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LABISKY Biological Notes No. 65 ILLINOIS NATURAL HISTORY SURVEY URBANA, ILLINOIS MAY, 1969 STATE OF ILLINOIS DEPARTMENT OF REGISTRATION AND EDUCATION NATURAL HISTORY SURVEY DIVISION *pajyuny Aor OIOYA SOUNOD oY} TOF ULY} TOYA OOUOp{sor JO SoRUNOD ITOq, TOF SYOOO Jo [[P{ [VuUOsvoS YOU} powodos Apoy!] AIOA sroyUNY OULOS OsNvoEq BONNGEySIpP jyuvsvoyd oprioejzeys Juosordorstur ATYaTS rONVHLSS37 [| NOINN NOSNOWE SN1d I'S 8vé6l Y3SLNNH ¥3d 0371 SHOOO NUNYE ( |_wosuawat} NOIMWA < FauOsMvuD ANJNODLNON HONONOG?W 43083N Ajqeqoid sdvur osoyy, *(6'8S6T wospoeqoy jaye) uosees Zuyuny gpEt 94} 49}ze siszuNYy 0} peyNqEysip sozreuuoNsonb uosvos Buyunyysod wo (yYysu) pure (peysyqndun ‘r1yow ‘Oo [ep se) uosvos Sununy juvsvoyd peET Oya 40J soreuuonsenb asusory Suyuny wWosz (7491) poure}qo vyep wosy poddvur sv stour[y ul syuvseoyd yo 9ouvpunqe puv uoRyNnquysid—T ‘BIA G314Od3y LON YN rO NVHL SS371 ae NIDANWES SN1d IZ § 2¢6) Y3LNNH Y¥3d G37 SAHN09 NOSH33430 INAV ZINIYMY 1) 0! QYOJMVHD WWHONIS43 Trends in Pheasant Abundance in Illinois: 1958 to 1968 The pheasant (Phasianus colchicus), first introduced into Illinois about 75 years ago (Robertson 1958:3), has established thriving populations in northern and east-central Illinois. It has never established perma- nent populations in the west-central and southern counties of the state. Although the pheasant has ex- hibited shifting patterns of population density within its occupied range in the state, it has persistently maintained its center of abundance in the cash-grain region of east-central Illinois since the mid-1930’s ( Fig. 1; Greeley et al. 1962:13-16; Labisky & Anderson 1965:128). The rapid increase in the acreage of row crops (corn and soybeans) in the east-central sector of the state during the 1960's (Joselyn et al. 1968: 217-218) has resulted in a marked reduction of pheas- ant nesting cover, particularly tame hay and small grains. These land-use changes have adversely affected pheasant abundance in this region (Labisky 1968:353- 389). The purpose of this paper is to document the patterns of distribution and abundance of pheasants in Illinois for 1968 and to compare these findings with similar statistics from 1958 and 1963. Some data from the 1958 and 1963 censuses have been presented, in part, in previous publications (Greeley et al. 1962; Labisky & Anderson 1965). ACKNOWLEDGMENTS The author extends his sincere appreciation to the hundreds of postmasters and rural mail carriers in Illinois who voluntarily participated in the 1968 and previous pheasant censuses and whose cooperation was outstanding. Clifford Edwards, President (Enfield, Ill.), and Vernon Maier, Secretary (Clifton, Ill.), Illinois Rural Letter Carriers Association, offered advice in organiz- ing the 1968 census. Officials of the U.S. Post Office Department, M. L. Stover (Washington, D.C.) and Donald L. Swanson and Norbert P. O'Donnell (Chica- go), expedited the project in numerous ways. William L. Preno, Illinois Department of Conserva- tion, Springfield, and my Illinois Natural History Sur- vey colleagues, William L. Anderson and Glen C. Sanderson, read the manuscript. Richard M. Sheets, Survey Illustrator, prepared the maps and designed the cover. Robert M. Zewadski, Survey Associate Editor, edited the manuscript. Dawn A. Labisky as- sisted in assembling the materials for distribution to the postmasters and rural mail carriers. METHODS Statistics on the distribution and abundance of pheasants reported in this paper were voluntarily col- This paper is published by authority of the State of Illinois, IRS Ch. 127, Par. 58.12. Dr. Ronald F. Labisky is an Associate Wildlife