Bonn zoological bulletin

BAZ E. museum
Nie AML 6 ENiG

Bonn Volume 59
: 2010
Zoological

Bulletin

formerly: Bonner zoologische Beitrage

ZFMK type catalogues: Vertebrata

An open access journal of organismal zoology, published by
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Bonn zoological Bulletin (BzB), formerly “Bonner zoologische Beitrage”, is published by the Zoologisches Forschungsmuseum
Alexander Koenig (ZFMK), Bonn. Two regular issues of BzB are published per year; supplements on focus topics are produced

in irregular succession.

Subscription price is 46 € per volume (year), including supplements and shipping costs. For subscription, back issues and insti-
tutional exchange, please contact the ZFMK library (ZFMK, Bibliothek, Frau Diane Steinebach, Adenauerallee 160, D-53113 Bonn,
Germany, tel. +49 228—9122-216, fax: +49 2289122212; d.steinebach.zfmk@uni-bonn.de). The online version of BzB is avail-
able free of charge at the ZFMK homepage: http://www.zfmk.de/web/Forschung/Buecher/Beitraege/index.en.html.

© 2010 Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany. ISSN 2190-7307
Produced by Eva-Maria Levermann, Kaiserstr. 129, D-53113 Bonn, Germany; emlevermann@netcologne.de.
Printed and bound by DCM, Werner-von-Siemens-Str. 13, D-53340 Meckenheim, Germany.

Bonn zoological Bulletin

Editor-in-Chief

Fabian Herder, Zoologisches Forschungsmuseum Alexander
Koenig (ZFMK), Ichthyology Section, Adenauerallee 160,
53113 Bonn, Germany,
tel. +49 228-9122—255, fax: +49 228-9122-212:;
f.herder.zfmk@uni-bonn.de

Editorial Board

Dirk Ahrens, Insects: Coleoptera, ZFMK,
tel. +49 228-9122—286, fax: +49 228-9122-332;
d.ahrens.zfmk@uni-bonn.de

Wolfgang Bohme, Amphibians and Reptiles, ZFMK,
tel. +49 228-9122—250, fax: +49 228-9122-212;
w.boehme.zfmk(@uni-bonn.de

Netta Dorchin, Insects: Diptera, ZFMK,
tel. +49 228-9122-292, fax: +49 228-9122-212;
n.dorchin.zfmk(@uni-bonn.de

Renate van den Elzen, Birds, ZFMK,
tel. +49 228-9122—231, fax: +49 228-9122-212;
r.elzen.zfmk(@uni-bonn.de

Bernhard Huber, Invertebrates except Insects, ZFMK,
tel. +49 228-9122-294, fax: +49 228-9122-212;
b.huber.zfmk@uni-bonn.de

Rainer Hutterer, Mammals, ZFMK,
tel. +49 228-9122—261, fax: +49 228-9122—212;
r.hutterer.zfmk(@uni-bonn.de

Gustav Peters, Mammals, ZFMK,
tel. +49 228-9122—262, fax: +49 228-9122-212;
g.peters.zfmk@uni-bonn.de

Bradley Sinclair, Canadian National Collection of Insects,
Ottawa Plant Laboratory — Entomology, CFIA, K.W. Neat-
by Bldg., C.E.F., 960 Carling Ave., Ottawa, ON, Canada
K1A 0C6, tel. + 1 613-759-1787, fax: + 1 613-759-1927;
bradley.sinclair@inspection.ge.ca

Dieter Stiining, Insects except Coleoptera and Diptera,
ZFMK, tel. +49 228—-9122—220, fax: +49 228-9122-212;
d.stuening.zfmk@uni-bonn.de

Advisory Board

Theo C. M. Bakker, Rheinische Friedrich-Wilhelms-Univer-
sitat, Institut fiir Evolutionsbiologie & Okologie, 53113
Bonn, Germany, tel. +49 228—-73-5130, fax: +49 228—-73-—
2321; t.bakker@uni-bonn.de

Aaron M. Bauer, Villanova University, Department of Biolo-
gy, 800 Lancaster Avenue, Villanova, PA 19085-1699,
USA, tel. +1 610-519-4857, fax: +1 610-519-7863;
aaron.bauer@villanova.edu

Wieslaw Bogdanowicz, Museum and Institute of Zoology,
Polish Academy of Sciences, Wilcza 64, 00-679 Warszawa,
Poland, tel. +48 22628-7304, fax: +48 22-629-6302;
wieslawb@miiz.waw.pl

Matthias Glaubrecht, Museum fiir Naturkunde Berlin, Leib-
niz-Institut fiir Evolutions- und Biodiversitatsforschung an
der Humboldt-Universitat zu Berlin, Invalidenstrasse 43,
10115 Berlin, Germany,
tel. +49 30-2093—8504/ 8400, fax: +49 030—2093-8565;
matthias.glaubrecht@mfn-berlin.de

Jeremy D. Holloway, The Natural History Museum, Depart-
ment of Entomology, Cromwell Road, London, SW7 5BD,
U.K.; j-holloway@nhm.ac.uk

Boris Krystufek, Slovenian Museum of Natural History, P.
O. Box 290, Ljubljana, Slovenia; boris.krystufek@zrs.upr.si

Wolfgang Schawaller, Staatliches Museum ftir Naturkunde,
Rosenstein 1, 70191 Stuttgart, Germany,
tel. +49 711-8936-221, fax: +49 711-8936—100;
schawaller.smns@naturkundemuseum-bw.de

Ulrich K. Schliewen, Department of Ichthyology, Bavarian
State Collection of Zoology, Mtinchhausenstr. 21, 81247
Miinchen, Germany, tel. + 49 89-8107-110;
schliewen@zsm.mwn.de

Michael Schmitt, Ernst-Moritz-Arndt-Universitat, Allge-
meine & Systematische Zoologie, Anklamer Str. 20, 7489
Greifswald, Germany,
tel. +49 3834864242, fax: +49 3834864098;
michael.schmitt@uni-greifswald.de

W. David Sissom, Dept. of Life, Earth and Environmental
Sciences, W. Texas A. & M. University, WITAMU Box
60808, Canyon, Texas 79016, USA, tel. +1 806-651-2578,
fax: +1 806-651-2928; dsissom@mail.wtamu.edu

Miguel Vences, Technical University of Braunschweig, Zoo-
logical Institute, Spielmannstr. 8, 38106 Braunschweig,
Germany, tel. + 49 531-391-3237,
fax: + 49 531-391-8198; m.vences@tu-bs.de

Erich Weber, Eberhard-Karls-Universitat, Zoologische
Schausammlung, Sigwartstr. 3, 72076 Tubingen,
tel. +49 70712972616, fax +49 7071—295170;
erich.weber@uni-tuebingen.de

Editorial

This special volume of the Bonn zoological Bulletin (BzB)
constitutes the second part of the ZFMK type catalogues.
It contains records of vertebrate type specimens housed
in the museum’s scientific collections, subdivided into the
sections Theriology (mammals), Ornithology (birds), Her-
petology (amphibians and reptiles), and Ichthyology (fish).
Together with the catalogues of the arthropod type spec-
imens published in volume 58 of the BzB, this work pro-
vides an overview of the most valuable assets of the mu-
seum’s scientific collections. Summarizing core data on
virtually all animal taxa that are represented by type spec-
imens in Bonn, these catalogues will serve taxonomists
as a valuable tool and guide when working in or referring
to the institute’s extensive collections.

The last comprehensive publication on vertebrate type ma-
terial in the ZFMK dates back to 1984, and was published
on the occasion of the 50th anniversary of Museum
Koenig as a public zoological museum. Since then, the
number of vertebrate types in the ZFMK increased con-
siderably (birds) or even doubled (mammals, amphibians,
reptiles and fishes). This increase underlines the impor-
tance of natural history museums as institutions that guar-
antee the maintenance of and access to scientific materi-
al, and highlights the need for continuous, substantial work
on basic taxonomy.

I thank all authors who contributed to the two volumes of
the type catalogues for their continuous efforts. Compil-
ing such catalogues and verifying the accuracy of the da-
ta therein are extremely time-consuming tasks that are of-
ten largely underestimated. I am especially happy for hav-
ing been able to cover virtually all groups of organisms
represented by type specimens in the ZFMK, thereby con-
tributing to an inventory of the fascinating animal collec-
tions available in Bonn.

With best wishes to all readers, authors and “BzB-Team”
members,

Tibi float

Fabian Herder (Editor-in-Chief, Fish Curator at ZFMK)
Bonn, December 2010

nia SOF

Bonn zoological Bulletin Volume 59

Bonn, December 2010

Type specimens of mammals (Mammalia) in the collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Rainer Hutterer & Gustav Peters

Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn, Germany;
E-mail: r.hutterer.zfmk@uni-bonn.de, g.peters.zfmk@uni-bonn.de

Abstract. An annotated catalogue of type specimens of Mammalia in the collections of the Zoologisches Forschungsmu-
seum Alexander Koenig in Bonn is given. By the end of 2010, the Bonn collections housed name-bearing types (five
syn-, 77 holo-, two lecto- and one neotype) for 85 taxa and para- and paralectotypes for 105 taxa. Casts of rare type spec-
imens of 3 extinct taxa are also listed. The collections also include vouchers for karyotypes and GenBank entries. A lec-
totype is selected for Microtus savii niethammericus Contoli, 2003, and replacement names are proposed for homonyms
of two preoccupied taxa, Apodemus maximus Thaler, 1972 (Muridae) and Sciurus vulgaris hoffmanni Valverde, 1967 (Sci-

uridae).

Key words. Taxonomy, types, vouchers, chromosomes, genetic sequences, mammal diversity.

INTRODUCTION

Biological collections of the world preserve the voucher
specimens on which our knowledge of the biodiversity on
our planet is based. Article 72.10 of the Code (ICZN 1999)
states that, “Holotypes, syntypes, lectotypes and neotypes
are the bearers of the scientific names of all nominal
species-group taxa (and indirectly of all animal taxa). They
are the international standards of reference that provide
objectivity in zoological nomenclature and must be cared
as such. They are to be held in trust for science by the per-
sons responsible for their safe keeping.” Recommendation
72F includes the following formulations “Every institu-
tion in which name-bearing types are deposited should ....
(2) take all necessary steps for their safe preservation, (3)
make them accessible for study, (4) publish lists of name-
bearing types in its possession or custody, and (5) so far
as possible, communicate information concerning name-
bearing types when requested.”

The collections of Mammalia in the Zoologisches
Forschungsmuseum Alexander Koenig house more than
90000 specimens representing about 1800 species, i.e.
about 33% of the known extant mammalian diversity.
These collections form part of the world holdings of mam-
malian archives (Genoways & Schlitter 1981) and have
contributed, among others, to the production of taxonom-
ic standard works such as the current world list of mam-
malian species (Wilson & Reeder 2005).

Bonn zoological Bulletin 59: 3-27

The first type catalogue of mammals in the Zoologisches
Forschungsmuseum Alexander Koenig was published
more than 25 years ago (Hutterer 1984). Here we present
an updated annotated list of the name-bearing types (holo-
type, syntype, lectotype, neotype), but also of paratypes
and paralectotypes, as these form part of the original type
series and are available for the selection of neotypes, if
required (Article 75, Recommendation 75A, ICZN 1999).
In a few cases we have also listed casts of type specimens
from other institutions in order to facilitate access to the
types of some extinct species. The collections also include
vouchers for chromosome counts and for genetic se-
quences. If available, such information has been men-
tioned but it was not possible to compile a full list of such
voucher specimens. As GenBank entries already play an
important role in the study of mammalian phylogeny and
taxonomy, cross-references to the vouchers on which the
genetic sequences were based will become more and more
important in order to ensure scientific objectivity and to
enable verification of correctness of specimen identifica-
tions when indicated. Figures listed for the respective orig-
inal description of a taxon are all figures in that publica-
tion which show features of the new taxon.

The present list is complied in the style of other modern
examples such as Lawrence (1993), Turni et al. (2007) and
Turn & Kock (2008). The systematics, sequence of or-

©OZFMK

4 Rainer Hutterer & Gustav Peters

ders and families within orders follows Wilson & Reed-
er (2005). Likewise, the taxonomy largely follows the re-
spective chapters in Wilson & Reeder (2005), particular-
ly those on soricomorphs (Hutterer 2005), rodents (Muss-
er & Carleton 2005), glirids (Holden 2005), sciurids (Thor-
ington & Hoffmann 2005) and artiodactyls (Grubb 2005).
We included no figures in this catalogue as photographs
of many of the ZFMK type specimens are available on-
line through the Global Biodiversity Information Facili-
ty (http://www.biologie.uni-ulm.de/cgi-bin/query_all/
query_all.pl?lang=d&pr=gbif-v1).

Abbreviations for museum collections used:

AMNH = American Museum of Natural History, New
York, USA,

BMNH = The Natural History Museum, London, UK,
CM = Carnegie Museum, Pittsburgh, USA,

FMNH = Field Museum of Natural History, Chicago,
USA,

HNHM = Hungarian Natural History Museum, Budapest,
Hungary,

HZM = Harrison Zoological Museum, Sevenoaks, UK,
IZEA = Institut de zoologie et d’écologie animale, Lau-
sanne, Switzerland,

MNHN = Museum National d’Histore Naturelle, Paris,
France,

NMP = National Museum of the Philippines, Manila, The
Philippines,

NMW = Naturhistorisches Museum Wien, Vienna, Aus-
tria,

QM = Queensland Museum, Brisbane, Australia,

PBS = Paimpont Biological Station of Rennes Universi-
ty, France,

RMCA = Royal Museum for Central Africa, Tervuren,
Belgium,

SMF = Senckenberg Museum, Frankfurt, Germany,
SMNS = Staatliches Museum fiir Naturkunde, Stuttgart,
Germany,

TM = Transvaal Museum, now Ditsong National Muse-
um of Natural History, Pretoria, South Africa,

ZFMK = Zoologisches Forschungsmuseum Alexander
Koenig, Bonn, Germany,

ZIN = Zoological Institute of the Academy of Sciences,
Moscow, Russia,

ZMB = Zoologisches Museum der Humboldt-Universi-
tat, Berlin, Germany.

Bonn zoological Bulletin 59: 3—27

CATALOGUE OF TYPE SPECIMENS
MONOTREMATA
Ornithorhynchidae

Obdurodon dicksoni Archer, Jenkins, Hand, Murray &
Godthelp, 1992

In: Platypus and Echidnas (Augee ML, ed.): 16, figs. 1-8
Material. ZFMK 95.167, cast of holotype skull QM
F20568, Middle Miocene sediments of Riversleigh, Aus-
tralia.

Current status. Obdurodon dicksoni Archer, Jenkins,
Hand, Murray & Godthelp, 1992.

Comments. Cast donated by I. Rocek in 1995. A full re-
construction of skull and dentary was provided by Muss-
er & Archer (1998).

AFROTHERIA
Chrysochloridae

Eremitalpa granti namibensis Bauer & J. Niethammer,
1960

Bonn. zool. Beitr. 10: 241, fig. 2 (10 Jan)

Holotype. ZFMK 59.513, cranium and mandible taken
from owl pellets, in good condition; Sossus Vley, Namib
Desert, Namibia, collected by G. Niethammer, Feb 1959,
field no. 47.

Paratypes. ZFMK 59.514 (40 skulls), and further 16 un-
catalogued skulls plus uncounted postcranial elements, all
from the same locality and collector.

Current status. Evemitalpa granti namibensis Bauer &
J. Niethammer, 1960.

Comments. Further paratypes are in the AMNH, NMVW
and TM.

MACROSCELIDEA
Macroscelididae

Elephantulus intufi omahekensis Lehmann, 1955
Bonn. zool. Beitr. 6: 171

Holotype. ZFMK 55.74, female, skin and skull; K1. Oka-
puta, Namibia, collected by W. Hoesch, 16 Oct 1954, field
no. M. 1130.

Paratypes. ZFMK 55.78, 55.120—55.125, 4 males, 3 fe-
males, skins and skulls, same locality and collector as
holotype, 14-18 Oct 1954.

Current name. Elephantulus intufi (A. Smith, 1836).

©ZFMK

Types in ZFMK: Section Theriology 5

CINGULATA
Dasypodidae

Cabassous chacoensis Wetzel, 1980

Ann. Carnegie Mus. 49: 335, fig. 2 (5 Sept)

Paratype. ZFMK 60.317, adult female, skin and skull;
Filadelfia, Paraguay, collected by J. Unger, 10 Dec 1958,
field no. 448.

Current status. Cabassous chacoensis Wetzel, 1980.
Comments. Holotype CM 67067, taken at 5—7 km W Es-
tancia Juan de Zalazar, Depto. Presidente Hayes, Paraguay
(Wetzel 1980).

LAGOMORPHA
Ochotonidae

Lagomys ater Eversmann, 1842

Addenda ad Celeberrimi Pallasii Zoographiam Rosso-
Asiaticam, Fasc. 3: 3.

Material. Possible paralectotype (see below), ZFMK
88.64, juvenile female, mount and skull (extracted in
1988); Russia, Altai, sold by J.G.W. Brandt 1851 to the
Zoologial Institute Heidelberg (No. 391), exchanged in
1988.

Current name. Ochotona alpina atra (Eversmann, 1842)
or Ochotona a. alpina (Pallas, 1773); see Lissovsky
(2003).

Comments. Johann Georg Wilhelm Brandt (1794-1856)
opened a warehouse for natural history specimens in Ham-
burg in 1827. He had world-wide connections to travel-
lors and collectors (Stresemann 1967). It is likely, but not
certain, that he purchased specimens from Eduard Evers-
mann (1794—1860), a German explorer and scientist who
held a professorship at the University of Kazan in Russia
for several decades and who sold specimens to German
museums such as the Berlin Museum (Gebhard 1964).
Lagomys ater was collected in the Altai Mountains, as stat-
ed on the label of the ZFMK specimen. Lissovsky (2003)
discussed the status of the Altai pika Ochotona alpina (Pal-
las, 1773) and the type locality and type series of O. a.
atra (Eversmann, 1842). There are three specimens of
Ochotona a. atra collected by Eversmann in 1841 in the
ZIN. Yakhontoy & Formozov (1992) designated ZIN
84398 as lectotype, and the other two specimens are para-
lectotypes. Provided that Eversmann distributed other
specimens from his type series among German museums,
ZFMK 88.64 could be a further paralectotype.

Bonn zoological Bulletin 59: 3-27

Leporidae

Pronolagus randensis waterbergensis Hoesch & Leh-
mann, 1956

Bonn. Zool. Beitr. 7: 32, fig. 6

Holotype. ZFMK 34.144, male, skin and_ skull;
Otjosongombe, Waterberg, Namibia, caught by W.
Hoesch, | June 1934, field no. XXXI.

Current name. Pronolagus randensis caucinus Thomas,
1929 (Hoffmann & Smith 2005),

ERINACEOMORPHA
Erinaceidae

Erinaceus algirus girbaensis Vesmanis, 1981

Bonn. zool. Beitr. 31: 207, figs. 1-4 (29 Jan 1981)
Holotype. ZFMK 79.464, skin and skull; Djerba Island,
Tunisia, collected May 1932, purchased from Fritsche.
Paratype. ZFMK 79.463, skin and skull, same data as
holotype.

Current status. Atelerix algirus girbaensis (Vesmanis,
1981) (Corbet 1988).

SORICOMORPHA
Soricidae

Congosorex verheyeni Hutterer, Barriere & Colyn, 2001
Bull. Inst. Royal Sci. Nat. Belg., Biol. 72, Suppl.: 10, figs.
1-6 (20 Dec 2001)

Holotype. ZFMK 99.932, female, specimen preserved in
spirit and skull; Mbomo, Pare National d’Odzala, Repub-
lic of the Congo, collected by members of the ECOFAC
team, 19 Oct 1996, field no. R16761.

Current name. Congosorex verheyeni Hutterer, Barrie-
re & Colyn, 2002.

Comments. Paratypes in MNHN, RMCA and in PBS.

Congosorex phillipsorum Stanley, Rogers & Hutterer,
2005

J. Zool., London 265: 271, figs. 3-8

Paratype. ZFMK 2009.631 (formerly FMNH 169493), al-
coholic specimen, skull removed; West Kilombero Scarp,
2062 m, Udzungwa Mts, Iringa District, Iringa Region,
Tanzania; collected by K.M. Howell, 15 Jul 2000, field
no. KMH 19125.

Current name. Congosorex phillipsorum Stanley, Rogers
& Hutterer, 2005.

Comments. Holotype in FMNH.

©ZFMK

6 Rainer Hutterer & Gustav Peters

Crocidura arabica Hutterer & Harrison, 1988

Bonn. zool. Beitr. 39: 64, figs. 3, 5, 6

Paratype. ZFMK 88.14, cranial fragment taken from owl
pellet; near Tawi Atair, Jebel Samhan, Dhofar, Oman, col-
lected by M.D. Gallagher, 9 Apr 1977; donated by D.L.
Harrison (HZM 19.8767).

Current status. Crocidura arabica Hutterer & Harrison,
L988.

Crocidura ariadne Pieper, 1979

Bonn. zool. Beitr. 29: 282, figs. 1-6 (30 Jan)

Holotype. ZFMK 78.100, cranial fragment taken from owl
pellet; Agia Pnevma, Crete, Greece, collected by Harald
Pieper, 23 Sept 1973.

Current status. Crocidura suaveolens caneae (Hutterer
1981).

Crocidura batakorum Hutterer, 2007

Lynx (Praha) n.s. 38: 11, figs. 5—8

Holotype. ZFMK 96.411, old adult male(?), specimen in
spirit, skull extracted, part of left braincase damaged; Tan-
abag River Valley near village Kalabayog (09°44’N,
118°43’E), ca. 60 km N Puerto Princesa, Palawan, Repub-
lic of the Philippines, 200 m, collected by S. Biichner, 10
Mar 1992.

Current name. Crocidura batakorum Hutterer, 2007.
Comments. Status confirmed by Esselstyn et al. (2009).

Crocidura (bottegi) obscurior Heim de Balsac, 1958
Mem. Inst. fr. Afr. noire 53: 328, fig. 24

Paratype. ZFMK 59.499, female preserved in spirit, skull
extracted and lost; “Foret Ziéla”, 500 m, Mount Nimba,
Guinea, collected by M. Lamotte, 24 Oct 1956, collector’s
no. 59.

Current name. Crocidura obscurior Heim de Balsac,
1958

Comments. Specimen donated to M. Eisentraut by H.
Heim de Balsac. Holotype probably in MNHN.

Crocidura canariensis Hutterer, Lopez-Jurado & Vogel,
1987

J. nat. Hist. 21: 1354, figs. 2-4

Holotype. ZFMK 86.75 (transferred to Museo Canario at
Las Palmas in 1987), adult male, specimen preserved in
spirit, skull and mandibles extracted; offspring of animals
collected at La Tiscamanita, N Tuineje, Fuerteventura, Ca-
nary Islands, Spain, by L. F. Lopez-Jurado, 30 Jan 1985.
Chromosome slides (IZEA 2331; Hutterer et al. 1990, fig.
2) and frozen tissue kept at IZEA, Lausanne.
Paratypes. ZFMK 86.76, 86.370, two females in alcohol,
same data as holotype; ZFMK 86.72-74, three skeletons,
same data as holotype; ZFMK 86.371, skin from La Oli-
va, Fuerteventura, Nov 1984; ZFMK 86.372, skin from
Malpais de los Arravales, Fuerteventura; ZFMK 86.374-

Bonn zoological Bulletin 59: 3-27

380, skulls from owl pellets collected near Gran Tarajal,
Fuerteventura, all collected by L. F. Lopez-Jurado.
Current name. Crocidura canariensis Hutterer, Lopez-
Jurado & Vogel, 1987.

Comments. Holotype ZFMK 86.75 is voucher for the
GenBank sequences EF525043 and EF525160 (IZEA
2331, Dubey et al. 2008). Paratype ZFMK 86.76 is vouch-
er for the chromosome count 2n = 36, NF = 56, NFa = 52
in Hutterer et al. (1992), table 1 (IZEA voucher 2332).

Crocidura flavescens bueae Heim de Balsac & Barloy,
1966

Mammalia 30: 631

Paratype. ZFMK 78.004, skin and skull of a juvenile fe-
male; garden of mission, Buea, ca. 1000 m, Cameroon,
collected by M. Eisentraut, 26 Jan 1954, field no. 54, re-
ceived in exchange from SMNS in 1978 (catalogue no.
SMNS 5561)

Current name. Crocidura olivieri bueae Heim de Bal-
sac & Barloy, 1966.

Comments. The holotype out of the same series (“femelle,
No. 53, Missionsgarten, Buea (Eisentraut ...)”) was in the
working collection of Heim de Balsac and today is in the
MNHN, Paris. In their short description the authors sta-
ted “Cotypes a choisir parmi les specimens de Buea, au
Musée de Bonn’. The series collected by Eisentraut near
Buea in 1954 and 1957 and studied by Heim de Balsac
ended up at the SMNS, though. Apart from the paratype
in the ZFMK collection which was received in exchange
in 1978, there are three more paratypes in the SMNS with
the collection nos. SMNS 5559 (field no. 35), 5560 (field
no. 49) and 6561 (field no. 11).

Crocidura flavescens hansruppi Hutterer, 1981

Bonn. zool. Beitr. 31: 226, fig. 3 (29 Jan)

Paratype. ZFMK 80.265, female, skin and skull; Jimma,
Kaffa Province, Ethiopia, collected by H. Rupp, 23 Aug
1973, collector’s no. 477; received in exchange from
Stuttgart Museum (SMNS 23703), Apr 1980.

Current name. Crocidura olivieri group, status unre-
solved (Hutterer 2005).

Comments. This form represents a dark chocolate-brown
forest form of the C. o/ivieri group in Ethiopia, the vari-
ation of which 1s still poorly understood.

Crocidura lamottei elegans Hutterer, 1986

Cimbebasia, Ser. A., 8: 29

Holotype. ZFMK 76.009, male, skin and skull; Bandia,
Senegal, collected by W. B6hme, W. Hartwig and H. UI-
lenbruch, 9-20 Jan 1976, field no 279.

Paratype. ZFMK 76.010, female, skin and skull, same
dara as holotype, field no. 234.

Current name. Crocidura lamottei elegans Hutterer,
1986.

©ZFMK

Types in ZFMK: Section Theriology aI

Crocidura lanosa Heim de Balsac, 1968

Biol. Gabonica 4: 309, figs. 1-4

Paratypes. ZFMK 89.458, 89.460, unsexed skins and
skulls; Uwinka, Rwanda, collected by U. Rahm, 10 Jun
1962, field no. E665, E657; ZFMK 89.459, female, skin
and skull; Lemera, Kivu province, Democratic Republic
of the Congo, collected by U. Rahm, 19 Oct 1962. Spec-
imens donated by U. Rahm in 1989.

Current name. Crocidura lanosa Heim de Balsac, 1968.

Crocidura littoralis stenocephala Heim de Balsac, 1979
Saugetierkundl. Mitt. 27: 258, figs. 4-6 (Nov)
Paratype. ZFMK 79.141, female, skin and skull; Kahuzi
Biega National Park, Kivu, Democratic Republic of the
Congo, collected by F. Dieterlen, 15 Nov 1972, field no.
K 2544; received in exchange from SMNS (SMNS cata-
logue no. 22806).

Current name. Crocidura stenocephala Heim de Balsac,
1979 (Hutterer 1982).

Comments. Holotype and remaining paratypes in SMNS.
The species was described in Dieterlen & Heim de Bal-
sac (1979).

Crocidura manengubae Hutterer, 1982

Bonn. zool. Beitr. 32: 241, figs. 1, 4 (Jan)

Holotype. ZFMK 81.443, male, skin and skull; Lake Ma-
nenguba, 1800 m, Bamenda Highlands, Cameroon, col-
lected by M. Eisentraut and assistants, 28 Dec 1966, field
no. 306.

Paratype. ZFMK 69.409, female, skin and skull; same lo-
cality and collectors, 3 Jan 1967, field no. 321.

Current name. Crocidura manengubae Hutterer, 1982.

Crocidura osorio Molina & Hutterer, 1989

Bonn. zool. Beitr. 40: 86, figs. 2, 3, 8, 13

Paratypes. ZFMK 89.54, specimen in spirit, skull extract-
ed; near Teror, Gran Canaria, Spain, collected by staff of
the Finca de Osorio, 1988, field no. 3.

Current name. Crocidura russula osorio Molina & Hut-
terer, 1989 (Hutterer 2005).

Comments. Holotype (field no. 42) deposited in the Mu-
seo Canario, Las Palmas de Gran Canaria (Molina, in litt.
2009).

Crocidura panayensis Hutterer, 2007

Lynx (Praha) n.s. 38: 6, figs. 1-4

Holotype. ZFMK 2003.1091, adult female, specimen in
spirit, skull extracted; Sibaliw, Municipality of Buruanga
(11°49.188°N, 121°58.064’E), NW peninsula, Panay, Re-
public of the Philippines, collected by E. Curio, 13 May
2000; transferred to the NMP (now NMP 5207).
Paratypes. ZFMK 2003.1092 — 2003.1094 (1 male, 2 fe-
males in spirit, skulls extracted), same locality (450-480
m) and collector, obtained 2001—2003.

Bonn zoological Bulletin 59: 3-27

Current name. Crocidura panayensis Hutterer, 2007.
Comments. Status confirmed by Esselstyn et al. (2009).

Crocidura suaveolens balcanica Ondrias, 1971

Z. Saugetierkd. 35: 379

Paratypes. ZFMK 66.337, female, skin and skull; Sara-
jevo, Bosnia and Herzegovina, former Yugoslavia, collect-
ed by G. Witte, 26 June 1962, field no. 383; ZFMK
66.335, 66.336, 1 female, 1 male, skins and skulls; Ombla
Valley, Croatia, former Jugoslavia, collected by G. Witte,
7 and 9 June 1962, field nos. 371, 380.

Current name. Crocidura suaveolens mimula Miller,
1901; see Dubey et al. (2007).

Comments. The holotype was collected near Kryoneri-
on, Attica, Greece.

Crocidura sicula aegatensis Hutterer, 1991

Bonn. zool. Beitr. 42: 247

Holotype. ZFMK 89.392, adult male, skin and skull; Casa
Sarraceno, Marettimo, Egadi Islands, Italy, collected by
F. Krapp, 23 Mar 1969, field no. 35/69.

Paratypes. 6 specimens, listed in detail by Krapp (1970);
1 from Levanzo (ZFMK 89.393, skull), Favignana
(ZFMK 89.394—-397, 4 skulls, ZFMK 89.398, skin and
skull).

Current name. Crocidura sicula aegatensis Hutterer,
1991 (Hutterer 2005).

Crocidura sicula calypso Hutterer, 1991

Bonn. zool. Beitr. 42: 247

Holotype. ZFMK 89.418, adult male, specimen preserved
in spirit and extracted skull; Is-Saqwi, Xlendi Valley, Go-
zo (Malta), collected by S.Schembri and D.M. Johnson,
22 Oct 1986.

Paratypes. ZFMK 89.400, specimen in spirit, skull re-
moved, same data as holotype; ZFMK 89.401-417, 2 un-
catalogued, 20 skulls from owl pellets, Gozo, collected by
J. Sultana, Oct 1981.

Current name. Crocidura sicula calypso Hutterer, 1991
(Hutterer 2005).

Comments. Further paratypes are in the collection of P.
J. Schembri, Malta, and in IZEA.

Crocidura suaveolens bruecheri Lehmann, 1977

Ric. Biol. Selvag., Suppl., 5: 198, fig. 2 (May)
Holotype. ZFMK 77.665, male, skin and skull; Tiriolo,
Catanzaro, Calabria, Italy, collected by E. von Lehmann,
6 Mar 1977, field no. 18.

Paratypes. 12 skins and skulls from Catanzaro Province:
3 males, 4 females, ZFMK 777.664, 77.666-77.671, Tiro-
lo; 3 males, ZFMK 77.661-77.663, Marcellinara; male,
ZFMK 77.660, Catanzaro; male, ZFMK 77.672, near
Petrizia; all collected by E. von Lehmann, Mar 1977.
Current name. Crocidura suaveolens mimula Miller,
1901; see Dubey et al. (1997).

©ZFMK

8 Rainer Hutterer & Gustav Peters

Comments. Described as C. s.

ESA):

brticheri (Lehmann

Crocidura tansaniana Hutterer, 1986

Bonn. zool. Beitr. 37: 27, fig. 3c

Holotype. ZFMK 85.194, adult male, skin and skull;
Amani, E Usambara Mountains, Tanga, Tanzania, collect-
ed by S.R.Telford, 17 June 1984, field no. SRT-TZ-12078.
Current name. Crocidura tansaniana Hutterer, 1986.

Crocidura telfordi Hutterer, 1986

Bonn. zool. Beitr. 37: 28, fig. 3d

Holotype. ZFMK 85.195, adult male, skin and skull;
Morningside, 1150m, Uluguru Mountains, Morogoro, Tan-
zania, collected by S.R.Telford, 17 Nov 1983, field no.
SRT-TZ-4806.

Current name. Crocidura telfordi Hutterer, 1986.
Comments. A further skin without skull (SRT-TZ-2397)
from Bondwa Peak, Uluguru Mountains, was mentioned
in the original description (Hutterer 1986) but excluded
from the type material.

Crocidura yankariensis Hutterer & Jenkins, 1980

Bull. Brit. Mus. (Nat. Hist.) Zool. 39: 305, figs. 1-7 (18
Dec)

Paratype. ZFMK 80.805, skull without occipital, with
both mandibles, from pellet; Futuk (9°50’N, 10°55’E),
Bauchi State, Nigeria, collected by A. Demeter, 7 Aug
1978, field no. 55.

Current name. Crocidura yankariensis Hutterer & Jen-
kins, 1980.

Comments. Holotype in HNHM.

Myosorex eisentrauti Heim de Balsac, 1968

Bonn. zool. Beitr. 19: 20, figs. 1-6 (10 Apr)

Holotype. ZFMK 69.372, male, skin and skull; Pico de
Santa Isabel, 2400 m, Bioko, Equatorial Guinea, collect-
ed by M. Eisentraut, 14 Oct 1966, field no. 41.
Paratypes. ZFMK 69.373, female, skin and skull, same
data as holotype, field no. 40; ZFMK 69.374, male, skin
and skull, Refugium, 200 m, 5 Oct 1966, field no. 2;
ZFMK, without catalogue no. (field no. 9), skull (skin giv-
en to Pater Basilio), Refugium, Bioko, 6 Oct 166, all col-
lected by M. Eisentraut.

Current name. Myosorex eisentrauti Heim de Balsac,
1968 (Hutterer 2005).

Myosorex eisentrauti okuensis Heim de Balsac, 1968
Bonn. zool. Beitr. 19: 24, figs. 4, 5 (10 Apr)

Holotype. ZFMK 69.376, male, skin and skull; Lake Oku,
Lager IV, 2100 m, Banso Highlands, Cameroon, collect-
ed by M. Eisentraut, 25 Jan 1967, field no. 410.
Paratypes. 11 skins and skulls, ZFMK 69.377-69.389,
Lake Oku (field nos. 460, 446, 377, 346, 461, 376, 340,

Bonn zoological Bulletin 59: 3—27

345, 416); Lake Manenguba (field nos. 277, 320), all col-
lected by M. Eisentraut in Jan and Feb 1967.

Current name. Myosorex okuensis Heim de Balsac, 1968
(Hutterer 2005).

Comments. Paratype ZFMK 69.389 (field no. 447) was
given to the Transvaal Museum Pretoria in Aug 1981. Ad-
ditional paratypes were in the working collection of H.
Heim de Balsac and subsequently passed to MNHN.

Myosorex eisentrauti rumpii Heim de Balsac, 1968
Bonn. zool. Beitr. 19: 27, figs. 4-6 (10 Apr)

Holotype. ZFMK 69.375, male, skin and skull; Lager V
near Dikume, 1100 m, Rumpi Hills, Cameroon, collect-
ed by M. Eisentraut, 28 Feb 1967, field no. 568.
Current name. Myosorex rumpii Heim de Balsac, 1968
(Hutterer 2005).

Myosorex kihaulei Stanley & Hutterer, 2000

Bonn. zool. Beitr. 49: 20, figs. 1, 2 (30 Dec)

Paratype. ZFMK 2009.633, female, skin and skull; 4 km
W 5 km N Chita, 1460 m, 8°28°30’S, 35°54’25”E, Ud-
zungwa Scarp Forest Reserve, Udzungwa Mts, Kilombe-
ri District, Morogoro Region, Tanzania, collected by W.T.
Stanley, 14 Aug 1995, field no. WTS 1877. Formerly
FMNH 155611.

Current name. Myosorex kihaulei Stanley & Hutterer,
2000.

Neomys anomalus rhenanus Lehmann, 1976

Bonn. zool. Beitr. 27: 160 (Oct)

Holotype. ZFMK 71.62, male, skin, skull lost; Firnbach
near Datzeroth, Westerwald, Germany, collected by E. von
Lehmann, 6 Apr 1971, field no. 360.

Paratypes. ZFMK 67.126, male, skin and skull, Dasburg,
Eifel, Germany, collected by J. Olert, 9 July 1967; ZFMK
61.440, 62.116, 70.110, 70.111, 4 skins and skulls, Roten-
boden, Liechtenstein, collected by E. von Lehmann, 27
July 1961, 25 Aug 1962, 23 May 1970; ZFMK 71.334,
71.335, 2 skins and skulls, Gamander, Liechtenstein, col-
lected by J. Olert, 5 Nov 1971; in addition several skulls
from pellets (not catalogued).

Current name. Neomys anomalus Cabrera, 1907.
Comments. Further paratypes from the private collection
of W. Issel, Augsburg, are now in SMNS.

Neomys fodiens niethammeri Buhler, 1963

Bonn. zool. Beitr. 14: 169

Neotype. ZFMK 94.525, female, skin, skull and skeleton;
islet below karstic spring, Ramales de la Victoria,
Cantabria, N Spain, collected by P. Buhler, 18-19 Sept
1970 (in captivity until 28 Dec 1970).

Current name. Neomys fodiens niethammeri Bihler,
1963.

Comments. Bihler (1996: 312) designated a neotype be-
cause the holotype, an isolated mandible, was lost. The

©OZFMK

Types in ZFMK: Section Theriology 9

taxon may eventually deserve species status (LOpez-Fuster
et al. 1990).

Paracrocidura schoutedeni camerunensis Heim de Bal-
sac, 1960

Bonn. Zool. Beitr. 10: 204, fig. 3 (10 Jan)

Holotype. ZFMK 61.760, male, skin and skull; Lager V,
ca. 600 m, above Mueli, N side of Mount Cameroon,
Cameroon, collected by M. Eisentraut, 7 Feb 1958, field
no. 568.

Current name. Paracrocidura camerunensis Heim de
Balsac, 1960 (Hutterer, unpubl.).

Sorex araneus hilleri Lehmann, 1966

Saugetierkundl. Mitt. 14: 131, fig. 2

Holotype. ZFMK 64.169, male, skin and_ skull;
Naturschutzgebiet Elmpter Schwalmbruch, Kreis Erke-
lenz, Bezirk Aachen, North Rhine-Westphalia, Germany,
collected by E. von Lehmann, 7 Apr 1964, field no. 29.
Paratypes. 6 males, 2 females (ZFMK 64.168, 64.170,
64.171, 64.577-580, 64.1193), same collector and local-
ity.

Current name. Sorex araneus huelleri Lehmann, 1966.
Comments. Two paratypes (ZFMK 64.1192, 64.1194)
were exchanged with NMW in May 1979.

Sorex araneus silanus Lehmann, 1961

Zool. Anz. 167: 214

Holotype. ZFMK 60.339, adult male, skin and skull;
Camigliatello Silano, Calabria, Italy, collected by E. von
Lehmann, 9 Aug 1960, field no. 71.

Paratypes. 4 males, 3 females, | indiv. sex undetermined,
ZFMK 60.336-60.338, 60.340-60.344, all same locality
and collector.

Current name. Sorex antinorii Bonaparte, 1840.
Comments. Taxonomy discussed in detail by Brinner et
al. (2002).

Sorex araneus wettsteini Bauer, 1960

Bonn. zool. Beitr. 11: 178 (30 Oct)

Paratypes. ZFMK 79.544 (male), 79.545 (female), skins
and skulls; siltation zone of Neusiedler See, Neusiedl am
See, Burgenland, Austria, collected by K. Bauer, 19 May
and 11 Apr 1951, field nos. KB 51/105, KB 51/44. Ob-
tained in exchange from NMW in 1979.

Current name. Sorex araneus wettsteini Bauer, 1960.

Sorex minutus becki Lehmann, 1963

Jb. Histor. Vereins Firstentum Liechtenstein 62: 187
Holotype. ZFMK 56.1004, male, skin and skull, Silum,
Liechtenstein, collected by E. von Lehmann and H.-J.
Kuhn, 11 Aug 1956.

Paratypes. ZFMK 49.43, male, skin and skull; Osterach
Valley, Allgau, Germany, 7 Sept 1949; ZFMK 50.144,
50.145, 2 males, skins and skulls, Bodman, Lake Con-

Bonn zoological Bulletin 59: 3-27

stance, Germany, 9 May 1949, all collected by G. Hein-
rich.
Current name. Sorex minutus Linnaeus, 1866.

Sorex etruscus Savi, 1822

Nuovo Giornale dé Letterati 1: 60, plate on p. 70
Syntypes. ZFMK 78.513, adult female, in spirit, skull re-
moved; Italy, earlier than 1846, sent to Géttingen Muse-
um by Paolo Savi (Berthold 1850: 13, “Gesch. des Hrn
Prof Savi” = donated by prof. Savi), where it was cata-
logued in 1846. Inscription on specimen label: “Crocidu-
ra etrusca (Savi) / 1846 / Savi’. ZFMK 78.514, juvenile,
complete specimen in spirit. Pisa, Italy, no date, sent to
Gottingen Museum by Paolo Savi; has 2 labels, a) one old
and faded, “Sorex etruscus ...”, b) a newer one, approx-
imately about 1850, “a. Crocidura etrusca / A.S. / Savi /
Pisa”.

Current name. Suncus etruscus (Savi, 1822).
Comments. Both specimens were detected on the occa-
sion of the take-over of parts of the mammal collection
of the old Géttingen Museum by ZFMK in 1978. The
meaning of “A.S.” on the label of the second specimen is
“Alte Sammlung” (=old collection). Therefore it already
was 1n the old collection when its first inventory was com-
piled about 1846.

Sylvisorex akaibei Mukinzi, Hutterer & Barriere, 2009
Mammalia 73: 130, figs. 2, 3

Paratype. ZFMK 2006.168 (R27986), young adult;
Masako Forest Reserve (00°36.304’N, 25°15.389’E), 388
m, Kisangani area, Tshopo district, NW Democratic Re-
public of the Congo, collected by I. Mukinzi and D.
Amundala, 2 Jun 2005.

Current name. Sy/visorex akaibei Mukinzi, Hutterer &
Barriére, 2009.

Comments. Holotype in RCMA.

Sylvisorex granti camerunensis Heim de Balsac, 1968
Bonn. zool. Beitr. 19: 35, figs. 10, 11 (10 Apr)
Holotype. ZFMK 69.358, male, skin and skull; Lager HI,
1800 m, Lake Manenguba, Cameroon, collected by M.
Eisentraut, 18 Dec 1966, field no. 271.

Paratypes. 6 skins and skulls, ZFMK 69.359, 69.361-
69.364, one specimen without catalogue no.; Lake Manen-
guba (field nos. 305, 295, 280), Dec 1966; Lake Oku, 2100
m (field nos. 354, 363, 375), Jan 1967, all collected by
M. Eisentraut.

Current name. Sy/visorex camerunensis Heim de Balsac,
1968; see Hutterer et al. (1992).

Sylvisorex morio isabellae Heim de Balsac, 1968

Bonn. zool. Beitr. 19: 31, figs. 8, 9 (10 Apr)

Holotype. ZFMK 69.365, male, skin and skull; Refugium,
2000 m, Bioko, Equatorial Guinea, collected by M. Eisen-
traut, 10 Nov 1966, field no. 32.

©ZFMK

10 Rainer Hutterer & Gustav Peters

Paratypes. 3 skins, 2 skulls, ZFMK 69.366, 69.367, one
specimen without catalogue no., Refugium (field no. 35),
Bioko, Oct 1966; Pico de St. Isabel, Bioko, 2400 m (field
no. 44), 14 Oct 1966, all collected by M. Eisentraut.
Current name. Sy/visorex isabellae Heim de Balsac,
1968; see Hutterer (2005).

Sylvisorex corbeti Hutterer & Montermann, 2009

Bonn. zool. Beitr. 56: 202, figs. 1, 2, 4, 5 (30 Sept)
Holotype. ZFMK 88.91, young adult female, flat skin and
skull; near Chappal Waddi (07°01 N, 11° 41’ E), 1900 m,
Gotel Mountains, SE Nigeria, collected by G. Nikolaus,
23 Mar 1988, field no. GN 60.

Current name. Sy/visorex corbeti Hutterer & Monter-
mann, 2009.

Comments. Previously identified as S. ollula by Hutter-
er et al. (1992).

Sylvisorex silvanorum Hutterer, Riegert & Sedlacek, 2009
Bonn. zool. Beitr. 56: 151, figs. 1-3 (30 Sept)
Holotype. ZFMK 74.430, young adult female, skin and
partial skull; near Lake Bambili, Mount Lefo, Bafut
Ngemba F. R. (5°55’ N, 10°14’ E), 1800-1900 m, Bamen-
da Highlands, NW Province, Cameroon, collected by W.
Bohme and W. Hartwig, 7 Mar 1974, field no. 98.
Paratypes. ZFMK 2003.1103, 2003.1104, 2 incomplete
skulls, removed from pellets of 7yto capensis; slope of the
lower Bambili volcano craters (5°55’ N, 10°14’ E), 2400
m, about 8 km SE city of Bamenda, Bamenda Highlands,
NW Province, Cameroon, collected by J. Riegert and O.
Sedlacek, Nov 2001.

Current name. Sy/visorex silvanorum Hutterer, Riegert
& Sedlacek, 2009.

Sylvisorex vulcanorum Hutterer & Verheyen, 1985

Z. Saugetierkd. 50: 267, figs. 1, 2

Paratype. ZFMK 85.193, specimen in spirit; Karisoke
(0°28’S, 29°29’E), 3100 m, Pare National des Volcans,
Rwanda, collected by W. Verheyen, 1982, field no. 3738.
Current name. Sy/visorex vulcanorum Hutterer & Ver-
heyen, 1985.

Talpidae

Talpa caeca steini Grulich, 1971

Acta Sc. Nat. Brno, Prague 5 (9): 38, figs. 1-4, 6-7, plate
II

Paratypes. ZFMK 66.307-66.316, 9 males, 1 female,
skins and skulls; Lovéen, Montenegro, former Yu-
goslavia, collected by G. Witte, 22-28 May 1962, field
nos. 350-360.

Current name. 7alpa caeca steini Grulich, 1971; see Hut-
terer (2005).

Bonn zoological Bulletin 59: 3—27

Talpa romana wittei Capolongo, 1986

Bonn. zool. Beitr. 37: 252

Holotype. ZFMK 66.298, male, skin and skull; Monte
Gargano, Apulia, Italy, collected by G. Witte, 28 July1961,
field no. 216.

Paratypes. ZFMK 66.291-297, 66.299-302; 9 males, 2
unsexed skins and skulls, all Monte Gargano, collected by
G. Witte, May—June 1061.

Current name. 7alpa romana wittei Capolongo, 1986; see
Hutterer (2005).

Talpa stankovici montenegrina KryStufek, 1994

Bonn. zool. Beitr. 45: 14

Paratypes. ZFMK 93.400 (male), 93.401 (female), flat
skins and skulls; sandy beach 5 m from the sea, Ulcinj,
Montenegro (former Yugoslavia), collected by B.
KryStufek & A. Stangelj, 14 Nov 1983, field nos. 3206,
3207.

Current name. Zalpa stankovici montenegrina Krystufek,
1994.

Comments. Donated by B. KryStufek 1993.

CHIROPTERA
Pteropodidae

Rousettus aegyptiacus occidentalis Eisentraut, 1960
Bonn. zool. Beitr. 10: 231, figs. 8, 13 (10 Jan)
Holotype. ZFMK 59.450, male, skin and skull; Mueli, ca.
600 m, N side of Mount Cameroon, Cameroon, collect-
ed by M. Eisentraut, 20 Feb 1958, field no. 648.
Paratypes. 28 skins and skulls, 11 specimens in spirit;
Lager V, above Mueli, 600 m (ZFMK 61.576, 61.578,
61.582, 61.584-61.586, 61.595-61.597, 61.600, 61.602),
Feb 1958; Lager I, above Buea, 1600 m, Mount Cameroon
(ZFMK 61.592, 61.593, 63.209a-f), Nov 1957; Lager VI,
Isobi, 30 m, W side of Mount Cameroon (ZFMK 61.577,
61.580, 61.581, 61.583, 61.587-61.591, 61.594,
61.598,61.599, 63.208a-e), Mar 1958; Lager IV, near Lake
Koto Barombi, 120 m, N side of Mount Cameroon
(ZFMK 61.575, 61.579, 61.601), Jan 1958; all collected
by M. Eisentraut.

Current name. Rousettus aegyptiacus unicolor Gray,
1870; see Koopman (1966).

Rhinolophidae

Hipposideros camerunensis Eisentraut, 1956

Zool. Jb., Syst. Okol. 84a: 526, figs. 6-8 (27 Dec)
Paratype. ZFMK 79.138, adult female; Buea, Cameroon,
collected by M. Eisentraut, 29 Apr 1954, field no. 506;
received in exchange from SMNS (SMNS catalogue no.
5196), Mar 1979.

©ZFMK

Types in ZFMK: Section Theriology 11

Current name. Hipposideros camerunensis Eisentraut,
1956.
Comments. Holotype and additional paratypes in SMNS.

Rhinolophus denti knorri Eisentraut, 1960

Stuttg. Beitr. Naturk. 39: 3, fig. 2 (1 June)

Paratypes. ZFMK 59.174, male, skin and skull; cave near
Nyembaro, Salung Plateau, 10 km W Kolenté, Guinea,
collected by H. Knorre, 15 Nov 1956, field no. 90; ZFMK
59.175, male, skin and skull, same locality and collector,
19 Nov 1956.

Current name. Rhinolophus (denti) knorri Eisentraut,
1960.

Comments. Holotype in SMNS.

Rhinolophus ferrumequinum martinoi Petrov, 1941
Zap. Russk. Nauch. Inst. Belgr.16: 59

Holotype. ZFMK 2002.458, female, skull; Pepeliste near
Krivolak, 400 m, 40 km SE Veles, S Serbia, collected by
B. Petrov, 9 Apr 1939, field no. BP 214.

Paratypes. ZFMK 98.078, female, skin and skull, same
data as holotype, field no. BP 217.

Current name. Rhinolophus ferrumequinum (Schreber,
1774), see Csorba et al. (2003).

Rhinolophus landeri guineensis Eisentraut, 1960
Stuttg. Beitr. Naturk. 39: 1, fig. 1 (1 June)

Paratypes. ZFMK 59.176, female, skin and skull; Tahiré,
bottom of Kelesi Plateau, Guinea, collected by H.
Knorre, 5 Oct 1956, field no. 29; ZFMK 59.177, male,
skin and skull, cave near Nyembaro, Kankuya Plateau, 8
km W Kolenté, Guinea, same collector, 23 Nov 1956, field
no. 137.

Current name. Rhinolophus guineensis Eisentraut, 1960;
see BOhme & Hutterer (1979).

Rhinolophus sumatranus K. Andersen, 1905

Proc. zool. Soc. London, 1905: 133 (16 May)
Paratype. ZFMK 78.526, male, complete specimen in
spirit; Sumatra, Dr. Lambrecht, without date; transferred
from Gottingen Museum (catalogue no. 61a) 1978.
Current name. Rhinolophus acuminatus sumatranus K.
Andersen, 1905; see Medway (1977: 48).

Rhinolophus ziama Fahr, Vierhaus, Hutterer & Kock,
2004

Myotis 40: 109, figs. 1-7

Holotype. ZFMK 99.934, adult male, specimen preserved
in spirit, skull and parts of skeleton; Reserve de la Bios-
phere du Massif du Ziama, western edge of Seredou,
Guinée Foresti¢re, Guinea, collected by H. Vierhaus, 12
Aug 1992, field no.HV 2590.

Current name. Rhinolophus ziama Fahr, Vierhaus, Hut-
terer & Kock, 2004.

Bonn zoological Bulletin 59: 3-27

Megadermatidae

Megaderma gigas Dobson, 1880

Proc. zool. Soc. London, 1880: 461, plate XLVI (15 June)
Holotype. ZFMK 78.525, specimen in spirit, skull extract-
ed; Wilson’s River, Mount Margaret, Central Queensland,
Australia, collected by Mr. Wilson, sent to Géttingen Mu-
seum by Dr. Schiitte 1879. Transferred to ZFMK in 1978;
Gottingen Museum catalogue no. 102a.

Current name. Macroderma gigas (Dobson, 1880).
Comments. When retrieved in 1978 the skull of the holo-
type was dissected on one side only. Therefore Dobson
must have obtained his figure by mirroring this side of the
skull. The only partial dissection of the skull also explains
the fact that in the original description the skull figured
has no nasal fossa. The skull was cleaned in 1979.

Phyllostomidae

Platyrrhinus chocoensis Alberico & Velasco, 1991
Bonn. zool. Beitr. 42: 238

Paratype. ZFMK 89.250, male, skin and skull; Conce-
sion Bajo Calima pulpopel, Municipio Buenaventura,
Valle, Colombia, collected by J.H. Gamba, 2 Mar 1986,
field no. 129, donated by M. Alberico in 1989.

Current name. Platyrrhinus chocoensis Alberico & Ve-
lasco, 1991.

Comments. Further paratypes in ZMB (Turni & Kock
2008), FMNH, MNHN and SMNS.

Molossidae

Dinops Cestoni Savi, 1825

Nuovo Giornale de’ Letterati, Pisa, 10: 235

Syntype. ZFMK 83.173, male, complete specimen in spir-
it; Tuscany, Italy. Label: “Nyctinomus cestonii(Savi)Dob-
son (In der Sammlg. Von Berthold als Dysopes Savii
angegeben) Prof. Savi ded. 1846 Toscana”. Transferred
to ZFMK from Gottingen Museum 1978.

Current name. Zadarida teniotis (Rafinesque, 1814).
Comments. Synonym of 7adarida teniotis. The specimen
was sent to the Géttingen Museum by P. Savi. The year
specification 1846 refers to the year of inventory taking
in Gottingen (cf. Berthold 1850), the specimen was col-
lected before. Additional specimens that can also be con-
sidered to represent syntypes are in the museums of Pisa,
Vienna, and London.

Vespertilionidae

Amblyotus atratus Kolenati, 1858
Sber. Math.-nat. Cl. kais. Akad. Wiss. 29: 252

©ZFMK

Syntype(?). ZFMK 77.1035, male, in spirit, skull extract-
ed; label 1: “Amblyotus atratus (Kolenati) / (Meteorus) /
Altvater Mountains”, label 2: “‘Vesperus atratus Kuhl / Alt-
vater /107a/ C. Koch”.

Current name. Epfesicus nilssoni (Keyserling & Blasius,
1839), see Ellerman & Morrison-Scott (1951).
Comments. The ZFMK specimen originates from the bat
collection of the renowned chiroperologist Carl Koch who
mainly collected in Hesse and Westphalia but also ex-
changed bats from all over the world or received them
from contemporaries. Koch very probably received the
specimen now in Bonn from his contemporary Kolenati.
This assumption is supported by the label entries of Kole-
nati’s genus (Amblyotus), subgenus (Meteorus) and
species (atratus) as well as the type locality “Altvater
Mountains”. C. Koch’s main work (Das Wesentliche der
Chiropteren mit besonderer Beschreibung der in dem Her-
zogtum Nassau und den angrenzenden Landestheilen vor-
kommenden Fledermause, 1862/63) contains a paragraph
on Meteorus atratus (Kolenati) on page 393 which docu-
ments that Koch saw a specimen. In addition he points out
the similarity of M. atratus with V. nilssoni.

Plecotus alpinus Kiefer & Veith, 2002

Myotis 39: 8, figs. 2-4 (Apr 2002)

Holotype. ZFMK 2001.325, male, skin and skull; Risto-
las (06°57’N 44°46’E), Haute Alpes, France, 1600m, col-
lected by P. Favre, 24 Aug 2001.

Paratypes. ZFMK 61.451, adult female, skin and skull,
Schaan (51°06’N, 06°28’E), Liechtenstein, collected by
E. von Lehmann, 23 Aug 1961; ZFMK 2001.237, adult
female, skin and skull, Fischertratten (46°56’N, 13°31’E),
768 m, Austria, collected by G. Reiter, 15 May 2000;
ZFMK 2001.328, juvenile male, dry skin with skull, Du-
vin (46°43’N, 09°12’E), 1080 m, Switzerland, collected
by M. Lutz, 29 Aug 2000.

Current name. Plecotus macrobullaris alpinus Kiefer &
Veith, 2002.

Comments. See Kock (2002) for a discussion of the pub-
lication date.

Plecotus auritus hispanicus Bauer, 1957

Bonn. zool. Beitr. 7: 312 (31 Jan)

Holotype. ZFMK 46.292, female, skin and skull; Lagu-
nilla, Bejar, Salamanca, Spain, collected by H. Griin, 7 Ju-
ly 1940.

Paratypes. 8 skins and skulls, 1 specimen in spirit; ZFMK
34.19 (Lagunilla, 3 Feb 1934), 46.288 - 46.291, 46. 293
(same locality, 6-10 July 1940), 35.73 (Linares de Riofrio,
2 May 1935), 53.36 (same locality, 3 Feb 1953), 30.66
(specimen in spirit, Mosqueruela, 3 Oct 1930), all collect-
ed by H. Griin.

Current name. Plecotus austriacus hispanicus Bauer,
1957; see Bauer (1960: 141-144).

Bonn zoological Bulletin 59: 3-27

12 Rainer Hutterer & Gustav Peters

Pipistrellus eisentrauti Hill, 1968

Bonn. zool. Beitr. 19: 45 (10 Apr)

Holotype. ZFMK 68.5, adult male, skin and skull; Lager
V, Dikume-Balue, Rumpi Hills, Cameroon, collected by
M. Eisentraut, 18 Feb 1967, field no. 498.

Paratype. ZFMK 68.6, adult female, skin and skull; Lager
II, ca. 1100 m, Mount Kupe, Cameroon, same collector,
30 Nov 1966, field no. 198.

Current name. Hypsugo eisentrauti (Hill, 1968).
Comments. Another paratype was given to BMNH.

Vespertilio Bonapartii Savi, 1838

Nuovo Giornale de’ Letterati, Pisa, 37: 226

Syntype. ZFMK 83.172, male complete specimen in spi-
rit; Tuscany, Italy. Label: “Vesperugo maurus Blas. (fe-
ste Dobson 1880. Vgl. Proc. Zool. Soc. Lond. June, 1880)
in der alten Samlg von Berthold als Vespertilio Savii be-
zeichnet. A.S. 1847. Toscana.” Transferred from G6ttin-
gen Museum to Museum Alexander Koenig in 1978.
Current name. Hypsugo savii (Bonaparte, 1837).
Comments. This specimen of Savi’s pipistrelle was sent
to the Géttingen Museum by Paolo Savi, as noted by
Berthold (1850) and was subsequently discussed by Dob-
son (1880). Vesperugo maurus Blasius, 1853 is another
synonym of H. savii; syntypes of this taxon are preserved
in the Natural History Museum of Braunschweig (Hev-
ers 2005).

CARNIVORA
Felidae

Felis catus jordansi Schwarz, 1930

Zool. Anz. 91: 223

Holotype. ZFMK 83.186, male, skin and skull (occiput
damaged); Santa Margarita, Majorca, Spain, collected by
J. Parpal, 13 Jan 1929.

Paratype. ZFMK 83.187, skin, in poor condition; San
Juan, Majorca, same collector, 9 June 1930.

Current name. Felis silvestris jordansi Schwarz, 1930.
Comments. Skull and skin of holotype are figured in Hal-
tenorth (1953). The paratype may be a domestic cat.

Viverridae

Genetta bourloni Gaubert, 2003

Mammalia 67: 95, figs. 4-6

Paratype. ZFMK 95.10, skin; Ziama Forest, Gboda, SE
Guinea, obtained from local people by W. Biitzler,
1993/94.

Current name. Genetta bourloni Gaubert, 2003.

©ZFMK

Types in ZFMK: Section Theriology 13

Genetta (Paragenetta) lehmanni Kuhn, 1960
Saugetierkundl. Mitt. 8: 154-160, figs. 1, 4, 5, 7, 9-12
Holotype. ZFMK 57.11, skull of an adult individual, sex
unknown; Kpeaple (6°36°N, 8°30’ W), Liberia, Feb 1956,
ex H. Himmelheber collection.

Paratype. ZFMK 61.966, skull of an adult individual, sex
unknown, same locality and collector, 1959; formerly in
the possession of Dr. H. Himmelheber.

Current name. Genetta johnstoni Pocock, 1908.
Comments. Synonym of Genetta johnstoni Pocock, 1908
(Kuhn 1965: 332). Genetta lehmanni 1s the type species
of the subgenus Paragenetta Kuhn, 1960.

Herpestidae

Crossarchus platycephalus Goldman, 1984

Can. J. Zool. 62: 1624

Paratypes. 6 skins, 6 skulls (2 damaged); ZFMK 61.739
(field no. 701), adult male, ZFMK 61.741 (field no. 700),
adult female, collected by M. Eisentraut, Lager VI, Iso-
bi, near Bibundi, W side Mount Cameroon, Cameroon, 5
Mar 1958, 30 m; ZFMK 61.742 (field no. 719), adult,
same locality and collector, 7 Mar 1958; ZFMK 61.740
(field no. 657), ZFMK 61.743 (field no. 563), adult males,
Lager V, above Mueli, 600 m, N side Mount Cameroon,
Cameroon, same collector, 21 Feb and 6 Feb 1958; ZFMK
69.150 (field no. 578), Lager V, Dikume, 1100 m, Rumpi
Hills, W Cameroon, same collector, 1 Mar 1967.
Current name. Crossarchus platycephalus Goldman,
1994.

Liberiictis Kuhni Hayman, 1958

Ann. Mag. nat. Hist. (13) 1: 449, fig. 1

Paratypes. 6 skulls, ZFMK 58.21 — 58.26; Gaplay (7°8’N,
8°28’ W), NE-Liberia, Jan 1957, ex H. Himmelheber col-
lection.

Current name. Liberiictis kuhni Hayman, 1958.
Comments. The original series consisted of 8 skulls. The
holotype is BMNH 58.507 and one paratype was in the
collection of the Department of Biogeography, Saarbriick-
en University.

Canidae

Vulpes vulpes toschii Lehmann, 1969

Z. Jagdwiss. 15: 28-31, figs. 14

Holotype. ZFMK 66.487, adult female, flat skin and skull;
Calvo area, Monte Gargano, Apulia, Italy, collected by M.
Facciorusso, 13 Oct 1961, collection G. Witte.
Paratypes. 4 skulls, 2 flat skins. ZFMK 56.741, male, flat
skin and skull, Foresta Umbra, Monte Gargano, collect-
ed by B. Hagen, 12 Sept 1955; ZFMK 56.742, skull,
humerus and femur, same locality and collector; ZFMK

Bonn zoological Bulletin 59: 3-27

66.488, female, skull, Calvo area, Monte Gargano,
800-900 m, collected by M. Facciorusso, June 1961, col-
lection G. Witte no. 214; ZFMK 66.489, damaged skull,
Foresta Umbra, Monte Gargano, collected by La Torre,
June 1961, collection G. Witte no. 243; ZFMK 68.205,
male, flat skin, Lungro, Calabria, Italy, through V. Skan-
derberg, winter 1967/68, collection E. von Lehmann.

Current name. Vulpes vulpes crucigera Bechstein, 1789.

ARTIODACTYLA
Cervidae

Alces pfizenmayeri Zukowsky, 1910

Wild und Hund 16, 45: 808.

Material. ZFMK 2009.632, a possible syntype or para-
lectotype; skull of adult male, junction of Aldan and Lena
River, Yakutsk Province, Russia; collected by E. Pfizen-
mayer (Tiblisi) in Nov 1909.

Current name. Alces americanus (Clinton, 1822).
Comments. Pfizenmayer (1910) published an account on
a large form of moose from northern Siberia in which he
figured five male skulls from the Aldan Range. Zukowsky
(1910: 808) subsequently commented on this article and
named the moose Alces pfizenmayeri. In 1914 he added
a more detailed account in which he fixed the skull shown
in Pfizenmayer’s (1910) figure 6 as the type; this skull had
been donated to the Berlin Museum and catalogued un-
der number A.19.09 (Zukowsky 1914: 38). A letter from
E. Pfizenmayer to Alexander Koenig dated 15 June 1911
(ZFMK Archives) tells that he sold other skulls from that
series to the museums of Vienna, Washington, Stockholm,
and Bonn; one he kept for himself. Zukowsky’s (1914) ac-
tion of fixing the Berlin specimen as the type can be tak-
en as a lectotype designation; the remaining specimens
will then be paralectotypes. The Bonn specimen was not
figured in the paper of Pfizenmayer (1910); its status is
therefore questionable. It is, however, one of the original
specimens obtained by Pfizenmayer from the type local-
ity. Grubb (2005) included the name as a synonym in A/-
ces americanus. Zukowsky’s name, however, is the ear-
liest available name for the Palaearctic range of the Amer-
ican Moose and is sometimes applied as technical name
for the Yakutsk Moose (Franzmann 1981).

Dama dama geiselana Pfeiffer, 1998

Eiszeitalter u. Gegenwart 48: 75, figs. 1C, 2, 6
Material. ZFMK 2009.629, cast of right antler of holo-
type (Landesmuseum fiir Vorgeschichte Halle, HK
97:14165 NN17); Pleistocene lake sediments of Neumark-
Nord, Geiseltal, Sachsen-Anhalt, Germany.

Current name. Dama dama geiselana Pfeiffer, 1998.
Comments. Cast taken from original by ZFMK staff.

©ZFMK

14 Rainer Hutterer & Gustav Peters

RODENTIA
Sciuridae

Sciurus ingrami sebastiani Miller & Vesmanis, 1971
Senckenberg. biol. 52: 378

Holotype. ZFMK 2009.622 (formerly Univ. Saarbrtick-
en 0535), adult male, skin and skull; rainforest above the
electric power station, 500 m, San Sebastiao Island
(23°50’S, 45°20’ W), Sao Paulo State, Brazil, collected by
E. & P. Miiller, E.B. Bohaler & F. Silva, Oct 1964, field
no. 234.

Paratypes. ZFMK 2009.623 - 2009.628, 6 skins, 3 skulls,
same locality and collectors, 1964-1967.

Current name. This taxon was overlooked by Thoring-
ton & Hoffmann (2005), and its status therefore not eval-
uated. They included ingrami as a subspecies in Sciurus
aestuans, however, Miller & Vesmanis (1971) demon-
strated that sebastiani was larger in cranial measurements
than ingrami from the mainland, and that its body coloura-
tion was darker. Here we tentatively accept the taxon as
a valid island form as Sciurus aestuans sebastiani Miller
& Vesmanis, 1971.

Comments. Holotype and paratypes were initially stored
at the Department of Biogeography at Saarbriicken Uni-
versity, then transferred to the University of Trier, and fi-
nally donated to the ZFMK in 2009. Two further
paratypes are in SMF (SMF 40557-8).

Sciurus vulgaris hoffmanni Valverde, 1967

Bol. R. Soc. Esp. Hist. Nat., Biol. 65: 225

Paratype. ZFMK 92.518, adult male, skin and skull; La
Perdiz, Sierra de Espuna, Alhama de Murcia, Spain, col-
lected by J. A. Valverde, 25 Apr 1967, field no. 979.
Current name. Sciurus vulgaris hoffmanni Valverde,
1967 is antedated by Sciurus aestuans hoffmanni Peters,
1864, a Central American squirrel (Peters 1864, Elliot
1904). Both homonyms were listed by Thorington & Hoff-
mann (2005) without comment. A replacement name is
therefore required for the strikingly large and pale squir-
rel of the Sierra de Espuna. It shall be further on known
as Sciurus vulgaris valverdei Hutterer nom. nov., in ho-
nour of its describer, J. A. Valverde.

Comments. Holotype is no. 985 in the Estacion Biolég-
ica de Donana, Sevilla. Measurements of paratype 979
were included in Valverde (1967); the specimen was do-
nated by J. A. Valverde to J. Niethammer and remained
in his private collection until it was passed to the ZFMK
in 1992.

Gliridae

Dryomys nitedula aspromontis Lehmann, 1964
Sitzber. Ges. naturf. Freude Berl. (N.F.) 4: 33, fig. 2

Bonn zoological Bulletin 59: 3—27

Holotype. ZFMK 63.510, adult male, skin and skull;
Gambarie d’ Aspromonte, Calabria, Italy, collected by E.
von Lehmann, 13 Aug 1963, field no. 92.

Paratypes. ZFMK 63.508, 63.509, 63.511 — 63.513, 3
males, 2 females, skins and skulls, same locality and col-
lector as holotype, 12-13 Aug 1963, field nos. 85-86,
93-95.

Current name. Dryomys nitedula (Pallas, 1778).

Dryomys nitedula diamesus Lehmann, 1959

Bonn. zool. Beitr. 10: 4 (7 Sept)

Holotype. ZFMK 58.211, male, skin and skull; Ivanova
Kurita, Montenegro, former Yugoslavia, collected by E.
von Lehmann, 16 Sept 1958, field no. 28.

Paratypes. ZFMK 58.210, 58.212, | male, | female, skins
and skulls, same locality and collector as holotype, 14-15
Sept 1958, field nos. 16, 23.

Current name. Dryomys nitedula (Pallas, 1778).
Comments. KryStufek (1985) discussed the status of the
species in former Yugoslavia.

Muscardinus avellanarius abanticus Kivanc, 1983
Bonn. zool. Beitr. 34: 421, figs. 2, 3 (14 Dec)
Paratypes. ZFMK 83.127, 83.128, females, skins and
skulls; Abant, Vil. Bolu, Turkey, collected by E. Kivane,
6 Aug 1980 and 5 July 1981, field nos. EK 84, EK63.
Current name. Muscardinus avellanarius abanticus Ki-
vanc, 1983.

Comments. Holotype in the collection of the Biology De-
partment in the Natural Sciences Faculty of Ankara Univ.

Muscardinus avellanarius kroecki G. Niethammer &
Bohmann, 1950

Neue Ergebnisse und Probleme der Zoologie (Klatt-Fest-
schrift), Leipzig, p. 661

Holotype. ZFMK 74.272, unsexed skin; Vitosa, 1500 m,
10 km S Sofia, Bulgaria, collected by A. Kroeck, 26 Aug
1944, field no. 81, at the same time no. 173/44 of the
Bohmann collection. Donated to Museum Koenig by
Bohmann in Oct 1974.

Current name. Muscardinus a. avellanarius (Storch
1978).

Nesomyidae

Petromyscus collinus kurzi Lehmann, 1955

Bonn. zool. Beitr. 6: 171 (31 Dec)

Holotype. ZFMK 55.75, male, skin and skull; Renoster-
berg, Kaoko-veld, Namibia, collected by W. Hoesch, 26
Mar 1955, field no. 1256.

Paratypes. ZFMK 55.79, 55.455, 55.456, skins and skulls,
same locality, collector and date as holotype.

Current name. Petromyscus collinus (Thomas & Hinton,
1925), see Musser & Carleton (2005).

©ZFMK

Types in ZFMK: Section Theriology 15

Cricetidae

Clethrionomys glareolus curcio Lehmann, 1961

Zool. Anz. 167: 219

Holotype. ZFMK 60.352, adult female, skin and skull;
Camigliatello Silano, Calabria, Italy, collected by E. von
Lehmann, 2 Aug 1960, field no. 29.

Paratypes. ZFMK 60.349 - 60.351, 60.353, 60.354,
60.356 - 60.366, 16 skins and skulls, same locality and
collector, July/Aug 1960.

Current name. Myodes glareolus (Schreber, 1780); see
Musser & Carleton (2005).

Clethrionomys glareolus garganicus Hagen, 1958

Z. Saugetierkd. 23: 54, fig. 1

Holotype. ZFMK 58.128, adult male, skin and skull;
Foresta Umbra, 800 m, Monte Gargano, Apulia, Italy, col-
lected by B. Hagen, 28 Aug 1955, field no. 24.
Paratypes. ZFMK 58.123 — 58.127, 58.129 — 58.143, 20
skins and skulls, same locality, collector and date, field
nos. 8, 19, 20, 22 — 26, 28 — 30, 32 — 42.

Current name. Myodes glareolus (Schreber, 1780); see
Musser & Carleton (2005).

Comments. Hagen (1958: 56, table 2) lists 24 specimens
in her description of which 21 were in the ZFMK collec-
tion in Apr 1979. The collection date of the holotype was
given by Hagen (1958: 55) as “July” but “VIII. 1955” is
recorded on its label; likewise all paratypes were collect-
ed in Aug 1955.

Clethrionomys glareolus pirinus Wolf, 1940

Mitt. K6nigl. Naturwiss. Inst. Sofia 13: 158 (20 Mar)
Holotype. ZFMK 39.91, adult female, skin and (damaged)
skull; Banderiza Hiitte, 1800 m, Pirin Mountains, Bulgar-
ia, collected by H. Wolf, 11 June 1938.

