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AE LIZARDS OF ECUADOR:

PeEpT CHECKLIST, DISTRIBUTION AND SYSTEMATIC REFERENCES

Omar Torres-Carvajal
Natural History Museum & Biodiversity Research Center
University of Kansas

SMITHSONIAN
HERPETOLOGICAL INFORMATION
SERVICE
NO. 131

2001

SMITHSONIAN HERPETOLOGICAL INFORMATION SERVICE

The first number of the SMITHSONIAN HERPETOLOGICAL INFORMATION SERVICE series
appeared in 1968. SHIS number 1 was a list of herpetological publications arising
from within or through the Smithsonian Institution and its collections entity, the
United States National Museum (USNM). The latter exists now as little more than
an occasional title for the registration activities of the National Museum of Natural
History. No. 1 was prepared and printed by J. A. Peters, then Curator-in-Charge
of the Division of Amphibians & Reptiles. The availability of a NASA translation
service and assorted indices encouraged him to continue the series and distribute
these items on an irregular schedule.

The series continues under that tradition. Specifically, the SHIS series prints and
distributes translations, bibliographies, indices, and similar items judged useful to
individuals interested in the biology of amphibians and reptiles, and unlikely to be
published in the normal technical journals. We wish to encourage individuals to
share their bibliographies, translations, etc. with other herpetologists through the
SHIS series. If you have such an item, please contact George Zug for its
consideration for distribution through the SHIS series. Contributors receive 25
free copies.

Single copies are available to interested individuals at $5 per issue. Libraries,
herpetological associations, and research laboratories are invited to exchange
their publications with us.

Please address all requests for copies and inquiries to George Zug, Division of
Amphibians and Reptiles, National Museum of Natural History, Smithsonian
Institution, Washington, D.C. 20560-0162, U.S.A. Please include a self-
addressed mailing label with requests.

INTRODUCTION

Several publications (Burt and Burt, 1933; Peters, 1967; Peters and Donoso-Barros,
1970; Miyata, 1982; Almendariz, 1992; Torres-Carvajal, 2000a; Uetz, 2001) provide lists of
Ecuadorian lizards. Of these, the most complete source of information is the checklist by
Peters (1967) because it provides both type specimen numbers (not found in other lists)
and bibliographic references. In addition, Peters (1967) and Peters and Donoso-Barros
(1970) are the only works that include taxonomic keys. On the other hand, Almendariz
(1992) and Torres-Carvajal (2000a) provided data on the general distribution of each
species within Ecuador; nevertheless, name of the species, author, and distribution is all
the information mentioned. None of these lists is a complete source of bibliographic,
taxonomic, and basic geographic information and most are now out of date.

The following checklist includes updated information on the taxonomy and
distribution of all known lizards (151 species) in Ecuador, as well as all systematic
references that I was able to find for each taxon. Any absent reference is an oversight, and
exclusion does not mean that the work is irrelevant.

Families and their included genera and species are arranged alphabetically. The
number of species known from Ecuador is given in parentheses after each familial name.
The following data are given for each species: current name; author; year of publication;
type specimen number(s); type locality (additional geographic information is indicated in
brackets where necessary); and distribution. For those species that were described in a
genus other than the current one, the original name is included. Systematic references and
general taxonomic comments are indicated where applicable. The distributions within
continental Ecuador used the geographical units of Coloma and Quiguango-Ubillus (2001)
that are presented in Table 1.

Table 1. Geographical units of continental Ecuador.

0-1000 m 1000-2000 m 2000-3000 m > 3000 m
West of the Andes western tropical western subtropical western temperate High
East of the Andes eastern tropical eastern subtropical eastern temperate panndean

Throughout the checklist, the catalogue numbers of type specimen(s) are preceded by
the institutional codes: AMNH (American Museum of Natural History, New York), ANSP
(Academy of Natural Sciences, Philadelphia), BMNH (British Museum of Natural History,
London), CAS (California Academy of Sciences, San Francisco), CAS-SU (Stanford
University Collection in the CAS), FMNH (Field Museum of Natural History, Chicago),
ICN (Instituto de Ciencias Naturales, Bogota), KU (University of Kansas, Natural History
Museum, Lawrence), MCZ (Harvard University, Museum of Comparative Zoology,
Cambridge), MECN (Museo Ecuatoriano de Ciencias Naturales, Quito), MHNG (Museum

d'Histoire Naturelle, Genéve), MHNP (Museum National d'Histoire Naturelle, Paris),
MRHN (Institut Royal des Sciences Naturelles de Belgique, Brussels), MZUT (Museo
Regionale di Scienze Naturali, Torino), NRM (Naturhistoriska Rijkmuseet, Stockholm),
NMW (Naturhistorisches Museum Wien), ZMB (Zoologisches Museum Berlin), OUMNH
(Oxford University Museum of Natural History), QCAZ (Museo de Zoologia, Pontificia
Universidad Catélica del Ecuador, Quito), UIMNH (Museum of Natural History,
University of Illinois at Urbana-Champaign), UMMZ (University of Michigan, Museum of
Zoology, Ann Arbor), USNM (National Museum of Natural History, Washington), UUZM
(Uppsala University Zoological Museum), ZFMK (Zoologisches Forschungsinstitut und
Museum Alexander K6nig, Bonn), ZMH (Zoologisches Museum Hamburg), ZSM
(Zoologische Staatsammlung, Munich).

ACKNOWLEDGEMENTS

I am grateful to Aaron Bauer, Wolfgang Bohme, Luis A. Coloma, Sammy de Grave,
Kevin de Queiroz, William E. Duellman, Richard Etheridge, Henry Fitch, Linda S. Ford,
Ned Gilmore, Jakob Hallermann, Marinus S. Hoogmoed, Sven Kullander, Colin McCarthy,
Linda Trueb and Luis Vinueza for their comments and help.

CHECKLIST OF LIZARDS OF ECUADOR

ANGUIDAE (1)
Diploglossus monotropis (Kuhl, 1820)
Original name: Scincus monotropis mihi
Type(s): Not located.
Type locality: Not reported.
Distribution: Western tropical.

GEKKONIDAE (21)
Gonatodes caudiscutatus (Giinther, 1859b)
Original name: Gymnodactylus caudiscutatus
Type(s): Syntypes (4): BMNH 1946.9.7.6-9
Type locality: Andes of western Ecuador.
Systematics: Kluge (1995).
Distribution: Western tropical, Galapagos.

Gonatodes concinnatus (O’ Shaughnessy, 1881)
Original name: Goniodactylus concinnatus

Type(s): Syntypes (3): BMNH 1946.9.7.10-12

Type locality: Canelos, [Provincia Pastaza], Ecuador.
Systematics: Kluge (1995).

Distribution: Eastern tropical.

