QE 841 .U62 PT . 1 'ERSITY OF KANSAS miscellaneous EUM OF NATURAL HISTORY publication No. 73 Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part I. Fossil Fishes B/ H.-P. Schultze, J. D. Stewart, A. M. Neuner, and R. W. Coldiron UNIVERSITY OF KANSAS LAWRENCE 1982 Ocfober 6, 1982 H 1 16 UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY The University of Kansas Publications, Museum of Natural History, beginning with volume 1 in 1946, was discontinued with volume 20 in 1971. Shorter research papers formerly published in the above series are now published as Occasional Papers, Museum of Natural History. The Miscellaneous PubUcations, Museum of Natural History, began with number 1 in 1946. Longer research papers are pub- lished in that series. Monographs of the Museum of Natural History were initiated in 1970. All manuscripts are subjected to critical review by intra- and extramural specialists; final acceptance is at the discretion of the Director. Institutional libraries interested in exchanging publications may obtain the Occa- sional Papers and Miscellaneous Publications by addressing the Exchange Librarian, University of Kansas Library, Lawrence, Kansas 66045. Individuals may purchase separate numbers of all series. Prices for all publications of the Museum may be obtained from the Publications Secretary, Museum of Natural History, University of Kansas, Lawrence, Kansas 66045. HARVARD UNIVERSITY "^ Library of the Museum of i m Comparative Zoology The University of Kansas Museum of Natural History Miscellaneous Publication No. 73 October 6, 1982 Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part I. Fossil Fishes By H.-P. ScHULTZE, J. D. Stewart, and A. M. Neuner, Museum of Natural History, University of Kansas Lawrence, Kansas 66045 and R. W. COLDIRON 1065 Miramonte Avenue, Los Altos, California 94022 The University of Kansas Lawrence 1982 L University of Kansas Publications, Museum of Natural History Editor: Robert M. Mengel Managing Editor: Joseph T. Collins Miscellaneous Publication No. 73 pp. 1-53 Published October 6, 1982 Museum of Natural History The University of Kansas Lawrence, Kansas 66045 U.S.A. OCT 2 ;: 1982 ARVAFJD Printed by University of Kansas Printing Service Lawrence, Kansas CONTENTS INTRODUCTION 1 SHORT HISTORY OF THE COLLECTION 2 ACKNOWLEDGMENTS 3 CATALOGUE 4 Class Chondrichthyes 4 Subclass Subterbranchialia 4 Subclass Elasmobranchii 4 Enterospira 16 Class Osteichthyes 17 Subclass Actinopterygii 17 Superordpr Chondrostei 17 Superorder Neopterygii 21 Division Pycnodontiformes 21 Division Ginglymodi 23 Division Halecostomi 23 Subdivision Halecomoi-phi 23 Subdivision Teleostei 25 Teleostei inc. sed. .-- 46 Subclass Sarcopteiygii 46 Superorder Actinistia 46 Superorder Choanata 47 Superorder Dipnoi 47 LITERATURE CITED 49 APPENDIX. GENERA AND SPECIES LISTED 51 INTRODUCTION The International Code of Zoological Nomenclature (1964, Art. 72D) recom- mends that each institution "publish lists of type-material in its possession or cus- tody." In accordance with this recom- mendation, the following catalogue of fossil fish types and of figured fossil fishes in the University of Kansas Museum of Natural History fossil vertebrate collec- tion (KUVP) is presented. The cata- logue includes figured specimens which could not be relocated in the collection. Specimens are grouped according to the following classifications: Zangerl 1981 (Chondrichthyes), SchaefFer 1973 (Chondrostei), Patterson and Rosen 1977 (Neopterygii) and Schultze 1981 A ( Sarcopterygii ) . Class Chondrichthyes Subclass Subterbranchialia Superorder Holocephalia Order Bradyodontida Suborder Chimaerina Subclass Elasmobranchii Order Xenacanthida Order Cladoselachida Order Eugeneodontida Superfamily Caseodontoidea Superfamily Edestoidea Order Petalodontida Order Euselachii Superfamily Protacrodontoidea Superfamily Ctenacanthoidea Superfamily Hybodontoidea Superfamily Batoidea Superfamily Squalimorphii ( Hexanchiformes, Squalifonnes, Pristiophorif ormes ) Superfamily Galeomoqihii Family inc. sedis Ptychodontidae Class Osteichthyes Subclass Actinopterygii Superorder Chondrostei Order Palaeoniscifomies Order Haplolepidida Order Tarrasiida Superorder Neopterygii Division Pycnodontiformes Division Ginglymodi Division Halccostomi Subdivision Halecomorphi Order Pachycormifonnes Order Amiiformes Order Aspidorhynchif ormes Subdivision Teleostei Order IchthyodectifoiTnes Suborder Ichthyodectoidei Order Osteoglossiformes Suborder Osteoglossoidei Order Elopiformes Suborder Elopoidei Order Anguilliformes Suborder Anguilloidei Order Leptolepiformes Order Culpeifomies Order Euteleostei Suborder Ostariophysi Suborder Protacanthopterygii Suborder Neoteleostei Infraorder Aulopifomies Infraorder Paracantliopterygii Infraorder Acanthopterygii Subclass Sarcopterygii Superorder Actinistia Order Coelacanthida Superorder Choanata Order Onychodontida Superorder Dipnoi The following information is included concerning each cited specimen: cur- rently used catalogue number and old number if used in publication; name published under; author and year of pub- lication; figure; preserved anatomical part(s) of the specimen; geologic stage, formation, member (see Zeller 1968); lo- cality (Bardack 1965A) and collector MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY ( where known ) . Within the Cretaceous, the specificity of assignments to geo- logical stage varies for old localities de- pending on our knowledge of each lo- cality. The areal distribution of outcrops correlated with the various stages is known and a locality can be assigned to stage if its location is well enough de- fined. Within one species, the specimens are arranged first after their type cate- gory and second in order of the appear- ance of the figures within the publica- tion. SHORT HISTORY OF THE COLLECTION The fossil vertebrate collection at the University of Kansas dates back to the 1870's. In 1877 and 1878, F. H. Snow guided two collecting parties to western Kansas, which brought back "41 sauri- ans, 6 pterodactyles and 117 fishes" (Snow 1880) from the Cretaceous of Gove County. However, these were not the first fossil vertebrates in the Museum of Natural Histoiy. They were preceded — as far as can be traced — by a large Bananogtnius from the Cretaceous of western Kansas, collected by F. F. Dins- more (1875), and a trackway from the Upper Pennsylvanian of eastern Kansas collected by F. B. Mudge (1873). Dur- ing the years following, new specimens were added by acquisition from profes- sional collectors such as E. P. West and C. H. Sternberg. In 1890 the situation changed drastically with the appoint- ment of S. W. Williston. Over the next few years