Paratypes. ZFMK 39.92 — 39.96, 5 skins and skulls, Ban-
deriza Hiitte and (ZFMK 39.96) Demianiza Valley, 1150
m, same collector, 9-13 June 1938.

Current name. Myodes glareolus (Schreber, 1780); see
Musser & Carleton (2005).

Pitymys bavaricus Konig, 1962

Senckenberg. biol. 43: 2, figs. 1, 2, 6, 8, 13

Paratypes. ZFMK 79.142, male, skin and_ skull;
Garmisch-Partenkirchen, Bavaria, Germany, collected by
C. Konig, Nov 1961, field no. 1149, received in exchange
from SMNS; ZFMK 2009.630, male, skin and skull; same
locality and collector, 10 Nov 1959, field no. 1194, ex
Coll. J. Niethammer.

Current name. Microtus bavaricus (Konig, 1962).
Comments. The German subpopulation is probably ex-
tinct.

Pitymys savii brachycercus Lehmann, 1961
Zool. Anz. 167: 223

Bonn zoological Bulletin 59: 3-27

Holotype. ZFMK 60.379, male, skin and skull; Camigli-
atello Silano, Calabria, Italy, collected by E. von Lehmann,
27 July 1960.

Paratypes. ZFMK 60.380 — 60.385, 6 skins and skulls,
same locality, collector, and date as in holotype.
Current name. Microtus brachycercus (Lehmann, 1961).
Comments. Elevated to species by Galleni et al. (1994).

Microtus anatolicus Krystufek & Kefelioglu, 2002
Bonn. zool. Beitr. 50: 8, figs. 2-4

Holotype. ZFMK 99.926, adult male, skin (tail missing)
and skull; Yapali Ko6yti, Cihanbeyli, Konya, Turkey, col-
lected by H. Kefelioglu, 20 July 1993.

Current name. Microtus anatolicus KryStufek & Kefe-
lioglu, 2002.

Comments. 12 paratypes were deposited in the Ondokuz
Mayis University at Samsun, Turkey.

Microtus (Terricola) savii niethammericus Contoli,
2003

Hystrix, Italian Journal of Mammalogy (n.s.) 14: 109
Lectotype. Here designated: ZFMK 2003.928, adult fe-
male (6 teats visible), flat skin and skull; Foresta Umbra,
12 km in direction Monte San Angelo (41.13N 15.58E),
750 m, Monte Gargano, Italy, collected by J. Niethammer,
23 Mar 1980; field no. 6124.

Paralectotypes. 19 skins and 20 skulls (11 females, 9
males), all from type locality Foresta Umbra between 700
and 750 m in 1980; ZFMK 2003.916-2003.927, 2003.929-
2003.936.

Current name. Microtus savii (de Selys-Langchamps,
1838); see Musser & Carleton (2005).

Comments. Contoli (2000: 40) named this taxon Micro-
tus (Terricola) savii niethammeri, a nomen nudum and
homonym of Microtus (Terricola) multiplex niethammeri
Brunet-Lecompte & Volobouev, 1994. In 2003, he re-
named it niethammericus and provided a short description
(3 pairs of teats in females, X chromosome acrocentric)
and type locality (Foresta Umbra), based on the work of
Niethammer (1981). He also listed a series of 21 syntypes
(ZFMK 2003.916 — 2003.936), based on the paper of Ni-
ethammer (1981) and on correspondence with one of us
(RH). No holotype was fixed. We selected a specimen as
the lectotype which exhibits one of the diagnostic traits
(3 pairs of teats), and which has a skull in good condi-
tion. Vouchers for the karyotype of 2n = 54 (1 male, | fe-
male; Niethammer 1981: 46, fig. 1) are among the para-
lectotypes.

Nesoryzomys fernandinae Hutterer & Hirsch, 1980
Bonn. zool. Beitr. 30: 276, fig. 2 (28 Dec)

Holotype. ZFMK 79.370, skull of an adult individual, in-
terparietale missing, tip of nasalia damaged, from ow! pel-
lets; Fernandina Island, 300 m, Galapagos Islands,
Ecuador, collected by U. Hirsch, June 1979.

©ZFMK

16 Rainer Hutterer & Gustav Peters

Paratypes. ZFMK 79.371 — 79.406, 36 skulls, ex pellets,
same locality, collector and date as holotype; further
paratypes deposited in AMNH, BMNH, FMNH and
Charles Darwin Station, Galapagos Islands.

Current name. Nesoryzomys fernandinae Hutterer &
Hirsch, 1980.

Comments. The species was recently collected alive
(Dowler et al. 2000).

Muridae

Acomys cilicius Spitzenberger, 1978

Ann. naturhist. Mus. Wien 81: 444, figs. lc, 2c, plate 1
(Feb)

Paratypes. ZFMK 68.265, male, skin and skull; 20 km
E Silifke, Turkey, collected by H. Mittendorf, 22 July
1968, field no. 181; ZFMK 68.266, 68.267, females, skins
and skulls, same locality and collector, 24 July 1968, field
nos. 193, 194.

Current name. Acomys cilicius Spitzenberger, 1978.
Comments. Holotype in NMW.

Apodemus agrarius henrici Lehmann, 1970

Saugetierk. Mitt. 18: 154, fig. 1 (May)

Holotype. ZFMK 46.441, male, skin and skull; Bild-
hausen near Munnerstadt, Bad Kissingen District, Ger-
many, collected by G. Heinrich, 10 Aug 1946, field no.
2956.

Paratypes. ZFMK 46.412, male, skin and skull, same lo-
cality, collector and date as holotype.

Current name. Apodemus agrarius (Pallas, 1771).

Apodemus flavicollis alpinus Heinrich, 1951 — Apodemus
flavicollis alpicola Heinrich, 1952

Zool. Jb., Syst., 80: 114; J. Mammal. 33: 260
Paratypes. ZFMK 49.20 — 49.25, 4 males, 2 females,
skins and skulls; Osterach Valley, Allgau, Germany, col-
lected by G. Heinrich between | and 14 Aug 1949.
Current name. Apodemus alpicola Heinrich, 1952
Comments. As alpinus was preoccupied Heinrich as-
signed the name a/picola to the same form.

Apodemus tauricus geminae Lehmann, 1961

Zool. Anz. 167: 228

Holotype. ZFMK 58.149, adult female; Monte Gargano,
Apulia, Italy, coll. by B. Hagen, Aug 1955, field no. 7.
Paratypes. ZFMK 58.144, 58.145, 58.148, 58.150 —
58.163, 58.156 — 58.164, 15 skins and skulls, same local-
ity, collector and date as holotype.

Current name. Apodemus flavicollis (Melchior, 1834);
see Musser & Carleton (2005).

Comments. According to the locality data published by
Hagen (1958) the terra typica can be specified as “Fores-
ta Umbra, Monte Gargano”.

Bonn zoological Bulletin 59: 3-27

Apodemus flavicollis sicilianus Lehmann & Schaefer,
1978

Ric. Biol. Selvag., Suppl. 5: 180, figs. 3, 4 (Nov)
Holotype. ZFMK 73.256, adult female, skin and skull;
higher range, Mount Etna, ca. 1600 m, Sicily, Italy, col-
lected by E. von Lehmann and H.-E. Schaefer, 12 Sept
1973, field no. 27 (A 7).

Paratypes. ZFMK 73.250 — 73.257, 4 males, 3 females,
skins and skulls, same loclaity, collector and date as holo-
type, field nos. 20-26, 28 (A 1-6, 8).

Current name. Apodemus sylvaticus (Linnaeus, 1758).
Comments. This name is missing in Wilson & Reeder
(2005).

Canariomys tamarani Lopez-Martinez & Lopez-Jurado,
1987

Donana, Publicacion ocasional 2:10

Material. ZFMK 2009.634, cast of holotype I-11, skull
without mandibles, rostrum and braincase damaged;
ZFMK 2009.635-638, casts of paratypes I-267, I-47, I-
516, 1-592, skull, upper toothrows, and 2 left mandibles;
all from Holocene dune sediments at La Aldea de San
Nicolas de Tolentino, Gran Canaria, Canary Islands, Spain,
collected by L.F. Lopez-Jurado and colleagues.
Current name. Canariomys tamarani Lopez-Martinez &
Lopez-Jurado, 1987.

Comments. The original type series is in the Museo Ca-
nario, Las Palmas de Gran Canaria, Spain. The species is
extinct.

Colomys goslingi eisentrauti Dieterlen, 1983

Bonn. zool. Beitr. 34: 89 (12 Sept)

Holotype. ZFMK 74.366, adult male, skin and skull; Ba-
fut-Ngemba Reserve, Mount Lefo, 1800-1900 m, SE Ba-
menda Highlands, W Cameroon, collected by W. Bohme
and W. Hartwig, 6 Mar 1974, field no. 88.

Paratype. ZFMK 74.365, female, skin and skull, same lo-
cality, collectors and date as holotype. Another paratype
was transferred to SMNS (now SMNS 32300).

Current name. Colomys goslingi eisentrauti Dieterlen,
1983.

Comments. The taxon deserves species status (Hutterer
& Dieterlen, unpubl. data).

Colomys goslingi ruandensis Dieterlen, 1983

Bonn. zool. Beitr. 34: 89 (12 Sept)

Holotype. ZFMK 68.548, adult male, skin and skull;
Rugege Forest, ca. 2300 m, Uinka, W Ruanda, collected
by F. Dieterlen, 21 Aug 1964, field no. D 3450.
Paratypes. ZFMK 68.549 — 68.551, skins and skulls,
same locality and collector as holotype.

Current name. Colomys goslingi ruandensis Dieterlen,
1983.

©ZFMK

Types in ZFMK: Section Theriology 17

Dasymys incomptus longipilosus Eisentraut, 1963

Die Wirbeltiere des Kamerungebirges, Parey, Hamburg
und Berlin, p. 132, fig. 30

Paratype. ZFMK 63.670, male, skin and skull; grasslands
above Musake Hiitte, Mount Cameroon, W Cameroon,
collected by M. Eisentraut, 15 Feb 1954, field no. 169;
received in exchange from SMNS (Mar 1979).

Current name. Dasymys incomptus (Sundevall, 1847), as
in Musser & Carleton (2005), but may be a different form
(Hutterer, unpubl.).

Comments. Holotype in SMNS.

Deomys ferrugineus poensis Eisentraut, 1965

Bonn. zool. Beitr. 16: 8 (1 Nov)

Holotype. ZFMK 64.1091, adult male, skin (prepared
from specimen in spirit) and skull; Refugio, Bioko, Equa-
torial Guinea, collected by Leon Martin Zambo, July 1964,
field no. 691.

Paratypes. ZFMK 66.796, in spirit, field no. 695; ZFMK
64.1092, male, skin and skull, field no. 692; ZFMK
64.1090, 64.1093, 64.1094, females?, all same locality,
collector and date as holotype.

Current name. Deomys ferrugineus Thomas, 1888; tax-
onomic status not evaluated.

Comments. One skin out of this series is figured in colour
in Eisentraut (1973: plate 1).

Desmodillus auricularis hoeschi Lehmann, 1955

Bonn. zool. Beitr. 6: 171 (31 Dec)

Holotype. ZFMK 55.76, male, skin and skull; Okatjongea-
ma, Namibia, collected by W. Hoesch, 22 Apr 1955, field
no. 1279.

Paratypes. ZFMK 55.80, 55.466, males, skins and
skulls, same locality and collector.

Current name. Desmodillus auricularis (Smith, 1834);
see Petter (1971).

Desmodillus auricularis wolfi Lehmann, 1955

Bonn. zool. Beitr. 6: 172 (31 Dec)

Holotype. ZFMK 55.77, male, skin and skull; Vogelwei-
de, Namibia, collected by W. Hoesch, 7 July 1954, field
no. 1020.

Paratypes. ZFMK 55.81, 55.472, females, skins and
skulls, same locality, collector and date.

Current name. Desmodillus auricularis (Smith, 1834);
see Lehmann (1960) and Petter (1971).

Grammomys caniceps Hutterer & Dieterlen, 1984
Stuttg. Beitr. Naturk. A, 374: 12, figs. Lh, 2h, 3, 4, Sa, 6a,
7 (31 Dec)

Holotype. ZFMK 84.76, male, skin, skull, baculum and
viscera in alcohol; Malindi (03°14’S, 40°05’E), Kenia,
collected by F. Frohlich, Sept 1982, in captivity until 24
May 1984.

Bonn zoological Bulletin 59: 3-27

Paratypes. 3 males (ZFMK 84.32, 84.34, 84.74) and 5
females (ZFMK 84.31, 84.33, 84.72, 84.75), all skins and
skulls, offpring of imported animals.

Current name. Grammomys caniceps Hutterer & Di-
eterlen, 1984.

Comments. A further paratype in Stuttgart (SMNS
38200). ZFMK 84.34 is voucher for the karyotype of 2n
= 56, NF = 78.

Gerbillus (Gerbillurus) vallinus tytonis Bauer & J. Ni-
ethammer, 1960

Bonn. zool. Beitr. 10: 255, figs. 4-6 (10 Jan)

Holotype. ZFMK 59.515, complete skull, from a mum-
my; Sossus Vley, Namib, Namibia, collected by G. Ni-
ethammer, 18 Feb 1959, field no. 1152.

Paratypes. ZFMK 59.516, 6 skulls out of barn owl pel-
lets, same locality, collector and date.

Current name. Gerbillurus tytonis (Bauer & J. Nietham-
mer, 1960)

Comments. The year of publication is often given as
1959, but the relevant journal issue was published 10 Jan
1960.

Gerbillus (Dipodillus) dasyurus leosolliticus Lehmann,
1966

Zool. Beitr. (N.F.) 12: 288 (25 Oct)

Holotype. ZFMK 65.174, female, skin and skull; Deit-el-
Hajar, 25 km SE Damascus, Syria, collected by H. Mit-
tendorf, 22 Jan 1965, Coll. Kumerloeve no. 185.
Paratypes. ZFMK 65.175 — 65.177, 2 males, 1 female,
skins and skulls, same locality and collector, Coll.
Kumerloeve nos. 191—193.

Current name. Dipodillus dasyurus (Wagner, 1842); see
Musser & Carleton (2005).

Gerbillus (Dipodillus) dasyurus palmyrae Lehmann,
1966

Zool. Beitr. (N.F.) 12: 288 (25 Oct)

Holotype. ZFMK 65.183, female, skin and skull; Palmyra,
Syria, collected by H. Mittendorf, 2 Jan 1965, Coll.
Kumerloeve no. 174.

Paratypes. ZFMK 65.184 — 65.186, 1 male, 2 females,
skins and skulls, same locality and collector, Coll.
Kumerloeve nos. 172, 173, 175.

Current name. Dipodillus dasyurus (Wagner, 1842); see
Harrison (1972) and Musser & Carleton (2005).

Gerbillus foleyi Heim de Balsac, 1936

Bull. Biol. Fr. Belg., Suppl. 21: 317, 389, fig. 15
Paratype. ZFMK 60.85, skin and skull; Beni-abbés, Al-
geria, collected by H. Heim de Balsac, without date.
Current name. Gerbillus gerbillus (Olivier, 1801).

©ZFMK

18 Rainer Hutterer & Gustav Peters

Hybomys eisentrauti Van der Straeten & Hutterer, 1986
Mammalia 50: 36, fig. 1

Holotype. ZFMK 74.377, adult male, skin and skull; Lake
Bambului, Mount Lefo, 1800 m, Cameroon, collected by
W. Bohme, 10 Mar 1974, field no. 115.

Paratypes. ZFMK 74.374, young adult male, 6 Mar 1974;
ZEMK 74.375, adult male, 6 Mar 1974; ZFMK 74.376,
young adult male, 9 Mar 1974; same collector and local-
ity as holotype; ZFMK 69.175, adult male; Lake Oku,
2100 m, Mount Oku, Cameroon, collected by M. Eisen-
traut, 19 Jan 1967; all specimens skin and skull.
Current name. Not resolved; Musser & Carleton (1995)
included it into Hybomys badius Osgood, 1936, while Mis-
soup (2010) used the combination Hybomys rufocanus
eisentrauti Van der Straeten & Hutterer, 1986.

Hybomys univittatus basilii Eisentraut, 1965

Zool. Ab: Syst, 922 20sfiss, 152

Holotype. ZFMK 63.670, adult male, skin and skull; Mo-
ca Valley, ca. 1200 m, Bioko, Equatorial Guinea, collect-
ed by M. Eisentraut, 16 Nov 1962, field no. 258.
Paratypes. 33 skins with skulls, ZFMK 64.275 — 64.279,
64.281 — 64.287, 64.289 — 64.308, 66.658, Fernando Pdéo
(=Bioko), Equatorial Guinea, same collector, Oct-Dec
1962; Moca Valley, field nos. 153, 154, 159, 166, 171, 172,
184-186, 198, 199, 200, 209, 225, 233, 253, 259, 264, 265,
277-279, 291, 295, 299, 689, 2 specimens without cata-
logue no.; Refugium, field nos. 359, 376-378, 459.
Current name. Hybomys basilii Eisentraut, 1965 (Van der
Straeten 1985).

Comments. In the original description Eisentraut (1965:
20) mentioned 34 more specimens in addition to the holo-
type. In Apr 1979 33 of these could be located in the
ZFMK collection. The missing specimen was very like-
ly given to Pater Basilio after whom this subspecies was
named. There are 15 more specimens in spirit in the
ZFMK collection. These were collected during Eisen-
traut’s 1962/63 expedition but were not considered in the
context of the description. Further paratypes were deposit-
ed in SMNS and AMNH.

Hylomyscus aeta grandis Eisentraut, 1969

Z. Saugetierkd. 34: 300

Holotype. ZFMK 69.731, adult female, skin and skull;
Lager IV, 2100 m, Lake Oku, Banso Highlands,
Cameroon, collected by M. Eisentraut, 22 Jan 1967, field
no. 391.

Paratypes. ZFMK 69.732 — 69.734, 1 male, 2 females,
skins and skulls, same locality and collector, 20-23 Jan
1967, field nos. 378, 394, 401.

Current name. Hylomyscus grandis Eisentraut, 1969 (see
Hutterer et al. 1992).

Bonn zoological Bulletin 59: 3-27

Hylomyscus alleni montis Eisentraut, 1969

Z. Saugetierkd. 34: 302

Holotype. ZFMK 66.372, adult male, skin and skull;
Refugium, 2000 m, Pico de Santa Isabel, Bioko, Equato-
rial Guinea, collected by M. Eisentraut, 22 Dec 1962, field
no. 406.

Paratypes. ZFMK 66.730, 66.731, 66.733 — 66.761,
66.691 — 66.693, 66.762 a, 35 skins and skulls, same lo-
cality, collector and date; ZFMK 69.664 — 69.669, 6 skins
and skulls, same locality and collector, 7 Oct 1966.
Current name. Uncertain; possibly Hylomyscus montis
Eisentraut, 1969 (RH, unpubl.).

Lemniscomys striatus mittendorfi Eisentraut, 1968
Bonn. zool. Beitr. 19: 7 (10 Apr)

Holotype. ZFMK 68.8, adult male, skin and skull; Lake
Oku, ca. 2100 m, Banso Highlands, W Cameroon, collect-
ed by M. Eisentraut, 30 Jan 1967, field no. 452.
Paratypes. ZFMK 69.258 — 69.262, 2 males, 3 females,
skins and skulls, same locality and collector, 29 Jan — 11
Feb 1967, field nos. 463, 453, 455, 443, 464.

Current name. Lemniscomys mittendorfi Eisentraut,
1968; see Van der Straeten & Verheyen (1980).
Comments. In the text of his description Eisentraut men-
tions the elevation of the type locality as ca. 2300 m, in
the locality register and on the specimen label it is listed
as 2100 m. The holotype is figured in colour in Eisentraut
(19735p1N3):

Lophuromys dieterleni Verheyen, Hulselmans, Colyn &
Hutterer, 1997

Bull. Inst. r. Sci. nat. Belg. Biol. 67: 173, figs. 2, 5
Holotype. ZFMK 68.289, adult male, skin and skull; Lake
Oku, Lager IV (06°12’N, 10°32’E), 2100m, Bamenda-
Banso Highlands, W Cameroon, collected by M. Eisen-
traut, 21 Jan 1967, field no. 384.

Paratypes. ZFMK 69.286, 69.287 (adult females),
69.288, 69.290 (adult males), skins and skulls, same lo-
cality and collector as holotype, 17 — 31 Jan 1967, field
nos. 459, 347, 369, 389.

Current name. Lophuromys dieterleni Verheyen, Hulsel-
mans, Colyn & Hutterer, 1997; see Musser & Carleton
(2005).

Comments. The status of this taxon was discussed by Mis-
soup (2010).

Lophuromys medicaudatus Dieterlen, 1975

Bonn. zool. Beitr. 26: 295, figs. 1-4 (Dec)

Paratypes. ZFMK 68.982 (male), 68.984 (female), skins
and skulls; Kahuzi, Democratic Republic of the Congo,
collected by F. Dieterlen, 11-13 Nov 1966, field nos. D
9069, D 9036; ZFMK 68.983, 68.985, 1 male, 1 female,
skins and skulls, Tshibati, Democratic Republic of the
Congo, same collector, 30 Mar 1966 and 7 Apr 1964, field
nos. D 7254, D 1884.

©ZFMK

Types in ZFMK: Section Theriology 19

Current name. Lophuromys medicaudatus Dieterlen,
1975
Comments. Holotype in SMNS.

Lophuromys roseveari Verheyen, Hulselmans, Colyn &
Hutterer, 1997

Bull. Inst. r. Sci. nat. Belg. Biol. 67: 167, figs. 2, 3
Holotype. ZFMK 69.297, female, skin and skull; Musake
Hiitte (04°08’N, 09°12’°E), Mount Cameroon, 1850-2200
m, Cameroon, collected by M. Eisentraut, 18 Mar 1967,
field no 656.

Paratypes. ZFMK 69.292 (adult male), 69.302 (adult fe-
male), skins and skulls, Musake Htitte, Mount Cameroon,
1850-2200 m, field nos. 655, 658; ZFMK 69.293, 69.296,
adult females, skins and skulls, Htitte II (04°11°N,
09°12’E), Mount Cameroon, 3000 m, Cameroon, field
nos. 716, 712; ZFMK 61.790, adult male, skin and skull,
Buea (04°09’N, 09°13’E), 1800 m, Cameroon, field no.
4; all collected by M. Eisentraut.

Current name. Lophuryomys eisentrauti roseveari Ver-
heyen, Hulselmans, Colyn & Hutterer, 1997; see Missoup
(2010).

Lophuromys sikapusi eisentrauti Dieterlen, 1979
Bonn. zool. Beitr. 29: 296, figs. 1, 4 (30 Jan)

Holotype. ZFMK 74.436, adult male, skin and skull; slope
of Mount Lefo, near Bamenda, Cameroon, collected by
W. Bohme, 6 Mar 1974, field no. 81.

Paratype. ZFMK 74.437, juvenile to subadult female,
skin and skull, same collector, locality and date as holo-
type, field no. 92.

Current name. Lophuromys eisentrauti Dieterlen, 1979.

Lophuromys sikapusi parvulus Eisentraut, 1965

Zool. Jb. Syst. 92: 24, figs. 3, 4

Holotype. ZFMK 63.669, adult female, skin and skull;
Moca Valley, 1200 m, Bioko, Equatorial Guinea, collect-
ed by M. Eisentraut, 28 Oct 1962, field no. 165.
Paratypes. ZFMK 64.481 — 64.483, 1 male, 2 females,
skins and skulls, same locality and collector as holotype,
30 Oct-16 Nov 1962, field nos. 173, 262, 178.

Current name. Lophuromys nudicaudus Heller, 1911.

Malpaisomys insularis Hutterer, Lopez-Martinez &
Michaux, 1988

Palaeovertebrata 18: 246, figs. 3—5, 7, plates 1, 2
Paratypes. ZFMK, yet uncounted and uncatalogued
mandibles, maxillaries, and skeletal elements; all from
type level CV-A1 at locality Cueva Villaverde near La Oli-
va (22°38’N, 10°12’30”W), Fuerteventura, Canary Is-
lands, Spain; age 1070 + 50 BP.

Current name. Malpaisomys insularis Hutterer, Lopez-
Martinez & Michaux, 1988.

Bonn zoological Bulletin 59: 3-27

Comments. Holotype in the Department of Palaeontol-
ogy, Universidad Complutense, Madrid (CV-188.1).
Paratypes were distributed to AMNH, BMNH, CM,
FMNH, SMF, and many other collections. Extinct species.

Mastomys verheyeni Robbins & Van der Straeten, 1989
Senckenberg. biol. 69: 4, figs. 1, 2

Paratypes. ZFMK 92.468 (male), 92.469 (female), both
preserved in spirit with skulls extracted; Guemé, N
Cameroon, both collected by members of RUCA, 23 Nov
1970, field nos. RUCA 162, 163.

Current name. Mastomys kollmanspergeri (Petter, 1957)
(Musser & Carleton 2005).

Comments. Specimens were donated by E. Van der
Straeten in 1992. Holotype and paratypes in SMF, further
paratypes in RMCA.

Mus sylvaticus maximus Burg, 1925

Pallasia 3: 70

Lectotype. ZFMK 83.169, adult female, skin and skull;
Bergell, Switzerland, collected by G. von Burg, Feb 1921.
Paralectotype. ZFMK 83.170, skin; Bergell, Switzerland,
collected by G. von Burg, Mar 1923.

Current name. Apodemus flavicollis (Melchior, 1834);
see Hutterer (1984).

Comments. The lectotype was designated by Hutterer
(1984). Musser & Carleton (2005) listed this name under
the synonymy of A. sy/vaticus (Linnaeus, 1758). Apode-
mus sylvaticus maximus (Burg, 1925) is a senior homo-
nym of the extinct taxon Apodemus maximus Thaler, 1972,
described from the Pliocene of Sicily (Thaler 1972). A sub-
stitute name is therefore proposed for the fossil species:
Apodemus pellegrinoides Hutterer nom. nov. (pro Apode-
mus maximus Thaler, 1972, not Apodemus sylvaticus max-
imus (Burg, 1925)).

Ocenomys hypoxanthus albiventris Eisentraut, 1968
Bonn. zool. Beitr. 19: 8 (10 Apr)

Holotype. ZFMK 68.9, adult male, skin and skull; Lager
IV, ca. 2100 m, Lake Oku, Banso Highlands, W
Cameroon, collected by M. Eisentraut, 23 Jan 1967, field
nowso7:

Paratypes. ZFMK 69.196 — 69.198, 1 male, 2 females,
skins and skulls, same locality and collector, 25 Nov
1966-2 Feb 1967, field nos. 171, 465, 395.

Current name. Included in Oenomys hypoxanthus
(Pucheran, 1855) by Musser & Carleton (2005), but this
form requires a re-consideration.

Comments. Holotype figured in colour in Eisentraut
(Bonn. zool. Monogr. 3: plate 3, 1973).

©Z7FMK

20 Rainer Hutterer & Gustav Peters

Otomys occidentalis Dieterlen & Van der Straeten, 1992
Bonn. zool. Beitr. 43: 386, figs. 2-4

Paratypes. ZFMK 69.218, female, skin and skull; Mount
Oku, 2100 m, Cameroon, collected by M. Eisentraut, 28
Jan 1967; ZFMK 88.139, ZFMK 88.140, females, skins
and skulls, Gangirwal, Gotel Mountains, Nigeria, collect-
ed by G. Nikolaus, 18 and 20 Mar 1988.

Current name. Ofomys occidentalis Dieterlen & Van der
Straeten, 1992

Comments. The holotype SMNS 41336 consists of a
skull, the skin is lost.

Praomys hartwigi Eisentraut, 1968

Bonn. zool. Beitr. 19: 8 (10 Apr)

Holotype. ZFMK 68.7, adult female, skin and skull; Lager
IV, Lake Oku, Banso Highlands, W Cameroon, collected
by M. Eisentraut, 24 Jan 1967, field no. 402.
Paratypes. ZFMK 69.1073, 69.1075, 69.1077 — 69.1079,
69.1081, 2 males, 4 females, skins and skulls, same lo-
cality and collector, 20 — 26 Jan 1967, field nos. 399, 396,
409, 377 a, 385, 386; ZFMK 69.1074, 69.1076, 1 male,
1 female, skins and skulls, Lager III, ca. 1800 m, Lake Ma-
nenguba, same collector, 2—4 Jan 1967, field nos. 316, 323.
Current name. Praomys hartwigi Eisentraut, 1968.
Comments. One paratype (ZFMK 69.1072) was trans-
ferred in exchange to SMNS.

Praomys hartwigi obscurus Hutterer & Dieterlen, 1992
Bonn. zool. Beitr. 43: 402, figs. 1, 2 (21 Oct)

Holotype. ZFMK 88.115, adult female, skin and skull;
Gangirwal, 2300 m, Gotel Mountains, SE Nigeria, collect-
ed by G. Nikolaus, 16 Mar 1988, field no. (GN5).
Paratypes.10 specimens, ZFMK 88.114, 88.116 — 88.124,
same locality and collector as holotype, 15—20 Mar 1988;
6 specimens, ZFMK 88.125 — 88.130, Chappal Waddi,
1900 m, same collector, 21—23 Mar 1988.

Current name. Praomys obscurus Hutterer & Dieterlen,
1992 (Musser & Carleton 2005).

Comments. Described in Hutterer, Dieterlen & Nikolaus
(1992). Additonal paratypes in SMNS.

Praomys petteri Van der Straeten, Lecompte & Denys,
2003

Bonn. zool. Beitr. 50: 333, figs. 2, 3 (24 Feb)
Paratypes. ZFMK 70.146 (male), 70.147 (female),
92.318 - 92.322 (2 males, 3 females), all preserved in spir-
it with skulls extracted; all from breeding colony of F. Pet-
ter (Paris), from animals collected 1968 at Boukoko, Cen-
tral African Republic.

Current name. Praomys petteri Van der Straeten,
Lecompte & Denys, 2003.

Comments. The specimens are from the breeding colony
of “ P. morio” which yielded the karyoptype of 2n = 42
(Petter 1965).

Bonn zoological Bulletin 59: 3-27

Rattus rattus povolnyi J. Niethammer & Martens, 1975
Z. Saugetierkd. 40: 344

Holotype. ZFMK 78.001, male, skin and skull; Dar-1-Nur,
near Dyalalabad, 1200 m, E Afghanistan, collected by J.
Niethammer, 18 Apr 1966, field no. A 1230.

Paratypes. ZFMK (not catalogued), 9 specimens, same
data as holotype, field nos. A 1223, A 1225, A 1229, A
1531, A 1534, A 1584 — 1587.

Current name. Rattus tanezumi Temminck, 1844; see
Musser & Carleton (2005).

Comments. All specimens were collected in houses (field
notes of J. Niethammer).

Thamnomys rutilans poensis Eisentraut, 1965

Zool. Jb. Syst. 92: 26, fig. 5

Holotype. ZFMK 63.668, adult male, skin and skull; Mo-
ca Valley, 1200 m, Bioko, Equatorial Guinea, collected by
M. Eisentraut, 7 Nov 1962, field no. 209 a.

Paratypes. ZFMK 64.484 — 64.487, females, skins and
skulls, field nos. 292 (29 Nov), 226 (9 Nov), 683 (3 Nov
1962), all same locality and collector as holotype; ZFMK
66.795, specimen in spirit, same locality and collector, 11
Nov 1962, field no. 229; ZFMK 64.486, skin and skull,
shrub willow area above Moca, ca. 1500 m, same collec-
tor, 11 Mar 1963, field no. 683.

Current name. Grammomys poensis (Eisentraut, 1965).
Comments. Grammomys poensis replaces the better
known G. rutilans which is not available (Hutterer, in
Musser & Carleton 2005).

Octodontidae

Octodon pacificus Hutterer, 1996

Z. Saugetierkd. 59: 28, figs. 1-6

Holotype. ZFMK 92.384, adult female, skin and skull; Is-
la Mocha, Arauco Province, Chile, collected by Francis-
co Behn, 16 Jan 1959, field no. L6. Skull lacks occipital
and bullae.

Paratypes. 3 skins and skulls, ZFMK 92.383, adult fe-
male, and ZFMK 92.385-6, two juveniles, collected by F.
Behn at the type locality beween 11 and 24 Jan 1959.
Current name. Octodon pacificus Hutterer, 1996.
Comments. The species is most probably extinct, the type
specimens obtained in 1959 being the first and last record
of a living animal. A survey of mammals on Isla Mocha
yielded no trace of this species, however, remains were
found in an archaeological excavation (Saavedra et al.
2003).

©ZFMK

Types in ZFMK: Section Theriology 2]

CONCLUSIONS

This catalogue presents the status of the ZFMK mammal
collections in 2010. 77 holotypes, 5 syntypes, 2 lectotypes,
and | neotype are listed. In addition, paratypes or para-
lectotypes for 105 taxa are included. The number of type
specimens will increase in the future, as more species of
mammals will be described, based on ZFMK material.
This is particularly true for Africa, where new species of
mammals are frequently being discovered (Hoffmann et
al. 2009).

Some issues concerning type material could not yet be re-
solved, but may be so in the future. For example, Groves
(2003) mentioned unspecified material in “Museum
Alexander Koenig (Bonn)” that served for his description
of Gazella benettii salinarum Groves, 2003. We were not
able to trace such material so far. The description of Sun-
cus hututsi Kerbis Peterhans & Hutterer, 2009 includes a
statement that one paratype will be deposited in the ZFMK
(Kerbis Peterhans & Hutterer 2009) but this has not been
done yet.

Numerous vouchers for karyotypes still have to be iden-
tified. For example, specimens of European and Asian Mus
and Vandeleuria were used by Gropp et al. (1969, 1970,
1973) to analyze their karyotypes, as were specimens of
Mus from Sicily by Lehmann & Radbruch (1977). Also
GenBank voucher specimens will have to be identified in
order to link published sequences with vouchers in the
Bonn collection. This is one of the next goals to be
achieved.

GEOGRAPHIC ORIGIN OF TYPE SPECIMENS
(EXCLUDING CASTS)

Afghanistan
Rattus rattus povolnyi J. Niethammer & Martens
Algeria
Gerbillus foleyi Heim de Balsac
Australia
Megaderma gigas Dobson
Austria
Sorex araneus wettsteini Bauer
Bosnia and Herzegovina (former Yugoslavia)
Crocidura suaveolens balcanica Ondrias
Brazil
Sciurus ingrami sebastiani Miller & Vesmanis
Bulgaria
Clethrionomys glareolus pirinus Wolf
Muscardinus avellanarius kroecki G. Niethammer &
Bohmann

Bonn zoological Bulletin 59: 3-27

Cameroon
Colomys goslingi eisentrauti Dieterlen
Crocidura flavescens bueae Heim de Balsac & Barloy
Crocidura manengubae Hutterer
Crossarchus platycephalus Goldman
Dasymys incomptus longipilosus Eisentraut
Hipposideros camerunensis Eisentraut
Hybomys eisentrauti Van der Straeten & Hutterer
Hylomyscus aeta grandis Eisentraut
Lemniscomys striatus mittendorfi Eisentraut
Lophuromys dieterleni Verheyen, Hulselmans, Colyn &
Hutterer
Lophuromys roseveari Verheyen, Hulselmans, Colyn &
Hutterer
Lophuromys sikapusi eisentrauti Dieterlen
Mastomys verheyeni Robbins & Van der Straeten
Myosorex eisentrauti okuensis Heim de Balsac
Myosorex eisentrauti rumpii Heim de Balsac
Oenomys hypoxanthus albiventris Eisentraut
Otomys occidentalis Dieterlen & Van der Straeten
Paracrocidura schoutedeni camerunensis Heim de
Balsac
Pipistrellus eisentrauti Hill
Praomys hartwigi Eisentraut
Rousettus aegyptiacus occidentalis Eisentraut
Sylvisorex granti camerunensis Heim de Balsac
Sylvisorex silvanorum Hutterer, Riegert & Sedlaéek
Central African Republic
Praomys petteri Van der Straeten, Lecompte & Denys
Chile
Octodon pacificus Hutterer
Colombia
Platyrrhinus chocoensis Alberico & Velasco
Czech Republic
Amblyotus atratus Kolenati
Democratic Republic of the Congo
Crocidura kivuana Heim de Balsac
Crocidura lanosa Heim de Balsac
Crocidura littoralis stenocephala Heim de Balsac
Lophuromys medicaudatus Dieterlen
Sylvisorex akaibei Mukinzi, Hutterer & Barriere
Ecuador
Nesoryzomys fernandinae Hutterer & Hirsch
Equatorial Guinea
Deomys ferrugineus poensis Eisentraut
Hybomys univittatus basilii Eisentraut
Hylomyscus alleni montis Eisentraut
Lophuromys sikapusi parvulus Eisentraut
Myosorex eisentrauti Heim de Balsac
Sylvisorex morio isabellae Heim de Balsac
Thamnomys rutilans poensis Eisentraut
Ethiopia
Crocidura flavescens hansruppi Hutterer

©ZFMK

i.)
NO

France
Plecotus alpinus Kiefer & Veith
Germany
Apodemus agrarius henrici Lehmann
Apodemus flavicollis alpinus Heinrich = Apodemus
flavicollis alpicola Heinrich
Neomys anomalus rhenanus Lehmann
Pitymys bavaricus Konig
Sorex araneus huelleri Lehmann
Sorex minutus becki Lehmann
Greece
Crocidura ariadne Pieper
Guinea
Crocidura (bottegi) obscurior Heim de Balsac
Genetta bourloni Gaubert
Rhinolophus denti knorri Eisentraut
Rhinolophus landeri guineensis Eisentraut
Rhinolophus ziama Fahr, Vierhaus, Hutterer & Kock
India
Gazella benettii salinarum Groves
Indonesia
Rhinolophus sumatranus K. Andersen
Italy
Apodemus flavicollis sicilianus Lehmann & Schaefer
Apodemus tauricus geminae Lehmann
Clethrionomys glareolus curcio Lehmann
Clethrionomys glareolus garganicus Hagen
Crocidura sicula aegatensis Hutterer
Crocidura suaveolens bruecheri Lehmann
Dinops cestoni Savi
Dryomys nitedula aspromontis Lehmann
Microtus savii niethammericus Contoli
Pitymys savii brachycercus Lehmann
Sorex araneus silanus Lehmann
Sorex etruscus Savi
Talpa romana wittei Capolongo
Vespertilio bonapartii Savi
Vulpes vulpes toschii Lehmann
Kenya
Grammomys caniceps Hutterer & Dieterlen
Liberia
Genetta (Paragenetta) lehmanni Kuhn
Liberiictis kuhni Hayman
Liechtenstein
Sorex minutus becki Lehmann
Malta
Crocidura sicula calypso Hutterer
Montenegro (former Yugoslavia)
Dryomys nitedula diamesus Lehmann
Talpa caeca steini Grulich
Talpa stankovici montenegrina KryS8tufek
Namibia
Desmodillus auricularis hoeschi Lehmann
Desmodillus auricularis wolfi Lehmann
Elephantulus intufi omahekensis Lehmann

Bonn zoological Bulletin 59: 3-27

Rainer Hutterer & Gustav Peters

Eremitalpa granti namibensis Bauer & J. Niethammer
Gerbillus (Gerbillurus) vallinus tytonis Bauer &
J. Niethammer
Petromys collinus kurzi Lehmann
Pronolagus randensis waterbergensis Hoesch &
Lehmann
Nigeria
Crocidura yankariensis Hutterer & Jenkins
Praomys hartwigi obscurus Hutterer & Dieterlen
Sylvisorex corbeti Hutterer & Montermann
Oman
Crocidura arabica Hutterer & Harrison
Paraguay
Cabassous chacoensis Wetzel
Republic of the Congo
Congosorex verheyeni Hutterer, Barriere & Colyn
Republic of the Philippines
Crocidura batakorum Hutterer
Crocidura panayensis Hutterer
Russia
Alces alces pfizenmayeri Zukowsky
Lagomys ater Eversmann
Rwanda
Colomys goslingi ruandensis Dieterlen
Sylvisorex vulcanorum Hutterer & Verheyen
Senegal
Crocidura lamottei elegans Hutterer
Serbia (former Yugoslavia)
Rhinolophus ferrumequinum martinoi Petrov
Spain
Crocidura canariensis Hutterer, Lopez-Jurado & Vogel
Crocidura osorio Molina & Hutterer
Felis catus jordansi Schwarz
Malpaisomys insularis Hutterer, Lopez-Martinez &
Michaux
Neomys fodiens niethammeri Buhler
Plecotus auritus hispanicus Bauer
Sciurus vulgaris hoffmanni Valverde (= Sciurus vul-
garis valverdei nom. nov.)
Switzerland
Mus sylvaticus maximus Burg
Syria
Gerbillus (Dipodillus) dasyurus leosolliticus Lehmann
Gerbillus (Dipodillus) dasyurus palmyrae Lehmann
Tanzania
Congosorex phillipsorum Stanley, Rogers & Hutterer
Crocidura tansaniana Hutterer
Crocidura telfordi Hutterer
Myosorex kihaulei Stanley & Hutterer
Tunisia
Erinaceus algirus girbaensis Vesmanis
Turkey
Acomys cilicius Spitzenberger
Microtus anatolicus Krystufek & Kefelioglu
Muscardinus avellanarius abanticus Kivan¢

©OZFMK

Types in ZFMK: Section Theriology 23

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©ZFMK

26 Rainer Hutterer & Gustav Peters

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©ZFMK

Bonn
zoological

Bulletin

formerly: Bonner zoologische Beitrage

ici
| =
| Bonn Volume 57
| Zoological oo10
| Bulletin
| Sst
Se Wisin
|
| Bonn AEST
Mies | zoological 2010

| SSSR Bulletin

Bonn
zoological
Bulletin

Bonn
zoological

An open access journal of organismal zoology, published by
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Bulletin

Index 2010
Volumes
57-59

Editor-in-Chief

Fabian Herder

Editorial Board

Dirk Ahrens
Wolfgang BoOhme
Netta Dorchin
Renate van den Elzen
Bernhard A. Huber
Rainer Hutterer

| Gustav Peters
. Bradley Sinclair

— Dieter StUning

Contents Volume 57

Arribas, Oscar J.:

Intraspecific variability of the Carpetane Lizard (/berolacerta cyreni [Muller & Hellmich, 1937])
(Squamata: Lacertidae), with special reference to the unstudied peripheral populations

from the Sierras de Avila (Paramera, Serrota and Villafranca)

LOZ —2i0

Barej, Michael F., lvan Ineich, Vaclav Gvozdik, Nathaly Lhermitte-Vallarino,

Nono Legrand Gonwouo, Matthew LeBreton, Ursula Bott & Andreas Schmitz:

Insights into chameleons of the genus Trioceros (Squamata: Chamaeleonidae) in Cameroon,
with the resurrection of Chamaeleon serratus Mertens, 1922

211-229

Bauer, Aaron M.:

A new species of Pachydactylus (squamata: Gekkonidae) from the Otavi Highlands of northern Namibia

257-266

Capula, Massimo & Claudia Corti:
Genetic variability in mainland and insular populations of Podarcis muralis (Reptilia: Lacertidae)

189-196

Carretero, Miguel A., Pietro Lo Casci, Claudia Corti, & Salvatore Pasta:
Sharing recources in a tiny Mediterranean Island? Comparative diets of
Chalcides ocellatus and Podarcis filfolensis in Lampione

111-118

Dubois, Alain & Roger Bour:
The distinction between family-series and class-series nomina in zoological nomenclature,

with emphasis on the nomina created by Batsch (1788, 1789) and on the higher nomencature of turtles

149-171

Gunther, Rainer:
Another new Cophixalus species (Amphibia: Anura: Microhylidae) from western New Guinea

231-240

Hita Garcia, Francisco, Georg Fischer, Patrick KUck, Birthe Thormann & Marcell K. Peters:
Tetramorium boehmei sp. n. — a new ant (Hymenoptera: Formicidae) species
from the Kakamega Forest, Western Kenya

359-366

Jirk, Miloslav, Andrei Daniel Mihalca, Petr Necas & David Modry:
An addition to the East African herpetofauna:
the first record of Tarentola annularis relicta (sSquamata: Gekkonidae) in Uganda

173-176

Joger, Ulrich & Ismail Bshaenia:
A new Tarentola subspecies (Reptilia: Gekkonidae) endemic to Tunisia

267-274

Koch, André, Mark Auliya & Thomas Ziegler:
Updated checklist of the monitor lizards of the world (Squamata: Varanidae)

127-136

Krause, Patrick & Wolfgang Bohme:
A new chameleon of the Trioceros bitaeniatus complex from Mt. Hanang, Tanzania, East Africa
(Squamata: Chamaeleonidae)

19-29

Kuhnel, Susanne, Sandy Reinhard & Andreas Kupfer:
Evolutionary reproductive morphology of amphibians: an overview

119-126

Lutzmann, Nicola, Jan Stipala, Ralph Lademann, Patrick Krause, Thomas M. Wilms & Andreas Schmitz:

Description of a new subspecies of Kinyongia uthmoelleri (Muller, 1938)
(Squamata: Chamaeleonidae) with notes on its captive propagation

281-288

Masseti, Marco:
Homeless mammals from the lonian and Aegean islands

367-373

Contents Volume 59

Bohme, Wolfgang:
A list of the herpetological type specimens in the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

79-108

Elzen, Renate van den:
Type specimens in the bird collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

29-77

Herder, Fabian, Jan Huylebrouck & Klaus Busse:
Catalogue of type specimens of fishes in the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

109-136

Hutterer, Rainer & Gustav Peters:
Type specimens of mammals (Mammalia) in the collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Publication dates:

Vol. 57 Issue 1: April 2010

Vol. 57 Issue 2: November 2010
Vol. 58: December 2010

Vol. 59: December 2010

Contents Volume 57

Arribas, Oscar J: 197-210
Intraspecific variability of the Carpetane Lizard (Iberolacerta cyreni [Miller & Hellmich, 1937})

(Squamata; Lacertidae), with special reference to the unstudied peripheral populations

from the Sierras de Avila (Paramera, Serrota and Villafranca) ra
Barej, Michael F., Ivan Ineich, Vaclay Gvozdik, Nathaly Lhermitte-Vallarino, 211-229
Nono Legrand Gonwouo, Matthew LeBreton, Ursula Bott & Andreas Schmitz:

Insights into chameleons of the genus Troceros (Squamata: Chamaeleonidae) in Cameroon,

with the resurrection of Chamaeleon serratus Mertens, 1922

Bauer, Aaron M.: 257-266
A new species of Pachydactylus (Squamata: Gekkonidae) from the Otavi Highlands of northern Namibia :
Capula, Massimo & Claudia Corti: 189-196
Genetic variability in mainland and insular populations of Podarcis muralis (Reptilia: Lacertidae)_

Carretero, Miguel A,, Pietro Lo Casci, Claudia Corti, & Salvatore Pasta: 111-118
Sharing recources in a tiny Mediterranean Island? Comparative diets of
Chalcides ocellatus and Podareis filfolensis in Lampione

Dubois, Alain & Roger Bour: 149-171
The distinction between family-series and class-series nomina in zoological nomenclature,
with emphasis on the nomina created by Batsch (1788, 1789) and on the higher nomencature of turtles

Giinther, Rainer: 231-240
Another new Cophixalus species (Amphibia: Anura: Microhylidae) from western New Guinea
Hita Garcia, Francisco, Georg Fischer, Patrick Kuck, Birthe Thormann & Marcell K. Peters:
Tetramorium boehmei sp. n. = a new ant (Hymenoptera: Formicidae) species

from the Kakamega Forest, Western Kenya

359-366

Jirkd, Miloslav, Andrei Daniel Mihalca, Petr Necas & David Modry
An addition to the East African herpetofauna:
the first record of Tarentola annularis relicta (Squamata; Gekkonidae) in Uganda

173-176

Joger, Ulrich & Ismail Bshaenia: 267-274
A new Tarentola subspecies (Reptilia: Gekkonidae) endemic to Tunisia

Koch, André, Mark Auliya & Thomas Ziegler: 127-136
Updated checklist of the monitor lizards of the world (Squamata: Varanidae)

Krause, Patrick & Wolfgang Bbhme:

A new chameleon of the Troceros bitaeniatus complex from Mt. Hanang, Tanzania, East Africa
(Squamata: Chamaeleonidae)

19-29

Kuhnel, Susanne, Sandy Reinhard & Andreas Kupfer: 119-126
Evolutionary reproductive morphology of amphibians: an overview

Lutzmann, Nicola, Jan Stipala, Ralph Lademann, Patrick Krause, Thomas M.
Description of a new subspecies of Kinyongia uthmoellen (Maller, 1938)

(Squamata: Chamaeleonidae) with notes on its captive propagation

Masseti, Marco:
Homeless mammals from the lonian and Aegean islands 367-373

Wilms & Andreas Schmitz: 281-288

Contents Volume 57

Nguyen, Quang Truong, Andreas Schmitz & Wolfgang Bohme: 15-17
Gekko ulikovskii Darevsky & Orlov, 1 1994: a junior synonym of Gekko badenii Szczerbak & Nekrasova, 1994 —

Niekisch, Manfred: 347-357
The history of reptiles and amphibians at Frankfurt Zoo

Pafilis, Panayiotes: 329-345
A brief history of Greek herpetology —

Rodel, Mark-Oliver, Laura Sandberger, Johannes Penner, Youssouph Mané & Annika Hillers: 177-188
The taxonomic status of Hyperolius spatzi Ahl, 1931 and Hyperolius nitidulus Peters, 1875
(Amphibia: Anura: Hyperoliidae)

Saikia, Munna Rani, Lakshmi Kanta Hazarika, Surajit Kalita & Ramen Lal Baishya: 91-98
Antennal sensilla of the rice hispa Dicladispa armigera (Olivier, 1808) (Coleoptera: Chrysomelidae)

Schmidtler, Josef Friedrich: 307-328
The taxonomic history of the Linnean genus Lacerta (Squamata: Sauria: Lacertidae)

in the mirror of book-illustration
Shoorcheh, Hossein Rajaei: 65-73
Life-history of Gnopharmia kasrunensis Wehrli, 1939 and G. colchidaria Lederer, 1870

(Geometridae, Ennominae) and their distribution in Iran, with first host-plant records for the genus

Sinclair, Bradley J.: 85-89
Proclinopyga ulrichi sp. nov.: the first fossil aquatic dance fly of the subfamily Clinocerinae
(Diptera: Empididae) —

Sonnenberg, Rainer & Eckhard Busch: 3-14
Description of Callopanchax sidibei (Nothobranchiidae: Epiplateinae), a new species of killifish from
southwestern Guinea, West Africa

StUning, Dieter 75-83
Two new species of the Plutodes costatus-group from the Philippines and Indonesia
(Lepidoptera, Geometridae, Ennominae)

Ullenbruch, Klaus, Olaf Grell & Wolfgang Bohme 31-54
Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae),
and a country-wide checklist

Vences, Miguel, Jorn K6hler, Angelica Crottini & Frank Glaw 241-255
High mitochondrial sequence divergence meets morphological bioacoustic conservatism
Boophis quasiboehmei sp. n., a new treefrog species from south-eastern Madagascar

Vogel, Gernot & Patrick David: 289-296
A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes Colubridae)

Wagner, Philipp & Thomas M. Wilms 297-306
A crowned devil: new species of Cerastes Laurenti, 1768 (Ophidia, Viperidae) from Tunesia,
with two nomenclatural comments _

Wilhelm, Gertha, Hans L. Nemeschkal, John Plant & Hannes F. Paulus 55-64
Fitness components in the relationship between Rhopalapion longirostre (Olivier, 1807)

(Insecta: Coleoptera: Apionidae) and A/cea rosea (Linnaeus, 1758) (Malvaceae)

Analysis of infestation balance of a herbivorous weevil and its host plant

Contents Volume 57

Wilms, Thomas M., Mohammed Shobrak & Philipp Wagner 275-280
A new species of the genus Tropiocolotes from Central Saudi Arabia (Reptilia; Sauria: Gekkonidae)

Ziegler, Thomas & Truong Quang Nguyen

137-147
New discoveries of amphibiens and reptiles from Vietnam

Book Reviews

FUnfstdck HJ, Ebert A & Weiss | (2010)
Taschenlexikon der Végel Deutschlands
(Kathrin Schidelko) 18

Moning C, Griesohn-Pfleger T & Horn M (2010)

Grundkurs Vogelbestimmung: Eine Einfahrung in die Beobachtung und Bestimmung unserer heimischen Vogel
(Darius Stiels) 99

Contents Volume 58

Blume, Christina, Claudia Etzbauer, Reinhard Gaedike, Uta Heidenreich, Ulrike Kleikamp, 169-215
Lena Schnug, Wolfgang Speidel & Dieter Stining

Type material of Lepidoptera and Trichoptera in the ZFMK collection, Bonn Se
Huber, Bernhard & Stefanie Lankhorst 217-226
Non-insect arthropod types in the ZFMK collection, Bonn
(Acari, Araneae, Scorpiones, Pantopoda, Amphipoda)

Lampe, Karl-Heinz, Dirk Rohwedder, Sigfrid Ingrisch & Birgit Rach: 89-167
Insect types in the ZFMK collection, Bonn: Blattodea, Dermaptera, Heteroptera, Homoptera”,

Hymenoptera, Mantodea, Orthoptera, Phasmatodea, Phthiraptera and Siphonaptera
Riede, Klaus & Mark Fichtner 3
Obituary to Karl-Heinz Lampe, Zoolagisches Forschungsmuseum Alexander Koenig:

Sinclair, Bradley J. & Netta Dorchin 49-88
Isoptera, Embioptera, Neuroptera, Mecoptera, Raphidioptera and Dipera types in 2FMK

Ulmen, Karin, Ruth Newzella, Lasse Hubweber, Michael Schmitt, Thorsten Klug & Dirk Ahrens: 5-48
Contribution to a catalogue of types preserved in the collection of
Zoologisches Forschungsmuseum Alexander Koenig (ZFMK): Coleoptera: 1. Checklist of taxa

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Types in ZFMK: Section Theriology D7

Turni H, Hutterer R , Asher R (2007) Type specimens of “in-
sectivoran” mammals at the Museum fiir Naturkunde, Berlin.
Zootaxa 1470: 1-33

Turni H, Kock D (2008) Type specimens of bats (Chiroptera,
Mammalia) in the collections of the Museum fiir Naturkunde,
Berlin. Zootaxa 1869: 1—82

Valverde JA (1967) Notas sobre vertebrados. III. Nueva ardilla
del S. E. espanol y consideraciones sobre las subespecies
peninsulares. Boletin de la Real Sociedad Espanola de His-
toria Natural, Seccion Biologia 65: 225-248

Van der Straeten E (1985) Note sur Hybomys basilii Eisentraut,
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33-45

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Volume 59

Bonn zoological Bulletin

pp. 29-77 Bonn, December 2010

Type specimens in the bird collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Renate van den Elzen

Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn, Germany;
E-mail: r.elzen.zfmk@uni-bonn.de.

INTRODUCTION

Primary types play an important role as name-bearing
specimens (ICZN, 1999) in the context of correctly ad-
dressing the whole spectrum of biodiversity. Names and
thus taxonomy are the first step to identify and arrange
the diversity of species in appropriate order. The impor-
tance of type specimens requires that the information about
types should be made globally available. This was the
main objective to present this checklist of syntypes, holo-
types, lectotypes and paratypes (including paralectotypes)
deposited in the skin collections of the ZFMK. A previ-
ous list of types in the ZFMK (Rheinwald & van den Elzen
1984) can be seen as a first step. The list is outdated, in-
complete and written in German only. Moreover, with the
progress of the digital catalogue of bird type specimens
in German research collections under GIBF-D-Vertebra-
ta (van den Elzen et al. 2005) and the increase of network-
ing between ornithological collections, several type series
were found to be spread over different collections and this
valuable information on whereabouts of type specimens
is added here. Also the new ICZN with its stricter rules
was applied to the former type list and helpful suggestions
from colleagues improved the revised version. The new
catalogue thus offers the state of knowledge of today. It
makes no claim to be complete or free of errors, but it is
thought to be a starting point for further discussions, ad-
ditions or amendments and other suggestions towards a
global register of scientific bird names and the collections
holding bird types.

Since the previous version of the ZFMK bird type cata-
logue 13 new taxa were added. The type checklist com-
prises today 322 primary names, 137 holotypes, 47 syn-
types and 95 lectotypes, as well as about 989 para- and
paralectotypes. A multitude of new names was proposed
by O. Kleinschmidt (and coauthors; 121), whose Collec-
tion (Coll. Kl.) was sold to the ZFMK in 1935. Several
other collections (of Harms, von Jordans, Dernedde,
Schiebel or Martens) were handed to the ZFMK between
1940 and 2006. These collections also included type ma-
terial that is considered here. And last not least the founder

Bonn zoological Bulletin 59: 29-77

of the ZFMK, Alexander Koenig, proposed eight new
names for birds, of which five are still recognized names
of bird species or subspecies today (Dickinson 2003).

The taxa presented in the checklist include all name-bear-
ing types, without evaluating their present acceptance. So
we list not only currently recognized taxa, but also syn-
onyms, regardless whether these are objective or subjec-
tive synonyms. The sequence of families follows the
Howard and Moore Checklist (Dickinson 2003). Also cur-
rent names were added as a service as in some cases the
original names cannot be assigned to current scientific bird
names without some difficulty. Current names follow
Dickinson (2003), but could not be applied with certain-
ty to all names proposed, as e.g. several of Kleinschmidt’s
taxon names are not listed in any forrunners of the Howard
and Moore Checklist (Dickinson 2003), e.g. Peters
Checklist (Mayr & Cottrell, 1979, 1986, Mayr & Green-
way 1960, 1962, Mayr & Paynter 1964, Paynter 1967,
1968, 1970, Peters 1931-1951, Traylor 1979), Hartert
(1910-1932) or other sources.

Abbreviations for Museum collections used:

AMNH = American Museum of Natural History, New
York, New York, U.S.A.

BMNH = Natural History Museum, Tring (= British Mu-
seum (Natural History), U.K.

CNHM = Chicago Natural History Museum, Chicago,
U.S.A.

DMD = Durban Museum, Durban, R.S.A.

MCZ = Harvard University, Massachusetts, U.S.A.
MfN = Museum ftir Naturkunde, Berlin, Germany (=
ZMB)

MHN = Museo de Historia Natural “Javier Prado”, Lima,
Peru

MTKD = see SNS

NHMW = Naturhistorisches Museum, Vienna, Austria

OZFMK

30 Renate van den Elzen

RSM = Royal Scottish Museum, Edinburgh, U.K.

SMF = Naturmuseum Senckenberg, Frankfurt/Main,
Germany

SMNS = Staatliches Museum fiir Naturkunde, Stuttgart
SNS = Senckenbergische Naturkundliche Sammlungen,
Dresden

UMB = Uberseemuseum, Bremen, Germany

ZMB = see MfN

ZMH = Zoologisches Museum Hamburg, Germany
ZFMK = Zoologisches Forschungsmuseum Alexander
Koenig, Bonn, Germany

ZSM = Zoologische Staatssammlung, Munich, Germany

CHECKLIST OF TYPE SPECIMENS

TINAMIDAE

Nothoprocta pentlandi niethammeri Koepcke, 1968
Bonn zool Beitr 19: 231.

Paratype: ZFMK 68.2060, male, Lomas of Lachay, ca. 90
km N of Lima, Peru. Captured as a chick, prepared as a
nine month old bird, 25.07.1960, leg. Montesa. Original
No Kp 1883 a, MHN Orn. 1117;

Other paratypes: Kp 1883 c, MHN Orn 2533, female, Lo-
mas of Lachay, 90 km north Lima, Peru; Kp 1883 d, male,
in 1968 in the zoological Garden Perque de las Leyndas,
Lima; Holotype: Kp. 1883 b, MHN Orn. 2534, male, Lo-
mas of Lachay, 90 km north Lima, Peru, in the MHN
Javier Prado, Lima. — Dickinson (2003): Nothoprocta
pentlandii niethammeri.

PHASIANIDAE

Perdix galliae Bacmeister & Kleinschmidt, 1918
J Ornithol 66: 255.

Lectotype: Coll. Kl. 8231, male, Aussonce, E France,
22.01.1917, leg. Monnard, ex Coll. Bacmeister;

Paralectotypes: Coll. KI. 8232 male, Thugny, Arrondisse-
ment de Rethel, E France, 02.12.1916, leg. Riist; Coll. K1.
8233 (male), Aussonce, 21.01.1917, leg. Monard; Coll. KI.
8234 Aussonce, 16.12.1916, found dead by Bacmeister,
all ex. Coll. Bacmeister.

Series of fourteen syntypes, whereabouts of other speci-

mens unknown; nom. nov. added as footnote. — Dickin-
son (2003): Perdix perdix perdix (Linnaeus, 1758).

Bonn zoological Bulletin 59: 29-77

Alectoris rufa laubmanni Jordans, 1928
Novit Zool 34: 306.

Holotype: ZFMK_ L.II.9.c?.a., male, Arta, Majorca,
03.04.1921, leg. A. von Jordans, No 2918.

Paratypes: ZFMK_ L.III.9.c?.B., male, Arta, Majorca,
31.03.1921; ZFMK L.III.9.c?-y., male, Salinas, Majorca,
03. 05. 1927; ZFMK L.III.9.c?.6, female, Salinas, Majorca,
03. 05. 1927; ZFMK L.HI.9.c?.¢, male, Salinas, Majorca,
03. 05. 1927; ZFMK L.HI.9. c?. ¢., male, Salinas, Majorca,
19. 05. 1927; ZFMK L.III.9.c?. female, 19.05.1927, Sali-
nas, Majorca, all leg. A. von Jordans. — Dickinson (2003):
Alectoris rufa intercedens (A. E. Brehm, 1857).

Francolinus coqui hoeschianus Stresemann, 1937
Orn Monatsber 45: 66.

Paratype: ZFMK 39.250, male, Waterberg-Plateau,
Namibia, 18.09.1936, leg. Hoesch No 628.

Holotype: MfN, Berlin (No 36.206) male, Otjosongombe,
Waterberg-Plateau, Namibia, 03.07.1936, leg Hoesch.

Three more paratypes probably also in MfN, Berlin. —
Dickinson (2003): Francolinus coqui coqui (A. Smith,
1836).

Francolinus hartlaubi crypticus Stresemann, 1939
Orn Monatsber 47: 62.

Paratype: ZFMK 39.967, female, Onguati, Kaokoveld,
Namibia, 03.09.1938, leg. Hoesch, No 1363.

Holotype at the MfN, Berlin (No 38.1343) — t. t. as
paratype; described from a series of six specimens. — Dick-
inson (2003): Francolinus hartlaubi crypticus.

Francolinus gariepensis stresemanni Hoesch & Niet-
hammer, 1940
J Ornithol 88, special issue: 89.

Paratype: ZFMK 39.966, female, Kakatswa, Namibia,
22.08.1938, leg. Hoesch No 1347.

Type specimen at the MfN, Berlin, also two more

paratypes (?). — Dickinson (2003): Francolinus levaillan-
toides levaillantoides (A. Smith, 1836).

©ZFMK

Types in ZFMK: Section Ornithology 31

Tetrao bonasia styriacus Jordans & Schiebel, 1944
Falco 40: 1.

Holotype: ZFMK 57.667, male, St. Georgen ob Judenburg,
Upper Styria, Austria, 01.11.1932, leg. Schiebel.

No paratypes defined in the large series. — Dickinson
(2003): Tetrastes bonasia styriacus.

Tetrao schiebeli Kleinschmidt, 1941
Falco 37: 18.

Lectotype: ZFMK 57.694, male, Zirknitz, Krain, Austria,
11.09.1927, ex. Coll. Schiebel.

No further paratypes defined or described out of a large
series of an unknown number of specimens. — Dickinson
(2003): Tetrastes bonasia schiebeli.

Tetrao Bonasa rhenana Kleinschmidt, 1917
Ornis Germanica (supplement of Falco) 1: 7; nomen
nudum

Tetrao Bonasa rhenana Kleinschmidt, 1944
Falco 40: 4.

Lectotypes: Coll. Kl. 8279, male, Niederhoppen a. d.
Wied, Germany 13. 12. 1897; Coll. Kl. 8280, female,
Niederhoppen a. d. Wied, Germany, 14. 12. 1897; Coll.
Kl. 8281, male, Niederhoppen a. d. Wied, Germany, 14.
12. 1897; Coll. Kl. 8282, female, Brochenbach a. d. Wied,
15.12.1897; Coll. Kl. 8283, male, Neustadt a. d. Wied,
Germany, 03.11.1898, alle leg. Schneider.

Name first mentioned in Ornis Germanica, but a nomen
nudum there. A large type series elaborately described in
Falco (1944: 4), but no type specimen designated and se-
ries not specified. Fourteen skins from ZFMK mentioned,
but not to be identified. — Dickinson (2003): Tetrastes
bonasia rhenana.

FALCONIDAE
Falco caucasicus Kleinschmidt, 1907

Falco 3: 62.

Lectotype: Coll. Kl. 7120, female ad., Wladikaukas, NE
Caucasus, 08.01.1907, per W. Schliiter;

Bonn zoological Bulletin 59: 29-77

Paralectotype: Coll. Kl. 7119, female ad., Prochladnaya,
Terek region, Caucasus, 17.01.1900, per W. Schliiter.

No types were designated in the original publication, in
the Catalogue (Kleinschmidt 1935—1943: 152) eight spec-
imens (Coll. K]. 7121-7125) are listed as caucasicus, Nr.
7120 marked as type in the catalogue, but Nr. 7119 la-
belled as “type specimen”. Illustrated in Berajah, Falco
Peregrinus: plate 27 (1914). — Dickinson (2003): Falco
peregrinus brookei Sharpe, 1873.

Falco rudoifi Kleinschmidt, 1909
Falco 5: 19.

Holotype: Coll. Kl. 7111, female , Hokodadi, N Japan,
04.02.1908, leg. Thielemann. — Dickinson (2003): Falco
peregrinus pealei Ridgway, 1874.

Falco Peregrinus rhenanus Kleinschmidt, 1912
Berajah, Falco Peregrinus: plate 09.

Iconotype: Plate 09 in Berajah.

Lectotype: Coll. KI. 7053, female, rock face ,,Rabenley™,
Altenahr/ Rhineland, 14.05.1897, leg. Kleinschmidt.

The species was described in 1912 by measurements giv-
en on the plate and in 1916 in Berajah (Kleinschmidt 1916:
42-43) in the text.

Several skin specimens are listed in the Catalogue
(Kleinschmidt 1935-1943: 149; 150), Coll. Kl. 7048, Coll.
KI. 7053, Coll.K1. 7059-7074, No. 7053 is designated as
, type specimen“ (Kleinschmidt 1935-1943: 154) and la-
belled as such. — Dickinson (2003): Falco peregrinus pere-
grinus Tunstall, 1771.

Falco Peregrinus scandinaviae Kleinschmidt, 1915
Berajah, Falco Peregrinus: plate 33.

Iconotype: plate 33 in Berajah.

Lectotype: Coll. Kl. 7051, female ad., Upland, Sweden,
03.05.1896, leg. Kolthoff, per Schliiter.

The species was described in 1915 by measurements giv-
en on the plate and in 1916 in Berajah (Kleinschmidt 1916:
46-47) in the text. Eight skin specimens are listed in the
Catalogue (Kleinschmidt 1935-1943: 149-151), Coll. K1.
7051, labelled as “type specimen”. — Dickinson (2003):
Falco peregrinus peregrinus Tunstall, 1771.

©ZFMK

Ww
NO

Falco Peregrinus Heinrichi Kleinschmidt, 1933
Kurzgef deutsche Rassenkunde: 24 and plate 2.

Holotype: Coll. KI. 7143, male, “Celebes”, 24.03.1933,
perished in the Zoological Garden Berlin, leg. Heinrich.
— Dickinson (2003): Falco peregrinus heinrichi.

Falco kreyenborgi Kleinschmidt, 1929
Falco 25: 33-35

Lectotype: Coll. Kl. 7144, male ad., caught at Punta Are-
nas Magallanos, S Chile; received on 22.10.1932 from the
Zoological Garden Minster via Carlos Strauch.

No types were designated in the publication, only in the
Catalogue (Kleinschmidt 1935—1943: 153) and on label.
Two other specimens are housed at the Zoological Muse-
um Minster. This form is assigned today as a light phase
to Falco peregrinus cassini Sharpe, 1873 (Ellis & Garat
1983, Auk 100: 269 et seq.)

Cerchneis tinnunculus canariensis Koenig, 1889
J Ornithol 37: 263.

Lectotypes: ZFMK_ A.II.12.a?.6., male, Villa Orotava,
Tenerife, 6.04.1889 and ZFMK A.II.12.a’.¢., female, Vil-
la Orotava, Tenerife, 10.02.1889, all leg. A. Koenig.

Further 16 specimens from Tenerife and the same year are
listed in the unpublished catalogue of the ZFMK: ZFMK
A.II.12.a?.a, male, Agua Manza, Tenerife, 12.01.1889 (in
the AMNH, New York); ZFMK A.JI.12.a?.B, male, Agua
Manza, Tenerife, 21 02.1889; ZFMK A.II.12.a?-y, male,
Puerto Orotava, Tenerife, 08.01.1889; ZFMK A.II.12.a7.¢,
female, Puerto Orotava, 28.01.1889; ZFMK A.II.12.a?.n,
female, Puerto Orotava, Tenerife, 08.01.1889; ZFMK
A.II.12.a?.6, female juv., Villa Orotava, Tenerife,
10.01.1889; ZFMK A.II.12.a7.1, female, Villa Orotava,
Tenerife, 10.01.1889; ZFMK A.II.12.a?.«, male juv., Icod
el Alto, Tenerife, 15.01.1889; ZFMK A.II.12.a?.X., male,
Orotava, Tenerife, 17.01.1889; ZFMK A.II.12.a?.u, fe-
male, Orotava, Tenerife, 23.01.1889; ZFMK A.II.12.a”.v.,
male juv., Icod el Alto, Tenerife, 27.01.1889; ZFMK
A.II.12.a7.€., female juv., Sta. Ursula, Tenerife,
07.02.1889; ZFMK A.II.12.a?.o., male juv., Villa Orota-
va, Tenerife, 10.02.1889; ZFMK A.II.12.a?.2., female juv.,
Puerto Orotava, Tenerife, 12.02.1889; ZFMK A.II.12.a7.p.,
male, Palma, 25.03.1889; all leg Koenig and ZFMK
A.II.12.a?.¢., female juv., bought from P. Schmitz, Estre-
ito, Madeira, 14.09.1889. Two specimens labelled as syn-
types in the collection and marked in the catalogue. —
Dickinson (2003): Falco tinnunculus canariensis.

Bonn zoological Bulletin 59: 29-77

Renate van den Elzen

Falco tinnunculus ultratinnunculus Kleinschmidt, 1929
Falco 25: 35.

?Holotype: Coll. Kl. 7451, male ad., Wladikaukas, Cau-
casus, 20.10.1913. No collector specified.

Type designated to “the very old male’, no specification
in the text or catalogue.

Paratypes: Coll. Kl. 7452, male ad., Wladikaukas,
21.02.1914; Coll. Kl. 7453 female ad., Wladikaukas,
4.10.1912.— Dickinson (2003): Falco tinnunculus tinnun-
culus Linnaeus, 1758.

Falco Hierofalco erlangeri Kleinschmidt, 1901
Aquila 8: 34—38, figured plate 1 and 3.

Holotype: Coll. KI. 7022, Djebel Sidi Ali ben Aoun, Tu-
nisia, 06.04.1897, leg. Hilgert.

Paratype: Coll. Kl. 7021, male, same locality, 12.04.1897,
leg. Hilgert; Coll. KI. 7023, female ad., Tanger, Moroc-
co, 14.08.1897, per Schliiter; 2 specimens from the Col-
lection of A. Koenig not to be traced; see also Falco Hi-
erofalco tschusii.

The subspecies was first illustrated by A. Koenig, (1898,
J. Ornithol: plate 09, figure 1—5) without him realizing the
new form. Described by Kleinschmidt from a larger se-
ries of six males and five females: two specimens from
the Hungarian National Museum; one from Museum Car-
lo Erlanger and two from living birds in the aviaries of
A. Koenig, Bonn. — Dickinson (2003): Falco biarmicus
erlangeri.

Falco Hierofalco tschusii Kleinschmidt, 1907
Falco 3: 103.

Holotype: Coll. Kl. 7023, female ad., Tanger, Morocco,
14.08.1897, per Schliiter.

In Catalogue defined by Kleinschmidt (1935-1943: 148)
as synonym of Falco biarmicus erlangeri, after consid-
ering it to be a paratype of F: H. erlangeri in 1901. — Dick-
inson (2003): Falco biarmicus tanypterus Schlegel, 1843.

Falco Hierofalco aralocaspius Kleinschmidt, 1939
Falco 35: 28.

Holotype: Coll. Kl. 7012, “female” ad., “Linkoran”,
10.01.1895, per Floericke.

©ZFMK

Types in ZFMK: Section Ornithology 33

In the brief description no details on material used are giv-
en. In the “Katalog” several specimens are recorded as ar-
alocaspius, Coll. Kl. 7009-7011, 7013-7014, Russia, Cau-
casus, but not marked as paralectotypes. — Dickinson
(2003): Falco cherrug cherrug.

Falco christiani-ludovici Kleinschmidt, 1917
Falco 13: 9.

Holotype: Coll. Kl. 7337, male juv., Wladikaukas, Cau-
casus, 12.11.1910, per Schliiter.

