Spring 2015 M The Quarterly of the Virginia Native Plant Society

Upside-down Anthers of Clethra Stand Out

By W. John Hayden, Botany Chair; illustrations by Sheila M. Hayden

or the most part, the flowers of
the 2015 VNPS Wildflower of
the Year, Clethra alnifolia (Sweet
Pepperbush), are unremarkable. Five
separate sepals, 5 separate petals, 10
stamens in 2 whorls, and a 3-carpel-
late superior ovary — an organization
that can only be considered prosaic
among the dicots. One floral feature,
however, stands out: the anthers in
the open flowers are upside-down!
(See Figure 1A.) Further, these
upside-down anthers open by pores
(Figures IB, 1C) rather than longi-
tudinal slits, as in most flowering
plants. These pores initially form on
what would normally be the lower-
most extremity of the anther, the

inversion of which brings the pores to
a forward position in the open flower.
The significance of these acrobatic
anthers — found not just in Sweet Pep-
perbush but through all 65 members
of the genus Clethra — is twofold:
anther inversion is essential to the
plant’s pollen-presentation strategy,
and the odd morphology taxonomi-
cally links Clethra with the large and
diverse heath family (Ericaceae).

Barnes (1880) provided one of the
first descriptions of anther inversion
in Clethra. In flower bud stages, the
anthers of Clethra alnifolia take on a
sagittate shape (Figure 1A), narrower
at the apex and flared outward
toward the base. Before the flower

bud opens, the future pore for pollen
release is located on the external
(petal) side of the anther, a position
technically described as extrorse. As
the bud opens, the staminal filaments
rapidly elongate, pushing the anthers
past the opening petals. At the same
time, the anthers begin to invert,
quickly becoming horizontal,
with their tips temporarily
pointing toward the center of the
flower. From the rapidly achieved
horizontal position, complete
inversion proceeds a bit more
slowly (over the course of a few
minutes), so that the narrow tip
of each anther now points toward
the base of the flower (Figures 1C,
ID). An additional consequence of
anther inversion is that the pores for
pollen release, formerly extrorse in
orientation, become introrse, i.e.,
positioned on the stigma side of the
now fully inverted anther.

Flow do these anthers invert?
Barnes (1880) attributes the anther
acrobatics of Clethra to cellular
processes in the connective, the tissue
located at the apex of the filament
that serves to attach (connect) anther
and filament together. In most plants,
connective cells are uniform, but in
Clethra , connective cells vary from
longer, thin-walled cells containing
oil droplets to shorter, thick-walled
cells devoid of oil. In unopened flower
buds, before anther inversion, the

For more on Clethra, see page 3

Fig. 1. (A) Flower of Clethra alnifolia. (B) Immature anther of Clethra, before inversion,
as viewed from external (petal) side; asterisk (*) indicates initial basal position of
dehiscence pore. (C) Mature anther of Clethra, after inversion, as viewed from internal
(stigma) side; asterisk (*) indicates ultimate terminal position of dehiscence pore.

(D) Mature anther of Clethra, after inversion, as viewed from external (petal) side.

(E) Mature inverted anther of Arctostaphylos uva-ursi, with appendages.

2

Sempervirem , Spring 2015

From the President

Gift to Natural

Heritage Caps Great Day

W e have a surprise for you,” said Stacey Loulies,

of the Virginia Department of Conservation and
Recreation, greeting a group of us from the Virginia Native
Plant Society, in Richmond for a check presentation. As a
result of the generosity of VNPS members and chapters,
we raised $11,542.75 for DCR’s Division of Natural
Heritage to search for “lost natural treasures” — rare plant
species that have not been seen for years at sites of earlier
observations. We had printed an oversize check for formal
presentation to DCR staff, including Director Clyde
Cristman. He thought it would be a good idea to include
Secretary of Natural Resources Molly Ward, so it was a
challenge to coordinate our schedules for Monday, January
26, at 11 a.m. at DCR’s offices.

So, what was the surprise? Well, a special guest had
accepted an invitation to participate in the check presen-
tation: Gov. Terry McAuliffe. The event was rescheduled
for 3 p.m. That left Suzanne Dingwell, Joanne Hutton,
and me with time to tour Natural Heritage’s herbarium
and zoology lab with chief biologist and Flora of Virgin-
ia co-author Chris Ludwig. Their plant specimens were
pressed in newspapers and labeled with the collection date
and location. In the zoology lab we saw mounted moth
specimens that showed their intricately detailed wings
and identifying characteristics. We are so fortunate to

VIRGINIA NATIVE
PLANT SOCIETY

SsTYlpCTVlTCnS (ISSN 1085-9632) is the quarterly newsletter of
the Virginia Native Plant Society, Blandy Experimental Farm, 400
Blandy Farm Fane, Unit 2, Boyce, Va. 22620, 540-837-1600, info@
vnps.org.

