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CONTENTS VOLUME IV.

or Netewcrthy Philippine Plents, V.

1.c. 169-£46.

or Notewcrthy Philippine Plants VI,

cr Neteworthy Philippine Plants,
VIL; Ise@« 4.(1909). Bat. 247-330.

cr Notewcrthy Philippine Plants,
VIII; 1.0.5 (2910) Bot, BG7+e67.

cr Notewcrthy Philippine Plants,
Im, Leese FLEA) Bats 259-587,

cr Notewcrthy Philippine Plants,
X, l.c. 9 (1914) Bot. 261-337.

cr Netewcrthy Philippine Plants,
KI, lo. 10 (4915) Bot. 1-84.

cr Neteworthy Philippine Plents, |
XII, 1.c. 10 (1915) Bot. 287-349.

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“ LIBRARY

YO 130%

THE PCIe INE

JOURNAL OF SCIENCE

Vou. I AUGUST 15, 1906 SuppLEMENT III

NEW OR NOTEWORTHY PHILIPPINE PLANTS, V.

By Evmer D. MERRILL.

(From the botanical section of the Biological Laboratory, Bureau of Science.)

INTRODUCTION.

The present article has been prepared as material and data became
available, as was the case with the previous papers published under the
above title! Our receipts of Philippine botanical material from various
sources approximate 5,000 numbers annually and the percentage of un-
described species and interesting forms is very large, as is to be expected
in a region botanically so little known as is this Archipelago. In making
the preliminary identifications, immediately after the receipt of the
material in the herbarium, many of the interesting forms are detected
and described, but in this paper, as in the preceding ones of the series,
only a portion of these forms are considered. Practically no work in some
groups can at present be done in Manila, owing to a lack of literature ;
in certain ones, all material is submitted to specialists for identification
and report. Frequently, specific identifications can not be made at once,
because of a lack of complete material, so that several hundred sheets,
apparently representing undescribed species, remain in the herbarium of
this Bureau and for one reason or another these will need to be considered
at a later date. In general, it is not considered good policy to describe
species based on a single specimen and many of the apparently un-
described plants at present in our herbarium have no corresponding
duplicate material.

1 Publications of the Bureau of Government Laboratories (1904-5), 6, 17, 29,

35.
44772 169

170

In this paper a large number of genera which were previously unknown
from the Philippines are for the first time reported, their representa-
tives in some cases being species heretofore undescribed and, in others,
well-known plants which for the first time have been encountered in
the Archipelago.

Rolfe * makes it clear that the Philippine flora has distinct connections
with that of the neighboring lands, but, at the time he wrote, compara-
tively little was known regarding this flora or that of the regions im-
mediately to the north and south. In the past twenty years much work
has been done by various authors; 1905 has seen the completion of
Forbes and Hemsley’s work on the flora of China,* and in 1898 Koorders
published his compilation of that of Celebes.*

Rolfe, in 1884, knew but three species common to the Philippines and
Celebes and confined to these two regions and only about sixteen which
he considered to represent northern or continental types; therefore it
has been thought advisable here to enumerate some of the more striking
representatives both of the southern connection with Celebes and of the
northern one with Formosa, Japan, and the Asiatic Continent, especially
as Usteri,® publishing as recently as the year 1905, has added nothing
to the twenty-year-old list of Rolfe.

The list of species confined to Celebes and the Philippines and com-
mon to both has been extended from the three mentioned in 1884 to
about sixty given below, while many others extending from the Philip-
pines through Celebes to neighboring islands, have been enumerated.
The list, so far as Celebes is concerned, is based largely on WKoorders’
work, cited above. Koorders® remarks that the relationship between
the Philippine and Celebes flora is very prominent, but does not discuss
the connections in detail.

In the following table the cross (+) indicates that the species is
found in Luzon and Celebes and, unless noted to the contrary in the
last column, that it is known only from the two regions. The dash (—)
indicates that the Philippine species is represented in Celebes by a
closely related one or a variety. The question mark indicates a Philip-
pine species doubtfully credited to: Celebes by Woorders. Where the
range of the species is known to extend beyond Celebes, its distribution is
viyen in the last column. Specimens of most of the species enumerated
are in the herbarium of this Bureau.

“On the Flora of the Philippine Islands and Its Probable Derivation, Journ.
Linn. Soe. Bot. (1884), 21, 283-316.

‘An Enumeration of All the Plants Known from China Proper, Formosa, Hai-
nan, Corea, the Luchu Archipelago, and the Island of Hongkong, ete, Journ.
Linn. Soc. Bot. (1886-1905), 23, 26, 36. :

'Verslag eener Botanische Dienstreis door de Minahasa, tevens eerste overzicht
der Flora van N. O. Celebes, Meded. ’s Lands, Plant. (1898), 19.

* Beitriige zur Kenntnis der Philippinen und ihrer Vegetation (1905), 3.

® Loc. cit., 258. :

ier

Table showing floristic relationship between the Philippines and. Celehes.

Species. nee Celebes. Other distribution.

POLYPODIACE:
Polypodium angustatum Blume____________
Polypodium vuleanicum Blume___.________
Polypodium heracleum Kunze _______--____
Polypodium setosum Blume ______----_____-
Polypodium stenophyllum Blume___-_____-
Polypodium decrescens Christ____---__--___
Lindsaya sarasinorum Christ ~----------____
Lindsaya retusa Mett=—--------==2--- == 52
Callipteris alismaefolia J. Sm__-----_-______
Diplazium deltoideum Christ____--_________
Athyrium sarasinorum var. philippinense
ChTISt= ©2823 25 Sos Se eee
Pteris heteromorpha Fée _-_----_------_-__-
Hymenolepis platyrhynchos Kunze ______--
Niphobolus sphaerostichus Copel__-----_-_-
Aspidium aculeatum var, batjanense Christ.
CYPERACE:
Cyperus pubisquama Steud ___-_-__-_________
ARACE:
Epipremnum elegans Engl __-------------_-
Pothoidium lobbianum Schott______________
Scindapsus argyraea Engl --_----------_-__-
Spathiphyllum commutatum Schott +. Amboina.
DIOSCOREACE:
Dioscorea n. sp
MYRICACE#:
Myrica javanica Blume
FAGACE:
Quercus campanoana Vid -__--_-----------_ oh de
QiercusiblancolgAy DC 2es== sss -nse =n + ae
Quercus celebica Blume ~_________-_--__---- ae te
ULMACE:
Celtis philippinensis Blanco ______~-_-_-___- +
URTICACE:
MIeuUs eCaRUlOCcanpa NL dees =a =e 5 een
Hicusichnysolepis#Mides=s-"=-"4- 5 seen
IGS ClUSIOIG GsyIMll Gane e en ak ee eee

=f Java.
Sunda Islands.
Java.
Java.
Java.

New Guinea.

pep tb pte eet

t++++4
+++ +4

Batjan.

4
Le

Moluceas, Ternate.

He ge
4

ae
+

Jaya.

Borneo.

Ficus manillensis Warb
Ficus minahassae Miq

+
+
+
hicusHforstenilMige-sae- 22. so. se ae
+
+

PITTOSPORACEE:
Pittosporum resiniferum Hemsl
LEGUMINOS#:
Clianthus binnendyckianus S. Kurz________
Dalbergia minahassae Prain_____-_---______
Gileditschis rolfe re Vic ea see es eae eee
Pithecolobium subacutum Benth____--_____
Pterocarpus echinatus Pers ________________-
Wallaceodendron celebicum Koord ________
RUTACE:
TUN Asla AMATAN DAN CO essen ee

+

ak

Timor, Borneo.

Ceram (?).

Java.

Halmaheira.

af

9

Table showing floristic relationship between the Philippines and Celebes—Continued.

Species.

BURSERACE.®: |
Garuga abilo)(Blco:) Meérr——-------_---=—_-_

MELIACE:

Reinwardtiodendron merrillii Perk___-----

Aglaia macrobotrys Turcz ------------------ |

Aglaia argentea Blume---------------------

Aglaia monophylla Perk ~------------------
MALPIGHIACE-E:

Ryssopteris dealbata Juss_------------------
EUPHORBIACEE: |

Actephila gigantea Koord____--------.-----

Cleistanthus’ eupreus Vid==—.<.-- --_---- ==}

Mallotus cumingii Muell. Arg__------------

Mallotus leucocalyx Muell. Arg_----..------

Glochidion album Boerl-_-_------------------

Macaranga hispida Muell. Arg -------------
ANACARDIACE:

Semecarpus perrottetii March _____------__-

Koordersiodendron pinnatum Merr ___--__-
SAPINDACE:

Cubilia blancoijBlume=—-=-=<-----=- 22 =e

Tristira pubescens! Merr =2==2=-0===-- 22
ELAEOCARPACE:

Elaecarpus cumingii Turcz ----_------------
TILIACE:

Columbia serratifolia DC ~----.-_--_--------
STERCULIACE:

Sterculia oblongata R. Br ~--.--==--=_-.-_-

Sterculia stipularis R: Br=__--22222-"S--
THEACEZ: |

Ternstroemia toquian F. -Vill -----.--------
MELASTOMATACE:

Medinilla cumingii Naud ---------------_--

Medinilla teysmannii Miq---.--------------

Astronia cumingiana Vid_------------------

Memecylon preslianum Triana__--_--_-___-
ARALIACEZE:

Tetraplasandra philippinensis Merr____----

Anompanax philippinensis Harms____—--_-|

Boerlagiodendron pulcherrimum Harms___
ERICACE:

Vaccinium microphyllum Blume____-~-----

Rhododendron apoanum Stein_----.----~--
MYRSINACEZE:

Rapanea avenis Mez____--------------------
SAPOTACEE:

Palaquium celebicum Burek ---------------
LOGANIACE:

Couthovia celebica Koord___------~- Sees

Fagraea plumeriaefolia A. DC_----.----~---

Strychnos celebica Koord__-.---------------
APOCYNACE:

Ichnocarpus ovatifolius A, DC__-.-~-------

Tabernaemontana cumingiana A, DC_____-

| Philip- |
| pines. |

Celebes.

Other distribution.

fe fee Ff

Amboina, New Guinea,

Samoa.

Java.

Moluceas.

New Guinea.

Boeroe, Ceram.

New Guinea.

| New Guinea (—).

Java.

Table showing floristic relationship between the Philippines and Celebes—Continued.
g. i PI

ie =< es tre SS

ee a
| Philip- | Celebes. | Other distribution.
|

Species. pines.

ASCLEPIADACE#:

Dischidiopsis philippinensis Schltr_________ i + ~ } New Guinea (—).
CONVOLVULACE: | |

Bonamia semidigyna var. farinosa Hallier__ + + | Banea, Lepar.
VERBENACEE:

Geunsia cumingiana Rolfe________________- ar | +

Premna cumingiana Schauer_______________ +4 +

Clerodendron blancoi Naves___--.-_---_____ + =
ACANTHACE®: |

Eranthemum zollingerianum Nees_________ =o of

Hemigraphis cumingiana Nees_____-__-____ + +

Hemigraphis rapifera Hallier_______________ 4 +

Hemigraphis primulifolia F.-Vill____-_-____ | oo + New Guinea.
BIGNONIACE®:

Nycticalos cuspidatum Miq-__--_-_---_.____- 4 | + Moluceas.
CUCURBITACE#: |

Momordica ovata Cogn 2 22-8 eee as | 4

Aes EA)

Notes on the above table.—Dioscorea sp. noy. is the species enumerated by
Koorders as D. glabra Roxb., but which is a distinct undescribed one according
to Dr. Prain in lit. and which has also been found in the Philippines. Gle-
ditschia rolfei Vid.; G. celebica Koord.,isasynonym. Pterocarpus echinatus Pers.
(P. vidalianus Rolfe) is also known from Salajar Island, south of Celebes.’
Wallaceodendron celebicum Woord., a monotypic genus, first found in Celebes and
later discovered in the central Philippines. Reinwardtiodendron merrillii Perk.,
a genus of two species, one Celebes and one Philippine. Koordersiodendron
pinnatum (Blanco) Merr., a monotypic genus known from the Philippines, Celebes,
and New Guinea. Cubilia blancoi Blume, a monotypic genus known from Luzon,
Celebes, Boeroe, and Ceram, Cubilia rumphii Blume being a synonym. <Anom-
panax philippinensis Harms., a genus comprising two species, one Celebes and
one Philippine. Vaccinium microphyllum Blume, see page 221.

Whereas the Celebes connections are very strikingly shown in the above
table, the semi-temperate or continental element in the highlands of
northern Luzon is nearly as prominent, as is shown by the following one.
The latter undoubtedly is far from complete and therefore, when more
extensive collections are made and our present material has more
thoroughly been worked over with reference to Asiatic types and com-
pared with them, the number of species will undoubtedly be greatly
enlarged.

The northern element in the Philippine flora—In the following table,
as in the preceding, the cross (+) indicates that the species is found in
Luzon and in the region for which the sign is entered, whereas the
dash (—) means that the Philippine species is represented by a closely
related form in the region indicated, or, in the last column, that the
general generic distribution is northern and not Malayan. The majority

7 Merrill: Publications of Bureau of Government Laboratories (1904), 17, 20.

174

of the genera so indicated are unknown from the Malayan region.
Specimens of all the species enumerated are in the herbarium of this
Bureau.

Table showing the northern element in the Philippine flora.

s ge) 5. ee
cS} retan | esrecall ees
| Species. 8 ® | 8a]
| . a = | = 5 zs
Pie ie Par ahi PeeaKcteiPaed be =
Wi ee =a (ee les eae eam eae peter fee
| aie |S |S|2)a}2/8 |e
|
POLYPODIACEE:
Cheilanthes argentea Hook -__------_- 5 eee + + a eS ol ie Ee
| Aspidium fauriei var, elatius Christ--| + |------ = jas ee ae — Wpeseeee — |------
Polypodium mengsteense Christ__--- Seta exes ee fae sfc) Neeson] So 2k os a ee en
Polystichum varium Pres] _-_-------- St ee + Sc A ees reser og ete be
Nephrodium erythrosorum Hook_---| + |------ + =f = fosececlest coe | See eee
Diplazium japonicum Thunb___----- oe ee + + SS ee ee eee
GLEICHENIACE®:
Gleichenia laevissima Christ ~------- 5 en ee ee ee ea eee eres ie BE eek
PINACE:
PAMUSMU SUIT Sie Ol = eens ween ee a a a a [eee ee ea Ne | Pe —
Pinus merkusii J. et DeVr ___---__-__ af NESS oe ies) sons s| hoes Seen see + _
GRVAIMIUN EH Alsen os Sa Se eee oe
Agrostis elmeri Merr_-.-------—--—--- 7 et — S25 SA — _
Calamagrostis filifolia Merr.--_-~--__- ae Nee ale ee | Se aS | ee ee _
Calamagrostis arundinacea var. nip-
POW CaM ACk=-e. wes ee = eee + — a eae Beene =
Poa luzoniensis Merr -_-___-.-._-_--- a | (Se ec el fe |-===-= as ea a —
Pollinia quadrinervis Hack ________- a ale | ere + + oh ose acl bakes ee
Polliniamudattrin’=—--—- = ae = Se Seana + 4 cope (EES eee | ae |
Pollinia imberbis var. willdenow- ;
IAN AAC Kae ene = eee eae ee SA Re eee |e eee ae = ee ee ee eee ee
Arthraxon microphyllum Hochst ~--| + |------]------ lestece Se ey ee ee ee ;
Eremochloa leersioides Hack ______ + |------|------ 4 |------|-----=[-nn=a-|---==-|=-mae
Brachypodium silvaticum Beauy. |
WATS TOW Keeton ss hee eee aL BA = ales — se lege —
Anthoxanthum luzoniense Merr ----| + |------|-=----|------ ee ee en meres _
Coelachne hackeli Merr -_---------_- 7 By) eae — — =e scan] a Sees eS _
CYPERACES: |

Carex, about:14 species 2252-23-22 2!) =p. |Sea se. os eee ea Se ee eee ee ee _
JUNCACER:

JUNCUS, 2 SpeCles =—s=- 32-65 e ee ee ee el ee foal ee ee ee -
LILIACEZE::

Aletris spicata Franch _--.--_-..------ + | 4- -+- | eer pees are) eee OS _

Disporum pullum Salisb —------------ | +- + -+ ate wile ees = em ee —

Ophiopogon japonicus Salish —.-___- vo + + oe heme 8 Sg S| Lie a —

Lilium philippinense Baker ____--__- ag eee) (t= eae | See eee | eerie Eee —
PIPERACE#: |

Saururus loureirii Deene______- eae, + nl Pt Oe 8 eee peered cee ------ _
SALICACEE: | | | |

Salix azaolana Blanco __----_-------- | + ad 2 _ Pes epee Bers 3 janes _-
MYRICACEA: |

Myrick THDIE Sy ie a peona cee adane + | 4 +} +4 Se See Pee dee 4

u '

175

‘Table showing the northern element in the Philippine flora—Continued.

‘ or 5 As
S 5 B
3 2 ee) se
. 2 |3¢| a
Species. ke o | 88 |32
2 | Ely) eral bey bre tie licks
: 2 : A 3) 3) A aise
Stbege aleseeiliees ve ciel Soult
Bef sc aeh Se Nar ea oB.ch a ollaag
4 By i) o A i) A S © © |
URTICACES: | |
Chamabainia squamigera Wedd ____| 4+ |----__|______ Bate [i =foopey te Selle Ra cae —
Debregeasia longifolia Wedd ____-___| Se | Sooo oe Sete le See ect al ca. Ok | POTN aL —
CARYOPHYLLACE: | |
Stellariavlaxa’ Merr2_-- - 2. st 225 243 Sn eee =f at EY | Mee oa || ey es ats _
RANUNCULACE#:
RaAMMNeUlUS ne. Spee s-— | 5 ee ree el ee le ee =
Anemone luzoniensis Rolfe_____----- abe |e cae | See: |e en Se ee ee eee =
BERBERIDACE®:
Berberis barandana Vidal -______-__- aa see |e eee jones | eeee tee eae ate SH =
Mahonia nepalensis DC__--_--------- eg Pees +4 Pe Va A 2 ee oe aes ==
CRUCIFERE:
Cardamine parviflora Linn —-__-______ an a a7 a Oe It ag + |=====- >=
CRASSULACEE: |
Sedum australis Merr_----_---------- Sat ese fer le oe J2eze5s| Se eo Gee. oh eee ae
SAXIFRAGACE:
INA esac iy oy (Oss TelanUern ts WaKol eel) Se Nee | S23 at ee he =
Deutzia pulchra Vid_-_--__---_-____- any ened Kee a jets | Ae ta se
Hydrangea lobbii Maxim_-_____-_____ Se ee eee ee (be as Bae A Me Le ed =
Itea macrophylla Wall —--_--_._--__- St pee ee=| eee | ae a eea sae ees eee ee =
Dichroa febrifuga Lour ___-____---__- Srey Sees ees + | aes eee + ar
ROSACE:
BragariaindicayAnd rss ss esse + + of + ep |e ne Si A aes [oe a ee fore
Rosa multiflora Thunb -_------_----- + | + |] + Ae eae |r eda Se Sg
Rubus tagallus Cham. & Seh —____--_- ae ie ae SFrte eat el es IL See =
Rubus rosaefolius Sm _____---__--___- Se See | eee = ae) es ee ee ee ee eo a
RUTACEE:
Boenninghausenia albiflora Reichb_| + os oo ar a? | Se ele eee aie SS
Skimmia japonica Thunb ___-------- = = + _— | Desk 3s SICA Ne Deere eee _
POLYGALACE:: |
Polygala glomerata Lour ___---_____- oa eee + fee | ost sal es | ee ==
CORIARIACE:
Coriaria intermedia Mats______------ + = — =) |Bibe st ess Ses eee _
ACERACE:
Acer philippinum Merr____-_-------- ap ee bee LS oe ae ee —_—
ACCmSp Inde tes 2s sae oo ee lee Se |------|------|------|------]------|------ =
VIOLACE#:
WiolaspainciniiDO 2s es a eee } + + fy (| Sak SH ee ee =
MELASTOMATACE:
SaTCOPVIAMUISH Spee eo en Se | ee ee == |ECSSESIBSs322 — =
OENOTHERACE: |
Epilobium, 2 species_________ ---____- yl re ee pS es S| Bes eS ae es ea ee Ss
HALORRHAGACE#:
Halorrhagis scabra, var. elongata
SCOT Ge wes oe sere NE A at RG os aes (eee “+ al |e Cl be She 8 eS OS
ARALIACE#: ,
Acanthopanax trifoliatum (L.) Merr_) 4 + + + a SE Se S| al eed be

176

Table showing the northern element in the Philippine fora—Continued.

|
|
Species.
; %
S S Fa] =
lac =| os 5
ERICACEE: i }
Rhododendron, 14 species ~---------- Stel bs oan) eee ee ESS
Vaccinium, 10 speties --------------- aan eer ene pa
Gaultheria, 3 species_----------:----- ee es eee je ae
MYRSINACE: |
Ambylanthopsis n. sp --------------- es eee Pee
PRIMULACELE:
Lysimachia microphylla Merr___---- oo end eee eee
Lysimachia ramosa Wall ~-------~--- SE | eae | tah | ee
Androsace saxifragifolia Bunge ____- sie as 2h ate
SYMPLOCACE-E:
Symplocos lancifolia 8. & Z ----_---- hal eosess | seee ee 4
OLEACE:
Fraxinus philippinensis Merr -----~- ie Pee ee et a
Ligustrum cumingianum Decne ----| + |_-----|------|------
GENTIANACE:
Gentiana, 3 species, 2=-===—-=----==_4- Sta fG| eget |e
Crawfurdia luzoniensis Vid__------_- See Oe oe ees ae
HYDROPHYLLACE#:
Ellisiophyllum reptans Maxim —_----| + |_____- By | eee ts
BORRAGINACELE:
Bothriospermum tenellum F.& M__-) + ae + Se
Trigonotis philippinensis Merr _____- rere eke) Ui | es eee ee
VERBENACE:
Callicarpa formosana Rolfe______---- =e au + —_
LABIATE:
J GPS C FWD OOH sta se RS A eg peers Peer! beset
Plectranthus diffusa Merr __________- a ee eel (ee | ae
Plectranthus'spss 253-2254 825 aay ate eee eee
Scutellaria luzonica Rolfe _-_-____- Ps ea 1h
SCROPHULARIACE#:
Alectra dentata O. Ktze ~-------____- Veet aac Be tao
Euphrasia borneensis Stapf —------__- mW | eee | oie Oa a
Sopubia trifida, Ham=-2--=-2.-- 2 ese. opis) ek Nie +-
RUBIACEE: |
Galium philippinensis Merr ___--___- Sf Pe ote ae ee
CAPRIFOLIACE®:
Lonicera rehderi Merr__------------- il sai aed el
Lonicera philippinensis Merr_______- se eal eee
Viburnum luzonicum Rolfe —-----_-- Sf Reena al ween
Viburnum odoratissimum Ker_ ~~, be Fae a +.
Viburnum sinuatum Merr__-----~--- oe Pe tree!
CAMPANULACE:
Lobelia nicotianaefolia Heyne —__-_- i eae) eee ae |
Pratia begoniaefolia Lind] __.-.-___- | -/ + H +-
Peracarpa Ny 6p) — beset eae | + |------|------|------

_ Northern India.

North America.

Europe.

or |

Sumatra,
Borneo.
Of general northern
generic distribution.

Java,

177

Table showing the northern element in the Philippine flora—Continued.

Ee eines
Species. : | z | = Be | ge
Se ie Sse AeA Tas Rea
A oa 5 oO Z Sb ly Fae WSs eye
| |
COMPOSIT #&: | | | |
INSters2 SpeCics seca se = ee ee often lee ste |Sesaee SN SINS) ape (os eae poe eo —
Artemesia parviflora Roxb ---------- So eee eee ese Pe ee ee eee cS
Ainslaea reflexa Merr _-------------- + | ile a ee | =|) 0a | See ee = |=
Carpesium cernuum Linn ----------- |} + peel Tl eee ted + + ee e2eeoe ates =
Cnicus wallichii Clarke_------------- | a eg ee ol oa Wirea, et eee cease | Oe ree _
Dichrocephala chrysanthemifolia | | | | |
PNG aes le AE eee et ee Lanse Fe Peed csc er mel. oes Rae -
Anaphalis cinnamomea Clarke------ Wet wees [eke ake || ab ees —|-=—=— =
Gnaphalium, 3 species___-_---------- Be aS ae | Joep ay] eee eee eeesaers leet [heres Vise a ad = |
Eupatorium lindleyanum DC-__------ | + + | + a ee ee pastes eos _
Lactuca thunbergiana Maxim____-_- jae =f fs is, Se) leeks see Widder Pea a! a
Lactuca brevirostris Champ -----_--- oa eee Si || | eee | ee ST ed ES —
Myriactis humilis Merr _____--------- [gh Nice el eet jetas aus fae eee lpr eee —
Seneciovs species seeea sea es pee ee |e es \ea ae Some Sree | eee Iisa eke —
Solidago virgaurea L __-------------- Nese (erew a We ee pes Sorry iret cae of slp | pa =
Vernonia philippinensis Rolfe______- + | = | as ae — | pee hou Rees | peer feetece|Pacees |
| | |

L — — ——————

Notes on the above table-—Chamabainia squamigera, a monotypic Asiatic genus.
Sedum australis is the most southern species of the genus known in the Eastern
Hemisphere. Polygala glomerata: This species is included, but the specimen
referred to is from Mindanao, not as yet having been found in Luzon. Several
other species of Polygala are known from the Philippines. Viola patricinii:
Three other species of Viola are represented in material recently collected in
northern Luzon, one of these has also been found in Mindanao. Sarcopyramis
is a monotypic Himalayan genus, the Luzon plant apparently representing an
undescribed form. Hpilobium: One or both forms may be undescribed. Rhodo-
dendron and Vaccinium include the total species known from the Philippines.
Ambylanthopsis, an Asiatic genus of two species from Bhotan and Assam, a
third undescribed species in Luzon. Ellisiophyllum reptans is a monotypic genus
previously known only from Japan. Peracarpa, an undescribed form, the genus
previously containing but one Himalayan species. Solidago virgaurea: This is
its most southern range in the Eastern Hemisphere.

PANDANACEE.
FREYCINETIA Gaudich.

Freycinetia rostrata Merrill sp. nov. § Pleiostigma.

A scandent, glabrous plant with terminal, globose heads, usually in
threes, and broadly-lanceolate, somewhat acuminate leaves 6 to 11 cm.
long, the margins with very few scattered small teeth. Branches brown-
ish gray, 5 mm. or less in diameter, obscurely angled. Leaves 1.5 to 2.4
cm. wide, narrowed below to the short flattened petiole, the margins often

175

recurved, nearly or quite entire below, above with few small scattered
teeth which are not spine-like. Heads about 1.5 em. in diameter,
globose, the peduncles from the apices of the branches 1 to 1.5 em. long.
Drupes many, but most of them aborted, narrowly ovoid, about 8 mm.
long, narrowed above into a rostrate apex about 2 mm. long. Stigmas 4.

MINDANAO, Lake Lanao, Camp Keithley (475 Mrs. Clemens) April, 1906.
PANDANUS Linn.

Pandanus clementis Merrill sp. nov. § Sussea. (PI. I.)

