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Var geetay ws ver joes: y sWeyee sey mh = j 7 ‘g | ~ wW = ~" wv. cUvy Je Hey bs yer” MT TI | vee tren peewee weet werent gs ehtyyy "4 ¥y Hy We Meey A Ae 10 oov hi u ett aoe uw § rh. rite b: we! 5 | wee ; TT 1% U ttib tid GILL Jove id On PET pe Te PY TTT rETLiGcIifC wf as. e / Se ea Oath ban pe ohne OCCASIONAL PAPERS ot etOR OF THE CALIFORNIA ACADEMY OF SCIENCES |e, VII _ SYNOPSIS cme OF California Stalk-Eyed : Crustacea BY ite’ 4 ae SAMUEL Jf Houwes 2 eR ot a : 2 ~ Prue ay) STAN Th baila BANS s\ Issued June 15; 1900 e: rhea | ee f eg , SAN FRANCISCO Pein! mene CALIFORNIA ACADEMY OF SCIENCES 1900 Occasional Papers ae ee, ea CALIFORNIA Ne eS %,\ “9 ACADEMY OF SCIENCES SAN FRANCISCO 1900 PUBLICATION COMMITQLEE: CHARLES H. GILBERT, Chairman. WILLIAM E. RITTER, G. P. RIXFORD: EDITORS OF ZOOLOGIGAL POLITICA OS, CHARLES H. GILBERT. WILLIAM E. RITTER. QL NUH M3AHGHZ Tnwert 2 SYNOPSIS OF California Stalk-Eyed Crustacea BY SAMUEL J. HOLMES igt i. eee J e VAN INS Dias FON* : 49 NY SMITHSOM Any ued June 15, 1900 is Oe 4 \ j - \ me fe ti 6 13 g Ce (om Sete ¥ 3 : ows aon, SEP 0 9 1986 LIBRARIES Ny SAN FRANCISCO CALIFORNIA ACADEMY OF SCIENCES 1g0o -_ , | ; PRINTED FROM THE HENDRIE PUBLICATION Funp. na ee eee PREFACE. The aim of the present paper is to give descriptions of the species of Stalk-eyed Crustacea which are found on the west coast of the United States. For the benefit of students of natural history, who have at present few readily accessible means of identifying the invertebrate animals of this coast, definitions of genera and higher divisions have been inserted, and a description given, in the Introduction, of those parts of the structure of the Decapod Crustacea which are most used for the purpose of classification. The endeavor has been made to enable those who have not previously studied the Crustacea to determine the specific names of the forms — here described. The rules of nomenclature of the American Ornithologists’ Union have been followed, except in case of the canon, not insisted on by the framers of the code, that specific names derived from the names of persons be written with a small letter. Localities from which I have examined spec- imens have usually been indicated by the note of exclamation, !. The larger portion of this paper was completed at the University of California in 1895, but it has received some additions and considerable revision since that date. I wish to express my indebtedness to Prof. Wm. Ki. Ritter of the University of California for many favors in connection with my work, and to Dr. H. P. Johnson and Dr.’ F. W. Bancroft for several valuable specimens and for assistance in many other ways. Through the kindness of Dr. Gustav Eisen I have had 6 PREFACE. the privilege of studying the collections of Crustacea in the California Academy of Sciences. The examination of Mr. Lockington’s type specimens, many of which are in the possession of the Academy, has made it possible to straighten out some questions which otherwise would scarcely have been capable of solution. To Mr. Rivers and Mr. Harford, formerly of the University of Cali- fornia, I am indebted for the privilege of studying the collections in the University museum. From Miss Mary J. Rathbun and Dr. James E. Benedict of the U.S. National Museum much valuable aid has been received which is gratefully acknowledged. Ss. J. HOLMES. UNIVERSITY OF MICHIGAN, Ann Arbor, Mich. January 20, 1900. SYNOPSIS OF CALIFORNIA STALK-EYED CRUSTACEA. BY SAMUEL J. HOLMES. INTRODUCTION. THE Crustacea are commonly divided into two princi- pal groups, the Malacostraca and the Entomostraca, although the primary divisions of the latter group may well be, as they often are, considered coordinate with the former. There is such great diversity of form among the Entomostraca that the group is scarcely definable except by negative characters. The number of segments composing the body is very variable and the openings of the genital ducts do not occur in the same segment in all the members of the group. There is no masticatory stomach and corresponding append- ages are modified in most diverse ways in different orders. The Malacostraca, on the other hand, while they include a great variety of forms, constitute a well defined subclass. Except in cases in which it is quite evident that certain segments are fused or lost, the body is composed of twenty (or in Nebalia of twenty-one) segments. The typical number of appendages is nine- teen. There isa pair of compound lateral eyes, and the stomach, except in a few parasitic species, is masticatory. The genital ducts of the male open on the last thoracic segment; those of the female on the antepenultimate one. The head is typically composed of five segments [7] 8 CALIFORNIA ACADEMY OF SCIENCES. and the thorax of eight, some or all of which may fuse with the head forming a more or less complete cephalo- thorax. The abdomen is sharply marked off from the thorax and consists of seven (in Nebalia of eight) seg- ments, all of which, except the last, the telson, may bear articulated appendages. The Malacostraca may be divided as follows:— Body composed of 21 segments and furnished with a bivalve shell. Leptostraca. Body composed of 20 segments and not furnished with a well defined carapace. : Kyes:sessilogss vant ruin cree meee Ane Arthrostraca.- Body composed of 20 segments and furnished with a well developed carapace which is never bivalved................-20- Thoracostraca.. SYNOPSIS OF THE THORACOSTRACA. Biyesvsessile. 2. hci 0 1 eras ys aia ee et Ne ee Cumacec.. Eyes stalked. Carapace rather small, not covering all of the thoracic segments. The thoracic feet dissimilar, the anterior ones devoid of exopods. Stomatopoda, Carapace covering some or all of the thorax but not joined to the . posterior segments. Thoracic feet usually similar, biramous. Schizopoda.. Carapace covering and joined to all of the thoracic segments. The three anterior thoracic feet generally biramous and modified into. moO tbh-paits 00.0... 2050 hs. pee oe ee ee ee eee Decapoda. As it is with the Decapoda that we are here mainly con- cerned, those parts of the structure of these animals which are most used for the purpose of classification will be briefly described. The exoskeleton of a Decapod crustacean may be considered as a series of rings, each of which, with the exception of the last, is typically furnished with a pair of articulated appendages. These rings and their appendages are modified in various ways in different regions of the body in relation to the different functions they are called upon to perform. CALIFORNIA STALK-EYED CRUSTACEA. 9 The segments in some parts become fused together so completely that they manifest no external signs of metamerism except the appendages they carry. The five segments forming the head are usually thus indis- tinguishably fused. The eight following segments, with the rare exception of the last one, are fused together and with the head, forming the cephalothorax; but the sutures between the rings are, in this case, generally visible on the ventral, or sternal.surface and at the sides. The abdomen is, in all cases, clearly marked off from the thorax, and is typicaily composed of seven segments. The number of joints may, however, be secondarily reduced by the concrescence or disappear- ance of some of the rings. Phe Carapace.—The carapace of the Decapod crustacea is considered to be a fold of the dorsal integument of one or more of the cephalic segments which extends backward over, and fuses with the dorsal portion of all of the thoracic segments and is produced downwards at the sides over the gills. The form of the carapace commonly found in the Macroura is shown in the fol- lowing cut:— “ber Fig 1.—Macrouran Carapace. a.s, antennal spine; b.7r, branchial region; c. g, cervical groove; h. s, hepatic spine; 7, rostrum; s. 0. s, suborbital spine; s. s, supra- orbital spine. 10 CALIFORNIA ACADEMY OF SCIENCES. The typical Macrouran carapace is elongated, and the anterior end is produced into a rostrum. The anterior region of the carapace is often furnished with teeth or spines which occur in the following positions: the supra- orbital spine, or spines, above the orbit; the suborbital spine on the anterior margin of the carapace below the orbit; the antennal spine below the last and behind the base of the antenna; the hepatic spine on the he- patic region. Any orall of these spines may be present or absent. There is generally visible a groove, known as the cervical groove, which extends downwards and forwards from the dorsal surface of the carapace, mark- ing the line of union of the head and thorax. The dorsal surface in front of this groove is termed the gastric region, the hepatic regions lying at the sides; immediately behind the gastric region is the cardiac, while the intestinal region les behind the latter at the posterior end of the carapace; the sides of the carapace behind the cervical groove are known as the branchial regions, since they form a covering for the branchie, or gills. In the Brachyura the carapace is wide and generally flattened, and the rostrum is small or absent. The regions are generally separated from each other by sulci, or grooves, and there are usually more regions marked out than in the Macroura, The cervical groove is usually visible and divides the gastric and hepatic from the cardiac and branchial regions. On the sides of the carapace below the lateral margins there are (1), the pterygostomian region on either side of the mouth area and (2), the sub-branchial regions below the branchial areas. The names of the regions which appear on the dorsal surface may be obtained from fig. 2. The dis- tinctness of these areas is subject to great variation in CALIFORNIA STALK-EYED CRUSTACEA. ha different forms. In some cases the outlines of all the regions are very clear, while in others they may be entirely obliterated. Other areas are sometimes given names but they are of little importance. 3 4 O Fig. 2.—Carapace of acrab. 3.—Mandible of acrayfish. 4.—First maxilla. 5.— Second maxilla. 6.--Third maxilliped. 6, basipodite; b. 7, branchial region; c, coxo- podite; c. g, cervical groove; cp, carpus; c. 7, cardiac region; d, dactyl; en, endopodite; ep, epipodite; ex, exopodite; h. r, hepatic region; 1, ischium; 7. r, intestinal region; m, merus; m.7T, mesogastric region: p, palp; p. g, protogastric areas; pr, propodus; rT, rostrum; wv, urogastric area. THE APPENDAGES. With the exception of the eyes and the first pair of antenne all of the appendages of a Decapod crustacean may be considered modifications of a common type 12 CALIFORNIA ACADEMY OF SCIENCES. consisting of a basal portion, the protopodite, and two terminal rami. The first basal joint, the coxopodite, or coxa, usually bears a branchial appendage, or epipo- dite; the second joint, the basipodite, carries the two rami; the inner ramus is called the endopodite, the outer the exopodite. In the Decapods the endopodite of the thoracic appendages is typically composed of five joints which, counting from the basipodite, are desig- nated as the ischium, merus, carpus, propodus, and dactyl. In the appendages which precede and in those which follow the thorax the endopodite is undivided, or composed of less than five true joints. The appendages will be described in their order from before backwards. The Eyes and Orbits.—The eyes in all of the Decapods are compound and situated at the end of movable stalks. These stalks, however, are probably not homologous with the other appendages. The orbits in the Macroura are generally represented by excavations in the anterior margin of the carapace, but in the Brachyura they form well defined cavities into which the eye-stalks can be withdrawn. The number and position of the orbital fissures are important characters for the purpose of classification. The First and Second Antenne.—The first antenne, or antennules, unlike the succeeding appendages, are not typically biramous. It is true that they may bear two terminal branches, or in some of the lower Macroura even three, but it is quite clear that this condition is a secondary one. The basal portion, or peduncle, nor- mally consists of three joints, in the first of which is located the auditory organ. In the Brachyura the antennules are small, and located in cavities called fossettes. CALIFORNIA STALK-EYED CRUSTACEA. 13 The ‘second antenne are usually larger than the first pair and situated behind or external to them. The exopod is represented by a scale, or acicle, attached to the second basal joint, but this structure is often absent, especially in the higher forms. The peduncle usually contains four or five joints and bears an annulated, terminal portion, the flagellum. The first joint contains the opening of the renal organ and is often immovably joined to the carapace. The Mundibles.—The appendages which function as mandibles are highly modified. The first joint (coxo- podite) is enlarged and modified for the purpose of mastication; this part is called the trunk and its inner edge is generally dentate and often divided into a lower portion, or molar process, and an upper cutting edge. The trunk bears a jointed appendage, or palp, which, in this group, never consists of more than three joints. The First and Second Maaxille.—The first maxille are always small and foliaceous. The two basal joints are flattened and produced inward; there is a small endo- podite; the exopodite is usually absent. In the second maxille the two basal joints are modified much like those of the first pair. There is a small endopodite outside of which is joined a large, flattened appendage (exopodite, scaphognathite) which serves to produce a current of water from the branchial chamber. The First Maaxillipeds.—These are the first thoracic appendages. The two basal joints are flattened and produced, forming the so-called lacinize, or masticatory ridges. The endopod is small and generally joined to a greater or less extent to the exopod; the latter is well developed and ends in an annulated, terminal portion, or palp. 14 CALIFORNIA ACADEMY OF SCIENCES. The Second Muzxillipeds.—The second maxillipeds are more like the third pair than the first; the endopod is well developed and generally contains the typical num- ber (five) of joints. The exopod is also well developed and furnished with a terminal palp. The epipod often bears one or more gills. The Third Maaillipeds.—The endopod of the third maxillipeds is usually composed of five joints. In the Brachyura the ischium and merus are flattened and generally fitted neatly to the buccal area; the last three joints are small and are termed the palp; in the Macroura the joints of the third maxillipeds are usually rounded. The exopod may be well developed, but is sometimes wanting. The Thoracic Legs, or Pereopods.—The thoracic legs, with rare exceptions, consist of seven joints, two or more of which are generally fused together. The exo- pod is wanting, except in some of the lower Macroura, where it is represented by a small, jointed appendage. One or more of the thoracic legs may be furnished with pincers or chele, in which case they are called chelipeds; the movable finger of the hand, or chela, is called the dactyl, the immovable one the pollex; the basal portion of the hand is known as the palm. The Abdominal Appendages, or Pleopods.—The pleo- pods in the Macroura are generally furnished with flattened rami adapted for swimming; the posterior pair is usually much larger than the others and forms with the telson a strong tail fin. In the Brachyura these appendages are never employed in swimming; in the females they are used for holding the ova; in the males only one or two of the anterior pairs are retained as sexual appendages and these are sometimes wanting. CALIFORNIA STALK-EYED CRUSTACEA. 15 The Gills.—The gills may be situated on the basal joints of the thoracic appendages (podobranchie), or on the articular membranes between the legs and the body (arthrobranchie), or on the sides of the thoracic segments (pleurobranchie). There are two types of gills—the trichobranchie, in which there are thread- like, branchial filaments arranged around a common stem, and the phyllobranchiz, in which the stem bears on each side a row of thin, transverse lamelle. A few other structures deserve mention in this con- nection, viz.:—the epzstoma, the area lying in front of the buccal, or mouth area and behind the antennules; the upper lip, a fleshy projection in front of the mandi- bles; the lower lip, a flap lying behind the mandibles; the thoracic sternum, or sternal plastron, the ventral por- tion of the thorax, and the terminal segment of the -Macrouran abdomen, the telson. Tribe BRACHYURA. Carapace broad. Orbits well defined. Antennules lodged in fossettes. Antenne short and devoid of an acicle. The external maxillipeds have the ischium and merus flattened and usually fitted neatly to the buccal area. First pair of legs always chelate; the following pairs always simple. The abdomen is small, shorter than the carapace, and flexed against the ventral surface of the thorax. The abdomen of the male is smaller than that of the female and bears but one or two pairs of appendages near the base; abdominal appendages in the female more numerous than in the male and © used for holding the ova. Tail fin absent. The: subdivisions of the Brachyura can usually be recognized by their general form. The Maioidea, or spider crabs, have the carapace narrowed in front where it is generally produced into a rostrum. The Cancroidea usually have a transverse carapace with a 16 CALIFORNIA ACADEMY OF SCIENCES. narrow front. The Grapsoidea may commonly be rec- ognized by their quadrate form, but there are many forms which do not exhibit this character. The Oxy- stomata may be distinguished by the form of the buccal area which is narrowed in front. None of these marks, however, are absolutely diagnostic, for it is difficult to separate the groups by single characters. Subtribe MAIOIDEA, or OXYRHYNCHA. Carapace narrowed in front and usually furnished with a rostrum Branchial regions large; hepatic regions small. Buccal area quadrate. Kpistoma usually large. The afferent channels open behind the pterygo- stomian regions in front of the chelipeds; efferent channels opening at the’ sides of the endostome. Antennules longitudinally plicated. Palp of the maxillipeds situated at the summit or antero-internal angle of the merus. Verges of the male on the coxz of the last pair of legs. This group includes the species known as the ‘‘ spider crabs’’ and ‘‘sharp-nosed crabs.”’ They are generally of sluggish habit and many of the species become over- grown with sea-weeds and other foreign growths, which make them difficult to detect in their natural habitat. The classification followed is that given in Miers’ revision of this group.’ Legion MAIINEA. Basal joint of the antennz well developed and inserted beneath the eyes, usually occupying a great part of the infra-orbital space. Family INACHID/AL. Orbits not defined. Preorbital and postorbital spines generally well developed. Eyes not retractile, or retractile against the sides of the cara- pace. Basal antennal joint generally slender. Rostrum simple or bifid. Fingers of the chele never excavated at the tips. 1 Jour, Linn. Soc., London, Vol. XIV, 1879. CALIFORNIA STALK-EYED CRUSTACEA. L7 Subfamily LEPTOPODIIN &. Eyes slender, not retractile, laterally projecting. Preocular and post- ocular spines small or wanting. Basal joint of the antenne slender throughout its length. Rostrum usually simple. Genus Podochela Sz. Carapace depressed, narrowly triangular, strongly produced in front. Gastric region narrow, tumid, rostrum short, entire, triangular or arcuate. Eyes long and laterally projecting. Basal antennal joint narrow, with a median, longitudinal groove; the apex narrow and not dentate. Flagellum slender -and visible from above. Merus of external maxillipeds much shorter than the ischium, the apices obtuse or produced, the inner one more or less incised; palp joined at the summit of the merus. Epistome very long. Chelipeds of moderate size; merus curved, the margins cili- ated. Ambulatory legs very long, subprehensile, and often furnished with a more or less cheliform hand. Dactyls of chelipeds very slender and often hooked, those of the remaining legs falciform. Sternum of female deeply concave and surrounded by a laminate margin. First abdominal segment quite large, second, third, and fourth very short; the last two or three segments fused together. Type.—P. grossipes St. Podochela Hemphillii (Lock: ) Microrhynchus Hemphillii Lock1netTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 30. Inachoides ( Microrhynchus) Hemphillii LocK1neTon, 1. ¢., p. 75. . Podochela tenuipes RatuBun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 224. Podochela Hemphilliti RatHBun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, Pp. 569. Q ‘ Carapace pyriform; gastric region prominent, rounded, and bearing tufts of curved sets; hepatic regions tumid, bearing two pointed tubercles, the larger one above and in front of the smaller; cardiac region separated by shallow grooves from the branchial and bearing a prominent elevation; branchial regions flattened or tumid. Rostrum triangular, acute, and bearing two double rows of curved setz above; it varies considerably in length, being sometimes broadly and sometimes narrowly triangular. The area between the two projecting rims of the orbits is concave. ‘There is no tooth at the posterior margin of the orbit, but there may be a small one a short distance behind it. Eye-stalks constricted at the middle and 2 18 CALIFORNIA ACADEMY OF SCIENCKS. bearing a few setz at the subacute tip. Basal antennal joint with a longi- tudinal ridge on its posterior half or two-thirds with a groove on either side. Ischium of maxillipeds with a shallow, longitudinal groove; merus narrower than ischium, the internal angle produced, and the surface strongly concave from side to side. Chelipeds in the male robust; merus incurved and having an outer spiny ridge; carpus with a posterior spine on the upper side; hand oblong, incurved, palm inflated, fingers gaping at the base and meeting along the distal half, or two-thirds of their length; there are generally several small, spiny projections on the surface of the hand, which are more conspicuous along the edges, especially the lower. In the female the chelipeds are smaller and more slender, the hands subcylindrical, .and the fingers nearly straight. Legs very long and slender, and furnished with tufts of long setw#, those on the upper side being curved; dactyls slender, falciform, those of the first pair about one- third the length of the propodi, and a little more slender but not more curved than the others; in the two posterior pairs the dactyls are about one-half the length of the propodi. Abdomen of the male six-jointed from the fusion of the last two joints; first two joints visible from above; the first joint is much longer but not so wide as the second; the second, third, and fourth joints decrease successively in width but are of nearly equal length; the fifth is longer than the preceding and the sixth is oblong, rounded at the tip, and much longer than any of the others; the sides of the abdomen behind the second joint are concave and at the middle of each joint there is a conical protuberance, the last joint bearing traces of two. In the female the abdomen is large and rounded and composed of five joints from the fusion of the last three; it fits closely over the thin, laminate rim which encloses the greater portion of the sternum. As the sternum and the ventral surface of the abdomen. are both hollowed out, there is thus ‘formed a quite capacious chamber for holding the ova. Length of male 26 mm.; breadth 17 mm.; length of first ambulatory Jegs 60 mm. San Diego (Lockington); Santa Catalina Island, dredged August, 1893! Off Capedt. Lucas and Gulf of California (Miss Rathbun); Magdalena Bay, Lower California! I have seen Lockington’s specimens from the latter locality, and Miss Rathbun’s types of tenuipes from Catalina. Subfamily INACHIN/. Eyes slender and retractile. Basal antennal joint usually slender throughout, not narrowing distally. Preocular spine often wanting; postocular distinct. Rostrum often bifid or two-spined. CALIFORNIA STALK-EYED CRUSTACEA. 19 Genus Oregonia Dana. Carapace flattened, not spinose. Spines of rostrum long, slender, and contiguous. Ambulatory legs of moderate length, the penultimate joint similar to the preceding one, not dilated and compressed. Near Hurypodius. Type.—O. gracilis DANA. Oregonia gracilis Dana. Oregonia gracilis Dana, Am. Jour. Sci. (2), Vol. XI, 1851, p. 270; Crust. U.S. Expl. Expd., Part I, 1852, p. 106, Pl. IIT, fig. 2. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 456. Bats, in Lord’s Natural- ist in Vancouver’s Island, Vol. IL, 1866, p. 269. LockineTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 74. Smiru, Rept. Prog. Geol. Sur. Canada, 1878-9, B, p. 209. Nerwcomses, Bull. Nat. Hist. Soe. Brit. Col., 1893, p. 21. Rarusun M., Proc. U.S. Nat. Mus., Vol. XVII, 1894, p, 59. Oregonia hirta Dana, Am. Journ. Sci. (2), Vol. XI, 1851, p. 270; Crust. U.S. Expl. Expd., Part I, 1852, p. 107, Pl. III., fig.3. Srirmpson, l. c., p. 456. LocKkineTon, l.c., p. 75. WHITEAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. Oregonia longimana Bate, Proc. Zool. Soc. London, 1864, p. 663; Ann. Nat. Hist. (3), Vol. XV, 1865, p. 487; in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866. Carapace subtriangular, more or less setose and roughened by minute prominences. Median, cardiac, and branchial regions tumid. The rostral horns may exceed the breadth of the interorbital space. Postorbital spines slender, acute, inclined forwards, and situated some distance behind the orbit. A prominence on the anterior side of the eye peduncle. The septum separating the antennulary fossettes is produced intoa spine. Maxillipeds setose, the ischium produced forward into a rounded process at the antero-internal angle. Chelipeds rather slender, the merus sub- cylindrical and roughened with minute tubercles; carpus rounded; hand long, slender, somewhat compressed, the margins obtuse; fingers slender, smooth, incurved. Legs subcylindrical and decreasing in length poste- riorly; dactyls slightly exceeding one-half the length of the propodi and terminated by long, corneous claws. Behring Sea to Oregon (Rathbun); Vancouver Island (Bate, Smith); Puget Sound (Dana); northern California! Like many other maioid crabs, this species varies greatly as regards the development of the chelipeds in 20 CALIFORNIA ACADEMY OF SCIENCES. the male. Speaking of a collection of specimens of this species from Vancouver’s Island, Professor Smith says: ‘ “eh: a2 CALIFORNIA ACADEMY OF SCIENCES. Hyas lyratus Dana. Hyas lyratus Dana, Am. Journ. Sci. (2), Vol. XI, 1851, p. 268; Crust. U.S. Expl. Expd., Part I, 1852, p. 86, Pi. I, fig. 1. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 450. Lockx1neton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 64. Muers, Challenger Reports, Vol. XVII, 1886, p. 47. Ratusun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 72, Pl. III. Newcomss, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 20. Carapace lyrate and furnished with blunt tubercles. Median region tumid, and generally furnished with two elevations, one near the middle and one behind; post-median region prominent, pustulate or roughened; cardiac region furnished with a subconical elevation; branchial regions elevated at the middle and crossed by a more or less evident line of rounded tubercles. Rostrum shorter than the width of the interorbital space, the cleft between the horns narrow. A tooth on the anterior margin of the optic peduncle. Basal antennal joint with a few small teeth on each mar- gin and a prominent, smooth tubercle at the antero-external angle; second joint dilated and almost entirely visible from above. A small tooth on the first joint below the articulation of the second. Maxillipeds granu- lated. Behind the eyes is an alate expansion, the anterior and posterior margins of which are generally nearly parallel, outer margin long and a little concave; anterior angle acute; a small tooth near the base of the convex posterior margin of this expansion, behind which the side of the carapace is quite deeply indented. Behind the alate expansion the margin of the carapace is furnished with a row of small tubercles. Merus of the chelipeds with a few small tubercles, which are more prominent at the angles; hands narrow, compressed, smooth or granulated, the upper mar- gin compressed and roughened by small tubercles; fingers nearly smooth. The abdomen of the male is widest at the second and third segments, behind which the sides run nearly parallel to the last joint, which is dis- tally truncated and over twice as broad as long. Abdomen of the female broadly elliptical, the joints increasing in length from the second to the last. Locality of specimens described:— 56° 40’ N. 169° 20’ W. 43 fms. 2 spms. 56° 58’ N. 170°09 > W.. 2a 7 ta 2 56° 34’ N. LOA aw G2 re al This species ranges from the extreme end of the Aleutian Islands to Puget Sound, whence it was first reported by Dana. > CALIFORNIA STALK-EYED CRUSTACEA. 30 Genus Hyastenus White. Carapace subpyriform, convex, smooth, tuberculated, or spiny. Spines of rostrum long, slender, and diverging from the base. Preocular spine small or obsolete. Eyes small, the peduncles slender and completely retractile within the small orbits which have a lateral aspect, and a fissure, or hiatus in the upper@and lower margins. Basal antennal joint large; flagellum usually exposed and visible from above at the sides of the ros- trum. Ambulatory legs elongated, subcylindrical and unarmed, the first pair usually much the longest. Abdomen of the male seven-jointed. This genus is divided by Miers into two subgenera, which, however, ‘‘are connected together by numerous gradations.” The one, Hyastenus, is characterized by having few or no spines on the carapace and by the absence of a spine at the external angle of the basal antennal joint. The other, Chorilia, has the carapace spiny and the basal antennal joint is furnished with a spine. Hyastenus (Chorilia) longipes is the type of the latter subgenus. Hyastenus (Chorilia) longipes (Dana). Chorilia longipes Dana, Am. Journ. Sci. (2), Vol. XI, 1851, p. 269; Crust. U.S. Expl. Expd., Part I, 1852, p. 91, Pl. I, fig.5. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 455. Lockineton, Proc. Cal. Acad. Scei., Vol. VII, 1877, p. 69. Wurruaves, Can. Nat. (2), Vol: VIII, 1878, p. 470. Ratusun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 572. Hyastenus (Chorilia) longipes Miers, Journ. Linn. Soc. London, Vol. XIV, p. 658; Proc. Zool. Soc. London, 1879, p. 27. Hyastenus japonicus Miers, Proc. Zool. Soc. London, 1879, p. 27, Pl. I, fig. 2; Challenger Reports, Vol. XVII, 1886, p. 56, (fide RatHBuN M.). Hyastenus longipes Miers, Challenger Reports, Vol. XVII, p. 56. Ratusun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 85, Pl. VII. NEewcomse, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 20. Carapace pyriform, inflated, and covered with sharp spines of unequal size. Median region tumid, with two median spines and a row of three spines on either side, the posterior one being the largest and situated a little behind. the middle of the interval between the two spines on the middle line; a prominent spine on the tumid hepatic region; several small 3 o4 CALIFORNIA ACADEMY OF SCIENCES. spines on the cardiac region and a small spine or tubercle near the pos- terior end of the intestinal. The spines on the branchial regions are numerous and variable, but there is usually a more prominent spine near the middle. Pterygostomian regions with a row of teeth or spines. Ros- trum long, the horns nearly straight, pubescent and divergent. Preorbital spine prominent, acute, postorbital acute and pointing obliquely down- wards. Basal antennal joint armed at its exterffal angle with a slender spine, the margin behind which is armed with two smaller spines. There may be a few minute spinules at the apex of the eye-stalk. Merus of the chelipeds furnished with small tubercles, which are more prominent on the angles; carpus furnished with small tubercles on the outer side, which are more or less plainly arranged in three or four rows; hands long, slen- der, compressed, palm subcarinated above, nearly smooth, and generally having a small tubercle on the outer side near the articulation; fingers long and slender. Dactyls of the slender ambulatory legs long and nearly straight. Abdomen of the male tapering from the third segment, the last joint longer than broad and rounded at the tip. Abdomen of the female broadly elliptical, the joints increasing in length from the third to the last. Insome females, either immature or sterile, the abdomen is narrowly elliptical and not nearly so wide as the thoracic sternum. The legs and many parts of the body are covered with a short pubescence. Miss Rathbun, who has carefully studied a large number of specimens of this species, gives the following account of its variations: ‘‘ This species ranges from 57° north latitude, off Kadiac, Alaska, to 32° north latitude, off San Diego, Calif., and in depth from 27 to 603. fathoms. It exhibits wide variations from Dana’s types, especially in more southern latitudes, where, as a rule, the carapace is very much swollen at the branchial regions, making the width much greater in proportion to the length; the second and third joints of the antenne are much more slender; the hepatic region is furnished with a sharp spine; and, lastly, the tubercles of the carapace are much more numerous and some of them spinous. These characteristics, if uniform, would be specific, but the two extremes intergrade to such an extent as to render impossible even a varietal separa- tion. The broad form is, with one exception, confined CALIFORNIA STALK-EYED CRUSTACEA. 35 to deep water; the typical longipes ranges from 27 fathoms in the north to 456 in the south. Variations exist In specimens from the same locality; for example: The broad forms may possess a hepatic spine or a tubercle; the antennal joints are narrow in some indi- viduals and wide in others. Occasional specimens of the narrow form have asharp hepatic spine. * * * The width of the typical form ranges from 0.71 to 0.8 of its length; of the wider form from 0.82 to 0.9 of its length; the length being measured from between the bases of the cornua.”’ | Genus Pelia Bell. Carapace subpyriform. Rostrum composed of two divergent spines which are united at the base. Orbits small, with a lateral aspect and with a superior and an inferior marginal hiatus; the upper orbital margin smooth and devoid of a preorbital spine. Eyes small, with slender pedun- cles. Basal antennal joint elongated, its distal portion visible from above at the sides of the rostrum; flagellum well developed. Merus of the max- illipeds with a notch at the articulation of the palp, legs of moderate length; first ambulatory legs much longer than the others; merus com- pressed, acute above, dactyls not denticulated below. Type.—P. pulchella BEuL. Near Pisoides, but having a narrower basal antennal joint, the distal end of which is visible from above. Differs from Pisa in the absence of spines on the under side of the dactyls. Pelia tumida (Lock.) Pisoides (?) tumidus Locx1neTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, pp. 30 and 67. Microphrys tumidus Miers, Chailenger Reports, Vol. XVII, 1886, p. 83. Carapace pyriform, rounded, tumid, covered with pubescence, but en- tirely devoid of spines. Median region rounded, smooth, much elevated, and furnished with a small rounded tubercle at the summit; branchial regions inflated; cardiac region with a small rounded elevation. Rostrum depressed, nearly one-half the length of the carapace and bifurcated for 36 CALIFORNIA ACADEMY OF SCIENCES. about half its length, the horns narrow, divergent, and slightly upturned at the tip. No preorbital tooth; postorbital small. Antero-lateral margin entire. Basal antennal joint considerably longer than wide and devoid of teeth with the exception of the one at the antero-external angle, which is plainly visible from above; tip of the peduncle reaching but beyond the apex of the notch between the rostral horns, not nearly reaching the tip of the rostrum as in pacifica; flagellum rather long. Merus of the maxil- lipeds distally truncated. Chelipeds unarmed; merus not nearly reaching the tip of the rostrum; hand oblong, inflated, the edges obtuse and paral- lel; fingers widely gaping, a tubercle on the inner margin of the dactyl near the base. Ambulatory legs compressed, pubescent, the margins fur- nished with a thick fringe of stiff setz; dactyls sharply curved at their sharp, corneous tips. Length 12 mm.; width 8 mm.; length of first ambulatory leg 12 mm. Described from a single male specimen from Magda- lena Bay, Lower California, preserved in the museum of the California Academy of Sciences. This specimen is, I believe, one that is mentioned by Lockington, l. c., peOv: San Diego (Lockington), Magdalena Bay, Lower Cali- fornia (Lockington)! Santa Catalina Island! Pelia pacifica Milne-Hdw. Pelia pacifica A. MILNE-EpwaRps, Miss. Sci. au Mex., Vol. I, Pt. V, 1875, p. 73, Pl. XV1, fig. 3. Ratusun M., Proc. U.S. Nat. Mus., Vol. XVI, 1894, p. 90; and Vol. XXI, 1898, p. 573. Near mutica, but having a shorter and wider carapace with the promi- nences on the gastric and cardiac regions more pronounced. Rostrum short and stout. Basal antennal joint shorter and wider than in mutica. Length of carapace 9 mm.; width 7 mm. Bay of Panama (M.-Edw.); Gulf of California, Mag- dalena Bay, Lower California! San Diego, Santa Cata- lina Island (Miss Rathbun). Genus Herbstia Milne-Edw. Carapace broadly triangular or subpyriform, tuberculated or spinose. Rostrum short, the horns acute, vertically compressed, and dilated at the base. Orbits shallow, with or without a preorbital spine. Eyes short and CALIFORNIA STALK-EYED CRUSTACEA. ot not entirely concealed when retracted. Basal antennal joint moderately dilated and armed with an antero-external spine, the distal portion not entirely covered by the rostrum. Merus of maxillipeds distally truncated and not produced at the outer angle. Ambulatory legs rather slender, subcylindrical, and of moderate length; dactyls nearly straight, acute. Abdomen in the male seven-jointed. This genus is divided into two subgenera as follows: Herbstia: Inferior margin of orbit not dentate. Ambulatory legs not spinose. Herbstiella St.: Inferior margin of orbit armed with a tooth or spine. Merus of ambulatory legs spinose. The subgenus Herbstiella to which our species belong is almost exclusively West American. Herbstia (Herbstiella) camptacantha St. Herbstia parvifrons Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 185 (not A. parvifrons RaNnDALL). Herbstiella camptacantha Stimpson, 1. c., Vol. X, 1871, p. 94. Herbstia camptacantha A. MILNE-EDWARDS, Miss. Sci. au Mex., Vol. I, Pt. Metig, 2a. XVI fio.8. Herbstia ( Herbstiella) camptacantha Mirrs, Journ. Linn. Soc. London, Vol. XIV, 1879, p. 655; Challenger Reports, Vol. XVII, 1886, p. 49. RATHBUN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 79: Fisheria depressa LOockINeTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 72. Carapace ovate, punctate, much flattened, and armed above with several small tubercles. Median region tumid, separated from the branchial and cardiac regions by deep sulci and armed, in front, with four tubercles in a transverse line, the middle tubercles separated from each other by about twice the distance that they are from the lateral ones; a median, rounded tubercle on the posterior portion of the median area; three, or four small tubercles on the cardiac region and about five on each branchial region; two tubercles, in a transverse line, on the intestinal area. Rostral horns flattened and divergent, forming more than one-half the length of the rostrum. Preorbital spine acute; there is a small spine, or tooth, on the margin of the orbit above the postorbital tooth, while below it there is a spine on the inferior orbital margin. The spine at the antero-external angle of the basal antennal joint is prominent, and on the margin behind it there are two spines, the posterior of which is sometimes reduced to a small, blunt tooth. Beside the postorbital tooth the antero-lateral mar- gin is furnished with about five spines, and there are several smaller ones 38 CALIFORNIA ACADEMY OF SCIENCES. on the postero-lateral margin. Pterygostomian region armed with several spines. Chelipeds in the adult male long; merus long, subcylindrical, armed above with numerous spines and usually having a few spines at the distal end of the lower margin; carpus with the outer surface tuberculated; hand large, smooth, elongated and compressed; fingers widely gaping. Ambulatory legs pubescent; merus armed above with numerous spines, and with one or more spines near the distal end of the lower margin; carpus with one or more spines above; propodi long, slender, subcylin- drical, unarmed, and about twice the length of the dactyls. Cape St. Lucas, Acapulco, Mexico (St.); Mazatlan, Magdalena Bay! Lower California; San Diego! Santa Catalina Island! San Pedro! San Clemente Island! This species differs from parvifrons in having the hand entirely devoid of spines. The chelipeds in the male become greatly elongated with age; the hand becomes narrower, the spines on the lower side of the merus become obsolete, while those on the upper side become blunt. In one specimen from Port Escondido the chelipeds were two and one-half times the length of the carapace; the merus was a little longer than the carapace and the hand about one-third longer. The carapace is covered with a pubescence which disappears to a greater or less extent with age. I have seen the two specimens of Fisheria depressa referred to by Lockington as ‘‘ No. 21” (1. ¢., p. 73). Herbstia parvifrons Randall. Herbstia parvifrons RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 107. GrpBEs, Proc. Am. Ass. Adv. Sci., 1850, p. 170. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 451. I have not seen this species nor do I believe that any- thing has been found since Randall’s specimen was taken which answers to his description, which is quoted below. ‘‘ Corpore depresso, in latere spinoso, pedibus spinosis, leviter dilatatis, manibus levigatibus, dactylis apice serrulatis.” CALIFORNIA STALK-EYED CRUSTACEA. 39 ‘‘ Body flattened above, punctured, somewhat hairy; sides spinous, especially anteriorly; rostrum very short; feet spinous above; hand of the anterior pair with only a few very small spines at the base; pincers with- out teeth, except toward their apices, where for one-third of their whole length they are hollowed out on their prehensile side, so that when closed they become interlocked at their extremities by means of their serratures. Length of carapax about one inch.” One specimen from Western America. Genus Chionecetes Kroyer. Carapace broad, depressed, more or less tuberculated or spinose. Ros- trum short, flattened and notched, not depressed. Orbits shailow, open above so that the short, thick eye-peduncles are visible from above when retracted. Nopreorbital spine; postorbital present. Basal antennal joint very narrow, with a terminal spine; second and third joints not dilated; flagellum short. Ambulatory legs more or less compressed and of moderate length. Abdomen seven-jointed. Type.—C. opilio (FABR.). Chionecetes opilio (Fabr.) Cancer phalangium O. Fapricius, Fauna Groenlandica, 1780, p. 234. Cancer opilioO. Fasricius, Danske Vid. Selsk. Skr. nye Saml., Band. ITI, 1788, p. 181, plate. Chionecetes opilio KR6YER, Naturh. Tidskrift (1), 2, 1838, p. 249. Chionecetes behringianus STIMPSON, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1856, p. 84; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 449. For further references see Rathbun; Proc: U.S. Nat. Mus., Vol. XVI, 1893, p. 74. This species extends from Greenland to Behring Sea and down the west coast of America as far as British Columbia, and may, not improbably, occur within our limits. The carapace is flattened and covered with wart-like tubercles. There is no deep depression sepa- rating the branchial regions. The ambulatory legs are shorter, stouter and less spiny than in the next species. 40 CALIFORNIA ACADEMY OF SCIENCES. Chionecetes Tanneri Rath. Chionecetes Tanneri RatHBUN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 76, Pl. IV, figs. 1-4; and Vol. XXI, 1898, p.573. NeEwcomss, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 19. Carapace a little wider than long, with the postero-lateral angles broadly rounded; branchial regions tumid and produced laterally, concealing the lateral margins. The carapace is covered with spines, instead of wart-like tubercles as in opilio, the most conspicuous of which are arranged in more or less irregular rows. A nearly transverse row of tubercles on the branchial regions, which, near the side of the carapace, curves around and runs obliquely forwards towards the orbit. Posterior margin of the carapace spinulous; above the outer portions.of this margin there begins another row of spinules, which is continued around the sides of the cara- pace, the spines becoming smaller anteriorly; at the bend in the row of spines on the branchial regions there begins another row of spines which runs forward to the anterior portion of the buccal area, the spines becom- ing smaller and more thickly set anteriorly, where they gradually spread out into a small patch of minute, irregularly placed tubercles next to the buccal area. An irregular row of spinules across the anterior portion of the median area. The upper margins of the orbit and the outer sides of the rostrum are spinulous. A small elevated patch of spinules on the cardiac region, in front of which is a deep depression, separating the branchial regions, which is rather narrow behind, but widens anteriorly, and divides into two broad depressions extending either side of the median area. Rostral horns upturned, narrower than in opilio, the notch between them more prominent. Basal antennal joint spinulous below. Maxillipeds more or less spinulous, and not reaching the anterior margin of the buccal area. Chelipeds spinulous, small in the female; hand nar- — row; fingers long. Legs spiny, becoming smoother toward the tips. A transverse spinulous or granulated ridge in front of the abdomen, in front of which the sternum is excavated. First two or three joints of the abdomen with a transverse spinulous or granulated ridge. The largest specimen in the series studied by Miss Rathbun gave the following measurements: Length from baseot rostral horned. we ee ae eee 119 mm. Width without spines....... MEARE ys CE Res Erne te ecienesene 130 mm. Iuenpth of sfirst.ambulatory ler ..'. 2 sak fees ee ee ae 316 mm. Length of merus of first ambulatory leg............... 134 mm. Miss Rathbun’s table of measurements of 18 specimens shows that in the smaller specimens the length more i al i arte Be et CALIFORNIA STALK-EYED CRUSTACEA. 41 nearly equals the breadth than in the larger ones. In one case the length and breadth were each 32 mm., but in no case was a specimen longer than broad. Deep water from Behring Sea to southern California; 29 to 1,625 fathoms (Miss Rathbun). This and preceding species seen and compared. Genus Scyra Dana. Carapace subpyriform, tuberculated, but not spinose. Rostrum com- posed of two flattened horns. Orbits small, with a fissure above and below, the lower and sometimes the upper one being open. Preorbital spine present. Basal antennal joint rather narrow, with a small spine at the antero-external angle, the two following joints compressed and not concealed by the rostrum. Merus of the maxillipeds distally truncated. Chelipeds in the male well developed, the hand compressed and carinated above; fingers acute. Legs moderately long, subcylindrical, the first pair not greatly exceeding the others in length; dactyls short and acute. Abdomen seven-jointed. Type.—S. acutifrons DANA. Scyra acutifrons Dana. Scyra acutifrons DANA, Am. Journ.-Sci. (2), Vol. XI, 1851, p. 269; Crust. U.S. Expl. Expd., Part I, 1852, p. 95, Pl. II, fig.2. . Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 455. LocxineTton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 69. Mrers, Journ. Linn. Soc. London, Vol. XIV, 1879, p. 663; Challenger Reports, Vol. XVII, 1886, p. 62. Situ, Rep. Prog. Geol. Surv. Canada, 1878-9, B, p. 210. Ratusun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 88. NErwcomBs, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 21. Scyra (sp. undet.) WHITEAVES, Can. Nat. (2), Vol. VIII, p. 471; (see Smith jas Carapace pyriform and furnished with rounded tubercles. Median region tumid and separated from the cardiac and branchial regions by a conspicuous depression; an acute tubercle near the center of the median region, behind which is a larger and more obtuse tubercle. Branchial regions tumid and bearing a large, projecting tubercle, in front of which is an elevated prominence which may bear several small tubercles, though it is often smooth. A large tubercle on the cardiac and a smaller one on the intestinal region. Rostrum short, the horns ovate-lanceolate. Two A2 CALIFORNIA ACADEMY OF SCIENCES. spines or teeth on the outer margin of the basal antennal joint behind the one at the anterior angle. Preorbital spine acute. Pterygostomian regions with several rounded teeth. Chelipeds of the male large, the merus subcylindrical (somewhat flattened below), and strongly pustulate, especially at the angles; the carpus is pustulate and may have several ridges on the outer side; hand long, narrow, compressed, the palm below the wide carina often inflated; fingers deflexed and, in old males, gaping at the base, in which case there is generally a large tooth near the base of the dactyl. Legs subcylindrical, more or less pubescent, the propodi sulcate on either side; dactyls considerably shorter than the propodi and furnished with sharp, corneous tips. The third, fourth, fifth and sixth abdominal segments in the male are of nearly equal length; the seventh is longer than broad and rounded at the tip; the abdomen tapers most at the third and fourth segments, the fifth and sixth being of subequal width. Length of iearapace:ignile sea cere tae eee eee here 30 mm. Width of carapacey 0 Wave a et Ra ee Meine, rae ee 26 mm. Leneth otcchelspedgy;) Wie rev seb Gaam oo aero e eee 55 mm. Lengthiof first ambulatorydles iy 3 2. nde oe nee 40 mm. Teneth*ot carapace: female. wa. nee sees oe eee 25 mm. Width of carapacesy: 3° *o. her eas eee eee 16mm. hénoth of chelipediay T= Os VOIR ase Oa ae eee ae Races 28 mm. Alaska, British Columbia, Oregon, Farallon Islands! southern California! Santa Catalina Island! San Diego! The individuals of this species vary considerably with age. In the old males the tubercles on the carapace are rougher and more prominent, the posterior tubercle on the branchial region projecting over the sides of the carapace; the horns of the rostrum become much widened at the base; and the chelipeds are much larger. In the females the tubercles on the carapace are smoother, those on the median region being small er obsolescent. Family PERICERIDA. Eyes retractile within the small, circular, and well defined orbits which are never incomplete as in the Maiid#w. Basal antennal joint well devel- oped, and constituting the greater part of the inferior wall of the orbit. CALIFORNIA STALK-EYED CRUSTACEA. 43 Subfamily PERICERIN2. Carapace more or less triangular. Rostrum well developed. Second _ joint of the antenna not dilated. Chelipeds with the fingers acute at the tips. Genus Libinia Leach. Carapace orbiculate-triangular, convex, spinose. Preocular spine dis- tinct. Rostrum of moderate length and emarginate only at the apex. Eye-peduncles short. Basal antennal joint moderately enlarged, the flagellum visible from above. Legs of moderate length. Abdomen of male and female seven-jointed. Type.—L. emarginata LEACH. Libinia emarginata Leach. Libinia emarginata Leacu, Zool. Mise. (2), 1815, p. 129. Ratusun R., The Fisheries of the U.S., Sec. I, 1884, p. 778, Pl. CCLXIX, fig. 4. ‘RatHeun M., Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 235, Pl. XXXI, fig. 2.. ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894. Libinia canalaculata Say, Journ. Acad. Nat. Sci. Phila., Vol. I, 1817, p. 76. Mitne-Epwarps, Hist. Nat. Crust., Vol. I, 1834, p. 300. STREETS, Proc. Acad. Nat. Sci. Phila., Vol. XXII, 1870, p. 105. Libinia afinis RaNDAbL, Journ. Acad. Nat. Sei. Phila., Vol. VIII, 1839, p. 106. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 455. STREETS, 1. c. For further references, see Rathbun M., /. c. Carapace suborbicnlar, much inflated, and covered with numerous smali, pointed tubercles. Median region with four or five median tuber- _ cles, besides many others. Hepatic regions not distended, and furnished with three quite prominent spines, the upper one forming the first of a series of five spines (or tubercles) which extends backward upon the branchial regions. A row of five tubercles behind the median region, making nine or ten median tubercles in all. Rostrum short, triangular, the apex furnished witha very small emargination. Length of male specimen 95 mm.; breadth 90 mm.; length of cheliped 160 mm.; of hand 84 mm.; of first ambulatory leg 158 mm.; of last leg 98 mm. Atlantic Coast from Maine to Florida; ‘‘ Western America” (Randall). The foregoing description is based on specimens from Wood’s Holl, Massachusetts. I have seen no specimens 44 CALIFORNIA ACADEMY OF SCIENCES. of this species from the Pacific Coast, and know of none having been found there since Randall’s specimen was taken. Mr. Streets, whose words imply that he has seen Randall’s type, says that this species ‘‘is undoubtedly nothing more than the young of canaliculata. That itis so will be evident to anyone who will take the trouble to compare them closely. It agrees with L. canaliculata in every respect excepting size.”’ Legion PARTHENOPINEA ‘¢ Basal antennal joint very small and imbedded with the next joint in the narrow hiatus between the front and the inner suborbital angle; the infra-ocular space being mainly occupied by the lower wall of the orbit.” Family PARTHENOPID&. Characters of the legion of which it is the only family. The species of this family are often markedly triangular in form; the carapace is usually angular and provided with conspicuous depressions separating the regions. In the characters of the antennz and also in several other points, the Parthenopidz approach the Cancroid Brachyura. Genus Heterocrypta St. Carapace triangular, transverse, the lateral margins greatly produced and concealing the ambulatory legs. A conspicuous depression separating the gastric from the cardiac and branchial regions. Rostrum simple. A strongly marked ridge on the pterygostomian regions. Chelipeds greatly developed, trigonal. Posterior margin of the carapace not produced, as in Cryptopodia. : fp Heterocrypta occidentalis (Dana). Cryptopodia occidentalis Dana, Am. Journ. Sci. (2), Vol. XVIII, 1854, p. 430, fig. in text. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 458. GrBBEs, Proc. Elliott Soc. Nat. Hist., Charleston, §.C., Vol. I, 1859, p. 36. A. M1~NE-Epwarps, Miss. Sci. au Mex., Vol. I, Ptsov,.p. 169. Lambrus frons-acutis Lock1neton, Proc. Cal. Acad. Sci., Vol. VII, 1877, D: GOL: CALIFORNIA STALK-EYED CRUSTACEA. 45 Carapace broadly triangular; rostrum triangular, subacute, not depressed. Median region narrow, the flattened upper surface bounded by two granu- lated ridges, which converge to a point behind. Cardiac region furnished with a three-sided, pyramidal elevation, the edges of which are usually granulated. Postero-lateral regions large, and furnished with an §-shaped, granulated crest, which extends from near the posterior end of the median region to the acute lateral angles of the carapace; in front of the anterior bend of this crest there is a pair of minute tubercles. Antero-lateral mar- gin straight or slightly concave in front, convex near the middle, the posterior portion passing outwards and backwards, arching over the legs; the teeth on the anterior part are small and irregular, but they become larger posteriorly, where they are furnished with secondary denticles. Postero-lateral margins transverse; posterior margin not produced over the abdominal segments. Outer portion of the orbit with a superior and an inferior fissure. Ischium of the maxillipeds smooth, the antero-in- ternal angle produced; merus small, the surface concave and bearing near the middle a prominent tooth. Chelipeds long, trigonal; the sides of the merus convex, the edges sharply granulate to dentate; carpus with three or four granular lines; hand about as long as the merus, the angles prominent and dentate and the sides concave; pollex short, deflexed; dactyl short, but longer than the pollex, and when closed its outer margin is nearly at right angies to the long axis of the hand. Ambulatory legs compressed, strongly carinated above; dactyls narrow, strongly sulcate, and corneous-tipped. Color reddish. Length 18 mm.; breadth 28 mm.; length of cheliped 50 mm. Monterey (Dana) ; Santa Catalina Island (Lock.)! This species is quite common in Avalon Harbor, Santa Catalina Island. I have examined a specimen in the California Academy of Sciences which I feel confident is Lockington’s type of frons-acutis. Genus Leiolambrus Milne-Hdw. Carapace subtriangular, smooth, with no tubercles or spines on the upper surface. The median and cardiac regions are not separated from each other by a deep depression, but are limited on either side by longi- tudinal sulci. Front short, truncated, and armed witha small, median tooth. The antero-lateral margins are denticulated and the posterior margin is furnished with teeth or spines. Lateral angles of the carapace prominent and acute. Orbits well defined and furnished with a fissure in the upper and lower margins. Antennules obliquely plicated. Ischium of the maxillipeds produced forward at the antero-internal angle; outer \ 46 CALIFORNIA ACADEMY OF SCIENCES. angle of the merus rounded. A ridge on the pterygostomian regions. Chelipeds trigonal, the angles dentate. Ambulatory legs slender, com- pressed; dactyls slender, depressed. Type.—L. punctatissimus (OWEN). ‘Leiolambrus punctatissimus (Owen). Parthenope punctatissima Owrn, Zool. Beechy’s Voyage, Orust., 1839, p. 81, Pl. XXIV, fig. 4. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 458. Lockineton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 78. Leiolambrus punctatissimus A. MILNE-Epwarps, Miss. Sci. au Mex., Vol. I, Pt. V, pp. 148 and 159. Carapace smooth, convex, minutely punctate; the median and cardiac regions forming a nearly continuous longitudinal elevation, bounded on either side by a conspicuous, longitudinal depression. Front truncated, having a small, median tooth, behind which the upper surface is marked with a short, longitudinal groove. Postorbital angle acute. Antero-lateral margins Tong, slightly arcuated, and furnished with teeth which become very small toward the anterior end. The sides of the carapace are pro- duced into a prominent, triangular tooth. Two teeth on the posterior mar- gin near the middle line; external to these a pair of larger marginal teeth; the margins between the external pair and the lateral angles of the carapace are concave. The whole posterior margin of the carapace is marked with a delicate, raised, jagged line; another fine raised line running upon the branchial regions from the external tooth of the posterior margin. Max- illipeds smooth, the ischium slightly narrowed distally, the antero-internal angle produced forward into a rounded lobe; merus broadly rounded at the antero-external angle and produced below the articulation of the palp. Chelipeds large; merus with the angles denticulated, the anterior edge bearing several rough teeth; carpus with the angles denticulated, the outer margin acuteand dentate; hand longer than the merus, the margins denticulated, the upper margin upturned near the end; pollex nearly longitudinal and almost straight, the inner margin dentate; dactyl curved and provided on the external portion of the base with two converging, denticulated ridges which meet and form a crest on the distal portion. Ambulatory legs smooth, compressed, and subequal in length; dactyls slender, lanceolate, longer than the propodi, and flattened in a plane at right angles to that of the preceding joints. Locality of specimen described, lat. 27° 45’ N., lon. 110° 45’ W.; 20 fathoms. SMEEN OF COEAPBOC:. viecis. val odieees 6 tem tae Ce Oe eee 7.25 mm. WV Gad REL CATA IAGO...) 5. ciecs s-s'% tes p bo ms Soe ae aed ete eee ee Ca 9.5 mm, NGO Oia AAD MRI. Bly sp cere's cscs as caves acer eas ota 7 mm CALIFORNIA STALK-EYED CRUSTACEA. 47 Subtribe CYCLOMETOPA, or CANCROIDEA. Carapace generally broader than long, wide and regularly arcuated in front, rarely quadrate or suborbicular in form; rostrum wanting. LEpi- stome short, transverse. Antennules longitudinal or transverse. Palp of the maxillipeds joined to the apex or inner angle of the merus. Branchiz nine, the efferent branchial channels as in the Maioidea. Verges of the- male in the coxe. Section CANCRINE. Ortmann. Carapace rounded or widened, a dentate antero-lateral margin sharply marked off from a postero-lateral margin devoid of teeth. Front with several teeth, one of which is median. -Antennules longitudinal or oblique. Genus Cancer JL. Carapace transverse, subelliptical, indistinctly areolated. Antero-lateral: margins regularly arcuated and armed with ten teeth. Front narrow, cut into five teeth or lobes. Eye-peduncles short; orbits small, with two. fissures in both upper and lower margins. Antennules longitudinal, or nearly so. Basal antennal joint somewhat enlarged and united with the front, thus excluding the short flagellum from the orbit. Merus of the. maxillipeds distally truncated and not produced at the antero-external angle. Chelipeds subequal, the hand generally costate on the outer surface.. KEY TO THE SPECIES. Carapace not undulated; size small.................. C. gracilis. Carapace undulated, large species. Autero-lateral teeth distant, the posterior one the RCS Une Fae ee Oe re ros 2 ate! NO eee cae oes C. magister. Antero-lateral teeth contiguous. Front flattened and produced............. C. productus. Front not flattened nor produced........ C. antennarius. Cancer productus Randall. Cancer productus RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839,. p.116. Dana, Crust. U.S. Nxpl. Expd:, Part I.’ 1852, p. 156, Pl. VII... fig. 3. Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 461. Bats, in Lord’s Nat. in Vancouver’s Island, Vol. II, 1866, p. 269. Lockineron, Proc. Cal. Acad. Sci , Vol. VII, 1877, p. 95. Smiru, Rep. Prog. Geol. Sur. Can- ada, 1879, B, p. 207. Ratrupun R., The Fisheries of the U.S., Sec.. 1, 1884, p. 771, Pl. CCLXII. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. VIL, 1894, p. 425. Canman, Ann. N. Y. Acad. Sci., Vol. XI,. 1898, p. 262. Platycarcinus productus GiBBEs, Proc. Am. Ass. Adv. Sci., 1850, p. 177.. Cancer perlatus Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88. 48 CALIFORNIA ACADEMY OF SCIENCES. Carapace usually wide, undulated above. Hepatic regions depressed. Front flattened, produced into five, generally obtuse teeth which lie in the same horizontal plane, the outermost pair not more widely separated from — the inner pair than these are from the median tooth. Superior margin of the orbit between the two fissures not produced into a tooth, as in anten- narius; external orbital tooth small, the next one behind it rounded; suc- ceeding teeth becoming larger and more acute posteriorly, the last tooth, in adult specimens, being the largest. Between the teeth the margin of the carapace is marked with short, closed: fissures. The tooth on the lower margin of the orbit small or absent. Flagellum of the antennules considerably shorter than the width of the front. Merus of the maxilli- peds obliquely truncated at the distal end and notched on the inner side at the articulation of the palp. Carpus of the chelipeds with a large tooth at the antero-internal angle and a smaller one behind the upper hinge joint; the outer surface is roughened with several irregular prominences; hand with a double row of tubercle-like teeth on the upper edge and four or five granuiated costz on the outer surface, the upper ones of which are the most obscure; fingers stout and tipped with black, the dactyl not cristate above. Ambulatory legs nearly naked, except the dactyls which are slender and a little longer than the propodi. Terminal abdominal segment in the female nearly equilaterally triangular, the sides concave. In the male this segment is more narrowly triangular than in the other sex. Color dark reddish above; below a dirty white. hength of large’specimen, males. =. 0.3.5 .,...-2..08s one 85 mm. Breadth of large specimen, male..............00.00 +e: 131 mm. Length of small specimen, female .................... 22.5 mm. Breadth of small specimen, female.................... 33.5 mm. Dr. Stimpson has given a table of measurements of specimens of different sizes and sex, showing that the males are wider than the females and the old males rel- atively wider than the young ones. The proportion of length to breadth was found to vary from 1:1.30 ina male .99 in. in length to 1:1.63 in a male of 2.95 in. in length. The young of this species differ in many respects from the old specimens. The carapace is much flatter and smoother, and ornamented above with numerous longitudinal, colored lines which entirely dis- appear in the adult. The teeth of the front are repre- sented by five subequal crenulations. The teeth on the antero-lateral margin are much less distinct, being CALIFORNIA STALK-EYED CRUSTACEA. 49 represented on the anterior portion by low crenulations which become more acute behind. In the adult the posterior tooth may curve forwards but not so sharply as in antennarius. Secondary denticles are nearly obsolete. The granules on the carapace are of unequal size, being generally larger and sharper on the median region. In the females the carapace is generally more convex than in the males. Queen Charlotte Island (Smith), Vancouver Island (Bate), Puget Sound (Dana), Oregon. Common on the entire California coast to Lower California. Cancer antennarius St. Cancer antennarius STtTIMPSON, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 462, Pl. XVIII. Locxk- INGTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 96. Smitu, Rep. Prog. Geol. Sur. Canada, 1879, B, p. 207. Ratusun R., The Fish- eries of the U.S., Sec. 1, 1884, p. 771, Pl. CCLXIII. Misrs, Challenger Reports, Vol. XVII, 1886. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894, p. 425.— Platycarcinus recurvidens Bats, Proc. Zool. Soc. London, 1864; Ann. Nat. Hist. (3), Vol. XV, p. 488; in Lord’s Nat. in Vancouver’s Island, Vol. II, 1866. Carapace undulated, smooth, widest at the penultimate antero-lateral tooth, the granulations on the surface minute and even. Front not pro- duced, the three median teeth separated from the outermost pair which is the largest. The middle tooth is smaller than those on either side and situated a little below them. A tooth on the superior margin of the orbit between the two fissures. The tooth on the basal antennal joint projects further forward than the lateral tooth of the front and there is quite a prominent tooth on the inferior orbital margin just external toit. The postorbital is the smallest of the antero-lateral teeth but is relatively much larger than in productus. The teeth of the antero-lateral margin are large and have the tips sharp (except when worn) and curved forward; on the margins of the teeth, especially the anterior margin, there are usually small, sharp, secondary denticles. There are but nine large antero-lateral teeth, the tenth being probably represented by a small projection behind the last large tooth. Flagellum of the antennz usually long, generally 4 — 50 CALIFORNIA ACADEMY OF SCIENCES. exceeding the width of the front. Distal margin of the merus of the max- illipeds nearly transverse and gently convex; inner margin notched. Chel- ipeds stout; carpus with a stout tooth at the antero-internal angle and a smaller tooth behind the superior hinge joint; the external surface is smooth in old specimens but roughened, granulated and pubescent in younger ones. The hand is longitudinally costate on the external surface in young specimens but the costxe disappear with age; the fingers are smooth and tipped with black. Ambulatory legs very hairy or nearly naked; the dactyls pubescent and a little stouter than in productus. Length of carapace, male..... OPO A PETAR A me Pdr Fe ot 76 | mm, Breadth of carapace, male, A. ek cee «eee ene 113.5 mm. Leneth of carapace; Temale.o vis, co. eee ere ee eee ve Tane Breadth of ‘carapace; femaletit2. gL ishla. eee ae 107. mm. The young of this species are relatively much less wide than the adults and have the carapace and append- ages thickly covered with hair. The hair on the upper surface disappears with age and in old specimens the appendages may become nearly naked. The postorbital tooth in young specimens is fully as large as those be- hind it. This is a well marked species and may easily be distinguished from the other species on our coast by the sharply recurved antero-lateral teeth. Queen Charlotte Island (Smith), Vancouver Island, Oregon, northern to Lower California! Common. Cancer magister Dana. Cancer irroratus RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 116. Cancer magister Dana, Proc. Acad. Nat. Sci. Phila., Vol. VI, 1852, p. 78; Crust. U.S. Expl. Expd., Part I, 1852, p. 151, Pl. VII, fig. 1. Srrmp- son, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 458. Coopsmr, Rep. Expl. Sur. Pac. Ocean, Vol. XII, 1860, p. 387. Locxineton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 94. Smita, Rep. Prog. Geol. Sur. Canada, 1878-9, B., p. 206. Ratusun R., The Fisheries of the U.S., Sec. 1, 1884, p. 770, Pl. CCLXI. Metacarcinus magister A. Mitnz-Epwarps, Ann. Sci. Nat. (4), T. XVIII, 1862, p. 33, and (5), T. I, 1864, p. 67; Nouv. Archiv. Mus. Hist. Nat. Paris, T.I, 1865, p.202, Pl. XIX, fig. 1. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894, p. 424. CALIFORNIA STALK-EYED CRUSTACEA. 51 Carapace gently convex, undulated, and covered with small granulations which are larger on the summits of the prominences. Two or three curved rows of light colored spots on the antero-lateral regions. Front not produced, the three median teeth small, the middle one being slightly the largest and projecting further forward than the others; outermost pair larger than the others, not reaching so far forwards, and separated from them by a considerable interval. No tooth on the superior orbital margin. Basal antennal joint large, its pointed extremity projecting much further forward than the outer tooth of the front. A small tooth on the inferior margin of the orbit just external to the basal antennal joint. Antero- lateral margin armed with ten distant teeth which decrease in size and become more closely set anteriorly; the teeth are triangular in shape and the margin between them is denticulated or sharply granulated; the pos- terior tooth is the largest of the series, its anterior margin is generally transverse and its posterior margin continuous with the postero-lateral margin of the carapace. Postero-lateral margin nearly straight (the anterior part slightly convex, the posterior a little concave) and devoid of a tooth or projection behind the large lateral tooth. Merus of the maxilli- peds very obliquely truncated at the distal end, the antero-internal angle rounded, and the inner margin concave (not deeply notched) behind the articulation of the palp. Chelipeds slightly unequal; merus armed with two teeth at the distal end, one on the margin, and one a short distance behind it; the outer surface of the carpus furnished with granulated ridges, the upper one running to a sharp spine at the antero-internal angle; upper surface of the hand furnished with a prominent crest, which is cut into numerous sharp teeth; outer surface with five longitudinal, granulated costz; fingers more slender than in the preceding species and furnished with more numerous teeth; dactyl armed with a dentate crest on the upper margin. Legs granulated, carpi sulcate above; dactyls strongly compressed, those of the fifth pair widened. Beesth of adult specimen... 223 5. 2B as delet. ocala 78 mm. Pema tOOL tut BPCCLINEM. on) soos sks « alee ae’ wierd Sane 106 mm. Alaska, Queen Charlotte Island (Smith), Puget Sound, Oregon, northern to Lower California. This species is common in and around San Francisco Bay, where it is caught in large numbers for food. It is the common crab of the markets. 52 CALIFORNIA ACADEMY OF SCIENCES. Cancer gracilis Dana. Cancer gracilis Dafa, Proc. Acad. Nat. Sci. Phila., Vol. VI, 1852, p. 73; Crust. U. S. Expl. Expd., Vol. I, 1852, p. 153, Pl. VII, fig. 2. Srimp- son, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 460. Coopsr, 1. c., 1860, p. 389. Lock1Nne- ToN, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 95. Newcomss, Bull. Nat. Hist. Soc. Brit. Col., 1892, p. 23. Carapace quite strongly convex, finely granulated and not undulated. Front not produced; the three median teeth reach further forward than the outer pair; the central tooth is smaller than the others but projects farther anteriorly. No tooth on the superior orbital margin. Postorbital tooth larger than the ones behind it, which are low, triangular and sub- equal in size. Asmall tooth on the nearly straight posterior margin. The pointed extremity of the basal antennal joint does not extend nearly so far forward as in the preceding species. A rounded lobe on the inferior orbital margin just external to the basal antennal joint. Merus of the maxillipeds distally rounded. Merus of the chelipeds with the distal extremity of the upper margin armed with two teeth as in magister; carpus ridged on the outer surface as in magister and havinga strong spine at the antero-internal angle, and another spine below it; the upper edge of the hand is acute and may bear a few teeth, but it is often edentulous; the outer surface furnished with granulated cost#, which sometimes become quite faint. The fingers resemble those of magister; the upper margin of the dactyl is acute but not dentate; legs long, nearly naked, the dactyls slender, compressed. Sex. Length of Carapace. Breadth. Proportion. Male 1.51 in: 2.30 in. 1: 1,52 ui 1.60 in. 2.40 in. 1: 1.50 ne 1.70 in. 2.00) 11, 1: 1.50 Female 1.90 in. 2.80 in. 1: 1.47 es 1.45 in. 2.13 in. De Shag The males therefore appear to be relatively wider than the females. Vancouver Island (Newcombe), Puget Sound, Toma- les Bay, San Francisco Bay! San Pedro! San Diego! This is one of the smallest species of the genus Cancer and it is also the rarest of our species. It is more closely allied to magister than to the other forms, but CALIFORNIA STALK-EYED CRUSTACEA. 53 may be easily distinguished from that species by the greater convexity and smoothness of the carapace, the character of the antero-lateral teeth and its smaller size. Genus Trichocarcinus Miers. (Trichocera DE HAAN preoccupied.) Carapace cancroid in form, the sides rounded; antero-lateral margins not sharply marked off from the postero-lateral; upper surface very uneven. Front with five teeth. Antennules nearly longitudinal. Antenna quite long, setose on all sides, the first joint elongated and occupying the inner orbital hiatus. Abdomen of the female seven-jointed; in the male five- jointed, the third, fourth and fifth joints fused. Crabs of small size. Type.—T. gibbosula (DE HAAN.) Although this genus was formerly placed among the Corystoid crabs it is in many respects, as Miers has observed, quite closely allied to the genus Cancer. Can- cer antennarius shows many characters in common with this genus. | Trichocarcinus Walkeri (new name). Trichocarcinus recurvidens WALKER, Trans. Liverpool Biol. Soc., Vol. XII 1898, p. 271, Pl. XV, figs. 1-10. ‘*Carapace 19 mm. long by 23 mm. wide, divided into a number of promi- nent areole of which the surfaces are flattened and granulated; they are separated by deep interspaces. ~ *~ * The antero- and postero-lateral margins are ill defined; the former has ten teeth (including the orbital) which are convex on the upper surface, widening distally, and inclined upwards, whence probably the specific name. * * * Eye-stalks short with a bifid calcareous appendage on the upperand innerside. * * * Differs from 7’. oregonensis (Dana) in the flatter, more sharply defined areolz, in the distal expansion of the teeth of the antero-lateral margin, and in the different form of the calcareous ocular appendage. Color of areolw bright red, chelipeds and legs flesh color, fingers black. One male.” Puget Sound. 54 CALIFORNIA ACADEMY OF SCIENCES. The specimen described by Mr. Walker was referred by him to Bate’s species only with great hesitation, as Bate’s ‘‘imperfect description would apply quite as well to T. oregonensis.”” The determination is made solely on the ground that Bate calls recurvidens a ‘‘pretty spe- cies’’—a term which applies better to Walker’s specimen than to the less attractive species of Dana. I do not believe, however, that Walker’s identification is correct, as Bate says that recurvidens ‘‘may be distinguished by the sharp points of the inner lateral teeth, granulated or minutely baccated along the margin and haying the apex recurved.” The teeth of the specimen described by Walker, instead of being sharp pointed, are distally widened with a wide, transverse, distal margin. The form that Mr. Walker describes has, therefore, been given a new specific name. Bate’s species may be 7’. oregonensis or, as Miers has suggested, the young of Cancer antennarius Stimpson. Trichocarcinus oregonensis Dana. Trichocera oregonensis DANA, Proc. Acad. Nat. Sci. Phila., Vol. VI, 1852, p. 86; Crust. U. S. Expl. Expd., Part I, 1852, p. 299, Pl. XVIII, fig. 5. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 464. Wurt- ‘EAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. Trichocarcinus oregonensis Miers, Proc. Zool. Soc. London, 1879, p. 34; Challenger Reports, Vol. XVII, 1886, p. 110. Smiru, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 207. Nerwcomsz, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 24. Waker, Trans. Liverpool Biol. Soc., Vol. IT, LS9Ss ps Zi, ele OVE seas Carapace elliptical, evenly rounded at the sides; areas distinct and ele- vated, especially in front, and covered with rounded granulations; sulci between the elevations wide, deep and smooth, especially on the anterior portion of the carapace. The front does not project beyond the line of the orbits; the outer pair of teeth (preorbital) are lobe-like, having a broad, eranulated edge, and are separated from the inter-antennary portion of the front by a deep, concave notch; the median portion of the front is very short, truncated, and obscurely divided by a slight incision on either side CALIFORNIA STALK-EYED CRUSTACEA. 55 of the center into a small median tooth and two larger lateral ones. A tooth between the two fissures in the superior margin of the orbit. Anten- nules nearly longitudinal, the fosettes extending nearly to the margin of the front. Basal antennal joint not strongly produced at its distal end; second joint broad, flattened, distally expanded and provided with a tuft of long set at the distal end; third joint subcylindrical and about as long as the preceding; flagellum long and hairy. A small tooth on the inferior margin of the orbit next to the inner fissure. Eye-stalks with a small tooth on the anterior surface. Sides of the carapace armed with ten to thirteen closely set, granulated teeth which extend upon the postero- lateral margin; in front of the rounded lateral angles the teeth are strongly upturned and bent forwards; alternate teeth are the more acute; the first sharp tooth the third from the front. Antero-external of the merus of the maxillipeds produced; the inner margin notched behind the articulation of the palp. Chelipeds very stout; merus about as wide as long; carpus granulated on the outer side, the granules becoming larger above where there may be a few small tubercles; a spine at the antero-internal angle below which there is a tooth; hand thick and high; the short upper edge of the palm bears two rows of small tubercles and on the outer surface there are five delicate, granulated lines, the two lower lines the finest and continued upon the short, thick pollex; both fingers dark colored. Legs hairy; dactyls about as long as the propodi. Sixth abdominal segment in the male nearly square, the last narrowly triangular. Length of carapace, male....... St AOI S ee | ae 17 mm. Brendthot carapace, Male: 6... 65 cooks a answr «cles aes 21 mm. Mutiny Bay, Alaska! Queen Charlotte Island, Van- couver’s Island, British Columbia, Gulf of Georgia, Puget Sound, northern to Lower California! From low tide mark to several fathoms in depth. Miss Mary Rathbun has recently united Trichocar- conus with the genus Cancer and reports two Japanese species, gzbbosulus (De Haan) and amphictus (= Tricho- carcinus dentatus Miers), from California.’ Oregonensis is closely allied to De Haan’s species. Section XANTHINI Ortmann. Carapace rounded or subquadrate. Front wide and generally divided by a median notch. Antennules oblique or transverse. 1See Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 581. 56 CALIFORNIA ACADEMY OF SCIENCES. Genus Cycloxanthops Rath. Front horizontal, produced, and divided by a median fissure into two deep, lamellate lobes, which are truncated and separated from the inner orbital angle by a rather deep fissure. Orbits small, with the superior margin marked by two fissures. The postorbital angle is only slightly prominent and is continuous with the antero-lateral margin, which is strongly curved behind. Basal antennal joint short, but joined at its internal angle to the front; the flagellum is inserted in the inner orbital hiatus. Merus of the maxillipeds subquadrate, the anterior margin not oblique. The abdomen in the male is composed of five free joints. Type.—C. sexdecimdentatus (Epw. & Lucas). The genus Cycloxanthus was established by Milne- Edwards’ to receive some species of fossil crabs. Find- ing subsequently that this species was a synonym of Zanthopsis M’Coy, Edwards applied the name Cyclo- xanthus, which he considered as having thus become ‘‘disponible,” to an entirely different genus of which he made sexdecumdentatus the typical species.” According to the rule, ‘‘ Once a synonym always a synonym,” the name Cycloxanthus is no longer available for use. Cycloxanthops novem-dentatus (Lock.) PL. = f- 2 : Xanthodes ? novem-dentatus LocKIN@Ton, Proc. Cal. Acad. Sci., Vol. VII, Li, Ps. o2- Xantho novem-dentatus Lockineton, [bid., 1877, p. 99. Kine@stey, Proc. Bost. Soc. Nat. Hist., Vol. XX, 1879, p. 153. Cycloxanthus californiensis RatHBUN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 237. Carapace wide, flattened, and more or less punctate behind; convex, granular and rugose in front. Median region divided into three well- defined regions by distinct sulci; branchial regions divided bya transverse sulcus arising from the lateral angles. Front horizontal, produced, eden- tate, with the anterior margin nearly transverse, but a little advanced in the center, and divided bya deep, closed, median notch which may become 1 dans d’Archiac, Hist. des progr. de la géologie, T. III, p. 304, K. 2Ann. Sci. Nat. (4), T. XX, 1863, pp. 278 and 307. CALIFORNIA STALK-EYED CRUSTACEA. 57 obliterated above by the fusion of the two sides; the anterior margin of the lobes may be straight, or, more often, slightly concave; seen from in front they are gently arched upward, or in some cases very strongly so. Antero-lateral margin armed with eight or nine small subacute teeth whose bases are generally narrower than the spaces between them; postorbital tooth small; next tooth also small, depressed and often lobate, and sepa- rated from the postorbital by a considerably greater interval than from the succeeding tooth; at the rounded sides of the carapace the teeth become larger, more acute, and more closely set. There is a tendency to alterna- tion in the size of the teeth; the fourth tooth, counting the postorbital, is larger than the fifth, and the sixth and eighth are larger than the seventh; ninth tooth small and situated behind the lateral angle; a small tenth tooth often occurs closely behind the ninth, On the outer margin of the orbit above the postorbital tooth there are two small, blunt teeth separated by a narrow fissure; preorbital tooth distinct; suborbital lobe acute. Basal joint of the antennules with a pronounced ridge on the outer surface with a groove on either side. Merus of the maxillipeds obliquely truncated at the anterior end. Merus of the chelipeds hairy above; carpus rugose, and furnished with two blunt teeth at the antero-internal angle; hand rugose above, and rather long, with the upper and lower margins nearly parallel; fingers long, sulcate, not gaping, and colored brown, with light colored tips. Legs with the margins hairy; dactyls longer than the pro- podi and terminating in nearly straight, corneous claws. Abdomen in the male narrow, the second joint a little overa half longer than wide, penul- timate joint longer than wide; last joint about as long as broad, with the distal end rounded. In the abdomen of the female the second joint is longer than the third or fourth, which are subequal and shorter than the fifth; sixth joint the longest, nearly equaling the third and fourth com- bined; last joint broader than long, and in adult specimens broadly rounded at the tip. The following measurements are from a series of specimens from San Diego, California:— Length of Carapace. Width of Carapace. Male. 25:79 mm. 38.75 mm. ae 25) =) mm on-oL) Tn, 24 mm. 35.25 mm. e oa mm. 35 mm. ss 22, am. a2. Wim. se 20 mm. 30 mm. Hg 18 mm. 26 mm. a 14.75 mm. 21 mm. BA 12 mm. 17 mm. 58 CALIFORNIA ACADEMY OF SCIENCES. Length of Carapace. Width of Carapace. Female. 26 mm. 40 mm. mn 21.75 mm. 32 mm. ms 15 mm. oe mm? ae 11 mm. —«16 mm. One large specimen from southern California (pre- cise locality unknown) measured 36.25 mm. by 56.75 mm. | San Pedro! San Diego! southern California! Catalina Harbor (Rathbun, M.), Guadalupe Island, Lower Cali- fornia (Rathbun, M.), San Clemente Island! This is a very common species on the coast of south- ern California. I have not seen Lockington’s type, which could not be found in the Academy, but there were other specimens in Lockington’s collection which bore the name Xantho vittata St. written above the ear- lier name Xantho novem-dentatus which had been partly scratched out. I have seen the types of Cycloxanthus californienis and compared them with specimens I had identified as novem-dentatus by a comparison with the specimens _in Lockington’s collection. There is, I believe, no doubt that the two species are identical. This species is closely allied to vittatus St., but I think we must accept Miss Rathbun’s conclusion, that it is distinct, although it was considered a synonym of the latter species by Milne-Edwards and subsequently by Lockington himself. At the National Museum in Wash- ington I had the opportunity of comparing specimens of vittatus from the western coast of Mexico with several specimens of this species. The former species differs in its longer and sharper lateral teeth, its 3 broader hands, and its narrower abdomen. CALIFORNIA STALK-EYED CRUSTACEA. 59 Cycloxanthops rugosa, sp. nov. Carapace very uneven, granulated, and thickly covered with small, cir- cular pits. Postorbital tooth small; antero-lateral teeth irregular. Ante- rior portion of the subhepatic region prominently granulated and pitted and not sharply separated from the strongly sloping anterior portion of the hepatic area. Front shorter and more depressed than in novem- dentatus. External maxillipeds much as in the preceding species; merus pitted and granulated. Carpus of the chelipeds strongly reticulated above; a prominent tooth at the antero-internal angle, below which is a smaller tooth. Hands narrow, strongly reticulated above and on the upper por- tion of the outer surface, the inner margin of the upper side furnished with several irregular tubercles. Ambulatory legs much as in novem- dentatus but less hairy. San Diego, California. Collected by A. U. Crawford. Described from a specimen in the museum of the University of California. Easily distinguished by the conspicuous pits and prominent granulations of the carapace. Genus Lophoxanthus A. Milne-Edwards. Carapace broader than long, flattened, the anterior portion of the antero- lateral margins straight, nearly devoid of teeth, and meeting the dentate posterior portion at an angle. Front short and divided by a median fis- sure into two broad, truncated lobes. Orbits nearly circular and marked with fissures which are closed or even obsolete. Basal antennal joint short and in contact with the infero-lateral process of the front, but not entering the inner orbital hiatus. Maxillipeds nearly quadrate, the merus distally truncated, the antero-external angle not produced. Ambulatory legs compressed and carinated or crested above. Abdomen in the male five-jointed. Type.—L. lamellipes (STIMPSON.) This genus is doubtfully distinct from Lophozozymus, being distin- guished chiefly by its more transverse carapace and the absence of con- spicuous teeth on the anterior portion of the antero-lateral margins. The three species of this genus here described are closely allied. In all these the areolation of the carapace is distinct, the postorbital tooth minute, the anterior half of the antero-lateral margin straight and furnished with a single low, obscure tooth, the posterior half nearly longitudinal and furnished with three, prominent, subequal teeth, the epistome with a ridge extending upon it from the sides, the inferior inner orbital tooth 60 CALIFORNIA ACADEMY OF SCIENCES. pointed, and the tooth external to the latter obtuse and separated from the postorbital by a fissure, the upper margin of the orbit furnished with two fissures, the merus of the maxillipeds rectangular, notched behind the articulation of the palp, and crossed by an oblique ridge, the chelipeds stout, the merus trigonous and dentate above near the distal end, the car- pus rounded and furnished with a tooth at the inner angle, the hands with a lobe pointing inward at the base of the upper side of the palm, the fingers irregularly dentate within but with the outer margins rounded; the ambulatory legs furnished above with lobulated crests on the earpi and propodi, the dactyls slender and villous, the second segment of the abdomen narrowed distaliy, the third segment wider at the base than the distal end of the second and touching the coxa of the fifth pair of legs, the last two segments of subequal width, the terminal joint rounded. Toph Ns bellus (Stimpson). Xantho bella Stimpson, Ann. N Lye. Nat. Hist., Vol. VII, 1860, p. 204, Pl. III, fig. 2. Bats, in Lord’s Nat. in Vntacon vere Island, Vol. II, 1866, p. 270. Newcomeg, Bull. Nat. Hist. Soc. Brit. Col., 18938, p. 24. Xanthodes Hemphillii Micor newex: Proc. Cal. Acad. Sci., Vol. VII, 1877, p- ol. Xanthodes Hemphilliana Lockineton, |. c., 1877, p. 100. Lophoxanthus bellus A. MILNE-Epwarps, Crust. Miss. Sci. au Mex., Pt. V, p. 256, Pl. XLVI, fig. 4. Lophozozymus ( Lophoxanthus) bellus Migrs, Challenger Reports, Vol. XVII, 1886, p. 115. Lophopanopeus bellus Ratupon M., Bull. Lab. Nat. Hist., Univ. Iowa, 1898, p. 272. Carapace flattened behind, convex in front, and transversely nearly plane; surface distinctly areolated, becoming roughened toward the antero- lateral margins. Front generally sinuate, and divided by a median fissure, the outer angles produced into small rounded lobes or teeth, which are separated from the adjacent supraorbitals by a fissure. Postorbital tooth small. The anterior half of the antero-lateral margin is nearly straight, meeting the posterior half at an angle, and furnished with a single low, inconspicuous tooth which is sometimes absent; posterior half furnished with three prominent, subequal, horizontally flattened teeth, the anterior one the least acute. Postero-lateral margins nearly straight, rapidly con- verging. The infero-internal angle of the orbit is produced into a promi- nent pointed tooth; a broad lobe-like tooth at the infero-external angle of the orbit separated from the postorbital above by a fissure. Merus of the maxillipeds subrectangular, the surface with two depressions separated by an oblique elevation. Chelipeds stout, the upper margin of the merus furnished with a few irregular teeth; surface of the carpus more or less roughened, the antero-internal angle furnished with a tooth which is CALIFORNIA STALK-EYED CRUSTACEA. 61 typically double, the lower cusp obscure; hands rounded above and fur- nished with a lobe projecting inward at the intero-proximal portion of the upper side; fingers black or dark brown. Abdomen in the male five-joint- ed, the last segment rounded and slightly wider than the preceding one, which is somewhat wider than long. Merus of the ambulatory legs acute above; carpus with a bilobed crest; crest of the propodus often with a lobe at the base; dactyls villous. Color purplish, the legs crossed with light- colored bars. Found among rocks at low tide from British Columbia to Monterey. Vancouver Island (Bate, Newcombe), Puget Sound (Stimpson)! northern California! Monterey (Stimpson, Lockington)! Locality. Length. Width. Puget Sound; male 19 mm. 26.5 mm. Puget Sound; female 15.75 mm. 22.5 mm. Monterey; male, type of Hemphillii Lock. 18.75 mm. 28 mm. Two specimens whose measurements are given by Stimpson were .54 by .80in.and .56by .89in. ‘‘Northern specimens,” says Stimpson, ‘‘are more transverse, rougher, more pubescent, and more sober in coloration than those found in warmer latitudes.’’ Possibly speci- mens of leucomanus or frontalis were confused with this species. I have seen Lockington’s type of Hemphillic (No. 35) which is preserved in the collection of the California Academy of Sciences. It isa typical bellus and as strongly transverse and as rough as the specimens I have seen from farther north. The specimen from Lower California which is figured as JL. bellus by A. Milne-Edwards (1. ¢.) probably belongs to a different species. i er ed sam (Lock. ) Xanthodes leucomanus LocKINcToX, Proc. Cal. Acad. Sci., Vol. VII, 1877, pp. 32 and 100. Lophoxanthus bellus A. MILNE-Epwarps, Crust. in Miss. Sci. au Mex., Pt. Rly 256 (in part). Lophopanopeus leucomanus RATHBUN M., Bull. Lab. Nat. Hist. Univ. Iowa, 1898, p. 272. | 62 CALIFORNIA ACADEMY OF SCIENCES. Closely allied to bellus, but having the anterior portion of the antero- lateral margin not so nearly transverse, the surface of the carapace more eroded, the lateral teeth narrower and more nearly conical, the ambulatory legs somewhat more slender and more prominently crested. The areola- tion of the carapace is plain; the front is produced more than in bellus, and divided {by a median notch, the lobes sinuate with the outer angles produced as in the preceding species. Orbital fissures and teeth as in bellus; lateral teeth of the carapace subconical, prominent and subequal, the anterior one least acute. Subhepatic regions eroded, especially near the orbits. Chelipeds unequal; merus irregularly dentate on the upper margin; carpus and upper surface of the hand generally eroded or furnished with a network of raised lines; hand with a lobe at the base as in bellus; fingers light to dark brown. Merus of the ambulatory legs with an acute upper margin which usually ends in a tooth near, but not at the distal end; carpus with a bilobed crest above; crest of the propodus with a lobe near the base. Dactyls rather slender. Locality. Length. Width. Monterey, male. 13.5 mm. 18.5 mm. Santa Catalina Island, male. 1235, mam. 17.75 mm. Southern California, male. 8.5 mm. 11.25 mm. es a Semale. 11.25 mm. 14.75 mm. West America, male. 13.5 mm. 18 mm. ue e female. 10 mm. 13) “mim, ae te Hs LO” Sma 13 tae La Paz, Lower California, male. 7.75 mm. 10° ama: ried | 168 os es * 6.5 mn. 8.5 mm. rae es ee es Female. 7) ome 9 mm. Monterey (Lockington)! Santa Catalina Island! San Diego and Santa Rosa Island (Lockington), La Paz, Lower California (Lockington)! ‘‘ Southern California,”’ two specimens (No. 17286 U.S. National Museum)! San Clemente Island! San Diego! This species varies considerably as regards the rough- ness of the carapace and legs. In some cases the ante- rior part of the carapace and the upper side of the car- pus and manus of the chelipeds are very much eroded, but occasionally they are nearly smooth; the tubercle on the middle of the surface of the maxillipeds is often CALIFORNIA STALK-EYED CRUSTACEA. 63 prominent, but in some specimens it is quite inconspic- uous. The crests on the ambulatory legs are also quite variable, and the tooth at the end of the sharp upper edge of the merus may be absent. These differences are probably due, in great measure, to age. The forms from near Japan which Miers identified with this species under the name Lophozozymus (Lopho- zanthus) bellus St. var. leucomanus Lock. belong, I believe, to a distinct species. The lateral teeth in Miers’ form are quite different from leucomanus; the hands, as shown in Miers’ figure, lack the lobe at the base of the palm; ‘‘the mobile finger is longitudinally carinated and sulcated above,” while in lewcomanus it is rounded and perfectly smooth; the pits on the upper side of the hand are arranged in longitudinal series, but in leucomanus this arrangement is not shown.' The types of lewcomanus are probably no longer pre- served. I have specimens from localities from which this species was reported by Lockington, and I have seen Lockington’s specimens from La Paz, Lower Cali- fornia.” These specimens are small and strongly eroded, but present no well-marked differences from the north- ern forms. _ None of: the specimens of leucomanus I have seen attain nearly the size reached by bellus. 1 See Challenger Reps., Vol. XVII, p. 115, Pl. XI, fig. 1. 2 See Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 100. 64 CALIFORNIA ACADEMY OF SCIENCES. Lophoxanthus frontalis (Rathbun). PLE, f- 846. | Lophozozymus (Lophoxanthus) frontalis RatHBuN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 236. Lophopanopeus frontalis RaATHBUN M., Bull. Lab. Nat. Hist., Univ. Iowa, 1898, p. 272. Closely allied to the two preceding species, especially to leucomanus. The shape of the carapace is nearly the same as in the preceding species, but the upper surface is smoother, not eroded, the lateral teeth more flat- tened horizontally and slightly upturned; the areolation is plain; the subhepatic regions are granulated but not eroded. Front considerably produced, sinuate, the median notch deep. Orbits and orbital teeth as in leucomanus. Maxillipeds granulated, the merus shaped as in the two pre- ceding species, and crossed by an oblique elevation. Chelipeds unequal; merus with afew teeth, on the upper margin; carpus rugose or nearly smooth, with a tooth at the-inner angle and a tubercle at the base on the — upper side; hands large, with a lobe pointing inward at the base of the upper side of the palm; fingers brown, the color extending far back on the hand. Legs rather slender, more or less hairy, the upper edge of the merus acute but having no well-marked tooth near the end; crests of the carpus and propodus as in leucomanus but less prominent; dactyls slender. Length of carapace 15 mm.; width 20 mm.; length of larger hand 16 mm. Described from a specimen from San Diego sent by Miss Rathbun (No. 19823, U. S. National Museum). San Diego (Rathbun)! Monterey (Rathbun). This species may readily be distinguished from bellus by its different shape; the portion of the carapace lying in front of a line connecting the tips of the first lateral teeth is about one-fourth the length of the whole cara- pace in bellus, and about one-third of this length in leucomanus and frontalis. The color of the fingers extends far back upon the hands in this species, while in both the preceding it does not extend further back than the base of the fingers. The front is much more produced than in bellus. The hands and carpal joints of the chelipeds are, like the carapace, much less eroded than in leucomanus, and the merus of the maxillipeds has the oblique ridge low and flattened and not raised into a tubercle in the center. CALIFORNIA STALK-EYED CRUSTACEA. 65 Genus Xanthias Rathbun. (Genus Xanthodes DANA, preoc.) Carapace transverse, distinctly areolated, more narrow and convex than in the genus Xantho, to which it is very closely allied. Antero-lateral margin not thin-edged or cristiform; teeth tuberculiform or even spinous. Posterior portion of the carapace depressed. Antennules transversely or obliquely plicated. Basal antennal joint short, and barely reaching the slender, narrow, infero-lateral process of the front, and not produced within the hiatus of the orbit; flagellum not excluded from the orbit. Chelipeds stout, the tips of the fingers acute (not excavated within). Ambulatory legs compressed or subcylindrical and devoid of a crest on the upper margin. Abdomen of the male five-jointed. Xanthias Taylori( St. ) Xanthodes Taylori Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1859, p. 208, Pl. III, fig. 3. Srreets and Kinestey, Bull. Essex Inst., Vol. IX, 1878, p. 105. A. Mintne-Epwarps, Crust. in Miss. Sci. au Mex., Bie. Pp. 200 Pl. XLV; fig. 3: Xantho spini-tuberculatus Lock1neTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, pp. 31 and 99. Carapace flat behind; in front quite strongly convex longitudinally but transversely nearly plane. Areolets distinct and embossed anteriorly; antero-lateral regions strongly lobulated with a prominent, smooth, rounded lobule above each marginal tooth. The lobes of the front are separated by a wide, rounded notch and furnished with a rounded tooth at the inner and outer angles, between which are several small, baccate protuberances. Closely behind and parallel with the anterior margin of the front is a baccated ridge interrupted in the middle by the prominent longitudinal groove which extends backwards from the median notch. Behind this ridge is another elevation not so distinctly baccated, which is separated by a transverse sulcus from the elevated anterior margins of a pair of lobules lying behind it. Median region divided into three distinct areas. Inner orbital angle prominent and separated by a deep notch from the outer angle of the front. Upper orbital margin with a rounded tooth bounded by a pair of sulci. Postorbital tooth small and continuous externally with a rounded protuberance; the latter is separated by asmooth sulcus from a pair of smooth, rounded prominences, one of which is situ- ated directly above the other. The three posterior teeth on the antero- lateral margin prominent, the last two acute and curved forwards, the first obtuse and generally more or less bifid. A small tooth usually present behind the one at the antero-lateral angle. A tooth below the infero- 5 66 CALIFORNIA ACADEMY OF SCIENCES. orbital fissure, and a prominent, subacute tooth at the infero-internal angle. Ocular peduncle with a few baccations in front. Antennules transversely plicated, the basal joint with a granulated or baccated ridge. Basal antennal joint short but touching the outer angle of the front. Merus of the maxillipeds subrectangu lar, transversely truncated in front, with the outer angle rounded and slightly produced. Chelipeds stout, more or less unequal; merus trigonal, with the distal portion of the upper margin armed with a few spine teeth; carpus thickly covered with promi- nent, rounded, smooth, glossy, rose-colored tubercles; hand oblong, with the palm longer than wide, the upper and outer surface covered with rose-colored tubercles like those on the carpus, arran ged in seven or eight longitudinal rows; fingers stout and colored black, the color not extending back upon the hand. Legs compressed, thickly covered with stiff setz and strongly spinous, especially on the merus and propodus; dactyls about as long as the propodi. Abdomen of the male five-jointed, the penultimate joint about as long as wide; last joint broadly rounded. Length of carapace 13 mm.; breadth 19.5 mm. Monterey (Stimpson)! Santa Rosa! San Pedro! San Diego! Santa Catalina Island! Magdalena Bay and San José Island tan ae ‘« This species,’ says Lockington, pear to be of common occurrence from Monterey southward to Mag- dalena, but to disappear, or at least to become rare, in localities further south.” Xanthias latimanus (Lock. ) X anthodes latimanus LocK1neTOoN, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 31. Xantho latimanus LockinerToy, 1. ¢c., p. 101. Lockington’s description of this species is as follows: ‘‘ Front sinuate, the inner angle of the orbit raised into a point; carapax but slightly transverse; teeth N. T.S. prominent and pointed, D and E almost obso- lete; areolation of medial and antero-lateral regions distinct, the former having the parts 2M and 3M entirely outlined. Hands subequal, the right somewhat the larger; movable fingers very long and curved abruptly downward; margin of manus continuous with the broad base of the fixed finger so as to form a sinuous sloping line; hinder feet compressed. This species may readily be identified by the delicate marbling of the carapace and chelipeds, and the downward bend of the movable fingers. Abdomen. of male five-jointed. ‘‘A single male specimen from San Diego. ‘* Length 0.73; breadth 0.88 [inch].” CALIFORNIA STALK-EYED CRUSTACEA. 67 I have never seen this species and the description is not sufficiently complete to enable one to determine whether or not it belongs in the genus in which Lock- ington placedit. Professor A. Milne-Edwards considers it identical with Stimpson’s Micropanope latimana. Milne-Edwards does not mention having seen latimana, and as he simply transcribes Stimpson’s description it is quite certain that he bases his identification solely upon the descriptions of the two authors. The identification is not improbably correct, but considering the brevity of both the descriptions, it is somewhat unsafe to unite the two species, especially since the characters mentioned by Stimpson are mainly those of which Lockington says nothing. Genus Pilumnus Leach. Carapace convex, little broader than long, and hairy above. Antero- lateral margins regularly arcuated, shorter than the postero-lateral, and armed with short spines. Front narrow, emarginate. Endostome liongi- tudinally carinated. Basal antennal joint short, barely reaching the infero-lateral frontal process and lying under the orbit. Ambulatory legs compressed, not carinated, and armed with spinules; dactyls slender, nearly straight. Abdomen of the male seven-jointed. Pilumnous spino-hirsutus (Lock. ) Acanthus spino-hirsutus LocKINGTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, pp. 32 and 102. Pilumnus spino-hirsutus STREETS and KINGSLEY, Bull. Essex Inst., Vol. IX, 1877, p. 107. KinesLry, Proc. Bost. Soc. Nat. Hist., Vol. XX, 1879, p. 154. A. MILNE-Epwarps, Crust. in Miss. Sci. au Mex., Pt. V, p. 278. Miers, Challenger Reports, Vol. XVII, 1886, p. 147. Carapace strongly convex, nearly smooth, but covered with stiff seta. Median frontal lobes truncated, separated by a prominent notch, and armed each with four or five spines; lateral lobes of the front small, sepa- rated from the median lobes by a deep notch and ending ina spine. Orbits with the upper, lower and outer margins armed with strong spines of unequal size, the two spines at the intero-inferior angle large and situ- ated on a kind of lobe. Besides the postorbital the antero-lateral margin is armed with three strong spines, and there is a small spine below the margin in front of the first of these. The basal antennal joint barely 68 CALIFORNIA ACADEMY OF SCIENCES. reaches the infero-lateral frontal process. Endostome plainly carinated. Pterygostomian regions more or less granulated. Chelipeds unequal, setose; merus scarcely as long as high, the margins finely denticulated; a spine at the supero-distal angle separated from a similar spine behind by a deep notch; outer surface of the carpus thickly covered with spines; upper and outer surface of the hand covered with several series of spines; lower margin finely denticulated; fingers dark colored. Legs setose; carpus and propodus armed above with spines; merus with a spine at the supero-distal angle; dactyls hairy, nearly straight, and subequal to the pro- podi. Terminal segment of the abdomen in the male triangular. San Diego, California (Lockington’s type)! Mulege Bay, Gulf of California! APPENDIX TO THE XANTHINI Ortmann. Genus Telemessus White. ) Carapace depressed, subpentagonal, broader than long and covered with seta. Front wide, the median portion cut into four teeth and separated by a sinus from the lateral portions which form the preorbital teeth. The antero-lateral and postero-lateral margins are dentate and meet each other at an angle. Orbits large; postorbital tooth prominent. Antennules folded longitudinally or nearly so in wide fossettes. Epistome with a triangular point extending forward between the antennules. Basal anten- nal joint broad, forming a part of the outer wall of the antennulary fossettes and produced on the outer side intoa lobe which occupies the inner hiatus of the orbit; flagellum quite long. Maxillipeds produced a little beyond the anterior margin of the buccal area; the merus has a more or less tri- © angular apex and bears the palp on the inner margin just behind the tip. Chelipeds setose and scabrous or spinous and rather short in both sexes; merus trigonous; hands compressed and furnished with spinous or tuber- culated ridges on the outer surface. Legs quite long, compressed, sca- brous or spinose; dactyls long, compressed, acute. Abdomen of the female seven-jointed, the sides of the sixth joint concave, leaving the genital © openings exposed. Abdomen of the male widest at the third segment and having the third, fourth and fifth joints fused. Type.—T'. cheiragonus (TILESIUvS). Unless Latreille’s bare mention in a list of genera of the name Cheiragone' without referring to it any species 1Familles nat. a Paris, 1825, p. 270. CALIFORNIA STALK-EYED CRUSTACEA. 69 constitutes the genus Chezragonus (or Cheiragone), Tele- messus has priority. According to the A. O. U. code, Cheiragone must be considered a ‘‘ nomen nudum.” , ‘ ‘i Telemessus cheiragonus (Tvleszus). Cancer cheiragonus TILEstus, Mem. Acad, St. Petersbourg, T. V, 1815, p. a4ti~ Pl. VEE) fie. (1, Telemessus serratus Waite, Ann. Nat. Hist., Vol. XVII, 1846, p. 497; Voy- age of Samarang, Crustacea, 1848, p. 14. Dana, Crust. U. S. Expl. Expd., Part I, 1852, p. 303, Pl. XVIII, fig. 8. SmirH, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 208. Platycorystes ambiguus BRANDT, Bull. Phys-math. Acad. St. Petersbourg, SI VIb, 1848. ni. 179: Platycorystes cheiragonus BRANDT, Middendorff’s Siberische Reise, Bd. II, Eh. fF. IS5i. p. 85. Cheiragonus hippocarcinoides BRANDT, Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 147. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 465. Telemessus serratus and T. cheiragonus Miers, Proc. Zool. Soc. London, 1879, p. 36. Telemessus cheiragonus BENEDICT, Proc. U. S. Nat. Mus., Vol. XV, 1892, ~p. 224, Pls. XXXV and XXXVI, figs. 2,3 and 4. Nzwcompz, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 24. WaLkeEr, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 373. Cheiragonus cheiragonus ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894, p. 420. Carapace depressed, areolated, furnished with granules and thickly cov- ered peculiar clavate setz. The teeth of the median lobe of the front are triangular, subequal, and extend forward to about the same distance. The preorbital teeth are large and acute and the rounded sinus between them and the median frontal lobe is armed with secondary teeth. Postorbital tooth acute. The margins of the orbit are denticulated and the upper margin is furnished with two small fissures; lower margin entire but deeply concave. Lobe of the basal antennal joint triangular. Merus of the chelipeds spiny; two rows of spines on the upper edge of the hand (the outer row may be reduced to tubercles) and below these four rows of spines or tubercles on the outer surface. Legs furnished with transverse, setose, granulated or, in some cases, spinous lines; dactyls strongly grooved, longer than the propodi. The abdomen of the male is abruptly contracted behind the third segment, a small part of the posterior margin of which projects nearly transversely. The antero-lateral margin of the 70 CALIFORNIA ACADEMY OF SCIENCES. carapace is furnished with four prominent triangular teeth including the postorbital and the one at the lateral angle; the posterior margins of these teeth are armed with several denticles and the three anterior teeth have each two denticles on the anterior margin; the fourth tooth has three nearly equally spaced denticles on the inner side of its anterior margin and a fourth at a longer interval from the others near the tip. Postero- lateral margin with two teeth. Body and appendages strongly setose. Length of carapace, 50 mm.; breadth, 64 mm. Northeastern Siberia, Aleutian Islands, Behring Sea, Sitka, Vancouver’s Island, Puget Sound, upper Cali- _ fornia. There is a specimen in the museum of the University of California labelled Gulf of California, but possibly this is wrong, as this species appears to be a northern one. This species may be distinguished from I. acuti- dens by its broader carapace, its larger and broader lat- eral teeth and by the median teeth of the front. Fora fuller list of localities and other facts concerning this peculiar species see Benedict |. c., pp. 224-227. Section PORTUNINEA Ortmann. Carapace more or less rounded, with the antero-lateral margin sharply marked off from the postero-lateral. The last pair of thoracic legs fitted for swimming. Verges of the male in the coxa. Genus Portunus Fabr. Carapace usually transverse, depressed, and marked with granulated lines. Front short and cut into five to eight teeth. Antero-lateral mar- gins arcuated and armed with nine teeth, all but the last (the largest) being of subequal size. Flagellum occupying the inner orbital hiatus. Merus of the chelipeds armed with spines on the inner margin; hand elongated, prismatic and costate. Type.—P. pelagicus (LINN.). : CALIFORNIA STALK-EYED CRUSTACEA. 71 Portunus Xantusii (St. ) Achelous Xantusii Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 222. Neptunus asper A. MILNE-Epwarps, Archiv. du Mus. Hist. Nat. Paris, T. X, 1860, p. 325, Pl. XXX, fig. 3. Neptunus Xantusii A. MILNE-Epwaprps, |. c., 1860, p. 429; Crust. in Miss. Sei, anu Mex. Pt. VY,p. 218, £1) XXXVI, fio. 1,. and Pi: XXXIx, fig. 3. Amphitrite paucispinis Lock1neToN, Proc. Cal. Acad. Sci., Vol. VII, 1877, pn: 107. Portunus Xantusii RatHpun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 593. Carapace markedly transverse, the upper surface pubescent. Gastric area with a median granulated line and a transverse line, near the middle, which is slightly concave in front; the posterior portion crossed by a short transverse line; a granulated line beginning near the posterior part of the gastric area, at first arching forwards, and then curving backwards near the side of the carapace and extending upon the large lateral spine; two short, oblique, parallel lines on the branchial areas; on the cardiac region there are two short transverse lines situated end to end, their inner extremities curving backwards and meeting to form a longitudinal, median line which is often obscure. Front short and not projecting beyond a line drawn across the tips of the acute triangular projections of the infe- rior margin of the orbit, the four frontal teeth are equally prominent, the two middle ones slightly narrower than the outer pair and separated from the latter by a slightly wider interval than they are separated from each other. The tooth at the inner angle of the orbit is double pointed. The postorbital tooth is larger than the one behind it and extends nearly as . far forward as the teeth of the front. The antero-lateral teeth are sharp and curved forward and show a tendency to alternate in size; the lateral spine is about three times as long as the tooth in front. Upper margin of the orbit with two fissures and a small tooth on the inner side of the outer fissure. A single fissure in the lower orbital margin external to which is a small cusp upon the under side of the postorbital tooth. Chelipeds pubescent, merus armed in front with four to seven spines; posterior mar- gin scabrous but not spiny, with the exception of a small spine at the dis- tal end; carpus: with several granulated ridges on the outer surface; a strong spine at the distal end of the upper margin and a smaller spine on the lower side at the distal end of the lowest granulated ridge; hand fur- nished with a spine at the external hinge joint; the granulated ridge on the upper margin terminates in a spine a little behind the distal end of the palm; four granulated ridges on the outer surface of the hand not counting the upper one; fingers strongly ridged, teeth dark colored; first ‘ i CALIFORNIA ACADEMY OF SCIENCES. x three pairs of legs slender; merus unarmed. In the male the first and second abdominal segments are free, the third, fourth and fifth coalesced; second and third joints transversely ridged and produced laterally into two acute processes which project beyond the margins of the other seg- ments; sides of the fourth segment strongly convex; last segment narrow and shorter than the preceding one. In the female, the second and third joints are much like those of the male; fourth and fifth transversely ridged, fifth and sixth of subequal length; last joint very small and triangular. bengethiof ‘carapace; male. ose ace ee ie eee ee eae pee 25.25 mm. Breadth of carapace, male, between tips of lateral spines...53 mm. Heweth otvcheliped; mailers. . 0. 38 ws = ciel oer er ae: 66 mm. eneth-of hand «male: 455 es4. she ore eae ete, Crete 35 mm. Another specimen measured 26 mm. by 52 mm. South America (Milne-Edwards), West Mexico, Pan- ama (Milne-Edwards), Cape St. Lucas (Stimpson), San Diego! Santa Catalina Island! San Pedro! The specimens of Amphitrite paucispinis which were sent by Lockington to Streets and Kingsley were, I believe, wrongly identified with Stimpson’s Achelous panamensis, instead of with A. Xantuszz Stimpson, to which species I feel sure they belong’. Judging from Stimpson’s descriptions, Xantusii and panamensis are closely allied, the latter species being distinguished by the presence of a spine on the merus of the ambulatory legs. Streets and Kingsley state that Lockington’s specimens agreed very well with Stimpson’s description of pana- mensis, except that the spine on the merus of the ambulatory legs was absent. I have examined some of Lockington’s. specimens of Amphitrite paucispinis, which are in the possession of the California Academy of Sciences, and they agree perfectly with Stimpson’s description of Achelous Xantusii, and A. Milne-Edwards’ excellent figures of that species in the ‘‘ Mission Scientifique au 1$ee Bull. Essex Inst., Vol. IX, p. 107. CALIFORNIA STALK-EYED CRUSTACEA. fies’ Mexique.” Panamensis apparently does not range so far north as Lower California, while XYantusii is common from southern California southward. Genus Callinectes Stimpson. Closely allied to Portunus. Carapace broad; front low. Merus of the external maxillipeds prominent and curved outward at the antero-external angle. Abdomen in the male narrow and 1-shaped. Callinectes bellicosus (Stempson). Lupa bellicosa (Stoat MS.) Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1859, p. 57. Callinectes bellicosus ORDWAY, Proc. Bost. Soc. Nat. Hist., Vol. VII, 1863, p.577. Ratusun M., Proc. U.S. Nat. Mus., Vol. XVIII, 1895, p. 365, Pls. XXII, XXIV. fig. 10; XXV, fig. 8; XXVI, fig. 8; and Vol. XXI, 1898, p. 596. Lupa bellicosa ? Lock1neTon, Proce. Cal. Acad. Sci., Vol. VII, 1877, p. 105. Callinectes bellicosus A. MILNE-EDWARDS, Miss. Sci. au Mex., Pt. V, 1879, p. 227, (var. of C. diacanthus). Carapace broad, convex, minutely granulated. Front with two distant spines, the margin between them sinuous. Preorbital teeth not so advanced as the lateral teeth of the front; postorbital tooth long, exceed- ing the preorbitals; subfrontal spine exceeding the lateral frontal teeth; a large tooth below the preorbital, which extends further forward than the others. Inner superior orbital fissure open. Teeth behind the postorbital broadly triangular, acute, with concave sides; last tooth about twice the length of the preceding one, its upper edge continued as a fine ridge for some distance on the carapace. Merus of the chelipeds trigonous, armed anteriorly with four or five spines; carpus with two or three external ridges and a few pointed tubercles near the anterior end; hand with a strong spine above the upper hinge joint at the proximal end of a tuber- culated costa; ridge on the posterior upper edge ending distally ina spine. Width 4.5 in.; length 2.5 in. Gulf of California, west coast of Lower California. A fine specimen of this species was taken by Miss Cook at Point Loma, California, which is the most northern locality from which it has been recorded. 74 CALIFORNIA ACADEMY OF SCIENCES. Subtribe CATOMETOPA, or GRAPSOIDEA. — Carapace wide in front, often subquadrate, sometimes subglobose, and truncated or arcuated anteriorly but not rostrate. The front is usually wide and bent downward. Epistome short, often almost linear. The palp of the maxillipeds may be articulated at the apex or at the inner or outer angle of the merus. Branchiz generally less than nine; efferent branchial channels as in the Maioidea. Verges in the male in the coxa, sternum, or in the cox of the last pair of legs, thence passing through channel in the sternum beneath the pleon. This group contains four families, the Gecarcinide, Ocypodide, Grapside and Pinnotheride. The first family, the Gecarcinide, includes land crabs and is not represented by any species within our limits, although there are several in Lower California and Mexico. Family OCYPODIDA. Carapace moderately convex, cancroid or trapezoidal in form, with the antero-lateral margins straight or rounded, and the branchial regions not greatly dilated. Front of moderate width or very narrow. Orbits and eye-stalks of moderate size or greatly developed. Palp of external maxil- lipeds joined to the antero-internal or rarely the antero-external angle of the merus. Dactyls of the ambulatory legs styliform and devoid of spines. The abdomen usually does not cover the whole width of the sternum between the last pair of legs. Species mostly small and littoral. Genus Uca Leach. Carapace trausverse, widest in front, the antero-lateral angles acute; dorsal surface smooth and granulated. Orbits large, extending to the lateral angles; eye-stalks long and slender. Merus of the maxillipeds generally transverse, smaller than the ischium, distally truncated, and not emarginate at the antero-internal angle; palp joined to the antero- external angle of the merus. Chelipeds in the male very unequal; hand in the larger cheliped very large, fingers longer than the palm. Smaller cheliped (both chelipeds in the female) feeble. CALIFORNIA STALK-EYED CRUSTACEA. 75 Uca crenulata (Lock. ) PLT, ¢.7-9. Gelasimus crenulatus LocKINGTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 149. Gelasimus vocator KINGSLEY (not Hrersst), Proc. Acad. Nat. Sci. Phila., 1880, p. 147. Gelasimus gracilis RatHBun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 244. Uca vocator ORTMANN (not Hersst), Zool. Jahrb. Abth. f. Syst., Bd. X, 1897, p. 352. ? : Uca gracilis RatHBUN M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 603. Carapace smooth, convex, with the H-shaped impression prominent. Front wide. The lower edge of the superior orbital margin is arched forward more strongly in the middle than the upperone. Lateral margins nearly parallel for a short distance behind the prominent antero-lateral angles of the carapace and then converging. Merus of the maxillipeds much broader than long and much shorter than the ischium. Merus of the iarge cheliped with the inner margin denticulated and the outer surface crossed by transverse granulated rugs; carpus with the outer surface granulated and the inner surface crossed by an oblique, usually granulated ridge; hand finely granulated on the outer surface; inner surface with a granulated or tuberculated ridge running from the lower margin to the carpal groove; there is often a line of granulations extending from the upper end of this ridge towards the upper margin of the hand; the carpal groove is deep and covered by the overhanging upper edge of the palm; fingers long and slender; the pollex is nearly straight or bent slightly upwards, and truncated at the tip; dactyl longer than the pollex, curving most strongly near the tip and generally armed with a tooth near the middle of the inner margin. Ambulatory legs pubescent, the merus transversely rugose. Abdomen of the male seven-jointed, the second joint very short. Todos Santos Bay near San Diego (Lockington); La Paz, Lower California! Guaymas! Sonora, Mexico! San Luis Gonzales Bay! Todos Santos Bay, La Paz, Lower California, San Diego (Miss Rathbun). The specimens of Miss Rathbun from La Paz and San Luis Gonzales Bay I have seen, and also Mr. Lockington’s types. The other specimens examined were in Lockington’s collection. This species is closely allied to pugnax Smith, but is distinguished by the more convex carapace, which is 76 CALIFORNIA -ACADEMY OF SCIENCKS. ° much wider behind; by the narrower abdomen in the male with the shorter second segment; by the more- slender fingers in the large cheliped, and the more slender merus joints in the ambulatory legs. It may be distinguished from coloradensis Rathbun by its darker and more convex carapace, the more slender merus joints in the ambulatory legs, and by the different curvature of the upper side of the palm in the large cheliped. It is a much smaller species than brevifrons Stimpson, and has a relatively wider and more convex carapace and very different upper orbital margins. Uca rectilata (Lock.) PLL, fF. 10-1. 