior margin (fig. 18). Tarsal formula: 5 5-5-4, 9 not known. Male copulatory organ (figs 25-27): Aedeagus slender, with hook-like proximal part, lateral margins sinuate, apex emarginate, ventral piece spatula-like. Parameres slender, gently narrowing toward apex and curved down. 358 FERNANDO ANGELINI & LUIGI DE MARZO HoLoTyPE d : China, Sichuan, Gongga Shan, above Camp 3, 3050 m, 22.VII.94, leg. A. Smetana (MHNG). PARATYPES: China, Sichuan, Gongga Shan, Lake above Camp 2, 2750 m, 24.VII.94, leg. A. Smetana, | d (MHNG), | 3 (AC). Discussion: Within the Chinese species of the /aevigatum group, A. gongga- ense n.sp. shares its large size only with A. rufescens n.sp.; it differs from the latter in the ratio of the 3rd/2nd antennal segment and the presence of microreticulation on the entire dorsum. Distribution: China (Sichuan). Agathidium (Agathidium) rufescens n.sp. Figs 19, 20, 28-31 Length 3.6-3.9 mm (holotype d: 3.8 mm). Dorsum reddish-brown, venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish- = = E Fics 17-24 Head and male metafemora of: 17-18, Agathidium gonggaense n.sp.; 19-20, A. rufescens n.sp.; 21-22, A. procerum n.sp.; 23-24, A. celatum n.sp. AGATHIDIINI FROM CHINA 359 brown. Microreticulation absent from head and pronotum, superficial on elytra; puncturation fine and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus moderately emarginate; clypeal line absent; eyes flattened (fig. 19). Antennal segment 3 2.3 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation absent; punctures small but clearly impressed, separated from each other by 3-4 times their diameter. Pronotum: 1.26 times as broad as head, slightly broader than long (W/L = 1.17), very convex (W/H = 1.37); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation absent; punctures smaller and less well impressed than on head, separated from each other by 1-6 times their diameter. Holotype: length 1.40 mm, width 1.65 mm, height 1.20 mm. Elytra: As broad as pronotum, as broad as long, slightly convex (W/H = 2.05); lateral outline with very weak humeral angle. Microreticulation slightly impressed, difficult to see, more distinct in the less coloured specimens; punctures as large those on head, sparser, separated from each other by 2-15 times their diameter. Holotype: length 1.60 mm, width 1.65 mm, height 0.80 mm. Metathoracic wings absent. Meso- and metasternum: median carina absent, lateral lines weak and incomplete, femoral lines complete. Legs: Male hind femora with pronounced tooth at posterior margin (fig. 20). Marsalstormula: d 5-59-45 2 5-4-4) Male copulatory organ (figs 28-30): Aedeagus slender, with proximal part simple, lateral margins sinuate, apex emarginate, ventral piece not bifid. Parameres slender, gently narrowing toward apex. Spermatheca (fig. 31): S-shaped; enlarged at the duct connection. HOLOTYPE d: China, Sichuan, Wolong Nat. Res., 900 m, 23.V.94, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 2 (MHNG), 1 4 (AC). Discussion: See under A. gonggaense n.sp. Distribution: China (Sichuan). Agathidium (Agathidium) procerum n.sp. Figs 21, 2273235 Length 2.15-2.30 mm (holotype d : 2.25 mm). Dorsum reddish-brown; venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish- brown. Microreticulation superficial, uniform on entire dorsum; puncturation absent. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus moderately emarginate; clypeal line absent; eyes flattened (fig. 21). Antennal segment 3 1.1 times as long as 2 and shorter than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation uniform, very super- ficial. Pronotum: 1.4 times as broad as head, moderately broader than long (W/L = 1.57), very convex (W/H = 1.46); anterior margin slightly curved; lateral outline 360 FERNANDO ANGELINI & LUIGI DE MARZO 31 i 30 Fics 25-31 Male copulatory organ (lateral view, dorsal and ventral view of apex) of: 25-27, Agathidium gonggaense n.sp.; 28-30, A. rufescens n.sp. Spermatheca of: 31, A. rufescens n.sp. Scale: 1 division = 0.1 mm. broadly rounded. Microreticulation as that on head. Holotype: length 0.70 mm, width 1.10 mm, height 0.75 mm. Elytra: As broad as pronotum, as broad as long, slightly convex (W/H = 2); lateral outline with very weak humeral angle. Microreticulation as that on head. Holotype: length 1.05 mm, width 1.10 mm, height 0.55 mm. Metathoracic wings absent. Meso- and metasternum: median carina absent, lateral lines absent, femoral lines complete; a small tubercle between the metacoxae. Legs: Male hind femora broadened distally (fig. 22). Tarsal formula: & 5-5-4, 2 5-4-4. Male copulatory organ (figs 32-34): Aedeagus slender, with spiral proximal part, lateral margins abruptly converging toward subacute tip, deeply bifid ventral piece. Parameres slender, gently narrowing towards apex. AGATHIDIINI FROM CHINA 361 Spermatheca (fig. 35): Basal and apical parts very different in shape and length. HOLOTYPE d: China, Sichuan, Wolong Nat. Res., 1500 m, 22.V.94, leg. S. Kurbatov (MHNG). PARATYPES: as holotype but 1700 m, 17.V.94, 1 d (AC); same but 18.V.94, 1 9 (MHNG), 1 2 (AC); same but 900 m, 23.V.94, 1 & and 2 2 (MHNG), 1 d (AC). Discussion: See under A. gonggaense n.sp. Within the laevigatum group, A. procerum n.sp. is closely related to A. celatum n.sp.; it differs from the latter in the head size, the ratio of the 3rd/2nd antennal segments, the ratio of the pronotum/head width and the tarsal formula in female. Distribution: China (Sichuan). 5 3 34 39 37 38 Fics 32-39 Male copulatory organ (lateral view, dorsal and ventral view of apex) and spermatheca of: 32- 35, Agathidium procerum n.sp.; 36-39, A. celatum n.sp. Scale: 1 division = 0.1 mm. 362 FERNANDO ANGELINI & LUIGI DE MARZO Agathidium (Agathidium) celatum n.sp. Figs 23, 24, 36-39 Length 2.2-2.3 mm (holotype ¢: 2.2 mm). Dorsum and venter reddish-brown; antennae uniformly testaceous; legs reddish-brown. Microreticulation present only on elytra; puncturation fine and sparse on head and pronotum. Sutural striae absent. Head: Widest just behind eyes: temple 1/2 as long as the eye; anterior-lateral margins not raised; clypeus deeply emarginate; clypeal line absent; eyes flattened, hardly distinct in dorsal view (fig. 23). Antennal segment 3 1.3 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures very small, superficial, separated from each other by 1-8 times their diameter. Pronotum: 1.2 times as broad as head, moderately broader than long (W/L = 1.5), moderately convex (W/H = 1.54); anterior margin sharply curved; lateral outline broadly rounded. Punctures larger and more distinctly impressed than on head, separated from each other by 2-6 times their diameter. Holotype: length 0.70 mm, width 1.05 mm, height 0.68 mm. Elytra: Slightly narrower than pronotum, as broad as long, moderately convex (W/H = 1.58); lateral outline with very weak humeral angle. Microreticulation superficial, uniform; puncturation absent, except for some very small punctures. Holotype: length 0.95 mm, width 0.95 mm, height 0.60 mm. Metathoracic wings absent. Meso- and metasternum: median carina weak, lateral lines absent, femoral lines complete: a small tubercle between the metacoxae. Metasternum very short. Legs: Male hind femora broadened distally (fig. 24). Tarsal formula: 4 5-5-4, 2 4-4-4. Male copulatory organ (figs 36-38): Aedeagus slender, with spiralled proximal part, lateral margins gently converging towards a arrow-like apex, ventral piece not bifid. Parameres slender, gently narrowing towards apex. Spermatheca (fig. 39): Basal part pear-shaped; apical part short and tapered. HOLOTYPE d : China, West Hubei, Shennongjia Nat. Res., 2000-2200 m, 7.VI.95, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 3-8.VI.95, 1 2 (MHNG), 1 2 (AC). Discussion: See under A. procerum n.sp. Distribution: China (West Hubei). dentatum group Agathidium (Agathidium) huaense n.sp. Figs 40, 45, 50-53 Length 2.9-3.1 mm (holotype d: 3.1 mm). Dorsum and venter reddish-brown; antennae uniformly testaceous; legs reddish-brown. Microreticulation almost absent, traceable on elytra; puncturation fine and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus slightly emarginate; clypeal line absent; eyes flattened (fig. 40). Antennal segment 3 1.5 times AGATHIDIINI FROM CHINA 363 as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Punctures small, superficial, separated from each other by 4-6 times their diameter. Pronotum: 1.27 times as broad as head, slightly broader than long (W/L = 1.33), moderately convex (W/H = 1.55); anterior margin slightly curved; lateral outline broadly rounded. Punctures smaller and more superficial than on head, separated from each other by 1-6 times their diameter. Holotype: length 1.05 mm, width 1.40 mm, height 0.90 mm. Elytra: As broad as pronotum, as broad as long, slightly convex (W/H = 2); lateral outline with very weak humeral angle. Microreticulation almost absent, only traceable; punctures larger than on pronotum, very superficial, separated from each other by 3-10 times their diameter. Holotype: length 1.30 mm, width 1.40 mm, height 0.70 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines incomplete. Legs: Male hind femora with pronounced tooth at posterior margin (fig. 45). Tarsal formula: & 5-5-4, 2 5-4-4. Male copulatory organ (figs 50-52): Aedeagus slender, with hook-like proximal part, lateral margins sinuate, apex truncate, ventral piece not bifid. Parameres slender, enlarged at apex. Spermatheca (fig. 53): Basal part pear-shaped; apical part short and tapered. HOLOTYPE d: China, Shaanxi, Mt. Hua, 500 m, 12.V.94, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 2 (MHNG), 1 4 and 1 9 (AC). Discussion: A. huaense n.sp. is closely related to A. fukiense Ang. & Dmz. and A. brunneipenne n.sp.; it differs from A. fukiense in the colour of antennae and the lack of metathoracic wings; it differs from A. brunneipenne n.sp. in the ratio of the 3rd/2nd antennal segments, the shape of the male hind femora and the tarsal formula. Distribution: China (Shaanx1). Agathidium (Agathidium) brunneipenne n.sp. Figs 41, 46, 54-57 Length 2.7-3.0 mm (holotype ¢: 2.9 mm). Head and pronotum reddish-brown, elytra either black or reddish-brown; venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish-brown. Microreticulation almost absent, traceable on elytra; punctures very small and sparse on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus deeply emarginate; clypeal line absent; eyes flattened (fig. 41). Antennal segment 3 1.1 times as long as 2 and shorter than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation absent; punctures small, superficial, hardly distinct, separated from each other by 4-10 times their diameter. Pronotum: 1.39 times as broad as head, slightly broader than long (W/L = 1.32), very convex (W/H = 1.46); anterior margin slightly curved; lateral outline broadly rounded. Microreticulation absent; puncturation as that on head. Holotype: length 1.00 mm, width 1.32 mm, height 0.90 mm. 364 FERNANDO ANGELINI & LUIGI DE MARZO 40 41 42 43 44 Fics 40-44 Head of: 40, Agathidium huaense n.sp.; 41, A. brunneipenne n.sp.; 42, A. inerme n.sp.; 43, A. lugubre n.sp.; 44, A. indubium n.sp. Elytra: Moderately narrower than pronotum, slightly longer than broad (W/L= 0.96), very convex (W/H = 1.36); lateral outline with very weak humeral angle. Microreticulation almost absent, only traceable; punctures as those on head. Holotype: length 1.25 mm, width 1.20 mm, height 0.88 mm. Metathoracic wings absent. Meso- and metasternum: median carina weak, lateral lines absent, femoral lines incomplete; a small tubercle between the metacoxae. Metasternum very short. Legs: Male hind femora broadened distally (fig. 46). Tarsal formula: & 4-4-4, 2 4-4-4. Male copulatory organ (figs 54-56): Aedeagus comparatively stout, with spiralled proximal part, lateral margins subparallel, apex broadly rounded, deeply bifid ventral piece. Parameres slender, gently narrowing towards apex. Spermatheca (fig. 57): Basal part pear-shaped; apical part slender. AGATHIDIINI FROM CHINA 365 HOLOTYPE à : China, West Hubei, Shennongjia Nat. Res., 2000-2200 m, 3-8.V1.95, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 4 3 and 13 9 (MHNG), 3 d and 3 2 (AC). Discussion: See under A. huaense n.sp. Distribution: China (West Hubei). Agathidium (Agathidium) inerme n.sp. Figs 42, 47, 58-60 Length 5.6-6.0 mm (holotype d: 5.95 mm). Dorsum black, venter reddish- brown, mesosternum lighter; antennae uniformly dark; legs reddish-brown. Micro- reticulation almost absent, traceable on elytra; puncturation fine on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus moderately emarginate; clypeal line absent; eyes prominent (fig. 42). Antennal segment 3 2.3 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter with 2 vesicles in 9th and 10th antennal segments; gutter without vesicles in the 7th antennal segment. Microreticulation absent; punctures small, impressed, separated from each other by 2-4 times their diameter. Pronotum: 1.48 times as broad as head, slightly broader than long (W/L = 1.37), very convex (W/H = 1.41); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation absent; punctures smaller and less well impressed than on head, separated from each other by 4-6 times their diameter, some very small punctures interposed. Holotype: length 2.00 mm, width 2.75 mm, height 1.95 mm. Elytra: Moderately narrower than pronotum, slightly longer than broad (W/L= 0.95), moderately convex (W/H = 1.79); lateral outline with very weak humeral angle. Microreticulation almost absent, only traceable; punctures as large as those on head, less well impressed, separated from each other by 3-8 times their diameter. Holotype: length 2.75 mm, width 2.60 mm, height 1.45 mm. Metathoracic wings absent. Meso- and metasternum: median carina weak, lateral lines complete, femoral lines complete. Legs: Male hind femora with pronounced distal tooth (fig. 47). Tarsal formula: 3 5-5-4, 2 not known. Male copulatory organ (figs 58-60): Aedeagus slender, with hook-like proximal part, lateral margins sinuate, apex truncate, deeply bifid ventral piece. Parameres slender, gently narrowing towards apex. HOLOTYPE d : China, West Hubei, Shennongjia Nat. Res., 2000-2200 m, 3-8.V1.95, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 d (AC). Discussion: A. inerme n.sp. is clearly different from all the Chinese Agathi- dium in its very large size and the ratio of the 3rd/2nd antennal segment. Distribution: China (West Hubei). Agathidium (Agathidium) lugubre n.sp. Figs 43, 48, 61-64 Length 3.95 mm (holotype d and paratype). Dorsum entirely black in the paratypes, reddish-brown at head in the holotype; mesosternum testaceous, meta- 366 FERNANDO ANGELINI & LUIGI DE MARZO 46 45 48 2 49 Fics 45-49 Male metafemora of: 45, Agathidium huaense n.sp.; 46, A. brunneipenne n.sp.; 47, A. inerme n.sp.; 48, A. lugubre n.sp.; 49, A. indubium n.sp. sternum reddish-brown, abdomen black; antennae uniformly testaceous; legs reddish- brown. Microreticulation almost absent, only some traces on elytra; puncturation fine and sparse on head and pronotum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins not raised; clypeus moderately emarginate; clypeal line absent; eyes prominent (fig. 43). Antennal segment 3 2.1 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation absent; punctures small, superficial, separated from each other by 2-6 times their diameter. Pronotum: 1.3 times as broad as head, slightly broader than long (W/L = 1.38), very convex (W/H = 1.4); anterior margin sharply curved; lateral outline broadly rounded. Puncturation as that on head. Holotype: length 1.32 mm, width 1.82 mm, height 1.30 mm. Elytra: Slightly narrower than pronotum, as broad as long, moderately convex (W/H = 1.7); lateral outline with very weak humeral angle. Punctures as large as those on head, less clearly impressed, more superficial, difficult to see. Holotype: length 1.70 mm, width 1.70 mm, height 1.00 mm. Metathoracic wings absent. Meso- and metasternum: median carina sharp, lateral lines absent, femoral lines complete. Legs: Male hind femora with weak distal tooth (fig. 48). Tarsal formula: d 5- 5-4, 2 5-4-4. AGATHIDIINI FROM CHINA 367 Male copulatory organ (figs 61-63): Aedeagus slender, with hook-like proximal part, lateral margins sinuate, apex truncate and slightly emarginate, deeply bifid ventral piece. Parameres slender, gently narrowing towards apex. Spermatheca (fig. 64): Basal part globose; apical part slender. HOLOTYPE d: China, Guangxi, 15 Km North Longsheng, 1000 m, 20.VI.95, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 & and 3 2 (MHNG), 1 d and I 9 (AC). Discussion: Agathidium lugubre n.sp. 1s closely related to A. yunnanicum Ang. & Svec and A. indubium n.sp. Its identification must be based on the male copulatory organ. Distribution: China (West Hubei). Agathidium (Agathidium) indubium n.sp. Figs 44, 49, 65-68 Length 3.7-4.0 mm (holotype d: 3.95 mm). Dorsum black, reddish-brown at head, venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish-brown. Microreticulation almost absent, traceable on elytra; puncturation fine and hardly distinct on entire dorsum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins slightly raised; clypeus slightly emarginate; clypeal line absent; eyes prominent (fig. 44). Antennal segment 3 2 times as long as 2 and longer than 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Microreticulation absent; punctures very small, superficial, hardly visible, separated from each other by 4-10 times their diameter. Pronotum: 1.4 times as broad as head, slightly broader than long (W/L = 1.28), very convex (W/H = 1.38); anterior margin slightly curved; lateral outline broadly rounded. Microreticulation absent; puncturation smaller and less well impressed than on head, hardly distinct, separated from each other by 5-15 times their diameter. Holotype: length 1.40 mm, width 1.80 mm, height 1.30 mm. Elytra: As broad as pronotum, moderately broader than long (W/L=1.09), slightly convex (W/H = 2.25); lateral outline with very weak humeral angle. Micro- reticulation almost absent, only traceable; puncturation absent, except for some very small punctures. Holotype: length 1.65 mm, width 1.80 mm, height 0.80 mm. Metathoracic wings absent. Meso- and metasternum: median carina short, lateral lines complete, femoral lines complete; a small tubercle between the meta- coxae. Legs: Male hind femora with sharp tooth at posterior margin (fig. 49). Tarsal formula: d 5-5-4, 9 5-4-4. Male copulatory organ (figs 65-67): Aedeagus slender, with hook-like proximal part, lateral margins subparallel, apex deeply emarginate, bifid ventral piece. Parameres slender, gently narrowing towards apex and curved up. Spermatheca (fig. 68): Basal part globose; apical part short. HOLOTYPE d : China, Sichuan, Mt. Emei, 1600 m, 28.IX.94, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 2 (AC). 368 FERNANDO ANGELINI & LUIGI DE MARZO Discussion: See under A. lugubre n.sp. Distribution: China (Sichuan). 53 51 52 57 Fics 50-57 Male copulatory organ (lateral view, dorsal and ventral view of apex) and spermatheca of: 50- 53, Agathidium huaense n.sp.; 54-57, A. brunneipenne n.sp. Scale: 1 division = 0.1 mm. Subg. Microceble Angelini & De Marzo, 1986 grouvellei group Agathidium (Microceble) corticinum n.sp. Figs 69, 70, 73-76 Length 2.65-2.80 mm (holotype d : 2.65 mm). Dorsum reddish-brown, darker at elytra, venter reddish-brown, mesosternum lighter; antennae with segments 7-10 AGATHIDIINI FROM CHINA 369 Fics 58-68 Male copulatory organ (lateral view, dorsal and ventral view of apex) of: 58-60, Agathidium inerme n.sp.; 61-63, A. lugubre n.sp.; 65-67, A. indubium n.sp. Spermatheca of: 64, A. lugubre n.sp.; 68, A. indubium n.sp. Scale: 1 division = 0.1 mm. 370 FERNANDO ANGELINI & LUIGI DE MARZO darker; legs reddish-brown. Head striolate only on clypeus, pronotum with some traces of microreticulation; puncturation fine and sparse on head and pronotum. Sutural striae absent. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus slightly emarginate, with a short groove and a small pit at each side; eyes hemisphe- rical (fig. 69). Antennal segment 3 1.2 times as long as 2 and as long as the 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal segments. Head striolate only on clypeus; punctures small, superficial, separated from each other by 4-6 times their diameter. Pronotum: 1.73 times as broad as head, slightly broader than long (W/L = 1.38), very convex (W/H = 1.47); anterior margin slightly curved; lateral outline broadly rounded. Microreticulation almost absent, only traceable; punctures smaller and less well impressed than on head, separated from each other by 8-10 times their diameter. Holotype: length 0.90 mm, width 1.25 mm, height 0.85 mm. Elytra: Slightly narrower than pronotum, longer than broad (W/L= 0.98), moderately convex (W/H = 1.71); lateral outline with very weak humeral angle. Microreticulation absent; puncturation absent, except for some very small punctures. Holotype: length 1.22 mm, width 1.20 mm, height 0.70 mm. Metathoracic wings present. Meso- and metasternum: median carina weak, lateral lines absent, femoral lines incomplete; a pronounced tubercle between the metacoxae. Legs: Male hind femora broadened distally (fig. 70). Tarsal formula: 4 5-5-4, 2 5-4-4. Male copulatory organ (figs 73-75): Aedeagus slender, with lateral margins gently converging towards a subacute tip, deeply bifid ventral piece. Bann slender, gently narrowing towards apex. Spermatheca (fig. 76): Basal part pear-shaped; apical part short. HOLOTYPE d: China, South Yunnan, Mengyang Nat. Res., 9.