Gonatodes humeralis (Guichenot, 1855)

Original name: Gymnodactylus humeralis

Type(s): Syntypes (6): MHNP 6745

Type locality: RioUcayali, Mission de Sarayacu [Provincia Pastaza, Ecuador], Peru.
Systematics: Avila-Pires (1995), Kluge (1995).

Distribution: Eastern tropical.

Hemidactylus mabouia (Moreau de Jonnés, 1818)

Original name: Gecko mabouia

Type(s): Lectotype: MHNP 6573 (Duméril and Duméril, 1851)

Type locality: Antilles; restricted to St. Vincent Island, Lesser Antilles, by Stejneger
(1904).

Systematics: Avila-Pires (1995), Kluge (1969).

Distribution: Eastern tropical.

Lepidoblepharis buchwaldi Werner, 1910

Type(s): Syntypes (2): ZMH 808

Type locality: Hacienda Clementina, Babahoyo [5 m, Provincia Los Rios], Ecuador.
Systematics: Kluge (1995), Vanzolini (1978).

Distribution: Western tropical.

Lepidoblepharis festae Peracca, 1897

Type(s): Syntypes (2): MZUT 2163

Type locality: San José de Cuchipamba [78°45' W, 03°01'S, 3000 m, Provincia Azuay],
Ecuador orientale.

Systematics: Kluge (1995), Vanzolini (1978).

Comments: Type species.

Distribution: Eastern tropical, eastern temperate.

Lepidoblepharis intermedius Boulenger, 1914

Type(s): Syntypes (2): BMNH 1946.8.26.62

Type locality: Pena Lisa, [Municipio] Condoto [Departamento Chocé], Colombia.
Systematics: Kluge (1995), Vanzolini (1978).

Distribution: Western tropical.

Lepidoblepharis grandis Miyata, 1985a

Type(s): Holotype: KU 142675

Type locality: 3.7 km east of village of Dos Rios [78°53' W, 00°12' S], 1190 m, Provincia
Pichincha, Ecuador.

Systematics: Kluge (1995).

Distribution: Western tropical, western subtropical.

Lepidoblepharis ruthveni Parker, 1926

Type(s): Holotype: BMNH 1946.8.27.3

Type locality: Chimbo [79°05' W, 02°09' S, 345 m, Provincia Chimborazo], Ecuador.
Systematics: Kluge (1995).

Distribution: Western tropical.

Lepidodactylus lugubris (Duméril and Bibron, 1836)

Original name: Platydactylus lugubris

Type(s): Lectotype: MNHN 5323 (Wells and Wellington, 1985)
Type locality: I'ile d' Otaiti [Tahiti].

Distribution: Western tropical, Galapagos.

Phyllodactylus barringtonensis Van Denburgh, 1912

Type(s): Holotype: CAS 12057

Type locality: Barrington Island, Galapagos Archipelago [Provincia Galapagos,
Ecuador].

Distribution: Galapagos.

Phyllodactylus baurti Garman, 1892
Type(s): Holotype: MCZ 9433
- Type locality: Las Cuevas, Charles Island [Provincia Galapagos, Ecuador].
Systematics: Van Denburgh (1912).
Distribution: Galapagos.

Phyllodactylus darwini Taylor, 1942

Type(s): Holotype: CAS 10848

Type locality: Chatham Island, Galapagos Islands [Provincia Galapagos, Ecuador].
Systematics: Van Denburgh (1912).

Distribution: Galapagos.

Phyllodactylus galapagensis Peters, 1869

Type(s): Holotype: ZMB 6550

Type locality: Galapagos-Inseln [Provincia Galapagos, Ecuador].
Systematics: Van Denburgh (1912).

Distribution: Galapagos.

Phyllodactylus gilberti Heller, 1903

Type(s): Holotype: CAS-SU 4549

Type locality: Wenman Island, Galapagos Archipelago [Provincia Galapagos,
Ecuador].

Systematics: Van Denburgh (1912).

Distribution: Galapagos.

Phyllodactylus leet Cope, 1889

Type(s): USNM 14957

Type locality: Chatham Island, Galapagos Islands [Provincia Galapagos, Ecuador].
Systematics: Van Denburgh (1912).

Distribution: Galapagos.

Phyllodactylus pumilus Dixon and Huey, 1970

Type(s): Holotype: USNM 167227

Type locality: 1 km south of Manta, Department [Provincia] Manabi, sea level,
Ecuador.

Distribution: Western tropical.

Phyllodactylus reissti Peters, 1862a

Type(s): Lectotype: ZMB 4567 (Dixon and Huey, 1970)
Type locality: Guayaquil [Provincia Guayas], Ecuador.
Systematics: Dixon and Huey (1970), Taylor (1942).
Distribution: Western tropical, western subtropical.

Pseudogonatodes guianensis Parker, 1935

Type(s): Holotype: BMNH 1946.8.27.8

Type locality: Upper Cuyini river, British Guyana.
Systematics: Avila-Pires (1995), Kluge (1995).
Distribution: Eastern tropical.

Sphaerodactylus scapularis Boulenger, 1902

Type(s): Holotype: BMNH 1946.8.30.70

Type locality: St. [San] Javier, [78°47' W, 01°04' N] 60ft [Provincia Esmeraldas],
Ecuador.

Systematics: Kluge (1995).

Distribution: Western tropical.

Thecadactylus rapicauda (Houttuyn, 1782)

Original name: Gekko rapicauda

Type(s): Not reported; restricted to Paramaribo, Suriname, by Hoogmoed (1973).
Type locality: West Indies.

Systematics: Avila-Pires (1995).

Distribution: Western tropical.

GYMNOPHTHALMIDAE (47)
Alopoglossus angulatus (Linnaeus, 1758)
Original name: Lacerta angulata
Type(s): Neotype RMNH 15200 (Hoogmoed, 1973)
Type locality: Brown's Mountain, Suriname.
Systematics: Avila-Pires (1995).
Distribution: Eastern tropical.

Alopoglossus atriventris Duellman, 1973

Type(s): Holotype: KU 126783

Type locality: Lago Agrio, 330 m, Provincia Napo [Sucumbios], Ecuador.
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical.

Alopoglossus buckleyi (O’ Shaughnessy, 1881)
Original name: Leposoma buckleyi

Type(s): Holotype: BMNH 1946.8.31.66

Type locality: Canelos [Provincia Pastaza], Ecuador.
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical, eastern subtropical.

Alopoglossus copii Boulenger, 1885

Type(s): Syntypes (4): BMNH 1946.8.31.58-59, 1946.9.8.17-18

Type locality: Pallatanga [Provincia Chimborazo] and Canelos [Provincia Pastaza],
Ecuador; herein restricted to Canelos, Provincia Pastaza, Ecuador.

