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THE  GBE  AN  W 

A newsletter  to  promote  communication  among  research  scientists 
concerned  with  the  systematics  of  the  Leguminosae/ Fabaceae. 

Editor:  Charles  R. Gunn,  Plant  Taxonomy  Laboratory,  BARC-EAST,  USDA,  Beltsville,  MD  20705 
Associate  Editor:  James  Lackey,  Botany,  NHB166,  Smithsonian  Institution,  Washington,  D.C.  20560 


7 " - 

• I 


4c. 


A 


N°.  15  May  1982 

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EDITORIAL 


The  Plant  Taxonomy  Laboratory  and  the  U.S.  National  Seed  Herbarium  are 
moving.  Please  use  our  new  address  for  all  Bean  Bag  correspondence: 

Lydia  R.  Poole,  Building  265,  BARC-EAST,  Beltsville,  MD  20705  USA. 

Notes  about  the  Bean  Bag:  The  Recent  Literature  column  will  only  report 

papers  of  general  taxonomic  interest  that  have  been  published.  Incomplete 
citations  will  not  be  published.  A complete  citation  includes  author(s), 
date,  title,  where  published.  Please  use  the  style  in  this  issue  of  the  Bean 
Bag.  Please  print  clearly.  Several  items  from  readers  could  not  be  read  and 
thus  could  not  be  put  in  this  issue.  We  appreciate  the  personal  comments  from 
our  Readers.  They  are  pleasant  to  come  across  during  the  rush  of  publishing. 

This  issue  of  the  Bean  Bag  was  data  banked  in  a PRIME  computer  and 
printed  by  a WANG  word  processer.  The  mailing  labels  also  were  issued  in  a 
similar  manner. 


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FLORA  NE OTROP I CA-LEGUMI NOS AE 
Roger  H.  Polhill 

The  time  has  come  to  draw  together  the  scattered  knowledge  and  vast 
research  potential  of  the  La t in-American  legume  flora  into  the  more  formalized 
format  of  FLORA  NEOTROPICA.  The  publication  of  Flora  Neotropica  Monographs 
began  in  1968  with  the  treatment  of  Swartzia  by  COWAN,  but  no  further  legume 
accounts  have  been  submitted. 

The  Organization  for  Flora  Neotropica  has  appointed  co-ordinators  for 
each  of  the  three  subfamilies,  and  anyone  interested  in  participating  is 
encouraged  to  make  contact  with  them: 


Caesalpinioideae : 


Kai  Larsen,  Botanical  Institute, 
University  of  Aarhus,  Nordlandsvej  68 
DK-8240  Risskov,  Denmark 


Mimosoideae:  Enrique  Forero,  Institute  de  Ciencias  Naturales 

Universidad  National 
Bogota,  Colombia 

Papilionoideae:  Roger  M.  Polhill,  Royal  Botanic  Gardens, 

Kew,  Richmond,  Surrey,  TW9  3AE  England 

The  co-ordinators  will  try  to  devise  a balance,  so  that  individual 
interests  in  single  genera  can  be  fostered,  and  so  that  collaborative  work  can 
be  devised  to  write  whole  tribes  with  a proper  consideration  of  generic 
structure  and  delimitation.  Clearly  both  considerations  are  effectively  met 
by  the  aefive  participation  of  more  people,  and  the  Organization  looks  to 
readers  of  The  Bean  Bag  for  practical  help  and  suggestions.  The  progress  of 
individual  projects  will  be  noted  in  these  pages  from  time  to  time,  and  the 
co-ordinators  propose  to  establish  a panel  of  advisers  from  among  those 
workers  most  actively  involved  with  neotropical  legumes. 
******************************************************************************* 

CYTOLOGICAL  GAPS 


You  are  encouraged  to  help  fill  the  cytological  gaps  in  GOLDBLATT 8 s 
survey  of  legume  genera.  Listed  below  are  the  genera  from  section  III  Gaps  in 
the  Record  (pp.  429-430)  from  GOLDBLATT 8 s chapter  in  Advances  in  Legume 
Systematics.  Please  review  this  list  and  either  send  GOLDBLATT  seed  samples 
with  herbarium  vouchers  or  provide  GOLDBLATT  with  published  chromosome  counts 
or  send  him  the  citation  if  found  in  an  obscure  publication.  All 
contributions  will  be  acknowledged.  (Editor's  note:  We  have  sent  GOLDBLATT 

seeds  of  Pachecoa.)  Send  all  material  to  Peter  Goldblatt,  Missouri  Botanical 
Garden,  P.0.  Box  299,  St.  Louis,  MO  63166,  USA.  (Telephone:  314-577-6100) 


