S
S934
Stuttgarter Beitrage zur Naturkunde
Serie A (Biologie)
Herausgeber:

Staatliches Museum ftr Naturkunde, Rosenstein 1, D-70191 Stuttgart

| Stuttgarter Beitr. Naturk. | Ser. A | Nr. 598 | 27ES: | Stuttgart, 15. 11. 1999

Cryptophagidae (Coleoptera) from the Himalayas,
with Descriptions ot New Species,
Keys and Remarks to some Languriidae”)

By Georgy Lyubarsky, Moscow

With 23 figures

Summary

Species of the Coleoptera family Cryptophagidae (Cryptophagus pedarius sp. n., Cryp-
tophagus honoratus sp. n., Himascelis nepalensis sp. n., Himascelis gracilis sp. n., Himascelis
turgidus sp. n., Atomaria accola sp. n., and Atomaria mentita sp. n.) are described from Ne-
pal. The males of Cryptophagus malaisei Bruce 1945, Cryptophagus anxiosus Grouvelle 1916,
Loberus sinuaticollis Bruce 1945, Atomaria torrida Johnson 1970 and the female of Atomaria
plecta Lyubarsky 1996 are described and illustrated for the first time. Several species of Cryp-
tophagidae as well as 2 species of Languriidae are recorded for the first time from Nepal. Iden-
tification keys for the species of the genera Cryptophagus, Himascelis and Atomaria from the
Himalayan Region are provided.

Zusammentassung

Arten der Kaferfamilie Cryptophagidae (Cryptophagus pedarius sp. n., Cryptophagus ho-
noratus sp. n., Himascelis nepalensis sp. n., Himascelis gracilis sp. n., Himascelis turgidus sp.
n., Atomaria accola sp. n. und Atomaria mentita sp. n.) werden aus Nepal beschrieben. Die
Mannchen von Cryptophagus malaisei Bruce 1945, Cryptophagus anxiosus Grouvelle 1916,
Loberus sinuaticollis Bruce 1945, Atomaria torrida Johnson 1970 und das Weibchen von Ato-
maria plecta Lyubarsky 1996 werden erstmalig beschrieben und abgebildet. Zahlreiche Arten
der Cryptophagidae ebenso wie 2 Arten der Languriidae werden erstmalig aus Nepal gemel-
det. Bestimmungsschlüssel für die Arten der Gattungen Cryptophagus, Himascelis-and Ato-
maria aus dem Himalaya werden beigefügt.

APR 12 2000

1 LIBRARIES
*) Results of the Himalaya Expeditions of J. MARTENS, no. ene 224 see: Boll. Mus.
reg. Sci. nat. Torino, 1999.- J. M. sponsored by Deutscher Akademischer Austauschdienst
and Deutsche Forschungsgemeinschaft.

| Me

ay ;
Oe.

a en I

2932
Stuttgarter Beitrage zur Naturkunde
Serie A (Biologie)
Herausgeber:

Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart

Stuttgarter Beitr. Naturk. Ser. A Nr. 598 DAS: Stuttgart, 15. 11. 1999
8 8

Cryptophagidae (Coleoptera) from the Himalayas,
with Descriptions of New Species,
Keys and Remarks to some Languriidae”)

By Georgy Lyubarsky, Moscow

With 23 figures

Summary

Species of the Coleoptera family Cryptophagidae (Cryptophagus pedarius sp. n., Cryp-
tophagus honoratus sp. n., Himascelis nepalensis sp. n., Himascelis gracilis sp. n., Himascelis
turgidus sp. n., Atomaria accola sp. n., and Atomaria mehtita sp. n.) are described from Ne-
pal. The males ‘of Cryptophagus malaisei Bruce 1945, Cryptophagus anxiosus Grouvelle 1916,
Loberus sinuaticollis Bruce 1945, Atomaria torrida Johnson 1970 and the female of Atomaria
plecta Lyubarsky 1996 are described and illustrated for the first time. Several species of Cryp-
tophagidae as well as 2 species of Languriidae are recorded for the first time from Nepal. Iden-
tification keys for the species of the genera Cryptophagus, Himascelis and Atomaria from the
Himalayan Region are provided.

Zusammentassung

Arten der Käferfamilie Cryptophagidae (Cryptophagus pedarius sp. n., Cryptophagus ho-
noratus sp. n., Himascelis nepalensis sp. n., Himascelis gracilis sp. n., H imascelis turgidus sp.
n., Atomaria accola sp. n. und Atomaria mentita sp. n.) werden aus "Nepal beschrieben. Die
Männchen von Cryptophagus malaisei Bruce 1945, Cryptophagus anxiosus Grouvelle 1916,
Loberus sinuaticollis Bruce 1945, Atomaria torrida Johnson 1970 und das Weibchen von Ato-
maria plecta Lyubarsky 1996 werden erstmalig beschrieben und abgebildet. Zahlreiche Arten
der Cryptophagidae ebenso wie 2 Arten der Languriidae werden erstmalig aus Nepal gemel-
det. Bestimmungsschlüssel für die Arten der Gattungen Cryptophagus, Himascelis.and Ato-
marıa aus dem Himalaya werden beigefügt.

APR 1 2 2000

| \

*) Results of the Himalaya Expeditions of J. MARTENS, no. 4 see: Boll. Mus.
reg. Sci. nat. Torino, 1999.- J. M. sponsored by Deutscher dei Austauschdienst
and Deutsche Forschungsgemeinschaft.

2 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

Contents

TS Mia trOG UGE Oia rs RN pete Pave IE 2
De Raxonomiepar ee A ee er ee eae 2
Dihamilys@ry ptophacidacu snc vey rs ee ip cre e eee 2
DEN PAntherophagusDejeanals2lu ar rene terra er eee 2
Dion @nyptophagusiklerbse 79020 Dr en ee 3
2.1.3. Key to the species of Cryptophagus from the Indian subcontinent ...... 9
ZA Hlenoticusallhomson 1868 cn eer ci ae eee 12
2.1.52. Mierambeilihomsen 1863. Lu. oo. ee ce er oh ae eee 12
2.156. Himascelis Sem Gupta l9/8 m... 20.0... oc. ee eee eee 12
2.17. Key to the Indian and Chinese species of Hımascehs 2.2... 2. 16
2.1.8, ‚GaenoseehssIhomson 18690 | Veh Jura. man. ce ae 17
211,92 Artomana Stephens 1330 Ba m ee oa eee 17

2.1.10. Key to Atomaria species from Nepal, based on JOHNSON (1970)
andıtherabove new tndings tom. 2... oe ee 23
Dale @urelins Casey. 1900 8. 2 0 es cae Seem eso ko 25
DA 2 VEphistemusiStephens 1829... u... ae ys ae ae os. ee 25
22 RamiulysWaneuridae wane <b tere ae esac. ono) asa ae 25
22242 Gryptophılus»Reitter 1 874i. ysis vue ok es an os ee 25
22.2. Ch overusisecomee 1861: er. g:s..crcde Se he eine pla ee ee 26
BMACKHOWled ements” ur. et error ee ny Shes ee 26
AR eferemces ee ee Rane A Winn Rien ae 26

1. Introduction

Both Central Asia and the Indian subcontinent represent one of the globe’s cen-
tres of generic and specific diversification of the Cryptophagidae. Numerous new
species have lately been described from there (JOHNSON 1970, LyuBARsKy 1997),
mainly in the genera Cryptophagus Herbst 1792, Micrambe Thomson 1863, Henoti-
cus Thomson 1868 and Atomaria Stephens 1830. Also a new genus from Nepal has
been described recently (LyuBaRsky 1998). In spite of this, the fauna of this region
is surely insufficiently known yet, though its general resemblance to that of central
and southern China can be suggested (LyUBARSKy 1996).

The present contribution is chiefly based on abundant material collected during joint expe-
ditions by JOCHEN MARTENS (Zoological Institute, University Mainz) & WOLFGANG SCHA-
WALLER (Staatliches Museum für Naturkunde Stuttgart, SMNS) mostly in Nepal but also in
the Indian Kashmir. The bulk of the specimens have been collected in different montane and
subalpine forest types by sifting litter and rotten wood and by using the extracting methods
“Berlese” and “Winkler”. The treated 36 Himalayan species of Cryptophagidae cover the al-
titudinal belt 1000-4000 m, the 2 species of Languriidae originate from the belt 950-1330 m.
The material including all holotypes is deposited in the SMNS, with only a few samples re-
tained for the collection of the Zoological Museum of Moscow University (ZMUM), Russia,
as indicated below.

2. Taxonomic part

2.1. Family Cryptophagidae
2.1.1. Antherophagus Dejean 1821

The Indian fauna of Antherophagus contains 2 species: himalaicus Champion
1922 and nigricollis Champion 1922 (Lyusarsky 1991).

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 3

Antherophagus himalaicus Champion 1922

Material: Nepal, Myagdi Distr., Myagdi Khola S Dobang, 2000-2400 m, 21. V. 1995 leg.
MARTENS & SCHAWALLER (1 ex. SMNS); - India, Ladakh, Kargil, 2950 m, 30. V.-7. VI. 1976
leg. MARTENS & SCHAWALLER (4 ex. SMNS, 3 ex. ZMUM); - Lahul, Keylong, 4000 m, 25. VII.
1989 leg. RıeDEL (1 ex. SMNS).

Distribution: India (Uttar Pradesh, Himachal Pradesh, Kumaon, Almora).
Some new Indian records of Antherophagus himalaicus are presented elsewhere
(Lyuparsky 1997). This is the first formal record of this species in Nepal.

Antherophagus sp.
Material: India, Lahul, Keylong, 3500-4000 m, 25. VI. 1989 leg. Rıeper (1 2 SMNS).

Remarks: This female is similar to Antherophagus himalaicus but it is smaller:
3.4mm (body length of himalaicus 4-5 mm), darker: dark red-brown (light yellow-
brown in himalaicus), the pronotum weakly narrowed basally. Perhaps this specı-
men represents a still undescribed species.

2.1.2. Cryptophagus Herbst 1792

The fauna of the Indian subcontinent contains 12 species of Cryptophagus:
anxiosus Grouvelle 1916, atratus Champion 1922, auropubens Grouvelle 1916, ben-
galensis Sen Gupta 1980, clavator Champion 1924, heteroclitus Lyuparsky 1997,
himalaicus Bruce 1952, johnsoni Sen Gupta 1980, lomus Sen Gupta 1980, martensi
Sen Gupta 1980, parallelicollis Grouvelle 1916, and simulator Grouvelle 1916. All
these species are distributed almost exclusively within the Indian subcontinent. In
addition, Cryptophagus aurovestitus Bruce 1945, and affinis Sturm 1845 (= laticollis
Lucas 1849, Reska 1994) have been recorded in India (LyuBARsKy 1997).

Cryptophagus anxiosus Grouvelle 1916 (Fig. 1)

Material: Nepal, Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16.-20.
IV. 1988 leg. MARTENS & SCHAWALLER (1 2 SMNS, 1 2 ZMUM); - Lalitpur Distr., Phulcho-
ki Mt., 1800-2000 m, 25. IV. 1995 leg. MARTENS & SCHAWALLER (1 2 SMNS); — Taplejung
Distr., Yamputhin, cultivated land, open forest, 1650-1800 m, 26. IV.—1. V. 1988 leg. MARTENS
& SCHAWALLER (1 2 SMNS);— Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest re-
mains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (1 ex. ZMUM); - Dailekh Distr., N Dai-
lekh, 1600 m, 1.-2. VI. 1998 leg. ScHAWALLER (10 ex. SMNS); — Dailekh Distr., Dailekh to
Mabuchin Pass, 2300m, 3.-4. VI. 1998 leg. SCHAWALLER (1 ex. SMNS); — Surkhet Distr.,
1600-2000 m, 28. V. 1998 leg. SCHAWALLER (3 ex. SMNS).