he guided parties of many students to the Cretaceous of Western Kansas, the Badlands of South Dakota, the lurassic of Wyoming, and other places. This productive collecting era ended with the departure of Williston to Chicago in 1902. C. E. McClung and H. T. Martin continued to collect, usu- ally at a slower pace, although the years 1910 and 1911 are an exception with the large parties in the Cretaceous of west- ern Kansas and the Oligocene-Miocene of northeast Colorado. McClung left the University of Kansas in 1912, and H. T. Martin took over the responsibility for the fossil vertebrate collection. His col- lecting focused on the Miocene and Pleistocene of Kansas, and the Oligo- cene-Miocene of northeast Colorado. These interests in the Pleistocene and Tertiary were continued by C. W. Hib- bard, who followed H. T. Martin as curator of the fossil vertebrate collec- tion. In 1931 and 1935 Miocene fishes from the diatomite of the Ogallala For- mation of Logan County western Kansas were collected by K.U. field parties (C. W. Hibbard, W. K. McNown and M. K. Elias, and D. H. Dunkle respec- tively). With the appointment of R. W. Wilson in 1947, collecting emphasis changed to fossil mammals from the early Tertiary of Wyoming, New Mexico and Utah. While Wilson was curator, F. E. Peabody collected at Ft. Sill, Okla- homa, and he excavated the important Pennsylvanian site near Garnett, Kansas, discovered in the early 1930's. Peabody left the Department of Zoology in 1956, and the museum was able to establish a second curatorial position for the collec- tion of lower fossil vertebrates. But even with the appointment of T. Eaton in 1958, the main additions to the collec- tion were still Cenozoic and Cretaceous mammals gathered by W. C. Clemens, C. C. Black, and L. D. Martin. Paleozoic fossils occurring in the eastern part of Kansas were only occasionally collected. The Sage nodus material from the Penn- sylvanian of northeast Kansas was col- lected by L W. Beede in 1899, and addi- tional extensive collecting was done by J. Chorn, 1974-79. The first braincases of Pennsylvanian palaeoniscoids were discovered by the geologists W. H. Twenhofel and C. Dunbar in 1914, al- though the locality at Garnett was first discovered and excavated by geologists. The latter locality was excavated further by Lane, McNown, D. H. Dunkle, C. W. Hibbard, and others in 1932 and 1936, before Peabody started extensive excava- tions in 1953 and 1954. In the 20's and TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 30's a large and important collection of fossil braincases of Pennsylvanian palae- oniscoids were brought together by a private collector, A. Bridvvell. In conclusion, the fossil vertebrate collection at the University of Kansas Museum of Natural History contains fos- sil mammals from a number of regions and horizons. Fossil lower tetrapods and fishes are mainly from the Cretaceous of western Kansas, and from the Pennsyl- vanian and Permian of eastern Kansas. Intensive collecting in these areas began again with L. D. Martin as curator at the beginning of the 1970's and II. -P. Schultze added a new emphasis on the Devonian since 1979. We were not able to relocate three of the 57 holotypes, four of the 55 syn- types, and 53 of the 355 figured speci- mens. Forty-one holotypes, 49 syntypes and all paratypes are valid, while the single neotype, 13 holotypes and two syntypes have been synonymized. Ten holotv'pes represent the type-species of the genus. ACKNOWLEDGMENTS We thank our colleagues L. D. Mar- tin, R. W. Wilson, J. Chorn, and G. Arratia, Museum of Natural History, Lawrence, and C. Poplin, Museum Na- tional d'Histoire Naturelle, Paris, for their help with various matters. We are especially grateful to R. Kurtz and the typing service of the Division of Biology for arranging and typing of this manu- script. The catalogue could not have been completed without special support for this project by P. S. Humphrey, Director of the Museum of Natural History. KUVP 888 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY CATALOGUE CLASS CHONDRICHTHYES Huxley 1880 Subclass Subterbranchialia Zangerl 1979 Superorder Holocephali Bonaparte 1832 Order Bradyodontida Woodward 1921 Suborder Chiniaerina Zangerl 1981 Family Deltodontidae Zangerl 1981 Dehodus Morris & Roberts 1862 ex Agassiz Ms. Deltodus attenuatus Branson 1905 Deltodus attenuatus sp. nov. — Branson 1905: p. 33, pi. 1, fig. 6; holotype, tooth plate. Upper Pennsylvanian, "Missouri Coal Measure"; near Kansas City, Missouri, collected by W. J. Parrish before 1885. KUVP 1136 KUVP 1023 KUVP 753 KUVP 1056 Family Chimaeridae Rafinesque 1815 Ichthypriapus Hibbard 1942 Ichthypriapu.s hu])bsi Hibbard 1942 This species was described as "?head clasper of a chimaeroid." It was found in close association with specimen KUVP 465 (not figured), which consisted of the large pectoral fins of a teleost, Protosphyraena gladius Cope. The hard tissue of the separately described piece is bone (not cartilage), a tissue atypical for chondrichthyes. Thus the piece has to be considered as truly part of the specimen KUVP 465. Ichthypriapus huhhsi gen. et sp. nov. — Hibbard 1942: pp. 237- 240; text-fig. 1-4; holotype, mistakenly considered as chimaerid ?head clasper ( see p. 00 ) . Upper Cretaceous, Santonian or Campanian, Niobrara Fm., Smoky Hill Chalk Member; Logan Co., Kansas; collected by C. Sternberg ?1912. Subclass Elasmobranchii Bonaparte 1838 Order Eugeneodontida Zangerl 1981 Superfamily Caseodontoidea Zangerl 1981 Family Caseodontidae Zangerl 1981 Caseodus Zangerl 1981 Dome-shaped symphysial teeth are characteristic for Caseodus ac- cording to Zangerl 1981, thus the teeth published by Eaton (1962) are referred to the new genus. Caseodus gigas (Eaton 1962) Fadenia gigas new species — Eaton 1962: p. 353; text-fig. 2; holo- type, 2 symphysial teeth. Fadenia gigas new species — Eaton 1962: p. 354; paratype pub- lished erroneously as KUVP 865, 2 symphysial teeth. Middle Pennsylvanian, Desmoinesian stage, label says: "4 feet from top of Cherokee Shale [=Cherokee Group], (Lexington coal) [=within Marmaton Group]"; at Lucas, Henry County, Missouri. Caseodus sp. Campodus variahihs — Eaton 1962: text-fig. 1; symphysial tooth. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS Upper Pennsylvanian, Virgilian Stage, Shawnee Group, Oread Limestone, Plattsmouth Limestone Member; SWJi, Sec. 27, T14S, R20E, N of Baldwin, Douglas Co., Kansas; collected by A. Bridwell. Superfamily Edestoidea Hay 1929 Family Agassizodontidae Zangerl 1981 Helicoprion Kaipinsky 1899 Helicoprion sp. KUVP 30606 Helicoprion sp. indet.