Distribution: Eastern tropical.

Alopoglossus festae Peracca, 1904

Type(s): Holotype: MZUT 2875

Type locality: Vinces [79°45' W, 01°32'S, 15 m, Provincia Los Rios], Ecuador.
Distribution: Western tropical.

Anadia petersi Oftedal, 1974

Type(s): Holotype: USNM 193221

Type locality: San Ramon, near Loja, Loja Province, Ecuador.
Distribution: Western temperate, eastern temperate.

Anadia rhombifer (Giinther, 1859b)

Original name: Cercosaurus rhombifer

Type(s): Holotype: BMNH 1946.9.1.4

Type locality: Andes of western Ecuador.

Systematics: Oftedal (1974).

Distribution: Western tropical, western subtropical, eastern subtropical.

Arthrosaura reticulata (O’ Shaughnessy, 1881)

Original name: Cercosaura reticulata

Type(s): Holotype: BMNH 1946.9.1.5

Type locality: Canelos, Ecuador.

Systematics: Avila-Pires (1995), Hoogmoed and Avila-Pires (1992).
Distribution: Eastern tropical.

Bachia trisanale (Cope, 1868)

Original name: Ophiognomon trisanale

Type(s): Holotype: ANSP 9637

Type locality: Napo [Ecuador] or upper Marafion [Peru].
Systematics: Dixon (1973).

Distribution: Eastern tropical.

Echinosaura horrida Boulenger, 1890

Type(s): Syntypes (2): BMNH 1946.8.31.60-61
Type locality: Ecuador.

Distribution: Western tropical.

Euspondylus guentheri (O’ Shaughnessy, 1881)
Original name: Ecpleopus guentheri

Type(s): Holotype: BMNH 1946.8.8.99

Type locality: Sarayacu [Provincia Pastaza], Ecuador.
Systematics: Peters (1959), Uzzell (1973).
Distribution: Eastern tropical.

Euspondylus maculatus Tschudi, 1845

Type(s): Syntypes (2): Not located.

Type locality: Not reported; restricted to Republica Peruana by Tschudi (1846).
Systematics: Peters (1959), Uzzell (1973).

Comments: Type species.

Distribution: Eastern tropical, eastern subtropical.

Iphisa elegans Gray, 1851

Type(s): Holotype: BMNH 1946.9.1.1

Type locality: Belém, Para [Brazil].
Systematics: Avila-Pires (1995), Dixon (1974).
Comments: Type species.

Distribution: Eastern tropical.

Leposoma parietale (Cope, 1885)
Original name: Mionyx parietalis
Type(s): Not located.

Type locality: Pebas, Peru.
Distribution: Eastern tropical.

Neusticurus cochranae Burt and Burt, 1931
Original name: Neusticurus ecpleopus cochranae
Type(s): Holotype: AMNH 28891

Type locality: San José de Sumaco, Ecuador.
Systematics: Uzzell (1966).

Distribution: Eastern tropical, eastern subtropical.

Neusticurus ecpleopus Cope, 1876

Type(s): Not located.

Type locality: Peru.

Systematics: Avila-Pires (1995), Uzzell (1966).
Distribution: Eastern tropical, eastern subtropical.

Neusticurus strangulatus (Cope, 1868)

Original name: Euspondylus strangulatus

Type(s): Holotype: ANSP 7538

Type locality: Not reported.

Systematics: Uzzell (1966).

Distribution: Eastern tropical, eastern subtropical.

Pholidobolus affinis (Peters, 1862b)

Original name: Ecpleopus affinis

Type(s): Holotype: ZSM 664/0

Type locality: Not reported.

Systematics: Hillis (1985), Montanucci (1973).

Distribution: Western temperate, Andean, eastern subtropical, eastern temperate.

Pholidobolus annectens (Parker, 1930)

Original name: Macropholidus annectens

Type(s): Holotype: BMNH 1946.8.31.34

Type locality: Vicinity of Loja city, 2200 m [Provincia Loja], Ecuador.
Systematics: Hillis (1985), Montanucci (1973).

Distribution: Western temperate, eastern temperate.

Pholidobolus macbrydei Montanucci, 1973

Type(s): Holotype: KU 140993

Type locality: 9.1 km north of and 2.3 km west of (by road) Biblian, near Hacienda
Oeste, 3070 m, Cafiar Province, Ecuador.

Systematics: Hillis (1985).

Distribution: Western temperate, Andean, eastern temperate.

10

Pholidobolus montium (Peters, 1862b)

Original name: Ecpleopus montium

Type(s): Lectotype: RMNH 3401 (Montanucci, 1973)

Type locality: Hochgebirge von Ecuador; restricted to Quito by Montanucci (1973).
Systematics: Hillis (1985), Montanucci (1973).

Comments: Type species.

Distribution: Western temperate, Andean, eastern temperate.

Pholidobolus prefrontalis Montanucci, 1973

Type(s): Holotype: KU 141093

Type locality: 4.9 km south of Tixan (by road), Chimborazo Province, Ecuador.
Systematics: Hillis (1985).

Distribution: Western temperate.

Prionodactylus argulus (Peters, 1862b)

Original name: Cercosaura argulus

Type(s): Holotype: ZMB 4555

Type locality: Mountainous regions around Sta. Fé de Bogota [Colombia].
Systematics: Avila-Pires (1995), Uzzell (1973).

Distribution: Eastern tropical, eastern subtropical.

Prionodactylus dicrus Uzzell, 1973

Type(s): Holotype: FMNH 36708

Type locality: Mapoto, 1300 m, [Provincia] Tungurahua, Ecuador.
Distribution: Eastern tropical, eastern subtropical.

Prionodactylus manicatus (O’ Shaughnessy, 1881)

Original name: Cercosaura manicata

Type(s): Syntypes (4): BMNH 1946.8.2.1, 1946.8.31.15-17

Type locality: Pallatanga [Provincia Chimborazo] and Canelos [Provincia Pastaza],
Ecuador.

Systematics: Uzzell (1973).

Distribution: Eastern tropical, eastern subtropical.

Prionodactylus vertebralis (O’ Shaughnessy, 1879)
Original name: Cercosaura vertebralis

Type(s): Holotype: BMNH 1946.8.31.35

Type locality: Intac, Ecuador.

Systematics: Uzzell (1973).

Distribution: Western tropical, western subtropical.

le

Prionodactylus oshaughnessyi Boulenger, 1885

Type(s): Syntypes (4): BMNH 1946.8.31.18-20, 80.12.8.17

Type locality: Pallatanga [Provincia Chimborazo] and Canelos [Provincia Pastazal],
Ecuador.

Systematics: Avila-Pires (1995), Uzzell (1973).

