Pat^^ 333.95 mo Terre F2t 1980 E Habitat Inventory in Southeastern Montana RESEARCH CONDUCTED BY: MONTANA DEPARTMENT OF FISH, WILDLIFE AND PARKS SPONSORED BY: BUREAU OF LAND MANAGEMENT Kj APR 6 1981 WONTANA STATE LIBRARV5 930 E Lyndale Ave. Helena, Montana 59601 TURN Peter R. FIartin ^OVEMBER 1930 ECOLOGICAL SERVICES DIVISION FEB 9 1983 « MONTANA STATE LIBRARY S 333.95 F2t C.I Martin Terreslnal wlldlile habitat invenlory I 3 0864 00035905 2 Abstract A wildlife inventory study on three areas of southeastern Montana (Otter Creek, Prairie Dog Creek and Hanging Woman Creek) encompassing approximately 105,200 acres was begun in April 1979. Field work was completed July 1, 1980. This report, will, among other things, be used to determine habitats unsuitable for mining according to the Secretary of the Interior's "unsuitability criteria." Game species observed on the Otter Creek study area include mule deer, white-tailed deer, antelope, sharp-tailed grouse and ring- necked pheasants. One hundred and ten bird species, including the golden eagle, Swainson's hawk, prairie falcon, upland sandpiper, mountain bluebird and Brewer's sparrow, were observed. Twenty-nine mammals including beaver, raccoon, black-tailed prairie dog and red fox were observed on the study area. Ten species of amphibians and reptiles were observed. The mule deer population structure in October 1979 was 8.7 percent bucks, 41.7 percent does and 49.5 percent fawns. Fawn production was extremely high at 118.6 fawns per 100 does. Antelope production, as observed in August 1979 was 64.4 fawns per 100 does. Twenty sharp-tailed grouse dancing grounds were located. Bird species composition was determined on grassland, sagebrush, ponderosa pine- juniper and riparian habitat types. The riparian type had the highest diversity, 31 species, while the sagebrush habitat type had the lowest diversity, 6 species. Prairie falcon aeries and red-tailed hawk nests were located. Small mammal trapping was conducted on 5 habitat types; grassland, sagebrush, ponderosa pine- juniper, deciduous tree riparian and marsh riparian. The deciduous tree riparian type had the highest number of captures, 63, and the sagebrush type had the lowest, 2. The Hanging Woman Creek area had mule deer, antelope, sage grouse, sharp-tailed grouse and ring-necked pheasants as resident game species. Seventy-eight bird species, including Cooper's hawk, ferruginous hawk, golden eagle and prairie falcon, were observed. Fifteen mammal species including the coyote, yellow-bellied marmot and black-tailed prairie dog were observed. Eight amphibian and reptilian species were observed. Over 1800 mule deer observations were made during the study period. Fawn production was observed to be 92.2 fawns per 100 does in October 1979. Antelope production, as observed in July 1979, was 67.4 fawns per 100 does. Three sharp-tailed grouse dancing grouncls and five sage grouse strutting grounds were documented. Bird species composition was determined on grassland and sagebrush habitat types. Numbers observed were highest on the sagebrush type while species diversity was highest on the grassland type. Small mammal trapping was conducted on the sagebrush and grassland habitat types. Fourteen prairie dog towns were located within or near the Hanging Woman study area. Mule deer and sharp-tailed grouse were the only resident game species observed on the Prairie Dog Creek study area. Seventy bird species, including turkey vulture, great horned owl, prairie falcon, and red-tailed hawk were observed. Only 10 mammal species, including coyote and porcupine, were observed. Five species of reptiles and amphibians were observed. Only 142 mule deer were observed during the study. None were observed during the fall season. They may have moved to the adjacent Tongue River bottoms in search of more mesic conditions. One sharp-tailed grouse dancing ground was located. Bird species composition was determined on ponderosa pine-juniper and riparian habitat types. The riparian type had the most observations and the greatest species diversity. Small mammal trapping was conducted in the ponderosa pine-juniper and deciduous tree riparian habitat types. Recommendations made included exclusion from leasing sharp-tailed grouse and sage grouse breeding areas, mule deer and antelope wintering areas and all creekbottoms and associated "riparian habitats. " • TABLE OF CONTENTS Page Introduct ion „ . . . . 1 Procedures 1 Game Mammals o 1 Game Birds 2 Songbirds . 2 Raptors 3 Non-gane Mammals 3 Study Area 3 Otter Creek. , 3 Hanging Woman Creek 5 Prairie Dog Creek 5 Results and Discussion » 5 Otter Creek 11 Mule Deer , . . 11 Population characteristics. 11 Distribution. 14 Vegetation type usage „ 14 Activity 19 Use of topography . . o 20 Use of exposure . o . , 29 Use of slope. , . . . „ . 21 Antelope . o 22 Population characteristicSo 22 Distribution. o . . 22 Vegetation type usage 27 Activity „ . . . 30 Use of topography 30 Use of exposure^ . „..„...... 31 Use of slope. . „ . » o . « . ^ . . . , » . » 32 Sharp-tailed Grouse. .„,..„ „.„...,. o 33 Ring-necked Pheasant . . o . o o . <. 35 Waterfowl ... ...... » „ .o. 35 Songbirds. o „ . <, »..„....<, » 35 Raptors. <,..„.„„..,. ..o , ....... . 37 Accipiters » ......... „ . . 37 Buteos. ..... o ...„..,.. o .. o . . o o 45 Harriers. ... .o. ....<,... o 45 Falcons. ..... ........ , 45 Owls . . o . „ 45 fon-gane Mammals 50 Amphibians and Reptiles 51 Hanging Woman Creek „ 53 Mule Deer , „ 53 Population characteristics 53 Distribution. 53 Vegetation type usage. 56 Activity. ....... ............ 60 Use of topography. 60 TABLE OF CONTENTS Page Use of exposure , » o . . o . » o o 60 Use of slope o - . o o 60 Antelope o » „ 60 Population characteristics. ..... o „ . 60 Distribution « 64 Vegetation type usage. . . . . „ . 66 Activity . . . „ » QQ Use of topography ..„. .... = . . . o . . 66 Use of exposure. . . » . . . . » „ „ <, «...„.„. 71 Use of slope o . » . o 71 Sharp-tailed Grouse. ... o ..„. .... ..„.,.. o ..... o . . 71 Sage Grouse ......... . 74 Ring-necked Pheasant ............ .......... 74 Waterfowl ..................... ................. . 74 Accipiters. .................. ............. . 77 Buteos ..................... o . o o .......... o . 77 Harriers. .... ....................... 83 Falcons. ................................... 83 Non-game Mammals. ............................... 83 Prairie Dog Creek. ................................... 88 Mule Deer. , . 88 Population characteristics. ... ............ . 88 Distribution. 88 Vegetation type usage. ..................... 88 iiL^LXVXLy«»«»o o«a*«<»oeooo*ooo »*aooo*9ooeo oe« OO Use of topography .......................... 92 Use of exposure ............................ 92 Antelope 94 Sharp-tailed Grouse. ............................ 94 Waterfowl ... 94 Songbirds ....................................... 94 Raptors 99 t\ \^ Kj a, yj (j C Xb**eooo«6«*cooeo»o*u*a*oo«uo*oooi,)ou •^C/ Harriers. .................................. 99 Owls 99 Non-game Mammals . . 101 Amphibians and Reptiles 102 Summary and Recommendations. .............................. 102 Literature Cited. ......................................... 105 Appendix. ................................................. iU/ 11 LIST OF TABLES Number 1„ Average group size of mule deer in the Otter Creek, Hanging Woman Creek and Prairie Dog Creek study areas. ... 12 2„ Mule deer, antelope and coyote aerial observations per hour in the Otter Creek study area « . 13 3. Mule deer population characteristics in the Otter Creek area in 1979„...„ „ 14 4. Seasonal use of vegetation types by mule deer in the Otter Creek area . » 19 5. Seasonal activity of mule deer in the Otter Creek area... 20 6. Seasonal use of topography by mule deer in the Otter Creek area „ . „ . . 21 7. Seasonal use of exposure by mule deer in the Otter Creek area ..o ...«...<. . 22 8. Seasonal use of slope by mule deer in the Otter Creek area* 22 9. Average group size of antelope in the Otter Creek, Hanging Woman Creek and Prairie Dog Creek study area..... 26 10. Antelope population characteristics in the Otter Creek area „ , , 27 11. Seasonal use of vegetation types by antelope in the Otter Creek area „ ^0 12. Seasonal activity of antelope in the Otter Creek area.... 31 13. Seasonal use of topography by antelope in the Otter Creek area .... 31 14. Seasonal use of exposure by antelope in the Otter Creek area. 32 15. Seasonal use of slope by antelope in the Otter Creek area. 32 16. Sharp-tailed grouse dancing grounds in the Otter Creek study area and number of males attending in 1979 and 1980 ..........;„..„„..... 33 17. Breeding status of the bird species observed on the Otter Creek study area 36 18. Numbers per census run, percent composition and breeding pair density of birds observed on the grass- land habitat census strip on the Otter Creek study area.. 39 111 LIST OF TABLES Number Page 19. Numbers per census run, percent composition and breeding pair density of birds observed on the sagebrush habitat census strip on the Otter Creek study area, ...... 40 20. Numbers per census run, percent composition and breeding pair density of birds observed on the ponderosa pine- juniper census strip on the Otter Creek study area. . „ . . „ „ 41 21. Numbers per census run, percent composition, and breeding pair density of birds observed on the riparian habitat census strip on the Otter Creek study area. . „ 42 22. Raptor nest sites on the Otter Creek study area. <, . . . 49 23. Mammals observed on the Otter Creek study area. . . . . >, o « . . . 50 24. Results of small-mammal trapping on the Otter Creek study area 51 25. Amphibians and reptiles observed on the Otter Creek study area. ........... ...» 53 26. Mule deer, antelope and coyote aerial observations per hour in the Hanging Woman Creek area. „ 55 27. Mule deer population characteristics in the Hanging Woman Creek area in 1979 55 28. Seasonal use of vegetation types by mule deer in the Hanging Woman Creek area, ..,..„.. 61 29. Seasonal activity of mule deer in the Hanging Woman Creek area. , , , 61 30. Seasonal use of topography by mule deer in the Hanging Woman Creek area ...,...,........„.,. ... ... ......... o..,.. 62 31. Seasonal use of exposure by mule deer in the Hanging Woman Creek area ...,...,,., .......... 63 32. Seasonal use of slope by mule deer in the Hanging Woman Creek area ,o,. .,,o .,o ........... 63 33. Antelope population characteristics in the Hanging Woman Creek area ;....,,.„. ..o .,.,..........,,., .o. 64 34. Seasonal use of vegetation types by antelope in the Hanging Woman Creek area. ... ............................. 67 35. Seasonal activity of antelope in the Hanging Woman IV LIST OF TABLES Number Page 36, Seasonal use of topof^raphy by antelope in the Hanging Woman C-reek area 72 37. Seasonal use of exposure by antelope in the Hanging Woman Creek area ..... . 72 33. Seasonal use of slope by antelope in the Hanging Woman Creek area „ „....„.„...„.„..„ 75 39. Sharp-tailed grouse dancing grounds in the Hanging V/oman Creek area and number of males attending in 1979 and 1980 „ » „ . . . 75 40. Sage grouse strutting grounds in the Hanging Woman Creek study area and number of males attending in 1979 and 1980 , » o . « » . » » . o . , 75 41„ Breeding status of the bird species found on the Hanging Woman Creek study area. . ... ..... ..... o . o ... o ... . 76 42. Numbers per census run, percent composition, and breeding pair density of birds observed on the grass- land habitat census strip on the Hanging Woman Creek study area „ o o . „ . . . . 78 43. Numbers per census run, percent composition and breeding pair density of birds observed on the sage brush habitat census strip on the Hanging Woman Creek study area , . . o . . . . 79 44. Raptor nests sites on the Hanging Woman Creek area 80 45. Mammals observed on the Hanging Woman Creek study area. 86 46. Results of small-mammal trapping on the Hanging Woman Creek study area 86 47. Hanging Woman Creek prairie dog town locations and size 37 48„ Amphibians and reptiles observed in the Hanging Woman Creek study area 87 49. Mule deer, antelope and coyote aerial observations per hour in the Prairie Dog Creek study area 89 50. Seasonal use of vegetation types by mule deer in the Prairie Dog Creek area ". 90 51. Seasonal activity of mule deer in the Prairie Dog Creek area 99 LIST OF TABLES Number Page 52. Seasonal use of topography by mule deer in the Prairie Dog Creek area 92 53. Seasonal use of exposure by mule deer in the Prairie Dog Creek area „ 93 54. Seasonal use of slope by mule deer in the Prairie Dog Creek Study area , 93 55. Breeding status of the bird species observed on the Prairie Dog Creek study area 95 56. Numbers per census run, percent composition and breeding pair density of birds observed on the ponderosa pine-juniper habitat census strip on the Prairie Dog Creek study area . . • „ . . . » . 96 57. Numbers per census run, percent composition and breeding pair density of birds observed on the riparian habitat census strip on the Prairie Dog Creek study area 97 58. Raptor nest sites on the Prairie Dog Creek study area... 99 59. Mammals observed on the Prairie Dog Creek study area„... 101 60. Results of small-mammal trapping on the Prairie Dog Creek study area » . . o . . . o 101 61. Amphibians and reptiles observed on the Prairie Dog Creek study area. » 102 Appendix Table 1. Parameters recorded at each observation... 107 VI LIST OF FIGURES Number Page 1 . Otter Creek study area 4 2. Otter Creek study area vegetation map 6 3. Hanging Woman Creek study area „ 7 4. Hanging Woman Creek study area vegetation map 8 5. Prairie Dog Creek study area 9 6. Prairie Dog Creek study area vegetation map 10 7. Spring mule deer distribution in the Otter Creek area. 15 8. Summer mule deer distribution in the Otter Creek area. 16 9. Fall mule deer distribution in the Otter Creek area... 17 10. Winter mule deer distribution in the Otter Creek area. 18 11. Spring antelope distribution in the Otter Creek area.. 24 12. Summer antelojDe distribution in the Otter Creek area. . 25 13. Fall antelope distribution in the Otter Creek area.... 28 14. Winter antelope distribution in the Otter Creek area.. 29 15. Sharp-tailed grouse dancing grounds in the Otter Creek area o «... o ........ ..^ ... o „. 34 16. Location of songbird census strips and small mammal traplines on the Otter Creek study area. ...<,. 38 17. Sightings of raptorial birds of special interest or concern on the Otter Creek study area. o ............ ^ .» 44 18. Nests and sightings of red-tailed hawks on the Otter Creek study area „ . „ . . . o . . . „ . 46 19. Nests and sightings of golden eagles on the Otter Creek study area „ o » . o „ . „ ... 47 20. Nests and sightings of prairie falcons on the Otter Creek study area o .. ..o ...... „ , „ 48 21. Prairie dog towns on the Otter Creek study area. ...... 52 22. Spring mule deer distribution in the Hanging Woman Creek area = .» .o „....„„..„...,..„,.„,.... 54 23. Summer mule deer distribution in the Hanging Woman Creek area o . o „ ........ , , 57 Vll LIST OF FIGURES Number Pa^e 24. Fall mule deer distribution in the Hanging Woman Creek area . ^ ..... 25. Winter mule deer distribution in the Hanging Woman Creek area , .... ..o ..... » ....<,.. 26. Spring antelope distribution in the Hanging Woman Creek area ... = »«,. . • • o • 29. Winter antelope distribution in the Hanging Woman 32. Nests and sightings of raptors of special interest or concern on the Hanging Woman Creek study area. . o ...... , 33. Nests and sightings of red-tailed hawks on the Hanging Woman Creek study areas ...„„. .... o.. ..... , 34. Prairie dog towns on the Otter Creek study area, 35. Mule deer observation on the Prairie Dog Creek study 58 59 65 27. Summer antelope distribution in the Hanging Woman Creek area. 68 28. Fall antelope distribution in the Hanging Woman Creek area ........ . 69 70 30. Sharp-tailed grouse dancing grounds, sage grouse strutting grounds and observations of both species during the study on Hanging Woman Creek. '^^ 31. Locations of bird census strips and small mammal trap- lines on the Hanging Woman Creek study area. .............. ^^ 82 84 85 91 36. Locations of bird census strips and small mammal trap- lines on the Prairie Dog Creek study area. .... .o. ........ . 98 37. Locations of red-tailed hawks, prairie falcons and Cooper's hawks on the Prairie Dog Creek study area. ...... .100 Appendix Figure 1. Ponderosa pine vegetation subtype. ........ 108 Appendix Figure 2. Ponderosa pine-sagebrush vegetation subtype ........ .......... . 108 Vlll LIST OF FIGURES Page Appendix Figure 3. Ponderosa pine-grassland vegetation subtype 109 Appendix Figure 4, Ponderosa pine-skunkbush vegetation subtype . , „ „ , . 109 Appendix Figure 5. Ponderosa pine-juniper vegetation subtype . . „ , „ 110 Appendix Figure 6. Ponderosa pine-snovvberry vegetation subtype, o » » » 110 Appendix Figure 7. Sagebrush/grassland-big sagebrush vegetation subtype Ill Appendix Figure 8. Sagebrush/grassland-silver sagebrush vegetation subtype Ill Appendix Figure 9. Sagebrush/grassland-grassland vegetation subtype, o , „ „ 112 Appendix Figure 10„ Sagebrush/grassland-deciduous shrub vegetation subtype o . . o » . . . . , . 112 Appendix Figure 11. Creekbottom-deciduous tree vegetation subtype o o „ . o . » , . . . „ . . . 113 Appendix Figure 12. Creekbottom-deciduous shrub vegetation subtype. o„. o ... o o.„. 0...0 ...„„.. , 113 Appendix Figure 13. Agricultural-grassland vegetation subtype o o o ,.,......„,,.„...._.„„„,.„„„ , 114 Appendix Figure 14, Agricultural-creekbottom vegetation subtypeo .. .0 ... o „.. o ».„.. ^ o ........... „ 114 IX c Southeast Montana Wildlife Inventory Study - Final Report INTRODUCTION In response to the worldwide energy situation, the United States government has begun to shift emphasis from oil to coal as our basic energy source. Because of this shift, interest has been expanded in the massive coal deposits of eastern Montana, In order to enhance orderly, responsible development of that resource and reasonable protection for Montana's considerable wildlife resource, the Montana Department of Fish, Wildlife and Parks conducted the southeastern Montana wildlife inventory study on three potential coal lease areas; Otter Creek, Hanging Woman Creek and Prairie Dog Creek. Information gathered in this study, contracted by the Bureau of Land Management (ELM), will be used to (1) update the BLM's land use planning system; (2) establish baseline data to assist in predicting impacts from coal development; (3) assess the reclamation potential of the habitat; and (4) determine habitats unsuitable for mining according to the Secretary's "unsuitability criteria. " This study was begun in April 1979 and field work was concluded in June 1980. The primary game species found in the three study areas are mule deer {Odocoi.tza'i hzm^onai], pronghorn antelope {Antilocapfia ame.fiicana] , sharp-tailed grouse {P&dioc^t^6 phaiiamllui) , sage grouse {CzntfLoce.Kcu6 u^opha6lana6) and ring-necked pheasants [Pha6.lana.i colchicu.6] . White-tailed deer {0docolt2.ui VA^^ginlanui) are also found on the study area. Non-game bird species of special interest or concern (Flath 1979b) found in the study areas were the Swainson's hawk. Cooper's hawk, ferruginous hawk, golden eagle, bald eagle, prairie falcon, upland sandpiper, burrowing owl, long-eared owl, mountain blue- bird, clay-colored sparrow. Brewer's sparrow and field sparrow. The black-tailed prairie dog {Cijnomiji tudovicianui,) was the only non-game mammal of special interest or concern (Flath 1979b) found during the study. PROCEDURES Game Mammals Observations were made during low-level fixed-wing aerial flights in a super cub airplane. Aerial surveys were conducted monthly and data were compiled on a seasonal basis as follows: winter (December-February), spring (March-May), summer (June- August) and fall (September-November). Each study area was completely covered by flying a grid transect at half-mile intervals, oriented parallel to the ma.lor drainage. Flight times were planned to coincide with feeding periods so that maximum numbers could be observed. This biased observations in favor of the more open habitat and vegetation types. Observations were also recorded during vehicular surveys and while walking through the study areas. At each observation vegetation type, activity, type of terrain, slope, exposure and time of day were recorded (Appendix Table 1). Examples of various vegetation types are shown in Apoendix Figures 1-14. Pertinent population data were also recorded. Each observation was assigned a grid coordinate number to determine distribution patterns and for future reference. Game Birds The primary emphasis was directed towards locating sharp-tailed grouse dancing grounds and sage grouse strutting grounds. The importance of these breeding grounds to these species cannot be understated. These grounds were found by driving, listening and looking from the ground and by flying low over likely terrain features and either seeing the birds on their arenas or flushing them into the air. Pheasant crow count routes were conducted during the spring breeding season to determine the density of cock pheasants (Kimball 1949). Ponds, sloughs, and creeks were visited during the spring and summer months to obtain waterfowl observations. Songbirds Species composition, relative abundance, and breeding densities of songbirds were determined using a strip-census method suggested by Hickey and Mikol (1979) for use in coal mine baseline studies. Two 100 m wide, 2500 m long strips were established in each of the four main habitats found in the study areas (grassland, sagebrush, ponderosa pine-juniper forest, deciduous tree riparian). Songbirds were mapped along each strip five times during the breeding season (early May - early July) in 1979 and three times in 1980. Two methods were used to estimate breeding bird densities as described by Hickey and Mikol (1979). The first method uses the sum of the average number per census run of singing (territorial) males of each species. This method may underestimate the number of birds for two reasons: 1) late migrants such as western wood pewees may not have been present during the earlier census runs; 2) some birds are silent and therefore undetected during some census runs. The second method, called the summation method, uses the sum of the highest counts of singing males for each species. This method may overestimate the number of birds during periods when migrants (which often sing) are present along with residents. It may underestimate the number of birds in most cases, since it is unlikely that all the resident males along a strip will be detected in one census run. A species list was compiled for each study area. Nomenclature follows the A.O.U. checklist (1957) and Supplements (1973,1976). Raptors Raptor nests were located using the methods described by Call (1978). Creekbottoms were searched in the early spring by airplane and from the ground for hawk and eagle stick nests. Suitable cliffs were examined by airplane and on foot for falcon aeries and eagle nests. Observations of all raptors except kestrels were mapped during the breeding season to aid in delineating territories. Intensive foot searches were employed when highly defensive adults were encountered. Prairie dog towns were checked during July for burrowing owl broods. Non-game Mammals Non-game mammals were sampled using the trapping method suggested by Flath (1979a) for use in baseline studies. Two traplines were set up on each of the four main habitats, and run for five days consecutively. One trapline in deciduous tree riparian habitat was heavily disturbed by raccoons, so an extra trapline was set up in this habitat and run for six days consecutively. One trapline was also set up in marsh riparian habitat and run for six days consecutively. Each trapline consisted of 25 stations spaced 10 m apart. Each station contained one Sherman live-trap, two mouse snap-traps, and one 5-pound coffee can pitfall trap. A mixture of peanut butter and rolled oats was used as bait. Live traps and snap traps which were sprung, but empty were subtracted from the total trap nights. Several pitfall trap-nights were also subtracted due to flooding. STUDY AREA Otter Creek The Otter Creek portion of the study area (Figure 1) is located east of the Tongue River near Ashland, Montana. It lies primarily in Powder River county with about 10 percent of its approximately 59,000 surface acres in Rosebud County. Otter Creek is the primary drainage system in the study area flowing from the southern border northeast to the Tongue River at Ashland. Double E coulee and Cook Creek provide for drainage of the northern portion of the area. Home Creek and Three-mile Creek are the major tributaries of Otter Creek in the southern portion. Several other creeks are also found in the area. The area south of U.S. Highway 212 is dominated by Otter Creek and its associated agricultural fields. Terraces adjacent to the creek and dissected mid-slopes comprise most of the area. Near the edge of the Custer National Forest, ponderosa pine uplands and steep sandstone bluffs are the dominant features. North of the highway, ponderosa pine breaks and creekbottoms are most prominent. The plateau areas between drainages have some agricultural development. Figure 1. Otter Creek study ar ea, A broad vegetation map (Figure 2) shows the three primary native vegetation types as they occur within the Otter Creek study area. These types are the ponderosa pine, the sagebrush-grassland and the riparian. Each type is very diverse with several subtypes, which are not depicted. The type boundaries are only approximations and include portions of the other vegetation types. The figure is presented only to give the reader a general feeling for the vegetative makeup of the area. A more detailed vegetation map has been published by the Western Energy and Land Use Team, U.S. Fish and Wildlife Service (U.S. D.I. 1978). Hanging Woman Creek The Hanging Woman Creek portion of the study area (Figure 3), is located 10 miles south of Birney, Montana and 3 miles north of the Montana-Wyoming border. It is about 12 miles east of the Tongue River Reservoir. It covers nearly 35,200 acres, all in Big Horn County. Hanging Woman Creek is the principal waterway and prominent terrain feature. It runs from the southern to the northern boundary. Horse Creek, East Fork Trail Creek, Corral Creek and First Creek are the major side drainages. Sagebrush and grassland vegetation types (Figure 4) constitute most of the ground cover in this study area. The only agricultural endeavor is hay production, primarily along Hanging Woman Creek bottoms . Prairie Dog Creek The Prairie Dog Creek portion of the study area (Figure 5), is located about 7 miles west of Birney. It is adjacent to and south of the Northern Cheyenne Indian Reservation, It lies approximately 11 miles northwest of the Tongue River Reservoiro Nearly 11,000 acres of Rosebud County are included in this study area. Prairie Dog Creek is the principal drainage and flows in a southeasterly direction across the area. Bull Creek borders on the north and Canyon Creek borders the area on the south. This area is characterized by high buttes with over 500 feet in relief from the plateau tops to the creekbottoms. Ponderosa pine is the dominant vegetation type (Figure 6) and there is practically no agricultural development. RESULTS AND DISCUSSION Each species or group of species is dicussed separately for each of the three study areas. Distribution maps are presented for the important game species and most of the non- game species of special interest or concern. LEGEND Permanent stream . . 'va/v Paved road Unpaved road — = Forest boundary Study area -• >- Vegetation map Ponderosa pine. jWi'* Sagebrush/ .