Actually considered as nom. nov. for Falco pallidus
(Sushkin, 1900), because Kleinschmidt presumed it to be
preoccupied two times by both Tinnunculus alaudarius
pallidus A.E. Brehm, 1866 and Falco pallidus (Schlegel
and Susemihl, 1839-1851). Contrary to Kleinschmidt,
Brehm’s name is a nomen nudum according to Vaurie
(1965) and Falco pallidus (Schlegel & Susemihl) refers
— like Falco pallidus Schinz, 1840 — to Circus pallidus
Sykes, 1832. — Dickinson (2003): Falco columbarius pal-
lidus (Sushkin, 1900).

Falco alfred-edmundi Kleinschmidt, 1917
Falco 13: 9.

Holotype: Coll. KI. 7341, female, N Iceland, 6.09.1909,
per Schliiter. — Dickinson (2003): Falco columbarius sub-
aesalon C. L. Brehm, 1827.

ACCIPITRIDAE

Accipiter Palumbarius caucasicus Kleinschmidt, 1922
Berajah 4: 5.

Lectotype: Coll. Kl. 7790, male ad., Wladikaukas, Cau-
casus, 11.11.1913;

Paralectotypes: Coll. Kl. 7791, male ad., Wladikaukas,
Caucasus, 15.01.1914; Coll. Kl. 7792, female,
Wladikaukas, 7.03.1914, all per Schliiter.

Described after three specimens with no type designated
in the original description, but only these three skins in
the collection. — Dickinson (2003): Accipiter gentilis mar-
ginatus (Piller & Mitterpacher, 1783).

Astur gentilis arrigonii Kleinschmidt, 1903
Orn. Monatsber. 11: 152.

Bonn zoological Bulletin 59: 29-77

Syntypes: Coll. Kl. 7788, female ?, St. Cosinus, Lanuset,
Sardinia, 14.04.1903; Coll. Kl. 7789, male ad., Leni, Bar-
bagia, Sardinia, 24.04.1902, leg. Meloni.

Described after five specimens, whereabouts of other spec-
imens unknown. — Dickinson (2003): Accipiter gentilis ar-
rigonil.

Accipiter gentilis kleinschmidti Jordans, 1950
Syll Biol Festschr Kleinschmidt: 178.

Holotype: ZFMK 46.343, male ad., Linares, Salamanca,
Spain, 16.05.1943, leg. Griin.

Paratypes: ZFMK 46.344, female, Linares, Salamaca,
Spain, 12.05.1943; ZFMK 46.345, female, Linares, Sala-
manca, Spain, 23.05.1938, all leg. Griin. — Dickinson
(2003): Accipiter gentilis arrigonii (Kleinschmidt, 1903).

Accipiter nisus peregrinoides Kleinschmidt, 1921
In Grote (1921): Aus der orn. Literatur Russlands (sup-
plement to Falco): 56.

Lectotype: Coll. Kl. 7926, female juv., Rossitten,
20.10.1921, leg. Thienemann.

No specification given in first description; mention of an
old female on migration, specimen illustrated in Proceed-
ings IV Intern. Ornithol. Congr. Not tracable. Also an old
male mentioned in first description from the collection of
A. Koenig, without specifying a locality or date. In Klein-
schmidt (1935—1943: 185) 39 specimens are listed under
peregrinoides. — Dickinson (2003): Accipiter nisus nisus
(Linnaeus, 1758).

Accipiter nisus Salamancae Jordans & Steinbacher, 1941
Ann Nat Hist Mus Wien 53: 238.

Holotype: ZFMK 38.528, female ad., Linares de Riofrio,
Salamanca, Spain, 10.06.1938, leg. Griin.

Paratypes: ZFMK 38.527, male, Linares de Riofrio, Sa-
lamanca, Spain, 30.08.1938; ZFMK 38.561, female,
05.08.1938, locality and collector as above;
ZFMK.38.562, female, 05.08.1938, locality and collector
as above; ZFMK A.II.9.a.yyyyyyyy, female, Mosquerue-
la, 06.10.1930; ZFMK A.II.9.a.AAAAAAK, female, Mo-
squeruela, 04.09. 1930; ZFMK A.I1.9.a.g€EEEEEE, male, Li-
nares, 21.08.1937; ZFMK A.II.9.a.0o0000000, female, Li-
naries des Riofrio, 02.08.1937, all leg. Griin. — Dickinson
(2003): Accipiter nisus nisus (Linnaeus, 1758).

©ZFMK

34 Renate van den Elzen

Accipiter wolterstorffi Kleinschmidt, 1901
Orn Monatsber 9: 168.

Holotype: Coll. Kl. 8037, female, Lanusei, Sardinia,
18.12.1900, leg. Wolterstorff.

Paratype: Coll. KI. 8038, pull. male, caught from the aerie,
Lanusei, Baulacci, 19.07.1900, leg. Wolterstorff.

Described from a larger series of another eleven speci-
mens, not specified; paratype identified by label in col-
lection. — Dickinson (2003): Accipiter nisus wolterstorf-

fi.

Accipiter nisus galliae Kleinschmidt, 1917
Falco 13: 24.

Lectotype: Coll. KI. 8032, female juv., Thugny, Ardennes,
France, 22.02.1917, leg. Bacmeister.

Paralectotype: Coll. Kl. 8033, female, Aussonce, Ar-
dennes, 05.01.1917.

No types defined, very sparse description. — Dickinson
(2003): Accipiter nisus nisus (Linnaeus, 1758).

Buteo zimmermannae Ehmcke, 1893
J Ornithol 41: 117.

? Holotype: Coll. Kl. 7606, female, Sassup6nen, Gumbin-
nen, East Prussia, (Russia), 21.04.1892, leg. Schwede.

Also Coll. Kl. 7607, male juv., identified as ““Cotypus”.
Described also from comparative material from the col-
lection of the MfN, Berlin.— Dickinson (2003): Buteo bu-
teo buteo (Linnaeus, 1758).

Buteo vulgaris hispaniae Jordans, 1940
Falco)35213,

Syntypes: ZFMK A.II.19.a°.a. and B., 2 males, Linares,
Salamanca, Spain, 27.04.1935 and Teruel, Mosqueruela,
18.09.1930, leg. Griin.

Paratypes: ZFMK A.II.19.a°y., female, Mosqueruela,
05.09.1930; ZFMK A.II.19.a°.6., female, Linares, 26.07.
1934; ZFMK A.II.19.a°.¢., female, Linares, 24.08.1934;
ZFMK 38.519, female, Linares, 02.07.1938; ZFMK
38.866, male, Linares, 05.08.1938, all leg. Griin. — Dick-
inson (2003): Buteo buteo buteo (Linnaeus, 1758).

Bonn zoological Bulletin 59: 29-77

Buteo vulgaris vulpicolor Kleinschmidt, 1938
Falco 34: 72.

Holotype: Coll. Kl. 7597, 0, Pipret Swamps, Russia,
14.08.1917, leg. Riidiger. — Dickinson (2003): Buteo bu-
teo vulpinus (Gloger, 1833).

Buteo vulgaris ultravulpinus Kleinschmidt, 1938
Falco 34: 73.

Holotype: Coll. Kl. 7619, Caucasus, lost. — Dickinson
(2003) presumably: Buteo buteo menetriesi Bogdanow,
1879.

Buteo vulgaris milanoides Kleinschmidt, 1938
Falco 34: 73.

Holotype: Coll. KI. 7628, ,” female, Naljtschik, 07.” Cau-
casus, (without year), lost. — Dickinson (2003): Buteo bu-
teo menetriesi Bogdanow, 1879.

Buteo vulgaris buteosimilis Kleinschmidt, 1938
Falco 34: 73.

Lectotypes: Coll. Kl. 7593, male juv., Bukowina, Roma-
nia, 19.09.1892, leg. Tschusi; Coll. Kl. 7594-7595, mat-
ed pair, 01.05.1871, Uman, S Russia, leg. Holtz, per C.
v. Erlanger.

Neither in the original description nor in the Catalogue
(Kleinschmidt 1935—1943) type specimens were defined;
lectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Buteo buteo buteo (Linnaeus,
1758).

Circus pygargus abdullae Floericke, 1896
Orn Monatsber 4: 155.

Lectotype: Coll. KI. 8097, male ad., Kermene, E Kaspi,
Turkmenia, 22.05.1896, leg. Floericke.

Described from a series of eight specimens, of which on-
ly four were skinned; Lectotype designated by Klein-
schmidt (1935-1943: 190). According to the label three
further specimens mentioned in the original description
are housed in the museum at Tiflis. Therefore, this spec-
imen should come from the Berlepsch collection.

©OZFMK

Types in ZFMK: Section Ornithology 35

OTIDIDAE

Choriotis arabs geyri Niethammer, 1954
Bonn zool Beitr 5: 193.

Lectotype: ZFMK 54.565, male, 250 km NW of Agades,
Niger (Sahara). 30.01.1954, leg. G. Niethammer No 511;
Second specimen of the description lost. — Dickinson
(2003): Choriotis arabs geyri.

Afrotis afra bohmeri Hoesch & Niethammer, 1940
J Ornithol 88; Sonderheft: 124.

Holotype: ZFMK 39.789, male, E side of the Brandberg,
Namibia, 23.09.1938, leg. G. Niethammer. — Dickinson
(2003): Afrotis afra (Linnaeus, 1758).

RALLIDAE

Gallinula chloropus vestigialis Clancey, 1939
Bull BOC 59: 70.

Paratype: ZFMK 50.131, male, Renfrewshire, Scotland,
15.01.1936, leg. Clancey.

Syntypes in Collection Clancey, RSME, Edinburgh and
Collection Ticehirst; t.t. Near Bardowie, Stirlingshire,
Scotland. A large series of unspecified skins from various
museums was investigated. — Dickinson (2003): Gallinu-
la chloropus chloropus (Linnaeus, 1758).

HAEMATOPODIDAE

Haematopus ostralegus occidentalis Neumann, 1929
Gefiederte Welt 58: 161.

Holotype: ZFMK M.VI.14.a'.a., male, Auskerry, Orkney
Islands, 03.10.1913, leg. Ticehurst No 1491. Type defined
by Neumann not until 1932 (Anz Orn Ges Bayern 2: 148).

— Dickinson (2003): Haematopus ostralegus ostralegus
Linnaeus, 1758.

CHARADRIIDAE

Charadrius hiaticula harrisoni Clancey, 1949
Limosa 22: 319.

Bonn zoological Bulletin 59: 29-77

Paralectotypes: ZFMK 50.318—50.319, male, female, Dal-
iburgh and Bornish, Outer Hebrides, 17. and 23.06.1947,
leg. Clancey, in exchange for other specimens handed to
the ZFMK.

In the original description only the holotype is designat-
ed. Status of ZFMK specimens taken from inventory and
labels. If not included in Ch. h. hiaticula Linné, 1758, one
might regard this form as nom. nov. for the preoccupied
Charadrius hiaticula major Seebohm, 1885. Its type ought
to be regarded then as lectotype for Seebohm’s subspecies,
for which no type specimen exists. Type stored within the
Collection Clancey, RSME, Edinburgh, t.t. Rudha Ard-
vule, Bornish, South Uist, Outer Hebrides. — Dickinson
(2003): Charadrius hiaticula hiaticula Linnaeus, 1758.

GLAREOLIDAE

Rhinoptilus africanus erlangeri Niethammer & Wolters,
1966
Bonn zool Beitr 17: 159.

Holotype: ZFMK 39.736, female ad., 40 km E Cape Cross,
Namibia, 25.09.1938, leg. G. Niethammer No 139.

Paralectotypes: ZFMK 39.737, female, Erongo,
28.10.1938; ZFMK 39.738, female, Omaruru,
27.09X.1938; ZFMK M..II.3.a.a., male, Kayas,

13.07.1934; ZFMK M.II.3.a.B., female, Kayas 15.07.1934;
ZFMK G.II.3.a.y., female, Ombutjomatemba, 30.08.1934;
ZFMK M.II.3.a.6., male, Ombutjomatemba, 08.03.1935,
all leg. Hoesch; ZFMK 66.660, female, Hohenfels, Wa-
terberg, 27.05.1931, leg. Maltzahn.

Described from a series of 14 specimens, of which five
paratypes are not in the ZFMK. The name Rhinoptilus a.
sharpei Erlanger, 1905, doesn’t correspond to the sub-
species above, it is in fact a synonym of Rhinoptilus
africanus africanus (Temminck, 1807). — Dickinson
(2003): Rhinoptilus africanus erlangeri.

ALCIDAE

Mormon arcticus meridionalis Jordans, 1914
Falco 10: 153.

Syntypes: ZFMK Q.I.7.a7.y., male, Bahia de Alcudia, Ma-
jorca, 07.04.1913; ZFMK Q.1.7.a. 6., female, Bahia de
Alcudia, Majorca, 07.04.1913; leg. A. von Jordans, No
999-1000. — Dickinson (2003): Fratercula arctica grabae
(C. L. Brehm, 1831).

©ZFMK

36 Renate van den Elzen

Uria aalge hyberborea Salomonsen, 1932
Ibis Ser. XII. 2: 130.

Holotype: ZFMK Q.I.1.a?, o., female ad., Bear Island,
Scandinavia, 16.07.1907, leg. Koenig.

Paratypes: Of the 55 paratypes mentioned only 30 still ex-
ist; some of them are part of the permanent exhibition (dio-
rama “Vogelfelsen” — “Bird rock”). — Dickinson (2003):
Uria aalge hyberborea.

PTEROCLIDIDAE

Pterocles lichtensteini targius Geyr, 1916
Orn Monatsber 24 : 56.

Lectotypes: ZFMK L.I.1.c?.B. and 6., female, Oued Ra-
ris, Algeria, 16.04.1914. leg. Geyr.

Paralectotypes: 11 more individuals, ZFMK L.I.1.c?.a.,
male ad., Oued Raris, Algeria, 16.04.1914; ZFMK
L.I.1.c*. y. male, Oued Raris, 16.04.1914; ZFMK
L.I.1.c*.e., male, Amgid, Algeria, 21.04.1914; ZFMK
L.L.1.c?.¢, male, Amgid, Algeria, 21.04.1914; ZFMK
L.I.1.c3.y., female, Amgid, Algeria, 21.04.1914; ZFMK
L.I.1.c*.6., male, Amgid, Algeria, 21.04.1914; ZFMK
L.IL.1.c3.., male, Oued Tidebar, Algeria, 24.04.1914 ;
ZFMK_ L.I.1.c?.«., male, Oued Tidebar, Algeria,
24.04.1914; ZFMK L.I.1.c?.A., male, Oued Tidebar, Alge-
ria, 25.04.1914; ZFMK L.I.1.c?.u., male, Oued Tidebar,
Algeria, 25.04.1914; ZFMK L.I.1.c?.v., female, Oued Ti-
debar, Algeria, 25.04.1914; all leg. Geyr.

Originally described from a series of thirteen specimens.
The type specimen labelled as such in the inventory of the
ZFMK, — ZFMK L.I.1.c3.a., male ad., Oued Raris,
16.04.1914, — doesn’t correspond to the type specimen
named in the original description concerning location and
date. The original description indicates a male, Quelle
Tahart, 21.02.1914 — this specimen is apparently lost. —
Dickinson (2003): Prerocles lichtensteini targius.

Pterocles senegallus remotus Neumann, 1934
Verh Orn Ges Bayern 20: 471.

Paratype: ZFMK I.1.2.a'.B., female, Kulchi, Kunaria, W
India, 20.02.1934, leg. Archer.

Holotype: male, Kunaria at Kutch, Nw-India, 20.02.1934,
leg G. Arcker; supposedly at the MfN, Berlin.

Bonn zoological Bulletin 59: 29-77

Described from a series of four males and two females,
whereabouts of other specimens not known. — Dickinson
(2003): Pterocles senegallus (Linnaeus, 1771).

COLUMBIDAE

Streptopelia turtur loéi Jordans, 1923
Falco 19, Sonderheft: 5.

Holotype: ZFMK K.I.4.a7.a. male, Albufera, Majorca,
07.06.1921., leg. A. von Jordans No 2914.

Paratypes: ZFMK K.1.4.a?.B., male, Alcudia, Majorca, 10.
05.1921; ZFMK K.I.4.a?. y., female, Alcudia, Majorca,
06.05.1921; ZFMK K.I.4.a?. 6., female, 31.05.1921; all
leg. A. von Jordans. — Dickinson (2003): Streptopelia tur-
tur turtur (Linnaeus, 1758).

Turtur turtur hoggara Geyr, 1916
Orn Monatsber 24: 59.

Holotype: ZFMK K.1.4.b'.a., male ad., Ideles, at the north-
ern edge of the Hoggara Plateau, Algeria, 24.03.1914, leg.
Geyr. Described from one single specimen. — Dickinson
(2003): Streptopelia turtur hoggara.

Columba livia targia Geyr, 1916
Orn Monatsber 24: 58.

Holotype: ZFMK K.I.1.a>2.a., male ad., Ain Tahart, Al-
geria, 21.02.1914;

Paratypes: ZFMK K.I.1.a>2.B., female, Ain Tahart, Alge-
ria, 21.02.1914; ZFMK K.I.1.a>2.y., male, Ideles, Algeria,
25.03.1914; ZFMK K.I.1.a>2.6., male, Amgid, Algeria,
14.02.1914; ZFMK K.I.1.ab2. ¢., female, Ain Tahart, Al-
geria, 21.02.1914; ZFMK K.I.1.a>2.C., female, Amgid, Al-
geria, 14.02.1914; ZFMK K.I.1.a>2.n., male, Ain Tahart,
Algeria, 24.04.1914; ZFMK K.I.1.a52.0., female, Amgid,
Algeria, 14.02.1914; ZFMK K.I.1.a>2.1., female, Ain Ta-
hart, Algeria, 21.02.1914; all leg. Geyr.

In the collection of the ZFMK eight specimens are regis-
tered as fargia, five males and three females, in the de-
scription only three males and two females are mentioned.
— Dickinson (2003): Columba livia targia.

OZFMK

Types in ZFMK: Section Ornithology 37

CUCULIDAE

Centropus heuglini Neumann, 1911
In Koenig, 1910: Verhandl. V. Intern. Orn. Kongr. Berlin:
504.

?Paratypes: ZFMK C.1.3.b.a.—6., 2 males, female, male
juv., Bahr-el-Gazal, Lado and Bahr-el-Zeraf, 28.02; 07.,
12. and 20.03.1910, Egyptian Sudan, leg. le Roi, Koenig,
Nolda.

The specimens listed above were labelled as syntypes by
Koenig in the ZFMK inventory; their status as paratypes
is questionable. Type specimen is the female of the SMNS,
Stuttgart, No 5662 collected by Heuglin at Bahr el Ghaz-
al, 02.1866, another paratype No 1916, a juvenile speci-
men from the collection of Herzog Paul v. Wiirttemberg,
Fazoqla, leg. 1843. — Dickinson (2003): Centropus
monachus fischeri Reichenow, 1887.

TYTONIDAE

Strix ernesti Kleinschmidt, 1901
Orn Monatsber 9: 168.

Holotype: Coll. KI. 6945, male, Loceri Ogliastra, Sardi-
nia, 17.04.1901.

Paratypes: Not defined, the Catalogue (Kleinschmidt
1935-1938) lists eight specimens as potential paratypes
for S. ernesti — Dickinson (2003): Tyto alba ernesti.

Tyto alba kleinschmidti Jordans, 1923
Falco 19, Sonderheft: 5.

Holotype: ZFMK A.II.13.a°.y., male, Alcudia, Majorca,
09.05.1921, leg. A. von Jordans No 3098.

Paratypes: ZFMK A_III.13.a°.a., male, Alcudia, Majorca,
07.04.1913; ZFMK A.III.13.a°.B., female, Alcudia, Ma-
jorca, 09.05.1913; ZFMK A.III.13.a°y., male, Alcudia,
Majorca, 09.05.1921; ZFMK A.III.13.a°.6., female, Alcu-
dia, Majorca, 22.05.1921; ZFMK A.III.13.a®.¢., male, Al-
cudia, Majorca, 24.05.1921; ZFMK A.III.13.a°.¢., male,
Alcudia, Majorca, 09.05.1921. — Dickinson (2003): Tyto
alba alba (Scopoli, 1769).

Bonn zoological Bulletin 59: 29-77

Strix hostilis Kleinschmidt, 1915
Falco 11: 18.

Syntypes: Coll. KI. 6952, male, Hurst, Sussex, England,
08.1894, and Coll. Kl. 6953, female, Bramber, Sussex,
(with parts of the skeleton), 10.09.1897, both per
Brazenor, Brighton. —Dickinson (2003): Tyto alba alba
(Scopoli, 1769).

Strix flammea rhenana Kleinschmidt, 1906
Berajah, Strix Flammea: 4: 20.

Lectotype: Coll. KI. 6920, female, Kornsand, Rhine near
Darmstadt, 03.11.1899, leg. Kleinschmidt.

Paralectotypes: Coll. Kl. 6921-6930, 4 males, 6 females,
same location, 10.1892; 03., 10., 11.1899, 09., 10.1901 and
04.1903, leg. Kleinschmidt.

Named by Kleinschmidt because Brehm’s type (see be-
low) is thought to refer to a migrant not native to Middle
Germany. No type designated in the original description;
type series designated in the catalogue (Kleinschmidt
1935-1943: 147); lectotype by Rheinwald & van den
Elzen (1984). — Dickinson (2003): Tyto alba guttata (C.L.
Brehm, 1831).

Tyto alba (Strix Flammeata) hauchecornei Klein-
schmidt, 1940
Falco 36: 59.

Syntype: Coll. Kl. 6978, 0, caught in Chile, perished in
the Cologne Zoological Garden.

The other syntype is housed in the SNS, Dresden, C
48381. — Dickinson (2003): Tyto alba tuidara (J. E. Gray,
1829).

STRIGIDAE
Otus leucotis Margarethae Jordans & Neubaur, 1932

Falco 28: 9.

Holotype: ZFMK A.III.10.e'.a., male, Zankab, Bahr-el-
Abiad, Sudan, 07.04.1910, leg. le Roi.

Paratypes: ZFMK A.III.10.e'.B., male, Zankab, Bahr-el-
Abiad, Sudan, 07.04.1910 ; ZFMK A.II.10.e!.y., female
ad., Zankab, Bahr-el-Abiad, Sudan, 07.04.1910. — Dickin-
son (2003): Ptilopsis leucotis (Temminck, 1820).

©ZFMK

38 Renate van den Elzen

Otus scops mallorcae Jordans, 1923
Falco 19: 5.

Holotype: ZFMK A.III.10.a'.¢., female, Alcudia, Majorca,
22.V.1921, leg. A. von Jordans No 3045.

Paratypes: ZFMK A.III.10.a'.B., male, Alcudia, Majorca,
05.04.1913; ZFMK A.HI.10.a'. y., male, Alcudia, Major-
ca, 07.04.1913; ZFMK A.III.10.a'.6., female, Alcudia, Ma-
jorca, 09.04.1913; ZFMK A.III.10.a'. ¢., male, Alcudia,
Majorca, 09.06.1921; ZFMK A.HI.10.a'. n., male, Alcu-
dia, Majorca, 11.06.1921; ZFMK A.II.10.a'. 0., male, Al-
cudia, Majorca, 13.06.1921, all leg. A. von Jordans.

Originally a series of eleven specimens is mentioned; one
exchanged with AMNH, New York, two specimens are un-
tracable. — Dickinson (2003): Otus scops mallorcae.

Athene noctua Griini Jordans & Steinbacher, 1941
An. Nat Hist Mus Wien 52: 234.

Holotype: ZFMK 40.238, female ad., Lagos, Portugal,
13.04.1939, leg. Grin.

Paratypes: ZFMK 40.230, male, Lagos, Portugal,
18.05.1939; ZFMK 40.231, female, Lagos, Portugal,
01.05.1939; ZFMK 40.232, female, Lagos, Portugal,
25.04.1939; ZFMK 40.233, female, Lagos, Portugal,
02.05.1939; ZFMK 40.234, female, Lagos, Portugal,
27.03.1939; ZFMK 40.235, female, Lagos, Portugal,
18.04.1939; ZFMK 40.236, male, Lagos, Portugal,
22.04.1939; ZFMK 40.237, female, Lagos, Portugal,
17.04.1939; ZFMK 40.239, female, Lagos, Portugal
19.05.1939, all leg. Griin. — Dickinson (2003): Athene
noctua vidalii A. E. Brehm, 1857.

Strix Athene sarda Kleinschmidt, 1907
Falco 3: 65.

Lectotype: Coll. Kl. 6748, female, Iini, Sardinia,
18.10.1901.

Paralectotype: Coll. Kl. 6749, female, Ogliastra, Sardinia,
04.04.1907.

Described from three specimens, no type specimen defined
in the original description, in Catalogue (Kleinschmidt
1935-1943: 143) only two specimens mentioned with sta-
tus as Holo- and Cotypus. — Dickinson (2003): Athene noc-
tua noctua (Scopolhi, 1769).

Bonn zoological Bulletin 59: 29-77

Strix Athene ruficolor Kleinschmidt, 1907
Falco 3: 65.

Lectotype: Coll. Kl. 6754, male, Marrakesch, Morocco,
28.12.1899; leg. Floericke.

Paralectotypes: Coll. Kl. 6755, female juv., Drenkat,
12.12.1899; Coll. Kl. 6756, 0 ad., Doukalla, 17.11.1899,
all leg. Floericke.

Described from nine specimens, no type designated in
original description or in his Catalogue, Kleinschmidt
(1938-1943: 144). — Dickinson (2003): Athene noctua
glaux (Savigny, 1809).

Strix saharae Kleinschmidt, 1909
Falco 5: 19.

Holotype: Coll. Kl. 6757, 0, Moulaina, Algerian Sahara,
20.03.1903, leg. Flukiger?

Paralectotypes: Coll. Kl. 6758, male, Biskra, Algeria,
12.12.1903; Coll. Kl. 6759, male, Biskra, Algeria,
15.03.1904; Coll. Kl. 6760, male, Biskra, Algeria,
30.01.1903, all leg. Fliikiger.

Only type designated in original description, no number
of paratypes given; paralectotypes designated in his Cat-
alogue (Kleinschmidt 1938-1943: 144). — Dickinson
(2003): Athene noctua saharae.

Bubo africanus kollmannspergeri Niethammer, 1957
Bonn zool Beitr 8: 287.

Holotype: ZFMK 57.1900, male, Guelta Basso, Ennedi,
11.09.1954, leg. Kollmannsperger.

Described from a single specimen. — Dickinson (2003):
Bubo cinerascens Guérin-Meéneville, 1843.

Strix aluco clanceyi Jordans, 1950
Syll Biol, Festschr. Kleinschmidt: 176.

Holotype: ZFMK A.III.12.a’.¢., male, Linares, Salaman-
ca, Spain, 03.04.1935, leg. Grin.

Paratypes: ZFMK A.III.12.a?.a., male, Mosqueruela near
Teruel, Spain, 23.09.1930; ZFMK A.III.12.a?.B., female,
Mosqueruela near Teruel, Spain, 29.09.1930; ZFMK
A.III.12.a?.y., male, Mosqueruela near Teruel, Spain,
09.09.1930; ZFMK A.III.12.a?.6., male, Mosqueruela near
Teruel, Spain, 29.09.1930; ZFMK A.III.12.a?.¢., female,

©ZFMK

Types in ZFMK: Section Ornithology 39

Linares, Spain, 07.04.1935; ZFMK 36.518, male, Linares,
Salamanca, Spain, 16.03.1938; ZFMK 46.294, female, La-
gunilla, Spain, 25. 12. 1940; ZFMK 46.295, female,
Linares, Spain, 25.12.1940; ZFMK 46.296, female,
Linares, Spain, 26.07.1941; all leg. Griin. — Dickinson
(2003): Strix aluco sylvatica Shaw, 1809.

Glaucidium bolivianum Konig, 1991
Okol Végel 13: 36, Abb. 10, p. 38.

Paratype: ZFMK 54.684, male, brown morph, Lambate,
La Paz province, Bolivia, 2300m, 14.09.1953.

The holotype is at the SMNS, Stuttgart. — Dickinson
(2003): Glaucidium bolivianum.

Glaucidium peruanum KO6nig, 1991
Okol Végel 13: 57, Abb. 29, p. 58.

Paratypes: ZFMK 54.1398, female, Lima, Peru,
01.12.1945; ZFMK 60.191, female, w Andean slopes, C
Peru, 1500m.

The holotype is stored at the BMHM, Tring. — Dickinson
(2003): Glaucidium peruanum.

CAPRIMULGIDAE

Nyctiphrynus ocellatus bergeni Niethammer, 1953
Bonn zool Beitr 4: 285.

Holotype: ZFMK 52.229, male ad., Quiquibey, Bolivia,
13.09.1951, leg. G. Niethammer.

Described from one specimen. — Dickinson (2003): Nyc-
tiphrynus ocellatus ocellatus (Tschudi, 1844).

Caprimulgus inornatus malbranti Niethammer, 1957
Bonn zool Beitr 8: 279.

Holotype: ZFMK 57.1901, male ad., rock between Wadi
Rei and Wadi Basso, Ennedi, Chad, 03.09.1957, leg. Koll-
mannsperger.

Of four males and four females collected, only the holo-

type has been prepared. — Dickinson (2003): Caprimul-
gus inornatus Heuglin, 1869.

Bonn zoological Bulletin 59: 29-77

APODIDAE

Streptoprocne zonaris kiinzeli Niethammer, 1953
Bonn zool Beitr 4: 286.

Holotype: ZFMK 52.231, male, Puri (1200 m), Yungas of
La Paz, Bolivia, 30.10.1951, leg. Ktinzel.

Paratypes: ZFMK 52.232, female, Puri (1200 m), Yungas
of La Paz, Bolivia, 30.10.1951; ZFMK 53.49, female, Puri
(1200 m), Yungas of La Paz, Bolivia, 18.11.1952, ZFMK
53.50, female, Puri (1200 m), Yungas of La Paz, Bo-
livia,18.11.1952, all leg. Kiinzel. — Dickinson (2003):
Streptoprocne zonaris kuenzeli.

TROCHILIDAE

Phaethornis koepckeae Weske & Terborgh, 1977
Condor 79: 143-147.

Paratype: ZFMK 83.66, female, Cerros del Sira, Cabez-
eras del Rio Llullapichis, Selva tropical, ca. 650 m AM-
SL, Peru, 24.07.1971, leg. Koepcke and Terborgh. Orig-
inal No Kp 1959c.

Holotype at the AMNH, No 820872, female ad., Cerros
del Sira, 9°30’S, 74°47°W, Depto. Huanuco, Peru. Two
remaining paratypes are stored in Hamburg (ZMH) and
Lima (MHN Javier Prado). — Phaethornis koepckeae is
systematically allocated in the proximity of Phaethornis
bourcieri and Phaethornis philippii. — Dickinson (2003):
Phaethornis koepckeae.

Thalurania nigricapilla Valdés-Velasquez & Schuch-
mann, 2009
Ornithol Anz 48: 144—145., Abb. 2

Holotype: ZFMK 81.384, adult male, Lake Calima (La-
guna de Calima, Embalse de Calima), (03°50’N,
76°30’W), 50km N of Cali, Valle del Cauca, Colombia,
leg. Schuchmann.

Paratype: ZFMK 81.383, adult male, forest NW of Yoto-
co on road to Puente Tierra at Lake Calima, vivinity of
Alto de los Torros (03°53’N, 76°27’ W), Valle del Cauca,
Colombia, leg. Schuchmann.

Only these two specimens were examined.

©ZFMK

40 Renate van den Elzen

Trochilus Adelomyia intergrediens Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 222.

Holotype: Coll. Kl. 8957, Central Cordillera, Rio Toche,
Quindiu-Pass and Canon del Monte Tolima, 1700-2400
m AMSL, Colombia, 20.09.1909, leg. Fassl.

Paralectotypes: Coll. Kl. 8958-8962, data like holotype.

In the short original description only the holotype is des-
ignated; paralectotypes designated by Schuchmann (1984).
— Dickinson (2003) presumably: Adelomyia melanogenys
cervina Gould, 1872.

Trochilus Adelomyia ultracervinus Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 222.

Holotype: Coll. K1. 8963, St. Antonio, W Cordillera, 2100
m AMSL, Colombia, 08.06.1909, leg. Fassl.

Paralactotypes: Coll. Kl. 8964-8968, data like holotype.

In the short original description only the holotype is des-
ignated; paralectotypes designated by Schuchmann (1984).
— Dickinson (2003) presumably: Adelomyia melanogenys
cervina.

Lafresnaya lafresnayi longirostris Schuchmann, Weller
& Wulfmeyer, 2003
Orn Neotrpical 14: 160.

Holotype: Coll. Kl. 9569, Madim, Paso del Quindio,
2500m, Colombia, 29.09.1909, leg. A. H. Fass.

Paratypes: Coll. Kl. 9570, Rio Toche, Paso del Quindio,
Tolima, 2100m, 10.10.1909; Coll. Kl. 9572, Volcancito,
Paso del Quindio, all leg. Fassl.

Coeligena lutetiae albimaculata Sanchez-Osés, 2006
Bull Brit Orn Cl 126: 6-7.

Holotype: ZFMK 85.87, male, Crater of Volcan Pichin-
cha, Ecuador (00°10’S, 78°33’ W), deposited by Schréder
in the Dernedde Collection November 1912.

Paratypes: ZFMK 85.76, 85.78-79, 3 males, Santo
Domingo de los Colorados, Ecuador; ZFMK 85.81—84, 4
females, Santo Domingo de los Colorados, Ecuador;
ZFMK 8589, Campamento Pailon, Carchi, Ecuador
(00°29°S, 77°55’ W), all deposited in the Dernedde Col-
lection by Schréder in November 1912.

Bonn zoological Bulletin 59: 29-77

Eustephanus galeritus robinsoni Dernedde, 1943
Falco 37: 18.

Lectotype: Coll. Dernedde 10248, male, Juan Fernandez
Islands, Chile, 04.1927, leg. Castillo.

See next taxon.
In the short original description no type specimen was de-

fined. Lectotype designated by Schuchmann (1984). —
Supposedly Dickinson (2003): Sephanoides fernandensis

fernandensis (King, 1831).

Eustephanus derneddei Kleinschmidt, 1943
Falco 37: 18.

=Eustephanus galeritus robinsoni Dernedde, 1943 — re-
named by Kleinschmidt, 1941 as Eustephanus derneddei
because of Ornysmia (=Eustephanus=Sephanoides) robin-
soni Delattre & Lesson, 1839. Type specimen Coll. Der-
nedde 10248. — Supposedly Dickinson (2003): Sepha-
noides fernandensis fernandensis (King, 1831).

Trochilus Exortis fassli Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 225.

Lectotype: Coll. KI. 9723, male ad., Monte Socorro, Ca-
li, W Cordillera, 3800 m AMSL, Colombia, 27.07.1909,
leg. Fassl.

Paralectotypes: Coll. Kl. 9724-9727, 4 males, locality like
holotype.

In the very short description Coll. Kl. Nos 9723-9727 are
referred to the new taxon, no specifications are given. Lec-
totype and paralectotypes designated by Schuchmann
(1984). — Dickinson (2003): Heliangelus exortis (Fraser,
1840).

Trochilus Caligula incommodus Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 225.

Lectotype: Coll. Kl. 9670, male ad., St. Antonio, W Cor-
dillera, 2000 m AMSL, Colombia, 17.04.1909, leg. Fassl.

Paralectotypes: Coll. KI. 9671-9699, 6 males, 23 females,
Sta. Magarita, W Cordillera, Colombia.

In the very short description Coll. Kl. Nos 9723-9727 are
referred to the new taxon, no specifications are given. Lec-
totype and paralectotypes desig. by Schuchmann (1984).
— Dickinson (2003): Ocreatus underwoodii incommodus.

©OZFMK

Types in ZFMK: Section Ornithology 4]

Ramphomicron microrhynchum bolivianum Schuch-
mann, 1984
Bull Brit Orn Cl 104: 7.

Holotype: ZFMK 81.367, male ad., Cocapata, Bolivia,
07.1980, leg. Cordier.

No paratypes exist. — Dickinson (2003): Ramphomicron
microrhynchum bolivianum.

Chalcostigma herrani tolimae Kleinschmidt, 1927
Falco 23/1: 5.

Holotype: Coll. KI. 9764, male ad., Monte Tolima, 4000
m AMSL, Colombia, 02.1910, leg. Fassl.

Paratypes: Coll. KL. 9765—9767, male ad., male immat.,
female, locality like holotype. — Dickinson (2003): Chal-
costigma herrani tolimae.

COLIIDAE

Colius macrourus laeneni Niethammer, 1955
Bonn zool Beitr 6: 56.

Holotype: ZFMK 54.468, male, Baguezan, 150 km N Aga-
des, Niger, 08.02.1954, leg. Laenen.

Paratypes: ZFMK 54.557 and 54.1051, 2 females, N
Agades, Niger, 31.01.1954, leg. Laenen.

Described from a series of four males and three females;
only three specimens found in the collection of ZFMK. —
Dickinson (2003): Urocolius macrourus laeneni (Niet-
hammer, 1955).

ALCEDINIDAE

Alcedo ispida, var. taprobana Kleinschmidt, 1894
Orn Monatsber 2: 126.

Lectotype: Coll. Kl. 6608, male, Kalawewe, Ceylon,
03.04.1889, leg. Geisler, per Schliiter.

Paralectotype: Coll. Kl. 6609, male, Kalawewe, Ceylon,
28.03.1889, leg. Geisler, per Schliiter.

The original description doesn’t indicate a holotype. It was
defined by Kleinschmidt (1935-1943: 140) in his Cata-
logue. — Dickinson (2003): Alcedo atthis taprobana.

Bonn zoological Bulletin 59: 29-77

MEROPIDAE

Merops orientalis flavoviridis Niethammer, 1955
Bonn zool Beitr 6:53.

Holotype: ZFMK 54.1031, male ad., Archei, Ennedi,
Chad, 09.04.1954, leg. G. Niethammer No 241.

Paratypes: ZFMK 54.556, male, N Agades, Niger (Sa-
hara), 04.02.1954 ; ZFMK 54.1030, male, Archei, Ennedi,
Chad, 10.04.1954; ZFMK 80.360, male, Archei, Ennedi,
Chad, 11.04.1954, all leg. G. Niethammer.

Described from a series of three males from Ennedi and
four males from Agades; three further specimens untrace-
able. — Dickinson (2003): Merops orientalis viridissimus
Swainson, 1837.

PICIDAE

Campethera tullbergi poensis Eisentraut, 1968
Bonn zool Beitr 19: 52.

Holotype: ZFMK 63.1342, female ad., Moka Valley, Fer-
nando Poo, 21.01.1963, leg. Eisentraut No 879.

Paratype probably in SMNS, Stuttgart. — Dickinson
(2003): preoccupied by Campethera nivosa poensis
Alexander 1903; Campethera tullbergi tullbergi (Sj6stedt,
1892).

Dendrocopus minor hispaniae Jordans, 1939
Falco 34: 52.

Holotype: ZFMK 38.508, male ad., Linares de Riofrio, Sa-
lamanca, Spain, 02.04.1938, leg. Griin.

Paratypes: ZFMK 38.509, male, Linares de Riofrio, Sala-
manca, Spain, 30.03.1938; ZFMK 38.511, female, Linares
de Riofrio, Salamanca, Spain, 25.03.1938; ZFMK 38.512,
male, Linares de Riofrio, Salamanca, Spain, 27.12.1934;
ZFMK 38.514, male juv., Linares de Riofrio, Salamanca,
Spain, 08.07.1936; ZFMK. 38.515, female juv., Linares
de Riofrio, Salamanca, Spain, 09.07.1936; ZFMK 38.516,
female, Lagunilla, Salamanca, Spain, 18.01.1934, all leg.
Grin.

ZFMK 38.512 and 38.513 exchanged with Natural His-

tory Museum, Warsaw. — Dickinson (2003) presumably:
Dendrocopos minor buturlini (E. Hartert, 1912).

©ZFMK

42 Renate van den Elzen

Dryobates minor Heinrichi Jordans, 1940
Mitt K6nigl Naturwiss Inst Sofia, 13: 131.

Holotype: ZFMK 38.614, male ad., Kamtschyja, Bulga-
ria, 06.06.1935, leg. Heinrich, PMZ Warsaw No 290.

Paratype: ZFMK 38.615, same locality, 08.06.1935, leg.
Heinrich.

Erronneously, registration number ZFMK 38.875 was pub-
lished for the type specimen in the original description. —
Dickinson (2003): Dendrocopos minor buturlini (E. Har-
tents ID):

Picus minor bacmeisteri Kleinschmidt, 1916
Falco 11: 14.

Lectotype: Presumably Coll. KI. 6463, male, La Tourau-
derie, Departement des Ardennes, France, 11.03.1916, leg.
Bacmeister.

Described from three males and one female. In the orig-
inal description no type specimen was defined, specimen
even not labelled as type; lectotype designated by Rhein-
wald & van den Elzen (1984). — It is the only specimen
with this name, in both the Collection Kleinschmidt in
Bonn and in the Catalogue (Kleinschmidt 1935-1943:
137). — Dickinson (2003) presumably: Dendrocopos mi-
nor hortorum (C. L. Brehm, 1831).

Picus Minor silesiacus Kleinschmidt, 1917
Ornis Germanica (supplement of Falco) 1: 5.

Lectotype: Coll. Kl. 6452, male, Rothhaus a.d. Neisse,
Silesia, 13.04.1894.

The very brief description “between Picus Minor typo-mi-
nor and hortorum” gives no type specimen. A single type
specimen was designated by Kleinschmidt (1935-1943:
136) in his Catalogue, no paralectotypes were defined. —
Dickinson (2003): Dendrocopos minor hortorum (C. L.
Brehm, 1831).

Picus major arduennus Kleinschmidt, 1915
Falco: 1 02:

Lectotypes: Coll. Kl. 6396, male, Departement des Ar-
dennes, France, 04.04.1916; Coll. Kl. 6397, female, La
Tourauderie, Departement des Ardennes, France,
04.04.1916, leg. Bacmeister.

Bonn zoological Bulletin 59: 29-77

No types defined in the description, only specified in Cat-
alogue (Kleinschmidt 1935-1943: 135). — Dickinson
(2003): Dendrocopos major pinetorum (C. L. Brehm,
1831).

Picus major hispanus Schliiter, 1908
Falco 4: 11.

Syntypes: Coll. KI. 6398, male, Sevilla, Spain, 14.01.1907;
Coll. KI. 6399 female, Espera, 12.02.1907; Coll. KI. 6400
female, Sevilla, 13.09.1906, per W. Schliiter.

The third female defined in the description is not in the
Coll. Kl. The male is labelled as “typus”. — Dickinson
(2003): Dendrocopos major hispanus (Schliter, 1908).

Dendrocopos maior thanneri le Roi, 1911
Orn Monatsber 19: 81.

Syntypes: ZFMK E.I.5.a°.¢., male, Mogan, Gran Canaria,
03.1909, and ZFMK E.1.5.a5t., female, Mogan, Gran Ca-
naria, 03.1909, leg. Thanner.

Paratypes: ZFMK E.1.5.a°.a., male, Mogan, Gran Canaria,
03.1909; ZFMK E.I.5.a5.y., male, Gran Canaria, 03.1909;
ZFMK E.I.5.a5.6., male, Mogan, Gran Canaria, 03.1909;
ZFMK E.I.5.a9.¢., female, Mogan, Gran Canaria, 03.1909;
ZFMK E.I.5.a5.n., female, Mogan, Gran Canaria, 03.1909;
ZFMK E.I.5.a°.0., female, Mogan, Gran Canaria, 03.1909;
ZFMK E.1.5.a9.«., male, Mogan, Gran Canaria, 04. 1909;
ZFMK E.1L.5.a5.4., Mogan, Gran Canaria, 04.1909, all leg.
Thanner.

Described from six males and six females, ZFMK
E.1.5.a5.B., exchanged with the AMNH (New York). The
syntypes are a breeding pair. — Dickinson (2003): Dendro-
copos major thanneri.

Picus (Dendrocopus) canariensis Koenig, 1889
J Ornithol 37: 263.

Syntypes: ZFMK E.I.5.a4.a. and B., male, female, Pinar
of Icod de la Vinos, Tenerife, 23.04. and 17.02.1889, leg.
Koenig.

Paratype: ZFMK E.I.5.a*-y., male, same locality as types,
26.01.1889.

In the original description the t. t. 1s indicated as “Canary
Islands”; it is restricted by Rheinwald & van den Elzen
(1984) to Tenerife. The type specimens have been shown
in the exhibition of the museum and were prepared as

OZFMK

Types in ZFMK: Section Ornithology 43

skins not before 1943. One paratype, ZFMK E.I.5.a+.6,
was exchanged with the AMNH, New York. — Dickinson
(2003): Dendrocopos major canariensis.

Dryobates major beicki Stresemann, 1927
Orn Monatsber 35: 134.

Paratype: Coll. Kl. 6417, female, Tschan-to, Kansu, Chi-
na, 30.1.1927, leg. Beick, No 337.

Type specimen at the MfN, Berlin, No 27.194, t. t. Lan-
hukou, Kansu, China. — Dickinson (2003): Dendrocopos
major beicki (Stresemann, 1927).

Mesopicos goertae koenigi Neumann, 1903
Orn Monatsber 11: 181.

Holotype: ZFMK E.1.9.a!.a., male ad., S Mutir, “Egypt-
ian Sudan”, 16.03.1903, leg. Koenig, No 375.

Description apparently based on only the type specimen

— Dickinson (2003): Dendropicos goertae koenigi (Neu-
mann, 1903).

Jynx torquilla tschusii Kleinschmidt, 1907
Falco 3: 103.

Holotype: Coll. Kl. 6560, male, Lanusei, Ogliastra, Sar-
dinia, 20.01.1907.

Obviously description from only this single specimen. —
Dickinson (2003): Jynx torquilla tschusii.
COTINGIDAE

Zaratornis stresemanni Koepcke, 1954

Publ Mus Hist Nat Javier Prado 16: 1.

Paratype: ZFMK 55.103, female, Zarate near San Bar-
tolomé. 60-70 km to Lima, Peru, 11.10.1953, leg.

Koepcke, No Kp 1019 h.

Holotype at the MHN Javier Prado, Lima, No Kp 1019 1.
t. t. as above. — Dickinson (2003): Zaratornis stresemanni.

Bonn zoological Bulletin 59: 29-77

FURNARITDAE

Synallaxis zimmeri Koepcke, 1957
Publ Mus Hist Nat Javier Prado 18: 1.

Paratype: ZFMK 56.1069, female, Colcabamba, tributary
river of Rio Cosma, 9°36’S, Peru, 17.08.1956, leg.
Koepcke, No Kp 1438 n.

Described from four specimens. Holotype MHN Orn
0584, Kp 1438 p, at the MHN Javier Prado, Lima, t. t. as
paratype; 2°¢ paratype MHN Orn. O585, Kp 1438 0, at
the MHN Javier Prado, Lima; 3" paratype KP 1450 a in
the AMNH, New York. — Dickinson (2003): Synallaxis
zimmeri.

Asthenes pudibunda grisior Koepcke, 1961
Am Mus Novitates 2028: 13.

Paratype: ZFMK 60.171, female, upper Pisco valley, be-
tween Huarhac and Manzanallo, Peru, 25.11.1957, leg.
Koepcke, No 1482 a.

Holotype at the MHN Javier Prado, Lima, Orn 0624, Kp
1371 s, t. t. Chuquibamba, Peru. — Dickinson (2003): As-
thenes pudibunda grisior.

Asthenes cactorum Koepcke, 1959
Beitr neotrop Fauna |: 243.

Paratype: ZFMK 56.1038, male, Pacaychacra, below
Chuquibamba, department of Arequipa, Peru, 30.01.1956,
leg. Koepcke, No 1374 b. See next taxon.

Described from a series of eight specimens. Holotype: at
the MHN Javier Prado, Lima, Kp 1341f. Atico, southern
Peru, 08.01.1956. — Dickinson (2003): Asthenes cactorum
cactorum.

Asthenes cactorum monticola Koepcke, 1965
Beitr neotrop Fauna 4: 162

Paratype: ZFMK 56.1038, see previous taxon.

Described from ten specimens. Holotype at the AMNH,
New York, male, George No 1742, t.t.24 km N Arequipa,
Peru. Our paratype has previously been a paratype of the
newly described species cactorum. — Dickinson (2003):
Asthenes cactorum monticola.

©ZFMK

44 Renate van den Elzen

Asthenes cactorum lachayensis Koepcke, 1965
Beitr neotrop Fauna 4: 164

Paratype: ZFMK 67.189, male, Lomas of Lachay, ca. 90
km N Lima, Peru. 24.10.1959, leg. Koepcke, No 1640 a.

Described from a series of eight specimens. Holotype:
MHN Orn 1902, Kp 1677 f at the MHN Javier Prado, Li-
ma, t. t. as above; two further paratypes at the AMNH,
New York and four at the LSUMZ. — Dickinson (2003):
Asthenes cactorum lachayensis.

Cranioleuca antisiensis zaratensis Koepcke, 1961
Publ Mus Nat Hist. Javier Prado 20: 1.

Paratype: ZFMK 55.108, female, Zarate near San Bar-
tolomé, 11°54’S, Peru, 10.X.1953, leg. Koepcke, No KP
1019 d.

Holotype MHN Orn 0641, KP 917 u, at the MHN Javier
Prado, Lima, t. t. as above. — Dickinson (2003) presum-
ably: Cranioleuca antisiensis palamblae (Chapman,
1923):

LANIDAE

Lanius excubitor galliae Kleinschmidt, 1917
Falco 13: 24.

Lectotype: Coll. Kl. 2161, male, Thenorgues, Departement
des Ardennes, France, 25.02.1916, leg. Dobritz, per
Bacmeister.

No type specimen designated in the poor description or
in the Catalogue (Kleinschmidt 1935—1943: 56). Lecto-
type designation by Rheinwald & van den Elzen (1984),
based on label in collection. It is uncertain whether the
specimens Coll. Kl. 2162—2164 may be considered as
paralectotypes. — Dickinson (2003): Lanius excubitor ex-
cubitor Linnaeus, 1758.

Lanius Excubitor tingitanus Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 57.

Lectotype: Coll. Kl. 2186, male, Tanger, Morocco,
08.05.1885, leg. Olcese;

Paralectotype: Coll. Kl. 2187, ? female, Tanger,
07.11.1898, leg. Floericke.

Only very poor specification in Catalogue given (Klein-
schmidt 1935-1943: 57), no types designated; lectotype

Bonn zoological Bulletin 59: 29-77

by Rheinwald & van den Elzen (1984), based on hand-
written information on label in collection. — Dickinson
(2003) presumably: Lanius excubitor excubitor Linnaeus,
1758.

Lanius senator fliickigeri Kleinschmidt, 1907
Falco 3: 68.

Lectotype: Coll. Kl. 2259, male, Lambése, Algeria,
1.06.1904, leg. Flikiger.

? Paralectotypes: Coll. Kl. 2257-2261, male, 2 females,
0, as type specimen, 06.1904 and 05.1903, leg. Fliikiger.

Kleinschmidt erroneously wrote Fliikiger with “ck”; no
type designated in first description; lectotypes by Rhein-
wald & van den Elzen (1984), based on label in collec-
tion. — Dickinson (2003) presumably: Lanius senator sen-
ator Linnaeus, 1758.

Lanius senator erlangeri Kleinschmidt, 1919
Falco 14: 16.

Lectotype: Coll. KI. 2264, male, Tunis, ,,April“, without
year, bought 21.02.1916 from Schliiter. Very poor descrip-
tion, type only designated in the Catalogue (Kleinschmidt
1935-1943: 58). — Dickinson (2003) presumably: Lanius
senator senator Linnaeus, 1758.

Lanius senator weigoldi Kleinschmidt, 1919
Falco 14: 16.

Lectotype: Coll. KI. 2243, male, Malaga, Spain, date hard-
ly legible, 01.or 07.03.1904.

Poor description, no material specified, no type specimen
designated; lectotype designated from the Catalogue
(Kleinschmidt 1935—1943: 58). — Dickinson (2003): La-
nius senator rutilans Temminck, 1840.

Lanius Senator italiae Kleinschmidt, 1922
Falco 18: 24.

Holotype: Coll. KI. 2244, male, Monte del Casentino, Tu-
scany, Italy, 16.07.1922.

Paratype: Coll. KI. 2245, male, Foggia, Italy, 25.04.1921.

Described from a series of six specimens, four from the
private collection of Schliiter. — Dickinson (2003): Lanius
senator senator Linnaeus, 1758.

©ZFMK

Types in ZFMK: Section Ornithology 45

Lanius collurio juxtus Clancey, 1951
Bonn zool Beitr 2: 83.

Paratype: male, Martlesham, Woodbridge, E Suffolk, Eng-
land, 31.05.1941, said to be held at the ZFMK lost; holo-
type in the Clancey collection, RSME, Edinburgh.

Described from a series of eight specimens; British pop-
ulation probably extinct. — Dickinson (2003): Lanius col-
lurio collurio Linnaeus, 1758.

Laniarius fuelleborni camerunensis Eisentraut, 1968
Bonn zool Beitr 19: 55.

Holotype: ZFMK 68.12, male, above Dikume-Balue,
Rumpi Hills, Cameroon, 25.02.1967, leg. Eisentraut No
1068.

Paratypes: ZFMK 61.1427, male, Cameroon Mountains,
09.02.1958; ZFMK 61. 1428, female, Cameroon Moun-
tains, 20.11.1957; ZFMK 61.1429, male, Cameroon
Mountains, 09.11.1957; ZFMK 68.952, male, Mt. Kupe,
01.12.1966; ZFMK 68.953, male, Lake Manenguba,
30.12.1966; ZFMK. 68.954, male, Dikume, Rumpi Hills,
22.02.1967; ZFMK 68.955, male, Dikume, Rumpi Hills,
23.02.1967; ZFMK 68.956, male, Dikume, Rumpi Hills,
25.02.1967; ZFMK 68.957, male, Dikume, Rumpi Hills,
02.03.1967; ZFMK 68.958, female, Dikume, Rumpi Hills,
17.02.1967, all leg. Eisentraut.

Described from a larger series of twenty-one males and
ten females, rest of series in SMNS, Stuttgart? — Dickin-
son (2003): Laniarius poensis camerunensis.

ORIOLIDAE

Oriolus chlorocephalus speculifer Clancey, 1969
Durban Museum Novit 8: 249 (April 1969)

Bull Brit Orn Cl 89: 108 (August 1969)

Holotype: ZFMK 68.2695, female, Gorongoza Mountain,
Mozambique, 25.10.1968, leg. Baddeley.

Announced as newly described subspecies from six spec-
imens in Durban Museum Novitates and described from
seven specimens in Bull Brit Orn Cl. Type specimen des-
ignated in August 1969 in the Bull Brit Orn Cl. Paratypes
DMD 24.675—78 and DMD 24.680-81, 4 males, 2 fe-
males, Gorongoza Mountain, 21.—24.10.1968, leg. Bad-
deley, at the DMD (Durban). The publication in the Dur-
ban Mus Novit was not meant to be an original descrip-

Bonn zoological Bulletin 59: 29-77

tion, rather the announcement of a new form, which was
released only “inadvertently” before the publication in the
Bull. Brit. Orn. Club. — Dickinson (2003): Oriolus chloro-
cephalus speculifer.

CORVIDAE

Pica galliae Kleinschmidt, 1917
Falco 13: 24.

Lectotype: Coll. Kl. 5681, male pull., Briquenay, Depar-
tement des Ardennes, France, 21.05.1916, leg. Bacmeis-
ter.

Paralectotypes: ? Coll. Kl. 5682-5689, 2 males, 4 females,
Departement Ardennes and Meuse, 22.09.1916 until
16.02.1917, leg. Bacmeister.

Not indicated as paratypes, listed in one series along with
No 5681; the latter designated as type specimen in the Cat-
alogue (Kleinschmidt 1935—1943: 119). No types defined
in the sparse description. — Dickinson (2003): Pica pica
pica (Linnaeus, 1758).

Corvus corax kretae Jordans & Steinbacher, 1943
Senckenbergiana 26: 73.

Holotype: ZFMK 46.21, male, Kanea, Crete, 26.02.1925,
leg. Schiebel.

Paratypes: ZFMK 57.676—57.681, female, 5 males, all Ka-
nea, Crete, 26.02.1925, leg. Schiebel.

Described from seven specimens. — Dickinson (2003) pre-
sumably: Corvus corax laurencei Hume, 1873.

Corvus corax jordansi Niethammer, 1953
Bonn zool Beitr 4: 74.

Holotype: ZFMK F.VIL3.g7-y., (No 10.325), male ad., Oli-
va, Fuerteventura, Canary Islands, 03.1912, leg. Thanner.

Paratypes: ZFMK_ F.VII.3.g’.a., B and ZFMK
F.VIL.3.g7.6—-., 5 males, 6 females, locality as type speci-

men.

Described from twelve specimens. — Dickinson (2003):
Corvus corax canariensis Hartert & Kleinschmidt, 1901.

©ZFMK

46 Renate van den Elzen

Corvus sardus Kleinschmidt, 1903
Orn Monatsber 11: 92.

Lectotype: Coll. KI. 5319, male, Boulvai, Ogliastra, Sar-
dinia, 09.03.1902, leg. Wolterstorff.

Paralectotype: Coll. KI. 5318, male, Ulassai, Ogliastra,
Sardinia, 18.01.1902, leg. Wolterstorff; uncertain paralec-
totypes: Coll. Kl. 5320-5325.

No types defined in the description, No 5319 only in 1943
designated as type in Catalogue (Kleinschmidt
1935-1943: 112). — Dickinson (2003) presumably: Corvus
corax corax Linnaeus, 1758.

Corvus corax hispanus Hartert & Kleinschmidt, 1901
Novit Zool 8: 45.

?Paratype: Coll. Kl. 5330, male ad., Murcia, Spain,
09.1855 or 1856 (unreadable), leg. A. E. Brehm.

Holotype supposedly at the BM, Tring (Hartert & Klein-
schmidt, 1901), t.t. Aguilhas, near Murcia. — Dickinson
(2003): Corvus corax hispanus.

Corvus bacmeisteri Kleinschmidt, 1919
Falco 14: 8.

Lectotype: Coll. KI. 5466, female juv., bill, claw and left
wing, Schwittersdorf, Saxony, 31.03.1917, leg. Klein-
schmidt

Designated as type and defined in the Catalogue (Klein-
schmidt 1935-1943: 116) as variation (aberratio) of
Corvus corone cornix Linnaeus, 1758.

Corvus sardonius Kleinschmidt, 1903
Orn. Monatsber. 11: 92.

Lectotype: Coll. Kl. 5481, ? female, Ogliastra, Sardinia,
09.01.1900, leg. Wolterstorff.

Poor description, type designated only in Catalogue
(Kleinscmidt 1935-1943: 116). If other specimens from
catalogue are paralectotypes is unclear. — Dickinson
(2003): Corvus corone sharpii Oates, 1889.

Corvus ultracollaris Kleinschmidt, 1919
Falco 14: 16.

Lectotype: Coll. Kl. 5663, male, Naryn, Tien Shan,
24.03.1910, per Schliiter.

Bonn zoological Bulletin 59: 29-77

Poor description; in Catalogue (Kleinschmidt 1935-1943:
119) type specimen designated, no other skins listed un-
der taxon name. — Dickinson (2003): Corvus monedula
soemmerringii J. G. Fischer, 1811.

Nucifraga caryocatactes wolfi Jordans, 1940
Mitt Konig! Naturwiss Inst Sofia 13: 63.

Holotype: ZFMK 38.605, male, Karlik, Rhodope Moun-
tains, Bulgaria, 25.08.1935, leg. Heinrich, Museum War-
saw, Inv. No 909.

Paratypes: ZFMK 38.606, female, Karlik, Rhodope
Mountains, Bulgaria, 26.08.1935 leg. Heinrich; ZFMK 38.
607, male, Pirin Mountains, Bulgaria, 06.06.1938; ZFMK
68.608, male, Pirin Mountains, Bulgaria, 07.06.1938;
ZFMK 68.609, male juv., Pirin Mountains, Bulgaria,
07.06.1938; ZFMK 68.610, female juv., Pirin Mountains,
Bulgaria, 07.06.1938; ZFMK 68.611, male, Pirin Moun-
tains, Bulgaria, 09.06.1938; ZFMK 68.612, female, Pirin
Mountains, Bulgaria, 10.06.1938; —ZFMK 38.613, male,
Pirin Mountains, 10.06.1938, leg. A. von Jordans. — Dick-
inson (2003): Nucifraga caryocatactes caryocatactes (Lin-
naeus, 1758).

Nucifraga hemispila interdictus Kleinschmidt & Wei-
gold, 1922
Falco 18: 2.

Paralectotype: Coll. K1. 5862, male, Tschili, protective for-
est, E of Peking, 02.09.1916.

Lectotype (labelled in handwriting): C23062 P,
02.09.1916, Hung-m6énntsze, forest reserve east of Bei-
jing, NO Tschili, 1400m, leg. Schott.

Lectotype in the SNS, Dresden, designated by Eck &
Quaisser (2004); paralectotype designated by Rheinwald
& van den Elzen (1984). — Dickinson (2003): Nucifraga
caryocatactes interdicta.

Garrulus glandarius hansguentheri Keve, 1967
Bull Brit Orn Cl 87: 40.

Paratype: ZFMK 51.10, 0, Cekmece, Turkey, 01.03.1947,
leg. Kosswig;

Holotype at the BMNH, Tring, No 1886, female, Tash-
Cupri, near Constantinople, 12.01.1922.

Other specimens consulted in ZFMK and BMNH, Tring.
— Dickinson (2003): Garrulus glandarius anatoliae See-
bohm, 1883.

©ZFMK

Types in ZFMK: Section Ornithology 47

Garrulus hilgerti Kleinschmidt, 1940
Falco 36: 8.

No type defined in the description, the Kleinschmidt Cat-
alogue (1935-1943: 121) or in the collection. Coll. Kl.
5868, female, Neustadt a.d. Wied, 22.05.1896, a breed-
ing bird with a clutch of three eggs Coll.KI. 5868a, are
the only specimens registered under that name. — Dickin-
son (2003): Garrulus glandarius glandarius (Linnaeus,
1758).

Garrulus ichnusae Kleinschmidt, 1903
Orn Monatsber 11: 92.

Lectotype: Coll. Kl. 5893, male, Ogliastra, Sardinia, 30.
12. 1902, leg Meloni, per Wolterstorff.

Paralectotype: ZFMK F.VII.8.a!.y., male ad., Ogliastra,
Sardinia, 03.02.1903, leg. A. von Jordans No 2811.

No type specimen was defined in the original description;
Lectotype designated from label in the collection (Rhein-
wald & van den Elzen 1984) as Catalogue (Kleinschmidt
1935-1943: 122) does not specify which specimen from
the series is the type. Paralectotype marked as such in the
inventory book of the ZFMK collection. — Dickinson
(2003): Garrulus glandarius ichnusae.

Garrulus albipectus Kleinschmidt, 1920
Falco 16: 28.

Lectotype: Coll. KI. 5898, male ?, Florence, 12. 1914, per
Schliiter.

Paralectotype: Coll. Kl. 5899, 0, Legi near Florence, no
date, per Schliiter.

The third specimen mentioned in the description doesn’t
exist anymore. No type specimen designated in descrip-
tion, or in Catalogue (Kleinschmidt 1935—1943: 122);
identified only from label (Rheinwald & van den Elzen
1984). — Dickinson (2003): Garrulus glandarius albipec-
tus.

Garrulus lambessae Kleinschmidt, 1912
Falco 8: 51.

Lectotype: Coll. Kl. 5921, male, Lambése, Algeria,
02.06.1904, leg. Fliikiger.

Paralectotypes: Coll. Kl]. 5922, male, Lambése, Cheliah,
Algeria, 01.06.1905; Coll. KI. 5923, male pull., Lambe-
se, Cheliah, Algeria, 01.07.1903, leg. Fliikiger.

Bonn zoological Bulletin 59: 29-77

No type specimen defined in the original description, on-
ly in Catalogue (Kleinschmidt 1935—1943: 123). — Dick-
inson (2003) presumably: Garrulus glandarius cervicalis
Bonaparte, 1853.

PARIDAE

Parus cristatus hispanus Jordans, 1933
Anz Orn Ges Bayern 2: 257.

Holotype: ZFMK G.X.1.g*.y., female, Mosqueruela near
Teruel, Spain, 02.10.1931, leg. H. Grin.

Paratypes: ZFMK G.X.1.g*.a., male, Mosqueruela near
Teruel, 19.12.1930; ZFMK G.X.1.g?.B., male, Mosque-
ruela near Teruel, 30.12.1930; ZFMK G.X.1.g?.6., female,
Mosqueruela near Teruel, 05.01.1931; ZFMK G.X.1.g7.C.,
male, Mosqueruela near Teruel, 07.10.1931; ZFMK
G.X.1.g?.u., male, Mosqueruela near Teruel, 06.10.1931;
ZFMK G.X.1.g?.«., female, Mosqueruela near Teruel,
07.10.1931; ZFMK G.X.1.g?.A., male, Mosqueruela near
Teruel, 02.01.1931; ZFMK G.X.1.g?.u., male, Mosque-
ruela near Teruel, 02.01.1931; ZFMK G.X.1.g?.v., 0, Mos-
queruela near Teruel, 02.01.1931, all leg. Griin.

Described from a series of thirteen specimens; three ex-
changed with AMNH, New York and an illegible institu-
tion. — Dickinson (2003): Parus cristatus weigoldi Tratz,
1914.

Parus cristatus bureschi Jordans, 1940
Mitt Konig] Naturwiss Inst Sofia 13: 90.

Holotype: ZFMK 38.718, female ad., Pirin Mountains
(above Bansko), Bulgaria, 05.06.1938, leg. Wolf.

Paratypes: ZFMK 38.719, female, Pirin Mountains,
04.06.1938; ZFMK 38.720, 0, Pirin Mountains,
05.06.1938; ZFMK 38. 721—38.724, 4 0, Pirin Mountains,
06.06.1938, leg. Wolf.

Described from twelve specimens; whereabouts of other

paratypes unknown. — Dickinson (2003): Parus cristatus
bureschi.

Parus Lophophanes nominandus Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 106.

Nomen nudum there, as it lacks description.

OZFMK

48 Renate van den Elzen

Parus rhenanus Kleinschmidt, 1900
Orn Monatsber 8: 168.

Holotype: Coll. Kl. 4536, male, banks of the River Rhine
between Mainz and Worms, 04.10.1894, leg. Lorentz.

Paralectotypes: Supposedly Coll. KI. 4537-4569, Korn-
sand, Ingelheim, Renthendorf, Guntershausen,
Knoblochsau, between 1894 and 1899, leg. Kleinschmidt.

Paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Parus montanus rhenanus.

Parus salicarius submontanus Kleinschmidt & Tschusi,
1913
Falco 9: 33.

Paralectotype: Coll. Kl. 4597, male, Gmunden, Upper
Austria, 6.1.1913, leg. Tschusi.

Holotypes at the NHMW, Vienna, breeding pair
29.03.1913 from Gmunden, ex. Coll. Tschusi.

Paralectotype designated on label, whole series from Coll.
Kl. 4597— 4603 referred to as “cotypes”. In original de-
scription only two specimens specified. — Dickinson
(2003): Parus montanus montanus Conrad von Balden-
stein, 1827.

Parus salicarius subrhenanus Kleinschmidt & Jordans,
1916
Falco 12: 17.

Holotype: Coll. Kl. 4512, female juv., foothills of Bonn,
06.12.1913, leg. A. von Jordans.

Paratype: Coll. K1. 4513, male juv., River Sieg near Bonn,
03.06.1911, leg. A. von Jordans No 116. — Dickinson
(2003): Parus montanus rhenanus Kleinschmidt, 1900.

Parus borealis tischleri Kleinschmidt, 1917
Falco 13: 23.

Lectotype: Coll. Kl. 4587, male, near Bartenstein, East
Prussia, 25.12,1916, leg. Tischler.

Paralectotypes: Coll. KI. 4588-4591, male, 2 females, o,
Bartenstein, 11.1909, 1910, 12.1916, leg. Tischler.

Very poor description, “between borealis and salicarius’’;
type specimens designated in the Catalogue (Kleinschmidt
1935-1943: 102). — Dickinson (2003): Parus montanus
salicarius C. L. Brehm, 1831.

Bonn zoological Bulletin 59: 29-77

Parus Salicarius natorpi Kleinschmidt, 1917
Falco 13: 23.

Lectotype: Coll. Kl. 4575, male, Emanuelsegen, Silesia,
17.10.1909, leg. Natorp.

Paralectotypes: Supposedly Coll. Kl. 4576-4582, 3
males, 2 females, 2 0, Emanuelsegen, Myslowitz,
Cieskowice, Bitschin, Roschkowitz, Silesia, collected in
02. 04.1909, 10.1909, leg. Natorp.

Very poor description with no types designated; lectotype
designated only in Catalogue (Kleinschmidt 1935-1943:
101), specimen labelled as type. — Dickinson (2003) pre-
sumably: Parus montanus borealis Selys-Longchamps,
1843.

Parus Salicarius supermontanus Kleinschmidt, 1921
Berajah, Parus Salicarius: 17.

Lectotype: Coll. Kl. 4634, male, Trient, Cembra valley,
Italy, 26.11.1897, leg. Marchi.

Paralectotypes: Presumably Coll. Kl. 4635-4640, 3 fema-
les, 3 0, Cembra valley, Trient, 11.1897, leg. Marchi.

Very poor description with no types designated; lectotype
designated only in Catalogue (Kleinschmidt 1935-1943:
102), specimen labelled as type. — Dickinson (2003) pre-
sumably: Parus montanus salicarius C. L. Brehm, 1831.

Parus Salicarius transsylvanicus Kleinschmidt, 1921
Berajah, Parus Salicarius: 17.

Lectotype: Coll. KI. 4644, male, surroundings of Kron-
stadt, Transylvania, Romania, 28.12.1896, leg. Hausmann,
per Schliiter.

Paralectotypes: ZFMK G.X.1.m!°.a., (20.186), male, da-
ta like type specimen, and Coll. Kl. 4645-4649, 4 males,
female, 08.1896, Kronstadt, leg. Hausmann, per Schliiter.

Very sparse description with no types designated; lecto-
type designated only in Catalogue (Kleinschmidt
1935-1943: 102), specimen labelled as type. — Dickinson
(2003): Parus montanus montanus Conrad von Balden-
stein, 1827.

Parus Salicarius schiebeli Kleinschmidt, 1937
Berajah, Parus Salicarius: 36.

Lectotype: Coll. KI. 4623, male, Zirknitz, Krain, Slowe-
nia, 03.01.1926, leg. Schiebel.

©ZFMK

Types in ZFMK: Section Ornithology 49

Paralectotypes: presumably Coll. Kl. 4624-4633, 5
males, 4 females, 0, all Zirknitz and surroundings,
09.1902, 1903, 11.1926, 12.1923 and 01.1924 and 1926,
leg. Schiebel. Further 37 specimens from the Coll.
Schiebel, possibly including ZFMK 38.139-38.159,
1926-1927, Zirknitz, leg. Schiebel.

No types designated in the original description; lectotype
designated only in Catalogue (Kleinschmidt 1935-1943:
102), specimen labelled as type (see also Schifter & van
den Elzen 1986). — Dickinson (2003): Parus montanus
montanus Conrad von Baldenstein, 1827.

Paris Salicarius styriacus Kleinschmidt, 1937
Berahjah, Parus Salicarius: 36.

Lectotype: Coll. KI. 4605, male, St. Georgen ob Juden-
burg, Styria, Austria, 23.10.1926, leg. Schiebel.

Paralectotypes: Coll. Kl. 4606-4622, 5 males, 9 females,
3 0, Bruck a. d. Mur, Turracher Hohe, Tamsweg, St. Wolf-
gang 1. Bachergebirge, Austria, leg. Schiebel, and 49 spec-
imens of the Coll. Schiebel, probably all at the ZFMK.
10.1919, 0.9-12.1921, 04.1922, 10., 11., 1926. — Dickin-
son (2003): Parus montanus montanus Conrad von Bal-
denstein, 1827.

Parus Salicarius weigoldi Kleinschmidt, 1921
Berajah, Parus Salicarius: 19.

Paralectotypes: Coll. Kl. 4697-4699, 2 males, female,
Atentsze, Mekong tributary, Janeti; Sungpan, E Tibet, 25.
and 29.09.1915 and 29.05.1914, leg. Weigold.

Described from a series, no types designated in first de-
scription or in Catalogue (Kleinschmidt 1935—1943: 103).
Lectotype designated in Eck & Quaisser (2004), lectotype
at the SNS, Dresden C 23803, t.t. Mauntschi, Szechuan.
— Dickinson (2003) probably: Parus montanus baicalen-
sis (Swinhoe, 1871).

Parus weigoldicus Kleinschmidt, 1921

Falco 17: 3, Abb. (Tafel) 2 in Falco 34: 62/63.
Paralectotypes: Coll. Kl. 4697-4699, s. previous taxon.
Nomen novum for Parus Salicarius weigoldi Klein-
schmidt, 1921, preoccupied by Parus weigoldi Tratz,

1914. — Dickinson (2003): Parus montanus baicalensis
(Swinhoe, 1871).

Bonn zoological Bulletin 59: 29-77

Parus Salicarius stotzneri Kleinschmidt, 1921
Berajah, Parus Salicarius: 20.

Paralectotype: Coll. KI. 4696, male, 30 km n. Balihandien,
NE Tschili, China, 30.04.1916, leg. Weigold, per Stotz-
ner.