A tree about 10 m. high, branched above, with ovoid, 3 to 4.5 em. long
heads, arranged in terminal condensed spikes, about 8 heads in each spike,
each head subtended by a large broadly-ovate to oblong-ovate, acute or
very shortly abruptly acuminate bract, the drupes indefimite, 10 to
12 mm. long. Ultimate branches about 1.5 em. in diameter. Leaves
60 em. long or more, 2 to 2.5 em. wide, the margins with small antrorse
spines throughout, the midrib above glabrous, beneath at the base with
a few retrorse spines and near the apex finely antrorsely denticulate,
the apex short acuminate. Heads red when mature, ovoid, obtuse,
obscurely obtusely 3-angled, 3 to 4.5 cm. long, 2.5 to 3 cm. in diameter,
the bracts gradually smaller upwards, 4 to 6 em. long, 3 to 4 cm. wide,
acute or very shortly acuminate, serrulate on the keel and margins above.
Drupes very many in each head, 10 to 12 mm. long, 3 to 4 mm. thick
above, gradually narrowed below, somewhat angular, the apical portion
convex, slightly ribbed. Stigma terminal, central, 1 to 1.5 mm. in
diameter.

Mryxpanao, Lake Lanao, Camp Keithley (Mrs. Clemens) March, 1906. Altitude
about 800 m.

GRAMINE4.
ANTHOXANTHUM Linn.

Anthoxanthum luzoniense Merrill sp. nov.

Erect, glabrous, 40 to 60 em. high, very fragant in drying. Culms
elabrous; nodes slightly pubescent with short appressed hairs; sheaths
usually exceeding the internodes, often loose, the margins above slightly
pubescent; ligule broad, ovate, obtuse, irregularly cleft or lacerate, hya-
line, about 4 mm. long; blades 11 to 18 em. long, 4 to 5 mm. wide,
elabrous, smooth, flat or revolute when dry. Panicles exserted, rather
dense, many flowered, 6 to 8 cm. long, 1 to 1.5 em. in diameter, the
lower branches about 3 em. long, appressed-ascending, densely flowered.
First glume hyaline except the median portion, ovate, acuminate, 3 to
| mm. long, 1-nerved ; second glume 5 to 6 mm. long, 3-nerved, glabrous,
oblong ovate, abruptly subtruncate-acuminate. Flowering glumes 3, the
first and second neuter, slightly pubescent, the first about 5 mm. long,
3-nerved, cleft at the apex for about 4 its length, awned in the cleft, the
awn scabrous, straight, about 1.2 mm. long; second flowering glume

MERRILL: NEw OR NOTEWORTHY) j PHIL. JOURN. Scr., Vou. I, Supp. III,
PHILIPPINE PLANTS, V. i \ Aveusr 15, 1906.

.
=. cone
;

Pirate |. PANDANUS CLEMENTIS MERR.

RS ts
wr r Me

al

179

about 4 mm. long, oblong, 1-nerved, cleft at the apex, awned from the
back at the lower 4, the awn straight, scabrous, about 6 mm. long; third
flowering glume broadly-ovate, ‘obtuse, hyaline, glabrous, nerveless or
nearly so, hermaphrodite, inclosing a very narrow palea. Anthers nar-
rowly oblong, about 2 mm. long.

Luzon, Province of Benguet, Pauai (4713 Merrill) November 8, 1905.

In the same locality and with the same associates as Poa luzoniensis, and
like that species a very interesting northern type. Hackel, who has kindly
examined specimens, states. that this species is apparently related to the British
Indian plant described by Hooker as Hierochloa clarkei, judging from the deserip-
tion. The present species differs from the well-known and widely distributed
Anthoxanthum odoratum Linn., in its glabrous outer glumes and especially in
the very short-awned first-flowering glume. It has the same characteristic odor
and much the appearance of that species.

ARUNDINELLA Raddi.

Arundinella setosa Trin. Diss. 2 (1824) 63; Hook. Tey Tl, Tehhes Whale 7 MTU)
70. Danthonia luzoniensis Steud. Syn. Pl. Gram. (1855) 245; Miq. Pl. Ind.
Bat. 3 (1859) 427; Vidal, Phan. Cuming. Philip. (1885) 159; Rev. Pl. Vase.
Filip. (1886) 282; Ceron Cat. Pl. Herb. (1892) 184; F.-Vill. Nov. App.
(1883) 319.

Puirerines (1415 Cuming) cotype of Danthonia luzoniensis Steud. Luzon,
Province of Benguet, Pauai to Baguio (4702 Merrill) November, 1905; Bued
River (4271 Merrill) November, 1905.

British India to Cochin China, and southern China.

This species is enumerated here only to call attention to the reduction of
Danthonia lugoniensis Steud. On receipt of a specimen of No. 1415 Cuming,
from the British Museum, I found the number to be an Arundinella and not a
Danthonia. A fragment was sent to Dr. Hackel, who pronounced it to be identical
with Arundinella setosa Trin. The genus Danthonia can therefore be excluded
from the Philippines.

CALAMAGROSTIS Adans.

Calamagrostis filifolia Merrill sp. nov. § Deyeucxia.

A densely tufted, erect grass 30 to 60 cm. high, with filiform leaves,
and strict, densely flowered panicles. Culms glabrous, many in each
tuft, erect, straight, glabrous; nodes dark, glabrous; sheaths, at least the
lower ones, shorter than the internodes, the upper one often inclosing the
immature panicles; ligule about 2 mm. long, subhyaline, irregularly cleft
or lacerate ; blades 10 to 20 cm. long, numerous at the base of the culms
and on the innovations, erect, somewhat stiff, revolute, 1 mm. wide or less.
Panicles at length exserted, pale green’ or slightly purplish, many
flowered, the lower branches appressed, often 4 cm. long, forming a
narrow somewhat spikelike, uninterrupted panicle. Empty glumes sub-
equal, lanceolate, acuminate, about 6 mm. long, both 1-nerved, scabrous
on the keel. Flowering glume lanceolate, 5 mm. long, minutely scabrous,
4-nerved, the nerves green, entire or very obscurely 2 to 3 toothed at
the apex. Palea equaling the glume. Anthers 1.2 to 1.5 mm. long.
Lodicules 1 mm. long. Callus hairs 1 mm. long or less; rhachilla about

-

180

1.2 mm. long, the hairs 2 mm. long or less. Awn of the flowering glume
inserted at about the lower fourth, 7 mm. long, scabrid, somewhat gen-
iculate and twisted below the geniculation.

Luzon, Province of Benguet, Mount Tonglon (4839 Merrill) November 12,
1905. In dry open soils at the summit of the mountain at about 2,250 m.
Hackel states that this species is closely related only to Calamagrostis nardifolia
(Griseb.) Hack. (Agrostis nardifolia Griseb.) of the Argentine Andes. A New
Zealand species Calamagrostis (Deyeuxia) setifolia has similar leaves but quite
other inflorescence and spikelets.

Hackel has suggested the name Calamagrostis filifolia forma cleistogama for
specimens No. 4537 and 4715 Merrill, the former from Mount Data, Lepanto,
and the latter from Pauai, Benguet, the specimens differing from typical C. filifolia
in having the 0.5 mm. long anthers persisting with the ripening fruit within
the closed glumes and palea and lodicules 0.7 mm. long.

If we exclude Deyeuxia quadriseta Benth., credited to the Philippines by
F.-Villar, and undoubtedly not a Philippine plant, the identification apparently
having been erroneous, then the above species is the first one of the genus to be
reported from the Philippines, the genus in general being a northern one, or at
least characteristic of the more temperate regions.

POA Linn.
Poa luzoniensis Merrill sp. nov.

An erect perennial grass from long rootstocks, not at all tufted.
Culms 30 to 40 em. high, glabrous, smooth, geniculate below; nodes
glabrous; sheaths equaling or shorter than the internodes, rather loose.
the lower ones dry and brown; ligule short, truncate, cartilaginous ;
blades 8 to 14 em. long, 1.5 to 2.5 mm. wide, flat, thin, glabrous, some-
what acuminate. Panicles lax, comparatively few flowered, slightly ex-
serted, about 9 em. long, the branches few, whorled, ascending or
spreading-ascending, the longer ones 3 cm. or less, filiform, each bearing
few spikelets. Spikelets about 3.5 mm. long, green or purplish, the
pedicels 2 mm. long or less; first glume lanceolate, acuminate, about
2.1 mm. long, 1-nerved, slightly scabrous on the keel; second glume
similar to the first but nearly 3 mm. long and 3-nerved. Flowering
elumes 3, ovate, acute, 5-nerved, about 3 mm. long, slightly silky hairy
on the keel below, otherwise glabrous, the paleas narrow, hyaline, about
2.5 mm. long, the rachilla glabrous.

LuzoN, Province of Benguet, Pauai (4712 Merrill) November 8, 1905.

This species was found in a large, open, natural meadow along the borders of
a cold stream at an altitude of about 2,100 m., being very rare in the locality,
and associated with Calamagrostis, Brachypodium, Bromus, Anthoxvanthum luzo-
niense, Agrostis elmeri, Ranunculus sp., Viola sp., various species of Carex and
other northern types. Single, scattered specimens only were found, the rhizomes
creeping rather extensively and generally rooting at the lower nodes. This
decidedly northern type is an interesting addition to the known boreal element
in the flora of northern Luzon, being the first native species of the genus to
be reported from the Philippines. Dr. Hackel, to whom specimens were sent,
states that it may be most closely related to the widely distributed Poa pratensis

Linn.

181

Poa annua Linn. Sp. Pl. (1753) 68; F.-Vill. Nov. App. (1883) 322.

Luzon, Province of Benguet, Bued River (4288 Merrill) November 14, 1905,
altitude 1,200 m. :

This species was reported from the Philippines by F.-Villar, who states that
he saw living specimens in Luzon and Panay. I have never seen specimens of
this grass from regions at low elevations in the Philippines, and it is possible
that F.-Villar’s record was based on an erroneous identification. Poa annua was
found along the new Benguet road near some of the construction camps, the
seeds undoubtedly having been introduced in American hay, used for food for
the horses and mules utilized in the construction work. It was well established,
and will undoubtedly persist.

Among other weedy plants collected on the same trip which were certainly
introduced by the same means are Trifolium pratense Linn., 7. repens Linn.,
Cerastium vulgatum Linn., Spergula arvensis Linn., Rumea acetosella Linn., and
Rumex crispus Linn. All of these plants were found in cultivated places or in
waste lands near construction camps, and some of them will undoubtedly persist.
With the exception of Rumex acetosella Linn., none of the above species have
previously been reported from the Philippines. Trifolium pratense Linn., has also
been collected in Mindanao, Camp Keithley (Irs. Clemens), April, 1906.

FLAGELLARIACE.
JOINVILLEA Gaudich.

Joinvillea malayana Ridley, Journ. Straights Branch Roy. As, Soc. 44 (1905)
199.

PALAWAN, Balsajan River (575 Foxworthy) March 3, 1906. In clearings in
forests at an altitude of about 650 m. Malayan Peninsula and Borneo.

No species of this small genus has previously been discovered in the Philip-
pines, the above specimen agreeing perfectly with authentic material of Joinvillea
malayana in the Herbarium of this Bureau, received from the Botanic Garden,
Singapere, collected at Perak. The species was described from material collected
in Perak and Selangor, Malayan Peninsula, and Mount Matang, Sarawak, British
North Borneo.

As noted by Ridley, this form is most closely related to one of the Hawaiian
species, there being two of the genus known from Hawaii, and one from New
Caledonia. For parallel distribution Ridley cites only the genus Cyrtandra of
the Gesneriacee, which reaches its maximum development in the Sandwich
Islands, extending to the Malayan Archipelago and Peninsula. For the Philip-
pines a parallel case of distribution is represented by Tetraplasandra philip-
pinensis Mer. of the Araliaceew (see p. 219). In this very characteristic genus,
one species is known from the Island of Palawan in the Philippines, one from New
Guinea, one from Celebes, and six from the Hawaiian Islands. Another case is
represented by the genus Nchistostege Hillebr., of the Polypodiacee, of its three
known species, one being from Hawaii and two from the Island of Mindanao,
southern Philippines.s Rolfe® cites Aspleniwm persicifolium J. Sm., as being
confined to the Philippines and Hawaii, but this species is apparently endemic
in the Philippines, as Hillebrand” considers the Hawaiian form distinct from
A. persicifolium.

’Copeland: Philip. Journ. Sct. 1 (1906), Suppl. 155.
” Rolfe: Journ. Linn. Soc. Bot. 21 (1884), 397.
1 Hillebrand: Fl. Hawaiian Isl. (1888), 591.

182

LILIACE AS.
ALETRIS Linn.

Aletris spicata (Thunb.) Franch. Journ. de Bot. 10 (1896) 199. Hypowis
spicata Thunb. Fl. Jap. (1825) 136. Aletris japonica (non Thunb.) Lamb.
Trans. Linn. Soe. 10 (1811) 407; C. H. Wright in Forbes & Hemsl. Journ.
Linn. Soc. Bot. 36 (1903) 76.

Luzon, Province of Benguet, Bugias (4685 Jerrill) October 28, 1905; Pauai
to Baguio (4801 Merrill) November 9, 1905.

Widely distributed in the highlands of Benguet Province, but not abundant,
growing on dry grassy slopes in thin pine forests 1,200 to 2,000 m. An interest-
ing addition to our knowledge of the northern element in the Philippine flora.
Japan and the Corean Archipelago to Central and Southern China and Formosa.

DISPORUM Salisb.

Disporum pullum Salisb. Trans. Hort. Soc. 1 (1812) 331; Wright in Forbes
& Hemsl. Journ. Linn. Soe. Bot. 36 (1904) 142.

Luzon, District of Lepanto, Mount Data (4857 Merrill) November, 1904. No
species of the genus has previously been reported from the Archipelago. It is
an interesting addition to the northern element in the Philippine flora. Not
abundant in the mossy forest at an altitude of about 2,200 m., the specimens in
fruit only. Japan to Formosa, China and temperate Himalaya, Java and Sumatra.

SALICACE.
SALIX Linn.

Salix azaolana Blanco, Fl. Filip. ed. 2 (1845) 539; ed. 3, 3 (1879) 188; Merr.
Govt. Lab. Publ. 27 (1905) 81. Salia tetrasperma Llanos (non Roxb.?)
Mem. Ac. Ciene. Mad. (1858); FI. Filip. ed. 3, 4 (1880) 106; F.-Vill. Nov.
App. (1883) 210.

Luzon, Province of Rizal, Bosoboso (1987, 2139 Ahern’s collector) November,

December, 1904, the former with staminate flowers, the latter with mature fruits.

Blanco’s description of this species is very short and imperfect, and a transla-
tion follows:

Salix (err. typ. Salis) azaolana, Azaola’s willow. Leaves alternate, incised,
serrate, attenuate at the apex, glabrous. Flowers dioecious. *“Staminate
flowers in aments. Calyx bifid. Stamens more than 12 (Azaola). Anthers
globose. *Pistillate flowers. Calyx as in the male. Achene one.==A_ tree,
the trunk as thick as a man’s body and which is found along the margins of
rivers in Calumpit. (Province of Bulacan, Luzon.) Given to knowledge by
P. Llanos. *Tiaun. :

Llanos, followed by F.-Villar, reduced this species to Saliva tetrasperma Roxh,,
the latter adding that he had seen specimens sent to him by Llanos and also
that he had seen living specimens at Quingua, Province of Bulacan, Luzon.

There is little doubt but that the specimens cited above represent Blanco’s
species, although his description is very imperfect; so far the only one of the
genus known from the Philippines and one not previously collected except by
Llanos and F.-Villar, no material of these early collections being preserved.
Although Blanco cites Calumpit as the locality from which his material came,
it seems probable that it was only sent to him from that town, having been
collected at some other locality. During a recent trip up the Pampanga River,
a stop of several days was made at Calumpit, and a careful search failed to

183

reveal the plant and I could find no natives who knew the word ‘Tiaun”? as a
plant name. However, as the country for many miles about Calumpit is under
rather close cultivation, it is apparent that the original vegetation of the region
has almost entirely been removed.

MORACE/®.
FICUS Linn.

Ficus anomala Merrill sp. nov. § Palacomorphe,

A tree about 10 m. high with elliptical-ovate, abruptly short acuminate
scabrous leaves, the receptacles globose to obovoid, yellow, scabrous, in
large fascicles on the trunk and larger branches. Branches brownish
gray, glabrous, the branchlets also glabrous except at the slightly ciliate
tips. Leaves submembranous, 10 to 16 cm. long, 6 to 11 em. wide, slightly
shining, harsh on both surfaces but not pubescent, the margins subentire
or very obscurely undulate-crenate, the base broad, rounded, truncate
or somewhat cordate, often somewhat inequilateral, the basal nerves 5
to 7, the outer ones short; primary lateral nerves about 7 on each side
of the midrib, prominent, the reticulations distinct, lax, subparallel ;
petioles 3 to 7 cm. long. Receptacles very numerous, often nearly con-
cealing the trunk and larger branches, in usually large fascicles of short,
stout, congested 1 to 2 cm. long branchlets, yellow when mature, scabrous,
nearly 2 cm. in diameter when fresh, considerably smaller when dry, the
pedicels ebracteolate, 1 to 2 cm. long. Male flowers numerous near the
ostiole, monandrous and with a rudimentary ovary, the perianth segments
4, free, enclosing and exceeding the stamen, oblong, about 2 mm. long,
slightly ciliate above. Filament stout, about 1 mm. long; anther
elliptical-ovoid, 1 mm. long. Rudimentary ovary less than 1 mm. long.
Gall flowers numerous, the perianth lobes 4, free, near, blunt, ciliate
above, much exceeding the ovary which is elliptical ovoid, 1.5 mm. long,
the style less than 1 mm. long, lateral. Fertile female flowers not seen.

Luzon, Province of Rizal, Montalban (5075 Merrill), March 10, 1906. <A tree
about 10 m. high and 30 em. in diameter growing along small streams in thickets
at an altitude of about 40 m. Anomalous for the section Palaecomorphe in that
the receptacles are not axillary but borne in large fascicles on the trunk and
branches as in many species of the sections Covellia and Neomorphe.

PARATROPHIS Blume.

Paratrophis caudata Merrill sp. nov.

A small tree about 8 m. tall, the young branches puberulent or gla-
brous, green, slender. Leaves broadly oblong-lanceolate or oblong-oblan-
ceolate, submembranous, glabrous, shining, the base obtuse, the apex
abruptly caudate-acuminate, the acumen obtuse, above with scattered
minute white glands, the margins irregularly crenate-serrate, 7 to 17
em. long, 2 to 5.5 em. wide; nerves 7 to 8 on both sides of the midrib,
prominent beneath, anastomosing; petioles 1 to 2.5 mm. long. Stam-
inate inflorescence axillary, solitary, spicate, amentiform, 4 to 6 cm. long,

184

white, densely many flowered, the rhachis puberulent with minute hairs.
Perianth 4-parted, the segments oblong-ovate, obtuse, about 1.5 mm. long.
Stamens 4; filaments inflexed in bud, exserted in anthesis, 2 to 3 mm.
long; anthers about 1 mm. long. Pistillate inflorescence axillary, spicate,
short, 1 to 1.5 em. long, few (about 8) flowered, the rhachis puberulent.
Perianth 4-parted, the segments imbricate, subequal, oblong-ovate, obtuse,
1.7 to 2 mm. long. Ovary sessile, oblong, exserted, 1-celled, glabrous ;
style 2 mm. long, bipartite for two-thirds its length.

MINDANAO, District of Davao (278 DeVore & Hoover), April, 1903. BASILAN
(2446 Hutchinson) December, 1905. ‘Common in forests above 15 m.” Hutch-
inson. Bagobo Hulas, in Basilan Olis.

A species characterized by its comparatively short inflorescence and caudate-
acuminate leaves, the margins of which are entire below and more or Jess
irregularly crenate-serrate above.

URTICACE.
CHAMABAINIA Wight.

Chamabainia squamigera (\Vall.) Wedd. in DC. Prodr. 16 (1869) 218. C. cus-
pidata Wight; Hook. f. Fl. Brit. Ind. 5 (1888) 580; C. H. Wright in
Forbes & Hemsl. Journ. Linn. Soc. Bot. 26 (1899) 489.

Luzon, District of Lepanto, Mount Data (4556 Merrill) November 4, 1905. In
the mossy forest at about 2,250 m. Western China to Northern India south to
Ceylon.

An interesting case of geographical distribution, no species of the genus having
previously been reported from the Philippines. So far as I am able to determine
from descriptions alone, the Philippine form is not distinct from the Asiatie
species to which it is here referred.

LORANTHACE.
LORANTHUS Linn.

Loranthus ahernianus Merrill sp. nov. § Dendropthoé.

Shrubby, branched, glabrous except the somewhat furfuraceous inflor-
escence. Branches glabrous, rugose, lenticellate, reddish brown when
dry, rather slender. Leaves opposite, glabrous, coriaceous, pale when
dry, dull, ovate to oblong ovate, the base rather broad, rounded or obtuse,
the apex acuminate, 6 to 10 cm. long, 2.5 to 5 em. wide, the midrib prom-
inent, the lateral nerves very irregular, obscure, petioles 1 em. long.
Inflorescence of solitary, axillary, lateral and terminal racemose panicles
6 to 8 em. long, rather many flowered, the axis and short branches fur-
furaceous, the 1.5 to 2 em. long peduncles subtended by a whorl of small
imbricated bracts, the branches mostly above the middle, about 5 mm.
long, each usually three flowered, the middle flower sessile, the two lateral
ones pediceled. Flowers slender, tubular, about 3.5 cm. long, yellow,
slightly enlarged below and at the apex, the subtending bracts ovate, that
of the sessile flower acuminate, 4 mm. long, those of the lateral pediceled
flowers smaller and often acute, the pedicels of the lateral flowers 3 to 4

185

mm. long. Calyx cylindrical, 4 mm. long, glabrous, the limb extended
somewhat beyond the ovary, truncate or obscurely 6-toothed. Corolla
about 3 cm. long, glabrous, the lobes linear, united for the lower 0.5 cm.,
the filaments inserted at about the middle, 9 mm. long, inappendiculate ;
anthers linear oblong about 5 mm. long.

Luzon, Province of Rizal, Bosoboso (2140 Ahern’s collector) December 15,
1904. A species apparently related to Loranthus viridis Mery., but differing in
many characters.

Loranthus cauliflorus Merrill sp. nov. § Stemmatophyllum, Arthrostemma.

Shrubby, glabrous throughout, the flowers arranged along one side of
the branches below the leaves on very short usually trianthous peduncles.
Branches dark brown, terete, glabrous, lenticellate. Leaves opposite,
coriaceous, 18 to 25 cm. long, 6 to 8 cm. wide, oblong-ovate to ovate-
lanceolate, dull, minutely densely wrinkled when dry, the base acute,
the apex acuminate ; nerves about 5 on each side of the midrib, ascending ;
petioles stout, 2 to 2.5 cm. long. Flowers rather numerous, red, the
peduncles stout, 2 mm. long or less, each bearing three, sometimes but
two or one, sessile flowers, each flower subtended by a broadly ovate
basal bract about 2.5 mm. long. Calyx cylindrical, 3 mm. long, the
limb short, truncate. Petals 5, free, 2 to 2.4 em. long, linear, 2 mm. wide
below, free, narrowed above, the recurved portion above the insertion of
the stamen about 8 mm. long. Anthers about 7 mm. long, sessile or
subsessile.

MinDANAO, Lake Lanao, Camp Keithley (55 Mrs. Clemens) January, 1906. <A
very characteristic species, at once recognizable by its peculiar inflorescence, the
flowers being borne along one side of the branches below the leaves.

Loranthus clementis Merrill sp. nov. § Dendropthoé.

Shrubby, glabrous throughout except the slightly puberulent flowers
and inflorescence. Branches brownish gray, stout, terete, with numerous
small lenticels. Leaves coriaceous, ovate to oblong-ovate, alternate, the
base acute, the apex obtuse, dull when dry, 8 to 12 cm. long, 4 to 6.5 cm.
wide, with two pairs of stout nerves from near the base of the leaf, the
upper pair extending nearly to the apex, the reticulations lax. Flowers
slender, somewhat curved, 2.5 to 3 em. long, 2 to 5 racemosely disposed
on each short peduncle, the peduncles numerous, fascicled in the axils
of the leaves. Calyx cylindrical, nearly 3 mm. long, somewhat fer-
ruginous puberulent, the limb produced above the ovary nearly 1 mm.,
5-toothed, the basal bract narrowly ovate, acute, about 1.5 mm. long, the
pedicel 1 to 2 mm. long. Corolla slender, somewhat curved, slightly
inflated above, the tube long, red and yellow, grayish or brownish scurfy
puberulent outside, the lobes 5, 10 to 12 mm. long, the reflexed portion
above the insertion of the stamens linear, about 8 mm. long. Filaments
4 to 5 mm. long; anthers about 2 mm. long.

Minpanao, Lake Lanao, Camp Keithley (Mrs. Clemens) March, 1906, altitude
about 800 m.

44772 2

186

Loranthus copelandi Merrill sp. nov. Dendropthoé § Budendropthoe.

Glabrous except the inflorescence, with 11 to 18 em. long lanceolate to
oblong-lanceolate petioled leaves, and many flowered axillary fascicled
racemes, the flowers red and yellow, about 4.5 cm. long. Branches
brownish gray, glabrous, terete. Leaves subcoriaceous, glabrous, minutely
wrinkled-rugose when dry, dull, 11 to 18 em. long, 2.5 to 3.5 em. wide,
narrowed above to the blunt apex and below to the somewhat attenuate
base, the midrib very stout, the lateral nerves 3 to 4 on each side of the
midrib, long, ascending, the reticulations obscure; petioles stout, about 1
em. long. Inflorescence shghtly ferruginous pubescent, 3 to 5 racemes
fascicled in each axil or in the axils of fallen leaves, 4 to 7 cm. long.
Flowers slender, the buds slightly pubescent, becoming glabrous except
the calyx, red below, yellow above, about 20 in each raceme. Pedicels
about 1 mm. long, puberulent, the bract narrowly ovate, acute, 1.5 mm.
long, puberulent. Calyx cylindrical, rather densely ferruginous puberu-
lent, 3 mm. long, with 5 small teeth. Corolla slender, somewhat curved,
slightly enlarged above, 4 to 4.5 em. long, 5-lobed, the lobes linear, about
1 em. long, becoming recurved. Anthers not narrowed below, 3 mm.
long, apiculate-acuminate ; filaments about 5 mm. long.

Luzon, Province of Benguet, Daklan to Kabayan (4407 Merrill) October 27,
1905. Parasitic on Pittosporum pentandrum (Blanco) Merr., in thickets at about
1,500 m. A species apparently closely related to Loranthus curvatus Blume,
differing in its vegetative characters and fascicled racemes.

Loranthus mindanaensis Merrill sp. nov. § Dendropthoé.

Shrubby, glabrous throughout. Branches terete, glabrous, light gray,
the younger branchlets brownish to olivaceous, rather slender. Leaves
coriaceous, glabrous, dull, ovate to oblong-ovate, 8 to 12 cm. long, 4 to
6 em. wide, the base broad, usually rounded, the apex blunt-acute, the
midrib stout; lateral nerves irregular, 3 to 4 on each side of the midrib,
ascending, not prominent, the reticulations lax; petioles stout, about
3mm. long. Inflorescence axillary, of solitary 3 to 5 cm. long racemose
panicles, the flowers in groups of threes at the ends of the very short
branchlets, the branchlets racemosely disposed, opposite, 2 to 3 mm. long.
Flowers sessile, glabrous, the bracts broadly ovate, about 1.5 mm. long.
obtuse. Calyx cylindrical, about 3.5 mm. long, the limb truncate, entire,
slightly exceeding the ovary. Corolla red, tubular, not enlarged below,
nearly 2 em. long, the lobes 6, linear, united for the lower 2 mm. Fila-
ments very short, less than 1 mm. long, inappendiculate, the anthers
linear, about 3 mm. long, the portion of the petals above the insertion of
the stamen about 5 mm. long, reflexed in anthesis.