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Aenictophyton,  Affonsea,  Afgekia,  Aganope,  Aldina,  Alepidocalyx,  Alexa, 
Alistilus,  Androcalymma,  Antheroporum,  Apaloxylon,  Aphanocalyx,  Apopbanesia , 
Arapatiella,  Archidendron,  Arcoa,  Arthrocarpum,  Arthroclianthus , Aubrevillea, 
Augouardia,  Austrodolichos . 

Balsamocarpon,  Baphiopsis,  Batesia,  Bathiaea,  Baudouinia,  Baukea, 

Behaimia,  Bembicidium,  Benedictella , Bocoa,  Bracteolanthus , Breniera, 
Browneopsis,  Bryaspis,  Buchenroedera , Burkilliodendron. 

Callistachys , Campsiandra,  Candolleodendron,  Carrissoa,  Cascaronia, 
Cedrelings,  Cenostigma,  Chadsia,  Chaetocalyx,  Chidlowia,  Chrysoscias, 
Clathrotropis,  Cleobulia,  Clitoriopsis , Cochlianthus , Coelidium, 

Condylostylis , Conzattia,  Coroya,  Coursetia,  Craibia,  Cranocarpus , 
Craspedolobium,  Cratylia,  Cruddasia,  Cryptosepalum,  Cyclocarpa,  Cyliocodiscus , 
Cymbosema,  Cynometra,  Cytisopsis. 

Dahlstedtia,  Dalbergiella , Dalhousiea,  Decorsea,  Dewevrea , Dicorynia, 
Dicraeopetalum,  Dicymbe,  Didelotia,  Didymopelta , Diphyllarium,  Diplotropis, 
Diptychandra , Discolobiutn,  Disynstemon,  Dolichopsis,  Droogmansia,  Dunbaria, 
Duparquetia,  Dysolobium. 

Elephantorrhiza , Eligmocarpus , Elizabetha,  Endertia,  Englerodendron, 
Eperua,  Eremosparton,  Erichsenia,  Etaballia,  Eurypetalum,  Exostyles. 

Fiebrigiella,  Fillaeopsis,  Fissicalyx,  Fordia. 

Geissaspis,  Gilletiodendron,  Goniorrhachis , Gonocytisus, 
Gossweilerodendron. 

Hallia,  Hannnatolobium,  Haplormosia,  Harpalyce,  Herpyza,  Hesperolaburnum, 
Hesperotharnnus , Heterostemon,  Humboldtia,  Humularia,  Hylodendron,  Hymenolobium. 

Indopiptadenia,  Isoberlinia,  Isomacrolobium. 

Jacqueshuberia . 

Kalappia,  Kaoue,  Kingiodendron,  Kostyczewa,  Koompassia. 

Lebruniodendron,  Lecointea,  Lemuropisutn,  Lennea,  Leonardoxa,  Leptodesmia, 
Leucomphalos , Leucostegane , Leycephyllum,  Librevillea,  Loesenera, 
Lophocarpinia,  Lovanafia,  Luetzelburgia , Luzonia,  Lyauteya. 

Macroberlinia , Macrolobium,  Macropsychanthus , Maniltoa,  Margaritolobium, 
Martiodendron,  Mastersia,  Mecopus,  Melanoxylon,  Melliniella,  Mendoravia, 
Michelsonia , Microberlinia , Mildbraediodendron,  Minkelersia,  Moldenhauera 
Monopetalanthus , Monopteryx,  Monoschisma,  Mora,  Muellera,  Myrocarpus. 

Neochevalierodendron,  Neocolletia,  Neocracca,  Neodielsia,  Neodunnia, 
Neoharmsia,  Neorudolphia , Nephrodesmus , Nesphostylis , Nissolia,  Notodon. 

Oddoniodendron,  Oreophysa,  Ormocatpopsis , Ostryocarpus , Otoptera, 

Oxystigma . 