^ Grassland ^. Riparian Figure 2, Otter Creek study area vegetation map. LEGEND Permanenf stream -'^^ County road State line : Study area -^ ^ oJadd\« MONTANA WYOMING Figure 3. Hangins Woman Creek study area. LEGEND Permanent stream County road State line Study a rea Vegetation map Ponderosa pine. Sagebrush/ Grassland Riparian MONTANA WYOMING Figure 4. Hanging Woman Creek study area vegetation nap. J.EGEND Permanent stream County road Cheyenne reserV^Ltjon^X- ^Study area .5 0 1 2 3 miles Figure 5. Prairie Dog Creek study area. u u to > E c 0 •H 4-> +j a> bJD 0) > 0) U aS >. -d =! +-> ^ CD 0) U u o n u (5 10 Care should be taken in interpretation of the tables and figures presented herein as they represent only fifteen months observations The winter period was exceptionally mild and the remainder of the months were marked by below normal precipitation in most instances. Otter Creek Mule Deer Population characteristics: During the study 831 mule deer were observed in the Otter Creek study area (Table 1). Average group size ranged from 2.4 during the summer months (1979) to 6.9 during winter. April 1980 had the highest individual monthly average as mule deer remained in herds prior to spring green-up. Deer were most scattered in May and June of 1980. Dusek (1978) reported group size variation from 2.0 in summer to 5.6 in winter in the Bull Mountains near Roundup. The indices of relative abundance (Odum 1959) have been used to determine population trends in southeastern Montana (Martin 1980). Mule deer observations per hour of aerial survey (Table 2) show the highest numbers during fall when an average 31.1 deer were observed. November was the best month for observation as 55.0 mule deer were observed per hour of flight. This coincides with the time of year that deer should be most numerous. February was the second most productive month, 40.4 deer observed per hour, reflecting the deer's need to feed more as the effects of winter are being manifested. The fewest deer were seen in May of 1980. This unusually low number, 2.8 per hour, is at least partially due to windy conditions during the flight. Low numbers per hour should be expected in spring and summer as the fawns have either not been born or are not traveling with the adults. Martin (1980) reported a range of 6.1 mule deer observations per hour in summer to 25.3 in fall m the Sarpy Creek area. Mule deer population structure was determined during September and October (Table 3). During these months fawns are traveling with the adults and are still small enough to be easily Identified. Bucks are also easily classified. While there has been some natural mortality among the young deer and some loss ot adults to poaching and other factors, the population is at Its yearly peak during this season. As can be seen production was extremely good this year with 118.6 fawns per 100 does observed in October prior to the hunting season. Considering the extremely hard winters the last two years, 1977-78 and 1978- 79, it seems that this herd came through in excellent condition. They apparently are considerably below the carrying capacity of this range They also appear to be on an increasing trend as Swenson (1978) reported that mule deer need only a 40 percent ^rLi''''''i ^? rr^'" stable. The October flight revealed nearly 50 percent of the population to be fawns. 11 a u c 0 U o c d E O & be a •H CIS X X 0) 0 u ■p ■p o 0) ■p f-l 0) 0 T3 CD I— I 3 E «H C/l 0 d 0) 0) ;h N ni •H m >- -a a =i :3 p 0 Cfl ^H M^ (U 0) 01 bC ^1 rt O ^ (U bfi > 0 < P CD It-; o o Q T— 1 a W p 1— ( 0 « H 1— ( < K o a w > a !h :3 0) 0 K) fn JD a o IX s < r-i s d o P ^ 0 Eh o z l-H r~ g 00 > < a Sh w ^ o o ^1 1 kyf 1 r-i r— 1 ^> d H p W O tf H u K • W m > Eh 0. 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C rH tC :3 13 3 I-D "-O 0) o o m o z Qj d a; n '-D ph 0^ o 00 CD CO CM CD ^ CM CM CD ^ t^ CM t^ "* I CM t- I I t- in CD 3 CO 0) >, C rH bJD 3 3 13 •^ -^ < CO > CO c o 0) X3 0) X Xi c d O ^- Cfl cti d >, > ^ (D ^ 0 > (1) > Jh CO CO o |Ol|O0| 12 cu u o u (D +-> o £i -P o Xi u CO c o •H +J ni > U O CO o 0) oS 0 -P o o o T3 C oS 0 a o iH 0) ■p a a - d ^H CD CD rt ■-H 3 P P o o 0) a Cfl +-> O 0) O 3 0 ■P o o u p c <; S Q CD O o o ai •H 0) i-H < 3 0) CD o CO I l> I o CO in Oi O) o cy> in in O CD ^ O CS] i-H O ^ I> o in CD O O CJ) (M 00 in CD CO CD CO CM I> CD rH O CD CM CO CO CMI O ^ CM in CO o c^ in '^ 00 CD o 0^ CM CO in o iH o C<1 CO CM CM CO CD O CM O O 00 00 00 CM 00 o o • • • CO rH CO C-^ CO I> o in 00 o O (-t I> CO -^ -H O CO CO rH CM O CM CD 00 CD CO 03 CO CM in oo CO m CM 00 o o l> o eg CO CO in o CO in o o CM CM CD CM CO O oa CM O O rH O O rH O O "^ O O I> CT; CM O O rH ■* in t> 1-i C r-t CJ) "^ o o CD '^r O CM O CO CM rH rji O O ^ CO 00 00 rH t> O O t> 05 CM CM '^f -* !> ^ CM C3^ O O O CM cy> rr CD CM eg CD 00 00 O in CD CM CO O rH CO C CD 00 O O CM O CM CD CD "^ O] CD O I> « • • rH c^i a> t> CO ^ '^ 00 in CM '^ in CD CM ^ CM in CO o C: lO r-i t> 00 CD rH CD CJ3 CO CO in in cv] CM CM CO CO 00 CD Cvl 00 Olio o O I I I I CD CO I I CO •<* I I O I I O '-i CM in I CD I I ^ I I in CD M< fcuO U u bo ?-i a Q) 0 C 0 •H s rH -p •iH p r-t ;-! g <-i c X; rH u ^ •H D. CD >> 3 ■p a •H cj -H a 0 >> 3 Jh >.co c rH hucn a-p > ph o a x^ ^ ^H Jh >,co C rH bJD CO a os 3 3 3 -:) Ph s <: s I-Tl ^ < CJ) o t> t^ 00 CT> Gi CJ) Cfi >> 0 > !h 3 W C 0 0 Td d E 0 w T3 c d o E ■iH P w d >> > 0 ;h > 0 ^^ m 3 ^ M O 0 o ST S?: lev] I 13 Table 3. Mule deer population characteristics in the Otter Creek area in 1979, Month Number Classified Total Bucks Does Fawns Fawns: 100 Bucks: Population Structure (%) Does Adults 100 Does Bucks Does Fawns September 52 5 24 23 95.8 79,3 20.8 October 103 9 43 51 118.6 98.1 20.9 9.6 46.2 44.2 8.7 41.7 49.5 Distribution: Spring distribution is shown in Figure 7. Most observations are located in a band across the center of the study area between Threemile and East Fork Otter Creeks. Several of the observations were in the 9-15 size category. More than half of the observations were in the 1-3 range. The deer are beginning to disperse over the area and break up the wintering groups. This dispersal and reduction of group size reaches its zenith during the summer season (Figure 8). Only three observations were of groups greater than three. Fall observations (Figure 9) show the deer beginning to group together as the number of observations in the 4-7 size category increased significantly. Also the deer were not as widespread as during the summer months. Three groups of deer appear to be forming: those north of highway 212; the central group stretching across the study area on a southwest-northeast axis centered on Home Creek; and a southern group extending from above Newell Creek to below Ten Mile Creek. The 1979-1980 winter in contrast to the two previous winters, was very mild with little snow fall. As a result, mule deer remained quite widespread and did not gather into large concentrations (Figure 10). Nearly all observations were in the 4-8 size category. One observation of a group larger than 15 animals was made and four groups in the 9-15 range were seen. Two major wintering areas were determined: one southwest of Otter Creek north of the Custer National Forest; the other east of Otter Creek and south of Threemile Creek. The remainder of the observations were too scattered to consider. It is likely that many deer wintered in the adjacent Custer National Forest. i Vegetation type usage: A considerab is apparent in usage of vegetation t percent of the spring 1979 observati pine type while only 15 percent util The weather pattern, deep snow and c 1979 spring which forced deer out of grassland types, explains the variat winter mule deer were able to utiliz pine and only 58 percent were observ extremely dry summer of 1980 forced coulees and draws of the pine type. le difference between years ypes (Table 4). Ninety-three ons were in the ponderosa ized the pine type in 1980. old in 1978 extending into the the creekbottoris and sagebrush/ ion. During the mild 1979-80 e vegetation types other than ed in the pine type. The deer to utilize the more mesic 14 LEGEND Permanent stream . '^a/^ Paved road Unpaved road — Forest boundary Study area -> <- Mule deer ob servations Group size 1-3 A ° / 4-8 a ' M 9-15 A 9 ^=V\J 16+ A * 3 Year 1979 198d Fipcuro 7. Spring nule deer distribution in th(^ Otter Creek area. 15 LEGEND Permanent stream . ''^v^ Paved road Unpaved road — = Forest boundary.. Study area -> -<- Mule deer observations Group size 1-3 ^ 4-8 -^ 9-15 * 16+ A Year 1979 198 Figure 8. Summer mule deer distribution in the Otter Creek area. 16 LEGEND Permanent stream ^^a/v Paved road Unpaved road Forest boundary — Study area Mule deer observations Group size 1 " 3 ^ o « ' 4-8 A o ^t 9-15 A e ^9 16+ * • -3 Year 1979 1980 i Fir.uro 0. Fall nu ] e deer distribution in the Otter Creek area. 17 LEGEND Permanent stream . aaa^ Paved road Unpaved road — Forest boundary. , Study area -• *- Year 1979 198 Mule deer ob servations Group s ize <*•" 1 - 3 A o / 4- 8 '5> . o 9- 15 A 9 ,^^ 16 + A . . • V Figure 10. Winter mule deer distribution in the Otter Creek area. 18 a u d X (D 0) u u CD 0 X! +-> U CD 0) T3 :3 S CD n >> +j c o •H +-> a +-> 0) b£ > o CD 3 C O a CD CD r-l n! ^H CD O e 00 1> E cs) csi 3 1— 1 m bjO C o •H OO CD ^ a o ai-i cMi w o ^H CX) CD 1 ■p C3) lO c t> CD •H C35 i-l & iH 1— 1 CD CDO t— 1 t> 00 ai O) 1— 1 fc iH u CD CT, g t- f— 1 g CT. T— 1 3 rH r—l CO M ^^ C Oil-Ill •r-l I> r-^ u CJi o a r—l r—l CO w CD a >. +-> 3 m c 0 •H +-> a +j CD hC CD > CO ^ I I CO oq i> I oj I OO CO t> CM 00 I CM CM CO in I I m in o t> 00 I cm! CD 1—1 -^ oq in o in O I rH C5 I CM CM I CM r-i ^ a CO rH o CM I rH I> I I CD CO CO I -^ -^ I CO CM CM in 't I I ^ CO 00 CO O) I ^ I '^ ^ CO I I I rH I rH I \t< CM I I I ^ CM CM in c d rH CO ■a (D CO c c d d •H u rH tH ft ^ O E CO d 0) rs — ^ o 0 CO tH c ni ;h s: -p C rt 3 •H CO x: CO +J •H ;h -p a O CO 3 u o ao E r-l "--OP a szT} s: >. 0 .c CO "O X5 ^ d xi 4^ o CO Cfl C 03 U T3 CO 3 G 0 0 (fi U) +^ ■'-1 O 3 rt 3 ^1 ^ C :3 o d M 0 O 3 0 ;h ;h J-l rH ^ CD CD 0 ^^ 13 rH oi ^H U {-, Xi be C) XI ^ X ax2 ft ^ T! CO CO -D U Q) X X < •d CD C/] c •H S 0 ■H CO 0 3 T3 0 0 C be rt 3 c O rH M O d rH 0 ^ iH C bX) 0 rH o a u X d C Oj oj 0 u d U x: d 0 cti ;-! oj ft CO O CO "-3 CO -P CO o a +-> ^ CO ■P ft CO CJ -P O O 0 0 £-> H H H 0 X 5 c 0 rH c x; & ■p CO 0 d 0 C O +-> ■o 0 C o to fl o •H -P CO d c o •H +-1 > o 0 tp CO o O -P C rH 0 ctS O -p ^ O 0 H ft HJCMl 19 The sagebrush/grassland type received its highest usage in the fall of 1979 and spring of 1980, 55 and 56 percent, respectively. Its lowest usage was noted in the spring of 1979 and the dry summer of 1980 with 7 and 19 percent of the observations respectively. Apparently the sagebrush/grassland type is not preferred by deer in either weather extreme, snow and cold or hot and dry. The creekbottom type usage ranged from none to nine percent over the course of the study. A goodly proportion of the Otter Creek bottoms have been developed into farmland. As a result only a few sections have adequate cover. Those remaining stretches with good riparian vegetation are extremely important to deer and other wildlife species. They should be protected in every possible instance. Combined creekbottom subtypes showed the highest usage, 21 percent of observations, during the 1979 summer. Deer utilize the agricultural areas quite heavily during the period of initial "greenup . " This occurred in the early summer months of 1979 and spring of 1980 when 15 and 24 percent of the deer were observed in agricultural fields in those seasons respectively. Very few observations were made in the agricultural types in any other season. Activity : Examination of mule deer activity at the time of observation (Table 5) gives some insight into the habits and condition of the deer herd. The high percentages of deer feeding in spring and summer indicate they were building up their fat reserves. A small, but substantial, percentage (over 25 percent) was observed feeding during fall and winter. The seasons of greatest stress, the cold spring of 1979, winter 1979-80 and the hot, dry 1980 summer, correlate with the greatest numbers observed lying down. Since most of the deer resorted to pine cover areas with their inherent observa- bility bias, the percentage actually lying down was no doubt underestimated. Table 5. Seasonal activity of mule deer in the Otter Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Activity lOli/ 111 188 165 218 27 Standing 362/ 37 63 44 11 15 Running 8 9 9 20 ^o/ 30 Lying 10 5 1 10 tr3/ 11 Feeding 47 50 27 26 83 44 \_l Total observations 2/ Percent of observations rounded to nearest whole number 3/ tr = trace; a percentage less than 0.5 20 Use of topography: Dissected midslopes were most heavily used by mule deer in all seasons of the study (Table 6), averaging nearly 80 percent for the six seasons. Two distinct peaks of lowland usage were noted, 19 percent in summer 1979 and 26 percent in spring 1980. These two seasons represent the initial "green-up" period for the respective years. A similar decrease in usage of the uplands types was noted from spring highs of 22 and 24 percent for 1979 and 1980 respectively to summer lows of 3 and 0 percent respectively. A lack of water at the higher elevations has been cited as being responsible for this shift in other areas of eastern Montana (Martin 1980, Knowles 1975). Table 6. Seasonal use of topography by mule deer in the Otter Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Topography lOli/ 111 188 165 218 27 24 - tr3/ - 50 89 22 7 4 4 Mesa-butte top 132/ 14 2 Mesa-butte steep slopes 9 2 5 3 Dissected mid- slopes 73 78 86 87 Alluvium/terrace - 5 2 - Flood plain - 14 3 8 !_/ Total observations 2/ Percent of observations rounded to nearest whole number 3^/ tr = trace; a percentage less than 0.5. Use of exposure: Seasonal use of exposure is shown in Table 7. Usage of northerly slopes decreased from its high, 41 percent, in the 1979 spring to its low, 17 percent, during the 1980 spring. Usage of southerly slopes was at or near 45 percent in all seasons except the 1979 summer and 1980 spring which have been discussed previously as being the green-up period. Easterly exposures were least used during the two spring seasons with 26 and 13 percent for 1979 and 1980 respectively. All other seasons usage levels were between 31 and 39 percent. Conversely, usage of westerly exposures were highest during the spring seasons, The 1979 spring saw 58 percent of mule deer observations on westerly slopes, the highest figure recorded during the course of the study. Numbers observed on flat lands were highest during the summer of 1979, 16 percent and the spring of 1980, 21 percent. 