— Chorn 1978: text-fig. 1, part of tooth whorl. Lower Pennian, Leonardian, Bone Spring Formation; 0.4 km E of Williams Ranch in Bone Canyon, Culberson Co., Texas; col- lected by F. R. West. Order Petalodontida Zangerl 1981 Petalodus Owen 1845 Petalodus ohioensis Safford 1853 KUVP 9972 Petalodus jewetti Miller, n. sp.— Miller 1957: pp. 82-83; text-fig. 1-2; holotype, tooth. Upper Pennsylvanian, Virgilian Stage, Shawnee Group, Oread Limestone, Plattsmouth Limestone Member; quarry at SEK, Sec. 20, T15S, R18E, Franklin Co., Kansas; collected by J. M. Jewett 1957. Megactenopetalus David 1944 Megactenopetalus kaihahanus David 1944 KUVP 57433 Megactenopetalus kaihahanus— Hansen 1980: text-fig. IB; cusp from upper dental element, cast. Upper Pennsylvanian, Virgilian Stage, Shawnee Group, Lecomp- ton Limestone, Beil Limestone Mbr.; SW^l, Sec. 13, T12S, R18E, Douglas Co., Kansas. Ctenoptychius Agassiz 1838 Ctenoptychius semicircularis Newberry and Worthen 1866 KUVP 14386 Ctenoptychius semicircularis Newbeny and Worthen — Martin 1913: pi. 21; 1 tooth. ? Pemiian (or Uppemiost Pennsylvanian); 30 miles E of Arkansas City, southern Kansas. Order Euselachii Hay 1902 Superfamily Protacrodontoidea Zangerl 1981 Holmesella Gunnell 1931 Holmesella ? sp. KUVP 1088 Holmesella ? sp.— 0rvig 1966: text-fig. IF-H (specimens illus- trated in text-fig. lA and 5A, B not relocated); pi. 1 and 2; scales. Upper Pennsylvanian, Missourian Stage, Kansas City Group, Tola Limestone, Muncie Creek Shale; 3 or 4 miles NW of Independence, Montgomery Co., Kansas; collected by R. C. Moore. Superfamily Hybodontoidea Zangerl 1981 KUVP 57406 hybodont— Maisey 1982: fig. 16 J,K; cephaUc spine. MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Upper Pennsylvanian, Virgilian, Bern Limestone Fomiation, Sol- dier Creek Shale Member after Sawin et al. 1979 (Biirlingame Limestone Member after Zeller 1968); creek bank 1 mi. N and % mi. E of Robinson, SWA, SWK, NW)^, Sec. 34, T2S, RISE, Brown Co. (mistakenly listed as Shawnee Co.), Kansas; collected by J. Chorn 1978.^ Superfamily Galeomorphii Compagno 1973 Family Odontaspididae Miiller and Henle 1839 Scapanorhynchus Woodward 1889 Scopanorliynchiis rhopJiiodon ( Agassiz 1843) KUVP ? Scapanorhynchus rhaphiodon Agassiz — Williston 19(X)A: pi. 8, fig. 4; pi. 14, fig. 5. Scapanorhynchus rhaphiodon Agassiz — Williston 1900B: pi. 26, fig. 4; pi. 32, fig. 5. Scapanorhynchus — Lane 1945: pi. 1, fig. 5 ( =Williston 1900A, pl. 14, fig. 5). Not relocated. See also Cretoxyrliina mantelH. Family Cretoxyrhinidae Gliickman 1958 Cretolamna Gliickman 1958 CretoJamna appendiculata (Agassiz 1843 KUVP 55227 Lamna, sp.— Williston 1900A: pl. 12, fig. 5. Clark Co., Kansas; col- Lamna, sp.— Williston 1900B: pl. 30, fig. 5. Isolated tooth. KUVP 55228 Lanma, sp.— Williston 1900A: pl. 12, fig. 6. Lamna, sp.— Williston 1900B: pl. 30, fig. 6. Isolated tooth. Lower Cretaceous, Albian, Kiowa Fm. lected by S. W. Williston. KUVP ? Lamna, sp.— Williston 1900A: pl. 12, fig. 12. Lamna, sp.— Williston 1900B: pl. 30, fig. 12. Tooth, not relocated. KUVP 55231 Lamna appendiculata Roemer— Williston 1900A: pl. 13, fig. 47; pl. 14, fig. 3A. Lamna appendiculata Roemer — Williston 1900B: pl. 31, fig. 47; pl. 32, fig. 3A. Lamna— Lane 1945: pl. 1, fig. 3 A (= Williston 1900A, pl. 14, fig. 3A). Tooth. Upper Cretaceous, Turonian, PEagleford Group; Dallas County, Dallas, Texas; collected by F. Hunston. KUVP ? Lamrui appendicidata Roemer — Williston 1900A: pl. 13, fig. 48; pl. 14, fig. 3B. Lamna appendicidata Roemer — Williston 1900B: pl. 31, fig. 48; pl. 32, fig. 3B. Lamna— Lane 1945: pl. 1, fig. 3B ( =Williston 1900A: pl. 14, fig. 3B). Tooth; not relocated. See also Cretoxyrhina niantelli. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS Cretoxijrhina Gliickman 1958 Cretoxyrhina mantelli ( Agassiz 1843) KUVP 13522 Scapanorh/nchus rluipiodon Agassiz— Williston 1900A: pi. 8, fig. 2; pi. 14, fig. 4. Scapanorhynchus rhaphiodon Agassiz — ^Williston 1900B: pi. 26, fig. 2; pi. 32, fig. 4. Scapanorhynchus— Lime 1945: pi. 1, fig. 4 (=Williston 1900A, pi. 14, fig. 4). Tooth. Upper Cretaceous, Senonian, probably Niobrara Fm., Smoky Hill Chalk Mbr.; western Kansas. KUVP ? Lamna appendicuhta Agassiz— Williston 1900A: pi. 8, fig. 3. Lamna appendictduta Agassiz — Williston 1900B: pi. 26, fig. 3. Tooth; not relocated. KUVP ? Oxyrhina mantelli Agassiz— Williston 1900A: pi. 8, fig. 5; pi. 14, fig. 2J. Isurus mantelli Agassiz— Williston 1900B: pi. 26, fig. 5; pi. 32, fig- 2J. Isurus— Lane 1945: pi. 1, fig. 2J (=Williston 1900A: pi. 14, fig- 2J). Tooth; not relocated. KUVP 64541 Oxyrhina mantelli Agassiz— Williston 1900A: pi. 13, fig. 41; pi. 14, fig. 2. Isurus mantelli Agassiz— Wilhston 1900B: pi. 31, fig. 41; pi. 32, fig. 2. Isurus— Lane 1945: pi. 1, fig. 2 (=Williston 1900A, pi. 14, fig. 2). Tooth. KUVP 64542 Oxyrhina mantelli Agassiz— Williston 1900A: pi. 13, fig. 42; pi. 14, fig. 2A. Isurus mantelli Agassiz— Williston 1900B: pi. 31, fig. 42; pi. 32, fig. 2A. Isurus— Lane 1945: pi. 1, fig. 2A (=Winiston 1900A, pi. 14, fig. 2A). Tooth. KUVP 64543 Oxyrhina mantelli Agassiz— Williston 1900A: pi. 13, fig. 43; pi. 14, fig. 2C. Isurus mantelli Agassiz— Williston 1900B: pi. 31, fig. 43; pi. 32, fig. 2C. Isurus— Lane 1945: pi. 1, fig. 2C (= Williston 1900A, pi. 14, fig. 2C). Tooth. KUVP 64544 Oxyrhina mantelli Agassiz— Williston 1900A: pi. 13, fig. 44; pi. 14, fig. 2D. Isurus mantelli Agassiz— Williston 1900B: pi. 31, fig. 44; pi. 32, fig. 2D. Isurus— Lane 1945: pi. 1, fig. 2D (= Wilhston 1900A, pi. 14, fig. 2D). Tooth. KUVP 64541-64544 Upper Cretaceous, Senonian, Niobrara Fm., Smoky Hill Chalk Mbr.; Gove Co., Kansas; collected by Cooper. MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP ? Oxyrhina manteUi Agassiz— Williston 1900A: pi. 13, fig. 45; pi. 14, fig. 2F. hums mantelli Agassiz — Williston 1900B: pi. 31, fig. 45; pi. 32, fig. 2F. Isiirtis— Lane 1945: pi. 1, fig. 2F (=Williston 1900A, pi. 14, fig. 2F). Tooth; not relocated. KUVP ? Oxi/rhina munteUi Agassiz— Williston 1900A: pi. 13, fig. 46; pi. 14, fig. 2G. Isiirus mantelli Agassiz — Williston 1900B: pi. 31, fig. 46; pi. 32, fig. 2G. Z5Mr«5— Lane 1945: pi. 1, fig. 2G (=Williston 1900A, pi. 14, fig. 20). Tooth; not relocated. KUVP 64545 Oxijrhina mantelli Agassiz— WilHston 1900A: pi. 13, fig. 50; pi. 13, fig. 2E. Isurus mantelli Agassiz — Williston 1900B: pi. 31, fig. 50; pi. 32, fig. 2E. hums— Lane 1945: pi. 1, fig. 2E (^WilHston 1900A, pi. 14, fig. 2E). Tooth. KUVP 64546 Oxyrhina mantelli Agassiz— WilHston 1900A: pi. 13, fig. 51; pi. 14, fig. 2F. hums mantelli Agassiz — Williston 1900B: pi. 31, fig. 51; pi. 32, fig. 2F. hums— Lane 1945: pi. 1, fig. 2F (^WilHston 1900A, pi. 14, fig. 2F). Tooth. KUVP 64547 Oxyrhina mantelli Agassiz— Williston 1900A: pi. 13, fig. 