HARVARD UNIVERSITY LEE LIBRARY OF THE Museum of Comparative Zoology DAI Ti PI] } N OTN Durst OA | sea \ Kran à Ù m \ N È k Ù Ci HUREN) TROT RARE NA | ii N fi i i US na RS ‘ n ti } RS no Ù | MOLE NN ny Ni y iy | ; { : $ if x WELL + Mi t { i fr ‘ Ni IT 3 Ti \ i \ o 2 sa D 2 N x x Pa 2 7 (I { * 3 en i Bl: i i ly i en iv yy ln VII a : ; VAGA Pi ¢ 1 x { 1 >) IDEE TARE AFLEVERING 7 * 1968 TIJDSCHRIFT VOOR ENTOMOLOGIE UITGEGEVEN DOOR DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING Afleveringen 1—3 verschenen 21 juni 1968 Da 4-5 5 10 oktober Pe 53 6-7 ds 27 december „, INHOUD VAN DEEL 111 CHRYSANTHUS, Fr., O.F.M. Cap. — Spiders from South New Guinea X. . . 49 Hire Ris LAMBERS, D. — A study of Newguenaphis Blanchard, 1939, with descriptions of new species (Homoptera, Aphididae) 2... ru 7257 LIEFTINCK, M. A. — A review of the genus Oligoaeschna Selys in Southeast Asia 137 LitH, J. P. van. — Contribution to the knowledge of Indo-Australian, South Pacific and East Asiatic Psenini. Genus Psen Latreille (Hymenoptera, Sphe- CIAO) neue ee ee So! ARE A SO, OBRAZTSOV +, NIKOLAUS S. — Die Gattungen der Palaearktischen Tortricidae. II. Die Unterfamilie Olethreutinae. 8. Teil und Schluss. Mit einem Gesamt- \ register zu allen Teilen, zusammengestellt von B. J. Lempke . . . . . 1 VECHT, J. VAN DER and F. M. A. van BREUGEL. — Revision of the nominate subgenus Sceliphron Latreille (Hymenoptera, con Ca on the Sceliphtonini spatte) RE Va D TAF ONTO Register APE AL TE TE ED I Ne 877 Eu Sano aalw As (eee FE k= pit FAN " M Ù ee. Y i KR k, © fr BN er ive x EK i di PA, STI eee ie 7 at eo a ' rich * SH. Ad Katy) a) \ Cubes BIS pe ware sf ” FR ibid A she È. mx allin dii alito Ul | yess salt gh won OB DIS abbé ban tat Tu a bui TA m Wise CN € = an me. RARE ECT er DEEL 111 AFLEVERING 1 1968 TIJDSCHRIFT VOOR ENTOMOLOGIE UITGEGEVEN DOOR DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING MUS. COMP. ZOOL, LIBRARY JUL 25 1968 HARVARD UNIVERSITY INHOUD : NIKOLAUS S. OBRAZTSOV +. — Die Gattungen der Palaearktischen Tortricidae. II. Die Unterfamilie Olethreutinae. 8. Teil und Schluss. Mit einem Gesamtregister zu allen Teilen, zusammengestellt von B. J. LEMPKE, p. 1—48, Abb. 1—5 und Tafeln 1—11. Tijdschrift voor Entomologie, deel 111, afl. 1 Gepubliceerd 21-VI-1968 NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING BESTUUR (BOARD) Voorzitter (Chairman) . . ey Gs Barendrecht Vice-Voorzitter (ice ze ee Wiebes Secretaris (Secretary) . . . . . . . . W. Hellinga PAGA essi un . … Weesperzijde 23 II, Amsterdam-O. Penningmeester (neh), e RAW]. abos Address u . . . + Van Baerlestraat 261, Amsterdam Bibliothecaris Tubes) Ro 00, CAEN Jeekel Addresse N 8 Zeeburgerdjk 21, Amsterdam.O: Kedene (Members)... em. JA S anse ASSEN aBesemer AFDELING VOOR TOEGEPASTE ENTOMOLOGIE (DIVISION OF APPLIED ENTOMOLOGY) BESTUUR (BOARD) Woorzitter (Chairman) seven VARE SH Besemer Secretaris (Secretary) er. an dee „Pe Gruys Address un neun en Nedenemdsesiraab 35, Restereni Eeden. (Members) … . . . o... 0.0. (IL. Bravenboer, J.J. Laarman, J.” Bo Manan Diether Publicaties van de Vereeniging (Publications of the Society) Subscription may be obtained from all booksellers or directly from the Librarian, Zeeburgerdijk 21, Amsterdam-O., except for Entomologia Experimentalis et Applicata, which is available through booksellers or from the Noord-Holland Editing Co., Post Office Box 103, Amsterdam. TIJDSCHRIFT VOOR ENTOMOLOGIE Redactie (Editing Board) . . . . . . Pater Chrysanthus, A. Diakonoff, C. A. W. Jeckel, M. A. Lieftinck, J. T. Wiebes Address . . . 4 2 2 ++ + + + Rijksmuseum van Natuurlijke Historie, Raamsteeg 2, Leiden. The Journal serves the publication of papers on Insecta, Myriapoda and Arachnoidea. It appears in separate issues, forming an annual volume of 350—400 pages. Subscription rate: D.Fl. 45.— (£ BNO $ 12.50) per volume. MONOGRAFIEEN VAN DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING Redactie (Editing Board) and address as for Tijdschrift voor Entomologie. The Monographs are intended for the pup ication of larger entomological papers on a single subject and will appear irregularly. - The following Monographs have been published: Hors série: F. T. Valck Lucassen et al., 1961. — Monographie du genre Lomaptera Gory & Percheron (Coleoptera, Cetoniidae), 299 pages, 739 figs., 2 pl, map. D.Fl. 50— (£ 5.— —, $ 13.90). No. 1. A. J. Besseling, 1964. — De Nederlandse Watermijten (Hydrachnellae Latreille, 1802) (The Hydrachnellae of the Netherlands), 199 pp. 333 figs., D.Fl.25— (£ 2.10.—, $ 6.95). DIE GATTUNGEN DER PALAEARKTISCHEN TORTRICIDAE II. DIE UNTERFAMILIE OLETHREUTINAE 8. Teil und Schluss *) VON NIKOLAUS S. OBRAZTSOV + **) (Mit Abbildungen 1—5 und Tafeln 1—11) mit einem Gesamtregister zu allen Teilen zusammengestellt von B. J. LEMPKE Amsterdam Abstract The present and last instalment of the second volume of the Revision contains a survey of the genus Ewcosma Hübner, 1823, divided into four subgenera: Ewcosma Hübner, 1823, Pygolopha Lederer, 1859, Palpocrinia Kennel, 1919 and Phaneta Stephens, 1852. Of these subgenera 48, 1, 1, and 69 species, respectively, are enumerated. A commentary dealing with 10 species, is added. A general alphabetical index to all parts of the Revision, starting with vol. 97, part 3 (1954) of this journal is attached, compiled by B. J. Lempke (p. 22), together with a list of posthumous papers of Dr. Obraztsov (p. 48). 88. Gattung Eucosma Hb., 1823 Typus generis (monotyp.): Eucosma circulana Hb., 1823 Phalaena Tortrix (part.): Linné, 1761, Fauna Suec.: 349. ?Phalaena (part.): Scopoli, 1763, Ent. Carn.: 234. ?Pyralis (part.): Fabricius, 1798, Suppl. Ent. Syst.: 476. Tortrix (part.): Haworth, 1811, Lep. Brit.: 455. ?Tinea (part.): Charpentier, 1821, Zinsler etc.: 120. Olethreutes (part.): Hübner, 1822, Syst.-alph. Verz.: 58. Eucosma Hübner, 1823, Zutr. Samml. Exot. Schm. 2: 28. Cydia (part.): Hübner, 1825, Verz. bek. Schm.: 375. Epiblema (part.): Hübner, 1825, op. cit.: 375. Ernarmonia (part.): Hübner, 1825, op. cit.: 375. Anchylopera (part.): Stephens, 1829, Syst. Cat. Brit. Ins. 2: 178. ?Aphelia (part.): Stephens, 1829, op. cit.: 180. Semasia (part.): Stephens, 1829, op. cit.: 180. Eupoecilia (part.): Stephens, 1829, op. cit.: 191. Grapholita (part.): Treitschke, 1829, Schm. Eur. 7: 232. Penthina (part.): Treitschke, 1830, Schm. Eur. 8: 26. *) Dieser Teil der Revision wurde mit Unterstützung des U.S. National Science Foundation zur Publikation vorbereitet. **) Im Leben: Research Associate, American Museum of Natural History, New York, U.S.A. Für eine Liste von posthumer Publikationen Dr. Obraztsov’s siehe Seite 48. 1 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (228) Gattung Eucosma Hb. Abb. 1, E. (E.) circulana Hb., Männchen, Kopf; Abb. 2, Geäder; Abb. 3—4, weibliche Genitalien; Abb. 5, männliche Genitalien. (229) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 3 Grapholitha (part.): Treitschke, 1830, op. cit.: 204. Cochylis (part.): Treitschke, 1830, op. cit.: 277. Carpocapsa (non Tr.): Curtis, 1831, Brit. Ent.: expl. t. 352. Anticlea (part.): Stephens, 1834, Illustr. Brit. Ent., Haust. 4: 114. Carpocapsa (Epiblema): Stephens, 1834, op. cit.: 122. ?Bactra (part.): Stephens, 1834, op. cit.: 125. Phoxopteris (non Tr.): Zetterstedt, 1840, Ins. Lap.: 987. Teras (non Tr.): Eversmann, 1840, Fauna Lepid. Volgo-Ural.: 520. Paedisca (part.): Freyer, 1842, Neue Beitr. Schm.kunde 4: 48. ?Peronea (part.): Duponchel, 1845, Cat. Méth. Lép. Eur.: 291. Antithesia (part.): Duponchel, 1845, op. cit.: 296. Catoptria Guenée, 1845, Ann. Soc. Ent. France (2) 3: 187. Typus generis (selectus): Tortrix cana Hw., 1811 (= Catoptria carduana Gn., 1845) (Fernald, 1908, Gen. Tortr.: 33). Nom. praeocc. per Catoptria Hb., 1825, Crambidae. Philalcea (part.): Westwood & Humphreys, 1845, Brit. Moths 2: 132. ?Ablabia (part.): Westwood & Humphreys, 1845, op. cit.: 140. Phaneta Stephens, 1852, List. Specim. Brit. Anim. 10: 32. Typus generis (monotyp.): Cochylis pauperana Dup., 1843. Lithographia (Notocelia) (part.): Stephens, 1852, op. cit.: 34. Grapholita : (Cydia) (part.): Stephens, 1852,, op. cit.: 93. Pygolopha Lederer, 1859, Wien. Ent. Mschr. 3: 123, 279. Typus generis (monotyp.): Penthina lugubrana Tr., 1830 (= Pygolopha tinacriana Ld., 1859). Grapholitha (Paedisca) (part.): Lederer, 1859, ibid.: 332. Grapholitha (Coccyx) (part.): Lederer, 1859, ibid.: 278. Calosetia Stainton, 1859, Man. Brit. Butt. & Moths 2: 271. Typus generis (monotyp.): Tortrix nigromaculana Hw., 1811. Ioplocama Clemens, 1860, Proc. Acad. Nat. Sci. Philad.: 360. Typus generis (monotyp.): Ioplocama formosana Clem., 1860. Poecilochroma (non Stph.): Clemens, 1860, ibid.: 353. Affa Walker, 1863, List Specim. Lepidopt. Ins. 27: 202. Typus generis (monotyp.): Affa bipunc- tella Wkr., 1863. Grapholitha (Semasia) (part.): Heinemann, 1863, Schm. Dtschl. u. Schweiz (2) 1 (1): 166. Steganoptycha (non Stph.): Clemens, 1865, Proc. Ent. Soc. Philad. 5: 137. Conchylis (non Sod.): Robinson, 1869, Trans. Amer. Ent. Soc. 2: 284. Exentera Grote, 1877, Canad. Ent. 9: 227. Typus generis (monotyp.): Exentera apriliana Grote, 1877. Rbyacionia (part.): Walsingham, 1879, Illustr. Lep. Het. B.M. 4: 75. Exenterella Grote, 1883, Canad. Ent. 15: 23. Typus generis (heredit.): Exentera apriliana Grote, 1877. Nom. substit. pro Exentera Grote, 1877. Crambus (non F.): Hulst, 1886, Trans. Amer. Ent. Soc. 33: 166. Thiodia (part.): Walsingham, 1897, Proc. Zool. Soc. Lond.: 125. Phtheochroa (ex. err.): Aigner-Abafi, 1898, Illustr. Zschr. Ent. 3: 312. Epinotia (part.): Fernald, 1903, Bull. U.S. Nat. Mus. 52 (1902): 464. Proteopteryx (non Wlsm.): Kearfott, 1907, Canad. Ent. 39: 155. Enarmonia (part.): Barnes & McDunnough, 1917, Check List Lepid. Boreal Amer.: 173. Palpocrinia Kennel, 1919, Mitt. Miinchn. Ent. Ges. 8: 66. Typus generis (monotyp.): Palpocrinia ottoniana Kenn., 1919. synon. nov. Ancylis (non Hb.): Meyrick, 1922, Exot. Micr. 2: 514. Phaneta (Semasia): Benander, 1928, Ent. Tidskr. 29: 142. Catoptria (Phaneta): Obraztsov, 1946, Zschr. Wien. Ent. Ges. 30: 39. Eucosmoides Obraztsov, 1946, ibid.: 38. Typus generis (monotyp. design.): Paedisca decolorana Frr., 1842. Catoptria (Semasia): Jäckh, 1953, Zschr. Lepid. 3: 43. Eucosma (Phaneta): Hannemann, 1961, Dahls Tierw. Dtschl. 48, Kleinschm. 1: 123. Eucosma (Catoptria): Kouznetzov, 1962, Bull. Soc. Ent. Mulhouse: 55. 4 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (230) Der Gattung Epiblema bis auf Folgendes ahnlich. Thorax glatt. Vorderflügel (Abb. 2) meistens nicht breit, distal ganz unwesentlich erweitert; Termen mäßig schräg, gerade oder zwischen den Adern M, und Cu, leicht eingezogen; Costalumschlag beim Männchen nicht stets vorhanden. Die Vorderflügelader S mehr oder weniger gerade, meistens ziemlich kurz; R, entspringt an oder kurz vor der Mitte der Mittelzelle; R, viel näher zu R, als zu R,; Innenader der Mittelzelle entspringt zwischen R, und R, oder fehlt; Adern R; bis Mz etwa gleich weit auseinander gestellt; Cu, der Mg stark genähert, verläuft dieser parallel, oder ist in der Mitte mehr oder weniger von dieser entfernt; Cu, entspringt kurz vor dem letzten Drittel der Mittelzelle; Basalgabel A, + 3 bis ein Drittel so lang wie die ganze Ader. Hinterflügel (Abb. 2) mit Costa im zweiten basalen Drittel leicht aufgebogen, dann etwas absteigend und in ihrem ganzen Verlauf gerade, oder fast gerade; Termen sanft konvex oder ziemlich flach. Die Ader S ganz sanft eingebogen; M; und Cu, gestielt oder zusammenfallend; Cu, entspringt kurz vor dem letzten Viertel der Mittelzelle. Männliche Genitalien (Abb. 5). Valva ohne Pulvinus; Aedoeagus in der Regel ziemlich plump, dick und verhältnismäßig kurz. Weibliche Genitalien (Abb. 3—4). Apophyses posteriores kürzer als die Apophyses anteriores; Ostium bursae liegt in einem mehr oder weniger tiefen Ausschnitt des Caudalrandes der Ventralplatte; Lamella post- vaginalis länglich, aus diesem Ausschnitt mehr oder weniger weit caudal hervortretend; kein Antrum vorhanden; Signa manchmal zu einem Stück reduziert oder fehlen. Encosma ist eine sehr artenreiche Gattung, die größte unter den Olethreutinae, und sie hat ihre Vertreter in allen faunistischen Gebieten. Trotz ihres großen Umfanges ist diese Gattung in ihren morphologischen Merkmalen doch ziemlich einheitlich und diese variieren nur unbedeutend von Art zu Art oder individuell. Sogar die übliche Aufteilung der Ewcosma-Arten in drei Gattungen (Ewcosma, Pygolopha und Phaneta = Thiodia auct., non Hb.) erwies sich nicht als gerechtfertigt und diese Gruppen sind nur mehr von subgenerischem Werte. Heinrich (1923) vermutete die Möglichkeit einer weiteren Aufteilung der Ewcosma in kleinere systematische Gruppen, hauptsächlich auf Grund der Cucullus-Form, aber bis jetzt gelang es nur Pelochrista Ld. als eine besondere Gattung abzutrennen. Die Artunterschiede der äußeren morphologischen Merkmale sind in Ezcosma meistens ganz unbedeutend und öfters zeigen sie Übergänge, die durch die infraspezifische Varia- bilität stark verdunkelt sind, wie z.B. der Abstand zwischen den Vorderflügeladern R, bis Cu, oder die Ursprungsstelle der Innenader der Mittelzelle, die bald R,, bald Ro näher steht und bisweilen fehlt die Ader sogar. Die von Kennel (1908, 1916) für lugu- brana angegebene starke Ausbiegung des mittleren Teils der Vorderflügelader Cu,, ihre Verkürzung und Krümmung beim Männchen in der Richtung zu M; ohne den Saum zu erreichen, erwies sich von rein individueller Natur und tritt, obwohl seltener, auch in den anderen Ezcosma-Arten auf. Die Hinterflügeladern R und M, können bei ein und der- selben Art entweder dicht beisammen entspringen oder gestielt sein. Der gemeinsame Stiel der Hinterflügeladern M; und Cu, ist manchmal an den beiden Flügeln ein und desselben Falters ungleich lang und kann zu einer vollständigen Verschmelzung dieser Adern führen. Ausnahmsweise kann diese Verschmelzung nur auf einem Hinterflügel geschehen. Als ein atavistisches Merkmal wurden bei einem /acteana-Minnchen im A.M.N.H. die Hinterflügeladern M, und Cu, an der Basis als deutlich getrennt, in der Mitte verschmolzen und gegen den Saum wieder getrennt beobachtet. Durchschnittlich sind die äußeren Merkmale artlich ziemlich beständig, aber bei mehreren, nicht unbedingt (231) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 5 nahe verwandten Arten ganz gleich und deshalb für diagnostische Zwecke meistens wenig geeignet. Die Genitalunterschiede sind bei den Ewcosma-Arten nicht immer auffallend, aber zusammen mit der Vorderflügelzeichnung diagnostisch sehr wichtig. Larvalmorphologisch sind die Ezcosma-Arten noch ganz ungenügend erforscht (Swatschek, 1958). Trotzdem kònnen ihre Raupen von denen der Pelochrista meistens dadurch unterschieden werden, daß bei Ewcosma die Borstengruppe III am 8. Abdomi- nalsegment ventrocranial vom Stigma oder mit diesem in gleicher Hôhe steht. Die Borsten II sind bei Ewcosma an demselben Segment weiter auseinander gestellt als die Borsten I. Die Unterschiede den Epiblema-Raupen gegenüber sind bei der Besprechung dieser letzteren Gattung näher eròrtert worden. Die von MacKay (1959) verôffentlichten Ergebnisse über die wenigen von ihr untersuchten nordamerikanischen Ezcosma-Raupen, die sie zwischen Ewcosma und “Thiodia” aufteilte, zeigen, daß eine gemeinsame larval- morphologische Charakteristik der ganzen Gattung vorläufig noch sehr kompliziert ist. Sie teilte die “Encosma”-Arten in drei Gruppen und die “T’hiodia"-Arten sogar in vier Gruppen, die, nach der Ansicht dieser Verfasserin, eine geringe Verwandtschaft zeigen. Von diesen erwies sich die 1. “Ewcosma'’-Gruppe, deren Raupen auf den Coniferen leben, von den ibrigen zweien besonders unterschieden. Die Raupen der letzteren fressen an verschiedenen Kräutern, an welchen sie in den Wurzeln und Stengeln bohren. In der 1. Gruppe ist die Spindelspitze abgerundet, wahrend sie in zwei anderen Gruppen deut- lich zugespitzt ist. Außerdem sind die Raupen der 1. Gruppe durch das Vorhandensein einer zusatzlichen Borste (IVa) auf dem 1. bis 7. Abdominalsegment charakterisiert. Die Entdeckung dieser Borste ist besonders interessant, da Swatschek (1958) sie bei den europäischen Arten nur in den Gattungen Rhyacionia und Clavigesta gefunden hat. Was den taxonomischen Wert der von MacKay aufgestellten “Ezcosma”-Gruppen betrifft, so ist die 1. Gruppe auch imaginalmorphologisch begründet und umfafit genitaliter ziem- lich einheitlich gebaute Arten, die trotzdem generisch nicht zu trennen sind. Dagegen erscheint die 2. Gruppe ganz bunt und enthält auch manche Pelochrista-Arten. Die Arten der 3. Gruppe sind von der 2. Gruppe imaginalmorphologisch nicht abtrennbar. Die Aufteilung der “Thiodia’-Arten in vier Raupengruppen findet auch wenig Unterstützung in der Imaginalmorphologie. Wie oben gesagt, können die Ewcosma-Arten nach ihren imaginalen Merkmalen in drei Untergattungen aufgeteilt werden. Dazu kommt noch Palpocrinia als die vierte Untergattung. Diese letztere wurde zunächst als eine eigene, aus einer einzigen Art bestehende und nur in einem männlichen Exemplare bekannte Gattung aufgestellt, unter den Tortricinae eingereiht (Kennel, 1919), später (Obraztsov, 1955) unter den Cnephasiini untergebracht und endlich (Obraztsov, 1965) richtig für eine Eucosmine anerkannt. Die vier Evcosma-Untergattungen sind wie folgt charakterisiert: 1. Untergattung Ezcosma Hb. s. str. (Typus subgeneris: Encosma circulana Hb., 1823; eine nearktische Art). Vorderflügel des Männchens mit einem Costalumschlag. Hinter- leibspitze des Weibchens einfach. 2. Untergattung Pygolopha Ld. (Typus subgeneris: Penthina lugubrana Tr., 1830). Vorderflügel wie in der vorigen Untergattung. Hinterleibspitze des Weibchens mit einem rundlichen Schuppenschopf. 3. Untergattung Palpocrinia Kenn., status nov. (Typus subgeneris: Palpocrinia otto- niana Kenn., 1919). Vorderflügel des Männchens ohne Costalumschlag, aber mit einem Haarbüschel an der Costa-Basis. Kopf, Labialpalpen, Tegulae und Unterseite des Thorax sehr lang behaart (Obraztsov, 1955: 163, fig. 279; 1956: 117; 1965: 25). 6 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (232) 4. Untergattung Phaneta Stph. (Typus subgeneris: Cochylis pauperana Dup., 1843). Vorderflügel in beiden Geschlechtern ohne besondere Auszeichnungen. Beschuppung des Kopfes, Körpers und ihrer Teile normal. KATALOG DER PALAEARKTISCHEN EUCOSMA-ARTEN Sg. Eucosma Hb., 1823 E. (E.) sordicomana (Stgr.) comb. nova* sordicomana Staudinger, 1859, Stett. Ent. Ztg. 20: 232 (Grapholitha). — Staudinger & Rebel, 1901: 117, No. 2096; Kennel, 1921: 571, t. 21 fig. 53 (2); diese Arbeit: Taf. 1 Fig. 1 (4- Genitalien). — Iberische Halbinsel. E. (E.) obesana (Kenn.) comb. nova obesana Kennel, 1901, Iris 13 (1900): 284 (Epiblema). — Staudinger & Rebel, 1901: 263, No. 2102bis; Kennel, 1921: 571, t. 21 fig. 54 (2). — Andalusien. E. (E.) medullana (Stgr.) comb. nova* medullana Staudinger, 1879, Horae Soc. Ent. Ross. 15: 254 (Grapholitha). — Staudinger & Rebel, 1901: 115, No. 2071; Kennel, 1921: 550, t. 21 fig. 7 (4); diese Arbeit: Taf. 1 Fig. 2 (4 -Genitalien). — Bulgarien; Mazedonien; Kleinasien; Syrien. E. (E.) conformana (Mn.) comb. nova* (1) conformana Mann, 1872, Verh. zool.-bot. Ges. Wien 22: 36 (Grapholitha); significantana Kennel, 1901, Iris 13 (1900): 282 (Epiblema). — Staudinger & Rebel, 1901: 121, No. 2175; 263, No. 2083bis (als significantana); Kennel, 1921: 567, t. 21 fig. 43, 44 (8 2); Hanne- mann, 1961: 126 nota, 220 fig. (4-Genitalien), t. 21 fig. 23 (Falter); diese Arbeit: Taf. 1 Fig. 3, Taf. 2 Fig. 1 (& 2 -Genitalien). — Ungarn; Kroatien; Dalmatien; Kärnten; Mazedonien; Griechenland; Kleinasien; Syrien; Palästina. E. (E.) clarescens Kuzn.* clarescens Kuznetzov, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 264, fig. 6 (4 2 -Geni- talien) (Ewcosma). — OstruBland (Provinz Uralsk). E. (E.) jerusalemana (Ams.) comb. nova* jerusalemana Amsel, [1933, Zoogeogr. 2: 123; nom. nud.}, 1935, Mitt. Zool. Mus. Berlin 20: 291, t. 11 fig. 87 (Falter), t. 17 fig. 62 (4-Genitalien) (Epiblema); diese Arbeit: Taf. 2 Fig. 3. — Palästina; Iran. E. (E.) persiae Raz. persiae Razowski, 1963, Acta Zool. Cracov. 8: 256, fig. 5—8 (Kopf, Vorderfligel, 4 2- Genitalien) (Eucosma). — Südwestiran. E. (E.) gypsatana (Kenn.) comb. nova* (2) gypsatana Kennel, 1921, Pal. Tortr.: 557, t. 21 fig. 25 (Epiblema). — Diese Arbeit: Taf. 1 Fig. 4 (4-Genitalien). — Kleinasien. E. (E.) albuneana (Z.) comb. nova albuneana Zeller, 1847, Isis: 725 (Paedisca); albuncana (err. typogr.): Lederer, 1859, Wien. Ent. Mschr. 3: 332 (Grapholitha, Paedisca). — Staudinger & Rebel, 1901: 116, No. 2079; Kennel, 1921: 557, t. 21 fig. 24 (4); diese Arbeit: Taf. 3 Fig. 1 (@-Genitalien). — Süd- frankreich; Spanien; Sardinien; Sizilien; Dalmatien; Istrien; Rumänien; Südrußland; Nordwest- afrika. E. (E.) albarracina Hartig (3) albarracina Hartig, 1941, Mitt. Münchn. Ent. Ges. 31: 156, t. 6 fig. 5 (Ezcosma). — Spanien (Albarracin). (233) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 7 E. (E.) lacteana (Tr.) lacteana Treitschke, 1835, Schm. Eur. 10 (3): 113, 256 (Grapbolitha); lacteanum: Meyer, 1901, Kranchers Ent. Jahrb. 18: 148 (Epiblema). — Staudinger & Rebel, 1901: 116, No. 2081; Kennel, 1921: 558, t. 21 fig. 27 (4); Hannemann, 1961: 132, fig. 260 ( Â-Genitalien), t. 13 fig. 20 (Falter). — Belgien; Frankreich; Mitteleuropa; Ungarn; Polen; Spanien. E. (E.) agnatana (Chr.) comb. nova agnatana Christoph, 1872, Horae Soc. Ent. Ross. 9: 15, t. fig. 12 (Grapholitha). — Staudinger & Rebel, 1901: 116, No. 2083; Kennel, 1916: 559, t. 21 fig. 29, 30 (3). — Ostrußland; Südukraine; Kleinasien. E. (E.) albidulana (H.S.)* albidulana Herrich-Schäffer, 1851, Syst. Bearb. Schm. Eur. 4: 245 (Semasia) [1848, Tortr.: t. 44 fig. 310; non bin.}; albidulata (laps.): Drenowsky, 1925, Zschr. wiss. Ins.biol. 20: 4 (Epiblema). — Staudinger & Rebel, 1901: 116, No. 2082; Kennel, 1921: 559, t. 21 fig. 28 (2); Toll, 1939: 247, t. 2 fig. 10 (&-Genitalien); Hannemann, 1961: 125, fig. 241 (4- Genitalien), t. 13 fig. 25 (Falter). — Deutschland; Österreich; Ungarn; Balkanhalbinsel; Süd- und Westfrankreich; Polen; Iran; Algerien; ?Ostchina. E. (E.) rigidana (Snell.)* (4) rigidana Snellen, 1883, Tijdschr. v. Ent. 26: 203, t. 12 fig. 4, 4a (Falter, Kopf) (Grapbolitha, Semasia). — Staudinger & Rebel, 1901: 116, No. 2089; Kennel, 1916: 527, t. 20 fig. 29 (9); Issiki, 1957: 59, t. 9 fig. 263 (2); diese Arbeit: Taf. 1 Fig. 5 (4-Genitalien). — Südost- sibirien; Korea; Japan. E. (E.) culmana (M.R.) comb. nova culmana Müller-Rutz, 1932, Mitt. Schweiz. Ent. Ges. 15: 198, Tekstabb. (4 9) (Epiblema). — Schweiz. E. (E.) coagulana (Kenn.)* coagulana Kennel, 1901, Iris 13 (1900): 278 (Epiblema). — Staudinger & Rebel, 1901: 263, No. 2085bis; Kennel, 1921: 560, t. 21 fig. 31, 32 (4 9); Razowski, 1963: 256, fig. 4 (&- Genitalien). — Transkaukasien; Taurus; Iran; Zentralasien. E. (E.) monstratana (Rbl.)* (5) monstratana Rebel, 1906, Iris 19: 235 (Epiblema). — Thomann, 1926: 164, t. 2 fig. 3 (Falter); Hannemann, 1961: 126 nota, 220 fig. (4-Genitalien), t. 22 fig. 1 (Falter); diese Arbeit: Taf. 2 Fig. 2, Taf. 3 Fig. 5, 6, Taf. 4 Fig. 1 (Falter; 4 9-Genitalien). — Schweiz; Vorarlberg; Südkarpaten. E. (E.) scutiformis Meyr.* scutiformis Meyrick, 1931, Bull. Sect. Sci. Acad. Roum. 14: 64 (Eucosma). — Clarke, 1958: 387, t. 192 fig. 3, 3a (Falter; ¢-Genitalien); diese Arbeit: Taf. 3 Fig. 2—4 ( 2 -Genitalien). — Ostchina. E. (E.) kemnerana (Lewin) comb. nova kemnerana Lewin, 1942, Opusc. Ent. 7: 70, fig. 3 (4-Genitalien) (Epiblema). — Schweden. E. (E.) cana (Hw.)* cana Haworth, 1811, Lep. Brit.: 456 (Tortrix); monetulana Hübner, [1814—1817, Samml. eur. Schm., Tortr.: t. 41 fig. 257; non bin.]; 1822, Syst.-alph. Verz.: 62 (Olethreutes); hohen- wartiana (non Schiff.): Treitschke, 1830, Schm. Eur. 8: 204 (Grapholitha); carduana Guenée, 1845, Ann. Soc. Ent. France (2) 3: 188 (Catoptria); marmorana [| ?Fabricius, 1798, Suppl. Ent. Syst.: 477 (Pyralis) ]: Werneburg, 1864, Beitr. Schm.kund. 1: 561 (Tortrix); scopoliana (part.): Stephens, 1852, List Spec. Brit. Anim. 10: 62 (Grapholita); hohenwarthana var. a.: Stephens, 1852, op. cit.: 62 (Grapholita); aspidiscana var. ?b: Stephens, 1852, op. cit.: 93 (Grapholita, Cydia). — Staudinger & Rebel, 1901: 116, No. 2086; Kennel, 1921: 562, t. 21 fig. 36 (9); 8 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (234) Pierce & Metcalfe, 1922: 72, t. 24 (4 9-Genitalien); Benander, 1950: 128, fig. 119 ( &-Geni- talien), t. 7 fig. 36 (Vorderflügel); Issiki, 1957: 59, t. 9 fig. 264 (Falter); Swatschek, 1958: 137, fig. 147, 149, 150 (Larvalmorphologie); Okano, 1959: 260, t. 174 fig. 24 (Falter); Hannemann, 1961: fig. 2392 (4-Genitalien; als hohenwartiana); Schantz, 1962: 2, fig. 1, 7 (2 8-Genitalien). — Großbritannien; Europa; Armenien; Kleinasien; Zentralasien; China; Japan; Nordwestafrika. f. (?) subvittana Stgr. subvittana Staudinger, 1922, Iris 5: 299 (Grapholita). — Staudinger & Rebel, 1901: 116, No. 20864; Kennel, 1921: 563. — Spanien; Sardinien; Nordwestafrika. E. (E) albicosta Falk.* albicosta Falkovitsh, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 279, fig. 14, 15 (6 9- Genitalien) (Ewcosma). — Transili-Alatau; Alai-Gebirge. E. (E.) fulvana (Stph.)* fulvana Stephens, [1829, Syst. Cat. Brit. Ins. 2: 180; nom. nud.}, 1834, Illustr. Brit. Ent, Haust. 4: 123 (Carpocapsa, Epiblema); pupillana (non Cl:): Haworth, 1811, Lep. Brit.: 455 (Tortrix); ?hohenwartiana (non Schiff.): Haworth, 1811, op. cit.: 455 (Tortrix); ?strigana Curtis, 1831, Brit. Ent.: expl. t. 352 (Carpocapsa); hohenwarthiana: Stephens, 1852, List Spec. Brit. Anim. 10: 62 (Grapbolita); jaceana (part.): Wocke, 1871, Stgr.-Wck. Cat. Lep. eur. Faun.: 252, No. 10464 (Grapholitha, Paedisca); fulvanum: Meyer, 1909, Kranchers Ent. Jahrb. 18: 147 (Epiblema); scopoliana (non Hw.): Benander, 1950, Svensk Ins.fauna 10: 128, fig. 119 (&-Genitalien) (Epiblema). — Staudinger & Rebel, 1901: 116, No. 2084; Kennel, 1921: 563, t. 21 fig. 35 (4); Pierce & Metcalfe, 1922: 72, t. 24 (4 9-Genitalien); Swatschek, 1958: 137 (Larvalmorphologie); Hannemann, 1961: fig. 240 (&-Genitalien; als jaceana); Schantz, 1962: 3, fig. 2, 10 (4 2-Genitalien); Kouznetzov, 1962, Bull. Soc. ent. Mulhouse: 55, fig. 5 (2-Genitalien). — Großbritannien; ganz Europa; Südwest- und Zentralasien; Nordchina. E. (E.) jaceana (H.S.)* jaceana Herrich-Schäffer, 1851, Syst. Bearb. Schm. Eur. 4: 248 (Semasia); fulvana (patt.): Barrett, 1873, Ent. Mo. Mag. 10: 5. — Staudinger & Rebel, 1901: 116, No. 2084 (part.; als fulvana); Toll, 1939: 247, 259, t. 2 fig. 11 (&-Genitalien). Mitteldeutschland; Sardinien; Siidpolen; Ostbaltikum. E. (E.) aurantiradix Kouzn.* aurantiradix Kouznetzov, 1962, Bull. Soc. Ent. Mulhouse: 55, fig. 4 (4 2 -Genitalien) (Ezcos- ma, Catoptria). — Südostsibirien (Provinz Amur). E. (E.) haberhaueri (Kenn.) comb. nova haberhaueri Kennel, 1901, Iris 13 (1900): 280 (Epiblema). — Staudinger & Rebel, 1901: 263, No. 2085zer; Kennel, 1921: 564, t. 21 fig. 37 (4). — Transili-Alatau; Alai-Gebirge; Kuldscha; Ferghana. E. (E.) hohenwartiana (Schiff.)* hohenwartiana Schiffermiller & Denis, 1776, Syst. Verz. Schm. Wien. Geg.: 129 (Phalaena Tortrix); scopoliana (non Schiff.): Haworth, 1811, Lep. Brit.: 456 (Tortrix); pupillana (non Cl.): Hübner, [1796—1799, Samml. eur. Schm., Tortr. t. 4 fig. 20; non bin.}, 1822, Syst.-alph. Verz.: 63 (Olethreutes); hohenwarthiana: Treitschke, 1829, Schm. Eur. 7: 232 (Grapholita); ?strigana: Westwood & Humphreys, 1845, Brit. Moth 2: 138 (Carpocapsa); ?cervana: Scopoli, 1763, Ent. Carn.: 234 (Phalaena); Zeller, 1855, Stett. Ent. Ztg. 16: 246 (Tortrix); ?hybridella (part.): Lederer, 1859, Wien. Ent. Mschr. 3: 278 (Coccyx); hohenworthiana (lapsus): Black- more, 1864, Ent. Mo. Mag. 1: 123; jaceana (part.): Wocke, 1871, Stgr.-Wck., Cat. Lep. eur. Fauna: 252, No. 1046a (Grapholitha, Paedisca); carduana (part.): Wocke, 1871, op. cit.: 252, No. 1047 (Grapholitha, Paedisca); scopolianum: Meyer, 1909, Kranchers Ent. Jahrb. 18: 147 (Epiblema); scopaliana (lapsus): Clutten, 1930, The Ent. 63: 114 (Catoptria); fulvana (non Stph.): Benander, 1950, Svensk Ins.fauna 10: 128, fig. 11r (&-Genitalien) (Epiblema); (235) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 9 hoenwartiana (err. typogr.): Bradley, 1959, Ent. Gaz. 10: t. 7 fig. 80 (Falter) (Ewcosma); cana (non Hw.): Hannemann, 1961, Dahls Tierwelt Dtschl. 48, Kleinschm. 1: fig. 245 (&- Genitalien) (Excosma). — Staudinger & Rebel, 1901: 116, No. 2085 (als scopoliana); Kennel, 1921: 561, t. 21 fig. 33 (4; als scopoliana); Pierce & Metcalfe, 1922: 72, t. 24 (3 2 -Geni- talien; als scopolina); Swatschek, 1958: 136 (Larvalmorphologie); Schantz, 1962: 4, fig. 3, 8 (& 2 -Genitalien). — Großbritannien; Europa; Ciskaukasien; Nordwestafrika. f. parvulana Stt. parvulana Stainton, 1858, Man. Brit. Butt. & Moths 2: 210 (Catoptria). — Staudinger & Rebel, 1901: 116, No. 2085a; Kennel, 1921: 561, t. 21 fig. 34 (4); Pierce & Metcalfe, 1922: 72, t. 24 (4 -Genitalien). E. (E.) balatonana (Osth.) comb. nova* (6) balatonana Osthelder, 1937, Iris 51: 109 (Epiblema); fulvana (non Stph.) Hannemann, 1961, Dahls Tierwelt Dtschl. 48, Kleinschm. 1: fig. 242 (4-Genitalien) (Ewcosma); synon. nov.: danicana Schantz, 1962, Notulae Ent. 42: 5, fig. 4, 11 (4 9-Genitalien) (Ezcosma). — Diese Arbeit: Taf. 4 Fig. 2—3 (Falter, ¢-Genitalien). — Dänemark; Deutschland; Österreich; Ungarn. E. (E.) afflicta Falk.* afflicta Falkovitsh, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 277, fig. 12, 13 (4 9- Genitalien) (Eucosma). — Sibirien (Provinzen Kemerovo und Krasnojarsk; Dschungar- und Transili-Alatau. E. (E.) expallidana (Hw.)* (7) expallidana Haworth, 1811, Lep. Brit.: 469 (Tortrix); rufana Stephens, [1829, Syst. Cat. Brit. Ins. 2: 180, No. 6983 (Semasia); nom. nud.}, 1834, Illustr. Brit. Ent., Haust. 4: 124 (Carpo- capsa, Epiblema); obumbratana Zeller, 1846, Isis: 240 (Grapholitha); westwoodiana Double- day, 1850, Synon. List Brit. Lep.: 26 (Catoptria); ibiceana Herrich-Schäffer, 1851, Syst. Bearb. Schm. Eur. 4: 249 (Semasia); lanceolana (part.): Lederer, 1859, Wien. Ent. Mschr. 3: 345 (Grapholitha). — Staudinger & Rebel, 1901: 116, No. 2088; Kennel, 1921: 566, t. 21 fig. 41 (2); Pierce & Metcalfe, 1922: 72, t. 25 (4 9-Genitalien); Benander, 1942: 44, fig. le, g (4 ®-Genitalien); Swatschek, 1958: 138, fig. 153, 154 (Larvalmorphologie); Hannemann, 1961: 126, fig. 244 (4-Genitalien), t. 15 fig. 2 (Falter); Schantz, 1962: 6, fig. 5, 9 (4 2- Genitalien). — Großbritannien; Nord- und Mitteleuropa; Norditalien; Spanien; Balkan; Polen; Ostbaltikum; Nordwest- und Ostrußland; Ukraine; Zentralasien (Alai-Gebirge); Südostsibirien. E. (E.) sublucidana (Kenn.) comb. nova sublucidana Kennel, 1901, Iris 13 (1900): 282 (Epiblema). — Staudinger & Rebel, 1901: 263, No. 2089bis; Kennel, 1921: 566, t. 21 fig. 42 (4). — Aragonien. E. (E.) scutana (Const.)* scutana Constant, 1893, Ann. Soc. Ent. France 62: 391, t. 11 fig. 3 (Grapholitha). — Stau- dinger & Rebel, 1901: 116, No. 2087; Kennel, 1921: 565, t. 21 fig. 39, 40 (4 9); Swatschek, 1958: 137, fig. 151, 152 (Larvalmorphologie); Hannemann, 1961: 126, fig. 243 ( &-Geni- talien), t. 15 fig. 3 (Falter); Razowski, 1961: 676, t. 88 fig. 12 ( 4 -Genitalien). — Südwest- und Süddeutschland; Schweiz; Südfrankreich; Mittelitalien; Sizilien. E. (E.) flavispecula Kuzn.* (8) flavispecula Kuznetzov, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 260, fig. 3a, 4a (& 9- Genitalien) (Eucosma). — Diese Arbeit: Taf. 5 Fig. 3, 5 (Falter). — Burgenland; Südtirol; Rumänien; Südukraine; Zentral- und Ostrußland; Sibirien. E. (E.) scorzonerana (Ben.)* scorzonerana Benander, 1942, Opusc. Ent. 7: 46, fig. If, h (4 2-Genitalien) (Epiblema). — Benander, 1950: 129, fig. 114 (&-Genitalien); Schantz, 1962: 8, fig. 6 (4-Genitalien); Kuznetzov, 1964 fig. 3b, 4b (4 ®-Genitalien). — Schweden; Finnland; Nordwestrußland; Ostbaltikum; Polen. 10 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (236) E. (E.) individiosana (Kenn.) comb. nova* (9) individiosana Kennel, 1901, Iris 13 (1900): 278 (Epiblema). — Staudinger & Rebel, 1901: 263, No. 2088guat.; Kennel, 1921: 556, t. 21 fig. 23 (4). — Algerien. E. (E.) cumulana (Gn.)* cumulana Guenée, 1845, Ann. Soc. Ent. France (2) 3: 189 (Catoptria); hornigiana Hornig, 1857, Wien. Ent. Mschr. 1: 72 (Grapholitha). — Staudinger & Rebel, 1901: 116, No. 2091; Kennel, 1921: 568, t. 21 fig. 46 (9); Hannemann, 1961: 125 nota, 220 fig. ( 4 -Genitalien), t. 11 fig. 6 (Falter). — Österreich; Ungarn; Rumänien; Mazedonien; ?Nordwestrußland. ssp. caradjai nom. nov. obscurana Caradja, 1916, Iris 30: 66 (Epiblema; praeoc. per F., 1798); cumulana (part.): Rebel, 1901, Stgr.-Rbl. Cat. Lep. Pal. Faun. 2: 116, No. 2091 (Epiblema). — Kennel, 1921: 568, t. 21 fig. 47 (8). — Südfrankreich; Spanien. E. (E.) umbratana (Stgr.) comb. nova* umbratana Staudinger, 1879, Horae Soc. Ent. Ross. 15: 253 (Grapholitha). — Staudinger & Rebel, 1901: 117, No. 2094; Kennel, 1921: 570, t. 21 fig. 50 (4); diese Arbeit: Taf. 5 Fig. 1 (&-Genitalien). — Mittelitalien; Sizilien; Albanien; Mazedonien; Kleinasien; Armenien. E. (E.) fervidana (Z.)* fervidana Zeller, 1847, Isis: 730 (Paedisca). — Staudinger & Rebel, 1901: 116, No. 2092; Kennel, 1921: 568, t. 21 fig. 48 (2); Hannemann, 1961: 128, fig. 249 (4-Genitalien), t. 12 fig. 3 (Falter); diese Arbeit: Taf. 5 Fig. 2 (&-Genitalien). — Niederösterreich; Kärnten; Dalmatien; ?Südkarpaten; Sizilien; Sardinien; Süditalien; Algerien. E. (E.) decolorana (Frr.)* decolorana Freyer, 1842, Neue Beitr. Schm.kunde 4: 48, t. 318 fig. 5 (Paedisca); ?hinnebergi- ana (non Fuchs): Krone, 1911, Jahresber. Wien. Ent. Ver. 21 (1910): 42 (Semasia). — Stau- dinger & Rebel, 1901: 116, No. 2090; Kennel, 1921: 567, t. 21 fig. 45 (4); Obraztsov, 1946: 38, fig. 2 (4-Genitalien); Hannemann, 1961: 128, fig. 250 (&-Genitalien) t. 11 fig. 6 (Falter). — Frankreich; Osten von Mitteldeutschland; Rhein; Österreich; Ungarn; Polen; Ukraine; Ost- und ?Nordwestrußland; ?Westchina (Bathang); ?Westpakistan (Karachi). E. (E.) kurdistana (Ams.) comb. nova kurdistana Amsel, 1959, Bull. Soc. Ent. Egypte 43: 58, t. 5 fig. 1 (4-Genitalien) (Epiblema). — Irak. E. (E.) victoriana (Kenn.) comb. nova* (10) victoriana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 86, t. 3 fig. 18, 19 (Epiblema). — Diese Arbeit: Taf. 6 Fig. 1—3 (Falter, 4-Genitalien). — Sibirien (Sajan). E. (E.) recentana (Zerny) comb. nova* fessana (non Mn.): Kennel, 1921, Pal. Tortr.: 548, t. 21 fig. 3 (4) (Epiblema); recentana Zerny, 1933, Iris 47: t. 1 fig. 9; 1934, ibid. 48: 16 (Epiblema). — Diese Arbeit: Taf. 6 Fig. 4 (Falter). — Nordlibanon; Mesopotamien. E. (E.) guentheri (Tgstr.) comb. nova* güntheri Tengström, 1869, Acta Soc. Faun. Flora Fenn. 10: 361 (71) (Grapholitha). — Staudinger & Rebel, 1901: 118, No. 2124; Kennel, 1921: 620, t. 23 fig. 21 (4); Toll, 1958: 74, fig. 10 (4 -Genitalien), t. 3 fig. 13 (Falter); diese Arbeit: Taf. 5 Fig. 4 (4 -Genitalien). — Finnland; Ostkarelien. Species incertae sedis E. (E.) atelosticta Meyr. atelosticta Meyrick, 1922, Exot. Micr. 2: 516 (Ewcosma). — Viette, 1952: 150 (Typus); Razowski, 1960a: 385, fig. 19 (®-Genitalien). — Ostchina (Schanghai). (237) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae M E. (E.) catharaspis (Meyr.) catharaspis Meyrick, 1922, Exot. Micr. 2: 514 (Ancylis); cataraspis Razowski, 1960, Polsk. Pismo Ent. 30: 386, fig. 20 (9-Genitalien) (Ewcosma). — Viette, 1952: 150 (Typus). — Ostchina (Schanghai). E. (E.) coeruleostriana Car. coeruleostriana Caradja, 1939, Iris 53: 11 (Ewcosma). — Nordchina (Schansi). E. (E.) disjectana (Kenn.) comb. nova disjectana Kennel, 1921, Pal. Tortr.: 558, t. 21 fig. 26 (Epiblema); disjectama (err. typogr.): Petersen, 1924, Lep.-Fauna Estl.: 459 (Epiblema). — ?Livland. E. (E.) erebantra Meyr. erebantra Meyrick, 1937, Iris 51: 181 (Ewcosma). — Yiinnan. E. (E.) ignotana (Car.) comb. nova ignotana Caradja, 1916, Iris 30: 64 (Semasia). — Südostsibirien (Kasakewitsch). Sg. Pygolopha Ld., 1859 E. (P.) lugubrana (Tr.)* lugubrana Treitschke, 1830, Schm. Eur. 8: 26 (Penthina); tinacriana Lederer, 1859, Wien. Ent Mschr. 3: 280, t. 2 fig. 1, 2 (Pygolopha); amandana (ex err.): Aigner-Abafi, 1898, Illustr. Zschr. Ent. 3: 312 (Phtheochroa); trinacriana (laps.): Rebel, 1901, Stgr.-Rbl., Cat. Lep. Pal. Faun. 2: 115 (Pygolopha). — Staudinger & Rebel, 1901: 115, No. 2066; Kennel, 1908: 53, t. 2 fig. 5—8 (Geäder, 9-Hinterleibspitze); 1921: 545, t. 20 fig. 62, 63 (4 2); Hannemann, 1961: 129, fig. 251-2515 (Kopf, Geäder, 4 -Genitalien). — Ungarn; Niederösterreich; Kara- wanken; Tschechoslowakei; Südtirol; Istrien; Mazedonien; Sizilien; Südfrankreich. f. tarica Hartig tarica Hartig, 1949, Bol. Assoc. Rom. Ent. 4: 2, t. 1 fig. 1 (Pygolopha). — Garda-See. Sg. Palpocrinia Kenn., 1919 E. (P.) ottoniana (Kenn.) comb. nova* ottoniana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 66, t. 2 fig. 20 (Falter), t. 4 fig. 5 (Kopf) (Palpocrinia). — Obraztsov, 1955: 163, fig. 279 (Kopf); 1956: 117; 1965: 25; diese Arbeit: Taf. 7 Fig. 1 (&-Genitalien). — Lob-Noor. Sg. Phaneta Stph., 1852 E. (P.) niveicaput (Wlsm.) comb. nova* niveicaput Walsingham, 1900, Ann. Mag. NH. (7) 6: 406 (Thiodia). — Issiki, 1957: 63, t. 10 fig. 296 (Falter); diese Arbeit: Taf. 6 Fig. 5, Taf. 7 Fig. 2—3 (& Q-Genitalien). — Japan. E. (P.) brachysticta Meyr.* brachysticta Meyrick, 1935, in Caradja & Meyrick, Mater. Microlep. Fauna chines. Prov.: 55 (Eucosma). — Diese Arbeit: Taf. 7 Fig. 4—5 (Falter, 4 -Genitalien). — Ostchina. E. (P.) nitorana Kuzn.* nitorana Kuznetzov, 1962, Bull. Soc. Ent. Mulhouse: 57, fig. 6 (4 9-Genitalien) (Eucosma, Phaneta). — Südostsibirien. E. (P.) pergratana (Rbl.) comb. nova* pergratana Rebel, 1914, Iris 28: 274, t. 4 fig. 11 (Steganoptycha). — Kuldscha. 12 E. TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (238) (P.) messingiana (F.R.)* messingiana Fischer v. Röslerstamm, 1837, Abb. Bericht. Ergänz. Schm.kunde: 89, t. 40 fig. 14-b (Grapholitha); wimmeriana (part.): Lederer, 1859, Wien. Ent. Mschr. 3: 337 (Grapho- litha). — Staudinger & Rebel, 1901: 114, No. 2034; Kennel, 1916: 524, t. 20 fig. 21 (8); Benander, 1950: 120, fig. 11f (&-Genitalien), t. 7 fig. 34 (Vorderflügel); Hannemann, 1961: 133, fig. 264 (&-Genitalien), t. 15 fig. 20 (Falter). — Nord- und Mitteleuropa; Ostbaltikum; Ungarn; Südpolen; Ukraine; Ostrußland; Südostsibirien; ?Spanien. . (P.) ursulana (Kenn.) comb. nova ursulana Kennel, 1919, Mitt. Miinchn. Ent. Ges. 8: 85, t. 3 fig. 17 (Semasia). — Ostrußland (Uralsk); Zentralsibirien (Ostsajan). . (P.) nessebarana (Soffner) comb. nova* nessebarana Soffner, 1962, Dtsche Ent. Zschr. (N.F.) 9: 140, fig. 5 (4 9-Genitalien) (Se- masia). — Ostbulgarien. . (P.) sparsana (Rbl.) comb. nova* sparsana Rebel, 1935, Bull. Soc. Lépid. Génève 7: 175 (Semasia). — Diese Arbeit: Taf. 7 Fig. 6 (Falter). — Kleinasien. (P.) pylonitis Meyr.* pylonitis Meyrick, 1932, Exot. Micr. 4: 307 (Eucosma). — Clarke, 1958: 384, t. 191 fig. 3—34 (Falter, 4 -Genitalien). — Kaschmir. (P.) pauperana (Dup.)* pauperana Duponchel, 1843, Hist. Nat. Lépid. France, Suppl. 4: 170, t. 64 fig. 10 (Cochylis); ?paykulliana (non F.): Haworth, 1811, Lep. Brit: 435 (Tortrix). — Staudinger & Rebel, 1901: 110, No. 1971; Kennel, 1916: 493, t. 19 fig. 40 (8); Pierce & Metcalfe, 1922: 70, t. 24 (4 Q-Genitalien). — England; Mittel- und Südeuropa; Polen; Kleinasien. (P.) muliebris Meyr. muliebris Meyrick, 1922, Exot. Micr. 2: 515 (Ewcosma). — Razowski, 19604: 385, fig. 11 ( 4-Genitalien). — Korea. . (P.) tundrana (Kenn.)* tundrana Kennel, 1900, Iris 13: 148, t. fig. 24, 25 (Semasia); cordulana (Kennel, il.) Rebel, 1917, Iris 30: 192 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2044bis; Kennel, 1916: 523, t. 20 fig. 20 (2); Osthelder, 1939: 90, t. 2 fig. 49 (2); Toll, 1939: 245, 259, t. 2 fig. 8 (&-Genitalien); Hannemann, 1961: 133, fig. 266 (4-Genitalien), t. 5 fig. 13, t. 21 fig. 7 (Falter). — Zentralasien; Sibirien; Mandschurei; Ostrußland; Ukraine; Südostpolen; Ungarn; Südbayern. (P.) brigittana (Kenn.) comb. nova brigittana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 83, t. 3 fig. 15 (Semasia). — Ost-Tannuola. . (P.) sybillana (Kenn.) comb. nova sybillana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 82, t. 3 fig. 14 (Semasia). — Sibirien (Sajan). (P.) amellana (Preiss.)* amellana Preissecker, 1930, Verh. zool.-bot. Ges. Wien 80: (20), fig. 3 rechts (Semasia). — Hannemann, 1961: 130 nota, 220 fig. (&-Genitalien), t. 21 fig. 21 (Falter); diese Arbeit: Taf. 6 Fig. 6, Taf. 8 Fig. 2, Taf. 9 Fig. 1 (Falter, 4 9 -Genitalien). — Österreich. . (P.) krygeri (Rbl.)* krygeri Rebel, 1937, Zschr. Österr. Ent. Ver. 22: 42, t. 2 fig. 3 (Semasia). — Worm-Hansen & Larsson, 1947: 174—195, 187 fig. 1—10 (Falter, Kopf, Beine, Puppe, Beschädigung), 188 (239) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 13 fig. 1—20 (Larvalmorphologie), 190 fig. 1, 3, 4 (4 Q-Genitalien); Hannemann, 1961: 129 nota, 220 fig. (4-Genitalien), t. 21 fig. 22 (Falter); diese Arbeit: Taf. 8 Fig. 4, 6, Taf. 9 Fig. 2 (Falter, 3 Q-Genitalien). — Dänemark. E (P.) tarandana (Möschl.)* tarandana Möschler, 1874, Stett. Ent. Ztg. 35: 165 (Grapholitha); taradana (err. typogr.): Heinrich, 1923, Bull. U.S. Natl. Mus. 123: 296 (Thiodia). — Staudinger & Rebel, 1901: 114, No. 2048; Kennel, 1916: 517, t. 20 fig. 7 (&); Heinrich, 1923: 52, t. 16 fig. 112 ( 4 -Geni- talien). — Labrador; Kanada; ?Dahurien. E. (P.) sinensis (Wlsm.) comb. nova* sinensis Walsingham, 1900, Ann. & Mag. N.H. (7) 6: 406 (Thiodia). — Diese Arbeit: Taf. 8 Fig. 3, 5 (Falter, ¢-Genitalien). — Ostchina. E. (P.) urbana (Kenn.) comb. nova urbana Kennel, 1901, Iris 13 (1900): 271 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2022bis; Kennel, 1916: 526, t. 20 fig. 26 (4). — Spanien; Kleinasien. E. (P.) glebana (Snell.) comb. nova* glebana Snellen, 1883, Tijdschr. v. Ent. 26: 206, t. 12 fig. 6, 64 (Falter, Kopf) (Grapholitha, Semasia). — Staudinger & Rebel, 1901: 114, No. 2024; Kennel, 1916: 527, t. 20 fig. 27, 28 (& 2); Issiki, 1957: 58, t. 9 fig. 262 (Falter); diese Arbeit: Taf. 8 Fig. 7 (4 -Genitalien). — Südostsibirien; Japan. E. (P.) gracilis (Fil.) comb. nova gracilis Filipjev, 1925, Jahrb. Martianov. Staatsmus. 2 (3): 61 (Semasia). — Sibirien (Minus- sinsk). E. (P.) cetratana (Kenn.) comb. nova cetratana Kennel, 1901, Iris 13 (1900): 264 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2032bis; Kennel, 1916: 510, t. 19 fig. 76 (&). — Originalfundort und Verbreitung un- bekannt. E. (P.) fraudulentana (Kenn.) comb. nova fraudulentana Kennel, 1901, Iris 13 (1900): 265 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2045bi5; Kennel, 1916: 510, t. 19 fig. 77 (2). — Alatau-Gebirge. E. (P.) intermediana (Kenn.) comb. nova intermediana Kennel, 1900, Iris 13: 146, t. f. 21 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2045ter; Kennel, 1916: 511, t. 19 fig. 78 (4). — Amur. E. (P.) malitiosana (Kenn.) comb. nova malitiosana Kennel, 1901, Iris 13 (1900): 266 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2054bis; Kennel, 1916: 511, t. 19 fig. 79 (&). — Mongolei (Uliassutai). E. (P.) lignana (Snell.) comb. nova lignana Snellen, 1883, Tijdschr. v. Ent. 26: 235, t. 12 fig. 5, 5a (Falter, Kopf) (Grapholitha, Semasia); messingiana (part.): Rebel, 1901, Stgr.-Rbl. Cat. Lep. Pal. Faun. 2: 114, No. 2034 (Semasia); elongata Filipjev, 1924, Jahrb. Martianov. Staatsmus. 2 (3): 24, 38 (Semasia). — Filipjev, 1930a: 6, t. 2 fig. 7 (Falter). — Sibirien (Irkutsk, Tschita). E. (P.) metana (Kenn.) comb. nova metana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 84, t. 3 fig. 16 (Semasia). — Ostchina (Schanghai). E. (P.) striatiradix Kuzn.* striatiradix Kuznetzov, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 262, fig. 5 (4 9- Genitalien) (Eucosma, Phaneta). — Südostsibirien; Minussinsk. 14 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (240) E. (P.) tripoliana (Barr.)* tripoliana Barrett, 1880, Ent. Mo. Mag. 17: 84 (Catoptria); aemulana (non Schl.): Pierce & Metcalfe, 1922, Genit. Brit. Tortr.: 71, t. 24 (& 9-Genitalien) (Phaneta). — Staudinger & Rebel, 1901: 114, No. 2032; Kennel, 1916: 519, t. 20 fig. 11 (2); Wolff, 1952: 61 (Syste- matik); Swatschek, 1958: 140 (Larvalmorphologie; als aemulana); Hannemann, 1961: 132, fig. 259 (Â-Genitalien), t. 20 fig. 11 (4). — England; Holland; Deutschland; Frankreich; ?Polen. E. (P.) aspidiscana (Hb.)* ?cervana Scopoli, 1763, Ent. Carn.: 234 (Phalaena); ?obscurana Fabricius, 1798, Suppl. Ent. Syst.: 476 (Pyralis); ?petiverella (non L.): Charpentier, 1821, Zinsler etc.: 120 (Tizea); aspidiscana Hübner, [1814—1817, Samml. eur. Schm., Tortr.: t. 41 fig. 256; non bin.}, 1822, Syst.-alph. Verz.: 58 (Olethreutes); ?zachana Hübner, [1814—1817, op. cit.: t. 38 fig. 243; non bin.}, 1822, op. cit.: 66 (Olethreutes); aspidana Frölich, 1828, Enum. Tortr. Würt.: 96 (Tortrix); dahlbomiana Zetterstedt, 1840, Ins. Lap.: 987 (Phoxopteris); nebritana (ex err.): Herrich-Schäffer, 1848, Syst. Bearb. Schm. Eur. 4: t. 33 fig. 241 (non bin.). — Staudinger & Rebel, 1901: 114, No. 2049; Kennel, 1916: 517, t. 20 fig. 8 (9); Pierce & Metcalfe, 1922: 71, t. 24 (& $-Genitalien); Higgins, 1934: 169, fig. 1 (Labialpalpus), t. 2 fig 3, 4 (3 2); Benander, 1928: 147, fig. 4m-o (Larvalmorphologie); 1950: 119, fig. 117 (4-Genitalien), t. 7 fig. 30 (Vorderflügel); Swatschek, 1958: 139, fig. 156 (Larvalmorphologie); Hannemann, 1961: 130, fig. 257 (&-Genitalien), t. 13 fig. 26 (Falter). — Großbritannien; ganz Europa und Sibirien; Zentralasien; Armenien; Kleinasien; Nordwestafrika. f. rubescana Const. rubescana Constant, 1895, Bull. Soc. Ent. France: LI (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 20494; Kennel, 1916: 517, t. 20 fig. 9 (2). E. (P.) saussureana (Ben.) comb. nova* saussureana Benander, 1928, Ent. Tidskr. 29: 142, fig. 4a-l ( $-Genitalien, Larval- und Pupal- morphologie) (Phaneta, Semasia). — Benander, 1950: 119, fig. 114 ( & -Genitalien), t. 7 fig. 31 (Vorderflügel). — Schweden; Finnland. E. (P.) gradensis (Galv.) comb. nova* gradensis Galvagni, 1909, Mitt. Naturwiss. Ver. Wien. Univ. 7: 248 (Semasia). — Diese Arbeit: Taf. 8 Fig. 1, 8 (Falter, 4 -Genitalien). — Dalmatien (Grado). E. (P.) catoptrana (Rbl.) status & comb. nov. catoptrana Rebel, 1903, Verh. zool.-bot. Ges. Wien 53: 91 (Semasia); rubescana (non Const.): Huggins, 1924, The Ent. 57: 14 (Catoptria); synon. nov.: heringiana Jäckh, 1953, Zschr. Lepid. 3: 43, fig. 1, 2 (Catoptria, Semasia). — Kennel, 1921: 719; Huggins, 1934: 169, fig. 1 (Labial- palpus), t. 2 fig. 1, 2 (4 2) (als rubescana); Bradley, 1959: 62, fig. 6 (4-Genitalien), t. 8 fig. 85 (Falter); Hannemann, 1961: 129, fig. 254 ( Â-Genitalien), t. 20 fig. 22 (Falter) (bei den beiden letzteren Autoren als heringiana). — England; Ostfriesische Inseln (Wangeroog); Osterreich; Spanien. E. (P.) conterminana (Gn.)* conterminana Guenée, 1845, Ann. Soc. Ent. France (2) 3: 189 (Catoptria); caecimaculata Duponchel, 1835, Hist. Nat. Lép. France 9: t. 249 fig. 5% (non bin.). — Herrich-Schäffer, 1848: t. 42 fig. 297; 1851: 247; Staudinger & Rebel, 1901: 114, No. 2051; Kennel, 1916: 519, t. 20 fig. 12 (2); Pierce & Metcalfe, 1922: 71, t. 24 (4 9-Genitalien); Swatschek, 1958: 140, fig. 158 (Larvalmorphologie); Hannemann, 1961: 130, fig. 258 (4-Genitalien), t. 13 fig. 23 (Falter). — Großbritannien; ganz Europa; Ciskaukasien; Armenien; Südostsibirien; Zentralasien. E. (P.) incinerana (Const.) comb. nova incinerana Constant, 1888, Ann. Soc. Ent. France (6) 8: 167, t. 4 fig. 6 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2050; Kennel, 1916: 520, t. 20 fig. 13 (4). — Süd- frankreich. (241) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 15 E. (P.) incana (Z.)