Described from a series, no type designated. Lectotype
designated in Eck & Quaisser (2004), Lectotype at the
SNS, Dresden C 23845, t.t. as above. Two more paralec-
totypes in MfN, Berlin and AMNH, New York. — Dick-
inson (2003) probably: Parus montanus baicalensis
(Swinhoe, 1871).

Parus communis jeholicus Kleinschmidt & Weigold,
1922
Falco 18: 2.

Paralectotypes: Coll. Kl. 4484-4485, 2 males, 30 km n.
Balihandien, NE Tschili, 29.04. and 03.05.1916, leg. Wei-
gold, per St6tzner.

Described from a series, but no information on specimens
given in original description. In Catalogue (Kleinschmidt
1935-1943: 100) the two specimens mentioned above are
designated as “cotypes”. Eck & Quaisser (2004) describe
a series of eight specimens at the SNS, Dresden as “syn-
types”; one specimen lost in 1945, t. t. north of Jehol. —
Dickinson (2003): Parus palustris jeholicus.

Parus sardus Kleinschmidt, 1903
Orn Monatsber 11: 186.

Holotype: Coll. Kl. 4766, male, Arzana, Ogliastra, Sardi-
nia, 13.10.1903, leg. Wolterstorff.

Three paratypes listed in the description are apparently
lost. — Dickinson (2003): Parus ater sardus.

Parus ater pinicolus Clancey, 1943
Bull Brit Orn Cl 63: 66.

Paratypes: ZFMK 50.112, male, Sutherlandshire, N Scot-
land, 31.08.1938; ZFMK 50.113, female, Sutherlandshire,
N Scotland, 09.09.1938; ZFMK 50.114, male, Sutherland-
shire, N Scotland, 05.09.1938; ZFMK 50.115, female,
Sutherlandshire, N Scotland, 31.08.1938, all leg. Clancey.

Holotype at the RSME, Edinburgh, t. t. Rothiemurchus

Forest, Inverness-Shire. — Dickinson (2003): Parus ater
britannicus Sharpe & Dresser, 1871.

©ZFMK

50 Renate van den Elzen

Parus ater martensi Eck, 1998
Zool Abhandl Staatl Mus Tierk Dresden 50 (8): 130.

Holotype: ZFMK 95.24, male, Chadziou, Khola-Tal,
Thakkhola, Mustang district, Nepal, 2600m, 30.10.1969,
leg. J. Martens.

Paratypes: ZFMK 74.459-—74.463, 5 males, Thakkola,
Thaksang above Tukche, 3150m, 08.07.1973, 10.03.1974;
ZFMK 74.467, 0 juv, Thakkola, Tkaksang obove Tukche,
02.07.1973; ZFMK 74.471—74.472, 2 females, Thakko-
la, Thaksang above Tukche, 3150m, 10 and 11.03.1974;
ZFMK 74.464—74.465, 2 males, Thakkola, Purano Marpha
(Alt Marpha), 3200 m, 15. and 16.03.1974; ZFMK
74.468—74.469, 2 females, Thakkola, Purano Marpha (Alt
Marpha), 3200 m, 16. and 19.03.1974; ZFMK 74.470, fe-
male juv., Thakkhola, at Nabrikot-Khola, 08.07.1973;
ZFMK 2006.41, female, Thakkola, Tukche, 3100m,
23.11.1969; ZFMK 2006.43, male, Parbat District between
Deorali and Chitre, 2850m, 02.05.1995; ZFMK 2006.44,
male, Mustang District, Purano Marpha, 3200 m,
10.05.1995, all leg. Martens.

Described from a series of 20 specimens; for the
paratypes no collection numbers were provided in the orig-
inal description, but date and finding locality; other
paratypes are in the bird collection of the BMNH, Tring.
The subspecies is already figured in Martens (1993: 43,
fig. d) and Martens & Eck (1995: plate 3, fig. d). — Dick-
inson (2003): Parus ater martensi.

Parus caeruleus touraudericus Bacmeister & Klein-
schmidt, 1920
J Ornithol 68: 101.

Lectotype: Coll. Kl. 4206, male, La Tourauderie, Dept. Ar-
dennes, France, 18.02.1916, leg. Bacmeister.

Paralectotypes: Coll. Kl. 4205, 4207-4210, 5 0, La Tou-
rauderie and Vitarville, Meuse, 02., 03., 10.1916, leg.
Bacmeister.

No types defined in the original description based on
twelve specimens; lectotype designated from the Cata-
logue (Kleinschmidt 1935-1943: 98) and handwriting on
label. Whereabouts of other six specimens unknown. —
Dickinson (2003): Parus caeruleus caeruleus Prazak,
1894.

Parus coeruleus balearicus Jordans, 1913
Falco 9: 43.

Holotype: ZFMK G.X.1.b®°.C¢., female, Valldemosa, Ma-
jorca, Baleares, 11.03.1913, leg. A. von Jordans No 622.

Bonn zoological Bulletin 59: 29-77

Paratypes: ZFMK G.X.1.b°.a. female, Palma, Majorca,
10.03.1913; ZFMK G.X.1.b°.B., female, Valldemosa, Ma-
jorcea, 11.03.1913; ZFMK G.X.1.b®.y., male, Lluch, Ma-
jorca, 22.03.1913; ZFMK G.X.1.b®¢., male, Palma-
Valldemosa, Majorca, 10.03.1913; ZFMK G.X.1.b°.n.,
male, Lluch, Majorca, 24.03.1913; ZFMK G.X.1.b°.6.,
male, Esporlas, Majorca, 20.05.1913; ZFMK G.X.1.b®.1.,
female, Esporlas, Majorca, 20.05.1913, all leg. A. von Jor-
dans.

Obviously described from a series of nine specimens list-
ed in the inventory book of the ZFMK, but only ZFMK
G.X.1.b°.¢., male, defined as “cotype”. One paratype ex-
changed with the MfN, Berlin. — Dickinson (2003): Parus
caeruleus balearicus.

Parus corsus Kleinschmidt, 1903
Orn Monatsber 11: 6.

Holotype: Coll. Kl. 4129, male, Ajaccio, Corsica,
18.01.1902.

Probable paralectotypes: Coll. KI. 4130-4137, 4 males, 4
females, Ajaccio, 18.01.—15.02.1902.

In original description no details on number of skins stud-
ied are given, no paratypes are defined. In his Catalogue
Kleinschmidt (1935-1943: 97) indicates nine specimens
from Corsica as “including type”, and lists additional six-
teen specimens from Sardinia. — Dickinson (2003):
Parus major corsus.

Parus wladiwostokensis Kleinschmidt, 1913
Falco 9: 33.

Holotype: Coll. Kl. 4162, male, Wladiwostok, NE Asia,
07.12.1900.

Paratype: Coll. KI. 4163, male, Wladiwostok, 18.09. with-
out year.

Only two specimens in Collection Kleinschmidt I. — Dick-

inson (2003) presumably: Parus major minor Temminck
& Schlegel, 1848.

Parus tschiliensis Kleinschmidt, 1922
Falco 18: 3.

Paralectotype: Coll. KI. 4164, male, 30 km n. Balihandi-
en, NE Jehol, Tschili, leg. Weigold, per Stotzner.

©ZFMK

Types in ZFMK: Section Ornithology 51

No type specimen designated in original description. Lec-
totype and other paralectotypes at the SNS, Dresden, lec-
totype specimen lost in 1945 (see Eck & Quaisser 2004).
— Dickinson (2003) presumably: Parus major minor Tem-
minck & Schlegel, 1848.

Parus major gotoensis Kleinschmidt, 1922
Falco 18: 2.

Lectotype: Coll. Kl. 4176, male, Goto Islands, Japan,
18.04.1906.

Paralectotype: Coll. Kl. 4177, male, Goto Islands,
01.04.1906.

Only two specimens existed for the description, their iden-
tity being uncertain as data were neither defined in the
original description nor a type designated in the Catalogue
(Kleinschmidt 1935-1943: 97). Lectotype and paralecto-
type designated by Rheinwald & van den Elzen (1984);
identification of type specimen taken from handwriting on
label, (Cotype = paralectotype; see also Dickinson et al.
2006: 88). — Dickinson (2003): Parus major kagoshimae
Taka-Tsukasa, 1919.

Parus major amamiensis Kleinschmidt, 1922
Falco 18: 2.

Lectotype: Coll. KI. 4178, female, Amami Oshima, Japan,
25.02.1906.

Described on the basis of one single specimen. Informa-
tion on type status derived from label (Rheinwald & van
den Elzen 1984). — Dickinson (2003): Parus major
amamiensis.

Parus major subtibetanus Kleinschmidt & Weigold, 1922
Abh Ber Mus Tierk Dresden 15: 11.

Paratype: Coll. KI. 4159, male, a three days’ journey from
Batang, Janeti, E Tibet, 29.09.1915, leg. Weigold.

Described from a larger series. Holotype and nine
paratypes listed in Eck & Quaisser (2004) for SNS, Dres-
den. Holotype and one paratype specimen lost in 1945.
Whereabouts of other specimens uncertain. — Dickinson
(2003) presumably: Parus major commixtus Swinhoe,
1868.

Bonn zoological Bulletin 59: 29-77

Parus maior mallorcae Jordans, 1913
Falco 9: 43.

Lectotype: ZFMK G.X.1.a3.a., male, Palma, Valldemosa,
Majorca, Baleares, 10.03.1913, leg. A. von Jordans No
707.

Paralectotypes: ZFMK G.X.1.a?.f., 0, Lluch, Majorca,
22.03.1913; ZFMK G.X.1.a3.y., male, Lluch, Majorca,
18.03.1913; ZFMK G.X.1.a3.6., 0, Lluch, Majorca,
22.03.1913; ZFMK G.X.1.a3.¢., female, Esporlas,
20.05.1913; ZFMK G.X.1.a3.¢., male, Fornalutx,
17.03.1913; ZFMK G.X.1.a3.n., male, Valldemosa,
19.05.1913; ZFMK G.X.1.a3.6., female, Esporlas,
12:03:19132 ZPEMK GX. 1-a3.1., male, Esporlas,
20.05.1913; ZFMK G.X.1.a3.«., female, Esporlas,
20.05.1913; ZFMK G.X.1.a3.2., female, Santany,
09.05.1913; ZFMK G.X.1.a3.u., male, Palma, Valldemosa,
10.03.1913; ZFMK G.X.1.a3.v., male, Fornalutx, Major-
ca, 17.03.1913, all leg. A. von Jordans.

Probably described from a series of thirteen specimens;
no type specimen designated in original description (“type
is in my collection’). Lectotypes designated by Rheinwald
& van den Elzen (1984) from handwritten label and col-
lection inventory. — Dickinson (2003): Parus major mal-
lorcae.

Parus major ecki Jordans, 1970
Zool Abh Mus Tierk Dresden 31: 213.

Holotype: ZFMK G.X.1.a?.p., male, Mt. Tortoli, Sardinia,
28.04.1926.

Paratypes: Coll. Kl. 4153, female, Lanusei, Ogliastra,
14.11.1902, and ZFMK G.X.1.a’.f., female, Talana,
Ogliastra, Sardinia, 31.01.1903; ZFMK G.X.1.a?.y., ma-
le, Bari, Ogliastra, Sardinia, 23.03.1903; ZFMK
G.X.1.a’.6., male, Loceri, Ogliastra, 06.02.1904; ZFMK
G.X.1.a*.¢., female, Loceri, Ogliastra, 06.02.1904; ZFMK
G.X.1.a?.¢., male, Arzana, Ogliastra, 24.02.1904; ZFMK
G.X.1.a?.n., male, Ilbono, Ogliastra, 22.03.1904; ZFMK
G.X.1.a?.0., female, Baunusei, Ogliastra, 03.03.1906;
ZFMK G.X.1.a?.t., female, Lanusei, Ogliastra, 06.05.1906,
all per O. Kleinschmidt; ZFMK G.X.1.a?.A., male, Oghia-
stra, Sardidien, 13.002.1904; ZFMK G.X.1.a?.u., male,
Ogliastra, Sardidien, 24.04.1906; ZFMK G.X.1.a?.v., ma-
le, Lanusei, 15.04.1924; ZFMK G.X.1.a?.€., male, Lanu-
sei, 18.08.1924; ZFMK G.X.1.a.o., male, Lanusei,
20.02.1925; male, Lanusei, ZFMK G.X.1.a?.2., Mt. Tor-
toli, Sardinia, 25.04.1926; all leg. A. v. Jordans.

Described from a series of twenty-five males and nine fe-
males; whereabouts of remaining paratypes unknown. —
Dickinson (2003): Parus major ecki.

©ZFMK

N
NO

Parus maior caspius Zarudny & Loudon, 1905
Orn Monatsber 13: 109.

Probable paralectotype ZFMK G.X.1.a5.a., male, Kuljam-
Chadschi-Ali, W Iran, 13.10. (= 26.10. Gregorian Calen-
dar) 1903, ex. Coll. Zarudny No 1645, Coll. Jordans No
1454.

No type defined in the original description; the individ-
ual is labelled as “cotype” by von Jordans and published
as “cotype” (Jordans 1923: 23). — Dickinson (2003): Parus
major karelini Zarudny, 1911. See next taxon.

Parus major karelini Zarudny, 1911
J Ornithol 59: 236.

Nomen novum for Parus maior caspius Zarudny &
Loudon, 1905, because of Parus caspicus S.G. Gmelin,
1774 (which is Motacilla cinerea). Presumed paralecto-
type is ZFMK G.X.1.a5.a., see previous taxon.

Parus maior zayrossiensis Zarudny & Loudon, 1905
Orn Monatsber 13: 108.

Paralectotype ZFMK G.X.1.a°.B., male, Samdalkal, W
Iran, 02.01.1904 (=20.12.1903 Gregorian Calendar),
Coll. Zarudny 1476a, Coll. A. von Jordans No 1455.

No holotype defined in the original description, the spec-
imen is labelled “cotype” by A. von Jordans, but not men-
tioned as in his collection (Jordans 1923); t. t. Zagros
Mountains, (name is misprint of Pm. zagrossiensis). —
Dickinson (2003): Parus major blanfordi Prazak, 1894.

Parus major niethammeri Jordans, 1970
Zool Abh Mus Tierk Dresden 31: 218.

Lectotypes: ZFMK 57.858 and 57.859, 2 males, Canea,
Crete, 12.02.1925, leg. Schiebel.

Paratypes: ZFMK 57.857, female, Canea, Crete,
17.02.1925; ZFMK 57.860, male, Canea, Crete,
14.02.1925; ZFMK 57.863, male, Canea, Crete,
26.03.1925; ZFMK 57.864, male, Canea, Crete,
20.03.1925; ZFMK 57. 865, male, Canea, Crete,
03.03.1925; ZFMK 57.866, male, Canea, Crete,
21.02.1925; ZFMK 57.867, female, Canea, Crete,
24.03.1925; ZFMK 57.868, female, Canea, Crete,
24.04.1925; ZFMK 57.869, female, Souda Bay,

14.02.1925; ZFMK 57. 870, male, Souda Bay, 24.03.1925;
ZFMK 57. 871, female, Souda Bay, 15.03.1915; ZFMK
57. 872, female, Souda Bay, 15.03.1925; ZFMK 57. 873,

Bonn zoological Bulletin 59: 29-77

Renate van den Elzen

male, Souda Bay, 10.05.1925; ZFMK 57. 874, female,
Souda Bay, 14.02.1925; ZFMK 57.875, male, Souda Bay,
26.04.1925; ZFMK 57.876, female, Souda Bay,
26.04.1925 and ZFMK 57.877, male, Wrysses, Crete,
03.04.1925, all leg. Schiebel.

One single specimen defined as type in the original de-
scription. However, in the collection of the ZFMK two
males collected at the same day and locality bear the hand-
written notice “type” by von Jordans. Thus, both speci-
mens are listed as lectotypes. Originally described from
eighteen males and thirteen females; ZFMK 57. 861-862
exchanged with AMNH, New York, whereabouts of oth-
er specimens unknown. — Dickinson (2003): Parus ma-
Jor niethammeri.

HIRUNDINIDAE

Cotyle ruspestris spatzi) Geyr, 1916
Orn Monatsber 24: 59.

Holotype: ZFMK B.HI.4.b!.y., male, Gara Djenrun, Al-
geria, 11.03.1914, leg. Geyr.

Paratypes: ZFMK_ B.III.4.b!.a., male, Oued Aghelil,
18.03.1914; ZFMK B.I.4.b!.B., Oued Amra, 05.04.1914;
ZFMK B.III.4.b!.6., female, Oued Amra, 05.04.1914;
ZFMK B.III.4.b!.¢., male, Amgid, 16.02.1914; ZFMK
B.III.4.b!.¢., male, Gara Dyjenrun,11.03.1914; ZFMK
B.I1.4.b!.n., male, Amgid, 16.02.1914; ZFMK
B.II1.4.b!.0., male, Gara Djenrun, 11.03.1914, all leg.
Geyr.

Described from eight specimens, series complete. — Dick-
inson (2003): Ptyonoprogne obsoleta spatzi.

!) misprint in original description, correctly: Cotyle ru-
pestris spatzi Geyr, 1916.

Hirundo Urbica vogti Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 61.

Lectotype: Coll. KI. 1995, male, Niedernhausen/ Oden-
wald, 02.06.1916, leg. Vogt.

Paralectotypes: Coll. KI. 1996-2001, 2 males, 2 females,
2 juv., Niedernhausen/ Odenwald, 25.05.—19.07.1916, leg.
Vogt.

Very brief description, no types designated. Lectotype and
paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Delichon urbicum urbicum
(Linnaeus, 1758).

OZFMK

Types in ZFMK: Section Ornithology 53

AEGITHALIDAE

Aegithalos caudatus expugnatus Bacmeister & Klein-
schmidt, 1916
Falco 12: 18.

Lectotype: Coll. Kl. 4870, male, Dept. Ardennes, E
France, 09.02.1916, leg. Bacmeister.

Paralectotypes: Supposedly Coll. Kl. 4871-4882, 7
males, 3 females, 2 0, La Tourauderie, Thugny, Dept. Ar-
dennes, 30.01.—08.04.1916, leg. Bacmeister.

No types defined in the original description; series indi-
cated as “types” in Catalogue (Kleinschmidt 1935-1943:
106), type specimen labelled as type. — Dickinson
(2003): Aegithalos caudatus europaeus (Hermann, 1804).

Acredula tephronota passekii Zarudny, 1904
Orn Monatsber 12: 164.

Paralectotype ZFMK G.X.2.a!9.a., male, Schalil, W Iran,
04.01.1904 (= 22.12.1903 according to the Gregorian Cal-
endar), leg. Zarudny No 1480, ex Coll. Jordans No 1586.
No type designated in original description; obviously no
large series existed. The specimen is published by Jordans
(1923: 25) as “I have a cotype from Schalil in Iran in my
collection” and labelled as “cotype” by A. von Jordans in
the ZFMK bird skin collection. — Dickinson (2003): Ae-
githalos caudatus passekii.

Aegithalos caudatus chlamyrhodomelanos Clancey,
1941
Ibis 14: 314.

Paratype: ZFMK 50.116, female, Suthershire, Scotland,
06.09.1938, leg. Clancey.

Holotype at the RSME, Edinburgh, t .t. Dornoch, Suther-
land. Described from a series of four specimens. — Dick-
inson (2003): Aegithalos caudatus rosaceus Mathews,
1937.

ALAUDIDAE
Eremophila alpestris aharonii Neumann, 1934

Anz Orn Ges Bayern 2: 333.

Paratype: ZFMK 37.142, female juv. ?, Ras Balbeck, Is-
rael, 18.04.1931, leg. Aharoni.

Bonn zoological Bulletin 59: 29-77

Type specimen at the MCZ, Harvard; others probably al-
so at Tring, UK and also at the MfN, Berlin? Described
from a series of thirteen specimens. — Dickinson (2003)
presumably: Eremophila alpestris bicornis (C.L. Brehm,
1831).

Galerida cristata jordansi Niethammer, 1955
Bonn zool Beitr 6: 57.

Holotype: ZFMK 54.517, male, Abbangarit, NW Agades,
Niger, 29.01.1954, leg. G. Niethammer No 480.

Paratype: ZFMK 54.1165, male, Abbangarit, 29.01.1954,
leg. G. Niethammer.

A third male is probably lost. — Dickinson (2003): Galeri-
da cristata jordansi.

Galerida cristata carthaginis Kleinschmidt & Hilgert,
1905
Orn Monatsber 13: 188.

Nomen novum for Galerida cristata macrorhyncha Er-
langer, 1899, nec Tristram, 1859.

Paratype: Coll. Kl. 1488-1491, 4 0, vicinity of Tunis,
Tunisia, 20. and 21.01.1899, leg. Blanc, original labels
lost.

Type specimen (Coll. Erlanger No 2937) at the SMF,
Frankfurt/Main?, t. t. like paratypes. — Dickinson (2003):
Galerida cristata carthaginis.

Galerida cristata gafsae Kleinschmidt & Hilgert, 1905
Orn Monatsber 13: 189.

Nomen novum for Galerida cristata arenicola Erlanger,
1899, nec Tristram, 1859. Paratypes: Coll.K1. 1492-1493,
2 males, Bir Sellousa, Tunesia, 07.03.1899, leg. Spatz.

Type specimen in Coll. Erlanger No 2950, leg. Erlanger,
t. t. Seggi, Tunesia, currently SMF, Frankfurt/Main. — Dic-
kinson (2003): ? Galerida cristata carthaginis.

Calerida cristata vamberyi Harms, 1907
Orn Monatsber 15: 49.
Holotype: ZFMK H.V.1.a!®y., male ad., Utsch-Adschi,

Trans-Caspia, 26.02.1900 (14.02.1900 according to the
Gregorian Calendar), leg. Harms.

©ZFMK

54 Renate van den Elzen

Paralectotypes: ZFMK_ H.V.1.a!®.B., male, Tedschen,
Transcaspia, 13.02.1900; ZFMK_ H.V.1.a!¢.6., male,
Utsch-Adschi, 27.02.1900; ZFMK H.V.1.a!°«., female,
Tedschen, 31.03.1900, all leg. Harms.

Only type specified in original description, paralectotype
designation in Rheinwald & van den Elzen (1984) The
type specimen is mentioned in Jordans (1923: 10). — Dick-
inson (2003): Galerida cristata magna Hume, 1871.

Galerida cristata apuliae Jordans, 1935
Orn Monatsber 43: 119.

Holotype: ZFMK H.V.1.a?3.a., male, San Severo, Apulia,
Italy, 20.02.1932.

Paratypes: ZFMK H.V.1.a23.B8., male, Cérignola, Apulia,
11.1931; ZFMK H.V.1.a23.y., female, San Severo, Apu-
lia, 20.01.1932.

Description based upon these three specimens. — Dickin-
son (2003): Galerida cristata apuliae.

Galerida schliteri Kleinschmidt, 1904
Orn Monatsber 12: 196.

Lectotype: Coll. KI. 1521, female ?, Chefia, vicinity of Bo-
ne, N Algeria, 10.1902, leg. Schliiter.

Paralectotypes: Coll. Kl. 1522-1523, 2 male, Béne,
10.1902 and 11.1904, leg. Schliter, Coll. Kl. 1524-1526,
3 male, Kerrata, Algeria, 28.04. and 21.05.1904, leg.
Flikiger.

No type specimen defined in the original description; lec-
totype designated in Catalogue (Kleinschmidt 1935-1943:
40). — Dickinson (2003): Galerida theklae ruficolor
Whitaker, 1898.

|Galerida desertorum Ch. L. Brehm, 1848
Okens Isis 7: 497.

Syntypes: ZFMK 64.251—64.252, male, female, (pair),
Hasenhaide/ Berlin, Germany, 28.03.1833, leg. Fehrmann.

See also Homeyer 1848: Orn Briefe: 47; letter from Ch.
L. Brehm to Homeyer dated 05.01.1835. Types today at
the “Brehm Gedenkstatte” in Renthendorf. — Dickinson
(2003): Lullulla arborea arborea (Linnaeus, 1758).]

Bonn zoological Bulletin 59: 29-77

Melanocorypha calandra hollomi Kumerloeve, 1969
Rev Fac Sci Univ Istanbul 34: 261.

Lectotype: ZFMK 68.1774, male, Yiiksekova, Turkey,
10.06.1968, leg. Kumerloeve.

Paralectotypes: ZFMK 68.1775, male, Yuksekova,
10.06.1968; ZFMK 68.1776, male, Yutksekova,
10.06.1968; ZFMK 68.1777, male, Yuksekova,
12.06.1968; ZFMK 68.1778, male, Yiiksekova,
12.06.1968; ZFMK 68. 1779, male, Yuksekova,

12.06.1968; ZFMK 68.1780, male, Van, 18.06.1968;
ZFEMK 68. 1781, male, Van, 22.06.1968; ZFMK 68.1782,
female, Van, 22.06.1968, leg. Kumerloeve.

No types designated in original description; lectotypes
specified in Rheinwald & van den Elzen (1984). — Dick-
inson (2003): Melanocorypha calandra calandra (Lin-
naeus, 1766).

Melanocorypha calandra dathei Kumerloeve, 1970
Beitr Vogelkunde 16: 246.

Holotype: ZFMK 68.1689, male, Ceylanpinar, Turkey,
10.05.1968, leg. Mittendorf No 54.

Paratypes: ZFMK 68.1690, male, Ceylanpinar, Turkey,
10.05.1968; ZFMK 68. 1691, male, Ceylanpinar, Turkey,
09.05.1968; ZFMK 68. 1692, male, Ceylanpinar, Turkey,
07.05.1968; ZFMK 68.1693, female, Ceylanpinar,
14.05.1968; ZFMK 60.970, male, Muratpasa, Syria,
22.05.1953; ZFMK 64.3056, male, Birecik, Turkey,
22.05.1964; ZFMK 64.3057, male, Birecik, Turkey,
22.05.1964; ZFMK 64.3064, male, Djabou, Syria,
08.06.1964; ZFMK 64.3065, male, Djabou, Syria,
08.06.1964, all leg. Mittendorf. — Dickinson (2003): Me-
lanocorypha calandra gaza Meinertzhagen, 1919

Calandrella brachydactyla woltersi Kumerloeve, 1969
J Ornithol 110: 324.

Holotype: ZFMK 68.1752, male, Amik G6li, Turkey,
03.05.1968, leg. Mittendorf.

Paratypes: ZFMK 68.1750, male, Bedirge, 02.05.1968;
ZFMK 68.1751, male, Amik Goli, 03.05.1968; ZFMK 68.
1753, female, Amik Golii, 03.05.1968; ZFMK 68.1754,
female, Amik Golii, 03.05.1968, all leg. Mittendorf. — Dic-
kinson (2003): Calandrella brachydactyla woltersi.

©OZFMK

Types in ZFMK: Section Ornithology 55

Calandrella rufescens niethammeri Kumerloeve, 1963
Vogelwelt 84: 147.

Holotype: ZFMK 62.950, male, Eber Gdlii, Turkey,
09.06.1962, leg. Mittendorf.

Paratypes: ZFMK 62.951, male, Eber Golii, 09.06.1962;
ZFMK 62.952 male, Eber Golti, 09.06.1962; ZFMK 62.
953, male, Eber Golii, 09.06.1962; ZFMK 62.954, male,
Eber Golu, 09.06.1962; ZFMK 62.955, male, Eber Goli,
09.06.1962, all leg. Mittendorf. — Dickinson (2003): Ca-
landrella cheleensis niethammeri.

Alaemon Margaritae Koenig, 1888
J Ornithol 36: 228.

Syntypes: ZFMK H.V.9.a!.y., male, near Gabes, Tunesia,
18.03.1887; ZFMK H.V.9.a!. ¢., female, near Gabes, Tune-
sia, 18.03.1887, leg. Koenig. — Dickinson (2003): Cher-
sophilus duponti margaritae (Koenig, 1888).

Ammomanes deserti kollmannspergeri Niethammer,
1955
Bonn zool Beitr 6: 59.

Holotype: ZFMK 54.1202, male, 20 km SE of Fada, Chad,
03.04.1954, leg. Kollmannsperger.

Paratypes: ZFMK 54.1201 male, E Fada, 04.1954;
ZFMK 54.1203, female, E Fada, 04.1954; ZFMK 80.376,
female, E Fada, 04.1954.

Described from four males and two females; whereabouts
of other specimens unknown. — Dickinson (2003): Ammo-
manes deserti kollmannspergeri.

Ammomanes phoenicura pallens \e Roi, 1912
Orn Monatsber 20: 6.

Syntypes: ZFMK H.V.6.b?.6., male, Bir Sani, Bajuda
steppe, “Egyptian Sudan’, 03.03.1903; ZFMK H.V.6.b”.7.,
female, Bir Sani, Bajuda steppe, Egyptian Sudan,
27.02.1903, leg. Koenig.

Paratypes: ZFMK_ H.V.6.b*.a., male, Bajuda steppe,
01.03.1903; ZFMK H.V.6.b’.B. female, Bajuda steppe,
26.02.1903; ZFMK H.V.6.b?. 6., female, Bir Sani, Baju-
da steppe, 27.02.1903; ZFMK H.V.6.b’.c., female, Bir
Sani, Bajuda steppe, 27.02.1903; ZFMK H.V.6.b*.¢., male,
Bajuda steppe, 02.03.1903; ZFMK H.V.6.b’.n., female,
Bajuda steppe, 02.03.1903; ZFMK H.V.6.b?.«., Chor Abu
Harrasch, 04.03.1903, all leg. A. Koenig.

Bonn zoological Bulletin 59: 29-77

One paratype in Tring, UK. — Dickinson (2003): Ammo-
manes cinctura arenicolor Sundevall, 1851.

Ammomanes grayi hoeschi Niethammer, 1955
Bonn zool Beitr 6: 185.

Syntypes/paralectotypes: ZFMK 66.895, male, female,
Namib, W Orupembe, Kaokoveld, Namibia, 27.06.1952
(=UMB 19887); ZFMK 66.896, female, Namib, W Oru-
pembe, Kaokoveld, 27.06.1952(=UMB 19888), all leg.
Hoesch.

Described from four specimens. In the original descrip-
tion male No 19889 of the UMB is defined as type spec-
imen, the ZFMK specimens are labelled as syntypes. List-
ed as syntypes in Rheinwald & van den Elzen (1984). The
second female mentioned in the original description is not
at the ZFMK. — Dickinson (2003): Ammomanes grayi
hoeschi.

Mirafra sabota uis Hoesch & Niethammer, 1940
J Ornithol 88, Sonderheft: 219

Holotype: ZFMK 39.983, female, Uis, pre-Namib, Namib-
ia, 22.09.1938, leg. Hoesch and Niethammer No 117.

Paratype: ZFMK 39.921, female, 70 km E of Uis,
27.09.1938, leg. Hoesch and Niethammer.

Two further specimens mentioned in the original descrip-
tion are not at the ZFMK. — Dickinson (2003): Mirafra
sabota naevia (Strickland, 1853).

Mirafra africanoides isseli Hoesch & Niethammer, 1940
J Ornithol 88, Sonderheft: 215.

Syntypes: ZFMK 39.897, male, south of Okahandja,
Damaraland, Namibia, 04.09.1938; ZFMK 39.898, fe-
male, south of Okahandja, Damaraland, Namibia,
04.09.1938, leg. Niethammer No 42 and without No.,
prep. Issel.

Subsequently defined as Mirafra africanoides omaruru

(Roberts, 1936) by Niethammer. — Dickinson (2003): Mi-
rafra africanoides harei Roberts, 1917.

©OZFMK

56 Renate van den Elzen

PYCNONOTIDAE

Phyllastrephus leucolepis Gatter, 1985
J Ornithol 126: 155.

Holotype: ZFMK 84.221, 0, 20 km NW Zwedru near Cay-
alla River, Grand Gedeh County, Liberia
(06°12’N,08°11°>W), 16.01.1984, leg Gatter.

Described from that specimen only. — Dickinson (2003):
Phyllastrephus leucolepis.

SYLVIIDAE

Regulus satelles Koenig, 1889
J Ornithol 37: 263.

Syntype: ZFMK G.X.6.a3.6., male, Pinar, Tenerife,
11.04.1889, leg. Koenig.

Synonym of Regulus teneriffae Seebohm, 1883, the syn-
type is obviously the only remaining from a series of four
specimens. — Dickinson (2003): Regulus regulus tenerif-
fae Seebohm, 1883.

Regulus regulus ellenthalerae Packert, Dietzen, Martens,
Wink & Kvist, 2006
J Avian Biol 37: 378.

Holotype: ZFMK G.X.6.a3.o., male, Hoya Grande, La Pal-
ma, 01.1905, leg. Thanner.

Paralectotypes: ZFMK G.X.6.a3.yy., female, El Paso, La
Palma, 01.1905; ZFMK G.X.6.a3.««., male, Pinar, Hier-
ro, 02.1905, all leg. Thanner.

Described from a larger series of another six males and
three females from La Palma, and two males, two females
from Hierro, all ZFMK, Bonn.

Regulus ignicapillus balearicus Jordans, 1923
Falco 1923:

Holotype: ZFMK G.X.6.b2.a., female, Lluch, Majorca,
24.03.1913, leg. A. von Jordans, No 1618.

Paratypes: ZFMK G.X.6.b2.6., female, Lluch, Majorca,
24.03.1913; ZFMK G.X.6.b2.8., female, Lluch,
24.03.1913; ZFMK G.X.6.b2.c., male, Fornalutx,
17.03.1913; ZFMK G.X.6.b2.0., male, Arta, 23.04.1913;

Bonn zoological Bulletin 59: 29-77

ZEMK G.X.6.b2.1., male, Palma-Valdemosa, 10.03.1913;
ZFMK G.X.6.b2.«., male, Lluch, 24.03.1913; all leg. A.
v. Jordans.

Described from fifteen individuals; ten registered in the
inventory of the ZFMK skin collection; of these three ex-
changed with NHM Louisiana, MfN, Berlin. Whereabouts
of the five missing specimens uncertain. — Dickinson
(2003): Regulus ignicapillus balearicus.

Cettia cetti salvatoris Jordans, 1914
Falco 10: 41.

Lectotype: ZFMK H.I.7.a!.a., male, Albufereta, Majorca,
27.03.1913, leg. A. von Jordans, No 742.

Paralectotypes: ZFMK H.I.7.a!.B., male, Albufereta, Ma-
jorca, 27.03.1913; ZFMK H.1.7. a!.0., male, Albufereta,
Majorca, 27.03.1913, all leg A. v. Jordans.

Originally described from a series of four specimens; lec-
totype designated by Rheinwald & van den Elzen (1984)
from information of ZFMK inventory; fourth specimen not
registered there, whereabouts unknown. — Dickinson
(2003) presumably: Cettia cetti cetti (Temminck, 1820).

Acrocephalus dumetorum gabrielae Neumann, 1934
Verh Orn Ges Bayern 20: 470.

Paralectotypes: ZFMK H.I.2.e3.6., male, Solak, Asia Mi-
nor, 27.05.1934; ZFMK H.I.2.e3.¢ female, Elmali, Asia
Minor, 11.06.1934, leg. G. Neuhauser.

Described from three males and one female. Synonym of
Hippolais pallida elaeica Lindermayer, 1843, see Orn.
Monatsber. 42: 178.

Hippolais pallida laeneni Niethammer, 1955
Bonn zool Beitr 6: 66.

Holotype: ZFMK 54.417, male ad., Bol,
15.04.1954, leg. Laenen.

Chad,

Paratypes: ZFMK 54.415, male, Bol, Chad, 31.03.1954;
ZFMK 54.429, , Bol, Chad, 16.04.1954, all leg. Laenen.

Described from three males, two females and one juve-

nile; whereabouts of other specimens unknown. — Dick-
inson (2003): Iduna pallida laeneni (Niethammer, 1955).

©ZFMK

Types in ZFMK: Section Ornithology 57

Hypolais icterina Borisi Jordans, 1940
Mitt K6nigl Naturwiss Inst Sofia 13: 103.

Holotype: ZFMK 38.796, male ad., Kamtschyja, E Bul-
garia, 07.06.1935, leg. Heinrich, Inv. No 307 (Museum
Warsaw).

Described from a series of twelve specimens; no paratypes
designated in original description. — Dickinson (2003):
monotypic: Hippolais icterina (Vieillot, 1817).

Melocichla mentalis incana Diesselhorst, 1959
Opusc Zool 36: 1.

Holotype: ZFMK 58.324, female, Momella-Meru Moun-
tains, Tansania, 17.01.1958, leg. Nagy.

Described from four females, one o. Further material
(paratypes) presumably at the ZSM, Munic. — Dickinson
(2003): Melocichla mentalis incana.

Cisticola iuncidis intermedia Jordans, 1923
Falco 19, Sonderheft: 3.

Holotype: ZFMK H.I.9.a!.o0., male, Albufera, Majorca,
11.05.1921, leg. A. von Jordans No 3050.

Presumed paratypes: ZFMK H.1.9.a!.«., male, Albufere-
ta, Majorca, 27.03.1913; ZFMK H.I.9.a!.2., male, Albu-
fereta, Majorca, 29.03.1913; ZFMK H.1.9.a!.u., male, Al-
bufereta, Majorca, 09.04.1913; ZFMK H.I.9.a!.v., male,
Estany de Gambas, 02.05.1913; ZFMK H.I.9.a!.€., fema-
le, Albufera, 31.03.1913; ZFMK H.1.9.a!.2., male, Albu-
fera, 10.06.1921; ZFMK H.I.9.a!.p., female, Alcudia,
07.05.1921; ZFMK H.1.9.a!.o., male, Alcudia, 04.05.1921;
ZFMK H.1.9.a!.t., female, Alcudia, 04.05.1921; ZFMK
H.1.9.a!.o., male, Alcudia, 06.05.1921; ZFMK H.1.9.a!.y.,
female, Albufera, 11.05.1921; ZFMK H.1.9.a!.y., fema-
le, Albufera, 16.05.1921, all leg. A. von Jordans.

Described from seventeen specimens, no paratypes des-
ignated or described in detail. In inventories of the ZFMK
bird collection only fifteen specimens registered, one spec-
imen exchanged with MfN, Berlin. — Dickinson (2003):
Cisticola juncidis cisticola (Temminck, 1820).

Eremomela icteropygialis laeneni Niethammer, 1955
Bonn zool Beitr 6: 65.

Holotype: ZFMK 54.390, male, Bol, Chad, 03.04.1954,
leg. Laenen.

Bonn zoological Bulletin 59: 29-77

Paratype: ZFMK 54.410, female, Bol, 15.04.1954, leg.
Laenen.

Described from three specimens; the second male listed
in the original description is not at the ZFMK. — Dickin-
son (2003): ?Eremomela icteropygialis alexanderi Sclater
& Mackworth-Praed, 1918.

Sylvia sarda balearica Jordans, 1913
Falco 9: 43.

Holotype: ZFMK H.1.212.b!.a., male, Isla Dragonera of
the W coast of Mallorca, 19.04.1913, leg. A. von Jordans
No 629.

Paratypes: ZFMK H.1.212.b!.B., male juv., E of Sta Pon-
za, Majorca, 15.05.1913; ZFMK_ H.I.212.b!.y., female,
Santany, 01.05.1913; ZFMK H.1.212.b!.6., male, San Tel-
mo, Andraitx, 15.04.1913; ZFMK H.1.212.b!.¢., male, Al-
cudia, 02.04.1913; ZFMK H.1.212.b!.C., female, Alcudia,
28.03.1913; ZFMK H.I.212.b!.n., male, Dragonera,
16.04.1913; ZFMK H.I.212.b!.1., male, Alcudia,
02.04.1913; ZFMK_ H.I.212.b!.«., male juv., Cabrera,
12.05.1913; ZFMK H.1.212.b!.A., male, Isla Dragonera,
16.04.1913; ZFMK H.1.21?.b!.u., male, Alcudia,
28.03.1913, all leg. A. von Jordans.

Described from a series of nineteen specimens (ten males,
seven females, two juveniles), twelve specimens registered
in the ZFMK inventory, one paratype exchanged with the
AMNH, New York. — Dickinson (2003): Sylvia sarda
balearica.

Sylvia communis traudeli Kumerloeve, 1969
Rev Fac Sci Univ Istanbul 34: 274.

Holotype: ZFMK 68.1906, male, Van, Turkey, 31.05.1968,
leg. Mittendorf No 152.

Paratypes: ZFMK 68.1904, male, Ceylanpinar,
09.05.1968; ZFMK 68.1905, male, Ceylanpinar,
17.05.1968; ZFMK 68.1907, male, Hakkari-Beytiissebap,
05.06.1968; ZFMK 68.1908, male, W of Yiiksekova,
12.06.1968; ZFMK 68.1909, male, W of Ytksekova,
12.06.1968; ZFMK 68.1910, male, Tatvan, 06.07.1968, all
leg. Mittendorf.

Described from ten specimens, the specimens from 1965

could not be identified with certainty. — Dickinson (2003):
Sylvia curruca curruca (Linnaeus, 1758).

©ZFMK

58 Renate van den Elzen

Sylvia melanocephala pasiphaé Stresemann & Schiebel,
1925
J Ornithol 58: 659.

Paratype: ZFMK 57.998, male, Suda near Canea, Crete,
26.04.1925, leg. Schiebel.

Holotype at the ZMB, Berlin; no other specimens speci-
fied. — Dickinson (2003): Sylvia melanocephala pasiphaé.

Sylvia semenowi Zarudny, 1904
Orn Jahrb 15: 220-221.

Paralectotype: ZFMK H.1.21.1-y., male, Karun Tschinebe,
W Iran, 25.01.1904 (= 12.01.1904 Gregorian Calendar),
ex. Coll. Zarudny No 1630, Coll. A. von Jordans No 2542.

Specimen belongs to the type series and 1s published as
“cotype” by Jordans (1923: 26); described from seven
specimens. Whereabouts of other specimens unknown. —
Dickinson (2003): Sylvia mystacea rubescens Blanford,
1817.

Sylvia borin Pateffi Jordans, 1940
Mitt Konig] Naturwiss Inst Sofia 13: 105.

Holotype: ZFMK 38.633, male ad., Bansko, Bulgaria,
05.06.1938, leg. A. von Jordans.

Paratype: ZFMK 38.634, male, Pirin Mountains near Ban-
sko, 13.06.1938, leg. A. von Jordans.

Described from these two specimens. — Dickinson (2003):
Sylvia borin woodwardi (Sharpe, 1877).
Sylvia clarae Kleinschmidt, 1901

Orn Monatsber 9: 167.

Holotype: Coll. Kl.
06.05.1899, leg. Spatz.

3608, male, Gafsa, Tunisia,

Paratype: Coll. Kl. 3609, female, Gafsa, 07.05.1899, leg.
Spatz. — Dickinson (2003): Sylvia borin borin (Boddaert,
1783).

Sylvia atricapilla koenigi Jordans, 1923
Falco 19, Sonderheft: 3.

Holotype: ZFMK H.1.21.d!.a., male, Arta, Majorca,
26.04.1913, leg. A. von Jordans No 977.

Bonn zoological Bulletin 59: 29-77

Paratypes: ZFMK H.1.21.d!.y., male, Esporlas,
20.05.1913; ZFMK H.I.21.d!.6., male, Arta, 23.04.1913;
ZFMK H.1.21.d!.¢., male, Valldemosa, 15.03.1913; ZFMK
H.1.21.d!.C., female, Valldemosa, 12.04.1913; ZFMK
H.1.21.d!.y., female, Esporlas, 12.03.1913; ZFMK
H.1.21.d!.0., male, Valldemosa, 12.04.1913; ZFMK
H.1.21.d!.1., male, Palma-Valldemosa, 10.03.1913; ZFMK
H.1.21.d!.*«., male, Albufera, 18.05.1921; ZFMK
H.1.21.d!.4., male, Valldemosa, 03.05.1921; ZFMK
H.1.21.d!.u., male, Alcudia, 20.06.1921; ZFMK
H.1.21.d!.v., male, Alcudia, 20.06.1921; ZFMK
H.1.21.d!.€., male, Alcudia, Majorca, 20.06.1921, all leg.
A. v. Jordans.

Described from twenty-two specimens, one exchanged
with the MfN, Berlin; the whereabouts of the remaining
skins are unknown. — Dickinson (2003): Sylvia atricapil-
la pauluccii Arrigoni, 1902.

TIMALIIDAE

Pnoepyga immaculata Martens & Eck, 1991
J Ornithol 132: 180.

Holotype: ZFMK 91.03, male, Thakkola, beneath Lete,
2530m, Mustang district, Nepal, 01.05.1980, leg. Martens.

Paratype: ZFMK 71.946, female, Thakkola, Chadziou
Khola, 2600 m, Mustang district, 30.06.1970, leg Martens.

Described from a series of two males and two females;
other paratypes at the ZSM, Munich and CNHM, Chica-
go; topotypical material at the ZFMK. — Dickinson (2003):
Pnoepyga immaculata.

Panurus biarmicus kosswigi Kumerloeve, 1958
Bonn zool Beitr 9: 197.

Holotype: ZFMK 58.460, male, Amik-Goéli, Turkey,
08.05.1953, leg. Kumerloeve.

Paratypes: ZFMK 58.461, male, Amik-G6li, Turkey,
06.05.1953; ZFMK 58.462, Amik-Golti, 06.05.1953;
ZFMK 58.463, female, Amik-G6lt, 08.05.1953, all leg.
Kumerloeve. — Dickinson (2003): Panurus biarmicus
kosswigi.

OZFMK

Types in ZFMK: Section Ornithology 59

Trochalopteron (Janthocichla) lineatum bilkevitchi
Zarduny, 1910
Orn Monatsber 18: 188.

Paralectotype: ZFMK H.1.208.a’.a., male, Chermadschan,
Buchara, Russland, 14.07.1910 (= 01.07. Gregorian Cal-
ender), leg. Zarudny, Coll. Harms No 1763, Coll. v. Jor-
dans No 1551.

Status derived from handwritten “Cotype” on label. —
Dickinson (2003): Garrulax lineatus bilkevitchi (Zardud-
ny, 1910).

ZOSTEROPIDAE

Zosterops poliogaster gerhardi van den Elzen & Konig,
1983
Bonn zool Beitr 34: 164.

Paratypes: ZFMK 78.195, female, Gilo, Imatong Moun-
tains, S Sudan, 06.01.1978; ZFMK 78.196, male, Gilo,
03.01.1978; ZFMK. 78.197, male, Gilo, 03.01.1978, all
leg. Nikolaus.

Holotype and two paratypes at the SMNS, Stuttgart (Holo-
type No 55.187, paratypes No. 55.185, 55.186). — Dick-
inson (2003) presumably: Zosterops poliogastrus kaffen-
sis Neumann, 1902.

TROGLODYTIDAE

Troglodytes troglodytes stresemanni Schiebel, 1926
Orn Monatsber 34: 14.

Holotype: ZFMK 57.1055, male, Elos, SW Crete,
24.04.1925, leg. Schiebel.

Described from seven specimens. A specimen labelled as
,cotype“ is housed at the MfN, Berlin (No 25.1444), topo-
typical material at the ZFMK (57.1054, 57.1056—57.1059,
Topolia, Homalo, 24.04.—05.05.1925). — Dickinson
(2003): Troglodytes troglodytes troglodytes (Linnaeus,
1758).

Troglodytes troglodytes weigoldi Jordans, 1923
Orn Monatsber 31: 14.

Paratype: ZFMK G.VII.1.a5.a., Favaios, Portugal,
20.03.1913, leg. Weigold.

Bonn zoological Bulletin 59: 29-77

Holotype and paratype at the MfN, Berlin, t .t. Oporto.

Described from three specimens. — Dickinson (2003) pre-
sumably: 7roglodytes troglodytes kabylorum E. Hartert,
1910.

Troglodytes troglodytes miilleri Jordans, 1928
Nov. Zool. 34: 283.

Holotype: ZFMK G.VII.1.a’. a., male, Valldemosa, Mal-
lorca, 12.04.1913, leg. A. von Jordans No 693.

Paratypes: ZFMK _ G.VII.1.a?.B., male, Alcudia,
28.03.1913; ZFMK G.VII.1.a?.y., female, Valldemosa,
14.03.1913; ZFMK G.VII.1.a7.6., male, Valldemosa,
12.04.1913; ZFMK G.VII.1.a’.e., male, Valldemosa,
2803 191325 ZEMIK NGaViliitas:G.) male; Alcudia;
28.03.1913; ZFMK G.VII.1.a*.n., 0, Alcudia, 28.03.1913;
ZFMK G.VII.1.a?.6., male, Lluch, 22.03.1913; ZFMK

G.VII.1.a7.1., male, Alcudia, 26.03.1913; ZFMK
G.VII.1.a?.«., male, Santany, 05.04.1913; ZFMK
G.VII.1.a7.r., male, Lluch, 18.04.1921; ZFMK

G.VII.1.a7.u., male juv., Albufera, 18.05.1921; ZFMK
G.VII.1.a.€., male juv., Alcudia, 04.06.1921; ZFMK
G.VII.1.a7.o., male juv., Alcudia, 04.06.1921; ZFMK
G.VII.1.a?.2., male, Alcudia, 18.05.1921; ZFMK
G.VII.1.a?.p., male, Valldemosa, 03.05.1921; ZFMK
G.VII.1.a7.o., Lluch, 09.04.1921; ZFMK G.VII.1.a?.t.,
Valldemosa, 28.03.1921; ZFMK G.VII.1.a7.v., female, Al-
cudia, 10.06.1927, all leg. A. v. Jordans.

Described from a series of twenty-seven individuals, twen-
ty registered in inventory of ZFMK bird skin collection,
one specimen exchanged with MfN, Berlin. Whereabouts
of other specimens unknown. — Dickinson (2003) presum-
ably: Troglodytes troglodytes kabylorum E. Hartert,
1910.

Troglodytes parvulus hyrcanus Zarudny & Loudon, 1905
Orn Monatsber 13: 107.

Paralectotype: ZFMK G.VII.1.a4. o., male, Kuljam-
Chadschi-Ali, W Iran, 26.10.1903 (= 13.10.1903 Grego-
rian Calendar), Coll. Zarudny No 1638, ex Coll. Jordans.

No information on material was given in the original de-
scription; no types designated. The whereabouts of the oth-
er specimens are unknown; the ZFMK specimen was la-
belled “cotype” by A. von Jordans subsequently and de-
fined as paralectotype by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Troglodytes troglodytes hyr-
canus.

©ZFMK

60 Renate van den Elzen

SITTIDAE

Sitta dresseri Zarudny & Buturlin, 1906
Orn Monatsber 14: 132.

Paralectotype: ZFMK G.IX.1.e!.a., male, Charadschi
(=Kharaji), W Iran, 14.04. (= 28.04. in the Gregorian Ca-
lendar) 1904, leg. Zarudny No 1631, ex coll. A. von Jor-
dans 1068.

Described from a series of unknown size; measurements
are given from eight specimens. Zarudny & Harms (1923)
mention twelve specimens collected, so according to
Mlikovsky (2007) the syntype series consisted of twelve
specimens. Specimen of ZFMK labelled by A. v. Jordans
as “cotype” and published in Jordans (1923: 18); lecto-
type in NHMW, Vienna, for details see Mlikovsky (2007).
— Dickinson (2003): Sitta tephronota dresseri.

Sitta tschitscherini Zarudny, 1904
Orn Jahrb 15: 218.

Paralectotype?, ZFMK G.IX.1.d3. a., male ad., Sarchun,
(Sar Khun), NW Iran, 21.12. 1903 (= 03.01.1904 in the
Gregorian Calendar), coll. Zarudny No 1484, ex Coll. A.
von Jordans 1064.

In original description three specimens from Ispahan (=Es-
fahan), Iran, presented with measurements, but no dates;
ZFMK specimen labelled by A. v. Jordans as “cotype”,
also registered as ““cotype” in inventory and published as
cotype in Jordans (1923: 18). Specimen not included in
Mlikovsky (2007). — Dickinson (2003): Sitta neumayer ts-
chitscherini.

Sitta europaea rubiginosa Tschusi & Zarudny, 1905
Orn Jahrb 16: 140.

Paralectotypes ?: ZFMK G.IX.1.a!!. a., male, Sermab-
Dasht, N Iran, 09.10.1903 (= 22.10.1903 in the Gregori-
an Calendar), leg. Zarudny No 1633, ex coll. A. von Jor-
dans No 1055; ZFMK G.IX.1.a!!. B., female, Sermab-
Dasht, N Iran, 9.10.1903 (= 22.10.1903 in the Gregorian
calendar), leg. Zarudny No 1823, ex coll. A. von Jordans
No 1056.

Type specimen designated in original description; male,
Asterabad, 09.10.1903, No 5501, Coll. v. Tschusi; prob-
ably in the NHMW, Vienna; paralectotypes labelled by A.
von Jordans as “cotypes”, also registered as such in in-
ventory of the ZFMK bird collection. — Dickinson (2003):
Sitta europaea rubiginosa.

Bonn zoological Bulletin 59: 29-77

Sitta reichenowi Kleinschmidt, 1917
Falco 13: 21.

Lectotype: Coll. Kl. 5058, male, Myslowitz, Silesia,
29.03.1911, leg. Natorp.

Very poor description, type designated only in Catalogue
(Kleinschmidt 1935-1943: 108). There also the numbers
Coll. Kl. 5059-5062 are listed under the same species
name and locality and may be paralectotypes. — [Dickin-
son (2003)]: Sitta europaea europaea> caesia

Sitta hassica Kleinschmidt, 1917
Falco 13: 22.

Lectotype: Coll. Kl. 5000, female, Kornsand/Rhine,
30.08.1889, leg. Kleinschmidt.

Paralectotype: Coll. KI. 5001, male, Knoblochsau/Rhine,
near Darmstadt, 17.09.1899, leg. Kleinschmidt.

Very poor description, no type specimens defined in e1-
ther description or in Catalogue (Kleinschmidt
1935-1943: 108). Lectotype and paralectotype designat-
ed by Rheinwald & van den Elzen (1984). — Dickinson
(2003): Sitta europaea caesia Wolf, 1810.

Sitta Auto-Sitta hoerningi Kleinschmidt, 1928
Berajah, Sitta Auto-Sitta: 6.

Holotype: Coll. Kl. 5024, female, Mehlis, Thuringian For-
est, 30.04.1928, leg. Horning.

Paralectotypes: Coll. Kl. 5021-5023, 2 males, female,
17.02.1920 and 30.04.1928, and 5025-5027, 2 females,
0, 22.09., 14.12.1919 and 22.09.1928, locality and collec-
tor like holotype.

In the original description only the holotype is designat-
ed, other topotypical specimens are not specified in de-
tail. Paralectotypes designated by Rheinwald & van den
Elzen (1984), based on entry in Catalogue (Kleinschmidt
1935—1943: 108). — Dickinson (2003): Sitta europaea cae-
sia Wolf, 1810.

Sitta Auto-Sitta dalmatina Kleinschmidt, 1928
Berajah, Sitta Auto-Sitta: 13.

Nomen novum for Sitta caesia longirostris A. E. Brehm,
1866; Verz. Sammlungen., p.6 (nomen nudum).

Holotype: Coll. Kl. 5013, male, Castelnuovo, Dalmatia,
24.03.1914.

©ZFMK

Types in ZFMK: Section Ornithology 61

Paratype: Coll. KI. 5014, female, Castelnuovo, Dalmatia,
11.09.1912. — Dickinson (2003): Sitta europaea cisalpi-
na Sachtleben, 1919.

Sitta siciliae Kleinschmidt & Jordans, 1932
Falco 28: 6.

Holotype: Coll. KI. 5012, male, Troina, Sicily, 18.09.1929,
per Rosenberg.

Paratypes: ZFMK G.IX.1.a°.a. and B., 2 females, Enna,
Sicily, 10.09.1929, per Rosenberg.

— Dickinson (2003): Sitta europaea cisalpina Sachtleben,
1919.

Sitta Auto-Sitta bacmeisteri Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 108.

Holotype: Coll. Kl. 4989, male, Vitarville, Departement
Meuse, France, 15.09.1916, leg. Bacmeister.

Only this specimen was labelled as “bacmeisteri”’, how-
ever, Coll. Kl. 4990-4997 are listed as “bacmeisteri” in
the Catalogue (Kleinschmidt 1938-1943), too. These
might therefore be recognized as paratypes. — Dickinson
(2003): Sitta europaea caesia Wolf, 1810.

Sitta europaea itschangensis Kleinschmidt & Weigold,
1922
Abh Ber Mus Tierk Dresden 15: 9.

Paralectotype: Coll. KI. 5121, male, near Itschang, Chi-
na, 14.11.1914, leg. Weigold.

Described from a series of nine specimens. The holotype
was at the SNS, Dresden C 23936 and was lost in 1945;
topotypical material exists in Dresden (Eck, 1970), des-
ignated by Eck & Quaisser (2004) as syntypes; paralec-
totype status designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Sitta europaea sinensis Ver-
reaux, 1871.

Sitta (europaea) tibetosinensis Kleinschmidt & Weigold,
1922
Abh Ber Mus Tierk Dresden, 15: 10.

Paralectotype: Coll. Kl. 5132, male, Samendo, E Tibet,
16.10.1915, leg. Weigold.

Described from three specimens; type specimen at the
SNS, Dresden. C 23942, lost in 1945; (Eck, 1970); third

Bonn zoological Bulletin 59: 29-77

specimen in Dresden (Eck & Quaisser 2004). Paralecto-
type status designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003) presumably: Sitta nagaensis
montium La Touche, 1899.

CERTHIIDAE

Certhia rhenana Kleinschmidt, 1900
Orn Monatsber 8: 169.

Holotype: Coll. Kl. 5222, male, Gunterhausen, banks of
the River Rhine between Mainz and Worms, 08.11.1897,
leg. Kleinschmidt.

Paralectotypes: Coll. KI. 5223-5227, 3 males, female, o,
Knoblochsau/Rhine near Darmstadt, 17.09.1899,
09.01.1898, 11.01.1898, 15.08.1892, leg. Kleinschmidt.

Only type specimen designated in the original description,
paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Certhia familiaris macro-
dactyla C. L. Brehm, 1831.

Certhia brachydactyla nigricans Jordans, 1931
Orn Monatsber 39: 181.

Holotype: ZFMK G.VHUI.1.b°.1., male, Mosqueruela, Ter-
nel, E Spain, 09.01.1931, leg. Griin.

Paratypes: ZFMK G.VIII.1.b°.B., male, Mosqueruela, E
Spain, 02.01.1931; ZFMK G.VIIL.1.b°%.«., 0, Mosqueru-
ela, Ternel, 05.01.1931; ZFMK G.VIII.1.b°.2., 0, Mos-
queruela, Ternel, 05.01.1931, all leg. Griin.

Described from six specimens collected in January. Fur-
ther two individuals from the same locality, dated
12.1930 and 02.1933, are labelled as “cotypes”, though
only birds from January are mentioned in the original de-
scription. — Dickinson (2003): Certhia brachydactyla
brachydactyla C. L. Brehm, 1820.

Certhia brachydactyla obscura Jordans, 1934
Alauda 5: 522.

Holotype: ZFMK G.VIII.1.b8.a., male, Lagunilly-Bejar,
Spain, 27.10.1933, leg. Griin.

Paralectotypes: ZFMK.G.1.b8.¢. male, Lagunilla,
19.10.1933; ZFMK.G.1.b8.c., female, Lagunilla,
20.10.1933, all leg. Griin.

©ZFMK

62 Renate van den Elzen

Described from six specimens, only the holotype is des-
ignated; in the footnote further five specimens are men-
tioned as newly arrived and “perfectly matching”. The five
specimens in the ZFMK skin collection, dated from De-
cember 1933 were therefore not recognized as paratypes
(Rheinwald & van den Elzen 1984). Two paratypes were
exchanged with the AMNH, New York, one handed to
Fouard. — Dickinson (2003): Certhia brachydactyla
megarhyncha C. L. Brehm, 1831.

STURNIDAE

Sturnus balcanicus Buturlin & Harms, 1909
Orn Monatsber 17: 56.

Syntypes: ZFMK F.III.1.a°. a., male, Barza, Romania,
01.06.1901, ex Coll. Harms No 1143; ZFMK FIII.1.a5.B.,
male, Fetsci, 20.03.1907, ex Coll. Harms 1243; ZFMK
F.III.1.a5. 6., 2 females, Cernovoda, 24.04.1906, ex Coll.
Harms No 1249.

Described from a series of twelve specimens; whereabouts
of the other specimens uncertain. — Dickinson (2003):
Sturnus vulgaris vulgaris Linnaeus, 1758.

TURDIDAE

Turdus merula mallorcae Jordans, 1950
Syll Biol, Festschr Kleinschmidt: 172.

Lectotype: ZFMK_ H.III.1.a8.6, male, Arta, Majorca,
22.04.1913, leg. A. von Jordans No 802.

Paralectotypes: ZFMK H.III.1.a8.a., male, Esporlas,
20.05.1913; ZFMK HLIII.1.a8.B., Arta, Majorca,
23.04.1913; ZFMK H.III.1.a8.y., female, Arta, Majorca,
24.04.1913; ZFMK H.III.1.a8.6., female, Santany,
07.05.1913; ZFMK H.III.1.a8.¢., male, Arta, Majorca,
22.04.1913, leg A. v. Jordans No 804; ZFMK H.III.1.a8.C.,
Santany, 07.05.1913; ZFMK H.II.1.a8.1., male, Alcudia,
Mallorca, 05.04.1913; ZFMK H.III.1.a8.«, male, Arta, Ma-
jorca, 22.04.1913, leg A. V. Jordans 803; ZFMK
H.JIT.1.a8.2., male, Arta, 03.04.1921; ZFMK H.III.1.a8.u..,
male, Arta, 31.03.1921; ZFMK H.III.1.a8.v., male juv., Al-
cudia, 09.06.1921; ZFMK H.III.1.a8.€., male juv., Alcu-
dia, 09.06.1921; ZFMK H.II.1.a8.0., male juv., Alcudia,
22.06.1921; ZFMK H.III.1.a8.2., female juv., Santany,
04.05.1913; ZFMK H.II.1.a8.p., male juv., Albufera,
18.05.1921, all leg. A. v. Jordans.

Bonn zoological Bulletin 59: 29-77

Described from seventeen specimens, sixteen still in the
ZFMK collections. Three males are registered in the in-
ventory under 22.04.1913; information on type status tak-
en from label in collection of ZFMK (Rheinwald & van
den Elzen 1984). One specimen has been exchanged with
the AMNH, New York. — Dickinson (2003) presumably:
Turdus merula merula Linnaeus, 1758.

Turdus hispaniae Kleinschmidt, 1909
Falco 5:22.

Lectotype: Coll. Kl. 3378, male, Malaga, Spain,
24.09.1903, per Schliter.

Paralectotypes: Coll. Kl. 3379-3380, 2 male, Malaga,
15.01.1904 and 15.04.1905, per Schliter.

No types designated in the original description; types and
paratypes designated by Kleinschmidt (1935-1943: 84) in
his Catalogue. — Dickinson (2003): Turdus merula meru-
la Linnaeus, 1758.

Turdus riidigeri Kleinschmidt, 1919
Falco 14: 15.

Holotype: Coll. Kl. 3367, female, Chambley, E France,
18.04.1918, leg. Riidiger.

Paralectotypes: Coll. KI. 3364-3366, male, male juv., fe-
male, Cote Chambley, E France, and “France”,
21.01.-18.04.1918, leg. Riidiger.

Only holotype specified; paralectotypes designated by
Rheinwald & van den Elzen (1984). — Dickinson (2003):
Turdus merula merula Linnaeus, 1758.

Turdus ultrapilaris Kleinschmidt, 1919
Falco 14: 16.

Lectotype: Coll. Kl. 3297, male, Issyk-Kul, W Asia,
05.01.1908, leg. Tancré.

Paralectotypes: Coll. KI. 3298-3299, male, female, Issyk-
Kul, 05.01.1908, leg. Tancré.

Rather poor description, no type defined. Information on
type status from Catalogue (Kleinschmidt 1935-1943: 83);
paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Turdus pilaris Linnaeus,
1758.

©ZFMK

Types in ZFMK: Section Ornithology 63

Turdus pseudohodgsoni Kleinschmidt, 1909
Falco 5: 20.

Holotype: Coll. Kl. 3262, male, Taschkent, Russia,
02.03.1908.

There was apparently no further material at hand for the
description. — Dickinson (2003): Turdus viscivorus visci-
vorous Linnaeus, 1758.

Turdus viscivorus hispaniae Jordans, 1950
Syll Biol, Festschr Kleinschmidt: 171.

Holotype: ZFMK 46.179, male, Linares de Riofrio, Spain,
08.04.1943, leg. Grin.

Paratypes: ZFMK 46.180, male, Linares, 01.04.1943;
ZFMK 46.181, male, Linares, 03.04.1942; ZFMK 46.182,
male, Linares, 04.04.1942: ZFMK 46.183, male, Linares,
10.04.1942; ZFMK 46. 184, female, Linares, 04.04.1942;
ZFMK 46.185, female, Linares, 04.04.1942 ; ZFMK 46.
186, female, Linares, 04.04.1942: ZFMK 187, female, Li-
nares, 04.04.1942; ZFMK 46.188, female, Linares,
04.04.1942, all leg. H. Griin.

Described from five males and four females; preoccupied
by Turdus hispaniae Kleinschmidt, 1909. — Dickinson
(2003): see following taxon.

Turdus viscivorus jordansi Eck, 1970

Zool Abh Mus Tierk Dresden 30: 135

Holotype: ZFMK 46.179, male, see previous taxon.
Nomen novum for 7urdus viscivorus hispaniae Jordans,
1950, this taxon being preoccupied by Turdus hispaniae

Kleinschmidt, 1909. — Dickinson (2003): Turdus vis-
civorus viscivorous Linnaeus, 1758.

MUSCICAPIDAE
Phoenicurus phoenicurus caesitergum Clancey, 1947

Bull Brit Orn Cl 67: 77.

Paralectotype. ZFMK 50.56, female, Northumberland,
England, 31.05.1942, leg. Clancey.

Holotype in the Coll. Clancey, RSME, Edinburgh, t. t.

Burnley, Lancashire, England. — Dickinson (2003):
Phoenicurus pheonicurus phoenicurus (Linnaeus, 1758).

Bonn zoological Bulletin 59: 29-77

Erithacus algeriensis Kleinschmidt, 1904
Orn Monatsber 12: 197.

Lectotype: Coll. Kl. 2921, male, Lambése, “in the
wood”, Algeria, 11.06.1904, leg. Fliikiger.

Paralectotypes: Coll. KL. 2922-2924, 3 male, Lambese,
9 11.06.1904, leg. Fliikiger.

No type defined in the original description; information
on type status taken from Catalogue (Kleinschmidt
1935-1943: 73). — Dickinson (2003): Phoenicurus pheon-
icurus phoenicurus (Linnaeus, 1758).

Phoenicurus ochruros aterrimus Jordans, 1923
Falco 19, Sonderheft: 8.

Lectotype: Coll. KI. 2804, male, Bellas, Portugal, 06.1906.

Paralectotypes: Coll. KI. 2805, female, Bellas, 06.1906;
ZFMK H. IL. 6. a’.f., male, Pinchao bei Aligo, 18.03.1913,
leg. Weigold.

No type specimens designated in original description; two
specimens in the Kleinschmidt Collection match the date
and locality provided in the description; type status des-
ignated in Catalogue Kleinschmidt (1935-1943: 72) and
also on label in the ZFMK collections. Second paralecto-
type designated by van den Elzen (1986). — Dickinson
(2003): Phoenicurus ochruros aterrimus.

Saxicola rubetra hesperophila Clancey, 1949
Limosa 22: 369.

Paralectotypes: ZFMK 50.312—50.315, male, 2 females,
female juv., Argyllshire, SW Scotland, 17.07 —18.08.1948,
leg. Clancey.

Holotype in the Coll. Clancey, RSME, Edinburgh, t. t.
Newton Mearns, E Renfrewshire. Only holotype designat-
ed in the original description. Information on paralecto-
types derived from ZFMK inventory and labels in the col-
lection; paralectotype status designated by Rheinwald &
van den Elzen (1984). — Dickinson (2003): Saxicola ru-
betra (Linnaeus, 1758).

Saxicola rubetra sengiini Kumerloeve, 1969
Rev Fac Sci Univ Istanbul 34: 277.

Syntypes: ZFMK 68.1873, female, Yiiksekova, Turkey,
10.06.1968; ZFMK 68.1874, male, Van-Ermaniz,
23.06.1968; ZFMK 68.1875, male juv., Van-Ermaniz

©ZFMK

64 Renate van den Elzen

23.06.1968, leg. Mittendorf, Nos 201, 274, 275. — Dick-
inson (2003): Saxicola rubetra (Linnaeus, 1758).

Saxicola torquata gabrielae Neumann & Paludan, 1937
Orn Monatsber 45: 15.

Lectotype: ZFMK 41.492 male, Olymp, S of Brussa, Asia
Minor, 20.07.1934. Paralectotype: ZFMK 41.493, male,
Olymp, S of Brussa, Asia Minor, 20.07.1934, both leg. G.
Neuhauser.

Described from a series of nine males, five from “W and
C Asia Minor“ and four from Rhodos. Two of our speci-
mens meet the locality and date of the defined “type” of
the original description, the specimen ZFMK 41.492 bears
a label with “type” in an old handwriting. The second
specimen 1s chosen as a paralectotype. The whereabouts
of the remaining specimens are not known. — Dickinson
(2003): Saxicola torquata rubicola (Linnaeus, 1766).

Saxicola transfuga Kleinschmidt, 1936
Nova Acta Leopold, N.F. 4: 377.

Lectotype: Coll. Kl. 2547, male, Schuster, W_ Iran,
22.03.1904.

Paralectotype: Coll. Kl. 2548, female, Mamed-abad, E
Iran, 19.07.1901.

Neither in the original description nor in the Catalogue
(Kleinschmidt 1935—1943: 66) type specimens were des-
ignated. Type status was taken from the handwritten la-
bel; lectotype and paralectotype designated by Rheinwald
& van den Elzen (1984). — Dickinson (2003): Oenanthe
pleschanka transfuga, but see Haffer (1977: 37).

Saxicola Meridionalis reiseri Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 65 and 69.

Lectotype: Coll. KI. 2534, male, Mosco, Bosnia - Herze-
govina, 23.04.1893.

Paralectotype: Coll. KI]. 2535, female, Trebinje, Bosnia -
Herzegovina, 24.04.1893.

Very poor description, but illustrated on page 70; no types
designated in Catalogue (Kleinschmidt 1935-1943: 65);
lectotype and paralectotype designated by Rheinwald &
van den Elzen (1984). — Dickinson (2003): Oenanthe his-
panica hispanica (Linnaeus, 1758).

Bonn zoological Bulletin 59: 29-77

Saxicola palearctica Kleinschmidt, 1920
Falco 17: 3.

Lectotype: Coll. Kl. 2490, male, Sirax, Trans-Caspia,
20.03.1905, per Schliter.

Kleinschmidt (1935—1943: 65) considers specimens col-
lected at the same time and the same place (Coll. KI.
2491-2495, 2 males, male juv., 2 females, Sirax,
20.—26.03.1905) as questionably similar to the type. No
type specimen designated in the poor original description;
lectotype designated in Catalogue (Kleinschmidt
1935-1943: 65). — Dickinson (2003): Oenanthe oenanthe
libanotica (Hemprich & Ehrenberg, 1833).

Saxicola Borealis obtusa Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 64.

Lectotype: Coll. KI. 2466, male, Gennargentu, Sardinia,
28.06.1907, leg. Meloni.

Paralectotypes: Coll. Kl. 2467-2475, 2 males, 4 females,
3 imm., Gennargentu, 26.—31.07.1903, leg. Meloni.

Very poor description, no types designated, information
taken from label in Kleinschmidt Collection; lecto- and
paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Oenanthe oenanthe libano-
tica (Hemprich & Ehrenberg, 1833).

Saxicola Borealis exilipes Kleinschmidt, 1943
Katalog meiner ornithologischen Sammlung: 65.

Lectotype: Coll. KI. 2485, male ad., Sarpa/ Wolga, Rus-
sia, 05.1909.

Paralectotype: Coll. Kl. 2486, 0 ad., Sarpa/ Wolga,
05.1909.

Very poor description, no types designated, information
taken from label in Kleinschmidt Collection; lecto- and
paralectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Oenanthe oenanthe oenan-
the (Linnaeus, 1758).

Oenanthe tractrac hoeschi Niethammer, 1955
Bonn zool Beitr 6: 188.

Paratype: ZFMK 66.899, female, Kaokoveld w Orupem-
be, Namibia, 27.06.1952, leg. Hoesch.

Holotype: UMB, Bremen No 19.905, male, Kaokoveld w
Orupembe, Namibia, 28.06.1952, leg Hoesch.

©OZFMK

Types in ZFMK: Section Ornithology 65

Described from these two specimens. — Dickinson (2003):
Cercomale tractrac hoeschi (Niethammer, 1955).

Cercomela familiaris hoeschi Niethammer, 1955
Bonn zool Beitr 6: 189.

Paratype: ZFMK 39.458, male, Brandberg, Namibia,
17.09.1938, leg. G. Niethammer No 94.

Holotype at the UMB (Bremen) No 19.907, t.t. Otju,
Kaokoveld. Preoccupied by Oenanthe tractrac hoeschi
Niethammer, 1955, because the species ¢ractrac has to be
placed within Cercomela, if this is accepted as separate
genus. Niethammer treated this form as synonym of
Oenanthe familiaris angolensis later. — Dickinson (2003):
Oenanthe familiaris angolensis Lynes, 1926.