7 Gelasimus rectilatus LockiIneTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 148. Gelasimus annulipes KINGSLEY (not MILNE-Epwarps), Proc. Acad. Nat. Sci. Phila., 1880, p. 148. Carapace wide, nearly flat transversely, but longitudinally strongly convex. Front broad and arched strongly forwards. The upper edge of the superior orbital margin is strongly curved; the lower edge runs close to the upper but is distinct; the lower margin of the orbit is prominently. denticulated, especiaily at the rounded outer angle. Postorbital angle prominent, acute, and directed obliquely outwards; the lateral margins of the carapace are straight, converging from the postorbital angles to the straight posterior margin. Maxillipeds with the ischium large, smooth, strongly convex; merus oblique, much wider than long, but narrower than the ischium and about one-third its length. A longitudinal groove on either side of the buccal area. Merus of the larger cheliped but slightly rugose with the angles rounded; carpus lightly granulated on the upper surface; hand similar to that of crenulata; the outer surface of the palm finely granulated and the lower edge margined; an oblique, granulated ridge on the inner surface extending from the lower margin to the deep carpal groove; no oblique ridge above this one as in crenulata; two parallel lines of granules behind the articulation of the dactyl, the posterior of which is continued upon the pollex; pollex tapering, a tooth near the middle, the extremity slightly excavated; dactyl curved more strongly toward the tip, which slightly overreaches the poilex. Ambulatory legs slender. Abdomen of the male seven-jointed, the first two joints short. CALIFORNIA STALK-EYED CRUSTACEA. Te henwthiet carapaces male si) oo0y eu ate nis 8.75 mm, Maetlt of earvapacoemulescs 621). 8 is A eee. 13.5 mm. Meneth Gi larger hand, male. co.cc) 80 hs Sek ee 19 mm. Tonweh of carapace, fenmles ty 6 522s ele Sele BAS 7.75 mm. Widta of carapace, female... 660.0565 ieee Be. \eoeTa2o mim: Described from Lockington’s types (Cat. No. 3112) in the collection of the California Academy of Sciences. West coast of Lower California (Lockington). This species differs from U. gibbosa (Smith) in having the surface of the carapace even, with the branchial regions not at all inflated, and in having seven, instead of five, abdominal segments in the male. U. stenodactyla is reported from San Diego by Ort- mann, who unites with this species gibbosa (Smith) and speciosa Ives’ . Genus Speocarcinus Stimpson. Carapace longitudinally convex both in front and behind, transversely nearly plane; sides converging behind. The space between the outer ends of the orbit exceeds one-half the width of the carapace. Antero-lateral margins dentate. Ocular peduncles of moderate length; eyes small. ‘Orbits and antenne similar to those of Panopeus. Palate devoid of a median elevation. Maxillipeds widely gaping; merus rather short and bearing the palp at the summit. Chelipeds short and stout. Ambulatory legs slender, smooth, with depressed, ciliated dactyls. Genital openings of the male in the sternum. Abdomen of the maie with the base much narrower than the last thoracic sternum; third, fourth and fifth segments coalesced. Type.—S. carolinensis STIMPSON. . Speocarcinus californiensis (Lock. ) Eucrate ? californiensis LocKineTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 33. Carapace strongly convex longitudinally, transversely plane, nearly smooth above, but minutely granulated towards the pubescent margins. Median region divided into three areas by pubescent sulci; a longitudinal, pubescent line on the branchial regions which are separated from the 1 See Zool. Jahrb. Abth. f. Syst., Bd. X, 1897, p. 356. 78 CALIFORNIA ACADEMY OF SCIENCES. hepatic by a transverse sulcus. Front over one-fourth the width of the carapace, the anterior margin straight and emarginate in the center. Upper margin of the orbit nearly straight, not interrupted by fissures; lower margin with an external fissure and a rounded inner lobe. - Ocular peduncles slender and about two-thirds as long as the width of the front and furnished anteriorly with two converging rows of long hairs. Antero- lateral margin strongly curved and furnished with three teeth (including the postorbital), the first two of which are thin-edged and lobate, the sec- ond one broadly rounded; the last tooth is acute. Buccal area widened in front. External maxiilipeds diverging anteriorly, the merus distally trun- cated, with the antero-external angle rounded and not produced. Chelipeds unequal; merus short, thick, trigonous, with the edges granulated, and furnished with a prominent tooth at the supero-distal angle; carpus with a spine at the antero-internal angle and a short, longitudinal, granulated ridge at the distal end of the outer surface; hands wide, much compressed, the outer surface nearly smooth but granulated near the upper and lower margins; upper edge acute and sharply granulated; fingers ridged, the pollex not deflexed. Legs smooth, glossy, pubescent; dactyls slender with strongly pubescent margins, the last pair and toa less extent the preceding pair upturned. First segment of the abdomen in the male short and wide, but much narrower than the last thoracic sternum; second segment a little longer but markedly narrower than the first; third seg- ment as wide proximally as the first, the sides converging rapidly to the coalesced fourth segment; remaining segments tapering gradually to the rounded extremity. Hength ‘of carapacer. s.65- soe e. ccc eee ene. Cee ete 16 mm. Width of carapace: Ur aue nc. eee Shee eee eee ee 21 mm. San Pedro, California. Lives in holes on muddy beaches. Quite closely allied to carolinensis St. from the Atlan- tic coast, but is distinguished by having three instead of five antero-lateral teeth, and by its longer eye- peduncles. From granulimanus Rath. it differs in the number of antero-lateral teeth, the smoother hands, and in having the antero-external angle of the merus rounded and not produced. Family GRAPSIDA. Carapace depressed or moderately convex, more or less quadrilateral, the lateral margins straight or slightly arcuate. Front usually broad, never very narrow. Orbits and eye-stalks of moderate size. Palp of the CALIFORNIA STALK-EYED CRUSTACEA. 79 maxillipeds joined to the summit or antero-external angle of the merus. Chelipeds in the adult male subequal and quite well developed; dactyls of the ambulatory legs styliform, compressed, and armed with strong spines. The base of the abdomen in the male usually covers the whole width of the last thoracic sternum. The species, with rare exceptions, are littoral or inhabitants of shallow water. Carapace striated. RE EERE PRIM taser oe, wad. 4 waalias Seats. Sieh Se © Pachygrapsus. Eins Womltemy GUEWALG © oe. lcs ck wan eae a bate Grapsodius. reap RIce Tn CO RTMAUCM. (22 ah cuiee chee shh ccs secels els Brachynotus. ; Genus Pachygrapsus Randall. Carapace trapezoidal, depressed, and marked with transverse striz. Front depressed and over one-half the width of the carapace. Inferior subocular lobe small, not reaching the front, thus allowing the antenne&x to enter the orbit. Eye-peduncles short and stout. Maxillipeds devoid of a *piliferous ridge and having a wide rhomboidal gap between them; merus as broad as or broader than long, distally truncated and bearing the palp at the summit. Chelipeds subequal, the merus trigonal, with the anterior margin distally dentated. Legs of moderate length, merus dilated and compressed, and furnished with an anterior subterminal tooth or spine; dactyls more or less spinulous. Antero-lateral margins entire or with one or two teeth. Abdomen in the male seven-jointed and covering the whole width of the last thoracic sternum. Type.—FP. crassipes RANDALL. Pachygrapsus crassipes Randall. Pachygrapsus crassipes RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 127. H. M1LnE-Epwarps, Ann. Sci. Nat. (3), T. XX, 1853, p. 166. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 467. LocxineTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 153. STREETS, Bull. U.S. Nat. Mus., No. 7, 1877, p. 115. Kinesuey, Proc. Acad. Nat. Sci. Phila., 1880, p. 199. Dr Man, Notes Leyden Mus., Vol. XII, 1890, p. 86, Tab. V, fig. 11. Ornrmann, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894, p. 707. Ratusun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 604. Carapace nearly square (the sides converging slightly posteriorly), moderately convex, the entire upper surfaee, except the cardiac and intes- tinal regions, transverselystriated. The front is broad, depressed, slightly ei} 80 CALIFORNIA ACADEMY OF SCIENCES. arcuated; a single tooth at the outer angle of the front; behind the front are four lobe-like prominences, the two middle ones the more conspicuous, and separated from each other by a groove whose sides are marked with sharp lines. A single tooth on the lateral margin behind the prominent postorbital. The notch at the infero-external angle of the orbit is deep. Orbits deep, the inferior margins strongly granulated. Antenne minute. Merus of the maxillipeds quite strongly produced at the antero-internal angle; first joint of the palp large and compressed. Merus of the cheli- peds short, trigonal, and transversely rugulose, the anterior margin laminate and armed at the distal end with three teeth; carpus finely rugose externally and bearing a tooth or tubercle at the antero-internal angle; hand in the adult male larger than all the preceding joints combined; a raised line usually present on the upper side of the palm and a delicate line on the lower portion of the outer surface. Merus joints of the legs transversely rugulose, strongly compressed and expanded, especially in the last three pairs, the postero-distal angles in the anterior pairs armed with a variable number of teeth; in the last or last two pairs they are rounded and usually edentate; dactyls short, compressed, and spinous on both margins. Third joint of the abdomen in the male strongly convex at the sides; following joints increasing successively in length. Length oftarapace,mmale:h. foie. 5) cele alk See 36.5 mm. Breadth /ofiearapace, male... ind. 2 2) tel ack cleo ra ele des 41 mm. hength oficarapace: temale 5s... cava. os eee ee 30 mm. Breadth of carapace, females icc. 4. 4-2 oie) 2 sie dee 34 mm. Japan (Stimpson, Ortmann), Sandwich Islands (Ran- dall), Oregon! northern California! San Francisco Bay! Monterey! San Diego! Gulf of California (De Man). A very common species on rocky shores. This species was originally reported by Randall from the Sandwich Islands, where it has not since been found. On the other hand, P. parallelus, which Randall reported as coming from the Columbia River, has not since been found on the west coast of the United States. It is probable, as Stimpson suggests, that the labels of the two species were accidentally exchanged. Several other species reported from this coast by Randall have not since come to light. Possibly mixing of labels has occurred in other cases also. CALIFORNIA STALK-BYED CRUSTACEA. 81 Genus Brachynotus De Haan. Carapace subquadrate, not markedly striated, and having two teeth behind the orbital angle. Front wide, not strongly deflexed. Maxillipeds without a wide rhomboidal gap; ischium transversely truncated; merus large and not produced at the antero-external angle; palp joined in a notch at the middle of the distal margin. Type.—B. sexdentatus (Risso). Brachynotus nudus (Dana). Pseudograpsus nudus Dana, Proc. Acad. Nat. Sci. Phila., 1851, p. 249; Crust. U. S. Expl. Expd., Part 1, 1852, p. 335, Pl. XX, fig. 7. Srmmp- son, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 469. Heterograpsus nudus Stimpson, Proc. Acad. Nat. Sci. Phila., 1858 p. 104. © Lockineron, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 152. Waurr- EAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. Smirns, Rep. Prog. Geol. Surv. Canada, 1878-9, B, p. 206. Ratusun R., The Fisheries of the U.S., Sec. 1, 1884, p. 765. Newcomsez, Bull. Nat. Hist. Soc. Brit. Col., 1895, p. 25. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. VII, 1894, p. 715. Wauxker, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 273. Heterograpsus sanguineus KINGSLEY (not De Haan), Proc. Acad. Nat. Sci. Phila., 1880, p. 208. Carapace posteriorly flat, smooth and punctate, anteriorly convex, un- dulated, and furnished with small, scattered granules. A curved line of light-colored pits extending from the anterior end of the H-shaped im- pression to the last antero-lateral tooth. Front bilobed, the median emargination shallow, but comparatively broad. The two transverse prominences behind the front are evenly rounded. Antero-lateral margins strongly arcuated and granulated; the second tooth behind the acute postorbital much smaller than the preceding ones. Posterior margin of the epistome granulated, the arcuated median portion separated from the subquadrate, projecting, lateral portions by a conspicuous, deep, smooth interval. On the pterygostomian regions a ridge running parallel with the antero-lateral margin and armed with numerous smooth, small, rounded denticles or granulations. Maxillipeds punctated; a groove behind the broad, raised, inner margin of the merus. Chelipeds smooth, and mottled above with small, round, red spots; antero-internal angle of the merus in the adult male produced into a rounded, smooth lobe; carpus with a subacute prominence on the inner side; hands smooth, with a fine longitudinal ridge along the lower side of the outer surface; fingers with small, corneous tips. In the males the hands are inflated and bear a large G 82 CALIFORNIA ACADEMY OF SCIENCES. patch of long, fine hair on the inner surface. Legs rather short, smooth, punctate and nude; dactyls short, stout, scabrous; those of the last pair less than two and one-half times (often less than twice) as long as wide, and upturned at the tip. © The color of this species is quite variable. It is gen- erally of a mahogany red, but may be purplish, dark red, or red marbled with white. I have seen some spec- imens with the upper side almost entirely white. But amid all the variations of color, the red spots on the chelipeds remain, so far as I can determine, an abso- lutely constant character. Young specimens present greater color variations than older ones. Length. Breadth. Maley yc. Van a eco ee APL BIN TNs soic.'s .. "ast ere em eneeiets 51.5 mm i ae Re ey Py ENE, 8 AS, TANUM Soi +. eer nyt pe EE 50 mm TEU a SANT 5 cing clital eae eae eas ice Pi Hie 18 CO Ree ae Rrra ORL TES, 26 mm Bema.) yb eee Pree i aate Dh MINTS 2, snp ee arc tenee Se 25.5 mm Sitka (Kingsley); Vancouver’s Island (Smith); Puget Sound; common from Oregon to Lower California; Gulf of California (Kingsley). Found among rocks near the shore. ' This species differs from sanguineus (De Haan), with which it has been united by Kingsley, in having a tuft of hair on the inner side of the hand in the male, and in the different inferior orbital margin. Brachynotus oregonensis (Dana). Pseudograpsus oregonensis DANA, Proc. Acad. Nat. Sci. Phila., 1851, p. 248; Crust. U. S. Expl. Expd., Part I, 1852, p. 334, Pl. XX, fig. 6. Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 468. Mintnz-Epwarps, Ann. Sci. Nat. (3), Zool., T. XX, p. 157. Cooprr, Rep. Expl. and Sur. to Pac. Ocean, Vol. XII, 1860, Book 2, p. 389. Srrmpson, Proc. Acad. Nat. Sci. Phila., 1858, p. 104. LockineTon, Proc. Cal. Acad. Sci., Vol. VII, 1876, p. 152. Kinastey, Proc. Acad. Nat. Sci. Phila., 1880, p. 209. RATHBUN, R., The Fisheries of the U. S., 1884, Sec. 1, p. 765. CALIFORNIA STALK-EYED CRUSTACEA. 83 Carapace more strongly undulated in front than in nudus but not so much flattened behind; the posterior portion is not punctate and the anterior portion may be sparsely granulated. Front four-lobed, the median lobes the most prominent. The median portion of the epistome is not granulated nor separated from the lateral portions by a deep smooth interval. The ridge on the pterygostomiau region furnished with several smooth, blunt teeth. The prominences behind the front are more conspic- uous than in nudusand sometimes end abruptly anteriorly. External max- illipeds smooth and punctated; surface of the merus next to the inner margin flat. Chelipeds smooth; carpus with a prominent inner angle; hands with a fine longitudinal ridge on the lower part of the lower surface which becomes very faint with age. In the adult male there is a rounded lobe on the antero-internal of the merus and a patch of long hair on the inner side of the hand. Legs hairy, the dactyls narrower than in nudus, those of the last pair upturned. The color is a dull gray mottled with ferruginous spots. The spots on the legs are small, but there may be blotches of considerable size on the carapace. The color is not so variable as in the preceding species although very young specimens are occasionally marked with large blotches of white. Length, 23 mm.; breadth, 28 mm. This species may readily be distinguished from the preceding by its dull color, the hairiness of the legs, the four-lobed front, and the absence of round red spots on the chelipeds so characteristic of that closely related species. Nudus is found chiefly among the rocks, while oregonensis prefers the mud flats, where it is usually found in abundance. Vancouver’s Island. Common on mud flats from Puget Sound to Lower California., It is also found among rocks near the shore. —— Grapsodius, gen.nov.. Carapace striated above, with the sides converging behind, and armed witha single tooth behind the postorbital. Front broad, not deflexed, but with the median portion depressed. Eye-peduncles short. Orbits with the posterior surface bulging outwards instead of concave. Maxillipeds narrow, widely gaping, and devoid of an oblique piliferous ridge; merus 84 CALIFORNIA ACADEMY OF SCIENCES. subcordate, shorter than the ischium, the antero-internal angle produced; palp joined near the middle of the distal margin of the merus. Dactyls spinulous. Abdomen of the male seven-jointed. Type.—G. eximius HoLMEs. / Grapsodius eximius, sp.nov. “© 2c. Carapace undulated in front and flattened behind where it is more strongly striated; sides strongly converging posteriorly. The front is over one-half the width of the carapace and has the outer angles more or less projecting and rounded; the anterior edge is thin and minutely granulated; viewed from above it is nearly straight, being slightly convex on either side of the middle where it is a little concave; viewed from in front it sags downward in the center. The orbits are remarkable in being swollen outward so that there is no hollow receptacle, as is usually the case, for the reception of the eyes; the superior orbital margin is marked by a fine ridge extending from the outer side of the front to the postorbital tooth; the inferior orbital margin is marked by a line of granules extend- ing from the lower side of the postorbital tooth to the buccal area. Max- illipeds slender and wide apart. The ischium is much longer than the merus but not so wide; merus with the outer margin convex and the antero-external angle broadly rounded; the inner margin is straight and the antero-internal angle is produced into a prominent narrow lobe; first joint of the palp strongly convex near the middle of the inner margin; exognath at the base about one-half as wide as the ischium and tapering regularly to the tip which reaches slightly beyond the middle of the merus. Chelipeds subequal; merus short, trigonal, the outer surface transversely striated, the inner margin produced into a laminate expansion which is distally truncated and dentate; carpus with a spine near the middle of the upper margin; hands smooth and inflated; the upper margin of the palm is broadly rounded, but bears a fine ridge; a very fine ridge on the lower side of the outer surface extending upon the pollex; fingers subcylindrical, not ridged or grooved, and armed within with small teeth. Merus of the ambulatory legs dilated and compressed much as in Pachygrapsus cras- sipes, with the upper margins acute and ending in a tooth a little behind the supero-distal angle; the infero-distal angle, in all but the last pair, is dentate; carpal joints with a few small spines near the distal end of the upper margin; propodi with the sides strongly convex and the upper and lower margins spiny; dactyls rather narrow, shorter than the propodi, strongly spinose above and below, and terminating in slender claws. The abdomen in the male is widest at the third segment, behind which it tapers to the tip, the sides converging more rapidly towards the posterior end; first segment much longer than the second; third segment about as long CALIFORNIA STALK-EYED CRUSTACEA. 85 as the fourth, the sides strongly convex; fifth segment scarcely longer than _ the fourth and shorter than the sixth; last segment triangular, acute. Length of carapace, 18.5 mm.; width of carapace, 21 mm.; width of front, 11.2 mm. San Diego, California. Collected by A. U. Crawford. ee This species is described from a single dried and somewhat imperfect specimen contained in the Museum of the University of California. The form and arrange- ment of the antenne and antennules could not be determined, nor could I make out whether or not the’ inferior orbital lobe was in contact with the front. The front, legs, maxillipeds, and striations on the carapace are similar to those of Pachygrapsus, but the character of the orbits separates it from that genus as. well as all the other genera of the Grapside. Family PINNOTHERID&. Carapace convex or depressed, often more or less membranaceous, the antero-lateral margins entire or very slightly dentate. Front narrow. Orbits and eye-peduncles very small. Buccal area convex anteriorly. The merus and often the ischium of the maxillipeds is weli developed, and the palp may be joined to the summit, antero-internal angle, or, rarely, the antero-external angle of the merus. Chelipeds usually small, or of moderate size. Ambulatory legs variable; dactyls generally styliform and not armed with spines. The members of this family are generally of small size, and most of them inhabit the shells of bivalved molluscs. Some species inhabit the intestine of echino- derms and others the tubes of annelid worms. The Pinnotheride are divided by Miers into four subfam- ilies, only two of which are found in our limits. Subfamily PINNOTHERIN~A. Carapace usually convex, subglobose, or transverse. Front not ros- trated. Ischium of the maxillipeds rudimentary or absent. The ambu- latory legs are all well developed (the last pair is sometimes small); dactyls often short. 86 CALIFORNIA ACADEMY OF SCIENCES. Genus Pinnotheres JLatr. Carapace smooth, subglobose, more or less membranaceous and scarcely wider than long. Orbits small, nearly circular. Front narrow, with the anterior margin nearly straight. Antennules transverse. Maxillipeds oblique, the merus large, usually curved, the last joint of the palp joined to the inner margin of the preceding one. Ambulatory legs subequal and of moderate length. Pinnotheres nudus Holmes. Pinnotheres nudus Houmss, Proc. Cal. Acad. Sci., 2d Ser., Vol. IV, 1895, p. 563, figs. 1-5. Carapace a little broader than long, subquadrate to orbicular in outline, curving downwards towards all the margins; surface smooth and naked. Front rounded, deflexed, not protruding. Orbits ovate. Antennules oblique. Maxillipeds oblique, neatly fitting the buccal area; merus broad, smooth, subquadrate, the outer margin produced into a broadly rounded, ~ laminate expansion; penultimate joint oblong and distally rounded; last joint spatulate, articulated near the base of, and extending somewhat beyond the preceding one. Chelipeds rounded, smooth, devoid of spines or teeth; hands narrow, rather thick; fingers subconical, subequal to the palm. The three anterior pairs of ambulatory legs subequal; dactyls nearly straight; those of the fourth pair longer and more slender than the others. Length, 20 mm.; breadth, 24 mm. Santa Cruz (Dr. Anderson’s Coll.)! Monterey! Pinnotheres pugettensis, sp. nov. Carapace soft, smooth, subpentagonal. Front triangular, acute, curved downwards, scarcely protruding beyond the general contour of the cara- pace. Orbits nearly circular. Antenne shorter than one-half the width of the front. Maxillipeds very oblique, strongly pubescent; merus nar- rower than in nudus, the outer margin convex; penultimate joint broad, subquadrate, distally truncated, last joint minute, joined slightly in ad- vance of the middle of the preceding one but scarcely reaching beyond the tip. Chelipeds smooth; merus short, the upper margin furnished with long hairs; hands narrow, elongated, rounded, smooth, the dactyls and inner side of the palm short/ pubescent; fingers subcylindrical, nearly straight, a little shorter than the palm, the tips strongly hooked, dactyl with a low tooth near the base of the inner margin. Ambulatory legs slender, increasing slightly in length posteriorly; propodi hairy above and EE CALIFORNIA STALK-EYED CRUSTACEA. 87 below; dactyls narrow, compressed, convex above, abruptly contracted near the tip into a short, curved claw; in the three anterior pairs the dactyls are shorter than the propodi and leave the lower margin nearly straight; in the last pair the dactyl is much longer than the propodus, much longer than the preceding dactyls, and has the lower margin con- eave. The outer surface of the palm is. brownish with light-colored reticulations. Length of carapace, 10 mm.; length of first ambulatory leg, 9.5 mm. Width of carapace, 10.5 mm.; length of last ambulatory leg, 10.6 mm. Described from a single female specimen found by Professor W. E. Ritter in the branchial cavity of a species of Cynthia from Puget Sound. Collection of the University of California. Genus Fabia Dana. Closely allied to Pinnotheres. Carapace smooth, more or less mem- branaceous, not much broader than long, and marked by a pair of longi- tudinal sulci which extend backward from the upper margin of the orbits, enclosing between them the median area. Front not rostrated. Eyes normal, not widely separated. Antennulary fossettes not widely separated. Maxillipeds with the merus large and the ischium rudimentary; third joint of the palp articulated on the inner margin of the preceding one. Ambulatory legs subequal and rather slender; dactyls short. Type.—f. subquadrata DANA. This genus differs from Pinnotheres in no important character except the longitudinal sulci on the carapace. Fabia subquadrata Dana. Fabia subquadrata Dana, Proc. Acad. Nat. Sci. Phila., 1851, p. 253; Geust, U.S. Expl. Expd., Part © 1852, p. 382, Pl. XXIV, fig. 5. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 470. Locx- INGTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 155. Smiru, Rep. Prog. Geol. Surv. Canada, 1878-9, B, p. 206. Nrewcomss, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 25. Carapace smooth, glossy, membranaceous, subquadrate in outline, with the angles broadly rounded; the space between the longitudinal sulci is longer than wide and slightly narrowed behind; antero-lateral margin rounded and marked by a raised line. Front very short and turned 88 CALIFORNIA ACADEMY OF SCIENCES. abruptly downwards, having no trace of a median or transverse groove. Eye-stalks short, stout, rounded and lodged in nearly circular orbits. Antennules obliquely plicated in very wide fossettes. Antennz minute. Maxillipeds very oblique; merus large, smooth, curved and distally rounded; the penultimate joint of the palp is broad, flattened, and bears the minute third joint near the middle of the inner margin. Chelipeds smooth and rather slender; hand long and narrow; palm about twice as long as wide, with the upper and lower margins parallel, and having one or two rows of hairs below, one of which reaches to the tip of the finger; fingers longitudinal, subcylindrical, nearly straight, and shorter than the palm; dactyl with a tooth a little behind the middle of the inner margin. Legs slender, glossy, and nearly naked, but the upper side of the merus and the lower side of the propodus are generally more or less pubescent; dactyls pubescent, about half the length of the propodi, and subuncinate at the tip. Color in life, whitish; carapace and abdomen largely covered with orange. Length of carapace, female, 11.5 mm.; width, 13 mm. The following measurements are given by Dana from his specimen from Puget Sound: Length of carapace, 5.75 lines; breadth, 6.75 lines; breadth between post-frontal sutures, 2.25 lines; length of third joint of fourth pair of legs, 2.75 lines, or about two-fifths the breadth of the carapace. Queen Charlotte Is. (Smith), Puget Sound (Dana), Farallon Is. and San Diego (Lockington), San Pedro! in shell of Tapes. It is found also in the shell of Pachy- desma crassitelloides and sometimes in the tests of Echini. Genus Pinnixa White (emended). Carapace much wider than long. Front narrow, nearly transverse. Orbits broadly ovate or nearly circular, with a wide inner hiatus which is partly occupied by the basal antennal joints. Antennules transversely or obliquely plicated in wide fossettes which communicate with each other beneath the front. Eye-stalks very short. Epistome linear-transverse. Ischium of the maxillipeds small; merus large, the distal portion of the outer margin convex; palp joined to the summit of the merus, the third joint joined on the inner side of the preceding one near the base. Cheli- peds of moderate size; merus trigonous; carpus smooth; hand large, com- pressed. Second ambulatory legs larger than the first; the third pairis the largest of all; fourth pair much shorter than the third and relatively stouter than the first and second. - The abdomen in both sexes is seven- jointed and narrower at the base than the width of the last thoracic sternum. Type.—P. cylindrica Say. CALIFORNIA STALK-EYED CRUSTACEA. So. Pinnixa occidentalis Rath. Pinnixa occidentalis RATHBUN, Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 248. Nerwcomss, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 26. Male: Carapace uneven, about twice as wide as long; median region tumid and bounded by sulci on either side and behind; a transverse crest on the cardiac region which is not interrupted in the middle but becomes lower and curves backwards. Behind this crest, which is more prominent in the male, the carapace slopes rapidly downward, being nearly at right angles to the surface in front of it. Front narrow, depressed, furnished with a median groove, and not projecting beyond the general contour of the carapace; a pair of prominences behind the anterior margin which is slightly produced at the center. The antero-lateral margin is marked by a sharp, more or less granulated line which begins a short distance external to the orbit and meets the postero-lateral margin ata little less than a right angle. Antenne longer than the width of the front. Maxillipeds nearly longitudinal; second joint of the palp narrow (over twice as long as wide), and tapering slightly from the base to the rounded tip; last joint narrowly spatulate (about four times as long as broad), joined near the base of and extending much beyond the preceding one; outer sur- face of the last two joints sulcated. Chelipeds stout; hand broad, ' smooth and shining, slightly widened distally, the upper and lower mar- gins rounded and scabrous; pollex broad, short, deflexed and furnished with one or more teeth on the inner margin; dactyl much curved, some- times having a minute tooth near the middle of the inner margin. The first pair of ambulatory legs is slender, having slender dactyls which are subequal to the preceding joint; second pair similar to the first but longer and stouter; third pair longer and much stouter than the second, but more slender. than in tubicola or littoralis; the dactyls slender, almost straight, and about equal to the propodi; fifth pair stouter than the first and nearly as long, reaching considerably beyond the merus of the preceding pair; dactyls slender, straight, and about equaling the propodi. The abdomen tapers gradually from the second segment to the last, which is rounded; the third, fourth, and fifth segments are subequal and slightly longer than the sixth. The appendages and sides of the carapace are more or less pubescent. The upper and lower margins and, to a less extent, the surfaces of the ambulatory legs, are scabrous. Length of Carapace. Breadth of Carapace. Length of Third Ambulatory Leg. (a ATA: 2 ono ak eels | IBY i it ot 0d ze a 19 mm ete sas sae Cea (. ODD eee 13 mm. Described from specimens received from Miss Rath- bun. | Monterey Bay and off Point Ano Nuevo, California; Magdalena Bay, Lower California (Rathbun). The locality of the specimens described is lat. 36° 47’ 50” N.; lon. 121° 49” W.; depth, 37 fathoms. CALIFORNIA STALK-EYED CRUSTACEA. 91 Pinnixa tubicola Holmes. Pinnixa tubicola Hotes, Proc. Cal. Acad. Sci., 2d Ser., Vol. IV, 1895, p. 569, Pl. XX, figs. 17 and 18. General form subcylindrical. Carapace about two and one-half times. as a loreal strongly curved downward towards the anterior margin and at the sides. A shallow, transverse depression behind the gastric area, behind which is a convex (not crested) transverse intumescence, from which the carapace curves sharply downwards towards the broad, slightly concave posterior margin. Front short, deflexed, not projecting beyond the general contour of the carapace. Outer portion of the antero- lateral margins defined bya ridge. The last joint of the palp of the maxillipeds is spatulate and joined near the base of, and extending con- siderably beyond the preceding one. Chelipeds rather small; hand oblong, longer than the preceding joints combined; fingers hooked at the tips, their inner margins meeting when closed. First pair of ambulatory legs. ’ slender, with slender dactyls which about equal the length of the propodi;. second pair much longer and stouter than the first, the dactyls relatively stouter than in the first pair and a little shorter than the propodi; third pair stouter and a little longer than the second and furnished with shorter and stouter dactyls, which are markedly shorter than the Eropealy fourth pair similar to but much shorter than the third. Length of carapace, 4mm.; width, 10 mm. Puget Sound! Trinidad! Cape Mendocino! Bodega Bay! San Pedro! San Diego! Pinnixa littoralis Holmes. Pinnixa littoralis Houtmess, Proc. Cal. Acad. Sci., 2d Ser., Vol. IV, 1895, p. 571, Pl. XX, figs. 14-16. Carapace naked, flattened above; a transverse depression behind the median region followed by a transverse intumescence. The front is not strongly depressed and projects beyond the general outline of the cara- pace; it is anteriorly truncated and furnished with a median groove. Maxillipeds similar to those of tubicola; the terminal joint extends only a short distance beyond the preceding one.. Chelipeds large, smooth; hand large, compressed, oblong, but widening slightly distally; pollex short, directed obliquely downwards and furnished at the tip with a notch, into which the point of the dactyl closes; dactyl curved, and when closed against the pollex, a large rounded space is left between them. Ambula- tory legs almost entirely naked, more slender than in tubicola, first pair but little shorter than the second, which is but little shorter than the 92 CALIFORNIA ACADEMY OF SCIENCES. third; the dactyls in all the legs are little, if any, more than one-half the length of the propodi and are curved at their corneous tips. Abdomen of the male tapering evenly from the base to the tip; the sides of the penul- timate joint are concave, those of the remaining joints straight. Length of carapace, 4.5 mm.; breadth, 9.5 mm.; length of third ambu- latory leg, 11 mm. Bodega Bay! near Fort Bragg, California! Pinnixa longipes (Lock. ). Tubicola longipes Locx1neTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 55. Pinnixa longipes Locxineton, Jbid., Vol. VII, 1877, p. 156. Streets and KINGSLEY, Bull. Essex Inst., Vol. IX, 1877, p. 107. Houtmes, Proc. Cal. Acad. Sci., 2d Ser., Vol. IV, 1895, p. 573, Pl. XX, figs. 19 and 20. Carapace considerably more than twice as wide as long, somewhat flat- tened above and furnished with a transverse depression behind the gastric area. Front slightly projecting, furnished with a median groove and a transverse groove behind the anterior margin. The last joint of the palp of the maxillipeds is spatulate and joined near the base of, and slightly exceeding the preceding joint. Chelipeds small, short, hairy; hands oblong, compressed. First two pairs of ambulatory legs slender (the second somewhat larger than the first), and furnished with slender, nearly straight dactyls which are about equal in length to the propodi; following pair of legs enormously developed; merus with a kind of flange on the posterior margin; dactyl stout, curved, much shorter than the propodus; last pair small and stout, scarcely reaching beyond the middle of the merus of the preceding pair; dactyl stout, curved, shorter than the propodus. Tomales Bay (Lockington)! San Pedro, Calif.! This species lives in the tube of an annelid worm (Clymenella). It forms the extreme point of modifica- tion of this peculiar genus. There is probably no other crab which has such great width relatively to its length, there is certainly no known species in which the fourth pair of pereopods is so enormously enlarged; and I believe there is no Brachyuran which exceeds it in smallness of size. | ee ee ee ee CALIFORNIA STALK-EYED CRUSTACEA. 93 Pinnixa faba Dana.) Pinnitheres faba Dana, Proc. Acad. Nat. Sci. Phila., 1851, p. 248; Crust. U.S. Expl. Expd., Part I, 1852, p. 381, Pl. XXIV, fig.'4. Bars, in Lord’s Nat. in Vancouver’s Island, Vol. II, 1866, p. 271. Pinnixa faba Stimpson, Journ. Bost. Soc.. Nat. Hist., Vol. VI, 1857, p. 470. Cooprr, Rep. Expl. and Surv. Pac. Ocean, Vol. XII, Book 2, 1860, p. 387. HAsweE tu, Cat. Australian Crust., p. 113. NEwcomBg, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 25. Carapace strongly convex, both longitudinally and transversely; no transverse ridge behind the gastric area. Antero-lateral margins marked by a low ridge which disappears near the orbits. Front slightly project- ing and divided into tworounded lobes. Antennuies transversely plicated. Antenne much longer than the width of the front. The penultimate joint of the palp of the maxillipeds projects considerably beyond the ter- minal one; both joints are more or less sulcate on the outer surface. Hands of the chelipeds flattened, widest just behind the articulation of the dactyl, and more or less pubescent on the inner side between the fingers; fingers gaping at the base; dactyl curved, acute and furnished with a tooth near the middle of the inner margin; and (generally) a row of hairs on the upper edge; pollex short, nearly straight, and obliquely truncated at the tip. First pair of ambulatory legs shorter than the sec- ond; dactyls subconical from a stout base and slightly curved; third pair longer and stouter than the second, the dactyl similar in form though stouter; last pair larger than in most of the species, the propodus reach- ing beyond the merus of the preceding pair; dactyls more slender than those of the third pair. Upper and lower edges of the ambulatory legs (generally) quite strongly pubescent. Abdomen of the male tapering evenly to the penultimate joint, which tapers more rapidly than the pre- ceding ones; last joint about as long as wide and rounded at the tip. Vancouver’s Island (Smith); Queen Charlotte Island (Newcombe); Puget Sound (Dana); Shoalwater Bay (Cooper); San Pedro, Calif. ! Dr. H. P. Johnson found several specimens of this species in the cloaca of a large species of Holothurian, Liosoma arenata St. It more commonly occurs in the shells of large bivalved molluscs. 94 CALIFORNIA ACADEMY OF SCIENCES. Pinnixa (Scleroplax) granulata (Rath.). Scleroplax granulatus Ratuspun M., Proc. U. S. Nat. Mus., Vol. XVI, 1893, p. 251. Carapace strongly convex, curving downward towards all the margins; sides rounded. Front very narrow (not one-fifth the width of the cara- pace), somewhat produced, almost transverse (slightly convex in the center), and uniformly convex above, having no median or transverse groove. No prominences behind the front. Median region not marked by sulci. Surface of the carapace more or less granulated anteriorly and near the margins, elsewhere smooth and punctate. Antero-lateral margin defined by a finely granulated ridge which runs parallel to the inferior margin of the carapace until very near the point, where it disappears, when it bends toward the lower margin but is not continued far enough to meet it. Orbits nearly circular. Antennules nearly transverse. Max- illipeds nearly longitudinal; merus broad, not curved; second joint of the palp narrow and longer than the merus, tapering gradually to its rounded tip; third joint rather narrow, spatulate, joined near the base of, and slightly exceeding the preceding joint; both of the last joints are grooved on the outer surface and fringed with long hair. Chelipeds of the female small, granulated, shorter than the following legs; hands pubescent, of ‘moderate size, margins rounded; palm more or less inflated; pollex nearly longitudinal; dactyl curved. Ambulatory legs slender, compressed, not markedly unequal; dactyls very long, slender, almost straight, about equaling the propodi in all the pairs, and furnished with long, sharp, corneous tips; the first pair is somewhat smaller than the second, which ‘is smaller than the third pair. Abdomen of the female smooth and shining. Length of carapace, female, 7.75 mm.; breadth, 10.75 mm.; length of third ambulatory leg, 11 mm. Ensenada, Lower California (Miss Rathbun); Bodega | Bay, Calif.! The genus Scleroplax cannot be sustained, I believe, “as some undoubted Pinniwas have a hard carapace and there are transition forms between granulata, in which the third pair of ambulatory legs is but little larger than the others, and the species in which the third pair is greatly developed. There are similar gradations between the very wide forms like longipes and the nar- rower species like granulata and faba. CALIFORNIA STALK-EYED CRUSTACEA. 95 Genus Parapinnixa Holmes. Carapace much broader than long, the anterior margin nearly straight. Frontal process defiexed. Orbits nearly round. Antennules transversely or obliquely plicated, the fossettes communicating with each other beneath thefront. Buccal areasmall, subtriangular. Maxillipeds with the ischium rudimentary, the merus large, triangular; palp tb jh three-jointed, the last joint joined to the tip of the preceding one. First pair of ambulatory legs the largest, the others diminishing successively in length, the last pair being quite small. Abdomen of female small, not nearly covering the sternal area. Type.—P. nitida (Lock.) Parapinnixa affinis, sp. nov. Closely allied to P. nitida, but the carapace is less than twice as wide as long, while in that species the width of the carapace is over twice its length; the surface of the carapaceis smooth and shining and the anterior margin straight. Front triangular, depressed, having a short median groove. Antennules oblique. Buccal area small, triangular, rounded in front, the posterior portion partly covered by a projection of the sternum. Chelipeds stout, smooth; hand thickened, smooth, rounded above and below; dactyl hooked at the tip and armed with a small tooth near the middle of the inner margin, the upper side smooth; pollex with two teeth at the tip. First pair of ambulatory legs larger than the others; dactyls short and stout. Next two pairs comparatively slender, having longer dactyls; last pair small, reaching about to the tip of the merus of the preceding pair; dactyls short and stout. Abdomen of the female widest at the third segment, behind which its shape is triangular, the tip broadly rounded. Described from a single female specimen collected at Dead Man’s Island, San Pedro, Calif., by Mr. F. W. Bancroft, July, 1895. Collection of the University of California. The maxillipeds in this species are similar to those of nitida. They were accidentally lost after I had removed them from the specimen; the palp appeared to have but two joints, but the examination was not made with sufficient care to make me feel sure of this point. 96 CALIFORNIA ACADEMY OF SCIENCES. Genus Cryptophrys Rathbun. Carapace subpentagonal, hard, convex, not much broader than long. Front produced. Orbits nearly circular and lodged in the sides of the front, only a small part of them visible from above. Antennules oblique or transverse. Buccal area much curved. Maxillipeds very oblique, the distal extremities (not as is usual the inner margins) being opposite each other and separated by a space in which the palpi are lodged; ischium _ rudimentary; merus large and curved; palp two-jointed and joined to the antero-external angle of the merus. Chelipeds moderately stout, the palm in the male broad and inflated. Ambulatory legs compressed; first three pairs subequal; fourth pair shortest. The base of the abdomen in the male does not cover the whole of the last thoracic sternum. Type.—C. concharum RATHEUN. Cryptophrys concharum Rath. Cryptophrys concharum RATHBUN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 250. Carapace smooth, slightly longer than broad; a faint sulcus behind the gastric region but no longitudinal sulci as in pubescens. Front emargin- ate. Antennules large, nearly transverse. Ocular peduncles short, stout, and completely filling the orbits. Antero-lateral margins marked bya ciliated line. Sides of the carapace broadly rounded. Merus of the max- illipeds with the outer margin curved, the distal portion almost transverse; the last joint of the palp four-sided, the extremity widened. Chelipeds and ambulatory legs margined by a row of coarse hairs. Ambulatory legs widened and quite strongly compressed; second pair a little longer than _the others; fourth pair overreaching the carpus of the preceding pairs; dactyls short, terminating in slender, curved, corneous tips. Abdomen in the male widest at the second joint; the large, compound segment has the margin at first convex and then slightly concave; the two following segments broader than long; terminal segment subrectangular but distally rounded; the last two segments may be partly coalesced. Length, 4.7 mm.; breadth, 4.2 mm. False Bay, San Diego County, Calif., from the mantle of Mya arenaria L. and Puget Sound from the mantle of Cardita borealis Conrad (Miss Rathbun). San Diego! Described from specimens from Puget Sound kindly sent me by Miss Rathbun. A specimen I collected at San Diego was found on the beach free from any host. CALIFORNIA STALK-EYED CRUSTACEA. 97 Subfamily ASTHENOGNATHIN. External maxillipeds with both ischium and merus well developed. Last pair of ambulatory legs not rudimentary or abortive. Genus Opisthopus Rathbun. **Carapace usually firm and unyielding; smooth, subquadrilateral, regions not defined; lateral margins regularly arcuated. Epistome very short. Abdomen seven-jointed, in the male not covering the sternum between the coxe of the last pair of ambulatory legs. EKye-peduncles short. Antennz small, situated at the inner orbital hiatus; basal joint small. Antennulz obliquely plicated. External maxillipeds with the ischium well developed, the merus broad, the palpus three-jointed, the ultimate joint articulated on the inner side of the penultimate. Chelipeds moderate; ambulatory legs subequal in length, joinis flattened.” Type.—O. transversus RaTHBUN. Opisthopus transversus Rath. Opisthopus transversus RaTHBUN M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 252. Carapace thin, convex, transverse. Front deflexed, with aslight median sulcus. Ischium of the maxillipeds broad; merus as broad as long, with the antero-external angle broadly rounded; palp large, last joint ‘‘ narrow, inversely spatulate, overreaching the penultimate joint. Chelipeds rather stout; merus broad, trihedral; palm a little longer than the fingers, thick, slightly compressed, margins rounded, lower margin convex.” Ambula- tory legs similar; second pair longest; fourth pair shortest; dactyls small, curved, little more than half the length of the propodi. Length of Carapace. Width of Carapace. 10 ge eae Aa Ra TMI eS ty! 5 cy a hee ee can te Ee 9.8 mm MSRATE oe hs 8 wd 1 ROMER: 0 13 ee GM aan SOc a Se eee Re 13; mm CPR rt, Smee eerie PES GT CARS Sh cane Shy. 2G 3 18 mm. Monterey and Point Loma (Miss Rathbun). One male from the shell of Lucapina crenulata Sowerby from Monterey. San Diego! / 98 CALIFORNIA ACADEMY OF SCIENCES. Subtribe OXYSTOMATA, or LEUCO- SOIDEA. Carapace very variable in form. Epistome rudimentary. Buccal area narrowed in front. Antennules longitudinal or oblique. The afferent branchial channels enter either behind the pterygostomian regions and in front of the chelipeds, or at the antero-lateral angles of the palate. Verges of the male in the sternum or the coxe. Family CALAPPIDAL® Afferent branchial channels opening behind the pterygostomian regions and in front of the chelipeds. The palp of the external maxillipeds is not entirely concealed by the merus. Verges of the male on the coxa of the last pair of legs. Genus Platymera Milne-Hdwards. Carapace convex, transversely elliptical, with a strong spine at the lat- eral angles; antero-lateral margins arcuated and dentate; postero-lateral margins not produced into lateral expansions. Front narrow. Orbits oval, deep, of moderate size, and furnished witha prominent fissure in the lower margin. The basal antennal joint is situated in the wide inner orbital hiatus and does not reach the front; flagellum small. Buccal cavity comparatively in front and more or less completely divided by a median ridge. The external maxillipeds do not completely cover the anterior portion of the buccal area; merus subequal to the ischium and excavated at the antero-internal angle; palp situated at the antero-internal angle of the merus and not completely covered over; the epipodite forms a broad, semilunar plate which covers the opening of the afferent branchial channels. Chelipeds large and, when folded, fitting closely to the body; merus trigonal; hands iarge, compressed, distally widened, surmounted by a laminate and dentate crest, and furnished with a longitudinal, granu- lated ridge near the lower side of the outer surface. Ambulatory legs slender, compressed, and furnished with long, styliform dactyls; first three pairs subequal in length, the fourth the shortest. Sternal plastron oval. The abdomen in the male is composed of five distinct joints, the third of which is furnished with a prominent, transverse posterior crest. Type.—P. Gaudichaudii MILNE-EDWARDS. : | oO SP Bt mp. ef CMm—ne—e, a tes - es. Se «~—--““~e* «a «4 CALIFORNIA STALK-EYED CRUSTACEA. 99 Platymera Gaudichaudii Milne-Edwards. Platymera CGaudichaudii MitNr-Epwarps, Hist. Nat. Crust., T. II, 1837, p. 108. Epwarps and Lucas, D’Orbigny’s Voy. ’Amer. Mérid. VI, Part I, 1843, p. 28, Pl. XIII, fig. 1. Faxon, Mem. Mus. Comp. Zool. Harvard Coll., Vol. XVIII, 1895, p. 32.. Ratusun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 610. Platymera californiensis RatuBpun M., Proc. U.S. Nat. Mus., Vol. XVI, 1893, p. 253. Carapace strongly convex, evenly granulated and furnished with several small, depressed tubercles. The anterior margin of the front is truncated and strongly concave when seen fromin front. Postorbital tooth very small. Antero-lateral margin regularly arcuated and armed with about fifteen small teeth separated by concave interspaces. Lateral spine large, straight, subcylindrical. Maxillipeds granulated; ischium dentated on the inner margin; merus very deeply excavated at the antero-internal angle. Merus of the chelipeds armed witha spine at the infero-distal angle, above which is a small tooth; carpus with some small tubercles on the outer surface; hands also furnished with several small tubercles on the outer surface; the ridge on the lower portion of the outer surface is very large and bears a small tooth near the proximal end; the superior crest is more or less hairy and armed with six teeth; a granulated ridge on the lower side of the hand; pollex short, subtriangular, flattened, depressed; the outer margin of the finger is prominent, granulated, and when closed, is at right angles to the palm. Upper margins of the ambuiatory legs granulated, either the first or the second pair may be the longer. Length of carapace, 41 mm.; breadth between tips of lateral spines, 79 mm.; length of lateral spine, 12 mm. Described from a specimen received from Miss Rathbun. Shit, Panama to.lats37. 0’ 30%.N., lon: 122° 33°:307 W. Depth, 29 to 204 fathoms. . Family LEUCOSIIDA. Afferent branchial channels opening at the antero-lateral angles of the palate. Palp of the external maxillipeds entirely concealed beneath the merus. Verges of the male on the sternum. ; . Genus Randallia Stimpson. Carapace subhemispherical; evenly rounded at the sides, with two lobes or teeth on the posterior margin. Front narrow, very short, with a con- cave anterior margin. Eyes small, and located in small, circular orbits 100 CALIFORNIA ACADEMY OF SCIENCES. which are provided with two or three marginal fissures. Antennules obliquely plicated, the laminiform basal joint capable of closing over and. concealing the following joints. Antenne minute, with the basal joint comparatively large. Merus of the maxillipeds subtriangular, shorter than the ischium, reaching nearly as far forward as the front; exognath broad and nearly as long as the endognath. Chelipeds rather long; merus sub- cylindrical; hand narrow; fingers acute and somewhat compressed. Am- bulatory legs of moderate length, the joints not dilated; dactyls styliform. The base of the abdomen in the male covers the last thoracic sternum; distal portion narrowed. Beles 7 Type.—h. ornata (RANDALL). Randallia ornata (Randall). Ilia ornata RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 129. Guiaia ornata GIBBES, Proc. Am. Ass. Adv. Sci., 1850, p. 186. Randallia ornata Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 85;. Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 471, Pl. XIX, fig. 3; Aun. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 69. Karusun M., Proc. U.S. Nat. Mus., Vol. XXI, 1898, p. 613. Carapace nearly smooth, but furnished anterioriy with a few scattered granules and a few larger granulations at the sides. A granulated intum- escence on the hepatic area and a blunt, granulated projection on the pterygostomian regions. A small tubercle on the posterior portion of the branchial region. Posterior margin of the carapace elevated, granulated, and furnished with two pointed tubercles. Maxillipeds granulated, the exognath considerably narrower than the endognath. Merus of the cheli- peds subcylindrical, pustulate; carpus granulated; hand long, palm inflated, and granulated on the upper margin; fingers slender, sulcate, generally longer than the palm, and finely dentate. Ambulatory legs sub- cylindrical; propodi with acute, granulated upper margins; dactyls longer than the propodi; those of the first pairs compressed; those of the last depressed. Abdomen of the male narrowly triangular, acute, tive-jointed, the third joint longer than the two following ones. Penultimate abdominal segment in the female large, smooth, convex, covering nearly the entire sternum; last joint small, oblong, distally rounded. The sternal surface in the female is deeply excavated and bounded bya prominent rim, against which the large segment of the abdomen closes. The inner side of this segment is hollowed out, thus forming a capacious chamber for holding the ova. At the anterior margin of the rim is a deeply concave notch which receives the small, terminal segment of the abdomen. The carapace is of a light color marbled with reddish patches which are larger in front. Length and breadth of a specimen each 48 mm. } : : ‘ 4 ; abc yt eee Se) a moe ep ier @ ‘ey ~~ CALIFORNIA STALK-EYED CRUSTACEA. LO Mendocino County, Calif.! Santa Barbara! Santa Catalina Island! San Diego! Magdalena Bay and off Abreojos Point, Lower California (Rathbun). Randallia bulligera Rathbun. Randallia bulligera RATHBUN M., Proc. U. S. Nat. Mus., Vol. XXI, 1898, p. 614, Pl. XLIV, fig. 6. A small species. Carapace a little longer than wide and covered with distinct, smooth, bead-like tubercles; a sulcus behind the tumid hepatic regions, and in front and at the sides of the intestinal area; posterior margin of the carapace with two, pointed, granulated tubercles; a granu- lated tubercle on the posterior end of the branchial area, and a single. tubercle on theintestinal region; frontal margin concave. Pterygostomian regions with a longitudinal, tuberculated prominence. Antero-lateral angles of the buccal area with three prominent lobes which project in front of the maxillipeds. Sternum and abdomen tuberculated. “Maxilli- peds with prominent tubercles. Merus of the chelipeds cylindrical, tuber- culated like the carapace; carpus and hand granulated. Ambulatory legs granulated; dactyls slender, longer than the propodi. Coloration much as in ornata. Magdalena Bay, Lower California (Miss Rathbun); off San Diego, 30 fathoms! Easily distinguished from ornata by the numerous bead-like tubercles on the carapace and the lobes at the anterior end of the buccal area. Tribe MACROURA. Body generally elongated, though sometimes broad. Abdomen gener- ally large and exceeding the carapace in length, the sixth segment usually bearing well developed pleopods which, with the telson, form a strong tail-fin. Eye-peduncles not lodged in weil defined orbits. Antennules generally elongated and not lodged in fossettes. Antenne usually furnished with a scale (exopod) attached to the second joint. The external maxilli- peds are typically pediform. Anyor none of the pereopods may be chelate. The vulvz of the female are in the coxa of the third pair of legs. 192 CALIFORNIA ACADEMY OF SCIENCES. KEY TO THE SUBDIVISIONS OF THE MACROURA. Abdomen agpmmetrical ! 5, sak haemo es coc Geer.) ae See Paguridea. Abdomen symmetrical. Exoskeleton calcareous; pereopods mostly six-jointed. None of the three anterior pairs of pereopods chelate.. ..Loricata. All of the three anterior pairs of pereopods chelate... Homaridea. First pair, or first two pairs of pereopods chelate; third pair never chelate. Last pair of pereopods reduced, slender, folded. Thoracic, sternum-bread 222 3.552) ene eee Galatheidea. Phoraeie sternum, Jinearay, \se5 48 sane eee Hippidea. Last pair of pereopods not greatly reduced..... Thalassinidea. Exoskeleton generally corneous; pereopods seven-jointed. Antennal scale large. Swimming forms with generally laterally compressed bodies. Third painvot pereopods chelate +) 2b ue seen eee Peneidea. Third pamotspereopods not chelate.) 22a.0.a.0 «see Caridea. Subtribe HIPPIDEA. Carapace ovate or subquadrate, comparatively smooth, the regions not well defined. Interantennulary region broad. Eyes small. Antennules generally well developed, with one flagellum elongate, the other short or absent. Antenne with a large five-jointed peduncle, with or without an acicle. Maxillipeds more or less operculiform. Dactyls of the ambula- tory legs flattened; fifth pair of legs slender and filiform. Thoracic sternum linear. Abdomen partially extended; telson large; penultimate abdominal joint with a pair of biramous lamellate appendages, which, however, do not form a caudal tin in connection with the telson. The males have no abdominal appendages, except the last pair. Pterygosto- mian regions free from the epistome. Family HIPPIDA. Antennal acicle small or absent. Maxillipeds operculiform, the merus broad; exognath wanting. First pair of legs not subchelate. Telson elongated, lanceolate. - Genus Hippa Fabr. Antennules of moderate length. Antennal flagellum very long, stout, multiarticulate and strongly ciliated. Maxillipeds with the ischium very small and the merus very large; the terminal joint narrow and compressed. Dactyls of the first pair of legs oval and lamellate. Type.—H. emerita L. _ . . | 4 CALIFORNIA STALK-EYED CRUSTACEA. 103 Hippa analoga St. Hippa emerita De Saussure, Rev. et Mag. de Zool., T. V, 1853, p. 367. ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. IX, 1896, p. 232. Hippa talpoidea DANA., Proc. Acad. Nat. Sci. Phila., 1854, p. 175. Hippa analoga Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 85; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 486. Murrs, Journ. fina. soc... London, (Zool.) Vol. XLV, 1879,.p., 324, Pl. .V, fig. 10. ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. VI, 1892, p. 537, Pl. XXVI, he. 2. Carapace oblong-oval, very convex, and marked with irregular, trans- verse, crenulated lines which become much less marked towards the sides and posterior end. An impressed, nearly straight, transverse line behind the front, and an impressed line which is concave in front on the median region. Median lobe of the front subtrianguiar but rounded at the tip; the lateral lobes are triangular and acute and project farther forward than the median lobe from which they are separated by round, smooth sinuses. Antero-lateral margins serrated. Eye-stalks long and very slender. Sec- ond joint of the antennal peduncle furnished with three spines of which the median is the largest and upturned at the tip; lowest spine smallest; superior spine more or less upturned; flagellum very long, curved and bent back under the body. Merus of the maxillipeds with the lobe at the antero-internal angle rounded. First pair of thoracic legs with a strong spine at the supero-distal angle of the carpus and at the infero-distal, angle of the propodus; dactyl ovate. Dactyls of the second and third pairs falcate, very broad at the base, and subacute at the tip. Telson narrowly triangular, acute; outer surface convex, smooth, glossy; margins raised and furnished with two rows cf hairs, the inner row lying on the surface. Length of carapace, 29 mm.; width, 23 mm.; length of telson, 19 mm.; width of telson,.9.5 mm.; length of abdomen when extended, 37 mm. The ratio of the length of the carapace to its breadth is quite variable. Oregon to Panama on sandy beaches. Family ALBUNEID. Anterior legs more or less perfectly chelate. The maxillipeds are sub- pediform or oniy moderately expanded, and furnished with an exognath. Telson not elongated, generally ovate and lamellate. Genus Blepharipoda Randall. Eye-peduncles very slender, elongated, cylindrical, and articulated in the middle. Antennules and antennez rather long, with multiarticulate flagella. Antenne devoid of an acicle. Merus of the maxillipeds narrow, similar to the carpus; carpus not produced at its antero-external angle. Type.—B. occidentalis RANDALL. 