21 Table 7, Seasonal use of exposure by mule deer in the Otte area. r Creek Exposure Spring 1979 lOll/ bummer 1979 111 Fall 1979 188 Winter '79-80 165 Spring 1980 218 Summer 1980 26 North South East West Northeast 62/ 10 14 24 14 12 12 11 12 21 12 13 17 5 39 10 14 21 5 6 10 15 7 30 6 1 27 23 23 4 Northwest 11 13 2 '~~ Southeast Q 16 2 — Southwest Flat 33 1 6 16 20 2 11 21 15 4 4 1/ Total observations 2/ Percent of observations rounded to nearest whole number Use of slope: Usage of the medium and steep slopes was hi-hest during the seasons of cold and snow. Eighty percent were ?TS^^«^°" Jhe steeper slopes in the 1979 spring season (Table 8). Sixty-eight and 71 percent were observed during ^nd liln ^"^H^'if ^""/^^^^"^ respectively. The summers of 1979 and 1980 with 54 and 66 percent respectively and the 1980 spring Zilt^ ^f'"''^"^ °^ *^^ ""^^ ^^^^ observations represent warm ^ weather slope usage patterns. Table 8. Seasonal area. use of slope by mule deer in the Ott er Creek Slope Spring 1979 1011/ Summer 1979 111 Fall 1979 188 Winter '79-80 165 Spring 1980 218 Summer 1980 26 Flat Gentle Medium Steep l2/ 19 65 15 16 38 31 15 2 30 36 32 29 39 32 21 54 19 6 8 58 35 1/ Total observations 2/ Percent of observations rounded to nearest whole number, 22 Antelope Population characteristics: During the six season study, 1784 antelope were observed in the Otter Creek study area (Table 9). Average group size ranged from a low of 3.0 in spring 1979 to a high of 18.4 during the winter months. January, in the dead of winter, recorded 25.7 antelope per group. Dispersal from winter herds occurred in April as the group size decreased from 13.4 in March to 5.0 antelope per group in April. Antelope were most easily spotted in December and February when over 91 antelope were observed per hour of aerial survey (Table 2). This is when the antelope are concentrated in large groups. Many antelope appeared to have left the Otter Creek area during January. The total number observed from the air, 77, was considerably below the 200 plus numbers observed in December and February. Perhaps this area represents marginal wintering habitat for antelope. High numbers per hour were also observed in July and August. This corresponds to the tine frame when antelope numbers should be at their seasonal peak just shortly after fawn recruitment. Production figures and population structure percentages are shown in Table 10. August data in 1979 show 64.4 fawns per 100 does. This compares to 68 and 59 fawns per 100 females in Fish, Wildlife and Parks hunting units 740 and 741 in 1978 and 1977 respectively, the latest year in which surveys were conducted (Wentland 1979). June of 1980 saw the percentage of fawns in the population remain stable at slightly over 20 percent, while the number of fawns per 100 does increased from 33.3 to 40.0 over June of 1979. The percentage of bucks was 28.9 in June of 1980, slightly lower than the average of 30.7 recorded for hunting unit 740 since 1963 (Wentland 1979) » Distribution : Antelope spring distribution is shown in Figure 11. There appears to be four widely scattered groupings of antelope observations. One is located on either side of Cook Creek north of highway 212. Another stretches along the southwest side of Otter Creek north of the Custer National Forest, This group appears to have the second largest number of sightings. Another group is found astride Threemile Creek on the eastern edge of the study area. The greatest number of observations occurred in the southern portion of the study area with Newell Creek as its approximate northern boundary. Many antelope are still in large herds at this time exhibited by the number of group sightings in the 9-15 and over 15 categories . During the summer months (Figure 12), antelope were more dispersed than during spring. The four groupings mentioned above seemed to 23 LEGEND Permanent stream Paved road Unpaved road Forest boundary Study area Figure 11. Spring antelope distribution in the Otter Creek area. 24 LEGEND Permanent stream ^^a/v Paved road Unpaved road — = Forest boundary Study area -n .- Antelope observations Group size ./ 1 " 3 a o < 4-8 & o ^or 9-15 A 16+ * Year 1979 198' Fif^ure 12. Summer antelope distribution in the Otter Creek area. 25 0 •H U •H e<5 $-1 P< •a o ■p 0) a 0 rH 0) +J c nS • tH en O d 0 0) fn N rt •H cn >> -o a 3 a +-> 0 m ^ bC^J 0 0) 0 bc ^H d O ;h 0 bC > O < P • 05 O iH ^ rt E- IX d +-> b£ O O Eh Q CO > o w o o d 6 O &: bO a IX bD c c- > d :^ u K =! 0 o CO o o IX 0 +-> Cfi > -P =H ^ O ^ 0 O Cfl c o o o o o o o CO o ^ CO CO iH r-l O rH CSl CD t> CSl CD O O CM CO CO CO r-t CO CO bo c ■rH rH ^H •H a a d o o o o o o o o o t> O Tj< in CD CO CO in CJi -sr CO CM rH '^ 05 rH in •^ CM 00 CO CD CM I> CD CD CO 1> CJ3 rH l> CM r-\ n^ (y> t> CO CSl OJ o o I o o I o o I 00 Tf< G> CO t> ^ CD I> l> l> CO CO CM CM -0| in o I r}< CO c o in o 05 CD O O CJ) O O rH O o o o CM rH O I o O I O O I O CO ca CD rH [> in oi CSl Oi ■^ rH ■* I> 00 r}< CM CO in 00 CO '^ CO I> !>• CM in -^j* CM CM rH rH I> O O t^ C<] CO CM as c^ in CM o in CM ^ 00 CM 0 0 >> =3 C •-> bJ3W P 13 3 •-D »-D < +J • d a+j > Ph 0 o o m o ^, 0 a •H O C X3 & 0 oi 0 P l-D P>H O O O O O C o o o rH O ^ CD ^ rH CD CO O rH O CO CM CM c^ in CD CM r}i -::}< ^ O O CO in CO •^ in CO rH CM l> CM rH CD in -st* rH CM CM o CM 00 ho c •H o -H a d Q^ d s <: s O I I O I I O I I CD CD • I I CD CO in i> I 00 I I CO n I I o [> I C<1 I I r-i CM 0 C ;3 3 CD CO 00 CO CO i-D >-:) < U 0 E E bDW 3 0 d U 0 > d >. 0 > Sh W C o 0 d E v. 0 C T3 O ctj ■H E +-> d CO > >> 0 > Cfl o o rH| O]|C0| 26 Table 10. Antelope population characteristics in the Otter Creek area. Number Classified Fawns ;:100 Bucks: Population Structure (%) Month Total Bucks Does Fawns Does Adults 100 Does Buclis Does Fawns 1979 June 96 16 60 20 33.3 26.3 26.7 16.6 62.5 20.8 July 213 26 116 71 61.2 50.0 22.4 12.2 54.4 33.3 Aug 165 22 87 56 64.4 51.3 25.3 13.3 52.7 33.9 Sept 81 18 43 20 46.5 32.8 41.9 22.2 53.1 24.7 19S0 June 59 17 30 12 40.0 25.5 56.7 28.9 50.8 20.3 blend together with a slight overall shift to the southern portion of the study area. The northern group still appears to be separated from the rest of the study area. While some larger herds remained intact, more observations in the 1-3 range were recorded than in any other season. Antelope fall distribution is shown in Figure 13. The number of observations are greatly reduced because there was only one fall season in the study and antelope are beginning to congregate into their winter herds. Except for a few scattered observations, the antelope seemed to be located in the southern portion of the study area with Threemile Creek as the approximate northern boundary. Five groups larger than 15 were observed and six groups in the 9-15 category were spotted. Figure 14 shows the winter antelope distribution. The pattern established during fall was continued as most of the observations were made south of Threemile Creek. A large majority of the antelope were in herds of over 16 individuals. The average group size was 18.4 (Table 9). Since the 1979-80 winter season was quite mild, antelope were free to roam about their range at will. Therefore the areas used during this study probably include more area than that used in more severe winter conditions. Vegetation type usage: Antelope were found primarily on the sagebrush/grassland vegetation types (Table 11). Ninety-seven percent of the antelope were observed on this type during the fall and winter seasons. The smallest percentage utilizing sagebrush types was seen in spring 1979, 84 percent. That spring 16 percent were observed in the ponderosa pine grassland subtype. 27 abc;de fgh LEGEND Permanent stream . -vaa. Paved road Unpaved road — Forest boundary Study area -> 1- Antelope observations Group size ; 1 " 3 a o * 4-8 ^ o J^ 9-15 A 16+ A • Year 1979 198' Figure 13. Fall antelope distribution in the Otter Creek area. 28 LEGEND Permanent stream Paved road Unpaved road Forest boundary Study area Fif^ure 14. 'Vinter antelope distribution in the Otter Creek area. 29 The only other subtype to receive more than mininal usage was the agricultural sagebrush/grassland subtype in the summer of 1980, where 9 percent of the antelope were observed. Table 11. Seasonal use of vegetation type; Otter Creek area. 3 by a ntelope in the Vegetation Subtypes Spring 1979 991/ Sunmer Fall ' 1979 1979 486 219 Winter '79-80 498 Spring 1980 412 Sunmer 1980 70 Ponderosa pine Sagebrush Grassland Skunkbush Juniper Snowberry 152/ 2 2 1 1 2 2 1 Total Ponderosa pine Sagebrush Deciduous shrub Grassland 16 34 49 5 43 53 48 44 1 39 58 4 51 40 1 24 66 Total Sagebrush/Grassland Tree Shrub 84 91 97 1 97 1 91 4 90 Total Creekbottom Ponderosa pine Sagebrush/grassland Creekbottom - 1 - tr3/ 2 3 1 4 1 1 9 Total Agricultural " 2 3 " 2 y 1/ Total observations 2/ Percent of observations rounded to nearest whole number 3/ tr = trace; a percentage 0.5 Activity: Antelope are more apt to run during aerial observation surveys than are mule deer. During the summer surveys, the highest percentage of running antelope were observed feeding in the spring months. More antelope were observed standing, 51 percent, in the winter time than in any other season. Use of topography: Antelope were most often observed on dissected mid-slope areas (Table 13) in all seasons. The percentage observed ranged from 74 in the summer of 1980 to 97 in the fall of 1979. Alluvium/terraces received use in every season except spring 1979 reaching its highest level, 21 percent, in summer 1980. Mesa-butte tops were slightly used every season except fall 1979. Mesa-butte steep sides and flood plains were seldom used. 30 Table 12. Seasonal activity of antelope in the Otter Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Activity 991/ 486 219 498 412 70 Standing 25 35 36 51 37 31 Running 34 45 39 25 18 44 Lying 16 8 8 9 15 6 Feeding 24 13 17 14 30 19 1/ Total observations 2/ Percent of observations rounded to the nearest whole number, Table 13. Seasonal use of topography by antelope in the Otter Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Topography 991/ 4 86 219 498 412 66 Mesa-butte top lO^./ 6-22 5 Mesa-butte steep sides _ 2 - - - Dissected mid- 90 90 97 95 86 74 slopes Alluvium/terrace - 2 3 2 8 21 Flood plain - 1 - 1 3 1^/ Total observations 2/ Percent of observations rounded to nearest whole number Use of exposure: Antelope were observed on all exposures in nearly every season (Table 14). The greatest usage occurred on southerly exposures in the spring of 1979, 57 percent. Flat lands generally had the least amount of antelope usage. This reflects their lack of usage of floodplains and plateau areas as discussed above. Northerly slopes were most highly used in the fall season, 56 percent of the observations, and least used in the spring of 1979. Easterly slopes were most often used in spring 1979 and summer 1980, 50 and 49 percent respectively. 31 Table 14. Seasonal use of exposure by antelope in the Otter Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Exposure 991/ 486 219 498 412 66 North 32/ 21 12 10 14 12 South 19 9 10 5 8 11 East - 14 9 7 15 3 West 6 15 — 14 6 — Northeast 25 14 27 24 24 23 Northwest 5 5 17 3 7 6 Southeast 25 12 3 6 8 23 Southwest 13 6 19 26 17 17 Flat 3 4 3 5 3 6 \_l Total observations 2/ Percent of observations rounded to nearest whole number, Use of slope: Antelope avoided the flat and steep slopes throughout the study (Table 15) with 6 percent being the highest percentage observed in any season. The spring and summer seasons of 1979 saw almost identical usage at gentle and medium slopes. In all other seasons, gentle slopes supported more antelope than medium slopes. Its highest level was 66 percent in the fall of 1979. « Table 15. Seasonal use of slope by antelope in the Otter Creek area. Spring Summer Fall Win ter Spring Summer 1979 1979 1979 '79 -80 1980 1980 Slope 991/ 486 219 498 412 66 Flat 32/ 4 3 5 3 6 Gentle 46 45 66 56 59 53 Medium 49 47 32 36 32 41 Steep 3 4 — 3 6 — \_l Total observations 2/ Percent of observations rounded to the nearest whole number. Sharp-tailed Grouse Twenty dancing grounds have been located in or near the Otter Creek study area (Figure 15). Attendance by male birds at known grounds averaged 15.3 in 1979 and 17.8 in 1980 (Table 16) The average number of birds in 1980 at the seven grounds located in 1979 was 23.0. This represents a 50.3 percent increase from 1979 to 1980. Obviously the overwinter survival was extremely high. There are approximately 0.22 dancing grounds per square mile in the study area. This compares with .09 grounds per square mile in the Sarpy Creek drainage (Martin 1980) and 0.12 per square mile in the Colstrip area (Schwarzkoph 1980). No broods were observed in the study area. One observation of approximately 150 birds was made along Home Creek north of dancing ground number 7 in early November. Table 16. Sharp-tailed grouse dancing grounds in the Otter Creek study area and number of males attending in 1979 and 1980. Dancing Ground Location T R S 5S 4 5E 23 NEi 5S 45E 3 SEi 4S 45E 33 SEi 4S 45E 12 NW^ 4S 45E 3 NWi 3S 45E 8 SEJ 3S 45E 35 NEi 2S 45E 32 NWi 3S 45E 5 NEi 3S 45E 3 SE^ 3S 44E 23 SE^ 3S 44E 36 SE^ 3S 46E 30 SWJ 3S 46E 31 SW;> 4S 45E 5 SEi 4S 45E 17 SEi 4S 4 5E 26 NWJ 4S 45E 36 SWi 5S 45E 15 NW^ 5S 45E 22 s\H Activity 1979 1980 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 12 9 12 20 1 13 40 13 20 33 36 5 24 30 36 33 9 7 5 20 2 16 6 20 16 16 9 Average males per ground 15.3 17.8 33 LEGEND Permanent stream 'va/v Paved road Unpaved road — = Forest boundary. Study area -< >- Sharp- tailed grouse dancing grounds • «, a Figure 15. Sharp-tailed grouse dancing grounds in the Otter Creek area. 34 Ring-necked Pheasant Ring-necked pheasants occur in the study area along all the major drainage ways. Highest numbers were observed along Otter Creek. A pheasant crow count route has been conducted south from the Otter Creek road - U.S. Highway 212 junction by the Montana Department of Fish, Wildlife and Parks since 1974. The average number of calls per two minute stop over the 20 mile route has been 18.9 (Knapp 1979), ranging from a low of 14.5 in 1978 to a high of 24.4 in 1977. Waterfowl Seven species of waterfowl game birds were observed on the Otter Creek study area (Table 17). Of these only the mallard was observed to breed in the area. It is possible that several of the others also breed within the study area, especially in good water years. Both spring seasons of this study were marked by drought conditions with 1980 being the most severe. Most of the reservoirs were dry while Otter Creek and some spring- fed ponds had water. Songbirds One hundred and ten species of birds, including game birds, were observed on the study area (Table 17). The Swainson's hawk. Cooper's hawk, golden eagle, bald eagle, prairie falcon, upland sandpiper, burrowing owl, long-eared owl, mountain bluebird, clay-colored sparrow, Brewer's sparrow, and field sparrow were listed by Flath (1979b) as species of special interest or concern. All of these except the bald eagle and burrowing owl, both listed as migratory birds of high federal interest (U.S. D.I. 1979), were known or suspected to breed in the study area. Habitat suitable for breeding burrowing owls existed in the study area, so breeding for this species