52; pi. 14, fig. 2B. hurus mantelli Agassiz — Williston 1900A: pi. 31, fig. 52; pi. 32, fig. 2B. hums— Lane 1945: pi. 1, fig. 2B (= WilHston 1900A, pi. 14, fig. 2B). Tooth. KUVP 64545-64547 Upper Cretaceous, Senonian, Niobrara Fm., Smoky HiH Chalk Mbr.; Gove Co., Kansas; collected by Cooper. KUVP ? Oxyrhina mantelli Agassiz— Williston 1900A: pi. 14, fig. 2H. Isurus manteUi Agassiz— Williston 1900B: pi. 32, fig. 2H. hurus— Lane 1945: pi. 1, fig. 2H (= WilHston 1900A, pi. 14, fig. 2H). Tooth; not relocated. KUVP ? Oxyrhina, mantelli Agassiz— Williston 1900A: pi. 14, fig. 21. hums mantelli Agassiz— Williston 1900B: pi. 32, fig. 21. hurus— Lane 1945: pi. 1, fig. 21 (=Williston 1900A, pi. 14, fig. 21). Tooth; not relocated. KUVP ? Oxyrhina mantelli Agassiz— Williston 1900A: pi. 14, fig. 2K. Isurus manteUi Agassiz— Williston 1900B: pi. 32, fig. 2K. hurus— Lane 1945: pi. 1, fig. 2K (=Williston 1900A, pi. 14, fig. 2K). TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS Tooth; not relocated. KUVP ? Oxyrhina mantelU Agassiz— Williston 1900A: pi. 14, fig. 2L. Imnis mantcUi Agassiz — Williston 1900B: pi. 32, fig. 2L. Isums—L^ne 1945: pi. 1, fig. 2L (^Williston 1900A, pi. 14, fig. 2L). Tooth; not relocated. KUVP ? Oxyrhina manteUi Agassiz— Williston 1900A: pi. 14, fig. 2M. hums manteUi Agassiz— Williston 1900B: pi. 32, fig. 2M. Isurus— Lane 1945: pi. 1, fig. 2M (^Williston 1900A, pi. 14, fig. 2M). Tooth; not relocated. KUVP 64548 Oxyrhina manteUi Agassiz— Williston 1900A: pi. 14, fig. 6. Isurus inantelli Agassiz — Williston 1900B: pi. 32, fig. 6. Istirus— Lane 1945: pi. 1, fig. 6 (= Williston 1900A, pi. 14, fig. 6). Tooth. Upper Cretaceous, Senonian, Niobrara Fm., Smoky Hill Chalk Mbr.; Gove Co., Kansas; collected by Cooper. KUVP ? Oxyrhina 7nantelli Agassiz— Williston 1900A: pi. 14, fig. 7. Isurus manteUi Agassiz — Williston 1900B: pi. 32, fig. 7. Isurus— Lane 1945: pi. 1, fig. 7 (^Williston 1900A, pi. 14, fig. 17). Tooth; not relocated. KUVP 64549 Oxyrhina 7nanteUi Agassiz— Williston 1900A: pi. 14, unnumbered. Isurus manteUi Agassiz — Williston 1900B: pi. 32, unnumbered. 75i/m.s— Lane 1945: pi. 1, unnumbered (=Williston 1900A, pi. 14, unnumbered). Tooth. Upper Cretaceous, Senonian, Niobrara Fm., Smoky Hill Chalk Mbr.; Gove Co., Kansas; collected by Cooper. Leptostyrax Williston 1900A We follow here the usage of Meyer (1974), who emended Lep- tostyrax Williston 1900A to include all primitive cretoxyrhinids with labial plications on most teeth, with the linguo-apical part of the root nearly pei-pendicular to the face of the crown, and with- out a nutritive groove. This includes species previously assigned to PUcatolamna Henrian 1975 (e.g. "Lamna" machrorhiza Cope, "Otodus" semipUcatus Agassiz, "Otodus" sulcatus Geinitz, and "Oxyrhina" crassidens Dixon). This usage was also accepted by Martin and Stewart 1977. Leptostyrax crassidens (Dixon 1850) KUVP 55230 Lamna appendiculata Roemer— Williston 1900A: pi. 13, fig. 49; pi. 14, fig. 3C. Lamna appendiculata Roemer — Williston 1900B: pi. 31, fig. 49; pi. 32, fig. 3C. Lamna— Lane 1945: pi. 1, fig. 3C (= Williston 1900 A, pi. 14, fig. 3C). Isolated tooth. Upper Cretaceous, Turonian, PEagleford Group; Dallas Co., Dallas, Texas; collected by F. Hunston. Leptostyrax m^acrorhiza (Cope 1875) KUVP 55249 Undetemiined lamnid— Williston 1900A: pi. 12, fig. 2. 10 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Undetermined lamnid— Williston 1900B: pi. 30, fig. 2. Isolated tooth. Lower Cretaceous, Albian, Kiowa Fm.; Clark Co., Kansas; col- lected by S. W. Williston. KUVP ? Undetermined lamnid— Williston 1900A: pi. 12, fig. 3. Undetermined lamnid— Williston 1900B: pi. 30, fig. 3. Tooth; not relocated. Cretoxyrhinidae indet. KUVP ? Undeten-nined lamnid— Williston 1900A: pi. 12, fig. 1. Undetemiined lamnid— WilHston 1900B: pi. 30, fig. 1. Tooth; not relocated. Family Anacoracidae Casier 1947 Squalicorax Whitley 1939 Squalicorax curvatus (Williston 1900A) KUVP ? Corax curvata, n. sp.— Williston 1900A: p. 41, pi. 12, fig. 7,8; Corax curvatus WiHist. in caption of pi. 12. Corax curvatus, n. sp.— WiUiston 1900B: pi. 30, fig. 7,8. Tooth; not relocated. Squalicorax kaupi ( Agassiz 1843) KUVP 64501 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, fig, Tooth. KUVP 64502 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— WilHston 1900B: pi. 31, Tooth. KUVP 64503 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, Tooth. KUVP 64504 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, Tooth. KUVP 64505 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— WilHston 1900B: pi. 31, Tooth. KUVP 64506 Corax falcatus Agassiz— Williston 1900 A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, Tooth. KUVP 64507 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— WilHston 1900B: pi. 31, Tooth. KUVP 64508 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, Tooth. KUVP 64509 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31 Tooth. KUVP 64510 Corax falcatus Agassiz— Williston 1900A: pi. 13, Corax falcatus Agassiz— Williston 1900B: pi. 31, Tooth. fig- 1. 1. fig. 2. fig. 2. fig. 3. fig. 3. fig. 4. fig. 4. fig. 5. fig. 5. fig. 6. fig. 6. fig. 7. fig. 7. fig. 8. fig. 8. fig. 9. fi^g. 9. fig. 10. fig- 10. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 11 KUVP 64512 KUVP 64513 KUVP 64514 KUVP 64521 KUVP 64522 KUVP 64524 KUVP 64526 KUVP 64530 KUVP 64533 KUVP 64537 KUVP 64539 KUVP KUVP 64501 ? Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 12. Corax fakatus Agassiz— Williston 1900B: pi. 31, fig. 12. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 13. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 13. Tooth. Corax jalcatus Agassiz — Williston 1900A: pi. 13, fig. 14. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 14. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 21. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 21. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 22. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 22. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 24. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 24. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 26. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 26. Tooth. Corax jalcatus Agassiz— Williston 1900 A: pi. 13, fig. 30. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 30. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 33. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 33. Tooth. Corax jalcatus Agassiz— Williston 1900A: pi. 13, fig. 37. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 37. Tooth. Corax jalcatus Agassiz— Wilhston 1900A: pi. 13, fig. 39. Corax jalcatus Agassiz— Williston 1900B: pi. 31, fig. 39. Tooth. ■64539 Upper Cretaceous, Santonian, Niobrara Fm., Smoky Hill Chalk Mbr.; western Kansas; collected by E. P. West. Corax jalcatus Agassiz — Williston 1900A: pi. 14, fig. 1-lL. Corax jalcatus Agassiz— Williston 1900B: pi. 32, fig. 1-lL. Cora.r— Lane 1945: pi. 1, fig. 1-lL (=Winiston 1900A, pi. 14, fig. ML). Teeth of one individual; not relocated. KUVP 64511 KUVP 64515 KUVP 64516 Squalicorax Corax jalcatus Corax jalcatus Tooth. Corax jalcatus Corax jalcatus Tooth. Corax jalcatus Corax jalcatus Tooth. cf. kaupi (Agassiz 1843) Agassiz— Williston 1900A: pi. 13, fig. 11. Agassiz— Wilhston 1900B: pi. 31, fig. 11. Agassiz— Williston 1900A: pi. 13, fig. 15. Agassiz— Williston 1900B: pi. 31, fig. 15. Agassiz— Williston 1900A: pi. 13, fig. 16. Agassiz— Wilhston 1900B: pi. 31, fig. 16. 12 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 64517 KUVP 64518 KUVP 64519 KUVP 64520 KUVP 64523 KUVP 64525 KUVP 64527 KUVP 64528 KUVP 64529 KUVP 64531 KUVP 64532 KUVP 64534 KUVP 64535 KUVP 64536 KUVP 64538 KUVP 64540 Cor ax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Corax falcatus Corax falcatus Tooth. Agassiz— WilUston 1900A: pi. Agassiz— WilUston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz— Agassiz— Agassiz— Agassiz— Agassiz— Agassiz— Agassiz— Agassiz— Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- Agassiz- 13, fig. 17. 31, fig. 17. 13, fig. 18. 31, fig. 18. 13, fig. 19. 31, fig. 19. -Williston 1900A: pi. 13, fig. 20. -Williston 1900B: pi. 31, fig. 20. -Wilhston 1900A: pi. -Williston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. -Williston 1900 A: pi. -Williston 1900B: pi. -Wilhston 1900A: pi. -Williston 1900B: pi. -Williston 1900A: pi. -Wilhston 1900B: pi. -Williston 1900A: pi. -Wilhston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. -Williston 1900 A: pi -Wilhston 1900B: pi. -Wilhston 1900A: pi. -Williston 1900B: pi. -Williston 1900 A: pi -Williston 1900B: pi. -Williston 1900A: pi. -Williston 1900B: pi. 13, fig. 23. 31, fig. 23. 13, fig. 25. 31, fig. 25. 13, fig. 27. 31, fig. 27. 13, fig. 28. 31, fig. 28. 13, fig. 29. 31, fig. 29. 13, fig. 31. 31, fig. 31. 13, fig. 32. 31, fig. 32. 13, fig. 34. 31, fig. 34. . 13, fig. 35. 31, fig. 35. 13, fig. 36. 31, fig. 36. , 13, fig. 38. 31, fig. 38. 13, fig. 40. 31, fig. 40. KUVP 64511-64540 Upper Cretaceous, Santonian, Niobrar Chalk Mbr.; western Kansas; collected by E. a Fni., Smoky Hill P. West. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 13 Squalicorax sp. KUVP ? CoraxP— Williston 1900A: pi. 6, fig. 14. Corax.?— Williston 1900B: pi. 24, fig. 14. Tooth; not relocated. KUVP ? Corax.P— Williston 1900A: pi. 6, middle figure. Com.T.?— Williston 1900B: pi. 24, middle figure. Centi-um; not relocated. Family Ptychodontidae Hay 1902, inccrtae sedis Ptychodus Agassiz 1839 Ptychodiis anomjmus Williston 1900 KUVP 55252a,d Ptychodus anonymus, n. sp.— Williston 1900A: p. 32; pi. 11, fig. 5-8; syntypes. Ptychodus anonymus Willist.— Williston 1900B: pi. 29, fig. 5-8; syntypes. Teeth. KUVP 55252e,f Ptychodus anonymus, n. sp.— Williston 1900A: p. 32; pi. 11, fig. 17, 18; syntypes. Ptychodus anonymus Willist.— Williston 1900B: pi. 29, fig. 17, 18; syntypes. Teeth. KUVP 55252h,g Ptychodus anonymus, n. sp.— Williston, 1900A: pi. 11, fig. 21, 22; syntypes. Ptychodus anonymus Willist.— Wilhston 1900B: pi. 29, fig. 21,22; syntypes. Teeth. KUVP 55252i,k,l Ptychodus anonymus, n. sp.: Psyntypes, teeth. These teeth could be the specimens figured by Williston 1900A: pi. 11, fig. 16,20,24 or Williston 1900B: pi. 29, fig. 16,20,24, but the identity between specimens and figures cannot be assured. Upper Cretaceous (PGreenhorn Limestone); Walnut Creek, Rush or Ness Co., Kansas; collected by J. L. Shearer. PPtychodus jajiewayii (Cope 1874) Specimens figured by Williston 1900A: pi. 12, fig. 9-11 (= Wil- hston 1900B: pi. 30, fig. 9-11) could not be relocated. Existing teeth detemiined as P. janewayii do not match the figures. Ptychodus 7nartini Williston 1900 KUVP 55271 Ptychodus martini, n. sp.— Williston 1900A: p. 32; pi. 10; holotype. Ptychodus martini Willist.— Williston 1900B: pi. 28, holotype. Teeth arranged by author as associated dentition. Presently 102 teeth (Williston 1900 cites 110) arranged in a plaster block unlike the published figure. Upper Cretaceous, Coniacian, Niobrara Formation, Smoky Hill Chalk Member; Smoky Hill River, Trego Co., Kansas; collected by H. T. Martin 1894. This assignment to Coniacian is based on our knowledge that P. martini became extinct in the Western Interior before the Coniacian/ Santonian boundary. The assignment is supported by scarce outcrops of Lower Santonian rocks along Smoky Hill River in Trego County. 14 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Ptychodus mortoni Mantell 1842 KUVP 55269 Ptychodus mortoni Mantell— Williston 1900A: pi. 7. Ptychodus mortoiii Mantell — Williston 1900B: pi. 25. 80 teeth arranged in a plaster block different from the pub- lished figure. Upper Cretaceous, Coniacian or Santonian, Niobrara Formation, Smoky Hill Chalk Member; probably Trego Co., Kansas; collection of J. Savage. P. inortoni extends from the Coniacian into the Middle Santonian in Kansas, therefore a more specific assignment of stage is not possible. KUVP 55270 Ptychodus mortoni Mantell— Williston 1900A: pi. 8, fig. 1; pi. 9, fig. 1,2. Ptychodus mortoni Mantell— Wilhston 1900B: pi. 26, fig. 1; pi. 27, fig. 1,2. 539 teeth arranged in a plaster block different from the pub- lished figures. Upper Cretaceous, Coniacian, Niobrara Fomiation, Smoky Hill Chalk Member; near Castle Rock, but Trego Co., Kansas; collected by E. S. Rose. Ptychodus occidentalis Leidy 1868 KUVP 55259 Ptychochis occidentalis Leidy— Williston 1900A: pi. 11, fig. 4; pi. 12, fig. 13. Ptychodus occidentalis Leidy— Wilhston 1900B: pi. 29, fig. 4; pi. 30, fig. 13. 1 tooth. Upper Cretaceous, Cenomanian, Graneros Shale Formation; Ells- worth Co., Kansas. Ptychodus polygyrus Agassiz 1839 KUVP 55237 Ptychodus pohjgyrus Buckl— Williston 1900A: pi. 11, fig. 9; pi. 12, fig. 14. Ptychodus polygyrus Buckl.— WilHston 1900B: pi. 29, fig. 9; pi. 30, fig. 14. 1 tooth. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; Smoky Hill River, ?Gove Co., Kansas. Ptychodus sp. KUVP 55260 Ptychodus, sp. indct.— Williston 1900A: pi. 11, fig. 1. Ptychodus, sp. indet.— Wilhston 1900B: pi. 29, fig. 1. 1 tooth. KUVP 55256a-b Ptychodus, sp. indet.— Williston 1900A: pi. 11, fig. 2,3. Ptychodus, sp. indet.— Williston 1900B: pi. 29, fig. 2,3. 2 teeth. KUVP 55255a-d Ptychodus, sp.— Williston 1900 A: pi. 13, fig. 53. Ptychodus, sp.— Wilhston 1900B: pi. 31, fig. 53. Ptychodus— Lane 1945: pi. 2 (= Williston 1900A, pi. 13, fig. 53). 