[X.94, 500 m, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 2 (MHNG), 1 2 (AC). Discussion: A. corticinum n.sp. does differs from the only other known Chinese species of the grouvellei group, A. melanarium Ang. & Svec, in the size, the microsculpture of head and pronotum, the colour of antennae and dorsum, the ratio of the 3rd/2nd antennal segment. Distribution: China (South Yunnan). andrewesi group Agathidium (Microceble) solutum n.sp. Figs 71, 72, 77-80 Length 2.6-2.8 mm (holotype d: 2.65 mm). Dorsum reddish-brown, venter reddish-brown, mesosternum lighter; antennae uniformly testaceous; legs reddish- brown. Microreticulation absent; puncturation fine and sparse on entire dorsum. Sutural striae absent. AGATHIDIINI FROM CHINA 371 CR a 69 71 70 72 FIGS 69-72 Head and male metafemora of: 69-70, Agathidium corticinum n.sp.; 71-72, A. solutum n.sp. Head: Widest at eyes; anterior-lateral margins distinctly raised; clypeus slight- ly emarginate, with a short groove and a small pit at each side; eyes hemispherical (fig. 71). Antennal segment 3 1.3 times as long as 2 and as long as the 4 and 5 combined. Hamann's organ: gutter without vesicles in 9th and 10th antennal seg- ments. Microreticulation absent; punctures small, superficial, separated from each other by 3-6 times their diameter. Pronotum: 1.65 times as broad as head, slightly broader than long (W/L = 1.28), very convex (W/H = 1.31); anterior margin sharply curved; lateral outline broadly rounded. Microreticulation absent; puncturation as that on head. Holotype: length 0.90 mm, width 1.16 mm, height 0.88 mm. Elytra: Slightly narrower than pronotum, moderately longer than broad (W/L= 0.91), moderately convex (W/H = 1.57); lateral outline with very weak humeral angle. Microreticulation absent; puncturation as that on head. Holotype: length 1.20 mm, width 1.10 mm, height 0.70 mm. Metathoracic wings absent. Meso- and metasternum: median carina weak, lateral lines absent, femoral lines incomplete; a pronounced tubercle between the metacoxae. Legs: Male hind femora simple (fig. 72). Tarsal formula: & 5-5-4, 9 5-4-4. Male copulatory organ (figs 77-79): Aedeagus stout, with twisted proximal part, lateral margins sinuate and converging towards a broadly rounded apex, deeply bifid ventral piece. Parameres slender, moderately enlarged at apex. Spermatheca (fig. 80): Basal part pear-shaped, prolonged towards the duct connection; apical part short. 372 FERNANDO ANGELINI & LUIGI DE MARZO HOLOTYPE d : China, Guangxi, 15 Km North Longsheng, 1000 m, 20.VI.95, leg. S. Kurbatov (MHNG). PARATYPES: as holotype, 1 ® (AC); same but 21.VI.95, 1 d (AC); same but 15- 22.VI.95, 3 2 (MHNG), 1 d (AC). Discussion: In China, the andrewesi group includes a total of 3 species, which can be distinguished from each other only on the basis of the male copulatory organ. Distribution: China (Guangxi). Fics 73-80 Male copulatory organ (lateral view, dorsal and ventral view of apex) and spermatheca of: 73- 76, Agathidium corticinum n.sp.; 77-80, A. solutum n.sp. Scale: 1 division = 0.1 mm. AGATHIDIINI FROM CHINA 373 Agathidium (Microceble) venustum Ang. & Dmz. Agathidium (Microceble) venustum Angelini & De Marzo, 1995: 250. Material: China, Guangxi, 15 Km North Longsheng, 1000 m, 22.V1.95, leg. S. Kurbatov, 3 exx. (MHNG), 2 exx. (AC). Distribution: China (Guangxi), Taiwan. New record for China. Agathidium (Microceble) manasicum Ang. & Dmz. Agathidium (Microceble) manasicum Angelini & De Marzo, 1986: 445; ANGELINI, 19922209: Material: China, Guangxi, 15 Km North Longsheng, 1000 m, 22.V1.95, leg. S. Kurbatov, 2 exx. (MHNG), 1 exx. (AC). Distribution: China (Guangxi), Thailand, India (Assam). New record for China. REFERENCES ANGELINI F., 1992 - Agathidium raccolti in Thailandia dal Dr P. Schwendinger con descrizione di 4 nuove specie (Coleoptera, Leiodidae). Revue suisse de Zoologie 99: 201-209. ANGELINI F. & L. DE Marzo, 1986 - Agathidium from India and Malaya: expedition of Geneva Natural History Museum (Coleoptera, Leiodidae). Revue suisse de Zoologie 93 (2): 423-455. ANGELINI F. & L. DE Marzo, 1995 - Agathidiini from Taiwan collected by Dr. A. Smetana (Coleoptera, Leiodidae, Agathidiini). Revue suisse de Zoologie 102 (1): 175-255. ANGELINI F. & Z. Svec, 1994 - Review of Chinese species of the subfamily Leiodinae. Acta Societas Zoologica Bohemoslava 58: 1-31. 4 ek è: i) Al BORN. ur LA. eet | my), 7 ry 4 P 1 L Bar I 4 ee FL I LIZON A, 1 ree 7 fy REVUE SUISSE DE ZOOLOGIE 105 (2): 375-382; juin 1998 The genus Laena Latreille (Coleoptera: Tenebrionidae) in Thailand, with descriptions of new species Wolfgang SCHAWALLER* Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart, Germany. The genus Laena Latreille (Coleoptera: Tenebrionidae) in Thailand, with descriptions of new species. - Laena burkhardti sp.n., Laena loebli sp.n., Laena masumotoi sp.n., Laena pseudosiamica sp.n. and Laena schwendingeri sp.n. from Thailand are described as new, another 5 known species are redescribed. All species of the genus Laena Latreille, 1829 are supposed to be endemic in that country. An identification key for all known 10 species from Thailand is provided. Key-words: Coleoptera - Tenebrionidae - Laena - New species - Thailand. INTRODUCTION The tenebrionid genus Laena Latreille, 1829 contains numerous species, distri- buted from eastern Europe and the Caucasus to Middle Asia (SCHAWALLER 1995a), the Himalayas (KASZAB 1977; SCHUSTER 1926, 1935), China, Vietnam (MASUMOTO 1995, 1996a, 1996b), Japan and southward up to continental Malaysia (SCHAWALLER 1995b). From Thailand, 5 species were known by now (KASZAB & CHUJO 1966, KASZAB 1973, MASUMOTO 1989, 1996b), but new collections mainly by Drs D. Burckhardt and I. Löbl from the Geneva museum show, that at least 10 species occur in Thailand. These are summarized alphabetically in the present paper. None of the species from Thailand could be found in huge material which I have seen from the Himalayas and from Malaysia. Neither are they conspecific with recently described species from China and Vietnam, so far it can be judged by the descriptions and by the comparison of single types. The species characters within the genus are discussed by SCHAWALLER (1995b), a natural subgeneric classification of the species- rich genus is still lacking. Thus, the very probably endemic Thailandese species are not compared herein with species from other regions, but separated by an own identi- fication key. The flightless species usually possess quite small areas. The species of Laena are characteristic elements of the soil fauna in mature forests of different composition, thus all Thailandese specimens have been collected by soil sifting in forests. However, in other regions some species also live in treeless alpine or steppe habitats. MATERIAL MHNG Muséum d'histoire naturelle Genève NSMT National Science Museum Tokyo SMNS Staatliches Museum für Naturkunde Stuttgart * Contribution to Tenebrionidae, no. 20. For no. 19 see: Stuttgarter Beitr. Naturk. (A) 566, 1997. Manuscript accepted 27.08.1997 376 WOLFGANG SCHAWALLER THE SPECIES OF LAENA FROM THAILAND Laena angkhangensis Masumoto, 1996 (Figs 1-3) Material: Thailand, Chiang Mai, Fang Distr., Doi Angkhang, 7.11.1989 leg. K. Masumoto, 1 male holotype NSMT. DESCRIPTION: Head with equal punctures, distance of punctures 1-3times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 8 facets. Pronotum shape Fig. 1; pronotum shining, punctures somewhat smaller and distinctly more scattered than on head, clothed only at all sides with setae as head; lateral margin with distinct and crenulate border, basal and distal margin unbordered; propleures with punctation distinctly smaller and with setation somewhat shorter than on pronotum. Elytra with 10 rows of punctures, second row with about 38 punctures, nearly all punctures with a long seta (Fig. 2); intervals gently convex, shining, scattered with small punctures, most with a long erect seta; basal part of elytra impressed along sutura; interval III with a distinct setiferous umbilicate pore at the tip, interval VII with a distinct setiferous umbilicate pore at the shoulders and interval IX with 4 setiferous umbilicate pores between the shoulders and the tip. Profemur, mesofemur and metafemur each with a distinct and acute spine. Aedeagus Fig. 3. Body length 7.5 mm. DISTRIBUTION: Known only from the type locality Doi Angkhang in northern Thailand. Laena burckhardti sp.n. (Figs 4-6) Holotype (male): Thailand, NE Bangkok, Khao Yai Nat. Park, Khao Khieo, 1150 m, 28.X1.1985 leg. D. Burckhardt & I. Löbl, MHNG. Paratypes: Same data as holotype, 9 ex. MHNG, 5 ex. SMNS. Derivatio nominis: Dedicated to Dr Daniel Burckhardt, Museum of Natural History in Basel, one of the collectors of the type series. DESCRIPTION: Head with equal and large punctures, distance of punctures 0.5-1times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 5-6 facets. Pronotum shape Fig. 4; pronotum shining, punctured and clothed with setae as head; lateral margin marked as a distinct waved line, basal and distal margins unbordered; propleures with punctation but without setation as on pronotum. Elytra scattered with large punctures as on head and pronotum, not forming distinct rows and intervals (Fig. 5), all punctures with a long and erect seta; space between punctures flat and shining, elytra with 2 distinct setiferous umbilicate pores at the shoulders and with 4 distinct setiferous umbilicate pores at the tip. Legs without peculiarities. Aedeagus Fig. 6. Body length 1.9-3.2 mm. Laena fangensis Masumoto, 1996 (Figs 7-9) Material: Thailand, Chiang Mai, Fang Distr., 22.V.1993, male holotype NSMT. DESCRIPTION: Head roughly punctured, distance of punctures 0.5-1times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 12 facets. Pronotum shape Fig. 7; pronotum shagreened, sowewhat uneven, roughly punctured as LAENA IN THAILAND 377 head, clothed only laterally with setae shorter than on head; lateral margin distinctly bordered, basal and distal margins unbordered; propleures with punctation distinctly smaller and sparser and with setation shorter than on pronotum. Elytra with 10 rows of punctures, lateral rows irregularly and fused, second row with about 50 punctures, punctures without distinct seta (only with microseta shorter than diameter of puncture) (Fig. 8); internal intervals feebly convex, external intervals convexer, interval IX distinctly ridged, shagreened, very sparsely scattered with small punctures, each with a short seta; basal part of elytra impressed along sutura; interval XI with 3 setiferous umbilicate pores between the shoulders and the tip. Profemur with a distinct spine at the dorsal anterior margin, metatibia with a distinct spine at the inner distal margin. Aedeagus Fig. 9. Body length 10.5 mm. DISTRIBUTION: Known only from the type locality Fang Distr. in northern Thailand. Laena loebli sp.n. (Figs 10-11) Holotype (female): Thailand, Chiang Mai, Doi Inthanon, 2450 m, 9.X1.1985 leg. D. Burckhardt & I. Lòbl, MHNG. Derivatio nominis: Dedicated to Dr Ivan L6bl, Muséum d'histoire naturelle in Geneva, one of the collectors of the type series. DESCRIPTION: Head with equal punctures, distance of punctures 2-5times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 7-8 facets. Pronotum shape Fig. 10; pronotum shining, punctured and clothed with setae as head; lateral margin distinctly bordered, basal and distal margins unbordered; propleures with Same punctation but without setation as pronotum. Elytra with 10 rows of punctures, second row with about 35 punctures, punctures without distinct seta (only with microseta shorter than diameter of puncture) (Fig. 11); intervals flat, shining, very sparsely scattered with small punctures, each with a long erect seta as on pronotum; interval VII with a distinct setiferous umbilicate pore at the shoulders and interval IX with 4 setiferous umbilicate pores between the shoulders and the tip. Legs without peculiarities. Aedeagus unknown. Body length 8.5 mm. Laena masumotoi sp.n. (Figs 12-14) Holotype (male): Thailand, Chanthaburi, Khao Sabap Nat. Park, 150-300 m, 23.-24.X1.1985 leg. D. Burckhardt & I. Lobl, MHNG. Paratypes: Same data as holotype, 2 ex. MHNG, 1 ex. SMNS. Thailand, Phetchaburi, Kaeng Krachan Nat. Park, 450 m, 19.X1.1985 leg. D. Burckhardt & I. Lobl, 1 ex. MHNG. Derivatio nominis: Dedicated to Dr Kimio Masumoto, Tokyo, who published important taxonomic contributions to the Tenebrionid fauna of Thailand and adjacent countries. DESCRIPTION: Head roughly punctured, distance of punctures 1-2times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 6-7 facets. Pronotum shape Fig. 12; pronotum shining, punctured and clothed with setae as head: lateral margin marked as a distinct waved line, basal and distal margins unbordered; 378 WOLFGANG SCHAWALLER propleures with punctation distinctly smaller and with setation somewhat shorter than on pronotum. Elytra with 10 rows of large punctures, second row with about 27 punctures, nearly all punctures with a seta only somewhat shorter than those on the intervals (Fig. 13); intervals convex, shining, scattered with small punctures, each with a long erect seta, on lateral intervals these punctures elevated, thus intervals crenulated; interval VII with a indistinct setiferous umbilicate pore at the shoulders, interval IX with 3 indistinct setiferous umbilicate pores between the shoulders and the tip. Legs without peculiarities. Aedeagus Fig. 14. Body length 2.8-4.0 mm. Laena pseudosiamica sp.n. (Figs 15-17) Holotype (male): Thailand, Kanchanaburi, Sai Yok Nat. Park, 100 m, 21.VII.1987 leg. P. Schwendinger, MHNG DESCRIPTION: Head roughly punctured, distance of punctures 1-2times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 6-7 facets. Pronotum shape Fig. 15; pronotum shining, punctures bigger than on head and partly confluenting, clothed with setae as head; lateral margin marked as a distinct waved line, basal margin deeper than disc of pronotum, distal margin unbordered; propleures with punctation somewhat smaller and with setation somewhat shorter than on pronotum. Elytra with 10 rows of large punctures, second row with about 25 punctures, nearly all punctures with a seta only somewhat shorter than those on the intervals (Fig. 16); intervals convex, shining, scattered with small punctures, each with a long erect seta, on lateral intervals these punctures elevated, thus intervals crenulated; interval IX with 2 indistinct setiferous umbilicate pores between the shoulders and the tip. Legs with all the femora with 2 spines situated on the internal and external margin, and with all the tibiae distinctly bended. Aedeagus Fig. 17. Body length 4.0 mm. Laena schwendingeri sp.n. (Figs 19-21) Holotype (male): Thailand, Chiang Mai, Doi Inthanon, 1650 m, 7.X1.1985 leg. D. Burckhardt & I. Löbl, MHNG. Paratypes: Same data as holotype, 2 ex. MHNG, 2 ex. SMNS. Thailand, Chiang Mai, Doi Suthep, 1450 m, 4.X1.1985 leg. D. Burckhardt & I. Löbl, 2 ex. MHNG. Mae Hong Son, Doi Chang, 20 km E Pai, 1930 m, 4.V1.1986 leg. P. Schwendinger, 1 ex. MHNG. Derivatio nominis: Dedicated to Dr Peter Schwendinger, who improved our knowledge about the soil fauna of Thailand with huge collections. DESCRIPTION: Head roughly punctured, distance of punctures 1-4times as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 6-7 facets. Pronotum shape Fig. 19; pronotum shining, punctures somewhat smaller and distinctly more scattered than on head, clothed with setae as head; lateral margin marked only as a feeble line or nearly absent, basal margin deeper than disc of pronotum, distal margin unbordered; propleures with punctation somewhat smaller and with setation somewhat shorter than on pronotum. Elytra with 10 rows of punctures, second row with about 25 LAENA IN THAILAND 379 punctures, nearly all punctures with a long seta only somewhat shorter than those on the intervals (Fig. 20); intervals convex, shining, scattered mainly laterally with small punctures, each with a long erect seta; basal part of elytra impressed along sutura; interval III with a distinct setiferous umbilicate pore at the tip, interval VII with a distinct setiferous umbilicate pore at the shoulders and interval IX with 4 setiferous umbilicate pores between the shoulders and the tip. Legs without peculiarities. Aedeagus Fig. 21. Body length 4.3-6.0 mm. Laena siamica Kaszab, 1973 (Fig. 18) Material: Notavailable. Remarks: From the outer appearance, the above described pseudosiamica sp.n. is quite similar to siamica and on the first view they might be considered as conspecific. However, siamica is said to have the femora with only 1 spine (in pseudosiamica sp.n. with 2 spines situated on the internal and external margin), and the parameres are quite prolonged (Fig. 18) (in pseudosiamica sp.n. triangular, Fig. 17). In particular the last difference is significant and can not be neglected, thus both must be considered as different species. DISTRIBUTION: Known only from the type locality Kachong forest in southern Thailand (not located). Laena thailandica Kaszab & Chujo, 1966 Material: Notavailable. Remarks: This species can be recognized by the short body length (3.5 mm) and the round shape of the elytra, the lacking of a dorsal setation, the shining surface and the pattern of the elytral punctures (foto in KASZAB & CHUJO 1966: plate 1,6). The original description points to distinct "knobs" on the elytral intervals VII and IX, which are very probably distinct setiferous umbilicate pores. The sex of the single type and the shape of the aedeagus are unknown. DISTRIBUTION: Known only from the type locality Khao Luang in Thailand (not located). Laena uenoi Masumoto, 1989 (Figs 22-24) (©, Material: Thailand, Chiang Mai, Doi Inthanon, 2500 m, 2.1.1981 leg. L. Deharveng, 2 ex. MHNG. Thailand, Chiang Mai, Doi Inthanon, 1650 m, 7.X1.1985 leg. D. Burckhardt & I. Löbl, 3 ex. MHNG. Thailand, Chiang Mai, Doi Inthanon, 2450-2500 m, 9.X1.1985 leg. D. Burckhardt & I. Löbl, 4 ex. MHNG, 4 ex. SMNS. DESCRIPTION: Head with punctures of different size, distance of punctures 2-Stimes as diameters, nearly all punctures with a long seta; diameter of eyes consists of about 6-7 facets. Pronotum shape Fig. 22; pronotum shagreened, punctured and clothed with setae as head; lateral and basal margins distinctly bordered, distal margin unbordered; propleures without punctation. Elytra with 10 rows of small punctures, second row with WOLFGANG SCHAWALLER 380 LAENA IN THAILAND 381 about 40 punctures, punctures without seta (Fig. 23); intervals slightly convex, shagreened, sparsely scattered with punctures as on pronotum, each with a long erect seta as on pronotum; interval III with a distinct setiferous umbilicate pore at the tip and interval IX with 4 setiferous umbilicate pores between the shoulders and the tip. Legs without peculiarities. Aedeagus Fig. 24. Body length 4.8-6.8 mm. Remarks: The shape of the parameres of the above listed material (Fig. 24) is somewhat different in comparison to the figures given by Masumoto (1989: Figs 3-4), nevertheless I am convinced of the conspecificity of both series. DISTRIBUTION: Known only from the type locality Doi Inthanon in northwestern Thailand. KEY TO THE LAENA SPECIES FROM THAILAND IL: All femora or only profemur at the inner side with | or 2 spines ............ D - MGLemMOnanwAthOUly SPINES AMPLES a ee Re Eee 5 De Only profemur with a spine at the inner side, metatibia with a spine at the innersside, elytralantesval IXedistncthy ridged] 22. uam: fangensis - All femora with | or 2 spines, metatibia without spines, elytral intervals equal. 3 3: All tibiae strongly curved, pronotum with rough punctation, punctures pantlyzcontluentinsbody.lensth40 3 man Eee ee eee + - All tibiae not distinctly curved, pronotum with sparser punctation with shiny spaces between the punctures, body length 7.5 mm ..... angkhangensis 4. All femora on the ventral side with a single spine on the internal margin, parameres lone: (REATO Ae en A tos. RE siamica - All femora on the ventral side with 2 spines situated on the internal and external margin, parameres triangular (Fig. 17).......... pseudosiamica sp.n. I: Elytra scattered with large punctures, not forming distinct rows and intervals SE OR ISO e e e ES OV burckhardti sp.n. - Blymaswithel0rdisunewrowslofpuneturese Dre RO 6 6. Elytra without any setation, shape of elytra nearly round.......... thailandica - Elytral intervals and sometimes also the punctures of the elytral rows withWlonessetaesshaperorelytralonerandeoyal 2 ir E 7 % Elytral intervals and sometimes also the punctures of the elytral rows WIHBIONP:Se (Ae Re RO te tie PR RR ll ge SOI 8 = Onilythevely(tralminternvals withilonessetac PE PP RR RR RE 9 Fics 1-24 Laena species from Thailand: dorsal view of head, pronotum and elytra; punctation and setation of elytra near sutura; aedeagus from lateral and parameres from dorsal. -- 1-3: angkhangensis, holotype male; 4-6: burckhardti sp.n., holotype male; 7-9: fangensis, holotype male; 10-11: loebli sp.n., holotype female; 12-14: masumotoi sp.n., holotype male; 15-17: pseudosiamica sp.n., holotype male; 18: siamica, redrawn after KASZAB 1973; 19-21: schwendingeri sp.n., holotype male; 22-24: uenoi, male. -- Scale: 4 mm (dorsal view), 2 mm (punctation and aedeagus). 382 WOLFGANG SCHAWALLER 8. Pronotum with the basal margin of pronotum distinctly deeper than disk, punctures of pronotum smaller and distinctly more scattered than on héadtbodyileneth 4S O(N 6 cas ec056040000 000000000 schwendingeri sp.n. - Pronotum with the basal margin on the same level as disk, punctation of pronotum as on head, body length 2.8-4.0 mm .............. masumotoi Sp.n. OR Lateral and basal margins of pronotum distinctly bordered, disk of pro- notum only with scattered punctation, propleures without punctation, bodyAlensthi4:8-6.8 mum. ee ere oe LIE uenoi - Lateral margin of pronotum only with a fine border, basal margin unbor- dered, disk of pronotum with dense punctation, propleures with same punctation aston disk body lensth’8. 57mm... 2. 