SPECIAL PUBLICATIONS THE MUSEUM TEXAS TECH UNIVERSITY Tiger Beetles of the Genus Cicindela in Arizona (Coleoptera: Cicindelidae) Judy Bertholf '"‘uS. COMP. ZOOL library harvard university April 1983 TEXAS TECH UNIVERSITY Lauro F. Cavazos, President Regents. — B. Joe Pevehouse (Chairman), John E. Birdwell, J. Fred Bucy, Clint Formby, Rex Fuller, Roy K. Furr, Nathan C. Galloway, Anne Phillips, and James L. Snyder. Academic Publications Policy Committee. — J. Knox Jones, (Chairman), Dilford C. Carter, Dale E. Cluff, John R. Darling, Murray C. Havens, Clyde Jones, John L. Kice, C. Reed Richardson, Donald W. Rude, Charles W. Sargent, Connie Steele, and Idris R. Traylor. The Museum Special Publications No. 19 44 pp. 15 April 1983 $7.00 Special Publications of The Museum are numbered serially, paged separately, and published on an irregular basis under the auspices of the Dean of the Graduate School and Director of Academic Publications, and in cooperation with the International Center for Arid and Semi-Arid Land Stu¬ dies. Copies may be purchased from Texas Tech Press, Sales Office, Texas Tech University Library, Lubbock, Texas 79409, U.S.A. Institutions interested in exchanging publications should address the Exchange Librarian at Texas Tech University. ISSN 0149-1768 ISBN 0-89672-097-7 Texas Tech Press, Lubbock, Texas SPECIAL PUBLICATIONS THE MUSEUM TEXAS TECH UNIVERSITY Tiger Beetles of the Genus Cicindela in Arizona (Coleoptera: Cicindelidae) Judy Bertholf No. 19 April 1983 TEXAS TECH UNIVERSITY Lauro F. Cavazos, President Regents. — B. Joe Pevehouse (Chairman), John E. Birdwell, J. Fred Bucy, Clint Formby, Rex Fuller, Roy K. Furr, Nathan C. Galloway, Anne Phillips, and James L. Snyder. Academic Publications Policy Committee. — J. Knox Jones, (Chairman), Dilford C. Carter, Dale E. Cluff, John R. Darling, Murray C. Havens, Clyde Jones, John L. Kice, C. Reed Richardson, Donald W. Rude, Charles W. Sargent, Connie Steele, and Idris R. Traylor. The Museum Special Publications No. 19 44 pp. 15 April 1983 $7.00 Special Publications of The Museum are numbered serially, paged separately, and published on an irregular basis under the auspices of the Dean of the Graduate School and Director of Academic Publications, and in cooperation with the International Center for Arid and Semi-Arid Land Stu¬ dies. Copies may be purchased from Texas Tech Press, Sales Office, Texas Tech University Library, Lubbock, Texas 79409, U.S.A. Institutions interested in exchanging publications should address the Exchange Librarian at Texas Tech University. ISSN 0149-1768 ISBN 0-89672-097-7 Texas Tech Press, Lubbock, Texas 1983 CONTENTS Materials and Methods . 6 Systematics . 6 Key to the Arizona Species of Cicindela . 7 Accounts of Species . 8 Cicindela debilis Bates . 8 Cicindela fulgida julgida Say . 9 Cicindela haemorrhagica Le Conte . 9 Cicindela haemorrhagica haemorrhagica Le Conte . 9 Cicindela haemorrhagica arizonae Wickham . !0 Cicindela hirticollis corpuscula Rumpp . 11 Cicindela hornii hornii Schaupp . 11 Cicindela lemniscata lemniscata Le Conte . 12 Cicindela lengi jordai Rotger . 12 Cicindela lepida Dejean . 13 Cicindela longilabns vestalia Leng . 13 Cicindela marutha Dow . 14 Cicindela nei’adica Le Conte . 14 Cicindela nevadica citata Rumpp . 14 Cicindela nevadica tubensis Cazier . 15 Cicindela nigrocoerulea nigrocoerulea Le Conte . 15 Cicindela obsoleta santaclarae Bates . 16 Cicendela ocellata ocellata Klug . 16 Cicindela oregona Le Conte . 17 Cicindela oregona mancopa Leng . 17 Cicindela oregona navajoensis Van Dyke . 17 Cicindela pimeriana Le Conte . 18 Cicindela praetextata praetextata Le Conte . 18 Cicindela fulgoris Casey . 19 Cicindela fulgoris fulgoris Casey . 19 Cicindela fulgoris erronea Vaurie . 20 Cicindela pulchra Say . 20 Cicindela pulchra pulchra Say . 20 Cicindela pulchra dorothea Rumpp . 21 Cicindela punctulata Olivier . 21 Cicindela purpurea Olivier . 22 Cicindela purpurea cimarrona Le Conte . 22 Cicindela purpurea graminea Schaupp . 23 Cicindela repanda repanda Dejean . 23 Cicindela sedecimpunctata sedecimpunctata Klug . 24 Cicindela sperata sperata Le Conte . 24 Cicindela tenuisignata Le Conte . 25 Cicindela terricola cinctipennis Le Conte . 25 Cicindela tranquebarica kirbyi Le Conte . 26 Cicindela tranquebarica lassenica Casey . 26 Cicindela viridisticta arizonensis Bates . 27 Cicindela wickhami Horn . 27 Cicindela willistom sulfontis Rumpp . 28 Acknowledgments . 28 Literature Cited . 29 Figures . 33 Tiger Beetles of the Genus Cicindela in Arizona (Coleoptera: Cicindelidae) Judy Bert ho If I iger beetles (Coleoptera: Cicindelidae) are active, predatory insects with a worldwide distribution. Bright coloration of the type, and largest genus, Cic¬ indela Linne, has made it the object of extensive collecting by both profes¬ sionals and amateurs. As a result, many manuscripts have been published concerning collecting techniques (Cutler, 1969; Ferris, 1969; Wendler, 1969; Willis, 1971). These insects are housed in many private and museum collec¬ tions throughout the world. According to Vaurie (1950b), there are three habitats in which tiger beetles are almost always found. These are sand hills, alkali lakes or flats, and sand banks. Habitats such as these, which are found throughout most of Arizona, plus warm temperatures, account for that state’s large tiger beetle fauna. In addition, the advent of stock water tanks in recent years has provided numerous ideal habitats for the tiger beetles. Sexually mature adult tiger beetles emerge in the spring and live for two to 10 months. The adult males can be easily distinguished by dense pads of hair on their front tarsi. The adult females have a large bladelike ovipositor for depositing eggs in the ground. The eggs are laid singly and after usually two weeks of incubation hatch into the first larval stage. This larval stage lasts a month, during which time the larvae dig a vertical burrow and prey on arthropods that happen by the entrance. The next two larval stages vary in length depending on the species and a number of environmental factors, including temperature and moisture. The third larval stage is followed by pupal ecdysis, which usually lasts two weeks, and is followed by the last ecdysis. Newly emerged adults remain in the burrow for several days as their pigment develops. There may be either one or two years between generations. If there is one year between generations, the beetles usually hibernate in the third larval stage. If there are two years between generations they may hibernate both years in the larval stage or one year as a larva and one as an adult. I he generations may overlap and color-changes have been observed (Shelford, 1908). In reviewing the Cicindela literature it became clear that although several general works have been published with zoogeograph ical emphasis, out knowledge of the distribution of many species is incomplete. I hese works have largely been arranged according to states, and, until recently, the southwest United States has been virtually ignored (Rumpp, 1956, 1961; Sumlin, 1976). 6 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Papers including information of Cicindela species in Arizona have been published concerning habitats (Davis, 1921; Payne, 1971; Sherman, 1908; Vaurie, 1950b), life-histories (Hamilton, 1925; Shelford, 1908), and biology and control (Frick, 1957). Species descriptions and Arizona distributional records have been scattered throughout the literature (Cazier, 1954; Duncan, 1958; Harris, 1913; Rumpp, 1977; Van Dyke, 1947; Vaurie, 1950). As a result, the total picture of the Cicindela fauna in Arizona has been unclear. Materials and Methods The present study began with an extensive literature search. Descriptions, localities, and habitat notes were reviewed and catalogued. These data were then compared to approximately 10,000 specimens from the following collec¬ tions: Arizona State University, University of Arizona, University of North¬ ern Arizona, University of Chicago, Midwestern State University, Texas Tech University, and several from the personal collection of Mr. Norman Rumpp. Morphological characteristics examined in the systematics of the genus included presence or absence of setae, number and size of teeth on the anterior margin of the labium, and color and size of maculations on the ely¬ tra. After the preliminary morphological study, the various species of Cicin¬ dela were mapped. During the summer of 1978, spot collecting was per¬ formed throughout Arizona at localities where distributions were unclear. The maps were then modified to include the new localities. (Detailed listing of all localities at which each taxon had been taken was found to be eco¬ nomically unfeasible; however, such lists are available directly from the author upon request.) Illustrations of the specimens were added, drawn at uniform scale for ease of comparison. (Illustrations and maps are grouped in the back of the volume.) Diagnoses, remarks, and keys were also included for further identification of the taxa. Thirty-seven nominal taxa of Cicindela occur in Arizona including one new combination, Cicindela haemorrhagica arizonae, and two new syn¬ onyms: Cicindela punctulata chihuahuae under Cicindela punctulata, and Cicindela tranquebarica cibecuei under Cicindela tranquebarica lassenica. Definitions for all terms used in the descriptions of species can be found in Torre-Bueno (1950). Systematics The family Cicindelidae can be distinguished from other beetles by the first abdominal segment, which is divided behind the coxae, and the head, which is as wide as or wider than the thorax. Three genera occur in Arizona: Amblychila Say, Cicindela Linne and Tetracha Hope. The genera may be separated by the following key: I Third joint of the maxillary palpi shorter than the fourth . Cicindela Third joint of the maxillary palpi longer than the fourth . 