pf eee Section of Wildlife Research, Illinois Natural History Survey, rbana, Ronald F. Labisky lected by rural mail carriers during 5-day census periods in April of 1958, 1963, and 1968. These censuses were restricted to the 74 northernmost counties of the state; Greeley et al. (1962:14), after conducting a statewide census in 1957, conservatively classified the remaining 28 of Illinois’ 102 counties as nonpheasant range. The censuses were conducted in April because Greeley et al. (1962:5) concluded: “The conditions for observing pheasants [in Illinois] are probably more nearly constant from year to year in April than in any other month.” The specific 5-day census periods (Monday through Friday) were April 21-25 in 1958 and April 22-26 in 1963 and 1968. The daily census period was between § AM and noon (CST) in all years. Approximately 5 days preceding each of the cen- suses, a packet of materials relating to the census was mailed to the postmaster of each post office having one or more rural routes in the 74 counties to be censused. The packet contained a letter of instruction to the postmaster and a letter of instruction plus a postal-card questionnaire for each rural mail carrier, The postmaster was requested to distribute the latter to each of his rural carriers. Each carrier was asked to report the counties and political townships in which his route was located, the length of his route in miles, and the number of pheasants (cocks and hens) observed along the route on each of the 5 consecutive days specified in the instructions. The number of miles driven and the number of pheasants seen during the 5-day census pe- riod were used to calculate the number of pheasants observed per 100 miles of driving in each township and county. When the route of a mail carrier extended into two or more townships, the number of miles driven and number of pheasants observed were divided equally among the number of townships that he re- ported on his questionnaire. The variables that influ- ence the counts of pheasants by rural mail carriers have been discussed by Greeley et al. (1962:4-5). The information reported by mail carriers during each census was the result of the initial request only; follow-up mailings to prompt the return of question- naires were unnecessary (Table 1). To exemplify, in 1968, 97 percent of all returned questionnaires had been received by May l—only 5 days after the last day of the census period. After each census all postmasters and mail carriers participating in the census were sent a summary of the census findings and a letter thanking them for their cooperation. FINDINGS The relative abundance of pheasants in the 74 north- ernmost counties of Illinois increased from 7.6 to 9.9 pheasants observed per 100 miles of driving (30 per- cent) between April, 1958, and April, 1963, and then declined to 5.5 birds per 100 miles (44 percent) by 4 Table 1.—Comparative statistics obtained from rural mail carrier censuses of pheasants in the 74 northernmost counties (including 1,257 townships) of Illinois during designated 5-day census periods in April of 1958, 1963, and 1968. Category April, 1958 April, 1963 April, 1968 Questionnaires mailed 1,368 1,320 1,256 Number retumed __1,053(77)® —-:1,202(91) 1,143(91) Number usable (of total ) t 1,150( 87) 1,105(88) Number of townships reported (of 1,257) 1,221(97) 1,222( 97) 1,214(97) Total miles driven 250,129 318,605 333,070§ Miles driven per township reported 205 261 268 Cocks observed 10,047 t 17,204 t 10,706 Hens observed 9,044} 14,466 7,545 Total pheasants observed 19,0914 31,6704 18,251 Sex ratio: hens per cock 0.9 0.8 0.7 Cocks per 100 miles 4.0 5.4 3.2 Hens per 100 miles 3.6 