Nancy Vehrs, President
Nancy Sorrells, Editor
Karen York, Office Manager

Original material in Sempervirens may be reprinted if credit is
given to the Virginia Native Plant Society, to Sempervirens, and
to the author of the material, if named. Readers are invited to send
letters, news items, and queries for consideration. E-mail items to
Nancy Sorrells at lotswife@comcast.net.

Next submission deadline: May 1, 2015

Commonwealth Connection: At the presentation were Ludwig,
Smith, me, Cristman, McAuliffe, Johnny Townsend, Hutton,
Dingwell, and Ward. (Photograph by Michaele White, Office of
the Governor)

have dedicated scientists who find, study, and catalog the
diverse species in our beautiful commonwealth.

After our DNH tour, Sue, Joanne, and I had busi-
ness at the Department of Environmental Quality, across
the street, where we met with Virginia Witmer from the
Coastal Zone Management Program. She is the force
behind Virginia’s native plant marketing campaign and
the production of the colorful native plant guides for the
Eastern Shore, Northern Neck, and Northern Virginia.
The Plant NoVA Natives guide was first printed last fall,
and we had revisions to discuss with Virginia for the sec-
ond printing, due out in March. While at DEQ, I received
word from Tom Smith, director of Natural Heritage, that
it was time for us to meet with the governor.

Hastily donning our coats, we joined the DCR group
to race several blocks uphill to the governor’s office. After
making our way through security, we were escorted up-
stairs to the governor’s conference room, where Secretary
Ward met us. As we settled around the conference table
with the big check in front of us, the governor joined us in
a carefully choreographed manner. We chit-chatted about
the fund-raiser, the important work of Natural Heritage,
and its close relationship with the VNPS. We also took the
opportunity to present the governor with a Plant NoVA
Natives guide, since he has a home in McLean.

Since the General Assembly Building was next-door,
we visited my state senator, Chuck Colgan, co-chairman
of the Senate Finance Committee. We urged him to vote
for a budget amendment that would increase funding
for the Natural Heritage Program. While we received no
promises, we later learned that GA had approved an addi-
tional $500,000. — Your President, Nancy Vehrs

Sempervirens ; Spring 2015

3

The Acrobatic Anthers of Sweet Pepperbush

Continued from page 1
long, thin-walled cells are located on
the petal side of the connective. As
described by Barnes, bud opening
brings relatively dry air into contact
with the thin-walled connective cells,
which collapse, presumably from
evaporative water loss. This collapse
of these thin-walled cells pulls the tip
of the anther downward, accomplish-
ing anther inversion as the bud opens.

So, if the anthers of Clethra did
not invert, pollen would be shed
extrorsely (petal side) from the anther
bottoms. But since they do invert,
pollen is shed introrsely (stigma side)
from the original bottom of the an-
ther that in open flowers occupies the
extreme tip of the stamen. Arguably,
pushing the pollen-releasing pores to
a foremost position makes sense for
efficient transfer to pollinators. In
Clethra alnifolia, pollen is shed soon
after flowers open. In other species,
for example Clethra acuminata,
pollen is shed before the flowers are
fully open. In either case, these flow-
ers are protandrous, shedding pollen
before stigmas become receptive, an
adaptation that promotes cross-polli-
nation. Later, styles elongate, pushing
stigmas beyond the level occupied by
the anthers (Figure 1A), at which point
stigmas become receptive to pollen.
Presumably, introrse orientation of the
inverted anthers allows for self-polli-
nation, which has been documented,
should cross-pollination fail.

Inverted porous anthers also occur
in the heath family, and the shared
presence of this unusual character
is part of the evidence indicating a
close relationship between these two
families. See, for example, the stamen
of Arctostaphyllos uva-ursi (Bear-
berry) in Figure IE. But the anthers
of Ericaceae are not identical to those

Clethra alnifolia in flower. (Photograph by
W. John Hayden)

of Clethra. In most Ericaceae, anther
inversion takes place much earlier
in flower development, when flower
buds are still quite small (Hermann
and Palser 2000). Further, anthers of
Ericaceae are usually equipped with
slender appendages (Figure IE) that
are absent in Clethra. These anther
appendages come in two forms: awns
(as defined in this context) are hollow,
elongate extensions of the pollen-
bearing region, whereas spurs are
solid and develop independently of
the pollen-bearing anther sacs.