MINDANAO, District of Davao, Davao (341 Copeland) March, 1904, on Theo-
broma cacao J, and Antidesma ghesambilla Gaertn.; Mount Apo (285 DeVore
& Hoover) May, 1903. A species related to Loranthus secundiflorus Merr., and

L. subalternifolius Merr., differing in vegetative characters from both.

187

Loranthus mirabilis Van Huerek et Muell. Arg. in Act. Soc. Helv. Sci. Nat.
55 (1872) 47. Stemmatophyllum cumingii Van Tiegh. Bull. Soe. Bot. Fr.
41 (1894) 505. Loranthus cumingianus Engl. Nat. Pflanzenfam. Nachtriige
(1897) 128, non L. cumingii Engl. 1. e.

PHILIPPINES, without locality (1966 Cuming) 1836-1840, in Herb. Bureau of
Science. BAsmLAn (3447 Hutchinson) December, 1905, parasitic on trees along
the seashore. MINDANAO, Davao (376 Copeland) March, 1904; Lake Lanao,
Camp Keithley (Mrs. Clemens) March, 1906.

The oldest name for this species is Loranthus mirabilis Van Huerck et Muell.
Arg., the description being published in 1872 as above cited. The name is
recorded in the first supplement to Index Kewensis, but the citation to the
original publication is not given, the reference being to Van Tieghem, Bull. Soe.
Bot. Fr. 41 (1894) 547, in obs. Van Tieghem |. c. states that he found this
number of Cuming’s Philippine collection in the herbaria of Van Huerck, De-
Candolle, and Boissier under the name L. mirabilis. He did not, however, find a
reference to the place of publication. Both Loranthus mirabilis and Stemmato-
phyllum cumingu (==Loranthus cumingianus) were based on the same number of
Cuming’s Philippine collection cited above. It was by chance only that I found
a reference to the original publication of Loranthus mirabilis, in reading over
the list of Mueller’s publications given by Briquet,’’ and I am indebted to Dr.
William ‘release, director of the Missouri Botanical Garden, for a copy of the
rather long description and discussion of the species, the original publication not
being available in Manila.

Loranthus secundiflorus Merrill sp. nov. § Dendropthoé.

Shrubby, less than 1 m. high, glabrous throughout except the minutely
puberulent inflorescence. Branches slender, terete, smooth, somewhat
enlarged and compressed below the nodes, olive brown when dry. Leaves
opposite, lanceolate, coriaceous, shining, sessile, 8 to 10 em. long, 2 to 3
cm. wide, paler beneath, the base obtuse or subacute, gradually narrowed
above to the slightly acuminate apex; lateral nerves obscure, irregular,
5 to 7 on each side of the midrib, the reticulations lax. Inflorescence
solitary, axillary, of 4 to 8 em. long racemose panicles, the flowers secund,
the axis and branchlets minutely puberulent, the branches of the in-
florescence opposite, very short, about 3 mm. long, racemosely disposed,
each with 2 or 3 sessile flowers at the apex, the bracts orbicular-ovate,
obtuse, about 2 mm. long. Calyx cylindrical, 4.5 mm. long, minutely
puberulent or subglabrous, the limb extending beyond the ovary and
rather prominently 6-toothed. Corolla glabrous, cylindrical, not en-
larged, about 2 cm. long, red, the lobes 2 mm. wide below, united for
the lower 2 min., the portion above the insertion of the stamens reflexed
in anthesis, 5 to 6 mm. long. Anthers sessile, linear, inappendiculate,
about 4.5 mm. long.

MINDANAO, Province of Surigao (237 Bolster) February 10, 1906. Parasitic
on various trees in open lands at an altitude of about 15 m. above the sea. <A
species recognizable by its secund flowers and sessile anthers.

1 Bull. Herb. Boiss. (1896), 4, 128.

188

Loranthus sessiliflorus Merrill sp. nov. § Phoenicanthemum.

Shrubby, entirely glabrous, with axillary solitary or fascicled spikes of
small sessile flowers, the whole inflorescence glabrous, red. Branches dark
reddish brown or nearly black when dry, terete. Leaves opposite, very
coriaceous, broadly-ovate to suborbicular-ovate, dull and minutely wrin-
kled on both surfaces when dry, the apex broad, rounded or obtuse,
rarely somewhat narrowed and acute, the base acute to subtruncate, 8 to
11 em. long, 5 to 8 em. wide, the nerves 4 to 5 on each side of the stout
midrib, obscure; petioles stout, about 1 cm. long. Spikes 1 to 4 or 5 in
each axil, usually 3, 4 to 7 em. long, many flowered. Flowers red,
sessile, about 7 mm. long, the subtending bract very thick, orbicular,
nearly 1.5 mm. in diameter. Calyx glabrous, cylindrical, about 2 mm.
long, the limb very slightly produced beyond the ovary, truncate. Petals
4, free to the base, linear, scarcely widened below, 5 mm. long, the upper
half curved-reflexed from the insertion of the stamens. Filaments about
1 mm. long, slightly rugose-wrinkled near the insertion below, the
anthers continuous, slightly wider than the filaments, oblong, 1.5 mim.
long. Style glabrous, about 5 mm. long. Immature fruit small, ovoid.

Luzon, Province of Bataan, Mount Mariveles (1171 Whitford) March, 1905;
Province of Benguet, Baguio (6057 Elmer) March, 1904. Mrinporo, Baco River
(4041 Merrill) March, 1905; (125 McGregor) March, 1905. Parasitic on various
trees, sea level to an altitude of about 1,500 m.

,

Loranthus subalternifolius Merrill nom. noy. Amylotheca cumingii Van Tiegh.
Bull. Bot. Soe. France, 41 (1894) 264. Loranthus cumingii Engl. Nat.
Pflanzenfam. Nachtr. (1897) 128, non L. cwmingianus Eng). 1. ¢.

Glabrous throughout. Branches light gray, terete, the branchlets
often olivaceous, somewhat thickened and compressed below the nodes,
smooth. Leaves coriaceous, subalternate, or the upper ones opposite,
oblong-ovate, usually pale when dry, somewhat shining 8 to 13 cm. long,
4 to 5.5 em. wide, usually gradually narrowed above to the acute, rarely
slightly acuminate apex, and more abruptly narrowed below to the
acute, often decurrent base; nerves about 6 on each side of the midrib,
obscure; petioles 1 to 1.5 cm. long. Inflorescence of solitary, axillary.
racemose panicles 4 to 7 em. long, the branches short, opposite, spreading,
racemosely disposed, about 3 mm. long, each bearing three sessile flowers
at the apex, the bracts reniform, rounded, about 2 mm. long. Calyx
cylindrical, about 5.5 mm. long, the limb strongly produced beyond the
ovary, truncate, somewhat spreading. Corolla tubular, about 18 mm.
long, golden yellow, somewhat inflated below, the lobes 6, linear, united
for the lower 3 mm. Filaments very stout, about 2 mm. long; anthers
linear, 2.5 mm. long, the portion of the petals above the insertion of the
stamens thickened, nearly 6 mm. long, spreading or reflexed in anthesis,

PHILIPPINES (1969 Cuming) 1836-1840. Luzon, Province of Benguet, Sablan
(6191 Blmer) April, 1904.

The above deseription is based on No. 6191 Hlmer, my specimen of Cuming’s
plant a cotype of Amylotheca cumingti Van Tiegh.=Loranthus cumingii Engl.,

189

being fragmentary, but sufficient I believe to show the identity of the two speci-
mens. Van Tieghem’s description being very short and imperfect, it has been
thought well to publish a longer one here.

Loranthus viridis Merrill sp. nov. § Dendropthoe.

Coarse, more or less branched, shrubby, glabrous throughout. Leaves
opposite, coriaceous, usually pale or yellowish when dry, elliptical-ovate,
ovate or even obovate, 6 to 11 cm. long, 4 to 6 em. wide, rounded or acute
at the apex, narrowed below to the acute, slightly decurrent base, shin-
ing above, dull beneath, the midrib stout, the lateral nerves obscure;
petioles stout, 2 to 3 cm. long. Cymes 5 cm. long or less, few flowered,
glabrous, solitary in the upper axils and terminal, the branches few, three
or four, whorled at the summit of the 1 to 2.5 em. long peduncles the
branches spreading, about 1 cm. long, each branchlet with two or three
flowers the pedicels about 2 mm. long, the bract subtending the flower
small, orbicular-ovate, rounded. Flowers pale green, about 3.5 em. long,
the buds somewhat clavate at the tips. Calyx cylindrical, glabrous, 5
to 6 mm. long, the limb truncate, or very obscurely 6 toothed, slightly
exceeding the ovary. Corolla slightly swollen below, the lobes 6, united
for the lower 0.5 cm., 3 mm. wide below, gradually narrowed upward
and 1 mm. wide above, the upper 1 em. reflexed from the insertion of the
stamens, much thickened and deeply channeled on the inner surface.
Stamens 8 mm. long, the antheriferous portion narrow, about 3.5 mm.
long, inappendiculate. Fruit fleshy, glabrous, ellipsoidal, about 12 mm.
long.

Luzon, Province of Bataan, Mount Mariveles (816, 1814, 2938 Borden) June,
September, 1904, and March, 1905. No. 1953 Cuming, Philippines, without
locality, of which a fragmentary specimen exists in our herbarium, appears to

belong here.
PHRYGILANTHUS Kichl.

Phrygilanthus obtusifolius Merrill sp. nov.

Glabrous throughout. Shrubby, much branched, about 1 m. long, the
branches slender, brown or yellowish when dry, rigid, terete, the younger
branches often somewhat compressed below the nodes, the nodes promi-
nent. Leaves opposite, oblong-obovate, 3 to 6 cm. long, 1 to 2.5 em. wide,
the:apex broad, rounded, gradually narrowed below to the cuneate base,
coriaceous, dull, finely wrinkled on both surfaces when dry, the nerves,
including the midrib, obsolete or nearly so; petioles 3 to 5 mm. long.
Cymes glabrous, axillary and terminal, few, usually 2-flowered, solitary
or two or three from each node, the peduncles slender, 1 to 2 cm. long,
the pedicels 3 mm. long. Flowers white about 1 cm. long the bract small,
rounded, about 1 mm. long. Calyx glabrous, cylindrical, truncate, 3 mm.
long, the limb slightly exceeding the ovary.- Petals 5, free, slightly
enlarged, and 2 mm. wide below, narrowed above, the upper three-fifths
curved-reflexed from the insertion of the stamens. Filaments 6 mm.
long; anthers oblong, obtuse, versatile. Style glabrous, about 12 mm.

190

long, slender. Fruit yellowish or bright red, glabrous, fleshy, ovoid, 6
to 7 mm. long.

Luzon, Province of Bataan, Lamao River, Mount Mariveles (1081, 134 Whit-
ford) February, 1905, May, 1904; (1813 Borden) September, 1904. Parasitic
on Hugenia and other trees in forests at about 600 m. about the sea.

The first species of the genus to be found in the Philippines, the other known
species being mostly confined to tropical America and Australia. In gross char-
acters and habit strongly resembling Loranthus nodosus (Van Tiegh.) Jngl.
Loranthus sp. Merr. Phil. Journ. Sci. Suppl. 1 (1906) 50.

OLACACEA®.
XIMENIA Linn.

Ximenia americana Linn. Sp. Pl. (1753) 1193; Mig. Fl. Ind. Bat. 1 (1855)
786; Masters in Hook. f. Fl. Brit. Ind: 1 (1875) 574; I.-Vill. Nov. App.
(1883) 45.

BASILAN (3467 Hutchinson) December, 1905. Along the seashore, common,

Moro, Pangungan. PALAWAN, Malinao River (3799 Curran) February, 1906.

This widely distributed species has previously been reported from the Philip-
pines only by Fernandez-Villar, who states that he saw living specimens in many
localities in the Province of Iloilo, Panay. So far as is known, the specimens
cited above are the only Philippine ones extant. Tropical shores of America,

Africa, Asia, and Malaya.

BERBERIDACE4.
MAHONIA Linn.

Mahonia nepalensis DC. Syst. 22 (1821) 21; Prodr. 1 (1824) 109; Fedde in
Engl. Bot. Jahrb. 31 (1901) 120. Berberis nepalensis Spreng. Syst. Veg. 2
(1825) 120; Hook. f. Fl. Brit. Ind. 1 (1872) 109; Forbes & Hemsl. Journ.
Linn. Soe. Bot. 23 (1886) 31.

Luzon, Province of Benguet, Baguio (5929 Hlmer) March, 1904; (18 Topping)
ebruary, 1903. An interesting addition to our knowledge of the Asiatie element
in the Philippine flora, the species having previously been known from the
mountains of British India, Java, China, and Japan. No. 5929 Himer has been
examined by Schneider, who has recently monographed the genus Berberis,” and
who verifies the above identification, although stating that in the absence of
flowers the determination can not be absolutely certain. ‘Topping’s specimen is
with flowers, and offers no characters by which I am able to distinguish the
Philippine form from the above species.

ANONACE.
UNONA Linn.

Unona merrittii Merrill sp. nov. § Stenopetalon.

A tree 20 to 25 m. high, glabrous throughout except the slightly
puberulent flowers, the flowers greenish yellow, very fragrant, in fascicles
from small tubercles on the branches below the leaves. Branches gray
or grayish brown, quite glabrous throughout, striate when dry. Leaves

1 Bull. Herb. Boiss (1905) II; 5.

191

oblong-lanceolate to elliptical-lanceolate, glabrous, subcoriaceous, shin-
ing above, pale and somewhat glaucous beneath, 10 to 14 cm. long, 2
to 3 em. wide, the base acute, slightly inequilateral, the apex acute or
obscurely sharp acuminate; nerves about 15 on each side of the midrib,
indistinct, scarcely more prominent than the secondary nerves and lax
reticulations ; petioles glabrous, 5 mm. long. Flowers in fascicles of 4
to 10 or more on the branches below the leaves, the pedicels slender,
glabrous, 1.5 to 2.5 cm. long. Calyx pubescent, the lobes orbicular-
reniform, rounded, about 1.2 mm. long, wider than long. Petals 6,
2-seriate, equal, narrowly oblong blunt, not or but slightly narrowed
above, puberulent, 1.6 to 2 cm. long, 4 mm. wide. Stamens about 1
min. long, nearly as wide, the connective very shortly produced, truncate,
concealing the cells. Ovaries many, glabrous, oblong, 1 mm. long, the
ovules 2, on the ventral suture; stigma elliptical-ovoid, equaling the
ovary in size. Fruit unknown.

Mrnporo, Bongabong River (1447 Whitford) February 23, 1906 (type); (3712
M. L. Merritt) March 20, 1906. Delta of the river slightly above sea level, in
forests. A species apparently related to Unona desmantha Hook. f. et Th., but

quite distinct from that species. The flowers have the same color and odor as
Canangium odoratum Baill.

MYRISTICACE.
MYRISTICA Linn.

Myristica nivea Merrill sp. nov.

A tree about 12 m. high, glabrous. Branches terete, striate, glabrous
except the innovations. Leaves chartaceous, oblong, the base rounded, the
apex acute, glabrous above and somewhat shining, beneath glabrous, white,
the midvein thick; nerves 14 to 15 on each side of the midrib, prominent,
curved, brown, anastomosing near the margin, the secondary and tertiary
nerves indistinct, the reticulations lax; petioles glabrous, suleate, rugose
when dry, 1 to 3 cm. long. Staminate inflorescence axillary, few flowered,
the peduncle very short, the pedicels 3 mm. long, ferruginous pubescent,
shorter than the flowers. Flowers subovoid, 7 to 9 mm. long, fasciculate,
outside rufous-pubescent, inside glabrous, three parted for the upper one-
fourth or one-fifth, the basal bract broadly ovate, obtuse, pubescent
outside. Anthers about 10, 3 to 4 mm. long, narrowly linear, connate,
glabrous, the stipe shorter than the column. Fruit solitary, short and
thickly pediceled, broadly ovoid, subtruncate or rounded at the base,
rounded at the apex, densely rufous pubescent or puberulent, 3 cm. long,
the pericarp thick, the aril laciniate nearly to the base, the areolae narrow.

BASILAN (3454 Hutchinson) December, 1905 (Staminate flowers). MINDANAO,
District of Zamboanga, San Ramon (Copeland) Mareh, 1905 (fruit).

A species apparently most closely related to Myristica mindanaensis Warb., of
the known Philippine species of this genus, differing from it in its fewer nerved
leaves, fasciculate, much larger flowers, and other characters. Myristica nivea

192

is well characterized by the very white under surface of its leaves and very
obscure, almost obsolete, lax, not parallel reticulations. It is known to the
natives of Basilan as Nyatnyat.

LAURACEZ.
ACTINODAPHNE Nees.

Actinodaphne philippinensis Merrill sp. nov.

A tree 15 to 20 m. high, quite glabrous except the inflorescence, with
penninerved oblong-ovate to elliptical-ovate, alternate, subcoriaceous
leaves which are somewhat glaucous beneath, and short, axillary, racemose
inflorescence, the clusters of flowers pediceled. Branches brown, gla-
brous, the younger ones nearly black when dry. Leaves 8 to 15 em. long,
4 to 8 em. wide, shining above, the base rounded or acute, rarely slightly
inequilateral, the apex acute or obscurely acuminate; nerves about 10
on each side of the midrib, prominent beneath, scarcely anastomosing,
the ultimate reticulations dense; petioles 2.5 to 4 cm. long. Racemes
axillary or from the axils of fallen leaves, rather densely ferruginous
puberulent, the axis 1 cm. long or less, the peduncles 1 cm. long or less,
puberulent, the bracts 6, orbicular ovate, concave, rounded, 6 to 7 mim.
long, veined, glabrous or slightly pubescent. Flowers about 7 in each
cluster, yellow, the perianth segments 6, sometimes 5, lanceolate, acute
or acuminate, about 4 mm. long, 1.5 mm. wide, somewhat pilose, the
tube about 1 mm. long. Stamens 9, all fertile, the anthers 4-locellate,
all introrse; filaments about 4 mm. long, pilose below, the inner three
glandular at the base. Pistillate flowers and fruits not seen.

°

Minporo, Bongabong River (1476, 1416 Whitford; 3667 Merritt) February,
March, 1906, the former with mature flowers, the latter two with buds only.
In forests, delta of the Bongabong River, slightly above sea level, the first species
of the genus to be reported from the Philippines. T., Bakan.

CRYPTOCARYA R. Br.

Cryptocarya acuminata Merrill sp. noy.

A tree 8 to 12 m. high with alternate ovate-lanceolate to oblong-lan-
ceolate, rather slenderly sharply acuminate penninerved leaves which are
paler and glaucous beneath, and axillary and terminal panicles which
are densely ferruginous pubescent with short shining hairs. Branches
light brown, rather densely ferruginous puberulent. Leaves subcor-
iaceous, shining and glabrous above, beneath glabrous or minutely puber-
ulent on the midrib and nerves when young, the base rounded or acute,
gradually narrowed above to the slender acuminate apex; nerves 8 to 10
on each side of the midrib, prominent beneath, ascending, parallel,
scarcely anastomosing, the reticulations indistinct; petioles ferruginous
puberulent, 1 to 1.5 em. long. Panicles 5 to 7 em. long, the peduncles
3 to 4 em. long. Perianth segments 6, elliptical ovate to oblong ovate,

193

acute or obtuse, subequal, densely ferruginous puberulent outside, about
2 mm. long, about equaling the tube. Fertile stamens 9, the filaments
short, somewhat pubescent, the anthers 2-locellate, those of the first and
second series introrse, those of the third series extrorse, the fourth series
of large stipitate, cordate, acuminate, staminodes. Ovary narrowly
ovoid, glabrous, narrowed above into the style. Fruit black when dry,
glabrous, or slightly puberulent, shining, subglobose, about 7 mm. in
liameter.

Mrnporo, Bongabong River (3673 Merritt; 1425 Whitford) March, February,
1906. In forests near the river slightly above sea level.

DEHAASIA Blume.

Dehaasia triandra Merrill sp. nov.

A tree 10 to 15 m. high, glabrous, only the three inner stamens fertile,
the outer series reduced to staminodes. Branches light gray, glabrous.
Leaves ovate-lanceolate to oblong-ovate, glabrous, subcoriaceous, some-
what shining on both surfaces, 10 to 20 em. long, 4 to 7 cm. wide,
narrowed below to the acute base and above to the sharply acuminate
apex; nerves 7 to 9 on each side of the midrib, curved ascending, rather
distinct beneath, the reticulations rather obscure, lax; petioles 1.5 to
2 em. long, nearly black when dry. Panicles several from the young
portions of each branchlet, glabrous, black when dry, 4 to 10 cm. long,
few flowered, the peduncles 5 cm. long or less, the branches usually
spreading, the flowers usually in threes at the tips of the branchlets.
Flowers small, hermaphrodite. Calyx segments 6, deciduous, broadly
ovate, acute, the inner three about 1.5 mm. long, the outer three some-
what smaller, slightly ciliate on the margins. Fertile stamens 3 only,
in the inner row, the filaments pubescent, glandular at the base, the
anthers 2-celled, extrorse, the stamens of the router series reduced to
flattened pubescent staminodes. Ovary narrowly ovoid, glabrous, about
1 mm. long, narrowed above into the 1 mm. long style. Fruit elliptical
to elliptical oblong, glabrous, 3 to 3.5 cm. long, 1.5 to 2 em. thick, the
pedicel when fresh 5 em. long or less, at least as thick as the mature
fruit, fleshy, bright red, when dry 2 to 5 em. long, less than 1 em. thick,
black.

Mrnporo, Bongabong River (3751 Merritt; 1459 Whitford) March, February,
1906. Maspatr (3068 Merrill) August, 1903. Luzon, Province of Rizal (2000
Ahern’s collector) November, 1904.

An anomalous species, differing from Dehaasia as described in its 3, not 9,
fertile stamens, but so evident are the fruit characters that I have not hesitated
to refer it to the above genus. In connection with this species I have carefully
studied the description of Salgada lawrifolia Blanco, referred by F.-Villar to
Busiderorylon borneense. The above specimens do not at all agree with Blanco’s

description, and to me it appears very doubtful if Salgada can be referred to
Busideroxylon with satety.

j 194
CRUCIFEREE.
LEPIDIUM Linn.

Lepidium ruderale Linn. Sp. Pl. (1753) 645; Hook. f. & Andr. in Hook. f. FI.
Brit. Ind. 1 (1872) 160.

Luzon, Province of Cagayan, Tuguegarao (188 Merrill) June 8, 1902. In dry,
open, waste places, a weed, apparently introduced, but well established. Not
previously reported from the Philippines. Kurope, Asia, North America, and
Australia.

CARDAMINE Linn.

Cardamine parviflora Linn. Sp. Pl. ed. 2 (1763) 914; DC. Prodr. 1 (1824)
152; Forbes & Hemsl. Journ. Linn. Soe. Bot. 23 (1886) 44.

Luzon, District of Lepanto, Balili (4609 Merrill) November 5, 1905, in a coffee
plantation, damp shaded soil at about 1,600 m.; Province of Benguet, Baguio
(5846 Elmer) March, 1904, in open damp soil along streams at about 1,400 m.

The first species of the genus to be reported from the Philippines, apparently
native, and not distinguishable from the widely distributed Cardamine parviflora
Linn., which extends from North America to Europe, northern Africa, and tem-
perate Asia southward to Formosa.

ROSACEA.
RUBUS Linn.

Rubus copelandi Merrill sp. noy.

An ascending plant 1 to 1.8 m. high, with trifohate leaves and solitary
axillary white flowers, or sometimes two or more pedicels from the upper
axils, the stems, petioles, and pedicels with usually slender, recurved
spines, and slender spine-like capitellate hairs. Stems terete, greenish
or reddish, not at all pubescent, the spines and intermixed capitellate
hairs rather numerous. Leaves alternate, trifohate, the petioles about
3 cm. long, spiny and. with capitellate stiff hairs, the stipules ovate-
lanceolate to oblong ovate, acuminate, nearly 1 cm. long, foliaceous,
capitellate hairy ; leaflets ovate to elliptical-ovate, acuminate, firm, brittle
when dry, glabrous on both surfaces, except the somewhat pubescent
midrib above, paler beneath, 4 to 7 em. long, 2 to 4 cm. wide, rather
abruptly acuminate, the base broad, rounded, that of the lateral leaflets
often inequilateral, the margins dentate or bidentate, the teeth small,
mucronate acuminate; nerves about 10 on each side of the midrib, rather
prominent, parallel, the midrib and often the lateral nerves with retrorse
spines beneath, petiolules of the terminal leaflets about 1.5 em. jong,
of the lateral ones 1 to 2 mm. Flowers white, 2.5 to 3 em. in diameter,
the pedicels 1 em. long or less, spiny and with capitellate hairs. Calyx
cinereous pubescent or puberulent, the lobes with few slender spines and
capitellate hairs, ovate lanceolate, acuminate, often 1 em. long, 5 mm.
wide or less. Petals elliptical, rounded, somewhat narrowed below,
about 12 mm. long, 8 mm. wide, distinctly veined. Stamens indefinite ;

195

filaments 4 to 5 mm. long, glabrous; anthers 1 mm. Ovaries many,
glabrous. Receptacle glabrous. Fruit red, ovoid about 1.5 em. long,
glabrous.

Luzon, Province of Benguet, Pauai (4810 Merrill) November 8, 1905. In
thickets near border of the mossy forest at about 2,200 m. A species characterized
by its trifoliate leaves, solitary large flowers, and peculiar capitellate, almost
spine-like, hairs.

Rubus luzoniensis Merrill sp. noy.

A scandent shrub 5 to 6 m. high, with simple leaves, and terminal lax
panicles, the branches, inflorescence, and under surface of the leaves
densely pale ferruginous pubescent. Branches brown, terete, stiff, with
scattered, small, usually somewhat retrorse spines, densely ferruginous
pubescent. Leaves coriaceous, ovate to elliptical-ovate, 5 to 10 em. long,
4 to 7 em. wide, the base broad, truncate to slightly cordate, the apex
acute or blunt, the margins rather finely dentate, not lobed, the upper
surface dull or shightly shining, somewhat pilose on the midrib and nerves
and also with few scattered long hairs on the lamina, becoming nearly
glabrous, the under surface pale and very densely pubescent, the midrib
and nerves also with scattered long hairs, the midrib with few scattered
spines ; nerves 6 to 7 on each side of the midrib, prominent, ascending,
nearly straight, the reticulations prominent, subparallel; petioles 1 to
2.5 em. long, densely pubescent and with small scattered spines. Inflores-
cence a terminal, rather lax panicle often 30 em. long, the rhachis,
branches and branchlets very densely ferruginous pubescent or pilose, and
with scattered small spines, the branches irregular, distant, usually spread-
ing, the lower ones often 15 cm. long, few flowered. Flowers white, short
pediceled, about 1.6 em. in diameter, the bracts ovate lanceolate, densely
pubescent, 5 mm. long or less. Calyx very densely ferruginous pilose,
the lobes ovate to oblong-ovate, acute, about 6 mm. long. Petals oblong
obovate, gradually narrowed below, obtuse, 6 to 7 mm. long, 3 to 4 mm.
wide. Stamens indefinite ; filaments glabrous, about 5 mm. long; anthers
0.5 mm. long. Ovaries many, somewhat pilose on one side aboye or
nearly glabrous, the receptacle pilose. Fruit unknown.

Luzon, District of Lepanto, Mount Data (4596 Merrill) November 4, 1905.
In the mossy forest at about 2,250 m. Sterile specimens of another species were

collected in the same locality, strongly resembling the above, but the leaves quite
glabrous or at least only slightly pilose beneath.

LEGUMINOSEZE.