Distribution: Eastern tropical, eastern subtropical.

Proctoporus anatoloros Kizirian, 1996

Type(s): Holotype: USNM 229691

Type locality: La Bonita [77°33' W, 00°27' S, ca.1500 mJ], Napo [Provincia Sucumbios],
Ecuador.

Distribution: Eastern subtropical.

Proctoporus balneator Kizirian, 1996

Type(s): Holotype: MHNG 2360.44

Type locality: Southern versant of [Volcan] Tungurahua [78°25' W, 01°32' S], 2750 m,
[Provincia Tungurahua], Ecuador.

Distribution: Eastern temperate.

Proctoporus cashcaensis Kizirian and Coloma, 1991

Type(s): Holotype: KU 135015

Type locality: Guaranda [79°00' W, 01°60' S], 2640 m, Provincia Bolivar, Ecuador.
Systematics: Kizirian (1996).

Distribution: Western temperate, Andean.

Proctoporus colomaromani Kizirian, 1996

Type(s): Holotype: QCAZ 2819

Type locality: 19.8 km west of Chillogallo, on road to Quito, 2810 m, Provincia
Pichincha, Ecuador.

Distribution: Western temperate.

Proctoporus hypostictus Boulenger, 1902

Type(s): Holotype: BMNH 1946.8.31.31

Type locality: Paramba, 3500 ft, [Provincia Imbabura], Ecuador.
Systematics: Kizirian (1996).

Distribution: Western tropical, western subtropical, western temperate.

Proctoporus labionis Kizirian, 1996

Type(s): Holotype: UMMZ 188631

Type locality: San Francisco de Las Pampas [78°58' W, 00°25' S, 1800 m], Provincia
Cotopaxi, Ecuador.

Distribution: Western subtropical.

i

Proctoporus meleagris Boulenger, 1885
Type(s): Holotype: BMNH 1946.8.31.33
Type locality: Western Ecuador.
Systematics: Kizirian (1996).
Distribution: Eastern subtropical.

Proctoporus oculatus (O’ Shaughnessy, 1879)

Original name: Ecpleopus oculatus

Type(s): Holotype: BMNH 1946.8.31.21

Type locality: Intac [=Intag, 78°36' W, 00°24’ N, 1200 m, Provincia Imbabura], Ecuador.
Systematics: Kizirian (1996).

Distribution: Western subtropical.

Proctoporus orcesi Kizirian, 1995

Type(s): Holotype: KU 146709

Type locality: 3.3 km (by road) ESE Cuyujua [78°00' W, 00°25' S], 2350 m, Provincia
Napo, Ecuador.

Systematics: Kizirian (1996).

Distribution: Eastern subtropical, eastern temperate.

Proctoporus petrorum Kizirian, 1996

Type(s): Holotype: USNM 196267

Type locality: Trail between Sevilla de Oro [78°38' W, 02°47’ S] and Méndez [78°18' W,
02°42' S], on the eastern slopes of the mountains between Cerro Negro [78°32' W,
02°45' S] and Pailas [78°31' W, 02°45'S] (tambos), Provincia Morona-Santiago,
Ecuador.

Distribution: Eastern temperate.

Proctoporus raneyi Kizirian, 1996

Type(s): Holotype: KU 142905

Type locality: 3.3 km (by road) ESE Cuyujua [78°00' W, 00°25" S], 2350 m, Provincia
Napo, Ecuador.

Distribution: Eastern temperate.

Proctoporus simoterus (O’ Shaughnessy, 1879)

Original name: Emphrassotis simoterus

Type(s): Holotype: BMNH 1946.9.1.6

Type locality: Intac [=Intag, 78°36' W, 00°24' N, 1200 m, Provincia Imbabura], Ecuador.
Systematics: Kizirian (1996).

Distribution: Western temperate, Andean.

13

Proctoporus stigmatoral Kizirian, 1996

Type(s): Holotype: USNM 229646

Type locality: Pailas [78°31' W, 02°45' S], a tambo on the trail between Sevilla de Oro
[78°38' W, 02°47' S] and Méndez [78°18' W, 02°42' S], on the eastern or northeastern
facing slope [Rio Negro Valley on northern versant of Cordillera Zapote Naida],
Provincia Morona-Santiago, Ecuador.

Distribution: Eastern temperate.

Proctoporus unicolor (Gray, 1858)

Original name: Riama unicolor

Type(s): Holotype: 1946.8.31.32

Type locality: Guayaquil [in error], Ecuador; herein restricted to Otavalo, 78°15' W,
00°13' N, 2554 m, Provincia Imbabura, Ecuador.

Systematics: Kizirian (1996).

Distribution: Western temperate, Andean.

Proctoporus vespertinus Kizirian, 1996

Type(s): Holotype: AMNH 22130

Type locality: [Pampa] Chitoqué, 6000 ft [Provincia Loja], Ecuador.
Distribution: Western subtropical.

Proctoporus vietus Kizirian, 1996

Type(s): Holotype: MHNG 2437.48

Type locality: [San Francisco de] Las Pampas [78°58' W, 00°25'S, 1800 m], Provincia
Cotopaxi, Ecuador.

Distribution: Western subtropical.

Ptychoglossus brevifrontalis Boulenger, 1912

Type(s): Holotype: BMNH 1946.8.31.63

Type locality: El Topo, Rio Pastaza [Provincia Pastaza], Ecuador.
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical, eastern subtropical.

Ptychoglossus bilineatus Boulenger, 1890
Type(s): Holotype: BMNH 1946.8.2.38
Type locality: Ecuador.

Systematics: Harris (1994).

Comments: Type species.

Distribution: Western tropical.

14

Ptychoglossus gorgonae Harris, 1994

Type(s): Holotype: ICN 5644

Type locality: Isla Gorgona, 78°09' W, 03°00' N, 10 m, Department of Cauca, Colombia.
Distribution: Western tropical.

Teuchocercus keyi Fritts and Smith, 1969

Type(s): Holotype: UIMNH 80451

Type locality: 4 km east of Rio Baba bridge, 24 km south of Santo Domingo de los
Colorados, 600 m, [Provincia] Pichincha, Ecuador.

Comments: Type species.

Distribution: Western tropical.

IGUANIDAE: CORYTOPHANINAE (1)
Basiliscus galeritus Duméril and Duméril, 1851
Type(s): Syntypes (2): MHNP 2130-31
Type locality: N.-Grenade.
Distribution: Western tropical, western subtropical.
Systematics: Lang (1989).

IGUANIDAE: HOPLOCERCINAE (7)
Enyalioides cofanorum Duellman, 1973
Type(s): Holotype: KU 146658
Type locality: Santa Cecilia, 340 m, Provincia Napo, Ecuador.
Distribution: Eastern tropical.