2 BERTHOLF— TIGER BEETLES OF ARIZONA 7 2. Eyes small, size 20-25 mm . . Amblychila Eyes large, size 25-35 mm . Tetracha Of the three genera, nearly all of the Arizona tigei heroic fauna belong to the genus Cicindela. In this genus there are 37 nominal taxa composed of six monotypic species and 31 subspecies of 23 polytypic species. New county records included in the Arizona distributions are designated by an asterisk. The majority of these new records are in the collection of Arizona State University and were collected by Dr. Mont A. Cazier. Key to the Arizona Species of Cicindela L Front trochanter with subapical seta or pit . 4 Front trochanter without subapical seta ot pit . 2 2. Apical dot or lunule present . 3 Apical dot or lunule absent . ocellata 3. Elytral margins slightly convex, widest at middle or basal third . haemorrhagic a Elytral margins nearly parallel, gradually widening to apical fourth . sedeampunctata 4. Middle trochanter with subapical seta or pit . 7 Middle trochanter without subapical seta or pit . 5 5. Elytron with maculations connected to form a submedian longitudinal stripe . lemniscata Elytron with maculations not forming a submedian longitudinal stripe . 6 6. Elytra impunctate, proepisternum bronze or green with bronze reflection . wickhami Elytra punctate, proepisternum deep green or blue . viridisticta 7. Clypeus glabrous or with very few erect setae . 11 Clypeus clothed with decumbent setae . 8 8. Antennae and legs without pigment, appearing pale tan . lepida Antennae and legs pigmented blue, green or cupreous . 9 9. First antennal segment with distal sensory setae only . 10 First antennal segment with decumbent setae below distal sensory setae . nevadica 10. Proepisternum blue to green . marutha Proepisternum bronze . sperata 1 1. Front of head bare except for ocular setae . 20 Front of head pilose medially . 12 12. Gena pilose . 13 Gena bare . 15 13. First antennal segment with a few erect setae or pits below distal sensory setae . 14 First antennal segment with distal sensory setae only . repanda 14. Apical elytral margins serrate, color cupreous . lengi Apical elytral margins not senate, color blue-black or green . purpurea 15. First antennal segment with several erect setae or pits below distal sensors setae . 16 First antennal segment with distal sensory setae only . hirticollis 16. Elytra immaculate or with maculations consisting of humeral dot and may have short transverse line . pimeriana Elytra maculations consisting of more than humeral dot and transverse lint' . 17 8 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY 17. 19. 20. 21. 22. 23. 23. 25. 26. 27. 28. Elytra dull . 18 Elytra shiny . ^ Labrum unidentate, proepisternum blue green with cupreous reflections . willistom Labrum tridentate, proepisternum rosy with some cupreous reflections . tranquebarica Proepisternum green with cupreous reflections, medium size (11-12 mm) . julgida Proepisternum blue with green reflections, large size (15-17 mm) . pulchra Elytra with subsutural row of fovea extending from base to near apex . 21 Elytra with obscure fovea, none at all, or lovea only at base . 22 Apical elytral margins serrate . punctulata Apical elytral margins not serrate . mgrocoerulea Proepisternum glabrous . debilis Proepisternum pilose, may be just a few setae near coxal margin . 23 Apical elytral margins serrate . 23 Apical elytral margins not serrate . 26 Middle band long and slender, and curving inward . tenuisignata Middle band short and wide or represented by a bulge . 25 Genae with sparse, decumbent setae, body robust . praetextata Genae without setae, body elongate . fulgoris Elytra immaculate . hornii Elytra with maculations . 27 First antennal segment pilose medially, below sensory setae . longilabns First antennal segment with distal sensory setae only . 28 Size large (19-21 mm), pronotum smooth and dull . obsolete. Size medium (10-11 mm), pronotum granulate and shiny . terricola Accounts of Species Cicindela debilis Bates (Figs. 3, 34) Cicindela debilis Bates, 1890:509; W. Horn, 1897:182; W. Horn, 1 903b;2 1 9: Harris, 1911:52; W. Horn, 1915:389; W. Horn, 1916:13; Leng, 1920:32; Cazier, 1954:287; Rumpp, 1977:177. Cicindela debilis var . segnis Harris, 1913:69; W. Horn, 1915:389; Cazier, 1953:287. Type data. — Type(s) from “Cuidad in Durango, Mexico” (Cazier, 1954); British Museum (Natural History), London. General distribution. — Parts of north-central Mexico into southwestern United States. Arizona distribution. — Cochise and Santa Cruz counties (Fig. 34). Diagnosis . — Color blue green dorsally; purple blue ventrally. Head finely rugose, bald; labrum unidentate; clypeus glabrous; gena bald; first antennal segment with sensory setae only. Thorax granulate, slightly pilose laterally; proepisternum bare, purple blue. Legs purple blue, with setae on front and middle trochanters. Abdomen bald to sparsely hairy. Elytra (Fig. 3) evenly punctate, rounded at apex with spine, widest at middle. Length 7-10 mm. BERTHOLF— TIGER BEETLES OF ARIZONA 9 Remarks. — The two varietal names of this species were synonymized by Cazier (1954). Th is species is very distinct and shows little variation in size, color, and maculation. Adults can be collected in sandy fields from July to November. Cicindela fulgida fulgida Say (Figs. 4, 55) Cicindela fulgida Say, 1823:141; Dunn, 1891:153; Wickham, 1899:217; Leng, 1902a: 139; Smyth, 1907-1908:182; W. Horn, 1916:7; Leng, 1920:41; Cazier, 1936:159; Vaurie, 19506:146; Wallis, 1961:51; Graves, 1963:494; Willis, 1967:153; Ferris, 1969:10. Type data. — Type(s) from “Missouri Territory,’’ near the mountains on the Platte and Arkansas rivers; Academy of Natural Sciences of Philadelphia; lost or destroyed. General distribution. — Southwestern and midwestern United States north¬ ward to Wyoming. Arizona distribution . — Coconino and Navajo counties (Fig. 35). Diagnosis. — Color red with green and cupreous reflections dorsally, green with cupreous reflections ventral ly. Flead rugose, pilose medially; labrum unidentate; clypeus glabrous; gena bare; first antennal segment with prox¬ imal erect setae. Thorax rugose, pilose laterally; proepisternum pilose, green with cupreous reflections. Legs green with cupreous reflections, with setae on front and middle trochanters. Abdomen pilose. Elytra (Fig. 4) glossy, densely punctate, serrate at apex, with spine, widest at apical fourth. Length 10-12 mm. Remarks. — Adults of this polytypic species can be collected in June and July along streams and mud flats in northeast Arizona. Members of this spe¬ cies are easily identified and show little variation in size and maculation. Cicindela haemorrhagica Le Conte Key to Subspecies of Cicindela haemorrhagica Le Conte Color black, black blue to black green, body robust . haemorrhagica Color reddish brown, body size reduced . arizonae Hybrid populations between haemorrhagica and arizonae have been found in southern Utah and northern Arizona (Rumpp, personal communication). Cicindela haemorrhagica haemorrhagica Le Conte (Figs. 5, 36) Cicindela carthagena var. haemorrhagica Le Conte, 1851:171; Le Conte, 1857:55; Bates, 1881- 1884:9; Schaupp, 1883-1884:105; Dunn, 1891:154; Blaisdell, 1892:48; G. Horn, 1892:27; G. Horn, 1894:306; W. Horn, 1897:179; Wickham, 1899:217; W. Horn, 19036:218; Smyth, 1907-1908:181; W. Horn, 1915:388; W. Horn, 1916:13; Leng, 1920:42; Moore, 1937:109; Hatch, 1938:236; LaRivers, 1946:139; Cazier, 1948:11. Cicindela haemorrhagica haemorrhagica : Leng, 1920:176; Cazier, 1954:282; Rumpp, 1954:131; Frick, 1957:503. 10 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Type data. — Type(s) from San Diego, San Diego County, California (Caz- ier, 1954); Museum of Comparative Zoology, Harvard University, Cam¬ bridge. General distribution . — From Baja California Norte extending northward through Washington and eastward into Texas. Arizona distribution . — Cochise, Graham, Maricopa, Pima, and Yuma counties (Fig. 36). Diagnosis . — Color black with some green reflections dorsally, black ven- trally. Head finely striate, bald; labrum unidentate; clypeus glabrous; gena bald; first antennal segment with sensory setae only. Thorax feebly rugose, slightly pilose laterally; proepisternum pilose, black. Legs black, front and middle trochanters without setae. Abdomen slightly pilose laterally. Elytra (Fig. 5) faintly and evenly punctured, apex finely serrate, with spine, widest at middle. Length 10-16 mm. Remarks. — This widely distributed subspecies occurs along streams, lakes, and alkali flats. Adults can be found from June to September. T here is much variation throughout its range; the maculations may be broad and complete or consist of broken lunules, in which case specimens resemble C. sedecim- punctata. The two can be distinguished from each other by characters in the key and by the broadness of the maculations found in C. haemorrhagica. Cicindela haemorrhagica arizonae Wickham (new combination) (Fig. 36) Cicindela rufiventris var. arizonae Wickham, 1899:226; Leng, 1902a: 178. Cicindela carthagena arizonae Smyth, 1907-1908:181; W. Horn, 1915:388; W. Horn, 1916:13; Leng, 1920:42. Type data. — Male syntypes from canyon of Colorado River (date unknown, R. Hayward; July 1982, Townsend); University of Iowa. General distribution. — Arizona and Utah. Arizona distribution. — Coconino County (Fig. 36). Diagnosis. — Color brown red with purple and green reflections dorsally, cupreous