Remarks: Cryptophagus anxiosus is similar to heteroclitus, but differs in being
monochromous and possessing a particular structure of the aedeagus. The aedeagus
of heteroclitus is a little longer, and the parameres are much longer than in anxzosus.
Cryptophagus anxiosus is also similar to callosipennis Grouvelle 1919, but differs by
the shape of the callosity which is with an acute tooth on top in callosipennis; in ad-
dition, callosipennis is usually either with a clear black band on the elytra or with
nearly completely black elytra, while anxiosus is usually monochromous, red-
brown, and only some specimens are with a very poorly-expressed, diffused, dark
spot on each side of the elytron. Cryptophagus zonatus Lyubarsky 1996, which 1s
known from China, is likewise bichromous, but it is larger in size, flatter, and the

4 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

callosity is acuminate at the tip (with an obtuse tooth in anxiosus). Aedeagus as in
iceule
Distribution: Nepal, N India (Darjeeling).

Cryptophagus auropubens Grouvelle 1916

Material: Nepal, Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest remains,
9-10. IV. 1988 leg. MARTENS & SCHAWALLER (1 2 SMNS).

Distribution: India, Nepal. This is the first formal record of this species in Ne-
pal.

Cryptophagus aurovestitus Bruce 1945

Material: Nepal, Myagdi Distr., Myagdi Khola N Dobang, 2800-3100 m, 22.-24. V. 1995,
leg. MARTENS & SCHAWALLER (1 2 SMNS).

Distribution: Nepal, India, Burma. This is the first formal record of this spe-
cies in Nepal.

Cryptophagus atratus Champion 1922

Material: Nepal, Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16-20.
IV. 1988, leg. MARTENS & SCHAWALLER (1 d SMNS).

Distribution: Nepal, India. This is the first formal record of this species in
Nepal.

Cryptophagus heteroclitus Lyubarsky 1997

Material: Nepal, Mustang Distr., right bank of Lethe Khola near Lethe, 2400 m, 5.-7. V.
1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS); - S Lethe, 2450-2600 m, 30. IV.-1. V. 1980
leg. MarTENs & Ausogsky (1 ex. SMNS); - Kaski Distr., above Pothana, 2000 m, 27.-29. IV.
1995 leg. MARTENS & SCHAWALLER (2 ex. SMNS, 3 ex. ZMUM); - Kaski Distr., above Dhum-
pus, 2100 m, Sarauja forest, 8.-10. V. 1980 leg. MartENs & Ausossky (1 2 SMNS); - Gorkha
Distr., Chuling Khola, Quercus semecarpifolia forest, 2800 m, 2.-3. VIII. 1983 leg. MARTENS
& SCHAWALLER (1 $ SMNS); — Panchthar Distr., Paniporua, mixed broadleaved forest,
2300 m, 16.-20. IV. 1988 leg. MARTENS & SCHAWALLER (2 ex. SMNS, 1 ex. ZMUM); - Panch-
thar Distr., Dhorpar Kharka, mature Rhododendron-Lithocarpus forest, 2700 m, 13.-16. IV.
1988 leg. MARTENS & SCHAWALLER (1 2 SMNS); — Lalitpur Distr., Phulchoki Mt.,
1800-2000 m, 25. IV. 1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS); - Sankhua Sabha
Distr., Arun Valley, Chichila, 1900-2000m, Quercus forest, bushes near village, 18.-20. VI.
1988 leg. MARTENS & SCHAWALLER (1 2 SMNS); - Taplejung Distr., Yamputhin, cultivated
land, open forest, 1800 m, 26. IV.-1. V. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS); —
Taplejung Distr., SE Yamputhin to Yamputhin, 1650-2000 m, forest mainly Alnus, 26.-30. IV.
1988 leg. MARTENS & SCHAWALLER (1 $ SMNS); — Taplejung Distr., Hellok in Tamur Valley,
2000 m, forest remains, bushes, 17. V. 1988 leg. MARTENS & SCHAWALLER (1 6 SMNS); - Tap-
lejung Distr., Omje Kharka NW Yamputhin, mature mixed braodleaved forest, 2300-2500 m,
1.-6. V. 1988 leg. MARTENS & SCHAWALLER (1 2 SMNS); — Ilam Distr., Mai Pokhari,
2100-2200 m, Castanopsis forest remains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (4 ex.
SMNS, 1 ex. ZMUM); — Ham Distr., Gitang Khola Valley, Alnus forest along river, 1750 m,
11.-13. IV. 1988 leg. MARTENS & SCHAWALLER (3 ex. SMNS, 1 ex. ZMUM); - Solukhumbu
Distr., below Pangum, 2500 m, 14.-15. V. 1997 leg. ScHAWALLER (1 ex SMNS); - Solukhumbu
Distr., Goyom above Sete, 3100m, 10. V. 1997 leg. SCHAWALLER (1 ex. SMNS); — Dailekh
Distr., Dailekh to Mabuchin Pass, 2300 m, 2.-4. VI. 1998 leg. ScHAWALLER (5 ex. SMNS); =
Dailekh Distr., N Dailekh, 1600 m, 1.-2. VI. 1998 leg. SCHAwALLER (1 ex. SMNS); - Surkhet
Distr., N Surkhet, 1600-2000 m, 28. V. 1998 leg. SCHAWALLER (2 ex. SMNS).

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 5)

Distribution: India, Nepal. This is the first formal record of this species in Ne-
pal.

Remarks: In addition to bichromous specimens with a black band on the elytra,
there are completely monochromous individuals which are more or less light brown,
without infuscated elytra. Eyes conical, prominent to varying degrees, but always
small and rather coarsely facetted. Pubescence varying from quite decumbent to
sparse and a little outstanding. Parameres very long, almost 1.5 times as long as api-
cal part of aedeagus, right paramere with 2-3 long setae, left one with 2-4 setae.

Cryptophagus himalaicus Bruce 1952

Material: Nepal, Dolakha Distr., SW Kalinchok Mt., 3100 m, 19.-23. IV. 1995 leg. Mar-
TENS & SCHAWALLER (3 ex. SMNS, 2 ex. ZMUM); — Mustang Distr., Purano Marpha, 3200 m,
9.-11. V. 1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS).

Distribution: India, Nepal.

Cryptophagus honoratus sp. n. (Figs 2-3)

Holotype (d): Nepal, Manang Distr. Marsyandi, 2200 m, above Bagarchap, Acer-Quer-
cus forest, 12-13. IV. 1980 leg. MARTENS & Ausogsky (SMNS).

Paratype: Nepal, Ilam Distr., Mai Pokhari, Gitang Khola Valley, 2500-2600 m, 28.-31.
III. 1980 leg. Martens & Ausogsky (1 2 ZMUM).

Description: Body broadly elongate (Fig. 2), convex; head, prothorax and ely-
tra reddish-brown, elytra slightly darker apically, with almost inclined pubescence.
Head transverse, normal in size, with prominent, hemispherical, somewhat finely fa-
cetted eyes, strongly and densely punctured. Antennae long, somewhat slender, club
well-marked, with half-club reaching beyond base of prothorax, segments 1-3 elon-
gated, with 3rd equal in length to 2nd; 5% slightly elongated, somewhat longer than
4th; 6th to 8th almost equal in length, subquadrate; 9t and 10‘ strongly transverse,
10th twice as broad as long; segments 9-11 equal in width. Prothorax distinctly
transverse, barely (0.7) broader than long, strongly and very densely punctured,
slightly narrowed basally, somewhat convex, sides slightly curved, finely bordered,
forming an angle at a minute but distinct lateral tooth situated near midway of later-
al side. Anterior margin of pronotum convex, anterior angle forming a prominent
callosity, latter occupying at most !/4 of side margin, with a small, elongate-oval
patch of bare surface visible from above, with an obtusangular caudolateral corner.
Lateral margin between callosity and lateral tooth straight, between lateral tooth and
posterior angle slightly convex. Basal groove narrow, basal margin with a prominent
medial lobe. Legs of male 5-5-4, of female 5-5-5. Scutellum small, transverse. Ely-
tra long, oval, humeral angles rounded, shoulders a little broader than prothorax at
lateral tooth, 1. 40-1.46 times longer than combined width and 2.6-2.7 times longer
than prothorax, moderately convex, with moderately strongly rounded sides and a
broadly rounded apex; punctation somewhat weaker and sparser than on prothorax.
Wings fully developed. Length 2.3-2.4 mm. Aedeagus as in Fig. 3.

Remarks: This new species belongs to the group which also includes Crypto-
phagus himalaicus Bruce 1952, simulator Grouvelle 1916 and parallelicollis Grou-
velle 1916. However, it differs from parallelicollis as follows: antennae longer, seg-
ment 5 usually oblong; eyes of normal size (small in parallelicollis); callosity forming
a strong concavity at lateral margin of pronotum and occupying !/4 of margin (thick-

6 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

ening of anterior angles only slightly prominent and not projecting on each side of
pronotum, occupying !/s of lateral margin of pronotum in parallelicollis). It differs
from Cryptophagus himalaicus by the following characters: Eyes finely facetted;
pronotum more strongly transverse, 0.6-0.7 times as broad as long; callosity with-
out tip, forming a concavity with lateral margin, patch of bare surface of callosity
visible from above; smaller size; structure of aedeagus different. It differs from Cryp-
tophagus simulator as follows: Callosity large, forming a strong concavity with lat-
eral margin of pronotum and occupying about '/, of lateral margin; club of antenna
more strongly transverse, segments 9 and 10 of antenna transverse, 10‘ strongly
transverse, 2.0-2.3 times as broad as long.

Cryptophagus malaise: Bruce 1945 (Fig. 4)

Material: Nepal, Kaski Distr., above Pothana, 2000 m, 27.-29. IV. 1995 leg. Martens &
SCHAWALLER (1 ex. SMNS); — Kaski Distr., above Dhumpus, Saranja Forest, 2100m, 8.-10. V.
1980 leg. Martens & Ausossky (2 2 2 SMNS); - Gorkha Distr., Chuling Khola, 2800 m,
Quercus semecarpifolia forest, 2.-3. VIII. 1983 leg. MARTENS & SCHAWALLER (1 d SMNS); —
Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16.-20. IV. 1988 leg. MarTENS
& SCHAWALLER (9 ex. SMNS, 4 ex. ZMUM); — Kathmandu Valley, Sheopuri Mts., Quercus
semecarpifolia forest, 2100-2300 m, 25. VI. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS);
— Parbat Distr., Chitre, 2400 m, 4. V. 1980 leg. MarTENs & Ausossky (1 ex. ZMUM); - Par-
bat Distr., between Chitre and Ghandrung, 2500-2600 m, 6. V. 1980 leg. MARTENS & AusoB-
sky (1 ex. SMNS); — Ramechap Distr., Mohabir Khola E Shivalaya, 2500-2600 m, 6.-7. V. 1997
leg. SCHAWALLER (1 ex. SMNS); - Solukhumbu Distr., Sanam 2700-2800 m, 22.-23. V. 1997
leg. SCHAWALLER (2 ex. SMNS); — Dailekh Distr., Dailekh to Mabuchin Pass, 2300 m, 3.—4. VI.
1998 leg. SCHAWALLER (2 ex. SMNS).

Distribution: NE Burma (Kambaiti), Nepal. This is the first formal record of
the species in Nepal.

Remarks: This species has been described from a single female holotype. The
new samples at hand allow both to elucidate variation and considerably alter/com-
plement the diagnosis of the apparently highly variable malaisez.

Description: Body 1.4-1.9mm long, usually monochromous dark brown,
rarely light brown, yet sometimes head and pronotum slightly darker than elytra.
Pubescence varying strongly from simple and loose, with some or all hairs slightly
outstanding to occasionally double, with both inclined and outstanding hairs
present. Eyes small, coarsely facetted, usually considerably prominent, conical. An-
tennae rather slender, thin, longest segment of flagellum usually second, sometimes
third; segments 4-8 slightly oblong, sometimes subquadrate to even slightly trans-
verse; club of antenna from rather weakly to strongly transverse, 10! segment
strongly transverse. Pronotum transverse, 1.4-1.6 times as broad as long. Lateral
margin of pronotum between callosity and lateral tooth straight or concave. Lateral
tooth small, situated at or in front of middle of lateral margin. Callosity small, weak-
ly thickened, flat, occupying !/7 of lateral margin, usually terminating in a small den-
ticle, but occasionally without it, forming an obtuse angle with lateral margin. Basal
margin of pronotum slightly rounded, either without or with weak medial lobe.
Protarsi of male thickened. Humeral angles of elytra regularly rounded, shoulders
with or without weak tooth. Different stages of wing reduction are observed: usual-
ly these are completely reduced, but there are specimens with weakly to even almost
fully developed wings. Aedeagus as in Fig. 4; it has never been described yet; with
characteristic apical lobes lacking teeth but possessing long, slender parameres (like
in heteroclitus), right paramere with three, left one with two long setae.