* incana Zeller, 1946, Isis: 239 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2046; Kennel, 1916: 520, t. 20 fig. 14 (4); Swatschek, 1958: 141 (Larvalmorphologie); Hannemann, 1961: 130, fig. 256 ( -Genitalien), t. 13 fig. 24 (Falter). — Mittel- und Südeuropa; Holland; Ostbaltikum; Polen; Ukraine; Rußland. E. (P.) aemulana (Schläg.)* aemulana Schläger, 1849, Ber. lepid. Tauschver. Jena: 39 (Grapholitha); latiorana Herrich- Schäffer, 1851, Syst. Bearb. Schm. Eur. 4: 248 (Semasia) [1848, Tortr.: t. 44 fig. 312; non bin.}; modestana Stainton, 1859, Man. Brit. Butt. & Moths 2: 242 (Grapholitha); ?decolorana (non Frr.): Barrett, 1885, Ent. Mo. Mag. 21: 109 (Catoptria). — Staudinger & Rebel, 1901: 114, No. 3031; Kennel, 1916: 518, t. 20 fig. 10 (9); Pierce & Metcalfe, 1922: 71, t. 24 (4 S- Genitalien; als latiorana); Wolff, 1952: 61, 62 (Systematik; als /atiorana); Swatschek, 1958: 140, fig. 157 (Larvalmorphologie; als latiorana); Hannemann, 1961: 130, fig. 255 ( &-Geni- talien), t. 13 fig. 22 (Falter). — England; Schottland; ?Dänemark; Holland; Mitteleuropa; Frankreich; ?Slowakei; ?Rumänien; Ostbaltikum; Polen; Nordwestrußland; ?Transili-Alatau. E. (P.) intacta (Wlsm.) comb. nova * intacta Walsingham, 1900, Ann. & Mag. NH. (7) 6: 405 (Thiodia). — Issiki, 1957: 59, t. 9 fig. 265 (Falter); diese Arbeit: Taf. 9 Fig. 3, Taf. 10 Fig. 2 (Falter, 3 -Genitalien). — China; Japan. E. (P.) cretaceana (Kenn.) comb. nova cretaceana Kennel, 1899, Iris 12: 42, t. 1 fig. 41 (Grapholitha). — Staudinger & Rebel, 1901: 121, No. 2170; Kennel, 1916: 521, t. 20 fig. 15 (4). — Spanien; Albanien. E. (P.) metzneriana (Tr.)* metzneriana Treitschke, 1830, Schm. Eur. 8: 277 (Cochylis). — Staudinger & Rebel, 1901: 114, No. 2043; Kennel, 1916: 522, t. 20 fig. 17 (4); Issiki, 1957: 59, t. 9 fig. 266 (Falter); Swatschek, 1958: 142 (Larvalmorphologie); Okano, 1959: 260, t. 174 fig. 25 (2); Hanne- mann, 1961: 133, fig. 265 (4-Genitalien), t. 15 fig. 14 (Falter). — Mitteleuropa; Holland; Belgien; Frankreich; Ungarn; Südtirol; Dalmatien; Polen; Ukraine; Süd- und Ostrußland; Iran; Sibirien; China; Korea; Japan; ?Nordwestafrika. f. osthelderi Dufr. osthelderi Dufrane, 1955, Bull. Inst. Roy. Sci. Nat. Belg. 31 (33): 3 (Eucosma). — Belgien. f. anserana Hein. status nov. anserana Heinemann, 1863, Schm. Dtschl. u. Schweiz (2) 1 (1): 169 (Grapholitha, Semasia). — Staudinger & Rebel, 1901: 114, No. 2028; Kennel, 1916: 521, t. 20 fig. 16 (4); Toll, 1939: 245, 259, t. 1 fig. 6, 7 (&-Genitalien). — Ungarn; Bulgarien; Südpolen; Ukraine; Ostrußland. f. joannisiola Duft. joannisiola Dufrane, 1955, Bull. Inst. Roy. Sci. Nat. Belg. 31 (33): 3 (Ewcosma). — Schanghai. E. (P.) wimmerana (Tr.)* wimmerana Treitschke, 1835, Schm. Eur. 10 (3): 256 (Grapholitha); wimmeriana Lederer, 1859, Wien. Ent. Mschr. 3: 337 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2044; Kennel, 1916: 522, t. 20 fig. 18 (4); Hannemann, 1961: 132, fig. 262 ( 4 -Genitalien), t. 20 fig. 21 (Falter). — Nordfrankreich; Norditalien; Ungarn; Ukraine; Ostrußland; Armenien. E. (P.) maritima (Westw. & Humphr.)* maritima Westwood & Humphreys, 1845, Brit. Moths 2: 138 (Carpocapsa); wimmerana (non Tr.): Doubleday, 1849, Synon. List Brit. Lepid.: 26 (Catoptria); ?lacteana (non Tr.): Stephens, 1852, List Spec. Brit. Anim. 10: 62 (Grapholita); caudidulana Nolcken, 1870, Arb. Naturf. Ver. Riga (N.F.) 3: 413 (Grapholitha). — Staudinger & Rebel, 1901: 116, No. 2080; Kennel, 16 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (242) 1916: 523, t. 20 fig. 19 (4; bei den beiden Autoren als candidulana); Pierce & Metcalfe, 1922: 70, t. 24 (4 9-Genitalien); Worm-Hansen & Larsson, 1947: 190, fig. 2 ( £-Genitalien; als candidulana); Benander, 1950: 119, fig. 11g (&-Genitalien), t. 7 fig. 33 (Vorderflügel); Swatschek, 1958: 142 (Larvalmorphologie); Hannemann, 1961: 133, fig. 263 (&-Genitalien), t. 15 fig. 23 (Falter). — England; Dänemark; Holland; Belgien; Mitteleuropa; Frankreich; Südmähren; Skandinavien; Finnland; Nordwestrußland; Ostbaltikum; Polen; Ukraine. E. (P.) apocrypha Falk.* apocrypha Falkovitsh, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 280, fig. 16, 17 (4 2- Genitalien) (Eucosma, Phaneta). — Kasachstan. ssp. tagarica Falk. tagarica Falkovitsh, 1964, Trudy Zool. Inst. Akad. Nauk SSSR 34: 282 (Eacosma). — Sibirien (Provinz Krasnojarsk). E. (P.) nigromaculana (Hw.)* nigro-maculana Haworth, 1811, Lep. Brit.: 436 (Tortrix); nigromaculana: Stephens, 1829, Syst. Cat. Brit. Ins. 2: 178, No. 6962 (Anchylopera); nigromaculata Wood, 1839, Ind. Ent.: 144, t. 32 fig. 962 (Anticlea); freyeriana Fischer v. Röslerstamm, 1839, Abb. Bericht. Ergänz. Schm.kunde: 132, t. 51 fig. 3 (Grapholitha); nigrimaculana Barrett, 1874, Ent. Mo. Mag. 11: 134; campoliliana (non Schiff.): Hannemann, 1961, Dahls Tierw. Mitteleur. 48, Microlep. 1: 132, fig. 261 (&-Genitalien), t. 13 fig. 19 (Falter) (Ewcosma, Phaneta). — Staudinger & Rebel, 1901: 110, No. 1972; Kennel, 1916: 496, t. 19 fig. 48 (2); Pierce & Metcalfe, 1922: 70, t. 24 (& 2-Genitalien); Swatschek, 1958: 139, fig. 155 (Larvalmorphologie). — Groß- britannien; Nordfrankreich; Mitteleuropa; Balkanhalbinsel; Südungarn. f. capitinivana Bruand albana Haworth, 1811, Lep. Brit.: 436 (Tortrix; nom. praeocc.); capitinivana Bruand, 1847, Mém. Soc. emul. Doubs 2, Cat. Microlép. Doubs: 45 (Grapholitha); nigromaculana (part.): Kennel, 1916, Pal. Tortr.: 496, t. 19 fig. 47 (9) (Semasia). ssp. ussuriana Car. ussuriana Caradja, 1916, Iris 30: 60 (Steganoptycha). — Südostsibirien (Kasakewitch). E. (P.) galactica nom. nov.* galactica Obraztsov: diese Arbeit; ?/ucimana (Kenn. il.) Caradja, 1916, Iris 30: 63 (Semasia; nom. nud.); luciana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8: 81, t. 3 fig. 13 (Semasia). — Diese Arbeit: Taf. 9 Fig. 4 (4-Genitalien). — Südukraine; Rumänien; Ostrußland; Sibirien. f. caliacrana Car. caliacrana Caradja, 1931, Mem. Sect. Stiint. Acad. Rom. (3) 7 (8): 329 (Semasia). E. (P.) sardoensis (Rbl.) comb. nova * sardoensis Rebel, 1935, Iris 50: 94 (Semasia). — Diese Arbeit: Taf. 11 Fig. 1 (Falter, &- Genitalien). — Sardinien. E. (P.) suomiana (Hoffm.) comb. nova suomiana Hoffmann, 1893, Stett. Ent. Ztg. 54: 142 (Grapholitha). — Staudinger & Rebel, 1901: 115, No. 2052; Kennel, 1921: 718; Benander, 1950: 120, fig. 114 ( &-Genitalien). — Schweden; Finnland. E. (P.) abacana (Ersch.) comb. nova * abacana Erschoff, 1877, Horae Soc. Ent. Ross. 12 (1876): 342 (Grapholitha); opulentana Christoph, 1881, Bull. Soc. Imp. Nat. Moscou 56 (3): 413 (Grapholitha; nom. praeocc.); abascana (err.): Kurentsov, 1950, Soobsch. Dalnevost. Fil. Akad. Nauk SSSR: 30 (Semasia). — Staudinger & Rebel, 1901: 114, No. 2047; Kennel, 1916: 506, t. 19 fig. 69, 70 ( 2); Issiki, 1957: 60, t. 9 fig. 277 (2). — Zentralasien; Süd- und Siidostsibirien; ?Südkarpaten (Retyezat- Gebirge). (243) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 17 E. (P.) oculatana (Kenn.) comb. nova oculatana Kennel, 1900, Iris 13: 143, t. fig. 19 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2047bis; Kennel, 1916: 507, t. 19 fig. 71 (2). — Ostsibirien (Irkutsk). E. (P.) pupillana (Cl.)* pupillana Clerck, 1759, Icones Ins.: t. 11 fig. 9 (non bin.) [Linné, 1761, Fauna Suec.: 349 (Phalaena Tortrix)]}; absinthiana [Hübner, 1796—1799, Samml. eur. Schm., Tortr.: t. 6 fig. 34; non bin.] Haworth, 1811, Lep. Brit.: 457 (Tortrix); absynthiana Hübner, 1822, Syst.-alph. Verz.: 58 (Olethreutes). — Staudinger & Rebel, 1901: 114, No. 2042; Kennel, 1916: 507, t. 19 fig. 72 (2); Pierce & Metcalfe, 1922: 71, t. 24 (4 ®-Genitalien); Benander, 1950: 118, fig. 11e (&-Genitalien), t. 7 fig. 32 (Vorderflügel); Swatschek, 1958: 141 (Larvalmorpholo- gie); Hannemann, 1961: 129, fig. 253 (4 -Genitalien), t. 13 fig. 21 (Falter). — Ganz Europa. ssp. saerdabana ssp. nova saerdabana Obraztsov, diese Arbeit: Taf. 10 Fig. 1—4, Taf. 11 Fig. 4 (Falter, 4 9-Genitalien). — Nordpersien; ?Armenien. E. (P.) mirificana (Peyer.)* mirificana Peyerimhoff, 1876, Ann. Soc. Ent. France (5) 6: 590, t. 12 fig. 18 (Grapholitha). — Staudinger & Rebel, 1901: 113, No. 2023; Kennel, 1916: 508, t. 19 fig. 78 (9); Hannemann, 1961: 129 nota, 220 fig. (4 -Genitalien), t. 22 fig. 2 (Falter). — Schweiz; Französische Alpen. E. (P.) tetraplana (Möschl.) comb. nova * tetraplana Möschler, 1866, Berl. Ent. Zschr. 10: 148 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2045; Kennel, 1916: 509, t. 19 fig. 74 (4); diese Arbeit: Taf. 11 Fig. 5 (4 -Genitalien). — Ostrußland; Armenien; Syrien. E. (P.) paetulana (Kenn.) comb. nova paetulana Kennel, 1900, Iris 13: 145, t. fig. 20 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2043bis; Kennel, 1916: 509, t. 19 fig. 75 (2). — Südural; ?Kleinasien. E. (P.) esmodes Meyr. esmodes Meyrick, 1937, Exot. Micr. 5: 99 (Eucosma). — Amsel, 1943: 312, t. 8 fig. 52 ( 4-Genitalien); Wiltshire, 1957: 141, t. 13 fig. 52 (&-Genitalien). — Irak. Species incertae sedis E. (P.) affectana (Kenn.) comb. nova affectana Kennel, 1901, Iris 13 (1900): 267 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2030bis; Kennel, 1916: 511, t. 19 fig. 80 (2). — Transkaukasien. E. (P.) albicostella (Trti. & Krüg.) comb. nova albicostella Turati & Kriiger, 1936, Mem. Soc. Ent. Ital. 15: 75, t. 11 fig. 25 (Semasia). — Kyrenaika. E. (P.) astragalana (Stgr.) comb. nova astragalana Staudinger, 1871, Berlin. Ent. Zschr. 14 (1870): 282 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2029; Kennel, 1916: 513, t. 19 fig. 83 (Falter). — Ostrufland. E. (P.) bactrana (Kenn.) comb. nova bactrana Kennel, 1901, Iris 13 (1900): 269 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2929ter; Kennel, 1916: 514, t. 20 fig. 1 (4). — Südspanien; Nordwestafrika; Iran; ?Ferghana. E. (P.) demissana (Kenn.) comb. nova demissana Kennel, 1901, Iris 13 (1900): 268 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2029bis; Kennel, 1916: 513, t. 19 fig. 84 (4). — Andalusien. 18 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (244) E. (P.) directa Meyr. directa Meyrick, 1912, Exot. Micr. 1 34 (Ewcosma). — Taurus. E. (P.) ephedrana (Chr.) comb. nova ephedrana Christoph, 1877, Horae Soc. Ent. Ross. 12 (1876): 291, t. 8 fig. 64 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2030; Kennel, 1916: 512, t. 19 fig. 82 (3). — Palästina; Hyrkanien; Westturkestan (Kopet-Dagh). E. (P.) giarabubensis (Trti.) comb. nova giarabubensis Turati, 1930, Atti Soc. Ital. Sci. Nat. 69: 76, t. 2 fig. 2. Vertikalreihe, unten (Semasia). — Kyrenaika. E. (P.) infirmana (Kenn.) comb. nova infirmana Kennel, 1900, Iris 13: 153, t. fig. 29 (Semasia). — Staudinger & Rebel, 1901: 262, No. 2030ter; Kennel, 1916: 512, t. 19 fig. 81 (2). — Zentralasien (Aschabad). E. (P.) luciana (Chret.) luciana Chrétien, 1908, Le Nat. 30: 245 (Epiblema). — Razowski, 1961: 677, t. 93 fig. 27 (2 -Genitalien). — Südfrankreich. E. (P.) lyrana (Snell.) comb. nova lyrana Snellen, 1883, Tijdschr. v. Ent. 26: 208, t. 12 fig. 7, 7a (Falter, Kopf) (Grapholitha, Semasia). — Staudinger & Rebel, 1901: 115, No. 2054; Kennel, 1916: 528, t. 20 fig. 33 (&). — Irkutsk; Amur. E. (P.) polyxena Meyr. polyxena Meyrick, 1937, Iris 51: 179 (Ewcosma). — Yünnan. E. (P.) resupinatana (Kenn.) comb. nova resupinatana Kennel, 1901, Iris 13 (1900): 270 (Semasia). — Staudinger & Rebel, 1901: 262, No. 1997bis; Kennel, 1916: 516, t. 20 fig. 5 (4). — Schweiz (Wallis). E. (P.) teliferana (Chr.) comb. nova teliferana Christoph, 1881, Bull. Soc. Imp. Nat. Moscou 56 (3): 415 (Grapholitha). — Staudinger & Rebel, 1901: 114, No. 2025; Kennel, 1916: 528, t. 20 fig. 32 (4). — Südussuri. E. (P.) verecundana (Car.) comb. nova verecundana Caradja, 1916, Iris 30: 63 (Semasia). — Alai-Gebirge. Appendix Hemimene marmarocyma Meyrick, 4, Tien-Mu-Shan, 5300’, June 1932 (Höne), Prip. No. 6681 (BM) ist abgebildet auf Taf. 10 Fig. 1, 4, Taf. 11 Fig. 2, diese Arbeit. Kommentar zum Katalog der Ewcosma-Arten 1. Eucosma (Eucosma) conformana (Mn.). — Typen der conformana: Lectotypus, Männchen (Genitalpräparat No. V. 38), „Type 1872 z.b.G.” (keine genaueren Angaben); Lectoparatypus, Männchen, Ragusa, Dalmatien, 1868 (Mann); beide im Wiener Naturhistorischen Museum. Typen der significantana Kenn.: Lectotypus, Mannchen (Genitalpräparat No. B. 33), Mazedonien (Kr.); Lectoallotypus, Weibchen (Genitalpräparat No. B. 34), Beirut, Syrien (CR.); beide im Z.M.B. 2. E. (E.) gypsatana (Kenn.). — Holotypus, Männchen (Genitalpräparat No. 5730), Akschehir, Anatolien, 1900 (Caradja, No. 2011; Walsingham, No. 71784); B.M. (245) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 19 3. E. (E.) albarracina Hartig. — Blieb mir unbekannt. Nach der Angabe des aufstellenden Autors gehört diese Art zur fulvana-expallidana-Gruppe und unterscheidet sich von diesen Arten durch die goldgelben Vorderflügel, einen hellen länglichen Fleck nach dem Costalumschlag und einen aus hellen und dunklen Schuppen gebildeten Fleck vor und unterhalb der Mitte der Mittelzelle des Vorderflügels. Das die Originalbeschreibung begleitende Photo stellt zweifellos eine Eucosma-Art dar, welche einen deutlichen Vorderflügelcostalumschlag hat und eher erinnert an albidulana oder albuneana, aber anscheinend mit keiner von diesen identisch ist. 4. E. (E.) rigidana (Snell.). — Lectotypus, Mannchen (Genitalpriparat No. 2638), Insel Askold, 1878 (D.; v. Hed.); M.L. Kennel (1916) hatte ein Weibchen und er stellte rigidana irrtümlicher- weise in die Gattung Semasia. 5. E. (E.) monstratana (Rbl.). — Lectotypus, Männchen (Genitalpräparat No. 2048), Schanfigg bei Chur, 1600 m, 16.VII.1903 (Stange; der Falter und seine Genitalien wurden von Hannemann, 1961, abgebildet); Lectoallotypus, Weibchen (Genitalpriparat No. V. 28), gleiche Zettelangaben; Wiener Naturhistorisches Museum. 6. E. (E.) balatonana (Osth.). — Als “var.” balatonana beschrieb Osthelder (1937) kleinere Exemplare einer Art aus Ungarn, die er mit expallidana konspezifisch zu sein vermutete. In seiner Sammlung (jetzt in der Z.S.M.) befinden sich mehrere Exemplare der balatonana, aber von einem besonderen Interesse sind jene zwei Männchen, die Osthelder als Typen bezeichnete. Eines von diesen, als ‘“Holotyp” bezettelt, stammt aus Puszta Peszér (12.VII.1929, L. Osthelder; Genital- präparat No. 3-Obr. 8/8 58) und kann nur als ein Pseudotypus betrachtet werden, da Osthelder in der Originalpublikation alle Exemplare aus dieser Lokalität der namenstypischen expallidana-Form zuzog. Nur die Exemplare aus Vörs (Komitat Somogy) behandelte er als balatonana. Deshalb bezeichne ich das zweite der beiden erwähnten Männchen als Lectotypus der balatonana, obwohl es von Osthelder als eine “Cotype’” bezettelt wurde. Der Fundortzettel dieses Exemplares lautet: “Hungaria, Vörs, Com. Somogy, 15.—30.VII.1932, E. Pfeiffer’ (Genitalpräparat No. M. 1053); die Vorderflügellänge ist 6,5 mm. Wie eine Genitaluntersuchung zeigt, ist dieser Lectotypus mit Ewcosma danicana Schantz kon- spezifisch. Im Einklang mit dem neuen International Code (1961, 1964: Artikel 15 und 45e) hat der Name balatonana eine Priorität vor danicana, obwohl er für die Bezeichnung einer Variation vorgeschlagen wurde. Dementsprechend muß danicana als ein jüngeres Synonym von balatonana zurücktreten. Der obenerwähnte Pseudotypus gehört zu Pelochrista (Pseudeucosma) modicana (Z.). Weitere aus der Sammlung Osthelder stammende, als balatonana bestimmte und mit danicana artlich zusammengehörige Exemplare (alle Männchen) sind: 1. Lochhauer Sandberg, Oberbayern, 19.VII.- 1938, am Licht (F. Daniel) (Genitalpräparat No. M. 1050); 2. Nyieb tor, Bator-Gebirge, West- ungarn, 9.VIII.1934 (F. Daniel) (Genitalpräparat No. M. 1051); 3. Vörs, Komitat Somogy, Ungarn, 9.V11.1931 (L. Osthelder). Zwei Männchen, die auch als balatonana bestimmt wurden, gehören zu Eucosma (Eucosma) fulvana (Stph.): Lochhauer Sandberg, Oberbayern, 3.VIII.1938, am Licht (F. Daniel) (Genitalpräparate No. M. 1049 und No. 877-Forster). Die sämtlichen Exemplare haben folgendes gemeinsam: alle sind nicht ganz frisch und gehören zu den Zwergen (Vorderflügellänge: 5—6,5 mm). 7. E. (E.) expallidana (Hw.). — Die von Osthelder (1937) als eine Form dieser Art beschrie- bene balatonana hat mit expallidana nichts zu tun und stellt eine andere, oben näher besprochene Art dar. 8. E. (E.) flavispecula Kuzn. — Kuznetzov (1964) verglich diese Art mit scorzonerana, aber m.E. steht sie eher der fu/vana näher und ist vielleicht ihre geographische Unterart. Aus dem Wiener Naturhistorischen Museum hatte ich folgende Exemplare, die ich als flavispecula identifiziere, zur Ansicht; ein Männchen aus Hansag, Burgenland, Österreich, 16.VII.1955 (F. Kasy); ein Männchen (Genitalpräparat No. 1344), Einrub, 25.VII.1921; ein Männchen von den Zitzmannsdorfer Wiesen, südlich von Weiden am See, Nordburgenland, 24.VI.1961 (F. Kasy); ein Männchen (Genitalpräparat No. 1372), Krs Balaton bei Vörs, Ungarn, 21.—29.V1.1936 (J. Klimesch); ein Weibchen (Genital- präparat No. 1379), Bozen, Südtirol, 24.VII.1913 (H. Zerny). Im A.MN.H. befindet sich noch ein weiteres Männchen (Genitalpräparat No. 758-Obr.), Illmitz, Burgenland, 11.VII.1954 (H. Reis- ser). Ähnliche Stücke sah ich auch aus der Südukraine (Nikolaew, Berdiansk). 20 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (246) 9. E. (E.) individiosana (Kenn.). — Holotypus, Männchen (Genitalpräparat No. B. 44), Lam- bèze, Algerien; Z.M.B. Die Abbildung dieses Exemplares bei Kennel (1921: t. 21 fig. 23) ist dunkler als das Original, das an Färbung anscheinend etwas eingebüßt hat. Die schwarzen Pünktchen der Spiegelstelle sind beim Holotypus auf den beiden Vorderflügeln etwas asymmetrisch geordnet und die Kennelsche Figur gibt deren Anordnung am linken Vorderfliigel wieder. Auf dem rechten Vorderfliigel stehen zwei von diesen Pünktchen in der Länge, nicht vertikal wie auf dem linken. . 10. E. (E.) victoriana (Kenn.). — Lectotypus, Männchen (Genitalpriparat No. B. 27), Sajan; Lectoallotypus, Weibchen (ohne Hinterleib), Sajan; beide im Z.M.B. Das Männchen ist etwas be- schidigt, aber die meisten Zeichnungselemente der Vorderfliigel sind gut erkennbar. Dieses unter- scheidet sich stark vom einzig bekannten Weibchen (Lectoallotypus) und die von Kennel (1919) veröffentlichte Abbildung des Männchens scheint nach einem anderen Exemplar angefertigt zu sein. Das Männchen gehört zweifellos zur Gattung Eucosma; das Weibchen, dessen Genitalien fehlen, macht für diese Gattung einen etwas fremden Eindruck und erinnert eher an eine Epiblema-Att. (247) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 21 GESAMTREGISTER ZU „DIE GATTUNGEN DER PALAEARKTISCHEN TORTRICIDAE” 1954-1968 zusammengestellt von B. J. LEMPKE Amsterdam Index der Familien, Gattungen und Untergattungen, mit deren Synonymen *) Ablabia 98: 159, 172, 173; 99: 120; 100: 325; MOA: 2375. 111 3 Acalla 98: 187, 189; 100: 333, 335 Acleris 97: 192, 206, 223; 98: 162, 179, 180, [187, 189, 190, 193, 194; 99: 128; (0023272336; 104:723257233, 234 Acroclita 102: 176; 107: 4, 5 Adoxophyes 97: 158, 160, 162, 164, 196, 198, [199, 201, 203; 98: 219; 100: 317, 335, [336; 104: 234 Aenectra 98: 195 Affa 111: 3 Agapeta 97: 189, 193, 199, 229; 98: 179, 187; [107: 17 Alceris 98: 190 Aleimma 98: 179, 180, 182, 187, 190; 99: 125; [100: 336; 104: 236 Allohermenias 107: 4; 110: 32, 33 Amelia 97: 189, 192, 193, 222; 98: 189 Amphisa 97: 219, 222, 227; 100: 335 Amphysa 97: 171, 219, 222; 98: 195 Anatropia 97: 225 Anchylopera 111: 1 Ancylis 104: 64; 107: 3, 4; 111: 3 Anisigona 97: 179 Anisotaenia 98: 152, 155, 159; 100: 334, 335 Anoplocnephasia 98: 159, 162 Anthichlidas 107: 3 Anticlea 102: 175; 111: 3 Antithesia 111: 3 Aphania 98: 189 Aphelia 97: 157, 160, 162, 163, 182, 185, 189, [191, 192, 193, 195; 98: 172, 181, 210; [100: 312, 336; 104: 236; 111: 1 Arathrophora 104: 64, 65 Archips 97: 159, 161, 163, 164, 166, 171, 172001758 176%. 178% 179) 1820183: [189, 193, 198, 199, 207, 209, 211, 12147219793:21555 17271815, 189195; [203; 100: 309, 310, 335; 104: 234, 235 Archipsini 97: 150; 98: 200 Arctephora 97: 149 Argrotoxa 98: 190 Argyrolepia 97: 168, 229; 98: 159, 185; 107: 6 Argyroploce 102: 176; 104: 65; 110: 32—34 Argyroptera 98: 172 Argyrotaenia 97: 152, 158, 160, 163, 168, 169, [171; 98: 148; 100: 309, 336; 104: 236 Argyrotosa 97: 193, 229; 98: 185, 187 Argyrotoxa 97: 168, 229; 98: 184, 185, 187, [189, 190; 100: 335 Argyrotoza 97: 211, 229, 231; 98: 181, 185, [187, 190; 103: 111; 104: 56 Ariola 97: 199 Articolla 110: 31 Artiphanes 110: 31 Fußnote Aspidia 108: 366 Asp1la®1102:176%.203;7103 2211152127 Aspis 108: 366 Astatia 97: 224 Asthenia 97: 214; 102: 176, 200; 103: 112 Asthenoptycha 97: 225 Atteria 97: 151, 154 Bactra 98: 190; 101: 233; 107: 28; 111: 3 Balbis 101: 236, 237 Barbara 107: 3, 11, 33, 39, 40, 41, 44 Bardia 108: 371 Batodes 97: 158, 161, 162, 163, 164, 198, 209, 1224225227: 098 (226; 1002 319) [335, 336; 104: 234 Begunna 98:195 Blastesthia 107: 3, 11, 36, 37, 38, 44, 47; [108: 371 Botropteryx 107: 17 Brachycnephasia 98: 162 Cacaecimorpha [Druckfehler!} 100: 312 Cacochroa 108: 366 Cacochroea 108: 366, 370, 371, 372, 375, 381 Gacoecia 97.1687 17415 1722 175717795 180; [182, 183, 185, 186, 193, 198, 199, [201, 214, 224; 100: 334, 335 *) Die fett gedruckten Zahlen geben die Seiten an, wo die betreffenden Namen behandelt werden. Vor den für die Wissenschaft neuen Namen steht ein *. 22 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 Cacoecimorpha 97: 159, 162, 163, 176, 180, [182, 224; 98: 209: 100: 335, 336; [104: 236 Callimosema 110: 66 Calosetia 111: 3 Capua 97: 159, 161, 162, 164, 174, 175, 180, MOR, DOS, 21102144 216) 2240225: [98: 195, 224; 100: 248, 249, 250 [318; 104: 234 Carpocampa 102: 176, 182 Carpocapsa 102: 175, 181, 203; 103: 111, 137, [139; 104: 56, 61, 64; 107: 29; [108: 366; 111: 3 Catamacta 97: 157, 158, 161, 162, 163, 174, [ 206; 98: 203; 100: 310, 335, 336 Catoptria 98: 159; 102: 176, 203; 103: 134, [137; 104: 56; 107: 17; 110: 66; 111: 3 Cerace 97: 152, 154, 155; 98: 199; 100: 334, [336 Ceraceopsis 97: 154 Cerasini 97: 150; 98: 198 Cerata 102: 176 Cheimaphasia 98: 167 Cheimatophila 98: 166, 173; 100: 336 Cheimonophila 98: 167 Choristoneura 97: 152, 158, 159, 162, 163, [171, 172, 182, 185, 186; 98: 202; [100: 310, 311, 335, 336; 104: 234, [235, 236, 237 Chresmarcha 97: 158 Chrosis 98: 185; 100: 334, 336 Cirriphora 101: 237; 104: 59, 60, 63, 68, 69, [70, 239 Clavigesta 107: 3, 11, 45, 46, 47; 111: 5 Clepsis 97: 151, 157, 158, 159, 160, 162, 163, 164, 182) 195) 193, 195, 196, 201, 206, [211, 214, 223; 98: 213; 100: 314, 335, [336; 104: 234, 235, 236, 237 Cnephasia 97: 168, 183, 193, 207, 209, 220, [227; 98: 149, 150, 151, 155, 156, [158, 159, 161, 162, 163, 164, 169, 170, 172, 17306175) 176, 177, 178, [190; 99: 109; 100: 320, 334, 335, 336; 1104: 237 107: 75429136 Cnephasiella 98: 148, 149, 151, 161, 162, 163; [99: 117; 100: 324, 336; 104: 237 Cnephasiini 97: 150; 98: 147; 99: 107 Coccyx 97: 186, 201; 98: 175; 101: 238; [102: 175, 200, 203; 103: 111; 107: 6, [11, 29, 36; 111: 3 Cochylis 97: 189, 229; 107: 17; 110: 66; [111: 3 Collicularia 101: 237; 103: 134, 135, 136; [104: 68, 69, 238 Commophila 97: 229; 103: 111 Conchylis 98: 156; 111: 3 Coptoloma 102: 176, 203 Cornicacoecia 97: 152, 159, 162, 163, 171, 7232708: 203; 100: 335; 104: 235 Corticivora 104: 63 Crambus 111: 3 Crobilophora 102: 176 Crobylophora 102: 176, 183 Crocidosema 101: 233; 107: 3, 4, 5 (248) Croesia 97: 229, 231; 98: 179, 187, 189, 190; [99: 127; 100: 327, 335, 336; 104: 234, [236 Crucimetra 107: 4 Cryptophlebia 101: 238; 104: 64, 65, 66, 67, 168, 70 Cydia 101: 238; 102: 175, 200, 203; 103: 112, [140; 111: 1, 3 Diamphidia 101: 236, 237; 104: 51, 52, 67, 69, [70, 239 Dichelia 97: 171, 180, 185, 193, 198, 205, 211, [220, 227; 98: 195 Dichrorampa 102: 176 Dichrorampha 101: 236, 237, 238, 239, 241, [242, 243, 244 (subgenus), 245 (sub- [genus); 102: 203; 103: 134; 104: 67, [69, 239; 107: 17; 110: 13—19 Dichroramphodes 101: 238, 242, 244, 254; [104: 240 Dicrorampha 101: 238 Dictyopteryx 97: 193, 211; 98: 179 Diurnea 98: 165, 167 Ditula 97: 224 Djakonovia 97: 189, 192, 195; 98: 212; [100: 314; 104: 236, 237 Dolophoca 98: 159, 162, 172; 100: 334, 335, [336; 104: 237 Dolophora 98: 159, 162 Doloploca 98: 149, 150, 159, 169, 173, 175; [99: 124; 100: 326 Drachmobola 101: 234 Eana 98: 149, 150, 151, 159, 161, 169, 170, [172, 173; 99: 120, 121; 100: 325, 336; [104: 237 Ecdytolopha 101: 238; 104: 66 Eclectis 98: 189 Enarmonia 101: 237, 238; 102: 175, 200, 203; [103: 134, 137, 140; 104: 56, 61, 62, 1163, 64, 68, 169; 7238511072930; [111: 3 Encelis 102: 175 Endopisa 101: 238; 102: 176, 203; 103: 112 Endopsia 102: 176 Endothenia 110: 32, 34 Endotritis 97: 160, 163 Enoditis 97: 213, 220, 222; 98: 225; 100: 335, [336; 104: 234 Enyphantes 98: 167 Epagoge 97: 159, 160, 162, 164, 171, 180, 189, [193, 205, 206, 209, 211, 213, 214, 216, [220; 98: 195, 223; 100: 318, 334, 336; [104: 234 Ephippiphora 98: 152; 101: 238; 102: 175, {203; 103: 111, 127, 140; 108: 366: [110: 14 Epibactra 107: 4, 20, 27, 28, 29 Epiblema 102: 203; 103: 111, 136; 108: 365, 1366, 367, 369, 370, 371, 372, 373. {374 375, 3772 A0 CES Epichorista 97: 189 Epigraphia 98: 173 Epinotia 102: 175, 203; 103: 111; 104: 232; [107: 3, 4, 11; 108: 366, 385; 111: 3 Episimus 101: 232, 233 (249) Ergasia 97: 216; 98: 179, 180, 182, 184, 185; [99: 126; 100: 326, 335, 336; 104: 236 Ericia 97: 179 Ericiana 97: 180 Eriopsela 108: 366 Erminia 102: 175 Ernarmonia 102: 175, 200, 203; 103: 134, 140; LOL ESC ACL NCA 111 1 Eucelis 102: 175; 103: 131, 140; 110: 31, 32, 33 Euchromia 107: 36; 108: 366 Eucosma 102: 176; 103: 112, 131; 104: 56; [107: 3, 17, 24, 29; 108: 365, 366, 13704 3719:3734374:5110% 24% 315032! [33, 34, 66; 111: 1, 2, 3, 4, 5, 6 Eucosmini 101: 234; 107: 2 Eucosmoides 111: 3 Eucosmomorpha 101: 232, 237; 103: 133; [104: 53, 55, 56, 57, 67, 68, 69, 70, [238 Eudemis 97: 168; 98: 152, 155, 159, 169; [102: 200; 103: 111 Euledereria 98: 149, 150, 169, 175; 99: 124; [100: 336; 104: 237 Eulederia 98: 175; 101: 233 Eulia 97: 168, 189, 193, 201, 204, 207, 209, {220; 98: 148, 149, 150, 151, 152, 156, [157, 158, 169, 177, 184; 99: 109; [100: 334, 336; 104: 236 Eupoecilia 111: 1 Eurydoxa 97: 154, 155; 98: 199 (subgenus) Euryptychia 108: 366 Euspila 102: 203, 207, 211; 110: 24—25, [238, 239 Eutrachia 97: 193; 98: 159, 165, 167, 172, [173, 187, 189 Evertia 107: 7 Evetria 107: 6, 10, 11, 29, 33, 36, 39, 44, 45 Exapate 98: 149, 150, 151, 167, 169, 170; [99: 119; 100: 325, 334, 335, 336; [104: 238 Exentera 111: 3 Exenterella 111: 3 Foveifera 107: 3, 11, 20 Glyphiptera 97: 193; 98: 190 Glyphipteryx 100: 318 Glyphisia 98: 190 Gnorismoneura 97: 158, 159, 160, 162, 164, [216, 228; 98: 227 Godana 97: 179 Goditha 101: 236 Grapholia 110: 66 Grapholita 101: 238; 102: 176, 182, 203; O3 ie 1362) 1047 56111: 15 3 Grapholitha 98: 169; 101: 236, 238; 102: 175, [183, 200, 203—215, 207 (subgenus); 103: 111, 1140125 012750129) 133, [134, 137, 140; 104: 53, 56, 59, 61, {63, 68, 69, 70, 238, 239; 107: 7, 17, [24, 27; 108: 366; 110: 18, 20, 21—26, [29, 66; 111: 3 Gravitarmata 107: 3, 11, 42, 43, 44 Gretchena 107: 3, 4 Griselda 107: 3, 4, 5 Gwendolina 101: 233; 107: 3, 4, 5 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 23 Gynoxypteron 98: 164, 165 Gypsonomoides 107: 3 Halonota 101: 238; 103: 112; 108: 366 Harmologa 97: 171, 172 Hastula 97: 160, 162, 164, 213, 220, 225, 227; [98: 226; 100: 319, 336; 104: 234, 237 Hedulia 101: 235 Hedya 108: 366 Hememe 101: 238 Hemene 101: 238 Hemerosia 102: 176; 103: 112 Hemimene 101: 238; 102: 175, 200, 203; [103: 111, 114; 110: 14, 15, 16, 17, 18; [111: 18 Hendecaneura 107: 3, 20 Hermenias 107: 4 Herpystis 107: 3, 4 Heterognomon 97: 182, 189, 193, 204, 227, [229; 98: 181, 184, 185, 187, 190 Heusimene 101: 238; 102: 175, 203; 103: 37, [111 Homona 97: 157, 159, 161, 162, 163, 176, 179, [180, 182, 213, 218; 98: 208; 100: 312, [335, 336; 104: 234 Hypostephanuncia 98: 172 Hysterosia 98: 187 Ioplocama 111: 3 Isotrias 97: 231; 98: 148, 150, 151, 154, 155, 1156:2.99381075. 1005319953357 339: 1104: 238 Kenneliola 102: 176, 183 Kundria 107: 4, 5 Lasperesia 102: 176; 104: 239 Laspeyresia 101: 233, 236, 238; 102: 175—200, [200, 203103: 112 014001201211 [133, 134, 140, 143; 104: 56, 61, 68, [69, 70, 238, 239; 110: 20, 21, 29, 30, 32933236 Laspeyresiini 101: 234, 235; 104: 238; 110: 13 Laspeyria 102: 183 Lathronympha 100: 309; 101: 237; 103: 131, [136, 137, 138, 139; 104: 68, 69, 238; [110: 29, 30, 31 Leguminivora 103: 129, 130, 131; 104: 68, 69; [110: 29, 30, 31 Lemmatophila 98: 165, 167, 173 Leontochroma 97: 157, 158, 161, 163, 164, [216, 218, 227; 98: 225; 100: 334, 336 Leptogramma 98: 190 Lespeyresia 102: 176 Lipoptycha 101: 238, 242, 243, 245, 246, 252; [104: 240; 110: 13, 14, 16, 17 Lipoptychodes 101: 238 Lithographia 102: 176; 111: 3 Lithosia 98: 167 Lobesia 98: 155; 101: 233; 102: 176; 104: 63 Lopas 98: 189 Lophoderus 97: 168, 189, 201, 204, 209; [98: 156, 157, 177 Loxotaenia 97: 168, 171, 175, 182, 183, 185, [193, 204, 211, 222 Lozotaenia 97: 160, 162, 163, 164, 166, 171, 75 01820183 0185 189719359203; [204, 205; 98: 169, 184) 1874195, 221; [100: 317, 336; 101: 234; 104: 236 24 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 *Lozotaeniodes 97: 151, 158, 159, 160, 162, [164, 201, 203; 98: 221; 100: 336; [104: 236 Mastumuraeses 110: 31 (Druckfehler!) Matsumuraeses 102: 183; 103: 131, 132, 133; [104: 67, 68; 110: 31—34 Melanalopha 101: 236 Melanalophinae 97: 149 Meritastis 97: 160, 162, 164, 186, 188, 189, [191, 231; 98: 210; 100: 335, 336 Metasphaeroeca 103: 112 Neoclyptis 97: 206 Neohermenias 10: 32, nota Neosphaleroptera 98: 148, 150, 161, 169, 170; [99: 119; 100: 336; 104: 236, 237 Nephodesme 97: 207, 214; 98: 159, 169, 170, [172; 98: 190 Nikolaia 97: 149 Norma 107: 4, 5 Notocelia 102: 176; 108: 366, 368, nota, 369, [1371003721573 375,381: 111: 3 Oenectra 98: 195 Oenophthira 98: 195; 100: 336 Ofatulena 101: 237 Olethreutes 97: 168, 171, 175, 183, 19837152, 157. 15952169; 0101: [102: 175, 200, 203; 103: 111, [104: 55, 61; 107: 7, 29, 36; 108: [110: 32, 33, 34, 66; 111: 1 Olethreutinae 97: 149; 101: 229, 230, 231; 110222175; 1003-0110; 104105140233: WOT OSG SN OGS MA Olethreutini 101: 235 Olinda 98: 152; 104: 238 Olindia 98: 148, 150, 151, 152, 155; 99: 107; {100: 319, 335, 336; 103: 124; 107: 11 Onectra 98: 195 Opadia 102: 203 Oporinia 98: 165 Oporopsamma 98: 172 Orchemia 102: 176 Orthotaenia 97: 168; 98: 152; 107: 6, 36 Oxapate 98: 167 Oxigrapha 98: 189, 190 Oxygrapha 98: 190 Oxypate 98: 167 Oxypteron 97: 220; 98: 149, 150, 151, 161, [164, 165; 99: 117; 100: 334, 336; [104: 237, 238 Paedisca 97: 168, 224; 102: 175; 103: 112, [137; 108: 366; 110: 66; 111: 3 Palla 103: 111 Palpita 98: 172 Palpocrinia 98: 149, 150, 151, 163, 164; 199: 117... 118: 3, 511 Pamene 1025176; 103: 112! 125, 127; 10451, [59; 110: 26 Pammene 101: 236; 102: 175; 103: 111, 113, MiA 15% 116, 121508106 1127; [104: 51, 59, 68, 69, 70, 238, 239; [108: 366, 377; 110: 24, 26, 35, 36 Pandemia 97: 164, 175 Pandemis 97: 152, 158, 160, 161, 163, 164, [165, 166, 168, 172, 174, 175, 179, [182, 193, 198; 98: 200; 100: 309, 310, DIA: 238; 136; 366; (250) [334, 335, 336; 104: 235, 237; 107: 3 Panoplia 97: 199 *Paraclepsis 97: 159, 160, 162, 164, 209, 213; [98: 157, 223; 100: 318, 336; 104: 236 *Paralipoptycha 101: 241, 242, 244, 255; {104: 240; 110: 13 Paramesia 97: 158, 161, 162, 164, 205, 206, {211, 213; 98: 190, 222; 100: 318, 336 [104: 234 Parapammene 103: 114, 125, 126, 143; 104: [68, 69, 239; 110: 35, 36 *Parapandemis 97: 152, 158, 160, 161, 163, [166; 98: 202; 100: 309, 336; 104: 235 *Parasyndemis 97: 159, 161, 163, 176, 185, [186; 98: 209; 100: 335, 336; 104: 235 Pardia 108: 366, 373 Pelochrista 108: 365, 370, 373, 374; 110: 66, ORE M9 Pentacitrotus 97: 152; 98: 198; 100: 336 Penthina 98: 152, 156; 102: 175, 176; 111: 1 Peridaedala 107: 4 Peronea 97: 192, 206; 98: 189, 193, 194; 00: 535360 Petrova 107: 11, 29, 30, 32, 33, 36, 38, 44; [108: 371 Phalaena 97: 164, 175, 189, 193, 199, 205, [209, 211, 219, 222, 229; 98: 155, 157, [158, 165, 167, 170, 181, 185, 187, [189, 194; 101: 238; 102: 175, 200, 203;103: 111,7 136: 1047556; [107: 7, 29, 36; 110: 13; 111: 1 Phanerophlebia 104: 66 Phaneta 111: 3, 6, 11 Philalcea 103: 111; 111: 3 Philedone 97: 159, 160, 161, 164, 193, 209, [214, 218, 219, 222, 223, 224, 227; [98: 225; 100: 334, 336; 104: 234 Philedonides 97: 161, 163, 164, 214, 222, 223; [98: 225; 100: 319, 336; 104: 234 Phloiophila 98: 190 Phoxopteris 111: 3 Phtheochroa 111: 3 Phthoroblastis 101: 238; 102: 176; 103: 112, [127; 110: 26 Phylacophora 98: 190 Platypeplus 104: 65 Poecilochroma 111: 3 Poedisca 98: 159 Pogonozada 104: 65 Polyortha 98: 194 Propira 98: 157 *Propiromorpha 97: 231; 98: 148, 149, 151, [152, 156, 157; 99: 108; 100: 336; 104: 236 Proteopteryx 111: 3 Proteoteras 107: 3, 4 Psammozesta 98: 164, 165 Pseudamelia 97: 195, 196; 98: 213; 100: 315; [104: 236, 237 Pseudargyrotoza 97: 159, 160, 161, 164, 228, [231; 98: 189, 227; 100: 319, 335, 336; [104: 236 Pseudatteria 97: 151 Pseudeucosma 110: 66, 72, 76; 111: 19 (251) *Pseudeulia 97: 151, 159, 161, 163, 164, 207, [209, 213; 98: 222; 100: 336; 104: 236 Pseudexentera 107: 4 Pseudococcyx 107: 4, 11, 33, 34, 35, 36, 38, 44 Pseudogalleria 104: 65, 66 Pseudoips 97: 164 *Pseudophiaria 104: 54, 55 *Pseudophiaris 101: 232, 237; 103: 133; [104: 53, 54, 55, 58, 67, 69 Pseudotomia 101: 238; 102: 175, 200, 203; F103 DI 1101270136 Pseudotomoides 102: 183, 200, 202; 104: 67, [238 Ptycholoma 97: 158, 159, 162, 164, 186, 193, [199, 201, 203, 214; 98: 157, 220; (1005) 31574 8335001011 #234; 0104-0235 *Ptycholomoides 97: 159, 161, 164, 185, 186; [98: 209; 100: 335; 104: 235 Pygolopha 98: 156; 111: 3, 4, 5, 11 Pyralis 97: 164, 175, 189 ,193, 199, 204, 205, [209, 211, 219, 228; 98: 155, 172, 173, [179, 181, 187, 189, 195; 101: 238; 1510254175%.203;,,103:4111., 127,136; 1104: 61; 107: 7, 29; 108: 366; 111: 1 Pyrodes 103: 112 Pyrsarcha 97: 160, 162, 164, :216,,. 223, 224; [98: 226 Retinia 107: 7, 10, 11, 29, 33, 36, 39, 42, 45, [201 Rhacodia 98: 187, 189, 190, 193 Rhocodia 98: 190 Rhopobota 107: 3, 4, 5 Rhyacionia 98: 152, 157, 190; 103: 134; 107: 3, (ARION TO 11, 202952 326; 1447452111273, 5 Ricula 101: 237 Satronia 101: 235, 236 Sciaphila 97: 168, 171, 183, 185, 193, 206, [207, 209, 214, 227; 98: 155, 159, 162, 15163%..164, 169) 172 175, 17775 1905 [107: 7; 108: 366; 110: 66 Scinipher 98: 167 Sciniphes 98: 167 Selania 101: 2372 (238% 102: 1776, 185, 203; [103: 139, 141, 143; 104: 68, 69, 70, [238 Semasia 101: 238; 102: 175, 203; 103: 111, [127, 131, 134, 136, 140; 104: 56, 61; 110723722 2437 110:726% 31-34, 766; RUES Sereda 101: 237 Sericoris 97: 168, 171, 201; 101: 238; 107: 136; [108: 366; 110: 13 Siclobola 97: 157, 185, 193, 196, 198; [98: 214; 100: 315; 104: 235, 236, 237 Sonia 107: 4; 108: 373 Sparganothinae 97: 149; 98: 194; 99: 154; [100: 309; 101: 230; 104: 232 Sparganothis 97: 193, 222; 98: 180, 189, 194, [195, 196; 99: 154; 100: 333, 334, [336; 104: 234 Sparganothris 98: 195 Sparganythis 98: 195 Spatalistis 97: 216; 98: 179, 185, 187, 189; [99: 127; 100: 335, 336; 104: 236 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 25 Sphaeroeca 103: 112 Sphaleroptera 98: 159, 162, 175; 100: 334, 336 Spilonota 102: 175; 103: 111; 107: 3, 45; [108: 366 Steganoptycha 98: 159; 102: 203; 103: 112; [111: 3 Stictea 97: 193; 101: 238; 102: 200 Stigmonota 102: 176, 203; 103: 112, 127, 140 Strepsicrates 107: 3, 4 Strophedra 101: 236; 103: 114, 126, 127, 128; [104: 53, 68, 69, 239 Strophosoma 103: 112, 127 Subepiblema 107: 34, 35 Suleima 107: 4 Syndemis 97: 159, 162, 163, 171, 176, 183, [185, 205; 98: 155, 159, 163, 172, 209; {100: 312, 335, 336; 101: 234; 104: 235 Synochoneura 97: 220, 227; 98: 148, 149, 150, 115152. 99:%.1073 1007 1336; 101-2255 Syricoris 97: 171; 98: 181; 107: 7 Talponia 101: 236 Teleia 98: 189 Teras 97: 175, 179; 98: 177, 187, 195; 104: 65; HLTO-NCG MERS Teratodes 97: 214 Thiodia 102: 176; 107: 3, 4, 17, 18, 20, 22, {i26,927, 108:°365;) 110332533 "1015; [4, 5 Thiodiodes 107: 3, 4, 20, 24, 25, 26, 27 Thirates 102: 175; 108: 366 Tia 101: 233 Tinea 101: 238; 102: 175, 200, 203; 103: 111; 1104755: 1072 29, 367 108: 360; 1110: 6: Ui Tortricinae 97: 148, 150; 98: 198; 100: 309; [101: 230; 104: 232 Tortricini 97: 148, 150; 98: 124; 99: 125 Tortricodes 97: 220; 98: 149, 150, 161, 164, [165, 166, 173; 99: 118; 100: 334, 335, [336; 101: 233; 104: 237 Tortrix 97: 164, 166, 168, 171, 172, 175, 179, 1F182, 18351857189, 193771987199, [201, 204, 207, 209, 211, 214, 219, 12224. 224.227.229:798:2152, 155.156, [157, 158, 159, 162, 163, 164, 165, 11692 1702 172217338175, 1277, 179, [180, 181, 182, 184, 185, 187, 189, (195: 99: 1125; 100:733473357 336; [101: 238; 102: 175, 200, 203; LOS LI; 1127. 1363. 1047" 555 NC10236 "10787, [17, 29, 36; 108: 366; 110: 20, 26, 66; ante a Trachysmia 98: 149.151. 1560, 1572169 91775, [177, 178; 99: 124; 100: 336; 104: 236 Trycheris 102: 176 Xanthosetia 97: 189, 193; 98: 195 Zacoriscini 97: 150 Zeiraphera 98: 152, 169; 102: 175, 203; [103.: 111; 107: 4 Zelotherses 97: 185, 189, 191, 192 (subgen.); [98: 210; 100: 313, 334, 335, 336; [101: 234; 104: 234, 235, 236, 237 Zomaria 101: 233 26 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (252) Index der Arten, Unterarten, Varietiten, Aberrationen und deren Synonyme A abacana 111: 16 abascama 111: 16 abdallah 99: 109 abietana 98: 189, 190; 99: 118, 138, 149; [100: 333; 104: 233 abietella 102: 192 abildgaardana 99: 135 abiscsana 99: 154 abiskoana 97: 220, 222; 99: 154; 100: 333; [104: 234 abjectana (ab.) 99: 150; 104: 233 abrasana 99: 116, 117; 100: 322; 104: 237 absconditana 108: 380 absinthiana 111: 17 absynthiana 111: 17 accentana (f.) 108: 375, 384 acceptana 108: 380 accinctana 98: 223 acclivana 98: 216 acclivella 108: 375 accuminatana 101: 248 accuratana 98: 212; 100: 314; 104: 237 acerana 98: 206; 100: 309, 310 acervosana (ab.) 98: 222 achatana 108: 366, 383 acleris 97: 206 acuminatana 101: 248; 104: 240 adamana 99: 118 adenocarpi 102: 188, 198; 104: 238 adiunctana 98: 221 adjunctana 98: 221; 102: 192; 104: 238 adspersana 99: 131 adulterina 98: 157 adulterinana 99: 108; 104: 236 advena 97: 152, 154; 98: 195 adumbratana 98: 207 aemulana 111: 14, 15 aeneana 102: 188 aeneus 98: 198 aeologlypta 102: 192 aeratana (Dichrorampha) 101: 255, 256, 259, [260, 261; 104: 240; 110: 14 aeratana (Pseudargyrotoza) 98: 228 aeriferana 97: 185, 186; 98: 209 aerosana 98: 214; 104: 237 aerugana (ab.) 99: 150, 151; 104: 233 aestivana 97: 179, 180 aetnana 98: 222; 100: 318 affectana 111: 17 affinisana 98: 224 affinatana 99: 134; 104: 234 afflicta 111: 9 agathana (f.) 99: 109 agenjoi 98: 214 agilana 101: 249, 254; 104: 240; 110: 19 agillana 101: 254 aglaoxantha 99: 127 aglossana 98: 212; 104: 237 agnatana 111: 7 agnotana 103: 117, 123 agraphana (ab.) 99: 128 agrestana 110: 69, 70, 71, 72, 75, 82 agricolana 99: 123; 100: 326 agrioma 99: 152 agyrana 103: 125 alaicana (Dichrorampha) 101: 251, 258; [110: 17 alaicana (ssp.) (sedana) 99: 110; 100: 320, [324; 104: 237 alatavica 110: 17 albana (ab.) (hastiana) 99: 137 albana (f.) (nigromaculana) 111: 16 albana (ab.) (variegana) 99: 135 albarracina 111: 6, 19 albatana 99: 112; 100: 321 albersana 104: 55, 57, 58, 238 albicans (ab.) 99: 116 albicapitana (ab.) (cristana) 99: 143 albicapitana (ab.) (hastiana) 99: 137 albicostana (ab.) (apiciana) 99: 148; 104: 234 albicostana (ab.) (cristana) 99: 143; 104: 232 albicostana (ab.) (hastiana) 99: 137 albicostella 111: 17 albidana 98: 223 albidulana 111: 7, 19 albidulata 111: 7 albimacula 101: 254, 260 albimaculana (ab.) 99: 137 albionana 107: 13 albipunctana (ab.) 99: 143, 146; 104: 232 albiscapulana 99: 147, 233 albisecundella 102: 209 albisparsana (ab.) 99: 137 albistriana (ab.) (hastiana) 99: 137, 138; [104: 234 albistriana (ab.) (hippophaëana) 99: 142 albistrigana 99: 147 albociliana 97: 185, 189, 191, 192; 98: 212; [104: 234 albodelineata (ab.) 99: 142 alboflavana 99: 143 albohamulana 108: 380 albomaculana (ab.) 99: 114 albonigrana 99: 143 alborufiscostana (ab.) 99: 144 albosparsana (ab.) 99: 113 albostriana (ab.) 99: 137 albovittana (ab.) (cristana) 99: 144; 104: 232 albovittana (ab.) (hastiana) 99: 138 albrechtella 108: 375, 384 albuginana 103: 118, 123, 124, 125, 126; [104: 239 albulana 99: 107, 108 albuncana 111: 6 albuneana 111: 6, 19 alcmaeonis 98: 206 alexiana 98: 218 alfredana 98: 203 algerana (ab.) 99: 115 alhamana 99: 116; 100: 324 alienana (ssp.) 102: 186 alniana 99: 119 alpestrana 101: 250, 251; 110: 18 (253) alpestris (ab.) 99: 122 alpha (ab.) 98: 227 alpicola (ab.) 99: 120 alpicolana 98: 169, 175; 99: 120, 124; 104: 237 alpicolana (f.) (variegana) 99: 135 alpigenana 101: 252, 256, 258; 104: 240; [110: 14, 16, 18, 19 alpinana 99: 132; 101: 245, 246, 254, 256; [104: 240 alsaticana (f.) 108: 378 alstroemeriana 99: 119 altaica (ssp.?) 98: 210 alternana 99: 113 alternella 99: 113, 118; 100: 322, 323 alticolana 99: 112, 113, 114, 115; 104: 237 alticolana (ssp.?) (consortana) 101: 249 amandana 111: 11 amasiana (ssp.) 98: 223 ambrosiana 101: 246; 110: 15 ambustana 98: 209 amellana 111: 12 ameriana 98: 204, 205, 206 americana 98: 206 amerina 98: 206 amerinana 98: 204 amethystana 107: 44 amoenana 108: 382 amplana (Archips) 98: 211; 100: 313; [104: 236 amplana (Laspeyresia) 102: 183; 190; 104: 239 amplidorsana 102: 185, 195 amseli 98: 212 amurensis (ssp.) 99: 130 amygdalana 102: 213; 103: 117, 118, 123, 143; [104: 239 anatoliana 107: 21, 22, 23 anatolica 99: 114 ancillana 99: 109; 104: 236 andreana 99: 112; 100: 321 angulana 98: 214 angulatana 108: 379, 386 angustilineata 98: 224 angustiorana 97: 209, 224, 225, 227; 98: 226; [104: 234 aniara 97: 199 anserana (f.) 111: 15 anthera 98: 195 anticipans 110: 29 antigona 98: 218 antiphila 99: 123; 100: 326 apenninicola 99: 113; 100: 321 apheliana 110: 76, 83, 84 aphorista 99: 152 apicalis 99: 127 apicatana 102: 192; 110: 25 apiciana 99: 147; 100: 332; 104: 233, 234 apiciana (ab.) (cristana) 99: 144 apiciana (ab.) (hastiana) 99: 138, 139 apiciana (ab.) (hippophaëana) 99: 142 apicivitta (ab.) 99: 140 apiletica 98: 208 apocrypha 111: 16 appenicola 99: 113; 100: 321 appenninicola 99: 113 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 27 approximana 99: 132; 100: 327 apriliana 111: 3 aquana 108: 382 aquilana (ab.) 99: 138, 140, 141; 104: 233 aquilina (ab.) 99: 146 arabescana 110: 68, 80, 86, 87, 88 arabica 102: 178, 185; 110: 20 archimedes (ssp.) 98: 199 arcigera 102: 210 arcticana 100: 311; 104: 235 arcticana (ab.) (hastiana) 99: 138 areolana 99: 107 argentana 98: 169, 172; 99: 120; 104: 237 argentana (ab.) (cristana) 99: 144 argentana (ab.) (hastiana) 99: 138 argentana (ab.) (variegana) 99: 135 argenteostrigana (ab.) (cristana) 99: 144 argenteostrigana (ab.) (hastiana) 99: 138 argentimicana (ab.) 103: 117, 123 argillaceana 98: 221 argutana 108: 384 argytana 102; 202,212; 1032 111, 117,118, [123; 104: 239; 108: 376 aristidana 110: 80, 87, 88 arquatana 98: 222; 104: 236 artemisiana 101: 249 articana (ab.) 99: 138 artificana 98: 223; 104: 234 ascoldana 99: 128; 104: 236 asiatica (ssp.) 98: 204; 100: 311 asinana 97: 207, 209; 98: 222; 99: 113; [104: 236 asperana 98: 189 asperana (ab.) (literana) 99: 150, 151 asperana (ab.) (variegana) 99: 135; 104: 233 aspersalis 99: 131 aspersana 99: 131; 100: 327; 104: 234 aspersana (ab.) (variegana) 99: 135 asphodilana 98: 213 aspiaiscana [recte: aspidiscana] 101: 255 aspidana 111: 14 aspidiscana (Dichrorampha) 101: 255 aspidiscana (Eucosma) 111: 7, 14 aspidiscana (Laspeyresia) 102: 190, 199 aspodilana 98: 213 asscelana 102: 184 asseclana 99: 114, 115; 100: 323; 102: 184; [108: 377, 385, 386 assinana 99: 113 assumptana 101: 236 astragalana 111: 17 atelosticta 111: 10 atlantis 99: 114 atra (ab.) 99: 138 atrana (ab.) 99: 144 atromargana 103: 117, 123 atropurpurana 101: 249; 103: 120, 124 atrosignana (ab.) 99: 148 attaliana (ab.) 99: 144 audouinana 99: 127 auduinana 99: 127 augustiorana 98: 226 aurana 100: 314; 102: 175, 176, 181, 187, 198; [103: 120; 104: 239 28 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 aurantania 103: 119 aurantiacum 97: 157, 216, 218 aurantiana 102: 193; 103: 118, 143; [104: 239 aurantiana (ab.) (aurana) 102: 187; 104: 239 aurantiradix 111: 8 aureolana 102: 212; 104: 239 aurichalcana 98: 180, 182; 99: 126; 104: 236 aurifasciana (ab.) 99: 108 auristellana (ab.) 99: 126 auroscriptana 102: 211, 215 austriacana (ssp.) 98: 214; 100: 315 autolitha 108: 381 autumnana 104: 233 autumnana (ab.) (apiciana) 99: 148 autumnana (ab.) (hastiana) 99: 138, 140, 141; [104: 233 autumnana (ab.) (scabrana) 99: 143 avellana 98: 200, 207, 214 azukivora 110: 33 B bactrana 111: 17 bacurana 99: 129 balatonana 111: 9, 19 balcanica 98: 218 baligrodana 108: 376 balticana (f.?) 101: 249 balticola (ab.) 98: 200 banghaasi 108: 375, 384 baracola 99: 152 barbarana 99: 117; 104: 237 basalticola (ab.) 99: 149; 104: 233 basilinea (ab.) 99: 139 bathyglypta 98: 207 bellana (ab.) 99: 122; 104: 237 bentleyana (ab.) 99: 144 bergmanniana 98: 187, 189, 190; 99: 127; [104: 236 bergstraesserella 100: 318 berolinensis 108: 380 besseri 98: 203 betulana 98: 204, 215; 100: 311, 316 betulifoliana 98: 216 bicoloria (ab.) 99: 138 bifasciana 97: 216; 98: 186, 189; 99: 127; [104: 236 bifidana (ab.) (ferrugana) 99: 133; 100: 329; [104: 234 bifidana (tripunctana) 99: 132 biformana 99: 120; 100: 325 bigeminata 102: 192 bimaculella 102: 187 bimaculosana 108: 380 binigrata 98: 206 bipartana 110: 23 bipartitana 102: 206, 211, 214 bipunctella 111: 3 birmensis (ssp.) 98: 196 biruptana 99: 121 biserialis 102: 209, 213 bistriana (apiciana) 99: 147 bistriana (ab.) (hippophaëana) 99: 142 bittana 110: 15 (254) bivittana (ab.) 99: 138 bizensis (ssp.) 99: 111 blackmoreana 102: 185, 195 blandana 99: 135 blandiana 99: 135 blasiana 101: 251; 110: 14 blepharana 101: 255 bleszynskii 99: 115 bleuseana 110: 68, 73 blockiana 103: 116, 122; 104: 239; [110: 26, 27, 28, 29 blockiella 110: 26, 27 bogodiana 99: 116; 100: 324 bollingerana (ab.) 99: 138; 104: 232, 233 borana 99: 135 boreana 99: 120 bornigiana 111: 10 borreoni 99: 120 boscana 99: 134, 136; 100: 330; 104: 233 bouliana 107: 12 bracatana 98: 218; 104: 235 brachiana 104: 234 brachiana (ab.) (ferrugana) 99: 133; [100: 329; 104: 234 brachiana (tripunctana) 99: 132 brachysticta 111: 11 brachytoma 98: 206 branderiana 98: 162, 204, 205, 206; 99: 115; [102: 199 breviornatana (ab.) 99: 154 breviplicana 98: 205; 100: 311 brevivittana (ab.) 99: 138 brigittana 111: 12 brisiacana (ab.) 99: 142 brunnea (ab.) (cristana) 99: 144 brunnea (ab.) (hastiana) 99: 138, 140 brunnea (ab.) (hippophaëana) 99: 142 brunneana (ab.) (comariana) 99: 129 brunneana (ab.) (cristana) 99: 144; 104: 232 brunneana (ab.) (hastiana) 99: 138 brunneana (ab.) (literana) 99: 151 brunneana (ab.) (mixtana) 99: 146 brunneana (ab.) (umbrana) 99: 137 brunneana (ab.) (viburniana) 98: 210; [100: 313 brunneastrigana (ab.) 99: 152 brunneoradiana (ab.) 99: 138 brunneostriana (ab.) 99: 138 brunnescens (ab.) (hastiana) 99: 138 brunnescens (ab.) (hippophaëana) 99: 142 brunnichiana 