ventrally. Head striate, bald; labrum unidentate; clypeus glabrous; gena bald; first antennal segment with sensory setae only. Thorax rugose, pilose laterally; proepisternum pilose, cupreous with red reflections. Legs cupreous, front and middle trochanters without setae. Abdomen pilose later¬ ally. Elytra granulate, apex serrate, with spine, widest at middle. Length 10- 12 mm. Remarks. — Cazier (1954) recognized C. haemorrhagica as a distinct species. Until now arizonae has remained as a subspecies under C. carthagena', how¬ ever, arizonae is more closely related to C. haemorrhagica as is evident by its smaller size and reduced maculations. Therefore, arizonae is now placed as a subspecies under C. haemorrhagica. Adults of this subspecies are found from May to August in the Grand Canyon. They have the markings of C. haemorrhagica but can be distin¬ guished by their brilliant red brown color. BERTHOLF— TIGER BEETLES OF ARIZONA 1 1 Cicindela hirticollis corpuscula Rumpp (Figs. 6, 37) Cicindela hirticollis corpuscula Rumpp, 1961:174. Type data. — Holotype, male, from Potholes, Imperial County, California (12 May 1946, N. L. Rumpp); the Norman L. Rumpp collection. General distribution . — Parts of southwestern and western United States. Arizona distribution. — Coconino*, Graham, Greenlee*, Maricopa, Navajo*, Pinal*, and Yuma counties (Fig. 37). Diagnosis. — Color dull bronze dorsally, reddish bronze ventrally. Head finely striate, pilose; labium unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax slightly rugose, pilose lat¬ erally; proepisternum pilose, cupreous. Legs reddish bronze, front and mid¬ dle trochanters with setae. Abdomen slightly pilose. Elytra (Fig. 6) shallowly punctate throughout, apex serrate, with spine, widest in anterior half. Length 11-12 mm. Remarks. — Adults of this subspecies can be collected from April to November. They are primarily located on sand banks along the Colorado River and its tributaries. This species somewhat resembles C. repanda but can easily be distinguished by its larger size and somewhat pointed abdo¬ men. Cicindela hornii hornii Schaupp (Figs. 7, 38) Cicindela anthracina G. Horn, 1880: 139 (preoccupied). Cicindela hornii Schaupp, 1883-1884:80, Leng, 1 902a: 1 27; W. Horn, 1903b:216; Smyth, 1907- 1908:188; W. Horn, 1916:9; Leng, 1920:41: Cazier, 1954:247; Rumpp, 1977:177. Cicindela ritten Bates, 1890:496; W. Horn, 1900:116; Leng, 1920:41. Type data. — Type(s) from Fort Bayard, Grant County, New Mexico (Caz- ier, 1954); type depository unknown. General distribution . — North-central Mexico northward into southwestern United States. Arizona distribution . — Cochise and Pima counties (Fig. 38). Diagnosis . — Color entirely glossy black to purple black to metallic green. Head finely rugose, bald; labium unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax smooth to slightly rugose, single row of short hair laterally; proepisternum slightly pilose. Legs pigmented, setae on front and middle trochanters. Abdomen glabrous. Elytra (Fig. 7) coarsely punctate near base, feebly serrate at apex, with weak spine, widest at middle. Length 13-15 mm. Remarks. — With the exception of color, the characters in this species are stable. They can easily be distinguished by their glabrous, immaculate appearance. This species is rare, but adults may be found at high altitude, from June to August, along rocky hillsides. They are somewhat difficult to catch. 12 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Cicindela lemniscata lemniscata Le Conte (Figs. 8, 39) Cicindela lemniscata Le Conte, 1854:220; Le Conte, 1857:59; Dunn, 1891:152; G. Horn, 1892:27; G. Horn, 1894:307; W. Horn, 1897:184; Wickham, 1899:224; Leng, 1902a: 174; W. Horn, 1903b:220; Smyth, 1905:252; Smyth, 1907-1908:187; Casey, 1913:16; W. Horn, 1915:384; W. Horn, 1916:11; Leng, 1920:41; W. Horn, 1926:170; Cazier, 1948:15; Vogt, 1949:6; Cazier, 1954:263; Rumpp, 1977:177. Type data. — Male, holotype, from San Diego, California to El Paso, Texas (1850-1852, T. H. Webb); Museum of Comparative Zoology, Harvard Univer¬ sity, Cambridge. General distribution . — Northwestern Mexican coastal states extending into parts of western and southwestern United States. Arizona distribution. — Cochise, Gila, Graham, Maricopa, Pima, Pinal, Santa Cruz, Yavapai, and Yuma counties (Fig. 39). Diagnosis . — Color cupreous red dorsally, cupreous red ventrally. Head striate to granulate-rugose, bald; labrum unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax rugose, slightly pilose laterally; proepisternum bare, blue. Legs cupreous with green reflec¬ tions, setae on front trochanters only. Abdomen pilose. Elytra (Fig. 8) deeply granulate, serrate at apex, with spine, widest at apical fourth. Length 7-10 mm. Remarks. — Throughout the summer, adults may be collected at night around electric lights, or during the day along streams, ponds, and lakes. They are easily identified and show little variation in size, maculation, and coloration. Cicindela lengi jordai Rotger (Figs. 9, 39) Cicindela lengi jordai Rotger, 1974:9. Type data. — Holotype, male, from Heart Canyon, 4 miles north of Aztec, San Juan County, New Mexico (29 May 1970, B. Rotger), collection of Reve¬ rend B. Rotger. General distribution. — Parts of southwestern United States. Arizona distribution. — Apache* County (Fig. 39). Diagnosis . — Color red with green and blue reflections dorsally, blue green with cupreous reflections ventrally. Head granulate, pilose; labrum trident- ate; clypeus glabrous; gena pilose. Thorax striate, pilose laterally; proepis¬ ternum pilose, blue green. Legs blue green with cupreous reflections, setae on front and middle trochanters. Abdomen pilose. Elytra (Fig. 9) granulate at base, smooth at apex, apex serrate with spine, widest at apical fourth. Length 12-15 mm. Remarks. — Adults of (his recently described subspecies are present from May to June along sandy banks of dry washes. Their large size and broad maculations make them distinctive. The only Arizona specimen was taken in June just west of Fort Defiance in Apache County. BERTHOLF— TIGER BEETLES OF ARIZONA 13 Cicindela lepida Dejean (Figs. 10, 40) Cicindela lepida Dejean, 1831:255; Le Conte, 1848:181; Le Conte, 1857:51; Jones, 1884:74; Leng and Beutenmuller, 1894:95; Wickham, 1899:224; Knaus, 1900:109; Leng, 1902a: 169; Leng, 1902b:239; Smyth, 1905:252; Griddle, 1907:105; Fall, 1907:155; Smyth, 1907- 1908:180; Shelford, 1908:168; Blatchley, 1910:27; Fox, 1910:75; Knaus, 1915:35; Dow, 1916:69; VV. Horn, 1916:18; Leng, 1920:43; Hamilton, 1925:32; Dawson and Horn, 1928:13; Cazier, 1936:28; Vaurie, 1950b: 148; Cazier, 1954:297; Wallis, 1961:67; Graves, 1963:501; Graves, 1973:191. Type data. — Type(s) from “America septentrionale”; University of Paris. General distribution. — Chihuahua, Mexico northward into southern Can¬ ada; extending eastward from Nevada through Gulf and midwestern states to north-Atlantic states. Arizona distribution . — Apache*, Coconino, Mojave, and Navajo* counties (Fig. 40). Diagnosis . — Color bronze with green reflections dorsally, green bronze centrally. Head striate, pilose; labium unidentate; clypeus with decumbent setae; gena bare; first antennal segment pilose medially, not pigmented. Thorax rugose, pilose laterally and medially; proepisternum pilose, green bronze. Legs not pigmented, pale tan, front and middle trochanters with setae. Abdomen densely pilose. Elytra (Fig. 10) with shallow punctations, apex obtuse, faint spine to no spine, widest at apical fourth. Length 9-12 mm. Remarks. — This monotypic species has an extensive range throughout the United States. In Arizona, adults can be found from June to August. They are easily distinguished by their light color and are most often collected on sandy slopes. Cicindela longilabris vestalia Leng (Figs. 1 1 , 34) Cicindela longilabris var. vestalia Leng, 1 902a: 121; Long, 1920:41; Hatch, 1938:235. Type data. — Lectotype, female, designated by Dahl (1941), from Maiden, Silver Bow County, Montana (17 June 1890, F. C. Bowditch); American Museum of Natural History, New York. General distribution. — Arizona northward to Montana. Arizona distribution. — Apache* and Coconino counties (Fig. 34). Diagnosis. — Color entirely black, brown, or green. Head granulate-rugose, bald; labrum tridentate; clypeus glabrous; gena bald; first antennal segment pilose medially. Thorax striate, pilose laterally; proepisternum bare to slightly pilose. Legs pigmented, front and middle trochanters with setae. Abdomen glabrous to slightly pilose. Elytra (Fig. 11) granulate, rounded to apex, with spine, widest at middle. Length 15-16 mm. Remarks. — Adults of this rather rare species occur at high elevations from June to August. Maculations are variable and so narrow that the elytra may appear immaculate. Specimens may occasionally be found on gravel along roadsides. 