Erithacus superbus Koenig, 1889
J Oya SZ M38.

Syntypes: ZFMK_ H.II.4.a3.B. male, Icod el Alto,
Tenerife, Canary Islands, 17.02. 1889; ZFMK H.II.4.a3.6.,
male, Villa Orotava, Tenerife, Canary Islands, 06.04.1889
— specimen lost, all leg. Koenig.

Paralectotypes: ZFMK H.II.4.a3.a., male, Villa Orotava,
22.01.1889; ZFMK H.II.4.a3-y., male, Villa Orotava,
06.04.1889; ZFMK H.II.4.a3.¢., female, Icod el Alto,
17.02.1889; ZFMK H.II.4.a3.¢., 0, Icod el Alto,
17.02.1889; ZFMK H.II.4.a3.7., female, Pinar, 11.04.1889;
ZFMK_ H.II.4.a3.6., 0, Pinar, 13.04.1889; ZFMK
H.I1.4.a3.1., female, 16.04.1889, all leg. Koenig.

In the original description no type specimen was defined;
in the inventory register B. and 6. are designated as type
specimens. The latter is currently untraceable in the col-
lection, supposedly also integrated in the permanent ex-
hibition. All type specimens of Koenig were mounted in
the first place and have been prepared as skins only after
1943. Specimens numbered a, y and ¢ are registered as
“cotypes” in the inventory (marked by crosses), but 5 more
specimens from the same localities and time period are
also registered. They have been chosen as paralectotypes
by Rheinwald & van den Elzen (1984). — Dickinson
(2003): Erithacus rubecula superbus.

Erithacus rubecula hispaniae Jordans, 1950
Syll Biol, Festschr Kleinschmidt: 174.

Holotype: ZFMK 46.232, male, Linares de Riofrio, Sala-
manca, Spain, 20.04.1943, leg. H. Grin.

Bonn zoological Bulletin 59: 29-77

Paralectotypes: ZFMK 46.233, female, Linares de Riofrio,
28.04.1942; ZFMK 46. 234, male, Linares, 12.04.1942;
ZFMK 46.235, male, Linares, 30.04.1942, all leg. H.
Grin.

Described from three males and one female collected “‘end
of April, beginning of May”; only holotype designated in
original description. Paralectotypes designated by Rhein-
wald & van den Elzen (1984), and information on status
taken from inventory of ZFMK _ bird collection.
— Dickinson (2003): Erithacus rubecula rubecula (Lin-
naeus, 1758).

Erithacus Dandalus sardus Kleinschmidt, 1906
Falco 2: 70.

Lectotype: Coll. KI. 2985, male, Pizzo Tricoli, Ogliastra,
Sardinia, 26.05.1905.

Paralectotypes: Coll. Kl. 2986-2991, 5 males, 0, Pizzo Tri-
coli and Lanusei, Ogliastra, 30.08.1904, 04.06.1905,
02.—03.09.1906.

No types defined in the original description; lectotype des-
ignated by Kleinschmidt in his Catalogue (1935-1943:
75); paralectotypes designated by Rheinwald & van den
Elzen (1984). One para(lecto)type in the SNS, Dresden
(Eck & Quaisser 2004). — Dickinson (2003): Erithacus
rubecula sardus.

Erithacus rubecula monnardi Kleinschmidt, 1916
Falco 12: 14.

Lectotype: Coll. Kl. 2981, 0 ad., La Tourauderie, Dept.
Ardennes, France, 24.02.1916, leg. Bacmeister.

Paralectotypes: Coll. Kl. 2982-2983, 2 0, La Touraude-
rie, 24.02. and 17.03.1916, leg. Bacmeister.

Originally described from three specimens, lectotype des-
ignated in Catalogue (Kleinschmidt 1935—1943: 75), al-
so paralectotypes. — Dickinson (2003): Erithacus rube-
cula rubecula (Linnaeus, 1758)

Luscinia megarhynchos luscinioides Jordans, 1923
Falco 19, Sonderheft: 3.

Holotype: ZFMK_ H.II.2.a°.a., male, Arta, Majorca,
22.04.1913, leg. A. von Jordans No 839.

Paralectotypes: ZFMK H.II.2.a?.B., male, Santany,
09.05.1913; ZFMK H.II.2.a?.y., male, Arta, Majorca,

©ZFMK

66 Renate van den Elzen

26.04.1913; ZFMK H.II.2.a°.6., male, Arta, Majorca,
26.04.1913; ZFMK HLII.2.a°.¢., female, Valldemosa,
19:05.1913: | -ZEMKo” HAll2%a2:'Ga males santany:
09.05.1913; ZFMK H.II.2.a?.n., male, Arta, Majorca,
25.04.1913; ZFMK H.II.2.a?.6., male, Santany,
01.05.1913; ZFMK H.II.2.a7.1., male, Santany, 01.05.1913;
ZFMK H.II.2.a?.«., male, Santany, 09.05.1913; ZFMK
H.II.2.a?.u., female, Alcudia, 06.06.1921; ZFMK
H.II.2.a?.v., male, Alcudia, 08.05.1921; ZFMK H.LII.2.a?.€,,
male, Albufera, 16.05.1921; ZFMK H.II.2.a°.2., male, Al-
cudia, 20.05.1921; ZFMK HLII.2.a?.p., male, Alcudia,
14.06.1921; ZFMK HLII.2.a°.0., male, Alcudia,
06.05.1921; ZFMK HLII.2.a?7., male, Alcudia,
06.05.1921, all leg. A. von Jordans.

Described from twenty-six specimens; only holotype spec-
imen designated, no paratypes defined. Of the original se-
ries seventeen still exist in the ZFMK collection, two more
specimens were exchanged with the AMNH, New York,
and Tring, UK; whereabouts of others unknown. Paralec-
totypes designated by Rheinwald & van den Elzen (1984)
based on specimens traced in the ZFMK inventory. —
Dickinson (2003): Luscinia megarhynchos megarhynchos
C.L. Brehm, 1831.

Luscinia megarhynchos caligiformis Clancey & Jordans,
1950
Auk 67: 361.

Paratype: ZFMK 50.316, male ad., Martlesham, Wood-
bridge, E Suffolk, England, 11.06.1941, leg. Clancey.

Holotype at the RSME, Edinburgh, t. t. as above. — Dick-
inson (2003): Luscinia megarhynchos megarhynchos C.L.
Brehm, 1831.

Luscinia svecica weigoldi Kleinschmidt, 1923
Abh Ber Mus Tierk Dresden 16: 43.

Paralectotype: Coll. Kl. 3092, male, Bago, 90 km n of Je-
hol, NO Tschili, 13.05.1916, leg. Weigold.

Holo(?)type C 23 427 at the SNS, Dresden, lost since
1945, other type material still exists (Eck & Quaisser
2004).

Described from a series of eight males and four females.
No types designated in the original description, three more
paratype specimens in the SNS, Dresden. ZFMK speci-
men designated as paratype by Kleinschmidt (1935-1943:
77) in his Catalogue. — Dickinson (2003): Luscinia sve-
cica svecica (Linnaeus, 1758).

Bonn zoological Bulletin 59: 29-77

Erithacus gaetkei Kleinschmidt, 1904
J Ornithol 52: 302.

Lectotype: Coll. Kl. 3045, male, Heligoland, “spring”
1896, leg. Kemp, per de Maes.

Paralectotypes: Coll. K1. 3046-3051, 2 males, male juv.,
female, Heligoland, “spring” and “autumn” 1896, leg.
Kemp, per de Maes.

No type specimen defined in the description; the ,,type
specimen“ designated in the Catalogue (Kleinschmidt
1935-1943: 76). There is also information given in a foot-
note that a larger series existed for the original descrip-
tion. — Dickinson (2003): Luscinia svecica svecica (Lin-
naeus, 1758).

Erithacus volgae Kleinschmidt, 1907
Falco 3: 47.

Holotype: Coll. Kl. 3097, male, Sarpa a.d. Wolga, Rus-
sia, 05.1906, per Schliiter.

No further material existed for the brief description. —
Dickinson (2003): Luscinia svecica volgae.

Muscicapa striata tyrrhenica Schiebel, 1910
Orn Jb 21: 102.

Syntype: ZFMK 57.1307, female, Aitone near Evisa, Cor-
sica, 19.05.1910, leg. Schiebel.
Aitone,

Paralectotype: ZFMK 57.1306, female,

26.05.1910, leg. Schiebel.

The other syntype, the mated male, is housed at the
NHMW, Vienna (NMW 70.694, male, Aitone near Evisa,
Corsica, 19.05.1910, leg. Schiebel), also some other para-
lectotypes; see Schifter & van den Elzen (1986). — Dick-
inson (2003): Muscicapa striata tyrrhenica.

Muscicapa striata cretica Schiebel, 1925
J Ornithol 58: 659.

Syntypes: ZFMK 57.1016, male, Kalessor near Candia,
Crete, 07.06.1925, and ZFMK 57.1013, female, Canea,
Crete, 15.05.1925, both leg. Schiebel.

A presumed paralectotype is ZFMK 57.1015, male, Homa-
lo, Crete, 04.05.1925; see also Schifter & van den Elzen
(1986). — Dickinson (2003): Muscicapa striata neuman-
ni Poche, 1904.

©ZFMK

Types in ZFMK: Section Ornithology 67

Muscicapa striata balearica Jordans, 1913
Falco 9: 43.

Lectotype: ZFMK G_II.1.a°.a., male, S Valldemosa, Ma-
jorca, 19.05.1913, leg. A. von Jordans No 654.

Paralectotype: ZFMK G.II.1.a*.y., female, Santany, Ma-
jorca, 02.05.1913, leg. A. von Jordans.

In the original description no type specimen is designat-
ed; information on type status is taken from collection la-
bels and inventory of ZFMK collection (Rheinwald & van
den Elzen 1984). There fourteen specimens (including the
type specimen) are listed, but not defined as type materi-
al: ZFMK G.II.1.a*.B.—C., 0—-€., four males, seven females,
Valldemosa, Sta. Ponsa, Santany, Arta, Isla Dragonera,
16.04.-19.05.1913, all leg. A. von Jordans. Three speci-
mens were exchanged with the bird collections in War-
saw, Berlin and Dresden. — Dickinson (2003): Muscica-
pa striata balearica.

Bradornis pallidus aquaemontis Stresemann, 1937
Orn Monatsber 45: 148.

Paralectotype: ZFMK 39.972, male, Waterberg-Plateau,
Namibia, 10.02.1937, leg. Hoesch No 734.

Holotype at the MfN, Berlin No 37.20, leg. Hoesch No
783.

Described from three specimens, only holotype designat-
ed; whereabouts of third specimen unknown. — Dickinson
(2003): Malaenornis pallidus aquaemontis (Stresemann,
1939):

CINCLIDAE

Cinclus aquaticus tschusii Kleinschmidt & Hilgert, 1907
Falco 3: 104.

Lectotype: Coll. KI. 3463, male, Neustadt a.d. Wied, Ger-
many, 09.04.1896, leg. Sch. I (Schneider the Elder?)

Paralectotype: Coll. KI. 3464, 0, Grafenbach, 09.1906.

No type defined in the original description; “Typus” and
“Cotypus” designated by Kleinschmidt in his Catalogue
(1935-1943: 85). — Dickinson (2003): Cinclus cinclus
aquaticus Bechstein, 1803.

Bonn zoological Bulletin 59: 29-77

Cinclus cinclus szetschwanensis Meise, 1928
Orn Monatsber 36: 138.

Paratype: Coll. Kl. 3484, male, Taukwan/ Min, Szechu-
an, 15.01.1915, leg. Weigold; second paratype at the MfN,
Berlin, Nr. 25.1158, Tsaupo near Wonnutschwan,
31.12.1914, leg Weigold.

Holotype at the MfN, Berlin, No 25.1157, Taukwan/ Min,
Szechuan, 15.01.1915, leg. Weigold. — Dickinson (2003):
Cinclus cinclus przewalskii Bianchi, 1900.

Cinclus cinclus amphitryon Neumann & Paludan, 1937
Orn Monatsber 45: 16.

Lectotype: ZFMK 41.434, male, Varsambek, Lasistan,
Turkey, 26.08.1934, leg. Neuhauser.

Paralectotypes: ZFMK 41.435, male, Varsambek, Lasis-
tan, Turkey, 26.08.1934; ZFMK 41.436, male, Varsambek,
26.08.1934; ZFMK 41.437, male juv., Varsambek,
26.08.1934; ZFMK 41.438, male juv., Varsambek,
28.08.1934; ZFMK 41.439, male, Varsambek, 27.08.1934;
ZFMK 41.440, male, Varsambek, 25.08.1934; ZFMK
41.441, male, Varsambek, 26.08.1934, all leg. Neuhauser.

Described from six adult males, one juv. male, two fe-
males, ZFMK 41.442 exchanged with the AMNH, New
York. Three specimens with the same date and finding lo-
cality match the “type” as presented in the original descrip-
tion. Information on type status taken from handwritten
label in collection (Rheinwald & van den Elzen 1984). —
Dickinson (2003): Cinclus cinclus caucasicus Madarasz,
1903.

NECTARINITDAE

Anthreptes rectirostris amadoni Eisentraut, 1973
Bonn zool Monogr 3: 248.

Holotype: ZFMK 68.625, male ad., Dikuma, Rumpi Hills,
W Cameroon, 04.03.1967, leg. Eisentraut No 1188.

Only holotype designated in the original decription, no de-
tails on material examined included; further three adult
males, two juvenile males, one female, with the same da-
ta, are potential paratypes. — Dickinson (2003): Anthreptes
rectirostris tephrolaemus (Jardin & Selby, 1851).

©ZFMK

68 Renate van den Elzen

Nectarinia preussi parvirostris Eisentraut, 1965
J Ornithol 106: 219.

Holotype: ZFMK 65.6, male ad., Refugio at the peak of
St. Isabel, Fernando Po (Bioko), 18.12.1962, leg. Eisen-
traut No 664.

Paralectotypes: ZFMK 63.949—63.959, 11 males, Moca,
10., 11. 1962; ZFMK 63.960—63.962, 2 males, female,
Refugium, 01.1963 and 12. 1962; ZFMK 63.963—63.968,
7 females, Moca, 10., 11. 1962, 02., 03. 1963.

Described from a series of nineteen males and eleven fe-
males; Eisentraut collected fourteen males and seven fe-
males. No paratypes were defined in the original descrip-
tion; paralectotypes designated by Rheinwald & van den
Elzen (1984). — Dickinson (2003): Cinnyris reichenowi
preussi Reichenow, 1892.

Nectarinia minulla amadoni Eisentraut, 1965
J Ornithol 106: 218.

Holotype: ZFMK 65.7, male ad., San Carlos, Fernando Po,
02.10.1962, leg. Eisentraut No 153.

Paralectotypes: ZFMK 63.1079—63.1082, 2 males, 2 fe-
males, San Carlos, 09., 10., 11. 1962 and ZFMK 63.1083,
male, Conception 03.1963, all leg. Eisentraut.

Described from six males and three females. Only holo-
type, no paratypes designated in original description; para-
lectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Cinnyris minullus amadoni.

PASSERIDAE

Passer montanus Hispaniae Jordans, 1933
Anz Orn Ges Bayern 2: 253.

Holotype: ZFMK L.11.3.f+.B., male, Oropesa, E Pyrenees,
Spain, 28.10.1931, leg. Griin.

Paralectotypes: ZFMK L.II.3.f*.y. to @., 10 males, 9 fe-
males, Oropesa, 08.—11. 1931 and 02., 10.-11. 1932, all
leg. Griin.

Described from twenty specimens, only type specimen
designated in original description; paralectotypes desig-
nated by Rheinwald & van den Elzen (1984) as nineteen
topotypical specimens were found in the collection. —
Dickinson (2003): Passer montanus montanus (Linnaeus,
1758).

Bonn zoological Bulletin 59: 29-77

Passer catellatus Kleinschmidt, 1935
Falco Skizzen 2: 1.

Lectotype: Coll. KI. 185, male, Sussex, England, 12. 1888,
leg. Rosenberg, per Brazenor Bros.

Paralectotypes: Coll. Kl. 186, male, Sussex, 05.12.1896,
leg. Rosenberg; Coll. Kl. 187 male, Tring, 25.12.1933;
Coll. KI. 188, female, Tring, 26.12.1933, both leg. Good-
son.

Described from four specimens, no type designated in the
description, lectotype designated in his Catalogue (Klein-
schmidt 1935-1943: 7), paralectotypes designated by
Rheinwald & van den Elzen (1984). — Dickinson (2003):
Passer montanus catellatus.

Passer hostilis Kleinschmidt, 1915
Falco 11: 19.

Lectotype: Coll. KI. 30, male, Tring, Zoological Garden,
England, 09.08.1897, leg. Kleinschmidt.

Paralectotypes: Coll. Kl. 28-29, 2 males, Tring, 07. and
09.08.1897, leg. Kleinschmidt.

No type specimens designated in the original description,
only type locality. Coll. Kl. 30 designated as type speci-
men in the Catalogue (Kleinschmidt 1935—1943: 2), in the
collection No 29 labelled as type. — Dickinson (2003):
Passer domesticus domesticus (Linnaeus, 1758).

Passer ahasver Kleinschmidt, 1904
Orn Monatsber 12: 7.

Holotype: Coll. Kl. 153, male, Marrakesh, S Morocco,
03.12.1899, leg. Floericke.

Description based on one single specimen. — Dickinson
(2003): Passer domesticus tingitanus Loche, 1867 .

Passer niger Radde, 1899
Die Sammlungen des Kaukasischen Museums, |: 246 and
260.

Paralectotype: Coll. KI. 133, male, Kumbaschinsk, 20 km
N of Lenkoran, Caucasus, 08.04.1896, leg. Floericke.

According to Kleinschmidt correct name of Passer domes-
ticus hyrcanus Zarudny, 1923; paralectotype designated
by Kleinschmidt (1935-1943: 5). — Dickinson (2003):
Passer domesticus hyrcanus Zarudny & Kudashevy, 1916.

©ZFMK

Types in ZFMK: Section Ornithology 69

Passer domesticus mayaudi Kumerloeve, 1969
Rev Fac Sci Univ Istanbul 34: 258.

Holotype: ZFMK 68.2004, male, Van, Turkey, 30.06.1968,
leg. Mittendorf No 336.

Paralectotypes: ZFMK 68.1989, male, Van, 30.05.1968;
ZFMK 68.1994, male, Van, 30.05.1968: ZFMK 68. 1995,
male, Van, 19.06.1968; ZFMK 68.1996, male, Van,
19.06.1968; ZFMK 68.1997, male, Van, 19.06.1968:
ZFMK 68.1998, male, Van, 19.06.1968; ZFMK 68.1999,
male, Van, 19.06.1968; ZFMK 68.2000, male, Van,
20.06.1968; ZFMK 68.2001, male, Van, 22.06.1968;
ZFMK 68.2002, male, Van, 27.06.1968; ZFMK 68.2003,
male, Van, 27.06.1968; ZFMK 68.2004, male, Van,
30.06.1968; ZFMK 68.2005, female, Van, 19.06.1968;
ZFMK 68.2006, female, Van, 27.06.1968; all leg Mitten-
dorf, collection Kumerloeve.

Described from a series of twelve males and two females
from Van; only holotype designated in original descrip-
tion; paralectotypes designated by Rheinwald & van den
Elzen (1984), here restricted to specimens collected at the
type locality. — Dickinson (2003): Passer domesticus bib-
licus Hartert, 1904

Passer domesticus balearoibericus Jordans, 1923
Falco 19, Sonderheft: 4.

Holotype: ZFMK L.II.3.a9.y., male, Valldemosa, Major-
ca, 01.05.1921, leg. A. von Jordans No 3060.

Paralectotypes: ZFMK I.II.3.a?.a., B., d-@., @., aa.—éé.,
20 males, 6 females, Lluch, Arta, Valldemosa, Sta. Pon-
sa, Esporlas, Alcudia, 03.05. 1913 and 03.—06.1921.

Described from a large series (thirty-seven specimens), on-
ly holotype designated in original description, paralecto-
types designated by Rheinwald & van den Elzen (1984);
from the series, twenty-six specimens are still in the
ZFMK collection. — Dickinson (2003): Passer domesti-
cus balearoibericus.

Passer fliickigeri Kleinschmidt, 1904
Orn Monatsber 12: 7.

Holotype: Coll. Kl. 169, male, Touggourt, Algeria,
16.04.1903, leg. Fliikiger.

Paratype: Coll. Kl. 170, male, Oasis Touggourt,
16.04.1903, leg. Fliikiger.

Bonn zoological Bulletin 59: 29-77

Description based on larger series, one paratype in the
SNS, Dresden (Eck & Quaisser 2004). According to Meise
(1936) the status of the taxon as a stibilized hybrid pop-
ulation of Passer domesticus x Passer hispanoliensis. Not
in Dickinson (2003).

Passer italiae schiebeli Rokitansky, 1934

Falco 30: 7.

Holotype: ZFMK 57.765, male, Canaea, Crete,
11.04.1925, leg. Schiebel.

Paratypes: ZFMK 57.774, male, Canaea, Crete,

26.04.1925, “Cotype I’; ZFMK 57.800, male, Suda, Cre-
te, 27.04.1925, “Cotype IT’, all leg. Schiebel.

Rokitansky described the taxon from a large series of fifty
males and twenty females in the collection of the ZFMK.
The paratypes are labelled as cotypes. — Dickinson (2003):
Passer domesticus domesticus (Linnaeus, 1758); (but see
Schifter & van den Elzen 1986, Stephan 2000).

Passer griseus laeneni Niethammer, 1955
Bonn zool Beitr 6: 75.

Holotype: ZFMK 54.331, male, Bol, E bank of Lake Chad,
01.04.1954, leg. Laenen.

Paralectotypes: ZFMK 54.332, female, Bol, 28.03.1954;
ZFMK 54.466—54.467, female, male, 8. and 15. 04.1954:
ZFMK 54.1243, male, Archei/ Ennedi, Chad, 05.04.1954
and ZFMK 80.326—80.328, 3 male, Fada/ Ennedi, 19. and
20.04.1954, leg. Laenen.

Described from a series of eleven males and seven fe-
males. Only the holotype was designated in the original
description. Paralectotypes designated by Rheinwald &
van den Elzen (1984). Whereabouts of other specimens
are unknown. — Dickinson (2003): Passer griseus laeneni.

Petronia petronia balearica Jordans, 1923
Falco 19, Sonderheft: 4.

Holotype: ZFMK_ [.II.4.a°.6., male, Lluch, Majorca,
21.04.1921, leg. A. von Jordans, No 3040.

Paralectotypes: ZFMK_ I.11.4.a°.a., female, Santany,
05.05.1913; ZFMK L.11.4.a°.B., female, Valldemosa,
30.04.1921; ZFMK L.11.4.a°y., female, Lluch, Majorca,
22.04.1921; ZFMK I.11.4.a°¢., ,maleArta, 04.04.1921;
ZFMK I.I1.4.a°.¢., male, Lluch, Majorca, 22.04.1921; n.,
female, Valldemosa, 02.05.1921; ZFMK_ 1.11.4.a®.x.,
male, Valldemosa, 29.04.1921, all leg. V. Jordans.

©ZFMK

70 Renate van den Elzen

Described from a series of ten specimens; only holotype
designated in original description, paralectotypes desig-
nated by Rheinwald & van den Elzen (1984). — Dickin-
son (2003): Petronia petronia petronia (Linnaeus, 1766).

PLOCEIDAE

Malimbus ballmanni Wolters, 1974
Bonn zool Beitr 25. 290.

Holotype: ZFMK 72.799, 0, NW of Tai, between Caval-
ly River and Keibli, SW Ivory Coast, 1972, leg. Ballmann.

The only specimen; further sightings in Sierra Leone,
Guinea and Liberia; synonym is Ploceus (Malimbus)
goloensis Field, 1979 (Malimbus 1: 2). Figured in Rhein-
wald & van den Elzen (1984: 113). — Dickinson (2003):
Malimbus ballmanni.

ESTRILDIDAE

Estrilda nonnula eisentrauti Wolters, 1964
J Ornithol 105: 355.

Holotype: ZFMK 61.1407, male, Camp VI, Isobi,
Cameroon Mountains, Cameroon, 10.03.1958, leg. Eisen-
traut No 886.

Paralectotypes: ZFMK _ 61.1398, female, Isobi,
05.02.1958: ZFMK 61.1400, female, Buea, 01.01.1958;
ZFMK 61.1411, male, Camp II, Cameroon Mountains,
26.11.1958, leg. Eisentraut.

Described from a large series of thirty-four specimens; in
the original description only the holotype is specified;
paralectotypes designated by Rheinwald & van den Elzen
(1984); the other specimens are presumably housed at the
SNMS, Stuttgart. — Dickinson (2003): Estrilda nonnula

eisentrauti.
Hypargos niveoguttatus baddeleyi Wolters, 1972
Bonn zool Beitr 23: 89.

Holotype: ZFMK 68.1297, female, Nacala, N Mozam-
bique, 12. 1967, leg. Baddeley.

Paratypes: ZFMK 68.1294—-68.1296, 3 males, Nacala, 12.
1967, leg. Baddeley.

Bonn zoological Bulletin 59: 29-77

Described from eight males and two females (incl. type
specimen); only the Nacala specimens are housed at the
ZFMK, the Mikinandi specimens may be stored in the
DMD, Durban. — Dickinson (2003): Hypargos niveogut-
tatus macrospilotus Mearns, 1913.

Amandava subflava niethammeri Wolters, 1969
Bonn zool Beitr 20: 66.

Holotype: ZFMK 66.618, male, Sa da Bandeira, Huila,
Angola, 20.03.1966, leg. Ferreira.

Paralectotype: ZFMK 66.619, female, Sa da Bandeira,
20.03.1966, leg. Ferreira.

Described from a larger series with material from sever-
al museum collections; only holotype designated in orig-
inal description; paralectotype designated by Rheinwald
& van den Elzen (1984), information taken from handwrit-
ten label in ZFMK collection and inventory. — Dickinson
(2003): Amandava subflava niethammeri.

VIDUIDAE

Hypochera lorenzi Nicolai, 1972
J Ornithol 113: 236.

Holotype: ZFMK 79.997, female, Emene near Enugu, E
Nigeria, 26.11.1971; in captivity at the Max-Planck-Insti-
tute for Behavioral Physiology, Seewiesen, until
19.04.1974, leg. Nicolai.

Described from only this specimen. — Dickinson (2003):
Hypochera wilsoni (E. Hartert, 1901).

PRUNELLIDAE

Prunella modularis interposita Clancey, 1943
Bull Brit Orn Cl 64: 14.

Paralectotypes: ZFMK 50.7—50.8, 2 females, Dornoch,
Sutherlandshire, Scotland, 02.09.1938 and 22.06.1942.

Described from fifteen specimens, only holotype desig-
nated in the original description; housed in Coll. Clancey,
RSM, Edinburgh, t.t. as paratypes. The paralectotypes
were designated by Rheinwald & van den Elzen (1984)
from handwritten information on label and ZFMK inven-
tory. — Dickinson (2003): Prunella modularis hebridium
Meinertzhagen, 1934.

©ZFMK

Types in ZFMK: Section Ornithology 7\

MOTACILLIDAE

Motacilla flava raddei Harms, 1909
Orn Monatsber 17: 2.

Holotype: ZFMK H.IV.1>.b!2.y., male, Aschabad, Trans-
Caspia, 15.05. (= 28.05. Gregorian Calendar), no year, leg.
Tancré, ex. Coll. Harms.

Paralectotypes: ZFMK H.IV.1>.b!2.a., B., 6., 3 males,
Aschabad and Sarepta, S Russia, 04., 05., no year, per Tan-
cré’s collector.

Described from a series, but only the type was defined and
designated in the original description. Paralectotypes des-
ignated by Rheinwald & van den Elzen (1984), informa-
tion taken from Jordans (1923: 15) and the inventory of
the ZFMK bird collection. — Dickinson (2003): Motacil-
la flava feldegg Michahelles, 1830.

Motacilla alba arduenna Kleinschmidt, 1916
Falco 12: 14.

Lectotypes: Coll. KI. 1775, male, La Tourauderie, Arden-
nes, France, 07.03.1916; Coll. KI. 1776, male, La Tourau-
derie, 07.03.1916 ; Coll. K1. 1777, male, La Tourauderie,
03.03.1916, all leg. Bacmeister.

No types were designated in the short original description;
series designated as lectotypes by Kleinschmidt in his Cat-
alogue (1935-1943: 43). — Dickinson (2003): Motacilla
alba alba Linnaeus, 1758.

Anthus Arboreus schliiteri Kleinschmidt, 1920
Falco 16: 16.

Syntypes: Coll. Kl. 1650-1651, male, female, Naryn,
Turkestan, 20.04.1906, Tienschan, 12.05.1908, leg. Schlii-
ter.

No further material existed. — Dickinson (2003): Anthus
trivialis trivialis (Linnaeus, 1758).

Anthus trivialis salomonseni Clancey, 1950

Dansk Orn Foren Tidsskrift 44: 43.

Paralectotypes: ZFMK 50.281 and 50.282, 2 males, Burn-

ley, Lancashire, SW Scotland, 08. and 15.06.1946, leg.
Clancey.

Bonn zoological Bulletin 59: 29-77

Described from a large series of forty-five specimens from
the breeding season; type specimen in the Coll. Clancey,
RSME, Edinburgh, t.t. Carmunnock, Lanarkshire. Para-
lectotypes designated by Rheinwald & van den Elzen
(1984). Information on status derived from ZFMK inven-
tory. — Dickinson (2003): Anthus trivialis trivialis (Lin-
naeus, 1758).

Anthus spinoletta hesperianus Clancey, 1942
Bull Brit Orn Cl 62: 57.

Paralectotypes: ZFMK 50.72—50.75, 4 males, Blackwa-
terfoot, Arran, W Scotland, 15. and 16.08.1940, leg.
Clancey.

In the original description only the holotype was desig-
nated, no information 1s given on further material: para-
lectotypes designated by Rheinwald & van den Elzen
(1984), information derived from ZFMK inventory. Type
specimen in the Coll. Clancey, RSME, Edinburgh. — Dick-
inson (2003): Anthus spinoletta spinoletta Linnaeus, 1758.

Anthus pratensis whistleri Clancey, 1942
Bull Brit Orn Cl 63: 6.

Paralectotype: ZFMK 50.78, female, Dornoch, Sutherland,
N Scotland, 05.09.1938, leg. Clancey.

In the original description only the holotype was desig-
nated, no information is given on further material: para-
lectotypes designated by Rheinwald & van den Elzen
(1984), information derived from ZFMK_ inventory.
Three further specimens are registered in the inventory,
but not defined as paratypes. Type specimen in Coll.
Clancey, RSME, Edinburgh. — Dickinson (2003): Anthus
pratensis whistleri.

Anthus richardi grotei Niethammer, 1957
J Ornithol 98: 449-450.

Holotype: ZFMK 57.8, male, Onguma, Etosha, Namibia,
18.09.1956, leg. Hoesch No 2682.

Paratypes: ZFMK 57.9 to 57.11, 3 males, Onguma, all
17.09.1956, leg. Hoesch.

Type specimen erroneously indicated as ZFMK 57.7 in the

original description. — Dickinson (2003): Anthus cinnamo-
meus grotel.

OZFMK

72 Renate van den Elzen

FRINGILLIDAE

Fringilla coerulescens Koenig, 1889
J Ornithol 37: 183.

Syntypes: ZFMK I.II.1.c?. a. and 8., 2 males, Palma, Ca-
nary Islands, 29.03.1889, leg. Koenig.

No types designated in original description; only two spec-
imens with appropriate data registered in ZFMK collec-
tion. — Dickinson (2003): Fringilla coelebs palmae Tris-
tram, 1889, because the name is preoccupied by Fringilla
coerulescens Vieillot, 1817 (=Estrilda caerulescens).

Fringilla tintillon madeirensis Koenig, 1890
J Ornithol 38: 271.

Syntypes: ZFMK LII.1.c.¢. and n., male, female, Cama-
cho, San Antonio da Serra, Madeira, 23.12.1888, leg.
Koenig.

Described from these two specimens. — Dickinson (2003):
Fringilla coelebs maderensis Sharpe, 1888.

Fringilla coelebs balearica Jordans, 1923
Falco 19, Sonderheft: 4.

Holotype: ZFMK _ L.II.1.a.¢., male, Lluch, Majorca,
25.03.1913, leg. A. von Jordans No 818.

Paralectotypes: ZFMK [II.1.a.a.-y., C-o., aa—es., 20
males, 6 females, Lluch, Valldemosa, Arta, Esporlas, Al-
bufera, Majorca, 03., 05.1913, 03., 05.1921, leg. A. von
Jordans.

Originally thirty-nine specimens collected and subspecies
described from this series; twenty-nine specimens are reg-
istered in the ZFMK inventory, two exchanged. Only type
specimen designated by author, paralectotypes designat-
ed by Rheinwald & van den Elzen (1984) from informa-
tion in ZFMK inventory. — Dickinson (2003): Fringilla
coelebs balearica.

Fringilla coelebs schiebeli Stresemann, 1925
J Orn 58: 658.

Paratype: ZFMK 57.878, male ad., Canea, Crete,
27.03.1925, leg. Schiebel (“Cotype”).

Holotype at the MfN, Berlin (ZMB 75.55) (see Schifter
& van den Elzen 1986). — Dickinson (2003): Fringilla coe-
lebs schiebeli.

Bonn zoological Bulletin 59: 29-77

Fringilla caelebs gengleri Kleinschmidt, 1909
Falco'5: 13.

Holotype: Coll. Kl. 399, male, Hampstead, England,
22.04.1869, ex Mus. Sharpe.

Only type designated in original description and in the Cat-
alogue (Kleinschmidt 1935-1943: 11). — Dickinson
(2003): Fringilla coelebs gengleri.

Fringilla montifringilla subcuneolata Kleinschmidt,
1909
Falco 5: 14.

Holotype: Coll. Kl. 348, male, without location data,
Japan, without date, leg. Kobe.

Paratypes: Coll. Kl. 349-351, as above.

All come from their winter quarters. — Dickinson (2003):
Fringilla montifringilla Linnaeus, 1758.

Pyrrhula hauseri Kleinschmidt, 1920
Falco 17: 4.

Holotype: Coll. Kl. 862, male, Nafels, canton Glarus,
Switzerland, 04.01.1899, leg. Hauser.

The taxon was obviously described from one single spec-
imen only. — Dickinson (2003) presumably: Pyrrhula
pyrrhula europaea Vieillot, 1816.

Pyrrhula pyrrhula wardlawi Clancey, 1947
Bull Brit Orn Cl 67: 76.

Paralectotype: ZFMK 50.100, male, Pertshire, Scotland,
14.06.1946, leg. Plowden-Wardlaw.

Holotype in Coll. Clancey, RSME, Edinburgh; t. t. Kin-
loch Rannoch, Pertshire, 02.06.1946.

Described from a series of fourteen males, ten females and
two juveniles. Only type designated in original descrip-
tion, paralectotype designated by Rheinwald & van den
Elzen (1984). — Dickinson (2003): Pyrrhula pyrrhula
pileata W. MacGillivray, 1837.

©ZFMK

Types in ZFMK: Section Ornithology 73

Pyrrhula pyrrhula paphlagoniae Roselaar, 1995
Songbirds of Turkey, Pica Press: 24—25.

Holotype: ZFMK 39.177, male, Karadere near Bolu,
Black Sea Coastlands, Turkey, 25.09.1934, leg. H. Réss-
ner.

A series of nine specimens was considered. The type spec-
imen is illustrated in Kirwan (2006: plates 7-8). — Dick-
inson (2003): Pyrrhula pyrrhula paphlagoniae.

Chloris sinica tschiliensis Jacobi, 1923
Abhandl Ber Mus Tierk Dresden 16: 25.

Syntype: Coll. Kl. 506, female juv., Beijing, China,
26.06.1916, leg. Weigold, per Stétzner.

The subspecies was described from a series of six males,
five females and the juvenile. The whereabouts of the oth-
er specimens are unknown, other syntypes apparently not
in the SNS, Dresden, as given in Rheinwald & van den
Elzen (1984), see Eck & Quaisser (2004).

Chloris chloris mallorcae Jordans, 1923
Falco 19, Sonderheft: 3.

Holotype: ZFMK 1.11.10.a°.0., male, Alcudia, Majorca,
14.05.1921, leg. A. von Jordans No. 3113.

Paralectotypes: ZFMK I.II.10.a°.a.—., u.—€., 1.-v., and y,,
15 males, 4 females, Santany, Alcudia, Arta, Esporlas, An-
draitx, Valldemosa, Majorca, Cabrera, Balearic Islands,
03.—05.1913, 05., 06. 1921, leg. A. von Jordans.

Described from a large series of twenty-seven males and
five females; only the holotype was designated in the orig-
inal description, paralectotypes designated by Rheinwald
& van den Elzen (1984). Only twenty-two specimens (in-
cluding the holotype) are found in the ZFMK collection,
two further were exchanged with the MfN, Berlin and the
AMNH, New York. The whereabouts of the remaining
specimens are unknown. — Dickinson (2003) presumably:
Carduelis chloris aurantiiventris (Cabanis, 1851),

Chloris chloris harrisoni Clancey, 1940
Ibis Ser. XIV, 3: 92.

Paralectotype: ZFMK 50.90, male, Renfrewshire, Scot-
land, 12.12.1937, leg. Clancey.

Holotype in Coll. Clancey, RSME, Edinburgh; Thornton-
hall, Lanarkshire, Scotland, 03.11.1937.

Bonn zoological Bulletin 59: 29-77

Described from a series of eighteen specimens; only the
holotype was designated by the author, paralectotype des-
ignated by Rheinwald & van den Elzen (1984), informa-
tion from ZFMK inventory and handwriting on label. —
Dickinson (2003): Carduelis chloris harrisoni.

Chloris chloris restricta Clancey, 1943
Bull Brit Orn Cl 63: 65.

Paralectotype: ZFMK 50.93, male, Wiltshire, England,
12.12.1942, leg. Clancey.

Described from a series of thirty specimens, only holo-
type designated in the original description; paralectotype
designated by Rheinwald & van den Elzen (1984), infor-
mation taken from inventory. Type specimen in Coll.
Clancey, RSME, Edinburgh. — Dickinson (2003): Cardu-
elis chloris harrisoni (Clancey, 1940).

Chloris chloris meridionalis Harms, 1910
Orn Monatsber 18: 121.

Syntypes: ZFMK I.II.10.a.€6., female, Colentina, Roma-
nia, 18.03.1904; ZFMK L1II.10.a.00., male, Colentina,
18.03.1904; ZFMK_ I.11.10.a.am., female, Fundeni,
05.03.1905; ZFMK_ I.II.10.a.cc., male, Colentina,
01.03.1906; ZFMK. L1II.10.a.co., male, Pantelimon,
13.02.1907, Coll. Harms No 1218-1222, Coll. A. von Jor-
dans No 1843-47. — Dickinson (2003): Carduelis chloris
muehlei Parrot, 1905.

Carduelis rhenanus Kleinschmidt, 1936
Katalog meiner ornithologischen Sammlung: 14.

Holotype: Coll.Kl. 514, 0, Kornsand/ Rhine, Germany,
03.05. 887.

Paratype: Coll. Erlanger 743, female, Rheinhessen, N of
Ingelheim, 15.05.1899.

Paratype probably at the SNS, Frankfurt/Main. — Dickin-
son (2003): Carduelis carduelis carduelis (Linnaeus,
1758).

Carduelis carduelis schiebeli Jordans & Steinbacher,
1943
Senckenbergiana 26: 75.

Holotype: ZFMK 46.22, female, Suda Bay near Kanea,
Crete, 15.03.1925, leg. Schiebel.

©OZFMK

74 Renate van den Elzen

Paralectotypes: ZFMK 57.1027—57.1033, 5 males, 2 fe-
males, Suda Bay, Topelia, lake of Kurna, 02. and
04.1925, all leg. Schiebel.

Described from a series of five males and three females;
only holotype defined by the authors. The paralectotypes
were designated by Rheinwald & van den Elzen (1984).
— (Dickinson 2003): Carduelis carduelis balcanica
Sachtleben, 1919.

Carduelis carduelis propeparva Jordans, 1923
Falco 19, Sonderheft: 4.

Holotype: ZFMK [.II.14.a8.o., male, Valldemosa, Major-
ca, 03.05.1921, leg. A. von Jordans No 3189.

Paralectotypes: ZFMK I.II.14.a8.a.—6., ¢.—-y,, 0., 1., T.—O.,
aa. and BB., 10 males, 10 females, Valldemosa, Lluch, Es-
porlas, Santany, Alcudia, Cap de Pinar, Albufera, Palma,
03.—05.1913 and 03. -05.1921, leg. A. von Jordans.

The subspecies was described from a large series of four-
ty-four specimens. In the original description only the
holotype was designated. Paralectotypes were designat-
ed by Rheinwald & van den Elzen (1984) based upon
twenty-six specimens registered in the inventory of the
ZFMkK bird collection; five specimens exchanged with the
AMNH, New York and Mayaud. — Dickinson (2003): Car-
duelis carduelis parva Tschusi, 1901.

Loxia Flavirostris parallelicolor Kleinschmidt, 1921
Berajah, Loxia Flavirostris: 1.

Lectotype: Coll. Kl. 730, male, Harwich, England,
21.01.1881, leg. Burton.

Paralectotype: Coll. Kl. 731, male, Rottingdean, Sussex,
England, 10.11.1896, leg. Brazenor Bros.

No types designated in the original description, designat-
ed subsequently by Kleinschmidt in his Catalogue (Klein-
schmidt 1935-1943: 19). — Dickinson (2003): Carduelis
flavirostris pipilans (Latham, 1787).

Acanthis flavirostris miniakensis Jacobi, 1923
Abhandl Ber Zool Mus Dresden 16: 25.

Syntype: Coll. Kl. 733, male, Tschuwo, Bameh, Szech-
wan, 11.11.1915, leg. Weigold per Stétzner No XII.235.

Described from a series of seven males, one male (mis-
spelling for female?) and two pulli. Further syntypes at

Bonn zoological Bulletin 59: 29-77

the SNS, Dresden, (see Eck & Quaissser, 2004). — Dick-
inson (2003): Carduelis flavirostris miniakensis.

Acanthis cannabina autochthona Clancey, 1946
Bull Brit Orn Cl 66: 84.

Paralectotypes: ZFMK 50.86—S0.87, 2 males, Lanarkshire,
SW Scotland, 09.11.1935; 13.03.1937.

Described from a series of five males, two females and
two juveniles; only holotype designated by the author;
paralectotypes designated by Rheinwald & van den Elzen
(1984) from information in inventory and handwritten la-
bel. Holotype (and other specimens?) in the Clancey Coll.,
RSME, Edinburgh. — Dickinson (2003): Carduelis
cannabina autochthona.

Acanthis cannabina obscura Jordans, 1923
Orn Monatsber 31: 13.

Paralectotype: ZFMK I.II.16.a°.a., female, Oporto, Por-
tugal, 15.03.1913, leg. Weigold.

Described from a series of three specimens; only the holo-
type was designated in the original description. Paralec-
totype designated by Rheinwald & van den Elzen (1984),
as specimen was labelled as “Cotype”. Type and third
specimen at the MfN, Berlin. — Dickinson (2003): Car-
duelis cannabina cannabina (Linnaeus, 1758).

Carduelis deichleri Kleinschmidt, 1936
Katalog meiner ornithologischen Sammlung: 17.

Lectotypes: Coll. KI. 632-634, 3 males, Holley, New York,
30.03., 04. and 11.04.1888, leg. Craven No 41-43.

In the very short description no types are designated, not
defined as types in the Catalogue (Kleinschmidt
1935-1943), but only these two specimens are registered.
Lectotypes designated by Rheinwald & van den Elzen
(1984). — Dickinson (2003): Carduelis flammea flammea
(Linnaeus, 1758).

Citrinella corsicana Koenig, 1899
Orn Monatsber 7: 120.

Lectotypes: ZFMK I.II.12.a!-y., female, Bocognano, Cor-
sica, 01.04.1896; ZFMK I.II.12.a!. 6., female, Bocognano,
Corsica, 04.04.1896; ZFMK I.II.12.a!. ¢., male, Bocog-
nano, Corsica, 06.04.1896, all leg. Koenig.

©ZFMK

Types in ZFMK: Section Ornithology 75

Described from six specimens, but no type specimen des-
ignated; ZFMK I.II.12.a!.a. and B. defined as type spec-
imens in the inventory register, but currently untraceable;
one specimen (C) exchanged with the AMNH, New York.
— Dickinson (2003): Serinus serinus (Linnaeus, 1766).

Serinus estherae renatae Schuchmann & Wolters, 1982
Bull Brit Orn Cl 102: 12.

Holotype: ZFMK 81.563, ? male, Rantekombola, Sulawe-
si, 06.08.1980, leg. Krause.

Described from this single specimen. — Dickinson (2003):
Serinus estherae renatae.

Loxia nidificans Kleinschmidt, 1919
Falco 14: 16.

Lectotypes: Coll. KI. 1008, 0, Wladikaukas, Caucasus,
without date; Coll. Kl. 1009, Wladikaukas, Caucasus,
without date;

Type specimens defined subsequently in Catalogue
(Kleinschmidt 1935-1943: 23). — Dickinson (2003): Lox-
ia curvirostra guillemardi, Madarasz, 1903.

EMBERIZIDAE

Emberiza citrinella caliginosa Clancey, 1940
Ibis 14: 94.

Paralectotype: ZFMK 50.106, male, Sutherlandshire,
shotland, 30.08.1938, leg. Clancey.

Holotype in the Coll. Clancey, RSME, Edinburgh, Dur-
noch, E Sutherlandshire, 02.09.1938.

In his original description Clancey did not define materi-
al examined, only the holotype was designated. Paralec-
totype designated by Rheinwald & van den Elzen (1984)
from information on ZFMK inventory. — Dickinson
(2003): Emberiza citrinella caliginosa.

Emberiza cirlus nigrostriata Schiebel, 1910
Orn Jahrb 21: 103.

Syntypes: ZFMK 57.1300 and 57.1298, 2 males,
Ghisonaccia, Corsica, 15. and 16.04.1910, leg. Schiebel.

Bonn zoological Bulletin 59: 29-77

Apparently no further material existed (see also Schifter
& van den Elzen 1986). — Dickinson (2003): Emberiza cir-
lus Linnaeus, 1766.

Fringillaria capensis cloosi Hoesch & Niethammer, 1940
J Ornithol 88, Sonderheft: 335.

Holotype: ZFMK 39.514, male, Brandberg, Namibia,
14.09.1938, leg. Niethammer No 88.

Paratypes: ZFMK 39.513 and 39.515, male, female,
Brandberg, 13. and 12.09.1938, leg. Niethammer.

Described from four specimens, one lost. — Dickinson
(2003): Emberiza capensis bradfieldi (Roberts, 1928).

Emberiza cia africana |e Roi, 1911
Orn Monatsber 19: 79.

Holotype: ZFMK _ 1.1.3.1+.a., male, Lambessa, Algeria,
13.06.1903, leg. Flukiger.

Described from thirteen specimens with no spcifications,
only the holotype was designated in the original descrip-
tion. Of the original material, probably only the holotype
still exists. — Dickinson (2003): Emberiza cia cia Lin-
naeus, 1766.

Emberiza cia flemingorum Martens, 1972
Bonn zool Beitr 23: 116.

Holotype: ZFMK 71.809, female, Barbung Khola, Tarang,
Nepal, 14.06.1970, leg. Martens, No 585.

Paratypes: ZFMK 71.810, male, Dhorpatan, 03.05.1970;
ZFMK 71.811, male, Gompa near Tarakot, 13.05.1970;
ZFMK 71.812, male, Ringmo at Lake Phoksumdo,
29.05.1970; ZFMK 71. 813, male, Ringmo at Lake Phok-
sumdo, 29.05.1970; ZFMK 71.814, male, Ringmo at Lake
Phoksumdo, 03.06.1970; ZFMK 71.815, female, Gompa
near Tarakot, 13.05.1970; ZFMK 71.816, female, Ring-
mo at Lake Phoksumdo, 24.05.1970; ZFMK 71.817, fe-
male, Barbung Khola, Tarang, Nepal, 14.06.1970, all leg.
Martens Nos 577-584. — Dickinson (2003): Emberiza cia

flemingorum.

©ZFMK

76 Renate van den Elzen

Emberiza tschusii witherbyi Jordans, 1923
Falco 19, Sonderheft: 4.

Holotype: ZFMK I.1.4.bb3.v., male, Albufera, Majorca,
13.06.1921, leg. A. von Jordans No 3140.

Paralectotypes: ZFMK I.1.4.bb3.a., male juv., Albufera,
10.06.1921; ZFMK_ 1I.1.4.bb3.B., male juv., Albufera,

13.06.1921; ZFMK I.1.4.bb?-y., female juv., Albufera,
10.06.1921; ZFMK_ 1.1.4.bb3.6., female, Albufera,
18.06.1921; ZFMK_ 1.1.4.bb3.s., female, Albufera,
13.06.1921; ZFMK 1.1.4.bb3.¢., female juv., Albufera,
01.06.1921; ZFMK_ I.1.4.bb3.n., female, Albufera,
13.06.1921; ZFMK_ 1.1.4.bb3.0., male, Albufera,
10.06.1921; ZFMK_ 1[.1.4.bb3.1., male, Albufera,
18.05.1921; ZFMK_ 1.1.4.bb3.x., male, Albufera,
10.06.1921; ZFMK_ 1I.1.4.bb3.4., male, Albufera,
10.06.1921; ZFMK_ 1[.1.4.bb3.., male, Albufera,
10.06.1921; ZFMK_ 1.1.4.bb3.€., male, Albufera,
13.06.1921; ZFMK_ 1I.1.4.bb3.0., male, Albufera,
10.06.1921; ZFMK_ 1.1.4.bb3.2., male, Albufera,
13.06.1921; ZFMK_ 1.1.4.bb3.p.,. male, Albufera,

10.06.1921, all leg. A. von Jordans.

Described from a series of seventeen specimens, two are
missing in the ZFMK collection. In the original descrip-
tion only the holotype was designated by v. Jordans; para-
lectotypes designated by Rheinwald & van den Elzen
(1984) from topotypical material. — Dickinson (2003): Em-
beriza schoeniclus witherbyi.

Emberiza schoeniclus zarudnyi Harms, 1909
Orn Monatsber 17: 1.

Holotype: ZFMK_ [.1.4.bb7.a., Kuldscha at Ili-River,
Turkestan, 01.05. without year, Tancré’s collector, Coll.
Harms No 504, Coll. A. von Jordans No 1364.

Type designated in original description, but no collection
number provided; information on type status taken from
Jordans (1923: 9) and handwritten label. Jordans mentions
a second specimen, which may be a para(lecto)type.—
Dickinson (2003): Emberiza schoeniclus pyrrhuloides Pal-
las, 1811.

Zonotrichia capensis illescasensis Koepcke, 1963
Beitr neotrop Fauna 3: 6.

Paratype: ZFMK 64.3809, female, N Revantazon at Cer-
ro Illescas, N Peru, 20.07.1958, leg. Koepcke No 1568 a.

Holotype at the MNH Javier Prado, Lima, Orn. 1156 (Kp
1568 b), other paratypes at the AMNH, New York and
CNHM, Chicago.

Bonn zoological Bulletin 59: 29-77

Described from a series of three males and three females.
— Dickinson (2003): Zonotrichia capensis illescasensis.

Phrygilus alaudinus humboldti Koepcke, 1963
Beitr neotrop Fauna 3: 11.

Paratype: ZFMK 64.3808, male, Quebrada Avip at Cer-
ro Illescas, N Peru, 23.V07.1958, leg. Koepcke No 1572
b.

Holotype at the MHN Javier Prado, Lima, Orn. 1162 (Kp
1575 d), other paratypes in MHN, Lima, CNHM, Chica-
go and AMNH, New York.

Described from a series of six males and six females. —
Dickinson (2003): Phrygilus alaudinus humboldti.

Sporophila obscura pacifica Koepcke, 1963
Beitr neotrop Fauna 3: 3.

Paratype: ZFMK 56.1055, male, Atiquipa near Chala,
(coast in S Peru), 5.1.1956, leg. Koepcke No 1332 a.

Holotype at the MHN Javier Prado, Lima, Orn 1159 (Kp
1789 n), other paratypes in the collections of MHN and
AMNH, New York.

Described from a series of four males and one juvenile
specimen. — Dickinson (2003): Tiaris obscurus pacificus
(Koepcke, 1963).

Acknowledgements. My thanks go to Carlos Sanchez-Osés for
lending me a helping hand with type material and to Stefanie
Rick for translating the German text and for patient proof read-

ing.

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list of the birds of the world. 34 edition, Christopher Helm,
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Eck S (1970) Von O. Kleinschmidt benannte oder mitbenannte
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(Dresden) 54: 233-316

©OZFMK

Types in ZFMK: Section Ornithology 77

Elzen R van den (1986) Nachtrage und Berichtigungen zum Ty-
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Hartert E (1912) (published 1921). Die Végel der palaarktischen
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Hartert E (1921) (published 1922). Die Végel der palaarktischen
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Hartert E (1923) Die Végel der palaarktischen Fauna: systema-
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Hartert E (1932) (published 1938). Die Végel der palaarktischen
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und der Mittelmeerregion vorkommenden Vogel. Ergaénzun-
gen. Friedlander, Berlin

Hartert E, Kleinschmidt O (1901) Verzeichnis der Brehm’schen
Sammlung. I. Die Formen von Corvus corax. Novitates zoo-
logicae 8: 40-48

ICZN (1999) International Code of Zoological Nomenclature.
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Jordans A von (1923) Uber seltenere und iiber fragliche Vogel-
formen meiner Sammlung. Falco 19, Sonderheft: 8—26

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birds recently described from Turkey, Evemophila alpestris
kumerloevei and Pyrrhula pyrrhula paphlagoniae, with re-
marks on geographical variation in related forms of Bullfinch
from the Balkans and Caucasus. Sandgrouse 28: 12—23

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World. Volume | (2"4 edition). Cambridge, Massachusetts

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van den (1984) Die Wirbeltiersammlungen des Museums Ale-
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Volume 8. Cambridge, Massachusetts

©ZFMK

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Bonn zoological Bulletin 59: 29-77

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OZEMK

Bonn zoological Bulletin Volume 59

pp. 79-108 Bonn, December 2010

A list of the herpetological type specimens
in the Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Wolfgang Bo6hme

Zoologisches Forschungsmuseum Alexander Koenig, Herpetology Section, Adenauerallee 160,
D-53113 Bonn, Germany; E-mail: w.boehme.zfmk@uni-bonn.de.

Abstract. In the herpetological collection of ZFMK 528 scientific species group names are represented by type materi-
al. Of these, 304 names are documented by primary type specimens (onomatophores) while for 224 further names sec-
ondary type specimens (typoids) are available, ranging chronologically from 1801 to 2010. The list is a shortened pred-
ecessor of a comprehensive type catalogue in progress. It lists name bearing types with their catalogue numbers includ-
ing information on further type series members also in other institutions, while secondary types are listed only by pres-
ence, both in ZFMK and other collections including holotype repositories. Geographic origin and currently valid names

are also provided.

Key words. Amphibians and reptiles, type list, ZFMK Bonn.

INTRODUCTION

A first ZFMK herpetological type catalogue was published
(Bohme 1974) three years after I had entered Museum
Koenig as a herpetological curator. It contained only 34
reptilian names documented by type material, 22 of which
were name-bearing type specimens (onomatophores), and
12 further names were documented by paratypes only.
Amphibians were not represented by type material at that
time. A second type catalogue was published ten years lat-
er, on the occasion of the 50‘ anniversary of Museum
Koenig as a public zoological museum (Bohme &
Bischoff 1984). It counted 147 names of amphibians (51)
and reptiles (96) of which 99 were onomatophores while
48 additional names were documented by secondary type
material (paratypes, paralectotypes). In addition, 22
names representing primary and secondary type specimens
of historical material which had belonged before to the Zo-
ological Museum in Gottingen (transferred to ZFMK in
1977) were listed as missing types because they were al-
ready lost before they ever reached ZFMK.

Within the present special BzB issue on type lists of the
vertebrate and invertebrate collections of ZFMK, the pres-
ent herpetological list is a much shortened predecessor of
a comprehensive, critically evaluated type catalogue of
ZFMK amphibians and reptiles which will be published
in the framework of a monograph dealing with the histo-
ry of herpetological research in Bonn since the founda-
tion of the Rheinische Friedrich-Wilhelms-Universitat in
1818 which predates the foundation of ZFMK, particular-
ly the creation of its own herpetological department

Bonn zoological Bulletin 59: 79-108

(currently section) in 1951, for many decades. Nonethe-
less, the present list does comprise some historical “pre-
ZFMK” material which has been obtained after 1971 from
smaller university museums, first of all from the Zoolog-
ical Museum of the University of Géttingen (1977). Sin-
gle taxa documented by type specimens were also received
by the Zoological Museum of the University of Kiel
(1973), and recently by the Biogeography Department of
the University of Saarbriicken (via University of Trier, in
2009).

The following catalogue comprises, with one exception
(the chameleon Kinyongia vanheygeni), only the extant
types while those types which went lost in Gottingen be-
fore the collection transfer to Bonn, and which have been
already mentioned in BOhme & Bischoff (1984), will be
carefully listed and commented on in the forthcoming
monograph mentioned above.

The delimitation and arrangement of families follows Frost
et al. (2006) for the amphibians, and BOhme & Sander
(2010) for the reptiles, plus the familial concept of Zaher
et al. (2009) for colubroid snakes. For dendrobatid frogs,
I follow the monograph by Grant et al. (2006, see also L6t-
ters et al. 2007); for Madagascar, I follow Glaw & Vences
(2007). Within their families, taxa are arranged in alpha-
betical order of genera, and within genera, arrangement
is again alphabetically. Names of authors are accompanied
by an initial letter of their forenames only when a fami-
ly name occurs more than twice.

©ZFMK

80 Wolfgang Bohme

Catalogue numbers are given here for primary types on-
ly, i.e. for holotypes, lectotypes and neotypes; syntypes
are also counted under this category because they are the
first candidates for lectotype selection and are thus like-
ly to contain future lectotypes; for secondary types, 1.e.
paratypes and paralectotypes, only their presence in
ZFMK and/or other collection 1s reported: due to the ne-
cessity to avoid redundancies with the future big cata-
logue. All names are given in their original combinations;
if taxonomic assignment or rank has changed, the current
name believed to be valid today is given. This shortened
preliminary type list thus offers the following informa-
tions:

— All names which are represented at ZFMK by type ma-
terial, in their original taxonomic combinations.

— Primary, name-bearing types and their catalogue num-
bers; also the repositories of additional secondary type
material, if at all existent.

— If only secondary types are involved at ZFMK, the col-
lection housing the respective onomatophore is given,
together with all collections holding additional second-
ary type material.

— The rough geographic origin of the respective taxa.

— The current name combination, if the original combi-
nation has changed.

For more details including full bibliographic references
of each name, for full locality and collector’s data, as well
as for taxonomic and/or historical comments the reader
is referred to the comprehensive monograph in progress.

Abbreviations for museum collections used:

AMNH = American Museum of Natural History, New
York

ANSP = Academy of Natural Sciences, Philadelphia
BEV = Laboratoire de Biogéographie et Ecologie des Ver-
tébrés, Ecole Pratique des Hautes Etudes, Montpellier
BIUB = Mongolian Academy of Sciences, Ulan Bataar
BM(NH) = British Museum (of Natural History), The Nat-
ural History Museum, London

CAS = California Academy of Sciences, San Francisco
CBF = Coleccion Boliviana de Fauna, La Paz

CET = Centro de Estudios Tropicales, Sevilla, Spain
CM = Carnegie Museum, Pittsburgh

CNHM = Chicago Natural History Museum

DZCHU = Department of Zoology, Charles University,
Prague

DZUL = Departamento de Zoologia, Universidad de La
Laguna, Tenerife

EBD = Estacion Biologica de Dofana, Sevilla

EPN = Escuela Politécnica Nacional, Quito

FMNH = Field Museum of Natural History, Chicago

Bonn zoological Bulletin 59: 79-108

GNHM = G6oteborgs Naturhistoriska Museet (see also
NHMG)

HLMD = Hessisches Landesmuseum, Darmstadt

IEBR = Institute of Ecology and Biological Resources,
Hanoi

IFAN = Institut Fondamental d’ Afrique Noire, Dakar
INHS = Illinois Natural History Survey [descended in part
from Illinois State Laboratory of Natural History], Uni-
versity of Illinois, Champaign, Illinois

INPA = Instituto Nacional de Pesquisas da Amazonia, Ma-
naus

IRSNB = Institut Royal des Sciences Naturelles de Bel-
gique, Bruxelles / (= KBIN) Koninklijk Belgisch Instituut
voor Natuurwetenschappen, Brussel

IZUG = Institute of Zoology, University of Genova
LACM = Natural History Museum of Los Angeles
County, Los Angeles

LBUM = Laboratoire de Biogéographie, Université de
Montpellier

LSUZM = Louisiana State University Zoological Museum
MCC = Museo Civico di Storia Naturale, Carmagnola
MCNSC = Museo Insular de Ciencias Naturales de San-
ta Cruz de Tenerife

MCZ = Museum of Comparative Zoology, Cambridge,
Massachusetts

MGL = Musée Guimet d’ Histoire Naturelle, Lyon
MHNCP = Museo de Historia Natural, Universidad Na-
cional de San Antonio Abad del Cusco, Cusco

MHNG = Muséum d’ Histoire Naturelle, Genéve
MHNSM = Museo de Historia Natural San Marcos, Li-
ma

MM = Museum Magdeburg

MNCN = Museo Nacional de Ciencias Naturales, Madrid
MNHN = Muséum National d’ Histoire Naturelle, Paris
MPEG = Museu Paraense “Emilio Goeldi’, Zoologia, Be-
lém, Para

MRAC = Musée Royal de |’ Afrique Centrale, Tervuren
MRSN = Museo Regionale di Scienze Naturali, Torino
MTKD (or MTD respectively) = Museum fir Tierkunde
Dresden

MUSM = Museo de Historia Natural San Marcos

MZB = “Museum Zoologicum Bogoriense” = Zoological
Museum, Bogor

MZUF = Museo Zoologico, Universita di Firenze, Flo-
rence

MZUN = Museo di Zoologia dell’Universita di Napoli,
Naples

MZUSP = Museu de Zoologia da Universidade de Sao
Paulo, Sao Paulo (also as MZSP and MSP)

NHCY = Natural History Collection Yemen, Sana’a
NHMG = Naturhistoriska Riksmuseet, Goteborg (see al-
so GNHM)

NKA = Museo de Historia Natural “Noél Kempff Mer-
cado”, Santa Cruz de la Sierra

NMB = Naturhistorisches Museum Basel

©ZFMK

Type specimens in ZFMK: Section of Herpetology 81

NMBE = Naturhistorisches Museum Bern

NMK = National Museums of Kenya, Nairobi

NMNW = National Museum of Namibia, Windhoek
NMP = National Museum Prague

NMSR = National Museum of South Rhodesia (= now
Zimbabwe), Harare

NMW = Naturhistorisches Museum Wien, Vienna
NUOL = National University of Laos, Vientiane
OMNH = Osaka Museum of Natural History, Laborato-
ry of Zoology, Osaka

PEM = Port Elizabeth Museum

PMNH = Pakistan Museum of Natural History, Islamabad
QCAZ = Museo de Zoologia, Pontifica Universidad
Catolica, Quito

QSMI = Queen Saovabha Memorial Institute, Thai Red
Cross Society, Bangkok

RMNH = Rijksmuseum van Natuurlijke Historie, today
“naturalis”, Leiden

SMF = Senckenberg-Museum Frankfurt/M.

SMNS = Staatliches Museum ftir Naturkunde, Stuttgart
SNHM = Staatliches Naturhistorisches Museum Braun-
schweig

SZE = Sistematik Zooloji Kiirsiisii, Ege Universitesi, Bor-
nova/Izmir

TFMC = Museo Insular de Santa Cruz de Tenerife

TM = Transvaal Museum, Pretoria

UADBA = Université d’ Antananarivo, Dépt. de Biolo-
gie Animale, Antananarivo

UF = Florida Museum of Natural History, Gainesville
UIMNH = University of Illinois Museum of Natural His-
tory, Urbana, Illinois, curated at INHS

ULABG = Coleccion de Anfibios y Reptiles, Laborato-
rio de Biogeografia, Universidad de Los Andes, Mérida
UMMZ = University of Michigan, Museum of Zoology,
Ann Arbor

UNLP = Universidad Nacional de La Plata, National Uni-
versity of La Plata

UNS = University of National Sciences, Ho Chi Min City,
Vietnam

URP = Museo de Historia Natural, Universidad Ricardo
Palma, Lima

USNM = United States National Museum, Washington
UTA = University of Texas at Arlington

UZMK = (see ZMUC)

VNMN = Vietnam National Museum of Nature, Hanoi
VNUH = Zoological Museum of the Vietnam National
University, Hanoi

ZIH = Zoological Institute, University of Halle

ZIK = Zoological Institute, Ukrainian Academy of Sci-
ences, Kiev (also as ZMUAS and IZANU)

ZIL = ZISP = Zoological Institute, Russian Academy of
Sciences, Leningrad =

ZISP = Zoological Institute, Russian Academy of Sci-
ences, St. Petersburg

ZIT = Zoological Institute, Grusinian Academy of Sci-

Bonn zoological Bulletin 59: 79-108

ences, Tbilisi

ZLU = Departamento de Zoologia de la Universidad de
La Laguna, Tenerife

ZMA = Zodlogisch Museum, Universiteit van Amsterdam
ZMB = Zoologisches Museum = Museum fiir Naturkun-
de, Berlin

ZMH = Zoologisches Museum der Universitat Hamburg
ZMMU = Zoological Museum, Moscow State Universi-

ZMUC = Zoological Museum, University of Copenhagen
ZRC = Zoological Reference Collection, National Univer-
sity of Singapore

ZSM = Zoologische Staatssammlung Minchen

LIST OF EXTANT TYPE SPECIMENS

AMPHIBIA

GYMNOPHIONA
Caeciliidae

Caecilia kaupii Berthold, 1859
Holotype: ZFMK 27684, no paratypes
Origin: Venezuela

Current name Potomotyphlops kaupii.

Oscaecilia koepckeorum M. Wake, 1984
Holotype: ZFMK 23392, no paratypes
Origin: Peru.

URODELA
Salamandridae

Cynops shataukokensis Freytag & Eberhard, 1977
Paratypes in ZFMK, Holotype SMF,

Origin: “China”

Current name: Cynops pyrrhogaster.

Mertensiella luschani antalyana Basoglu & Baran,
1976

Paratype in ZFMK, further paratypes and holotype in SZE,
Origin: SW Turkey

Current name: Lyciasalamandra antalyana.

Mertensiella luschani atifi Basoglu, 1967
Paratypes in ZFMK and in SZE, Holotype in SZE
Origin: SW Turkey

Current name: Lyciasalamandra atifi.

©ZFMK

82 Wolfgang Bohme

Mertensiella luschani basoglui Baran & Atatir, 1980
Paratypes in ZFMK amnd SZE, Holotype in SZE
Origin: SW Turkey

Current name: Lyciasalamandra luschani basoglui.

Mertensiella luschani billae Franzen & Klewen, 1987
Holotype: ZFMK 45766, paratypes in ZFMK and ZSM
Origin: SW Turkey

Current name: Lyciasalamandra billae.

Mertensiella luschani fazilae Basoglu & Atattr, 1974
Paratypes in ZFMK and SZE, Holotype in SZE
Origin: SW Turkey

Current name: Lyciasalamandra fazilae.

Mertensiella luschani finikensis Basoglu & Atatiir,
1975

Paratypes in ZFMK and SZE, Holotype in SZE

Origin: SW Turkey

Current name: Lyciasalamandra luschani finikensis.

Mertensiella luschani helverseni Pieper, 1963
Paratypes in ZFMK and SMF, Holotype in SMF
Origin: Karpathos Id., Greece

Current name: Lyciasalamandra helverseni.

Paramesotriton caudomaculatus Seidel, 1981
Lectotype: ZFMK 38536, paralectotypes in ZFMK
Origin: China

Current name: Paramesotriton caudopunctatus.

Salamandra algira tingitana Donaire-Barroso & Bo-
gaerts, 2003

Paratypes in ZFMK, Holotype in MNCN

Origin: N Morocco

Salamandra salamandra alfredschmidti G. Kohler &
Steinfartz, 2006

Paratypes in ZFMK, SMF and SNHM

Origin: Asturias, Spain

Salamandra salamandra gigliolii Eiselt & Lanza, 1956
Paratypes in ZFMK, NMW, MM anfd MZUF, Holotype
in MZUF

Origin: Italy

Salamandra salamandra (gallaica) crespoi Malkmus,
1983

Holotype: ZFMK 36842, Paratypes in MTKD

Origin: S Portugal

Current name: Salamandra salamandra crespoi.

Salamandra salamandra longirostris Joger & Steinfartz,
1994
Paratypes in ZFMK and MM, Holotype in HLMD

Bonn zoological Bulletin 59: 79-108

Origin: S Spain
Current name: Salamandra longirostris.

Triton ophryticus Berthold, 1846
Lectotype: ZFMK 27793, paralectotype lost
Origin: Georgia

Current name: Ommatotriton ophryticus.

Triturus vulgaris kosswigi Freytag, 1955

Paratypes in ZFMK, MM and ZISP, Holotype in MM
Origin: N Turkey

Current name: Lissotriton vulgaris kosswigi.

Tylototriton vietnamensis Bohme, Schottler, Nguyen Q.T.
& J. Kohler, 2005

Holotype: ZFMK 80637, paratypes in ZFMK

Origin: Vietnam

Plethodontidae

Oedipina uniformis Keferstein, 1868
Holotype: ZFMK 27833, no paratypes
Origin: Costa Rica

SALIENTIA
Megophryidae

Leptolalax pictus Malkmus, 1992
Holotype: ZFMK 51762, paratypes in ZMB
Origin: Sabah, Malaysia

Limnodynastidae

Limnodynastes peronii var. rugulosus Keferstein, 1867
Syntypes: ZFMK 28332-337

Origin: NSW, Australia

Current name: Limnodynastes tasmaniensis

Platyplectrum superciliare Keferstein, 1867
Holotype: ZFMK 28331, no paratypes
Origin: “Australia”

Current name: Limnodynastes dorsalis

Myobatrachidae
Crinia georgiana var. laevipes Keferstein, 1867
Syntypes: ZFMK 28246—247

Origin: NSW, Australia
Current name: Crinia signifera

OZFMK

Type specimens in ZFMK: Section of Herpetology 83

Pseudophryne coriacea Keferstein, 1867
Syntypes: ZFMK 38201—207
Origin: NSW, Australia

Uperoleia marmorata var. laevigata Keferstein, 1867
Lectotype: ZFMK 26309, paralectotypes in ZFMK
Origin: NSW, Australia

Current name: Uperoleia laevigata

Pelodrydadiae

Hyla dentata Keferstein, 1868
Holotype: ZFMK 28808, no paratypes
Origin NSW, Australia

Current name: Litoria dentata

Ayla (Litoria) freycineti var. unicolor Keferstein, 1867
Syntypes: ZFMK 28788—789

Origin: “Australien”

Current name: Litoria freycineti

Hyla (Litoria) mystacina Keferstein, 1867
Holotype: ZFMK 28810, no paratypes
Origin: “Australien”

Current name: Litoria mystacina

Ayla schuettei Keferstein, 1868
Holotype: ZFMK 28809, no paratypes
Origin: “Sydney”, Australia

Current name: Litoria adelaidensis

Amphignathodontidae

Gastrotheca piperata Duellman & J. Kohler, 2005
Holotype: ZFMK 66838, paratype in ZFMK
Origin: Bolivia

Hylidae

Argenteohyla altamazonica Henle, 1981
Holotype: ZFMK 29993, paratype in ZFMK
Origin: Peru

Current name: Trachycephalus venulosus

Dendropsophus salli Jungfer, Reichle & Piskurek, 2010
Paratypes in ZFMK, MNK and SMNS, holotype in NVK
Origin: Bolivia.