101: 248 brunnichianum 102: 190 brunnistriana (ab.) 99: 142 buddhana 110: 76, 84 bugnionana 101: 238, 252; 104: 240; [110: 18, 19 buoliana 107: 6, 8, 10, 11, 12, 14 buraetica 99: 124; 100: 326; 104: 237 burgasensis 98: 218 burgasiensis 98: 218 burgisana 98: 218 burgnionana 101: 252 buringerana (ab.) 99: 138, 139 buvati (ab.) 99: 123 (255) byringerana (ab.) 99: 138, 141; 104: 233 byringiana 99: 131 C cacaleana 101: 242, 253; 104: 246 cacuminana 101: 243, 251 caeca (ab.) (rigana) 99: 125 caecana 102: 211, 215; 104: 238 caecimaculana 110: 67, 70, 71, 72, 76, 79, 81 caecimaculanum 110: 76 caecimaculata 110: 76; 111: 14 caementana 110: 80 caeruleoatrana (ab.) 99: 135 caerulescens 99: 152; 104: 232 caesareana 98: 217 caledoniana 98: 193; 99: 130; 104: 234 caliacrana 111: 16 calidoniana 99: 130 caliginosana 101: 248, 249 callimachana 99: 111; 100: 320 callisphena 102: 192; 110: 24 calvicaput 98: 224; 100: 318 campoliliana 111: 16 campositella 102: 211 cana 108: 381; 111: 3, 7, 9 canariensis 98: 215; 104: 236 cancellatana 101: 250, 257; 104: 240 candana 99: 152 candidana (ab.) 99: 121; 100: 325; 104: 237 candidulana 111: 16 canescana 99: 120; 100: 320, 325; 104: 237 canescana (ab.) (literana) 99: 149; 104: 233 canescens 99: 121 cantiana 99: 120 capillana 99: 111 capillitana 101: 247 capillitiana 101: 247 capitinivana (f.) 111: 16 cappana 98: 201 capparidana 102: 179, 186, 197; 104: 238; [110: 36 caprana 98: 218 caprimulgana 98: 203 caprionica (ab.) 99: 112 capsigerana 97: 178; 98: 203; 100: 310 capsigeranus 104: 235 capuana 99: 144 capucina (ab.) 99: 144 capuzina (ab.) 99: 144 *caradjai (ssp.) (Eucosma) 111: 10 caradjana (Aphelia) 98: 210 caradjana (Laspeyresia) 102: 192 caradjana (Thiodia) 107: 21, 22, 23, 24; [108: 365 carduana 111: 3, 7, 8 carinthiaca 99: 108 cariosana 103: 142 carpiniana 98: 201 carpophaga 104: 64, 67 caryocryptis 99: 125 castaneana (mixtana) 99: 146; 104: 233 castaneana (ab.) (schalleriana) 99: 134; 104: 233 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 29 castiliana (ssp.) 99: 108; 100: 319; 104: 238 cataclystiana 107: 4 cataxesta 98: 201 catharaspis 111: 11 catoptrana 111: 14 caucasica 101: 250 caudana 99: 151, 152 caudana (ab.) (emargana) 99: 152; 104: 232 caudidulana 111: 15 celsana 98: 216; 100: 316 cembrella 107: 13 centralasiae (ssp.) 102: 184 centrana 99: 132; 100: 327 centrovittana (ab.) (hastiana) 99: 139; [104: 233 centrovittana (ab.) (hippophaëana) 99: 142 centrovittata (ab.) 99: 139 cerasana 102: 208 cerasana (ab.) (ribeana) 98: 200; [100: 309, 312; 104: 235 cerasema (ab.) 98: 200 cerasivora 102: 192 ceretana 98: 223 cerusana 99: 136 cervana 111: 8, 14 cervinana 102: 192 cesareana 98: 217; 104: 235 cetratana 111: 13 chanana 110: 79 chantana (ab.) 99: 144 chapmani 99: 118 characterana 98: 205; 99: 124; 100: 326; [104: 237 charlottana (ab.) 99: 144 chavanneana 101: 251, 252; 110: 18, 19 chelias 110: 24 chlorograpta 98: 201 chlorotypa 98: 206 chondrillana 97: 152, 166, 168; 98: 202; [100: 309; 104: 235 chrétieni 108: 379 chrétieni (ssp.) *98: 212 ;100: 313 christophana 99: 127; 103: 119, 124; [104: 236, 239 christophi (ssp.) *98: 211; 100: 313 christophi (ssp.) (paleana) *98: 211; [100: 313; 104: 236 chrysantheana 98: 161; 99: 113; [100: 322, 323; 104: 237 chrysanthemana 99: 113, 116 chrysitana 98: 211, 217 ciliana 99: 131 cinareana 99: 109; 100: 320; 104: 237 cinctana 97: 209, 211, 213; 98: 157, 223; [100: 318; 104: 236 cinerana 99: 121, 154; 100: 333; 101: 246 cinerascens 101: 249, 257 cinerea (ab.) 99: 136 cinereana 97: 222; 99: 109, 154; 100: 333 cinereanum 99: 129 cinerosana 101: 247, 257; 103: 136; 104: 240; [110: 18 cingulana 99: 108 cinnamomea 98: 205 30 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 cinnamomeana 97: 166; 98: 200; [100: 316, 317; 104: 235 circiana (f.) 108: 378 circulana 111: 1, 2, 5 circumclusana 98: 220 circumflexana 108: 375 circumfluxana 108: 384 cirrana (ab.) 99: 135; 104: 233 cirsiana (f.) 108: 378 citrana 99: 111; 107: 17, 18, 20, 21, 22, 23, 24 clanculana 103: 117, 123; 104: 238 clara (ssp.) 98: 196 clarana 99: 122 clarescens 111: 6 clarkiana (ab.) 99: 145 clavigerana 102: 192; 108: 375, 384 clercana 99: 121 cleuana (ab.) 99: 115 cnicicolana 108: 379 coagulana 111: 7 cocciferana 103: 118, 124 coecana 102: 211 coecimaculana 110: 76 coeruleana (ab.) 99: 125 coeruleosparsa 102: 208 coeruleostriana 111: 11 coffearia 98: 208 cognatana 98: 202; 102: 189, 198; 104: 238 colfaxiana 107: 39, 40 colossa (ssp.) 99: 120 colquhounana (f.) 99: 122, 123; 100: 325; [104: 237 colquouhana 99: 123 comariana 98: 193; 99: 129; 104: 234 combustana 104: 232 combustana (ab.) (cristana) 99: 145 combustana (ab.) (hastiana) 99: 139; [104: 233 commodestana 110: 74 communana 98: 161; 99: 112, 115, 117; [100: 321; 104: 237 compacta (ab.) 98: 206 comparana (= tripunctana) 99: 132 comparana (ab.) (comariana) 99: 129 comparana (ab.) (latifasciana) 99: 129; [104: 234 composana 102: 211 compositella 102: 203, 207, 211; 104: 238 compressana 98: 221 comstockiana 107: 30 conchyloides 99: 128 conciliata 110: 24 concinnana 102: 210; 107: 12 confidana 110: 72, 82 confixana 99: 140; 100: 330 confixana (ab.) (abietana) 99: 149; 104: 233 confluens (ab.) (crataegana) *98: 205; [100: 311; 104: 235 confluens (f.) (foenella) 108: 375 confluens (ab.) (virgaureana) 99: 115 confluentana (ab.) 99: 112 conformana 111: 6, 18 confusa (ab.) 99: 139 confusana 108: 377 (256) congelatella 98: 167, 169, 170; 99: 119; [104: 238 congeminata *110: 80, 86 congenerana 98: 204 congruana 98: 220 congruentana 98: 214; 104: 235 coniana 101: 250 conicolana 102: 179, 181, 187; 104: 238 coniferana 102: 179, 182, 189, 198; 104: 238 conjugana 103: 119 conjunctana 102: 184, 195 conradii 99: 119 consequana 102: 184 consimilana 98: 214, 215 consimilana (ab.) 99: 144; 104: 232 consortana 101: 249, 253, 257; 104: 240; [110: 17 conspectana 102: 210 conspersana 99: 113, 116, 122, 132; 100: 322; [104: 237 constanti 98: 227; 104: 234 contaminana 99: 131; 100: 327 contemptrix 98: 206 conterminana 111: 14 continentana 98: 212; 100: 314 contrasignata 108: 380 conwagana 97: 228, 231; 98: 227; 100: 319; [104: 236 conwayana 98: 227; 100: 319 coprana (ab.) 99: 154; 100: 333; 104: 233 cordulana 111: 12 coriacana 98: 217; 104: 236 coriacanus 98: 217 corneliana 110: 76, 84 cornucopiae 102: 192; 104: 238 corollana 102: 175, 177, 179, 181, 187; [104: 238 coronana 99: 137, 138, 143; 104: 233 coronana (ab.) 99: 139, 141 coronillana 102: 212, 215; 104: 239 corticana 107: 33 coruscana 107: 13 corylana 97: 152, 165, 166; 98: 200, 205; [100: 317; 104: 235 cosmophorana 102: 181, 185, 196; 104: 238 costana 98: 215, 222; 100: 315 costialba (ab.) #99: 149; 100: 333 costimacula (ab.) 99: 141 costimaculana 99: 132, 133, 14; 100: 329; [104: 234 costimaculana (ab.) (lipsiana) 99: 147 costipunctana 103: 121, 123, 125; 108: 376 costistrigana (ab.) *99: 147; 100: 332 cottiana 99: 123; 100: 326; 104: 237 crassana 99: 147; 104: 234 crassana (ab.) (apiciana) 99: 148 crassifasciana 99: 112 crataegana 98: 201, 204, 205; 100: 310, 311 crataeganus 104: 235 crataegophila 103: 118, 123; 110: 29 craterana 97: 225 cretaceana (Cnephasia) 99: 113 cretaceana (Eucosma) 111: 15 cretana 98: 223; 108: 374, 378 (257) crispinana 98: 213; 100: 315 cristalana (ab.) 99: 144; 104: 232 cristana 98: 180, 189, 190; 99: 135, 139, 143, [144, 145, 146; 100: 332; 104: 232, 233 cristata 107: 33 critica 110: 24 criticana 98: 205 criticanus 104: 235 croatica 110: 78 croceana 98: 200, 214, 215, 221; [100: 316, 317, 318 crocopa 110: 30 crocopepla 99: 152 cruciana (ab.) 99: 107 crustulana 110: 74 cuencana 99: 108; 100: 319 culmana 98: 213; 111: 7 cumulana 110: 70; 111: 10 cupidiana 98: 221 cupidinana 98: 221; 100: 317, 318; 104: 236 cuprea (ab.) (aurana) *102: 187, 198 cuprea (ab.) (hastiana) 99: 139 cupressana 97: 201, 203; 98: 221; [103: 116, 122; 110: 26, 27, 28 cupressanus 104: 236 cupressivorana 99: 109, 112, 113, 121; [104: 237 curvifasciana (ab.) 99: 108; 100: 319 curtisana (ab.) 99: 194 cyaneana (ab.) 99: 135; 104: 233 cyanescana 99: 122 cyclidias 110: 31 cynosbana 108: 382, 383 cynosbatana 108: 382 cynosbatella 108: 366, 382 cytisana 103: 119, 124; 104: 239 D daedaleana 101: 247 dagestana 110: 80, 88 dahlbomiana 111: 14 dahliana 102: 191 daldorfana 103: 120 daldorfiana 103: 120 daleana 98: 211 dalecarliana 102: 185 dandana 102: 185 danicana 111: 9, 19 danilevsky *101: 254; 102: 216 danilevskyi 101: 260; 102: 216 dannehli 102: 182, 189, 198 darvaza (ssp.) 99: 120; 100: 325 dativa 107: 10, 12, 15 decaryi (ab.) 99: 115 decolorana (Cnephasia) 99: 113 decolorana (Eucosma) 111: 3, 10, 15 decolorana (ssp.) (succedana) 102: 184, 195 decorana 102: 184 decorata 102: 186 decoratana 102: 179, 186, 197 decosseana 99: 131, 132; 104: 233 decretana 97: 178; 98: 204; 100: 311, 316 decretanus 104: 235 definita (ssp.) 100: 324 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 31 definitana (f.) 110: 78 deflexana 102: 186 delibatana 98: 207 delicatana 99: 152; 104: 232 delineana 102: 211; 110: 24, 25, 26 delitana 99: 112 demissana 111: 17 denigratana 108: 376 densana 98: 205 densana (ab.) (apiciana) 99: 148 denticana 99: 154 derivana 99: 121, 122; 100: 325; 104: 237 dernina 110: 74 desfontainana (ab.) 99: 144, 146; 104: 232 detractana 98: 208 deyana 102: 188, 198 diana 102: 186 dicastica 110: 35 dichrorhamphana 103: 121; 104: 239 difficilana 102: 206, 212, 215; 104: 239 diffusana (= Grapholitha fissana) 102: 209 diffusana (Hastula) 98: 227; 104: 234 diffusana (ab.) (chrysantheana) 99: 114 dilutana (ab.) 99: 109 dimidiana (= Pammene rhediella) 103: 120 dimidiana (= Acleris rhombana) 99: 131 dimidiatana 102: 206, 207, 211, 214; [104: 239; 110: 23 directa 111: 18 directana (ab.) 99: 114 discedana 110: 14 discretana 100: 318; 102: 210; 104: 239 disjectana 111: 11 disparana 98: 207 disparanus 104: 235 disquei 110: 80, 86 disquéi (ab.) *99: 133; 100: 329 dissimilana 98: 204 dissonana 99: 135 distinctana (Cnephasia) 99: 116 distinctana (Dichrorampha) 101: 247, 249, [255; 104: 240 distorta 104: 65, 66 districta 98: 214 diticinctana 98: 228 diurneana 99: 122 diversana 97: 152, 171; 98: 202, 203; [100: 309, 310; 104: 235, 237 divisa 102: 192 divisana (=Grapholitha fissana) 102: 209 divisana (ab.) (apiciana) 99: 148 divisana (ab.) (hastiana) 99: 139, 141; [104: 233 dohrniana (ab.) 98: 213; 100: 315; 104: 236 dominicana 99: 123; 100: 326 donelana 98: 210 dorana 98: 216; 100: 316 dorsana 102: 191, 203, 204, 207, 209, 210, [211, 212, 214; 103: 117; 104: 239; 110; 21 dorsialba (ab.) *99: 149; 100: 333 dorsilunana 102: 186 dryochyta 98: 218 dryonephela 99: 109 duercki 110: 69, 70, 73, 75 32 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 dumeriliana 98: 226 dumetana 97: 166; 98: 201, 214; 104: 237 dumicolana 98: 214; 100: 315; 104: 235 dumonti (ssp.) 99: 122 duplana 107: 10, 13, 16, 38, 47 duplicana 102: 181, 185, 191, 200; 104: 239; [110: 21 duratella 99: 119; 100: 325; 104: 238 dürcki 110: 73 dzhungarica 101: 246 E eatoniana 98: 216; 100: 316; 104: 235, 236 ectypana (ab.) 99: 125; 104: 236 ecullyana 98: 172; 99: 123 edrisiana 110: 74 eductana 98: 207 effractana 99: 149 effractana (ab.) (emargana) 99: 152 effusana 102: 183; 108: 375 eidmanni 101: 242, 256, 261 elegantana 102: 212; 110: 70, 73, 82, 83 elevana 99: 142 elongata 111: 13 elsana 98: 216 elutana 110: 32, 33, 34 emargana 98: 180, 189, 190, 193; 99: 151, 152; [104: 232 emptycta 98: 201; 100: 309 endrosias 102: 211; 110: 26 engadinensis 103: 121 enitescens 99: 153 epanthista 102: 192 ephedrana 111: 18 ephippana 99: 143; 103: 118, 124 epicharana 101: 252 epicyrta 98: 206 epidola 110: 32 erebana (ab.) 99: 149 erebantra 111: 11 erebina (ab.) 98: 228 erectana 102: 212 eremica 99: 118 eremicum 99: 118 ericicolana 102: 189 erschoffi 98: 220; 104: 235 erutana 99: 129; 104: 234 erutana (ab.) (schalleriana) 99: 134; [104: 233, 234 erythrana (ab.) 98: 200 esmodes 111: 17 eulogiana 97: 168; 98: 202; 100: 309 eurychorana 101: 247; 110: 18 euryomis 98: 219 eutaeniana (ab.) 99: 147; 104: 233 euxina 97: 189, 192, 195; 98: 212; 100: 314 evanidana 98: 207 evanidanus 104: 235 eversmanni 110: 80, 87, 88 evisa (ab.) 99: 122 exalbescens 98: 224 excavana 104: 232 excavana (ab.) (emargana) 99: 152 exigua 99: 118 (258) exiguana 99: 118 exiguanum 99: 118 exiguanus 104: 237 eximia 101: 254; 110: 17, 18 eximiana 99: 138 exosticha 102: 193; 110: 23, 24 expallidana 111: 9, 19 expressa 99: 130 expressana 108: 380 exquisitana 102: 190; 104: 238 exsucana 99: 153; 104: 234 externana 98: 215 extinctana 110: 36 exuberans 99: 127 exulis 97: 216, 228 F faaborgensis (ab.) 99: 129 facetana 98: 162; 99: 110; 104: 237 fackiana 101: 249 faeneana 108: 375 fagana 99: 130 fagi (f.) 98: 221; 100: 318 fagiglandana 102: 181, 183, 190, 199; [104: 239 falcana 103: 133, 134; 110: 32, 33, 34 fallaciana 98: 216 falsana (ab.) 99: 134; 104: 233 famula (ab.) 99: 134 farfarae 108: 374, 376, 385 fasciana 98: 201, 202; 99: 107, 139; [103: 119, 124; 104: 239 fasciana (ab.) (comariana) 99: 129 fasciana (ab.) (emargana) 99: 152; 104: 232 fasciana (ab.) (fimbriana) *99: 147; 100: 332 fasciana (ab.) (hastiana) 99: 139 fasciata 98: 219; 100: 317; 104: 239 fasciculana 98: 208 favillaceana 97: 209, 214, 225; 98: 224; [99: 130; 100: 315, 318, 319, 320; 104: 234 ferrugana 99: 109, 131, 132, 133, 154; [100: 327, 328; 104: 234 ferruganum 99: 132 ferruginea (ab.) 99: 139 ferrumixtana 99: 132; 100: 328 ferugana 98: 211; 99: 109 fervidana 111: 10 fessana 108: 365 filana 102: 212 filipjevi (Acleris) *99: 147; 100: 332 filipjevi (ab.) (canescana) 99: 121 filipjevi (Dichrorampha) 101: 245 fimana 102: 209, 213; 104: 239; 1110221002924 fimbriana 99: 147; 100: 332; 103: 117; [104: 233 fimbriana (f.) (coffearia) 98: 208 fimbriana (ab.) (tripunctana) 99: 132 fingalana 99: 108 fiorii 108: 381 fisana 102: 210 fischeri (ssp.?) 98: 211; 100: 313 fissana 102: 209, 210; 104: 239 fissurana 99: 133; 100: 328, 329 (259) flammeana (ab.) 99: 144 flavana 98: 200, 210, 211, 215, 223; 99: 136 flavana (ab.) (cristana) 99: 145 flavana (ab.) (hastiana) 99: 139 flavana (ab.) (literana) 99: 151 flavana (ab.) (lorquiniana) 99: 148; 100: 332 flavicapitana 99: 139 flavicostana (ab.) 99: 139 flavidorsana 101: 245, 246, 256; 104: 240; [110: 15 flavidorsana (ab.) (hastiana) 99: 139 flavillaceana 98: 224 flavipunctana 102: 183 flavispecula 111: 9, 19 flavodorsana (ab.) 99: 139 flavostriana (ab.) 99: 144 flavovittana (ab.) 99: 145 flexana 103: 128, 129 floricolana 103: 119 fluidana 108: 377, 378, 386 fluxana 104: 235 fluxana (ab.) 98: 216; 100: 316 focnella 108: 375 foederatana 97: 176, 180; 98: 208; 104: 234 foenana 108: 375 foenella 108: 365, 367, 372, 375, 384 foenellum 108: 375 folskaleana 99: 128 fônella 108: 375 formosana (Enarmonia) 104: 61, 62, 63, 64, [238 formosana (Eucosma) 111: 3 formosana (Lozotaeniodes) 98: 221 formosana (ssp.) (stipatana) 98: 200 formosanus (Enarmonia) 104: 236 forscaeleana 99: 128 forscaleana 99: 127 forskaeleana 99: 127 forskähleana 99: 127 forskahliana 99: 127 forskaleana 98: 190; 99: 127 forskäleana 98: 189; 99: 127; 104: 236 forskaliana 99: 128 forskäliana 99: 127 forskoleana 99: 127 forskoliana 99: 127 forsskaleana 99: 128 forsterana 97: 201, 204, 211; 98: 221; [100: 318; 104: 236 forsteri 101: 250 fragariana 98: 166 fragosana 99: 110; 100: 320; 104: 237 fragrosana 99: 110, 117 fraudulentana 111: 13 fraxinana (ab.) 103: 118, 124; 104: 239 freii (ab.) 99: 114 freyeriana 111: 16 fuchsiana 108: 377 fucsana 98: 212; 104: 236 fuligana 99: 119 fuliginosana (nybomi) 98: 218 fuliginosana (ab.) (spectrana) 98: 215 fultana 110: 72 fulturata 99: 113; 100: 321 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 33 fuluana 102: 187 fulvana (Archips) 98: 204 fulvana (Eucosma) 111: 8, 9, 19 fulvana (ab.) (cristana) 99: 144 fulvana (ab.) (literana) 99: 151 fulvanum 111: 8 fulvella 102: 187 fulvifrontana 103: 120 fulvimitrana 107: 38 fulvipunctana 107: 38 fulvocristana (ab.) 99: 144; 104: 232 fulvoliterana (ab.) 99: 150; 104: 233 fulvomaculana (ab.) 99: 150; 104: 233 fulvomixtana (ab.) 99: 150, 151; 104: 233 fulvopunctana (ab.) 99: 144 fulvorittana (ab.) (cristana) 99: 144 fulvostriana (ab.) 99: 144 fulvostrigana 110: 78 fulvovittana (ab.) (cristana) 99: 144; [104: 232 fulvovittana (ab.) (hastiana) 99: 139 fumatana (ssp.?) 98: 211; 100: 313 fumiferana 97: 172 funebrana 102: 203, 208; 104: 238 furvana 99: 139 fusca 101: 252; 110: 17 fuscana (rosana) 98: 206 fuscana (ab.) (comariana) 99: 129 fuscana (ab.) (cristana) 99: 145 fuscana (ab.) (emargana) 99: 152; 104: 232 fuscana (ab.) (variegana) 99: 135 fuscociliana 98: 202 fuscocupreana 98: 220 fuscotogata 99: 128; 104: 234 fustulana 110: 74 fuvillaceana 99: 130 G gaditana 98: 225 gaditanus 104: 234 gailicolana 103: 118 galacteana (ab.) 99: 132 galactica 111: 16 galiana 98: 210; 100: 312 gallarum 103: 117, 123 gallicana 102: 176, 181, 182, 184, 186, 197; [104: 239 gallicana (ssp.) (genitalana) 99: 116 gallicolana 103: 118, 123, 124 gammana 108: 379 gavisana 97: 157, 174, 175 gelata 99: 119 gelatana 99: 119 gelatella 99: 119 geleana 98: 210 gemellana 101: 248; 104: 240 geminana 102: 185; 107: 12 geministriata 102: 193 gemmana 107: 12, 38 gemmatella 107: 11 gemmiferana 102: 204, 205, 207, 213; [104: 238 generosana 102: 182, 190, 199; 104: 238; [110: 20 34 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 geniculana 102: 210 genitalana 99: 116; 100: 322, 324; 104: 237 georgiella 99: 120 germana 103: 120, 124; 104: 239 germarana 99: 134; 101: 248; 102: 208 germmana 103: 120, 124 gerningana 97: 209, 214, 219; 98: 204, 205, [210, 225; 104: 234 giarabubensis 111: 18 gibbosana (ab.) 99: 145 gigantana (ssp.) (dorsana) 102: 210, 214 gigantana (= luticostana) 98: 203 gilvana 98: 202, 205, 206; 99: 132 gilviciliana 102: 178, 180, 181, 182, 185, 19G; [104: 238 glaphyrana (ab.) 102: 190 glareana 99: 119 glaucana 102: 180, 189, 198; 104: 239 glebana 103: 139; 111: 13 glycinivorella 103: 129, 130, 131, 139; {110: 29, 30 gnomana 97: 201, 205, 206, 211; 98: 215, [217, 222, 223; 100: 315, 316, 318; [104: 234 gnomana (ab.) (ferrugana) 99: 133 gnomana (ab.) (tripunctana) 99: 132 goliana 99: 120 gossipiana 98: 207 gouana 99: 120 govana 99: 120 gracilis (Dichrorampha) 101: 253; 110: 17, 18 gracilis (Eucosma) 111: 13 gradana 102: 193; 104: 63, 238 gradensis 111: 14 graeca (ssp.) 102: 191, 200; 104: 239 graecana (ssp.) 99: 111 graminana 101: 248 grammana 110: 76 granadana (ab.) 100: 316 granadanus (ab.) 98: 216 grandaevana 108: 370, 371, 372, 373, 375, 381 grandis 98: 162; 99: 110 graphana 108: 377 graphanum 108: 377 graphitana 98: 228 gratana 99: 111 gratiana 98: 223 gravana 98: 201; 100: 309 grisea (filipjevi) 99: 147; 100: 332 grisea (ab.) (hastiana) 99: 139, 141; [100: 330, 332 grisea (ab.) (hippophaëana) 99: 142 griseana (= canescana) 99: 121 griseana (ab.) (emargana) 99: 152 griseana (ab.) (hastiana) 99: 139 griseana (ab.) (lipsiana) 99: 147 griseana (ab.) (literana) 99: 150 griseana (ab.) (mixtana) 99: 146 griseana (ssp.?) (succedana) 102: 184, 195 griseicoma 98: 218 griseis (ab.) 99: 143; 100: 331 griseofasciana 99: 139 griseolana 110: 70, 78 griseovariegata 99: 141 (260) griseovitta (ab.) 99: 139 grisescana 99: 120 grossana 102: 183, 190, 199 grossulariana 98: 200 grotiana 97: 209, 211, 213, 214; 98: 223; [100: 318; 104: 234 gruneriana 101: 252, 255; 104: 240 griineriana 101: 252 grunertiana (ab.) 102: 189, 198; 104: 238 grunneriana 101: 252 guadarramana 101: 256 gueneana (Cnephasia) 98: 161; 99: 110, 111; [100: 320, 321; 101: 254; 104: 237 guenéeana (Dichrorampha) 101: 238, 242, [244, 254, 257; 104: 240 guentheri 111: 10 gueriniana 103: 142 gumpiana (ab.) 99: 145 gumpinana (ab.) 99: 145 gundiana 102: 211 guttana 98: 195 gypsatana 111: 6, 18 H haberhaueri 111: 8 halliana (ab.) 99: 130; 104: 233 hamatana 102: 209, 213; 104: 239 hapalactis 99: 153 harpeana 101: 250, 252; 104: 240; 110: 18, 19 hastana 107: 11, 20 hastiana 98: 189; 99: 107, 134, 136, 137, 138, [139, 140, 141, 142, 148; 100: 329, 330, [332; 107: 11; 104: 232, 233 haworthana 99: 130 hebenstreitella 98: 207; 100: 312 heegerana 101: 248, 258; 102: 187; 104: 240 hegeriana 101: 248 heinemanni *99: 110; 100: 320; 104: 237 helenica 99: 116; 100: 324 heliapoda 98: 227 hellenica 98: 161; 99: 116; 100: 324 helvolana 97: 185, 193, 195, 196; 98: 213; [100: 314, 315; 104: 236 hemicriptana 97: 225 hemidoxa 102: 193 heparana 97: 166; 98: 201, 205; 100: 309; [104: 235 hepatariana 110: 78 hepaticana 108: 377, 379, 386 hepaticanum 108: 379 heperana 98: 201 herbaecolor 98: 194 herbosana 101: 251 heringiana 111: 14 hermanniana 98: 204 hermineana 98: 209 herrichiana 107: 39, 41, 42 herrichiana (ab.) (fasciana) 103: 119, 124; [104: 239 heteroidana 97: 198, 199 heteropa 110: 24 heuseriana (ssp.?) 101: 250 hexaphora 103: 116, 121 hinnebergiana (Eucosma) 111: 10 (261) hinnebergiana (ab.) (Laspeyresia) 102: 185 hinnebergiana (f.) (Pelochrista) 110: 78 hippophaëana 99: 141, 142; 100: 330; [104: 233 hispanica 99: 113; 100: 321 hispidana 99: 130, 147; 104: 232 histrionana 97: 176, 185, 186; 98: 203, 209; [104: 235 hochenwartiana 108: 375 hoenwartiana 111: 9 hoffmanseggana 98: 227 hoffmanseggiana 98: 227 hofmannseggana 98: 227 hohenwarthana 111: 7, 8 hohenwarthiana 111: 8 hohenwartiana (Eucosma) 111: 7, 8 hohenwartiana (Lathronympha) 103: 139 hohenworthiana 111: 8 holmiana 98: 189; 99: 128; 104: 234 homotorna 110: 35 homsana 99: 118 homsanum 99: 118 honorana (ab.) 103: 119; 104: 239 hoplista 110: 34 horridana 99: 108, 124 hübneri (ab.) 99: 138 hübneriana 110: 66, 79 huebneriana 110: 79 humana 99: 125 humerana 99: 116 hybnerana 98: 205 hybridana 99: 107, 108, 119; 100: 320; [104: 238 hybridella 111: 8 hyemalis 99: 146 hyemana 99: 113, 118; 100: 332 hyemella 99: 119 hyerana 97: 220, 227; 98: 226; 100: 319; [104: 234, 237 hypericana 100: 309, 318; 103: 136, 139 hypericano 103: 139 hyperptycha 99: 125 hypsicrates 97: 216, 223, 224; 98: 226 hyringerana (ab.) 99: 138 I ianthinana 102: 207 iaquiniana 101: 245 iberica 98: 213; 100: 315 ibiceana 111: 9 icterana (ab.) (paleana) 98: 211; 100: 313; [104: 236 ictericana (ab.) (longana) 99: 111; 104: 237 ictericana (ab.) (paleana) 98: 211 idana 98: 216 idonatana 110: 68, 76, 84 ignavana 99: 118 ignavanus 104: 237 ignoratana 98: 227; 104: 234 ignotana 111: 11 ilicetana 102: 184 ilipulana 102: 184, 195 illipeda 104: 66, 67 illotana 98: 208; 100: 312; 104: 234 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 35 illustrana 98: 202 illutana 102: 189; 104: 238 illutanana 102: 189 imitator 98: 217; 100: 316 immaculana (= germana) 103: 120 immaculana (ab.) (murinana) 98: 203; [104: 235 immaculata (Dichrorampha) 101: 254, 260 immaculata (,,Hemimene”) 101: 260 immaculatana (ab.) 98: 221 impar 97: 220; 98: 161, 164, 165; 99: 118; [104: 238 imperfectana 98: 211; 104: 236 imperita 99: 126 implicatana 102: 181, 182, 187, 197; 104: 238 impunctana (ab.) 99: 120; 100: 325 impura (ab.) 99: 141 incana 111: 15 incanana 99: 121; 100: 324, 325; 104: 237 incarnatana 108: 382 incertana 98: 161, 162, 163; 99: 117; 100: 323, [324; 104: 237 incinerana 111: 14 incisana 102: 207 includana 102: 209 incognitana 101: 242, 248, 254, 257 inconditana 98: 224; 104: 234 inconspicua 108: 376, 384; 110: 17 inconspiqua 101: 252 incursana 101: 252; 104: 240 indecorana 104: 232 indignana 97: 216; 99: 126; 104: 236 individiosana 111: 10, 20 indivisa 110: 20, 21 indodatana 110: 76, 80, 84 infidana 110: 68, 69, 70, 71, 72, 82 infidanum 110: 72 infirmana 111: 18 infuscanus 99: 109 informosana 102: 193 infuscata (Dichrorampha) 110: 16 infuscata (ab.) (incanana) 99: 121 infuscatana 108: 381 ingentana: 97: 178; 98: 205; 100: 310, 311 ingentanus 98: 205; 104: 235 inignana 110: 76, 84 iniquana 110: 76, 84 inopinata 102: 208 inopinatana 98: 202 inornata (ab.) 98: 203 inquilina 103: 111, 117; 104: 239 inquinatana 102: 176, 181, 183, 191; 104: 239 insignana 99: 135 insincera 98: 218 insolatana 98: 209; 99: 111 instratana 102: 190; 104: 239 insubrica (ssp.) 99: 108; 100: 319, 320 insulana 103: 119, 143; 104: 239 insulana (ab.) (cristana) 99: 145 insulana (ab.) (hastiana) 99: 143 intacta (Eucosma) 111: 15 intacta (Grapholitha) 102: 211 interjectana 99: 114; 100: 323 interjunctana (ab.) 99: 114 36 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 intermediana (ab.) (paleana) 98: 211; [100: 313; 104: 236 intermediana (Eucosma) 111: 13 intermediana (hastiana) 99: 145 internana 102: 212, 215; 104: 238 interrogationana 108: 375, 384 interruptana 102: 181, 186, 191, 199, 200; [104: 239 interscindana 102: 188, 198; 104: 238 inulivora 108: 377, 385 inumbratana 98: 210; 104: 236 invalidana 98: 208 iridescens 102: 194 irrita 103: 139 irrorana 99: 136 irrorana (ab.) (hastiana) 99: 139 irrorana (ab.) (literana) 99: 150, 151; [104: 233 isacma 102: 211; 110: 24, 25, 26 ishidai 98: 218 ishidaii 98: 218 isocyrta 98: 206 isshikii (ab.) 99: 126 italica 99: 122 interponana 110: 15 interruptana 110: 20, 21 isocampta 110: 35 jaceana 111: 8 J jacquiniana 101: 245; 102: 191; 108: 376 jansoniana (ab.) 99: 145 janthinana 102: 203, 206, 207, 213; 104: 239 japonica 99: 136 jaquiniana 101: 245, 246 jaspidana 108: 383 jecorana 98: 201; 104: 235 jerusalemana 111: 6 jesonica 102: 193; 110: 22, 23, 24 joannisana 99: 108 joannisi (Croesia) 99: 128 joannisi (Eana) 99: 121; 100: 325 joannisi (ab.) (hastiana) 99: 140 joannisiana 98: 227; 100: 319; 104: 234 joannisiola (f.) 111: 15 jodocana 110: 76, 82 jottrandi (ssp.) 98: 209 juliana 103: 119, 124 juncta (ab.) 99: 115 junctana 108: 383 junctistrigana 102: 193; 104: 239 jungiana 102: 209, 210 jungiella 102: 206, 209, 213; 103: 129; [104: 239 juniperana 103: 115, 116, 122, 143; 104: 238 K kemnerana 111: 7 kenneli 99: 116; 100: 324; 104: 237 kenneli (ab.) (congelatella) 99: 119 kenteana 110: 81, 88 klimeschi 101: 243, 252; 110: 16 klimeschiana 101: 255 koenigana 102: 193 koenigiana 102: 193 (262) kokeilana 103: 117, 123 kollarana 102: 202 kollariana 110: 76 krausiana 102: 206, 209, 213; 104: 238 kurdistana 111: 10 krygeri 111: 12 L labatiana (ab.) 98: 213; 100: 315; 104: 236 labecula 99: 129 labeculana (ab.) 99: 129; 100: 327; 104: 234 labyrinthicana 110: 68, 80, 188, 226 lacordairana 99: 135; 104: 233 lacordairiana 99: 135 lacteana 111: 4, 7, 15 lacteanum 111: 7 lacunana 98: 201 laetana 99: 112 laetulana 108: 381 laevigana 98: 200, 206 lafauriana 98: 203; 104: 235 lafauryana 97: 171, 172 lambessana (ssp.) 102: 188 lamprana (ab.) 99: 137; 104: 232 lanceolana (Eucosma) 111: 9 lanceolana (ab.) (succedana) 102: 184 lapponana 97: 213; 98: 203; 104: 234 lapponana (= abiskoana) 99: 154 larsana 110: 15 larseni 102: 207, 213 larseni (ab.) (cristana) 99: 144, 146 larseni (ab.) (spectrana) 98: 215 lasiandra 104: 67 lasithicana 101: 250 latericiana 110: 69, 77, 78, 79, 85 lathyrana 102: 186, 206, 209; 103: 117; [104: 238 latifasciana 98: 193; 99: 128, 129, 137; [100: 327; 104: 234 latifasciana (ab.) (comariana) 99: 129 latiflavana 101: 246, 256 latior (ab.) 99: 115 latiorana (Eucosma) 111: 15 latiorana (ab.) (spectrana) 98: 215 latofasciana (ab.) 99: 139 laurana 98: 216 laviceana 98: 221 lavidorsana 101: 246 leachana (ab.) 99: 139 leacheana 98: 220; 99: 139 lecheana 97: 199, 201, 203; 98: 220; 104: 235 lediana 102: 186, 203, 207 leechi 99: 127 lefebvriana 99: 143 leguminana 102: 186; 104: 239 lemvigiana (ab.) 99: 139 lepidana 98: 202 lepidulana 102: 190; 110: 20 leplastriana 102: 186; 103: 139, 141, 142; [104: 238 leporinana (ab.) 99: 149; 104: 233 leprosana 99: 138, 139, 142, 143 leptograpta 98: 218 lerneana 107: 20; 110: 78 leucitis 102: 193 (263) leucodora 110: 35 leucogrammana 102: 193; 104: 238 leucophaeana (ab.) 99: 139; 104: 233 leucophracta 98: 228 leucotaeniana 99: 117 leucotreta 104: 67 levigana 98: 206 libanicolana (ssp.) *107: 21, 22, 23 liberana 99: 138 lichenana (ab.) 99: 145 lignana 111: 13 ligulana 101: 243, 250; 104: 240; 110: 19 lineata 99: 124; 100: 326; 104: 237 lineolana 110: 72 linophagana (f.) 99: 115; 100: 324 liotoma 98: 214 lipsiana 99: 129, 131, 147; 100: 332; 104: 233 lipsiana (ab.) (maccana) 99: 150 liratana 98: 214; 100: 315; 104: 235 literana 98: 189, 190, 193; 99: 150, 151; [104: 233 lithargyrana 99: 133; 100: 328, 329 lithargyranum 99: 133 lithosiana 98: 227; 104: 234 litterana 99: 150 littoralana 108: 379 littorana 102: 184 lividana (ab.) 99: 130 livoniana 98: 222; 99: 109 livonica (ab.) 99: 122 lobarzewskii 103: 117, 123, 205, 209, 213; [104: 238 loefflingiana 99: 125 loeflingiana 98: 179, 180, 181, 182, 190; [99: 125 loeflingianum 104: 236 loewiana 99: 111 loderana 102: 209 logaea (ssp.) 107: 13, 16 löfflingiana 99: 125 logiana 98: 162; 99: 115, 117, 134, 136, 138, [141; 100: 327, 330; 104: 233 longana 98: 159, 161; 99: 111; 100: 320, 321; [104: 237 longepartitana 99: 140 longiana 99: 134 longicellana 98: 206; 100: 311 longipalpana 99: 153; 104: 234 longulana 99: 132 luteana (ab.) 99: 127 lorquiniana 99: 148; 100: 332; 104: 234 lotana (ssp.) 102: 186 lotinana 97: 175 loxiana 103: 120 lubricana 98: 190; 99: 148; 100: 332; 104: 234 lucia (ab.) 99: 112 luciana 111: 16, 18 lucicostana 104: 236 lucidana (ab.) 99: 148 lucinana 111: 16 luctiferana 103: 120 luctuosana 99: 116; 100: 324 luctuosana (= tetragona) 108: 382 luctuosana (f.) (scutulana) 108: 378 luctuosanum (f.) 108: 378 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 37 ludicra 110: 24 luedersiana 103: 116, 121; 104: 239 lugubrana (Eucosma) 111: 3, 4, 5, 11 lugubrana (ab.) (alpicolana) 99: 124 lunana 98: 226; 100: 319 lunulana 101: 245; 102: 210 luridalbana 99: 111 lusana 98: 213 luteolana 98: 211; 99: 154 luticostana 98: 203; 100: 310; 104: 236 lutiplaga (ab.) 99: 149; 104: 233 lutosana 99: 111 lyrana 111: 18 lythargyrana 99: 133; 100: 328 M mabilliana 98: 222; 104: 236 macana 99: 149 maccana 98: 190, 193; 99: 137, 149; 104: 233 maccana (ab.) (abietana) 99: 149 macilentana 103: 121 macrogrammana 103: 136 macrorris 108: 379, 386 mactana 98: 219 maculiferana 110: 79 maculipactana 99: 153; 104: 233 maculostriana (ab.) 99: 140 maderae 103: 142; 104: 238 magnana 99: 120 magnanima 98: 208 magnificana 98: 220; 100: 317; 104: 235 major (Laspeyresia) 102: 184, 191, 200 major (Thiodia) 107: 21, 23, 24 malacotorna 98: 224 malcolmiae 103: 142 malitiosana 111: 13 malivorana 99: 153 malivorella 102: 193 mancipiana 110: 66, 67, 69, 70, 71, 72, 73, 82 maraschana (ssp.) 99: 110; 100: 320 marcidana 98: 224; 99: 110; 100: 320 margaritalis 99: 120 margarotana 107: 41, 42, 43, 44 marginata (ab.) 98: 227 marginestriana 102: 193 marginula (ab.) 98: 227 mariana 102: 189 maritima 111: 15 marmaranthes 110: 35 marmarocyma (Dichrorampha) 101: 251 marmarocyma (Eucosma) 111: 18 marmaroxantha 110: 76 marmorana 111: 7 maroccana 99: 123 masoniana (ab.) 99: 145 maura (ab.) 102: 191 mayrana (ab.) (hastiana) 99: 139, 140; [104: 233 mayrana (ab.) (hippophaéana) 99: 142 mayrana (ab.) (scabrana) 99: 143 mediana 102: 187 mediocris (ab.) 99: 115 mediterranea (Epiblema) 108: 382 mediterranea (Cnephasia) (,,race’’) 99: 110 38 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 medullana 111: 6 megerleana 102: 210 meincki 99: 129 melaleucana 103: 117 melanatma 98: 224 melanaula 110: 34 melaneura 110: 33, 34 melicrossis 102: 194 melissa 98: 224 melstediana (f.) 108: 377 menciana (f.) 97: 179; 98: 208 mendiculana 108: 379 menotoma 98: 206 meridionalis 99: 109 merlana (ab.) 99: 145 mesoloba 98: 225 mesomelana 99: 116 messingiana (Eucosma) 111: 12, 13 messingiana (Thiodia) 107: 20 metacritica 110: 33 metallicana (ab.) 99: 140 metana 111: 13 metria 110: 74 metzneriana 111: 15 micaceana 98: 206 micaceana (ab.) (succedana) 102: 184 micantana 98: 220; 104: 235 microgrammana 103: 134, 135, 136; 104: 238 micromys 98: 214 micronca 98: 225 micropterana 108: 381 mienshani 99: 116; 100: 324 miniatana (ab.) 107: 13, 15 ministrana 97: 220; 98: 152, 157; 99: 109; [104: 236; 107: 11 ministranus 99: 109 minor 99: 111 minorana 99: 112, 117 minorana (ab.) (consortana) 101: 249 minusculana 99: 117 minuta 98: 224; 100: 318 mirana 107: 21 mirificana 111: 17 mitterbergeriana (ab.) 99: 149; 104: 233 mixta (ab.) 99: 140 mixtana 99: 125, 134, 146, 147; 100: 332; [104: 233 mixtana (ab.) (hastiana) 99: 140 mixtana (ab.) (literana) 99: 151; 104: 233 modeeriana 98: 214; 99: 131; 100: 315 moderiana 98: 214 modestana (aemulana) 111: 15 modestana (modicana) 110: 77 modestana (montanana) 101: 251 modestana (rectifasciana) 99: 108; 100: 319 modestana (rigana) 99: 124 modicana (Eucosma) 111: 19 modicana (Pelochrista) 110: 77, 84 moeschleriana 97: 213; 104: 234 moldovana 102: 212, 215; 104: 239 molesta 102: 208, 213 mollitana 110: 74, 82 molybdana 102: 183, 191; 104: 239 monetulana 111: 7 (264) monochromana 99: 110, 120; 100: 320 monstratana 111: 7, 19 monstruosana 110: 34 montana 98: 227; 103: 120 montanana 101: 243, 245, 251; 101: 258; [104: 240; 110: 14, 18, 19 montedorea (ssp.) 98: 222 monticola 99: 125 monticolana (montanana) 101: 251 monticolana (pseudoalpestrana) 101: 250 monticolana (m. alt.) (rigana) 99: 124; [104: 236 montserrati (ab.) 99: 121 mortuana 108: 378 motacillana 103: 119 mucidanum 99: 133 mughiana 107: 13, 39 muliebris 111: 12 mullerrutzi 101: 252 mulsantiana 107: 38 multipunctana 99: 133; 100: 329 multipunctata (ab.) 99: 151 multistriana 102: 188, 198 mulzeriana 99: 136 mundana 102: 212; 104: 238; 110: 26 murinana 97: 171; 98: 203; 104: 235 musculana 97: 176, 183; 98: 209; 99: 114; [100: 322; 104: 235 musculinana (ssp.) 98: 209; 104: 235 myriopa 98: 195 myrrhophanes 99: 126 N nannodes 103: 121; 104: 239 napaea 99: 142 nebritana (Eucosma) 111: 14 nebritana (Grapholitha) 101: 236; 102: 185, [196, 206, 208, 213; 104: 238 nebulana 98: 206; 99: 130 nefrana 99: 109 negatana 102: 185, 195; 104: 238 neglectana 98: 216; 100: 316; 104: 235 nemorana 99: 107 neogena 98: 164; 99: 118 neogenum 99: 118 nervana 99: 123; 100: 326 nessebarana 111: 12 neurophaea 98: 218 nevadensis 99: 123; 100: 326 niconditana 98: 224 nigrana (ab.) (cristana) 99: 145 nigrana (ab.) (hastiana) 99: 140 nigricana 101: 236; 102: 176, 180, 181, 182, [185, 208; 104: 238 nigricana (ab.) (abietana) 99: 149 nigricaudana 97: 168; 98: 202; 104: 235 nigrimaculana 111: 16 nigriradix 99: 131 nigritana 102: 194; 104: 239 nigrobasis (ab.) (hastiana) 99: 209 nigrobasis (scabrana) 99: 211 nigrobrunnea 101: 253; 110: 17 nigrociliana 102: 211, 215 nigrocostana (ab.) 99: 145 (265) nigrocristana (ab.) 99: 145 nigrodivisana (ab.) 99: 140 nigrofasciana (ab.) 99: 150; 104: 233 nigromaculana (ab.) (Acleris) 99: 151 nigromaculana (Eucosma) 111: 3, 16 nigromaculata 111: 16 nigropunctana 102: 200 nigropunctana (ab.) (cristana) 99: 145 nigropunctata (ab.) (hyerana) 98: 227 nigroruficostana (ab.) 99: 145 nigrosignana (ab.) 99: 142 nigrostriana 102: 194; 104: 238 nigrostrigana (ab.) 99: 140 nigrosubvittana (ab.) 99: 145 nigrovittana (ab.) 99: 140 nimbana (fagiglandana) 102: 190 nimbana (fasciana) 103: 119 nitidana 103: 129; 104: 239 nitidulana 103: 129 nitorana 111: 11 niveana 99: 134, 136, 137 niveicaput 111: 11 niveosana (ssp.) 99: 120; 100: 325, 327; [104: 237 nivisellana 99: 131, 136 nobiliana (ab.) 98: 221; 104: 236 nordlandiae (f.) 98: 221; 100: 318 notana (ab.) 99: 133; 100: 329 notata 103: 119 notatana 99: 150 nougatana 102: 183 novana 108: 378 novickii 101: 246 novimundi 102: 185 nubiferana 97: 182 nubilana 98: 169, 170; 99: 119; 100: 324; [104: 236, 237 nubilea 99: 118 nuraghana 99: 111; 100: 321 nybomi 98: 218 nyctemerana 99: 135 O obcaecana (ab.) 102: 211, 215; 104: 238 obesana (Eucosma) 111: 6 obesana (Petrova) 107: 33 oblimatana 98: 218; 104: 236 obliquana 98: 205, 215 obliterana (= unifasciana) 98: 215 obliterana (ab.) (ligulana) 101: 250 obliterana (ab.) (tripunctana) 97: 206; [99: 132; 104: 233, 234 oblongana 97: 172 obnuptana (f.) 110: 78 obraztsovi 110: 17 obscura (ab.) (hastiana) 99: 140 obscura (ab.) (hippophaéana) 99: 142 obscura (ab.) (micaceana) 98: 206 obscura (ab.) (ribeana) 98: 200; 104: 235 obscura (ab.) (rosana) 98: 207 obscurana (Dichrorampha) 101: 249 obscurana (Epiblema) 108: 377, 385 obscurana (Eucosma) 111: 14 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 39 obscurana (Laspeyresia) (aurana) *102: 187, [198 obscurana (Laspeyresia) (— cosmophorana) [102: 185 obscurana (Pammene) 103: 112, 117, 118, 123; [104: 238 obscurana (ssp.) (cumulana) 111: 10 obscurana (ab.) (pascuana) 99: 115 obscurana (ssp.) (sedana) 99: 110; 100: 320 obscurana (ab.) (rhombana) 99: 131; 104: 234 obscurana (ab.) (ulmana) 99: 107 obscuratana 101: 247, 257 obsoletana 99: 114, 115, 116, 138; 100: 323, [324 obsoletana (ab.) (hastiana) 99: 140; 104: 233 obsoletana (ab.) (lecheana) 98: 220; 104: 235 obsoletana (ssp.) (musculana) 98: 209 obstinatana 110: 74 obtusana 99: 135, 136 obumbratana 111: 9 occultana 107: 12 ochracea (ab.) (emargana) 99: 152 ochracea (ab.) (rosana) 98: 207; 100: 311 ochreaceana 98: 224 ochreana 97: 191; 98: 211, 221, 223; 99: 132; [100: 312; 104: 235 ochreana (ab.) (apiciana) 99: 148 ochreana (ab.) (cristana) 99: 145 ochreana (f.) (farfarae) 108: 377 ochreapunctana (ab.) 99: 145 ochreocervina 103: 134; 110: 32; ochreofasciana (ab.) 99: 140 ochreoservina 110: 32, 33 ochreostriana (ab.) (apiciana) 99: 148 ochreovittana (ab.) 99: 140 ochriclivis 97: 220, 227; 98: 151; 99: 107 ochsenheimeriana 103: 120; 104: 239 octomaculana 99: 114, 122; 104: 237 oculatana 111: 17 ocydroma 99: 133; 100: 328 olbienana 102: 207 oleraceana 99: 114 olgana 99: 126 olivacea (ab.) (hastiana) 99: 140 olivacea (ab.) (montanana) 101: 251 olivana (ab.) 99: 151 olseniana 99: 148 ombrodelta 104: 64, 67 ongana 99: 111 onustana 98: 199 opacana 99: 139 opacana (ab.) (abietana) 99: 149; 104: 233 operana 98: 204 operosa 99: 126 oporana 97: 178, 179; 98: 200, 204, 205, 207, [100: 310 opulentana 102: 188; 111: 16 orana 98: 219 oricasis 99: 110, 117; 100: 320, 324 orientana (Cnephasia) 99: 110; 100: 320; [104: 237 orientana (Epiblema) 108: 383, 387 orientana (ssp.) (rosana) 98: 207; 100: 311 orientata 110: 24 ornamentana 110: 80, 85, 87 40 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 ornata 103: 120, 125 ornatana 104: 64 orobana 102: 210, 213; 104: 239; 110: 21 orphnocycla 99: 148; 100: 332 orthioterma 98: 194 orthoxyana 99: 112; 100: 321 osmana 107: 41, 42 osseana 98: 172, 173, 226; 99: 120; 100: 325; [104: 237 osthelderi (Acleris) 99: 142 osthelderi (Cnephasia) 99: 114 osthelderi (f.) (Eucosma) 111: 15 ottoniana (Eucosma) 111: 3, 5, 11 ottoniana (Palpocrinia) 98: 163; 99: 117 ovulana 103: 119 oxyacanthana 98: 206; 99: 119 oxycedrana 103: 115, 116, 122, 143; 104: 238 oxydroma 99: 133 oxystaura 103: 121; 110: 36 oxytropides 102: 188 oxytropidis 182: 88 B pactolana 102: 181, 189, 198; 104: 238 padana 98: 201 paetulana 111: 17 paleana 97: 185, 191, 192; 98: 210, 211; [100: 313; 104: 236 pallasana 107: 12 palleana 98: 211 pallens (ssp.) (xylosteana) 98: 205; 104: 235 pallens (ab.) (hyerana) 98: 227; 100: 319 pallida (ab.) 99: 120 pallidifrontana 102: 212 pallifrontana 102: 212; 104: 239 palmoni 99: 117 pamira 102: 184, 195 panzerana 103: 120 paradiseana 99: 153; 104: 232 paradoxa 99: 119 paraliana 99: 122; 100: 325 parastrepta 102: 194; 110: 29 parisiana 99: 136 partitanum 98: 165; 99: 118 parvana (ab.) 99: 114 parvulana (Strophedra) 103: 129 parvulana (f.) (Eucosma) 111: 9 pascivana 99: 115 pascuana 97: 220; 98: 161, 162; 99: 115, 117; [100: 321, 323, 324; 104: 237 pasivana 98: 162; 99: 115; 100: 323 pasquagana 98: 201 pasquayana 98: 201 passadenana 107: 10 passivana 99: 115 pauperana 111: 3, 6, 12 pavonana 102: 194 paykulliana 111: 12 pectinana 98: 225 pedana 108: 381 pedemontana (ssp.) (aspersana) 99: 132 pedemontana (ssp.) (hybridana) 99: 108; [104: 238 pencleriana 102: 190 penkleriana 102: 190 (266) pentagonana 108: 379 pentalychna 102: 194 pentheriana 101: 253 pentziana 99: 122 penziana 98: 169, 170, 172, 173; 99: 114, 120, [122; 100: 325 peramplana 98: 211; 100: 313 perdicina 98: 199 perfracta 104: 66 perfuscana (ab.) 99: 119 pergratana 111: 11 perlepidana 102: 209 permixtana 99: 146 permutana 99: 136; 104: 233 permutatana 99: 136 pernix 99: 143 perochreana (ab.) 98: 209; 100: 312; 104: 236 perplexana (ab.) 99: 129; 104: 234 perplexana (chrysantheana) 99: 113 perpulchrana 98: 222; 104: 236 persiae 111: 6 persicana 102: 194; 110: 21 persimilana 98: 219; 104: 235 personatana 99: 117; 104: 237 perterana 99: 113, 116, 117 petasitis 108: 381 petiverana 101: 245, 246 petiverella 101: 242, 245, 246, 248, 257; petiverella (Dichrorampha) 101: 242, 245, 246, [248, 257; 103: 114; 104: 240; 110: 15 petiverella (Eucosma) 111: 14 petiverellum 101: 245 pettitana 98: 195 petulantana 104: 51, 52, 53, 54, 239 peyerimhoffi (ab.) 99: 114 pfeifferi 102: 189, 198 pfisteri (Dichrorampha) 101: 252; 110: 16 pfisteri (Pelochrista) 110: 74 pflegeriana 99: 125 pflugiana 108: 375, 378 pfugiana 108: 378 phacana 102: 213; 103: 117 phaeana 98: 211 phaeoli 110: 33 phalacris 102: 184, 195 pharaonana 104: 59, 60, 61, 239 phaseoli 103: 131, 132, 133, 134; 110: 32, 33, [34, 35 phasmatica 97: 189; 98: 210 phaulomorpha 102: 194 piceana 97: 172, 178, 179; 98: 204, 205; [100: 310; 104: 234 pictana 108: 380 pierrelovyana 98: 223 pierretana 108: 377 pietruskii 108: 377 pillerana 99: 154 pilleriana 97: 222; 98: 180, 194, 195, 196, [210; 99: 154; 100: 333; 104: 234 pinana 98: 209 pinetana 102: 190, 199; 103: 116; 104: 239 piniana 107: 14, 17 pinicola 107: 12 pinicolana 107: 10, 12 pinivorana 107: 10, 13, 15 (267) pisana 102: 185 pistaciana (ab.) (mabilliana) 98: 222 placata 99: 153 placida (ssp.) 98: 216; 100: 316 placidana (Acleris) 99: 136 placidana (Thiodia) 107: 21, 23 plagiferana 98: 212 planifrontana 102: 186 platynotana 99: 153 plumbagana 101: 238, 239, 246, 251, 254, 257; [104: 240 plumbagana 101: 247 plumbana (Acleris) 99: 134 plumbana (Aleimma) 99: 125 plumbana (Dichrorampha) 101: 241, 244, [255, 256, 259, 260, 261; 104: 240; [110: 13, 14 plumbatana 103: 116 plumbeana (ssp.) (argentana) 99: 120 plumbeana (ab.) (hastiana) 99: 140 plumbeofasciana (ab.) 99: 140 plumbeolana 98: 220; 99: 125; 104: 235 plumbeostriana (ab.) 99: 140 plumbiferana 102: 180, 181, 184, 195; [104: 238 plumbosana (ab.) 99: 134; 104: 233 plusiana 101: 250 podana 97: 176, 178; 98: 204, 205; [100: 310, 311 podanus 104: 235 podoliensis 101: 242, 253 poecilana 108: 376 policolana 104: 237 polita 99: 118 politana (Argyrotaenia) 98: 202; 100: 309 politana (Dichrorampha) 101: 245, 249, 254 politana (Lathronympha) 103: 139 politum 99: 118 pollinis (ab.) 107: 48 polyxena 111: 18 pomana 102: 188 pomonella 101: 233; 102: 175, 179, 181, 182, [183, 188, 198; 104: 239 pontica 103: 115, 116, 122, 143 populana 103: 111, 112, 118; 104: 239 porphyrocentra 99: 153 porrectana 99: 108 postchantana (ab.) 99: 145 posticana (Blastesthia) 107: 35, 38 posticana (Clepsis) 98: 215; 100: 315 postmaculana (ab.) 99: 140 potentillana (ab.) 99: 129; 104: 234 pouzatella (ab.) 98: 222 praecana 97: 220, 222; 98: 225; 104: 234 praeclarana 98: 212; 100: 314; 104: 236 praedorsana 102: 210, 214 praefactana 110: 78, 85 praefloratana 98: 202; 100: 309; 104: 235 praeviella 99: 124; 100: 326; 104: 237 pratana 99: 120 pretiosana (ab.) 99: 128; 104: 234 prismatica 102: 194 privatana 98: 219 probolias 98: 224 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 41 prochantana (ab.) 99: 145 procristalana (ab.) 99: 145 prodromana 97: 209, 214; 98: 226; 100: 319 prodromanus 104: 234 productana 98: 215 profana (ab.) 99: 145, 146 profanana (ab.) 99: 144, 145, 146; 104: 232 profundana 108: 376, 378, 386 profundana (ab.) 99: 145 pronuba 98: 209 pronubana 97: 176, 180, 182, 224; 98: 209; [100: 312; 104: 236 pronuhana 104: 236 prospera 110: 31 prostriana (ab.) 99: 145 proteana (ab.) (comariana) 99: 129; [104: 234 proteana (ab.) (tripunctana) 99: 132; 104: 233 provinciana (ab.) 99: 147; 104: 233 provittana (ab.) 99: 145 provocata 97: 175; 98: 203; 100: 310 proxanthovittana (ab.) 99: 146 proxima 101: 245; 110: 15 proximana (Acleris) 99: 153 proximana (Laspeyresia) 102: 185 proximana (Pammene) 103: 120 pruinosana (ab.) 99: 140; 104: 233 prunifoliae 102: 208 prunivorana 102: 209, 213 pryerana 108: 379, 386 pryerana (ab.) (affinitana) 99: 135 pseudoalpestrana 101: 250, 258; 104: 240 pseudoalternella (ab.) 99: 116 pseudochrysantheana (ab.) 99: 114 *pseudocinerosana (ssp.) 101: 243, 248, 257 pseudocommunana (ab.) 99: 117 *pseudoconfixana 99: 140; 100: 330 *pseudocoronana (ab.) 99: 143; 100: 331; [104: 232, 233 pseudologiana (ab.) 99: 141 pseudolongana 99: 120 pseudomayrana (ab.) 99: 142 *pseudomodestana (ab.) 99: 108; 100: 319; [104: 238 pseudomorpha 110: 24 pseudonectis 110: 30 pseudorthoxyana (ab.) 99: 112 pseudotypica 99: 115; 100: 324 psorana (ab.) (hastiana) 99: 139 psorana (ab.) (scabrana) 99: 143; 100: 331; [104: 233 ptichogrammos 98: 193 ptychora 110: 30 pudendana 107: 13 pudicana 102: 186 puellana 99: 108 pulchella 103: 120, 125 pulchellana 97: 152, 168; 98: 202; 100: 309; [104: 236 pulchra 98: 220 pulmonariana 99: 115 pulverana 98: 213; 99: 136 pulverula 102: 194 pumicana 98: 161; 99: 111; 100: 321; [104: 237 42 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 punctana (ab.) 99: 145 puncticostana 103: 120 puncticostata 103: 120 punctiseriata 97: 195 punctulana 98: 169, 173, 175; 99: 124; [100: 326; 104: 237 punctulella 99: 124 punicae 98: 207 pupillana 111: 8, 17 purdeyana (ab.) 99: 145 purdeyi 107: 48 purpurana (ab.) 99: 148 purpureana 103: 120, 143; 104: 239 pusillana (Epagoge) 98: 224; 100: 318 pusillana (Grapholitha) 102: 209 pusillana (Lathronympha) 103: 139 putaminana (f.) 102: 188, 198; 104: 239 pylonitis 111: 12 pyrana 98: 207; 100: 312 pyrastrana 98: 204 pyrenaea (ssp.) 99: 123 pyrenaica 99: 121 pyrivora 102: 179, 182, 189, 198 pyrivorana 99: 153; 100: 333; 104: 233 pyrophagana (f.) 99: 115; 100: 323 Q quadarramana 101: 256 quadrana 108: 376 quadratana 108: 376 quadridentana 99: 153 quadripunctana 99: 120 quadripunctata 99: 120 quadristriana 102: 211; 110: 25 quadrocellana 102: 194 quaestionana 101: 246, 254; 110: 15, 25 quaestonana 101: 245 querceti 103: 120 quercinana 98: 190; 99: 133; 100: 329; [104: 234 questionana (alpinana) 101: 245 questionana (ab.) (alpinana) 101: 246; {104: 240 quinquefasciana 99: 128 quinquemaculana (ab.) 98: 217; 104: 236 R radiana (ab.) (apiciana) 99: 148 radiana (ab.) (ferrugana) 99: 133; 100: 329 radiana (ab.) (hastiana) 99: 139, 140, 141; [104: 233 *radiana (ab.) (scabrana) 99: 143; 100: 331; [104: 233 radiana (ab.) (umbrana) 99: 137 *raebeli (ssp.) 98: 211 ragatzana (ab.) 99: 142; 104: 233 ragazana 99: 142 ramostriana (ab.) 99: 140; 104: 233 (hier steht remostriana !) rasana (f.) 98: 221 rastrata 99: 122; 100: 325 ratifera 102: 185 ravana 108: 381 ravulana 103: 117, 123; 108: 376 reaumurana (ab.) 102: 190, 199; 104: 239 (268) recentana 108: 365; 111: 10 recreantana 102: 194; 104: 239 rectifasciana 97: 231; 98: 155; 99: 107, 108; [100: 319, 320; 104: 238 rectilinea (ab.) 99: 114 rediana 103: 120 rediella 103: 121 redimitana 103: 129 reducta (ab.) 99: 112 refrigescens 102: 186; 110: 20 regiana 103: 119; 104: 239 regisborissi 98: 212; 100: 314 reisseri 103: 118, 124 reisseri (= Cnephasia fragosana) 99: 110 rejectana 101: 252; 104: 240; 110: 13, 14 relictana 107: 12 remostriana (ab.) 104: 233 (Druckfehler fiir ramostriana!) repandana 99: 149 repletana 104: 67 resedana *102: 186, 196; 103: 143 (fig.) resinana 107: 33 resinella 107: 13, 33, 38 resplendana 101: 256; 104: 240; 110: 13 resupinatana 111: 18 retiana 98: 222 retucilana 98: 221 reticulana (Adoxophyes) 97: 196, 199, 201, [203, 219; 100: 317; 104: 234 reticulana (= Aphelia ochreana) 98: 211; [100: 313 