Enyalioides heterolepis (Bocoutrt, 1874)

Original name: Enyalius heterolepis

Type(s): Lectotype: MHNP 4067 (Guibé, 1954)
Type locality: Amerique tropicale.
Comments: Type species.

Distribution: Western tropical.

Enyalioides laticeps (Guichenot, 1855)

Original name: Enyalius laticeps

Type(s):Holotype: MHNP 6821

Type locality: Fonte Boa, upper Amazon, Brazil.

Systematics: Avila-Pires (1995).

Comments: I follow Avila-Pires (1995) in not recognizing proposed subspecies.
Distribution: Eastern tropical.

15

Enyalioides microlepis (O’ Shaughnessy, 1881)

Original name: Enyalius microlepis

Type(s): Syntypes (2): BMNH 1946.8.9.5-6

Type locality: Sarayacu [78°09' W, 03°00' N, Provincia Pastaza], Ecuador.
Distribution: Western tropical.

Enyalioides oshaughnessyi (Boulenger, 1881)
Type(s): Holotype: MRHN 2009

Type locality: Ecuador.

Distribution: Eastern tropical, eastern subtropical.

Enyalioides praestabilis (O’ Shaughnessy, 1881)

Original name: Enyalius praestabilts

Type(s): Syntypes (2): BMNH 1946.8.9.15, 80.12.8.37

Type locality: Pallatanga [Provincia Chimborazo] and Canelos [Provincia Pastaza],
Ecuador.

Distribution: Eastern tropical, eastern subtropical.

Morunasaurus annularis (O’ Shaughnessy, 1881)
Original name: Hoplocercus annularis

Type(s): Holotype: BMNH 1946.8.10.35

Type locality: Canelos [Provincia Pastaza], Ecuador.
Systematics: Kohler et al. (1999).

Distribution: Eastern tropical.

IGUANIDAE: IGUANINAE (4)
Amblyrhynchus cristatus Bell, 1825
Type(s): Holotype: OUMNH 6176
Type locality: Mexico [in error]; restricted to Narborough [Isla Fernandina] by
Eibl-Eibesfeldt (1956).
Systematics: de Queiroz (1987).
Distribution: Galapagos.

Conolophus pallidus Heller, 1903

Type(s): Holotype: CAS-SU 4749

Type locality: Barrington Island, Galapagos.
Systematics: de Queiroz (1987).
Distribution: Galapagos.

16

Conolophus subcristatus (Gray, 1831)
Original name: Amblyrhynchus subcristatus
Type(s): Not located.

Type locality: Galapagos.

Systematics: de Queiroz (1987).
Distribution: Galapagos.

Iguana iguana (Linnaeus, 1758)

Original name: Lacerta iguana

Type(s): Syntypes (?): NRM 114, other syntypes not located.

Type locality: Indies; restricted to island of Terre de Haut, Les Iles de Saintes,
Departement de La Guadeloupe, French West Indies, by Lazell (1973).
Systematics: de Queiroz (1987).

Distribution: Western tropical.

IGUANIDAE: POLYCHROTINAE (33)
Anolis aequatorialis Werner, 1894
Type(s): Holotype: NMW 16233
Type locality: Ecuador.
Distribution: Western subtropical.

Anolis binotatus Peters, 1863

Type(s): Holotype: ZMB 4685

Type locality: Guayaquil [Provincia Guayas, Ecuador].
Distribution: Western tropical.

Anolis biporcatus Williams, 1966

Type(s): Holotype: MCZ 78935

Type locality: Northern Gorgona Island, 5-45 m, [Departamento] Cauca, Colombia.
Distribution: Western tropical.

Anolis bitectus Cope, 1864

Type(s): Syntypes (2): BMNH 1946.8.5.83, 1946.8.20.11
Type locality: West Ecuador.

Distribution: Western tropical.

Anolis bombiceps Cope, 1876
Type(s): Not located.

Type locality: Nauta, Peru.
Distribution: Eastern tropical.

17

Anolis chloris Boulenger, 1898

Type(s): Holotype: BMNH 1946.8.13.20

Type locality: Paramba [Parambas, 78°21' W, 00°48' N, 780 m, Provincia
Imbabura, Ecuador].

Distribution: Western tropical.

Anolis chocorum Williams and Duellman, 1967

Type(s): Holotype: KU 96934

Type locality: Rio Tuira at Quebrada La Plata, 100 m, Darién, Panama.
~ Distribution: Western tropical, western subtropical.

Anolis fasciatus Boulenger, 1885

Type(s): Syntypes (2): BMNH 1946.8.20.13, other syntype not located.
Type locality: Guayaquil [Provincia Guayas], Ecuador.

Distribution: Western tropical.

Anolis festae Peracca, 1904

Type(s): Syntypes (3): MZUT 2872

Type locality: Balzar [Provincia Guayas], Ecuador.

Distribution: Western tropical.

Anolis fitchi Williams and Duellman, 1984

Type(s): Holotype: KU 142865

Type locality: 16.5 km (by road) north-northeast of Santa Rosa, 1700 m,
Provincia Napo, Ecuador.

Distribution: Eastern tropical, eastern subtropical.

Anolis fraseri Giinther, 1859b

Type(s): Lectotype: BMNH 1946.8.8.47 (Williams, 1966)
Type locality: Andes of Western Ecuador.

Systematics: Williams (1966).

Distribution: Western tropical, western subtropical.

Anolis fuscoauratus D’ Orbigny, 1837

Original name: Anolis fusco-auratus

Type(s): Holotype: MHNP 2420

Type locality: Chile; restricted to RioMarmore, between Loreto and the
confluence of RioSara, Moxos Province, Bolivia, by D’ Orbigny (1847).
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical, eastern subtropical.

18

Anolis gemmosus O’ Shaughnessy, 1875

Type(s): Holotype: BMNH 1946.8.13.32

Type locality: Not reported.

Distribution: Western subtropical, western temperate.

Anolis gracilipes Boulenger, 1898

Type(s): Syntypes (5): BMNH 1946.8.13.9-12, NMW 12816
Type locality: Paramba [Parambas, 78°21' W, 00°48' N, 780 m,
Provincia Imbabura, Ecuador].

Distribution: Western tropical.

Anolis granuliceps Boulenger, 1898

Type(s): Syntypes (13): BMNH 1946.8.13.25-30, MZUT 2357,

NMW 12813 (2), 12814 (3), UMMZ 59002

Type locality: Paramba [Parambas, 78°21' W, 00°48' N, 780 m, Provincia
Imbabura, Ecuador].

Distribution: Western tropical.

Anolis lynchi Miyata, 1985b

Type(s): Holotype: MCZ 124406

Type locality: Santo Domingo de los Colorados, 600 m, Pichincha Province, Ecuador.
Distribution: Western tropical.

