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SYNOPSIS
OF
THE LICHENES
OF NEW ENGLAND, THE OTHER NORTHERN STATES,
AND BRITISH AMERICA.
BY EDWARD TUCKERMAN, A.M.
FELLOW OF THE AMERICAN ACADEMY OF ARTS AND SCIENCES ; CORRESPONDING
MEMBER OF THE BOTANICAL SOCIETY OF EDINBURGH, AND OF THE
ROYAL BOTANICAL SOCIETY OF RATISBON.
CAMBRIDGE :
GEORGE NICHOLS,
BOOKSELLER TO THE UNIVERSITY.
1848.
CAMBRIDGE:
METCALF AND COMPANY,
PRINTERS TO THE UNIVEaSITY.
^^ -t-^^^A•
.^
PREFACE
-r
A^ The study of any group of plants, with a view to the true
relations of the species brought together in it, and their real his-
^ tory, requires the investigation of a great number of individual
^ states, and this often through a series of years. And there is,
-"S perhaps, no family of plants in which extensive and continuous
^ observation is more important than in the Lichenes. The present
work is prepared, therefore, only as an Index and Introduction
3-to the history of the plants described in it ; and it has been
^ my object rather to state results of foreign study, than to offer
the necessarily imperfect conclusions of my own limited re-
\5" search. Use has constantly been made of the Lichenographia
" Europma Reformata of Fries, whose characters of the sections
■^ and genera I have, for the most part, adopted entire, or with
\— only occasional amplification ; and the profound descriptions of
the Lichenographia, as here applied to our species, constitute
almost the whole descriptive value of the present enumeration.
The exceptions to this will, I hope, meet with the indulgence
which the extreme difficulty of any inquiries in a field as much
neglected as this has been, among us, seems to demand.*
* The later works of Fries have also been consulted, and, beside these, prin-
cipally those of Turner & I3orrcr, Eschweiler, and Schsrer, and, for American
*:>ci«>«.^ a
IV PREFACE.
The geographical hmits of the work include the countries
generally north of 40^ N. lat. ; but only occasional reference has
been made to those regions of Arctic America which do not
belong to the British possessions, and I have had, also, less
re|ard to the extreme southern boundary, which is everywhere
characterized by the appearance of southern species. The great-
er, or more northern, portion of this district is naturally distinct
and similar throughout, and its Lichenose vegetation seems, in-
deed, almost represented by that of New England alone. It is
probable, however, that a large proportion of the species in-
habitmg any part of the district, if we except its southwestern
extremity, are described here.
For the particular citations of New York Lichens, my princi-
pal authorities have been the Catalogue of the Plants of JVew York
of Dr. Torrey, 1819, and the Synoptical View of the Lichens of
JVeio York by Mr. Halsey, 1824. For citations of species from
Pennsylvania, &c., besides Dillenius (as revised in Fries's fndex
Dillenianus) I have been almost wholly indebted to the Catalogue
of the Plants of JVorth America of Muhlenberg, 1818, and his
specimens now existing in that part of the herbarium of Acharius
which is at Upsal, as well as in the herbaria of Willdenow and
Floerke at Berlm. For the Canadian and Newfoundland Lichens,
I have availed myself of those described in the Flora of ]\Ii-
chaux, as compared with the specimens in his herbarium at Paris,
the species enumerated by Mr. De la Pylaie in his Voyage a Vile
species chiefly, the Synopsis and the Lichenographia of Acharius. The termi-
nology is wholly that of Fries, and its peculiarity will be found mostly to consist
in a strict etymological use of the whole force of the terms he employs. These
terms are, then, possibly, the best expressions of the knowledge they contain, and,
in this view, as well worth studying as any other part of systematic science, tlie
design of which is to teach, not current names for its objects, but llieir history.
PREFACE. V
de Terre-JVeuve, and the specimens of those collected by him,
Mr. Despreaux, and others, preserved in the Royal Herbarium
at Berlin, in that of Professor Kunth, and in those of the late
Baron Delessert and of Dr. Montagne, at Paris. To the vast
herbarium of Sir W. J. Hooker I am indebted, not only for
numerous Lichens of Canada and Newfoundland, but for a large
collection of the arctic species obtained in the different voyages
of Parry, Franklin, and others ; and to these, and his various
Enumerations, together with those of Mr. Brown, Sir John Rich-
ardson, and Dr. Greville, I owe most of the arctic citations.
The late venerable Mr. Menzies also favoured me with a nearly
entire set of the Lichens collected by him on the Northwest
Coast of America.
The genera separated from Lichenes by Fries, and referred to
his family Byssacece, have not yet been fully studied in this coun-
try ; but these plants are so closely related to Lichenes, that I
have enumerated our ascertained species, as an appendix, at
the end.
The present occasion does not permit me to offer more than
general acknowledgments to the eminent botanists whose kind
consideration has encouraged the progress of this work. But I
cannot conclude it without expressing my indebtedness to the
great kindness and liberality of Sir W. J. Hooker, and of Wil-
liam Borrer, Esq., the learned lichenographer of Britain ; to my
much respected friend, Dr. Klotzsch of Berlin, without whose
liberal assistance in the study of the Floerkean herbarium it could
not have been undertaken ; and to my esteemed friend Dr. Gray,
at whose instance it has been prepared.
Cambridge, i/h February, 1848.
LICH^NES.
Perennial, aerial Algse, vegetating only under the influence of moist-
ure, which is imbibed by the whole surface, propagated by spores
(sporidia), and also by the cells (gonidia) of the green layer.
Thallus (universal receptacle, Ach.) composed of three layers, viz. :
the cortical, the medullary, and the gonimoiis ; evolved from a hypo-
thallus (the elementary state in which the layers are confused, and dis-
cernible afterwards as cylindrical cells, and also as fibres on the under
side of foliaceous Lichenes, and forming the base, closely adnate to the
matrix, in crustaceous ones), typically horizontal or vertical. The
horizontal thallus is either crustaceous (often somewhat lobed at the
circumference or squamulose), or foliaceous (becoming sometimes in
degenerate states crustaceous). The vertical thallus is either com-
pressed {suifoUaceous), or terete {fruticulose) ; of both of which the
Jilamentous thallus and the pendulous thallus are degenerations. In
Cladonia and Stereocaulon a vertical thallus (podetium) arises from
the primary horizontal thallus, and is itself often besprinkled with a
kind of secondary horizontal thallus in the form of leaf-like scales. —
Lichenes are reproduced in two ways ; 1. by gonidia, the (normal-
ly green) cells of the green (gonimous) layer, which appear on the
surface as irregularly shaped powdery masses (soredia), and propa-
gate either on the original thallus, forming foliaceous or squamulose
expansions, or external to the original thallus, forming new individ-
uals of the parent thallus ; and 2. by sporidia, consisting of sub-
globose >or elliptical cells, which are either naked or contained in
other elongated more or less vertical cells (asci), and immersed in
the ihalaraium (or fructification proper), and propagate new indiyid-
4 LICHENES OF THE NORTHERN STATES
uals of the species. The thalamium is either rounded, gelatinous-
waxy, and tlie asci converging (nucleiform) ; or flattened at length into
a rigid, persistent, or afterwards collapsing lamina {subdisciform) ;
or originally disciform (open); and is itself contained in a receptacle
[exciple), either of the same color with and like the thallus [thalline
exciple), or of different color and nature {proper exciple). The whole
fructification constitutes the apothecium, which is typically round, though
also occurring normally oblong and linear [lirellaform), and is either
excavated with a contracted margin {urceolate) ; or slightly concave
with an elevated margin (scuteUiform) ; or very concave-scutelliform
{cyalhiform) ; or very concave-scutelliform and pervious {infundibuli-
form, a term applied also to the pervious cup-bearing podetia of Cla-
doniae) ; or goblet-shaped and stipitate {craleriform) ; or dilated, flat,
and without prominent margin {peltcBform, of which the reniform is
a variation) ; or convex with repressed margin (cephaloid) ; or be>
tween scuteUiform and peltseform {disciform) ; or between scuteUi-
form and cephaloid {tuber culate). When the thalline exciple is pro-
longed below into a footstalk, it is said to be pedicellate ; a proper
exciple in like manner prolonged is said to be stipitate. When Ihe
proper exciple is originally and typically closed, the apothecium re-
ceives the name of perithecium. In the Angiocarpi several thalamia
are sometimes contained in the same exciple {composite apothecia) ;
and in the Gymnocarpi, in like manner, several disks are sometinies
confluent {symphycarpeous apothecia). The colors of the thallus in
Lichenes are disposed by Fries in four series : — 1. from pale green
becoming glaucous ; 2. from yellowish green becoming ochroleucous ;
3. from dark green becoming fuscous or olivaceous ; 4, from pale
yellow-green becoming lemon- colored. Each series has its peculiar
variations. The glaucous runs into pale green, cerulescent, and
white ; the fuscous into dark green, olivaceous, cinereous, grayish-
fuscous, and dark chestnut ; the ochroleucous into yellowish green
and albescent; the lemon-colored into pale yellow, orange-red, and
vermilion-red.
and british america. o
Synopsis of the Genera.
Div. I. GYMNOCARPI, Schrader, Fries.
Apothecia open, disciferous. Thalamium originally disciform, or
becoming so, contained in a thalline exciple or a proper exciple ; disk
normally persistent, ascigerous ; sometimes originally pulveraceous-
coUapsed.
Tribe I. PARMELIACE^E, Fr. — Apothecia rounded, from con-
cave becoming explanate, scutelliform, rarely peltate. Disk
somewhat waxy, persistent, contained in a thalline exciple.
Subtribe 1. Usnee.!;, Eschw. — Disk open. Thallus subvertical, or
pendulous-sarmentose, centripetal, without apparent hypothallus.
1. UsNEA. Apothecia peltate ; thallus with a solid medullary layer.
2. EvERNiA. Apothecia scutelliform ; thallus fistulous, or with a
cottony medullary layer.
3. Ramalina. Apothecia orbiculate-subpeltate ; disk pale, of nearly
the same color with the thallus.
4. Ceteaeia. Apothecia scutellate-peltate, oblique.
Subtribe 2. Paebielie^;, Eschw. — Disk at first closed, becoming
at length discoid-open. Thallus horizontal, centrifugal, with a
hypothallus.
5. Nephroma. Apothecia reniform, adnate to the lobes beneath.
6. Peltigera. Apothecia peltseform, adnate to the upper side of
the elongated lobes.
7. SoLORiNA. Apothecia adnate to the disk of the thallus.
8. Sticta. Apothecia scutelliform ; thallus with cyphellEe, or discol-
ored spots, on the under side.
9. Parmelia. Apothecia scutelliform ; thallus without veins or cy-
phellee beneath.
10. Thelotrema. Apothecia urceolate-scutelliform, a discrete inte-
rior exciple veiling a rigescent disk.
11. Gyalecta. Apothecia urceolate, an elevated and discrete colored
margin bordering a nigrescent disk.
Tribe II. LECIDEACE/E, Fr. Apothecia rounded, a persistent
disk contained in an open proper e.xciple, which it finally cov-
ers, and becomes convex, cephaloid, and immarginate.
O LICHENES OF THE NORTHERN STATES
12. STEREOCAnLON. Apothecia cephaloid ; podetia mostly solid.
13. Cladonia. Apothecia inflated ; podetia fistulous.
14. B^OMYCEs. Apothecia capitate, globose, immarginate, velate.
15. BiATORA. Apothecia disciform, solid, with a \va.xy (originally
paler) exciple. ,^-r . /'^ if
Stones and moist rocks in alpine districts. Newfoundland, Despreaux
in herb. Deless. ! Bory in herb. Kunth ! fertile. Northward to Arctic
America, Herl. Hook. ! Melville Island, R. Br. (Parry's First Voy.).