4 teeth. KUVP 55255, 55256, 55260 Upper Cretaceous, Turonian, Greenhorn Limestone Foi-mation; Salt Creek, Russell Co., Kansas; collected by J. L. Shearer. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 15 KUVP 55264 Ptijcliodus, spp.— Williston 1900A: pi. 11, fig. 19. Ttijchodus, spp.— Williston 1900B: pi. 29, fig. 19. 1 tooth. Upper Cretaceous; Western Kansas. Specimens to Williston 1900A: pi. 11, fig. 23,25 and 27 (=Wil- liston 1900B: pi. 29, fig. 23,25 and 27) could not be relocated. Ttijchodus lohipplei Marcou 1S58 KUVP 55254a-f Ptt/chodus whippleiji Marcou— Williston 1900A: pi. 11, fig. 10-15. Ptychodus whippJeyi Marcou— Williston 1900B: pi. 29, fig. 10-15. 6 teeth. Upper Cretaceous (PEagleford Group); Dallas, Texas; collected by F. Springer. KUVP 55253 Ptychodus, spp.— Williston 1900A: pi. 11, fig. 26. Ptychodus, spp.— Williston 1900B: pi. 29, fig. 26. 1 tooth. Upper Cretaceous; western Kansas. Elasmobranchii inc. sed. Listracanthus Newberry and Worthen 1870 Listracanthus eliasi Hibbard 1938 KUVP 884 Listracanthus eliasi n. sp.— Hibbard 1938: pp. 169-70; pi. 19, holotype, spine. KUVP 885, 886 Listracanthus e/Za^/— Hibbard 1938: pp. 169-170; paratype, spine, part and counterpart. Upper Pennsylvanian, Virgilian Stage, Wabaunsee Group, Cedar- vale Shale, few feet above Elmo coal; Carpenter's coal mine, 2 mi. W and 3 mi. S of Burlington Junction, Nodaway Co., Missouri; collected by M. K. Elias 1934. Listracanthus ? hystrix Newberry and Worthen 1870 KUVP 30604 probably L. hystrix— Chorn and Reavis 1978: text-fig. 2 (30604-1) + 3C (30604-2), densely packed spines (see below). Middle Pennsylvanian, Desmoinesian Stage, Mamiaton Group, Fort Scott Limestone, Little Osage Shale Member; SEJi, NEJl, NW)4, Sec. 19, T25S, R25E, Bourbon Co., Kansas; collected by E. A. Reavis 1974. Petrodus McCoy 1848 Petrodus sp. KUVP 30604 Petrodus— Choxn and Reavis 1978: text-fig. 3A,B (30604-5), denticle. Chorn and Reavis (1978) consider Petrodus denticles congeneric with Listracanthus spines. Middle Pennsylvanian, Desmoinesian Stage, Marmaton Group, Fort Scott Limestone, Little Osage Shale Member; SEK, NE)4, NWK, Sec. 19, T25S, R25E, Bourbon Co., Kansas; collected by E. A. Reavis 1974. Physonemus McCoy 1848 Physonemus mirahilis (St. John and Worthen 1875) KUVP 55274 Xystracanthus grandis Moore, n. sp.— Moore 1929: pp. 239-242; 16 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY pi. 40, fig. 1-3; holotype of X. grandis, spine [for synonymy see Zidek'l977, and Chorn and Frailey 1978]. Upper Pennsylvanian, Cisco Group, Graham Fomiation, Wayland Shale Member; 5 miles west of Eastland, Eastland Co., Texas; collected by R. C. Moore. KUVP 24011 Physonemus mirabilis—Chom and Frailey 1978: text-fig. lA-D; dorsal spine. Upper Pennsylvanian, Missourian Stage, Coffeyville Formation; approximately 7.5 km north of Coffeyville, Montgomery Co., Kansas; collected by B. Foreman 1975. KUVP 61 KUVP 17851 KUVP 17852 KUVP 17853 KUVP 17854 KUVP 17855 KUVP 17856 KUVP 17857 KUVP 17858 KUVP 17859 KUVP 17860 KUVP 17863 Physonemus striatus ( Moore 1929 ) Xystracanthus striatus Moore n. sp. — Moore 1929: 40, fig. 4,5; holotype, spine. Pennsylvanian; eastern Kansas. p. 242-243; pi. Enterospira Heteropolar "spiral coprolite" — Williams 1972: pi. 1; fig. 1; com- plete specimen. Heteropolar "spiral coprolite" — Wilhams 1972: pi. 1; fig. 2; com- plete specimen. Heteropolar "spiral coprolite"— Wilhams 1972: pi. 1; fig. 3; broken specimen. Heteropolar "spiral coprolite"— Williams 1972: pi. 1; fig. 4; broken specimen. Heteropolar "spiral coprolite" — Williams 1972: plete specimen. Heteropolar "spiral coprolite" — Williams 1972: plete specimen. Heteropolar "spiral coprolite" — Williams 1972: plete specimen. Heteropolar "spiral coprolite" — Williams 1972: plete specimen. Heteropolar "spiral coprohte" — Williams 1972: section. Heteropolar "spiral coprolite"— Williams 1972: pi. 1; fig. 10; longi- tudinal section. —Williams 1972: pi. 4, fig. 3-4; pi. 5, fig. 3; cross section 1. — Williams 1972: pi. 5, fig. 2; cross section 5. — Williams 1972: pi. 5, fig. 4; cross section 4. — Williams 1972: pi. 7, fig. 4; cross section 7. —Williams 1972: pi. 8, fig. 1; cross section 10. —Williams 1972: pi. 8, fig. 2; cross section 2. —Williams 1972: pi. 8, fig. 3; cross section 8. —Williams 1972: pi. 8, fig. 4; cross section 12. —Williams 1972: text-fig. 10-1; cross section 6. KUVP 17866 —Williams 1972: pi. 6, fig. 1-2; cross section 10. KUVP 17867 —Williams 1972: pi. 6, fig. 3-4; cross section 2. —Williams 1972: pi. 7, fig. 1-2; cross section 4. KUVP 17868 —Williams 1972: pi. 7, fig. 3; cross section 2. Williams (1972) regards the "spiral coprolites" as enterospira of Xenacanthus. pi. 1; fig. 5; com- pl. 1; fig. 6; com- pl. 1; fig. 7; com- pl. 1; fig. 8; com- pl. 1; fig. 9; cross TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 17 KUVP 17862 KUVP 17864 KUVP 43074 KUVP 43073 KUVP 43071 KUVP 43072 Lower Pennian, Wolfcampian, Matfield Shale, lower 3 meters of the Wymore Shale Member; western side of roadcut 1.7 km south of junction Kansas Highway 177 with 113 in NE)4, NW)^, Sec. 35, T9S, R7E, Riley County, Kansas; collected by M. E. Williams 1969. —Williams 1972: pi. 2, fig. 2; section 8. —Williams 1972: pi. 4, fig. 1-2; section 6. The enterospira probably belongs to Xenacanthus as do the above cited specimens judging from "striking resemblance" (Williams 1972, p. 13). Lower Permian, Wolfcampian; near Wichita Falls, Texas; collected by Kansas Geological Survey. —Williams 1972: pi. 5, fig. 1, cross section 10. —Williams 1972: text-fig. 10-2; cross section 13. The enterospira probably belongs to Xenacanthus. Lower Permian, Wolfcampian, Wichita Group, Putnam Fomiation; near Bellevue, Clay Co., Texas; donated by A. S. Romer. Enterospira — Stewart, J. D. 1978: text-fig. 4-1; complete specimen. Enterospira — Stewart, J. D. 1978: text-fig. 4-2; complete specimen. Enterospira — Stewart, J. D. 1978: text-fig. 4-3; broken specimen. Stewart, J. D. 1978 Enterospira section. KUVP 43068 Enterospira— Stewart, J. D. 1978: section. Enterospira— Stewart, J. D. 1979: section. KUVP 43070 Enterospira— Stewart, J. D. 1978: section. Enterospira— Stewart, J. D. 1978: section. KUVP 43069 Enterospira— Stewart, J. D. 1978: text-fig. 4-4; mounted cross text-fig. 4-7; mounted cross text-fig. 5-3,4; mounted cross text-fig. 5-1,2; mounted cross text-fig. 6-2,3,4; mounted cross text-fig. 6-1; mounted cross section. Assigned by J. D. Stewart 1978 to "small selachians previously undetected in the fomiation." Upper Cretaceous, Coniacian, Niobrara Fomiation, Smoky Hill Chalk Member; E)2, Sec. 27, T14S, R26W, Gove County, Kansas; collected by J. D. Stewart 1976. KUVP 17861 Spiral valve of Squalus acoMf/u'as- Williams, 1972: pi. 2, fig. 1; cross section 1. —Williams, 1972: pi. 3, fig. 1; cross section 2. — WilHams, 1972: pi. 3, fig. 2,3,4; cross section 3. Embryo. Recent; locality unknown. CLASS OSTEICHTHYES Howes 1894 Subclass Actinopteiygii Klein 1885 Superorder Chondrostei Miiller 1846 Order Palaeoniscifomies Hay 1929 Family Palaeoniscidae Owen 1860 Cocconiscus White and Moy-Thomas 1940 Cocconiscus baldwini Poplin 1974 18 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 55598 (12487-11-31)* Cocconiscus haUwini n.g., n. sp.