104 CALIFORNIA ACADEMY OF SCIENCES. Blepharipoda occidentalis Randall. Blepharipoda occidentalis RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 131, Pl. VI. Guispes, Proc. Am. Ass. Adv. Sci., 1850, p..187. Dawa, Crust. U.S. Expl. Expd., Part I, 1852, p. 406. Srimpe- gon, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 486; Proc. Acad. Nat. Sci. Phila., 1858, p. 230. Muers, Journ. Linn. Soc. London, Vol. XIV, 1879, p. 334. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. TX, 1896; p. 222: Carapace oblong, scabrous in front, but smooth and punctate behind. The middle of the carapace is elevated, forming a longitudinal ridge which ends anteriorly in a spine immediately behind the transverse postfrontal impression. A transverse impression behind the median region and an oblique one on either side terminating at the last lateral spine. Front with a median spine separated by a rounded sinus from the more promi- nent triangular spine-tipped, lateral teeth. Antero-lateral margin with three large spines and a smaller fourth spine some distance further back. The last joint of the eye-peduncles is but little longer than the preceding one. Antennules with the upper flagellum long (over one-haif the length of the carapace) and strongly ciliated; lower flagellum scarcely as long as the peduncle. Peduncle of the antennz long (nearly one-half the length of the carapace), the last two joints cylindrical and ciliated on the outer surface; flagellum curled, strongly ciliated on one side, and somewhat — shorter than the peduncle. Ischium of the maxillipeds dentate on the inner margin and: produced at the antero-internal angle; exognath reach- ing considerably beyond the middie of the merus. Chelipeds strong; merus short, stout, with one or more spines on the lower side; carpus scabrous externally, with the supero-distal angle produced into a large, compressed, triangular, spine-tipped tooth; at the base of this tooth the upper margin is armed with a variable number of spines, and there may be several small spines on the anterior edge; hand flattened, the outer surface scabrous and furnished with two spines, one near the base, the other behind the gap between the fingers; a spine near the middle of the lower margin, in front of which there may be several smaller ones; pollex broad, triangular, much compressed, with the inner margin armed with a variable number of spines; dactyl rather slender, curved, and spiny on the outer edge. The ambulatory legs have the anterior margins of the carpi spinulous and the antero-distal angles produced; dactyls subfalci- form, but varying greatly in shape in the different pairs. The slender last: pair of legs ends in a small, well developed chela. The lateral expansions of the fifth abdominal segment are subacute. The inner ramus of the uropods is narrower than the outer and articulated near the middle of the side of the peduncle, the outer one joined to thetip. Telson short, CALIFORNIA STALK-EYED CRUSTACEA. 105 suborbicular, thick and convex in the middle, but with the sides laminate. Length of carapace, 46.5 mm.; breadth of carapace, 35 mm. Length of carapace, 47 mm.; breadth of carapace, 36 mm. San Diego (Randall)! Monterey (Miers); Santa Monica, Calif.! Estero Bay! San. Quentin Bay, Lower California! Genus Lepidopa Stimpson. Eye-peduncles lamellate, compressed, almost squamiform; cornes rudi- mentary. Antennules long. Antennz with a very small accessory joint; flagellum very short. Maxillipeds with the fourth joint produced at the antero-external angle into a lobe which reaches to or beyond the distal extremity of the fifth (penultimate) joint. Type.—L. scutellata (FABR.) Lepidopa myops St. Lepidops myops Stimpson, Ann. N.Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 241. Mrers, Journ. Linn. Soe. (Zool.) London, Vol. XIV, 1879, p. 333, Pl XIV, fig. 16: Lepidopa myops ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. IX, 1896, p. 226. Carapace oblong, with an obtuse median ridge, and marked with trans- verse grooves. Front trilobed, the median lobe rounded and shorter than the lateral lobes, which are acute. A spine at the antero-lateral angles. Posterior margin of the carapace deepiy concave in the center. Ocular peduncles oblong, broadly rounded in front, the cornea when present minute and located on the margin near the antero-lateral angle. Anten- nules ciliated and over twice the length of the carapace. Antenne shorter than the carapace; a thin lamina on the inner side of the basal joint; acicle minute; flaelglum scarcely as long as the peduncle. Chelipeds stout; hand high, and strongly compressed; pollex very short and curved; finger rather slender, curved, and, when closed, lying nearly at right angles to the long axis of the hand. The dactyls are compressed, falcate, dissimilar. The second, third, and fourth abdominal segments are furnished with wide, wing-like expansions which diminish in length and breadth pos- teriorly. Telson about as wide as long and rounded at the sides and apex. Length of carapace, 15 mm.; breadth, 17 mm. Cape St. Lucas (Stimpson); Lower California! San Diego (five specimens)! The small teeth on the median frontal lobe mentioned by Stimpson are present on only a part of the speci- mens I have seen. 106 CALIFORNIA ACADEMY OF SCIENCES. Subtribe GALATHEIDEA. Rostrum present. Body flattened, the carapace divided by a lateral (anomoural) line. Abdomen flattened, the sides of the segments generally laterally produced. ‘Tail-fin well developed. Pterygostomian regions free from the epistome. External antennz four-jointed, the second and third joints fused; acicle small or absent. First pair of pereopods chelate; the last pair chelate, rudimentary, and folded in the branchial chamber. Legion PORCELLANINEA. Carapace broad, smooth. Antennules concealed. Antennz long and generally furnished with a scale. Ischium of the maxillipeds broad and flattened; merus broad, with a prominent inner lobe. Chelipeds large, often flattened; first three pairs of ambulatory legs well developed; last pair slender, inflexed. Abdomen loosely bent under the trunk. These crabs in the form of the carapace, the bend of the abdomen, and in their flattened maxillipeds resem- ble the true Brachyura; but these resemblances are not indicative of true affinity for both groups had, in all probability, an independent origin from Macrouran ancestors. Genus Petrolisthes St. Carapace subovate, depressed, generally longer than wide. Front trian- gular, entire or dentate, usually depressed, the margins more or less undulated. Eyes of rather large size. First basal joint of the antennz very short, not reaching the upper margin of the carapace; the second joint is flattened and more or less cristate; flagellum long. Chelipeds broad and flattened; the carpus elongated, with or without teeth on the inner margin. The dactyls of the ambulatory legs are short and stout and terminate in a single claw. Type.—P. violaceus (GUERIN). ——— = CALIFORNIA STALK-EYED CRUSTACEA. 107 Petrolisthes cinctipes (Randall). Porcellana cinctipes RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 136. | Porcellana rupicola Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 85; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 480, Pl. XIX, fig.2. Bate in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 276. Petrolisthes rupicolus Stimpson, Proc. Acad. Nat. Sci. Phila., 1858, p. 227. Locki1neTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 396. Nrwcomsg, Bull. Nat. Hist. Brit. Col., 1893, p, 30. Petrolisthes cinctipes ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. X, 1897, p. 278. Carapace longer than wide, and quite distinctly areolated; the anterior portion is roughened by minute prominences and the branchial regions are transversely finely striated. Front triangular, depressed, blunt, trans- versely concave above, with a rounded notch on either side of the base next to the orbit. Antero-lateral margin marked by araised line. Ocular peduncles somewhat flattened; superior margin of the orbit slightly con- cave. Antennal peduncle granulated; the flagellum may exceed one and one-half times the length of the carapace, and is furnished with a few sete which are longer than the width of the joints. Ischium of the max- illipeds finely rugose, the distal margin transverse and broadly rounded; the outer border is prolonged into a lobe; the lobe on the inner side of the merus is prominent and distally rounded. Chelipeds granulated above; a small tooth at the antero-distal angle of the merus; carpus slightly tapering towards the distal end; no teeth on the anterior margin but the inner angle is prominent and evenly rounded; the posterior surface of the carpus is rugose, the upper edges of the projecting ruge forming a rough ridge along the posterior margin which terminates distally in a tooth; hands strongly flattened, outer edge acute, nearly straight along the mid- dle; tips of the fingers curved. Ambulatory legs more or less rugose, the merus expanded and nude; propodi hairy and furnished with a few short spinules near the distal end of the posterior margin; dactyls short, hairy, curved, acute, and generally furnished with a few minute spines below. The posterior pair of sutures in the telson is oblique. Length, 20.5 mm; breadth, 19.75 mm. Vancouver’s Island (Bate, Newcombe); WWumboldt County, Calif.! San Francisco Bay! Monterey! Santa Barbara! Santa Catalina Island! San Diego! San, Miguel Island; west coast of Lower California and Gulf of Cali- fornia (Lockington). 108 CALIFORNIA ACADEMY OF SCIENCES. This is one of the most common of our California crabs and is generally found under rocks at low tide. According to Ortmann (l.c.) ‘‘Die in Philadelphia aufbewahrten Originale Randall’s zeigen, dass cinctipes dieselbe Art ist wie rupicola. Der von Randall ange- gebene fundort ‘Sandwich-Ins.’ beruht offenbar auf der unter seinem Material vorgekommenen Verwechs- lung der Localitaten.”” I am unable to follow Ortmann in uniting eriomerus St. with this species. The two. forms occur together on a great part of the California coast and keep perfectly distinct. Petrolisthes eriomerus ‘S?. eee ae Petrolisthes eriomerus Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. X, p. 119. Locxineton, Ann. Nat. Hist. (5), Vol. II, 1878, p. 397. Closely allied to cinctipes. Carapace smooth and punctate, though there are often minute prominences, especially anteriorly, which are not so marked as in the preceding species. Front not so strongly deflexed as in cinctipes. The antennz may be over twice the length of the carapace and are devoid of sets. Distal extremity of the ischium of the maxillipeds more or less truncated. Chelipeds longer and smoother than in cinctipes; the carpus is narrower and has the sides parallel; the antero-internal angle is much less prominent and the ridge on the posterior margin rougher. The merus joints of the ambulatory legs are hairy and less dilated than in the preceding species. The last pair of sutures on the telson is trans- verse or nearly so. Humboldt County, Calif., to San Francisco Bay! Point Mendocino (Stimpson)! I have collected numer- ous species at the latter, or type locality. Cinctopes is also abundant in the same place. | Genus Pachycheles Stimpson. Carapace round-ovate, or suborbicular, and not longer than wide, the posterior portion of the subbranchial region subquadrate and separated from the larger anterior portion by a membranous interval. Front but little produced, subacute, edentulous. First joint of the antennal peduncle ee —— = CALIFORNIA STALK-EYED CRUSTACEA. 109 produced and joined to the margin of the carapace; second joint at some distance from the orbit. Chelipeds thick and roughened; carpus short. Dactyls short and terminating in a single claw. Type.—P. grossimanus (GUERIN). Pachycheles rudis St. Pachycheles rudis Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1862, p. 76; Proc. Acad. Nat. Sci. Phila., 1858, p: 228. LockinerTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 404. Nerwcomss, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 30. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. X, } 1897, p. 294. Carapace nearly smooth, quite strongly convex longitudinally, and finely striated on the branchial regions. Front short, deflexed, entire, and hairy above. Sides of the carapace evenly rounded and marked by a raised line. Sinus in the posterior margin deep and subacute in the middle. Superior margin of the orbit concave but not raised; outer angle of the orbit acute. Distal end of the ischium of the maxillipeds evenly rounded; inner lobe of the merus prominent, rounded. Chelipeds unequal, with a rough, irregular, granulated, or tuberculated upper surface, which is more or less hairy; merus with an anterior tooth; carpus short, about as broad as long, the anterior margin laminate, angular, and often furnished with one or more teeth; the posterior margin is convex and the upper surface bears two more or less evident longitudinal, granulated ridges; hands broad, subtriangular, very uneven above, and furnished with a rounded protuberance near the middle; the lower surface is glossy and very finely granuiated at the center, but the granules become larger towards the mar- gins; fingers stout, granulated. Ambulatory legs stout and more or less rugose; dactyls short, curved, and furnished with two or three spines below; posterior margins of the propodi with a few spines. Length, 14.5 mm.; breadth, 15.25 mm. British Columbia (Newcombe); Humboldt County, Calif., to San Francisco Bay! Monterey! Santa Rosa Is.! Santa Catalina Is.! Found under rocks at low tide. Lockington states that he found young specimens of this species in a bottle of material, without label, from Lower California. 10) CALIFORNIA ACADEMY OF SCIENCES. Pachycheles pubescens, sp. nov. Carapace suborbiculate, not so convex longitudinally as in rudis, and more or less distinctly areolated in front, where it is marked by small, transverse punctures. Branchial regions transversely striated. Sinus of the posterior margin of the carapace subacute and not nearly so deep as in the preceding species. Front entire moderately deflexed, somewhat: pro- duced and rounded in the center. The membranous interval separating the parts of the subbranchial region is nearly vertical. Eyes large. An- tennal peduncle compressed. Superior orbital margin concave; postorbital tooth acute. The ischium of the maxillipeds is finely transversely rugose and the distal margin is more or less truncated. Upper surface of the chelipeds not rugose or tuberculated, but coarsely granulated and setose; eranulations large, smooth, shining, generally transverse, those on the — hand larger, becoming more acute towards the outer side, where they give rise toa denticulated margin; a tooth on the anterior margin of the merus; carpus generally a little wider than long, the anterior margin laminate and cut into three or four teeth armed with secondary denticles, which are most numerous on the proximal tooth; hand similar in shape to that of rudis; fingers stout, the inner margins an Legs stout, setose. Pro- podi and dactyls spinulous below. Length of carapace, 15 mm.; width, 15 mm.; length of carpus, 11.5 mm.; width, 11.5 mm.; length of hand, 2 miss widths 13 mm. Drake’s Bay, Calif.! Farallon Islands! Humboldt County, Calif.! This species differs from Pachycheles setimanus (Lock- ington), the types of which I have examined, in its somewhat narrower carapace, more prominent front and broader pollex. Collection of the University of California. Legion GALATHEINEA. Carapace elongated, generally rugose, and quite distinctly areolated; ros- trum prominent, acute. Ocular peduncles short and stout, and lodged in incomplete orbits. Antennules not lodged in fossettes. Antennz with the second and third joints often coalesced; flagellum long. Maxillipeds subpediform, the ischium and merus narrow, Chelipeds often slender and elongated. Fifth ambulatory legs feeble and inflexed. Abdomen broad and bent under the body; in the female the second to fifth segments CALIFORNIA STALK-EYED CRUSTACEA. TF generally bear appendages; in the male there are often sexual appendages on the first two segments, and the appendages on the three following segments may be well developed or rudimentary; the last pair of append- ages is always well developed and forms, with the telson, a strong tail-fin. Genus Munida Leach. Carapace rugose, generally spinous, the cardiac area distinctly defined; the infero-lateral regions are not swollen out and are separated from the dorsal surface by a well defined margin. Rostrum slender and styliform, with a supraorbital spine on either side of the base. Ocular peduncles free; eyes normal. Chelipeds and ambulatory legs elongated and slender. One or more of the abdominal segments usually furnished with a series of spinules on the anterior dorsal margin. Type.—M. rugosa (FABR.). ‘This genus is represented off the coast of California by a species Munida (Grimothea) gregaria, the Galathea gregaria of Fabricius, which still appears to be of uncer- tain systematic position. Miers considers it the young of Munida subrugosa, while Henderson thinks it doubtful that such is the case." Owen? reports this species from “off San Francisco.” Genus Pleuroncodes St. Near Munida. Carapace rugose, with the infero-lateral margins swollen out so that the sutures (anomoural iines) are visible from above. Rostrum styliform, with a supraorbital tooth on either side of the base. Ocular peduncles free; eyes large, normal. Insertion of the antennez not con- cealed beneath the antero-lateral angles of the carapace. Merus of the maxillipeds unarmed, the penultimate joint slightly dilated. Type.—P. monodon (MILNE-EDWARDS). 1 See Miers’ Catalogue of New Zealand Crustacea, p. 168 ; Henderson, Challenger Reports Vol. XXVII, p. 124; and Milne-Edwards and Bouvier, Ann. Sci. Nat. (7), T. XVI, 1894 pp. 256 and 314. 2 Zoology of Beechy’s Voyage, Crustacea, 1839, p. 87. 14 CALIFORNIA ACADEMY OF SCIENCES. Pleuroncodes planipes Si. Pleuroncodes planipes Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 245. A. Mi~ne-Epwarps, and Bouvier, Ann. Sci. Nat. (7), T. XVI, 1894, pp. 248 and 245. Closely allied to P. monodon from the coast of Chili. Carapace crossed by setose striz but, with the exception of a few spinules behind the supra- orbital teeth, devoid of spines. Rostrum long, very slender, scabrous above, and continued back upon the carapace as a carina; supraorbital teeth spine-like. A spine at the rounded antero-lateral angle of the cara- pace, behind which there area few spines on the margin. Ocular ped- uncles not reaching the tip of the rostrum; eyes large. Mavxillipeds with a small spine at the antero-internal angle of the ischium; penultimate joint moderately dilated. Chelipeds long, slender, and spinous; merus slender, trigonous, spinulous, especially on the edges, and generally ex- ceeding the tip of the rostrum; carpus spinous; hand long, narrow, spinulous, with the upper and lower margins parallel; fingers slender, straight, longer than the palm, the tips curved. Ambulatory legs sca- brous, ciliated, with the penultimate joints flattened. Abdomen devoid of ‘spines. ‘‘This species,” says Stimpson, ‘‘lives in the open ocean, and is sometimes found in vast quantities in the Pacific off the American Coast. It was taken by Mr. Grayson in N. lat. 24°, W. lon. 130°.; In March? iao: it was thrown ashore in considerable numbers at Mon- terey, California.” Ninety miles southwest of San Francisco, Calif.! 150 miles southwest of Cape St. Lucas, Lower California, numerous specimens! Subtribe PAGURIDEA. Carapace with a lateral (anomoural) line. Pterygostomian regions free from the epistome. External antenne with a five-jointed peduncle and, generally, with an acute, movable acicle. First pair of pereopods large, chelate, often unequal; last pair small and generally more or less chelate. Abdomen asymmetrical, the lower and often the upper side membranous; the appendages generally reduced or absent, and commonly occurring only on one side. , al 6 ea lr ce CALIFORNIA STALK-EYED CRUSTACEA. 113 Legion LITH ODINEA. Carapace broad, often resembling that of the Brachyura. Rostrum generally well developed. Third ambulatory legs subequal to the preced- ing pair; last pair feeble, chelate, and folded in the branchial chamber. Sternum wide. Abdomen bent under the thorax; in the male it is devoid of appendages, but in the female the first segment mary bear a small pair, and each of the four following segments often has an appendage only on the left side. KrY TO THE GENERA OF Lithodinea. Carapace smooth and produced laterally into two wings, which Bumeea) tho-amDUlalory LOSS .6c cui. .s0 hs pacie eo dww enw an Cryptolithodes. Carapace not so produced. Carapace devoid of spines or tubercles on the upper surface. Carapace much flattened, hairy...... ss ek IRS rn ee Hapalogaster. Carapace not much flattened; acicle curved......... Gdignathus. Carapace not much flattened; acicle straight........ Dermaturus. ' Carapace with spines or tubercles above. Carapace with deep pits above.............000 .0e Phyllolithodes. Carapace not deeply pitted. : Abdomen membranous. Carapace spiny.... Acantholithodes. Abdomen furnished with calcareous plates. Carapace tuberculated ........0 00)5...006 Lopholithodes. RRA SRDEACEE PRT 0 ofa. Sh0 EE oo oe ete ths Soh tists Avera ius Lithodes. Genus Hapalogaster Brandt. Carapace flattened and marked with chitinous, uncalcified lines; lateral margin with teeth or spines. Last joints of the external maxillipeds dilated. Chelipeds depressed, very unequal; hands more or less trigonal, moving horizontally. Ambulatory legs depressed. Abdomen soft, loosely inflexed, the basal segment with a transverse, calcareous plate on either side. Type.—H. Mertensii BRANDT. Hapalogaster cavicauda St. Hapalogaster cavicauda Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 81, Pl. I, fig. 7; Proc. Acad. Nat. Sci. Phila., 1858, p. 232. ScHALFEEW, Mélan. biol., Tome XIII, p. 326, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, p. 332. Boas, Vid. Selsk. Skr., 6 Rekke, Natur. og Math., Afd. 1, 2, 1894, p. 194, Pi. VI, fig. 200. Bouvisr, Ann. Sci. Nat. (7), Tome XVIII, 1894, p. 166, Pl. XI, fig. 2, and Pl. XII, figs. 1, 15, 27 and 29; Ibid. (8), Tome I, 1896, p. 2, et seq. 8 114 CALIFORNIA ACADEMY OF SCIENCES. Body and legs much flattened and densely covered with short hair. Carapace subcordate, nearly smooth, but roughened or granulated at the insertions of the hairs. Front short, acute, narrowly triangular. Post- orbital tooth large, acute, and extending nearly as far forward as the*ront. Superior orbital margin concave and furnished with a small, acute tooth which is separated from the postorbital by a deep fissure. The portion of the antero-lateral margin in front of the cervical groove is convex, sub- laminate, edentate, and separated from the portion behind the groove by an incision; two marginal teeth at the origin of the sutures. Ocular peduncles short, hairy, and swollen at the base, where they are furnished with a small, smooth tubercle. Antenne about as long as the carapace; acicle ovate, acute, very thin, more or less transparent, and margined by hairs on both sides; a smooth, slender spine joined beneath, and reaching nearly to the tip of the acicle. Second joint of the palp of the maxillipeds with a rounded inner lobe; last joint rounded distally and somewhat flat- tened. Chelipeds flattened, not tuberculated, anterior margin of the merus produced and cut into two teeth; carpus produced into a tooth at the pos- tero-distal angle and bearing a single spine at the proximal end of the anterior margin; larger hand with one or more small, calcareous tubercles on inner face behind the articulation of the dactyl; inner margin of the smaller hand produced into a rounded lobe behind the base of the dactyl; the fingers of the smaller hand are excavated within, the edges corneous and armed with small, calcareous denticles. Ambulatory legs much flat- tened and very hairy, the anterior margins deeply incised, forming four or five closely approximated teeth on each of the larger joints; postero-distal angle of the carpi produced into a tooth or lobe which is very prominent on the third pair; dactyls short, flattened, the curved, corneous tips turned obliquely to the plane of compression. The abdomen is broad and bent abruptly at about the third segment, the reflexed portion not so hairy as the upper side; the calcareous plates on the basal segment are wide and separated by a membranous interval in which there is no median plate. In the female the left side of the abdomen is coriaceous and seg- mentally incised. Length of carapace from tip of rostrum, 17 mm; breadth, 19 mm.; length of large hand, 28 mm.; length of small hand, 20 mm. Northern to southern California! Monterey (Stimp- son)! Found under rocks at low tide. Common. : CALIFORNIA STALK-EYED CRUSTACEA. 115 Hapalogaster Mertensii Brand. eS ' Hapalogaster Mertensii BRANDT, Mélanges biologiques, Tome I, 1850, p. 58. Srimpson, Journ. Bost. Soc. Nat. Hist, Vol. VI, 1857, p. 480. ScHALFEEW, Mélang. biol., Tome XIII, 1892, p. 327, figs. 4 and 5a, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, 1892, p. 333. Newcomesz, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 28. Bouvisr, Ann. Sci. Nat. (8), Tome I, 1896, p. 4 et seq. Carapace with scattered tufts of seta on the upper surface. Rostrum acuminate, longer than in cavicauda. A large, sharp spine at the outer orbital angle, and a smaller sharp spine on the superior orbital margin, separated from the last by an incision. No marginal spines in front of the cervical groove, but behind it there are five slender spines which decrease in size posteriorly. Merus of the large cheliped armed in front with a pair of large spines having a small spine on either side; carpus armed with several spines subserially arranged; hand narrow, with three rows of large spines on the outer surface and a row of smaller spines on the upper edge; in the smaller hand there are four spines on the outer border, three on the inner edge, a row of three spines on the upper surface near the inner side, and a single spine between this row and the outer edge near the gap between the fingers. Ambulatory legs armed with sharp spines on the anterior margin. Described from five specimens in the National Museum from Kadiak, Alaska (No. 19471), collected by W. J. Fisher. Sitka (Brandt); Kadiak (Schalfeew)! British Columbia (Bouvier); near Victoria (Newcombe). SE a ee: lr Hapalogaster Grebnitzkii Schalfeew. Hapalogaster Grebnitzkii SCHALFEEW, Mélang. biol., Tome XIII, 1892, p. 329, figs. 3a and 3b, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, 1892, p. 335. Bouvier, Ann. Sci. nat. (8), Tome I, 1896, p. 4 et seq. Closely allied to Mertensii. Carapace devoid of prominent tufts of sete. Rostrum prominent, acuminate; a large spine at the outer orbital angle and a sharp spine on the superior orbital margin which is relatively larger than the corresponding spine in the preceding species. Margin of the carapace behind the cervical groove armed with five spines, which decrease in size posteriorly. Legs armed with setose spines. The merus of the large cheliped is similar to that of Mertensii, but the carpus has ee ee = a eS -e \ 116 CALIFORNIA ACADEMY OF SCIENCES. fewer spines above; hand shaped as in Mertensii, but having only two rows of spines on the outer surface and a row of small spines on the inner edge; there is a broad, smooth interval on the outer surface, which in Mertensii is armed with the median row of spines. The number and arrangement of the spines on the smaller hand is the same as in the pre- - ceding species, with the exception that there is no spine behind the gap between the fingers. ' Described from numerous specimens in the National Museum from Humboldt Bay (No. 19443, Albatross col- lection); Behring Is., Kadiak Is. (Schalfeew). Genus Dermaturus Brandt. Carapace more or less lyrate, devoid of prominent spines or tubercles and marked with transverse strie; lateral margins unarmed. Rostrum triangular, simple. A small tooth on the margin of the orbit within the outer orbital angle. First and second joints of the antenne with a spine at the outer angle; acicle oblong, straight; last two joints of the peduncle subcylindrical. Last two joints of the maxillipeds not dilated. Chelipeds very unequal, not flattened; hands rounded, fingers excavated within. Ambulatory legs subcylindrical; dactyls spiny below and armed with long, sharp claws. Abdomen soft, loosely inflexed. Type.—D. Mandtii BRANDT. a cu oe Brandt. PL. £.146. Dermaturus Mandtii BRANDT, "Bull, phys..math. Acad. imp. sci. St. Petersb., Tome VIII, 1850, p. 50; Mélanges biologiques, Tome JI, 1850, p. 58. Stimpson, Proc. Acad. Nat. Sci. Phila., 1858, p. 282. Bouvier, Ann. Sci. nat. (7), Tome XVIII, 1895, p. 173; Jbid. (8), Tome I, 1896, p. 19. Hapalogaster Mandtii SCHALFEEW, Mélang. biol., Tome XIII, 1892, p. 382, figs. 2 and 5c, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, 1892, p. 338. Carapace narrowly lyrate, not much wider behind than in front, the upper surface unarmed, but marked with prominent, transverse striz, the anterior edges of which are furnished with short, appressed setz. TRos- trum prominent triangular, having a minute knob of granules just behind the acute tip. Outer orbital angles acute, reaching about as far forward as the middle of the rostrum; a small, but prominent tooth on the trans- verse posterior margin of the orbit a short distance internal to the outer angle, Eye-peduncles with transverse, setose ridges. The spine at the CALIFORNIA STALK-EYED CRUSTACEA. a ky outer angle of the second joint of the antenna is long and narrow, reach- ing beyond the middle of the acicle; acicle narrow, oblong, straight, the tip rounded, the margins, especially the inner, setose. The last two joints of the maxillipeds are not dilated. Chelipeds with transverse granulated ridges which are furnished with a fringe of appressed setx on the distal side; merus of the large cheliped with 5-6 teeth on the anterior margin; carpus armed in front with three teeth, the proximal one being quite prominent; hand crossed by double rows of granules which become broken up into separate groups on the pollex; fingers with tufts of setx; merus of the small cheliped armed in front with several spines which decrease rapidly in size towards the distal end; anterior margin of the carpus with three teeth, the proximal one much the largest; hand narrow, anterior margin of the palm with four small teeth at the ends of lines of granules; fingers hairy, longer than the palm; a group of granules above the base of the dactyl. Ambulatory legs crossed with setose strie; claws long and sharp. Described from two specimens obtained from the National Museum. Pribyloff Is. (Brandt); Kadiak Is.; Behring Is. (Schal- feew); slat. 50°. 304°.30° N.; lon. 162° 31% 45", W.; .19 fathoms! Genus Cdignathus Benedict. Carapace lyrate, convex, devoid of spines or tubercles and furnished with a short simple rostrum. Antennal acicle flattened, the inner margin thick and concave, the outer thin and convex; last two joints of the peduncle subcylindrical; flagellum long. Maxillipeds subpediform; the last two joints dilated. Chelipeds unequal; tips of the fingers excavated within. First three pairs of ambulatory legs subequal, subcylindrical; dactyls with curved, corneous claws. Abdomen soft, thick, broad, and loosely inflexed, the basal, and to a less extent, the last two segments strengthened by calcareous plates. Type.—O. Brandti (SCHALFEEW). This genus is closely allied to Dermaturus with which it has been united by Bouvier. It differs from Derma- twrws in having a more or less crescent-shaped acicle, in the different marking of the carapace, and in not having the last two joints of the maxillipeds dilated. 118 CALIFORNIA ACADEMY OF SCIENCES. aes seal a Brandti (Schalfeew). Hapalogaster Brandti sit Ashes Mélang. biol., Tome XIII, 1892, p. 330, figs. 1 and 50, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, 1892, p. 336. dignathus Gilli BENEpDIcT, Proc. U.S. Nat. Mus., Vol. XVII, 1894, p. 487. Dermaturus Gilli Bouvier, Ann. Sci. nat. (8), Tome I, 1896, p. 19. Carapace lyrate, convex above, and covered with flat scaie-like plates which are setose on the anterior margin. Rostrum short, triangular, acute and curved downwards. Outer orbital angle subacute, the inner orbital tooth represented by a minute prominence. Upper surface of the antennal acicle furnished with minute spines; lower surface smooth. Chelipeds very unequal and covered with low, granulated, wart-like tubercles; in the larger cheliped the anterior margin of the merus is produced and cut into three or four teeth; the anterior margin of the carpus is furnished with a jagged tooth near the base; the hand is large and swollen, the fin- gers are somewhat gaping at the base, the tips excavated within. In the smaller cheliped the process on the anterior side of the merus is gener- ally cut into two small teeth; a tooth on the anterior margin of the car-. pus; the hand is narrower and less tumid than in the large cheliped; the fingers are deeply excavated within and meet along the entire inner margin; outer margin corneous like the tip and armed with a few small teeth. Ambulatory legs more or less hairy; a group of small spines on the infero- distal angle of the propodi; dactyls ending in black, curved, acute tips and arroed below with five or six black spines. Abdomen covered with a short pubescence; the margins fringed with long hairs; the calcareous plates on the last two segments inconspicuous. Four two-jointed, uni- . ramous appendages on the left side of the abdomen in the female. The margin of the left side of the abdomen in the female is hardened and seg- mentally incised, but this is not the case in the male. Length of carapace from tip of the rostrum to the posterior emargina- tion, 15.5 mm.; breadth, 15.5 mm.; length of large cheliped, 40 mm.; length of small cheliped, 28 mm. é Alaska (Benedict)! British Columbia! Humboldt _ County, Calif., to San Francisco Bay! ote Se wee ur) This species is often found on the rocks among t mussels. It may be easily recognized by its lyrate carapace covered by the very peculiar scale-like plates and the very unequal chelipeds with their wart-like tubercles. Through the kindness of Mr. Bencaict I have exam- ined the types of his Gillc (No. 18525, U.S. Nat. Mus.). CALIFORNIA STALK-EYED CRUSTACEA. 119 (dignathus inermis (St. ). Hapalogaster inermis Stimpson, Ann. N.Y. Lye. Nat. Hist., Vol. VII, 1860, p. 243. ScHALFEEW, Mélang. biol., Tome XIII, 1892, p. 326, du Bull. Acad. imp. sci. St. Petersb., Tome XXXV, 1892, p. 332. NewcomeseE, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 28... Bouvisr, Ann. Sci. Nat. (7), Tome XVIII, 1895, p. 172. _ Dermaturus inermis Bouvier, Ann. Sci. Nat. (8), Tome I, 1896, p. 19. ‘“‘ Carapace longer than broad, smooth above except where some minute transverse, setose scabrosities or minute squame are sparsely distributed, occurring most abundantly toward the sides. Margins unarmed, smooth. No sinus at the juncture of the cervical suture with the lateral margin. Rostrum conyex above, aimost carinated; apex scarcely acute. Inner orbital tooth minute, inconspicuous. Outer orbital tooth (antero-exterior angle of the carapax) far less prominent than the rostrum. Acicle rather short and broad, irregularly somewhat dentated along the exterior margin. Feet subcylindrical, almost naked, rugose above with minute and some- what rounded setose tubercles; dactylus three-fourths as long as the penult joint and armed with a long unguiculus. Plates of the first joint of the abdomen narrow. ; , ‘*Length of carapax, 0.4; breadth posteriorly, 0.38 inch.” Puget Sound (Stimpson); Queen Charlotte Is. (New- combe); Sitka (Schalfeew). — Judging from Stimpson’s description, O. Brandtz is very closely related to this species and not improbably will prove identical with it. The setose squame in the specimens of Brandti I have seen were thickly set in- stead of ‘‘sparsely distributed,” the upper surface of the carapace being almost entirely covered by them, except a small, narrow area along the median line, where they were small and scattered. Stimpson’s specimen was probably immature and the character that separates Brandti may be due toage. In all the characters except the abundance and distribution of the squame Brandti agrees with Stimpson’s description. Schalfeew consid- ered his Brandti closely allied to, if not identical with unermis, and Bouvier, who had a specimen of the latter 120 CALIFORNIA ACADEMY OF SCIENCES. at hand, states that it agrees perfectly with the descrip- tion and figures of Brandti. The specimen Bouvier called znermis, however, may not have been that species, for if it agreed perfectly with Schalfeew’s description and figures of Brandtz, it could not agree perfectly with Stimpson’s description of iermis. The figure of Brandti shows the squame on the carapace to be thickly set. Genus Acantholithodes Holmes. rd 7 ‘4 Carapace lyrate, flattened, and covered with setose spines. Rostrum prominent and terminated by strong spines. The first basal joint of the antenne has one or more spines on the outer side; the second basal joint is produced forwards on the outer side into a.long, pointed process whose outer margin is armed with several strong spines; acicles pointed, spiny, joined to the end of the second joint above the spiny process; fourth and fifth basal joints subcylindrical. The ischium of the maxillipeds is widened, produced forward at the antero-internal angle, and dentate on the inner margin; the iast two joints of the palp are not markedly dilated. The legs are covered with setose spines; chelipeds of moderate size; more or less unequal; fingers excavated within and furnished with calcareous teeth and corneous extremities. First of the three pairs of ambulatory legs subequal. Abdomen soft, the integument spiny; the first and last two segments strengthened by calcareous plates. 3 Type.—A. hispidus (STIMPSON). Acantholithodes hispidus (St.). Dermaturus hispidus Stimpson, Ann. N. Y. Lye. Nat. Hist., Vol. VII, 1860, p. 242. WuHiItEAves, Can. Nat. (2), Vol. VIII, 1878, p. 471. BouvigR, Ann. Sci. Nat. (7), Tome XVIII, p. 174 et seq., Pl. XI, figs. 3and 16; 1894, Pl. XII, figs. 2, 16, and 31; /bid. (8), Tome I, 1896, p. 19. Acantholithodes hispidus Houmss, Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 575. Carapace flattened, lyrate, covered with long setose spines, which become somewhat larger towards the margins. Median region tumid and separated from the cardiac by a very deep, transverse sulcus; a prominent depression between the median and branchial regions. Rostrum quite large and end- ing in three spines; a large fourth spine above and behind the median one. CALIFORNIA STALK-EYED CRUSTACEA. wi Eye-peduncles hispid and retractile against the anterior margin of the carapace. Antennules large. Antennz shorter than the carapace; first basal joint with a spine on either side; second joint produced into a long acute process whose outer margin is armed with four or five spines; acicle narrow, acuminate, armed externaily with numerous spines and about equaling the lower spiny process. Ischium of the maxillipeds with two small, subconical teeth on the outer surface near the antero-internal angle; merus hispid, not dilated, and longer than the ischium; last joint of the palp flattened below. Chelipeds unequal, very spiny; two or three very large spines on the anterior margin of the merus and carpus; in the larger hand the palm is inflated, the fingers deflexed, shorter than the palm, excavated within, corneous tipped, and furnished with calcareous, molar-like teeth. In the smaller hand the palm is not so strongly inflated, the fingers are longer and straighter and more deeply excavated within; the corneous tips are more extensive and the calcareous teeth are smaller and more numerous. Ambulatory legs spiny, somewhat compressed; dactyls about two-thirds the length of the propodi. Abdomen short, broad, and soft, the spines not so large as on the carapace; on the basal segment there is a median plate and two wide lateral ones; the plate on the penultimate segment is oblong; the last one is small, rounded, and - about as long as wide. Length of carapace from tip of rostrum, 58mm.; width of carapace, 56 mm. — Monterey (Stimpson)! Vancouver’s Is. (Whiteaves); iswaliornia.’! © A rare species. Stimpson’s description was drawn from a small female which was somewhat broader than long, .85 in. by .9in. Our description is taken from a single male from Monterey collected by Dr. Ritter. Genus Phyllolithodes Brandt. Carapace triangular, and provided with rounded tubercles which sur- round an excavated area on the dorsal surface; lateral margins armed with spines. Rostrum prominent, ending in two cornua. Acicle of the an- tenn cut into prominent lobes. Chelipeds unequal, spinose. Ambula- tory legs subcylindrical and armed withspines. Abdomen furnished with calcareous plates in which is a central membranous area. Type.—P. papillosus BRANDT. 122 CALIFORNIA ACADEMY OF SCIENCES. Phyllolithodes papillosus Brandt. Phyllolithodes papillosus BRANDT, Bull. phys.-math. Acad. St. Petersb., Tome VII, 1849, p. 175. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p..476; Proc. Acad. Nat. Sei. Phila., 1858, p. 2315, Amm: N. Y. Lye. Nat. Hist., Vol. VII, 1862, p.80; WuitTEAvEs, Can. Nat. (2), Vol. VIII, 1878, p. 471. Bouvirr, Ann. Sci. Nat. (7), Tome XVIII, 1895, p. 174, Pl XI) fig. 125 Pl. XI, figs. 14 and 25; Pl) XE news Ibid. (8), Tome I, 1896, p. 22. Petalocerus bellianus WuitE, Proc. Zool. Soc. London, 1856, p. 134, Pl. XLII; Ann. Nat. Hist. (2), Vol. XV, 1855, p. 307. Barts, Proc. Zool. Soc. London, 1864; in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 275. Carapace triangular and covered with strawberry-like tubercles which form a smaller elevated triangle whose sides are parallel to the sides of the carapace; a pair of large, deep pits on the dorsal surface. Rostrum prom- inent, divided into two, blunt, diverging, obliquely compressed horns, and bearing on the upper side a high, bilobed crest. On the under side of the rostrum there is a spiny process directed forwards and curving slightly upwards, above the base of which is a pair of much smaller spines. Antero-lateral margin constricted behind the hepatic regions. A subacute spine at the outer orbital angle which is directed forwards, downwards, and slightly inward; two spines of unequal length just external to the ‘postorbital; behind. the constriction, the antero-lateral margin is armed with three, long, slender, rather blunt spines which exceed the anterior ones in length. The postero-lateral angles of the carapace are produced outwards into flattened expansions which bear three blunt projections, the anterior one of which is elongated, slightly compressed, pointing upwards, outwards and forwards; the posterior projection is short and rounded. Posterior margin of the carapace transverse and furnished with a row of rounded tubercles. Antennal acicle furnished with three smooth, flattened, subequal, spatulate processes. The chelipeds are unequal and thickly covered with long, slender, more or less compressed, blunt spines, which are largest on the merus and carpus and gradually diminish in size toward the tip and lower margin of the hand; the two hands are similar; fingers stout, corneous-tipped, and excavated, the outer sides furnished with numerous tufts of seta. Ambulatory legs subequal, subcylindrical, and thickly covered with long spines similar to those of the chelipeds. Basal abdominal segment nearly vertical and deeply pitted on either side like the following segments. Length of carapace from tip of the rostral horn, 52 mm.; width between tips of postero-lateral horns, 51 mm. Kadiak Is. (Brandt); Vancouver’s Is.( Bate, Whiteaves) ; Monterey (White); ‘‘ California ’’, several specimens! CALIFORNIA STALK-EYED CRUSTACEA. 123 Phyllolithodes bicornis (Bate). f Petalocerus bicornis Bats, Proc. Zool. Soc. London, 1864, p. 664; in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 271. Phyllolithodes bicornis BouviER, Ann. Sci. Nat. (8), Tome I, 1896, p. 22. This species is evidently closely allied to papillosus. The carapace is ‘anteriorly produced into two, horizontal, horn-like processes. The sec- ond pair of antennz have a compound scale, consisting of two large and two short compressed processes, and a third joint is furnished with two or three sharp, short strong processes.’’ This species differs from papillosus, according to Bate, ‘‘in having a horizontal, bifurcate rostrum to the carapace, being more distinctly tuberculated, and in the pereopoda being more strongly spinated.”’ ; Color, yellow, with purple between the tubercles. ‘‘ Dredged in Esquimalt Harbor in ten fathoms of water.” The character of the rostrum does not distinguish this species from papillosus, as I have seen in the latter species all stages between prominent rostral spines and short, blunt knobs. Of the other distinguishing char- acters I cannot speak. Had not Bate specimens of papillosus from near the same locality for comparison, one would be inclined to regard this species as a syno- nym of the preceding one. Genus Cryptolithodes Brandt. Carapace transverse, nearly smooth, high in the middle and produced laterally into two wing-like expansions which cover the ambulatory legs. Rostrum broad, flattened, deflexed. Antero-lateral margins arcuate, often dentate, and meeting the postero-lateral margins at an angle. Ocular peduncles quite slender and approximated at their enlarged bases. An- tennules separated by a considerable interval, the first joint enlarged. First basal joint of the antenne very short; second joint with a crest above and a large process on the outer side; acicle broad and laminate; flagellum rather short. Ischium of the maxillipeds small, the inner margin denticu- lated; merus with the outer surface flattened; joints of the palp compressed. Chelipeds unequal and more or iess covered by the carapace; hands stout, compressed, the fingers excavated within. The three anterior pairs of oN 124 CALIFORNIA ACADEMY OF SCIENCES. ambulatory legs are subequal, crested above for a part of their length; dactyls with curved, acute tips. Abdomen flattened, triangular, fitting neatly into a depression in the sternum; behind the entire basal segment the abdomen bends abruptly, and the three following segments are divided into a median row of plates with a series of lateral plates on either side; the penultimate segment is devoid of lateral plates. and conceals the minute, triangular telson. Type.—C. typicus BRANDT. Cry ptolithodes ty picus Brandt. Cryptolithodes typicus BRANpDT, Bull. phys-math. Acad. St. Petersb., Tome VII, 1849, p. 175; and Tome XI, 1853. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857; p. 472, Pl. XIX; Proc. Acad. Nat. Sei: Phila., 1858, p. 231. Bats, Proc. Zool. Soc. London, 1864; in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 271. Bouvier, Ann. Sci. Nat. (7), Tome XVIII, 1895, p. 189, and (8), Tome I, 1896, p. 28. Carapace very wide. Rostrum very much deflexed, not distally widened, ‘the upper surface rounded, the anterior extremity truncated with the angles rounded. Lateral angles of the carapace acute; the posterior mar- gin bulges outward very slightly in the middle. Seen from below the carapace is strongly coneave and not at all flattened out towards the sides. The depression between the cardiac and median region is shallow. Teeth on the posterior margin of the carapace obscure; those on the antero- lateral margin small. Orbital sinus much less deep than in sitchensis. Chelipeds very low, rounded tubercles; large hand with the outer surface tuberculated and the upper side rounded; small hand acute above. The merus joints of the ambulatory legs are furnished below with a deep groove bounded by the sharp, produced, lower margins. Abdomen crossed by transverse ridges. Described from a small, dried specimen in the U. 8. National Museum (No. 3475) collected by Dall at Bel- koffsky Bay, Alaska (25 fathoms). Northern California (Brandt); ee (Stimpson); Vancouver’s Is. (Bate). im ‘al 7 w) ssh i ee ie CALIFORNIA STALK-EYED CRUSTACEA. 125 Cryptolithodes sitchensis Brandt. Pa Ot det oe Cryptolithodes sitchensis BRANDT, Bull. phys.-math. Acad. imp. sci. St, Petersb., Tome XI, 1853, p. 133; Mélanges biologiques, Tome I, p. 654. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 476; Proc. Acad. Nat. Sci. Phila., 1858, p. 231. NeEwcomse, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 26, Pl. 3: Bouvisrr, Ann. Sci. Nat. (7), Tome XVIII, 1895, p. 189, Pl. XI, figs. 11 and 15, Pl. XII, fig. 26; Ibid. (8), Tome I, 1896, p. 28. Cryptolithodes alta-fissura BatTr, Proc. Zool. Soc. London, 1864, p. 665; Ann. Nat. Hist. (3), Vol. XV, p. 485; in Lord’s Nat. in Vancouver’s Ts. VOloLE, 18665) p< 271. Carapace transversely subelliptical; median region highand surmounted by a ridge or crest, which extends to the middle of the front anteriorly; a pair of transverse ridges extending upon the hepatic region from the sides of the median elevation. Behind the median region the carapace is much inflated and rounded and furnished with a rounded elevation behind the depression at the posterior end of the median area. Median ridge obsoles- cent on the posterior portion of the carapace. Surface of the carapace smooth, microscopically very closely punctated, and furnished here and there with very small, rounded tubercles which, in some specimens, are almost entirely absent. The rostrum is deflexed at an angle of about 45 degrees and widened towards the distal end, which is transverse and fur- nished with a small median tooth. The lateral expansions of the carapace are produced forwards to about two-thirds the length of the front, forming a large deep, rounded, orbital sinus on either side. Theantero-lateral mar- gins are strongly arcuated, curving most sharply a little in front of the middle, the posterior third being almost straight and much more nearly longitudinal than transverse; the antero-lateral teeth are eight to ten in number and separated by concave interspaces of variable length; they are small and variable, often subacute, but sometimes represented by small undulations. The lateral angles of the carapace much exceed 90 degrees, and the lateral teeth are larger than those in front. Posterior margin strongly arched, and generally devoid of teeth, although there may be one - or two small ones near the lateral angles. A very prominent longitudinal crest on the pterygostomian regions. The ocular peduncles are smooth, bent upwards, and flattened above, the bases much enlarged. Thesuperior crest on the second joint of the antenne is short and high and the lami- nate outer process is triangular, produced at the postero-external angle, and extending forward in a sharp point considerably exceeding the fourth joint of the peduncle; the fifth joint is strongly compressed, having a high, thin crest on the upper side, which is produced at the distal end; flagellum nearly naked, scarcely as long as the peduncle; the acicle is very \/ /» 126 CALIFORNIA ACADEMY OF SCIENCES. thin, unarmed, about as wide as long, concave above, very strongly curved externally, the upturned inner margin somewhat irregular and nearly straight. The ischium of the maxillipeds is small, trigonal, the outer angle furnished witha small tooth; of the two edges which face the middle line, the inner one is the longer and finely denticulated, the outer one is furnished with two rounded teeth; the merus is smooth, flattened, strongly produced at the outer angle, the inner and outer margins sharp, the outer surface transversely concave; the joints of the palp are strongly compressed and crested above. Chelipeds stout; inner margin of the ischium acute; inner margin of the merus with a compressed (occasion- ally subconical) tooth; the carpus has a ridge on the outer surface, an acute projecting distal margin, and a tooth over the upper hinge joint of the hand; hands smooth, compressed, with a single, rounded, longi- tudinal ridge a little below the middle of the outer surface; the high upper crest is produced at the distal end; fingers short and stout, dactyl with a superior crest which is most prominent at the proximal end. Ambulatory legs smooth, naked, strongly compressed, with the upper edges acute; the lower side of the merus is flattened, not deeply erooved as in typicus; the acute inferior margin of the propodi ends dis- tally in a short spine; dactyls spinulous below, about three-fourths the length of the propodi. The sides of the abdomen in the male are nearly straight, but in the female they are convex and more or less unequal; the basal joint is nearly vertical, with a depression on either side of the cen- ter; the posterior side more strongly concave in the female than in the male; the sutures of the median and lateral pieces alternate, those between the lateral pieces being transverse in the male but oblique on one side in © the female, the penultimate segment is oblong, distally widened, and concave at the distal end; the telson is very small and fits into the space between the bases of the chelipeds. Length of Carapace. Width of Carapace. Male? 22 oocaeee Aj. SU ye tes ACRE ace Sunes eae 64 mm. ps aie See Wee: Ee 3G .9 GMM Yio Ani oalc oe ee eine ieee 53 mm Se NTR T ait 3D, THM. 1 4% «fees «ch kc eee 50 mm aoe eRA NT Sat ey UNIAN ie i ioe ee 47 mm Fe yes Ng eet a! 1G: ’ Me Ae eye Cee eee Eee 23;. mm MemaAles: a. eke BGO TORTS Jae ant ee ete er ane 49.5 mm Bor Bee ede DD,» SHINER so rs eae pe po 36 mm. The males I have seen in a fresh condition were all of a uniform, bright red color} Two/( females) seen were red but with a purplish tinge irregularly marked with blotches of a lighter color} This species is often seen at low tide on the sides of rocks where, at some distance, CALIFORNIA STALK-EYED CRUSTACEA. 127 it might readily be mistaken for a species of bright red incrusting sponge which is found in similar situations. Near Sitka (Brandt); Alaska! Vancouver’s Island (Bate); Victoria; Queen Charlotte Island (Newcombe); British Columbia (Bouvier); Port ‘Townsend! Cape Mendocino! Point Arena, Calif.! In young specimens the posterior margin of the cara- pace is straighter than in adults; the ridge on the outer surface of the hand is plainer and there may be one or more additional ridges more or less plainly visible. In females the posterior portion of the carapace is very tumid, and the posterior margin bulges backwards more strongly than in the males. Cryptolithodes brevifrons Miers. Cryptohthodes brevifrons MixERs, Proc. Zool. Soc. London, 1879, p. 48, Note. Bouvier, Ann. Sci. Nat. (8), Tome 1, 1896, p. 28. . This species is described in a foot-note as follows: ‘¢ There is a dried specimen in the [British] Museum from Vancouver’s Island which closely resembles C. typicus, but is distinguished by the form of the rostrum, which is obtusely triangular, and does not project be- yond the anterior margin of the carapax. This I propose to call C. brevifrons.” Genus Lopholithodes Brandt. Carapace broad, pentagonai or hexagonal, convex above; with the mar- gins and upper surface armed with setose tubercles. Rostrum short, spiny. Acicle of the antennz with the margins and generally the upper surface spiny. The terminal joint of the mandibular palp is elongated and flat- tened, and bent backward so as to lie between the concave inner faces of the mandibles. Chelipeds unequal, tuberculated; carpus with a promi- nent lobe on the inner side. Ambulatory legs short, tuberculated, and capable of being folded under the carapace. Basal segment of the abdomen KH) 128 CALIFORNIA ACADEMY OF SCIENCES. entire, the three following segments with lateral plates and also small marginal plates on one or both sides; penultimate joint devoid of lateral plates; telson very small. Type.—L. Mandtii BRANDT. Large crabs of rough aspect, confined to the Pacific Coast of North America. In this genus the chelipeds and ambulatory legs, when folded, fit neatly together,-thus affording pro- tection to the mouth-parts and under side of the body in a very effective manner. Until the present time Lopholithodes Brandt has been ranked as a synonym of Echinocerus White, the date of Brandt’s genus being given as 1849 and that of White’s as 1848. But while Vol. VII of the Bulletin of the St. Petersburg Academy of Sciences bears the date 1849, the number containing the description of Lopholithodes bears the date of publication June 29, 1848. The part of the Proceedings of the Zoological Society of London in which White’s. description of Hchinocerus appeared was received from the printer November 14, 1848. The name Lopholithodes, therefore, has priority. ! Lopholithodes Mandtii Brandt. Lopholithodes Mandtii BRanvtT, Bull. phys.-math. Acad. St. Petersb., _ Tome VII, 1849, p. 174. Echinocerus cibarius WHITE, Proc. Zool. Soc. London, 1848, p. 47, Pls. Land III; Cat. of Crust. in Brit. Mus., p. 56. Stimpson, Journ. Bost. Soc. Nat. Hist. Vol. V1, 1857, p. 477; Proc. Acad. Nat. Sci. Phila., 1858, p. 231. Bars, in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 276. WHITEAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. NErwcomesz, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 26. Bouvier, Ann. Sci. Nat. (7), Tome XVIII, 1894, p. 184, Pl. XI, fig. 138, Pl. XII, figs. 13 and 24; Ibid. (8), Tome I, 1896, p. 27. Ctenorhinus setimanus GIBBONS, Proc. Cal. Acad. Sci., Vol. I, 1855, p. 48. Echinocerus sentimanus STIMPSON, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist. Vol. VI, 1857, p. 477; Proc. Acad. Nat. Sci. Phila., 1858, p. 231. Bouvier, Ann. Sci. Nat: (7), Tome XVIII, 1894, p. 184; /bid. (8), Tome I, 1896, p. 27. CALIFORNIA STALK-EYED .CRUSTACEA. 129 Carapace strongly convex, wider than long, the whole surface—the large tubercles as well as the intervening spaces—roughened by small short, setose tubercles which vary greatly in size from low, nearly smooth swellings to rough, subconical projections. Median region very promi- nent, having a subconical apex a little in front of the middle and a cluster of small, rounded, or subacute tubercles near the posterior end; a large, subconical elevation on the cardiac region in front of which, on either side, is a depressed, comparatively smooth area; median and cardiac areas separated by a comparatively smooth sulcus; a large subconical elevation on the branchial regions and two smaller tubercles on the posterior mar- gin of the carapace. A peculiar smooth wart-like prominence on either side of the median area. Rostrum short and consisting of a strong sub- conical spine above the base of which is a knob bearing two lateral spines or tubercles with (generally) a spine or tubercle above and below the notch between them. A deep, rounded sinus on either side of the front which serves as an orbit, external to which is an acute spine; beyond this spine is a large spine which extends nearly as far forwards as the rostrum and in the sinus between the latter spine and the postorbital there is generally a small spine or tooth. The antero-lateral margin is armed with a variable number (about eight) of prominent spines and several smaller ones; the middle portion is convex, flattened, and produced. A large, knob-like prominence at the postero-lateral angles which is separated from the last antero-lateral spine by a conspicuous sinus. Ocular peduncles two-jointed, thickly set with spines above, and not nearly reaching the tip of the ros- trum. Acicle of the antenne narrow, tapering, much exceeding the tip of the peduncie, and furnished with about thirty strong, smooth, sharp spines. Ischium of the maxillipeds armed within with dark-colored teeth; a tooth on the outer surface near the antero-internal angle; exognath somewhat exceeding the merus. Chelipeds short, the first and second joints with irregular tuberculous projections on the under side; one or more prominent spines on the antero-internal angle of the merus; the lobe on the inner side of the carpus is very large and acute and margined with sharp, spine-like tubercles; spines on the outer surface of the hands large and numerous; four or five large spines on the upper margin of the larger hand and three or four on that of the smaller. Ambulatory legs subequal, strongly tuberculated, quadrangular at the base; carpi and pro- podi subcylindrical but very irregular; dactyls short and stout. The basal abdominal segment is strongly concave behind, especially in the female, and nearly at right angles to the carapace; there are two prominent tuber- cles near the middle and several on the margins; the remaining segments are studded with numerous subconical tubercles; between the median plates there are small, transverse, secondary plates which may be divided by longitudinal fissures; penultimate segment oblong and distally concave. 9 130 CALIFORNIA ACADEMY OF SCIENCES. The abdomen of the male is subtriangular behind the basal segment; that of the female rounded at the sides and tip and bent to one side. Length, 200 mm.; breadth, 245 mm.; length of larger cheliped, 202 mm.; length of first ambulatory leg, 195 mm. Some specimens are widest between the tubercles at the postero-lateral angles, while others are widest between the tips of the last spines on the antero-lateral margin. Sitka (Brandt); Vancouver’s Is. (Bate); near the mouth of the Columbia River (White); Farallon Is- lands! Monterey! * yo Lopholithodes foraminatus, St.) Echinocerus foraminatus Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 79. NeEwcomsBgz, Bull. Nat. Hist. Soc. Brit. Col., 1893, p. 27, Pl. III. Bouvier, Ann. Sci. Nat. (7), Tome XVIII, 1894, p. 184; Ibid. (8), Tome I, 1896, p. 27. Carapace depressed, wider than long, the tubercles on the upper surface mostly small and subconical; median region elevated, furnished with a conical tubercle near the middle, and a group of smaller tubercles near the posterior end; cardiac region with a few subconical tubercles; an irregular line of tubercles extending from the median area to the postero-lateral angles of the carapace; the greater portion of the upper surface is covered with depressed, more or less papillose elevations. A smooth spot on either side of the median area asin Mandtii. Rostrum rather short, with the median spine much as in Mandtii and a cluster of spiny tubercles above the base. External to the orbits there are three spines in a nearly trans- verse line, behind which the antero-lateral margin is at first concave, then strongly convex; the spines on the convex portion of the margin are large and subconical; behind the convex part the margin is concave and devoid of spines; posterior margin of the carapace arcuated and studded with conical tubercles. Eye-stalks spinulous. Antennal acicle with strong spines on the margins but aimost devoid of spines on the upper surface. Chelipeds very spiny; merus with the antero-internal angle produced and witha flattened, nearly semicircular area at the supero-distal angle; carpus with the inner lobe prominent, and with the outer edge excavated, form- ing a smooth, deep, rounded sinus which, when approximated to the shallower corresponding sinus on the anterior edge of the carpus of the first pair of ambulatory legs, forms a smooth, nearly circular hole; hands quite similar to those of Mandtii. Ambulatory legs and abdomen much as in the preceding species. CALIFORNIA STALK-EYED CRUSTACEA. 