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS

Oo

LE

4

Figs 1-7. 1. Cryptophagus anxiosus, male genitalia; — 2. C. honoratus sp. n., dorsal view; —
3. C. honoratus sp. n., male genitalia; — 4. C. malaisei, male genitalia; — 5. C. pe-

darius sp. n., dorsal view; — 6. C. pedarius sp. n., male genitalia; — 7. Loberus s1-
nuaticollis, aedaegus and paramere.

8 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

Cryptophagus pedarius sp. n. (Figs 5-6)

Holotype (8): Nepal, Ilam Distr., N Mai Pokhari, Gitang Khola Valley, 2500-2600 m,
28.-31. III. 1980 leg. Martens & Ausogsky (SMNS).

Paratypes: Same locality and date as holotype (1 2 SMNS, 1 2 ZMUM); — Nepal, Gor-
kha Distr., Chuling Khola, Meme Kharka, 3300-3400 m, 5.-6. VIII. 1983 leg. MARTENS &
SCHAWALLER (1 d SMNS); — Sankhua Sabha Distr., Thudam, mixed forest mainly Betu-
la/Rhododendron, 3550-3650 m, 25.-27. V. 1988 leg. MARTENS & SCHAWALLER (1 d SMNS,
1 6 ZMUM).

Description: Body braodly elongate (Fig. 5), convex; head and prothorax dark
brown, elytra light brown, latter with almost inclined white, sometimes slightly
loose pubescence. Head transverse, normal in size, with small, prominent, conical,
somewhat strongly facetted eyes, strongly and densely punctured. Antennae short,
with a poorly delimited club, with half-club reaching beyond base of prothorax; seg-
ments 1-3 slightly elongated, with 3" equal in length to 24; 4th to 8th subquadrate to
transverse, 9th and 10 transverse, latter 1.7-2.0 times as broad as long; segments
9-11 equal in width. Prothorax transverse, moderately (0.60-0.63) broader than
long, strongly and densely punctured, slightly narrowed apically and basally, some-
what convex with slightly angular sides, its sides finely bordered, lateral tooth dis-
tinct, situated at or near midway of lateral side. Anterior margin or pronotum con-
vex, anterior angle forming a flat callosity, latter occupying !/5 of side margin, with a
small, very narrow, elongate-oval patch of bare surface invisible from above, usually
with a pointed caudolateral corner, yet sometimes latter not pointed. Lateral margin
rounded, straight or concave between callosity and lateral tooth, strongly convex or
sinuate between lateral tooth and posterior angle. Basal groove narrow, basal margin
with a slightly prominent medial lobe. Legs of male 5-5-4, of female 5-5-5. Scutel-
lum small, transverse. Elytra short, oval, humeral angles rounded, each with a small
tooth, shoulders a little broader than prothorax at lateral tooth, 1.30-1.35 times
longer than combined width and 2.2-2.4 times longer than prothorax, moderately
convex, with strongly rounded sides and a broadly rounded apex; punctation some-
what weaker and sparser than on prothorax. Wings absent. Length 1.7-2.1 mm. Ae-
deagus as in Fig. 6.

Remarks: This species is very similar to Cryptophagus malaisei, yet it differs in
the following characters: body larger, 1.7-2.1 mm long; coloration bichromous: head
and pronotum dark brown, elytra light brown; pubescence white, simple, decum-
bent, sometimes slightly outstanding; antennae thick, segments 4-8 transverse, club
poorly defined; callosity usually forming a stronger concavity than in malaise, larg-
er, occupying !/s of lateral margin of pronotum; basal margin or pronotum with a
medial lobe; shoulder of elytron with a small tooth; aedeagus with teeth on apical
lobes, each paramere with two setae. The trio heteroclitus, malaisei, and pedarius
sp.n. seems to form an apparently homogenous group of particularly closely related
species.

Cryptophagus affinis Sturm 1845

Material: Nepal, Mustang Distr., Lupra, in Apodemus wardi nest, 29. IX. 1997 leg. VoL-
zır (2 ex. ZMUM); — Mustang Distr., Purano Marpha, 3200 m, 9.-11. V. 1995 leg. MARTENS &
SCHAWALLER (1 d SMNS); — Lalitpur Distr., Phulchoki Mt., 1800-2000 m, 25. IV. 1995 leg.
MARTENS & SCHAWALLER (1 ex. SMNS); — India, Kashmir, Tangmarg, Pir Panjal Mts., 2600 m,
21-25. V. 1976 leg. MARTENS & SCHAWALLER (18 ex. SMNS, 4 ex. ZMUM).

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 9

Distribution: Holarctic, including India and Nepal. This is the first formal re-
port of this species from India.

Remarks: In Central Europe, Cryptophagus affinis has fully developed wings
(RESKA 1994). In Kirghizia, central part of Central Asia, it displays various degrees
of wing reduction: there are completely wingless specimens as well as individuals
with underdeveloped wings. In the Kopetdagh Mts. (Turkmenia), this species is re-
presented by completely or nearly completely winged samples. Nearly all specimens
from India and Nepal appear wingless, except for one individual from Lalitpur Dis-
trict. This pattern is likely to be due to wind force in these regions. In Kashmir as
well as over the Tibetan Plateau, it is known to be very windy. In valleys which cut
through Nepal in a meridional direction, in particular in the Kali Gandaki Valley
(Mustang Distr.), strong winds dominate all days, usually calming down only at
night. So perhaps it is this pattern that leads to wing reduction in affinis in these re-
gions, just as it happens also in malaisei and pedarius.

Cryptophagus robustus Bruce 1959

Material: Nepal, Dolpo, Ringmo, Lake Phoksumdo, 3600-3900 m, 10.-15. VI. 1973 leg.
MARTENS (1 ex. SMNS, 1 ex. ZMUM).

Distribution: Nepal, Afghanistan. This is the first formal record of the species
in Nepal.

Remarks: Differs from the original description (BRUCE 1959) as follows: wings
completely reduced; eyes small, conical; elytra covered with inclined pubescence;
body length 2.4-2.9 mm; left paramere with three setae, right with two large setae
and two small hairs on top.

Cryptophagus simulator Grouvelle 1916

Material: Nepal, Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16.-20.
IV. 1988 leg MARTENS & SCHAWALLER (1 ex. ZMUM); - Taplejung Distr., upper Simbua
Khola Valley, near Tseram, 3250-3350 m, mature Abies-Rhododendron forest, 10.-15. V. 1988
leg. MARTENS & SCHAWALLER (1 ex. SMNS); - Dailekh Distr., Dailekh to Mabuchin Pass,
2300 m, 3.—4. VI. 1998 leg. SCHAWALLER (1 ex. SMNS).

Distribution: Nepal, India. This is the first formal report of the species from
Nepal.

2.1.3. Key to the species of Cryptophagus from the Indian subcontinent

1 Club of antenna large, 11t segment of the same length as or even longer than I and 10th
combined; pubescence decumbent; callosity forming an obtuse angle with lateral margin,
without pointed caudolateral corner, patch of bare surface on callosity visible from above;
pronotum 0.7-0.8 times as long as broad; lateral tooth situated in middle of lateral margin
REMOTE. Eee ee 2

- Club normal, 11! segment of antenna shorter than 9!" and 10% combined ........... 3

2 Body flatter. Antennae thick, segments 9 and 10 strongly transverse, 10° much wider than
9th, while 11th unusually large, oval, narrowed from middle on, longer than segments 8-10
combined. Callosity not protruding beyond lateral margin. Body length 3mm. India

NE SE ran Br ER REE ra AS en RISE hen ore CNC Soke clavator

— More convex. Antennae slender, club large: segments 9 and 10 transverse, 9! more weak-

10

STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

ly, 10 more strongly so, 10: almost twice as wide as 9°", while 11% wide and as long as 9%
and 10t combined. Callosity projecting beyond margin, occupying about !/4 of lateral
margin. Sides of pronotum slightly curved, lateral margin straight between lateral tooth
and basal angle, slightly concave between callosity and tooth. Base of pronotum with in-
distinct longitudinal wrinkles. Body length 2.7 mm. Aedeagus as depicted elsewhere (Lyu-

BARS Kevall SITES) AN epalaindıayBurmaseer ee Deere aurovestitus
Monochromous, dark, black of blackish-brown. Pronotum moderetely transverse. Eyes
normals hemisphericalwtinely tacetted rn re ee ae 4
Lighter, red or brown, sometimes light with dark spots on elytra ............-.-0-5 5

Antennae long, slender, all joints of flagellum oblong, club moderately transverse, seg-
ments 9 and 10 1.33 times as broad as long. Eyes normal, finely facetted, hemispherical.
Pronotum slightly transverse, 0.7—0.8 times as long as broad. Lateral margin of pronotum
slightly angulated, almost parallel. Callosity small, occupying !/5 of lateral margin, acumi-
nate. Lateral tooth of pronotum small, situated at or in front of middle of lateral margin.
Body oblong, narrow, with simple inclined pubescence. Body length 1.8-2.2 mm. Aedea-
gus as depicted elsewhere (LYUBARSKY 1997: Fig; 2) India nee atratus
Antennae less slender, segments 4, 7 and 8 subquadrate, club well-marked, 9: and 10%
segments distinctly transverse, 1.6 and 1.7 times as wide as long, respectively. Sides of pro-
notum angulated, callosity large, occupying !/4 of lateral margin, slightly prominent, with
a wide patch of bare surface, ending up in an obtuse angle, non-acuminate, more or less
smoothly turning into lateral margin of pronotum. Lateral tooth in middle of lateral mar-
eine B odyalengeh2amimney Nepaly Indiawer me erw ee auropubens

Eyes small, conical, slightly prominent. Lateral tooth small (occasionally absent), situated
at or in front of middle of lateral margin of pronotum. Pronotum slightly transverse, 0.7
timesiassloneas broad: 2. es ee a EEE SIE eee 6
Eyes ofmorimalisize, hemispherical) aa. 2. 2. 2 N eee 10

Eyes finely facetted. Body stout, wide, elytra 2.1 times longer than pronotum, pubescence
inclined. Antennae long, all segments of flagellum oblong, 3'4 longest, club moderately
transverse, Ih and 10th segments 1.3 times as broad as long. Pronotum moderately trans-
verse, 0.7-0.8 long as broad, strongly narrowed basally. Callosity strongly prominent,
forming an acute angle with lateral margin, with or without tip. Patch of bare surface of
callosity invisible from above. Lateral margin of pronotum strongly angulated, with a
small lateral tooth in middle. Paramere as depicted elsewhere (BRUCE 1959: Fig. 1D). Body
lensch 24 3:9 mmy Nepales Attelianiscanl sen ee ees eis ener eee robustus
Eyes coarsely facetted. Pubescence loose, outstanding, sometimes double, i.e. in addition
to decumbent pubescence, outstanding hairs present as well. Body length less than
2.4mm. Antennae shorter, segments 4-8 frequently transverse, 3™4 shorter than 1s‘ and fre-
quently not. longersthan sh nn capt ea beeen oes eck cone eee 7

Wings fully developed. Coloration often bichromous, elytra with a black band, sometimes
reduced to two vague spots. Callosity a little larger, occupying !/4-V/s of lateral margin of
pronotum, usuallysnon=acumimate, vn an deem eae eee eo CCE 8
Wings usually absent or reduced. Elytra uniform light to dark brown, with neither bands
nor spots. Pubescence often outstanding. Callosity a little less, occupying !/7—/s of later-
al margin of pronotum and usually ending up as pointed ..............++2e+eeeee 9