Anolis maculiventris Boulenger, 1898

Type(s): Syntypes (2): BMNH 1946.8.13.33-34

Type locality: Paramba [Parambas, 78°21' W, 00°48’ N, 780 m, Provincia
Imbabura, Ecuador].

Distribution: Western tropical.

Anolis nigrolineatus Williams, 1965

Type(s): Holotype: MCZ 38940

Type locality: Machala, El Oro Province, Ecuador.
Distribution: Western tropical.

Anolis nitens Cope, 1864

Type(s): Holotype: BMNH 1946.8.8.55
Type locality: Tropical America.
Systematics: Avila-Pires (1995).
Distribution: Eastern tropical.

19

Anolis ortonti Cope, 1868

Type(s): Holotype: ANSP 11404

Type locality: Napo [Ecuador] or upper Marafion [Peru].
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical.

Anolis parilis Williams, 1975

Type(s): Holotype: UIMNH 82901

Type locality: Rio Baba, 2.4 km south of Sto. Domingo de los Colorados,
[Provincia] Pichincha, Ecuador.

Distribution: Western tropical.

Anolis peraccae Boulenger, 1898

Type(s): Syntypes (6): BMNH 1946.8.13.13-17, MZUT 2358
Type locality: Chimbo and Rio Peripa, Ecuador.
Distribution: Western tropical, western subtropical.

Anolis princeps Boulenger, 1898

Type(s): Syntypes (4): BMNH 1946.8.13.4, 1946.8.12.99, 1946.8.8.45—46
Type locality: Carondelet, Rio Lita, Paramba.

Distribution: Western tropical.

Anolis proboscis Peters and Orcés-V., 1956

Type(s): Holotype: MCZ 54300

Type locality: Cunuco, 5 km NW Mindo, 1200 m, Pichincha Province, Ecuador.
Systematics: | Williams (1965, 1979).

Distribution: Western subtropical.

Anolis punctatus Daudin, 1802b
Type(s): Holotype: MHNP 2340
Type locality: South America.
Systematics: Avila-Pires (1995).
Distribution: Eastern tropical.

Anolis trachyderma Cope, 1876
Type(s): Holotype: ANSP 11363
Type locality: Nauta, Peru.
Systematics: Avila-Pires (1995).
Distribution: Eastern tropical.

20

Anolis transversalis Duméril and Duméril, 1851

Type(s): Holotype: MHNP 2499

Type locality: Amérique méridionale; restricted to Sarayacu [Provincia Pastaza],
Ecuador, by Guichenot (1855).

Systematics: Williams (1966).

Distribution: Eastern tropical.

Phenacosaurus orcesi Lazell, 1969

Type(s): Holotype: MCZ 38937

~ Type locality: Mt. Sumaco, Napo Pastaza [Napo] Province, Ecuador.
Systematics: Williams and Mittermeier (1991).

Distribution: Eastern subtropical.

Phenacosaurus vanzolinii Williams et al., 1996

Type(s): Holotype: MECN 0309

Type locality: south of La Alegria, 14 km (by road) from La Bonita
(77°38' W, 00°27' N), Sucumbios Province, Ecuador.

Systematics: Lazell (1969).

Distribution: Eastern subtropical, eastern temperate.

Polychrus marmoratus (Linnaeus, 1758)

Original name: Lacerta marmorata

Type(s): Syntypes (2): UUZM 14, 31

Type locality: Hispania [in error]; restricted to the vicinity of Paramaribo,
Suriname, by Hoogmoed (1973).

Systematics: Avila-Pires (1995).

Distribution: Eastern tropical.

Polychrus femoralis Werner, 1910

Type(s): Holotype: Lost (Hallermann, pers. comm.).
Type locality: Guayaquil [Provincia Guayas], Ecuador.
Distribution: Western tropical.

Polychrus gutturosus Berthold, 1846

Type(s): ZFMK 21341

Type locality: Popayan [76°37' W, 02°27' N, 1738 m, Departamento del
Cauca], Colombia.

Distribution: Western subtropical.

Zi

Polychrus spurrellii Boulenger, 1914

Type(s): Syntypes (2): BMNH 1946.8.8.33-34
Type locality: Pefia Lisa, Condoto, Colombia.
Distribution: Western tropical.

IGUANIDAE: TROPIDURINAE (26)
Microlophus albemarlensis (Baur, 1890)
Original name: Tropidurus albemarlensis
Type(s): Syntypes (11): USNM 15003-13
Type locality: Tagus Cove, Albemarle Island [Isla Isabela], Galapagos Archipelago.
Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus bivittata (Peters, 1871)

Original name: Tropidurus (Craniopeltis) bivittata

Type(s): Not located.

Type locality: Chatman Island [Isla San Cristobal], Galapagos Archipelago.
Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus delanonis (Baur, 1890)

Original name: Tropidurus delanonis

Type(s): Syntypes (17): USNM 1501418, 15020-25, 14926-30; MCZ 22664
Type locality: Hood Island [Isla Espanola], Galapagos Archipelago.
Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus duncanensis (Baur, 1890)

Original name: Tropidurus duncanensis

Type(s): Syntypes (4): USNM 1494144

Type locality: Duncan Island [Isla Pinzon], Galapagos Archipelago.
Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus grayjii (Bell, 1843)

Original name: Leiocephalus grayii

Type(s): Syntypes (2): BMNH 1946.8.29.73-74

Type locality: Chatman [Isla San Cristobal] and Charles [Isla Floreana] Islands,
Galapagos Archipelago.

Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

22

Microlophus habelii (Steindachner, 1876)

Original name: Tropidurus(Craniopeltis) pacificus (var. habelii)

Type(s): Holotype: NMW 13891

Type locality: Indefatigable [Isla Santa Cruz] or Bindloe [Isla Marchena] Islands,
Galapagos Archipelago.

Systematics: Frost (1992),Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus occipitalis (Peters, 1871)

Original name: Tropidurus (Laemopristus) occipitalis
Type(s): Holotype: ZMB 26397

Type locality: Peru.

Systematics: Dixon and Wright (1975), Frost (1992).
Distribution: Western tropical, western subtropical.

Microlophus pacificus (Steindachner, 1876)

Original name: Tropidurus (Craniopeltis) pacificus

Type(s): Not located.

Type locality: Indefatigable [Isla Santa Cruz] or Bindloe [Isla Marchena] Islands,
Galapagos Archipelago.

Systematics: Frost (1992), Van Denburgh and Slevin (1913), Wright (1983).
Distribution: Galapagos.

Microlophus peruvianus (Lesson, 1831)

Original name: Stellio peruvianus

Type(s): Holotype: MHNP 6873

Type locality: Callao and Payta, Peru.