Hyla coffea J. Kohler, Jungfer & Reichle, 2005
Paratypes in ZFMK and CBF, holotype in CBF
Origin: Bolivia

Current name: Dendropsophus coffea

Bonn zoological Bulletin 59: 79-108

Hyla creolica Werner, 1899

Syntypes: ZFMK 28569-583

Origin: Colombia

Current name: Dendropsophus labialis

Hyla delarivai J. Kohler & Lotters, 2001
Paratypes in ZFMK and HLMD, Holotype in CBF
Origin: Bolivia

Current name: Dendropsophus delarivai

Hyla elkejungingerae Henle, 1981

Holotype: ZFMK 33352, paratypes in ZFMK
Origin: Peru

Current name: Osteocephalus mimeticus

Ayla felixarabica Gvozdik, Moravec, Kliitsch & Kotlik,
2010

Paratypes in ZFMK, ZMH and NMP, holotype in NMP
Origin: Yemen

Ayla joannae J. Kohler & Lotters, 2001

Paratypes in ZFMK, CBF, KU and HLMD, holotype in
CBF

Origin: Bolivia

Current name: Dendropsophus joannae

Hyla lineomaculata Werner, 1899
Holotype: ZFMK 28585, no paratypes
Origin: Trinidad

Current name: Scinax ruber

Ayla servalina Werner, 1899
Holotype: ZFMK 28584, no paratypes
Origin: Colombia

Current name: Dendropsophus labialis

Hypsiboas nympha Faivovich, Moravec, Cisneros-Here-
dia & J. Kohler, 2006

Paratypes in ZFMK, AMNH, KU, DFCH-USFQ, QCAZ
and NMP, holotype in DFCH-USFQ

Origin: Ecuador

Ololygon pedromedinae Henle, 1991
Holotype: ZFMK39737, no paratypes
Origin: Peru

Current name: Scinax pedromedinae

Osteocephalus deridens Jungfer, Ron, Seipp & Almen-
dariz, 2000

Paratypes in ZFMK, SMF, EPN and QCAZ, holotype in
EPN

Origin: Ecuador

©ZFMK

84 Wolfgang Bohme

Osteocephalus fuscifacies Jungfer, Ron, Seipp & Almen-
darez, 2000

Paratypes in ZFMK, EPN, MHNG and QCAZ, holotype
in EPN

Origin: Ecuador

Osteocephalus mutabor Jungfer & Hédl, 2002
Paratypes in ZFMK, EPN, MHNG and SMF, holotype in
EPN

Origin: Ecuador

Osteocephalus oophagus Jungfer & Schiesari, 1995
Paratypes in ZFMK, NMW, MPEG, AMNH, MZUSP, IN-
PA and SMNS

Origin: Amazonian Brazil

Phyllomedusa camba De \a Riva, 2000

Paratypes in ZFMK, KU, NKA and USNM, holotype in
NKA

Origin: Bolivia

Phyllomedusa iheringii Boulenger, 1885
Syntype: ZFMK 28560, further syntypes in BMNH
Origin: Rio Grande do Sul, Brazil

Centrolenidae

Hyalinobatrachium carlesvilai Castroviejo-Fisher, Padi-
al, Chaparro, Aguayo & De la Riva, 2009

Paratypes in ZFMK, MHNCP and MNCN, Holotype in
MNCN

Origin: Bolivia

Strabomantidae

Eleutherodactylus ashkapara J. Kohler, 2000
Paratype in ZFMK, holotype in CBF

Origin: Bolivia

Current name: Yunganastes ashkapara

Eleutherodactylus ibischi Reichle, Lotters & De la Riva,
2001

Paratype in ZFMK, holotype in CBF

Origin: Bolivia

Current name: Oreobates ibischi

Eleutherodactylus llojsintuta J. K6hler & Lotters, 1999
Paratypes in ZFMK, CBF and NKA, holotype in CBF
Origin: Bolivia

Current name: Pristimantis llojsintuta

Eleutherodactylus nebulosus Henle, 1992
Holotype: ZFMK 27634, no paratypes

Bonn zoological Bulletin 59: 79-108

Origin: Peru
Current (substitute) name: Pristimantis caliginosus

Eleutherodactylus olivaceus J. Kohler, Morales, Lotters,
Reichle & Aparicio, 1998

Paratypes in ZFMK, CBF, MUSM and USNM

Origin: Bolivia

Current name: Pristimantis olivaceus

Eleutherodactylus samaipatae J. Kohler & Jungfer, 1995
Holotype: ZFMK 59600, no paratypes

Origin: Bolivia

Current name: Pristimantis samaipatae

Eleutherodactylus pluvicanorus De la Riva & Lynch,
1997

Paratypes in ZFMK, CET, UTA and SMF, holotype in
NKA

Origin: Bolivia

Current name: Yunganastes pluvicanorus

Ischnocnema sanderi Padial, Reichle & De la Riva, 2005
Paratypes in ZFMK, CBF, MNCN and NKA, holotype in
CBF

Origin: Bolivia

Current name: Oreobates sanderi

Phyllonastes carrascoicola De la Riva & J. Kohler, 1998
Holotype: ZFMK 59573, paratypes in ZFMK

Origin: Bolivia

Current name: Noblella carrascoicola

Pristimantis koehleri Padial & De la Riva, 2009
Paratypes in ZFMK, MNCN and NMK, holotype in
MNCN

Origin: Bolivia

Pristimantis reichlei Padial & De la riva, 2009
Paratypes in ZFMK, MNCN and NMK, holotype in NUK
Origin: Bolivia

Pristimantis yuruanensis Rodder & Jungfer, 2008
Holotype: ZFMK 87278, paratypes in SMNS and ULA-
BG

Origin: Venezuela

Leptodactylidae
Adenomera griseigularis Henle, 1981
Holotype: ZFMK 31800, no paratypes

Origin: Peru
Current name: Leptodactylus griseonotus

©ZFMK

Type specimens in ZFMK: Section of Herpetology 85

Leptodactylus raniformis Werner, 1899
Holotype: ZFMK 80600, no paratypes
Origin: Colombia

Leptodactylus gracilis delattini P. Miller, 1968
Holotype: ZFMK (recently acquired, to be catalogued
soon)

Origin: S Brazil

Ceratophryidae

Telmatobius hockingi Salas & Sinsch, 1996
Paratypes in ZFMK and URP, holotype in URP
Origin: Peru

Telmatobius mayoloi Salas & Sinsch, 1996

Paratypes in ZFMK, URP, MHNSM and KU, holotype in
URP

Origin: Bolivia

Cyclorhamphidae

Odontophrynus cordobae Martino & Sinsch, 2002
Holotype: ZFMK 73460, transferred to UNLP, paratypes
in ZFMK

Origin: Argentina

Dendrobatidae

Ameerega boehmei Lotters, Schmitz, Reichle, Rodder &
Quennet, 2009

Paratypes in ZFMK, CAS, CM, LACM and NKA, Holo-
type in NKA

Origin: Bolivia

Dendrobates claudiae Jungfer, Lotters & Jérgens, 2000
Holotype:FMK 73561, paratypes in ZFMK

Origin: Panama

Current name: Ranitomeya claudiae

Dendrobates histrionicus Berthold, 1845

Lectotype: ZFMK 28123, paralectotypes in ZFMK and
AMNH

Origin: Colombia

Current name: Oophaga histrionica

Dendrobates lehmanni Myers & Daly, 1976
Paratypes in ZFMK and AMNH

Origin: Colombia

Current name: Oophaga lehmanni

Bonn zoological Bulletin 59: 79-108

Dendrobates nubeculosus Jungfer & Bohme, 2004
Holotype: ZFMK45354, no paratypes
Origin: Guyana

Dendrobates typographus Keferstein, 1867
Holotype: 28115, no paratypes

Origin: Costa Rica

Current name: Oophagus pumilio

Dendrobates variabilis Zimmermann & Zimmermann,
1988

Paratypes in ZFMK, holotype in SMNS

Origin: Peru

Current name: Ranitomeya variabilis

Dendrobates vicentei Jungfer, Weygoldt & Juraske, 1996
Holotype: ZFMK 61100, paratypes in ZFMK and SMNS
Origin: Panama

Current name: Oophaga vicentei

Epipedobates ardens Jungter, 1989
Holotype: ZFMK 49084, paratype in ZFMK
Origin: Peru

Current name: Ameerega cainarachi

Epipedobates bilinguis Jungfer, 1989
Holotype: ZFMK 49073, paratypes in ZFMK
Origin: Ecuador

Current name: Amereega bilinguis

Epipedobates rubriventris Lotters, Debold, Henle, Glaw
& Kneller, 1997

Holotype: ZFMK 64838, paratypes in ZFMK and ZSM
Origin: Peru

Current name: Ameerega rubriventris

Phyllobates azureiventris Kneller & Henle, 1985
Holotype: ZFMK 41507, paratypes in ZFMK
Origin: Peru

Current name: Hyloxalus azureiventris

Phyllobates melanorrhinus Berthold, 1845

Lectotype: ZFMK 28139, paralectotypes in ZFMK and
AMNH

Origin: Colombia

Current name: Phyllobates bicolor

Phyllobates terribilis Myers, Daly & Malkin, 1978
Paratype in ZFMK and AMNH, Holotype in AMNH
Origin: Colombia

Phyllobates (Hypodiction) palmatus Werner, 1899
Holotype: ZFMK 28132, no paratypes

Origin: Colombia

Current name: Rheobates palmatus

©ZFMK

86 Wolfgang Bohme

Prostherapis variabilis Werner, 1899
Syntypes: ZFMK 28134—-138

Origin: Colombia

Current name: Colostethus subpunctatus

Bufonidae

Atelopus cruciger vogli L. Miller, 1935
Paratypes in ZFMK and ZSM, holotype in ZSM
Origin: Venezuela

Atelopus flaviventris Werner 1899
Holotype: ZFMK 28107, no paratypes
Origin: Colombia

Current name: Atelopus ignescens

Atelopus reticulatus Lotters, Haas, Schick & Bohme,
2002

Holotype: ZFMK 76247, further paratype in ZFMK
Origin: Colombia

Atelopus subornatus Werner, 1899
Lectotype: ZFMK 28104, paralectotypes in ZFMK
Origin: Colombia

Atelopus tricolor Boulenger, 1902
Lectotype: ZFMK 28103, paralectotypes in BMNH
Origin: SE Peru

Bufo ehlersi Werner, 1899

Holotype: ZFMK 21414, no paratypes
Origin: Colombia

Current name: /ncilius coniferus

Bufo pentoni tihamicus Balletto & Cherchi, 1973
Paratypes in ZFMK and IZUG, Holotype in IZUG
Origin: Yemen

Current name: Amietophrynus tihamicus

Bufo pithecodactylus Werner, 1899
Holotype: ZFMK 27999, no paratypes
Origin: Colombia

Current name: Rhinella marina

Bufo pseudoraddei baturae Stéck, Schmid, Steinlein &
Grosse, 1999

Paratype in ZFMK, holotype in ZSM

Origin: N Pakistan

Bufo scorteccii Balletto & Cherchi, 1970

Paratypes in ZFMK and IZUG, Holotype in IZUG
Origin: Yemen

Bonn zoological Bulletin 59: 79-108

Bufo siculus Stéck, Sicilia, Belfiore, Buckley, Lo Brut-
to, Lo Valvo & Arculeo, 2008

Paratypes in ZFMK, ZMB, MVZ, MZPA and NME, holo-
type in Museum of Terrasini, Palermo

Origin: Sicily, Italy

Bufo stanlaii Lotters & Kohler, 2000

Paratypes in ZFMK, USNM and ZSM, Holotype in CBF
Origin: Bolivia

Current name: Rhinella stanlaii

Bufo verrucosissimus circassicus Orlova & Tuniyev,
1989

Paratype in ZFMK, Holotype in ZMM

Origin: Russia

Bufo viridis turanensis Hemmer, Schmidtler & Bohme,
1978

Paratypes in ZFMK, MTKD, ZSM and ZISP, Holotype in
MTKD

Origin: Tadjikistan

Nectophrynoides mirei Perret, 1971

Paratypes in ZFMK and MHNG, Holotype in MHNG
Origin: Cameroon

Current name: Wolterstorffina mirei

Werneria ibundji Rodel, Schmitz, Pauwels & Bohme,
2004

Paratype in ZFMK, Holotype in IRSNB

Origin: Gabon

Werneria submontana
Holotype: ZFMK 69999, paratypes in ZFMK
Origin: Cameroon

Microhylidae

Anodonthyla emilei Vences, Glaw, J. Kohler & Wollen-
berg, 2010

Paratypes in ZFMK and ZSM, holotype in ZSM

Origin: Madagascar

Anodonthyla jeanbai Vences, Glaw, J. Kohler & Wollen-
berg, 2010

Paratypes in ZFMK and ZSM. holotype in ZSM

Origin: Madagascar

Anodonthyla moramora Glaw & Vences, 2005
Paratypes in ZFMK, ZSM, ZMA and UADBA, holotype
in ZSM

Origin: Madagascar

©ZFMK

Type specimens in ZFMK: Section of Herpetology 87

Anodonthyla nigrigularis Glaw & Vences, 1992
Holotype: ZFMK 53745, paratypes in ZFMK and ZSM
Origin: Madagascar

Anodonthyla theoi Vences, Glaw, J. Kéhler & Wollenberg,
2010

Paratypes in ZFMK and ZSM, holotype in ZSM

Origin: Madagascar

Hypopachus seebachi Keferstein, 1867
Syntypes: 28389-392

Origin: Costa Rica

Current name: Hypopachus variolosus

Microhyla marmorata Bain & Nguyen Q.T., 2004
Paratypes in ZFMK, AMNH and FMNH, holotype in
IEBR

Origin: Vietnam

Pachybatrachus petersii Keferstein, 1868
Holotype: ZFMK 28388, no paratypes
Origin: “Neu-Siid-Wales, Australien”
Current name: Uperodon systoma

Platypelis occultans Glaw & Vences, 1992

Holotype: ZFMK 53735, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Cophyla occultans

Plethodontohyla coronata Vences & Glaw, 2003
Holotype: ZFMK 57459, paratypes in ZSM and VADBA
Origin: Madagascar

Current name: Rhombophryne coronata

Plethodontohyla mikanika Vences, Raxworthy, Nuss-
baum & Glaw, 2003

Paratypes in ZFMK, ZSM, UMMZ and MNHN, holoty-
pe in UMMZ

Origin: Madagascar:

Rhombophryne mangabensis Glaw, J. Kohler & Vences,
2010

Paratypes in ZFMK, ZSM and UADBA, holotype in ZSM
Origin: Madagascar

Scaphiophryne boribory Vences, Raxworthy, Nussbaum
& Glaw, 2003

Paratypes in ZFMK, MRSN, UMMZ and ZSM, holoty-
pe in UMMZ

Origin: Madagascar

Scaphiophryne gottlebei Busse & Bohme, 1992

Holotype: ZFMK 53543, paratypes in ZFMK and ZSM
Origin: Madagascar

Bonn zoological Bulletin 59: 79-108

Stumpffia gimmeli Glaw & Vences, 1992
Holotype: ZFMK 52541, paratypes in ZFMK and ZSM
Origin: Madagascar

Stumpffia pygmaea Vences & Glaw, 1991
Holotype: ZFMK 52541, paratypes in ZFMK and ZSM
Origin: Madagascar

Stumpffia tetradactyla Vences & Glaw, 1991
Holotype: ZFMK 52547, paratypes in ZFMK and ZSM
Origin: Madagascar

Hyperoliidae

Heterixalus mandrakata Glaw & Vences, 1991
Holotype: ZFMK 52557, paratypes in ZFMK and ZSM
Origin: Madagascar

Hyperolius dintelmanni Lotters & Schmitz, 2004
Holotype: 67871, paratypes in ZFMK
Origin: Cameroon

Hyperolius nienokouensis Rodel, 1998

Holotype: ZFMK 65392, paratypes in ZFMK, ZMUC and
SMNS

Origin: Ivory Coast

Hyperolius veithi Schick, Kielgast, Rodder, Muchau, Bur-
ger & Lotters, 2010
Holotype: ZFMK 89607, paratypes in ZFMK and ZMUC
Origin: D.R. Congo

Kassina schioetzi Rodel, Grafe, Rudolf & Ernst, 2002
Paratype in ZFMK, Holotype in SMNS
Origin: Ivory Coast

Morerella cyanophthalma Roédel, Assemian, Kouamé, To-
hé & Perret, 2009

Paratypes in ZFMK, MHNG, SMNS and ZMB, holotype
in MHNG

Origin: Ivory Coast

Arthroleptidae

Cardioglossa alsco H.W. Herrmann, P. Herrman, Schmitz
& Bohme, 2004

Holotype: ZFMK 77678, paratypes in ZFMK

Origin: Cameroon

Cardioglossa gratiosa peternageli Bohme & B. Schnei-
der, 1987

Holotype: ZFMK 40805, paratypes in ZFMK

Origin: Cameroon

©ZFMK

88 Wolfgang Bohme

Leptodactylodon wildi Amit & Dowsett-Lemaire, 2000
Paratypes in ZFMK, MNHN and BMNH, Holotype in
MNHN

Origin: Cameroon

Leptopelis crystallinoron Lotters, Rodel & Burger, 2005
Holotype 73139, no paratypes
Origin: Gabon

Leptopelis mackayi J. Kohler, Bwong, Schick, Veith &
Lotters, 2006

Paratypes in ZFMK and NMK, holotype in NMK
Origin: W Kenya

Leptopelis spiritusnoctis Rodel, 2007

Paratypes IN ZFMK, ZMB, SMNS, PEM and MOR, ho-
lotype in ZMB

Origin: SE Guinea

Phrynobatrachus nlonakoensis

Holotype: ZFMK 80970, paratypes in ZFMK
Origin: Cameroon

Current name: Arthroleptis nlonakoensis

Ptychadenidae

Ptychadena arnei Perret, 1997
Paratypes in ZFMK and ZMUC, Holotype in ZMUC
Origin: Senegal

Phrynobatrachidae

Phrynobatrachus kakamikro Schick, Zimkus, Channing,
J. Kohler & Loétters, 2010

Paratypes in ZFMK, BMNH and NKM, Holotype in NKM
Origin: W Kenya

Phrynobatrachus phyllophilus Rodel & Ernst, 2002
Paratypes in ZFMK, ZSM, ZMUC and SMNS, Holotype
in SMNS

Origin: Ivory Coast

Petropedetidae

Petropedetes euskircheni Barej, R6del, Gonwouo,
Pauwels, Bohme & Schmitz, 2010

Holotype: ZFMK 75582, paratypes in ZFMK, IRSNB and
ZMB

Origin: Cameroon

Bonn zoological Bulletin 59: 79-108

Petropedetes juliawurstnerae Barej, R6del, Gonwouo,
Pauwels, Bohme & Schmitz, 2010

Holotype: 67360, paratypes in ZFMK, ZMB and IRSNB
Origin: Cameroon

Petropedetes vulpiae Barej, R6del, Gonwouo, Pauwels,
Bohme & Schmitz, 2010

Holotype: ZFMK 81623, paratypes in ZFMK, IRSNB,
MHNG and ZMB

Origin: Cameroon

Dicroglossidae

Fejervarya iskandari Veith, Kosuch, Ohler & Dubois,
2001

Paratypes in ZFMK and FMNH, Holotype in MNHN
Origin: Java, Indonesia

Mantellidae

Aglyptodactylus laticeps Glaw, Vences & Bohme, 1998
Holotype: ZFMK 59972, paratypes in ZFMK and ZSM
Origin: Madagascar

Aglyptodactylus securifer Glaw, Vences & Béhme, 1998
Holotype: ZFMK 59975, paratype in ZSM
Origin: Madagascar

Blommersia galani Vences, J. Kohler, Pabijan & Glaw,
2010

Paratypes in ZFMK, ZSM and ZMA

Origin: Madagascar

Boophis albilabris occidentalis Glaw & Vences, 1994
Holotype: ZFMK 57383, paratype in ZSM
Origin: Madagascar

Boophis albipunctatus Glaw & Thiesmeier, 1993
Holotype: ZFMK 53617, no paratype
Origin: Madagascar

Boophis albipunctatus sibilans Glaw & Thiesmeier, 1993
Holotype: ZFMK 53618, paratype in ZSM

Origin. Madagascar

Current name: Boophis sibilans

Boophis andreonei Glaw & Vences, 1994)
Holotype: ZFMK 57393, paratypes in ZFMK and ZSM
Origin: Madagascar

Boophis blommersae Glaw & Vences, 1995

Holotype: ZFMK 57398, paratypes in ZFMK and ZSM
Origin: Madagascar

©ZFMK

Type specimens in ZFMK: Section of Herpetology 89

Boophis boehmei Glaw & Vences, 1992
Holotype: ZFMK 53642, paratypes in ZFMK and ZSM
Origin: Madagascar

Boophis bottae Vences & Glaw, 2002
Paratypes in ZFMK, holotype in ZSM
Origin: Madagascar

Boophis burgeri Glaw & Vences, 1994
Holotype: ZFMK 53641, paratypes in ZFMK an ZSM
Origin: Madagascar

Boophis englaenderi Glaw & Vences, 1994
Holotype: ZFMK 57390, paratypes in ZFMK an ZSM
Origin: Madagascar

Boophis entingae Glaw, J. Kohler, De la Riva, Vietes &
Vences, 2010

Paratypes in ZFMK and ZSM, holotype in ZSM

Origin: Madagascar

Boophis feonnyala Glaw, Vences, Andreone & Vallan,
2001

Paratypes in ZFMK, Holotype in ZSM

Origin: Madagascar

Boophis haematopus Glaw, Vences, Andreone & Vallan,
2001

Paratypes in ZFMK and MRSN, Holotype in ZSM
Origin: Madagascar

Boophis haingana Glaw, J. Kohler, De la Riva & Vieites,
2010

Paratypes in ZFMK, ZSM and UADBA, Holotype in ZSM
Origin: Madagascar

Boophis jaegeri Glaw & Vences, 1992
Holotype: ZFMK 52569, no paratypes
Origin: Madagascar

Boophis liami Vallan, Vences & Glaw, 2003

Holotype: ZFMK 60123, paratypes in ZFMK, NMBE,
ZSM and UADBA

Origin: Madagascar

Boophis luciae Glaw, J. Kohler, De la Riva & Vieites,
2010

Paratypes in ZFMK, ZMA, ZSM and UADBA, Holotype
in ZSM

Origin: Madagascar

Boophis luteus septentrionalis Glaw & Vences, 1994
Holotype: ZFMK 57386, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Boophis septentrionalis

Bonn zoological Bulletin 59: 79-108

Boophis marojezensis Glaw & Vences, 1994
Holotype: ZFMK 57401, paratype in ZSM
Origin: Madagascar

Boophis picturatus Glaw, Vences, Andreone & Vallan,
2001

Paratypes in ZFMK, Holotype in ZSM

Origin: Madagascar

Boophis piperatus Glaw, J. Kohler, De la Riva, Vieites
& Vences, 2010

Paratypes in ZFMK, UADBA and ZSM, Holotype in ZSM
Origin: Madagascar

Boophis pyrrhus Glaw, Vences, Andreone & Vallan, 2001
Paratypes in ZFMK, ZMA and MRSN, Holotype in ZSM
Origin: Madagascar

Boophis roseipalmatus Glaw, J. Kohler, De la Riva,
Vieites & Vences, 2010

Paratypes in ZFMK and ZSM, Holotype in ZSM
Origin: Madagascar

Boophis rufioculis Glaw & Vences, 1997
Holotype: ZFMK 60079, Paratypes in ZFMK and ZSM
Origin: Madagascar

Boophis sandrae Glaw, J. Kohler, De la Riva, Vieites &
Vences, 2010

Paratypes in ZFMK, ZMA and ZSM, Holotype in ZMA
Origin: Madagascar

Boophis schuboeae Glaw & Vences, 2002
Holotype: ZFMK 62299, paratypes in ZFMK and ZSM
Origin: Madagascar

Boophis tampoka J. Kohler, Glaw & Vences, 2007
Paratypes in ZFMK, HLMD, FGZC, UADBA and ZSM,
Holotype in ZSM

Origin: Madagascar

Boophis tasymena Vences & Glaw, 2002
Paratypes in ZFMK, Holotyepe in ZSM
Origin: Madagascar

Boophis xerophilus Glaw & Vences, 1997
Holotype: ZFMK 59987, paratypes in ZFMK and ZSM
Origin: Madagascar

Mantella aurantiaca rubra Staniszewski, 1996
Lectotype: ZFMK 68868, paralectotypes
Staniszewsk1) lost

Origin: Madagascar

Current name: Mantella aurantiaca

(coll.

©OZFMK

90 Wolfgang Bohme

Mantella bernhardi Vences, Glaw, Peyrieras, Bohme &
Busse, 1994

Holotype: ZFMK 57164, paratypes in ZFMK

Origin: Madagascar

Mantella crocea Pintak & Bohme, 1990
Holotype: ZFMK 45007, paratypes in ZFMK and ZSM
Origin: Madagascar

Mantella expectata Busse & Bohme, 1992
Holotype: ZFMK 53540, paratypes in ZFMK
Origin: Madagascar

Mantella madagascariensis haraldmeieri Busse, 1981
Holotype: ZFMK 25351, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Mantella haraldmeieri

Mantella viridis Pintak & Bohme, 1988
Holotype: ZFMK 47900, no paratypes
Origin: Madagascar

Mantidactylus ambohitra Vences & Glaw, 2001
Paratypes in ZFMK, MNHN and MTKD, holotype in
ZSM

Origin: Madagascar

Current name: Gephyromantis (Duboimantis) ambohitra

Mantidactylus charlotteae Vences & Glaw, 2004
Paratypes in ZFMK and ZSM, Holotype in ZMA
Origin: Madagascar

Current name: Mantidactylus (Chonomantis) charlotteae

Mantidactylus cornutus

Holotype: ZFMK 53691, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Gephyromantis (Duboimantis) cornutus

Mantidactylus corvus Glaw & Vences, 1992

Holotype: ZFMK 57430, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Gephyromantis (Phylacomantis) corvus

Mantidactylus enki Glaw & Vences, 2002

Holotype: ZFMK 62293, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Gephyromantis enki

Mantidactylus fimbriatus Glaw & Vences, 1994
Holotype: ZFMK 57441, paratype in ZSM
Origin: Madagascar

Current name: Spinomantis fimbriatus

Bonn zoological Bulletin 59: 79-108

Mantidactylus kathrinae Glaw, Vences & Gossmann,
2000

Holotype: ZFMK 62264, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Guibemantis kathrinae

Mantidactylus kely Glaw & Vences, 1994
Holotype: ZFMK 57444, no paratypes
Origin: Madagascar

Current name: Blommersia kely

Mantidactylus madinika Vences, Andreone, Glaw & Mat-
tioli, 2002

Paratypes in ZFMK, MRSN and ZSM, Holotype in ZSM
Origin: Madagascar

Current name: Wakea madinika

Mantidactylus massi Glaw & Vences, 1994
Holotype: ZFMK 57442, paratype in ZSM
Origin: Madagascar

Current name: Spinomantis massi

Mantidactylus moseri Glaw & Vences, 1997
Paratypes in ZFMK, holotype in ZSM

Origin: Madagascar

Current name: Gephyromantis (Duboimantis) moseri

Mantidactylus phantasticus Glaw & Vences, 1997
Holotype: ZFMK 60000, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Spinomantis phantasticus

Mantidactylus rivicola Vences, Glaw & Andreone, 1997
Holotype: ZFMK 57428, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Gephyromantis (Vatomantis) rivicola

Mantidactylus sarotra Glaw & Vences, 2002

Paratypes in ZFMK, UADBA-FG and ZSM, holotype in
ZSM

Origin: Madagascar

Current name: Blommersia sarotra

Mantidactylus schilfi Glaw & Vences, 2000

Holotype: ZFMK 59885, paratypes in ZSM and MNHN
Origin: Madagascar

Current name: Gephyromantis (Duboimantis) schilfi

Mantidactylus (Laurentomantis) striatus Vences, Glaw,
Andreone, Jesu & Schimmenti, 2002

Paratypes in ZFMK and FACZ (MRSN), holotype in ZSM
Origin: Madagascar

Current name: Gephyromantis (Laurentomantis) striatus

©ZFMK

Type specimens in ZFMK: Section of Herpetology 9]

Mantidactylus tandroka Glaw & Vences, 2001
Paratypes in ZFNK, ZSM and MNHN, holotype in
MNHN

Origin: Madagascar

Current name: Gephyromantis (Duboimantis) tandroka

Mantidactylus thelenae Glaw & Vences, 1994
Holotype: 57422, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Gephyromantis thelenae

Mantidactylus timidus Vences & Glaw, 2005
Paratypes in ZFMK and ZMA, holotype in ZMA
Origin: Madagascar

Current name: Guibemantis timidus

Mantidactylus tschenki Glaw & Vences, 2001
Paratypes in ZFMK and MRSN, holotype in ZSM
Origin: Madagascar

Current name: Gephyromantis (Duboimantis) tschenki

Mantidactylus zavona Vences, Andreone, Glaw & Ran-
drianirina, 2003

Paratypes in ZFMK, MRSN, PBZT and UADBA, holo-
type in ZSM

Origin: Madagascar

Current name: Gephyromantis (Duboimantis) zavona

Mantidactylus zipperi Vences & Glaw, 2004

Holotype: ZFMK 60090, paratypes in ZSM and MRSN
Origin: Madagascar

Current name: Mantidactylus (Chonomantis) zipperi

Mantidactylus zolitschka Glaw & Vences, 2004
Holotype: ZFMK 60110, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Mantidactylus (Ochthomantis) zolitschka

Tsingymantis antitra Glaw, Hoegg & Vences, 2006
Paratypes in ZFMK, ZSM and UADBA, holotype in ZSM
Origin: Madagascar

Rhacophoridae

Philautus quyeti Nguyen Q.T., Hendrix, Bohme, Vu &
Ziegler, 2008

Holotype: ZFMK 82999, paratype in VNUH

Origin: Vietnam

Rhacophorus orlovi Ziegler & J. Kohler, 2001
Holotype: ZFMK 71368, paratypes in ZMFK, MTKD,
CRES and ZMB

Origin: Vietnam

Bonn zoological Bulletin 59: 79-108

Ranidae

Amunirana fonensis Rodel & Bangoura, 2004
Paratypes in ZFMK and SMNS, Holotype in SMNS
Origin: Guinea

Amolops amoropalamus Matsui, 1986

Paratypes in ZFMK and OMNH, Holotype in OMNH
Origin: Sabah, Malaysia

Current name: Meristogenys amoropalamus

Hylarana amnicola Perret, 1977

Paratypes in ZFMK and MHNG, Holotype in MHNG
Origin: Cameroon

Current name: Amnirana amnicola

HAylarana asperrima Perret, 1977

Paratypes in ZFMK and MHNG, Holotype in MHNG
Origin: Cameroon

Current name: Amnirana asperrima

Rana aragonensis Palanca Soler, Rodriguez Vieites &
Suarez Martinez, 1995

Paralectotypes in ZFMK and MNHN, Holotype in
MNCN

Origin: Spain

Current name: Rana temporaria

Rana balcanica H. Schneider, Sinsch & Sofianidou, 1993
Holotype: ZFMK 52825, paratypes in ZFMK

Origin: Greece

Current name: Pelophylax kurtklari

Rana epeirotica H. Schneider, Sinsch & Kyriakopoulou-
Sklavounou, 1984

Holotype: ZFMK 41679, paratypes in ZFMK

Origin: Greece

Current name: Pelophylax epeiroticus

Rana holtzi Werner, 1898y

Syntypes: ZFMK 28913914, one further syntype in
NMW

Origin: S Turkey

Rana levantina H. Schneider, Sinsch & Nevo, 1992
Holotype ZFMK 52836, paratypes in ZFMK
Origin: Israel

Current name: Pelophylax bedriagae

Rana shqiperica Hotz, Uzzel, Giinther & Heppich, 1987
Paratypes in ZFMK, ANSP, UIMNH, ZMB, BO
(Beograd) and MHNG, holotype in ANSP

Origin: Albania

Current name: Pelophylax shqipericus

©ZFMK

92 Wolfgang Bohme

TESTUDINES
Chelidae

Chelodina reimanni Philippen & Grossmann, 1990
Paratypes in ZFMK, MTKD and UFNM, holotype
MTKD

Origin: West Papua, Indonesia

Current name: Chelodina novaeguineae

Bataguridae

Cuora cyclornata Blanck, McCord & Le, 2006
Holotype: ZFMK 71348, no designated paratypes
Origin: Vietnam

Current name: Cuora trifasciata

Geoemyda (trijuga) wiroti Nutaphand, 1979
Holotype: ZFMK 46510, no paratypes
Origin: Thailand

Current name: Melanochelys trijuga edeniana

Emydidae

Emys orbicularis (galloitalica) capolongoi Fritz, 1995
Paratypes in ZFMK, BMNH, HCC, MCZ, MGL, MHNG,
MNHN, MTKD, MZUF, MZUN, NMB, SMF, ZMB and
ZSM, holotype in MNHN

Origin: Sardinia, Italy

Current name: Emys orbicularis capolongoi

SQUAMATA
Iguanidae

Amblyrhynchus cristatus venustissimus Eib|-Eibesfeld,
1956

Paratypes in ZFMK, SMF, HLMD and “Haus der Natur”
Salzburg, holotype in SMF

Origin: Galapagos Ids., Ecuador

Polychrotidae

Anolis albimaculatus Henle & Ehrl, 1991
Holotype: ZFMK 34263, no paratypes
Origin: Peru

Current name: Anolis boettgeri

Anolis latifrons Berthold, 1845
Holotype: ZFMK 21342, no paratypes
Origin: “Provinz Popayan”, Colombia

Bonn zoological Bulletin 59: 79-108

Norops bicaorum G. Kohler, 1996

Paratypes in ZFMK, SMF, UNAH, UF and IRSNB, ho-
lotype in SMF

Origin: Utila Id., Honduras

Polychrus gutturosus Berthold, 1845
Holotype: ZFMK 21341, no paratypes
Origin: “Provinz Popayan”, Colombia

Sceloporidae

Liolaemus buergeri Werner, 1907
Syntypes: ZFMK 26706—707
Origin: Chile

Uromastycidae

Lacerta aegyptia Forskal, 1775
Neotype: ZFMK 44216

Origin: Egypt

Current name: Uromastyx aegyptia

Leiolepis belliana rubritaeniata Mertens, 1961
Paratypes in ZFMK and SMF, holotype in SMF
Origin: Thailand

Current name: Leiolepis rubritaeniata

Leiolepis boehmei Darevsky & Kupriyanova, 1993
Holotype: ZFMK 24651, paratypes in ZFMK and ZISP
Origin: S Thailand

Leiolepis guentherpetersi Darevsky & Kupriyanova, 1993
Paratypes in ZFMK, ZISP and NHRM, holotype in ZISP
Origin: Vietnam

Leiolepis triploida Peters, 1971

Paratypes in ZFMK, MCZ and ZMB, holotype in MCZ
Origin: “Malayisch-thailandisches Grenzgebiet auf der
Malayischen Halbinsel”, Malaysia

Uromastyx alfredschmidti Wilms & Bohme, 2000
Holotype: ZFMK 24643, paratypes in MHNP, MHNG and
NMW

Origin: S Algeria

Uromastyx geyri L. Miller, 1922
Neotype: ZFMK 9230, paratypes in ZFMK and ZSM
Origin: S Algeria

Uromastyx leptieni Wilms & Bohme, 2000
Holotype: ZFMK 52398, paratypes in BMNH
Origin: United Arab Emirates

Current name: Uromastyx aegyptia leptieni

©ZFMK

Type specimens in ZFMK: Section of Herpetology 93

Uromastyx yemenensis Wilms & Schmitz, 2007
Holotype: ZFMK 47861, paratypes in ZFMK and MTKD
Origin: Yemen

Uromastyx yemenensis shobraki Wilms & Schmitz, 2007
Holotype: ZFMK 48681, paratypes in ZFMK and MHNG
Origin: Yemen

Current name: Uromastyx shobraki

Agamidae

Acanthocercus guentherpetersi Largen & Spawls, 2006
Paratypes in ZFMK, BMNH and ZMG, holotype in
BMNH

Origin: Eritrea

Agama agama mucosoensis Hellmich, 1957
Paratypes in ZFMK and ZSM, holotype in ZSM
Origin: Angola

Current name: Agama mucosoensis.

Agama chernovi Ananjeva, Peters & Rzhepakowsky,
1981

Paratypes in ZFMK, MTKD, ZMB and ZISP, holotype in
ZISP

Origin: Tadzhikistan

Current name: Laudakia bochariensis.

Agama finchi Bohme, Wagner, Malonza, Lotters & J.
Kohler, 2005

Paratypes in ZFMK and NMK, holotype in NMK
Origin: W Kenya

Agama lebretoni Wagner Bare} & Schmitz, 2009 (“2007”)
Holotype: ZFMK 87698, paratypes in ZFMK, MHNG and
ZMB

Origin: Cameroon

Agama pakistanica Baig, 1989
Paratypes in ZFMK and PMNH
Origin: N Pakistan

Current name: Laudakia pakistanica.

Agama savignii Duméril & Bibron, 1837
Neotype: ZFMK 77470

Origin: Egypt

Current name: 7rapelus savignii.

Agama stoliczkana altaica Peters, 1971
Paratypes in ZFMK, ZMB, ZISP, BIU-B and ZIH
Origin: Mongolia

Current name: Laudakia stoliczkana altaica.

Bonn zoological Bulletin 59: 79-108

Calotes (Bronchocela) intermedius Berthold, 1842
Syntypes: ZFMK 27099-101

Origin: Java, Indonesia

Current name: Bronchocela jubata.

Calotes nigriplicatus Hallermann, 2001 (“2000”)
Holotype 26379, no paratypes
Origin: Moluccas, Indonesia

Gonocephalus denzeri Manthey, 1991
Holotype: ZFMK 50527, paratype in ZFMK
Origin: Sarawak, Malaysia

Current name: Gonocephalus bornensis.

Lacerta agama Linnaeus, 1758
Neotype: ZFMK 15222
Origin: Cameroon

Current name: Agama agama.

Lacerta stellio Linnaeus, 1758
Neotype: ZFMK 2063

Origin: Delos Id., Greece
Current name: Laudakia stellio.

Laudakia pakistanica auffenbergi Baig & Bohme, 1996
Holotype: ZFMK 54471, paratypes in ZFMK and PMNH
Origin: N Pakistan

Laudakia pakistanica khani Baig & Bohme, 1996
Holotype: ZFMK 54468 (“54469”), paratypes in ZFMK
and PMNH

Origin: N Pakistan

Pseudocalotes ziegleri Hallermann, Nguyen Q.T., Orlov
& Ananjeva, 2010

Paratypes in ZFMK, IEBR and ZISP, holotype in ZISP
Origin: Vietnam

Stellio caucasius triannulatus Ananjeva & Atayev, 1984
Paratypes in ZFMK and ZISP

Origin: Turkmenia

Current name: Laudakia microlepis triannulata.

Trapelus schmitzi Wagner & Bohme, 2007
Holotype: ZFMK 2590, no paratypes
Origin: Tchad

Chamaeleonidae
Bradypodion oxyrhinum Klaver & Bohme, 1988
Holotype: ZFMK 46406, paratypes in ZFMK and ZMUC

Origin: Tanzania
Current name: Kinyongia oxyrhina.

©ZFMK

94 Wolfgang Bohme

Bradypodion tavetanum boehmei Lutzmann & Neéas,
2002

Holotype: 63373, paratypes in ZFMK

Origin: Kenya

Current name: Kinyongia boehmei.

Calumma glawi Bohme, 1997
Holotype: ZFMK 62909, paratype in ZFMK
Origin: Madagascar

Chamaeleo (Trioceros) balebicornutus Tilbury, 1998
Holotype: ZFMK 63049, paratypes in ZFMK

Origin: Ethiopia

Current name: Trioceros balebicornutus.

Chamaeleo bitaeniatus Fischer, 1884

Syntype: ZFMK 66060, further syntypes in ZMH
Origin: Kenya

Current name: Jrioceros bitaeniatus.

Chamaeleo (africanus) calcaricarens Bohme, 1985
Paratypes in ZFMK and BMNH, holotype in BMNH
Origin: Ethiopia

Current name: Chamaeleo calcaricarens.

Chamaeleo camerunensis L. Miller, 1909
Paralectotype in ZFMK, levctotype in ZSM
Origin: Cameroon

Current name: Trioceros camerunensis.

Chamaeleo eisentrauti Mertens, 1968

Holotype: ZFMK 5781, paratypes in ZFMK, SMF and
MHNG

Origin: Cameroon

Current name: 7rioceros eisentrauti.

Chamaeleo harennae fitchi Necas, 2004
Holotype: ZFMK 81201, paratype in ZFMK
Origin: Ethiopia

Current name: Trioceros harennae fitchi.

Chamaeleo laterispinis brookesiaeformis Bohme, 1982
Holotype: ZFMK 30785, no paratypes

Origin: Tanzania

Current name: Trioceros laterispinis.

Chamaeleo marsabitensis Tilbury, 1991
Paratypes in ZFMK and BMNH

Origin: N Kenya

Current name: 7rioceros marsabitensis.

Chamaeleo narraioca Neéas, Modry & Slapeta, 2003
Holotype: ZFMK 73956, paratypes in ZFMK and NMP
Origin: N Kenya

Current name: Trioceros narraioca.

Bonn zoological Bulletin 59: 79-108

Chamaeleo necasi Ullenbruch, Krause & Bohme, 2007
Holotype: ZFMK 76922, paratypes in ZFMK and MRAC
Origin: Togo

Chamaeleo (Trioceros) ntunte Necas, Modry & Slapeta,
2005 (“2003”)

Holotype: ZFMK 73963, paratypes in ZFMK

Origin: N Kenya

Current name: Trioceros ntunte.

Chamaeleo quadricornis gracilior Bohme & Klaver,
1981

Holotype: ZFMK 15291, paratypes in ZFMK, MHNG and
RMB

Origin: Cameroon

Current name: 7rioceros quadricornis gracilior.

Chamaeleo wiedersheimi perreti Klaver & Bohme, 1992
Paratypes in ZFMK, SMF and ZMB, holotype in MHNG
Origin: Cameroon

Current name: 7rioceros perreti.

Furcifer timoni Glaw, J. Kohler & Vences, 2009
Paratypes in ZFMK, ZSM and UADBA, holotype in ZSM
Origin: Madagascar

Kinyongia asheorum Necas, Sindaco, Koreny, Kopecna,
Malonza & Modry, 2009

Paratypes in ZFMK, MCC and NMK, holotype in MCC
Origin: Kenya

Kinyongia vanheygeni Necas, 2009

Holotype: ZFMK 88383, lost before ever reaching
ZFMK, no paratypes

Origin: Tanzania

Rhampholeon (Rhinodigitum) acuminatus Mariaux &
Tilbury, 2006

Paratypes in ZFMK, PEM and MHNG, holotype in MH-
NG

Origin: Tanzania

Rhampholeon (Rhinodigitum) beraduccii Mariaux &
Tilbury, 2006

Paratypes in ZFMK and MHNG, holotype in MHNG
Origin: Tanzania

Rhampholeon moyeri Menegon, Salvidio & Tilbury, 2002
Paratypes in ZFMK, MTSN and NMZB, hoilotype in
MZSN

Origin: Tanzania

Rhampholeon uluguruense Tilbury & Emmrich, 1996
Paratypes in ZFMK, NMZB and ZMB, holotype in ZMB
Origin: Tanzania

©ZFMK

Type specimens in ZFMK: Section of Herpetology 95

Trioceros hanangensis Krause & Bohme, 2010
Holotype: ZFMK 82369, paratypes in ZFMK
Origin: Tanzania

Diplodactylidae

Eurydactylodes agricolae Henkel & Bohme, 2001
Holotype: ZFMK 71022, paratype in ZFMK
Origin: New Caledonia

Platydactylus vieillardi Bavay, 1869
Neotype: ZFMK 46981

Origin: New Caledonia

Current name: Eurydactylodes vieillardi.

Eublepharidae

Goniurosaurus araneus Grismer, Viets & Boyle, 1999
Paratypes in ZFMK and HLMD, holotype in HLMD
Origin: Vietnam

Goniurosaurus catbaensis Ziegler, Nguyen Q.T.,
Schmitz, Stenke & Résler, 2008

Paratypes in ZFMK MHNG and VNUH, holotype in
IEBR

Origin: Vietnam

Phyllodactylidae

Gecko fascicularis Daudin, 1802

Neotype: ZFMK 35631

Origin: Libya

Current name: Zarentola mauritanica fascicularis.

Phyllodactylus delsolari Venegas, Townsend, C. Koch &
Bohme, 2008

Paratypes in ZFMK, UF and MUSM, holotype in UF
Origin: N Peru

Phyllodactylus thompsoni Venegas, Townsend, C. Koch
& Boéhme, 2008)

Paratypes in ZFMK and MUSM, holotype in UF
Origin: N Peru

Phyllopezus maranjonensis Koch, Venegas & Bohme,
2006

Paratypes in ZFMK and MHNSM, holotype in MHNSM
Origin: N Peru

Bonn zoological Bulletin 59: 79-108

Tarentola annularis relicta Joger, 1984

Holotypus: ZFMK 25963, paratypes in BMNH, USNM
and UZMK

Origin: S Sudan

Tarentola boehmei Joger, 1984

Holotype: ZFMK 26133, paratypes in ZFMK, MNHN,
CNHM and USNM

Origin: Morocco

Tarentola boettgeri bischoffi Joger, 1984
Paratypes in ZFMK and SMF, Holotype in SMF
Origin: Selvagens Ids., Portugal

Tarentola boettgeri hierrensis Joger & Bischoff, 1983
Holotype: ZFMK 24906, paratypes in ZFMK
Origin: Hierro, Canary Ids., Spain

Tarentola darwini Joger, 1984 (“1983”)
Holotype: ZFMK 37256, paratypes in BMNH and ZSM
Origin: Sao Tiago, Cape Verde Islands

Tarentola delalandii hoggarensis Werner, 1937
Neotypus: ZFMK 36609

Origin: Algeria

Current name: 7arentola ephippiata hoggarensis.

Tarentola ephippiata senegambiae Joger, 1984
Holotype: ZFMK 19979, paratypes in ZFMK
Origin: Senegal

Tarentola ephippiata nikolausi Joger, 1984
Holotype: ZFMK 38431, paratype in BMNH
Origin: NE Sudan

Tarentola gomerensis Joger & Bischoff, 1983
Holotype: ZFMK 35207, paratypes in ZFMK, ZSM,
NMW, ZMH and MNHN

Origin: Gomera Id., Canary Ids., Spain

Tarentola mauritanica juliae Joger, 1984

Holotype: ZFMK 26132, paratypes in ZFMK, CNHM,
ZSM and USNM

Origin: Morocco

Tarentola neglecta geyri Joger, 1984
Holotype: ZFMK 2134, paratypes in ZFMK and BMNH
Origin: Algeria

Tarentola parvicarinata Joger, 1980

Holotype: ZFMK 22157, paratypes in ZFMK, BMNH,
MNHN, USNM, MHNG, LBUM, IFAN and NHMG
Origin: Mali

©ZFMK

96 Wolfgang Bohme

Sphaerodactylidae

Lepidoblepharis conolepis Avila-Pires, 2001
Paratypes in ZFMK and MHNG, holotype in MHNG
Origin: Ecuador

Pristurus samhaensis Rosler & Wranik, 1999
Paratypes in ZFMK, MTKD and HLMD, holotype in
MTKD

Origin: Socotra Id., Yemen

Pristurus schneideri Rosier, J. Kohler & Bohme, 2008
Holotype: ZFMK 86654, paratypes in ZFMK, HLMD and
NHCY

Origin: Yemen

Pristurus obsti Rosler & Wranik, 1999
Paratypes in ZFMK, MTKD, HLMD and CPHR
Origin: Socotra Id., Yemen

Sphaerodactylus buergeri Werner, 1900
Holotype ZFMK 20712, no paratypes
Origin: Trinidad

Current name: Sphaerodactylus molei.

+ Sphaerodactylus dommeli Bohme, 1984
Holotype ZFMK 66238, paratype in SMNS
Origin: Dominican Republic

Tropiocolotes wolfgangboehmei Wilms, Shobrak &
Wagner, 2010

Holotype: ZFMK 43668, paratype in ZFMK

Origin: Saudi Arabia

Gekkonidae

Alsophylax (Altiphylax) boehmei Szczerbak, 1991
Holotype: ZFMK 38773, paratype in ZFMK
Origin: N Pakistan

Current name: A/tiphylax stoliczkai.

Cyrtodactylus consobrinus kinabaluensis Malkmus,
1989

Holotype: ZFMK 46741, no paratype

Origin: Sabah, Malaysia

Current name: Cyrtodactylus consobrinus.

Cyrtodactylus cattienensis GeiBler, Nazarov, Orlov,
Bohme, Trung, Nguyen Q.T. & Ziegler, 2009

Paratypes in ZFMK IEBR, VNUH, ZMMU and ZISP, ho-
lotype in IEBR

Origin: Vietnam

Cyrtodactylus cryptus Heidrich, Résler, Vu, Bohme &
Ziegler, 2007

Bonn zoological Bulletin 59: 79-108

Holotype ZFMK 86037, paratypes in ZFMK and VNUH
Origin: Vietnam

Cyrtodactylus kotschyi buchholzi Beutler & Gruber, 1977
Holotype: ZFMK 9956, paratypes in ZFMK and NMW
Origin: Siphnos Id., Greece

Current name: Cyrtopodion kotschyi buchholzi.

Cyrtodactylus kotschyi schultzewestrumi Beutler & Gru-
ber, 1977

Paratypes in ZFMK, and ZSM, holotype in ZSM
Origin: Valaxa Id., Greece

Current name: Cyrtopdion kotschyi schultzewestrumi.

Cyrtodactylus kotschyi tinensis Beutler & Frér, 1980
Paratypes in ZFMK, ZSM and ZMB, holotype in ZSM
Origin: Tinos Id., Greece

Current name: Cyrtopodion kotschyi tinensis

Cyrtodactylus phongnhakebangensis Ziegler, Rosler,
H.W. Herrmann & Than, 2003 (“2002”)

Holotype: ZFMK 76193, paratypes in ZFMK, VNUH and
MTKD

Origin: Vietnam

Cyrtodactylus pseudoquadrivirgatus Rosler, Vu, Nguyen
Q.T., Ngo & Ziegler, 2008

Holotype: ZFMK 83895, paratypes iN ZFMK, IEBR and
UNS

Origin: Vietnam

Cyrtodactylus roesleri Ziegler, Nazarov, Orlov, Nguyen
Q.T., Vu, Dang, Dinh & Schmitz, 2010

Holotype: ZFMK 89377, paratypes in ZFMK, IEBR,
MNHG, VNUH, ZISP and ZMMU

Origin: Vietnam

Cyrtodactylus wayakonei Nguyen Q.T., Kingsada, R6s-
ler, Auer & Ziegler, 2010

Paratypes in ZFMK, MTKD and NUOL, holotype in
IEBR

Origin: Laos

Cyrtodactylus ziegleri Nazarov, Orlov, Nguyen N.S. & Ho
(2008)

Paratypes in ZFMK, ITBCZ, ZISP and ZMMU, holoty-
pe in ZISP

Origin: Vietnam

Dixonius aaronbaueri Ngo & Ziegler, 2009
Paratypes in ZFMK, UNS and ZMB, holotype in UNS
Origin: Vietnam

Gekko canhi Roésler, Nuyen Q.T., Doan, Ho, Nguyen T.T.
& Ziegler, 2010

©ZFMK

Type specimens in ZFMK: Section of Herpetology 97

Paratypes in ZFMK and VNMN, holotype in IEBR
Origin: Vietnam

Gekko grossmanni Ginther, 1994
Paratypes in ZFMK and ZMB, holotype in ZMB
Origin: Vietnam

Gekko scientiadventura Rosler, Ziegler, Thanh, H.W.
Herrmann & Bohme (2005 (“2004”)

Holotype: ZFMK 76198, paratypes in ZFMK and VNUH
Origin: Vietnam

Gekko siamensis Grossmann & Ulber, 1990
Paratypes in ZFMK, MTKD and pers. coll. W.G.
Origin: Thailand

Gekko ulikovskii Darevsky & Orloy, 1994
Paratypes in ZFMK and ZISP, holotype in ZISP
Origin: Vietnam

Current name: Gekko badenii.

Gymnodactylus gastropholis Werner, 1917

Holotype: 27094, paratypes in ZFMK, ZMH and CAS
Origin: Iran

Current name: Cyrtopodion gastropholis.

Gymnodactylus kotschyi rumelicus L. Miller, 1939
Neotype: ZFMK 24327, paratype in ZFMK

Origin: Bulgaria

Current name: Cyrtopodion kotschyi rumelicus.

Hemidactylus lamaensis Ullenbruch, Grell & Bohme,
2010

Holotype: ZFMK 77072, no paratypes

Origin: Benin

Hemidactylus pseudomuriceus Henle & Bohme, 2003
Holotype: ZFMK 42409, paratypes in ZFMK
Origin: Ivory Coast

Hemidactylus turcicus lavadeserticus Moravec & Bohme,
1997

Paratypes in ZFMK and NMP, holotype in NMP
Origin: Syria

Current name: Hemidactylus lavadeserticus.

Hemidactylus turcicus spinalis Buchholz, 1954
Holotype: ZFMK 33227, paratypes in ZFMK
Origin, Baleartic Ids., Spain

Homopholis (Blaesodactylus) antongilensis Bohme &
Meier, 1980

Holotype: ZFMK 21772, no paratypes

Origin: Madagascar

Current name: Blaesodactylus antongilensis.

Bonn zoological Bulletin 59: 79-108

Pachydactylus atorquatus Bauer, Barts & Hulbert, 2006
Paratypes in ZFMK, NMNW and TM, holotype in TM
Origin: Namibia

Pachydactylus capensis katanganus De Witte, 1953
Paratypes in ZFMK and IRSN
Origin: D.R. Congo

Paroedura bastardi ibityensis Rosler & Kriiger, 1998
Paratypes in ZFMK, CPHR, MNHM and MTKD, holo-
type in MTKD

Origin: Madagascar

Current name: Paroedura ibityensis.

Paroedura lohatsara Glaw, Vences & K. Schmidt, 2001
Paratypes in ZFMK and ZSM, holotype in ZSM
Origin: Madagascar

Phelsuma berghofi Kriiger, 1996
Holotype: ZFMK 62991, no registered paratypes
Origin: Madagascar

Phelsuma borbonica (agalegae) mater Meier, 1995
Holotype: ZFMK 57901, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Phelsuma borbonica.

Phelsuma leiogaster isakae Meier, 1993

Holotype: ZFMK 54981, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Phelsuma modesta.

Phelsuma leiogaster trautmanni Meier, 1993
Holotype: ZFMK 55122, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Phelsuma modesta.

Phelsuma lineata elanthana Kriger, 1996
Holotype: ZFMK 47227, paratypes in ZFMk and ZSM
Origin: Madagascar

Phelsuma madagascariensis (madagscariensis) boehmei
Meier, 1984

Holotype: ZFMK 32107, paratypes in ZFMK and ZSM
Origin: Madagascar

Current name: Phelsuma madagascariensis boehmei.

Phelsuma madagascariensis (sundbergi) ladiguensis
Bohme & Meier, 1981

Holotype: ZFMK 33411, paratypes in ZFMK and ZSM
Origin: La Digue Id., Seychelles

Current name: Phelsuma sundbergi ladiguensis.

©ZFMK

98 Wolfgang Bohme

Phelsuma nigristriata Meier, 1984
Holotype: 40317, paratypes in ZFMK and ZSM
Origin: Mayotte Id., Comoro Ids.

Phelsuma pusilla hallmanni Meier, 1989

Holotype: ZFMK 54500, paratypes in ZFMK, SMF and
ZSM

Origin: Madagascar

Phelsuma quadriocellata lepida Kriiger, 1993
Holotype: ZFMK 54500, paratypes in ZFMK, SMF and
ZSM

Origin: Madagascar

Phelsuma quadriocellata parva Meier, 1983
Holotype: ZFMK 19315, paratypes in ZFMK and ZSM
Origin: Madagascar

Phelsuma robertmertensi Meier, 1980
Holotype: ZFMK 29900, paratypes in ZFMK and ZSM
Origin: Mayotte Id., Comoro Ids.

Phelsuma seippi Meier, 1987
Holotype ZFMK 46702, no catalogued paratypes
Origin: Madagascar

Phelsuma v-nigra anjouanensis Meier, 1986
Holotype: ZFMK 42077, paratypes in ZFMK and ZSM
Origin: Anjouan, Comoro Ids.

Phelsuma yv-nigra comoraegrandensis Meier, 1986
Holotype: ZFMK 42047, paratypes in ZFMk and ZSM
Origin: Grande Comore Id., Comoro Ids.

Phelsuma v-nigra pasteuri Meier, 1984

Holotype: ZFMK 40310, paratypes in ZFMK and ZSM
Origin: Mayotte Id., Comoro Ids.

Current name: Phelsuma pasteuri.

Stellio fimbriatus J.G. Schneider, 1797
Neotype: ZFMK 36503

Origin: Madagascar

Current name: Uroplatus fimbriatus.

Uroplatus giganteus Glaw, Kosuch, Henkel, Sound &
Bohme, 2006

Paratypes in ZFMK, ZSM and UADBA, holotype in ZSM
Origin: Madagascar

Uroplatus henkeli Bohme & Ibisch, 1990
Holotype ZFMK 48229, paratypes in ZFMK and ZSM
Origin: Madagascar

Uroplatus sikorae sameiti Bohme & Ibisch, 1990
Holotype: ZFMK 46005, paratypes in ZFMK, SMF and

Bonn zoological Bulletin 59: 79-108

ZSM
Origin: Madagascar

Amphisbaenidae

Lepidosternon infraorbitale Berthold, 1859
Holotype: ZFMK 26546, no paratypes
Origin: Bahia, Brazil

Current name: Leposternon infraorbitale.

Gymnophthalmidae

Echinosaura brachycephala Kohler, Bohme & Schmitz,
2004

Paratypes in ZFMK, MHNG and SMF, holotype in MH-
NG

Origin: Ecuador

Teiidae

Tupinambis teguixin buzioensis P. Miller, 1969
Holotype: ZFMK 89199, paratypes in ZFMK
Origin: Buzio Id., SE Brazil

Current name: 7upinambis merianae buzioensis.

Tupinambis teguixin sebastiani P. Miller, 1968
Holotype: ZFMK 89193, paratypes in ZFMK
Origin: Sao Sebastiao Id., S Brazil

Current name: 7upinambis merianae sebastiani.

Lacertidae

Acanthodactylus busacki Salvador, 1982

Paratypes in ZFMK, BMNH, FMNH, CM, CAS, MNHN
and EBD, holotype in BMNH

Origin: Morocco

Algyroides marchi niethammeri Buchholz, 1964
Holotype: ZFMK 4541, no paratypes

Origin: Spain

Current name: A/gyroides marchi.

Eremias afghanistanica Bohme & Szczerbak, 1991
Holotype: ZFMK 8584, paratype in ZFMK
Origin: Afghanistan

Eremias barbouri K.P. Schmidt, 1927

Paratypes in ZFMK and AMNH, holotype in AMNH
Origin: Mongolia

Current name: Eremias argus barbouri.

©OZFMK

Type specimens in ZFMK: Section of Herpetology 99

Eremias lalezharica Moravec, 1994
Paratypes in ZFMK and NMP, holotype in NMP
Origin: [ran

Eremias strauchi kopetdaghica Szczerbak, 1972
Paratypes in ZFMK and NMP, holotype in NMP
Origin: Turkmenia

Eremias velox suphani Basoglu & Hellmich, 1968
Paratypes in ZFMK, SZE and ZSM, holotype in SZE
Origin: Turkey

Current name: Eremias suphani.

Gallotia atlantica mahoratae Bischoff, 1985
Holotype: ZFMK 35034, paratypes in ZFMK
Origin: Fuerteventura Id., Canary Ids., Spain

Gallotia galloti eisentrauti Bischoff, 1982
Holotype: ZFMK 34800, paratypes in ZFMK and MNHN
Origin: Tenerife, Canary Ids., Spain

Gallotia galloti insulanagae Martin, 1985
Paratypes in ZFMK, ZLU and MCNSC, holotype in ZLU
Origin: Tenerife, Canary Ids., Spain

+ Gallotia goliath bravoana Hutterer, 1985

Holotype: ZFMK 42392, paratypes in ZFMK

Origin: Late Pleistocene of La Gomera Id., Canary Ids.,
Spain

Current name: Gallotia simonyi bravoana.

Gallotia intermedia Hernandez, Nogales & Martin, 2000
Paratypes in ZFMK (still to be received), DZUL, TFMC,
holotype in DZUL

Origin: Tenerife, Canary Ids.

+ Gallotia simonyi gomerana Hutterer, 1985

Holotype: ZFMK 42399, paratypes in ZFMK

Origin: Late Pleistocene of La Gomera Id., Canary Ids.,
Spain

Current name: Gallotia simonyi bravoana

Lacerta agilis brevicaudata Peters, 1958

Paratypes in ZFMK, ZMB, MM and ZISP, holotype in
ZISP

Origin: Armenia

Lacerta agilis ioriensis Peters & Muskhelishvili, 1968
Paratypes in ZFMK, ZIT and ZMB, holotype in ZIT
Origin: Georgia

Lacerta bendimahiensis Schmidtler, Eiselt & Darevsky,

1994
Paratypes in ZFMK and ZSM, holotype in ZSM

Bonn zoological Bulletin 59: 79-108

Origin: Turkey
Current name: Darevskia bendimahiensis.

Lacerta cappadocica muhtari Eiselt, 1979

Paratypes in ZFMK, SZE, NMW, ZSM, ZMH, BMNH,
FMNH and CAS

Origin: Turkey

Current name: Apathya cappadocica muhtari.

Lacerta cappadocica schmidtlerorum Eiselt, 1979
Paratypes in ZFMK, NMW and ZSM, holotype in ZSM
Origin: Turkey

Current name: Apathya cappadocica schmidtlerorum.

Lacerta cyanisparsa Bischoff & Schmidtler, 1999
Paratypes in ZFMK, SMF, ZSM and CAS, holotype in
ZSM

Origin: Turkey

Current name: Phoenicolacerta cyanisparsa.

Lacerta derjugini abchasica Bischoff, 1982
Paratypes in ZFMK and MM, holotype in MM
Origin: Abchasia, Georgia

Current name: Darevskia derjugini abchasica.

Lacerta derjugini barani Bischoff, 1982

Paratypes: ZFMK, MM, SZE and CAS, holotype in MM
Origin: Georgia

Current name: Darevskia derjugini barani.

Lacerta derjugini boehmei Bischoff, 1982

Paratypes: ZFMK, MTKD, MM, ZMB, AMNH and US-
NM, holotype in MTKD

Origin: Russia

Current name: Darevskia derjugini boehmei.

Lacerta derjugini orlowae Bischoff, 1984
Holotype: ZFMK 38047, paratypes in MTKD
Origin: Georgia

Current name: Darevkskia derjugini.

Lacerta erhardii pachiae Wettstein, 1937
Paralectotypes in ZFMK and NMW, lectotype in NMW
Origin: Pachia Id., Cyclades, Greece

Current name: Podarcis erhardii pachiae.

Lacerta erhardii syrinae Wettstein, 1937
Paralectotypes in ZFMK and NMW, lectotype in NMW
Origin: Syrina Id., Cyclades, Greece

Current name: Podarcis erhardii syrinae.

Lacerta erhardii weigandi Gruber & Schultze-Westrum,

1971
Holotype: ZFMK 10914, paratypes in ZFMK and MTKD

©ZFMK

100 Wolfgang Bohme

Origin: Piperi Id., Northern Sporades, Greece
Current name: Podarcis gaigeae weigandi

Lacerta hieroglyphica Berthold, 1842
Syntypes: ZFMK 21220-221

Origin: W Turkey

Current name: Podarcis siculus hieroglyphicus.

Lacerta kulzeri petraea Bischoff & Miller, 1999
Holotype: ZFMK 65019, paratypes in ZFMK, JUMR,
HUJ and SMF

Origin: Jordan

Current name: Phoenicolacerta kulzeri petraea.

Lacerta lepida nevadensis Buchholz, 1963
Holotype 3581, paratypes in ZFMK
Origin: SE Spain

Current name: Timon lepidus nevadensis.

Lacerta lilfordi conejerae L. Miller, 1927
Neotype: ZFMK 11898

Origin: Conejera Id., Balearic Ids., Spain
Current name: Podarcis lilfordi conejerae.

Lacerta lilfordi jordansi L. Miller, 1927

Paratypes in ZFMK and ZSM, holotype (formerly in
ZSM) destroyed in WW II

Origin: La Guardia, Balearic Ids., Spain

Current name: Podarcis lilfordi jordansi.

Lacerta muralis adolfjordansi Buchholz, 1962
Holotype: ZFMK 1854, paratypes in ZFMK and ZMB
Origin: Ananes Id., Cyclades, Greece

Current name: Podarcis milensis adolfjordansi.

Lacerta muralis var. coerulea Eimer, 1872

Syntypes: ZFMK 21322-333, further syntypes (Univer-
sity of Tubingen) seem to be lost.

Origin: Capri Id., Italy

Current name: Podarcis siculus coeruleus.
Lacerta oertzeni budaki Eiselt & Schmidtler, 1987
(“1986”)

Paratypes in ZFMK and NMW, holotype in NMVW
Origin: Turkey

Current name: Phoenicolacerta oertzeni budaki.

Lacerta parvula adjarica Darevsky & Eiselt, 1980
Paratypes in ZFMK, ZISP, MHNG, MHNB, SMF, MNH
and ZIK

Origin: Abchasia, Georgia

Current name: Darevskia parvula adjarica.

Bonn zoological Bulletin 59: 79-108

Lacerta peloponnesiaca lais Buchholz, 1960

Holotype: ZFMK 12115, paratypes in ZFMK, NMW, MM
and MTKD

Origin: Peloponnesos, Greece

Current name: Podarcis peloponnesiacus lais.

Lacerta peloponnesiaca phryne Buchholz, 1960
Holotype: ZFMK 12263, paratypes in ZFMK and NMW
Origin: Peloponnesos, Greece

Current name: Podarcis peloponnesiacus lais.

Lacerta peloponnesiaca thais Buchholz, 1967
Holotype: ZFMK 12261, paratypes in ZFMK, NMW and
MTKD

Origin: Peloponnesos, Greece

Current name: Podarcis peloponnesiacus thais.

Lacerta pityusensis caragolensis Buchholz, 1954
Holotype: ZFMK 12497, paratypes in ZFMK
Origin: Caragolé, Balearic Ids., Spain

Current name: Podarcis pitvusensis caragolensis.

Lacerta pityusensis characae Buchholz, 1954
Holotype: ZFMK 12467, paratypes in ZFMK and MTKD
Origin: Characa Id., Baleatric Ids., Spain

Current name: Podarcis pityusensis characae.

Lacerta pityusensis hortae Buchholz, 1954

Holotype: ZFMK 12410, paratypes in ZFMK and MTKD
Origin: Hort Id., Balearic Ids., Spain

Current name: Podarcis pityusensis hortae.

Lacerta pityusensis puercosensis Buchholz, 1954
Holotype: ZFMK 12411, paratypes in ZFMK and MTKD
Origin: Puercos Id., Balearic Ids., Spain

Current name: Podarcis pityusensis puercosensis.

Lacerta pityusensis purroigensis Buchholz, 1954
Holotype: ZFMK 12409, paratypes in ZFMK and MTKD
Origin: Purroige Id., Balearic Ids., Spain

Current name: Podarcis pityusensis purroigensis.

Lacerta pityusensis sabinae Buchholz, 1954
Holotype: ZFMK 12607, paratypes in ZFMK
Origin: Sabina Id., Balearic Ids., Spain
Current name: Podarcis pityusensis sabinae.

Lacerta pityusensis torretensis Buchholz, 1954
Holotype: ZFMK 12408, paratypes in ZFMK and MTKD
Origin: Torretas Id., Balearic Ids., Spain

Current name: Podarcis pitvusensis torretensis.

Lacerta rudis bischoffi Bohme & Budak, 1977
Paratypes in ZFMK and SZE, hoilotype in SZE
Origin: NE Turkey
Current name: Darevskia rudis bischoffi.
©ZFMK

Type specimens in ZFMK: Section of Herpetology 101

Lacerta rudis chechenica Eiselt & Darevsky, 1991
Paratypes in ZFMK and ZISP, holotype in ZISP
Origin: Chechnya-Ingushetia, Russia

Current name: Darevskia rudis chechenica.

Lacerta sapphirina Schmidtler, Darevsky & Eiselt, 1994
Paratypes in ZFMK, ZDSM, CAS and NMW, holotype
in ZSM

Origin: E Turkey

Current name: Darevskia sapphirinica.

+ Lacerta siculimelitensis Bbhme & Zammit-Maempel,
1983

Paratypes in ZFMK and Coll. Zammit-Maempel, holoty-
pe in NUNHM

Origin: Malta

Current name: “Lacerta” (inc. sed.) siculimelitensis.

Lacerta trilineata pamphylica Schmidtler, 1975
Paratypes in ZFMK, ZSM and NMW, holotype in ZSM
Origin: S Turkey

Current name: Lacerta pamphylica.

Lacerta uzzelli Darevsky & Danielyan, 1977
Paratypes in ZFMK, CAS and ZISP, holotype in CAS
Origin: NE Turkey

Current name: Darevskia uzzelli.

Lacerta viridis guentherpetersi Rykena, Nettmann &
Mayer, 2002

Paratypes in ZFMK, NMW, MTKD and ZMB, holotype
in NVW

Origin: Evvia Id., Greece

Lacerta viridis paphlagonica Schmidtler, 1986
Paratypes in ZFMK and ZSM, holotype in ZSM
Origin: N Turkey

Nucras caesicaudata Broadley, 1972
Paratypes in ZFMK and UM, holotype in UM
Origin: Zimbabwe

Ophiops macrodactylus Berthold, 1842
Holotype: ZDFMK 21219, no paratypes

Origin: W Turkey

Current name: Ophisops elegans macrodactylus.

Podarcis dugesii jogeri Bischoff, Osenegg & Mayer, 1989
Holotype: ZFMK 44193, paratypes in ZFMK, SMF and
BMNH

Origin: Porto Santo, Madeira Ids., Portugal

Current name: Jeira dugesii jogeri.

Bonn zoological Bulletin 59: 79-108

Podarcis dugesii selvagensis Bischoff, Osenegg & May-
er, 1989

Holotype: ZFMK 41561, paratypes in ZFMK, SMF and
BMNH

Origin: Selvagens Ids., Portugal

Current name: Teira dugesii selvagensis.

Podarcis lilfordi carbonerae Perez Mellado & Salvador,
1988

Holotype: ZFMK 11985, paratypes in ZFMK

Origin: Carbonera Id., Balearic Ids., Spain

Podarcis lilfordi codrellensis Perez Mellado & Salvador,
1988

Paratypes in ZFMK, and MNCN, holotype in MNCN
Origin: Codrell Id.,Balearic Ids., Spain

Takydromus (Platyplacopus) kuehnei vietnamensis Zieg-
ler & Bischoff, 1999

Holotype: ZFMK 70342, paratypes in ZFMK

Origin: Vietnam

Zootoca vivipara carniolica Mayer, BOhme, Tiedemann
& Bischoff, 2000
Holotype: ZFMK 68438, paratypes in NMW and MCSNT
Origin: Slovenia

Zootoca vivipara louislantzi Arribas, 2009

Paratypes in ZFMK, MNCN, NMW and Coll. O. Arribas,
holotype in MNCN

Origin: N Spain

Gerrhosauridae

Gerrhosaurus nigrolineatus ahlefeldti Hellmich &
Schmelcher, 1956

Paratypes in ZFMK, ZMH and ZSM, holotype in ZSM
Origin: Angola

Zonosaurus brygooi Lang & Bohme, 1990
Holotype: ZFMK 46789, paratypes in ZFMK and ZSM
Origin: NW Madagascar

Zonosaurus madagascariensis haraldmeieri Brygoo &
Bohme, 1985

Holotype: ZFMK 19335, paratypes in ZFMK and NU W
Origin: N Madagascar

Current name: Zonosaurus haraldmeieri.

OZFMK

102 Wolfgang Bohme

Scincidae

Ablepharus budaki anatolicus Schmidtler, 1997
Paratypes in ZFMK, ZSM, NMW and ZDEU, holotype
in ZSM

Origin: S Turkey

Ablepharus chernovi eiselti Schmidtler, 1997
Paratypes in ZFMK, ZSM, ZDEU and NMW
Origin: S Turkey

Chalcides chalcides mertensi Klausewitz, 1954
Paratypes in ZFMK, SMF and ZMB, holotype in SMF
Origin: Morocco

Current name: Chalcides mertensi.

Chalcides ghiarai Caputo & Mellado, 1992

Holotype: ZFMK 34549, paratypes in ZFMK, MZUF and
Coll. Mellado

Origin: Chafarinas Ids., Morocco

Current name: Chalcides parallelus.

Chalcides simonyi Steindachner, 1891
Syntypes: ZFMK 53098—100, further syntypes in NMW
Origin: Fuerteventura, Canary Ids., Spain

Cophoscincopus greeri Bohme, Schmitz & Ziegler,
2000

Holotype: ZFMK 57599, paratypes in ZFMK, BMNH,
FMNH. MNHN, RMNH, MCZ and ZMB

Origin: SE Guinea

Corucia zebrata alfredschmidti G. Kohler, 1997
Paratypes in ZFMK, SMF and MTKD, holotype in SMF
Origin: Bougainville, Solomon Ids., Papua New Guinea

Eugongylus haraldmeieri Bohme, 1976
Holotype: ZFMK 15888, paratype in ZFMK
Origin: New Caledonia

Current name Geoscincus haraldmeieri.

Eumecia anchietae wittei Laurent, 1964

Paratypes in ZFMK, IRSNB, RMNH and CM, holotype
in IRSNB

Origin: D.R. Congo

Isopachys borealis Lang & Bohme, 1990
Holotype: 45709, paratypes in ZFMK
Origin: Thailand

Lankascincus taylori Greer, 1991

Paratypes in ZFMK, BMNH, CM, FMNH and USNM,
holotype in BMNH

Origin: Sri Lanka

Bonn zoological Bulletin 59: 79-108

Leiolopisma greeri Bohme, 1979
Holotype: ZFMK 25456, no paratypes
Origin: New Caledonia

Current name: Lioscincus greeri.