Regardless of their internal struc-
ture and how they originate, anther
appendages in Ericaceae are critical
for pollination. Typical ericaceous
flowers are urceolate (urnlike) and
pendent, which forces pollinators to
probe the flower from below. As the
insect, typically a bee, rummages
around, it inevitably contacts one or
more anther appendages, which act
as tiny levers, causing pollen to sift
out of the porous anthers onto the

probing pollinator; when it moves on
to another flower, some of that pollen
will be deposited on its stigma.

As is often the case in compar-
ative biology, the details are a little
more complicated than the simple
overview described above. Ericaceae
is a large family, and intermediate
states of anther modifications have
been documented. For example, in
Arbutus and Enkianthus, anther
inversion happens late, as the flower
buds open, pretty much as in Cleth-
ra. In Cassiope, partial anther inver-
sion happens a little earlier, shortly
before bud break (Hermann and
Palser 2000) and reaches completion
as the flower becomes fully open.
Anthers of Enkianthus also differ
from those of all other Ericaceae in
that they open by longitudinal slits,
not pores. And some of the most
familiar members of Ericaceae have
nonurceolate corollas; examples are
Rhododendron (azaleas and rhodo-
dendrons) and Kalmia (Mountain
Laurel). These genera have com-
pletely different pollen-presentation
strategies and, correlated with that
fact, they have no anther appendag-
es. So the situation regarding anther
structure and development is rela-
tively complicated in Ericaceae. Nev-
ertheless, the acrobatic anthers of
Clethra provide — conceptually — a
reasonable starting point for inter-
preting ericaceous anther diversity
and the peculiar pollination mecha-
nism characteristic of those members
of the family that possess urceolate
corollas.

Prosaic flowers? Clethra ? Hardly!
Its acrobatic inverted anthers earn
Sweet Pepperbush a place of distinc-
tion among flowering plants, and a
noteworthy addition to our Society’s
roster of beloved native plants. ❖

4

VNPS Legend Looks Back

A Visit with Nicky

N icky Staunton has achieved legendary status in the Virginia Native
Plant Society. She is a charter member who served three terms as
president and many years in other offices. An accomplished amateur
botanist, she is also an extraordinary artist. She produced the whimsical
watercolors that illustrate the new book Isabella’s Peppermint Flowers,
an introduction to botany for children. Nicky was president of the
Prince William Wildflower Society when I joined in 1987 and has been
an inspiration to me and many others. Join us in our visit. — Society
President Nancy Vehrs

How did your love of
wildflowers and native
plants start ? Can you trace
it to a single event or time?
Did any individuals play a
significant role ?

At the age of 5, when
I was told to go outside
and play, the flowers in
Grandmother’s garden were
my playmates. I remember
sitting at the base of a huge
old red oak and watching
ants, and lolling on top
of a low evergreen hedge
around the garden and
watching cloud formations.
In 1957, at Shenandoah
National Park, I learned
“wildflower” in a natural
setting. I remember a
sign that said don’t pick
wildflowers. Until
then they had just been
flowers to me. Native
plants became separate
from wildflowers in 1989,
as the VWPS [Virginia
Wildflower Preservation
Society] became the VNPS.
Eventually native plants and
nonnative plants became
defined. I became president

of VNPS in 1989 as the
VNPS Board of Directors
changed the name to reflect
native plant and habitat
conservation issues.

Many significant
individuals definitely
played a role:

• Grandparents who
believed in sending me
outdoors to play in nature.

• My husband Loren’s
encouragement to “see”
and photograph plants,
beginning in 1957
with my first serious
cameras. Currently,

Lou, our daughter, is my
encourager to capture
what I see with my camera
and botanical art.

• Charlotte Turner, artist
and art professor at
Northern Virginia
Community College,
Manassas, who taught me
several years of drawing
with graphite and pen-
and-ink. That built on my
studies of art history at
Mary Washington.

• Marion Lobstein in the
1980s, professor of botany

at NVCC, who taught me
how to identify any plant I
saw through keying, using
Newcomb’s Wildflower
Guide. Marion included
me as the illustrator for
Susan Leopold’s Isabella’s
Peppermint Flowers. Our
goal was to plant a seed
to spark a child’s interest
in basic botany through
one species of plant within
Virginia history.