14 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Cicindela marutha Dow (Figs. 12, 41) Cicindela sperata var. marutha Dow, 1911:272; Leng, 1920:42. Cicindela sperata var. rubicunda Harris, 1911:55. Cicindela marutha : Cazier, 1954:296. Type data. — Syntypes from Fort Wingate, McKinley County, New Mexico (J. Woodgate); type depository unknown. General distribution. — Chihuahua, Mexico northward into southwestern Fhiited States. Arizona distribution . — Apache*, Cochise, Coconino*, Maricopa*, Mojave*, and Navajo* counties (Fig. 41). Diagnosis . — Color green to blue green or cupreous red dorsally, blue green ventrally. Head granulate-striate, pilose; labium unidentate; clypeus pilose; gena bare; first antennal segment with sensory setae only. Thorax striate, pilose laterally and medially; proepisternum pilose, blue green with cupre¬ ous reflections. Legs green with cupreous reflections, front and middle tro¬ chanters with setae. Abdomen pilose laterally. Elytra (Fig. 12) punctate basal to near smooth apical, somewhate obtuse; apex serrate, with spine, widest at apical third. Length 13-14 mm. Remarks. — From June to August adults of this monotypic species can be found on alkali flats or near seepage (see discussion under C. sperata spe¬ rata). Cicindela nevadica Le Conte Key to Subspecies of Cicindela nevadica Le Conte Color reddish brown, markings broad . tubensis Color brownish bronze, with green micropits, middle band narrow . citata Cicindela nevadica citata Rumpp (Fig. 42) Cicindela nevadica citata Rumpp, 1977:170. Type data. — Holotype, male, from eight kilometers WSW Willcox, Cochise County, Arizona (20 July 1970, N. L. Rumpp); type no. 12528, Cali¬ fornia Academy of Sciences, San Francisco. General distribution . — Sonora, Mexico northward into Arizona. Arizona distribution . — Cochise County (Fig. 42). Diagnosis. — Color brown bronze with green reflections dorsally, brown with green reflections ventrally. Head granulate, pilose; labium unidentate; clypeus pilose; gena pilose; first antennal segment pilose medially. Thorax feebly granulate, pilose laterally and medially; proepisternum pilose, bronze. Legs green with cupreous reflections, front and middle trochanters with setae. Abdomen pilose laterally. Elytra punctate with blue green micropits, subsutural row of fovea, apex obtuse, serrate, with spine, widest at apical third to middle. Length 10-1 1 mm. BERTHOLF— TIGER BEETLES OF ARIZONA 15 Remarks. — This recently described subspecies is found in (he Sulphur Springs Valley in southeastern Arizona. It can be distinguished from C. nevadica tubensis by its bronze green color. Adult specimens have been col¬ lected in July or August along playa lakes. Cicindela nevadica tubensis Cazier (Figs. 13, 42) Cicindela nevadica tubensis Cazier, 1 936:24; Vaurie, 1951:7. Type data. — Holotype, female, from Tuba City, Coconino County, Ariz¬ ona (5 July 1937, R. P. Allen); American Museum of Natural History, New York. General distribution. — Southwestern United States. Arizona distribution. — Apache, Coconino, and Navajo Counties (Fig. 42). Diagnosis . — Color cupreous red dorsally, cupreous with some green reflec¬ tions ventrally. Head granulate-striate, pilose; labium unidentate; clypeus pilose; gena pilose; first antennal segment pilose medially. Thorax striate, pilose medially and laterally; proepisternum bronze, pilose. Legs cupreous with some green reflections, front and middle trochanters with setae. Abdo¬ men pilose laterally. Elytra (Fig. 13) with shallow green punctures, rounded to apex, with spine, widest at middle. Length 11-12 mm. Remarks. — This subspecies can be distinguished for C. nevadica citata by its cupreous red color. Adults are found from June to August in large popu¬ lations along bodies of water. Members of this species take flight when approached but with care can be caught by hand. Cicindela nigrocoerulea nigrocoerulea Le Conte (Figs. 4, 43) Cicindela nigrocoerulea Le Conte, 1848:181; Leng, 1 902a: 1 23; W. Horn, 1903b:213; Smyth, 1905:252; Smyth, 1907-1908:180; Casey, 1909:267; W. Horn, 1916:9; Leng, 1920:41; Caz¬ ier, 1954:246. Type data. — Type(s) from Arkansas River, probably in Colorado (Cazier, 1954); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution. — North-central Mexico extending into southwestern United States. Arizona distribution . — Apache*, Cochise, Coconino*, Gila, Graham, Maricopa*, Navajo*, Pima, Pinal*, Santa Cruz*, and Yavapai counties (Fig. 43). Diagnosis. — Color blue to green to black with cupreous reflections dor- sally, purple blue ventrally. Head finely rugose, bald; labium tridentate; cly¬ peus glabrous; gena glabrous; first antennal segment with sensory setae only. Thorax finely rugose, pilose laterally; proepisternum pilose, blue green. Legs blue green, front and middle trochanters with setae. Abdomen sparsely pilose. Elytra (Fig. 14) with base heavily punctate, apex almost smooth, sub- sutural row of fovea, apex rounded to small spine, widest at middle. Length 10-14 mm. 16 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Remarks. — This species shows variation throughout its range. Adults may be immaculate or have complete maculations (Fig. 14). Specimens found along the Puero River have a reddish outline along the edge of the elytra. Color may range from brilliant blue to dull green to black. Adults are found from June to October in fairly dry areas near alkaline soil. I his species can be differentiated from C. punctulata because its elytra are widest at middle and not serrate at apex. Cicindela obsoleta santaclarae Bates (Figs. 15, 44) Cicindela obsoleta santaclarae Bates, 1 890:493; W. Horn, 1 903 b: 2 1 6; Smyth, 1907-1908:188; Casey, 1909:268; W. Horn, 1916:9; Leng, 1920:41; Cazier, 1954:249. Type data. — Type(s) from Santa Clara, Chihuahua, Mexico (Cazier, 1954); British Museum (Natural History), London. General distribution. — Durango, Mexico northward into Arizona and New Mexico. Arizona distribution. — Apache*, Cochise, Gila*, Navajo*, Pima*, Pinal*, Santa Cruz, and Yavapai* counties (Fig. 44). Diagnosis. — Color entirely green to blue green to black to red. Head finely striate, bald; labium tridentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax smooth to finely rugose, pilose lat¬ erally, proepisternum pilose. Legs pigmented, front and middle trochanters with setae. Abdomen sparsely pilose. Elytra (Fig. 15) with minute punctures at base, smooth at apex, apex serrate to smoothly rounded, widest at middle. Length 20-22 mm. Remarks. — Adults of this species can be easily distinguished by their large size. Somewhat rare, they can be found at high elevations, from June to Sep¬ tember, in short grass. Maculations vary and may consist of lunules or may be broken into dots. Cicindela ocellata ocellata Klug (Figs. 16, 40) Cicindela ocellata Klug, 1834:33; Graves, 1973:180. Cicindela flavopunctata Chevrolat, 1833-1835:2; Bates, 1881-1884:10; Bates, 1890:505; Dunn, 1891:152; W. Horn, 1897:177; W. Horn, 19036:217; Knaus, 1906:147; Smyth, 1907- 1908:185; W. Horn, 1915:387; W. Horn, 1916:12; Leng, 1920:42; Vogt, 1949:1; Cazier, 1954:275. Type data. — Type(s) from Mexico (Cazier, 1954); probably part of the Walther Horn Collection at Berlin-Dahlem, East Berlin. General distribution. — Northward from Central America and southern and central Mexico into southwestern United States. Arizona distribution. — Cochise, Gila*, Graham, Pima*, Pinal*, and Santa Cruz* counties (Fig. 40). Diagnosis. — Color cupreous brown with blue green reflections dorsally, cupreous bronze ventrally. Head granulate, bald; labium unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax BERTHOLF— TIGER BEETLES OE ARIZONA 17 smooth to faintly rugose, pilose laterally; proepisternum cupreous, pilose. Legs green with cupreous reflections, front and middle trochanters without setae. Abdomen pilose. Elytra (Fig. 16) shallowly to deeply punctate, serrate at apex, with spine, widest at apical third. Length 9-10 mm. Remarks. — Adults of this species occur along the southern border of Ariz¬ ona from June to October. They are often taken at electric lights, or in sandy areas near water. Specimens show little variation with respect to size, color, and maculation. Cicindela oregona Le Conte Key to Subspecies of Cicindela oregona Le Conte Color dark purple . Color cupreous . . . maricopa navajoensis Cicindela oregona maricopa Leng (Figs. 17, 45) Cicindela oregona maricopa Leng, 1902a:H7; Smyth, 1907-1908:181; W. Horn, 1916:7; Leng, 1920:41. Cicindela oregona mormonella Casey. 1924:15. Cicindela oregona provensis Casey, 1924:15. Cicindela oregona nephiana Casey, 1924:16. Type data. — Lectotype, male, designated by Dahl (1941), from Phoenix, Maricopa County, Arizona; American Museum of Natural History, New York. General distribution. — California westward to New Mexico. Arizona distribution. — Cochise*, Coconino, Gila*, Greenlee*, Maricopa, Mohave*, Navajo*, Pima*, Pinal, and Yavapai counties (Fig. 45). Diagnosis . — Color blue green to purple dorsally, blue green ventrally. Head granulate, bald; labium tridentate; clypeus glabrous; gena bare; first antennal segment pilose medially. Thorax granulate-striate, pilose laterally; proepisternum pilose, blue green. Legs blue green, front and middle tro¬ chanters with setae. Abdomen sparsely pilose. Elytra (Fig. 17) granulate, finely serate at apex, with spine, widest at middle. Length 11-14 mm. Remarks. — Maculations in this subspecies show little variation, whereas coloration may range from blue green to purple. Adults occur from April to October along streams. Cicindela oregona navajoensis Van Dyke (Fig. 45) Cicindela oregona navajoensis Van Dyke, 1947:155. Type data. — Holotype, male, from 15 miles WNW Kayenta, Navajo County, Arizona (17-24 June 1933, H. N. Hultgren); type number 5864, Cali¬ fornia Academy of Sciences, San Francisco. General distribution. — Parts of southwestern United States. Arizona distribution. — Coconino* and Navajo counties (Fig. 45). 