is possible in the future. Four breeding bird census strips were located on the Otter Creek study area, one each in grassland (without sagebrush), sage- brush, ponderosa pine, juniper, and boxelder-ash riparian habitats (Figure 16). Tables 18-21 list the percent composition and average number of birds observed per census run for each of the bird species found on the four breeding bird census strips. The average number and the highest number of singing males observed on each strip are also listed along with breeding pair density estimates derived from them. Estimates were not made for red crossbills, which were present as large, non-breeding flocks during the censusing. Western meadowlarks dominated the grassland and sagebrush census strips. Chipping sparrows, lark sparrows, and western wood pewees were the three most abundant species on the ponderosa pine census strip. House wrens and yellowwarblers were the two 35 Table 17. Breeding status of the bird species observed on the Otter Creek study area. Species Breeding Status Species Breeding . Status 1. Pied-billed grebe 2. Great blue heron 3. Whistling swan 4. Canada goose 5. Mallard 6. Gadwall 7. Pintail 8. Green-winged teal 9. Blue-winged teal 10. Northern shoveler 11. Turkey vulture 12. Sharp-shinned hawk 13. Cooper's hawk 14. Red-tailed hawk 15. Swainson's hawk 16. Rough-legged hawk 17. Golden eagle 18. Bald eagle 19. Marsh hawk 20. Prairie falcon 21. American kestrel 22. Sharp-tailed grouse 23. Ring-necked pheasant 24 . Turkey 25. American coot 26. Killdeer 27. Common snipe 28. Upland sandpiper 29. Spotted sandpiper 30. Solitary sandpiper 31. Willet 32. Black tern 33. Mourning dove 34. Yellow-billed cuckoo 35. Black-billed cuckoo 36. Great horned owl 37. Burrowing owl 38. Long-eared owl 39. Poor-will 40. Common nighthawk 41. White-throated swift 42. Belted kingfisher 43. Common flicker 44. Red-headed woodpecker 45. Hairy woodpecker 46. Downy woodpecker 47. Eastern kingbird 48. Western kingbird 49. Cassin's kingbird 50. Says phoebe Bi/ 51. t 52. M 53. M 54. B 55. M 56. M 57. M 58. M 59. M 60. b 61. b 62. b 63. B 64. b 65. W 66. B 67. M 68. b 69. B 70. B 71. B 72. B 73. b 74. b 75. b 76. b 77. b 78. b 79. M 80. M 81. M 82. B 83. b 84. b 85. B 86. M 87. B 88. B 89. b 90. b 91. b 92. B 93. B 94. B 95. b 96. b 97. B 98. b 99. B 100. Least flycatcher b Western wood pewee b Horned lark b Violet green swallow b Tree swallow b Barn swallow B Cliff swallow B Black-billed magpie B Common crow b Pinyon jay b Black-capped chickadee b White-breasted nuthatch b Red-breasted nuthatch b House wren B Canyon wren b Rock wren b Gray catbird b Brown thrasher b American robin B Mountain bluebird B Townsend's solitaire b Water pipit M Bohemian waxwing W Loggerhead shrike b Starling b Solitary vireo b Warbling vireo b Yellow warbler B Yellow-rumped warbler b Ovenbird M Common yellowthroat b Yellow-breasted chat b House sparrow b Western meadowlark B Yellow-headed blackbird M Red-winged blackbird b Northern oriole B Brewer's blackbird b Common grackle B Brown-headed cowbird b Western tanager b Black-headed grosbeak b Lazuli bunting b Evening grosbeak W Cassin's finch W Pine siskin W American goldfinch b Red crossbill b Rufous-sided towhee b Lark bunting b 36 Table 17 continued. Breeding Breeding Species Status Species Status 101. Grasshopper sparrow b 106. Chipping sparrow B 102. Vesper sparrow B 107. Clay-colored sparrow b b 106. B 107. B 108. b 109. 103. Lark sparrow B 108. Brewer's sparrow b 104. Dark-eyed junco b 109. Field sparrow b 105. Tree sparrow W 110, White-crowned sparrow M ]^/ Breeding status B - Hard evidence of breeding b - Circumstantial evidence of breeding t - Occurs, but no evidence of breeding W - Overwintering observations M - Migratory observation most abundant species on the riparian census strip. The riparian census strip supported the highest number of species and breeding bird density, followed by the ponderosa pine census strip. The sagebrush census strip supported the lowest number of species and breeding bird density, although the grassland census strip supported densities nearly as low. The higher number of species observed on the grassland strip than on the sagebrush strip was probably due to greater proximity of the grassland census strip to ponderosa pine habitat. Raptors Eleven species of hawks and three species of owls were observed on the study area. Inconspicuous species such as screech owls and saw-whet owls may also have occurred in the study area. Goshawks were observed within ten miles of the study area, and suitable nesting habitat for them was present in the study area. This species may have been overlooked because of poor visibility in the dense forests it prefers. Accipiters: Cooper's hawks were observed twice in the study area in 1980 (Figure 17). One of the observations was during the nesting season (June). One sharp-shinned hawk was observed on 1 June 1980. Both of these species probably nested in the study area. 37 LEGEND Permanent stream '^^^^ Paved road Unpaved road — = Forest boundary Study area -> >- Bird census strip D Small-mammal frapline. Q^* Grassland G «, Sagebrush. . S ^* Ponderosa pine P ^.° Riparian R Figure 16. Location of songbird census strips and snail nammal traplines on the Otter Creek study area. 38 C 3 i Di m w o 1— 1 m a; 3 00 CO O 03 CO CO -d • rH (0 Oi 1— I u O rf c rH ■H IZ S (D JD -P M ^-t W C >> 0 OJ -H c £1 M d O) >. bJ3 C t> rH iM 00 in CD +-> • ■^^ -H C O) r-i rH CO •H Oj ffi K) -H 1—1 m 0) c u (D cij TJ c >. m 3 U 73 a; ci o f- cr: O o •H 3 . r-l. 00 • • • • nS +-> O rt w Oi I> O CD CO l> a en ^; s 3 I—I rH M^ 0 bo C C 0) bfl C 0) •H (D rt -H u 73 ^ ^H b.0 03 O 00 ^ O CM 0) U (D C ^ t> • • • • • CJ > -H (D C5 in t^ rp CO O ^ ;^ < CO Ph 1— 1 CM ^ (n +J ■O 4-> c o rt 0) "x: O C +J 00 CDOrHCO^rH"^ r-{ CSl '^CO CM M o c O) '^ CM rH c •H C o rH rH +J -n^ ■H •r- +-I -P CT, a. 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OQ ^fetJCCKCJCQSK H 73 0 C •H oj -P +J X c 0 0 O CO ^ -P 0 c a 3 o 0 o c o o c c rt - x: +-> +j c 0 CO CO CO 0 0 u ►J a a 43 B C 1 D E F c; H i LEGEND Permanent stream Paved road Unpaved road Forest boundary . . Study area Sightings: Cooper's hawk . . Swainson's hawk Long -eared owl . Burrowing owl x aA/^ Fi^w Paved road Unpaved road Forest boundary Study area Golden eogle nests . Nos. 12-1 Sightings: 1979 0 1980 Figure 19. Nests and sightings of golden eagles on the Otter Creek study area. 47 LEGEND Permanent stream . . ^w Paved road Unpaved road — ^ Forest boundary Study area -• >- Nests Nos.lO- Sightinqs o Probable nest KJ «. Fif^ure 20. Nests and sightings of prairie falcons on the Otter Creek study area. 48 o 00 I Q -p o W) ^ 0 ^ >> a n o (D 0 cu c -a > > > J -g ■P -P q o Ci 03 Cj c c c ■P 0 (D > > ■H -H P' P> O O C C fcfl bjo bJD 43 pi rti I 0) I ^ C P o o •H >> pi 0 •* O (D ^f O M C tp 0 1 I > 0 (P -H > > p ■H -H O P> P> d o o c pi ;3 tp o o pi CO O 3 >> o 'H -s o 73 13 C M rt o -Si 0 1 > > 0 •p •H > pi P> •H o O Pi rt c^ O c a 0 0 > > •H -H P> Pi O O piLiP^MhH<;p(p{pi<;<;i— i •H P O < CD C5 M R pi pi fa£ O CO M rH 0 0 ■a > x) > -d 0 -H 0 -H 0 bp pi hppi •^ q "o o 0 d 0 oj C rH G o q pLHi-(pHi-(p!H 0 c o CO 0 p> •H CO o p> 0 P a u p> I en CO 0 •H o CO 2^8 pi p> 0 0 0 0 U U P> P" OPp O O Of-iPrHiPrH ^^s^ii^j^;^^^^ x: x; ^ % S S 3 S •5 S ^ "ij ^ "^ TO Co 03 eg TO TO x: x: x: i2 x: x: c c o o 0 0 0 0 0 '8^'8'g'S'SI'g'S^^^ ^^^^^ 0 0 0 0 0 H -H -H -H -H -H 0 0 d •H -H -H d d d d d Cti P P P Pi P P> I I I I I I I "O "p X3 t3 73 T3 T) pi ;^ ;-i I T3 C C C C C 0 0 0 0 0 T3 TJ T3 T3 T3 Oj d 1— I 1— I i-H tp ip ^^^^^^^i^^^S88S3 CM CM ■§ H P CO IP in rHCN)COrtiinCDt>00(j50rH(MCO'^"l>CO 49 Non-game Mammals Twenty-nine species of mammals were observed in the study area (Table 23). Mink [Huite-la vlion] and yellow-bellied marmots {Hafimota llavlv z.ntnli>] were observed in the region and probably occurred in the study area. One dead hoary bat was found in riparian habitat. Other species of bats undoubtedly were present in the study area. The blacktail prairie dog was listed by Flath (1979b) as a species of special interest or concern. Two small towns were located (Figure 21). Table 23. Mammals observed on the Otter Creek study area, 1 . Masked or common shrew 2. Hoary bat 3. Raccoon 4. Shorttail weasel 5. Badger 6. Striped skunk 7. Coyote 8. Red fox 9. Bobcat 10. Blacktail prairie dog 11. Thirteen-lined ground squirrel 12. Least chipmunk 13. Red squirrel 14. Northern pocket gopher 15 . Wyoming pocket mouse 16. Beaver 17. Western harvest mouse 18. Deer mouse 19. Bushytail woodrat 20. Meadow vole 21. Prairie vole 22. Muskrat 23. House mouse 24. Porcupine 25. Whitetail jackrabbit 26. Desert cottontail 27. Mule deer 28. Whitetail deer 29. Antelope La6iafia6 cim>ie.LLi Pfioctjon totoK Ha6te.la t^mlnza Taxidza taxui He.phiti6 maphltl^ Cani6 latfian6 \/iilpe.6 vulpe.6 Lynx fiaffU'S Cijnomij6 ludovlci.anui Sp2.fimophilu6 tfi-Ldzczmlimatuii Eutam-la^ ml.nln}Li6 Tam.la6c^ufiui hLid6oni(iui> Thomomij.!) talpoldei, PtfLognathai, {^aiclatai) Ca&tofi canade,ni,i6 Re.ith^odontomij6 rmgatotii ?(i^omtj.{>cu6 manicatatui, tizotona cimfidCL M^C'lotui penn6 ijlvanicui H-icfLotiL6 ochfioga^t^fi Ondatfia zibethicui Mai mu6culu6 Enzth-Lzon do liatum Lzpai toMnizndi-i Sijtvilagai aadabonii OdacoZlzui htm-ionu6 Odocoileu6 vA.n^gX.nZanU't) Ant-Lloaapfici amzK-icana. m Six small-mammal traplines were run in the study area, one each in grassland, sagebrush, ponderosa pine-juniper, and marsh riparian habitat, and two traplines in boxelder-ash riparian habitat (Figure 16). The marsh riparian habitat consisted of a cattail [Typha], bulrush [SalfLpui] area around a stockpond. Table 24 lists the total trap nights, trapping success, and numbers of each species caught in each habitat. The highest 50 number of species were caught in marsh riparian habitat and the highest trapping success was in boxelder-ash riparian habitat. The trapline in sagebrush habitat met with very poor trapping success. This may not have been typical of all sagebrush habitat in the study area. Three of the ?ziomyi,ca6 captured in riparian habitat were suspected to be white-footed mice (P. £eacopa4 ) . They were counted as P. manicalata6 in the data until identification of the specimens is positively verified. Table 24. Results of small-mammal trapping on the Otter Creek study area. Ponderosa Boxelder- Grass- Sage- Pine Ash ^larsh land Brush Juniper Riparian Riparian Total captures Trap nights Captures/100 trap nights Number of species caught Species: VoAomtjiiCuA, manic-uXaXvU) K^ltlTAodontomiji mzQatotli) HicfLotui p2.nn6ytvayu.ciU> f^UcAotui odhfioqaiit^n. Mai ma6cu£a6 SpzumopkiZuui) tfujd.(ic.2mZA,n2.atvu> Eutam^a^ minlmLii Vzn.0Qnathiii> {^aicAOtiUi SofLZx clneAzuA 11 448 2. 2 12 63 14 486 468 806 541 0.4 2.6 7. 8 2.6 2 2 4 5 1 7 29 2 1 ' 2 1 22 6 6 Amphibians and Reptiles Four species of amphibians and six species pf reptiles were seen on the Otter Creek study area (Table 25). The snapping turtle was listed by Flath (1979b) as a species of special interest or concern. Snapping turtles were fairly common along Otter Creek. Only one sagebrush lizard was encountered, in ponderosa pine-juniper habitat with large boulders. The other species were common to abundant in the study area. LEGEND Permanent stream . . '^aa- Paved road Unpaved road ' — Forest boundary Study area -> <■ Prairie dog towns c? Fif^ure 21. Prairie dog towns on the Otter Creek study arcm. 52 Table 25. Amphibians and reptiles observed on the Otter Creek study area. 1. Woodhouse toad 2. Tiger salamander 3. Northern chorus frog 4. Leopard frog 5. Snapping turtle 6. Painted turtle 7. Sagebrush lizard 8. Racer 9. Bull snake 10. Prairie rattlesnake Ba{)0 woo dhou-itA. Ambif-itoma tigfiZnum P6zadacKi6 t>i-ii>(i>ilata Rand plplzn^ Ch^tijdna iitfipzYitlna. ChfiijfiZmyii picta SdztopoKViii gfLacioia6 Colubzfi con&tfi-ictoit P-ituophl.!, catzni{)t/i Cfiotatai, vifildii Hanging Woman Creek Mule Deer Population characteristics: During the fifteen months of field work, 1830 mule deer were observed in the Hanging Woman Creek study area (Table 1). Group size averages were largest during the winter season, 10.2, and smallest during the summer time, 2.0. Deer gradually separated from their winter herds during April and May in both years reaching their smallest sized groups in June. Winter herds were formed in December. Hanging Woman Creek supported the highest density of mule deer of the three study areas. Observations per hour of aerial survey ranged from a summer low of 19.4 to a winter high of 74.8 (Table 26). During the January flight, 144.4 mule deer were seen per hour of flight. The highest number seen in Otter Creek per hour was 55.0 (Table 2). The population structure (Table 27) shows a slightly lower percentage of fawns than observed in the Otter Creek area. Does comprised 47.8 percent of the population in October. This is slightly higher than the 41.7 percent observed in the Otter Creek area (Table 3). While the fawn production was lower than that observed on Otter Creek, it was still very good at 92.2 fawns per 100 does. This area has been classified as having stable population levels (Swenson 1978) of mule deer in low nutritional condition (Swenson and Knapp 1979). These data seem to indicate an under-harvested mule deer herd. Distribution: Spring mule deer distribution is shown in Figure 22. The deer are fairly evenly distributed throughout the study area. The largest grouping of observations stretches across 53 LEGEND Permanent stream -'^^ County road State line Study a rec Mule deer observations Group size 1 ~ 3 A O 4-8. A o 9-15 A e 16+ A • Year 1979 1980 MONTANA WYOMING Figure 22. Spring mule deer distribution in the Hanging Woman Creek area. 54 Table 26. Mule deer, antelope and coyote aerial observations per hour in the Hanging Woman Creek area. Month Hours Aerial Observations Mule Deer Ant . Coyotes Observat ions/Hour Mule Deer Ant. Coyote Coyot Mule Deer es per 100; Antelope April May 2.25 2.25 143 123 110 146 5 2 63,6 54.7 48.9 64.9 2,2 0.9 3.5 1.6 4.5 1,4 Spring 4.5 266 256 7 59.1 56.9 1.6 2,6 2.7 June July Aug 2.3 2.3 2.1 33 46 46 61 197 121 3 5 3 16.5 20.0 21.9 26.5 85.7 57.6 1.3 9 '^ 1.4 7.9 10.9 6.5 4.9 2.5 2.5 Sumner 6,7 ISO 379 11 19.4 56.6 1.6 8.5 2.9 Sept Oct Nov 2.3 2.25 -y 89 135 191 237 14 5 38.7 60.0 83.0 105.3 6.1 2.2 15.9 3.7 7.3 2.1 Fall 4.55 224 428 19 49.2 94.1 4.2 8.4 4.4 Dec Jan Feb 1.9 1.6 2.1 65 231 123 449 274 381 2 4 6 34.2 144.4 58.6 236.3 171.3 181.4 1.1 2.5 2.9 3.1 1.7 4.9 0.4 1.5 1.6 