Marion also supported
my representing VNPS
on the Llora of Virginia
Project board of directors
[on which Nicky served
from 2001 to 2013], which
was an opportunity to
ignite and fuel enthusiasm
in VNPS members to
support the Llora Project
through contributions of
funds and now as the Llora
Project moves to the next
level of producing its App.

You were VNPS
president for three terms
and a member of the
Executive Committee
for many years. What

Nicky Staunton

are some of the VNPS
accomplishments you’re
proudest of?

Satisfaction would
better describe my feelings
as I revisit my years
with VNPS. Satisfaction
by being a facilitator
for development of
membership, partnerships,
and programs. Our
mission of conserving
wildflowers and wild
places, now interpreted as
conserving native plants
and community habitats,
is the unique focus of our
members and, especially,
committed leaders of the
chapters and the state
organization.

VNPS members are the
“mind, hands, and feet” of
VNPS: professional and
nonprofessional botanists
(education); gardeners,
both professional and non-
professional (horticulture,
Master Gardeners);
natural-resource managers
and users (conservation,
including advocacy), field
trips, photography and art,

Sempervirens ; Spring 2015

5

Staunton, right, and VNPS Past President Sally Anderson, at
a tree dedication honoring Sally at the State Arboretum of
Virginia. (Photograph by Nancy Sorrells)

inventory and species
descriptions, botanical
history and heroes, original
illustrations of many of the
plants, and the standard
for all future knowledge
of flora habitats, as well
as efforts to communicate
this knowledge through
electronic channels.

• The Registry Program
to recognize rare natural
habitat lands that do not
yet qualify for acquisition
by the Virginia Natural
Heritage Program or The
Nature Conservancy.

• Shift of focus from rescue
or relocation of plants to
natural area protection.

What are you doing, now
that you no longer serve on
the VNPS and Flora Project
boards?

I am focusing on
botanical art. Through my
working career, there never
seemed time enough. The
advocacy for our native
plants and their habitats is
ingrained in my nature by
now and will continue.

Where do you see the
VNPS going in the next 10,
20, or 50 years?

Whatever years VNPS
exists depends on the total
commitment of members
to supporting a strong
leadership and growth,
along with the drive to
educate ourselves and
others to appreciate and
want to protect natural
habitats and our native
plants, along with their
fauna, including humans.
Frankly, I believe 10 years

plus the botanists, Master
Naturalists, and other
members with statewide
credibility. A strong
organization does not
reflect accomplishments
of one person or programs
developed while a person
was in office. It does reflect
how the person can effect
program success with those
members.

All of the following
satisfactions grew from
members coming together
for native plant and
habitat preservation, work
that began in the early

1900s by the Wild Flower
Preservation Society, a small
group of botanists, Garden
Club of America members,
and hiker naturalists, with
offices in Washington. It
ceased in 1945.

• Flora of Virginia, my
most recent satisfaction of
leading VNPS, is that of
the successful challenge to
VNPS members to support
the publication of the Flora
of Virginia, by financial
and personal skills and
talents. The Flora gives
Virginians their own book
containing their botanical

will have to see VNPS grow
in strength, or there will not
be another 20 or 50 years.
The WFPS of the early
1900s lasted 45 years with a
small base but did not reach
the strength of VNPS or
have to face the issues of the
continued demise of native
plants and habitats. VNPS
can continue its prime
purpose of conserving
wildflowers and wild
places if individuals lead its
activities to attain that goal.

What advice can you give
to our members who aspire
to your level of knowledge
of native plants?

• Get a copy of the New-
comb Wildflower Guide
for the field;

• Keep a copy of the Flora
of Virginia to have in the
car and at home;

• Be alert in the field to see
not only plants you know,
but especially plants you do
not know; and

• Make it your driving
pleasure to find, identify,
and note threats to plants,
so you can protect them for
future generations.

Any final thoughts?

The effort of conserv-
ing, educating, and living
with native plants is never
complete, but it’s satisfying
to have accomplished any
part. Finally, I am grateful
for each of you who has
joined, served, and worked
on the organization and
programs to keep our
simple focus of conserving
our native plants and their
habitats. ❖

6

Sempervirem , Spring 2015

Sempervirens: A New Look for VNPS

Y ou are reading the first issue of
Sempervirens, the newsletter of
the Virginia Native Plant Society. You
will notice that the publication has
changed in purpose, in content, in
look, and, of course, in name. First,
the name. The Board felt it needed
a new one. In this
day and age, when
we tweet at the
drop of a hat, a
printed newsletter
cannot be a bulle-
tin, which, by
definition, offers timely or urgent
information. What’s more, the name
Bulletin carried no reference to the
mission of VNPS.