INDIGOFERA Linn.

by

Indigofera linifolia Retz. Obs. 4 (1779-1791) 29; Baker in Hook. f. FI. Brit.
Ind. 2 (1876) 92.

Luzon, Province of Benguet, Ambuklao to Daklan (4387 Merrill) October 25,
1905. On dry, rocky banks in open grass lands at an altitude of about 900 m.
Not previously reported from the Philippines. Abyssinia to Afghanistan, British
India, Malaya, and northern Australia.

196
MUCUNA Adans.

Mucuna acuminata Merrill sp. nov. § Carpopogon.

A long slender climber reaching a height of from 12 to 15 m. with
slender glabrous branches, glabrous trifoliate leaves and short or long
peduncled pendulous inflorescence, the flowers pale green or nearly white,
5 to 5.5 em. long, the pods about 20 cm. long, rather strongly abruptly
acuminate, without transverse plates and nearly glabrous, broadly winged
down both sutures. Petioles 8 to 10 cm. long, glabrous or very slightly
pubescent, the petiolules about 5 mm. long; leaflets ovate to elliptical
ovate, 10 to 18 em. long, 6 to 11 cm. wide, abruptly caudate-acuminate.
the base rounded or subcordate, the terminal one regular, the lateral
ones inequilateral glabrous, or when young with few appressed hairs ;
stipels subulate, nearly 5 mm. long. Peduncles slender, 20 to 60 em.
or more in length, the flowers paniculate at the apex, the branches
5 em. long or less, the pedicels 1 to 1.5 cm. long, pubescent; bracts and
bracteoles early deciduous. Calyx cup-shaped, 1 cm. long, about as broad,
densely ferruginous and cinerous pubescent, but without stinging hairs,
suboblique, the teeth short, broad. Keel and wings subequal, the former
somewhat inflexed at the apex, the wings clawed, auricled at the base,
pubescent on the margins below; standard about 2.5 cm. long, wider
than long, retuse. Pod oblong, 18 to 21 cm. long, 5 to 5.5 em. wide,
4 to 6 seeded, black when dry, shining, reticulate, but without plaits and
stinging hairs, often somewhat pubescent below, the base acuminate, the
apex abruptly apiculate-acuminate, the acumen 1.5 to 2 cm. long, the
marginal wings about 1 cm. wide.

Minporo, Baco River (4069 Merrill) March, 1905 (type); (220, 322 McGregor)
April-May, 1905. ‘In thickets along the river. Possibly No. 2955 Ahern’s col-
lector, from Bosoboso, Province of Rizal, Luzon, is the same, but the specimens
are with immature flowers only.

A species closely related to Mucuna gigantea DC., differing from that species
in its larger pods, which are without stinging hairs and which are rather long
apiculate-acuminate, rather larger flowers and leaves, the leaflets also differing
in shape and size. Differing from J. longipedunculata Merr., in its much smaller
flowers.

Mucuna luzoniensis Merrill sp. nov. § Stizolobium.

Scandent, the trifoliate leaves 20 cm. long or less, the short peduncled
racemes 15 to 40 em. long, densely flowered, the flowers black-purple,
about 3.5 em. long, the pods flattened, scarcely turgid, not plaited or
winged, straight, 9 em. long, 1.5 cm. wide, densely covered with réddish-
brown pungent stinging hairs. Petioles 8 to 10 em. long, pubescent,
the petiolules very densely pubescent, about 5 mm. long; terminal leaflet
broadly ovate, rounded-truneate at the base, the apex rounded, apiculate,
the lateral leaflets somewhat larger, 8 to 9 em. long, 6 to 8 em. wide,
irregularly broadly truneate at the base, the apex rounded, apiculate,

LOY

beneath very densely grayish pubescent, above pubescent, especially on
the nerves; nerves rather prominent, 6 to 7 pairs. Racemes densely
cinereous pubescent and with brownish stinging hairs. Pedicels about
5 mm. long. Calyx about 1 cm. long, the teeth acute or acuminate,
cinereous pubescent and with numeruos long, brownish red, stinging hairs.
Corolla black-purple, about 3.5 em. long; standard about 2 cm. long ;
wings slightly shorter than the keel, the latter somewhat inflexed at the
apex. Pod flat, straight, abruptly hooked at the apex, with about 5
seeds.

Luzon, Province of Union, Bauang (5999 Elmer) February, 1904, distributed
as Mucuna sericophylla Perk., from which it differs in its apiculate, not retuse
leaflets, somewhat smaller flowers, and much broader pods.

Mucuna lyonii Merrill sp. nov. § Stizolobium. Negretia mitis Blanco (non
Beauv.) Fl. Filip. ed. 1 (1837) 588; ed. 2 (1845) 410; ed. 3, 2 (1878)
388; Naves_l. c. t. 405 bis. Mucuna nivea F.-Vill. (non DC.) Nov. App.
(1883) 63.

A scandent annual vine with somewhat pubescent, with apiculate
leaflets, axillary racemes 30 to 40 cm. long, of very pale green, nearly
white flowers 4 to 4.5 em. long and narrowly oblong pods 10 to 12 cm.
long, 2 cm. wide, densely covered with brown or gray hairs, not plaited
or winged. Stems striate, pubescent with appressed grayish hairs. Leaves
25 to 40 cm. long, the leaflets three, membranous, the terminal one broadly
ovate, regular, the apex acute or obtuse, the base subtruncate-acute, 11 to
14 cm. long, 9 to 11 cm. wide, the lateral leaflets 20 cm. long and 12 cm.
wide, or less, oblique-truncate at the base, much broader on one side
of the midrib than on the other, the apex apiculate, glabrous and shining
above, beneath paler and with scattered appressed rather long erayish
hairs; petioles 30 cm. long or less, somewhat pubescent; petiolules
nearly 1 cm. long, rather densely pubescent ; stipels subulate, about 2
mm. long. Racemes 30 to 40 em. long, many flowered, more or less
appressed grayish pubescent. Flowers in groups of threes, the inter-
nodes 1.5 to 2 cm. long, the pedicels about 8 mm. long. Calyx pale
green, rather densely appressed grayish pubescent, the upper tooth
triangular-ovate, acute, 7 to 8 mm. long, the lower lanceolate, acuminate,
1 cm. long, the two lateral ones lanceolate, acute or acuminate, 5 mm.
long. Standard broadly elliptical-ovate, 2.5 em. long, 2 cm. wide, obtuse,
minutely retuse; wings 4 cm. long, 1.5 cm. wide, obtuse, auriculate at
the base, claw short; keel equaling the wings, the upper 1 cm. inflexed,
auricled at the base, the claw short. Ovary hirsute. Pods slightly
turgid, somewhat curved, mucronate at the apex, slightly compressed
between the seeds; seeds about 6.

Description from plants raised in Manila by W. S. Lyon of the Bureau of

Agriculture, the seeds from Pampanga Province, Luzon, known to the Pampangans
as Sabual. Blanco gives only the Spanish names Habas and Garbanzos.

198

PTEROCARPUS Linn.

Pterocarpus klemmei Merrill sp. nov.

A large tree with glabrous shining leaves, terminal panicles and yel-
low flowers, the pods densely dark brown velvety pubescent when young
and with many short soft spine-like processes in the. central portion.
Branches lenticellate, glabrous. Leaves alternate, about 20 cm. long,
the rhachis slightly and sparingly pubescent with very short hairs;
leaflets alternate, about 10, thinly coriaceous or submembranous, ovate to
oblong-ovate, acuminate, the base broad, rounded, 5 to 7 cm. long, 3 to 4
cm. wide, glabrous and shining on both surfaces; nerves about 8 on each
side of the midrib, the reticulations close, distinct; petiolules 2 to 3 mm.
long. Panicles about 20 em. long. Flowers yellow, about 1.5 cm. long.
Calyx about 9 mim. long, the lobes short, acute or acuminate, pubescent
with few scattered short hairs. Pods (immature) oblong-ovate, about
3 em. long, thin, the stipe short, the pedicel about 1° cm. long, the
periphery of the pod from stipe to style nearly straight or slightly
convex, the style subulate, both surfaces densely dark brown velvety
pubescent, shining, the wings without spine-like processes, but the pod
proper with numerous soft spine-like processes 2 mm. long or less.

Luzon, Province of Cagayan, Palanan (4275 W. Klemme) June, 1906.

A species in vegetative and floral characters resembling Pterocarpus indicus
Willd., and P. echinatus Pers., differing from both in its densely velvety young
pods, from the former in having soft, spine-like processes on the pods and from
the latter in having fewer and shorter, soft, spine-like processes instead of
numerous stiff spines. The fourth species of the genus to be found in the Phil-
ippines, possibly a fifth being represented by No. 1002 Clark, Masbate, this
specimen having densely velvety young pods without spine-like processes.

SINDORA Miq.

Sindora supa Merrill sp. noy. Sindora wallichii var. intermedia F.-Vill. (non
Bak.) Nov. App. (1880) 71. Sindora wallichii Vidal (non Benth.) Sinopsis,
Atlas (1883) 24. t. 48. f. C.; Rev. Pl. Vase. Filip. (1886) 118; Ceron Cat.
Pl. Herb. (1892) 70; Ahern Import. Philip. Woods (1901) 80.

A tree reaching a height of 25 m. and a diameter of 140 em. with
equally pinnate leayes, coriaceous glabrous leaflets and densely pubescent
calyx lobes which have a few straight or curved spines on the upper half.
Branches and branchlets glabrous. Leaves with a glabrous rhachis 6 to
7 em. long; stipules foliaceous, 1 em. long, acute, the base rounded or
auriculate, glabrous or nearly so; leaflets 2, or mostly 3-jugate, the lower
pair somewhat smaller than those above, oblong-ovate, 5 to 8 em. long,
2.5 to 4 em. wide, very coriaceous, entirely glabrous or with few scattered
hairs on the under surface, especially on the midrib, the apex rounded,
the base acute; nerves numerous, close, faint; petiolules 4 mm. long,
glabrous. Flowers in rather dense axillary and terminal panicles 10. to
|

5 em. long, acute,

the rhachis densely pubescent; bracts 4 mm. long,

jubescent: pedicels 2 mm. long, each with two lanceolate acute pubescent
5

199

bracteoles 4 mm. long. Calyx tube short, the lobes four, thick, 1 em.
long, densely pubescent within with appressed yellowish hairs, outside
densely cinereous puberulous and in the upper half with few straight or
curved pubescent spines about 3 mm. long. Petal 1, as long as the calyx
lobes, densely appressed pubescent on the margins below. Stamina!
sheath and filaments hairy.. Ovary hirsute. Pod broadly ovate, flat-
tened, rounded at the base, the apical beak very small or nearly obsolete.
Valves dehiscent, woody, uniformly armed on the outside with strong
straight spines 5 mm. long or less and more or less densely ferruginous
pubescent, becoming quite glabrous in age. Seeds usually four, ovate,
hard, black, with an arillate funicle.

Luzon, Province of Tayabas, Pagbilao (2611 Merrill) May, 1903; Lagumanoe
(2596 Merrill) March, 1903; (23 Ware) September, 1903; (910 Whitford) Sep-
tember, 1904; (201 Merrill) Decades Philip. Forest Flora, coll. Hunt, May, 1903;
Atimonan (859 Bath) June, 1904; Guinayangan (860 Bath) June 1, 1904; (2021
Merrill) April, 1903; Baler (1010 Merrill) August, 1902.

A species related to, and previously identified with Sindora wallichii Benth.
(S. wallichiana Benth.) of the Malayan Peninsula, differing from that species’
in its glabrous leaves, and larger pods. Dr. Prain, director of the Royal Gardens,
Kew, has examined some of the material cited above and informs me that this
species is not identical with Bentham’s S. wallichiana. I have accordingly de-
seribed the Philippine plant as a distinct species, using for the specific name the
Tagalog name “Supa,” by which this important timber tree is universally known
in the Philippines.

The timber of this tree is hard and of a yellowish or reddish color, being used
in naval and general constructions, and is frequently substituted for the more
valuable “Ipil” wood (Intsia bijuga O. Ktze.). From 1900 to 1904 supa ranked
fourteenth in amount received in the local lumber markets, with a total of
177,189 feet B. M., its average price for sawed lumber being $81.50, United States
currency per 1,000 feet B. M.

In addition to being a valuable timber tree, supa also yields considerable quan-
tities of straw-colored or light-yellow, somewhat fragrant oil which burns with
a clear flame. This has been discussed by Clover” sub Sindora wallichii.

From a report submitted to the Chief of the Forestry Bureau by Mr. Kobbe,
forester, the following extracts are taken: :

“This oil (supa) is secured from the trunk of the living tree and not from
the fruit or dead wood. The tree is usually hacked with bolo cuts as high as a
man can reach and the oil runs down the channels so formed, into some vessel
so placed as to catch the product. The oil seems to be a product of the entire
woody portion of the tree and does not flow from any particular portion such
as the sap wood only. If an auger hole be bored into the heart of a living
tree, as much as 10 liters of oil is frequently obtained from the one hole. When
the trees are slashed for gathering the oil, the first that exudes is set on fire,
the heat causing a great increase in the flow of oil.”

“The oil is not widely used. There is a demand for it for the manufacture of
paint, especially for use on ships, varnish for sailboats, ete., and as an illumi-
nating oil.”

Tagalog, Supa, in Baler also Manapo.

18 Phil. Journ. Science (1906), 1, 192.

200
RUTACEA®.
ATALANTIA Correa.

Atalantia linearis (Blanco). Limonia linearis Blanco, Fl. Filip. ed. 1 (1837
357. Limonia monophylla Blanco, |. c. ed. 2 (1845) 252; ed. 3, 2 (1878)
103; Merrill, Govt. Lab. Publ. 27 (1905) 28, non Linn. Atalantia mono-
phylla ¥.-Vill. Noy. App. (1880) 27, non Correa.

A shrub 1 to 3 m. high. Branches light gray, glabrous, the young
branchlets greenish, pubescent or puberulent, terete. Leaves alternate,
linear to narrowly linear-oblong or lnear-lanceolate, glabrous, shining,
coriaceous, 2 to 7 em. long, 3 to 10 mm. wide, the base acute, the apex
blunt, retuse, entire, the margins often somewhat recurved, the midrib
prominent, the lateral nerves numerous, scarcely more distinct than the
dense reticulations; petioles glabrous or pubescent, 3 mm. long or less.
Inflorescence of terminal and axillary panicles, 3 to 5 cm. long, pubescent,
the branches short, few flowered. Flowers white, short pediceled, about
6.5 mm. long. Calyx short, regularly 5-lobed, the lobes imbricate,
broadly suborbicular, rounded, about 2 mm. long, 2.5 mm. wide, pubes-
cent, the margins ciliate. Petals 5, free, oblong, the apex rounded, 6
to 6.5 mm. long, about 3 mm. wide, somewhat narrowed at the base.
Stamens 5; filaments broad, 4 mm. long; anthers about 1.4 mm. long.
Ovary glabrous, 3-celled, sessile, the disk shallow. Fruit glabrous,
globose, white, glandular-punctate, about 8 mm. in diameter.

Luzon, Province of Rizal, Montalban (3392 Ahern’s collector) November 28,
1905; (5044 Merrill) March 8, 1906, the former in flower, the latter with mature
fruit. On cliffs and boulders at an altitude of about 40 m. along the river,
frequently in situations submerged at high water associated especially with
Bugenia mimica Merr., and sometimes with Homonoya riparia Lour. T., Dayap
na monti (Blanco), Aguhw (Ahern’s collector).

A species at once recognized by its very narrow leaves, which are retuse at the
apex. Blanco’s original description of this species is very short, his material
being from the Island of Maricaban, Province of Batangas, Luzon, flowering in
the month of July. In this description he states that the leaves are minutely
serrate, probably from the fact that in specimens with recurved leaf margins,
the slightly raised veins on the upper surface appear like minute teeth. The
note following the deseription of the species in the first edition of the Flora
de Filipinas is excluded in the second, this note referring to a similar form
observed by Blanco in the Province of Bulacan, Luzon. I am of the opinion
that the form here described is identical with Blaneo’s Limonia linearis, and
accordingly his specific name is adopted and the species is redeseribed under
Atalantia.

Atalantia retusa Merrill sp. nov.

A shrub about 3 m. high, with oblong elliptical to elliptical-ovate,
subcoriaceous, glabrous, strongly retuse leaves, racemose inflorescence
and 5- to 7-merous flowers, the stamens 10 to 15, free, the ovary 1-celled.
Branches brownish gray, glabrous, the branchlets green, glabrous. Leaves
5 to 9 cm. long, 2.5 to 4 em. wide, shining, broad at both ends, scarcely

narrowed above, the base rounded; nerves numerous, anastomosing ;

201

petioles 8 mm. long or less, the spines short or wanting. Racemes in the
upper axils, 1.5 cm. long in anthesis, densely flowered, puberulent.
‘alyx somewhat puberulent, the teeth 5 to 7, short, broad, regular, their
margins ciliate, obtuse. Petals 5 to 7, glabrous, oblong, obtuse, 5 to 5.5
mm. long, 2.5 mm. wide. Stamens 10 to 15, unequal, the filaments free,

5

2.5 to 3.5 mm. long; anthers broadly ovate, slightly exceeding 1 mm. 1
length. Ovary oblong, glabrous, 1-celled, 1-ovuled; style much shorter
than the ovary, including the stigma about 1 mm. long. Disk thickened,
ring formed. Fruit (immature) ovoid, glabrous.

PALAWAN, Puerto Princesa (3609 Curran) January 30, 1906; (195 Bermejos)
December, 1905. In old clearings and thickets, not common, at about 20 m.
above sea level.

A species apparently related to Atalantia disticha (Blanco) Merr., differing
from the latter in its leaves not being narrowed above, and in its flower characters.
In some eases a very rudimentary second cell was observed in the ovary, showing
that the species is perhaps sometimes with 2-celled ovaries. The very short
style is another aberrent character, as well as the variable number of calyx
teeth, petals, and stamens. In other species of the genus the ovary is from 2 to
5 celled, the petals and calyx teeth 3 to 5, and the style equaling or longer
than the ovary.

SKIMMIA Thunb.

Skimmia japonica Thunb. Fl. Jap. (1784) 62; Skan in Curt. Bot. Mag. IV. 1
(1905) t. 8038.

Luzon, District of Lepanto, Mount Data (4552 Merrill) November, 1905.

This species is generally distributed, but not abundant, in the mossy forest
above 2,200 m. on Mount Data, and along the high ridges between Loo and Pauai
in Benguet Province, and will doubtless be found on most of the high mountains
of northern Luzon. Vidal’ has reported .an undetermined species of Skimmia
from Lepanto, which is probably identical with the specimens here determined
as S. japonica, while in a manuscript list of identifications of Loher’s Philippine
plants, made at Kew, No. 237 Loher is determined as S. japonica, From the
material at hand I am unable to distinguish the Philippine form from Skimmia
japonica, so well figured and described in Curtis’s Botanical Magazine. The
specimens available have immature flowers and mature fruits, the inflorescence
being racemose and the flowers 5-merous. The discovery of Skimmia in the
Philippines is a very interesting addition to the northern element in the Phil-
ippine flora, the present species being previously known only from Japan and
Formosa.

POLYGALACE.
POLYGALA Linn.

Polygala glomerata Lour. Fl. Cochinch. (1790) 426; DC. Prodr. 1 (1824)
326: Benth. Fl. Hongk. (1861) 44; A. W. Benn. in Hook. f. Fl. Brit. Ind. 1
(1872) 206; Forbes & Hemsl. Journ. Linn. Soe. Bot. 23 (1886) 60.

Mrypanao, Lake Lanao, Camp Keithley (36 Mrs. Clemens) February, 1906.
A species not previously reported from the Philippines, according to Bennett
extending from the Eastern Himalayan region to the Eastern Archipelago and

4 Rey. Pl. Vase. Filip. (1886), 75.

oe

202

China, and according to Forbes and Hemsley from Southern China to the Hastern
Himalayan region southward through the Malayan Peninsula. Jing,’ however,
does not record the species from the Malayan Peninsula.
Polygala luzoniensis Merrill sp. nov. i

A perennial herb, prostrate, branched from the base, the branches
slender, pubescent, 10 to 30 cm. long. Leaves oblong-ovate, acute, firm,
1 to 1.5 em. long, 5 mm. wide or less, the base obtuse or rounded, rarely
subacute, the petiole very short, margins somewhat recurved, the nerves
3 to 4 on each side of the midrib, not prominent, glabrous or slightly
pubescent on the midrib on both surfaces. Flowers in short few flow-
ered axillary racemes, the bracts small, caducous. Outer sepals sub-
equal, 2 mm. long, oblong, obtuse, their margins with few hairs, the inner
sepals petaloid, elliptical-oblong, hyaline, rounded, 5 mm. long, 3 mm.
wide, 7-nerved from the base, the reticulations not anastomosing, gla-
brous, all persistent in fruit. Corolla about 5 mm. long, shghtly pubescent
below, the lateral petals connate for 2 mm., the free portions oblong,
obtuse, 3 mm. long, 1 mm. wide. Keel calyptrate, quite entire, crested,
the crest 1.5 mm. long, many cleft, purplish. Staminal tube, free portion,
1.5 mm. long, the filaments united throughout, the anthers sessile on the
end of the tube, oblong, about 0.4 mm. long. Ovary glabrous; style
curved, 2 mm. long, glabrous; stigma lateral. Fruit flattened, orbicular,
5 mm. in diameter, glabrous, membranous, reticulate-veined, somewhat
emarginate, the wing 1 mm. wide. Seed oboyoid, 2 mm. long, brown,
uniformly but not densely villous, the strophiole with three hyaline
appendages, one 2 mim. long, the other two about 1 mm. long.

Luzon, Province of Benguet, Baguio to Ambuklao (4368 Merrill) October, 1905.
On dry banks in grass lands at an altitude of about 1,600 m.

Polygala septemnervia Merrill sp. noy.

An erect or suberect much branched annual herb 20 to 30 em. high,
with small persistent bracts, and palmately 7-nerved, strongly reticulately
veined orbicular-ovate to subelliptical inner sepals. Branches more or
less pilose. Leaves lanceolate to oblong-lanceolate or linear-lanceolate,
2.5 to 6 em. long, 3 to 7 mm. wide, membranous, the margins usually
strongly revolute, apex and base acute, with few scattered hairs on the
upper surface and margins, the lateral nerves obscure; petioles about
1 mm. long. Racemes axillary and terminal, few or many flowered,
the flowers nodding, the axis slightly pubescent, the pedicels slender 2
to 8 mm. long, bracts persistent, lanceolate, acuminate, pubescent, about
1 mm. long. Sepals strongly reticulate-veied, the three outer ones
somewhat unequal, oblong to elliptical ovate, obtuse, 2 to 2.5 mm. long,
their margins minutely ciliate, the two inner ones petaloid, suborbicular
ovate to subelliptical, rounded, 4 mm. long in anthesis, 5 to 7 mm. long
in fruit, membranous, strongly palmately 7-nerved, the reticulations

Material for the Flora of the Malayan Peninsula (1889).

203

anastomosing, the margins slightly ciliate. Corolla about 4 mm. long,
pale or purplish, the lateral lobes 2.8 mm. long, 1.3 mm. wide, obovate,
obtuse, reticulate, the keel about 3 mm. long, crested, the crest purple,
fimbriate, 1 to 1.3 mm. long. Filaments slender, about 1 mm. long:
anthers oblong-obovoid, about 0.8 mm. long. Ovary sparingly ciliate,
compressed ; style glabrous, about 3 mm. long the stigma lateral. Cap-
sule elliptical, flattened, membranous laxly reticulate veined, about
4.5 mm. long, with very narrow marginal wings which are sparingly
ciliate. Seed elliptical or oblong elliptical, black, 3 mm. long, stro-
phiolate, densely hirsute especially, above with, grayish or brownish hairs,
the strophiole nearly white, about 0.8 mm. long, 3-appendiculate.

Luzon, Province of Benguet, Ambuklao to Daklan (4401 Merrill) October 25,
1905, in open grass lands at about 1,000 m.; Bued River (4263 Merrill) October
20, 1905, on open rock slopes at about 1,000 m.

SALOMONIA Lour.

Salomonia cylindrica Blume. Fpirhizanthe cylindrica Blume; Mig. Fl. Ind.
Bat. 1 (1859) 2: 128. ¢. 15. |

Luzon, Province of Benguet, Sablan (6054 Elmer) April, 1904.

This species is here first credited to the Philippines, and is apparently rare,
as Mr. Elmer found but three individuals, and no other collector has as yet
detected it in the Philippines. It was previously known from Java and Sumatra.
The Philippine specimens are evidently referable to Blume’s species, rather than
to the more widely distributed Salomonia aphylla Griff., which extends from
China to Borneo, Malacea and Tenasserim.

EUPHORBIACEZ.
BACCAUREA Lour.

Baccaurea gracilis Merrill sp. noy. § Pierardia,

A tree 6 to 10 m. high, quite glabrous throughout, with membranous
oblong to oblong-lanceolate leaves, slender fascicled racemes of staminate
flowers, the pistillate inflorescence short, racemose, both from the slender
branches below the leaves and in the axils of the lower leaves. Branches
light gray or brown, slender, terete, lenticellate, glabrous. Leaves
shining on both surfaces, 10 to 16 cm. long, 3 to 5 cm. wide, short
acuminate, the base acute, entire, glabrous; primary nerves 10 to 12 on
each side of the midrib, spreading, distant, the secondary nerves and
reticulations evident, rather lax; petioles rather stout, about 5 mm. long.
Staminate inflorescence: Racemes slender, glabrous, 5 to 9 cm. long, 1 to
4 in the axils of the lower leaves or of fallen leaves, the flowers glomerate,
about 10 in each glomerule, the glomerules scattered, bracts small, the
pedicels very short. Sepals 5, imbricate, glabrous, orbicular, about
1.3 mm. long. Petals 0. Stamens 5, the filaments very short; anthers
1mm. long. Rudimentary ovary present, glabrous, 2-cleft, the disk want-
ing. Pistillate inflorescence: Racemes 5 mm. long or less, sometimes
reduced to fascicles in the axils of fallen leaves. Sepals 5. Ovary

204

glabrous, 3-celled, each cell 2-ovuled. Flowers one in each bract, subses-
sile. Fruit about 4 mm. long, somewhat flattened vertically, 6 mm. wide,
obscurely 3-lobed, 3-celled, dehiscent, each cell 2-seeded, the pedicel in
infrutescence slender, 1 to 1.5 cm. long.

PALAWAN, Puerto Princesa (218, 258 Bermejos) December, 1905; Panacan
Point (3828 Curran) March. 1906. In forests near the seashore.

CLAOXYLON Juss.

Claoxylon elongatum Merrill sp. nov. § Huclaoxrylon.

A tree about 10 m. high with oblong to oblong-lanceolate, acuminate
leaves and slender elongated staminate inflorescence much exceeding the
leaves. Branchlets slightly pubescent. Leaves membranous, rigid, 12
to 20 em. long, 4 to 6 em. wide, scabrous, the base acute, the apex rather
slenderly acuminate, the margins distantly serrate, the upper surface
glabrous, dull, scabrid with numerous densely disposed small white
dots, the lower surface paler, slightly pubescent on the nerves, becoming
nearly glabrous; lateral nerves prominent beneath, 5 to 6 on each side of
the midrib, curved-ascending, the reticulations distinct; petioles 4 to 7
cm. long, slightly pubescent. Staminate inflorescence slender, axillary,
solitary, 13 to 30 cm. long, many flowered, the axis rather densely hirsute
pubescent. Flowers glomerate, white, sessile or short pediceled, 3 or 4
in the axil of each bract, the glomeruli scattered, the bracts densely
pubescent, 1 to 1.5 mm. long. Sepals 3, oblong ovate, acute, glabrous
or slightly ciliate on the margins, about 3 mm. long. Stamens about
25: filaments glabrous, about 1.5 mm. long; anther cells basifixed, about
0.5 mm. long. Disk glands scattered among the stamens, ovate to
obovate, obtuse, densely ciliate above, about 0.8 mm. long. Pistillate
flowers and fruits not seen.

