Wy Nya ee Shier ei \\ ) } V ' PVA : 1 ne i] ie Ny ‘ii WRAP vk uit iy, y eG " HUN cr Coit yh my i i) ii att ny i: a Bie Nuc THE SCIENTIFIC MONTHLY A Hite ii Wn WS Ay t i AF f ( vee ah ay why ay 4) 1 Ki! +) i inh ' Ae dh 1 ea fps THE SCIENTIFIC MONTHLY EDITED BY J. McKEEN CATTELL VOLUME XV JULY TO DECEMBER, 1922 NEW YORK THE SCIENCE PRESS 1922 4) net i i My 4 ih f fe me “) \ 4 | ui \ ba Y WW, AWN) mn i ’ My if if) \ yl Wh vii ’ hil AYA « } Ch oe Cv : \ h Ay } | . ve oH YA i ebpamient, 1922 (0) \ WS Na Ae an ‘ aN, "| - THE SCIENCE PRESS “ Ae \)? eS RE | i “ am ) ! \ WAV Fat Al) ) THOMAS J. oir THS AND SONS i } ) ! nt Yi i Me | uh p mAh THE SCIENTIFIC MONTHLY JULY. 1922 SOME ASPECTS OF THE USE OF THE AN- NUAL RINGS OF TREES IN CLIMATIC STUDY’ By Professor A. E. DOUGLASS UNIVERSITY OF ARIZONA I. AFFILIATIONS ATURE is a book of many pages and each page tells a fas- N cinating story to him who learns her language. Our fertile valleys and craggy mountains recite an epic poem of geologic con- flicts. The starry sky reveals gigantic suns and space and time without end. The human body tells a story of evolution, of com- petition and survival. The human soul by its sears tells of man’s social struggle. The forest is one of the smaller pages in nature’s book, and to him who reads it too tells a long and vivid story. It may talk in- dustrially in terms of lumber and firewood. It may demand preservation physiographically as a region conserving water sup- ply. It may disclose great human interests ecologically as a phase of plant succession. It may protest loudly against its fauna and parasites. It has handed down judicial decisions in disputed mat- ters of human ownership. It speaks everywhere of botanical lan- guage, for in the trees we have some of the most wonderful and complex products of the vegetable kingdom. The trees composing the forest rejoice and lament with its suc- cesses and failures and carry year by year something of its story in their annual rings. The study of their manner of telling the story takes us deeply into questions of the species and the indi- vidual, to the study of pests, to the effects of all kinds of injury, especially of fire so often started by lightning, to the closeness of grouping of the trees and to the nearness and density of competing 1 Address of the President of the Southwestern Division of the American Association for the Advancement of Science, Tucson, Arizona, January 26, 1922. 6 THE SCIENTIFIC MONTHLY vegetation. The particular form of environment which interests us here, however, is climate with all its general and special weather conditions. Climate is a part of meteorology, and the data which we use are obtained largely from the Weather Bureau. Much helping knowledge needed from meteorology has not yet been gar- nered by that science. For example, the conditions for tree growth are markedly different on the east and west sides of a mountain or on the north and south slopes. The first involves difference of exposure to rain-bearing winds, and the second means entirely dif- ferent exposure to sun and shade. The latter contrast has been studied on the Catalina Mountains by Forrest Shreve. Again, the Weather Bureau stations are largely located in cities and therefore we can not get data from proper places in the Sierra Nevada Mountains of California, where the Giant Sequoia lives. Consider- ing that this Big Tree gives us the longest uninterrupted series of annual climatic effects which we have so far obtained from any source, it must be greatly regretted that we have no good modern records by which to interpret the writing in those wonderful trees, and, so far as I am aware, no attempt is yet being made to get complete records for the future. In reviewing the environment, one must go another step. One of the early results of this study was the fact that in many different wet climates the growth of trees follows closely and sometimes fundamentally certain solar variations. That means astronomical relationship. It becomes then an interesting fact that the first two serious attempts to trace climatic effects in trees were made by astronomers. I do not know exactly what inspired Professor Kapteyn, the noted astronomer of Groningen, Holland, to study the relation of oak rings to rainfall in the Rhineland, which he did in 1880 and 1881 (without publishing), but for my own ease I can be more explicit. It was a thought of the possibility of determin- ing variations in solar activity by the effect of terrestrial weather on tree growth. This, one notes, assumed an effect of the sun on our weather, a view which was supported twenty years ago by Bigelow. But the possible relationship of solar activity to weather is a part of a rather specialized department of astronomical science, called astrophysics. And there is a great deal of help which one wants from that science, but which one ean not yet obtain; for example, the hourly variations in the solar constant. I would like to know whether the relative rate of rotation and the relative tem- peratures of different solar latitudes vary in terms of the 1l-year sunspot period. These questions have to do with some of the theories proposed in attempting to explain the sunspot periodicity. THE ANNUAL RINGS OF TREES 7 We do not know the cause of the 1l-year sunspot period. Here then is work for the astronomers. Yet another important contact has this study developed. The rings in the beams of ancient ruins tell a story of the time of build- ing, both as to its climate and the number of years involved and the order of building. This is anthropology. It will be mentioned on a later page. Viewed through the present perspective, there is one way of expressing the entire work which shows more clearly its human end, a contact always worth emphasizing. If the study works out as it promises, it will give a basis of long-range weather forecasting of immense practical value for the future and of large scientific value in interpreting the climate of the past. This statement of it earries to all a real idea of the central problem. II. YEARLY IDENTITY oF RINGS The one fundamental quality which makes tree rings of value in the study of climate is their yearly identity. In other words, the ring series reaches its real value when the date of every ring ean be determined with certainty. This is the quality which is often taken for granted without thought and often challenged without real reason. The climatic nature of a ring is its most ob- vious feature. There is a gradual cessation of the activity of the tree owing to lowered temperature or diminished water supply. This causes the deposition of harder material in the cell walls, producing in the pine the dark hard autumn part of the ring. The growth practically stops altogether in winter and then starts off in the spring at a very rapid rate with soft white cells. The usual time of beginning growth in the spring at Flagstaff (eleva- tion 7,000 feet) is in late May or June and is best observed by Dr. D. T. MacDougal’s ‘‘Dendrograph,’’ which magnifies the diameter of the tree trunk and shows its daily variations. This spring growth depends upon the precipitation of the preceding winter and the way it comes to the tree. Heavy rains have a large run-off and are less beneficial than snow. The snow melts in the spring and supplies its moisture gradually to the roots as it soaks into or moves through the ground. There is evidence that if the soil is porous and resting on well cracked limestone strata, the moisture passes quickly and the effect is transitory, lasting in close proportion to the amount of rain. Trees so placed are ‘‘sensitive’’ and give an excellent report of the amount of precipitation. Such condition is commonly found in northern Arizona over a limestone bed rock. If the bed rock is basalt or other igneous material the soil over it is apt to be clay. The rock and the clay sometimes hold water until the favorable season is past and the tree growth de- 8 THE SCIENTIFIC MONTHLY pends in a larger measure on other factors than the precipitation. For example, the yellow pines growing in the very dry lava beds at Flagstaff show nearly the same growth year after year. It is sometimes large, but it has little variation. Such growth is “‘com- placent.’’ Yearly identity is disturbed by the presence of too many or too few rings. Surplus rings are caused by too great contrast in the seasons. The year in Arizona is divided into four seasons, two rainy and two dry. The cold rainy or snowy season is from De- cember to March, and the warm tropical summer, with heavy local rainfall, occurs in July and August. Spring and autumn are dry, the spring being more so than the autumn. If the snowfall of winter has not been enough to carry the trees through a long dry spring, the cell walls in June become harder and the growing ring turns dark in color as in autumn. Some trees are so strongly affected that they stop growing entirely until the following spring. A ring so produced is exceptionally small. But others near-by may react to the summer rains and again produce white tissue be- fore the red autumn growth comes on. This second white-cell structure is very rarely as white as the first spring growth and is only mistaken for it in trees growing under extreme conditions, such as at the lowest and dryest levels which the yellow pines are able to endure. Such is the condition at Prescott or at the 6,000- 7,000 ft. levels on the mountains about Tucson. A broken and scat- tered rainy season may give as many as 3 preliminary red rings before the final one of autumn. In a few rare trees growing in such extreme conditions, it becomes very difficult to tell whether a ring is formed in summer or winter (that is, in late spring or late autumn). Doubling has become a habit with that particular tree— a bad habit—and the tree or large parts of it cannot be used for the study of climate. But let us keep this clearly in mind: This superfluous ring formation is the exception. Out of 67 trees collected near Prescott, only 4 or 5 were discarded for this reason. Out of perhaps two hundred near Flagstaff, none have been discarded for this reason. Neodly a hundred yellow pines and spruces from northwestern New Mexico have produced no single ease of this difficulty. The sequoias from California, the Douglas firs from Oregon, the hem- locks from Vermont and the Scotch pines from north Europe give no sign of it. On the other hand, 10 out of 16 yellow pines from the Santa Rita Mountains south of Tueson have had to be dis- earded and the junipers of northern Arizona have so many sus- picious rings that it is almost impossible to work with them at all. Cypress trees also give much trouble. THE ANNUAL RINGS OF TREES 9 The other difficulty connected with yearly identity is the omis- sion of rings. Missing rings occur in many trees without lessening the value of the tree unless there are extensive intervals over which the absence produces uncertainty. A missing ring here and there ean be located with perfect exactness and causes no uncertainty of dating. In fact, so many missing rings have been found after care- ful search that they often increase the feeling of certainty in the dating of rings. Missing rings occur when autumn rings merge together in the absence of any spring growth. This rarely if ever occurs about the entire circumference of the tree. There are a few cases in which, if the expression may be excused, I have traced a missing ring entirely around a tree without finding it. I have observed many eases in which the missing ring has been evident in less than 10 per cent. of the cireumference. Some are absent in only a small part of their circuit. I have observed change in this respect at different heights in the tree, but have not followed that line of study further. It is beautifully shown in the longitudinally bi- sected tree. One sees from this discussion what the probable errors may be in mere counting of rings. In the first work on the yellow pines the dating was done by simple counting. Accurate dating in the same trees (19 of them) later on showed that the average error in counting through the last 200 years was 4 per cent., due prac- tically always to missing rings. A comparison in seven sequoias between very careful counting and accurate dating in 2,000 years shows an average counting error of 35 years, which is only 1.7 Der cent. Full confidence in yearly identity really comes from another source. The finding of similar distribution of large and small rings in practically all individuals of widely seattered groups of trees over great periods of time has been evidence enough to make us sure. This comparison process of groups of rings in different trees has received the rather clumsy name of ‘‘cross-identifieation.’’ Cross-identification was first successful in the 67 Prescott trees, then was carried across 70 miles to the big Flagstaff groups. Later it was found to extend 225 miles further to southwestern Colorado with extreme accuracy, 90 per cent. perhaps. This is over periods of more than 250 years. Catalina pines from near Tueson have a 50 per cent. likeness to Flagstaff pines. There are many points of similarity in the last 200 years and many differences. Santa Rita pines are less hke the Flagstaff pines than are the Catalinas. In comparison with the California sequoias, differences become more common. The superficial resemblance to Arizona pines is 5 or 10 10 THE SCIENTIFIC MONTHLY per cent. only. That is, out of every 10 or 20 distinctive rings with marked individuality, one will be found alike in California and Arizona. For example, A. D. 1407, 1500, 1580, 1632, 1670, 1729, 1782, 1822 and 1864 are small in Arizona pines and Califor- nia sequoias. While only a few extreme individual years thus match, there are correspondences in climatic cycles to which atten- tion will be called later. Cross-identification is practically perfect amongst the sequoias stretching across 15 miles of country near General Grant National Park. Trees obtained near Springville, some 50 miles south, show 50 to 75 per cent. resemblance in details to the northern group. This was far more than enough to carry exact dating between these two localities. Cross-identification in some wet climate groups was extremely accurate. A group of 12 logs floating in the river- mouth at Geffle, Sweden, showed 90 to 95 per cent. resemblance to each other. The range was 100 to 200 years and there were no uncertain years at all. The same was true of some 10 tree sections on the Norwegian coast and of 13 sections cut in Eberswalde m Germany. A half dozen sections cut in a lumber yard in Munich did not cross-identify with each other. A group of 5 from a lumber yard in Christiania was not very satisfactory. The vast majority, however, have been absolutely satisfactory in the matter of cross- identification. Nothing more is needed to make the one ring a year ideal perfectly sure, but if there were, it would come in such tests as frequently occur in checking the known date of cutting or boring, with a set of rings previously dated. That has been done on many occasions in Arizona and California. To give final assurance, the record in the yellow pine was compared with statements of good and bad years, and years of famine, flood and cold, reported in Bancroft’s ‘‘History of Arizona and New Mexico,’’ and it was found that his report identified with the character of the growth in the corresponding years of the trees. Three results may be noted before leaving this important sub- ject. Deficient years extend their character across country with more certainty than favorable years. |) | OS SR DR feared S i=} @ 3 ee ; a Bt ge cS ie 3 I or e < Po w Ee Ps r © @ ws 5 C) ¢ 5 a ° 2 B cs) = 5 2 o % 4 o cy uo Lal @ Stelocyttarous = = 5 Reotinidal laterinidal E 3 g a g & a! e f <| : 4 Q & g E See * 3 o 5 © ~ ie 3 5 a gq 3 =z ° g f ¢ E g : a : - 3 - ii] 2d v o 2 8 ° 5 o g " g er o oe e bal e i] °o e | c > FIG. 29 Phylogenetic tree of the various genera and families of Vespoids. (After Ducke, with modifications). 88 THE SCIENTIFIC MONTHLY pamphlets of his library. The Pison, under natural conditions, builds elliptical clay cells and provisions them with spiders, whereas the species of Trypoxylon nest in hollow twigs and the interstices of wall but use the same kind of prey. All these species adapted themselves to the glass tubes in the same manner. Each of them plugged the end of the tube with clay and divided the lumen into successive cells by building simple clay partitions across it. After thecells had been provisioned, Bordage observed that the first of them were longer by half a centimeter and con- tained more prey than those provisioned later, and he was able to show that the larve in the larger, more abundantly provisioned cells produced female, the others male wasps. Similar observations have also been published by Roubaud on the Congolese Odynerus (Rhynchium) anceps, which makes clusters of straight, tubular galleries in clay walls and divides each gallery into several cells by means of clay partitions. In this case also the first cells are much longer than the later, though there is no difference in the quantity of small caterpillars allotted to the different eggs. But Roubaud was able to prove experimentally that even when the amount of food is so greatly decreased that the larve produce adult wasps of only half the normal size, their sex is nevertheless in no wise affected. It would seem therefore that the mother wasp must discriminate between the deposition of a fertilized, female- producing and that of an unfertilized, male-producing egg, and regulate the size of the cell and in some instances also the amount of provisions accordingly. In the accompanying diagram (Fig. 29), taken from Ducke but somewhat modified, I have indicated the hypothetical family tree of the solitary and social Vespoids. The genera below the heavy horizontal line are solitary, and among them Eumenes and Odynerus seem to be nearest to the original ancestors, because they are very similar to the social forms in having longitudinally folded wings and in other morphological characters. It will be seen that there are six independent lines of descent to the social forms above the heavy line and that the genera plotted at different levels represent various stages of specialization as indicated by the nature of the materials and types of structure of the nests. With the doubtful exception of a few Stenogastrine, all the social wasps make paper nests consisting wholly or in part of one or more combs of regular hexagonal cells, in which a number of young are reared simultaneously. (To .be continued) THE PROGRESS OF SCIENCE 89 THE PROGRESS OF SCIENCE' INTERNATIONAL COOPERATION IN INTELLECTUAL WORK Steps have been taken toward the formation of a committee of the League of Nations on international cooperation in intellectual work. Eleven of the twelve members have been appointed and as none of them is an American, it is expected that the vacancy will be offered to an American scholar. The committee so far chosen con- sists of Henri Bergson, the French philosopher and author of ‘‘ Creative Evolution ’’; Curie, the Polish discoverer of radium; Albert Einstein, the German mathematician who propounded the theory of rela- tivity; Gilbert Murray, professor of Greek at Oxford; Miss Bonnevie, pro- fessor of zoology at Christiania; D. B. Bannerjee, professor of polit- ical economy at Calcutta; A. De Cas- tro, of the medical faculty of the University of Rio de Janeiro; J. Destree, former minister of science and art in the Belgian cabinet; G. De Reynold, professor of French litera- ture at Berne; F. Ruffini, professor of ecclesiastical law at Turin, and L. De Torres Quevedo, director of the electro-medical laboratory of Madrid. The first meeting of this committee is set for August 1, and a prominent position on the program of work out- lined is given to measures that will facilitate the interchange of scientific information and the development of higher education in the countries par- ticipating. With regard to the organization of intellectual work from an inter- national standpoint the report adopt- ed by the council of the League of Nations when the committee on inter- Madame 1 Edited by Watson Davis, Science Service. national cooperation in work was organized says: intellectual We are all agreed that the League of Nations has no task more urgent than that of examining these great factors of international opinion—the systems and methods of education and scientific and philosophical research. It would be unthinkable that the league should endeavor to improve the means of exchange of material products without also endeavoring to facilitate the international, exchange of ideas. No association of nations can hope to exist without the spirit of reciprocal intellectual activity be- cween its members. For example, it is clear to all how much the league would benefit by any new measures which by establishing a more definite parallelism between the diplomas of the various countries and a more frequent exchange of chairs between professors of various nationalities would lead to a more active interchange of teachers and students between nations. A_ still greater benefit would result from measures which permitted a more rapid and more accurate communica- tion of all work undertaken simul- taneously in the field of scientific re- search in various parts of the world. There is no question of detracting from the originality of national workers whose very diversity is essen- tial for the general progress of ideas. On the contrary, the object is to en- able each of these national thinkers to develop his ideas with greater force and vitality, by making it possible for him to draw more fully upon the common treasure of knowledge, meth- ods and discoveries. As a part of the work of the League of Nations, a ‘‘ Handbook of International Organizations’’ has re- cently been issued, which lists 315 societies, associations, bureaus, com- mittees and unions, all of them inter- national in some aspect. It is an interesting collection of religious, scientific and other sorts of organiza- tions, the association interested in lawn tennis being listed with the entomological, meteorological international 90 THE SCIENTIFIC MONTHLY PROFESSOR SANTIAGO RAMON Y CAJAL The distinguished Spanish histologist who retires from the chair of histology and pathological anatomy at the University of Madrid on reaching his seventieth year. THE PROGRESS OF SCIENCE ot and other scientific societies. Such a directory is a necessary preliminary of the activities of the committee on international cooperation in intellec- tual work. CALENDAR REFORM RerorM of the calendar has been much discussed during the past decade or more, for the inconveniences and inconsistencies of the present calen- dar are obvious. The two schemes which are receiy- ing the largest amount of attention are the international fixed calendar plan and the Swiss plan. The former, first publicly proposed by Moses B. Cotsworth of Vancouver in 1894, provides for thirteen months in the year, with twenty-eight days to the month, every date being at- tached to the same day of the week in every month. New Year’s Day is a zero day called January 0, and is a full holiday. The extra day in leap year is a similar holiday inserted as July 0.. The extra month, which, of course, does not. add to the actual length of the year, is introduced be- tween June and July, and is called *“Sol.’? Easter is to be fixed by the Christian churches on some date be- tween March 21 and April 26, this stabilizing an event whose drifting causes inconveniences and losses in business and social life. The Swiss plan has been advocated largely by astronomers. It also sets aside each New Year’s Day and each leap-year day as independent legal holidays. The other 364 days are divided into four quarters of 91 days each, each quarter containing one month of 31 days and two months of 30 days, thus keeping twelve months as at present. The international fixed plan recently received the unanimous approval of a convention held in Washington by those interested in calendar reform. The American sec- tion of the International Astronom- ical Union, after considering both the calendar Swiss plan advocated by its com- mittee on calendar reform and the fixed calendar plan, recently refused to take action on the matter. The question of calendar reform was taken up at a meeting of the International Association of Acad- emies held in St. Petersburg in 1913, and a committee was appointed on that occasion ‘‘to study questions relative to the unification and sim- plification of the calendars and the fixing of the date of Easter.’’ This committee would have made a report in 1916, but for the war. Another discussion of this subject took place at the International Geographical Congress held in Rome in 1913. In June of the same year the World Congress on International Associa- tions, which met at Brussels, passed a resolution urging the governments of the world to adopt a universal eal- endar. Three of the International Congresses of Chambers of Commerce have given expression to ‘the same de- sire. Finally, just before the out- break of the world war, the Interna- tional Congress on the Reform of the Calendar held its sessions at Liége, and not only agreed to urge the adop- tion of a universal and improved eal- endar but also made plans for a for- mal conference, which was to have been convoked in Switzerland at the invitation of the Swiss government. but was never held. In the future there may come a conference of nations that will adopt a new and more logical calendar as easily as standard time was estab- lished by an international conference at Washington about forty years ago. INVISIBLE SUN-SPOTS Dr. GEORGE ELLERY HALE, director of the Mount Wilson Observatory, has announced the discovery of invis- ible sun-spots. In 1908 Dr. Hale found that a sun-spot is a great whirl- ing storm, similar to a_ terrestrial tornado, but on a gigantic scale, often vastly larger than the earth. The ex- TOOHOS OITdNd SHTHONV SOT V DNIGNYLLV SNIML "sovoud PHOM PIM THE PROGRESS OF SCIENCE 93 pansion of the hot solar gases, caused by the centrifugal action of the whirl, cools them sufficiently to produce the appearance of a dark cloud, which we call a sun-spot. If this cooling is not great enough to produce a visible darkening of the surface, the whirl- ing storm may still be present, though invisible to the eye. Such invisible whirls have now been detected by their magnetic effect on the light emitted by the luminous vapors within them. Magnetic fields in visible sun-spots were first found by Dr. Hale in 1908. They are due to the whirl of electri- fied particles in the spot vortex, just as the magnetic field of an electro- magnet is produced by the whirl of electrons through its wire coils. The magnetic field in a sun-spot is recog- nized by the effect it produces on the lines in the spectrum. A line due to iron vapor, for example, is split into three parts by the powerful magnetic field in a large spot. In a very small _ spot, where the magnetic field is much weaker, the line is not split up but is merely widened. Invisible spots were discovered by exploring promising regions of the sun where signs of disturbance, such as facule or clouds of calcium vapor, are present. A special polarizing ap- paratus moves back and forth across the slit, while the iron line is watched through a very powerful spectroscope. The presence of a weak magnetic field, showing the existence of an invisible spot, is betrayed by a slight oscillation of the corresponding part of the line, caused by its widening successively to right and left as the polarizing apparatus oscillates over the slit. Ten invisible spots have been found since November by this method by Messrs. Hale, Ellerman and Nichol- son with the 150-foot tower telescope and 75-foot spectroscope on Mount Wilson. Some of them foreshadow the birth of a visible spot, which finally appears to the eye several days after the first indications of the whirl have been found, Others correspond to the period of decay, and permit a spot to be traced for some time after it ceases to be visible. In other cases the invisible spot never reaches ma- turity, which means that the cooling produced by expansion never becomes great enough to produce perceptible darkening of the sun’s disk. TWINS AGAIN THE popular interest in twins seems to have considerable vitality. Every year brings into the public press and magazines some news item or article concerning multiple births. Just a year ago the whole country was stirred by the announcement of the birth of quadruplets in New Haven, Connecticut. (By the way, they have all passed their first birth- day). Recently the newspapers car- ried full accounts of the death of the conjoined Blazek twins of Chicago, recalling the older days when the Siamese twins were in the prints and broadsides. Now comes Los Angeles, with photographic evidence that in one school ‘building are enrolled as many as nine pairs of twins. And on the heels of the City of Angels comes the City of Churches, Brooklyn, with a contingent of ten pairs of twins, all attending Public. School 77. Some statistician may soon find for us a rural school in which 30 per cent. or more of the entire enrollment are twins. - After all, twins are more common than we ordinarily suppose; and our interest in them far exceeds their rarity. Wappeus found that more than one child was born in 1.17 per cent. of 20,000,000 cases of labor. Pre-war Prussian statistics showed that twins occurred once in 89, trip- lets once in 7,910, and quadruplets onee in 371,125 labors. This does not, of course, mean that all survive. The hazards of birth and of both prenatal and neonatal life are greater for plural than for singular pregnancies. TOOHOS Ol Tdnd NATMOOUd V ONIGNALLVY SNIML ‘SO1OUd PLOA\ apt Ay oe RRA i A AI sean wewe do aoe “Po een ne THE PROGRESS OF SCIENCE 95 A comparison of international statis- tics appears to indicate that multiple pregnancy is more common in cold than in warm climates. We should, therefore, not be Brooklyn has a higher record than Los Angeles! If the figures quoted by Williams can be trusted, Russia is in this special sense over twice as prolific as Spain. In Russia, multiple birth occurred once in 41.8 labors, as compared with once in 113.6 labors in Spain. The accompanying photographs raise some interesting questions in regard to the distribution of sex among twins. In the aggregate 38 children are pictured, of which 18 are boys, and 15 of these happen to be in the California group. This ap- proximates the equal or nearly equal divisions which we should expect when all the twin boys and girls in the land are counted. But there are other interesting questions.. Suppose the Brooklyn group were playing helter- skelter in the school yard. Would it be possible for a stranger to select all the pairs of twins and match each to the appropriate co-twin? It hap- pens that there is such a predom- inance of same-sexed similar twins that this could be readily done. Sup- pose, however, that the Los Angeles group were scrambled in the same manner, would it be possible by in- spection to pair off all the twins? This is doubtful. It would be par- ticularly difficult to make a confident decision in the case of the four chil- dren in the back row, beginning with the third from the left. This diffi- culty brings out clearly the fact that there are. marked differences as well as resemblances between twins. We, of course, always expect at least some degree of family resemblance, but even this may not be obvious to or- dinary observation. The problem of twin resemblance is discussed by Dr. Arnold Gesell, professor of child hygiene, Yale Uni- in two recent articles on surprised versity, that: ‘*Mental and Physical Correspond- ence in Twins,’’ published in the April and May numbers of THE ScrentTiFic MontHiy. He describes a remarkable case showing extensive, detailed correspondence both in phys- ical and psychological characters in a pair of gifted girl twins. He notes, however, that pathological deviation in the process of twinning may pro- duce monstrous degrees of individual difference even in twins derived from Biologists recognize two major classes of human twins: (1) Duplicate or uni-oval, who are always of the same sex, closely re- a single egg. semble one another and presumably originate from one fertilized egg. All but one of the Brooklyn group ap- parently belong to this class; (3) Fraternal or bi-oval twins, who may or may not be of the same sex, who show ordinary family resemblance and are in all. probability derived from two separate eggs. Two pairs, at least, of the Los Angeles twins belong to this category. Statistics based on a large series of cases indicate that one pair of twins in every three pairs born consists of a boy and a girl, and that about two out of every five pairs in which the members are of the same sex are uni-oval in Since there are only two pairs of two sexed or ‘‘pigeon’’ twins in the combined group of 18 pairs in the photographs, we must again be cautious in drawing general deductions from the pictures. Do resemblances decrease with age? Such a deduction might be drawn from the Los Angeles photograph, but it would not be well supported by the facts. Two pairs of twins of the fraternal type happen to include the older children in this group, and re- semblances are less marked in this type. The fundamental correspond- ences, both physical and mental, to be found in twins unquestionably have a hereditary basis and are only in @ secondary way affected by time and experience. Time may, by a cumula- tive process, accentuate a differentia~ origin. 96 | THE SCIENTIFIC MONTHLY tion of twin personalities dating from childhood or youth, but the primary differences and resemblances are due to original nature. The study of twins does not shake our confidence in the importance of education and surroundings, though it impresses upon us at every turn the decisive sig- nificance of inheritance and the law- fulness of the mechanics of develop- ment. The popular interest in twins is a wholesome one, because twins are a key to many biological and psycho- logical principles at the basis of human welfare. SCIENTIFIC ITEMS WE record with regret the death of Henry Marion Howe, emeritus pro- fessor of metallurgy in Columbia University; of John Sandford Shear- er, professor of physics at Cornell University; of George Simonds Boul- ger, the English writer on botany; of Ernest Solvay, known for his process for the manufacture of soda; and of Cc. L. A.-Laveran, of the Pasteur Institute. AMONG five busts unveiled in the Hall of Fame for Great Americans at New York University on May 20 was one of Maria Mitchell, the gift of her Wiliiam Mitchell Kendall, and the work of Emma 8. Brigham. President Henry Noble McCracken, of Vassar College, where Miss Mitch- ell was professor’ of astronomy from 1865 to 1888, unveiled the bust. Dr. Ray LYMAN WILBUR, president of Stanford University, has elected nephew, been president of the American Medical Association for the meeting to be held next year at San Fran- cisco. THE Croonian lecture was delivered before the Royal Society on June 1, by Dr. T. H. Morgan, professor of zoology in experimental Columbia University. His subject was ‘‘The mechanism of heredity.’’ Dr. W. W. CAMPBELL, director of the Lick Observatory, has been elect- ‘ed president of the International As- tronomical Union in succession to M. Baillaud, director of the Paris Ob- servatory. The Astronomical Union. held its triennial meeting in Rome in May and will hold its next meeting in Cambridge, England. Iv is announced that the contest of the will of Amos F. Eno will be set- tled out of court by the payment of about four million dollars to Columbia University. The 1915 will, which has been twice broken by juries but both times upheld by courts on appeal, gave the residuary estate to Columbia University. The will made bequests of $250,000 each to the Metropolitan Museum of Art, the American Mu- seum of Natural History, the New York Association for Improving the Condition of the Poor, and the New York University. Had the will been broken finally, these institutions would have received nothing. Whether they receive the full $250,000 each under the settlement, or what propor- tion of the total they receive, is not disclosed. The Society of Mechanies and Tradesmen reeeived $1,800,000 under the 1915 will, and had that will been broken would have received $2,000,000 under an earlier will. This institution could not therefore be called upon to sacrifice anything in order to satisfy the heirs, and will receive the full $1,800,000. WE much regret that there was an error in the inscriptions of the illus- trations of the note on Hesperopithe- cus in the last issue of this journal. Fig. 2 on page 589 is the important type tooth, whereas Fig. 1 is the sec- ond molar of Hesperopithecus which serves to confirm the first of the type. ee ee THE SCIENTIFIC MONTHLY AUGUST, 1922 THE GEOLOGIC EVIDENCE OF EVOLUTION By Professor EDWARD W. BERRY THE JOHNS HOPKINS UNIVERSITY NE of the outstanding, possibly the only difference between man and the other animals is his ability to profit by the ex- perience and accumulated wisdom of the race, and yet, despite this characteristic, each generation seems to produce its quota of anti- vaccinationists, anti-evolutionists and believers in a flat earth. We may still entertain the hope that the race is becoming more rational when we recall that it has taken about three centuries to convince the Anglo-Saxon and a few other races among the countless mil- lions of the globe that the earth is not flat, so that to-day only the leader of Zion City (Voliva) among leading cosmologists defends the pentateuchal view. I do not wish to be thought of as sneering at any one’s beliefs, and I fully realize that there are a great many earnest Christian men and women who are perturbed at anything that they think, rightly or wrongly, will shake the foundations of their faith, who are puzzled by the present outspoken opposition to evolution, and who wish to know what is the truth. No truer article of faith was ever penned than the motto of the Johns Hopkins University— Veritas vos liberabit—and to you seekers after truth I would like to explain away certain misconceptions, before undertaking to show you that the record of earth history is the record of evolu- tion, and not to be disputed by honest people. Evolution *,jnot a theory of origins, nor an article of scientific faith, but an i Aisputable fact. We could not teach geology with- out teaching e*olution. One of the difficulties to the layman is the confusion of evolution—the record of the past and present history of organisms—with the various theories that have been proposed to explain its factors or mode of operation. Let me emphasize that evolution, the record, is in an altogether different category from the theories such as Darwinism, Lamarckianism, or any other VOL. XV—7 98 THE SCIENTIFIC MONTHLY ism that has been advanced to explain its working. You may flout all the theories or you may advocate one of a dozen different theories, but this has nothing to do with the history of life. We, in geology, spend much time in going over the history of organisms, but pay but shght attention to the theories—at least in our teach- ing. A simple illustration of the once universal and now fortunately less frequent clerical reaction to evolution will make clear what I am driving at. Evolution was regarded as a dangerous heresy, inimical to Christianity, contrary to Genesis, which was regarded as a scientific account of the origin of the earth and its inhabitants. Do these people claim that the hundreds of varieties of horses, dogs, chickens and pigeons go back to the Garden of Eden, or were in Noah’s ark, or that all the horticultural varieties of flowers, shrubs and vegetables were in Mother Eve’s kitchen garden? Not at all! They are more or less familiar with the cattle breeders or the Burbank method of artificial selection. Their objection to evolu- tion rests on the assumption that man is of a different stuff from the brute world—as if they had had no experience with congre- gations or legislative assemblies. It is the implied collateral rela- tionship with monkeys, and the tradition engendered by medieval art that the devil has a tail that offends their dignity. The statement that the human species is descended from monkeys is merely polemical obscurantism and the playing on prejudices that started with Bishop Wilberforcee—soapy Sam as he was called by some of his contemporaries—and is a sort of Bryanesque smoke screen. As to lineage, man is not at all closely related to the existing monkeys or apes. They are the culmination of different lines of evolution, and this statement is especially true of the monkeys. That their ancestry in the far distant past ap- proximated the human line or indeed may have merged with it millions of years ago in early Tertiary times is quite another matter. I find nothing in Genesis either for or against evolution. The language, to be sure, is not explicit (dust of the earth), but the special creation of man as opposed to the evolutionary creation is entirely an egoistical interpretation that is supposed, quite wrongly it seems to me, to add dignity to ourselves, and RS of a cloth with the idea that the earth is the center of the univeyse—all the earth (homocentric) centering in man, and all the universe revolving around the earth—man’s temporary abode. It is a most curious revelation in the workings of the human mind that so many good people grow indignant over the idea that man was made from a long line of animal ancestry as degrading; and yet who do not GEOLOGIC EVIDENCE OF EVOLUTION 99 quarrel with the facts that each human starts his or her individual life as a single cell, and during the nine months preceding birth passes through a series of stages that roughly epitomize the main stages of evolution, even to possessing a rudimentary tail like an ape. Five hundred years ago we should have said that embryology was the invention of the devil to test the faith of the elect—exactly a reason that was once advanced to explain the fossils in the rocks. To-day most of us know better, and we find in the truth of creation far more to reverence than in the anthropomorphic deity of the childhood of the races. In approaching the geological record of evolution, I will state only facts and leave fundamental causes severely alone. The mechanism of evolution we leave to experimental biology, and I do not advocate any theories of explanation. Here is evolution. Here are the myriad of forms that moved across the stage of the past and were the actors in the drama of life. In geology, to bor- row a simile from written history or philology, we are dealing with the original documents in so far as they were preserved as fossils, and in their actual order of succession. In approaching the geological record, the time conception is most important, and I can best illustrate this by a brief recital of the progress of knowledge concerning fossils. It is only in compara- tively modern times that fossils have been recognized as the remains of animals and plants that had once been alive. The early Greeks were sane enough to recognize this apparently obvious relationship, and we find Xenophanes, 500 B. C., speculating on the fossils found in the quarries of Syracuse, Sicily. But during the middle ages there was no end to the discussion regarding the nature and origin of fossils. What seems strange in this year of grace may really not have been so strange in the days when the universally held belief was that of spontaneous generation, a flat earth created in six days, and the only past submergence of the land that of Noah’s flood. Was it not the same ‘‘plastie foree’’ in nature which traced the frost patterns and the moss agate that fashioned the fossils, and was there not every gradation from shells and bones that exactly resemble recent ones to mere stones of similar form and appearance? We now know that the mineral replaces the organic matter sf a fossil. Was it strange to have believed three or four hundred years ago that the process was the reverse— from the mineral toward the organic? At any rate many strange theories were evolved to explain the fossils. One tells us that fossil shells were formed on the hills by the influence of the stars. Others called up a stone-making spirit. Others believed that fossils were the models made by the Creator in perfecting his handiwork before 100 THE SCIENTIFIC MONTHLY he essayed the task of making living organisms. I am quoting en- tirely the views of devout churchmen. Others believed that fossils were mere ‘‘figured stones,’’ or were the abortive products of the germs of animals and plants that had lost their way in the earth, or that they were the invention of the devil to test the faith. Even ~ after the belief that fossils were the remains of animals and plants had become well established, it was assumed that they had been killed by Noah’s flood and stranded on the mountain tops—an in- terpretation suggested by Martin Luther in 1539 as secular proof of the correctness of the scriptural account. This flood theory found numerous advocates throughout the seventeenth and even far into the eighteenth century. It passed through various phases of opinion. At first, the fossils were regarded as similar to those still living in the vicinity—a natural enough belief when the uni- versal acceptance of the Mosaic cosmology and a world but 6,000 years old is borne in mind. Later, when the differences in the fos- sils became apparent, it was assumed that they had been swept to Europe and buried by the waters of the flood and represented forms still existing in the tropics. With the progress of knowledge of tropical organisms this last view became untenable, and it was thought that the fossils represented forms that had been exter- minated by the flood, and from this it was but a slight step to the once popular belief that there had been no thistles or weeds or noxious insects in the Garden of Eden, that all creation had become base with the fall of man. Gradually it came to be recognized that fossils were not only frequently unlike recent organisms, but that they were very ancient, and not merely antediluvian, but pre- Adamitie—a view first advocated by Blumenbach in 1790. We are still far from a ehronology. Granting that fossils were the traces of once living organisms and antedated Adam—what of it? When Guettard (Jean Etienne Guettard, 1715-1786) made one of the first geological maps, it wasn’t really a geological map in the modern sense, but a map of what he called mineral bands (like a modern soil map). He had no idea of geological succession or of structure. The credit of recognizing fossils as the modals of crea- tion we owe to the genius of William Smith (1769-1839) and to the orderly arrangement of the Mesozoic rocks of the English Mid- lands. Smith journeyed about for years in this. region, where the suecession of fossiliferous strata is an open book. In his work of building canals, roads and drains, he observed that each bed con- tained fossils, some of which were peculiar to it, and he found that he could recognize the same horizons and the same succession at many different localities. This important generalization has since been verified and end- GHOLOGIC EVIDENCE OF EVOLUTION 101 lessly extended. The contained fossils furnish the surest guides to the age of the sedimentary rocks that geology knows. To the biologist these facts have a deeper meaning, for they show that dur- ing the vast lapse of time, to be measured in tens or hundreds of millions of years, the living population of the globe has undergone almost continuous change, old simple forms becoming extinct, and newer, more specialized, forms taking their place, the change being, in general, from lower to higher, in other words—evolution. That God rested from his six days’ task of creation just 4004 years B. C. is so absurd that I have yet to meet a person of normal mind who believes in Archbishop Usher’s chronology. There have been many attempts to determine the age of the earth in years— ealeulations of the rate of cooling of molten bodies, the rate of re- tardation by tidal friction, the thickness of the sedimentary rocks, the amount of dissolved salts added to the oceans by the rivers of the world, the condition of the radium minerals in igneous rocks. All methods contain unknown variables and are merely estimates. A favorite method has been to measure the thickness of a composite section of the sedimentary rocks, for the whole VU OIOZONAD ARCHEOZOIC ERA (Pritaal tile) of unic Age eliulur life nt life) (Ancic PALFEOZOIC ERA PROTEROZOIC ER (Pr a FIG. 1. GEOLOGIC TIME CLOCK 102 THE SCIENTIFIC MONTHLY record is not complete in any one section—seas and sediments shifted with the incessant change in geographic pattern. If they had not, we should have such a perfect record of evolution with no missing chapters that we should be able to establish geological time boundaries between rock formations or biological boundaries between animal and plant groups. The measurement of thicknesses has this advantage, that where- as its results expressed in years are not accurate, its results ex- pressed in relative ratios of duration for the different geological periods are fairly so. I have sought to show the totality of geo- logical time reckoned in this way in the form of the face of a clock in which the dial represents the total thickness of sedimentary rocks divided among the different geological periods in the proper ratios. With this perspective I wish to pass in review in an un- technical way some of the facts of evolution. Obviously, one can not go into details in a brief hour, nor present the links in the chain of evidence, or talk about the septa and sutures of the ammonites, the pygidia of trilobites; or the frontal, parietal, temporal and other bones of the vertebrates. Show us one species changing into another, and we shall be- lieve in evolution, says the bigot, expecting to see an Alice-through- the-looking-glass transformation of cats into dogs or rabbits into porecupines, not realizing what a species is, or the slowness with which very obvious new characters are acquired as measured in terms of human years. If they had been present through any 70 years of geological time, they would have seen no more evidence of evolution than they see to-day. The first man to see the trans- formation of species was Waagen,? an Austrian geologist and paleontologist, who, in 1869, in the successive layers of fossiliferous Jurassic rocks, observed the minute and inconspicuous changes of form in a definite direction, resulting as they increased in magni- tude in the gradual emergence of successive new species of am- monites (Oppelia). These observed grades of difference or nuances (Waagen termed them mutations) are the more gradual and in- eonspicuous the more abundant the material studied, or the finer our analysis of it. This observed gradual evolution of adaptive characters is quite the opposite of Darwin’s theoretical idea of the natural selection of chance variations, and its abundant verification among all groups of fossil organisms wherever an abundance of successive faunas or floras are available for study is one of the reasons why paleontologists have never been strong Darwinists, but 2 Waagen, Wilhelm Heinrich: Die Formenreihe des Ammonites subradi- atus. Versuch einer Paliontologischen Monographie. Geognostisch-Palaon- tologische Beitrage. Bd. 2, Hft. 2, pp. 179-256, pls. 16-20, November, 1869. GEOLOGIC EVIDENCE OF EVOLUTION 103 have emphasized the environment as the main stimulus of variation. Discontinuity is observed only in characters where continuity is impossible, as in changes in the number of teeth or vertebra. I could spend days showing you these evolutionary series of trilobites, brachiopods, crinoids, molluses, ete., but they are not especially convineing without fullness of knowledge and presentation, and are not nearly so impressive to a lay audience as the more ob- viously discerned, but identical, series among the higher verte- brates. There is probably no group of organisms as ideal for evo- lutionary studies as are the Ammonites—extinct relatives of the pearly Nautilus. Their shells are preserved in tens of thousands in the Mesozoic and earlier rocks. From the time the embryo formed its first shell until death, each successive stage is preserved in calcite within the enrolled shell. If you would see the size, form and details of ornamentation of a baby, adolescent or mature shell, all you have to do is to break away the outer shell. No other fossils furnish a complete life history with each individual. Moreover, the repetition of phylogeny during ontogeny is beauti- fully shown, as well as the inheritance of acquired characters, so RECENT CENOZOIC SALURIAN ORDOVICIAM FIG. 2. THE EVOLUTION OF THE CEPHALOD PHYLUM 104 THE SCIENTIFIC MONTHLY that we know the ammonite descent much better than we do that of many still existing groups of organisms. The main outlines of the evolution of the Cephalopods, to which group the ammonites belong, is shown in its chronological setting in Figure 2. Observe the gradual transformation of straight camerated shells becoming curved, then loosely coiled, then tightly coiled, giving rise to forms with angulated floors to the living chamber, the parent stock waning with the rise of the daughter stock, and represented to-day by a single living type, the pearly Nautilus. This daughter stock waxing great during Mesozoic times, we know 10,000 different species, gradually reaching overspecialization or racial senility, displayed in the progressive uncoiling and bizarre ornamentation of the shells, and finally passing off the stage altogether. A second main line of descent leads from the ancient Nautilus stock in the direction of animals whose soft parts outgrew their shells, retain- ing them within the mantle. This second line waxed abundant during Mesozoic time, and then waned in competition with its more perfected progeny, being represented in existing oceans by the single form, Spirula, which in its extreme youth lives in a tiny chambered shell like that of its remote ancestors, but soon outgrows this shell, and for the rest of its life carries this eloquent witness of its ancestry within the hind end of its body. You might remain incredulous before a single Spirula, but when you ean trace throughout the records of hundreds of thousands of years the gradual subordination and progressive decrease in relative size of the shell and increase of the soft body, the meaning is unmis- takable, and to corroborate the correctness of our reading of his- tory, we have the more modern group of squids and cuttles with all of the morphological features of the Spirula stock, which solved their problem by modifying the now useless shell into an internal axis of support and are otherwise entirely soft bodied and often of large size; and, finally, the latest evolved group—the Octopoda— smaller less active forms, having slight need for the axis of the more elongated and actively swimming squids, have lost all traces of the ancestral shell. Another great phylum of invertebrate animals (Echinoderma) starfishes, sea urchins and ecrinoids, have a wonderful abundance of fossil forms and well-ascertained relationships. Their history shows a worm like ancestor developing a plated exoskeleton of many irregular pieces; the progressive reduction in number and the assumption of definite form of these pieces—the radial sym- metry impressed by the habit of stalked attachment—the various lines of descent which sought to increase the food gathering mechanism by extending the parts concerned over the test or rais- GEOLOGIC EVIDENCE OF EVOLUTION 105 STELLEROIDES EOMITRPOIDEA Articulete DEVONIAN 0 OQ nN SILURIAN PALEO Agelacrinidae ne ORDOVICIAN CAMBRIAN PRE CAMBRIAN Carpoides ij Qyathooyetidae Diplevrvie ' ' { Vorme FIG. 0. THE EVOLUTION OF THE ECHINODERM PHYLUM ing them on long arms—the reversed orientation of the errant urchins and starfishes—the one time dominance of specialized erinoids—the late evolution and present abundance of the free- swimming forms with flexible skeletons—the intermediate or syn- thetic character of the earlier forms, especially well shown in the Ordovician to Lower Carboniferous ancestors of the starfishes and serpent stars—all afford an excellent chapter in nature’s record of evolution. CENOZOIC CRETACEOUS JURASSIC TRIASSIC LOWER CARBONIFEROUS DEVONIAN PALEOZOIC ORDOVICIAN CAMBRIAN ATRTRETATRA ‘wm ISTPHOVOTARTAOEA PRE : S CAMBRIAN soe yes FIG. 4. THE EVOLUTION OF THE BRACHIOPOD PHYLUM 106 THE SCIENTIFIC MONTHLY A group of invertebrates unknown to the layman, but im- mensely important to the paleontologist, whether he be interested merely in chronology or in evolution, is the Brachiopoda—the bivalved lamp shells of the ancients. The stock is very ancient and shows the most intricate series of gradating forms from the ancient hingeless Atremata, long since extinct except for a single family, which in Ordovician times modified its stalk of attachment into a burrowing organ, and from that time to the present has lived on practically unchanged in an unchanging environment of foul mud inimical to higher forms of life, sharing with the similarly reduced representatives of the two other primitive groups a record of un- modified habits or form in an unchanging environment that has enabled them to come down to the present, although all of their early relatives have long since passed off the stage of existence. Contrast the dwindling history of these families as represented by the black of their life lines with the series of forms, each step in whose history we have represented, of those which perfected hinge mechanisms—a protective device, and internal hard parts—loops and spirals for greater efficiency in collecting food and oxygen. We can see these structures grow until at the present time the few unchanged survivors of the more primitive orders are outnumbered fifty to one by the loop-bearing forms which retained the habit of protruding their so-called arms in search of food, whereas the spire-bearing forms that developed along with them in Paleozoic times had their arms fastened to their spiral supports and non- protrusible, and hence faded out of existence in the earlier half of Mesozoic times. ENOZOIC CRETACEOUS a JURASSIC ae ee CARBONIFEROUS LOWER CARBONIFEROUS = | me | ORDOVICIAN aga v CAMBRIAN PALEOZOIC aspBeLroa FIG. 5. THE EVOLUTION OF THE ARTHROPOD PHYLUM GEOLOGIC EVIDENCE OF EVOLUTION 107 A fourth great group is the Arthropoda, embracing the hosts of articulated animals whose relationships are shown, with the early evolution of the aquatic types—trilobites, crustaceans and Limulus-like forms. They exhibit the early efflorescence of the less specialized as to parts, and less protected as to armor—the trilo- bites; the relative late evolution of their terrestrial descendants— the spiders and centipedes, and the latest appearance of the aerial forms with specialized larval stages—the insects. Most interesting to see displayed among these myriad of diverse forms insects, spiders, crabs and ticks, their community of origin and the impress of their remote trilobite-like ancestry. Hither these myriads of slightly differing forms in progressive or retrogressive series represent evolution, or each slightly changed faunal and floral assemblage represents an independent act of special creation. These are absolutely the only alternatives, and the advocates of special creation, little as they seem to realize it, have to assume a creator, who every few years during a period of hundreds of millions of years mechanically fashioned new sets of organisms. Not only so, but each new set was fashioned surpris- ingly like their predecessors, sometimes with vestigial, useless or even harmful organs. It seems to me that this only logical appli- cation of the special creation hypothesis is a reductio ad absurdum, a bare statement of which is sufficient to demonstrate its obvious falsity. I would offer for the religiously inclined Henry Drum- mond’s dictum that evolution was God’s method of creation. The complete epitome of vertebrate evolution showing the range in time and relative abundance deserves a word of comment. I should like the critics of evolution to explain why the most primitive vertebrates appear twice as far back in the record as any of the others, and why the different classes appear in the actual order from the less to the more evolved—from lower to higher—the fish- like amphibians appearing during the Devonian, the reptiles dur- ing the Upper Carboniferous, and the two lines to which the latter gave rise, the mammals and birds in the upper Triassic and upper Jurassic, respectively. . Is it not most unfortunate for evolutionary sceptics that the most ancient fossil bird should be one of the best and most spec- tacular fossils—feathers and all preserved with great fidelity in the fine-grained lithographic stone of Solnhofen—and should rep- resent virtually a modified and partially feathered reptile, 25 per cent. reptile and 75 per cent. bird. About the size of a crow, the head was billess and the jaws were armed with true teeth, the wings had three free-clawed fingers, the tail was long and lizard- _ like, of 20 vertebree, with pinnate feathers and not consolidated 108 THE SCIENTIFIC MONTHLY with digitate feathers, the hind legs were wide apart and far back, with distinct tibia and fibula as in the reptiles, with the three pelvic bones distinct as in reptiles with no body feathers, the latter on only the wings, legs and tails—with feeble flight and obvious volplaning habits. (Archeopteryx or lizard tailed bird of the upper Jurassic.) Before taking up man, I have time to consider but two among the many groups of mammals whose history is almost completely known. You doubtless think of elephants in North America only in connection with zoological gardens or circuses, and yet the ele- phants were a most conspicuous element of the American fauna from the middle Miocene to the end of the Pleistocene, and numer- ous bones and teeth have been found here in Maryland. They lived in America much longer than has the human race and much longer than the bears which we commonly think of as character- istically American. The elephants were originally immigrants from the old world. They occupy to-day a somewhat isolated position among hoofed mammals and display a curious but readily under- standable mixture of specialized and primitive characters. Their specializations are in head and teeth, their conservatism is in body and limbs. To understand their ancestry, we must understand the five or six African and Asian species of the present. Their most obvious feature is the long trunk or proboscis that gives the name Proboscidia to the order. This trunk is simply an elongated nose, although it did not come into existence in the way Kipling relates. Aside from the trunk the tusks mark the elephant. These are simply much modified upper incisor teeth. The dental formula is then - e— pm— m— This is not the whole story of the teeth, however, for, if you examine an elephant’s teeth, you will rarely find more than a single immense functional grinder in each jaw ramus—the milk molars, developing serially 1, 2, 3, and followed in turn by the molars 1, 2, 3 during life—the worn ones being pushed forward and out, a contrivance for increasing the elephant’s life span, for an animal is only as long lived as its teeth. The mechanies of trunk and tusk support have specialized the head ; eranial bones are thickened and lightened, hence the difficulty of shooting an elephant in the brain. The neck is shortened to bring the head weight nearer the withers. The body is long and massive with large shoulder and hip bones. The feet are short and broad with the nail-like hoofs around the edge. Toes are five but not all hoofed (Indian 5 in front, 4 behind; East African 4 in front, 3 behind). Limb adaptations are those common to all heavy ani- mals of other stocks. Most quadrupeds have knee and elbow per- GEOLOGIC EVIDENCHK OF EVOLUTION 109 manently bent. Great weight necessitates the straightening of the limb and individual bones and the shifting of the articular sur- faces from an oblique to a right angled position. Weight of tusks causes a shortening and heightening of the skull. Shortening brings the weight arm of the lever nearer the fulcrum at the neck, and heightening lengthens the power arm and affords attachment for the increased musculature. (Modern tusks weighing 239 lbs. each are recorded.) The lengthening of the trunk makes it un- necessary for the mouth to reach the ground for food and water. The earliest known fossil elephant, only a potential elephant, was of upper Eocene age and comes from near Lake Moeris in the Fayim, and was consequently christened Moeritherium. It was small and somewhat suggestive of a tapir. The skull was long and narrow, the trunk was merely a snout, the neck was moderately long and the limbs were slender. The teeth were the most signi- J Panui 3 eant feature. Formula i> cy pms m = vw First upper incisor was 3 small and simple, the second was a downwardly directed small tusk. The third and the canine were non-functional and there were 6 grinders, simple and quadritubereular (4 cusps and 2 crests). In the lower jaw the incisors were procumbent. The first long, the second an enamelled tusk with worn chisel edge; the third and canine already gone and 6 grinders. The second stage of elephant evolution was Paleomastodon of the lower Oligo- cene of the same region. Several species are known, ranging in size from that of a modern tapir to a half grown Indian elephant. Tooth, formula = c pms ms canines have gone; the incisors are reduced to a single tusk in each jaw ramus, 2. e., two upper and two lower tusks. All the grinders are functional, but they have increased in complexity and now consist of six cusps and three erests. The trunk was still short, the head still long and narrow, the limbs heavier, but still relatively light. The elephants now spread into southern Asia and over Europe during the lower Miocene, giving rise to various collateral lines of evolution along their different routes of dispersal. They increased greatly in size and became more elephantine in appearance. They reached North America during the middle Miocene, and these four-tusked forms spread from Nebraska to Florida. The old-world stock shortened the chin and lost the lower pair of tusks during the Pliocene, giving rise to the mastodons and mammoths of the late Pliocene and Pleistocene, which reinvaded North America and ranged south- ward: to the straits of Magellan. Our mastodon survived much later than the European mastodon, and the males sometimes show vestigial tusks in the lower jaw. The mammoth was the contempo- 110 THE SCIENTIFIC MONTHLY rary of early man in Europe as the many excellent carvings of the stone age show, and probably also in North America, as some- what vaguely pictured carved bone and associated flints indicate. They were so common over the northern hemisphere at that time that we have records of 1,685 fossil tusks, averaging 150 lbs., be- ing exported from Siberia in a single year. Between 1820 and 1833, trawlers out of Happisburg, Norfolk, dredged 2,000 elephant molars from the submerged old land of the North Sea. (We had three true elephants in America during the Pleistoeene—the North- ern or Hairy Mammoth, the Southern or Columbian Mammoth, and the Imperial Mammoth, the latter standing 18 feet at the shoulder.) The family tree of our noblest of domesticated animals—the horse—has been called the example de luxe of evolution, since no animal stock is more completely known or has a more spec- tacular history. Long domesticated the modern animal is found almost everywhere that man can live, and of many breeds. As wild animals, horses are found only in the Old World in moderis times—the arid plains of Central Asia and Africa. There are several species—horses, asses, zebras and quaggas—very uniform in tooth and skeletal characters, but strikingly different in appear- ance, because of the superficial difference in coloration and in the development of forelock, mane, tail and ears. They differ from all living animals in having a single toe on each foot. Their re- motest ancestors were small five-toed plantigrade animals as were all of the earliest mammals. Hosts of fossil species are known, some extinct side lines especially adapted to certain environments, like the small mountain horses or the forest-dwelling and softer ground-inhabiting forms. Others were a part of the progressive line. The earliest known fossil horse you would not recognize as a horse. How do we know it was? By tracing backward step by step from the known. Nearly every stage of this ancestry is now complete, and we are as certain of the remote Tertiary form as we are of the present cart horse. The earliest well-known ancestral horse is the tiny Eohippus or Dawn horse of our early Hocene. It was about the size of a fox terrier, 7. ¢., 11 to 14 inches high, with a short neck, long body, arched back, short legs and small teeth. The front feet had 4 functional toes, and a splint repre- senting the first or thumb. The hind feet had 3 functional and 2 splints representing the first and fifth. It is significant that at that time the ancestral horse line is so generalized that a layman could not distinguish it from the contemporaneous ancestral rhinoceroses or tapirs. The second-stage Protorohippus of the middle Eocene was GEOLOGIC EVIDENCE OF EVOLUTION Ee about the size and proportions of a whippet hound. The thumb splint had now disappeared from the front foot, and the little finger splint from the hind foot. The weight was beginning to center on the middle toe, but it required two or three million years more to completely suppress the lateral toes. If there were time, we might pass in review each stage of horse evolution—the Epihippus of the upper Eocene, the Mesohippus of the Oligocene, about the size of a sheep, the Miocene, Protohippus, Pliohippus, Neohipparion, etc. The upper Miocene Protohippus is in the direct line and may be briefly characterized. About 40 inches high, longer head, longer teeth, deeper jaws, shortened body, longer legs and feet, only the third toe normally reaching the ground, but the second and fourth were complete ‘‘dew claws’’ and helped to support the weight on soft ground. There were many varieties of three-toed horses, and in the late Tertiary they had spread pretty well over the world, being found in South America, Europe and Asia, as well as in North America. By Pleistocene time the horses had become monodactyl, varied, abundant and wide ranging. So countless were the herds that the Sheridan formation of the West was long known as the Equus beds from the abundance of their fossil remains. When, however, America was discovered, horses had become extinct in the western hemisphere as well as in native tradition, although their bones are found associated with flint im- plements, pottery and fire refuse. They appear to have first been domesticated during the Neolithic, that is about 7000 B. C. in Europe, but probably at a much earlier date in Asia. Our modern work horse is descended directly from the European Neolithic horse, which was much like the Celtic pony. Descendants of this low-bred primitive race were distributed over Eurasia, where they are still represented by the Norwegian and Mongolian ponies. All the earlier horses of written history belonged to this type. It was improved by importations from Libya—the Arabs, for example, getting stallions and brood mares from Barbary, where the stock had suffered no ill effects during the Pleistocene glaciation, there having been no severity of climate in northern Africa. The course of evolution in the horses was not confined to the feet. It may be summarized as follows: Along with the disappearance of side toes went increase in length of leg and foot, especially the distal portion. Increased length of the lower leg and foot increased length of stride and, as the chief muscles are in the upper leg, the center of gravity was changed very little, consequently the swing was about as rapid but mechanical strain was greatly increased, so that strengthening at the expense of flexibility by consolidation of the lower leg and 112 THE SCIENTIFIC MONTHLY arm bones and conversion of ball and socket into pulley joints (ginglymoid) occurred. Lengthening of limbs for speed in grazing animals necessitates lengthening of the neck. Loss of toes was a hard ground adaptation for speed. The lengthening of the teeth which caused the deepening of the jaws was an adaptation for hard food and ensured more thorough mastication and a longer life span. Increase in size, although demanding an increased food supply, is a better defence against enemies or competitors. The evolution of the horse was from forest and swamp to grassy plains and went hand in hand with the evolution of the environment. Since monkeys are unaccountably not fashionable and we are very fond of horses here in Maryland, { show you for comparison a skeleton of a modern horse and man. Not only in the structure of all his physical parts, bone for bone, muscle for muscle, and nerve for nerve, is man fundamentally like the other mammals, but his specific organic functions are identical. We have the same diseases; we are similarly affected by the same drugs—in fact the whole wonderful advance of physiology and experimental medicine is built up on this truism. Have you ever thought of the countless generations of meat-eating humans involved in the specialization of the two human tape worms—the one passing its intermediate stage in beef and the other in pork and of which man alone is the host of the adult stage. The pre-humans were not meat-eaters, and we should not fail to take into account the improvement in nutrition in shortening the digestive processes and the stimulating properties of the proteins and their split products that a change in diet gave our ancestors the energy for other things. I have already mentioned the remoteness of man’s relationship with the existing monkeys and apes. Unfortunately, we have but slight knowledge of the earlier stages which remain hidden in the unexplored regions of Asia and Africa, to which much evidence points as the original homes of a majority of the mammalian stocks that appeared in Europe and North America during the Tertiary. But we know much of our less remote fossil ancestors. Evidences of their slowly advancing skill in the fashioning of weapons and implements, in the discovery of the bow and the uses of fire are innumerable, and their skeletal remains are found over a period estimated at from 250,000 to over a million years. We know at least two, perhaps three extinct genera of men and at least five distinet human species. All the existing races of man—white, black, red and yellow—belong to the single zoological species which we modestly call Homo sapiens. 