3. C. aculeata, Fr. Th. fruticulose, rigid, subfistulous, lacunose-
compressed, very much and irregularly branched, dark-brownish-chest-
nut, branches divaricate, black-spinulose ; apoth. terminal, peltate, den-
ticulate, disk of the same color. Fr. Lichenogr. p. 35. Cornicularia,
Ach.
On the earth in alpine and subalpine districts. White Mountains,
fertile. Northward to Arctic America, R. Br., Hook. .'
4. C. Richardsonii, Hook. Th. subfoliaceous, canaliculate, divari-
cate-bipinnatifid, naked or sparingly black-denticulate, dark-chestnut ;
apoth. marginal, subpedicellate, margin granulate or irregular, disk
yellowish-brown. Hook, in Frankl. Narr. p. 761, & Icon, t. 31.
Barren grounds north of Great Slave Lake, Rich. (herb. Hook. ! &
herb. Grev, !). Prostrate.
5. C. Islandica, Ach. Th. subfoliaceous, sublinear, canaliculate,
ciliate-spinulose, olivaceous-chestnut ; apoth. obliquely scutellate, ad-
nate to the upper side of the lobes, very entire, disk dark-chestnut.
AND BRITISH AMERICA. 15
Fr. Lichenogr. p. 36. — p. plalyna, Fr. ; laciniae broader, flattish,
waved. Fr. I. c. — ■/. crispa, Ach. ; lacinice narrow, crisped, with
connivent margins. Fr. I. c. ^ I . i^.l ^fi loif .
On the earth in alpine and subalpine districts, and at lower eleva-
tions northward, abundant and fertile; y not found elsewhere. Also
degenerate and sterile on hill-sides, and in sandy fields near the coast,
throughout New England. New York, Torrey. Pennsylvania, Mulil.
6. C cucuUata, Kch. Th. subfoliaceous, sinuate-laciniate,ochroleu- t, /)v, /
cous, sanguineous-fuscous at the base, margins connivent and waved ;
apoth. adnate to the under side of the lobes, disk pale-flesh-colored.
Fr. Lichenogr. p. 37.
On the earth in alpine and subalpine districts. White Mountains,
fertile. Northward to Arctic America, Rich.
7. C. nivalis, Ach. Th. foliaceous, erectish, lacunose-reticulate,
lacerate-laciniate, ochroleucous, yellowish at the base ; lacinise canalic- '
ulate-patulous, crisped ; apoth. marginal, crenulate, yellowish-flesh-col-
ored. Fr. Lichenogr. p. 38.
On the earth in alpine and subalpine districts. White Mountains,
fertile. Northward to Arctic America, R. Br. (Scoresby).
§11. Mem.hr anacecB, Fr. Thallus coriaceous-membranaceous,
the sterile fronds subdepressed.
8. C. glauca, Ach. Th. membranaceous, foliaceous, expanded,
sinuate-lobed, ascendant, glaucous (and cinerascent) ; becoming black
on the under side ; apoth. terminal, peltate, dark-reddish-chestnut. Fr.
Lichenogr. p. 38. — «. fertilis, Fr. ; laciniae elongated, channelled,
becoming whitish on both sides, or spotted with white. Fr. I. c. —
j5. sterilis, Fr. ; lacinis shorter, wider, subdepressed, the under side
fuscous-black. Fr. I. c.
Trunks of trees, stones, &c., in mountain forests, and elsewhere;
New England. Northward to Newfoundland, Pylaie.
9. C. sepincola, Ach. Th. membranaceous, foliaceous, ascendant, !''- I
laciniate, from green becoming olivaceous-fuscescent ; paler beneath ;
lacinise plane (the margins sometimes crisped, pulverulent), fertile ones
short ; apoth. adnate to the upper side of the lobes, dark-fuscous. Fr.
Lichenogr. p. 39.
Trees and dead wood. Branches of dwarf firs, with C. pinastri.
White Mountains, fertile. Arctic America, Rich. Hudson's Bay,
Herh. Banks ! Northwest Coast, Menzies !
16 LICHENES OP THE NORTHERN STATES
10. C. ciliaris, Ach. Th. subcoriaceous, foliaceous, reticulate-lac-
unose, greenish glaucous becoming fuscescent ; whitish-fuscescent be-
neath ; lacinise ascendant, crisped, ciliate or black-denticulate ; apoth.
elevated, blackish-fuscous, with a crenate margin. Ach. Syn. p. 227.
Trunks of trees, and old rails, common and fertile ; ascending to
subalpine districts, where it is often very small, and resembles the last ;
New England. New York, Halsey. Pennsylvania, Mulil. !
11. C. lacunosa, Ach. Th. cartilagineous-coriaceous, foliaceous,
round-lobed, rugose-reticulate-cellulose, glaucescent; whitish on the un-
der side, or spotted with white ; lacinise ascending, the margins crenate,
crisped, black-denticulate ; apoth. large, elevated, dark-reddish, entire.
Ach. Meth. 295, t. 5, f. 3, Syn. p. 227. Lichen cavernosus, Menz.
herh. — /?. Ailantica, Tuckerra. ; cartilagineous-membranaceous, lacu-
nose-reticulate ; apoth. at length perforate. C. lacunosa, Hals. Lich.
N. Y., I. c. & Auct. Amer. C. Tuckermanii, OaJces in Sill. Jour.
Trunks of trees, and old rails. — a. Northwest coast, Menzies ! —
§, Lake Superior to New England, fertile. New York, Halsey. Penn-
sylvania, Mulil. The plant of Menzies differs from ours considerably,
but more specimens of the Oregon Lichen are required, to settle the
distinctness of the two.
12. C. placorodia, Tuckerm. Th. subcartilagineous, foliaceous, of
narrow, at first smooth and discrete, at length convex, concrete, and
plicate lobes, finally besprinkled with black grains or wholly isidioph-
orous, pale livid-glaucous; on the underside fuscescent, rugose, smooth,
sparingly fibrillose ; lacinife crisped, crenate; apoth. marginal, peltate
on the ascending lobules, from pale fuscous becoming dark chestnut,
with an inflexed crenate margin, at length explanate. Parmelia pla-
corodia, Ach. ! Syn. p. 196.
Trunks (normal), Chelmsford, Russell! and common on rails, when
(like C. ciliaris, C. lacunosa, and others) it assumes a Parmeliaceous
aspect. From Parmelia it appears to me distinct, in its marginal, ob-
liquely affixed apothecia, and its smooth, reticulate-rugose under-side.
Acharius was acquainted only with the rail-Lichen.
13. C. aurescens, Tuckerm. Th. subcoriaceous, foliaceous, plane,
sinuate-lobed, yellowish-green ; beneath whitish with pale fuscescent
fibres ; margins of the lobes elevated, crisped, black-denticulate ; apoth.
large, elevated, chestnut, with a thin crenulate margin.
Trunks and branches of Coniferae, New Hampshire. And old rails,
Massachusetts.
AND BRITISH AMERICA. 17
14. C. Oakesiana, Tuckerm. Th. subcoriaceous, foliaceous, de-
pressed, linear-laciniate, from green becoming yellow ; fuscous on the
under side, with scattered coarse fuscous fibres ; laciniae plane, with
elevated, black-ciliate (or more commonly pulverulent) margins ; apoth.
marginal, elevated, rufous-fuscous, somewhat entire. Tuckerm. Lich,
N. E. in Bost. Jour. Nat. Hist. 1841, p. 445.
Trees and rocks in mountain forests, New England ; fertile.
15. C. viridis, Schwein. Th. membranaceous, foliaceous, round-
lobed, lacunose-reticulate, glaucous-green ; pale yellow on the under
side ; margins waved, black-denticulate ; apoth. chestnut-brown, with
an infle.xed, lobate-dentate margin. Schwein. in Hals. Lich. N. Y. I. c.
p. 16.
Cedars, Massachusetts. New York, Halsey. Certainly very near the
next ; and the Massachusetts Lichen here referred to it is perhaps noth-
ing but a state of C. juniperina, (5.
16. C. juniperina, Ach. Th. membranaceous, foliaceous, ascen-
dant, sublacunose, lacerate-laciniate, bright yellow ; on the under side
subreticulate, of the same color; laciniae concave, crisped, black-den-
ticulate ; apoth. adnate to the lobes in front, disk fuscous, margin cren-
ulate. Fr. Lichenogr. p. 40. C. juniperina, Ach. Syn. p. 226, & C.
Tilesii, Ach. J Syn. p. 228. — /S. virescens, Tuckerm. ; glaucous-green
becoming pale yellowish, pale beneath.
On trees, and on the earth, Arctic America, Rich., Hook. ! — /?,
cedars and other trees, and rails, on the coast of Massachusetts, Rus-
sell ! and southward to New York, Torrey, and Pennsylvania, Muhl.
Our jS can be compared only with the low-country Lichen of the North
of Europe, from which it appears to differ as described. The alpine
European forms, and our own arctic ones, recede variously from this
type.
17. C. pinastri, Sommerf Th. membranaceous, foliaceous, de-
pressed, round-lobed, greenish-yellow ; laciniae plane, not denticulate
(with crisped and powdery margins in the sterile plant) ; ' apoth. mar-
ginal, disk yellowish-brown, margin obtuse.' Fr. Lichenogr. p. 40. C.
juniperina, ^. pinastri, Ach. Tuckerm. Lich. N. E. I. c.
Subalpine shrubs and rocks ; also trees in mountain woods and
swamps, infertile ; New England. Northward to Arctic America,
Rich.
3
18 LICHENES OF THE NORTHERN STATES
V. NEPHROMA, Ach.
Apothecia reniform, plane, not velate, adnale to the under side of
the thallus, with an elevated thalline margin. Thallus membrana-
ceous, softish, somewhat villous on the under side.
Nephroma is constituted a section of Peltigera in the Lichenogra-
phia of Fries, but in his Flora Scanica, 1835, and his Summa Fl. Scand.
1845, these genera are recognized as distinct ; as they are also by Mon-
tagne. Fee (Crypt. Exot. Suppl. p. 8) remarks that they differ also
in their thecae.
1. N. arciicum, Fr. Thallus coriaceous-membranaceous, smooth,
ochroleucous ; on the under side subvillous, becoming black ; fertile
lobules somewhat elongated, erectish ; apothecia dark orange-red.
Peltigera arctica, Fr. Lichenogr. p. 42. N. polaris, Ach. Tuckerm.
Lich. N. E. I. c.
Eocks among mosses, and on dwarf firs, in alpine and subalpine
districts. White Mountains, fertile. Abundant in Newfoundland, and
forming patches of two or three feet in extent, Pylaie ! in herb.
Kunth. Northward to Greenland, Brasen (Fl. Dan.), and elsewhere
in Arctic America, Rich.
2. N. resujiinaiMOT, Ach. Th. cartilagineous-membranaceous, smooth,
from glaucous becoming fuscescent ; pale and downy on the underside,
which is sparingly besprinkled with whitish soredia ; apoth. rufous-^fus-
cous. Ach. Syn. p. 241.
Trunks, often of mountain ash, in mountain forests, luxuriant and
fertile ; New England. New York, Halsey. Arctic America, Rich.
Darker on rocks, where it is frequently quite small.
3. N. parile, Ach. Th. membranaceous, suborbiculate, softish,
livid-fuscous ; on the under side naked, rugulose, dark ; (the lacinife
often sorediiferous, and pulverulent at the margins), fertile lobules
short ; apoth. dark-fuscous. Ach. Syn. p. 242.