Winter 5.6 419 1104 12 74.8 197.1 2.1 2.9 1.1 March April May 2.9 2.3 2.0 151 199 131 298 138 136 4 3 02/ 52a 86.5 65.5 102.8 60.0 68.0 1.4 1.3 0.0 2.6 1.5 0,0 1,3 2.2 0.0 Spring 7.2 431 572 7 66.8 79.4 1.0 1.5 1.2 June July Aug 2.3 87 119 4 37.8 51.7 1,7 4,6 3.4 Sunmer 2.3 87 119 4 37.8 51.7 1.7 4.6 3.4 1/ No surveys made 2/ No observations made on surveys Table 27. Mule deer population characteristics in the Hanging Woman Creek area in 1979. Month Number Classified Total Bucks Does Fawns Fa^vnsrlOO Does Adults Bucks: 100 Does Population Structure (%) Bucks Does Fawns September October 77 8 38 31 134 11 64 59 81.6 67.4 92,2 78.7 21.1 17.2 10.4 49.4 40.3 8.2 47.8 44,0 the ridse from Waddle Creek to Corral Creek west of Hanging Woman Creek. Another major group was observed east of Hanging Woman between Horse Creek and Trail Creek. V/hile deer were still observed in most portions of the study area, a noticeable converging on Hanging Woman Creek occurred during the summer season (Figure 23). Mule deer were staying close to their primary water supply, i.e. Hanging Woman Creek. The greatest concentration of observations was located east of Hanging Woman Creek south from Horse Creek through the mouth of Trail Creek to the southern boundary of the study area. Practically all the observations were in the 1-3 group size category. The number of fall observations is greatly reduced from spring and summer because group sizes are bigger and only one year's data were gathered (Figure 24). The greatest numbers observed were in the vicinity of coordinates 0, 26. Most observations were in the central and southern portions of the study area. Winter observations are mapped in Figure 25. Four wintering areas are evident. The largest, both in numbers of deer and observations, is located along the ridge between Horse Creek and Trail Creek. The second largest group is found astride Corral Creek in the southwest portion of the study area„ Two smaller groups, one north of Horse Creek and one astride Trail Creek in the southeast corner, make up the remainder of the mule deer wintering areas. Practically all of the observations were of large groups of mule deer. Vegetation type usage: The ponderosa pine type was most heavily used in the spring of 1979 when 48 percent of the mule deer were observed in that type (Table 28). The winter season saw 44 percent of the deer in the pine subtypes. The least number of deer were observed in the ponderosa pine type during the fall season. Sagebrush/grasslands were most often used by mule deer in all seasons and averaged 56.8 percent over the course of the study. Seventy-three percent of the deer were observed in the sagebrush/grassland type in the spring of 1980. This type covered the largest amount of surface in the study area (Figure 4) . Creekbottoms were extremely important during the summer and fall months. This type, with a very small percentage of surface area, accounted for 17 and 23 percent of the summer observations in 1979 and 1980 respectively. During the summer of 1980, the driest season of the study, fully 30 percent of the mule deer observations were made in creekbottom subtypes. This includes the agricultural creekbottom type which is primarily hay fields. 56 LEGEND Permanent stream ^^^ County road State line Study area -< >- ^ad*^^ Mule deer observations Group size 1-3 A 4-8. A 9-15 * 16+ A Year 1979 1980 Figure 23. Sumner nule deer distribution in the Hanging Woman Creek area. 57 LEGEND Permanent stream -'^^ County road State line Study area Mule deer observations Group size 1 ~ 3 A o 4-8. A o 9-15 A 9 16+ A Year 1979 1980 -MQNIANA WYOMING Figure 24. Fall mule deer distribution in the Hanj^^ing Woman Creek area. 58 LEGEND Permanent stream -'^^ County road State line Study area Mule deer observations Group size 1-3 -^ o 4-8. £^ O 9-15 A e 16+ A • Year 1979 1980 MOJilAJvlA. WYOMING mi les Figure 25. Winter nule deer distribution in the Hanging; Wonan Creek area. 59 Activity: The lack of ponderosa pine cover made deer easier to observe as they lay out in the open„ For that reason more deer were observed lying in the Hanging Woman study area than in the Otter Creek area (Table 29). Most deer were observed standing in all seasons except spring 1980 when more deer were observed feeding. Use of topography: Use of the dissected mid-slopes was the highest in every season (Table 30). The highest percentage of observations using the hillside category was during the winter season, 89 percent. This topographic area is most likely to offer shelter from the wind and, at the same tine, be free of snow cover. The lowland features, alluvium/terrace and flood plains, had peak use during summer and fall seasons. Deer were forced onto the riparian bottom lands in search of water and succulent forage. These types received heaviest usage, 31 percent, during the driest season, summer of 1980. Use of exposure: Deer were observed on every aspect in every season (Table 31) . They were fairly evenly distributed with a few exceptions. During the spring 1979 season 48 percent of the deer were observed on southerly slopes. This occurred just after a long hard winter and the southern slopes were the first to become free of snow and offer new growth to eat. The summer seasons were marked by substantial increases in the usage of flat, lowland categories. Again, deer were reacting to the lack of moisture in the upland areas. The mild nature of the 1979-80 winter was demonstrated by the 62 percent usage figure on northerly slopes. Use of slope: Seasonal use of slopes by mule deer in the Hanging Woman Creek area is presented in Table 32 » Usage of steep slopes was highest during the winter months when 32 percent usage was observed. The lowest percentage was observed during the summer of 1980, 6 percent. More deer v/ere observed on medium slopes than any other in all seasons except winter when gentle and steep slopes had more deer. Medium slopes averaged 41 percent of mule deer observations during the study. Flat land areas received the most during the two summer seasons with 28 and 37 percent for 1979 and 1980 respectively o Antelope Population characteristics: During the six seasons of study, 3,308 antelope were observed on the Hanging Woman Creek area (Table 9). Average group size ranged from 3.4 antelope per group during the first spring season to 44.2 antelope per group in the winter months. January was by far the largest when 3 groups averaged 91.3 antelope per group were observed. 60 • Table 28. Seasonal use of vegetation types by mule deer in the Hanging Woman Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Vegetation Subtypes 3241/ 144 227 419 612 103 Ponderosa pine Sagebrush Grassland Skunkbush Juniper Snowberry Total Ponderosa Pine Sagebrush Deciduous shrub Grassland Total Sagebrush/grassland Tree Shrub Total Creekbottom Ponderosa pine Sagebrush /grass land Creekbottom Total Agricultural - 4 3 10 2 5 3l2/ 10 4 26 14 6 6 3 5 - 6 6 tr3/ 3 4 2 1 1 11 3 - 5 2 7 - - 1 - - - 48 24 17 44 25 24 40 28 40 34 62 26 — — — 15 6 9 7 28 26 7 5 8 47 57 65 56 73 43 _ 3 8 _ .» 4 3 15 9 - 2 19 3 17 18 - 2 23 - 1 - - - 3 2 1 — — - 7 10 l_l Total observations 2_/ Percent of observations rounded to nearest whole number 3/ tr = a percentage 0.5 Table 29. Seasonal activity of mule deer in the Hanging Woman Creek area. Spring Summer Fall Winter Spring Summer 1979 3241/ 1979 1979 '79-80 1980 1980 Activity 144 227 419 611 103 Standing Running Lying Feeding 422/ 67 45 63 34 50 17 10 9 6 8 21 15 2 36 21 10 6 27 21 10 10 48 22 l_l Total observations 2/ Percent of observations rounded to nearest whole number 61 Table 29. Seasonal activity of nule deer in the Hanging Woman Creek area,, Spring Summer Fall Winter Spring Summer 1979 1979 1979 •79-80 1980 1980 Activity 3241/ 144 227 419 611 103 Standing 422/ 67 45 63 34 50 Running 17 10 9 6 8 21 Lying 15 2 36 21 10 6 Feeding 27 21 10 10 48 22 j^/ Total observations 2/ Percent of observations rounded to nearest whole number Table 30. Seasonal use of topography by mule deer in the Hanging Woman Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Topography 3241/ 144 227 419 611 103 Mesa-Butte top 71/ 8 6 1 8 14 Mesa-Butte Steep 11 - 5 4 3 1 Slopes Dissected mid-slopes 73 72 70 89 79 55 Alluvium/terrace 5 4 3 6 7 15 Flood plain 3 15 16 - 3 16 \l Total observations 2/ Percent of observations rounded to nearest whole number 62 Table 31. Seasonal use of exposure by mule deer in the Hanging Woman Creek area. Spring Summer Fall Winter Spring Summer 1979 324-/ 1979 1979 •79-80 1980 1980 Exposure 144 199 419 611 103 North 13 19 19 18 13 16 South 29 6 4 8 9 14 East 4 6 4 13 14 6 West 10 13 25 5 10 6 Northeast 7 12 12 25 12 10 Northwest 8 5 2 19 7 3 Southeast 5 5 2 3 9 9 Southwest 14 6 12 2 17 1 Flat 10 28 20 7 8 37 1_/ Total observations 2/ Percent of observations rounded to nearest whole number Table 32. Seasonal use of slope by mule deer in the Hanging Woman Creek area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Slope 3241/ 144 199 419 611 103 Flat 10 28 18 7 9 37 Gentle 21 19 22 34 36 9 Medium 50 32 45 26 43 49 Steep 19 21 15 32 12 6 ]_/ Total observations 2/ Percent of observations rounded to nearest whole number 63 Antelope were more dense than nule deer in the Hanging Woman study area (Table 26) with greater numbers observed in every season except the initial spring survey. The number of antelope observed per hour of flight ranged from the 50 's in spring and summer to a whopping 197,1 during the winter season. In December, 449 antelope were observed on the study area in a little less than 2 hours flight time. That this area is a major antelope winter range is undeniable. and production figures are shown in tage of fawns in the population in 1979 34.6 percent) and decreased in each oduction figures (1979) were also 67.4 fawns per 100 does were observed, oes were observed in June of 1980. The conducted before most fawns were born, res compare to an average of 61.3 fawns Wildlife and Parks hunting unit 742 8 (Went land 1979). The Hanging Woman ontained within hunting unit 742. Popu lation structure Tabl e 33. The percen was highest in July ( succeeding month. Pr high est in July when Sixt y fawns per 100 d June 1979 flight was These production figu per 100 does in Fish, from 1963 through 197 Cree k study area is c Table 33. Antelope population characteristics in the Hanging Woman Creek area. Population Structure Total Bucks Does Fawns: 100 Fawns Does Adults Bucks: 100 Does Percentage Month Bucks Does Fawns 1979 June 93 22 65 6 9.2 6.9 33.9 23.7 69.9 6.4 July 240 34 123 83 67.4 52.9 27.6 14.2 51.3 34.6 Aug 138 22 73 43 58.9 45.3 30.1 15.9 52.9 31.1 Sept 201 40 101 60 59.4 42.6 39.6 19.9 50.2 29.9 1980 June 122 34 55 33 60.0 37.1 61.8 27.9 45.1 27.0 Distribution : Antelope spring distribution is shown in Figure 26. Antelope were observed throughout the study area with groups fairly evenly distributed within the size categories. A concentration of observations is evident just north of Trail Creek. Another group of observations is centered in the vicinity of Horse Creek. The entire western portion of the study area from above PK Creek south to Waddle Creek was covered with observations. 64 Permanent stream County road State line Study area Antelope observations Group size 1 ~ 3 A o 4-8 A o 9-15 A ffl 16* A Year 1979 1980 MOMIANA_ WYOMING Figure 26. Spring antelope distribution in the Hanging Wonan Creek area. 65 During the summer months, the antelope used less of the Hanging Woman Creek study area as they condensed into three major use areas (Figure 27). One large group of observations was made in the northeast portion of the study area north of Horse Creek. Another group was concentrated in the First Creek and PK Creek drainages. The third major area was located along the north side of Trail Creek and extending northward along the east side of Hanging Woman Creek approx- imately 2 miles north of the mouth of Trail Creek. Very few observations were made in the Corral Creek drainage. Fall antelope observations revealed a major shift in antelope usage patterns (Figure 28). Many observations were made in the Corral Creek drainage. Antelope were noticed to be staying closer to the main water sources, PK Creek and Hanging Woman Creek. There was a definite shift to the south and west portions of the study area. Winter antelope observations are shown in Figure 29. The major wintering area is seen to be along the north side of Trail Creek extending northward along the east side of Hanging Woman Creek. The PK Creek area is another wintering range. Antelope were also observed north of Horse Creek and between Corral and First Creeks. This was a mild winter with very little snow cover. Therefore the antelope were able to pick which ever area suited them. They may have used entirely different areas under more harsh circumstances. Practically all of the observations were of large groups of antelope. Vegetation type usage: The fact that antelope are heavily dependent on sagebrush is evident in Table 34. Antelope usage of the sagebrush/grassland type averaged 91 percent and ranged from 81 to 99 percent during the six seasons of study. Antelope were observed in ponderosa pine subtypes in all seasons but in very low numbers. A significant percentage, 15 percent, were observed on the agricultural creekbottom type during the fall season. These antelope were on harvested hay meadows near water . Activity: The activity of antelope at the time of observation is recorded in Table 35. More antelope were observed standing in the spring seasons, 34 percent in 1979 and 45 percent in 1980, than the other four seasons which ranged from 22 to 28 percent. The fewest numbers were observed running during the spring season as antelope seemed to be conserving energy. The relatively large percentage of antelope observed feeding and lying down in the 1979 spring is indicative of the very severe winter just completed. No antelope were observed lying down or feeding during the winter aerial surveys. Use of topography: Antelope were most often observed in the rolling portions of the dissected midslopes (Table 36) and least often seen on steep mesa-butte hillsides. A few Table 34„ Seasonal use of vegetation types by antelope in the Hanging Woman Creek area. Total Sagebrush/grassland 94 92 81 93 90 Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Vegetation Subtypes 3141/ 471 524 1104 720 175 Ponderosa pine _ 1 tr ^ ^ _ Sagebrush l2/ 2 3 2 4 1 Grassland 3 1 - - 3 1 Skunkbush - 1 - - - - Juniper - trl/ - - - - Snowberry - - - - - - Total Ponderosa pine 4 5 3 2 7 1 Sagebrush 56 50 37 67 55 45 Deciduous shrub - - - - - - Grassland 39 42 44 26 35 54 99 Tree Shrub Total Creekbottom Ponderosa pine Sagebrush/ grass land Creekbottom Total Agricultural 15 15 2 tr tr J_/ Total observations 2/ Percent of observations rounded to nearest whole number 3/ tr= trace; a percentage 0.5 Table 35. Seasonal activity of antelope in the Hanging Woman area. Activity Spring Summer 1979 1979 314i./ 471 Fall Winter 1979 '79-80 524 1104 Spring 1980 720 Summer 1980 175 Standing 342/ 28 26 25 45 22 Running 19 51 56 75 30 39 Lying 21 9 tr3/ — 7 7 Feeding 26 12 19 - 18 23 !