131 ‘‘ Near San Francisco” (Stimpson); Farallon Islands! Victoria (Newcombe). Easily recognized by the remarkable foramen between the chelipeds and the first ambulatory legs. Genus Lithodes Lair. Carapace broad, more or less depressed and spiny. Rostrum generally prominent and armed with spines. The large basal segment of the abdo- men may be entire or composed of three or more plates; the three follow- ing segments are furnished with median, lateral, and marginal plates. Lithodes Rathbuni Benedict. Lithodes Rathbuni BENeEpicT, Proc. U.S. Nat. Mus., Vol. XVII, 1894, p. 482. Paralithodes ? Rathbuni Bouviser, Ann. Sci. Nat. (8), Tome I, 1896, p. 23. Carapace armed with four spines on the gastric area and four (two long and two short ones) on the cardiac; branchial region with six spines. Rostrum composed of five branches, the main stem sharply bent upward and deeply bifurcate; the lower spine projecting nearly horizontally. The right cheliped is rather slender; the carpus has more than twelve spines; palm with two rows of four spines each on the median, outer surface. Ambulatory legs slender and very spiny. Antennal acicle very long and slender. Known from a single male specimen taken off San Simeon Bay, Calif., in 211 fathoms. Lithodes californiensis Benedict. Lithodes californiensis BENEDICT, Proc. U.S. Nat. Mus., Vol. XVII, 1894, p. 483. Paralithodes? californiensis Bouvier, Ann. Sci. Nat. (8), Tome I, 1896, p. 23. Very closely allied to LZ. Rathbuni, but having the spines on the cara- pace shorter and stouter and the rostrum simply bifid instead of divided into two, well-developed horns. 132 CALIFORNIA ACADEMY OF SCIENCES. Known from two female specimens taken off Santa Cruz Island, Calif., in 155 fathoms. There are several other species of Lithodes found further north on our coast, viz., L. brevipes, camtschati- cus, equispinus, and Couesi. Panamensis occurs on the west coast of Central America. Legion PAGURINEA. Tue HERMIT CRABS. Carapace elongated, generally membranaceous behind the cervical groove. Rostrum small. Abdomen generally long and unprotected by calcareous plates. First two pairs of ambulatory legs well developed, last two pairs small. Last abdominal segment furnished with a pair of ap- pendages. Species usually inhabiting the coiled shells of Mol- luses. Family PAGURIDA. Antennular peduncle of moderate size, the first joint short and stout, second and third slender and cylindrical; both flagella small. The ped- uncle of the antenna is subcylindrical; acicle lanceolate. Gills phyllo- branchiate. Abdomen soft, asymmetrical. ‘ Genus Pagurus Fabr. Front with a rostral projection. Ocular peduncles with the basal scale of moderate size and wide apart. Antennal acicle slender and elongated; flagellum long and naked. Chelipeds rarely subequal, the right usually the larger. Fourth pair of pereopods subchelate. First and second abdominal segments without genital appendages. Males without a pro- truded vas deferens. Type.—Pagurus Bernhardus (LinN.). It has been recently pointed out by Mr. J. EH. Bene- dict’ that the generic name Hupagurus will have to be superseded by the older name Pagurus. ‘‘I believe,” lAnn, Nat. Hist. (6), Vol. XVIII, July, 1896, p. 99. CALIFORNIA STALK-EYED CRUSTACEA. 133 says Mr. Benedict, ‘‘ that the genus Pagurus, as now constituted, does not contain a single one of the original species placed in it by Fabricius; but be that as it may, a valid and therefore imperative reason for making the change lies in the fact that Bernhardus was designated as the type of Pagurus by Latreille in 1810'.” The old genus Pagurus was divided by Brandt in his work on Crustacea in Middendorff’s Siberische Reise? into sev- eral subordinate sections- or subgenera, one of which containing Bernhardus and its allies was called Hupagu- rus or the true Pagurus. Brandt’s subgenus Lupagurus was afterwards raised to the rank of a genus with Bern- hardus as the typical species, thus depriving the older genus of its type. Hupagurus must, therefore, give way to the older name. The species of the genus Pagurus are very numerous. and often difficult to determine. The west coast of North America seems especially rich in these forms. The Albatross collections in museums of the University of California and the California Academy of Sciences have been of especial value to mein studying our species of this genus, and at the U. S. National Museum, through the kindness of Mr. Benedict, I have examined some species from this coast not previously met with. At one place or another I have seen all the species of Pagurus here described. A few Atlantic species have been reported from this coast: P. Kroyeri from Puget Sound by Stimpson, and Bernhardus from the north Pacific by Brandt. The Bernhardus as Mr. Benedict suggests (zn Iztt.) may be aleuticus or alaskensis. The genus Pagurus is divided by Mr. Benedict into four subgenera as follows: 1Consid. Gener. Crust., p. 421. 2Bd.II, TheilI, p. 105. 134 CALIFORNIA ACADEMY OF SCIENCES. Subgenus Eupagurus. Pagurus ‘‘ with the large surface of the left hand horizontal. Species not inclined to be hairy.” . Typical species, P. Bernhardus (Liny.). P. aleuticus, alaskensis and ochotensis Gelsne here. All are large species with the upper surface of the left hand triangular and the edges prominent. Subgenus Trigonocheirus. Pagurus ‘‘with the outer face of the left hand oblique and more or less triangular. In some cases it is flattened, in others concave, or very much swollen as in hirsutiusculus. The species are frequently very hairy.” Typical species P. trigonocheirus STIMPSON. This group includes the greater number of the spe- cies here described, viz., trigonocheirus, capillatus, Dalli, Brandti, Rathbuni, Tanneri, confragosus, Newcombet, Kennerlyi, hirsutiusculus, Samuelis, minimus, granosi- manus, and Hemphillir. In many of the species the left hace is small and rounded, presenting no distinctly outlined upper face. In others the left hand is trigonous, with a well-defined, triangular face bounded by prominent edges. The first five species mentioned are very closely allied. In all three the chelipeds are very unequal, the carpus of the larger one oblong with the sides flattened and the upper surface furnished with short spines or spiny granules; the hand is oblong, evenly convex above, and armed like the carpus; the smaller hand has a prominent, triangular supero-external face. Rathbwni can be dis- tinguished by the peculiar round pubescent patch on the upper side of the carpus. Tannerz and confragosus are distinguished from all the others by the ridge on the upper surface of the larger hand. Newcombez can be CALIFORNIA STALK-EYED CRUSTACEA. 135 recognized by the bright red tips of the chelipeds and dactyls. Kennerlyi has the spines on the upper surface exceptionally long and slender. Hirsutiusculus, granosi- manus, Samuelis and Hemphillii are small, littoral spe- cies. They are the forms most commonly met with along the shore at low tide. Granosimanus may readily be distinguished from the other three by its rounded, median, frontal projection and its broader left hand. Hirsutiusculus may be recognized by its strong pubes- cence, the short anterior portion of the carapace, and short eye-stalks. Subgenus Elassocheirus. Pagurus ‘‘with the left hand very smail, in some cases almost rudi- mentary. The large hand is always wide or broadly ovate, excepting in munitus and Gilli where the carpus is exceedingly wide and the hand mod- erately so. Eye-scale semicylindrical, grading to triangular, sharp and pointed. Species not hairy; many of them smooth and glabrous.” Typical species, P. tenuimanus (DANA). Besides the type the only species of this group in our limits are mwnitus and californiensis. Subgenus Labidocheirus. Pagurus ‘‘with the manus of the left cheliped cylindrical.” Typical species, P. splendescens (OWEN). Pagurus alaskensis (Benedict). Eupagurus alaskensis Bunepict, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 2. Anterior portion of the carapace a little wider than long and sparingly pubescent. Median tooth of the front triangular, acute, and produced considerably beyond the lateral ones, which are rounded, but furnished with a small spine. Ocular peduncles short, stout, constricted in the middle, and about one-half the length of the anterior portion of the carapace. Eye-scales subovate, with the apex blunt or subacute, the upper surface concave and the subterminal spine visible from above. Antennal acicle trigonous, reaching much beyond the tips of the ocular peduncles; 136 CALIFORNIA ACADEMY OF SCIENCES. the upper surface flat, the inner margin spinulous and pubescent. Cheli- peds spinulous, markedly unequal, and almost devoid of pubescence. Merus of the larger cheliped oblong, trigonous, the upper surface with a few transverse, granulated, or spinulous ruge@ near the distal end; carpus with the inner and outer sides flattened, the upper surface convex and spinous, with a row of stout spines on the inner edge; hand oblong, slightly wider than the carpus; inner side flattened; upper surface mod- erately convex and covered with acute granulations; outer margin defined by a row of spiny granules; fingers armed within with tuberculous teeth. Merus of the smaller cheliped compressed; carpus with the inner side flattened, the supero-lateral surface convex and covered with spiny gran- ules; outer margin raised; upper surface convex; upper internal margin armed with a row of strong spines, external to which is a similar row on the upper surface; hand oblong, flattened, and covered with sharp gran- ules; outer margin raised; upper surface convex, the inner surface of the palm and dactyl flattened; dactyl much longer than the palm, the upper surfacerounded. Ambulatory legs with the merus and propodus spinulous above, the dactyls long, curved, twisted, becoming thinner but not de- creasing in width towards the tip, and pectinated below with slender, closely set, parallel spinules. - : Concerning the color of this species Mr. Benedict says: ‘‘ In alcoholic specimens the general color above is a light purple with iridescent reflec- tions; below light tinged with reddish. A red streak runs around the prehensile edge of the thumbs and behind the dactyls to the inner margins of the hands. There is an oblong patch of red on the outer distal mar- gins and on the upper surface of the merus joints of the cheliped. The lower outer surface of the carpal joints of ambulatory legs is pointed with » red. The propodal joints and dactyls are longitudinally streaked with red. ) This species is very close to Bernhardus but is easily distinguished by its broader and shorter left hand, by the wide dactyls of the ambula- tory legs, by the acicula, and by its pearly iridescence.” Alaska; off Washington in 31 to 38 fathoms (Bene- dict). Pagurus aleuticus (Benedict). ELupagurus aleuticus BengepictT, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 3. Closely allied to alaskensis. 'The teeth on the front are sharper, the lateral ones acute. Eye-peduncles stouter and over one-half the length of the anterior portion of the carapace. Antennal acicle similar to that of alaskensis but broader at the base and not projecting so far beyond the tips of the eye-peduncles. The chelipeds are very similar to those of alaskensis but the spines are more prominent; a prominent line of spiny granulations a4 CALIFORNIA STALK-EYED CRUSTACEA. 187 on the upper surface of the dactyl, which is represented only by an obtuse ridge in the preceding species. On the dactyl of the smaller hand there is an oblique, smooth surface visible from above which is not found in alaskensis, and the supero-external surface of the carpus of the smaller cheliped is much more flattened and the outer edge of the hand more ele- vated and more sharply spinose. The dactyls are similar in form to those of alaskensis, but are easily distinguished by the deep sulci on the upper side bounded by sharp, minuteiy spinulous edges. Aleutian Is. (Benedict). Pagurus ochotensis (Brandt). Eupagurus ochotensis BRANDT, Middendorff’s Sibirische Reise, Bd. II, Th. 1, 1851, p. 108. Stimpson, Proc. Acad. Nat. Sci. Phila., 1858, p. 248. Bernhardus armatus DANA, Proc. Acad. Nat. Sci. Phila., 1851, p. 269; Crust. U.S. Expl. Expd., Part I, 1852, p. 442, Pl. XXVII, fig. 2. ELupagurus armatus STIMPsoN, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 484. Bate in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 278. WHITEAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. Anterior portion of the carapace wider than long; the three projections of the front about equally advanced. Ocular peduncles short and stout, but little over one-half the length of the anterior portion of the carapace. Eye-scales acute, witha prominent, subterminal tooth. Acicle slender, flattened above, and extending far beyond the eyes. Chelipeds spiny, the carpus of the larger one with the sides flat and sharply granulated, the upper surface evenly rounded and spiny, with a row of longer spines along the inner margin; hand oblong, evenly rounded above, and spiny like the carpus; dactyl with a row of spines on the outer edge and another row near the middle of the outer surface. Smaller cheliped reaching a little beyond the base of the dactyl of the larger one; carpus and hand spiny above, the inner edge of the palm somewhat raised-and armed with two rows of spines, the outer spines being the shorter. The meral, carpal, and propodal joints of the ambulatory legs are spinous above; the dactyls are spinous, twisted, grooved on each side, and nearly as long as the two preceding joints. A large species. Described from specimens (No. 14951) in the U.S. National Museum. Puget Sound (Dana); Sitka! San Diego, Calif.! 138 CALIFORNIA ACADEMY OF SCIENCES. Pagurus trigonocheirus (St.) Hupagurus trigonocheirus Stimpson, Proc. Acad. Nat. Sci. Phila., 1858, pp. 237 and 249. Murpocu, Rep. Int. Polar Expd. to Pt. Barrow, Part IV, 1885, p. 138, Plate I, figs.1, la and 1b. Bernepict, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 1. The three prominences of the front are subequal, the lateral ones end- ing inaminute spine. The anterior portion of the carapace is longer than wide and more or less pubescent; a tuft of hairs on the median pro- jection of the front. Eye-stalks rather stout and much shorter than the width of the carapace. LEye-scales subovate, acute, more or less pubes- cent and not at all channeled above. Acicle rounded, pubescent, and reaching the tip of the ocular peduncle. Chelipeds pubescent; right cheli- ped with the ischium denticulated on the lower margin; merus trigonal, supero-lateral surface much rounded and scabrous; lower surface spinu- lous; the supero-distal margin spiny; carpus thick, oblong, the upper surface convex and covered with short spines; inner and outer surfaces flattened and less strongly spinose; hand oblong, slightly narrower than the distal end of the carpus; the upper surface convex and covered with short spines; inner margin obtuse; outer margin subacute, except near the base, and spinulous; lower surface spinulous near the margins; fingers shorter than the palm; dactyl narrower than the pollex, with a spinulous, subacute outer margin and a longitudinal row of short spinules on the upper surface. Smaller cheliped reaching beyond the carpus of the larger one; merus compressed, spinulous below; carpus compressed, distally widened, especially on the lower side, the upper margin spinulous; hand trigonous, the supero-lateral face wide, flattened, subspinulous or granu- lar, the edges acute; the upper edge is the longer, with the posterior por- tion convex and spinulous, the spinules becoming smaller on the pollex; dactyl narrow and nearly smooth. Ambulatory legs scabrous or spinulous on the margins; the two small posterior pairs are smooth and strongly pubescent. Length, 3 inches. Arctic Ocean and Behring Sea (Stimpson); Alaska! Pagurus capillatus (Benedict). Eupagurus capillatus BENEDICT, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 8. Anterior portion of the carapace a little longer than wide. Median frontal lobe broadly rounded. Ocular peduncles more than two-thirds as long as the anterior portion of the carapace, and slender. Acicle not reaching to the tip of the eye. Hand of larger cheliped twice as long as CALIFORNIA STALK-EYED CRUSTACEA. 139 wide and rather sparsely set with sharp, stout spines. Hand of smaller cheliped narrow, slender, with the supero-external surface narrower than in trigonocheirus, and the edges not so strongly produced. Dactyls of the ambulatory legs slightly twisted. Very near pubescens from the Atlantic. Foran enu- meration of the differences, see Benedict, l. c., p. 9. Alaska; Washington in 40 fathoms (Benedict). Pagurus Brandti (Benedict). EHupagurus Brandti BeNepict, Proc. U. S. Nat. Mus., Vol. XV, 1892, p. 9. Median portion of the front triangular and produced scarcely beyond the lateral teeth. Eye-stalks rather stout. Acicles scarcely extending beyond the eyes. Hand of the larger cheliped twice as long as wide. Hand of the smailer cheliped as in trigonocheirus. Ambulatory legs almost entirely unarmed; dactyls slender, slightly twisted and longer than the _propodi. Color of alcoholic specimens ‘‘ everywhere reddish. Spines everywhere red or red pointed,” Aleutian Islands (Benedict); Captain’s Harbor, Una- laska, 80 fathoms! Pagurus Dalli (Benedict). Eupagurus Dalli BExevict, Proc. U. 8. Nat. Mus., Vol. XV, 1892, p. 9. Very closely allied to Brandti. Median lobe of front triangular, acute, and produced further beyond the lateral teeth thanin Brandti. EHye- peduncles more slender than in Brandti or trigonocheirus. Acicle reaching but little beyond the eyes. The carpus of the small cheliped is not so high, and the supero-external surface of the hand is not nearly so wide at the base as in the last two species. The dactyls of the ambulatory legs are long, more slender than in Brandti, and not twisted. Aleutian Islands, Unalaska (Benedict); Iluliuk Har- bor, 10 fathoms! ‘‘ This species,” says Bouin ‘is closely allied to Kroéyeri, trigonocherrus, and Dalli. The small chelipeds of all have the triangular outer face, and in general the species resemble each other very much. With a large series of all I find substantial specific characters. 9 > 140 CALIFORNIA ACADEMY OF SCIENCES. Brandti and Dalli are found together in a depth of from 15 to 85 fathoms in the Aleutian Island region. They grow to a length of three inches. They are separated at sight by a sharply defined band of white on the distal end of the merus joints of Dall, which is entirely want- ing in Brandt.” Pagurus Rathbuni (Benedict). EHupagurus Rathbuni Benepict, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 14. Near trigonocheirus, but with the median projection of the front acute and projecting considerably beyond the lateral teeth. Eye-peduncles stout, not two-thirds as long as the anterior portion of the carapace. Eye- scales ovate, concave above, with a subterminal spine. Chelipeds resem- bling those of trigonocheirus; the hand of the larger one over twice as long as wide. Hand of the smaller cheliped much narrower than in trigono- cheirus, the edges not nearly so prominent; fingers curved downwards. Ambulatory legs spinulous. above; dactyls twisted, longer than the pro- podi, and nearly devoid of spinules with the exception of a few on the lower side near the tip. Alaska. This species is distinguished from all the other hermit crabs of the coast by the possession of a circular patch of fine hairs on the antero-internal angle of the upper surface of the carpus of the large cheliped. It is also distinguished from the four preceding species by its larger median frontal prominence. °* Pagurus Tanneri (Benedict). Eupagurus Tanneri BENEpIcT, Proc. U. S. Nat. Mus., Vol. XV, 1892, p. 10. Anterior portion of the carapace as wide as long. Median frontal tooth triangular, much larger and projecting much further forward than the lateral teeth, which are broadly triangular, subacute, and furnished with a terminal spine. Ocular peduncles stout; cornez dilated; scales ovate and furnished with a subterminal spine which is visible from above, the upper surface more or less concave, The outer spine of the second basal antennai joint reaches to, or beyond the middle of the fourth joint; acicle CALIFORNIA STALK-EYED CRUSTACEA. 141 slender and exceeding the tips of the ocular peduncles. Chelipeds spinu- lous and pubescent; merus oblong, the supero-lateral surface rounded and scabrous, or spinulous; lower surface spinulous, and devoid of marked prominences; carpus thick, oblong, the inner and outer surfaces flat; upper surface convex, spinous, with the inner and outer edges defined by rows of spines; hand about as wide as the carpus and over twice as long as broad; on the upper surface behind the gap between the fingers is a longitudinal ridge, which is continued forward upon the pollex though diminishing in size; the ridge bifurcates behind, forming two lower, less marked ridges which diverge toward the posterior margin; on either side of this ridge the surface of the hand is depressed and less spiny than elsewhere; fingers flattened and bent downwards; a row of short spinules on the outer margin of the dactyl and a similar row near it on the upper surface. Merus of the smaller cheliped compressed, spinulous below; carpus compressed, but widened at the distal end of the lower side, having a rounded, spinulous process at the antero-external angle; upper edge with two series of spines; hand trigonous, supero-external face subtrian- gular, deeply concave; edges prominent, spinulous; a row of spinules extending from the proximal margin to the upper ridge, which it meets a short distance from the base; the surface below the upper edge is concave and limited below by a row of spinules; fingers curved downwards; dactyl slender, nearly smooth. Ambulatory legs subequal, spinulous above; dactyls slender, longer than the propodi, quite strongly twisted, and obsoletely spinulous. Alaska; off Washingt n from 167 to 559 fathoms (Benedict). Pagurus confragosus (Benedict). Eupagurus confragosus BrNEpIcT, Proc. U.S. Nat. Mus., Vol. XV, 1892, pi 1. Very near Tanneri. The median lobe of the front is acute but not so prominent as in that species. Ocular peduncles short, stout, distally widened, and not two-thirds the length of the anterior portion of the carapace. Chelipeds much like those of TYanneri; the larger hand is slightly wider, with the outer margin more curved; the triangular area between the two diverging ridges of the upper surface is wider, and the more prominent inner ridge is interrupted at aboutits posterior third, the two parts changing direction at this point; the elevated apex of the tri- angle where the ridges meet is not continued forward as a high crest as in Tanneri, but the ridge begins to descend immediately toward the pollex; fingers much flattened, but scarcely curved downward. Smaller hand scarcely distinguishable from that of Tanneri. Ambulatory legs subequal; dactyls slender, curved, twisted, nearly devoid of spinules. 142 CALIFORNIA ACADEMY OF SCIENCES. ‘¢ The two species,” says Benedict, ‘‘ are readily separated by the inner side of the raised triangle of the larger hand. In this species it cuts off the inner depression of the upper surface from the proximal margin of the palm, while in Z'anneri this depression is allowed to reach the margin by a curvature of the ridge.” Alaska! off Washington and the mouth of the Colum- bia River in 68-178 fathoms (Benedict). Der VAY MAY Pagurus:-Newcombei (Benedict). Hupagurus Newcombei BENEDIcT, Proc. U.S. Nat. Mus., Vol. XV, 1892, Dali: Anterior portion of the carapace longer than wide. Median frontal tooth short and obtuse; the lateral teeth broadly rounded and slightly promi- nent. LEye-stalks moderately slender and a little over one-half the length of the anterior portion of the carapace. Eye-scales subtriangular, slightly concave above, the subterminal spines prominent. Antennal peduncles longer than the eye-stalks; acicle nearly reaching the tip of the eye. Chelipeds very unequal; merus of the larger one stout; the outer surface is convex and nearly smooth, but there are some transverse spinulous ruge near the supero-distal angle; two prominent tubercles on the lower side; carpus oblong, scarcely one-half longer than wide, and of nearly equal width throughout; outer surface flat; inner surface a little concave; upper surface evenly convex, thickly covered with spiny granules, and bordered on the inner side by a row of short spines, internal to which is a parallel row of smaller spines; hand no wider and scarcely longer than the carpus, and covered above with spiny granules; outer edge acute, especially on the pollex, and armed with short spines; fingers with minute, corneous tips; dactyl margined with spiny granules. Small cheliped pubescent; merus compressed, and more or less spinulous below; carpus armed with ten or twelve long, curved spines on the upper edge; outer surface with several spines on the upper side and spiny granules further down; hand longer than the carpus, with a triangular, spinulous, supero- external face; a sulcus on the posterior portion of the upper edge. The carpus and propodus of the ambulatory legs are armed above with spines, which are much smaller in the second pair, where they may be absent on the propodi; dactyls spinous below and about as long as the propodi. Color in alcohol: chelipeds reddish, the tips of the fingers bright red. The ambulatory legs are spotted with red; the base of the dactyls and the distal end of the propodi are bright red. British Columbia (Benedict); Sitka! Port Townsend! J ee CALIFORNIA STALK-EYED CRUSTACEA. 143 Pagurus Kennerlyi (Sz. ). Eupagurus Kennerlyi Stimpson, Proc. Acad. Nat. Sci. Phila., 1864, p. 153. BENEDIcT, Proc. U. S. Nat. Mus., Vol. XV, 1892, p. 19. WALKER, Trans. Liverpool, Biol. Soc., Vol. XII, 1898, p. 275. Anterior portion of the carapace a little longer than wide. Median tooth of the front triangular, low, subacute; laterai teeth rounded, with a prom- inent submarginal spine making them appear acute. Eye-peduncles moderately slender and fully two-thirds the length of the anterior portion of the carapace; cornex not large and scarcely dilated. Eye-scales sub- ovate, dorsally concave, with a prominent subterminal spine, which is visiblefrom above. Antennal peduncle exceeding the eyes; acicle rounded above and seldom reaching the tip of the eye. Chelipeds very unequal, pubescent, and spiny; merus nearly trihedral, the outer surface convex and nearly smooth; carpus short, distally widened; inner surface smooth, glossy, more or less concave; outer surface flattened, nearly smooth; upper face covered with scattered hairy prominences; inner edge furnished with seven or eight slender curved spines; a parallel row of three or four spines near the distal end; hand oblong, wider and much longer than the carpus; upper surface with seven to nine rows (including the marginal rows) of spines, a tuft of long hairs arising from the base of each spine; fingers corneous tipped; outer margin of the dactyl with a prominent row of Short spines and a similar row on the upper surface. Merus of the left cheliped with the lower side spinulous; carpus compressed, with a double row of spines on the upper surface; hand with the supero-external sur- face widened at the base and thickly set with strong spines; upper edge with a smooth sulcus. Ambulatory legs pubescent, the upper sides devoid of spines except on the carpus of the anterior pair; dactyls not twisted, about equalling the propodi and armed below with slender spines. Alaska (Benedict); Puget Sound (Stimpson)! Kadiak ist late 55/35". 30° No; lon 162° 31° 45" We; 29 dathoms! Pagurus hirsutiusculus (Dana). _ Bernhardus hirsutiusculus Dana, Proc. Acad. Nat. Sci. Phila., 1851, p. 269; Crust. U. S. Expl. Expd., Part 1, 1852, p. 443, Pl. XXVII, fig. 3. ELupagurus hirsutiusculus STIMPSON, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 484; Proc. Acad. Nat. Sci. Phila., 1858, pp. 237 and 249. Carapace and legs pubescent; anterior portion of the carapace a little wider than long; median tooth triangular and acute; lateral teeth obso- lete. Ocular peduncles short, about one-half the length of the anterior portio n of the carapace; cornew scarcely dilated. FEye-scales oblong / 144 CALIFORNIA ACADEMY OF SCIENCES. obtuse, concave on the upper side, with the subterminal spine visible from above. Antero-external process of the basal antennal joint short and stout, not reaching the middle of the fourth joint of the peduncle, and double pointed; acicle rounded, pubescent on the inner side, and not reaching the middle of the last joint of the peduncle. Chelipeds very unequal; merus of the larger one trigonal, with the angles rounded and the surfaces convex and marked with small, transverse, granulated, pubes- cent rugs; carpus oblong, distally widened, the sides rounded and the upper surface very convex; surface covered with pubescent, granulated ruge similar to those on the merus, which become broken up, in many places, into separate granules; hand oblong, nearly twice as long as wide, the upper surface evenly convex and granulated; fingers short and stout, the dactyl with a row of granules on the upper side parallel to the outer edge. Smaller cheliped with the merus and carpus compressed and gran- ulo-scabrous; hand small, narrow, with the sides rounded and having no prominent angles; palm more or less inflated, with often a trace of a groove near the proximal end of the upper surface; surface of the hand granulated; fingers pubescent with corneous edges. Ambulatory legs granulo-scabrous, but not spinous, except on the lower side of the dactyls and the distal end of the propodi; dactyls nearly straight, almost as long as the propodi, and not twisted. Alaska! Behring Is.! Puget Sound; Oregon; northern to southern California! Common on the shore at low tide. Pagurus Samuelis (Stimpson). Hupagurus Samuelis Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 86; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 482; Proc. Acad Nat. Sci. Phila., 1858, p. 250; Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 90, Pl. I, fig. 8. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. Vi, 1892, p,. 301 ,/Pl) Xit, fig: 12. Closely allied to hirsutiusculus. Body and legs quite strongly pubes- cent. Anterior portion of the carapace considerably longer than wide. Median tooth of the front quite prominent and sharp; lateral teeth repre- . sented only by a slight convexity in the margin. Ocular peduncles mod- erately stout, somewhat constricted in the middle, and about one-half as long as the anterior portion of the carapace. Ocular scales subovate and dorsally concave, with the subterminal spine visible from above. Anten- nal peduncle exceeding the eyes by fully one-half the length of the last joint; acicle rounded above, pubescent on the inner margin, not reaching the middle of the last joint of the peduncle, but nearly reaching the tip of 7 q } | CALIFORNIA STALK-EYED CRUSTACEA. | 145 the eye-stalk. Chelipeds very unequal, the larger one elongated, granu- lated, nearly devoid of pubescence, but furnished with minute sete which arise from the base of the granules; merus compressed, the angles rounded, length about one-half greater than the height; outer surface convex and more or less granulo-rugose toward the distal end; carpus in the adult a little over one-half longer than wide, the upper surface evenly convex and rounded off to the sides, the outer of which is granulated and more or less flattened; inner side somewhat more strongly granulated and slightly concave; on the posterior portion of the carpus there are several trans- yerse, granulated ruge which become gradually replaced anteriorly by isolated, rounded granules; hand oblong (but not twice as long as wide), broader than the carpus, becoming more elongated with age; upper sur- face evenly convex and granulated like the carpus, the granules becoming longer on the pollex; outer margin acute, granulo-deuticulate, the gran- ules becoming less sharp with age; fingers shorter than the palm; outer margin of the dactyl granulo-denticulate; a granulated line on the upper surface parallel to the margin; tips of the fingers corneous in young, but calcareous in old specimens. Smaller cheliped pubescent, not reaching the base of the dactylin adults, aud often scarcely reaching the base of the hand; merus spinulous below; carpus compressed, the upper edge armed with nine or ten sharp spines external to which is a parallel row of smaller spines; the granulations on the lower surface become more or less spiny towards the anterior margin; hand granulated, narrow, oblong, not shorter than the merus; palm rounded, having no prominent ridges or angles; fingers corneous tipped. Ambulatory legs pubescent, the propodi granulo-spinose above; dactyls rather stout, spinous below, and markedly shorter than the propodi. The dactyls are of a bluish color with a longi- tudinal reddish stripe on the sides; distal end of the propodi bluish. Patrick’s Point! Mendocino County! Pescadero! Mon- terey (St.)! Santa Catalina Is.! San Pedro! San Diego! Japan (Stimpson, Ortmann). I very strongly doubt that the #. Samuelis reported from Japan by Stimpson and Ortmann is the same as our California species. . Pagurus minimus, sp. nov. Anterior part of the carapace about as wide as long; median projection | of the front triangular, acute, the lateral ones rounded. LEye-scales pointed, with a very prominent subterminal spine. Ocular peduncles stout, cylindrical, a little flattened distally, and about two-thirds the length of the anterior portion of the carapace, reaching about the middle of the 10 146 CALIFORNIA ACADEMY OF SCIENCES. last joint of the antennal peduncle. Acicle short, slender, nearly reaching the tip of the cornex, but not reaching the middle of the last joint of the peduncle. Chelipeds pubescent; merus of the larger one compressed, the angles rounded; carpus one-half longer than broad, distally widened, the upper surface rounded, not sharply marked off from the lateral faces, and armed with short spines which are inclined forwards; inner face convex, but less so than the outer, which is evenly rounded; hand oblong, widen- ing distally to a short distance beyond the base of the dactyl; the base of the hand is armed with spines, which are inclined forwards like those of the carpus, and is strongly convex longitudinally, giving the hand the appearance of being bent downwards; pollex broad, the outer edge evenly rounded, sharp, upturned, and armed with anteriorly directed spines, the upper surface smooth and concave; dactyl broad, widest a little beyond its articulation, the outer margin sharp, spiny, evenly curved, the upper _ surface nearly smooth and concave; inner margin of both fingers furnished _with large, white, tubercular teeth; lower side of the hand evenly convex from side to side, the distal two-thirds longitudinally nearly straight, tapering evenly to the tip of the pollex. Hand of small cheliped narrow, rounded, the upper face oblique; fingers longer than the palm. Ambula- . tory legs rather slender and pubescent; dactyls slender, curved, tapering from the base, spiny below, and longer than the propodi. | | General color reddish, with spots of darker red; larger cheliped a darker red than the rest of the body, especially at the distal end; ocular peduncles with a median, transverse, light-colored band. Length, three-eighths of an inch. Described from a single female specimen dredged in 30 fathoms off San Diego, February 11, 1899. The Specimen carried numerous pale green eggs. This species is best distinguished by the character of the larger hand, which is convex at the narrow base, both longitudinally and transversely, while the distal portion is nearly smooth, longitudinally straight, but transversely concave. Collection University of Cali- fornia. Pagurus granosimanus (St.) Hupagurus granosimanus Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 90. Smitu, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 211. Anterior portion of the carapace a little longer than wide. Median tooth of the front broad and rounded; lateral teeth represented only by slight convexities of the margin. Ocular peduncles moderately stout, and ! nts sy a % 4 . “ , ot ee! ee ee ee te ~~ f oe ee ee ee we oS ) Piha yaaa f22 DIM C4 CALIFORNIA STALK-EYED CRUSTACEA. 147 about two-thirds the length of the anterior portion of the carapace; cornea “transverse and scarcely dilated. Eye-scales subovate, more or less con- cave on the-dorsal surface, the subterminal spine visible from above. Antennal aeielé reaching but little beyond the eyes, exceeding them by much less than half the length of the terminal joint; acicle not nearly reaching the cornex or the middle of the last joint of the peduncle. Chel- ipeds very unequal, the right one elongated in adults, devoid of pubes- cence, and strongly granulated; merus a little less than one-half longer than high, the anterior portion of both lower,edges denticulated; lower surface furnished with two rounded eeties outer surface convex and roughened as in Samuelis; carpus oblong, more than one-half longer than wide, but not twice as long; upper surface evenly convex and strongly granulated; sides flattened; upper edges rounded; hand oblong, nearly twice as long as wide, slightly wider and about one-sixth longer than the carpus, the upper surface evenly convex and granulated, the granules becoming larger on the pollex; outer margin acute, especially on the pollex, and granulo-dentate; dactyl nearly as long as the palm, the outer edge acute, granulo-dentate; tips of the fingers calcareous; smaller cheliped nearly devoid of pubescence; merus compressed; carpus rather stout, the upper margin sharp and armed with 5-7 spines; the upper part of the external surface is more or less spinulous, the lower portion granulo- rugose; hand oblong, wider than the carpus, and longer than the carpus or merus; the supero-external surface rather wide at the base and strongly granulated; supero-internal edge marked proximally by a ridge, external to which is usually a slight groove. Ambulatory legs spinous on the upper edges; dactyls spinous below and armed with smaller spines above, shorter than the propodi, not twisted, and armed with long, corneous claws. : Color in alcohol reddish; legs dark and marked with small bluish spots behind the small tufts of sete; granules bluish. Monterey (Stimpson)! San Pedro! San Diego! Pesca- dero! Patrick’s Point, Humboldt County, Calif.! Puget Sound! Pagurus Hemphillii (Benedict). Eupagurus Hemphillii Benepict, Proce. U. S. Nat. Mus., Vol. XV, 1892, Do 16. Closely allied to granosimanus and Samuelis. Carapace glossy, the ante- rior portion much longer than broad. Median tooth acute, the lateral teeth represented by a convexity in the margin. Eye-stalks rather slender and about two-thirds the length of the anterior portion of the carapace. Eye-scales subovate, more or less concave above, and subacute, the sub- terminal spine visible from above. The antennal peduncle somewhat K - convex, thickly granulated, with the edges rounded, there being no defi- 148 CALIFORNIA ACADEMY OF SCIENCES. exceeds the eye-stalk but not by one-half the length of the last joint; acicle small, much shorter than the eye-stalk, not reaching the posterior * third of the last joint of the peduncle. Chelipeds markedly unequal; merus strongly compressed, the outer surface quadralateral, nearly smooth, but marked more or less as in Samuelis; anterior portion of both lower margins denticulated; a small tubercle at the lower posterior angle; carpus oblong, deep, twice as long as wide, the sides flattened, the upper surface nite lateral margins; hand oblong, no wider than the carpus at the base, but widening distally, being broadest near the base of the dactyl; upper surface evenly convex and granulated like the carpus; outer edge acute; dactyl shorter than the palm. Merus of the left cheliped with fhe outer part of the lower margin spinulous; carpus with nine or ten spines on the sharp upper edge, external to which isa parallel row of much smaller spinules; hand much compressed, scarcely wider than the carpus, its broad, outer face nearly parallel with the inner one; a groove on the upper surface extending from the posterior margin nearly to the dactyl and mar- gined on either side by spiny granules. Ambulatory legs slender, much compressed, the carpus and propodus of the anterior pair (and especially _ the right member) spinulous on the upper surface; dactyls spinulous. below, scarcely as long as the propodi and not twisted. Color in alcohol, bright red, the tips of the dactyls light oalpneee ‘‘California’’ (Benedict); Humboldt County, Calif.! Pagurus tenuimanus (Dana). Bernhardus tenuimanus DANA, Proc. Acad. Nat. Sci. Phila., 1851, p. 2693. Crust. U. S. Expl. Expd,, Part I, 1852, p. 447, Pl. XXVII, fig. 7. Hupagurus tenuimanus Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 483; Proc. Acad. Nat. Sci. Phila., 1858, p. 237. Smuirn, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 211. Brnepicr, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 1. Waker, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 274. - Carapace smooth, naked. Median process but little more prominent. than the lateral teeth which end in a minute spine. Eye-scales narrowly oblong, subacute, channeled above, and having a minute subterminal spine not visible from above. First three joints of the antennal peduncle armed as in munitus; acicle rounded, the inner side pubescent, the tip. reaching as far as, or a little beyond the ocular peduncles; flagellum long. Chelipeds very unequal; merus of the larger cheliped short, trigonous, with the sides convex, the length and breadth subequal; surface more or less tuberculated, especially near the supero-distal angle, which is furnished with several marginal spines; extero-distal angle with a few spines; carpus. CALIFORNIA STALK-EYED CRUSTACEA. 149 stout, a little longer than wide, the upper surface strongly convex and thickly covered with small, sharp tubercles; inner edge spinous; hand very wide and strongly flattened, with the edges: produced into prominent, _ thin expansions, which make the hand much wider than the carpus; both surfaces thickly covered with small tubercles; inner edge of the hand convex, denticulated, and curved downward in the middle; fingers strongly compressed and hollowed out above, both sides minutely tuberculated or granulated. Left cheliped with the merus flattened, denticulated on the upper margin and at the infero-distal angle; carpus compressed, upper surface granulated or tuberculated, the inner edge acute and spinous; hand flattened, similar in shape to that of munitus, but the raised margins are denticulated; upper surface granulated or minutely tuberculated, the proximal margin overlapped by, instead of overlapping, the distal margin of the carpus; lower surface smooth; fingers more or_less pubescent. | Ambulatory legs) nearly naked, and marked with red spots; upper margins spinulous; dactyls slightly twisted, considerably longer than the propodi, the margins and anterior surface spinulous; sides of the preceding joints smooth and glossy. Length of large cheliped, 81 mm.; merus, 19 mm.; width of same, 16 mm.; carpus, 26 mm.; width of same, 24.5 mm.; hand, 34 mm., greatest width of same, 29 mm.; length of smaller cheliped, 51 mm. Puget Sound (Dana); Kadiak Is., Alaska! Pagurus californiensis (Benedict). Eupagurus californiensis BENEDICT, Proc. U. S. Nat. Mus., Vol. XV, 1892, p. 21. Faxon, Mem. Mus. Comp. Zool. Harvard College, Vol. XVIII, 1895, p. 55, Pl. XI, fig. 2-2e. Median tooth of the front low and scarcely in advance of the broadly triangular lateral teeth. Ocular peduncles moderately stout and about - three-fourths as long as the anterior portion of the carapace, which is nearly as wide as long. LEye-scales oblong, subacute, deeply channeled above, and curved downward, the subterminal spine small. Acicle rounded, pubescent on the inner margin, and not reaching the tips of the ocular peduncles. Chelipeds of exceptionally unequal size; larger cheliped glossy; merus trigonous, the surfaces convex and nearly smooth, but slightly roughened with scattered, minute prominences; carpus much widened at its distal end where its breadth about equals its length; upper surface convex and furnished with scattered, depressed granulations; outer © margin raised and finely granulated; inner margin prominent and armed with several short, unequal spines; outer surface more or less flattened; inner surface oblique and slightly concave; hand flattened, much wider - 150 CALIFORNIA ACADEMY OF SCIENCES. than the carpus, and nearly as broad as long; upper surface evenly convex and covered with sharp granulations of unequal size; outer margin acute, denticulated, and evenly,curved from the tip of the pollex to the base; inner margin produced, acute, more strongly denticulated than the outer, and ending in a prominent angle some distance from the base of the dactyl; fingers compressed; pollex very wide and furnished with several. large granulations; dactyl with the upper margin sharply granulated and ~ furnished witha granular line on the surface, between which and the outer margin the surface is deeply concave. Merus of the smaller cheliped compressed, narrow, and smooth, except on the lower side, which is spinu- lous toward the distal end; carpus compressed, armed above with tworows of spines, the included surface being slightly concave; hand small, oblong, with no prominent angles, a little longer than the carpus, but not exceed- ing the merus. Ambulatory legs glossy, nearly smooth; a spine at the supero-distal angle of the carpus projecting over the propodus; dactyls thin, pubescent, not twisted, and spinulous -below, exceeding the length of the propodi. ‘Color of alcoholic specimens near orange; the ambulatory legs crossed by bands of a lighter color. There isa band of lighter color near the middle of the ocular peduncles. , California (Benedict); Santa Catalina Island! Dredged in the harbor of Avalon. Pagurus munitus (Benedict). Hupagurus munitus BENEDIcT, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 19. Carapace smooth, nearly naked, the anterior margin with three nearly equal teeth. Eye-scales oblong, channeled above. First joint of the antennal peduncle with a spine or tooth at the interior antero-inferior angle; second joint with a spine at the antero-internal angle, the infero- distal produced forward into a prominent spine which is generally double pointed; third joint with a spine at the apex; acicle rounded, pubescent on the inner side, and about reaching the tips of the ocular peduncles. Chelipeds very unequal, merus of the right cheliped short, trigonous; ¢ar- pus broader than long, the margins produced into two large, thin, wing- like expansions and curved so that the inner surface is concave; hand smooth, much narrower than the distai end of the carpus, and about twice as long as wide, the edges subparallel; outer edge siightly concave; fingers punctate, shorter than the palm. In the smaller cheliped the car- pus is elongated and compressed, the upper margin spiny; hand oblong, smooth, much flattened, the acute edges upturned, making the upper sur- face markedly concave; proximal margin acute and overlapping the distal raw .wwr hs a es CALIFORNIA STALK-EYED CRUSTACEA. “on FB end of the carpus; fingers smooth, more or less pubescent below, the dactyl provided with a crest which is a continuation of the elevated mar- gin of the hand; the opposite margin is continued as a similar crest upon the pollex. Ambulatory legs slender, glossy, nearly naked, the upper margins of the last three joints more or less spinulous; dactyls slightly longer than the propodi. Alaska! Genus Paguristes Dana. Front with a distinct rostral projection. Ocular peduncles long and slender, the scales of moderate size. Antennules long. Antennal acicle robust, flagellum ciliated. Chelipeds equal or subequal, the fingers mov- ing in a horizontal plane. Fourth pair of pereopods not chelate. The first two abdominal segments in the male and the first segment in the female bear sexual appendages. Type.—P. longirostris DANA. Paguristes turgidus (St. ). Eupagurus turgidus Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 86. Clibanarius turgidus Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI; 1857, p. 484, Pl. XXI, fig. 1. Bate in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 278. WHuITEAVES, Can. Nat. (2), Vol. VIII, 1878, p. 471. Paguristes turgidus STIMPSON, Proc. Acad. Nat. Sci. Phila., 1858, p. 236; Ann. N. Y. Lyc. Nat. Hist., Vol. VII, 1860, p. 86. Smitu, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 211. Watker, Trans. Liverpool, Biol. Soc., Vol. XII, 1898, p. 275. Carapace pubescent, the anterior portion longer than wide. Front tri- dentate, the median tooth much the largest. Ocular peduncles over one- third the length of the anterior portion of the carapace. Antennal acicle armed with several dark-tipped spines on the inner margin; flagellum short. Chelipeds hairy above, the upper surface covered with dark-tipped spines; fingers corneous tipped. Ambulatory legs hairy, the dactyls a little longer than the propodi. Attains a length of about three inches. Puget Sound (Stimpson); Vancouver’s Is. (Smith); Farallon Islands! Santa Catalina Is.! Boetiires ee sp. nov. 26 Rostrum long, narrow and acute. Ocular scales cut into three spines: Ocular peduncles pubescent, enlarged at the base, and exceeding the peduncle of the antennz Acicle pubescent, reaching nearly to the tip of 152 CALIFORNIA ACADEMY OF SCIENCES. the peduncle, and having the outer margin armed with short spines; sec- ond joint of the peduncle produced, external to the acicle, into a process which is cut into three spines. Legs pubescent. Chelipeds equal, the merus nearly smooth; carpus with a flattened, subtriangular upper surface, the inner and outer margins of which are armed with three or four short, — stout spines; hands with the upper surface coarsely granulated; the short inner margin armed with three short, stout spines; a longitudinal intu- mescénce on the upper surface upon which are two rows of smali tubercles; fingers excavated within and having black, corneous tips. The upper margins of the carpi and propodi of the ambulatory legs are spinulous; dactyls scabrous, slender, nearly straight, and longer than the propodi. Length one-half inch. Three specimens taken at White’s Point near San Pedro, ‘Calif., July, 1895. I synly pe USNIM 52933 , Collection University of California. | Paguristes Bakeri, sp. nov. Carapace hairy, the anterior portion nearly as wide as long, eroded in front and towards the sides, and having a few short spinules near the antero-lateral margins; front with three, subequal, triangular teeth. Ocu- lar peduncles slender, cylindrical, two-thirds the length of the anterior portion of the carapace, and having a line of hairs on the upper side. Ocular scales armed with several marginal spines which become larger towards the tip. Peduncle of the antennzx nearly three-fourths the length of the eye-stalk; second basal joint with a spine at the antero-internal | ‘angle; the antero-external angle produced forward as a large, triangular, acute process which is armed with a few short spines on the outer margin. ~ Acicle rather small, awl-shaped, hairy, reaching a little beyond the middle of the last joint of the peduncle, the inner margin armed with spines and the outer margin furnished near the tip with two spines which point obliquely upwards. Chelipeds equal; merus trigonous, the upper ante- rior end hirsute and spiny and the two lower margins armed with short spines; carpus short, hirsute and spiny above, the inner margin of the upper side armed with about six spines which increase in size anteriorly; outer face small, hard portion of the lower side reduced to a narrow, trans- verse bar, inner face flattened, with a concavity on its lower side, and armed with a few spines above and anteriorly; hands short and broad, widest across the base of the pollex, the upper surface flattened, hairy, armed with numerous spines, inner edge straight, the outer very strongly curved, inner faces flattened and armed with small spines, four spines on the inner margin behind the base of the dactyl; pollex very broad at base; CALIFORNIA STALK-EYED CRUSTACEA. 153 lower surface of hand tumid below base of pollex and armed with short spines; an angle of 90° between its upper and inner faces. Ambulatory legs hairy; carpus of the anterior pair spiny above; propodus armed below with short spinules and above with seven or_eight rather strong spines; dactyls flattened, much longer than the propodi, both margins closely set with spines; second pair much less spiny than the first. There is a pair of very small, simple, two-jointed appendages on the first abdominal segment in the female, but the succeeding four segments have an appendage only on the left side; of these the first three (those on the second, third, and fourth segments) are well-developed and biramous, but the appendage on the fifth segment is smaller and reduced to a single branch. In the male the pair of appendages on the first abdominal seg- ment is larger than in the female and the inner faces are channeled; the second segment has a pair of appendages, and the third, fourth and fifth ' segments have a uniramous appendage only on the left side. General color dark reddish; legs more or less colored with blue. Leneth of female 2.5 in.; of males 4.5 in. and 5 in. Described from three specimens from San Diego, Calif.: one a female loaned by the San Diego Natural. History Society, and two large males. For the oppor- tunity of examining the latter I am indebted to Dr. Fred Baker of San Diego. This species is allied to P. fecundus Faxon, but is dis- tinguished by having a less prominent rostrum, much longer antennal flagellum, shorter dactyls on the sec- ond and third pereopods, and small but well-marked spines instead of ‘‘obscure teeth,’ on the terminal lobes of the telson. Holopagurus, gen. nov. Rostrum not prominent. Ocular peduncles moderately long and slender; basal scales close together. Antennal acicle rather short; flagellum hairy. Left cheliped larger than the right; fingers of both hands moving horizon- tally, their inner margins not excavated. Fourth pair of legs more or less chelate. First abdominal segment of the male devoid of appendages; sec- ond, third and fourth segments with an appendage only on the left side. Telson entire. Type-—H. pilosus. 154 CALIFORNIA ACADEMY OF SCIENCES. Holopagurus pilosus, sp. nov. Anterior portion of the carapace slightly wider than long, eroded at the front and sides, and having a few short spinules near the antero-lateral margin. Frontal margin transverse, rostral projection represented by a small prominence which does not exceed the lateral ones. Ocular pedun- eles cylindrical, rather slender, and about two-thirds the length of the anterior portion of the carapace; cornez not dilated; basal scales very - wide at base, their inner sides produced forwards into a subacute, triangu- lar lobe. Antennular peduncle about equalling the eye-stalks. Peduncle of the antenne shorter than the eye-stalks, the second joint with a short spine at the antero-external angle but none at the antero-internal angle; last joint of the peduncle short, less than twice as long as wide; acicle straight, ciliated, evenly tapering to an acute tip, convex and roughened above, and not reaching the tip of the peduncle; flagellum shorter than the carapace and thickly set below with long hairs. Chelipeds of similar form, furnished with long hairs and short spines; merus trigonous, the lower side flattened, the upper edge sharp; carpus armed with short, scat- tered spines on the upper surface which is convex, rounding off insensibly to the small outer face; inner face flattened and separated from the upper surface by a rather sharp edge which is armed with sharp spines and turns outward near the base-more sharply on the left cheliped than on the right; hands horizontally flattened, the upper surface gently convex and covered with scattered spines which are larger and more thickly set near the rounded margins; the larger hand is widest across the base of the fingers, beyond which it is quite abruptly contracted; the pollex is rather - narrow a Short distance beyond the base, its inner and outer margins rounded, the tip corneous; dactyl subconical, the tip corneous and strongly curved, its surface thickly set, like the pollex, with short, corneous spines. The smaller hand is relatively narrower than the larger one, the inner and outer faces parallel; pollex not so much contracted beyond the base as in the larger hand, its outer margin evenly convex near the base and not slightly concave like that of its fellow. Ambulatory legs hairy and armed with spines on the upper sides of the merus, carpus and propodus; _propodi subcylindrical, nearly smooth below; dactyls long, gently curved, flattened towards the tip, subcylindrical at the base, about twice the length of the upper side of the propodi, strongly twisted, and armed only with very short, corneous spinules. Dactyl of the fourth pair of legs extending for about one-third its length beyond the propodus. Coxe of the last pair of legs separated by an oblong process of the sternum on the posterior side of which is a sharp, median groove. Abdominal append- ages long and hairy. The posterior portion of the carapace is more or less calcified, especially over the cardiac area, and the branchial regions are reticulated with mem- branous lines. CALIFORNIA STALK-EYED CRUSTACEA. 155 General color yellowish white; the antenns are blue and the anterior portion of the carapace has more or less of the same coloration; a peculiar reddish coloration occurs on different parts of the body, notably on either side of the cardiac area of the carapace. Length four inches; large cheliped three-eights of an inch longer than the smaller one. One large male specimen and several small ones dredged in about 25 fathoms off San Diego, Calif. Col- lection University of California. Family PARAPAGURIDE. Similar to the Paguride but having trichobranchie, instead of phyllobranchie. Deep sea forms. Genus Parapagurus Smith. Resembles Pagurus. Third maxillipeds widely separated at the base. Chelipeds very unequal. Sternum between the first and second pereopods narrow. Eleven pairs of branchizx, two at the base of the maxillipeds and each of the first three pereopods, and three at the base of the fourth. Well developed and symmetrical pairs of appendages on the first and sec- ond abdominal segments in the male. Type.—P. pilosimanus SMITH. Parapagurus Mertensii (Brandt). Pagurus Mertensii BRANDT, Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 112. Eupagurus Mertensit Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 483; Proc. Acad: Nat. Sci. Phila., 1858, p. 237. Brnepict, Proc. U.S. Nat. Mus., Vol. XV, 1892, p. 2. Anterior portion of the carapace sparsely hairy and about as wide as long. Median frontal tooth prominent, acute, the lateral teeth small. Ocular scales ovate, acute, concave above. LEye-stalks short, about one- half the length of the anterior portion of the carapace. Acicle slender, rounded, sparsely ciliated, curved downwards, much exceeding the eye- stalks, but not reaching the tip of the antennal peduncle. Chelipeds pubescent, spiny, very unequal; right cheliped: very large; merus armed with a row of short spines on the inner and outer margins of the lower “i 156 CALIFORNIA ACADEMY OF SCIENCES. side; carpus long, the inner and outer margins spiny, and the convex upper surface armed with two rows of short spines; outer face more or less flattened; lower side bulging and armed with a few short spines; hand long, narrow, dorso-yentrally flattened, and bent inward at a slight angle to the carpus; the rounded upper surface furnished with small, sub- serially arranged granules which become more prominent on the fingers; ‘inner and outer margins sharp, granulo-denticulate and parallel; lower surface of the palm with a broad, shallow concavity which extends nearly to the dactyl. Left cheliped slender; merus spiny below; carpus subcy- — lindrical, with three rows of spines on the upper surface; hand narrow, much longer than the carpus; palm very short; fingers long, narrow, and curved downwards. Ambulatory legs long, compressed, and very slender; dactyls exceedingly long and slender and armed below with numerous spines. Described from a single female specimen obtained from the U. S. National Museum from Mr. Benedict, who pointed out’ that it was not a true Pagurus but belonged among the trichobranchiate forms. Northern California, Nitkasund, Kadiak Is., Atcha, Kamtschatka (Brandt); lon. 33° 55’ 30” N.; lat. 119° 53° 30" W.; 19 fathoms! ; This species affords one of those curious cases of com- mensalism with colonies of hydroids sometimes found among deep sea pagurids. The colony of hydroids cov- ering the shell in which the crab lives forms a mem- branous growth which extends beyond the boundary of the shell and, in course of time, according to Mr. Bene- dict, causes the shell to disappear, leaving its inhabitant “with a membranous, instead of a calcareous domicile. In the specimen which I have, the covering is partly formed by the tip of a broken shell, but mainly by an extension of the membranous growth formed by the colony of hydroids. This arrangement certainly affords the crab the advantage of allowing it to grow, without — its having to undergo the troublesome experience of changing its lodgings. 5 ) rw €@ Sa 7 ‘+ © 2 é , e bet cat = s ae ea ee ‘al CALIFORNIA STALK-EYED CRUSTACEA. 157 Subtribe THALASSINIDEA. Carapace short, compressed, and marked with two longitudinal sutures. Rostrum small or wanting. Both pairs of antenne elongated and furnished . with long peduncles, those of the outer pair five-jointed and usually devoid of an acicle. First pereopods more or less chelate; second pair often chelate; third pair always simple. Last segment of the thorax movable. Abdomen long, the segments not overlapping, the side plates feebly de- veloped. Tail-fin well developed. Branchiz variable. Family CALLIANASSID.. Carapace strongly compressed. Rostrum flattened. Eyes small. First pair of chelipeds generally unequal; second pair with or without chele. Side plates of the abdomen absent. Telsonand uropods broad and devoid of cross furrows. Burrowing crustaceans with soft, elongated bodies. Genus Upogebia Leach. Rostrum short and tridentate. Eyes small, with cylindrical peduncles. No antennal scale. External mavxillipeds pediform. First thoracic legs subequal and subchelate, the pollex small; the remaining pairs not chel- ate. Abdomen long, with subquadrate joints; pleopods broad, the second pair similar to the others; uropods broad and foliaceous; telson broad, subquadrate, foliaceous. Podobranchiw and mastigobranchie wanting. Type.