reticulana (rhombana) 99: 131 reticulana (ab.) (sparsana) 99: 130; 104: 233 reticulata 98: 219; 100: 317, 327 reticulata (ab.) (rhombana) 99: 131; 104: 234 retiferana (Clepsis) 98: 215; 100: 315; [104: 236 retiferana (Gravitarmata) 107: 42, 44, 45 retiferana (Rhyacionia) 107: 13 retractana 97: 176, 180; 98: 208; 100: 312 reynana 98: 226 rhaeticana (Dichrorampha) 101: 251; [104: 240; 110: 18, 19 rhaeticana (Rhyacionia) 107: 13, 15 rhediana 103: 120, 121 rhediella 103: 111, 112, 120; 104: 58, 239 rheediana 104: 58 rheediella 103: 121; 104: 58 rhezelana 102: 18 rhodochropa 99: 126 rhodopa 98: 199 rhodophana 97: 231; 98: 152, 156, 157; [99: 108; 104: 236 rhombana (Acleris) 97: 192; 98: 189; [99: 131; 100: 317, 327; 104: 234 rhombana (Aphelia) 98: 210; 100: 312 rhombana (Clepsis) 98: 217 rhombicana 98: 217, 225 rhombicanus 104: 234 rhopica 97: 216 rhytmologa 98: 219 ribeana 97: 166; 98: 200; 100: 309, 310; [104: 235 rielana 99: 121 * (269) rigana 98: 157, 169, 175, 177; 99: 124; [104: 236 rigidana 111: 7, 19 rilana 101: 251 rimosana 108: 380, 386 rivulana 103: 117 rjabovi 101: 254 roborana (Archips) 98: 205 roborana (Epiblema) 108: 382 rogana 97: 196; 98: 213; 100: 315; 104: 236 rolandriana 98: 213; 100: 315; 104: 237 romanana (ab.) 99: 150; 104: 233 rosaceana 98: 207; 100: 311; 104: 235 rosae 108: 383 rosaecolana 108: 383 rosana 97: 178; 98: 204, 205, 206, 207; [99: 127, 131; 100: 309, 311 rosanus 104: 235 roscidana 99: 130; 104: 233 roseidana 99: 130 roseticolana 102: 185, 208 rosinana 102: 195; 110: 26 rossiana 99: 143 rotundana 108: 380, 387 rubellana (ab.) 99: 148 rubescana (f.) 111: 14 rubiana 108: 383 rubicundana 97: 222; 99: rubidana 99: 154 rubidana (ab.) (ferrugana) 99: 133; [100: 329; 104: 234 154; 104: 234 rubrana 98: 201 rubromaculata (ab.) 98: 205 rufana (Acleris) 99: 129, 132, 133, 147, 148; [100: 328, 332 rufana (Eucosma) 111: 9 rufana (ab.) (hastiana) 99: 138 ruficostana (ab.) 99: 144, 145; 104: 232 ruficristana (ab.) 99: 145 rufifasciana (ab.) 99: 138 rufillana 102: 186 rufimaculana (ab.) (caledoniana) 99: 130 rufimaculana (ab.) (hastiana) 99: 140 rufimixtana (ab.) 99: 140 rufinigrana (ab.) 99: 145 rufistrigana (ab.) 99: 140 rufivittana (ab.) 99: 140 rungsi 102: 185, 196 rureana 97: 175 ruriana 98: 214; 100: 315 rushana 110: 80, 88 rusticana 98: 213, 224; 100: 314, 318; [108: 376 S sabulana (ab.) 99: 132 sabulosa 100: 324 *saerdabana 111: 17 sagittana (ab.) (cristana) 99: 145 sagittana (ab.) (hastiana) 99: 140 salicetana 101: 246 saltitans 102: 183 salvana 102: 185; 103: 116, 121; 104: 239 sappadana 104: 54, 55 sapporensis 97: 154; 98: 199 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 43 sardiniana107: 21, 23 sardivola (ssp.) 99: 151 sardoensis 111: 16 sareptana 107: 27, 28, 29 sarmatana 108: 377 sarthana 98: 216; saturana 98: 226 saturnana (= montanana) 101: 251; 110: 14 saturnana (ab.) (plumbana) 101: 255, 261; [104: 240 sauberiana (ab.) 98: 204; 104: 235 saussureana 111: 14 scabrana 99: 128, 136, 137, 139, 141, 142, [143; 100: 330; 104: 232 scabrana (ab.) (emargana) 99: 152; 104: 232 scabrana (ab.) (literana) 99: 158 schalleriana 98: 190; 99: 128, 129, 132, 134; [100: 327, 329; 104: 233, 234 schatzmanni 101: 251 schawerdae 98: 222 schawerdai 99: 124 schlemmerella 99: 124 schrankiana 102: 209 schreberiana 99: 128 schumacherana 99: 107; 100: 319 scintillana 102: 206, 211, 215; 104: 238; [110: 66 100: 316; 104: 235 scopaliana 111: 8 scopoliana (Epiblema) 108: 365, 375 scopoliana (Eucosma) 111: 7, 8,9 scopolianum 111: 8 scorzonerana 111: 9, 19 scotana (ab.) 99: 137; 104: 233 scoticana (ab.) 99: 140 scriptana 104: 64 scrophulariana (ab.) 98: 210; 100: 313; [104: 236 scrutatrix 97: 195 scutana 111: 9 scutellana (Aphelia) 98: 212; 100: 314 scutiformis 111: 7 scutulana (Epiblema) 108: 374, 376, 377, [378, 386 sebastianiae 102: 183 seclusa 99: 126 sedana 98: 161; 99: 109, 110; 100: 320, 324; [104: 237 sedatana 101: 255, 261 seditiosa 98: 206 seeboldi 107: 25, 27 seeboldianus 98: 225 seeboldiana104: 234 segetana (ab.) 99: 111; 104: 237 segnis 99: 126 selasana (ab.) 99: 133; 100: 329; 104: 234 selectana 103: 125, 126, 143; 104: 239 selenana 102: 210; 104: 239 selliferana 102: 206, 210, 214; 104: 239 semialbana 97: 196; 98: 214; 100: 315; [104: 235 semiana (ssp.?) 98: 215 semibruneata 99: 110 semibrunneata 99: 110 semicinctana 102: 180, 181, 191, 199; [104: 239 44 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 seminigra (ab.) 99: 112 seminotata 103: 121 semirombana 99: 134 semistriana (ab.) 99: 145 semistructa 98: 219 semitexta 99: 153 semiustana (ab.) 99: 145 senecionana 98: 213; 100: 314; 108: 379 senectana 101: 243, 247, 257; 104: 240 separana (f.) 108: 376, 384 separatana 102: 189 sepiana (ab.) 99: 145 sequana 101: 243, 245, 248; 104: 239, 240 sequana (ab.) (cristana) 99: 145; 104: 232, [233 seriana 110: 76, 84 sericana 101: 253; 104: 240 sericana (ab.) (cristana) 99: 146; 104: 232 servillana 102: 176, 178, 180, 181, 187, 198; [104: 238 servilleana 102: 187 sescuplana 98: 219 severana 98: 216; 104: 235 shanghainana 98: 220 sheldonana (ab.) 99: 140 sheljuzhkoi (Acleris) 99: 149; 100: 332 sheljuzhkoi (Dichrorampha) 101: 253, 260 shepherdana 99: 134; 104: 234 simulata 107: 13, 16 siciliana 98: 216; 104: 236 siennicolor (ab.) 99: 114 signana (ab.) 99: 139 significantana 111: 6, 18 similana (Cnephasia) 100: 323 similana (Epiblema) 108: 376, 377, 379, 385, [386 similana (ab.) (apiciana) 99: 148; 104: 233 similis 99: 131 simillimana 102: 212 simonyi 98: 219; 104: 235 simplicana 101: 249, 254, 258 simpliciana 101: 249; 104: 240; 110: 15 simploniana 108: 376, 380 simpsoni (f.) 102: 188, 198 simulana (f.) 97: 179; 98: 208 simutata 107: 14 sinana 102: 205, 212, 215; 104: 238 sinapina 98: 217; 100: 316; 104: 235 sineana 98: 221 sinensis (ssp.) (Cerace) 98: 200 sinensis (Eucosma) 111: 13 sinicana 108: 375, 384 sinuana 99: 112, 113, 115, 121 slavana (ab.) 101: 245 snellenana 103: 121, 125 socialis (f.) 98: 208 solandriana 100: 314; 108: 366, 383 solida 98: 206 sorbiana 97: 178; 98: 207; 100: 310, 311; [104: 235 sordicomana 111: 6 soriana 98: 215; 104: 237 southiana (ab.) 99: 146 spadiceana 107: 13 spadiceana (ab.) (cristana) 99: 144, 146 (270) spargotis (ssp.) 98: 208 sparsana 98: 193; 99: 130, 142, 143; [100: 317, 319, 327, 333; 104: 233 sparsana (ab.) (cristana) 99: 145 sparsana (Eucosma) 111: 12 sparsana (ab.) (hastiana) 99: 138, 140 speciosana (ab.) 99: 142 spectrana 97: 185, 196; 98: 215; 100: 315, [316; 104: 235 spectrana (ab.) (boscana) 99: 136; 104: 233 spectrana (ab.) (logiana) 99: 137 spinetorum 103: 118 spiniana 103: 118, 124; spixiana 98: 227 splendana 102: 183, 190, 199; 104: 239 splendana (ab.) (rosana) 98: 207; 100: 311 splendidana 102: 190; 103: 116 splendidulana 103: 116; 104: 239 sponsana 99: 130, 153; 100: 327 squamana (ab.) (literana) 99: 150, 151; [104: 233 squammulana (ab.) (literana) 99: 150, 151 squamulana (ab.) (literana) 99: 150, 151; [104: 233 104: 239 stachi 99: 131 stagnana 98: 213 *staintoni 98: 219; 100: 316; 104: 235 staudingeri 99: 152 steinerana 98: 217 steineriana 98: 213, 217; 100: 315; 104: 236 steliferana 101: 254 stelliferana 101: 245 stelviana 99: 120 stenochorda 98: 226; 100: 319 *stenoptera (ab.) 98: 211; 100: 313 stephensiana 99: 113 stereomorpha 98: 223; 100: 318 stibiana 99: 127; 100: 326; 104: 236 sticticana (Epiblema) 108: 376, 378, 385 sticticana (Pammene) 103: 118 stigmatana 98: 211; 104: 236 stipatana 97: 152, 154, 157; 98: 195 stramentana 99: 108; 104: 238 straminea 98: 222 straminea (ab.) (gnomana) 98: 222 stramineana 98: 213; 100: 316 stramineana (ab.) (strigana) 98: 217; [104: 235 strandiana (ab.) 102: 210 strangulana 103: 119 stratana 99: 111 striana (strigana) 98: 217 striana (ab.) (cristana) 99: 144, 145, 146; [104: 232 striana (ab.) (hastiana) 99: 140 *striana (ab.) (scabrana) 99: 143; 100: 331 striata (ab.) (lorquiniana) 99: 148 striatana 102: 189, 199 striatana (ab.) (lorquiniana) 99: 148 striatiradix 111: 13 striatulana 98: 219 strigana (Clepsis) 98: 217; 100: 316; [104: 235, 236 strigana (Dichrorampha) 101: 254 strigana (Eucosma) 111: 8 (271) strigana (Latronympha) 103: 136, 138, 139; [104: 238 strigana (ab.) (hastiana) 99: 139 strigulana 99: 147 striolana 98: 217; 104: 235 strobilana (Dichrorampha) 101: 246 strobilana (Pammene) 103: 116 strobilana (Pseudotomoides) 102: 202 strobilella (Pammene) 103: 111, 116, 117 strobilella (Pseudotomoides) 102: 200 strobilellus 102: 201, 202; 104: 238 strobitella 102: 202 stroemiana 108: 378, 385, 386 stryriacana 99: 113 (Druckfehler für styriacana) styriacana 99: 113 stygiana (ab.) 98: 209 styriacana 99: 113, 114, 120, 121 styx (ab.) 99: 112 suavana (ab.) 99: 151; 104: 233 subalboflammana (ab.) 99: 146 subaurantiana 98: 227 subcapucina (ab.) 99: 146 subcapuzina (ab.) 99: 146 subchantana (ab.) 99: 146 subclarana (f.) 98: 201 *subcoprana (ab.) 99: 154; 100: 333 subcostana 98: 215; 104: 236 subcristalana (ab.) 99: 146 subcristana (ab.) 99: 140; 104: 233 subdivisana (ab.) 99: 141 suberana 103: 116 subfasciana (ab.) (hastiana) 99: 141 subfasciana (ab.) (ministrana) 99: 109; [104: 236 subfascianus 99: 104 subferruginea (ab.) 99: 138 subfulvovittana (ab.) 99: 146 *subgrisea (ab.) 99: 141; 100: 330 subhastiana (ab.) 99: 141 subjectana 98: 162; 99: 117 subjunctana 98: 221; 104: 236 sublimana 108: 380 sublucidana 111: 9 subnigrana (ab.) 99: 146 subradiana (ab.) 99: 141 subrigidana 108: 381 subrufana (ab.) 98: 205; 104: 235 subscabrana (ab.) 99: 141 subsequana 101: 247, 251 subsidiaria 98: 208 subsponsana (ab.) 99: 142 substriana (ab.) 99: 141, 146; 104: 232 substrigana 98: 217 subterminana 110: 74 subtiliana 110: 78 subtripunctulana 99: 131 subunicolorana (ab.) 99: 146 subvittana (f.) (cana) 111: 8 subvittana Sheldon (ab.) (hastiana) 99: 141 subvittana Stephens (ab.) (hastiana) 104: 232 succedana 102: 181, 184, 185, 195; 104: 238 succineana 110: 70, 78, 84 sucineana 110: 78 sudorana (ab.) 99: 147 sudoriana (ab.) 99: 147; 104: 233 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 45 suffusana 108: 382, 383 suffusana (ab.) (literana) 99: 150 suffusana (ab.) (maccana) 99: 149 sulfatarana (ab.) 99: 120 sulphurana 107: 17, 21, 23 sumptuosana (Laspeyresia) 102: 195 sumptuosana (Pseudargyrotoza) 98: 228 suomiana 111: 16 suppurpuratum 97: 218 suspectana 103: 118, 123; 104: 239 sutschana 98: 219; 100: 317 suttneriana (ab.) 99: 125; 104: 236 sybillana 111: 12 sylvana 98: 202 sylvestrana 107: 13, 38, 46, 47, 48 sylvicolana 101: 246, 250, 255, 256, 261 T taeniodesma 98: 225 tagarica (ssp.) 111: 16 tamerlana 101: 253, 258; 110: 17 tanacetana 101: 247 tanaceti 101: 247, 251, 253, 255, 258 tapaishani 98: 213 taradana 111: 13 tarandana 111: 13 tarica (f.) 111: 11 taurominana 99: 111; 100: 321 teleopa 107: 12, 14, 15 teliferana 103: 139; 111: 18 tenebrana 102: 195; 104: 238 tenebriosana 102: 185 tenebrosana 102: 185, 208; 104: 238 tephromorpha 99: 147; 100: 332 terebrana 99: 110; 100: 320 termias 98: 206 terreana 98: 224; 100: 315 terstrigana 102: 212 teshionis 98: 208 tessulatana 107: 33, 34, 35, 36 testaceana (ferrugana) 99: 133 testaceana (ssp.) (decretana) 98: 204 testaceanus (ssp.) (decretana) 104: 235 tetragonana 108: 379, 382 tetragrammana 102: 212, 215 tetraonis 98: 199 tetraplana 111: 17 tetraploca 104: 67 textana 98: 200 tharsaleopa 98: 204; 100: 310 tholera 110: 79 thuriferana 103: 116, 122 thurificana (ssp.) 107: 12 tianshanica 99: 116; 110: 16 tibetana (ssp.) (Exapate congelatella) 99: 119 tibetana (Pelochrista) 110: 80, 85 tibialana 108: 375 tigricolor 99: 126 timana 110: 22, 23 tinacriana 111: 3, 11 tofina 99: 111; 100: 321 tolana (ab.) 99: 146 tolerana 110: 76 tomiana 103: 117, 123; 104: 238 46 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 toreuta 102: 182 tornimaculana 110: 68, 73, 77, 82 torquana 99: 149 torrana 101: 255, 260 tortricea 99: 118 tortricella 97: 220; 98: 161, 165; 99: 118 tortricellus 104: 237 tragoditis 98: 223 transapiciana (ab.) (cristana) 99: 146 transapiciana (ab.) (hastiana) 99: 141 transcaspica (ssp.) 100: 324 transitana (ab.) 98: 202, 203 translucida 99: 126 transversana (ab.) (cristana) 99: 146 transversana (ab.) (hastiana) 99: 141 trasias 102: 195 trauniana 103: 111, 113, 119; 104: 239 treitscheana 100: 318 treitschkeana 98: 223; 100: 318; 104: 234 treueriana 99: 136, 137 treueriana (ab.) (literana) 99: 150, 151 treveriana 99: 136, 137 triana 99: 132 tricentra 110: 30 trichocrossa 110: 24, 31 trichosema 104: 67 trichroa 97: 175 tricolorana (ab.) 99: 150, 151; 104: 233 tricuneana (ab.) 102: 194; 104: 239 trifasciana 98: 209; 99: 107 trigeminana 108: 376, 377 trigeminanum 108: 376 trigonana (Acleris) 99: 134 trigonana (Pammene) 103: 118, 124 trigonana (Tortrix) 99: 126 trigonella 102: 191 trimaculana (Acleris) 99: 133; 100: 329 trimaculana (Epiblema) 108: 382, 383 trimaculata (ab.) 99: 150 trinacriana 111: 11 trinotana 103: 119 tripoliana 111: 14 tripsiana 98: 219 tripunctana (Acleris) 99: 132; 100: 327, 328, [329; 104: 233, 234 tripunctana (Epiblema) 108: 366, 382 tripunctana (ab.) (logiana) 99: 137 tripunctulana (ab,) 99: 133; 100: 329 trisignana 110: 66, 70, 74 trisignana (ab.) (logiana) 99: 137 tristana (Acleris) 99: 130, 134 tristana (Grapholitha) 102: 208 tristana (ab.) (hastiana) 99: 141; 104: 233 tristana (f.) (hepaticana) 108: 379 tristana (ssp.) (internana) 102: 212, 215 tristana (ab.) (maccana) 99: 149 tristana (ab.) (sparsana) 99: 130 tristrami 99: 110; 104: 237 trivia 98: 216 trivittana (ab.) 99: 141 trophiodes 103: 134; 110: 32, 33 tscheliana 107: 21 tshetverikovi 110: 16 tshimgana 101: 246 tundrana 111: 12 (272) tunicatana 99: 153 turatiana (ab.) 98: 222 turbidana 108: 371, 373, 381 turbitana 108: 381 turiana 110: 74, 75, 82, 83 turionana 107: 11, 12, 38 turionella 107: 11, 12, 37, 38 tussilaginana 108: 381 typhlodes 101: 247 tyrrhaenica 99: 123 U ubagonana 98: 213 uddmanniana 108: 366, 369, 372, 375, 383, [387 uhagonana 98: 213 uhagoni (ab.) 98: 213; 100: 315; 104: 236 ulicana 101: 255 ulicetana (ab.) 102: 184, 195; 104: 238 uliginosana (ab.) 99: 148; 104: 234 ulmana 98: 152; 99: 107; 100: 319; 103: 124; [104: 238; 107: 11 ulmana (f.) (boscana) 99: 136; 104: 233 ulmicola 99: 136; 100: 330 ulotana (ab.) 99: 146 umbraculana (f.) 110: 72 umbrana 99: 137; 104: 232 umbratana 111: 10 umbrosa 97: 186, 188, 189, 191, 231 undana 98: 203 undosa 101: 245 undulana 98: 203; 99: 154; 100: 333; [104: 233 undulata 99: 128 unguicella 104: 64 unicolor 101: 250 unicolor (ab.) (chondrillana) 98: 202 unicolor (ab.) (gnomana) 98: 222 unicolor (ab.) (hastiana) 99: 141 unicolorana 97: 195, 196; 98: 213; 100: 315; [104: 236 unicolorana (ab.) (cristana) 99: 146 *unicolorana (ab.) (fimbriana) 99: 147; {100: 332 unicolorana (f.) (foenella) 108: 376, 384 unicolorana (ab.) (rhombana) 99: 131 unifasciana 97: 185, 193, 196, 198; [98: 215, 216; 100: 316, 317; 104: 235, 236 uniformana 99: 116 uniformata (ab.) 99: 140 uniformis 99: 143 uniformis (ab.) (variegana) 99: 135 unipunctana 98: 210; 100: 312, 313 unitana 98: 210 uralensis 101: 255 urbana 103: 139; 111: 13 ursulana 111: 12 urticana 98: 203 ussuriana (ssp.) (Eucosma) 111: 16 ussuriana (ssp.) (Eucosmomorpha) 104: 58 ustomaculana (ab.) (cristana) 99: 146 ustomaculana (ab.) (hastiana) 99: 141 ustulana (ab.) (cristana) 99: 146 ustulana (ab.) (hastiana) 99: 141 (273): V vacivana 101: 255 valderiana (f.) 99: 109; 100: 320 vallifica 98: 208; 100: 312 y vana 103: 141, 142, 143; 104: 238; 110: 36 variana 98: 206 variegana 99: 135; 100: 330; 104: 233 variegana (ab.) (hastiana) 99: 141; 104: 233 variegata (ab.) 99: 142 variostriana (ab.) 99: 142 vaughaniana (ab.) 99: 146 venansoni (ab.) 99: 121 verbascana 98: 211; 100: 313 verecundana 111: 18 vermicularis 98: 226 vernana (= snellenana) 103: 121 vernana (ab.) (argyrana) 103: 117, 123; [104: 239 vertheimsteini 99: 123 veterana 99: 142 veterana (ab.) (hastiana) 99: 141; 104: 232 vetulana 99: 112, 130, 321; 104: 237 vexilla 102: 195 viardi 99: 122 viburnana 98: 210, 221 viburnana (ab.) (schalleriana) 99: 134 viburniana 97: 185, 189, 191, 192; [98: 210, 211, 221; 100: 312; 104: 236 vicariana 98: 207 viciana 102: 185 victoriana 111: 10, 20 viduana 98: 202 vigeliana 103: 129 vinculana 98: 215; 100: 316 violaceana 99: 134 violascens 98: 227 virescana 98: 219 virgaureana 98: 161, 163; 99: 112, 114, 115, [122; 100: 323; 104: 237 virgaurenana 99: 114 virginana 98: 161, 162; 99: 112 virginiana 104: 237 virgulana (ab.) 99: 132 viridana 98: 180, 181, 182, 184, 211; 99: 125; [104: 236 viridochraceum 97: 218 vitis 99: 154 vittana (ab.) (cristana) 99: 146; 104: 232 vittana (ab.) (hastiana) 99: 137 voluta 97: 189 vulgana 98: 224 vulgaris (ab.) 99: 114 vulnerata (Laspeyresia) 102: 193 N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 47 vulnerata (Pentacitrotus) 97: 152 vulpeculana 98: 204; 100: 311 vulpisana (ab.) 98: 201; 104: 235 W wahlbaumiana 99: 112, 113 wahlbomiana (Cnephasia) 99: 110, 112, 113, AUS NU IO 0:23 2250323 wahlbomiana (Epiblema) 108: 377 wahlbomiana (Sciaphila) 98: 162 walbomiana 100: 323; 99: 115 walkerana 98: 226 walkeri 98: 226 walkeriana 98: 226 walsinghami 107: 10, 13, 15 washiyai 107: 12, 15 wassiana 98: 218 webbiana (ab.) 99: 146 weberana 104: 64 weirana 103: 128; 104: 239 wertheimsteini 98: 172; 99: 123 westriniana 98: 205 westwoodiana 111: 9 wilkinsoni 99: 113; 100: 322, 323 wimmerana 111: 15 wimmeriana 111: 12, 15 wöberana 104: 64 woeberana 104: 64 woeberiana 104: 61, 64 wolfschligeriana (ab.) 99: 148 x xanthocosma 98: 195 xanthoides 98: 195; 99: 154; 100: 333 xanthomitra 100: 318 xanthovittana (ab.) 99: 144, 146 xylosteana (Archips) 97: 166, 175, 179; [98: 201, 204, 205, 206; 107: 11, 12 xylosteana (Acleris) 99: 131 xylosteanus 104: 235 xylostiana 98: 205 xylotoma 98: 215; 104: 236 Z zachana (Dichrorampha) 101: 255 zachana (Eucosma) 111: 14 zebeana 102: 188; 104: 238 zelleri 103: 117, 123 zelleriana 108: 381 zermattana (ssp?) 101: 254 zygogramma 102: 195; 110: 29, 30 48 10. 1 12: TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 (274) POSTHUMOUS PUBLICATIONS OF THE LATE Dr. N. S. OBRAZTSOV . Neotropical Microlepidoptera, IX. Revision of genus Pseudatteria (Lepidoptera: Tortricidae). — Proc. U.S. Nat. Mus. (no 3535) 118: 577—622, Fig. 1—12, Pl. 1—43, 1966. . Neotropical Microlepidoptera XI. Revision of genus Idolatteria (Lepidoptera: Tortricidae). — Loc. cit. (no. 3543) 119: 1—12, PI. 1—8, 1966. . On the correct usage of two names of the genus-group in the Tortricidae, with the proposal of a new generic name (Lepidoptera).— Entomologist's Gazette 17: 141—147, 1966. . Genera Tortricidarum. Check list of the genera and subgenera belonging to the families Tortricidae (Ceracidae, Chlidanotidae, Schoenotidae and Olethreutidae included) and Phalo- niidae. — Journ. New York Ent. Soc. 75: 1—11, 3.V.1967. . Some apocryphal species of the Tortricinae (Lepidoptera, Tortricidae). — Loc. cit. 75: 34, 1966. . Notes on the genus Homonopsis V. Kuznetsov (Lepidoptera, Tortricidae). — Ent. Berichten 27: 173—175, 1967. . Die Gattungen der palaearktischen Tortricidae III. Addenda & Corrigenda, 2, Teil. Notes on the Palaearctic Laspeyresiini. — Tijdschrift voor Ent. 110: 13—36, Fig. 1—2, Pl. 1—2, 1967. . Die Gattungen der palaearktischen Tortricidae. II. Die Unterfamilie Olethreutinae, 7. Teil. Tribus Eucosmini (Heinr. 1923). Fortsetzung. — Tijdschrift voor Entomologie 110: 65—88, Fig. 1—24, Pl. 3—10. . Die Gattungen der palaearktischen Tortricidae. II. Die Unterfamilie Olethreutinae. Tribus Eucosmini (Heinr. 1923), 8. Teil und Schluss, mit einem Gesamtregister auf alle Teile, zusam- mengestellt von B. J. Lempke. — The present article. In press: Descriptions and records of South Asiatic Laspeyresiini (Lepidoptera, Tortricidae). Notes on and descriptions of Aphelia, Clepsis and Choristoneura species (Lepidoptera, Tortri- cidae). Records and descriptions of Palaearctic and South Asiatic Laspeyresiini (Lepidoptera, Tortri- cidae). TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT Taf. 1. Eucosma-Arten. Abb. 1, Eucosma sordicomana Stgr., &, Prip. No. M 742, Mti Simbruini, Italien (Mus. München). Abb. 2, Eucosma medullana Stgr., è, Ohrid Umgebung, Mazedonien, 700 m, 27.VI.1954 (J. Thurner), Prip. No. 67 (Mus. Wien). Abb. 3, Epiblema significantana Kennel, Lectotypus, 4, Präp. No. B33, Mazedonien (Kr.) (Mus. Berlin). Abb. 4, Epiblema gypsatana Kennel, Typus &, Präp. 5730, “Achr.”, Anatolien, 1900 (Caradja, No. 2011, WIsm. No. 71784). (BM). Abb. 5, Grapholitha rigidana Snellen, 4, Lectotypus, Insel Askold, 1878, Prip. No. 2638 (Mus. Leiden) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 2 i Taf. 2. Eucosma-Arten. Abb. 1, Epiblema significantana Kennel, Allolectotypus, 9, Beirut, Syrien, Präp. B. 34 (Mus. Berlin). Abb. 2, Epiblema monstratana Rebel, Lectoallotypus, ©, Schaufigg bei Chur, 1600 m, 16.VII.1903 (G. Stange), Präp. No. V. 28 (Mus. Wien). Abb. 3, Epiblema jerusalemana Amsel, ©, Paratypus, Kasr el Jehud, Jordan, 7.1II.1930 (H. G. Amsel) (Mus. Miinchen). N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 3 5 Taf. 3. Eucosma-Arten. Abb. 1, Eucosma albuneana Zeller, ®, Span, Marokko, Ketama, 1500—1600 m, Mitte VI.1941 (W. Marten) (Mus. Wien). Abb. 2, Ewcosma scutiformis Meyr., Kwanshien, China, VII.1928 (F. M. Frank), 9, Präp., 6858. Abb. 3, Dasselbe, Detail von Bursa copulatrix, stärker vergrößert. Abb. 4, Dasselbe, Detail von Ostium und Antrum, stärker vergrößert. Abb. 5, Epiblema monstratana Rebel, Lectoallotypus, ®, Schaufigg bei Chur, 1600 m, 16.VII.1903 (G. Stange), Falter. Abb. 6, Dasselbe, &, Falter N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 4 3 Taf. 4. Eucosma-Arten. Abb. 1, Epiblema monstratana Rebel, Lectotypus, 3, Präp. No. 2048, Schaufigg bei Chur, 1600 m, 16.VII.1903 (G. Stange) (Mus. Wien). Abb. 2, Epiblema expallidana balatonana Osth., Lectotypus, 4, Präp. M. 1053, Vörs, Comit. Somogy, Ungarn, 9.VII.1931 (L. Osthelder), Falter. Abb. 3, Dasselbe, Lecto- typus, Genitalien (Mus. Miinchen) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 5 Taf. 5. Eucosma-Arten. Abb. 1, E. umbratana Stgr., 4, Armenien, Präp. M. 856 (Mus. München). Abb. 2, Eucosma fervidana Zeller, 4, Wippach, 1854, Präp. No. V. 62 (Mus. Wien). Abb. 3, Eucosma flavispecula Kuzn., Illmitz, Burgenland, Österreich, 11.VII.1954 (H. Reisser). (A.M.N.H.). Abb. 4, Eucosma güntheri Tgstr., “Eppelsh.”’, Prip. No. 23-Obr. 8/8.1960 (Mus. München). Abb. 5, Eucosma (Eucosma) flavispecula Kuzn., dasselbe wie Abb. 3, Genitalien. N. S. OBRAZISOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 6 Taf. 6. Eucosma-Arten. Abb. 1, Epiblema victoriana Kennel, Lectoallotypus, 2 (kein abdomen), Sajan (Mus. Berlin). Abb. 2, Dieselbe Art, Lectotypus, 4, Sajan, Präp. No. B. 27 (Mus. Berlin). Abb. 3, Dieselbe Art, 4, Lectotypus, Sajan, Falter. Abb. 4, Epiblema recentana Zerny, Lecto- typus, 4, N. Libanon, Cedern b. Bechamé, 1900 m, 12—19.VI.1931 (H. Zerny). Abb. 5, Thiodia niveicaput Wals., Lectoallotypus, 9, Japan, Pryer, 1886 (No. 70100), Prip. 6325 (BM). Abb. 6, Semasia amellana Preissecker, Lectotypus, 4, Bisamberg b. Wien, 7.VIII.1903, Präp. No. V. 39 (Mus. Wien) N. S, Opraztsov : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 7 Taf. 7. Eucosma-Arten. Abb. 1, Palpocrinia ottoniana Kennel, &, Holotypus, Lobnoor, Präp. B 1 (Mus. Berlin). Abb. 2, Thiodia niveicaput Wlsm., 4 Präp. No. 6325 (BM). Abb. 3, Dieselbe Art, 9, Japan, Pryer, 1886, No. 70102, No. 6306 (BM). Abb. 4, Eucosma brachyclista Meyr., 4, Nanking, China 28.IX.1933 (K.), Präp. No. 6843 (BM). Abb. 5, Dieselbe, Falter. Abb. 6, Semasia sparsana Rebel, Holotypus, 4, Ankara, VI.1934 (H. Noack), Präp. No. V. 25 (Mus. Wien) , Japan, Pryer, 1886, No. 70099, N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 ‘8 ‘qqv "(uapIaT SJN) TEST ‘ON ‘derd LLSTIA IT muy wel eursyss] ‘sndAjopaT ‘2 ‘Jus purgajs vqigogdrio ‘L ‘qqy aeg ‘Pp ‘QQV SIM [gay Modkıy vısvwas ‘9 ‘qqy (WEI) 8SLS ‘ON ‘derd ‘¢ ‘qqv SIM ‘2 ‘SndÂJOIOH “wisi szswours vrpoIgT, ‘Ss “qqy (ULM SN) FE “A CON ‘dead ‘ CEGI'IA‘GI (DIN) SECLIAT SINO Pur vısmuagsy ‘puejjoelS ‘ssuweig Ory, ‘snd403227 ‘È TPM 2425047 vısvwas ‘y -qqy (Nd) (ISTO9 “ON ‘Y999T) O8STTA'T ‘eUIUD 'S ‘SueryayD ‘odeur ‘9 ‘sndAyoqaT “USA Sısuauıs PIPOIGL “€ ‘QQV “(UPI SON) 6€ “A ‘ON "deg ‘COGT'IIIA L “2 ‘sndA0PIT ‘TONPASSTAIT Pur vıspwas ‘T qqy ‘(UM 'snW) (IUSEA]ED) COGI UTNSTUIFT ‘puejuasn y ‘opero josur ‘9 ‘snd4jeredop9T ‘tseayey sysuaprrd visvutas ‘1 ‘QQV ‘uarry-ru)somd ‘8 Jer alaearktischen Tortricidae P Die Gattungen der S. OBRAZTSOV : N. TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 9 Taf. 9. Eucosma-Arten. Abb. 1, Semasia amellana Preissecker, Lectoallotypus, 9, Mödling, Oster- reich, 19.VIII.1908 (Richardhof), Präp. No. V. 40 (Mus. Wien). Abb. 2, Semasia krygeri Rebel, 9, Lectotypus, “410, Bramnes, Sjaelland, Artemisia maritima Galle, 2.VI.1935 (Kryger), 19.VI.1935”, Prip. No. V. 34 (Mus. Wien). Abb. 3, Thiodia intacta Wlsm., Holotypus, 4, Japan, 1886, Pryer, No. 70172 (BM). Abb. 4, Eucosma (Phaneta) galactica Obr. n.n., Holotypus, 4, Emba, Uralsk, Prip. M. 1042 (Mus. Miinchen) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae PLAAT 10 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 1 “qqv SI sorjea “IGO vurgopsops Ivurjpidud (viaurqd) vuisoma ‘y ‘qq (ME) 1899 ON ‘derd ‘(2u0H) ZE6I ounf ‘00€ ‘uIyD ‘UEUS-NN-UAIL ‘2 “IAN rulsomwwsvu auowtwop ‘€ ‘AY '6SLS ON ‘deig ‘TLIOL ‘ON (1414) 9881 ‘uede( ‘9 ‘sndAJoJoH este priv PIPOIYIL ‘Tt ‘qqv "(vaypunyy Sn) (1235704 A ® J2jplajd A) LEGTIIAYI- OT “WH 0007 0061 ‘ueqrepur A ‘TEL € -gepies “so ‘SU sImqug “S EISI24 ‘© ‘sndhojoH “IQO vurqvpsavs vurndnd (vrourgd) vwsosng T “qqy ‘UANV-PMSOIMA “OL JEL . nn. Die Gattungen der palaearktischen Tortricidae N. S. OBRAZTSOV : TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 1, 1968 PLAAT 11 Taf. 11. Eucosma-Arten. Abb. 1, Semasia sardoensis Rebel, Lectotypus, 6, Uras, Sardinien, 16.X.1934 (Predota), Präp. No. V. 37 (Mus. Wien). Abb. 2, Hemimene marmarocyma Meyrick, 4, Tien-Mu- Shan, China, 5300’, June 1932 (Hone), Präp. No. 6681 (BM). Abb. 3, dasselbe wie Abb. 1, Falter. Abb. 4, Eucosma (Phaneta) pupillana saerdabana Obr., Allotypus, ®, Persia S., Elburs Mts., Särdab- Tal, Vandarban, 1900-2200 m, 10—14.V11.1937 (E. Pfeiffer & W. Forster), Präp. 2-Obr. 8/8/58 (Mus. München). Abb. 5, Ewcosma (Phaneta) tetraplana Möschl., 3, Achyr Dagh. s., Bertiz Jaila, 1800 m. 9—13.VI.1929, Marash, N. Syrien (E. Pfeiffer) (Mus. München) N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae pro ETTI Sen ad C È 3 (E? - | 7 | > Oo Ü > DES DEEL 111 AFLEVERING 2 1968 TIJDSCHRIFT VOOR ENTOMOLOGIE UITGEGEVEN DOOR DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING MUS. COMP. ZOOL, LIBRARY. JUL 25 1968 HARVARD UNIVERSITY INHOUD : FR. CHRYSANTHUS O.F.M. CAP. — Spiders from South New Guinea X, pp. 49—74, Figs. 1—95. Tijdschrift voor Entomologie, deel 111, afl. 2 Gepubliceerd 21-VI-1968 NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING BESTUUR (BOARD) Voorzitter (Chairman) . . on oe wae Ga Barendtecht Vice-Voorzitter (ize Biases) RE ek ET LWW EDES Secretaris (Secretary) . . . . . . . . W. Hellinga Add hersen . . « . . Weesperzijde 23 II, Amsterdam-O. Penningmeester Ceres) RDA ENT GI Kabos Address) 2: . . . . + Van Baerlestraat 261, Amsterdam Bibliothecaris Cube) RENO ne VELA ENTRER El Addis CN ER BU Leeburverdyen ol) Amsterdam-O: Teden (Members) e ON NT VA anse ADF MEL Besemer AFDELING VOOR TOEGEPASTE ENTOMOLOGIE (DIVISION OF APPLIED ENTOMOLOGY) BESTUUR (BOARD) Voorzitters (Chairman) meenten APE ME Besemer, Seeretaris A(Sectetary)a. 1. 2... EE SSP Gruys Adaness m ne ne Nederermasestraal 35, Kesteren: Leden (Members) .-. . . : . . 2... .L. Bravenboer, J. J. Laarman, J. B Me vans Dinther Publicaties van de Vereeniging (Publications of the Society) Subscription may be obtained from all booksellers or directly from the Librarian, Zeeburgerdijk 21, Amsterdam-O., except for Eniomologia Experimentalis et Applicata, which is available through booksellers or from the Noord-Holland Editing Co., Post Office Box 103, Amsterdam. TIJDSCHRIFT VOOR ENTOMOLOGIE Redactie (Editing Board) . . . . . . Pater Chrysanthus, A. Diakonoff, C. A. W. Jeekel, M. A. Lieftinck, J. T. Wiebes Address . . : 2 2 2 20200020. Rijksmuseum van Natuurlijke Historie, Raamsteeg 2, Leiden. The Journal serves the publication of papers on Insecta, Myriapoda and Arachnoidea. It appears in separate issues, forming an annual volume of 350—400 pages. Subscription rate: D.Fl. 45.— (£ 4.10.—, $12.50) per volume. MONOGRAFIEEN VAN DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING Redactie (Editing Board) and address as for Tijdschrift voor Entomologie. The Monographs are intended for the publication of larger entomological papers ‘on a single subject and will appear irregularly. The following Monographs have been published: Hors serie: F. T. Valck Lucassen et al, 1961. — Monographie du genre Lomaptera Gory & Percheron (Coleoptera, Cetoniidae), 299 pages, 739 figs, 2 pl., map. D.Fl. 50.— (£5——, $ 13.90). No. 1. A. J. Besseling, 1964. — De Nederlandse Watermijten (Hydrachnellae Latreille, 1802) (The Hydrachnellae of the Netherlands), 199 pp, 333 figs., D.Fl. 25 — (£2.10—, $ 6.95). È È B $ i \ SPIDERS FROM SOUTH NEW GUINEA X by FR. CHRYSANTHUS O.F.M. Cap. Oosterhout (N.B.) Abstract In the present paper, being the last part of this series, 21 species belonging to the family Salticidae are dealt with. All have been collected by Br. Monulf in the environs of Merauke (1956-1957) and Mindiptana (1958—1965). The species discussed are: Linus fimbriatus (Doleschall), Bavia aericeps Simon, Diolenius amplectens Thorell, Cytaea frontaligera (Thorell), C. nimbata (Thorell), C. mitellata (Thorell), Euryattus bleekeri (Doleschall), E. porcellus Thorell, Trite longula (Thorell), Sandalodes bernsteini (Thorell), Bathippus macrognathus (Thorell), B. papuanus (Thorell), Palpelius beccarii (Thorell), Plexippus paykullit (Audouin), Zenodorus durvillii (Walckenaer), Mopsus mormon Karsch, Poecilorchestes decoratus Simon, Cosmophasis bitaeniata (Keyserling), C. micarioides (L. Koch), Dendryphantes laticeps Strand and Menemerus bivittatus (Dufour). À list of all species discussed in this series is added. Acknowledgements I wish to express my sincere thanks to the following colleagues who kindly sent me material for study from their respective museums: Dr. L. van der Hammen (Rijks- museum van Natuurlijke Historie, Leiden), Dr. O. Kraus (Senckenberg Museum, Frankfurt a.M.; at some places indicated as SMF in the text), Dr. G. Rack (Zoologisches Museum, Hamburg), Dr. D. Guiglia (Museo Civico di Storia Naturale, Genova), Dr. M. Hubert (Muséum National d'Histoire Naturelle, Paris), Mr. D. J. Clark (British Museum, Natural History, London), and Dr. J. Cooreman and Mr. J. Kekenbosch (In- stitut royal des sciences naturelles de Belgique, Bruxelles). À special word of thanks is due to Dr. L. van der Hammen for his continuous help and advice in my study of the New Guinean spiders during the last ten years. Furthermore I am grateful to the Netherlands Organization for the Advancement ot Pure Research (Z.W.O.) and to the Consiglio Nazionale delle Ricerche, for grants which enabled me to study the rich Thorell collection in Genoa. ECRIBELLATAE (concluded) SALTICIDAE PLURIDENTATI Boethinae Linus Peckham, 1885 Linus fimbriatus (Doleschall, 1859) Fig. 1—6 Doleschall, 1859, Act. Soc. Sci. Ind.-Neerl. 5: 22, Pl. 5 Fig. 8, 28 (Salricus). Thorell, 1878, Ann. Mus. civ. stor. nat. Genova 13: 269, & (Siris). , 1881, ibid. 17: 499, 2 (Siris). Simon, 1901, Hist, nat. Araignées 2: 411, Fig. 435—443, 4. 49 50 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Fig. 1—6, Linus fimbriatus (Doleschall). 1, 9; 2, id. abdomen, ventral view; 3, id. cephalothorax, lateral view; 4, id. tibia, lateral view; 5, id. epigyne; 6, id. vulva. Fig. 711. Bavia aericeps Simon. 7, 9; 8, id. abdomen, ventral view; 9, id. epigyne; 10, 6, left palp, ventral view; 11, id. part of left palp, lateral view. Fig. 1—3: X 7; 4: X 5; 5: X 35; 6: X 60: 7, “Sir X 3: 10, 11: X 40 FR. CHRYSANTHUS : Spiders from South New Guinea X il Three females from Merauke (1956—1957) are identical with the female of this species in the Doleschall collection (Rijksmuseum van Natuurlijke Historie, Leiden), probably the holotype; their body length varies from 7.5—9.0 mm. The type locality is Amboina, the species has also been recorded from Ceylon to New Guinea and Australia, and even from Madagascar (Roewer, 1954: 935; Bonnet, 1957: 2482). Thiodininae Bavia Simon, 1877 Bavia aericeps Simon, 1877 Fig. 711 Simon, 1877, Ann. Soc. ent. France (5) 7: 61, 4. L. Koch, 1879, Arachn. Austr. 1 (2): 1146, Pl. 99 Fig. 6, 7, 2 8 (Acompse suavis). Simon, 1901, Hist. nat. Araignées 2: 470, Fig. 529—531, 4. In the Zoologisches Museum, Hamburg, two syntypes of L. Koch’s Acompse suavis are preserved, viz. one adult and one young female. Simon established the synonymy of this species with his Bavia aericeps; in “Berichtigungen”, published in 1883 at the end of “Die Arachniden Australiens” (vol. 1 (2): 1477), Keyserling said: “Herr E. Simon war so liebenswürdig mir mitzuteilen, dass Acompse suavis L. K. p. 1146, identisch sei mit der von ihm beschriebenen Bavia acriceps”. I could compare a Bavia female (12 mm) from Merauke (1956—1957) with Koch's syntypes: they are identical, my specimen being somewhat darker; Koch mentioned this darker form. His figure of the epigyne (Fig. 6d) is misleading: in the adult syntype it is as in my Fig. 9; in Fig. 6a he omitted the lateral notches in the labium. The male syntype is lost. Three males (11—14 mm) from Mindiptana (1959) fully agree with Koch’s description and figures; the pattern is as in the female, the colours are darker. The type locality of aericeps is Manila (Philippines); Koch's specimens of szavis originated from Huahine, Rayatea, Tahiti. The geographical distribution extends from Sumatra and the Philippines into the Pacific area (Roewer, 1954: 979; Bonnet, 1955: 871). Diolenius Thorell, 1869 Diolenius amplectens Thorell, 1881 Fig. 12—18 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 412, 9. Several Diolenius females from Mindiptana (1958—1965) fully agree with Thorell’s description and with types from the Aru Is. and Ramoi Riv. (Vogelkop, New Guinea) (Genoa Museum); the central part of the epigyne may be uniform brownish yellow. The body length varies from 6—8 mm. Strand (Ads. senckenb. Naturf. Ges. 34 [1911]: 179) mentioned two females from Terangan (Aru Is.). Male. About an equal number of Diolenius males were collected in the same years and the same locality as the females: they certainly belong to amplectens, no other species of this genus being present in our collection. In nearly all details they are like the females; the abdomen slenderer and marked by two longitudinal grey bands (Fig. 16); first legs more slender and twice the length of those of the female (Fig. 15, 17; 2 leg: X 20; & leg: X 10); the metatarsus with short spines (in the figure I omitted the dense fringe of rather long hairs, running along the underside of the tibia, 52 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 : a ELLE pn gan dek - | erts Fig. 12—18. Diolenius amplectens Thorell. 12, $ ; 13, id. epigyne; 14, id. vulva; 15, id. left I leg, lateral paraxial surface; 16, 6, abdomen; 17, left I leg, lateral paraxial surface; 18, id. left palp, ventral view. Fig. 19, 20. Cytaea frontaligera (Thorell). 19, ®; 20, 4. Fig. 21. C. nimbata (Thorell), 2. Fig.) 12,76, 20,2125 7132 X 70; 14: X 120: 15: X 20: 172 NON OMIS EDS AS XD Fr. CHRYSANTHUS : Spiders from South New Guinea X 53 in order to show the strong spines — the same with the female leg). The palp (Fig. 18) strongly resembles that of D. phrynoides (Walckenaer, 1837); in that species, however, the spermal duct of the bulbus runs in a flat curve towards the tip. Measurements: cephalothorax, length 3.3 mm, width 2.6 mm; abdomen, length 4.2 mm, width 1.7 mm; legs, I 20, II 8, III 8, IV 10 mm. Some males are smaller, e.g. total body length 6 mm FISSIDENTATI Cytaeinae Cytaea Keyserling, 1882 Cytaea frontaligera (Thorell, 1881) Fig 10202224 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 607, 9 & (Plexippus). Several females (8—11 mm) and three males (6—7 mm) from Merauke (1956— 1957) belong to this species, known from the Aru Is. and Queensland (Roewer, 1954: 1025; Bonnet, 1956: 1371). They fully agree with Thorell’s description and with the type material (Genoa Museum). The central yellow band on the abdomen of the females may be much wider because the dark reddish brown squamiform hairs are easily rubbed off. The epigyne is some- what variable: the distance between the two sperm canals may be greater and their strongly curved tips are not always discernible. In the male the pattern of the abdomen may be the same as in the female. Cytaea nimbata (Thorell, 1881) Fig. 21, 25—30 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 600, & (Plexippus). A large number of Cytaea specimens (about 30 males and as many females from Mindiptana, 1958—1965; one 4, one ® from Merauke, 1956—1957) belong to a second species. Two Cytaea species resemble each other to a high degree, viz. C. sinuata (Doleschall, 1859) and C. nimbata Thorell, 1881 (cf. Thorell, loc. cit.). The Doleschall collection (Rijksmuseum van Natuurlijke Historie, Leiden) does not contain specimens of Salticus sinuatus, nor of S. floricolus Doleschall, 1859, a synonym of sinzatus. This species has a wide distribution: it is known from Sumatra and the Philippines to Australia (Roewer, 1954: 1025; Bonnet, 1956: 1372). Of C. nimbata only three males are known: two from Hatam in the Arfak Mountains, one from Andai, Vogelkop, all in New Guinea (type material, Genoa Museum) (Roewer, 1954: 1026; Bonnet, 1958: 3717 [Plexippus}). I was able to compare my specimens with these types: the males are completely iden- tical. Moreover, they possess one of the few characters given by Thorell, through which the nimbata male differs from the sinuata male. In nimbata the circular hollow of the bulbus with the spiral of the embolus occupies nearly 4/5 of the bulbus, whereas in sinuata in occupies scarcely more than half its width (loc. cit.: 603). The body length of the males varies from 7—8 mm. Female (Fig. 21). Cephalothorax: length 4.0 mm, width 3.4 mm; a broad black band containing the eyes surrounds a brown central field, the remaining part is yellowish brown but for the greater part it is covered with black, somewhat flattened 54 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Fig. 22—24. Cytaea frontaligera (Thorell). 22, 2, epigyne; 23, id. vulva; 24, &, left palp, ventral view. Fig. 25—30. C. nimbata (Thorell). 25, 9, epigyne; 26, id. vulva; 27, 4; 28, id. left palp, ventral view; 29, id. part of left palp, dorso-lateral view; 30, id. dorsal view. Fig. 22, 25: X 70; 23, 26: X 60; 24, 28—30: X 40; 27: X 7 FR. CHRYSANTHUS : Spiders from South New Guinea X 55 hairs, which are easily rubbed off. Measurements of the legs: I 10.5, II 9.5, III 9.0, IV 8.5 mm. Abdomen: length 4.5 mm, width 2.5 mm; yellow, covered with black squamiform hairs, except for the central band; these hairs, too, are easily loosened; underside greyish yellow; in some specimens, however, with a dark brown central band as in the male. Epigyne: Fig. 25, in darker specimens the details are not so clear as in our figure. Vulva: Fig. 26; the complexity of the sperm canals explains the fact that in the epigynes the visible part of these canals is not always the same. The body length varies from 9—11 mm. Cytaea mitellata (Thorell, 1881) Fi2 1231122 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 604, & (Plexippus). Thorell based his description on three males, one from Aru Is., one from Yule 1. and one from Ternate. Three males in our collection (Mindiptana, 1958, 1965) are conform to the description and the type material (Genoa Museum) ; two of them are not so dark as in Fig. 31; their body length lays between 6 and 7 mm. There are no further records of this species (Roewer, 1954: 1026; Bonnet, 1958: 3717 [Plexippus}). Euryattus Thorell, 1881 Euryattus bleekeri (Doleschall, 1859) Fig. 33—36, 41, 42 Doleschall, 1859, Act. Soc. Sci. Ind.-Neerl. 5: 17, Pl. 3 Fig. 6, & (Salticus). Thorell, 1878, Ann. Mus. civ. stor. nat. Genova 13: 260, 9 & (Plexippus). , 1881, ibid. 17: 631, 9 4 (Plexippus). Keyserling, 1881, Arachn. Austr. 1 (2): 1299, Pl. 111 Fig. 1, ® (Hasarius albescens). , 1881, ibid.: 1300, PI. 111 Fig. 2, 9 (H. pauperatus). , 1881, ibid.: 1307, Pl. 111 Fig. 6, 2& (H. chrysostomus) syn. nov. Simon, 1903, Hist. nat. Araignées 2: 815, Fig. 962, 4. Doleschall’s collection (Rijksmuseum van Natuurlijke Historie, Leiden) does not contain a specimen of his Salticus bleekeri. Therefore a comparison of some 30 Euryat- ins specimens, males and females, from Merauke (1956—1957) and Mindiptana (1958—1965) with the holotype was not possible; they fit rather well the short des- cription and poor figure. The specimens in our collection perfectly agree with Thorell’s elaborate description of the male (1878: 260) and sufficiently with his much shorter description of the female (1878: 263, note). I studied the material discussed by him in his papers (Museo Civico di Storia Naturale, Genoa): in all details they are identical with my specimens. Simon gave some characters of this species (“très commun à Amboine et en Nouvelle Guinée”) and a figure of the characteristic dentation of the cheliceral furrow: our specimens agree with all of them. Moreover they are identical with some specimens in the Senckenberg Museum, Frankfurt a.M., identified by Strand, and with a pair from New Guinea in the Leiden Museum (det. probably Reimoser). Variability. — Males. A rather broad band of white hairs is generally present along the borders of the cephalothorax and so is the band of yellow hairs along the abdomen; these hairs, however, may nearly totally have been rubbed off; the remaining part of the abdomen is uniform dark brown or bears some brownish yellow patches, sometimes 56 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Fig. 31, 32. Cytaea mitellata (Thorell). 31, &; 32, id. left palp, ventral view. Fig. 33—36. Euryattus bleekeri (Doleschall). 33, 2; 34, 35, id. epigyne; 36, &, left palp, ventral view. Fig. 37—40. E. porcellus Thorell. 37, ®; 38, id. epigyne; 39, &, left palp, ventral view; 40, id. tarsal part. Fig. 3133: 7: 32,36, 402 X 40; 34,35, 382070 375 << 33) 393 a0 FR. CHRYSANTHUS : Spiders from South New Guinea X 57 united into a wedge-shaped band on the posterior half. The legs are unicoloured brown or variegated with broad yellowish bands. The variability of the dimensions is remarkable: the six males from Merauke measure 5.5, 7 (3 ex.), 7.5 and 8.5 mm; the ten males from Mindiptana: 6, 7, 7.5, 8 (2 ex.), 9, 9.5 (2 ex.) and 10 (2 ex.) mm. Thorell (1881: 631) says: “Long. 4.5—9 mm”. The palpi of the large specimens are somewhat longer than those of the smaller: tibia 0.68 (0.59) mm, tarsus 0.81 (0.68) mm; bulbus and tibial apophyse are exactly the same, only slightly larger. Females. Colour as in the males. The two females from Merauke measure 6 and 7 mm; of the ten females from Mindiptana one is 8 mm, the others are from 9.5— 10.5 mm. The epigyne and the vulva of the small Merauke specimens (Fig. 34, 41) are smaller than those of the large Mindiptana specimens (Fig. 35, 42); the structure, however, is the same in both. The sidewards curved furrows in the epigyne are not always as sharply marked as in our figures, the straight furrows are often rather vague; in the female described by Thorell (Genoa Museum) they are very near to each other and form a triangle with the anterior border of the epigyne (1878: 264). Synonymy. — Simon (1903: 815, nota 3) remarked about E. bleekeri: “Les Hasarius albescens | 2 }, pauperatus (9) et pumilo [sic!} (4) Keys. en sont peut-être synony- mes”. Roewer (1954: 1026) and Bonnet (1956: 1816, 1817) followed him as to pauperatus and pumilio, but retained albescens as a good species. À comparison of our specimens with the descriptions, figures and type specimens (Hamburg Museum) of H. albescens from Rockhampton (Queensland) and H. pau- peratus from Port Mackay (Queensland) have convinced me that both are synonyms of bleekeri. Keyserling’s Fig. la of epigyne of albescens suggests a pointed anterior border; in the type, however, this border is regularly curved as in our Fig. 35 of bleekeri; the epigyne of pauperatus is identical with this figure. The abdomens of both type specimens are in rather bad condition and do not give further indications. H. pumilio Keyserling (Arachn. Austral. 1 (2) [1881]: 1317, PI. 112 Fig. 3) from Peak Downs (Queensland), holotype in the Hamburg Museum, belongs without any doubt to a different species and almost certainly to a different genus: e.g., the palp resembles that of Dendryphantes laticeps Strand (Fig. 87). In his discussion of the genus Plotius, near to Euryattus, Simon (1903: 818) wrote: ‘Jen connais deux espèces: P. curtus E. Sim. de l’île Halmahera et breviusculus E. Sim. de Ceylon, et je lui rapporte le Hasarius chrysostomus Keys. du Queensland”. He cer- tainly did not see the male types of this species (three specimens, Hamburg Museum — the female is lost): they do not possess the characters of Plotius but clearly those of Euryattus, e.g. the characteristic dentation of the cheliceral furrow (Simon, loc. cit.: 812, Fig. 962); the male palp is almost identical with that of Ewryattus senex (Simon, loc. cit.: 812, Fig. 961). After having studied Keyserling’s description and figures of H. chrysostomus and the male types from Rockhampton I am certain that also this species is a synonym of E. bleekeri. The type locality of bleekeri is Amboina; its area extends from that island to Australia (Roewer, 1954: 1026; Bonnet, 1956: 1817). 58 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 NEN fi Le Fig. 41, 42. Euryattus bleekeri (Doleschall), 9, vulva. Fig. 43—45. Trite longula (Thorell). 43, 4; 44, id. left palp, ventral view; 45, id. left palp, lateral view. Fig. 46—50. Sandalodes bernsteini Thorell. 46, 4; 47, id. lateral view; 48, 49, id. left palp, ventral view; 50, id. chelicerae, frontal view. Fig, 51. Bathippus macrognathus (Thorell), 6. Fig. 52—54. B. papuanus Thorell. 