18 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Diagnosis . — Color cupreous dorsally, metallic green ventrally. Head granulate-striate, bald; labium tridentate; clypeus glabrous; gena bare; lirst antennal segment pilose medially. Thorax granulate-striate, pilose laterally; proepisternum pilose, green with cupreous reflections. Legs cupreous, front and middle trochanters with setae. Abdomen bare to sparsely pilose. Elytra with green punctures, serrate at apex, with spine, w’idest at middle. Length 11-14 mm. Remarks. — This subspecies has the maculations of C. oregona mancopa but can be distinguished by its cupreous color. Adults are found in northern Arizona, from June to July, along streams. They are sometimes confused with C. repanda repanda, from which they differ by their usually broken humeral and apical lunules and their first antennal segment, which is pilose medially. Cicindela pimeriana Le Conte (Figs. 18, 46) Cicindela viatica Le Conte, 1857:62 (preoccupied). Cicindela pimeriana Le Conte, 1866:363; Leng, 1 902a: 127; W. Horn, 1903b:215; Smyth, 1907- 1908:186; W. Horn, 1916:7; Leng, 1920:41; Cazier, 1954:242; Rumpp, 1977:177. Cicindela pimeriana var. cochisensis Casey, 1909:274. Type data. — Type(s) from Sonora, Mexico (Cazier, 1954); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution. — Sonora, Mexico, northward into Arizona and New Mexico. Arizona distribution. — Cochise County (Fig. 46). Diagnosis . — Color entirely blue to blue green, glossy. Head granulate to finely rugose, pilose; labium tridentate; clypeus glabrous; gena bare; first antennal segment pilose medially. Thorax finely rugose, pilose laterally; proepisternum pilose, blue green. Legs blue green, front and middle tro¬ chanters with setae. Abdomen pilose. Elytra (Fig. 18) shallowly punctate, serrate at apex, with spine, widest at apical third. Length 11-13 mm. Remarks. — This monotypic species occurs in the southeastern corner of Arizona. Adidts can be found from June to August, near water. Specimens show little variation in size and shape; however, their maculations vary from a humeral spot and a transverse line to immaculateness. Despite this diver¬ sity, their glossy blue-green color makes them easy to distinguish from other species. Cicindela praetextata praetextata Le Conte (Fig. 47) Cicindela praetextata Le Conte, 1854:220; Le Conte, 1857:58; Schaupp, 1833-1884:104; W. Horn, 1897:183; Acciavatti, 1980:233-236. Cicindela circumpicta praetextata-. Leng, 1902a: 171. Cicindela californica praetextata: W. Horn, 1 903b:2 1 9; Casey, 1913:33; W. Horn, 1915:391; W. Horn, 1916:14; Leng, 1920:42; Cazier, 1948:24. Cicindela praetextata: Cazier, 1954:288. BERTHOLF— TIGER BEETLES OF ARIZONA 19 Type data. — Syntypes, males, from San Diego, California, to El Paso, Texas (1850-1852, T. H. Webb); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution. — Arizona, California, and Utah. Arizona distribution. — Gila, Graham, Maricopa, Mohave, Pinal, Yavapai, and Yuma counties (Fig. 47). Diagnosis . — Color light brown dorsally, bronze ventrally. Head striate, bald; labium unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax smooth to slightly rugose, pilose medially and laterally; proepisternum pilose, bronze. Legs bronze, front and middle trochanters with setae. Abdomen pilose. Elytra evenly punctate, serrate at apex, with spine, widest at apical third. Length 12-14 mm. Remarks. — This subspecies has the maculations of C. fulgoris fulgons (big. 19) but can be distinguished by its dull, light brown color. From June to August, adult specimens are commonly collected on sandbanks through¬ out central Arizona. Cicindela fulgoris Casey Key to the Subspecies of Cicindela fulgoris Casey Color cupreous red or cupreous green . fulgoris Color dull green or blue with no cupreous reflections . erronea Cicindela fulgoris fulgoris Casey (Figs. 19, 47) Cicindela praetextata ssp .fulgoris Casey, 1913:34. Cicindela praetextata ssp . stnngens Casey, 1913:34. Cicindela fulgoris Acciavatti, 1980:233-236. Type data. — Type(s) from El Paso, El Paso County, Texas; United States National Museum, Washington, D.C. General distribution. — Southwestern United States. Arizona distribution . — Cochise and Navajo* counties (Fig. 47). Diagnosis . — Color cupreous red, may have green reflections dorsally, bronze with green reflections ventrally. Head granulate, bald; labium uni¬ dentate; clypeus glabrous; gena bare; first antennal segment with sensory- setae only. Thorax rugose, pilose medially and laterally; proepisternum pilose, bronze. Legs bronze or green, front and middle trochanters with setae. Abdomen densely pilose. Elytra (Fig. 19) punctate throughout, serrate at apex, with spine, widest at apical third. Length 1 1-12 mm. Remarks. — From July to September, adults of this subspecies can be found on sand and alkali flats. They can be distinguished from C. praetextata prae¬ textata by their red color and more elongate size. 20 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Cicindela fulgoris erronea Vaurie (Fig. 47) Cicindela californica viridicyanea Vaurie, 1 950a: 1 (preoccupied). Cicindela californica erronea Vaurie, 1951:12; Rumpp, 1956:131. Cicindela praetextata erronea Vaurie (new combination, Rumpp, 1957:144). Cicindela fulgoris erronea Vaurie: Acciavatti, 1980:233-236. Type data. — Holotype, male, from YVillcox, Cochise County, Arizona (1 September 1947, F. H. Parker); American Museum of Natural Ffistory, New York. General distribution. — Arizona. Arizona distribution . — Cochise County (Fig. 47). Diagnosis. — Color blue or green dorsally, blue ventrally. Head striate, bald; labium unidentate; clypeus glabrous, gena bald; first antennal segment with sensory setae only. Thorax striate, pilose laterally; proepisternum pil ose, blue green. Legs blue with green reflections, front and middle tro¬ chanters with setae. Abdomen pilose. Elytra evenly punctate throughout, ser¬ rate at apex, with spine, widest at apical third. Length 12-13 mm. Remarks. — Adults of this subspecies are found only in the Sulphur Springs Valley of Arizona. Their blue green color makes them easy to iden¬ tify. Specimens show little variation with regard to size and shape, although their maculations also resemble those of C. fulgoris fulgoris (Fig. 19). They can be collected from July to September on mud flats at the water’s edge. Cicindela pulchra Say Key to the Subspecies of Cicindela pulchra Say Color red with violet reflections, immaculate or with reduced maculations . pulchra Color red with green reflections, maculations usually connected along margin . dorothea Cicindela pulchra pulchra Say (Figs. 20, 48) Cicindela pulchra Say, 1823:142; Ulke, 1875:813; Wickham, 1899:213; Smyth, 1907-1908:188; W. Horn, 1916:7; Leng, 1920:41. Type data. — Type from “Missouri Territory,’’ along the Platte and Arkan¬ sas rivers near the mountains; Academy of Natural Sciences of Philadelphia; lost or destroyed. General distribution. — Texas northward into Wyoming. Arizona distribution . — Apache County (Fig. 48). Diagnosis . — Color purple-violet with green reflections, glossy dorsally, blue to blue green ventrally. Head granulate, pilose; labium tridentate; cly¬ peus glabrous; gena bare; first antennal segment pilose medially. Thorax smooth to slightly rugose, pilose laterally; proepisternum pilose, blue green. Legs blue green, front and middle trochanters with setae. Abdomen sparsely pilose. Elytra (Fig. 20) punctate, rounded at apex, faint spine, widest at api¬ cal fourth. Length 16-18 mm. BERTHOLF— TIGER BEETLES OF ARIZONA 21 Remarks. — Adults ot this subspecies occur from July to August on bate sand and gravel. Coloration ranges from violet to deep purple to red, and maculations may consist of isolated spots or be absent. Members of this sub¬ species are easily recognizable. Cicindela pulchra dorothea Rumpp (Figs. 21, 48) Cicindela pule lira dorothea Rumpp, 1977:170. Type data. — Holotype, male, from 5.2 to 6.2 kilometers SE of Wilcox, Cochise County, Arizona (26 July 1946, N. L. and D. H. Rumpp); type number 12529, California Academy of Sciences, San Francisco. General distribution. — Arizona. Arizona distribution. — Cochise County (Fig. 48). Diagnosis . — Color red orange with blue green reflections, glossy dorsally, blue to blue green ventrally. Head slightly rugose, pilose; labrum tridentate; clypeus glabrous; gena bare; first antennal segment pilose medially. Thorax rugose, pilose laterally; proepisternum pilose, blue green. Legs blue to blue green, front and middle trochanters with setae. Abdomen slightly pilose. Ely¬ tra (Fig. 21) punctate, smooth and rounded at apex, with spine, widest at apical fourth. Length 15-16 mm. Remarks. — In July and August, this orange red subspecies has been col¬ lected on the open flatlands of southeastern Arizona. Maculations are varia¬ ble and may be connected along the outer margin or reduced to dots. This subspecies can be distinguished from C. pulchra pulchra by its color, broad maculations, and smaller size. Cicindela punctulata Olivier (Fig. 22) Cicindela punctulata Olivier, 1790:27; Gould, 1834:54; Le Conte, 1848:182; Leng and Beuten- muller, 1894:93; Wickham, 1899:218; Leng, 1902a:158; Leng, 1902b:239; W. Horn, 1 903 b:2 1 7 ; Brimley, 1906:81; Smyth, 1907-1908:181; Griddle, 1907:105; Shelford, 1908:168; Blatchley, 1910:27; Fox, 1910:75; Johnson, 1915:307; Knaus, 1915:35; Gold¬ smith, 1916:447; W. Horn, 1916:10; Leng, 1920:41; Blanchard, 1921:396; Hamilton, 1925:41; Dawson, 1928:6; Cartwright, 1935:73; Vogt, 1949:6; Vaurie, 1950b: 150; Cazier, 1954:251; Wallis, 1961:61; Graves, 1963:503; Willis, 1967:154; Graves, 1973:117. Cicindela punctulata chihuahuae Bates, 1890:500; Leng, 1920:41; Fanner, 1928:269; Cazier, 1954:251; new synonym. Type data. — Type(s) from New Jersey (Cazier, 1954); Museum National d’Histoire Naturelle, Paris; destroyed. General distribution. — Widespread in North America, from north-central Mexico northward to southern Canada. Arizona distribution. — Apache, Cochise, Coconino, Gila*, Maricopa, Mohave,* Navajo*, Pima*, Santa Cruz*, and Yavapai counties. Diagnosis.