1 should say that our knowledge of the exact stages between non-human ape-like animals and man is as complete as was the knowledge of the evolution of the horse GEOLOGIC EVIDENCE OF EVOLUTION 113 at the time of the founding of this university when Huxley lec- tured on the evolution of the horse. At the present rate of dis- covery (Piltdown man in 1911, Foxhall man in 1919 and Broken Hill man in 1920), another generation will not pass before the story is complete. ; Before relating what we now know of this story, I should like to refer to how we arrive at estimates of age in this part of the geological column—estimates which are as exact as the earlier dates of what is called the historic period. During geologic time immediately preceding the present there is conclusive evidence of a mantle of ice spreading over northwestern Europe and northern North America. This was not a single episode but a long en- during succession of glacial stages and milder interglacial stages— some of which were much longer than the time that has elapsed since the last ice sheet shrunk away from the Baltic or from the valley of the St. Lawrence. Naturally the deposits and moraines of the last ice sheet are fresher and less disturbed than the similar traces of the earlier ice sheets. By counting the annual layers in the clays in the wake of the shrinking ice of the last glaciation, we can trace and date its gradual withdrawal from the plains of Germany across the Baltic to:the Scandinavian uplands, and the more broken clay layers in the valleys of the Connecticut, Hudson and Champlain give the story for this country. Using this period of time as a unit, we calculate from a variety of criteria the dura- tion of the earlier glacial and interglacial stages. The oldest known man-like animal comes from distant Java and dates from the beginning of the Pleistocene, or from 250,000 to 1,000,000 years ago, or more precisely, twenty-five times as long ago as the interval since the last ice sheet extended across Long and Staten Islands here in the eastern United States. Evidence of human, or if you prefer so to call it, pre-human, industry in the form of rudely chipped flints and a knowledge of fire occur still earlier, and if the recent discovery of the Foxhall man in East Angelia is properly dated, we shall have unmistakable evidence of man in the late Pliocene. Our knowledge of the ape man of Java is on a sounder footing. First of all, he came from Asia along with the greater part of the considerable variety of animals and plants that are found fossil with him. The motive power was the less hospit- able climates in Asia resulting from the gradual uplifting of its ereat mountain areas in the late Tertiary. The fauna and flora ineluding the ape man drifted to the southeast down the broad valleys that at that time of emergence made a single land mass of the Malayan region. The anatomical features of the ape man are technical. Our interest centers on the brain ease and the fact that VOL. XV—8 114 THE SCIENTIFIC MONTHLY he was a ground inhabiting biped and not arboreal. The cranial capacity has been variously estimated between 850 and 950 ee as compared with 1,300 to 1,700 of the Neanderthal man of the third Interglacial period, or 650 ce, the greatest ape brain in the gorilla, which has twice the body weight of a man. The lower frontal- lobe region of this brain case shows conclusively that Pithecan- thropus possessed speech—not sounds or signals expressive of emo- tional states, but that he was capable of transmitting ideas and in- formation. In the painstaking models of MeGregor, he has man- aged to superimpose on the obvious inheritance of the brute a look of fleeting intelligence and a dumb prophetic gaze that gives promise of the great things of the then far off future, and I con- fess to feeling a more tremendous thrill in the contemplation of that empty brain case than any other fossil has invoked. A long gap in the record brings us to Sussex, England, and Eoanthropus or Dawn man of Piltdown. Discovered in 1911 the usual ignorance resulted in the destruction of most of the skeleton, as it did also in the wonderfully interesting find in the Broken Hill Mine of Rhodesia, so that only a few fragments of skull, 3 teeth, and a portion of the jaw were saved. Subsequently more frag- ments of other individuals have rewarded the most patient and painstaking search. If there is a wise Providence overhanging the world it is certainly watching over the paleontologists instead of their critics, which is rather surprising if paleontologists are as bad as they are sometimes painted, for these later finds are exactly the pieces needed to supplement the earlier, and to justify Smith Woodward’s conelusions. The Piltdown man probably lived during the long and warm second Interglacial period. With him are found very primitive worked flints of the type known as pre-Chellean, together with bones of the rhinoceros, hippopotamus, beaver and deer. The skull is about twice as thick as a modern and 50 per cent. thicker than a Neanderthal skull. Its capacity was about 1,300ce. The jaws are protruding, the chin receding, the nose flattened and the canine teeth very prominent; in fact, although the skull and brain are essentially human and denote the power of speech, the jaws and teeth are much like those of a young chimpanzee, as are certain muscular attachments of the neck and temporal regions. About the same age as the Piltdown man is the so-called Heidel- berg man, based on a single jaw found in 1907 associated with a large fauna at the base of the Mauer sands, 79 feet below the sur- face. This jaw is exceedingly massive with receding chin, but human dentition, and is generally regarded as merely an extinct species Homo heidelbergensis, although some students would erect a distinet genus, Paleonthropus, for its reception. It seems clearly GEOLOGIC EVIDENCE OF EVOLUTION 115 to foreshadow the Neanderthal race of the third Interglacial period. Passing over implements representing the evolution of human in- dustry and confining our attention to actual human bones, we must now jump from the time of the Piltdown and Heidelberg men over a blank interval, estimated at from one to two hundred thousand years, to the Neanderthal race. I say race advisedly, because some hostile critics have waxed humorous or satirical over the type skull-eap found in the Neander valley near Diisseldorf, as if that were the whole story. The earliest find of this race was a female skull found at Gibraltar in a cave in 1848, but the significance of which was not recognized until 1887. The Neander skull-cap with thigh bones and other fragments was discovered in 1856, and their description was received with indifference even by Darwin and Huxley, and it was not until a generation later, when two com- plete skeletons were found at Spy near Dinant in Belgium, 1887, that recognition of their significance became general. The appear- ance of this race in western Europe was contemporaneous with the wane of the last warm forest and meadow fauna of the Pleistocene and with the invasion of animals heralding the approach of the fourth glaciation. Hence the Neanderthal race dwelt in caves. Wells writes picturesquely, but not especially accurately, of their jackal-like habits, but the Neanderthalers were hardy, and appear to have utilized the bison, wild cattle, horse and deer for food, ousting cave bear and eave hyenas—the successive layers in the caves often tell an eloquent story of the struggles between man and beast for possession. Fire played its part and old hearths are abundantly preserved. Spears and throwing stones appear to have been the weapons used. The abundance of skeletons of this race is due to their cave habit and hence their better chance for preserva- tion. Over an interval of something like 50,000 years, if not much longer, preceding Neanderthal times, we have abundant evidence of human industry in the pre-Chellean and Chellean cultures repre- sented by flint implements, but these open-air nomads either threw their dead to the hyenas or buried them in the river terraces on which they dwelt, where the chances of preservation and fossiliza- tion were remote. Homo neanderthalensis, primigenius or mousteriensis as it has been called, has been discovered at over twenty different localities. Skeletons of men, women and children and of many individuals have been collected, so that the earlier critics of the type material who pronounced them merely pathological, 7. e., a diseased modérn man, are completely and absolutely refuted. In many of their im- portant features this race was more ape-like than human, but their teeth were decidedly human; they possessed the power of speech, 116 THE SCIENTIFIC MONTHLY fashioned skins and weapons, were skilled in the use of fire, and practised ceremonial burial, placing implements with their dead, the first appearance in the geological record of a belief in a future existence, so that we can not cut them off from us and say they were apes and not men. Let us get a good picture of this race that lived in Europe longer than have the Anglo-Saxons. They were short and thickset—the tallest skeleton indicates a height of 5 feet, 514 inches; with very broad shoulders and muscular robust torso, big hands, short fingers, and not entirely perfected thumb joints. They were clumsy on foot, with ape-like legs, in that the shin is relatively short and the thigh long (shin 76 per cent. of thigh). Their knees were habitually bent, and they were squatters instead of sitters when resting or working, as shown by the facets on the ankle bone (astragulus). The forearm was relatively short, like the modern Eskimos, Lapps and Bushmen. That they were far removed from contemporary apes is shown by their arms being but 68 per cent. the length of their legs. In apes the reverse prevails—the chim- panzee’s arm is 104 per cent. the leg length. The position of the foramen magnum, and the neck vertebra indicate stooped shoulders with the head held well forward, and a spinal column curved like that of a modern baby. The head was massive, with deep face, retreating forehead from heavy overhanging brows (platycephal- ic), with broad flat nose, long upper lip, prognathous jaws and re- ceeding chin. The skull was thick, but capacious. The jaw was similar to, but less massive than, that of the Heidelberg man of the second Interglacial. Let me point out that if you should find a modern skull with some of the ape-like features of the Neander- thal skull, it would prove nothing. Some of these ape-like features do occur in recent rare individuals of the lower races—they are all present in the Neanderthal skulls that have been discovered. I have said that the skull was capacious—the limits of variation are 1,300 to 1,700 ce (existing man, 950 to 2,020 ec). The size of the Neanderthal brain was therefore entirely human, but I need not emphasize that a large head does not necessarily offer anything except a field for tonsorial art, and what critics fail to take into account is that the Neanderthal brain, although it had quantity, lacked quality—its proportions were decidedly different from a modern brain—those parts conceriied with the higher faculties were less developed and with simpler convolutions—this is not infer- ence, but is based on the actual configuration of the interior of the brain case (we even know that they were right-handed). Over 50 sites of Neanderthal industry are known in western Europe (see map) and their implements increased in variety and improved in technique as the years passed, but not to any remarkable degree. GEOLOGIC EVIDENCE OF EVOLUTION ALG Some anthropologists hold that the Neanderthal race is represented by the Briinn and Piedmont races of the upper Paleolithic, others that they were exterminated by the arrival in western Europe of a new race from Asia about 25,000 years ago. This progressive race of Homo sapiens, the same species as ourselves, appears to have come from Asia Minor through Tunis into Spain, and per- haps along the northern shores of the Mediterranean as well. Their successive cultures are known as the Aurignacian, Solutrian, Magdalenian and Azilian, and the development of their industry and art has been traced with the most detailed precision. They were hunters, and followed in the trail of the wild ass, Elasmo- ‘there, steppe horse and various other Asiatic immigrants. Asso- ciated with fourteen Cro Magnons skeletons in the Grotto on the Riviera near Mentone are two negroid skeletons. (I. will not stop to describe this negroid Grimaldi type.) The fourth Glacial period had not yet closed when the Cro-Magnons apeared in Eu- rope, but the climate was dryer—the summer temperate, but the winters severe. Most of the stations where their remains have been discovered were in caves or rock shelters, but several open camps have been discovered, as at Solutré which was probably a summer assembling of hunters. This remarkable race was tall, the average height 6 ft. 144 inches, with large chest, relatively long legs, re- markable lengthening of the forearm and shin, wide short face, ' prominent cheek bones, narrow pointed chin, narrow skull, aquiline nose and shallow orbits. They were vigorous and fleet-footed, practiced ceremonial burial, had much improved implements in- eluding the bow and arrow and stone lamps, with brains 1,500 to 1,880 ec. They show an appreciation of animals and have been called the Greeks of the old stone age because of their art, which included drawing, engraving, paintings and bas reliefs on cavern walls and floors, and the carving of soapstone, bone and ivory. Their history shows fluctuations in art and industry, in particular their flint workmanship declined with the introduction of bone im- plements. During the climatic fluctuations concerned with the oscillations of the shrinking glaciers and concomitant geographic changes, both their culture and physical vigor show a décline. Their history covers a period of from 10,000 to 15,000 years, and during this time there probably was some intermixture of other blood. Disharmoniec skulls, 7. e., broad face and narrow skull, are still found in the Dordogne and at a few other localities and near by is the primitive agglutinative language of the Basques. Some conclude that these represent late survivals of the Cro-Magnon race. They were followed by fishing races and the first broad headed types, the Maglemose culture (possibly Teutonic) around the Baltic, the Mediterranean (known as the Tardenoisian), and 118 THE SCIENTIFIC MONTHLY the Alpine (Furfooz Grenelle) along the Danube (painted peb- bles). This was from 7,000 to 10,000 years ago, and the so-called Campignian culture of this time is transitional to the Neolithic or New Stone Age of polished stone. The rest of the history belongs more to Anthropology and Archeology. The Robenhausian culture of the Swiss and other lake dwellings about 7000 B. C. shows permanent dwellings, domes- tication of animals and cultivation of crops with use of pottery: the Copper Age extended from 3000 to 2000 B. C.; the Bronze Age in Europe from about 2000 to 1000 B. C., in Orient 4000 to 1800 B. C.; the Iron Age (earlier or Hallstatt culture) in Europe from 1000 to 500 B. C., in the Orient from 1800 to 1000; and the latter — Iron Age from 500 B. C. to Roman times in Europe.' Note the cumulative rapidity of the advance as compared with slowness of change in earlier stages. Although very much remains to be discovered we know enough to assure the layman that man has had a long evolutionary history extending over tens if not hundreds of thousands of years. Does this knowledge breed cynicism and irresponsibility. What answer does science give on this point? Since late Paleolithic time, 7. e., toward the close of the Old Stone age, 25,000 years ago, man’s evolution biologically has been slight and to some extent retro- grade. Skull bones and teeth have changed but little. It was during this period of slight physical change that our race has made the most astonishing progress, and the hope is natural that there is no limit to the betterment of the race by the exercise of wisdom, altruism and idealism—the spiritual graces if you choose so to call them. 1 Figures from Obermaier. SOCIAL LIFE AMONG THE INSECTS Pts SOCIAL LIFE AMONG THE INSECTS’ By Professor WILLIAM MORTON WHEELER BUSSEY INSTITUTION, HARVARD UNIVERSITY LeEctTuRE II. Part 2. Wasps SoLITARY AND SOCIAL Authorities on the classification of the social wasps now divide them into five subfamilies, namely the Stenogastrine, which are confined to the Indomalayan and Australian Regions, the Ropali- diine, confined to the tropics of the Old World, the Polistine, which are cosmopolitan, the Epiponine, possibly comprising two independent lines of descent from Eumenes-like and Odynerus- like ancestors respectively and constituting a large group, mostly confined to tropical America, with a few species in the Ethiopian, Endomalayan, Australian and North American regions, and the Vespine, which are recorded from all the continents except South America and the greater portion of Africa south of the Sahara. These five families may be briefly characterized before considering some of the peculiarities of social organization common to most or all of them. (1). The Stenogastrine evidently represent a group of great interest, because they form a transition from the solitary to the social wasps, but unfortunately our knowledge of their habits is very incomplete. F. X. Williams has recently published observa- tions on four Philippine species, and though his account is frag- mentary, it nevertheless reveals some peculiar conditions. He shows that the single genus of the subfamily, Stenogaster, includes both solitary and social forms and that all of them exhibit a mix- ture of primitive and specialized traits. The species all live in dark, shady forests and make very delicate, fragile nests with particles of decayed wood or earth. S. depressigaster (Figs. 30 E and F) hangs its long, slender, cylindrical nests to a pendent hair-like fungus or fern. The structure consists of tubular, inter- twined galleries and cells, with their openings directed downwards. The colony comprises only a few individuals probably the mother wasp and her recently emerged daughters. The eggs are attached to the bottoms of the cells as in all social wasps and the larve are fed from day to day with a gelatinous paste, which Williams be- lieves may be of vegetable origin. In the cells the older larve and the pupe hang head downwards. Another social species, S. vari- 1 Lowell Lectures. 120 THE SCIENTIFIC MONTHLY pictus, constructs a very different nest, consisting of cells made of sandy mud mixed perhaps with particles of decayed wood and attached side by side in groups to the surfaces of rocks and tree- trunks (Fig. 30 G). In this case also the cell-openings are directed downward. A nest may consist of thirty or more cells in several FIG. 30 Nests of Stenogastrine wasps from the Philippines. A. Stenogaster micans var. luzonensis, female. B. Completed nest of same; C. Nest’ with only the basal portion completed; D. Nest of Stenogaster sp., with umbrella-like “suards”; E. Nest of S. depressigaster; F. diagram of same showing arrange- ment of cells and passage-ways. The numbers indicate the cells. The tops of the passage-ways are shown in two planes by series of parallel lines. G. Nest of S. varipictus on the bark of a tree. (After F. X. Williams). SOCIAL LIFE AMONG THE INSECTS 121 FIG. 31 Suspended and naked comb of a very primitive African Epiponine wasp, Be- lonogaster junceus, with young cells above and old cells containing larve be- low; natural size. Most of the wasps have been removed but two are seen bringing food-pellets to the larvae. (Photograph by E. Roubaud). rows. There are only a few wasps in a colony, and when the larve are full-grown the cells are sealed up by the mother as in the solli- tary wasps. But after the young have emerged the cells may be used again as in many of the social species. Williams describes and figures the nests of two solitary species, one an undetermined form, the other identified as S. micans var. luzonensis. The nest of the former (Fig. 30 D) is suspended, like that of depressigaster, from some thin vegetable fibre and appears to consist of particles of decayed wood. It is a beautiful, elongate structure of seven tubular, ribbed cells, arranged in a zigzag series with their openings below and two peculiar umbrella-like dises around the supporting fibre. These dises ‘‘remind one a good deal of the metal plates fastened to the mooring lines of vessels and serving as rat guards. Their function in the ease of the nest may be an imperfect protec- tion from the ants, or perhaps they may serve as umbrellas, though neither they nor the cells are strictly rain proof.’’ They may pos- sibly be rudiments of the nest envelopes which are so elaborately 122 THE SCIENTIFIC MONTHLY developed in many of the higher social wasps. The mother wasp attends to several young simultaneously, and when their develop- ment is completed seals up the cells. S. micans var luzonensis - (Fig. 30 B) makes the most remarkable nest of all. It is attached to some pendent plant filament under an overhanging bank or under masses of dead leaves supported by twigs or vines and is made of ‘‘moist and well-decayed wood chewed up into a pulp and formed into delicate paper which is not rain proof.’’ The basal portion of the nest (Fig. 30 C) is a single comb of about 20 regu- lar, hexagonal cells, enclosed in a pear-shaped covering which is longitudinally grooved and ribbed on the outside and constricted below to form a filigree-work, funnel-like aperture surrounded on one side by a spear-shaped expansion. This species seems also to have been observed in Ceylon by E. E. Green, who remarks that ‘the nest seems to be the property of one pair only’’ of wasps. Two other species, S. nigrifrons of Burma and melleyr of Java, are also recorded as social. They make nests consisting of a few pendent, hexagonal-celled combs attached to one another by slen- der pedicels. All of the descriptions indicate that the colonies of the social species of Stenogaster must consist of very few indi- viduals, and there is nothing to show that the female offspring FIG. 32 Nest of Polybioides tabida from the Congo, with the involucre partly re- moved. (After J. Bequaert from a photograph by H. O. Lang). SOCIAL LIFE AMONG THE INSECTS 123 differ in any way from their mother or that they assist in caring for the brood. Even in the ease of S. varipictus, Williams remarks: ‘‘In a small way, it seems to be a social wasp; one to several insects FIG. 33 A. Nest of Polybioides melaena of the Congo. B. The same partly destroyed, showing the pendent combs, which have cells on both sides. (After J. Bequaert, from a photograph by H. O. Lang). 124 THE SCIENTIFIC MONTHLY attend to a cell group. It may be, however, that each female has her own lot of cells in this cell group.’’ Future investigations may show that none of the species of Stenogaster is really social — in the same sense as are the four other subfamilies, though they approach the definitively social forms in using paper in the con- struction of the nest, in sometimes making combs of regular hexa- gonal cells and in caring for a number of larve at the same time. (2). The Epiponine are a large and heterogeneous group, comprising a much greater number of genera (23) than any other subfamily of social wasps, and ranging all the way from very primitive forms like Belonogaster to highly specialized forms like Chartergus and Nectarina. Great differences are also apparent in the architecture of the nest, which in the more primitive genera consists of a single naked comb of hexagonal cells attached to some support by a peduncle (Fig. 31), and in the more advanced forms ' of a single comb or of several combs superimposed on one another and enclosed in an envelope with an opening for ingress and egress (Figs. 32 and 33). The combs are in some cases peduneulate (stelocyttarous), in others attached directly to the support or to the envelope (phragmocyttarous). In nearly all cases the nest is made entirely of paper, but in a few tropical American species some clay may be added. It is always above ground and attached to the branches or leaves of trees, to the underside of some shelter (roofs, banks, ete.). In primitive forms like Belonogaster (Fig. 31), as a rule, a single fecundated female starts the nest by build- ing a single pedunculate cell and then gradually adding others m circles concentrically to its periphery as the comb grows, but not infrequently the foundress may be joined by other females before the work has progressed very far. Each larva is fed with pellets of malaxated caterpillars till it is full grown when it spins a con- vex cap over the orifice of its cell and pupates. The emerging females are all like the mother in possessing well-developed ovaries and in being capable of fecundation. In other words, ali the fe- males of the colony are physiologically equal, and even such differ- ences in stature as they may exhibit have no relation to fertility. The colonies are small, the nests having usually only about 50 to 60 cells, rarely as many as 200 to 300. In larger colonies there is a certain rude division of labor since the older females devote themselves to egg-laying, the younger to foraging for food and nest materials and the recently emerged individuals to feeding the larve and caring for the nest. The males, too, remain on the comb, but behave like parasites and exact food whenever it is brought in by the foraging females. Belonogaster is described as a polygynous wasp because each of its colonies contains a number SOCIAL LIFE AMONG THE INSECTS 125 of feeundated females. When it has reached its full development the females leave in small companies and found new nests either singly or together. This phenomenon is known as ‘‘swarming’’ and occurs only in the wasps of the tropics where it seems to be an adaptation to the favorable climatic conditions. In the higher South American genera of Epiponine, however, the females are not all alike but are differentiated into true females, or queens, 1. €., individuals with well-developed ovaries and capable of fe- cundation, and workers, 7. e., females with imperfectly developed ovaries and therefore sterile or capable only of laying unfertilized, male-producing eggs. Many of these wasps, according to H. and R. von Ihering and Ducke, are polygynous and regularly form new colonies and nests by sending off swarms of workers with one or two dozen.queens. The colonies often become extremely popu- lous and comprise hundreds or even thousands of individuals. Some of the species (Nectarina, Polybia) have a habit of storing a considerable amount of honey in their combs, while others are known to capture, kill and store within the nest envelope, and even in the combs, quantities of male and female termites or male ants as a supply of food to be drawn on when needed. (3). The Ropalidiine are a small group of only three genera, the best known of which is Ropalidia. These are primitive wasps which build a single naked comb lke that of Belonogaster and feed their young with pellets of malaxated insects. The colonies are small and polygynous, but, according to Roubaud, true workers ean be distinguished, though they are few in number compared with the true females. Swarming seems to occur in some species. (4). The Polistine are represented by only two genera. One of these, Polistes, is cosmopolitan and, lke Ropalidia and Belono- easter, makes a single, naked comb, suspended by a central or ev- centric pedunele to the underside of some shelter. As there are several common species in Europe and the United States, the habits of the genus are well known. The nest is usually established and in its incipient stages constructed by a single female, or queen. A eertain number of her offspring are workers though they seem often to lay male-producing eggs. True females are rather nu- merous in the colonies of some species, which may therefore be regarded as polygynous, and some of the tropical forms may, per- haps, swarm. In temperate regions, however, the Polistes colony is an annual development and usually not very populous. The young females are fecundated in the late summer and pass the winter hidden away under bark or in the crevices of walls, whence they emerge in the spring to found new colonies. Several of the species, even in temperate regions, are known to store small quan- tities of honey in their combs. 126 THE SCIENTIFIC MONTHLY (5). Like the Polistine, the subfamily Vespine includes only two genera, Vespa and Provespa. The species of the former, the only genus besides Polistes that occurs in the north temperate zone, are the largest and most typical of social wasps. So far as known the species are strictly monogynous. The nest, founded by a single female, consists at first of a small pendent comb, lke that of Polistes, but while there are still only a few cells a more or less spherical envelope is built around it. The eggs first laid produce workers, which are much smaller than the mother and incapable of feeundation. They remain with the parent, enlarge the comb and envelope and, to accommodate the rapidly inereasing brood, build additional combs in a series from above downward, each new comb being supported by one or more peduncles attached to the comb above it (stelocyttarous). At first large numbers of workers are produced, but later in tlie summer males and females appear. Owing to the greater size of the females, the cells in which they are reared are considerably larger than the worker cells. After the mating of the males and females the colony perishes, with the exception of the fecundated females, which hibernate like the fe- males of Polistes and during the following spring found new colo- nies. In the Vespine, therefore, a very distinct worker caste has been developed, though its members occasionally and perhaps regu- larly lay male-producing eggs. The species of Vespa are usually divided into two groups, one with long, the other with very short cheeks. In Europe and North America the long-cheeked forms as a rule build aerial nests above ground, the short-cheeked forms in cavities which they excavate in the ground. The colonies may often be very populous by the end of the summer (3,000 to 5,000 individuals). After this hasty sketch of the five subfamilies of, the social wasps we may consider a few of their fundamental behavioristic peculiarities, especially the trophic relations between the adults and larve, the origin of the worker caste, its ultimate fate in cer- tain parasitic species and the question of monogyny and polygyny. In all these phenomena we are concerned with effects of the food- supply and therefore of the external environment. The feeding of the larve by Vespa and Polistes queens and workers with pellets made of malaxated portions of caterpillars, flies or other insects has often been described and can be readily witnessed in any colony kept in the laboratory. The hungry larve protrude their heads with open mouths from the orifices of the cells, like so many nestling birds, and when very hungry may actually scratch on the walls of the cells to attract the attention of the workers or their nurses. The feeding is not, however, a one-sided SOCIAL LIFE AMONG THE INSECTS 127 affair, since closer observation shows that the wasp larva emits from its mouth drops of sweet saliva which are eagerly imbibed by the nurses. This behavior of the larve has been observed in all four subfamilies of the higher wasps by du Buysson, Janet and Roubaud. Du Buysson says that the larve of Vespa ‘‘secrete from the mouth an abundant liquid. When they are touched the liquid is seen to trickle out. The queen, the workers and the males are very eager for the secretion.. They know how to excite the off- spring in such a way as to make them furnish the beverage.’’ And Janet was able to prove that the secretion is a product of the sali- vary or spinning glands and that it flows from an opening at the base of the lower lip. ‘‘This product,’’ he says, ‘‘is often imbibed by the imagines, especially by the just emerged workers and by the males, which in order to obtain it, gently bite the head of the larva.’’ Most attention has been bestowed on this reciprogal feed- ing by Roubaud, from whose interesting account of Belonogaster, Ropalidia and Polistes I take the following paragraphs: ‘‘All the larve from birth secrete from a projection of the hypopharynx, on the interior surface of the buccal funnel, an abundant salivary liquid, which at the slightest touch spreads over the mouth ina drop. All the adult wasps, males as well as females, are extremely eager for this salivary secretion, the taste of which is slightly sugary. It is easy to observe, especially in Belonogaster, the insistent demand for this larval product and the tactics em- ployed to provoke its secretion. ‘“ As soon as a nurse wasp has distributed her food pellet among the various larve, she advances with rapidly vibrating wings to the opening of each cell containing a larva in order to imbibe the salivary drop that flows abundantly from its mouth. The method employed to elicit the secretion is very easily observed. The wing vibrations of the nurse serve as a signal to the larva, which, in order to receive the food, protrudes its head from the orifice of the cell. This simple movement is often accompanied by an imme- diate flow of saliva. But if the secretion does not appear the wasp seizes the larva’s head in her mandibles, draws it toward her and then suddenly jams it back into the cell, into which she then thrusts her head. These movements, involving as they do a stimulation of the borders of the mouth of the larva, compel it to secrete its salivary liquid. ‘‘One may see the females pass back and forth three or four times in front of a lot of larve to which they have given nutriment, in order to imbibe the secretion. The insistence with which they perform this operation is such that there is a flagrant disproportion between the quantity of nourishment distributed among the larve 128 THE SCIENTIFIC MONTHLY by the females and that of the salivary liquid which they receive in return. There is therefore actual exploitation of the larve by the nurses. ‘‘The salivary secretion may even be demanded from the larva without a compensatory gift of nourishment, both by the females that have just emerged and by the males during their sojourn in the nest. The latter employ the same tactics as the females in compelling the larve to yield their secretion. They demand it es- pecially after they have malaxated an alimentary pellet for them- selves, so that there is then no reciprocal exchange of nutritive material. ‘‘It is easy to provoke the secretion of the larve drtificially. Merely touching the borders of the mouth will bring it about. The forward movement of the larve at the cell entrance, causing them to protrude their mouths to receive the food pellet, is also easily induced by vibrations of the air in the neighborhood of the nest. It is only necessary to whistle loudly or emit shrill sounds near a nest of Belonogaster to see all the larve protrude their heads to the orifice of the cells. Now it is precisely the vibrations of the air created by the rapid agitation of the bodies of the wasps and repeated beating of their wings that call forth these movements, either at the moment when food is brought or for the purpose of obtaining the buccal secretion which is so eagerly solicited.’’ Roubaud has called the interchange of food here described ““oecotrophobiosis,’’ but for reasons which I cannot stop to discuss, I prefer to use the word ‘‘trophallaxis.’’ It will be seen that the larve have acquired a very definite meaning for the adult wasps of ll the castes and that through trophallaxis very close physiolog- ical bonds have been established, which serve to unite all the mem- bers of the colony, just as the nutritive blood stream in our bodies binds all the component cells and tissues together. We found that even in forms like Synagris cornuta the larva has acquired a mean- ing for the mother. In this case Roubaud has shown that the mother while malaxating the food-pellet herself imbibes its juices before feeding it to the larva, and that ‘‘the internal liquids having partly disappeared during the process of malaxation, the prey is no longer, as it was in the beginning, soft and juicy and full of nutriment for the larva. It is possible, in fact, to observe that the caterpillar paté provided by the Synagris cornuta is a coarse paste which has partly lost its liquid constituents. There is no exag- ceration in stating that such food would induce in larve thus nourished an increase of the salivary secretion in order to compen- sate for the absence of the liquid in the prey and facilitate its di- evestion.’’ It is here that the further development to the condition SOCIAL LIFE AMONG THE INSECTS 129 seen in Belonogaster and other social wasps sets in. The mother finds the saliva of the larva agreeable and a trophallactic relation- ship is established. As Roubaud says, ‘‘the nursing instinct having evolved in the manner here described in the Humenids, the wasps acquire contact with the buccal secretion of the larva, become ac- quainted with it and seek to provoke it. Thence naturally follows a tendency to increase the number of larve to be reared simulta- neously in order at the same time to satisfy the urgency of ovi- position and to profit by the greater abundance of the secretion of the larve.’’ As I shall endeavor to show in my account of the ants and termites, trophallaxis is of very general significance in the social life of insects. It seems also to have an important bearing on the development of the worker caste. Both queens and workers arise from fertilized eggs, and the differences between them are com- monly attributed to the different amounts of food they are given as larve. There seems to be much to support this view in the social wasps. As Roubaud points out in the passages quoted, the larve are actually exploited by the adult wasps to the extent of being compelled to furnish them with considerable quantities of salivary secretion, often out of all proportion to the amount of solid food which they receive in return. Owing to this expenditure of substance and the number of larve which are reared simulta- neously, especially during the earlier stages of colony formation, they are inadequately nourished and have to pupate as rather small individuals, with poorly developed ovaries. Such individ- uals therefore become workers. This inhibition of ovarial devel- opment, which has been called ‘‘alimentary castration,’’ is main- tained during the adult life of most workers by the exigencies of the nursing instincts. The workers have to complete and care for the nest, forage for food and distribute most of it among their larval sisters. All this exhausting labor on slender rations tends to keep them sterile. In other words, ‘‘nutricial castration’’ (de- rived from nutriz, a nurse, to use Marchal’s terms, takes the place in the adult worker of the alimentary castration to which it was subjected during its larval period. It is only later in the develop- ment of the colony, when the number of workers and consequently also the amount of food brought in have considerably increased, and the labor of foraging and nest construction have correspond- ingly decreased for the individual worker, that the larve can be more copiously fed and develop as fertile females, or queens. At that season, too, some of the workers may develop their ovaries, but as the members of the worker caste are incapable of fecunda- tion, they can lay only male-producing eggs. That this is not the VOL. XV—9 130 THE SCIENTIFIC MONTHLY whole explanation of the worker caste will appear when we come to consider the much more extreme conditions in the ants and ter- mites, but it may suffice to explain the conditions in the social wasps and social bees. Parasitism is another phenomenon which seems to indicate that a meager or insufficient diet is responsible for the development of the worker caste. Although parasitic species are much more nu- merous among the bees and ants, I will stop to consider very briefly a few of those known to occur among the wasps. ) , cat can ae | fat Dan hat ran Tat fan Ga (ie ke) sat man that than ok: ee Words on Chart at Left. vine of oa violet love —f NW visit give voice lived et ) very lives ACS have over . hive clover five seven average. They were taught, in the East Stroudsburg State Normal Training School, by two cadet teachers and myself, no one of us having ever before taught a child to read. There was some diffi- culty also in procuring the netessary type and other materials for keeping the experiment going.’ At the end of a month (spending a little over an hour a day on the subject), twenty-three sounds had been introduced, and the pupils were attacking new words with fair success. A week later, the brighter pupils were separated from the rest and began 1In reporting this experiment, I wish to make acknowledgment of the receipt of financial aid by means of which it was promoted from the Amer- ican Association for the Advancement of Science. Acknowledgment of substantial assistance of a different kind is due to Mrs. La Rue, without whose help the necessary reading material could not have been composed, illustrated and printed. VOL. XV.—18. 274 THE SCIENTIFIC MONTHLY reading such stories as ‘‘The Little Red Hen’’ without the aid of the teacher. At the close of eleven weeks, our advanced class had learned all the symbols, had read about one hundred fifty pages of the Fonoline Primer, and could readily master, independently, any new word of not more than five or six symbols (that is, five or six sounds when spoken), unless it involved some pecuhar diffi- culty. As few words in the first grade vocabulary reach this length, Mie CAT AND THE MOUSE = | | i. hn) A wee mouse was eating. Cc, map) c*) el. A cat saw her. ii ae baa The cat said, © “T must have that mouse.” —~-)| (| =p) Then away she went. Fic. 2. Showing fonoline used interlineally to aid in the introduction to a-b-e¢ English. The words that have no fonoline beneath them had already been mastered by. the pupils before reaching this story. THE SHORTHAND ALPHABET 275 we thought it best to pass from this grade of attainment to the study of a-b-c English. At the end of fifteen weeks, the slower section also (containing, it will be remembered, some retarded pupils) having covered all their symbols and read over one hun- | dred pages of the Primer, proceeded to the study of a-b-c English. Passing from fonoline to ordinary English introduced prac- tically no new problems except those which are always incident to the teaching of reading in English, and we of course used our “‘nerfect’’ phonetic alphabet to aid in the mastery of the imper- fect, partially unphonetic one. The first means employed was that of interlinear printing, placing the a-b-c English above and the corresponding fonoline just below as a key to pronunciation, as shown in the figure. As soon as a word had appeared in the a-b-c type a few times, it was left without the fonoline aid to pronunciation beneath it, whereupon the pupil either remembered it or was forced to go back and find it where it had last appeared. At the close of the year, our pupils had accomplished, so far as we were able to judge, substantially the.same amount of work in a-b-e English, after spending the first ten or fifteen weeks on fonoline reading, as they would have done had they spent the whole year on a-b-c English; that is, their achievements were on a level with those of preceding classes, the time devoted to reading re- maining unchanged. Our advanced elass won the special com- mendation of the State examiner, who had no knowledge of how the grade had been taught. We are inclined to believe that fonoline forms’a good introduc- tion to a-b-e English, and that if it could replace the usual system of diaeritical marking, time would ultimately be gained through its use. We consider it quite safe to assert that if a pupil of average intelligence and application were given a year of instruction in reading fonoline (especially if there were devoted to reading the two hours per day commonly assigned to it in our city schools), such a pupil would then be able to read anything (printed in that alphabet) which he was capable of understanding. Beyond re- views, no further work in reading would be necessary for one so taught except to train him in the apt expression of those thoughts and feelings which would come to him with maturity. And he would not only know how to read: he would be able to find in the fonoline dictionary any ordinary word that he could pronounce. Further, he could ‘‘spell,’’ both orally and in writing (fonolne characters) any word that he could turn his tongue to. Let us now give our attention to the educational and social advantages that would be ours if such an alphabet as fonoline were brought into common use. Let us keep in mind, too, that 276 THE SCIENTIFIC MONTHLY fonoline is advantageous beyond any other phonetic alphabet; for it bears a unique relation to Pitmanic shorthand, the most speedy and efficient means yet devised by the human brain for passing its thoughts down through hand and pen and so recording them on paper. First, then, does fonoline present an alphabet which adequately represents the sounds of spoken English? We can sum up this matter admirably by quotations from Max Muller: ‘‘What I like in Mr. Pitman’s system of spelling is exactly what I know has been found fault with by others, namely, that he does not at- tempt to refine too much, and to express in writing those endless shades of pronunciation, which may be of the greatest interest to the student of acoustics, or of phonetics, as applied to the study of living dialects, but which, for practical as well as for scientific philological purposes, must be entirely ignored . . . . Out of the large number of sounds, for instance, which have been eata- logued from the various English dialects, those only can be recog- nized as constituent elements of the language which in and by their difference from each other convey a difference of meaning. Of such pregnant and thought-conveying vowels, English possesses no more than twelve. Whatever the minor shades of vowel sounds in Enelish dialects may be, they do not enrich the language, as such; that is, they do not enable the speaker to convey more minute shades of thought than the twelve typical single vowels If I have spoken strongly in support of Mr. Pitman’s system, it is : chiefly because it has been tested so largely and has stood the test well.’” Next, if the number of our characters is correct, is their form satisfactory? As to the advantages of simplicity, perhaps the work of Broea and Sulzer can be accepted as authoritative. These in- vestigators concluded that both our letters and the words of which they are composed would be more easily recognized and quickly read if they were simplified in form. ‘‘Practically,’’ they report, ‘‘the recognition of a letter demands an expenditure of energy that is' greater as its form is more complex. Thus we read a V, a T, orvsan Li more easily than an E or a B. From the standpoint of speed of reading and also of the cerebral fatigue caused by the act it would be better to employ simpler letters than those now used. .We have thus been led to seek the least complex possible forms, and we have concluded that, for capital letters, they are those shown in Fieure 3. For the small letters, where there 2 From an article in the Fortnightly Review of April, 1876, as quoted in The Life of Sir Isaac Pitman, by Alfred Baker, p. 206. THE SHORTHAND ALPHABET 206 O'S Ges O I -HnAS ets ie oe eee -~JFE< 10-16 The hydrogen atom consists of one proton and one electron. The unit of mass here employed is a mass 1/1846th part less than that of the hydrogen atom itself. The precise value of this mass in grams does not here interest us any more than does the precise value of the unit electric charge expressed in terms of the con- ventional electrostatic units. Every atom consists of a positively charged nucleus in which is concentrated nearly all the mass of the atom, and this nucleus is surrounded by a number of electrons sufficient to neutralize the nuclear charge. The atom as a whole is, of course, electrically neutral. For hydrogen, the positive nucleus consists of but one proton; but, in all other atoms, the positive nucleus contains what are called intranuclear electrons as well as protons. The value of the net positive charge on the nucleus of any particular type of atom is equal to the atomic number of the atom, that is, the ordinal number the atom would receive if all our 92 elements were numbered in order of increasing atomic weight, or, better, because of the three standard disorderly cases of potassium-argon, nickel- eobalt and iodine-tellurium, in order of rank in the periodic tabu- lation of the elements. The roll of the chemical elements, as we shall see, was called first, if incompletely, by one Moseley in 1914, and, of the 92 now on the roll, five elements are yet absent. The number of extranuclear or planetary electrons in chemical atoms, therefore, runs from 1 for hydrogen, 2 for helium, 3 for lithium, ete., up to 92 for uranium, the heaviest atom not yet extinct. The number of planetary electrons is the same as the atomic number of the atom. The outside diameter of atoms is of the order of 1 to 510-5 em., or 100,000 times the diameter of the electron, so that it is evident that the spacing of the nucleus and extranuclear electrons in an atom is a very open one, more so than in our solar system, in which, also, our central sun is inordinately large for true relative dimen- sions. So far as the volumes of the discrete structural units, pro- 366 THE SCIENTIFIC MONTHLY tons and electrons, are concerned, therefore, atoms are filled chiefly with emptiness or void. Granting that the planetary electrons are so few in number, never over 92, and so light in weight, it is obvious that, as has been said, the main mass of the atom must be concentrated in its nucleus. To illustrate the make-up of an atomic nucleus, consider, for example, the case of the atom of fluorine. Its atomic weight is about 19. Therefore its nucleus must contain 19 protons, which would furnish a positive charge of +19. But its atomic number is 9, and hence its nuclear charge is +9. To reduce a positive charge of 19 to 9 will require the presence of 10 electrons in the nucleus, ten negative units of electrostatic charge thus offsetting ten of the positive units. As regards the picture of the atom developed so far, there is much unanimity. The kind of questions not yet solved are those about the detailed constitution of the nucleus, and the arrangement of the extranuclear electrons. Further study might therefore be divided into (1) study of the nucleus and its constitution, and (2) study of the extranuclear portion of the atom and its possible arrangements of electrons, with the orbits or oscillations in which they are engaged. But we shall not follow this program. Having placed before you a broad idea of the present-day conception of the structure of atoms, I wish now to go back and look at some of the methods of study that have led to the knowl- edge we have gained. ELECTRONS AND PROTONS Our knowledge of the electron has been reached chiefly through a study of the discharge of electricity through gases, with which the name of Sir J. J.-Thomson is so closely associated. We should not forget, however, that, as early as the seventies of last century, Sir William Crookes observed streamers of light emerging from the cathode of highly evacuated discharge tubes. These he con- sidered were composed of matter in a new, fourth state which he called ‘‘radiant matter.’’ Certain German investigators thought the streamers were due rather to an ether wave motion analogous to light. It was not till 1895 that Sir J. J. Thomson showed that Crookes had been correct. The cathode rays are now known to consist of streams of swiftly moving electrons. Like any other particles that carry a charge, these electrons are deflected from a straight line course by either a magnetic or an electrostatic field properly applied, and, by using known field strengths of both kinds and measuring the resulting deflections of the electrons, the re- lation e/m between their charge and their mass ean be ascertained. This is a method of measurement of very fundamental importance. Very high speed electrons of different, definite speeds are ejected MODERN STUDY OF THE ATOM 367 ‘ by certain ‘‘radioactive’’ atoms at the moment when such atoms break up, and a study of e/m for these electrons has shown that, if the charge is the same, the apparent mass increases with the measured speed of the electrons in a manner that harmonizes pre- cisely with what would be anticipated on theoretical grounds if their mass were entirely what is called electromagnetic, or entirely due to their charge. Knowing that the mass of an electron is entirely electromagnetic in origin, we can then, assuming a spherical form, calculate by electromagnetic rules its radius, which comes out with the value noted above. Individual electrons have been isolated by Millikan and others, and their charge evaluated precisely in electrostatic units. No electric charge that was a fractional part of that of an electron was ever encountered in this work, but only whole number multi- ples of it. From this it appears that negative electricity comes in small unit amounts or grains, like pepper. In other words, nega- tive electricity, like matter, is atomic. At present we know less about the proton, or unit positive charge, than about the electron; its much greater mass, if electromagnetic, gives it a correspond- ingly smaller radius, as noted above, because, for the same charge on a spherical surface, electromagnetic mass varies inversely as the radius of the sphere. ORIGIN OF X-RAYS The electrons in a discharge tube, such as an X-ray bulb, can be made to move with very high speeds, and so to carry much energy. They are stopped abruptly by striking the atoms that constitute the target in the bulb, and, by transferring some of their own energy, stimulate radiation of short wave light on the part of certain of the extranuclear electrons belonging to these atoms. They likewise themselves emit general wave radiation as they slow down. The very existence of these so-called X-rays, emerging from such targets, was not observed until 1896, and they were not known to consist merely of short wave length light until 1912. X-Rays AND CRYSTAL STRUCTURE I may remind you that one of the most successful methods of studying and analyzing common light is by means of a Rowland diffraction grating, which consists of a large number of fine lines ruled exceedingly closely together on a transparent or else re- flecting surface. Such a grating will perform its analytical fune- tion only provided the ruling or grating space is sufficiently close compared with the wave length of the light being tested. A picket fence is a possible Rowland grating, but is far too coarse to give 368 THE SCIENTIFIC MONTHLY useful results. So likewise a real Rowland grating is far too coarse to give appreciable diffraction of X-rays, because these are of such short wave length. In 1912 it occurred to the Swiss Laue that in ordinary crystalline substances we have ready-made very minutely spaced diffraction gratings, of skeletal type, indeed, and in three instead of only two dimensions of space. Stated other- wise, he thought the atoms of which the crystal is built might fur- nish centers to diffract the X-ray light, and these atoms are already arranged by Nature in rank, file and column with perfect regularity and with the requisite close spacing. Laue’s expecta- tions were brilliantly fulfilled, and the method has been developed by the Braggs in England, by Dr. Hull at Schenectady and by many others. Because it is based on a relationship between the spacing of the layers of atoms and the wave length of the light diffracted by them, this powerful modern method of analysis may be em- ployed in either of two ways, namely, to study (a) either the light wave length or (b) the spacing of the atoms. A single relationship connects these two unknowns. To get a value to start from, the atomic spacing in, say, sodium chloride can be computed from the known number and plausible crystallographic arrangement of atoms in one cubic centimeter of crystalline salt. Hence we can find the wave length of some particular easily-generated mono- chromatic X-ray light, which we can thenceforth use conveniently as our yard-stick in measuring atomic spacings in other crystals. RESULTS FROM THE X-RAY SPECTROMETER fa). Moseley in 1914 tried many different elements as targets in the X-ray bulb, analyzed the resulting X-rays by means of a erystal, and found that each element emitted general X-radiation, but also its own characteristic sets of X-rays of but a few kinds, that is, each element emitted a characteristic X-ray spectrum, consisting of but a few spectral lines peculiar to the element. Se- lecting a particular type of emitted light (K, L, ete., series) for making comparisons amongst the elements, Moseley arrived at the remarkable result that the square roots of the wave frequencies varied in progressive stepwise fashion from element to element, giving a series of equal whole-number steps that would, if com- plete, run approximately from 1 for hydrogen to 92 for uranium, just as the atomic numbers do. Any missing step was conspicu- ous, and indicated clearly an element lacking. From this we have learned that we must search for just five new elements, and that we must adjust our atomic numbers to allow for the elements that are missing. MODERN STUDY OF THE ATOM 369 It may here be remarked that the ordinary old style light spectra emitted by the chemical elements in the flame, are, spark, sun or stars are of much more complex character than these sim- pler X-ray spectra. The latter give us information in regard to those extranuclear electrons which, after suffering displacement, gain positions of stability nearest to the nucleus; while the former longer wave light spectra inform us of the relatively slower radia- tions of those electrons that reach stations of stability in the outer regions of the atom. Within the last few years, both types of spectra have been so extended as to overlap, and the former out- standing gap of four octaves of light frequency has been com- pletely bridged. (b). The distances apart of layers of atoms in erystals are of an order smaller than one millionth of an inch. Small as are these distances, however, they are measurable by the X-ray spectrometer with an accuracy much better than one hundredth of one per cent. Given the crystallographic data of a crystal and its atomic arrangement, one may, by assuming the atomic weights of the constituent atoms, determine the density of the crystal by X-ray analysis with an accuracy superior to that of the more usual older methods. Conversely, if the density of the erystal is known, but the atomic weight of one of the constituent atoms is unknown, this may similarly be determined. Of greater interest, however, is the employment of the X-ray spectrometer to determine the arrangement of the atoms in a erystal. This can be done with a good deal of assurance for crystals belonging to the erystallo- graphic systems of higher symmetry. Simple substances, such as the pure metals, usually crystallize in such systems, and have been subjects of fruitful study. Especially interesting to the organic chemist is the erystal lattice of diamond, in which each carbon atom is surrounded by four others at equal distances from it and in the directions of the four corners of a tetrahedron from its center of gravity. It has heretofore been tacitly assumed that the nuclei of the atoms in the crystal acted as simple point centers for diffracting light; but closer study reveals evidence of the posi- tion of certain extranuclear electrons which are in all probability those that constitute the valence bonds of the carbon. It is further apparent that these valence electrons, if not stationary, at least patrol stations other than the nucleus. This is confirmatory of the Lewis-Langmuir theory of atomic structure, to be mentioned shortly. Of even greater interest to the organic chemist are the recent studies by means of the X-ray spectrometer of the possible arrangement of the atoms in naphthalene, anthracene and allied substances. Viewed from one standpoint, the carbon atoms in Vol. XV.—24. 370 THE SCIENTIFIC MONTHLY diamond le at the corners of hexagons bent at their corners so as to fit on puckered surfaees, or surfaces composed of V-like corru- gations lke galvanized roofing without rounded bends. In spite of its different crystalline form, very different hardness, ete., graphite, the other crystalline form of carbon, exhibits an identical arrangement of carbon atoms in puckered hexagonal formation, but with the planes containing the hexagons further apart. As- sumption of this same corrugated hexagonal structure, which ap- pears so favorable for carbon atoms, satisfies the experimental re- sults obtained with crystals of naphthalene in the X-ray spec- trometer. One can not say that the organic chemist’s customary graphic formula for naphthalene and its allies has been inde- pendently proyed correct; but it has at least been shown that this time-honored formula is in complete harmony with the new ex- perimental facts. ARRANGEMENT OF EXTRANUCLEAR ELECTRONS IN ATOMS Passing from the arrangement of atoms in molecules to the arrangement of the extranuclear or planetary electrons in indi- vidual atoms, we reach a field where the experimental facts, chem- ical or spectroscopic, are more difficult of present interpretation, and where, at present, we are largely being guided and, I should add, stimulated by hypothesis. The hypothesis at present most acceptable to the chemist, doubtless because it was designed to fit the chemical rather than the spectroscopic facts, is that of Lewis and Langmuir; and this may be most easily visualized by the aid of models of atoms. Here we have the electrons patrolling definite- ly localized stations which are arranged in concentric shells round the nucleus. Similarity in number and arrangement of stations in the outermost shell makes for similarity of chemical properties, and so accounts for the well-known family resemblances of a chemical kind among the atoms. Furthermore, for many of the commoner atoms at least, the number of electrons in the outermost shell that gives the greatest stability is, on this ‘‘octet’’ theory, eight. If we personified such atoms, their chief ideal in life would be to secure, by hook or crook, an outside shell of precisely eight electrons; and this ideal motivates all the chemical reactions of such atoms. The two possible mechanisms by which the octet ideal is achieved are by reciprocal lending and borrowing or else by sharing electrons, corresponding to the conceptions of electro- valence and covalence, respectively. Atoms that already have eight electrons in their outer shells, like neon and argon, have no motive for any chemical action, and, as is well known, are entirely inert. By these new conceptions of valence, certain molecular structures are anticipated to be closely similar (isosteric) which . MODERN STUDY OF THE ATOM 371 would be entirely dissimilar according to the older ideas of valence ; and the close similarity of crystal form (isomorphism) actually observed in many such cases, a stumbling block and even a reproach to the chemical crystallographers of but ten years ago, strongly confirms the correctness of the newer views in these instances. ’ Less interested in the chemistry of the atoms, Bohr and his followers have been more concerned in devising an atomic model that will explain the kinds of energy radiated by an atom as light when one of its extranuclear electrons falls from a location of higher to one of lower potential energy, this light having a fre- queney which depends not on the final environment reached by the electron but rather on the energy made available by the fall. This theory is brilliantly successful, but hitherto only in the two sim- plest cases. If the planetary electrons all revolve in simple ellipses round the nucleus, there is insufficient localization of fields of force around the atom to satisfy the valence demands of the chemist, even although the orbits be not in the same plane. To me, the possibility of twisted or looped orbits round the nucleus would offer more satisfaction ; as also would a complex electron to account for the facts of radiation. . Because both these types of hypothesis as to the arrangement and activities of the extranuclear electrons are at present in a somewhat speculative stage, I prefer to pass on to tell of other matters closer to the facts reached by the modern methods of study. SPONTANEOUS DISINTEGRATION OF ATOMS: ISOTOPES All of our atoms of atomie weight over 206 are observed to have the proclivity to disintegrate. The nucleus of the atom of Ura- nium-1l, for example, has a mass about 238 and a charge of +92. Oceasionally, such an atomic nucleus, for cause utterly unknown, suffers a cataclysm in which the nucleus of a helium atom is ex- pelled with enormous speed. (A helium atom consists of a minute nucleus built of 4 protons bound together by 2 electrons, which is surrounded by two planetary electrons). This expelled portion, endowed with terrific energy of motion, is called an alpha-particle. Because it lacks the two planetary electrons of the helium atom, it bears a double positive charge, and the moving alpha-particle can therefore be studied in the magnetic and electrostatic fields and the ratio measured of its charge to its mass, by which measure- ment its nature was divulged. On picking up two _ planetary electrons the alpha-particle becomes a helium atom. The precise volume has been measured of helium gas generated in this wise from radioactive material ejecting alpha-particles in numbers that can be counted one by one, and so has been enumerated directly 372 THE SCIENTIFIC MONTHLY the number of helium atoms per cubic centimeter of the helium gas collected. After throwing out its alpha-particle, the residual portion of the Uranium-1 atom, having lost 4 protons, possesses a mass 234 instead of 238. It has lost from its nucleus also 2 electrons, and therefore a net charge of (4—2) or +2. Its atomie number is thus smaller by 2, that is 90, and its ordinal position in the periodic table is two places below and to the left of Uranium-l. It is, in fact, a new element, named Uranium-X,, which resembles the element Thorium. But it is a short-lived element, and soon expels from its nucleus an electron, or beta-particle, yielding a residual product, called Uranium-X,, of the same atomic weight 234, but of atomic number 91. This disrupts with loss of another beta- particle even sooner than the last, producing the atom of an element, called Uranium-2, of atomic weight still 234, but of atomic number 92. But 92 is the atomic number of Uranium-l from which we started. Thus Uranium-1 and Uranium-2 have the same nuclear charge, +92, and must have the same arrangement of extranuclear electrons, for this is ordered by the nuclear charge, and not appreciably by the nuclear mass. Thus, the outsides of U-1 and U-2 will be identical, and on the outside of an atom do its chemical properties depend. Consequently, U-1 and U-2, falling in the same ordinal position in the periodic system, are identical chemically and therefore inseparable by chemical means. Such elements are called isotopes. They have the same nuclear charge. Elements with the same nuclear mass are called isobars, as U-X,, U-X.,, and U-2. After a chain of successive disintegrations involving losses of 8 and 6 alpha-particles (mass 4) respectively, both uranium (mass 238) and thorium (mass 232) produce atoms with the nuclear charge of lead. On account of their different parentage, however, these two types of lead atoms would be expected to have masses of 206 and 208 respectively. They have each been isolated, from uranium and thorium minerals respectively, and found to possess atomic masses closely as expected. Most of the known radioactive or spontaneously disintegrating elements have atomic weights of 206 or over. But rubidium and potassium are also radioactive, as are one of the constituents of common brass and also the metal platinum, although these latter atomic species disintegrate at an exceedingly slow speed and emit alpha-particles so lacking in energy that they are difficult to de- tect. There is no reason why the habit of disintegration should not be general among elements. In any case, we are clear that isotopic atoms, whether formed in a process of disintegration or in MODERN ‘STUDY OF THE ATOM 373 a process of evolution, should be identical chemically, and so, like birds of a feather, should be found together. The question then arises, how many of our elements are mixtures of isotopes, indis- tinguishable by chemical difference? The question can be answered only by physical methods. If the isotopes are heterobaric, of dif- ferent mass, then they can be identified and separated because of their difference in mass. Speed of diffusion or of free evaporation depends on mass, and so the elements mereury and chlorine have both been shown to consist of mixtures of heterobaric isotopes. THE Mass SPECTROGRAPH But the most fertile method yet employed to recognize hetero- baric isotopes is that devised by Sir J. J. Thomson, depending on the fundamental fact, already mentioned above, that different values of the ratio charge to mass of a charged particle moving in a high vacuum will give rise to different degrees of deflection in the magnetic and electrostatic fields. A gas or vapor particle con- taining the element under investigation is therefore given a posi- tive charge in the discharge tube, and its resulting deflection is studied. Such a particle may acquire more than one unit charge, but such multiple charges cause no confusion. Refinement of this beautiful method in the hands chiefly of Aston has shown us that the elements Li, B, Ne, Mg, Si, Cl, A, K, Ca, Ni, Zn, Br, Kr, Rb, Sn, Xe, Hg, are all, at least as we have them ordinarily available on this planet, mixtures of isotopes; whereas H, He, Be, C, N, O, F, Na, P, As, I, and Cs have been studied and proved simple. Pub- lished results on no others are yet available. Only those elements which can readily be obtained in stable gaseous form have yet been investigated; but as soon as any of the less volatile elements or their compounds are obtained in suitable gas form, additional results of interest will be forthcoming. Because his measurements in the mass spectrograph of the masses of most atoms had an ac- curacy of 0.1 per cent., Aston was enabled to discover that the atomic weights of the chemical elements investigated, save hydro- gen, are invariably whole numbers on the scale oxygen=—16, to the degree of accuracy mentioned. The fractional value 35.46 found by chemical analysis for chlorine, for example, is explained by that element’s consisting of certain proportions of two isotopes of masses 35.0 and 37.0 respectively. Thus is the century old hypothesis of Prout, that all atoms have masses that are whole number multiples of the mass of the hydrogen atom, resurrected and rehabilitated, but with a unit of mass almost 0.8 per cent. less than that of the hydrogen atom. Mathematical rigor in this ‘“whole number rule’’ is, however, not to be expected for a reason that will now be referred to. 374 THE SCIENTIFIC MONTHLY A NEw PossIBLE SouRcE OF CosMIcAL ENERGY Hydrogen, in Aston’s mass spectrograph, appears to have an atomic mass of 1.008 (0O=16), precisely as found by the chemists. Hydrogen, however, is unique among the elements in having no electrons but merely one proton in its nucleus. In all other atomic nuclei there are electrons packed close to the protons, and this close packing of charges of opposite sign is expected, by electrical theory, to influence the electromagnetic mass of the complex by an amount dependent on the closeness of packing. If we could build a helium atom from the materials which are correctly furnished by four hydrogen atoms, there would ensue a loss of mass of about 0.8 per cent., since the atomic weight of helium is 4.00. This mass, however, can not be destroyed, but must appear, according to the relativity theory, as an equivalent amount of energy. The quan- tity of energy concerned in the transformation of 1 gram of hydro- gen to helium, namely 8 milligrams, or about the weight of one fifth of a postage stamp, corresponds to what, as electrical energy. at ten cents per kilowatt hour, would cost $20,000. ~ If such a synthesis of helium from hydrogen may be supposed to be going on in the sun, we have a much needed explanation of the sun’s present brightness at his known old age. EXPERIMENTAL UTILIZATIONS OF ALPHA-PARTICLES IN INVESTIGATION The alpha-particles expelled from radioactive elements are by far the most energetic entities with which we are yet acquainted. The swifter kinds have, for unit mass, an energy of motion 400 million times greater than that of a rifle bullet. Being, helium nuclei without cireumambient planetary electrons, they are very small, and readily shoot through atoms, knocking out their extra- nuclear electrons right and left. The damaged atoms soon pick up other electrons to fill the gaps, and thus suffer no permanent change. Alpha-particles pass through thin glass, leaving no hole. Very occasionally, an alpha-particle will make a bullseye collision with the nucleus of an atom. The consequence of this collision, in the case of nuclei of some light atoms, is that the nucleus struck is disintegrated, with the expulsion of a single, swift-moving pro- ton. The experiment succeeds in the case of B, N, F, Na, Al, and P nuclei. In the ease of aluminium, Rutherford found that the single protons expelled have an energy of motion that is 40 per cent. greater than that of the alpha-particle missile that struck the atom. What remains of the aluminium nucleus is still under investigation, but it is permanently changed and is certainly no longer aluminium. Thus, in transmuting an element, we obtain free energy. The alchemists expressed this allegorically when they identified the philosopher’s stone, which would transmute metals, MODERN STUDY OF THE ATOM 375 with the elixir of life, a source of ever fresh life or energy. Our enthusiasm for this transmutation is duly restrained by the know]l- edge that only about two per million alpha-particles make bullseyes on the aluminium nuclei. In the case of the heavier atoms with their more highly charged nuclei, the alpha-particle, of the speed hitherto at our disposal, apparently loses too much energy in the approach to be able to effect disintegration of the nucleus. Its path is sometimes bent back in a large angle deflection, by an elaborate study of which in the case of gold the existence of the minute, positively charged nucleus of atoms was first established. The nuclear charges of gold, platinum, silver and copper have each been directly evaluated by this method, and agree with the atomic numbers of these atoms to 1 per cent., the known value of the experimental error. THE DURATION AND PossIBLE REPETITIONS OF GEOLOGICAL TIME The unchanging and, so far as all experiment goes, unchange- able spontaneous disintegration of heavy atomic nuclei has geo- logical interests both in view of the time periods involved and also of the energy emitted in the process. Thorium disintegrates into thorium-lead, which is stable, at a rate which we have been able to ascertain. The ‘‘range’’ or distance travelled through matter by the expelled alpha-particles is strictly related to the rate of the various disintegrations, and such ranges and rates have ever re- mained constant for thorium and its descendants as evidenced by the constancy of the diameters of the range ‘‘halos’’ surrounding microscopic thorium inclusions in rocks of various ages. The diameters of these halos agree, also, with the ranges in air of the alpha-particles as studied to-day, and so the thorium clock has ever run at the same rate. Some thorium minerals contain lead whose measured atomic weight shows that it has practically all been derived from the decay of thorium atoms. From the ratio of the number of thorium atoms remaining to the number of those originally present, many of which are now represented by lead atoms which they produced, one can compute the age of the min- eral, which thus gives a date to early paleozoie times of 150 million years back. Similar calculation from uranium minerals gives a period over 900 million years, but there is reason to believe that the uranium clock formerly ran fast, or, rather, that it appeared to run fast owing to the former presence of a third isotope of uranium, now almost extinct, of speedier rate of decay than what to-day we call U-1. Rock analysis shows that, assuming percentage composition” similar to that on the surface, there is enough radioactive material 376 THE SCIENTIFIC MONTHLY in a depth of only 12 miles of the earth’s crust to supply by its daily disintegration all the heat the earth radiates daily into space. If an appreciable amount of radioactive material exists below this depth, as seems certain, then heat must slowly be accumulating within the earth’s non-conducting crust. Eventually, therefore, if no compensating heat-absorbing process is taking place within, an unstable state will be reached when the underlying incandescent material will perforce evert itself to the exterior and there dis- burden itself of its accumulated heat by radiation into space at a very rapid rate proportional to the fourth power of the tempera- ture. This is the earth’s incandescent epoch. When the crust has cooled down again sufficiently, a new geological epoch of per- haps 200 million years may begin, to be followed in turn by an- other incandescent epoch, and so on, alternately, but more and more slowly, until the radioactive materials, if not regenerated, have by disintegration lost their available energy. This alternation the Brahmans have symbolized in their cosmogony as the indraw- ing and outbreathing of the breath of Brahma. THE PROGRESS OF SCIENCE 377 THE PROGRESS OF SCIENCE CURRENT COMMENT By Dr. Epwin E. SLtosson Science Service EVOLUTION WORKING BACKWARD ONcE farmers planted the nubbins of their corn and the potatoes that were too small to sell. Now they know better. They cut up their finest potatoes to plant, and every grain of their seed corn is pedigreed as care- fully as a Colonial Dame. The result is seen in the doubled yield in pota- toes richer in starch and corn richer in protein. Modern agriculture is fertilized by science. The most backward branch of biol- ogy is the infant science of sociology. It is only just beginning to get its eyes open, to see things; in time, per- haps it will be able to do things, like the older sciences. But there is need of haste. The age of instinct is pass- ing, the reign of reason has not come. Man has been pushed up to his _pres- ent position. He has succeeded in slackening the pressure. Will he go forward rationally, of his own free will, or sink back until again he falls under the sway of the blind and merciless forces of the struggle for existence ? A decrease in the birth rate is not necessarily a misfortune to a country. Very likely, for instance, the British Isles have now all the population they can support in comfort under present economic conditions. The alarming thing about it is that the breeding is from the poorest stock instead of the best. Whatever objective standard one may take this is true. A statis- tical study of the population of Great Britain showed that in the dis- tricts where there was the most over- crowding, the cheapest type of labor, the lowest degree of culture and edu- cation, the highest percentage of pauperism and lunacy, the greatest criminality and the highest death rate from tuberculosis and infantile diseases, there the number of chil- dren was greatest in proportion to the possibly productive wives. It is a clear case of the survival of the un- fittest, the reversal of evolution. No race can maintain its efficiency and virility against sueh reactive forces. The future of a country depends ultimately upon the character and ability of its people. Increase of wealth, advance of science, improve- ment in education, discoveries in .san- itation, juster social conditions, all the achievements and hopes of the present age will be of little benefit to posterity if there is a decline in the native quality of the race. It would be disastrous to hand over a more perfect and complicated govern- mental machine to inferior engineers. ' One seventh of the present generation will be the parents of one half of the next. Therefore, two generations of selection, natural or designed, would completely transform the character of a nation. Is this seventh composed of the best men and women that we have? This is what is going to determine whether civilization shall advance or retrograde. Galton’s ideal of euge- nics may be too much in advanée of the age to be practical, but at least something could be done to awaken the people to the imminent dangers of dysgenies. ATOMS OF LIGHT THE discovery of the X-rays in 1895 acted like the discovery of gold in an unexplored country. It opened the way to the exploration of a field of unsuspected wealth of new knowl- edge and to the radical reconstrue- 3 78 THE SCIENTIFIC .MONTHLY Wide World Photos PROFESSOR A. L. HERRARA The distinguished biologist of Mexico, who is visiting the biological institu- tions of the United States. THE PROGRESS OF SCIENCE tion of some of our time-honored and fundamental conceptions. up to us the atom, the ne plus ultra of the chemist, and showed within it a system of revolving bodies far more numerous and complicated than the solar system. Already our knowledge of these electrons, whose existence was unsuspected a few years ago, is greater than our knowledge of the molecules, and we can study them with much more facility be- cause they carry charges of electricity which betray their presence in the minutest number. A single electron can be detected while the smallest number of gas molecules which can be discerned with the spectroscope is about ten million million. The tendency of the times is to ex- tend the atomic theory into new tields, to speak of atoms of electricity, of energy and of light. The corpuscle, the smallest known particle of nega- tive electricity, is only one seventeen- hundredth the mass of the atom of hydrogen. The smallest unit of posi- tive electricity, on the other hand, seems to be equal to the atom of hy- drogen. It is possible, however, that this positive particle may be a com- plex of many positive and negative particles and that the individual pos- itive corpuscle when isolated as the negative one has been may prove to be equally minute. The discovery of the stores of energy compact in the atom in the form of the electrostatic po- tential energy of its negative cor- puscles gives one a peculiar sensation. It is like finding out that there is a barrel of gold and a dynamite bomb in the cellar of the house. But a gram of hydrogen would be capable of developing more heat than the burning of thirty-five tons of coal. Since energy is wealth we have every- where enough to make us all rich ““Heyond the dreams of avarice’? for- ever, but we have no way of unlock- ing this storehouse. This may be for- tunate for us since Professor J. J. enormous It opened. 379 Thomson, of Cambridge, says, ‘‘if at any time an appreciable fraction were to get free the earth would ex- plode and become a gaseous nebula.’’ Professor Thomson, in compensation for our natural disappointment at being frightened off these preserves by such a terrifying spring-gun, re- minds us that on every sunny acre 7,000 horse-power of radiant energy from our solar dynamo is going to waste and that it is neither impossi- ble nor dangerous to utilize it. THE LITTLE ENEMIES OF MAN EARLY in the history of the human race man learned how to conquer the mastodon. He has yet to learn how to master the microbe. Whales and elephants are now almost extinet, but mice and flies still increase and mul- tiply, and the bacteria, smallest and most dangerous of all, find new ways of attacking us. It is only within the last few years. that has learned which his greatest enemies are, and he has not yet found weapons against them. The explorer in tropical jungles used to fear the lions, tigers and pythons;' now he man protects himself most carefully against the mosquitoes and _ tsetse. Mars has afflicted the human race less than Beelzebub. Although we theoreticaliy accept the conclusion of science that a man’s foes are those of his own household, we are not yet aroused to the neces- sity of waging war in earnest against them. We have a secretary of navy and we give him millions for defense, but we have no secretary of sanita- tion, though that is a more necessary office. It is quite improbable that any American will be killed by an invading army this year, but our land is invaded by millions of mosquitoes and flies armed with deadly weapons and certain to slaughter thousands. Years of study and experimentation before we learn but will be necessary how to fight our already enough has been done to show insect foes, 380 THE SCIENTIFIC. MONTHLY Photograph by Harris and Ewing DR. GEORGE P. MERRILL Head curator of geology in the United States National Museum, to whom the National Academy of Sciences has awarded the J. Lawrence Smith Medal for his investigations of meteorites. THE PROGRESS OF SCIENCE what can be accomplished if we go about it in the right way. Many of the sanitary measures of the past we now know to be crude, clumsy and misdirected, yet they are fixed in the popular mind and remain on our statute books. People still talk about the dangers of miasma and sewer gas, and think a deodorizer is a dis- infectant. We are far from acting up to our lights. The housewife wages war against vermin, but she does not realize that they are more dangerous than trolley cars. She gets more ex- cited at the discovery of a moth than a fly, although the former only attacks clothing, not its contents. We have drain pipes in our walls to carry off disease, but beside them are con- veniently arranged passages by which | roaches can carry diseases from flat to flat, so that everybody has a fair chance to cateh whatever is going. Our windows are hospitably open to the malarial mosquito and typhoid- bearing fly. Over our clothing on the street cars crawl unmentionable insects carrying unmentionable dis- eases. In the fashionable hotel and restaurant the napery and porcelain are immaculate and the waiters are scrupulous; what goes on behind the sereen and in the market is another story. “We have got past the days when we kept the pig in the parlor, but we still keep the dog in the parlor, which is quite as bad. On the street we see the pet dog gnawing a decaying bone and nosing the foulest spot to be found, and a moment later he is cuddled in the arms of his fair | and fastidious mistress and licking her cheek. We have yet to realize that it is the dogs which are not mad that are the more dangerous. They injure more people by their kisses than their bites. In primitive days man had to asso- ciate with the lower animals. He needed dogs and horses and he very properly made friends of them. He is now learning how to’ do without 381 them, and he should, like a snob who has risen in the world, exclude them from his circle of intimates. The house is not intended for a zoological garden. Insects and animals may be our worst enemies. THE BIRTHPLACE OF THE EELS THE final chapter in the life story of the eel has been written by the Danish expedition under Dr. Joh. Schmidt, which has recently returned to Copenhagen. The breeding grounds have been found between the Bermudas and the Leeward Islands, Where the sea reaches a depth of more than a mile. The origin and mode of reproduc- tion of the common eel have been for centuries a matter of speculation. It has long been observed that large eels migrate toward the sea in autumn and that in the spring little elvers are found under stones on the seashore and ascend the streams in vast num- bers. A group of small transparent salt water fishes, known as Lepto- cephali, were described in 1763, but no one guessed that they were in any way related to the eels. In 1864, Theo. N. Gill, of the Smithsonian Institution, published the conclusion that these Lepto- cephali are the young or larve of the 2els, and this was confirmed through direct observation by Yves Delage in 1886. Beginning the following year, Professor Grassi made careful studies of the development of the eel in Sicily, observing the transformation of Leptocephali into the conger and other genera of eels, and in 1894 the larva of the common eel was dis- covered. It was evident that the spawning of mature eels occurred in the sea, and now the place has been discov- ered by Dr. Schmidt. The European species deposit their eggs to the south. and east of the Bermudas, while the-American species breeds to the south and west of the islands. Wide World Photos The ship Dana, returning to Copenhagen with the Danish deep sea expedition which found the breeding place of the eel near the Bermudas. ' Wide World Photos The Dana at Elsinore, where it was boarded by Prince Valdemar of Denmark and Prince George of Greece, both of whom are seen in the door of the cabin, while the leader of the expedition, Dr. Johs. Schmidt, director of the Carlsberg Laboratory, is seen at the extreme left. THE PROGRESS The former make a three-year migra- tion to the shores of Europe from the North Sea to Italy, while the latter journey to the American coast from New. England to the south in a few months or a year. The Leptocephali after their trans- formation into elvers ascend the and sometimes travel over- or up streams land from stream the faces of dams and along the sides of rocks in search of sufficient water. The eels lve for fresh waters, the period being from five to In the eels to stream years in as many as twenty or thirty. autumn some of the mature travel back to the sea, the males then being from twelve to eighteen inches in length, the females never less than eighteen. At the original breeding places they. spawn and-die. THE TOTAL SOLAR ECLIPSE OF: SEPTEMBER 21 By IsaseLt M. LEwIs Science Service Some of the points at eclipse expeditions were located on September 21 are the Maldive Isl- ands in the Indian Ocean, Christmas Island about 250 miles south of the west end of Java, Wallal on the western Goast of Australia, Cordillo Downs in‘central Australia, to which instruments and supplies were trans- ported by camel trains from Ade- which OF SCIENCE 383 laide, South Australia and Goondi- windi in the southern part of Queens- land. The longest duration of total- ity was five minutes and seconds at Wallal. The Kodiakanal Observatory expe- dition from South India in charge of Director Evershed was in the Maldive Islands. On Island the eclipse was awaited by expeditions the Royal Observatory of Greenwich and the combined expedi- tion Holland which were joined by observers from Java. The British has been on the island since the last of nineteen Christmas from and from Germany expedition March making extensive preparations for testing the Einstein theory of relativity. It is essential for this purpose to photograph the field of stars in which the sun will be found at the time of eclipse several months tt as the Einstein theory requires, the rays of light from stars near the sun are deflected from their course at the time of eclipse owing to the attrac- before or after the eclipse date. ‘ tion of the sun’s mass, a comparison of photographs taken when the sun is in this field of stars at eclipse with photographs taken months previous when the sun was not in the field will show ithe displacement of several the star images required by the theory. A number of eclipse expeditions From Nature. Shadow track during total solar eclipse of September 21, 1922. 