52, 4; 53, id. left palp, lateral view; 54, id. ventral view. Fig. 41, 42: X 60; 43, 46, 47: X 7; 44, 45, SZ RZ NO SE TOS SKS CSSS SES Ses 0 FR. CHRYSANTHUS : Spiders from South New Guinea X 59 Euryattus porcellus Thorell, 1881 Fig. 37—40 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 660, 9 4. Simon, 1903, Hist. nat. Araignées 2: 813, Fig. 959, 9 4. A female (14 mm) and a male (13 mm) from Merauke (1956—1957) fully agree with Thorell’s description and his type material in the Museo Civico di Storia Naturale, Genoa. The type locality is Yule I., and the species has been found on New Guinea only (Roewer, 1954: 1027; Bonnet, 1956: 1817). Trite Simon, 1885 Trite longula (Thorell, 1881) Fig. 43—45 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 454, & (Marptusa). A male of this species (7 mm) from Merauke (1956—1957) is in all details identical with the holotype from Cape York, North Australia (Genoa Museum); the species was also recorded from Lord Howe I. (Rainbow, Rec. S. Austr. Mus. 1 [1920]: 267). UNIDENTATI Hyllinae Sandalodes Keyserling, 1883 Sandalodes bernsteini (Thorell, 1881) Fig. 46—50 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 619, & (Plexippus). The description of this species was based on three males, two from Andai (Vogelkop, New Guinea) and one from the Aru Is.; it had not been found since and the female is still unknown (Roewer, 1954: 1067; Bonnet, 1957: 2246 [Hyllus}). A male (5 mm) from Mindiptana (1958) is conform to the description and the types in the Genoa Museum. Plexippinae Bathippus Thorell, 1892 Bathippus macrognathus (Thorell, 1881) Bigs, 51, 55 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 531, & (Plexippus). Six males (8—10 mm) from Merauke (1956—1957) correspond with Thorell’s description and his type material from Fly River (South New Guinea) and the Aru Is. (Genoa Museum); the species was also collected on the Kei Is. (Roewer, 1954: 1076; Bonnet, 1955: 854). The female is still unknown but it may have been described as a separate species, probably even in another genus, 60 DEEL 111, AFL. 2, 1968 TIJDSCHRIFT VOOR ENTOMOLOGIE, Pisi 4a eG aß si Fig. 56—60. Palpelius A) beccarii (Thorell). 56, 9; 57, id. epigyne; 58, id. vulva; 59, &, left palp, ventro-lateral view; Fig. 55. Bathippus macrognathus (Thorell), &, left palp, ventral view. 60, id. ventral view, Fig. 61—63. Plexippus paykullii (Audouin). 61, 9; 62, id. epigyne; 63, &. Fig, SONO UD 36: 0141635 CT STD OMS 8:0 12059 X 2062: Fr. CHRYSANTHUS : Spiders from South New Guinea X 61 Bathippus papuanus (Thorell, 1881) Fig. 52—54 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 526, ® & (Plexippus monstrouzieri Lucas var. papuana). Roewer, 1938, Mém. Mus. Hist. nat. Belg. (hors série) 3 (19): 87, Fig. 67—69, &. According to Simon (1903, Hist. nat. Araignées 2: 731) Thorell’s var. papuana differs from monstrouzieri Lucas, 1869. The type material (several males, one female, Genoa Museum) originated from the Aru Is. and Fly River; the species has also been recorded from Manoi (Solomon Is.); the type locality of monstronzieri is New Cale- donia (Roewer, 1954: 1077; Bonnet, 1955: 855). One male (9 mm) from Mindiptana (1965) is in agreement with Thorell’s des- cription and type material, and with the figures given by Roewer. Palpelius Simon, 1903 Palpelius beccarii (Thorell, 1881) Fig. 56—60 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 582, 9 4 (Plexippus). , 1881, ibid.: 588, 9 (P. dearmatus) syn. nov. Simon, 1903, Hist. nat. Araignées 2: 735, Fig. 842, &. Thorell described two closely related species, viz. Plexippus beccarii and P. dearmatus: the first species, several males and a few females, from Ternate, Ceram, New Guinea (Andai, Ramoi and Fly River), Yule I., Aru Is. and Somerset (Cape York, Australia); the second species, some females only, from Yule I., Aru Is. and Somerset. His des- cription of dearmatus is very short; apart from a few colour differences he saw small differences in the epigyne only: (1) in dearmatus the two egg-shaped parts are united over nearly their whole length, in beccarii they do not touch each other or do so along the anterior half only, (2) in dearmatus the width of the receptacula is about 1.5 times that of the egg-shaped parts, whereas in beccarii the width of both is equal. About dearmatus, however, he remarked: “‘vix modo varietas est P. beccarit”’ (it is scarcely a variety of P. beccarii). Our collection contains three females from Merauke (1956-1957), four from Min- diptana (1958, 1965) (10—11 mm) and three males (8—9 mm) from Mindiptana (1958, 1965). I could compare them with type specimens in the Museo Civico di Storia Naturale, Genoa. The males are identical with the beccarii males. As to the females: the egg-shaped parts of the epigyne are as in dearmatus, or as in beccarii ot they may show features of both (cf. Fig. 57, 58): the width of the receptacula varies from 1.1 to 1.5 times the width of the egg-shaped parts; the females from Merauke do not differ from those from Mindiptana. P. dearmatus, therefore, must be considered a synonym of beccarii. The species has not been found outside the region mentioned above (Roewer, 1954: 1084; Bonnet, 1958: 3303). Plexippus C. L. Koch, 1846 Plexippus paykullii (Audouin, 1827) Fig. 61—65 Audouin, 1827, Explic. Planch. Arachn. Savigny Descr. Egypt. (2) 22: 409, PI. 7 Fig. 22, 4 (Attus). Doleschall, 1859, Act. Soc. Sci. Ind.-Neerl. 5: 14, PI. 9 Fig. 5, ® (Salticus culicivorus). 62 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 JN \ SS Fig. 64, 65. Plexippus paykullii (Audouin). 64, 2, vulva; 65, 4, left palp, ventral view. Fig. 66—69. Zenodorus durvillii (Walckenaer). 66, 9; 67, id. epigyne; 68, id. vulva; 69, &, left palp, ventral view. Fig. 70. Mopsus mormon Karsch, 6, left palp, ventral view. Fig. 64: X 50; 65, 69, 70: X 40; (SR) << 7/8 (GS DK DI 68: X 9120 Fr. CHRYSANTHUS : Spiders from South New Guinea X 63 Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 501, 9 3 (Menemerus). Simon, 1903, Hist. nat. Araignées 2: 711, 734, Fig. 839—841, 28. , 1937, Arachn. France 6 (5): 1242, 1271, Fig. 2023, 2024, 28. As this species is nearly cosmopolite there are many more useful descriptions and figures (cf. Roewer, 1954: 1086; Bonnet, 1958: 3717). Doleschall’s lan does not contain the type of his Salticus culicivorus. The epigyne is somewhat variable: when the skin is lightly sclerotized the whole vulva (Fig. 64) is discernible, when it is heavily sclerotized the epigyne consists of a dark brown shield without details; usually the vulva is more or less discernible. Our collection contains several males (6—9 mm) and females (8—10 mm) from Merauke (1956—1957) and Mindiptana (1958—1965). Zenodorus Peckham, 1886 Zenodorus durvillii (Walckenaer, 1937) Fig. 66—69 Walckenaer, 1837, Hist. natur. Ins. Apt. 1: 459, & (Attus). Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 653, 9 & (Ephippus). Keyserling, 1883, Arachn. Austral. 1 (2): 1422, PI. 120 Fig. 4, 5, 9 & (Ephippus). Squamiform hairs with a pink gloss, scattered over cephalothorax and abdomen, render durvillii a beautiful species. It is known from New Guinea and adjacent islands and from Australia (Roewer, 1954: 1108; Bonnet, 1959: 4960), and it is common in the environs of Merauke and Mindiptana: many females (9—11 mm) and males (7—10 mm). Thyeninae Mopsus Karsch, 1878 Mopsus mormon Karsch, 1878 Fig. 70—74 Karsch, 1878, Mitt. Münchn. Entom. Ver. 2: 31, 4. Thorell, 1881, Ann. Mus. civ. stor. nat. Genova 17: 462, 9 4. Simon, 1903, Hist. nat. Araignées 2: 686, Fig. 816—818, 9 4. In his “Katalog der Araneae” Roewer suggested that the male only was described (1954: 1109) though he mentioned Simon, who gave a description of the female and added: “très répandue en Nouvelle Guinée et dans le Nord de l'Australie” (1903: 686). There are many females (12—15 mm) and males (12—15 mm) in my collection, all from Merauke (1956—1957), only one male from Mindiptana (1965). The species was recorded from several places in New Guinea, adjacent islands and Cape York (Australia) (Thorell, loc. cit.: 466; Roewer, 1954: 1109; Bonnet, 1957: 2985). In Queensland and New South Wales the closely related M. penicillatus (Karsch, 1878) is found. Strongly contrasting with the female, where the cephalothorax is brownish yellow and the abdomen is yellowish white, the male is beautifully coloured: the abdomen is yellowish white with two black longitudinal stripes like in the female (Strand [1911, Abb. senckenb. naturf. Ges. 34: 185} remarked that in living specimens the abdomen is green). The cephalothorax is orange, eye region lively red, frontal part of ‘the cephalothorax, chelicerae and underside of the stout first legs dark wine-red, the femora 64 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Fig. 7174. Mopsus mormon Karsch. 71, ®; 72, id. epigyne; 73, id. vulva; 74, 4. Fig. 75—77. Poecilorchestes decoratus Simon. 75, ® ; 76, id. lateral view; 77, id. epigyne. Fig. 78—81. Cosmo- phasis bitaeniata (Keyserling). 78, ® ; 79, id. epigyne; 80, 4, left palp, ventral view; 81, id. tibial apophyses, lateral view. Fig. 71, 742. 2@ 3:12,79 312x170; 732 1205 752170 TI 1503 738 XK 715280 K-40 FR. CHRYSANTHUS : Spiders from South New Guinea X 65 with some squamiform white hairs, the tips of the chelicerae and the greater part ot the first legs orange; the anterior median eyes have a greenish lustre. A rather broad fringe of snow-white hairs extends from the lateral eyes to about the middle of the cephalothorax, whereas long black hairs form a tuft between the anterior lateral eyes. Coccorchestinae Poecilorchestes Simon, 1901 Poecilorchestes decoratus Simon, 1901 Fig. 75-77 Simon, 1901, Hist. nat. Araignées 2: 648, Fig. 765—767, &. , 1902, Ann. Soc. entom. Belg. 46: 34, 6. A comparison of a female of this family (Mindiptana, 1965) with the description and figures given by Simon and with the male type from Dorey (—Manokwari, North New Guinea) in the Muséum national d’Histoire naturelle, Paris, have convinced me that it belongs to the same species. The remarkable shape of the cephalothorax, the position of the eyes and of the striking snow-white patches on cephalothorax, abdomen and legs, caused by squamiform hairs, the very rugose skin of the cephalothorax: all these characters are common to both; the colour only is somewhat different: the male is dark reddish brown, in the female there is a purplish hue over the dark brown colour. The measurements of the female are: cephalothorax, length 1.7 mm, width 1.6 mm; abdomen, length 1.9 mm, width 1.6 mm; legs. I 2.5, II 2.3, III 2.1, IV 2.6 mm. Epigyne: Fig. 77. Heliophaninae Cosmophasis Simon, 1901 Cosmophasis bitaeniata (Keyserling, 1882) Fig. 78—81 Keyserling, 1882, Arachn. Austral. 1 (2): 1365, PI. 115 Fig. 8, 4 (Sobara). , 1882, ibid.: 1374, PI. 116 Fig. 5, ® (Selaophora rubra). My collection contains a couple of this beautifully orange-coloured species (Merauke, 1956-1957); the male (8 mm) is identical with the holotype of Sobara bitaeniata from the environs of Sydney (Hamburg Museum). The female paratype is juvenile (“Das leider unentwickelte Weib’); in his description Keyserling mentions Tafel 115 Fig. 9 “femina, Cephalothorax”, but there is no Fig. 9 on this plate. The holotype ot Selaophora rubra from Cape York is lost; the female in my collection (7 mm) fully agrees with Keyserling’s description and figures. The species is also known from the Aru Is. (Roewer, 1954: 1152; Bonnet, 1956: 1241). Strand (Abh. senckenb. naturf. Ges. 34 [1911]: 180, PI. 4 Fig. 22, Pl. 6 Fig. 84) described C. orsimoides from the Kei Is.; the female holotype (SMF 2436) is smaller than C. bitaeniata, viz., 5 mm; it seems to me, however, that the epigynes are identical; a study of the vulvae, which are only partly discernible through the skin, could give certainty. The male of orsimoides is unknown. 66 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Fig. 82—85. Cosmophasis micarioides (L. Koch). 82, 2; 83, id. epigyne; 84, id. vulva; 85, 4, left palp, ventral view. Fig. 86—88. Dendryphantes laticeps Strand. 86, & ; 87, id. left palp, ventral view; 88, @, epigyne. Fig. 89—92. Menemerus bivittatus (Dufour). 89, 9; 90, id. epigyne; 91, 92; &, abdomen. Fig. 82, 86, 89, 91, 92: X 7; 83: X 150; 84: X 120; 85, 87: x 40; 88; 90: x70 FR. CHRYSANTHUS : Spiders from South New Guinea X 67 Cosmophasis micarioides (L. Koch, 1880) Fig. 82—85 L. Koch, 1880, Arachn. Austral. 1 (2): 1178, PI. 102 Fig. 3, 4 (Amycus). Strand, 1911, Abh. senckenb. naturf. Ges. 34: 180, Pl. 4 Fig. 16, PI. 6 Fig. 85, 2. There is a large series of females (5—7 mm) and males (5—7 mm) in my col- lection from Merauke (1956—1957) and Mindiptana (1958—1965); the males agree with Koch’s description and figures and with the male holotype in the Zoologisches Museum, Hamburg; the females are identical with several females, identified by Strand, in the Senckenberg Museum, Frankfurt a.M. The colour of the female is rather variable: it may be multicoloured, viz., orange, yellow, black and silver-coloured, sometimes these colours have partly faded away, or even the greater part of the coloured squamiform hairs are rubbed off, resulting in an almost uniform tawny colour of the abdomen. Koch’s figure of the male does not give a good idea of its colour-pattern: the contrast between the dark and the lighter parts is much too strong. In reality there is only a faint silver-coloured bloom in the middle and on the sides of the dark greyish brown abdomen and some greenish and purplish metallic lustre on the cephalothorax. The colours may for the greater part, have disappeared, as in the female. Strand’s C. maculiventris (Abb. senckenb. naturf. Ges. 34 [1911]: 180, PI. 4 Fig. 23 — SMF 2433) from Terangan, Aru Is., is almost certainly a synonym of micarioides: (1) colours and pattern are the same, (2) the characteristic light spots on the under- side of the abdomen are discernible in nearly all specimens of our collection, especially in the vividly coloured ones, (3) the not yet completely developed epigyne (“Die nicht reife Epigyne erscheint in Flüssigkeit als ein weisslicher, vorn gerundeter Querfleck, der mehr als doppelt so breit wie lang ist und durch eine schwarze Mittellängslinie geteilt zu sein scheint’) fits very well the developed form. The type locality of micarioides is Port Mackay (Queensland) and the species has also been recorded from other parts of Australia, from New Guinea and adjacent islands (Roewer, 1954: 1153; Bonnet, 1956: 1243). Dendryphantinae Dendryphantes C. L. Koch, 1837 Dendryphantes laticeps Strand, 1911 Fig. 86—88 Strand, 1911, Abh. senckenb. naturf. Ges. 34: 183, PI. 4 Fig. 25, PI. 6 Fig. 91, 4. Two males (7 and 8 mm) from Merauke, 1956—1957, fully correspond with Strand’s description and figures and with the holotype from Terangan, Aru Is. (SMF 2452). The species is also known from the Kei Is. (Roewer, 1954: 1190; Bonnet, 1956: 1396); the female, however, had not yet been found. A female, also from Merauke, certainly belongs to the same species. In nearly all details it is identical with one of the two males (Fig. 86): cephalothorax and all appen- dices are dark wine-red, palpi and first legs are flattened, the sternum bears a blunt knob, the abdomen is flattened and the sides are wrinkled. The colour of the abdomen is lighter, but in this respect the female rather resembles the other male, where all colours are lighter. Measurements of the female: cephalothorax, length 2.7 mm, width 68 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 2.4 mm; abdomen, length 4.0 mm, width 2.5 mm; legs I 5.6, II 4.3, III 3.5, IV 4.5 mm. Epigyne: Fig. 88. Marpissinae Menemerus Simon, 1868 Menemerus bivittatus (Dufour, 1831) Fig. 89—95 Dufour, 1831, Ann. Sci. nat. 22: 369, PI. 11 Fig. 5, 2 (Salticus). Doleschall, 1859, Act. Soc. Sci. Ind.-Neerl. 5: 15, Pl. 9 Fig. 4, 2 (Salticus convergens). Thorell, 1878, Ann. Mus. civ. stor. nat. Genova 13: 232, 9 & (Icius convergens). F. Pickard-Cambridge, 1901, Arachnida #2 Biol. Centr. Amer. Zool.: 250, Pl. 21 Fig. 18, 19, Q & (Marpissa melanognatha). Simon, 1937, Arachn. France 6 (5): 1210, 1262, Fig. 1932, 1933, 2 4. Berland & Millot, 1941, Mém. Mus. Hist. nat. Paris 12: 246, Fig. 49, 50 A-C, 2 4. There is a long list of synonyms and there exist several other useful descriptions and figures of this nearly cosmopolite species (cf. Roewer, 1954: 1263; Bonnet, 1957: 21.64.) In our collection there are some 20 females (8—10 mm) and four males (6—8 mm) from Merauke (1956—1957) and Mindiptana (1959, 1965). In the female the pattern of the abdomen is often less distinct than in our Fig. 89: the dark brown hairs are easily rubbed off. As the epigyne is surrounded by many hairs and sometimes partly filled with a resinous concretion (‘Begattungszeichen”), its shape may be not as clear as in our Fig. 90. In the abdomen of the male the lateral streaks are lighter than in the female, whereas the central streak is darker (Fig. 91, 92). The bulbus of the male palp may be smooth or with a more or less deep groove (Fig. 94, 95; cf. Pickard- Cambridge, loc. cit. and Berland & Millot, loc. cit.). Fig. 93—95. Menemerus bivittatus (Dufour). 93, 9, vulva (X 60); 94, 95, &, left palp, ventral view (X 40) FR. CHRYSANTHUS : Spiders from South New Guinea X 69 Concluding remarks The collection made by Br. Monulf in the environs of Merauke (1956—1957) and Mindiptana (1958-1965) contains about 200 species. In the present series of papers (Chrysanthus, 1958—1968) 151 species are dealt with; 110 are known species, 41 are described as new to science of Arachnology. Of 19 species I could describe the male, hitherto unknown, of 6 species I could give a description of the unknown female. At least 15 species were not yet recorded from New Guinea; 21 species were recorded for the first time after their original descriptions, these descriptions being published 50, or even 80 years ago. This fact is not so surprising because after the publications by Kulczynski, Strand and Hogg, all between 1911 and 1915, scarcely any collecting, followed by publications, occurred in New Guinea: I, therefore, do not agree with Bonnet (1961: 438) who seems to doubt the validity of species, described from a single specimen so many years ago, if no specimens have been found since. About 50 species of the collection have not yet been discussed: part of them have been obtained after my publication of the study of their group, the others need further investigation. I hope to include them in a new series of papers on New Guinean Spiders, dealing with material from other collections. 70 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 LIST OF SPECIES discussed in “Spiders from South New Guinea”, I—X The signs Qn, dn denote first description of female or male, respectively; NG, means first records for New Guinea; Ch., means Chrysanthus, referring to the papers listed under References, below. ECRIBELLATAE SICARIIDAE Scytodes fusca Walckenaer, 1837. Ch., 1967b: 91, @ 8 S. longipes Lucas, 1845. Ch., 1967b: 89, 2 4 S. pallida Doleschall, 1859. Ch., 1967b: 91, 2, NG S. tardigrada Thorell, 1881. Ch., 1967b: 91, 9, NG PHOLCIDAE Artema atlanta Walckenaer, 1837. Ch., 1967b: 92, 9 & Crossopriza lyoni (Blackwall, 1867). Ch., 1967b: 96, 2 6, NG Psilochorus nigromaculatus Kulczynski, 1911. Ch., 1967b: 96, 9n (second record) ZODARIIDAE Storena zebra Thorell, 1881. Ch., 1967b: 98, & THERIDIIDAE Achaeranea decorata (L. Koch, 1867). Ch., 1963: 743, 9, NG A. hammeni Chrysanthus, 1963. Ch., 1963: 746, 9 A. krausi Chrysanthus, 1963. Ch., 1963: 744, 2 8 A. meraukensis Chrysanthus, 1963. Ch., 1963: 746, 9 & A. mundula (L. Koch, 1872) papuana Chrysanthus, 1963. Ch., 1963: 741, 9 Argyrodes argentatus Cambridge, 1880. Ch., 1963: 739, 9 A. fissifrons Cambridge, 1869. Ch., 1963: 737, & A. flavescens Cambridge, 1880. Ch., 1963: 739, 9 6 A. kulczynskii Roewer, 1942 (= A. argenteolus Kulczynski, 1911). Ch., 1963: 741, 2 gn (second record) A. miniaceus (Doleschall, 1857). Ch., 1963: 739, 9 & Latrodectus geometricus C. L. Koch, 1841. Ch., 1963: 737, 9 Lithyphantes bertkaui (Thorell, 1881). Ch., 1963: 737, ® &n (second record) Phoroncidia levii Chrysanthus, 1963. Ch., 1963: 735, 2 Theridion diadematum Chrysanthus, 1963. Ch., 1963: 749, 9 T. rufipes Lucas, 1846. Ch., 1963: 748, 9 & FR. CHRYSANTHUS : Spiders from South New Guinea X Vil ARGYOPIDAE Anepsion wichmanni (Kulczynski, 1911). Ch., 1961: 207, ® &n (second record) Arachnura melanura Simon, 1867. Ch., 1961: 203, 9, NG Araneus cyrtarachnoides (Keyserling, 1887). Ch., 1960: 36, 9 3 . dehaani (Doleschall, 1859). Ch., 1960: 31, 2 . granti Hogg, 1914. Ch., 1960: 36, ® gn (second record) . inustus (L. Koch, 1871). Ch., 1960: 41, 9 gn, NG . laglaizei (Simon, 1877). Ch., 1960: 39, 9 . lugubris (Walckenaer, 1842). Ch., 1960: 36, 9 & . rufofemoratus (Simon, 1884). Ch., 1960: 33, 9 4 . simillimus Kulczynski, 1911. Ch., 1961: 199, 9 . theisi (Walckenaer, 1842). Ch., 1960: 39, 9 4 . transmarinus (Keyserling, 1865) (= productus L. Koch, 1867). Ch., 1960: 30, 2 & Argyope aetherea (Walckenaer, 1842). Ch., 1958: 237, 242, 9 gn A. catenulata (Doleschall, 1859). Ch., 1958: 240, 9 4 A. chloreides Chrysanthus, 1961. Ch., 1961: 197, 9 A. mindiptanensis Chrysanthus, 1961. Ch., 1961: 195, 9 A picta Tt Koch: 1871 Eh’, 1953:7237,2242,, 975 Carepalxis tricuspidata Chrysanthus, 1961. Ch., 1961: 205, 9 Cyclosa camelodes (Thorell, 1878). Ch., 1961: 201, 9 C. insulana (Costa, 1834). Ch., 1961: 199, 9 4 C. micula (Thorell, 1892). Ch., 1961: 203, 9 C. mulmeinensis (Thorell, 1887). Ch., 1961: 203, 9 C. velata Chrysanthus, 1961. Ch., 1961: 201, ® ¢ Cyrtarachne friederici Strand, 1911. Ch., 1961: 209, 9 C. tricolor (Doleschall, 1859). Ch., 1961: 209, 9 Cyrtophora beccarii (Thorell, 1878). Ch., 1960: 25, 9 4 C. cicatrosa (Stoliczka, 1869). Ch., 1960: 28, 9 gn . cylindroides (Walckenaer, 1842). Ch., 1959: 200; 1960: 25, ® & . moluccensis (Doleschall, 1857). Ch., 1959: 199, 9 . monulfi Chrysanthus, 1960. Ch., 1960: 26, 9 . unicolor (Doleschall, 1857). Ch., 1959: 201, 2 Gasteracantha brevispina (Doleschall, 1857). Ch., 1959: 203; 1960: 25, 9 gn G. taeniata (Walckenaer, 1842). Ch., 1959: 202; 1960: 23, 9 gn G. theisi (Guérin, 1838). Ch., 1959: 203, 9 3 Larinia phthisica (L. Koch, 1871). Ch. 1961: 205, 9, NG I tavida, (L. Koch, 1871). Ch, 1961:7205, 2 Nephila maculata (Fabricius, 1793). Ch., 1959: 197; 1960: 23, 2 4 N. malabarensis (Walckenaer, 1842). Ch., 1959: 199, 9 Poecilopachys verrucosa (L. Koch, 1871). Ch., 1961: 209, 9 Poltys illepidus C. L. Koch, 1843. Ch., 1961: 211, 9 P. sigillatus Chrysanthus, 1961. Ch., 1961: 211, 9 SA AA à à À à A SESIA TETRAGNATHIDAE Eucta javana Thorell, 1890. Ch., 1963: 733, 9 Leucauge argentata (Cambridge, 1869) marginata Kulczynski, 1911. Ch., 1963: 731, 9 72 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 L. granulata (Walckenaer, 1842). Ch., 1963: 731, 9 & L. grata (Guérin, 1838). Ch., 1963: 729, 4 Mesida humilis Kulczynskki, 1911. Ch., 1963: 727, ® &n (second record) Tetragnatha mandibulata Walckenaer, 1842. Ch., 1963: 733, 9 & T. rubriventris Doleschall, 1857. Ch., 1963: 735, 9 & Tylorida striata (Thorell, 1877). Ch., 1963: 727, 9. PISAURIDAE Dolomedes chroesus Strand, 1911. Ch., 1967a: 422, 9 &n (second record) D. facetus L. Koch, 1876. Ch., 1967a: 423, 9 4, NG Perenethis unifasciata (Doleschall, 1859). Ch., 1967a: 421, 9 4 LYCOSIDAE Anomalomma cinctipes Simon, 1898. Ch., 1967a: 425, 9 A. muju Chrysanthus, 1967. Ch., 1967a: 425, 9 & Artoria parvula Thorell, 1877. Ch., 1967a: 423, 9 Varacosa hickmani Roewer, 1954. Ch., 1967a: 423, 2 4 n, NG (second record) V. papakula (Strand, 1911). Ch., 1967a: 424, & Sn (second record) V. tanna (Strand, 1913). Ch., 1967a: 424, 9, NG OXYOPIDAE Oxyopes macilentus L. Koch, 1878. Ch., 1967a: 419, 9 4 O. papuanus Thorell, 1881. Ch., 1967a: 419, 9 & O. tapponiformis Strand, 1911. Ch., 1967a: 421, 9 (second record) CLUBIONIDAE Chetracanthium lanceolatum Chrysanthus, 1967. Ch., 1967a: 407, 9 C. marplesi Chrysanthus, 1967. Ch., 1967a: 403, 9 & C. wiehlei Chrysanthus, 1967. Ch., 1967a: 405, 9 & Clubiona ericius Chrysanthus, 1967. Ch., 1967a: 409, 9 & . kowong Chrysanthus, 1967. Ch., 1967a: 407, 9 4 meraukensis Chrysanthus, 1967. Ch., 1967a: 411, 9 2 . pantherina Chrysanthus, 1967. Ch., 1967a: 413, 9 . papuana Chrysanthus, 1967. Ch., 1967a: 413, 9 . procera Chrysanthus, 1967. Ch., 1967a: 415, 9 subula Chrysanthus, 1967. Ch., 1967a: 415, 4 Matidia chlora Chrysanthus, 1967. Ch., 1967a: 415, @ M. muju Chrysanthus, 1967. Ch., 1967a: 417, 4 9 Da aa EUSPARASSIDAE Clastes freycineti Walckenaer, 1837. Ch., 1965: 369, 2 8 Heteropoda atriventris Chrysanthus, 1965. Ch., 1965: 356, 9 H. erythra Chrysanthus, 1965. Ch., 1965: 362, ® & H. fusciventris Chrysanthus, 1965. Ch., 1965: 366, 2 & FR. CHRYSANTHUS : Spiders from South New Guinea X 73 H. mindiptanensis Chrysanthus, 1965. Ch., 1965: 359; 1967a: 403, & H. rubra Chrysanthus, 1965. Ch., 1965: 360, 2 4 H. venatoria (Linné, 1767). Ch., 1965: 356, 9 & Isopeda goliath Chrysanthus, 1965. Ch., 1965: 348, 9 I. meraukensis Chrysanthus, 1965. Ch., 1965: 346, 2 & Olios artemis Hogg, 1915. Ch., 1965: 353; 1967a: 401, 9 n (second record) O. fimbriatus Chrysanthus, 1965. Ch., 1965: 355; 1967a: 403, 9 O. insignifer Chrysanthus, 1965. Ch., 1965: 353, 9 3 O. rubriventris (Thorell, 1881). Ch., 1965: 351; 1967a: 401, 9 Pandercetes isopus Thorell, 1881. Ch., 1965: 368, 9 gn Thelcticopis cuneissignata Chrysanthus, 1965. Ch., 1965: 349, 9 THOMISIDAE Borboropactus divergens (Hogg, 1914). Ch., 1964: 89, 2 (second record) Cymbacha saucia L. Koch, 1874. Ch., 1964: 95, 9, NG Diaea limbata Kulczynski, 1911. Ch., 1964: 102, 9 4 (second record) D. sticta Kulczynski, 1911. Ch., 1964: 102, 9 4 n (second record) Hedana bonneti Chrysanthus, 1964. Ch., 1964: 87, 9 Lycopus atypicus Strand, 1911. Ch., 1964: 92, @ (second record) Misumena alpha Chrysanthus, 1964. Ch., 1964: 97, 9 M. beta Chrysanthus, 1964. Ch., 1964: 99, 9 M. gamma Chrysanthus, 1964. Ch., 1964: 99 9 M. lorentzi Kulczynski, 1911. Ch., 1964: 95, 9, misidentified Philodromus planus (L. Koch, 1875). Ch., 1964: 91, 3 n, NG Runcinia acuminata (Thorell, 1881). Ch., 1964: 99, 9, NG Senoculifer dentibulbis Balogh, 1936. Ch., 1964: 91, & (second record) Stephanopis monulfi Chrysanthus, 1964. Ch., 1964: 89, 9 Tharrhalea albipes L. Koch, 1875. Ch., 1964: 9 4, NG Thomisus spectabilis Doleschall, 1859. Ch., 1964: 102, 9 & Tmarus homanni Chrysanthus, 1964. Ch., 1964: 94, 9 SALTICIDAE Bathippus macrognathus (Thorell, 1881). Ch., 1968: 59, 4 B. papuanus (Thorell, 1881). Ch., 1968: 61, 4 Bavia aericeps Simon, 1877. Ch., 1968: 51, 9 4 Cosmophasis bitaeniata (Keyserling, 1882). Ch., 1968: 65, 2 4 C. micarioides (L. Koch, 1880). Ch., 1968: 67, 2 & Cytaea frontaligera (Thorell, 1881). Ch., 1968: 53, 9 4 C. mitellata (Thorell, 1881). Ch., 1968: 55, & (second record) C. nimbata (Thorell, 1881). Ch., 1968: 53, & Qn (second record) Dendryphantes laticeps Strand, 1911. Ch., 1968: 67, 3 Qn Diolenius amplectens Thorell, 1881. Ch., 1968: 51, 9 gn Euryattus bleekeri (Doleschall, 1859). Ch., 1968: 55, 9 4 E. porcellus Thorell, 1881. Ch., 1968: 59, 9 4 Linus fimbriatus (Doleschall, 1859). Ch., 1968: 49, 9 Menemerus bivittatus (Dufour, 1831). Ch., 1968: 68 9 & 74 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 Mopsus mormon Karsch, 1878. Ch., 1968: 63, 9 4 Palpelius beccarii (Thorell, 1881). Ch., 1968: 61, 9 4 Plexippus paykullii (Audouin, 1827). Ch., 1968: 61, 9 & Poecilorchestes decoratus Simon, 1901. Ch., 1968: 65, gn Sandalodes bernsteini (Thorell, 1881). Ch., 1968: 59, & (second record) Trite longula (Thorell, 1881). Ch., 1968: 59, 4, NG Zenodorus durvillii (Walckenaer, 1837). Ch., 1968: 63, 9 8 CRIBELLATAE Deinopis camela Thorell, 1881. Ch., 1967b: 100, 9 gn (second record) Fecenia angustata (Thorell, 1881). Ch., 1967b: 102, 9 gn F. buruana Reimoser, 1936. Ch., 1967b: 104, 9 ¢ F. maforensis Simon, 1906. Ch., 1967b: 104, 9 Psechrus argentatus (Doleschall, 1857). Ch., 1967b: 105, 9 Titanoeca fulmeki Reimoser, 1927. Ch., 1967b: 102, 9 gn (second record) Uloborus albofasciatus Chrysanthus, 1967. Ch., 1967b: 98, 9 & U. geniculatus (Olivier, 1789). Ch., 1967b: 98, 9 3 U. hilaris Simon, 1906. Ch., 1967b: 98, 2 REFERENCES Bonnet, P. 1945—1961. Bibliographia Araneorum. Toulouse, Chrysanthus, Fr. 1958. Spiders from South New Guinea I. Nova Guinea, new series 9: 235—243. 1959. Id. II. lc. 10: 197—206. 1960. Id. III. Nova Guinea. Zoology, no. 3 : 21—42. 1961. Id. IV. Lc. no. 10: 195—214. 1963. Id. V. Lc. no. 24: 727—750. 1964. Id. VI. Ic. no. 28: 87—104. 1965. Id, VII. l.c. no. 34: 345—369. 1967a. Id. VIII. Ic. no. 37: 401-426. 1967b. Id. IX. Tijdschr. Ent. 110: 89—105. 1968, Id. X. lc. 111 (this paper). C. F. 1942. Katalog der Araneae. I. Bremen. 1954. Id. II. Bruxelles. IN M M vo w M SI SI vo ir Hd © oO 4 5 an A Die head DEEL 111 AFLEVERING 3 1968 TIJDSCHRIFT VOOR ENTOMOLOGIE UITGEGEVEN DOOR DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING MUS. COMP. ZOOL. LIBRARY JUL 25 1968 HARVARD UNIVERSITY INHOUD : F. LukoscHus. — Neue Krätzmilben von einheimischen Insektivoren (Psorergatidae : Trombidiformes), p. 75—88, Abb. 1-41. Tijdschrift voor Entomologie, deel 111, afl. 3 Gepubliceerd 21-VI-1968 NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING BESTUUR (BOARD) Voorzitter (Chairman) . MIO Gay Barendrecht Vice-Voorzitter (Vice- President) BU art ny MER Wiebes Secretaris (Secretary) . . . . .:. . . W. Hellinga Address mmm: . . « . . Weesperzijde 23 II, Amsterdam-O. Penningmeester er) DER NE SO WORST Ma DOS Address, St NE Wann Baenlest naats 2 Ovis Amsterdam Bibliothecaris natans) A TOEN eN le Cera Weinfeekel Address VON tenn aen eh Zeeburserdmkr 21, Amsterdam-Q: Pedenu (Members) ) 9) . 0 a0 une eae mjanse,, ANSE ET sBesemen AFDELING VOOR TOEGEPASTE ENTOMOLOGIE (DIVISION OF APPLIED ENTOMOLOGY) BESTUUR (BOARD) Wooezitter | (Chairman) AN ay EN PA CE SH NBesermnen Seetetaris (Secretary) NV wile at) ce aa te ba Grays Address... .... . +. … Nedereindsestraat 35, Kesteren. Leden (Members) . . . . . . . . . L. Bravenboer, J. J. Laarman, J. B. M. van Dinther Publicaties van de Vereeniging (Publications of the Society) Subscription may be obtained from all booksellers or directly from the Librarian, Zeeburgerdijk 21, Amsterdam-O., except for Entomologia Experimentalis et Applicata, which is available through booksellers or from the Noord-Holland Editing Co., Post Office Box 103, Amsterdam. TIJDSCHRIFT VOOR ENTOMOLOGIE Redactie (Editing Board) . . . . . . Pater Chrysanthus, A. Diakonoff, C. A. W. Jeekel, M. A. Lieftinck, J. T. Wiebes Address . . . 2 . . 2 + + + + Rijksmuseum van Natuurlijke Historie, Raamsteeg 2, Leiden. The Journal serves the publication of papers on Insecta, Myriapoda and Arachnoidea. It appears in separate issues, forming an annual volume of 350—400 pages. Subscription rate: D.Fl. 45.— (£ 4.10.—, $12.50) per volume. MONOGRAFIEEN VAN DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING Redactie (Editing Board) and address as for Tijdschrift voor Entomologie. The Monographs are intended for the publication of larger entomological papers on a single subject and will appear irregularly. The following Monographs have been published: Hors série: F. T. Valck Lucassen et al., 1961. — Monographie du genre Lomaptera Gory & Percheron (Coleoptera, Cetoniidae), 299 pages, 739 figs, 2 pl., map. D.FI. 50.— (£5—.—, $ 13.90). No. 1. A. J. Besseling, 1964. — De Nederlandse Watermijten (Hydrachnellae Latreille, 1802) (The Hydrachnellae of the Netherlands), 199 pp. 333 figs., D.FI.25— (£2.10—, $6.95). NEUE KRATZMILBEN VON EINHEIMISCHEN INSEKTIVOREN (PSORERGATIDAE : TROMBIDIFORMES) von F. LUKOSCHUS Zoölogisch Laboratorium der Katholieke Universiteit, Nijmegen Zusammenfassung Es werden drei neue Psorergates (Psorergates)-Arten, die intraepithelial bei Insektivoren para- sitieren, beschrieben und in ihrer systematischen Stellung zu den bislang bekannten 18 Arten der Untergattung Psorergates (Psorergates) diskutiert. Die neuen Arten Psorergates (Psorergates) talpae, crocidurae und sorici wurden bei den niederländischen Wirten Talpa europaea europaea L., Crocidura russula russula (Hermann) und Sorex araneus araneus L. und bei den gleichen Wirtsarten in Südfrankreich und Nordspanien gefunden. Einleitung In einer vorangegangenen Untersuchung wurde Psorergates (Psorergates) desmanae Lukoschus, 1968 von Galemys pyrenaicus E. Geoffroy Saint Hilaire als erste Art dieser Familie von einem Insektivoren beschrieben. Alle übrigen Arten der Untergattung sind Parasiten von Rodentia (Tyrell, 1883; Michael, 1889; Till, 1960; Fain, 1961; Fain, Lukoschus & Hallmann, 1966; Lukoschus, Fain & Beaujean, 1967). Die neuen Arten wurden von frischgefangenen Wirten aus mehreren Fanggebieten des Zoologischen Institutes Nijmegen, sowie von Alkoholmaterial des Zoologischen Labo- ratoriums, Leiden, des Laboratoire Arago in Banyuls-sur-Mer (Pyr. Or.) und des Centro Pirenaico de Biologia Experimental in Jaca (Huesca) abgesammelt. Den Direktoren der Institute sei auch an dieser Stelle fiir ihr Entgegenkommen gedankt. Typen und Paratypen wurden in den folgenden Museen und Instituten deponiert: Rijksmuseum van Natuurlijke Historie, Leiden; British Museum (Natural History), London; Muséum National d’Histoire Naturelle, Paris; U.S. National Museum, Wash- ington; Zoologisches Staatsinstitut und Museum, Hamburg; Institut de Médecine Tro- picale Prince Léopold, Antwerpen; Zoologisches Institut der Akademie der Wissenschaf- ten, Leningrad; Departamento de Zoologia del Suelo y Entomologia aplicada, Madrid; Zoölogisch Laboratorium der Katholieke Universiteit, Nijmegen, im Text abgekürzt als Leiden, London usw. Beschreibungen Psorergates (Psorergates) talpae spec. nov. (Abb. 1-19) Mit den Charakteren der Familie Psorergatidae Dubinin (1955) und der Untergat- tung Psorergates Tyrell (Fain, 1961). Weibchen (Holotype). — Körperform und Kôrpergliederung wie bei den anderen Arten des Genus. Länge einschliesslich Gnathosoma 129 y, bei 31 gemessenen Paratypen im Mittel 136 u (126—144). Breite 101 y, bei den Paratypen © 105 u (99—117). 75 76 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 2, 1968 = Abb. 1—6. Psorergates (Psorergates) talpae spec. nov., Weibchen (Holotype). 1. Ventralansicht, 2. Bein II, Tibia-Tarsus dorsal, 3. Krallenpaar stark vergrôssert, 4. Dorsalansicht, 5. Gnathosomahaar dorsal, 6. Gnathosomahaar seitlich. F. Luxoscuus: Krätzmilben von Insektivoren 77 Ventralansicht (Abb. 1). Die weichhäutige Ventralfläche ist regelmässig fein gewellt. Die Anordnung der Wellung ist auch bei 43 daraufhin beobachteten Paratypen gleich- artig. Die Ventralhaare von 3—4 y stehen 8 u (7—11 bei den Paratypen) weit aus- einander. Die Epimeren I sind am hinteren Ende halbkreisförmig nach aussen gebogen. Der halbkreisformige Epimerenteil ragt kielartig aus der Ventralfläche heraus. Die Epimeren II-IV und der vordere Teil der Epimeren I liegen im Körperinneren dicht unter der gewellten Ventralflache. Die Wellung geht über die Epimeren hinweg. Ter- minalhaare von ca. 65 u. Die Beine sind ventral, weit seitlich eingesetzt. Alle Trochanteres besitzen nach unten und zum vorderen Körperdrittel gerichtete kräftige, an der Spitze leicht abgerundete Apophysen. Distal von den Apophysen stehen 9—11 u lange Haare. Der Basalteil des Trochanters ist stärker chitinisiert. Diese Chitinversteifung liuft gegenüber der Femur- apophyse in eine kleine abstehende Hautduplikatur aus. Der Femur aller Beinpaare trägt auf der Aussenseite auf einer kleinen stärker chitinisierten Platte ein Haarpaar und an seiner hinteren Ventralseite eine weit nach unten gerichtete spitze Apophyse. Bei den Haarpaaren ist das äussere Haar etwas länger. Es misst auf Femora I-III 28—36 u, auf Femur IV 33—39 u. Auf der hinteren Ventralseite steht auf einem kleinen Höcker von Genu I-III ein dünnes 3 y langes, und von Genu IV ein sehr dickes 50—55 y langes Haar. Die kurze Tibia trägt ventral vorn einen kurzen Dorn (fehlt auf Tibia IV) und dorsal median ein längeres Haar. Der Tarsus tragt ventral vorn einen kurzen, deut- lich gekriimmten Dorn, dorsal lateral zwei Haare. Im Gegensatz zu den bislang be- schriebenen Arten der Untergattung von Rodentia, bei denen die Dorsalhaare der Tarsi etwa gleichlang ausgebildet sind, messen die setae d.p. (fehlt auf Tarsus IV) 16—20 y, die setae d.a. dagegen nur 4 u (Abb. 2). Die beiden kräftigen einspitzigen Krallen (Abb. 3) und das zweiteilige Haftläppchen sind ventral am Tarsusende angesetzt. Die Tarsi I und II besitzen dorsal einen grösseren Sinneskolben und ein winziges Sinneshaar in einer kleinen Hautduplikatur. Dorsalansicht (Abb. 4). Das vorn auffallig eingebuchtete Rückenschild ist gut chiti- nisiert und bis zum Schildrande fein punktiert. Die weichhäutige Rückenfläche ist regel- mässig fein gewellt. Die Schildlänge beträgt (paramedian gemessen) 86 u, bei den Paratypen @ 86 y (80—96), die Breite 72 u, bei den Paratypen O 73 u (70—76). Einige Paratypen besitzen am Schildhinterrande weniger ausgeprägte Einbuchtungen. Die Schildhaare stehen auf kleinen, hellen unpunktierten Höfen. Die drei seitlichen Haarpaare, die deutlich vom Schildrande entfernt stehen, messen 4—5 y, die vorderen paramedianen sind sehr kurz, jedoch als Haare noch deutlich zu erkennen. Auf der Schildfläche sind in regelmässiger Anordnung 12 dunkle Punkte vorhanden. Bei 25 daraufhin untersuchten 9 9 fehlen in einem Falle die Punkte in Stellung a, bei einem anderen die in Stellung d. Gnathosoma und Mundteile sind wie bei den anderen Psorergates-Arten aufgebaut. Das 6 u lange Gnathosomahaar ist flach und am Ende mehrfach eingekerbt (Abb. 5). Die breitere, aufliegende Deckschuppe ist in mehrere feine Spitzen ausgezogen. Der Aufbau des komplizierten Haares ist aus Seitenansichten (Abb. 6) zu ersehen. Die Variabilitit des Haarrandes wird durch die Abb. 12—15 gezeigt. Das hintere Palptibia- haar von ca. 4 u (Abb. 19) ist am Ende abgesetzt und etwas dünner. Männchen (Allotype). — Kôrperlänge einschliesslich Gnathosoma 120 y, bei 7 Paratypen @ 115 u (108—120), Breite 99 u, bei den Paratypen @ 94 u (90—99). Ventralansicht wie beim Weibchen, jedoch nur zwei Terminalhaare auf einer lang- 78 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 16017 18919 im Abb. 7—19. Psorergates (Psorergates) talpae spec. nov. 7. Männchen (Allotype) dorsal, 8. Sklerit der Terminalhaare der Allotype, 9. Genitalregion und Penis einer Paratype, 10. Larve ventral, 11. Beinkralle stärker vergrössert, 12—15. Gnathosomahaare der Larve (12), der Protonymphe (15), der Deutonymphe (14) und des Männchens (13), 16—19. hintere Palptibiahaare der Larve (16), der Protonymphe (17), der Deutonymphe (18) und des Weibchens (19). F. LukoscHus : Krätzmilben von Insektivoren 79 Tabelle I. Messwerte und Determinationsmerkmale von Psorergates-Weibchen von Insektivoren. Angaben in u. Art: desmanae talpae crocidurae sorici Anzahl gemessene Exemplare: 12 31 10 10 Körperlänge im Durchschnitt 156 136 129 123 Körperlänge Minimum 135 126 123 117 Körperlänge Maximum 168 144 138 135 Körperbreite im Durchschnitt 127 105 104 95 Körperbreite Minimum 108 99 96 93 Körperbreite Maximum _ 140 117 112 99 Terminalhaare ca. 90 ca. 65 hi, SIS 52—60 Ventralhaare 5—6 3—4 2 4 Ventralhaarabstand ON UT 13—18 9—12 Trochanterhaare 10—12 9—11 8 9—10 Anzahl Femurhaare IV 2 2 2 1 Femurhaare I-III 20—28 28—36 20—25 20—25 Femurhaare IV 36—45 33—39 25—33 28—33 Genuhaare I-III 5 3 3 2 Genuhaare IV 51—60 50—55 30—35 30—35 Tarsus II d.a. 6 4 3 3—4 Tarsus II d.p. 15—18 16—20 0 14—18 Schildlänge im Durchschnitt 91 86 82 81 Schildlänge Minimum 87 80 78 76 Schildlinge Maximum 96 96 85 87 Schildbreite im Durchschnitt 84 73 69 66 Schildbreite Minimum 75 70 66 63 Schildbreite Maximum 90 76 72 72 Genitalöffnung 9—12 10—11 12 10—12 Palptibiahaar 5—6 4 3 D=) Gnathosomahaar 9—10 6 7 9 Anzahl Krallenspitzen 1 1 3 2 eiförmigen stärker chitinisierten Platte, die auf einem medianen Längswulst liegt (Abb. 8). Haare etwas kürzer als beim Weibchen, Messdaten in Tabelle II. Dorsalansicht (Abb. 7). An der Mitte des Vorderrandes des Riickenschildes eine ovale Genitalöffnung mit zwei Paaren winziger Härchen auf kleinen, hellen, runden Schild- chen. Das vordere Haarpaar steht 10 u, das hintere 9 u weit auseinander. Der zweispitzige Penis misst 25 y (25—29 bei den Paratypen), seine dorsal aufgeschlitzte Scheide 18 y. Die Seitenansicht des Penis (Abb. 9) von einem Quetschpräparat macht den Aufbau deutlich. Auf der Schildfläche sind auch bei allen Paratypen 12 dunkle Punkte in gesetzmässiger Anordnung vorhanden. Entwicklungsstadien. — Die dünnschaligen, fast runden Eier messen 85 y: 79 u (@ aus 10). Die Larven von durchschnittlich 93 y Länge und 78 y Breite zeigen bei gleichem Habitus wie bei anderen Arten der Untergattung eine auffällige regelmässige Wellung der Ventralseite (Abb. 10) und der Dorsalseite. Die Krallen der Beinendglieder sind 80 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 Tabelle II. Messwerte und Determinationsmerkmale von Psorergates-Männchen von Insektivoren. Angaben in y. Art: desmanae talpae crocidurae sorici Anzahl gemessene Exemplare: 5 7 10 10 Körperlänge im Durchschnitt 137 115 -111 AI Körperlänge Minimum 132 108 105 102 Körperlänge Maximum 146 120 114 115 Körperbreite im Durchschnitt 112 94 90 87 Körperbreite Minimum 105 90 85 84 Körperbreite Maximum 120 99 93 935 Terminalhaare 72—89 51—54 42—48 52—60 Ventralhaare ASS 3 2—3 3—4 Ventralhaarabstand 7—8 7—10 12—14 10—14 Trochanterhaare 9—10 7—8 8—9 7 Anzahl Femurhaare IV 2 2 2 1 Femurhaare I-III 15—20 25—28 18—23 20—25 Femurhaare IV 25—36 30—33 20—25 20—25 Genuhaare I-III 4 3 3 2 Genuhaare IV 35—40 40—45 30—33 27—34 Tarsus II da. 5 3 2 3 Tarsus II d.p. 14—15 14—16 0 15 Schildlänge im Durchschnitt 87 73 Uy 7a Schildlänge Minimum 84 72 75 70 Schildlänge Maximum 90 74 84 78 Schildbreite im Durchschnitt Ti 64 67 59 Schildbreite Minimum 75 62 64 57 Schildbreite Maximum 81 66 70 62 2. Schildhaar 4—4,5 4 3—4 3—4 Palptibiahaar 5 3,5 3 2 Gnathosomahaar IV 6 TS TS Penis 28—31 25—29 24—27 36—38 Penisscheide 23—24 18 15—16 NINA Genitalhaarabstand vorn 9—10 10—11 12—13 12—14 Genitalhaarabstand hinten 10—11 9 7—8 5 Anzahl Krallenspitzen 1 1 5 2 dreispitzig (Abb. 11). Auf den Beinen I und II ist je ein Sinneskolben vorhanden. Die Gnathosomahaare (Abb. 12) sind in der gleichen Grösse wie bei den Adulten. ausgebildet (Abb. 5, 6 von 9, 13 von &). Die Art unterscheidet sich dadurch von allen bislang beschriebenen Arten von Rodentia. Die dorsalen hinteren Palptibiahaare (Abb. 16) sind ebenfalls fast in der gleichen Form und Grösse wie bei den Adulten vorhanden. Die Protonymphen von durchschnittlich 111 u Länge und 90 u Breite besitzen gleich- artig gebildete Gnathosomahaare (Abb. 15) und Palptibiahaare (Abb. 17). Im Ven- tralbereich der Ventralseite ist die Wellung längsgerichtet. Die Existenz der morphologisch etwa gleichgebauten Deutonymphe geht aus Häu- F. LukoscHus : Krätzmilben von Insektivoren 81 tungsstadien hervor, in denen sich eine Form mit vier Paaren kurzer, zweigliedriger Beine in einer gleichartigen Form entwickelt. Die Deutonymphen sind im Durchschnitt 130 u lang und 103 u breit. Deutonymphen, in denen sich Adulte entwickeln, zeigen Gnathosomahaare wie Abb. 14 und Palptibiahaare wie Abb. 18. Die Wellung der Ventralseite ähnt der des Weibchens. Bei gutgestreckten Präparaten kònnen Proto- und Deutonymphe durch die Stellung der Beine IV unterschieden werden. Bei der Deuto- nymphe ist der Abstand der Beine voneinander fast doppelt so gross wie bei der Protonymphe. Wirt und Verbreitung. — Die Typenserie entstammt einer Talpa europaea europaea L., die am 22.VI.1967 bei Nijmegen gefangen wurde. Weitere parasitierte Tiere der gleichen Wirtsart: 4 aus der Umgebung des Typenfangplatzes 6.VI.1967—24.I.1968, 7 aus der Umgebung von Villanua, Prov. Huesca, Spanien, 11.VII.1966—20.V.1967 (Alkoholmaterial des Centro Pirenaico de Biologia Experimental, Jaca). Einfluss der Parasiten auf den Wirt. — Die Milben sitzen oberflächlich in der Epidermis des äusseren Gehörganges. Sie verursachen dort Hyperceratosis und Hyper- funktion der Talgdrüsen. Bei den bislang untersuchten, nur schwach parasitierten Wirten wurde ein Befall anderer Körperstellen nicht festgestellt. Typen. — Holotype 9 und Allotype 3 in Leiden. Paratypen 9 und ¢ in London, Paris, Washington, Antwerpen, Nijmegen. Paratypen 9 in Hamburg, Leningrad, Madrid. Psorergates (Psorergates) crocidurae spec. nov. (Abb. 20—32) Mit den Charakteren der Familie Psorergatidae Dubinin (1955) und der Unter- gattung Psorergates Tyrell (Fain, 1961). Weibchen (Holotype). — Lange einschliesslich Gnathosoma 135 u, bei 10 gemes- senen Paratypen @ 129 u (123—138). Breite 108 y, bei den Paratypen O 104 y (96—112). Ventralansicht (Abb. 23). Die weichhäutige Ventralfläche ist im medianen Bereich schwach längsgewellt. Die sehr kurzen Ventralhaare (2 u) stehen 16 u (13—18 u bei den Paratypen) weit auseinander. Die Epimeren I sind am Ende nach aussen gebogen und mit einer feinen nach vorn laufenden Linie eiförmig geschlossen. Ein halbkreis- formiger Epimerenteil ragt kielartig aus der Ventralfläche heraus. Die unter der Korper- oberfläche liegenden Epimeren II-IV besitzen an ihren inneren Enden langovale helle Flächen. Die zwischen den beiden Längswülsten langgestreckte Genitalöffnung von 12 y ist in ihrem vordersten Teil durch eine Hautduplikatur der Ventralfläche über- deckt. Die Terminalhaare sind etwa 55 y lang. Die Beine sind ventral, weit seitlich eingesetzt. Alle Trochanteres haben nach unten und zum vorderen Körperdrittel gerichtete kräftige spitze Apophysen. Trochanterhaare ca. 8 p. Der stärker chitinisierte Basalteil des Trochanters läuft gegenüber der Femur- apophyse in eine sehr kleine, abstehende Hautduplikatur aus. Der Femur aller Beinpaare trigt auf der ventralen hinteren Seite ein Haarpaar und eine nach unten abstehende, spitze Apophyse. Bei den Haarpaaren ist das äussere Haar etwas länger. Es misst auf Femur I-III 20—25 y, auf Femur IV 25—33 y. Die Haare auf Genu I-III sind sehr kurz (3 u), auf Genu IV sehr kräftig und 30—35 y lang. Die kurze Tibia trägt ventral vorn einen kurzen gekriimmten Dorn (fehlt auf Bein IV) und dorsal median ein längeres Haar. Die Tarsi haben ventral vorn einen kurzen kräftigen Dorn und dorsal sehr fein, 3 „ lange setae d.a. (Abb. 24). Die dorsale posteriore setae sind nicht vorhan- 82 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 Abb. 20—25. Psorergates (Psorergates) crocidurae spec. nov. 20. Holotype Weibchen dorsal, 21. Palptarsus einer gequetschten Paratype, 22. Gnathosomahaar der Holotype, 23. Holotype Weibchen ventral, 24. Bein II, Tibia-Tarsus dorsal einer Paratype, 25, Krallenpaar stärker vergrössert, F. LugkoscHus : Krätzmilben von Insektivoren 83 den. Die beiden kräftigen dreispitzigen Krallen (Abb. 25) und das zweigeteilte Haft- läppchen sind ventral am Tarsusende angesetzt. Auf Tarsus I und II ist dorsal je ein dickerer Sinneskolben und ein winziges Sinneshaar in einer kleinen Hautduplikatur vorhanden. Dorsalansicht (Abb. 20). Das am Vorderrande auffällig eingebuchtete Rückenschild ist gut chitinisiert und bis zum Schildrande deutlich fein punktiert. Die Schildlänge (von der Verbindungslinie der vorderen Spitzen an gemessen) beträgt 82 u, bei den Paratypen @ 82 u (78—85), die Breite 69 u, bei den Paratypen @ 69 u (66—72). Die Schildhaare stehen auf kleinen unpunktierten Höfen, seitliche Schildhaare 4 u, vordere paramediane Haare 1 u. Auf der Schildfläche sind in regelmässiger Anordnung 12 dunkle Punkte vorhanden. Gnathosoma und Mundteile sind wie bei anderen Psorergates-Arten aufgebaut. Abb. 21 zeigt die Ausbildungsform der Palptarsuskrallen von einer gequetschten Paratype. Das ca. 7 u lange, abgeflachte Gnathosomahaar ist am Ende mehrfach eingekerbt, die aufliegende breitere Deckschuppe ist am Rande in 5—7 abgestumpfte Enden ausgezogen (Abb. 22). Die feinen hinteren Palptibiahaare messen nur 3 u. Männchen (Allotype). — Körperlänge einschliesslich Gnathosoma 111 y, bei 10 gemessenen Paratypen @ 111 x (105—-114). Körperbreite 90 y, bei den Paratypen D 90 u (85—93). Ventralansicht wie beim Weibchen, jedoch nur zwei Terminalhaare auf einem lang- gestreckten stärker chitinisierten Schildchen (Abb. 28). Körper und Beinbehaarung etwas kürzer als beim Weibchen (Messdaten in Tabelle IT). Dorsalansicht (Abb. 26). An der Mitte des Vorderrandes des Rückenschildes liegt die langovale Genitalöffnung inmitten eines Bereiches, der stärker chitinisiert und dichter punktiert ist. Die winzigen Härchen an den Seiten der Genitalöffnung stehen in kleinen, unpigmentierten hellen Höfen. Das vordere Haarpaar steht 12—13 n, das hintere 7—8 y weit auseinander. Auf der Rückenschildfläche sind 10 dunkle Punkte in gesetzmässiger Anordnung vorhanden. Der Penis misst 24—27 y, seine Scheide 15—16 u. Die Ausbildungsform des Penis wird aus Seitenansichten (Abb. 27) deutlich. Gnathosoma, Mundteile, Palptibiahaare und Gnathosomahaare (Abb. 29) unterscheiden sich nicht vom Weibchen. Entwicklungsstadien. — Die etwa runden, dünnschaligen Eier (O aus 20) messen 85 u: 76 y. Larven und Nymphen sind wie bei anderen Psorergates-Arten ausgebildet. Die Larven messen (© aus 10) 85 : 70 y. Palptibiahaare und Gnathosomahaare (Abb. 32) unterscheiden sich nicht wesentlich von der Ausbildung der Adulten. Die Proto- nymphen besitzen dicht beieinander stehende Beine IV. In der Grösse der Gnathoso- mahaare (Abb. 31) und der Palptibiahaare unterscheiden sie sich nicht wesentlich von anderen Entwicklungsstadien. Die Protonymphen sind im Durchschnitt (10) 100 y lang und 74 n breit. Die Deutonymphen haben weiter auseinander stehende Beine IV. Sie messen (© aus 10) 126 : 94 y. Die Gnathosomahaare (Abb. 30) sind wie bei den anderen Entwicklungsstadien ähnlich wie bei den Adulten aufgebaut, die Deckschuppe ist etwas verkürzt. Wirt und Verbreitung. — Die Typenserie entstammt einer Crocidura russula russula (Hermann), die am 12.VI.1967 bei Nijmegen gefangen wurde. Acht Tiere der gleichen Wirtsart von mehreren Fangplitzen in der Umgebung von Nijmegen, ein Wirt von Banyuls (Alkoholmaterial des Laboratoire Arago, Banyuls) und je ein Wirt von Lerida und Madrid (Alkoholmaterial des Centro Pirenaico de Biologia Experimental, Jaca) waren ebenfalls parasitiert. 