— Color brown to green to blue to purple dorsally, blue green to purple ventrally. Head granulate-rugose, bald; labrum unidentate; clypeus 99 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY glabrous; gena bare; first antennal segment with sensory setae only. I hoi ax granulate to nearly smooth, pilose laterally; proepisternum pilose, cupreous to blue green. Legs green with cupreous to blue green reflections, front and middle trochanters with setae. Abdomen pilose. Elytra (big. 22) punctate throughout, subsutural row of green fovea, serrate at apex, with spine, wid¬ est at apical fourth. Length 10-14 mm. Remarks. — In the past, the subspecific name chihuahuae was given for the high altitude green to purple color form, and punctulata for the low altitude green to brown form of this species. There has since been extensive collect¬ ing and mapping (Mayr, 1969:178) of this common species at high, low, and intermediate altitudes. These studies have shown wide ranges of color at nearly every altitude, clearly examples of color variation. Therefore, chihua¬ huae is placed in synonymy. Adult specimens of this common species can be collected throughout most of Arizona from May to September along roadsides, alkali flats, or near water. They are strong fliers and often difficult to catch. Elytra may be immaculate or maculations may consist of isolated spots (see discussion under C. mgrocoerulea). Cicindela purpurea Olivier Key to the Subspecies of Cicindela purpurea Olivier Color black to black green with maculations usually complete around margin . cimarrona Color green with cupreous reflections, maculations not complete around margin . graminea Cicindela purpurea cimarrona Le Conte (Figs. 23, 49) Cicindela purpurea cimarrona Le Conte, 1868:49; Leng, 1 902a: 1 34; Smyth, 1907-1908:183; Leng, 1920:40. Type data. — Syntypes, six male(s), female(s) from south of Raton Moun¬ tain, Colfax County, New Mexico (1867, S. Lewis); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution. — Arizona, Colorado, and New Mexico. Arizona distribution . — Graham County (Fig. 49). Diagnosis. — Color entirely black or green. Head rugose, pilose, labrum tridentate; clypeus glabrous; gena pilose; first antennal segment pilose medially. Thorax granulate, pilose laterally; proepisternum slightly pilose. Legs pigmented, front and middle trochanters with setae. Abdomen sparsely pil ose. Elytra (Fig. 23) granulate-punctate, smooth at apex, with weak spine, widest at apical fourth. Length 12-15 mm. Remarks. — This subspecies has been collected at over 8,000 feet in the Pinaleno Mountains. Adults occur from June to August on bare soil. Speci¬ mens show little variation in size, color, and maculation, and are easily iden¬ tified. BERTHOLF— TIGER BEETLES OF ARIZONA 23 Cicindela purpurea graminea Schaupp (Figs. 24, 49) Cicindela purpurea graminea Schaupp, 1883-1884:89; Dunn, 1891:153; Wickham, 1899:215; Lcng, 1902a: 132; Smyth, 1907-1908:180; Leng, 1920:40; Hamilton, 1925:27; Dawson and Horn, 1928:9; LaRivers, 1946:135. Type data. — Type(s) from Kansas and California; type depository unknown. General distribution. — Western United States northward into southwestern Canada. Arizona distribution. — Apache*, Coconino, and Greenlee* counties (Fig. 49). Diagnosis . — Color dark green dorsally, dark blue with some green reflec¬ tions ventrally. Head granulate-striate, pilose; labium tridentate; clypeus glabrous; gena pilose; first antennal segment pilose medially. Thorax granu¬ late, pilose laterally; proepisternum pilose, rosy. Legs green with cupreous reflections, front and middle trochanters with setae. Abdomen sparsely pilose. Elytra (Fig. 24) shallowly punctate, smooth at apex, with weak spine, widest at apical fourth. Length 10-12 mm. Remarks. — Adults occur from July to August throughout most of Arizona along barren soil or sandy beaches. They are easily identified by their colora¬ tion and narrow maculations. Cicindela repanda repanda Dejean (Figs. 25, 50) Cicindela repanda Dejean, 1825:74; Jones, 1884:74; Dunn, 1891:154; Leng and Buetenmuller, 1894:93; Wickham, 1899:215; Knaus, 1900:109; Leng, 1902a: 1 47: Hood, 1903:113; Smyth, 1905:252; Moore, 1906:338; Griddle, 1907:105; Smyth, 1907-1908:184; Shelford, 1908:170; Blatchley, 1910:27; Fox, 1910:75; Davis, 1912:18; Goldsmith, 1916:447; Leng, 1920:40; Blanchard, 1921:396; Hamilton, 1925:31; Dawson and Horn, 1928:7; W. Horn, 1930:81; Cartwright, 1935:71; Hatch, 1938:233; Vaurie, 1 950b: 151; Willis, 1967:154; Chaves, 1969:86; Graves, 1973:168. Cicindela unijuncta Casey, 1897:299. Type data. — Types(s) from “America septentrionale”; University of Paris. General distribution. — Widespread in the United States and southern Can¬ ada. Arizona distribution. — Maricopa County (Fig. 50). Diagnosis. — Color brown with some green reflections dorsally, green blue with bronze reflections ventrally. Head granulate, pilose; labium tridentate; clypeus glabrous; gena pilose; first antennal segment with sensory setae only. Thorax rugose, pilose laterally; proepisternum rosy, pilose. Legs cupreous with green reflections, front and middle trochanters with setae. Elytra (Fig. 25) granulate-punctate, subsutural row of green fovea, serrate at apex, with weak spine, widest at middle. Length 11-12 mm. Remarks. — Adults of this species are found throughout the summer along sandy beaches. Although markings are stable they are often confused with 24 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY other species (see discussion under C. hirticolhs corpuscula and C. oregona navajoensis). Cicindela sedecimpunctata sedecimpunctata Klug (Figs. 26, 51) Cicindela sedecimpunctata Klug, 1834:32; Le Conte, 1857:57; Schaupp, 1883-1884.106; Bates, 1881-1884:259; Bates, 1890:503; W. Horn, 1897:172; Smyth, 1907-1908:181; Casey, 1913:15; Cazier, 1954:270. Cicindela rufiventris var. sedecimpunctata-. Leng, 1902a: 178; W. Horn, 1 903b:2 1 6; W. Horn, 1915:386; W. Horn, 1916:12. Type data. — Type(s) from Mexico (Cazier, 1954); probably from the Walther Horn Collection at Berlin-Dahlem, East Berlin. General distribution . — Guanajuato, Mexico northwestward into south¬ western United States. Arizona distribution . — Cochise, Coconino*, Gila*, Graham, Greenlee, Navajo*, Pima, Pinal, and Santa Cruz counties (Fig. 51). Diagnosis. — Color brown with some blue and green reflections dorsally, bronze ventral ly. Head striate, bald; labium unidentate; clypeus glabrous; gena bald; first antennal segment with sensory setae only. Thorax rugose, pilose laterally; proepisternum pilose, bronze with blue green reflections. Legs bronze with green reflections, front and middle trochanters without seta. Abdomen pilose. Elytra (Fig. 26) evenly punctate throughout, serrate at apex, with spine, widest at apical fourth. Length 10-13 mm. Remarks. — Adults occur in large numbers on mud flats or along sandy shores. They may be found with C. haemorrhagica but can be differentiated by size and coloration (see discussion under C. haemorrhagica). Members of this species are active from May to November and can be easily collected during the hottest part of the day as they conceal themselves under debris in the shade at the water’s edge. Cicindela sperata sperata Le Conte (Figs. 27, 52) Cicindela sperata Le Conte, 1857:50; G. Horn, 1876:239; Wickham, 1899:206: Leng, 1 902a: 1 67 : W. Horn, 19036:219; Smyth, 1907-1908:185; W. Horn, 1916:17; Leng, 1920:42; Vogt, 1949:1; Cazier, 1954:296. Type data. — Type(s) from “Rio Grande, at various places’’ (Cazier, 1954); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution . — Northern Mexico northward into southwestern United States. Arizona distribution . — Apache*, Cochise*, Coconino, Navajo*, and Yuma counties (Pig. 52). Diagnosis. — Color brown cupreous dorsally, green ventrally. Head rugose, pilose; labrum unidentate; clypeus pilose; gena bare; first antennal segment with sensory setae only. Thorax striate, pilose medially and laterally; proep¬ isternum pilose, bronze. Legs bronze, front and middle trochanters with BERTHOLF — TIGER BEETLES OF ARIZONA 2.5 setae. Abdomen pilose. Elytra (Fig. 27) closely punctate throughout, serrate at apex, with spine, widest at apical third. Length 12-14 mm. Remarks. — Adults occur from June to August on mud flats and at water banks. Superficially, they resemble C. marutha but are distinguished by the following characters: deeper punctations on the elytra, bronze proepister- num, and the striate appearance of the thorax. Cicindela tenuisignata Le Conte (Figs. 28, 46) Cicindela tenuisignata Le Conte, 1851:171; Dunn, 1891:152; Wickham, 1899:220; Leng, 1902a: 153; W. Horn, 1903b:218; W. Horn, 1916:10; Leng, 1920:41; La Rivers, 1946:138; Vogt, 1949:6; Cazier, 1954:256. Cicindela psilogramma Bates, 1890:507. Type data. — Type(s) from “Colorado Desert,’’ probably California (Cazier, 1954); Museum of Comparative Zoology, Harvard University, Cambridge. General distribution. — Northern Mexico (including Baja California Norte) northward into parts of western and southwestern Lhiited States. Arizona distribution. — Cochise, Coconino*, Gila*, Graham, Maricopa, Navajo*, Pima*, Pinal*, Yavapai*, and Yuma* counties (Fig. 46). Diagnosis . — Color brown bronze dorsally, metallic green with bronze reflections ventrally. Head striate, bald; labium unidentate; clvpeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax granulate, pilose laterally; proepisternum bronze, pilose. Legs bronze, front and middle trochanters with setae. Abdomen pilose. Elytra (Fig. 28) punctate, serrate at apex, with spine, widest at apical fourth. Length 11-12 mm. Remarks. — Adults of this monotypic species are found throughout most of Arizona from May to November. They show little variation and are easily identified by a narrow descending middle band. They most often occur in ditches and on alkali flats. Cicindela terricola cinctipennis Le Conte (Figs. 29, 38) Cicindela pusilla var. cinctipennis Le Conte, 1848:182; Le Conte, 1857:45; Schaupp, 1 883- 1884:82; Dunn, 1891:154; Wickham, 1899:206; Leng, 1902a: 155; W. Horn. 1915:390; W. Horn, 1916:14; Leng, 1920:42; Wallis, 1961:64; Rumpp, 1961:171. Cicindela terricola cinctipennis : Huber, 1969:19. Type data. — Type(s) from the Rocky Mountains; Museum of Comparative Zoology, Harvard University, Cambridge. General distribution . — Arizona north to Alberta and Saskatchewan. Arizona distribution . — Apache*, Coconino, and Navajo counties (tig. 38). Diagnosis.