Rocks. White Mountains, not uncommon. And on the coast, Mr.
Oakes. Fertile.
4. N. Helveticum, Ach. Th. cartilagineous-membranaceous, some-
what rigid, glaucous-fuscescent ; on the under side tomentose, becoming
black; margins of the lobes and of the apothecia fimbriate-toothed ;
fertile lobules somewhat elongated ; apoth. blackish. Ach. Syn. p. 242.
AND BRITISH AMERICA. 1§
Trees and rocks, fertile, New England. A small rock-form occurs
(N. aspera, Tuckerm. Lich. N. E. 1. c), analogous to a similar one
of N. resupinatum.
VI. PELTIGERA, Hoffra.
Apothecia orbiculate, peltseform, plane, adnata to the upper side of
elongated lobes of the thallus, or more rarely marginal ; with a thin
margin of the thallus. Thallus coriaceous-membranaceous, venose on
the under side.
1. P. malacea, Ach. Thallus spongy, soft, smooth, round-lobed, fus-
cous-cinerascent, clothed 'on the under side with a dense blackish to-
mentum becoming white towards the margins ; apothecia ascendant,
rounded, margin crenulate. Fr. Lichenogr. p. 44.
Mountainous districts ; on the earth and on shrub firs near the limit
of trees, and on rocks at lower elevations. White Mountains.
2. P. aphthosa, Hoffm. Th. coriaceous, smooth, besprinkled with
warts, bright green (and glaucescent) ; reticulated with blackish veins,
and fibrillose on the under side ; apoth. large, eiscendant, round, with
a somewhat lacerate margin. Fr. Liclienogr. p. 44.
Rocks among mosses, and on the earth. Common in mountain for-
ests} New England. New York, Torrey. Pennsylvania, Mm/jZ. North-
ward to Newfoundland, Pylaie; and Arctic America, Rich., R. Br.
3. P. canina, Hoffm. Th. membranaceous, flaccid, scrobiculate,
subtomentose, fuscous-green (and cinerascent, and, hoary) ; the under
side whitish and reticulated with pale fuscous veins; apoth. ascendant,
rounded, at length semi-revolute, vertical. Fr. Lichenogr. p. 45.
On the earth, rocks, and mossy trunks, common in New England.
New York, Torrey. Pennsylvania, Muhl. Northward to Greenland,
Gieseke.
4. P. rufescens, Hoffm. Th. coriaceous, soft, subtomentose, cinere-
ous-virescent (and cinereous, and Tufescent) ; fuscous-fibrillose on the
under side, and reticulated with black-fuscous veins ; lobes rather
narrow, with elevated and crisped margins ; apoth. at length vertical,
oblong, revolute. Fr. Lichenogr, p. 46. Pellidea spuria, Ach. Tuckerm,
Lich. N. E. I. c.
On the earth, rocks, and trunks among mosses ; New England,
Thallus smaller and thicker than in the last.
20 LICHENES OF THE NORTHEKN STATES
5. P. polydactyla, HoflVn. Th. papyraceous, very smooth, shining,
plumbeous-virescent (and gray), on the under side almost naked, re-
ticulated with spongy fuscous veins; (fertile lobules often very numer-
ous ;) apoth. ascending, finally revolute. Fr. Lichenogr. p. 46. — (J.
sailata, Fr. ; margins often crisped (or powdery) ; apoth. at first trans-
versely oblong, at length erect and revolute. Fr. I. c. Pellidea scutata,
Ach. ' -L' r ■• , .; /,
Eocks and trunks among mosses, abundant in mountain forests ;
New England. New York, Halsey. Pennsylvania, Mulil. The va-
riety (9 may be taken for the next species, which has a different thallus.
P. reticulata, Hook. ms. (herb. Borr. I), from the Northwest Coast, is
near this, but apparently a distinct species. I have not seen the de-
scription.
6. P. horizontalis, Hoffm. Th. coriaceous, lacunulose, smooth, fus-
cous-virescent ; the under side reticulated with black veins ; apoth.
transversely oblong, plane, horizontal. Fr. Lichenogr. p. 47. . '/ . .; H
Rocks and trunks, among mosses, less common than the last; New
England. New York, Torrey. Pennsylvania, Muhl. Margins of the
thallus sometimes crisped,and the under side scarcely venose (var. lo-
phyra, Ach.).
7. P. venosa, HoffVn. Th. coriaceous (small), fan-shaped, simple,
green (and cinereous) ; white on the under side, and variegated with
fuscous-black, divaricately branched veins ; apoth. adnate to the thal-
lus, round, horizontal. Fr. Lichenogr. p. 48. •
On the earth, in woods. Pennsylvania, Muhl. New York, Torrey !
Northwest Coast, Menzies !
VII. SOLORINA, Ach. ^
Apothecia suborbiculate, depressed, adnate to the disk of the thallus,
covered originally with a thin membrane, which forms at length an
evanescent margin, ' subgelatinous within.' Thallus coriaceous-mem-
branaceous, foliaceous, venose or lanuginous beneath.
Eschweiler (Syst. p. 21, & Lich. Brasil. in Mart. Fl. Bras. 1833, p. 60)
considers this genus very distinct from Peltigera in the peculiar evolu-
tion of its apothecia. The apothecia of some species of Peltigera are
indeed velate, and this is the case with nearly all, according to Fries ;
but these groups differ also in their thecse, as shown by Eschweiler and
by Fee, and in a somewhat different habit. Montagne (Bet. Zeitung,
AND BRITISH AMERICA. 21
1, p. 476), Flotow (Ibid. p. 613), Fee (Crypt. Exot. 1. c), and J. D.
Hooker (Lich. Antarct. in Hook. Jour. Bot.) have enlarged the present
genus by the addition of some interesting tropical and other species.
1. S. crocea, Ach. Thallus coriaceous, lobed, obscurely green be-
coming cinnamon-colored ; on the under side saffron-colored, with rather
obscure, branched, anastomosing veins; apothecia applanate, immargi-
nate, dark-chestnut. Ach. Syn. p. 8. Peltigera, Fr. Lichenogr. p. 48.
On the earth, Arctic America. Greenland, Dill. North of Point
Lake, Rich. (Herb. Hook. !). V;, /,• zi-
2. S. saccala, Ach. Th. membranaceous, subimbricate, from green
becomiDg greenish-cinerascent ; on the under side whitish and fibrillose ;
apoth. applanate, finally saccate-depressed, blackish-fuscous. Ach. Syn.
p. 8. Peltigera, Fr. Lichenogr. p. 49.
Rocks (limestone). New York, B. D. Greene, Esq. Newfoundland,
Pylaie. Northward to Bear Lake, Herl. Hook. ! Solorina orbiculata,
Menz. herb. I from the Pacific coast, appeared to me a distinct, but I
believe it is an undescribed species.
Vm. STICTA, Ach.
Apothecia scutelliform, adnate to the margin or disk of the thallus,
somewhat oblique, the margin free beneath. Disk at first closed, nu-
cleiform ; becoming at length elevated and explanate. Thallus expand-
ed from a centre, foliaceous, coriaceous-cartilagineous, lobate, villous
on the under side, and having on this side small, regular urceolate cav-
ities {cyphella), or where these are wanting soredia, or discolored
spots.
A mostly tropical genus, with many West Indian and South Ameri-
can species, some of which are represented in the extreme southern
parts of the United States.
1. S. mirata, Ach. Thallus subcoriaceous, reddish-brick-colored ;
on the under side lanuginous, reddish-yellow at the circumference, and
besprinkled with small, irregular, often sorediiform, yellow cyphellae ;~
laciniae rounded, sinuate-cut, the margins undulate, crisped, and yellow-
pulverulent ; ' apothecia marginal, disk plane, fuscous-red, margin in-
flexed.' Delis. Stict. p. 49. .'~'
Among mosses on rocks and trees. (Southern States ! and Texas !
infertile.) Ohio .' The Southern Lichen probably occurs within our
limits.
22 LICHENES OF THE NORTHERN STATES
2. S. crocata, Ach. Th. submembranaceous, scrobiculate, greenish-
glaucous-fuscescent ; on the under side lanuginous, liver-colored at
the circumference, with minute, pale-lemon-colored cyphellse ; laciniae
short, rounded, crenulate, with yellowish-pulverulent margins ; ' apoth.
scattered, fuscous-black.' Delis. Siict. p. 56. .
Rocks among mosses, New England, infertile ; less common in the
Northern mountains. S. Feei, Delis. 1. c. p. 44, from North America,
is perhaps a Southern species.
3. S. sylvatica, Ach. Th. coriaceous-membranaceous, laciniate-lobed,
lacunulose, greenish-fuscous ; tomentose, and subfuscous-cinerascent
beneath, with urceolate, whitish cyphellse ; lobes somewhat truncate,
rounded, crenulate ; ' apoth. marginal, peltate, rufous-fuscous.' Fr.
Lichenogr. p. 51.
Rocks, among mosses. Pennsylvania, Muhl., New York, Halsey. S.
fuliginosa, Ach., differs principally in its round-lobed, rugose fronds,
frequently isidioid-efBorescent, and its (normal) sessile, orbiculate apo-
thecia. The described apothecia of S. sylvatica depend upon the figures
of Dillenius, Wulfen, &c. The species is now unknown in a fertile state.
4. S. quercisans, Ach. Th. cartilagineous, laciniate, plane, pale-ru-
fous-fuscous ; somewhat tomentose, and subfuscous-nigrescent beneath,
with urceolate (membranaceous), whitish cyphellae ; lobes subimbricate,
oblong, rounded, crenulate ; ' apoth. scattered, disk somewhat concavo-
plane, with a thin entire margin.' Delis. Slid. p. 84. Lobaria, Michx.
Pennsylvania, Herb. Montague! Mossy rocks. New York, Russell!
The specimens from Mr. Russell seem to me to differ from S. sylvatica
in the characters indicated by Delise, and to agree with his S. querci-
zans, as they also do with my brief notes on the specimen (from Car-
olina) in herb. Michaux. S. Beauvoisii, Delis. 1. c. p. 83, constituted
on a North American Lichen, seems hardly distinct from the present.
5. S. scrobiculala, Ach. Th. coriaceous,- suborbiculate, Ielx, scro-
biculate, leaden-gray (and glaucescent) ; lanuginous on the under side,
with naked, white spots ; laciniae rounded, somewhat entire (commonly
sorediiferous) ; ' apoth. scattered, from rufous becoming fuscous.' Fr.
Lichenogr. p. 53. ,"f-Y» . p 'b f THf . "» •
Trunks, and rocks among mosses. New England ; infertile. North-
ward to Newfoundland, Pylaie. S. limbata, Ach., a species resembling
this, but with urceolate, true cyphelte, very possibly occurs with us.
6. S. anthraspis, Ach. Th. cartilagineous-membranaceous, lacu-
AND BRITISH AMERICA. Za
nose-reticulate, broadly round-lobed, cinereous-virescent ; rugulose and
somewhat villous beneath, with small, white, sorediiform cyphellse ;
lobes somewhat crenate; apoth. scattered, disk at length convex, black,
and excluding the entire thalline margin. Ach. Syn. p. 233.
On the earth, among mosses ; Northwest Coast, Menzies ! New
York, Halsey. The upper surface resembling that of S. pulmonaria.
7. S. pulmonaria., Ach. Th. coriaceous, lax, lacunose-reticulate,
dark green (and olivaceous) ; tomentose on the under side, with naked,
white spots; lacinice elongated, discrete, sinuate-lobed, retuse-truncate ;
apoth. submarginal, rufous. Fr. Lichenogr. p. 53. Lichen pulmona-
rius, L.