45 2.3 Total pheasants per 100 miles 7.64 9.9 5.5 © Percentages are given in parentheses. } Data are not available. t These figures are slightly different from those appearing in Labisky and Anderson (1965:129, Table 1); the figures in the earlier publica- tion were in error. § 8,233 miles of driving were, for the lack of township designations, assigned only to counties. April, 1968 (Table 1). Statistics for hens only, em- ployed to avoid sex-ratio differences among years, re- vealed similar trends in abundance. The number of hens observed per 100 miles increased from 3.6 in 1958 to 4.5 in 1963 (25 percent) and then declined to 2.3 hens per 100 miles in 1968 (49 percent). Thus, pheasants increased in abundance by about one-fourth between 1958 and 1963 and declined in numbers by almost one- half in the succeeding 5 years, 1964-1968. Pheasants were only about three-fourths as abundant in 1968 as in 1958. Although the overall trend in pheasant abundance in Illinois during the past decade was downward, not all portions of the occupied range suffered declines in abundance. In fact, the pheasant range can be sepa- rated into three major sectors with respect to changes in abundance that have occurred within the past 10 years. The east-central portion of the state showed substantial gains (about 50 percent) in abundance be- tween 1958 and 1963; however, the marked decline (about 60 percent) in abundance between 1963 and 1968 more than cancelled the earlier gains, resulting in a net loss of pheasants in Illinois’ prime range (Table 2; Fig. 2-4). All counties of northern Illinois, except Carroll, were characterized by substantial losses in pheasant abundance between 1958 and 1968; the losses incurred between 1958 and 1963 were particularly se- vere. In contrast, the southern and southwestern por- tions of the range occupied by pheasants in Illinois were characterized by very substantial gains in abundance between 1958 and 1968. Table 2.—Relative abundance of pheasants as reported by rural mail carriers in the 74 northernmost counties of Illinois in April 1958, 1963, and 1968. The remaining 28 counties, located in southern Illinois, are not occupied by pheasants. Percentage Change County Total Pheasants Pheasants Per 100 Miles County Rank Reported Per 100 Miles a8 198 158 to to to 1958 1963 1968 1958 1963 1968 1958 1963 1968 1963 1968 1968 Livingston 1 1 1 3,634 8,181 3,006 56.4 99.1 33.3 +76 —66 —4l Logan 16 N7/ 2 305 345 1,543 8.3 9.7 SLL +17 +221 +275 Ford 2 2 3 1,569 2,696 1,144 50.7 75.8 29.2 +50 —61 —42 Woodford Wf U 4 718 852 787 15.6 21.8 21.4 +40 —2 +37 Moultrie 37 15 5 27 301 Bede I Sry oe +707 +460 +1,193 Piatt 12 6 6 305 986 402 11.8 34.6 17.7 +193 —49 +50 McLean 4 4 7 2,071 3,868 1,324 27.7 43.1 17.0 +56 —61 —39 Champaign 5 5 8 1,641 3,018 1,371 25.7 35.6 16.8 +39 —53 —35 Iroquois °3) 3 9 1,991 3,674 1,373 28.9 43.4 ibsz¢ +50 —64 —46 De Witt 19 Vil 10 190 547 405 6.5 16.1 12.2 +148 —24 +88 Putnam 13 9 11 81 131 105 10.8 19.1 10.6 +77 —44 —2 Kankakee 8 8 12 706 973 538 15.3 19.7 10.1 +29 —49 —34 Vermilion 6 16 13 966 728 632 17.3 ili tsil 9.8 —36 —12 —43 Grundy 10 10 14 323 429 283 12.5 gi 9.3 +42 —47 —26 La Salle 11 14 15 873 1,233 820 12.4 13.8 9.1 +11 —34 —27 Douglas 15 13 16 204 487 244 8.7 14.3 8.6 +64 —40 —l Macon 42 19 17 27 326 421 0.7 6.6 8.5 +843 +29 +41,114 Marshall 9 12 18 328 395 176 14.6 15.0 8.2 +3 —45 —44 De Kalb 14 2 19 367 261 396 10.4 5.2 8.0 —50 +54 —2 Tazewell 21 18 20 284 343 356 6.1 8.5 6.5 +39 —24 +7 Mason 30 20 21 66 148 184 2.7 5.4 6.4 +100 +19 +137 Edgar 24 24 22, 190 214 258 5.0 4.2 5.3 —16 +26 +6 Kendall 17 22, 23 135 131 148 7A 48 5.0 —35 t4 —32 Christian 52 32 24 9 67 204 0.2 1.2 3.5 +500 +191 +41,650 McHenry 18 25 25 280 200 183 7.0 3.9 3.4 —44 —13 —5l Coles 31 29 26 60 86 128 2.0 2.3 Se) +15 +39 +60 Lee 20 26 hi 243 147 131 6.2 3.1 2.8 —50 —10 —55 Will 26 27 28 208 179 181 4.6 3.0 iy f —35 —10 —Al The drastic decline in pheasant abundance in east- central Illinois, which has harbored perhaps as many as two-thirds of the state’s pheasant population during the past 30 years, is of critical interest. In both 1958 and 1963, the east-central counties of Livingston, Ford, Iroquois, McLean, and Champaign were, in that order, the five top-ranked counties with respect to relative abundance of pheasants (Table 2). The relative abun- dance of pheasants observed in these counties, collec- tively, was 37.9 and 59.4 birds per 100 miles in 1958 and 1963, respectively. In 1968, these same five coun- ties had a collective relative density of 22.4 pheasants per 100 miles, a decline of 62 percent from 1963. Fur- 5 thermore, by 1968 McLean, Champaign, and Iroquois counties had slipped from their traditional positions among the five top-ranked counties to occupy positions 7, 8, and 9, respectively. Notably, the only segment of range in Illinois to register a measure of abundance exceeding 100 pheasants per 100 miles of driving in 1958, 1963, and 1968 was that located at the junction of Champaign, Vermilion, Douglas, and Edgar counties (Fig. 24). Northern Illinois, characterized by secondary centers of pheasant abundance in the 1940's (Fig. 1), has, in the intervening years, suffered a greater proportionate loss in pheasant numbers than any other sector of the Table 2.—( Continued ) Percentage Change County Total Pheasants Pheasants Per 100 Miles County Rank Reported Per 100 Miles [656 Te 168 by to to to 1958 1963 1968 1958 1963 1968 1958 1963 1968 1963 1968 1968 Menard 43 34 29 8 18 49 0.6 1.0 DS +67 +4130 +283 Boone 23 35 30 112 21 35 5 1.0 2:3; —80 +4130 —55 Ogle 35 Al 31 76 26 119 1.5 0.5 2.0 —67 +300 —33 Carroll 33 33 32, 40 39 65 19 iil 2.0 —42 +82 +5 Kane 28 28 33 116 98 67 3.6 2 Se —25 —37 —52 Sangamon AT 38 34 17 43 101 0.3 0.6 1n5: +100 +4150 +400 Cook 29 30 35 92 61 54 2.8 1.6 1.4 —43 —12 —50 Cass 53 36 36 4 18 32 0.2 0.8 1.2 +300 +50 +500 Du Page 25 23 37 93 117 35 49 4.3 12, —12 —72 —76 Lake 32 31 38 60 66 54 2.0 1.2 ila —40 —8 —45 Bureau 38 42 39 66 29 70 1.3 0.5 1.0 —62 +100 —23 Winnebago 36 37 40 46 22 39 es) 0.6 1.0 —60 +67 —33 Stephenson 22 44 Al 201 18 38 5.4 0.3 0.9 —94 +200 —83 Whiteside 48 47 42 15 13 39 0.3 0.3 0.8 0 +4167 +167 Shelby 54 39 43 11 35 40 0.2 0.5 0.7 +150 +40 +250 Morgan 57 48 44 2 5 14 0.1 0.2 0.4 +100 +100 +300 Mercer 55 43 45 2 15 14 0.1 0.5 0.4 +400 —20 +300 Henry 34 46 46 88 19 16 1.8 0.3 0.3 —83 0 —83 Montgomery 60 54 47 3 4 16 0.1 0.1 0.3 0 +200 +200 Jo Daviess 39 45 48 37 15 11 12 0.3 0.2 —75 —33 —83 Stark 27 40 49 63 9 7 4.2 0.5 0.2 —88 —60 —95 Macoupin® 59 50 0 1 8 0.0 0.0+ 0.1 —} et + Cumberland 50 58 51 6 2 3 0.3 0.1 0.1 —67 0 —67 Fulton 64 52 1 0 5 0.0+ 0.0 0.1 — + aa Rock Island 59 52 53 3 2 3 0.1 0.1 0.1 0 0 0 Peoria 46 49 54 16 6 3 0.4 0.1 0.1 —75 0 —75 Knox 62 57 55 2 3 2 0.0+ 0.1 0.1 + 0 + Effingham 41 56 56 24 2 2 0.8 0.1 0.0+ — _ -— Henderson 45 57 10 0 1 0.5 0.0 0.0-4 0 _ Hancock 58 0 0 2 0.0 0.0 0.0-+ 0 ee +. Jasper 40 50 59 33 4 2 1.0 0.1 0.0+ _ — _ Warren 49 55 60 9 3 2 0.3 0.1 0.0-+4 — — — Crawford 61 0 0 1 0.0 0.0 0.0+ 0 +- te Adams 62 0 0 1 0.0 0.0 0.0-+- 0 + + McDonough 44 63 19 0 1 0.6 0.0 0.0+- — + — Bond 64 0 0 1 0.0 0.0 0.0+- 0 + + Clark 56 51 65 3 5 1 0.1 O01 0.0+ 0 — — Fayette 61 66 4 0 1 0.1 0.0 0.0+- _ + = Clay 51 5 0 0 0.3 0.0 0.0 _ 0 — Greene 58 53 2 3 0 0.1 0.1 0.0 0 _ — Jersey 60 0 1 0 0.0 0.0+- 0.0 + — 0 Madison} 61 0 1 0 0.0 0.0+ 0.0 + _ 0 Pike 63 1 0 0 0.0+ 0.0 0.0 _ 0 — Brown 0 0 0 0.0 0.0 0.0 0 0 0 Schuyler 0 0 0 0.0 0.0 0.0 0 0 0 Scott 0 0 0 0.0 0.0 0.0 0 0 0 ° No rank was assigned to counties from which pheasants were absent in year of census. ; ; + Percentage change presented only as ++ (gain), — (loss), or 0 (no change) for those counties from which pheasants were reported absent (including 0.0 + ) in any of the three census periods. 