384 were located at Wallal, West Aus- tralia, owing to the generosity of the Australian government in placing at the disposal of the eclipse expeditions a transport of the Australian navy. Some of the expeditions that ac- cepted this offer of the Australian government are the Crocker eclipse expedition of the Lick Observatory, California, in charge of Professor W. W. Campbell; an expedition from the University of Toronto which in- cluded Dr. R. K. Young, of the Do- minion Astrophysical Observatory, Victoria, B. C., and an expedition from the Observatory of Perth, West Australia. The transport left Free- mantle, the port of Perth, the last of August and will bring members of the expeditions back to that port after the eclipse. The chief object of several of the expeditions was to test the EHinstein theory which requires that stars near the sun that are visible when the sun’s rays are temporarily blotted out shall be displaced from their nor- mal positions by amounts depending upon their angular distances from the rim of the sun. It will be re- called that the deflections both in direction and amount theory were obtained by the British observers at Principe, Africa, and Sobral, Brazil, at the time of the total solar eclipse of May, 1919. This is the first opportunity that has been afforded since that date to obtain an additional test of this prediction of the relativity theory. required by SCIENTIFIC ITEMS WE record with regret the death of William S. Halsted, professor of sur- gery at the Johns Hopkins Medical School; of Rollin D. Salisbury, pro- fessor of geographical geology at the University of Chicago; of Dr. Harold C. Ernst, professor of bacteriology in the Harvard Medical School; of Ste- phen Smith, distinguished for his contributions to public health, who THE SCIENTIFIC MONTHLY had nearly reached his hundredth birthday; of Arthur Ransome, the English authority on public health, who died at the age of ninety-two years; of W. H. Hudson, the English ornithologist and writer on natural history, and of Edward M. Eidheer, formerly expert in the Austrian bureau of chemistry. THE British Association for the Advancement of Science held its ninetieth annual meeting at Hull from September 6 to 13 under the presidency of Sir Charles Sherring- ton, professor of physiology at Ox- ford and president of the Royal So- ciety. Professor Mangin, director of — the Paris Museum of Natural His- tory, presided over the meeting of the French Association for the Advance- ment of Science held at Montpellier from July 24 to 29.—The Association of German Scientific Men and Physi- cians held its hundredth meeting at Leipzig from September 18 to 24. One of the public addresses was by Professor Albert Einstein. A pRIZE of $25,000 to be awarded annually to a chemist of the United States for contributions to chemistry was announced by the Allied Chem- ical and Dye Corporation of New York, at the recent Pittsburgh meet- ing of the American Chemical So- ciety. THe French Senate has unanimous- ly voted 2,000,000 franes to observe the hundredth anniversary of the birth of Louis Pasteur, which will take place this year. The Senate in voting the appropriation described Pasteur as the ‘‘symbol of French science. ’’ Tur late Prince of Monaco has bequeathed sums of one _ million francs each to the Academy of Sciences, the Academy of Medicine, the Institut Océanographique, the Institut de Paléontologie Humaine of Paris, and the Musée Océanographi- que of Monaco. THE SCIENTIFIC MONTHLY NOV EMBER. 1922 SOCIAL LIFE AMONG THE INSECTS’ By Professor WILLIAM MORTON WHEELER BUSSEY INSTITUTION, HARVARD UNIVERSITY Lecture IV—Ants, THEIR DEVELOPMENT, CAsTES, NESTING AND FEEDING Hasits N one occasion several years ago when I was about to lecture on ants in Brooklyn, a gentleman introduced me to the audi- ence by quoting the sixth to eighth verses of the sixth chapter of Proverbs, and then proceeded in utter seriousness to give an inti- mate account of their author. He said that Solomon was the greatest biologist the Hebrews had produced, that he had several large and completely equipped laboratories in which he busied himself throughout his reign with intricate researches on ant behavior and that the 700 wives and 300 concubines mentioned in the Bible were really devoted graduate students, who collaborated with the king in his myrmecologieal investigations. The gentleman deplored the fact that the thousand and one monographs embody- ing their researches had been lost, and concluded by saying that he was delighted to introduce one who could supply the missing information. As he had consumed just forty-three minutes with his account of Solomon and his collaboratrices, I had to confess my inability to ‘‘deliver the goods’’ in the remaining seventeen. From what recondite sources of biblical exegesis the Brooklyn gentleman drew his information I have never been able to ascertain, but I am sure that Solomon’s few myrmecological comments, which have come down to us from about 970 B. C., are very accurate—far more accurate than that story of Herodotus, written some 500 years later, of the gold-digging ants of India, which were as large as leopards, and whose hides were seen by Nearchus in the camp of Alexander the Great, and whose horns were mentioned by Pliny as hanging, even in his time, in the temple of Hercules at Erythre. 1 Lowell Lectures. Vol. XV.—25. , 386 THE SCIENTIFIC MONTHLY This and the many other ant stories invented or disseminated by ancient and modern writers are certainly not devoid of interest, but the actual behavior of the insect is so much more fascinating that you will pardon me for not dwelling on them. The Formicide constitute the culminating group of the stinging Hymenoptera and have attracted many investigators for more than a century and especially during the past thirty years. Unlike the honeybee these insects make no appeal to our appetites nor even to that vague affection which we feel for most of the common denizens of our forests, fields and gardens, but only to our in- quisitiveness and anxiety. Hence the vast literature which has been written on the ants may be said to have been prompted by scientific, philosophic or mere idle curiosity or by our instinct of self-preservation. In the presence of the ant we experience most vividly those peculiar feelings which are aroused also by many other insects, feelings of perplexity and apprehension, which Maeterlinck has endeavored to express in the following words: ‘‘The insect does not belong to our world. Other animals and even the plants, despite their mute lives and the great secrets they enfold, seem not to be such total strangers, for we still feel in them, not- withstanding all their peculiarities, a certain terrestrial fraternity. They may astonish or even amaze us at times, but they do not completely upset our calculations. Something in the insects, how- ever, seems to be alien to the habits, morals and psychology of our globe, as if it had come from some other planet, more monstrous, more energetic, more insensate, more atrocious, more infernal than our own. With whatever authority, with whatever fecundity, un- equalled here below, the insect seizes on life, we fail to accustom ourselves to the thought that it is an expression of that Nature whose privileged offspring we claim to be. . . . No doubt, in this astonishment and failure to comprehend, we are beset with an indefinable, profound and instinctive uneasiness, inspired by be- ings so incomparably better armed and endowed than ourselves, con- centrations of energy and activity in which we divine our most mys- terious foes, the rivals of our last. hours and perhaps our suc- CeSSOrs. he The similarities which the ants, as one of several families of aculeate or stinging Hymenoptera, necessarily bear to the wasps and bees, are so overlaid by elaborate specialization and idiosyn- erasies that their primitive vespine characters are not very easily - detected. I wish to dwell on some of these specializations, but be- fore doing so, it will be advisable to give under separate captions a brief summary of what I conceive to be the fundamental pe- culiarities of the ants: SOCIAL LIFE AMONG THE INSECTS 387 (1) The whole family Formicide consists of social insects, that is, it includes no solitary nor subsocial forms such as we found among the beetles, wasps and bees. We are therefore unable to point to any existing insects that might represent stages leading up to the social life of the ants. Within the family, nevertheless, we can distinguish quite a number of stages in a gradual evolution of social conditions from very simple, primitive forms, whose colonies consist of only a few dozen individuals, with a compara- tively feeble caste development, to highly specialized forms with huge colonies, comprising hundreds of thousands of individuals and an elaborate differentiation of castes. (2) The number of described species of ants is approximately 3,500, but if we include their subspecies and varieties, many of which will probably be raised to specific rank by future, less con- servative generations of entomologists, we shall have more than double the number. This is far in excess of the number of all other social insects, including both the groups I have already con- sidered and the termites. The ants are therefore the dominant social insects. (3) This dominance is shown also by their geographical dis- tribution, which is world-wide. There are ants everywhere on the land-masses of the globe, except in high aretic and antarctic lati- tudes and on the summits of the higher mountains. The number of individual ants is probably greater than that of all other insects. With few exceptions, the termites are all confined to tropical or subtropical countries, and the number of social wasps and bees in temperate regions is very small. (4) We found that the social wasps arose from the Eumenine solitary wasps and the bees from the solitary Sphecoids. All the authorities agree that the ants had their origin in neither of these ancestral stocks, but among the Scolioids, a distinct offshoot of the primitive Vespoids. Of the four modern families of the Scolioids, the Psammocharide, Thynnide, Mutillide and Scoliide, the last seems to be most closely related to the ants. Since they must be traced to ancestors which were winged in both sexes, the Thynnids and Mutillids, which have wingless females, are excluded, and the family Psammocharide is not very closely allied to the Formicide. (5) The ants, unlike the social wasps and bees, are eminently terrestrial insects. They inherited and seem very early to have exaggerated the terrestrial habits of their primitive Scolioid ances- tors. The majority of the species in all parts of the world still nest in the soil. Many of them later took to nesting in dead or de- eaying’ wood, and more recently a number of species, especially in the rain-forests of the tropics, have become arboreal and nest by preference in the twigs of trees and bushes or construct paper or 388 THE SCIENTIFIC MONTHLY silken nests among the leaves and branches. The terrestrial habit led to a permanent phylogenetic suppression of the wings in the workers, an ontogenetic loss of the wings in the queens and a diminution of the eyes in both of these castes. A few very archaic ants still possess large eyes like the wasps and bees, but in the great majority of species, which are more or less subterranean, and therefore practically cave-animals during much of their lives, the eyes have dwindled, and in many species have almost or com- pletely disappeared. The great abundance of ants in the desert, savanna and prairie regions of the globe indicates that they arose during some period of the Mesozoic, perhaps during the Triassi¢ or Liassic, when the climate was warm but arid. Their extensive adaptation to low, damp jungles, with their rank vegetation, seems to have developed during the Cretaceous or early Tertiary. The ants therefore resemble the solitary wasps, which are still con- spicuously abundant in hot, arid regions. Both groups are repre- sented by only a small number of species in cool, moist regions, like New Zealand, the British Isles and certain mountain ranges, like the Selkirks of British America. (6) In the social wasps and bees we found that the worker, | FIG. 54 Stigmatomma pallipes, a primitive, subterranean Pcnerine ant of the United States. The winged individuals are virgin queens and are very similar to the workers. Nearly twice natural size. (Photograph by J. G. Hubbard and OWS strong) SOCIAL LIFE AMONG THE INSECTS 389 or sterile caste, though distinctly differentiated, is, nevertheless, very much like the queen, or fertile female. In ants the differ- ences are much greater. Even when, as in many primitive ants (Fig. 54), the worker resembles the queen in size and form, it never possesses wings, and in most ants the two castes are so dis- similar that they have often been described as separate species. The male ant, too, is much less like the queen than is the corre- sponding sex among the social wasps and bees (Fig. 57). It is evident, therefore, that all three castes are more, highly specialized. In many ants, as we shall see, the worker, queen and male may each become differentiated into two or more castes, a phenomenon which is nowhere even suggested among the wasps and bees. (7) Very long and intimate contact with the soil has made the ants singularly plastic in their nesting habits. While most social wasps and bees construct elaborate combs with very regular, hexa- gonal cells of such expensive substances as paper and wax, the ants merely make more or less irregular galleries or chambers in the soil or dead wood or if they construct paper or silken nests avoid a rigid type of architecture. Hence the great variability of nest- ing habit in the same species. This plasticity and saving of time and labor are very advantageous, because they enable the insects, when conditions of temperature or moisture become unfavorable or when bothersome enemies settle too near the nest, to change their habitation readily and without serious loss to the colony. Espinas long ago noticed the importance of the terrestrial habits of ants. He says: ‘‘Ants owe their superiority to their terrestrial life. This assertion may seem paradoxical, but consider the excep- tional advantages afforded by a terrestrial compared with an aérial medium in the development of their intellectual faculties! In the air there are the long flights without obstacles, the vertigim- - ous journeys far from real bodies, the instability, the wandering about, the endless forgetfulness of things and of oneself. On the earth, on the contrary, there is not a movement that is not a con- tact and does not yield precise information, not a journey that fails to leave some reminiscence; and as these journeys are deter- minate, it is inevitable that a portion of the ground incessantly traversed should be registered, together with its resources and its dangers, in the animal’s imagination. Thus there results a closer and much more direct communication with the external world. To employ matter, moreover, is easier for a terrestrial than an aérial animal. When it is necessary to build, the latter must, like the bee, either secrete the substance of its nest or seek it at a distance, as does the bee when she collects propolis, or the wasp when she gathers material for her paper. The terrestrial animal has its building materials close at hand, and its architecture may be as 390 THE SCIENTIFIC MONTHLY varied as these materials. Ants, therefore, probably owe their social and industrial superiority to their habitat.’ (8) The plasticity of ants is shown even more clearly in their care of their young, which are not reared in separate cells but in clusters and lie freely in the chambers and galleries of the nest where they can be moved about and easily carried away or hidden when the colony is disturbed or the moisture and temperature conditions are unfavorable. Like their continual contact with their physical environment, their intimate acquaintance with their young in all their stages has been an important factor in the high psychological development of the Formicide. (9) A similar plasticity characterizes their feeding habits. As a group they feed on an extraordinary range of substances: the bodies and secretions of other insects, seeds, delicate fungi, nectar, the saccharine excreta of plant-le, scale insects, ete. Some species seem to be almost omnivorous. (10) All this adaptability, or plasticity in nesting and feeding habits is, of course, an expression of a very active and enterprising: disposition and has resulted in the formation of a vast and intri- cate series of relationships between ants and other organisms, in- cluding man. These restless, indefatigable, inquisitive busybodies, forever patrolling the soil and the vegetation in search of food, poke their noses, so to speak, into the private affairs of every living thing in their environment. Nor do they stop at this; they actually draw many organisms, by domesticating them or at any rate at- taching them to their nests or bodies, into the vortex: of their cease- © less, impudent activities. Nearly every week during the past twenty years I have received from some entomologist somewhere on our planet one or more vials of ants with a request for their . identification, often because they had been found associated with some insect or plant which the sender happened to be investi- gating. In the next lecture-I shall describe a number of . the strange partnerships into which ants have entered as a result of their inordinate and unappeasable appetites. As my time is limited I shall select for discussion only a few of the topics suggested in the foregoing summary, namely, the main taxonomie divisions of the family Formicide, polymorphism, or the development of castes, the origin and growth of colonies, the structure of the alimentary canal in adult and larval ants and the evolution of the feeding habits. In their main outlines, at least, the phylogenetic relationships of the various subdivisions or subfamilies of the Formicide have been clearly established. There are seven of them: the Ponerine, Cerapachyine, Doryline, Pseudomyrmine, Myrmicine, Dolicho- SOCIAL LIFE AMONG THE INSECTS 391 FORMICINAE MY PMICINAE DOLICHODERINAE PSEUDOMYRMIHAE PONERINAE CERAPACHYINAE DORYLINAE SCOLIOID ANCESTORS. FIG. 55 Ancestral tree showing the putative phylogenetic relations of the family Formicide as a whole and of its subfamilies to one another. derine and Formicine. The Ponerine constitute the primitive, basic stock of the family and have given rise to the six other sub- families, which are represented in the ancestral tree (Fig. 55) as so many branches. Their thickness roughly indicates their vigor or comparative development and their height their degree of spe- cialization and dominance in the existing fauna. All the sub- famihes are well represented in the tropies of both hemispheres, but in the north temperate region nearly all the species belong to the two largest and highest subfamilies, the Myrmicine and Formicine. In temperate North America and Eurasia there are very few Dolichoderine and Ponerine and no Cerapachyine nor Pseudomyrminez. A small number of Doryline extend as far north as Colorado, Missouri and North Carolina (35° to 40°) and to about the same latitude on the southern shores of the Mediterranean. With the exception of a series of peculiar parasitic genera, which are represented only by males and females, all ants possess a sharply defined worker caste. In primitive groups, like the 392 THE SCIENTIFIC MONTHLY Ponerine, Cerapachyine and Pseudomyrmine, the worker is nearly as large as the queen but lacks the wings and has therefore a more simply constructed thorax, the compound eyes are smaller and the simple eyes or ocelli are minute or absent. In the three sub- families mentioned the worker is monomorphie, that is, it always has the same form though it may vary somewhat in size. In the four remaining subfamilies (Doryline, Myrmicine, Dolichoderine and Formicine) we find the same uniformity of the worker in many species, but in a considerable number it has become highly variable, or polymorphic, as a result of agencies which have acted independently in each subfamily or even within the limits of a single genus (Figs. 57 and 58). In such eases the workers can be arranged in-a graduated series, beginning with large, huge- headed individuals more like the queen in stature, and ending with minute, small-headed individuals, which may be very much smaller than the queen. Such a series exhibits not only great morpholog- FIG. 57 Ns A small Myrmicine harvesting ant of Texas, Pheidole instabilis, with poly- morphic worker caste. a, soldier; f, worker; b to e, forms intermediate be- tween the soldier and worker (lacking in most other species of the huge genus Pheidole) ; g, queen (dedlated) ; h, male. The figures are all drawn to the same scale. SOCIAL LIFE AMONG THE INSECTS 393 FIG. 58 Portion of a colony of a common Formicine ant (Camponotus americanus), comprising virgin, winged queens and workers, the latter showing the un- stable polymorphism in stature and size of head, characteristic of most species of the genus. (Photograph by J. G. Hubbard and O. S. Strong.) ical but also great functional differences among its members. The largest individuals commonly act as policemen or defenders of the colony, but in some species their powerful jaws enable them to erush seeds or the hard parts of inseets, so that the softer parts may be exposed and eaten by the smaller individuals (Fig. 57). The latter excavate the nest, forage for food, nurse the young and in some species devote all their energies to the cultivation of the fungus gardens. In a graduated series like the one described we usually call the largest workers ‘‘maxime,’’ the smallest ‘‘minime’’ and the intermediate forms ‘‘medie,’’ the word ‘‘operariz’’ (workers) being understood in each ease. Now in some ants only the two extremes, the maxime and the minima, of the polymorphic series proved to be serviceable to the colony, so that all the intermediate forms (medizw) have been eliminated, leaving the worker caste distinctly dimorphic. In such ants we eall the maxime ‘‘soldiers’’ (milites) and the minime ‘‘workers’’ (operari#). This condition has been attained in several genera and subgenera among the Myrmicine and Formicine (Pheidole (Fig. 57), Oligomyrmex, Colobopsis, ete.). In still-other genera, ‘ 394 THE SCIENTIFIC MONTHLY where soldiers were not needed or were too expensive to rear and maintain, on account of their great size and appetites, they too have been eliminated and the worker caste is represented only by the tiniest individuals of the originally polymorphie series (Care- bara, Tranopelta, Pedalgus, Solenopsis, ete.). There is therefore an enormous difference in these ants in size and structure between the queen and the only surviving worker form of the species. In Carebara, e. g., the queen is several thousand times as large as the worker! Nevertheless, both are merely extreme female forms of the same species and may, of course, develop from the eggs of the same mother. But the worker is not the only caste that has become dimorphie. In some species there are two distinet forms of queen, in others two distinct forms of male. In these cases one of the forms is winged, the other usually apterous. And here again, by suppres- sion of the winged female or winged male, the wingless form may become the only surviving fertile form of its sex in the species. All these developments are interesting because they indicate that the distinctions among the various eastes have arisen gradually by eontinuous or fluetuating variations and that the survival and persistence of some of them and the elimination of others have led to the sharply discontinuous series of castes which we find in many ants. : It is obvious that some of the differences between the various castes, especially those in size, are due to differences in the amount of food consumed during the larval stages, but the profounder morphological differences which separate the queens, soldiers and workers, must be due to other causes. We must suppose either that the food administered to the larve differs in quality or that there are several different kinds of eggs, some of which develop into fertile, other into sterile forms. In a sense the latter would be mutations, like the various sterile forms of the evening primrose, which make their appearance generation after generation from some of the seeds of the fertile forms. In the ease of the ants, however, we find that the workers not infrequently lay viable eggs, and though they are never fertilized’ and generally develop into males, the latter may mate with queens and thus be a means of establishing a representation of the characters of their worker mothers in the germ-plasm of the species. The peculiar anomalies known as gynandromorphs, that is, individuals partly male and partly. female, which occasionally oceur among ants, also indicate that the queens, soldiers and workers arise from as many different kinds of eggs, since there are three different kinds of gynandro- morphs, exhibiting respectively combinations or mosaics of male and queen, male and soldier and male and worker characters. It SOCIAL LIFE AMONG THE INSECTS 395 » is difficult to see how such perfectly definite combinations could be produced by larval feeding, and it is equally difficult to account for them as the results of internal secretions. In the present state of our knowledge we can only surmise that the differences between the queen and worker castes were originally ontogenetic and de- termined by feeding, as they still are in the social wasps and bees, but that in the ants the germ-plasm has somehow been reached and modified, so that an hereditary basis for caste differentiation has been established. The ant colony may be initiated and developed by one of two different methods which I shall call the independent and the de- pendent. The former is peculiar to the nonparasitic; the latter to the parasitic species. Leaving an account of the ants which employ the dependent method for the next lecture, I would say that the great majority of ants establish their colonies in essentially the same manner as Vespa and the bumble-bees. The winged, virgin queen, after fecundation during her nuptial flight, descends to the ground, rids herself of her wings and seeks out some small cavity under a stone or piece of bark, or excavates a small cell in the soil. She then closes the opening of the cell and remains a voluntary prisoner for weeks or even months while the eggs are growing in her ovaries. The loss of the wings has a peculiar effect on the voluminous wing-museles in her thorax, causing them to break down and dissolve in the blood plasma. Their substance is earried by the circulation to the ovaries and utilized in building up the yolk of the eggs. As soon as the eggs mature, they are laid and the queen nurses the hatching larve and feeds them with her saliva till they*pupate. Since she never leaves the cell during all this time and has access to no food, except the fat she stored in her abdomen during her larval life and her dissolved wing-muscles, the workers that emerge from the pupe are all abnormally small. They are, in fact, always minime in species which have a poly- morphic worker caste. They dig their way out through the soil, thus establishing a communication between the cell and the outside world, collect food for themselves and their mother and thus enable her to lay more eggs. They take charge of the second brood of egos and larve, which, being more abundantly fed, develop into larger workers. The population of the colony now increases rapidly, new chambers and galleries are added to the nest and the queen devotes herself to digesting the food received from the workers and to laying more eggs. In the course of a few years numerous males and queens are reared and on some meteorologic- ally favorable day the fertile forms from all the nests of the same species over a wide expanse of country escape simultaneously into the air and celebrate their marriage flight. This flight provides 396 THE SCIENTIFIC MONTHLY not only for the mating of the sexes but also for the dissemina- tion of the species, since the daughter queens, on descending to the ground, usually establish their nests at some distance from the parental colony. It will be seen that the queen ant, like the queen wasp and bumble-bee, but unlike the queen honeybee, is the perfect female of her species, possessing not only great fecundity but in addition all the worker propensities, as shown by her ability to make a nest and bring up her young. But as soon as the first brood of workers appears, these propensities are no longer manifested. That they are not lost is shown by the simple experiment of removing the queen’s first brood of workers. Then, provided she be fed or have a sufficient store of food in her body, she will at once proceed to bring up another brood in the same manner as the first, although she would have manifested no such behavior under normal con- ditions. As already stated, this independent method of colony forma- tion is the most universal and is followed alike by tropical and extratropical ants. It is undoubtedly the primitive method and, as we shall see, the one from which the dependent method has been derived. It differs from that of Vespa and Bombus, nevertheless, in leading to the formation of perennial colonies even in temperate and boreal regions. The queen ant may, in fact, live from 12 to 17 years and although, like other aculeates, she is fecundated only once, may produce offspring up to the time of her death. Unhke the queen honeybee she is never hostile to her own queen daughters, and in many species of ants some of these daughters may return after their marriage flight to the maternal colony ‘and take a very active part in increasing its population. In this manner the colony may become polygynic or pleometrotic, and in some instances may contain a large number of fertile queens. When such a colony grows too large it may separate into several, the queens emigrating singly or in small companies, each accompanied by a detachment of workers, to form a new nest near the parental formicary. This behavior is exhibited by the well-known mound-building ant (Formica exsectoides) of our New England hills. You will notice that its mounds usually occur in loose groups or ¢lusters and that the workers of the different nests are on friendly terms with one another and sometimes visit back and forth. We may, of course, eall the whole cluster a single (polydomous) colony, but it really differs from a number of colonies only in the absence of hostility between the inhabitants of the different mounds. In certain tropical ants, like the Doryline (Figs. 59 and 60), however, I am inclined to believe that the only method of colony formation is by a splitting of the original colony into as many parts as it con- SOCIAL LIFE AMONG THE INSECTS 397 FIG. 59 Argentinian legionary (Doryline) ant Eciton (Acamatus) strobeli. Workers showing polymorphism and male, photographed to the same scale as the four smaller workers. (Photograph by Dr. Carlos Bruch.) FIG. 60 Dorsal arid lateral view of the wingless queen (dichthadiigyne) of Eciton (Acamatus) strobeli. Same scale as Fig. 59. (Photcgraph by Dr. Carlos Bruch.) 398 THE SCIENTIFIC MONTHLY tains young queens. These huge, clumsy creatures (Fig. 60) are always wingless and must therefore be fecundated in the nest, and since the colonies, which comprise hundreds of thousands of work- ers, are nomadic and keep wandering from place to place, they must become independent entities as soon as they are formed. We possess no accurate data on the age that ant colonies may attain. Some of them certainly persist for 30 or 40 years and probably even longer. In such old colonies the original queen has, of course, been replaced by successive generations of queens, that is, by her fertile daughters, grand-daughters and great-grand- daughters, and the worker personnel has been replaced at a more rapid rate, because the individual worker does not live more, and in most instances lives considerably less, than three or four years. The feeding habits of ants are so varied and complicated that it will be advisable before considering them to describe the struc- ture of the alimentary canal in both adult and larva. The mouth- parts of the adult are of the generalized vespine type and consist FIG. 61 Sagittal sections through the heads of ants. A, of queen Lasius niger with the mouth open (After Janet). B, of queen Camponotus brutus with the mouth closed. #, tongue; 0, oral orifice; ph, pharynx; h, infrabuccal pocket; pe, pellet in situ, made up of solid particles of food refuse and strigil sweep- ings. Note stratification in the substance of the pellet, indicating successive meals or toilet operations. » SOCIAL LIFE AMONG THE INSECTS 399 of a small, flap-like upper lip, or labrum, a pair of strong, usually toothed mandibles, a pair of small maxille and a broad. lower lip, or labium. The maxille and labium are each provided with a pair of joimted, sensory appendages, the palpi. The mandibles, which are really the ant’s hands, vary greatly in shape in different genera and are used not only in securing the food but also in many other activities, such as digging in the earth or wood, transporting other . ants or the young, fighting, leaping, etc. Liquids are, of course, merely imbibed and swallowed, but solid food is seized and crushed with the mandibles and the juices or smaller particles licked up with the tongue, which is a roughened pad at the tip of the lower lip (Fig. 61¢) just anterior to the opening of the duct of the salivary glands. The small particles thus collected are carried back into a small chamber or sae, the infrabuceal pocket (Fig. 61h), which lies immediately below and anteriorly to the mouth-open- ing (0). This pocket is an important structure since it serves as a receptacle not only for the more solid particles of food but also for the dirt, fungus-spores, ete., which the ant. collects during her toilet operations, for the ant is an exquisitely cleanly insect and ‘devotes much of her leisure to licking and burnishing her own smooth or finely chiseled armor and that of her nest-mates. More- over, the tip of the fore tibia is furnished with a beautiful comb or strigil which can be opposed to another comb on the concave — inner surface of the fore metatarsus. The ant cleans her legs and antenne by drawing them between these combs, which are then drawn across the mouth, with the result that any adhering dirt is carried off into the infrabuccal pocket. In this manner the dirt and the solid or semisolid food particles are combined and the whole mass moulded in the infrabuceal pocket into the form of a roundish oblong pellet (Fig. 61B pe). After any liquid which it may contain has been dissolved out and sucked back into the mouth, the pellet is cast out, so that no solid food actually enters the alimentary canal. All adult ants therefore subsist entirely on liquids. The alimentary canal proper is a long tube extending through the body and divided into sections, each with its special function. The more anterior sections are the mouth eavity, the pharynx (Fig. 61 ph), which receives the duets of certain glands, and the very long, slender gullet, which traverses the posterior part of the head, the whole thorax and the narrow. waist, or pedicel of the abdomen as far as the base of its large, swollen portion, the gaster. Here the gullet expands into a thin-walled, distensible sac, the erop, which is used for the storage of the imbibed liquids. At its posterior end the crop is separated from the ellipsoidal stomach by a peculiar valvular constriction, the proventriculus. The hindermost sections of the alimentary tract are the intestine and 400 THE SCIENTIFIC MONTHLY the large, pear-shaped rectum. The crop, proventriculus and stomach are the most interesting of these various organs. Forel calls the crop the ‘‘social stomach,’’ because its liquid contents are in great part distributed by regurgitation to the other members of the colony and because only a small portion, which is permitted to pass back through the proventricular valve and enter the stom- ach, is absorbed and utilized by the individual ant. That the erop functions in the manner deseribed can be readily demonstrated by permitting some pale yellow worker ant to gorge herself with syrup stained blue or red with an aniline dye. The ant’s gaster will gradually become vividly colored as the crop expands. Now if the insect be allowed to return to the nest, other workers will come up to it, beg for food with rapidly vibrating antenne and protrude their tongues, and very soon their crops, too, will become visible through the translucent gastric integument as they fill with the stained syrup. Then these workers in turn will distribute the food by regurgitation in the same manner till every member of the colony has at least a minute share of the blue or red ecropful of the first worker. The alimentary tract of the helpless, legless, soft-bodied ant erub or larva is much simpler than that of the adult. The mouth- parts are similar but more rudimentary. As a rule, the mandibles are less developed but in some larve they are strong, dentate and very sharp. The lower lip is fleshy and protrusible and provided with sensory papille instead of palpi, and the unpaired duct of the long, tubular and more or less branched salivary glands opens near its tip. The mouth-opening is broad and its lining in many species is provided with numerous transverse ridges beset with very minutes spinules (Fig. 62C). Larger, pointed projections or imbrications may also cover the basal portions of the mandibles. All these spinules and projections are probably used in triturating the food but perhaps when rubbed on one another they may also produce shrill sounds for the purpose of apprising the worker nurses of the hunger or discomfort of their charges. The gullet is long and very slender and opens directly into the large stomach, which throughout larval life is closed behind, that is, does not open into the intestine. A communication with the more posterior por- tion of the alimentary tract is not established till the larva is about to pupate. Then all the undigested food which has accumulated in the stomach since the very beginning of larval life is voided as a large black pellet, the meconium. In the larve of the Pseudomyrmine (Figs. 62, 68 and 64) there are certain very peculiar additional structures which may be briefly described. The head is not at the anterior end of the body as in other ant larve but pushed far back on the ventral surface so that it is surrounded by a great hood formed from the three thoracic SOCIAL LIFE AMONG THE INSECTS 401 / GAEE LL/ ey CLE AED: 4 FIG. 62 Larva of Pseudomyrma gracilis. A, ventral; B, lateral view; C, head and adjacent portions of same enlarged; D, sagittal section through anterior por- tion of larva. o, oral orifice; x, exudatoria; t, trophothylax, or pocket, which holds the pellet; (pe), deposited by the worker nurses and which is eaten by the larva. Note the hooked dorsal hairs of the larva, which serve to suspend it from the walls of the nest. a, mouth cavity, more enlarged to show the fine spinules (also seen in C), which serve to triturate the pellet and probably also as a stridulatory organ. Vol. XV.—26. 402 THE SCIENTIFIC MONTHLY FIG. 63 A, ventral; B, lateral view of the first larval stage (“trophidium”) of the Ethiopian Pachysima latifrons, showing the peculiar appendages (“exuda- toria”) surrounding the head. These belong to the three thoracic and the first abdominal segments. segments, and the first abdominal segment, which lies immediately behind the head, has in the midventral line a singular pocket, the trophothylax (t). Furthermore, each side of this segment and each ventrolateral portion of the several thoracic segments is de- veloped as a peculiar protuberance or appendage, which functions as a blood-gland, or exudatorium (2). Unlike the adult ants the larve can devour solid food, though they are often fed, at least in their youngest stages, with liquids regurgitated on their mouths by the worker nurses. The larve of the Pseudomyrmine are fed with the pellets (pe) from the in- frabueeal pocket, which are placed by the workers in the tropho- thylax where they are within easy reach of the mandibles and can be gradually drawn into the mouth, triturated and swallowed. Some primitive ants (Ponerine, some Myrmicine, ete.) actually feed their young with pieces of insects or entire small insects, which are simply placed on the ventral surface of the larva within reach of its mouth-parts. In a former lecture I referred to the fact that the larve of the social wasps, either before or after feeding, produce droplets of a sweet salivary secretion, which are eagerly imbibed by the adult wasps, and I designated this interchange of food between adult and larva as trophallaxis. I have recently made some ob- servations which show that the ant larve also produce secretions which appeal to the appetites of their nurses. These secretions are more varied than in the wasps. Certain ant larve undoubtedly SOCIAL LIFE AMONG THE INSECTS | 403 supply their nurses with saliva, but many or all sweat a fatty secretion through the delicate general integument of the body, and the larval Pseudomyrmine produce similar exudates from the papille or appendages above described. Although these various substances are produced in very small quantities they are of such qualities that they are eagerly sought by the adult ants. This explains much of the behavior which has been attributed to ma- ternal affection on the part of the queen and the workers, such as the continual licking and fondling of the larvae, the ferocity with which they are defended and the solicitude with which they are removed when the nest is disturbed. In other words, a de- eidedly egoistic appetite, and not a purely altruistic maternal anxiety for the welfare of the young constitutes the potent ‘‘drive’’ that initiates and sustains the intimate relations of the adult ants to the larve, just as the mutual regurgitation of food initiates and sustains the similar relations among the adult workers themselves. I am convineed that trophallaxis will prove to be the key to an understanding not only of the behavior I have briefly outlined Second, third and fourth (adult) larval stages of Pachysima latifrons, show-: ing the gradual dwindling of the exudatoria. A and B show the trophothy- lax (t); and B also shows the food pellet pe; which is the pellet formed in the infrabuccal pocket of the worker nurse; x, exudatcrium. See Figs. 62 and 63. 404 THE SCIENTIFIC MONTHLY but also of the relations which ants have acquired to many kinds of alien organisms. In the accompanying diagram (Fig. 65) I have endeavored to indicate how trophallaxis, originally developed as a mutual trophic relation between the queen ant and her brood, has expanded with the growth of the colony, like an ever-widening vortex, till it involves, first, all the adults as well as the brood Trophic Relations of Ants to Plants. Myrmecophytes Trophic Relations of Ants to Insects outsida the Rest. Trophallaxis batween Ants of Different Species, irophallaxis between Ants and True Guests, Trophailaxis botween Adult Ants, Trophallaxis between Mother or Adult Workers and Larval Brood, Mutual Regurgi- tation of Food Symphily Social Parasitism Trophobiesis FIG. 65 See text for explanation. and therefore the entire colony; second, a great number of alien insects that have managed to get a foot-hold in the nest as scavengers, predators and parasites (symphiles) ; third, alien social insects, that is, other species of ants (social parasites) ; fourth alien insects that live outside the nest and are ‘‘milked’’ by the ants (trophobionts), and fifth, certain plants that are regularly visited or even inhabited by the ants (myrmecophytes). These extranidal relationships, represented by the two outer rings in the diagram are, of course, incomplete or one-sided, since the organisms which they represent are not fed but merely cared for or protected by the ants. In my next lecture I shall have more to say about some of these relationships. (To be continued) WATER 405 WATER’ By Professor LAWRENCE J. HENDERSON HARVARD MEDICAL SCHOOL a task of science is one that may be fulfilled quite directly a by providing useful information of a practical sort for our immediate needs, or it may furnish us with general ideas which, if we are skilful enough and intelligent enough, we may use to ex- plain a great variety of phenomena. These general ideas when combined make up our conception of the universe. The latter function of science is the more important one. It is the one that brings science close to philosophy. It is the one that arouses the ereatest interest and enthusiasm in those who pursue the task of science. When we look for generalizations of this kind we always seek simple explanations of nature, confident that they may be used as guides to our action in particular cases and as means for gaining an ever increasing control of the world that we live in. Everything in the history of science justifies us in saying that it is the general interpretation of nature which, especially in the last hundred years, has given man his extraordinary increase in ma- terial power, an increase in power which he has used far from wisely no doubt, but which for better or for worse he possesses. In the ancient world there were a certain number of advances of this kind. The history of geometry, in spite of the labors of the last century and in spite of Einstein, is to a great extent ancient his- tory. The few necessary general conceptions that the Greeks found out enabled them to develop a complete science of geometry, enabled them to be good surveyors, enabled them to begin to understand how to lay out the heavens and the earth in their astronomy and in their geography, enabled them to see how to make buildings, and to do a great many practical things. But in the main the really important, the really general ideas of science, are modern, and of course the most interesting ones have to do, at least so most people will think, not with statie things, not with space taken by itself, but with dynamic things, with the question of what is happening in the world. On the whole, the most interesting and the most important generalizations of science 1This lecture was one of the public series given in Boston on Sunday afternoons recently under the auspices of the Harvard Medical School. 406 THE SCIENTIFIC MONTHLY will be judged by most men to be those that tell them what is hap- pening in the world, how things happen and how those happenings may be, if possible, changed to their own advantage, and how they may be interpreted so as to yield a philosophy. Of course everyone knows that the first step in this direction in modern times, the first step of great importance, was the inter- pretation of the phenomena of our solar system. It was the labors of Copernicus, of Galileo, and of Newton that enabled men to look at the sun and the moon and the planets, and to think quite clearly and quite simply and without any sort of difficulty at all about what their movements and their apparent changes in position and in form indicated. But, once more, it is not so much our solar system as it is our own earth that interests us, and until we come down to the earth and examine the happenings on the surface of the earth we are necessarily a long way from those things that interest us most. How shall we state abstractly what is happening on the earth? That is one of the great questions for science as well as for phil- osophy, for sociology, and for all the other intellectual activities of man. What is happening on the earth? Of course there are an indefinite number of answers to that question; but if we seek the strictly scientific answer of physical science I think there can be very little doubt that it is possible to put it in a few words which make our interpretation of the changes in nature surrounding us surprisingly simpler than they otherwise would be. Men in antiquity and in all ages have vaguely perceived an answer to this question. Perhaps many of you have heard of the famous remark of Thales, who is often thought of as the founder of philosophy and mathematics and science, that water is the origin of all things. This statement seems to many people to-day, I suppose, ridiculous, or at all events very exaggerated, but it is in fact much less ridiculous and much less exaggerated than you may suppose. A little later Empedocles and Aristotle, extending the elements from one to four, included water along with earth, air, and fire as the elements of which the world is made up. No doubt the word ‘‘element’’ is used in a sense a little different from our ordinary idea of an element to-day, but we still speak of the Aris- totlean idea when we talk about exposure to the elements. Of these elements water is in many respects the most interesting. Never- theless, while it was always clear in antiquity, as it has been in modern times, that water is very important in the world that sur- rounds us, its real importance is a comparatively recent discovery. In the first place, it was only a century and a half ago when WATER 407 Cavendish and Watt and others first found out that water is not itself what the chemist now calls an element, but that it is a com- pound, a compound of hydrogen with oxygen, a substance that can be formed by burning hydrogen gas in air or in oxygen. With this discovery a new road was opened, not only for chemistry and for physiology, but for the general interpretation of nature. Even before that time men had begun to understand the meteorological eycle, to perceive that there is going on on the earth a physical process which may be very briefly described as the evaporation of water from the ocean, from lakes and streams, and from the moist land, its dissemination as water vapor throughout the atmosphere by the winds, its precipitation as rain and snow and hail and dew, its collection into streams, but not a rapid collection, for it persists for long periods of time in the soil and underground, and finally its return to the ocean and reevaporation. That had been per- ceived, as I say, quite clearly before the discovery of the chemical nature of water. After that discovery another thing became apparent, that all that is happening in living organisms, in animals, and in plants on the earth, may be regarded in one sense as a part of this great meteorological cycle, this great cycle of water in its evaporation and condensation and flowing back through definite channels; for, as you know, our rivers persist, our streams persist, the circulation takes place in a very definite and very definitely canalized way. As I say, when men had made out the chemical nature of water they were able to see that water and earbonie acid are taken up by the plant, taken, if you will, out of the meteorological cycle and built up into sugar and starch, and then, sooner or later, turned into a great variety of substances, the substance of the plant, in short. These in turn serve as the food of the herbivorous animals, of cattle and sheep, and they are not greatly modified when built up into the body of the ox or the sheep. Next they may be con- sumed by man, or by a carnivorous animal, and are made over into the materials of his body. But sooner or later they are turned out into inorganic nature, excreted, as water and carbonic acid, after being burned in the body, once more to become a part of the meteorological cycle. That, I submit to you, is a fair statement, which no doubt needs a great deal of enlargement, a fair statement in terms of physics and chemistry, of what is happening on the surface of the earth. What is happening on the surface of the earth is, first of all, a circulation of water, a circulation of water which we recognize in the fall of rain, in the flow of streams, which we recognize equally in the growth of plants and in the feeding of animals on plants, 408 THE SCIENTIFIC MONTHLY and in the burning up of material in the body of the animal—of course a circulation of water and of other things, other things which in the total are very numerous and certainly very difficult to know in all cases, but fundamentally a circulation of water. I have not touched upon the energy of the process, the side of the question that is represented by the energy that goes into the water to evaporate it off the surface of the ocean, that goes into the green leaf to turn water and carbonic acid into sugar and starch, or the energy that comes out again in the form of muscular activity and body heat in you as you live, when you convert starch, sugar, and other things formed from them in the plant, back to water and carbonic acid. But let us say in the beginning that what is happening on the surface of the earth is this circulation of water, and let us add that it is a circulation which is driven by the energy of the sun, which is driven both in the one eyele, in the meteorologi- eal cycle, by the sun’s heat evaporating the water, and in the other, the organic cycle, by the sun’s energy making possible the forma- tion of sugar and starch from water and earbonie acid. In the first place, this process depends upon the great amount of water that there is on the earth. There is enough, if the earth had a perfectly smocth surface, to cover the whole earth, to a depth of two to three miles. How much of this is circulating it is very difficult to say. The amount is large. For instance, at the Equator evaporation takes off a layer of perhaps about seven feet every year from the tropical ocean. Of course the evaporation is less in other localities, and it is much less where there are no great bodies of water, but that will perhaps give you some idea of the magnitude of the process. How much is the energy involved in this process? How much work is done? How much energy of the sun does it take to evapo- rate so much water? A few square miles of the tropical ocean are taking up solar energy in the process of the evaporation of water to such an extent that if that energy were available in the bodies of the people of the United States, it would run all their bodies; that is to say, all the work that we are capable of doing, plus all the heat that we produce, is equivalent to an amount of energy which is no greater than that involved in evaporating the water off a very few, perhaps ten, square miles of the surface of the tropical ocean. So you see that not only is there a vast amount of water being evaporated, but that amount of water carries with it, involves in its evaporation, a prodigious amount of energy. Another way of studying the process is to try to form an idea of the run-off of the rivers of the earth. It has been estimated that the run-off is yearly, for all the rivers of the earth, about 6,500 WATER 409 cubic miles of water; that is to say, if you were to take a band a little more than two miles wide from Boston to San Francisco, and imagine it to rise up one mile into the air, that would give you a solid equivalent to the volume of water that pours back into the oceans yearly. And that of course is only a fraction of the water that evaporates yearly, because a great deal that evaporates and falls to the ground never reaches the ocean at all. This prodigious amount of water carries with it to the ocean what seems to be an almost equally prodigious amount of other material. There are something like 5,000,000,000 tons of dissolved substance in the river waters going back to the ocean yearly, and nobody knows how much undissolved sediment there may be. This dissolved material has been leached out of the earth, it is being leached out of the earth all the time, and the record of it is the accumulation of dissolved substances in the ocean as well as that long continued action of sedimentation which is on the whole the greatest of geological processes. In the sea, as a result of this, nearly all the chemical elements are contained in solution, most of them, to be sure, in such small amounts that they can not be measured, but so many of them are there that the sea water is quite a different thing from anything else in the world, and can not be imitated. Nobody ever made an artificial sea water that would serve the purposes of sea water as an environment for simple marine forms, as well as sea water itself. You may make as careful an analysis as you please of the sea water. You may put into pure distilled water all the things that you ean find in the sea water, and then if you try to make organisms live there they will not live in it—some of them, at least, will not—as well as they will live in genuine sea water. There is a good deal of resemblance between the salts dissolved in our blood and the salts dissolved in sea water. This has led Quin- ton to speculate about the use of sea water for medical purposes. He has made many experiments by diluting it to the proper degree and injecting it into the body. It has been suggested by Pro- fessor MacCallum that our blood is, so to speak, descended from sea water, that in the course of evolution somehow or other the fluids of the body originated as sea water. Of course single-celled organisms have no blood. When multi-cellular beings came into existence, where did the fluid which bathes the cells come from, provided multi-cellular individuals did develop from the uni- cellular? It is not a wild assumption to suppose that sea water furnished the inter-cellular liquid, that it was the material that first surrounded the several cells making up the complex organism. If you look into the whole story of comparative physiology that 410 THE SCIENTIFIC MONTHLY idea, while it must not be pushed too far, seems not an extravagant one, and if so, it is interesting to reflect that in that original simple fluid, simple compared with our blood in most respects, there were nevertheless a vast number of substances that had been leached out of the earth’s crust in the course of millions of years. This peculiarity of sea water depends upon the fact that water is, among all the liquids that we know, on the whole the best sol- vent, the one that can dissolve the greatest number and the greatest variety of substances of all kinds. That of course is a statistical statement. There are some things that water can not dissolve which can be dissolved by a great many other things; but by and large, taking everything that we know, taking all the chemical sub- stances that we know, there is not anything which on the whole is a better solvent, or capable of dissolving a larger number of things, or greater quantities of them, than water. Well, that is one of the very decisive facts in the meteorological eyele. It is one of the great factors in determining the geological action of water, in determining a large part of the evolution of the surface of the earth. And, on the other hand, in the organic eyele, in men, in animals, and in plants, water, always present in large quantities—your own body is three quarters water—con- tains a great number of things dissolved in it. This solvent action is certainly no less important in physiology than it is in geology. Not only does the water in your bodies, the water in the blood, the water inside the cells, the water in the lymph, contain a great many things dissolved in it, but it is almost exclusively in solution that things penetrate into your body. That of course may seem to you a strange statement. You are well aware that you eat many more or less solid substances—anybody ean swallow a lump of sugar, which is certainly a solid—but in fact what is geometric- ally inside your body is not necessarily physiologically inside your body. The digestive tube is physiologically not inside the body, and the process of digestion turns most substances that are swal- lowed in solid form into soluble products, which are dissolved in water and then in solution pass the real physiological barrier, the wall of the intestine, and so enter the body. This is no less true of the substances that are excreted from the body. The waste products are turned out of the body in solution, and if it were not possible to turn them out in solution it is very diffieult to imagine by what kind of physiological device it would be possible to carry on the activities of really active bodies and get rid of the waste products that have been burned up in the course of their activity. This great circulation of water has another very important influence upon the world, and upon the world of life, and that is WATER 411 its effect upon climate. Every one knows how much more steady is the state of the weather on small islands at great distances from continents than in most other places. Every one knows how much milder the climate is, how much cooler in summer and warmer in winter, at the sea shore than a comparatively small number of miles inland. This phenomenon depends upon water. How does it depend upon water? What is the effect that water exerts in that respect? Well, there are several factors. In the first place, it takes a great deal of heat to raise the temperature of water, or, as the physicist says, the specific heat of water is high. If you take, for instance, a pound of water and a pound of almost anything else—there are a few substances that are harder to heat than water—and heat them over a carefully regulated flame for a certain length of time, and measure the rise in temperature, you will find that the rise in temperature of the water is less than that of the other substance. As I say, there are a few exceptions, but there are very few. The result is that an ocean or a lake absorbs heat, and does not itself rise very much in temperature. Again, the evaporation of water takes up heat. Every one knows that. Every one knows that in order to evaporate water away at all rapidly you must heat it, and the amount of heat that is taken up in this evaporation of water is greater than in the evaporation of anything else; that is to say, you have got to put more heat into water in order to boil away or to evaporate, let us say, a pound of it, than you have in order to evaporate a pound of anything else. Thus the more rapid the evaporation the more effective the resistance of water to the rise of temperature, and for that reason the cooler the climate in the marine region com- pared with the climate in a region where there is no water to evaporate. This is one of the most important of all economic factors on the earth. It is a factor that, as much as any other one, per- haps, determines whether a given part of the earth is or is not really favorable for a high and active and prosperous civilization. But these factors are no less important in your own body than they are in determining the climate. As you know, you are con- stantly producing a considerable amount of heat, and that heat has somehow or other to be got rid of. If it were not, your body tem- perature would continue to rise. Well, in the first place, if your body were not mostly water, and therefore such that it takes a great deal of heat to raise its temperature one degree, a little ex- ercise might be impossible. If it were not for the fact that it takes so much heat to raise the temperature of the body a little, on ac- eount of the presence of water as its principal constituent, if it took only that amount of heat which is necessary upon the average 412 THE SCIENTIFIC MONTHLY to raise the temperature of most substances one degree, running a mile might be quite sufficient to produce a coagulation of all the albuminous material in the body, and therefore death. That is to say, the regulation of the temperature of the human body rests first of all upon this fact: that you have to heat water so much in order to raise its temperature a little, and then, in the second place, upon the effect of evaporation. The effect of evapo- ration is to cool the body very much, because you have to put a lot of heat into water in order to evaporate it, and since evapora- tion is the only way of cooling the body when the temperature of the environment rises to the temperature of the body this is a matter of the first importance, as everybody knows from his own experience in hot summer weather. It is a priceless advantage in the economy of your body that so much heat is taken up in the evaporation of water during sweating. One of the factors that greatly influence the circulation of water on the earth is the way in which it clings in the soil. Indeed water clings better, on the whole, than any other substance. This is due to the phenomenon which everybody has heard of as eapil- larity, and which is well illustrated by the action of a sponge in soaking up water. If you study this process you will find that it is easy to represent the sticking power of a liquid in the soil or in any finely divided matter by the height to which it can rise in a small capillary tube, in a very fine tube. Water rises to a very great height relatively to other substances in such a tube, and for the same reason it sticks very tight in the soil and water rises to a great height in the soil. That is one reason why great portions of the earth are habitable, or at least can grow crops. If water did not stick as well as it does, a large portion of the fertile earth would be sterile. But here again we have come upon a property of water which is just as important in the body as it is in the meteorological cycle. The living cell may be compared to a microscopic swamp, to a swamp of inconceivably fine dimensions. There is water running through it, and it consists of a very intricate meshwork of only partly known nature. In this swamp—this microscopic swamp, if I may call it that—these same forces, these same capillary forces, are of decisive importance. And here again it is the particularly ereat capillary activity of water that is one of the factors that determine the nature of physiological processes. There is one more point that I want to refer to about the in- organic, the meteorological cycle, because it is a particularly in- teresting and important one, although this one seems to have no highly important direct bearing upon our own bodies and our own WATER 413 life process. What causes rainfall? It takes a good many things to produce a precipitation of rain, but we may begin at the begin- ning and say that at least you can not have rain falling out of the air unless the air has become supersaturated with water vapor. The way to bring about that condition is to cool air that is pretty moist, because the amount of water vapor that the air can hold varies with the temperature. For instance, if you go down to the freezing point, to 32 degrees Fahrenheit, the amount of water va- por when the air is quite saturated is almost exactly one half of the amount of water vapor that is in the air at about 50 degrees Fahrenheit. Suppose, then, you have air that is three quarters saturated at 50 degrees Fahrenheit. Cool it down to 32 degrees Fahrenheit and it becomes not three quarters but three halves saturated; that it to say, it has 50 per cent. more water than it ean hold. That water will come out in the form of rain. This difference, a difference of 100 per cent. in the amount of water vapor that the air can hold, is far greater than the difference in the amount of the vapor of any other substance that the air can hold. That is to say, if you had any other substance at 50 degrees Fahrenheit, any other substance whatever, and the air were just three quarters saturated with it, and you cooled it down just to 32 degrees Fahrenheit, then you would not have reached the satura- tion point and the vapor would not fall out. If there were not this property, rain would be a comparatively rare occurrence. Moreover, unless rain falls, unless water comes out of the air, there can not be further evaporation, because you ean only evaporate into air which is more or less empty, so far as the water is concerned; that is to say, the whole meteorological eycle, the circulation of water, depends upon the rate of precipita- tion quite as much as upon the rate of evaporation, and the rate of precipitation of water is great because of this property of water. I think I have perhaps said enough to make clear to you some- thing of the natural importance of water. Such are the reasons why it is possible to say that what is happening on the earth in the last analysis is a circulation of water and the results of that cireu- lation. This is true not merely because there is a great deal of water on the earth, but also because water is a remarkable sub- stance that has a good many unique peculiarities. But in dis- cussing these properties, I have thus far spoken only about what may be ealled the physical peculiarities of water. Water is quite as important a substance chemically, and indeed I think we may say that the most important clues that we find in the properties of water to our understanding of what is happening in our every day life are to be found in the chemical properties of water rather 414 THE SCIENTIFIC MONTHLY than in its physical properties. This leads us back to the great French chemist Lavoisier, the most eminent victim of the French Revolution and the man who perhaps made the greatest contribu- tion to science of the eighteenth century. Lavoisier, studying the process of burning, of combustion, of oxidation as we call it, discovered that when things burn they combine with oxygen. Those of you who are not chemists and who have not studied chemistry will of course always feel a certain discomfort when one talks about the union of atoms. Atoms can not be seen. They are hard to imagine. But somehow or other the burning of tin for instance or the burning of mereury, is the combination of one or more atoms of the one ele- ment with one or more atoms of the other element. Tin and oxygen combine to form the oxide of tin. The atoms somehow or other fasten themselves together. That is what Lavoisier found out. He did not express it in terms of atoms, but he saw that these elements combined. So it is in the case of the burning of hydrogen, and water is a compound of hydrogen with oxygen. Atoms of hydrogen, two in number, combine with one atom of oxygen to form H,O, according to our present way of stating the ease. Well, having discovered the nature of combustion, Lavoisier turned to all kinds of cases where there was combustion. He proved, for instance, with Laplace, the great astronomer, in one of the most remarkable collaborations in the history of science, that in the last analysis our life activity, so far as chemical, is oxidation. The oxygen that we breathe into our lungs is com- bined with the various elements that make up our foods, and then the products are turned out as carbonic acid gas, which is nothing but the oxide of carbon, the result of burning carbon, and water. Then, reflecting further upon this, reflecting not only upon the chemistry of the process but also vaguely upon the energy that was involved, so far as it could be represented by the heat of the process, Lavoisier saw the real nature of the process, Lavoisier saw the real nature of the organic cycle and stated it clearly. He per- ceived that plants draw from the air that surrounds them and from the mineral kingdom the necessary materials of their organ- ization; animals take from the plants that which the plant has formed, directly in the case of the herbivorous animals, indirectly in the ease of the carnivorous animals, and build it up into their bodies, and finally the processes of combustion, fermentation, and putrefaction are continuously returning to the inorganic world the materials that were taken up originally by the piant. Of course WATER 415 these materials are not merely water, nor merely water and carbon dioxide, but also other substances. But note this. In the organic cyele water and earbon dioxide are free; the other things are fixed. Those things that are free in the physical sense are also free in the economic sense, under a wide variety of circumstances. You don’t have to pay, at least in New England and in many parts of the world, for the water and carbon dioxide which are the principal foods of the plant; all that you have to pay for are the so-called fertilizers, which make up an infinitesimal fraction of the material taken up by the plant. In- deed, if it were not for the mobility of water and carbon dioxide in this cycle that Lavoisier first understood, there would be no vegetation on the mountain tops. How could the material get there? The mountain tops would necessarily be bare. Not only the mountain tops, but everything would be bare except the waters of the earth. It is because water and carbon dioxide circulate, circulate rapidly and penetrate everywhere and stick when they get there, that there is a widespread vegetation, that there is an intense organic activity on the earth. But now what of this process that goes on in the plant? la- voisier could not understand it. It goes on in the green leaf, and the green leaf is truly the symbol of life. It is the starting point of life. It is the factory in which are created the materials of your body and mine, of the body of every living thing. These materials are created by the conversion of water and carbonic acid gas into sugar and oxygen through the influence of solar energy and with the fixation of that energy. Sunshine, water and earbon dioxide, if one may speak very loosely, are the components in the green leaf from which sugar or starch and oxygen are produced. The oxygen is turned out—the oxygen, please note, that had combined pre- viously in your body with hydrogen and earbon in the burning that is the essential process of your own activity. In the leaf this process is reversed. The energy of the sun is fixed in the leaf, and that is the source of the driving force of your body and of the driving force of every animal body. And not only that, it is the source of our fuels generally. If you will reflect for a moment, and remember that not only wood, but coal and petroleum, gasoline, what you will that is combustible upon this earth, is stored solar energy that has been fixed in the green leaf in the past, you will realize the extraordinary economic importance of this process. You will then realize that the cycle is, in fact, not only from the standpoint of the material changes, but also from the standpoint of energy, of horse power, of what you 416 THE SCIENTIFIC MONTHLY must buy and pay for as a source of any sort of material activity, the decisive factor on the earth. Since we can increase this process of photosynthesis, since there are still portions of the earth that are not adequately utilized, it is conceivable that one of the solutions of the problem of the increas- ing demand for energy may be to grow more available energy, for example, in the Amazon basin, where there are forests that it does not pay to cut at the present time. We might, for instance, turn a vast amount of solar energy that is not being utilized at the pres- ent time, or that is being expended in a manner that we can not ourselves turn to account into starch and sugar, into indus- trial aleohol, and so get a substitute for gasoline. That is an idea that has been in the minds of chemists, of course, for many years. One does not know how economic conditions will develop. At all events, we have here the clue to an understanding of the sources of energy on the earth. Aside from the fixation of energy in the organic cycle, and aside from the water power and other sources of energy in the inorganic cycle, there is little enough of any kind of energy that is available. You might perhaps have expected me to say something about water in medicine, since this is a medical school lecture. Water is © indeed important in medicine, but not, I suspect, in a manner that makes it possible for a lecturer to explain in two words its importance. There are diseases involving water. Of course dropsy involves the physiology of water in a remarkable degree. And there are processes that might be regarded as in a certan sense the opposite of dropsy, such as the curious dehydration of sick babies. In many eases they lose water, and it is difficult, or impossible, to get it back again. I can only say that perhaps because these are in some respects simple phenomena—i say in some respects—we know just enough about them to know that they are so complicated that it is really difficult to explain them or even to understand them at all. And so I shall, I fear, have to omit the more practical and immediate bearings of the physics and chemistry of water upon the organism, especially under pathological conditions. It is the principal constituent of our bodies, it is the principal substance that enters our bodies, it is the principal substance that leaves our bodies, and it is, as I have said, that substance whose movement in the inorganic world and in the organic world constitutes the first, the most fundamentally important, activity in the world that we live in. INTELLIGENCE TESTS 417 INTELLIGENCE TESTS OF CERTAIN IMMIGRANT GROUPS By Professor KIMBALL YOUNG UNIVERSITY OF OREGON HE influx of the foreigner into this country only became a Ah serious problem with the shifting of the Old World source of the immigration from North to South Hurope. The early situation in our national history was relatively simple. We had up to the Revolution, and forty years beyond, what constituted a genuine colonization. In fact, it was only after 1820 that a definite count was made of immigration. Even from this date until the Civil War, though the arrival of new-comers from Europe was con- stantly accelerated, the type of migration made for colonization of our free land and permanent citizenship.’ In the two decades following the outbreak of the Civil War, the curve of yearly in- crease of immigration began to rise very rapidly, yet the source of the stream still remained Northern and Western Europe and the British Isles. The destination of this immigration, however, began to be more and more the rising industrial centers of our country and decreasingly so the rural districts—although it appears that the peasantry of Seandinavia and Germany often continued into the free lands of the Far West, the industrial workers of Great Britain, Germany and the bulk of the Irish went into the urban centers more particularly. Wherever this immigration went, it nevertheless fitted fairly readily into our socio-economic folkways and mores, and the biological amalgamation of the ‘‘Older Immi- eration,’’ as it has been called, was in line with the racial stocks? already in the country. 1The existence of free land in America has had considerable influence upon our theories of government, our attitudes toward property, freedom and the socio-economic order generally. The significance of free land for racial amalgamation and cultural assimilation has been no less important, but less often noted. 2 Race is used in this article in the popular rather than in the strict anthro- pological sense. Properly speaking there are no ‘‘races’’ in Europe, but only ‘*sub-races.’’ Cf. Retzius, ‘‘The So-Called North European Race of Man- kind.’’ Jr. Roy. Anthrop. Institute, 39: 277-313. Cf. also introductory chap- ter in Reuter, E. B., The Mulatto (1918) on use of term ‘‘race.’’ Vol. XV.—27. 