Trunks in mountain forests, fertile. Also on rocks, where various
sterile forms are found. Among these is S. linita, Ach., quoted by
Delise as from the United States, which has occurred at the White
Mountains, with all the features of the Swiss Lichen. New England.
New York, Torrey. Pennsylvania, Muhl. Newfoundland, Pylaie.
8. S. glomerulifera, Delis. Th. coriaceous-cartilagineous, thick, or-
bicular, appressed, smooth, from pale green becoming glaucescent ;
villous on the under side, with scattered, excavated cyphellse (which
are often wanting) ; lacinise elongated, sinuate-lobed ; apoth. scattered,
dark-reddish-chestnut, with a rugose, persistent margin. Delis. Siict.
p. 129. Tuckerm. Further Enum. I. c. Parmelia, Ach.
Trunks of trees, and rocks, fertile ; New England. Pennsylvania,
Muhl. in herb. Willd. ! Northward to Newfoundland, Pylaie. The
green glomerules of the European Lichen always wanting in ours.
Young plants of this species may be taken for the next.
9. S. herhacea, Ach. Th. membranaceous, appressed, smooth, ob-
scurely green (and glaucescent) ; on the under side paler, lanuginous,
the membranaceous, hoary cyphelte rare ; lacinise sinuate-repand,
rounded at the apices ; apoth. scattered, rufous, margin crenulate. Ach.
Syn. p. 341. Parmelia, Ach, Syn. p. 198.
Trunks and rocks. Pennsylvania, Muhl. New York, Torrey, Hal-
sey. Arctic America, Rich.
IX. PARMELIA, Ach., Fr.
Apothecia scutelliform, orbicular, adnate horizontally to the disk of
the thallus, with an equal thalline margin. Disk at first connivent-
24 HCHENES OF THE NORTHERN STATES
closed, somewhat waxy. Thallus expanded horizontally from a centre,
two-sided, of various form, upon a hypothallus. Fr. Lichenogr. p. 56.
Synopsis of the Sections.
Sect. I. The fibrillose hypothallus adnate to the foliaceous thallus,
which is discrete from the matrix.
Subsect. I. Imbkicaeia, Fr. — Apothecia elevated, subpedicellate,
regular ; disk very thin, naked, placed upon the gonimous layer.
Thallus imbricate-foliaceous ; often black-dotted from abortion
of the apothecia. — Sp. 1 - 24.
Subsect. II. Physcia, Fr. — Apothecia at first closed, at length de-
hiscent. Disk thickish, waxy, placed upon the medullary layer,
Thallus normally foliaceous ; ascendant or stellate ; fibrillose
on the under side.
* Thallus normally ascendant, or loosely decumbent ; apoth. some-
what obliquely marginate. — Sp. 25.
** Thallus normally stellate-appressed ; apoth. plane. — Sp. 26 -
33.
Sect. II. Thallus subfoliaceous, at length compacted into a conglom-
erate, subgranulose crust ; arising from a fibrillose (rarely ob-
solete) hypothallus, which is adnate to the matrix.
Subsect. III. Pyxine, Tuckerm. — Apothecia erumpent, at first
closed, palish ; becoming patellffiform, and, with the altered
thalline margin, black ; finally cephaloid, excluding the mar-
gin. Thallus subfoliaceous, imbricate-laciniate, at length crus
taceous-concrete at the centre, on a black, fibrillose hypothal
lus. — Sp. 34.
Subsect. IV. Amphiloma, Fr. — Apothecia erumpent, somewhat cor
onate with an accessory thalline margin. Disk wa.xy, thickish
naked. Thallus foliaceous, somewhat monopliyllous, rounded
at length crustaceous-compact at the centre, placed on a spongy-
pannose hypothallus. — Sp. 35-38.
Subsect. V. PsoROMA, Fr. — Apothecia for the most part two-formed
adnate or immersed ; arising in the one case from the thallus,
with a crenate-thalline margin ; and in the other from the hy
pothallus, with an entire proper margin. Disk waxy. Thallus
of discrete, foliaceous squamules, arising from a common hy
pothallus ; often at the centre, or wholly, concrete in a subgran
ulose crust. — Sp. 39 - 41.
AND BRITISH AMERICA. 25
Sect. in. Thallus crustaceous, lobed at the circumference, or wholly
squamulose-effigurate. Hypothallus smooth, adnata to the ma-
trix, often confused with the thallus.
Subsect. VI. Placodium, Fr. — Apothecia plano-scutelliform, elevat-
ed, disk without proper margin, naked. Thallus as above.
(Thalline margin often colored like the disk.) — Sp. 42-47.
Subsect. VII. Psora, Fr. — Apothecia innate, at first somewhat ur-
ceolate, afterwards scutelliform. Disk with a proper margin
(visible at least in the younger apothecia), normally at first
csesious-pruinose. Thallus as above. — Sp. 48 - 50.
Sect. IV. Thallus crustaceous, uniform. Circumference similar, or
the hypothallus sometimes fibrillose-radiant.
Subsect. VIII. Patellaria, Fr. — Apothecia regular, scutelliform,
sessile, the thalline margin persistent. Lamina of the disk
somewhat plane, without proper margin. Thallus crustaceous,
adnate to an indeterminate, mostly black hypothallus. Disk not
csesious-pruinose. — Sp. 51-66.
Subsect. IX. Ukceolaria, Fr. — Apothecia innate in the crust, or
immersed in protuberant warts. Lamina urceolate, or protu-
berant, verrucoeform, blackish, normally caesious-pruinose, mar-
ginate. Thallus crustaceous ; the whitish hypothallus confused
with the thallus, or often fibriUose and radiant. — Sp. 67-70.
Sect. I. The fibrillose hypothallus adnate to the foliaceous thallus.
Subsect. I. Imbricaria, Fr.
Series 1. G I au c es cen t e s , Fr.
1. P. crinila, Ach. Thallus submembranaceous, suborbicular, glau-
cous-fuscescent (the whole thallus, as well as the apothecia, beset with
isidioid granules and branchlets) ; black and somewhat smooth on the
under side, and here and there black-fibrillose ; lobes plane, with some-
what ascendant, erose-crenate, ciliate margins ; apothecia (imperforate)
marginal, subpedicellate,cyathiform, with a thin, inflexed,crenulate mar-
gin, at length explanate, large. Ach. ! Syn. p. 196. P. perforata, §. Fr.
Trunks, &c., fertile ; New England. New York, Torrey. Penn-
sylvania, Muhl. There appear to be indications of other differences
beside the isidioid efflorescence to distinguish this from P. perforata.
The latter is perfectly normal with us.
4
26 LICHENES OF THE NORTHERN STATES
2. P. perforata, Ach. Th. membranaceous, smooth, greenish-glau-
cescent ; on the under side black, with dark fibres ; lobes rounded, as-
cendant, subcrenate, ciliate ; apoth. large, rufous, elevated, infundibu-
liform ; disk perforate, at length explanate, margin very entire. Fr.
Lichenogr. p. 5S. ' '■
Trees, particularly on the coast, luxuriant and fertile. Also on
stones, &c., in sterile states. Pennsylvania and Virginia (from Bartram
and Mitchell), DHL, the original stations of the Lichen. New Eng
land. New York, Torrey. Northwest Coast, MeJi^ies (Herb. Smith!)
3. P. perlata, Ach. Th. submembranaceous, suborbicular, greeii'
ish-glaucous ; on the under side blackish-fuscous, scarcely fibrillose
lobes rounded, plane, not ciliate ; apoth. elevated, dark red, cyathl
form, at length explanate, margin thin, entire. Fr. Lichenogr. p. 59.
— ^. olivetorum, Ach. ; margins of the lobes elevated, crisped, whitC'
pulverulent. Ach. Syn. p. 198. ■ ' '
Trunks and rocks in mountainous districts, fertile ; and common also
in sterile forms ; New England. New York, Hahey.
4. P. scortea, Ach. Th. subcoriaceous, orbicular, smooth, glaucous-
white ; on the under side black, hispid-fibriUose ; lobes longish, sinu-
ate-crenate, incised ; apoth. rufous-fuscous, margin somewhat entire.
Ach. Syn. p. 197.
Stones and trunks, fertile ; New England. New York, Hahey.
Pennsylvania, Muhl. Less common than the next, with which Fries
unites it.
5. P. tiliacea, Ach. Th. membranaceous, orbicular, smoothish,
glaucous-cinerascent ; on the under side blackish-fuscous, with black
fibres ; lobes sinuate-laciniate, the external ones rounded, crenate ;
apoth. subfuscous, margin veiy entire. Ach. Syn. p. 199.
Trunks, fertile, very common ; New England. New York, Halsey.
Pennsylvania, Muhl. Nova Scotia, Menzies !
6. P. Borreri, Turn. Th. cartilagineous-membranaceous, orbicti-
lar, smoothish, glaucous-cinerascent (with round, marginate soredia);
on the under side fuscescent, fuscous-fibrillose ; lacinia? rounded at the
apices, naked ; apoth. chestnut, margin inflexed, entire. Fr. Lichen-
ogr. p. 60. — (3. rudecta, Tuckerm. ; soredia immarginale ; the whole
thallus beset with isidioid granules and branchlets. P. rudecta, AcL !
Stjn. p. 197. / ' .; , J- «
AND BRITISH AMERICA.
27
Trunks, &c., fertile ; New York, Halsey. — /?, New England. Penn-
sylvania, Muhl. The anamorphous development called by Sommerfelt
Lecidea Parmeliarum, and referred by Acharius to Endocarpon, occurs
not unfrequently in this species, as well as in the next.
7. P. saxatilis, Ach. Th. subcartilagineous, reticulate-lacunose,
glaucous-cinerascent ; black and fibrillose beneath ; lacinise sinuate-
lobed, plane, subretuse ; apoth. dark-chestnut, margin at length cre-
nate. Fr. Lichenogr. p. 61. — « ; lacinias irregularly imbricate, nar-
rower. Ach. Lichenogr. p. 469. — (i. rosceformis, Ach. ; th. orbicular,
lobes wider, besprinkled commonly with elongated, marginate soredia ;
apoth. smaller, less e.xplanate. Ach. I. c. p. 471. — /. omphalodes, Fr. ;
th. smoothish, shining, dark purplish-fuscous, laciniee subtruncate. Fr.
Lichenogr. p. 62. Parmelia omphalodes, Ach. Syn. p. 202. /ci^,p^lf // C
Rocks and stones, and, somewhat less commonly, on trees and rails, ' -" ' .'
fertile ; New England. New York, Torrey. Pennsylvania, Muhl.
Northward to Arctic America, Rich. — y. Arctic America.
8. P. aleurites, Ach. Th. membranaceous, orbicular, contiguous,
rugose-plicate, glaucescent (at length furfuraceous) ; on the under side
pale, with fuscous fibres ; lobes discrete at the circumference, plane,
rounded, cut-crenate ; apoth. dark-fuscous, margin at length crenulate.
Fr. Lichenogr. p. 62. >
Dead wood, and firs, in mountainous districts, fertile; and on rocks,
sterile. The sterile plant is also common on rails, &c., on the coast.
New England. New York, Hahey. Fries refers to this species the
P. obsessa, Muhl. Catal., and Ach. Syn. p. 213.
9. P. lavigata, Ach. Th. membranaceous, suborbicular, smooth,
glaucescent ; black, and fibrillose on the under side ; lacinise multifid,
linear, plane, cut, divaricate (often sorediiferous) ; apoth. chestnut,
margin very entire. Ach. Syn. p. 212. ,
Trunks (very common on beech in mountainous districts), fertile.