2 ne ee loebli sp.n. ACKNOWLEDGMENTS For the loan of this interesting material and for the possibility to keep some duplicates for the museum in Stuttgart I thank Dr Ivan Löbl (Geneva). Dr S. Nomura kindly loaned type material for comparison from the National Science Museum (Tokyo) and last, but not least, Dr K. Masumoto (Tokyo) contributed with some helpful comments. REFERENCES KASZAB, Z. 1973. Tenebrioniden (Coleoptera) aus Nepal. Acta zoologica Academiae scientarum hungaricae 19: 23-74. KASZAB, Z. 1977. Tenebrionidae der Nepal-Expeditionen von Dr. J. Martens (1969-1974) (Insecta: Coleoptera). Senckenbergiana biologica 57: 241-283. KASZAB, Z. & CHuJO, M. 1966. Coleoptera from southeast Asia V. Family Tenebrionidae. Memoirs of the Faculty of Education, Kagawa University 2 (140): 51-56. MASUMOTO, K. 1989. A new Laena (Coleoptera, Tenebrionidae) from northwest Thailand. Elytra 17: 61-64. MASUMOTO, K. 1995. A new Laena (Coleoptera, Tenebrionidae) from northern Vietnam. Bulletin of the national Science Museum Tokyo (A) 21: 33-36. MASUMOTO, K. 1996a. New tenebrionid beetles of the tribes Strongyliini, Misolampini and Adeliini (Coleoptera) from northern Vietnam. Bulletin of the national Science Museum Tokyo (A) 22: 33-43. MASUMOTO, K. 1996b. Fourteen new Laena (Coleoptera, Tenebrionidae) from China, Vietnam and Thailand. Bulletin of the national Science Museum Tokyo (A) 22: 165-187. SCHAWALLER, W. 1995a. Revision der Laena-Arten Mittelasiens (Insecta, Coleoptera, Tene- brionidae). Spixiana 18: 65-73. SCHAWALLER, W. 1995b. Neue Laena-Arten (Coleoptera: Tenebrionidae) aus Malaysia. Stutt- garter Beiträge zur Naturkunde (A) 523: 1-16. SCHUSTER, A. 1926. Bestimmungstabelle der Laena-Arten aus dem Himalaya und den angren- zenden Gebieten. Mit Beschreibungen neuer Arten. Koleopterologische Rundschau 12: 31-54. SCHUSTER, A. 1935. Neue Laena-Arten aus dem Himalaya (Col., Fam. Tenebrionidae). Annals and Magazine of natural History (ser. 10) 16: 437-466. REVUE SUISSE DE ZOOLOGIE 105 (2): 383-394; juin 1998 Die Gattung Stenus Latreille in Vietnam (Coleoptera, Staphylinidae)* Volker PUTHZ c/o Limnologische Fluss- Station des Max- Planck- Instituts für Limnologie Damenweg 1, D-36110 Schlitz, BRD. The genus Stenus Latreille in Vietnam (Coleoptera, Staphylinidae). - List of 67 Stenus- species known from Vietnam including descriptions of 3 new species: Stenus (Hypostenus) voluptabilis sp. n., S. (Parastenus) perrotianus sp. n., S. (P.) unguliventris sp. n.), new synonyms (S. Para- stenus) horridulus Ryvkin, 1992 and S. (P.) unguinosus Ryvkin, 1992 = S. (Hypostenus) deliculus Ryvkin, 1992; S. (P.) mangpuensis Cameron, 1943 = S. (P.) salebrosus L. Benick, 1942), and new species- records. The shape of paraglossae is used for the first time as a taxonomic character. Key-words: Coleoptera - Staphylinidae - Stenus - Vietnam - Taxonomy - paraglossa. EINLEITUNG Die letzte Zusammenstellung der Stenus- Arten Vietnams ist 1981 erfolgt, in der Zwischenzeit kamen einige Arten hinzu. Das Naturhistorische Museum Genf erhielt durch den Ankauf der Sammlung Curti (in der sich coll. Ochs und coll. Perrot befinden) älteres Material aus Vietnam, das mir zur Bearbeitung vorgelegt wurde. Darin fanden sich drei neue Arten, die im Folgenden beschrieben werden, eine Serie von Stenus deliculus Ryvkin, dessen Männchen bisher unbekannt war und einige Erstnachweise. Ich nehme deshalb die Gelegenheit wahr, den gegenwärtigen Kennt- nisstand der Stenus- Fauna dieses ostasiatischen Landes festzuhalten und fiige noch eine weitere Synonymie hinzu, die ich jiingst herausfand. Insgesamt liegen aus Vietnam jetzt 67 Stenus- Arten und - Rassen vor. Zum ersten Mal verwende ich in dieser Arbeit ein bisher unbeachtetes Merkmal für die Taxonomie: die Gestalt der Paraglossen (vgl. u. und Figs 8, 9). - Als Abkürzungen gelten die in meinen früheren Arbeiten gebrauchten. Stenud (Hypostenus) voluptabilis sp. n. Diese neue Art erinnert auf den ersten Blick an Vertreter der Spezies um S. amoenus L. Benick, ist aber mit diesen nicht eng verwandt. Wegen ihres Kopfbaues * 251. Beitrag zur Kenntnis der Steninen. Manuskript angenommen an 27.08.1997 384 VOLKER PUTHZ und wegen ihres lateral spitzen 9. Sternums steht sie - soweit ich sehe - isoliert, ich kann keinen orientalischen Stenus nennen, der eine ganz ähnliche Gestalt besässe. Glänzend, schwarz, Elytren mit einer grossen orange- gelben Makel, Vorder- körper ziemlich grob und dicht, Abdomen etwas weniger grob und mässig dicht punktiert; Beborstung kurz, anliegend. Fühler gelb, ab der Mitte gebräunt, Kiefer- taster gelb, Beine rötlichgelb, die Knie deutlich verdunkelt, Tarsenglieder zum Teil verdunkelt. Oberlippe braun, Clypeus und Oberlippe mässig dicht beborstet. 9. Ster- num apikolateral spitz (Fig. 1). Länge: 3,0 - 3,8 mm (Vorderkörper: 2,0-2,1 mm). ?-Holotypus: Vietnam (Tonkin): Hoa Binh, H. Perrot (MHNG). Paratypen: 2 dd, 1 2:40 km NW An Khe, Buon Luoi, 620-750 m, 14°10’N, 108°30’E, 28.3.-12.4.1995, Pacholatko & Dembicky (NHMW, coll. Puthz). Vermutlich gehört hierher auch das £ von Tam Dao, da ich 1981 (11) als “spec. amoenus- Gruppe” gemeldet habe, das mir aber jetzt nicht vorliegt (Museum Leningrad). PM des HT: Kbr.: 33; AA: 16,5; Pbr: 24,5; Plg: 30; Ebr: 37; Elg: 40; Nlg: 33. Kopf deutlich schmäler als die Elytren, Stirn vergleichsweise breit, mit zwei deutlichen Längsfurchen und einem spiegelglatten, rundlich erhobenen Mittelteil, dieser hinten so breit wie jedes der Seitenstiicke; Stirn insgesamt mit 5 Spiegelflecken, die Punktierung im übrigen ziemlich grob und dicht, mittlerer Punktdurchmesser so gross wie der basale Querschnitt des 3. Fühlergliedes. Fühler mässig lang, zurückgelegt mit dem 11. Glied den Pronotumhinterrand überragend, vorletzte Glieder etwa doppelt so lang wie breit. Pronotum in der vorderen Mitte am breitesten, daselbst eine kurze Strecke fast parallelseitig, vorn schräg eingezogen, hinten konkav- eingeschnürt verengt; Oberfläche gleichmässig gewölbt und ganz gleichmässig grob und dicht punktiert, mittlerer Punktdurchmesser so gross wie der apikale Querschnitt des 3. Fühlergliedes, Punktabstände fast überall kleiner als die Punktradien. Elytren etwa quadratisch, mit deutlichen Schultern, hinten rundlich eingezogen, Naht- und Schulter- eindrücke flach; die grosse organge- gelbe Makel ist etwas kürzer als die halbe Elytrenlänge (18: 20), vom Vorderrand etwa um 2/3 ihrer Länge, vom Hinterrand gut um 1/3 ihrer Länge getrennt, innen erreicht sie fast die Naht, aussen erstreckt sie sich bis auf den Deckenabfall, ohne den Seitenrand zu erreichen; die Scheibenpunktierung der Elytren ist deutlich etwas gröber als diejenige des Pronotums, Punktabstände wenig grösser. Abdomen nach hinten deutlich verschmälert, basale Quereinschnürungen der ersten Segmente sehr tief, 7. Tergit mit breitem, apikalem Hautsaum (= makroptere Art); Punktierung vom ziemlich grob und ziemlich dicht, nach hinten feiner und weitläufiger, auf dem 6. Tergit sind die Punkte so gross wie eine mittlere Augenfacette, ihre Abstände 1,5 - 2 x so gross wie die Punkte, das 10. Tergit trägt mehrere, mässig feine Punkte. Beine schlank, Hintertarsen 3/5 schienenlang, 1. Glied deutlich länger als die beiden folgenden zusammen mittlere Glieder breit gelappt. Nur das 8. Tergit an der Basıs mit Netzungsspuren, die restliche Oberfläche spiegelglatt. MANNCHEN: Vordersternite grob und dicht punktiert, 7. Sternit in der hinteren Mitte fein und ziemlich dicht punktiert und beborstet, nicht eingedrückt. 8. Sternit mit dreieckigem Ausschnitt etwa im hinteren Fünftel. 9. Sternit (Fig. 1), apikolateral mit STENUS IN VIETNAM 385 spitzem Zahn und feinem, langen Borstenpinsel. 10. Tergit breit abgerundet. A e do e - agus (Fig. 2), Medianlobus gerundet- konisch verengt, Apikalpartie ventral mit zwei Zähnchenleisten, Basalteil im Innern mit zwei diinnen Versteifungsleisten (Ausstülphilfen) und einem breittubigen Innensack; Parameren gut so lang wie der Medianlobus, apikal leicht löffelförmig und daselbst mit zahlreichen langen Borsten versehen. WEIBCHEN: 8. Sternit breit abgerundet. Valvifer alikal abgestutzt mit kurzem Lateralzahn und dünnem Borstenpinsel. 10. Tergit breit abgerundet. Stenus voluptabilis sp. n. - “der, der Vergnügen macht” - unterscheidet sich durch die Kombination der Merkmale: erhobene, geglättete Stirnmitte, Kurze, unauf- fällige Beborstung und die Gestalt des 9. Sternums sowie die des 10. Tergits von allen orientalischen Arten; von den wegen ihrer Makeln allenfalls ähnlichen Vertretern um S. amoenus L. Benick (S. alumoenus Rougemont, S. amenous Rougemont, S. amoe- nulus Puthz und S. aneomus Rougemont, vermutlich auch von S. chlorostigma L. Benick) durch geringere Grösse und den Kopfbau: diese verglichenen Arten besitzen grössere Augen, ihre Stirnmitte ist relativ schmäler und ganz flach, die Stirn zeigt keine auffälligen 5 Spiegelflecken. Durch grössere Elytrenmakel und den Stirnbau lässt sich die neue Art auch leicht von S. changi Puthz und S. delectus Puthz trennen. Stenus (Hypostenus) deliculus Ryvkin Stenus (Parastenus) deliculus Ryvkin, 1992: 51 ff. figs. Stenus (Parastenus) horridulus Ryvkin, l.c.: 45 ff. figs, syn. n. Stenus (Parastenus) unguinosus Ryvkin, l.c.: 49 ff. figs, syn. n. Diese Art gehört zu den abweichenden (Hypostenus-) bzw. (Parastenus-) Arten mit ausserordentlich langer, abstehender Beborstung und unterschiedlich starker Seitenrandung des Abdomens. Solange keine phylogenetische Analyse der Gattung Stenus vorliegt, stelle ich diejenigen Arten dieser monophyletischen Gruppe, die eine vollständige, wenn auch oft nur linienförmig- schmale Seitenrandung des Abdomens haben, zu Parastenus und nenne den Komplex “cirrus”- Komplex, diejenigen Spezies aber, bei denen die abdominale Seitenrandung nur am 3. Segment und dann wieder als Schnittlinie vom 7. Segment an auftritt, bei denen also die Segmente 4-6 ohne jede Spur einer Seitenrandung sind, stelle ich - per definitionem - zu Hypostenus und nenne diesen Komplex ab jetzt den “falsus”- Komplex. Stenus deliculus Ryvkin gehört in den falsus- Komplex. Für die Beschreibung seiner drei Taxa hat der genannte Autor 7 Weibchen zur Verfügung gehabt, von denen 5 von Tam Dao stammen. In der Sammlung OCHS fanden sich nun 4 8 8,4 ? © ebenfalls von Vietnam (Tonkin): Tam Dao, H. Perrot, die ich mit den Holotypen von 5. horridulus und S. deliculus sowie einem Paratypus von S. unguinosus ver- gleichen konnte. Die von RYVKIN präparierten Spermatheken liegen in den Präparaten verschieden und sind bei S. deliculus zerbrochen. Die unterschiedliche Kontur des 8. Sternits und der Valvifera (l.c. S. 48) liegen im Rahmen der Variationsbreite. Die in 386 VOLKER PUTHZ den Beschreibungen bemerkten Unterschiede in Bezug auf die Mikroskulptur sind nicht vorhanden: es handelte sich bei dem Material aus Moskau um verschmutzte Exemplare! Und was die Punktierungsunterschiede angeht, so finde ich solche auch in der mir vorliegenden Serie von Tam Dao. Als erster Revisor wähle ich den Namen S. deliculus als denjenigen, den diese Spezies kiinftig tragen soll, die beiden anderen Taxa werden eingezogen. PM eines d von Tam Dao: Kbr: 32,5; AA: 15,5; Pbr: 23; Plg: 25; Ebr: 27; Elg: 29 [bei anderen Stücken sind die Elytren so lang wie breit]; Nlg: 22. Vorder- körperlänge: 1,8 mm, Gesamtlänge: 3,2-4,0 mm (stark ausgezogen bis 4,5 mm). MÄNNCHEN: Vordersternite ohne besondere Merkmale, 7. Sternit median sehr fein und viel dichter als an den Seiten punktiert und beborstet. 8. Sternit mit flach- runder Apikalausrandung etwa im hinteren Zweiundzwanzigstel (3: 67). 9. Sternit (Fig. 11). 10. Tergit breit abgerundet. Aedoeagus (Fig. 12) mit gerundeter Api- kalpartie, umfangreichen, stark sklerotisierten Ausstülphaken und breittubigem Innen- sack. WEIBCHEN: 8. Sternit zur Hinterrandmitte kaum vorgezogen, breit abgerundet. Valvifer apikolateral spitz. Spermatheka (Fig. 5; aber je nach Lage auch andere Gestalten). 10. Tergit breit abgerundet. Stenus deliculus Ryvkin unterscheidet sich von den ungemakelten Arten des falsus- Komplexes wie folgt: Vom ebenfalls sehr grob punktierten, glänzenden S. splendidulus Puthz durch längere Elytren and vorn erheblich gröber punktiertes Abdomen, von S. punctifer Naomi durch breiteren Kopf sowie glänzenderen, gröber punktierten Vorderkörper, von S. falsus L. Benick (dessen Aedoeagus ich hier erst- malig abbilde: Fig. 10) und S. amamiensis Naomi durch erheblich grôbere Vorder- körperpunktierung, viel gröber punktiertes Abdomen und (meist) längere Elytren, von allen durch den Aedeoagus. Stenus (Parastenus) perrotianus sp. n. Diese neue Art ist die Schwesterart des aus Indien beschriebenen S. pseudopic- tus Cameron, einer schlanken Art mit ausserordentlich langen Fühlern. Die äusseren Unterschiede sind gering, aber die Spermatheken differieren so deutlich, dass ich dieses einzige Weibchen für eine neue Art halten muss. Schwarz mit Metallschimmer, jede Elytre mit grosser, ovaler, schräger, orange- gelber Makel in der hinteren Aussenhälfte. Punktierung der Stirn grob und sehr dicht, die des Pronotums grob, sehr dicht, leicht rugos, diejenige der Elytren grob, ebenfalls sehr dicht, in der Aussenhälfte längs- zusammenfliessend. Abdomen ziemlich grob und dicht, aber nicht gedrängt punktiert. Fühler gelb, die Keule wenig dunkler. Kiefertaster gelb. Paraglossen oval (vgl. u.). Beine einfarbig rötlichgelb. Oberlippe schwarzbraun, heller gesäumt, Clypeus und Oberlippe wenig dicht beborstet. Länge: 6,2 - 7,3 mm (Vorderkörper: 3,6 mm). ® - Holotypus: Vietnam (Tonkin): Tam Dao, H. Perrot (MHNG). PM des HT: Kbr: 49; AA: 27; Pbr: 34; Plg: 44; Ebr: 50; Elg: 56; Nlg: 43. Hin- tertarsen: Hinterschienen = 46: 65, Hintertarsen: 22- 10- 6- 7- 8. STENUS IN VIETNAM 387 MANNCHEN: unbekannt. WEIBCHEN: 8. Sternit am Hinterrand deutlich rund vorgezogen, die Seiten davor leicht konkav. Valvifer mit langem Apikolateralzahn (Fig. 7). Spermatheka (Fig. 7), die langen Schläuche mit 5 Windungen. 10. Tergit breit abgerundet. Stirn wie bei S. pseudopictus, ihre Mitte mit flacher, aber deutlicher Erhebung, die hinten verflacht und insgesamt unterhalb des Augeninnenrandniveaus liegt; Punkte so gross wie der mittlere Querschnitt des 3. Fühlergliedes, Punktzwischen- räume fast überall viel kleiner als die Punktradien, keine Glättungen. Fühler sehr lang, zurückgelegt überragen mehr als die letzten 4 Glieder den Pronotumhinterrand, das 9. Glied ist dreimal, das 10. doppelt so lang wie breit. Pronotum erheblich länger als breit mit deutlicher Quereinschniirung kurz hinter der Mitte, die Seiten nach vorn fast gerade verengt, nach hinten konkav verengt; in der Vorder- und in der Hinterhälfte einige Längseindrücke: insgesamt erscheint das Pronotum also bemerkenswert (wenn auch nicht sehr tief) uneben; Punktierung durchschnittlich so grob wie auf der Stirn, aber nicht so gleichmässig grob und, obwohl überall sehr dicht, auch unterschiedlich dicht, in der Nähe des Vorder- und des Hinterrandes auch leicht zusammenfliessend. Elytren seitlich lang gerundet, uneben: Nahteindruck tief, je ein mittlerer und ein hinterer Eindruck ziehen sich von der Naht schräg nach aussen, der Schultereindruck ist weniger auffallig; die schrägstehende ovale Makel im hinteren Aussenviertel der Elytren ist so lang wie das 3. Fühlerglied und etwa so breit wie das 4. Fiihlerglied; Punktierung gròber als am Pronotum, in der Innenhälfte dicht, aber überwiegend getrennt, in der Aussenhälfte, vor allem um die Elytrenmakel herum, lang zusammen- fliessend. Abdomen mit schmalen, ventrad abfallenden Paratergiten, diejenigen des 4. Segmentes fast so breit wie das 1. Hintertarsenglied (also schmäler als das 2. Fiihle- glied), mit einer Reihe feiner, wenig dicht stehender Punkte versehen; 7. Tergit mit apikalem Hautsaum (= makroptere Art); die Punktierung ist deutlich dichter als bei S. pseudopictus, so sind zum Beispiel die Punktabstände in der Mitte des 3. Tergits iberall kleiner als die Punkte (bei der verglichenen Art mehrfach so gross wie oder deutlich grösser als diese), auf dem 6. Tergit sind die Punktabstände höchstens 1,5 mal so gross wie die Punkte (bei S. pseudopictus oft doppelt so gross). Nur das 4. Tarsenglied ist lang und schmal gelappt. Die ganze Oberseite ist deutlich genetzt. Von S. pseudopictus Cameron unterscheidet sich die neue Art auch durch ihre Spermatheka, die 5 statt 2 Windungen aufweist (vgl. Fig. 6). Da es in der Orientalis zahlreiche prinzipiell ähnliche, gemakelte Arten gibt, möchte ich ausführlich angeben, wie sich diese neue Art von ihnen unterscheidet. Dabei verwende ich hier erstmalig ein Merkmal, das bisher taxonomisch noch nicht berücksichtigt worden ist: die Form der Paraglossen. In seinen beispielhaften Arbeiten hat O. BETZ diese zu Klebpolstern umgestalteten Paraglossen von zahlreichen Stenus- Arten untersucht und dabei einen sogenannten (ovalen) “Grundtyp” (Fig. 8) von anderen unterschieden. Unter den orientalischen Parastenen besitzen nun zahlreiche Arten einen auffällig vom Grundtyp abweichenden spitzkegeligen (koniformen) Typ, den BETZ von S. cf. luteolunatus publiziert hat (Fig. 9). Dies ist bisher die einzige Spezies, von der dieser besondere Paraglossenbau mitgeteilt worden ist. Ich habe nun festgestellt, daB sich anhand der beiden genannten Paraglossentypen (ich will sie ab 388 VOLKER PUTHZ jetzt mit den Begriffen “oval” und “koniform” bezeichnen) auch an trockenem Mate- rial die orientalischen Parastenen leicht in zwei Gruppen unterscheiden lassen, Gruppen, in denen äusserlich sonst recht ähnliche Spezies stehen. Man muss aller- dings, um dieses Merkmal festzustellen, den Kopf von der Unterseite betrachten und - in seltenen Fällen - sogar die “Zunge” (das Praementum) herauspräparieren, was leicht gelingt, wenn man bei gekochtem Material den Kopf abtrennt, ihn mit der einen Nadel in der hinteren Offnung festhält, mit der anderen Nadel hinter dem Mentum einsticht und Mentum + Praementum nach vorn herauszieht. Bisher konnte ich noch nicht alle orientalischen Arten auf dieses Merkmal hin untersuchen. Was aber nun die Unterscheidung der hier beschriebenen neuen Art von anderen, relativ grossen, gemakelten Arten angeht, so lassen sich erst einmal die folgenden Spezies wegen ihrer koniformen Paraglossen klar von S. perrotianus trennen: S. abdominalis Fauvel, S. bicolon Sharp, S. bilunatus Puthz, S. biplagiatus Puthz, S. brachati Puthz, S. chakratianus Cameron, S. cham Puthz, S. commaculatus Puthz, S. coronatus L. Benick, S. diversus, L. Benick, S. jenisi Hromadka, S. languor L. Benick, S. leileri Puthz, S. lopchuensis Cameron, S. luteonotatus Puthz, S. luteo- maculatus Rougemont, S. malabarensis Cameron, S. rafflesi Rougemont, S. rouge- monti Puthz (die Subspezies einzelner Arten eingeschlossen). Von S. diversus L. Benick unterscheidet sich die neue Art aussenden durch feiner und nicht gedrängt-dicht punktiertes Abdomen, vor allem durch feiner punktierte Paratergite, von S. invocatus Ryvkin (der ebenfalls von Tam Dao be- schrieben wurde, mir bisher aber nicht vorgelegen hat) durch viel längere Fühler (diese sollen bei S. invocatus, zurückgelegt, den Pronotumhinterrand kaum über- ragen), durch erheblich breiteren Kopf und durch bedeutendere Grösse, von den folgenden Arten, die alle den ovalen Paraglossentyp besitzen, so: von S. subthora- cicus Rougemont durch schmälere, nur in einer Reihe punktierte Paratergite und viel grössere Elytren mit relativ kleinerer Makel, von S. signatipennis Puthz durch viel längere Fühler, von S. miwai Cameron durch feiner punktiertes, längeres Pronotum und breitere Paratergite, von S. grandimaculatus L. Benick durch längeres Pronotum und breitere Paratergite. Stenus (Parastenus) unguliventris sp. n. Diese neue Art ist die Schwesterart des S. habropus Puthz, dem sie zum Verwechseln ahntich sieht; eine ausführliche Beschreibung hätte deshalb nur wieder- holenden Charakter. Ich beschränke mich also auf das Wesentliche. Schwarz, metallisch schimmernd, Kopf grob und wenig dicht punktiert, Pronotum sehr grob und sehr dicht, leicht rugos punktiert, Elytren mit sehr grober, Fics 1-7 9. Sternit (1) Ventralansichten der Aedoeagi (2, 3), Valvifer (4, 7) und Spermatheken (5- 7) von Stenus (Hypostenus) voluptabilis sp. n. (PT, 1, 2), S. (Parastenus) unguliventris sp. n. (PTT, 3, 4), S. (H.) deliculus Ryvkin (5), S. (P.) pseudopictus Cameron (PT, 6) und S. (P.) perrotianus sp. n. (HT, 7).- Mass- Stab = 0,1 mm (1= 2, 5; 3= 4, 6, 7). 389 STENUS IN VIETNAM 390 VOLKER PUTHZ stark rugoser Skulptur (die Punkte sind in langen, gedrehten Furchen verbunden). Abdomen mässig grob bis fein und ziemlich weitläufig punktiert. Die ganze Oberseite dicht genetzt. Beborstung sehr kurz, anliegend, kaum auffällig. Fühler gelblich, zur Spitze etwas gebräunt, Kiefertaster gelb, Beine rötlichgelb, Schenkelspitzen und Tarsengliedspitzen etwas dunkler. Clypeus und Oberlippe schwarz (-braun), wenig dicht beborstet. Paraglossen oval. 10. Tergit in beiden Geschlechtern breit abgerundet. Länge: 6,5 - 7,5 mm (Vorderkörper: 3,6 mm). 4 - Holotypus und 1 ®, 2 8 d- Paratypen: Vietnam (Tonkin): Tam Dao, H. Perrot (MHNG, coll. Puthz). 1 ¢: Indien: Meghalaya: W Garo Hills, Nokrek N.P., ca. 1100 m, 25° 29,6’ N, 90° 9,5’ E, 9.-17.5.1996, Jendek & Sousa (NHMW). PM des HT: Kbr: 50.5; AA: 26; Pbr: 37; Plg: 40; Ebr: 51; Elg: 57; Nlg: 44. Hintertarsen: Hinterschienen = 62: 68. Hintertarsen: 28- 12- 8- 9- 11. Bei 9- Para- typen kann der Kopf deutlich breiter als die Elytren sein! MANNCHEN: Schenkel gekeult, Hinterbeine mit Trochanterzahn, Mittelschienen mit starkem Apikaldorn, Hinterschienen mit schwachem Präapikaldorn. Metasternum breit-dreieckig eingedrückt/ abgeflacht, wenig grob und mässig dicht auf flach genetztem Grund punktiert, vordere Mitte (zwischen den Mittelhiiften) plattenartig- abgesetzt erhoben und daselbst dicht skulptiert und lang beborstet, Hinterhüftum- randungen glatt. 4. Sternit median leicht abgeflacht, 5. Sternit in der hinteren Mitte flach eingedrückt, 6. Sternit daselbst mit tiefem Eindruck, Hinterrand flach aus- gerandet, 7. Sternit mit langem Mitteleindruck, dieser proximal tiefer als distal, dichter als an den Seiten punktiert und beborstet, Hinterrand flach ausgerandet. 8. Sternit mit schmalem Apikalausschnitt im hinteren Viertel. 9. Sternit apikolateral mit sehr langem, krallenförmig leicht ventrad gebogenem Zahn (Name !). Aedoeagus (Fig. 3), Medianlobus mit knopfförmiger Spitze, Apikalpartie mit zwei kräftigen Ventralzähnen, Innenkörper aus membranösen Strukturen bestehend. WEIBCHEN: Metasternum abgeflacht, ohne auffallende Merkmale zwischen den Mittelhüften. 8. Sternit am Hinterrand breit und stumpf vorgezogen. Valvifer mit krallenförmigem Apikalzahn (Fig. 4). Keine sklerotisierte Spermatheka, jedoch mit einem länglichen, plattenartigen Sklerit in Höhe der Valvifer. Von Stenus habropus Puthz lässt sich die neue Art sicher nur durch den Aedoeagus trennen. Äusserlich unterscheidet sie sich von ihm durch breiteren Kopf, weniger grobe und weniger dichte Stirnpunktierung und durch etwas feiner und weitläufiger punktiertes Abdomen. Da jedoch über die Variationsbreite dieser Arten zu wenig bekannt ist, müssen diese Skulpturunterschiede mit Vorsicht betrachtet werden, zumal auch schon in der Typenserie hier bemerkbare Unterschiede auftreten.