418 THE SCIENTIFIC MONTHLY With the ever-increasing concentration of industrial controls and the accompanying specialization, and withal simplification of production processes due to the introduction of machine methods, the demand for cheaper, semi-skilled and unskilled labor became imperative. The rise of Germany to industrial power, the reach- ing of a point of stable population growth in Great Britain and the Scandinavian countries eliminated them as sources of this cheap labor. In consequence, we have a noteworthy change to the South and Southeast of Europe, and even to Western Asia, for this sup- ply. In the short span of twenty years the shift in the center of gravity of the immigration to this country became the ‘‘common talk’’ of economies, sociology, political science and demography. The following table illustrates the now familiar facts: TABLE I38 SHOWING THE SHIFT IN THE SOURCE OF IMMIGRATION TO THE UNITED STATES (In terms of percentages of total from various sources) Country Year Percent. Year Percent.. Year Pereenw Great Britain. el. 1882 22.8 1907 8.8 1914 6.0 Mer rien iy; (ps ess eee 1882 31.7 1907 2.9 1914 3.0 SKOeA CORN tena NN meee Sars te aa 1882 13.3 1907 3.9 1914 2.0 Total Western European COUNTS) \eoete aia... 1882 71.3 1907 17e¥ 1914 4 vase Mitaalig | VES At eee geen 1882 4.1 1907 22.2 1914 26.5 PATI SETO- ENUM GAT; Yeteer sens deen onan 1882 3.7 1907 26.3 1914 25.6 RUSsian) AME eeseereeee eee 1882 2.7 1907 20.1 1914 23.0 Total Southeast and Eastern OD jm oo) oYe hte ane sess ee a 1882 18.2 1907 75.5 1914 75.6 The complete significance of this change in the source of our immigration we do not yet know. If the theory of General F. A. Walker be true, we may doubt whether immigration has been as important or necessary for the growth of the country, since 1850 anyway, aS some persons imagine. Had we had no immigration from the middle of the last century on, according to Walker, our present population would be as large as it is to-day, but racially we should be a much more homogeneous people, or, at least, in the way of becoming so. Now the meaning of the coming of the ‘‘New Immigration,’’ as that from Southern Europe has been called, in the matter of racial mixture is an unknown quantity. The writer believes, how- ever, that there is accumulating evidence from studies in general 3 Table compiled from Ellwood, C. A., Modern Social Problems (1913) and reports of immigration. The term ‘‘Russian Empire’’ in the table included Russian Poland and other sections now cut away from the former empire. Racially most of the contribution from Russia was Hebraic or Polish rather than strictly Russian. INTELLIGENCE TESTS 419 intelligence of certain immigrant groups, at least, which material, coupled with the results of researches in the inheritance of mental traits, should cause us to consider rather carefully the bearing which these facts may have on features of the racial mixture that will surely come out of the shift in the source and nature of im- migration.* The present paper will deal specifically with certain samples of the South European immigration in terms of general intelligence. The special question is: Leaving aside physical characteristics, differences in emotional traits and cultural contacts, what are the significant findings of psychology concerning the general intelli- gence of certain of these immigrant groups that have come to us so overwhelmingly in the past thirty years? The nature of general intelligence need not detain us here. The uninformed should refer to the long series of important studies upon this question initiated by Binet in France, and expanded by Stern and his pupils in Germany, and particularly by Goddard, Terman, Thorndike and Yerkes in the United States, and more recently still taken up by a group of English psychologists—Webb, Burt, Hart and others. There are obviously several problems as to the exact nature of so-called general intelligence, its growth, its importance in successful life career, and especially as to its in- nateness (that is wm potentia). On the latter point, the evidence at hand seems sufficiently valid to establish that general as well as specific abilities are transmitted by heredity. The precise biological nature of the units involved we do not yet know. Special talents may actually turn out to be due to the presence of separate units in the germ plasm, and all-around ability may be due to a conflux of several factors—multiple and difficult to segregate in the chromo- somes. Whether in fact, general intelligence is shown to be simply a convergence of special traits in certain patterns, as Thorndike might assume, or whether due to a more general factor as Spear- man seems to hold is unimportant for our present paper. Though the entire data on the mechanism of inheritance are not at hand, and even admitting that the subtle influences of environment have not always been completely segregated and controlled in the studies made, the writer believes that our social programs for education, Americanization, amalgamation of the foreigner racially, can not nonchalantly ignore these important implications. 4 Studies of the second and third generation of certain immigrant stocks are beginning to accumulate, but so far we lack psychological measures of the mental capacity of these various generations,—more especially of the strains that have gone into the mixtures. The writer, however, is in much sympathy with such studies as Drachsler’s and others. 420 THE SCIENTIFIC MONTHLY The studies of Galton, Woods, Pearson, Davenport, Thorndike, Earle, Starch, and others revealing the inheritance of intellectual traits appear reliable enough, irrespective of the peculiar biological form which subsequent research may reveal, to make the present status of the mentality of immigrant groups, if known, rather prognostic, to say the least, of what the mentality of the future generations of these peoples will be. In other words, the evidence appears rather conclusive for the inheritance of mental abilities, and if general intelligence tests reveal a given level of intelligence in an immigrant group may we not assume that we can predict something of the mental endowment which such a group will add to the future mixtures with other racial groups? It is well recognized by all persons interested in racial men- tality and has been recently reiterated by Major Leonard Dar- win® that what we want is a high average intelligence in the masses, not a small group of selected superior intelligences, with the bulk of the population of low or inferior intelligence. Out of the con- stantly changing matrix of a high average intelligence will arise the superior-minded persons who will be able to make the outstand- ing cultural contributions of the future, while the good average mentality of the masses makes for solid eitizenship, appreciation of high cultural values and successful group spirit. Any mixture which will lower the level of intelligence of a group and restrict the variability of the same will be deleterious. For instanee, if the average intelligence of certain of the South Huropean stocks which have come to us in the past twenty-five years should prove to be as high as that of the older American stock (i. e., of North Huro- pean ancestry) the problem of mixing the older ana the newer stocks, so far as general intelligence is eoneerned, will not be se- rious. However, on the other hand, if the Latins, say, who come to us should prove to be but four fifths as intelligent on the average and less variable when compared with the North European off- spring in this country, the racial mixture between the two may be damaging to the welfare of the country. With the methods of testing general intelligence that have come out of psychological investigation and with the implication of heredity in mental endowment, let us review the significant studies which throw light on the matter of immigrant mentality. On the basis of these we may then be able to note certain important bearings of these findings on the problem of immigration as it relates to racial mixture and the general intelligence of our future population. 5 Darwin, L., ‘‘The Field of Bugenie Reform.’’ THE Screntimric MonrHLY, 13: 392. ee se Bg a a he er = = a eT ee INTELLIGENCE TESTS 421 The writer will review three or four studies made during the past five years which deal with racial stocks in various parts of this country and will then add notes on his own contribution which grew out of a rather extended investigation of certain South Eu- ropean groups in California. In one chapter of the published data from the psychological testing of men in the army® is shown the relative standing in the mental tests of the foreign-born men of various nationalities found in that portion of the army which was tested. Each man tested in the army was given a final letter rating ranging from A to HE depending upon his raw score in either the alpha, beta or individ- ual test. The alpha, as is now well-known, was devised for literates of ten years mental age and above, the beta was devised primarily for men of low-grade mentality, but was also used for illiterates or for men who did not understand the English language well enough to handle alpha. The individual tests were exclusively used for low-grade mentalities who did not score well or at all in the group tests (alpha and beta.) The men of high mentality were given ratings of A and B. Men in these classes were considered, other things being equal, as being prospective officer material. Grades C and D made up men of the grades of privates, but those of D rating were discovered to be rather poor military ‘‘risks,’’ to borrow a term from insurance practice. The following nationality groups are selected as samples from a longer list showing the rank order of countries according to the percentage of men in each nationality group receiving their final letter ratings in D, D—, and E, and also in A and B. TABLE II? SHOWING THE PERCENTAGE OF MEN OF SAMPLE NATIONALITIES RECEIVING FINAL LETTER RATINGS UNDER THE GIVEN CLASSIFICATIONS Nationality Per cent. D, D—and E Per cent. A and B MRotalpawWihite: (Dirac tis cer oo ee een eee 24.1 12.1 SMe One TONY COUTEEICS erste se ee eee eee nee an Ey 4.0 Giirye em 20 Ch SoS ee re Sem 2: 8.7 UBHi RODMIAI Yh 2a nace fs css eaut das atde cee 15.0 8.3 ROW Ge Orit cae oie: Fc ae Oe se eee ea 19.4 4.3 PARIS Ted eM eee oss LE 11 ee Se Se he 3.4 Diy rfe Eolya 6 EA Mes nade wl ete enone nNOH OTe CU Se NP re 39.4 4.1 EVEL SST Moses ee aed ik Gs eee EL as ee 60.4 2.7 VITA Dt RE tet Se Lege eee Ee LL lh a 63.4 8 LECALH Xa In tade ADAM Ore eMen Ree amen site Let ete Alf te yy Wea 69.9 3) 6 Memoirs of the National Academy of Sciences, 15. Psychological Ex- amining in the United States Army, Pt. III, pp. 693-700. ‘op. cit. p. 696: The omitted nationality groups fall variously in the range of the percentages given in this table. The totals for the white draft and for the total foreign nationalities are given for comparison. 422 THE SCIENTIFIC MONTHLY In general the superiority of the Nordic stocks over the others is evident in this table. No assertions of language handicap can be brought against these findings, for the men who were unable to manipulate the verbal test, alpha, were given the performance test, beta, or the individual performance tests. Over 85 per cent. of the Italian group, more than 80 per cent. of the Polish group and 75 per cent. of the Greeks received their final letter grades from the beta or other performance examinations. The corre- sponding percentages for the Northwestern Europeans were: for the English, about 8 per cent, for the Germans slightly over 30 per cent., for the Swedish about 58 per cent. It is interesting to compare these figures with those compiled for the Southern and Northern negro draftees: 82.1 per cent. of the former received final letter ratings of D grade or lower; for the latter the like percentage was 47.2. It will be enlightening to recall these comparisons of the negro draft with the Italians and others when reference is made to Miss Murdoch’s investigation. Dickson’ in his study of first grade school children in certain publie schools in California, while his number of cases was small, found important differences in the intelligence quotients of the various groups when tabulated unaer racial stocks. A summary of his data follows: TABLE III SHOWING DIFFERENCES IN INTELLIGENCE QUOTIENTS IN FIRST GRADERS BY RACIAL STOCK Race No. of cases Median I.Q. SOP EUTRL Set eac Ee aie eee os oo ec, Se eae nomi lee ee ema 37 78 PP OTUU UCSC (ee cseee ese coset ore coehea ca saenteh scans eecocnsabecsnpncct en stueteces 23 84 italian’ (chiefly, Southern® [taiiy)) es) -2 eee seen 25 84 INiorth- Hr o pea mpm pO rrisse es 22 22 ee oe eee ees 14 105 American (North-European ancestry).......................--. 49 106 With the average intelligence as measured by careful Binet tests revealing such differences in children of the first grade (the chil- dren were practically all of the same chronological age) would not the educability, the future economic success, and the contributions to national culture and well-being be differentially in favor of those of higher intelligence? True, the overlapping in such groups is considerable. The figures do not mean that all the Spanish do but four fifths as well as the children of North European stock. Some of the Spanish children have intelligence quotients above the average of the latter groups, but, on the other hand, the lower quotients of the Spanish group fall below the quotient of the lowest American child. So, too, the Portuguese and Italian groups over- 8 V. Dickson, Unpublished Ph.D. thesis (Stanford, 1919), Tables from notes of Dr. L. M. Terman. Dickson’s study will soon be off the press. INTELLIGENCE TESTS 423 lap the other groups greatly. It must never be ignored that in dealing with averages in intelligence tests, the matter of varia- bility must also be taken into account.® The objection might well be raised that the differences in these racial groups found by Dickson are due to accident of environment which would be eliminated later. Would not, for example, school contacts wipe out, in large part, these differences? Might not home infiuences, such as use of the foreign tongue by parents and chil- dren, affect the standing of the Latin children in these tests? With a view to investigating these and allied questions, Miss Thomson’? five years later undertook the re-testing with Binet tests of the same children whom Dickson examined. In a period five eighths the total length of time demanded by our elementary public school curriculum one would imagine that any grave handicaps of language or family background would be largely overcome. After correcting Dickson’s tables on the basis of chance errors in chrono- logical age in his group, and eliminating, of course, from her group the eases which could not be found for re-testing, she presents the following table: TABLE IV SHOWING THE CHANGE IN MEDIAN I. Q. IN FIVE YEARS FOR RACIAL GROUPS Race 1916 median I.Q. 1920 median I. Q. Total South Eur op ears ee See eee eee ee eee eee 88.0 85.5 WOREW PUGS) fae. ee ele a eee 88.3 74.0 Americans) (from! One) SCHOO) ese see seeeeee sess 111.0 110.5 Americans (from second school)............:..-.....----- 102.3 95.0 In another table somewhat differently arranged showing the ancestry of the pupils she gives the following from her own data: TABLE V SHOWING THE MEDIAN I. Q. OF CERTAIN RACIAL GROUPS IN PUBLIC SCHOOLS Race Median I. Q. Range of I. Q. South’; Huropean=::::..-f2\2.2 22 ee eee eee 81.0 51 to 117.5 INonthiebire pea born! 10.25 k) ec areas soe 97.5 79 to 114.0 American born (North Huropean ancestry )............ 102.0 62 to 139.9 While the number of cases is small for wide conclusions, it is apparent that the American group not only has the highest median 9It is unfortunate that the P. E.’s of these distributions were not at hand. 10 Thomson, Mildred, ‘‘ Validity of Stanford-Binet Tests as a Basis of Prediction of School Suecess’’ (A.M. thesis—Stanford University, 1920— unpublished). It must be noted that Miss Thomson was unable to secure more than two-thirds of the same children as Dickson tested, but there is no reason for believing this reduction in the number of cases was due to any- thing other than chance. Speaking in terms of intelligence, there is as much likelihood of an American family moving away as a foreign one. 424 THE SCIENTIFIC MONTHLY I. Q., but that it is the most variable in range of I. Q. Miss Thomson notes: It seems that as a whole the tests are as accurate a judgment of the mental capacity of the low foreign clement as of the American children... As to the matter of the alleged language handicap of the children of foreign parentage she comments: Had language difficulty caused the first low median, the central tendency to change would now be plus. (That is, the overcoming of the language handicap by additional years in school would raise the median I. Q.) .... It seems evident that although the tests involve the use and understanding of language, low scores result not from the failure to understand, but from the failure to comprehend. Miss Murdoch" has studied four racial groups in certain publie schools in New York City in order to discover any mental differ- ences in these groups with special reference to hght on the problem of Americanization. Her groups included Hebrew, Italian, ‘* Ameri- ean’’ and negro boys in the upper grades. The children all under- stood Enelish, the Italian group being especially hand-picked, that is to say, only the boys who in the opinion of the school authorities did not suffer from any language handicap were tested. Miss Mur- doch believes this selection of Italians was in favor of the brighter boys in this nationality. The children were not selected by age, but by grade, ranging from the fifth to eighth, inclusive, which means that the age groups ten, eleven, twelve and thirteen were most frequently represented. This grade selection, however, elimi- nated from the comparison all the pupils ten, eleven, twelve and thirteen who were still in the lower grades. This means that the selection was likely in favor of the more intelligent pupils of these age groups. The groups came from two different sections of New York City, both described as rather ‘‘undesirable.’’ The writer of this article says: At least, between Jewish and Italian boys and between white and colored boys respectively, no allowance for difference in neighborhood environment need be made. The test used was devised by Pressey along the lines of the army alpha. Close to 500 cases each of Jewish, American, and Italian boys and 225 negro boys were examined. The Hebrew and Ameri- ean children had practically the same averages in the tests. The Itahans and negroes while running somewhat together, with the latter slightly better, were decidedly inferior to the other two groups as measured by these tests. Taking the twelve-year-olds 11 Murdoch, K., ‘‘A Study of Race Differences in New York City.’’ School and Society, 11 (1920): 147-50. INTELLIGENCE TESTS 425 who were tested as a sample—in the Italian group 13.5 per cent. equal or exceed the 50th percentile or median of the Jewish boys; 24 per cent. of the negroes exceed the median of the Jewish; and 50 per cent. of the Americans likewise. For the three age-groups having the largest numbers in the total group studied, i. e., the eleven, twelve, and thirteen-year-olds—the average is 15.5 per cent. for the Italians, 30.5 per cent. for the negroes, 53.7 per cent. for the ‘‘Americans’’ equaling or exceeding the median for the Hebrews.” Whether the children studied according to age in relation to test results, or by grade location the Italians maintained ‘‘their position at the foot of the four races.’’ Language can not account for the low scores of the Italian, for, as already noted, those who were suffering from difficulties with the English language were excused from the tests. Miss Murdoch writes: The fact remains that Italians who were thought by their teachers, prin- cipal, and the neighborhood social workers to be laboring under no language handicap were found to be very inferior to the other three races. However, since the test used by Miss Murdoch was based upon verbal concepts, in part at least, the accusation may still be made that the Italan home does not furnish the English tools to its chil- dren so well as the Hebrew, negro, or American. Let us look fur- ther to see what additional studies have shown. In order to test the matter of alleged language handicap and to investigate the general question of mental differences in immigrant groups in this country, the writer of the present article undertook in the years 1919 and 1920 to apply psychological tests to the ehil- dren of certain South European and Latin American immigrants— viz.: South Itahans, Portuguese, and Spanish-Mexicans—and to compare the same with groups of children of Northern and Western European aneestry.** Over 90 per cent. of the Latin stock (as we may call the former for brevity) studied were born in this country and practically all of the children of the latter stocks, or Non-Latin as they have been called. The survey was made in several cities and towns of the west-central portion of the state of California. Several measures of mental differences, direct and indirect, were made. Such outside criteria as the grade location, teachers’ 12'The number of negroes at these ages is too small to warrant any wide conclusion concerning the intelligence of negroes compared with Italians. The negroes were a much more highly selected group perhaps than the Italians, further, the Northern negro is on the average higher than the average intelli- gence of the entire race in this country. 18 This study will be published soon in The University of Oregon Publi- cations under the title: ‘‘Mental Differences in Certain Immigrant Groups.’?’ 426 THE SCIENTIFIC MONTHLY estimates of intelligence and of the quality of schoolwork were made and compared with the results of the psychological tests themselves. Of the latter there were two batteries of so-called general intelligence tests: the first, the army alpha, is a verbal test based quite extensively upon language and dealing with con- cepts of verbal nature; the other, a slightly modified army beta, adapted to school children who understand at least a modicum of English necessary for instructions. Rather than selecting the school children from certain grades the writer selected all the twelve-year-old children in the schools which avoided the biased sampling that must have arisen had children in one or two grades only been selected. By this method, then, all the twelve-year-olds from the entire public school systems in the communities surveyed were studied irrespective of their grade location. Incidentally the manner in which these twelve-year-olds distributed themselves through the grades indicates once again the fact that chronological age is no criterion to the pedagogical age of a child, of whatever racial stock. This is no place for a detailed report of the findings in this study, but the pertinent facts for the problem of immigrant men- tality will be briefly given. It was found that the actual grade lo- eation of the children is the best single criterion of their intelli- gence aside from the mental tests themselves. The comparative standings of the Latin with the non-Latin groups shows that the modal grade for the latter is the low seventh, for the South Italians, the high fifth, for the Spanish-Mexicans, the high fourth, for the Portuguese, the fifth.1* On the basis of the teachers’ estimates of general intelligence on a seven-rank scale the Italians are on the average .8 of one class-rank below the ‘‘Americans,’’ the Portu- guese are over one class-rank below the latter, and the Spanish- Mexicans over one and one quarter class-ranks below. Coming directly to the results of the tests themselves, the im- portant fact for us is that of overlapping of intelligence in the groups. The following tables give first the absolute differences as measured by the tests, then the measurement of the overlapping: TABLE VI (A) SHOWING THE DIFFERENCES IN AVERAGES AND IN VARIABILITY IN GROUPS Race group Median Alpha P.E. dist. Median Beta P. E. dist. eAmervean (7) | Uae eee oo o2) Od: 18.48 68.88 7.03 Ufa ma. 2222 se eee ke 24,92 13.90 54.75 10.90 Portugtese |) yoo eh 22.00 15.65 52.03 9.88 Spanish-Mexiean. ¢ s2c-c-- coe. -conc-0c0: 23.67 12.82 52.96 10.76 14 The Portuguese group was not segregatable into half grades. INTELLIGENCE TESTS 427 (B) ALPHA RESULTS SHOWING PERCENTAGES OF FOLLOWING RACIAL GROUPS EXCEEDING THE QUARTILES OF THE AMERICAN GROUP (Actual and theoretical)15 Per cent. exceed. Per cent. exceed. Per cent. exceed. American Q1 American Median American Q3 Race group (Actual) (Theor.) (Actual) (Theor.) (Actual) (Theor.) C1 DET Vl 24.50 22.66 7.00 4.95 1.00 eats) PBOMUWGOUCSE! -cc-cndeseceneas 22.50 23.88 10.00 6.55 2.00 1.07 Spanish-Mexican _ ...... 21.50.... 17.36 6.80 2.81 75 .20 (C) BETA RESULTS (as under B) LUG 5 aor 33.40 31.21 18.00 16.11 8.25 6.94 IPOTUUSUCSO! | <.sccessn-c-n- 29.00 25.46 13.75 11.51 4.15 4.09 Spanish-Mexican ...... 30.40 27.76 15.50 13.79 6.40 5.59 From the average for the American twelve-year-olds it looks at once as though measured by the alpha the Latins are decidedly inferior to the Americans. The beta averages, while more nearly alike for the Latins and Americans, still show a marked superiority for the latter. It must be noted that the range of the beta units is just half that of the alpha, hence the differences in the actual figures in the table are not indicative of the real differences re- vealed. It is still true, however, that the Latins approach their competitors more closely in the beta than in the alpha tests. The latter is based, as we noted, upon verbal concepts, while the latter is a so-called performance test operated independently of lan- guage. Were it not for the differences shown in other measures of intelligence, as, for instance, grade location, teachers’ estimates of intelligence, and school marks, it might be imagined that the differences brought out in beta are more truly significant of the differences in ability than alpha. I have shown elsewhere that this is not the case,'® but space prevents a lengthy discussion of the matter here. The following points summarize these facts: First, the beta tests proved too easy for the American twelve- year-olds who were above their years in intelligence. This means that the tests were not diagnostic enough for these children and that there was a piling up of high scores for these children at the upper end of the scale. This narrowing of the spread of their actual abilities is the reason for the lower P.E. of the Americans in beta as compared with the Latins. Had the tests been more suited to their real mental capacities, some of them would have scored higher and the average for their group would have risen, and their variability as a group would have been greater. For the Latins the tests proved well suited for their capacities. 15 The figures tabulated under ‘‘theoretical’’ are the computed measures of overlapping when the reliability of the tests are known. Cf. Kelley, T. L., ‘*The Measurement of Overlapping.’’ Jr. Educ. Psychol., 10: 458ff. 16 Discussed in writer’s study mentioned in footnote No. 13 above. 428 THE SCIENTIFIC MONTHLY Second, checking the alpha by correlation against the outside eriteria, such as, grade location, teachers’ estimates of intelligence, and the school marks, it was found that the alpha, the verbal test, was more diagnostic of the mentality and hence educability of the Latins and the Americans both than was the beta. In addition these correlations go to indicate that the asserted language handicap under which the foreign children are supposed to labor does not exist, at least so extensively as imagined.’ As Miss Thomson put it the failure in the tests and in school is not based so much on failure to understand as failure to comprehend. The measures of overlapping (sections B and C of above table) reveal more significant facts of mental differences in these groups than the differences in average score. If these tests are at all true measures of differences, and there is certainly much evidence that they constitute the best measure of mentality science has yet pro- duced, may we not assume that the extent of the overlapping among the groups clearly concerns an important problem before us, name- ly: the relation of these differences to future racial mixture which is subsequent to our enormous immigration? If there is anything preponderant in mental heredity it appears that this question must be answered. Taking the South Italian group as a sample of the Latins and near Latins (as we may term the Spanish-Mexicans),—if they equalled the American group, 75 per cent. of them should exceed the first quartile or 25 per cent. of the latter. As a matter of fact in alpha less than 25 per cent. of them exceed the lowest 25 per cent. of the Americans. So, too, with the median or second quartile, but 7 per cent. of the South Italians exceed the lower 50 per cent. of the Americans, and but 1 per cent. exceed the upper 25 per cent. of the latter. It may be that alpha is unjust to the Latins even though there is considerable evidence that it is on the whole quite adequate for diagnosis of mental differences. It may even be that the language difficulty which a proportion of the Latins eertainly had when they entered the public school has persisted until its effects are 17 It is rather interesting to recall that thirteen years ago Ayres in his book, Laggards in Our Schools, pointed out that language difficulty did not operate as a factor in school retardation. He wrote: ‘‘Wherever studies have been made of the progress of children through the grades, it has been found that ignorance of the English language does not constitute a serious handicap.’’ (p. 116). Of course no one would argue that in children newly arrived in this country from foreign countries speaking another language would not be handicapped in the period of acquiring our tongue. But beyond that period, the trouble in retardation lies in other directions. . INTELLIGENCE TESTS 429 felt as late as five or six years after entry.'* Therefore for those who would still maintain that alpha is too distinctly advantageous to American children, the overlapping in beta, in spite of the fact that the evidence is clear that this test is unfair to the American twelve-year-olds, still shows the latter superior. But 33 per cent. of the Italians exceed the lowest 25 per cent. of the Americans, but 18 per cent. exceed the lower 50 per cent. and’ but slightly over 8 per cent. the upper 25 per cent. There is no evidence that the beta was in any way unfair to the Latins either in content or presenta- tion. All the children were tested by the writer personally in small eroups and in each session members of all the racial stocks were present. The Portuguese and the Spanish-Mexicans compare less favorably with the Americans than the South Italians. We may now conveniently summarize the results of the experi- mental findings on this subject before passing to interpretative paragraphs. The army tests on recruits and draftees showed that the Latins and other Southern European (also certain Eastern European) eroups were decidedly inferior to the men who were natives of Northern and Northwestern Europe. The Dickson-Thomson studies have presented good evidence that the intelligence of the racial eroups, at least of those investigated, does not improve by eduea- tion, and further that in the samples which they study the Latin eroups are on the average eight tenths (.8) the average intelligence of the American children of North European ancestry. Murdoch’s survey revealed that as measured by the Pressey seale of general intelligence, the Italian groups which she studied were decidedly inferior to the American and Hebrew ehildren tested, and even slightly lower than the negro children measured at the same time. Her twelve-year-old group (which we noted was a selected’ sample) compares quite noticeably with the writer’s. In her results she reports that 13.5 per cent. of the Italians equaled or exceeded the median of the Americans. The writer’s figure for a similar meas- urement of overlapping was but 7 per cent., but the difference may be due partly to the nature of the test and partly to the fact that his sample was not at all handpicked. It may be due to actual differences in the intelligence of the group. The important point, however, is the likeness of the two studies. 18 The writer in answering those who maintain that all differences in mentality (barring the extreme feeble-minded) are due to environment and opportunity, is always puzzled to know how these persons account for the dis- tribution of children at 12 years of age, some in third, fourth, fifth, sixth and seventh grades, when they all started in school at the same time. 430 THE SCIENTIFIC MONTHLY One may use norms which were developed’® for translating the alpha scores of school children into Binet mental ages. If this is done it is found that the Latins measured are on the average two years retarded as compared with the American groups studied. Again the caution must be repeated that the range of scores in each eroup is wide, meaning that the differences are relative not abso- lute. Space forbids further comment on the studies themselves, but we must turn before concluding to a few paragraphs on the probable bearing of these findings upon the problem of immigra- tion and the mixture of immigrant stocks in this country. If the researches into mental heredity have the validity which we suspect, then the differences shown in these investigations just cited are simply reflections of what is found in the general adult population. The findings of racial differences in immigrant stocks revealed in the army lend weight to this assumption. Further it is interesting that the preliminary testing of the Japanese and Chinese made in California indicates that these Oriental immi- erant stocks compare very favorably with the American popula- tion of North European ancestry in the same neighborhoods. Surely the language handicap is of greater potency in the Oriental than in the European. So it seems that these methods of testing have enough pertinence to make us pause and consider the meaning of their findings. As noted at the outset of this paper, we have shifted the source of our immigration from North to South and East Europe, and further we have drawn more and more upon a low type of peas- antry as compared with the rather good average industrial and agricultural classes that formerly came to us from German, Sean- dinavian and Anglo-Saxon countries. As we noted above*® the problem of mixing racial stocks is one of producing high average intelligence in the masses so as to lay the biological chances for the production of each generation of the superior persons who will make the greatest advances for their generation. If the mentality of the South Europeans who are flooding this country is typified by the mentality of the three groups studied by the writer and others, the problem of future standard of living, high grade citizenship and cultural progress is serious. The writer does not attempt to say whether these findings are 19 From norms developed by Drs. 8. C. Kohs and W. M. Proctor at Stan- ford University (1918-19). Cf. Proctor, W. M., ‘‘Psychological Tests and Guidance of High School Pupils.’’ Jr. Educ. Research, Monog. 1, No. 1, pp. 70 (1921). 20 supra, p. 5. INTELLIGENCE TESTS 431 typical of the entire racial groups from which these samples came. None of these studies is an investigation of the whole population of Portugal, South Italy, Spain or Mexico. They are surveys of these stocks as actually found in the usual immigrant settlements in this county. However, the writer stands firmly upon the ground that the evidence points to the fact that these samples are very typical of the immigrants who reach this country from these Euro- pean sources. Is not this after all the significant thing? For in- stance 85 per cent. of the total immigration from Italy in the past 30 years has been from the same districts of South Italy from which the largest sample of the writer’s own investigation and that of Mr. Dickson and Miss Thomson came. The indirect evidence is that Miss Murdoch’s Italian subjects were also from South rather than North Italy. In fact it is not impossible that the Italian in California is even a little superior on the average to his confrére who has neither energy nor ambition to get out of the industrial centers where most of these people first settle in our country. Of the Portuguese the writer can only venture a guess that he is much like the Portuguese found elsewhere in the United States, princi- pally in New England. The Spanish-Mexican groups appear typical of the immigrants of this stock. Does not the evidence accumulating, one phase of which has been reviewed here, point conclusively to the fact that a continued deluge of this country of the weaker stocks of Europe will ulti- mately affect the average intelligence of the population of this country? It is comment everywhere that the better stocks are losing ground compared with the poorer in the matter of offspring. The more intelligent classes are practicing, in one way or another, a conscious control of the number of their children. The result of this must mean a shifting in the average intelligence of the popu- lation toward that of the poorer stocks. Now hybridization of stocks is already taking place. It does not seem to be true that the inferior stocks always mix with their own kind—the history of the Kallikaks and such like proves that these stocks are con- stantly sending out their tenacles into the higher biological strains. Have we not been caught in the myth of the Melting Pot and in a sort of pious wish that all was well and that nothing could happen to us? The general economic progress of the past fifty years may have made us myopic of the larger meaning of these accompanying changes in our population. As Irving Fisher put it :?4 Mechanical inventions . . . have given us more and more room for expan- sion and we have mistaken this progressive conquest of nature for a progres- sive improvement in ourselves. 21 Fisher, I.: ‘‘Impending Problems of Eugenies.’’ THe Screntiric MonrTsLy, 13: 215. 432 THE SCIENTIFIC MONTHLY Certain sentimentalists have talked elibly of assimilation of racial stocks. This presupposes the capacity of one stock com- pletely to take over another and make as the original stock was. There may be cultural assimilation (even this is doubtful in a technical sense), but there can not be biological assimilation. What we have is amalgamation, and as Conklin and others have pointed out, amalgamation means a hybrid stock, a stock compounded out of the elements of the two or more strains running into the new generation. It is true that biologically the mixing of the North and South European is hardly analogous to the mixing say of the white and the black races or the whites and the mongoloids. Biologically as previously noted?? the HKuropean stocks are actually sub-races. Furthermore in reference at least to the latter stocks the problem is not an all or none principle. If these tests of general intelligence are at all significant it is evident at once that there is considerable overlapping in the various groups. For the future of this country a careful selection of the best in all the European stocks might be thought desirable. However, it is a common assumption of breeders and eugenists that for the production of a stable, homogeneous, strong stock isolation, group inbreeding and some occasional, but not too great, influx of exotic strains is necessary.??> The incoming of even somewhat distinct racial stocks might be profitable if of a high average intelligence and wide variability. What we want, then, in brief is such a selection of Huropean peoples that they will add variety to our population but not lower its intelligence. We have, of course, the comparable problem of pre- venting the continuance of inferior lines in the present population in this country without adding any more congenital liabilities to our people. In conclusion the writer wishes to submit a few comments upon the practical phases of immigration problems which he believes grow out of these partial but significant investigations of mentality In immigrant stocks. (1.) There must be a change in public opinion as to the de- sirability of large numbers of immigrants. We acquired the habit previous to 1914 of pointing proudly each year to the rising influx of foreigners into our land. The economic exploitation of cheap, unintelligent labor from abroad has fastened a serious racial as well as social-economic problem upon us. It has resulted in con- siderable ethnie displacement. The ‘‘Older Immigration’’ has con- 22 Supra, p. 2—ref. Retzius, ete. *3 Cf, East and Jones, Inbreeding and Outbreeding, p. 264. INTELLIGENCE TESTS 433 stantly lost ground in the face of the ‘‘New.’’ The public opinion of this country needs arousing to opposition to the policy of eco- nomic stimulation of immigration to this country for the profits of the few at the expense of the general well-being. (2.) This means that immigration should be controlled in the interests of the national welfare. The present law is inadequate, unjust and ill-administered. To base the percentages of admission on the number of immigrants of various nationalities in this coun- try in 1910 is merely to continue relatively the same evils as here- tofore. At present there seems to be some evidence that desirable people from Northern Europe and certain English colonies are being kept out while the ratio for Italians and Southern and Southeastern Europeans is large because of the ‘‘high’’ tides of immigration which they reached previously. Moreover, a literacy test is not an adequate criterion of the kind of persons we want. Due to inequality of educational opportunities in Europe, it is likely that persons of good intelligence may be barred because they can not read or write while persons really subnormal, but possessed of a modicum of education are able to pass the meagre demands on their reading and writing abilities at the gates of the country. Davenport has suggested a study of family strains in Europe, but this is expensive and impractical of execution. The writer believes a set of well-worked out physical and psy- chological tests applied to all applicants for entrance into this country would assist in rejecting those whom we do not want. While the psychological tests are not perfect, they are far superior to any other scientific means which are available. The remarkable success of mental testing in our military organization during the war should dispel serious doubts as to the practical value of psycho- logical tests for determining intelligence rankings and from that predicting success at tasks such as modern complex society de- mands. It would well repay our government to spend a consider- able sum if necessary (say half the cost of one battleship) to de- vise under expert advice a set of tests to fit the needs of the im- migration bureau. With adequate time at their disposal a body of experts could arrange a battery of tests, with norms, superior to anything yet at hand. Tests could be devised which would take into account the different linguistic and cultural backgrounds of the applicants, and further the non-verbal test offers possibility of expansion that is still unknown. On the basis of such tests, legal enactment could determine the standards to be set. In addition, if necessary, some differential standards on absolute numbers ad- missible any year could be laid down. It seems to me that there is not a better piece of service for the National Research Council than Vol. XV.—28. 434 THE SCIENTIFIC MONTHLY an attack upon this problem with an effort to secure the national government’s support and adoption. True, there remains after such a program, if it is ever accepted, the entire matter, noted already, of the inferior strains in the pop- ulation now present in our country. Were we to set out on a sen- sible program regarding the immigrant, we should be led ulti- mately into another analogous one concerning the inferior stocks already extant in our population. Linking up these two programs with a sane educational policy we might look forward to a true national greatness. For who doubts that the contributors to a high culture must be a high-minded race? On the other hand, if we continue our present muddling, ir- rational, hit-and-miss method of dealing with these two related problems and the country becomes more and more inundated with inferior stocks, the questions of American citizenship, education and cultural progress will be increasingly difficult of solution. In fact, ultimately such problems as we now see them will be sub- merged in the low standards of life and culture which arise out of a lessened average intelligence in the general population. The picture may be pessimistic. It should arouse us to action. To those who are pragmatists and meliorists the matter is not hopeless, except in so far as the individuals and communities in- volved fail to recognize the gravity of the matter and naively close their eyes to the realities. Blind leaders of the blind, we may well shake our heads in serious meditation. But if the public conscious- ness of the country, under good leadership, realizes the problem soon enough there is no reason why we should not successfully solve this issue and assure our country’s place of leadership in the world. CONCEPTUAL THINKING 435 CONCEPTUAL THINKING’ UNIVERSITY OF PITTSBURGH By Dr. WALTER LIBBY N the recent newspaper controversy concerning evolution, one of Mr. Bryan’s supporters displayed some impatience at the emphasis placed by scientists on resemblances. He protested in the name of logic against what seemed to him an undue insistence on resemblance, resemblance, resemblance! But unless we take account of the similarity among phenomena, how are we to ar- range and classify the data of physics, chemistry, botany and zoology, or arrive at the concepts of which the propositions and syllogisms of the logician are composed? We can have no science of distinct existences ununited by the bond of likeness. It is only by virtue of resemblances that we are enabled to pass from the observation of particulars to the consideration of universals. Bain and other psychologists are so far from belittling the ability to discover the bond of similarity among phenomena, often apparently unlike, that they regard it as characteristic of the man of genius. For James, genius is the possession of similar association to an extreme degree. To the type of genius that notices the identity underlying cognate thoughts belong the men of science, and it is in the concept that the conscious identification takes place. Conception, or the cognition of the universal aspects of phe- nomena, can be illustrated from the history of the biological sciences. For example, what Linnaeus ealled the ‘‘System of Nature’’ was in reality a system of concepts. His eclassifi- cation of plants, though it prepared the way for more natural classifications, was crude, because based on superficial similarities. He, as Harvey-Gibson says, ‘‘elaborated a complex and beautifully arranged and catalogued set of pigeon-holes and forced the facts that Nature presented to him into these pigeon-holes, whether they fitted the receptacles or not.’’ His zoological concepts were like- wise inadequate. Suffice it to recall his vague use of the terms Insecta, Vermes and Chaos. The likenesses revealed in animal structure by the comparative anatomists from Hunter to Cuvier and Owen led to a sharper definition of concepts and a more satis- factory classification. Paleontology afforded new materials for 1 The second lecture in a series entitled ‘‘The Psychology and Logie of Research’’ given before the Industrial Fellows of the Mellon Institute, Feb- ruary 14 to May 2, 1922. 436 THE SCIENTIFIC MONTHLY comparison. Lamarck introduced the term Invertebrate. Embry- ology and the use of the microscope led to fresh observations of likeness and to the establishment of the natural affinities of species and genera. Here might be mentioned particularly the discovery of the notochord—the key, as it has been ealled, of vertebrate anatomy—and the consequent use of the term Chordata. The coming of evolution made possible a phylogenetic classification of organisms and a subtler differentiation of biological concepts. The dominance of conceptual thinking in the classificatory aspects of science is fairly obvious. For example, in adopting the terms Quercus alba, Q. rubra and Q. salicifolia, the seventeenth- century taxonomist merely associated a definite nomenclature with the distinct concepts of Virginian woodmen, who had observed the likeness of American and British oaks. In other aspects of science, where the importance of the concept is far less obvious, it is none the less real. In the study of human anatomy by means of dis- section it is generalized or conceptual knowledge that one seeks and retains. Anomalous or exceptional structures—such as a triceps muscle in place of a biceps, or an over-developed panniculus carnosus—are either disregarded and forgotten, or remembered as anomalies and exceptions. In any ease, what the student retains, after two years spent in the dissecting-room, is a generalized knowl- edge of the structure of the human body and not a memory of the particular cadavers that seemed to occupy the focus of his atten- tion. The case is somewhat similar with your natural history mu- seum. The biological collections consist wholly of dead symbols of living things. Bones are frequently represented by masses of limestone or silica. A single specimen may do duty for a species, genus or order. The mere external surface—the shape—may alone be preserved. Only one stage in an animal’s development may be exhibited, or only one posture—the Megatherium pulling down a tree or a dinosaur in a more or less characteristic pose. Hach speci- men has value not as representing an individual but as symboliz- ing a group. Hach concrete object, like a word in a catalogue, serves to recall a concept. The importance of cultivating the habit of conceptual think- ing has been definitely recognized since the time of Plato, and Plato’s master, Socrates. The majority of people, according to the Platonic dialogues, are the slaves of their senses and never attain to a system of clear concepts. They fail to translate their perceptions into conceptions and to pass from the sensible world to the supersensible or intelligible world. Plato valued the sciences with which he was best acquainted—mathematies and astronomy— because they tend to wean the mind from what is sensory and transitory to what is conceptual and eternal. The concept triangle, CONCEPTUAL THINKING 437 the triangle of definition, is more real than any of its visible repre- sentations. Discipline in conceptual thinking is perferable to utility. ‘‘You amuse me,’’ he writes, ‘‘by your evident alarm lest the multitude should think that you insist upon useless studies. Yet, indeeed, it is no easy matter, but, on the contrary, a very difficult one, to believe that in the midst of these studies an organ of our minds is being purged from the blindness, and quickened from the deadness, occasioned by other pursuits—an organ whose preservation is of more importance than a thousand eyes; because only by it can truth be seen.’’ Plato’s philosophy is interesting for us because modern science is Just such a system of clear concepts as he had in mind, and it is a mistake to interpret his intelligible world as something invisible to-day but accessible to the senses at some future time. The scien- tist must turn away from the sensuous world of the artist and the child to the intelligible world of mathematics, physics and biology ; from the eight or seven little boys seated on a fence to cube root and prime numbers; from the panorama of many-colored nature to the conceptual world of elements, atoms, ions and electrons. The sciences are a sort of shorthand way of conceiving phenomena. They do not give us nature in its richness and fullness. In the words of Mephistopheles, ‘‘Gray, dear friend, is all theory, and green the golden tree of life.’’ Picturesqueness is sacrificed by the scientist for the sake of clearness and economy of thought. He easts his net now for one kind of fish, now for another. For an artist like Monet, water may be a shimmering variegated surface ; for the child it may mean ‘‘to drink;’’ while for the chemist it may be H,O, or, if he conceives it as an aleohol, H.OH. Of course science has no monopoly of clear conceptions. In in- sisting on the value of an education in conceptual thinking, Plato had in mind the training of leaders in ethics and statesmanship, and thus anticipated by a few centuries the publicists and phil- osophers who to-day advocate, as a novelty, the application of the intellect in social and political reform or proclaim the cultivation of the scientific habit of mind as the sole means of maintaining and advancing contemporary civilization. He would have recognized Lincoln as a man of clear conceptions, gained through a unique self-education, by living in close contact with man and nature, by reading a few books with extraordinary care, by poring over the statutes of Indiana, studying grammar, arithmetic and sur- veying, conning the dictionary, passing in review the life of Wash- ington and the history of the United States, following with a frontiersman’s imagination the exploits of Robinson Crusoe, ab- sorbing the wisdom of Aesop’s Fables, weighing the moral prin- ciples of The Bible and The Pilgrim’s Progress. Indeed, 438 THE SCIENTIFIC MONTHLY clear conceptual thinking, the scientific habit of mind, was con- sciously cultivated by the moral philosophers of Greece, notably by Socrates, whose chief distinction is that he subjected to critical examination such concepts as virtue, temperance and justice. Speaking of this pioneer work, Stout says: ‘‘It is only at a late stage of mental development that an attempt is made to distinguish an identical or persistent element of meaning pervading the vary- ing significations of a word. When the attempt is made, it con- stitutes an epoch in the history of thought. It is the beginning of definition and of the scientific concept.’’ Conception, or thinking the same in like circumstances, has its complement in discrimination. Association by similarity is offset by dissociation, integration by disintergration, synthesis by analysis, and the observation of congruity by the observation of incongruity. In this respect, there is a marked difference between one individual and another. Experiment shows that one student may recognize readily ten shades of gray where his classmate has the greatest difficulty in discerning any difference in shade what- ever. Similarly, one mind is alive to shades of meaning that make no impression on another. Aristotle, the greatest of all scientific intellects, trained for twenty years in the school of Plato in con- ceptual thinking, the most acute among the Greeks, as he has been ealled, in noting differences and making distinctions, carried his investigations into almost all the realms of knowledge. His suc- cess was most marked in fields where his aptitude in the employ- ment of general concepts was supported by a wealth of observa- tional data. This is particularly remarkable in his researches in biology. In the Historia Animaliwm, for example, one discovers the trained thinker bringing order out of chaos by the application of the intellect to the facts of experience, so called. It is the cus- tom among historians of a conservative type to belittle the medieval followers of Aristotle, and above all the scholastics. But even to scholasticism modern science is deeply indebted for the develop- ment of logic in general, and for the definition and differentiation of concepts in particular. The use of language is the indispensable concomitant of clear conceptual thinking. We can not think of one of the lower ani- mals advancing very far in logical thought. He is bound down for the most part to the fleeting images of things, and lacks the word (logos) by which they might be made permanent and inde- pendent of the continuum of experience. The intellectual develop- ment of the child proceeds as a rule pari passu with its command of appropriate terms. ‘‘Out of hundreds of English-speaking children,’’ says Terman, ‘‘we have not found one testing signifi- cantly above age who had a significantly low vocabulary; CONCEPTUAL THINKING 439 and, correspondingly, those who test much below age never have a high vocabulary. Occasionally, however, a subject tests somewhat higher or lower in vocabulary than the mental age would lead us to expect. This is often the case with dull children in cultured homes and with very intelligent children whose home environment has not stimulated language develop- ment. But even in these cases we are not seriously misled, for the dull child of fortunate home surroundings shows his dullness in the quality of his definitions, if not in their quantity; while the bright child of illiterate parents shows his intelligence in the aptness and accuracy of his definitions.’’ Terman thus makes it clear that intelligence is not to be gauged by the extent of one’s vocabulary, but by the exactness with which concepts are defined. In the interests of the progress of both science and democracy, it is important that training in the precise use of words—especially the derivatives of those languages from which we draw our general eoncepts—should not lag behind other conditions of the develop- ment of the immature. As Walter Lippmann says, ‘‘ Education that shall make men masters of their vocabulary is one of the central interests of liberty.’’ Franklin’s success, both as a states- man and a scientist, was in no small measure owing to the severe drill in the use of the English language to which he subjected himself. Two other self-educated research men, namely, John Hunter and Michael Faraday, who, on first thoughts, might seem to bear witness against the view that language training is of im- portance in scientific investigation, prove on examination to fur- nish testimony in corroboration. John Hunter received little if any schooling. He turned in contempt from the opportunity of studying the classics at Oxford. Although, after reaching ma- turity, he was brought, through the closest association with his brother, William Hunter, in contact with scholarly traditions, he never overcame the defects of his early education. We are in- debted to him for such concepts as ‘‘arrested development’’ and ‘secondary sexual characters,’’ but his pages are strewn with terms like ‘‘the stimulus of death,’’ ‘‘the stimulus of imperfec- tion,’’ and ‘‘sympathy,’’ to which he assigned a significance now impossible to recover. The lack of language training, in spite of Hunter’s genius and vivid personality, was detrimental to his influence as a lecturer and writer. Owing to this shortcoming, science has not yet reaped the full harvest of his tireless energy in research. For example, the recapitulation theory, as stated by him, seems to-day little more than a literary curiosity, though it may have influenced the progress of embryology through the inter- pretation of the younger Meckel. The case of Faraday was some- what different from that of Hunter. Having received an elemen- 440 THE SCIENTIFIC MONTHLY tary education, Faraday became apprenticed to a book-binder. For years he spent his leisure time in reading scientific works. At the age of twenty-one he gained the favor of Sir Humphry Davy | by a lucid report of some lectures delivered by Davy at the Royal Institution. Faraday became Davy’s assistant, traveled on the Continent with his patron, studied foreign languages, and made definite efforts to acquire the oratorical arts of Davy, a recognized master of scientific diction. Faraday’s opportunities for language training, however, came just a little too late. He sometimes con- fessed his difficulty in formulating the ideas that occurred to him. He sought aid at the University of Cambridge and was indebted to Whewell for such terms as ‘‘electrolysis,’’ ‘‘electrolyte,’’ ‘‘ion,’’ ete. Language permits us to summarize nature, to express it schematically, to seize upon certain aspects of it—that is, to analyze phenomena with certain purposes in view. For Priestley the part of the atmosphere that supports life was ‘‘pure dephlogisti- eated air.’’ Lavoisier substituted a new term and a new concep- tion, viz., ‘‘oxygen.’’ Davy spent a great deal of time proving that Lavoisier had a false conception of the element discovered by Priestley. We retain the name after having modified the con- cept. This we do with the greater freedom, seeing that the classical term ‘‘oxygen’’ is not self-explanatory, as is the analogous term ‘“Sauerstoff.’’ Gases were known in the last quarter of the eight- eenth century as ‘‘kinds of air,’’ or ‘‘factitious airs.’’ As late as 1766, Cavendish ealled hydrogen ‘‘inflammable air.’’ In 1783 and 1785, he made experiments that justify the conceptions ex- pressed by the terms ‘‘hydrogen’’ and ‘‘nitrogen.’’ It was almost impossible to think clearly concerning earth, air, fire, and water, the so-called elements, without having the terms ‘‘oxygen,’’ ‘‘nitro- gen,’’ ‘‘hydrogen,’’ ete., as symbols of the concepts corresponding. Counting, measuring, weighing—the application of mathe- maties—must be regarded as among the best means of sharpening up our conceptual thinking. One classical example is Lavoisier’s use of the balance in establishing the nature of combustion and giving phlogiston the quietus. ‘‘About a week ago,’’ he wrote on November 1, 1772, ‘‘I discovered that sulphur in burning, so far from losing weight, rather gains it; that is to say, that from a pound of sulphur more than a pound of vitriolic acid may be ob- tained, allowance being made for the moisture of the air. It is the same in the ease of phosphorus. The gain in weight comes from the prodigious quantity of air which is fixed during the com- bustion and combines with the vapors. This discovery, which I have confirmed by experiments that seem to be decisive, has made me believe that what is observed in the combustion of sulphur and CONCEPTUAL THINKING 441 phosphorus may equally well take place in the case of all those bodies which gain weight on combustion or calcination. I am per- suaded that the gain in weight of the metallic caleces is owing to the same cause.’’ Lavoisier followed up this work by the caleina- tion of tin in 1774, and in the same year—after Priestley’s dis- covery of ‘‘pure dephlogisticated air’’—by the oxidation of mer- eury. In 1777, Lavoisier stated: that in all cases of combustion heat and light are evolved; that bodies burn only in oxygen (or air éminement pur, as he at that time ealled it) ; that oxygen is used up by the combustion, and the gain in weight of the substance burned is equal to the loss of weight sustained by the air. The differentiation of terms and concepts is so necessary an accompaniment of the advance of science that no collection of examples can be regarded as adequate or as even fairly repre- sentative. Though Lavoisier in 1777 succeeded in giving to the concept ‘‘combustion’’ a much more clearly defined meaning than had attached to the ‘‘fire’’ of the ancient philosophers or the ‘*flame’’ of Francis Bacon, in 1789 he still included ‘‘ecaloric’”’ and ‘‘light’’ in his table of elements. In spite of the definition by Robert Boyle of the concept ‘‘element,’’ and the attempt of New- ton to determine the meaning of ‘‘atom,’’ these ideas, inherited from the remote past, were at the close of the eighteenth century about to enter on a new series of transformations. In the seven- teenth century Boyle’s contemporary, John Ray, ascribed to the term ‘‘species’’ a definite, if not a final, significance, and Syden- ham, seeking to establish by clinical observation distinct species of disease, succeeded in differentiating measles from smallpox, in defining chorea, in modifying the significance of the term ‘‘hysteria,’’ ete. Progress in science may involve lessening or in- creasing the extension of a familiar term, determining anew the distinction between familiar terms, and introducing new clearly defined terms. Pasteur’s studies in molecular asymmetry involved a reconsideration of the terms ‘‘tartrate’’ and ‘‘racemate’’ and a delimitation of the concepts which each of these terms expressed. An advantage is gained by substituting the unfamiliar ‘‘neuras- thenia’’ for the familiar ‘‘neryousness,’’ partly because the new term is unambiguous and partly because it is devoid of every popu- lar connotation. In fact, our scientific terminology has become so much a thing apart that one may overlook the relationship be- tween a common term like ‘‘weight’’ and a more technical term like ‘‘mass.’’ The researches of Schleiden and Schwann, which led up to the statement of the cell theory, were affected and, to some extent, vitiated by traditional conceptions concerning ‘‘cellular tissue’’ and the ‘‘cell.’’ Robert Hooke was the first to use the term ‘‘cell’’ 442 THE SCIENTIFIC MONTHLY in describing organic structure. He had examined charcoal, cork, and other vegetable tissues under the microscope and described them in 1665 as ‘‘all perforated and porous, much like a honey- comb.’’ He could discover no passages between the minute cavities or cells, though he took it for granted that the nutritive juices to be seen in the cells of green vegetables had some means of egress. Hooke’s observations were verified by his contemporaries. Grew, in describing the microscopic structure of plants, mentioned the infinite mass of ‘‘little cells or bladders’’ of which certain parts are composed, and Malpighi described the cuticle of the plant stem as consisting of ‘‘utricles’’ arranged horizontally. Caspar Wolff in his doctor’s thesis (Theoria Generations, 1759) reported the observation of cells and ‘‘little bubbles’’ which developed in the homogeneous layers of the embryo. In the works of Bichat, the founder of histology, the term ‘‘cellular tissue’’ was used, as in- deed it is to-day, to indicate a certain kind of connective tissue. Treviranus and Link described the cells in vegetable tissues in 1804, the latter maintaining that they are closed vesicles incapable of communicating with each other. Professor John H. Gerould has recently pointed out, in the pages of The Scientific Monthly, the important part taken by Lamarck, Mirbel (the disciple of Caspar Wolff), and others in the development of the conception of the ‘‘cell’’ and of ‘‘cellular tissue.’’ After the appearance of Moldenhawer’s Contributions to the Anatomy of Plants (1812), which demonstrated that the cavities of vegetable cells are separated from each other by two walls, the attention of observers was di- verted from the cell contents to the cell wall. The consequent misconception of the nature of the cell was in part corrected by Robert Brown’s discovery of the cell nucleus and by the later dis- covery of protoplasm. It was before the full significance of the eell contents was realized that the cell theory was conceived by Schleiden and Schwann. It is evident that advances in scientific thinking imply the use of clear concepts and clear terms. The term ‘‘neuron,’’ employed by the early Greeks in the sense of ‘‘thong’’ or ‘‘sinew,’’ was ap- plied by the anatomists of the fourth century B. C. to the tendon as well as to the nerve. A considerable treatise alone would suffice to trace its subsequent meanings and those of its derivatives and at the same time to give an account of the investigations that from the time of Herophilus and Erasistratus have contributed to the elucidation of the concepts in question. The terms that represent to-day the so-called chemical elements have no doubt undergone a similar series of transformations in meaning. Distillation, erystalli- zation, and other refining processes had to be brought into play before the concept—the spirit, the essence, the thing in itself— could be realized. “WHO’S WHO” AMONG AMERICAN WOMEN 443 “WHO’S WHO” AMONG AMERICAN WOMEN By Professor STEPHEN S. VISHER and GERTRUDE HOVERSTOCK INDIANA UNIVERSITY, BLOOMINGTON ATTELL has made some stimulating statistical studies of the more eminent scientific men, including the distribution of their birthplaces and of their present residence.t Some years ago a study of the distribution of the first ten thousand persons in ‘“Who’s Who in America’’ appeared in this journal.? The conclu- sions drawn from these studies, while not to be seriously doubted, are so interesting that it appears worth while to test them by a similar study of a different group of notable people— the 1,687 women included in the last edition cf ‘‘Who’s Who in America’’ (Vol. XI, 1920-21), especially the 1,582 women concern- ing whom biographical data are given. The distribution of the place of birth of 1,551 women who gave this information is indicated by districts in Table 1, as is also the ratio between eminent women and the general population of 1880. TABLE 1 BIRTHPLACES OF WOMEN IN ‘‘WHO’s WHO IN AMERICA’’ Number per 100,000 at District Number Per cent. 1880 census JeMiSy f= LOH vd Fe 6 ee ace pecs 333 21.5 8.3 Middle) -Atlantic...:.2<-.224..2..000 eee 511 33.0 4.3 Mast North Central... 22.2.2 222 4 ee 347 22.4 3.1 Wiest North: ‘Central... ees 99 6.2 0.5 OTLEY ae tt ee tl ee 101 6.5 1.6 — Mountain and: Pacific: ....25--2.-0o ee 49 3.2 2.8 IGEGUPTE . COMNETIOS a aasanncere ee 112 7.2 a This table reveals the prominent share New England has had in the production of eminent women, and the small share which the southern and western halves of the nation have had. ‘‘Who’s 1J. McKeen Cattell: ‘‘Families of American Men of Science.’’? The Popular Science Monthly, May, 1915, and Tue Screntiric MonrTHLy, October, 1917 (reprinted in ‘‘ American Men of Science,’’ third edition, 1921). An earlier study based on the starred scientists in the first edition is reprinted in the second edition of ‘‘ American Men of Science,’’ 1910. 2 Scott Nearing: ‘‘The Geographical Distribution of American Genius,’’ The Popular Science Monthly, Vol. 85, pp. 189-199, 1914. 444 THE SCIENTIFIC MONTHLY Who in America’’ is published in Chicago and is edited by an Ohioan. Of scarcely less interest than the variation among the districts’ is the variation among the individual states in the number of famous women they have produced. Table two shows for each of the leading six states the number and proportion of eminent women. TABLE 2 Six LEADING STATES IN THE PRODUCTION oF EMINENT WOMEN Per cent. Ratio per of total 100,000 of Number now eminent general popula- residing in State Native women tion in 1880 the state New) Monks se suey An il 291 19.0 5.8 550 Massachusetts .............-...--. 171 11.2 9.5 237 TO: COA DUES) 117 7.6 Safi 46 Pennsylvamia — ......2..1--4..----- 113 7.4 2.7 90 HD eKa wk yag ante beehe N ULE Ak ie Una 90 5.8 3.0 107 JO oko beens yy MmAe LU Leh UE eat 43 2.8 2.1 20 IMGimn Gs Ota hare ee ees Le 43 2.8 6.6 24 Table three gives the number of eminent women born in each state and the number now living there. TABLE 3 BIRTHPLACE AND RESIDENCE OF EMINENT WOMEN BY STATES State Native Resident VAST ATED UTIN EU een ee Ee vee aera 19 10 PN Ay 0% Ys) US EN LeU e a e 0 1 VeNo oho 0 (SEEN AtOY 2 tae aU IRE LENA 5 2 Galli orm ayer see hele 8 UL A 28 78 Wolo ste ae RIE Le ee eee 0 14 Cf chal ea kT Fie Ne RPA aioe Me GRAN AC SL YOU 54 62 TDL even Te yee ee ee CTA A Oa 5 i EN Koy ea Webi) Vee AU A OOS 3 6 GOT Oey ie ee ee 9 13 LECH ahh eae hee a a UI A RR HRMS Pa IOS TA 1 1 OM ayy Me SENOS AERA ET SOV TELE 90 107 Mei cnn ciuaeesee eee UY NORTE SNOPES Wh RAED 43 20 CIFOSUVIEL NR re oem Coe eo CL MPM 24 11 TERCERA epee ET AN ae BS 15 9 rom inc isya geet tk aN rane Oe 32 22 faye bE Wee H ae | yeu ee Raa PR ee ONE AY yet te 14 2 Tea ie ee 2 Lg 43 21 ANTEER gh 1 be EAR a HC SR 28 80 IMPARSA CHISTES Ny cscee clos uhs UMN Meee 171 237 UVB Fee: 7 NN MUR 8 POE oP atl Ae 33 17 IVETE SOEs Fee ch ART TU eey 18 24 D4 GESFSAESI EH 0) 0} bi) Maa CRs CUNEO A ea LEE 10 2 NU Dokst= {040 ftp) y SS UR eRe ER EL UE ed 36 25 BA's roy enysezly ok: i) Rac ese eae Mee a Lee aM TS DB 2 INFO TS Sai eee eT Ue 5 3 “WHO'S WHO” AMONG AMERICAN WOMEN 445 INIGK ENG ESAS ao ober eo MUU Ui Ede D Ye ae 2 2 New? Elampshire 2s eee ao eueteeeeoe 27 14 ING) WOR SOs? cso AD oe 33 58 UNTerenit Wee O20. uot eee OLE LR aN RN ql 2 ING Wi ip WOT oe. SURAT aga ea aa 291 550 BNowtbyan @aro lim aye ee aera i Mee ff 11 NOTH MD AKO ba LA eI eR a 2 2 ( O}LATCG) NH Cus AIAG Rca Nan Eat Re TT 7, 46 (OSIM aN(a ja FA pear OM MRD Aram aie ON 1 1 CO hefcf (0) 0 aan A Bee LUAU OY 4 7 IRONS LV ANT A, | 22ctke CUE a eee 113 90 bod ete lslan Gis hs a One 16 11 South Carolina ee UN ae, 4 3 South Dakota) Wee es if 0 WENN ESSE (o/s. 2.221 Se a NE ee 20 10 MIN OSA patie eT Sle UNC aU 6 5 [Utrera 2 2S Lee 3 3 NVIG@EINVONIE iy Bees oe a sace ee ee a cea 22 3 AU(itcfea bits A piectimereenyeer epee MeN eer 35 16 PiU COM) encase A aes ag ae of 12 VG S bry VAI PEIN ea ea 2 We cee a die bee 6 5 IWS COTS LMNs |) nse 02a LEAN MD AI an 32 15 IWVay OTe NCU aa ee ee 0 3 Countries! outside Wey See ee eens 112 48 INOG) Movie mes Qh a ME Ate A 136 Motegi ees 2 a eee ee ee 1,687 1,687 This table indicates that Colorado, Wyoming and Arizona, having a population of 260,000 in 1880, produced no women who have been included in this issue of ‘‘Who’s Who in Amerieca.’’ Furthermore, Idaho, South Dakota, New Mexico and Oklahoma, having a total population of 451,000 in 1880, have each produced only one. Thus these seven states with a population in 1880 of 712,000 are represented in ‘‘Who’s Who’’ by only four women. On the other hand, New England, which had a searcely larger popu- lation in 1880, contributed 333 eminent women. Similarly, the Southern States, with an 1880 population somewhat greater than that of New England and the Middle Atlantic States combined, produced only 99 famous women in contrast with 844 from these northern states. Not only were few eminent women born in the South about 1880, but still fewer now reside there, 63 vs. 99. The North Atlantic and New England States have attracted many no- table women with the result that 1,148 of the 1,687 women listed now live there. In other words, these states produced 54.5 per cent. of the eminent women, but now have 68 per cent. of the nation’s total. This great centralization of production of famous women and of their present distribution may be due to the following in- 446 THE SCIENTIFIC MONTHLY fluences: The presence of more educational institutions in the northeast, and the greater emphasis placed on education there. Unquestionably there are sectional differences in ideals. In parts of the South, for example, an intellectually ambitious woman is not in favor. The fact that men outnumber women in the West tends to encourage early marriage in the West. Relatively sparse popula- tion and more recent occupation also tend to cause life to be on a somewhat more primitive plane, with less opportunity or incentive for the type of achievement recognized by inclusion in ‘‘Who’s Who.’’ Western women who do not marry early are more likely than eastern women to have opportunity to become school teachers, clerks or business women. Selective emigation certainly helps explain the distribution of the birthplaces of the eminent. As a rule, the highly intellectual type do not become frontiersmen. Pioneering ealls for physical vigor and daring rather than high education or unusual intelli- gence. Furthermore, the highly intelligent type generaily are in fair circumstances, and it usually is the poor who emigrate, not the well-to-do. Hence, for a number of reasons, there is a ten- dency for the intellectual type of people not to emigrate, unless it be as missionaries, but to remain where they can make the most use of their ability and education. Thus many have remained in the older states, or have moved into the older communities in the Middle West, rather than going to the Newer West. Consequent- ly, few infants possessing unusual intellectual endowment are born on the frontier or in the Newer West. The presence of nearly ten million negroes in the South reduces the South’s contribution of eminent women in proportion to its total population, for no negress is included in this volume of ‘“Who’s Who.”’ The climate of the North is more favorable for mental activity than is the often rather enervating climate of the South. It like- wise favors physical vigor and thus increases accomplishment. Climate also doubtless has played a part in reducing the produc- tion of eminent women in parts of the West. While the climate of the arid and semi-arid west may possibly favor intellectual activity and physical vigor, it can not be disputed that the fre- quent droughts, unseasonable frosts, etc., have tended powerfully to encourage the emigrating of the exceptionally alert and re- sourceful people. Such people tend to go into regions where the opportunities are less uncertain. In addition to place of birth and residence, note was taken of occupation, education and state of marriage. It was found that “WHO’S WHO” AMONG AMERICAN WOMEN 447 53 per cent. of the last 950 women in ‘‘Who’s Who”’ are married. Many occupations are followed. Eighteen chief types were listed. The most important eight, with the number engaged in each, and the per cent. of the total are shown in Table 4. TABLE 4 OCCUPATIONS OF WOMEN IN ‘‘WHO’s WHO IN AMERICA’’ Occupations Number of women Per cent. of women AWiritOls 573.2. 2-2 33 de Sa eee | 714 45.3 EEL tL OI 2a posses kt nen ee 244 15.6 Social, workers) =..2.- so) ee 127 8.0 JF) ChE Ae MA a ree ALN ik. tos ht ADM lily 7.5 ICEL CSSOS) (oie. ee eu £2 Mea one 63 3.9 SILC) ee ae AO. Ik SSIES 46 2.9 PEGG ONS) eee? be.2 Wes Sb en aD 34 2.1 PIV SICI ATS, | rec. ck te ee eee 28 1.7 In addition to these eight, there are lawyers, politicians, religious workers, librarians, scientists, lecturers, explorers, musicians, busi- ness women and those interested in home economics. Thus there is a very wide range of activities. From Table 4 and other data a few conclusions appear war- ranted. One is that women receive recognition for writing more readily than in most activities. Nearly one half of the 1,582 women whose biographies are given are writers. On the other hand, comparatively few teachers have attained the fame of the type indicated by inclusion in ‘‘Who’s Who,’’ most of the ‘‘edu- cators’’ included being administrators such as deans and presi- dents. Indeed, a considerable number of women are holding ad- ministrative positions. The higher education of women was also noted, and it was found that 88 per cent. report training above the secondary school. Of the two groups attending college or college and university, exactly one half of the women report training in women’s colleges. TABLE 5 Type of training Number of women Per cent. of women (Oa Weve) Fase pe Bo eS eo oor 440 27.9 AUUREV GPS)! foo 2 prea eee eee wiles 216 13.7 S| LTTE) ERR a oe ena chap ice Soe Soe 466 29.5 College and university... 2._.2.-2.-- 205 13.0 Collegevand special... ee 39 2.4 University and specials... 200-22. o-oo. 23 1.5 NOME MMENELONC Cte ccaterrsenns noeen see meee 190 12.0 This table indicates that few women who do not take advan- tage of existing opportunities for higher education (beyond the secondary or high school) now attain national fame. 448 THE SCIENTIFIC MONTHLY GO TO THE BEE A CONTRIBUTION TO THE PERMANENCE OF ARISTOCRACY By Dr. DAVID STARR JORDAN STANFORD UNIVERSITY 667 FYHE Leisure Classes, the Chief Support of the Nation they adorn’’—such was the topic of a remarkable sermon de- livered not long ago to an audience of superior people, by the Reverend Vicar of Girlington, England, whose actual name I do not give lest my feeble words fail to interpret his lofty ideals. From the press notices that came under my eye, it appears that the admi:ed and admirable vicar finds his mission in the salvation of British aristocracy through its complete restoration to the ranks of leisure. In his judgment the aristocrats or superior persons serve society best by standing as examples of human perfectibility. This is the end they should seek, through utter surcease from all worry, all effort and all personal hopes and desires. The vicar would indeed make of the upper classes a group, not of hereditary rulers, but of elect examplars of what humanity may become, a condition to be open to a chosen few to whom is granted release from the sordid side of life. From such relief the great body of the British people are of course excluded—not from any fault or deficiency of their own, but simply because there is not leisure enough to go around. Thus for the chosen the mass must live; the many gather honey for the few to enjoy. But in fair- ness—and this I take it is the vital part of the vicar’s contention— all should have an equal chance in the beginning. A rationally organized society would then consist of two classes which for convenience the vicar might call the laborers and the leisurers. To the former belongs the capitalist as well as the workman; all indeed who work with hand or tongue or pen or brain labor alike. The leisurer alone enjoys that perfect serenity which comes from fearing nothing, wanting nothing, hoping for nothing. True happiness rests on a division of duty. A natural cleavage lies between those who create and those who enjoy, each condition having its own peculiar delight. Between the two yawns a great gulf which society crosses only at its peril. The vicar’s discourse harks back to the words of Solomon, ‘‘Go to the ant; consider her ways and be wise,’’ an injunction inciting alike to modesty and to thrift, two virtues of which the ant is a GO TO THE BEE 449 model. She knows her place and keeps it. Aspiration goes with aviation ; having no wings, she never tries to fly. But in addressing the leisurer the vicar would modify his text: ‘““Go rather to the bee; consider his ways and be wise.’’ Physically the bee resembles the ant, but his social system is organized on a more exalted basis. With him, the leisure class is unquestionably the chief support of the society it adorns. And now to make clear the vicar’s appeal, I find it necessary to amplify the too meager report given in the Girlington Guardian, and without holding the speaker closely responsible, I may draw for the moment from the fascinating observations of the noted apiarist of Brussels, Maurice Maeterlinck. Two salient facts at once appear: first, bee society maintains its own aristocracy ; second, its leisurers, having no hereditary claim for distinction, are chosen by lot and by no effort of their own. As the young bees are about to hatch, their faithful nurses construet a few cells of extra size and feed the occupants on a special food, the “‘royal jelly’’ of the apiarist. These selected individuals, the ‘““queens,’’ then grow up in an atmosphere of leisure. To produce the harmonious and perfect bee for which the toiling workers exist is the culmination of the apiarian system. In carrying the analogy into human society, the essential point (as I think the vicar would agree) is that among men as among bees, no injustice shall be done. Leisurer and laborer must both exist, but as both are of one lineage, each should have an equal chance for the great prize of existence. That ‘‘the rank is but the guinea’s stamp, the man’s the gold’’ is literally true. Royal jelly and a royal cell make the bee-aristocrat. The difference between queen and worker is purely one of bringing up; the two are of the same blood, the queen becomes regal without effort of her own. By like means human society may breed its aristoecrats—so reads the lesson to be learned from the bee. The queen exists not for her own sake, nor by inherited right. Neither should a lord among men, his only true function being to round out the humanity of his fellows, show what man has it in him to be if brought up without work or worry, marred by no trace of struggle, no fear of defeat, by nothing which wrinkles the brow, makes callous the hands, nor hardens the heart! Lacking this perfect ideal, end and aim, humanity can never realize itself. Perfection, by the very nature of things, is denied the laborer. Yet how vital it is that perfection should exist! Plainly, however, only a few can be leisurers, though in life’s grand lottery all may start alike. The needs of the hour eall for the ‘‘man with the hoe,’’ the woman at the washtub. Some must toil and spin that the human lily may be properly arrayed. Mark, too, the perfect rose, Vol. XV.—29. 450 THE SCIENTIFIC MONTHLY ‘‘its own excuse for being,’’ yet dependent on stem and leaves and roots for very existence. Moreover, its glowing petals are but leaves transformed and perfected. Thus from the lily or the rose one may draw the same lesson as from the bee and the ant. In his implied criticism of the British aristocracy of the re- nowned Victorian era, the vicar proceeds with becoming modesty. It would seem indeed presumptuous in the holder of a living, the eift of a gentleman of the county, to say one word in dispraise of conditions as they are. The very essence of organized religion is conservatism. Yet some one must turn his face towards the light, and every student of history knows that the ‘‘Decline of Aristocracy,’’ as defined by one of its illustrious examples, Mr. Arthur Ponsonby, is now imminent. The only way of avoiding decadence is to place aristocracy on a democratic basis, as it were, to open its doors to all, but at the same time keep the passage narrow so that few can enter. ‘‘Strait is the gate, and narrow is the way’’ that leadeth to perfection. The Victorian system is open to eavil on three sides: it is hereditary : it is open to invasion by wealth, it is used as a reward for achievement. As for heredity, one easily sees that mere aristrocratic descent does not guarantee a perfect leisurer. ‘‘Blue blood’’ tends to run thin, and in-and-in breeding plays havoe with the human stock as well as with dogs and horses. Furthermore, inheritance smacks of privilege, and the iconoclastic Georgian Age will have none of it, No youth is now willing to follow his father’s trade. The son of a tailor no longer sits cross-legged on a bench if fate has taken him early in hand, the son of a non-conformist preacher becomes prime minister—and one never knows where it is all going to stop! But any healthy boy, caught young enough, is soon aligned with the customs and opinions of his entourage. Set among gentlemen, noth- ing is easier than to act like a gentleman! As a matter of fact every wise butler soon acquires a manner as lordly as that of his master, whom he often instructs in the higher etiquette. Frequently he has had longer experience, and has given to items of behavior more serious thought. A sympathetic moisture of the eye and a gentle lowering of the voice in delicate moments may come as natural to him as toa bishop. Stephen Leacock of Montreal, a noted American tourist, gives an amusing account of his call upon an ambassador of the United Kingdom. At the end of a discussion concerning the weather and the future of the Empire, he was still doubtful as to whether he had met the ambassador or his butler. To test the matter, therefore, he slipped a gold sovereign into the hand of his host. The coin was accepted, but in such a detached and dignified GO TO THE BEE 451 manner that the visitor’s uncertainty was not dispelled by his maneuver. With a ‘‘young person,’’ the process of adaptation is even more rapid and sure. The charwoman’s daughter brought up as a lady yields to none in ladylike perfection. Napoleon even went so far as to propose that women should have no hereditary rank at all, but content themselves with the title of their consorts. Presumably his experience with the stalwart insistence of his otherwise plebeian sisters whom he had personally ennobled determined his attitude in that regard. But most of us are familiar with the career of the Honorable Lady Burnett, a woman of humble origin as her friends admit—once a milkmaid or (according to traducers) a barmaid, who became through an exalted marriage ‘‘the arbitress of the elegances for all the region about her husband’s manor house.”’ From the Morning Post one learns that ‘‘she used, queen-like, to reign; nay, pour at tea in the newest and tightest of white gloves,’’ and with that undefinable je ne sais quot which marks the true aristocrat ! An aristocracy has been defined as ‘‘a social superstructure reared on a foundation of bestness.’’ There is but one permanent basis for ‘‘bestness.’’ This is implied in the thousand year old motto of the aristocratic Winchester College, ‘‘Manners makyth man,’’ or in the modern vernacular, ‘‘Handsome is as handsome does.”’ The vicar, himself of humble birth but aristocratic connections, admits freely that it is too late in the day to lay stress on lineage. We all claim Adam as a forbear, and not one of us has ever had even a single ancestor weak enough to die in infancy. Leisurer or laborer, each has weathered the storm; in that sense, all are noble alike. Genealogists also affirm that we unmixed English people are all of Plantagenet stock, most of us through the first three Edwards, descended from William, Alfred and Charlemagne. Indeed, one eminent authority classes all Englishmen together as the ‘‘inbred descendants of Charlemagne,’’ and Charlemagne (as we know) was at the head of a League of Nations, being at once King of France, Emperor of Germany, and Overlord of Europe. Noble blood indeed is ours, but unfortunately so many share it that it can not serve as a test of aristocracy. Again, large numbers of our so-called nobility have arisen through long years of struggle directed toward that end. Social extrication is a form of hard work, and labor of whatever kind mars the soul as it wrinkles the brow. A furrowed face is of itself a badge of serfdom. A mind too alert, an ambition too urgent, tends to defeat the purpose of social adornment. To diffuse sweet- ness and light is not an arduous occupation. Indeed to give ‘ 452 THE SCIENTIFIC MONTHLY thought to it or to make it a result of definite effort is to fail in the most important element. The leisurer who goes down into the East End to mend the manners of the poor would do better to confine himself to awarding the conventional goose and bottle of ale at the happy season of Christmas. Those who have pursued this policy enjoy a devout gratitude height- ened by its rarity. If life for the laborer was all ale and goose —or beer and skittles, as vulgarly paraphrased—no one would know his place, and the barriers betwixt laborer and leisurer so carefully built up in the long centuries of England’s glory would be completely broken down. Browning paints the ideal leisurer as a king who ‘‘lived long ago in the morning of the world’’ with a forehead ‘‘calm as a babe new born.’’ This is an enchanting ideal, but the further descrip- tion (so familiar that I need not quote it here) does not fit their lordships of to-day. No one would take it for a portrait of North- eliffe, Carson, or Birkenhead, whose bustling activities keep the realm in turmoil. An aristocracy founded on labor of mind or hand is not a elass of leisurers. Like my Lord Bottlebrush and the ‘‘Duchess of Draggletail’’ in Thackeray’s satire, however high the circle in which they move, their manifest lack of noble repose only swells the confusion. Moreover, no genuine aristocracy can be founded solely on wealth. The ‘‘bounder-nobility’’—as an irreverent press styles them—are noble in their own eyes alone. In an exalted circle money should never be thought of, much less mentioned! The bee queen builds no cells and gathers no honey. No increment of beebread or royal jelly is due to her own activities, or received by inheritance. Queen and environment are alike parts of one sys- tem. So it should be with a true aristocracy. No quest of gold, no promotion of enterprise, no regret over the past, no worry for the future, no will to know, no call to govern, no mission to control, no fear of loss, no hope of gain, ought to intrude to break the perfect peace. Kept in place by a reverent society, the Lord of Leisure need only pose as the glint of a sunbeam across the trail of the toilers, merely strew flowers along the pathway of life, in short be like Roses in their bloom Casting their petals ever on the grass Over the way the Beautiful must pass. II To all this there is an intensely practical side. If aristocracy is to endure (and without it this would be a dreary world indeed) it must be constituted aright. It must reject the false bases of heredity, effort and wealth. It must not be the reward of dis- GO TO THE BEE 453 tinction, nor be attainable by any competitive examination. Its door should open to all on equal terms, even though straight the gate and narrow the way. The present House of Lords, now swollen to seven hundred members, including almost everybody able and willing to pay the price, will die of its own accord. Let it alone. Let the climbers of yesterday keep on climbing, while we remove the ladders behind them. Let the men who replenish the party treasuries receive in the name of our gracious Kine— our sole true aristocrat—whatsoever honors their patriotism de- serves; but for the good of society let us build a new class in a new way. Should we fail in that endeavor the iconoclasts of the day will cast us all into the melting-pot from which no leisurer returns, an upshot the vicar would sadly deprecate. To select noblemen by primogeniture is a process comparable to choice by loaded dice; the cast is made before the heir is born. Nothing could be more undemocratic, nothing in reality more un- fair to men and women of the race in general. Then let our lords be chosen by lot from among the people at large, let us pick a cer- tain number of babies to be our future leisurers, and feed them on royal jelly* or the nearest parallel to that condiment our gracious King may secure. All expense involved should of course be carried by the people. In the United Kingdom are some 48,000,000 men and women. Let each pay alike; the assessment would then be so small that no one would even notice it. Let each contribute say a penny yearly for social inflorescence—for the perfected blossom of humanity. Such trifling levy would amount in the aggregate to £200,000 sterling, which, judiciously invested, would yield an assured annual income of £10,000, a sum quite adequate to provide for a leisurer through- out life. And the contribution by everybody of one shilling, a tax still absurdly small, would support twelve members of the new aristocracy each year. The necessary sum once collected, the infant chosen should be entirely healthy, and so attested by a Court physician accustomed to the needs of the leisure classes. It ought also to be a manchild, and its future mate, having no title in her own right, would be- come a ‘‘Lady’’ by courtesy, even as now the wife of the knighted erocer or jockey is recognized as ‘‘Lady’’ Jones or Atkins because her husband has been touched by the flat of the King’s sword and allowed to write ‘‘Sir’’ before his name. Only through an impartial selection may aristocracy and democracy be satis- 1 This expression is of course purely figurative, because no product of Cross and Blackwell serves our indicated purpose. It is with the general problem of perfected environment that we have to deal. 454 THE SCIENTIFIC MONTHLY factorily reconciled; and only when begun before effort, am- bition or deterioration has set in can the budding leisurer be ade- quately trained by nurse and butler in the thoughts and manners proper to a perfect lord. Otherwise, less lovely traits might have become stiffened beyond remedy. As his lordship grows up, the necessary allowance should come to him in regular sums only. He should never forestall, never hoard, never gamble, never speculate, never go into trade, never run into debt, never have anything left over after Christmas! He should support society as society has supported him; but chiefly he and the lady he may happily choose must remain through life ‘‘on the hills as gods together, careless of mankind.”’ True, as the noble Lord Tennyson once observed, ‘‘kind hearts are more than coronets,’’ but there is nothing in the plan to inhibit possession of both at once. A coronet, moreover, may be very be- coming as well as very welcome to My Lady. For our new-made lord will never marry for money nor as a rule where money is— all dowry acquired being turned over to the state; and what more exquisite pleasure than that of a young maiden unexpectedly chosen for the high distinction of a coronet! Let us also notice the amazing widening of the possible range of choice when no dowry need be sought. Doubtless a new title ought to be devised for the consummate flower of leisure. Lord, Duke, Earl, Knight—all these hark back to the discarded emblems of war, ‘‘the faded fancies of an elder world.’’ The vicar himself, I believe, was undecided, but the plan developed from hints given in his discourse should not fail just because a suitable name is not immediately forthcoming. The hellenistie term, ‘‘Bianthine,’’ ‘‘flower of life,’’ would be appro- priate, but it seems rather long, used as we are to the abrupt Saxon “‘Lord and Lady,’’ or the Norman ‘‘Sir.’’ ‘‘Flovite’’ (flos-vite) might do; represented by the letters F. V. it would be a natural contrast to M. P., and a pleasing reminiscence of F. F. V., the designation of certain Elite of the United States of America. This again suggests that the word Elite itself, a good Norman ex- pression much valued by our Gallic allies, may be the term we are seeking. In any ease, the title selected should in some way indicate one chosen from among many, first among equals, the bloom of exist- ence, the triumph of aspiring democracy reaching the goal of per- fection amidst the leaven and the levelling of the commonry. And we hope that the admirable vicar may find ample support in his noble crusade to make the British peerage once more a counsel of perfection. THE FOOD RESOURCES OF THE SEA 455 THE FOOD RESOURCES OF THE SEA By GEORGE W. MARTIN ASSISTANT PROFESSOR OF BOTANY, RUTGERS COLLEGE, NEW BRUNSWICK, N. J. RIMITIVE man got his food as his competitors did—that is to say, he picked it up or killed it where he could find it. Very early in his civilized career he ceased to be a hunter and began to cultivate the land; in fact, the beginnings of civilization and of agriculture were contemporaneous. Since that time the pressure of increasing populations demanding to be fed has been a prolific source of human strife. There are not lacking economists who would maintain that the recent catastrophic war in Europe was the direct result of an increasing demand for food on the part of the rapidly multiplying German nation. JEHast, in THE Screntiric MontrHiy for December, 1921, points out that the agri- cultural resources of the United States can in all probability not support in reasonable comfort more than two hundred million people and that the present indications are that our population will reach that figure within the next century. Furthermore, we ean not count on importing food indefinitely, since by the time our own population reaches its limit, the now scantily peopled parts of the world will produce little or no food in excess of the needs of their own greatly augmented populations. His is but one of many voices warning us that there is a limit to the number of human beings whom the earth can support, and however we may disagree with the various estimates as to what the limit may be, we can not doubt that it exists, and that, historically speaking, we are rapidly approaching it. The purpose of this article is, however, not to consider this question in detail, but merely to point out one source of food of which the possibilities are still largely unrealized. If we were to-day still depending upon the chase as the main source of our food, most of us would be dead, or, rather, we should never have been born. Yet so far as the oceans, which cover three fourths of the surface of the earth, are concerned, we have made little essential advance over the methods of the primitive fisher- men. The flocks and herds of the sea still roam freely in their native haunts, and we cast our lines and nets over their feeding grounds, and catch what we can. Our operations are on a larger 456 THE SCIENTIFIC MONTHLY seale, it is true, than those of our predecessors, our tackle is su- perior, our nets larger and stronger, and, by equipping our fishing vessels with steam or gasoline power, we have enlarged the area we can cover. Once the fish are landed, we have an elaborate system of distribution and marketing, so that cities a thousand miles inland can have fish a day out of the ocean shipped to them in fast refrigerated cars. There is also a little direct utilization of the plants of the sea. In certain parts of the world, notably in China and Japan, and to a less extent in HKurope, a few of the alez are eaten by man or his domestic animals, or gathered to be utilized in some minor industry. But so far as the actual cultiva- tion of the sea’s resources as distinguished from their mere ex- ploitation is concerned, we have made only the feeblest beginning. The reasons for this are, of course, the uncontrollability of the sea as compared with the land; its instability; the vastness of the oceans and the relative inaccessibility of much of their area; and especially the difficulty of attempting to control living organisms in the sea, out of man’s natural element, as they may be controlled on the land where he is at home. Yet if the demand becomes in- sistent enough we ean not doubt that methods will be devised which will give us the desired results. To question that would be to admit that man has neared the culmination of his evolutionary career and is preparing to bequeath the mastery of the earth to his successor, whoever that may be. The bulk of the food supply which we have come to expect the ocean to furnish us is animal. Animals that live in the sea are, however, no less dependent upon plant life for their food than are land animals. We all know that the beef we eat is built up from the grass and grain upon which the cattle have fed; and, while there are many animals that feed upon other animals only, sooner or later the cycle goes back to the green plants. This is because the green coloring matter contained in plants is the only substance known that can so combine carbon and water as to form the carbohydrates that are the fundamental materials of which all living beings, whether plants or animals, are constructed. The plants living in the sea are then the equivalents, so far as the life of the sea is concerned, of the land plants. Like land plants, they not only need carbon and water, but certain mineral salts. Of these, those that furnish them with nitrogen and phosphorus are most important, since they are most apt to be present in in- sufficient quantity and thus to be limiting factors. The plants in the sea are continually using these substances and are continually dying or being eaten by animals. Sooner or later there comes a time when by the death and decay of the plant or animal most of THE FOOD RESOURCES OF THE SEA 457 these materials are returned to solution. Part, however, have been transformed into insoluble compounds and are lying inert in the depths of the ocean. Thus there is a constant loss of nutrient salts. This loss is replaced by drainage from the land, the great rivers carrying constantly into the ocean an almost incredible amount of dissolved minerals in addition to suspended matter. Another important requirement of plants is light. Plants on land receive the full benefit of the sun’s rays as we know them. Plants living under water receive only a portion of the rays that reach the land. Part of the light that strikes the water is reflected, and the part that penetrates the water is gradually absorbed in passing through that medium, the red and yellow rays first, the blue and violet last. This differential absorption is reflected in the curious and well-known vertical distribution of marine alge according to color—the green kinds growing in shallow water, the browns in an intermediate zone and the reds in the deepest water, although there are, of course, numerous exceptions to this general rule of distribution. Another property of light is that it is re- fracted by water, and the greater the angle at which the rays strike the water the greater will be the refraction. In the tropics, where the rays are practically vertical, the amount of refraction is in- significant, but in high latitudes, where the rays strike the water at a sharp angle, the refraction is marked, as a result of which the rays are bent into a more nearly vertical direction, thus inereas- ing their penetration in depth and partly compensating for the unfavorable angle at which they strike the water. The penetra- tion is also markedly affected by the amount of suspended matter and the number of microorganisms present in the water. Helland- Hansen was able to show that in the Atlantic Ocean south of the Azores, on a bright summer’s day, light is abundant at a depth of 100 meters, still including at that depth a few red rays. At 500 meters the red rays have completely disappeared, but blue and ultra-violet rays are still plentiful, and may be detected at 1,000 meters, but have completely disappeared at 1,700 meters. Ht is not probable, however, that under the most favorable conditions photosynthesis may be carried on at depths greater than 200 meters. Temperature is less directly important in the sea than on land since there is no great danger of injurious extremes being reached. Indirectly, its importance lies in the fact that carbondioxide is much more soluble in cold water than in warm, (Fig. 1) and it is probably this, rather than the direct influence of temperature which accounts for the fact that the most luxuriant development of plant life is in the colder waters of the earth. 458 THE SCIENTIFIC MONTHLY 0.6 o4 02 CO; at re = 3 re) 10° 20 aio) 40° C. = 32° 50° G8? 86° 104° F. Fig. 1. SOLUBILITY OF CARBON DIOXIDE IN WATER. CUBIC CENTIMETERS OF C09 SOLUBLE IN 1 C. C. OF WATER, REDUCED TO NORMAL TEMPERATURE AND PRESSURE On the basis of habitat, marine plants may be divided into two great groups. On all coasts, and in shallow waters near the coasts, there is usually a conspicuous zone of ‘‘seaweed.’’ This is known as the benthos, from a Greek word denoting bottom, because it occurs attached to the bottom on the shelf of shallow water sur- rounding the coasts. The plants of the benthos are extremely varied and range in size from microscopic to very large, the giant kelps of the northern Pacific sometimes reaching a length of nearly four hundred feet. We are accustomed to think of these plants as alow, and most of them belong to one or another of the great algal groups, but the most important of all the plants of the benthos is not an alga but a flowering plant, belonging to the pondweed family, and not very distantly related to the grass family, which is the most important economic family of land plants. This is Zostera marina, the common eel-grass. Zostera marina or some of its near relatives occurs on almost all the ocean shores of the globe, being absent only in the extremely hot and extremely cold portions. It grows in water varying from a little over a meter in depth at low tide, to depths of fifteen or twenty meters in cer- tain clear Mediterranean waters; and in waters varying in salinity from that of the sea water of the open ocean to that of brackish bays and estuaries of less than half full salinity. It prefers mud, but will grow nearly as well in sand. It will not grow, however, where the bottom is composed of loose stones nor where the wave action is severe, and these localities are inhabited by the true alge. The windrows of dead ‘‘seaweed’’ commonly found east up on our Atlantic shores are very largely composed of Zostera. A few other flowering plants occur in salt or brackish water, but they are relatively unimportant. THE FOOD RESOURCES OF THE SEA 459 The algx of the benthos occur, as previously stated, in three rather indefinitely limited zones. In the quiet waters of shallow bays we find a great many green and blue-green alge. Such forms as Ulva lactuca and Enteromorpha intestinalis, both often called sea lettuce, are typically found in such situations. The branching, feathery masses of Cladophora and the thick green felt of Vaucheria are also often prominent. The blue-green alge are en- tirely microscopic, but often occur in great abundance. A species of Lyngbya frequently forms felted mats two or three inches thick and many feet across. Masses of Spirulina as large as a dinner plate are common. The plants which grow between tide levels or just below low tide, at places where there is considerable wave action, belong mostly to the brown alge. Various rockweeds, belonging to such genera as Fucus and Ascophyllwm, are common in northern waters, and the gulfweed, Sargassum, is typical of the forms growing in the warmer waters. below low tide level the great kelps—the Laminarias and their allies—are the largest plants of the ocean, the larger species occurring, however, only in the cooler waters. The alge of the deeper waters are mainly the reds, and the farther south we go the greater becomes the preponderance of the red algw. It must be remembered, however, that red, brown, green and blue-green algw are more or less inter- mingled at all depths and in all latitudes. All the plants we have been considering are alike, however, in that they occur only near the shores, except in cases where they have been torn loose from their fastenings and carried by currents into the open sea. The Sargasso Sea of the Atlantic Ocean is merely an area outside of the track of the great oceanic currents and therefore constituting a huge eddy in which such material accumulates, growing vegeta- tively to a certain extent and finally dying and sinking. If the only marine plants were in the benthos, in spite of the local luxuriance of its growth, the great mass of the ocean would be a desert, incapable of supporting anything like the amount of life which actually exists in it. There is, however, another great group of living organisms, the plankton. The plankton comprises those plants and animals that are neither attached to the bottom nor able to swim against a current, but normally live floating in the water and carried by it from place to place. Some of the animals of the plankton are of rather large size—such, for example, as the Portuguese man o’war and some of the larger jellyfish. Most of them, and all the plants, are microscopic. The plants, in fact, are all unicellular, although the cells are often united into filaments or colonies of various shapes. Two groups of plankton organisms are of special importance: the diatoms and the peridines. 460 THE SCIENTIFIC MONTHLY The diatoms are unicellular plants occurring in all waters, but especially abundant in the colder parts of the ocean, growing in enormous numbers in the Arctic and Antarctic regions and reach- ing in temperate waters their maximum development in the cooler months of the year. They are unicellular organisms, enclosed m two silicious shells, one fitting over the other much as the lid fits the bottom of a pill box. In shape they are extremely varied, and the shells are usually marked with intricate patterns and often decorated with spines and knobs. The peridines constitute a curious group of unicellular organ- isms intermediate between plants and animals. Some of them are holophytie or plantlike in their method of nutrition, others are saprophytic, absorbing dead organic material dissolved in the water, some are parasitic in the body cavity of small animals, such as copepods; a large number are holozoic, capturing and ingesting their food just as animals do. The peridines very largely replace the diatoms in the tropical seas, and in temperate waters attam their maximum numbers in the warmer months when the diatoms are at a minimum. This seasonal distribution is very possibly cor- related with the diminution of the carbon dioxide content of the warmer waters. The peridines may be naked, or provided with a simple or elaborate armor of cellulose, the same material as that of which plant cell walls are composed. Whether armored or not, they are always provided with two flagella, one, hair-hke, which trails behind them, the other a ribbon-like undulating structure typically encircling the body in a special groove provided for it and giving to the organism a rotatory motion. In spite of their minute size it may fairly be said that the dia- toms and the holophytic peridines are to the ocean what the grasses are to the land. What they lack in size is more than com- pensated for by their extreme abundance and rapidity of reproduc- tion. Under favorable conditions hundreds of thousands of them may exist in a liter, so that the water is distinctly colored and is almost soupy to the touch. A third group of important plankton organisms, entirely holo- phytic in their nutrition, is known as the Coccolithophoridae. These are brown-pigmented forms secreting very regular calcareous shells and occurring in the warmer parts of the open ocean. Most of them are extremely minute and while they are known to occur in extremely great abundance at times, because of their small size little is definitely known about their distribution or relative im- portance. In addition to these groups of organisms, the zoospores and gametes of algwe are liberated in enormous number at certain sea- sons of the year, and add their contribution to the food supply of THE FOOD RESOURCES OF THE SEA 461 the plankton, while there are numerous brown and green flagel- lates belonging to other than the three groups mentioned. The phytoplankton exists everywhere at the surface of the ocean and for a considerable depth below the surface. Certainly it is fairly abundant as far down as a hundred fathoms in those parts of the ocean where the light approaches the vertical. It is, however, much more abundant near land than in mid-ocean, mainly because the organisms near land are in more immediate contact with the nutrient materials washed from the land into the sea, and tend to exhaust the supply before these materials reach the open ocean. The ratio between the number of plankton organisms near land as compared with the number in the open seas has been given as fifty to one, but this estimate, for reasons which will be apparent later, is probably much too high. It is not enough to know life in the ocean qualitatively. It was long ago realized that it must be analyzed quantitatively. The earliest quantitative studies were those of Hensen, beginning about 1880, and modern work on the problem may be said to begin with him. He devised the forerunner of the present-day plankton net and attempted to account for the production of the sea by his eol- lections. His nets missed all the smaller organisms and failed to secure many of the larger ones. Hensen realized this and at- tempted to allow for it in his caleulations but his results were nec- essarily very imperfect. He succeeded in imparting new impetus to such studies, however, and deserves the credit due to a pioneer worker in a difficult field. The amount of food required by the animals of the sea is so much in excess of the amount shown to be produced by these early methods that Piitter (1907-1909) argued that the nutrition of marine animals was on an entirely different plane from that of land animals, and that a Jarge number of them, especially the smaller ones, absorbed dissolved organic matter di- rectly from the water without the mediation of plants. Piitter’s arguments have not been generally accepted and more recent studies have invalidated many of them. Nevertheless, it is pos- sible that something of this sort is more general than we realize. Mitchell has recently (1917) reported on experiments strongly in- dicating, although perhaps not proving, that so highly organized an animal as an oyster can utilize dextrose dissolved in sea water, transforming it into glycogen and storing it. In 1911 Lohmann showed that many of the organisms that pass through the plankton net may be secured and studied by the use of a centrifuge. This, again, marked a great advance. He showed that these smaller organisms, for which he proposed the convenient term ‘‘nannoplankton,’’ existed in enormous numbers in parts of the sea which the net collections seemed to show were barren, 462 THE SCIENTIFIC MONTHLY and found that certain animals—Appendicularia e. g.—were very efficient collectors of these minute forms, sometimes feeding on them almost exclusively. Gran reports a similar experience in the Straits of Gibraltar. Net collections showed very little plankton in the water, but the stomachs of Salpae when examined were found to be crammed with forms too small to be retained by the nets. Even centrifuging fails to get many of these forms since they are often lighter than water or as light, or are so delicate that the operation destroys them. Allen has devised a dilution culture method similar in principle to those used in making bacterial counts. He adds to a quantity of filtered sea water a few drops of a nutrient solution, sterilizes it, and adds to it a known quantity of the sea water to be investigated—say, one cubic centimeter. After thorough shaking, this is poured into about a hundred sterile flasks and put in a north light for a few days. The flasks are then examined and the organisms that have developed in them identi- fied, and the number of different kinds that are found in each flask are recorded. A given species is recorded only once from each flask and sinee at least one individual of the species must have been introduced to permit such development, the count is still too small. By actual experience this gave results of 464 organisms per cubic centimeter or 464,000 to a liter. Bacteria were disregarded. The centrifugal method applied to the same water gave a count of 14,450 to a liter. Allowing for duplication in some of the flasks and for the failure of many forms to develop under laboratory conditions Allen concludes that a count of 1,000,000 organisms per liter would be conservative. That is, one organism to each cubic millimeter. Assuming an average size equivalent to a sphere with CO Sick TIE. a diameter of five microns, that is not excessively crowded, as the diagram will show. (Fig. 2.) In this country, Moore, fol- lowed by Grave, attempted to } calculate the ‘‘food value’’ or areas of water on the basis of similar plankton counts, work- ing with special reference to the nutrition of oysters. They found out what organisms occurred in greatest numbers in the oysters’ ie aie stomachs, studied the rate of Fig. 2. COMPARATIVE SPACE OCCUPIED feeding and then proceeded to BY AN ORGANISM 6 MICRONS IN pDIAM- collect and analyze samples of ETER IN A CUBIC MILLIMETER : (1/1,000,000 tareR) of WATER water over actual or potential THE FOOD RESOURCES OF THE SEA 463 oyster beds, finding out just how many of these organisms occurred to the liter. They then carefully calculated the volume of each organism and by multiplying the volume by the number of times a given organism occurred in a unit volume of water and adding the totals, the ‘‘food value’’ was secured. This method gave much valuable information, but the results can not be regarded as final for several reasons. For one thing, organisms differing so widely as diatoms and peridines are not comparable on the basis of their volumes, as has been pointed out by Brandt and Juday. Again, in a locality where oysters are not naturally erowing, their introduction often materially increases the supply of food, since their shells form substrata and their excreta help in the nourishment of numerous food organisms. Finally, they took no account of the food value of the nannoplankton nor of the detritus, which may be very considerable. The most elaborate attempts to calculate the production of the sea have been those of the Danish biologist Petersen and his asso- eiates. As a result of their studies, these workers have come to the conclusion that the plankton plays a very small part in the nutrition of the animals of the sea and that the fundamental food of all marine forms in northern waters at any rate is the ‘‘dust- fine detritus’’ of the sea bottom, derived primarily from the eel- erass, Zostera. As an indication of the degree of progress which Kattegat the figures indicate thousands of tons Bl is Plankton Plaice ete. 5 Cod etc. © Herring etc. 7 nt tt | Predato CR : a4 igs ce 1 Useful animals Useless animals 5000 Zostera 24000 Fic. 3. ANIMAL LIFE IN THE KATTEGAT DERIVED FROM ZOSTERA, AFTER PETERSEN 464 THE SCIENTIFIC MONTHLY these workers have made, the preliminary conclusions in the life in the Kattegat may be cited. The Kattegat is the rather shallow body of water between eastern Denmark and Sweden, having an extreme length of 150 miles and an extreme breadth of about 90 miles. It is assumed that about half of the total amount of Zostera annually produced in this area is washed elsewhere by the currents. The balance, estimated at 24,000,000 tons, serves as the basis for the animal life of the area. (See chart, Fig. 3.) The useless animals, that is, those that are of no value to man and do not serve as food for fish, feeding directly on the Zostera, amount to about 5,000,000 tons. Useful animals, mainly those capable of serving as food for fish, are estimated at 1,000,000 tons. These are not all utilized by food fish, however. Starfish account for per- haps 200,000 tons; 500,000 tons are eaten by the larger gastero- pods and crustaceans, of which only a part are consumed by fish ; while plaice and other flatfish consume about 50,000 tons, producing 5,000 tons of human food annually. Cod are much less economical, since they get their food at third hand, so to say, and each ton of the 6,000 tons produced annually represents about one hundred times as much of the original synthesized organie food. On the other hand, the cod help to keep down the predatory gasteropods and crustaceans. The herring is the most important food fish feeding on the plankton, (mainly on copepods) and it in turn is eaten by the cod. Perhaps the most striking feature brought out by these figures is the comparatively trifling amount of human food finally produced from such a large amount of organic material. So much, then, for the life naturally existing in the ocean. How may our utilization of it be more intelligently directed? Ob- viously the most economical use of it as food would be for man or his domestic animals to eat it directly. So far as the use of alge as food for human beings is concerned, we cannot expect, for the present at least, any considerable increase over the insignificant amount now consumed in the United States. In some countries algw have been used as food for stock for centuries, and recent experiments have indicated that cheap and effective treatments may make possible a substantial increase in such use. It has re- cently been shown that the kelps exhibit wide variation in their carbohydrate as well as in their iodine content during the growing season and future utilization, to be profitable, must take such fluctuations into account so that the plants may be harvested at the proper season. We would not think of doing anything else in the case of a land plant. And it may be mentioned in passing that the old method of burning kelps for potash and iodine content THE FOOD RESOURCES OF THE SEA 465 cannot be regarded as anything but a wasteful process even when temporary conditions permit it to compete with other sources of these products, as during the world war. Recent experiments in Sweden have suggested that dry distillation will yield, not only the potash and iodine formerly sought, but numerous other products, including illuminating gas, acetic acid, methyl alcohol, formic acid, acetone and creosote. -Since, however, man prefers to harvest the plant life of the sea indirectly, those animals which feed directly on the plants are able to increase with less waste and at a more rapid rate, considered in total populations, than those which feed on other animals. Most of our food fish, for example, feed on smaller fish; these in turn feed upon small crustaceans and the latter eat the microscopic plants and detritus, so that in many instances the fish we eat are removed three or four steps, perhaps more, from the original food source. This is more significant than may seem apparent at first glance, since it involves an enormous waste. Before any organism can grow, the energy needed merely to live must be supplied, and by the time a crustacean is eaten by a minnow, or a minnow by a food fish, it will, on the average, have consumed a quantity of food several times its own weight. These facts are well brought out in the diagram and statistics of Petersen, previously quoted. The edible shellfish, however—oysters, clams, mussels and the like— feed for the most part directly on the marine plants and this is one reason why the extension of the shell fisheries represents so much promise. _ The carp is one of the few edible fish which lives directly on vegetable food. In the United States most people do not, it is true, regard this species as particularly edible but since it is largely eaten in Europe and raised for the purpose, it will serve as an excellent example of what such a fish may produce. The statistics on this fresh water fish are particularly valuable because they are not subject to the sources of error which hamper attempts to meas- ure the productivity of the sea. The amount of fish produced. in carp ponds has been calculated as ninety-five pounds on the average each year per acre. Brandt calculated the productivity of Kiel Harbor as eighty-nine pounds per acre annually, but the latter figures are much less exact. The average yield of beef on good land in the United Kingdom is seventy-three pounds per acre an- nually. The beef is much superior in food value, pound for pound, but it is also much more costly to produce. There are many inlets of the sea where conditions are almost as readily controllable as they are in the fresh water ponds. ‘Such, for example, are the shallow enclosed sounds on the Atlantic coast of the United States Vol. XV.—30. 466 THE SCIENTIFIC MONTHLY —Great South Bay in Long Island, Barnegat Bay in New Jersey, Albemarle and Pamlico Sounds in North Carolina and parts of Chesapeake Bay. These enclosed areas are at present producing a great deal of human food, but only a small part of what they might produce under proper management. On a larger scale, the Baltic and North Seas of Europe are similar regions where, if not sea culture, at least an intelligent harvesting of the seas’ resources is yearly becoming more possible because of the careful studies which have been made by the English, German and Scandinavian biolo- cists, whose countries are chiefly interested in the matter. Before we can expect to make substantial advance we must have a much more comprehensive knowledge of the ecology of the sea than we have at present. For example, we may class the animals of the sea roughly into those which are valuable to man and those which are not, but when it comes to determining to which of these classes any particular species belongs, difficulties arise. The dif- ference which direct and indirect utilization of food makes has already been pointed out. Some useful species which get their food at second or third hand, that is, by eating other animals, prey largely on forms that would not otherwise be converted into human food; some, on the other hand, eat species that are good human food. Thus, drumfish are good human food, but they may at times consume large numbers of oysters which are still more valuable to man than they are. Starfish, which are utterly useless forms, alsu attack oysters, but this case offers no perplexities. A very large number of animals are useless in the sense that they occupy space and consume food which might otherwise be utilized by useful species. The snails, sponges, sea anemones and some of the mus- sels of our northern waters belong to this group. However, in the case of most marine animals only a complete account of their life histories, together with the life histories of their associated forms is sufficient to enable us to know whether they are, in the long run, valuable or harmful from the human standpoint and whether it would be wise for us to attempt to overturn the balance which we find them maintaining in nature. Another phase demanding careful study is the effect on the marine life of the waste materials which are constantly being poured into our waters, especially in the vicinity of our great cities. So far as industrial wastes and oil are concerned, the effect is wholly bad, and the questions at issue are: How much of this dis- charge is necessary? How can the pollution best be restricted to the necessary minimum? The sewage problem is more complex. The addition of large amounts of rich nitrogenous fertilizing mate- rial to our waters could be made a great source of wealth if it were THE FOOD RESOURCES OF THE SEA 467 properly utilized, and even under our present hit or miss methods it results in a marked enrichening of the marine flora in favored localities. On the other hand, the danger of disease transmission is so well recognized and is illustrated by so many striking exam- ples that large areas of sewage-polluted waters are eliminated as sources of food in whole or in part. How to utilize this valuable asset without endangering public health or spoiling the recreational value as well as the food value of our coastal waters is one of the biological problems of our day. Finally, it may be reiterated that shellfish culture offers the most immediate hope for effective utilization of the sea’s re- sources. The economy of direct utilization of plant food by these animals has been emphasized. Most shellfish, ike land crops, stay where they are planted. Even the seallop, which can swim about after a fashion, is restricted in its movements and could readily be controlled. Oyster culture is already a great and important industry but it has not nearly approached its possibilities. Clam culture is still in an embryonie stage and scallop culture has as yet merely been suggested. When some of the problems confront- ing the establishment of these industries have been solved we may hope to have acquired additional information concerning the ecology of the sea which will help us in our approach to the more difficult problems of the future. It is recognized that there are numerous economic phases in- volved in attempts to increase the productivity of the sea, but con- sideration of these would be beyond the seope of this paper. When social and economic forces demand additional food at reasonable prices, the biologists must be prepared to show where and how this may best be obtained. SSOULVATV ALOOS FHL 10 GvVaH THE ALBATROSS 469 OUR GREAT ROVERS OF THE HIGH SEAS— THE ALBATROSS By Dr. R2 WY SHUFELDT FELLOW OF THE AMERICAN ORNITHOLOGISTS’ UNION, WASHINGTON, D. C. N going over the literature devoted to ornithology, we find that but a small part of it refers to the birds known as Albatrosses. Alexander Wilson, the famous American ornithologist, never once mentions any of them in his work; and Audubon, who had splen- did opportunities to study them in nature as well as in museums and private collections, touched upon those he had heard of, or studied skins of, in the lightest possible manner. In volume VIIT of his work in my library, I note that he devotes but a single para- graph to the description of the genus (Diomedea). Apart from the deseription of characters, he gives but three and a half lines to the Yellow-nosed Albatross; a few lines more to the Black-footed Albatross, and four lines and a half to the Dusky Albatross—the last-named being the only one he figures. He was indebted to a ‘‘Mr. Townsend’’ for skins of all these species, the latter having collected them ‘‘not far from the mouth of the Columbia River.’’ As to the Black-footed Albatross, Audubon says: ‘‘It is clearly distinct from the other two described in his work, namely, the Dusky and the Yellow-nosed; but I have received no information respecting its habits. Not finding any of the meagre notices or descriptions to which I ean refer to agree with this bird, I have taken the liberty of giving it a name, being well assured that, should it prove to have been described, some person will kindly correct the mistake.’ He named it Diomedea nigripes, the Black- footed Albatross, and it is the name we have for the species to-day. In the last A. O. U. ‘‘Check-List”’ (1910), in addition to the bird just mentioned, we recognize four other species as belonging to the North American avifauna, namely, the Short-tailed Albatross (D. albatrus) ; the Laysan Albatross of Rothschild (D. immuta- bilis), and the Yellow-nosed and the Sooty Albatrosses (7. culmi- natus and P. palpetrata). These are all Pacifie Ocean birds, though the Yellow-nosed species is said to have ‘‘accidentally oceurred in the Gulf of St. Lawrence.”’ Personally, I do not recall ever having seen an Albatross in 470) THE SCIENTIFIC MONTHLY nature; only a few of our ornithologists have, and, as just stated, neither Wilson nor Audubon fared any better. However, I have earefully examined quite a number of them in the collections of - the United States National Museum; and not long ago, Dr. Charles W. Richmond, Assistant Curator of the Division of Birds of that institution, kindly loaned me the head of a specimen of the Sooty Albatross. It had no artificial eyes, and apparently was simply a head and nothing more—not even bearing any label or history. This head I photographed on side view, reducing it about one fourth, furnishing the print with an eye. That print is here reproduced as an illustration to my article. Audubon’s deseription of the beak of this species is so obscure as to be of but little value. The plumage of the head—a rich snuff-brown—is soft and com- posed of fine feathers, and there is a narrow white stripe of short feathers surrounding the posterior half of the eye-ld on either side of the head. The beak is glossy black and formed as shown in the eut. . There is in existence a wonderful literature on the Albatrosses, especially when we consider how few species there are compara- tively speaking. The old figures of them in the works are often very erude; while, upon the other hand, some fine photographie repro- ductions in different works are wonderfully fine and of great value. Among these are the remarkable photographs obtained by the Hon. Sir Walter Rothschild of the immense numbers of the Laysan _Albatrosses, nesting on the island of that name; of the dreadful practice of carting away the eggs of that species, taken at the same place, and many others. Then Mr. Dudley Le Souef, Director of the Melbourne Zoological Garden, has furnished us with a fine photograph of the White-capped Albatross on its egg, which latter, according to Professor Moseley, is held in a sort of pouch to be found. between the legs of the bird. Some species of these birds have a ‘‘tip to tip’’ measurement of the wings of no less than eleven feet and a few inches. It is a well-known fact that in the southern seas, where sailors have fallen overboard, they have been attacked in the water by one or more of these giants of the feathered race, and a poem on an ineident of this sort would quite offset the experience of the ancient mariner who shot the albatross, which furnished Coleridge with the mate- rial for his famous verses. The marvelous flight of one of these birds has been graph- ieally described by Mr. Froude, who tells us that “‘the albatross wheels in circles round and round, and forever round the ship— now far behind, now sweeping past in a long, rapid curve, like a perfect skater on an untouched field of-ice. There is no effort; ? THE ALBATROSS 471 watch as closely as you will, you rarely or never see a stroke of the mighty pinion. The flight is generally near the water, often close to it. You lose sight of the bird as he disappears in the hollow between the waves, and catch him again as he rises over the crest ; but how he rises and whence comes the propelling force, is to the eye inexplicable: he alters merely the angle at which the Wings are inclined; usually they are parallel to the water and horizontal; but when he turns to ascend or makes a change in his direction, the wings then point at an angle, one to the sky, the other to the water.’’ The bird of this group usually referred to in prose or poetry is the Wandering Albatross (D. exulans) ; and it is said that speci- mens of it have been collected having an alar extent of no less than twelve feet. It is a bird with extraordinary power of flight, and Professor Hutton has well deseribed the remarkable power’ these birds possess in that direction. ‘‘Suddenly he sees something floating in the water,’’ says this authority, ‘‘and prepares to alight; but how changed he now is from the noble bird but a moment be- fore, all grace and symmetry! He raises his wings, his head goes back, and his back goes in; down drop two enormous webbed feet, straddled out to their fullest extent; and with a hoarse croak, be- tween the ery of a raven and that of a sheep, he falls ‘souse’ into the water. Here he is at home again, breasting the waves like a cork. Presently he stretches out his neck, and with great exertion, of his wings runs along the top of the water for seventy or eighty yards, until, at last, having got sufficient impetus, he tucks up his legs, and is once more fairly launched in the air.’’ Another distinguished British writer on this subject, Professor Moseley, in describing their mating habits informs us that ‘‘when an albatross makes love, he stands by the female on the nest, raises his wings, spreads his tail and elevates it, throws up his head with the bill in the air, or stretches it straight out forwards as far as he ean, and then utters a curious ery. . . . Whilst uttering the ery, the bird sways his neck up and down. The female responds with a similar note, and they bring the tips of their bills lovingly to- gether. This sort of thing goes on for half an hour or so at a time.”’ There is great danger of the entire genus of Albatrosses be- coming entirely extinct in the comparatively near future, and for several very good reasons. In the first place, many are shot and killed by passengers and others from the decks of vessels of all descriptions sailing on the high seas. This practice claims its quota every year, and no use is ever made of the poor birds thus ruthlessly slain. Again, many are caught with hook and line, but 472 THE SCIENTIFIC MONTHLY these are usually released after being hauled aboard and giving an exhibition of their walking powers on the deck of the vessel. Another practice leading to their extinection—outranking all the others—is seen in the wholesale collection of their eggs for the markets of the western coasts of the Americas. The eges used to be gathered, and stifl may be, on their breeding grounds, more particularly on the Island of Laysan, by the cartload, none being left for the perpetuation of the species. At least this was not looked out for in the early days of this most reprehensible trade. Whether it is still going on I am unable to say; but should it be, steps ought to be taken to bring it to an end. In our bird fauna, the nearest relatives of the Albatrosses are the Fulmars, the Petrels and Shearwater. All these species possess the peculiar anatomy of the external nostrils—being designated in the vernacular as the Tube-nosed Swimmers, while in technical science the name Tubinares stands for the group. THE PROGRESS OF SCIENCE 473 THE PROGRESS OF SCIENCE CURRENT COMMENT By Dr. Epwin E. Stosson Science Service RELATIVITY AND THE ECLIPSE | On September 21 the theory of relativity was put to the proof. After the results of the photographs then taken have been measured we may perhaps know whether Einstein is to be ranked with Copernicus and New- -ton, among those who have revolu- tionized man’s conception of the uni- verse, or whether he will be regarded merely as the author of an ingenious mathematical theory of limited appli- cability to reality. For the last three years the theory of relativity has been the topic of lively discussion extending far beyond the scientific circle, for the public realized that some interesting issues were somehow involved in its incom- prehensible mathematics. More than a thousand books and uncountable articles have been published on Ein- | stein; all sorts, pro and con, physical | and metaphysical, experimental and | speculative, serious and Prizes have been offered for explana- tions in ordinary language. Personal, political, religious and racial pas- sions and prejudices have been aroused. Einstein was the first Ger- man scientist to be welcomed since the war, in England, France and the United States, but in his own country he has to go into hiding to escape assassination by the junkers. It is a remarkable example of how the progress of science may continue spite of political conflict that during the world war Einstein should have sat quietly in his study in Ber- lin thinking out his theory and that during the world war English astron- omers should have been quietly study- ing his work and preparing to put it in frivolous. | _ cautious creatures and not all of tion to the test at the earliest oppor- | tunity. This opportunity came on May 29, 1919, when there was a total eclipse of the sun. For Einstein had pre- dicted that when the stars about the darkened disk of the sun were photo- graphed they would appear as though pushed out of their positions. This is one of the consequences of his theory of relativity, which is designed to supplant, or at least to supple- ment, Newton’s theory of gravita- tion. According to Einstein, a ray of light from a star passing close by a heavy body like the sun is drawn out of its straight course a little, some- what as though it were a stream of material particles, but to a greater extent than Newton’s would allow for. To an astronomer looking up at the star along this crooked path and not making allowance for the bend, it would seem that the star had been moved away from the sun a minute distance (1.74 seconds of are). Of course this effect is the same at all times, but it can only be observed when the sun’s disk is com- pletely shadowed from us by the moon’s coming between. law So the British astronomer royal, Professor Eddington, sent out two eclipse expeditions in 1919 to points where the eclipse could be observed, one to the west coast of Africa and the other to the east course of Brazil. When he to develop and measure up his photographs, he found that the stars about the dark- ened sun were displaced in the direc- and close to the pre- dicted by Einstein. This was good evidence in stein’s favor, but came amount Ein- are them scientists ‘BSunox ‘iq pue ssoy ‘qd “V “ueyO ‘V ‘OD “Ad ‘say ‘SWEPY “A ‘DO “Aq ‘suepy “SII “WeUO SST Md ‘f¢ ‘f¢ ‘1ejduniy ‘¢ ‘A Ad ‘sputysoy ‘O ‘d ‘f ‘soavorsiey Lf ‘yorn®) “TH “T[oaxe TRITVM LOZ “VIpBIJSNy UW104S0 AA ‘yjaog wWory oanpivdop toy} OF rortd poydeasoyoyd ‘Az1vd asd ALUVd USdIION NVITVUISAVY NaaLsdM HAL qpqdwep “MoM “AG ‘Teqdwep ‘Smoot “fF “A ‘LO}ISSON ‘OD ‘ox 0} 4fo, ‘surpueyg SOJOUd uvyo ‘SII :8ulyyIg ‘wuNN “IW ‘YD ‘Q100W ‘H ‘£ ‘id ‘89}8X “S ‘O W VIN “ff 23U8M )9 OY} FO Scoqtuow IT, ePIA PILOA © THE PROGRESS OF SCIENCE were ready to accept so startling a theory as this without further con- firmation. The weather was very cloudy in Africa, and the only good photograph obtained at the South American station was one taken with a four-inch lens and showing seven stars around the sun. But there has been no other total eclipse observable till this year and this is not so good a one. There were no bright stars near the sun, in fact only one visible to the naked eye among those close enough to the sun so that their displacement could be measured. But there were four or five faint stars that may have been caught on a sensitive plate with a good telescope. Unfortunately too, the eclipse oe- eurred in a highly inconvenient part of the earth. Its track was along the Indian Ocean and through the heart of Australia where there are no observatories and few people. The best point was on Christmas Island, lying west of Australia and south of Java. This island only measures eight by twelve miles and has a pop- ulation of about 250, according to the latest census. But it was selected by the British, German and Dutch ex- peditions for it was in the middle of the track of the eclipse. The dark- ness there lasted five minutes and as- tronomers can do much in five min- utes. It appears, however, from cable despatches that the weather conditions were bad. An expedition from the Lick Observatory, Califor- nia, was stationed on the west coast of Australia, and the Observatory of Adelaide sent a party into the arid interior of Australia, which involved five weeks of travel by camel train but which was pretty certain to get cloudless weather. In Australia the weather was favorable. If the astronomical expeditions now in the field bring home con- firmation of. the results of the eclipse of 1919,- then we may have to get used to all sorts of queer ideas, be- | but 475 sides crooked beams of light in empty space. We may have to give up the force of gravitation and the ether and the constancy of mass and the distinction between matter and en- ergy. We may get to talking about the curvature of time, the weight of heat, kinks in space, atoms of energy, four dimensions, world-lines and a finite universe. We may be called upon to come to conceive of arrows that shrink and bullets that get heavier the faster they travel; of clocks that go slower the faster they travel and of a future that turns back and tangles itself up in the present. ' TANGLING UP THE TIME LINE EINSTEIN’S theory of relativity is like a magician’s bag. There seems to be no end to the queer things that can be pulled out of it. The more it is studied the more paradoxical it appears. The latest thing I have seen is the queerest, the idea that the future may get tangled up in the present or even in the past. It is all worked out mathematically in a book just trans- lated from the German, Weyl’s ‘¢Time-Space-Matter.’’ Too mathe- matical for most of us, but the point in plain language is this: Here is a line representing the course of time extending from the dim past into the indefinite future: Past Future The present iis the point where I stand, looking both ways like Janus not seeing any end in either direction. I am continually moving or being moved straight along the time road from left to right. Every instant I step from the past into the future. Every instant a bit of time is taken from the future and added to the past, though neither gets any smaller or larger since both are in- finite. The past time and the future time are permanently separated by the moving present where I am and Copyright Harris and Ewing PRESIDENT SAMUEL W. STRATTON of Standards since its establishment in sachusetts Institute of Technology. Director of the Bureau 1901, now elected president ef the Mas THE PROGRESS OF SCIENCE 477 there seems no chance of the two kinds of time ever getting mixed up for they extend in opposite directions. But waitt—here’s a disconcerting idea. If I roll up the paper I ean make the future touch the past. I can overlap them. I can put A.D. into B.C. and what becomes of chro- nology then? We are used to this curving of ap- parently straight lines in space ever since 1492 when men found that they were not living on a flat earth but on a sphere. If I travel straight east from this town I shall eventually come back to it from the west. How far I shall have to go depends upon where I live. If my home were on the equator, I should have to travel 25,000 miles to get to my starting point. If it were near one of the poles, I could do this astonishing stunt in the course of a morning’s walk. Now, according to LHinstein, the time line is like the space lines. The framework of the world is measured by four dimensions, three of -space and one of time, namely, the up- right-left, to-fro, past-future lines. But these are not rigidly fixed. They may be bent and distorted like a bird cage that has been twisted and crushed, though every wire remains intact and conn ected to the other - wires just the same. Wherever there is a bit of matter, wherever there are electrical or mag- netic forces, there the time and space lines are more or less distorted. Einstem, reasoning from this idea, saw that a ray of light from a star, passing close by a heavy body like the sun, would not travel straight, but would be bent a little out of its course. The eclipse of 1919 brought the first chance to test Einstein’s idea, and the astronomer royal of Great Britain went to Brazil and took a photograph of the shadowed sun and seven stars about it. And the seven stars seemed shoved out of their customary places just as if in down, the region around the sun the space and time were puckered up in the way Einstein said they “were. When the eclipse of September 21, 1922, came, eight parties of astronomers were on the watch to see if the observations of three years before were confirmed. We have not heard their verdict yet, but, if their photographs measure up according to Einstein formula we shall have to get accustomed to the idea that time—like the tariff—is a local issue; that time measurements like space measurements are relative, not absolute, and that we are not sure of the constancy of our stand- ards of measure in either case. When two things happen in our presence we may be pretty sure which comes first. But if one event is here and another in Mars we can not be sure about priority with any conceivable system of clocks and signals. What seems past from one standpoint may seem future from another, for the time line may not run straight. Is your present condition in any the result. of your future actions? Can the light of a match be seen before the match is lit? Such a thing is conceivable in the generalized theory of relativity though, like most other conceivable things, it does not occur or is never known to occur in reality. But it is hard to get used to this strange new notion that the future may curl around in some sort of a circle and so come into the past. Did £ say ‘‘new’’? It was. a slip of the pen. For the idea is old. I open a volume of Egyptian antiqui- ties and I see carved on a monument of the Pharaohs a serpent with its tail in its mouth, the symbol of eter- nity, of which time is a segment. But what the Egyptians merely guessed at Einstein is putting to the proof. way HOW THE CHEMIST MOVES THE WORLD THE chemist provides the motive power of the world, the world of man, not the sinanimated globe. THE SCIENTIFIC MONTHLY World Wide Photos THE QUEST AT PLYMOUTH The Quest, the vessel of the Shackleton-Rowett Antarctic Expedition, on its arrival in Plymouth Harbor on September 16, after the expedition on which it embarked on September 17 of the previous year Archimedes said he could move the world if he had a long enough lever. The chemist moves the world with molecules. The chemical reactions of the consumption of food and fuel furnish the energy for our muscles and machines. If the only get control of the electron, he will be in command of unlimited en- chemist can ergy. For in this universe of ours power seems to be in inverse ratio to size and the mightiest. When we handle particles smaller than the atom, we can get behind the elements and may effect more mar- velous transformations than The smaller the building blocks, the minutest things are ever. greater the variety of buildings that. can be constructed. The chemistry of the past was a kind of cooking. The chemistry of the future will be more like astronomy; but it will be a new and more useful sort of astron- astronomer might omy such as an employ if he had the power to re- arrange the solar system by annexing a new planet from some other system or expediting the condensation of a nebula a thousand times. The chemist is not merely a manipulator of molecules; he is a manager of mankind. His discoveries and inventions, his economies and creations, often transform the condi- tions of ordinary life, alter the rela- tions of national power and shift the currents of thought, but these revo- lutions are effected so quietly that the chemist does not get the credit for what he accomplishes, and indeed does not usually realize the extent of his sociological influence. For instance, a great change that has come over the world in recent made conditions so unlike those existing in any previous period that historical precedents have no application to the present prob- lems, is the rapid intercommunica- years, and has THE PROGRESS OF SCIENCE tion of intelligence. Anything that anybody wants to say be com- municated to anybody who wants to hear it anywhere in all the wide world within a few minutes, or a few days, or at most a few In the agencies by which this is accom- plished, rapid transit by ship, train or automobile, printing, photography, telegraph and wireless, chemistry plays an essential “an months. telephone, wired or part, although it is so unpretentious a part that it rarely receives recog- nition. For instance, the expansion of literature and the spread of en- lightenment, which put an end to the Dark Ages, are ascribed to the inven- tion of movable type by Gutenberg, or somebody else, at the end of the fourteenth But the credit belongs rather to the unknown chem- century. ist who invented the process of mak- paper. The stamped their bricks and lead pipes with type, but printing had to wait ing ancient Romans > 479 more than a thousand years for a supply of paper. Movable not the essential feature of printing, type is for most of the printing done now-a- days is not from movable type, but We could if necessary do away with type and from solid lines or pages. press altogether, and use some pho- tographic method of composition and reproduction, but we could not do without The of wood-pulp paper has done more for paper. invention the expansion of literature than did the invention of rag paper 600 years ago. Print is only an imperfect repre- sentation of the sound of speech, a particularly imperfect representation in the case of English because we not tell half the words sound from their spelling. But the phonograph gives us sounds directly, and the extended the range of a speaker, un- can how audion and the radio have til now a speaker may have an audi- OFFICERS OF THE QUEST W orld Wide Photos After the death of Sir Ernest Shackelton, Commander Frank Wild succeeded him as leader of the expedition. eenter is Commander Wilson. He is shown second from the left. Mr. Wilding is shown on the right In the with the camera, 480 ence covering a continent and in- cluding generations yet unborn. What these inventions do for sound, pho- tography has done for sense of light. By means of them man is able to transcend the limita- tions of time and space. He can make himself seen and heard all round the earth and to all future the sister years. SCIENTIFIC ITEMS WE record with regret the death of Alexander Smith, formerly professor of chemistry at the University of Chicago and Columbia University; of Alice Robertson, formerly professor of zoology in Wellesley College; of David Sharp, formerly curator of the | Museum of Zoology of the University of Cambridge and editor of the Zoological Record; of F. T. Trouton, emeritus professor of physics in the University of London, and of HE. Bergmann, director of the Chemisch- Technische Reichsanstalt, Berlin. Sir ERNEST RUTHERFORD, Caven- dish professor of physies at the Uni- versity of Cambridge, has been elect- ed president of the British Associa- tion for the Advancement of Science THE SCIENTIFIC MONTHLY in succession to Sir Charles 8. Sher- rington. The meeting next year will be at Liverpool; the following year the meeting will be in Toronto. Dr. Rospert A. MILLIKAN, chair- man of the board of the California Institute of Technology and director of the Norman Bridge laboratory of physics, has been appointed a mem- ber of the committee .on intellectual cooperation of the League of Nations’ to succeed Dr. George E. Hale, direc- tor of the Mt. Wilson Observatory, who has resigned from the committee owing to the state of his health. Proressor W. L. Bragg, of Man- chester University, who, ‘together with his father, Sir William Bragg, was awarded the Nobel Prize for physics in 1915, delivered on Sep- tember 6 the lecture in Stockholm as prescribed by the statutes of the | Nobel Institution. TaIs Siliman Memorial Lectures at Yale University will be year’s | delivered by Dr. August Krogh, pro- | fessor of zoophysiology in Copen- hagen University. Professor Krogh has taken for his general topic ‘‘ The Anatomy and Physiology of Capil- laries.??°_ , THE SCIENTIFIC MONTHLY DECEMBER, 1922 THE VEGETATION OF AUSTRALIA AND NEW ZEALAND By Professor D. H. CAMPBELL STANFORD UNIVERSITY HOSE parts of the world which for one reason or another are completely isolated show very plainly the effects of this isola- tion upon the animals and plants which inhabit them. The degree of specialization in these organisms is to a certain extent an index of the length of time the region has been shut off. A comparison of these organisms with those of other regions may throw light upon such problems as the changes in the distribution of land and water upon the earth’s surface in the course of ages, and thus be of great interest to the geologist and geographer as well as to the biologist. If we compare the lands of the northern hemisphere, as they now exist, with the principal land masses of the southern hemis- phere, we find the former to be very much more extensive than the latter. In the north there is a marked preponderance of land in the polar and subpolar regions, which merge into the temperate regions in both the American and Eurasian continents. In the southern hemisphere there is an extensive almost absolutely barren polar continent, but the regions corresponding to the subarctic land masses of the north are entirely occupied by water; and the south temperate regions are completely separated from the antare- tie continent by a wide stretch of sea. Moreover, the temperate regions of Australasia, South Africa and South America are widely separated from each other by the Atlantic, Pacific and Indian oceans. In extent the temperate re- gions of the south are much less than those of the northern hemis- phere. As might be expected, this condition of things is accom- panied by a much greater diversity in the temperate floras of the southern hemisphere than is the case in northern latitudes. This perhaps reaches its maximum in the Australasian region, the com- pletely isolated Australian continent and the islands of New Zea- Vol. XV.—31. 482 THE SCIENTIFIC MONTHLY land having extremely specialized floras which are of very great interest to the student of plant geography. While Australia and New Zealand are usually grouped together geographically as ‘‘Australasia,’’ they differ much from each other in their vegetation, although having more or less in common. New Zealand is separated from Australia by over a thousand miles of sea, and many of the most characteristic Australian types are quite absent, and others only very sparingly represented. Owing to its very much greater size and range of climate, Australia, as might be expected, possesses a much more extensive flora than the rela- tively small islands of New Zealand. The completely isolated continent of Australia is almost exactly the size of the continental United States exclusive of Alaska. The Australian climate, however, is very different. The north- ern portion of Australia is within the tropics, the tip of York Peninsula being only 11 degrees from the equator, while the south- ernmost part of the continent scarcely touches the fortieth degree of latitude. The adjacent island of Tasmania extends about three degrees further south. The climate is therefore much warmer on the whole than that of the United States or Europe, the coolest regions in the south having a climate comparable to that of Califor- nia or the Mediterranean. At the north a true tropical climate prevails. The topography of Austraha is much less varied than that of the United States. There are no mountains comparable to our great western ranges, and there is a marked dearth of large rivers and lakes. The principal mountain masses are close to the eastern coast, a succession of mountain ranges and highlands extending from the York Peninsula to eastern Victoria and Tasmania. In Queensland there are some definite mountain ranges, but for the most part the high land is a plateau sloping gradually westward, with more or less definite escarpments toward the east. These escarpments sometimes exhibit abrupt gorges cut by the streams. These are well shown in the Blue Mountains west of Sydney. The highest point in Australia is Mt. Kosciusko, 7,300 feet, situated in New South Wales near the Victoria border. This highland region and the adjacent coastal areas have for the most part a good rainfall, but there are no large rivers. The heaviest rainfall is in the coastal region of North Queensland where at certain stations it may exceed two hundred inches annually and averages one hundred and fifty. Inland, however, the rainfall diminishes rapidly, and a third of the continent is said to have ten inches or less annually and another third less than twenty. This means that two thirds of the area of Australia must be classed as desert or semiarid, and VEGETATION OF AUSTRALIA 483 much of it unsuited to agriculture, although vast areas are more or less adapted to grazing, which at present in much of the com- monwealth is the most important industry. There is a more or less marked wet and dry season in most of Australia, as on our own Pacifie coast. In the south most of the rain falls during the winter months, May to September; in the north the heaviest rains fall in the summer. June is the wettest month in the south, January in the north. Northern Australia, lying entirely within the tropics, has for the most part a genuinely tropical climate, hot and humid in the coastal districts. In the more elevated regions of the plateau, how- ever, there may be sharp frost during the winter months, June to August. In August of last year I observed bananas and other tender plants cut down by frost at an elevation of 2,000 to 3,000 feet, in latitude 17°. On the coast, however, frost is quite unknown, and the forest shows a genuine tropical luxuriance. The wettest region in Australia is in northeast Queensland, on the coast, about latitude 17°. In this region a short range of pre- cipitous mountains rises directly from the coast to a height of over 5,000 feet, the highest land in the state. At the foot of this range, the precipitation is very heavy. One place, Babinda, which I visited in August, 1921, had already registered over two hundred inches for the year, and it rained almost incessantly during my stay. The low swampy forest about Babinda was almost impenetrable, the trees loaded down with creepers of various kinds, among which the rattan palms were only too conspicuous. Throughout the east- ern tropics the thickets of rattans are a great hindrance to progress in the forest, as their tough, horribly spiny twining stems make absolutely impenetrable tangles, natural barbed-wire barriers. Climbing Aroids and species of Vitis and Piper are also abundant as well as various other lianas. In these wet lowland jungles, the palms reach their fullest de- velopment, forming a conspicuous and beautiful feature of .the vegetation. One of the commonest and most attractive species is Archontophoenixz Cunninghaniana, often cultivated under the name Seaforthia elegans, and one of the most beautiful of all palms, with its smooth slender trunk and crown of graceful feathery leaves. No feature of the Australian vegetation is more beautiful than the groves of these lovely palms. Serew-pines (Pandanus) abound in this region and there are also a number of species of Cyeads. Australia is especially rich in these’ ancient plants. The most widespread genus is Macrozamia, of which there are several svecies, the genus having representatives in all the states. The two other Australian genera, Cycas and THE SCIENTIFIC MONTHLY Pic. 1. TROPICAL RAIN-FOREST, NORTH QUEENSLAND VEGETATION OF AUSTRALIA 485 Bowenia, are confined to tropical Queensland. The latter genus, peculiar to Australia, differs much in appearance from any living Cycads, in its solitary bi-pinnate leaves, rather suggesting a bracken fern. In the dryer parts of the Queensland coast the rain-forest is replaced by a more or less mixed forest, composed in part of Eucalyptus, and in part of tropical rain-forest types, like Ficus. A forest of this type may be seen occupying the sandy soil in the neighborhood of Cairns, the principal port of North Queensland. A feature of the coast in this district is the mangrove formation along, the shore and the banks of the streams flowing into the sea. Several genera are represented, the most important being the wide- spread Rhizophora and Avicennia. Some interesting ferns were noted in this region, the most striking being a gigantic Angiopteris which was seen in several places in the vicinity of Babinda. Immediately back of the coast the land rises rapidly to a plateau reaching an extreme elevation of about 4,000 feet, but averaging 2,000 to 3,000 feet over most of its extent. This table-land has an ample rainfall, and on the better soils develops a fine forest which yields extremely valuable timber. Much of the timber has been destroyed, but there are still some remnants which are accessible, and these are really magnificent examples of tropical forest growth. This tropical rain-forest 1s known in Queensland by the very inappropriate name of “‘Serub”’ and is confined to the rich basaltic and alluvial soils. The trees of this forest are mainly of Malayan affinity, and are tall with lofty straight trunks yielding a large amount of fine timber. Some of them, especially the Kauri (Agathis Palmerston‘) and ‘‘Red Cedar’’ (Cedrela toena) reach a very large size. The latter was formerly abundant and sometimes: attained a diameter of upwards of ten feet. It has been largely exterminated, but an oceasional fine specimen may still be seen, and the same is true of the Kauri. i Belonging to the same family (Meliaceae) as the cedar are several species of Flindersia, which are locally known as “‘hickory,”’ ‘“maple,’’ ‘‘beech,’? and other woods not in the least related to them. Other characteristic trees are Elaeocarpus, (Tiliaceae), Aleurites Moluccana, widespread throughout Polynesia; Sideroxy- lon (Sapotaceae), Eugenia (Myrtaceae) and others. The charac- teristic Australian family Proteaceae is represented in the rain- forest by several species of Grevillea, Stenocarpus, Macadamia and other genera. Grevillea robusta of southern Queensland is often grown in California as an ornamental tree. This upland forest has much finer trees than the lowland forest NTIFIC MONTHLY 1 u THE SOLE 486 TRUNK OF GIANT FIG, NEAR YUNGABURRA, NORTH QUEENSLAND Fie. 2. VEGETATION OF AUSTRALIA 487 near the coast, but the palms and some other tropical types are almost entirely absent, and the development of the epiphytes and hanas is not so marked, although these are by no means absent. Many of the large trees, as is so common in rain-forests everywhere, show a conspicuous development of buttresses at the base of the trunk. The giants of the forest are species of Ficus, the size of which is amazing. As in most tropical lands the genus is well represented in northern Australia, some species extending as far south as Sydney. Like so many other species of Ficus, these giant Queens- land figs begin life as epiphytes, the descending roots finally coalescing more or less completely, and strangling the host tree. The descending roots are produced in great numbers and in one tree that was seen, the huge conical trunk formed by the united roots was said to measure 120 feet in circumference at the ground, and the enormous spreading crown was in proportion. While the predominant forest on the plateau is ‘‘scrub,’’ there are large areas occupied almost exclusively by open Eucalyptus forest. This Eucalyptus forest is the dominant type of vegetation over much of Australia, but in the region in question is restricted to areas of sandy soil. The line between the ‘‘scrub’’ and the Eucalyptus forest is often very sharply marked, and is probably determined by the difference in the soil. In southern Queensland, in the neighborhood of Brisbane, the Euealyptus forest predominates, though there are also areas oc- eupied by ‘‘serub,’’ but many of the strictly tropical species of North Queensland are absent. Probably the most striking tree of South Queensland is the ‘“*Bunya’’ (Araucaria Bidwilli) a coniferous tree confined to a relatively small area in this region. It reaches a large size, and is valuable for its timber. The big seeds were much prized as food by the aboriginals. This handsome tree is frequently cultivated in California, where it seems very much at home. A second species, A. Cunninghami, is much more widely diffused, and was seen in extensive pure stands on some of the islands off the coast of Queens- land. An analysis of the constituents of the scrub vegetation of Queensland and New South Wales shows that it is largely made up of genera widespread through the Indo-Malayan region, or closely related to these, and may very properly be considered a part of the ereat Malayan flora. Such types as the figs, palms, screw-pines, Araceae, many epiphytic ferns and orchids are characteristic of the whole Indo-Malayan region; and as it is evident that northeast Australia was connected at no very distant period with the great island of New Guinea, it is pretty certain that this portion of the Australian flora is derived from the north. THE SCIENTIFIC MONTHLY 488 BOTANICAL GARDENS, BRISBANE BUNYA PINES (ARAUCARIA BIDWILLI), 9 vo. Fic. VEGETATION OF AUSTRALIA 489 This Malayan flora is best developed in northeast Queensland, some of the forms like the pitcher plants (Nepenthes) and certain genera of palms (Borassus, Areca, Caryota, ete.) being confined to the York Peninsula, Australia’s northernmost extension, which is separated from New Guinea by only about 100 miles of water. Some of the Malayan types, like Cedrela and two or three palms, and a considerable number of others, extend southward to the borders of Victoria; but this vegetation is confined to regions of ample rainfall and rich soils, and the number of these Malayan types diminishes rapidly toward the south. While the scrub vegetation is made up for the most part of the Malayan types there are a number of genera probably of Australian origin. Such are the fine trees Tristanea (allied to Euealyptus) the silky oak (G@revillea robusta), Stenocarpus and Macadamia of the Proteaceae, a family which reaches its maximum development in Australia. The luxuriant serub-forest disappears as one proceeds inland, and with the diminishing precipitation is replaced by the open Eucalyptus forest. Still further inland in Queensland are exten- Sive open grass lands or prairies which afford pasturage to great herds of cattle.’ To the south of Queensland is the state of New South Wales, the first colony to be established in Australia. The coastal region is a continuation of that of southern Queensland and has much the same vegetation as the latter, but the Malayan elements diminish toward the south, where there is an increasing proportion of true Australian types such as Euealyptus and Acacia. The scrub, how- ever, retains a decidedly tropical aspeet, with tall palms and tree- ferns in abundance. Much the greater part of the state, however, is far too dry for such forest growth, and is occupied by a very different type of vegetation. This is almost purely of Australian origin and is more or less decidedly xerophytie in character. The predominant trees are various species of Eucalyptus forming open forests with the sandy soil between occupied by a great variety of low shrubs, often with extremely showy flowers. Herbaceous plants are less conspicuous, although there are coarse grasses and a good many perennial plants growing from tubers, corms or bulbs. The Myrtaceae, so abundant in Australia, have numerous species of Leptospermum and Melaleuca; the Leguminosae include many species of Acacias, ‘‘ Wattle’’ in the vernacular, and a bewildering array of showy Papilionaceae ; several beautiful species of Boronia 1 Maiden, J. H.: ‘‘ Australian Vegetation,’’ p. 207. Federal Handbook for Australia, Melbourne, 1914. MONTHLY SCIENTIFIC THE 490 AANGAS ‘SNUGUVD TVOINV.LO ‘; DIT . VEGETATION OF AUSTRALIA 491 ‘and Eriostemon (Rutaceae) and many other striking and unfa- miliar flowers abound. The Proteaceae, as everywhere in Aus- tralia, are much in evidence, the most abundant being species of Hakea, Banksia and Grevillea. To this family belongs one of the most gorgeous of Australian flowers, the ‘‘Waratah,’’ whose mag- nificent scarlet flowers are the pride of New South Wales. Another very striking plant peculiar to New South Wales may be men- tioned—the giant torch lily (Doryanthes excelsa), bearing an enormous cluster of great scarlet lilies on a stout stalk ten or fifteen feet in height. Victoria, the smallest state, occupies the southeast corner of Australa, and is about the size of Kansas. It is the best eulti- vated, and apparently the most prosperous state of the common- wealth. Much of the state has a climate adapted to the cultivation of most crops of the north temperate zone, and better suited to the North European settlers than the hotter parts of Australia. Its smaller size and more uniform rainfall result in a lesser variety of vegetation than in the larger states; but in the mountain dis- triects of the east are found the tallest trees in Australia, close rivals of the California redwood. These forests of giant gums with their heavy undergrowth of tree-ferns and other luxuriant vegetation -are among the finest in the world. Where the forest has been cleared, the land is some of the best in the commonwealth. The distinctive Australian flora is seen at its best in West Australia. This immense state occupies the entire western third of the continent, and is almost completely separated from the eastern states by exvensive deserts, and is itself very largely a region of extremely low rainfall. There is, however, a small region occupying the extreme southwest portion, which has a fairly heavy rainfall, and this district possesses a flora which for variety and beauty has scarcely a rival anywhere in the world. Travelling overland from Victoria one traverses the rather un- interesting state of South Australia, and then proceeds by the recently completed line over the desert to West Australia. This desert is not unlike certain parts of our own western arid regions, often suggesting parts of Nevada or Arizona. While ex- ‘tensive tracts show only sparse salt-bush (Atriplex, Kochia), much “resembling the sage-brush deserts of Nevada or Utah, more often _there is a fairly heavy growth of small trees, interspersed with low shrubs, and sometimes bunch grasses and a few flowering herbaceous plants. The commonest trees are, as usual, species of Eucalyptus, but other abundant trees are species of Casuarina, whose thin leafless twigs simulate the needles of a pine. These curious trees, while not exclusively Australian, being also found in the Malayan region, reach their maximum development in Australia. 492 THE SCIENTIFIC MONTHLY 3 & Fe * : { i ’ Fig. 5. GIANT GUM-FOREST, VICTORIA VEGETATION OF AUSTRALIA 493 The commonest shrubs are species of Acacia and dwarf Eucalyptus, the former at the time of my visit being covered with masses of golden bloom, which enlivened the prevailing dull gray green tints of the foliage. A species of sandal-wood grows in this region as well as a number of other interesting trees and shrubs. Comparatively few showy flowers are seen, aside from the Acacias. Occasionally masses of pretty pink and white everlast- ings are encountered, and a gorgeous scarlet pea (Clianthus Dampier). As the western coast is approached the country becomes some- what less arid, and presently there appears along the railway line an increasing profusion of beautiful flowers, until before Perth, the terminus of the railway, is reached, the train travels through a veritable garden of brilliant bloom. The beauty and variety of this wonderful floral display must be seen to be appreciated. While some of the flowers, such as the great variety of pea-shaped blos- soms, suggest familiar northern types, many are entirely strange with little suggestion of relationship with any northern genera. Whole families, quite unknown to this northern botanist, are richly represented. Thus the Goodeniaceae, a characteristic Australian family, has a large number of extremely showy species of Goodenia (yellow), Dampiera and Leschenaultia (blue), one of the latter, L. formosa, of a wonderful blue that would put to shame an Alpine gentian. Ground orchids are very abundant, some of them of great beauty. They belong largely to special Australasian genera, Caladenia, Diuris, Thelymitra and other quite unfamiliar ones. The little sundews of northern bogs are here represented by an extraordinary assemblage of species, some slender, half climbing plants four to five feet high with flowers the size. of small roses. Pink Boronias and Tetratheea (Tremandraceae), yellow Hib- berties (Dilleniaceae) are a few of the many beautiful novelties among the lower growing species; while Banksias, Hakeas and Grevilleas of the Proteaceae; Leptospermum, Callistemon, and Melaleuca of the Myrtaceae, are the predominant larger growths. Of the Monocolyledons, aside from the orchids already referred to, and various grasses and sedges, there are a number of attractive species. The Iridaceae are represented by species of Patersonia with pretty blue or purple flowers. Of the lily-family are several species of Thysonotus, with delicate fringed petals, and Burchardia, whose umbels of pretty white blossoms suggest an Allium or the Californian Brodiza. Peculiar, if not beautiful, were the extraordinary grass-trees, or ‘‘black-boys,’’ as they are commonly ealled in the West. The larger species develop a stout trunk and somewhat resemble an THE SCIENTIFIC MONTHLY 494 Photograph furnished by Mr. C. E. Lane-Poole. GRASS-TREES (XANTHORRH@A PREISSI1), WEST AUSTRALIA 6. Fie. VEGETATION OF AUSTRALIA 495 arborescent Yucea, but the leaves, which are very numerous, are slender and more or less drooping. The insignificant flowers are borne on a elub-like spike, sometimes six or eight feet high. The plants are said to flower especially freely after a recent fire, and a grove of these strange plants with hundreds of these upright flower-spikes is one of the most striking botanical sights of Aus- tralia. A related genus, Kingia, is confined to western Australia. Among the most extraordinary flowers of West Australia are the ‘‘Kangaroo Paws’’(Anigozanthus). These flowers are of the most bizarre coloring—bright green and scarlet, yellow and black, red and yellow, or pure green. The genus is unknown outside West Australia. The only Gymnosperm noted was a Cyead, Macro- zamia Fraseri. This is very common, and is regarded as a serious pest, as animals are often poisoned by eating the young foliage in time of drought. Throughout the less arid parts of West Australia, this wonderful floral display may be seen in the spring, August to November. It perhaps reaches its culmination in the Albany dis- trict on the south coast. Certainly the variety of flowers near Albany surpasses anything the writer has seen in any part of the world. I was unable to visit the Island of Tasmania, which differs much in its topography and climate from the mainland of Austra- lia, and is much more like New Zealand in these respects. It is: very mountainous and in many parts, especially in the west, the rainfall is extremely heavy. This heavy precipitation and rela- tively low temperature resemble the climatic conditions in the south island of New Zealand, and there is a considerable degree of resemblance in the vegetation of the two regions. In common with New Zealand there is an important element of the flora closely related to, or even identical with, South American species. Some of these ‘‘Fuegian’”’ plants are found also in the parts of the adjacent state of Victoria and also as Alpines in the moun- tains further north. The most striking of these are the evergreen beeches (Nothofa- gus spp.)which are a notable constituent of the flora of southern Chile and also of New Zealand. These are the sole representatives of the Cupuliferae (oaks, beeches, ete.) found in Australasia. The visitor to Australia is immediately impressed by the prc- dominance of the Eucalyptus forest, and indeed this is the com- monest tree genus. While much of this open forest is extremely monotonous and unattractive, it must be remembered that among the more than two hundred species there are some of the stateliest and mést beautiful trees known anywhere. The great Karri forests of West Australia and the giant gum forests of Victoria, as well as some of the Eucalypts from the rich mountain forests 496 THE SCIENTIFIC MONTHLY of New South Wales and Queensland, are some of the most mag- nificent the writer has ever seen. In the spring, when the new foliage is developing, many species’ show beautiful golden and ruddy tints in the young leaves that are in strong contrast with the gray-green of the adult foliage of most species. In the arid regions of the interior there are dwarf species shrubs of moderate size remarkably resistant to drought. The flowers of some species of Eucalyptus are very beautiful and produced in great profusion. As in so many Myrtaceae the numerous stamens form the showy part of the flower and are pure white, pink or searlet in color. The splendid F. ficifolia with bril- liant scarlet stamens is a favorite ornamental tree in parts of Cali- fornia. The Myrtaceae, aside from Euealyptus, are very largely de- veloped in Australia, being second in number of species in the Australian flora,? more than eight hundred having been described. Allied to Eucalyptus are Tristanea, Angophora and Synearpia, all fine trees of large size. In the moister and warmer areas of the coast are members of the widespread genera Myrtus, Eugenia and Barringtonia, the latter entirely tropical in its habitat, a very beautiful tree with large glossy leaves and big white flowers. The genus is common throughout the Malayan region and the southern islands of Polynesia. More characteristically Australian and represented by many species are the genera Leptospermum and Melaleuca, very widely distributed and often forming extensive thickets. Some of the Melaleucas are small trees; the Leptospermums are as a rule shrubs of medium size. The flowers are usually white and produced in great profusion, so that some species are very attractive when in flower and prized as garden ornaments. Other characteristic Myrtaceae are the showy red ‘‘bottle-brushes’’—species of Callis- temon and the pretty fringed flowers of the West Australian Verti- cordias. First in number of species in the Australian flora is the great family of Leguminosae, with over one thousand species. Acacia leads with upwards of four hundred, ranging from tiny shrubs a few inches in height to large trees. The Acacias are popularly known as ‘‘wattle,’? and in the spring the profusion of golden bloom of many species makes them very conspicuous. Some of these Australian wattles are common in cultivation and often ealled ‘‘Mimosa.’’ The majority of the Australian Acacias are of the ‘‘phyllodineous’’ type, 7. e., the feathery leaf-lamina is sup- 2 Maiden, loc. cit., p. 166. VEGETATION OF AUSTRALIA 497 pressed and the flattened leaf-stalk, or ‘‘phyllode,’’ looks like a simple lanceolate leaf. The section Papilionaceae, or Pea family, contributes a host of showy flowers to the floral show. Nearly all of these exhibit very brilliant colors, pink, crimson, ‘scarlet, orange, yellow, blue and purple, and the flowers are borne in profusion. Many belong to strictly Australian genera—e. g., Chorizema, Gastrolobium, Jacksonia, ete., and comparatively few are in cultivation. The third family, in point of numbers, the Proteaceae, has not a single representative in the United States, and is almost entirely absent from the northern hemisphere. About two thirds of the species belong in Australia, and South Africa is next in number of species. The Proteaceae are mostly shrubs of moderate size, but a considerable number are arborescent, becoming forest trees. Of these trees, the most important are the species of Grevillea Banksia, Stenocarpus, Macadamia and several others peculiar to the rain-forests of Queensland. The handsome Grevillea robusta is a fine tree frequently seen in California, and a few other species of Grevillea and Hakea are less commonly seen in gardens; but many fine species, well worth cultivation, are still to be seen only in the wild. Grevillea is the largest genus and is widespread in Australia. The flowers are often very showy, pink, scarlet or yellow. Hakea, next in number of species, has as a rule rather inconspicuous flowers. Few Australian trees are more peculiar in habit than some of the Banksias, whose stiff serrate leaves and huge oblong heads of yellow flowers are most peculiar and striking. The great majority of the Proteaceae are xerophytic, but a few inhabit the scrubs of New South Wales and Queensland. Perhaps the finest flowers among the Proteaceae belong to the ‘‘Waratah’’ of New South Wales, previously referred to. Other important families, nearly or quite confined to Australia, are the Tremandraceae, Goode- niaceae, Candolleaceae and Casuarinaceae. Reference has already been made to some of the Australian Gymnosperms, which are extremely interesting. The Cycads have already been mentioned, as well as the Araucarias and Kauri of Queensland. The coniferous types of the northern hemisphere are absent, the nearest relation being the genus Catlitris, which is re- lated to the cypresses. The Yew family or Taxaceae, however, is remarkably developed in the southern hemisphere and has a number of extremely inter- esting forms in Australia and especially Tasmania. Podocarpus, of which a small number of species occur in the warmer parts of the northern hemisphere, is the most important Australasian genus, Vol. XV.—32. 498 THE SCIENTIFIC MONTHLY and comprises a number of large and valuable trees, as well as some smaller ones. Species occur in all the states. Certain genera, absent from the mainland, are found in Tas- mania and New Zealand. Such genera are Phyllocladus and Dacrydium, as well as several others. Ferns and their relatives are scarce or entirely wanting in a very large part of Australia, owing to the prevalence of arid and semi-arid conditions unsuited to these moisture-loving plants. There are, however, regions where they abound and are an important feature of the vegetation. The ubiquitous bracken- fern (Pteridium aquilinum) often covers large tracts of open land, as in northern regions; and in the moist gullies of the Blue Mountains of New South Wales and the forests of Victoria or in the rain-forests of the north there is a rich assortment of Pteridophytes, including some very fine treeferns, interesting Lyecopods and the curious Psilotum and Tmesipteris, whose life- histories, which long baffled the botanist, have at last been revealed through the labors of Lawson and Holloway. In the rain-forests are many epiphytic species, of which the extraordinary stag-horn ferns (Platyeerium) are the most con- spicuous; but there are also a good many of the beautiful and delicate filmy ferns (Hymenophyllaceae). NEw ZEALAND New Zealand comprises two large islands of about equal size and several adjacent ones of very much smaller dimensions. The northernmost point of the North Island is about 34° south latitude, and the South Island extends to south latitude 47°. The total area of the islands is about 100,000 square miles. New Zealand presents a marked contrast to Australia, both m its topography and climate. Its relatively small area results in a climate of distinctly insular character, with very much less range of temperature and precipitation than is the case in con- tinental Australia. Owing to its higher latitude, the climate as a whole is rather cool, but severe cold is rare in the lowlands. It is comparable with the climate of Britain, but especially in the North Island is considerably warmer. Owing to the proximity of the sea, there is less difference between North and South than might be expected. Thus between Auckland in the North Island and Invereargill, about ten degrees further south, there is less than ten degrees difference in the average temperature. For the most part rain is abundant and well distributed, and much of the country shows a luxuriant growth of forest. There are certain regions, however, notably the Canterbury Plain of the South Island, which have a relatively scanty rainfall and are VEGETATION OF AUSTRALIA 499 mostly destitute of trees. These grass-covered plains may be com- pared to the prairies of the mid-west of the. United States. The topography of New Zealand is for the most part exceed- ingly rugged, with much higher mountains than those of Australia. In the North Island are extensive voleanie formations, some of which are still active. In the Rotorua district, familiar to tourists, are numerous hot springs and geysers much like those of the Yel- _lowstone and in addition there are active voleanic craters. In the neighborhood of Auckland are a number of very perfect extinct cones, and on the west coast is Mt. Egmont, over eight thou- sand feet high. To the south lies the Wellington district, extreme- ly rugged in character. The harbor of Wellington, surrounded by steep mountains, opens into Cook’s Strait, separating the North and South Islands. The South Island shows less extensive evidences of voleanie activity than the North Island. It is distinguished by the lofty snow-clad range of the Southern Alps near the west coast, culmi- nating in the majestic Mount Cook, over twelve thousand feet high, snow covered for most of its height and with extensive glaciers reaching nearly to its base. The southwest coast is in- dented by numerous fiords, which are said to present a magnificent spectacle. The Southern Alps exercise a great influence on the climate of the South Island, intercepting a very large part of the moisture from the seaward side. Between the mountains and the coast there are stations with as much as two hundred inches of rain annually, while Christchurch on the east coast has only about twenty-five inches, and there are a few stations with even a lighter precipitation. This dry region is mostly destitute of trees, the ground being covered with coarse tussock-grasses. The contrast between these dry grasslands and the densely forested regions of rainy Westland is most striking. To the south of the great mountain range the conditions are more uniform, and the whole southern end of the South Island is covered with forest. The North Island originally was almost completely covered with heavy forest, in which the most important tree was the Kauri pine (Agathis australis). Very little of this splendid forest re- mains, and the Kauri is almost extinct. A few small tracts have recently been reserved, and I had an opportunity of visiting one of these in the extreme northern part of the island. This new park is of limited extent, but is a typical example of the magnificent Kauri forest which once covered the desolated regions now oe- cupying most of the surrounding country. The Kauri is entirely different in appearance from any conif- by S en) a z, S ~S <2) ~~ Ry ~ i Ss = DO WD Sa) Ry By KAURI FOREST, NORTH I LAND, NEW ZEALAND VEGETATION OF AUSTRALIA 501 erous tree with which the American botanist is familiar. In its younger stages it shows the symmetrical pyramidal habit of most conifers, but the early branches finally fall off, leaving a perfectly smooth cylindrical bole with very little taper. This columnar trunk may reach a height of sixty to eighty feet, or even more, with a diameter of eight to ten feet, or it is said of twice this size. At the top there are several enormous diverging branches forming an immense spreading crown which overtops the other trees of the forest and gives the tree a most characteristic appearance. The interior of the Kauri forest, with the huge smooth gray columnar trunks, is most impressive, and only rivalled by the great coniferous forests of the Pacific Coast, or the Cyptomerias of Japan. The New Zealand Kauri must rank as one of the giants of the vegetable kingdom. Associated with the Kauri are a number of other trees, inelud- ing several other Conifers or rather Taxads, as these belong to the Yew family. Of the latter the most important are the ‘‘Totara’’ (Podocarpus Totara) and ‘‘Rimu’’ (Dacrydiwm cupressinum), both valuable timber trees. The curious Phyllocladus trichomi- noides with flattened twigs (cladodes) looking like small fer::- leaves, is not uncommon in this region. Some other characteristic trees are Weinmannia sylvicola (Saxifragaceae), said to be the commonest tree in New Zealand, and Beilschmiedia tarare, be- longing to the Lauraceae. A number of fine shrubs, e. g., Coprosma, Pittosporum, Notho- panax and others are common, and as in all New Zealand forests ferns are much in evidence. The abundant and beautiful tree- ferns lend a special charm to the New Zealand forest. The finest of these is Cyathea medullaris, which may reach a height of up- wards of fifty feet and is the finest tree-fern with which I am ac- quainted. ‘Other interesting ferns are several species of Gleichenia, and the climbing fern, Lygodium articulatum, which is said to climb to the top of lofty trees. Filmy ferns (Hymenophyllaceae) are common in the damp shady woods, but are hardly as abundant or luxuriant as in the rain-forests of the South Island. Epiphytes abound in the rain-forests and include many con- spicuous mosses and liverworts as well as ferns and various flower- ing plants. Among the latter are a number of orchids, but these are mostly inconspicuous species, far inferior in beauty to some of the fine Australian epiphytic orchids. Perhaps the most conspicu- ous epiphyte is a very common liliaceous plant, Astelia solandera, forming great tufts of stiff sword-shaped leaves on the trunk or branches of many trees. It very often is seen on the slender stem of the Nikau palm, forming great bunches completely surround- 502 THE SCIENTIFIC MONTHLY Fic. 8. TREEFERNS, NEW ZEALAND VEGETATION OF AUSTRALIA 503 ing the trunk. This palm (Rhopalostylis sapida), is the only palm native to New Zealand. © Where the land has been cleared it is often invaded by the ubiquitous bracken (Pteridiwm aquilinum), and another plant which quickly takes possession is the ‘‘Manuka’’ (Leptospermum scopartum), closely resembling some of the Australian species. When in bloom the shrub is decidedly ornamental, with myriads of pretty white flowers. In open, more or less swampy districts, all over New Zealand, two extremely characteristic plants can hardly fail to attract at- tention. These are the native ‘‘flax’’ (Phormium tenax) and the ‘“eabbage-tree’’ (Cordyline australis), often grown in California under the name Yuecca-palm. The flax yields an abundant and valuable fibre, which is manufactured on an extensive scale and is one of the most important products of the country. At the time of my visit both of these striking plants were in flower. The flax sends up from its tuft of broad leaves, five or six feet high, scapes about twice as tall, bearing racemes of tubular red flowers which are much frequented by honey-sucking birds. The stately Yucea- like Cordylines bear immense panicles of small white fragrant flowers. Among the most widespread trees of New Zealand are several species of Metrosideros, a genus occurring throughout Polynesia and Australasia. M. robusta, the ‘‘Rata,’’ is a very beautiful tree with glossy green leaves and bright red flowers. Like Eucalyptus, to which it is not very distantly related, the bright-colored stamens are the showy part of the flower. Another species, M. tomentosa, also with showy flowers, is abundant about Auckland, and several other species, some of them climbers, occur in various parts of New Zealand. WM. robusta begins life as an epiphyte, the trunk of the older tree being made up of the united descending roots, as is so often the case in many species of Ficus. The southern portion of the North Island has a somewhat dif- ferent vegetation from that of the Auckland district. The Kauri is quite absent from this region, and in some districts there are forests of evergreen beeches much like those of the South Island. In the immediate vicinity of Wellington the forest shows many of the same trees as that further north, e. g., Podocarpus, Dacry- dium, Metrosideros, Weinmannia, Beilschmiedia and _ others. Among these is one of the two species of Proteaceae found in New Zealand. This is a handsome tree, Knightia excelsa, some- what resembling the Australian Banksias. One of the most interesting small trees is a Fuchsia, F. excorticata, New Zealand having three species of this otherwise ex- elusively South American and Mexican genus. 504 THE SCIENTIFIC MONTHLY Photograph by Dr. L. Cockayne. Fic. 9. CABBAGE-TREES (CORDYLINE AUSTRALIS) AND NEW ZEALAND FLAX (PHORMIUM TENAX) Shrubby Compositae are common in New Zealand. The genus Olearia, much like Aster, is well represented, some species having fine flowers, others like O. ilicifolia with handsome evergreen foliage. Another peculiar genus is Raouwlia, which includes the ‘‘vegetable sheep,’’ R. eximia. Some other characteristic shrubs noted near Wellington are species of Melicytus, Elaeocarpus, Myrsine and Sophora. SNS. tetraptera, with brillant yellow, very conspicuous flowers, is one of the few really showy New Zealand shrubs. Wellington has an attractive if not large botanical garden of which the most interesting feature is a small ravine in which are erowing many of the native trees and shrubs as well as some fine ferns and liverworts. Some of the treeferns are very tall and make a fine show. Of the liverworts the most notable is the re- markable Monoclea Forsteri, a giant among liverworts. This species is abundant about Wellington and also in various part of the South Island. The only other species known occurs in tropical America. In the botanical gardens in Wellington are some fine exotic conifers, including a number of Californian species. One of these, the Monterey pine (P. radiata) is extensively planted in New Zea- VEGETATION OF AUSTRALIA 505 land, where it grows with extraordinary rapidity and furnishes a large amount of timber. Across the harbor from Wellington at Day’s Bay is a consider- able extent of forest made up almost exclusively of two species of evergreen beeches (Nothofagus fusca and N. Menziesii). This forest is much more open than the mixed forest which prevails over most of the country near Wellington. These beeches, except for their much smaller leaves, are not unlike the true beeches of northern forests. They also remind one, in general habit, of the tree alders of the Pacific Coast. ‘Cook’s Strait, separating the North and South Islands, does not seem to form an appreciable barrier to the migration of plants between the islands, there being little difference in the vegetation on the two sides of the Strait. Probably the separation of the two islands took place at a comparatively recent date, so that there © has not been time for any marked change in the vegetation. The important city of Christchurch is surrounded by the famous Canterbury Plain, an open grassland which like our west- ern prairies is admirably adapted to agriculture. The trip across the South Island from Christchurch to the west coast is full of interest to the botanist and includes some magnificent scenery. I had the good fortune to be accompanied by Dr. L. Cockayne, the well-known botanist, whose knowledge of the native flora is both extensive and accurate.* The Canterbury Plain, where it has not been cultivated, is covered with tussocks of coarse grass, and this is true also of the lower slopes of the mountains on the eastern side. The most abundant species is Festuca Novae Zealandeae, but Poa caespitosa was another common and conspicuous species. The change from this open grassland to the first beech forest is very abrupt, and marks the beginning of the western rainy dis- trict. The beech forest is very dense, and composed exclusively of the mountain beech (Nothofagus Cliffortiana). At the summit of Arthur’s Pass, about 3,000 feet elevation, the increasing moisture becomes more evident. The open stony ground supports a heavy growth of herbaceous plants and low shrubs. In this formation is found perhaps the most beautiful of all new Zealand flowers, Ranunculus Lyallu. This fine plant has very large undivided peltate leaves, and clusters of pure white flowers two inches or more in diameter, borne on stout stalks a foot or more in height. Another charming flower is Owrisia macrocarpa, with large flowers something like Mimulus. 3 Dr. Cockayne’s book, ‘‘ New Zealand Plants and Their Story,’’ Welling- ton, 1919, is an admirable account of New Zealand vegetation. 506 THE SCIENTIFIC MONTHLY The sub-alpine serub of this region is composed of a number of very characteristic species. The most casual observer cannot fail to note the Dracena-like Dracophyllum Traversu, a small tree with clusters of reddish leaves at the tips of the straggling branches. In spite of its Yucca-like habit, this is a heath of the family Epacridaceae. Various shrubby Veronicas, a genus devel- oped to a remarkable degree in New Zealand, are abundant and several shrubby Compositae (Olearia, Celmisia, Senecio) abound. A curious leafless leguminous shrub, Carmichelia sp., 1s noted and a species of Gaultheria, and an Araliad (Pseudopanax lineare) are not uncommon. Along the roadside the mountain flax (Phormiwm Colensov) is frequent. The descent on the west side, through the magnificent Otira Gorge, is one of the finest pieces of scenery in New Zealand. The very steep walls of the gorge are densely covered with luxuriant forest from crest to base. The very heavy rainfall of this district is attested by the luxuriant rain-forest which reaches its maximum development on the west side of the range. At first there is some admixture of beech, but this finally disappears and in the typical Westland rain- forest is quite absent. The banks along the roadside show a constantly increasing pro- fusion of ferns, liverworts, and moisture-loving herbs, like violets, Hydrocotyle and the interesting Gunnera, a genus particularly de- veloped in New Zealand. ‘ Tree-ferns, which had not been seen at the higher elevations, inerease in size and numbers as the lowlands are approached and in the lowland forest form a conspicuous and beautiful feature. The Westland rain-forest is one of extraordinary luxuriance. The extremely heavy precipitation and mild temperature result in a rich profusion of vegetation that has all the aspects of a genuine Malayan rain-forest. Composed of exclusively evergreen trees and shrubs, draped with lianas and epiphytes and interspersed with thousands of noble tree-ferns, it was hard to believe that this forest was in latitude 43°, corresponding in the United States to the latitude of Buffalo. This forest is of the type called ‘‘Taxad’’ by Cockayne, the most important trees belonging to the taxaceous genera Podocarpus and Dacrydium. In ‘the swampy areas the ‘‘white-pine’’ P. dacrydioides predominates, a very tall tree with fine straight trunk yielding valuable timber. Of the angiospermous trees the most abundant is Weinmanma sylvicola, a tree of Malayan affinity, and its relative, Quintima acutifolia, both belonging to the Saxi- frage family. A very common large shrub is Aristotelia racemosa, with rather attractive pinkish flowers, and other common shrubs are species of Coprosma, Metrosideras lucida, and Pseudopanax crassifolia. VEGETATION OF AUSTRALIA 507 Ferns and other Pteridophytes abound in these wet forests. Various species of Lycopodium are abundant, and also the curious Tmesipteris. Of the tree-ferns, the commonest is Dicksonia squarrosa, sometimes twenty to thirty feet high. Less abundant is Hemitelia Smithu. Of the abundant epiphytic growths the most beautiful are the filmy ferns (Hymenophyllaceae) which in these Westland forests attain their finest development. Another ex- tremely beautiful fern is Todea (Leptopteris) swperba. As might ‘be expected, these supersaturated forests are a veri- table garden of mosses and liverworts which drape the trees and form a thick carpet on the ground and big cushions over every prostrate log and stump. Great tussocks of Sphagnum are com- mon about the pools, and now and then one encounters colonies of the giant Dawsonia superba, the last word in moss development. The abundance and luxuriance of the liverworts is astounding ; indeed, it is doubtful if anywhere else in the world is a richer growth of these interesting plants to be found. Of the lanas the most interesting is Preycinetia Banksit, a distinctly tropical genus belonging to the secrew-pine family, abun- dant throughout Polynesia and the Malayan region. Everywhere in the New Zealand rain-forests there is a rich development of epiphytes and climbing plants. Of the epiphytes there are two eategories, those that begin life as epiphytes, but later become rooted in the ground, and those which retain perma- nently the epiphytic habit. Among the latter are many ferns, Lycopods, Mosses and liverworts, as well as a good many flowering plants like Peperomia, various orchids, Astelia, ete. Of the ferns the filmy ferns or Hymenophyllaceae are especially numerous and beautiful. Several species of New Zealand trees begin life as epiphytes. The seeds germinate in the branches of some tree and presently the young plant sends out roots which descend the trunk of the host-tree until they reach the earth. In course of time these de- scending roots coalesce in a more or less solid trunk and the host- tree may be completely strangled in the process. The ‘‘rata’’ (Metrosideros robusta) is the best known of these temporary epiphytes. Others are Dracophyllum arboreum and Griselinea littoralis. Some of the climbing plants are great woody lianas, whose stout cables are thrown from tree to tree. One of the biggest of these is a huge bramble, Rubus australis, whose stems, sometimes six inches in diameter at the base, reach the tops of the tallest trees. Freycinitia climbs by means of roots, clinging to the trunks of trees, and some species of Metrosideros have a similar habit. Other common climbing plants are species of Clematis, Par- 508 THE SCIENTIFIC MONTHLY sonsia and Miihlenbeckia and the climbing fern Lygodium articu- latum. Compared with Australia there is a remarkable scarcity of brilliantly colored flowers, a large proportion of the plants having white or greenish flowers. The bright red flowers of some species of Metrosideros, Clianthus puniceus and the native flax, bright yellow of Sophora tetraptera, the blue of many Veronicas and the blue or purple of some of the Compositae are the most marked exceptions to the rule. WEEDS As in all countries where the white man has settled there have come with him many plant immigrants, some of which are not entirely weleome. These weeds hail from many lands. In the hotter and dryer parts of Australia they may come from such tropical countries as India, Brazil or Africa, while in the more temperate regions of southern Australia and New Zealand they are largely from northern Europe and America, e. g., thistles, sorrel, dock, plantain and other familiar weeds. Parts of Australia have been invaded by species of prickly pear (Opuntia) from America, which are a very serious pest. It has been said that in Queensland 30,000,000 acres of land have been invaded by one species which has caused immense damage. From America have also come species of cockle-bur (Xanthium) and Stramonium, as well as several other pestilent weeds. In the moister cooler regions of Australia and New Zealand, the common European blackberry, sweet brier and gorse have escaped from cultivation and become very persistent and troublesome weeds. A number of plants from the Cape, whose climate is very sim- ilar to that of Australia, have become completely naturalized. It is not uncommon to see the familiar calla growing in ditches and low ground, and several of the beautiful Iridaceae from South Africa—Ixia, Sparaxis, Watsonia and Homeria—very often are seen growing along the railway embankments. The latter is said to be poisonous and may perhaps be called a weed. This name may also be given to the ‘‘Cape-weed’’ (Cryptostemma calendu- lacea), a daisy-like Composite which is extremely abundant. CoNCLUSION Attention has already been called to the evident close relation- ships existing between the Australian ‘‘scrub’’ floras and those of the Indo-Malayan regions. The rain-forests of Queensland and New South Wales may be looked upon, with little question, as the remnants of a much more extensive flora which occupied these regions when they were united with New Guinea and separated VEGETATION OF AUSTRALIA 509 from the ancient Western Australian continent. It is generally believed that in the latter, which was probably much larger than at present, the ancestors of the characteristic types which now dominate the flora of the greater part of modern Australia had their origin. In the old Western continent, completely isolated from other lands, there was an extraordinary development of a comparatively small number of families. The most conspicuous examples of this are the Myrtaceae, with over 200 species in Eucalyptus alone; Leguminosae, especially Acacia with over 400 species, and many peculiar Papilionaceae; Proteaceae with over 600 species (Gre- villea, Hakea, Banksia, ete.). A few families, e. g., Candol- leaceae, Goodeniaceae, are almost exclusively Australian and es- pecially abundant in Western Australia. These peculiar Australian plants are largely xerophytic, and after the union of Eastern and Western Australia it may be as- sumed that the extreme aridity and poor soils of much of the central part of the continent would be much more favorable to these Western xerophytes than to the Malayan types of the Hast which have evidently been largely evicted by the drought-resistant West Australian immigrants, and are now restricted to compara- tively limited areas where there is good soil and abundant mois- ture. The autochthonous types have for the most part remained in Australia. Eucalyptus, Acacia, a few Proteaceae and some others are represented in the savannahs of Southern New Guinea and the dryer portions of the Malay Archipelago; and a few genera range through Polynesia; but the great majority of the true Australian species are unknown outside the Australian continent. In the southeast, and especially in Tasmania, there is a marked infusion of plants whose relationships are with the Andean and Fuegian vegetation of South America. Most of these occur also in New Zealand. . Comparing New Zealand with Australia, there is found a good deal in common in the floras of the northern districts, 7. e., the Malayan rain-forest vegetation. This type is, however, of very much greater importance in New Zealand, where in spite of a much cooler climate, a large proportion of the trees and shrubs are more _or less closely related to Malayan ones. There is strong evidence of former connections with the tropical regions to the North, and it is quite as likely that the Malayan genera which New Zealand shares with Australia have been de- rived from the North and not directly from Australia. The distinctively Australian genera are relatively few in New Zealand, and a striking feature of the flora is the absence of such 510 THE SCIENTIFIC MONTHLY predominant Australian genera as Eucalyptus and Acacia. The family Myrtaceae, with over 800 species in Australia, has barely twenty in New Zealand, only one genus, Leptospermum, being typically Australasian. The Proteaceae, which reach their maxi- mum in Australia, with more than 650 species, have only two representatives in New Zealand. There are, however, a consider- able number of Epacridaceae, and several Australian genera of orchids, as well as Compositae and Leguminosae. It has been suggested, however, that some of these forms might be of New Zealand origin and migrants into Australia. Most of the ferns common to the two countries are widespread Austrahan-Malayan species, but mention should be made of one, viz., Todea barbara, common to northern New Zealand and New South Wales and also found in South Africa. The Fuegian genera already referred to are mostly shared by Australia and New Zealand. Of these there are twenty-two genera common to the two countries, among which may be mentioned Astelia, Muehlenbeckia, Drimys, Nertera and the evergreen beeches, Nothofagus.* There are sufficient resemblances between the floras of Austra- lia and South Africa to indicate some former land connections between the two, but it is probable that the connection was severed at a very remote period. In South Africa, as in Australia, there is a remarkable devel- opment of Proteaceae, but there are no genera common to the two, indicating a very long period of separation. The true heaths (Ericaceae) which are a marked feature of the South African flora are replaced in Australia by the Epacridaceae. It has been suggested® that the two families are offshoots of a common stock, differentiated since the disappearance of a former land connection. It is pretty well agreed that at one time all the great southern land masses were connected more or less completely. The name ‘Gondwana Land’’ has been given to an assumed great southern continent, existing in late Permian time, and embracing a large part of the present South American, African and Australian conti- nents, as well as parts of India and Malaya. Just how long these connections remained is not entirely clear, but if they persisted into the Cretaceous, or early Tertiary, this would explain many of the apparently anomalous facts of the present distribution of the floras of the southern hemisphere. We have also to take into account the great antarctic conti- nent. At present this is practically destitute of any terrestrial 4 Cockayne, loc. cit., p. 206. 5 Maiden, loc. cit., p. 181. VEGETATION OF AUSTRALIA 511 vegetation, but there have been found fossils indicating the former presence of a vegetation related to that of South America and New Zealand. Further investigation may show that there was a north- ward extension of the present antarctic continent, with climatic conditions much more favorable for vegetation, than exists at present. If further discoveries of fossils should show that, as in the northern hemisphere during the Tertiary, there was also in the south a practically uniform flora encircling the globe, this would make comprehensible both the resemblances and differences now existing in the floras of the southern land-masses. Migrants from this common southern flora later completely shut off in the present widely separated countries, would in course of time show greater or less divergence from each other, depend- ing upon the amount of change in their environment. It might be expected that in the cool humid elimate of New Zealand, the evolution of the primordial southern types would be very different from those subjected to the hot and arid conditions of Western Australia, which is supposed to be the birthplace of most of the strictly Australian plant-types. 512 THE SCIENTIFIC MONTHLY EASY GROUP THEORY By Professor G. A. MILLER UNIVERSITY OF ILLINOIS HE report that the title of the chair of ‘‘differential and fi integral caleulus’’ in the University of Paris has re- cently been changed to ‘‘the theory of groups and the eal- culus of variations’’ may tend to create a desire on the part of a larger group of scientific men to understand the essence of a mathematical group and the role which the group concept is assuming in modern mathematical develop- ments. The fundamental importance of the concepts of differential and integral calculus in various fields of science has long been recognized, and the change of title noted above does not imply that the theory of groups and the calculus of variations tend to supplant the differential and integral caleulus. It does, however, imply that the former subjects are also sufficiently fundamental and fruitful to merit prominent recognition in a leading mathe- matical center. Paris may justly claim a very large share in the early develop- ment of group theory. A Parisian, A. L. Cauchy, is commonly regarded as the founder of this subject, while other Parisians, in- eluding A. T. Vandermonde and E. Galois, did important pioneer- ing work in this field. Although J. L. Lagrange is commonly re- garded as a French mathematician his pioneering work in group theory was done before he settled in Paris. The first separate treatise on this subject was written by a Parisian. This work ap- peared in 1870 under the title Traité des substitutions et des équations algébriques by C. Jordan. In group theory as well as in differential and integral caleulus the first extensive formal or abstract developments are due to Eng- lish and German mathematicians. In the case of the latter subjects the authors of these developments, viz., Newton and Leibniz, are commonly regarded as its founders. Fortunately this has not yet been done as regards the former subject. Notwithstanding the fundamental importance of formal developments and _ abstract formulations the real life of mathematics abides in its contact with the conerete. In the case of group theory this contact has been em- phasized especially by S. Lie, F. Klein and H. Poincaré. The rapidly increasing prominence of this theory during the last half EASY GROUP THEORY 513 century is largely due to the writings of these three men, who have been leaders also in various other lines of mathematical ac- tivity. A dominating mathematical concept, like a dominating per- sonality, has a charm of its own, and creates a kind of atmosphere which is as invigorating as that of a real university or that which emanates from any group of real scholars. A fundamental notion of a mathematical group is that ‘‘there is no new thing under the sun.’’ It is true that we speak here of generators and generational relations, but the objects which are generated were members of the group since the beginning of time and will remain members thereof throughout eternity. We study the group to perceive this same- ness in its various forms and to understand the relative properties of the various elements which unite to make a single element of the group. The non-technical meaning of the term group suggests little as regards its technical meaning except the invariance of the number of its elements. In a non-technical group one usually thinks of the elements as units which may or may not have the power of reproduction. In the former case the elements thus produced are usually new elements of the group. In the technical group the elements have necessarily the power of uniting, but when they unite they neither produce any new element nor lose their own identity. The union merely exhibits the possible decomposition of an element of the group, or the different ways of securing e pluribus unum. Union, unity and stability constitute the trium- virate of the theory of groups. The stability here noted is not the stability of statics but the stability of dynamics. It is a kind of invariance under transformations. What is perhaps of most interest in this connection as regards the non-mathematician is the question why the concept to which we referred above is so fundamental in mathematics. It is well known that mathematical developments have been largely guided by the desire to secure intellectual penetration into the workings of nature. Do we find in nature numerous instances of the union of elements of the same kind to produce an element of this kind which is really not new but belongs to a totality which has been clearly defined? For instance, one may think of the totality of the transformations of space subject to the condition that the distance between every pair of points in space remains invariant. It is evident that if one such transformation is fol- lowed by another the two together are equivalent to a single trans- formation of the totality in question. Hence we say that this totality constitutes a group. Vol. XV.—33. 514 THE SCIENTIFIC MONTHLY It is also clear that the totality of the natural numbers when they are combined by addition has the property that no new num- ber arises from the combination of any two of them, or from the combination of one with itself. In both this totality and in the totality of transformation noted in the preceding paragraph it is evident that if x, y, zg represent three elements of the totality and if any two of them are supposed to be known then the third is completely determined by the following equation: cy=2 The term group is commonly used in mathematics in the restricted sense that the third of any three elements is completely determined by such an equation where any two of them are supposed to be given. Moreover, it is usually assumed that when any three ele- ments are combined the associate law is satisfied, but it is not assumed that the commutative law is necessarily satisfied when two of them are combined. Even when these restrictions are imposed there are instances of groups almost wherever one turns. It is true that in the vege- table and the animal kingdoms one sees new elements arising in pro- fusion, and mathematics is naturaliy called on to deal also with such conditions, but if one looks deep enough here there seems to be a union of elements without loss of identity of the elements, and there seems to be nothing new in the profound formal physical sense. Hence one may see some significance in the following state- ment made by Poincaré shortly before his death: ‘‘The theory of groups is, so to say, entire mathematies, divested of its matter and reduced to a pure form.’’ In the groups noted above the number of elements is infinite. As instances of a finite group we may consider the six movements which transform a fixed equilateral triangle into itself and the eight movements which transform a fixed square into itself. Such special instances are, of course, of little interest to the general scientist except as illustrations. The thing that may be supposed to command the interest even of the educated layman is the fact that these very evident considerations appertain to a profound mathematical theory. Group theory did not arise from such obvious considerations. After it was partially developed as an autonomous science some of its more obvious applications received special attention. A certain degree of difficulty seems to create the most favorable atmosphere for scientific developments. In group theory this atmosphere was created by the n roots of the algebraic equation of the following form: Ce = det eg 0 EASY GROUP THEORY 515 It is customary to speak of x as the unknown and to call this an equation of degree m in one unknown. As a matter of fact the equation has n roots and all of these are unknown, so that it is really an equation in n unknowns. The constant coefficients a,, @,... a, were known to be symmetric functions of these n unknowns be- fore the subject of group theory was developed. The mysterious appearance of » unknowns to take the place of the one which presents itself openly is perhaps sufficient to create an atmosphere suitable for scientific endeavors. At any rate, it was in this atmosphere that our subject arose and hence we shall note here a few of the early steps in its development. It is true that these steps were taken by men who seemed to have no idea that they were dealing with notions which had the widest applications in other fields of mathematical endeavor. In fact, the early ex- plorers of group theory died long before any one realized that the notions with which they were dealing were destined to permeate a large part of the science of mathematics. The n roots of an equation of degree n with constant coefficients constitute a group in the non-technical sense of the term, but the group of this equation is something very different and hes much deeper. It relates to a certain totality of permutations of these nm roots, or substitutions on these n roots, leaving invariant every possible rational function of these roots which is equal to a con- stant, and having the property that if a rational function of these roots is invariant under these substitutions it is equal to a con- stant. These fundamental properties of the n roots of the equation in question were first noted by E. Galois, a French mathematician of great renown, although he died before reaching the age of twenty-one years. They were sufficiently difficult to create an atmosphere suitable: for the development of our subject as an autonomous science. » A study of the permutations of 7 things might at first appear to promise little of importance. It is true that before the time of Galois attention had been called to this subject by Lagrange and Vandermonde in connection with the question under what condi- tion a rational function on n variables can be expressed rationally in terms of another such function, but it was not until long after the days of Galois that mathematicians began to realize the funda- mental importance of this subject in the study of a large variety of mathematical questions. When the substitutions arising from these various permutations were studied by themselves they were seen to combine according to laws which are found almost every- where when the data are sufficiently connected to admit mathe- matical treatment. 516 THE SCIENTIFIC MONTHLY The reader who has given little attention to mathematical de- velopments may be inclined to ask, If the notion group is so fundamental why did the ancients and even such eminent later thinkers as Descartes and Newton pay no explicit attention to it? Why does the theory of groups not have an ancient prototype like differential and integral calculus, whose prototype is found in the method of exhaustion of the ancient Greeks? Does it appear rea- sonable that a subject founded only three quarters of a century ago should really deserve such a dominating position in modern mathematics as is claimed for group theory in what precedes the present paragraph? Does the history of mathematics present any other instance of the sudden rise of a dominating concept extend- ing into practically all the large branches of mathematics? The fact that the last of these questions must be answered in the negative tends to enhance the interest in the others. This negative answer calls also for a word of caution for there is danger that the reader might infer from it that the subject of group theory has greater merit than really belongs to it. It seems that mathe- matical developments have always been guided by the group con- cept. In the words of Poincaré ‘‘the ancient mathematicians em- ployed groups in many cases without knowing it.’’ The main question in mathematical developments is that one gets on the right road. The ability to explain why one has chosen this road is of secondary importance. In fact, the best teacher is frequently unable to give a good account of his methods while a much in- ferior teacher may be able to talk glibly about methods. Group theory is largely a method and those who are studying this subject by itself may be compared with those who are devoting their attention to methods of teaching. Just as the latter are not necessarily the best teachers so the former are not necessarily the best mathematical investigators. Possibly the bacteria which have tended to make the teachers colleges such a prominent feature of our modern universities have also caused the emphasis on group theory in modern mathematics. Just as some of our best teachers have never read a work on methods of teaching so some of the best mathematical investigators have never secured a speaking acquaint- ance with the notion of group. In both cases the real essence of ‘the subject has been acquired unconsciously, or, at least, without the development of a formal language relating thereto. The fact that modern mathematicians emphasize the group con- cept while the ancient and medieval mathematicians did not do this does not imply a change of mental attitude. Naturally the modern mathematicians secured a somewhat deeper insight into various subjects and thus discovered evidences of groups EASY GROUP THEORY 517 which were unknown to the older mathematicians. Inspired by these groups they developed also the theory relating to the groups which the ancients used implicitly, but it is questionable whether the modern mathematicians would have developed a theory re- lating to the latter if they had not been inspired by the former. At any rate, they did not take any steps towards such a theory before they had this additional motive. These observations may serve as partial answers to questions raised above relating to the late development of our subject as an autonomous science. The heading of the present article suggests that some of the de- velopments of our subject can not be properly called easy. In fact, by far the larger part of these developments presuppose a rather extensive technical knowledge and hence they are unsuited for a popular article. Among all the scientists the mathematician works usually at the greatest distance from his postulates, and hence he has the greatest difficulty to exhibit the results of his toil to the public in the hope of securing appreciation, which he craves with the others of his fellowmen. In group theory this distance has become especially long even for a mathematical subject, but this theory does extend also into the experiences of all thoughtful per- sons. The present article aims merely to direct attention to the richness of the mathematical developments which have contact with these particular experiences and thus to secure an easier approach to some of this richness. If a group contains a finite number of elements this number is called the order of the group. For instance, the 24 different move- ments of space which transform a cube as a whole into itself but interchange some of its parts constitute a group of order 24. The most elementary group of a given order g is cyclic; that is, it is composed of the powers of a single one of its elements. For instance, the g numbers which satisfy the condition that the gth power of each of them is equal to unity constitute this group of order g, where g is any natural number. It is evident that the 24 different movements which transform a eube into itself are not powers of a single one of them. Hence they constitute a non- eyche group of order 24. In fact, none of the elements of this group has to be raised to a higher than the fourth power in order to obtain unity, or the identity. One of the fundamental problems of abstract group theory is the determination of all the possible groups of a given order g. It was noted above that there is one and only one eyclie group of every possible order g. When g is a prime number there is no other group of this order. This is also sometimes the case when g is composite but there is no upper limit to the number of groups 518 THE SCIENTIFIC MONTHLY which may have the same composite order. That is, it is always possible to find a number g such that the number of the different. abstract groups of order g exceeds any given finite number. In addition to the two groups of order 24 noted above there are 13 others, which were first completely determined in 1896 by the pres- ent writer. The lowest order for which the number of groups exceeds the order is 32. There are 51 distinet groups of this order. The verification of several of these statements would carry us beyond easy group theory. They may serve, however, to exhibit a type of inquiry relating to our subject. Fortunately, some of the most important and far-reaching phases of this subject are also the easiest. For instance, the group of all the transformations of space which leave invariant the distances between every pair of points can easily be comprehended. Those geometric figures which can be transformed into each other under this group may be called equivalent and we may confine our attention to the study of geometric properties which remain invariant under the transforma- tion of this group. We thus obtain a body of knowledge commonly ealled Euclidean Geometry, but this term is also often used with different meanings. Following the custom introduced by Gauss some still use it to denote all the geometry in which the parallel postulate is assumed. It is clear that in geometry it is undesirable to endeavor to study every figure as an individual since one could not make much progress in this way. What people have always done in this sub- ject is to confine their attention to invariants under certain infinite groups of transformations. It is true that the ancients did not specify these groups and that we do not usually do this now in a first course in elementary geometry, but for the advanced student, at least, the developments become much clearer if this specification is explicitly made. If we add to the transformations noted above those which do not preserve the size of the figures but do preserve their angles, so that all similar figures are regarded as equivalent, we obtain a larger group, which has been ealled the principal group of geometry. The body of knowledge relating to the invariants of this group is commonly known as Elementary Geometry. In particular, all circles are equivalent in this geometry and all squares are also equivalent here. Some writers eall this geometry Euclidean Geometry. This is done, for instance, on page 61, volume 2, Paseal’s Repertorium der hoheren Mathematik, 1910. These observations relating to the groups of geometry may serve also to support the implication noted above that group theory is often a kind of mathematical luxury. One can frequently get along without a knowledge of this subject where a knowledge EASY GROUP THEORY 519 thereof would add greatly to the intellectual comfort. The fact that the mathematical world traveled far without making explicit use of this subject which now receives so much emphasis in work closely related to that which they were doing is perhaps best ex- plained by viewing the matter from this standpoint. It need searcely be added that group theory has also served to point out the way to easier methods of attack and to more powerful means of penetration, but this applies more especially to the more difficult group theory and hence lies outside the domain to which the head- ing of the present article relates. In the opening sentence of this article we alluded to the fact that in the University of Paris there is now a chair entitled ‘‘the theory of groups and the calculus of variation.’’ This should not be construed to mean that the developments of these two subjects have as yet much in common. In fact, there are few large mathe- matical subjects whose developments exhibit as little explicit use of group theory as those of the ealeulus of variations. Possibly the title noted above indicates that there will soon be a change in this direction. This title also raises the question whether our larger American universities should not have more chairs devoted to special subjects. The creation and occasional renaming of such chairs would tend to direct attention to leading investigators in various fields, an attention which often needs cultivation on the part of administrative officers. 520 THE SCIENTIFIC MONTHLY THE HISTORY OF THE CALORIE IN NUTRITION By MILDRED R. ZIEGLER (FROM THE SHEFFIELD LABORATORY OF PHYSIOLOGICAL CHEMISTRY IN YALE UNIVERSITY, NEW HAVEN, CONN.) See nomenclature of a science is of vital importance; it is intimately bound up in the subject itself. Special technical words are used to describe the phenomena which are being studied. Lavoisier emphasized the importance of nomenclature in his ‘“Traité Elémentaire de Chimie’’ (1789) when he stated: ‘‘ Every branch of physical science must consist of three things; the series of facts which are the objects of the science ; the ideas which repre- sented these facts, and the words by which these ideas are ex- pressed. Like three impressions of the same seal, the word ought to produce the idea, and the idea to be a picture of the fact . . .; we can only communicate false or imperfect impressions of these ideas to others, as long as precise terms are lacking.’’ The calorie as defined in the science of nutrition is a measure of food value. The significance of the word calorie as thus used to-day is not the same as its import when first introduced into the French language. To the student of nutrition the word connotes something quite different from the term as employed by the physicist. It is the amount of a food substance which on combus- tion in the body will yield energy—heat or work—equivalent to a calorie as understood by the physicists. Even the term as em- ployed by the latter to-day has undergone a change from its orig- inal meaning (1845). As first defined the calorie represented the amount of heat required to raise one kilogram of water through one degree centigrade. It is well known that the same term now means the amount of heat required to raise one gram of water through one degree centigrade. The calorie, then, has had—and still does have—two somewhat unlike meanings according as the gram or kilogram of water is the unit of mass, the temperature of which is being changed and the energy required for this change is being considered. Both of these meanings are preserved in the scientific literature of to-day, the larger unit being designated as a large calorie and written Calorie. HISTORY OF THE CALORIE 521 It is quite evident that either the small calorie or the large Calorie may be the more convenient unit with which to express heat change depending upon the magnitude of the change being studied. The physicist in expressing the amount of energy sup- plied in the form of heat required to raise one gram of water from its freezing point to its boiling point obviously is dealing with a relatively small energy change which if expressed in heat units, as discussed above, is in the neighborhood of one hundred calories. In such eases the little calorie is the convenient unit to employ; on the other hand experiments in the field of nutrition have demonstrated that the energy changes are of such magnitude that the Calorie is the more convenient unit for expressing them quantitatively. The derivation of the word is very interesting ; it has developed from the old French word ‘‘ealorique’’ which was derived from the Latin term ‘‘calor.’’ Lavoisier introduced ‘‘calorique,’’ which was defined at that time as an elastic fluid containing the hidden cause of the sensation of heat and to which the phenomena of heat were attributed. This meaning was abandoned with the theory to which it belonged. As far as a review of the literature reveals Bouchardat (1845) was probably the first to define it according to the modern physical conception. In his ‘‘Physique élémentaire”’ we read: ‘‘Unité de chaleur—On designe sous le nom d’unité de chaleur ou de calorie celle qui est nécessaire pour élever 1 kilogram d’eau d’un degré du thermométer centigrade.”’ The idea of measuring heat in this way had been in general use for some time, but the word ealorie had not been introduced earlier. Pouillet (1832) in his ‘‘Physique’’ defined the heat liberated by combustion of different substances as ‘‘Elévation de temperature que 1 gr. de chaque substance en se brilant avec l’oxigéne com- muniquerait a 1 gr. d’eau.’’ He ascribed the value 6195° to aleohol which checks closely with its caloric value as now known. Although the French had coined the word in the year 1845, it apparently was not in general use for some time; for in 1852 Favre and Silbermann! in an article on heat wrote: ‘‘We shall repeat that the unit which we have adopted is that adopted by all physicists, that is, the quantity of heat necessary to raise 1 gram of water 1 degree and which they eall unit of heat or calorie.’’ The Germans evidently took the word from the French, but it is difficult to determine at what time. Gmelin (1817-19) in his 1 Favre and Silbermann: Ann. de chim. et phys., 1852, Ser. 3, xxxiv, 357. 522 THE SCIENTIFIC MONTHLY ‘‘Handbuch der Chemie’’ uses the same idea as the calorie unit but refers to it as did Pouillet. As late as 1871 Senator? deemed it necessary in an article on heat production and metabolism to define the word ‘‘calorie’’ in a footnote. According to a contributor to Murray's ‘‘New English Dic- tionary on Historical Principle’’ (1888), the word calorie was first introduced into English in 1870 by T. L. Phipson in his trans- lation of ‘‘The Sun’’ by the French astronomer, Amédée Guillemin. Before this time the English had spoken of heat units which re- ferred to Joule’s mechanical equivalent of heat, viz., 1 kilog. of water raised 1°C—423 metrekilogs. This was the term used in Frankland’s® classic treatise ‘‘On the Origin of Muscular Power.’’ The derivation of the word calorie has not revealed its mean- ing; for that it is necessary to consider the history of animal heat, combustion and the potential energy of the foodstuffs. To the old scientists animal heat offered a difficult problem, surrounded with much mystery. They could not explain it by any known chemical or physical laws. They did not assign any cause to it, but de- seribed it as an innate quality, something ‘‘vital’’ situated in the heart and distributed to the body by the blood vessels (Plato, Aristotle, Galen). The principal function of the blood was to distribute the heat, while the great function of respiration was to cool this distributing medium. Mechanical and chemical notions were accorded to heat production. As a history of the calorie is concerned only with the chemical theories, the mechanical ones deseribed by Haller (1757)*, Boerhaave (1709)*° and contempo- raries will not be discussed here. Willis (1670)* was probably the first to consider an idea of combustion in heat production. He said that there was a ‘‘combustion’’ in the blood dependent upon fermentation excited by the combination of different chemical sub- stanees. All the chemists of the time considered animal heat a product of a ‘‘fermentation’’ occurring in the blood while in the heart. Willis’ term ‘‘combustion’’ was not in accord with the modern conception in which oxygen is essential, as this element had not been discovered. A more correct opinion was enunciated by Mayow (1674)* who had experimented on the elements of the 2 Senator: Centralbl. f. d. med. Wiss., 1871, IX, 737, 753. 3 Frankland: London, Edinburgh, and Dublin Phil. Mag. and Jour. of Sc., 1866, xxxii, 182. 4 Haller: ‘*Hlementa Physiologie,’’ 1757. 5 Boerhaave: ‘‘Aphor. cum Notis Swieten,’’ pp. 382-675. 6 Willis: ‘‘De Accensione Sanguinis,’’ 1670. 7 Mayow: ‘‘Tractatus Quinque,’’ Oxonii, 1674. HISTORY OF THE CALORIE 523 air and described a ‘‘nitro-aerial spirit’’ (oxygen). He held that the function of respiration was not to cool the blood but to enable this fluid to absorb nitro-aerial gas for generating heat. Mayow’s theory did not appear to make any considerable impression upon his contemporaries. It was not until Joseph Black’s (1755)® ex- periments on fixed air were performed that more correct ideas prevailed. He showed that the gas expired from the lungs is the same as that produced by combustion of fuel, thus establish- ing a relationship between combustion and respiration. Black was forced to relinquish his theory by his contemporaries who claimed that if the lungs were the sole seat of combustion the amount of heat produced therein would be so great that the vitality of the organ would be destroyed. The production of heat by a combustion of foodstuffs was merely suggested at this time (1677) with no air of conviction that such could be the ease. It appears that Descartes should re- ceive the credit for first suggesting the correct theory regarding heat production in the animal body in his ‘‘De Homine’’ (1662). He thought that the change produced in the food in the stomach was analogous to the heat produced when water is poured upon lime or aqua fortis on metals. Hunter (1761) in his dissertation ‘On Blood”’ incidentally remarks that ‘‘the source of heat is in the stomach.’’ He had previously expressed dissatisfaction with all prevailing theories of animal heat. Hunter’s idea is to be re- garded merely as his suggestion, with as yet no clear-cut data to justify its assumption of the rank of an established theory. It remained for Lavoisier, the father of modern chemistry, to prove by his carefully performed experiments that animal heat is not caused by any mystical ‘‘vital force’”’ as the ancients believed, but is a phenomenon analogous to the burning of a candle, namely, the combustion of carbon. He repeated, verified and added ex- periments to those of Black and Priestley, and explained more ex- plicitly than had ever been done before the source of animal heat. In 1780 in his ‘‘Mémoire sur la Chaleur’’ published in collabora- tion with Laplace he gave in detail the theory of heat essentially as we have it to-day. By means of the calorimeter the heat evolved during the formation of a definite quantity of carbonic acid was compared with that produced during the formation of the same amount of this compound during respiration. These investigators saw that in such transformations identical amounts of heat were produced. They concluded that animal heat is derived from the oxidation ‘of the body’s substance. 8 Robison: ‘‘Joseph Black,’’ Edinburgh, 1803. 524 THE SCIENTIFIC MONTHLY The publications of Lavoisier’s time and before that had spoken of heat as ‘‘chaleur.’’ Pouillet (1832), the physicist, remarked: *“Scientists confused the cause of heat with its effect. They called it chaleur, fluide igné, matiére du feu. Finally in reforming the chemical nomenclature Lavoisier, Bertholet, Morveau and Foureroy have called it calorique. This term was adopted by the physicists of the time; and they reserved the word ‘chaleur’ to designate the science which treats of the properties, effects and laws ‘du calorique.’’’® An interesting use of ‘‘calorique’’ is found in the ‘‘Mémoire sur la Respiration des Animaux’’ published in 1789 by Lavoisier and Seguin. They state, ‘‘1. Que le calorique (matiére de chaleur) est un principe constitutif des fluides (Sous ce nom générique nous comprenons les airs et les gaz.) et que ec’est a ce principe qu’ils doivent leur état de’expansibilité, leur élasticité, et plusieurs autres des propriétés que nous leur connaissons.’’!