10. P. sinuosa, Ach. Th. membranaceous, suborbicular, smooth,
glaucescent ; black, and fibrillose on the under side ; lacinia; linear,
wider at the circumference, sinuate-pinnatifid, the sinuses wide, circu-
lar ; apoth. somewhat plane, fuscous, margin thin, very entire. Ach.
Syn. p. 207. ,, h Tuckerm. Lich. N. E. I. c.
Scaly bark of old pines ; New England.
21. L. mdancheima, Tuckerm. Cr. cartilagineous, areolate-verru-
cose, becoming somewhat lobulate, glaucous-white, confused with the
hypothallus ; apoth. appressed, somewhat plane, disk equalling the very
thin margin, at length convex, scarcely excluding the margin, very
black, polished, and shining.
Trunks ; and very common on rails on the coast of Massachusetts
(Ipswich, Mr. Oakes, Lynn, Hingham, &c.), and occurring on dead
wood at the White Mountains. Disk sometimes a little pallescent, but
the margin always very black.
22. L. sahuletorum, Fr. Cr. cartilagineous, at first contiguous, be-
coming rimose-areolate, granulate and somewhat lobulate, cinerascent
or fuscous, confused with the hypothallus ; apoth. produced from the
crust, horny ; exciple annular, with an evanescent margin ; disk naked,
often fuscescent. Fr. Lichenogr. p. 339. Lichen s. Lecidea muscorum,
Auct. quortmd.
On the earth, decaying wood and mosses, stones, and trees, ascend-
ing to alpine districts ; New England. New York, Torrey. Pennsyl-
vania, Muhl. Arctic America, Rich.
23. L. arctica, Sommerf. Granules of the crust cartilagineous, at
first discrete, papillseform, persistent, fuscescent-cinereous ; apoth. im-
mixed, somewhat immarginate, csesious-pruinose, horny and cinerascent
within. Fr. Lichenogr. p. 342. Sommerf. Suppl. Fl. Lapp. p. 156.
Upon mosses in alpine districts ; White Mountains.
24. L. milliaria, Fr. Granules of the crust at first discrete, fuscous,
and cinEreous-white, often deliquescent and leprous ; apoth. produced
among the granules, globose, somewhat immarginate, naked ; exciple
AND BRITISH AMERICA. 69
cupular ; disk at length rugulose and tuberculate, blackish within. Fr,
Lichenogr. p. 3-12. — «. terresiris, Fr. — jS. soj-a/i'Zis, Fr. — •/. ligni-
aria, Fr. ! Lichen dubius, E. Bot. t. 2347 (e JFV.). L. duhia, Turn.
4- Borr. in Hook. Br. Fl. 2, p. 176. Tuckerm. Lick. N. E. I. c.
On old rails (y), common ; New England.
Tribe III. GRAPHIDACE^, Fr.
XVII. UMBILICARIA, Hoffm.
Apothccia superficial ; an originally closed thalline exciple convert-
ed into a carbonaceous proper exciple, becoming more or less open, of
various form. Disk horny, ascigerous, at length chinky, or gyrose-pli-
cate, with an incurved margin. Thallus horizontal, carlilagineous, fo-
liaceous, somewhat monophyllous, affixed by a central point.
This most natural genus can, perhaps, still be retained in the place
given it in the Liclienographia Europaa, though I have, in pursuance
of Fries's suggestion (1. c. p. 347), confirmed by all the observations
that I have been able to make, preferred to alter the generic character,
and make it indicate more fully the relations of the group. It appears
to me as analogous to Biatora as to Sticta ; and as the former genus is
considered to indicate a Lecideaceous type, irrespective of its approxi-
mations to Parmelia, so Umbilicaria may perhaps be taken as typically
representative of a peculiar (perhaps properly lirellseform, or Graphi-
daceous) type, irrespective of the approach which some of the species
make to the characters of Parmeliacese.
Sect. I. PATELLAT.E. Apothecia orbiculate-patellseform ; disk at length
chinky, plicate, or proliferous-papillate.
1. U. mammulala, Ach. (sub Gyroph.). Thallus membranaceous,
smooth, irregularly round-lobed and somewhat crenate, fuscous-nigres-
cent ; on the under side very black, papillose-granulate, and fibrillose ;
apothecia elevated, orbicult^e; maVgin rather thick; disk plane, chinky,
becoming at length convex, and proliferous-papillate. Gyrophora mam-
mulala, Ach. Syn. p. 67. G. mamiUata, Muhl. Cafal. p. 105.
Rocks. Pennsylvania, ilfu/jZ. (North Carolina, il/;-. Cwr^w .') Very
distinct from the ne.\t.
70 LICHENES OF THE NORTHERN STATES
2. U. Pennsylvanica, Hofftn. Th. coriaceous-membranaceous, pap-
ulose, dark-fuscous ; on the under side papillose-granulate and nigres-
cent ; apoth. elevated, orbiculate ; margin rather thin ; disk plane, but
becoming at length convex, chinky, and plicate. Hoffm. PI. Licit. 3,
J). 5, Sf t. G9, f. 1, 2. Lecidea, Ach. Meth. p. 86. Gyrophora, Acli.
Lichenogr. p. 227. Ach. Syn. p. 67. Hook, in Rich. App. Frcmkl.
Narr. p. 759. U. puslulata, Michx. ! Fl. 2, p. 322, non Hoffm.
Kocks. Mountains of Pennsylvania, Muhl. New York, Halsey.
New England, common, and fertile. Canada, Michaux !
3. v. pustulata,Hofrm. Th. coriaceous, papulose, cinerascent; on
the under side smooth, and reticulate-lacunose ; apoth. appressed, or-
biculate-patellfflform, somewhat simple ; margin obtuse. Fr. Lichenogr.
p. 351. Hook..' Br. Fl. 2, p. 219. Gyrophora, Ach. — ^. papulosa,
Tuckerm. ; apoth. at length subpedicellate, irregularly proliferous-pap-
illate, excluding the margin. Gyrophora papulosa, Ach. Lich. Univ.
p. 226. Ach. Syn. p. 67. U. lavis, Pers. {ex Ach.). Gyroph. Imlla-
ta, Willd. herb. .'
Rocks, a. New York, Halsey. — /S, Nova Scotia, 'used for dyeing
reds and browns'; Gov. Wentworth, 1795, Herb. Smith! New-
foundland, Bory in herb. Kunth ! New York, Torrey. Pennsylvania,
Muhl. ! New England, common and fertile, and ascending to alpine
districts, where it is often smaller, thicker, and glaucous-pruinose. /J
does not seem to afford any constant characters to distinguish it from
the European Lichen but the luxuriant development of the apotbecia.
In the var. papillata, Hampe ! a Cape of Good Hope Lichen, the apo-
tbecia are papillate, and perhaps also by a proliferous growth of the
patellceform apothecium ; but this variety, though in other respects re-
sembling ours, is distinct from it. The small, fruticulose tufts almost
characterizing this species in Europe, which I have also observed in
the Swedish U. vellea, are generally wanting in the American plant,
which is almost always normal and fertile.
4. U. anihracina (SchsEr.), Fr. Th. coriaceous, not papulose, black ;
on the under side smooth and black-pruinose ; apotli. elevated, orbicu-
late-patellseform, simple ; margin tumid, disk somewhat plane and even.
Fr. Summ. Fl. Scand. U. atro-pruinosa, Scha:r. in Ser. Mus. {cit.Fr.).
Fr. Lichenogr. p. 351. Lecidea, Schxr. ! Spicil. 1, p. 104. Lichen
anihracinus, Wulf. — a ; th. smooth and even above. Schcer. I. c. Fr.
I. c. — /?. tessellata, Schser. ; th. above finely rimose-areolate or punc-
AND BRITISH AMEKICA. 71
tate-vernicose, rugose at the central point. Seller. 1. c. Fr. 1. c. —
y. reticuluia, Schaer. ; th. reticulate-rugose above. Schar, I, c. Fr.
1. c.
Rocks in alpine districts. «, Newfoundland, Bory in herb. Willd. !
— y. Bear Lake, and elsewhere in Arctic America, Hook.! (Parry's
Sec. and Third Voy.).
5. U. ■polyphylla, HofTm. Th. coriaceous-cartilagineous, smooth,
corrugated, fuscous-black ; on the under side very black and glabrous ;
' apoth. sessile, at first patellasform, marginate, becoming at length con-
vex, and concentrically plicate.' Fr. Lichenogr. p. 352. Gyrophora,
Hook. ! Br. Fl. 2, p. 217. Lichen, L. Gyrophora glabra, Ach. —
(3. deusta, Fr. ; th. thinner, furfuraceous-flocculose ; somewhat lacunose
and paler on the under side. Fr. 1. c. Umbilicaria deusta, Hoffm.
Gyrophora, Ach. Lichen, L.
Rocks on mountains; a, alpine; — (5, descending. White Moun-
tains, infertile. Northward to Newfoundland, Pylaie, and Greenland,
Herh. Banks .'
6. U. prohoscidea, DC, Stenh. Th. submembranaceous, reticulate-
rugose, olivaceous-fuligineous ; on the under side pale and fibrillose ;
apoth. somewhat elevated, orbiculate-patelteform, becoming at length
convex, very gyrose, or proliferous-papillate, somewhat excluding the
margin. Fr. Lichenogr. p. 354. Gyrophora, Ach. Hook. ! Br. Fl.
2, p. 219. — (5. lomala, Ach. ; th. indurated, complicated, plicate-ru-
gose ; obsoletely fibrillose beneath. Ach. Syn. p. 65. Hook, in Rich.
I. c. p. 758, & Icon, t. 30, f. 4. — y. arclica, Ach. ; th. incrassated,
rugose ; glabrous beneath. Ach. I. c. Fr. I. c.
Alpine and subalpine rocks. White Mountains ; and Chin of Mans-
field in the Green Mountains, fertile. Northward to Arctic America,
Rich.
7. U. cylindrica, Ach. (sub Gyroph.). Th. subcoriaceous, rigid,
smoothish, livid, cinereous-pruinose, ciliated with elongated, rigid, ra-
mose, black fibres (or naked) ; on the underside somewhat pale-ochro-
leucous ; apoth. pedicellate, orbiculate-patelljeform, plane, becoming at
length hemispherical, gyrose-plicate, scarcely excluding the margin.
Gyrophora cylindrica, Ach. Hook.! Br. Fl. 2, p. 21S. Lie/ten, L.
U. prohoscidea, /S, Fr. Lichenogr. p. 356.
Alpine rocks. A single specimen from Bear Lake, Herb. Hook. !
72 LICHENES OF THE NORTHERN STATES
is perhaps referable to this species, which has escaped notice, but prob-
ably occurs within our limits.
8. U. /tirsu/a, Ach. (sub Gyroph.). Th. coriaceous, softish, pulver-
ulent, cinerascent and white ; on the under side from pale-fuscous be-
coming blackish, very hirsute with large, softish, at first pale, branched
fibres (at length subfibriUose-scabrous and black) ; apoth. marginal, ap-
pressed, becoming patellfeform, and at length convex, and subglobose,
gyrose-plicate, with a thin margin. Gyrophora hirsuta, Ach. ! Syn. p.
69. U. vellea, y. hirsuta, Fr. Lichenogr. p. 358. — /S. depressa ; th. at
length rigid ; apoth. somewhat impressed, plane, with a thick margin.
U. vellea, (3. depressa, Fr. I. c. V. depressa, /J. spadochroa, Scliar. !
Tuckerm. Lich. N. E. I. c. {sub Gyroph. spadochroa).