1 Richland County was censused in place of Madison County in 1958. 6 state’s pheasant range (Fig. 2-4). Although the pheas- ant population in northern Illinois had already declined by 1958, it diminished even more during the past decade. The abundance of pheasants in all counties of northern Illinois, except Carroll, diminished substan- tially between 1958 and 1968; losses in abundance dur- ing these 10 years exceeded 50 percent in Jo Daviess, Stephenson, Boone, McHenry, Kane, Cook, Du Page, Lee, Henry, and Stark counties (Table 2). Pheasant populations in northern Illinois would have plummeted even lower had not such counties as Carroll, Stephen- son, Winnebago, Boone, De Kalb, and Ogle actually registered gains in abundance between 1963 and 1968. The most spectacular change in pheasant abundance that has occurred in Illinois within the past decade has been the increase in pheasant numbers in a block of nine counties—Logan, Moultrie, De Witt, Macon, Mason, Christian, Menard, Sangamon, and Cass—sit- uated south and southwest from east-central Illinois. Increases in abundance among these nine counties, all of which had densities of more than one pheasant per 100 miles of driving in 1968, ranged from twofold to sixteenfold during the 10 years, 1958-1968 (Table 2; Fig. 2-4). Notably, two of these counties—Logan and Moultrie—advanced from their respective county rank- ings of 16 and 37 in 1958 to rankings of 2 and 5 in 1968; these advances in ranking were due not only to gains in abundance recorded in the two counties but also to declines in abundance in many of oe 4 i il ite - feo Le i i fi ile ide APRIL, 1958 PHEASANTS PER 100 MILES MB 100! PLUS 50.1—100.0 10.1 — 50.0 1.1 —10.0 0.1 - 1.0 LESS THAN OI Fig. 2.—Distribution and abundance of pheasants in Illinois as mapped from township statistics obtained by a rural mail carrier census, April, 1958 (after Labisky and Anderson 1965). Counties below the heavy line were not censused. Ba (0) naa =] the previously top-ranked counties. Several counties that adjoin the nine-county sector, namely Piatt, Taze- well, Edgar, and Coles, also posted substantial, though proportionately smaller, gains in pheasant abundance during the past decade. No decisive spread of pheasants into unoccupied range along the southern and western boundaries of the contiguous pheasant range has occurred within the past decade (Fig. 24). Yet pheasant abundance in many segments of this so-called “marginal” range, par- ticularly in the south, has increased in recent years. The several small populations of pheasants beyond the con- tiguous range could be precursory, self-maintaining flocks, but are probably the artifacts of local releases of artificially propagated birds. DISCUSSION Examination of records documenting the distribution and abundance of pheasants since their introduction into Illinois (Greeley et al. 1962), leaves little doubt that the state’s all-time high in pheasant abundance was attained in 1962 or 1963. This conclusion is sub- stantiated by the fact that the calculated autumn kill of cocks by hunters in Illinois was greater in these years— 989,000 in 1962, and 1,066,000 in 1963—than in any pre- ceding or following year through 1968 ( William L. Pre- no, Illinois Department of Conservation, unpublished ). The low in pheasant numbers in Illinois during the past ST Bue = tia (I APRIL, 1963 PHEASANTS PER _1OOMILES 100.1 PLUS 50.I — 100.0 10.1 -50.0 1 —10.0 0.1 — 1.0 LESS THAN O.I Fig. 3.