—U. stellata (MONTAGU). Upogebia pugettensis (Dana). Gebia pugettensis Dana, Proc. Acad. Nat. Sci. Phila., 1852, p. 19; Crust. U. S. Expl. Expd., Part I, 1852, p. 510, Pl. XXXII, fig. 1. Srirmpson, > Journ. Bost. Soc Nat. Hist., Vol. VI, 1857, p. 488, Pl. XXI, fig. 2. LockineTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 299. Gebia californica Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88. Upper portion of the carapace in front of the cervical groove flattened, scabrous, and hairy, and marked with three longitudinal grooves, the median groove being the shortest; front with the median tooth large, horizontal, triangular, the lateral teeth short. A minute marginal tooth or spine-generally present a short distance below the lateral teeth. Antero- lateral and postero-lateral angles rounded. Eye-stalks short, reaching very little further forward than the lateral teeth of the front. Antennules less 158 CALIFORNIA ACADEMY OF SCIENCES. than one-half the length of the antenna, the flagella subequal. Peduncle | of the antennz exceeding that of the antennules; flagellum ciliated and longer than the carapace. Chelipeds equal; merus spinulous and hairy below; carpus with the upper and lower edges spinulous, and having a . spine at the antero-internal angle and another spinea short distance below it; hand with two parallel, scabrous, hairy lines on the upper edge and a transverse granulated line at the proximal end of the inner surface; a line | of hairs on the outer surface which is continued obliquely across the carpus; lower side of the palm scabrous and hairy; pollex short, bent downwards, and having a tooth near the middle; dactyl obliquely com- pressed, incurved, acute, projecting, when closed, far beyond the tip of the pollex. Uropods short, the inner ramus distally rounded, truncated, the outer rounded. Telson entire, rectangular, wider than long. Length, 112 mm. : Queen Charlotte Is. (Smith); Vancouver’s Is. (Bate); Puget Sound (Dana); Bodega Bay! San Francisco Bay! Monterey! Santa Catalina Is.! San Pedro! San Quentin Bay, Lower California (Lockington). This species can generally be found wherever muddy beaches occur. It inhabits holes of one or more feet in depth, many of which are left uncovered at low tide, when the creatures can easily be dug out. Specimens are often taken, however, at a depth of several fathoms. There is a peculiar parasitic crustacean, Phyllodurus abdominalis St., that is often found upon this species, being usually attached to the base of the abdominal appendages. Sometimes also a small bivalve mollusc, Pythina rugifera, is found attached to this crustacean by its byssus. The tooth on the pollex is sometimes absent, as it was in the specimen described by Dana. ‘‘ This tooth,” says Stimpson, ‘‘is a prominent character in all the very numerous specimens in the Smithsonian Museum, but it is obsolete in the specimen described by Dana, although actual comparison shows them to be the same.” In some small specimens from Catalina Island the small CALIFORNIA STALK-EYED CRUSTACEA. 159 marginal spine beneath the lateral teeth of the front was absent, although they agreed with specimens from . northern California in every other essential feature. Genus Callianassa Leach. Rostrum absent or reduced to a small point. Eye-peduncles flattened; cornea median, small or absent. Antennular flagella never shorter than the preceding joint. No-antennal scale. External maxillipeds operculi- form. First pair of pereopods very unequal and furnished with well- developed chelw#; second pair small and chelate; third pair with the penultimate joint broadly expanded; fifth pair subchelate. Second pair of abdominal appendages in the female slender, the succeeding ones broad, foliaceous, and fringed with ciliated hairs. Caudal appendages wide. Gills with flattened filaments. -In this genus, according to Bate, only arthrobranchiz are present, with the exception of a rudimentary masti- gobranchia on the third maxilliped. . Callianassa californiensis Dana. a i 27 Callianassa californiensis DANA, Proc. Acad. Nat. Sci. Phila., 1854, p. 175. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 489, PI. XXI, fig. 4. A. Minnr-Epwaxps, Nouv. Archiv. Hist. Nat. Paris, Tome VI, 1870, p. 82. LocxineTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 301. Callianassa occidentalis Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88. Front very short and rounded, with a small, triangular tooth on either side between the bases of the ocular peduncles and the antennz. Ocular peduncles subtriangular, approximated at the bases, but diverging towards the acute tips which are somewhat upturned. Antennulary flageila sub- equal. Antenne from one-half to two-thirds the length of the body, the peduncle nearly equalling that of the antennules. Chelipeds in the adult male with one arm enormously developed; ischium of the larger cheliped . slender, compressed, incurved, distally widened, and finely denticulated on the acute lower margin; merus about as long as the ischium (generally a trifle longer) but stout, curved, smooth, almost naked except on the cili- ated margins, and furnished with a prominent lobe at its infero-proximal angle; carpus very wide, a little longer than the merus, the outer surface smooth. naked, glossy, and evenly convex as if forming a part of the sur- face of a cylinder; margins acute, ciliated, and minutely serrulated; upper \ ¢ ~S 160 CALIFORNIA ACADEMY OF SCIENCES. margin produced into a thin expansion overhanging the smooth inner surface; lower margin produced in a similar manner but not nearly so | strongly; posterior margin below the articulation of the merus produced backwards into a broadly and evenly rounded lobe; hand narrower and a little shorter than the carpus, broadest at the base, with the ciliated mar- gins more or less incurved towards the proximal end; a prominent hiatus between the fingers; a prominent upturned tooth between the bases of the fingers; fingers a littie longer than the palm and furnished with tufts of — cilia; dactyl longer than the pollex, minutely denticulated on the inner margin, and bent abruptly near the tip, forming a sharp hook. In the ‘smailer cheliped the ischium resembles that of the larger one but is scarcely denticuiated below; merus widest near the middle, shorter than the ischium, and devoid of the lobe at the base, but having an obsolescent tooth near the middle of the lower margin; carpus very long and narrow, with both surfaces convex; hand narrow, relatively much longer than the hand of the larger cheliped, but shorter than the carpus; palm oblong, the margins parallel, the fingers not gaping; dactyl longer than the poilex and not hooked at the tip. The smaller cheliped in the female is like that of the male but the larger cheliped is relatively smaller and of a different form; the ischium is like that of the smaller cheliped and the merus much the same but stouter and furnished with a more prominent inferior tooth; the carpus is similar in shape to that of the other cheliped but relatively shorter, scarcely one-half longer than broad, and but little longer than the merus; both surfaces convex, the margins not produced as in the male; ‘ @ hand as wide as the carpus, palm longer than broad, with the margins subparallel; fingers a little shorter than the palm; dactyl a little dilated near the base and curved at the tip. Ischium and merus of the ambula- tory legs fringed with long hairs below; hand ovate, ciliated, much com- pressed, fingers longer than the palm. Caudal appendages distally trun- cated. Telson scarcely shorter than the uropods and furnished with a rounded emargination at the tip. Length of specimen, male, 61 mm.; length of larger cheliped, 50 mm.; of small cheliped, 32 mm.; width of carpus of large cheliped, 13 mm.; leneth of same, 155 mm.; length of hand to tip of dactyi, 10 mm.; of merus, 12 mm.; of ischium, 10.5 mm.. Length of larger cheliped of a female, 31 mm.; of smaller cheliped, 28 mm. California (Dana); Mutiny Bay, Alaska (Lockington); Puget Sound (Stimpson); Shoalwater Bay (Cooper); Bodega Bay! San Francisco Bay! Tomales Bay! sig Barbara! San Pedro! 2 ae “oe ~~ 7 CALIFORNIA STALK-EYED CRUSTACEA. 161 Callianassa longimana St. Me, £2 Callianassa longimana Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 86; Journ. Bost. Soc, Nat. Hist., Vol. VI, 1857, p. 490, Pl. XXI, fig. 5. Cooprer, Rep. Expl. and Sur. to Pacific Ocean, Vol. XII, Book 2, 1860, p. 388. Bats, in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 279; Challenger Reports, Vol. XXIV, 1888, p. 49. A. Mitnge-Epwarps, Nouv. Archiv. Hist. Nat. Paris, Tome VI, 1870, p. 83. Lockineton, Ann. Nat. Hist. (5), Vol. II, 1878, p. 302. Rostrum small but subacute, with two small, lateral teeth as in californi- ensis. Ocular peduncles and antennules much as in the preceding species. Antenne about one-half as long as the body. The chelipeds in the adult male are narrower than in californiensis; the ischium is of similar shape, with the lower margin finely denticulated and the outer surface more or less scabrous; merus similar to that of californiensis, but the upper margin is not convex and the lobe at the base is more prominent; carpus oblong, with the margins subparallel, ciliated, thin, but not produced nearly so much as in californiensis; outer surface smooth, glossy, and very convex; hand oblong, longer than the carpus, with thin, ciliated, parallel margins and a smooth, convex, glossy, outer surface; fingers shorter than the palm and furnished with tufts of cilia and not widely gaping when closed; the dactyl is dilated near the base, hooked at the tip, and projects but little beyond the pollex; upper edge and inner surface more or less granulated. The smaller cheliped resembles that of californiensis; merus widest near the middle, shorter than the ischium, often having a small tooth near the middle of the lower margin; carpus narrow, much longer than the ischium, convex and glossy on both sides, the margins parallel; hand of the same width as the carpus but scarcely as long; palm longer than broad; fingers long, ciliated and contiguous. The larger cheliped in the female is scarcely distinguishable from that of the female of californiensis. The smaller chelipeds are also very similar in the two species, but in longimana the hand and carpus are somewhat narrower. Ambulatory legs as in califor- niensis. 'Telson with an obtuse emargination at the tip. Uropods distally truncated and slightly exceeding the telson. Color whitish. Length of a male specimen, 39 mm.; length of large cheliped, 39 mm.; ischium, 8 mm.; merus, 8.5 mm.; carpus, 10.5 mm.; hand, 13.5 mm.; length of small cheliped, 22.6 mm.; carpus, 6.5 mm.; width of same, 2.1 mm.; length of merus, 4.2 mm.; hand, 6.2 mm. Vancouver’s Island (Bate); Shoalwater Bay (Cooper); Puget Sound (Stimpson); San Francisco eeenh San Pedro! 11 , G2 CALIFORNIA ACADEMY OF SCIENCES. San Diego! Santa Barbara Island! Santa Catalina Is.! San Quentin Bay, Lower California (Lockington). It is remarkable that a species which resembles cali- forniensis in almost all other respects should differ from it so markedly in the chelipeds of the adult male. The females are very much alike, but they can be distin- guished by the differences in the rostrum and the rela- tive stoutness of the smaller chelipeds. Callianassa gigas Dana. Callianassa gigas Dana, Proc. Acad. Nat. Sci. Phila., 1852, p. 19; Crust. U.S. Expl. Expd., Part I, 1852, p. 512, Pl. XXXII, fig: 3. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857,.p. 489, Pl. XXI, fig. 3. A. MILNE-EDwarps, Nouv. Arch. Hist. Nat. Paris, Tome VI, 1870, p. $l. Looxineton, Ann. Nat. Hist. (5), Vol. II, 1878, p. 302. Front more or less triangular. Large hand strongly compressed and smooth; fingers similar, shorter than one-half the length of the hand and not gaping, the dactyl arcuate, acute; carpus more than one-half the length of the hand; merus narrow, toothed below at the base. Telson but little shorter than the uropods. Length 43 inches. Puget Sound (Dana). Callianassa affinis, sp. nov. Of tr, 6: 29 3p Male: Front obscurely tridentaté. cular peduncles oblong, subacute, the inner margins not diverging toward the tip; cornea in front of the middle of the peduncle. Antennulary flagella subequal. Antennz about one-half the length of the body. Ischium of the larger cheliped dentate below; merus stout, with a prominent lobe on the under side near the base, the lower side of which is denticulated; carpus short, with the pos- tero-inferior angle broadly rounded and the margin not produced as it is in californiensis; hand fully twice the length of the carpus; palm oblong, both inner and outer faces convex; dactyl longer than the pollex, the extremity hooked and the inner margin furnished with a few stout teeth. Smaller cheliped slender; merus widest at the middle; carpus narrow, as long as the hand; fingers pubescent. First pair of ambulatory legs cili- ated below. Telson broadly rounded and shorter than the uropods. Point Loma, Calif.! San Clemente Is.! July, 1895. Collection University of California. CALIFORNIA STALK-EYED CRUSTACEA. 168 Subtribe HOMARIDEA. Carapace not strongly compressed and furnished with a more or less flattened rostrum. Antennal acicle present. First three pairs of pereo- pods chelate, the first pair large; last five pairs of pereopods with seven joints. Abdomen rather wide, the segments overlapping and laterally produced. Tail-fin well developed. Family ASTACID/E. Carapace subcylindrical; rostrum well developed. Fifth thoracic seg- ment movable. First abdominal segment in the male (and generally also in the female) furnished with sexual appendages. Telson and the outer branch of the uropods divided by a cross furrow. Terrestrial or fresh water species confined to the northern hemisphere. Genus Astacus Fabr. This genus is distinguished from Cambarus chiefly by the possession of a pleurobranchia on the last thoracic segment. The Astacidz west of the Rocky Mountains belong, without exception, to the genus Astacus, while the numerous species inhabiting the Mississippi valley and the eastern states belong to the closely allied genus Cambarus. Astacus klamathensis St. Astacus klamathensis Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 87; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 494. BatTE in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 278. Hagen, Ill. Cat. Mus. Comp. Zool. Harvard Coll., No. 3, 1870, p. 93, Pl. ITI, fig. 169. LocxineTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 303. Faxon, Proc. Amer, Acad. Arts and Sci., Vol. XX, 1884, p. 151; Mem. Mus. Comp. Zool. Harvard Coll., Vol. X, No. 4, 1885, p. 131, Pl. VI, figs..1 and 2; Proc. U.S. Nat. Mus., Vol. VIII, 1885, p. 356; Jbid.,, Vol.. XII, 1890, p. 634; Jbid., Vol. XX, 1898, p. 665. A rather small species with ‘a smooth, punctated carapace. Rostrum not so long as in J'rowbridgii, with the sides converging anteriorly and the dorsal surface concave and marked with an obscure median ridge; term. inal tooth acuminate and shorter than in TVrowbridgii; lateral teeth acute or subacute and not produced into spines; a single spine on either side of 164 CALIFORNIA ACADEMY OF SCIENCES. the base of the rostrum, the posterior pair seen in Trowbridgii. being absent. Dorsal area about twice as long as its width across the middle. Chelipeds much like those of Trowbridgii but there is no spine at the antero-internai angle of the carpus. Antero-lateral angles of the abdomi- nal segments rounded; postero-lateral angles subacute. Length, 95 mm. Klamath Lake! Sikon Creek; Des Chutes River; Walla Walla, Wenos Valley, Spokane Falls, Washing- ton (Faxon); British Columbia in streams east of the Cascades (Bate); Eel River, Humboldt County, Calif.! Astacus Gambelii (Girard). Cambarus Gambelii GIRARD, Proc. Acad. Nat. Sci. Phila., 1852, p. 90; Ibid., 1853, p. 380. Astacus Gambelii AGassiz, Proc. Acad. Nat. Sci. Phila., 1853, p. 375. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 492. Hacen, Ill. Cat. Mus. Comp. Zool. Harvard Coll., No. 3, 1870, p. 90, Pls. I, III, and XI. lLocxk1neTon, Ann. Nat. Hist. (5), Vol. II, 1878, p. 303. Faxon, Proc. Amer. Acad. Arts and Sci., Vol. XX, 1884, p. 152; Mem. Mus. Comp. Zool. Harvard Coll., Vol. X, No. 4, 1885, p. 186; Proc. U.S. Nat. Mus.,-Vol. VIII, 1886, p. 356; /bid., Vol. XX, 1898, p. 666. Carapace obese, punctate. Rostrum short, acute, concave above, with the sides serrated and converging to the tip; a small spine on either side of the base behind which there may be a trace of a second pair. Dorsal area over twice as long as the width across the middle. Chelipeds large; merus much compressed, with the lower margins spinulous, the spine at the antero-internal angle small or absent; a spine on the upper side near the distal end; carpus devoid of spines, the upper edge with a longitud- inal median depression; hands large, broad, scabrous, the upper surface of the palm thickly pilose on either side of the middle; lower side of the palm inflated. Ogden River, Utah; Idaho; Mouth of Yellowstone River, Montana; Wyoming. This species has been reported from California, but its occurrence there is not entirely certain. 4 ie —————— ee 6s te Oa + CALIFORNIA STALK-EYED CRUSTACEA. 165 Astacus Trowbridgii S¢?. Astacus Trowbridgii Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p- 87; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 493. Coopnr, Rept. Expl. and Sur. Pacific Ocean, Vol. XII, Book 2, 1860, p. 388. Hagen, Ill. Cat. Mus. Comp. Zool. Harvard Coll., No. 3, 1870, p. 93, Pl. III, fig. 171, and Pl. X. Faxon, Proc. Amer. Acad. Arts and Sci., Vol. XX, 1884, p. 152; Mem. Mus. Comp. Zool. Harvard Coll., Vol. X, No. 4, 1885, p. 134; Proc. U.S. Nat. Mus., Vol. VIII, 1885. p. 356. Ibid., Vol. XX, 1898, p. 666. Carapace obese, punctate, nearly smooth, but roughened by minute prominences at the sides, and furnished with two pairs of spines at the base of the rostrum. Rostrum long, concave above, the sides very slightly converging anteriorly, each ending in an acute, upturned spine separated by a rounded notch from the long, slender, acuminate median spine which is also somewhat upturned. Antennules with a small spine on the lower side of the basal joint near the tip; peduncle shorter than the rostrum. Antenne shorter than the body, with a strong spine on the outer side of the outer basal joint; acicle acuminate, nearly equaling the rostrum. Epistome triangular. Chelipeds long, but shorter than the body; merus compressed and armed below with a double series of spines, the outer series containing the less number; a strong spine at the infero-distal angle; a spine at the anterior end of the outer surface and another on the upper surface a little behind the anterior end; aspine at the antero-internal angle of the carpus and another on the anterior margin of the lower side; . hands large, rough and punctate, with minute set in the punctures; fin- gers narrow, longer than the palm, irregularly dentated within, and spinu- lous toward the distal end of the outer margins; inner margin of the dactyl convex, the outer slightly concave. The lateral wings of the abdominal segments have the antero-distal angles rounded and the postero-distal angles subacute. Telson longer than broad, the terminal division nearly semicircular. Cardiac area wide. Length of body, 141 mm.; of cheliped, 106 mm.; of hand, 54 mm. Columbia River, Oregon! Streams emptying into Shoalwater Bay (Cooper); Monterey (Faxon). The speci- men from the latter locality was a large female ‘‘ said to have been taken from a bunch of seaweed in salt water.’’ ~ 166 CALIFORNIA ACADEMY OF SCIENCES. Astacus leniusculus Dana. Astacus leniusculus DANA, Proc. Acad. Nat. Sci. Phila., 1852, p. 20; Crust. U.S. Expl. Expd., Part I, 1852, p. 524, Pl. XXXIITI, fig. 1. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 493. Hagen, Ill. Cat. Mus. Comp. Zool.. Harvard Coll., No. 3, 1870, p. 94. LocxkinerTon, Ann, Nat. Hist. (5), Vol. II, 1878, p. 304. Faxon, Proc. Amer. Acad. Arts and Sci., Vol. XX, 1884, p. 151; Mem. Mus. Comp. Zool. Harvard Coll., Vol. X, No. 4, 1885, p. 132, Pl. VI, fig. 4; Proc. U.S. Nat. Mus., Vol. VIII, 1886, p. 356. Jbid., Vol. XX, 1898, p. 666. Closely allied to Trowbridgii. Carapace punctate. Rostrum trispi- nose, the median spine long and slender; two pairs of spines at the base. Cardiac area rather wide. Carpus of the chelipeds unarmed with the exception of the antero-internal angle; merus denticulated and armed with a dorsal spine. Length, 4 inches. Columbia River (Dana); San Francisco County, Calif. (Faxon). Astacus nigrescens St. Astacus nigrescens StIMpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 87; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 492. Haaen, Til. Cat. Mus. Comp, Zool. Harvard Coll., No. 3, 1870, p. 92, Pl. III. Lockineton, Ann. Nat. Hist. (5), Vol. II, 1878, p. 303. Huxtey, The Crayfish, p. 244, fig.62. Faxon, Proc. Amer. Acad. Arts and Sci., Vol. XX, 1884, p. 152; Mem. Mus. Comp. Zool. Harvard Coli., Vol. X, No. 4, 1885, p. 185; Proc. U. S. Nat. Mus., Vol. VIII, 1886, p. 356; Ibid., Vol. XII, 1890, p. 634. ‘Margins of the rostrum nearly parallel, denticulated with five or six small sharp spines on either side; the two anterior thoracic spines rather long. Dorsal area between the branchial regions as wide as in A. Gambelliz, from which this species differs in its smaller and more slender hands, which _arealso without pubescence. The lateral angles of the abdominal segments are sharp, and the caudal segment has two slender spines on each side. Color, blackish. Length three inches.” Unalaska, Alaska; Fort Steilacoom, Washington; San Joaquin Slough, Coyote Creek, Santa Clara County; Alameda Creek, Alameda County, California (Faxon). CALIFORNIA STALK-EYED CRUSTACEA. 167 Astacus oreganus Randall. Astacus oreganus RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIII, 1839, p. 138, Pl. VII. DrKay, Zoology of New York, Part 6, Crustacea, 1844, p. 238. Erzicuson, Wiegmann’s Archiy. f. Naturg, (1), Bd. XII, 1846, p. 375. Stimpson, Journ. Bost. Soc. Nat. Hist.; Vol. VI, 1857, p. 495. HaGeEn, Ill. Cat. Mus. Comp. Zool. Harvard Coll., No. 3, 1870, p. 95. Faxon, Mem. Mus. Comp. Zool. Harvard Coll., Vol. X, No. 4, 1885, p. 133. Randall’s description of this species is as follows:— ‘*Body fuscous, granulated, carpus with a sharp spine at the anterior angle; arm produced into a spine on each side anteriorly; thorax behind the front with five spines placed three before and one on each side behind the lateral ones; a large reddish spot on each side posteriorly; front little reflected on the sides, terminating in a long, slender spine and having a short, marginal spine on each side. Lon. 4”. Testa granulata, bimacu- lata, fronte valde producta.”’ | Columbia River. Randall’s specimen of this species has, unfortunately, been lost and nothing answering to his description has since been found. If the species belongs to the genus Astacus it differs from all the Astacz that are now known in the possession of a median thoracic spine. The fig- ure accompanying Randall’s description is so obviously inaccurate that it has no scientific value whatever. Dr. Hagen suggests that oreganus may be the same as lenz- usculus, as both species come from the same locality. Faxon considers the species indeterminate. The species described by Randall as Nephrops occi- dentalis was doubtless wrongly reported, through an exchange of labels, from Western America, instead of the Sandwich Islands. Ortmann, who puts the species in the genus Lnoplometopus unites with it H. pictus A. Milne-Edwards.' 1See Kingsley, Bull. Essex Inst., Vol. XIV, 1883, p. 131, Pl. II, fig: 1, and Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. X, 1897, p. 274. 168 CALIFORNIA ACADEMY OF SCIENCES. Subtribe LORICATA. / Body not strongly compressed and often depressed. Rostrum broad, short or absent. Outer antennez devoid of an acicle, the first joint of the peduncle fused with the epistome. All of the pereopods six-jointed, and none of them chelate except the last pair which is sometimes subchelate in the female. First abdominal segment devoid of appendages in both sexes. Posterior portion of the telson and uropods flexible. Gills tri- chobranchiate. Genus Panulirus White. Carapace spiny. Ocular peduncles small, free. Upper portion of the antennular segment wide, nearly horizontal, fused with the carapace, the surface furnished with spines, the sides smooth where they are rubbed by the bases of the widely separated antenne. Flagella of the antennules long. Rostrum absent. A pair of spines on the anterior margin of the carapace over the eyes. Epistome devoid of a longitudinal furrow. Fifth pereopods of the female subchelate. Pfeffer discards the name Panulirus on account of its similarity to Palinurus, the former name being derived from the latter by simply transposing the position of some of the letters. This does not appear to me a suffi- cient reason for discarding a generic name. Panulirus interruptus (Randall). Palinurus interruptus RANDALL, Journ. Acad. Nat. Sci. Phila., Vol. VIZI, 1839, p. 187. GiBBEs, Proc. Amer. Ass. Adv. Sci., 1850, p. 194. Panulirus interruptus Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 88; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 491. Ratupoun, R., The Fisheries of the U. S., Sec. 1, 1884, p. 780, Pl. 320. Ortmann, Zool. Jahrb. Abth. f. Syst., Bd. X, 1897, p. 260. Senex interruptus ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. VI, 1892, pe: | Carapace subcylindrical and covered with short spines. Rostral horns subparallel, compressed, and curved forwards. Upper portion of the antennulary segment with two pairs of spines, the margins raised and polished. The peduncle of the antennules slightly exceeds that of the antenn#; first joint about as long as the next two; second joint about three-fourths as long as the third; flagella longer than the peduncle, the “~~ CALIFORNIA STALK-EYED CRUSTACEA. 169 outer one setose on one side except near the base. Peduncle of the antenne armed with short, strong spines; flagellum spinulous, compressed at the base, and exceeding the body in length. Epistome with the trun- cated anterior end armed with five or more spines, the median and outer- most spines the largest. Ambulatory legs smooth, pubescent distally, the coxa of the last pair armed with a pair of spines. Abdominal seg- ments furnished with a pair of transverse dorsal setose sulci which do not meet in the middle line except on the sixth segment. Telson spinulous at the base. Southern California to Mexico. This is the common ‘«spiny lobster’ of the markets. P. gracilis Streets, which has been united with inter- ruptus, differs from the latter, according to Ortmann who has examined the types of both species, in having no transverse sulci on the segments of the abdomen. Subtribe CARIDEA. Body generally laterally compressed, the integument usually of flexible corneous texture. Carapace not united to the epistome. Antennules with a three-jointed peduncle, usually furnished with an external basal spine - and two or three flagella. Antennal scale generally well developed. Ex- ternal maxillipeds generally pediform. FEither, both, or neither of the first two pairs of pereopods may be chelate, the three posterior pairs always simple. Abdomen long, the sides produced downwards. Caudal fin well developed. Gills phyllobranchiate. The Caridea are divided by Bate into four groups:— 1. Crangoninea, with the characters of the single family Crangonida. 2. Polycarpinea: Carpus of the second pereopods annulated, or divided into secondary joints. Four families, Nikidw, Alpheidw, Hippo- lytide, Pandalide. : 3. Monocarpinea: Carpus of second pereopods undivided. This group is divided by Bate into eleven families, only two of which are represented in our limits, the Atyide and the Palemonide. 4, Haplopodinea: All of the thoracic legs similar. Single family, Hec- tarthropida. 170 CALIFORNIA ACADEMY OF SCIENCES. Family CRANGONID. Mandibles slender, incurved, not divided at the apex, and devoid of a palp. External maxillipeds pediform. First pair of pereopods stouter than the second and subchelate, the dactyl closing against the margin of the hand, the pollex spiniform; second pair slender, simple or chelate, the carpus entire; all of the succeeding pairs simple. Genus Crangon Fabr. Rostrum very short and dorsally flattened. Eyes free. Antennules biflagellate. First pair of pereopods stouter than the second and but little longer. Third pair of pereopods slender, subequal to the first pair in length, and terminating ina styliform dactyl; the last two pairs are stouter than the third, and have long, more or less flattened dactyls. Branchiz five. Type.—C. crangon (LINN.). Crangon nigricauda Si. Hh, f- 31 Crangon vulgaris OWEN (non ABR.), Zool. Beechy’s Voyage, 1839, p. 87. Dana, Crust. U. S. Expl. Expd., Part I, 1852, p. 536. Bats, in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 278. Smiru, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 212. Kinestey, Bull. Essex Inst., Vol. XIV, 1883, p. 129. Ratusun R., The Fisheries of the U.S., Sec. I, 1884, p. 817. Murpocu, Rep. Int. Polar Expd. to Pt. Barrow, Pi. IV, 1885, p. 188. Bats, Challenger Reports, Vol. XXIV, 1888, p. 485. Crangon nigricauda STIMPSON, Proc. Cai. Acad. Sci., Vol. I, 1856, p. 89; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 496, Pl. XXII, fig. 6. Srreets and Kinestey, Bull. Essex Inst. Vol. IX, 1877, p. 108.. Kines ey, /bid., Vol. X, 1878, p.54. Lockinerton, Jbid., Vol. X, 1878, - p. 159. Crangon (Steiracrangon) nigricauda KINAHAN, Trans. Roy. Irish Acad., Vol. XXIV, 1862, p. 57; Proc. Roy. Irish Acad., Vol. VIII, 1864, p. 68. Crangon alaskensis Lock1neTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 34. KiINnG@sLey, Bull. Essex Inst., Vol. X, 1878, p. 54. Crangon crangon affinis ORTMANN, Proc. Acad. Nat. Sci. Phila., 1895, p. 180. Crangon vulgaris LINN., var. afinis DE Haan, Walker, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 275. Carapace with a spine on the gastric area and one on each hepatic region, the three situated in a transverse line. Rostrum short, grooved above, and rounded at the tip. Suborbital and antennal spines present, the latter the larger. Thoracic sternum furnished with a spine between the bases of ee ea ad CALIFORNIA STALK-EYED CRUSTACEA. 171 the third pair of pereopods. The process on the onter side of the base of the antennules is broad, irregularly ovate, and about reaches the tip of the first joint; flagella longer than the peduncle, the inner one (and generally the outer) exceeding the tip of the antennal scale. Antenne as long as, or longer than the body; acicle about two-thirds the length of the cara- pace, with the antero-internal angle produced and rounded, extending as far forwards as (often further than) the spine at the outer angle. The external maxillipeds reach about as far forwards as the tip of the antennal scale. The first pair of chelipeds generally extend about as far forwards as the tips of the maxillipeds; merus with a spine near the middle of the lower side; hand oblong, about twice as long as wide, with the edges sub- parallel; the margin against which the finger closes is regularly convex and more nearly transverse than longitudinal; spinous pollex directed obliquely forwards. Third pair of pereopods slender and a little longer than the second. Antepenultimate abdominal segment carinated above; the sixth segment shows a tendency to become carinated and often has a slight median groove. Telson grooved above, subacute at the tip, and exceeding the sixth plus one-half of the fifth abdominal segment. Uropods narrow, rounded at the tip, and equalling (commonly slightly exceeding) the telson. An acute tooth on the sixth segment on either side of the base of the telson and a spine at the posterior inferior angles, between which there is a median ventral spine which is curved backwards; the posterior three-fourths or more of the ventral surface of this segment is marked by a groove. Color in life dark gray with a blackish tail. - Alaska to Lower California. This species differs from C. crangon of the Atlantic in having the fifth and sixth abdominal segments cari- nated above and in the groove on the under side of the sixth segment. It is also distinct from C. affinis. Affinis has a longer rostrum, a much narrower antennal scale, a much narrower and more acute telson. In the speci- men of affinis figured by Bate’ the distal end of the hand is concave and is furnished with a tooth next to the pollex. The corresponding portion of the hand in nigricauda is evenly convex. I have examined the types of Lockington’s Crangon alaskensis. 1 Challenger Reports, Vol. XXIV, F1. LXXXVI, fig. 1. x Gia? CALIFORNIA ACADEMY OF SCIENCES. Crangon franciscorum St.. Crangon franciscorum Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 89; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 495, Pl. XXII, fig. 5. KINAHAN, Trans. Roy. Irish Acad., Vol. XXIV, 1862, p.47; Proc. Roy. Irish Acad., Vol. VIII, 1864, p. 68. Cooper, Rep. Expl. and Sur. Pac. Ocean, Vol. XII, Book 2, 1860, p. 388. Kinestry, Bull. Essex Inst., Vol. IX, 1877, p. 54; /bid., Vol. XIV, 1883, p. 129, Pl. I, fig. 7. Rata- BUN R., The Fisheries of the U.S., Sec. I, 1886, p.818. SHarp, Proc. Acad. Nat. Sci. Phila., 1893, p. 125. Ortmann, Jbid., 1895, p. 181. Near nigricauda but more slender; the number and position of the thoracic spines are the same as in that species and the rostrum has almost the same form. Flagella of the antennules markedly unequal, the inner much exceeding, the outer not reaching the tip of the acicle. The antennz nearly equal, or slightly exceed the length of the body; acicle narrow, about three-fourths the length of the carapace, with the antero-internal angle rounded and not produced beyond the spine at the outer angle. Maxillipeds scarcely reaching so far forwards as the tip of the acicle. Chelipeds more slender than in nigricauda; merus with a spine near the middle of the lower side; hand narrowly oblong, more than three times as long as broad, slightly wider and more inflated near the base; the mar- gin against which the finger closes long, convex, and nearly longitud- inal; the spinous pollex projects nearly transversely. The remaining pairs of legs similar to those of nigricauda. A well developed sternal spine. Antepenultimate abdominal segment rounded above, not at all carinated, with a pair of small spines at the posterior margin at the supero- lateral angles; sixth segment rounded, and devoid oi a carina or groove above, sulcate below, with the spines at the posterior end the same in number and position as in the preceding species. Telson rounded, not grooved above, and tapering to an acute tip. Uropods as in nigricauda, generally slightly exceeding the telson. Color ‘‘ dark and light yellowish gray mottled. Eyes salmon- colosen in life.”’ Length, 54 mm.; length of cheliped, 18 mm.; of hand, 8 mm.; of antennal scale, 11 mm.; of carapace, 14.5 mm.; width of hand, 2 mm. Puget Sound; Shoalwater Bay (Cooper); Tomales Bay and Monterey (Stimpson); San Francisco Bay (Stimp- son)! This shrimp is very common in San Francisco Bay, where it is caught in large quantities for the markets. CALIFORNIA STALK-EYED CRUSTACEA. 173 Crangon nigromaculata Lock. Pl. SRS A es Crangon nigromaculata LocKINGTON, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 34. Crangon nigricauda STREETS and KinesLey, Bull. Essex Inst., Vol. IX, _ 1877, p. 108.2Kinestey, Bull. Essex Inst., Vol. X, 1878, p. 54.) LocxineTon, /bid., Vol. X, 1878, p. 159. | i) : Crangon vulgaris SMiTH, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 212. KINGSLEY, Bull. Essex Inst., Vol. XIV, 1882, p. 129. Crangon “aes afinis ORTMANN, Proc. Acad. Nat. Sci. Phila., 1895, p. 180. | 4% — Closely allied to nigricauda, but possessing the following distinguish- ing marks. It attains a considerably larger size, the rostrum is generally more nearly acute; the antennal scale is narrower and longer, being three- fourths the length of the carapace and subequal to the telson; the hand is a little longer, with the anterior margin more nearly transverse; the ante- penultimate abdominal segment is rounded (not carinated) above; the telson is acute at the tip, rounded above, and equaling (often slightly exceeding) ~ the uropods, which are narrower than in nigricauda. The most charac- teristic and striking mark on this species isa large, colored spot on each side of the sixth abdominal segment. This spot has a bluish center sur- rounded bya black ring, outside of which is another ring of a yellowish color. The sides of the preceding abdominal segments are marked with small, yellow spots.) The chelipeds generally reach but little beyond the middle of the antennal acicle. In none of the specimens examined did they reach to the tip. With the exception of these differences this species . agrees with nigricauda in all the characters mentioned in describing that form. Although this species was regarded as a ‘‘ color variety” of nigricauda by Kingsley and subsequently ranked as a synonym of that species by its own author,’ I am convinced that it should be restored to its rank as a distinct species. I have collected large numbers of nigricauda and nigromaculata from various localities on the coast of California, and have compared numerous specimens of both sexes and of all ages with consider- able care. The large color mark on the sixth abdominal segment appears to be an absolutely constant character; 1Bull. Essex Inst., Vol, X, p. 159. A 174 CALIFORNIA ACADEMY OF SCIENCES. there is nothing corresponding to it or approaching it in any specimen of nigricauda I have seen. Both species are often brought up in the same dredge load and they can easily be separated at sight. The two species occur side by side from northern to Lower California without any notable approach to intergradation. Crangon alba, sp. nov. Closely allied to nigricauda, from which it differs in the following points: The antennal acicle is more narrowed at the tip and the mem- branous portion does not project forward. Hand slightly wider in the middle. The fifth and sixth segments of the abdomen are evenly rounded above and not at all crested. The lower side of the sixth abdominal seg- ment is not grooved. The telson is evenly rounded above and devoid of a sulcus. Color nearly white. Dredged at Monterey, November, 1895. Collection University of California. ” Crangon ice ae om sp. nov. Resembling WN ae oe ‘the spines on the thorax the same in num- ber and position, with the exception that there is no trace of a spine on the gastric area. Body stouter than in franciscorum. The rostrum is longer than in nigricauda, narrow, grooved above and tapering to a slen- der, acute tip, which is curved strongly downwards and much compressed Jaterally. The antennular peduncle does not reach so far forward as the middle of the antennal scale, the process on the outer side of the basal joint similar to that of nigricauda; the second joint of the peduncle is broader than long and much shorter on the outer than on the inner side; inner flagellum longer (but not one-half longer) than the outer, which about equals ora little exceeds the tip of the acicle. Antenne shorter than the body; acicle about two-thirds the length of the carapace and narrow at the distal end, with the antero-internal angle not at all pro- duced. The external maxillipeds reach nearly to the tip of the antennal scale. The anterior chelipeds closely resemble those of nigricauda; a spine near the middle of the under side of the merus; hand oblong, about twice as long as wide, slightly widened distally, with the distal margin convex and oblique; spinous pollex oblique; finger fitting closely against the margin when closed. Second pair of pereopods more slender and a little longer than the third; last two pairs subequal. Sternal spine well CALIFORNIA STALK-EYED CRUSTACEA. 175 developed. Fifth abdominal segment evenly rounded above, having no trace of a carina; sixth segment rounded above, neither grooved or cari- nated, with the spines at the posterior end the same in number and posi- tion as in nigricauda, and with no groove on the lower side. Telson short, about equaling the length of the preceding segment and scarcely more than one-half the length of the carapace, the upper surface rounded (not grooved), the tip somewhat obtuse. The uropods are similar to those of nigricauda and extend considerably beyond the tip of the telson. Each of the abdominal segments in the male bears a median spine on the ven- tral side. Length, 55 mm.; length of carapace, 15 mm.; of antennal acicle, 10 mm.; of telson, 8 mm.; of sixth abdominal segment, 7.5 mm. Two specimens, male and female, dredged at Trini- dad, Humboldt County, Calif., together with numerous specimens of nigricauda. Easily distinguished from the other species on the coast by the absence of a median thoracic spine, the short telson, and the acute rostrum. Collection Uni- versity of California. Crangon munitus (Dana). Crangon munitus DANA, Proc. Acad. Nat. Sci. Phila., 1852, p. 20; Crust. U.S. Expl. Expd., Part 1, 1852, p. 536, Pl. XXXIII, fig. 5. Srimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 497. Kinesiry, Bull. Essex Inst., Vol. X, 1878, p. 54; WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 275. Crangon (Sclerocrangon) munitus ORTMANN, Proc. Acad. Nat. Sci. Phila., 1895, p. 179. Abdomen not sculptured. Carapace seven-carinate, the median carina armed with two small spines. Lateral keels of the carapace smooth and, excepting the second from the median carina, ending in front in spines. Epimera of the abdominal segments without spinules. Puget Sound (Dana). I have examined the specimens from Magdalena Bay, Lower California, which Lockington! has identified with munitus, and cannot agree with him in referring them to that species. - 1Bull. Essex Inst., Vol. X, p. 159. 176 CALIFORNIA ACADEMY OF SCIENCES. Crangon munitellus Walker. Crangon munitellus WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, pi27o;( Pl. XVI, fig. (1: Body stout. Rostrum concave above and rounded at the tip. Carapace with a median carina with two teeth, and two short, parallel carinz on either side terminating anteriorly in a tooth. Antennal acicle short and broad, with a strong central rib. Abdomen abruptly contracted at the fourth segment; ‘‘sixth segment darker colored than the others; a dark, transverse band on the caudal appendages. ‘Length 25 mm.” ‘‘Near C. munitus Dana, but differs in its much smaller size and in the second carina from the median termi- nating in a tooth half-way to the orbital margin, while in C. munitus it reaches the margin and has no tooth.” Puget Sound. Genus Paracrangon Dana. Rostrum elongate; eyes free. Second pair of pereopods wholly absent; fourth and fifth pairs acuminate, gressorial. Type.—P. echinatus Dana. PLL. f 3h echinatus Dana. Pt, 4.36 +37 Paracrangon echinatus DaNa, Proc. Acad. Nat. Sci. Phila., 1852, p. 20,; Crust. U. S. Expl. Expd., Part 1, 1852, p. 538, Pl. XXXIIT, fig. 6. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p.497. Kinas- LEY, Bull. Essex Inst., Vol. X, 1878, p. 55. Wuirkavess, Can. Nat. (2), Vol. VIII, 1878, p. 471. SmitruH, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 212. Faxon, Mem. Mus. Comp. Zool. Harvard Coll., Vol. XVIII, 1895, p. 181. Ortmann, Proc. Acad, Nat. Sci. Phila., 1895, p. 189. ‘‘Beak elongate, obliquely porrect, bidentate at apex, unidentate above near middie, at base below in front a long curved spine. Carapax mul- tispinous, along middle of back unequally four-toothed, either side 5-7 spinous. Abdomen above partly keeled, somewhat sculptured, sides acute. Hand elongate, immovable finger long and very slender. Fourth and fifth pairs of legs nearly naked, subequal. Length of body, one and three-fourths inches. Length of beak half as long as carapax or rather longer than the line of it along the back. The oe i i i i i i CALIFORNIA STALK-EYED CRUSTACEA. 177 exterior maxillipeds are very slender and short hairy. The fourth abdom- inal segment has a tooth and inside of it an emargination either side of middle, and the fifth is nearly similar.” Puget Sound, Oregon, obtained by dredging (Dana). Campbell Is. (Whiteaves); Vancouver’s Is. (Smith); Cal- ifornia and Japan (Faxon, Miers). Genus Sclerocrangon Sars. Carapace sculptured and armed with spines in the median line. Ros- trum securiform, expanded below. First chelipeds stout; second pair slender, elongated, with a narrow hand and short dactyls. Abdomen gen- erally sculptured and carinated above. Type.—S. boreas (PHIPPS). Sclerocrangon boreas (Phipps). Cancer boreas Puipps, Voyage North Pole, 1774, p. 190, Pl XII, fig. 1. Cancer homaroides Fasr. O., Fauna Greenlandica, 1780, p. 241; Mohr. Isl. Naturh. 1786, p. 108, No. 245, Pl. V. Crangon boreas FasR., Syst. Ent. Suppl., p. 409. Saxninse, Appendix to Parry’s Ist Voyage, No. 10, 1821, p. 57. Ross and Owen, Crust. in Appendix to Ross’ 2nd Voyage, 1835, p. lxxxi. Cheraphilus boreas KINAHAN, Proc. Roy. Irish Acad., Vol. VIII, 1864, p. 68. Miers, Ann Nat. Hist. (4), Vol. XX, 1877, p. 57. Murvpocu, Rep. Int. Polar Expd. to Pt. Barrow, 1885, p. 139. Sclerocrangon boreas Sars, Den Norske Nordhavs-Expedition, VI, Zoologi, Crust., Part 1, p. 15. Sreppine, Crustacea, p. 228. OrrT- MANN, Zool. Jahrb. Abth. f. Syst,, Bd. V, 1890, p. 532. Crangon (Sclerocrangon) boreas ORTMANN, Proc. Acad. Nat. Sci. Phila., 1895, p. 178. Carapace with three spines on the dorsal carina; a toothed carina on the branchial regions. Rostrum triangular, acute. Abdomen sculptured, the anterior segments with a median dorsal carina, the sixth segment with a double carina with a sulcus between the ridges. Epimera of the abdom- inal segments with only one spine each. Telson sulcate. Arctic Europe and America, Labrador, Alaska, Si- beria, ‘‘California,’ (Ross and Owen l. c.). 12 178 CALIFORNIA ACADEMY OF SCIENCES. Genus Nectocrangon Brandt. Rostrum wanting. Eyes hidden by the carapace. Second pair of pereo- pods chelate; dactyls of the last two pairs dilated and fitted for swimming. Branchiz five on each side; none on the second maxilliped. Type.—WN. lar (OWEN). Nectocrangon lar (Owen). Crangon lar OwEN, Zool. Beechy’s Voyage, 1839, p. 88, Pl. XXVIII, fig. 1. Argis lar KroéyEerR, Nat. Hist. Tidskr., IV, 1842-3, p. 255, Pl. V, figs. 45-62. Nectocrangon lar BRanpt, in Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 115. Srimpson, Proc. Acad. Nat. Sci. Phila., 1860, p. 25;: Ann. N.¥. liyc. Nat. Hist., Vol. X, 1873,.p. 125. KINncStey Sia Essex Inst., Vol. X, 1879, p. 55. Smiru, Rep. Prog. Geol. Sur. Can-- ada, 1878-9, B, p. 212; Trans. Conn. Acad., Vol. V, 1879, p. 61. Murpocg, Rep. Int. Polar Expd. to Pt. Barrow, 1885, p. 139. ORTMANN,, Proc. Acad. Nat. Sci. Phila., 1895, p. 181. 4 , F Median carina of the carapace armed with two spines behind the one at the anterior end. First five abdominal segments with a median dorsal carina, that of the fifth segment ending behind in a tooth or spine; the two carine of the sixth segment end posteriorly ina small spine. Telson acute, the carinz armed with two or three pairs of spines near the tip.. Circumpolar: Arctic Ocean (Owen, Stimpson); Green- land (Kréyer); Labrador, Nova Scotia, Vancouver’s Island (Smith); Pt. Barrow (Murdoch). Owen’s statement, that in this species ‘‘ the second joint of the superior antenna is dilated and spiniform externally” is an error. He mistook the first joint for the second; the second is cylindrical. A specimen in the U.S. National Museum (No. 7889) from Alaska agrees very closely with several specimens I have seen from the Atlantic. The ridges on the cara- pace are a little plainer than in the Atlantic forms; the carine on the abdomen are a little less acute and havea more decided tendency to fade out behind; the carina. of the fifth segment ends behind in a tooth, but those of the sixth segment do not. In specimens from the North. CALIFORNIA STALK-EYED CRUSTACEA. 179 Pacific (Station 3441 Albatross Collection) the carine of the first four abdominal segments fade out before reaching the posterior end, but that of the fifth segment ends behind in a tooth, and the two carine of the sixth segment terminate posteriorly in small, acute teeth. In specimens from Vancouver’s Island, according to Pro- fessor Smith, ‘‘the dorsal carina on the third, fourth, and fifth segments of the abdomen is broad and rounded, or flattened, and scarcely reaches the posterior edges of the segments, and the two carine on the sixth segment are rounded and fade out in the same way before reach- ing the posterior extremity of the segment.” All of the Pacific specimens I have seen of this species came from farther north than Vancouver’s Island, and it is an interesting fact that they are intermediate in character, as in locality, between specimens from the Atlantic and the forms described by Professor Smith. Nectocrangon alaskensis Kingsley. Nectocrangon alaskensis Kinestey, Bull. Essex Inst., Vol. XIV, 1882, p. 128, OrTMANN, Proc. Acad. Nat. Sci. Phila., 1895, p. 182. Carapace with four equally spaced, median spines and a small tooth between the first spine at the anterior end and the second. Orbits elon- gated. Antennular peduncles reaching about to the middle of the acicle. The abdomen has no carina on the first four segments except a slight one on the anterior portion of the first; the fifth segment has a prominent carina, which ends behind inaspine. The two carinz on the sixth seg- ment end behind ina spine. Telson acute, with three pairs of spines near the tip. Alaska (Kingsley); Station 3257 Albatross Collection, numerous specimens! Family NIKID/& Rostrum horizontal. Ocular peduncles free. Mandibles devoid of a cutting edge and palp. First pair of pereopods simple or chelate. Second pair chelate, longer than the first and more slender, the carpus multiar- ticulate. 180 CALIFORNIA ACADEMY OF SCIENCES. Genus Hippolysmata Stimpson. Carapace furnished with an elongated, compressed, dentate rostrum which is nearly horizontal. Antennules furnished with two long flagella. Mandibles strongly incurved, devoid of a palp, the apex undivided. Max- illipeds elongated, furnished with an exognath, and an epipodite, the ter- minal joint slender. First four pairs of pereopods furnished with an epipodite; the first pair is a little stouter than the others and chelate, the hand oblong; second pair filiform, multiarticulate and chelate. Abdomen smooth above. Type.—H. vitiata STIMPSON. Kingsley in his Revision of the Genera of Crangon- ide, Atyide and Palemonide’ states, in defining this genus, that the first four pairs of feet are provided with an exopodite. This is probably a clerical error, the word exopodite having been substituted for epipodite. Stimpson says in his definition of this genus: ‘‘ Pedes 1 mi-4 ti epipodo instructi.”’ ” Hippolysmata californica St. PL. £.38. Hippolysmata californica Stimpson, Proc. Chicago Acad. Sci., Vol. I, 1866, p. 48; Ann. N. Y. Lyc. Nat. Hist., Vol. X, 1873, p. 123. Kinesuey, Bull. Essex Inst., Vol. X, 1878, p. 56. Hippolyte lineata Lockx1neTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 35. SHARP, Proc. Acad. Nat. Sci. Phila., 1893, p. 116. Rostrum slender, strongly ridged on the sides, bent downwards near the base, about one-half as long as the carapace, scarcely exceeding the ante- penultimate basal joint of the antennules; it is armed above with six or seven teeth, the last tooth situated at considerably more than the usual interval from the preceding one and at about the anterior third of the car- apace; below the rostrum is armed with three teeth; a strong spine above and a smaller one below the base of the antenna on the anterior margin. The spine on the outer side of the basal joint of the antennules reaches about two-thirds as far as the tip; last joint of the peduncle nearly as long as the preceding one; flagella subequal and longer than the body. Antennal acicle elongated, exceeding the tip of the antennular peduncle, and wide at the tip, where it is transversely truncated or broadly rounded. External 1Proc. Acad. Nat. Sci. Phila., 1879, p. 418. 28ee Proc. Acad. Nat. Sci. Phila., 1860, p. 26. ) CALIFORNIA STALK-EYED CRUSTACEA. 181 maxillipeds equalling or exceeding the tips of the antennal scale, the terminal joint elongated and spinulous at the tip. In the first chelipeds the hand is narrow, a little longer than the carpus; fingers about two- thirds as long as the palm. Second pair of pereopods very long and slender; ischium stouter than the merus and about as long, and more or less annulated towards the tip; merus divided into something over twenty annulations; carpus about as long as the merus and ischium combined and divided into about thirty-two annulations; hand minute, oblong, the fingers scarcely as long as the palm. The three posterior pairs of pereo- pods are subequal in length, much longer than the first pair, but shorter than the second; merus joints stouter than the distal ones and armed with scattered spinules; propodi elongated, minutely spinulous below; dactyls very short and armed below with spines. Postero-lateral angle of the fifth abdominal segment acute; that of the fourth segment rounded. Tel- son subacute, much shorter than the uropods, and armed on the rounded dorsal surface with two pairs of spinules. ’ This beautiful species is Very conspicuously marked with longitudinal stripes of drab and red or reddish brown. Length from tip of rostrum to tip of telson, 47 mm.; length of carapace from tip of rostrum, 17 mm.; length of acicle, 9.6 mm.; of first pair of chelipeds, 15 mm.; of second pair, 25 mm.; of third pair, 22 mm.; of tel- son, 7 mm. San Diego (Stimpson)! Santa Catalina Is. (Locking- ton)! San Pedro! Found in tide pools. I have examined the type specimens of Lockington’s Hippolyte lineata. They unquestionably belong to the above species. Family ALPHEID/. Rostrum small or absent. Eye-peduncles short and concealed beneath a projection of the frontal margin of the carapace. Mandibles deeply divided at the apex and furnished with a palp. First pair of pereopods chelate and larger than the second pair which is slender, chelate and has the carpus annulated. Telson broad and rounded. Genus Alpheus Fabr. Carapace compressed, rounded above. Rostrum short or absent. Eyes completely hidden beneath the carapace. Antennules biflagellate. Man- dibles stout, the apical process narrow, palp two- or three-jointed. 182 CALIFORNIA ACADEMY OF SCIENCES. Anterior chelipeds stout, often very unequal. Second pair filiform, carpus cylindrical, annulated, hands small. Type.—A. rapax FABR. KEY TO THE SPECIES oF Alpheus. Front with a rostrum. A spine on the infero-distal angle of the merus of the ~ dent, Res am bulatony plegs 2:55. icuecaekesokaie & ny eed ese oe ee A. clamator. No meral spine. No spine on the second basal antennal joint......... A. barbara. A spine:on the basal antennal joint. Front trispinose. Fingers of larger hand longitudinal; dactyl working nearly vertically...........A4. equidactylus. i Fingers of large hand not longitudinal; dactyl short, curved, working horizontally...A. bellimanus.— Front with but one spine.:.......2.2.... =. A. californiensis. ~ Front devoid of a rostrum; dactyl joined to the lower side of | i the hand (Genus Beteus Dana). -h av lara Front emarginate in the center; hands oval............... A. equalis. Front not emarginate in the center; hands long and WAEVOW oc rateaeee nicer te hlens co etclice Cakee tents eae anette A. longidactylus. Alpheus clamator Lock. LL, fre 37 V Go. Alpheus clamator LockineTon, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 43; Ann. Nat. Hist. (5), Vol. I, 1878, p. 469. Kinestey, Bull. Essex Inst., Vol. X, 1878, p..58; Jbid., Vol. XIV, 1883, p. 117. Alpheus transverso-dactylus Kinestry, Bull. U.S. Geol. Sur., Vol. IV. No. 1, 1878, p. 196; Bull. Essex Inst., Vol. X, 1878, p. 58. Alpheus candei KINGSLEY (not GUERIN), Bull. Essex Inst., Vol. XIV, 1883, p. 124. Front trispinose, the median spine short and separated by a deep sulcus from the small, acute, lateral spines which project nearly as far forward. Antero-lateral angle of the carapace devoid of a spine. Basal spine of the antennules broad, scarcely reaching the extremity of the first joint; second joint about twice as long as the third; outer branch of the flagellum much shorter than the inner one, the slender terminal portion shorter than the basal part; inner flagellum longer than the carapace. Second joint of the antenna with a spine below the articulation of the acicle; acicle narrow, nearly as long as the peduncle, with the outer side concave and ending in a strong spine which is separated by a deep, narrow cleft from the mem- branous inner portion; flagellum longer or shorter than the body. Max- illipeds about equaling the antennal peduncle. Chelipeds very unequal; eS CALIFORNIA STALK-EYED CRUSTACEA. 183 merus of both trigonous, compressed, smooth, the upper surface edge ending abruptly, forming a sharp angle or tooth; carpus of smaller cheli- ped rounded, with a tooth at the anterior end of the upper side; carpus of the large cheliped very short, especially below, and devoid of teeth; large hand oblong, compressed, and greatly developed, being considerably larger than the thorax; proximal portion smooth and nearly naked, the distal portion deeply sculptured; a little in front of the middle of the palm the upper and lower edges are crossed by deep, transverse sulci, which are nearly opposite each other and are continued towards each other on the inner surface of the hand, their ends being separated by a shallow, longi- tudinal depression; the upper transverse sulcus is curved abruptly forwards a little external to the upper edge of the hand and joins a deep, transverse sulcus on the outer face behind the articulation of the finger; this last sulcus is continuous atits lower end witha longitudinal sulcus which ends near the middle of the palm; the transverse sulcus crossing the lower edge of the hand is curved abruptly forwards a little external to the lower mar- gin, forming a longitudinal sulcus which extends to the end of the pollex; this sulcus is separated from the longitudinal one above by a ridge which narrows distally and terminates in a curyed spine or tooth just below the articulation of the dactyl and behind a short groove uniting the lower lon- gitudinal sulcus with the one above it; dactyl flattened, inclined down- wards, but working horizontally, the outer margin strongly convex, the in- ner concave near the thickened tip, but widened into a concave prominence near the base; the postero-external angle is flattened, forming a smooth sur- face which, when the finger is opened, abuts against a similarly flattened prominence behind it; pollex deeply grooved for the reception of the dactyl, ending below in a short, curved tooth. The small hand is several times smaller than the larger one and not so deeply sculptured; the fingers are similar, longitudinal, and about as long as the palm, which is oblong and compressed; a small carina on the upper edge of the dactyl which does not reach the tip; a spine behind the articulation of the dactyl. Carpus of the second pair of legs five-jointed, the first two joints subequal, each about as long as the fourth and fifth combined; third and fourth joints subequal, each shorter than the fifth; hand narrow, fingers slightly longer _ than the palm, whichis subequal to the last joint of the carpus. The two following pairs stout, the merus large and armed with a spine near the infero-distal angle; carpus with a spine at the infero-distal angle and another further back; propodi with five or six spines below. Last pair of legs much less stout than the preceding pairs; merus and carpus devoid of a spine at the infero-distal angle; spines on the propodi feeble; dactyls of all the pairs short, curved, with a small spine behind the tip. Telson furnished above with a broad, shallow groove and two pairs of spinules. Length of body, 31 mm.; of carapace, 11 mm.; of large hand, 18 mm.; of small hand, 11 mm. 184 CALIFORNIA ACADEMY OF SCIENCES. Farallon Is.! Monterey! Santa Barbara! San Pedro! San Diego! San Bartholome Bay, Lower California (Lockington). Kingsley* unites his Alpheus transverso-dactylus with A. candei Guerin from the Bermuda Islands and the West Indies, at the same time listing A. clamator Lock. as a distinct species. But as Kingsley’s description of transverso-dactylus agrees perfectly with clamator, and as his specimens came from a locality where that species is abundant, there is no doubt in my mind that Kingsley’s species is identical with the latter form. Having de- scribed as clamator*? what was really a distinct species, A. barbara Lock., Kingsley made out of specimens which really belonged to clamator a new species, transverso- dactylus. I am unable, moreover, to follow Kingsley when he subsequently unites his transverso-dactylus with A. candei. Candei, according to Guerin,’ ‘‘ a la piece basilaire des antennes externes prolongée en une longue pointe qui atteint la moité de la longeur de l’appendice lamelleaux,”’ while in clamator this spine does not reach one-third the length of the acicle. In candei ‘les cuisses des troisieme et quadrieme pattes n’ont pas de dents,” while in clamator both bear a prominent spine- tooth. The rostrum is smaller in candez than in clama- tor, and there are differences in the form of the larger hand. Alpheus bellimanus Lock. Alpheus bellimanus Pic ree Pico. Cal. Acad. Sci,, Vol. VII, 1877, p. 34; Ann. Nat. Hist. (5), Vol. I, 1878, p. 470. Kinesiey, Bull. U.S. Geol. Sur., Vol. IV, No. 1, 1878, p. 199; Bull. Essex Inst., Vol. X, 1878, p. 59; Ibid., Vol. XIV, 1883, p. 111. 1 Bull. Essex Inst., Vol. XIV, p. 124. 2Bull. U. 8. Geol. Sur., Vol. IV, No. 1, p. 197. 3 De Sagra’s Hist. de Cuba; Crustacés, 1857, p. 50, Pl. II, figs. 9 and 9a. CALIFORNIA STALK-EYED CRUSTACEA. 