— Poplin, 1974: p. 118; text-fig. 34; pi. 32; holotype, partial skull. KUVP 55600 (12688-12-12) Cocconiscus haldwini n.g., n. sp.— Poplin, 1974: text-fig. 47; pi. 33, fig. 4,5; pi. 40, fig. 2; paratype, upper and lower jaw. KUVP 55599 (12486-10-9) Cocconiscus boJdwini n.g., n. sp.— Poplin, 1974: pi. 33, fig. 1,2.3; paratype, partial skull. KUVP 55603 (12698-22-69) Cocconiscus haldwini n.g., n. sp.— Poplin, 1974: pi. 34, fig. 1,2; cleithiimi. KUVP 55601 (12695-19-5) Cocconiscus haldwini n.g., n. sp.— Poplin, 1974: text-fig. 39; pi. 34, fig. 3,4; maxillary. KUVP 55602 (12698-22-9) Cocconiscus haldwini n.g., n. sp.— Poplin, 1974: text-fig. 46; pi. 40, fig. 1; lower jaw. Upper Pennsylvanian, Douglas Group, Lawrence Fonnation, Has- kell Limestone Member; Douglas County, Kansas (precise locality for every specimen see ) . Kansasiella Poplin 1975 (=Kansasia Poplin 1974) Kansasiella eat oni (Poplin 1974) (Type-species) KUVP 22070 £/o/i/c/jt/i{/.s?— Miller and Swineford 1957: pi. 2, figs. A,B; endo- cranium. Miller and Swineford 1957: specimens to pi. 1, figs. A-D, pi. 2, fig. C, pi. 3, fig. A,B,D could not be identified in the collection. KUVP 55611 (12992-31-8) Kansasia eatoni n.g., n. sp.— Poplin, 1974: p. 23; pi. 10; pi. 12; pi. 13; pi. 15, fig. 1; pi. 27, fig. 1,3; holotype endocranium. KUVP 55643 (12698-22-76) Kansasia eatoni n.g., n. sp.— Poplin, 1974: text-fig. 4; pi. 1, fig. 2; pi. 4, fig. 3; paratype, partial skull. KUVP 55641 (12995-33-8) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 1,2; paratype, maxillary and palatoquadrate. KUVP 55642 (12985-26-26) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 5; paratype, cleithrum. KUVP 55612 (12992-31-9) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 2,4; paratype, occipital part of endocranium. KUVP 55613 (12995-33-20) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 2,3; pi. 30, fig. 1,2; paratype, endocranium. KUVP 55614 (12684a-B-6) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 27, fig. 2; paratype, endocranium. KUVP 55616 (12695-19-106) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 3; pi. 19; pi. 28, fig. 1; paratype, endocranium. KUVP 55615 (12695-19-62) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 3,4; paratype, endocranium. KUVP 55649 (12696-20-1) Kansasia eatoni n.g., n. sp.— Poplin, 1974: text-fig. 5; pi. 1, fig. 1; frontal. KUVP 55645 (12692-16-12) Kansasia eatoni n.g., n. sp fig. 1; preoperculum. KUVP 55644 (12692-16-6) Kansasia eatoni n.g., n. fig. 2; preoperculum. ** Numbers in ( ) are the published numbers of the speeimens. pl. 2, pl. 4, pl. 9, pl. 23, pl. 14; pl. 15, pl. 21, xt- Poplin, 1974: pl. 3, .sp.— Poplin, 1974: pl. 3, TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 19 KUVP 55651 KUVP 55648 KUVP 55646 KUVP 55647 KUVP 55650 KUVP 55634 KUVP 55635 KUVP 55640 KUVP 55622 KUVP 55626 KUVP 55618 KUVP 55633 KUVP 55652 KUVP 55638 KUVP 55636 KUVP 55631 KUVP 55632 KUVP 55617 KUVP 55627 KUVP 55624 KUVP 55619 KUVP 55639 KUVP 55637 KUVP 55620 sp.— Poplin, 1974: sp.— Poplin, 1974: sp.— Poplin, 1974: pi. 3, pi. 3, pl. 4, pl. 4, pl. 4, pl. 5, 12987-26-45) Kansasia eatoni n.g., n. fig. 3,4; maxillaiy. 12495-19-71) Kansasia eatoni n.g., n. fig. 4; palatoquadrate. 12694-18-2) Kansasia eatoni n.g., n. fig. 1; parasphcnoid. 12695-19-42) Kansasia eatoni n.g., n. sp. — Poplin, 1974: fig. 2; parasphcnoid. 12985-26-8) Kansasia eatoni n.g., n. sp. — Poplin, 1974: fig. 4; cleithrum. 12988-27-64) Kansasia eatoni n.g., n. sp.— Poplin, 1974: sections 157, 237, 313, 367; pl. 6, sections 441, 519, 579, 645 (error 12988-22-64) pl. 7, sections 665, 701, 729, 749; pl. 8, sections 825, 843, 855, 877 (error on page 21, 12988-67-64); endocranium. 12992-31-35) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 3; endocranium. 13357-35-26) Kansasia eatoni n.g., n. sp.— Poplin, 1974: endocranium. 12695-19-58) Kansasia eatoni n.g., n. sp. — Poplin, 1974: fig. 2; endocranium. 12697-D-3) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 1 (error 12697-D-8); pl. 29, fig. 1; endocranium. 12690-14-1) Kansasia eatoni n.g., n. sp. — Poplin, 1974: fig. 2; endocranium. 12988-27-32) Kansasia eatoni n.g., n. sp.— Poplin, 1974: endocranium. 13357-35-22) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 1,2; pl. 29, fig. 2; endocranium. 13361-39-7) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 3; endocranium. 12996-32-4) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 4; endocranium. 12987-26-86) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 1; endocranium. 12987-26-87) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 2; endocranium. 12687-11-10) Kansasia eatoni n.g., n. sp.— Poplin, 1974: fig. 3; endocranium. 12698-22-13) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 21, fig. 1,2; pl. 23, fig. 1; endocranium. 12695-19-84) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 22, fig. 1; pl. 30, fig. 3; endocranium. 12491-15-3) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 22, fig. 2; endocranium. Gl [error G-11] ) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 24; endocranium. 12995-33-17) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 25, fig. 1; endocranium. 12691-15-34) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pl. 25, fig. 2; endocranium. pl . 9, pl. 11, pl. 15, pl. 16, pl. 16, pl. 17; pl. 18, pl. 18, pl. 18, pl. 20, pl. 20, pl. 20, 20 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 55629 (12987-26-72) Kansasia eatoni n.g, n. sp.— Poplin, 1974: pi. 25, fig. 3; pi. 2(8, fig. 2; endocranium. KUVP 55621 (12695-19-60) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 26, fig. 1; endocranium. KUVP 55625 (12695-19-112) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 26, fig. 2; endocranium. KUVP 55630 (12987-26-81) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 26, fig. 3; endocranium. KUVP 55628 (12698-22-44) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 26, fig. 4, pi. 28, fig. 3; endocranium. KUVP 55623 (12488-12-3) Kansasia eatoni n.g., n. sp.— Poplin, 1974: pi. 29, fig. 3; endocranium. Upper Pennsylvanian, Douglas Group, Lawrence Fomiation, Has- kell Limestone Member; Douglas County, Kansas (precise locality for every specimen see ) . Palaeoniscid indet. KUVP 56232 (12987-26-76) Palaeoniscid indet.