Callosity rather large, moderately prominent, occupying approximately !/, of lateral mar-
gins, terminating in an obtuse angle without tooth. Antennae short, segments of flagellum
slightly elongated or subquadrate, 5‘ not longer than 6t*. Club well-defined, 9: and 10%
segments strongly transverse, each 1.5 times as broad as long. Pronotum narrowed basal-
ly, strongly transverse, lateral tooth small, situated in front of middle of lateral margin.
Body red, head and pronotum darker, red-brown, elytra ligther, yellow-brown, occasion-
ally vague dark spots on sides of elytra. Body length 2.2-2.4 mm. Parameres rather short,
shorter than aedeagus. Aedeagus as in Fig. 1. Nepal, India .................. anxiosus
Elytra usually with a black band, latter sometimes splitting into two spots near external
edge of each elytron, sometimes completely vanishing. Callosity occupying !/5 of lateral
margin, forming a weak obtuse angle with lateral margin, sometimes with small tooth. An-

10

11

12

13

14

15

16

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS iil

tennae rather short, joints of flagellum a little elongated or subquadrate. Club moderately
transverse, not well-defined, 9 segment very slightly transverse, 10th moderately trans-
verse, 1.5 times wider than 9". Body length 1.9-2.2 mm. Parameres very long, longer than
aedeagus, latter as depicted elsewhere (Lyusarsky 1997: Fig. 6). Nepal, India heteroclitus

Body shortly oval, monochromous, red-brown, sometimes pronotum a little darker, dark
brown, and elytra light brown. Pronotum widest in middle, together with callosities nar-
rower than at lateral tooth, slightly narrowed from lateral tooth to basal angles, seeming-
ly expanded in basal half (sometimes sides almost straight). Antennae thin, slender, seg-
ment 9 and 10 rather transverse. Callosity narrow, with or without a narrow patch of bare
surface, occupying !/7 of lateral margin of pronotum, usually with a small tooth, occasion-
ally without tooth. Basal margin of pronotum rounded, with or without slight medial
lobe. Punctation of pronotum and elytra very dense. Aedeagus as in Fig. 4. Body length
ike IEomnrNepaleBunmia Ara. tyres ive ial eezaen allen, ner malaisei
Body bichromous: head and pronotum dark brown, elytra light brown. Pubescence
white, simple, inclined, sometimes slightly outstanding. Antennae thick, segments 4-8
transverse, club not well-defined, weakly transverse. Callosity usually more strongly
prominent than in malaisei, larger, occupying !/s of lateral margin or pronotum. Basal
margin of pronotum with a medial lobe. Each shoulder with a small tooth. Aedeagus as
in Fig. 6: male genitalia differing from those of malaisei by teeth on apical lobes of ae-
deagus and two setae both on right and left paramere. Body length 1.7-2.1 mm. Nepal

EEE tAdTRt 2 Cath hs ER A N Re et pedarius sp. n.

Pubescence double. Callosity small, slightly prominent, not or weakly concave at lateral
margin of pronotum, with a small tooth. Lateral tooth small ..................-.. 11
Pubescence simple, all hairs inclined. Callosity usually larger, more strongly concave at lat-
SEAM AK MOM PLOMOEMMINNG Bee 15

Eyes rather finely facetted. Antennae slender, segments 9 and 10 moderately transverse.
Lateral margin of pronotum slightly rounded, more strongly narrowed basally than api-
cally, lateral tooth at or in front of middle of lateral margin. Wings frequently reduced.

Bodyslenechi117- > dumm lOlarctic foe yd seis es ess on og es as ee oes affinis
Eyes more coarsely facetted. Wings fully developed. Size over 2.5mm ................ 12
Lateral tooth situated approximately in middle of lateral margin of pronotum ...... 13
Lateral tooth situated a little in front of middle of lateral margin of pronotum. Pronotum
cistinetiystransverse. Elytraamonochromous, red... 14
Lateral margin between lateral tooth and basal angle of pronotum slightly concave. Pro-

notum slightly transverse, subquadrate. Elytra infuscate around scutellum. Body length
ZAM ARS UENO, sister NRE Bi Sere nae Semen ranean We ap. ce Anam bY Sa A < lomus
Pronotum distinctly transverse. Lateral margin between lateral tooth and basal angle of
pronotum straight or slightly convex. Elytra monochromous, red. Body length 2.8 mm.
ING D6 See a Se ne Ue J nun nO he ee jobnsoni

Lateral margin between lateral tooth and basal angle of pronotum slightly concave. Body
entire mmYIndawa en re a nn Sen RENO dee bengalensis
Lateral margin between lateral tooth and basal angle of pronotum straight or slightly con-
exes o chygleng tayeoimana Nepal ans. heist ee ee: martensi

Callosity large, forming a strong concavity with lateral margin of pronotum, occupying
about !/, of lateral margin. Club of antenna strongly transverse, segments 9 and 10 trans-
verse, 10‘ strongly transverse, 2.0-2.3 times as broad as lose. 2 Sue 16
Callosity small, forming a weak concavity with lateral margin of pronotum, occupying
1/5-1/4 of lateral margin. Pronotum usually strongly transverse, 0.6-0.7 times as long as

ORO AGERE VE ee ieh, Hh che os Ml eae A iO a tow aly 17

Eyes coarsely facetted, hemispherical. Pronotum slightly transverse, 0.65-0.85 times as
long as broad, strongly narrowed basally. Callosity large, forming a strong concavity with
lateral margin, with a tooth and a well-expressed patch of bare surface. Lateral tooth
usually at or, sometimes, in front of middle of lateral margin. Body length 2.1-3.0 mm.
Aedeagus as depicted elsewhere (BRUCE 1952: pl. XI, IIIA). India ......... himalaicus

12 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

- Eyes finely facetted. Pronotum strongly transverse, 0.6-0.7 times as long as broad. Cal-
losity without tooth, forming a strong concavity with lateral margin, patch of bare sur-
face visible from above. Aedeagus as in Fig. 3. Body stout, length 2.3-2.4mm. Nepal

ee EEE ee See AS neice ene ia SEE Vi ara I honoratus sp. n.

17 Callosity rather small, taking up !/5 of lateral margin, without tooth. Club of antenna
well-defined, segment 10 about 2 times as broad as long. Pronotum parallel-sided, lateral
margin straight, slightly narrowed basally. Lateral tooth small, situated at or in front of
middle of lateral margin. Lateral margin between lateral tooth and basal angles slightly
Concavesbodyslenethi22—2-/ nanny cae were see ieee parallelicollis

— Callosity large, occupying !/4 of lateral margins, forming a concavity with lateral margin
of pronotum, with a small tooth. Club of antenna slightly transverse, segment 10 about 1.5
times as broad as long. Pronotum strongly narrowed basally, its lateral margin rounded,
with a tooth in middle, convex between lateral tooth and basal angle. Body length
2.2-2.9mm. Aedeagus as depicted elsewhere (LyuBarsky 1997: Fig. 4). India . simulator.

2.1.4. Henoticus Thomson 1868

Henoticus serratus nepalensis Johnson 1975

Material: Nepal, Dolakha Distr., SW Kalinchok Mt., 3100 m, 19.-23. IV. 1995 leg. Mar-
TENS & SCHAWALLER (1 d SMNS).

Distribution: Nepal, India.

Henoticus indicus Grouvelle 1916
Material: Nepal, Kathmandu Valley, Phulchoki Mt., 2600-2650 m, 21.-22. III. 1980 leg.
MARTENS & Ausogsky (1 2 SMNS).
Distribution: Nepal, India. This is the first formal record of the species in Ne-
pal.
Henoticus regificus Johnson 1975

Material: Jumla Distr., Khali Lagna Pass, 3500 m, 16.-17. VI. 1998 leg. SCHAWALLER (6 ex.
SMNS, 3 ex. ZMUM); - Dailekh Distr., Dailekh to Mabuchin Pass, 2300 m, 3.-4. VI. 1998 leg.
SCHAWALLER (2 ex. SMNS).

Distribution: Nepal.

2.1.5. Micrambe Thomson 1863

Micrambe sp.

Material: Nepal, Manang Distr., Marsyandi, above Bagarchap, 2200 m, Acer-Quercus, IV.
1980 leg. Martens & Ausossky (1 ex. SMNS); - Gorkha Distr., Chuling Khola, 2800 m,
Quercus semecarpifolia forest, 2-3. VIII. 1983 leg. MARTENS & SCHAWALLER (2 ex. SMNS).

2.1.6. Himascelis Sen Gupta 1978

The genus Himascelis is currently known to contain two species: brunneus Sen
Gupta 1978, distributed in India (West Bengal, Himachal Pradesh, Kashmir), Nepal
and Bhutan, and kashmirensis Sen Gupta & Pal 1980, from India (Kashmir) (SEN
Gupta 1978; SEN Gupta & Pat 1980). Regrettably, I have failed to receive their
types for re-examination.

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 13

13 45

Figs 8-15. 8. Himascelis chinensis, head and pronotum; - 9. H. similis, head and pronotum;
— 10. H. nepalensis sp. n., head and pronotum; — 11. H. nepalensis sp. n., aedea-
gus; - 12. H. gracilis sp. n., head and pronotum; - 13. H. gracillis sp. n., aedeagus;
— 14. H. turgidus sp. n., head and pronotum; — 15. H. turgidus sp. n.; aedeagus.

14 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

Recently, Nikıtsky (1996) has described two Chinese species of Sternodea Reitter
1875: chinensis Nikitsky 1996 and similis Nikitsky 1996. The types of these species
have been revised and, as a result, both Sternodea chinensis and Sternodea similis ap-
pear to actually belong to Himascelis.

In addition, three new congeners have been found in Nepal, with their descrip-
tions and a general key to species of Himascelis presented in this paper.

An overview of the known species belonging to Himascelis shows this genus to be
indeed closely related to Sternodea. Yet the new species described herein fail to fit
exactly in the modern generic diagnoses as given by LEsCHEN (1996). For example,
the longitudinal mesosternal line is equally weakly developed in both genera. In ad-
dition, in both genera the transverse line of the pronotum is present in some species
but absent in some others. Despite this, the antennal grooves present in Himascelis
make it readily distinguishable from Sternodea.

Himascelis chinensis (Nikitsky 1966) comb. n. (Fig. 8)

Remarks: The holotype in ZMUM has been re-examined. As a result, the spe-
cies proves to be actually referred to Himascelis, because the holotype displays well-
developed antennal grooves and all other characters of this genus. Pronotum with a
transverse line (Fig. 8). The original description (Nixkirsky 1996) is inaccurate as ın
fact this species differs from congeners in having no boss on the frons.

Distribution: China (Shaanxi).

Himascelis similis (Nikitsky 1996) comb. n. (Fig. 9)

Remarks: The holotype and paratype, both in ZMUM, have been restudied. As
a result, this species is likewise referred to Himascelis, because the antennal grooves
are well-developed and all other traits are characteristic of this genus. Pronotum
with a transverse line (Fig. 9).

Distribution: China (central Sichuan).

Himascelis nepalensis sp. n. (Figs 10-11)

Holotype (d): Nepal, Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest re-
mains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (SMNS).

Paratypes: Same locality and data as holotype (3 ex. SMNS, 1 ex. ZMUM); - Nepal, Ilam
Distr., Gitang Khola, 2550 m, 28.-31. III. 1980 leg. Martens & Ausogsky (1 ex. SMNS); -
Taplejung Distr., upper Tamur Valley, resthut/side-valley, broadleaved forest with bamboo,
2450 m, 19. V. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS, 1 ex ZMUM); - Panchthar
Distr., Dhorpar Kharka, mature Rhododendron-Lithocarpus forest, 2700 m, 13.-16. IV. 1988
leg. MARTENS & SCHAWALLER (2 ex. SMNS); — Panchthar Distr., Paniporua, 2300 m, mixed
broadleaved forest, 16.-20. IV. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS); — Parbat
Distr., Chitre, 2400 m, 4. V. 1980 leg MARTENs & Ausossky (1 ex. SMNS); - Gorkha Distr.,
Chuling Khola, Djinski Kharka, 3400 m, Abies forest, 4.-5. VIII. 1983 leg. MARTENS &
SCHAWALLER (1 d ZMUM).