—Distribution and abundance of pheasants in Illinois as mapped from township statistics obtained by a rural mail carrier census, April, 1963 (after Labisky and Anderson 1965). Counties below the heavy line were not censused. UEUURE il decade occurred in 1966; the 1968 breeding * population In the 5 years following the pheasant high of 1963, was estimated to be about 10 percent above that of the abundance of pheasants in Illinois declined by 1966 (William L. Preno, unpublished). about 50 percent. Much of this decline in abundance, ge | 4p | Sek, \ eae , c. + ACOUPI =) foe P= APRIL, 1968 - PHEASANTS PER 100 MILES ae 100.1 PLUS Re 50.1 — 100.0 (MM) 10.1-S0.0 4 1.1-10.0 (i 01-10 [__] LESS THAN O1! HAMILTON | WHITE Fig. 4.—Distribution and abundance of pheasants in Illinois as mnERe sd from to Sat statistics obtained by a rural mail carrier census, April, 1968. Counties b« 15 ow the heavy line were no ee snsus 8 particularly in the prime pheasant range in east-central Illinois, resulted from the rapid decrease in acreages of tame hays and other suitable nesting habitats. The increase in acreages planted to the major cash-grain crops, corn and soybeans, has brought about this de- crease in nesting habitats (Labisky 1968:353-389; Joselyn et al. 1968:217-218). The drought of 1964 de- pressed pheasant production and also contributed to the decline in abundance. It is most unlikely that substantial gains in pheasant abundance will occur in east-central Illinois in future years unless there is a reversal of agricultural trends, which seems most improbable. In fact, it is doubtful that pheasants will be able to maintain existing popu- lation levels for long under today’s farming practices. To illustrate, the obvious fragmentation of Illinois’ good pheasant range in 1968, as evidenced by the intersper- sion of areas of lower pheasant abundance with those of higher abundance (Fig. 4), suggests that further declines in abundance will be forthcoming. Livingston County, the top-ranked pheasant county in the state, shows signs of being cleaved from northwest to south- east by a trough of lower pheasant abundance. Other top-ranked counties, notably Champaign, Iroquois, and McLean, exhibited noticeable range fragmentation be- tween 1963 and 1968 (Fig. 3 and 4). In 1968 the only sector of pheasant range in the state showing encouraging increases in pheasant numbers was in that block of counties (Logan, Moultrie, De Witt, Macon, Mason, Christian, Menard, Sangamon, and Cass) constituting the southwestern portion of IIli- nois’ occupied pheasant range. Gains in pheasant num- bers in this area, much of which was classified as medi- ocre to poor with respect to abundance as recently as the early 1960's, were unusually great (Table 2; Fig. 2-4). In fact, within this sector of pheasant range a secondary center of abundance has appeared in north- ern Logan County. Not only did this southwest sector of range contain more pheasants in 1968 than it ever had, but it also contained the highest density of pheas- ants ever recorded along such a broad front so far south in Illinois. Similarly, a surge in pheasant abun- dance, beginning in the late 1950's, occurred in an area of traditionally poor pheasant range in southwestern Iowa (Klonglan 1962); this buildup in pheasant num- bers has already persisted for a decade. A decisive southward shift in pheasant abundance has occurred in Illinois from 1948 to 1968. Northern Illinois, which contained strong secondary centers of pheasant abundance