Luzon, Province of Laguna, Mount Maquiling (5141 Merrill) March 17, 1906.
In forests on steep slopes at about 150 m.

Claoxylon purpureum Merrill sp. nov. § Huclaoxylon.

A shrub about 5 m. high, with shining, few nerved, usually purplish,
glabrous leaves and short, few flowered, axillary racemes, dioecious.
Branches light gray, glabrous, the ultimate branchlets often slightly
pubescent. Leaves oblong ovate to elliptical oblong, or obovate, sub-
membranous, the apex short acuminate, often abruptly so, narrowed
below to the acute base, the margins distantly sinuate crenate or dentate
to subentire; nerves 5 to 6 on each side of the midrib, distant, prominent,
curved-ascending, interarching, the reticulations prominent, lax ; petioles
0.5 to 1.5 em. long, glabrous or nearly so. Inflorescence slender, 4 em.
long or less, axillary, slightly pubescent. Female flowers few. Ovary
2-celled, each cell 1-oyvuled; style cleft nearly to the base, the two arms
recurved, about 2 mm. long. Male flowers: Sepals 3, valvate, elliptical,
ovate, about 2 mm. long, acute. Petals 0. Stamens about 16; filaments
about 1 mm. long; anther cells spreading or ascending, about 0.7 mm.

205

long; disk glands about 0.8 mm. long, among the filaments, slightly
ciliate at the apex, the staminate flowers apparently but one in each
bract. Fruit glabrous, purplish, 2-celled, each cell 1-seeded, about
6 mm. long.

Luzon, Province of Benguet, Suyoe to Pauai (4689 Merrill) November 7, 1905,
staminate and pistillate flowers from different plants; Baguio (6307 Hlmer)
May, 1904; Mount Santo Tomas (6545, 6548 Hlmer) June, 1904. Abundant in
the mossy forests in the mountains and on high ridges above 2,000 m.

MACARANGA Thouars.

Macaranga dipterocarpifolia Merrill sp. nov. § Mappa.

A shrub or small tree reaching a height of about 7 m. Branches
reddish brown, glabrous. Leaves ovate, glabrous, 13 to 20 em. long,
8 to 12 cm. wide, subcoriaceous, glabrous, penninerved, the base broad,
truncate, somewhat auriculate at the insertion of the petiole and with
several large glands at the base on the upper surface, the margins
irregularly repand-crenate, usually dull, the lower surface with numerous
small glands, usually paler than the upper surface, 7 to 9 nerved from
the base, the lower nerves short; lateral nerves prominent, spreading,
subparallel, 10 to 13 on each side of the midrib, the lower pair (basal)
sending stout branches toward the margins, the reticulations subparallel,
distinct ; petioles 5 to 10 em. long, glabrous. Male inflorescence glabrous,
paniculate, from the upper axils, 10 to 15 cm. long. Flowers numerous,
yellowish, fasciculate in the axils of the bracts, the bracts linear,
5 to 7 mm. long or less, enlarged and with 1 to 3 or 4 large disk-like
glands above, often somewhat enlarged below and with one or two lobes.
Sepals 3, 1 to 1.2 mm. long, glabrous, elliptical-ovate, acute. Stamens
7 to 9; filaments 1 mm. long or less; anthers 4-celled. Pistillate flowers
and fruit not seen.

Luzon, Province of Benguet, Sablan (6137 Hlmer) April, 1904. Not common on
dry open ridges.

A species resembling Macaranga cumingii Muell. Arg., but with much larger
leaves and more numerous stamens, the shape and venation of the leaves resem-
bling some species of Dipterocarpus, from which character the species is named.
The same species has been collected (sterile) by Whitford, in the canyon of the
Lamao River, Province of Bataan, Luzon, altitude 700 m.

CORIARIACEAD.
CORIARIA Niss.

Coriaria intermedia Matsum. Bot. Mag. Tokyo 12 (1898) 62.

Luzon, Province of Benguet, Suyoe to Pauai (4800 Merrill) November 7, 1905.
In ravines at about 2,000 m. Formosa.

Specimens of the above number were sent to Dr. J. Matsumura, of the Botan-
ical Institute, Imperial University, Tokyo, Japan, for comparison with the type
of his Formosan species, and after comparing the specimens, he expresses the
opinion that the Luzon plant is identical with his Coriaria intermedia. Specimens
collected in Benguet by Vidal, and recorded by him as “C. sp. (aff. C. japonica

206

A. Gray)”™ are undoubtedly referable to Coriaria intermedia Matsum. The
thirteen known species of the genus have a peculiar geographical distribution,
extending from the Mediterranean region to the mountains of British India,
China, Japan and Formosa, and from New Guinea to New Zealand, Mexico, and
South America. The presence of this Formosan species in Benguet adds another
very characteristic species to the known northern element in the Philippine flora.

RHAMNACE/H.
ZIZYPHUS Juss.
Zizyphus cumingiana Merrill sp. nov. Zizyphus atl. Z. baclei DC., Vidal, Phan.
Cuming. Philip. (1885) 104.

A scandent shrub reaching a height of 10 m., with broadly-ovate to
oblong-ovate, inequilateral, nearly glabrous, 3-nerved leaves, the lateral
nerves branched, solitary spines and sessile or short-peduncled cymes
about equaling the petioles. Branches dark brown, the older ones lenti-
cellate, glabrous, the branchlets slender, more or less ferruginous pubes-
cent, the spines stout, recurved, short. Leaves submembranous, shining,
glabrous, or when young the nerves on both surfaces somewhat pubescent,
the margins obscurely crenate or dentate, the teeth often apiculate, 4
to 6 em. long, 2 to 3.5 em. wide, the base rather strongly inequilateral,
broad, rounded, sometimes subcordate, the apex acuminate, the acumen
blunt, often obscurely apiculate; nerves 3, the lateral ones scarcely
reaching the apex of the leaf and sending rather bold, curved-ascending
branches toward the margin, the lower branches often similating basal
nerves, so that sometimes the leaves appear to be 4+ or 5-nerved; petioles
6 to 8 mm. long, usually somewhat pubescent. Cymes about 8 mm.
long, few flowered, slightly ferruginous-pubescent. Flowers greenish
white, about 4 mm. in diameter, the pedicels 2 to 4 mm. long. Calyx
glabrous or with very few hairs outside, the lobes ovate, acute, nearly
2 min. long, keeled within. Petals about 1.5 mm. long, obovate-spatulate,
cucullate, clawed. Disk glabrous, 10-crenate. Fruit obovoid, glabrous,
black when dry, 1-celled, 1-seeded, 6 to 7 mm. long.

PHILIPPINES (453 Cuming) 1836-1840. Luzon, Province of Tayabas, Mount
Banajao (994 Whitford) October, 1904. Minporo, Bongabong River (3616 Jer-
ritt) February, 1906. PALAWAN (Paragua) Puerto Princesa (354 Bermejos)
February, 1906; E-wi-ig River (690 Merrill) February, 1903. In forests and
thickets from the sea level to 650 m. <A species apparently related to Zizyphus
cellidifolius DC., and Z. timorensis DC.

TILIACEZ.
TRICHOSPERMUM Blume.

Trichospermum trivalvis Merrill sp. noy.

A small tree 6 to 8 m. high with oblong-ovate, cordate, often somewhat
inequilateral serrate leaves and 3-valved triangular capsules. Branches
nearly black when dry, glabrous, terete, the younger parts somewhat

Rey. Pl. Vase. Filip. (1886), 102.

207

~

ferruginous pubescent. Leaves 10 to 20 em. long, 5 to 12 em. wide,
submembranous, the base broad, cordate, usually somewhat inequilateral.
the apex acuminate, the margins rather finely irregularly serrate, pubes-
cent on the nerves and midrib above, paler and rather densely stellate
pubescent beneath, the base 5-nerved, the lateral nerves prominent, 5
to 6 on each side of the midrib, ascending, the reticulations rather lax ;
petioles 5 to 10 mm. long, densely pubescent. Inflorescence of axillary,
peduncled cymes 4 to 7 cm. long, the peduncles, branches, pedicels, and
calyces densely uniformly ferruginous stellate pubescent, the peduncles
1.5 to 3 em. long. Pedicels about 5 mm. long. Sepals free, oblong,
acute, 7 to 8 mm. long, about 2.5 mm. wide, densely stellate pubescent
outside, glabrous inside. Petals 5, about 6 mm. long, 1.5 to 2 mm. wide,
acute, glabrous except at the thickened base which is rather densely
pubescent. Stamens indefinite, inserted on the inside of a short densely
villous disk; filaments’slender, glabrous, 3 to 4 mm. long; anthers broad,
about 0.6 mm. long. Ovary sessile, oblong, densely villous, 3-celled,
each cell many ovuled. Style about 1.5 mm. long, expanded upward
into the broad somewhat cleft stigma. Capsule nearly 1 em. long,
triangular, 3-valved, the style persistent, the valves pale and glabrous
inside, densely ferruginous villous outside. Seeds many, 1.5 to 2 mm.
long, compressed, the hairs of the arillus long.

Luzon, Province of Rizal, Antipolo (398 Ahern’s collector) February, 1904;
Bosoboso (1161 Ahern’s collector) June, 1904. Minporo, Bongabong River (1391
Whitford) January, 1906.

The first species of the genus to be reported from the Philippines, differing from
all other described ones in its 3-valved capsules. Of the three other known species
of the genus one is found in Java, one in the Fiji Islands, and one in Perak and
the Nicobar Islands.

MALVACEA.
ABELMOSCHUS Medik.

Abelmoschus luzoniensis Merrill sp. nov. A. moschatus Perk. Frag. Fl. Philip.

(1904) 111, in part.

An erect, herbaceous, somewhat branched annual, the vegetative por-
tions densely velutinous pubescent and with scattered simple or stel-
lately disposed bristle-like hairs, the bracteoles 5 to 7, ovate-lanceolate.
Branches terete, the pubescence and bristles ciereous to fulvous, the latter
few or many. Leaves 9 to 15 cm. long, cordate, deeply palmately 5-lobed,
rather densely pubescent on both surfaces and with few or many stellately
disposed bristle-like hairs, the outer lobes short, the others 7 to 13 em.
long, lanceolate, rather coarsely lyrately lobed, acuminate; petioles 8 em.
long or less, densely pubescent and with few bristles; stipules linear,
about 7 mm. long. Flowers forming terminal racemes, yellow, the
pedicels equaling the petioles, similarly pubescent. Bracteoles ovate
lanceolate to oblong, 5 to 7, at length deciduous, 1 to 2.5 em. long,

208

6 to.10 mm. wide, acute, densely pubescent outside and with few stel-
lately disposed hairs inside. Calyx about 2 cm. long, cleft down one
side, toothed at the apex, deciduous, densely pubescent outside, slightly
so within. Corolla 6 cm. long or less, yellow. Capsule oblong ovoid,
acute, 5-angled, 4 to 5 em. long, about 2 cm. thick, the valves some-
what pubescent outside and beset with simple bristle-like hairs, inside
slightly pilose. Seeds many, pubescent, about 3.5 mm. long.

Luzon, Province of Rizal, Tanay (2328 Merrill) May, 1903; Antipolo (1307
Merrill) February, 1903; Bosoboso (1864 Ahern’s collector) September, 1904.
Apparently a distinct species not at all closely related to Abelmoschus moschatus
Moench., to which the first two numbers cited were referred by Perkins. Distin-
guished at once from the latter species by its broad bracteoles.

Abelmoschus multilobatus Merrill sp. noy.

An erect branched herb 1 to 2 m. high, the vegetative portions beset
with few or many simple bristle-like hairs each.from a small papilla,
otherwise glabrous, with deeply, narrowly palmately 7-lobed leaves, the
lobes again coarsely irregularly lyrately lobed. Branches stout, terete,
the bristles usually numerous. Leaves 12 to 20 em. long, deeply cordate,
the lobes narrow, lanceolate, acuminate, 9 to 17 cm. long, the lobules
3 cm. long or less, often coarsely irregularly toothed; petioles 18 cm.
long or less, usually hispid; stipules linear, about 8 mm. long. Flowers
yellow, their pedicels 10 em. long or less. Bracteoles 6 to 8, lanceolate,
about 1.5 em. long, 5 mm. wide, acuminate, with few scattered hairs and
bristles, becoming nearly glabrous. Calyx rather densely pubescent out-
side, somewhat pilose within, about 2 mm. long, deciduous, the apex
rather coarsely toothed. Petals yellow, 6 to 8 em. long. Ovary pubescent.
Staminal column antheriferous throughout. Capsule (immature) ovoid,
densely beset with simply disposed bristles, exceeding the bracteoles.

Luzon, Province of Bataan, Lamao River (1820, 1902 Borden) September,
November, 1904; (6739 Elmer) November, 1904: Province of Union, Bauang (14
Fenix) October, 1905. All the above were distributed as Abelmoschus moschatus
Moench., and the material from the Province of Bataan was so enumerated by
the author." The species is, however, very distinct from A. moschatus Moench.

DILLENIACE 2.
SAURAUIA Willd.

Saurauia clementis Merrill sp. noy.

A shrub or small tree, the branches, leaves, and inflorescence with
numerous linear, subulate chaffy bristles. Branches terete, brown, the
bristles many, short and long intermixed. Leaves membranous, slightly
shining, elliptical-obovate to oblong-obovate, 10 to 16 cm. long, 4 to 7
em. wide, rather dark above, paler beneath, somewhat narrowed below
to the acute, rarely obtuse base, more abruptly narrowed above to the
short acuminate apex, both surfaces with numerous subulate bristles

“Phil, Journ, Science Suppl. (1906), 1: 92.

209

which are more numerous beneath, especially on the midrib and nerves,
the margins subentire or bristly toothed above; nerves 7 to 8 on each side
of the midrib, rather prominent beneath; petioles about 1 cm. long,
densely bristly. Inflorescence axillary, peduncled, cymose, few flowered,
4 to 8 em. long, densely bristly throughout, the bracts lanceolate, acum-
inate, 8 to 9 mm. long, about 2 mm. wide. Sepals 5, imbricate, ovate,
acute or acuminate, about 8 mm. long, the exposed outer surfaces thickly
beset with subulate dark purple bristles about 3 mm. long, the outer
three sepals coriaceous, the inner two membranous. Corolla about 10
mm. long, the lobes obovate, rounded, strongly irregularly retuse.
Stamens 20; anthers 3 mm. long. Ovary glabrous, 3-celled; styles 3,
6 mm. long, united for the lower 1 mm.

Mindanao, Lake Lanao, Camp Keithley (57 Mrs. Clemens) January, 1906.

Saurauia longistyla Merrill sp. nov.

A tree about 10 m. igh, nearly glabrous throughout, with oblong
leaves and fasciculate or solitary flowers on the branches below the
leaves, the ovary 3-celled, the styles 3, united below. Branches light
gray, glabrous, striate, the tips with few appressed, pale, triangular
to narrowly ovate scales. Leaves 20 to 28 em. long, 6 to 9 cm. wide,
glabrous above, paler and quite glabrous beneath or with few scattered
appressed scales, firm, shining, narrowed below to the acute base, the
apex short acuminate, the margins above serrate or spinulose-serrate,
entire below; nerves about 14 on each side of the midrib, prominent
beneath, scarcely anastomosing, the reticulations subparallel, distinct ;
petioles 1 to 2.5 em. long, with few appressed scales, becoming glabrous
or nearly so. Flowers solitary or two or three in a fascicle in the axils
of fallen leaves, white or pink, about 2 cm. in diameter, the pedicels
0.5 to 2 cm. long, with few appressed scales, the bracts about 8 mm.
long, 8 mm. wide, acute or obtuse, oblong, glabrous. Sepals 5, glabrous,
concave, elliptical to obovate, 8 to 10 mm. long, 6 to 8 mm. wide, rounded
at the apex, subequal. Corolla lobes oblong-obovate, 10 mm. long, 6
mim. wide, glabrous, irregularly retuse at the apex. Stamens about 30,
inserted on the base of the corolla; filaments about 4 mm. long, anthers
2 mm. long, 1.5 mm. thick. Ovary glabrous, 3-celled; styles 3, united
for the lower 2 to 3 mm., the free portions 7 to 8 mm. long.

PALAWAN (Paragua), San Antonio Bay (835 Merrill) February 18, 1903. In
forests at about 500 m. :

Saurauia luzoniensis Merrill sp. nov.

A shrub, the branches, inflorescence, calyx lobes and petioles, minutely
ferruginous pubescent and beset with setiform scales or scale-like hairs,
the cymes axillary, solitary, 1 to 3 cm. long, few, 3 to 5-flowered, the
ovary cells and styles 3. Branches brown, the older ones becoming
glabrous. Leaves oblong to broadly oblong-oblanceolate, membranous,
10 to 17 em. long, 3 to 6 em. wide, somewhat shining when dry, paler
beneath, the midrib, nerves and reticulations beneath with few scattered

210

’

appressed scale-like hairs, the upper surface quite glabrous or with few
scales on the midrib, the margins spinulose serrate, often obscurely so,
the apex short acuminate, narrowed below to the acute base; nerves about
13 on both sides of the midrib, distinct beneath, anastomosing, the reticu-
lations netted, distinct; petioles 1 to 2 cm. long. Cymes few flowered,
strigose, the bracts and bracteoles linear, acute or apiculate, 3 to 4 mm.
long, 1 mm. wide. Flowers pink, about 13 mm. in diameter. Sepals
5, equal, oblong-ovate of oblong-obovate, 6 mm. long, 4 mm. wide,
rounded at the apex, outside ferruginous pubescent or puberulent and
beset with pale, strigose, awl-shaped, scale-like hairs 1 to 2 mm. long.
Corolla glabrous, the lobes 5, oblong-ovate, irregularly retuse at the apex,
about 7 mm. long, 4 to 5 mm. wide. Stamens 20; filaments more or less
united, about 2.5 mm. long; anthers nearly 3 mm. long. Ovary minutely
densely pubescent, 3-celled; styles 3, about 3 mm. long, slightly united
at the base.

Luzon, Province of Rizal, Bosoboso (2902, 3101 Ahern’s collector) March, May,
1905.

VIOLACEZE.
RINOREA Aubl.

Rinorea palawanensis Merrill sp. nov. § Prothesia.

A shrub 2 to 4 m. high with membranous, nearly glabrous, oblong to
obovate-oblong, acuminate leaves and few, axillary, fascicled or race-
mosely disposed flowers, the anther cells with a short mucro, the con-
nective with a large, ovate appendage. Branches gray, becoming gla-
brous, the younger ones somewhat pubescent. Leaves alternate, 15 to 27
em. long, 6 to 9 em. wide, irregularly crenate-dentate to subentire, shin-
ing, paler beneath, slightly pubescent on the nerves and midrib beneath.
becoming quite glabrous, usually rather prominently acuminate, narrowed
below to the rather abruptly acute, often somewhat inequilateral base ;
nerves 12 to 13 on each side of the midrib, distinct, distant 1.5 to 3 em..,
anastomosing, the reticulations lax, distinct. Flowers fascicled or in
short few flowered racemes, about 5 mm. long, greenish, the inflorescence
pubescent, the pedicels 6 mm. long or less, the sepals and petals accrescent
in fruit and persistent. Sepals narrowly ovate, acute, densely hirsute
pubescent outside, less so within. Petals narrowly ovate, acute, 3 to 4
mm. long, somewhat pubescent outside. Stamens from the margin of
the disk; anthers broadly ovate, 1.5 to 1.7 mm. long, each cell tipped
with short mucro 0.3 mm. long or less, the connective with a broad, ovate,
acute, subhyaline appendage from the middle, nearly 1.5 mm. long.
Ovary somewhat ferruginous hirsute; style 2 mm. long, narrowed aboye.
Capsule ovoid, 1 em. long, obtusely obscurely 3-angled, glabrous or with
few ferruginous hairs. Seeds few, glabrous, shining.

PALAWAN, Puerto Princesa (252 Bermejos) December, 1905; (3529 Curran)
January, 1906. In forests.

211

LECYTHIDACE.
BARRINGTONIA Forst.

Barringtonia curranii Merrill sp. nov. § Stravidium.

A tree 4 to 6 m. high, glabrous except the inflorescence, with large or
very large oblanceolate leaves, elongated racemes, and 4-merous flowers
which are about % em. across when open, the fruits elliptical-obovoid,
crowned by the calyx lobes, terete or very obscurely 4-angled, 7 cm. long,
about 4 em. thick. Branches thickened, light gray, the leaf scars rather
large. Leaves alternate, crowded at the apices of the branchlets, sub-
coriaceous, rather pale when dry, shining, 20 to 60 cm. long, 6 to 12 cm.
wide, the margins obscurely minutely but distantly toothed, the apex
acuminate, gradually narrowed below to the attenuate base, the petiole
stout, less than 1 em. long; nerves 18 to 24 on each side of the midrib,
very prominent, the reticulations evident. Racemes pendulous (1 m.
long according to Bermejos) stout, ferruginous-puberulent. Flowers
pink or red, sessile. Calyx tube about 8 mm. long, obscurely 4-angled.,
densely ferruginous mealy puberulent, the lobes 4, broadly ovate or
orbicular-ovate, rounded, less than 1 cm. long, densely ferruginous mealy
puberulent, the margins slightly ciliate. Petals 4, obovate, rounded,
3.5 em. long, about 2 cm. wide. Stamens indefinite, the filaments 4 to 5
em. long. Ovary 4-celled; style about 7 cm. long. Fruit wrinkled when
dry, somewhat ferruginous puberulent, with a single large seed.

PALAWAN. near Puerto Princesa (3596 Curran) January 30, 1906; (225
Bermejos) December 14, 1905. In forests and old clearings 20 m. or more above
the sea level.

Barringtonia revoluta Merrill sp. nov. § Butonica,

A tree about 12 m. high, quite glabrous, with oblong-lanceolate to
oblong-oblanceolate entire leaves, the margins strongly revolute, elon-
gated, slender, recemose inflorescence and 3-merous flowers, the calyx
tube sharply 3-angled, the lobes and petals 3, and the ovary 3-celled.
Branches slightly thickened, strongly rugose when dry, brown or erayish.
Leaves alternate, crowded at the ends of the branchlets, subcoriaceous.
pale and shining when dry, 15 to 24 em. long, 3 to 6 cm. wide, the apex
acute or slightly acuminate, gradually narrowed below to the attenuate
base, the petiole proper stout, rugose, less than 1 em. long; nerves about
14 on each side of the midrib, the reticulations rather lax, distinct.
Racemes 40 to 50 em. long or more, the flowers few, scattered, their
pedicels about 1 cm. long. Flowers immature. Calyx tube narrowed
below to the pedicel, sharply 3-angled, closed in bud, splitting into three
broadly ovate, rounded lobes, which are sometimes minutely apiculate
at the apex. Petals 3, free, elliptical ovate, rounded. Stamens indefi-
nite: anthers 0.8 mm. long. Ovary 3-celled, each cell with 3 or 4
pendulous ovules from the upper inner angle. Fruit and mature flowers
not seen.

212

PALAWAN, Iwahig Penal Settlement (3507 Curran) January 7, 1906. <A tree
not common along the river, slightly above sea level.

A very characteristic species, readily recognized by its entire strongly revolute
leaves and 3-merous flowers.

COMBRETACE/).
COMBRETUM Linn.

Combretum sexalatum Merrill sp. noy.

A scandent shrub, nearly glabrous, with opposite, oblong-ovate leaves,
terminal and axillary panicles, the flowers crowded at the ends of
the panicle branches, 4-merous flowers and prominently membranously
6-winged fruits, apparently dehiscent. Branches glabrous, brownish or
grayish, terete, the ultimate branchlets somewhat pubescent. Leaves
membranous or chartaceous, shining, the base acute or rounded, the apex
short broadly acuminate, entire, 8 to 10 cm. long, 4 to 7 em. wide, when
very young more or less glandular-lepidote, beco: sing quite glabrous, or
the nerve axils beneath barbellate; nerves 7 to 8 on each side of the
midrib, somewhat prominent, the reticulations distinct; petioles 1.5 to
2 em. long. Panicles 8 to 13 em. long, the axis and branches densely
pubescent, the lower branches subtended by leaves, the flowers densely
racemosely disposed at the ends of the branchlets. Flowers yellow, ob-
scurely glandular lepidote, the pedicels 4-angled, gradually merged into
the calyx, the flower and pedicel about 12 mm. long, the buds acute.
Calyx gradually wider upward, expanded above and villous at the throat
within, the limb 4-lobed, the lobes reflexed, ovate, sharply acute, 3 mm.
long. Petals 4, oblong, truncate, shghtly exceeding 2 mm. in length.
Stamens 8, exserted; filaments nearly 6 mm. long; anthers nearly 1 mm.
long. Style 10 to 12 mm. long. Fruit apparently dehiscent, including
the wings elliptical in outline, 3 to 5 cm. long, 2.5 to 4 cm. wide, the seed-
bearing portion 1.3 em. long or less, the back of each valve with a thin
membranous wing about 1.5 em. long, 0.5 mm. wide, the margins of the
3 valves expanded into membranous, reticulately veined wings quite
surrounding each valve, 1.5 to 2 em. wide, the free edges forming 6
longitudinal wings.

Luzon, Province of Rizal, Antipolo (3130 Ahern’s collector) June, 1905
(flower) ; Bosoboso (1868, 3321 Ahern’s collector) September, 1904, September,
1905 (fruit). The last number cited has much broader leaves and larger fruits
than the others.

MYRTACEAL.
EUGENIA Linn.

Eugenia mimica Merrill sp. nov. § /ambosa.
A glabrous shrub 1 to 3 m. high with subcoriaceous, lanceolate,
oblanceolate or oblong-oblanceolate, acute or blunt leaves, terminal cymose

panicles 6 em. long or less, the flowers small, white, the buds narrowly

213

oblong-obovoid or club-shaped. Branches gray or brownish, terete, the
branchlets sharply 4-angled. Leaves 4.5 to 9 em. long, 1 to 1.5 em. wide,
shining above, paler and dull beneath, narrowed above to the acute or
blunt apex and below to the narrow acute base, the margins somewhat
recurved ; primary nerves about 10 on each side of the midrib, distant,
irregular, spreading,
ginal nerve, the reticulations evident, netted; petioles 2 to 4 mm. long.
Flowers usually in groups of threes at the tips of the branchlets of the
inflorescence, the lower branches sometimes 3 to 4 ecm. long, the flowers

not prominent, anastomosing and forming a mar-

including the stamens 10 to 12 mm. long, narrow, the buds about 8 mm.
long. Calyx truncate about 4 mm. in diameter at the mouth, cup-shaped,
narrowed below into a 4 to 5 mm. long pseudostalk, glabrous. Petals 4.
free, orbicular, 2.5 to 3 mm. in diameter, caducous. Stamens indefinite ;
filaments about 4 mm. long; anthers 0.8 mm. long. Fruit oblong-ovoid
to ellipsoid, glabrous, dark purple when mature, about 1 cm. long, 7 to
8 mm. thick, crowned by the persistent calyx rim, with a single large seed,
the pericarp thin, crustaceous when dry.

Luzon, Province of Rizal, Montalban (5034 Merrill) March 8, 1906; (3431
Ahern’s collector) November, 1905.