Description: Body uniform brown, elliptical-oval, stout, 1.65-1.75 times as
long as wide. Length of body 1.2-1.4mm. Eyes small, strongly prominent laterally,
coarsely facetted. Frons coarsely punctate, with a triangular boss (Fig. 10). Antenna
(Fig. 10) stout, flagellum broader than length of claw joint; basal joint cylindrical,
distinctly longer than thick; second much narrower than basal, cylindrical; third 1.2
times longer than second, slightly curved, club-like; each of 4t to 7th joints some-

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 15

what wider than long; 8!" and 9 transverse, 3 times as wide as long; two apical joints
forming a large club, 10‘ approximately twice as wide as long, apical joint 2 times
longer than previous one. Pronotum subtrapezoidal, approximately twice as wide as
long; lateral sides strongly narrowing in apical half; posterior margin weakly but dis-
tinctly bisinuate; basal corners sharply angulate. Punctation of pronotum coarse and
moderately dense. Transverse line consisting of punctures, weak, undistinguished
(like in Sternodea baudii Reitter 1875 and some specimens of Sternodea raddei Reit-
ter 1876, LyuBarsky 1987), extending to sides of pronotum. Scutellum transverse.
Hind tarsi of male 4-segmented. Elytral base slightly wider than pronotal base; ely-
tral sides arcuate in anterior half; elytral punctation coarse. Pubescence somewhat
outstanding, long; hairs approximately 2 times longer than claw joint. Wings absent.
Aedeagus as in Fig. 11.

Himascelis gracilis sp. n. (Figs 12-13)

Holotype (8): Nepal, Taplejung Distr., upper Tamur Valley, resthut/side-valley, broad-
leaved forest with bamboo, 2450 m, 19. V. 1988 leg. MARTENS & SCHAWALLER (SMNS).

Paratypes: Same locality and data as holotype (1 ex. SMNS); — Nepal, Parbart Distr.,
between Deorali and Chitre, 2700 m, 1.-2. V. 1995 leg. MARTENS & SCHAWALLER (1 ex.
SMNS); - Sankhua Sabha Distr., above Pahakhola, 2600-2800 m, Quercus semecarpifolia-
Rhododendron forest, 31. V.- 3. VI. 1988 leg. MARTENS & SCHAWALLER (3 ex. SMNS, 3 ex.
ZMUM); — Manang Distr., above Bagarchap, Acer-Quercus forest, 2400 m, 13.-14. IV. 1980
leg. MARTENS & Ausogsky (1 ex. SMNS).

Description: Body uniform yellowish-brown, elliptical-oval, elongate,
1.70-1.87 times as long as wide. Length of body 1.2-1.4mm. Eyes small, strongly
prominent laterally, coarsely facetted. Frons coarsely punctate, with a triangular
boss (Fig. 12). Antenna (Fig. 12) slender, flagellum somewhat narrower than length
of claw joint; basal joint cylindrical, distinctly longer than wide; second much nar-
rower than basal, cylindrical; third 1.2 times longer than second, straight; each of 4th
to 7th joints somewhat wider than long; 8!h and 9 transverse, 2.5-3.0 times as wide
as long; two apical joints forming a large club, 10th approximately twice as wide as
long, apical joint 2 times longer than previous one. Pronotum subtrapezoidal, ap-
proximately twice as wide as long; lateral sides strongly narrowing in apical half;
posterior margin weakly but distinctly bisinuate; basal corners sharply angulate.
Punctation of pronotum weak and moderately dense. Transverse line consisting of
punctures, more or less well-developed, extending to sides of pronotum. Scutellum
transverse. Hind tarsi of male 4-segmented. Elytral base slightly wider than prono-
tal base; elytral sides arcuate in anterior half; elytral punctation weak. Pubescence al-
most decumbent, shorter than in previous species; hairs approximately 1.5 times
longer than claw joint. Wings absent. Aedeagus as in Fig. 13.

Himascelis turgidus sp. n. (Figs 14-15)

Holotype (3): Nepal, Panchthar Distr., Dhorpar Kharka, mature Rhododendron-Litho-
carpus forest, 2700 m, 13.-16. IV. 1988 leg. MARTENS & SCHAWALLER (SMNS).

Paratypes: Same locality and data as holotype (1 d SMNS, 1 d ZMUM); - Nepal, Taple-
jung Distr., Omje Kharka NW Yamputhin, mature mixed broadleaved forest, 2300-2500 m,
1.-6. V. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS, 1 ex. ZMUM).

Description: Body uniform brown or dark brown, elliptical-oval, elongate,
1.75-1.85 times as long as wide. Length of body 1.35-1.50 mm. Eyes small, strongly

16 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 598

prominent laterally, coarsely facetted. Frons coarsely punctate, with a triangular
boss (Fig. 14). Antenna (Fig. 14) slender, flagellum somewhat narrower than length
of claw joint; basal joint cylindrical, distinctly longer than wide; second much nar-
rower than basal, cylindrical; third 1.5 times longer than second, slightly curved,
club-like; each of 4* to 7!" joints somewhat wider than long; 8!" and Ih transverse, 3
times as wide as long; two apical joints forming a large club, 10 approximately
twice as wide as long, apical joint 1.5 times longer than previous one. Pronotum sub-
trapezoidal, approximately twice as wide as long; lateral sides strongly narrowing in
apical half; posterior margin weakly but distinctly bisinuate; basal corners sharply
angulate. Punctation of pronotum coarse and moderately dense. Transverse line con-
sisting of punctures, weak, undistinguished, extending to sides of pronotum. Scutel-
lum transverse. Hind tarsi of male 4-segmented. Elytral base slightly wider than pro-
notal base; elytral sides arcuate in anterior half; elytral punctation coarse. Pu-
bescence somewhat outstanding, long; hairs approximately 2 times longer than claw
joint. Wings absent. Aedeagus as in Fig. 15.

2.1.7. Key to the Indian and Chinese species of Himascelis

Remarks: The pairs brunneus/chinensis and kashimirensis/similis are poorly distin-
guished, chiefly because material is too limited (1-2 specimens). In particular, both kashmi-
rensis and chinensis are known only from the female holotypes. The aedeagi of brunneus and
similis are generally very similar.

1 8th and, especially, 9t* joint of antenna strongly transverse (Figs 10, 12, 14), 9t approxi-
mately 3 times as wide as long. Second joint shorter than third one. Transverse line con-
sisting of punctures usually weak and undistinguished = 2.22... ...1.2..22.. 2 oseeeee 2

— 8th and 9% joint of antenna weakly transverse (Figs 8-9), I approximately 1.5 times as
wide as long. Second joint equal in length to third one. Antenna slender, its 3 joint
straight, weakly club-like. Pronotum strongly punctured. Transverse line consisting of
puncturesisometimes well-developed) eae 2 2 ee eae ee ae ee 4

2 Antenna stout (Fig. 10), flagellum somewhat broader than length of claw joint. Third joint
of antenna clearly club-like, slightly curved, 1.25 times longer than second joint. 11 joint
approximately 2 times longer than previous one. Pronotum strongly punctured. Pu-
bescence of elytra somewhat outstanding, longer, approximately 2 times longer than claw
joint. Body brown, stout, 1.65-1.75 times as long as wide. Aedeagus as in Fig. 11. Length
1.2-1.4.mm.; Nepal 2 ee Sunita ener ee nepalensis sp. n.

— Antenna slender, flagellum somewhat narrower than length of claw joint ............ 3

3 Third joint of antenna approximately straigth (Fig. 12), about 1.25 times longer than se-
cond joint. 11!" joint approximately 2 times longer than previous one. Pronotum slightly
punctured. Pubescence of elytra decumbent, shorter, somewhat less than 2 times longer
than claw joint. Body light brown, elongate, 1.70-1.87 times as long as wide. Aedeagus as
inte; 13) ensthsl.2_ 1 AmmyNepalbey ee een rea ae eer gracilis sp. n.

— Third joint of antenna slightly curved (Fig. 14), club-like, 1.5 times longer than second
joint. 11% joint approximately 1.5 times longer than previous one. Pronotum strongly
punctured. Pubescence of elytra outstanding, shorter, approximately 2 times longer than
claw joint. Body brown or dark brown, elongate, 1.75-1.85 times as long as wide. Aedea-
gusidsunvhia-5)) Wengthel:35 1650, mm Nepales rer erik er turgidus sp. n.

4° Fkronsiwith aitriangularjboss (Kies 9) 2.2.2.0 een eee eee eee 5
Frons without triangular boss (Fig. 8)

5 Body yellow-brown to black-brown, elongate, 1.90-1.95 times as long as wide. Third
joint of antenna equal to or slightly longer than any of joints 4-6. Elytra 1.19 as long as
wide. Length 1.33-1.45 mm. India, Nepal, Bhutan .............0.00000000e brunneus

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS WW,

- Body red-brown, elongate, 1.9 times as long as wide. Third joint of antenna as long as
joints 4-6 combined. Elytra 1.22-1.30 times as long as wide. Length 1.3-1.5mm. China
(eentral’Sichvan)sa. nee en me A Saori AM BT aired. Stel Senet one similis

6 Pubescence more or less decumbent. Body red-brown, stout, 2.3 times as long as wide.
11‘ joint of antenna approximately 1.3 times longer than 10‘ joint. Length 1.36 mm. In-
CINE a Se ae cae 0 ae MR Ro es ee Tons estan Bis tole htts, aut doa es oe ea kashmirensis
— Pubescence of elytra more or less outstanding, less than about 2 times longer than claw of
joint. Body red-brown, stout, 1.73 times as long as wide. 11‘ joint of antenna slightly
longer than preceding one. Length 1.3 mm. China (Shaanxi) ................ chinensis.

2.1.8. Caenoscelis Thomson 1863

From Nepal, only a single species has been described: Caenoscelis franzi Sen Gup-
ta & Basak 1985 (SEN Gupta & Basar 1985). Caenoscelis jakesi Johnson 1972 has
been described from Afghanistan (Herat).

Caenoscelis franzi Sen Gupta & Basak 1985 (Fig. 16)

Material: Nepal, Ilam Distr., N Mai Pokhari, Gitang Khola Valley, 2500-2600 m, 28.-31.
III. 1980 leg. MARTEns & Ausossky (1 d ZMUM); - Parbat Distr., between Deorali and
Chitre, 2700 m, 1.-2. V. 1995 leg. MARTENS & SCHAWALLER (1 2 SMNS); - Panchthar Distr.,
Paniporua, 2300 m, mixed broadleaved forest, 16.-20. IV. 1988 leg. MARTENS & SCHAWALLER
(1 2 SMNS).

Distribution: Nepal.

Remarks: This distinctive species is similar as Caenoscelis ferruginea (Sahlberg
1820) and Caenoscelis jakesi on account of the structure of the antennal club. Cae-
noscelis franzi differs from them in the structure of the aedeagus (Fig. 16), the feebly
convex body, the dark coloration and the reduced lateral submarginal line of the pro-
notum.

2.1.9. Atomaria Stephens 1830

A key to the Indian species of Atomaria has been published by JOHNSON (1970).
The fauna of the Indian subcontinent contains 11 species of Atomaria: frugi Lyubar-
sky 1997, incertula Johnson 1970, khumbuensis Johnson 1970, klapperichi Johnson
1970, lewis: Reitter 1877, obliqua Johnson 1970, pudica Johnson 1970, torrida John-
son 1970, tristis Johnson 1970 (all from the subgenus Anchicera Thomson 1863); gra-
cilicornis Reitter 1887, prolixa Erichson 1846 (both from Atomaria s. str.). Six species
have been recorded in Nepal (JOHNSON 1970, 1971): incertula, khumbuensis, lewis,
obliqua, torrida, and gracilicornis.

Atomaria barani Brisout 1863, fasciata Kolenati 1846, peltata Kraatz 1853, pusil-
la (Paykull 1798), munda Erichson 1846, klapperichi, obliqua, lewisi, and gracilicor-
nis have been found in Afghanistan (JOHNSON 1972). From central China, Atomaria
plecta Lyubarsky 1966 and Atomaria angellata Lyubarsky 1996 have been described
(LyuBARSKyY 1996).