Pachecoa,  Pachyelasma,  Padbruggea,  Paloue,  Paloveopsis,  Panurea, 
Paralbizzia,  Paramachaerium,  Paratephrosia,  Parryella,  Peekelia, 
Pellegriniodendron,  Peltogyne,  Petaladenium,  Phylacium,  Phylloxylon, 
Piptadeniopsis , Plagiocarpus , Plagiosiphon,  Platycelyphium,  Platycyamus, 
Platypodium,  Platysepalum,  Podocytisus,  Podopetalum,  Poecilanthe,  Poeppigia, 
Poissonia,  Poitea,  Polystemonanthus , Prioria,  Prosopidastrum,  Pseudeminia , 
Pseudoeriosema , Pseudoentada , Pseudomacrolobiura,  Pseudoprosopis , 

Pseudosindora  Psoralea  (sens.  str.). 

Ramirezella , Ramorinoa,  Recordoxylon,  Rhodopis,  Rhynchotropis , 

Riedeliella,  Robyns iophy ton. , 


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Sakoanala,  Salweenia,  Sartorxa,  Sauvallella,  Schef f lerodendron, 
Schranckiastrum,  Scorodophloeus , Sellocharis,  Sindoropsis,  Sinodolichos , 
Soemmeringia,  Spartidium,  Spathionema,  Spatholobus,  Spirotropis, 

Stachyothyrsus , Stahlias  Stenodrepanum,  Stracheya,  Streblorrhiza , Stuhlmannia, 
Sweetia,  Sympetalandra . 

Talbotiella,  Tessmannia,  Tetraberlinia , Tetrapterocarpon,  Teyleria, 
Thylacanthus , Trif idacanthus . 

Uleanthus,  Umtiza,  Urariopsis,  Uribea. 

Vatairea,  Vataireopsis , Vatovaea,  Vavilovias  Vermifrux,  Vouacapoua. 
Wagatea,  Weberbauerella,  Whitfordiodendron. 

Xerocladia* 

Yucaratonia . 

Zenia,  Zenkerellas  Zollernias  Zuccagnia. 

**********★************************************************************■***■****'* 

PIGEONPEA  NEWSLETTER 

The  INTERNATIONAL  PIGEONPEA  NEWSLETTER  has  been  issued.  Number  1 (1981) 
may  be  obtained  from  Donald  G.  Faris,  Pulse  Improvement  Program,  ICRISAT, 
Patancheru  P.0. , Andhra  Pradesh  502  324,  India.  Also  available  are  separate 
publication?  on  pigeonpea  and  chickpea. 

NITROGEN  FIXING  TREE  ASSOCIATION 

The  Nitrogen  Fixing  Tree  Association  (NFTA)  was  established  in  1981 
to  promote  international  research  and  development,  communication,  and 
utilization  of  nitrogen-fixing  trees  for  fuel,  fertilizer,  forage,  feed, 
forests,  and  other  products.  Annual  dues  of  U.S  $5.00  should  be  sent  to  NFTA, 
P.0.  Box  680,  Waimanalo,  Hawaii  96795  USA. 

******************************************************************************* 
GLEANINGS  FROM  THE  DATA  COLLECTION  SHEETS 

ALBUQUERQUE  is  studying  Cajanus  cajan  and  would  welcome  legume  reprints. 

ANDERSON  has  completed  studies  of  Acacia  (mainly  Australian)  and  Prosopis 
(mainly  American)  exudates  and  is  starting  a similar  study  of  African 
Acacia  exudates.  Recently  returned  from  field  study  of  Acacia  plantations 
in  the  Sudan.  Welcomes  water-soluble  gum  exudates  from  any  legume. 

AQUINO  is  a new  Reader. 

ARAGAO  DE  OLIVEIRA  is  a new  Reader  who  has  completed  a study  of  Desmodium  for 
the  state  of  Rio  Grande  do  Sul,  Brazil  and  is  starting  a study  of 
Aeschynomene  for  same  state.  Needs  literature  on  the  Hedysareae. 


-5- 


ARORA  is  working  on  the  Asiatic  Vigna  species  with  E.  R.  Nayoz,  K.  L.  Mehra, 
and  CHANDEL.  Recommends  study  of  Cajanus,  Atylosia,  Dunbaria,  and 
Dolichos.  Will  be  collecting  wild  Vigna  species  in  India.  Plans  to  study 
Mucuna,  especially  phytogeography  of  Asiatic  species.  His  name  and 
address  were  omitted  from  the  BB-14  Directory. 