Systematics: Dixon and Wright (1975), Frost (1992).
Distribution: Western tropical.

Stenocercus aculeatus (O'Shaughnessy, 1879)

Original name: Leiocephalus aculeatus

Type(s): Syntypes (4): BMNH 1946.8.12.33-36

Type locality: Moyobamba, [Departamento San Martin], Peru.
Systematics: Torres-Carvajal (2000b).

Distribution: Eastern tropical, eastern subtropical.

Stenocercus angel Torres-Carvajal, 2000

Type(s): Holotype: QCAZ 3733

Type locality: 8 km NE El Angel on road to Tulcan (77°52' W, 00°40' N),
Provincia Carchi, Ecuador.

Distribution: Andean.

23

Stenocercus carrioni Parker, 1934

Type(s): Holotype: BMNH 1946.8.11.83

Type locality: Zamora, 3250 ft, [Provincia Zamora-Chinchipe], Ecuador;
questioned by Cadle (1991) and Torres-Carvajal (2000b).

Systematics: Cadle (1991), Fritts (1974), Torres-Carvajal (2000b).
Distribution: Western subtropical.

Stenocercus chota Torres-Carvajal, 2000

Type(s): Holotype: QCAZ 3768

Type locality: Panamerican Highway 5 km east of Chota (78°01' W, 00°28' N),
Valle del Chota, Provincia Imbabura, Ecuador.

Distribution: Western subtropical.

Stenocercus festae (Peracca, 1897)

Original name: Liocephalus festae

Type(s): Neotype: QCAZ 4059 (Torres-Carvajal, 2000b)

Type locality: Sevilla de Oro [78°39' W, 02°48' S], 2630 m, Provincia Azuay, Ecuador.
Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western temperate, Andean, eastern temperate.

Stenocercus guentheri (Boulenger, 1885)

Original name: Liocephalus guentheri

Type(s): Syntypes (20): BMNH 58.7.25.16-16a, 59.9.20.6, 60.6.16.18, 60.6.16.20-21,
71.2.7-10, 71.4.16.53 (nine specimens), 80.12.8.53

Type locality: Guayaquil [Provincia Guayas, Ecuador; in error], Sarayacu [Provincia
Pastaza; in error] Ecuador, Western Ecuador, and Colombia; restricted to San
Antonio de Pichincha, 2500 m, Provincia Pichincha, Ecuador, by Fritts (1974).
Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western temperate, Andean, eastern temperate.

Stenocercus haenschi (Werner, 1901)

Original name: Liocephalus haenschi

Type(s): Holotype: ZMB 16595

Type locality: Balsapamba [79°10' W, 01°47' S], 750 m, [Provincia Bolivar], Ecuador.
Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western tropical.

24

Stenocercus humeralis (Giinther, 1859a)

Original name: Microphractus humeralis

Type(s): Syntypes (2): BMNH 1946.8.11.76-77

Type locality: Andes of Ecuador; restricted to Loja [79°12' W, 04°00 S], 2150 m,
Provincia Loja, Ecuador, by Fritts (1974).

Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western temperate, eastern temperate.

Stenocercus iridescens (Gunther, 1859b)

Original name: Liocephalus iridescens

Type(s): Syntypes (6): BMNH 60.6.16.2-7

Type locality: Andes of western Ecuador.
Systematics: Torres-Carvajal (2000b).

Distribution: Western tropical, western subtropical.

Stenocercus limitaris Cadle, 1998

Type(s): Holotype: AMNH 22183

Type locality: Alamor [80°02' W, 04°02’ S], 1325 m, Provincia Loja, Ecuador.
Systematics: Torres-Carvajal (2000b).

Distribution: Western subtropical.

Stenocercus ornatus (Gray, 1845)

Original name: Leiocephalus ornatus

Type(s): Holotype: BMNH 1946.8.29.72

Type locality: Guayaquil [Provincia Guayas, Ecuador; in error]; restricted to
Loja [79°12' W, 04°00 S], 2150 m, Provincia Loja, Ecuador, by Fritts (1974).
Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western subtropical, western temperate, eastern subtropical,
eastern temperate.

Stenocercus rhodomelas (Boulenger, 1899)

Original name: Liocephalus rhodomelas

Type(s): Syntypes (4): BMNH 1946.8.29.77-80

Type locality: Ofia [Provincia Azuay], Ecuador.

Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western tropical, western subtropical, western temperate.

Stenocercus simonsii Boulenger, 1899

Type(s): Syntypes (2): BMNH 1946.8.11.73-74

Type locality: Ofa, 6500 ft, [Provincia Azuay], Ecuador.
Systematics: Cadle (1991), Fritts (1974), Torres-Carvajal (2000b).
Distribution: Western subtropical, western temperate.

25

Stenocercus varius Boulenger, 1885

Type(s): Holotype: BMNH 71.4.16.53

Type locality: Not reported; restricted to Tandapi [78°47' W, 00°25' S], 1460 m,
Provincia Pichincha, Ecuador, by Fritts (1974).

Systematics: Fritts (1974), Torres-Carvajal (2000b).

Distribution: Western subtropical, western temperate.

Tropidurus flaviceps (Guichenot, 1855)

Original name: Doryphorus flaviceps

Type(s): Holotype: MHNP 6882

Type locality: Sarayacu [Provincia Pastaza, Ecuador], Peru.
Systematics: Avila-Pires (1995).

Distribution: Eastern tropical.

Tropidurus plica (Linnaeus, 1758)

Original name: Lacerta plica

Type(s): Holotype: NRM 112

Type locality: Indiis; restricted to vicinity of Paramaribo, Suriname, by
Etheridge (1970).

Systematics: Avila-Pires (1995), Etheridge (1970), Frost (1992).
Distribution: Eastern tropical.

Tropidurus umbra (Linnaeus, 1758)

Original name: Lacerta umbra

Type(s): Syntypes (3): NRM 111 (2 specimens), UUZM 73

Type locality: Meridionalibus; restricted to vicinity of Paramaribo, Suriname,
by Etheridge (1970).

Systematics: Avila-Pires (1995), Etheridge (1970), Frost (1992).

Distribution: Eastern tropical.

SCINCIDAE (1)

Mabuya nigropunctata (Spix, 1825)

Original name: Lacertus mabouya

Type(s): Neotype: MPEG 15248 (Avila-Pires, 1995)

Type locality: Santa Rita, Municipio de Maraa, left margin of Rio Japura
(Lago Parica), Amazonas, Brasil.

Systematics: Dunn (1936).

Distribution: Eastern tropical.

26

TEIIDAE (10)
Ameiva ameiva (Linnaeus, 1758)
Original name: Lacerta ameiva
Type(s): Syntypes (3): NRM 120, 124; UUZM 12
Type locality: America; restricted to the confluence of Cottica river and
Perica Creek, Suriname, by Hoogmoed (1973).
Systematics: Avila-Pires (1995).
Distribution: Eastern tropical.