Atomaria fasciata Kolenati 1846

Material: N Afghanistan, Badakshan, Dehgul, 3000 m, 15. VII. 1973 leg. Kasakov (1 d
ZMUM); - E Kazakhstan, Dzhungarsky Alatau Mts., Tastau Mt., Jumak, Kokmoin River,

18 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 598

1800 m, Abies forest, litter under stones, 4. VIII. 1991 leg. TiscHECHKIN (1 d ZMUM); - same
data, Kara-Unghur River, under stones, 3546m, 8. VIII. 1991 leg. TiscHECHKIN (1 9
ZMUM); - same data, Sanyk-Tas Mt., Mynteks River, forest, litter under stones, 2100 m, 11.
VIII 1991 leg. TiscHECHKIN (2 ex. ZMUM); - Kirghizia, N Tyup, Shaty area, 30. VII. 1987
leg. KurBaTov (1 ex. ZMUM); - Tajikistan, Hissar Mt. Range, 1900 m, 1. IX. 1975 leg. Yanu-
SHEV (1 ex. coll. YANUSHEV); — Zaalaisky Mt. Range, 28. VII. 1975 leg. YAnUsHEv (1 ex. coll.
YANUSHEV).

Distribution: Mediterranean (S Europe, N Africa), Ukraine (Kharkoy, Cri-
mea), Caucasus (Stavropol Prov., N Ossetia, Georgia, Armenia, Azerbaijan), Kirghi-
zia, E Kazakhstan, Tajikistan, Afghanistan.

Remarks: Although Atomaria fasciata is distributed as far as Central Asia and
Afghanistan in the east, it has not been found yet down to the southern border of the
Palaearctic (Himalayas), hence remaining purely Palaearctic. The specimens from
the Dzhungarsky Alatau Mts. are the easternmost record of this species.

Atomaria lewisi Reitter 1877

Material: Nepal, Dolakha Distr., SW Kalinchok Mt., 3100 m, 19.-23. IV. 1995 leg. Mar-
TENS & SCHAWALLER (2 ex. SMNS, 2 ex. ZMUM); - Kaski Distr., above Pothana, 2000 m,
27-29. IV. 1995 leg. MARTENS & SCHAWALLER (4 ex. SMNS); - Lalitpur Distr., Phulchoki Mt.,
1800-2000 m, 25. IV. 1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS); — Mustang Distr.,
right bank of Lethe Khola near Lethe, 2400 m, 5.-7. V. 1995 leg. MARTENS & SCHAWALLER
(1 ex. SMNS); — Lamjung Distr., Senghe, bridge above Marsyandi, 150m, 10. IV. 1980 leg.
Martens & Ausogsky (1 ex. SMNS); - Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis
forest remains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS); - Ilam Distr., Mai
Pokhari, 2150-2250 m, 23.-25. VIII. 1983 leg. Martens & Daams (1 ex. SMNS); - Ilam
Distr., Mai Pokhari, 2100 m, 31. III.-1. IV. 1980 leg. Martens & Ausopssky (1 ex. SMNS); —
Ilam Distr., between Mai Pokhari and Ilam, 1330 m, 1. IV. 1980 leg. MARTENS & Ausossky
(1 ex. SMNS).

Distribution: Cosmopolitan.

Atomaria incertula Johnson 1970

Material: Nepal, Kaski Distr., above Pothana, 2000 m, 27.-29. IV. 1995 leg. MARTENS &
SCHAWALLER (30 ex. SMNS, 12 ex. ZMUM); - Kaski Distr., above Dhumpus, 2100 m, Sarau-
ja forest, 8.-10. V. 1980 leg. MarTENs & Ausossky (1 ex. SMNS); - Ilam Distr., Mai Pokhari,
cultivated land, 2100-2200 m, 25.-27. III. 1980 leg. Martens & Ausossky (3 ex. SMNS); —
Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest remains, 9.-10. IV. 1988 leg. Mar-
TENS & SCHAWALLER (17 ex. SMNS); — Ilam Distr., Mai Pokhari, 2150-2250 m, 23.-25. VIII.
1983 leg. MARTENS & Daams (1 ex. SMNS); - Ilam Distr., road from Ilam to Birtamode, Kar-
phok, 1700m, cultivated land with bushes, 2. IV. 1980 leg. MarreNs & Ausossky (1 ex.
SMNS); - Lalitpur Distr., Phulchoki Mt., 1800-2000 m, 25. IV. 1995 leg. MARTENS & SCHA-
WALLER (1 ex. SMNS); - Myagdı Distr., Myagdi Khola, Dobang, 2400 m, 25. V. 1995 leg. Mar-
TENS & SCHAWALLER (2 ex. SMNS); - Myagdi Distr., N of Bega Deorali, 2400 m, 16.-17. V.
1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS); — Manang Distr., Marsyandi, 2200 m, above
Bagarchap, Acer-Quercus, 12.-13. IV. 1980 leg. MARTENS & Ausossky (1 ex. SMNS); — Ma-
nang Distr., Marsyandi, 2550m, Thimang-Bagarchap, Tsuga-Acer-Rhododendron forest,
14-17. IV. 1980 leg. Martens & Ausogsky (1 ex. SMNS); — Panchthar Distr., Paniporua,
2300m, mixed broadleaved forest, 16.-20. IV. 1988 leg. MARTENS & SCHAWALLER (2 ex.
SMNS); — Panchthar Distr., between Deorali, Puspati and Sheldoti, 2500-2800 m, Tsuga-
Lithocarpus forest, 28. VII. 1983 leg. MARTENS & Daas (1 ex. SMNS); — Kathmandu Valley,
Nagarjung, Jamacok, 1900-2100m, 18. VIII. 1983 leg. MARTENS & SCHAWALLER (1 ex.
SMNS, 1 ex. ZMUM); - Kathmandu Valley, Baneshwar, cultivated land, gardens, 1400 m, 30.
II1.-2. IV. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS); - Kathmandu Valley, Sheopuri
Mts., Quercus semecarpifolia forest, 2100-2300 m, 25. VI. 1988 leg. MARTENS & SCHAWALLER

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 19

16

21

Figs 16-23. 16. Caenoscelis franzi, aedeagus; — 17. Atomaria torrida, aedeagus; — 18. A. acco-
la sp. n., dorsal view; — 19. A. accola sp. n., aedeagus; — 20. A. accola sp. n., sper-
matheca; — 21. A. mentita sp. n., dorsal view; — 22. A. mentita sp. n., aedeagus; —
23. A. plecta, spermatheca.

(3 ex. SMNS); - Taplejung Distr., Yamputhin, cultivated land, open forest, 1650-1800 m, 26.
IV.-1. V. 1988 leg. MARTENS & SCHAWALLER (2 ex. SMNS, 1 ex. ZMUM); - Dolakha Distr.,
SW Kalinchok Mt., 3100 m, 19.-23. IV. 1995 leg. MARTENS & SCHAWALLER (6 ex. SMNS, 3 ex.
ZMUM); — Mustang Distr., right bank of Lethe Khola near Lethe, 2400 m, 5.-7. V. 1995 leg.
MARTENS & SCHAWALLER (2 ex. SMNS); — Mustang Distr., Thaksang, 3150 m, Pinus excelsa-
Abies forest, 26.-29. IV. 1980 leg. Martens & Ausossky (1 ex. SMNS); - Sankhua Distr.,
Arun Valley, Chichila, 1900-2000 m, Quercus forest, bushes near village, 18.-20. VI. 1988 leg.
MARTENS & SCHAWALLER (5 ex. SMNS, 2 ex. ZMUM); - Bhojpur Distr., Dilkharka, 2100 m,
26. V. 1997 leg. Hauser (1 ex. SMNS); - Solukhumbu Distr., below Pangum, 2500 m, 14.-15.
V. 1997 leg. SCHAWALLER (1 ex. SMNS); - Surkhet Distr., N Surkhet, 1600-2000 m, 28. V. 1998
leg. SCHAWALLER (3 ex. SMNS); — Dailekh Distr., Talpokhari S Dailekh, 1800 m, 29. V. 1998

20 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

leg. SCHAWALLER (1 ex. SMNS); — Dailekh Distr., N Dailekh, 1600m, 1.-2. VI. 1998 leg.
SCHAWALLER (1 ex. SMNS).

Distribution: India, Pakistan, Nepal.

Remarks: A single specimen (label: road from Ilam to Birtamode, Karphok,
1700 m, 2. IV. 1980) has the left antenna with only 10 segments, because the joints 3
and 4 are fused, this combined antennomere is very long, somewhat longer than first
joint.

Atomaria gracilicornis Reitter 1887

Material: Nepal, Dolakha Distr., SW Kalinchok Mt., 3100 m, 19.-23. IV. 1995 leg. Mar-
TENS & SCHAWALLER (3 ex. SMNS, 2 ex. ZMUM); — Mustang Distr., Purano Marpha, 3200 m,
9.-11. V. 1995, leg. MARTENS & SCHAWALLER (1 ex. SMNS).

Distribution: Caucasus, Siberia up to the Far East, India, Nepal.

Atomaria prolixa Erichson 1846

Material: Nepal, Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16.-20.
IV. 1988 leg. MARTENS & SCHAWALLER (1 ex. ZMUM).

Distribution: Palaearctıc.

Atomaria torrida Johnson 1970 (Fig. 17)

Material: Nepal, Ilam Distr., Mai Pokhari, 2150-2250 m, 23.-25. VIII. 1983 leg. MARTENS
& Daans (1 ex. SMNS); - Ilam Distr., Mai Pokharı, 2100-2200 m, Castanopsis forest remains,
9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (6 ex. SMNS, 3 ex. ZMUM); - Panchthar Distr.,
Paniporua, 2300 m, mixed broadleaved forest, 16.-20. IV. 1988, leg. MARTENS & SCHAWALLER
(4 ex. SMNS, 2 ex. ZMUM); - Sankhua Sabha Distr., Arun Valley between Mure and Hurure,
mixed broadleaved forest, 2050-2150 m, 9.-17. VI. 1988 leg. MARTENS & SCHAWALLER (1 ex.
SMNS).

Distribution: India, Nepal.
Remarks: The male of Atomaria torrida is illustrated here for the first time
(reas):

Atomaria obliqua Johnson 1970

Material: Nepal, Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16-20.
IV. 1988 leg. MARTENS & SCHAWALLER (8 2 2 SMNS, 3 22 ZMUM); - Taplejung Distr.,
above Yamputhin, left bank of Kabeli Khola, bushes, open forest, 1800-2000 m, 27.-29. IV.
1988 leg. MARTENS & SCHAWALLER (1 d ZMUM); - Taplejung Distr., Omje Kharka NW
Yamputhin, mature mixed broadleaved forest, 2300-2500 m, 1.-6. V. 1988 leg. MARTENS &
SCHAWALLER (1 2 SMNS); - Taplejung Distr, Yamputhin, ascent to Pass Deorali,
2100-2600 m, cultivated land, bushes, 16. V. 1988 leg. MARTENS & SCHAWALLER (1 2 SMNS);
— [lam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest remains, 9.-10. IV. 1988 leg.
MARTENS & SCHAWALLER (3 2 2 SMNS, 1 d ZMUM); - Ilam Distr., Mai Pokhari, Gitang
Khola Valley, 2500-2600 m, 28.-31. III. 1980 leg. MARTENS & SCHAWALLER (1 & SMNS); —
Tam Distr., Mai Pokhari, cultivated land, 2100-2200 m, 25.-27. III. 1980 leg. MarTENs & Au-
soBsky (1 2 SMNS, 1 d ZMUM); - Kaski Distr., above Pothana, 2000 m, 27.-29. IV. 1995 leg.
MARTENS & SCHAWALLER (4 22 SMNS, 1 2 ZMUM); — Kathmandu Valley, Nagarjung,
Jamacok, 1900-2100 m, secondary forest, 18. VIII. 1983 leg. MARTENS & SCHAWALLER (1 @
SMNS); — Kathmandu Valley, Nagarjung, Jamacok, 1400-1600 m, secondary forest, 18. VII.
1983 leg. MARTENS & SCHAWALLER (1 2 SMNS); - Surkhet Distr., N Surkhet, 1600-2000 m,
28. V. 1998 leg. SCHAWALLER (2 ex. SMNS).