— Color bronze with blue green reflections dorsally, blue green ventrally. Head striate, bald; labium tridentate; c lvpeus glabrous; gena bald; first antennal segment with sensory setae only. 1 horax granulate-striate, pilose laterally and medially; proepisternum pilose, green. Legs green with bronze reflections, front and middle trochanters with seta. Abdomen pilose 26 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY laterally. Elytra (Fig. 29) punctate throughout, smooth at apex, with spine, widest at apical fourth. Length 9-12 mm. Remarks. — From July to September, adults of this species may be collected in short grass along the edges of alkali flats, or on muddy shores. I hey resemble no other species and are easily distinguished. Cicindela tranquebarica kirbyi Le Conte (Figs. 30, 53) Cicindela obliquata Dejean, 1825:72; Blaisdel 1, 1892:48; Leng, 1902a: 145 (preoccupied). Cicindela tranquebarica kirbyi Le Conte, 1866:362; Wickham, 1899:206; W. Horn, 1916:6; Leng, 1920:40; Dawson and Horn, 1928:8; Cazier, 1936:27; La Rivers, 1946:136. Type data. — Type(s) from “America septentrionale”; Lnited States National Museum, Washington, D.C. General distribution . — Texas northward into southern Canada. Arizona distribution. — Apache*, Coconino, Gila#, Navajo*, and Mari¬ copa* counties (Fig. 53). Diagnosis. — Color coppery brown with some blue green reflections dor- sally, blue green with cupreous reflections centrally. Head striate-granulate, pilose; labium tridentate; clypeus glabrous; gena bald; first antennal segment pilose medially. Thorax granulate, sparsely pilose laterally; proepisternum pilose, rosy. Legs bronze, front and middle trochanters with setae. Abdomen pilose laterally. Elytra (Fig. 30) granulate at base, punctate at apex, serrate at apex, weak spine to no spine, widest at apical fourth. Length 13-15 mm. Remarks. — Adults of this subspecies may be found from April to June along lakes, stream, and in grassy fields near water. Specimens show little variation. They can be distinguished from C. willistoni sulfontis by charac¬ ters in the key, and from C. tranquebarica lassenica by their brown colora¬ tion. Cicindela tranquebarica lassenica Casey (Fig. 53) Cicindela tranquebarica lassenica Casey, 1914:22. Cicindela tranquebarica cibecuei Duncan, 1958:43; new synonym. Type data. — Type locality reported as probably California (Casey, 1914). Rumpp (personal communication) has indicated Casey’s material is identical to populations below the Mogollon Rim, Gila and Navajo counties, Ari¬ zona; United States National Museum, Washington, D.C. General distribution. — Arizona. Arizona distribution. — Gila and Navajo counties (Fig. 53). Diagnosis. — Color entirely navy blue to black. Head striate-granulate, pil ose; labium tridentate; clypeus glabrous; gena bald; first antennal segment pilose medially. Thorax striate, pilose laterally; proepisternum pilose. Legs pigmented, front and middle trochanters with setae. Abdomen pilose later¬ ally. Elytra granulate at base, punctate at apex, serrate at apex, with weak spine to no spine, widest at apical fourth. Length 13-15 mm. BERTHOLF— TIGER BEETLES OF ARIZONA 27 Remarks. — Norman Rumpp (personal communication) lias compared types ol tranquebarica lassenica Casey to tranquebarica cibecuei Duncan, and found them to be the same. Casey listed the type locality as probably California, but it is actually one of the populations below the Mogollon Rim in east-central Arizona. Following Rumpp’s suggestion, I propose to place tranquebarica cibecuei in synonymy under tranquebarica lassenica. Adults of this subspecies have a distinctive coloration and are found on dry sand above creek beds from April to June. Cicindela viridisticta arizonensis Bates (Figs. 31, 35) Cicindela viridisticta arizonensis Bates, 1881-1884:1; W. Horn, 1935:65; Cazier, 1956:284. Cicindela arizonensis: W. Horn, 19036:219; W. Horn, 1915:388: W. Horn, 1916:13; Leng, 1920:42. Cicindela viridisticta: Schaupp, 1883-1884:103; Leng, 1902a: 1 15; W. Horn, 1903a:182; Smyth, 1907-1908:188; Rumpp, 1977:178. Type data. — Type(s) from Arizona (Cazier, 1954); British Museum (Natural History), London. General distribution. — Chihuahua, Mexico northward into Arizona. Arizona distribution . — Cochise, Gila#, Maricopa*, Pima, and Santa Cruz* counties (Fig. 35). Diagnosis . — Color brown bronze dorsally, greenish blue ventrally. Head granulate-striate, bald; labium unidentate; clypeus glabrous; gena bare; first antennal segment with sensory setae only. Thorax granulate-striate, pilose laterally, proepisternum bald, deep green or blue. Legs green with cupreous reflections, front trochanters with setae, middle trochanters without. Abdo¬ men sparsely pilose. Elytra (Fig. 31) evenly, deeply punctate, subsutural row of fovea, serrate at apex, with weak spine, widest at apical fourth. Length 7-8 mm. Remarks. — Adults of this subspecies show little variation throughout their range. They occur from July to October in short grass along streams, ditches, and ponds. Superficially, this subspecies resembles C. wickami, but it can be distinguished by the subsutural row of fovea and a more convex appearance. Cicindela wickhami Horn (Figs. 32, 48) Cicindela wickhami W. Horn, 1 903a : 1 82; Smyth, 1907-1908:180; Leng, 1920:42; W. Horn, 1926:170; Cazier, 1948:18; Cazier, 1954:284. Cicindela lernniscata: Smyth, 1907-1908:188 (misidentification). Type data. — Types(s) from Tucson, Pima County, Arizona (Cazier, 1954); Berlin-Dahlem, East Berlin. General distribution. — Northwestern Mexico (including Baja California Norte) northward into Arizona. 28 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Arizona distribution . — Cochise, Pima, Pinal, and Santa Cruz counties (Fig. 48). Diagnosis . — Color cupreous green to cupreous red dorsally, cupreous, green, and blue ventrally. Head granulate-striate, bald; labrum unidentate; clypeus glabrous; gena bald; first antennal segment with sensory setae only. Thorax granulate-striate, pilose laterally and medially; proepisternum bald, bronze or green with bronze reflections. Legs brown with cupreous reflec¬ tions, front and middle trochanters with setae. Abdomen pilose. Elytra (Fig. 32) with shallow green punctations, weakly serrate at apex, with no spine to weak spine, widest at apical fourth. Length 6-8 mm. Remarks. — Adults may be collected from June to July in bare areas next to water (see discussion under C. viridisticta arizonensis). Cicindela ivillistoni sulfontis Rumpp (Figs. 33, 50) Cicindela ivillistoni sulfontis Rumpp, 1977:170. Type data. — Holotype, male from 5.6 kilometers WSW of Willcox, Cochise County, Arizona; type number 12530, California Academy of Scien¬ ces, San Francisco. General distribution . — Arizona. Arizona distribution . — Cochise County (Fig. 50). Diagnosis . — Color dark blue green, green bronze or dull brown dorsally, blue green or green with blue and cupreous reflections ventrally. Head granulate-striate, pilose; labrum unidentate; clypeus glabrous; gena bald; first antennal segment pilose medially. Thorax rugose, pilose laterally; proepisternum pilose, blue green with cupreous reflections. Legs blue green or green with cupreous reflections, front and middle trochanters with setae. Abdomen pilose. Elytra (Fig. 33) with even green punctations throughout, weakly serrate, with spine, widest at middle. Length 13-14 mm. Remarks. — Adults of this species occur from mid-July to August on mud flats in the Sulphur Springs Valley. Maculations show little variation, but occur in two color forms, green and brown (see discussion under C. tran- quebanca kirbyi). Acknowledgments The author is deeply indebted to several people for their support and advice: Dr. Raymond Jackson, Texas Tech University; Dr. William P. Morrison, Texas Tech University; and Dr. David Foster, University of Kentucky. Thanks are also due to Mr. Norman Rumpp for speci¬ mens and advice, to Arizona State University for the loan of specimens, to Mr. Bruce Hall for helping with the drawings, to Miss Kim Gazier and Mr. Martin Mortenson for help in the field, and to the Interlibrary Loan Office at Texas Tech University. Most importantly, the author wishes to give special thanks to Dr. Mont Cazier, Arizona State University, and Dr. Oscar Francke, Texas Tech University, fot their support, advice, and friendship. This study was sup¬ ported in part by the Institute for Museum Research, Texas Tech Elniversity. BERTHOLF— TIGER BEETLES OE ARIZONA 29 Literature Cited Acciavatti, R. E. 1980. A review of Cicindela praetextata from the southwest United States (Coleoptera: Cicindclidae). Southwestern Entomol. 5(4): 131-1 45. Bates, H. W. 1881-1884. Biologia Centrali-Americana, Coleoptera, Cicindelidae, 1 ( 1 ) : 1 - 1 8. - . 1890. 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Tiger beetles of the genus Cicindela in southwest Nevada and Death Val¬ ley, California, and descriptions of two new subspecies. Bull. S. California Acad. Sci., 55:131-144. - . 