- Das oben genannte Weibchen aus den Garo Hills, Indien, kann ich nicht sicher von der neuen Art unterscheiden und stelle es deshalb mit Vorbehalt zu ihr. Das von mir 1981 aus Yunnan gemeldete Weibchen (Museum Leningrad) liegt mit jetzt nicht vor; auch hier kann nicht ausgeschlossen werden, dass es zu S. unguliventris gehört. - Von anderen (Parastenus-) Arten mit linienartig- schmal gerandetem Abdomen und spinnendünnen Extremitäten lässt sich die neue Art durch ihre Grösse, fehlende Elytrenmakeln, weitläufige Abdomenpunktierung und langen “krallenförmigen” Api- kolateralzahn des 9. Sternums trennen. Bei S. rugosipennis Cameron und noch STENUS IN VIETNAM 391 473% 15KV WD: 38MM S:25692 P:00007 Di 15KY WD:16MM 5:26793 P:69992 186UM—________________——--++»y9 NU ai n Fics 8-9 Paraglossen von Stenus: ovaler Typ (8, Stenus comma LeConte, Blick von der Seite auf die Spitze des Praementums; unterhalb der Paraglossen die Labialpalpen) und koniformer Typ (9, S. cf. luteolunatus Puthz, Blick auf die Spitze des Praementums mit Paraglossen und Labial- palpen). Die REM- Aufnahmen wurden mir liebenswiirdig von Dr O. Betz, Kiel, zur Verfiigung gestellt, wofür ich auch an dieser Stelle herzlich danke. unbeschriebenen taiwanesischen Arten ist dieser Zahn weniger auffällig und erheblich kürzer; diese Arten besitzen auch eine sklerotisierte Spermatheka. Nicht so ist das aber bei den beiden näher verwandten, aber viel kleineren S. guenai Rougemont und S. calcariventris Puthz, bei denen das Metasternum zwischen den Hinterhiiften eben- falls, jedoch nicht erhoben- abgesetzt, dicht skulptiert ist; beiden Arten fehlt ebenfalls eine sklerotisierte Spermatheka. Stenus (Parastenus) salebrosus L. Benick Stenus salebrosus L. Benick, 1942: 25 ff. Stenus mangpuensis Cameron, 1943: 3 syn. n.; PUTHZ 1978: 130. Von beiden Taxa konnte ich die Typen, darunter männliche Stücke, untersuchen und feststellen, dass sie konspezifisch sind. S. salebrosus war aus Burma beschrieben, S. mangpuensis aus Indien, Darjeeling District. Ausser den schon mitgeteilten Stiicken sah ich inzwischen noch: | © Sikkim: Lacher 9000’, from moss, leaf mould, earth. Pine wood, 29.2.1952, T. Clay (BMNH); 1 d China: Yunnan: 100 km W Baoshan, Gaoliongshan N. Res., 14.-21.6.1993, Jendek & Sousa (NHMW): neu für China! Die Typen unterscheiden sich etwas durch den Grad der Kurzfliigligkeit, die Stiicke aus Darjeeling zeigen eher trapezoide Elytren, die aus Burma eher recht- eckige; dies ist aber nur Ausdruck der Variabilität. S. salebrosus gehört zu den Arten mit koniformen Paraglossen. 392 VOLKER PUTHZ = 4 = || = I = A SS A, 10 11 Fics 10-12 Ventralansichten der Aedoeagi und 9. Sternit des Männchens: Stenus (Hypostenus) falsus L Benick (Chinkiang, 10), S. (H.) deliculus Ryvkin (Tam Dao, 11, 12). Mass- Stab = 0,1 mm. LISTE DER BISHER AUS VIETNAM BEKANNTEN Stenus- ARTEN Sofern nicht in Klammern anders angegeben, sind die aufgezählten Taxa schon in meinen Arbeiten von 1981 und 1983 genannt. Ich gebe nur bei solchen Taxa Fundorte an, bei denen es sich um Erstnachweise fiir Vietnam handelt. Von den übrigen Arten liegen durch die Aufsammlungen Perrots zahlreiche weitere Funde vor, die ich hier nicht verzeichne. Stenus s.str. + Nestus S. formosanus L. Benick . insignatus Puthz louwerensi Cameron . megacephalus Cameron (inkl. spec. À, PUTHZ 1980) melanarius annamita Fauvel . puberulus fukiensis L. Benick venator Fauvel: neu! 1 ® S. Vietnam: 14 km SW Bao Loc, 16-29.5.1994, Pacholatko & Dembicky (NHMW). spec. B. (rugicollis- Gruppe): von beiden Arten sind immer noch keine Männchen bekannt. LA La La La La La a STENUS IN VIETNAM 393 Hypostenus S. S. S. S. La La La La La La La La La La La La La La La La Ca La La La La La La La La La amoenulus Puthz amoenus L. Benick angusticollis Eppelsheim arachnoides Bernhauer: Neu! 1 d: Da Lat, cam Ly area, rainforest, swept and Beaten, No. 691, 4.12.1994, S. Mahunka et al. (Museum Budapest). Dieses Männchen lässt sich geni- taliter und skulpturell nicht vom javanischen S. arachnoides unterscheiden; die äusseren Körperumrisse sind jedoch etwas anders: der Kopf ist so breit wie die Elytren, während er bei javanischen Exemplaren meist deutlich breiter ist als dieselben (z.B. 39,5 : 35). Es kann nicht ausgeschlossen werden, dass auch enge Beziehungen zum Komplex um S. velox Sharp bestehen, doch liegt zur Klärung dieser Frage zur Zeit noch nicht genügend Material vor. . basicornis Kraatz . basicornis subtropicus Cameron . bicuspis Puthz . bispinoides Puthz (PUTHZ 1985) . cicindeloides (Schaller) . chlorostigma L. Benick . currax Sharp . cursorius L. Benick (RYVKIN 1992) . deliculus Ryvkin (RYVKIN 1992) . echiniventris Puthz . elegantulus Cameron . fistulosus L. Benick . flavidulus paederinus Champion . flavittatus Champion . flavovittatus obliteratus Cameron . flavovittatus sinuatus Cameron: Neu! 1 4 (Tonkin) Thank Moi, Perrot; 1 d Bac Kan, Perrot (MHNG); 1 9 N. Vietnam: Sapa (Lao Cai) 22°20’N, 103°50’E, 26.5-10.6, Horak (NHMW). S. frater L. Benick Si S. hirtellus Sharp: neu! 1 d, 1 9: Saigon, 12.11. und 13.12.1950, J. Barbier (MNP); 1 d gastralis Fauvel Hanoi, Perrot; 3 2 2: Hoa Bink, Perrot (MHNG, coll. Puthz). Bisher aus Japan und Taiwan gemeldet. . loebli Puthz: neu! 1 5, 1 2: Hanoi (Coll. Fauvel, IRScNB); 1 9: Annam) Muong Sen, Perrot (MHNG) . lorifer Puthz . vegetus Puthz: Von Tam Dao, Perrot, liegen jetzt 1 d, 2 2% vor, die meine unsichere Meldung von 1983 (als S. nitidulus Cam.) bestätigen (MHNG, coll. Puthz). . oblitus Sharp . piliferus Motschulsky . pulchrivestis Puthz (PUTHZ 1991) . pustulatus Bernhauer: neu! 1 3: (Tonkin): Kep, Perrot (MHNG). . puthzianus Rougemont: neu! 1 © (Tonkin) Yen Bay, Perrot (MNP). . spenceri Puthz . topali Puthz . tuberculicollis Cameron . verticalis L. Benick . voluptabilis sp. n. “Hemistenus” Si depressus secessus Puthz Parastenus abdominalis Fauvel belli Fauvel: neu! 1 2: (Tonkin) Kep, Perrot (MHNG); | 2: S. Vietnam: 40 km NW An Khe, Buon Luoi, 620-750 m, 14°10’N, 108°30’E, 28.3-12.4.1995, Pacholatko & Dembicky 394 VOLKER PUTHZ (NHMW). Bisher gut aus Südindien bekannt, aber auch aus Nordindien (Haldwani District) gemeldet. Die Stiicke aus Vietnam belegen eine weitere Verbreitung in der Orientalis. Diese Art besitzt übrigens ovale Paraglossen. bicolon posticus Fauvel biplagiatus Puthz: neu! 1 2 (Tonkin) Tam Dao, Perrot (MNP). cham Puthz gestroi callifrons L. Benick: neu! 1 © (Tonkin) Tam Dao, Perrot (MNP). gestroi grandiculus L. Benick gestroi stigmatipennis L. Benick invocatus Ryvkin (RYVKIN 1992) languor L. Benick (RYVKIN 1992) maculifer Cameron marginifer Puthz pallidipes Cameron perfidiosus Puthz perroti Puthz perrotianus Sp. n. signatipennis Puthz: neu! 1 & (Tonkin): Tam Dao, Perrot (MHNG) stigmaticus Fauvel tenuimargo Cameron vietnamensis Puthz virgula Fauvel unguliventris sp. n. LITERATUR BENICK, L. 1942. Entomological Results from the Swedish Expedition 1934 to Burma and British India, Coleoptera: Staphylinidae, gesammelt von René Malaise. Ark. Zool. 33 A 17: 1-48. BETZ, O. 1994. Der Fangapparat bei Stenus spp. (Coleoptera, Staphylinidae): Bau, Funktion, Evolution. Dissertation Bayreuth. BETZ, O. 1996. Function and evolution of the adhesion- capture apparatus of Stenus species (Coleoptera, Staphylinidae). Zoomorphology 116: 15-34. CAMERON, M. 1943. Description of new Staphylinidae (Coleopt.) (cont.). Proc. R. ent. Soc. Lond. (B) 12: 1-5. PUTHZ, V. 1978. Steninae. Coleoptera, Staphylinidae) aus Indien. Ann/s hist.-nat. Mus. natn. hung. 70: 123-134. PuTHz, V. 1981. Steninen aus Jünnan (China) und Vietnam (Coleoptera, Staphylinidae) 182. Beitrag zur Kenntnis der Steninen. Reichenbachia 19: 1-21. PuTHz, V. 1983. Alte und neue Steninen aus Hinterindien und China (Coleoptera, Staphy- linidae) 194. Beitrag zur Kenntnis der Steninen. Reichenbachia 21: 1-13. PUTHZ, V. 1985. Revision der Gruppe des Stenus (Hypostenus) bispinus Motschulsky (Col. Staphylinidae). Dr. ent. Z., N.F. 32: 75-100. PUTHZ, V. 1991. Über indo- australische Steninen II (Insecta, Coleoptera, Staphylinidae) 222. Beitrag zur Kenntnis der Steninen. Ent. Abh. Mus. Tierk. Dresden 54: 1-46. RYVKIN, A.B. 1992. To the fauna of the staphylinid genus Stenus (Coleoptera, Staphylinidae) of Vietnam /n: Insect systematics and ecology of Vietnam. (Medvedev, L.N. ed.) - Moscow: 41-54 (russisch). REVUE SUISSE DE ZOOLOGIE 105 (2): 395-463; juin 1998 Aleocharinae della Cina: Parte II (Coleoptera, Staphylinidae) Roberto PACE Via Vittorio Veneto 13, I-37032 Monteforte d’Alpone (Verona), Italia. Aleocharinae from China: Part II (Coleoptera, Staphylinidae). - In this paper further 67 species are described as new to science. These new species belong to following tribes: Falagriini (20 n. sp.), Deremini (2) and Athetini, part I (45). Three genera are described as new, assigned to following two tribes: Deremini (Kamptomerus n. gen. near Longiprimi- tarsus), Athetini (Enkoilogeneia n. gen. near Hydrosmecta, and Amphi- bolusa n. gen. near Amidobia). Two new combinations are proposed. The main diagnostic caracters are illustrated. Key-words: Coleoptera - Staphylinidae - Aleocharinae - Taxonomy - China. INTRODUZIONE Sono qui descritti i taxa ritenuti nuovi per la scienza raccolti prevalentemente dal Dr Ales Smetana del “Centre for Land and Biological Resources Research” di Ottawa e dal collega stafilinidologo Guillaume de Rougemont di Londra in occasione di lunghi viaggi o soggiorni in Cina. Le specie note o nuove per la Cina sono elencate nella prima parte di questa stessa serie (PACE 1998). Le nuove specie appartenengono alle tribù Falagriini, Deremini e Athetini. Specie di quest’ultima tribù saranno descritte anche e in numero maggiore nella terza parte della presente serie di articoli. La trattazione separata in due delle specie della tribù Athetini si è resa necessaria al fine di non aumentare il numero di pagine e di tavole, sì da rendere il lavoro difficilmente pubblicabile. I disegni delle tavole sono stati da me eseguiti in tutte le loro fasi. Gli holotypi delle nuove specie sono depositati nel Museo di Storia Naturale di Ginevra (MHNG). FALAGRIINI Cordalia occipitalis sp. n. Figg. 1-4 Holotypus 3, China, Beijing, Xishan, IX.1992, de Rougemont leg. (MHNG). Paratypi: 4 dd e3 9 9, stessa provenienza; 1 9, Beijing, B.N.V., Malaise trap, 10.VI- 10.VII.1993, de Rougemont leg.; 3 9 9, China, Luayang, 10.IX.1994, de Rougemont leg. (140° Contributo alla conoscenza delle Aleocharinae). Manoscritto accettato il 13.02.1998 306 ROBERTO PACE DESCRIZIONE. Lunghezza 2,4 mm. Corpo debolmente lucido, giallo-rossiccio con capo nero-bruno e uriti liberi quarto e quinto bruni; antenne bruno-rossicce con i due antennomeri basali rossicci; zampe giallo-rossicce. Il capo presenta una punteggiatura distinta, ma assente a metà della fronte, e una depressione mediana che termina in una fossetta basale. La superficie del capo non presenta reticolazione, come il resto del corpo. Il fine solco mediano del pronoto raggiunge solo la meta della lunghezza del pronoto che è coperto di fini e fitti tubercoletti. La superficie delle elitre è come quella del pronoto. Tubercoletti salienti coprono la superficie degli uroterghi. Edeago figg. 2-3, spermateca fig. 4. COMPARAZIONI. La nuova specie si pone tassonomicamente in posizione intermedia tra C. vestita (Boheman, 1858), presente pure in Cina e C. chinensis Pace, 1993a, dello Yunnan. Dalla prima si distingue esternamente per l'assenza di lunga pubescenza ai lati del pronoto e delle elitre, per la presenza di una fossetta occipitale (assente in vestita); dalla seconda per la presenza della fossetta occipitale (e non discale come in chinensis) e per l'assenza di profonda depressione discale del pro- noto. Per la forma della spermateca, la nuova specie può essere più affine a chinensis, ma è ben distinta da essa perché ha bulbo distale poco sviluppato, con introflessione apicale breve, mentre in chinensis ha tale introflessione assai profonda in bulbo dilatato. ETIMOLOGIA. La profonda fossetta nella regione occipitale ha suggerito il nome della nuova specie. Cordalia yunnanensis sp. n. Figg. 5-7 Holotypus 4, China, Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. (MHNG). Paratypi: 2 d d, stessa provenienza; | d, China, Yunnan, Xishuangbanna, Jinghong, II. 1993, de Rougemont leg. DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido e giallo-rossiccio con capo, elitre tranne la base e gli uriti liberi quarto e metà basale del quinto bruni; antenne brune con l’antennomero basale bruno rossiccio e l’undicesimo giallo-rossiccio; zampe rossicce. L’avancorpo è coperto di tubercoletti finissimi, l'addome di tubercoletti ben salienti e netti. L'intera superficie del corpo è priva di reticolazione. Il solco mediano del pronoto è profondo su una superficie convessa. Edeago figg. 6-7. COMPARAZIONI. La nuova specie è affine a C. chinensis Pace, 1993a per la forma dell’edeago. Ma quest’ organo nella nuova specie è più di un terzo meno sviluppato, con due spine dell'armatura genitale interna corte e robuste, mentre le corrispondenti in chinensis sono assai lunghe e sottili. Inoltre l'apice dell’edeago della nuova specie è ogivale, mentre in chinensis è a punta stretta e arrotondata. Anche esternamente la nuova specie è differenziata da chinensis. Le antenne della nuova specie sono brune con undicesimo antennomero giallo-rossiccio, mentre in chinensis le antenne sono unifor- memente rossicce. Il pronoto della nuova specie è convesso con solco esteso dal margine anteriore fino alla base e ha distinti punti basali, mentre in chinensis il pronoto ha un’ampia depressione mediana, un solco mediano non raggiungente per notevole tratto la base del pronoto stesso e assenza di robusti punti basali del pronoto. ALEOCHARINAE DELLA CINA 397 Bot 111, | nie i pui f UAEVTECUANE SUN ba Fi, Fico. 1-7 Habitus, edeago in visione laterale e ventrale e spermateca. 1-4: Cordalia occipitalis sp. n.: 5-7: Cordalia yunnanensis sp. n. 398 ROBERTO PACE Cordalia funebris sp. n. Figg. 8-11 Holotypus 6, China, Yunnan, Xishuangbanna, Mengdian, 26.1.1993, de Rougemont leg. (MHNG). Paratypi: 2 2 2, stessa provenienza; 4 es., Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. DESCRIZIONE. Lunghezza 2,8 mm. Corpo lucidissimo privo di reticolazione, con capo e pronoto neri, elitre e addome rossicci con uriti liberi quarto e quinto bruni; antenne bruno-rossicce; zampe rossicce. L’avancorpo è coperto di punteggiatura finissima, l’addome di tubercoletti fini. Il capo presenta una larga depressione mediana dal disco all’area occipitale. Il solco mediano del pronoto è profondo, posto nel fondo di una larga depressione a forma di lettera V molto larga. Edeago figg. 9-10, spermateca fig. 11. COMPARAZIONI. Per il colore nero del capo e del pronoto e per la forma della sperma- teca, la nuova specie è comparabile con C. rougemonti Pace, 1993a, pure della Cina. E° tuttavia facilmente distinguibile se si osservano gli antennomeri 4° a 10° della nuova specie, non o appena trasversi, mentre in rougemonti sono tutti fortemente trasversi. Inoltre il pronoto della nuova specie è lungo quanto largo, con netti punti basali, mentre in rougemonti è nettamente trasverso, privo di punti basali e con ampia concavità mediana posteriore, assente sul pronoto della nuova specie. Inoltre l’introflessione apicale del bulbo distale della spermateca è meno profondo nella nuova specie e profondissima in rougemonti. ETIMOLOGIA. La nuova specie è chiamata funebris per il colore nero del capo e del pronoto, tipico colore adottato nei funerali occidentali. Melagria beijingensis sp. n. Figg. 12-15 Holotypus d, China, Beijing, B.N.U., at light, V-VI.1993, de Rougemont (MHNG). Paratypi: 2 9 9, China, Yingtaogou, III.1993, de Rougemont. DESCRIZIONE. Lunghezza 1,9 mm. Corpo lucido, senza reticolazione, bruno con margine posteriore delle elitre largamente giallo; antenne brune con i due antennomeri basali giallo-bruni; zampe gialle. La punteggiatura del capo e delle elitre è fine e distinta. Il pronoto e l’addome sono coperti di tubercoletti distinti e salienti. Il pronoto mostra una larga concavità mediana posteriore. Edeago figg. 13-14, spermateca MONS" COMPARAZIONI. Per i caratteri esterni e dell’edeago, la nuova specie si mostra affine a M. gratella Erichson, 1840, della Mesopotamia. E’ distinta per avere l’edeago tozzo, con apice, in visione laterale, corto e largo e non edeago snello con apice, in visione laterale, molto stretto e ricurvo al lato ventrale come in gratella. Inoltre le antenne della nuova specie sono brune, con i due antennomeri basali giallo-bruni e le elitre sono brune marginate posteriormente di giallo, mentre in gratella le antenne sono interamente giallo-rossicce e le elitre dello stesso colore con una macchia discale scura. 399 ALEOCHARINAE DELLA CINA E E So FIGG. 8-15 Habitus, edeago in visione laterale e ventrale e spermateca. 8-11: Cordalia funebris sp. n.; 12-15: Melagria beijingensis sp. n. 400 ROBERTO PACE Melagria marginata sp. n. Figg. 16-17 Holotypus 9, China, Beijing, Panshan, 8.V.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Corpo lucidissimo e nero con elitre brune, con margine posteriore e metà posteriore della sutura gialli; antenne brune; zampe giallo- brune, le posteriori bruno-rossicce. La punteggiatura del capo è fine. Tubercoletti fini, meno densi sulle elitre, coprono la superficie del corpo al di fuori del capo. Il pronoto ha una fossetta posteriore mediana davanti allo scutello. Spermateca fig. 17. COMPARAZIONI. La nuova specie è affine alla precedente nuova specie M. beijingensis. Ne è distinta per le antenne interamente brune, con antennomeri nono e decimo nettamente trasversi, per le tempie parallele, per il pronoto nettamente trasverso, per il margine giallo delle elitre risalente lungo la sutura e per la spermateca con bulbo distale corto e trasparente (nero quasi opaco in beijingensis). Falagria (Leptagria) salamannai sp. n. Figg. 18-20 Holotypus d, China, Zhejiang, Tienmushan, 2.1X.1994, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,6 mm. Corpo lucido e bruno con capo e uriti liberi 3°, 4° e base del 5° bruno-rossicci; antenne brune con i due antennomeri basali giallo-rossicci; zampe gialle. La punteggiatura del capo è poco distinta e assente sulla fascia mediana. Le due fossette frontali sono deboli. I tubercoletti della superficie del pronoto sono fini al di fuori dell’area anteriore mediana dove sono grossolani. Quelli delle elitre sono più salienti e più fitti intorno allo scutello e quelli degli uroterghi sono ben distinti. Edeago figg. 19-20. COMPARAZIONI. La nuova specie è molto affine a F. sichuanensis Pace, 1993a, pure della Cina, nota sulla sola femmina. La nuova specie se ne distingue per gli occhi meno sviluppati, per la presenza di due fossette frontali e per la presenza di grossolani tubercoli anteriori mediani del pronoto, assenti in sichuanensis. ETIMOLOGIA. La nuova specie è dedicata al Prof. Giovanni Salamanna dell’ Università di Genova, noto ditterologo, per tanti anni direttore delle pubblicazioni della Società Entomologica Italiana, in segno di riconoscenza per l’aiuto dimostrato nei miei confronti per la pubblicazione di miei lavori. Falagria (Leptagria) occulta sp. n. Figg. 21-23 Holotypus 9, China, Yunnan, Xishuangbanna, Sanchahe, elephant res., 24.1.1993, de Rougemont (MHNG). DESCRIZIONE. Lunghezza 2,9 mm. Corpo lucido e bruno-rossiccio con capo e uriti liberi 4° e 5° bruni; antenne brune con i due antennomeri basali bruno rossicci; zampe gialle. La punteggiatura del capo è finissima e quella del pronoto è svanita. Il capo presenta una debole depressione longitudinale mediana. Il solco mediano del pronoto è profon- dissimo: alla sua estremità posteriore si notano alcuni tubercoletti assai salienti. Le elitre mostrano tubercoletti assai salienti solo alla base. Edeago figg. 22-23. ALEOCHARINAE DELLA CINA 401 a | ‘i FIGG. 16-21 Habitus, spermateca ed edeago in visione laterale e ventrale. 16-17: Melagria marginata sp. n.: 18-20: Falagria (Leptagria) salamannai sp. n.; 21: Falagria (Leptagria) occulta sp. n. 402 ROBERTO PACE COMPARAZIONI. La nuova specie avendo occhi molto sviluppati, tempie assai sfuggenti e granuli alla base delle elitre radi, si distingue esternamente da F. densi- pennis Cameron, 1939a, da F. assamensis Pace, 1985a e da F. sichuanensis Pace, 1993a che hanno occhi meno sviluppati e tempie arrotondate. Per gli occhi assai sviluppati e le tempie molto sfuggenti, la nuova specie si avvicina tassonomicamente a F. vilis Kraatz, 1859, dell’ India, nota sul solo tipo femmina, ma vilis ha antennomeri 5° a 10° nettamente trasversi, pronoto non sinuato ai lati davanti agli angoli posteriori e 1 tubercoletti basali delle elitre più fitti. Falagria (Myrmecocephalus) xishanensis sp. n. Figg. 24-25 Holotypus 9, China, Beijing, Xishan, IX.1992, de Rougemont leg. (MHNG). Paratypus: | 2, China, Henan, Luoyang, 10.IX.1994, de Rougemont leg. DESCRIZIONE. Lunghezza 3,2 mm. Capo e pronoto opachi, resto del corpo lucido. Corpo bruno-rossiccio con elitre gialle e uriti liberi 4° e 5° bruni; antenne brune con i due antennomeri basali giallo-rossicci e 1 tre seguenti bruno-rossicci; zampe gialle con metà distale dei femori mediani e posteriori bruno-rossiccia. La reticolazione della superficie del capo e del pronoto è quasi vigorosa, con punteggiatura o tubercoletti non visibili. Il pronoto ha un profondo solco mediano. I tubercoletti basali delle elitre sono appena più salienti di quelli visibili sul resto della superficie elitrale. I tuber- coletti dei due uroterghi basali sono distinti, quelli sui restanti uroterghi sono quasi indistinti. Spermateca fig. 25. COMPARAZIONI. Per le elitre gialle e la forma della spermateca, la nuova specie appare affine a F. pallipennis Cameron, 1939a, dell’India. Se ne distingue per avere le antenne brevi, con antennomeri 8° a 10° lunghi quanto larghi, per gli occhi più lunghi delle tempie che sono sfuggenti e per il pronoto sinuato davanti agli angoli posteriori che sono non evidenti. In pallipennis le antenne sono lunghissime, con gli antenno- meri 8° a 10° molto più lunghi che larghi, gli occhi più corti delle tempie che sono largamente arrotondate e il pronoto non sinuato ai lati, con angoli posteriori sporgenti. Inoltre la parte della spermateca, escluso il bulbo distale, è nettamente più corta e più larga nella nuova specie rispetto all’analoga parte della spermateca di pallipennis. Falagria (Myrmecocephalus) zhejiangensis sp. n. Figg. 26-29 Holotypus d, China, Zhejiang, Tienmushan, 29.1V.1993, de Rougemont leg. (MHNG). Paratypi: 5 es., stessa provenienza; 9 es., stessa provenienza, ma in data 2.IX.1994, de Rougemont leg.; 2 9 2, China, Shaanxi, Nanwutai, 17.1X.1995, de Rougemont leg. DESCRIZIONE. Lunghezza 4,1 mm. Capo e pronoto opachi, resto del corpo lucido. Corpo bruno-rossiccio con i due uriti basali gialli e i restanti bruni; antenne rossicce; zampe giallo-rossicce con metà distale dei femori mediani e posteriori bruna. La superficie del capo e dell'addome è coperta di reticolazione nettissima e regolare, cioè composta di maglie di grandezza uniforme. Il capo ha un debole e largo solco mediano. Il fine solco mediano del pronoto sta nel fondo di una depressione a larga V. Le elitre presentano superficie a reticolazione molto superficiale e tubercoletti più ALEOCHARINAE DELLA CINA 403 qa EL 01 mm Emme na a CSS Fico. 22-29 Edeago in visione laterale e ventrale, habitus e spermateca. 