The new name addresses those
shortcomings. The word semper-
virens means evergreen — always
flourishing, forever vigorous, all of
which, it is hoped, describe VNPS.
Besides its botanical meaning,
though, evergreen has a journalistic
sense, referring to copy that has a
longer shelf life, and this reflects a
change in editorial focus. Of course,
important events will be put in the
spotlight, but look for more articles
you’ll want to reread in a year or
three, with more botanical and

Flora Meets Fall

T hank you to Virginia Native Plant
Society members and chapters
that donated to the Flora of Virginia
Project in response to our fall
mailing. We met our $25,000 goal,
and that is tremendously exciting. It
will come as no surprise that many
Flora supporters are VNPS members,
and we are so grateful! As you know,
all our funding comes from donations
and grants, so your help is especially
important and valued.

ecological information, visits with
people who have been instrumental
in the success of VNPS, and close-
ups of research and conservation in
Virginia. This was already under
way, and we hope it will be even more
noticeable in this and future issues.

Sempervirens will be published
four times a year instead of five. The
fifth issue was instituted as a sort of
legislative alert, but today it can’t do
that as effectively as other means.
Timely or urgent communications
are now the niche of our website, our
Facebook page, or the telephone.

The word sempervirens, of
course, is the specific epithet for a
number of our native plants, the star
probably Lonicera sempervirens, our
Wildflower of the Year for 2014,
a.k.a. Coral Honeysuckle. We
considered leaving the initial “s”
lowercase in the title as it appears in
a species name but felt a capital “S”
would give the title more weight,

Other than concerted efforts like
the VNPS’s annual drive in 2013
earmarking the Flora App, gifts to
the Flora Project are not restricted
to a particular program. Grants,
however, are specific, and we have
been working hard applying for
grants to cover various efforts, but
especially the App, our top priority.
Nevertheless, this fall’s gifts are
supporting work that is very much
related to the App, such as writing

and because it just looked better.

The design of Sempervirens is airier,
cleaner, newer.

There remains the matter of how
Sempervirens was named. You will
recall that there was a contest, the
prize an original illustration by Lara
Call Gastinger
from the Flora of
Virginia. A lot of
good nominations
came in. The
Executive Com-
mittee voted first,
narrowing the field. Then the full
Board voted on the candidates the
EC picked. You know how it turned
out. But who nominated it? Well,
um, that would be this reporter. And
this reporter is returning the prize to
the Board to give away. There will
be a raffle at the Annual Meeting.

The person whose ticket is drawn
will receive his or her choice of Lara’s
illustrations, if available. (Lara
illustrated virtually the entire Flora,
i.e., everything but the graminoids,
the ferns, and the fern allies.) Thank
you for your suggestions, and we
hope you will be as excited as we
are about Sempervirensl — Bland
Crowder, Publications Chair

those proposals and work toward the
volunteer project in which the Flora
descriptions will be added to the App
database (see the last issue for details).

A good number of VNPS mem-
bers have volunteered to work on this
part of the App project. I will be in
touch with you. We’re still working
on the particulars, and I will pipe up
if we need more people. Thanks!

— Bland Crowder, executive
director, Flora of Virginia Project

Sempervirens

Funding Goal. Thanks, Members!

Sempervirens ; Spring 2015

7

How a Property Is Chosen
To Be a Natural Area Preserve

L ast time, I discussed how the Vir-
ginia Natural Heritage Program
determined which native plants were
rare and to be included as natural
heritage resources. The logical next
step is to explain how a property is
selected to become part of your state
Natural Area Preserve System. In
25 short years, your Natural Area
Preserve System has grown from zero
natural area preserves in 1990 to 62
today, with the addition in January
of Dundas Granite Flatrock Natural
Area Preserve, in Brunswick County.
This system of protected lands
includes some of the finest forests,
savannas, beaches, dunes, barrens,
woodlands, natural ponds, swamps,
marshes, and caves in Virginia —
and on earth.