185 Allied to clamator. Front trispinose, the median spine the largest and not separated from the lateral spines by sulci. Antennules as in clamator; the acicle of the antennezx about reaches the tip of the peduncle and is sim- ilar to that of clamator but the cleft between the spine and the inner por- tion is not so deep as in that species. Maxillipeds pubescent, reaching to the tip of the antennal peduncle. Chelipeds unequal, dissimilar; resem- bling those of clamator, but presenting the following differences: The transverse sulcus on the lower side of the larger hand is prolonged poste- riorly on the inner side of the lower edge as a longitudinal sulcus, a feature not present in that species; there is no longitudinal sulcus on the inner surface of the hand separating the inner ends of the transverse sulci, and the upper, transverse sulcus is not so deep as in clamator. The smaller hand is sculptured much like the larger one but the sulci are not so deep, nor the ridges so prominent; the pollex is rather slender, a little shorter than the palm, and nearly longitudinal, curving a little downwards and having a ridge on the outer side a little above and parallel with the outer edge, the surface above which is somewhat excavated for the recep- tion of the lower margin of the dactyl; dactyl contracted at the base but expanded a short distance beyond into a high, thin, rounded lamina, the plane of which is nearly vertical; the lower edge is nearly straight when seen from the side but curved when seen from below and ends in a short tooth which is curved downward and inward; its postero-external angle has a flattened projection which, when the finger is opened, abuts against a flattened projection of the palm as in the larger hand. Carpus of the second pair of legs five-jointed, the first joint about as long as the next two combined, second and fifth joints subequal and each about as long as the third and fourth together; hand narrow, fingers a little longer than the palm which is somewhat shorter than the last joint of the carpus. The two following pairs of legs not so stout as in clamator, neither the merus nor carpus with a spine at the infero-distal angle; propodi armed below with seven or eight pairs of spines; dactyls short, curved and devoid of a subterminal spine. Length, 33 mm.; length of carapace, 12 mm.; of large hand, 17 mm.; of small hand, 13 mm. Monterey; Santa Barbara! San Pedro! San Diego! Alpheus barbara Lock. Alpheus clamator Kines.ey (not Locki1neTon), Bull. U.S. Geol. Sur., Vol. iy, No: Tiss, p. 197: ; Alpheus barbara Lockineton, Ann. Nat. Hist. (5), Vol. I, 1878, p. 471. KINGSLEY, Bull. Essex Inst., Vol. XIV, 1883, p. 117. I have seen no specimens of this species. It was described by Kingsley from a single imperfect specimen 186 CALIFORNIA ACADEMY OF SCIENCES. which he erroneously considered to belong to clamator Lock. Afterwards Lockington, perceiving that Kings- ley’s clamator was not true to name, rechristened the species A. barbara. The following is Kingsley’s descrip- tion: ‘‘Basal spine of antennule stout, short, not reaching second joint of peduncle; third joint half as long as preceding. Antenne without spine on the basal joint. Antennal scale narrow, the spine at the antero- exterior angle acute, slender, reaching the end of the antennular peduncle. External maxillipeds rather broad, extending slightly beyond the antennal scale. Feet of first pair unequal. Merus smooth, with a very slender spine on the distal portion above. Larger hand compressed, a constriction of each margin at about the middle, a spine above the articulation of the dactylus, behind which a sulcus runs obliquely across the superior mar- gin. Asecond spine on the outside; thumb slender; dactylus compressed, semicircular in outline viewed from the side, slightly longer than the thumb. Smaller hand with both margins constricted; upper margin of palm tuberculate; a spine above the articulation of the dactylus; fingers about equal to the palm, completely closing. Ischium and merus of second pair equal; carpus five-jointed, first two joints equal, and each as long as the third and fourth, which are also equal; fifth joint nearly as longas the first. Merus joints of posterior pairs without spines; propodal joints spinulose beneath; dactyli slender. Santa Barbara, Cal. (W. G. W. Harford).”’ Alpheus californiensis, sp. nov. PLE, £42, pl WL, F-93 VEY. Anterior portion of the carapace convex and furnished with a fine longi- tudinal line. Rostrum short, acute, continuous behind with a short carina, which is separated from the ocular lobes on either side by a deep eroove. Ocular lobes rounded in front and devoid of a tooth or spine. Antero-lateral angle of the carapace broadly rounded. Basal antennular spine broadly ovate, acuminate, and scarcely reaching the tip of the first joint of the peduncle; third joint of the peduncle less than half the length of the second; outer flagellum nearly one-half as long as the inner one, the slender, terminal part about one-half the length of the basal portion; inner flagellum about one-half the length of the body. Antennz about as long as the body; peduncle slightly exceeding that of the antennules; a small spine on the infero-distal margin of the second basal joint; acicle not reaching the tip of the peduncle, outer margin slightly concave and terminating in a strong spine, which is separated from the membranous portion by a deep, narrow cleft. Maxillipeds somewhat exceeding the CALIFORNIA STALK-EYED CRUSTACEA. 187 antennal peduncle. Anterior chelipeds very unequal; merus of both members compressed, and devoid of a spine at the supero-distal angle; carpus without a tooth or spine; hand of large cheliped large, oblong, compressed, upper and lower sides constricted near the middle, the upper constriction formed by a short, transverse groove a short distance behind the dactyl; pollex curved upwards and inwards at the tip and deeply exca- vated at the base for the reception of the dactyl; dactyl compressed, shorter than the palm, the outer margin strongly convex, the inner more or less concave and having a flattened process near the base which fits into the excavation in the pollex. Smaller hand long, narrow, pubescent, somewhat compressed and devoid of sculpturing; fingers narrow, straight, subequal to the palm, the tips hooked and crossing when closed. First joint of the carpus of the second pair of legs long, exceeding the second, which is about as long as the next two combined; third and fourth joints subequal, each a little shorter than the fifth; hand shorter than the last two joints of the carpus. The two following pairs of legs are stouter than the last pair; merus devoid of a spine; propodi strongly spinose below. The sides of the first five abdominal segments are rounded. Telson broadly rounded at the tip and armed above with two pairs of spinules. Length, 37 mm.; length of carapace, 12.6 mm.; of large hand, 15 mm.; of small hand, 9 mm. San Pedro, Calif.; dredged in the harbor, July, 1895. Collection University of California. 4546 Alpheus equidactylus LocKINGT Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 35; Ann. Nat. Hist. (5), Vol, 1, 1878, p. 472. KinesLzy, Bull. U. S. Geol. Sur., Vol. IV, No. 1, 1878, p. 199; Bull. Essex Inst., Vol. X; 1878, p. 59; Jbid., Vol. XIV, 1883, p. 111. Alpheus e uidactylus Lock. Piaf. Carapace smooth, convex longitudinally on the upper side. Front tri- spinose, the median spine considerably larger than the lateral ones and not separated from them by sulci. Basai spine of the antennules long, reach- ing nearly to the tip of the second joint of the peduncle; last two basal joints short and subequal; outer flagellum shorter than the more slender inner one, the slender part much longer than the basal portion. Second joint of the antenna with a spine below the acicle; acicle rather wide, longer than the peduncle. Maxillipeds sparingly pubescent and about reaching the tip of the antennal scale. Anterior chelipeds smooth, naked, unequal, but of similar form; merus with the angles rounded and devoid of spines or teeth; carpus short, similar to that of the larger cheliped in clamator; hands oblong, narrow, compressed, smooth; a transverse sul- cus crossing the upper edge of the hand behind the dactyl, from which a 188 CALIFORNIA ACADEMY OF SCIENCES. narrow, deep, longitudinal suleus extends backwards on the upper edge in the larger hand almost to the posterior margin, but about two-thirds as far in the smaller one; fingers longitudinal, subequal (the dactyl slightly the longer), and working nearly vertically; pollex rather slender, tapering, the tip curved upwards and the inner margin armed near the base with two or three small teeth; dactyl, with one or two very small teeth near the base of the straight inner margin, the tip curved and crossing that of the pollex when closed; smaller hand more slender than the larger one, the fingers somewhat narrower, otherwise very similar. Carpus of the second pair of legs five-jointed, the first joint about as long as the four following ones combined, second, third and fourth joints subequal, the fifth a little longer; hand narrow, the fingers longer than the palm. Posterior pairs of legs slender; merus and carpus devoid of spines; dactyls with very slender tips. Sixth abdominal segment with a postero-lateral spine. Tel- son tapering, rounded above where it is furnished with two pairs of spin- ules, the tip rounded. Length of body, 18 mm.; of carapace, 6.5 mm.; of larger hand, 7 mm.; of smaller hand, 5 mm. Monterey (Lockington)! Santa Barbara! Mr. Kingsley has united A. equidactylus Lock. with A. heterochelis Say,' probably on Lockington’s authority, for he says: ‘‘Mr. Lockington informs me that his A. equidactylus, from Monterey, Cal., presents no appreci- able differences from specimens of A. heterochelis, from Florida, that I sent him.’ There is evidently a mis- take here, for heterochelis and equidactylus are so differ- ent that it is scarcely credible that they should be regarded as members of the same species by anyone who had carefully compared them. Lockington’s spe- cies was described from a single, imperfect, dried speci- men which had lost one of the anterior chelipeds, which Lockington in his later description of the same speci-- men was inclined, in opposition to: his previous opinion, to regard as the larger member. Both hands in this species are elongated much like the smaller hand in heterochelis and, from a superficial comparison, Lock- ington probably concluded that his specimen was a 1 Proc. Acad. Nat. Sci. Phila., 1887, p. 329. CALIFORNIA STALK-EYED CRUSTACEA. 189 heterochelis with the larger hand lost. The two species may, however, readily be distinguished aside from the great differences in the larger chelipeds, as heterochelis has no ocular spines, while in equidactylus the front is ‘‘trirostrate, without sulcus between rostrum and ocular spines” (Lockington). I have examined Lockington’s type which has now lost both its hands. Alpheus Bee) equalis (Lock. ). Beteus equimanus adenine Poe aL. Acad. Sci., Vol. VII, 1877, p. 43. Alpheus Harfordi Kinesiry, Bull. U.S. Geol. Sur., Vol. IV, No. 1, 1878, p. 198; Bull. Essex Inst., Vol. X, 1878, p. 58; Jbid. Vol. XIV, 1883, p. 124 Pl, fie 4 Alpheus equalis KinesLry, Bull. U.S. Geol. Sur. Vol. IV, No. 1, 1878, p. 199. Beteus equalis LockineTon, Ann. Nat. Hist. (5), Vol. I, 1878, p. 478. Carapace moderately compressed, the dorsal side nearly straight. Front devoid of spines, emarginate in the middle and rounded in front of the eyes. Basal spine of the antennules long and slender, reaching beyond the middle of the second joint; second joint about twice the length of the third; the inner flagellum may exceed the length of the carapace; outer flagellum much shorter than the inner one, with the slender terminal por- tion subequal fo the basal part. Second joint of the antennz with a spine below the articulation of the acicle; peduncle about equaling that of the antennules; scale narrowly ovlong, nearly reaching the tip of the peduncle, the outer margin nearly straight and ending in a strong spine which is separated by a deep, narrow cleft from the membranous portion; flagellum shorter than the body. Maxillipeds scarcely reaching the tip of the an- tennal peduncle. Anterior chelipeds subequal; merus trigonous, com- pressed, scarcely more than twice as long as high; carpus devoid of a tooth or spine; hands oval, vertical, smooth, strongly compressed, and devoid of grooves or inequalities; dactyl slender, articulated on the lower side of the hand, and working vertically, the inner margin straight or somewhat concave near the base; pollex much wider than the dactyl; the entire inner margin may be straight, or there may be a deep, rounded notch near the base; tips of the fingers curved and crossed when closed; inner margins pubescent, the rest of the hand naked. Carpus of the second pair of legs five-jointed, the first joint about as long as the three following ones com- bined, which are subequal and shorter than the fifth; hand narrow, palm nearly as long as the fingers or the last joint of the carpus. Posterior legs 190 CALIFORNIA ACADEMY OF SCIENCES. slender; dactyls short, with a subterminal spine. Postero-lateral angle of the sixth, abdominal segment acute, that of the two preceding segments subacute. Telson tapering, rounded at the tip, and armed on the convex dorsal surface with two pairs of spinules. Color in life a dark purple. Leugth of body, 19 mm.; of carapace, 6 mm.; of hand, 6 mm. Catalina Island (Lockington)! Santa Barbara (Kings- ley); Point Arena! This species is found under the mantle of the Aba- lone, Haliotis rufescens, but it is not confined to that habitat, for I found several specimens upon some sea- urchins that were brought up from several feet of water at Catalina Island. Their color was a dark purple like the specimens described by Lockington and resembled the color of the sea-urchins in whose spines they were entangled when captured. At Point Arena I captured from under a rock at low tide a single specimen, which was nearly white. Alpheus (Betzus) longidactylus (Lock. ). Beteus longidactylus Lockineton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 35; Ann. Nat. Hist. (5), Vol. I, 1878, p. 480. Alpheus longidactylus Kinestty, Bull. U. 8. Geol. Sur., Vol. IV, No. 1, 1878, p. 198; Bull. Essex Inst., Vol. X, 1878, p. 58; Jbid., Vol. XIV, 1883, p. 124. Carapace quite strongly compressed and smooth, with no trace of a ros- trum or ocular spines, and scarcely notched at the center. No spines or teeth on the anterior margin of the carapace. Basal spine of the anten- nules slender, elongated, about reaching the tip of the second joint; last two basal joints subequal; inner flagellum a little longer than the carapace, the outer one considerably shorter than the inner and contracted near the middle, the terminal portion slender. Antenne witha spine at the infero- distal angle of the second basal joint; scale with a prominent terminal spine which nearly reaches the tip of the peduncle, the latter reaching about as far forward as the peduncle of the antennules. Maxillipeds nearly reaching the tip of the antennal peduncle. Chelipeds elongated, similar; merus with rounded angles, the distal end widened, the outer surface with a broad, oblique sulcus; a short, transverse groove at the CALIFORNIA STALK-EYED CRUSTACEA. 191 supero-distal angle, behind which the upper margin ends abruptly bu: does not terminate in a spine; the surface is granulated, the lower side granulo-spinulous; carpus short, rounded; hand large, oblong, compressed, scabrous, the margins rounded, the length much exceeding that of all the preceding joints combined; fingers slender, widely gaping, longer than the palm, the tips furnished with small, curved, corneous claws, which are crossed when the fingers are closed; pollex witha large tootha little behind the middle of the inner margin and a small, round tooth at the base. Carpus of the second pair of legs five-jointed, the first joint about as long as the three following ones combined; second, third, and fourth joints sub- equal, the fifth a little longer; hand oblong; fingers about as long as the palm, which is about one-half longer than wide and nearly as long as the last joint of the carpus. Postero-lateral angles of the fourth and fifth abdominal segments acute. Telson tapering, rounded at the tip, with two pairs of spinules above. On the outer side of the distal end of the peduncle of the uropods there isa pair of prominent spines, behind which is a small third spine; aspine near the outer angle of the uropods. ‘‘ Color of carapax ) of dried specimen green with nuances of russet and olive. The fingers of larger hand are light red, the tips green.” | Length, 1.12 inch; length of larger hand, 0.56; of smaller hand, 0.36. San Diego (Lockington)! San Pedro! This description is taken from Lockington’s type specimen (No. 96), which is preserved in the California Academy of Sciences. The specimen was dried and had lost most of its ambulatory legs. The unequal size of the chelipeds is exceptional. Since writing the above description I have collected numerous specimens of this species at San Pedro, Calif., where it is found in abundance in tide pools on a rocky ledge near the entrance to the harbor. The hands are similar and generally equal and very much larger in adult males than in young males and females. The anterior margin of the carapace in some specimens is slightly convex, in others straight, while in a few it is slightly concave. In many adult females, and to a less extent in the adult males also, the dorsal surface of the carapace is bulged upwards, owing, doubtless, to the enlargement of the ovaries or testes. Many of the 192 CALIFORNIA ACADEMY OF SCIENCES. specimens were infested with a parasitic Isopod attached to the under side of the abdomen. The color varies in living specimens from olive green to olive brown; the legs are reddish, and in many specimens there is a light-colored dorsal stripe along the middle of the body. The specimens were taken June 25, 1895; most of the females were carrying ova. | Family HIPPOLYTIDA. Rostrum generally prominent. Eyes not covered by the carapace. Man- dibles with or without a cutting edge and palp. First pair of legs chelate and stouter than the second pair which is also chelate and has the carpus annulated. The genera of this family need a thorough revision. The genus Hippolyte has long been a sort of receptacle for a large and somewhat miscellaneous assemblage of forms. Bate has instituted several new genera for many of the species but several more will doubtless be required. Most of the Pacific species of this family fall naturally into a group which I have characterized as the genus Heptacarpus. Genus Hippolyte Leach. Carapace produced into a laterally compressed, serrated rostrum which is excavated at the under side of the base. A supraorbital and antennal spine present. Ocular peduncles short. First joint of the antennular peduncle excavated above and armed with an external basal spine; second and third joints subcylindrical, the terminal one supporting two flagella, the outer of which is the shorter and more robust. Antennal scale with a spine at the distal end of the outer margin. Third maxillipeds with a short exognath and having the last joint armed with spinules at the tip. First pereopods short and stout with the carpus excavated at the extrem- ity to receive the base of the hand. Second pair slender, a little longer than the first, and having the carpus three-jointed. The three following pairs of legs decrease slightly in length posteriorly; dactyls spinulous below. Abdomen smooth, the third segment posteriorly produced in the median dorsal line and somewhat arcuate. Branchie five. Type.—fH. varians LEACH. CALIFORNIA STALK-EYED CRUSTACEA. 193 Bate’s diagnosis of this genus was based on an exam- ination of Leach’s type specimens of varzans. I have slightly modified Bate’s description, as some of the spe- cies which Bate himself refers to Hippolyte contradict the characters of the genus as he defines it. Hippolyte californiensis Holmes. 2). Hippolyte californiensis Hotmes, Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 576, figs. 21-26. A long slender species. Rostrum slender, slightly upturned, a little longer than the carapace; upper margin armed with 3-5 teeth; lower margin with 4-5 teeth; base of the rostrum rounded and not continued upon the carapace. A supraorbital spine. Peduncle of the antennules about one-half as long as the rostrum; outer flagellum much shorter than the slender inner one, the last few joints much narrowed. First pair of chelipeds very short; hand broad and thick at the base, which fits into a depression in the carpus. Second pair of chelipeds more slender but much longer than the first pair; carpus three-jointed, the first joint the longest. Abdomen not crested or carinated. Telson truncated and spinulous at the tip. Length, 38 mm. Bodega Bay, Calif., in the eel-grass! San Pedro! San Diego! Specimens from the last named locality are con- siderably smaller than those from Bodega Bay. Hippolyte Layi Owen. Hippolyte Layi Owen, Zool. Beechy’s Voyage, 1839, p. 90, Pl. XXVII, fig. 3. Branpt, in Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 117. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 499. Bates, in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 279. Kines- LEY, Bull. Essex Inst., Vol. X, 1878, p. 62. Lock1neton, Jbid., 1878, p. 161. I have seen no specimens which I can refer to H. Layi. Owen’s short description of this species is as follows: ‘* Hip. rostro acuminato, supra multi-serrato, ante medium subtus quadri-serrato. Long. corp. unc. 24. Color ruber.” A little more light on the subject is given in the discussion of the affin- ities of this species. ‘‘ In addition to the difference in the serrations of 13 ; 194 CALIFORNIA ACADEMY OF SCIENCES. the rostrum, which probably varies in different individuals of the same species (although the number was constant as above quoted in three specimens of each species which was preserved), Hip. Layi has a longer and narrower rostrum, extending to the extremity of the superior anten- ne; the inferior seta of these antenne& is also proportionately longer. Hip. affinis has a spine above the eye, at the root of the rostrum, which is want- ing in the Hip. Layi; whilst this species has the fifth and sixth abdominal segments unispinous inferiorly on each side, Hip. affinis having the sixth segment only so armed. “From the Alpheus Polaris of Sabine (Zool. Appendix to Parry’s Voyage, p. 238, Pl. II, fig. 5) which is a Hippolyte of Leach, and from the species characterized by the latter author in the Malacostraca Podoph. Brit., the above species differ in the forms and proportions, as well as in the serra- tions of the rostrum.” Monterey (Owen); Vancouver’s Island (Bate); Alaska (Lockington). It is impossible to determine from Owen’s description | and figure in what genus this species belongs. It is certainly not a Hippolyte; it may be a Heptacarpus, but until more is known concerning it, it is best, I think, to leave its name unchanged. 7 Hippolyte affinis Owen. Hippolyte affinis Owen, Zool. Beechy’s Voyage, 1839, p. 90, Pl. XXVII, fig. 4. Branpt, in Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 117. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, 498. KinesLey, Bull. Essex Inst., Vol. X, 1878, p. 62. Owen’s short description of this species is as follows: ‘‘ Hip. rostro antennis superioribus breviore, supra multi-serrato, ante medium subtus sex-serrato. Long. corp. 14. Color ruber.” Rostrum shorter than the antennules, multi-serrate above, and armed below with six teeth in front of the middle. Length of the body 13 inch. Color red. Monterey (Owen). There are several species of Hippolyte reported from Vancouver’s Island by Professor Smith: viz., H. Gav- mardit Milne-Edw.; H. Grenlandica, under which are CALIFORNIA STALK-EYED CRUSTACEA. 195 included H. armata and H. cornuta of Owen; H. Phippsit Kroyer, with which Smith unites H. turgida Kroyer, H, vibrans St. (46 var.), and H. ochotensis Brandt; and H. spinus, which is now the type of Bate’s genus Spiron- tocaris. All of these are circumpolar species, the west- ern representatives of which I have not seen. Another species of Hippolyte from Vancouver’s Island has been described by Bate as H. esquimaltiana. Itis not, however, a true Hippolyte, but the description is. not sufficiently complete to enable one to determine just where it belongs. Hippolyte sitchensis Brandt has been reported from California by Stimpson, but a remark in Stimpson’s description of H. picta discloses the fact that what he called sztchensis was probably another species. “«Picta,”’ says Stimpson, ‘‘ differs from H. sztchensis in its longer external maxillipeds and non-carinated abdo- men.” It is certainly here implied that sztchensis has a carinated abdomen which, however, is not the case. Brandt in his description of that species says: ‘‘Annulus abdominis tertius facie dorsali ecarinatus,”’ and further on that this species differs from H. gibba ‘‘ durch den Mangel des Riickenkieles des dritten Bauchgiirtels.” Heptacarpus, gen. nov. Carapace carinated anteriorly and furnished with a dorsally serrated rostrum; the anterior margin is furnished with two spines, one just below the orbit, the other (generally the smaller) at the antero-lateral angle. No supraorbital spine. Ocular peduncles short, stout, single-jointed, and furnished with an ocellus on the posterior side. Antennules with a well developed outer basal spine; flagella two, a thick outer one terminating in a slender portion,and a slender (generally longer) innerone. Antenne long, with a well developed scale, the outer margin of which ends in a spine. Mandibles strongly incurved, furnished with a slender apical process and a two-jointed palp. External maxillipeds devoid of an exognath. Anterior chelipeds stout, the carpus not excavated in front as in Hippolyte; hand oblong, the fingers ending in small curved claws or hooks. Second pair of 196 CALIFORNIA ACADEMY OF SCIENCES. chelipeds filiform; carpus seven-jointed; hand small and narrow. Pos- terior legs of subequal length; carpus produced distally over the propodus; propodi and dactyls spinulous below. Telson tapering and armed above with several pairs of spines. Branchiz five. Type.—H. palpator (OWEN). Beside the characters mentioned in the diagnosis of the genus, all of the species I have seen agree in pos- sessing the following features: There is a spine at the antero-external angle of the oblong first joint of the antennular peduncle and another spine on the lower margin of the inner side a short distance behind the distal end; the second joint of the peduncle has a spine on the outer side, and the third joint has a spine above the articulation of the thick flagellum, and there is generally a small spine above and internal to the articu- lation of the slender one. On the second joint of the antenna there is a blunt tooth above and a sharp spine below the outer edge of the scale. The tips of the external maxillipeds are armed with a circle of spines. The carpus of the anterior chelipeds has a short, trans- verse groove behind the inner distal margin. Heptacarpus palpator (Owen). PLL, 48S Y9, : Hippolyte palpator OwEn, Zool. Beechy’s Voyage, 1839, p. 89, Pl. XXVIII, tig.3. Branpt in Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 117. Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 89; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 499. KinesLry, Bull. Essex Inst., Vol. X, 1878, p. 62. Lockineron, Jbid., Vol. X, 1878, p. 160. ? Hippolyte Hemphillii Locxineton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 35; Bull. Essex Inst., Vol. X, 1878, p. 160. Kineastey, Jbid., Vol. X, 1878, p. 63. Rostrum slender, horizontal, considerably shorter than the carapace and armed above with six teeth, the last two (or three) of which are on the cara- pace; the lower margin may be devoid of teeth, or there may be a small tooth near the tip. The outer basal spine of the antennules reaches. nearly to the tip of the second joint of the peduncle; the upper distal margin of the first basal joint is armed with four spines, the spine at the ae, wy CALIFORNIA STALK-EYED CRUSTACEA. 197 outer angle being separated by a considerable interval from the other three which are close together. The spine above the articulation of the slender inner flagellum is very short; outer flagellum much shorter than the inner one. Antennal acicle tapering, longer than the telson; peduncle reaching about to the middle of the acicle which is in advance of the tip of the ros- trum; fiagellum often longer than the body, Mandibles with an oblong, thin, apical process which is armed at the distal end with several small teeth; palp two-jointed, about as long as the apical process behind which it normally lies. Maxillipeds very large, extending far beyond the tip of the antennal scale even in small specimens, while in adults the antepenul- timate joint often reaches the tip of the acicle, the last joint extending beyond it for a distance greater than the length of the carapace; the outer margin of the antepenultimate joint is spinulous; last joint flattened, about four times the length of the preceding one and armed with a circle of spines at the tip and several smaller spines on the inner margin; there is a small epipodite but no exognath. Anterior chelipeds reaching con- siderably beyond the tip of the antennal scale; hand narrowly oblong. Second pair of chelipeds longer than the first; first joint of the carpus longer than the second; third joint longer than the first or fourth; fifth subequal to the fourth, longer than the sixth, which is much shorter than the seventh; hand about as long as the last three joints of the carpus. Posterior legs moderate, two or three spines on the outer side of the merus near the distal end; dactyls short, stout, about one-fourth the length of the propodi. Ail of the abdominal segments are rounded above; postero- lateral angles of the fourth segment rounded or subacute, having a small spine-tooth; those of the fifth segment acute and ending inaspine. Tel- son somewhat flattened above, where it bears four pairs of spinules; tip broadly rounded and furnished with a minute median spine’ between which and the spines at the outer angles there are on either side about four marginal sets. Monterey (Owen)! San Francisco Bay (Stimpson)! San Pedro! Santa Catalina Island! San Diego! Magda- lena Bay (Lockingtén). Brandt refers some specimens from Alaska to this species. He says: ‘‘Wosnesenski brachte von der Insel Kadjak vier Exemplare einer Hippolyte mit die durch die langen, aiisseren Maxillarfiisse und mehrere andere Kennzeichen H. palpator ungemein ahneln. Ich méchte sie daher vorlaufig zu ihr ziehen, obgleich, abweichend von palpator der Thorax derselben oben zwei-bis dreiz- ahnig, der Stirnschnabel oben meist zwei-, zuweilen 198 CALIFORNIA ACADEMY OF SCIENCES. einzahnig, unten zahnlos und nur so lang wie die Augen erscheint, wahrend die inneren Antennen die Deckschuppen der aitissern Fuhler deutlich tiberragen. Die gréssten Individuen messen von der Stirn zur Schwanzspitze 2”, 4", die Lange des Thorax betragt 94.” I think it more probable that the species Brandt de- scribes belongs to H. brevirostris (Dana) than to H. pal- pator (Owen). I have never seen any specimen of pal- pator attain nearly such a large size as the specimen measured by Brandt. The other characters mentioned by Brandt agree perfectly with brevirostris. The locality _at which Brandt’s specimens were taken also favors this supposition, as brevirostris appears to be a more north- ern species than palpator, having been taken as far north as Vancouver’s Island (Smith). Heptacarpus brevirostris (Dana). i i Wieeae: brevirostris DANA, Proc. Acad. Nat. Sci. Phila., 1852, p. 24; Crust. U.S. Expl. Expd., Part I, 1852,.p. 566, Pl. XXXVI, fig. 5. Stimpson, Proc. Cal. Acad. Sci., Vol. I, 1856, p. 89; Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p.500; Proc. Acad. Nat. Sci. Phila., 1860, p- 33. Bats, in Lord’s Nat. in Vancouver’s Is., Vol. II, 1866, p. 279. SmituH, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 214. KINGSLEY, Bull. Essex Inst., Vol. X, 1878, p. 61. Lockinerton, Jbid., Vol. X, 1878, p. 160. Hippolyte palpator BRANDT, Middendorff’s Siberische Reise, Bd. II, Th. By 18a1, pally. Spirontocaris brevirostris WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 276. Allied to palpator, but larger and more robust. Carapace crested for a. little more than its anterior third; rostrum short, not reaching the tip of the first joint of the peduncle of the antennules and armed above with five or six teeth, the last three or four of which are on the carapace, the pos- terior one being a little in advance of the middle; lower side of the ros- trum devoid of teeth. The basal spine of the antennules reaches beyond the first joint of the peduncle; the spine above the articulation of the inner flagellum is very short; outer flagellum reaching beyond the antennal scale; inner flagellum about one-half longer than the outer. Antennal CALIFORNIA STALK-EYED CRUSTACEA. 199 scale broad, short, about equal to the length of the telson; the peduncle extends about two-thirds the distance to the tip of theacicle. Maxillipeds very large, similar to those of palpator ; in adults the tip of the antepe- nultimate joint may reach the tip of the antennal scale. The first pair of chelipeds often reaches beyond the acicle. The relative lengths of the joints of the carpus of the second pair are about the sameas in palpator; hand as long as the last three carpal joints. Posterior legsstouter than in palpator but otherwise very similar. Postero-lateral angle of the fourth abdominal segment broadly rounded but furnished with a minute spine-tooth; that of the fifth segment acute. Telson dorsally flattened, having four or five pairs of spinules; tip rounded, wider than in palpator, and having several marginal setz between the small median spine and the postero-lateral spines. Length, 49 mm.; length of carapace, 16 mm.; of external maxillipeds, 32 mm. Sitka (Brandt); Vancouver’s Island (Smith); Puget Sound (Dana); Humboldt County, Calif.! San Francisco Bay (Stimpson)! Differs from palpator in its larger, stouter body, shorter rostrum, much shorter and broader antennal scale, and stouter posterior legs. De ore (st. [ge +S 23-5 2. Hippolyte Taylori Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 500. Kinestey, Bull. Essex Inst., Vol. X, 1878, p. 61. LocKine- ton, Jbid., Vol. X, 1878, p. 160. A short, plump species very closely allied to brevirostris, but the rostrum is shorter, reaching but very little beyond the anterior margin of the cara- pace and armed above with five or six teeth, the last three or four situated on the carapace; the teeth become more closely set and more strongly inclined forwards anteriorly, the end of the rostrum appearing curved downwards, although the lower margin is nearly horizontal; there is but one spine on the upper distal margin of the first joint of the antennules; the telson is narrower than in brevirostris, and the distance between the small median spine at the tip and the spines at the postero-lateral angles is not nearly so great, and there are only one or two marginal sete on either side of the apex, instead of four or five as in the preceding species. None of the specimens of Yaylori I have seen attain nearly the size reached by brevirostris. The rostrum is rarely so long as in that species, and when it is so it maintains its characteristic appearance. ~~ 7\ 200 . CALIFORNIA ACADEMY OF SCIENCES. o San Francisco Bay! Pescadero! Monterey! Magdalena Bay, Lower California (Lockington). Although this species is very close to brevirostris, a comparison of several specimens of both species from different localities shows that they are distinct. Taylori is more southern in its range than brevirostris, but both species occur together in San Francisco Bay. eee pictus (St. ). PLUG, f SE FES. Hippolyte picta Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. X, 1873, p. 125. Kinasuey, Bull. Essex Inst., Vol. X, 1878, p. 62. A small species, less robust than brevirostris or Taylori. Rostrum slender, straight, horizontal or slightly upturned and about two-thirds the length of the carapace; upper margin with six or seven quite evenly spaced . teeth, the last two of which are on the carapace; lower margin with two to four teeth near the tip. The tip of the peduncle of the antennules reaches the middle of the antennal acicle but not the tip of the rostrum; the outer basal scale of the antennules reaches beyond the first joint of the ped- uncle but scarcely reaches the tip of the second joint; no spines on the distal margin of the first basal joint except the one at the external angle; no spine above the base of the slender flagellum; outer flagellum reaching a short distance beyond the tip of the antennal scale; inner flagellum about one-half longer than the outer. The antenne may exceed the length of the body; acicle oblong, slightly tapering, and generally exceeding the tip of the rostrum. Maxillipeds shorter and more slender than in palpa- tor, but reaching beyond the tip of the rostrum even in quite small speci- mens; last joint in adults about four times the length of the preceding one. Anterior chelipeds not reaching the tip of the antennal scale. The first and second joints of the carpus of the second pair are subequal; third joint about equal to the first two combined; fourth joint subequal to (or somewhat shorter than) the third, longer than the fifth, which exceeds the sixth; seventh joint considerably longer than the sixth; hand shorter than the last three joints of the carpus. Posterior legs moderately stout; the distal portion of the outer surface of the merus joints armed with a row of spines, which in the first pair often extends more than half way to the base; dactyls short, stout, about one-fourth the length of the propodi, and armed below with spines, the largest of which are near the tip. None of the abdominal segments are carinated above; postero-lateral angle of the fourth abdominal segment obtuse but having a small tooth; that of the fifth segment acute and produced intoa spine. Telson rounded above and CALIFORNIA STALK-EYED CRUSTACEA. 201 armed with four or five pairs of spinules; a small median spine at the tip, between which and the spines at the lateral angles there are one or two marginal setz. Carapace beautifully marked with oblique crimson- bands; legs barred with crimson, the dactyls and tips of the propodi light colored. Monterey (Stimpson); Santa Catalina Island! San Pedro! San Diego! This beautiful little species is commonly found in tide pools. It is abundant at Monterey and San Pedro on rocky shores. Heptacarpus paludicola, sp. nov. PE ef. SET EF Closely allied to picta. Rostrum slender, horizontal, about as long as the carapace, armed above with 6-8 sharp, evenly spaced teeth, the last tooth on the anterior fourth of the carapace; lower margin armed with 2-4 teeth on the distal third or two-fifths. Antennular peduncle reaching two-thirds as far as the tip of the rostrum; basal spine extending beyond the tip of the first joint; upper distal margin of the first basal joint armed with a single spine at the outer angle. Antennal peduncle not reaching the middle of the acicle;.acicle oblong, slightly tapering, closely resem- bling that of picta, and equalling the rostrum; flagellum as long as the body. Anterior chelipeds not reaching the tip of the acicle; hand narrow; palm fully as long as the carpus and about three times as long as wide. Second chelipeds about reaching the tip of the acicle; first joint of the carpus but little longer than the second, third and fourth subequal and each longer than the first, about equalling the fifth and sixth combined, fifth a little longer than the sixth and slightly shorter than the seventh; hand scarcely longer than the last two joints of the carpus. Posterior legs very slender; distal portion of the outer surface of the merus joints armed with a row of spines; dactyls short, about one-fourth the length of the propodi, narrower than in picta but similarly armed. Postero-lateral angles of the fourth abdominal segment subacute and furnished with a small tooth; that of the fifth segment acute. Sixth segment a little longer than in picta. Telson as in the preceding species. Color uniform green. Humboldt Bay! Shelter Cove! Bodega Bay! June, 1894. Large numbers of this species were caught in a trawl in the eel-grass in Humboldt Bay and Bodega Bay. The color is almost exactly that of the eel-grass in which 202 CALIFORNIA ACADEMY OF SCIENCES. they live. This species may be distinguished from picta by its larger size, different color, longer and more slender rostrum and the much more slender ambulatory legs. Collection University of California. Heptacarpus cristatus (Si. ). TE, f.5¥ 54. Hippolyte cristata Stimpson, "Proc. Acad. Nat. Sci. Phila., 1860, p. 33. KINGSLEY, Bull. Essex Inst., Vol. X, 1878, p. 62. Spirontocaris cristata WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 277. Less robust than Yaylori. Rostrum rather narrow, about two-thirds the length of the carapace, arched over the eyes where the teeth are most thickly set, and armed above with 5-8 teeth, the posterior two or three being on the carapace, while the most anterior one is situated some dis- tance behind the tip; lower margin with two or three teeth near the tip. The basal spine of the antennules reaches a little beyond the tip of the first joint. Antennal scale considerably longer than the rostrum and narrower than in palpator or Taylori; flagellum longer than the body. Maxillipeds reaching to or beyond the tip of the acicle; hand of the anterior chelipeds long and narrow. Second pair of chelipeds reaching the tip of the acicle; first joint of the carpus slightly longer than the sec- ond; third about as long as the first two; fourth, fifth and sixth decreasing successively in length; seventh longer than the fourth; hand nearly as long as the last three joints of the carpus. Posterior legs more slender than in palpator, the merus joints armed with spines on the distal portion of the outer surface; dactyls long, slender, evenly tapering, about half the length of the propodi. Abdomen moderate, none of the segments cari- nated above; postero-lateral angles of the fourth segment rounded but gen- erally bearing a minute spine; those of the fifth segment acute and having a larger spine. Telson rounded above and having three pairs of dorsal spinules. San Francisco Bay (Stimpson); Monterey! San Diego! Easily distinguished from the other species described here by its long, slender dactyls. Heptacarpus carinatus, sp. nov. T'L.IL, , 60. Carapace small in Teme to the abdomen; antennal spine well developed, larger than the spine at the antero-inferior angle. Rostrum rather narrow, evenly tapering, about equalling the carapace in length, , CALIFORNIA STALK-EYED CRUSTACEA. 203 the upper margin armed with 4-6 small teeth, the posterior one on the carapace, the anterior one'a little behind the middle of the rostrum; lower margin armed with 4-6 teeth, the anterior tooth commonly near the tip. Ocular peduncles with an ocellus on the posterior side. The spine on the outer side of the base of the antennules reaches about to the tip of the first joint; antennal scale narrow, about equalling the length of the rostrum, the sides parallel, the inner, membranous portion projecting much beyond the outer distal spine; flagellum about as long as the body; the peduncle reaches about as far forward as the penultimate joint of the peduncle of the antennules. The external maxillipeds scarcely reach as far forward as the middle of the rostrum; last joint flattened, somewhat twisted, the tip armed with dark colored spines. The anterior chelipeds do not reach as far forward as the maxillipeds; hand thick at the base. Second pair of chelipeds reaching beyond the tip of the antennal peduncle; first and third joints of the carpus subequal and longer than the second; fourth and fifth joints subequal and shorter than the third; sixth joint the shortest of ail; seventh about as long as the two preceding ones combined; hand narrow, the paim a little longer than the last joint of the carpus. Posterior legs rather stout, the extero-distal angles of the merus joints armed with 1-3 spines, the last pair seldom having more than one. The third segment of the abdomen is posteriorly produced and crested. Tel- son with four pairs of spines on the dorsal surface. Dredged in large numbers in shallow water in Mon- terey Bay, November, 1895. The color varies with the surroundings. Specimens among the bright green sea-weeds are of a uniform bright green color, while other specimens living only a few yards away among the red sea-weeds imitate almost , exactly the color of the alge that surround them. Collection University of California. USN M 59579 - Heptacarpus tenuissimus, sp. nov. e A very slender species. Anterior half of the carapace crested. Suborb- ital and antennal spines well developed. Rostrum longer than the cara- pace, horizontal and very slender, scarcely higher than wide, gently arching over the eyes, and armed above with four spines, the posterior spine on the carapace, the anterior one near the middle of the rostrum, the distal half of the upper margin being smooth; lower side of the rostrum with five teeth. Basal spine of the antennules scarcely reaching the tip of the first joint. Acicle long and narrow, about equalling the rostrum, the 204 CALIFORNIA ACADEMY OF SCIENCES. sides parallel; peduncle not reaching the middle of the acicle. Mazxillipeds rather short. Anterior chelipeds scarcely reaching the tip of the antennal peduncle. The three posterior pairs of legs are-slender and have short dactyls. Abdomen slender, the third joint rounded above but produced backwards in the middle. Telson narrow and armed above with seven pairs of minute spinules. Length, 25 mm. One specimen collected at Monterey by Dr. H. P. Johnson. USNM £6808 Easily distinguished from all the other species of Heptacarpus by its very slender, elongated body and very narrow rostrum. Collection University of California. Heptacarpus Herdmani (Walker). Spirontocaris Herdmani WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 277, Pl. XVI, fig. 2; Ann. Mag. Nat. Hist. (7), Vol. II, 1898, DG: Carapace with a subocular spine and a small tooth at the antero-inferior angle. Rostrum horizontal, a little shorter than the carapace, the dorsal carina continued back upon the carapace nearly to the middle; ‘‘ upper margin with five teeth, of which two are on the thorax, the second, third. and fourth close together, the distance from the fifth to the point equal to the length from the second to the fourth. Lower margin with one tooth near the point.” Maxillipeds reaching considerably beyond the tip of the acicle. First pair of legs reaching beyond the tip of the acicle, ‘‘ the propodus as wide and more than twice as long as the carpus.” ‘‘Abdom- inal segments having the lower margin rounded in the first four, acute in the fifth.” ‘One female with ova. Length, 30 mm.” Heptacarpus ? stylus (S?.). Hippolyte stylus Stimpson, Proc. Acad. Nat. Sci. Phila., 1864, p. 154. Kinestey, Bull. Essex Inst., Vol. X, 1878, p. 62. x Body slender. Carapace smooth, not crested, except for a short distance in front. Rostrum slender, styliform, perfectly straight and equal to the carapace in length; the upper side is armed with four or five teeth near the base, while the anterior two-thirds is devoid of teeth; lower side with five or six teeth. An antennal spine present but no supraorbital or ptery- gostomian spine. Antennal scale oblong, scarcely shorter than the rostrum CALIFORNIA STALK-EYED CRUSTACEA. 205 and obliquely truncate at the end. Maxillipeds very small, reaching only to the end of the antennal peduncle, or to the basal third of the ros- trum; there is an epignath but no exognath. None of the pereopods have an epipod. Telson with four pairs of spinules on the dorsal surface. Length, 14 inch. Straits of Fuca (Stimpson). Heptacarpus ? Suckleyi (Sz. ). Hippolyte Suckleyi Stimpson, Proc. Acad. Nat. Sci. Phila., 1864, p. 154. KiIn@sLey, Bull. Essex Inst., Vol. X, 1878, p. 62. Carapace with the anterior half crested and sloping forwards. An an- tennal and pterygostomian spine present but no supraorbital spine. Ros- trum scarcely as long as the carapace, rather broad and curved, with a slender, acute tip; upper margin six-toothed, beginning at the anterior third of the rostrum; lower margin four-toothed. Maxillipeds reaching nearly to the tip of the acicle and devoid of an exognath or epignath. Pereopods long, the last pair nearly reaching the tip of the rostrum; first pair only provided with an epipod; dactyls of the last three pairs elon- gated, with only one terminal claw. Abdominal segments with smooth _ Margins, the superior margin of the third segment obtuse. Length, 1} inch. Puget Sound (Stimpson). Heptacarpus? gracilis (St. ). Hippolyte gracilis Stimpson, Proc. Acad. Nat. Sci. Phila., 1864, p. 155. KINGSLEY, Bull. Essex Inst,, Vol. X, 1878, p. 62. A slender species. Carapace crested on the anterior third. Rostrum very slender, scarcely higher than wide and a little longer than the cara- pace; the upper side is armed with four teeth over the eye, in front of which it is smooth to the tip; lower side armed with four minute distant teeth. An antennal and a pterygostomian spine present. The thick flageilum of the antennules reaches the tip of the rostrum. Acicle a little longer than the rostrum. The maxillipeds reach the middle of the ros- trum and have no exognath. Pereopods very slender and devoid of an epipod. Abdomen very long and strongly geniculated, the third segment compressed and prominent, penultimate joint much elongated. Length, 1} inch. Puget Sound (Stimpson). Found in deep water. 206 CALIFORNIA ACADEMY OF SCIENCES. Genus Spirontocaris Bate. Carapace anteriorly carinated and produced into a deep, laterally com- pressed rostrum which is serrated on both margins. There are two or more supraorbital spines, an antennal spine, and a spine at the antero- inferior angle of the carapace. First and second antenne much as in Hippolyte. Mandibles with a broad molar process, a slender apical process, and a small, two-jointed palp. Third maxillipeds furnished with a small exognath and having the tip of the last joint armed with spinules. First pair of pereopods short, robust; second pair slender, the carpus seven- jointed. Posterior pereopods subequal, similar, and furnished with biun- guiculate dactyls which are spinulous below. Type.—S. spinus (SOWERBY). Spirontocaris prionata (St. ). Hippolyte prionata Stimpson, Proc. Acad. Nat. Sci. Phila., 1864, p»153. Kinestty, Bull. Essex Inst., Vol. XIV, 1883, p. 127, Pl. T1gm@eies- SHARP, Proc. Acad. Nat. Sci. Phila., 1893, p. 117. Spirontocaris prionata WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 277. Carapace large, the anterior margin furnished with two spines, of which the upper one is the larger. There are two or three supraorbital spines in a longitudinal series; the upper side is crested nearly to the posterior margin and cut into three large teeth whose transverse anterior margins are armed on either side with severalsmallspines. Rostrum-very broad, widest a little behind the tip, the upper margin thickly set with small spines, while the lower margin is furnished with four or five larger spines. An- tennules short, the external basal spine large, reaching about to the tip of the peduncle; the first joint of the peduncle is longer than the next two and has a spine at the antero-external angle; second joint with a large spine on the outer side; third joint with a slender spine above the articu- lation of the larger flagelium and a similar spine near the articulation of the slender one; thick flagellum longer than the slender one, with a slen- der tip about one-third the length of the proximal portion. Antenne somewhat shorter than the body; acicle subtriangular, about reaching the tip of the rostrum; peduncle not reaching the middle of the acicle. Man- dibles stout, cutting edge long, slender, distally four-toothed, and slightly longer than the palp; palp two-jointed, the last joint narrow and about twice as long as the first, Maxillipeds very stout and exceeding the tip of the rostrum, the last two joints bent downwards; the antepenultimate joint is somewhat widened at the tip where it bears, on the upper side, two small prominences, each of which bears a spine and numerous sete; last joint over twice the length of the preceding one, the tip armed with a circle of dark-colored spines; exopod small, scarcely reaching the tip of el le CALIFORNIA STALK-EYED CRUSTACEA. 207 the antepenultimate joint. First pair of chelipeds not reaching the middle of the antennal scale; hand oblong, thick, the tips of the fingers hooked. Posterior legs stout, setose; dactyls about half the length of the propodi. Abdomen short, contracted behind the fifth segment; all the segments are rounded above but the third is produced backwards in the middle into a rounded, triangular process; the postero-lateral angles of the fourth and fifth segments are produced into an acute tooth. Telson flat above where it is armed with four pairs of spinules; tip armed with two pairs of spines (the inner pair being the longer) and a short, median spine or tooth. Puget Sound (Stimpson); Marmot Isles, Alaska (Sharp); Monterey! Spirontocaris bispinosus, sp. nov. Carapace with two supraorbital spines, a triangular,tooth on the margin below"the orbit, and a short distance below this a spine behind the base of the antenna; antero-inferior angle of the carapace with an obsolescent spinule. Rostrum a little longer than the carapace, strongly upturned, with the basal half high and laminate, but abruptly contracted near the middle into a slender, styliform process; the rostrum is continued back- ward over the anterior two-thirds of the carapace as a carina; the upper margin is armed with 10-12 teeth which decrease in size and become more closely set anteriorly, there being several small teeth crowded together where the rostrum is abruptly narrowed, the two posterior teeth on the carapace; distal half of the upper margin devoid of teeth; lower margin armed with three or four low teeth on the basal half and a single tooth on the slender process. Eyes pyriform with very large cornez. Basal spine of the antennules reaching the middle of the second joint of the peduncle. Antenne much longer than the body; acicle not nearly reaching the tip of the rostrum; peduncle reaching middle of acicle. Mandibles with a slen- der apical process; palp two-jointed, subequal to the apical process, the first joint broad; second joint narrow, about twice as long as the first. Maxillipeds not quite reaching the tip of the acicle; exognath slender, not nearly reaching the tip of the antepenultimate joint. First chelipeds rather slender, nearly reaching the tip of the antennal peduncle; carpus not distally excavated and longer than the palm of the narrow elongated hand. Second chelipeds exceeding the acicle, carpus seven-jointed, third joint longer than the first two. Posterior legs long and slender, merus armed externally with spines, dactyls very slender. All the abdominal segments are rounded above. Telson rounded above and armed with three pairs of dorsal spines. Puget Sound, June, 1898, five specimens. Collection University of California. 208 CALIFORNIA ACADEMY OF SCIENCES. Spirontocaris lamellicornis (Dana). Hippolyte lamellicornis Dana, Proc. Acad. Nat. Sci. Phila., 1852, p. 24; Crust. U. S. Expl. Expd., Part 1, 1852, p. 567, Pl. XXXVI, fig. 6. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 498. Kines- LEY, Bull. Essex Inst., Vol. X, 1878, p. 62. Spirontocaris lamellicornis Wane, TTADR crepe Biol. Soc., Vol. XII, 1898, pi 277. ' ‘*Beak long, broad, lamellate, peoaoam nearly to posterior margin of thorax, bifid at apex, undulate above, four spines upon cephalothorax and about six upon proper beak, teeth of pines unequal, all nearly equally spaced; outline of beak below triangulately salient, two-dentate. Inner antenne little longer than the beak. A¥iterior feet very slender, but little stouter than the next pair. Second pain hardly shorter than the third, carpus elongate, seven-jointed, third joint quite long. Tarsi of following pairs nearly unarmed, a few very minute spinules towards the base. Outer maxillipeds spinulous at apex, last joint pubescent above. ‘*Length of body one and one-half to two inches. The four dorsal spines are rather larger than those of the beak proper; and the first and last of the latter (not counting the bifid tip), are much smaller than the others; the outline of the whole is separately arcuate along the back and along the beak, with a concave outline between the two parts. The naked tarsus is peculiar. The base of the inner antenna is hardly as long as half the beak, and the flagella scarcely reach beyond the tip of the beak. The second and third joints of the abdomen have the lateral margin trian- gulate or obtusely pointed, and in the next two this margin is acute. The third joint of the carpus of the second pair of feet is twice as long as the first and second jounts together. The six posterior legs are nearly naked. “Dungeness, in the Straits of De Fuca, Northwest America,’ Species not seen. Family PANDALIDA. Rostrum long, slender and spiny. Eyes free. Mandibles with a two- or three-jointed palp. First pair of pereopods simple; second pair chelate, with the carpus annulated. CALIFORNIA STALK-EYED CRUSTACEA. 209 Genus Pandalus Leach. Carapace smooth. Rostrum armed above and below with teeth and continued backward upon the carapace asa carina. Second pair of pereo- pods with the carpi unequal and multiarticulate. Type.—P. annulicornis. LEAcH. Pandalus Dane St. O.TE, $.6/ tbr Pandalus Dane Stimpson, Proc. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 87; Journ. Bost. Soc. Nat. Hist., Vol. VI, p. 502, Pl. XXI, figs. 6 and 7, 1857. Coopsrr, Rep. Expl. and Sur. to Pac. Ocean, Vol. XII, Book 2, 1860, p. 389. Kinestey, Bull. Essex Inst., Vol. X, 1878, p. 64. Smitu, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 214. Ratusvun, R., The Fisheries of the U. S., Sec. ], 1884, p. 821. SHarp, Proc. Acad. Nat. Sci. Phila., 1893, p. 115. Rostrum slender, about as long as the carapace and quite strongly upturned; upper margin armed with 10-12 sharp teeth, the posterior five of which are on the carapace, the last one a little behind the middle, the anterior half of the upper margin of the rostrum smooth; apex, trifid; lower margin armed with six or seven teeth which decrease in size ante- riorly, the posterior tooth large and curved strongly forwards. Subocular spine well developed; antennal spine small. The appendage on the outer side of the basal joint of the antennules is small, lamellate, rounded at the tip and not reaching as far as the middle of the joint; inner flagellum longer than the outer one. Antennal scale narrow, tapering, not nearly so long as the rostrum; two spines on the outer side of the second joint of the peduncle; flagellum longer than the body. The maxillipeds do not reach the tip of the rostrum and may fall short of or exceed the tip of the acicle. First pair of legs shorter than the maxillipeds, the merus pro- duced forwards at the infero-distal angle into a spine. Second pair elon- gated, very unequal, the longer member with a sharp crest on the infero- proximal portion of the ischium; the merus and distal end of the ischium are closely annulated; carpus very long, slender, divided into very numerous annulations and channeled along one side; hand oblong, nar- row, the fingers scarcely as long as the palm; the shorter cheliped is vari- -able in length, exceeding or not equalling the tip of the rostrum; the ischium is not distally annulated and the merus is obscurely annulated, but the annulations of the carpus are distinct; hand similar to that of the longer cheliped. The three following pairs of legs are spinous and rather stout, decreasing in length posteriorly, the dactyls short and spinulous below. The postero-lateral angle of the fifth abdominal segment is rounded, but furnished with a minute tooth; that of the fifth segment 14 210 CALIFORNIA ACADEMY OF SCIENCES. subacute and furnished with a similar tooth. Telson a littie shorter than the uropods, furnished with a shallow superior groove, at the proximal end of which is a small tuft of hairs, and armed above with seven or eight pairs of spinules. Length, 124 mm.; length of carapace including rostrum, 58 mm.; ros- trum, 30.5 mm.; left cheliped, 67 mm,; right cheliped, 42 mm.; third pereopods, 65 mm. Alaska (Sharp); Vancouver’s Is. (Smith); Puget Sound (Cooper); San Francisco Bay (Stimpson)! Pandalus pubescentulus Dana. Pandalus pubescentulus Dana, Proc. Acad. Nat. Sci. Phila., 1852, p. 24; Crust. U. S. Expl. Expd., Part 1, 1852, p. 568, Pl. XXXVI, fig. 8. Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 501. Kryes- LEY, Bull. Essex Inst., Vol. X, 1878, p. 63. Smiru, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 214. *‘Carapax densely very short pubescent, margin below the eye with two spines.. Beak longer than the basal scale of outer antenne; ensiform, somewhat recurved, but apex not raised above level of back, 16-18 toothed above, teeth small and continued nearly to middle of back, towards apex unarmed, apex bifid, below seven-toothed. Feet nearly naked, third, fourth and fifth pair decreasing regularly in length, anterior pair but little longer than the first three joints of second pair. ‘‘ Length of body, 5 inches; of carapax, 24 inches; of beak to the poste- rior tooth on the back, 1% inches; of beak to the orbital sinus, its proper base, 1¢ inches. ‘‘ Straits of De Fuca, at Dungeness, Oregon.” This species has been reported from Vancouver’s Is. by Smith. Pandalus Gurneyi St. Pandalus Gurneyi Stimpson, Ann. N. Y. Lyc. Nat. Hist., Vol. X, 1873, p. 128. Kinesuey, Bull. Essex Inst., Vol. X, 1878, p. 63. ‘“‘A large species, of the same size as P. borealis, etc. Surface of the carapax marked with shallow pits in clusters; not pubescent. Rostrum more than one-half longer than the carapax and unarmed above, except near the base, where the crest has eight or nine teeth, four of which are on the carapax; these teeth are small and rather distant. Below the ros- trum is armed with nine teeth, the two teeth next the base being rather CALIFORNIA STALK-EYED CRUSTACEA. yah close together, large, and hook-shaped, but not broad. The feet of the third pair are rather short, not reaching the extremity of the rostrum; they terminate in well-formed, subcheliform hands. ‘¢ Found at Monterey, Calif., by A.S. Taylor, Esq.” Pandalus franciscorum Kingsley. Pandalus franciscorum Kinestey, Proc. Acad. Nat. Sci. Phila., 1878, p. 94; Bull. Essex Inst., Vol. X, 1878, p. 63. “““ Carapax with a minute pubescence; antennal and branchiostegal spines acute; rostrum a fifth longer than the carapax, extending a fourth its length beyond the antennal scales, considerably recurved, ten or eleven teeth above, of which five are on the carapax and the remainder on the basal portion of the rostrum; distal half of the rostrum above smooth, the apex being minutely bifid or trifid; below with seven to nine teeth, the posterior being the largest. Third joint of the antennular peduncle a third longer than the preceding; flagella about as long as the carapax. Basal joint of the antenne with a spine on the outside and another below; antennal scale long and proportionately narrower than in P. borealis Kroyer; flagellum longer than the body. External maxillipeds falling short of the extremity of the antennal scale. Second pair of feet unequal, the shorter extending further forwards than the external maxillipeds. Posterior pairs stout, armed with spines below. Fifth and sixth abdomi- nal segments with a spine at the infero-posterior angle. Telson narrow, a shallow furrow on its upper surface, apex obtusely triangular. ** Length, 110 mm.; carapax, 52 mm. ‘‘ San Francisco, Calif., W. G. W. Harford.” Family ATYIDE. Mandibles stout and divided, but not deeply so, and devoid of a palp. First two pairs of pereopods chelate, the tips of the fingers furnished with hairs. Rostrum variable. Fresh water forms. Syncaris, gen. nov. Carapace not carinated in front and armed with supraorbital and anten- nal spines. Rostrum long and slender, armed with teeth on one or both sides, and rounded above at the base. Antennules biflagellate, the outer flagellum with a thickened basal portion. External maxillipeds furnished with an exognath. Two or more pairs of pereopods furnished with exo- pods. First pair of chelipeds short and rather stout; carpus short and PA? CALIFORNIA ACADEMY OF SCIENCES. distally excavated to receive the propodus. Second pair of chelipeds longer and more slender than the first, the carpus long and not distally excavated. The three following pairs of pereopods are longer than the- chelipeds; dactyls very short and spinulous below. All the abdominal segments rounded above. Tip of the telson truncated or broadly rounded. Type.