84 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 Sp 50u 100 u Abb. 26—32. Psorergates (Psorergates) crocidurae spec. nov. 26. Allotype Männchen dorsal, 27. Penis einer Paratype, 28. Sklerit mit Terminalhaaren der Allotype, 29—32. Gnathosomahaare der Allotype (29), der Deutonymphe (30), der Protonymphe (31) und der Larve (32). Abb. 33—34. Psorergates (Psorergates) sorici spec. nov. 33. Dorsalnsicht der Allotype Männchen, 34. Sklerit mit Terminalhaaren der Allotype F. LukoscHus: Krätzmilben von Insektivoren 85 Einfluss des Parasiten auf den Wirt. — Die Milben sitzen in der Epidermis des spärlich behaarten Tibiabereiches der Hinterbeine und am Rande der Ohrmuschel und der Ohrmuschelfalten. Sie verursachen dort eine Hypertrophie von Epidermis und Bindegewebe. Bei stärkerem Befall treten blutdurchsetzte Krusten auf. Sekundärinfektio- nen sind bei alterem Befall wahrscheinlich. Abb. 41 zeigt ein Foto des abgeschnittenen Ohres des Typenwirtes. Auf der Fläche der Ohrmuschelfalte ist ein Befall grösserer Ausdehnung vorhanden, der einige kleine Blutkrusten enthält. Der Rand ist zu acht, teilweise grösseren gallartigen Epidermiswucherungen ausgewachsen. Typen. — Holotype © und Allotype & in Leiden. Paratypen @ und 4 in London, Paris, Washington, Hamburg, Antwerpen, Leningrad, Madrid, Nijmegen. Psorergates (Psorergates) sorici spec. nov. (Abb. 33—40) Mit den Charakteren der Familie Psorergatidae Dubinin (1955) und der Untergat- tung Psorergates Tyrell (Fain, 1961). Weibchen (Holotype). — Lange einschliesslich Gnathosoma 123 u, bei 10 gemessenen Paratypen © 123 u (117—135). Breite 96 u, bei den Paratypen © 95 u (93—99). Ventralansicht (Abb. 38). Die weichhäutige Ventralfläche ist nur vor der Genital- öffnung schwach gewellt. Die Ventralhaare von 4 u Länge stehen 9 u (9—12) weit auseinander. Der vordere Teil der Epimeren I liegt wie die Epimeren II-IV im Kôrper. Der hintere Epimerenteil I ragt in der Form eines fast geschlossenen Kreises kielartig weit aus der Ventralfläche heraus. Es entsteht der Eindruck von Saugnäpfen. Die Terminalhaare messen 54 u (52—60). Die ca. 10 n lange Genitalöffnung liegt lang- gestreckt zwischen zwei Längswülsten. Die Beine sind ventral, weit seitlich eingesetzt. Die Trochanterapophysen sind vorn abgestumpft. Die Hautduplikatur der Chitinversteifung der Trochanterbasis ist klein und wenig abstehend. Trochanterhaare 9—10 u. Der Femur trägt eine weit nach unten abstehende Apophyse und auf Beinpaar I-III zwei Haare, auf Femur IV nur ein Haar. Das äussere wenig längere Haar misst auf Beinpaar I-III 25 u, auf Bein IV 30 u. Auf einem kleinen Höcker von Genu I-III steht ein sehr kurzes (2 „) Haar, von Genu IV ein dickes, 35 u langes Haar. Die Tibia trägt ventral vorn einen diinnen gekriimmten Dorn (fehlt auf Tibia IV) und dorsal median ein längeres Haar. Der Tarsus hat ventral vorn einen stärkeren gekriimmten Dorn und dorsal lateral zwei Haare ungleicher Länge (Abb. 39); die dorsale anteriore setae messen 3—4 u, die dorsale posteriore setae 14—18 y (d.p. fehlt auf Tarsus IV). Die kräftigen zweispitzigen Krallen (Abb. 40) und das zweigeteilte Haftlippchen sind ventral am Tarsusende angefiigt. Tarsus I und II besitzen dorsal einen grösseren Sinneskolben und ein winziges Sinneshärchen in einer kleinen Hautduplikatur. Dorsalansicht (Abb. 35). Das gut chitinisierte und bis zum Rande gleichmässig punktierte Rückenschild ist am Vorderrande eingebuchtet. Die Linge von der Verbin- dungslinie der beiden vorderen Spitzen an gemessen beträgt 79 u, bei den Paratypen © 81 u (76—87), die Breite 64 y, bei den Paratypen @ 66 u (63—72). Die Schild- haare stehen auf kleinen unpunktierten Höfen deutlich vom Schildrande entfernt. Die seitlichen Schildhaare sind 4—5 y lang, die vorderen 1 u. Auf der Schildfläche sind in regelmässiger Anordnung auch bei allen beobachteten Paratypen 12 dunkle Punkte vorhanden. Gnathosoma und Mundteile sind wie bei anderen Psorergates-Arten aufgebaut. Das ca. 9 y lange flache Gnathosomahaar ist an seinem Ende unterschiedlich lang gelappt 86 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 Su 36,3740 Abb. 35—40. Psorergates (Psorergates) sorici spec. nov. 35. Holotype Weibchen dorsal, 36. Gnathosomahaar der Holotype seitlich, 37. Gnathosomahaar dorsal, 38. Holotype ventral, 39. Bein II, Tibia und Tarsus dorsal, 40. Krallenpaar stärker vergrössert. F. LukoscHus : Krätzmilben von Insektivoren 87 (Abb. 37). Der Aufbau aus aufliegender breiter gezähnter Deckschuppe und flachem Haar geht aus Seitenansichten (Abb. 36) hervor. Die dünnen hinteren Palptibiahaare sind nur 2—3 u lang. Männchen (Allotype). — Körperlänge einschliesslich Gnathosoma 114 u, bei 10 gemessenen Paratypen @ 111 u (102—115). Breite 90 u, bei den Paratypen © 87 u (84—93). Ventralansicht wie beim Weibchen, jedoch mit nur zwei Terminalhaaren auf einem langgestreckten stärker chitinisierten Schildchen (Abb. 34). Haare etwas kürzer als beim Weibchen (Tabelle II). Dorsalansicht (Abb. 33). Das gut chitinisierte und punktierte Rückenschild besitzt hinter dem 2. Haarpaar typische, bei allen Paratypen gleichförmig ausgebildete Ein- buchtungen, in die sich die weichhäutige, regelmässig fein gewellte Rückenhaut hin- einzieht. Die Genitalòffnung ist an den Vorderrand des Rückenschildes verlagert. Die inneren (hinteren) Genitalhaare kommen dadurch auf ihren langovalen hellen Schild- chen an den Vorderrand des Schildes. Die inneren Härchen stehen 5 u, die äusseren 12—14 y weit auseinander. Die Seitenhaare des Schildes stehen auf grösseren hellen Höfen. Der 36—38 y lange Penis ist sehr dünn und spitz, er ragt bei allen Paratypen weit aus der Genitalöffnung heraus. Die Penisscheide misst nur 11—14 u. Auf der Schildfläche sind auch bei allen Paratypen 12 dunkle Punkte in regelmässiger Anord- nung vorhanden. Bei einigen Paratypen sind zusätzlich Punkte in Stellung d vorhanden. Entwicklungsstadien. — Die dünnschaligen fast runden Eier sind im Durchschnitt 81 u : 78 y gross. Die Entwicklungsstadien sind wie die Adulten im Vergleich zu P. talpae und crocidurae etwas mehr langgestreckt. Wirt und Verbreitung. — Die Typenserie entstammt einem Sorex araneus araneus pee Abb. 41. Abgetrennte Ohrmuschel von Crocidura russula (Typenwirt) mit starkerem Psorergates- Befall. Erläuterung im Text. Foto H. J. M. Spruyt. 88 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 3, 1968 L., der am 26.V.1967 bei Nijmegen gefangen wurde. Sechs weitere Tiere der gleichen Wirtsart aus der Umgebung von Nijmegen und zwei aus dem Alkoholmaterial des Zoologischen Laboratoriums, Leiden, waren ebenfalls parasitiert. Einfluss der Parasiten auf den Wirt. — Die Milben sitzen oberflächlich in der Epidermis der Ohrmuschel und des schwachbehaarten Tibiabereiches der Hinterbeine. Sie verursachen dort Hyperceratosis. Blutschorfe an den Hinterbeinen weisen darauf hin, dass der Wirt sich stärker belästigt fühlt. Typen. — Holotype @ und Allotype 4 in Leiden. Paratypen 9 und ¢ in London, Paris, Washington, Hamburg, Antwerpen, Leningrad, Madrid, Nijmegen. Abgrenzung gegenüber benachbarten Arten In einer vorangegangenen Untersuchung (Lukoschus, Fain & Beaujean, 1967) wurden die bis dahin bekannten 17 Arten der Untergattung Psorergates (Psorergates) nach ihren Merkmalen in Gruppen einander nahestehender Arten zusammengefasst. Die Typenart der Familie P. (P.) simplex Tyrell (1883) konnte dabei nicht berücksichtigt werden, da die Beschreibung dafür nicht ausreichend ist und die Typenexemplare ver- loren gegangen sind. In den Tabellen I und II sind die Messdaten von ® und & der an Insektivoren parasitierenden Arten zusammengefasst. Fett und kursiv weisen dabei auf wichtige Determinationsmerkmale hin. Den Arten auf Insektivoren ist gemeinsam: 1. sehr kurze setae dorsal anterior auf den Tarsen; 2. vorn stark ausgebuchtetes Rückenschild; 3. alle Schildhaare deutlich vom Schildrande entfernt; 4. Gnathosomahaare gekerbt oder gelappt; 5. Terminalhaare beim Männchen auf einem längeren, stärker chitinisierten Sklerit. Diese Merkmale unterscheiden die Arten von Insektivoren deutlich von den Arten, die Rodentia parasitieren. Die Arten P. (P.) desmanae, talpae, crocidurae und sorici können deshalb als Insektivoren-Gruppe innerhalb der Untergattung Psorergates sensu Fain (1961) zusammengefasst werden. Die Gesetzmässig angeordneten dunklen Punkte auf dem Riickenschild verdienen weitere Beachtung. Es handelt sich dabei nicht um stark reduzierte Haare, wie sie bei den Arten P. P. pitymydis und deomydis beobachtet wurden. Literatur Dubinin, W., 1955. Acariens der rongeurs d’U.R.S.S. II. Psorergatidae. Trav. Acad. Sci. U.R.S.S. (Zool.) 59: 150—152. Fain, A., 1961. Notes sur le genre Psorergates Tyrell. Description de Psorergates ovis Womersley et d'une espèce nouvelle. Acarologia 3 : 60—71. Fain, A., F. Lukoschus & P. Hallmann, 1966. Le genre Psorergates chez les muridés. Description de trois espèces nouvelles (Psorergatidae: Trombidiformes). Acarologia 8: 251—274. Lukoschus, F., A. Fain & M. M. J. Beaujean, 1967. Beschreibung neuer Psorergates-Arten. Tijdschr. Ent. 110 : 133—181. Lukoschus, F., 1968. Psorergates desmanae spec. nov., eine neue Krätzmilbe von Galemys pyrenaicus (Psorergatidae: Trombidiformes). Bull. Mus. Nat. d'Hist. Nat., Paris 40 (1) (im Druck). Michael, A. D., 1889. On some unrecorded Acari found in Great Britain. J. Linn. Soc. (Zool.), London 20 : 400—406, pl. 26. Till, W. M., 1960. Psorergates oettlei n.sp. A mange-causing mite from the multimammate rat (Acarina: Psorergatidae). Acarologia 2 : 75—79. Tyrell, J. B., 1883. On the occurrence in Canada of two species of parasitic mites. Proc. Canad. Inst. Toronto n.s. 1: 342—343, pl. IV. tay UC PS oN AA x LIS } Cona: at 4 1 } x 4 a J | . | AL GY | 2 US, COMP, VOL DEEL 111 AFLEVERING 4 LIBRARY 1968 NOV 5 1908 HARVARD TIJDSCHRIFT 5% VOOR ENTOMOLOGIE UITGEGEVEN DOOR DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING INHOUD : J. P. van LitH. — Contribution to the knowledge of Indo-Australian, South Pacific and East Asiatic Psenini. Genus Psen Latreille (Hymenoptera, Sphecidae), pp. 89—135, Figs. 1—44. Tijdschrift voor Entomologie, deel 111, afl. 4 Gepubliceerd 10-X-1968 NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING BESTUUR (BOARD) Voorzitter (Chairman) . . RPG" Barendrecht Vice-Voorzitter (Vice- President) ei ur Sl Wtebes Secretaris (Secretary) . . . . . . . . W. Hellinga Address io ee a ti Weesperzijide 123 11, Amsterdam: O. Penningmeester (Treasurer) . . . . . W.J. Kabos PAGA ressa . . . + + Van Baerlestraat 261, Amsterdam Bibliothecaris (Libre) ON NEN IG SAINTE D Teekel Address . . . . 2.2.2 202020. 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The Journal serves the publication of papers on Insecta, Myriapoda and Arachnoidea. It appears in separate issues, forming an annual volume of 350—400 pages. Subscription rate: D.Fl. 45 — (£4.10—, $12.50) per volume. MONOGRAFIEEN VAN DE NEDERLANDSCHE ENTOMOLOGISCHE VEREENIGING Redactie (Editing Board) and address as for Tijdschrift voor Entomologie. The Monographs are intended for the publication of larger entomological papers on a single subject and will appear irregularly. The following Monographs have been published: Hors série: F. T. Valck Lucassen et al., 1961. — Monographie du genre Lomaptera Gory & Percheron (Coleoptera, Cetoniidae), 299 pages, 739 figs, 2 pl, map. D.Fl. 50— (£5——, $ 13.90). No. 1. A. J. Besseling, 1964. — De Nederlandse Watermijten (Hydrachnellae Latreille, 1802) (The Hydrachnellae of the Netherlands), 199 pp., 333 figs, DFI. 25 — (£2.10.—, $ 6.95). DE j we MUS. COMP. ZGOL. LIBRARY NOV 5 1968 CONTRIBUTION TO THE KNOWLEDGE ORANDO- AUSTRALIAN, SOUTH PACIFIC AND EAST ASIAMERPSENINI GENUS PSEN LATREILLE (HYMENOPTERA, SPHECIDAE) by J. P. VAN LITH ABSTRACT New species and subspecies of the subgenus Pser Latreille from the Indo-Australian and South Pacific regions are described: aspites, nepalensis, sauteri, elisabethae madrasiensis, heinrichi, regalis, marjoriae, paulus paulus, paulus baduriensis, paulus subtilis, vadosus, bishopi, simlensis, sedlaceki. Of a few species described earlier from only one sex the opposite sex was found. A new key is given for the subgenus Pser, now including species from Indo-Australia, the South Pacific Islands and East Asia. A new subgenus, Punctipsen, is erected to receive P. exaratus (Eversmann) of which a new subspecies, exaratus indicus, is described. A remarkable new species, inflatus, has been collected by the Third Archbold Expedition, 1938—1939, to New Guinea. It may belong to the subgenus Mimesa Shuckard, a taxon not yet recorded from Indo-Australia. During the last few years a relatively great number of Psenini became available for study. Most of these Psenini were collected in the Indo-Australian and South Pacific area for the Bernice P. Bishop Museum, Honolulu. The directors of the museum of the Humboldt-University, Berlin, and of the Deutsches Entomologisches Institut, Eberswalde, had the kindness to send me their Psenini, including those collected by H. Sauter in Formosa (also his types) and by G. Heinrich in Celebes. Especially the forms from Celebes proved to be very interesting, some being strongly different from the other Indo-Australian Psen Latreille and Psenulus Kohl. The new species of Psen from Celebes are described in this paper; the genus Psenulus will be discussed in a future publication. Dr. G. R. Ferguson, Scarsdale, and the Rijksmuseum van Natuurlijke His- torie, Leiden, obtained some good specimens from South India from P. S. Nathan, including a new subspecies of Psen exaratus (Eversmann), which is the first record of this species from Southern Asia. Through the kind intermediary of Prof. Dr. J. van der Vecht and Dr. V. Gupta I received some fresh material from North India. Dr. C. R. Baltazar, Manila, and Dr. Henry K. Townes, Ann Arbor, sent me some Psenini from the Philippines and Japan. Our knowledge of the Psenini from New Guinea and Papua could be much improved, to which the material collected for the British Museum (Natural History) by Miss L. E. Cheesman has also contributed. Important work has been done by Prof. Dr. K. Tsuneki, Fukui, Japan, who studied Sphecidae, including a number of Psenini, from the Ryukyus and Formosa. In my keys to the subgenera Psen and Punctipsen I have tried to include the species treated in his publications. I express my gratitude to all these institutions and entomologists. Special thanks are due to Dr. K. V. Krombein for allowing me to study his material from Ambon, New Guinea and Papua, although he had originally reserved this for his own publications on Sphecidae from this region. 89 As 90 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Now 65 species and subspecies of the subgenus Psen are known from East Asia and Indo-Australia. From Formosa, the Ryukyus and the South Pacific Islands 15 species and The subspecies are added here, making a total of 80 (64 species and 16 subspecies). subgenera Punctipsen, Mimumesa, Mimesa, Pseneo and Odontopsen count together 15 species and 9 subspecies. The extension of my study to the South Pacific Psenini revealed interesting close relationships between forms of the group of P. elisabethae occurring in the area from South India to Samoa. The names of the institutions and entomologists are abbreviated as follows: BISH — Bernice P. Bishop Museum, Honolulu BM — British Museum (Natural History), London BPIM — Bureau of Plant Industry, Entom. Research Section, Manila Coll. Ferguson — Dr. G. R. Ferguson, Scarsdale, New York, U.S.A. ML — Rijksmuseum van Natuurlijke Historie, Leiden OUM — Oxford University Museum, Hope Department of Entomology, Oxford Coll. Townes — Dr. Henry K. Townes and Dr. M. Townes, Ann Arbor, U.S.A. UDE — University of Delhi, Department of Entomology, Delhi USNM — United States National Museum, Washington D.C. ZMB — Zoologisches Museum der Humboldt-Universitàt, Berlin a ol KEY TO THE INDO-AUSTRALIAN, SOUTH PACIFIC AND EAST ASIATIC SPECIES OF THE SUBGENUS Psen LATREILLE . Tarsi of fore legs of male laterally enlarged into a broad plate (fig. 1). Clypeus with broad and smooth ivory-white margin. Underside of head with. long white hairs. Mesosternum short. Legs largely reddish; gaster including petiole reddish. Female unknown. (S. ‚Gelebes) wer CITI IE aspites sp. n. Fore tarsi of male without plate. Clypeus without ivory-white margin. Mesosternum longer, ini RETI III SITI IATA LITI TITTI 2 Female with long and pointed interantennal tooth (lateral view); pygidial area broadly triangular, densely punctate. Mid basitarsi of male broadened ......... 3 Interantennal tooth less strong. Basitarsus of mid legs of male normal ......... 5 All tarsal segments of mid legs of male of abnormal shape. Face of female silvery; third antennal segment at most three times as long as it is broad at apex. Antennal segments 6—13 of male with tyloidea. (Europe, Siberia, China, Mongolia, Korea and Japam)ae. ner. ee ee lege. II ater Only basitarsus of mid legs of male with hook-shaped protuberance ............ 4 Antennal segments 6—12 (or 13) of male with tyloidea. Face of female golden; third antennal segment more than three times as long as it is broad at apex. (Japan) u... Re RAR CR III aurifrons Antennae of male without tyloidea. Female unknown. (Japan) ...... yasumatsui Posterior margin of gastral tergites with fringe of long hairs. ..................... 6 No distinct fringe of long hairs indd ndgn 20er EE 13 Back of propodeum in both sexes with irregular carination. Scape of antennae thick, further antennal segments normally slender. Pygidial area of female densely punctate. Antennae of male with tyloidea. Underside of petiole smooth in both sexes: \Group).0f P. emangenalusı EOS secre een eee ee eee 7 \O 10. IM. 12% 13: 14. J. P. van LrrH: Oriental species of the genus Psen 91 Back of propodeum, especially of females, with parallel striation. Antennae thick, flagellum almost as broad as scape. Pygidial area of female broadly triangular, dull, an irregular row of punctures along the margin. Antennae of male without tyloidea. Underside of petiole irregular. Group of P. curvipilosus ............ ul Posterior margin of tergites with fringe of black hairs. Gaster black with bluish shinesabemale, unknown (Malay) en ee nen ee pilosus Fringe of hairs on posterior margin of tergites paler ................................. 8 . First gastral tergite reddish-brown laterally. Posterior margin of tergites with fringe of long fuscous-golden hairs. Face of female pale golden. Length of female 13=—I4ymmy of malen lE 12 mm. (Japan). en le nete tee dzimm Nordistinceredion first) gastral tersite inte 2e eee cesser nnt 9 . Posterior margin of tergites with dense fringe of golden-brown, slightly bent hairs. Face of female pale golden. Length about 9 mm. Male unknown (E. Nepal) ...... ARR LITRO PARENT SOSTARE OLII RT nepalensis sp. n. Hairs on posterior margin of tergites paler .......................................... 10 Posterior margin of tergites with dense fringe of yellowish-grey long hairs. Disk of tergites with dense pubescence. Face of female golden. Length about 12 mm. Malemunknow ten (Formosa) teen sauteri sp. n. Posterior margin of tergites with inconspicuous fringe of yellowish stiff hairs. Disk of tergites normally pubescent. Face of both sexes silvery. Length of female about li0*mm*ofimaletaboutt8:5tmmi (Java, Borneo) tn emarginatus Curved long hairs along margin of gastral tergites and pubescence on disk of tergites golden-brown. Pubescence of face pale golden in both sexes. (Java) … … scat doder ed JENGA ARANEOS OO curvipilosus Curved hairs almost black, pubescence on disk of fourth, fifth and sixth gastral terpites greyish Pubescencerofs face SILVER) rer eee eee cesse 12 Hind tibiae brown. Larger. Punctures on second gastral tergite indistinct. In female end of petiole coarsely punctate ventrally, but punctures not deep. In male underside of petiole not or slightly concave on either side of median carina, very coarsely striato-punctate there. Carinae on either side of median ventral carina usually of irregular length. (Sumatra, Malaya, Formosa?) … … lieftincki lieftincki Hind tibiae darker, nearly black. Smaller. Punctures on second gastral tergite more distinct but still fine. In female end of petiole below much finer punctate. In male underside of petiole concave on either side of median carina, few punc- tures only. Carinae on either side of median ventral carina usually of about equal mein (ONE) asso ersehen lieftincki minor Mandibles extremely broad. Interantennal carina often ending in a transverse, much protruding carina. Pygidial area of female triangular, dull, an irregular row of punctures along the margin. In female, sometimes also in male, part of the longitudinal carinae of the enclosed area of the propodeum distinctly continued on to back of propodeum. Antennae of male probably always without tyloidea. Group OL PEUR TIM eine nere nie ein 14 Mandi lest Ema Nee attente deelen « 19 Petiole more or less quadrate in cross section; sides depressed, consequently with distinct longitudinal edges above and below. Median ventral carina present 15 Petiole rounded below and only upper lateral edges distinct or petiole almost eylindkieal 28. ordende eenden sasenondendeneaneenernensvdsressensenseemersend enn nrenverseenee 16 97. 115) 16. 19. 20. 21° TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Posterior end of petiole about 1 2/3 times as broad as base of petiole. Reticulate carination on back of propodeum rather coarse. Ventral median carina sharper. Punc- turation of frons very fine but distinct, sparser on vertex. Female unknown. (SIDERNO IIS éagosaupncengoc oc 0e ussuriensis Petiole more slender, sides almost parallel. Carination on back of propodeum less coarse. Frons and vertex almost impunctate. Ventral median carina of petiole of malesweaker»Racerot temale’silvery.2 (Japaner m en ee tsunekit Legs dark, tibiae brown. Frons and vertex with extremely fine puncturation. Back of propodeum with very coarse and irregular carination; a few of the longitudinal carinae of the enclosed area distinctly continued on to the back. Female unknown. (CASSa OOO TTT ASSAMENSIS Less, atleastitibiae, patty ed... TE 17 Only tibiae and tarsi reddish. Femora nearly black, underside brown. Antennae dark brown, underside paler, scape dark. Face of female golden. Male unknown. (Burma) is IT vechti birmanicus Legs completely ‘or largely *reddish o IP 18 Scape of antennae reddish, also underside of flagellum. All legs, except trochan- ters, completely red. Sides of petiole finely punctate. Face of female golden. Male Unknown. (JAVA) vu... scp se doc edge Se ge vechti vechti Scape of antennae black with iridescent shine, flagellum slightly brownish. Fore and mid femora largely black. No tyloidea. Face of male silvery. Female unknown. (Eotmosa) iI shirozui Behind enclosed area of propodeum on both sides of median sulcus a smooth area, at most finely coriaceous. In case the smooth area is narrow and indistinct (cf. P. politiventris and P. orientalis) the dorsal side and back of propodeum forming an obtuse angle (cf. P. refractus if propodeum more or less smooth only dorso- laterally, behind enclosed area and second recurrent vein ending in third sub- marginal cell). Pygidial area of female (as far as known) with one row of punctures ‚along Margin ,...................0. ect: RL 20 No smooth or coriaceous area behind enclosed area of propodeum, near median sulcus; carinae on back of propodeum extending to enclosed area. Dorsal side and back of propodeum always curved in lateral view. Pygidial area of female densely Pünetate pica inni ATO TIE 63 Smooth (or coriaceous) areas behind enclosed area of propodeum broad, oblong, covering a large part of back of propodeum. Propodeum in lateral view usually gently curved, in some species somewhat angular. Second recurrent vein of fore wings never ending in third submarginal cell. Antennae of male without tyloidea 21 Smooth areas narrower and for the greater part situated on the horizontal part of the propodeum. Propodeum in lateral view more angular, distinctly divided into an almost horizontal dorsal part and an almost perpendicular back part. Second recurrent vein of fore wings in some species ending in third submarginal cell 38 Pygidial area of female broadly triangular, surface coriaceous, one row of punctures along margin. Mesosternum with acetabular carina about as long as half the dis- tance between the epicnemial carinae (in male even longer). Upper part of pro- podeum not coriaceous. (Group:0fP.7/{44 LOI 22 Pygidial area of female elongate triangular, smooth and shining. Acetabular carina 22), 26. Lijs 28. J. P. vAN LitH: Oriental species of the genus Psen 93 shorter (except in male of P. coriaceus and P. heinrich). Group of P. elisabethae BARA NE LL CARE RCI PNA A UE Glare i RT NOIR 23 Gaster, especially first tergite, finer and less densely punctate, pubescence sparser. Legs darker, in female hind tibiae only at knees and apices somewhat brownish. Length about 7.5—9 mm. (Java, Krakatau I., Bangka I., Sumatra, Ceylon, S. India) ARCA LRO Pl Ad RL nitidus nitidus Gaster somewhat stronger pubeseent especially on first tergite, also puncturation there slightly stronger. In female basal third of hind tibiae yellowish-brown. Benesthraboue 83mm: (Formosa) snes nno nitidus takasago Margin of gastral tergites 2—-5 impunctate, this margin broadest medially; in male almost amrew /minutes piliferousi punctures we.) Inn 24 Only margin of fifth tergite impunctate, puncturation on tergites 2—4 almost feachine apical marginy butjsometimes | Very Sparse nt. sn 32 Pétioleandlesstimcludinottrochanters redire PRE CREER nee 25 Pétioleblack#alsonlecstlareelVablackRiOb ro RER ner en 28 Also first two gastral tergites red. Disk of first gastral tergite very sparsely punctate. Pubescence of face silvery, of head and thorax reddish-brown. Length about 10 mm; petiole about seven times as long as wide in the middle. Male unknownia(Solomomtiis) PEER AAA SO. RIA. bishopi sp. n. Gastralitter cites blacks n Tec AE EEE FORTIS SIE, 26 Punctures of scutum relatively coarse, irregularly placed, partly almost contiguous in rows but interspaces often two or three times diameter of punctures. Scutum shining. Antennae entirely black or dark brown. Petiole brighter reddish. Face of female sil- very, rest of body with greyish pubescence. Length about 6.5—11 mm, petiole about seven times as long as wide in the middle. (New Guinea, Papua) .................. eee iaia, Taka ka RARE A AO paulus paulus sp. n. Bene relatively finer. Usually larger species. Either scape of antennae reddish omfacelsomewhatlbrassyaBetiolerdarksreddishue. ern EM O. 27 Scutum with somewhat leaden shine, puncturation fine and regular. Scape reddish. Pubescence of face silvery, of thorax golden-brown. Length about 9.5 mm, petiole about seven times as long as wide in the middle. Female unknown. (Solomon Is.) ARRA senad A SAS: paulus subtilis subsp. n. Puncturation fine on anterior part of scutum, coarser on posterior half. Scape ot antennae dark brown. Face of female brassy, pubescence of scutum brownish. Length about 10—12 mm, petiole about seven times as long as wide in the middle. Male unknown. (New Guinea: Japen I.) ............ paulus baduriensis subsp. n. Puncturation of frons and scutum very coarse (as in P. coriaceus). Post-ocellar area and frons raised. Smooth area of propodeum partly with coriaceous sculpture. Hind basitarsi at least dorsally black-brown, tibial spur of mid and hind legs white. Pygidial area narrow. Face silvery. Length about 10.5 mm, petiole about eight times as long as broad in the middle. Male unknown. (Philippines) … … marjoriae sp. n. Puncturation of frons and scutum much finer. Hind basitarsi usually paler; tibial spurs yellowish-brown ...............0 000 29 AII tarsi yellowish-brown, also apex of hind tibiae and pubescence of vertex and scutum brownish. Both recurrent veins of fore wings ending in second submarginal cell. Length about 9—12.5 mm, petiole about 6—7.5 times as long as wide in themid dle (News Ireland): en Et novahibernicus 94 30. le 33; 34. aps 36. 37. TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Tarsi yellowish-brown or darker, but pubescence of vertex and scutum whitish 30 Tarsi and apex of hind tibiae yellowish-brown as in P. novahibernicus. Enclosed area of propodeum hardly depressed and indistinctly separated from smooth parts of propodeum. Length about 8—9 mm, petiole about seven times as long as wide inthe ymiddlew@Solonmo mils) ET AE vadosus sp. n. Tarsi more or less darkened, apex of hind tibiae dark. Enclosed area normally depressed, sande dee tene AR AN SIE PER EEE 31 Tarsi of male darkened, of female largely testaceous. Fasciculate hairs of third and fourth gastral sternites of male brown. Genitalia of male large. Petiole about 7—8 times as long as wide in the middle. (Samoa) ..............-. ni bryani Tarsi of male testaceous, of female darkened dorsally. Fasciculate hairs of male yellow. Petiole about 7.5 times as long as wide in the middle. (New Hebrides) MIRA RR E e I ee STIER cheesmanae (I could not find any distinct difference in the puncturation of the scutum of P. bryani ( & ) and P. cheesmanae (2 and &) and am unable at the moment to distinguish between the females of these species). . Legs including trochanters red. Petiole dark reddish-brown, first and second gastral tergites laterally somewhat reddish-brown. Pronotal tubercles reddish. Face with appressed golden pubescence, frons and mesosternum with dense, short golden pubescence. Length about 12 mm, petiole 6.5—7.5 times as long as wide in the middle. Male unknowa-+(Sc mR TIE E regalis sp. n. At most tibiae and ‘tarsi reddish, petiolenblack it Tt ver. a een 33 Petiole about eight times as long as wide in the middle. Greater part of femora dark brown or black. Tibiae and tarsi reddish. Scape of antennae dark. Face silvery. Length about 8—10 mm. (Java, Sumatra) .................. elisabethae elisabethae Petiole about five or six times as long as wide in the middle. If petiole much longer, tibiae not reddish and carination on back of propodeum very coarse 34 Areas behind enclosed area of propodeum with fine but conspicuous coriaceous sculpture. Vertex raised. Puncturation of scutum very coarse. Inner side of fore tibiae and all tarsi testaceous (sometimes hind tarsi darkened). Spurs of mid and hind tibiae yellowish-brown. Length about 8—10 mm, petiole about six times as long Vas: awiden in the middle (Philippines) Sr. ann. ee regen cortaceus No coriaceous sculpture. Puncturation of scutum finer ........................... 35 Wettex distinctly raisedsbehindnocellinn att 2.2.2. CERN ne E 36 Vertex mot distinetlyhraisede. nn. PR I ne sees ee Oe eee ees 37 Puncturation of frons and scutum strong, frons and interocellar area raised. Ca- rination on back of propodeum very strong. Tibiae dark brown, tarsi paler brown. Length nearly 12 mm, petiole 7—8 times as long as broad in the middle. Female unknown. h(Gelebesi) we. sys! el EREN DELETE ER heinrichi sp. n. Puncturation much finer. Vertex raised behind ocelli. Tibiae reddish-brown, tarsi paler. Length about 9 mm, petiole about five times as long as broad in the middle. Male unknown.) ((Aimboml an a ee TIERT amboinensis Face silvery in both sexes. Pubescence of propodeum greyish. Pygidial area of female as in P. elisabethae elisabethae. Length about 9—10.5 mm, petiole about five times as long as wide in the middle. (S. India) ......... verversen Eee ROL Aa Pa. N te elisabethae madrasiensis subsp. n. Face of female with golden pubescence. Pubescence of propodeum brownish. Py- 38. 39) 42. 43. 44. 45. J. P. van LITH: Oriental species of the genus Psen 95 gidial area of female slightly broader. Length about 9 mm, petiole about 5.5—6 times as long as wide in the middle. (Malaya, Pulau Tioman) ........ Meen ia OI AE ORI AMES. Oe: elisabethae auricomus Second recurrent vein of fore wings ending well in third submarginal cell. Man- dibles reddish except dark tips. Pygidial area narrow, smooth and shining, a few punctures along the margin. Petiole almost cylindrical. Group of P. rufiventris 39 Second recurrent vein of fore wings interstitial or ending just in third submarginal cell. Mandibles mostly dark, often median part reddish-brown. Pygidial area of female triangular, surface finely reticulate, a few punctures along the margin; if the pygidial area is narrow, the surface is smooth or almost smooth. Petiole cylin- drical or with lateral groove and ventral carina. Group of P. orientalis ...... 44 Basal half of clypeus much raised, gaster red but petiole and first and third gastral tergites for the greater part brown-black. Face pale golden. Back of propodeum with swhitish thairssuMalesunknownt.((Saindia ME e rufiventris Basalwhaliofe clypeuse mot ymuch straisedie ern ta 40 . Petiole red, remainder of gaster red or partly dark brown. Face of female silvery 41 Petiole black, gaster completely black or first tergites somewhat reddish on the sides@Eace ofetemalencol dent try. A ESE ORO Ae nee. FROCIO, 42 Gaster almost completely red, tergites only slightly darkened. Red band before anterior margin of clypeus. Male unknown. (W. Java) ...... rubicundus rubicundus Gaster much darkened, only apical margin and sides of second, third, fourth and fifth tergites ferruginous. Clypeus almost completely dark. Male unknown. (E. VASE seats mans. Lo Gentoo ME ANNEE RNA, di rubicundus lawnensis Back of propodeum not only carinate but also punctate. Gaster completely black. Antennal segments 8—9 about twice as long as broad. Face including frons with dense golden pubescence. Back of propodeum with whitish hairs. Male unknown. (Luzon) écrans ROANNE road AA eN DA dada, nigriventris Bacco fapropodelmenot, punctate. en PR OA 43 Back of propodeum with whitish hairs. Face of female golden, of male silvery. Sides of gaster reddish. Antennae of female short, segments 8—9 about one and one-half times as long as they are broad. Antennae of male without tyloidea. (Japan) ns. Earle. ARI EMED AA, ORSI, 0 RON richard si Back of propodeum with golden-brown hairs. Face golden. Gaster completely black. Lower half of clypeus red. Frons angularly protruding, horizontal side of head long. Antennae long, no tyloidea. Smooth area behind enclosed area of propodeum indistinct. Female unknown. (Mindanao) … an gulifrons Pygidial area of female very narrow, almost gutter-shaped, shining. In female median part of clypeus distinctly raised, anterior margin with a deep triangular emargination. Tarsi red. Face with golden appressed pubescence. In male median part of clypeus not distinctly separated from lateral parts, tarsi dark brown and pubescence of face silvery. Antennal segments 4—13 with tyloidea. Both sexes without distinct smooth area behind the enclosed area of propodeum. Scutum with strong puncturation. Petiole cylindrical. (Java) ..................... terri venus Pygidial area of female slightly or much broader. Median part of clypeus of female less distinctly separated from the sides. Scutum finer punctate in both sexes, except in P. triangulatus. Face of female silvery or, if golden, pygidial area broad ...... 45 Base of second gastral tergite of male with triangular, minutely reticulate area. 96 46. 47. 48. 49. 50. Da TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Scutum shining, with strong punctures in rows. Antennal segments 3—13 with tyloidea. Female unknown.) (Java, Sumatra) Et IE TER triangulatus No, reticulate area on saster. (some males unknown) ron AE 46 Very long acetabular carina, nearly reaching epicnemial carina. Petiole without distinct ventral keel. Antennal segments 7—12 with subelliptic tyloidea. Female unknown... (Ryukyuails-) ans Heee u toe A RER hirashimai Acetabular carinasshortor.absent.... une. stre a MEN CREVER PER PIE 47 Pygidial area of female very narrow, basal half somewhat convex and almost shining, apex emarginate, bilobed, lobes not much raised. Face of female silvery (forimales see also couples 2) Mar ass. RAR 48 Pygidial area broader, more flat and dull, apex almost straight (always?) and ends of lateral keels often distinctly raised into a blunt tooth .................. DZ Petiole without lateral carinae. Anterior margin of clypeus protruding, slightly raised, with two triangular teeth. Legs dark. Male unknown. (N. India) Lerici pelata DI DIREI simlensis sp. n. Petiole in both sexes almost quadrate in cross section, sides with two distinct lateralkeelsic nie Be aka lv ee 49 Smooth area behind enclosed area of propodeum (near median sulcus) narrow. Male with tyloidea on antennal segments 3—12 (on third segment incomplete). Length about 8.5—10 mm. (China, Tibet, Formosa) ...... hakusanus seminitidus Larger. forms kan ria lavabo Lola eta eee err 50 Pygidial area narrower. Puncturation of scutum finer. Smooth area behind enclosed area of propodeum broad. Third antennal segment of female 3.2 times as long as broad at apex, fourth segment almost twice as long as broad. In male antennal segments 3—10 or 3—11 with tyloidea (on third segment incomplete). (Japan) Mellini ile dice ali hakusanus hakusanus Pygidial area slightly broader. Puncturation of scutum stronger ............... 51 Puncturation of scutum remote, stronger than in P. hakusanus hakusanus. Area behind enclosed area of propodeum smooth. Third antennal segment 2.7, fourth segment 1.8 times as long as broad at apex. Length about 12—13 mm. Male unknowal(Sorca)it a koreanus koreanus Puncturation of vertex and scutum finer than in preceding subspecies but still stronger than in P. hakusanus hakusanus. Area behind enclosed area of propodeum with fine longitudinal striation. Third antennal segment three times, fourth segment twice as long as broad at apex. Length about 11.3 mm. Male unknown. (Formosa) Robredo koreanus formosensis Scutum with plumbeous hue, very finely punctate. Petiole in both sexes rounded above, in female with lateral depression and two lateral edges, ventral keel not sharp. Frons of female finely punctate. Antennal segments 5—6 of male with tyloidea. Third and fourth gastral sternites of male with fasciculate hairs. (India) bennen steen id de a atl La pean ee ote eee PEPE TERE orientalis Scutum. with, much. stronger «punctures Wem. eee RENEE DO, Petiole distinctly quadrate in cross section with sharp ventral keel and normally punctate scutum. Antennal segments 5—6 of male with tyloidea. Face silvery in both sexes... Hain an. See SETTE 54 Petiole without distinct ventral keel. Antennae of male without tyloidea or segments 3—9, 3—12, 4—10 or 4—12 with narrow carina (not all males known) ... 55 54. DD: 56. DI Do: D). 60. 61. 62. J. P. VAN LrrH: Oriental species of the genus Psen 97 Frons below ocelli with punctato-striate sculpture. Vertex and scutum finely but distinctly punctate. (Ussuri region, Japan, Korea) .................. affinis affinis Frons below ocelli stronger striate. Vertex and scutum slightly stronger punctate. Pygidial area of female slightly broader. Smooth area behind enclosed area ot propodeum of female slightly broader. In female carination of propodeum stronger. (CHINA) kun era sen IRE O I LORI AE affinis grabami Kegs dark at most tarsı, reddishiorgreddis boy ne ns ae ee 56 Femora and tibiae or tibiae and tarsi red. Face of female golden pubescent 60 Petiole short, less than twice as long as first gastral tergite, sides depressed. Pygidial area of female rather broad, with irregular, basally double row of punctures. Face silvery. Fore and mid tarsi reddish. Male unknown. (S. China) TIA IT SITA tante IA Rene i kulingensis Penolerlongereylindricali.. 33.33 RIE. SI ISS TÀ, SERI, I] DI Face of female silvery. Pubescence of scutum brownish. Most antennal segments about twice as long as broad in the middle, segments 4—12 with tyloidea. Third and fourth gastral sternites of male with fasciculate hairs. (India) fuscinervis Face of female golden (female of P. longicornis Tsuneki unknown). In male antennal segments without tyloidea or segments 3—9 with tyloidea or segments LOTTERIA IE cs. cat oh aa cic CATO OI ARI o INA Lo 58 Scutum with close and strong puncturation. Behind enclosed area a broad smooth margin. Pubescence of scutum brownish. Apex of pygidial area emarginate. In male antennae without tyloidea and only fourth gastral sternite with fasciculate haits: (Burma) nn bid, tens. seis le Haare Sek IT nets oR REEL re Meike yomasanus Scutum with finer puncturation. Smooth margin behind enclosed area narrower. Pubescence of scutum greyish. Antennae of male with tyloidea ............... 59 Apex of pygidial area of female truncate. Puncturation of scutum anteriorly with fine and close puncturation, posteriorly more coarsely punctate. Petiole subcircular or subquadrate, laterally at most weakly furrowed. In male antennal segments 3—9 with tyloidea; fourth gastral sternite only with fasciculate hairs. (Formosa) ...... Sti Mens prod MIR ne PIRO LIO IR ONORI JE His TE alishanus Female unknown. Petiole roughly quadrangular in cross section, laterally with broad furrow margined by carinae. Antennal segments long, 3—12 with tyloidea; third and fourth gastral sternites with fasciculate hairs. (Formosa) longicornis Legs including trochanters pale red, tarsi somewhat darkened, also lower outer side of fore femora. Temples and thorax with whitish hairs. Frons below ocelli striato-punctate. Petiole reaching beyond hind femora. Antennal segments 4-10, rarely 4—11 of male with narrow tyloidea. (Luzon) … politiventris politiventris Outer side of fore femora entirely darkened or all femora darkened ......... 61 Legs red, only outer side of fore femora darkened. Pubescence on vertex, thorax and gaster golden-brown. Frons below ocelli strongly striato-punctate. Petiole reaching beyond hind femora. Male unknown. (Mindanao) … politiventris bellus Femora brown or black. Hairs on propodeum whitish ........................... 62 Femora brown, tibiae and tarsi pale red. Sides and back of thorax with whitish hairs. Frons below ocelli finer punctate than in nominate form, interspaces some- what opaque. Petiole not reaching as far as hind femora, cylindrical. Punctures of pygidial area somewhat larger than in nominate form. Male unknown. (Malaya) UE AA rte AET. elt TI MAI SOIT se etes politiventris pahangensis 98 63. 64. 65. 66. 67. Tike TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Femora black, apices of femora, entire tibiae and tarsi ferruginous to reddish- yellow. Petiole with distinct lateral furrow bordered on both sides by carinae. Antennae of male without tyloidea. Gastral sternites 3—4 of male with fasciculate ee (EIDOS OTO RIT E tanot Scutum except at the sides coarsely striato-punctate or with very strong punctures. Interantennal tooth reduced to a. flat round tuberele”...............2. nme: 64 Scutum shining, withesnormalspunctutation tds. Heten. A I eee ereeE 65 Scutum with strong punctures and also longitudinally striate. Legs reddish. Pygidial area of female triangular, with large punctures. Antennal segments 6—13 of male with broad tyloidea. Face of female golden, of male silvery. (Philippines) ... bakeri Scutum with stronger punctures, interspaces often less than width of one puncture; a few longitudinal striae. Legs, petiole, first gastral tergite and second tergite except dark margin, reddish. Antennal segments 6—13 of male with tyloidea. Face of male silvery. Female unknown. (New Guinea) .................. sedlaceki sp. n. Second recurrent vein of fore wings ending in third submarginal cell. Median anterior part of clypeus of female strongly raised and separated from the depressed lateral parts by a sharp edge. Antennae of male with broad tyloidea. Group ot RSM SIONI TEA 66 Second recurrent vein of fore wings interstitial or ending in third submarginal cell. Median part of clypeus of female not separated from the lateral parts by a sharp edge (a short carina in P. ruficrus) ............- "0" 69 Petiole of female without distinct lateral carinae or groove. Raised part of clypeus short. Only hind margin and sides of first tergite and sides of second and third tergite reddish, in both sexes. Face of female silvery. No smooth area on propodeal deelivity. (CHOO E matalensis Petiole of female with distinct lateral groove. Raised part of clypeus long. In P. refractus (both sexes) propodeum with an almost smooth dorso-lateral area behind TRASCINATA RARA TE 67 Face of female golden. Only sides of first gastral tergite somewhat reddish. Py- gidial area of female broader, less densely punctate. Fifth and sixth antennal segments of male about twice as long as broad. (S. India) ............ krombeini Face of female silvery. First or first and second gastral tergites red. Pygidial area of female narrower, more densely punctate .........:................2.0 scree 68 . First and second gastral tergites red. Male unknown. (India) … refractus refractus First gastral tergite red, second tergite with lateral red spots. Fifth and sixth antennal segments of male about as long as broad. (S. India) refractus meridianus Gaster including “petiole black. enne essere renee erates 70 Petiole red; if black, following segments at least partly red .................. 