Lepidothyris hinkeli Wagner, Bohme, Pauwels &
Schmitz, 2009

Holotype: ZFMK 55701, paratypes in ZFMK, IRSNB and
MHNG

Origin: Rwanda

Lepidothyris hinkeli joei Wagner, B6hme, Pauwels &
Scgmitz, 2009

Holotype: ZFMK 64410, paratypes in MHNG and
IRSNB

Origin: People’s Rep. Congo

Lygosoma boehmei Ziegler, Schmitz, Heidrich, Vu &
Nguyen Q.T. (2007)

Holotype: ZFMK 86359, no paratypes

Origin: Vietnam

Lygosoma meleagris hackarsi De Witte, 1941
Paratypes in ZFMK and IRSNB

Origin: Kivu, D.R. Congo

Current name: Leptosiaphos hackarsi

Mabuya macrophthalma Mausfeld & Bohme, 2002
Holotype: ZFMK 71717, paratype in ZFMK
Origin: Java, Indonesia

Current name: Eutropis macrophthalma

Mabuya perroteti upembae De Witte, 1953

Paratypes in ZFMK, IRSNB, MCZ, BMNH, MZB/UM,
HUJ, CM, RMNH, holotype in IRSNB

Origin: D.R. Congo

Current name: 7rachylepis perroteti upembae

Mabuya quinquetaeniata langheldi Sternfeld, 1917
Paratypes in ZFMK and ZMB, holotype in ZMB
Origin: Cameroon

Current name: Trachylepis langheldi

Mabuya quinquetaeniata pulcherrima De Witte, 1953
Paratypes in ZFMK, IRSNB, BMNH and CM, holotype
in IRSNB

Origin: D.R. Congo

Current name: 7rachylepis pulcherrima

Mabuya quinquetaeniata scharica Sternfeld, 1917
Paratypes in ZFMK and ZMB, holotype in ZMB
Origin: Chad

Current name: Trachylepis quinquetaeniata scharica.

©ZFMK

Type specimens in ZFMK: Section of Herpetology 103

Panaspis chriswildi Bohme & Schmitz, 1996
Holotype: ZFMK 59537, no paratype

Origin: Cameroon

Current name: Lacertaspis chriswildi.

Panaspis ianthinoxantha Bohme, 1975
Holotype: ZFMK 15349, paratypes in ZFMK
Origin: NW Cameroon

Current name: Leptosiaphos ianthinoxantha

Riopa erythropleuron Mertens, 1968
Holotype: ZFMK 5814, no paratypes
Origin: W Cameroon

Current name: Leptosiaphos pauliani

Sphenomorphus aquaticus Malkmus 1991
Holotype: ZFMK 52214, paratypes in ZFMK
Origin: Sabah, Malaysia

Current name: 7ropidophorus beccarii.

Sphenomorphus buenloicus Darevsky & Nguyen V.S.,
1983

Paratypes in ZFMK and ZISP, holotype in ZISP

Origin: S Vietnam

Sphenomorphus rufocaudatus Darevsky & Nguyen
V.S., 1983

Paratypes in ZFMK and ZISP, holotype in ZISP

Origin: S Vietnam

Current name: Scincella rufocaudata.

Trachylepis dichroma Ginther, Whiting & Bauer, 2005
Paratypes in ZFMK and ZMB, holotype in ZMB
Origin: Tanzania

Trachylepis loluiensis Kingdon & Spawls, 2010
Holotype: ZFMK 89371, paratypes in ZFMK and in Coll.
Kingdon

Origin: Lolui Id., Lake Victoria, Uganda

Tropidophorus boehmei Nguyen Q.T., Nguyen T.T.,
Schmitz, Orlov & Ziegler, 2010

Paratypes in ZFMK, IEBR, VNMN, MHNG, ZISP, and
VNUH, holotype in VNMN

Origin: N Vietnam

Tropidophorus noggei Ziegler, Vu & Bui, 2005
Holotype: ZFMK 83668, paratypes in ZFMK and VNUH
Origin: Vietnam

Typhlacontias punctatissimus brainei Haacke, 1997

Paratypes in ZFMK, TM and SMWN, holotype in TM
Origin: Namibia

Bonn zoological Bulletin 59: 79-108

Anguidae

Ophiodes intermedius Boulenger, 1894
Syntypes 26407, further syntypes in BMNH
Origin: Paraguay

Ophisaurus apodus thracius Obst, 1978

Paratypes in ZFMK and MTKD, holotype in MTKD
Origin: Bulgaria

Current name: Pseudopus apodus thracius.

Varanidae

Monitor doreanus Meyer, 1874
Neotype: ZFMK 52922

Origin: W Papua, Indonesia
Current name: Varanus doreanus.

Stellio salvator Laurenti, 1768
Neotype: ZFMK 22092

Origin: Sri Lanka

Current name: Varanus salvator.

Tupinambis indicus Daudin, 1802
Neotype: ZFMK 70650

Origin: Ambon Id., Moluccas, Indonesia
Current name: Varanus indicus.

Varanus boehmei Jacobs, 2003
Holotype: ZFMK 77837, paratypes in ZFMK and ZMA
Origin: Waigeo Id., W Papua, Indonesia

Varanus caerulivirens Ziegler, Bohme & Philipp, 1999
Holotype: ZFMK 68874, paratypes in ZFMK and SMF
Origin: Halmahera Id., Moluccas, Indonesia

Varanus cerambonensis Philipp, BOhme & Ziegler,
1999

Holotype: ZFMK 70617, paratypes in ZFMK, RMNH and
ZMB

Origin: Seram Id., Moluccas, Indonesia

Varanus cumingi samarensis Koch, Gaulke & Bohme,
2010

Holotype: ZFMK 64713, paratype in ZFMK

Origin: Samar Id. Philippines

Varanus doreanus finschi Bohme, Horn & Ziegler, 1994
Holotype: ZFMK 26347, paratypes in ZFMK and ZMB
Origin: New Britain Id., Papua New Guinea

Current name: Varanus finschi.

©ZFMK

104 Wolfgang Bohme

Varanus juxtindicus Bohme, Philipp & Ziegler, 2002
Paratypes in ZFMK and ZMUC, holotype in ZMUC
Origin: Rennell Id., Solomon Ids.

Varanus lirungensis Koch, Arida, Schmitz, Bohme &
Ziegler, 2009

Paratypes in ZFMK, MZB and ZMA, holotype in MZB
Origin: Salibabu Id., Talaud Ids., Indonesia

Varanus macraei Bohme & Jacobs, 2001
Holotype: ZFMK 74558, paratypes in ZFMK and SMF
Origin: Batanta Id., West Papua, Indonesia

Varanus melinus Bohme & Ziegler, 1997
Holotype: ZFMK 65737, paratypes in ZFMK
Origin: Sula Ids., Indonesia

Varanus palawanensis Koch, Gaulke & Bohme, 2010
Paratypes in ZFMK and SMF, BMNH, MNHN and
ZMUC, holotype in SMF

Origin: Palawan, Philippines

Varanus panoptes horni Bohme, 1988

Holotype: ZFMK 19290, paratypes in ZFMK, ZMA and
ZSM

Origin: W Papua, Indonesia

Varanus rainerguentheri Ziegler, Bohme & Schmitz,
2007

Holotype: ZFMK 85404, paratype in USNM

Origin: Halmahera Id., Moluccas, Indonesia

Varanus rasmusseni Koch, Gaulke & Bohme, 2010
Paratype in ZFMK, holotype in ZMUC
Origin: Tawi Tawi UID., Sulu Ids., Indonesia

Varanus yemenensis Bohme, Joger & Schatti, 1989
Holotype: ZFMK 46500, paratypes in ZFMK, MNHG and
BMNH

Origin: Yemen

Typhlopidae

Rhinotyphlops debilis Joger, 1990
Holotype: ZFMK 33684, paratype in HLMD
Origin: Central African Republic

Current name: Letheobia debilis.

Rhinotyphlops episcopus Franzen & Wallach, 2002

Holotype: ZFMK 74224, paratypes in ZFMK and ZSM
Origin: E Turkey

Bonn zoological Bulletin 59: 79-108

Boidae

Boa carinata J.G. Schneider, 1801
Lectotype: ZFMK 35503

Origin: Ambon Id., Moluccas, Indonesia
Current name: Candoia carinata.

Pythonidae

Boa reticulata J.G. Schneider, 1801
Neotype: ZFMK 32378

Origin: Peninsular Malaysia
Current name: Python reticulatus.

Python bivittatus progschai Jacobs, Auliya & Bohme,
2009

Holotype: ZFMK 87841, paratypes in ZFMK

Origin: Sulawesi, Indonesia

Python reticulatus jampeanus Auliya, Mausfeld, Schmitz
& Bohme, 2002

Holotype: ZFMK 73475, paratypes in ZFMK

Origin: Tanajampea Id., Indonesia

Python reticulatus saputrai Auliya, Mausfeld, Schmitz
& Bohme, 2002

Holotype: ZFMK 73473, paratype in ZFMK

Origin: Selayar Id., Indonesia

Calamariidae

Calamaria thani Ziegler & Le, 2005
Holotype: ZFMK 82920, no paratypes
Origin: Vietnam

Colubridae

Ablabes tessellatus Berthold, 1859
Syntypes: ZFMK 36913—914

Origin: Suriname

Currentt name: Mastigodryas boddaerti.

Boiga bourreti Tillack, Ziegler & Le, 2004
Holotype: ZFMK 82921, no paratypes
Origin: Vietnam

Calamaria degenhardtii Berthold, 1845
Holotype: ZFMK 36828, no paratypes
Origin: “Provinz Popayan”, Colombia
Current name: Stenorrhina degenhardtii.

©ZFMK

Type specimens in ZFMK: Section of Herpetology 105

Coluber florulentus perreti Schatti, 1988

Paratypes in ZFMK, BMNH and USNM, holotype in
MNHG

Origin: N Cameroon

Current name: Platyceps florulentus perreti.

Coluber gemonensis gyarosensis Mertens, 1968
Holotype: ZFMK 4332, patratypes in ZFMK and SMF
Gyaros Id., Cyclades, Greece

Current name: Hierophis viridiflavus.

Coluber najadum kalymnensis B. Schneider, 1979
Holotype: ZFMK 24933, paratype in Coll. B. Schneider
Origin: Kalymnos Id. Dodekanes, Greece

Current name: Platyceps najadum kalymnensis.

Coluber rubriceps thracius Rehak, 1985

Paratypes in ZFMK and DZCHU, holotype in DZCHU
Origin: Bulgaria

Current name: Platyceps collaris.

Coluber schmidtleri Schatti & McCarthy, 2001
Paratypes in ZFMK, BMNH, ZISP, holotype in FMNH
Origin: Iran

Current name: Platyceps schmidtleri.

Dispholidus typus kivuensis Laurent, 1955

Paratypes in ZFMK, IRSNB, CM and RMNH, holotyxpe
in MRAC

Origin: Kivu, D.R. Congo

Current name Dispholidus typus.

Dispholidus typus punctatus Laurent, 1955

Paratypes in ZFMK, IRSNB, CM, FMNH and RMNH,
holotype in MRAC

Origin: D.R. Congo

Current name: Dispholidus typus.

Eirenis barani Schmidtler, 1988
Paratypes in ZFMK, ZSM and NMW, holotype in ZSM
Origin: Turkey

Eirenis barani bischofforum Schmidtler, 1997
Paratypes in ZFMK, ZMR and ZSM, holotype in ZSM
Origin: Turkey

Ejirenis levantinus Schmidtler, 1993

Paratypes in ZFMK, NMW, MHNG and SZE, holotype
in ZSM

Origin: S Turkey

Eirenis modestus cilicius Schmidtler, 1993

Paratypes in ZFMK, NMW, MNHG and ZSM, holotype
in ZSM

Origin: S Turkey

Bonn zoological Bulletin 59: 79-108

Eirenis punctatolineatus kumerloevei Eiselt, 1970
Holotype: ZFMK 5990, no paratypes
Origin: Akdamar Id., Lake Van, E Turkey

Elaphe porphyracea laticincta Schulz & Helfenberger,
1998

Paratypes in ZFMK, MHNG and CAS, holotype MHNG
Origin: Peninsular Malaysia

Current name: Oreocryptophis porphyraceus laticinctus.

Elaphe quatuorlineata parensis Cattaneo, 1999
Paratypes in ZFMK and in Coll. Cattaneo, holotype in
Coll. Cattaneo

Origin: Paros Id., Cyclades, Greece

Lycodon ruhstrati abditus Vogel, David, Pauwels,
Sumontha, Norval, Hendrix, Vu & Ziegler, 2009
Holotype in ZFMK, paratypes in ZFMK, MNHN and
ZMB

Origin: Vietnam

Macrocalamus schulzi Vogel & David, 1999
Holotype: ZFMK 51159, paratype in ZFMK
Origin: Peninsular Malaysia

Orthriophis taeniurus callicyanous Schulz, 2010
Holotype: ZFMK 81450, paratype in ZFMK
Origin: Vietnam

Platyceps saharicus Schatti & McCarthy, 2004
Paratypes in ZFMK, BMNH, FMNH, HUJ, MCZ, MH-
NG, MZUF, NMW, and TAU, holotype in FMNH
Origin: Egypt

Telescopus fallax christianus Buchholz, 1955
Holotype: ZFMK 9233, paratype in ZFMK
Orgin: Christiana Id., Cyclades, Greece
Current name: Telescopus fallx pallidus.

Zamenis andreana Werner, 1917

Syntype: ZFMK 31600, further syntype n CAS
Origin: Fars, Iran

Current name: “Coluber” (s.1.) andreanus.

Lamprophiidae
Pseudoboodon boehmei Rasmussen & Largen, 1992

Paratypes in ZFMK and BMNH, holotype in BMNH
Origin: SW Ethiopia

©ZFMK

106 Wolfgang Bo6hme

Psammophiidae

Malpolon monspessulanus saharatlanticus Geniez,
Cluchier & De Haan, 2006

Paratypes in ZFMK and BEV, holotype in MNHN
Origin: SW Morocco

Psammophis elegans univittatus Perret, 1961
Paratypes in ZFMK and MHNG, holotype in MHNG
Origin: N Cameroon

Psammophis sibilans rukwae Broadley, 1966
Paratypes in ZFMK and NMSR, holotype in NMSR
Origin: S Tanzania

Current name: Psammophis rukwae.

Psammophis sibilans trinasalis Werner, 1902
Lectotype: ZFMK 30503, paralectotypes in NUW
Origin: Namibia

Current name: Psammophis leightoni trinasalis.

Rhamphiophis acutus wittei Laurent, 1956

Paratypes in ZFMK, ASU, CM, FMNH, LSUMZ, IRSNB
and NMZB/UM, holotype in MRAC

Origin: D.R. Congo

Current name: Psammophylax acutus acutus.

Dipsadidae

Apostolepis multicincta Harvey. 1999
Paratypes in ZFMK and NKA, holotype in NKA
Origin: Bolivia

Atractus paravertebralis Henle & Ehrl, 1991
Holotype: ZFMK 39705, no paratype
Origin: S Peru

Clelia langeri Reichle & Embert, 2005
Paratypes in ZFMK & NKA, holotype in NKA
Origin: Bolivia

Duberria ancoralis Berthold, 1842
Holotype: 36992, no paratypes

Origin: “Surinam” (in error)

Current name: Farancia erythrogramma.

Liophis miliaris intermedius Henle & Ehrl, 1981
Holotype: ZFMK 34243, paratypes in ZFMK
Origin: S Peru

Current name: Liophis reginae.

Psammophis molochina Berthold, 1846
Holotype: ZFMK 36923, no paratypes
Origin: “Brasilien”

Current name: Philodryas nattereri.

Bonn zoological Bulletin 59: 79-108

Homalopsidae

Enhydris gyii Murphy, Voris & Auliya, 2005
Holotype: ZFMK 65824, paratypes in ZFMK and ANSP
Origin: N Kalimantan, Indonesia

Natricidae

Amphiesma andreae Ziegler & Le, 2006
Holotype: ZFMK 83747, no paratypes
Origin: N Vietnam

Amphiesma leucomystax David, Bain, Nguyen Q.T.,
Orlov, Vogel Vu & Ziegler, 2007

Holotype: ZFMK 71702, paratypes in ZFMK, ZISP,
MNHN, IEBR, AMNH and FMNH

Origin: Vietnam

Natrix megalocephala Orlov & Tuntyev, 1987
Paratypes in ZFMK and ZISP, holotype in ZISP
Origin: Abchasia, Georgia

Current name: Natrix natrix scutata.

Tropidonotus chinensis Berthold, 1859
Holotype: ZFMK 33443, no paratypes
Origin:China”

Current name: Sinonatrix annularis.

Tropidonotus tigrinus Boie, 1826
Syntypes: ZFMK 8406-407

Origin: Japan

Current name: Rhabdophis tigrinus.

Elapidae

Elaps bertholdi Jan, 1859

Holotype: ZFMK 36402, no paratype
Origin: Australia

Current name: Simoselaps bertholdi.

Naja melanoleuca subfulva Laurent, 1955

Paratypes in ZFMK, IRSNB and FMNH, holotype in
MRAC

Origin: D.R. Congo

Current name: Naja melanoleuca.

Naja nigricollis atriceps Laurent, 1955

Paratypes in ZFMK and IRSNB, holotype in MRAC
Origin: D.R. Congo

Current name: Naja nigricollis.

Naja nubiae Wiister & Broadley, 2003
Paratypes in ZFMK, BMNH, MHNG, MNHN and

©ZFMK

Type specimens in ZFMK: Section of Herpetology 107

NHRM, holotype in MRAC
Origin: Sudan

Hydrophiidae

Hydrophis sibauensis Rasmussen, Auliya & Béhme, 2001
Holotype: ZFMK 64891, paratypes in ZMUC and MZB
Origin: N Kalimantan, Indonesia

Viperidae: Viperinae

Bitis parviocula Bohme, 1977
Holotype: ZFMK 16803, no paratypes
Origin: Ethiopia

Cerastes boehmei Wagner & Wilms, 2010
Holotype: ZFMK 58054, no paratypes
Origin: Tunisia

Echis multisquamatus Tcherlin, 1981
Paratypes in ZFMK and ZISP, holotype in ZISP
Origin: Turkmenia

Current name: Echis carinatus multisquamatus.

Vipera barani Bohme & Joger, 1983
Holotype: ZFMK 35444, no paratypes
Origin: N Turkey

Vipera wagneri Nilson & Andrén, 1984
Holotype: ZFMK 32495, no paratypes
Origin: NW Iran

Current name: Montivipera wagneri.

Viperidae: Crotalinae

Agkistrodon halys boehmei Nilson, 1983
Holotype: ZFMK 8648, no paratype
Origin: Afghanistan

Current name: Gloydius halys boehmei.

Triceratolepidophis sieversorum Ziegler, H.W. Her-
rmann, David, Orlov & Pauwels, 2000

Holotype: ZFMK 71262, no paratype

Origin: Vietnam

Current name: Protobothrops sieversorum.

Trigonocephalus schlegelii Berthold, 1845
Holotype: ZFMK 32554, no paratypes
Origin: “Provinz Popayan”, Colombia
Current name: Bothriechis schlegelii.

Trimeresurus fucatus Vogel, David & Pauwels, 2004
Paratypes in ZFMK, BMNH, IRSNB, MNHN, QSMI,
ZRC, ZSM and Coll. G. Vogel, holotype in MNHN
Origin: Thailand

Bonn zoological Bulletin 59: 79-108

Trimeresurus gumprechti David, Vogel, Pauwels & Vi-
dal, 2002

Paratypes in ZFMK, MNHN, PSUAA, and RFI, holotype
in MNHN

Origin: Thailand

Trimeresurus nebularis Vogel, David & Pauwels, 2004
Paratypes in ZFMK, IRSNB, MNHN, ZRC and Coll. G.
Vogel

Origin: Thailand

Trimeresurus vogeli David, Vidal & Pauwels, 2001
Paratype in ZFMK, holotype in MNHN
Origin: Thailand

CROCODYLIA
Crocodylidae

Crocodylus porosus J.G. Schneider, 1801
Paralectotypes: ZFMK 40955—956, lectotype in ZMB
Origin: Java, Indonesia.

Acknowledgements. I am indebted to my colleagues Michael
Franzen and Frank Glaw (ZSM) as well as Jorn Kohler (HLMD)
and Thomas Ziegler (Cologne Zoo), and also Philipp Wagner and
Nguyen Quang Truong (currently ZFMK) for reading parts of
the type list. Cordial thanks are again due to Ursula Bott (ZFMK)
for her constant help and cooperation, especially in regard to
ZFMK’s herpetological collection and the type catalogue.

REFERENCES

Bohme W (1974) Die Typusexemplare der Herpetologischen
Sammlung des Museums Alexander Koenig, Bonn. Bonner
zoologische Beitrage 25: 165-176

Bohme W, Bischoff W (1984) Reptilien und Amphibien in:
Rheinwald G (ed.) Die Wirbeltiersammlungen des Museums
Alexander Koenig. Bonner zoologische Monographien 19:
151-213

Bohme W, Sander PM (2010) Sauropsida (without birds) in:
Westheide W, Rieger R (eds.) Spezielle Zoologie, Teil 2: Wir-
bel- oder Schadeltiere. Heidelberg (Spectrum Verl.), 2°4 edi-
tion, pp. 362-421

Frost DR, Grant T, Faivovich J, Bain RH, Haas A, Haddad CFB,
De Sa RO, Channing A, Wilkinson M, Donnellan SC, Rax-
worthy CJ, Campbell JA, Blotto BL, Moler P, Drewes RC,
Nussbaum RA, Lynch JD, Green DM, Wheller WC (2006)
The amphibian tree of liefe. Bulletin of the American Muse-
um of Natural History 297: 1-370

Glaw F, Vences M (2007) A field guide to the amphibians and
reptiles of Madagascar, K6In (Vences & Glaw Verlags-GbR),
3'd edition, 495 pp.

Grant T, Frost DR, Caldwell JP, Gaghardo R, Haddad CFB, Kok
PJR, Means DB, Noonan BP, Schargel WE, Wheeler WC
(2006) Phylogenetic systematics of dart-poison frogs and their
relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin
of the American Museum of Natural History 299: 1-262

©ZFMK

108 Wolfgang Bohme

Lotters S, Jungfer KH, Henkel FW, Schmidt W (2007) Pfeilgift- vanced snakes (Serpentes, Caenophiodia) with an emphasis
frésche. Biologie, Haltung, Arten. Chimaira, Frankfurt, 668 on South American xenodontines: a revised classification and
pp. desceription of new taxa. Papéis Avulsos de Zoologia 49:

Zaher H, Gobbi Grazziotini F, Cadle JE, Murphy RW, Moura- 115-153

Leite JC de, Bonatto SL (2009) Molecular phylogeny of ad-

Bonn zoological Bulletin 59: 79-108 ©ZFMK

Volume 59

Bonn zoological Bulletin

pp. 109-136 Bonn, December 2010

Catalogue of type specimens of fishes in the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Fabian Herder!2, Jan Huylebrouck! & Klaus Busse!
'Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn
?Corresponding author. E-mail: f-herder.zfmk(@uni-bonn.de.

Abstract. Here we provide a checklist of type specimens housed in the fish collection of the Zoologisches Forschungsmu-
seum Alexander Koenig, Bonn (ZFMK). The catalogue includes type locality and sampling data, lists type material of
the taxa represented in ZFMK deposited in other collections, provides remarks on the current status of the material, and
covers the relevant bibliography. In total, 126 species are represented, 80 of these by holo- or lectotypes.

Key Words. Type specimens, taxonomy, fish, ichthyology, type catalogue, ZFMK Bonn.

INTRODUCTION

The ichthyology section of the Zoologisches
Forschungsmuseum Alexander Koenig was founded in
1954, and currently contains more than 41,000 fish spec-
imens representing > 1500 species. The latter number is
however based on the current state of ongoing digitaliza-
tion, and is most likely a rather serious underestimation
of the collection’s inventories. Among other material, the
collection contains material collected by the museum
founder A. Koenig since 1879, specimens obtained from
South America by the institute’s first fish curator K. H.
Liiling, collections of Indian fish from marine and brack-
ish habitats collected by G. A. von Maydell in the 1950s,
and important collections of freshwater fish from Central
Vientnam collected by J. Freyhof and colleagues at the be-
ginning of the last decade. The name-bearing types in the
ichthyology section partially reflect the collection focus,
composed in prominent parts by e.g. Asian loaches, South
American characins, and catfishes from both regions. The
ZFMkK collection also houses substantial numbers of kil-
lifish types from Africa and South America, most desig-
nated either by L. Seegers or R. Sonnenberg.

The present type catalogue includes, besides of taxon
names and collection references, in brief some crucial da-
ta from the original descriptions and the current status at
time of publication; these include also additional type ma-
terial housed in other collections.

Bonn zoological Bulletin 59: 109-136

MATERIAL AND METHODS

The order of fish families in the present catalogue wide-
ly follows Eschmeyer’s (1998) “Catalog of Fishes”. With-
in families, species appear in alphabetical order of the
genus as used in the original description. If not stated oth-
erwise, taxon names are valid as originally published,
widely based on the online database of the “Catalog of
Fishes” (http://research.calacademy.org/ichthyology/cat-
alog/fishcatmain.asp) in November 2010. The original ref-
erence is provided in brief for each single taxon. As far
as available in the description, the present catalogue in-
cludes besides of collection numbers also data on size, sex,
locality (in increasing order, 1.e. from habitat data to coun-
try name), coordinates, collection date and collector(s). In
addition, we included for fish taxa represented by types
in ZFMK also the relevant data of other type material, in-
cluding collection and number (if available), and in most
cases the same relevant data as outlined above. If not stat-
ed otherwise, size data refer to mm standard length. Num-
bers in parentheses following collection numbers refer to
the number of specimen in the respective series or lot. Da-
ta on sex refer to the composition of individuals as orig-
inally published; specific data describing the amount of
males and females within certain series are not included.
As far as possible, locality data were translated to Eng-
lish; coordinates are provided as they appear in the orig-
inal description.

©ZFMK

110 Fabian Herder, Jan Huylebrouck & Klaus Busse

Abbreviations for museum collections used:

AM = Albany Museum, Grahamstown, Zambia

AMNH = American Museum of Natural History, New
York, USA

AM PF = Albany Museum, Grahamstown, Zambia
ANSP = Academy of Natural Sciences, Philadelphia, USA
BM = British Museum (Natural History), London, UK
BMNH = Natural History Museum (formerly British Mu-
seum (Natural History)), London, UK

CAS = California Academy of Sciences, San Francisco,
USA

CAZ = Collection Axel Zarske, Dresden, Germany
CENAREST = Research station in Batoke, Cameroon
CIRA UTB = Centro de Investigation de Recursos Aquati-
cos, Universidad Técnica del Beni, Trinidad, Bolivia
CMK = Collection Maurice Kottelat, Cornol, Switzerland
FMNH = Field Museum of Natural History, Chicago, USA
INPA = Instituto Nacional de Pesquiasas da Amazonia,
Manaus, Brazil

IRET = Institut de Recherche en Ecologie Tropicale, Li-
breville, Gabon

IRSNB = Institut royal des Sciences naturelles de Belgi-
que, Bruxelles, Belgium

IBTS = Taxonomy Research Group, Research Centres in
Biology Field, University of Bucharest, Tr. Savulescu,
Bucharest, Romania (formerly: ISBB)

IZUA = Instituto de Zoologia, Universidad Austral de
Chile, Valdivia, Chile

KU = Museum of Natural History, Lawrence, Kansas,
USA

MCP = Pontificia Universidade Catolica do Rio Grande
do Sul, Museu de Ciéncias e Tecnologia, Rio Grande do
Sul, Porto Alegre, Brazil

MCZ = Museum of Comparative Zoology, Harvard Uni-
versity, Cambridge, Massachusetts, USA

MGAB = Muzeul de Istorie Naturala “Grigore Antipa’,
Bucharest, Romania

MHNG = Muséum d’Histoire naturelle de Geneve,
Geneva, Switzerland

MNHN(P) = Muséum national d’Histore naturelle, Paris,
France

MNHNC = Museo Nacional de Historia Natural, Santia-
go, Chile

MRAC = Musée Royal de I’Afrique Centrale, Tervuren,
Belgium

MTD = Staatliche Naturhistorische Sammlungen, Muse-
um ftir Tierkunde, Dresden, Germany

MTD F = Staatliche Naturhistorische Sammlungen, Mu-
seum ftir Tierkunde, Dresden, Germany

MZUSP = Museu de Zoologia da Universidade de Sao
Paulo, Sao Paulo, Brazil

NHMW = Naturhistorisches Museum, Vienna, Austria
NMW = Naturhistorisches Museum Wien, Vienna, Aus-
tria

Bonn zoological Bulletin 59: 109-136

NRM = Naturhistoriska riksmuseet, Stockholm, Sweden
RMNH = Rijksmuseum van Natuurlijke Histoire, now Na-
turalis - National Natuurhistorisch Museum, Leiden, the
Netherlands

RUSI = J.L.B. Smith Institute of Ichthyology, Graham-
stown, South Africa

SMF = Senckenberg Museum, Frankfurt/Main, Germany
UMMZ = Museum of Zoology, University of Michigan,
Ann Arbor, USA

USNM = National Museum of Natural History, Smithson-
ian Institution, Washington, D.C., USA

ZFMK = Zoologisches Forschungsmuseum Alexander
Koenig, Bonn, Germany

ZMA = Zodlogisch Museum, Universiteit van Amsterdam,
the Netherlands

ZMB = Zoologisches Museum der Humboldt-Universi-
tat, Berlin, Germany

ZMH = Zoologisches Museum Hamburg, Germany
ZMM = Zoological Museum, Moscow State University,
Moscow, Russia

ZRC = Zoological Reference Collection, National Univer-
sity of Singapore, Singapore

ZSM = Zoologische Staatssammlung, Munich, Germany

KNERITIDAE

Kneria rukwaensis Seegers, 1995
Ichthyol Explor Freshwaters 6: 109

Paratypes: ZFMK 17477-17491 (15), 28.0-45.5 mm, 32
km northwest of Tunduma on road to Sumbawanga, af-
fluent of the Momba River, Chiwanda River at Chiwan-
da, western Lake Rukwa drainage, Mbeya Region, west-
ern Tanzania, 09°10’S 32°34’E, 29 Jul 1992, U. Renninger
& L. Seegers.

Type material in other collections: Holotype: ZMB
32352, male, 50.1 mm, same data as ZFMK 17477-17491;
Paratypes: ZMB 32353 (3), 33.7-52.9 mm, same data as
holotype; ZMB 32354 (5), 32.7-62.3 mm, same locality,
30 Jun 1989, U. Renninger & L. Seegers; ZMH 8400 (11),
19.4-42.6 mm, small brook, crossing road from Tundu-
ma to Sumbawanga 58 km northwest of Tunduma, west-
ern Lake Rukwa drainage, Mbeya Region, western Tan-
zania, 30 Jul 1989, U. Renninger & L. Seegers; ZMB
32355 (2), 46.0-51.5 mm; BMNH 1993.3.29: 7-10 (4),
27.0-38.3 mm, same locality, 27 Jul 1991, U. Renninger
& L. Seegers; MRAC 93-38-P-1—11 (11), 21.0-52.3 mm,
same locality, 29 Jul 1992, L. Seegers; ZMH 8501 (11),
22.5-63.0 mm, Sau River at Kisumba, 20 km south of
Sumbawanga on road to Mbala, Kalambo drainage, Ruk-
wa Region, western Tanzania, 24 Jul 1993, L. Seegers;
BMNH 1993.3.29: 1-6 (6), 17.8-36.7 mm, Luiche Riv-
er, some km north of Sumbawanga near junction Sum-

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 11]

bawanga-Mbala and Sumbawanga-Mpanda, western Lake
Rukwa drainage, Rukwa Region, western Tanzania, 2 Jul
1989, U. Renninger & L. Seegers; MRAC 93-38-P-12 (1),
34.0 mm, Mfwizi River, 110 km north of Sumbawanga
on road to Mpanda, northwestern Lake Rukwa drainage,
Rukwa Region, western Tanzania, 2 Jul 1989, U. Ren-
ninger & L. Seegers; MRAC 191042 (1), 39.9 mm, Sais
River at Lumi confluence, western Lake Rukwa drainage,
Mbala distr., northern Zambia, 15 Oct 1967, H. Matthes;
MRAC 93-38-P-13—27 (15), 17.5—61.1 mm, Sau River at
Kisumba, 20 km south of Sumbawanga on road to Mbala,
Kalambo drainage, Tanganyika basin, Rukwa Region,
western Tanzania, 24 Jul 1993, L. Seegers; MRAC 93-38-
P-28—38 (11), 17.0-35.0 mm, Pala River, 55 km south of
Sumbawanga at the road to Mbala, Kalambo drainage,
Tanganyika basin, Rukwa Region, western Tanzania, 24
Jul 1993, L. Seegers; MRAC 93-38-P-39-46 (8),
21.2—-29.0 mm, Kanyele River, 61 km south of Sumbawan-
ga on road to Mbala, Kalambo drainage, Tanganyika basin,
Rukwa Region, western Tanzania, 24 Jul 1993, L.
Seegers; MRAC 93-38-P-47-49 (3), 23.9-29.0 mm,
brook, 5 km north of the Tanzania/Zambia border on road
from Mbala to Sumbawanga, Kalambo drainage, Tan-
ganyika basin, Rukwa Region, western Tanzania, 24 Jul
1993, L. Seegers; RUSI uncat. (26), 16.9-35.4 mm, brook,
Luiche affluent, at the road from Muze to Sumbawanga,
4 km before reaching the main road from Mpanda to Sum-
bawanga at Ntendo, Lake Rukwa drainage, Rukwa Re-
gion, western Tanzania, 23 Jul 1993, L. Seegers; RUSI
50340 (2), 48.5-52.4 mm, Mfwizi River, 142 km south
of Mpanda on road from Mpanda to Sumbawanga, Lake
Rukwa drainage, Rukwa Region, western Tanzania, 6 Jul
1994, L. Seegers; ZMA 109763 (2), 32.9-36.0 mm,
Mwandwisi brook near Chipanda, Lake Rukwa drainage,
Abercorn-District (=Mbala), northern Zambia, 30 Jul
1966, H. Matthes; ZMA 109764 (11), 14.2—23.3 mm,
Mwaa-Rivulet near Kawimbi, Lake Rukwa drainage,
Abercorn-District (=Mbala), northern Zambia, 5 Jul
1966, H. Matthes; ZMA 109765 (437), 17.0-45.2 mm, Lu-
mi River near Kawimbi, Lake Rukwa drainage, Abercorn-
District (=Mbala), northern Zambia, 30 Jul 1966, H.
Matthes.

CYPRINIDAE

Rasbora rubrodorsalis Donoso-Biichner & Schmidt, 1997
Z Fischk 4: 90

Paratypes: ZFMK 18459-18460 (2), Cha Am, 208/209 km
main road 4, Thailand, Nov 1989, R. Donoso-Btichner;
ZFMK 18461 (1), Cha Am/Hua Hin, 212 km main road
4, Thailand, Nov 1989, A. Waser.

Bonn zoological Bulletin 59: 109-136

Type material in other collections: Holotype: ZMB
32549, same data as ZFMK 18459-18460; Paratypes:
ZMB 32550-32551, same data as ZFMK 18461; ZMB
32420-32422 (3), Nam Ngum Dam reservoir, 20 km
northeast of Vientiane, Laos, K. Lung & J. Topfer [no col-
lection date provided].

Tanichthys micagemmae Freyhof & Herder, 2001
Ichthyol Explor Freshwaters 12: 216

Holotype: ZFMK 39098, 18.8 mm, stream Bau Dung, a
tributary of River Ben Hai, Quang Binh Prov., Vietnam,
17°07°24”N 106°58712”E, 5 Apr 2000, J. Freyhof & F.
Herder.

Paratypes: ZFMK 39099-39118 (20), 13.4-23.1 mm, same
data as holotype.

BALITORIDAE

Hemimyzon ecdyonuroides Freyhof & Herder, 2002
Ichthyol Explor Freshwaters 13: 54

Holotype: ZFMK 22136, 41.7 mm, River Pako about 50
km north of Kontum, Kontum Prov., Vietnam, 14°39.60’N
107°46.98’E, 22 Feb 1999, J. Freyhof & D. V. Serov.

Paratypes: ZFMK 22137-22146 (10), 32.6-42.1 mm, same
data as holotype.

Type material in other collections: CMK 16011 (4),
38.1-41.5 mm, same data as holotype.

Nemacheilus banar Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 135

Holotype: ZFMK 28588, 65.8 mm, middle River Pako
about 50 km north of Kontum, Kontum Prov., Vietnam,
14°36.60’N 107°46.98’E, 22 Feb 1999, J. Freyhof & D.
Serov.

Patatypes: ZFMK 26528-26589 (60), 22.1-77.7 mm,
same data as holotype; ZFMK 26274-26278 (5), 38.1-43.4
mm, River Dakabla at Kontum, Kontum Prov., Vietnam,
14°20.73’N 108°00.03’E, 20 Feb 1999, J. Freyhof & D.
Serov; ZFMK 26322-26324 (3), 38.4-42.4 mm, River
Dakpne 30 km northeast of Kontum, Kontum Prov., Viet-
nam, 14°30.35’N 108°14.92’E, 20 Feb 1999, J. Freyhof
& D. Serov; ZFMK 28647, Stream Pako 15 km west of
Kontum, Kontum Prov., Vietnam, 14°23.23’N
107°54.15°E, 23 Feb 1999, J. Freyhof & D. Serov; ZFMK
26590, 64.5 mm, Stream Iasia 25 west of Kontum, Kon-

©ZFMK

112 Fabian Herder, Jan Huylebrouck & Klaus Busse

tum Proy., Vietnam, 14°23.63’N 107°48.64’E, J. Freyhof
& D. Serov; ZFMK 26591-26592 (2), 69.6-80.4 mm, Riv-
er Ba at An Khe, Gia Lai Prov., Vietnam, 13°57.23’N
108°39.15’E, 24 Feb 1999, J. Freyhof & D. Serov.

Type material in other collections: CMK 16009 (5), 34.1-
68.5 mm, same data as holotype.

Nemacheilus cleopatra Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 138

Holotype: ZFMK 27155, 68.9 mm, River Sol near Plao
Ganong, about 60 km south of Pleiku, Province Gia Lai,
Vietnam, 13°31.82’N 108°18.63’E, 19 Feb 1999, J. Frey-
hof & D. Serov.

Paratypes: ZFMK 20656-26081 (25), 38.4—75.3 mm, same
data as holotype.

Type material in other collections: CMK 16023 (3),
44.2—60.8 mm, same data as holotype.

Schistura antennata Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 140

Holotype: ZFMK 27934, 73.6 mm, stream at Son Kim,
Ha Tinh, Prov., Vietnam, 18°24.25’N 105°11.10°E, 11 Apr
2000, J. Freyhof, F. Herder & D. Serov.

Paratypes: ZFMK 27935-27941 (7), 33.7—70.3 mm, same
data as holotype;

Remarks: Additional type material in other collections:
CMK 16673 (3), 50.8—66.5 mm, same data as holotype.

Schistura bachmaensis Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 143

Holotype: ZFMK 27942, 44.3 mm, Stream Suoi Voi Strom
at Thua Luu, 55 km south of Hue, Thua Thien Hue Prov.,
Vietnam, 16°14.58’N 107°59.39’E, 19 Mar 2000, J. Frey-
hof, F. Herder & D. Serov.

Paratype: ZFMK 27942 (1), 49.0 mm, same data as holo-
type.

Type material in other collections: CMK 16674 (1), 56.6
mim, same data as holotype.

Schistura carbonaria Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 145

Bonn zoological Bulletin 59: 109-136

Holotype: ZFMK 26593, 57.3 mm, stream Khe Vin An at
Nam Dong village about 40 km south of Hue, Thua Thien
Hue Prov., Vietnam, 16°11.58’N 107°43.76’°E, 15 Mar
1999, J. Freyhof & D. Seroy.

Paratypes: ZFMK 26594-26653 (60), 25.8-58.5 mm, same
data as holotype; ZFMK 26654-26664 (11), 48.8-60.8
mim, River Rao Nho at Hong Ha village about 50 km west
of Hue, Thua Thien Hue Prov., Vietnam, 16°17.35’N
107°21.59°E, 16 Mar 1999, J. Freyhof & D. Serov; ZFMK
26665-26669 (5), 48.1-50.6 mm, stream Mau about 30
km west of Hue, Thua Thien Hue Prov., Vietnam,
16°19:97’N 107°25.29°E, 16 Mar 1999, J. Freyhotiép)
Serov.

Schistura dalatensis Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 150

Holotype: ZFMK 23906, 87.2 mm, River Dai Tan about
30 km south of Da Lat, Lam Dong Prov., Vietnam,
11°46,64°N 108°19,28’E, 21 Jan 1999, J. Freyhof & D.
Serov.

Paratypes: ZFMK 23641-23652 (12), 29.8-49.1 mm,
small stream running to River Da Dung south of Da Lat,
Prov. Lam Dong, Vietnam, 11°46.86’N 108°25.04’E, 19
Jan 1999, J. Freyhof & D. Serov; ZFMK 23759-23761
(3), 41.7-48.0 mm, River Da Dung 25 km south of Da Lat,
Lam Dong Prov., Vietnam, 11°45.48’N 108°23.78’E, 21
Jan 1999, J. Freyhof & D. Serov.

Type material in other collections: CMK 15999 (8),
30.8—-51.8 mm, 11°46.86’N 108°25.04’E, same data as
ZFMK 23641-23652.

Schistura huongensis Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 155

Holotype: ZFMK 26810, 57.2 mm, stream Keh Vinh An
at Nam Dong village about 40 km south of Hue, Thua
Thien Hue Prov., Vietnam, 16°11.58°N 107°43.76°E, 15
Mar 1999, J. Freyhof & D. Serov.

Paratypes: ZFMK 26811-26826 (16), 28.2-55.5 mm,
same data as holotype; ZFMK 26827-26848 (22),
30.8-56.9 mm, River Rao Nho at Hong Ha village about
50 km west of Hue, Thua Thien Prov., Vietnam,
16°17.35’N 107°21.59’E, 16 Mar 1999, J. Freyhof & D.
Serov; ZFMK 26670-26672 (3), 51.0-54.5 mm, stream
Mau about 30 km west of Hue, Thua Thien Hue Prov.,
Vietnam, 16°19.97’N 107°25.29’E, 16 Mar 1999, J. Frey-
hof & D. Serov; ZFMK 38460-38678 (19), stream run-
ning to River Huong, 40 km west of highway No. 1, Thua

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 113

Thien Hue Prov., Vietnam, 16°06.81’N 107°38.81’E, 19
Mar 2000, J. Freyhof, F. Herder & D. Serov.

Type material in other collections: CMK 16014 (4),
47.7-51.9 mm, same data as ZFMK 26811-26826.

Schistura kontumensis Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 160

Holotype: ZFMK 26687, 55.2 mm, stream Iasia about 25
km west of Kontum, Kontum Prov., Vietnam, 14°23.63’N
107°48.64’E, 23 Feb 1999, J. Freyhof & D. Serov.

Paratypes: ZFMK 26688-26747 (60), 23.0-55.0 mm,
same data as holotype.

Type material in other collections: ZSM 28869 (12),
39.7-51.4 mm, same data as holotype.

Schistura namboensis Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 163

Holotype: ZFMK 24094, 55.3 mm, small stream at Ea
Nuol about 20 km northwest of Buon Ma Thout, Dac Lac
Prov., Vietnam, 12°41.58’N 107°57.74’E, 26 Jan 1999, J.
Freyhof & D. Seroy.

Paratypes: ZFMK 24095-24177 (83), 30.5—60.8 mm,
same data as holotype.

Schistura psittacula Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 170

Holotype: ZFMK 27015, 63.1 mm, middle River Cam Lo
about 20 km west of Dong Ha, Quang Tri Prov., Vietnam,
16°47.00’°N 106°53.86’E, 14 Mar 1999, J. Freyhof & D.
Serov.

Paratypes: ZFMK 27016-27110 (95), 18.9-62.1 mm, same
data as holotype; ZFMK 38371-38400 (30), 22.3-63.8
mm, stream running to River Cam Lo, 25 km west of
Dong Ha, Quang Tri Prov, Vietnam, 16°45.71°N
106°51.63’E, 18 Mar 2000, J. Freyhof, F. Herder & D. Se-
Tov.

Type material in other collections: ZSM 28867 (10),

37.3-56.4 mm, same data as holotype; CMK 16013 (10),
32.1-55.9 mm, same data as holotype.

Bonn zoological Bulletin 59: 109-136

Schistura sokolovi Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 172

Holotype: ZFMK 27116, 67.7 mm, River Azun about 30
km east of Pleiku, a confluent to River Ba, Gia Lai Prov.,
Vietnam, 14°02.34’N 108°21.07’E, 24 Feb 1999, J. Frey-
hof & D. Serov.

Paratypes: ZFMK 27117-27122 (6), 55.8-67.0 mm, same
data as holotype; ZFMK 27123-27149 (27), 32.0—-95.3
mm, stream at Kon Ha Nung about 50 km north of An
Khe, a confluent to River Ba, Gia Lai Prov, Vietnam,
14°20.45’N 108°34.81°E, 25 Feb 1999, J. Freyhof & D.
Serov.

Type material in other collections: ZSM 28868 (6),
40.4-88.1 mm, same data as ZFMK 27123-27149; CMK
16012 (5), 45.8-87.6 mm, same data as ZFMK
27123-27149.

Schistura susannae Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 179

Holotype: ZFMK 28382, 44.5 mm, stream Mong Mo
south of Hai Van Pass at national highway No. 1, Quang
Nam Danang Prov., Vietnam, 16°09.67’N 108°07.73’E,
14 Mar 2000, J. Freyhof, F. Herder & D. Serov.

Paratypes: ZFMK 28383-28412 (30), 16.7—-51.8 mm,
same data as holotype.

Type material in other collections: CMK 16333 (6),
32.8-42.6 mm, same data as holotype.

Schistura thanho Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 182

Holotype: ZFMK 21047, 76.9 mm, River Vinh Thanh,
about 65 km northwest Quy Nhon, Binh Dinh Prov., Viet-
nam, 14°16.19’N 108°45.69°E, 23 Mar 1999, J. Freyhof
& D. Serov.

Paratypes: ZFMK 21048-21069 (22), 36.7-82.5 mm,
same data as holotype; ZFMK 21074 (1), 42.8 mm, stream
Xem, a confluent to River Vinh Tanh, about 50 km north-
west of Quy Nhon, Binh Dinh Prov, Vietnam, 14°05,00’N
108°47,01°E, 23 Mar 1999, J. Freyhof & D. Serov.

Type material in other collections: CMK 16019 (3),
43.1—70.8 mm, same data as holotype.

©ZFMK

114 Fabian Herder, Jan Huylebrouck & Klaus Busse

Schistura yersini Freyhof & Serov, 2001
Ichthyol Explor Freshwaters 12: 186

Holotype: ZMFK 23579, 63.2 mm, small stream running
to River Da Dung south of Da Lat, Lam Dong Prov., Viet-
nam, 11°46.86’N 108°25.04’°E, 19 Jan 1999, J. Freyhof
& D. V. Serov.

Paratypes: ZFMK 23580-23640 (45), 29.6-65.0 mm,
same data as holotype; ZFMK 23576-23577 (2), 30.9-31.3
mm, stream Suoi Vang north of Da Lat, Lam Dong Prov.,
Vietnam, 11°59.53’N 108°21.95’E, 19 Jan 1999, J. Frey-
hof & D. V. Serov; ZFMK 23661-23684 (24), 30.7—58.5
mm, stream running to River Dai Tan south of Da Lat,
Lam Dong Prov., Vietnam, 11°46.26’N 108°19.16’E, 20
Jan 1999, J. Freyhof & D. V. Serov; ZFMK 23687-23752
(66), 24.5-50.3 mm, River Dai Tan at Lam Ha village
south of Da Lat, Lam Dong Prov., Vietnam, 11°47.49°N
108°14.12’E, 20 Jan 1999, J. Freyhof & D. V. Serov;
ZFMK 23753-23758 (6), 32.2-53.4 mm, River Da Dung
25 km south of Da Lat, Lam Dong Prov., Vietnam,
11°45.48’ 108°23.78°E, 21 Jan 1999, J. Freyhof & D. V.
Serov; ZFMK 23762-23905 (144), 29.2—-60.3 mm, River
Dai Tan about 30 km south of Da Lat, Lam Dong Prov.,
Vietnam, 11°46.64’ 108°19.28’E, 21 Jan 1999, J. Freyhof
& D. V. Serov.

Type material in other collections: CMK 15998 (16),
29.2—59.2 mm, same data as holotype.

Sewellia albisuera Freyhof, 2003
Ichtyol Expor Frehswaters 14: 226

Holotype: ZFMK 32913, 44.3 mm, River Thu Bon about
16 km west of Than My, Quang Nam Da Nang Prov., Viet-
nam, 15°40.33’N 107°48.45’W, 12 Mar 2000, J. Freyhof
& D. Serov.

Paratypes: ZFMK 32910-32912 (3), 38.7-52.4 mm, same
data as holotype.

Type material in other collections: AMNH 231609 (10),
33.8—64.0 mm, River Nuac Ta at waterfall, Try My dis-
trict, Quang Nam Da Nang Prov., Vietnam, 29 Mar 1999,
M. L. J. Stiassny, R. A. Arrindell et al.; AMNH 231611
(3), 34.1-40.2 mm, River Tranh, Try My district, Quang
Nam Da Nang Prov., Vietnam, 4 Apr 1999, M. L. J. Sti-
assny, R. A. Arrindell et al.; AMNH 231616 (2), 56.0—57.4
mm, River Nuac Ta, Try My district, Quang Nam Da Nang
Prov., Vietnam, 29 Mar 1999, M. L. J. Stiassny, R. A. Ar-
rindell et al.; AMNH 231622 (1), 39.7 mm, River Nam
Nin, Try My district, Quang Nam Da Nang Prov., Viet-
nam, 3 Apr 1999, M. L. J. Stiassny, R. A. Arrindell et al.

Bonn zoological Bulletin 59: 109-136

Sewellia breviventralis Freyhof & Serov, 2000
Ichtyol Expor Frehswaters 11: 231

Holotype: ZFMK 20955, 50.7 mm, River Pako about 50
km north of Kontum, Kontum Prov., Vietnam,
14°39°60"N 107°46°98”E, 22 Feb 1999, J. Freyhof & D.
V. Serov.

Paratypes: ZFMK 20956-20961 (6), 37.8—50.1 mm, same
data as holotype.

Sewellia marmorata Serov, 1996
Vop Ikhtiol 36: 273

Paratypes: ZFMK 18543-18544 (2), Kon Ha Nung, Vi-
etnam, 13°57’N 107°32’E, 25 Jan 1991, D. Serov.

Type material in other collections: Holotype: ZMM P-
19760, same data as ZFMK 18543-18544; Paratypes:
ZMM P-19761 (11), same data as ZFMK 18543-18544.

Sewellia patella Freyhof & Serov, 2000
Ichtyol Expor Frehswaters 11: 235

Holotype: ZFMK 20962, 36.8 mm, stream Azun about 30
km east of Pleiku, Gia Lai Prov., Vietnam, 14°02’34”N
108°21°07”E, 24 Feb 1999, J. Freyhof & D. V. Serov.

Paratypes: ZFMK 20963-20980 (18), 21.6-48.2 mm,
same data as holotype; ZFMK 20981-20986 (6),
30.9-43.6 mm, River Dakpne 30 km northeast of Kontum,
Kontum Prov., Vietnam, 14°30°35”N 108°13’92”E, 21
Feb 1999, J. Freyhof & D. V. Serov; ZFMK 20988-20989
(2), 34.7-40.2 mm, stream near Buon Loi village, Gia Lai
Prov., Vietnam, 14°14’93”N 10°35’20”E, 25 Feb 1999, J.
Freyhof & D. V. Serov.

HEMIODONTIDAE

Pterohemiodus luelingi Géry, 1962
Bonn zool Beitr 12: 332

Holotype: ZFMK 1226, (ex MKI 59-60/QC/12), Quisto
Cocha, 15 km west of Iquitos, Peru, 1 May 1959, K. H.
Liiling

Paratype: ZFMK 1227, (ex MKI 59-60/QC/13), same da-
taas ZFMK 1226.

Type material in other collections: Paratype: MHNG
2150.6 (ex Géry M.202.3), same data as holotype.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 115

Current status: Synonym of Hemiodus atranalis Fowler,
1940 (Langeani in Reis et al. 2003: 98).

LEBIASINIDAE

Pyrrhulina zigzag Zarske & Géry, 1997
Das Aquarium 31 (No. 336): 14

Paratypes: ZFMK 19909-19912 (4), 21.2-25.6 mm, Ya-
rina Cocha, Rio Ucayali, Peru, 8 Aug 1974, K. H. Liiling.

Type material in other collections: Holotype: MTD F
17705, creek on road from Campo Verde to Nueva Re-
quena, Pert, 28 Aug 1996, P. Debold, K. Schade, I. Sei-
del, H. Windhorst, S. Wlach & A. Zarske; Paratypes: MTD
F 10963-10966 (4), Cocha Nazareth, tributary stream of
Rio Ucayali close by Requena, Peru, 18 Sep 1979, H.-J.
Franke; MTD F 10967-10969 (3), brook at the Carretera
Central, 21 km from Pucallpa, Pera, 21 Aug 1981, H.-J.
Franke; MTD F 17247-17254 (8), same data as holotype;
MTD F 17318-17333 (16), at Quebrada Manatai, Peru,
28 Aug 1996, P. Debold, K. Schade, I. Seidel, H. Wind-
horst, S. Wlach & A. Zarske; MTD F 17414-17425 (13),
Tierra Roja, 30 Aug 1996, P. Debold, K. Schade, I. Sei-
del, H. Windhorst, S. Wlach & A. Zarske; MTD F
17660-17663 (4), km 34 of the street Pucallpa-Campo
Verde, Peru, 28 Aug 1996, P. Debold, K. Schade, I. Sei-
del, H. Windhorst, S. Wlach & A. Zarske; Additional
paratypes at ZMB, ZMA, MHNG, and private collection
of Géry [no collection numbers provided].

CHARACIDAE

Aphyocharacidium bolivianum Géry, 1973
Beitr neotrop Fauna 8: 101.

Holotype ZFMK 2933, brook between upper Rio Chaparé
and Rio Chimoré, Rio Mamoré drainage, eastern of To-
dos Santos, Central Bolivia, 9 Oct 1966, K. H. Luling.

Type material in other collections: Paratypes: ZSM [no
number provided] (1), same data as holotype; ZSM [no
number provided] (1), Arroyo Chichigambo, near Espiritu
upper Yacuma, Rio Mamoré drainage, north of Rio Cha-
paré, 15 Aug 1955, K. P. Baumert [both paratypes current-
ly not in ZSM; at least one inventoried at MHNG as MH-
NG 2232.073; to be returned].

Astyanax villwocki Zarske & Géry, 1999
Mitt Hamb Zool Mus Inst 96: 199

Paratype: ZFMK 20781, 43.4 mm, Chaparé-Chimoreé, Bo-
livia, 1968, K. H. Liiling.
Bonn zoological Bulletin 59: 109-136

Type material in other collections: Holotype: MTD F
22400, 113.8 mm, Rio Pacal, Rio Pachitea basin, Depar-
tamento Ucayali, Pert, Sep 1973, W. Villwock; Paratypes:
ZMH 9175 (2), 55.0-55.8 mm, same data as holotype;
MTD F 22116 (1), 100.3 mm, same data as holotype, pri-
vate collection of J. Géry 1022.1 (1), same data as holo-
type; MTD F 17737-17741 (5), 32.9-59.4 mm, San Ale-
jandro on the Carretera Federico Basadre (km 111), Dept.
Ucayali, Peru, 31 Aug—2 Sep 1996, A. Zarske et al.; MTD
F 17614-17615 (2), 42.6-47.0 mm, pond on km 25 of the
road Campoverde-Tournavista, Dept. Ucayali, Peru, 30
Aug 1996, A. Zarske et al.; MNHN 1999-0318 to 0320
(3), 40.0-46.3 mm, Chapare-Coni, Bolivia, 1982, G.
Loubens & L. Lauzanne; MNHN 1999-0321 to 0323 (3),
41.1-46.0 mm, Rio San Juanito, Dept. Beni, Bolivia, 13
Jul 1982, G. Loubens; private collection of J. Géry
1022.2—3 (2), 38.6-51.5 mm, Rio San Juanito, Dept. Beni,
Bolivia, 13 Jul 1982, G. Loubens; MHNG 2603.17 (1),
94.1 mm, Bosque Nacional von Humboldt, middle Rio
Catilona, 86 km from Pucallpa, Dept. Ucayali, Pert, 31
May 1977, P. de Rham; MHNG 2603.18 (6), 29.3-43.8
mm, Bosque Nacional von Humboldt, Dept. Ucayali, Peru,
24 Jun 1978 [no collector provided].

Bryconamericus guyanensis Zarske, Le Bail & Géry,
2010
Vertebrate Zoology 60: 4

Paratypes: ZFMK 41764-41773 (10), 27.9-39.2 mm,
crique Eau Claire at 7 km of Saiil, Mana, French Guyana,
3 Jun 1983, Duraton & Demarty.

Type material in other collections: Holotype: MNHN
2010-0036, 43.2 mm, same data as ZFMK 41764-41773;
Paratypes: MTD F 31932-31936 (5), 18.7—30.1 mm, Bas
Maroni, crique Balatée, French Guyana, 9 Oct 1979, P.
Planquette (lot 1); MNHN 2010-0037 (2), 23.4-38.1 mm,
Village Antecume Pata, Haut Maroni, French Guyana, 15
Dec 1985, P. Planquette & P.-Y. Le Bail; 15.12.1985 (lot
2); MNHN 2010-0038 (12), 33.8-46.8 mm, crique Eau
Claire at 7 km of Satil, Mana, French Guyana, 3 Jun 1983,
Duranton & Demarty (lot 3); MTD F 31937-31949 (13),
31.8-43.1 mm, same data as ZFMK 41764-41773; MH-
NG 2722.008 (10), 31.6-41.0 mm, same data as ZFMK
41764-41773; ZMB 33936 (10), 31.0-41.4 mm, same da-
ta as ZFMK 41764-41773; MTD F 31950-31951 (2),
27.3-33.5 mm, Saut Ananas, crique No. 3, French
Guyana, 04°06’04”N 53°31°23”'W, 22 Sept 1995, P.-Y.
Le Bail, P. Planquette & P. Keith (lot 4); MTD F
31988-31993 (6), 21.4—-7.9 mm, Saut Ananas, French
Guyana 1957, Géry; MNHN 2010-0039 (14), 24.2-42.3
mm, Saut Capiaie, French Guyana, 04°05’°39”N
53°31731”W, 24 Sept 1995, P.-Y. Le Bail, P. Planquette
& P. Keith (lot 5); IRSNB 21735 (9), 36.2-56.7 mm,

©ZFMK

116 Fabian Herder, Jan Huylebrouck & Klaus Busse

Crique Colon, affluent de la Crique Lézard, near Citron,
French Guyana, 04°44’ 19”N 53°57°33”W, 13 Nov 1982,
P. Planquette & P.-Y. Le Bail.

Characidium heinianum Zarske & Géry, 2001
Zool Abh Mus Tierk Dresden 51: 231 Figs 1-3

Paratypes: ZFMK 39131-39135 (5), Rio Iourupuru, trib-
utary stream of Rio Mamoré, near the cross of the river
with the road Ramon, about 73 km north of Trinidad, Dep.
Beni, Bolivia, 14°12°53.8”S 64°56’26.8” W, 14-15 Jun
2000, A Zarske, G. Hein, T. Tonoki & J. Zapata.

Type material in other collections: Holotype: MTD F
24825, same data as ZFMK 39131-39135; Paratypes:
MTD F 24826-24830 (5); MTD F 24831-24832 (2), ali-
zarin preparation, same data as ZFMK 39131-39135;
MTD F 24826-24830 (5), same data as ZFMK
39131-39135; CIRA-UTB 968 a-e (5), same data as
ZFMK 39131-39135; ZMB 33236 (5); ZSM
29465-29469 (5), same data as ZFMK 39131-39135;
MHNG 2483 1-32 (2), same data as ZFMK 39131-39135;
MHNG 2618.90 (5), same data as ZFMK 39131-39135;
Private collection Géry 1043.1—5.2001 (5), same data as
ZFMK 39131-39135; CAZ 361 (5), same data as ZFMK
39131-39135.

Cheirodon luelingi Géry, 1964
Beitr neotrop Fauna 4: 19

Holotype: ZFMK 1417, small creek between Cano Yari-
na and Zapote Cocha, lower Rio Pacaya, Pert, 29 Sep
1959, K. H. Liling.

Paratype: ZFMK 1418, same data as holotype.

Type material in other collections: Paratypes: MHNG
2150.16—17 (2), same data as holotype.

Hasemania crenuchoides Zarske & Géry, 1999
Spixiana 22: 91]

Paratypes: ZFMK 20543-20547 (5), 28.6—31.88 mm, Cor-
rego Planaltina upper Rio Sao Bartolomeu, northeast of
Brasilia, Districto Federal, Goias, Brazil, Aug 1965, H.
Schultz.

Type material in other collections: Holotype: MZUSP
52732, male, 65.8 mm, same data as ZFMK 20543-20547;
ZSM 22656 (55), same data as ZFMK 20543-20547;
MTD F 21538-21545 (8), therefrom 3 alizarin prepara-
tions, same data as ZFMK 20543-20547; private collec-

Bonn zoological Bulletin 59: 109-136

tion of J. Géry 1010.1—-S5 (5), same data as ZFMK
20543-20547; MHNG 2594.44 (5), same data as ZFMK
20543-20547; ZMB 23959 (5), same data as ZFMK
20543-20547.

Hemigrammus luelingi Géry, 1964
Beitr neotrop Fauna 4: 7

Holotype: ZFMK 1304, small creek between Cano Yari-
na and Zapote Cocha, lower Rio Pacaya, Peru, 29 Sep
1959, K. H. Liiling.

Paratype: ZFMK 1305, same data as holotype.

Type material in other collections: Paratypes: MHNG
2150.14-15 (2).

Hyphessobrycon gracilior Géry, 1964
Beitr neotrop Fauna 4: 12

Holotype: ZFMK 1329, small creek between Cano Yari-
na and Zapote Cocha, lower Rio Pacaya, Peru, 29 Sep
1959, K. H. Luling.

Paratype: ZFMK 1330, same data as holotype.

Type material in other collections: Paratypes: MHNG
2150.12—13 (2).

Hyphessobrycon guarani Mahnert & Géry, 1987
Bonn zool Beitr 38: 307

Paratypes: ZFMK 15521-15523 (3), Monte Carlo, Misio-
nes Alto Parana, Argentina, Sep 1979, K. H. Liiling.

Type material in other collections: Holotype: MHNG
2366.99, 30.0 mm, Rio Alto Parana at Puerto Bertoni, Pa-
raguay, 13 Nov 1982, P. Berner, C. Dlouhy & V. Mahnert;
Paratypes: MHNG 2366.100 (77), 17.9-30.5 mm, there-
from 6 alizarin preparations, same data as holotype;
MHNG 2367.1 (34), Pirapyta, Lago Itaipu, Alto Parana,
19 Feb 1984, C. Dlouhy; MHNG 2367.2 (41), Itabo, La-
go Itaipu, 19 Oct 1984, C. Dlouhy; MHNG 2367.3 (1),
same data as ZFMK 15521-15553.

Hyphessobrycon nigricinctus Zarske & Géry, 2004
Zool Abh Mus Tierk Dresden 54: 33, Figs 1-3

Paratype: ZFMK 39847 (1), 27.4 mm, Import bred spec-
imen out off Rio Reducto, tributary stream of Rio Ma-
nuripi, near the city Alegria, Madre de Dios, Peru, ded Apr
2004, P. & M. Hoffmann.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 117

Type material in other collections: Holotype: MTD F
28465 (1), 32.8 mm, import specimen, same data as
ZFMK 39847; Paratypes: MTD F 28457-28459 (3),
29.5—32.7 mm, import specimens, same data as ZFMK
39847, 19 Jul 2004, R. Numrich; Private collection of J.
Géry, 1065.1.2004 (1), same data as ZFMK 39847.

Hyphessobrycon pando Hein, 2009
Bull Fish Biol 10: 2

Holotype: ZFMK 41452, 35.4 mm, Rio Manuripi, near to
Puerto Cardenas, Depertamento Pando, Bolivia,
11°20.083’S 67°44.576’W, 12 May 2001, G. Hein, H.
Langert, J. Zapata & A. Zarske.

Paratypes: ZFMK 41453-41456 (4), 27.4-36.7 mm, sa-
me data as holotype.

Type material in other collections: CIRA-UTB 973 a—c
(3), 29.3-32.0 mm, same data as holotype; MTD F
30756-30758 (3), 28.3-30.0 mm, same data as holotype.

Remarks: Additional material: ZFMK 41457-41459 (3),
same data as holotype, cleared and stained. Date in refe-
rence “31.12.2008”, but printed in 2009.

Hyphessobrycon procerus Mahnert & Géry, 1987
Bonn zool Beitr 38: 311

Paratypes: ZFMK 15518-15519 (2), Rio Guayrau-gua,
Depto. Caaguazu, Paraguay.

Type material in other collections: Holotype: MHNG
2385.68, male, 32.9 mm; Paratypes: MHNG 2385.69 (28),
therefrom 2 alizarin preparations; MNHN 1988-0468 (2);
MZUSP 45714 [ex MHNG 2385.69] (2); USNM 290841
[ex MHNG 2385.69] (3).

Hyphessobrycon tenuis Géry, 1964
Beitr neotrop Fauna 4: 10

Holotype: ZFMK 1327, small creek between Cano Yari-
na and Zapote Cocha, lower Rio Pacaya, Peru, 29 Sep
1959, K. H. Liiling.

Paratype: ZFMK 1328, same data as holotype.
Type material in other collections: Paratypes: MHNG
2150.8-11 (4), same data as holotype; ZMA 113.853 (1),

same data as holotype; ANSP 139708 (1), same data as
holotype.

Bonn zoological Bulletin 59: 109-136

Knodus borki Zarske, 2008
Vertebrate Zoology 58: 160

Paratype: ZFMK 41480 (1), 29.46 mm, near Iquitos, Pert,
specimen from commercial import, 2008, D. Bork don.

Type material in other collections: Holotype: MTD F
31357, 33.99 mm, same data as ZFMK 41480; Paratypes:
MTD F 31358-31362 (5), 29.74-33.13 mm, same data as
ZEMK 41480; NMW 95153 (1), 30.76 mm, same data as
ZFMK 41480; MHNG 27098.085 (1), 29.37 mm, same
data as ZFMK 41480; ZMB 33872 (1), 31.55 mm, same
data as ZFMK 41480.

Moenkhausia margitae Zarske & Géry, 2001
Zool Abh Mus Tierk Dresden 51: 7, Figs 1—2

Paratype: ZFMK 26448 (1), Campo Verde, Ucayali, Pe-
ru, 3 Sep 1978, K. H. Luling.

Type material in other collections: Holotype: MTD F
17256, Quebrada on road Campo Verde-Nueva Requena
(about half way), Ucayali, Peru; Paratypes: MTD F
17257-17258 (2), same data as ZFMK 26448; MHNG
2610.01, Bosque Nacional A. von Humboldt, Carretera
Central 86 km from Pucallpa in direction to Lima, Uca-
yali, Peri (No. 13), 29 May 1977, P. de Rham; MHNG
2610.02 (16) (No. 14), same data as MHNG 2610.01;
MHNG 2610.03 (13) (No. 14), same data as MHNG
2610.01; MHNG 2610.04 (13) (No. 15), same data as
MHNG 2610.01; private collection of J. Géry
1029.1—3.2000 (3), 37.2-52.0 mm, waters at km marker
85 of the road from Puerto Inca to Tingo Maria, Ucaya-
li, Peru, 9 Aug 1985, Staeck & Linke; private collection
of J. Géry 1029.4.2000 (1), 44.0 mm, Rio Neshuya, 60 km
southeast of Pucallpa, Ucayali, Peru, Feb 1984, Lombart.

Oxybrycon parvulus Géry, 1964
Beitr neotrop Fauna 4: 16

Holotype: ZFMK 1331, small creek between Cafio Yari-
na and Zapote Cocha, lower Rio Pacaya, Pert, 29 Sep
1959, K. H. Liiling.

Paratypes: ZFMK 1332-1357 (26), same data as holotype.
Type material in other collections: Paratypes: MHNG
2150.18-27 (10); ZMA 113.861 (2); ANSP 139709 (1);

all same data as holotype.

Remarks: Type species of Oxybrycon Géry, 1964.

©ZFMK

118 Fabian Herder, Jan Huylebrouck & Klaus Busse

Petitella georgiae Géry & Boutiére, 1964
Vie et Milieu Supl 17: 474

Paratype: ZFMK 1419, muddy bank of the Amazon Riv-
er, upstream of Iquitos, Distr. Loreto, Peru, Dec 1959, K.
H. Liling.

Type material in other collections: Holotype: MHNG
2150.28. (ex J. Géry collection M.314,1), male, near La-
gunas, lower Rio Huallaga, Distr. Loreto, Peru, beginning
of 1959, V. Manuel; Paratypes: MHNG 2150.29—64 (37),
same data as holotype; MHNG 2150 6588 (31), near
Iquitos, Distr. Loreto, Pert, beginning 1960, collector un-
known; ZSM 26332 (3), same data as holotype; MNHNP
1980-1447 (2), Distr. Loreto, Peru, 24 Feb 1961, P. Pi-
ron.

Piabarchus torrenticola Mahnert & Géry, 1988
Rev fr Aquariol 15: 3

Paratype: ZFMK 15520 (ex MHNG 2385.72) (1), Salto
Pirareta, Cordillera, Paraguay, 12 Nov 1985, C. Dlouhy,
Cl. Vaucher & V. Mahnert.

Type material in other collections: Holotype MHNG
2385.70, 45.7 mm, Salto Pirareta, 21 Mar 1985, C.
Dlouhy, F. Baud, & V. Mahnert; Paratypes: MHNG 2385
72 (59), same data as ZFMK 15520; MHNG 2385.73 (4),
smaller tributary stream running in Rio Piribebuy, 10 km
south of Tobati, Prov. Cordillera, 19 May 1985, C. Dlouhy,
F. Baud & V. Manhnert; MHNG 2385.74, Mountain
stream (afflux) running into Lago Ypacarai, Prov.
Cordillera, ded. H. Bleher.

Serrasalmus sanchezi Géry, 1964
Beitr neotrop Fauna 4: 27

Holotype: ZFMK 1216, Cafio Yarina, Rio Pacaya, creek
in lower Rio Ucayali, Pert, 31 Aug 1959, K. H. Liiling.

Paratype: ZFMK 1217, same data as ZFMK 1216.

Type material in other collections: Paratype: MHNG
2150.7, same data as ZFMK 1216.

Tucanoichthys tucano Géry & R6émer, 1997
Aqua, J Ichthy Aquat Biol. 2: 66 Figs 1, 5—6

Paratypes: ZFMK 18585-19589 (5), brook draining to
Igarape Yavuari river, tributary stream of river Uaupés, up-
per river Negro, Amazonas, Brazil, 0°14°317N
68°03°48" W, May 1996, U. Romer et al.

Bonn zoological Bulletin 59: 109-136

Type material in other collections: Holotype: MZUSP
51322 (10); MHNG 2588.41 (3); MTD F 18469-7; SMF
28208 (1); all same data as ZFMK 18585-19589.

Tyttobrycon dorsimaculatus Géry, 1973
Beitr neotrop Fauna 8: 118.

Holotype: ZFMK 2916, male, 19 mm, “Hoffmann-lago-
on”, Rio Chaparé (tributary of Rio Mamoré), lower To-
dos Santos, Central Bolivia, about 16°45’S 65°07’W, 7
Oct 1966, K. H. Luling.

Paratypes: ZFMK 2624 (1), 12 mm, same data as holo-
type; ZFMK 2952-2956 (5), small specimens, Rio Cha-
paré, ravine 4 km below Todos Santos, 8 Oct 1966, K. H.
Liling; ZFMK 2960-2962 (3), ravine 6 km below San
Francisco on the Rio Chipiriri, Mamoré-system, Central
Bolivia, 3 Oct 1966, K. H. Luling; ZFMK 2963-2979
(18), stagnant arm of the Rio Chipiriri 1.5 km below San
Francisco, Mamoré-system, Central Bolivia, 4 Oct 1966,
K. H. Luling.

Type material in other collections: Paratypes: private col-
lection of J. Géry [no collection numbers provided] (28).

Tyttobrycon spinosus Géry, 1973
Beitr neotrop Fauna 8: 121 Figs 23—26

Holotype: ZFMK 3000, brook between Rio Chaparé and
Rio Chimoré, Rio Mamoré system, east of Todos Santos,
Central Bolivia, 9 Oct 1966, K. H. Liling.

Paratype: ZFMK 1420 (1), same data as holotype.

Type material in other collections: Paratypes: Private col-
lection J. Géry [no collection numbers provided] (2), same
data as holotype.

DIPLOMYSTIDAE

Diplomystes camposensis Arratia, 1987
Bonn zool Mon 24: 45

Paratypes: ZFMK 14800 (1), 39°44°34”S 72°30°56”W,
Rio San Pedro close to Taco 3, Chile, 14 Dec 1985, K.
Busse; ZFMK _ 14801-14805 (5), 39°467°34”S
72°2710”W, Lago Rinihue (Desagiie), Prov. Valdivia,
Chile, Jan 1986, K. Busse.