22-23: Falagria (Leptagria) occulta sp. n.; 24-25: Falagria (Myrmecocephalus) xishanensis sp. n.; 26-29: Falagria (Myrmeco- cephalus) zhejiangensis sp. n. 404 ROBERTO PACE distinti in avanti che all’indietro. L’urotergo basale mostra una carena mediana nel fondo del solco trasverso basale. La pubescenza dei tre uroterghi basali è rada, quella dei restanti uroterghi è fitta. I due uroterghi basali sono privi di reticolazione, il terzo l’ha estremamente svanita, il quarto, il quinto e il sesto l'hanno netta. Edeago figg. 27-28, spermateca fig. 29. COMPARAZIONI. Per la forma della spermateca e la struttura dell’edeago la nuova specie va associata a F. flavocinta Kraatz, 1859, dello Sri Lanka e a F. latesulcata Cameron, 1939b, di Giava (nec latesulcata Cameron 1939a dell’India = F. assa- mensis Pace, 1985a). Queste due specie non hanno l’apice dell’edeago bisinuato come quello della nuova specie e il sacco interno presenta un lungo tubulo in flavocinta e una robusta lama in /atesulcata. Inoltre il pronoto della nuova specie è molto più stretto rispetto quello delle due citate specie. Ischnopoda (Caliusa) hebeiensis sp. n. Figg. 30-32 Holotypus d, China, Hebei Beidaihe, 29.V.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,8 mm. Corpo lucido e nero; antenne brune con i due antennomeri basali e la base del terzo rossicci; zampe bruno-rossicce con tarsi giallo- rossicci. Il corpo ha superficie priva di reticolazione. I tubercoletti della superficie del capo e delle elitre sono salienti, quelli del pronoto sono molto salienti e quelli dell'addome sono radi. Edeago figg. 31-32. COMPARAZIONI. La nuova specie è simile a /. chinensis (Pace, 1993a), comb. n. (olim Tachyusa chinensis). Ne è differente per l’edeago meno sviluppato, privo di lama ricurva del sacco interno, presente in hebeiensis sp. n. e per addome più ristretto alla base. Ischnopoda (Caliusa) gilvipes sp. n. Figg. 33-37 Holotypus 9, China, Beijing, Panshan, 8.V.1993, de Rougemont leg. (MHNG). Paratypus: 1 d, China, Zhedang, Tianmushan, 29.1V.1993. DESCRIZIONE. Lunghezza 2,9 mm. Corpo lucido e nero; antenne brune con i due antennomeri basali bruno-rossicci; zampe brune con i tarsi gialli. L'intera superficie del corpo è priva di reticolazione. I tubercoletti della superficie del capo sono estremamente superficiali, quelli del pronoto sono distinti, quelli delle elitre sono netti e meno fitti di quelli del pronoto, quelli dell'addome sono salienti. Edeago figg. 34- 35, spermateca figg. 36-37. COMPARAZIONI. La nuova specie è affine a /. chinensis (Pace, 1993a). Ne è distinta per l’edeago meno sviluppato e non ricurvo al lato ventrale e per la spermateca molto meno sviluppata, con introflessione apicale del bulbo distale più profondo e più stretto (larga e poco profonda in chinensis). Ischnopoda (Caliusa) turfanensis sp. n. Figg. 38-39 Holotypus 9, China, Xinjiang, Turfan, 40 m, 11.X.1993, de Rougemont leg. (MHNG). ALEOCHARINAE DELLA CINA 405 ready AU I ry a EE Enti te FıGG. 30-39 Habitus, edeago in visione laterale e ventrale e spermateca. 30-32: Ischnopoda (Caliusa) hebeiensis sp. n.; 33-37: Ischopoda (Caliusa) gilvipes sp. n.; 38-39: Ischnopoda (Caliusa) turfanensis sp. n. 406 ROBERTO PACE DESCRIZIONE. Lunghezza 2,9 mm. Corpo lucido e bruno con elitre giallo-rossicce con base bruna e con margine posteriore degli uroterghi liberi 1°, 2° e 3° e restanti uroterghi bruno-rossicci; antenne brune; zampe rossicce. L’avancorpo è privo di reticolazione come i tre uriti basali, i restanti uriti presentano una reticolazione distinta. La punteggiatura del capo è molto superficiale e assente sulla linea mediana. I tubercoletti della superficie del pronoto sono distinti, quelli delle elitre sono poco salienti e quelli degli uroterghi sono un po’ allungati longitudinalmente. Spermateca fig. 38. COMPARAZIONI. Le elitre giallo-rossicce con base bruna e la caratteristica forma della spermateca, rendono unica questa nuova specie nell’ambito del genere a cui è attri- buita. Ischnopoda (Caliusa) finitima sp. n. Figg. 40-43 Holotypus 9, Hong Kong, N. T., IV.1996, de Rougemont leg. (MHNG). Paratypi: 2 9 9, stessa provenienza. DESCRIZIONE. Lunghezza 2,2 mm. Corpo lucido e bruno-rossiccio con capo e quarto urite libero bruni; antenne brune con i due antennomeri basali e la base del terzo giallo-rossicci; zampe giallo-rossicce. L'intero corpo è privo di reticolazione. La punteggiatura del capo è indistinta e quella del pronoto è finissima e svanita. Il capo ha un solco mediano e il pronoto una fossetta mediana posteriore trasversa. I tubercoletti che coprono la superficie delle elitre sono fini ed estremamente svaniti, quelli dell'addome sono al contrario distinti. Edeago figg. 41-42, spermateca fig. 43. COMPARAZIONI. La nuova specie, in base alla forma della spermateca, sembra tassono- micamente affine a /. song (Pace, 1993a), comb. n. (Olim Tachyusa song), pure della Cina. Se ne distingue per il pronoto più trasverso e per la parte prossimale della spermateca nettamente più lunga. Gnypeta beijingensis sp. n. Figg. 44-48 Holotypus 9, China, Beijing, Xiaolongmen, 1.VII.1993, de Rougemont leg. (MHNG). Paratypi: 8 es., China, Shanxi, Wutaishan, 4-5.VI.1993, de Rougemont leg; 1 d e 1 9, China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e bruno con capo nero-bruno e con margine posteriore dei due uriti basali largamente rossicci; antenne brune con i due antennomeri basali e la base del terzo rossicci; zampe anteriori rossicce, zampe intermedie rossicce con femori bruno-rossicci, zampe posteriori brune con tarsi gialli. La reticolazione della superficie del capo è distinta, quella del pronoto è superficiale e quella delle elitre e degli uriti liberi 4° e 5° è estremamente svanita; assente è sui due uroterghi basali. I tubercoletti della superficie del capo sono fini e non fitti, quelli del pronoto e dell’addome sono salienti e quelli delle elitre sono distinti. Edeago figg. 45- 46, spermateca figg. 47-48. COMPARAZIONI. La nuova specie è affine, ma ben distinta da G. modesta Bernhauer, 1915 di Sumatra, Thailandia, Birmania, Bali e Hong Kong, per l’habitus snello (tozzo ALEOCHARINAE DELLA CINA 407 Ficc. 40-48 Habitus, edeago in visione laterale e ventrale e spermateca. 40-43: Ischopoda (Caliusa) finitima sp. n.; 44-48: Gnypeta beijingensis sp. n. 408 ROBERTO PACE in modesta), per l’edeago con robusta armatura genitale interna (armatura appena distinta in modesta) e per il bulbo prossimale della spermateca ben conformato (indistinto in modesta). Gnypeta chinensis sp. n. Figg. 49-52 Holotypus ¢, China, Yunnan, Xishuangbanna, Chayanhe F. P., 24.1.1993, de Rouge- mont leg. (MHNG). Paratypi: 1 d e 6 9 9, stessa provenienza; | ©, stessa provenienza, ma Mengadien, 26.1.1993, de Rougemont leg.; 1 4, stessa provenienza, ma Jinghoo, 3.1.1993, de Rougemont. DESCRIZIONE. Lunghezza 2,8 mm. Avancorpo debolmente opaco, addome debolmete lucido. Corpo bruno con margine posteriore delle elitre giallo e con uriti liberi 4° e 5° neri; antenne brune con i due antennomeri basali e la base del terzo giallo-rossicci, undicesimo rossiccio; tarsi gialli, tibie rossicce e femori bruni. L’intero corpo è coperto di tubercoletti fini tra loro contigui. Edeago figg. 50-51, spermateca fig. 52. COMPARAZIONI. La nuova specie è simile a G. yaoana Pace, 1992 della Thailandia, a motivo della forma dell’edeago e della spermateca. Ne è distinta, tra l’altro, per l’apice dell’edeago largo e arcuato e non stretto e a lati sinuati come in yaoana e per il bulbo distale della spermateca più lungo, con la parte prossimale della stessa spermateca molto più breve rispetto la medesima parte della spermateca di yaoana. Gnypeta samchunensis sp. n. Figg. 53-56 Holotypus 9, Hong Kong, XII-I.1996, de Rougemont leg. (MHNG). Paratypi: 16 9 2, Hong Kong, Kadoorie Agricultural Research Centre, flight inter- ception trap, 19-31.V.1996, de Rougemont leg.; 1 2, Hong Kong, Tai Po, Malaise trap, V.1966, de Rougemont leg.; 25 es., Hong Kong, N.T., V.1996, vegetable refuse, de Rougemont; 11 es., Hong Kong, N.T. IX.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 3,3 mm. Avancorpo debolmente lucido, addome lucido. Corpo bruno con elitre bruno-rossicce aventi il margine posteriore finemente giallo; antenne brune con i quattro antennomeri basali e l’apice dell’undicesimo giallo- rossicci; zampe gialle con femori giallo-bruni. L'intero corpo è coperto di tubercoletti superficiali. La reticolazione del capo è assente sul disco, quella del pronoto e delle elitre è svanita e quella dell'addome è assente. Edeago figg. 54-55, spermateca fig. 56. COMPARAZIONI. Anche questa nuova specie, come la precedente, è simile a G. yaoana Pace, 1992 della Thailandia. Ne è distinta per avere la sporgenza ventrale dell’edeago poco saliente (molto in yaoana), l’apice dello stesso organo ogivale (e non sinuato come in yaoana) e per il bulbo distale della spermateca più sviluppato, con presenza di introflessione apicale del bulbo distale dello stesso organo (introflessione assente nella spermateca di yaoana). ETIMOLOGIA. Il nome della nuova specie deriva da Samchun, il fiume di Hong Kong. ALEOCHARINAE DELLA CINA 409 Ficc. 49-53 Habitus, edeago in visione laterale e ventrale e spermateca. 49-52: Gnypeta chinensis sp. n.; 53: Gnypeta samchunensis sp. n. 410 ROBERTO PACE Gnypeta pagodarum sp. n. Figg. 57-58 Holotypus 9, China, Yunnan, Xishuangbanna, Mengdian, 26.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 3,3 mm. Avancorpo debolmente opaco, addome lucido. Corpo nero pece con elitre brune con base e stretto margine posteriore rossicci, con i due uriti basali di un giallo sporco e con il sesto bruno; antenne nero pece con i due antennomeri basali e la base del terzo e l’undicesimo giallo-rossicci; zampe gialle con femori anteriori bruni e medi e posteriori nero-bruni. Una distinta reticolazione è presente solo sugli uroterghi. Il capo e il pronoto sono coperti di tubercoletti fittissimi, contigui e netti. Il capo ha una debole impressione discale. I tubercoletti della superficie delle elitre sono distinti e meno fitti di quelli del capo e del pronoto, quelli dell’addome sono superficiali, ma sempre fitti. Spermateca fig. 58. COMPARAZIONI. In base alla forma della spermateca, la nuova specie sembra affine a G. modesta Bernhauer, 1915, di Sumatra, Giava, Hong Kong, ecc. Ne è distinta per avere tutti gli antennomeri più lunghi che larghi (antennomeri 4° a 10° da lunghi quanto larghi a trasversi in modesta) e per il bulbo distale della spermateca piriforme e privo di introflessione apicale (e non ovale e con introflessione apicale come in modesta). Gnypeta immodesta sp. n. Figg. 59-61 Holotypus d, China, Yunnan, Xishuangbanna, Chayanhe F.P., 24.1.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,7 mm. Avancorpo debolmente opaco, addome lucido. Corpo nero con pronoto e base dell'addome bruni e con elitre brune con omeri e margine posteriore giallo-bruni; antenne brune con i due antennomeri basali rossicci; zampe anteriori rossicce con tibie gialle, quelle medie e posteriori gialle con femori bruni. L'intero corpo è privo di reticolazione. I tubercoletti che coprono la superficie del capo sono fitti e poco salienti, quelli del pronoto sono svaniti e quelli dell'addome sono fini e distinti. La punteggiatura delle elitre è superficiale. Edeago figg. 60-61. COMPARAZIONI. Esternamente la nuova specie è pressoché identica a G. modesta Bernhauer, 1915, tranne che per il colore delle elitre e delle zampe. La differenza più vistosa che separa le due specie è l’apice dell’edeago in visione ventrale che nella nuova specie è stretto, mentre in modesta è molto largo. Gnypeta lucidula sp. n. Figg. 62-63 Holotypus ©, Hong Kong, NT. vegetable refuse, V.1996, de Rougemont leg. (MHNG). Paratypi: 1 ©, Hong Kong, Tai Po, VII.1996, de Rougemont leg.; 3 © 2 Hong Kong, Kadoorie Agricultural Research Centre, VIII.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 2,6 mm. Corpo lucido e nero; antenne nero-brune con i due antennomeri basali e la base del terzo gialli; femori bruni, tibie e tarsi giallo-rossicci. Sulla superficie del corpo non vi è traccia di reticolazione. La punteggiatura del capo 411 ALEOCHARINAE DELLA CINA 01mm FIGG. 54-58 Edeago in visione laterale e ventrale, spermateca e habitus. 54-56: Gnypeta samchunensis Sp. n.; 57-58: Gnypeta pagodarum sp. n. ROBERTO PACE 412 0,1 mm 63 F1GG. 59-63 Habitus, edeago in visione laterale e ventrale e spermateca. 59-61: Gnypeta immodesta sp. n.; 62-63: Gnypeta lucidula sp. n. ALEOCHARINAE DELLA CINA 413 è svanita, quella del pronoto è molto superficiale e quella delle elitre è distinta. Tubercoletti molto salienti stanno sugli uroterghi. Spermateca fig. 63. COMPARAZIONI. Specie simile a G. rougemonti Pace, 1984a, della Brimania, data la forma simile della spermateca. Se ne distingue per gli occhi più sviluppati e per le elitre poco più larghe del pronoto (molto più larghe del pronoto in rougemonti). Il bulbo distale della spermateca è troncoconico con calotta sferica, mentre in rouge- monti è ellittico. La parte prossimale della stessa spermateca è breve nella nuova specie e lunga in rougemonti. Gnypeta yunnanensis sp. n. Figg. 64-67 Holotypus d, China, Yunnan, Dali, 9.11.1993, de Rougemont leg. (MHNG). Paratypi: 13 es., stessa provenienza. DESCRIZIONE. Lunghezza 2,8 mm. Corpo lucido e nero; antenne nero-brune; zampe nere con ginocchia gialle e tarsi rossicci. Non è presente una reticolazione sulla superficie del corpo, tranne che sul disco del capo dove la reticolazione è distinta nel fondo di una fossetta. La punteggiatura del capo è fitta, distinta e assente sulla linea mediana longitudinale. I tubercoletti della superficie del pronoto, che ha una fossetta mediana posteriore profonda, sono finissimi e distinti come quelli delle elitre e dell'addome. Edeago figg. 65-66, spermateca fig. 67. COMPARAZIONI. La presenza di una fossetta discale del capo e il particolare colore delle tibie, distinguono la nuova specie da G. hauseri Bernhauer, 1940 della Cina. DEREMINI Demerinda hongkongensis sp. n. Figg. 68-73 Holotypus 4, Hong Kong, XII.1995-1.1996, flight interception trap, de Rougemont leg. (MHNG). Paratypi: 2 d d e 1 9, stessa provenienza. DESCRIZIONE. Lunghezza 3,2 mm. Corpo lucido. Capo nero-bruno, pronoto bruno- rossiccio, elitre giallo-brune, addome rossiccio con uriti 4° e base del 5° bruni; antenne brune con i due antennomeri basali giallo-bruni; zampe gialle. La reticolazione del capo è estremamente svanita, quella del pronoto è superficiale, quella delle elitre è distinta e quella dell’addome è assente. I tubercoletti della superficie del capo e del pronoto sono fini e distinti, quelli delle elitre sono svaniti e quelli dell’addome sono salienti. Il pronoto presenta un profondo solco mediano e uno altrettanto profondo a ciascun lato: essi non raggiungono il margine anteriore, ma solo quello posteriore. Edeago figg. 69- 70, spermateca fig. 73, labio con palpo labiale 72, mento fig. 73. COMPARAZIONI. La nuova specie è distinta da D. termitophila Cameron, 1927, dell’India, per gli antennomeri 4° e 5° nettamente trasversi (lunghi quanto larghi in termitophila), per la presenza di un largo solco mediano del pronoto (fine e corta linea longitudinale mediana impressa e una fossetta davanti allo scutello in fermitophila). 414 ROBERTO PACE ORTEN UL CUR 2 AH m 5 1 BROS" SER, h ee =i E 44 E (al = 4 n À VIII Et ra | nn 1 005mm Tal NE MISE FO DU n QUE sl UN di Imm Fico. 64-71 Habitus, edeago in visione laterale e ventrale e spermateca. 64-67: Gnypeta yunnanensis sp. n.; 68-71: Demerinda hongkongensis sp. n. ALEOCHARINAE DELLA CINA 415 Kamptomerus gen. n. Figg. 74-78 DIAGNOSI. Il nuovo genere è da distinguere dal genere Longiprimitarsus Eichelbaum, 1915, dell’Africa orientale, e Demerinda Cameron, 1927, dell’ India, per la presenza di due profonde foveole sul pronoto (fig. 74) e per il margine anteriore del mento profondamente arcuato all’indietro (fig. 78). DESCRIZIONE. Corpo poco depresso, fittamente pubescente, soprattutto sulle elitre e sull’addome; tempie non marginate; palpi labiali di tre articoli; iigula larga e divisa all'estremità; paraglosse sporgenti in avanti; palpi mascellari di quattro articoli; mento profondamente incavato al margine anteriore; processo mesosternale acuto: la sua punta raggiunge quella del processo metasternale; mesocoxe contigue; formula tarsale 4-5-5; primo tarsomero posteriore molto lungo. TYPUS GENERIS. Kamptomerus bifoveolatus sp. n. ETIMOLOGIA. Il nome del nuovo genere significa “Lato arcuato” e allude a quello anteriore del mento. Il genere grammaticale evidentemente è maschile. Kamptomerus bifoveolatus sp. n. Figg. 74-78 Holotypus d, Hong Kong, Tai Po, III. 1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido, senza reticolazione, tranne che sulle elitre dove la reticolazione è distinta (forse le elitre non appartengono a questo esemplare perché recuperate staccate nel liquido di conservazione in provetta). Capo ed elitre bruni, pronoto rossiccio, addome giallo-rossiccio con gli uriti liberi 2° e 3° bruno-rossicci, tranne il margine posteriore, e con il 4° bruno; antenne giallo-brune con i tre antennomeri basali gialli; zampe gialle. La punteggiatura del capo e del pronoto è netta. Tubercoletti fini e distinti coprono le elitre (forse non pertinenti all’esemplare) e l'addome. Edeago figg. 75-76, labio con palpo labiale fig. 77, mento fig. 78. ATHETINI (parte I) Brachyusa rougemonti sp. n. Figg. 79-81 Holotypus d, China, Shaanxi, Nanwutai, 17.1X.1995, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,7 mm. Corpo debolmente opaco e nero pece, comprese le antenne; zampe gialle con femori rossicci. L'intera superficie del corpo è coperta di tubercoletti fitti e di pubescenza sericea fitta su un fondo a reticolazione distinta. Il pronoto mostra un solco mediano posteriore. Edeago figg. 80-81. COMPARAZIONI. La nuova specie è ben distinta da B. velox Cameron, 1939a, dell’ India, per avere le elitre non marginate di giallo posteriormente e per molti caratteri differenziali dell’edeago, tra cui l’apice di quello della nuova specie che è acuto, mentre quello di velox è ogivale e largo. 416 ROBERTO PACE Fico. 72-78 Labio con palpo labiale, mento, habitus, edeago in visione laterale e ventrale. 72-73: Deme- rinda hongkongensis sp. n.; 74-78: Kamptomerus bifoveolatus gen. n., sp. n. ALEOCHARINAE DELLA CINA 417 ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore, il collega Guillaume de Rougemont, noto studioso di Staphylinidae di Londra. Outachyusa chinensis sp. n. Figg. 82-83 Holotypus 9, China, Yunnan, Ruili, ca. 700 m, 3.11.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,0 mm. Corpo debolmente opaco e bruno con margine posteriore delle elitre giallo; antenne brune; zampe gialle. L'intera superficie del corpo è coperta di fitta pubescenza sericea e di fittissimi tubercoletti che simulano una reticolazione. Spermateca fig. 83. COMPARAZIONI. La nuova specie è distinta da O. nepalensis Pace, 1991a, per le elitre poco più larghe del pronoto (molto più larghe del pronoto in nepalensis) e marginate di giallo all'indietro (uniformemente picee in nepalensis). Inoltre la spermateca della nuova specie è meno sviluppata, con distinta introflessione apicale del bulbo distale (assente in nepalensis). Hydrosmecta cooteri sp. n. Figg. 84-87 Holotypus 4, China, Zhejiang Prov., Anji County, ca. 500 m, Long Wang Shan N.R., 17.V.1996, J. Cooter leg. (MHNG). Paratypus: 1 ©, stessa provenienza. DESCRIZIONE. Lunghezza 2,9 mm. Corpo poco convesso, debolmente lucido e nero pece; antenne nero-brune; zampe gialle. La reticolazione della superficie del capo e del pronoto è netta e regolare, quella delle elitre è svanita e quella dell’addome è a maglie un po’ trasverse distinte. La punteggiatura del capo (assente sulla linea mediana) e del pronoto è estremamente superficiale. I tubercoletti delle elitre sono fini ed estremamente svaniti e quelli dell'addome sono distinti. Edeago figg. 85-86, sper- mateca fig. 87. COMPARAZIONI. In base alla forma simile della spermateca, la nuova specie è tassono- micamente avvicinabile a H. aquarum Pace, 1985b, del Nepal. Ne è ben distinta per avere gli occhi lunghi quanto le tempie (occhi molto più corti delle tempie in aquarum), antenne più lunghe e per la robusta introflessione apicale del bulbo distale della spermateca (assente in quello di aguarum). ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore Jonathan Cooter di Hereford (Gran Bretagna) noto studioso di Liodidae. Hydrosmecta perignota sp. n. Figg. 88-90 Holotypus d, China, Beijing, B.N.U., at light, V-VI.1993, de Rougemont leg. (MHNG). (0 DESCRIZIONE. Lunghezza 2,2 mm. Corpo debolmente lucido e bruno, comprese le antenne; zampe gialle. Il corpo è coperto di pubescenza sericea fitta e non ha reticolazione, tranne che sul capo dove è estremamente svanita. Tubercoletti fini e fitti coprono la superficie di tutto il corpo. 418 ROBERTO PACE Fico. 79-83 Habitus, edeago in visione laterale e ventrale e spermateca. 