° There is no reference to a heat unit, calorie, in Lavoisier’s publiea- tions. He defines the amount of heat in terms of the weight of ice melted in a given experiment. Although the early physiologists suggested the correct source of animal heat, it is significant that they considered heat a simple chemical reaction, as stated by Deseartes. Apparently they had no idea of a combustion as understood in the modern sense occur- ring in the tissues, for they did not consider the source of animal heat in starvation. Even physiologists of 1803 like Richerand do not consider starvation in their texts. Lavoisier taught us the true cause of animal heat, but he could not explain the theory as to-day because the law of the conserva- tion of energy had not been formulated. Joule (1844) by his masterly researches evaluated the mechanical equivalent of heat, and largely on this basis Mayer (1845) gave expression to the law of conservation of energy, the application of which was made by Helmholtz. Energy cannot be destroyed or created. As even the ancient Greek Democritus (370 B. C.) once said, ‘‘Nothing can ever become something nor can something become nothing’’—ez nihilo nihil fit, et in nihilum nihil potest revertt. Dulong, Depretz, Regnault, Reiset, Pettenkofer and Voit con- tributed considerable to the study of animal heat and in turn helped to perfect the calorimeter. It remained for Rubner’* to 9¢¢Chaleur’’? probably would be translated thermodynamics and ‘‘du ealorique’’ heat. 10 Lavoisier et Seguin: ‘‘ Mémoire sur la Respiration des Animaux,’’ 1789. 11 Rubner: Zeit. f. Biologie, 1883, xix, 313; 1885, xxi, 250; 1886, Sadi 40; 1894, xxx, 73. HISTORY OF THE CALORIE 525 prove that physiological activity in the animal body is no excep- tion to the operation of the law of conservation of energy. He showed that chemical change is the cause of animal heat. He discovered that the animal is a living calorimeter in which food, when burned, changes into another form of energy. MRubner, experimenting with dogs, measured the body’s income and output of energy, and determined the relation of heat produced by oxida- tion of the foodstuffs ingested to the products which are given out. This figure he compared with the energy produced as measured by the calorimeter and found an agreement within one per cent. This remarkable result has since been confirmed by other investigators who have been aided by modern methods, the refinements of which are difficult to describe adequately. Rubner’s important conclusions are that energy is not destroyed or created by the living organism; that the foodstuffs are oxidized within the body in a manner simi- lar to their oxidations in a chemical laboratory. The law of con- servation of energy therefore applies to the animate as well as to the inanimate world. When calories are mentioned in nutrition, it is from the point of view of food fuel value. The calorie value of a diet is a factor of great importance in nutrition. Frankland (1866) was the first to determine this for various foodstuffs by oxidizing them in a calorimeter. He did not express the results in ‘‘calories’’ but rather as ‘‘heat units’’ (loc. cit.) which had the same value. Stoh- mann (1879)'? and Rubner (1883) were apparently the first to use the term calorie as it is now applied in the science of nutrition. Rubner made three outstanding contributions regarding the calorie in nutrition: (1) He applied its present day meaning to the term; (2) he determined the caloric value of protein, fat and carbohydrate, figures which are widely used in determining the energy content of a diet; (3) he drew the distinction between the absolute and physiological heat values of foods. By absolute heat value he meant the amount of heat yielded by a substance when oxidized in a bomb calorimeter; the amount of heat produced by the substance in question when burned within the animal body he regarded as its physiological heat value. These values may or may not be identical, a fact which is of fundamental importance in the science of nutrition. Calorie as a mere word explains nothing. It is a symbol for an idea, however, which, as we have seen, has undergone changes brought about by the development of several sciences. Ancient 12 Stohmann: Journ. f. prakt. Chem., 1879, xix, 115. 526 THE SCIENTIFIC MONTHLY and hazy notions regarding the phenomenon of fire and combus- tion first contributed to this concept, placed on a firmer foundation by the clarifying influence of Lavoisier. The broad generalization regarding force in nature, receiving its impression in the law of the conservation of energy, played its réle in the evolution of the idea and should probably be regarded as the most important factor contributing to the development of this notion as it exists to-day. The history of the calorie in nutrition, therefore, is wrapped up in the history of nutrition itself and the fundamental natural sciences upon which this branch of knowledge rests. SOCIAL LIFE AMONG THE INSECTS 527 SOCIAL LIFE AMONG THE INSECTS’ By Professor WILLIAM MORTON WHEELER BUSSEY INSTITUTION, HARVARD UNIVERSITY Lecture IV—Ants, THerr DEVELOPMENT, Castres, NESTING AND Frepine Hasits M There is throughout the animal kingdom, as I believe Espinas was the first to remark, a clear correlation on the one hand be- tween a solitary life and carnivorous habits and on the other hand between social habits and a vegetable diet. The beasts and birds of prey, the serpents, sharks, spiders and the legions of predacious insects all lead solitary lives, whereas the herbivores, rodents, granivorous and frugivorous birds and plant-eating snails and insects are more or less gregarious. Man himself is quite unable to develop populous societies without becoming increasingly vege- tarian. Compare, for example, the sparse communities of the carnivorous Esquimaux with the teeming populations of the purely vegetarian Hindoos. The reasons for these correlations are obvious, for plants furnish the only abundant and easily obtainable foods and, at least in the form of seeds and wood, the only foods that are sufficiently stable to permit of long storage. In the previous lectures I have shown that the social beetles and bees are strictly vegetarian and that the social wasps, though descended from highly predatory ancestors, are nevertheless becoming increas- ingly vegetarian like the bees. The ants exhibit in the most strik- inv manner the struggle between a very conservative tendency to retain the precarious insectivorous habits of their vespine ancestors and a progressive tendency to resort more and more to a purely vegetable regimen as the only means of developing and maintain- ing populous and efficient colonies. Anthropologists have dis- tinguished in the historical development of human societies six successive stages, designated as the hunting, pastoral, agricultural, commercial, industrial and intellectual. Evidently the first three, the hunting, pastoral and agricultural, are determined by the na- ture of the food and represent an advance from a primitive, mainly flesh-consuming to a largely vegetarian regimen. Lubbock showed that the same three stages occur in the same sequence in the phylo- genetic history of the ants. At the present time we are able to give even greater precision to his outlines of this evolution. 1 Lowell Lectures. 528 THE SCIENTIFIC MONTHLY All the primitive ants are decidedly carnivorous, that is preda- tory hunters of other insects. That this must have been the char- acter of the whole family during a very long period of its history is indicated by the retention of the insectivorous habit, in a more or less mitigated form, even in many of the higher ants. Always striving to rear as many young as possible, always hungry and exploring, the ants early adapted themselves to every part of their environment. They came, in fact, to acquire two environments, each peopled by a sufficient number of insects, arachnids, myrio- pods, ete., to furnish a precarious food-supply. Most of the ants learned to forage on the exposed surface of the soil and vegetation and became what we call epigeic, or surface forms; while a smaller number took to hunting their prey beneath the surface of the soil and thus became hypogeic, or subterranean. Many of the latter are very primitive but their number has been repeatedly recruited from higher genera, which by carrying on all their activities within the soil have found a refuge and surecease from a too strenuous competition with the epigwic species. We have here some very interesting cases of convergence, or parallel development, since the underground habit has caused the workers, which rarely or never leave their burrows, to lose their deep pigmentation and become yellow or light brown and to become nearly or quite blind. As will be evident in the course of my discussion, the tendency towards vegetarianism is apparent among both the epigzic and hypogeic forms. The ants belonging to the oldest and most primitive subfamilies, the Ponerine, Doryline and Cerepachyine and also to many of the lower genera of Myrmicine, feed exclusively on insects and therefore represent the hunting stage of human society. Owing to the difficulty of securing large quantities of the kind of food to which they are addicted, many of the species form small, de- pauperate colonies, consisting of a limited number of monomorphic workers. Many of these species lead a timid, subterranean life. In the size of their colonies, which may comprise hundreds of thou- sands of individuals, the Dorylinz alone constitute a striking ex- ception, but one which proves the rule. These insects, known as driver, army or legionary ants and very largely confined to Equa- torial Afriea and tropical America, are strictly carnivorous, but being nomadic and therefore foraging over an extensive territory, are able to obtain the amount of insect food necessary to the growth and maintenance of a huge and polymorphic population. They are the famous ants whose intrepid armies often overrun houses in the tropics, clear out all the vermine and compel the human inhabitants to leave the premises for a time. In Africa they have been known to kill even large domestic animals when they were tethered or penned up and thus prevented from escaping. SOCIAL LIFE AMONG THE INSECTS 529 The pastoral stage is represented by a great number of Myrmi- cine and especially of Formicine and Dolichoderine ants which live very largely on ‘‘honey-dew.’’ This sweet liquid, concerning the origin of which there was much speculation among the an- cients, is now known to be the sap of plants and to become accessible to the ants in two ways. First, it may be excreted by the plants from small glands or nectaries (‘‘extrafloral nectaries’’) situated on their leaves or stems, where it is eagerly sought and imbibed by the ants. Second, a much more abundant supply is made ac- cessible by a great group of insects, the Phytophthora, comprising the plant-lice, seale-insects, mealy-bugs, leaf-hoppers, psyllids, ete., which live gregariously on the surfaces of plants. These Phytoph- thora pierce the integument of the plants with their slender, pointed mouth-parts and imbibe their juices which consist of water containing in solution cane sugar, invert sugar, dextrin and a small _ amount of albuminous substance. In the alimentary canal of the insects much of the cane sugar is split up to form invert sugar and a relatively small amount of all the substances is assimilated, so that the excrement is not only abundant but contains more invert and less cane sugar. This excrement or honey-dew either falls upon the leaves and is licked up by the ants or is imbibed by them directly while it is leaving the bodies of the Phytophthora. Many species of ants have learned how to induce the Phytophthora to void the honey-dew by stroking them with the antenne, protect and care for them and even to keep them in specially constructed shelters or barns. Some ants have acquired such vested interests in certain plant-lice that they actually collect their eggs in the fall, keep them in the nests over winter and in the spring dis- tribute the hatching young over the surface of the plants. Lin- neus was therefore justified in calling the plant-lice the dairy- cattle of the ants (‘‘hw formicarum vacce’’). This dairy business is, in fact, carried on in all parts of the world on such a scale and with so many species of Phytophthora that it constitutes one of the most harmful of the multifarious activities of ants. Their irre-. pressible habit of protecting and distributing plant-lice, scale- insects, ete., is a source of considerable damage to many of our cultivated plants and especially to our fruit-trees, field and garden crops. Ants mostly attend Phytophthora on the leaves and shoots of plants, but quite a number of species are hypogwic and devote themselves to pasturing their cattle on the roots. Thus our com- mon garden ant (Lasius americanus) distributes plant-lice over the roots of Indian corn. The habit of keeping Phytophthora was probably developed in- dependently in many different genera, and it is easy to see how the habit of feeding by mutual regurgitation among the ants them- selves might have led to the behavior I have been describing. Cer- Wol, XV .—34. 530 THE SCIENTIFIC MONTHLY tainly the genera that have developed trophallaxis among the adult members of their colonies are the very ones which most assiduously attend the Phytophthora. And it is equally certain that the latter habit is very ancient, because it was already established among the ants of the Baltic Amber during Lower Oligocene times and that, as we have seen, was many million years ago. The dairying habit has led to an interesting specialization in certain species: known as ‘‘honey ants,’’? which inhabit desert re- gions or those with long, dry summers. These ants have found it very advantageous to store the honey dew collected during periods of active plant growth, and as they are unable to make cells like those of wasps and bees, have hit upon the ingenious device of using the crops of certain workers or soldiers for the purpose. In all ants, as we have seen, the crop is a capacious sac, but in the typical honey ants it becomes capable of such extraordinary distention that the abdomen of the individuals that assume the role of ani- mated demijohns or carboys, becomes enormously enlarged and per- fectly spherical. Such ‘‘repletes’’ (Fig. 66) are quite unable to walk and therefore suspend themselves by their claws from the ceilings of the nest chambers. When hungry the ordinary workers stroke their heads and receive by regurgitation droplets of the honey dew with which they were filled during seasons of plenty. The condition here described, or one of less gastric distention, has been observed in desert or xerothermal ants in very widely sepa- rated regions and belonging to some nine different genera of Myrmicine, Formicine and Dolichoderine (Myrmecocystus and Prenolepis in the United States and Northern Mexico, Melophorus, Camponotus, Leptomyrmex and Oligomyrmex in Australia, Plagio- FIG. 66 Replete of honey ant (Myrmecocystus melliger) from Mexico. a, lateral aspect of insect; b, head from above. SOCIAL LIFE AMONG THE INSECTS bol lepis and Aéromyrma in Africa and Pheidole in Australia and the southwestern United States). A more direct vegetarian adaptation is seen in many Formi- cide that inhabit the same desert or xerothermal regions as the honey ants. In such regions insect food is at no time abundant and is often so searce that the ants are compelled to eat the seeds of the sparse herbaceous vegetation. At least a dozen genera, all Myrmicine, illustrate this adaptation: Pogonomyrmex, Veromes- sor, Novomessor and Solenopsis in America, Messor, Oxyopomyr- mex, Goniomma, Tetramorium and Monomorium in the southern Palearctic region, Meranoplus in the Indoaustralian, Cratomyrmex and Ocymyrmex in the Ethiopian region and Pheidole (Fig. 57) in the warmer parts of both hemispheres. It was at one time be- lieved that some of these ants actually sow around their nests the grasses and other herbaceous plants from which they gather the seeds, but this has been disproved. They are merely collected, husked and stored in special chambers or granaries in the more superficial and dryer parts of the formicary. Emery has shown that as food the proteids are preferred to the starchy portions of the seeds and are also fed to the larve. Messor barbarus, the ant to which Solomon refers, is one of these harvesters. Probably none of them disdains insect food when it can be had. Neverthe- less the adaptation to crushing hard seeds is so pronouneed in cer- tain genera that the mandibles have become distinctly modified. Their blades have become broader and more convex and the head has been enlarged to accommodate the more powerful mandibular muscles. In certain forms (Pheidole, Messor, Novomessor, Holeomyrmex) the soldiers or major workers seem to function as the official seed-ecrushers of the colony. The harvesting ants can hardly be regarded as true agricul- turists because they neither sow nor cultivate the plants from which they obtain the seeds. Yet there is a group of ants which may properly be described as horticultural, namely the Attiini, a Myrmicine tribe comprising about 100 exclusively American species and ranging from Long Island, N. Y., to Argentina, though well represented by species only within the tropies. The tribe includes several genera (Cyphomyrmex, Apterostigma, Sericomyrmex, Myrmicoerypta, ete.) the species of which are small and timid and form small colonies with monomorphie workers, while others (Atta and Acromyrmex) are large and aggressive and form very popu- lous colonies with extremely polymorphic workers. The Attas or parasol ants inhabit the savannas and forests of South and Central America, Mexico, Cuba and Texas. Their extensive excavations result in the formation of large mounds and often cover a consid- erable area (Fig. 67). According to Branner, a single mound of THE SCIENTIFIC MONTHLY 532 Csojzuny “ff “S Aq yde1s0j0y ) "SUXOT, ‘BILOPIA Je (DUDrA} DIFP) JUL suTyNd-fea] UeXIT, of} JO ISON LQ ‘oIt 4 af SOCIAL LIFE AMONG THE INSECTS 533 the common Brazilian Atta serdens may contain as much as 265 cubic meters of earth, and the population of a colony of this species, according to Sampaio, may number from 175,000 to 600,000 in- dividuals. Of course, the size of the mounds varies with the depth of the excavations, which are much shallower in the rain-forests than in the dry savannas. From their mounds the ants make well-worn paths through the surrounding vegetation and frequently defoliate bushes or trees, cutting large pieces out of their leaves and carrying them like banners to their nests. The pieces are then cut into smaller fragments and built up on the floors of the large nest chambers (Figs. 68 and 69) in the form of sponge-like vollenweidert, c Argentinian leaf cutter, Atta 68 FIG. Vertical section through the center of a nest of the Carlos Bruch). (Photograph by Dr. showing the chambers containing the fungus gardens. 534 THE SCIENTIFIC MONTHLY masses, which become covered with a white, mould-like fungus mycelium (Figs. 70 and 71). The latter is treated in some un- known manner by the smallest, exclusively hypogeic caste of work- ers, so that the hyphe produce abundant clusters of small, spheri- cal dwellings, the bromatia (Fig. 72), which are eaten by the ants and fed to their larve. Each species of Attiine ant cultivates its own particular fungus and no other is permitted to grow in the nest. That the bromatia are really anomalous growths induced FIG. 69 Portion of nest of Atta vollenweideri shown in Fig. 68, Bruch. ) ed to show the sponge-like fungus- (Photograph by Dr. Carlos more enlarg About one eighth natural size . gardens im situ in the chambers. EE SOCIAL LIFE AMONG THE INSECTS 535 by the ants is indicated by the fact that they do not appear when the fungus is grown in isolation on artificial media. Alfred Moel- FIG. 70 Portion of fungus garden of the Texan leaf-cutting ant (Atta texana). About one half natural size. FIG. 71 Fungus garden built in a Petri dish by a colony of Apterostigma in British Guiana. Natural size. (Photograph by Mr. Tee-Van.) 536 THE SCIENTIFIC MONTHLY FIG. 72 Modified mycelium (bromatium) of fungus cultivated by the Argentinian Moellerius heyeri. The globular swellings of the hyphz are produced by the ants. (After Carlos Bruch.) ler, who was the first to cultivate these fungi, regarded them as belonging to the Agarics and named one of them Rozites gongylo- phora. Wither the ants prevent the mushrooms from appearing, or, more probably the subterranean conditions under which the mycelium is cultivated are unfavorable to their development. Moeller was also unable to obtain the mushrooms in his cultures, but found those of Rozites growing on the surface of an aban- doned Acromyrmex nest. That the fungi cultivated by the various Attiini belong to several different genera is shown by Bruch and Spegazzini who have recently been able to identify the mushrooms of the fungi cultivated by several Argentinian Attimi. Acromyr- mex lundi, e. 2. cultivates Vylaria micrura Speg., Moellerius heyert, Poroniopsis bruchi Speg. and Atta vollenweideri, a gigantic Agaric, Locellina Mazzuchu Spee. (Fig. 73). The lower genera of the Attiini differ in many particulars from such highly specialized forms as Atta and Acromyrmex. Their nests are smaller and there are differences in the gardens and the substratum, or substances on which the fungi are grown. The species of Trachymyrmex suspend the garden from the ceiling of the nest chamber instead of building it on the floor, and in some species of Apterostigma it is enclosed in a spherical envelope of dense mycelium, so that, except for its larger size, it much resem- bles the silken egg-case of a spider. These ants and others, such as Cyphomyrmex and Myrmicoerypta, use the excrement of other insects, especially of caterpillars, as a substratum for the gardens, and one species, Cyphomyrmex rimosus, cultivates a very peculiar funeus (Tyridiomyces formicarum Wheeler), which does not grow SOCIAL LIFE AMONG THE INSECTS 537 FIG. 73 a, Locellina Maszzuchii the gigantic fruiting phase (pileus 30 to 42 cm. in diameter!) of the fungus cultivated by the Argentinian leaf-cutting ant (Atta vollenweideri) ; b, section of same; c, basidia; d, spores. (After C. Speg- gazzini. ) in the form of a mycelium but of isolated, compact bodies, resem- bling little pieces of American cheese, and consisting of yeast- like cells. The same or a very similar fungus is grown by the species of Mycocepurus. How do all these Attiine ants come into possession of the various fungi which they cultivate with such consummate skill. The ques- tion is, of course, twofold, since we should like to know how the individual colony obtains its fungus and how the ancestors of the existing Attiini first acquired the fungus-growing habit. The former question has been answered by the very interesting investi- gations of Sampaio, H. von Ihering, J. Huber and Goeldi on the Brazilian Atta sexdens and of Bruch on the Argentinian Acromyr- mex lundi. The virgin queen of these species, before leaving the parental nest for her marriage flight, takes a good meal of fungus. The hyphex, together with the strigil sweepings from her own body and, according to Bruch, also some particles of the substratum, are packed into her infrabuccal pocket, where they form a large pellet, which she retains till she has mated, thrown off her wings and made a small chamber for herself in the soil. She then casts the pellet on the floor of the chamber where its hyphe begin to proliferate in the moist air and draw their nutriment from the extraneous materials with which they are mingled (Fig. 744). The queen carefully watches the incipient garden and accelerates its growth by manuring it with her feces (C and D). She begins to lay eggs (Fig. 76 A) and even breaks up some of them and adds them to the garden, which soon becomes large enough to form a kind of nest for the intact and developing eggs (Fig. 74 B to F). 538 THE SCIENTIFIC MONTHLY The young larve on hatching proceed to eat the mycelium and eventually pupate and emerge as small workers, which break through the soil, bring in pieces of leaves and add them to the garden. The care of the latter then devolves on the workers and the queen henceforth devotes herself to laying eggs. The colony is now established and its further development is merely a matter of enlarging the nest, multiplying the gardens and increasing the population. Thus Atta and Acromyrmex transmit their food- plants from generation to generation in a very simple manner, that is, merely by the queen’s retaining, till she has established her nest chamber, the infrabuccal pellet consisting of her last meal in the colony in which she was reared. And there is every reason FIG. 74 Stages in the development of the fungus garden by the queen of the Argen- tinian Moellerius hyeri. A, pellet of substratum 36 hours after its ejection from the queen’s infrabuccal pocket. The hyphe have begun to grow. B, same pellet after 3 days, with 4 eggs; C, same pellet after 8 days, showing droplets of feces with which the queen manures the hyphe. D, same pellet after 12 days, also showing droplets of feces; E, small fungus garden after 30 days, with 32 eggs; F, same after 40 days. The magnification of all the figures is very nearly 10 diameters. (Photographs by Dr. Carlos Bruch.) SOCIAL LIFE AMONG THE INSECTS 539 BGs. 75 A, an infrabuccal pellet of the queen Moellerius heyeri after cultivation for 36 hours on gelatine. X1o. B, eggs and pellets made of filter paper by a queen Moellerius heyeri that had failed to develop a fungus garden. X1o. (Photo- graph by Dr. Carlos Bruch.) to suppose that the same method of transmitting the fungus from the maternal to the daughter colonies is practiced by all the other genera of the tribe. Of course, the answer to the question as to how the ancestors of the Attiini acquired their food-fungi in the first place must be purely conjectural. Yet certain observation by Professor I. W. Bailey and myself seem to indicate from what simple beginnings the elaborate fungus-growing habits may have been evolved. An examination of the infrabueeal pellets of the most diverse ants shows that in nearly every case they contain fungus spores or pieces of mycelium collected from the surfaces of their bodies or from the walls of the nest. Moreover, many ants have a habit of casting their pellets on the refuse heaps, or kitchen-middens of their nests, and Professor Bailey finds that in the case of certain African 540 THK SCIENTIFIC MONTHLY FIG. 76 Behavior of the queen of Moellerius heyeri. A, photographed in the act of laying an egg. The incipient fungus garden in which the egg will be placed is shown to the left resting on the floor of the nest chamber; B, queen placing an egg in the fungus garden which is sticking to the glass wall of the artificial nest. C, queen photographed in the act of placing a droplet of feces in the fungus garden. Magnification 5 diameters. (Photographs by Dr. Carlos Bruch. ) SOCIAL LIFE AMONG THE INSECTS 541 DIGS 77. Two friendly queens of Moellerius heyeri caring for a single incipient fungus garden, which is adhering to the glass wall of the artificial nest X5. (Photo- graph by Dr. Carlos Bruch.) Crematogasters that live in the moist cavities of plants (Plectronia, Cuviera) the refuse heaps consist very largely of such ejected pellets and produce a luxuriant growth of aérial hyphe which are cropped by the ants. From such a condition it is, perhaps, only a short step to the establishment of small gardens consisting at first of the pellets and later of these and accumulations of extraneous ma- terials, such as the feces of the ants, those of caterpillars and beetles, vegetable detritus, ete., which might serve to enlarge the substratum and increase the growth of the fungus. The selection of particular species of fungi and their careful culture and transmission are evidently specializations that must have been established before the stages represented by even the most primitive existing Attiimi could have been attained. Whatever may have been the processes whereby the ancestral Attiini developed the fungus-growing habit, it must have originated in the more humid portions of the tropics, since nearly all the more primitive species of the tribe are still confined to the rain-forests. But certain species soon found that by sinking their galleries and chambers to a greater depth in the soil they could easily carry on their fungus farming even in arid regions. Thus some species of Moellerius, Trachymyrmex and Cyphomyrmex have come to live in the dry deserts of Arizona, New Mexico and northern Mexico, and as they can always find in such localities enough vegetable ma- terial for the substrata of their gardens, they have attained to a eontrol of their environment and food-supply, which even the hu- man inhabitants of those regions might envy. 542 THE SCIENTIFIC MONTHLY THE MARINE FISHERIES, THE STATE AND THE BIOLOGIST By WILL F. THOMPSON ANY of the great marine fisheries of the world lie within the M jurisdiction of more than a single sovereign country. The great North Sea fisheries, those of the Grand Banks, the salmon fisheries of the Fraser River (now nearly extinet), and the halibut fisheries of the North Pacific might be cited as examples. It would seem that this divided ownership has reacted most disastrously upon their care. Responsibility seems in such cases to be simply lost, not divided, and the net effect is that no one cares to saerifice his own interests to maintain the fishery for the benefit of all. The splendid scientific work done in the North Sea contrasts vividly with the relative futility of the movement to conserve the vanishing bottom fisheries there by regulatory laws. There is simply no ma- chinery capable of overriding the selfish interests of the few in each country, supplemented as it always is by the general suspicion one nation seems to think it necessary to have of every other nation. The case seems far more hopeful, where there is no division of authority. And in many of our great states fisheries exist which are entirely under the legal control of a single state government. That is true in California, where there is not a single fishery common to both its own water and the waters of another state or country save of the sparsely inhabited desert of Lower California, to whose less exploited fisheries vessels often go from Southern California ports. There is thus no possibility of shirking respon- sibility—the eare of its fisheries devolves upon California alone by virtue of the Constitution of the United States and of its geograph- ical position. No question of nationalism can be involved, only that of sectionalism. As a result, the failure to conserve the fisheries for the people of the entire state can result only from faulty organization of public opinion and the lack of real proof of the necessity of conservation. The securing of this proof of the condition of the fisheries has in California, as everywhere else, been recognized as a legitimate funetion of the responsible government, and the proper execution of that function is vital to the success of any popular movement toward conservation. Unlike the forests and the mines, private ownership has never been granted in the fisheries, save in the case MARINE FISHERIES 543 of oysters and those of certain fresh waters, and for that reason aroused popular opinion is entirely likely to control in the end. But powerful interests have grown up who will vigorously object to curtailment of their activities. Something tantamount to legal proof is necessary before what seems to them confiscation may be indulged in. And they have in the past shown a vitality which augurs ill for any but well-based movements toward conservation. The general policy of conservation is, moreover, largely sup- ported by men among the public who are not trained scientists and do not know the value of evidence. Their conclusions as to the existence of depletion may carry weight where, as in the case of birds and mammals, any honest man may observe conditions with his own eyes, and where powerful interests are not placed in jeopardy. But in the marine fisheries this is not true, for the fish are not easily observed and the evidence must come from statistical proof of comprehensive character. Under such circumstances the ery of conservation, raised hysterically and hastily, as is done even by scientists at times, must in the long run lead to failure and the injury of the cause at stake. And measures of regulation or re- striction passed in response to pleas made on an insecure basis must in the end fail to justify themselves. So the acquirement of real knowledge, while a protection to the men legitimately engaged in exploitation, is equally such to the cause of conservation itself, for it should not only prevent this lack of balance and undue regu- lation, but it should prevent the growth of interests which must later be curtailed. This necessity of knowledge was acknowledged by the fishery authorities of the State of California when they instituted the present system of observing the fisheries. Their action in this re- gard was based on the following facts: First, that enough ac- curate knowledge already exists to prove the susceptibility of marine fisheries in general to overfishing; second, that proof is required in the case of each individual fishery, and that there is no way of knowing the strain a species will stand save by submitting it to one; third, that such a course of action implies the duty of the state to maintain a constant and intelligent ward over its fisheries ; and finally, that such a ward is possible and that it implies con- tinuous and prolonged statistical and biological investigations. In regard to the first point, the existence of proof that marine fisheries are exhaustible, we must turn to the oldest and best known of fisheries, namely those in North European seas. Contrary to the opinions of many, these great fisheries cannot justifiably be called ancient. The use of steam vessels began in 1880; the otter trawl first came into use in 1895, laying open to exploitation the depths of the ocean below fifty fathoms; while the means of mar- 544 THE SCIENTIFIC MONTHLY keting and the extent of the demand increased equally with the recent great industrial expansion. The latter involved the de- velopment of railroads, the refrigeration of food products, the use of cans for their preservation, fast steamships to carry them, the growth of city life as a market, ete. Meanwhile, as cited by Jen- kins (‘‘The Sea Fisheries,’’ 1920) as an illustration of the trend of the times, the number of fishing vessels in the port of Aberdeen, Seotland, increased 258 per cent. in the period 1897-19038, and ac- cording to the estimate of the same authority, agreeing with that of others, the efficiency of each steam trawler of to-day exceeds eight times that of the sailing trawler it displaced (aside from the independence the steamer has of weather and distance). The fisheries in other parts of the world are still more recent, and show the same great increase in apparatus although not necessarily in ecateh. If these facts are considered, it is im- possible to doubt that, unless civilization comes to an abrupt pause, with the destruction of our highly developed transportation and of our industrialism which builds towns and markets, we are on the brink of an era of exploitation of our fisheries, and not at the erest of such an era. And the existence of overfishing now be- comes a serious problem, for if the fisheries do show depletion, it is indeed a serious question whether they will, even in their more stable parts, survive the coming strain. Faith in our destiny and that of the world implies care of our resources of fish. That they do show depletion in certain fisheries is now proved. The most clearly ascertained instances at present are those of the bottom fisheries. Thus the halibut in both the Atlantic and the Pacific has decreased with great rapidity. But the bottom fisheries of the North Sea for plaice and other allied species are, as Gar- stang (‘‘The Impoverishment of the Sea,’’ 1900) says, “‘not only exhaustible, but in rapid and continuous process of exhaustion.’’ This conclusion has been seconded and supported by men who have, in the various countries around the North Sea, actually had the examination of the statistics in their care, as Heincke, Fulton, Thompson and others, such as Jenkins and Allen. And if these bottom fisheries already show exhaustion, since they are more sta- tionary, are most highly valued, and were first sought for, it is not to be expected that ‘‘pelagic’’ fish will show otherwise upon the imposition of greater strain, even though they are more abun- dant. But in this connection, it must, indeed, be remembered that there is no accurate means of determining whether pelagic fish actually are more abundant than other fish in the ocean—although we do know that the cod and the herring, for instance, are not numberless, as some estimates have made them. Aeainst such a view there has been urged the objection that MARINE FISHERIES 545 the fisheries seem to be prospering and to be continuing on a firm basis. That fact may be granted, however, without conflicting with the above conclusions. It may be admitted that the total yield of the fisheries does not everywhere seem to decline, but it can be proved that continuously greater toil is required to obtain it. That this decreasing yield for the effort involved does not attract more attention should be understandable when it is considered that the cost of catching fish is but a fraction of the cost of distribution. Thus the fisherman may receive eight cents per pound where the retailer asks forty cents, and doubling the fisherman’s price would add but a fifth to the retailer’s price. The cost of obtaining the fish could be multiplied many fold without seriously affecting the final price to the consumer. The latter is, moreover, willing to pay high prices for a product to which he has become accustomed, and the rarer it is the more he will pay. The increase in initial cost does not seem, in fact, to be of the greatest importance. There is also this fact to be taken into consideration, that there are influences which actually counteract the effect of increasing scarcity in raising the initial cost. The accompaniments of that intensified exploitation which results in depletion are the constant broadening of the fishing grounds, the inclusion of more than one species of fish and of inferior quality in the catch of the boats, the development of means of preservation, the constant improve- ment of gear and the increase in quantity of apparatus. All these things tend to eliminate the great and sudden fluctuations in amount of yield which are characteristic of fisheries confined to one species or one locality. These variations in yield render the ex- ploitation of the fisheries expensive and uncertain because the periods of abundance must be made to pay interest upon the capital and to maintain the organization during periods of scarcity. Their elimination as a result of intense fishing undoubtedly does reduce the cost of fish to the consumer, perhaps to the extent that for a while the influence of depletion in raising the initial cost will not be felt. But in the end that very fact may defeat the natural safeguard which should protect a species, namely, the lack of profit in carry- ing on a fishery when it comes dangerously near to exhaustion. It becomes possible to prolong a fishery because other species are taken; the by-product becomes the mainstay of the business and the depleted species is kept under a strain for which it could not itself pay. If it were not for cod, perhaps the halibut fishery of Iceland might have long since collapsed; and if it were not for the cheaper round fishes, the flat fishes in the North Sea might be pur- sued far less rigorously. On the Pacifie coast, the tuna and sardine Vol. XV.—35. 546 THE SCIENTIFIC MONTHLY fishery of California may have been of considerable assistance to the albacore fishery. In fact, the objection often raised to the pos- sibility of over-fishing that the fisheries are prospering and that they would immediately cease to prosper should depletion occur is not a valid one. But that they would ultimately fail as a result of over-fishing seems sure. Another basis for scepticism as to the reality of the fact of pos- sible exhaustion has been the seeming boundlessness of the sea and its resources. But every fisherman knows that the areas within which fish of a given species are found are very limited, perhaps less so in the eases of ‘‘pelagic’’ fishes than in those of ‘‘demersal,’’ vet highly limited nevertheless. And the scientist will testify to the sharp limitations which temperature, depth, salinity and eur- rents place upon every species, so that it is in reality only a very small part of the ocean which yields our commercial fishes. They are, in facet, limited largely to the area of the coastal regions or the continental shelf, where there is drainage from the land, and to comparatively small parts of that shelf. In so far as this pro- ductive area is concerned, Gran and others have remarked that it corresponds in general with the distribution of the minute plankton organisms which are vastly more abundant where coastal water is found; and upon these plankton organisms fish must necessarily exist in the final analysis. And even where conditions are thus favorable, and the fisheries are highly developed, as in the North Sea, Allen (‘‘Food from the Sea,’’ 1917) estimates that an acre yields but fifteen pounds of fish per year while pasture land yields seventy-three pounds of beef. In accordance with these facts the experiments which have been made in marking fish and observing the frequency of recapture have shown that the fishermen are able to take, and do take, a very high percentage of the bottom fish in the North Sea. What they do with other fish, such as the herring and the sardine, or in any other regions, is for the most part un- known. It is therefore a mistake to assume that the resources of the sea are inexhaustible, or that over-fishing characterizes small areas easily replenished from without. There is, indeed, no manner of gauging in advance the product- ivity of the ocean, in so far as edible fish are concerned. It is in the first place obvious to students of the matter that the amount of food present for fishes does not determine abundance, any more than the amount of grass did determine the abundance of the buf- falo on our plains, or of deer in our forests. But it is certain that the rate of reproduction varies widely, and with it the relative resistance to depletion of the species of fish. Such matters as egg production, length of life, varying mortality at different stages and time of sexual maturity must all be taken into account, together MARINE FISHERIES 547 with the sharp limitations provided by climatic and geographical conditions. Moreover, the relative amount of competition for the available food is unknown, although we do know that the com- mercial fishes are probably but a small part of the population to be supported by the sea. And even if the abundance of the species were a gauge to its resistance to a strain—which it does not neces- sarily have to be—there is thus far no method of accurately ascer- taining the abundance of any one species of fish or of all together save within limited areas of the ocean. It seems, indeed, that there is no method of measuring the amount of fishing a species will stand save by submitting it to a strain. The only hint which can be obtained concerning the limits of the fisheries in California come from a comparison of the pro- ductive area with that of the North Sea, where the bottom fisheries show decline. It is, however, very hard to define the productive area, save by the width of the continental shelf. The area within the one hundred fathom line in the North Sea is approximately 130,000 squares miles (nautical), while off the coast of California it is about 7,500 square miles. In the former ease this area is about 300 miles wide and 450 long, but in California the average width is but 8.4 miles, much of this rocky or unsuitable for bottom fishes. In this connection it must be recollected that, as cited above, Gran and other authorities regard the presence of coastal water with land drainage in it as essential to the production of abundant planktonic life. Such water is abundant off the coast of Europe, but the California coast is more arid in nature, especially the south- ern portion. However, the great fisheries of California are of the ‘‘nelagic’’ type, regarding which such speculation may be limited in value. Nevertheless, it is probably safe to say, when all is taken into consideration, that these fisheries are far more limited in pro- portion to length of coast line than is the case in the North Sea, and hence much more susceptible to overfishing. As has been said above, the possibility and actuality of over- fishing have been definitely proved, yet it seems true that there-is no arbitrary limit which can be economically assigned to any fishery. It would be indeed sheer waste to impose a limit below what might be safely taken and the alternative is plain, to allow the imposition of all the strain the species will carry. It is, asa matter of fact, the only politically practical course of action, at the same time being the correct one from the scientifie stand- point. But it must not be forgotten that the acceptance of such a fact implies the serious duty of close observation and prompt ac- tion, in ease of overfishing, by the government in control. That is 548 THE SCIENTIFIC MONTHLY clearly recognized by the fishery authorities of California and is the mainspring of their actions. These things having been recognized as true, it followed that a careful survey of measures necessary for such observations was in order, and this has been made in so far as possible. For such purposes the great mass of literature published by the various countries around the North Sea was available, especially that issued by the ‘‘Conseil Permanent International pour 1’Exploration de la Mer,’’ or inspired by it. It soon became evident that it was impossible for the State of California to undertake the many lines of general inquiry into the varying conditions of the sea and its life which had been investigated more or less by these European countries. That would have been tunnelling the mountain by re- moving it in its entirety. It was necessary for the state to limit its efforts to those fields which had been shown to bear directly on the ascertainment of the condition of the fisheries; namely, the measurement of the variance in abundance of the fishes in the sea, the effects of fishing upon it and the biological criteria of over- fishing. A eareful perusal of much of the hydrographic and planktonic work demonstrated its remoteness from the work in hand despite its undoubtedly great ultimate value, and showed that most of the immediate questions could be solved to the re- quired degree without their aid. There were necessary certain biological studies upon the fishes themselves, but above all a statis- tical study of the fisheries and the fish. This method of approach, as Johan Hjort has most appropri- ately said of a certain phase of it, is regarding the study of the fisheries in a similar light to the study of the vital statistics of mankind. It involves primarily the taking of what amounts to a comparative census from year to year in order to test the relative abundanece—not the actual abundance—of fish; then to determine whether such great fluctuations as appear are due to natural causes or to overfishing. For this program, the legislature of the state has passed laws taxing the fisheries industries fifty cents per ton of raw fish used for canning, and has definitely specified the duty of the agents of the state. It is unnecessary to give the details of these laws, but something as to their operation will be of use. Every commercial transaction involving the first sale of fish is accompanied by the giving of a receipt by the buyer upon a form issued by the fish and game commission and of this receipt one copy is returned to the commission and another kept by the dealer. There are, therefore, actual records of all fish taken for profit, according to the boat and to the day. This unique system has been most successful in its operation for the last three years, avoiding “ MARINE FISHERIES 549 what we now know were widely erroneous estimates in statistics; while the fresh fish dealer has frequently for the first time a record of his own dealings. The results obtained have continuity, and are in such detail that market conditions, changes in apparatus or fishing fleets, ete., may be readily discounted. So every commer- cial fishing boat becomes in effect a means of testing the abundance of fish, and it is possible to segregate the effects of scarcity of fish from the effects of those economic changes which alter the total yield. This appears the necessary procedure from the experience of investigators in the North Sea, and is preferable to the limited experimental fishing which is possible. We do, in fact, feel con- fident that we will have a relatively accurate and sensitive record of the variations in abundance of fish in the ocean, when studied in connection with biological facts. This scientific collection of statistics is the starting point and the foundation for further investigations. The interpretation of the evidence drawn therefrom is the duty of the biologists engaged by the commission; for the great fluctuations in abundance of fish which may be shown must be analyzed and their true nature dis- eovered. Such natural fluctuations are very likely to be mistaken for depletion from overfishing; or, perhaps, if of opposite trend, as a contradiction of any theory of overfishing when they are in truth, as we have said, due to natural causes, and depletion may exist despite the temporary obliteration of the evidence. There must, as a consequence, be developed and utilized those biological criteria which distinguish depletion due to excessive fishing. The biological knowledge necessary for the use and formulation of such eriteria includes among other things the determination of age, the discovery of migrations and in so far as possible the correlation of abundance with natural physical conditions. One may justi- fiably call it ecology on a vast scale. Granted a fair knowledge of these criteria, it is not exceeding the reasonable to hope that the fishery authorities will be able to give warning when depletion is occurring—and, indeed, unless a degree of confidence can be placed in the competency of the work, the exploitation of the fisheries should not be allowed to proceed freely, nor can freedom be had from the constant fear of ruthless exploitation. There is, in addition, a need on the part of legislators for com- petent data upon which measures of regulation may be based. The imposition of arbitrary and reckless restrictions should be prevented by the acquisition of proper knowledge as soon as pos- sible. At present many of our fishery laws are untenable from a scientific standpoint, save in so far as they actually operate to re- duee the take. And even if it be said that legislatures will not take proper action, it would be a defeatist’s attitude to take to 500 THE SCIENTIFIC MONTHLY fail to provide them proper knowledge upon which they might take action. There are a great many legislators who will act along the line of their best knowledge, and more who will respond to intelli- gent pressure on the part of the public. In thus accepting conservation as a major policy because of its dependence upon the legal powers of the state, the program adopted in California has not been oblivious of the fact that the work for that purpose has a very definite bearing upon some of the greatest problems of exploitation. As an example, the abundance of fish is subject to great natural fluctuations beyond the control of man. The return from the fisheries vary greatly from day to day, from season to season, and from year to year. The resultant waste is an exaggerated case of the same kind which the electrical engineer meets when he is faced with the ‘‘peak load’’ er maxi- mum use of electricity during a short period each day. Just as apparatus must be available to carry this ‘‘peak load,’’ so must the fish canners or dealers maintain the machinery and organiza- tion for brief periods of maximum supply and longer ones of scarcity as well as variations in demand which are disconcerting both to the dealers and consumers. The meat packers, their rivals, need not do this. The understanding of these fluctuations so that regularly recurring ones may be expected, others foretold and provision made to meet or avoid them, is without doubt one of the most neglected functions of government scientists. The proper study of depletion necessitates just such an understanding of these changes as will serve the industry. It must be acknowledged that in adopting such a program, in- stalling such a system of statistics and founding a California state fisheries laboratory at San Pedro to care for the biological science of the subject, the state of California is experimenting. It still remains to be seen whether popular support will be rendered the project, either on the part of scientific men or the general public. The field seems to be one in which the scientist, particularly the biologist, should weleome a chance to show how his work can be applied to the needs of humanity ; but, aside from this, basic prin- ciples of animal life and behavior are really involved to such an extent as to satisfy the most academic of men and are attacked with the aid of vast masses of material unobtainable through any other source than the commercial fisheries. On the part of the public, it would seem that only a failure to understand or lack of faith in the competency of the work could lead to lack of support. It is sincerely to be hoped that this effort to approach the prob- lems of conservation upon a rational and well-balanced basis will meet with the reception its sincerity deserves. DE ANOPLURIS 5oL DE ANOPLURIS By Professor G. F. FERRIS STANFORD UNIVERSITY HE distinguished professor had attained some portion of his distinction by reason of years of work spent upon a small and little studied group of insects. His wife and small daughter chanced into his laboratory one day and discovered a student examining one of these insects through a microscope. The small daughter demanded a look and then came the inevitable question, ‘‘ What is that?’’ ‘‘That,’’ said the unsophisticated student, ‘‘is a louse.’’ There was a moment of pained silence and then came the gentle but unmistakable rebuke from the professor’s wife, ‘‘We always eall them Anoplura.’’ Now these very insects are the subject of my present discourse and lest I again offend the delicate sensibilities of any one I have disguised my intentions by a title to which not even the most fastidious should be able to take exception. To be sure it means the same thing as something else that might have been used, but after all there 7s something in the name by which a thing is called. Even the scratching soldier, from whom one would least expect any delicacy in such matters, conceals the identity of these insects under the euphemistic titles of ‘‘seam squirrels’’ and ‘‘cooties.’’ That the deference thus accorded them reduces in the slightest degree the frequency or the painfulness of their attentions may be doubted, but at least the victim is enabled thereby to retain a bit more of the shreds of his self-respect. In fact under the name of ‘‘cooties’’ these insects may quite properly become a subject even of parlor conversation. The word carries a faintly humorous connotation. One may without risk of immediate social ostracism speak of the great wads of hair that girls wear over their ears as ‘‘cootie coops.’’ True, such an ex- pression might not be looked upon with favor in the most refined circles, but we need only reflect upon what would happen were the wording changed a bit to see what a concession has been gained for it to be used at all. It is perhaps a fortunate thing that this has happened, for even entomologists, who should have put all squeamishness behind them, have been more or less reticent in speaking of these particular 552 THE SCIENTIFIC MONTHLY insects. The remark of an author writing in 1842 that ‘‘in the progress of this work, however, the author has had to contend with repeated rebukes from his fellows for entering upon the illustra- tion of a tribe of insects whose very name was sufficient to create feelings of disgust’’ might almost be justified to-day. In fact, m a rather recent entomological book it is said that ‘‘from their habits lice are not popular insects even for entomologists to take up,’’ and certainly the amount of attention they have received has never been large. Yet I confess to being a student of these insects and I do so without hesitation and without apologies. Not to me are they merely ‘‘disgusting parasites.’’ Not to me is the term ‘‘louse man,’’ with which my botanical and chemical and even entomo- logical acquaintances, with a misguided sense of humor, see fit to address me, a term of reproach. For, know you, there are very few who can merit it, scarcely more than half a dozen men in all the world in fact. To us it is a title of distinction, an evidence that we few have been able to avoid the well-beaten paths of the butter- fiy and beetle hunters and strike out into a but little explored country. For us it is a country of much interest and—dare I say it?—even of some beauty. And it is my hope that in these pages I may lead others to see in these disgusting parasites, these cooties, some of the things that we, their devotees, are able to see. For a proper understanding of these parasites, these lice, as I shall not hesitate henceforth to eall them, it should be explained that there are really two quite different sorts of them. One sort, known as the bird lice or the biting lice, is found chiefly on birds, although there are a few species on mammals, while all the species of the other sort, the sucking lice, occur on mammals. There is a very great difference in the manner by which the species of these two groups obtain their food. The biting lice feed by biting off and chewing up bits of hair or feathers or skin seales while the sucking lice feed by inserting their beaks through the skin and sucking up the blood of their host. Such a difference in habit is a very important thing, for with it is most intimately bound up the matter of the potentialities of the insects for harm to the animal upon whieh they live. It is now a firmly established and generally recognized fact that many of the most important diseases of man and of other animals as well are transmitted by insects. In by far the majority of cases the insects concerned are forms that live upon blood, that actually pierce the skin of the animal upon which they feed. Thus in feed- ing upon successive individuals these insects may transfer disease- producing organisms from one individual to another. This poten- DE ANOPLURIS 553 Oo tiality for evil is inherent in every blood-sucking form, and its pos- sibilities are realized to a high degree in those blood-sucking lice that live upon man, the familiar ‘‘cooties’’ of the war period. Under the conditions that usually prevail in armies it is impos- sible for soldiers to keep themselves free from these insects. Thus it was that certain diseases which are transmitted by the lice be- came especially prevalent during the late war. Typhus and trench fever are transmitted by lice, and as far as known only by lice, and the measures for the control of these diseases were directed chiefly against their insect carriers. The tremendous losses due to these diseases undoubtedly had a profound effect upon the fight- ing strength of the various armies. Who can say to what extent the course of the war may have been influenced by them? There are a few other diseases of man that are known to be carried by lice, and they have been suspected of carrying several others. It is known also that an epidemic disease of certain small Asiatic rodents is transmitted by the sueking louse that occurs upon these animals, and it is highly probable that there are many other cases of the same sort. On the other hand the biting lice, although far exceeding the sucking lice in numbers of species, are not known to be the carriers of any diseases. If they are abundant upon their host they may cause injury merely by the irritation of their crawling about or by the matting of the hairs or feathers to which their eggs are glued. Otherwise they are of no concern to their host. But however important and interesting this connection of lice with the transmission of disease may be it is not the only thing about them that is worthy of consideration. This connection with disease is simply a fact, and after all facts are not always as inter- esting as theories, even though they may be more important. Any one should be able to travel the plain and open road of fact, but there is more pleasure in the narrow and devious trail of theory that occasionally takes the traveler up into the high places—and threatens always to lead him into a bog from which the utmost of mental agility may not be sufficient to extricate him! The most interesting thing about lice is not these highly important facts of hygiene. It is that they may be made to yield a contribution to biological theory. The starting point of this contribution is the fact that by far the majority of all the different species of lice, both of the biting and sucking groups, are found upon a single species of animal or at the most upon a few very closely related species. It is a curious fact that although horses and cattle and sheep have for many hun- dreds of years been in close contact in their stables each has re- 554 THE SCIENTIFIC MONTHLY tained its particular kinds of lice. There are at least four species of lice upon domestic cattle, but these do not occur upon horses or sheep. There are at least three kinds upon sheep, but none of these has been taken from horses or cattle. There are at least two kinds on horses, but neither of these has been taken from eattle or sheep. These instaaces may be paralleled by many others. The in- ference is obvious. It is evident that under normal conditions each species of louse ‘‘prefers’’ to feed upon a particular kind of ani- mal. In other words it is adapted to feed upon the blood or epi- dermal structures of this particular host and does not find the blood or the epidermal structures of another kind of host a suitable food. Furthermore these insects are very reluctant to leave their host and even after its death may be found clinging to the hairs or feathers. Under experimental conditions lice have been fed upon animals very different from those upon which they normally live, and it is true that sometimes under natural conditions they may be found upon an animal on which they do not belong. Still this does not change the fact that usually each species of louse sticks pretty closely to a certain kind of animal, passing from one individual to another in the nest or at mating time. Thus it is that the parasites are in a way inherited by the young from their parents—heirlooms, if I may be pardoned an atrocious pun. The next fact of interest is that the same species of louse may be found upon distinct but closely related species of birds or mam- mals in widely separated parts of the world. Thus we have upon the kingfisher in North America a louse that is the same as one on a kingfisher in Egypt. Another species is found upon various species of hawks throughout the world. Another is found on seals in both the Pacific and Atlantic Oceans. Another is found on ground squirrels in Siberia and in North America. Now all these animals are so widely separated that certainly it is impossible for the louse of the African kingfisher to transfer directly to the North American kingfisher or for the louse of the Atlantic seal to transfer directly to the Pacific seal or for a trans- fer to take place in any one of the many other cases of this sort that might be mentioned. Then how has the parasite managed to get upon both of its widely separated hosts? There are enough facts of this character to demand some at- tempt at a logical explanation. There must be a reason for them. If we remember that these parasites are normally passed down from one generation to another as a sort of racial inheritance and if we follow far enough the train of reasoning that is thus initiated we can not but conclude that at some time this African kingfisher DE ANOPLURIS 559 and the North American kingfisher, or the North American ground squirrel and the Siberian ground squirrel, or the members of any such pairs as we may name, were together as a single species. We have a set of facts that can not reasonably be explained unless we accept the theory of evolution. The conclusion is inevitable that at some time these kingfishers, or whatever they may be, had a common ancestor and that the shifting of land masses or climates or some other cause has left part of the descendants in one corner of the world and some in another and that there by various evo- lutionary processes they have become sufficiently different to be recognizable as different species. One of the most remarkable examples of the working out of such a change is that of the llama and the camel. The paleontologists tell us that at one time there were many more species of camels and camel-like animals than there are now and that these animals first appeared in the New World. At the present time, however, the only remaining representatives of this group are the llamas of South America and the camels of Asia and Africa. Yet, sepa- rated by half the world—these cousins, or forty-second cousins, still show their relationship and their common ancestry not only in their bedily structure but in their parasites as well, for on both there is found the same species of louse. We can extend the list of facts much farther and still the answer of common ancestry is the only reasonable explanation of them. The louse of the domestic hog has its nearest relative in the bush pigs of Africa. The lice of one kind of squirrel are more closely related to those of other squirrels than they are to the lice of other animals. The lice of the domestic chicken have their nearest rela- tives in the many different species of lice upon the other chicken- like birds. Like nearly all theories, however, this one must allow fer certain exceptions. We can lay down a general rule but it is almost too much to expect it to work always, at least if we are dealing with such things as living organisms. It is one of the difficulties in the way of the study of living things that they refuse to stay put. They simply will not go always into the little pigeon holes that we block out for them. Like men they must at times assert their indi- viduality by breaking our trifling little laws; they at times demand the right of living their own lives in their own way. In some in- stances related species of lice are found upon animals that are certainly not very closely related and sometimes different species of lice are found on animals that really ought to have the same kind. So it must simply be admitted that in some cases other in- fluences have been at work. Still the broad facts remain as I have pictured them. 556 THE SCIENTIFIC MONTHLY All this leads finally and directly to the question, ‘‘What about the lice of man?’’ And here the evolutionist may rub his hands together in satisfaction. For here—in the most interesting place of all—the theory works! The lice of man do indeed find their nearest relatives in the lice of the apes and monkeys. In faet, it is even possible that lice of some of the apes are really the same species that occurs on man. Nor can the argument that here we are probably dealing with one of those exceptions that I have men- tioned hold, for the facts throughout are entirely too consistent with each other. So the evolutionist who, heedless of the fair name of his species, would derive man from some ape-like ancestor finds here another bit of support for his theories. He finds another stone for the de- fending wall that piece by piece has been built about them. Nor will this wall, like the walls of Jericho, crumble before the blasts of its enemies’ trumpets. Not even before the most silver tongued of them. I would like to close this with a moral, but morals have gone out of fashion and then it is obvious enough, anyway. TOPOGRAPHICAL MAPS 55 ~] TOPOGRAPHICAL MAPS OF THE UNITED STATES By Professor WILLIAM MORRIS DAVIS HARVARD UNIVERSITY HE United States Geological Survey is still engaged in pre- paring a topographic map of our country. Progress thus far made is summarized on a large two-sheet map of the United States, on a scale of 1:240,000 or 40 miles to an inch, which serves as an index for all the quadrangles surveyed and published up to 1921; but, as a matter of fact, there are large areas in certain western states which are here marked off as surveyed, but which are represented only by maps published nearly 40 years ago, on so small a seale and of such inferior workmanship that their areas must soon be surveyed again more worthily. The contrast between the vague, sketchy contours of those early maps and the minutely intricate and apparently accurate contours of the newer maps marks the advance in topographic standards during the interval of nearly half a century. But Americans as a rule are still topographically uneducated. They are accustomed to ‘‘flat’’ maps, on which the form of the land surface, the ‘‘relief’’ as it is technically called, is either not represented at all, or else so badly represented that it might better remain unrepresented. Automobilists are coming to know some- thing of the ascents and descents on the roads that they follow; but most of them are still so inexperienced in or distracted from the observation of the landscape that they do not look at it closely or attentively; and even if they do, they hardly see what they look at. The driver of a car of course should not be expected to turn his attention far to the right or left; but his fellow travellers may do so, and they would be greatly aided in seeing the country they traverse by carrying along the topographic maps of their route. The cost of the maps is very low; an inquiry addressed to the di- rector of the U. S. Geological Survey at Washington will bring information concerning maps published for any desired part of the country. Tf distance lends enchantment to some views, appreciation lends enjoyment to many others, and appreciations of landscape views is greatly increased by the possession of a good map. As examples 558 THE SCIENTIFIC MONTHLY of the contrasts between different parts of the country, look at the map of the Brasua Lake quadrangle, next west of Moosehead lake in Maine, where the brooks, many of them called ‘‘streams,”’ have a well-defined flow only in their steep descents from the up- lands, while in the lower lands they are for the most part either delayed in swamps or stopped in lakes; or of the Williamsport quadrangle, Pennsylvania, where the drainage is so well developed that neither lake nor swamp is to be found, and where the single or double ridges, running in the zigzag pattern of the Alleghenies, prevail with occasional enclosed limestone valleys, of which Nippe- nose is a perfect type; or of the Rives Junction quadrangle, Mich- igan, where the surface is agitated in the minute inequalities of morainie topography with many kettles and ponds; or of the Craig quadrangle, Missouri-Nebraska, where the boundary between the state of Missouri and Nebraska follows a former course of the Missouri river, which has now changed its channel to the right or left, thus inconveniently leaving patches of each state on the wrong side of the river; or of the Natchez quadrangle, where the uplands east of the Mississippi are cut into a labyrinth of intricately branch- ing ravines as they fall off to the broad flood plain in which the great river swings in large meanders; or of La Sal Vieja quad- rangle on the coastal plain of Texas, where the smooth surface has no valleys and very few hills, but is pitted by countless depressions, small and large, holding wet or dry lakes. The variety of topog- raphy is infinite; the lover of mountain and valley, of forest and stream will find no end of enjoyment in striving to apprehend its many expressions. None of the maps are more remarkable than those which repre- sent the slopes of the great voleanic island of Hawaii. Several of these have already been published, and four more soon to be com- pleted are now issued as ‘‘advance sheets, subject to correction, ’’ on a seale of 1:31,680 with 10-foot contours incompletely drawn. Two of these sheets include parts of the Kau Forest Reserve and a streteh of the Voleano Road that leads from Hilo on the east coast southwestward to the cauldron of Kilauea. All these sheets reveal admirably the long continuity of the gradual slope by which the voleano descends from its great height, the minute ravines incised with sub-parallel courses down the slope, the occasional ragged lava surfaces where the contours are given a minutely ser- rate pattern, the occasional oblique or radial scarps which seem to indicate fractures and displacements in the huge mass, and most striking of all a vast bulging or convex slope, skirted around its southeastern base by the Volcano Road, which contradicts the general idea that voleanic slopes are concave. If these sheets are TOPOGRAPHICAL MAPS 559 continued so that, when mounted together, they may include a large share of the island, they will afford an unequaled illustration of voleanic topography on a large scale. One of the several ways in which the newer maps are improved over the earlier ones is in the addition of submarine contours, with the same vertical intervals as those on the land, for quadrangles on the ocean and lake coasts. Thus the Cape San Martin quad- rangle, California, shows the bold slopes of the Santa Lueia range, which descends to the Pacific with crewded 50-foot contour lines, to be adjoined by a gently inclined sea-floor plain with wide-spaced 50-foot contour lines across a breadth of from two to four miles off shore, before a moderate slope to deeper water begins. In strong contrast therewith, the Portsmouth quadrangle shows the sea bottom off the coast of New Hampshire and Maine to be almost as undulating as the land, although, perhaps because soundings are seattered, the texture of the submarime undulations is drawn in a coarser pattern than that of the terrestrial surface. The manifest reason for the contrast between these samples of Pacific and At- lantiec borders is that the shallow sea bottom along the California coast has been uninterruptedly subjected to normal marine agencies —waves and currents—by which land-derived detritus is smoothly distributed ; while the sea bottom near the New England eoast has been recently, as the earth counts time, subjected to glaciation. Another novelty on the recent maps is the addition of the num- bers and subdivisions of the rectangles, over 900 in all, into which the whole country has been divided by the War Department. These rectangles measure one degree of latitude on the sides and one degree of longitude at the top and bottom; they are numbered from north to south in successive columns, beginning on the Pacifie coast. Each rectangle is divided into north and south halves; and eaeh half into four quarters (I, northwest; II, northeast; III, south- west; IV, southeast). Thus the Conejos, Colo., quadrangle of the Geological Survey nomenclature, on a scale of 1:96,000, is the 298-S-II & IV quadrangle of the War Department. When the seale is large, the numerical nomenclature is somewhat unhandy ; thus the Firebaugh, Calif., quadrangle of the Survey on a scale of 1 :31,680, is the 60-N-II-W/2-SW/4 quadrangle of the War Depart- ment. Outline maps of the states have been called for and prepared in recent years on a scale of 1:500,000. All are now completed, except that Nevada, Utah, Colorado and New Mexico are in press, and Texas is yet to be drawn. These maps will doubtless be issued eventually with contours, but for the present the map of such a state as South Dakota shows only its rivers and streams, many of 560 » THE SCIENTIFIC MONTHLY which are printed in broken lines to show their intermittent flow, its railroads, its county and village names, and its township and section rectangles as marked off years ago by the Land-Survey preliminary to selling the public lands. Many of our state maps in the central and western parts of the country are based on those crude surveys, which served their purpose well enough when they were honestly made in plain country, but ia rough country the case was different. One of the recently issued Geological Survey maps of a quadrangle in a mountainous state explains in a legend at the bottom of the sheet that the township and sectional rectangles of the Land Survey for a part of the area are omitted ‘‘because land plats and topography can not be reconciled and no [section] corners can be found.’’ This recalls a story of early days in California, told by the late Professor Brewer of Yale, who was in the 60’s a member of the California Geological Survey. A desperado, at last captured after many deeds of violence, was about to be hanged by a vigilance committee. When asked if he had anything on his mind which he wished to confess, he said he had; but it was not his manifold murders that troubled him. The only misdeeds to which he owned up with remorse had been committed while he was an assistant on the Land Survey; the law required that the corners of the square-mile sections should be marked with wooden posts, charred at one end and driven into the ground; the desperado confessed that, in a district where wood was scarce, he had marked the section corners with burnt matches. We are only about half- a-century climb up from that rung in the ladder of our civilization. One of the most characteristic signs of our ascent to a higher level is the preparation of a large number of excellent maps of our do- main, some examples of which are noted above. IMMIGRATION LEGISLATION 561 WHAT NEXT IN IMMIGRATION LEGISLATION ? By Professor ROBERT DE C. WARD HARVARD UNIVERSITY THe ImpERATIVE NECESSITY OF FURTHER ‘‘PERMANENT’’ LEGISLATION HE present 3 per cent. immigration restriction law will ex- pire on June 30, 1924. What shall take its place? The con- ditions which led to its adoption will still exist. We are facing a permanent tendency toward rapidly increasing and _ steadily deteriorating immigration, and millions of prospective immigrants Overseas are impatiently waiting for the thirtieth of June, 1924, when they will rush in, in a seething chaotic mob, unless Congress takes steps to stop them. THE OPPOSITION TO RESTRICTION To any program for restriction there is certain to be active, well-organized and heavily financed opposition. This opposition is centered in (1) certain racial groups which are interested, not in the future of America, but in the increase of their own race in America; (2) employers who want foreign labor so cheap that it is dear at any price; who put pocketbook above patriotism; (3) the steamship companies, who believe themselves to have vested rights in the United States as a dumping-ground for their human cargoes, and (4) those who have been well termed ‘‘the incurable senti- mentalists.’’ Every effort is now being made to misconstrue our present laws; to distort and misrepresent their effects; to make them unworkable and unpopular. These laws are being subjected to organized attack by ‘‘interested’’ individuals, alien racial groups and hyphenated societies, and certain influential newspapers which carry heavy steamship advertising. All of these are bent on making any restriction whatever appear unreasonable, unjust and inhumane. It is very important that the character, the motives and the tactics of this opposition should be known. Our GENERAL IMMIGRATION LAW MUST BE MAINTAINED AND ; ENFORCED There can be no question that our general immigration law of 1917 must be maintained. This law names some thirty classes of Vol. XV.—36. 