Rocks. Common in mountainous, and ascending to alpine districts.
New England, fertile. Northward to Arctic America, R. Br. The
Nevsr England Lichen does not appear to differ from those of Sweden
and Switzerland, unless, perhaps, in attaining to a larger size, and, like
the foreign ones, is near the U. vellea of Sweden, which differs in its
tumid-marginate, papillate apothecia. Of the last species I have not
seen American specimens, unless, with Schaerer, and in accordance
also with the earlier view of Fries, we consider the present species as
a variety of it.
9. U. Dillenii, Tuckerm. Th. coriaceous, rather rigid, smooth, from
glaucous-fuscescent becoming dark-fuscous ; on the under side black,
and closely hirsute with short, black, crowded fibres (or lacerate, and
papillose-scabrous) ; apoth. convex, at first orbiculate and concentri-
cally plicate, becoming at length lirellale, with a thin (canaliculate)
margin. Lichenoides coriaceum lalissivio folio, SfC, Dill. Muse. p.
545, 4- t. 82, /. 5. U. vellea, Michx. .' Fl. 2, p. 323, ^ Aucl. Amer.
Rocks. Paiqualian Mountain, New Jersey,' J. Bartram (Dill.).
Canada, Micliaux ! Newfoundland, Her J. iJ/o/j/ag-ne .' Pennsylvania!
Muhl. New York, Torrey. Veiy common in New England, and fer-
tile. The apothecia are often abortive (very small, and forming some-
times a continuous black crust) ; but in a single specimen from the
White Mountains they are perfect, and agree with the minute descrip-
tion in Michaux, whose Lichen was certainly the same with that of
Dillenius. The species is widely diffused in North America, and pre-
serves its peculiar features from Newfoundland to the Allcghanies of
Pennsylvania ; contrasting in this respect with the more limited and
AND BRITISH AMERICA. 73
northern U. hirsuta. It was considered certainly distinct, in 1841, by
Montagne. Linnaeus cites the figure of Dillenius under his Lichen vel-
leus, and his description includes also U. hirsuta, the differences in the
apothecia being disregarded ; but the specimen that I saw in the Lin-
nsean herbarium was the L. vellea of Sweden, which I have collected
abundantly in that country, and which seems to me very distinct from
the present.
Sect. II. LiKELLAT^. Apothecia somewhat lirellseform, becoming at
length angulate-patellate, or finally crowded together in a hemispher-
ical, subimmarginate, lirellate tubercle.
10. U. hyperlorea, Hoffm. Th. coriaceous-membranaceous, papu-
lose-rugose, dark-olivaceous-fuscous, and blackish ; on the under side
lacunose, smooth, and fuscous-nigrescent ; apoth. appressed, originally
somewhat lirelteform, at length angular, substellate-multiform, plicate
and papillate, with an apparent margin. Fr. Lichcnogr. p. 353. Gy-
rophora, Acli. Floerk. ! Berl. Mag. cil. Fr.
Alpine and subalpine rocks (and perhaps a flocculose state, /?. deus-
ta, Enum. Lich. N. Amer., descending). White Mountains ; Chin of
Mansfield and other of the Green Mountains, fertile. Arctic America,
Rich. Eocky Mountains, Herb. Hook. ! In separating this section of
the genus from the other, I have endeavoured to indicate the features
of difference that seem, at the first view, to distinguish the lirellate
from the patellate apothecia ; but I am uncertain how far the proposed
characters are constant. The ternary division, incidentally proposed
by Fries (Lichenogr. p. 349), suggested the present; but my present
acquaintance with the species has not enabled me to adopt the former
entire.
11. U. erosa, Hoffm. Th. carlila. Lichen-
ogr. p. 394. — /5 ; cr. less perfect. Fr. ! I. c. C. roscidum, /?. Schcer. !
Tuckerm. Emim. Lich. N. Anier. p. 55. Icon, E. Bot. t. 1540.
Decaying wood (jS), Arctic America, Rich. (Herb. Hook. !).
AND BRITISH AMERICA. 81
Sect. II. Apothecia sessile; without crust; parasitical.
13. C. turbinatum, Pers. Parasitical; e.xciple from globose becom-
ing turbinate, sessile, free, shining-black, the disk impressed, with a
thickish, inflexed margin. Fr. Lichenogr. p. 402. C. sessile, DC.
Turn. 6f Borr. Lick Brit. p. 128. Icon, E. Bot. t. 2520.
On the crust, and in the verrucas of Pertusaria pertusa, Ach. New
England. New York, Torrey. Pennsylvania, Muhl.
XXII. CONIOCYBE, Fr.
Apothecia stipitate, spherical, suberose, without margin, bursting at
the ape.x and becoming at length entirely pulverulent, and concealing
the proper exciple. Thallus crustaceous.
C. nigricans, Fr. Crust very thin, leprous, white ; stipes naked,
from whitish becoming black ; apothecia globose, naked, black. Fr.
Liclienogr. p. SS4.
Rough bark of hemlock and rock-maple ; New England. It is with
hesitation that I refer our plant to the European species, though it ap-
pears to agree with a specimen from Flotow. The genus is at once
distinguishable from the other genera of the tribe, and several other
species, as C. furfuracea, with yellow-pulverulent apothecia, and C.
pallida, with pale, white-pruinose apothecia, not improbably occur
with us.
Div. II. ANGIOCARPI, Schrad., Fr.
Tribe I. SPH^ROPHORACEiE, Fr.
XXIII. SPHiEROPHORON, Pars.
Apothecia terminal, spherical, the thalline exciple at first closed, be-
coming at length lacerate-dehiscent. Nucleus globose, within cottony-
carlilagineous, without powdery with naked, black sporidia. Thallus
vertical, fruticulose, crustaceous-cartilagineous without, solid within.
1. S. compressum, Ach. Thallus fruticulose, whitish, irregularly
branched, compressed, fibrillose-ramulose; apothecia globose-depressed,
11
82 LICHENES OF THE NORTHERN STATES
at length disciform, with a reflexed margin. Fr. Lichenogr. p. 404.
Turn. 8f Borr. Licit. Brit. p. 1 15. Icon, E. Bot. t. 1 14.
Rocks and on the earth in alpine districts. Canada, fertile, Herb.
Hook. ! Arctic America ! Rich.
2. S. ghbiferum (L.), DC. Th. fruticulose, somewhat terete, with
erectish, fibriUose-ramulose branches, chestnut ; apoth. globose, with
an inflexed margin. DC. Fl. Fr. Lichen glohiferus, L. S. Coralloi-
des, Pers. Fr. Lichenogr. p. 405. Turn. ^ Borr. Lich. Brit. p. 110
{excl. (S). Icon, E. Bot. t. 115.
On the earth in alpine and subalpine districts ; and descending,
northward. White Mountains, fertile. Eastport, Maine, Russell !
Newfoundland, Pylaie. Arctic America, Hook. !
3. S. fragile, Pers. Th. densely csespitose, fruticulose, dichoto-
mously branched, somewhat cinereous ; branches terete, fastigiate, na-
ked ; apoth. turbinate-globose, with an inflexed margin. Fr. Lichenogr.
p. 405. Schar. ! Spicil. p. 7. Icon, E. Bot. t. 2474.
Alpine rocks. White Mountains, fertile. Northward to Arctic
America, Hook. Rarely somewhat compressed.
Tribe II. ENDOCARPACE^, Fr.
XXIV. ENDOCARPON, Hedw.
Apothecia included in the thallus, globose ; a membranaceous, thin,
pale thalline exciple inclosing a gelatinous, colored, deliquescent nu-
cleus ; ostioles somewhat prominent. Thallus horizontal, cartilagine-
ous-foliaceous, subpeltate.
1. E. minialum, Ach. Thallus cartilagineous-coriaceous, rigid, pale-
yellowish-fulvescent, becoming cinerascent and glaucous-pruinose ; on
the under side naked, at length somewhat rugose, fulvescent, at length
black ; ostioles somewhat prominent, fuscous-nigrescent. Fr. Lichen-
ogr. p. 'iOS. — /?. complicatum, Schter. ; ca:spitose-polyphyllous ; lobes
ascendant, imbricate and complicate, cinereous ; on the under side
dark-fuscous. Schar. ! Spicil. p. 59. Fr. I. c.
Rocks. New York, Halsey. Pennsylvania, Muhl. Arctic Ameri-
ca, iitc/j. — (S, near water, New England. New York, ifa/sey. Fries,
AND BRITISH AMERICA. 83
as well as Sprengel, refers E. gtaucum, Acli. (North America, Ach.),
to the variety « of the present species. I have not found this variety,
but the next species is near to it.
2. E. Muhlenhergii, Ach. Th. cartilagineous-coriaceous, thick,
from greenish-orlaucous becoming fuscescent, very finely rugose and
somewhat chinky ; on the under side fuscous-black ; ostioles convex.
Ach. Syn. p. 101.
Rocks. North America, Ach. West Point, New York, Russell !
(Cf. Ach. Syn. pp. 101, 103.)
3. E. Jluvialile, DC. Th. cartilagineous-membranaceous, flaccid,
lobed, green, becoming fuscescent when dry ; lobes rounded, somewhat
auriculate-lobulate, on the under side naked, reficulate-rugulose, pale-
fuscous, becoming black ; ostioles somewhat prominent, black- Fr. !
Lichenogr. p. 409. E. miniatum, y. aquaticum, Schcer. I Spicil. p. 60.
jE. Weberi, Ach. — /?. fulvo-fusaim, Tuckerm. ; th. thick, subcoria-
ceous, submonophyllous, with auriculate-lobulate, somewhat infle.\ed
margins, fuscous-fulvescent ; on the under side reticulate-rugose, dark-
fulvous-fuscous becoming black ; ostioles scarcely prominent, dark-red-
dish nigrescent.
Rocks (granite), suffused with water; New England. New York,
Halsey. Newfoundland, Pylaie. — /S, alpine. Lake of the Clouds,
White Mountains, at an elevation of five thousand feet. Fries re-
marks, in comparing the present species with E. miniatum, «, that
monophyllous specimens of the former are always minute ; but in /5
these occur nearly as large as average specimens of the latter. The
very brief indication given by Persoon (Act. Wetterav.) of his E.
Americanum answers to our variety.
4. E. pusillum, Hedw. Th. cartilagineous, squamulose-foliaceous,
smooth, brownish-olivaceous, pale on the under side, arising from a
black, fibrillose hypothallus ; ostioles black, somewhat prominent, per-
tuse. Fr. Lichenogr. p. 411. E. Hedwigii, Ach., ^ E. lachneum 4*
squamulosiim, Ach. (e Fr.).
On the earth, and rocks, especially of the more recent formations.
Pennsylvania, Muhl. New York, Halsey. Apparently wanting in the
granite region of New England.
5. E. latevirens, Turn. Th. thin, membranaceous, irregularly orbic-
ular, somewhat concave, round-lobed, grass-green, margins very entire,
84 LICHENES OF THE NORTHERN STATES
inflexed, the under side white at the edges. E. viride, Ach. Verruca-
ria latevirens, Borr. in E. Bot. Suppl. t. 2658.
On the earth in alpine districts. White Mountains. Arctic Ameri-
ca, Rich. The apothecia are unknown, and the plant is a very doubt-
ful member of the present genus. Fries regards it a metamorphosis
of the squamules of Cladonia.
XXV. SAGEDIA, Ach., Fr.
Apothecia included in the thallus, globose ; nucleus gelatinous, de-
liquescent, and, as well as the membranaceous, thin exciple, becoming
at length blackish ; ostioles discrete, attenuated into a thin neck, and
dilated at the apices, pertuse. Thallus horizontal, subcrustaceous.