_/ Total observations 2/ Percent of observations rounded to nearest whole number 3/ tr = trace; a percentage 0.5 67 LEGEND Permanent stream ^^^^ County road =— State line Study area Antelope observations Group size 1 "" 3 A O 4-8 A o 9-15 4 • 16* A Year 1979 1980 MQMIANA WYOMING Fif^uro 27. Sumner antelope distribution in the Hans^ins Woman Creek area. 68 LEGEND Permanent stream County road State line Study a rea Antelope observations Group size 1 ~ 3 A o 4-8 A G 9-15 A « 16* A Year 1979 1980 MONTANA WYOMING Figure 28. Fail antelope distribution in the Hanging Woman Creek area. 69 Permanent stream County road State line Study area Antelope observations Group size 1-3 4-8 9-15 16* Year LEGEND A o A o A e A • 1979 1980 MONTANA WYOMING Fif^ure 29. V/inter antelope distribution In the Hant;-ins? Woman Creek area. 70 antelope were observed on the plateau tops in all seasons with the most, 13 percent, occurring during the winter months. Antelope were also seen on the lowland topographic features during all seasons. The fall and winter seasons, with 18 and 15 percent of the antelope respectively, had the most antelope observations of the lowland topographic categories. Use of exposure: Antelope were observed on every exposure in every season except the north exposure during the winter months (Table 37). There were more antelope on southerly slopes than on northerly slopes in every season. Forty-two to 51 percent of the antelope were on the southern exposures in all seasons except summer 1980 when 70 percent were observed on the southerly slopes. Usage of the northerly slopes was relatively uniform, ranging from 19 to 32 percent. There were more antelope on easterly exposures than westerly in every season. Usage of flat lands was highest during the fall and winter months. Use of slope: Antelope use of gentle and medium slopes was predominant throughout the study (Table 38) ranging from 82 to 97 percent of total observations. In the summer of 1989 medium slope usage was at its highest point, 66 percent and gentle slope usage was at its lowest point, 31 percent. Their positions were reversed in the 1979 spring when 51 percent of the antelope were observed on gentle slopes and 44 percent were seen on medium slopes. Flat land usage was low in all seasons except fall with a 16 percent usage level. The only season with much usage of steep slopes was winter with 12 percent. Sharp-tailed Grouse Three active sharp-tailed grouse dancing grounds were located during the study (Figure 30). They average 14.7 male birds per ground in 1980 (Table 39). In an earlier study, Knapp (1977) reported three dancing grounds which were not located at this time. Sharp-tailed grouse have been documented as abandoning grounds in other southeastern Montana studies (Martin 1980). Sharp-tailed grouse were observed in several locations some distance from known dancing grounds (Figure 30). Because these grouse are known to remain close to dancing grounds it is possible that grounds exist near PK Creek in the northeast portion of the study area, somewhere in the Horse Creek drainage, along the ridge between Trail Creek and East Fork Trail Creek and along the South Fork Corral Creek drainage. 71 Table 36. Seasonal use of topography by Woman Creek area„ antelope in the Hanging Topography Spring Summer Fall 1979 1979 1979 3141/ 471 524 Winter '79-80 1104 Spring 1980 720 Summer 1980 175 Mesa-Butte top Mesa-Butte steep sides Dissected mid-slopes Alluvium/Terrace Flood plain 12/ 13 8 92 90 79 72 87 99 5 3 8 15 5 1 tr3/ 2 10 — _ _ !_/ Total observations 2/ Percent of observations rounded to nearest whole number 3/ tr = trace; a percentage 0.5 Table 37. Seasonal use of ex posure by ante] ope in the Hanging Woman Cree ■k area. Spring Summer Fall Winter Spring Summer 1979 1979 1979 '79-80 1980 1980 Exposure 3141/ 471 524 1104 720 175 North 142/ 9 4 11 9 South 9 17 23 6 20 41 East 17 23 12 22 8 6 West 5 4 6 3 5 2 Northeast 12 13 12 1 18 8 Northwest 6 4 3 20 1 5 Southeast 11 7 18 31 19 14 Southwest 25 18 6 6 12 15 Flat 2 4 16 12 6 1 1/ Total observations 2/ Percent of observations rounded to nearest whole number, « 72 LEGEND Permanent stream County road State line Study area -. ^ v>jadd\« Grouse Sharptail Sage Breeding grounds • Observations Winter b a Spring g o Summer o o Fal I a A _WQNIANA WYOMING Firjure 30. Sharp-tailed grouse dancin;; grounds, sage grouse strutting grounds and observations of both species during the study on Hanging Wonan Creek. 73 Sage Grouse Sage grouse were the most abundant game bird species in this area. Five strutting grounds were located on or near the Hanging Woman Creek study area (Figure 30). These are concentrated in the southern portion along the Corral Creek, Trail Creek and Hanging Woman Creek bottom lands. The largest of the grounds, the Corral Creek ground, had 73 birds (male and female) using it one morning during the 1980 breeding season. The average number of cocks observed at the four grounds within the study area boundary in 1980 was 23.9 birds (Table 40). Sage grouse were observed in all seasons during the study. The Corral Creek sagebrush bottoms and uplands appear to be an important wintering area for sage grouse. Twelve sage grouse broods, averaging 49 young per brood, were observed in the Hanging Woman study area in the summer of 1979. This represented good recruitment (Wallestad 1975) and coupled with an easy winter resulted in a large population increase o The number of cocks present on the two grounds observed both years increased from 36 to 72 (Table 40). This amounted to a 100 percent increase. Possibly the 1979 numbers observed were somewhat lower than the actual population levels. Ring-necked Pheasant Pheasants were observed along the Hanging Woman Creek bottom. A pheasant crow count survey was conducted during the 1979 breeding season. Birds were heard on 4 of the 12 stops. The average number of calls per stop was a relatively low 0.5. This low population density is probably the result of a scarcity of deciduous shrubs in the creekbottom vegetation types. Waterfowl Six waterfowl species were observed on the Hanging Woman Creek study area (Table 41). The most common species observed was the mallard. Only one brood of ducks was observed. That was a group of 13 young blue-winged teals with one adult feeding in a road side pond in the early summer of 1979. As in other portions of southeastern Montana, the Hanging Woman Creek area suffered through severe drought conditions in the spring and summer of 1980. Consequently the reservoirs were dry and offered little or no breeding habitat for waterfowl. Songbirds Seventy-eight species of birds, including game species, were observed on the study area (Table 41). The goshawk. Cooper's hawk, ferruginous hawk, golden eagle, bald eagle, prairie falcon, upland sandpiper, burrowing owl, mountain blue bird, and Brewer's 74 Table 38. Seasonal use of slope by antelope in the Hanging Woman Creek area. Slope Spring Summer 1979 1979 3141/ 471 Fall 1979 524 Winter Spring '79-80 1980 1104 720 Summer 1980 175 Flat Gentle Medium Steep 22/ 4 51 45 44 47 3 3 16 34 48 2 4 6 37 42 47 51 12 1 1 31 66 1 !_/ Total observations 2/ Percent of observations rounde d to nearest whole number Table 39. Sharp-tailed grouse dancing grounds in the Hanging Woman Creek study area and number of males attending in 1979 and 1980. Grnii nd Lo cation Males 1979 Ob served 1980 Number T R S 1 2 3 8S 8S 9S 43E 44E 43E 30 29 10 SEJ SWJ NW* 10 16 12 16 Table 40. Sage grouse strutting grounds in the Hanging Woman Creek study area and number of males attending in 1979 and 1980. Ground Lo cation No. of 1979 Mai es Observed Number T R S 1980 1 9S 43E 15 NWi 20 16 2 9S 4 3E 5 NEi 16 56 3 9S 4 3E 14 SEi _ 8 4 9S 44E 6 SEi « 15 5 9S 4 2E 13 SEi — 2 75 Table 41. Breeding status of the bird species found on the Hanging Woman Creek study area„ Species 1. Great blue heron 2. Mallard 3. Gadwall 4« Pintail 5. Green-winged teal 6. Blue-winged teal 7. American wigeon 8. Goshawk 9. Cooper' s hawk 10. Ferruginous hawk 12. Golden eagle 13. Bald eagle 14. Marsh hawk 15. Prairie falcon 16o American kestrel 17. Sharp-tailed grouse 18. Sage grouse 19. Ring-necked pheasant 20. Turkey 21. American coot 22. Killdeer 23. Common snipe 24. Upland sandpiper 25. Spotted sandpiper 26= Solitary sandpiper 27. Willet 28. Greater yellowlegs 29. Lesser yellowlegs 30. Wilson's phalarope 31 „ Northern phalarope 32 o Mourning dove 33. Great-horned owl 34. Burrowing owl 35. Poor -will 36. Common nighthawk 37. Wliite-throated swift 38. Belted kingfisher 39o Common flicker Breeding Status Species Breeding Status tl/ 40. b 41. M 42. M 43. b 44. B 45. M 46. b 47. b 48. b 49 „ B 50. M 51. b 52. B 53. B 54. B 55. B 56 » b 57. b 58. M 59. b 60. b 61. b 62, b 63. M 64. M 65. M 66. M 67. M 68. M 69. b 70. B 71. B 72. b 73. b 74. b 75. b 76. B 77. 78. Red-headed woodpecker b Hairy woodpecker b Eastern kingbird b Say's phoebe B Western wood pewee b Horned lark b Violet-green swallow b Barn swallow B Cliff swallow B Black-billed magpie B Common crow b Pinyon jay b Clark's nutcracker t Black-capped chickadee b House wren b Rock wren b Sage thrasher b American robin b Mountain bluebird b Sprague's pipit M Loggerhead shrike B Starling b Yellow warbler b Yellow-rumped warbler b Common yellowthroat b Western meadowlark B Red-winged blackbird b Northern oriole b Brewer's blackbird b Common grackle b American goldfinch b Red crossbill b Rufous-sided towhee b Lark bunting b Grasshopper sparrow b Vesper sparrow b Lark sparrow b Chipping sparrow b Brewer's sparrow b 1/ Breeding status: B = Hard evidence of breeding b = Circumstantial evidence of breeding t = Occurs, but no evidence of breeding W = Overwintering observations M = Migratory observation 7« sparrow were listed by Flath (1979b) as species of special interest or concern. The ferruginous hawk, golden eagle, bald eagle, prairie falcon and burrowing owl are also listed as migratory birds of high federal interest (U. S.D.I. 1979). All were suspected or known breeders in the study area except bald eagles. Two breeding bird census strips were located on the Hanging Woman Creek study area, one each in grassland (without sagebrush) and sagebrush habitats (Figure 31). Ponderosa pine-juniper and riparian habitats were not sampled on the Hanging Woman Creek study area. These habitats probably supported most of the same species as found in similar habitat on the Otter Creek study area. Tables 42 and 43 list the percent composition and average number of birds observed per census run for each of the bird species found on the two breeding bird census trips. The average number and highest number of singing males observed on each strip are also listed, along with breeding pair density estimates derived from them. The majority of the birds seen on the grassland census strip were western meadowlarks. Brewer's sparrows, vesper sparrows and western meadowlarks in nearly equal proportions dominated the sagebrush census strip. The sagebrush census strip supported fewer species, but a higher breeding pair density than the grassland census strip. Raptors Nine species of hawks and two species of owls were found on the Hanging Woman Creek study area. Short-eared owls were seen within ten miles of the study area. Inconspicuous species such as screech owls and saw-whet owls may have been overlooked. Accipiters : One goshawk was observed on the study area on 9 May 1980 (Figure 32). It was suspected to have nested in the area. Cooper's hawks were observed on several occasions along pine ridges, where they were probably nesting (Figure 32). One bird acted defensive, but the nest was not found. Buteos: Bald eagles were observed along Hanging Woman Creek twice in the early spring (Figure 32). They were both migrants. Ferruginous hawks were seen several times at the southern end of the study area in 1979 (Figure 32). They were suspected to be nesting in or near the study area. Red-tailed hawks were the most common buteo. Seven nests were active in 1979 and five nests were active in 1980 (Figure 33),, Eight nests were located in cottonwood trees and two were in ponderosa pine trees (Table 44). 77 en O >. d •H U m os c T3 (D ■H a a c c o •H ^ ■a (D fcUD CD C £1 he C TS oj C K erf 0) -X! 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Locations of bird census strips and snail nammal traplines on the Hanging Woman Creek study area. 81 LEGEND Permanent stream County road State line Study area Raptor nests Nos. 11-15 1979 Sightings • 1980 Sightings. . . x Gotden eagle Bald eagle Prairie falcon Burrowing owl MQMIANA Goshawk. '^'W'' Cooper's hawk. . . (O Ferruginous hawk ^/\ Finjure 32. Nests and sightings of raptors of special interest or concern on the Hanging Woman Creek study area. 82 Golden eagles were frequently observed on the study area. Three nests were located on cliffs in the same area (Figure 32). One of the nests was active in 1980 and fledged one female eagle (Table 44). Many other suitable nest sites existed for eagles in large pine trees on ridges and large cottonwoods in the creekbottoms. Sone nests may not have been detected due to the difficulty of finding stick nests in ponderosa pine forests. Most of the golden eagle observations were located around two ridges covered with ponderosa pine forest, indicating possible nests. Harriers : 'larsh hawks were common along the hay meadows on Hanging Woman Creek. They probably nested on the study area, but no nests were located. Falcons: One prairie falcon aerie was located on a cliff (Figure 32). Other suitable nest potholes were located on cliffs in the north end of the study area. Prairie falcons were usually seen hunting over the hay meadows along Hanging Woman Creek. Kestrels were the most common falcon in the study area. They mainly utilized dead pine snags found along ridges for nesting. Owls : Great horned owls were common in the study area. Two nests were found in 1979, one in an old magpie nest and the other in a pothole on a cliff. One pair of burrowing owls nested on a prairie dog town in 1979 and 1980 (Figure 32). They fledged five young in 1979. No brood counts were made in 1980. Another burrowing owl was sighted along a road at the north edge of the study area on 26 June 1980 (Figure 32). It may have been associated with a prairie dog town located just off the study area. Non-game Mammals Fifteen species of mammals were observed on the study area (Table 45). The blacktail prairie dog was listed by Flath (1979b) as a species of special interest or concern. Two small-mammal traplines were located in the study area, one in grassland and one in sagebrush habitat (Figure 31) . Table 46 lists the total trap nights, trapping success, and numbers of each species caught in each habitat. The sagebrush habitat supported the most species and the highest trapping success, Blacktail prairie dog towns are shown in Figure 34o Exact locations and approximate size are given in table 47„ All towns were searched for sign of black-footed ferrets with negative results. LEGEND Permanenf stream -^-^ County road State line Study area -. ^ vjodd^* Nests Nos.l-lO Sightings: 1979... D MQMIANA 1980 o WYOMING Fifcure 33. Nests and si'^htings of red-tailed hawks on the Hanging V/ornan Creek study areas. 