79-81: Brachyusa rougemonti Sp. n.; 82-83: Outachyusa chinensis sp. n. Imm ALEOCHARINAE DELLA CINA F1GG. 84-90 84-87: Hydrosmecta cooteri sp. n.; 88-90: Hydrosmecta perignota Sp. n. 419 420 ROBERTO PACE COMPARAZIONI. In base alla forma simile dell’edeago, la nuova specie sembra tassonomicamente vicina a H. interiecta Pace, 1985b, pure presente in Cina. Il pronoto e le elitre della nuova specie sono meno trasversi. Tra le molte differenze dell’edeago, l’apice in visione ventrale è ogivale nella nuova specie e a punta stretta in interiecta. Hydrosmecta rougemonti sp. n. Figg. 91-95 Holotypus d, Hong Kong, XII.1995-1.1996, de Rougemont leg. (MHNG). Paratypi: 12 es. stessa provenienza; 1 d e 1 ©, Hong Kong, Kadoorie Agricultural Research Centre, flight interception trap, 18-31.V.1996; 2 es., Hong Kong, Tai Po, VII.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 2,1 mm. Corpo lucido, depresso e bruno con capo, lati delle elitre e uriti liberi 4° e 5° nero-bruni; antenne brune con l’antennomero basale rossiccio e il secondo bruno-rossiccio; zampe gialle. L’intero corpo è coperto di pubescenza sericea e di tubercoletti fitti, tranne che sui tre uroterghi apicali. Il capo ha un largo solco mediano. Un appiattimento mediano della superficie è visibile sul pronoto. Edeago figg. 92-93, spermateca fig. 94, sesto urotergo libero del maschio fig. 95. COMPARAZIONI. La nuova specie è affine a H. newarica Pace, 1988, del Nepal, ma gli occhi sono più lunghi delle tempie (in newarica occhi più corti delle tempie) ed è assente una larghissima introflessione apicale del bulbo distale della spermateca. ETIMOLOGIA. La nuova specie è dedicata al collega Guillaume de Rougemont, noto studiose di Stafilinidi di Londra. Hydrosmecta longwangensis sp. n. Figg. 96-97 Holotypus ®, China, Zhejiang Prov., Anji County, ca. 500 m, Long Wang Shan N.R., 12.V.1996, J. Cooter leg. (MHNG). DESCRIZIONE. Lunghezza 2,4 mm. Corpo lucido e nero pece con base dell’addome bruna; antenne brune; zampe gialle. La reticolazione del capo è assente, quella del pronoto è superficiale, quella delle elitre distinta e quella degli uroterghi è a maglie trasverse e nette: solo sul quinto urotergo libero la reticolazione è un po’ superficiale. La punteggiatura del capo è fitta e fine, quella del pronoto e delle elitre è assente. Gil uroterghi sono coperti di tubercoletti svaniti. Il disco del capo è impresso. Spermateca fig. 96. COMPARAZIONI. La nuova specie è affine a H. newarica Pace, 1988, del Nepal. Ne è distinta per avere gli occhi lunghi quanto le tempie (e non molto più corti delle tempie come in newarica) e per le parti apicale e distale della spermateca più corte rispetto alla parte mediana della stessa spermateca. Enkoilogeneia gen. n. Figg. 98-101 Diagnosi. Il nuovo genere è affine al genere Hidrosmecta Thomson, 1858. Infatti la ligula è pressoché identica. Tuttavia è chiaramente distinto per la profonda incavatura ALEOCHARINAE DELLA CINA 421 | 01 mm 01 mm Fico. 91-97 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 91-95: Hydrosmecta rougemonti sp. n.; 96-97: Hydrosmecta longwanensis sp. n. 422 SANDOR MAHUNKA del lato anteriore del mento che perciò ha angoli anteriori spinti molto in avanti. Anche il tipo di spermateca è di tipo differente. DESCRIZIONE. Antenne corte, con antennomeri 4° a 10° assai trasversi; tempie a profilo divergente all'indietro e marginate solo all’indietro; palpi labiali di tre articoli; ligula larga e divisa in due lembi uniti; paraglosse poco sporgenti in avanti (fig. 101); mento profondamente incavato al lato anteriore (fig. 100); palpi mascellari di quattro articoli simili a quelli del genere Hydrosmecta; processo mesosternale corto non insinuato tra le mesocoxe che sono tra loro contigue; formula tarsale 4-5-5; sperma- teca fig. 99. TYPUS GENERIS: Enkoilogeneia rougemonti sp. n. ETIMOLOGIA. Il nome del nuovo genere significa “Mento incavato”. Genere grammaticale femminile. Enkoilogeneia rougemonti sp. n. Figg. 98-101 Holotypus ©, Hong Kong, Tai Po, flight interception trap, V.1996, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Corpo lucido e giallo-rossiccio con uriti liberi 4° e base del 5° rosicci; antenne rossicce con 1 tre antennomeri basali gialli; zampe gialle. La reticolazione del capo, del pronoto e dell’addome è distinta, quella delle elitre è superficiale. La superficie dell’avancorpo è coperta di tubercoletti fini e svaniti, quella dell'addome di tubercoletti distinti. Spermateca fig. 99, mento fig. 100, labio con palpo labiale fig. 101. Geostibida smetanai sp. n. Figg. 102-105 Holotypus d, Taiwan, Kaohsiung Hsien, Peinantashan trail, ridge at 2800 m, 3.VII. 1993, A. Smetana leg. (MHNG). Paratypi: 1 d e 1 9, stessa provenienza. DESCRIZIONE. Lunghezza 2,7 mm. Capo e pronoto debolmente opachi, elitre e addome lucidi. L'intero corpo, comprese antenne e zampe è giallo-rossiccio. La reticolazione del capo è netta, quella del pronoto è vigorosa, quella degli uriti liberi 1°, 2°, 3° e 6° è estremamente svanita e quella degli uroterghi liberi 4° e 5° è distinta. Gli occhi sono composti di 15 ommatidi ben salienti. Il capo ha un fine solco mediano. La plica periscultellare del maschio è molto saliente. I tubercoletti della superficie del capo sono assai svaniti, quelli del pronoto sono distinti e quelli delle elitre sono salienti, tranne che nell’area posteriore esterna dove sono svaniti. Il 5° urotergo libero ha tubercoletti allungati molto salienti, sulla metà posteriore. Edeago fig. 103-104, sper- mateca fig. 105. COMPARAZIONI. La nuova specie è affine, ma ben distinta da G. himalayiensis Pace, 1984b, del Nepal, per gli occhi maggiormente ridotti, per il pronoto appena trasverso (molto trasverso in himalayiensis), per la presenza di due pliche periscutellari basali delle elitre del maschio (assenti in himalayica) e per l’edeago più sviluppato, con parte apicale più stretta in visione ventrale. ALEOCHARINAE DELLA CINA 423 Ar: (RE lin E E n a o o Fico. 98-105 Habitus, spermateca, mento, labio con palpo labiale ed edeago in visione laterale e ventrale. 98-101: Enkoilogeneia rougemonti gen. n., sp. n.; 102-105: Geostibida smetanai sp. n. 424 ROBERTO PACE ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore, il Dr Ales Smetana, noto studioso di Staphylinidae di Ottawa. Aloconota gonggensis sp. n. Figg. 106-107 Holotypus ®, China, Sichuan, Gongga Shan, above camp 2, 2800 m, 26.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,2 mm. Corpo lucido e nero-bruno con elitre giallo-brune e uriti liberi 4° e 5° neri; antenne nere con i due antennomeri basali bruni; zampe rossicce. La reticolazione del capo è estremamente superficiale, quella del pronoto e delle elitre è distinta e quella dell’addome è a maglie molto trasverse e svanite sui tre uroterghi basali e distinta sui restanti. La punteggiatura del capo è estremamente sva- nita e assente sulla linea mediana. I tubercoletti della superficie del pronoto sono dis- tinti, quelli delle elitre sono indistinti e quelli dell'addome sono svaniti. Spermateca fig. 107. COMPARAZIONI. In base alla forma della spermateca, la nuova specie sembra tasso- nomicamente affine ad A. chakratiana (Cameron, 1939a), (olim Atheta (Metaxya) chakratiana Cameron, 1939a, comb. n.) dell’India, tuttavia se ne differenzia per il lunghissimo undicesimo antennomero, per il pronoto non fortemente ristretto all'indietro, per la debole introflessione apicale del bulbo distale della spermateca (profonda introflessione in chakratiana) e per il bulbo prossimale dello stesso organo molto più dilatato di quello corrispondente della spermateca di chakratiana. Aloconota lanzhouensis sp. n. Figg. 108-110 Holotypus ®, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIN.1994, A. Smetana leg. (MHNG). Paratypus: 1 ®, stessa provenienza. DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e nero, comprese le antenne; zampe giallo-rossicce con femori posteriori di un giallo sporco. La reticolazione della superficie del capo è distinta, quella del pronoto e delle elitre è netta e quella degli uroterghi è a maglie molto trasverse ed estremamente superficiali. L’avancorpo è coperto di tubercoletti fini e distinti, addome mostra tubercoletti svaniti. Spermateca figg. 109-110. COMPARAZIONI. La nuova specie è simile ad A. inaequalis Cameron, 1944, dell’India. Se ne distingue per il corpo meno robusto, per gli antennomeri 6° a 10° nettamente trasversi (appena trasversi in inaequalis) e per le elitre poco più larghe del pronoto (molto più larghe del pronoto in inaequalis). Di A. inaequalis non è nota la femmina. Aloconota beijingensis sp. n. Figg. 113-114 Holotypus ®, China, Beijing, Xiaolongmen, 1.VII.1993, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 4,0 mm. Corpo lucido e bruno con elitre giallo-brune ed estremità addominale bruno-rossiccia; antenne brune con antennomero basale ros- 425 ALEOCHARINAE DELLA CINA iS ee \ ee ru i Gay (REIN HN nt! \ qui il a \ Schi Imm 1mm 112 005 mm Figc. 106-112 : 108-110: Aloconota lanzhouensis Habitus e spermateca. 106-107: Aloconota gonggensis sp. n. sp. n.; 111-112: Aloconota sulcifrons (Stephens). 426 ROBERTO PACE siccio; zampe giallo-rossicce. La reticolazione della superficie del capo e dell’addome è distinta, quella del pronoto e delle elitre è molto netta. I tubercoletti della superficie del capo sono distinti e assenti sulla linea mediana, quelli del pronoto e delle elitre sono superficiali e quelli dell'addome sono distinti. Il capo ha un debolissimo solco occipitale e il pronoto presenta un debolissimo solco mediano. Spermateca fig. 114. COMPARAZIONI. In base alla forma e alla taglia del corpo, la nuova specie può essere avvicinata tassonomicamente ad A. beesoni Cameron, 1939a, dell’India. Se ne distingue per le elitre giallo-brune e non bruno-scure, nettamente più larghe e poco più lunghe del pronoto, e non più larghe e un quarto più lunghe del pronoto come in beesoni. Non è nota la femmina di beesoni. Lasiosomina lii sp. n. Figg. 119-120 Holotypus 9, China, Hebei Yongniang, 6.X.1995, Shuqiang Li leg., (MHNG). DESCRIZIONE. Lunghezza 1,8 mm. Avancorpo opaco, addome debolmente lucido. Corpo bruno con pronoto bruno-rossiccio e con larga fascia marginale posteriore delle elitre ed estremità addominale gialle; antenne bruno-rossicce con i due antennomeri basali rossicci; zampe gialle. La reticolazione del capo e del pronoto è distinta, quella delle elitre e dell’addome è netta. L’intero corpo è coperto di pubescenza fitta d’as- petto sericeo. Il pronoto ha un'impressione mediana posteriore. Spermateca fig. 120. COMPARAZIONI. Finora il genere Lasiosomina Pace, 1990, era noto solo delle Filippine. La nuova specie è distinta da L. collaris Pace, 1990, appunto delle Filippine, per il colore bruno-rossiccio del pronoto e non giallo paglierino come in collaris, per gli occhi più sviluppati e per la parte posteriore delle elitre gialla al margine (elitre uniformemente brune in collaris). Non è nota la femmina di collaris. Amischa beijingensis sp. n. Figg. 121-124 Holotypus d, China, Beijing, B.N.U., at light, V-VI.1993, de Rougemont leg. (MHNG). Paratypi: 1 3 e 1 9, Beijing, Yingtaogou, III.1993, de Rougemont leg. DESCRIZIONE. Lunghezza 1,8 mm. Corpo depresso con capo e addome bruno-rossicci, pronoto giallo-rossiccio ed elitre e margine posteriore degli uroterghi liberi 1° e 2° giallo-bruni; antenne bruno-rossicce con i due antennomeri basali giallo-rossicci; zampe gialle. La reticolazione della superficie dell’avancorpo è coperta di retico- lazione distinta. Il disco del capo è appiattito e vi è nella regione occipitale una debolissima impressione longitudinale. Le elitre sono coperte di tubercoletti finissimi e distinti. Sui tre uroterghi basali i tubercoletti sono più salienti di quelli sui restanti uroterghi. Edeago figg. 122-123, spermateca fig. 124. COMPARAZIONI. La nuova specie è simile ad A. kashmirica Cameron, 1939a, dell’ India, dato che ha pronoto giallo-rossiccio, occhi ridotti e tipo di edeago. Ne è distinta per avere il sesto urotergo libero del maschio più largamente e poco profondamente incavato al margine posteriore (profondamente incavato in kashmirica), per il pronoto meno trasverso e di un rossiccio più chiaro, per il quarto antennomero trasverso Imm ALEOCHARINAE DELLA CINA 497 01 mm 114 118 01 mm 117 116 Fico. 113-118 Habitus, spermateca ed edeago in visione laterale e ventrale. 113-114: Aloconota beijingensis sp. n.; 115-118: Tomoglossa luteicornis (Erichson). 428 ROBERTO PACE Fico. 119-124 Habitus, spermateca ed edeago in visione laterale e ventrale. 119-120: Lasiosomina lii sp. n.; 121-124: Amischa beijingensis sp. n. ALEOCHARINAE DELLA CINA 429 (lungo quanto largo in kashmirica). Ma è l’edeago che presenta molteplici caratteri differenziali, il più evidente dei quali è l’apice appuntito in visione ventrale nella nuova specie, mentre in kashmirica è tronco. La spermateca della nuova specie è molto più sviluppata, con parte prossimale avvolta a spirale nettamente più lunga rispetto alla parte corrispondente della spermateca di kashmirica. Amischa rougemonti sp. n. Figg. 125-128 Holotypus d, China, Shanxi, Wutaishan, 4-5.VI.1993, de Rougemont leg. (MHNG). Paratypi: 1 ©, stessa provenienza; 1 d, China, Gansu, Yonghai, ca. 20 Km SW Yuzhong, 2700-2800 m, 9.VII.1994, Smetana leg. DESCRIZIONE. Lunghezza 2,2 mm. Corpo lucido e bruno con metà posteriore del 5° urite libero e il 6° bruno-rossicci; antenne brune con antennomero basale bruno- rossiccio; zampe giallo-rossicce. La reticolazione della superficie del capo è svanita, quella del pronoto e delle elitre è distinta e quella dell'addome è netta. Il capo presenta una punteggiatura estremamente svanita, quasi indistinta e il disco piatto. I tubercoletti del pronoto e delle elitre sono superficiali, quelli dei quattro uroterghi basali sono salienti e quelli del quinto superficiali su un fondo a reticolazione netta solo su questo quinto urotergo. Edeago figg. 125-127, spermateca fig. 128. COMPARAZIONI. La nuova specie è ben distinta dall’affine A. kashmirica Cameron, 1939a, dell'India, per il differente colore del corpo: bruno con estremità addominale rossiccia, invece di bruno-rossiccio con pronoto rossiccio, come in kashmirica. I più evidenti caratteri differenziali si osservano nell’edeago che, tra l'altro, è profon- damente e strettamente ricurvo al lato ventrale e ha apice appuntito, in visione ven- trale, mentre l’edeago di kashmirica non è ricurvo al lato ventrale e il suo apice è tronco. ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore, il noto studioso di Staphylinidae londinese, Guillaume de Rougemont. Amischa nana sp. n. Figg. 129-130 Holotypus 2, China, Yunnan, Xishuangbanna, Sanchahe, elephant res., 24.1.1992, de Rougemont leg. DESCRIZIONE. Lunghezza 1,6 mm. Capo e pronoto debolmente lucidi, resto del corpo lucido. Corpo nero con estremità addominale nero-bruna; antenne nere; zampe giallo- brune. La reticolazione della superficie del capo e delle elitre è distinta, quella del pronoto e dell'addome è netta, nel fondo dei solchi trasversi basali. I tubercoletti della superficie del capo e delle elitre sono distinti, quelli che coprono il pronoto sono un po’ confusi nella reticolazione e quelli degli uroterghi sono superficiali. Spermateca fig. 130. COMPARAZIONI. Per la taglia ridotta del corpo e per la forma della spermateca, la nuova specie sembra simile ad A. kathmanduensis Pace, 1991a, del Nepal, ne è distinta nettamente per avere le elitre più lunghe del pronoto (elitre più corte del pronoto in kathmanduensis) e per il bulbo distale della spermateca meno sviluppato e ROBERTO PACE 430 127 126 130 01 mm 01 mm FicG. 125-130 Habitus, edeago in visione laterale e ventrale e spermateca. 125-128: Amicha rougemonti Sp. n.; 129-130: Amischa nana sp. n. ALEOCHARINAE DELLA CINA 43] con introflessione apicale del bulbo distale della stessa spermateca, profonda e larga (introflessione profonda e stretta in kathmanduensis). Geostiba (Indatheta) hongkongensis sp. n. Figg. 131-135 Holotypus d, Hong Kong, XII.1995-I.1996, flight interception trap, de Rougemont leg. (MHNG). Paratypi: 5 es., stessa provenienza; 5 es., Hong Kong, Tai Po, III.1996, de Rougemont leg.; 12 es. Hong Kong, N.T., IV.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 2,5 mm. Corpo lucido e giallo-rossiccio con metà posteriore delle elitre, tranne il margine posteriore, e metà basale del quarto urite libero, bruni; antenne bruno-rossicce con i tre antennomeri basali e l’undicesimo giallo-rossicci; zampe gialle. La reticolazione della superficie del capo e del pronoto è superficiale, quella delle elitre è estremamente svanita e quella dell’addome assente. La punteggia- tura del capo è assai superficiale e assente sulla fascia longitudinale mediana. Tubercoletti fini e superficiali stanno sulla superficie del resto del corpo. Edeago figg. 132-133, spermateca fig. 134; piastra apicale di paramero fig. 135. COMPARAZIONI. La nuova specie è affine a G. rougemonti Pace, 1993a, pure della Cina. Se ne distingue per gli occhi lunghi quanto le tempie e non molto più corti delle tempie come in rougemonti, per le elitre meno accorciate, per l’edeago di un terzo meno sviluppato e in visione ventrale con parte apicale stretta (larga in rougemonti). Inoltre le spine dell’armatura genitale interna dell’edeago sono nettamente più robuste di quelle dell’edeago di rougemonti. Geostiba (Indatheta) pervolans sp. n. Figg. 136-139 Holotypus d, Hong Kong, XII.1995-1.1996, flight interception trap, de Rougemont leg. (MHNG). DESCRIZIONE. Lunghezza 2,1 mm. Corpo lucido. Capo bruno, pronoto giallo-rossiccio, elitre brune con omeri e angoli posteriori esterni bruno-rossicci, addome giallo- rossiccio con uriti liberi 3°, 4° e base del 5° bruni; antenne brune con i tre antenno- meri basali gialli e l'undicesimo bruno-rossiccio; zampe gialle. Una reticolazione è presente solo sul disco del capo e sulla fascia longitudinale mediana del pronoto. La punteggiatura del capo è fitta, superficiale e assente sulla fascia mediana. Il pronoto è coperto di tubercoletti fini e poco distinti. La punteggiatura delle elitre e dell'addome è distinta. Edeago figg. 138-139, piastra apicale di paramero fig. 136. COMPARAZIONI. Tutte le specie del sottogenere /ndatheta Cameron, 1939a, mostrano occhi molto più corti delle tempie, tranne la specie hongkongensis sp. n. sopra de- scritta. Inoltre la nuova specie mostra elitre molto più larghe del pronoto che denun- ciano una vocazione della specie al volo attivo (infatti è stata catturata al volo). Le elitre sono poco più larghe del pronoto nelle restanti specie. L’edeago della nuova specie ha l'armatura genitale interna priva di spine. Per questo carattere la nuova specie è avvicinabile a G. notabilis (Cameron, 1939a), dell’India, ma il maschio di questa specie presenta un tubercolo mediano sui due uroterghi liberi basali e due carene mediane posteriori accostate tra loro sul sesto urotergo libero, caratteri questi (N°) ROBERTO PACE 195 136 0,05mm Fico. 131-136 Habitus, edeago in visione laterale e ventrale, spermateca e piastra apicale di paramero Geostiba (Indatheta) hongkongensis sp. n.; 136: Geostiba (Indatheta) pervolans sp. n. 005mm 131-135: ALEOCHARINAE DELLA CINA 433 assenti sull’addome del maschio della nuova specie. Non vi è dubbio sulla sua appartenenza al sottogenere /ndatheta, per la forma delle piastre apicali dei parameri, unica in questo sottogenere (fig. 136). Geostibasoma satyrus sp. n. Figg. 140-143 Holotypus d, Hong Kong, XII.1995-I.1996, de Rougemont leg. (MHNG). Paratypi: 7 es., stessa provenienza: | ©, Hong Kong, Tai Po, V.1996, de Rougemont leg.: 1 ©, Hong Kong, N.T., IV.1996, de Rougemont leg. DESCRIZIONE. Lunghezza 2,7 mm. Corpo lucido e bruno con elitre giallo-brune con base di un giallo sporco e con margine posteriore dei tre uroterghi basali rossiccio; antenne nero-brune con i due antennomeri basali rossicci; zampe gialle. Su capo e pronoto non vi è reticolazione che sulle elitre è distinta e sull’addome è netta. La punteggiatura del capo è assai superficiale e assente sulla fascia mediana. I tuber- coletti che coprono la superficie del pronoto sono svaniti, quelli delle elitre sono distinti e quelli dell’addome superficiali. Edeago figg. 141-142, spermateca fig. 143. COMPARAZIONI. L'attribuzione di questa nuova specie al genere Geostibasoma Pace, 1985c, della Nuova Zelanda, si basa sulla forma simile della ligula e della spermateca. La nuova specie è distinta da G. antipodum (Bernhauer, 1941), della Nuova Zelanda, per avere gli occhi lunghi quanto le tempie (occhi molto ridotti in antipodum), per la presenza di una lamina ventrale dell’edeago (assente nell’edeago di antipodum) e per il bulbo distale della spermateca allungato e non subsferico come in antipodum. ETIMOLOGIA. Per lo sviluppo in lunghezza notevole dell’apice dell’edeago, la nuova specie prende nome dai satiri, divinità minori della motologia greca caratterizzati dalla loro lussuria. Paraloconota montium sp. n. Figg. 144-147 Holotypus 4, China, Gansu, M. ts 25 KM E Xiahe, 3000 m, 5.VIII.1994, A. Smetana leg. (MHNG). Paratypus: 1 ©, stessa provenienza. DESCRIZIONE. Lunghezza 4,1 mm. Corpo lucido e nero con elitre nero-brune; antenne brune con i quattro antennomeri basali bruno-rossicci; zampe rossicce. La retico- lazione della superficie dell’avancorpo è netta, quella dell'addome è a maglie molto trasverse e distinte. I tubercoletti che coprono la superficie del capo sono pressoché indistinti, al contrario quelli del resto della superficie del corpo sono salienti. Il pronoto ha un debole solco mediano posteriore. Edeago figg. 145-146, spermateca fig. 147. COMPARAZIONI. In base alla struttura della spermateca e dell’edeago, la nuova specie è avvicinabile tassonomicamente a P. coiffaiti (Pace, 1984a), comb. n. (olim Liogluta), del Nepal, nonostante i caratteri dell’esoscheletro. Infatti P. coiffaiti ha zampe lunghe e pronoto fortemente ristretto all’indietro. Ciò non si osserva nella nuova specie. Il bulbo distale della spermateca della nuova specie è enormemente largo, mentre in coiffaiti lo è poco. L’apice dell’edeago della nuova specie è largo, mentre quello di coiffaiti è strettissimo. 434 ROBERTO PACE 141 142 SS 143 FIGG. 137-143 Habitus, edeago in visione laterale e ventrale e spermateca. 137-139: Geostiba (Indatheta) pervolans sp. n.; 140-143: Geostibasoma satyrus sp. n. 435 ALEOCHARINAE DELLA CINA 01 mm 145 146 147 Ol mm Imm Fico. 144-148 Habitus, edeago in visione laterale e ventrale e spermateca. 