ARROYO.  See  KEIGHERY. 

BAUDOIN . See  MARECHAL. 

BERMUDEZ  has  completed  a study  of  legumes  of  forest  value  in  Colombia. 

CARRILLO-FUENTES  is  a new  reader  and  has  finished  a study  of  Peruvian  species 
of  Phaseolus  and  will  be  working  on  the  Faboideae  of  the  Valle  del  Rimac. 
Needs  legume  sheets  and  seeds.  Will  be  collecting  Phaseolus  along  the 
coast  and  mountains  of  Peru.  Would  like  to  obtain  selected  descriptions 
of  species  of  Phaseolus. 

CHANDEL.  See  ARORA. 

CHEN  has  completed  the  Bauhinia  for  Flora  Republicae  Popularis  Sinicae  and 
has  started  study  of  Dalbergia  for  flora. 

CLEMONS  is  a new  Reader. 

COMBES,  who  will  be  working  with  LABEYRIE,  is  a new  Reader. 

CORBY  is  now  an  Honorary  Research  Fellow  and  continues  to  write  about  legume 
root  nodules. 

CUBERO  has  contributed  to  two  books  and  recommends  the  study  of  free 

amino-acids  in  Vicia.  Needs  seeds  of  Vicia  faba  (small  seeded  varieties) 
and  V.  melanops  and  has  seeds  for  distribution  of  Vicia,  Lupinus 
(Mediterranean),  Cicer.  Will  be  in  North  Africa  during  summer  of  1982 
collecting  legumes. 

CUSATO  is  a new  Reader. 

DELBOS  is  a new  Reader. 

ESTEVE  GARCIA  is  a new  Reader. 

EVANS  is  conducting  a chemotaxonomic  survey  of  the  Tephrosieae,  especially 
nonprotein  amino  acids  as  part  of  his  Ph.  D.  thesis  on  Toxins  in  the 
Tephrosieae.  He  would  like  to  exchange  material. 

FERGUSON  and  R.  Strachan  have  completed  a manuscript  on  the  pollen  morphology 
of  Indigofera. 


-6- 


GIANGUALANI  continues  her  study  of  Argentine  Vicia  species  even  though  she 
moved  to  Brazil. 

GILL  has  had  a cytomorphology  study  of  selected  Cassia  species  in  Nigeria 
published  in  Adansonia  and  has  a similar  paper  on  arborescent  legumes  in 
press  in  Silvae  Genetica.  Has  started  studies  of  Nigerian  legume  pollen, 
tracheary  elements,  Cassia  seed  germination,  and  effect  of  temperature  on 
seed  germination  of  Pentaclethra  macrophylla.  Suggests  study  of  tropical 
legume  seed  anatomy.  Needs  viable  seeds  of  African  legumes. 

GRETHER-GONZALEZ  is  working  with  Mimosa  for  Flora  de  Nicaragua  and  will  be 
collecting  in  northwest  Mexico  during  August  and  September,  1982. 

GUNN  notes  with  a mixture  of  weariness  and  pleasure  that  the  U.S.  National 
Seed  Herbarium  is  moving  to  larger  quarters  in  order  to  accommodate  an 
expansion  in  staff  and  herbarium  cases.  For  the  first  time  all  cases 
will  be  in  one  room  with  ample  adjacent  work  space. 

HEISER  has  for  distribution  a limited  number  of  copies  of  his  1981  Presidental 
address  to  the  Botanical  Society  of  America,  titled  Taxonomy  and  Ecology  of 
North  American  Gleditsia.  He  does  not  plan  to  publish  this  in  the  near 
future. 

JOHNSON  has  started  separate  revisions  of  Acanthoscelides  (Coleoptera: 
Bruchidae)  for  southern  Mexico  and  Central  America,  and  northern  South 
America. 

KEIGHERY  has  studies  of  pollination  of  Jansonia  in  press  in  Western  Australia 
Naturalist,  pollination  of  Cupulanthus  (completed),  and  phytogeography  of 
western  Australian  Fabaceae  (completed).  Has  started  separate  studies  of 
pollination  biology  and  chromosome  numbers  of  western  Australian  legumes. 
Also  started  a study  of  legume  breeding  systems  (a  resume  of  this  work  was 
used  by  ARROYO  in  her  chapter  of  the  Proceedings  of  the  International 
Legume  Conference. 