in the late 1940's, had, by 1968, exhibited greater proportionate declines in pheasant numbers than any other major region of the state. As pheasant abundance declined in northern Illinois, it increased in the prime range of east-central Illinois— reaching an all-time high about 1963. Yet in 1968, east- central Illinois, despite sustaining a 60-percent decline in the pheasant population from that of 1963, still con- tained the state’s center of pheasant abundance as it had in the preceding 30 years. However, during the 1960's a comparatively spectacular gain in abundance occurred in a large and contiguous segment of range (4-90587—5M—5-69 ) located south-southwestward from east-central Illinois and north of the southern boundary of the contiguous pheasant range in Illinois. If pheasant abundance con- tinues to dissipate in east-central Illinois as a result of current agricultural practices, the southwestern portion of range may replace east-central Illinois as the state’s center of pheasant abundance. Even though popula- tion abundance has increased along the southernmost boundary of the pheasant’s range in Illinois in recent years, there has been no real southward extension of the bird’s range in the state. SUMMARY The relative abundance of pheasants per 100 miles of driving in the 74 northernmost counties of Illinois, as reported by rural mail carriers during April censuses, was 7.6 in 1958, 9.9 in 1963, and 5.5 in 1968. Thus, pheasants increased in abundance by about 30 percent between 1958 and 1963 and then declined by about 44 percent between 1963 and 1968. The all-time high in pheasant abundance in Illinois was probably reached in 1962 or 1963. Although the number of pheasants in Illinois has de- clined during the past decade, not all portions of the state’s pheasant range have suffered similarly. Pheasant abundance in east-central Illinois, the state’s prime pheasant range since the 1930's, increased about 50 percent between 1958 and 1963 and then declined by about 60 percent between 1963 and 1968. Pheasants in northern Illinois were less than half as abundant in 1968 as in 1958. In contrast, a severalfold increase in pheasant abundance occurred between 1958 and 1968 in a large, contiguous unit of range immediately south- southwestward from east-central Illinois. Notably, Illinois has experienced a significant, pro- gressive southward shift in pheasant abundance within the past 20 years. Yet, despite the recent surge in pheas- ant abundance in the southern portion of the range occupied by pheasants, there has been no permanent southward extension of the birds’ range in the state. LITERATURE CITED GREELEY, FREDERICK, RoNALD F. Lasisky, and Stuart H. Mann. 1962. Distribution and abundance of pheasants in Illinois. Illinois Natural History Survey Biological Notes 47. 16 p. Josetyn, G. BLam, JOHN E. WaRNocK, and STANLEY L. Etter. 1968. Manipulation of roadside cover for nesting pheasants—a preliminary report. Journal of Wildlife Management 32(2):217—233. KLONGLAN, EuGENE DAWAyYNE. 1962. Ecology of pheasant production in southwestern Iowa. Ph.D. Thesis. Iowa State University of Science and Technology, Ames. 343 p. Lapisky, RonaLp F. 1968. Ecology of pheasant populations in Illinois. Ph.D. Thesis. University of Wisconsin, Madison. 511 p. Labisky, RONALD F., and WitittaM L. ANDERSON. 1965. Changes in distribution and abundance of pheasants in Illinois: 1958 versus 1963. Illinois State Academy of Science Transactions 58(2):127—135. Robertson, WILuiAM B., JR. 1958. Investigations of ring- necked pheasants in Illinois. Illinois Department of Conservation Technical Bulletin 1. 137 p. Ne eae ee SS * Maceo icy ee aS Sel se aT