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS

Distribution: India, Nepal, Afghanistan.

Remarks: The Indian subcontinent and China lodge a group of several similar
species of Atomaria with a more or less well-developed dark band on the elytra. Ato-
maria fasciata joins this group which also includes obliqua, plecta and angellata. The
above material from Nepal shows that some differences claimed to exist between an-
gellata from China, and obliqua appear unreliable. Atomaria angellata differs from
obliqua by the more strongly elongated 1st antennal segment, by the more stout club
and by the shape of the apex of the aedeagus. It is noteworthy that usually in the fe-
males of obliqua from Nepal the structure of the club tends to be more similar to
that of angellata. As compared to angellata, on the average obliqua is characterized
by the less transverse pronotum (obliqua: 1.20-1.35, angellata: 1.3-1.4) which is
much more densely punctuate (in obligua, the distance between the punctures is
much less than the puncture diameter; in angellata, the distance between the punc-
tures is about 0.5 of the puncture diameter), and by the somewhat longer elytra (in
oliqua, 1.3-1.4 times longer than the overall width of both elytra; in angellata, this
ratio is 1.2-1.3) which are reddish in obliqua and straw-yellow in angellata. The ely-
tra are somewhat indistinctly infuscate in khumbuensis and frugi. The same group
also includes one more, new species described below.

Atomaria (Anchicera) accola sp. n. (Figs 18-20)

Holotype (d): Nepal, Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis forest remains,
9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (SMNS).

Paratypes: Same locality and date as holotype (8 ex. SMNS, 4 ex. ZMUM); — Nepal,
Panchthar Distr., Paniporua, 2300 m, mixed broadleaved forest, 16.-20. IV. 1988 leg. MARTENS
& SCHAWALLER (1 ex. SMNS).

Description: Body length 1.4-1.7 mm, body covered with long decumbent sil-
very pubescence, hairs somewhat longer than claw joint. Head dark brown, prono-
tum yellow (straw-coloured), elytra lighter, brownish-yellow with an irregular
brownish-black transverse band across the disk, this band forming a V-shape astride
the suture (Fig. 18), similar to that in Atomaria obliqua. Antennae and legs reddish-
yellow, club of antenna slightly infuscate. Antennae long and slender, as in Fig. 18;
all segments elongate, segment 1 longer than 2, 2™4 longer than 3", latter equal to 7*,
4th equal to 6t or 8th. Segment 5 most elongate except 1. Club narrow and not well-
marked, segment 9 slightly elongate, 10 subquadrate. Pronotum transverse,
1.40-1.55 times as broad as long, broadest across the middle, sides strongly curved
and narrowed apically and basally; side borders single, visible from above only in ba-
sal half. Pronotal surface shining, not shagreened, densely covered with deep punc-
tures separated from their nearest neighbours by a half-diameter on the disk; hind
angles obtuse; pronotal disk convex; base of pronotum with a narrow transverse de-
pression; hind margin finely bordered, not or barely produced caudomedially. Ely-
tra strongly arched, broadest about the middle, 2.25-2.60 times as long as pronotum
and 1.15-1.23 times as long as broad combined; surface shining, not shagreened,
densely punctured, the punctures in basal half similar in size to those on pronotal
disk and separated from their lateral neighbours by approximately one diameter on
the average. Wings fully developed. Aedeagus as in Fig. 19. Spermatheca as in Fig. 20.

Remarks: The new species is similar to Atomaria obliqua but differs in size, be-
ing somewhat longer; antennae longer, all segments elongate, segment 7 longer, al-
most as long as 34; head dark (head of obliqua lighter); pronotum more transverse,

DY, STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

punctation more dense, pronotum and elytra more shining, elytra as densely and
strongly punctured as pronotum. Aedeagus of accola sp. n. with a beak-like promi-
nence, arms of tegmen narrower than in obliqua.

Similar to Atomaria torrida in antennal structure and appearance, but differs in
coloration, segment 5 of antenna longer than 3"4 (segment 5 of torrida nearly as long
as 3rd, 5th segment of accola sp. n. somewhat longer than that of torrida), punctation
of pronotum and elytra more strong and dense, pubescence longer.

The new species differs from Atomaria plecta and angellata by the antennae
which are considerably longer (7'h segment very long), the larger punctation of the
pronotum and elytra, and by the structure of the aedeagus.

Atomaria (Anchicera) mentita sp. n. (Figs 21-22)

Holotype (d): Nepal, Sankhua Sabha Distr., above Pahakhola, 2600-2800 m, Quercus
semecarpifolia-Rhododendron, 31. V.-3. VI. 1988 leg. MARTENS & SCHAWALLER (SMNS).

Paratypes: Same locality and data as holotype (1 2 SMNS, 1 d ZMUM); - Nepal, Gor-
kha Distr., Chuling Khola, 2800 m, Quercus semecarpifolia forest, 2.-3. VII. 1983 leg. Mar-
TENS & SCHAWALLER (1 2 SMNS).

Description: Body length 1.25-1.40 mm, light brown (head sometimes dark
brown), covered with short decumbent silvery pubescence, hairs somewhat shorter
than claw joint. Eyes normal, head weakly and sparsely punctured, punctures separ-
ated from their nearest neighbours by 1.0-1.5 diameters on the disk. Antennae short
and stout, as in Fig. 21; segments 1-3 slightly elongate, segment 1 longer than 2nd,
latter longer than 3"4, segments 4-8 not elongate, 5‘ and 7 subquadrate, 4*, 6th and
8th transverse. Club stout, well-marked, segments 9 and 10 strongly transverse. Pro-
notum transverse, 1.38 times as broad as long, broadest across the middle, sides
slightly curved and slightly narrowed basally; side borders single, visible from above
only in basal half. Pronotal surface shining, not shagreened, extremely densely cov-
ered with deep punctures separated from their nearest neighbours by a half-diame-
ter on the disk; hind angles obtuse; pronotal disk convex; base of the pronotum with
a narrow transverse depression; hind margin finely bordered, not or barely pro-
duced caudomedially. Elytra short, strongly arched, broadest about the middle,
1.9-2.0 times as long as pronotum and 1.15-1.20 times as long as broad combined;
surface shining, not shagreened, densely punctured, the punctures in basal half sim-
ilar in size to those on pronotal disk and separated from their lateral neighbours by
approximately one diameter on the average. Wings absent. Aedeagus as in Fig. 22.

Remarks: This wingless species is similar to the Caucasian Atomaria circassica
Reitter 1888 and cephennoides Reitter 1887 because of the short and stout antennae
and the rather short elytra. Differs by the body narrower, strongly punctured and
the pubescence short and decumbent. Differs from the wingless Nepalese Atomaria
khumbuensis by the lighter body, the shorter antennae, the more strongly transverse
segments of the club, and the poorly curved sides of the pronotum.

Atomaria klapperichi Johnson 1970

Material: Nepal, Dolakha Distr., SW Kalinchock Mt., 3100 m, 19.-23. IV. 1995 leg. Mar-
TENS & SCHAWALLER (1 d SMNS); — Ilam Distr., Mai Pokhari, 2100-2200 m, Castanopsis for-
est remains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (1 6 SMNS); - Kaski Distr., above
Pothana, 2000 m, 27.-29. IV. 1995 leg. MARTENS & SCHAWALLER (1 d ZMUM).

Distribution: Nepal, India, Afghanistan. This is the first report in Nepal.

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 23

Atomaria pudica Johnson 1970

Material: Nepal, Ilam Distr. road from Ilam to Birtamode, Karphok, 1700 m, cultivated
land with bushes, 2. IV. 1980 leg. MARTEns & Ausogsky (1 d ZMUM); - Iam Distr., Mai
Pokhari, 2100-2200 m, Castanopsis forest remains, 9.-10. IV. 1988 leg. MarTENS & ScHa-
WALLER (3 ex. SMNS, 1 ex. ZMUM); - Dolakha Distr., SW Kalinchok Mt., 3100 m, 19.-23. IV.
1995 leg. MARTENS & SCHAWALLER (1 ex. SMNS).

Distribution: Nepal, India. This is the first report in Nepal.

Atomaria khumbuensis Johnson 1971

Material: Nepal, Mustang Distr., Thaksang, 3150 m, Pinus excelsa-Abies forest, 26.-29.
IV. 1980 leg. Martens & Ausossky (8 ex. SMNS, 3 ex. ZMUM).

Distribution: Nepal.

Atomaria frugi Lyubarsky 1997

Material: India, Kashmir, Tangmarg, Pir Panjal Mts., 2600 m, 21.-25. V. 1976 leg. Mar-
TENS & SCHAWALLER (1 $ SMNS); — Nepal, Ilam Distr., Mai Pokhari and Ilam, 1330 m, 1. IV.
1980 leg. Martens & Ausogsky (1 d ZMUM).

Distribution: India, Nepal.

Atomaria plecta Lyubarsky 1996 (Fig. 23)

Material: Nepal, Manang Distr., Marsyandi, above Bagarchap, Acer-Quercus forest,
2200 m, 12.-13. IV. 1980 leg. MARTENS & Ausossky (6 ex. SMNS, 3 ex. ZMUM); - Taplejung
Distr., Omje Kharka, NW Yamputhin, mature mixed broadleaved forest, 2300-2500 m, 1.-6.
V. 1988 leg. MARTENS & SCHAWALLER (2 22 SMNS); — Ilam Distr., Mai Pokhari,
2100-2200 m, Castanopsis forest remains, 9.-10. IV. 1988 leg. MARTENS & SCHAWALLER (1 2
SMNS).

Distribution: China (Shaanxi), Nepal.

Remarks: Certain intraspecific variation has been elucidated from the above ma-
terial. Thus, the 7 antennal segment varies in shape from subquadrate to slightly
elongated; the apex of the paramere plate can be rounded (usually) or slightly con-
cave in the middle. The spermatheca is as in Fig. 23. Atomaria khumbuensis, frugi,
and plecta are similar, the differences being shown in the key.

2.1.10. Key to Atomaria species from Nepal, based on JOHNSON (1970)
and the above new findings

I Vınes absent 1.0 Kr ra ya ca he ah ated ita ea aa fee ae east ANd Eee 2
SMM inSsspresemt Er aa ee ee a be ee ce ag cies) sta oes eile ale cago 3

2 Elytra reddish-brown, with a vaguely darker area on either side of the disk in some speci-
mens. Antennae somewhat stout, segment 5 a little longer than 3", 7th transverse, 9" and
10th rather strongly transverse. Sides of pronotum more strongly curved. Paramere plate
zoundedlapıcallyalseneth1r93_ 142mm. Seer eg. khumbuensis

— Lighter body colour. Antennae short and stout, segments 5 and 7 subquadrate, club stout,
well-marked, segments 9 and 10 strongly transverse. Sides of ponotum almost subparallel,
ablittlexcumvedleemetiill 2 5— le 4 Omani 2.2... 0000 ee nennen mentita sp. n.