84 LEGEND Permanent stream -^-^ County road State line Study area . ^^ Prairie dog town 0) O !-, - c E c 5. -o Z O (0 D 0) O jC «- Q. U O U1 OJ xi -p c o CO 0) d •H iH a a u -p 1— 1 Oj £ g d E i—i 1—1 a E m •c c rt m Ci, •H • U re! ■P OJ M u d 0) :3 >. in TJ c 3 0) +-> o W T3 ^ U Q •H OJ /3 ;h t_) tH o be O m Q c 0 QJ •H •H -P ^ rel •H O rt 0 u hJ Ph • CD CO tu ^ S bD Raptors Six species of hawks and one species of owl were observed on the study area. Inconspicuous species such as screech owls and saw-whet owls may have been overlooked. Goshawks and sharp-shinned hawks were observed within ten miles of the study area, and suitable nesting habitat for them was present in the study area. Turkey vultures were common, and there were many suitable nesting areas for them in the study area. Accipters : One Cooper's hawk was seen just outside the study area boundary on 11 June 1980 (Figure 37). It was nesting on or near the study area. Buteos : Two active red-tailed hawk nests were found on the study area in 1979. One of these was also active in 1980 (Figure 37). One nest was in a ponderosa pine tree and the other nest was in a cottonwood tree (Table 58). Harriers: Marsh hawks were seen several times hunting over sagebrush habitat. They may have nested in the study area. Falcons : Kestrels were the most common raptor in the study area (Figure 37) » One was active in 1979 and the other was active both in 1979 and 1980 (Table 58). Suitable nest potholes are present on several other cliffs in the study area. Ow 1 s : Great horned owls were occasionally seen in the study area. One nest was found in a pothole on a cliff in 1979. Table 58. Raptor nest sites on the Prairie Dog Creek study area. Nest Activity No. Species Substrate 1979 1980 1. Red-tailed Cottonwood 1 young fledged 1 young fledged hawk tree 2. Red-tailed Ponderosa 2 young fledged Status unknown hawk pine tree 3. Prairie Cliff Active - no Inactive falcon production data 4. Prairie Cliff Fledged 4 young Fledged 4 young falcon QQ X! 4-> C 0) c o E (1) a. 0) - ._ ^ c S !£ i ^ ^ .-. u u CO a 2: 2^ a o m a xi QJ a o o u -o c m c o o r-\ lA Ponderosa Pine Juniper 11 470 2.3 1 11 Deciduous Tree Riparian 12 406 3.0 3 1 4 7 101 Amphibians and Reptiles Three species of amphibians and two species of reptiles were observed on the study area (Table 61). All were common to abundant. Other species were probably present, but not encountered. Table 61. Amphibians and reptiles observed on the Prairie Do^ Creek study area. 1. Eastern toad 2. Woodhouse toad 3. Northern chorus frog 4. Racer 5. Bull snake BaffO Moodhou^^Z ?6e.adackfii6 tni^zKlata Colubufi coni>tKictofL P-ltuoph-is catznl^zn. SUMMARY AND RECOHMEinDATIONS The Hanging Woman Creek study area has the most abundant populations of big game animals, mule deer and antelope. Both species are quite dense and in good condition. Both species seem to be non-migratory and spend most of the year on relatively small home ranges. The Hanging Woman Creek area is excellent sage grouse habitat and supports a thriving sage grouse population, Sharp-tailed grouse and ring-necked pheasants are also present in fair numbers. While mule deer and antelope numbers in the Otter Creek area do not approach the levels in Hanging Woman, they are more than adequate. Many deer seem to commute into the Otter Creek study area from the surrounding Custer National Forest to feed, returning to the forest for cover and rest. The lack of ample expanses of sagebrush severely limit antelope numbers and distribution in the Otter Creek area. The lack of sagebrush is further evidenced by the fact that no sage grouse grounds were located in the study area. Otter Creek does provide excellent habitat for sharptail grouse and ring-necked pheasants and has large populations of these tv/o bird species. Prairie Dog Creek has no antelope or sage grouse populations. Only one sharp-tailed grouse dancing ground was found within one mile of the study area, none actually on the area. Mule deer were the most common game species observed and they were not seen in nearly half of the monthly aerial surveys. Prairie Dog Creek is easily the poorest of the three study areas in terms of game species diversity and density. ^Ci9. The study areas currently support good populations and numbers of species of raptors, songbirds, and sraall-mammals . Fifteen species of special interest or concern currently breed on or adjacent to the study areas. Efforts should be made to minimize the impacts of coal development on nongame wildlife through protection, restoration, and enhancement of habitat. Protection of feeding and roosting areas was important as well as protection of nesting areas. The following measures should be undertaken whenever possible to maintain, enhance or protect wildlife populations: 1. Exclude from leasing all sharptail and sage grouse breeding areas and appropriate buffer zones. These buffer zones should be at least one-half mile in radius or more to include nesting cover and wintering areas. 2. Exclude from leasing all major mule deer and antelope winter ranges. 3. Exclude from leasing all creekbottoms and associated "riparian" habitat types. Protection of "riparian habitat" which is defined as mesic areas with deciduous trees and shrubs, is very important. A large proportion of the riparian vegetation on the study areas is found on side drainages that are not currently protected as floodplains. Two very important examples are Home Creek (on the Otter Creek study area) and Prairie Dog Creek. Many other small patches of riparian vegetation are scattered on smaller drainages and by stockponds. Riparian vegetation comprises the smallest area of native vegetation types on the study areas. Its importance cannot be understated. It plays an extremely important role during the hot and dry summer and fall seasons in eastern Montana. In most instances, these riparian areas are the only source of water to both wildlife and domestic livestock. All of the animals congregate on these bottom lands at this tine. During this period of intra and interspecific competition for forage and water, all might be lost if this habitat type were altered or destroyed. Riparian vegetation supports the highest densities and diversity by far, of songbird and small mammals, and supported nearly a third of the raptor nests. This habitat was also heavily used by migrating songbirds, and roosting raptors. Restoration of riparian habitat after mining may be difficult due to the slow growth of trees and shrubs (compared to grasses) and high sensitivity to damage from livestock grazingo Most of the riparian habitat on the study areas are already 103 beirif? damaged by livestock. Riparian habitat should be excluded from mining until its restoration has been demonstrated and assured. Cliffs and rock formations, especially those made of sandstone, should be protected or enhanced during miningo All cliffs with suitable potholes and ledges for raptor nests should be protected, even if they are not presently being used by nesting raptors. These cliffs are being used as roosting areas for raptors, and may be used in the future for nesting. Cliffs often support a large number of snags which are heavily used as nest trees by mountain bluebirds^ Preserve nests that are regularly used by golden eagles or other special interest raptors. Raptors commonly build several nests within their territories and alter- nate between them from year to year (Call 1978). A nest may be "active" even though it hasn't been used in several years. All known raptor stick nests, even those originally built by red-tailed hawks, should be monitored for use by special interest raptors until mining actually takes place. Raptors often rebuild nests originally built by other species. Minimize disturbance of active raptor nests during the breeding season (April-July). Disturbance during incubation can cause egg mortality from predation or exposure when the incubating bird is flushed, or total abandonment of the nest. Disturbance of nests with nestlings can cause mortality from exposure, interrupted feedings, and premature fledging. 104 Literature Cited American Ornithologist's Union. 1957„ Checklist of North American Birds, 5th ed. American Ornithologist's Union, Ithaca, New York, 691 pp. 1973. 32nd supplement to the American Ornithologist's Union checklist of North American birds. Auk. 90(2 ): 411-419 . 1976. 33rd supplement to the American Ornithologist's Union checklist of North American birds. Auk. 93(4 ): 875-879. Call, M.W. 1978. Nesting habitats and surveying techniques for common western raptors. Tech. Note NOo TN-316. U.S. Bureau of Land Management., Denver Service Center, Denver, Colorado. 115 pp. Dusek, G.L. 1978. Bull Mountains Coal Field Study. Mont. Dept. of Fish and Game and Consolidation Coal Co. Final Report. 120 pp. Flath, D.L. 1979a„ Evaluation of sampling techniques for small mammal community species composition. Paper presented at the 59th annual meeting of the American Society of Mammologists. Corvalis, Oregon. June 1979. . 1979b. Nongame species of special interest or concern. Wildlife Division, Mont. Fish and Game. 73 pp. Hickey, J.J. and S. Mikol. 1979. Estimating breeding bird densities on coal lands in Montana and Wyoming. U.S. Fish and Wildlife Service, Biological Services Program. FWS/WELUT-79/03. March 1979. Kimball, J.W. 1949. The crowing count pheasant census. J„ Wildl, Manage. 13( 1) : 101-120 . Knapp, S.J. 1977. Birney-Decker wildlife study. Mont, Dept » of Fish and Bureau of Land Management final report. 163 pp. 1979. Upland game bird surveys and inventory - Region 7. Proj . W-130-R-10. Job No. II-7. Rept . Mont. Dept, of Fish, Wildlife and Parks. Unpubl. 13 pp. Knowles, C.J. 1975. Range relations of mule deer, elk and cattle in a restoration grazing system during summer and fall. M.S. Thesis, Mont. St. Univ., Bozeman. 79 pp. Martin, P.R. 1980. Sarpy Basin Wildlife Ecology Study. Mont. Dept. of Fish, Wildlife and Parks and Cormorant Corp. Final Report. 125 pp. 105 Odum, E.P. 1959. Fundamentals of ecology. Second edition. Press of W.Bo Saunders Company. Philadelphia. 546 pp. Schwarzkoph, W.F. 1980o Pers. Comm. Biolop;ist. Western Enerf?;y Company. Colstrip„ Swenson, J.E. 1978. Big game survey and inventory (deer) - Region 7, including an analysis of deer production data, 1960-1977o Proj. No„ W-130-R-9, Job No. 1-7. Rept. Mont » Dept. of Fish and Game. Unpubl . 35 pp„ . and S.J. Knapp. 1979. Big game survey and inventory (deer) - Region 7. Proj. No. W-130-R-10, Job No. 1-7. Rept. Mont. Dept. of Fish, Wildlife and ParkSo Unpubl. 32 pp. United States Department of Interior (U.S.D.I.). 1978. Wildlife habitat thermatic map - Birney Quadrangle and Lame Deer Quadrangle. Western Energy and Land Use Team, Fish and Wildlife Service and HRB - Singer Inc., Contract No. 14-16-009-78- 011. .1979o Unsuitability criterion 14 guidelines for the Powder River coal production region. Inst. Memo. No. 80-126. Bureau of Land Management, Washington, D.C. Wallestad, R. 1975. Montana sage grouse. Life history and habitat requirements of sage grouse in Central Montana^ Mont„ Depto of Fish and Game. 66 pp. Wentland, H.J. 1979. Big game surveys and inventory - Region 7. Proj. No. W-130-R-10. Job No. 7. Rept. Mont. Dept, of Fish, Wildlife and Parks. Unpubl. 28 pp. 106 APPENDIX il Appendix Table ]. Parameters recorded at each observation. A. Vofjetation type and subtyy^e 1. Ponderosa pine a. ponderosa pine b . sagebrush c . {TT'assland d. skiiiikbush e. juniper f . snowberry 2. SafTebn;sh/grass]and a. sagebrush b. deciduous shrub c. grassland 3. Creek taotton a. tree b. shrub /grass 4. Agricultural a. ponderosa pine b. sagebrush/grassland c. creek botton B. Activity 1. Standinf!^ 2. Running 3. Lying 1 . Feeding C. Topr^?n"aphy 1. Mesa-butte top 2. Hesa-butte steep sides 3. Dissected mid-slopes 4. Alluvium/terrace 5. QuTent flood plain 1). Expos-ure 1. North 2. South 3. East 4. West 5. Northeast 6. Northwest 7. Southeast 8. Southwest 9. Flat Slope" 1. Flat 2, Gentle 3. Medium 4, Steep 107 Appendix Figure 1. Ponderosa pine vegetation subtype, Appendix Figure 2. Ponderosa pine-sagebrush vegetation subtype. 108 Appendix Figure 3. Ponderosa pine-grassland vegetation subtype. t Appendix rigure 4. Ponaorosa pine-skunkbusli vegetation subtype. 109 Appendix Figure 5. Ponderosa pine-juniper vegetation subtype. « Appendix Figure 6. Ponderosa pine-snowberry vegetation subtype. 110 Appendix Figure 7. Sagetarush/grassland-big sagebrush vegetation subtype. Appendix Figure o. Sagebrush/grassland-silver sagebrush vegetation subtype. Ill Appendix Figure 9. Sagebrush/grassland-grassland vegetation subtype. Appendix Figure 10. Sagebrush/grassland-deciduous shrub vegetation subtype. 112 f- Appendix Figure 11. Creekbotton-deciduous tree vegetation subtype. Appendix Figure 12, Creekbottom-deciduous shrub vegetation subtype. 113 Appendix Figure 13. Agricultural-grassland vegetation subtype. Appendix Figure 14. Agricultural-creekbottom vegetation subtype. 114