74 Interocellar and postocellar areas distinctly raised. Antennal segments 6—13 of male with broad tyloidea. No acetabular carina. Second recurrent vein of fore wings interstitial. Face of male silvery. Female unknown. Legs dark. (Philippine Is.) roan ar ANNE IME AAA AH Be nade AGhodeboceaaoadotGccounbonoadsddosoc melanosoma Interocellar and postocellar areas not or only slightly raised. Antennal segments 6—12 or 7—12 of male with tyloidea (male of P. opacus unknown) ...... zi Second recurrent vein of fore wings ending well in third submarginal cell. Propo- deum behind enclosed area with fine, more or less parallel carinae. Pygidial area of female broadly triangular, with four or six longitudinal rows of punctures. Face of female silvery. ……...s.svvorsenonsen veneneneenenenerernenersenenernenosnenen 72 UR UD: 74. 1D: 70. Une 78. J. P. VAN LitH: Oriental species of the genus Psen 99 Second recurrent vein of fore wings interstitial. No parallel carinae behind enclosed ENGEN 6900000000480600000900080000.00960040900040o gun GE 7EHRE SGADBRERTIENE OS RHISDOSCO HUNDEN 73 Legs dark. Second recurrent vein of fore wings ending well in third submarginal cell #Malenunknowa. (Ehilippineals er opacus opacus Legs partly ferruginous. Second recurrent vein of fore wings ending close to base ofgcell Male unknown (Ryukyaipls5) A ende Opacus gressitti Propodeum on both sides behind enclosed area not smooth but with some indistinct carinae and irregular tubercles. Antennal segments 6—12 of male with tyloidea. Female unknown (Morotal) Er ER RE II eee carbonarius Propodeum behind enclosed area in male with normal carination, in female much finer. Pygidial area of female broad with many large punctures, margin impunctate. Antennal segments 7—12 of male with oval tyloidea. Face of female golden, ot MASSIVE KAV) Ine nee A ase di RO an daelen beiremi Petiole and following two or three tergites red. Legs red. Interocellar and posto- cellar areas much raised. Pygidial area of female striato-punctate, margin dull and impunctate. Face of female golden, of male silvery, rest of body with golden-brown pubescence. Length of female about 12—13 mm, of male about 11 mm. (Celebes) CO FOGICE OO AIPIDINTII TO TTOG OOC GO TOO III GIOIA ETA IACGCIORION se descieeisecselsleniuse seen oe toxopewsi Petiole red but no tergites fully red or petiole black and at least second tergite énturelomalmostentirelheredSmallersspeciese 5 ns AN ee 75 Petiole black, gastral tergites 1—4 for the greater part red. Second recurrent vein of fore wings ending well in third submarginal cell. Interocellar and postocellar areaspnotymuchsraised nl en INNS IE 76 Petiole black or red, at most two gastral tergites entirely or partly red. Second KechiEcenty vemeapmostlypainterstitial ues. stanti RINO Tah Face of female silvery. Mesosternum, coxae and femora conspicuously silvery pubescent. Red: gastral tergites 1—4 entirely, fifth tergite partly and sternites 2—3 entirely; legs reddish brown but femora partly darkened. Pygidial area of female broadly triangular, densely and coarsely punctate, also medially, margin reticulate Male unknown: A(Malaya)e eee eee ee brinchangensts Face of female golden. Red: gastral tergites 1—4, first, third and fourth tergites partly darkened, and sternites 2—4. Legs also red, fore and mid femora darkened posteriorly. Pygidial area broadly triangular, coarsely punctate with a narrow median impunctate lines Malesunknown-@N- India) tte eurypygus First and second gastral tergites red. Petiole black. Second and following gastral tergites with conspicuous short white pubescence, hairs longer and somewhat curved backwards at hind margins. Face of female silvery. Pygidial area of female rather broadly triangular, densely punctate, margin impunctate. Male unknown. ON AAE AR RE PA SR INR, MISES ONE TA KURS rufoannulatus Gästen, darkerw pettole»blacc. torte dii BRAIN ROSI des 78 Petiole black. First gastral tergite of female with red apical margin, second gastral tergite with two lateral spots; male with more red on these tergites and on following segments. interocellar and postocellar areas much raised. Second recur- rent vein of fore wings about interstitial. Pygidial area of female densely and almost fully striato-punctate. Antennal segments 6—13 of male with broad tyloidea. Face golden in both sexes, rest of body golden-brown. (Luzon) .................. DREI E I Te D OEE EE aureohirtus aureohirtus 100 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 sin Petiole: red. tis Mi RR EEn ELS LG RE TPE SEE ARE 79 79. Gaster slightly darker but otherwise as in preceding form. Female unknown. (Negroös)) E RIRE RAR ADN aureobirtus rufopetiolatus — Gastral tergites black. Interocellar and postocellar areas not much raised. Second recurrent vein of fore wings usually interstitial. Antennal segments 6—13 of male with tyloidea. Pygidial area of female broadly triangular, with many large punc- tures. Legs pale red. Face of female golden, of male silvery. (N.E. New Guinea and Papua) Ci. VE ET EE NN RTRT ruficrus Psen (Psen) ater (Fabricius) 1794, Fabricius, Entom. Syst. 4: 457 (Sphex atra). In the material of the museum of the Humboldt University, Berlin, I found two females from Manchuria: Harbin, 25—30 June 1935, collected by V. Kardoff. The genitalia of the male of P. ater are relatively large, the stipites are ear-shaped, without inner membranous lobes: fig. 34—35 (undated specimen, probably from Europe). Psen (Psen) aspites spec. nov. Male. — Head and thorax black, scutum with steely reflection, back of propodeum with bronze gloss. Mandibles with dark red tips. Anterior margin of clypeus (about one third) and labrum ivory-white. Palpi testaceous. Scape of antennae dark yellow, following segments yellowish-red, segments 5—11 partly black on underside and last segment almost completely black. Humeral tubercles yellowish-white. Legs reddish, in- cluding coxae and trochanters; some parts of fore legs yellowish-white: trochanters, underside of femora, ends of tibiae and the first four, broadened, tarsal segments except outer tips which are black. Gaster including petiole reddish. Clypeus smooth and shining, anterior margin with a slight emargination. Anterior margin of labrum almost straight. Frontal carina ending below in a slightly thicker part, no distinct tooth. Frons immediately above the antennae shining somewhat depressed to receive the heavy scapes, rest of frons and vertex with fine and remote puncturation. Mandibles normal. Scape of antennae at least one and one-half times as broad as the broadest part of flagellum. Third antennal segment about three times as long as broad, ventrally with a deep longitudinal depression, fourth segment about as long as broad, following segments much shorter than broad, last segment pointed. Segments 4—13 strongly concave ventrally and with sharp sides. Scutum and scutellum with fine and remote puncturation. Enclosed area of propodeum of normal type, median two longitudinal carinae diverging. Behind this area propodeum on both sides somewhat raised and with some fine carination, back of propodeum medially depressed, with rather fine reticulate carination, on either side of the very narrow longitudinal sulcus the carinae are partly transverse and parallel. Sides of pro- podeum, metapleura and mesopleura smooth and shining, hypo-epimeral area distinct. Underside of epicnemial area receding backwards, consequently mesosternum short and fore and hind margins of mesopleura parallel. Fore legs of unusual shape; trochanter nodiform, femur much broadened, underside flattened into a triangle with an angle of about 120 degrees, apical half of tibia somewhat broadened, tarsal segments 1—4 J. P. VAN LITH : Oriental species of the genus Psen 101 enlarged, forming a large thin shield on outer side (fig. 1). Mid tibiae broadened medially. Second recurrent vein of fore wings ending in third submarginal cell. Petiole cylindrical, reaching about as far as end of femora. Gaster with minute and remote puncturation, second sternite on base with a few larger punctures. Upper part of clypeus and supra-clypeal area with silvery appressed pubescence, also frons below ocelli on either side with a ring of apressed silvery hairs enclosing a bare shining spot. Basal half of mandibles with long white hairs. On both sides of hypostoma a patch of long white hairs like a beard. Temples with short silvery pubescence. Vertex with long yellowish-grey hairs. Pronotum, lower part of epicnemial areas, mesosternum and mesopleura with short silvery pubescence. Scutum and propodeum with long, yellow- ish-grey pubescence. Shield of fore tarsus with fringe of fine long and pale hairs. Gaster with sparse pubescence, a few long hairs near the margin of the sternites. Apical margin of sternites 3—4 with golden fasciculate hairs, in two bundles on each sternite. Length about 9 mm. Female unknown. S. Celebes: 1 4 (holotype), Talassa (Maros) 300 m, Oct. 1931, coll. G. Heinrich (ZMB). This is the first Psen known to me with so much enlarged fore tarsi. They remind of the tarsal shield of some Crabronidae. The ivory-white margin of the clypeus is also peculiar. In view of some other characters, such as the course of the epicnemial carinae, the frontal carina and the fasciculate hairs on the gastral sternites, I have provisionally placed this wasp in a separate group of the subgenus Psen. To avoid the risk of irre- parable damage I have not examined the genitalia of this valuable specimen. P. petiolatus Smith of which only the holotype (a female collected in 1864 in Misool or Celebes) is known, appears to be very similar. But according to the notes I made when examining this specimen it has distinct inner epicnemial carinae and acetabular carina and no recurved outer epicnemial carinae, so that I am rather inclined to place it in the subgenus Mimumesa. Group of Psen emarginatus Representatives of this group were already known from Malaya, Java and Japan. P. emarginatus from Java is now also found in Borneo. The area of distribution of the group was further extended by two new species from Nepal and Formosa. They are all closely related, the differences being mostly in the colour of the gastral fringes and of the clypeal pubescence. Psen (Psen) emarginatus Van Lith 1959, Van Lith, Zool. Verh. Leiden 39 : 43—44 (Java). New record: 1 2, British N. Borneo, Mt. Kinabalu, Kambaranga, 2140 m, 22—30 Oct. 1958, coll. T.C. Maa (BISH). This is the first record of the genus Psen from Borneo. The wasp was collected together with its prey, an Issid (Homoptera). As far as I can see it is not different from the females from Java. Pygidial area of one of the paratypes (female) from Central Java is depicted on fig. 3. 102 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 aspites nepalensis ruficrus sauteri ruficrus Fig. 1—7. Species of Indo-Australian Psen s.str. 1, fore tarsus of P. aspites sp.n., 6; 2—5, pygidial area of female; 2, P. nepalensis sp.n.; 3, P. emarginatus Van Lith; 4, P. sauteri sp.n.; 5, P. ruficrus Van Lith; 6—7, anterior margin of clypeus; 6, P. sauteri sp.n.; 7, P. ruficrus Van Lith. J. P. VAN LITH: Oriental species of the genus Psen 103 Psen (Psen) nepalensis spec. nov. Female. — Black; outer half of mandibles dark red, palpi yellowish-brown, under- side of antennae dark brown. Tarsi brown. Tibial spurs of hind legs yellowish-white. Veins of wings and tegulae dark brown. Protruding median part of clypeus with rounded emargination. Disk of clypeus regularly convex, densely punctate, margin impunctate. The interantennal carina ends between the antennae in a short tooth, indistinctly connected with the antennal sclerites by a fine carina. Frons coarsely punctate with narrow, shining interspaces, on the upper part against the eyes an elliptic, slightly raised, almost impunctate area, vertex sparsely punctate but more strongly and denser behind the ocelli. Mandibles normal. Antennae long, third segment about five times as long as it is broad at the apex, segments 4—6 more than twice as long as broad, segments 7—11 less than twice as long as broad, gradually shorter, last segment about twice as long as it is broad at the base. Scutum with strong punctures, these sometimes in rows, but interspaces mostly greater than diameter of punctures. Puncturation of scutellum and metanotum much more widely spaced. Enclosed area of propodeum depressed, shining, with longitudinal carinae, median two slightly diverging and with some irregular carination in between. Back of propodeum with coarse irregular carination, which is finer just behind the enclosed area. A bent carina separates the back of the propodeum from the sides which show some fine rugae and sparse fine punctures on their posterior half. Mesopleura sparsely and finely punctate, posterior margin smooth, metapleura entirely smooth. Back of hind femora with median longitudinal line of extremely fine punctures with short hairs. Above this line the posterior surface of the femora is completely smooth and shining, lower half shining with a few long hairs. Mid tibiae normal. Hind tibiae with a row of white spines. The first recurrent vein of the fore wings ends well in the second submarginal cell, the second recurrent vein ends about half this distance into the third submarginal cell. Petiole nearly flat above, rounded below, sides with sharp longitudinal edge above from base to end and a shorter one below on the apical half. Gaster finely but distinctly punctate, second tergite laterally at base much coarser. Pygidial area broadly triangular with rounded apex, surface with fine reticulate sculpturation, median part somewhat convex and with rows of large punctures which are open towards the rear. Broad margin of pygidial area without punctures, also the narrow median line is impunctate (fig. 2). Face with very pale golden pubescence and a number of long, erect whitish hairs. Pubescence on upper side of head and body brownish, whitish on ventral side. Pubes- cence of gastral tergites, especially of third and fourth tergites, dense and long. Poste- rior margin of tergites with dense fringe of golden-brown, slightly bent hairs. Pygidial area and sides of last tergite with stiff hairs, directed backwards. Length about 9 mm. Male unknown. East Nepal: 1 9 (holotype), Taplejung distr. above Sangu, 6500 feet, evergreen scrub, 5—13 Oct. 1961, R. L. Coe, E. Nepal Expedition 1961-1962, BM 1962— 177 (BM). This wasp belongs to the group of P. emarginatus and resembles the equally strongly pubescent Japanese species P. dzimm, from which it differs in the darker first gastral 104 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 tergite and legs and in the smaller size. There is also some difference in the relative length of the antennal segments. Further studies of both sexes are necessary to ascer- tain the relationship between the two forms. The Javanese P. emarginatus is much less densely pubescent and the fringes on the hind margin of the tergites are less conspicuous. The tibial spurs of the fore and mid legs are whitish with a small brown apex, whereas in P. nepalensis these spurs are completely yellowish-white. There is no important difference between the pygidial areas of P. nepalensis and P. emarginatus (fig. 2 and 3). The punctures of P. emarginatus are larger and they are more in longitudinal rows whilst the irregular longitudinal im- punctate areas are slightly raised. P. sauter: from Formosa is larger, the pubescence is paler and its antennal segments are longer. In P. pilosus from Malaya the fringes on the hind margin of the tergites are black. Psen (Psen) sauteri spec. nov. Female. — Black, gaster with bluish shine; median part of mandibles yellowish-red, outer tips dark red, anterior margin of clypeus reddish transparent, labrum paler red, underside of antennae brown, palpi testaceous. Tegulae reddish-brown. Femora dark brown, fore side of fore and mid tibiae and base of hind tibiae paler brown, tarsi yellowish-brown but hind tarsi darkened dorsally, tibial spurs yellowish-brown. Veins of wings brown. Protruding part of clypeus with triangular emargination (fig. 6). Surface of clypeus densely punctate, impunctate margin with fine transverse striation. Interantennal carina ending between antennae in a distinct tooth, which is connected with the underside of the antennal sclerites by a distinct carina. Frons below ocelli densely and coarsely punctate, on the upper part against the eyes an elliptic, slightly raised, impunctate area. Vertex finely and sparsely punctate, interocellar area somewhat denser punctate. Man- dibles normal. Antennae long and slender, third segment more than five times as long as broad at apex, fourth and fifth segments about three times as long as broad, segments 6—8 twice as long as broad, segments 9—11 nearly twice as long as broad, last segment more than twice as long as it is broad at base. Scutum with coarse punctures, often in rows, interspaces mostly smaller than diameter of punctures, or as large as diameter of punctures, on central part of scutum a few rugae with rows of punctures. Puncturation of scutellum and metanotum much finer and sparser. Enclosed area of propodeum rather long, with longitudinal carinae. Posterior half of median two carinae somewhat diverging, basal half between these carinae finely irregularly carinate. Back of propodeum somewhat rounded in lateral view with coarse reticulate carination which is finer just behind enclosed area, antero- dorsolaterally a small, almost smooth area. A bent carina separates the back of the propodeum from the sides, which have a few short carinae and are finely punctate on the posterior part. Mesopleura with sparse and fine puncturation, mesosternum with fine and dense puncturation, metapleura smooth. Back of hind femora smooth, with longitudinal line of fine piliferous punctures. Mid tibiae slightly bent but not much thickened; a row of four long yellowish thorns on outer half and two slightly shorter thorns at apex, basitarsus about four times as long as broad with a comb of five long spinés. Hind tibiae with a row of short brownish thorns. The first recurrent vein of the J. P. van LITH: Oriental species of the genus Psen 105 fore wings ending well in the second submarginal cell, second recurrent vein ending about half this distance into third submarginal cell. Petiole flattened on posterior dorsal half, sides flattened but without sharp carinae. Gaster densely and finely punctate, second tergite at base laterally coarser, narrow apical margin densely punctate. Pygidial area (fig. 4) broadly triangular with rounded apex, surface with fine coriaceous sculpture and with about six irregular rows of large punc- tures; margin impunctate, also base medially where sculpture is finer. Second gastral sternite with dense and rather strong puncturation, median part somewhat depressed and shining, with only a few punctures on its basal half, also base of this tergite impunctate. Median part of ventral disk of first sternite (petiole) impunctate. Hind margin of gastral sternites 3—5 impunctate. Last sternite with smooth keel on apical half. Face with beautiful golden, appressed pubescence and a few long, whitish, erect hairs. Pubescence on vertex and on dorsal side of thorax light brown. Temples, dorsal side of pronotum, mesosternum and underside of legs with silvery-white pubescence. Propodeum with long whitish hairs. Gastral tergites 1—5 with yellowish-grey, dense and soft pubescence, these hairs partly rather long, hind margins of tergites with a dense row of longer yellowish hairs, directed backwards and with bent tips. Pygidial area and sides of last tergite with long, backwards-directed bristle-hairs. Length about 12 mm. Male unknown. Formosa: 1 @ (holotype), Taihorin, Aug. 1910, coll. H. Sauter (ZMB). P. sauteri differs from P. dzimm Tsuneki (Japan) in the dark first tergite and paler apical fringes of the tergites, from P. emarginatus Van Lith (Java and Borneo) in the denser pubescence and golden face, and from P. nepalensis Van Lith (E. Nepal) in the paler fringes, longer antennal segments and denser puncturation of the scutum. P. pilosus Van Lith (Malaya) has black apical fringes of the tergites. Group of Psen tsunekiz Psen (Psen) shirozui Tsuneki 1966, Tsuneki, Etizenia Fukui Univ. 14: 10—11 (Formosa). Tsuneki placed this species in the group of P. tsunekii Van Lith. It is indeed closely related to P. vechti Van Lith, also in regard to the structure of the propodeum. Both forms have small transverse depressions at the posterior margin of the enclosed area ot the propodeum, which are deeper than the parts of the enclosed area between the longitudinal carinae. In the description of P. vechti this remarkable character was omitted. The longitudinal carinae of the enclosed area pass between these transverse depressions, continuing on to the back of the propodeum. In P. vechti they are very regular, parallel, surpassing the central part of the back. In P. shirozui the carinae seem to be shorter and less parallel, but of this species only the male being known and of P. vechti only the female, it is uncertain whether this is a matter of sexual difference. Group of Psen nitidus Psen (Psen) nitidus nitidus Van Lith 1959, Van Lith, Zool. Verh. Leiden 39 : 28—30. P. nitidus was first recorded from Java, Bangka and Krakatau, moreover one female was mentioned from Ceylon. Its occurrence in S. India is now well confirmed, nine 106 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 specimens having been collected there. The discovery of two females, collected in C. Sumatra is therefore not surprising. I could not find any important difference between the Indian and the Indonesian material, except some variation in the colour of the legs. The base of the hind tibiae, however, is not as pale as in the subspecies from Formosa. In the female from Cinchona the median two carinae of the enclosed area are less diverging so that the area in between is narrower, but probably this is a deformation. The genital apparatus of the Indian male agrees with that of the Indonesian spec- imens. New records from S. India: 2 © and 1 &, Kerala State, S. Malabar, Walayar Forests, 1000 ft, Sept.-Oct. 1955/1956, coll. P. S. Nathan (Coll. Ferguson); 1 9, Walayar Forests, 700 ft., Oct. 1959; 2 4, Madras State, Nilgiri Hills, Devala, 3200 ft., Oct.- Nov., 1960; 2 4, Madras State, Anamalai Hills, Kadamparai, 3500 ft., May 1963; 1 9, Anamalai Hills, Cinchona, 3500 ft., May 1960, all coll. P. S. Nathan (ML). Northeast Sumatra: 2 2, Bandar Baru, 30 May 1912, coll. Buttel-Reepen (ZMB). Psen (Psen) nitidus takasago Tsuneki 1967, Tsuneki, Etizenia Fukui Univ. 24 : 8—9. Tsuneki considers this form from Formosa as a subspecies. The bases of the hind tibiae of the female are yellowish-brown (dark brown in the nominate subspecies) and the pubescence, especially on the first gastral tergite, is somewhat denser, the punc- turation on this tergite being slightly stronger than in the nominate subspecies. The differences are, however, very small and the colour of the base of the hind tibia of the female is variable. New records from Formosa: 1 9, Oct. 1910, and 4 ¢, April and Sept. 1910, Tai- horin, coll. H. Sauter (ZMB). Group of Psen elisabethae P. elisabethae (Java) and a closely related species, P. coriaceus (Philippine Is.), were described in 1959. They are characterized by the smooth areas of the propodeum and the shining, elongate-triangular pygidial area of the female. The antennae of the males have no tyloidea. Later I found that the genital apparatus of the male of P. elisa- bethae is distinctly different from that of the other species of the subgenus Psen. In the course of subsequent studies I recognized at least fourteen more forms which should be placed into this group. The group has a very extended area of dispersal but the differences between the species are still relatively slight. At the moment they are known to occur from India to Samoa, and from the Philippine Is. to Java. It is one of the most important groups of species in the Indo-Australian and Pacific region and certainly the most interesting one. A number of islands are still to be explored and also the southeastern part of continental Asia. I expect that collectors on future expeditions will discover a great many new forms which will enable us to draw more reliable con- clusions in regard to the specific or subspecific status of the forms described so far. Psen (Psen) elisabethae Van Lith A male collected in Viet-Nam, Mt. Lang Bian, 1500—-2000 m, 19 May—8 June 1961, in a Malaise trap by N. R. Spencer (BISH), could not yet be classified satis- factorily. It may be a new subspecies. J. P. van LiTH: Oriental species of the genus Psen 107 Psen (Psen) elisabethae auricomus Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 31 (Malaya, Pulau Tioman). A drawing (fig. 9) of the pygidial area of one of the paratypes ( 9 ) is given here to complete the original description. Psen (Psen) elisabethae madrasiensis subspec. nov. 1965, Van Lith, Zool. Verh. Leiden 73 : 31—32 (P. elisabethae subsp.). In 1965 I briefly mentioned a female of P. elisabethae subsp. from S. India. The study of another female and two males from the same region has proved that the S. Indian form is indeed quite different from the nominate subspecies. The S. Indian specimens are easily distinguished from the nominate subspecies by the much shorter petiole and the heavier legs. In this respect they resemble P. amboinen- sis in which, however, the median part of the vertex is much more raised while the puncturation of vertex and scutum is weaker; moreover the pygidial area of the female of the latter is broader (fig. 10). In P. elisabethae madrasiensis the pygidial area (fig. 11) resembles that of the nominate subspecies (fig. 8). Female. — First two antennal segments reddish (in the Javanese specimens these parts usually almost completely very dark brown). Base of flagellum ventrally reddish or brown. Legs darker than in the nominate form but fore side of fore femora and apices of all femora reddish-brown; mid and hind tibiae partly, mostly on fore side, dark brown, also thorns on outer side of hind tibiae dark. Fore tibiae almost completely red (in P. elisabethae auricomus from Malaya on the contrary, the legs are paler than in the nominate subspecies, the femora being also reddish or reddish-brown). Puncturation of vertex stronger, median part only slightly raised. Puncturation ot tergites, notably of second tergite somewhat finer, but still distinct and reaching to outer apical margin. Petiole much shorter, about five times as long as broad in the middle, the basal part before the bend about one third of total length, in the nominate form approximately one-fourth of total length. Pubescence of face silvery (in P. elisabethae auricomus pubescence distinctly golden). Length about 10.5 mm. Male. — Underside of antennae more brown. Legs much more slender than in female. Easily distinguished from nominate subspecies by shorter petiole. Length about 9 mm. South India: 1 @ (holotype), Madras State, Nilgiri Hills, 3500 ft., coll. H. L. Andrews (OUM); 2 4 (allotype and paratype), Madras State, Anamalai Hills, Cinchona, May 1959, coll. P. S. Nathan (Coll. Ferguson); 1 9 (paratype), Madras State, Anamalai Hills, Kadamparai, 3500 ft., May 1963, coll. P. S. Nathan (ML). Psen (Psen) amboinensis Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 32—33. The third known female, recorded here, seems to confirm the constancy of the characters mentioned in the original description. This strengthens my opinion that the present form represents a distinct species, closely related to P. elisabethae s.str. Ambon I.: 1 2, Oct. 1949, coll. M. A. Lieftinck (ML). 108 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 elisabethae auricomus amboinensis madrasiensis marjoriae coriaceus regalis Fig. 8—14. Pygidial area of female of species of Indo-Australian and South Pacific Psen s.str. 8, P. elisabethae elisabethae Van Lith; 9, P. elisabethae auricomus Nan Lith; 10, P. amboinensis Van Lith; 11, P. elisabethae madrasiensis subsp.n.; 12, P. marjoriae sp.n.; 13, P. coriaceus Van Lith; 14, P. regalis sp.n. J. P. van LitH: Oriental species of the genus Psen 109 Psen (Psen) heinrichi spec. nov. Male. — Black; palpi yellowish-brown, fore side of fore and mid tibiae and tarsi of all legs light brown, spurs of hind tibiae yellowish-white. Veins of wings dark brown. Anterior margin of clypeus with slight emargination and two blunt teeth. Clypeal disk densely punctate. Interantennal carina ending between antennae in a low tooth, flattened laterally and rounded in lateral view. This tooth is connected with the under- side of the antennal sclerites by a fine carina. Frons distinctly punctate, a small impunc- tate area against the eyes; frons much raised on both sides of the median carina, raised parts confluent with raised interocellar area. A sulcus connects depression of anterior ocellus with transverse sulcus behind posterior ocelli. Vertex with sparse but deep punctures, area behind ocelli distinctly raised. Mandibles normal. Third antennal segment three times as long as it is broad at apex, fourth segment twice as long as it is broad at apex, following segments gradually shorter, twelfth segment as long as broad, last segment about one and one-half times as long as it is broad at base, no tyloidea. Scutum shining with deep punctures, interspaces equal to or twice as large as diameter of punctures, sometimes larger. Puncturation of scutellum sparser, metanotum with finer puncturation. Enclosed area of propodeum deep, with oblique longitudinal carinae, median two carinae diverging. Lower part of longitudinal sulcus on back of propodeum broad. Smooth area on both sides behind enclosed area not large, about twice as broad as long, towards the longitudinal sulcus distinctly raised. Back of propodeum with very strong reticulate carination, enclosing some large smooth multi-angular areas. Sides of pro- podeum almost smooth, separated from carination of back by a long bent carina. Meso- pleura and mesosternum with fine puncturation. Acetabular carina about half as long as distance between ends of epicnemial carinae. Epicnemial carina connected with meso- pleural signum. Interepicnemial area not with distinct transverse striation. Legs normal, back of hind femora smooth with longitudinal row of fine piliferous punctures, upper half bare, lower half with a few long hairs. Both recurrent veins ending in second submarginal cell. Petiole flattened dorsally, sides depressed with short upper basal keel and distinct lower keel. Ventrally an indistinct keel posteriorly and a few punctures, each bearing a long hair. Length of petiole about 7--8 times the width in the middle. Gastral tergites with fine and wide-spread puncturation; puncturation much denser at margin of first tergite and sides of base of second tergite. Fifth tergite with smooth margin. Second gastral sternite shining with a few hair-bearing punctures. Appressed pubescence of face silvery, also with a few long erect hairs. Rest of body with a greyish-white pubescence which is longest on back of propodeum. Apical margin of first tergite and sides of base of second tergite with dense and short whitish pubescence; following tergites with a few long hairs on disk. Apical margin of sternites with a row of long and stiff hairs, on disk some shorter pubescence. Last sternite with dense brownish pubescence, laterally margined by a row of piliferous punctures. Apical margin of third and fourth gastral sternites medially with long erect fasciculate hairs, on each sternite divided into four separate bundles. Length about 12 mm. Female unknown. S.W. Celebes: 1 4 (holotype), Bonthain, Wawa Karaeng, 1100 m, Sept.-Oct. 1931, coll. G. Heinrich (ZMB). 110 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 The pubescence of the gastral tergites is not dense as in P. nitidus, but the smooth areas of the propodeum are rather short and the acetabular carina is long. Although these latter two characters would suggest the group of P. nitidus, I prefer to place this male provisionally into the group of P. elisabethae. The decision whether it is correctly placed or whether it belongs to the group of P. nitidus, depends upon the structure of the pygidial area of the female. Psen (Psen) regalis spec. nov. Female. — Head and thorax black. Antennae reddish (dorsally slightly darkened), mandibles reddish except dark tips, palpi testaceous, anterior margin of clypeus reddish- brown. Tegulae and part of pronotal tubercles reddish. Veins of wings reddish-brown. Legs including trochanters and apices of coxae red. Petiole, sides of first and second gastral tergites, ventral plate of petiole and second sternite dark reddish. Clypeus with slight emargination, densely punctate, anterior margin with fine trans- verse striation. The interantennal carina ends below in a distinct tooth which is connected with the antennal sclerites by fine carinae. A small smooth area near oculi and vertex somewhat raised. Frons between ocelli and antennae with fine puncturation, frons laterally and also vertex with remote fine puncturation. Mandibles normal. Scutum densely punctate, punctures very fine on anterior third part and at the sides, stronger on posterior part, interspaces there sometimes less than diameter, but usually larger than diameter of punctures. Scutellum with remote coarse punctures, metanotum much finer punctate. Enclosed area of propodeum normally depressed, between median two carinae some irregular rugae, behind each lateral part of enclosed area a distinct smooth area, back of propodeum with very coarse reticulate carination. Mesopleura with indistinct puncturation. Metapleura smooth. Short acetabular carina, in front ot ‘and behind this carina some fine transverse striation. Back of hind femora with longitudinal line of many fine punctures, each with a very short fine hair. Outer side of hind tibiae with a longitudinal row of short thorns. Second recurrent vein of fore wings ending in second submarginal cell near apex. Petiole almost cylindrical, slightly flattened dorsally and anterolaterally, about 6.5— 7.5 times as long as wide in the middle. Gastral tergites with fine puncturation, inclu- ding hind margin, except on hind margin of first tergite where the puncturation is dense. Hind margin of fifth tergite impunctate, bordered by a few large punctures. Pygidial area elongate-triangular, shining, a few punctures along the lateral margins (fig. 14). Gastral sternites with a few strong punctures in front of the impunctate hind margin. Pubescence of face golden, appressed and also with a few long hairs. Head and thorax with long golden-brown hairs, mesosternum with dense, short, golden pubescence. Gastral tergites with short golden-brown pubescence; a few long hairs on disk of second gastral sternite, following sternites with a few long hairs arising from large punctures along the impunctate hind margin. Length about 12 mm. Male unknown. Solomon Islands: 1 9 (holotype), Vella Lavella, Ulo crater, 10 m, 17 Dec. 1963, 1 2 (paratype), Kolombangara, Gollifer’s Camp, 700 m, 23 Jan. 1964, both coll. P. Shanahan, Malaise trap (BISH). J. P. van LiTH: Oriental species of the genus Psen dl This beautiful species which certainly belongs to the group of P. elisabethae, is easily recognized by its golden face and the brown petiole. Psen (Psen) coriaceus Van Lith 1959, Van Lith, Zool. Verh. Leiden 39 : 34—36. The female which was recorded in my earlier paper (1959) as a paratype of P. coriaceus from Mt. S. Tomas, is now considered as a new species: P. marjoriae. A more detailed figure of the pygidial area of the female of P. coriaceus is given here (fig. 13, holotype). The acetabular carina is shorter than in P. »itidus, but relatively long in comparison with other Psen. In the male this carina is even longer than half the distance between the epicnemial carinae. The scutum of the female from Negros, recorded in 1959, is much finer punctate than in the holotype; a male from the more southern Mindanao Island, on the contrary, is more coarsely punctate than the allotype from Mindoro. I am hesitating to consider these specimens as separate subspecies, as long as we have no further material available. Psen (Psen) marjoriae spec. nov. 1959, Van Lith, Zool. Verh. Leiden 39 : 34—36 (partim). Female. — Black; scutum, scutellum and metanotum with somewhat plumbeous shine. Fore side of fore tibiae and tarsi (except base of basitarsus) of fore and mid Jegs brown. Ventral side of hind basitarsus brown, following tarsal segments apically somewhat brownish. Apical spurs of mid and hind tibiae whitish. Veins of wings dark brown. Clypeus with broad margin which is emarginate as in P. coriaceus and dull, im- punctate; disk of clypeus densely punctate. Interantennal tooth as in P. corzaceus. Cen- tral part of frons irregularly raised and coarsely punctate, sometimes punctures in dis- tinct rows; area near eyes smooth, not distinctly raised as in coriacens. Vertex much finer punctate, postocellar and interocellar areas raised. Antennae gradually thickened towards apex. Scutum shining, coarsely punctate, punctures sometimes in rows. Posterior two-thirds of tegulae smooth. Scutellum more sparsely punctate; metanotum almost impunctate. Epicnemial carinae as in coriaceus. Acetabular carina short. Epicnemial areas smooth, less distinctly separated from inter-epicnemial area, which is finely and densely punctate. Enclosed area of propodeum somewhat depressed, shining but with traces of coriaceous sculpture; distinct longitudinal carinae. Smooth areas behind enclosed area partly with coriaceous sculpture, not as distinct as in coriaceus. Posterior part of propo- deum irregularly, not very coarsely, carinate. Sides of propodeum shining with some fine sculpture along posterior carina. Hind tibiae dorsally with a row of fine, black spines. Second recurrent vein of fore wings ending in second submarginal cell. Petiole about eight times as long as broad in the middle, cylindrical, dorsally slightly flattened. Hind margin of gastral tergites 2—5 smooth, impunctate, broadest medially and gradually narrowing towards the sides. A few punctures stronger than those on the disk border the impunctate margin of the fifth tergite. Rest of tergites finely, not densely punctate. Greater part of second gastral sternite smooth, almost impunctate and with broad smooth margin; following sternites finely reticulate at base. Last segment with a few large punctures laterally. Pygidial area as in P. elisabethae elisabethae but punctures smaller (fig. 12). 112 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Face with silvery, appressed pubescence and also with a few long erect hairs. Rest of body with greyish-white pubescence, on scutum yellowish-grey. Punctures of pygidial area each with backwards-directed bristle. Last tergites, and all sternites, with a few long and stiff hairs separating the smooth apical margin from the disk. Base and apex of hind tibiae with dense white pubescence on inner side. Length about 10.5 mm. Philippine Is.: 1 2 (holotype), Mt. Sto. Tomas, 7200 ft, near Baguio (Luzon), 27 Dec. 1952, H.M. and D. Townes (coll. Townes). This specimen closely resembling P. coriaceus was thus far mistaken for the latter as I did not earlier pay attention to the puncturation of the gastral tergites and their impunctate margins. The same broad impunctate margin of the gastral tergites was also found in a few more species of the group of P. elisabethae and of P. rufiventris. P. marjoriae is easily distinguished from P. coriaceus by the puncturation of the tergites and by the whitish tibial spurs. According to information received from Dr. Clare R. Baltazar the temperature on Mt. Santo Tomas, which is about 7200 feet high, is sometimes as low as 36—45 degrees Fahrenheit and temperate or subtemperate trees belong to its flora, such as naturally growing pines and various species of oak. Mt. Makiling, where P. coriaceus has been found, is lower, about 300 feet, the tem- perature near the summit is usually between 70 and 80 degrees and the flora is quite different. A female from Baguio, 15 km away from Mt. Sto. Tomas (there is no record of the altitude at which this specimen has been taken) agrees with the description of P. coriaceus. The hind basitarsi are dark dorsally but all tarsi are generally paler than in P. marjoriae. The hind margins of the gastral tergites are richly punctured. I regret that only one specimen of P. marjoriae is known thus far and I have been hesitating about its systematic status. However, especially in view of the conspicuous impunctate margins of the gastral tergites, I feel justified to describe it as a distinct species. It is to be hoped that more and ample material will become available in the near future. I have the pleasure to dedicate this species to Dr. Marjorie Townes who, together with the other members of the Townes family, collected so many interesting Philippine Psenini. Psen (Psen) novahibernicus Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 33—34. The original description was based on three males. Here follows the description of the female, which was collected together with the holotype. Black, greater part with bluish shine. Mandibles somewhat reddish near apex, fore side of fore tibiae (also in male), ends of all tibiae and all tarsi yellowish-brown, basitarsi somewhat paler, tibial spurs straw-yellow. Veins of wings black. Anterior margin of clypeus as in male. Interantennal carina and puncturation of frons and vertex as in male. Postocellar and interocellar area slightly raised. Structure of scutum, scutellum and propodeum as in male, back of propodeum with few carinae, so that a few large enclosed areas are formed, each with very fine sculpture on the surface. Precoxal suture connected with subpleural signum. Legs less slender. Both J. P. van LITH: Oriental species of the genus Psen 113 recurrent veins of fore wings ending in second submarginal cell. Petiole about six times as long as it is broad in the middle, anterior half laterally somewhat flattened, also dorsally somewhat flattened, no distinct lateral carinae, underside rounded, thick- ened apically. Gaster very shining. First tergite very sparsely punctate on disk, apical margin very finely punctate. Tergites 2—5 sparsely punctate, apical margins impunctate, separated from the disk by a line of piliferous punctures, these punctures stronger on fourth and fifth tergites. Pygidial area narrow (fig. 15), laterally with a row of punc- tures, each with a bristle pointing backwards. Sternites 2—4 sparsely punctate with smooth margin which is bordered laterally by a line of punctures, each with a back- wards-directed bristle. Second sternite strongly shining, fifth sternite with fine coriaceous sculpture, more densely punctate. Sixth sternite densely punctate, finely so on base, coarsely on apical part, medially with an impunctate keel. Length about 12.5 mm. New Ireland: 1 2, Schleinitz Mts., Lelet Plateau, Oct. 1959, coll. W. W. Brandt (BISH). Psen (Psen) paulus spec. nov. Female. — Black, palpi testaceous, legs including trochanters and petiole reddish. Underside of antennae brown. Mandibles reddish-brown. Labrum red. Tegulae and veins of wings dark brown. Clypeus densely punctate, broad anterior margin finely transversely striate and weakly emarginate. The fine interantennal carina ends between the antennae in a distinct tooth which is connected with the antennal sclerites by fine carinae. Frons distinctly raised on either side of the median carina, remotely punctate. Vertex with fine and sparse puncturation, postocellar area slightly raised. Mandibles normal. Antennal segments 9—11 about as long as broad, last segment slightly more than one and one-half times as long as its breadth at base. Scutum with relatively coarse punctures, often in rows with interspaces much smaller than diameter of punctures, otherwise interspaces more than diameter. Anteriorly and antero-laterally the punctures are much finer. Scutellum finer and more sparsely punctate, metanotum still finer. Enclosed area of propodeum distinct, a few oblique longitudinal carinae, median two carinae much diverging. Behind the smooth areas some coarse reticulate carination. Mesopleura with indistinct puncturation, metapleura smooth. Legs normal, hind tibiae with a row of thorns on outer side. Second recurrent vein of fore wings ending near the end of the second submarginal cell. Petiole cylindrical, dorsally somewhat flattened, about seven times as long as broad in the middle. Gastral tergites with sparse puncturation, hind margins of segments 2-5 impunctate. Pygidial area (fig. 18, holotype and 19, Hollandia) elongate-trian- gular, shining, with a few punctures along the margin. Sternites with sparse puncturation. Sixth sternite with a few large punctures. Face silvery pubescent with a few long erect hairs, vertex and scutum with greyish pubescence, propodeum with long whitish hairs, gaster with sparse fine pubescence. Gastral tergites 4 and 5 and all sternites with a few long hairs near hind margin. Sixth sternite with dense, short brownish pubescence. Male. — Apical thorn brown. Hind tibiae without thorns. Hind margins of tergites 2 etc. with a few fine punctures. Gastral sternites 3 and 4 with long dark brown fasci- culate hairs at hind margin in two bundles on each segment. Genitalia Fig. 36—37. 114 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 New Guinea: 1 ® (holotype), N.E. New Guinea, Wau, Morobe District, 1050 m, 16 Oct. 1961, Malaise trap, coll. J. Sedlacek; 1 4 (allotype), Karimui, 1080 m, 14 July 1963, coll. M. Sedlacek (BISH); 1 2, Hollandia, July 1938, Neth.