144-147: Paraloconota montium sp. n.; 148: Paraloconota fengicola sp. n. 436 ROBERTO PACE Paraloconota fengicola sp. n. Figg. 148-151 Holotypus d, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VI11.1994, A. Smetana leg. (MHNG). Paratypi: 19 es., stessa provenienza; 1 9, China, Gansu, Dalijia Shan, 60 Km W Linxia, 3475 m, 11.VII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 4,0 mm. Corpo lucido e nero-bruno con elitre e apice addominale bruno-rossicci; antenne brune con i quattro antennomeri basali bruno- rossicci; zampe giallo-rossicce. La reticolazione che copre la superficie dell’avan- corpo è nettissima, quella dell’addome è a maglie molto trasverse e distinte. I tubercoletti diffusi sulla superficie del capo sono poco salienti, quelli sul resto della superficie del corpo sono distinti. Il pronoto ha una depressione mediana posteriore. Edeago figg. 149-150, spermateca fig. 151. COMPARAZIONI. Poiché l’edeago ha l’apice largo e la spermateca ha la parte prossi- male avvolta in numerose spire, la nuova specie è ben differente da P. coiffaiti (Pace, 1984a), del Nepal, che mostra apice dell’edeago strettissimo e parte prossimale della spermateca con una sola spira. ETIMOLOGIA. Il nome della nuova specie significa “Colei che abita i picchi montani”. Infatti “feng” in cinese significa picco montano. Paraloconota almaatensis sp. n. Figg. 152-155 Holotypus d, Kazakhstan, Alma Ata, 1000 m, 18.IX.1994, de Rougemont leg. (MHNG). Paratypi: 16 es., stessa provenienza. DESCRIZIONE. Lunghezza 3,6 mm. Corpo lucido e nero pece; antenne rossicce; zampe giallo-rossicce. La reticolazione del capo è vigorosissima, quella sul resto del corpo è netta o distinta. Il capo e le elitre sono coperti di tubercoletti quasi indistinti, il pronoto e l'addome li hanno fini e molto superficiali. Edeago figg. 154-155, sper- mateca fig. 153. COMPARAZIONI. Tra le varie specie di Paraloconota Cameron, 1939a, quella che presenta i due lembi dell’apice dell’edeago strettissimi come quelli dell’edeago della nuova specie, è P. jaloriensis Cameron, 1939a, dell’India. Tuttavia l’edeago della nuova specie non è profondamente ricurvo come quello di jaloriensis, nè la “crista apicalis” dell’edeago della nuova specie è molto sviluppata quanto quella di jalo- riensis. Paraloconota gansuensis sp. n. Figg. 156-157 Holotypus 9, China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994, A. Smetana leg. (MHNG). Paratypus: | ®, stessa provenienza, ma 60 Km W Linxia, 3475 m, 11.VII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,9 mm. Corpo debolmente lucido e nero; antenne bruno- rossicce con antennomero basale bruno; zampe bruno rossicce con femori bruni. La reticolazione del capo e del pronoto è vigorosa, quella delle elitre e dell'addome ALEOCHARINAE DELLA CINA 437 0,1 mm Fico. 149-153 Edeago in visione laterale e ventrale, spermateca ed habitus. 149-151: Paraloconota fengicola sp. n.; 152-153: Paraloconota almaatensis sp. n. 438 ROBERTO PACE distinta, fine sulle elitre e a maglie nettamente trasverse sull’addome. L’avancorpo è coperto di tubercoletti indistinti o poco distinti, l'addome invece presenta tubercoletti salienti. Spermateca fig. 157. COMPARAZIONI. La nuova specie è simile a P. smetanai Pace, 1991b, del Nepal, per la presenza di una fossetta discale del capo e per il largo bulbo distale della spermateca. Tuttavia la parte prossimale della spermateca della nuova specie è estremamente sviluppata in lunghezza, ciò non si osserva nella parte prossimale della spermateca di smetanai che ha questa porzione corta e robusta. Paraloconota yonghaiensis sp. n. Figg. 158-160 bis Holotypus d, China, Gansu, Yonghai, ca. 20 Km SW Yuzhong, 2700-2800 m, 9.VIH. 1994, A. Smetana leg. (MHNG). Paratypi: 1 d e 2 9 9, stessa provenienza. DESCRIZIONE. Lunghezza 4,1 mm. Corpo lucido e nero; antenne e zampe nero-brune, tarsi rossicci. La reticolazione della superficie del capo e del pronoto è netta, quella delle elitre è distinta e quella dell'addome è svanita sui tre uriti basali e netta e a maglie poligonali irregolari sul resto degli uroterghi. Il capo e le elitre presentano punteggiatura quasi indistinta, il pronoto ha tubercoletti della superficie fini e super- ficiali e due profonde fossette sulla metà posteriore. Edeago figg. 159-160, sperma- teca fig. 160 bis. COMPARAZIONI. Soprattutto in base alla forma della spermateca, la nuova specie può essere tassonomicamente assai vicina a P. almorensis Cameron, 1939a, dell’India. Tuttavia l’edeago di almorensis è profondamente arcuato al lato ventrale e i suoi due lembi apicali sono larghi, mentre l’edeago della nuova specie è poco ricurvo al lato ventrale e i suoi due lembi apicali sono strettissimi. Inoltre le due profonde fossette del pronoto della nuova specie sono assenti sul pronoto di almorensis. Paraloconota difficilis sp. n. Figg. 161-162 Holotypus 2, China, Gansu, M. ts 25 Km E Xiahe, 3000 m, 5.VIII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,1 mm. Corpo lucido e nero con estremità addominale bruno-rossiccia; antenne brune con antennomero basale nero-bruno; zampe rossicce con femori bruno-rossicci. La reticolazione della superficie del capo è netta, quella del pronoto e delle elitre è quasi vigorosa e quella dell’addome a maglie molto trasverse distinte. I tubercoletti della superficie del capo sono salienti e assenti sulla fascia mediana, quelli del pronoto e delle elitre sono molto salienti, quelli dei tre uroterghi basali sono distinti e quelli sui restanti uroterghi sono svaniti. Spermateca fig. 162. COMPARAZIONI. In base alla forma della spermateca, la nuova specie appare affine a P. naddiana Cameron, 1939a, dell’India. Ma la nuova specie non ha pronoto poco trasverso come quello di naddiana, ma lo è molto. Inoltre la spermateca della nuova specie è molto più piccola, con introflessione apicale del bulbo distale subconica e non ovale come in naddiana. ALEOCHARINAE DELLA CINA 439 IRRE er | a be) bs SANDI SES A pu PAR ALES cae Imm Ficc. 154-158 Edeago in visione laterale e ventrale, habitus e spermateca. 154-155: Paraloconota almaatensis sp. n.; 156-157: Paraloconota gansuensis sp. n.; 158: Paraloconota yonghaiensis sp. n. 440 ROBERTO PACE Emmelostiba chinensis sp. n. Figg. 163-165 Holotypus d, China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 2,2 mm. Corpo lucidissimo e nero, comprese le antenne; zampe nero-brune con tarsi rossicci. La reticolazione sulla superficie del capo è assente, quella del pronoto è superficiale, quella delle elitre è molto svanita e quella dell’addome è assente sui tre uroterghi basali e a maglie poligonali irregolari distinte sui restanti uroterghi. La punteggiatura dell’avancorpo è superficiale, quella dell’addome è distinta. Il capo è privo di punteggiatuta sulla linea mediana e il pronoto presenta un debole solco mediano posteriore. Edeago figg. 164-165. COMPARAZIONI. Tra le poche specie del genere Emmelostiba Pace, 1982, aventi elitre più lunghe del pronoto, la più affine sembra essere E. brachycephala (Cameron, 1939a), dell’ India. La nuova specie però ha il quarto antennomero molto trasverso, mentre quello di brachycephala è lungo quanto largo e l’edeago della nuova specie ha “crista apicalis” molto più sviluppata, mentre in brachycephala la “crista apicalis” è quasi del tutto assente e presso essa sta una lamina sporgente, assente sull’edeago della nuova specie. Liogluta sinensis sp. n. Figg. 166-169 Holotypus d, China, Gansu, Dalijia Shan, 60 Km W Linxia, 3475 m, 11.VII.1994, a. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e nero, antenne comprese; zampe rossicce con femori bruni. L'intero corpo è coperto di reticolazione netta. Il capo presenta punteggiatura fitta e superficiale e una profonda fossetta discale. I tuber- coletti della superficie delle elitre sono poco distinti, quelli del pronoto e dell'addome sono distinti. Il pronoto mostra un appiattimento dorsale mediano. Edeago figg. 167- 168, sesto urotergo libero del maschio fig. 169. COMPARAZIONI. La nuova specie è affine a L. nepalica Scheerpeltz, 1976, del Nepal., sia per la struttura dell’edeago, che per i caratteri del sesto urotergo libero del maschio. L’edeago della nuova specie è meno sviluppato e ha armatura genitale interna robusta ed estesa, mentre quella di nepalica è evanescente e corta. I rilievi laterali del margine posteriore del sesto urotergo libero del maschio della nuova specie sono sottili, mentre quelli di nepalica sono robusti. Inoltre il disco del capo della nuova specie ha una profonda fossetta, mentre quello di nepalica ne è privo. Liogluta xiahensis sp. n. Figg. 170-172 Holotypus d, China, Gansu, M. ts 25 Km E Xiahe, 3000 m, 5.VIII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,8 mm. Corpo lucido e nero, antenne comprese; zampe rossicce con femori bruni. La reticolazione del capo è netta sull’impressione discale e svanita sul resto della superficie. La reticolazione del pronoto è netta, quella delle elitre è vigorosa e quella dell’addome è a maglie debolmente trasverse e svanite. La ALEOCHARINAE DELLA CINA 44] E E s Fico. 159-165 Edeago in visione laterale e ventrale, spermateca e habitus. 159-160bis: Paraloconota yongha- iensis sp. n.; 161-162: Paraloconota difficilis sp. n.; 163-165: Emmelostiba chinensis sp. n. 442 ROBERTO PACE Fico. 166-170 Habitus, edeago in visione laterale e ventrale e sesto urotergo libero del maschio. 166-169: Liogluta sinensis sp. n.; 170: Liogluta xiahensis sp. n. ALEOCHARINAE DELLA CINA 443 punteggiatura del capo è superficiale. I tubercoletti sulla superficie del pronoto e delle elitre sono poco distinti. Edeago figg. 171-172. COMPARAZIONI. Per i caratteri dell’esoscheletro e soprattutto del sesto urotergo libero del maschio, la nuova specie appare molto simile a L. subumbonata Cameron, 1939a, dell’ India. Tuttavia l’edeago della nuova specie è profondamente arcuato al lato ventrale e un’armatura genitale del suo interno è un tubulo cortissimo. Questi caratteri non sono presenti nell’edeago di subumbonata che non è arcuato al lato ventrale e ha un’armatura genitale composta tra l’altro di una membrana coperta di dentini e di una piastra terminante a punta triangolare. Liogluta inverecunda sp. n. Figg. 173-176 Holotypus d, China, Gansu, Dalijia Shan, 60 Km W Linxia, 3475 m, 11.VII.1994, A. Smetana leg. (MHNG). Paratypi: 3 dd e 1 9, stessa provenienza; 1 d e 4 9 9, China, Gansu, M. ts 25 Km E Xiahe, 3000 m, 5.VIII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e nero, comprese le antenne; zampe bruno-rossicce con tarsi rossicci e femori bruni. La reticolazione del capo è svanita, quella del pronoto molto superficiale, quella delle elitre è netta e quella dell’addome è a maglie poligonali irregolari distinte. La punteggiatura del capo è distinta e assente sulla linea mediana, quella del pronoto è svanita. I tubercoletti della superficie delle elitre sono poco distinti. Il disco del capo è impresso. La pubescenza delle zampe è molto lunga. Edeago figg. 174-175, spermateca fig. 176. COMPARAZIONI. La nuova specie è probabilmente affine a L. verecunda Cameron, 1939a, dell’India, a motivo della forma simile della spermateca. Tuttavia la nuova specie mostra occhi più sviluppati e bulbo distale della spermateca non ovale come in verecunda, con introflessione apicale del bulbo distale della stessa spermateca triangolare a base stretta, ma triangolare a base larga. Liogluta xiaheorum sp. n. Figg. 177-180 Holotypus d, China, Gansu, M. ts 25 Km E Xiahe, 2805-2925 m, 3.VHI.1994 A. Smetana leg. (MHNG). Paratypi: | 2, stessa pronenienza; | d; stessa provenienza, ma 3000 m, 5.VIII.1994, A. Smetana leg.; 1 d e 2 2 9, Gansu, Xinlong Shan; ca. 70 Km S Lanzhou, 2225-2380 m, 7.VII.1994, A. Smetana leg. Descrizione. Lunghezza 3,3 mm. Corpo lucido e nero con elitre giallo-rossicce: antenne nere; zampe di un giallo sporco con femori bruni. Il capo e il pronoto sono privi di reticolazione. Quella delle elitre è distinta, quella dei quattro uroterghi basali è a maglie molto trasverse e distinte e quella sul quinto urotergo libero è a maglie meno trasverse di quelle sui precedenti uroterghi e nette. Il sesto urotergo libero del maschio presenta robusti granuli davanti a un rilievo marginale semiellittico. La punteggiatura ombelicata del capo è svanita e assente su una larga fascia longitudinele mediana, quella del pronoto è fine e quella delle elitre è indistinta. Edeago figg. 178-179, spermateca fig. 180. 444 ROBERTO PACE Fs 4 PA; CN sai, I ET lt lt AE I: Fısc. 171-176 Edeago in visione laterale e ventrale, habitus e spermateca. 171-172: Liogluta xiahensis sp. n.; 173-176: Liogluta inverecunda sp. n. ALEOCHARINAE DELLA CINA 445 COMPARAZIONI. La nuova specie è simile a L. subumbonata Cameron, 1939a, dell’ India, ma gli antennomeri 6° a 10° sono più trasversi, le elitre meno larghe rispetto alla larghezza del pronoto e il margine posteriore del sesto urotergo libero del maschio è ispessito per intero (solo ai lati in subumbonata). L’edeago della nuova specie ha taglia nettamente minore e se visto ventralmente è fortemente ristretto nel quarto anteriore (largo in subumbonata). L'enorme introflessione apicale del bulbo distale della spermateca permette di distinguere la nuova specie dalla femmina di subumbonata che ha introflessione apicale del bulbo distale della spermateca breve. Liogluta gonggana sp. n. Figg. 181-185 Holotypus d, China, Sichuan, Gongga Shan, above camp 3, 3050 m, 22.VII.1994, A. Smetana leg. (MHNG). Paratypi: 112 es., stessa provenienza; 3 es., stessa provenienza, ma above camp 2, 2800 m, 26.VII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,7 mm. Avancorpo debolmente lucido, addome lucido. Capo bruno, pronoto ed elitre giallo-bruni, addome nero con base e apice giallo- rossicci; antenne bruno-rossicce con i sei antennomeri basali giallo-rossicci; zampe giallo-rossicce. La reticolazione della superficie del capo è nettissima sul disco e superficiale sul resto dell’epicranio, quella del pronoto è pure nettissima, quella delle elitre è distinta e quella degli uroterghi è composta di maglie molto trasverse e svanite. I tubercoletti che coprono la superficie del capo sono assai fini, poco fitti e assenti sulla fascia longitudinale mediana, quelli del pronoto sono fini e netti, quelli delle elitre svaniti e quelli dell’addome distinti. Sesto urotergo libero del maschio fig. 182, edeago figg. 183-184, spermateca fig. 185. COMPARAZIONI. La nuova specie è simile a L. phylhygroides Cameron, 1939a, dell’ India e del Nepal. Se ne distingue per la forma del margine posteriore del sesto urotergo libero del maschio, fig. 182, e per l’edeago maggiormente sviluppato con parte apicale, in visione ventrale, molto più stretta del bulbo basale dell’edeago stesso e non appena più stretta come nell’edeago di philhygroides. L’introflessione apicale del bulbo distale della spermateca è profonda in philhygroides e breve nella nuova specie. Liogluta lacustris sp. n. Figg. 186-189 Holotypus d, China, Sichuan, Gongga Shan, lake above camp 2, 2750 m, 27.V11.1994, A. Smetana leg. (MHNG). Paratypus: 1 9, China Sichuan, Gongga Shan, above camp 2, 2800 m, 26.VII.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,7 mm. Corpo lucido e nero con elitre giallo-brune; antenne interamente nere; zampe giallo-rossicce. La reticolazione della superficie del capo è distinta, quella del pronoto è netta, quella delle elitre è quasi vigorosa e quella degli uroterghi è a maglie molto trasverse ed estremamente svanite: solo sul quinto urite libero essa è distinta. La punteggiatura del capo e del pronoto è distinta e quella delle 446 ROBERTO PACE Fico. 177-184 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 177-180: Liogluta xiaheorum sp. n.; 181-184: Liogluta gonggana sp. n. 01 mm ALEOCHARINAE DELLA CINA 447 185 189 01 mm F1GG. 185-189 Spermateca, habitus ed edeago in visione laterale e ventrale. 185: Liogluta gonggana sp. n.; 186-189: Liogluta lacustris sp. n. 448 ROBERTO PACE elitre è molto svanita; sul capo è assente sulla fascia longitudinale mediana. Edeago figg. 187-188, spermateca fig. 189. COMPARAZIONI. Il profilo ventrale dell’edeago della nuova specie è simile a quello dell’edeago di L. nepalica Scheerpeltz, 1976, del Nepal. Tuttavia, in visione ventrale, la parte distale dell’edeago della nuova specie è bruscamente incavata a ciascun lato: ciò non si osserva nella parte apicale dell’edeago di nepalica. Inoltre l'armatura genitale interna dell’edeago di nepalica è evanescente, al contrario di ciò che si osserva nell’edeago della nuova specie. Liogluta gansuensis sp. n. Figg. 190-192 Holotypus d, China, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,6 mm. Corpo lucido e nero- bruno con capo e uriti liberi 4° e 5° neri; antenne nere con 1 tre antennomeri basali bruno-rossicci; zampe gialle. La reticolazione del capo è netta sul disco e svanita ai lati, quella del pronoto e delle elitre è assente e quella degli uroterghi è a maglie molto trasverse e molto superficiali. La punteggiatura dell’avancorpo è distinta, quella dell’addome è netta. Essa è assente su una fascia mediana del capo. Edeago figg. 191-192. COMPARAZIONI. Poiché ha l’edeago esile, la nuova specie è probabilmente tassono- micamente vicina a L. philhygroides Cameron, 1939a, dell’ India, che ha pure edeago esile. La nuova specie si distingue da essa per avere il bulbo basale dell’edeago meno sviluppato della parte restante dello stesso organo (bulbo basale molto più sviluppato in philhygroides) e per la parte apicale dell’edeago molto stretta in visione ventrale (larga in phylhygroides). Liogluta hezuoensis sp. n. Figg. 193-194 Holotypus 9, China, Pass btw Hezuo-Amgog, 3300 m, 12.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,0 mm. Corpo lucido e nero, antenne comprese; zampe rossicce con femori bruni. La reticolazione della superficie del capo è svanita, quella del pronoto è netta, quella delle elitre è nettissima e quella degli uroterghi è a maglie trasverse distinte. Il capo presenta punteggiatura assai svanita e una larga depressione discale. I tubercoletti della superficie del pronoto sono fini e distinti come quelli degli uroterghi, quelli delle elitre sono indistinti. Spermateca fig. 194. COMPARAZIONI. La spermateca della nuova specie è simile a quella di L. kulliorum Pace, 1991b, del Nepal. Se ne distingue per il bulbo distale della stessa spermateca meno sviluppato, con introflessione apicale brevissima (profonda in kulliorum) e per avere gli occhi lunghi quanto le tempie (occhi assai ridotti in kulliorum). Liogluta dalijiensis sp. n. Figg. 195-196 Holotypus 9, China, Gansu, Dalijia Shan, 46 Km W Linxia, 2980 m, 10.VII.1994, A. Smetana leg. (MHNG). ALEOCHARINAE DELLA CINA 449 CALA PAS GA ; IAN BEN Imm Figc. 190-196 Habitus, edeago in visione laterale e ventrale e spermateca. 190-192: Liogluta gansuensis sp. n.; 193-194: Liogluta hezuoensis sp. n.; 195-196: Liogluta dalijiensis sp. n. 450 ROBERTO PACE DESCRIZIONE. Lunghezza 4,8 mm. Corpo lucido e nero, antenne comprese; zampe rossicce con femori bruno-rossicci. La reticolazione della superficie del capo è svanita, quella del pronoto è distinta, quella delle elitre è netta e quella dell’addome è a maglie molto trasverse e molto svanite. Il capo presenta una punteggiatura molto svanita e il disco impresso. I tubercoletti sparsi sul pronoto sono poco distinti e quelli delle elitre sono ben conformati. Spermateca fig. 196. COMPARAZIONI. Per la forma della spermateca è probabile l'affinità tassonomica della nuova specie con L. franzi Pace, 1991b, del Nepal. Ne è distinta per gli occhi lunghi quanto le tempie (occhi assai ridotti in franzi) e per la parte prossimale della sperma- teca molto più lunga nella nuova specie. Liogluta attenuata sp. n. Figg. 198-199 Holotypus 2, China, Gansu, Xilong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIH.1994, A. Smetana leg. (MHNG). Paratypi: 3 2 2, stessa provenienza. DESCRIZIONE. Lunghezza 3,6 mm. Corpo lucido e nero con elitre e i tre uriti basali nero-bruni; antenne nere; zampe giallo-rossicce. La reticolazione della superficie delle elitre è nettissima, quella sul resto del corpo è distinta. Le maglie di retico- lazione degli uroterghi sono molto trasverse. La punteggiatura del capo è svanita e assente sulla fascia mediana, quella del pronoto è distinta. Tubercoletti poco distinti coprono la superficie delle elitre. Spermateca fig. 199. COMPARAZIONI. In base alla forma della spermateca, la nuova specie è forse affine a L. philhygroides Cameron, 1939a, dell India. Se ne distingue per le elitre meno larghe rispetto alla larghezza del pronoto e per una reticolazione molto trasversa e distinta degli uroterghi, assente in philhygroides. Il bulbo distale della spermateca della nuova specie è nettamente più largo che lungo (quasi sferico in philhygroides). Liogluta langmusiensis sp. n. Figg. 200-201 Holotypus ©, China, Sichuan, Langmusi, 3500-3600 m, 13.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,8 mm. Corpo lucido e bruno-rossiccio con capo e uriti liberi 4° e 5° neri; antenne brune con i cinque antennomeri basali rossicci; zampe giallo-rossicce. La reticolazione della superficie del capo è distinta e quella sul resto del corpo è netta. Le maglie di reticolazione degli uroterghi sono molto trasverse. I tubercoletti che coprono la superficie del capo sono distinti, quelli del pronoto e delle elitre sono salienti. Spermateca fig. 201. COMPARAZIONI. La nuova specie ha la spermateca di forma simile a quella della spermateca di L. franzi Pace, 1991b, del Nepal. Se ne differenzia per la parte prossi- male della stessa spermateca più sviluppata. Inoltre gli occhi della nuova specie sono poco più corti delle tempie, mentre in franzi sono molto più corti e la reticolazione molto trasversa degli uroterghi è netta nella nuova specie e superficiale in franzi. ALEOCHARINAE DELLA CINA 451 199 \\ 01 mm Imm 201 Fico. 197-202 Habitus e spermateca. 