KINGSOLVER  has  completed  a study  of  bruchids  (Rhipibruchis ) associated  with 
South  American  Prosopis  and  started  separate  studies  of  bruchids 
( Scutobruchus ) associated  with  South  American  Prosopis  and  bruchids 
associated  with  Brazilian  Parkia.  Would  like  bruchids  reared  from  South 
American  Prosopis  and  Parkia.  Will  identify  and  return  insects. 

KRUKOFF.  See  STIRTON. 

LABEYRIE.  See  COMBES. 

LACKEY  spent  most  of  April  and  part  of  May  in  Botswana. 

LANGENHEIM  has  completed  a study  of  photosynthetic  responses  of  Hymenaea  and 
Copaifera  seedlings  while  at  the  Australian  National  University,  Canberra. 
Has  started  comparative  studies  of  secondary  chemical  and  lepidopteran 
interactions  in  Hymenaea  and  Copaifera  of  southwestern  Brazil. 


-7- 


LEELAVATHI  with  MALVEY  has  finished  several  studies:  Trichomes  in 

Caesalpinioideae  and  Faboideae,  styloids  in  Fabaceae,  and  stomata  in 
Fabaceae.  Interested  in  working  with  medicinal  and  toxic  legumes.  Wants 
seeds  of  Stylosanthes,  Canavalia,  and  Rhynchosia. 

LERSTEN  has  completed  a SEM  survey  of  seed  coat  anatomy  and  surface  topography 
among  tribes  of  the  Faboideae. 

MALVEY.  See  LEELAVATHI . 

MARECHAL  and  BAUDOIN  have  prepared  a descriptor's  list  for  evaluating  the  lima 
bean. 

MOHLENBROCK  with  D.  R.  Bissing  has  started  an  anatomical  study  of  New  World 
Pithecellobium.  Also  started  revision  of  Enterolobium  and  Apios  priceana, 
and  continuing  with  studies  of  Stylosanthes  and  Zornia.  Needs  wood  samples 
of  North  American  Pithecellobium. 

MUINZ  is  a new  reader  working  on  endangered  Cuban  species  with  Attila 
Borhidi.  Will  be  collecting  in  Cuba. 

NEILL  has  completed  field  studies  of  Central  American  Erythrina  (variability, 
pollination,  seed  predation,  chromosome  counts).  As  part  of  dissertation 
will  start  a hybridization  program  at  the  Pacific  Tropical  Botanical 
Garden,  Hawaii.  Needs  seeds  of  certain  Erythrina  species,  expecially  from 
southern  Mexico  and  tropical  Africa.  Offers  Erythrina  seeds  and  cuttings, 
which  will  remain  viable  even  if  mailed.  Will  be  collecting  in  Central 
America  during  the  winter  of  1983. 

NOZZOLILLO  has  finished  a biometric  study  of  Medicago,  based  on  the  thesis  of 
Debbie  Classen  with  SMALL,  to  be  published  in  the  Canadian  Journal  of 
Botany.  She  is  the  1981-82  President  of  the  Phytochemical  Society  of 
North  America.  Meeting  and  symposium  (Mobilization  of  Reserves  in 
Germination)  to  be  held  at  the  University  of  Ottawa,  August  2-6,  1982. 

POLHILL.  Sent  Flora  Neotropical  data  in  separate  column.  See  also  STIRTON. 

REID  is  a new  Reader. 

RODRIGUEZ  PEREZ  completed  a planting  study  of  Phaseolus  acutifolius  var. 
latifolius  in  Teneriffe. 

SENESSE  is  studying  caesalpinioid  pollen. 

SHARMA  is  studying  the  relationship  of  pollution  to  white  clover  and  alfalfa. 
Will  be  collecting  in  Central  Asia. 

SMALL.  See  NOZZOLILLO. 

SOLEDAD  is  a new  Reader. 

STANKEVICZ  (mispelled  in  BB-14  Directory  as  Stankovicz)  has  completed  a study 
of  Vicia  subvillosa.  Will  be  studying  Section  Orobus  of  Lathyrus. 