562 THE SCIENTIFIC MONTHLY aliens who are excludable on mental, physical, moral or economic grounds. ‘These include the insane, the idiot and the feeble- minded; those who have loathsome or dangerous contagious dis- eases ; criminals; prostitutes ; persons physically incapacitated from earning a living, illiterates, etc. In fact the enumeration of the undesirable classes is so complete that, if the law had been and were always rigidly enforced, our immigration ‘‘problem’’ would give us far less trouble than it does. Our law bars criminals, but our court and institution records show a large excess of foreign- born. Our law bars the insane, but our insane hospitals, especially in the northeastern States, are filled with aliens. Our law bars those suffering from loathsome and dangerous contagious dis- eases, and those suffering from physical disability that may affect ability to earn a living, but political ‘‘pull’’ often suffices to admit over the doctor’s certificate, against the express provision for ex- clusion. Our law debars paupers; yet an insignificant number of - aliens is debarred on these grounds, although the majority of those now arriving come without money, and are not productive laborers. In a recent paper on the deportation system of several States, Dr. H. H. Laughlin, of the Eugenics Record Office, Cold Spring Harbor, N. Y., brought together some startling facts. A recent survey shows that in 1916 the several states expended on an average of 17.3 per cent. of their total governmental expenditures in main- taining custodial and charitable institutions. This percentage varied from 5.4 in Alabama to 30.5 in Massachusetts. A survey of 460 state institutions for the several types of the socially inadequate, with a total of 210,835 inmates, recently (1922) completed by the Committee on Immigration and Naturalization of the House of Representatives, found 21.14 per cent. of these fifth of a million inmates to be of foreign birth and 44.09 per cent. to be of foreign stock—that is, of foreign birth or who have at least one parent foreign born. Thus if, on the average, it costs the same in the institutions to maintain native-born and foreign-born inmates, then currently the several states are expending approximately 7.62 per cent. of their total revenues in caring for degenerate and dependent human forejgn stock. This is the logical outgrowth of the asylum idea which has pervaded the American immigration poliey. The proper enforcement of our general immigration law in- volves not only a very careful and deliberate scrutiny of all ar- riving aliens, but also a systematic and thorough round-up of all aliens already in this country who are deportable because they have become public charges, or who have been found to belong to certain other specified classes of undesirables which are by law subject to deportation. Never yet since the law of 1917 has been on our statute books has it been strictly enforced. It is to the credit of the present administration that a distinct improvement in IMMIGRATION LEGISLATION 563 this respect was made during the past year. And it should be re- membered that strict enforcement leads to a certain extent to self- enforcement, for the more aliens are debarred as undesirable, the fewer such attempt to get in. Tar PERCENTAGE LIMITATION MUST BE MADE PERMANENT The present 3 per cent. law is not perfect, but it has on the whole worked successfully, and has fully justified its enactment. It is reasonably generous in permitting the reuniting of families; in allowing unrestricted entry to tourists and other excepted classes, and it has kept our ports open to a fairly considerable inflow of newcomers, for it should be remembered that it permits an annual immigration of over 350,000. It has undoubtedly worked hard- ships in some cases, but most of the newspaper stories of such hardships have either been intentionally exaggerated, or have been untrue. Public sympathy is easily aroused by a single instance of real or fictitious hardship. The far more vital problem of how the present character of immigration is to affect the American race of the future is more remote, and attracts less attention. The ad- ministration of the law is fortunately in the hands of officials who are enforcing it with justice and humanity. There is one point in connection with the 3 per cent. law which is often lost sight of. For a good many years before the war, aliens from southern and eastern Europe largely outnumbered those from northern and western Europe. Under the new law these numbers are nearly equalized, so that if all nationalities fill their allotted quotas, the so-called ‘‘newer’’ immigration can not contribute more than about one half of our annual inflow. This fact is biologically of great significance. In the fiscal year ending June 30, 1922, deducting emigrants from immigrants, we gained in Nordic stock, and lost in the natives of southern and eastern Europe. Those who attribute solely to the present percentage restriction the need of labor in certain industries are either wholly ignorant of the facts, or are intentionally trying to mislead the public in the effort to break down all restrictions and to flood the country with cheap labor. In this connection it should be realized that (1) there has been a very considerable emigration of alien labor dur- ing the recent period of business depression and unemployment; (2) if all countries filled up their quotas, which they have not been doing during the past year, there would be an annual inflow of over 350,000; (3) the countries of northern Europe have fallen much farther below the quotas than those of southern and eastern Europe, most of the latter having exhausted their quotas, thus showing that the intelligent and skilled labor of northern Europe 564 THE SCIENTIFIC’ MONTHLY has not been disposed to emigrate to the United States; (4) the immigration of aliens who are natives of any countries of the New World is not subject to the provisions of the law; (5) a considerable proportion of our immigration under the 3 per cent. law has been made up of sweat-shop workers, peddlers and small shop-keepers, not of strong, sturdy, intelligent laborers. This is clearly not the fault of the law, but results from the present ten- dencies of immigration. Thus the ‘‘need of labor’’ is by no means to be attributed solely, or even largely, to the percentage law. Fur- thermore, there is little doubt that northern and western Europe will fill up its quotas during the coming year, as immigration from those countries is increasing. The relation of immigration restric- tion to the rising scale of wages was so clearly stated by Honorable Albert Johnson during the closing days of the session of Congress recently ended that we can not do better than to quote from his remarks on this point. Every good American sympathizes with workingmen in their effort to obtain decent wages and decent conditions. . . . Restriction is an absolutely necessary supplement to a protective tariff. Immigration must be curtailed until all workers, native and foreign-born, whether in the basie or other industries, get wages and have conditions commensurate with American standards and ideals if we are to maintain those standards, ideals, and, in fact, our very civilizatjon. Just as soon as wages in those industries rise to the point where the breadwinner can rear and support his family in keeping with American standards, the native-born will reinvade those industries from which they have been driven by the ruinous competition of imported cheap labor, often inducted into conditions amounting to slavery, and when wages do rise to that level the native supply will quite meet the demand, just as native labor does in every other country. This is the situation in a nut-shell. As a result of the awakening on the part of our people to the effects of the practically unrestricted new immigration from south- ern and eastern Europe and Asia, for the first time, and against bitter opposition, the principle of numerical limitation has been established by overwhelming majorities in Congress in a manner which gives equal treatment to all the nationalities which make up our population as far as is consistent with the maintenance of what we know as America. This principle, which has been long and strongly advocated by leading authorities on immigration, should be made our permanent immigration policy. Our own country, foreign countries, the steamship companies—all have be- come more or less adjusted to a definite numerical limitation of our alien immigrants. The machinery is in operation, and works remarkably smoothly, as much so as any restrictive legislation ever works. In reenacting a percentage law, whether it be the present 3 per cent., or the 2 per cent. which has been suggested in certain IMMIGRATION LEGISLATION 565 bills lately introduced into Congress; and whether the percentage quotas be based on the census of 1910 or an earlier census, it would be wise to make the law somewhat more elastic. Reasonable provision should be made to prevent the breaking up of families. This could readily be accomplished by treating the immediate members of a family as a unit whenever some members could be admitted without exceeding the quota while the remain- ing ones would otherwise be excluded because exceeding the quota. Exceptions in favor of bona-fide tourists, of students and of pro- fessional classes are necessary. Further, a maximum number of 500 or 600 could wisely be set for the admissible aliens from cer- tain countries whose quotas are very small under the 3 per cent. provision, and from which we receive highly desirable immigrants. The quota from Australia, for example, in the present fiscal year is only 279 and that from New Zealand and Pacific Islands is only 80. With these changes, a percentage law such as the present one would involve very few hardships. At the same time if the excep- tions were carefully drawn, there would be no danger of our being swamped by any such flood of aliens as swept in upon us before the war, and as will, in even greater volume come in again unless we take steps to prevent it. It can not be too strongly emphasized that, while the original argument in favor of the 3 per cent. law was economic, the real, fundamental, lasting reason for its continuanee is biological. This side of the matter was so clearly and forcibly presented in an editorial note in the October number of the World’s Work that we can not do better than to quote that statement here: If America is to realize its fullest possibilities, it must exércise the prin- ciple of selection. Up to the present time it has ignored this method. Our policy of opening our gates to all comers has really meant that we have recognized no distinctions among peoples, that we have refused to admit that one presented better material for citizenship than another, and that we have pinned our faith on the existence of some wonder-working alchemy in the American atmosphere which could transmute an inferior race into a superior one. But the teaching of all history, as well as the experiments of the biolog- ical laboratory, show the absurdity of any such easy-going philosophy, and the nation has reached the point where it should base its future upon scien- tific and historical fact. This is really the argument in favor of the three per cent. immigration law. It does not directly apply this principle of selection, it is true; that is, it does not in so many words limit immigration in future to particular races and particular nations. Yet indirectly it does accomplish a result which is not dissimilar. It takes the population of 1910 as representing the propor- tions of different peoples which, under the practical limitations of the prob- lem, may be regarded as furnishing the desirable racial composition of the future United States. The great majority of that population came from the countries of northwestern Hurope—Germany, Scandinavia, Great Britain and Ireland. There are few who have studied the matter who do not regard these 566 THE SCIENTIFIC MONTHLY peoples as the most desirable elements with which to construct the nation. By limiting future arrivals to three per cent. of these stocks, therefore, the law does provide that the American people of the future, as well as of the present, shall be chiefly from the races of northwestern Europe. That is the reason why this law, or one based upon the same principles, should represent the permanent policy of the republic. OBJECTIONS TO A FuAt NUMERICAL LIMITATION WITHOUT PROPER SELECTION The sole purpose of such a numerical limitation as that em- bodied in the present 3 per cent. law is to cut down numbers. Be- cause immigration was largely of an undesirable quality we cut it down. The 8 per cent. law certainly does let in a smaller amount of bad stock, but does not improve the stock, physically, mentally or morally. It doubtless shuts out some highly desirable immi- grants because the quotas are mostly filled with undesirable ones. Furthermore, being based on nationality, 7. e., country of birth, and not on race, it has made possible a disproportionate immigra- tion of Jews to the exclusion of thousands of non-Jewish aliens. This fact comes about because of the extraordinary activities of Jewish relief societies, both in this country and in Europe. These organizations take all the steps necessary to enable their co-religion- ists in Europe to emigrate to the United States, such as procuring their passports, purchasing their passage tickets, and earing for them en route to the ports of embarkation. Thus the annual quotas from several European countries are largely filled with Jews. In this way the ‘‘flat’’ percentage restriction has been worked with injustice to non-Jewish aliens who desire to come to the United States. It is because a flat percentage restriction works only quantita- tively and not qualitatively that it is absolutely necessary to main- tain and to enforce our general immigration law of 1917, as urged above. OVERSEAS INSPECTION OF PROSPECTIVE IMMIGRANTS IS NOT PRACTICABLE There seems to be so many and such obvious advantages, both to the prospective immigrant and to the United States, in having some sort of examination overseas, that there is at present a wide- spread demand for such inspection. This is no new agitation. It has marked the history of immigration literature and debate for at least thirty-five years. Such foreign inspection would seem to be our only way of looking into the antecedents, habits and character of our intending immigrants; of picking out those who by heredity and education are best fitted to become American citizens; of eliminating, at the source, all those who, under our general immi- IMMIGRATION LEGISLATION 567 gration law, are physically, mentally, morally or economically un- desirable. Overseas inspection suggests itself as a humane method of stopping most of the inadmissible aliens before they start on their voyage, and it should be welcomed by the steamship com- panies, for it would mean that few rejected aliens would have to be taken back at the companies’ expense. The plan recently most widely advocated is that the United States should establish an immigration office in each consulate to function in connection with the work of viséing passports, the im- migration inspectors to certify as to an alien’s admissibility to this country before his passport is viséd. On its face this plan seems sensible, wise and humane. It seems to offer a simple and practical solution of the immigration problem. Yet there are so many objections to it, and so many obstacles in the way of its accomplishment, that for years all committees of Congress which have considered it, as well as the Immigration Com- mission of a decade or so ago, and leading authorities on immigra- tion, have been forced to abandon it. A first objection to overseas inspection is that it would necessitate a very large increase in the number of immigration inspectors and medical officers, with the resulting heavy expense. If an undesirable alien is to be stopped before he leaves home and if the antecedents and character of our prospective immigrants are to be accurately ascertained, we must have our inspectors and doctors at all the thousands of places all over Europe and western Asia from which our immigrants come. For it is obvious that a United States immigration inspector at a consulate in Hamburg, é. g., making an examination there of, say, a thousand Jews coming from all parts of Poland, would be no better able to determine their eligibility there than on their arrival at Ellis Island. Of course, those who might be declared inadmis- sible by the inspector at a European port, or at some inland city where we have a consulate, would be saved the voyage across the Atlantic, but complete information concerning any alien could only be obtained in his home town or hamlet. In the second place, overseas inspection would divide the responsibility between the officials abroad and those at our own ports, for there is no question that we must, under any and all conditions, always maintain our inspection service at our ports. In all doubtful cases, each in- spector, the one abroad and the one here, would throw the respon- sibility upon the other. Thirdly, overseas inspection, supple- mented by home inspection, would work hardship on the aliens because it would never be certain that all with overseas certificates would be allowed to land on a second examination here. Lastly, whenever overseas inspection of prospective immigrants has been seriously considered by Congress, certain foreign govern- 568 THE SCIENTIFIC MONTHLY ments have objected to it on the ground that this country would thereby be assuming extra-territorial sovereignty not in accord- ance with treaty rights. Delicate diplomatic problems are here in- volved. No scheme for foreign inspection could be devised which did not use the existing machinery of consular offices. As the Honorable Albert Johnson has recently said, ‘‘Our consular offices are established under authority of trade and commercial treaties, each of which sets forth specifically what functions may be earried on by consular employees. Examination of persons who contem- plate migration to the United States is not included. It follows that if we are to set up a plan for examination of immigrants over- seas we must of necessity revise many of our trade and commercial treaties.’? This is the situation in which the United States finds itself in this matter of overseas inspection. It is obvious that the interests of the United States and those of foreign countries are absolutely opposed in this matter of immi- eration selection. We want the sound, able-bodied, intelligent. We do not want the defective, the delinquent, the physically unfit. The former are the ones most desired at home. The latter, foreign governments would not regret to have emigrate. It is, therefore, readily understood why these governments may not be too ready to acquiesce in any new arrangement whereby we can select the best and refuse the worst of their people. The present passport and visé system gives foreign governments the power to designate and to allow to emigrate those persons only whose presence in their own countries is not desired. The selection of our future citizens is therefore not in our own hands. In spite of the present obstacles in the way of our establishing overseas inspection, it might perhaps be possible, through ordinary diplomatic channels, without the necessity and the delays of nego- tiating any new treaties, to come to an amicable working agree- ment—a Gentlemen’s Agreement, in short—with the governments of foreign countries from which our immigrants come, whereby, by international cooperation, the United States could make some sort of a preliminary examination of intending immigrants before they sail. If the present administration could bring about such an agreement, it would take a long step in the settlement of this most difficult and important national problem. The sympathetic atti- tude of the present Secretary of Labor on this question has been clearly indicated. Thus, in an address given in Boston on June 14 last, he said (The Boston Herald, June 15) ‘‘We must bar out those who menace our national life and our national institutions. Much could be done by providing for inspection of prospective immigrants in Europe before they undertake the long journey IMMIGRATION LEGISLATION 569 across the Atlantic. I would insist upon the most rigid tests of blood, physical, mental and moral stamina before admitting a single immigrant.’’ With this view all patriotic and thinking Americans must surely agree. Overseas inspection, however, desirable as it would be, should not and could not in any way replace the other restrictive and se- lective measures advocated in the foregoing discussion. We im- peratively need a stricter enforcement of our general immigration law, and a permanent percentage limitation with the amendments above suggested. ADDENDUM: If we want the American race to continue to be predominantly Anglo-Saxon-Germaniec, of the same stock as that which originally settled the United States, wrote our Constitution, and established our democratic institutions; if we want our future immigration to be chiefly made up of kindred peoples from north- ern and western Europe, easily assimilable, literate, of a high grade of intelligence, able to understand, appreciate and intelligently support our form of government, then the simplest way to accom- plish this purpose is to base the percentage limitation upon an earlier census than that of 1910, 2. e., before southern and eastern Europe had become the controlling element in our immigration. In an important discussion of ‘‘The Immigration Problem,’’ in Scribner’s Magazine for September, 1922, which came to the pres- ent writer’s attention after he had completed the foregoing article, Professor Roy L. Garis, of Vanderbilt University, suggested that our permanent legislation be based upon the percentage principle, but that we admit 3 per cent. of the different nationalities of for- eign-born in the United States as shown by the census of 1890. This, as Dr. Garis rightly says, ‘‘is a simple yet practical solution, based on historical facts.’’ If instead of 3 per cent. we should admit 5 per cent. of the foreign-born resident here in 1890, the an- nual total for all Europe would be 400,000, in round numbers. Of these, about 200,000 would be admissible from northwestern Eu- rope; 50,000 from Scandinavian Europe; 165,000 from Central Europe; 10,000 from eastern Europe; 10,000 from southwestern Europe, and 2,000 from southeastern Europe. Such a law would result in bringing in a large preponderance of immigrants who present no difficulties of assimilation; who do not give rise to our immigration “‘problem.’’ It would thus be automatically selective, as well as numerically restrictive. If we are to maintain the physi- cal and mental standards of our race; if we are to make America safe for democracy, to keep America for Americans, there is no more logical or practical method than this. ; 570 THE SCIENTIFIC MONTHLY A MODERN MECCA By Professor A. E. KENNELLY HARVARD UNIVERSITY HE ancient Arabian city of Mecca is in one sense the most re- markable in the world. It is the birthplace of Mahomed, the Resoul ’Illah, or prophet of God, and it contains the Kaaba, or cen- tral shrine of the Mahomedan faith. No giaour may enter there. Very few Christians have ever seen the place. The explorer Bur- ton, for instance, one of the most famous oriental linguists, pene- trated there, under elaborate disguise, at the risk of his life. Over two hundred millions of persons—about one eighth of the population of the globe, or nearly double the population of the United States—are reputed Mahomedans. The great majority of these Mahomedans are devout worshippers, according to the ritual of the Koran. Several times a day, notably at dawn and sunset, the faithful, scattered over Asiatic and African lands, unfold their prayer-carpets, lay them in the assumed direction of Mecca, and prostrate themselves thereon in prayer. This custom of diurnal obsequience towards the prophet’s holy city has continued for cen- turies with very little change. Mecca has thus maintained a mar- vellous hold upon the daily thoughts and acts of a vast number of human beings. In their psychology, the Kaaba is the common and universal point of reference. Their lives are lived in perpetual procession towards that one place. Every year, about one hundred thousand pilgrims of zeal journey from afar to visit it. He who has thus attained to the holy city acquires with pride the title of Hajji, which entitles the owner to social respect. Although the modern world has created no new Mecea in any such sense as Mahomedanism conveys, yet in a certain very re- markable way it has created a scientific Mecca at Sévres, in the beautiful park of St. Cloud, on the Seine, near Paris, and close to the famous old Sévres porcelain factory. Here, on a plot of land which France has given to the world, by extraditing it from French boundaries, is an unpretentious building, called the ‘‘ Bureau Inter- national des Poids et Mesures,’’ especially constructed in 1875, for enabling accurate measurements of standard lengths and weights to be carried on quietly, and at a nearly uniform temperature, within its walls. Hight meters beneath the surface of its courtyard is a vault containing a closed steel safe, wherein are treasured the A MODERN MECCA 571 world’s most valued reference standards of length and weight— the international meter and kilogram—together with certain ‘‘témoins’’ or ancillary duplicates. The building is mainly devoted to comparisons between these fundamental standards and the cor- responding replica of other nations, through the medium of work- ing copies. It has no upper stories and is surrounded by the forest park. Fortunately for the even tenor of its duties, it ordinarily escapes attention from the ubiquitous tourist, despite its proximity to the much visited porcelain museum. Neither Baedeker nor ‘‘Guide Bleu’’ refers to it, and it finds no place on the list of ob- jects to attract the interest of the visitor; yet it may properly be described as a latter-day temple for the guardianship of the ‘‘ Lares and Penates’’ in the world of weights and measures. The expense of its upkeep is shared, in definitely prescribed proportions, among the twenty-eight foremost countries of the world. The need for an international clearimg house and depositary for the world’s standard meter becomes evident from a considera- tion of industrial needs alone, as well as from a cursory glance at the history of the subject. If a steel bar, say one yard in length, is ordered, without any further specifications, from a smithy, we might reasonably expect the bar to be delivered true to length within one per cent., more or less; that is to a precision of one part in 100, or 107. This may be described as a precision of the second order. It does not mean that the smithy could not furnish a higher degree of precision in length, if the need existed; but merely that in the ordinary course of business, a yard bar at the smithy might well be interpreted to include bars longer or shorter than a yard by as much as one per cent. Moreover, the measurement of the bar, by the act of laying a yard-stick alongside it, would be an operation lasting only a few moments. Nevertheless, if the tolerance of something more than one per cent. were clearly admitted, the moral certainty of the measure within those limits would be very great. Any intel- ligent man receiving the bar from the smithy, and laying a eredit- able yard-stick beside it, could see at a glance that the two were nearly alike in length; so that unless some question as to the amount of tolerance were raised, he would feel a high degree of assurance that—in the common use of language—the bar was a yard long. The rough bar from the smithy might next be sent to a machine shop with an order that it be trimmed, smoothed and ecut to a finished length of say 35 inches; but without any specifications as to tolerance in precise length. In the operations of smoothing and finishing, the mechanic entrusted with the order, would probably 572 THE SCIENTIFIC MONTHLY ? spend a little time in adjusting its length to ‘‘35 inches.’’ One per cent. he would regard as unworkmanlike, and he would probably aim at a precision say of one per mil; 7%. e., one part in 10%, or of the third order. If he were put upon his mettle, he might be will- ing to attempt a precision of the fourth order, or 1 in 10*; but the extra time involved in such an effort might not be justifiable. Even to reach the third order, he would have to make the length correct within 0.035 inch, and much more time would be needed in the measurement than was spent at the smithy in attaining the second order. In the first place, an ordinary rough yardstick would no longer suffice. A graduated steel tape or straight edge would be necessary. Moreover, the operation of juxtaposition and reading off the length would take much longer than before. Finally, in spite of this increased mental and physical care in the task of measurement, the degree of moral certitude as to the re- liability of the result will probably not have increased propor- tionately. In repeating the measurement, the mechanic will begin to arrive at slightly different results and the effect of the dis- crepancies disconcerts the judgment. Of course, unless the me- chanic made a gross error or mistake in reading the tape, he would feel convinced that the finished bar was much nearer to ‘‘35 inches’’ than one per cent.; but upon the new plane of third- order precision engaging his attention, he might feel less assurance of success than his predecessor, who measured only to the second order of precision at the smithy. If now the finished 35-inch bar were sent to a superior work- shop, to be assembled perhaps in some fine piece of mechanism, after being adjusted to say 34.8 inches, at a room temperature of 20° Centigrade, with a tolerance of 0.004 inch, this would entail refinishing the ends of the bar and measuring its new length to one part in 10*, or to fourth order precision. For the purpose of making this measurement, a special gauge might have to be pre- pared. If the bar in the assembled mechanism had to be made interchangeable with similar bars coming from other machine shops, the gauge might have to be adjusted to a precision of the fifth order in length, at a special workshop for the construction of precise gauges. In order to reach fifth order precision in the gauge at the special workshop, a standard measuring rule of yet higher precision would be needed there. It would not be unreason- able to require a precision of the 514th order in that special work- shop standard. That standard would probably be compared and calibrated against a still finer standard at the Bureau of Standards in Washington, where a precision of the 6.5th order, or one per 3,000,000 might be readily obtainable. Finally, to keep the stand- A MODERN MECCA 573 ards of the different national laboratories of the world in mutual agreement, an international standard is maintained at Sévres, where the precision attainable is of the seventh order. It is clear that every measure of length, either in the world of business or in the world of science, has its order of precision, over the entire range from the first to the seventh, depending upon its construction and purpose. No one expects a higher order of pre- cision than the particular purpose of the measure in question de- mands; because the effort and expense involved in securing an extra order of precision is relatively great. There are certain lines of industry, notably in gauge making for fine tools, where fifth order precision is necessary. Not many decades ago, this was the highest order scientifically attainable in measures of length. The advance from the fifth to sixth order demanded an immense amount of scientific and industrial effort. Progress had to be made in the mathematics of accidental errors, in metallurgical chemistry to provide improved materials, in physics to learn the laws of length variation in material standards, in tools, to fashion the improved parts, in workmanship and experience to handle the new tools. At the present date, we can look for the seventh order precision in the comparison of the various national meters with the inter- national meter at Sevres; but although this suffices for practically all industrial needs, it is inadequate for certain scientific require- ments. Certain problems of the Einstein theory, for instanee, might find solution, if the eighth or ninth order of precision in the measurement of length were attainable. If material civilization advances, we may hope to secure one higher order of length precision at Sévres in the course of another eentury. This would probably add one order to national scientific length measurement all over the globe. Apart from questions of moral or spiritual development, an estimate of the world’s eivi- lization might be furnished, based upon the order of precision realizable in the certificates of meter-bar comparisons furnished by the bureau at Sévres. The permanence and inviolability of the international meter are clearly of importance to all nations. The control of the Savres bureau has been vested, since 1875, in an international body com- posed of delegates from the twenty-eight leading national govern- ments that are parties to the bureau’s maintenance. The inter- national meter and kilogram are deposited at the bureau in such a manner that seven successive keys have to be used in order to reach them. Three of these keys are in regular service for the doors of the building; but the four others belong to the special vault in which the standards are preserved, and are placed in the 574 THE SCIENTIFIC MONTHLY custody of as many different officers of the international commit- tee. Some of these officers live abroad; so that the keys are usually kept far apart. It has thus only been possible to open the vault at the regular six-year meetings, when the assembling officers pro- duce their respective keys. The inconvenience of so inflexible a modus operandi manifested itself during the world war. At the outbreak of the war, one of the officers, entrusted with a key, resided _ in Germany. That key, being in an enemy country, was inac- cessible. If Sévres had been subjected to bombardment, it would have been necessary to remove the standard meter and kilogram to a place of safety, and this would have involved breaking into the vault. In order to provide against such a contingency in future, a new set of regulations has been brought into effect; whereby a duplicate of each vault key is deposited with the Insti- tut de France, and so that, under special emergency, the vault can be opened by its authority. One of these six-year meetings of the International Conference took place recently in Paris under the presidence of M. Emile Picard, the permanent secretary of the French Academy of Seiences. On October 6th, 1921, the delegates met at Sévres and at a specified hour formed themselves into a visiting and attesting committee, under the leadership of M. Ch. Ed. Guillaume, the Di- rector of the Bureau. The committee members line up across the courtyard in column by twos. At the signal, the procession enters the main and east door of the Bureau, and passes along a corridor, in the half light, to a descending stone stairway. Incandescent lamps light up, and we descend to a basement floor. Again, down another stone stair- way, to a sub-basement. At this level, the temperature changes but little all the year round. We now face the first of the three steel vault doors in the east wall. It opens to the corresponding official key. Behind it are two other steel doors, which are suc- cessively unlocked, revealing the vault beyond. This is about 4 meters long in an easterly direction, 3 meters wide and three high. An electric incandescent lamp, that has been idle for eight years, is turned on and we can see the interior clearly. The walls and ceiling are lined with white enamelled brick. Opposite to the en- trance against the eastern wall is a table supporting a steel safe. There is nothing else in the vault except an auxiliary table against the north wall. The director produces the last of the seven keys and unlocks the safe. Its doors swing open and disclose two shelves. On the upper shelf are three meter-bar cases, a mini- mum-maximum thermometer and a hygrometer. On the lower shelf is a row of five glass double bell-jars. Inside of each is a shin- A MODERN MECCA 575 ing cylindrical standard kilogram. The director calls for a reading of the instruments, which have been shut up in the safe since 1913. The min-max, thermometer register 10.6°—13.2° Centigrade, or a total range of only 2.6 degrees during those eight years. The hygrometer shows 88 per cent. humidity. The director ealls atten- tion to the international meter and kilogram, with their control duplicates. He carefully lifts out of the safe the central case, containing the primary standard or prototype meter, and lays it on the auxiliary table. He opens the case and reveals the brightly gleaming meter bar within. Like all the other standard bars, its section is of a special X shape, so designed as to offer the maximum stiffness, or resistance to sagging in the middle, when the bar is supported at its two ends. Of course, a stout platinum-iridium bar, a little more than forty inches long, and built with any rea- sonable shape of cross-section, would sag very little at the center when the bar is supported at its ends. Nevertheless an extremely small sag would be apt to alter, in perceptible degree, the apparent leneth of the bar, as measured on its surface. By giving this X shape to the section, and cutting a flat strip of surface at the middle of the groove in the X, the meter length is marked off along this flat strip, where the change in length due to any possible central sag becomes quite negligible. The meter bar is not graduated, or marked off into equal divisions like an ordinary rule. There is merely a fine scratch cut with a diamond point across the bar near each end. The international meter is defined as the distance be- tween these two fine line scratches, when the bar is at the tempera- ture of melting ice. No hand touches the bar, which lies face down in its case. It is the final reference standard, and does not need to be used except as a final arbiter, in case differences should arise among the work- ing standards. Its duty is merely to remain steadfast—to pre- serve its dimensions unchanged. Only a few of the witnesses can occupy the vault at one time, and the air in it becomes oppressive. As soon as those within have recognized the contents of the safe, they leave the vault and make room for others. In about half an hour, the standards are replaced on their shelves, the safe is shut and locked, the light is turned off and the vault doors closed. The world’s standards of length and weight thus resume their wonted repose, until the next awakening by a visiting committee, probably six years hence. This visiting com- mittee, however, returns to daylight and reforms in the courtyard, where it is photographed by a moving-picture machine, to record the passage of its members. 576 THE SCIENTIFIC MONTHLY * The measurement laboratories of the Bureau are also visited by the committee. These laboratories have specially constructed walls, to keep the temperature within them nearly uniform. In one of them, standard meter bars are compared and certified. The bar to be tested is laid horizontally on end supports in a tray of water, alongside of the working standard of the laboratory, whose error with respect to the international meter of the vault has been carefully determined. Being composed of platinum-iridium, the ~ bars do not tarnish under water; although they have to be cleansed from deposits of dust or organic matter—of which more anon. The immersion of the bars in water is desirable during the tests, in order to enable their temperature to be more closely determined, each bar being only true to length at a certain temperature, and correction being necessary for observed deviation of temperature from the standard. A microscope is supported on a stone pillar over each end of the bar, in such a manner that the cross-hair of the eyepiece can be brought over the delicate line marks on the ends of each bar in turn, the operations being regularly repeated many times accord- ing to a definite schedule. The observations, as finally collected and reduced, reveal the difference between the length of the tested bar and that of the working standard. Readings can be taken to the eighth order, or to 0.01 micron (10° meter, about 1/2,500,000th inch; but the final result is only depended upon to the nearest 0.1 micron, or ten-millionth of a meter, a precision of the seventh order. The micron is the one-millionth of a meter, and is the unit of length in practically universal use among scien- tific workers with the microscope, all the world over. Some excitement was recently aroused at the Bureau, by the discovery that two working standard meter bars, in use there during many years, although retaining their lengths almost un- changed with respect to each other, had both lengthened in that time by about 0.4 micron, or 1/65,000th of an inch, the change of length being progressive. After much search, it is believed that this minute lengthening of each bar has been gradually brought about through the constantly repeated cleansing of the polished working surface by the gentle rubbing of a cloth. The rubbing had always been directed, as a matter of habit, along the bar tu- wards the end. In this way, it is supposed that the walls of the fine diamond scratch across the bar, near each end, were slowly pushed over and away from the middle of the bar, by 0.2 micron, in the course of long usage. It has since become the prescribed routine to clean the surface of a meter bar by rubs delivered alter- nately in opposite directions. ; A MODERN MECCA 577 Another laboratory is devoted to the measurement of standard weights. In each corner of this room a delicate balance weighing machine is mounted, while the observer sits in the middle of the room at a distance of four meters from each and all of them. The reason for his having to sit so far from his work is that if he were to take up a position comfortably close to the balance he operated, the warmth from his body would probably vitiate the measure- ments seriously for the degree of precision aimed at, in spite of the usual windows and shields with which such a balance is ordi- narily provided. His respectful distance from the subject of his investigation suggests the old rhyme: “Who suppes with ye Deville Shoulde have a longe spoone.’’ With the aid of an ingenious arrangement of four parallel and horizontal brass rods, running from the balance to the observer, and which he can rotate in different ways, he is able to weigh the test kilogram mass against the mass of a standard kilogram, on the seales four meters off, and to read the balance through a telescope. One of these four-rod combinations enables him to observe the difference of weight between say the tested mass on the left-hand seale, and the standard mass on the right-hand scale; then to clamp the beam, next to lift off the two weights and reverse their relative positions, bringing the tested mass to the right and the standard mass to the left; and finally to repeat the weighing in this reversed relation, all at four meters’ distance. A standard kilogram is a solid eylinder of polished platinum- iridium, with its circular edges slightly rounded. The mass of such a cylinder can be measured, after making all corrections, to the nearest hundredth of a milligram; or to the 10-* kilogram—a pre- cision of the eighth order. It is curious that the available pre- cision of mass determination should thus be one order greater than that available in length determination in our national standards, at this period of the world’s history. It is evident that a blow, or severe mechanical shock, adminis- tered to any standard meter bar, might readily alter its length and render its use unreliable. Great care has to be taken in the na- tional laboratories of the various countries not to let a standard meter bar fall. Occasionally, a national standard meter bar is brought back to Sevres from some distant part of the world for recomparison at the Bureau. In such a ease, it is always entrusted to a careful messenger, and usually the chief of the national labo- ratory takes it himself. At last October’s Paris meeting two na- tional meter bars came back for retesting, one from Washington, D. C., and the other from Japan. In each ease, the director of the Vol. XV.—37. 578 THE SCIENTIFIC MONTHLY a national laboratory brought it in person. In thé American in- stance, Dr. Stratton brought the meter bar from the Bureau of Standards at Washington, with the aid of an assistant. The spe- cial case containing this meter always remained in the care of one or the other, being carried from place to place with more care than a mother ordinarily devotes to the handling of her baby. A tumble that might not hurt a baby might injure a meter bar. In the Japanese instance, Dr. Tanakadate arrived at Sévres, after a jour- ney from half way round the world, carrying his meter case im his arms. The way in which the precious national standard was ear- ried in each instance brings to mind the story of the Italian knight, the founder of the well-known Pazzi family, who, in one of the erusades, returning from Jerusalem on horseback, always sat baekwards, or facing the tail of his horse, because he tended con- stantly a little fire in a brazier, whose embers he had first lighted from the sacred fire in Jerusalem. Journeying painfully and slowly, he was determined to bring the fire still alive to his native Italian city. His unusual gait and attitude naturally aroused much astonishment on the journey, and the name ‘‘pazzo,’’ or crazy, clung, so the story goes, to him and his family thereafter. The reason for so much care being warranted, moreover, in the American instance was that both by act of Congress and by ex- ecutive order, the U. S. Yard is defined and maintained as a cer- tain definite fraction of the international meter, as is also the U. S. pound avoirdupois as another definite fraction of the inter- national kilogram. Consequently, all American measures are linked with the fundamental units at Sévres. Although the Wash- ington Bureau of Standards has more than one official copy of the meter; yet it is advantageous to have one of the copies checked and recompared at ‘‘Mececa’’ from time to time. After the international bureau at Sevres was organized in 1875, and set to work making and certifying copies of the international meter and kilogram, 30 standard meter bars in platinum-iridium were finished and approved by 1889. The bar which agreed most closely with the pre-existing standard meter of the ‘‘Archives”’ was adopted as the ‘‘international meter’’ for deposit in the vault, and the others were distributed by lot among the various contributing governments. Meter-bars Nos. 21 and 27 fell to the share of America, as well as Kilograms Nos. 4 and 20. The work of measuring and intercomparing the 30 standard bars, previous to their acceptance and distribution by the international commit- tee, took over three years to accomplish. Meter No. 27 and kilogram No. 20 were brought to Washing- ton, under seal, by an officer of the U. S. Coast and Geodetic Survey, - A MODERN MECCA 579 and delivered to President Harrison at the White House on Jan- uary 2nd, 1890. President Harrison, after breaking the seals and opening the cases himself, signed for their receipt and certified to the reception in good order of these national standards. These exercises were conducted with befitting ceremony, a special recep- tion, followed by a social function and dance, being arranged at the White House for the occasion. As originally planned, the meter was decimally derived from the dimensions of our universal mother earth, so as to be the ten- millionth of the distance from the North Pole to the Equator, on a meridian carried through Paris. In this way, if the meter should become lost or entangled in dispute, it might be re-established by gveodesy. It would not, of course, be necessary to start with a tape- line from the north pole. It would fortunately suffice to measure the length of a portion or portions of an are of the meridian, be- tween terrestrial stations whose difference in latitude should be determined with the requisite degree of precision by astronomical methods. A series of meridian measurements were actually con- ducted about the year 1799, for the purpose of arriving at the length of the standard meter, although the best scientific instru- ments available at that date were distinctly inferior to those now in regular use. A marked advantage of this decimalized meridian basis for the meter is that with decimalized angles, which are slowly but surely winning their way to favor through ease in their calculation, the kilometer, or ten-thousandth of the earth quadrant, becomes the nautical as well as the terrestrial mile, thus bringing both land and sea under the dominion of the same standards. Re- cent measurements have shown that the actual standard meter is a little short of the theoretical meridian meter. With respect to the meridian through Paris, the standard meter is stated to be approximately 0.2 millimeter, or 200 microns, too short. This error is insignificant and negligible from the standpoint of sailor’s charts, or navigational requirements; but it is an enormous error from the standpoint of national standard intercomparison at Sévres. By the time that the shortcomings of the standard meter with respect to the theoretical meter became known, it was too late to change the standard. Moreover, supposing that the stand- ard meter of to-day were corrected to the best known value of the theoretical meter; there can be no doubt that in the course of another century of progress in geodesy, the meter-bar thus cor- rected would have to undergo a new correction at that date. prob- ably much less than the 200 microns now known; but yet very large with respect to inter-comparison work. The result would be that we should never have a final meter, and we should always 580 THE SCIENTIFIC MONTHLY have to examine into the date of a meter-bar in order to be able to use it with a high degree of precision. For practical purposes, therefore, the world has been compelled to accept a standard meter whose precision with respect to the theoretically decimalized meridian is only of the 3.5th order, in order to secure a precision of the 7th order in replication and dissemination. In any ease, the probable precision of redetermination by astronomical measures is at present much less than that attainable by direct meter-bar com- parison. Consequently, if the standard meter should be lost in any one part of the world, it could always be re-established by reference to the international meter at Sevres. Will it always be necessary to make periodical comparisons be- tween national meter-bars and the international standards at Sevres? Perhaps not. Already a method has been developed, which, with the aid of the Michelson interferometer, enables the number of waves of cadmium red light to be counted in the length of one meter. This number has been stated to be 1,553,164.1, optically measurable with a precision of the seventh order, or prac- tically the same as that of direct mechanical comparison. It may, therefore, be found, after sufficient experience has been acquired, that instead of sending a standard bar from Washington to Sevres for calibration and certificate, it may only be necessary to use, in the Washington laboratory, the right kind of cadmium light, and to count off the right number of its wave lengths—a number exceed- ing a million and a half—in order to arrive at the length of one international meter, to the requisite degree of precision. More- over, it 1s possible that yet another method may be found for de- riving the meter locally, without making a pilgrimage to ‘‘Mececa’’ for it. At present, however, it is necessary to make the pil- erimage. If then, we examine to-day any scale, divided rule, or measure of length, we always, either consciously or unconsciously, look towards Sevres. The rule has always been marked off and gradu- ated by comparison with a standard, and usually this standard has been more precise than the copy. If the graduated rule is very rough and imperfect, with a correspondingly low order of pre- cision, we do not have to go far to find a possible standard of reproduction for it; but the finer and more nearly accurate the rule, the higher up in the scale of development we must search for a possible progenitor. The very finest in any country are only obtainable by direct comparison with the national standards. These, in turn, have been derived from Sévres. Thus there is al- ways a lineal succession of standards of length and of mass to every local inch, foot-rule or pound standard that is worthy of the name. THE PROGRESS OF SCIENCE THE PROGRESS OF SCIENCE CURRENT COMMENT By Dr. Epwin E. SLOSsoN Science Service REWARDS FOR WORKING IN- SIDE THE ATOM Two Englishmen, one Dane and one German, are the winners of Nobel prizes in physics and chemistry for 1921 and 1922. The names just an- nounced from Stockholm are Albert Einstein, of Berlin; Neils Bohr, of Copenhagen; Frederick Soddy, of Ox- ford, and Francis William Aston, of Cambridge. This is a striking illus- tration of the unity of science in spite of national divisions, for these four scientists have been in uncon- sidered cooperation trying to solve the same question, the most funda- mental problem of the universe, what is the atom made of. The atom was originally supposed to be the smallest thing possible, the ultimate’ unit of the universe. The ancient Greeks, who were the first to think about the question, concluded that if you kept on cutting up matter into smaller and smaller pieces you must come at length to something too small to be further sub-divided, so they called this smallest of all pos- sible particles the ‘‘atom’’ which means ‘‘uncutable.’? The modern chemist took over this old Greek idea to serve for the combining weights of the elements and likewise assumed that the atom was the limit. But early in the present century, Professor J. J. Thomson, of Cam- bridge, found radioactive matter giv- ing off particles more than a thousand times smaller than the smallest atom, and for this discovery he received the Nobel prize of 1905. This opened up a new field of research that has been diligently prosecuted ever since, especially by British scientists. Pro- fessor Soddy has not only done a large part of this work but he has given a good popular account of what it means in his book, ‘‘Science and Tite.” ? Chemists used to suppose that all the atoms of the same element were exactly alike in weight and every other way, wherever it came from, but this fixed idea has been upset. Soddy found, for instance, that lead from thorium ores is eleven per cent. heavier in its atomic weight than lead from uranium ores. Soddy named these different forms ‘‘isotopes.’’ What are listed in chemical text- books as atomic weights and were supposed to be unvarying turn out to be in many cases averages of several isotopes. Mereury, for instance, which is listed as having an atomic weight of 200.5 consists of six iso- topes with weights varying from 197 to 204. Aston devised an ingenious way of making the atoms record their own atomic weights. He drives a stream of positively chargea particles be- tween the poles of a powerful magnet which deflects them in the degree of their relative weights. When the di- viding streams strike a photographic plate they leave their tracks and from these the mass of the various isotopes can be determined. Chlorine has al- ways been a puzzle to chemists be- cause its atomic weight figured 35.46 instead of a whole number. But sub- jected to the scrutiny of Aston’s ap- paratus it is found to be a mixture of two kinds of chlorine atoms, one weighing exactly 35 and the other exactly 37. The Scandinavian scientist, Bohr, was the first to venture on a picture of the new fashioned atom. We had DR. FRIDJOF NANSEN The distinguished Norwegian Arctic explorer and man of science, who has recently been occupying himself with the relief of sufferers from the Russian famine and is now engaged in similar work in Asia Minor. THE PROGRESS OF SCIENCE been accustomed to think of atoms as round hard balls, but according to Bohr they are more like miniature solar systems with a positive elec- trical nucleus in the center and one or more negative electrical particles, called ‘‘electrons,’’ revolving around it at tremendous speed. Here is where Einstein comes in, for, while the planets moving majes- tically in their orbits obey Newton’s law of gravitation, the electrons, which travel almost as fast as light, deviate from Newton’s law in pro- portion to their speed and follow the formula of Einstein instead. Accord- ing to Newton the mass of a body re- mains the same whatever its motion. According to Hinstein, the mass in- creases with its velocity. The differ- ence between them is inconsiderable for any ordinary speed, but when we are dealing with electrons moving at the rate of 100,000 miles a second it becomes important. The public has associated Einstein exclusively with astronomy because his theory has been tested at a time of eclipse, but the theory of relativity has applica- tions quite as revolutionary and much more practical in earthly chemistry and physics. HOW THE CHEMIST MOVES THE WORLD THE chemist provides the motive power of the world, the world of man, not the inanimate globe. Archimedes said he could move the world if he had a long enough lever. The chemist moves the world with molecules. The chemical reactions of the consumption of food and fuel furnish the energy for our muscles and machines. If the chemist can only get control of the electron, he will be in command of unlimited energy. For in this uni- verse of ours power seems to be in in- verse ratio to size and the minutest things are mightiest. When we handle particles smaller than the atom we can get behind th: elements and may effect more marvel- 583 lous transformations than ever. The smaller the building blocks the great- er the variety of buildings that can be constructed. The chemistry of the past was a kind of cooking. The chemistry of the future will be more like astronomy; but it will be a new and more useful sort of astronomy, such as an astronomer might employ if he had the power to rearrange the solar system by annexing a new planet from some other system or ex- pediting the condensation of a nebula a thousand times. The chemist is not merely a mani- pulator of molecules; he is a manager of mankind. His discoveries and in- ventions, his economies and creations, often transform the conditions of or- dinary life, alter the relations of na- tional power, and shift the currents of thought, but these revolutions are effected so quietly that the chemist does not get the credit for what he accomplishes, and indeed does not usually realize the extent of his sociological influence. For instance, a great change that has come over the world in recent years and has made conditions so un- like those existing in any previous period that historical precedents have no application to the present prob- lems, is the rapid intercommunication of intelligence. Anything that any- body wants to say can be communi- cated to anybody who wants to hear it anywhere in all the wide world within a few minutes, or a few days, or at most a few months. In the agencies by which this is accom- plished, rapid transit by ship, train or automobile, printing, photography, telegraph, and telephone, wired or wireless, chemistry plays an essential part, although it is so unpretentious a part that it rarely receives recogni- tion. For instance, the expansion of literature and the spread of enlight- enment, which put an end to the Dark Ages, is ascribed to the invention of movable type by Gutenberg, or some- body else, at the end of the four- 584 teenth century. But the credit be- longs to the unknown chemist who in- vented the process of making paper. The ancient Romans stamped their bricks and lead pipes with type, but printing had to wait more than a thousand years for a supply of paper. Movable type is not the essential fea- ture of printing, for most of the printing done nowadays is not from movable type, but from solid lines or pages. We could if necessary do away with type and press altogether, and use some photographic method of composition and reproduction, but we could not do without paper The in- vention of wood-pulp paper has done more for the expansion of literature than did the invention of rag paper 600 years ago. Print is only an imperfect repre- sentation of the sound of speech, a particularly imperfect representation in the case of English because we can not tell how half the words sound from their spelling. But the phono- graph gives us sounds directly, and the audion and the radio have extend- ed the range of a speaker, until now a speaker may have an audience covering a continent and including generations yet unborn. What these inventions do for sound, photography has done for the sister sense of light. By means of them man is able to transcend the limitations of time and space. He can make himself seen and heard all round the earth and to all future years. THE OOST OF NIAGARA Ir a man stood on the banks of the Mississippi at the time of the spring freshet, when the stream was carry- ing down to the Gulf fences, pigs, chickens, furniture and, occasionally, a house, he would be seriously con- cerned over the loss of the property of those who had so little to lose, and perhaps exert himself to save some of it; but the continuous calamity of Niagara arouses in him no feelings of a nature to mar his THE SCIENTIFIC MONTHLY enjoyment. He shows the same esthetic appreciation of a sublime and beautiful spectacle and the same indifference to its cost as Nero at the burning of Rome. It is easier to comprehend how much it is costing us to keep up Nia- gara as a spectacle if we put the waste in concrete terms. Various en- gineers have estimated that it would | be possible to get from Niagara Falls over 5,000,000 more horse-power than is now utilized. In one of the large steam plants of New York City the cost of power is $50 a year per horse- power. Taking these figures as suf- ficiently close for our purpose the water that goes over the Falls repre- sents the annihilation of potential wealth at the rate of some $250,000,000 a year or nearly $30,000 an hour. We are told that there are some millions of people in poverty and poorly nourished in this country, yet here is wasted the equivalent of 250,000 loaves of bread an hour. We may see with our mind’s eye 600,000 nice fresh eggs dropping over the precipice every hour and making a gigantic omelet in the whirlpool. If calico were continuously pouring from the looms in a stream 4,000 feet wide like Niagara River it would represent the same destruction of property. If a Oarnegie Library were held under the spout it would be filled with good books in an hour or two. Or we can imagine a big department store float- ing down from Lake Erie every day and smashing its varied contents on the rocks 160 feet below. That would | be an exceedingly interesting and di- verting spectacle, quite as attractive to the crowd as the present, and no more expensive to maintain. Yet some people might object to that on the ground of extravagance who now object to the utilization of the power of the falling water. It must not be supposed that I am insensible to the beauties of nature or ignore their esthetic and cultural THE PROGRESS OF SCIENCE value. On the contrary, I would wish to enhance the interest and impres- siveness of Niagara Falls by making it a rarer spectacle. The reason why people fail to appreciate the beauty of the clouds, of the sunset and of the landscape from their windows is because these are so common. If a bouquet of fireworks were shot off at eight o’clock every night we would not care to look at them. Of course the Falls would be turned on for all legal holidays and as often as there was sufficient demand for it. On such occasions those who wished to go down the current in barrels could en- joy their favorite sport. Weddings would naturally be arranged to come off at a time when the Falls fell. At the hours when the water was pro- hibited from making a run on the banks, rambles over the eroded rocks and worn channels would be of great interest to the geologist and the tour- ist. Couples and groups could be photographed at the Falls then, as they are now, by posing them in front of a painted screen. Many more people would see Nia- gara and their enjoyment of it would be much greater if it could be seen only on fete days. Thinking they could see it any time, thousands of people have neglected it in favor of some passing show. Of course, there is something im- pressive in the thought that the flood pours thundering into the abyss all of the time regardless of sight-seers. But if one has not sufficient imagina- tion to find an equal emotional value in the contemplation of the varied life and industry it supports as it pours through the penstocks and spins the turbines he can swell with satisfac-. tion on the thought of the thousands of years when it was of no use to any- body. In 18938, when Lord Kelvin stood on the brink of Niagara, he was not so much impressed by its grandeur as he was saddened by the sight of such an enormous waste of power, tauri. 585 and’ he expressed the hope that he would live to see it all utilized, an observation which was much ridiculed at the time by hard-hearted sentimen- talists and unimaginative poets. To them Niagara was a mere spectacle, but to the great scientist, who had devoted his life to the study and ex- position of the law of the conserva- tion of energy, it was much more. His prophetic eye could see the poor who might be enriched, the homes that could be made happy, the hungry who might be fed, the naked who might be clothed, and the toiling mil- lions who might be relieved of their burdens by the water dashing upon the rocks below for the amusement of idle tourists. LOOK OUT FOR ALPHA CEN- TAURI As if we did not have enough to worry about, what with winter com- ing on and coal so short and clothing so high, here comes along Professor Ellsworth Huntington, of Yale, with a book on ‘‘Climate Changes’’ which warns us that the stars in their courses may fight against us. He has a theory that the glacial epochs and the lesser disturbances of the earth’s climate are largely due to prior dis- turbances in the- sun’s atmosphere and these in “turn may be caused by the approach or increased activity of certain stars. All the stars, includ- ing our sun, are in radio communica- tion with one another, and when one flares up over something it arouses responsible excitement in all the others within range. Then, too, the stars are not ‘‘fixed,’’? as we used to think, but are wandering about in various directions, and when two stars come close enough together they be- come mutually inflamed by the proxi- mity and may become permanently attached. Now the nearest star to us is the brightest one in the Centaur constel- lation, therefore named Alpha Cen- It is only about 25 trillion 586 THE SCIENTIFIC MONTHLY Wide World Photos THE ASTROPHYSICAL OBSERVATORY BUILT FOR PROFESSOR ALBERT EINSTEIN AT POTSDAM miles away and its light takes four | sunspots, and would affect the and a third years to reach us. Alpha | weather on the earth. Centauri is not only big and bright The dates when the two bright and relatively near, but it is triple | spheres of Alpha Centauri were near- and variable. Its two main com- | est together and most radiant are ponents are like two suns the size of | 81.2 years apart and these fall on ours, revolving around one another | the years 1388, 1469, 1550, 1631, every 81.2 years. When they are | 1713, 1794, 1875 and 1956.° Com- closest they are 1,100,000,000 miles | paring these with the records of sun- apart and when their orbits separate | spots, which have been kept only for most widely they are three times as | the last century and a half, we see far as that from each other. It is | that such evidences of solar disturb- when the twin stars are nearest that | ances were most evident in periods we should expect them to be most | ending in 1794 and 1875, and that active in sending out light waves and | another period of high solar activity electrons. These reaching the sun | started in 1914 and may be expected might set up wild whirlings in the | to end about 1956. solar atmosphere, which would appear | If this theory of stellar influence to us as an unusual abundance of | is true we may expect something to THE PROGRESS OF SCIENCE happen somewhere between 1950 and 1956. What it will be Professor Huntington does not venture to sur- mise, but he reminds us that in the years preceding 1388, when Alpha Centauri was active, Europe was a very uncomfortable place to live in. There were droughts and floods, fa- mines and freezings. The Baltic was frozen so that horse sleighs could cross from Germany to Sweden, and the Danube and the Rhine sometimes inundated the cities on their banks and sometimes nearly dried up. There are more serious grounds for suspecting Alpha Centauri of a malign influence on the earth for that star was nearest to the earth 28,000 years ago, being then only 3.2 light- years away. Now this is the date that geologists have set for the end of the last Great Ice Age so the ap- proach and proximity of Alpha Cen- taurl may have had something to do with that spell of cold weather which came near freezing out the human race. The world is even yet con- valescing from the chills of the Gla- 587 cial Epoch. Greenland which onee was really green with ferns and figs is still covered by an ice cap. We need not fear another glacial age from the same cause for Alpha Centauri is now 4.3 light-years away and leaving us at the rate of thirteen miles a second. But Sirius is due in this vicinity in 65,000 years and that would be quite as—I should say, might be equally—bad for us. But Professor Huntington en- deavors to console us by reminding us that the human race not only sur- vived several such periods of climatic stress, but has come out of them in each case stronger and better for the struggle for existence. He is a firm believer in the value of stormy weather. He is a New Englander. NEW LIGHT ON THE ORIGIN OF LIFE Was the first living being a plant or animal? How could either origi- nate out of non-existing matter? These that have They could are questions hitherto baffled scientists. World Photos A LABORATORY OF THE ASTROPHYSICAL OBSERVATORY BUILT FOR PROFESSOR EINSTEIN Apepousypg ye Aurdwoy orsyoopy pexouay oy} Jo SOlIOPVIOGVT YIIvasoy oy} UT ZLAWNIGLS ‘d SATHVHO ‘Wd CNV NOSIGH ‘V SVNOHL ‘UN 80104 PLO A apr4f THE PROGRESS OF SCIENCE trace back, more or less satisfactorily, the lines of development of plants and animals to the simplest and most primitive forms of life, but there they ran up against an insurmount- able wall, on the near side of which was the world of living organisms and on the far side the world of inert mineral and inorganic matter. We all know that non-living matter can be converted over into living matter for we do that ourselves when- ever we eat or breathe. We all know that green plants have the power of building up sugar and starch and wood (the so-called carbohydrates) out of the water of the soil and earbon dioxide of the air, for we can see them do it any sunny day. But it is life only that can bring into the living organism this inorganic material. Water and carbon dioxide, plain ‘‘soda water,’’ do not spon- taneously change over into sugar or start to grow into a plant. It re- quires green colored granules of the leaves, called chlorophyll, to effect this transformation. But chlorophyll is a very compli- cated chemical compound. It is formed ouly by green plants as they develop in the sun’s rays from white sprouts. So the plant must exist be- fore chlorophyll is formed. But, on the other hand, a plant could not exist unless it got its energy from the sugar and other stuff stored up previously by some chlorophyll-bear- ing plant. Even the simplest green plant can not Hve and grow on its nutritive salts in the sunshine unless it has a bit of plant-stuff to feed on as a starter. We might surmise as a way out of the dilemma that animal life came first on the earth, and, in decaying, supplied the primitive plants with the necessary organic food stuff. But here we are blocked because animals are parasites of plants. They live on the sugars and so forth that the green leaves have stored up by means of sunshine. 589 So this was the perplexing situa- tion. Plants can feed on animals or other plants. Animals can feed on plants or other animals. But where could the first animals or plants get their food when there was nothing but mineral matter in the world? It was worse than the old question, which came first, the hen or the egg? But of late we are beginning to get light on the problem. The wall between the living and non-living is crumbling. Certain sugars and pro- teins, such as the plant forms that we eat, can now be made in the lab- oratory out of inorganic material. Artificial cells have been constructed that grow and crawl and feed them- selves and stick out feelers and sub- divide very much like living cells. It has been found that ultra-violet rays, that is, light of such short waves that it can not be seen, can convert water and carbon dioxide into sugar as chlorophyll does. These short waves are not con- tained in the sunshine that reaches the earth to-day, but it is found that ordinary rays may act the same way in the presence of certain substances such as iron rust in water. These same energetic rays are able to incor- porate the nitrogen of mineral salts into compounds like the protein of the living cell. So here we see the possibility that the action of the sun- light on the sea in primordial periods —or even in the present—might pro- duce sufficient food to give a single cell a start in life and enable it to grow and multiply and develop into - other and higher forms. But how this primal cell got to going in this way the biologists are only beginning to surmise. Dr. E. J. Allen, at the recent Hull meeting of the British Association for the Ad- vanecement of Science, ventures the theory that the first organism was of the animal sort and spherical shape, but that it gradually grew a tail or whip that enabled it to rise to the sunny surface of the sea whenever it 590 sank below and that it there acquired the chlorophyll by which it could make its own food out of the air and water. This is far from knowing what did happen in those early days, but it is a great advance to be able even to speculate as to how it might have happened since not many years ago it seemed that it happen at all. could not SCIENTIFIC ITEMS WE record with regret the death of Robert Wheeler Willson, emeritus professor of astronomy at Harvard University; of Guy Henry Cox, for- merly professor of geology at the Missouri School of Mines; of Dr. Chauncey William Waggoner, head of the department of physics in West Virginia University; of F. T. Trou- ton, emeritus professor of physics in the University of London, and of E. Bergmann, director of the Chemisch- Technische Reichsanstalt, Berlin. THE Henry Jacob Bigelow medal of the Boston Surgical Society was presented to Dr. William W. Keen, of Philadelphia ‘‘for conspicuous contributions to the advancement of surgery,’’ on the evening of October 25, when Dr. Keen addressed the so- ciety on ‘‘Sixty years of surgery, 1862-1922.’ On the occasion of the celebration of the fiftieth anniversary of the Dutch Zoological Society there were admitted as honorary members: Pro- fessor O. Abel, Vienna; Professor M. Caullery, Paris; Professor L. Dol- lo, Brussels; Professor B. Grassi, Rome; Professor V. Hacker, Halle; Professor 8. J. Hickson, Manchester ; Professor N. Holmgren, Stockholm; Professor T. H. Morgan, New York; Dr. F. Sarasin, Basle, and Dr. J. Schmidt, Copenhagen. Foster Haun, the chemical labora- tory of the University of Buffalo, THE SCIENTIFIC MONTHLY designed especially to meet the needs of the electro-chemical, hydro-elec- tric, dye and steel industries on the Niagara frontier, was dedicated on October 27 in connection with the in- stallation of Dr. Samuel P. Capen, of Washington, as chancellor of the university. Dr. Edgar F. Smith, president of the American Chemical - Society, and Dr. Edwin E. Slosson, of Science Service, were speakers at the ceremony. The laboratory, erected at a cost of a million dollars, is the gift of O. E. Foster, of But- falo. In the will of Prince Albert of Monaco, who died on June 26 last, there are several gifts for scientific purposes. His farm at Sainte Su- zanne is left to the French Academy of Agriculture, and the wish is ex- pressed that the estate should re- main a place for agricultural experi- ments, to demonstrate what science can obtain from sterile lands. Dr. Jules Richard will receive 600,000 francs to enable him to complete literary and scientific works in prog- ress, including the results of the oceanographic cruises and the prepa- ration of the Bathymetric Chart of the Oceans. The proceeds of the sale of the yacht Hirondelle, all books and publications of a scientific nature, as well as certain personal effects, will go to the Oceanographic Institutes at Paris and Monaco, while the In- stitute of Human Paleontology in Paris is to receive any personal ef- fects relating to the work carried on there. The Paris Academy of Sciences will receive a million franes, the income of which is to provide a prize to be awarded every two years, the nature of the prize to be indicated by the academy, according to the needs of the moment; a like sum is bequeathed to the Academy of Medi- cine for a similar prize. INDEX TO VOLUME XV 591 INDEX NAMES OF CONTRIBUTORS ARE PRINTED IN SMALL CAPITALS Albatrosses, Our Great Rovers of the High Seas, R. W. SHUFELDT, 469 Alcohol and Gasoline, 188 Alpha Centauri, Look out for, 585 ANDERS, JAMES M., City Parks and Playgrounds as Health Agents, 42 Anopluris, G. F. Frrris, 551 Ants, Wi~tt1am Morton WHEELER, 385, 527 Astronomy in Canada, Orro Kuorz, 215 Atom, Modern Study of, ALAN W. C. Menzigs, 364; of Light, 377; Re- wards for Working Inside the, 581 Austraha, Vegetation of, D. H. CAMPBELL, 481 : Bacot, Martyr to Science, ARTHUR H. SMitH, 359 Bees, Solitary and Social, WILLIAM Morton WHEELER, 235, 320 Berry, Epwarp W., Geologie Evi- dence of Evolution, 97 Brown, W. Norman, Tar-Baby Story at Home, 228 Bureau of Weights and Measures, International, A. E. KENNELLY, 570 Calendar Reform, 91 Calorie in Nutrition, History of MILDRED R. ZIEGLER, 520 CAMPBELL, D. H., Vegetation of Aus- tralia and New Zealand, 481 Canada, Astronomy in, Orro Kuorz, 215 Character Reading from External Signs, Knigur DuNuLap, 153 Chemist, how he moves the world, 483 City Parks and Playgrounds as Health Agents, JAMES M. ANDERS, 42 ? Cost of Niagara, 583 DapouriAn, H. M., Some Problems of Progress, 348 DANForRTH, RALPH E., Path as Factor in Human Evolution, 338 Darwin, Charles, ADDISON GULICK, 132 Davis, W. M., Reasonableness of Science, 193; Topographical Maps of United Staites, 557 De Anopluris, G. F. Frrris, 551 Dovuetass, A. E., Annual Rings of Trees in Climatic Study, 5 DunuLAPp Knicut, Reading Character from External Signs, 153 Easy Group Theory, G. A. MILLER, 512 Eclipse, Relativity and the, 473 Kels, Birthplace of, 381 Einstein and Time Line, 475 EVERMANN, BartoN WARREN, Con- servation and Utilization ot Nat- ural Resources, 289 Evolution, Geologic Evidence of, Ep- WARD W. Berry, 97; Working Backward, 377 FELT, E. P., Kxterminating Insects, Possibility of, 35 Ferris, G. F., De Anopluris, 551 Finnish Poetry, EUGENE VAN CLEEF, 50 Food Resources of the Sea, GEORGE W. Martin, 455 Galen, Claudius, Mentality and Cos- mology of, JONATHAN WRIGHT, 144 Gasoline and Alcohol, 188 GaTEs, CuirrorD E., Polynesians, Caueasians of Pacific, 257 Glands and Complexes, 191 Go to the Bee, Davin Starr JORDAN, 448 Group Theory, Easy, G. A. MILLER, 512 GULICK, ADDISON, Charles Darwin the Man, 132 HANNA, G. Dauuas, Reindeer Herds of Pribiloff Islands, 181 Health Agents, City Parks and Play- grounds as, JAMES M. ANDERS, 42 HENDERSON, LAWRENCE J., Water, 405 HOVERSTOCK, GERTRUDE, and STEPHEN S. VisHer, ‘‘Who’s Who’’ among American Women, 443 Imagination, Scientific, WALTER Lipsy, 263 Immigration: Restriction, ROBERT DeC. WarD, 313; Intelligence Tests of Immigrant Groups, KIMBALL Youne, 417; Legislation, Rosert DEC. Warp, 561 Insects, Social Life among, WILLIAM Morton WHEELER, 68, 119, 235, 320, 385, 527;Exterminating, E. P. FELT, 35 Intelligence, LIGHTNER WITMER, 57 International Intellectual Coopera- tion, 89 JORDAN, Davin Srarr, Go to the Bee, 448 Keeping Cool, 187 KENNELLY, A. E., Modern Mecea, 571 Kuiorz, Orro, Astronomy in Canada, 215 592 THE SCIENTIFIC MONTHLY e La Rue, DanizL WotrorD, The Shorthand Alphabet and the Re- forming of Language, 271 Lippy, WALTER, Conceptual Think- ing, 435 Life, Origin of, 587 Light, Atoms of, 377 Macrosiphum Solanifolii, Epitn M. Patcu, 166 Maps, Topographical, of United States, Wi~tiam Morris Davis, 557 Marine Fisheries, State and Biologist, WILLIAM F.. THoMpPson, 542 MartTIN GEORGE W., Food Resources of Sea, 455 Menzies, ALAN W. C., Modern Study of Atom, 364 Mecea, Modern, A. E. KENNELLY, 571 Microbes and Man, 379 MinuEr, G. A., Easy Group Theory, 512 Mind-Cleaning, 287 Naming, New Inventions, 285 Natural Resources, Conservation and Utilization of, Barton’ WARREN EVERMANN, 289 New Zealand, Vegetation of, D. H. CAMPBELL, 481 Niagara, Cost of, 584 Nutrition, History of Calorie in, MinprRED R. ZIEGLER, 520 Origin of Life, 584 Patou, Epira M., Marooned in a Potato Field, 166 Path as Factor in Human Evolution, RautpH E. DANFORTH, 338 Polynesians, Caucasians of Pacific, CLIFFORD E, GATES, 257 Potato Field, Marooned in, EpirH M. PatcH, 166 Problems of Progress, Some, H. M. DADOURIAN, 348 Progress of Science, 89, 187, 282, 377, 473, 581 Reindeer Herds of Pribiloff Islands, G. DALLAS HANNA, 181 Relativity and Eclipse, 473 Science, Reasonableness of, W. M. Davis, 193; of Keeping Cool, 187 Scientific Items, 96, 191, 288, 384, 480, 585; Imagination, WALTER Lippy, 263 Shorthand Alphabet and the Reform- ing of Language, DANIEL WOL- FORD LA Rug, 271 SHUFELDT, R. W., Albatrosses, Our Great Rovers of the High Seas, 469 SmirH, ARTHUR H., Bacot, Martyr to Science, 359 Solar Eclipse of September 21, 383 Sunspots, Invisible, 91 Tar-Baby Story at Home, W. Nor- MAN Brown, 22 ; Thinking, Conceptual, WALTER LisBy, 435 THOMPSON, WILLIAM F., Marine Fisheries, State and Biologist, 542 Tricks of Mediums, 285 Tropies, Variability vs. Uniformity in, STEPHEN SARGENT VISHER, 22 Twins, 93 Topographical Maps of United States, Wui~uiAM Morris Davis, 557 VAN CLEEF, EUGENE, Finnish Poetry, Nature’s Mirror, 50 Variability vs. Uniformity in Tropics, STEPHEN SARGENT VISHER, 22 Vegetation of Australia and New Zealand, D. H. CAMPBELL, 481 VISHER STEPHEN SARGENT, Variabil- ity vs. Uniformity in Tropics, 22; and GERTRUDE HOVERSTOCK, ‘“‘Who’s Who’? among American Women, 443 WALLIS WiLtson D., Why do we Laugh? 343 Ward, Rosert DEC., Some Thoughts on Immigration Restriction, 313; What Next in Immigration Legis- lation, 561 Wasps, Solitary and Social, WILLIAM Morton WHEELER, 68, 119 Water, LAWRENCE J. HENDERSON, 405; Value of, 282 What is Intelligence and Who has it? LIGHTNER WITMER, 957 WHEELER, WILLIAM Morton, Social Life among Insects, 69, 119, 235, 320, 385, 527 ‘‘Who’s Who’’. among American Women, STEPHEN S. VISHER and GERTRUDE HOVERSTOCK, 443 Why do we Laugh? Winson D. WALLIS, 343 Witmer, LigHTtNER, What is Intelli- gence and Who has it? 57 WRIGHT JONATHAN, Mentality and Cosmology of Claudius Galen, 144 YounG, KimBauu, Intelligence Tests of Immigrant Groups, 417 ZIEGLER, MiLpRED R., History of Calorie in Nutrition, 520 . en .) BINDING LIST NOVA 1933 Q The Scientific monthly 1 $817 veld er: hysical & Applied Sci, Serials PLEASE DO NOT REMOVE CARDS OR SLIPS FROM THIS POCKET UNIVERSITY OF TORONTO LIBRARY d/h 417) 413) i x i a pharma - a Lo fe-emcts Nie ea Be 4 = PER = oe iets ch sta a — ' a - + S x ~ en — > ST - ‘ > * < ¥ : 2 ay il aor * ~ . bs a ‘ en 3 ) 1 - “4 — > - a ‘ 7 =~ - s « us “ - 1 oe ‘ +4 > i t To ae ? - ; ai -; ¥ ‘ a % : > 3 Pe - { = » . - - ” - t sen 2 if - . » ‘ : 1 ; > a ‘ s . ‘ t = ¢ a z A . Fete Savy teeny es tas