1957. Notes on the Cicindela praetextata calijornica tiger beetle complex. Descriptions of a new subspecies from Death Valley, California. Bull. S. California Acad. Sci., 58:144-154. - . 1961. Three new tiger beetles of the genus Cicindela from southwestern ETited States (Cicindelidae-Coleoptera). Bull. S. California Acad. Sci., 60:165-187. - . 1977. Tiger beetles of the genus Cicindela in the Sulphur Springs Valley, Arizona, with descriptions of three new subspecies. Proc. California Acad. Sci., 4 1 (4 ) : 1 69- 1 82. Say, T. 1823. Descriptions of the Coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. J. Acad. Nat. Sci. Philadelphia, 3:139-331. Schaupp, F. G. 1883-1884. Synoptic tables of North American Coleoptera: Cicindelidae. Bull. Brooklyn Entomol. Soc., 6:73-126. Shelford, V. E. 1908. Life-histories and larval habits of the tiger beetles (Cicindeli¬ dae). Linnean Soc. J. Zook, 30:157-184. Sherman, F. 1908. Notes on tiger beetle elevations. Entomol. News, 19:360-362. Smyth, E. G. 1905. Notes on collecting Cicindelidae. Trans. Kansas Acad. Sci., 19:252-260. - . 1907-1908. Notes on collecting Cicindelidae-II. Kansas Acad. Sci., 21:180-188. Sumlin, W. D. 1976. A new subspecies of Cicindela politula from west Texas and a note on Cicindela cazieri (Coleoptera: Cicindelidae). J. Kansas Entomol. Soc., 94(4):52 1 -526. Tanner, V. M. 1928. The Coleoptera of Zion National Park, Utah. Ann. Entomol. Soc. Amer., 21:269-280. 32 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY I orre-Bueno, J. R. de la. 1950. A glossary of entomology. Brooklyn Entomol. Soc., Brooklyn, 345 pp. Van Dyke, E. C. 1947. New species of Coleoptera from western North America. Pan-Pacific Entomol., 23(4): 155. Vaurie, P. 1950a. Font new subspecies of the genus Cicindela (Coleoptera, Cicindeli- dae). Amer. Mus. Novitates, 1458:1-6. - . 1950b. Notes on the habits of some North American tiger beetles. J. New York Entomol. Soc., 58:143-153. - . 1951. Five new subspecies of tiger beetles of the genus Cicmdela and two corrections. Amer. Mus. Novitates, 1479:1-12. Vogt, G. B. 1949. Three new Cicindelidae from south Texas with collecting notes on other Cicindelidae (Coleoptera). Bull. Brooklyn Entomol. Soc., 44(1): 1-9. Wallis, J. B. 1961. Cicindelidae of Canada. Univ. of Toronto Press, Toronto, 74 pp. W endler-Funaro, C. de. 1969. “Tiger hunting” in every land. Cicindela, 1 (2): 15-18. Wickham, H. U 1899. The habits of American Cicindelidae. Proc. Davenport Acad. Nat. Sci., pp. 206-228. \\ illis, H. L. 1967. Bionomics and zoogeography of tiger beetles of saline habitats in the central Elnited States (Coleoptera: Cicindelidae). Kansas Univ. Sci. Bull., 47:145-313. - . 1971. Collecting Cicindelidae in the northwest. Cicindela, 3:41-51. Address of author: J. Bertholf, Entomology Hall, Purdue University, West Lafayette , Indiana -17907. Received 16 July 1981 , accepted 20 April 1982. BERTHOLF— TIGER BEETLES OF ARIZONA 33 basal dot humeral dot posthumeral dot marginal dot extra dot supplementary dot discal dot anteapical dot .apical dot Figs. 1-2. — Types of elytral markings. 34 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY 8 Eigs. 3-8— Dorsal view of Cicindela species: 3, C. debilis Bates: 4, C. fulgida fulgida Say 5 C haemorrhagica haemorrhagica Le Conte; 6. C. hirticollis corpuscula Rumpp; 7, C. hornii hor- nn Schaupp; 8, C. lemniscata lemniscata Le Conte. BERTHOLF— TIGER BEETLES OF ARIZONA 35 Fics. 9-14. — Dorsal view of Cicindela spec ies: 9, C. lengi jordai Rotger; 10. C. lepida Dejean; 1, c. longilabris vestalia Leng; 12, C. marutha Dow; 13. C. nevadica tubensis Cazier; M. C. igrocoerulea nigrocoerulea Le Conte. 36 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY I-igs. 15-19. Dorsal view of Cicindela species: 15, C. obsoleta santaclarae Bates; 16, C. ocel- lata ocellata King; 17, C. oregona maricopa Leng; 18, C. pimeriana Le Conte; 19, C. fulgoris fu Igor is Casey. BERTHOLF— TIGER BEETLES OF ARIZONA 37 Figs. 20-24. — Dorsal view of Cicindela species: 20, C. pulchra pule lira Say; 21, C. puli lira dorothea Rumpp; 22, C. punctulata punctulata Olivier: 23, C. purpurea cimarrona Le Conte; 24, C. purpurea graminea Schaupp. 38 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY BERTHOLF— TIGER BEETLES OF ARIZONA 31 Figs. 31-33.— Dorsal view of Cicindela species: 31, C. viridkticta arizonensis Bates; 32, C wickhami W. Horn; 33, C. luillistom sulfontis Rumpp. 40 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Figs. 34-37. — Arizona distribution of Cicindela species: 34, C. debilis Bates (wavy lines), and C. longilabris vestalia Leng (straight lines); 35, C. fulgida fulgida Say (wavy lines), and C. vir- idisticta anzonensis Bates (straight lines); 36, C. haemorrhagica arizonae Wickham (wavy lines), and C. haemorrhagica haemorrhagica Le Conte (straight lines); 37, C. hirticollis corpuscula Rumpp. BERTHOLF— TIGER BEETLES OF ARIZONA 41 lines), and C. terricola cinctipennis Le Conte (straight lines); 39, C. lengi jordai Rotger (wavy lines), and C. lemniscata lemmscata Le Conte (straight lines); 40, C. ocellata ocellata Klug (wavy lines), and C. lepida Dejean (straight lines); 41, C. marutha Dow. SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY 42 Figs. 42-45. — Arizona distribution of Cicindela species: 42, C. nevadica citata Rumpp (wavy lines), and C. nevadica tubensis Cazier (straight lines); 43, C. nigrocoerulea nigrocoerulea Le Conte; 44, C. obsoleta santaclarae Bates; 45, C. oregona navajoensis VanDyke (wavy lines), and C. oregona maricopa Leng (straight lines). BERTHOLF— TIGER BEETLES OF ARIZONA 43 Figs. 46-49. — Arizona distribution of Cicindela species: 46, C. pimeriana Le Conte (wavy lines), and C. tenuisignata Le Conte (straight lines); 47, C. fulgoris fulgoris Casey (wavy lines), C. praetextata praetextata Le Conte (straight lines), and C. fulgoris erronea Vaurie (dots); 48. C. pulchra pulchra Say (wavy lines), C. wickhami W. Horn (straight lines), and C. pulchra doro- thea Rumpp (dots); 49, C. purpurea cimarrona Le Conte (wavy lines), and C. purpurea grami- nea Schaupp (straight lines). 44 SPECIAL PUBLICATIONS MUSEUM TEXAS TECH UNIVERSITY Figs. 50-53. — Arizona distribution of Cicindela species: 50, C. willistoni sulfontis Rumpp (wavy lines), and C. repanda repanda Dejean (straight lines); 51, C. sedecimpunctata sedecim- punctata King; 52, C. sperata sperata Le Conte; 53, C. tranquebarica lassenica Casey (wavy lines), and C. tranquebarica kirbyi Le Conte (straight lines). Copies of the following numbers of Special Publications of The Museum may be purchased from Texas Tech Press, Sales Office, Texas Tech University Library, Lubbock, Texas 79409, U.S.A. Institutions interested in exchanging publications should address the Exchange Librarian at Texas Tech University. No. 1 Watkins, L. C., J. K. Jones, Jr., and H. H. Genoways. 1972. Bats of Jalisco, Mexico, 44 pp., 3 figs . $2.00 No. 2 Krishtalka, L. 1973. Late Paleocene mammals from the Cypress Hills, Alberta, 77 pp., 21 figs . $4.00 No. 3 West, R. M. 1973. Review of the North American Eocene and Oligocene Apatemyidae (Mammalia: lnsectivora), 42 pp., 20 figs . $2.00 No. 4 Gardner, A. L. 1973. The systematics of the genus Didelphis (Marsupialia: Didelphidae) in North and Middle America, 81 pp., 14 figs . $4.00 No. 5 Genoways, H. H. 1973. Systematics and evolutionary relationships of spiny pocket mice, genus Liomys, 368 pp., 66 figs . $10.00 No. 6 Northington, D. K. 1974. Systematic studies of the genus Pyrrhopappus (Compositae, Cichorieae), 38 pp., 14 figs . $2.00 No. 7 King, M. E., and I. R. Traylor, Jr., eds. 1974. Art and environment in native America. 169 pp . $8.00 No. 8 Pence, D. B. 1975. Keys, species and host list, and bibliography for nasal mites of North American birds (Acarina: Rhinonyssinae, Turbinoptinae, Speleognathinae, and Cv- toditidae), 148 pp., 728 figs . $6.00 No. 9 Bowles, J. B. 1975. Distribution and biogeographv of mammals of Iowa, 184 pp., 62 figs. . $8.00 No. 10 Baker, R. J., J. K. Jones, Jr., and D. C. Carter, eds. 1976. Biology of bats of the New World family Phyllostomatidae. Part 1., 218 pp . $8.00 No. II Foster, D. E. 1976. Revision of North American Trichodes (Herbst) (Coleoptera: Cleri- dae), 86 pp.. 17 figs . $4.00 No. 12 Mitchell, R. W., W. H. Russell, and W. R. Elliott. 1977. Mexican eyeless characin fishes, genus Astyanax: environment, distribution, and evolution. 89 pp., 21 figs . $5.00 No. 13 Baker, R. J., J. K. Jones, Jr., and D. C. Carter, eds. 1977. Biology of bats of the New World family Phyllostomatidae. Part II., 364 pp . $16.00 No. 14 Francke. O. F. 1978. Systematic revision of diplocentrid scorpions (Diplocentridae) from circum-Caribbean lands, 93 pp., 146 figs . $7.00 No. 15 Carter. D. C.. and P. G. Dolan. 1978. Catalogue of type specimens of Neotropical bats in selected European museums, 136 pp . $8.00 No. 16 Baker. R. J., J. K. Jones. Jr., and D. C. Carter, eds. 1979. Biology of bats of the New World family Phyllostomatidae. Part III., 441 pp . $20.00 No. 17 Wilhelm, R. B., J. R. Choate, and J. K. Jones, Jr. 1981. Mammals of Dicreek National Wildlife Refuge, South Dakota, 39 pp., 3 figs . $4.00 No. 18 Martin, C. O., and D. J. Schmidly. 1982. Taxonomic Review of the Pallid Bat, Antro- zous pallid us (Le Conte), 48 pp.. 1 1 figs . $7.00 No. 19 Bertholf, J. 1983. Tiger beetles of the genus Cicindela in Arizona (Coleoptera: Cieindeli- dae), 44 pp., 53 figs . .00 iPECIAL PUBLICATIONS THE MUSEUM TEXAS TECH UNIVERSITY IVhistling-ducks: Zoogeography, Ecology, Anatomy Zric G. Bolen and Michael K. Rylander mu 3. COMP. ZOOL LIBRARY May 1983