A shrub growing on cliffs and boulders along the river at an altitude of about
40 m., associated with and similating in habit Atalantia linearis (Blanco) Merr.
Like Atalantia linearis and Homonoya riparia, this species is often found on
ledges and boulders subject to overflow in times of high water, and like the
former species is abundant along the river at Montalban in the limestone region,
but is found only immediately bordering the stream. So close is the resemblance
in habit and leaf characters between this species and Atalantia linearis that
Ahern’s collector, Ramos, brought in both under one number as being the same
species.

MELASTOMATACEA.
ASTRONIA Blume.

Astronia lagunensis Merrill sp. nov.

A shrub 4 to 6 m. high, nearly glabrous throughout except the in-
florescence, with oblong-elliptical, acuminate, 7-nerved leaves, and fur-
furaceous inflorescence, the orbicular petals about 5 mm. in diameter.
Branches rather stout, obscurely 4-angled, brown, glabrous. Leaves
subcoriaceous, glabrous and green on both surfaces, not furfuraceous,
14 to 18 cm. long, + to 8 cm. wide, equally narrowed below to the acute
base and above to the acuminate apex, the transverse nerves numerous,
prominent beneath; petioles stout, 5 to 6 cm. long, when young slightly
furfuraceous, becoming quite glabrous. Panicles pyramidal, 5 to 7 em.
long, 5 to 8 em. wide below, the axis and opposite branches brown fur-
furaceous, the lower branches spreading, 4 cm. long or less, the upper
ones gradually shorter. Flowers yellowish white and red. Calyx broadly
funnel-shaped, about 4 mm. long, 5 to 6 mm. in diameter above, slightly
furfuraceous, with 5 broad acute teeth less than 1 mm. long. Petals 5,

214

orbicular, 5 mm. in diameter. Stamens 10; filaments strap-shaped, 4
mm. long, 1 mm. wide; anthers dolabriform, 2 mm. long. Style about
6 mm. long.

Luzon, Province of Laguna, Mount Maquiling (5144 Merrill) March 17, 1906.
Rather common in the mossy forest at an altitude of about 1,100 m. A species
apparently related to Astronia macrophylla Blume, differing from that species
in its smaller leaves and much larger flowers.

MEDINILLA Gaudich.

Medinilla bolsteri Merrill sp. noy.

An erect glabrous shrub, 2 to 3 m. high, with opposite, petioled, 5-
nerved leaves, the transverse nerves evident, and fascicled or solitary lax
cymes 10 to 15 em. long from tubercles on the trunk, the flowers 5-merous.
Branches light gray or greenish, glabrous, terete. Leaves membranous
or chartaceous, elliptical to oblong-elliptical, short acuminate, the base
acute, 13 to 20 em. long, 7 to 9 em. wide, 5-nerved, the exterior sub-
marginal nerves from the base, the inner pair from the midrib 2 to 3
cm. above the base, the transverse nerves somewhat prominent. Pe-
duncles to the cymes slender, 4 to 5 em. long, the branches slender,
opposite or whorled, divaricate, the flowers mostly subumbellately dis-
posed at the tips of the branchlets, the pedicels 0.5 to 1 em. long,
ebracteolate. Flowers pink. Calyx cup-shaped, 4 mm. long, subtruncate
or with 5 shallow teeth. Petals 5, obtuse, irregularly obliquely oblong
obovate, 8 to 9 mm. long, 3 to 3.5 mm. wide. Stamens 10, subequal ;
filaments 4 mim. long; anthers 3 mm. long, acuminate. Style slender,
glabrous, 7 mm. long.

Luzon, Province of Rizal, Bosoboso (3262 Ahern’s collector), August 21, 1905
(type): Province of Cagayan, Tabug (173 7’. H. Bolster) August 15, 1905. In
shaded thickets by streams at about 250 m.

Medinilla dolichophylla Merrill sp. nov.

A glabrous shrub 3 to 5 m. high with 4-merous flowers, terminal and
lateral long peduncled inflorescence, and lanceolate leaves 20 to 35 em,
long, the lateral longitudinal nerves faint. Branches quadrangular, the
younger ones strongly 4-winged, the wings not undulate. Leaves mem-
branous to subcoriaceous, + to 5.5 cm. wide, pale, shining, gradually nar-
rowed above to the somewhat acuminate apex, the base acute, decurrent ;
midrib stout, the lateral longitudinal nerves four, scarcely more prom-
inent than the distant transverse nerves, the reticulations lax; petioles
about 1.5 em. long. Cymes pendulous, terminal and from the branches
below the leaves, the peduncles 15 to 20 em. long, few flowered, ebrac-
teolate. Calyx 4 to 5 mm. long, truncate, cylindrical or subcampanulate.
Petals 4, white, irregularly obovate, narrowed below, about 14 mm. long,
S mm. wide. Stamens 8, subequal; filaments 6 mm. long; anthers 6
to 7 min. long, linear, tips purplish, base white, the middle portion yellow,
the dorsal basal spur 1 mm. long, anteriorly 2-tuberculate. Style about

12 mm. long. Fruit ovoid, about 1 cm. long. Seeds oblong, rounded
at the apex, about 1.5 mm. long.

Luzon, Province of Benguet, Baguio (4332 Merrill) October 22, 1905; (44
Topping) February, 1903.

A shrub characterized by its elongated, faintly nerved leaves and pendulous
inflorescence, growing in damp shaded ravines along small streams at an altitude
of about 1,500 m.

Medinilla myriantha Merrill sp. noy.

A shrub 4 to 5 m. high, quite glabrous throughout, with opposite, 7
to 9-nerved sessile leaves and terminal many flowered panicles, the flowers
4-merous. Branches terte, rather slender, the younger ones obscurely
angled, the nodes densely barbellate. Leaves subcoriaceous, elliptical
ovate, 9 to 13 cm. long, 5.5 to 7 cm. wide, the apex abruptly short
acuminate, the base cordate, often broadly so; nerves 7, prominent,
sometimes with an additional pair of outer short nerves at the base.
Panicles 10 to 20 cm. long, pink, many flowered, the branches verticillate,
the lower ones about 9 cm. long, gradually shorter upwards, the bracts
minute, subulate. Flowers numerous, pink. Calyx urceolate-cylindrical,
4 mm. long, truncate. Petals 4, oblique, about 8 mm. long. Stamens
8, subequal; filaments 4 mm. long; anthers nearly 5 mm. long, the spur

less than 1 mm. long.

Luzon, Province of Laguna, Mount Maquiling (5129, 5133 Merrill) March 17,
1906. Abundant in the mossy forest 900 to 1,100 m. A species apparently
related to Medinilla intermedia Blume, differing from that species in its more
numerously nerved, .sessile, cordate leaves and larger flowers.

Medinilla teysmanni Mig. Ann. Mus. Bot. Lugd. Bat. 1 (1864) 217; Cogn. in
DC. Monog. Phan. 7 (1891) 593.

MINDANAO, Camp Keithley, Lake Lanao (1 Mrs. Clemens) December, 1905.
A species previously known only from Celebes and New Guinea, the specimen
cited above differing from the species, as described, in its somewhat larger flowers
but in no essential characters so far as I can detect.

PACHYCENTRIA Blume.

Pachycentria formicaria Merrill sp. nov.

A glabrous or nearly glabrous epiphytic shrub 50 cm. high or less with
3-nerved leaves, the transverse nerves very obscure or obsolete, short
petioles, and terminal corymbose inflorescence, the roots with oblong to
ovoid thickened bulb-lke swellings 1 to 3 em. in diameter. Branches
light gray to brownish, terete, glabrous or the younger branchlets
minutely furfuraceous. Leaves coriaceous, oblong to elliptical or ovate
oblong, pale when dry, dull, 6 to 12 cm. long, 1.5 to 4.5 em. wide, entire,
the base acute, the apex shortly acuminate, the lateral basal nerves
slender, ascending, submarginal, not reaching the apex; petioles 3 to 10
mm. long. Corymbs about 5 cm. long, nearly as wide, many flowered,
glabrous or nearly so, the bracteoles minute. Flowers pink. Calyx
narrow, urceolate, about 4 mm. long, constricted above the ovary, the

limb nearly 2 mm. long, with 4 rounded teeth. Petals 4, ovate-lanceolate,
about 5 mm. long, 2.2 mm. wide in the middle, narrowed below and to
the acuminate apex. Stamens 8; filaments about 2 mm. long; anthers
lanceolate, about 2 mm. long, rostrate, the basal dorsal spur about
0.5mm, long. Ovary 4-celled; style 5 mm. long.

MInpdANAO, Lake Lanao, Camp Keithley (433, 361 Mrs. Clemens) March, 1906;
(3920 Hutchinson) March, 1906.

A typical Malayan genus, of which no species has previously been reported from
the Philippines, the present species being well characterized by its thickened
bulb-like roots, which are inhabited by colonies of ants, in this respect resembling
the symbiosis found in the rubiaceous genera Hydnophytum and Myrmecoidea.

HALORRHAGACE.AE.
HALORRHAGIS Forst.

In Schindler’s recent monograph of this family '’ no species of the
genus Halorrhagis are credited to the Philippines. Schindler recognizes
59 species of Halorrhagis, of which 57 are confined to Australia and New
Zealand, one to the Asiatic continent, and one extending over the entire
range of the genus, from New Zealand and Australia to Japan, China,
and Bengal. Three species of the genus are represented in our Philip-
pine collections, one of which is apparently undescribed.

1. Calyx tube papillose-punctate, the ribs scabrous.
2. Stamens 6 to 8; inflorescence paniculate ; leaves lanceolate,

about. 1:15) "Cn: .OM Boece nee ee een en ey ee (1) H. scabra elongata
2, Stamens 4; inflorescence racemose ; leaves elliptical-oblong,
LCi, WOME OLA LSS Sis: eee ee nee ee Sensaenot tees (2) H. philippinensis
1. Calyx tube quite glabrous, not papillose-punctate, the ribs not
SCaADLOuUSee eee SELLY IEE er ee EN eR aR es ae Sane Vereen (3) H. micrantha

(1) Halorrhagis scabra (Koenig) Benth., var. elongata Schindler in Engler’s
Pflanzenreich 23 (1905) 28.

CuLIon (459 Merrill) December 13, 1902. In damp deserted rice lands near
the sea level.

This variety has previously been known only from southern China, Hongkong,
and Annam, although another variety of the same species is found in the Khasia
Mountains, British India. The Philippine plant here referred to, Halorrhagis
scabra, agrees very closely with the description and with specimens from Hong-
kong kindly supplhed me by the Director of the Hongkong Botanie Garden.

(2) Halorrhagis philippinensis Merrill sp. nov.

A seabrous herb 8 to 20 cm. high, erect or spreading, more or less
branched from the base, with simple racemose inflorescence, the flowers
solitary in the axils of the bracts, the calyx manifestly papillose-punctate,
scabrous on the ribs but not parallel callose, the stamens four. Branches
slender, appressed-pilose, 4-angled. Leaves elliptical oblong, coriaceous,
sparingly pilose, 5 to 10 cm. Jong, 2.5 to 5 em. wide, short petioled or

subsessile, mucronate acuminate or acute, the base acute, the margins

* Engler’s Pflanzenreich 23, 1905.

217

usually serrate, strongly cartilaginous. Racemes simple, the pedicels
less than 1 mm. long; bracts foliaceous, ovate-lanceolate, 2 to 3 mm.
long, sparingly scabrous-pilose; bracteoles 2, membranous, linear, spar-
ingly pilose, 1 to 1.5 mm. long. Calyx 2 mm. long, the tube manifestly
papillose-punctate, 8-ribbed, 1 mm. long, the ribs scabrous, the lobes 4,
erect, ovate-lanceolate, acute, 1 mm. long, navicular, somewhat pilose on
the keel. Stamens 4; anthers 1.2 mm. long. Fruit grayish, somewhat
shining.

‘Luzon, Province of Benguet, Baguio to Ambuklao (4357 Merrill) October 24,
1905: District of Lepanto, Suyoe to Cervantes (4444 Merrill) October 30, 1905;
Mount Data (4553 Merrill) November 4, 1904. On dry, open, grassy slopes in
thin pine forests, 1,500 to 2,100 m.

(3) Halorrhagis micrantha (Thunb.) R. Br. Schindler 1. ¢. 42.

Necros, Canlaon Voleano (Banks) March, 1902. Minpanao, District of Davao,
Mount Apo (1050 Copeland) April, 1904; (1436 Copeland) October, 1904, det.
Rolfe. “Grows all over the summit but abundant only about pools, 2,600 to
3,000 m.” Copeland.

A species not previously reported from the Philippines, extending from Bengal
to Japan, Malaya, Australia, and New Zealand. (

MYRIOPHYLLUM Linn.

Myriophyllum spicatum Linn. Schindler 1. ¢. 90.

MrinpANnao, Lake Lanao, Camp Keithley (450 Mrs. Clemens) April, 1906.
Altitude about 800 m.

No species of the genus has previously been reported from the Philippines.
Widely distributed in the tropical and temperate regions of the world, but not
recorded by Schindler from the Malayan region.

ARALIACEE.

ACANTHOPANAX Decne. et Planch.

Acanthopanax trifoliatum (Linn.). Zanthoxylum trifoliatum Linn. Sp. PI.
(1753) 270. Panazx aculeatum Ait. Hort. Kew. 3 (1789) 448. <Acanthopanax
aculeatum Seem. Journ. Bot. 5 (1867) 238; Forbes & Hemsl. Journ. Linn.
Soc. Bot. 23 (1888) 229. Aralia trifoliata Meyen.

Luzon, Province of Benguet, Bugias (4661 Merrill) October 28, 1905. In
thickets at an altitude of about 1,500 m.

An interesting addition to our knowledge of the relationship of the Philippine
flora to that of the Asiatic continent. The above specimen agrees very closely
with material of this species supplied me by the director of the Botanie Garden,
Hongkong, with Formosan specimens received from the Imperial University, Tokyo,
and with sterile specimens in Herb. Bureau of Science from a cultivated plant
in the Botanical Garden at Buitenzorg, Java. The Philippine plant is, however,
a scandent shrub, while in all the references to Acanthopanax aculeatwm Seem.,
and synonyms of that species which I have been able to consult, the species is
described as a shrub or small tree. However, I can not, from the descriptions
and material available, find any other characters by which the Philippine plant
can be distinguished, and have accordingly referred it to the above species, which
extends from’Japan and Formosa to China and Eastern India.

44772 4

218
SCHEFFLERA Forst.

Schefflera luzoniensis Merrill sp. nov. § Luschefflera, Heptapleurum.

A scandent shrub, quite glabrous except the deciduously pubescent
inflorescence, with 5-foliolate leaves, the leaflets caudate-acuminate, and
terminal panicle branched from the base, the branches ascending, elon-
gated, the flowers disposed in 3 to 5 flowered peduncled umbels. Branches
light gray, glabrous. Petioles 6 to 8 em. long, glabrous, slender ; leaflets
firm, shining, elliptical-lanceolate to narrowly obovate-lanceolate, abruptly
long caudate-acuminate, the base acute, 4 to 6 em. long, 1.5 to 2 cm. wide;
primary nerves 6 to 7 on each side of the midrib, not prominent, the
secondary nerves and reticulations nearly as evident, anastomosing and
forming a marginal nerve, the leaf margin entire, recurved; petiolules
1.5 to 2. em. long. Panicles 10 em. long or less, branched from the base,
the branches about 8 cm. long, more or less flocculose pubescent, becoming
subglabrous, the bracts and bracteoles very caducous, the branchlets
racemosely disposed, 5 to 10 mm. long, each bearing 3 to 5 umbellately
disposed flowers, the pedicels 3 to 4 mm. long. Calyx about 1.5 mm.
long, truncate. Petals 5, oblong-ovate, 2 mm. long, acute, apex inflexed,
free, falling separately. Stamens 5; filaments 2 mm. long; anthers
broadly elliptical-ovate, about 0.8 mm. long. Ovary 5-celled; style 0.
Fruit unknown.

Luzon, Province of Tayabas, Mount Banahao (928 Whifford) October 19, 1904.
Scandent on trees in forests at about 1,100 m.

Schefflera microphylla Merrill sp. nov. § Nuschefflera, Heptapleurum.

Subscandent, 2 to 3m. high; glabrous throughout, with alternate, long
petiolate, digitately 5 to 8-foliolate leaves, the leaflets lanceolate, acum-
inate, 5 cm. long or less, the flowers umbellately disposed in- lax
terminal panicles. Branches slender, terete, light gray or brownish.
Petioles slender, 4 to 5 em. long; leaflets entire, firm, shining, 3.5
to 5 cm. long, 4 to 6 mm. wide, gradually narrowed above to the
long caudate-acuminate apex, the base acute; nerves few, not prominent ;
petiolules 5 to 8 mm. long. Panicles few branched from the base, 6
to 8 em. long, the bracts and bracteoles wanting or very caducous,
glabrous; branchlets spreading, few, slender, 1 to 1.5 em. long, the
flowers in 4 to 6 flowered umbels at the ends of the branchlets, the pedicels
t to 6 mm. long. Calyx truncate or obscurely toothed, 1.5 mm. long or
less. Petals 5, triangular-ovate, acute, 1.5 mm. long, adherent by their
apices and falling as a whole. Stamens 5; filaments about 1 mm. long;
anthers broadly ovate, nearly 1 mm. long. Ovary 5-celled; style not
developed. Fruit elliptical-ovoid, glabrous, about 5 mm. long, 5-ridged.

Luzon, District of Lepanto, Mount Data (4605 Merrill) November, 1905. A

subscandent shrub growing on rotten trunks and mossy trees in the wet forest
at about 2,200 m., characterized by its small lanceolate leaflets.

219
TETRAPLASANDRA A. Gray.

Tetraplasandra philippinensis Merrill sp. nov.

A tree 8 to 12 m. high, quite glabrous throughout, with 2 to 8 or
more jugate leaves, the leaflets distantly obscurely crenate-serrate, and
7-merous flowers. Leaves alternate, the lower ones much larger than
those near the apices of the branches, the latter often reduced to 20 em.
or less in length and 2 to 3 jugate, the former 80 cm. or more in length,
8 or more jugate, the common petiole stout, joimted. Leaflets oblong-
ovate to oblong, glabrous and shining above, somewhat dull beneath,
10 to 16 cm. long, 4 to 8 em. wide, the base rounded or acute, often
inequilateral, the apex usually very abruptly short acuminate, the margins
with few distant, scattered crenate-serrate, glandular teeth; nerves about
10 on each side of the midrib, irregular, the secondary nerves nearly as
prominent; petiolules 1 to 1.5 em. long. Flowers in peduncled umbels
of 6 to 10 flowers each which are arranged in large racemose panicles,
the peduncles about 2 em. long in anthesis, 5 cm. long in infrutescence,
whorled or alternate; pedicels 4 to 5 mm. long. Calyx cup-shaped or
somewhat funnel-shaped, 2.5 to 3 mm. long, the limb slightly produced,
truneate. Petals 7, lanceolate, acute, crested on the inside at the apex,
about 3.5 mm. long, 1.2 mm. wide. Stamens about 28, several seriate,
the inner much smaller; filaments 2 mm. long or less; anthers about 1
mm. long, the cells but slightly united. Ovary 7-celled; stigma sessile.
Fruit ovoid, glabrous, about 8 mm. long, boldly 7 to 10 ridged.

PALAWAN, Casuarina Point (3809, 3844 Curran) March, 1906. Back of the
beach and in forests along the seashore.

No species of the genus has previously been reported from the Philippines; of
the 8 described species, one is from New Guinea, one from Celebes, and the remain-
ing 6 are from the Hawaiian Islands. The species here described as new is
apparently most closely related to the New Guinea species, Tetraplasandra
paucidens Miq., but differs from the latter in many characters.

ERICACEZE.
DIPLYCOSIA Blume.

Diplycosia scandens Merrill sp. nov.

A scandent more or less hispid pseudo-epiphytic shrub reaching a
height of 4 m., with ovate or narrowly-ovate, short acuminate or acute
leaves and fascicled flowers. Branches slender, gray or brown, the
younger ones rather- strongly beset with long brown appressed bristly
hairs. Leaves coriaceous 3 to 4.5 cm. long, 1.5 to 2.5 em. wide, the
base rounded, entire, gradually narrowed above, glabrous on the upper
surface or with very few hairs, beneath paler and with scattered bristly
hairs; petioles about 3 mm. Jong, densely bristly hairy; nerves 2 on
each side of the midrib, both below the middle of the leaf. Flowers

axillary, 3 to 6 in each axil, the pedicels pubescent, about 4 mm. long,
2-bracteolate at the apex, the bracteoles 1 mm. long, broader than long,
ciliate. Calyx teeth broad, ovate, obtuse, about 1 mm. long, ciliate.
Corolla white, about 5 mm. long, 3.5 mm. in diameter, the lobes broadly
ovate, acute, reflexed, nearly 2 mm. long. Stamens 10; filaments 2 mm.
long, not dilated below, glabrous; anthers about 1.2 mm. long, not
horned. Ovary superior, glabrous, globose, 5-celled, cells many ovuled ;

style about 2.2 mm. long.

Luzon, District of Lepanto, Mount Data (4597 Merrill) November 4, 1905.
Pseudo-epiphytic on mossy tree trunks in ravines, mossy forest at about 2,250 m.
The first species of the genus to be reported from the Philippines, unless
Gaultheria (Diplycosia) luzonica A. Gray, Proc. Am. Acad. 5 (1862) 324, proves
to be a true Diplycosia. Gray’s description is so short that I have been unable
to locate his species, the type being from the collections made by members of
the Wilke’s U. S. Exploring Expedition from “Luzon, in the Majaijai Mountains”
(Mount Banajao). Dr. B. L. Robinson informs me that the type is not in the

Gray Herbarium. <A second, apparently closely related but glabrous species, is
represented by No, 5932 Elmer, Province of Benguet, Luzon, the specimens with
fruit only.

RHODODENDRON Linn.

Rhododendron nortoniae Merrill sp. noy.

An epiphytic shrub with lanceolate to oblong-lanceolate, coriaceous,
acuminate leaves which are densely brown lepidote beneath, and terminal
umbellate-like fascicles of tubular crimson flowers about 4.5 cm. long.
Branches terete, brown, the branchlets densely covered with round, dark-
brown scales. Leaves 9 to 15 em. long, 1.5 to 3.5 em. wide, narrowed
below to the acute base and above to the rather long slender apex, entire,
the margins slightly recurved, glabrous and shining above, densely
covered with small round dark-brown scales beneath; lateral nerves 5 to
6 on each side of the midrib, distant, indistinct; petioles densely lepidote,
stout, 1 cm. long or less. Umbels terminal, about 12 flowered, the
bud bracts coriaceous, glabrous except the slightly pubescent margins,
ovate to oblong-ovate, acute, 1 to 1.7 em. long. Pedicels 1.3 cm. long,
glabrous, the bracts oblong to spatulate, the bracteoles filiform, caducous.
Calyx a disk about 3 mm. in diameter, obscurely 3-toothed, the teeth
rounded, small. Corolla 4.5 em. long, scarlet, the tube cylindrical, about
3.5 em. long, scarcely inflated or enlarged above, slightly curved, 6 to 7
mm. in diameter, the limb 5-lobed, the lobes broadly ovate to obovate,
rounded, 1 em. long. Stamens 10; anthers oblong, 3.5 mm. long. Ovary
narrowly oblong, the style elongated filiform, densely brown lepidote.

MinpdANAO, Lake Lanao, Camp Keithley (500 Mrs. Clemens) April, 1906.
Altitude about 800 m. Epiphytie on a large tree, associated with an epiphytic
Vaccinium.

At the collector’s request this distinct species is named in honor of Miss Norton,
of the Pacifie Grove (Calif.) Museum, the collector’s interest in botanical work
having been largely inspired by her.

Japa
VACCINIUM Linn.

Vaccinium microphyllum Blume. Bijdr. (1826) 851; Mig. Fl. Ind. Bat. 2
(1856) 1063.

MtInDANAO, District of Davao, Mount Apo (1037, 1417 Copeland) April, October,
1904. A terrestrial shrub on the wind-swept summit 2,600 to 3,000 m., the shoots
not ascending more than 10 cm. from the ground except in the more sheltered
places.

Vaccinium microphyllum Blume, was described from a specimen in Reinwardt’s
herbarium from the mountains of Celebes, the original description being very
short, but applying closely to the specimens cited above. A specimen of No, 1417
Copeland was sent to Dr. Treub, director of the Botanical Garden at Buitenzorg,
Java, for verification of the above identification. Dr. Valeton, who examined
the material, states that there is no authentic specimen of Vaccinium microphyllum
in the herbarium at Buitenzorg, but that there is a sterile specimen collected by
Teysmann in Ternate and identified as V. microphyllum by Scheffer, with which
the Mount Apo plant agrees, except that the leaves of the latter are somewhat
smaller than in the Ternate plant. The Mount Apo plant is apparently quite dif-
ferent from the species described by Beccari’ and Clarke” as Diplycosia micro-
phylla (Blume) Bece., and by King and Gamble * as Vaccinium microphyllum
Blume, but I am of the opinion that Blume’s description applies more closely
to the Mount Apo plant than to the descriptions of Malayan Peninsula and
3ornean material by the authors cited above. According to Valeton, the spe-
cimen reported from Celebes by Koorders * as Vaccinium microphyllum “Reinw.,”
is not Blume’s species, the flowers being racemose in Koorders’s specimens.

PRIMULACE.
LYSIMACHIA Linn.

Lysimachia microphylla Merrill sp. nov. § Lerousxia.

A small ascending uniformly and rather strongly hirsute-pubescent
annual 5 cm. high or less, with opposite, petioled, glandular-punctate,
obtuse or rounded leaves and short pediceled yellow flowers. Stems
slender, densely hirsute-pubescent with pale spreading hairs, simple or
slightly branched from the base. Leaves opposite, orbicular-ovate to
ovate, sometimes subreniform-orbicular, 5 to 7 mm. long, often nearly as
wide, the apex rounded or blunt, the base subtruncate to acute, glandular
punctate, uniformly hirsute-pubescent on both surfaces; petioles 1.5 to
2 mm. long, pubescent. Flowers solitary in the upper axils, the pedicels
pubescent, about 2 mm. long. Calyx cleft nearly to the base, the lobes
linear-lanceolate, slightly acuminate, about 3.5 mm. long, 1 mm. wide,
glandular punctate, strigose pubescent outside. Corolla yellow, 3.5 mm.
long, glabrous, the tube less than 1 mm. long, the lobes elliptical-ovate to

obo)

elliptical-oblong, obtuse or acute, glandular punctate. Filaments 1.5

19 Malesia (1878), 1, 212.
*° Hook. f. Fl. Brit. Ind. (1882), 3, 456.
2 Mat. Fl. Mal. Penin. (1905), 3, 273

3) 4
2 Weded.’s Lands Plant. (1898), 19, 514.

222
mm. long; anthers blunt, 1 mm. long. Capsule 4 to 5 mm. long, ovoid,
somewhat hirsute above, regularly dehiscing by 5 valves. Seeds many,
triangular, rugose, brown, nearly 1 mm. long.

Luzon, Province of Benguet, Bugias (4668 Merrill) October 28, 1905; Daklan
to Kabayan (4405 Merrill) October 27, 1905. On damp banks along the trail
in open grass lands and in thin pine forests.

A species well characterized by its small leaves, which are glandular punctate
with round spots.

SAPOTACE.
SIDEROXYLON Linn.

Sideroxylon luzoniense Merrill sp. noy.