S. cinerea, Fr. Crust cinereous, at length, pruinose, somewhat foli-
aceous at the circumference ; on the under side spongy, black ; osti-
oles superficial, spheroidal. Fr. Lichenogr. p. 413. Endocarpon, Pers.
E. iephroides, « ^ jS, Ach. Syn.
(On the earth. Fr.) New York (rocks), HaZsey. We have perhaps
a Sagedia, on rocks, in New England.
XXVI. PERTUSARIA, DC.
Apothecia verrucEeform, formed from the thallus, including (1 — 00)
naked, waxy-gelatinous, colored nuclei. Thallus crustaceous, often
passing into soredia and isidia.
1. P. pertusa, Ach. (sub Porina). Crust cartilagineous, glaucous-
white; apothecia depressed-hemispherical, irregular; ostioles depressed,
discrete, the perfect ones black-papillate. Fr. Lichenogr. p. 420. Po-
rina pertusa, Ach. Lichen pertusus, L. Perlusaria communis, DC.
— * sorediifera; crust sterile, sorediiferous. Fr. I. c. Variolaria sp.
Ach. — ** coccodes ; crust isidioid, papillose-ramulose. Fr.l. c. Isi-
dium coccodes, Ach. — /S. areolata, Fr. ; crust thicker, rimose-areolate,
verrucose, often sterile and sorediiferous. Fr. I. c. Variolaria Flo-
fowiana, Floerk. .' — /. leucosloma, Fr. ; apothecia with white ostioles,
the black papillfE deficient. Fr. I. c. Porina lexicosloma, Ach. —
S. leioplaca, Fr. ; crust very smooth ; apothecia imperfect, chinky-de-
hiscent. Fr. I. c. Porina leioplaca, Ach.
Trunks and dead wood; — .?, stones ; New England. New York
(a, y, and 8), Halsey. Pennsylvania (« and (5), Muhl.
AND BRITISH AMERICA. 85
2. P. faginea. Cr. tartareous-cartilagineous, cinereous-white, the
circumference zonate, often thin, polished, and somewhat bluish ; apoth.
hemispherical, bursting into mealy soredia. Lichen faguieus, L. Sf
And. (e JFV.). Variolaria muJtipuncta, Turn, in Linn. Trans. 9, p.
127, t. 10,/. 1. V. faginea, Floerk. .' P. sorediata, Fr. — i3. orhi-
culata; apoth. lax, esplanate; the nuclei e.tpanded into a submembra-
naceous, denudate, flesh-colored disk, which at length falls out, leaving
the sorediiform verrucse. P. communis, ^. sorediata, c. orhiciilala, Fr.
Lichenogr. p. 422. Variolaria faginea, communis, ^ corallina, Auct.
var.
Trunks, dead wood, rocks, and stones ; New England and westward.
New York, Torrey. Pennsylvania, Muhl. Arctic America, Rich.
The Variolarise have been illustrated most largely by Turner and Bor-
rer, in the Lichenographia Britannica, and by the first-mentioned au-
thor in the Linncean Transactions. That they are sorediiferous states
of various crustaceous Lichens has been shown at great length by
Me^'er, Wallroth, and Fries, and this view is confirmed by the observa-
tions of Eschweiler and of Schserer. To the present species, and the
last, most of our common Variolarice are to be referred.
3. P. papillafa, Ach. (sub Porina). Cr. smooth, chinky, whitish ;
apoth. convex, hemispherical ; ostiole solitary, elevated, papillaeform,
with a rufescent pore. Ach. Syn. p. 111.
Trunks. New England. Pennsylvania, Muhl.
4. P. glohilaris, Ach. (sub Porina). Cr. of very numerous, subglo-
bose, and ramulose, glaucescent granules; apoth. (infrequent) globose,
smooth, with a solitary, impressed, punctiform, black ostiole. Ach.
Syn. p. 112.
Upon mosses, Pennsylvania, Muhl., Ach.
5. P. hymenia. Cr. cartilagineous, pale-sulphureous or grayish, bor-
dered by a black line ; apoth. hemispherical-depressed, with a solitary,
depressed ostiole, or more often dehiscent, marginate, and somewhat
scutelliform, the discoid centre black-dotted. Turn. Sj- Borr. Lich. Brit,
p. 185, sui Thelotr. Lichen hymenius, Ach. Prodr. P. Wulfenii, DC.
Fr. Lichenogr. p. 424. Porina fallax, Ach. Syn.
Trunks. New England. New York, Halsey. Pennsylvania, Muhl.
86 LICHENES OF THE NORTHERN STATES
Tribe III. VERRUCARIACEiE, Fr.
XXVII. CONOTREMA, Tuckerm.
Perithecia mostly solitary, horny, black, at first pertuse, becoming
at length open, with a coarctate, inflexed margin, including a depressed
nucleus, which is elevated at the centre into a somewhat marginate
disk. Thallus crustaceous.
C. urceolalum, Tuckerm. Crust thin, smooth, rugose-rimose, glau-
cous-white, bordered by a black line ; perithecia scattered, at first cov-
ered by the crust, finally superficial, conoidal, white- pruinose. Lecidea
iirceolata, Ach. Lichenogr. p. 671. Ach. Syn. p. 27. Pyrenula ente-
roleuca, Spreng. in Hals. Lich. N. Y. I. c. Thelolrema enteroleuca,
Schwein. in Hals. I. c. Verrucaria enteroleuca., Spreng. Syst. 4, p. 243.
Tuckerm. Lich. N. E. I. c. Icon, Hals. I. c. t. 1, /. 1.
Trunks. North America, Sioartz. {ex Ach.). Pennsylvania, Muhl.
in herb. Willd. ! New York, Halsey. New England, very common.
Probably the Lecanora urceolata of Muhl. Catal., but the above-cited
specimen in the herbarium of Willdenow is without name. The Li-
chen appears to me an aberrant form of the present tribe. Thelotre-
ma ? atratum, Fee Crypt. Exot. t. 13, f. 4, seems to be distinguished
from Thelotrema precisely as the present genus (passing over the other
essential differences) is, by its black proper exciple, but the structure
of the nucleus in the former plant removes it from ours.
XXVIII. VERRUCARIA, Pars.
Perithecia hemispherical-globose, solitary, horny, black, closed, with
a simple or papillceform ostiole ; becoming sometimes at length subscu-
telliform, or rarely inclosed in a thalline verruca. Nucleus gelatinous,
hyaline, deliquescent. Thallus crustaceous.
* Saxicola. Crust somewhat tartareous^
1. V. rwpes^m, Schrad. Crust tartareous-compact, contiguous, whit-
ish ; perithecia (small) entire, globose, somewhat sunk, umbonate with
the naked ostiole, at length collapsing and scutelliform ; nucleus hya-
line. Fr. Lichenogr. p. 436. Hook. Br. Fl. 2, p. 152. V. Schi-aderi,
Ach. Icon, E. Bot. t. 1711, /. 2.
Rocks and stones (limestone). Pennsylvania, Muhl.
AND BRITISH AMERICA. 87
2. V. elceochroa, Tuckerm. Cr. applanate, rimose-areolate, oliva-
ceous ; perith. with a wide base, globose, emerging and conical at the
apex, becoming at length depressed and umbilicate.
Rocks (limestone), Ohio, Mr. Lea ! Apparently related to V. elaei-
na, Borr. (E. Bot. Suppl. t. 2623, f. 2), and V. olivacea, Fr. (Lichen-
ogr. p. 43S), but very different from V. olivacea, Pers. (Borr. 1. c. t.
2596, f. 1), which is a bark-Lichen.
3. V. nigrescens, Pers. Cr. somewhat gelatinous-tartareous, chinky,
fuscous-nigrescent, within white ; perith. entire, globose, covered by the
crust and verrucose-prominent, subpapillate ; nucleus whitish. Fr. Li-
chenogr. p. 43S.
Eocks and stones (limestone). New England. New York, Halsey.
4. V. umbrina, Wahl. Cr. verrucose-granulate, or sraoothish, from
fuscescent at length dark-brown ; perith. entire, globose, somewhat
prominent above the crust, papillate. Fr. Lichenogr. p. 441.
Eocks and stones (granite), near water ; New England. We have
doubtless other sa.xicoline species, but they occur often in imperfect
states, and are easily overlooked. I have an alpine Verrucaria, with
large perithecia, from the White Mountains, but the crust is deficient.
** Cortical a. Crust innate in the matrix, often deficient.
5. V. nitida, Schrad. Cr. innate in the matrix, smooth, greenish,
olivaceous, or fuscous ; perith. entire, covered, becoming at length
somewhat prominent, persistent, ostioles subpapillate ; nucleus fluxile.
Fr. Lichenogr. p. 443. Borr. in E. Bot. Suppl. t. 2607, f. 1.
Trunks ; the hue varying with the different epidermis of the matrix ;
New England. Pennsylvania, Muhl. V. composita, Schwein. in Hals.
Lich. N. Y. 1. c. p. 9, has apothecia clustered, forming dark spots, but
I have not been able to find in my specimens, which agree apparently
with the description, any constant characters to separate it from the
present.
6. V. alha, Schrad. Cr. innate in the matrix, becoming at length
denudate, white ; perith. subglobose, entire, denudate, persistent, im-
mersed at the base, ostiole papillate, or pertuse. Fr. Lichenogr. p. 444.
— ^ ; cartilagineous, smoothish ; perith. smaller. Fr. I. c. V. glabra-
ta, Ach.
Trunks. New England. Perithecia prominent.
7. V. gemmata, Ach. Cr. innate in the matrix, effuse, smoothish,
88 LICHENES OF THE NORTHERN STATES.
white-hoary ; perith. hemispherical, dimidiate (not immersed at the
base), persistent ; nucleus whitish. Fr. Lichenogr. p. 444.
Trunks. New England. New York, Halsey.
8. V. epidermidis, Fr. Cr. innate in the matrix or obsolete ; perith.
dimidiate, the base patent, innate-superficial, at length collapsing, and,
together with the nucleus, applanate-depressed. Fr. Lichenogr. p. 447.
— « ; perith. larger, orbiculate. Fr. I. c. V. analepta, Ach. — ^ ; pe-
rith. larger, elliptical. Fr. I. c. V. Cerasi ^ epidermidis, Ach. —
y; perith. small, punctiform (with the habit of the next species). Fr.
I. c. V. sligmatella, Ach. part.
Trunks, mostly on smooth bark ; New England, and westward.
New York (« and ^), Halsey. Arctic America, Rich.
9. V. punctiformis, Pers. Cr. innate in the matrix or obsolete ; pe-
rith. innate-superficial, semiglobose, subdimidiate, the base infle.xed ;
nucleus globose. Fr. Lichenogr. p. 447. V. sligmatella, Ach. pari.
Trunks on smooth bark ; New England. New York, Torrey. Penn-
sylvania, Muhl. Arctic America, Rich.
10. V. pulla, Ach. Cr. smoothish, blackish-fuscous ; perith. minute,
hemispherical, glabrous, subpapiUate, black within. Ach. Syn. p. 83.
Bark of Dirca palustris, Ach., who compares it with V. carpinea,
which is referred to the last species by Fries.
Tribe IV. LIMBORIACEiE, Fr.
XXIX. PYRENOTHEA, Fr.
Perithecia round, carbonaceous, closed, pertuse at length with a sim-
ple ostiole, and protruding the somewhat gelatinous, bursting nucleus,
finally dehiscent, explanate, and empty. (A disciferous state occurs in
a single species.) Thallus crustaceous.