3 Body form narrow, elongated, pronotum and elytra with the sides subparallel, very feebly
curved at most; antennae close together at base, segment 1 about twice as long as broad.
(AGAR ATES SUR) OH so Fe GRC eevee ans ot ote ee Oe eo hn Ome oe ian 4

2A

10

11

STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 598

Body form proportionately broader and shorter, more curved on sides; antennae usually a
little more strongly separated at base, segment 1 about as long as broad in most species:
few species’ have it twice as longeas broadi(Amebicera) 2... u... 0 er 5

Base of pronotum with a transverse depression; pronotum somewhat sparsely punctured,
surface not shagreened; elytra somewhat coarsely and sparsely punctured, not or feebly
shagreened; brownish species, elytra generally with a lighter spot from humeri to disk; an-
tennae rather long, segments 9 and 10 clearly but not strongly transverse; more convex
species. Length 1.4-1.8mm. Apical half of penis with a large Y-shaped sclerite internally

ig RRR ER RE N 00.40.0000 prolixa
Base of pronotum without transverse depression; pronotum generally more finely and
densely punctured and shagreened; elytra more finely and densely punctured, strongly
shagreened; dark brownish species, elytra often paler but lacking a humeral-discal streak;
antennae rather short, segments 9 and 10 about as long as broad; more depressed species.
Length 1.6-1.8 mm. Apical half of penis without large Y-shaped internal sclerite .......

sige Hb arate iia alors evade ives Desi cinta ine Bis GEER Se CRIA La gracilicornis

Antennal segments 9 and 10 strongly transverse, club very broad and well-marked; seg-
ment 7 clearly transverse, 6" and 8! more so; body form very short, elytra rather strong-
ly roundedjon'sidessireddish speciest-pj15 ances oon aciiee See i oe eee 6
Antennal segments 9 and 10 more weakly transverse, club narrower and not so well-
marked; segment 7 slightly elongate, 6!" and 8» quadrate to feebly transverse; body form
generally not so short, more weakly rounded on'sides\.........)..-- 13 eee 7

Antennal segment 1 nearly twice as long as broad and about 1.5 times as long as segment
2; elytral pubescence a little outstanding. Apical prolongation of penis broader than long,
shorter sides more narrowed basally=enn ene tea ae re ee eee lewisi
Antennal segment 1 about as long as broad, not or but little longer than segment 2; elytral
pubescence generally slightly more outstanding. Apical prolongation of penis longer, as
long as or slightly longer than broad, sides more subparallel ................ incertula

Head, pronotum and elytra monochromously reddish-yellow/brown, darker, at most
withithe'suture,somewhat intuseated 22 ols eine iene eee ee 8
Head and pronotum reddish-yellow, elytra a dirty yellow (straw-coloured) with a more
or less irregular brownish-black transverse band across the disk, this forming a V-shape
astride the suture. ann an. sunt e em RL 10

Antennae rather stout, segments 9 and 10 weakly transverse, club narrow and indistinct;
pronotum 1.41-1.46 times as broad as long; base of pronotum with a deep and well-de-
fined transverse depression; body more convex, sides of the elytra more strongly curved.
Eength1:57- 1.65: mm sr Ma Nae a ee torrida
Antennae more slender, segments 9 and 10 more strongly transverse, club broader and
more distinct; pronotum 1.46-1.64 times as broad as long; base of pronotum with a very
narrow and poorly-developed depression; body more depressed, sides of the elytra more
weakly curved: sn. ne ee de a ek, nee >

Size larger, 1.54-1.65 mm; antennal segment 1 about as long as broad, subequal in length
to segment 2; elytral pubescence rather long and outstanding, hairs 0.05-0.06 mm long;
pronotum 1.52-1.64 times as broad as long, breadth 0.74-0.80 mm; sides of elytra very
wealelyscurved ee Sacre ital ata RT EINE a klapperichi
Size smaller, 1.41-1.52 mm; antennal segment 1 a little longer than broad, slightly longer
than segment 2; elytral pubescence shorter and more depressed, hairs 0.04-0.05 mm long;
pronotum 1.41-1.52 times as broad as long, breadth 0.54-0.59 mm; sides of elytra more
obviously curved; elytral suture generally clearly infuscated .................. pudica

Segment 7 of antennae strongly elongated, 9% subquadrate or very feebly transverse. Ely-
traleVoshape transverse band distinct. 9 Messe ene ee eee 11
Segment 7 of antennae transverse, 9% joint strongly transverse. Elytral V-shape transverse
bandunmdistinct, 3 3.3 deat ee ange lesa ee ee sca ee 12

Antennae shorter, 1°! antennal segment slightly longer than wide and about equal in length
to 2nd; 5th antennal segment equal to or slightly longer than 2"4, Pronotum somewhat less

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 25

transverse (width/length ratio 1.45-1.60), its sides smoothly rounded, not angular. Size
somewhausmaller (130 145mm) sr re ee obliqua
— Antennae longer, 1% antennal segment much longer than wide and 24, 5th antennal seg-
ment longer than 2nd. Pronotum somewhat more transverse (width/length ratio
1.40-1.55), its sides strongly rounded, angular in the middle. Punctation more dense; pro-
notum and elytra more shining. Size somewhat greater (1.4-1.7mm) ...... accola sp. n.

12 Antennae stout, segment 7 subquadrate or a little longer, 9 transverse. Pronotum slight-
ly narrowing towards base, with obtuse posterior angles. Elytra with a darker area on each
side of the disk or with wide transverse band. Paramere plate widely rounded or slightly
simulate apically-|Wengeln 1.2 Ilomm nn ya. ae ne ee plecta

- Antennae slender, segment 7 transverse, 9 subquadrate. Pronotum strongly narrowing
towards base, with sharp posterior angles. Elytra light reddish-brown, monochromous.
Paramere plate clearly sinuate apically. Length 1.3-1.4mm .............-..+.-. frugi.

2.1.11. Curelius Casey 1900

Curelius japonicus (Reitter 1877)

Material: Nepal, Dhading Distr., W Samari Banjyang/Topal Khola, 1000m, forest re-
mains, 23. VII. 1983 leg. MARTENS & SCHAWALLER (1 ex. ZMUM); - Iam Distr, N Mai
Pokhari, Tai Gitang Khola, 2500-2600 m, 18.-31. III. 1980 leg. Martens & Ausossky (1 ex.
SMNS); - Ilam Distr., Mai Pokhari, 2100-2200 m, 25.-27. III. 1980 leg. MARTENS & AusoB-
sky (1 ex. ZMUM).

Distribution: Europe (Spain), E Asia (from Japan to Java), Africa (from Ethi-
opia to Zimbabwe), America (from Ohio, USA to Brazil) JOHNSON 1989).

2.1.12. Ephistemus Stephens 1829

Ephistemus splendens Johnson 1971

Material: Nepal, Sankhua Sabha Distr., Arun valley, Chichila, Quercus forest, bushes near
village, 18.-20. VI. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS, I ex. ZMUM); - Gorkha
Distr., Darondi Khola, between Naya Sangu and Gorkha, 1200 m, 14. VIII. 1983 leg. Mar-
TENS & SCHAWALLER (1 ex. SMNS); - Kathmandu valley, Mt. Sheopuri, Quercus semecarpifo-
lia forest, 2100-2300 m, 25. VI. 1988 leg. MARTENS & SCHAWALLER (1 ex. SMNS).

Distribution: India. First record in Nepal.

2.2.Family Languriidae

2.2.1. Cryptophilus Reitter 1874
Cryptophilus integer (Heer 1838)

Material: Nepal, Iam Distr., Bililate near Ilam, 1330 m, remains of trees around spring,
moist soil, 8. IV. 1988 leg. MARTENS & SCHAWALLER (6 ex. SMNS, 1 ex. ZMUM); - Ilam
Distr., between Mai Pokhari and Ilam, 1330 m, 1. IV. 1980 leg. MARTENS & Ausopsky (1 ex.
SMNS, 1 ex. ZMUM); - Chitwan Distr., Chitwan National Park, Sauraha, 150 m, 31. V—4. VI.
1997 leg. SCHAWALLER (3 ex. SMNS).

Distribution: Palaearctic, India, Nepal, S America. This is the first formal re-
port of the species from Nepal.

26 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 598

2.2.2. Loberus Leconte 1861

Loberus sinuaticollis Bruce 1945 (Fig. 7)

Material: Nepal, Taplejung Distr., Kabeli Khola Valley below Limbudin, 950 m, river
bank with bushes, 22.-23. IV. 1988 leg. MARTENS & SCHAWALLER (1 ¢ SMNS).

Distribution: Nepal, Burma. This is the first formal record of the species in
Nepal.

Remarks: This species was originally described from two females deriving from
Burma (Tenasserim Coast, Ale Mekane, 90km E Moulmein, 200 m) (Bruce 1945).
The aedeagus is thus depicted here for the first time (Fig. 7).

3. Acknowledgments

I am very grateful to Dr. W. SCHAWALLER (Stuttgart) for sending me this material for study
and for arranging the publication, to Dr. S. GoLovatrcu (Moscow) for checking the English
of an earlier draft, and to the Russian Foundation of Basic Research for awarding me a subsid-
iary grant (97-0448661).

4. References

Bruce, N. (1945): Entomological results from the Swedish Expedition 1934 to Burma and
British India. - Ark. Zool. (36A) 4: 1-5; Uppsala.
- (1952): Coleoptera Cryptophagidae in the British Museum. — Ann. Mag. nat. Hist. (12)
5: 167-188; London.
— (1959): Einige Cryptophagiden (Col.) aus Afghanistan nebst Beschreibung von Crypto-
phagus robustus sp. n. — Opusc. ent. 24: 18-20; Lund.
JOHNSON, C. (1970): Atomariinae (Col., Cryptophagidae) from the northern parts of the In-
dian sub-continent with descriptions of seven new species. - Entomologist’s mon. Mag.
107: 224-232; London.
- (1971): Cryptophagidae — Atomariinae from Nepal (Col.). - Senckenberg. biol. 52:
437-439; Frankfurt/M.
- (1972): Beiträge zur Kenntnis der Fauna Afghanistans. Atomariinae, Cryptophagidae,
Coleoptera. — Cas. moray. zemsk. Mus. 56-57: 261-265; Brno.
- (1989): Cryptophagidae (Coleoptera) of the Arabian Peninsula. — Fauna of Saudi Ara-
bia 10: 108-112; Basel.
LescHEN, R. A.(1996): Phylogeny and revision of the genera of Cryptophagidae (Coleoptera:
Cucujoidea). — Kans. Univ. Sci. Bull. 55: 549-634; Lawrence.
Lyusarsky, G. (1987): Species review of the genus Sternodea Reitter. — Bull. Mosk. obshch.
ispyt. prir., biol. 92: 53-62; Moscow. [in Russian]
— (1991): Review of Palaearctic species of the genus Antherophagus Latr. (Coleoptera,
Cryptophagidae). - Zool. Zhurn. 70: 63-67; St. Petersburg. [in Russian]
— (1996): Cryptophagidae and some Languriidae from palaearctic China. — Russ. ent. J. 4:
45-53; Moscow.
— (1997): Cryptophagidae and Languriidae from India (Coleoptera, Clavicornia). —
Entomofauna 18: 49-60; Ansfelden.
- (1998): Scytomaria, a new genus of Cryptophagidae (Coleoptera) from the Nepal Hi-
malayas. — Stuttgarter Beitr. Naturk. (A) 572: 1-6; Stuttgart.
Nixitsky, N. (1996): New Coleoptera from China. — Zool. Zhurn. 75: 1366-1373; St. Peters-
burg. [in Russian]
Reska, M. (1994): Bestimmungstabellen der mitteleuropäischen Arten der Gattungen
Micrambe Thomson und Cryptophagus Herbst (Insecta: Coleoptera: Cryptophagidae).
— Annln naturhist. Mus. Wien 96B: 247-342; Wien.
SEN Gupta, T. (1978): A new genus of Atomariinae (Coleoptera: Cryptophagidae) with de-
scriptions of two new species from India. - Orient. Insects 12: 275-279; New Delhi.

LYUBARSKY, CRYPTOPHAGIDAE FROM THE HIMALAYAS 27,

SEN Gupta, T. & Basak, P. K. (1985): Discovery of the genus Caenoscelis Thomson (Coleo-
ptera: Cryptophagidae) from Indian region and description of a new species from
Nepal. - Bull. zool. Surv. India 7: 41-43; Calcutta.

SEN Gupta, T. & Pat, T. K. (1980): Ergebnisse der Bhutan Expedition 1972 des Naturhisto-
rischen Museums in Basel. Coleoptera: Fam. Cryptophagidae, genus Himascelis. - Ent.
basiliensia 5: 42-44; Basel.

Author’s address:

GEORGY Lyusarsky, Zoological Museum of Moscow State University, B. Nikitskaya 6,
RUS-103009 Moscow, Russia.

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