-Ind—American New Guinea Exp. 1938—39, coll. L. J. Toxopeus (ML). Papua: 1 9, Inonda, Horanda District, April 1943, coll. W. G. Bodenstein (USNM); 1 4, Mondo, 5000 ft., Febr. 1934, coll. L. E. Cheesman, BM 1934—321 (BM). The name paulus is used because of the small size of both holo- and allotype, which have a length of about 6.5 mm. Afterwards I received a male from Papua of about 8 mm and two females from Papua and Hollandia of about 10.5 and 11 mm. This great difference in size is rather unusual. The male from Papua differs in a few minor details from the allotype: the fine puncturation of the scutum antero-laterally is denser and the pubescence of the scutum is somewhat brownish. P. paulus s.str. is easily recognized by the red legs and petiole and the coarse punc- turation of the scutum. The subspecies from Japen Island has a brassy face and finer punctures and the subspecies from the Solomon Islands has a much finer puncturation of the scutum (pubescence of face silvery). Psen (Psen) paulus baduriensis subspec. nov. Two females from Japen Island differ distinctly from the females of the nominate form from New Guinea and Papua. They are larger (length about 10 and 12 mm), the antennal segments are slightly longer (segments 9—11 about one quarter longer than they are broad at base; last segment about three quarters longer than broad at base); the puncturation of the scutum is somewhat finer than in the nominate subspecies and the appressed pubes- cence of the face is more brassy than silvery. Petiole about seven times as long as wide in the middle. Pygidial area Fig. 20, holotype. New Guinea: 2 2 (holotype and paratype), Japen Island, Mt. Baduri, 1000 ft, August 1938, coll. L. E. Cheesman, BM 1938—593 (BM). Psen (Psen) paulus subtilis subspec. nov. Male. — Differs from the nominate subspecies in the reddish colour of the scape of the antennae, the finer puncturation and the leaden shine of the scutum, and the brownish hairs of head and thorax, the long hairs of the propodeum being very con- spicuous. Genitalia Fig. 40—41. Length about 9.5 mm. Female unknown. Solomon Islands: 1 4 (holotype), Kolombangara, Collifer's Camp, 700 m, 23 Jan. 1964, Malaise trap, coll. P. Shanahan (BISH). Psen (Psen) vadosus spec. nov. Female. — Black with a slight metallic-blue shine; fore side of fore tibiae and ex- treme apices of all tibiae brown, tarsi yellowish-brown. Middle part of mandibles dark reddish-brown. Palpi testaceous. J. P. van LITH : Oriental species of the genus Psen 115 Clypeus convex, with dense superficial puncturation and broad, almost smooth, im- punctate margin which is slightly emarginate. The interantennal carina ends between antennae in a low tubercle connected with underside of antennal sclerites by a fine carina. Frons with some puncturation on both sides of median carina, against the eyes a slightly raised smooth area, on either side of frontal carina frons somewhat protruding. Vertex with fine and remote punctures, postocellar area raised. Mandibles normal. An- tennae gradually thickened towards apex, segments 9—11 about as long as broad. Last segment about one and one-half times as long as broad at base. Scutum with distinct punctures, irregularly placed, sometimes in short rows but interspaces usually two or three times diameter of punctures, central part of scutum largely impunctate, scutellum and metanotum with a few fine punctures. Enclosed area of propodeum with less longitudinal carinae than usual (six long carinae and also a few incomplete carinae), median two carinae much diverging. Enclosed area not much depressed, posteriorly not distinctly bordered and gradually passing into the smooth areas behind the enclosed area. Back of propodeum with mostly fine, irregularly reticulate carination. Mesopleura with very fine remote punctures. Acetabular carina reaching about halfway the distance between the median line and the end of the epicnemial carina. Legs normal, back of hind femora smooth with median longitudinal line of very fine piliferous punctures. Second recurrent vein of fore wings interstitial. Petiole about seven times as long as broad in the middle, almost cylindrical, with upper lateral edge; sides anteriorly somewhat depressed. Gastral tergites with sparse and fine puncturation, margin of tergites 2—5 impunctate, sixth tergite with elongate triangular pygidial area (Fig. 16, holotype), which is very shining and has a few punctures along the margin. Gastral sternites with a few fine punctures, second sternite almost smooth. Appressed pubescence of face silvery, temples and mesosternum also with silvery pubescence. Rest of body with greyish pubescence, a few long stiff hairs along margin of tergites and sternites, sternite 6 with dense yellowish-grey pubescence, partly with long, backwards directed hairs. Length about 8.5 mm. Male. — Legs paler: femora more or less brownish, mid and hind tibiae much paler brown. Sides of petiole somewhat more depressed so that almost two carinae are formed. Antennal segments 9—12 slightly longer than broad, last segment about one and one- half times as long as its width at base. Second gastral sternite with smooth posterior margin, bordered by a few long stiff hairs. Margins of following sternites finely pubescent and with a few long stiff hairs, margins of third and fourth sternites each with two bundles of fasciculate hairs. Sixth sternite with a patch of dense short brownish pubescence and a great number of long, backwards directed stiff hairs. Genitalia Fig. 42. Length about 8.5 mm. Solomon Is.: 2 @ (holotype and paratype), Santa Ysabel, Tamatahi, 450 m, 2 July 1960, coll. C. W. O'Brien; 1 4 (allotype), Bougainville S., Buin, 2 June 1956, coll. J. L. Gressitt; 1 4 (paratype), Bougainville, Mumurai, 400 m, 7 June 1956, coll. J. L. Gressitt (all BISH). The sexes come from different islands; this may explain the difference in the colour of the legs. P. vadosus differs from its relatives in the enclosed area of the propodeum not being 116 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 distinctly separated from the smooth areas on the propodeum and in the long acetabular carina. Psen (Psen) bryani Perkins & Cheesman 1928, Perkins and Cheesman, Insects of Samoa 5 (1): 28—29 (Psen bryani). I have studied the allotype ( 4 ) in the collection of the British Museum (Natural History), but not the female. In the material received from the Bernice P. Bishop Museum I found three males which undoubtedly also belong to this species. Their tarsi are dark. Hind margins of gastral tergites 2—5 impunctate. Genital apparatus resembling that of P. elisabethae but of extraordinary large size (Fig. 38 and 39). The only characteristics I could find to distinguish this species from P. cheesmanae from the New Hebrides are the darker tarsi of the male and probably also its darker fasciculate hairs of the gastral sternites. New records: Samoa: 1 4, Tutuila, Afono trail, March 1930, coll. O. T. Fullaway; 2 &, Tutuila, Mt. Alava, 500 m, 20—24 Febr. 1965, Malaise trap, coll. G. A. Samuelson (BISH). Psen (Psen) cheesmanae Krombein 1949, Krombein, Proc. Hawaiian Ent. Soc. 13 (3): 364—365 (4). Krombein based his description on two males, collected by Miss L. E. Cheesman. In the collection of the British Muscum (Natural History) I found three females, also collected by Miss Cheesman, same locality and same period, which are probably the opposite sex of this species. The hind margins of the gastral tergites 2—5 are impunctate, as in the male holotype. The puncturation of the scutum of these females and of the holotype resembles to a great extent that of the male of P. bryani Perkins & Cheesman. Pygidial area Fig. 17. New Hebrides: 3 9, Santo, Aug.—Sept. 1929, coll. L. E. Cheesman, BM 1929— 537 (BM). Psen (Psen) bishopi spec. nov. Female. — Head and thorax black; middle part of mandibles reddish, palpi testa- ceous, underside of scape reddish-brown, apex of antennae below somewhat brownish, tegulae reddish-brown. All legs including trochanters reddish. Petiole, first and second gastral tergites except darkened apical margin, second sternite and apical margin of following sternites reddish. Rest of gaster black. Veins of wings dark brown. Clypeus raised below, densely punctate, anterior margin with light emargination, surface of this margin shining with very fine transverse striation (just visible under 30 x magnification). Interantennal carina ending in a low triangular tooth which is connected with the underside of the antennal sclerites by a fine carina. Only central part of frons with dense superficial puncturation, rest of frons and vertex shining, with few punctures. Postocellar area slightly raised. Mandibles and antennae normal. Scutum with fine puncturation on fore part, punctures stronger on posterior half, interspaces often a few times larger than diameter of punctures, on posterior half punc- tures partly in rows. Scutellum and metanotum smooth, with a few punctures. Enclosed area of propodeum but slightly concave, latero-posteriorly not distinctly separated from the smooth surface behind the enclosed area and with long oblique longitudinal carinae, J. P. van LiTH: Oriental species of the genus Psen 15107 median two carinae strongly diverging. Behind enclosed area on each side of median sulcus a large smooth area, partly on horizontal part of propodeum. Back of propodeum with irregular, not very strong reticulate carination. Mesopleura almost smooth, meta- pleura smooth and shining. No distinct acetabular carina. Both recurrent veins end in the second submarginal cell. Legs normal, back of hind femora smooth with a longitudinal line of minute piliferous punctures. Petiole cylindri- cal, dorsally somewhat flattened, about 7 times as long as wide in the middle. Gastral tergites and sternites with very few fine punctures, broad hind margin of tergites 2—5 impunctate. Pygidial area elongate-triangular, smooth with a few punctures along the margin (fig. 21). Face with silvery appressed pubescence and a few long erect hairs, dorsum and back of thorax with golden-brown long pubescence, pubescence of mesosternum paler. Gaster almost bare, a few long stiff hairs near margin of sternites. Length about 10 mm. Male unknown. Solomon Islands: 1 9 (holotype), Kolombangara, Collifer’s Camp, 700 m, 24 Jan. 1964, coll. P. Shanahan (BISH). Easily distinguished from other species of the group by the red petiole and first gastral tergites and by the almost impunctate gaster. Group of Psen orientalis This group is even larger than that of P. elisabethae, including the three species just described by Tsuneki from Formosa now counting 20 forms. All have the characteristic shape of the propodeum. The females of most of the species belonging here have a broad and opaque pygidial area, but in P. hakwsanus and a few allied species the pygidial area is narrower and more or less shining. The number of tyloidea on the antennae of the males varies, sometimes they are even absent. Psen (Psen) terrigenus Van Lith 1959, Van Lith, Zool. Verh. Leiden 39: 30—34 (@). 1965, Van Lith, Zool. Verh. Leiden 73 : 39. This species was described from three females from West Java, no male being known at that time. A male, collected in East Java was discovered recently in the collections of the British Museum (Natural History), London. The puncturation of this specimen is strong, as in the females. The genitalia resemble those of P. orientalis; this confirms my earlier supposition (1965) that P. terrigenus belongs to the group of that species. Fore and mid tarsi dark brown, hind tarsi paler (in the female tarsi reddish brown). Clypeus convex, median part of anterior margin with shallow triangular emargination, not distinctly separated from lateral parts. Frons with dense puncturation, punctures distinct and mostly in rows. Frontal line fine but distinct. Antennal segments 4—13 with broad oblong, longitudinal, shining tyloidea; tyloidea of segments 4—11 about as long as segment, broadest on first segments, on the 12th segment ending before apex, on the 13th segment very small. Propodeum generally similar to that of female but carination on back coarser and 118 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 | vadosus cheesmanae novahibernicus bishopi baduriensis paulus (Hollandia) Fig. 15—21. Pygidial area of female of species of Indo-Australian and South Pacific Psen s.str. 15, P. novahibernicus Van Lith; 16, P. vadosus sp.n.; 17, P. cheesmanae Krombein; 18, P. paulus sp.n. (N.E. New Guinea); 19, P. paulus sp.n. (Hollandia); 20, P. paulus baduriensis subsp.n.; 21, P. bishopi sp.n. J. P. van LitH: Oriental species of the genus Psen 119 extending higher, leaving a narrow, almost smooth area laterally behind the enclosed area. Acetabular carina (with a few parallel carinae) not yet reaching halfway the epicnemial carinae. Face with silvery pubescence. Hind margin of third and fourth gastral sternites with fringe of fasciculate yellowish-golden hairs. Length about 8.5 mm. East Java: 1 4, May 1938. Tengger Highlands, Nongkodjadjar, 1100 m, coll. Mrs. M. E. Walsh (BM). Psen (Psen) hakusanus seminitidus Van Lith 1934, Gussakovskij, Ark. Zool. 27A (21): 7 (Mimesa kobli). 1965, Van Lith, Zool. Verh. Leiden 73 : 40—41 (Psen (Psen) seminitidus). 1966, Tsuneki, Etizenia Fukui Univ. 14: 9—10 (Psen (Psen) hakusanus seminitidus) 8. 1967, Tsuneki, Etizenia Fukui Univ. 24: 2, 9. In 1966 Tsuneki studied a male from Formosa which he provisionally identified as P. hakusanus seminitidus. He noticed some differences from the females reported from China in 1965 (Van Lith). According to Tsuneki the puncturation is as close as in P. hakusanus s.str. and the petiole has no ventral median carina. Tsuneki therefore suggested that it is a different geographical race. In his recent paper Tsuneki (November 1967) also recorded a female from Formosa and Sauter collected there two females which are very similar to P. hakusanus seminitidus. They differ, however, in the slightly narrower pygidial area as compared with the figure in my paper of 1965. The puncturation is less dense than in P. hakusanus s.str. The ventral median keel of the petiole is distinct, as in the specimens from the continent. Length about 9 mm. A further study based on more extensive material would be very welcome. New records from Formosa: 2 9, Hoozan, Nov. 1910, coll. H. Sauter (ZMB). Psen (Psen) kulingensis Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 41—42 (S. China). Two females from Japan are very probably conspecific with P. kulingensis described from S. China. The petiole is slightly longer, the legs and wing veins are somewhat paler than in the paratype available for comparison. The pygidial area is less broad. Perhaps these specimens form a new subspecies. They differ from P. affinis Gussa- kovskij, from Japan, in the absence of a ventral keel of the petiole and in the somewhat coarser puncturation of the scutum. Japan: 2 9, Chuzenji, July—Aug. 1913, 4000—5000 ft. coll. F. Muir (BISH). Psen (Psen) simlensis spec. nov. Female. — Black; median part of mandibles reddish-brown, antennae somewhat brownish. Palpi and tarsi reddish-brown. Spurs of tibiae yellowish-brown. Ventral side of gaster partly dark reddish-brown. Veins of wings dark brown. Clypeus convex, shining, with fine punctures, median part of anterior margin pro- truding and somewhat raised, margin smooth, slightly emarginate (Fig. 23, holotype). Frontal carina fine, ending between antennae in a distinct tooth, which is connected 120 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 with antennal sclerites by very fine, bent carinae. Frons above antennae with fine punctures, upper part of frons and vertex shining, with sparse and minute punctures. Frons distinctly raised in front of each of the posterior ocelli, also postocellar area somewhat raised. Antennae normal, segments 9—11 about as long as they are broad, twelfth segment slightly more than one and one-half times as long as its width at base. Scutum shining with fine punctures, interspaces mostly a few times as large as dia- meter of punctures, sometimes only just as large. Scutellum and metanotum shining, with scarce puncturation. Enclosed area of propodeum with oblique carinae, median two carinae almost parallel, surface between latter two carinae somewhat irregular. A smooth area, broadest on the sides, behind the enclosed area, on dorsal side of propodeum, narrowing towards the median sulcus. Back of propodeum with irregular reticulate carination. Median sulcus with a few transverse carinae. Mesopleura and mesosternum smooth and shining, with minute punctures. No acetabular carina. Mid tibiae with a short longitudinal row of three or four long thorns at apex, hind tibiae dorsally with a longitudinal row of short, slightly bent, pale thorns. Second recurrent vein of fore wings interstitial. Petiole almost cylindrical, dorsally somewhat flattened. Tergites and sternites, except narrow hind margins, with sparse and fine punctures. Hind margin of fifth gastral sternite emarginate, sixth sternite densely punctate, with a narrow, smooth and shining median keel. Pygidial area elongate-triangular (Fig. 22), surface with very fine coriaceous sculpture, a few fine punctures along the lateral carinae. Pubescence of face silvery, appressed, with a few long erect hairs. Pubescence of rest of body greyish, long on propodeum, sparse on gaster. Sternites 2—5 with a few long and stiff hairs before hind margin. Sixth sternite with dense and short pubescence. Length about 8.5 mm. Male unknown. North India: 1 @ (holotype), Simla (Himachal Pradesh or Simla, Calcutta), 12 Aug. 1918, Bruneti coll. 1927—184 (BM); 1 ® (paratype), Mussoorie, 7500 ft., 24 June 1962, No. 22, coll. V. Gupta (UDE). There is no doubt that Simla in the state of Himachal Pradesh is concerned. P. simlensis resembles the species of the subgroup of P. hakusanus, which have also a narrow and more or less shining pygidial area, but it is easily distinguished by the almost cylindrical petiole. Psen (Psen) orientalis Cameron 1890, Cameron, Mem. Proc. Manch. Lit. Phil. Soc. (4) 3 : 269, ® (Psen orientalis; Madras, India). 1902, Cameron, JI. Bombay Nat. Hist. Soc. 14: 289, 2 and & (Psen reticulatus; Deesa (!), India). 1965, Van Lith, Zool. Verh. Leiden 73 : 42—44. After the study of a fine series of fresh specimens, all females, from Simla (type- locality of P. reticulatus Cameron), a supplementary description can now be given. Female. — Black, also legs and veins of wings. Only the tips of the mandibles, the tibial spurs and the claws of the tarsi are dark reddish. Palpi dark brown. Hind tibiae dorsally with short reddish spines. Mid tibiae and basitarsi, also apex of hind tibiae and hind basitarsi with longer, pale spines. Apex of pygidial area not always with reddish spot. Clypeus slightly raised before the impunctate anterior margin. Frontal carina ending in a small tooth and an angular carina, which is not connected on either side with the antennal sclerites. Postocellar area slightly raised. Third antennal segment almost twice J. P. van LITH : Oriental species of the genus Psen 121 as long as fourth segment. Dorsal part of propodeum bordered by a fine high carina. Below this carina a narrow area without carinae but with piliferous punctures. Petiole laterally with distinct upper and lower edges and on posterior half of ventral side with unsharp carina which is more distinct on ventral plate of petiole. Scutum with very fine and short brownish pubescence and long whitish hairs. Meso- sternum with short white pubescene and long white hairs. New records: N. India, Simla Hills: Narkanda, 9000 ft., 6 9, 3 Oct. 1962, coll. Gupta, No. 27; 3 9, 5 Oct. 1962, coll. Gupta, No. 30 and 31; Chini bungalow, 8600 ft, 2 9, 11 and 13 Oct. 1962, coll. Gupta, No. 40 (UDE). As I explained in 1965 in my opinion P. orientalis and reticulatus are conspecific but a series of fresh material, males as well as females, from South India would be welcome to confirm this. Group of Psen refractus Psen (Psen) refractus Nurse 1903, Nurse, JI. Bombay Nat. Hist. Soc. 15: 11—12, ® (Mt. Abu, N. India). 1965, Van Lith, Zool. Verh. Leiden 73 : 50—51. Four females from Nilgiri Hills (S. India) are slightly darker than the type from N. India. The first and second gastral tergites are red, but the posterior margin of the second tergite is darkened medially. Base of third tergite red laterally. Apex of petiole below and whole ventral plate of petiole red, also second sternite. Clypeus completely black. Femora and greater part of tibiae black. A male from the same locality is a little more red than the females: second tergite completely red, also basal half of third sternite. Last antennal segment brown below. Structure of antennae as in P. refractus meridianus (Van Lith, 1965). As the differences between the females from Mt. Abu and from Nilgiri Hills are so unimportant, I do not think that the specimens from Nilgiri Hills belong to a dif- ferent subspecies. P. refractus meridianus from Kodaikanal, Palni Hills (two females and four males, Van Lith, 1965) is much darker and this may indeed be a distinct subspecies. The Palni Hills are not so far distant from the Nilgiri Hills, but the wasps were collected there at the altitude of 6500 feet whereas those from the Nilgiri Hills were found at 3200—3400 feet. Moreover these mountainous regions seem to be well separated geographically by plains. New records: Nilgiri Hills, South India, 1 9, Singara, 3400 ft, May 1954, coll. P. S. Nathan (coll. Ferguson), 3 9 and 1 4, Devala, 3200 ft., Oct. 1960, coll. P. S. Nathan (ML). Psen (Psen) krombeini Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 52—55. New records: 11 9, S. India, Madras State, Anamalai Hills, Kadamparai, 3500 ft. May 1963, coll. P. S. Nathan (ML). Group of Psen aureohirtus Psen (Psen) aureohirtus aureohirtus Rohwer 1921, Rohwer, Phil. JL Sc. 18 : 322—323 (Psen (Mimesa) aureohirta). 1959, Van Lith, Zool. Verh. Leiden 39 : 49—50 (Psen (Psen) aureohirtus). 1965, Van Lith, Zool. Verh. Leiden 73: 56. New record: Philippine Is., 1 4, Baguio (Luzon), April 1917, coll. F. X. Williams (BISH). 122 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 Psen (Psen) toxopeusi Van Lith 1959, Van Lith, Zool. Verh. Leiden 39: 47—49 (9). The original description was based on three females collected in 1936. In the material from the Berlin Museum I found two females, also collected in S. W. Celebes, by G. Heinrich, in 1931. In these specimens the acetabular carina is not distinctly present. The third gastral sternite is partly darkened. A male, taken at the same locality, is very similar. Antennae long, sixth segment with narrow longitudinal carina, seventh segment with broader tyloidea, segments 8—12 with almost circular depression, last segment with small round tubercle. Hind margin of third and fourth gastral sternites with long, brown and fasciculate hairs. Clypeus with appressed silvery pubescence, supra-clypeal area with appressed, pale golden pubescence, frons with erect golden pubescence. Genitalia resembling those of P. melanosoma (Fig. 44). Length about 11 mm. New records from S. W. Celebes: 2 @ and 1 ¢, Bonthain, Wawa Karaeng, 1100 m, Sept.-Oct. 1931, coll. G. Heinrich (ZMB). Group of Psen ruficrus Psen (Psen) ruficrus Van Lith 1965, Van Lith, Zool. Verh. Leiden 73 : 62—63 (€). A female collected in the close neighbourhood of the locality where two males were found six years earlier, is undoubtedly conspecific. It has no distinct bluish shine on head and thorax and the orange-red colour is slightly more extended: legs including a great part of coxae, petiole with ventral plate, lower margin of first tergite. Mandibles except dark tips and labrum orange-red. Upper side of scape of antennae red, most of the following segments with dark red spot on ventral side. Veins of wings dark brown. Lower part of clypeus protruding and with sharp lateral carinae (resembling the clypeus of the females of the group of P. refractus), anterior margin with triangular emargination (Fig. 7), surface of disk densely punctate. The fine interantennal carina ends in a low smooth triangle which is connected with the antennal sclerites by fine carinae. Frons distinctly punctate between antennae and ocelli, smooth laterally. Vertex with a few fine punctures, interocellar area stronger punctate and somewhat raised. Antennae slightly thickened towards apex. Mandibles normal. Scutum, scutellum and metanotum with fine remote punctures. Propodeal enclosure with distinct longitudinal carinae, median two carinae strongly diverging. Upper part of back of propodeum rounded laterally, vertical part flat, completely and coarsely cari- nate; sulcus narrow. Hind tibiae dorsally with longitudinal row of reddish thorns, at base with a short second row, apex with four longer thorns. Second recurrent vein of fore wings interstitial (in right wing ending just in second submarginal cell, owing to deformation). Petiole cylindrical except short anterolateral carinae. Gaster almost com- pletely smooth with very few minute punctures, margins completely smooth. Pygidial area large, broadly triangular, base convex, surface coriaceous with 7—8 irregular rows of large punctures, narrow margin impunctate (Fig. 5). Face with golden pubescence, cheeks more silvery pubescent, rest of body with long J. P. vAN LITH: Oriental species of the genus Psen 123 26 exaratus santoro Daf 28 indicus == RK 29 CAPTER LRQ Sean NN ES indicus 32 taiwanus inflatus inflatus taiwanus Fig. 22—23. Pygidial area and anterior margin of clypeus of Psen (Psen) simlensis sp.n. (9) Fig. 24—25, face and pygidial area of Psen (Mimesa?) inflatus sp.n. (2). Fig. 26—33, sub- species of Psen (Punctipsen) exaratus (Eversmann), all males. 26, mid tarsus of P. exaratus s.str.; 27, mid tarsus of P. exaratus santoro Yasumatsu; 28—29, mid basitarsus and mandible of P. exaratus indicus subsp.n.; 30—31, dorsal and ventral view of genitalia of P. exaratus indicus subsp.n.; 32—33, mid basitarsus and fore basitarsus of P. exaratus taiwanus Tsuneki. 124 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 brownish hairs, scutum darker. Gaster very sparsely haired. Pygidial area with back- wards directed hairs. Length about 11 mm. New Guinea: 1 2, N.E. New Guinea, Daulo Pass, 2400 m, 15 May 1963, coll. J. Sedlacek (BISH). I am more convinced now that P. ruficrus is closely related to P. melanosoma of the group of P. aureohirtus. The pygidial area of the female, the golden clypeal pubescence and the extent of red colour also point to a relationship with P. toxopeusi, of the same group, although the vertex is much less raised. Contrary to P. toxopeust the median part of the clypeus is raised in the female of P. ruficrus and carinate laterally. Therefore I hesitate to place it definitely in the group of P. aureohirtus and prefer to consider it provisionally as an intermediate form. The species also occurs in Papua, where Miss L. E. Cheesman collected a fine series of males: 9 3, Mondo, 5000 ft., Febr.-March 1934, BM 1934-321 (BM). Group of Psen bakeri Psen (Psen) sedlaceki spec. nov. Male. — Head and thorax black; reddish are: mandibles except dark red tips, legs including trochanters and hind coxae. Claws of legs, margins of pronotal tubercles, tegulae and veins of wings brown. Scape of antennae brown, palpi testaceous. Petiole, first gastral tergite, second gastral tergite except brown margin, sides of third tergite and entire second gastral sternite pale reddish. Rest of gaster black. Clypeus dull, densely and finely punctate, anterior margin with transverse striation and triangular emargination. The interantennal carina ends in a small flat tubercle. Frons and interocellar area with fine but distinct puncturation, vertex smooth and shining, postocellar area not raised. Antennae rather long, segments 7—12 with large broad-oval tyloidea, segments 6 and 13 with a small shining tubercle. Last segment about twice as long as broad at base. Greater part of scutum with very coarse punctures, on posterior half partly in striae. On the sides of the scutum the punctures are much finer. Scutellum with a few distinct punctures, metanotum much finer and more densely punctate. Enclosed area of propo- deum with strong posterior carina, two median of the longitudinal carinae strongly diverging. Back of propodeum with coarse reticulate carination: sides with a few oblique carinae. Mesopleura and hypo-epimeral area with fine punctures. A short but distinct acetabular carina. Interepicnemial area with fine transverse striation. Legs normal. Second recurrent vein of fore wings interstitial. Petiole long, cylindrical. Gaster with minute punctures. Pubescence of face silvery, appressed and with a few long erect hairs. Head and thorax greyish pubescent. Mesosternum with dense silvery pubescence. Gaster with very fine, short and sparse pubescence, a few long hairs near the hind margin of the last tergites and sternites. Hind margin of third and fourth sternites each with two bundles of golden-brown long fasciculate hairs. Sixth sternite with dense, short brownish pubescence. Genitalia Fig. 43. Length about 10.5 mm. Female unknown. J. P. VAN LITH: Oriental species of the genus Psen 125 N. E. New Guinea: 1 ¢ (holotype), Karimui, 1080 m, 14 July 1963, coll. M. Sedlacek (BISH). This species seems to be closely related to the Philippine P. bakeri Rohwer, which has an entirely black gaster and petiole. They share the following characters: coarse puncturation and striation of scutum, reduced interantennal tooth and triangular emargin- ation of clypeus. The female of P. bakeri has a golden pubescent face. Punctipsen subgen. nov. De Beaumont (1937) placed P. exaratus (Eversmann) in the subgenus Psen but he admitted that its systematical position was somewhat doubtful. Indeed the lack of fasciculate hairs on the margins of the third and fourth gastral sternites of the male distinguish P. exaratus from any other species belonging to the subgenus Psen. In both sexes the longitudinal groove and the strong punctures, which are sometimes fused into short grooves, of the petiole are properties lacking in Psen s.str. Only in oriental repre- sentatives of the subgenus Mzmwmesa Malloch the posterior end of the petiole has a few large punctures. Moreover, the genital apparatus of the male differs considerably from the genitalia studied thus far in the subgenus Psen s.str. Therefore I place P. exaratus in the present new subgenus. The nominate subspecies occurs or has occurred probably in the whole palaearctic area. Four subspecies have been found in the Indo-Australian region; one of these is described here as new from South India which is the first record of P. exaratus from South Asia. As the oriental representatives seem to be allopatric and the differences are not important, I prefer to maintain their subspecific status as proposed by Tsuneki (1966). The differences between /arwanus Tsuneki and the four other subspecies are greater, however, than the differences between the latter subspecies (mid basitarsus ot male, pubescence of gaster). Punctipsen is very closely related to Psen s.str., the epicnemial carinae being contin- uous with the anterior precoxal suture (carina). The hypo-epimeral area is distinct, the upper longitudinal half of the back of the hind femora is smooth and separated from the lower half by a broad line of minute piliferous punctures. The frontal carina 1s complete. The basitarsi of fore and mid legs of the male and mostly also its mandibles have special structures but these characters are not of subgeneric value. The stipites of the genitalia are not nearly flat as in the subgenus Psen but swollen, also their tips, and there are no membranous lobes. Type species of Punctipsen: Mimesa exarata Eversmann. KEY TO THE SUBSPECIES OF Psen (Punctipsen) exaratus (The females of a few subspecies being unknown, this key is only reliable for the identification of the males) IAD = interantennal distance; AOD = antenno-ocular distance. 1. Mandibles of male ventrally with a long tooth. Basitarsus of mid legs of male with short median tooth, apical tooth long. In both sexes ratio eye-temple 1.0—1.2; ratio IAD-AOD 1.8—2. Punctures on gastral tergites 1 and 2 sparse and weak. In female apical segments of antennae ferruginous beneath. (W. Europe, Caucasus, Siberia, PROTEIN) pati) wiser CEE exaralus exaratus 126 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 — Mandibles of male ventrally without tooth or with vestigial tooth. Ratios different 2. Mandibles of male without ventral tooth. Median tooth of mid basitarsus of male about half as long as apical tooth. In both sexes ratio eye-temple about 1.6; ratio (IAD-AOD about 3.2—4. In female antennae completely black. (Amami-Oshima VE EE TORRES ART Ne oan ASA Re dodd elec exaralus Santoro ==" Mandibles of male with vestigial ‘tooth 0 eren CESSE EEE 3 3. Median and apical teeth of mid basitarsus about equally long. Ratio eye-temple about 1.25; ratio IAD-AOD about 2.15. First gastral tergite with very fine punc- tures. Underside of antennae yellowish-brown. Pubescence of gaster greyish-white, not long and erect on segments 3—5. Female unknown. (S. India). .................. exaratus indicus subsp. n. — Median tooth of mid basitarsus of male at most half as long as apical tooth. Ratio eye-temple ‘about 1:3; TAD-AOD 23 Ze ne 4 4. Median tooth of mid basitarsus about half as long as apical tooth, situated about halfway basitarsus. Underside of antennae dark ferruginous. Pubescence of gaster not very dense or erect. Female unknown. (Okinawa) exaratus intermedius — Median tooth of mid basitarsus of male inconspicuous, situated about two-thirds of length of basitarsus from base. Punctures on gastral tergites 1 and 2 more abun- dant and more distinct. In female underside of antennae partly ferruginous, in one of the males light brown. Pubescence yellowish-grey, dense, long and erect, especially on gastral tergites 3—5. (Formosa) .................. exaralus taiwanus Psen (Punctipsen) exaratus taiwanus Tsuneki 1966, Tsuneki, Etizenia Fukui Univ. 14: 6 (9). 1967, Tsuneki, Etizenia Fukui Univ. 24: 2 (2). As Tsuneki was able to study only a single female from Formosa, he doubted the validity of the new form. Through the kindness of Dr. Königsmann, Humboldt- University in Berlin, I received several females and males collected by H. Sauter in Formosa more than half a century ago. Now I am completely satisfied that this form from Formosa is indeed a distinct subspecies. The male has a very short (vestigial) tooth on the ventral side of the mandibles but it is easily distinguished from the male ot P. exaratus intermedius Tsuneki from the Okinawa Islands by the very indistinct tooth of the mid basitarsus and the smaller distance between this tooth and the apex. Male. — Fore and mid legs almost completely brown (in female more darkened); underside of basal part of antennae more yellowish (in female brown below). Anterior margin of clypeus with a simple triangular emargination. Mandibles ven- trally with a very small tooth (cf. Fig. 29). Scape of antennae thick; antennal segments 6—13 with distinct flat tyloidea, most distinct on segments 8—11. Frons and vertex slightly raised, with distinct punctures. Scutum with strong punctures, interspaces often smaller than diameter of punctures. Rugae on upper part of back of propodeum coarse, more or less parallel. Both recurrent veins of fore wings ending in second submarginal cell. Basitarsus of mid legs with long tooth at apex; an indistinct tooth at about two- thirds of length of basitarsus from base (Fig. 32). Basitarsus of fore legs Fig. 33. Petiole dorsally with deep punctures in coarse rugae, sides and underside each with two J. P. van LITH: Oriental species of the genus Psen 127 distinct longitudinal carinae. Apex of petiole latero-ventrally also with a number of short, irregular carinae. Dense, yellowish-grey pubescence; apical margin of gastral tergites 2—5 with a fringe of long pale hairs with bent tips. Length about 9—10 mm. The female was amply described by Tsuneki. I should like to add that the second gastral sternite has a slight but broad median longitudinal depression. Base and hind margin of tergites 3—5 are densely punctate. The petiole has indeed only two lateral carinae; the ventral side is rounded (it has two carinae in the male) with coarse punctures and small rugae on apical half. New records from Formosa: 3 9, Taihorin, May and Oct. 1910; 2 &, Taihorinsho, Aug. 1909, and Taihorin, Oct. 1910, coll. H. Sauter (ZMB). Psen (Punctipsen) exaratus indicus subspec. nov. Male. — Black, fore side of scape brownish, underside of antennae yellowish- brown. Mandibles yellowish-red with darker tips. Fore and mid femora reddish except dorsal side of basal half. Fore and mid tibiae reddish, slightly darkened on outer side. Mandibles with vestigial tooth (Fig. 29) on ventral side, like that of P. exaratus inter- medius Tsuneki from Okinawa. Frons and vertex slightly raised. Puncturation of frons, vertex and scutum distinct but not very strong. Propodeum on upper part of back with a number of rugae, not very strong and somewhat irregular. Antennal segments 6—13 with indistinct broad tyloidea. Inner tooth of mid basitarsus much longer than in the males of the other four forms, nearly as long as apical tooth; an imaginative line connecting the apices of the teeth of the mid basitarsus is about parallel with the axis of the basitarsus (Fig. 28) whilst in the other subspecies these lines are distinctly diverging (Fig. 26—27). Both recurrent veins of fore wings ending in second submarginal cell. Petiole not rugose, with deep and large punctures, sides and ventral side each with two carinae. Apex of ventral side of petiole not with additional irregular rugae. Median part of second gastral sternite somewhat depressed longitudinally. Pubescence greyish, face silvery. Genitalia Fig. 30—31. Length about 8—9 mm. Female unknown. S. India: 2 & (holotype and paratype), Madras State, Anamalai Hills, Kadamparai, 3500 ft., May 1963, coll. P. Susai Nathan (ML). The present male is easily distinguished from that of the nominate subspecies by the vestigial tooth of the mandibles and from the other forms of P. exaratus by the long median tooth of the mid-basitarsus. Subgenus Mimesa Shuckard The subgenus Mimesa has not yet been recorded from the Indo-Australian region. Two females collected in New Guinea during the Third Archbold Expedition in 1939 share a few important characters with palaearctic Mimesa (sensu De Beaumont, 1937) and they are either the first representatives of this subgenus in the Indo-Australian area or they belong to a new subgenus. The characters referred to are: the distinctly 128 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 paulus 37 43 vadosus toxopeusi sedlaceki Fig. 34—44. Male genitalia of species of Indo-Australian, South Pacific and East Asiatic Psen s.str. 34—35, dorsal and ventral view of P. ater (Fabricius); 36—37, ventral and dorsal view of P. paulus sp.n. (left part); 38—39, ventral and dorsal view of P. bryani Perkins and Cheesman; 40—41, ventral and dorsal view of P. paulus subtilis subsp.n.; 42, dorsal view of P. vadosus sp.n. (right part); 43, dorsal view of P. sedlaceki sp.n.; 44, dorsal view of P. toxopeusi Van Lith (left part) J. P. VAN LitH: Oriental species of the genus Psen 129 bordered epicnemial areas, the course of the epicnemial carinae, the incomplete frontal carina, and the lack of carinae connecting the antennal sclerites with the interantennal tooth. The pygidial area is shaped as in oriental Mimumesa, being shining with a few punc- tures along the margin. The hypo-epimeral area is not carinate or punctate as in palaearc- tic Mimesa, but it is extended laterally into a right cone with the entire hypo-epimeral area as its base. The lower half of the mesopleura and the whole mesosternum are coar- sely and transversely striate. The petiole is also different from that of palaearctic Mimesa, being long and cylindrical as in oriental Psen and Mimumesa. The longitudinal carinae of the enclosed area of the propodeum continue somewhat on the back of the propodeum, as in Mimumesa auratus and tridentatus. Although in some details the two females also resemble the oriental species of Mimu- mesa, | prefer to relegate them provisionally in the subgenus Mimesa, because of the structure of the epicnemial areas and the lack of an acetabular carina. Psen (?Mimesa) inflatus spec. nov. Female. — Head and thorax black; head with somewhat metallic-blue shine, prono- tum dorsally and notably pronotal tubercles with violet-bronze reflections, also hypo- epimeral cones somewhat bronze-coloured; propodeum with paler bronze-coloured re- flections tending to brassy. Mandibles except their dark base yellowish, with dark red tips. Labrum yellowish. Palpi reddish. Scape of antennae reddish, also underside of antennae, upper side of antennae brownish. Ends of femora and the entire tibiae and tarsi reddish, hind tibiae and tarsi somewhat darkened. Spurs of tibiae yellowish-white. Tegulae reddish. Veins of wings dark brown. Petiole including ventral plate and sides of first gastral tergite dark reddish, margins of gastral tergites and of sternites 3—5 and the entire last segment reddish. Clypeus (Fig. 24, holotype) densely punctate, median part much protruding with slight angular emargination and fine transversely striate surface. Frontal carina only distinct on lower half of frons, ending below in a tooth which in frontal view has an elliptic depression on its lower part. No carinae between this tooth and the antennal sclerites. Frons densely punctate, punctures in shallow rugae. Vertex much shining with a few distinct punctures, postocellar area not raised. Temples and cheeks almost im- punctate. Antennal segments long, third segment more than four times as long as wide at apex, segments 10 and 11 at least one and one-half times as long as they are broad and last segment slightly more than twice as long as broad at base. Scutum with strong puncturation, punctures in rows in shallow rugae. Scutellum anteriorly raised with strong punctures and a few irregular transverse carinae on its fore part; posterior part with a number of parallel longitudinal carinae. Metanotum with indistinct punctures and traces of longitudinal carinae. Enclosed area only slightly depressed, without any distinct posterior border or carina. Longitudinal carinae ot enclosed area continuing on to back of propodeum, a few continuing even further until they are lost in the irregular reticulate carination on lower end of propodeum. Median two longitudinal carinae of enclosed area slightly diverging and then narrowing into the median sulcus on back of propodeum. Sides of propodeum with irregular sculpture and a few oblique carinae. Epicnemial areas shining with minute punctures, distinctly separ- ated from interepicnemial area, which is lower. Outer epicnemial carinae not bent back- 130 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 wards. The subpleural signum forms a pointed tubercle. Interepicnemial area and mesosternum with coarse transverse, almost parallel rugae of which a few continue on to the lower part of the mesopleura. Upper half of mesopleura almost smooth. Hypo- epimeral area distinctly bordered, swollen and extending laterally into a high cone, which in dorso-lateral view is as high as the length of the tegulae. Subalar area (under tegula) also somewhat swollen, its ventral side bordered by a sharp carina. Legs normal, back of hind femora smooth and shining, hind tibiae with longitudinal row of short spines. First recurrent vein of fore wings ending near middle of second submarginal cell, second recurrent vein interstitial. Petiole cylindrical, reaching beyond hind femora when stretched. Gastral tergites and sternites with a few fine punctures, hind margin impunctate. Pygidial area (Fig. 25, holotype) elongate triangular, at base carinae bent somewhat inwards, surface smooth and shining, apical part with some fine, transversely striate sculpture, along the lateral margins a few fine punctures. Face below antennae with golden pubescence, appressed and also with a few long erect hairs. Lower part of frons with golden pubescence, head and thorax dorsally with brownish hairs, ventral side and back of thorax with long whitish hairs. Gaster with very few hairs, in front of hind margins of sternites and of tergites 4—5 a few long and stiff hairs. Length about 9.5 mm. Male unknown. Central New Guinea: 1 9 (holotype), Mist Camp, 1800 m, 10 Jan. 1939, 1 9 (paratype), Top Camp, 2100 m, 9 Febr. 1939, Neth.-Ind—American New Guinea Expedition, coll. L. J. Toxopeus (holotype ML, paratype USNM). Some particulars regarding the localities where this highly interesting form was collected have been published by Toxopeus (1940, 1941). The camps were on the northern side of the Nassau Range near the Sahuweri, an affluent of the Idenburg River. The climate was not favourable. At Top Camp the sun did not shine more than a few hours in the morning; it was misty in the afternoon, stormy during the night. Mist camp was even worse, the sun shining during an hour or less in the morn- ing, the rest of the day being foggy. J. P. vAN LITH: Oriental species of the genus Psen oil LIST OF THE EAST ASIATIC, INDO-AUSTRALIAN AND SOUTH PACIFIC SPECIES OF THE GENUS Psen LATREILLE WITH THEIR RANGE OF DISTRIBUTION Subgenus Psen Latreille Group of ater P. ater (Fabricius), 2 4 — Europe, Siberia, China, Mongolia, Korea, Japan P. aurifrons Tsuneki, 9 4 — Japan P. yasumatsui Gussakovskij, 4 — Japan Group of aspites P. aspites sp. n., 3 — Celebes Group of emarginatus P. emarginatus Van Lith 9 & — Java, Borneo P. dzimm Tsuneki 2 ¢ — Japan P. pilosus Van Lith, & — Malaya P. nepalensis sp. n., 9 — Nepal P. sauteri sp. n., Q — Formosa Group of curvipilosus P. curvipilosus Van Lith, 9 4 — Java P. lieftincki lieftincki Van Lith, Q & — Sumatra, Malaya, ? Formosa P. lieftincki minor Van Lith, 9 4 — Malaya Group of tsunekii P. tsunekit Van Lith, 9 & — Japan P. ussuriensis Van Lith, 4 — Siberia P. shirozui Tsuneki, & — Formosa P. vechti vechti Van Lith, 9 — Java P. vechti birmanicus Van Lith, 9 — Burma P. assamensis Van Lith, 4 — India Group of nitidus P. mitidus nitidus Van Lith, 9 4 — Java, Krakatau, Bangka, Sumatra, South India P. nitidus takasago Tsuneki, 9 & — Formosa Group of elisabethae P. elisabethae elisabethae Van Lith, 9 & — Java, Sumatra P. elisabethae auricomus Van Lith, 9 4 — Malaya, Pulau Tioman P. elisabethae madrasiensis subsp. n., 2 & RS india P. amboinensis Van Lith, 9 — Ambon P. heinrichi sp. n., & — Celebes P. regalis sp. n., 9 — Solomon Islands P. coriacens Nan Lith, 98% — Philippines P. marjoriae sp. n., 9 — Philippines P. novahibernicus Van Lith, 9 & — New Ireland P. paulus paulus sp. n., 9 & — New Guinea, Papua „m ISS) N VVD ve u Ewe RS ov nio ed os) Lone Vv Dv DCS DID TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 111, AFL. 4, 1968 paulus baduriensis subsp. n., 9 paulus subtilis subsp. n., 4 vadosus Sp. n., 2 4 bryani Perkins & Cheesman, 9 4 cheesmanae Krombein, 9 @ bishopi sp. n., 2 New Guinea: Japen Island Solomon Islands Solomon Islands Samoa New Hebrides Solomon Islands Group of rufiventris rufiventris Cameron, 9 rubicundus rubicundus Van Lith, 9 rubicundus lawuensis Van Lith, 9 angulifrons Van Lith, & nigriventris Van Lith, 9 richardsi Tsuneki, 9 & S. India W. Java E. Java Mindanao Luzon Japan Group of orientalis terrigenus Van Lith, 9 & hakusanus hakusanus Tsuneki, 9 & hakusanus seminitidus Van Lith, 9 & koreanus koreanus Tsuneki, 9 koreanus formosensis Tsuneki, 9 hirashimai Tsuneki, & kulingensis Van Lith, 9 simlensis sp. n., 9 orientalis Cameron, 9 & affinis affinis Gussakovskij, 9 4 affinis grahami Van Lith, 9 & yomasanus Van Lith, 9 & tanoi Tsuneki, 9 & fuscinervis (Cameron), 9 & politiventris politiventris Rohwer, 9 & politiventris bellus Van Lith, 9 politiventris pahangensis Van Lith, 9 dlishanus Tsuneki, 9 & longicornis Tsuneki, 4 triangulatus Van Lith, & Group of refractus refractus refractus Nurse, 9 refractus meridianus Van Lith, @ & krombeini Van Lith, 9 & matalensis Turner, 9 @ Java Japan China, Tibet, Formosa Korea Formosa Ryukyu Islands S. China, ? Japan N. India India Siberia, Korea, Japan China Burma Formosa India Luzon Mindanao Malaya Formosa Formosa Java, Sumatra N. India, S. India S. India S. India Ceylon Group of aureohirtus aureohirtus aureohirtus Rohwer, 9 & aureohirtus rufopetiolatus Van Lith, 3 toxopeusi Van Lith, & melanosoma Rohwer, & carbonarius (Smith), ¢ Luzon Negros Celebes Philippines Morotai © VV © © © D a Ma AS) os) VV DP gs) Bs) Gsi Se) SS) GS BS) oS D