P. leucocephala, Fr. Crust smooth, glaucescent ; perithecia subglo-
bose, naked, black, coronate with the white, persistent, globuliform nu-
cleus. Fr. Lichenogr. p. 450. — ^. Lecidina, Fr. ; crust somewhat
leprous ; disk dilated-scutelliform, rigescent, covered for the most part
with a dense pale-yellowish-cinereous bloom. Fr. I. c. Lecidea able-
Una, Ach.
Trunks (/?), Arctic America, Rich.
COLLEMACEiE.
Filamentous, or foliaceous gelatinous-conglutinate plants without dis-
crete layers. Sporidia included in asci, and immersed in a thalamium,
which is contained either in a thalline exciple or a proper exciple.
Several genera are included here formerly referred to Lichenes, but
separated by Fries, and with other genera constituted a distinct family,
intermediate between Lichenes and aquatic Algse. Collema and Lep-
togium may be said to have the thallus of Phyceae with the apothecia
of Lichenes, and Ephebe is considered by Fries nearly related to the
Byssi .
Synopsis.
Tribe L COLLEMEjE, Fr. — Thallus gelatinous-conglutinate,
caulescent or foliaceous.
1. Collema. Apothecia scutelliform, with a thalline exciple.
2. Leptogium. Apothecia scutelliform, with a proper exciple.
Tribe II. EPHEBIDEjE. — Thallus filamentous, not gelatinous.
3. Ephebe. Apothecia scutelliform, with a thalline exciple.
Tribe I. COLLEMEJ:, Fr.
I. COLLEMA, Hoffm.
Apothecia at first subglobose, becoming at length discoid-open and
scutelliform, with a thalline exciple. Thallus corneous-gelatinous,
somewhat pulpy, of a moniliform-filamentous texture, variously lobed.
* Thallus imbricate-plicate, becoming thick and turgid when wet.
1. C. pulposum, Ach. Thallus thick, suborbicular, very compact,
blackish-green, of numerous, somewhat imbricate, plicate, rather entire
or repand-crenate, erectish lobes, those of the circumference larger,
somewhat appressed ; apothecia somewhat crowded, slightly concave,
rufous, with an elevated, irregular margin. Ach. Syn. p. 311. Schcer.f
12
90 COLLEMACEa: OP THE NORTHERN STATES
Spicil. 2, p. 538 {sub Parmelia). C. crislalum, Borr. in Hook. Br. Fl.
2, p. 208. Icon, Wulf. in Jacq. Coll. 3, p. 139, t. 12, /. 1.
Upon rocks, among mosses. Pennsylvania, Muhl. I have not ob-
served this species in the granite region of New England.
2. C. plicaiile, Ach. Th. thick, orbicular, black-green ; lobes ru-
gose-plicate, ascending, laciniate ; apoth. concave, of nearly the same
color with the thallus, with a thick, elevated margin. Ach. Syn. p. 314.
Hook. Br. Fl. 2, p. 209. Schar. Spicil. 2, p. 543 {stib Parmelia).
Exs. Schleich. ! Lich. Helv.
Eocks (limestone, Schser.), New York, Russell ! I have seen only a
small fragment, but it appears to belong to this rather than to the pre-
ceding species.
3. C. tenax, Ach. Th. rather thick, suborbicular, glaucous-green,
of somewhat plane, rounded, cut, or crenate lobes ; apoth. scattered, at
first urceolate, becoming rather elevated, concave, rufescent, with a
somewhat entire margin. Ach. Syn. p. 314. — (?. pallescens, Ach.;
th. yellow-virescent, pale beneath, the lobes irregular, densely compli-
cated, irregularly crenate, ascending ; apoth. submarginal. Ach. I. c.
Rocks among mosses, Pennsylvania, Muhl. ! New York, Spreng.!
The cited specimens belong probably to the variety (S of Acharius, but
were not considered to differ from a by Floerke.
4. C. fasciculare, Ach. Th. suborbicular, imbricate-plicate, olive-
green, the lobes dilated upward, waved, those of the circumference
rounded, cut-crenate ; apoth. marginal, at length elevated-subpedicel-
late, fasciculate, the disk somewhat convex, rufous. Ach. Syn. p. 317.
Fr.! Lich. Suec. 50. Icon, E. Bot. t. 1162.
Trunks and rocks, New England. Pennsylvania, Muhl.
5. C. pustulatum, Ach. Th. substellate, lacerate-laciniate, the la-
cinise expanded, plane, irregularly crenate, besprinkled above with paler
pustules which pass at length into apothecia ; disk punctiform, black.
Ach. Syn. p. 351. Parmelia leucoderrna, Willd. herb. 7
Upon mosses. North America, Ach., who says it is a minute species,
very distinct from the last. Pennsylvania (P. leucoderrna, Willd.),
Muhl. !
6. C. granulatum, Hook. Th. foliaceous, membranaceous, corru-
gated, granulated on both sides, imbricate-complicate, blackish-olive,
the lobes somewhat rounded, waved and crisped, rather entire ; apoth.
AND BRITISH AMERICA. 91
scattered, sessile, blackish-fuscous, margin entire. Hook. Br. Fl. p. 2,
211. Lichen granulatus, Huds. (e Hook.). Colhma furvtim, Ach.
Exs. Schar. ! Lich. Heh>. 413, 414. Icon, E. Bol. t. 1757.
Stones and trunks. New England. New York, Hdlsey. Pennsyl-
vania ? Muhl.
•• Thallus thin, foliaceous, gelatinous-membranaceous, lobed princi-
pally at the circumference.
7. C. melcmum, Ach. Th. foliaceous, somewhat stellate, blackish-
green, the lobes elongated, radiant, multifid, with elevated, waved and
crisped, crenate margins ; apoth. submarginal, somewhat plane, at length
rufescent, with a subgranulate margin. Ach. ! Syn. p. 315. — /S. jaco-
haifolium, Ach. ; lacinise narrow, strict, lacerate-pinnatifid ; apoth. mar-
ginal, with an entire margin. Ach. I. c. Exs. Schar. ! Lich. Helv. 422.
Rocks among mosses and trunks. New England.
8. C. palmatum, Ach. Th. csspitose-pulvinate, brownish-green, of
crowded, erectish, palmate-divided, somewhat linear and terete laciniae ;
apoth. rufous-fuscous. Ach. Syn. p. 319.
On the earth, and trunks. Pennsylvania, Muhl.
9. C. nigrescens, Ach. Th. somewhat monophyllous, membrana-
ceous, expanded, round-lobed, rugose-plicate, olivaceous-nigrescent;
costate-lacunose beneath ; apoth. (minute) central, crowded, at length
convex, rufous-fuscous, with a very entire margin. Ach. Sy/i. p. 321.
Hook. Br. Fl 2, p. 211. Exs. Schar. ! Lich. Heh. 410.
Eocks and trunks. New England. New York, Halsey. Pennsyl-
vania, Muhl.
10. C. Jlaccidum, Ach. Th. foliaceous, membranaceous, smooth,
flaccid, blackish-green ; lobes somewhat ascending, rounded, rather en-
tire, undulate-plicate ; apoth. scattered (small), somewhat plane, rufous.
Ach. Syn. p. 322. Hook. Br. Fl. 2, p. 211. Exs. Schcer. ! Lich. Helv.
412.
Rocks and trunks. New England.
11. C. tUTueforme, Ach. Th. foliaceous, membranaceous, somewhat
rugose, blackish-green, besprinkled with fuliginous powder; lobes ob-
long, deeply cut, sinuate-laciniate, obtuse, flexuous, crisped, subcrenate ;
apoth. scattered, somewhat plane, fuscous, with a very entire margin.
Ach. Syn. p. 322.
92 COLLEMACE^ OF THE NORTHERN STATES
Rocks (especially limestone, Ach.). Pennsylvania, Muhl. in herb.
Willd. ! The specimen appeared to me to agree with an original one
from Acharius. Schserer refers the species to C. flaccidum. It is
said to occur in Massachusetts.
12. C. pulchellum, Ach. Th. membranaceous, orbicular, plane,
somewhat laciniate, round-lobed at the circumference, plicate-papulose
and dark-green above, beneath paler and deeply lacunose ; apotb.
crowded, elevated, the disk urceolate, pale, the margin thin, coarctate,
very entire, at length somewhat rugulose. Ach. Syn. p. 321.
Trunks and rocks. New England. Pennsylvania, Muhl. ! Ohio,
Mr. Lea !
13. C. saturninum, Ach. Th. rosulate, blackish-green, glaucous
and subtomentose beneath, the lobes broad, oblong, rounded, waved,
very entire ; apoth. scattered, somewhat plane, rufous, with a thin, en-
tire margin. Ach. Syn. p.Z2Q. Hook. Br. Fl. 2, p. 211. Exs. Schar. !
Helv. 423.
Trunlcs and stones. New England. Arctic America, JlicA. I have
omitted several species of this genus, which require more observation.
II. LEPTOGIUM, Fr.
Apothecia rounded, becoming discoid-open and scutelliforra, some-
what pedicellate, with a proper exciple. Thallus gelatinous-membra-
naceous, subdiaphanous, texture cellulose.
1. L. Tremelloides, Fr. Thallus foliaceous, membranaceous, very
thin and somewhat diaphanous, smooth on both sides, or powdery above,
lead-colored ; lobes oblong, rounded, very entire ; apothecia scattered,
elevated, plane, rufous-fuscous, with a paler margin. Fr. Fl. Scan. p.
293. Collema, Ach. Hook. Br. Fl. 2, ;;. 213.
Rocks and trunks, New England. New York, Torrey. Pennsylva-
nia, Muhl.
2. L. lacerum, Fr. Th. foliaceous, membranaceous, very thin and
somewhat diaphanous, glaucous-fuscescent, the lobes small, subimbri-
cate, lacerate-laciniate, denticulate-ciliate ; apoth. (small) scattered,
subsessile, somewhat concave, rufous, with a paler margin. Fr. ! Fl.
Scan. p. 293. Collema, Ach. Hook. Br. Fl. 2, p. 213.
On the earth, and rocks, among mosses. New England. New
York, Halsey. Pennsylvania, Muhl.
AND BRITISH AMERICA. 93
3. L. Burgessii, Fr. Th. membranaceous, subimbricate, glaucous-
fuscescent, somewhat spongy and downy beneath, the lobes rounded,
sinuate-laciniate, crisped and minutely lacerate-dentate at the margins ;
apolh. depressed ; disk somewhat concave, fuscous, with an elevated,
lacerate-dentate or foliose margin. CoUema, Ach. Syn. p. 320. Hook. !
Br. Fl. 2, p. 211. Icon, E. Bot. t. 300.
Trunks. Mountains of New England.
Tribe II. EPHEBIDE^.
III. EPHEBE, Fr.
Apotbecia formed from the thallus, from concave becoming plane,
and at length convex, black, the margin evanescent. Thallus filamen-
tous, not gelatinous.
E. puhescens, Fr. Thallus decumbent, softish, terete, black, the
branches entangled, capillaceous ; apothecia of the same color. Fr.
Fl. Scan. p. 294. Cornicularia, Ach. — /?. Jibrillosa, Ach. ; thallus
obscurely fuscous, smoothish, very delicate, branched, somewhat hir-
sute with numerous, flexuous, branched, subclavate fibres. Cornicula-
ria puiescens, var. Jibrillosa, Ach. Syn. p. 302.
Rocks and stones ; — a, in alpine districts. Greenland, Di'ZZ. White
Mountains. — /S, North America, Ach.
4