The goal of natural area protec-
tion is to secure habitats of natural
heritage resources. The Virginia
Natural Area Preserves Act author-
izes the Virginia Department of Con-
servation and Recreation to acquire
and manage a system of natural area
preserves. Natural area preserve
dedication provided in the act affords
the strongest form of protection in
Virginia for biodiversity. Natural area
protection requires understanding
the many factors that may affect the
quality of a site, threats to resource
values, landowner attitudes and
interests, which resource-protection
tools are most appropriate, and
what may be accomplished with the
resources available.

The public and private lands
that have been dedicated as natural
area preserves have been selected to
protect the rarest and most threat-
ened elements (plants, animals,
communities) of Virginia’s natural
heritage. A basic requirement is that
each natural area must support
viable populations or examples of
Virginia’s and the planet’s most
significant natural heritage
resources. Our work, as part of the
NatureServe network, provides that
important global data. Considera-
tions used to determine whether
a site qualifies for dedication as a
natural area preserve include:

1. Global and state rarity ranks of the
communities and species on the site.

2. Whether its size and condition can
meet protection and management
needs of the elements present.

3. The degree to which the elements

The Pinnacle Natural Area Preserve, in
Russell County, supports at least nine rare
species and two rare natural communities.
(Photograph © DCR-DNH, Gary P. Fleming)

present on the conservation site are
adequately protected on other lands.

4. The number and quality of rare-
species populations and exemplary
natural communities.

5. Physical or functional proximity of
the site to other protected lands.

6. Potential costs of managing the site
and of restoring conditions favorable
for natural heritage resources.

7. The level of threats from invasive
species, climate change, and other
factors.

Members of the Virginia Natural
Heritage Program staff have estab-
lished scoring criteria for these
attributes, and candidate sites are
evaluated, scored, and compared
in the office and the field to reach
a decision as to whether an area
qualifies. Then it depends on a
willing landowner and available
resources.

Thanks to the great work over
the years of many partners, includ-
ing the Virginia Native Plant
Society, your Natural Area Preserve
System’s 55,372 acres support 760
mapped locations (populations) of
458 different exemplary natural
communities and rare plant and
animal species. Of the $84 million
protection cost for the 62 properties,
$42 million has been in state funds,
primarily the 1992 and 2002 Natural
Area bonds, and the other $42 mil-
lion has been raised from various
sources mostly matched by the bond
funds. You might be wondering what
is next for additions to the system.
Sadly, there are only very limited state
funds for the purchase of new natural
area preserves, so progress will be
slow until a new bond is approved or
another source is identified.

Torn Smith is director of the Vir-
ginia Natural Heritage Program.
Learn more about our natural area
preserves at www.dcr.virginia
.gov/natural_heritage/index.shtml.

Sempervirem , Spring 2015

September 11-13

Mark Your Calendar for the Annual Meeting

T he Shenandoah
and Upper James
River chapters
will host the 2015
Annual Meeting of
the Virginia Native
Plant Society at
Staunton’s Frontier
Culture Museum,

September 11-13.

This will be a
wonderful time to
visit the Great Valley
of Virginia, where
the headwaters of
both the James River
and the Shenandoah
River arise. Public
lands, including national forests
and Shenandoah National Park,
form the Valley boundaries to
the east and west. The common-

1

m

Cowbane Prairie Natural Area Preserve, in Augusta County, will be the site
of one of the field trips at the Annual Meeting. It features wet and mesic
prairies and 11 rare plant species. (Photo by Nancy Sorrells)

wealth’s premier agricultural region
poses special challenges to native
species and restoration efforts.

Speakers and field trips will

highlight the unique
habitats and native
plants — common and
globally rare species —
found there. Trips
will include visits to

_ l ± ^ Deep Run Ponds, Sink

Hole Ponds, Augusta
Springs, and Reddish
Knob. Herbaria and
gardens are also on
the list of interesting
places you can visit.

So, come join these
two VNPS chapters in
the Valley of Virginia!
The next issue of
Sempervirens will
include a registration form and
details about the meeting. Watch
your mailbox in early July. ❖

jadud papAaaj uo paruijq

•A|§UjpjODDE M3U3J pUE |3qE| Suj|JEUU
jnoA uo a}Ep uopEjjdxa aip a}ou asEajd

VA ‘bjnquosujen
'ON lllflygd

aivd

86e}sod s n
uoiieziueBjo mcud-uON

^JO’SdUA'MMM

0Z9ZZ VA AoAog
z Jiun ‘3UE1 UUB,J Apuejg 00i7
tujnq imuomijadx^ Xpun|g
lUBJd UTUT§JIy\

A13IDOS INVld
3AI1VN VINIOdIA