198-199: Liogluta attenuata sp. n.; 200-201: Liogluta langmusiensis sp. n.; 202: Liogluta granulipyga sp. n. 452 ROBERTO PACE Liogluta granulipyga sp. n. Figg. 202-205 Holotypus d, China, Beijing, Xiaolongmen, 1100-1500 m, 1.VII.1993, de Rougemont leg. (MHNG). Paratypi: 2 d d e 1 9, stessa provenienza. DESCRIZIONE. Lunghezza 3,8 mm. Corpo lucido e bruno con elitre ed estremità addominale bruno-rossicci; antenne brune con antennomero basale, la metà basale del secondo e la base del terzo rossicci; zampe gialle. La reticolazione del disco del capo è distinta e composta di maglie ampie, quella sul resto della superficie del capo è svanita, quella sul pronoto è netta, quella delle elitre è distinta, quella dei quattro uroterghi basali è a maglie trasverse e distinte e quella sul quinto urotergo libero è a maglie poligonali irregolari nette. Il sesto urotergo libero del maschio è coperto di granuli robusti e di reticolazione vigorosa a maglie ampie. Il capo presenta una pun- teggiatura svanita e assente sulla fascia mediana e un'impressione discale. I tuber- coletti della superficie del pronoto sono fini e salienti, quelli delle elitre sono robusti e molto salienti. Edeago figg. 203-204, spermateca fig. 205. COMPARAZIONI. La forma della spermateca indica che la nuova specie può essere affine a L. langmusiensis sp. n. sopra descritta. Ma la nuova specie ha la sutura delle elitre appena più corta della lunghezza del pronoto, mentre /angmusiensis ha lunghezza della sutura delle elitre nettamente più corta della lunghezza del pronoto. Gli occhi lunghi quanto le tempie permettono di distinguere la nuova specie da L. franzi Pace, 1991b, che mostra occhi assai ridotti. Liogluta claripennis sp. n. Figg. 206-207 Holotypus 9, China, Zhejiang, Tianmushan, 29.1V.1993, de Rougemont leg. (MHNG): DESCRIZIONE. Lunghezza 4,2 mm. Corpo lucidissimo e nero con elitre giallo-rossicce con zona periscutellare, lato esterno e sutura bruni; antenne nere con antennomero basale bruno; zampe rossicce. La reticolazione del disco del capo è distinta e svanita sul resto della superficie del capo stesso, quella del pronoto è superficiale, quella delle elitre è distinta e quella sui due uroterghi basali è molto svanita, sui restanti uroterghi è a maglie molto trasverse, ondulate e superficiali. Il capo presenta una punteggiatura fine e distinta soprattutto ai lati e assente sulla fascia mediana. Il pronoto mostra una punteggiatura svanita. Spermateca fig. 207. COMPARAZIONI. Grazie alla forma molto simile della spermateca è possibile affermare che la nuova specie è affine a L. subumbonata Cameron, 1939a, dell’ India. Tuttavia la nuova specie mostra la sutura delle elitre appena più corta della lunghezza del pronoto, mentre la sutura delle elitre di subumbonata è nettamente più lunga della lunghezza del pronoto che è inoltre più trasverso. La parte prossimale della spermateca della nuova specie oltre la curva prossimale è molto più lunga della parte corrispondente della spermateca di subumbonata. Liogluta ceraillita sp. n. Figg. 208-209 Holotypus 9, China, Yunnan, Ruili, ca. 700 m, 3.1.1993, de Rougemont leg., (MHNG). ALEOCHARINAE DELLA CINA 453 FIGG. 203-207 203-205: Liogluta granulipyga sp. n.; 206-207: Liogluta claripennis sp. n. 454 ROBERTO PACE DESCRIZIONE. Lunghezza 3,3 mm. Corpo lucidissimo e nero con elitre giallo-brune con lati esterni bruni; antenne nere; zampe gialle con femori giallo-bruni. Solo sul disco del capo la reticolazione è distinta: sul resto della superficie del corpo è da svanita ad estremamente svanita. Sui quattro uroterghi basali la reticolazione è a maglie molto tresverse, sul quinto urotergo libero le maglie non sono trasverse. La punteggiatura del capo è superficiale e assente su una fascia mediana. Il pronoto ha punteggiatura fine. Tubercoletti fini e poco distinti coprono la superficie delle elitre. Spermateca fig. 209. COMPARAZIONI. La nuova specie in base alla forma della spermateca può essere affine a L. franzi Pace, 1991b, del Nepal, ma ha taglia della spermateca minore e più robusta, con parte prossimale più corta e distintamente dilatata (non dilatata in franzi). Anche gli occhi più lunghi delle tempie permettono di distinguere la nuova specie da franzi che mostra occhi molto più corti delle tempie. Dacrila smetanai sp. n. Figg. 210-213 Holotypus d, China, Gansu, Dalijia Shan, 60 Km W Linxia, 3475 m, 11.VII.1994, A. Smetana leg. (MHNG). Paratypus: 1 ®, stessa provenienza. DESCRIZIONE. Lunghezza 2,8 mm. Corpo lucido e nero; antenne nero-brune; zampe giallo-brune. La reticolazione della superficie del capo e del pronoto è netta, quella delle elitre è distinta e quella dell’addome è a maglie molto trasverse evidenti. I tubercoletti della superficie del capo sono distinti e assenti sulla fascia mediana, quelli del pronoto sono salienti e quelli delle elitre sono svaniti. Edeago figg. 211-212, spermateca fig. 213. NOTA. Questa specie è attribuita al genere Dacrila Mulsant & Rey, 1874, a motivo essenziale della forma della ligula che è divisa in due lembi fino alla base dove sono uniti, per le tempie arcuate e per la struttura dell’armatura genitale interna dell’ede- ago. Finora il genere Dacrila non era noto in Cina. COMPARAZIONI. La nuova specie si distingue da D. fallax (Kraatz, 1858), della regione paleartica occidentale, per i lati del pronoto sinuati davanti agli angoli posteriori e per gli antennomeri 4° a 10° più lunghi. L’edeago della nuova specie, oltre a essere meno profondamente e più largamente arcuato al lato ventrale, è bruscamente ristretto, se visto dal lato ventrale, e non profondamente e strettamente arcuato al lato ventrale e non ristretto al lato ventrale come in fallax. Inoltre la spermateca della nuova specie ha la parte prossimale lunghissima e flessa in due parti: tale parte nella spermateca di fallax è assente. ETIMOLOGIA. La nuova specie è dedicata al suo raccoglitore, il noto studioso di Staphylinidae Dr Ales Smetana di Ottawa. Dacrila setigera sp. n. Figg. 214-218 Holotypus ¢, China, Sichuan, Langmui, 3500-3600 m, 13.VII.1994, A. Smetana leg. (MHNG). Paratypi: 5 es., stessa provenienza; 1 4, China, Sichuan, Gongga Shan, above camp 3, 3300-3350 m, 23.VII.1994, A. Smetana leg. ALEOCHARINAE DELLA CINA 455 Figc. 208-213 Habitus, spermateca ed edeago in visione laterale e ventrale. 208-209: Liogluta ceraillita sp. n.; 210-213: Dacrila smetanai sp. n. 456 ROBERTO PACE DESCRIZIONE. Lunghezza 2,7 mm. Avancorpo debolmente lucido, addome lucido. Corpo nero, antenne comprese; zampe nero-brune. La reticolazione della superficie del capo è nettissima, quella del pronoto è vigorosa, quella delle elitre svanita e quella dell'addome è a maglie molto trasverse ed estremamente superficiali. I tubercoletti che coprono il capo sono distinti e assenti sulla fascia mediana, quelli del pronoto sono netti quelli delle elitre ben salienti e quelli dell’addome sono ben visibili. Edeago figg. 215-216, spermateca fig. 217, sesto urotergo libero del maschio fig. 218. COMPARAZIONI. La nuova specie è distinta da D. smetanai sp. n. sopra descritta, per i lati del pronoto non sinuati davanti agli angoli posteriori, per gli antennomeri 5° a 10° più larghi che lunghi, per l’edeago poco profondamente arcuato al lato ventrale e per l’introflessione apicale del bulbo distale della spermateca. Taxicera smetanai sp. n. Figg. 219-220 Holotypus ®, China, Gansu, pass btw Hezuoc-Amgog, 3300 m, 12.VII.1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 3,0 mm. Corpo lucido e nero, comprese le antenne; zampe anteriori con tarsi e tibie giallo-rossicci e femori giallo-bruni, medie con tarsi e tibie bruno-rossicci e femori bruni, posteriori brune con tarsi giallo-bruni. La reticolazione della superficie del capo e del pronoto è svanita, quella delle elitre è distinta e quella dell’addome è a maglie trasverse nette. La punteggiatura del capo e del pronoto è superficiale, quella delle elitre è poco distinta. Spermateca fig. 220. COMPARAZIONI. La nuova specie, presentando ligula divisa, con una setola a ciascun lato e spermateca corta, va attribuita al genere Taxicera Mulsant & Rey, 1873. Il genere era finora sconosciuto in estremo oriente. La nuova specie si distingue da T. deplanata (Gravenhorst, 1802) della regione paleartica occidentale, per avere il pronoto coperto da tubercoletti superficiali (netti in deplanata) per il differente colore del corpo (pronoto, elitre ed estremità addominale giallicci in deplanata) e delle zampe: zampe giallo-brune invece di zampe giallo-rossicce con femori bruni come nella nuova specie. Amidobia smetanai sp. n. Figg. 221-222 Holotypus ©, China, Sichuan, Gongga Shan, above camp 3, 3300-3350 m, 23.VII. 1994, A. Smetana leg. (MHNG). DESCRIZIONE. Lunghezza 4,0 mm. Corpo lucido e nero con elitre nero brune; antenne nere; zampe rossicce con femori bruni. L’avancorpo è coperto di reticolazione dis- tinta, l'addome da reticolazione molto trasversa e svanita. La punteggiatura dell’avan- corpo è poco distinta: è assente sulla linea mediana del capo. Spermateca fig. 222. COMPARAZIONI. La nuova specie è attribuita al genere Amidobia Thomson, 1858, per la forma della ligula e per la struttura della spermateca che è minuscola rispetto alla taglia corporea e lineare come nelle specie del genere. La nuova specie è distinta da A. talpa (Heer, 1842) della regione paleartica occidentale, per la taglia corporea molto ALEOCHARINAE DELLA CINA 457 Fico. 214-220 Habitus, edeago in visione laterale e ventrale, spermateca e sesto urotergo libero del maschio. 214-218: Dacrila setigera sp. n.; 219-220: Taxicera smetanai sp. n. 458 ROBERTO PACE maggiore (4,0 mm invece di 1,8 mm), per gli antennomeri 7° a 10° poco trasversi (molto trasversi in falpa) e per la spermateca priva di parte prossimale ricurva come in talpa. Amphibolusa gen. n. Figg. 223-229 Dragnosi. Il nuovo genere è affine al genere Amidobia Thomson, 1858, per la forma della ligula, ma distinta per la presenza di setole apicali sui due lembi della ligula stessa, per il pronoto ristretto in avanti, per il primo tarsomero posteriore lunghissimo e per la struttura della spermateca più simile a quella di alcune specie di Oxypoda Mannerheim, 1831, che presentano, come nel nuovo genere, microscultura all’interno della parte mediana della stessa spermateca. DESCRIZIONE. Palpi labiali di tre articoli, fig. 228, ligula robusta, divisa in due lembi fino a metà, l’apice di ciascun lembo con setole corte; paraglosse non sporgenti in avanti; palpi mascellari di quattro articoli, fig. 229; tempie marginate; pronoto ristretto in avanti; processo mesosternale acuto, insinuato fino a metà delle mesocoxe che sono fra loro contigue; formula tarsale 4-5-5, primo tarsomero posteriore molto lungo rispetto ai successivi tre; edeago di grande taglia, figg. 224-225, spermateca simile a quella di certe specie di Oxypoda. TYPUS GENERIS: Amphibolusa smetanai sp. n. ETIMOLOGIA. Il nome del nuovo genere significa “Colei che è ambigua”, infatti pre- senta caratteri morfologici esterni ed interni sia degli Athetini che degli Oxypodini. Amphibolusa smetanai sp. n. Figg. 223-229 Holotypus d, China, Sichuan, Langmui, 3500-3600 m, 13.VII.1994, A. Smetana leg. (MHNG). Paratypi: 1 d e 1 2, stessa provenienza; 1 2, China, Gansu, M. ts 25 Km E Xiahe, 2805-2925 m, 3.VIIL.1994, A. Smetana leg. DESCRIZIONE. Lunghezza 3,7 mm. Avancorpo debolmente lucido, addome lucido. Corpo bruno con addome nero-bruno; antenne brune con i tre antennomeri basali bruno-rossicci; zampe giallo-rossicce. La reticolazione del capo è distinta, quella del pronoto netta, quella delle elitre svanita e quella dell'addome assente. Il capo presenta superficie coperta di distinti tubercoletti e il disco debolmente impresso. I tubercoletti che coprono il pronoto sono salienti e quelli delle elitre sono talmente fitti da dare un aspetto scabro alla superficie. La pubescenza sui tre uriti basali è fittissima, d’aspetto sericeo, quella del quarto urite è fitta, quella del quinto a sesto è rada. Edeago figg. 224-225, spermateca fig. 226, mento fig. 227. Nehemitropia chinicola sp. n. Figg. 230-233 Holotypus d, China, Zhejiang, Tianmushan, 23.1V.1993, de Rougemont leg. (MHNG). Paratypi: 12 es., stessa provenienza. DescRIZIONE. Lunghezza 3,7 mm. Corpo lucido. Capo bruno, pronoto bruno chiaro, elitre giallo-brune con angoli posteriori esterni bruni, i due uriti basali rossicci, il terzo ALEOCHARINAE DELLA CINA 459 222 226 0,1 mm Fico. 221-226 Habitus, spermateca ed edeago in visione laterale e ventrale. 221-222: Amidobia smetanai sp. n.; 223-226: Amphibolusa smetanai gen. n., sp. n. 460 ROBERTO PACE 233 005mm FIGG. 227-233 Mento, labio con palpo labiale, maxilla con palpo mascellare, habitus, edeago in visione laterale e ventrale e spermateca. 227-229: Amphibolusa smetanai gen. n., sp. n.; 230-233: Nehemitropia chinicola sp. n. ALEOCHARINAE DELLA CINA 461 bruno, i successivi nero-bruni; antenne nero-brune con i tre antennomeri basali rossicci; zampe giallo-rossicce. La reticolazione della superficie del capo è svanita, quella del pronoto e delle elitre è distinta e quella dell'addome è estremamente super- ficiale e a maglie trasverse. La punteggiatura del capo è fitta e superficiale. Il pronoto e le elitre sono coperti di tubercoletti fini e salienti. Edeago figg. 231-232, spermateca 11233} COMPARAZIONI. La nuova specie è nettamente differente da N. jiniana Pace, 1993a, pure della Cina, come da N. sordida (Marsham, 1802) a diffusione subcosmopolita, per la forma dell’edeago e della spermateca. In jiniana l’edeago non è così profondamente arcuato al lato ventrale, come quello della nuova specie e la sperma- teca è robusta sia in jiniana che in sordida e non esile come quella della nuova specie. ADDENDA All’elenco delle specie note o nuove per la Cina, dato nella parte I della presente serie di lavori sulle Aleocharinae della Cina, va aggiunta la seguente specie: Aloconota sulcifrons (Stephens, 1832) figg. 111-112 Homalota sulcifrons Stephens, 1832: 121 Atheta (Aloconota) sulcifrons: BENICK 1954: 139 Aloconota sulcifrons: LOHSE 1974: 96 1 2, China, Gansu, Xinlong Shan, ca. 70 Km S Lanzhou, 2225-2380 m, 7.VIII.1994, A. Smetana leg. Specie cosmopolita. RINGRAZIAMENTI Rivolgo i miei più sentiti ringraziamenti a coloro che mi hanno affidato in studio le Aleocharinae della Cina oggetto del presente lavoro e frutto di recenti raccolte: il Dr Ales Smetana di Ottawa, i colleghi Guillaume de Rougemont di Londra, Jonathan Cooter di Hereford (Gran Bretagna), Garry Ades, Graham Reels di Hong Kong e il Dr Shugiang Li di Stuttgard (Germania). Per il prestito di tipi e di materiale di confronto ringrazio molto il Dr A.F. Newton del “Field Museum of Natural History” di Chicago, il Dr P.M. Hammond del “Natural History Museum” di Londra, il Dr L. Baert dell’ “Institut Royal des Sciences Naturelles de Belgique” di Bruxelles, il Dr H. Sch6nmann del “Naturhistorisches Museum” di Vienna et il Dr L. Zerche del D.E.I. di Eberswalde. 462 ROBERTO PACE BIBLIOGRAFIA BENICK, G. 1954. Revision der Untergattung Aloconota C. G. Thomson (Gattung Atheta, Staphylinidae). Entomologische Blätter 50: 133-174. BERNHAUER, M. 1915. Neue Staphyliniden aus Java und Sumatra. Tijdschrift voor Entomologie 58: 213-243. BERNHAUER, M. 1940. Neuheiten der paläarktischen Staphylinidenfauna (Coleoptera, Staphy- linidae). Mitteilungen der Miinchner Entomologischen Gesellschaft 30: 622-642. BERNHAUER, M. 1941. Neue Staphyliniden aus Neuseeland (New Zealand). Fol. Zool. Hydro- biol. 11: 26-38. BOHEMAN, C.H. 1858. Coleoptera, in: Svenska Fregatt Eugenies Resa Omkring Jorden, Zool., Insecta. CAMERON, M. 1927. Descriptions of two new genera of termitophilous Staphylinidae from India. The Entomologist’s monthly Magazine 63: 222-224. CAMERON, M. 1939a. The Fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae 4: 410 pp., London. CAMERON, M., 1939b. Fauna Javanica. The Staphylinidae collected by Mr C.F. Drescher. Tijdschrift voor Entomologie 82: 1-29. CAMERON, M. 1944. Descriptions of new Staphylinidae (Coleoptera). The Entomologist’s monthly Magazine 76: 103-106. EICHELBAUM, F. 1915. Verbesserung und Zusätze zu meinem Katalog der Staphyliniden- Gattungen aus dem Jahre 1909. Archiv fiir Naturgeschichte 81: 98-121. ERICHSON, W.F. 1840. Genera et Species-Staphylinorum Insectorum Coelopterorum Familiae: 954 pp., Berlin. GRAVERHORST, J.L.C. 1802. Coleoptera Microptera Brunsvicensia: 206 pp., Brunsvigae. HEER, O. 1842. Fauna Coleopterorum Helvetica: 652 pp., Turici. KRAATZ, G. 1858. Naturgeschichte der Insecten Deutschlands. Coleoptera: 1080 pp., Berlin. KRAATZ, G. 1859. Die Staphyliniden-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv für Naturgeschichte 25: 1-196. Louse, G.A. 1974. Die Käfer Mitteleuropas. Band 5 Staphylinidae II (Hypocyphtinae und Aleocharinae): 304 pp., Krefeld. MANNERHEIM, C.G. 1831. Précis d’un nouvel arrangement de la Famille del Brachélytres de Ordre des Insectes Coléoptères. Mémoires de l’Académie des Sciences de St. Petersbourg 1: 415-501. MULSANT, M. E. & C. REY. 1873. Description de divers Coléoptères Brévipennes nouveaux ou peu connus. Opuscula Entomologica 15: 147-189. MULSANT, M. E. & C. REY. 1874. Tribu des Brévipennes: Famille des Aléochariens: Septième Branche: Myrmédoniaires. Annales de la Société Agr. de Lyon 6: 33-738. PACE, R. 1984a. Aleocharinae della Thailandia e della Birmania riportate da G. de Rougemont. Bollettino del Museo civico di Storia naturale di Verona 11: 427-468. PACE, R. 1984b. Aleocharinae dell’Himalaya. Annales de la Société entomologique de France 20: 309-339. PACE, R. 1985a. Aleocharinae dell’ Himalaya raccolte da Guillaume de Rougemont. Bollettino del Museo civico di Storia naturale di Verona 12: 165-191. PACE, R. 1985b. Aleocharinae riportate dall’ Himalaya dal Prof. Franz, Parte I. Nouvelle Revue d’Entomologie (N.S.) 2: 91-105. PACE, R. 1985c. Aleocharinae attere della regione australiana e neozelandese raccolte dal Prof. Franz. Fragmenta Entomologica 18: 105-114. PACE, R. 1988. Aleocharinae dell Himalaya raccolte da Marc Tronquet e Georges Ledoux. Bollettino del Museo civico di Storia naturale di Verona 14: 403-419. ALEOCHARINAE DELLA CINA 463 PACE, R. 1990. Aleocharinae delle Filippine. In: Berti, N. (ed.), Miscellanées sur les Staphylins. Mémoires du Muséum national d'Histoire Naturelle 147: 57-113. PACE, R. 1991a. Aleocharinae nepalesi del Museo di Ginevra. Parte IV: Autaliini ed Athetini (Coeloptera, Staphylinidae). Revue suisse de Zoologie 98: 107-158. PACE, R. 1991b. Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini. Revue suisse de Zoologie 98: 803-863. PACE, R. 1992. Aleocharinae della Thailandia. Bollettino del Museo civico di Storia naturale di Verona 16: 227-268. PACE, R. 1993a. Aleocharinae della Cina. Bollettino del Museo civico di Storia naturale di Verona 17: 69-126. PACE, R. 1998. Aleocharinae della Cina: Parte I. Revue suisse de Zoologie 105 (1): 139-220. SCHEERPELTZ, O. 1976. Wissenschafftliche Ergebnisse der von Prof. Dr H. Janetschek im Jahre 1961 in das Mt-Everest-Gebiet-Nepals unternommenen Studienreise (Col. Staphy- linidae). Khumbu Himal 5: 3-75, Innsbruck & Miinchen. STEPHENS, J.F. 1832. Illustrations of British Entomology. Mandibulata 5: 1-240, London. THOMSON, C.G. 1858. Försök till uppställning af Sveriges Staphyliner. Ofv. Kon. Vet.-Akad. Förh. 15: 27-40. REVUE SUISSE DE ZOOLOGIE Tome 105 — Fascicule 2 KLIMASZEWSKI, Jan & Stewart B. PECK. A review of Aleocharine Rove Beetles from the Galäpagos Islands, Ecuador (Coleoptera: Staphy- Hinidae PAMÉOCChATINAC) ER a en en er BAEHR, Martin. Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae)...... VASILEVA, Gergana P., Boyko B. GEORGIEV & Todor GENOV. Redescrip- tion of Hymenolepis hoploporus Dollfus, 1951, with the erection of the new genus Dollfusilepis (Cestoda, Hymenolepididae).......... HUBER, Jean H. A new Cyprinodont species with a uniquely-colored female, Aphyosemion hera n. sp. (Cyprinodontiformes, Pisces), from Northwestern GabOMeeno8 cas cg Hole ood cone eke SI NO DALENS, Henri. Endémisme pyrénéen: sur une nouvelle espèce épigée du genre Oritoniscus: O. rousseti n. sp. (Crustacea, Isopoda, Oniscidea). COMELLINI, André. Notes sur les Psélaphines néotropicaux (Coleoptera, Staphylinidae, Pselaphinae) 10 - un nouveau genre et quatre nouvelles especesidella wi buideSIMEtOpioSIM EE RE LEO ea ANGELINI, Fernando & Luigi DE MARZO. Agathidiini from China, with description of 14 new species (Coeloptera, Leiodidae)............. SCHAWALLER, Wolfgang. The genus Laena Latreille (Coleoptera: Tene- brionidae) in Thailand, with descriptions of new species. .......... PUTHZ, Volker. Die Gattung Stenus Latreille in Vietnam (Coleoptera, Sap iy DIRES) Eee go RR PACE, Roberto. Aleocharinae della Cina: Parte II (Coleoptera, Staphy- lidia OR TI I SR: 221-260 261-318 319-329 331-338 339-343 345-350 351-373 375-382 383-394 395-463 REVUE SUISSE DE ZOOLOGIE Volume 105 — Number 2 KLIMASZEWSKI, Jan & Stewart B. PECK. A review of Aleocharine Rove Beetles from the Galapagos Islands, Ecuador (Coleoptera: Staphy- limidacsSAleéocharimae). +... ALTRA O BAEHR, Martin. Leleupidiini from the Oriental Region. 2. The genus Gunvorita Landin (Insecta, Coleoptera, Carabidae, Zuphiinae)...... VASILEVA, Gergana P., Boyko B. GEORGIEV & Todor GENOV. Redescrip- tion of Hymenolepis hoploporus Dollfus, 1951, with the erection of the new genus Dollfusilepis (Cestoda, Hymenolepididae).......... HUBER, Jean H. A new Cyprinodont species with a uniquely-colored female, Aphyosemion hera n. sp. (Cyprinodontiformes, Pisces), from mnorthwesterm! Gabon: =! «u... 2 24024 20m ce N DALENS, Henri. Pyrenean endemism: O. rousseti a new epigean species (Grustacca@isopodaf@niscidea) 9301) 0. MRI ee COMELLINI, André. Notes on Neotropical pselaphines (Coleoptera, Staphy- linidae, Pselaphinae) 10 - A new genus and four new species of the telbe-Metopiasini.. er... 21er RAI, OI CO ANGELINI, Fernando & Luigi DE MARZO. Agathidiini from China, with description of 14 new species (Coeloptera, Leiodidae)............. SCHAWALLER, Wolfgang. The genus Laena Latreille (Coleoptera: Tene- brionidae) in Thailand, with descriptions of new species........... PUTHZ, Volker. The genus Stenus Latreille in Vietnam (Coleoptera, Stapp lays dae) vse ek hex errata ear nt RE PACE, Roberto. Aleocharinae from China: Part II (Coleoptera, Staphy- NMAC a ee me hie ON ce em a co 0 oo Indexed in CURRENT CONTENTS, SCIENCE CITATION INDEX Pages 221 261 SIG) | 359 345 391 313 383 395 PUBLICATIONS DU MUSÉUM D'HISTOIRE NATURELLE DE GENÈVE CATALOGUE DES INVERTÉBRÉS DE LA SUISSE Fasc. 1. 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