—S. pacifica (HOLMES). This genus is intermediate between Atyephyra and. Xiphocaris as defined by Ortmann.’ The latter genus, according to Ortmann, contains besides the type, X. elongata, the Ephyra? compressa of De Haan which Von Martens had placed in Atyephyra and for which Miers. had constituted provisionally the genus Paratya. Mr. Ortmann recognizes but one species of Atyephyra, viz..,. Desmarestit (the Caridina Desmarestii of Joly), which Brito Capello described as Atyephyra rosiana, making the new genus Atyephyra to receive it. Desmaresti being generically distinct from Caridina in the strict. sense becomes, therefore, the type of Atyephyra. Our two species differ from A. Desmarestz in not having the carpus of the second pair of pereopods distally exca- vated, in having the anterior part of the carapace rounded above instead of armed with a dentate carina and in having a prominent suborbital spine. S. Trews approaches Atyephyra, however, in having exopods only on the first two pairs of pereopods instead of the first four as in pacifica. From Xiphocaris elongata both of ° our species differ in having supraorbital spines, in the less number of exopods, and in having the carpus of the first pair of chelipeds excavated. They approach more nearly X. compressa (De Haan), for in that species supraorbital spines are present, though there are exo- pods on all the pereopods, and the carpus of the anterior chelipeds is not hollowed out. 1 Proc, Acad, Nat. Sci. Phila., 1894, p. 399. ——— CALIFORNIA STALK-EYED CRUSTACEA. 4 hy It was owing to the affinities of my pacifica with X. compressa (De Haan) that I was led to put the former species in the genus Miersia, the Hphyra? compressa (De Haan) having been placed in Miersia in Kingsley’s revision of the genera of the Atyide. Mr. Kingsley, however, was wrong in referring compressa to Miersia, for, as Bate has pointed out, the type of that genus, WM. pelagica, is marine and belongs properly in a different family. Syncaris pacifica (Holmes). Miersia pacifica Houmess, Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 577, Pl. XXI, figs. 27 and 28. Rostrum slender and about as long as the carapace; upper margin armed with one or two spines, the lower with 5-9 spines. A pair of supraorbital spines. Antennular flagella subequal, the outer one with a thickened basal portion. Antennal scale reaching about to the tip of the rostrum. All the pereopods except the last pair furnished with exopods. The first pair of legs is short, the carpus short and distally widened, the end exca- vated to receive the basal prominence of the palm. Second pair of legs longer than the first; carpus longer than the hand. The three following pairs are subequal; dactyls short. Telson tapering to a truncated or broadly rounded tip. Length, 5 cm. Sonoma County, Calif., (L. E. Ricksecker coll.) Types in the Museum of the California Academy of Sciences. Syncaris Trewi, sp. nov. Pt.20-, [6% Carapace with supraocular a subocular spines; a tooth at the _ antero-inferior angle. Rostrum slender, slightly upturned, and about one-half the length of the carapace; the upper side is rounded and devoid of teeth, the lower margin armed with 3-5 teeth on its distal half. The basal spine of the antennules reaches slightly beyond the tip of the first joint; flagella subequal and about as long as the carapace exclusive of -the rostrum; the basal third of the outer flagellum is markedly thicker than the distal portion. Antenne ‘nearly as long as the body; acicle oblong, about equalling the rostrum, sides nearly parallel, tip broadly rounded and projecting considerably beyond the spine at the end of the outer © Oita CALIFORNIA ACADEMY OF SCIENCES. margin. Maxillipeds not reaching the tip of the rostrum, the exognath not reaching the tip of the penultimate joint. Anterior chelipeds short; car- pus short and excavated to receive the propodus. Second pair of chelipeds much longer than the first, carpus much longer than the hand and not distally excavated. The succeeding pereopods are longer than the cheli- peds; merus and carpus with a few spines on the outer side; dactyls of the first two pairs about one-fourth the length of the propodi and armed below with numerous spines; dactyls of the last pair longer than those of the preceding pairs and of a different shape, the lower margin finely pec- tinated nearly to the terminal claw behind which are two spines. Telson rounded above and tapering to a broadly rounded or truncated extremity; upper surface armed with two pairs of short spines. Uropods longer than the telson. Length 1.5 in. Described from four specimens collected in a small stream near San Gabriel, Los Angeles County, Calif., by Mr. N. C. Trew. All the exopods in some of the specimens were broken off, but in others they were present on the first two pairs of pereopods. I have found no trace of exopods on any of the posterior pairs; they may have been present and become broken off but I do not think this\is probable. Collection University of California. USNM BLR07T +E Genus Caridina Milne-Edwards. Carapace with a well developed rostrum. Pereopods devoid of exopods. Carpus of the first exopods distally excavated; that of the second pair not excavated. Type.—C. typus Mitng-EDWwaRbs. Caridina pasadene Kingsley. Caridina pasadene Kines.tery, Bull. Essex Inst., Vol. XXVII, 1897, p. 98, Bl, hes. 1-7: Carapace not carinated anteriorly and furnished with a suborbital and an antennal spine. Rostrum three-fourths the length of the carapace, smooth above, bifid at the apex, and occasionally armed with a single tooth on the lower margin. Basal scale of the antennules reaching slightly beyond the tip of the first joint; inner margin of the first joint CALIFORNIA STALK-EYED CRUSTACEA. 215 armed with a small spine. Antenne about two-thirds the length of the body; acicle narrow, reaching slightly beyond the tip of the antennular peduncle. Exognath of the external maxillipeds slender, not reaching the tip of the merus. First chelipeds short, rather stout; second pair twice the length of the first, the carpus much longer than the hand. Posterior pereopods elongated, dactyls curved, spinulous below. Length, 32-39 mm. Anh From streams near Pasadena, Calif., where it is said to becommon. This species bears some resemblance to the form from near the same locality which is here described as Syncaris Trewi. It differs from the latter in having fewer teeth on the lower side of the rostrum, and, according to Kingsley’s fig. 1, in the absence of a supraorbital spine. Kingsley’s fig. 6, which represents a mandible with a two-jointed palp, probably refers to the mandible of Naushonia and not, as is stated in the explanation of the figures, to that of Carzdina. Family PALEMONIDA. Body compressed. Carapace dorsally rounded and furnished with a long, laterally compressed rostrum which is generally armed with teeth. Ocular peduncles well developed. Antennules with the first joint hollowed on the upper surface and furnished with a well developed basal spine; flagella two, one of which is frequently branched. Antenne long, with a long acicle, the outer margin of which terminates in a tooth or spine. Mandibles with a molar tubercle and a cutting edge, with or without a palp. Third maxillipeds pediform. First two pairs of pereopods chelate; the carpus of the second pair not annulated. Genus Palemon Fabdr. Rostrum long, deep, dentate above and below. Frontal margin of the carapace armed with two teeth, the oneabove the other. No hepatic spine. Ocular peduncles pyriform and provided with an ocellus. Antennules with two long flagella, the outer one branched. Mandibles with a three- jointed palp. External maxillipeds slender, with a short exognath. First pair of chelipeds slender; second pair larger than the first; the carpus elongated. 216 CALIFORNIA ACADEMY OF SCIENCES. Palemon Ritteri Holmes. Palemon Ritteri Houmes, Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 579, Pl. XXI, figs. 29-35. Carapace crested on the anterior third or half. A spine beneath the’ angular suborbital projection of the anterior margin of the carapace and another spine behind the base of the antenna and above the rounded in- feriorangle. No hepaticspine. Rostruma little longer than the carapace, tapering gradually from the widest portion to the acute tip; upper margin armed with seven or eight teeth, the posterior one of which is situated on the gastric region; lower margin with three teeth. Antennal scale oblong, shorter than the rostrum. First pair of chelipeds slender, not reaching beyond the tip of the rostrum; carpus devoid of a spine; hand slender. Second pair somewhat larger than the first; carpus not half as long as the preceding joint; hand linear and slightly bent. Length, 4.5 cm. San Diego, Calif. (Dr. Ritter). Genus Anchista Dana. Rostrum long and slender. Eyes prominent. Antennules with one flagellum partly divided. Mandibles devoid of a palp. External maxilli- peds slender. Second pair of pereopods long, slenderand equal. Dactyls slender, long,-and nearly straight. Type.—A. gracilis DANA. - Anchista tenuipes, sp. nov. Rostrum about as long as the carapace, rather deep near the middle, armed above with six or seven teeth, the last one or two on the carapace; lower side armed with three or four teeth; the last dorsal spine is near the anterior third of the carapace. A supraorbital, an antennal and a hepatic spine present; antero-lateral angle of the carapace rounded and devoid of spines. Ocular peduncles large and furnished with an ocellus between the cornea and the proximal part of the stalk. Antennular peduncle shorter than the antennal scale; first joint witha small spine at the antero-external angle, the basal spine small, not reaching the middle of the joint; second and third joints of subequal length; flagella longer than the peduncle, the lower one very slender. Antenns about as long as the body; a spine on the outer angle of the second basal joint; acicle oblong, equalling or exceeding the rostrum. Maxillipeds slender, not nearly reaching the tip of the acicle; exognath reaching considerably beyond the antepenultimate | joint. Anterior pereopods very slender, and reaching scarcely beyond the CALIFORNIA STALK-EYED CRUSTACEA. 217 carpus of the second pair; carpus slightly longer than the merus; hand © very narrow, shorter than the carpus; fingers straight, subequal to the palm. Second pair of pereopods slender and greatly elongated, the carpus reaching beyond the rostrum; ischium, merus and carpus of subequal length, the latter furnished with two teeth at the distal end; hand long, very narrow, subcylindrical, exceeding the length of the carpus and merus combined; fingers slender, straight, about-two-thirds as long as the palm, . the tips hooked and crossed when the fingers are closed. The ambulatory legs are very slender, nearly naked, and subequal. The postero-lateral angle of the fifth abdominal segment is rounded, that of the sixth acute. Telson with two pairs of dorsal spinules. Length of body, 18 mm.; of second chelipeds, 16 mm.; of hand, 7 mm. ; of carpus, 3 mm. Santa Catalina Island, August, 1893; three specimens. Closely allied to A. ensifrons Dana from north of Borneo, but is distinct. Collection University of California. Subtribe PEN AIDEA. Gills dendrobranchiate. Third pair of pereopods and frequently the first and second pairs chelate; the first pair never very large and stout, usually smaller than the others. Fourthand fifth pairs of pereopods always devoid of chelz. Family PEN EID. Carapace generally compressed and produced backwards at the sides. Rostrum compressed and usually continued back upon the carapace as a carina. Eye-stalks generally two-jointed. Antennules biflagellate; first basal joint with an external spine and an unjointed appendage on the. inner side; upper surface excavated. Antennz long, with a large acicle. Mandibular palp never more than two-jointed. Third maxillipeds long and pediform. First three pairs of pereopods chelate and similar. Abdo- men long and compressed, more or less carinated above; the sides of the first segment produced so as to overlap the sides of the carapace in front and those of the second segment behind. Genus Peneus Fabr. Body strongly compressed. Rostrum prominent, serrated. Eye-stalks two- or three-jointed. Flagella of the antennules not longer than the carapace. Mandibular palp two-jointed. First three pairs of pereopods 218 CALIFORNIA ACADEMY OF SCIENCES. ‘ chelate and furnished with small exopods and increasing in length pos- teriorly. No podobranchizw. All the abdominal appendages bear two foliaceous branches except the last pair which, in the male, carries attached to the base a large, membranous appendage, which in the female is reduced to a rudiment. Peneus californiensis, sp. nov. Peneus canaliculatus HOLMES (not OLIVIER), Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 581. fl ze, ¢ 64-67 Rostrum slender. acuminate, slightly arched, the upper margin ciliated and armed with nine or ten teeth, the last tooth on the gastric region and separated by an unusually wide interval from the preceding tooth; lower margin with two small teeth; the rostrum is continued backward upon the carapace as a carina nearly to the posterior end, its dorsal surface sulcate posterior to the last tooth; on either side a prominent groove which ter- minates abruptly a short distance in front of the posterior margin of the carapace; the outer margins of these grooves spread outward near the last rostral tooth and are continued anteriorly npon the lateral ridges of the _rostrum. ‘Above the base of the antenne there is a strong marginal spine which is continued backwards into a carina; above this is a wide, oblique groove, at the posterior end of which is a small hepatic spine; a narrow groove, running obliquely downwards, joins the wide groove immediatély above this spine. Antero-lateral angle of the carapace rounded, with a sinus above. Antennuldr peduncle equalling or a little exceeding the rostrum, the inner appendage of the first joint long, narrow and ciliated; flagella much shorter than the peduncle. Antennal scale reaching about as far forward as the tip of the rostrum; the flagellum may exceed the - length of the body. Chelipeds becoming more slender posteriorly and increasing in length, owing mainly to the increase in the length of the carpus; the second joint in the first two pairs has a spine at the distal end of the lower margin, and there is a second spine in the first pair on the lower side of the ischium; the chelze become longer and narrower pos- teriorly; the first is nearly as long as the carpus, while the third is scarcely _ half as long as that joint; two posterior pairs of pereopods subequal. The three anterior abdominal segments are not carinated above; the fourth is carinated at least on the posterior portion; fifth segment carinated, but not ending posteriorly ina tooth; the carina on the sixth segment is acute, with a groove on either side, and terminates in a tooth at the posterior end. Telson acute, about as long as the preceding segment, devoid of lateral spines, and furnished with a conspicuous dorsai groove, which extends from the anterior end to the tip. ‘The uropods extend far beyond the telson: a os CALIFORNIA STALK-EYED CRUSTACEA. 219 Length from tip of rostrum to tip of telson, 182 mm.; rostrum, 24 mm.; carapace including rostrum, 4] mm.; acicle, 27 mm.; external maxillipeds, ' 45 mm.; first chelipeds, 35 mm.; carpus and hand of same, each 8 mm.; carpus of second chelipeds, 14 mm.; hand of same, 9 mm.; carpus of third chelipeds, 23 mm.; hand of same, 10.5 mm.; sixth abdominal segment, 25 mm.; height of same, 19 mm. Specimens examined, one male from near Anaheim, Calif., and two specimens, male and female, from San Francisco Bay. Closely allied to P. braziliensis but having shorter and stouter legs. This species is also closely allied to canaliculatus, but, so far as I can obtain information concerning the latter species, our California forms differ from it in having a tooth at the posterior end of the sixth abdominal segment, in having the longitudinal ridges on the carapace spreading somewhat in the mid- dle instead of running parallel; the anterior chelipeds are a little shorter than Bate’s figure represents them to be in the typical form of canaliculatus. Possibly the California forms will later be connected with braziliensis or canaliculatus, but it is best, 1 believe, at present to regard them as forming a distinct species. Collection California Academy of Sciences. Suborder STOMATOPODA. Carapace small, leaving the last three or four segments of the thorax uncovered. Rostrum separated from the carapace by a movable joint. Eye-peduncles situated like the antennules on a movable segment. First five pairs of thoracic appendages not biramous, the second pair large and chelate, last three pairs biramous and gressorial. Abdomen large, gener- ally depressed; the first five pairs of pleopods with external gills. Uro- pods large and forming with the telson a strong tail-fin. Genus Pseudosquilla Dana. Abdomen smooth and strongly convex above, the sixth segment not fused with the telson. Telson with well developed submedian spines 220 CALIFORNIA ACADEMY OF SCIENCES. with movable tips, and one to four intermediate denticles between the submedian and intermediate spines. The dactyl of the large chelipeds is not strongly dilated at the base and has few or no marginal spines. Pseudosquilla Lessonii (Guerin). Squilla Cerisiti GUERIN, Voy. Coquille, Crust., 1830, p. 40 (S. Lessoniz on plate). ‘ Squilla spinifrons OwEN, Proc. Zool. Soc. London, 1832, p. 6. Squilla Lessoniti MILNE-Epwarps, Hist. Nat. Crust., Tome II, 1837, p. 527. Waits, List. Crust. Brit. Mus., 1847, p. 84. Squilla monoceros Mitnr-Epwarps, Hist. Nat. Crust., Tome II, 1837, p. 526. NicoLet in Gay’s Hist. Chile, III, Crust., 1849, p. 224. | Pseudosquilla Lessoniti Dana, Crust. U. S. Expl. Epd., Part 1, 1852, p. 662. Muirrs, Ann. Nat. Hist. (5), Vol. V, 1880, p. 113. BigELow, Proc. U.S. Nat. Mus., Vol. XVII, 1894, p. 502. Pseudosquilla marmorata Lockineton, Proc. Cal. Acad. Sci., Vol. VII, 1877, p. 33. Rostrum with acute outer angles and a long, median spine. Hand with three marginal spines; finger with two spines near the base. Sixth abdom- inal segment with four longitudinal carine which end posteriorly in spines. Telson with a median carina which ends posteriorly in a spine and five smaller carine on either side; the posterior margin has three large spines on either side of the triangular, median notch; two spinules between the sub- median spines and the next large spine in front; a singie spinule between the following pair of large spines. Basal prolongation of the uropod end- ing in one large spine with two spines on its inner margin; outer margin of the outer ramus of the uropods armed with spines which increase in size posteriorly, the last spine much exceeding the preceding ones. Chili, S. lat. 00° 46’; W. lon. 89° 42”; Wilmington, Calif. (Bigelow); Lower California; San Diego (Lock- ington)! Santa Catalina Island! Pseudosquilla stylifera (M.-Hdw.). Gonodactylus styliferus MILNE-EpwaRps, Hist. Nat. Crust., Tome II, 1837, p. 530. NIcoLET in Gay’s Hist. Chile, III, Crust., 1849, p. 225. Pseudosquilla stylifera Miers, Ann. Nat. Hist. (5), Vol. V, 1880, p. 112. BicELow, Proc. U.S. Nat. Mus., Vol. XVII, 1894, p. 502, fig. 3. Rostrum longer than broad, acute, but not ending in a spine. Dactyl of the large chelipeds devoid of spines. Sixth abdominal segment longi- tudinally carinated. Telson with a median carina and a lateral one on CALIFORNIA STALK-EYED CRUSTACEA. 221 either side; posterior margin with three large spines on either side of the median notch; a rounded denticle between the submedian and interme- diate spines. Chili; San Pedro (Bigelow); Santa Catalina Island! Point Mendocino, Calif. ! Suborder SCHIZOPODA. Carapace rather small and generally not covering all the segments of the thorax. Eyes stalked. Mandibles generally furnished with an elon- gated palp. Mavxillipeds similar to the succeeding appendages of the thorax which are furnished with well developed exopods. Ova carried ‘beneath the thorax, with or without marsupial plates. Tail-fin well developed. Family MYSID. Carapace small and membranaceous. Pereopods generally of similar form and slender. Branchiz entirely absent. Females with a marsupial pouch, Pleopods generally rudimentary in the female and often soin the male. Inner ramus of the uropods usually furnished with an auditory organ. Genus Mysis Latr. Body siender. Antennal scale setose on both sides and not ending in a spine. Thoracic legs with the propodi subdivided. Fourth pair of pleo- pods in the male with the outer ramus greatly elongated; third pair of pleopods generally unlike those of the female, the remaining pairs simple and rudimentary as in the other sex. Type.—M. oculata Fase. Mysis costata, sp. nov. Le) f. fF Rostrum broadly triangular, enfingina spine. Ocular peduncles short, stout, and slightly flattened horizontally. First joint of the antennular peduncle about one-half ionger than wide; second joint very short; outer flagellum about twice as long as the inner one and furnished with a few long sete near the base; the sensory organ of the male is well developed and furnished with a large, dense tuft of long curved hairs. Peduncle of the antenne reaching about to the middle of the acicle but not reaching the tip of the peduncle of the antennules; second joint with a spine 222 CALIFORNIA ACADEMY OF SCIENCES. external to the base of the acicle; acicle narrow, tapering slightly towards the narrowly rounded tip and ciliated on both margins. Trunk of the mandibles very short and thick; penultimate joint of the palp widest a short distance from the base, whence it tapers to the distal end; second joint narrow, shorter than the preceding one, thickly setose, and having a large, curved, claw-like seta at the tip. The three pairs of maxillipeds end in a claw, the last joint of the third pair five-jointed; the propodi of the succeeding thoracic appendages are 5-7-jointed and end in a claw. Pleopods of the female simple and rudimentary. First three pairs of pleopods in the male simple and rudimentary, asin the female, and of similar form and nearly equal size; fourth pair with the exopod long, slender, smooth and curved, extending backwards beyond the tip of the sixth abdominal segment, the endopod short and furnished with long, plumose sete; fifth pair of pleopods similar to, but slightly longer than the first three pairs. Telson very narrowly triangular, channeled above, and tapering to the narrow tip which is armed with strong spines; the margins are armed with strong spines, between which are several smaller ones. Inner caudal lamella narrow, widened at the base where the audi- tory organ is situated, rounded at the tip, and about equalling the telson; outer lamella about one-fourth longer than the inner one. The first abdominal segment is crossed with three and the following segments with two transverse ridges; first ridge on the sixth segment with a prominent median spine, second ridge with a pair of small median spines, and at the posterior margin of the segment there isa small median spine with a rounded lobe on either side. Two pairs of marsupial lamella. Length, 10 mm. Numerous specimens taken ina tow net at San Pedro, Calif., July, 1896. Collection University of California. Genus Neomysis Czerniavsky. Very closely allied to Mysis, from which it differs in having the long, narrow, antennal scale, which is setose on both margins, end in a sharp Spine. Type.—WV. vulgaris (THoMpPson). Neomysis mercedis Holmes. Neomysis mercedis HOLMES, Proc. Cal. Acad. Sci. (2), Vol. VI, 1897, p. 199, Pl, XIX, figs. 1-10. Rostrum rounded. Acicle narrow, elongated, tapering gradually to an acute tip, and furnished with thickly set, plumose setz on both margins. ™ Ce a A a i i eG CALIFORNIA STALK-EYED CRUSTACEA. 223 The distal portion of the thoracic legs is divided into 8-10 setiferous articulations, the last joint furnished with a claw. In the fourth pair of pleopods in the male the endopod reaches about to the middle of the slender, elongated exopod; terminal joint of the exopod short and bearing two large, sparingly plumose setx at the tip. Telson triangular, tip slightly emarginate, margins armed with several spines. Inner ramus of the uropods widened at the base and tapering to an acute tip which reaches beyond the telson; outer ramus much longer than the inner one, the sides subparallel to the rounded tip. Almost colorless, with the exception of several large, irregularly branch- ing pigment spots. Lake Merced, Calif. Collected by Dr. H. P. Johnson. Neomysis franciscorum, sp. nov. Anterior margin of the carapace produced forwards into a wide quadrate lobe with a straight anterior margin and rounded angles. A strong spine at the antero-lateral angles of the carapace, above which the margin is concave. Eye-peduncles large and pyriform. Antennular peduncle reaching about one-third the length of the acicle. First joint of the antenns with a spine at the antero-external angle; peduncle one-fourth the length of the acicle, which is very narrowly lanceolate and ciliated on both margins. Epistome produced anteriorly into a spine. Last joint of the mandibular palp narrow and somewhat shorter than the preceding one which is much widened. Thoracic legs similar, the propodi divided into from ten articulations in the first, to fourteen in the last pair. All the pleopods in the female short and simple. Telson narrow, concave above, and tapering evenly to a point, the margins armed with numerous spines. Inner uropods narrow, tapering to an acute tip, and about equalling the telson. Length, 1.25 in.; length of carapace, 3, in.; of telson, ;°; in. A single specimen taken in San Francisco Bay. This species is closely allied to Neomysis Rayw (Mur- doch) from Point Barrow, but differs in having the telson acute instead of ‘‘ truncated,’ and in having the terminal portions of the thoracic legs divided into a greater number of articulations. Collection University of California. 224 CALIFORNIA ACADEMY OF SCIENCES. Genus Callomysis Holmes. — AY Carapace with rounded angles and a deep posterior emargination. An- tennal scale oblong, the outer margin naked and ending in a spine; inner and distal margins ciliated. Distal portions of the thoracic legs divided into numerous articulations. Pleopods in the female rudimentary but biramous; pleopods in the male short and small except the third pair, the. outer ramus of which is much elongated. Telson armed with lateral spines and having a spinous emargination at the tip. Type.—C. maculata HOLMES. Callomysis maculata Holmes. Callomysis maculata Houtmess, Proc. Cal. Acad. Sci. (2), Vol. IV, 1895, p. 582, Pl. XXI, figs. 37-44. Rostrum subtriangular. Last joint of the peduncle of the antennz about as wide as long; acicle oblong, shorter than the peduncle, the tip truncated. Maxillipeds (first thoracic appendages) with the exopod longer than the endopod, the terminal portion divided into about fifteen articula- tions; the following pair of appendages much like the preceding but with a longer exopod; the endopods of the remaining pairs are longer than the exopods and have the terminal portion divided into 10-13 articulations. Pleopods rudimentary in the female, the rami of the first pair subequal; in the following pairs the outer ramus is minute. In the male the inner rami are shorter than the outer, the exopod of the third pair very long and slender; fourth and fifth pairs subequal and a little shorter than the first two pairs. Telson oblong, the sides armed with about eight strong spines; apical.emargination with several slender spines. Uropods subequal and scarcely exceeding the telson. Nearly transparent, with large black pig- ment spots having numerous, irregularly branching radiations. Length, 15 mm. Trinidad, Calif.; taken on a sandy beach in which the animals burrow as the waves recede. The females taken in June had their marsupial pouches filled with eggs or larve. Genus Heteromysis Smith. Body comparatively short. Carapace emarginate behind, not covering all the segments of the thorax. Eyes small. Antennal scale small, lamel- liform, with both margins setose. Third maxillipeds much larger than CALIFORNIA STALK-EYED CRUSTACEA. 225 the following appendages, propodal joint entire and spinous, terminal joint unguiform; the remaining legs slender, the terminal portion multiarticu- late and devoid of a claw. Pleopods in both sexes small, simple, and sim- ilar. Telson deeply cleft at the apex. Type.—H. formosa SMITH. Heteromysis odontops Walker. Heteromysis odontops WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 278, Pl. XV, figs. 3-6; Ann. Mag. Nat. Hist. (7), Vol. II, 1898, p. 276. Body rather slender; rostrum subacute. Eyes stout, with a tooth on the distal end of the anterior margin of the peduncle. Legs behind the second pair with a terminal segment divided into eight articulations. Telson with the lateral margins slightly concave and armed with about twenty-four spines which extend over the whole length; tip of the telson with a deep, narrowly triangular cleft whose sides are armed with about thirty rather long spines. Inner uropod ‘‘considerably shorter than the outer, with four spines at the proximal end of the inner margin.” The inner margin of the antennal acicle is figured as devoid of setw—a charac- ter in which this species differs from the next. Puget Sound. Heteromysis spinosus, sp. nov. (L.TL, $.73-76. Body quite robust and somewhat depressed. Rostrum triangular, acute. First and third joints of the antennular peduncle of subequal length; second joint very short, the inner side longer than the outer; sensory organ represented by a minute papilla; inner flagellum short, not one-half the length of the outer one. Antennal scale small, oblong-oval, ciliated on both sides and at the rounded extremity; peduncle reaching about to the tip of the acicle, the last joint about three-fourths the length of the preceding one; flagellum about equal to the outer flagellum of the anten- nules. Trunk of the mandibles short and strongly curved; first joint of the palp very short, second long, curved; the third joint much narrower and much shorter than the preceding one. Third maxillipeds large and stout, the penultimate joint with three pairs of spines on the inner mar- gin; claw strong and curved. The succeeding thoracic appendages are slender; the terminal portions divided into 8-10 articulations and ending in a slender claw. The pleopods are uniramous, rudimentary, and of similar form. Telson triangular, with a deep, triangular, terminal notch, 15 226 CALIFORNIA ACADEMY OF SCIENCES. the margins of which are closely set with small spinules; sides of the telson armed with spinules throughout their length. Inner caudal lamella about three-fourths the length of the outer one, but much ionger than the telson and scarcely widened at the base where the small auditory organ is located. Both lamellew are strongly ciliated and rounded at the tip. Length, one-half inch. A single specimen taken at San Pedro, Calif., June, 1896. ; Collection University of California. Genus Mysidopsis Sars. Carapace comparatively small and not covering all the thoracic segments. Antennal scale lanceolate, setose on both edges, with a short apical articu- lation. First and second thoracic appendages powerfully developed; legs unequal, comparatively short and stout, the propodal joint divided into three articulations, the terminal one having a slender setiform claw. Pleopods in the male all natatory. Telson comparatively short, the apex entire or cleft. Type.—M. didelphys (NoRMAN). Mysidopsis elongata, sp. nov. 4,1E, §-77/ - Fo ; t A smail, slender species with along, narrow, somewhat depressed abdo- men. Rostrum rounded. Hye-peduncles reaching about to the middle of the antennal scale. In the stout antennular peduncle of the male the first joint is nearly as wide as long, the second joint much wider than long, and the third somewhat longer than wide and furnished with a well developed sense organ below and a foliate appendage on the inner side. In the more slender peduncle of the female the first joint is over twice as long as wide, the second longer than broad, the third nearly as long as the first and devoid of the appendage found in the male; the inner flagellum in both sexes is shorter than the outer one, which does not attain half the length of the body. Antenne short, the acicle narrow, tapering, but not ending in a spine, and having both edges fringed with plumose set; ped- uncle slender and about two-thirds the length of the acicle. The pleopods of the female are small, slender and uniramous, while those of the male are large, biramous, and of similar form. Sixth abdominal segment about as long as the two preceding ones combined. ‘Telson not one-half as long as the preceding segment and not reaching beyond the middle of the inner ramus of the uropods; its general form is triangular, but the sides converge more strongly in the basal half; the tip is broadly rounded and CALIFORNIA STALK-EYED CRUSTACEA. Prey | armed with numerous, short, closely set spinules, which do not extend in front of the posterior third of lateral margins. The inner rami of the uropods are shorter than the sixth abdominal segment, somewhat widened at the base where the auditory organ is situated, behind which the sides are nearly parallel to the abruptly rounded tip; outer ramus narrow, about , one-half longer than the inner one and abruptly rounded at the end. Several ‘specimens caught at San Pedro, Calif., by Dr. H. P. Johnson, December 29, 1895. Collection University of California. Genus Siriella Dana. -Carapace small, rostrate. Eyes normal. Antennal scale with the outer margin naked and produced distally into a spine. Legs subequal, the propodi entire or only two-jointed and furnished with a transverse row of set on either side of the strong, sharp, falciform claw. Pleopods rudi- mentary in the female, larger and biramous in the male and often fur- nished with two spirally twisted appendages at the base of the inner ramus. Outer ramus of the uropods larger than the inner, having an imperfect articulation near the tip, and the outer margin armed with spines. Telson elongated, the margins spinous and the apex entire. Audi- tory organ well developed. The species of this genus are mostly pelagic and many of them have a very wide range. Siriella pacifica, sp. nov. Rostrum triangular, acute. Eyes large. Peduncle of the antennules reaching nearly to the tip of the antennal scale, the first joint larger than the next two and concave above; second joint wider than long and bear- ing two large plumose setx at the distal end; third joint about twice as long as wide, with a single, large plumose seta near the middle of the inner margin and several similar setz at the tip; inner flagellum about two-thirds the length of the outer, Antennal scale oblong, the straight outer margin ending in a spine; distal margin rounded, produced beyond the outer spine, and furnished, like the inner margin, with long plumose setz. Upper lip subcordate and produced at the anterior end into a long narrow spine. Propodi of the thoracic legs two-jointed with a transverse row of set at the distal end on either side of the base of the dactyl. Dactyls claw-like, tapering to a sharp, curved tip, and having a transverse or oblique row of setz at about the proximal third of their length. 228 CALIFORNIA "ACADEMY OF SCIENCES. Pleopods in the male rather small. Telson narrow, channeiled above, somewhat constricted behind the base, the sides gently rounded and tapering from near the middle to the narrowly rounded tip; the sides and tip armed with spines; three large spines on each margin of the slightly widened base, behind which is a short interval at the constriction devoid of spines, beyond which the margin is closely set with spines to the tip; two somewhat larger spines at the distal end between which are three very short spines and a pair of plumose sete. Inner ramus of the uropods narrow, scarcely expanded at the base, distally rounded, slightly exceed- ing the telson, setose on both margins and furnished with strong spines along the inner side. Outer ramus larger and much wider than the inner, the outer margin armed with (10-14) spines which begin about the anterior . fourth of its length and extend, gradually increasing in size, to the articu- lated portion, where the ramus is somewhat constricted and beyond which the margin is furnished with plumose seta. Length 11 mm. The color of several specimens preserved in formalin is white or a very pale pink. Pigment is scarce but on the posterior marsupial lamella of the females there is a pair of large, irregularly branched pigment spots, the two anterior lamella having each a single pigment spot. Several specimens caught in a tow net at San Diego, Calif., October 15, 1898. A single female specimen taken at Santa Catalina Island, August 3, 1893. The marsupial pouch contained young embryos in speci- mens from both these localities. Collection University of California. This species resembles Siriella Thompsoniz Milne- Edwards, but may readily be distinguished by the two- jointed propodi of the thoracic legs. Family EUPHAUSIID. Carapace rather small, not calcareous. Branchiz arborescent, situated on the cox of the thoracic legs. Thoracic iegs generally similar in form, the hinder pairs somewhat rudimentary. No marsupial lamella. Pereo- pods well developed and natatory in both sexes. Inner ramus of the uropods devoid of an auditory organ. Telson long and slender, with two long, spiniform appendages attached in front of the tip. CALIFORNIA STALK-EYED CRUSTACEA. 229 Genus Thysanoessa Brandt. Carapace rostrate, the antero-lateral angles produced. Eyes large, the peduncles short. Antennules with the first joint flattened and devoid of a dorsal, leaf-like appendage; flagella short. Maxillipeds slender and not greatly elongated. First pair of legs much longer than the rest; merus and carpus elongated; propodus compressed, the margins furnished with strong sete; last joint very short and setose. Penultimate pair of legs very small; endopod two-jointed. Last pair of legs quite rudimentary. Type.—T. longipes BRANDT. Thysanoessa spinifera, sp. nov. LIE, § £1 Rostrum lanceolate from a triangular base, carinated above, the carina extending back some distance upon the carapace. A prominent spine on either side of the base of the rostrum. A small spine a very short dis- tance above the acute antero-lateral angle of the carapace which is pro- duced forwards; a sinus between this angle and the spines at the base of the rostrum. Eyes large, nearly spherical, the cornea occupying most of the surface. Antennules stout, the first joint flattened, about reaching the tip of the rostrum, the antero-external angle armed with a small spine; second joint prismatic, about two-thirds the length of the first but a little longer than the third. Antennz scarcely half the length of the body, the last joint of the peduncle about three-fourths the length of the third, and reaching nearly to the tip of the acicle; acicle narrowly oblong, the sides nearly parallel, the tip reaching the tip of the second basal joint of the antennules; outer margin slightly concave, ending distally in a spine; inner margin slightly convex and furnished with long seta; the distal end is broadly rounded, transverse, and furnished with set# like the inner margin. Mandibles stout; terminal joint of the palp oblong-elliptical, the inner margin setose, the setz increasing in length towards the tip, the terminal seta long and stout. Maxillipeds moderately long; first joint of the endopod short and tapering distally; second joint elongated, slen- der, about as long as the following joints combined; third joint longer than the fourth; last joint tapering, about three-fifths the length of the preceding one, the tip bearing a very long seta. First pair of legs (gnath- opods) comparatively short for the genus, exopod two-jointed, the first joint not reaching the tip of the first joint of the endopod; second joint of the endopod slender and elongated, furnished with fine cilia on one margin; third joint furnished distally: with long sete and about as long as the two following joints combined; fourth joint with both margins fur- nished with strong setz; terminal joint very short, furnished with four or five large setz# and several smaller ones. Second pair of legs with the 230 CALIFORNIA ACADEMY OF SCIENCES. exopod reaching the tip of the first joint of the endopod; third joint of the endopod about equalling the two following; last joint about one-fourth the length of the preceding one. The four succeeding pairs of pereopods are similar to the first but decrease successively in length posteriorly. The penultimate pair of pereopods is very small; the endopod is two- jointed, the tip of the first joint nearly reaching the tip of the exopod. The last pair of pereopods is probably represented by a minute styliform appendage on the last tuft of branchis. Abdomen slender, the lateral angles acute, the first five segments carinated above, the carinz on the fourth and fifth segments ending posteriorly in a long spine; sixth seg- ment rounded above but ending posteriorly in a small spine which is _ curved downwards. Telson long, deepiy channelled above, the anterior fourth with the sides nearly parallel, the posterior portion tapering to the acute tip; subapical spines extending beyond the tip; a pair of marginal spines near the middle of the dorsal surface and another pair above the bases of the subapical spines. Uropods subequal to the telson, the inner ramus narrowly lanceolate; outer ramus much broader and slightly longer than the inner, the outer angle subacute. . Length, 30 mm. Taken near Fort Bragg, Calif., June, 1894. This species may be distinguished from 7’. longipes Brandt by its more slender rostrum, differently shaped antennal scale, by having the third basal joint of the > antennules shorter, instead of longer than the second, and in having the first pair of pereopods considerably shorter. The cornee are nearly round and not divided by a constriction into an upper and a lower portion as in 7. gregaria and macroura described by Sars. Collection University of California. Thysanoessa gregaria Sars. Thysanoessa gregaria SARS, Forhandl. Vidensks. Selsk., Christiania, No. 7, 1883; Challenger Reports, Vol. XIII, 1885, p. 120, Pl. XXI, figs. 8-17; Pl. XXII, figs. 1-30. Rostrum large, triangular, acuté; antero-lateral angle of the carapace produced into a sharp, triangular process; lower margin of the carapace with a small tooth behind the middle. Eyes very large, subglobose, the upper part of the large cornea constricted; an incision between the smaller CALIFORNIA STALK-EYED CRUSTACEA. yak upper portion of the cornea and the peduncle. First basal joint of the antennules elongated, flattened, and armed with a spine at the antero- external angle; second and third joints of subequal length; flagella shorter than the peduncle. Second basal joint of the antennz longer than wide, with a slender spine at the antero-external angle; third joint very short; last two joints very slender, the terminal one not quite reach- ing the tip of the narrow acicle. Legs short, diminishing in length pos- teriorly, the first pair much longer than the others, and geniculated, the terminal joint very short. Abdomen smooth above, none of the segments produced into spines. Telson very narrow, the apex acuminate, the sub- apical spines smooth and reaching a short way beyond the tip; dorsal surface with two pairs of small spines. Inner ramus of the uropods very narrow and about reaching the tip of the telson; outer ramus broader and somewhat shorter than the inner, the postero-external angle acute. Preanal spine in the form of a broad plate, the posterior margin of which is armed with spinules. Length, 18 mm. Numerous specimens taken in tow at Avalon, Santa Catalina Island, August 13, 1893. This is a pelagic species and is very extensively dis- tributed in the Atlantic and Pacific oceans, both north and south of the equator. rE Z) CALIFORNIA STALK-EYED CRUSTACEA. . 793s APPENDIX. A few species have recently been reported as occurring in the area covered by this paper which have not been mentioned in the preceding pages. Some new species also have been added, and certain changes made in nomenclature after the proof of most of the paper had been corrected and returned. For the sake of complete- ness a Short account of these contributions is given in the following notes. Philyra pisum De Haan. Philyra pisum DE HAAN, Fauna Japonica, Crust., 1850, p. 131, Pl. XXXITI, fig.7. ORTMANN, Zool. Jahrb. Abth. f. Syst., Bd. VI, 1892, p. 582, . Pl, XXVI, fig. 16. Cauman, Ann. N. Y. Acad. Sci., Vol. XI, No. 13, 1898, p. 262. This species has been recently reported from Puget Sound by Calman. ‘‘A single male specimen,” says Calman, ‘‘lacking both chelipeds and some of the am- - bulatory legs, is referred to this species. I have been able to compare it with two specimens dredged in Yoko- hama Bay by Professor D’Arcy Thompson, and also with three specimens from the Strassburg Museum iden- tified by Dr. Ortmann (1. c.), and sent to us by the great kindness of Professor L. Déderlein, by whom they were collected in the same neighborhood. The resem- blance in both cases is so exceedingly close that in spite of the imperfection of the Puget Sound specimen I have no hesitation in adding P. pisum to the list of species inhabiting both sides of the North Pacific.” | 234 CALIFORNIA ACADEMY OF SCIENCES. Pagurus Middendorffii Brandt. Pagurus (Hupagurus) Middendorfii BRanpt, Middendorff’s Siberische Reise, Bd. II, Th. 1, 1851, p. 108, Pl. V, figs. 1-16: HLupagurus Middendorffii Stimpson, Journ. Bost. Soc. Nat. Hist., Vol. VI, 1857, p. 482; Proc. Acad. Nat. Sci. Phila., 1858, p. 250. Canman, Ann, N. Y. Acad. Sci., Vol. XI, No. 13, 1898, p. 260. This species has recently been reported from Puget Sound by Calman. It was reported by Brandt from Okhotsk Sea and Sitka, Alaska, and later by Stimpson from Japan. Pagurus splendescens Owen. Pagurus splendescens OwEN, Zodlogy Beechy’s Voyage, 1839, p. 81, Pl. XXV, figs. 1 and la. Hupagurus splendescens CALMAN, Ann. N. Y. Acad. Sci., Vol. XI, No. 13, 1898, p. 260. | This species also has been found by Calman to range as far south as Puget Sound. It is quite different from the other species of Pagurus here described. The cara- pace is very short and broad and much harder than in the other species, the whole surface being ‘‘ granulate, much resembling the back of a toad.” The front is tri- dentate and the rostrum is very long, reaching beyond the middle of the short, stout eye-stalks. The antero-. - lateral angles of the branchial regions are produced forwards into a prominent process. The antenne are longer than the chelipeds and the acicle is acute and ‘‘notched along the upper part.” The chelipeds)are elongated, very unequal in size, hirsute and granulate below; ‘‘above they/bear long rows of small tubercles and reflect hues of green and gold with metallic lustre; a pink hue is also-reflected in some positions,”’ Dactyls of the left hand long and curved downwards. Ambula- tory legs longer than the chelipeds, the dactyls twisted. Kamtschatka (Owen). CALIFORNIA STALK-EYED CRUSTACEA. 235 Spirontocaris spinus (Sowerby). Cancer spinus SOWERBY, Brit. Misc., 1806, p. 47, Pl. XXI. Hippolyte spinus-@wen, Appendix to Ross’s Second Voyage, 1835, p. Ixxxiii; Pl, HE fig. 2. > Spirontocaris spinus Bats, Challenger Reports, Vol. XXIV, 1888, p. 596, Pls. CVI and CVII. - For further references and synonyms, see Batr, l. c. A short, robust species. Rostrum beginning at the posterior end of the carapace as a carina, which is strongly arched and armed with four or five large teeth; the lamina or rostrum proper is high, much shorter than the carapace, more or less truncated at the tip, and armed above with several teeth which decrease in size anteriorly; lower side strongly curved distally and armed with one or more teeth. Two supra-orbital teeth on either side, the posterior one the larger. A prominent antennal spine and a smaller spine at the antero-inferior angle of the carapace. Outer spine at the base of the antennules large, about reaching the tip of the third joint of the peduncle. Acicle extending beyond therostrum. Maxillipeds stout, ex- ceeding the rostrum, the terminal joint flattened, about four times the length of the preceding joint, the tip armed with spines; exognath not reaching the tip of the antepenultimate joint. First pair of chelipeds rather slender. Third carpal joint of the second pair longer than the first and second combined, but not twice as long. Dactyis of the ambulatory legs short and spiny below. Third segment of the abdomen produced backward into a prominent beak. This species has been reported from very near our limits, and as it is very closely allied to S. lamellicornis (Dana), and will probably be found to occur along with that species, this description is inserted in order to dis- tinguish the two forms. I have not seen any represen- tatives of S. spinus from the Pacific coast, but Dana’s ‘description and figures of lamellicornis agree quite closely with specimens of spznus that I have from the north Atlantic. It is possible that the so-called western rep- resentatives of spznus belong to Dana’s species, for the differences between the two forms are not greater than those often found between widely separated members of an extensively distributed species. The third joint of the carpus of the second pair of chelipeds is somewhat 236 CALIFORNIA ACADEMY OF SCIENCES. 4 larger in lamellicornis than in spinus, and the rostrum is also larger. The chief difference, however, between the two species seems to be in the dactyls of the ambu- latory legs; in spinus they are rather stout and strongly spiny below, while in lamellicornis they are more slender and nearly devoid of spines, having only a few minute spinules near the base. | Spirontocaris grenlandica (Fabr. ) Astacus grenlandicus FABRICIUS, Syst. Ent., 1775, p. 416. Hippolyte grenlandica Mirrs, Ann. Nat. Hist. (4), Vol. XX, 1877, p. 62. Sm1tTH, Rep. Prog. Geol. Sur. Canada, 1878-9, B, p. 214; Trans. Conn. Acad. Sci., Vol. V, 1880; p. 85, Pl. X, fig. 2. ~ CammAN, AnneeNee Acad. Sci., Vol. XI, 1898, p. 260. Spirontocaris grenlandica WALKER, Trans. Liverpool Biol. Soc., Vol. XII, 1898, p. 276. This species, which has previously been mentioned as having been reported by Professor 8. I. Smith from Queen Charlotte’s Islands, has been found both by Walker. and by Calman to occur in Puget Sound. ‘The carina on the carapace extends about to the posterior margin and is armed with four large teeth; the rostrum is short and very slender, scarcely higher than wide, and pro- jects nearly straight in front; the upper margin is armed with from two to four spines, the lower with two or three. Maxillipeds robust, exceeding the acicle. Anterior chelipeds stout. Ambulatory legs rather stout, spiny, the dactyls short and spiny below. Sides of the seg- ments of the abdomen acute. I have not seen western representatives of this species. Miss M. J. Rathbun has recently made Lophoxanthus — bellus (Stimpson) the type of a new genus Lophopano- peus,! in which are included also Lophoxanthus leuco- manus (Lockington) and de: frontalis (Rathbun). In 1 Bull. Lab. Nat. Hist. State Univ. Iowa, 1898, p. 272. CALIFORNIA STALK-EYED CRUSTACEA. 237 her key to the Cyclometopa of North America’ Miss Rathbun gives the names of three new species of Lopho- panopeus from the Pacific coast, Lophopanopeus Locking- toni and diegensis from San Diego, Calif., and L. Heathii from Monterey. Fuller accounts of these species, it is stated, are soon to appear in the Proceedings of the United States National Museum. Lophopanopeus Heathii has the carpus of the chelipeds smooth, or nearly so, as in fron- talis, but the color of the pollex does not extend back upon the hand. JDvzegensis and Lockingtona have the carpus very rough as in leucomanus. Lockingtoni is said to be distinguished from leucomanus and diegensis by the slightly bilobed carpal joints of the ambulatory legs; diegensis has the carpus of the chelipeds covered with tubercles and not with reticulating ridges as in leucomanus. A new species of Cancer, 0. Jordani from Monterey, is also announced in the same paper. In this species the fronto-orbital width is nearly half the width of the carapace. The carapace is slightly areolated and the antero-lateral teeth are strong and projecting. The carpus of the chelipeds has two spines at the inner angle, the one above the other. Cancer Anthonyz is reported in the key from the region between Monterey and San Diego; it was originally described from Playa Maria Bay, west coast of Lower California.” Miss Rath- bun does not recognize the genus T7'richocarcinus as dis- tinct from Cancer. | Reference may be made to two articles contributed by Prof. J. S. Kingsley to the ‘‘ Synopses of North Ameri- can Invertebrates,” published by the American Natural- ist.° The first article consists of a key to the species of 1 American Naturalist, Vol. XXXIV, Feb., 1900, p. 131. 2Proc. Biol. Soc. Washington., Vol. XI, 1897, p. 111 3 American Naturalist, Sept. and Oct., 1899. 238 CALIFORNIA ACADEMY OF SCIENCES, Caridea of North America. The species occurring — within or near our limits of which figures are given are the following: Alpheus cequalis (=Harfordi); Caridina pasadene; Crangon boreas; Crangon munitellus; Crangon munitus; Crangon franciscorum; Hippolyte affinis; Hip- polyte californiensis; Hippolyte Garimardit; Hippolyte Lay; Hippolyte Phippsir; Hippolyte polaris; Heptacarpus brevirostris, Herdmani and palpator (as Hippolyte); Hip- polysmata californica; Palemon Ritteri; Pandalus Dane; Pandalus pubescentulus; Paracrangon echinatus; Syncaris pacifica (as Acanthephyra). The large cheliped of Alpheus Candei is figured, but whether the specimen from which the figure was drawn came from the eastern or western coast of North America is uncertain. The figure resem- bles the cheliped of Alpheus clamator, and may have been taken from a specimen of that species (see p. 184). The species here described as Syncaris pacifica is placed by Kingsley in the genus Acanthephyra. The type of Acanthephyra, viz., A. armata A. Milne-Edwards? is, however, a marine species belonging properly to a differ- ent family from that which includes Syncaris. In A. armata the abdomen is carinated and the rostrum extends back as a carina upon the carapace; there is no supra- orbital spine, and the mandible is provided with a two- jointed palp. In Syncaris neither the carapace nor the abdomen is carinated, a supra-orbital spine is present, and the mandible, as in the other genera of the Atyide, is devoid of a palp. The second article consists of a key to Astacoid and Thallassinoid Crustacea. Figures are given of the chelipeds of Upogebia pugettensis, Callianassa gigas, lon- gemana and californiensis. 1lAnn. Sci. Nat. (6), T XI, 1881, Art. No. 4, p. 12. 1852. 1864. 1865. 1865a. 1888. 1841. 1853. 1855. 1892. 1892. 1894. 1891. 1893. 1894. 1880. 1892. 1894. 1895. CALIFORNIA STALK-EYED CRUSTACEA. 239 BIBLIOGRAPHY. Acassiz, L. Communication on Cambarus Gambelii. Proc. Acad. Nat. Sci. Phila., Vol. VI, p. 375, Bate, C. SPENCE. 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Lobb, at Monterey in California in the Winter of 1850. Ann. Nat. Hist. (2), Vol. XV, p- 307; also in Proc. Linn. Soc. London, Vol. II, p. 329. Some Remarks on Crustacea of the Genus Lithodes, with a Brief Description of a Species Apparently Hitherto Unrecorded. Proc. Zool. Soc. London, Vol. XXIV, p. 132. WuitkEAves, J. F. On Some Marine Invertebrata from the West Coast of North America. Canadian Naturalist (2), Vol. VIII, p. 464. WIEGMANN, A. Beschreibung einiger neuen Crustaceen des Ber- liner Museums aus Mexiko und Brazilien. Archiv. fiir Natur- geschichte, Bd. II, p. 145. INDEX TO GENERA AND SPECIES OF CALIFORNIA STALK-EYED CRUSTACEA. Names of new genera and species in heawy faced type; synonyms in italics. ACANTHEPHYRA armata..............- 238 PUL ELCSee peee eae ee ell afeteferaifeus ah faincstsesnes-¢ 238 AcanGholithodes) nicccssceces se 600% 113, 120 ASTONCES coc cela Sets tatinsiie sieve mre oie a 120 Acanthonychin®................. Sans 21 Acanthus spino-hirsutus ........2..00+ 67 Achelous panamensis................. 72 MAGINALPESUS Pete yaiatces thstars cietose ous fate eines 71, 72 Albuneide...... Sa a orgs abo te dose De 1038 J ATES LEE (6 ERE eee Ser ry AEE Ine 169, 181 I AUS are Seen BE oie iaadeiiataraetel = 181 SEEM EN UTED CUED UL Scie 01 2/2505 = ol 0 choi, a1 <1 clare) wfeber= 189 SEPARA 3 ciel aces: 01s So) ost 182, 189, 238 DAL DATOS ch ves vaso oie 182, 184, 185, 186 GUND ATITLG io) ae etadee a sisiasers tats «oct 182, 184 californiensis..........-- ,..182, 186 CUTE Ctra Ste Sect Ri aehaylers 182,184, 238 clamator....182, 184, 185, 186, 187, 238 GGUIGACEYVIUSE 6.05305 0'- <0 182, 187, 188 BEV OT LU rte ota! cee x pote SS ae 15 ' Vittata cc\.- se leie « Di OSCE ESOS pm ON 180 Gonodactylus styliferus...........00.--- 220 Happolyte Oe ener Sri Te oe [SEs 2 eerie F 74, 78 Fg aagt erence she See Nees oe Grapsodius..... Tat See 79, 88 POCA ape Pore eine Ree che oles aatoletaicieroreie 195 ee eadiane. NE AOS: Ait aes 84 WHACTORIFID sac ckacdsnaocoseadn onoC 198 Ton aor 16, 74 CalMOrmiensis’ ncn. oe 6 1938, 238 Petite. ech 100 CUOMBhognine aon A Ob Od 00nn DOOODOa.e 195 (GOODS Gaia GOCOSODO LOCO GO DOO. Uade 202 EVADTOUIS PULESCEDB: yo 60 2.05 aceaces 190 iphone ie i oy ERMERIO PARED. ooo lek. ae! casino. cts ale 118 aie a a ea ae a ha ee ENT Te ea ea ne 118 CGT SSS EST a asia iges. sir ar 208 oo hee ee 113, 115 GQUENLONAUCO seo) = tlt te lal 194, 236 Bog a ee A ae ire a 115 Hemphillii...... bb saotc obvogpomaS 196 =) eS a Phe os 119 Lamellicormise rae canes ole fecundus ...... etabatelatetal tate a) tatetad Uetots 153 | pilOsitianus. 3.2555 ace eee eos 155 longirostris......... wees cceecsenes LOU (PPA Fa PINTIKA: n0.0% 30 0senadedeeeee reece: 95 PPBEI WAM a (alnopcin's ace Waele date eters 151 MAUS ncn Taxes Soak sx ere: 95 turgidus....... Sekhar =cpay ek oroeevatererapecot 151 nitida...... Neuse et TO petal ang ce 95 Pagurus ..... wreeeeo ee 132, 134,135, 155 | Paratya...... See re Se FR 212 alasKensis.......... -133, 134,135, 126 | Parthenope punctatissima.......-...... 46 CUE ETO 7 | A A oe AS 133, 134, 186 Parthenopide,.............. Bee iey: i 8 Bernhardus........... 132, 133,134, 136 | Parthenopinea............e.e0..0- eet Be. Brandti . ....-.....-..-2++--.- 134, 1BD | Pelia cee cece cece cee eens 31, 35 GALTEOTNUCNGIA «5 scr sec aae sites 2 135,149 | RIRURTCS oes ene ea ae ob ~. US5 Capillatus .............02.0000. 134,138 | Dae ed, 4 oc satoe Hidodiace 223 bier eG COTETARORUS i. «15c,aboinwre Fee .. 1384, 141 malehioliac. ~. see Res eee Een Ras Day ~.3 i. alasfaria) foam naj absicuesssaiet siete) 134,189 tumida........ RS oo ee 35 Gilli........ 22.00... 00. tees eeeeee LAE 51) GE ee en ee ae a ee A 217 granosimanus ................ 134,146 | Penwidea. ..... EMAea AE 3 hte 102, 217 Hemphillii ................... 134,147 | Penmwus ............ -£t SPREE E eee 217 hirsutiusculus........... 134,143, 144 braziliensis............ Feat iler: 219 Lccrelel ey hig Ge iy ere ee ee 134,148 californiensis ...............-. 218 REGRBEU cet a. > s” “ . SS eee wee A a are > b. ‘ .e4) M w., j J \ , y = . Pep ry _ % oa a Fy A 3 1 19, Oy a mA A Ma ; , A Sewers: & % ee e “ ie : ih f at A nts pels 4 : : « . ” af ee f m ‘ Miee. -\s a OP oe ; Sheet Wiss tes Rares 12") 4 a ‘a ati , , A," ayes Vie ee rR We y ar oe : ioe . tied ‘ s] w) > oahee a rei Wel Nia) be, Lia 1 sas | 0a" a - . . A ar i Bis mr 6/944, 260 Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. 43. 44. 45. 46. 47. 48. 49. 50. 51. 52. 53. 04. 55. 56. 57. 58. 59. 60. CALIFORNIA ACADEMY OF SCIENCES. EXPLANATION OF PLATE III. Alpheus californiensis, large hand. as ef small hand. de equidactylus, large hand. ac sy - small hand. “ equalis, hand. Heptacarpus palpator, carapace. i acicle. a) brevirostris, carapace. “ se acicle. CY Taylori, carapace. a ie acicle. a pictus, carapace. i ‘* acicle. ed paludicola, carapace. oe i acicle. o cristatus, carapace. s ce first ambulatory leg. ob carinatus. . Oc. Papers, Can Acan. Scr. VoLVIL. [Houmes| PLate IIT. HOLMES, DEL. LUE. BRITTON & REY, GL ETL TROT TIE TL OLE LEO LIETD — Fs ATC RR Bee IS Spe Tay Ty RS Le ee ee a See Sete 262 61. . 62. . 63. . 64. . 65. . 66. sey: g. 68. . 69. 70. EY MOM ee ya Wi. 76: nay: 18! 79. ig. 80. 81: CALIFORNIA ACADEMY OF SCIENCES. EXPLANATION OF PLATE IV. Pandalus Dane, carapace. e «¢ acicle. Syncaris Trewi, carapace. Peneus californiensis, carapace. ae i hand and carpus of first pereopods. a ne hand and carpus of second pereopods sf ae hand and carpus of third pereopods - abdomen. a Bg acicle. Mysis costata, telson and outline of uropods. fe sé antenna. a Gs fourth pleopod of a male. Heteromysis spinosus, antenna. os “ third maxilliped. “ ee thoracic leg. aot e telson and outline of uropods. Mysidopsis elongata, antennule of the male. a on antennule of the female. fe ts antenna. us ic telson and uropods. Thysanessa spinifera. LUTH.BRITTIN & REY, SE Homes| Erate IV A Se 61. 62. = i = az ee 2 : oO (=) - af ££ FF am *® BP ap rrAee alae y Nn a aan Y 4 mal, | 54 | anes se” porirep! ms & bbe & & GO | A. maT h eA paPra. | lai mle) ft , Py YTTITL 1) Wee a: pl o 1h 46% m Abpe | : & Tame? —- san, apr! snneime lad a i 4 ) io y@> "Ap ¢ ai = mel Y Yes = Cc © => a shinsphaae = i Sa 4 ; . a; | fe ef re p Ral TT ce f wn | T | a This Pea, Shake a T Aas ? 8 Ra NON gy { wee Pel . ands RAR Seeciiee Am a Tidtt : al aa oe C ai F 4a *AAAQA, f ~-a¢g Lo Maa - aa B ‘acee vl 4 ) af CUCL ne fraaa, ; & Ba ane Aa, “e, pate a Ay 5 aA NT a} ASSET laaaal chp Ts ate 3 i . ’ = ger VR gy 1p Lan re oak Raf * Annee ame ~~ ay — Ne ana’ rn mas 5 ; : 45 ~, f shapes | nT a Nir i Ny, pau ha LTA cer “ ppahisian be gpa’? Nunes p48 98a band ir rit mA LL 4 anes: ] ahac acini =. a PR» pe v1] ay & ~ iv ae se > SeeeeeeEs “ . = ee Wome ow ‘ : - @e@e2 3.36 ‘? ‘ - ony ue ’ ° * . sliefeteas ta 6. Bi Oh O—8 . te ce sk . .