Type material in other collections: Holotype: IZUA 3302,
169.6 mm, Lago Rinihue, 2 Oct 1983, A. Arriagada & A.
Klink; Paratypes: IZUA 3303 (2), Lago Rinihue, 4 Nov

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 119

1975; IZUA 2807 (1), Lago Rinihue (IZUA-material pro-
bably destroyed in the blaze of 2008); SMF 7579 (1), Rio
Valdivia, 11 Dec 1964, H. Canmpos; MCZ 54388 (1), Rio
San Pedro near Los Lagos, Prov. Valdivia, 22 Mar 1979,
G. Arratia; CAS 55428 (2), Lago Rinihue, 16 Mar 1979,
C. Eigenmann; KU 19210 (1) (flare example), Rio San Pe-
dro near Purey, 19 Feb 1967, G. Arratia & H. Diaz; KU
19209 (3), Rio San Pedro near Los Lagos, Feb 1976, G.
Arratia & H. Diaz; KU 19257 (1), Rio Curalelfa, Los La-
gos, Feb 1976, G. Arratia & H. Diaz; private collection
of G. Arratia 110276, 120276, 130276, 140276 (5) (flare
examples), Rio San Pedro, G. Arratia; private collection
of G. Arratia 111285 (1), Rio San Pedro near confluence
of Quinchilca,10 Dec 1985, K. Busse; private collection
of G. Arratia 011986 a (2), Desagiie Lago Rinihue, Jan
1986, K. Busse; private collection of G. Arratia 011986
b (3), Lago Rinihue, Jan 1986.

BAGRIDAE

Bagrus koenigi Pietschmann, 1932
Zool Anz 100: 92

Holotype: ZFMK (1899) 209, Nile, upper Egypt, 1899,
A. Koenig.

Current status: Synonym of Bagrus docmak Forskal, 1775
(Risch 1986: 11)

Pseudobagrus nubilosus Ng & Freyhof, 2007
Ichthyol Explor Freshwaters 18: 10, Fig. 1; 9-26

Paratypes: ZFMK 31245-31246 (2), 43.9-49.0 mm,
14°12°37.2”N 108°45’42.6’E, Nuoc Choe stream, a trib-
utary of Song Vinh Tranh, about 60 km northwest of Qui
Nhon, Binh Dinh Prov., Vietnam, 23 Mar 1999, J. Frey-
hof & D. V. Serov.

Type material in other collections: Holotype: UMMZ
246370, 48.5 mm, same data as ZFMK 31245-31246;
Paratypes: UMMZ 246320 (2), 49.0-50.8 mm, same da-
ta as ZFMK 31245-31246.

Pseudomystus sobrinus Ng & Freyhof, 2005
Copeia 2005:745

Holotype: ZFMK 27167, 63.2 mm, 12°53’05’N
109°23’70”E, stream about 15 km south of Tuy Hoa Phu
Yen Prov., Vietnam, 26 Feb 1999, J. Freyhof & D. V.
Serov.

Bonn zoological Bulletin 59: 109-136

Paratypes: ZFMK 24068-24073 (6), 38.3-59.0 mm,
Crong Hang (stream) near the village of M’Drac, about
60 km west of Ninh Hoa, tributary of Song Ba, Dac Lac
Prov., Vietnam, 12°46’10”N 108°42’76”E, 28 Jan 1999,
J. Freyhof & D. V. Serov; ZFMK 24166 (1), 86.3 mm,
market of Azun Pa, Gia Lai Prov., 30 Apr 2000, J. Frey-
hof, F. Herder & D. V. Serov; ZFMK 27168-27183 (17),
30.6-79.8 mm, same data as holotype; ZFMK
27184-27185 (2), 38.0-42.5 mm, stream about 30 km
west of Tuy Hoa, Phu Yen Prov., Vietnam, 13°03’08”N
109°05’00”E, 2 Mar 1999, J. Freyhof & D. V. Serov.

Type material in other collections: UMMZ 245016 (3),
46.3-47.8 mm; ZRC 49172 (2), 42.6-52.5 mm; all same
data as ZFMK 27184-27185.

AMPHILIIDAE

Leptoglanis wamiensis Seegers, 1989
Aquar Terrar Z 42: 285

Holotype: ZFMK 15899, Kisangata creek, tributary of the
Wami river at Mvumi, 32 km southwest of Kidete on the
Kimamba-Kilosa road, Morogoro, Tanzania, 3 Jan 1989,
B. Kilian & L. Seegers.

Paratypes: ZFMK 15900-15902 (3), same data as holo-
type.

Current status: Synomym of Zaireichthys rotundiceps
Hilgendorf, 1905 (Ferraris 2007: 29)

AKYSIDAE

Akysis clavulus Ng & Freyhof, 2003
Ichthyol Explor Freshwaters 14: 312

Holotype: ZFMK 27159, about 10 km west of Ninh Hoa,
Song Duc My, Khanh Hoa Prov., Vietnam, 12°32’22”"N
109°01°38”E, 10 Feb 1999, J. Freyhof & D. V. Serov.

Paratypes: ZFMK 25506-25516 (11), 23.8-31.0 mm,
same data as holotype; ZFMK 223265 (1), stream near
Kanh Vinh about 34 km from Nha Trang, tributary stream
of Song Cai, 12°16’42”’N 107°54’56”E, 17 Dec 1998, J.
Freyhof & D. V. Serov; ZFMK 27506-27509 (4),
25.5—31.4 mm, stream near grave of Yersin, about 20 km
south of Nha Trang in a tributary stream of Song Cai,
12°10°73”"N 103°04’10”E, 13 Feb 1999, J. Freyhof & D.
V. Serov; ZFMK 27156-27158 (3), 21.2—-28.0 mm, and
ZFMK 27160-27165 (6), 25.0-27.7 mm, stream Giang
May, about 30 km south of Nha Trang, a tributary of Song
Cai, Knanh Hoa Prov., Vietnam, 12°12712”N

©ZFMK

120 Fabian Herder, Jan Huylebrouck & Klaus Busse

108°55°48”E, May 2000, J. Freyhof, F. Herder & D. V.
Serov.

Type material in other collections: Paratypes: UMMZ
238018 (1), 35.0 mm; ZRC 46177 (3), 26.8-30.0 mm; all
same data as ZFMK 27159.

SISORIDAE

Glyptothorax filicatus Ng & Freyhof, 2008
Zootaxa 1873: 12

Holotype: ZFMK 34398, 83.3 mm, 12 km east of A Lu-
oi, stream draining to Se Sap River, Thua Thien Hue
province, Vietnam, 16°20.57’N 107°9.41°E, 20 Mar 2000,
J. Freyhof, F. Herder & D. Serov.

Paratype: ZFMK 34399 (1), 91.9 mm, same data as holo-
type.

Type material in other collections: Paratype: UMMZ
245668 (1), 69.0 mm, same data as holotype.

Glyptothorax strabonis Ng & Freyhof, 2008

Zootaxa 1873: 16

Paratypes: ZFMK 35285-35286 (2), 84.0—95.5 mm, mar-
ket in Phong Nha, Quang Binh province, Vietnam, 7 Apr
2000, J. Freyhof, F. Herder & D. Serov.

Type material in other collections: Holotype: UMMZ
245669, 80.1 mm, same data as ZFMK 35285-35286.
Oreoglanis infulatus Ng & Freyhof, 2001

J Fish Biol 59: 1165; Figs 1-3

Holotype: ZFMK 35719, 76.1 mm, stream at Son Kima,
tributary of Song Lam, Ha Tinh Prov., Vietnam,
18°24°25"N 105° 117107 ES MApr 1999. J: FreyhotiF:
Herder & D. V. Serov.

Paratypes: ZFMK 35720-35725 (6), 40.3—72.4 mm.
Type material in other collections: Paratypes: UMMZ
238025 (6), same data as ZFMK 35719.
HEPTAPTERIDAE

Imparfinis pseudonemacheir Mees & Cala, 1989
Proc konink nederl Akad Wetensch C 92: 388

Bonn zoological Bulletin 59: 109-136

Paratypes: ZFMK 17818-17819 (2), at the street of Pu-
callpa to Tournavista, Distr. Loreto, Peru, 3 Sep 1978, K.
H. Liling.

Type material in other collections: Holotype and
paratypes: RMNH 30489 (3) (largest specimen is the holo-
type); Rio Tucuragua near Caicara, Bolivar, Venezuela, 25
Mar 1986, B. Roman; RMNH 30487 (4), Rio Guayuriba
(Puerto Lopez), Meta, Colombia, 20 Feb 1972, P. Cala;
ICN-NHM 1403 (4), same data as RMNH 30487.

CALLICHTHYIDAE

Corydoras lacerdai Hieronimus, 1995
Z Fischk 3: 110

Paratypes: ZFMK 18551, 33.6 mm; ZFMK 18552, 27.8
mm; ZFMK 18553-18556 (4), 21.0-26.0 mm; all Rio
Ribeira da Terra Firme between Canavieiras and Ilheus,
Brazil, 1 Jan 1993, M. T. C. Lacerda, S. S. Valério & D.
Byron.

Type material in other collections: Holotype: MZUSP
47682, female, 33.6 mm; Paratypes: MZUSP 47683 (1),
33.0 mm and MZUSP 47684 (1), male, 25.2 mm; MZUSP
without number (12), 14-23 mm; ZMA 121.255 (4), pri-
vate collection of Hieronimus (4); all same data as ZFMK
Ihesyay le

Corydoras pinheiroi Dinkelmeyer, 1995
Aquarist aktuell 3: 60

Paratypes: ZFMK 18499, 18500, 18540-18542 (5), trib-
utary stream of Rio Ribeirao, near the village of Guayara-
Mirim, Est. Rondonia, Brazil, 3 Oct 1994, M. T. C La-
cerda, Y. Nakamura & V. Alvez.

Type material in other collections: Holotype: MZUSP
48099, male, 57.0 mm; Paratypes: ZMA 121.269 (3); all
same data as ZFMK 18499, 18500, 18540-18542.

Corydoras sarareensis Dinkelmeyer, 1995
Aquarist aktuell 3: 61

Paratypes: ZFMK 18504-18506 (3), Rio Sararé, right
hand side tributary stream of Rio Guaporeé, Est. Mato
Grosso, Brazil, 1994, M. T. C. Lacerda.

Type material in other collections: Holotype; MZUSP

48100, male, 43.8 mm; Paratypes: ZMA 121.268 (4); all
same data as ZFMK 18504-18506.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 121

Corydoras seussi Dinkelmeyer, 1996
Aqu aktuell 1996: 25

Paratypes: ZFMK 18628-18653 (26), small side stream
at Rio Pacas Novos, side branch of Rio Mamoré near Gua-
jar-Mirim, Est. Rod6nia, Brazil, Sep 1999 (date of shipp-
ment), T. C. Lacerda.

Type material in other collections: Holotype: MZUSP
49322, female, 60.8 mm; Paratypes: MZUSP 49323 (10);
ZMA 121.570 (10); all same data as ZFMK 18628-18653.

Corydoras similis Hieronimus, 1991
Z Fischkunde 1: 41

Holotype: ZFMK 16400, 35.3 mm, near the city of
Ariqumes, a tributary of Rio Madeira, federal state of
Rodonia, Brazil, commercial import.

Paratypes: ZMA 120.746, same data as ZFMK 16400.

LORICARIIDAE

Ancistrus pirareta Muller, 1989
Rev Suisse Zool 96: 891

Paratypes: ZFMK 16281-16282 (2), Salto Pirareta, Piri-
bebuy, Afflux of Rio Tebicuary-mi, Cordillera, Paraguay,
21—23 Mar 1985, F. Baud, C. Dlouhy & V. Mahnert.

Type material in other collections: Holotype: MHNG
2450.10, male, 82.3 mm, same data as ZFMK
16281—16282; Paratypes: MHNG 2268.56 (78) (2 of the-
se are alizarin preparations), MCP (2) [no number provi-
ded], USNM 307765 (2), ZMA 120.329 (5) and ZSM
PISS O);

Ancistrus piriformis Muller, 1989
Rev Suisse Zool 96: 887

Paratypes: ZFMK 16279-16280 (2), Rio Acaray beneath
the barrier lake, Alto Parana, Paraguay, 15 May 1982, C.
Dlouhy.

Type material in other collections: Holotype: MHNG
2450.11, male, 80.8 mm, same data as ZFMK
16279-16280; Paratypes: MHNG. 2268.58—59 (88), (2 of
these are alizarin preparations), same sampling locality as
ZFMK 16279-16280; MCP (2), USNM 307766 (2), ZMA
120.328 (5) and ZSM (2); all same data as ZFMK
16279-16280; MHNG 2160.25 (1), 52.7 mm and 2268.20
(1), 72.5 mm, 7 Oct 1980.

Bonn zoological Bulletin 59: 109-136

Pseudohemiodon (Planiloricaria) cryptodon \sbriicker,
197]
Bonn zool Beitr 21: 278

Holotype: ZFMK 1865 (ex ZFMK 1/66/1717), Rio Uca-
yali near Pucallpa, Per, Sep—Oct 1966, K. H. Liiling.

Remarks: Type species of the subgenus Planiloricaria |s-
briicker, 1971.

Current status: Valid as Planiloricaria cryptodon Isbrtick-
er, 1971 (Isbriicker 1980).

GALAXIIDAE

Brachygalaxias gothei Busse, 1982
Bonn zool Beitr 33: 72

Holotype: ZFMK 11110, male, Fundo Porvenir, Talca,
Chile, 9 Nov 1981, K. H. Gothe & K. Busse.

Paratypes: ZFMK 11111-11130, 11137-11158 (42), same
data as holotype.

Type material in other collections: Additional paratypes
in the collections of BM 1982.11.10: 865-866 (2) (ex
ZFMK 11131-11132); IZUA 2721 (2) (ex ZFMK
11133-11134); MNHNC P6.055—P6.056 (2) (ex ZFMK
11135-11136).

Galaxias bullocki Regan, 1908
Ann Mag nat Hist London 8: 372

Paralectotypes: ZFMK 12911 (1), male, Maquehue near
Temuco, Chile, 6 Apr 1907, D. S. Bullock; ZFMK 12912
(1), female, same data as ZFMK 12911. Both specimens
ex BMNH.

Type material in other collections: Lectotype: BMNH
1968.9.16.1, female, same data as ZFMK 12911; Paralec-
totypes: BMNH 1968.9.16.2-47, same data as ZFMK
12911 (design. Busse 1984, Bonn zool Mon 19: 217).

Current status: Valid as Brachygalaxias bullocki (Regan,
1908) (McDowall 1971: 60).

SALMONIDAE

Coregonus fontanae Schulz & Freyhof, 2003
Ichthyol Explor Freshwaters 14: 212

©ZFMK

122 Fabian Herder, Jan Huylebrouck & Klaus Busse

Holotype: ZFMK 22805, 118.8 mm, Lake Stechlin, Bran-
denburg, Germany, 53°09’N 13°04’E, 23 Apr 2001, M.
Schulz.

Paratypes: ZFMK 22806-22820 (15), 98.3-118.6 mm,
same data as holotype.

Type material in other collections: Paratypes: ZMB 33328
(20), 90.0-125.9 mm, same data as holotype.

NOTHOBRANCHITIDAE

Aphyosemion (Mesoaphyosemion) etsamense Sonnen-
berg & Blum, 2005
Bonn zool Beitr 53: 213

Holotype: ZFMK 39832, male, 32.4 mm, small river at
the village Etsam I crossing the road N5 from Medoneu
to Kougouleu, western slopes of the Monts de Cristal,
Gabon, 0°46°34,1”"N 10°24’03”E, 29 Jul 2002, T. Blum,
G. Fleck & R. Sonnenberg.

Paratypes: ZFMK 39833-39842 (10), same data as holo-
type; ZFMK 39843 (1), male, north of the village Assok,
Gabon, 0°43°36,9”"N 10°21°58,1”E, 11 Sept 2002, T.
Blum & P. Sewer; ZFMK 39844-39846, same locality as
holotype, Aug 1999, preserved after approximately 2 years
in aquarium, F. Bitter, T. Blum & P. Sewer.

Type material in other collections: MRAC A4-42-P-1-4,
same data as holotype; IRET (2), same data as holotype.

Callopanchax sidibei Sonnenberg & Busch, 2010
Bonn zoll Bull 57: 6

Holotype: ZFMK 41613, male, 58.2 mm, small river and
adjacent pools and ditches in a secondary forest, close to
the small village Bombokore, about 3 km from the clos-
est larger village Fandi, Kindia region, Guinée maritime,
Guinea, 9°32.71’N 13°14.51?W, 25 Oct 2008, E. Busch
& B. Wiese.

Paratypes: ZFMK 4161441615 (2), 47.0 (male)—S55.4 (fe-
male) mm, same data as holotype; ZFMK 41616-41628
(13), 39.0-58.9 mm, same data as holotype.

Type material in other collections: Paratypes: MRAC
2010-02-P-1-5 (5), 46.3-59.9 mm, same data as holotype.

Bonn zoological Bulletin 59: 109-136

Chromaphyosemion koungueense Sonnenberg, 2007
Zootaxa 1591: 7

Holotype: ZFMK 39949, male, 24.3 mm, river Koungué,
a small river in the forest near the village Koungué Ndon-
ga at the road from Edea to Dizangue and Ndonga,
Cameroon, 03°48736.9”"N 09°55’03.9”E, 4 Jan 2000, U.
Kampf, R. Sonnenberg & A. Trankner.

Paratypes: ZFMK 39950-39955 (6), same data as holo-
type; ZFMK 39956-39957 (2), males, small creek cross-
ing the road Mbambou to Mouanko, some kilometers in
oil palm plantation in direction Mouanko, Cameroon,
03°42°48.8°N 09°51°28.2”E, 2003, H. Kullmann & M.
Volker; ZFMK 39958-39959 (2), males, near the road
from Edea to Kribi, Cameroon, 03°32’28.9"N
10°05712.1”E, 6 Jan 2000, E. U. Kampf, R. Sonnenberg
& A. Trankner; ZFMK 39960-39964 (5), small creek
crossing the road from Edea to Kribi near the village
Koukoué, Cameroon, 03°43718.177N 10°06’26.9”E, 14
Dec 2001, H. Kullmann, B. Misof & R. Sonnenberg;
ZFMK 39965-39969 (5), small creek near the road Edea
to Knibi, Cameroon, 03°36’29.2”N 10°06’26.2”E, 14 Dec
2001, H. Kullmann, B. Misof & R. Sonnenberg.

Type material in other collections: CENAREST (2), same
collection data as ZFMK 39960-39964.

Chromaphyosemion melinoeides, Sonnenberg, 2007
Zootaxa 1591: 16

Holotype: ZFMK 40000, male, 25.1 mm, the first stream
crossing the road from Kribi to Ebolowa after the village
Akok in direction Ebolowa, Cameroon, 02°47730.4”N
10°17°00.0”E, 17 Jan 2000, U. Kampf, R. Sonnenberg &
A. Trankner.

Paratypes: ZFMK 39987-39988 (2), same data as holo-
type; ZFMK 39989-39997 (10), near the village Fenda at
the road from Kribi to Ebolowa, Cameroon, 02°49’02.7”N
10°22’50.6”E, 17 Jan 2000, U. Kampf, R. Sonnenberg &
A. Trankner; ZFMK 39998-39999 (2), males, small creek
Mintang near the village Fenda on the road from Kribi to
Ebolowa, Cameroon, 02°49’36.0”N 10°22’39.0”E, 8 Jan
1998, W. Eberl, U.Kampf & A. Kliesch; ZFMK
40001-40003 (3), small creek crossing the road from Kribi
to Ebolowa, near the village Njabilobé, Cameroon, 7 Jan
1998, W. Eberl, U.Kampf, A. Kliesch; ZFMK 40004 (1),
male, road from Kribi to Ebolowa, east of Akok,
Cameroon, 02°47756.2”N 10°18’55.3”E, 17 Jan 2000, U.
Kampf, R. Sonnenberg & A. Trankner.

Type material in other collections: CENAREST, Batoke,
Cameroon (1), male, same data as ZFMK 39998-39999;

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 123

CENAREST, Batoke, Cameroon (1), female, same data
as ZFMK 40004.

Chromaphyosemion omega Sonnenberg, 2007
Zootaxa 1591: 12

Holotype: ZFMK 39970, male, 31.3 mm, small river Ngo
Njock near the village Ndog Nyang, old road from Edea
to Douala, Cameroon, 03°59712.4”"N 10°05°50.3”E, 5 Jan
2000, U. Kampf, R. Sonnenberg & A. Trankner.

Paratypes: ZFMK 39971-39980 (10), same data as holo-
type; ZFMK 39981-39986 (7), creek crossing the old road
from Edea to Douala, near Bonepoupa, Cameroon,
04°06°30.17°N 10°01°12.6”E, 9 Oct 2001, H. Kullmann,
B. Misof & R. Sonnenberg.

Type material in other collections: CENAREST, Batoke,
Cameroon (1), male, same data as holotype; CENAREST,
Batoke, Cameroon (1), female, same data as ZFMK
39981-39986.

Episemion krystallinoron Sonnenberg et al. 2006
Zootaxa 1361: 6

Holotype: ZFMK 39942, male, 27.9 mm, river near vil-
lage of Nkineén, crossing the road N5 from Médoneu to
Kougouleu, northern part of the Monts de Cristal, Gabon,
OO Se0GSgNe10241°33.57E, 29 Jul 2002, TF Blum, G.
Fleck & R. Sonnenberg.

Paratypes: ZFMK 39943-39945 (3), same data as holo-
type; ZFMK 39948 (1), male, same data as holotype;
ZFMK 39946-39947 (2), river near Médouneu, at road
NS, northern part of the Monts de Cristal, Gabon,
1°00°06.0”N 10°45’50.9”E, 24 Aug 1999, T. Blum, F. Bit-
ter & P. Sewer.

Type material in other collections: MRAC 2006-09-P-1
(1), male, same data as holotype; IRET (uncat.) (1), male,
same data as holotype.

Fundulopanchax kamdemi Akum, Sonnenberg, Zee &
Wildekamp, 2007
Zootaxa 1532: 43

Paratypes: ZFMK 39900-39917 (18), small forest stream
emptying in swampy pools, Korup Nationalpark,
Cameroon, 05°00’20”N 08°47°01”E, Dec 2001, C.
Akum; ZFMK 39918-39931 (14), juv., same data as
ZFMK 39900-39917.

Bonn zoological Bulletin 59: 109-136

Type material in other collections: Holotype: MRAC
A2—003—P—0001, male, small pools under forest cover,
Korup Nationalpark, Cameroon, 05°05’N 08°52’E, 12 Jan
2001, G. Chiambeng; Paratypes: MRAC A2-003-P-
0002-0003 (2), males, same data as holotype; MRAC 92-
144-P-0384—0386 (3), 40.5—-47.3 mm, west of science
camp, Korup Nationalpark, Cameroon, 05°05’N 08°52’E;
19 Dec 1992, A. Kamdem Toham; IRAD Research Sta-
tion Batoke, Limbe, Cameroon (4), small forest stream
emptying in swampy pools, Korup Nationalpark,
Cameroon, Dec 2001, C. Akum.

Nimbapanchax leucopterygius Sonnenberg & Busch,
2009
Zootaxa 2294: 11

Holotype: ZFMK 41481, male, 43.6 mm, Foret Classeé
de Dieke, southeastern Guinea, 7°28.70’N 8°50.45’W, 16
May 2006, E. Busch & B. Wiese.

Paratypes: ZFMK 41482-41483 (2), females, same data
as holotype.

Type material in other collections: MRAC 73-39-P-
1897-1904 (8), N’zerekore, Guinea, Nov 1963—Feb
1964, H. Olsen; MRAC 75-26-P-01—04 (4) Salayea,
Liberia, 7°37°N 9°30’W, 1971, E. Roloff; MRAC 81-19-
P-114—116 (3), riviertje buiten Déoule, zijriviertje van de
Kilo op secundaire weg van Man naar Biankouma (rivulet
near Déoulé, small tributary of the Kilo River, on second-
ary road from Man to Biankouma), Cote dIvoire, 24 Jul
1980, H. De Breuyn.

Remarks: Type species of Nimbapanchax Sonnenberg &
Busch, 2009

Nimbapanchax melanopterygius Sonnenberg & Busch,
2009
Zootaxa 2294: 14

Holotype: ZFMK 41484, male, 42.9 mm, Cavally River,
southeastern Guinea, 7°41.83’N 8°26.48’W, 15 May 2006,
E. Busch & B. Wiese.

Paratypes: ZFMK 41485-41487 (3), females, same data
as holotype; ZFMK 41488-41493 (6), near Kementa in
the Konkere river, southeastern Guinea, 7°43.83’N
8°26.49’W, 15 May 2006, E. Busch & B. Wiese.

Type material in other collections: MRAC 2008-16-P-1-4

(4), same data as ZFMK 41488-41493; ZSM 37485 (4),
same data as ZFMK 41488-41493.

©ZFMK

124 Fabian Herder, Jan Huylebrouck & Klaus Busse

Nothobranchius kafuensis Wildekamp & Rosenstock,
1989
Aquar Terrar Z 42: 417

Holotype: ZFMK 13951, male, 50.2 mm, seasonal pool
in Kafue National Park, 5 km south of Chunga-camp, 100
m from Kafue River, Zambia, 26°00’E 15°03’S, 20 Feb
1983, K. Oestergaard.

Paratype: ZFMK 13952 (1), female, 36.0 mm, same da-
ta as holotype.

Type material in other collections: Paratypes: MRAC 89-
07-P-1 (1), male, 43.8 mm, same data as holotype; AM
PF 1542 (5), males, seasonal pool of Kafue river system,
Kafue National Park, Zambia, 25°45’E 16°10’S, G. Bell-
Cross.

Nothobranchius luekei Seegers, 1984
Aquar Terrar Z 37: 250

Holotype: ZFMK 13551, male, 40.4 mm, bridge across
Mbezi at road to Kibiti, about 40 km south of Dar es
Salaam, East Tanzania, Jan 1983, L. Seegers & H. J. Wis-
chmann.

Paratypes: ZFMK 13552, female, 27.8 mm, same data as
holotype; ZFMK 13553-13556 (4), 22.8-27.0 mm, F1
aquarium bred.

Type material in other collections: Paratypes: BMNH
1984.3.22.1—2 (2), 27.1-29.0 mm, same data as holotype.

Nothobranchius rubripinnis Seegers, 1986
Aquar Terrar Z 39: 19

Holotype: ZFMK 13941, male, 26.8 mm, bridge across
Mbezi at road to Kibiti, about 40 km south of Dar es
Salaam, East Tanzania, Jan 1983, L. Seegers & H. J. Wis-
chmann.

Paratypes: ZFMK 13942-13946 (5), 19.3—29.6 mm, same
data as holotype; ZFMK 13947-13949 (3), females,
21.7-24.1 mm, few hundred meters northeast of River
Mbemkuru at road from Lindi to Kilwa, East Tanzania,
30 Jun 1985 [label in conflict with original description,
showing same collection data as holotype].

Type material in other collections: Paratypes: ZMB
31546-31547 (4), 22.0—26.9 mm, same data as holotype;
NRM (4), 21.2—27.1 mm, same data as holotype; MRAC
KTZ 85/28 (4), 21.7—25.2 mm, few hundred meters north
of River Mbemkuru at road from Lindi to Kilwa, East Tan-
zania, 30 Jun 1985, [no collector provided].

Bonn zoological Bulletin 59: 109-136

Paranothobranchius ocellatus Seegers, 1985
J. Deutsche Killifisch Gem, Suppl 1: 47

Holotype: ZFMK 13927 (but see below), male, 50.0 mm,
swamp between Mtanza and northern entrance of Selous
Game Reserve, north of Rufiji, East Tanzania, 31 Dec
1980, G. Eggers, C. Kasselmann & L. Seegers.

Remarks: The author has received collection number
ZFMK 13927 prior to description and deposition of the
specimen in Bonn. ZFMK did however not receive the
specimen so far; deposition of the holotype has accord-
ingly not taken place. Type species of Paranotho-
branchius.

Current status: Valid as Nothobranchius ocellatus
(Seegers, 1985) (Lazara 2001: 225).

Roloffia banforensis Seegers, 1982
Aquar Terrar Z 35: 449

Holotype: ZFMK 12155, male, 26.8 mm, Comoé draina-
ge, 15 km north of Banfora, 85 km southwest of Bobo
Dioulasso, Obervolta, Oct 1980, B. Bauer.

Paratypes: ZFMK 12156-12162 (7), same data as holoty-
pe.

Type material in other collections: Paratypes: MNHN 60-
131 (7), road from Banfora to Sindou, Obervolta, 9 Nov
1959, J. Arnoult & d’Auberton; SMF 17679-17681 (3),
21.0-36.2 mm, aquarium bred.

Current status: Valid as Scriptaphyosemion banforense
(Seegers, 1982) (Lazara 2001: 289).

RIVULIDAE

Cynolebias pantanalensis Seegers, 1988
Aquar Terrar Z 41: 31

Holotype: ZFMK 15534, male, 21.6 mm, close to road
Transpantaneira between Poconé and Corumba (Mato
Grosso), about 45 km south of Poconé, drainage of Rio
Sararé and Rio Paraguai, Mato Grosso, Brazil, about
56°50’W 16°43’S, | Aug 1987, W. Staeck et al.

Paratypes: ZFMK 15535-15541 (7), 23.9-26.1 mm, same
data as holotype; ZFMK 15542, female, 24.1 mm, small
inlet of Rio Cuiaba, about 5 km north of Porto Cercado
at road Poconé to Porto Cercado, Brazil, about 56°23’W
16°33’S, 30 Mar 1986, W. Staeck.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 125

Type material in other collections: Paratypes: MZUSP [no
collection number provided] (4), 17.2-20.4 mm, same da-
ta as holotype.

Current status: Synonym of Plesiolebias glaucopterus
Costa & Lacerda, 1988 (Costa 1991: 375).

Pterolebias rubrocaudatus Seegers, 1984
Bonn zool Beitr 35: 243

Holotype: ZFMK 13413, male, 31.5 mm, Rio Tambopa-
ta area, 7 km from Puerto Maldonado, Rio Madre de Dios
drainage, Southeast Pert, 25—28 Apr 1981, K. Elgner &
E. Lenkenhoff.

Paratype: ZFMK 13414, female, 29.4 mm, same data as
holotype.

Current status: Valid as Aphyolebias rubrocaudatus
(Seegers, 1984) (Costa 1998: 78).

Pterolebias staecki Seegers, 1987
Aquar Terrar Z 40: 202

Holotype: ZFMK 14597, male, 52.7 mm, swampy lago-
on of Lago Janahuaca, south of Rio Amazonas, not far ups-
tream of confluent with Rio Negro, southwest of Manaus,
Brazil, about 60°16’ W 3°25’S, 20 Mar 1986, W. Staeck.

Paratypes: ZFMK 14598, female, 40.8 mm, same data as
holotype; ZFMK 14599-14601 (3), 30.5—41.2 mm, same
data as holotype.

Type material in other collections: MZUSP [no collection
number provided] (2), 39.1-33.5 mm, aquarium bred.

Current status: Valid as Moema staecki (Seegers, 1987)
(Costa 1993: 623).

Pterolebias wischmanni Seegers, 1983
J Deutsche Killifisch-Gemeinschaft 15: 68

Holotype: ZFMK 11534, male, Ucayali drainage 120 km
south of Pucallpa, Peru, Jul 1981 (preserved Feb 1982),
H. J. Wischmann & H. Fries.

Paratypes: ZFMK 11535-1536 (2), females, same data as
holotype.

Type material in other collections: Paratypes: BM
1983.6.9.2-82 (3), males, aquarium bred of holotype and
a paratype.

Bonn zoological Bulletin 59: 109-136

Current status: Valid as Aphyolebias wischmanni (Seegers,
1983) (Costa 1998: 78).

Rivulichthys luelingi Meinken, 1969
Bonn zool Beitr 20: 423

Holotype: ZFMK 2091, male, 36.0 mm, hollow closeby
Rio Chaperé, about 4 km from village Todos Santos, East
Bolivia, 8 Oct 1966, K. H. Liiling.

Paratypes ZFMK 2092-2095 (11), same data as holotype.

Current status: Synonym of Prerolebias longipinnis Gar-
man, 1895 (Thomerson 1984: 528).

Rivulus bolivianus Seegers, 1988
Bonn zool Beitr 39: 173

Holotype: ZFMK 14602, male, 24.1 mm, lagoon “Suarez”,
Rio Mamoré drainage, about 4 km southeast of Trinidad,
Bolivia, 24 Aug 1985, H. Linke & W. Staeck.

Paratypes: ZFMK 14603-14609 (7), same data as holo-
type.

Current status: Synonym of Rivulus beniensis Myers, 1927
(Costa in Reis et al. 2003: 534).

Rivulus caudomarginatus Seegers, 1984
Zool Beitr 28: 307

Holotype: ZFMK 12848, male, Greta Funda, south of Rio
de Janeiro, Brazil, Jul-Aug 1981, R. A. Ferreira.

Paratypes: ZFMK 12849 (1), female, same data as holo-
type; ZFMK 12850-12854 (5), males, aquarium bred;
ZFMK 12855-12859 (5), females, aquarium bred.

Type material in other collections: Paratypes: SMF
17864 (4), aquarium bred; BM 1983.6.9. 16—19 (4), aquar-
ium bred; ZMA 119096 (4), aquarium bred; ZMB
31349(4), aquarium bred; MZUSP [no number published]
(4), aquarium bred.

Current status: Valid as Kryptolebias caudomarginatus
(Seegers, 1984) (Costa 2004: 107).

©ZFMK

126 Fabian Herder, Jan Huylebrouck & Klaus Busse

Rivulus derhami Fels & Huber, 1985
Rev fr Aquariol 12: 33

Paratypes: ZFMK 2071—2090 (20), Universidad Agraria
de Tingo Maria, Pert, May 1966, K. H.-Liiling; ZFMK
7813 a (66), same locality as ZFMK 2071-2090, 28 Apr
1972, K. H. Liiling.

Type material in other collections: Holotype: MNHN
1985-577, male, 38.5 mm, area of the Universidad Agraria
de Tingo Maria, Pert, 23 Aug 1981, P. de Rham;
Paratypes: MNHN 1985-578 (2), females, same data as
holotype; MNHN 1985-579 (9), same data as holotype;
MNHN 1985-580 (5) aquarium bred, strock collected by
Liling; MNHN 1985-581 (4), 3 km east of Tingo Maria,
18 Oct 1976, A. C. Radda; MNHN 1985-582 (6), aquar-
ium bred, strock collected by P. de Rham.

Rivulus erberi Berkenkamp, 1989
Das Aquarium 23 (No. 238): 211

Paratypes ZFMK 14820-14822 (3), 35.0-39.4 mm, 7 km
east of Coca (= Puerto Francisco Orellana), near Rio Na-
po, Napo Prov., East Ecuador, 4°30°00”S 76°54’00” W,
8 Mar 1983, H. Heinemann, H. Heidt & K. Wilke.

Type material in other collections: Holotype: SMF
18432, male, 40.2 mm, same data as ZFMK 14820-14822;
Paratypes: SMF 18433, 26.7-35.0 mm, same data as
ZFMK 14820-14822; MHNG 22281.04 (2), 19.4-35.0
mm, same data as ZFMK 14820-14822.

Rivulus luelingi Seegers, 1984
Zool Beitr 28: 264

Holotype: ZFMK 11671, male, near Rio Pirai at Joinville,
Southeast Brazil, 25 Oct 1982, K. H. Luling.

Paratypes: ZFMK 11672, female, 20 km southwest of
Joinville, Southeast Brazil, 22 Sept 1980, K. H. Liiling;
ZFMK 11673-11682 (10), same locality as holotype, 10
Oct 1980.

Type material in other collections: Paratypes: SMF
17585 (9), same data as holotype; ZMB 31347 (3), same
data as holotype; BM, 1983.6.9. 14-15 (2), same data as
holotype.

Bonn zoological Bulletin 59: 109-136

POECILIDAE

Aplocheilichthys maculatus lacustris Seegers, 1984
Rev Zool afr 98: 76

Holotype: ZFMK 12323, male, Kibiti, about 140 km south
of Dar es Salaam, Tanzania, 26 Dec 1981, L. Seegers &
W. Clafen.

Paratypes: ZFMK 12324-12326 (3), same locality as holo-
type, 31 Dec 1980, L. Seegers, C. Kasselmann & G. Eg-
gers; ZFMK 12327-12334 (8), same locality, 26 Dec
1981, L. Seegers & W. ClaBen, fixed Aug 1982.

Type material in other collections: Paratypes: SMF
17689 (4), males, and SMF 17690 (1), female, aquarium
bred, fixed in Aug 1982; MRAC 83-21-P-1—4 (4), males
and MRAC 83-21-P-5 (1), female, aquarium bred, fixed
in Aug 1982; ZMB 31319 (4), aquarium bred, fixed in Oct
1982.

Current status: Valid as Lacustricola lacustris (Seegers,
1984) (Huber 1999: 58).

ADRIANICHTHYIDAE

Oryzias hadiatyae Herder & Chapuis, 2010
Raffles Bull Zool 58: 271

Paratypes: ZFMK 41540-41550 (11), 31.6-40.3 mm,
Lake Masapi, southern shore between 2°50.837’S
121°21.116’E and the lake’s outlet, approx. 600 m west-
wards of this position, Larona drainage, South Sulawesi
Province, Sulawesi, Indonesia, 15-16 Apr 2004, F.
Herder, J. Pfaender, J. Schwarzer & R. K. Hadiaty.

Type material in other collections: Holotype: MZB
18491, 34.2 mm, male, same data as ZFMK 41540-41550;
Paratypes: MZB 18492-18506 (15), 22.5-44.6 mm, same
data as ZFMK 41540-41550; ZSM 39760-39766 (7),
28.4-43.7 mm, same data as ZFMK 41540-41550; ZRC
51984 (4) 28-46.1 mm, same data as ZFMK
41540-41550.

CICHLIDAE

Aequidens patricki Kullander, 1984
Rev fr Aquariol 11: 2

Paratypes: ZFMK 10766-10767 (2), males, 97.7—104.5
mm, tributary of Rio Pachitea, Quebrada Pacal, near Pan-
guana, basin of Rio Pachitea, dept. Huanuco, Peru, about
09°37°S 74°56’W, 11 Sep 1976, K. H. Liiling; ZFMK

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 127

10769 (1), sex indet., 37.6 mm, Quebrada near Pangua-
na, basin of Rio Pachitea, about 09°37’S 74°56’ W, 10 Sep
1976, K. H. Ltiling; ZFMK 10770-10771 (2), juv.,
17.9-20.2 mm, tributary of Rio Llullapichis, basin of Rio
Pachitea, Sira mountains; 17 Sep 1976, A. Hanrieder &
W. Foersch; all dept. Huanuco, Peru.

Type material in other collections: Holotype: MHNG
2163.93 (1), female, 116.2 mm, small brook from a drying
pool, along the road from Pucallpa to Tingo Maria, 15 km
west of the village of Aguaytia, tributary of Rio Aguay-
tia, Coronel Portillo Prov., depto. Ucayali, Peru, about
09°02’S 75°31°W, 24 Aug 1981, P. de Rham, H. Nijssen,
H. J. Franke & C. Villanueva; Paratypes: NRM
A82/1976376.3458 (1), female, 93.8 mm (former number
ZFMK 10768), same data as ZFMK 10766-10767; addi-
tional paratypes in following collections: IRSNB 700 (1);
MHNG 2163.94 (1); NRM 17760[ex A82/1976376.3458];
ZMH 6427 (1), 6435 (2) (see Kullander 1986: 383).

Apistogramma arua Romer & Warzel, 1998
Aqua J Ichth and Aquat Biol 3: 46

Paratypes: ZFMK 18599 (1), female, 34.3 mm, same da-
ta as holotype, A. Kriiger, F. Warzel, M. Loew & R. Zgor-
niak; ZFMK 18601 (1), male, 34.5 mm; ZFMK 18605 (1),
female, 11.6 mm, 2.5 km upstream of the village of Arua
in a small right-bank affluent of Rio Arua, Para State,
Brazil, 14 Sep 1995, A. Kriiger, F. Warzel, M. Loew &
R. Zgorniak [specimens 18601 and 18605 were preserved
in Oct 1995 after keeping in aquaria].

Type material in other collections: Holotype: MZUSP un-
cat., male, 44.4 mm, about 2.5 km upstream of the village
of Arua in a small right-bank affluent of Rio Arua, tribu-
tary of Rio Arapiuns, Para State, Brazil, about 02°39°27”S
55°43°24”W, 14 Sep 1995, A. Kriiger & F. Warzel [pre-
served in May 1996 after keeping in aquaria].

Apistogramma baenschi Romer, Soares & Wohler, 2004
Das Aquarium 38 (No. 422): 16

Paratypes: ZFMK 39828-39831 (4), tributary of Rio
Shanushi, lower Rio Hualaga system, Peru, near 6°03’S
77°53’ W, Mar 2003, J. Melgar.

Type material in other collections: Holotype: MTD F
26965, same data as ZFMK 39828-39829. Paratypes:
CAS 219381—86 (6), 219388—92 (5); MTD F 26964 (1),
F 28102—08 (7), F 28109-14 (6); SMF 29765a—b (2),
29765c—d (2).

Bonn zoological Bulletin 59: 109-136

Apistogramma eunotus Kullander, 1981
Bonn zool Beitr 32: 183.

Holotype: ZFMK 10772, male, 49.9 mm, way to Aguay-
ta near Pucallpa, Pert, 3 Sep 1978, K. H. Luling.

Paratypes: ZFMK 10773, female; ZFMK 10774, juy.,
same data as holotype.

Type material in other collections: Paratypes: BM
1913.7.30: 56-57 (2), sex indet., Rio Ucayali, Pert, no
date, Mounsey; MCZ 15807 (2), Tabatinga Solimoes, East
Amazon river system, Sep—Oct 1865, Thayer-Expedition:
D. Bourget; MHNG 1583.50, female, Jenaro Herrera, Rio
Ucayali-system, distr. Loreto, Peru, 18 Oct 1977, P. de
Rahm; NRM 1971365: 3057, female, Lago Matamata,
distr. Loreto, Peri, 10 Oct 1971, T. Hongslo; NRM
1976312: 0914-0923 (10), “Cano Comprido”, San Sebas-
tian, Rio Yavari, distr. Loreto, Peru, 27 Jul 1976, T. Hongs-
lo; NRM 1876311: 1164-1168 (5), 26 Jul 1976, same da-
ta as previous specimens.

Apistogramma linkei Koslowski, 1985
Bonn zool Beitr 36: 151

Holotype: ZFMK 13323, male, 36.9 mm, northwest of
Santa Cruz, Bolivia, [type locality unclear]; Original de-
scription and label: In lagoons along the road between the
villages Portachuelo and Bellavista, 76 km northwest of
Santa Cruz, Bolivia, about 68°25’W 16°20’S, small wa-
ter bodies along the road and small and low watercourse
which crosses the road and lagoon at the road 2 km east-
ern of the village Japacani at Rio Japacani, northwest of
Santa Cruz, Bolivia, about 68°50’W 16°15’S, Jul 1983,
H. Linke & W. Staeck.

Paratypes: ZFMK 13324-13327 (4), 24.9-28.6 mm;
ZFMK 13328-13362 (35), all same data as holotype;
ZFEMK 13320-13322 (3), 18.5-27.0 mm, in a course of
a stream 4 km in front of Okinawa, west of Montero
Guabira in direction to Rio Grande, about 62°50’W
17°12’S, Jul 1983, H. Linke & W. Staeck; ZFMK
13363-13367 (5), 21.1-30.3 mm; ZFMK 13368-13393
(26), 16.0-24.4 mm, in lagoons and backwater along the
road from Trinidad to west, about 1 km in front of the vil-
lage of Pto. Amacen at the Rio Mamoreé, Bolivia, about
64°58’ W 14°53’S, 11 Jul 1983, H. Linke & W. Staeck;
ZFMK 13394-13399 (6), 21.5—27.0 mm, in lagoons and
remains of river water at the road from Trinidad to east,
about 10 km in direction to Peroto, east of the town
Trinidad, Bolivia, about 64°48’W 14°49’S, 12 Jul 1983,
H. Linke & W. Staeck; ZFMK 2268-2273 (6), pond like
backwater, 1.5 km downstream of San Francisco, Rio
Chipiriri, Bolivia, 4 Oct 1966, K. H. Liling; ZFMK 2303

©ZFMK

128 Fabian Herder, Jan Huylebrouck & Klaus Busse

(1), 25.2 mm, brook between Rio Chaparé and Rio
Mamore, Bolivia, 9 Oct 1966, K. H. Luling & A. Meyer.

Type material in other collections: Paratypes: MZUSP
28726 (8), 17.7—25.7 mm; NRM A 83/1983273.3046 (7),
16.3-34.7 mm, all same data as holotype; BMNH 1985,
1. 28 (5), 20.1—23.9 mm; IRSNB 724 (6), 18.6—25.0 mm;
NRM A 83/1983281.3054 (7), 16.3—25.1 mm; ZMA
119.629 (6), 18.6—23.7 mm, all west of the town Trinidad,
in lagoons and backwater along the road from Trinidad
to west, | km in front of Pto. Amacen at the Rio Mamo-
ré, Bolivia, about 64°58’ W 14°53’S, 11 Jul 1983, H. Lin-
ke & W. Staeck; NRM A 84/1983166/3061 (3), 19.7—27.2
mm, Laguna Suarez, near Trinidad, Rio Mamoré system,
depto. Beni, Bolivia, 23 Apr 1983, G. Loubens & L. Lau-
zanne; NRM A 84/1982118.3062 (5), 27.6-42.1 mm,
Arroyo San Juan, near Trinidad, depto. Beni, Rio Mamo-
ré system, Bolivia, Mar 1982, G. Loubens & L. Lauzan-
ne; IRSNB 695 (former IRSNB 19.975) (1), female, 25.6
mm, tributary to the Rio San Miguel, Rio Surucusi, road
from Ascesion, 14 km north of Limon, Rio Guaporé
system, depto. Santa Cruz, Bolivia, 9 Nov 1977, J.-P. Gos-
se.

Apistogramma luelingi Kullander, 1976
Bonn zool Beitr 27: 259

Holotype: ZFMK 2283, male, 26.4 mm, downstream of
Todos Santos, Bolivia, 3 Oct 1966, K. H. Liling & A.
Meyer.

Paratypes: ZFMK 2284-2291 (8), 18.2—-29.0 mm, same
data as holotype.

Apistogramma martini Romer, Hahn, Romer, Soares &
Wohler, 2003
Das Aquarium 37 (No. 406): 15

Paratypes: ZFMK 39412-39421 (10), small tributary of
the Rio Nanay near the village of Alvarenga (formerly
Puerto Alianza), distr. Santa Maria, Maynas Prov. (former-
ly Iquitos), dep. Loreto, Peri, about 74°25°40”W
03°31°10’S, 19 Oct 2002, M. Mortenthaler.

Type material in other collections: Holotype: MTD F
26849 (1), female, 30.2 mm, same data as ZFMK
39412-39421; Paratypes: MTD F 26850-26858 (9); MTD
F 26945-26963 (19); MTD F 26957-26958 (2); CAS
216883-87 (5), males; CAS 216878—82 (5), females; SMF
2933 1la—e (5), males; SMF 29332a-—e (5), females; ZMB
33329-33333 (5), males; ZMB 33334-33338 (5), females;
all same data as ZFMK 39412-39421.

Bonn zoological Bulletin 59: 109-136

Apistogramma mendezi Romer, 1994
Aqua J Ichth and Aquat Biol 1: 5

Holotype: ZFMK 17458, male, 34.5 mm, Rio Salgado, ne-
ar Barcelos do Rio Negro, Dept. Amazonas, Brazil, about
0°01’°S 63°04’W, 19 Oct 1991, U. Romer.

Paratypes: ZFMK 17459 (1), female; ZFMK 17462 (1),
male; 17502 (1), male; 17461 (1), male [cleared and stai-
ned], all same data as holotype, U. Romer, A. Schneider
& W. Windisch; ZFMK 17460 (1), male; ZFMK 17463
(1), male; ZFMK 17501 (1), male; ZFMK 17504 (1), ma-
le, all same data as holotype; ZFMK 17503 (1), male, sa-
me locality as holotype, 22 Aug 1992, M. Geismann, S.
Leissner, U. Romer & A. Schneider [ZFMK 17460, 17463,
17501, 17504 were preserved after 6 months of keeping
in aquaria].

Type material in other collections: Paratypes: INPA UR
91024 (1); INPA UR 91027 (1); INPA UR 91169 (1); IN-
PA UR 91209 (1); all same data as holotype, U. Romer,
A. Schneider & W. Windisch; INPA UR 91170 (1), male;
INPA UR 91172 (1), male; all same locality as holotype,
22 Aug 1992, M. Geismann, S. Leissner, U. R6mer & A.
Schneider; INPA UR 91205 (1); INPA UR 91217 (1); all
same data as holotype [preserved after six months of keep-
ing in aquaria].

Apistogramma panduro Romer, 1997
Buntbarsche Bulletin 182: 12

Holotype: ZFMK 18610, male, 37.5 mm, small forest
brooks east of Jenaro Herrera aside the road heading to
the Brazilian border at kilometers 26 and 27, Rio Ucay-
ali-system, Prov. Loreto, Peru, Jun 1996, professional col-
lectors of exporter Jesus Victoriano Panduro Pinedo, ded-
icated in early June 1996 after import by Aquarium Glaser
(Rodgau, Germany), preserved immediately after import
by F. Schafer and U. Romer.

Paratypes: ZFMK 18580-18582 (3), 25.6-29.5 mm;
ZFMK 18609 (1), male, 43.7 mm; ZFMK 18611-18616
(6), 25.7-48.4 mm; ZFMK 18620 (1), male, 38.3 mm, all
same data as holotype.

Type material in other collections: Paratypes: SMF
28206—08 a—d (6), same data as holotype.

Apistogramma paucisquamis Kullander & Staeck, 1988
Cybium 12: 190

Paratypes: ZFMK 15496-15500 (5), arquipiélago das
Anavilhanas, small shallow bay on the right bank of Rio

©OZFMK

Type specimens in ZFMK: Section of Ichthyology 129

Negro, Estado do Amazonas, Brazil, 27 Mar 1986, W.
Staeck.

Type material in other collections: Holotype: MZUSP
36952 (1), male, same data as ZFMK 15496-15500;
Paratypes: NRM A 86/1986134.3587 (15); USNM 289281
(5); MZUSP uncat. (4), all together 16.4-28.9 mm, same
data as holotype; NRM A 86/1986134.3594 (5), males,
26.2—31.2 mm, baylet with small clear water creek at right
bank of Rio Negro, arquipiélago das Anavilhanas, Brazil,
27 Mar 1986, W. Staeck; MZUSP 28210 (57); NRM A
83/1980077.3058 (4), downstream of Rio Dara, central
lake in Rio Negro, 17 Feb 1980, M. Goulding.

Apistogramma rubrolineata Hein, Zarske & Zapata, 2002
Das Aquarium 36 (No. 402): 16

Paratypes: ZFMK 39241-39242 (2), 23.5-26.4 mm, near
the village of Puerto Cardenas, Rio Manuripi, Departa-
mento Pando, Bolivia, about 11°20°083”S 67°44°576”"W,
12 May 2001, G. Hein, H. Langert, J. Zapata & A. Zarske.

Type material in other collections: Holotype: MTD F 26
477, 40.0 mm; Paratypes: CIRA UTB 970 a-e (5),
17.0-25.7 mm; MTD F 26 478-26-486 (9), 14.5-40.0
mm; NMW 94 645-94 646 (2), 26.5—28.0 mm, all same
data as ZFMK 39241-39242.

Apistogramma similis Staeck, 2003
In Staeck: Cichliden Lexikon 3: 76

Paratypes: ZFMK 39800-39801 (2), forest brook at the
Laguna La Brava, Rio Yata system, Beni Prov., Bolivia,
2001, W. Staeck.

Type material in other collections: Holotype: MTD F
27038, 38.1 mm; Paratypes: MTD F 27039 (1); MTD F
27482-27484 (3); additional paratypes at NVW and ZMB
[no collection numbers provided]; all same data as ZFMK
39800-39801.

Apistogramma staecki Koslowski, 1985
Bonn zool Beitr 36: 146

Holotype: ZFMK 13400, 20.8 mm, male, in lagoons on
both sides of the road, 10 km south of Trinidad in the di-
rection of El Colegio and Loreto, Bolivia, about 64°51’ W
14°57’S, 12 Jul 1983, H. Linke & W. Staeck.

Paratypes: ZFMK 13401—13404 (4), 17.7-20.8 mm, same
data as holotype; ZFMK 13405-13410 (6), 19.5-20.7 mm,
in lagoons with a remain of river water, about 10 km east
of Trinidad in direction of Peroto, Bolivia, about 64°48’ W
14°49’S, 12 Jul 1983; same collectors as holotype.

Bonn zoological Bulletin 59: 109-136

Type material in other collections: SMF 18855 a—h (8),
14.7-20.4 mm; MZUSP 28725 (2), females, 15.0—19.4
mm; NRM A 83/1983282.3053 (7), 15.7—20.9 mm, all
same data as holotype.

Apistogramma tucurui Staeck, 2003
In Staeck: Cichliden Lexikon 3: 112

Paratype: ZFMK 39802 (1), brook in Rio Tocantins sys-
tem, at highway 150, 10 km south of Jacunda, Para, Brazil,
1989, W. Staeck.

Type material in other collections: Holotype: MTD F
27046; Paratypes: MTD F 27047 (1); MTD F
27487-27488 (2); additional paratypes at NVW and ZMB
[no collection numbers provided]; all same data as ZFMK
39802.

Bujurquina huallagae Kullander, 1986

In Kullander (1986): Cichlid Fishes of the Amazon Riv-
er drainage of Peru. Swedish Museum of Natural Histo-
ry, Stockholm: 290

Paratypes: ZFMK 2152-2168 (17), lower course of Rio
Subte, downstream of Tingo Maria, about | km in front
of the mouth into the Rio Huallaga, 30 Jun 1965; ZFMK
2184-2187 (4), lower Rio Monzon, 10 km upstream of
Tingo Maria, 3 Jun 1966; ZFMK 2318 (1), brook near Tin-
go Maria, just before the hospital, 500 m in front of the
mouth into the Rio Huallaga, 2 Jun 1966; ZFMK
8659-8664 (6), near the hospital, Tingo Maria, 12 Nov
1974; ZFMK 2190 (1), left bank of Rio Huallaga, oppo-
site to Tingo Maria, 22 Jun 1966; all Huallaga river sys-
tem, departamento Huanuco, Peru, all leg. K. H. Liling.

Type material in other collections: Holotype: ANSP
158171 (1), male, 75.4 mm, about 0.25 miles above Rio
Huallaga, vicinity of Tingo Maria, Rio Huallaga drainage
system, dept. Huanuco, Pert, 30 Sep 1955, Catherwood
Expedition, C. C. G. Chaplin, M. H. Hohn; additional 45
paratypes in following collections: ANSP 139109 (1),
139110 (11), 139112 (3), 139114 (5), 139116 (3), 139143
(1), 158172 (9); MHNG 2233.66 (1), 2205.01 (4),
2205.06 (4); NRM 17594 (1) (see Kullander op. cit.:
375-376).

Cichlasoma amazonarum Kullander, 1983
Revision of the South-American cichlid genus Cichlaso-
ma. Swedish Museum of Natural History, Stockholm: 117

Paratypes: ZFMK_ 13451-13460 (10), brook Aguajal,
Ucayali river system, 3 Sep 1978; ZFMK 2225-2230 (6),
Chauahuan Cocha, Ucayali-system upstream of Pucallpa,

©ZFMK

130 Fabian Herder, Jan Huylebrouck & Klaus Busse

Aug 1966; ZFMK 2300-2301 (2), Yarina Cocha, Ucay-
ali-system, 19 Jul 1966; ZFMK 1125-1126 (2), Cafio Ya-
rina, depto. Loreto, Pert; 31 Aug 1959; ZFMK 8655-8656
(2), between Paca Cocha and Yarina Cocha, 9 Aug 1974;
ZFMK 8657-8658 (2), Quebrada near Yarina Cocha, 9
Aug 1974; ZFMK 2191-2192 (2), Yarina Cocha, 16 Jul
1966; ZFMK 2205-2211 (7), Yarina Cocha, 19 Jul 1966;
ZFMK 2233 (1), Rio Ucayali, opposite of Pucallpa, 12 Jul
1966; ZFMK 1118-1120 (3), bank of Amazon upstream
of Iquitos, 8-16 Aug 1959; ZFMK 7802-7806 (5), Quis-
to Cocha, Amazon system, depto. Loreto, 10 May 1972;
all from Peru, all leg. K. H. Liiling.

Type material in other collections: Holotype NRM (SOK)
3651, floating meadow near mouth, right bank tributary
of the Rio Amptyacu, Sacarita del Tuyé, little upstreams
of Pebas, Ampiyacu system, depto. Loreto, Peru, 12 Aug
1981, S. O. Kullander & A. Hogeborn; additional
paratypes in following collections: ANSP, BM, FMNH,
IRSNB, MCZ, MHNG, MNHNP, MZUSP, NHMW,
NRM, USNM and ZMA [there are discrepancies between
published numbers and museum records].

Cichlasoma boliviense Kullander, 1983
Revision of the South-American cichlid genus Cichlaso-
ma. Swedish Museum of Natural History, Stockholm: 165

Paratypes: ZFMK 2199 (1), downstream of Pto. San Fran-
cisco, Rio Chipiri, Mamoré system, depto. Santa Cruz, Bo-
livia, 4 Oct 1966, K. H. Liiling; ZFMK 2275-2282 (8),
and ZFMK 2299 (1), Todos Santos “Hoffmann-lagoon”’,
Rio Chapare, Mamoré system, Bolivia, 7 Oct 1966, K. H.
Luling; ZFMK 2219-2221 (3), 4 km downstream of To-
dos Santos, Bolivia, 8 Oct 1966, K. H. Liiling; ZFMK
2212-2213 (2), downstream of Pto San Francisco, Boli-
via, 3 Oct 1966, K. H. Liiling; ZFMK 2305-2310 (6), 4
km downstream of Todos Santos, Bolivia, 30 Sep 1966,
K. H. Lilling; ZFMK 13461—13464 (4), Rio Tambopata
near Puerto Maldonado, Madre de Dios system, Pert,
25—28 Apr 1981, E. Lenkenhoff; ZFMK 13465 (1), Puer-
to Maldonado, Peru, 6 Apr 1981, E. Lenkenhoff.

Type material in other collections: Holotype: IRSNB 677
(1), male, 75.9 mm, Rio Uruguaito, River Guaporé basin,
tributary of Rio Quizer, 13 km south of San Xavier, dep-
to. Santa Cruz, Bolivia, 4 Nov 1977, J.-P. Gosse; addition-
al paratypes in following collections: ANSP, BM, FMNH,
IRSNB and USNM.

Crenicichla sedentaria Kullander, 1986
Cichlid Fishes of the Amazon River drainage of Peru.
Swedish Museum of Natural History, Stockholm: 83

Bonn zoological Bulletin 59: 109-136

Paratypes: ZFMK 2141 (1), branch of Rio Huallaga near
Aspusana, 70 km downstream of Tingo Maria, Pert; 19
Jun 1966, K. H. Liling; ZFMK 2142-2143 (2), Rio Hual-
laga near Aspusana, same data as ZFMK 2141; ZFMK
10765 (1), branch of Pacal near Panguana, Llullapichis,
Pachitea, Pert, 11 Sep 1976, K. H. Luling.

Type material in other collections: Holotype: USNM
229057 (1), probable male, 104.4 mm, near San Pedro de
Longin, Rio Yamushimas, Rio Pachitea drainage system,
depto. Pasco, Pert, 24 Jul 1981, P. Bayley. The total num-
ber of paratypes amounts to 83 specimens in the follow-
ing collections: ANSP (59 in 8 lots); BMNH 1969.7.15.55
(1); 1969.7.23.4 (1); CAS 57455 (1); FMNH 84259 (2);
84268 (3); FMNH (1) [no number provided]; MCZ 48326
(1); MHNG (3 in 2 lots); NRM 12293 (2); 13577 (2);
13778 (1); USNM 229053 (1); ZMH 6443 (2).

Neolamprologus cylindricus Staeck & Seegers, 1986
Aquar Terrar Z 39: 448

Holotype: ZFMK 14595, 83.6 mm, Lake Tanganyika,
south of the mouth of the Kalambo River, near Chipwa,
Zambia, 8°38’S 31°14’E, 3 Aug 1986, W. Staeck.

Paratype: ZFMK 14596 (1), 66.1 mm, same data as holo-
type, L. Seegers.

Paratheraps breidohri Werner & Stawikowski, 1987
Aquar Terrar Z 41: 20

Holotype: ZFMK 15524, juv., 59.6 mm, Presa de Angos-
tura, below the dam wall, Chiapas, Mexico, 16 Apr 1987,
U. Werner, R. Stawikowski, E. Sosna & I. Kranz.

Paratypes: ZFMK 15525-15531 (7), 40.3—73.5 mm, same
data as holotype.

Remarks: Type species of Paratheraps Werner &
Stawikowski, 1987

Current status: Valid as Vieja breidohri (Werner &
Stawikowski, 1987) (Burgess 2000: 50).

Pseudocrenilabrus multicolor victoriae Seegers, 1990
Aquar Terrar Z 43: 100

Holotype: ZFMK 16252, male, 60.8 mm, stream which
connects two marsh districts on both sides of the road
Kampala-Masaka, southwest of Mpugwe, about 14 km
notheast of Masaka, distr. Masaka, Southwest Uganda,
about 0°36’N 31°41’E, [no collection date provided], U.
Renninger & L. Seegers.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 131

Paratypes: ZFMK 16253-16255 (3), 51.9-69.8 mm, same
data as holotype; ZFMK 16256-16263 (8), 18.9-49.3 mm,
way from Lake Nabugabo to Masaka, 10 km northwest
of Lake Nabugabo, West Uganda, 26 Mar 1988; ZFMK
16264-16265 (2), males, 30.9-32.4 mm, stream crossing
road to Kabale about 27 km west of Mbarara and 6 km
east of Kabwohe, West Uganda, 26 Mar 1988; ZFMK
16266-16268 (3), 33.2-49.3 mm, near Kyenjojo, 2 km in
front of the branch of the road from Fort Portal to Kam-
pala and Hoima, West Uganda, 2 Apr 1988; ZFMK
16269-16272 (4), 26.3-41.9 mm, west of Ngogwe, road
which leads south of the main street Kampala-Jinja from
Mukono via Nkokonjeru to Jinja, Uganda, 5 Apr 1988;
ZFMK 15313 (1), male, 30.2 mm, in Ahero in a pond west
of the branch of the federal road Al and B1 to Kisumu,
Kericho and Kisii, Kenia, | Jan 1987, all same collectors
as holotype.

Type material in other collections: Paratypes: ZMB [no
collection numbers provided] (3), 52.2—63.1 mm, same da-
ta as holotype; ZMB [no collection numbers provided] (2),
44.2—62.2 mm, same data as ZFMK 16256-16263; SMF
23436 (3), 34.5-58.1 mm, same data as ZFMK
16256-16263; SMF 23437 (5), juv., 18.3-32.0 mm, same
data as ZFMK 16264-16265.

Theraps coeruleus Stawikowski & Werner, 1987
Aquar Terrar Z 40: 499

Holotype: ZFMK 15501, 77.0 mm, little tributary to Rio
Mizol Ha, at Francisco Madero, west of the road from
Palenque to Ococingo, about 30 km south of Palenque,
Mexico, 21 Apr 1987, I. Kranz, E. Sosna, R. Stawikows-
ki & U. Werner.

Paratypes: ZFMK 15502-15505 (4), 60.5—66.6 mm, same
data as holotype.

Theraps rheophilus Seegers & Staeck, 1985
Aquar Terrar Z 38: 500

Holotype: ZFMK 13928 (1), probable male, 88.7 mm,
Chiapas, 6 km south of Palenque, under the bridge of the
road to Ococingo in Chiapas, Rio Notutun, Mexico, 2 Apr
1983, L. Seegers & W. Staeck.

Paratypes: ZFMK 13929-13930 (2), 76.5—84.1 mm, same
data as holotype.

Type material in other collections: Paratypes: NRM 17598
(2); NMW 81184 (2); all same data as holotype.

Bonn zoological Bulletin 59: 109-136

Current status: Synonym of Theraps lentiginosus Stein-
dachner, 1864 (Kullander in Reis et al. 2003: 643).

Tilapia eisentrauti Trewavas, 1962
Bonn zool Beitr 13: 168

Paratypes: ZFMK 732 (1), female, Lake Barombi-ma-
Mbu, Cameroon, 1954, M. Eisentraut.

Type material in other collections: Holotype, male, 65.0
mm, and paratype, female, BM 1961.10.18.11—14.

Remarks: Type species of Konia Trewavas, 1972.

Current status: Valid as Konia eisentrauti (Trewavas,
1962) (Teugels 1991: 206).

Uaru fernandezyepezi Stawikowski, 1989
Bonn zool Beitr 40: 21

Holotype: ZFMK 15757, 180 mm, branch of Rio Atabapo,
5 km upstream of its mouth into the Rio Orinoco,
Venezuela, 29 Apr 1988, H. J. Kopke.

Paratype: ZFMK 15758 (1), 169 mm, same data as holo-
type.

ELEOTRIDAE

Mogurnda pulchra Horsthemke & Staeck, 1990
Aquar Terrar Z 43: 160

Holotype: ZFMK 16247: female, 60.3 mm, small river in
the drainage basin of the Brown River, at the Hiritano
Highway, 24 km north of Port Moresby, Papua New Gui-
nea, about 09°16’S 147°14’E, Jul 1988, W. Staeck.

Paratypes: ZFMK 16248-16251 (4), 22.0-5S8.1 mm, same
data as holotype.

GOBIIDAE

Knipowitschia cameliae Nalbant & Otel, 1995
Ocrot Nat Med Inconj Bucaresti 39: 51

Paratypes: ZFMK 22151-22152 (5), 1 adult or subadult
and 4 very small specimens, small lake in sand spit, 1 km
north of Portita lighthouse, between Lake Golovitsa and
Black Sea, Portita, Dobrogea, Romania, 8 Jun 1994, Nal-
bant & Otel.

©ZFMK

132 Fabian Herder, Jan Huylebrouck & Klaus Busse

Type material in other collections: Holotype: IBTS 4587,
same data as ZFMK 22151-22152; Paratypes: IBTS 4586
(30), IBTS 4588 (6), ICDD uncat. (54), MGAB uncat.
(51).

Remarks: ZFMK 22151-22152 are out of IBTS 4588.

OSPHRONEMIDAE

Betta climacura Vierke, 1988
Das Aquarium 22: 336

Holotype: ZFMK 15532, male, 66 mm, small clearwater
brooks about 60 air-miles southwest of Brunei, near Ram-
payoh, West Borneo, Jul 1983, T. Schulz.

Paratype: ZFMK 15533 (1), female, 65 mm, same data as
holotype.

Current status: Synonym of Betta akarensis Regan, 1910
(Kottelat et al. 1993: 161).

Betta persephone Schaller, 1986
Aquar Terrar Z 39: 298

Holotype: ZFMK 14226, male, Asian highway no. 2, 3 km
north of Ayer Hitam, Malaysia, 20 Feb 1985, D. Schaller.

Paratypes: ZFMK 14227-14238 (12), same data as holo-
type.

Type material in other collections: ZMB 31614 (4);
ZSM/CMK 5944 (4); same data as holotype.

Betta tussyae Schaller, 1985
Aquar Terrar Z 38: 350

Paratypes: ZFMK 14224-14225 (2), about 77 km south
of Kuantan at the road parallel to east coast, 17 km south
of Pekan, about | km from sea, Pahang State, Malaysia,
1984, P. Nagy; ZFMK 14195 (1), male, 16 km south of
Kuantan, Malaysia, 22 Feb 1985, D. Schaller; ZFMK
14196-14207 (12), same data as ZFMK 14195; ZFMK
14208-14212 (5), same data as ZFMK 14195.

Type material in other collections: Holotype: ZRC 38451
(ex ZSM 27336); Paratypes: ZSM/CMK 5945 (5);
ZSM/CMK 6345 (11); ZMB 31607 (6); ZMB 31609 (1).

Remarks: Originally the specimen ZFMK 14195 was de-
posited as holotype by the author in Bonn, but it cannot
be identical to the specimen which was characterized as

Bonn zoological Bulletin 59: 109-136

holotype by measurements in the original description be-
cause the ZFMK specimen is smaller. Nevertheless this
specimen belongs to the series of paratypes.

Macropodus erythropterus Freyhof & Herder, 2002
Ichthyol Explor Freshwaters 13:162

Holotype: ZFMK 38945, 58.9 mm, stream at Quyét Tién,
Quang Binh Prov., Vietnam, 17°11.54’N 106°37.53’E, 8
Apr 2000, J. Freyhof & F. Herder.

Paratypes: ZFMK 38946-38961 (16), 23.7—-5S7.3, same da-
ta as holotype; ZFMK 38962-38973 (12), stream at Lé
Ninh, Quang Binh Prov., Vietnam, 17°14.12’N
106°37.51°E, 8 Apr 2000, J. Freyhof & F. Herder; ZFMK
35462-36465 (4), stream at Lé Ninh, Quang Binh Prov.,
Vietnam, 17°13.16’N 106°39.08’E, 8 Apr 2000, J. Frey-
hof & F. Herder.

Remarks: Winstanley & Clements (2008) were not able
to follow unambiguously the diagnostic colour characters
used by Freyhof & Herder (2002) for discriminating
erythropterus from M. spechti, and suggested that M. ery-
thropterus constitutes a junior synonym of M. spechti. As
their analyses did however (i) not incorporate material
from the terra typica (streams Quyét Tién and Lé Ninh,
Quang Binh Province, Vietnam), (11) were based on only
limited numbers of obviously mostly immature specimens
(Nota) = 22 ex., without reference to maturity or sex;
SLinax = 42.3 mm SL; four samling locations incorporat-
ed: AK 27598, 6 ex.; 13.7—38.0 mm SL; AK 27599, 3 ex.,
19.1-26.0 mm SL; AK 27600, 3 ex., 26.0-34.1 mm SL;
AK 27601, 10 ex., 26.5-42.3 mm SL; compare the sizes
of specimens examined by Freyhof & Herder (2002) as
listed above), and (111) covered only locations from a part
of the species distribution (only locations in the Quang Tri
Province in Vietnam), we argue that this conclusion is pre-
mature. Macropodus erythropterus is, like many other
teleost fish species, distinguished from other species of its
genus mainly by colour characters present in adult spec-
imens. As long as these are not incorporated properly in
critical analyses, we consider the database for judging
about species status insufficient. Hence, we suggest that
M. erythropterus should be treated as valid species.

Parosphromenus nagyi Schaller, 1985
Aquar Terrar Z 38: 302

Lectotype: ZFMK 14186, Asian highway route 18, 16 km
south of Kuantan, Malaysia, 22 Feb 1985, D. Schaller.

Paralectotypes: ZFMK 14187-14191 (5), same data as lec-
totype.

©ZFMK

Type specimens in ZFMK: Section of Ichthyology 133

Type material in other collections: Paralectotypes: ZMB:
31605 (2); CMK 5949 (2).

Remarks: In Schaller’s description the animals were still
alive; later Schaller deposited the ZFMK specimens as
holotype and paratypes. In the original description no
holotype was mentioned, so that all specimens are con-
sidered as syntypes (Schaller & Kottelat 1990). Schaller
and Kottelat (1990), designated a lectotype, the remain-
ing syntypes became accordingly paralectotypes.

CHANNIIDAE

Channa bleheri Vierke, 1991
Das Aquarium 25 (No. 259): 22

Holotype: ZFMK 16555, male, 109 mm, in muddy re-
mains of pools of river beds which are dried out during
the dry season, upper Diburu near Guijam, Assam, India,
12-13 Nov 1987, H. Bleher & D. Nopak.

Paratype: ZFMK 16556 (1), female, 82 mm, same data as
holotype.

Acknowledgements. We are greateful to Mario Milanovic for
assistance in assembling catalogue data.

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ples of reference style are:

Kottelat M, Whitten T, Kartikasari SN, Wirjoatmodjo S (1993)
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of ricefishes, Oryzias and relatives (Beloniformes, Adrianichthyi-
dae). Zoological Journal of the Linnean Society 154: 494-610

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(U.S. Department of Agriculture, Forest Service, Rocky Moun-
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phology of two cryptic bat species: Pipistrellus pipistrellus and
P. pygmaeus —a 3D geometric morphometrics approach with land-
mark reconstruction. Acta Chiropterologica 11: 113-126

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re
1

Contents “wil

Hutterer, Rainer & Gustav Peters: 3
Type specimens of mammals (Mammalia) in the collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Elzen, Renate van den: 29
Type specimens in the bird collections of the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Bohme, Wolfgang: 79
A list of the herpetological type specimens in the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Herder, Fabian, Jan Huylebrouck & Klaus Busse: 109
Catalogue of type specimens of fishes in the
Zoologisches Forschungsmuseum Alexander Koenig, Bonn

Cover illustration:

Picus (Dendrocopus) canariensis Koenig, 1889 (= Dendrocopos major canariensis (Koenig, 1889)),
Syntype, ZFMK E.1.5.a4.a

(this volume, pp. 29-77)

Ministerium fiir Innovation,
Wissenschaft und Forschung
des Landes Nordrhein-Westfalen

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/% i Leibniz
@. Gemeinschaft