Ameiva bridgesti (Cope, 1869)
Original name: Holcosus bridgest
Type(s): Holotype: ANSP 9651
Type locality: Ecuador.
Distribution: Western tropical.

Ameiva edracantha Bocourt, 1874

Type(s): Holotype: MHNP 4202

Type locality: Amérique occidentale.

Distribution: Western tropical, western subtropical.

Ameiva orcesi Peters, 1964

Type(s): Holotype: USNM 149655

Type locality: 0.5 km NE Abdon Calderon, Azuay Province, Ecuador.
Distribution: Western subtropical.

Ameiva septemlineata Duméril and Duméril, 1851
Type(s): Holotype: MHNP 4198

Type locality: Amerique meridionale.

Distribution: Western tropical, western subtropical.

Callopistes flavipunctata (Duméril and Bibron, 1839)
Original name: Aporomera flavipunctata

Type(s): Syntypes (2): MHNP 8298

Type locality: South America.

Distribution: Western tropical.

Dicrodon guttulatum Duméril and Bibron, 1839
Type(s): Holotype: MHNP 1116

Type locality: Peru.

Distribution: Western tropical.

27,

Dracaena guianensis Daudin, 1802a

Type(s): Holotype: MHNP 8385

Type locality: Cayenne, French Guiana; restricted to Amapa, Brazil, by
Hoogmoed and Lescure (1975).

Systematics: Avila-Pires (1995).

Distribution: Eastern tropical.

Kentropyx pelviceps Cope, 1868

Type(s): Holotype: ANSP 9556

Type locality: Napo or upper Amazon of Ecuador.
Systematics: Avila-Pires (1995), Gallagher and Dixon (1992).
Distribution: Eastern tropical.

Tupinambis teguixin (Linnaeus, 1758)

Original name: Lacerta teguixin

Type(s): Lectotype: UUZM 13 (Presch, 1973)

Type locality: Indiis; restricted to the vicinity of Paramaribo, Suriname,
by Presch (1973).

Systematics: Avila-Pires (1995), Presch (1973).

Distribution: Eastern tropical.

28

LITERATURE CITED

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Avila-Pires, T. C. S. 1995. Lizards of Brazilian Amazonia (Reptilia: Squamata). Zool. Verh.
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Baur, G. 1890. Das Variieren der Eidechsen-Gattung Tropidurus auf den Galapagos Inseln
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Bell, T. 1825. On a new genus of Iguanidae. Zool. J., 2:204-207.

Bell, T. 1843. Reptiles In C. Darwin, Zoology of the Voyage of the H.M.S. Beagle, under the
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Berthold, A. A. 1846. Mittheilungen tiber das zoologischen Museum zu Gottingen. I.
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Boulenger, G. A. 1881. Description of a new species of Enyalius in the Brussels Museum.
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Boulenger, G. A. 1885. Catalogue of the lizards in the British Museum (Natural History).
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Boulenger, G. A. 1890. First report on additions to the lizard collection in the British
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Boulenger, G. A. 1898. An account of the reptiles and batrachians collected by Mr. W.F.H.
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Boulenger, G. A. 1899. Descriptions of new reptiles and batrachians collected by Mr. P. O.
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Boulenger, G. A. 1902. Descriptions of new batrachians and reptiles from north-western
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preserved in the British Museum. Ann. Mag. Natur. Hist., 10(8):420-424.

Boulenger, G. A. 1914. On a second collection of batrachians and reptiles made by Dr. H.
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Burt, C. E. and M. D. Burt. 1933. A preliminary checklist of the lizards of South America.

Trans. Acad. Sci. St. Louis, 28(1):v+104pp.

29

Cadle, J. E. 1991. Systematics of lizards of the genus Stenocercus (Iguania: Tropiduridae)
from northern Peru: new species and comments on relationships and distribution
patterns. Proc. Acad. Natur. Sci. Phil., 143:1-96.

Cadle, J. E. 1998. New species of lizards, genus Stenocercus (Iguania: Tropiduridae), from
western Ecuador and Peru. Bull. Mus. Comp. Zool., 155(6)257—297.

Coloma, L. A. and A. Quiguango-Ubillus. 2001. Anfibios de Ecuador: lista de especies y
distribucion altitudinal. [on line]. Ver. 1.3 (2 April 2001). Museo de Zoologia, Pontificia
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<http://www.puce.edu.ec/Zoologia/anfecua.htm>

Cope, E. D. 1864. Contributions to the herpetology of tropical America. Proc. Acad. Natur.
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Cope, E. D. 1868. An examination of the reptilia and batrachia obtained by the Orton
expedition to Equador and the upper Amazon, with notes on other species. Proc.
Acad. Natur. Sci. Phil.: 96-140.

Cope, E. D. 1869. Sixth contribution to the herpetology of tropical America. Proc. Acad.
Natur. Sci. Philadelphia: 305-313.

Cope, E. D. 1876. Report on the reptiles brought by Professor James Orton from the middle
and upper Amazon, and western Peru. J. Acad. Natur. Sci. Phil., 8(2):159-183.

Cope, E. D. 1885. Catalogue of the species of batrachians and reptiles contained ina
collection made at Pebas, upper Amazon, by John Hauxwell. Proc. Amer. Phil. Soc.,
23:271-287.

Cope, E. D. 1889. Scientific results of explorations of the U.S. Fish Commission Steamer
Albatross. III.-Report on the batrachians and reptiles collected in 1887-'88. Proc. U.S.
Natl. Mus., 12(769):141-147.

Daudin, F. M. 1802a. Histoire naturelle, générale et particuliére des reptiles. 2:1-432.

Daudin, F. M. 1802b. Histoire naturelle, générale et particuliére des reptiles. 4:1-397.

de Queiroz, K. 1987. Phylogenetic systematics of iguanine lizards. A comparative
osteological study. Univ. California Pub. Zool., 118:xii+203pp.

Dixon, J. R. 1973. A systematic review of the teiid lizards, genus Bachia, with remarks on
Heterodactylus and Anotosaura. Misc. Pub. Mus. Natur. Hist. Univ. Kan., 57:1-47.

Dixon, J. R. 1974. Systematic review of the microteiid genus Iphisa. Herpetologica,
30(2):133-139.

Dixon, J. R. and R. B. Huey. 1970. Systematics of the lizards of the gekkonid genus
Phyllodactylus of mainland South America. Natur. Hist. Mus. Los Angeles Co. Contr.
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Dixon, J. R. and J. W. Wright. 1975. A review of the lizards of the iguanid genus Tropidurus
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