A small tree about 5 m. high with oblong-ovate to broadly oblong-
lanceolate or elliptical-oblong, acuminate leaves, beneath shining and
rather densely ferruginous-pubescent, and axillary fascicled flowers, the
staminodes elongated, entire. Branches dark brown or nearly black,
lenticellate, glabrous, the young parts rather densely cinerous and ferru-
ginous pubescent. Leaves 7 to 9 em. long, 2 to 4 em. wide, coriaceous,
elabrous above, the apex short acuminate or acute, gradually narrowed
below to the acute base; nerves somewhat prominent beneath, 6 to 7 on
each side of the midrib, the reticulations obscure ; petioles densely pubes-
cent, 1 to 1.5 em. long. Flowers in axillary fascicles of from 2 to 5,
mostly on the young branchlets, numerous, sessile. Calyx cup-shaped,
| mm. long, densely ferruginous pubescent, 5-toothed, the teeth blunt,
broad, less than 1 mm. long. Corolla 5 mm. long, glabrous, 5-lobed,
lobes oblong-ovate, truncate, about 3.5 mm. long, 2 mm. wide below.
Staminodes linear or linear-lanceolate, entire, blunt, 2.5 to 3 mm. long,
less than 1 mm. wide. Filaments 2 mm. long; anthers broadly ovoid,
1.3 mm. long. Ovary ferruginous-pubescent, 5-celled.

Luzon, Province of Rizal, Antipolo (127 I’. W. Foxworthy) January 16, 1906.
In forest on Mount Kaysipot, at about 700 m.

OLEACE.LE. ;
JASMINUM Linn.

Jasminum populifolium Blume, Mus. Bot. Lugd. Bat. 1 (1850) 276; Miq. FI.
Ind. Bat. 2 (1856) 538; F.-Vill. Nov. App. (1883) 128; J. luzoniense
Vidal, Phan. Cuming. Philip. (1885) 185; Rey. PI. Vase. Filip. (1886) 180;
Ceron, Cat. Pl. Herb. (1892) 111; Usteri, Beitr. Ken. Philip. Veg. (1905) 120.

Blume cites no collector in the original diagnosis of his species, simply giving
the locality “Im insulis Philippinis.” It seems probable that he had before him
the same number of Cuming’s plant that Vidal later deseribed as Jasminum

luzoniense, Wis description applies very closely to a specimen of No, 1029

Cuming in our herbarium, on which number Vidal’s species was based. I have

accordingly reduced J. luzoniense to the much earlier J. populifolium,

Jasminum triphyllum Merrill sp. nov. § 7'rifoliata.

A seandent shrub, glabrous or nearly so throughout, with opposite or
subopposite, trifoliate leaves and axillary and terminal paniculate cymes.
Branches slender, glabrous, the younger ones greenish-brown. Leaflets
subcoriaceous, glabrous, ovate, slightly shining, short acuminate or
merely acute, the base broad, rounded, the terminal one 6 to 9 em. long,
3 to 4.5 em. wide, the lateral ones somewhat smaller but similar in
shape; nerves irregular, not prominent, about 6 on each side of the
midrib, the base sub 5-nerved, the marginal pair often obscure; petioles
1.5 to 2 em. long, the petiolule of the terminal leaflet 1.5 cm., of the
lateral leaflets 1 cm. or less. Inflorescence equaling or exceeding the
leaves, slightly pubescent, the peduncles 2 to 4 cm. long, the bracts small,
ovate, the bracteoles minute. Flowers white. Calyx tube cup-shaped,
about 2 mm. long, truncate or with very obscure teeth. Corolla slender,
the tube 8 mm. long, slightly enlarged above, glabrous, the lobes 5,
spreading, orbicular-ovate, obtuse, about 2.5 mm. in diameter. Filaments
short; anthers oblong, slightly acuminate, about 2.3 mm. long. Stigma
bifid, the lobes nearly 3 mm. long.

PALAWAN, Puerto Princesa (261 Bermejos) December, 1905. In forests.

A species apparently related to Jasminum parviflorum Deene., differing espe-
cially from the latter in its glabrous branches and orbicular-ovate, obtuse, not
lanceolate, acuminate, corolla lobes.

GENTIANACE.
COTYLANTHERA Blume.

Cotylanthera tenuis Blume. Bijd. 2 (1825) 707; Mig. FI. Ind. Bat. 2 (1856)
735; Dunal, in DC. Prodr. 13 (1852) 674.

Luzon, Province of Laguna, Mount Maquiling (5149 Merrill) March 17, 1906.
In forests at about 900 m.

The above identification is presumably correct, although the original and
subsequent descriptions of the species, and the descriptions of Hophylon lobbii
A. Gray, and HF. tenellum A. Gray, the former at least being apparently a
synonym of Blume’s species, are all so short that absolute identification is
impossible without comparison with the type material. The Philippine plant
when fresh, is of a pale lavender color throughout, 4 to 6 em. high, unbranched,
or with at most one branch, with solitary flowers and 3 to 5 pairs of bracts.

GENTIANA Linn.

Gentiana apoensis Merrill sp. nov. § Chondrophylla.

A low, erect, often densely tufted, more or less branched perennial
2 em. high or less, glabrous, the leaves all similar, distichous, imbricated.
Leaves ovate-lanceolate, glabrous, acute or obscurely acuminate, 5 to 6
mm. long, about 2 mm. wide, sessile, crowded, the nerves obscure. Flow-
ers solitary at the ends of the short branches. Calyx 5 mm. long
including the teeth, the latter lanceolate, acuminate, apiculate, 2 to 2.5
mm. long. Corolla tubular, about 7 mm. long, 5-lobed, the lobes broadly

ovate, acute or obscurely acuminate, about 1.5 mm. long, the tube
plaited, the teeth of the folds rounded or obtuse, entire or nearly so.
Filaments 2 mm. long; anthers 0.9 mm. long. Capsule shghtly exserted,
stalked, the stalk about 5 mm. long, the capsule broadly ovoid, com-
pressed, about 4 mm. long, the valves rather strongly 3-nerved.

MINDANAO, District of Davao, Mount Apo (1161, 1432 Copeland) April, October,
1904; (371 DeVore & Hoover) May, 19038. “Occasional in turf, but inconspicuous,
not common,” Copeland. Summit of the mountain at about 3,000 m.

No species of this genus have previously been reported from the Philippines.
Gentiana diversifolia Merrill sp. nov. § Chondrophylla.

An erect annual 2 to 3 cm. high or less, fasciculately branched, the
stems glabrous, single. Lower leaves orbicular-ovate to orbicular-obovate,
10 to 12 mm. long, 10 mm. wide, acute, apiculate, narrowed below, gla-
brous, the margins very minutely denticulate, the leaves of the branches
narrowly lanceolate, acuminate, apiculate, 6 to 8 mm. long, about 2
mm. wide, hyaline margined, the margins ciliate. Branches with one
or two flowers. Calyx including the teeth about 6 mm. long, the teeth
narrowly lanceolate, acuminate, apiculate, the margins ciliate, about
3 mm. long. Corolla tubular 7 to 8 mm. long, the teeth 5, narrowly
ovate, blunt, their margins incurved, 2 mim. long or less, the tube plaited
between the teeth, the teeth of the folds less than 1 mm. long, narrowly
ovate, entire, acute. Filaments about 1.5 mm. long; anthers 1 mm. long.
Capsule stalked, slightly exserted, the stalk broad, about 5 mm. long, the
capsule about 4 mm. long, orbicular-ovate, margined, compressed, obtuse.
Seed subglobose to ovoid, brown, minute, about 0.2 mm. in diameter.

Luzon, Province of Benguet, Kabayan (4443 Merrill) October 27, 1905. On
damp, rocky banks, in open lands. Rave, altitude about 1,300 m.

Gentiana luzoniensis Merrill sp. nov. § Chondrophylla.

An erect simple or slightly branched annual 3 cm. high or less, the
leaves similar, not imbricated, the stem and branches glandular. Leaves
sessile, elliptical-ovate, acute or obtuse, glabrous, 6 mm. long or less, 3.5
mm. wide or less, the nerves few, obscure. Flowers white, solitary at the
ends of the branches. Calyx about 4.5 mm. long including the teeth, the
teeth lanceolate, acuminate, about 1.5 mm. long. Corolla 8 mm. long,
tubular-campanulate, 5-lobed, the lobes ovate, acuminate, 1.5 to 2 mm.
long, the tube plaited between the lobes, the teeth of the folds ovate, often
slightly toothed, 1 mm. long or less. Filaments 4 mm. long; anthers
0.8 mm. long. Stigmas 1 to 1.5 mm. long, recurved. Capsule slightly
exserted, stalked, narrowly obovoid, compressed, margined, about 5 mm.
long. Seeds narrowly ovoid, acute, minutely reticulate, about 0.5 mm.
long.

Luzon, District of Lepanto, Mount Data (4558 Merrill) November 4, 1905.
On dry, open, grassy slopes in thin pine forests at about 2,100 m., rare.

225
GESNERIACE.
CYRTANDRA Forst.

Cyrtandra villosissima Merrill sp. noy.

A shrub very densely ferruginous villous throughout with long soft
hairs, the pubescence felted on the younger branches, inflorescence,
petioles and under surface of the leaves. Branches brown, terete, the
older ones less densely villous than the ultimate branches. Leaves thick,
oblong-ovate to ovate-lanceolate, 11 to 20 cm. long, 3.5 to 8 em. wide,
narrowed above to the rather slender acuminate apex, and below to the
acute or acuminate strongly inequilateral base, densely softly ferruginous
villous throughout, paler beneath, the upper surface less densely pubes-
cent than the lower, the margins dentate above, densely ciliate villous ;
nerves 12 to 14 on each side of the midrib; petioles densely felted villous,
4 em. long or less. Cymes axillary, several flowered, about 3 cm. long,
the bracts linear, 1 cm. long or less, densely felted villous lke the rest
of the inflorescence. Calyx lobes linear, 1.5 cm. long, 1 mm. wide, per-
sistent, densely villous. Corolla about 18 mm. long, somewhat hirsute
outside, tubular, subequally 5-lobed, the lobes orbicular-ovate, about 4
mm. long, pilose outside. Stamens 2, inserted at about the middle of the
tube, included; filaments about 2 mm. long; anthers 2.6 mm. Ovary
oblong, densely pilose; style pilose, included, 2-cleft; stigmas broadly
elliptical-ovate. Disk cup-shaped, glabrous, 1 mm. long. Capsule ob-
long, about 1 em. long, 4.5 mm. thick, hirsute. Seeds indefinite, oblong,

about 0.38 mm. long, glabrous, brown.

Mrypanao, Lake Lanao, Camp Keithley (51 Mrs. Clemens) January, 1906.
A very characteristic species, recognizable by its very dense, felted, ferruginous
soft, villous pubescence, perhaps most closely related to Cyrtandra mollis DeVriese.

MONOPHYLLEA R. Br.

Monophyllea lowei C. B. Clarke in DC. Monog. Phan. 5 (1883) 183.

Mrnpanao, District of Zamboanga, San Ramon (1565 Copeland) December 11,
1904. In rocky gorges in forests at about 160 m.

No species of this Malayan genus has previously been reported from the
Philippines, the above specimens differing from Monophyllea lowei in its some-
what smaller leaves, but I am unable to separate it by other characters, the
original description of Clarke’s species being rather short. Borneo.

PARABOEA Ridl.

Paraboea luzoniensis Merrill sp. nov.

Stems stout, 5 to 7 em. long, somewhat floccose, gray. Leaves many,
crowded, long petioled, elliptical-ovate to elliptical-obovate, membranous,
7 to 11 em. long, 4 to 7 em. wide, subentire, the apex broad, rounded,
rarely somewhat acute, the base acute or subtruncate, often oblique, the
upper surface glabrous or nearly so, the lower surface paler, more or less
densely floccose pubescent; nerves 6 to 8 on each side of the midrib,

226

ascending; petioles 2 to 7 em. long, somewhat pubescent. Cymes many,
axillary, long peduncled, lax, glabrous, the peduncles 12 em. long or less,
slender, dichotomously branched above, the bracts and bracteoles linear
to lanceolate, 5 mm. long or less, the pedicels slender, 1 cm. long or less.
Flowers pale blue, 1.5 cm. in diameter. Calyx 5-cleft nearly to the base,
the lobes linear, 3 to 3.5 mm. long, less than 1 mm. wide, glabrous.
Corolla 5-lobed, the lobes ovate, rounded, about + mm. long, the tube
short, broad. Stamens 2; filaments + mm. long; anthers 2 mm. long,
much broader than long. Ovary glabrous; style subcapitate. Capsule
glabrous, linear, 2 to 2.5 em. long, dehiscing into 4 straight valves.
Seeds numerous, narrow, acute, brown, 0.4 mm. long.

Luzon, Provinee of Cavite, Maragondong (4178 Merrill) July 30, 1905.
Abundant locally on cliffs in ravines 100 to 400 m.

| have followed Ridley*® in retaining Paraboea as a genus distinet from
Didymocarpus Clarke (Roettlera Vahl.).

TRICHOSPORUM Don.

Trichosporum littorale Merrill sp. nov.

A scandent epiphyte, glabrous except the slightly pubescent branches
and inflorescence, with opposite, orbicular-ovate to elliptical-ovate leaves,
3 em. long or less, and terminal flowers about 5 cm. long. Branches
gray, slender, terete, somewhat pubescent. Leaves coriaceous, 2 to 3 em.
long, 1.5 to 2.5 em. wide, entire, the base broad, rounded or slightly
cordate, the apex acute or obtuse; nerves obsolete; petioles slightly
pubescent, about 2 mm. long. Flowers purplish, several at the end of
each branchlet, the pedicels 1.5 to 2 cm. long, slightly pubescent. Calyx
nearly 3 cm. long, cylindrical, somewhat inflated, glabrous or slightly
ciliate at the apex, 5-toothed, the teeth ovate, acute or obtuse, about 38 mm.
long. Corolla about 5 cm. long, inflated at the base, then narrowed, and
inflated again above. Capsule unknown.

MINDANAO, District of Davao, Malita (647 Copeland) March 26, 1904. Grow-
ing on tree trunks over the beach along the seashore, characterized by its short,
broad, somewhat cordate leaves and relatively large calyx.

Trichosporum ovatum Merrill sp. noy.

A scandent plant, apparently an epiphyte, with opposite, glabrous,
fleshy, coriaceous, ovate, acute leaves 3 em. long or less, the flowers fas-
cicled at the apices of the branches, 3 to 3.5 cm. long. Branches slender,
gray, somewhat pubescent with weak scattered hairs, emitting rootlets.
Leaves opposite, 1.5 to 3 ecm. long, 1.5 to 2 em. wide, very thick, fleshy
when fresh, acute, the base broad, rounded, entire, the nerves obsolete ;
petioles about 2 mm. long, slightly ciliate. Flowers fascicled, 5 or 6 at
the end of each branchlet, the pedicels, calyx, and corolla uniformly hir-
sute outside with scattered hairs, the pedicels slender, about 7 mm. long.
Calyx glabrous inside, cylindrical, 1 cm. long, subequally 5-toothed,

Journ. Straits Branch R. A. Soe. (1905), 43, 63.

227
the teeth short, broad, obtuse, 1.5 mm. long or less. Corolla appar-
ently purple, tubular, curved, slightly inflated above, contracted some-
what at the throat, about 3 em. long, the lobes ovate, rounded, 5 to
6 mm. long. Stamens 4, slightly exserted, the anthers 2.5 mm. long.
Ovary glabrous; style slightly pubescent below the stigma. Fruit un-
known.

Minpanao, Lake Lanao, Camp Keithley (316 Mrs. Clemens) February, 1906.
Altitude about 800 m. A species probably of the section Holocalyx,
Trichosporum copelandi Merrill sp. noy.

A scandent epiphyte, glabrous except the inflorescence, with opposite,
oblong leaves 5 to 7 em. in length, and axillary and terminal inflorescence,
the flowers quite the same as in Trichosporum ovatum. Branches
slender, gray, glabrous. Leaves coriaceous, 1.5 to 2.5 cm. long, the base
rounded or obtuse, the apex acute, pale, somewhat shining, the nerves
obsolete ; petioles rugose, 4 to 5 mm. long. Peduncles axillary and ter-
minal, solitary or fascicled, somewhat pilose, two or more flowered.
Flowers purple, in size, shape, and pubescence quite the same as in
Trichosporum ovatum. Capsules linear, 10 cm. long. Seeds wanting.

MINDANAO, District of Zamboanga, San Ramon (1619 Copeland) February 5,
1905. An epiphytic vine in forests along the river at about 200 m. A species
apparently closely related to the preceding, similar in floral characters, but
differing in its much larger, very differently shaped leaves, longer petioles, and
glabrous branches, the inflorescence being also lateral as well as terminal.
Trichosporum rubrum Merrill sp. noy.

A scandent epiphyte reaching a length of from 4 to 6 m., rather
strongly hirsute pilose with pale often appressed hairs, the leaves lan-
ceolate to oblanceolate, the flowers red. Branches gray, the older ones
glabrous, the younger branchlets densely appressed hirsute-pilose. Leaves
subcoriaceous, 4 to 8 cm. long, 1.5 to 2 em. wide, pale beneath, the upper
surface rather densely appressed hirsute-pilose, the lower surface densely
so on the midrib and nerves, gradually narrowed below to the slender
acute base, more abruptly narrowed above to the acute apex, the margins
entire, hirsute; nerves about 4 on each side of the midrib, ascending ;
petioles 1 to 4 cm. long, densely appressed hirsute. Flowers fascicled in
the tipper axils, the pedicels hirsute, 2 cm. long or less. Calyx hirsute,
deeply 5-cleft nearly to the base, the lobes linear, about 6 mm. long,
1.5 mm. wide, blunt. Corolla nearly 3 em. long, bright red, tubular,
slightly curved, densely uniformly hirsute pilose with pale 2 mm. long
hairs, the lobes elliptical-ovate, rounded, 4 to 5 mm. long. Filaments
glabrous; anthers broad, 2 mm. long. Disk truncate or obscurely toothed,
glabrous, cup-shaped, 2 mm. long. Ovary glabrous. Old capsules pale,
elabrous, linear, 8 cm. long, the seeds not seen. :

Luzon, District of Lepanto, Mount Data (4581 Merrill) November 4, 1905.
On mossy tree trunks in ravines at about 2,200 m.

228
HYDROPHYLLACEE.
ELLISIOPHYLLUM Maxim.

Ellisiophyllum reptans Maxim. Bull. Acad. Pétersb. 16 (1871) 223.

Luzon, Province of Benguet, Pauai (4738 Merrill) November 8, 1905. In the
mossy forest at about 2,200 m. Japan.

This monotypic genus has previously been known only from Japan, and the
discovery of the species in the Philippines is of special interest, especially in
connection with the now known, strong, northern floristic element in the highlands
of Northern Luzon. The identification has been made from the generic deserip-
tions in Bentham and Hooker, Genera Plantarum, and in Engler und Prantl,
Natiirlichen Pflanzenfamilien, as the original description of the genus and species
is not available in Manila.

BORRAGINACEH.
BOTHRIOSPERMUM Bunge.

Bothriospermum tenellum (Hornem.) Fisch. & Mey. Ind. Sem. (1835) 24;
Clarke in Hook. f. Fl. Brit. Ind. 4 (1883) 167. Anchusia tenella Hornem.
Hort. Hain. 1 (1813-1815) 176.

Luzon, Province of Benguet, Baguio (5774 Hlmer) March, 1904. In waste
places about old rice lands. An interesting addition to our knowledge of the
northern element in the Philippine flora, Northern India to Manchuria, China,

Formosa, and Japan.
TRIGONOTIS Stev.

Trigonotis philippinensis Merrill sp. noy.

An erect or somewhat spreading few branched herb 25 em. high or
less, with oblong-elliptical short acuminate or apiculate leaves, white
flowers in terminal ebracteolate racemes, and glabrous tetrahedral shining
nutlets. Branches more or less appressed hispid pubescent. Leaves
coriaceous, 1 to 1.8 cm. long, 8 mm. wide or less, appressed hispid
pubescent on both surfaces with scattered white hairs, the petioles 1 em.
long or less, the nerves obsolete. Racemes terminal, solitary, about 10
cm. long, appressed hispid pubescent, ebracteolate, the pedicels short,
elongated to 5 or 6 mm. in fruit. Calyx nearly 3 mm. long, 5-cleft, the
lobes narrowly ovate, acute, hispid, nearly 2 mm. long. Corolla white,
4 mm. long, the tube nearly 2 mm. long, the 5 lobes spreading, broadly
obovate, rounded, the throat with 5 rather broad scales. Stamens 5,
included, the filaments very short, the anthers oblong, about 0.8 mm.
long. Ovary glabrous; style simple, slender, about 1 mm. long. Carpels
|, with 4 acute edges, black, glabrous, shining, about 1 mm. long, the sear
small, basal.

Luzon, Province of Benguet, Pauai to Baguio (4700 Merrill) November 9, 1905.
Growing on damp mossy cliffs just below the limits of the mossy forest at an
altitude of about 1,900 m., rare.

An interesting addition to our knowledge of the northern element in the Phil-
ippine flora, no species of the genus having previously been reported from the
Philippines, all which had been known being confined to the Asiatic continent,

Japan, and the Luehu Archipelago,

VERBENACEA®.
PREMNA Linn.

In F.-Villar’s Novissima Appendix to the third edition of Blanco’s
Flora de Filipinas, 11 species of Premna are enumerated. Of these
eleven species, but two are here enumerated under the same names ag those
used by F.-Villar, Premna adenosticta Schauer and P. cumingiana
Schauer. From synonomy and other sources I have been able satis-
factorily to reduce five species, P. foetida F.-Vill., non Reinw.; P. pubes-
cens F'.-Vill., non Blume; P. vestita Schauer; P. tomentosa F.-Vill., non
Willd., and P. mucronata ¥.-Vill., non Roxb., while judging from the
native names cited the species enumerated as Premna viburnoides Wall., is
P. nauseosa Blanco. Premna philippinensis Turez, is a species of Vitex,
V. turczaninowii Merr.** Premna gaudichaudii Schauer is enumerated
by F.-Villar, but is not credited to the Philippines, this with P. marian-
narum, the remaining species of Villar’s list, should be excluded from
the Philippine flora, unless at some future time material is collected that
agrees with the descriptions of these species.

1. Calyx equally or subequally 4-toothed.
2.A tree; leaves long acuminate, pale; corymbs densely stel-
avg el SO MD CS COI ps oe acon ee ee Se se ee (1) P. adenosticta
2.Scandent shrubs; leaves acute or short acuminate, dark
when dry; corymbs glabrous or pubescent, never stellate
pubescent.
3. Leaves broadly elliptical-ovate, 10 to 20 cm. long;
inflorescence 12 to 14 em. in diameter, crisped
DUDES COM ieee. te estee sete coronene ee ee Ag) ee ot (2) P.membranacea
3. Leaves oblong, 5 to 9 em. long; inflorescence 4 to 6
cm. in diameter, ferruginous pubescent, the pubes-
Cenee mot stellate Om Crisp edges se eaten ene eee (3) P. oblongifolia
1. Calyx subequally 5-toothed, or bilabiate, one lip entire and the
other 2-toothed, or one lip 2-toothed and the other 3-toothed.
2. Leayes ample, rufous or ferruginous stellate pubescent... (4) P. cumingiana
2. Leaves glabrous or pubescent, never stellate pubescent.
3. Scandent ; leaves pubescent beneath .__. ERAS os Senn tP ones op a: 2 (5) P. subscandens
3. Erect trees or shrubs.
4. Leaves uniformly and usually densely pubescent

beneath.
5. Leaves 6 cm. long or less; petioles 5 mm.
longs OT: 6Ss.2).b eee eee eee kek Wee (6) P. depauperata
5. Leaves 10 to 20 em. long; petioles 2 to 6
CYL ODS: » os: a5 eee eee ee eee a Prrcra Ocet ee ds (7) P. odorata

4. Leaves glabrous, or if pubescent only so. on the
nerves and midrib.
5. Leaves 6 to 9 cm. long.
6. Leaves coriaceous ; inflorescence dense (8) P. congesta
6. Leaves membranous; inflorescence
rather ‘lax: Mice See ees oe ee Se (9) P. nauseosa
5. Leaves 9 to 18 em. long; inflorescence more
or less lax.
6. Leaves usually oblong or oblong-
ovate, acute or obtuse; nerves 4 to
5, glabrous, or at least pubescent
Only inthe vaxils beneatiheercs-- (10) P. integrifolia

* Govt. Lab. Publ. (1906), 35, 77.

250

6. Leaves ovate, acuminate; nerves 5 to
6, pubescent on both surfaces; co-
Olay SDA LON See ee eee (11) P. nitida

6. Leaves usually broadly ovate, acum-
inate; nerves 6 to 7, glabrous, or
pubescent on the upper surface or
on both; corolla 5.5 to 6 mm. long..(12) P. subglabra

(1) Premna adenosticta Schauer in DC. Prodr. 11 (1847) 630; Mig. Fl. Ind.
Bat. 2 (1856) 892; F.-Vill. Nov. App. (1883) 159; Vid. Phan. Cuming:
Philip. (1885) 134; Rev. Pl. Vase. Filip. (1886) 209; Ceron Cat. Pl. Herb.
(1892) 131.

PHILIPPINES (1230 Cuming) 1836-1840, cotype. Luzon, Province of Rizal.
Bosoboso (2863 Ahern’s collector) March, 1905; San Mateo (1122 Ahern’s col-
lector) May, 1904; Antipolo (435 Ahern’s collector) February, 1904; Province of
Camarines, Pasacao (103 Ahern) Mareh, 1902. Endemic.

(2) Premna membranacea Merr. -Govt. Lab. Publ. 35 (1906) 65.
Luzon, Province of Rizal, Bosoboso (1165, 3102 Ahern’s collector) June, 1904,
June, 1905. T., Alamag. Endemic.

(3) Premna oblongifolia Merr. 1. ¢. 29 (1905) 48.
Luzon, Province of Benguet, Baguio (5990 Elmer) March, 1904; Province of
Tayabas (Infanta) (816 Whitford) September, 1904. Hndemice.

(4) Premna cumingiana Schauer in DC. Prodr. 11 (1847) 634; Vidal, Cat.
Pl. Prov. Manila (1880) 39; Rev. Pl. Vase. Filip. (1886) 209; Phan.
Cuming. Philip. (1885) 134; F.-Vill. Nov. App. (1883) 159; Miq. Fl. Ind.
Bat. 2 (1856) 778; Koorders Meded. ’s Lands Plantent. 19 (1898) 559.
Premna cordata Blanco, Fl. Filip. ed. 1 (1837) 489, non R. Br. Premna
tomentosa Blanco, |. ¢., ed. 2 (1845) 342; ed. 3; 2 (1878) 269; F.-Vill.
Noy. App. (1883) 159, non Willd. Premna cardiophylla Schauer, 1. ¢. 638;
Miq. Fl. Ind. Bat. 2 (1856) 900.

PHILIPPINES (778, 1280 Cuming) 1836-1840, cotypes. Luzon, Province of
Rizal, San Mateo (1850 Ahern’s collector) September, 1904; (112 Merrill)
Decades Philip. Forest. FI., coll. Ahern’s collector, May, 1904, distributed as
Geunsia cumingiana; Bosoboso (2676 Merrill) June. 1903; (3133 Ahern’s col-
lector) June, 1905; Provinee of Camarines, Pasacao (803 Ahern) 1902. MuInN-
DANAO, District of Davao (689 Ahern) June, 1901; (109°DeVore & Hoover) Apvril,
1903.

A species known only from the Philippines and Celebes. Premna cordata
Blanco, non R. Br.==Premna tomentosa Blanco, non Willd., although very imper-
fectly described by Blanco, is certainly referable to Schauer’s species. Blanco’s
short description applies well to the specimens above cited, especially his expres-
sion “El involucro universal de la umbela, el de la parcial y el de la florecita, dos
hojuelas alesnadas,” by “involucro” Blanco meaning the bracts and _ bracteoles.
Premna cardiophylla Schauer, is only a new name for Blanco’s Premna cordata,
and is included by the former in his monograph under doubtful or not sufliciently
known species. ‘T., Maguilac; according to Blanco, Malaapi.