— Poplin 1974: cf. text-fig. 46B; pi. 40, fig. 3; mandible. KUVP 56241 (12492-16-4) Palaeoniscid indet.— Poplin 1974: text-fig. 48; man- dible. KUVP 56254 (12987-26-39) Palaeoniscid indet.— Poplin 1974: pi. 35, fig. 1 right frontal. KUVP 56115 (12985-24-33) Palaeoniscid indet.— Poplin 1974: pi. 35, fig. 2 right ceratohyal. KUVP 56243 (12985-24-15) Palaeoniscid indet.— Poplin 1974: pi. 35, fig. 3 right palatoquadrate. KUVP 56246 (12986-25-27) Palaeoniscid indet.— PopHn 1974: pi. 35, fig. 4 right palatoquadrate. KUVP 56249 (12987-26-36) Palaeoniscid indet.— Poplin 1974: pi. 35, fig. 5 left palatoquadrate. KUVP 56240 (13358-36-5) Palaeoniscid indet.— Poplin 1974: pi. 36, fig. 1,2 left mandible. KUVP 56227 (12987-26-16) Palaeoniscid indet.— Poplin 1974: pi. 36, fig. 3,4 left mandible. KUVP 56255 (12988-27-61) Palaeoniscid indet.— Poplin 1974: pi. 37, fig. 1 hyomandibula. KUVP 56234 (12988-27-53) Palaeoniscid indet.— Poplin 1974: pi. 37, fig. 4 mandible. KUVP 56257 (12994-32-8) Palaeoniscid indet.— Poplin 1974: pi. 39, fig. 1 scales. KUVP 56256 (12484-B-3) Palaeoniscid indet— Poplin 1974: pi. 39, fig. 2; scale KUVP 56158 (12994-32-10) Palaeoniscid indet— Pophn 1974: pi. 39, fig. 3 scale. KUVP 56259 (12995-33-21) Palaeoniscid indet.— Poplin 1974: pi. 39, fig. 4,5 scale. Upper Pennsylvanian, Douglas Group, Lawrence Formation, Has- kell Limestone Member; Douglas County, Kansas (precise locality for every specimen, see ). TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 21 Palaeoniscoid B KUVP 56374 Palaeoniscoid B— Poplin 1981: text-fig. lA-C, 3/4A; natural cast of endocranial cavity. KUVP 56371 Palaeoniscoid B— Poplin 1981: text-fig. 3/4B; natural cast of endo- cranial cavity. KUVP 56373 Palaeoniscoid B— Poplin 1981: text-fig. 3/ 4C; natinal cast of endo- cranial cavity. KUVP 56372 Palaeoniscoid B— Poplin 1981: text-fig. 3/4D; natural cast of endo- cranial cavity. Upper Pennsylvanian, Douglas Group, Lawrence Formation, Has- kell Limestone Member; Douglas County, Kansas (precise locality for every specimen, see ). Localities for figured specimens of Cocconiscus haldwini, Kan- sasiella eatoni, Palaeoniscoid B, and palaeoniscids indet.: 55617 Stiu-dy Fanu, 2 mi. NE of Baldwin, Douglas County, Kansas. 3)2 mi. East of Vinland, Douglas Co., Kansas. 55623 NW)^, Sec. 27, T14S, B20E, Douglas Co., Kansas. 55615, 55616, 55621, 55622, 55624, 55625, 55647, 55648 Palmyra Township, N and NE of Baldwin, Douglas Co., Kansas. 55603, 55627, 55628, 55643, 56371, 56372 NE)^, Sec. 21, T14S, R20E, Douglas Co., Kansas. 55612, 55635, 56374 Week's Fanu, N of Baldwin, Douglas County, Kansas. 55637, 55641, 56259 NEJi, Sec. 27, T14S, R21E, Douglas Co., Kansas. 56256 East of Oak Hill Cemetery, Lawrence, Douglas Co., Kansas. 55620, 55639, 56240 Douglas Co., Kansas. McQueen Fanu, NW of Baldwin, Douglas Co., Kansas. Road between Sec. 25 and 36, T13S, R19E, Douglas Co., Kansas. 55630, 55631, 55632, 56227, 56232, 56249, 56254 NWK, Sec. 26, T14S, R20E, Douglas Co., Kansas. 55634, 56234, 56255 Sec. 16, T14S, R20E, Douglas Co., Kansas. 56257, 56258 SWK, Sec. 13, T14S, R21E, Douglas Co., Kansas. Sec. 7, T14S, R21E, Douglas Co., Kansas. 55652 SW)1, Sec. 8, T14S, R21E, Douglas Co., Kansas. 55650, 56115, 56243 SE)1, Sec. 16, T14S, R20E, Douglas Co., Kansas. 55645, 56241 SE)4, Sec. 26, T14S, R20E, Douglas Co., Kansas. NW)4, Sec. 21, T14S, R20E, NE Queen Fann, Douglas Co., Kansas. Kelly Fai-m, NE of Baldwin, Douglas Co., Kansas. Al Andrew's Farm, East Road, Baldwin, Douglas Co., Kansas. NE of Baldwin, Douglas Co., Kansas. Collector: A. Bridwell, Baldwin, Kansas, in the years 1920-1940 and earlier; few specimens as early as 1893. KUVP 55598, KUVP 55599 KUVP 55600, KUVP 55601, KUVP 55602, KUVP 55611, KUVP 55613, KUVP 55614, KUVP 55619, KUVP 55618 KUVP 55626 KUVP 55629, KUVP 55633, KUVP 55636, KUVP 55638 KUVP 55640, KUVP 55642, KUVP 55644, KUVP 55646 KUVP 55649 KUVP 56246 KUVP 56373 Superorder Neopterygii Regan 1923 Division Pycnodontiformes Berg 1940 The pycnodonts are clearly ncopterygians (Patterson 1973), but they cannot be classified within any of the divisions of neoptery- 22 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY gians. Therefore, they are here provisionally elevated to Division rank until shown otherwise. Family Pycnodontidae Owen 1846 Coelodus Meckel 1856 Coelodtis hrownii Cope 1894 KUVP 55420 Coelodus brownii Cope— Williston 1900A: pi. 6, fig. 12; holotype. Coelodus brownii Cope— Williston 1900B: pi. 24, fig. 12; holotype. Left prearticular. Lower Cretaceous, Albian, Kiowa Fonnation; near Belvidere, Kiowa Co., Kansas; collected by N. C. Gould. Coelodus stantoni Williston 1900 KUVP 887 Coelodus stantoni Wilhst.— Williston 1900A: p. 28; pi. 6, fig. 11; pi. 8, fig. 6; holotype. Coelodus stantoni Willist.— Williston 1900B: pi. 24, fig. 11; pi. 26, fig. 6; holotype. Right prearticular. Lower Cretaceous, Albian, Kiowa Fomiation; Kiowa Co., Kansas. Coelodus streckeri Hibbard 1939 KUVP 946 Coelodus streckeri, sp. nov.— Hibbard 1939: p. 373, pi. 41, fig. 1, 2; holotype. Fossil Pycnodont teeth— Lane 1945: pi. 3 (=Hibbard 1939, pi. 41); holotype. Vomer. Upper Cretaceous, Turonian, Carlile Shale Fm., Faii-port Chalk Mbr.; SEK, Sec. 19 or NEK, Sec. 30, TllS, R14W, 2'A mi. south and M mi. east of Paradise, Russell Co., Kansas; collected by Ernest Streaker 1938. Micropycnodon Hibbard and Graffham 1945 Syn. Pycnomicrodon Hibbard and Gralfliam 1941 Micropycnodon kansasensis (Hibbard and Graffham 1941) (Type-species) KUVP 1019 Pycnomicrodon kansasensis sp. nov.— Hibbard and Graffham 1941: p. 72, text-fig. 1; pi. 1, fig. 1-4; holotype. Micropycnodon kansasensis (Hibbard and Graffham) — Hibbard and Graffham 1945: p. 404 ( new name) . Micropycnodon kansasensis (Hibbard and Graffham) — Dunkle and Hibbard 1946: pi. 9, fig. 1-4; holotype. Anterior part of head. Upper Cretaceous, Coniacian, Niobrara Fm., probably Smoky Hill Chalk Mbr. (published as Fort Hays Limestone Mbr.); Sec. 32, T7S, R19W, 1 mi. W of old Webster townsite, Rooks Co., Kansas; collected by Allen Graffham 1939. KUVP 7030 Micropycnodon kamasensis (Hibbard and Graffliam )— Dunkle and Hibbard 1946: text-fig. 1-2, pi. 7+8; head and trunk. Upper Cretaceous, Coniacian, Niobrara Fm., probably Smoky Hill Chalk Mbr. (published as Fort Hays Limestone Mbr.); Sec. 21, T13S, R21W, 5 mi. south and 4J2 mi. east of Ogallah, Trego Co., Kansas; collected by G. F. Sternberg ca. 1935. TYPE AND FIGURED FOSSIL FISHES AT UNIVERSITY OF KANSAS 23 Pycnomicrodon Hibl)ard and GrafFliam 1941 see Micropycnodon Ilibbard and Graffham 1945 Division Ginglymodi Cope 1