-8- 


STIRTON  has  completed  a study  of  the  7 species  complex  known  as  Eriosema 
squarrosum.  Is  starting  a revision  of  the  ca  125  species  of  African 
Psoraleeae.  He  will  annotate  and  needs  seeds  of  African  Psoralea.  He  was 
awarded  a five  year  fellowship  to  study  New  World  Leguminosae  (excluding 
the  Mimosoideae) . This  fellowship  has  been  made  possible  by  a Trust  Fund 
set  up  by  KRUKOFF  and  administered  by  the  Missouri  Botanic  Garden.  He 
will  be  based  at  Kew  and  will  work  under  POLHILL.  While  details  are  not 
established,  the  generic  circumscription  of  New  World  Sophoreae  have  a 
high  priority. 

WEDER  with  H.-D.  Belitz  and  F.  Lynen  have  completed  comparative  studies  on  the 
inhibitory  action  of  some  legume  seeds  against  human  and  bovine 
proteinases.  Has  started  research  on  proteinase  inhibitors  in  Lens 
culinaris,  and  has  started  with  D.  R.  Murray  an  application  of  proteinase 
inhibitor  pattern  studies  on  the  taxonomy  of  the  Pulchellae  group  of 
Acacia. 

ZINDLER-FRANK  with  H.  T.  Horner  has  two  papers  in  press:  Calcium  oxalate 

crystals  and  crystal  cells  in  leaves  of  Rhynchosia  caribaea  in  Protoplasma; 
and  histochemical , spectroscopic,  and  dif fractometric  identification  of  the 
two  hydration  forms  of  calcium  oxalate  crystals  in  three  legumes  and 
Begonia  in  Canadian  Journal  of  Botany. 

INIA,  Centro  Regional  de  Canarias  is  a new  Reader. 
******************************************************************************** 


Arco,  M.  del.  1982.  Sobre  la  presencia  de  Genista  L.  en  la  isla  de  La  Palma 
(I.  Canarias):  Genista  benehaoavensis  (Bolle  ex  Svent)  M.  del  Arco,  comb, 

nov.  (Fabaceae-Genisteae) . Vieraea  11  (1-2) : 267-282. 

Arco,  M.  del,  and  ACEBES.  1982.  Teline  salsoloides  M.  del  Arco  & J.  R. 

Acebes,  sp.  nov.  (Fabaceae-Genisteae),  una  nueva  especie  para  la  isla  de 
Tenerife  (I.  Canarias).  Vieraea  11(1-2) :251-266. 

BAHADUR  and  RAO.  1981.  Seedling  hardedness  in  Fabaceae.  Proc.  Ind.  Acad. 
Sci.  (Plant  Sci.)  90:231-236. 

BARNEBY.  1981.  New  species  of  Dalea  section  Parosela  (Leguminosae: 

Amorpheae)  from  Peru  and  Mexico.  Brittonia  33(4):508-511. 

BASSLER.  1981.  Revision  of  Lathyrus  section  Lathyrostylis  Fabaceae.  Feddes 
Repert  92(3) : 179-254. 

Bazilevskaya,  N.  A.  1980.  A key  to  the  weed  genera  of  the  order  Fabales 
based  on  their  fruits.  Moscow  Univ.  Biol.  Sci.  Bull.  35(2):48-51. 

CARRILLO-FUENTES . 1982.  Las  Leguminosas  del  Valle  de  Rimac  (Mimosoideae  and 

Caesalpinioideae . Bol.  Soc.  Peruana  Bot.  7(1-2). 

CARTER  and  RUDD.  1981.  A new  species  of  Acacia  (Leguminosae:  Mimosoideae) 
from  Baja  California  Sur,  Mexico.  Madrono  28:220-225. 

COWAN  and  STAFLEU.  1981.  Rose  and  Britton:  From  Brittonrosea  to  Cassia. 

Brittonia  33(3) :285-293. 

Crankshaw,  D.  R. , and  LANGENHEIM.  1981.  Variation  in  terpenes  and  phenolies 
through  leaf  development  in  Hymenaea  and  its  possible  significance  to 
herbivary.  Biochem.  Syst.  Ecol.  9:115-160. 


RECENT  LEGUME  LITERATURE 


-9- 


DELGADO  SALINAS.  1981.  Macroptilium  gibbosi folium  (Ortega)  A.  Delgado,  the 
correct  name  for  M.  heterophylliim  (Willd.)  Marechal  & Baudet  (Fabaceae: 
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