(5) Premna subscandens Merrill sp. noy.

A scandent shrub, the leaves, branchlets, and inflorescence more or
less pubescent, with subequally 5-toothed or obscurely 2-lipped calyces,
the corollas 4-lobed. Branches light gray or brown, glabrous, obscurely
angled, the lenticels few, the younger branchlets pubescent, sometimes

densely so. Leaves ovate, membranous, 6 to 10 em. long, 3 to 7 em.

Zen

wide, entire or distantly obscurely toothed above, acuminate, the base
broad, rounded or cordate, sometimes acute, thinly pubescent on both
surfaces, more densely so beneath, often densely pubescent on the neryes
and midrib, except the latter sometimes becoming glabrous or nearly so,
shining, usually dark colored when dry; nerves 5 on each side of. the
midrib, somewhat prominent, the reticulations somewhat lax: petioles 1
to 4 em. Jong, pubescent. Cymes terminating the branches and short
lateral branchlets, many flowered, somewhat dense, 5 to 14 cm. across,
all parts except the corollas uniformly densely softly grayish brown
pubescent, the linear bracts 5 to 7 mm. long, densely pubescent, the
bracteoles similar but much smaller. Flowers white or greenish,
fragrant. Calyx densely softly pubescent, cup-shaped, 3 mm. long,
5-toothed, the teeth subequal, rounded or acute, 1 mm. long or less, or
sometimes obscurely 2-lipped, one lip 2-toothed, the other 3-toothed.
Corolla 5 mm. long, glabrous outside, villous within, 2 lipped, one lip
ovate, entire, nearly 2 mm. long, the other coarsely 3-lobed, the lobes
nearly equaling the entire lip. Stamens 4, didynamous; filaments 4 to 5
mm. long, villous below; anthers about 0.4 mm. long. Ovary glabrous ;
stvle 5.5 mm. long.

Luzon, Province of Rizal, Antipolo (449, 443 Ahern’s collector) April, 1904;
Bosoboso (1874 Merrill) April, 1903. Ticao (1095 Clark) May, 1904. T., Alag-
baguin, Alagao-baguin. V., Uradqao.

(6) Premna depauperata Merrill sp. noy.

An undershrub, 1 m. high or less, with short petioled, coriaceous
oblong-ovate to elliptical-ovate, small leaves, densely ferruginous pubes-
cent inflorescence and 5-toothed calyx. Branches terete, light brown,
mostly densely ferruginous pubescent, lenticellate. Leaves 3 to 6 cm.
long, 2 to 3.5 cm. wide, shining above, entire, the margins recurved, acute,
the base broad, rounded-cordate, the upper surface sparingly strigose
pubescent, the lower surface somewhat densely pubescent, glandular;
nerves 4 to 5 on each side of the midrib, distinct beneath, anastomosing ;
petioles densely pubescent, 5 mm. long or less. Corymbs terminal, 4
to 5 em. long, nearly as wide, densely ferruginous pubescent throughout,
densely flowered, the lower bracts small, foliaceous, the upper and brac-
teoles linear. Calyx cup-shaped, nearly 3 mm. long, sparingly strigose
pubescent, subequally 5-toothed, or obscurely 2-lipped with one 2-toothed
and one 3-toothed lip. Corolla 4 mm. long, pubescent outside, villous
on the throat within, 4-lobed, one lobe lip-like, exceeding the others.
Stamens not exceeding the corolla lobes; anthers 0.5 mm. long; filaments
glabrous. Ovary glabrous, ovoid or globose; style 3.5 mm. long. Fruit
glabrous, ovoid, about 4 mm. long.

CULION (603 Merrill) December, 1902. On dry open grassy hillsides 30 to
40 m. above the sea. A species well characterized by its small size, small leaves,
which are cordate at the base, and short petioles.

232

(7) Premna odorata Blanco FI. Filip. ed. 1 (1837) 489; ed. 2 (1845) 341; ed.
3, 2 (1878) 268; Schauer in DC. Prodr.’11 (1847) 638; Mig. Fl. Ind. Bat. 2
(1856) 900; Merr. Govt. Lab. Publ. 27 (1905) 68; 1. ce. 35 (1906) 76.
Premna vestita Schauer, 1. c. 631; Mig. 1. c. 892; F.-Vill. Nov. App. (1883)
159; Vidal, Phan. Cuming. Philip. (1885) 134; Rev. Pl. Vase. Filip. (1886)
209; Sinopsis, Atlas (1883) ¢. 74. f. H.; Merr. Forest. Bur. Bull. 1 (1903) 51.
Premna serratifolia Blanco 1. ¢. ed. 2 (1845) 269; ed. 3,1. ¢. Premna tomen-
tosa F.-Vill. 1. ec. ex syn. Blanco, non Blume. Premna foetida F.-Vill., 1. ¢.
non Reinw., ex syn. Blanco.

PHILIPPINES (599 Cuming) 1836-1840, cotype of Premna vestita Schauer.
Luzon, Manila (22, 3421 Merrill) April, 1902; November, 1903; (718 Ahern)
April, 1901: Province of Bataan, Dinalupihan (1514 Merrill) January, 1903;
Lamao River (2590 Meyer) February, 1905; Mariveles (748, 780 Ahern) Jan-
uary, 1902: Province of Pampanga (37 Parker) May, 1904: Province of Rizal,
Antipolo (24 Merrill) Decades Philip. Forest FI., coll. Ahern’s collector, February,
1904: Province of Tayabas, Lucena (2892 Merrill) June, 1903: Province of
Union, Bauang (5561 Elmer) February, 1904: Province of Benguet, Sablan (6154
Hlmer) April, 1904. Minporo, Baco (1218 Merrill) January, 1903. GUIMARAS
(235 Gammill) January, 1904.

Premna vestita is placed by Schauer in the section with 4-toothed calyces, but
in two specimens of Cuming’s No. 599 in our herbarium, on which number the
species was based, the calyces are distinctly 5-toothed. This is certainly the
most abundant and widely distributed species of the genus in the Philippines,
and is the one utilized by the natives as noted by Blanco. Blaneco’s deserip-
tion applies very closely, and I am of the opinion that there can be no doubt
as to the correctness of the identification. Premna serratifolia Blanco is reduced
to P. odorata, as Blanco states that the species can be distinguished from the
latter only by the serrate leaves. Nos. 1218 and 3421 Merrill and No. 37 Parker
show this character, but no other characters on which I am able to separate
this form as a distinct species. As a result of the reduction of Premna_ ser-
ratifolia to P. odorata, I have also so reduced P. foetida F.-Vill., to which F.-Villar
referred P. serratifolia Blanco. For the same reason F.-Villar’s Premna pubescens
is reduced, he having referred Premna odorata Blanco to Blume’s species. From
F.-Villar’s remark, however, it is possible that he referred specimens of Premna
cumingiana to Premna pubescens. Abundant and widely distributed in the Phil-
ippines. T., Alagao. V., Adgao.

(8) Premna congesta Merrill sp. noy.

A shub or small tree with small ovate to elliptical-ovate nearly glabrous
leaves 2.5 to 7 em. long, and densely flowered terminal cymes 3 to 5
em. in diameter, the calyx 2-lipped, the corolla 4-lobed. Branches terete
lenticellate, ight gray or brownish, becoming quite glabrous, the younger
parts with few scattered, weak hairs. Leaves subcoriaceous, 1.5 to 4.5
cm. wide, shining and glabrous above, or the nerves somewhat pubescent,
paler beneath and minutely punctulate, glabrous, or the axils of the
nerves barbellate, and sometimes the midrib somewhat pubescent, entire
or somewhat crenate above, acute or obtuse, rarely very short acuminate,
the base rounded; nerves about 4 on each side of the midrib, somewhat
prominent; petioles slender, 1 cm. long or less, somewhat pubescent.
Cymes pubescent with few weak, scattered hairs, the branches spreading
or ascending, the bracts and bracteoles linear to lanceolate, 1.5 to 3 mm.

235

long, somewhat pubescent, the short pediceled flowers subtended by about
3 linear bracteoles. Calyx glabrous, cup-shaped, 2 mm. long, 2-lipped,
one lip coarsely 2-toothed, the other truncate, subentire or minutely 2
to 3 toothed. Corolla 4 to 4.5 mm. long, tubular, 4-lobed, three lobes
equal, ovate, about 1.5 mm. long, obtuse, the fourth somewhat larger,
glabrous outside, villous within. Stamens 4, didynamous ; anthers about
0.5 mm. long. Ovary glabrous; style slender, nearly 4 mm. long.

Luzon, Province of Rizal, Bosoboso (3281 Ahern’s collector) August 6, 1905.
T., Alacaas. According to the native collector, the flowers are reddish in color.

(9) Premna nauseosa Blanco Fl. Filip. ed. 1 (1837) 489; Schauer in DC.
Prodr. 11 (1847) 638; Mig. Fl. Ind. Bat. 2 (1856) 900; Vidal, Phan.
Cuming. Philip. (1885) 134; Rev. Pl. Vase. Filip. (1886) 209; Merrill,
Govt. Lab. Publ. 27 (1905) 68. Premna mucronata F.-Vill. Nov. App.
(1883) 159; Schauer, 1. e. 635, in- part; Mig. FI. Ind. Bat. 2 (1856) 897,
in part, with reference to No. 1367 Cuming. Premna leucostoma Naves, FI.
Filip. ed. 3, pl. 346, non Miq.

PumiprInEs (1367 Cuming) 1836-1840, in Herb. Bureau of Science. LUZON,
Province of Zambales, Subie (2191, 2915 Merrill) May, 1903: Province of Bataan,
Mount Mariveles (387 Whitford) June; (6841 Elmer) November ; (769, 1271,
1275, 1613 Borden) May to August, 1904: Province of Tarlac (705 Ahern) May,
1901: Province of Rizal, Antipolo (24 Guerrero) June, 1903; (1682 Merrill)
March, 1903.

A species widely distributed in Luzon, usually somewhat pubescent, in which
character Blanco’s short and imperfect description does not apply. It is, however,
the only species at present known to me that has a somewhat disagreeable odor,
the character on which Blanco’s specific name was based. This species is
enumerated by Schauer as doubtful, or not sufticiently known. According to
Vidal, Nos. 693 and 1353 Cuming also represent it, and it is possible that
No. 1451 Cuming is also the same, a fragment of the latter number existing in
our herbarium. From the native names cited, Premna viburnoides Vidal, Cat.
Pl. Prov. Manila (1880) 134; F.-Vill. Nov. App. (1883) 159, non Wall., should
be referred to this species, Premna nauseosa being quite universally known to
the Tagalogs as Molauain aso, literally “dog molave,” Molauain or Molave being
the Tagalog name for the valuable timber trees of the genus Vitex, the timber of
Premna nauseosa being considerably utilized for certain constructions, and some-
what resembling Molave.

(10) Premna integrifolia Linn. Mant. (1767) 252; Clarke in Hook. f. Fl. Brit.
Ind. 4 (1885) 574; Forbes & Hemsl. Journ. Linn. Soe. Bot. 26 (1890) 255;
Schum. und Lauterb. Fl. Deutsch. Schutz. Siidsee (1901) 523.

Luzon, Province of Camarines, Pasacao (141, 138, 801 Ahern) 1902: Province
of Tayabas (Infanta) (760 Whitford) September, 1904: Province of Zambales,
Subic (2199 Merrill) May, 1903: Province of Principe, Baler (1083 Merrill)
October, 1902: Province of Bataan, Lamao River (2043 Borden) October, 1904.
Mrnporo, Calapan (899 Merrill) April, 1903; Baco (1173 Merrill) January.
Apo Island, Mindoro Strait (427 Merrill) December, 1902. Mrnpanao, District
of Davao (350 Copeland) March, 1904; (138, 226 DeVore & Hoover) April, 1903.

A species apparently confined to the seashore or near it, widely distributed
in the Philippines. Tropical Asia to Malaya, New Guinea and Polynesia. T.,
Alagao, Alagao dagat. (Dagat in Tagalog ocean ) .

44772—_5

234

(11) Premna nitida K. Sch. Fl. Kaiser Wilhelms Land (1889) 120; Sch. und
Lauterb. Fl. Deutsch Schutz. Siidsee (1901) 523; Usteri, Beitr. Kennt.
Philip. Veg. (1905) 123.

This species is reported from Negros and Panay by Usteri, but I have seen
no Philippine specimens to which Schumann’s description applies. The species
is known from New Guinea and doubtfully from Celebes.

(12) Premna subglabra Merrill sp. noy.

A tree 7 to 12 m. high, becoming nearly glabrous, the slightly pubescent
calyces obscurely 2-lipped, one lip entire or very obscurely 2 to 3 toothed,
the other 2-toothed. Branches hght gray or brown, glabrous, terete, the
branchlets reddish brown, obscurely angled, deciduously short-pubescent,
becoming glabrous or nearly so, Leaves ovate to oblong-ovate, char-
taceous, entire, short acuminate, the base broad, rounded, subtruncate to
somewhat cordate, or sometimes acute, shining, glabrous above except
the somewhat pubescent midrib and nerves, paler beneath, obscurely
minutely glandular punctate, glabrous, or the nerves and midrib very
slightly pubescent, the axils sometimes barbellate; nerves prominent

n

beneath, ascending, 6 to 7 on each side of the midrib, the reticulations
distinct ; petioles 2 to 7 em. long, usually pubescent on the upper surface,
otherwise glabrous. Corymbs terminal, spreading, many flowered, 9 to
14 cm. long, 10 to 20 em. wide, more or less ferruginous pubescent
throughout, the ultimate branchlets rather densely so, the bracts linear,
5 to 8 mm. long, the bracteoles similar but much smaller. Flowers
greenish. Calyx slightly pubescent, becoming glabrous, cup-shaped, 2
mm. long. Corolla 5.5 to 6 mm. long, glabrous outside, villous within,
2-lipped, one lip short, entire or retuse, the other 3-lobed, the middle
lobe much exceeding the lateral ones. ‘Stamens slightly exceeding the
corolla lobes; filaments villous below; anthers about 0.6 mm. long.
Ovary glabrous; style 4 to 5 mm. long. Fruit globose, dark purple when
mature, glabrous, about 3 mm. in diameter.

Luzon, Province of Tayabas, Atimonan (670, 671 Whitford) August 19, 1904.
Mrinporo, Pinamalayan (2165 Merrill) May, 1903; Pola (2243 Merrill) May,
1903. In forests along streams below 100 m. T., Alagao.

LABIAT AL.
COLEUS Lour.

Coleus macranthus Merrill sp. nov. § Solenostemonoides.

An erect branched herb 1 to 2 m. high, more or less glandular fer-
ruginous puberulent, with long petioled, ovate to oblong-ovate, rather
strongly dentate leaves, narrow many flowered panicles, the flowers pure
white to purple, often 2 cm. long. Branches brown when dry, densely
glandular puberulent to nearly glabrous. Leaves membranous, 4 to 15
em. long, 3 to 7 em. wide, the base often broad and subtruncate, some-
times acute, decurrent, the apex short acuminate, the margins entire

235

near the base, above prominently and regularly dentate, dull, the nerves
and midrib on both surfaces usually puberulent, paler beneath and with
numerous small punctate glands; nerves about 10 on each side of the
midrib, prominent; petioles 2 to 7 cm. long. Panicles narrow, 15 to
25 cm. long, the branches verticillate, spreading, branched, glandular
puberulent, 2 em. long or less; bracts deciduous, ovate, acuminate, 7 to
8 mm. long. Calyx glabrous within, glandular puberulent outside, in
fruit about 10 mm. long, the upper lip broadly ovate, acute or slightly
acuminate, flat, about 4 mm. long, the lateral teeth ovate to elliptical,
rounded, about 2.5 mm. long, the lower lobe oblong, 7 mm. long, 2 toothed,
the teeth lanceolate or linear lanceolate, acuminate, about 1.5 mm. long.
Corolla 1.5 to 2 cm. long, slightly puberulent, the lower lip nearly 1 cm.
long, the upper short, 3-lobed. Nutlets ovoid, 1.7 mm. long, smooth,
glabrous, shining.

Luzon, District of Lepanto, Mount Data (4502, 4483, 4505 Merrill) November,
1905. In the mossy forest at about 2,200 m., abundant.

Var. crispipila, n. var.

Similar to the species, flowers white, the inflorescence puberulent and
with many crisped ferruginous hairs, the leaves also with few or many
crisped hairs on both surfaces and margins.

Luzon, Province of Benguet, Suyoe to Pauai (4780 IMerrill) November, 1905.
On high ridges in the mossy forest at about 2,200 m.

PLECTRANTHUS IJ/’Her.

Plectranthus diffusus Merrill sp. noy.

A much branched, diffuse, lax herb 1.5 to 2.5 m. high with many
flowered narrow lateral panicles forming large compound terminal leafy
panicles, more or less pubescent throughout with pale or ferruginous
hairs. Branches angled, gray or brownish, pubescent. Leaves ovate to
oblong-ovate, 2 to 6 em. long, 1 to 3 em. wide, membranous, acuminate,
the base acute, entire below, above rather prominently serrate-dentate,
with scattered crisped hairs on both surfaces; nerves, 5 to 6 on each side
of the midrib; petioles 1 cm. long or less. Branchlets of the in-
florescence opposite, several flowered, 1.5 cm. long or less, the bracts
foliaceous, gradually reduced upwards. Calyx somewhat hirsute, about
2 mm. long, subequally 5-toothed, the teeth acute, less than 0.5 mim.
long. Corolla deep purplish blue, 5 to 6 mm. long, straight, somewhat
pubescent outside, the limb gibbous, 2-lpped, the upper lip short, 3-lobed,
the lower one entire about 3 mm. long. Stamens free, filaments very
slightly pubescent below. Calyx in fruit scarcely accrescent, reflexed.
Nutlets ovoid or elliptical, glabrous, smooth, about 1 mm. long.

Luzon, District of Lepanto, Mount Data (4554 Merrill) November 4, 1905.
In damp shaded ravines along streams at about 2,000 m.

!

236

SOLANACEZ.
SOLANUM Linn.

Solanum inaequilaterale Merrill sp. noy.

An unarmed, more or less stellate pubescent shrub 2 to 3 m. high,
with alternate subentire or undulate leaves and extra-axillary cymes.
Branches light gray, glabrous, the younger ones black when dry, more or
less densely stellate pubescent. Leaves membranous or submembranous,
oblong-ovate, acute or acuminate, the base acute or obtuse, inequilateral,
the lamina on one side of the midrib extending somewhat down the
petiole, the margins entire or somewhat undulate, glabrous above, more
or less densely stellate pubescent beneath, the young leaves very densely
so, the older ones subglabrous; nerves 5 to 6 on each side of the midrib ;
petioles 2 to 3 cm. long. Cymes peduncled, stellate pubescent, densely
or laxly flowered, about 5 cm. long. Calyx densely stellate pubescent,
about 4 mm. long, subtruncate or obscurely 5-toothed. Corolla pale
purple, 11 mm. long, stellate pubescent outside, the tube 2 mm. long,
the 5 lobes oblong to oblong-lanceolate, acute, about 3 mm. wide.
Stamens 5, oblong, about 5 mm. long. Ovary 2-celled, glabrous or
slightly pubescent at the apex, the style glabrous, about 6 mm. long.
Fruit globose, glabrous, shining, bright red when mature, about 7 mm.
in diameter, the calyx not enlarged. Seed 2 to 2.5 mm. in diameter.
Inflorescence becoming nearly or quite glabrous in fruit.

Luzon, Province of Benguet, Suyoe to Pauai (4807 Merrill) November 7, 1905.
Border of the mossy forest on high ridges at about 2,000 m. No. 6204 Elmer,
from Sablan, the same province, appears to be a form of this species with thin,
less pubescent leaves and longer, lax cymes.

SCROPHULARIACE4.
ALECTRA Thunb.

Alectra dentata (Benth.) O. Kuntze Rev. Gen. Pl. (1891) 458. Hymenosper-
mum dentatwnm Benth. in Wall. Cat. 3963. Alectra indica Benth. in DC,
Prodr. 10 (1846) 339; Hook. f. Fl. Brit. Ind. 4 (1884) 297; Forbes &
Hemsl. Journ. Linn. Soc. Bot. 26 (1890) 20]. Melasma indica Wettst. in
Engl. und Prantl. Nat. Pflanzenfam. 4 (1891) 3B: 91.

Luzon, Province of Benguet. Baguio to Ambuklao (4350 Merrill) October 24,
1905. Dry open grass Jands in thin pine forests at about 1,500 m. Not previously
reported from the Philippines.

Southern China to the mountains of Burma, India, and Mauritius.

EUPHRASIA Linn.

Euphrasia borneensis Stapf, Trans. Linn. Soc. Bot. I]. 4 (1894) 210. pl. 16,
f. 1-16.

Luzon, Province of Benguet, Suyoe to Pauai (4720, 4722 Merrill) November 7,
1905. On dry open grassy ridges at about 2,200 m.

No species of the genus has been previously reported from the Philippines,
Kuphrasia borneensis having previously been known only from the type locality,
Mount Kanabalu, British North Borneo. The specimens cited above differ from
Euphrasia borneensis as described by Stapf in some slight minor characters, but

237

these I believe are scarcely sufficient to warrant the distinguishing of the Philippine
plant as a distinct species. Gaultheria borneensis Stapf, also originally described
from Mount Kinabalu specimens, and later reported from the Philippines by
Rendle,” was also found near Pauai (4796 Merrill) .

SOPUBIA Hamilt.

Sopubia trifida Hamilt. in D. Don Prodr. Fl. Nepaul. (1802) 88; Hook. f. Fl.
3rit. Ind. 4 (1884) 302; Benth. in DC. Prodr. 10 (1846) 522; Forbes &
Hemsl. Journ. Linn. Soe. Bot. 26 (1890) 202.

Luzon, Province of Benguet, Baguio (6518 Hlmer) June, 1904. On grassy
hillsides in thin pine forests at an altitude of about 1,500 m. Flowers yellowish,
the throat pink or purplish.

No representative of the genus has previously been reported from the Philip-
pines, the above species extending from southern China to British India and
Ceylon.

VANDELLIA Linn.

Vandellia grandiflora Merrill sp. nov.

An erect or spreading, simple or shghtly branched, more or less hirsute
herb 10 em. high or less, the calyx teeth 5, equal, less than half as long
as the calyx tube, the corolla about 12 mm. long. Branches, leaves.
petioles, pedicels, and calyces sparingly hirsute with scattered white hairs.
Leaves opposite, ovate to oblong-ovate, 0.5 to 1.5 em. long, short petioled,
acute, the base obtuse, the margins usually coarsely serrate. Flowers
solitary, pediceled, axillary, the pedicels 1.5 to 2 em. long. Calyx in
anthesis oblong, 6 mm. long, not keeled or winged, reticulate, equally
5-toothed, the teeth lanceolate, acuminate, about 2.5 mm. long. Corolla
pale blue or purplish, the tube cylindrical, enlarged above, upper lip
broad, about 4 mm. long, cleft at the apex, the lower lip 6 to 7 mm. long,
3-lobed, the middle lobe the largest. Stamens 4, the two posterior
included, the two anterior longer and prominently appendaged near the
base of the filaments. Ovary and style glabrous. Fruiting calyx about
8 mm. long, oblong, the teeth scarcely longer than in anthesis. Capsule
oblong, glabrous, tipped by the style, equaling or slightly shorter than the
calyx. Seeds many, flattened, ovate, about 0.5 mm. in diameter.

Luzon, District of Lepanto, Mount Data (4572 Merrill) November 4, 1905:
Province of Benguet, Baguio to Ambuklao (4359 Merrill) October 24, 1905;
Suyoe to Pauai (4735 Merrill) November 7, 1905. On dry open grassy slopes in
thin pine forests 1,600 to 2,000 m.

BIGNONIACEZ.
NYCTICALOS Teysm. et Binn.

Nycticalos cuspidatum (Blume) Miq. Ann. Mus. Bot. Lugd. Bat. 3 (1867)
249, t. 8. f. B. Tecoma cuspidata Blume, Rumphia 4 (1848) 35.
PALAWAN, Puerto Princesa (254 Bermejos) December, 1905. In forests; flowers
yellow. Celebes and the Moluceas.
An undetermined species of this genus has previously been reported from Luzon

* Journ. Bot. (1896) 34, 355.

238

by Ceron,** No. 3395 Vidal, but our specimen cited does not appear to be suf-
ficiently distinct from Miquel’s species. The Palawan specimens differ from
Nycticalos cuspidatum as described by Miquel in having shorter petioles, and
slightly larger calyces, while the leaves are acute, scarcely cuspidate.

RADERMACHERA UHassk.

Radermachera biternata Merrill sp. nov.

A small tree about 8 m. high, quite glabrous throughout, with bipin-
nate leaves, elliptical-ovate, usually obtuse leaflets and few flowered
panicles much shorter than the leaves, the flowers about 5.5 em. long.
Branches gray or brownish, the younger parts black when dry. Leaves
20 em. long or less, opposite, biternate; leaflets 5 to 9 cm. long, 2.5 to
5 em. wide, subcoriaceous, shining, the apex rounded obtuse or broadly
acute, the base acute, the margins revolute; primary nerves about 10 on
each side of the midrib, rather distinct beneath, the reticulations netted,
rather close; petiolules of the lateral leaflets 1.5 cm. long or less, of the
terminal ones about 3 cm. long. Inflorescence much reduced, the rhachis
3 cm. long or less, the branches very short or none. Flowers few. Calyx
about 1 em long, closed in bud, in anthesis unequally 3-lobed, the lobes
short, acute. Corolla 5 to 5.5 em. long, the tubular portion less than
1 em. long, about’ 3.5 mm. in diameter, enlarged-ventricose above, the
lobes about 1.5 em. long, rounded, entire, glabrous pink or pale purple.
Stamens about 2 em. long. Style 2.5 cm. long. Fruit unknown.

CuLion (568 Merrill) December 24, 1902. BuSUANGA (3491 Curran) Decem-
ber 31, 1905, both of these islands belonging to the Calamianes Group, between
Mindoro and Palawan.

A species growing in open grassy valleys slightly above the sea level well char-
acterized by its much reduced inflorescence and large flowers.

RUBIACEZE.
GALIUM Linn.

Galium philippinense Merrill sp. nov. G. ciliare Elm. Leaf. Philip. Bot. (1906)
4, non Hook.

A tufted erect or diffuse, much branched perennial, more or less ciliate-
pilose herb 10 to 30 em. high, the leaves in whorls of fours, 3-nerved
from the base, the cymes short, lateral, few flowered. Branches 4-angled,
more or less pilose when young, but not hispid, becoming nearly glabrous.
Leaves elliptical-ovate, sessile or nearly so, 5 to 8 mm. long, 2 to 4 mm.
wide, acute at both ends, more or less ciliate-pilose with long scattered
white hairs, 3-nerved. Cymes about 1 em. long, mostly 5-flowered, the
peduncles 1 em. long or less, glabrous or nearly so, the bracts foliaceous,
oblong-lanceolate, about 2 em. long. Flowers white, corolla rotate,
0.8 mm. long, 4-lobed, the lobes ovate, obtuse. Calyx tube ovoid. Fruit
about 1 mm. in diameter, glabrous or somewhat rugose when dry.

OCb. Pls Herds Ul 9A) Le ls

239

Luzon, Province of Benguet, Mount Santo Tomas (6557 Elmer) June, 1904:
District of Lepanto, Mount Data (4531 Merrill) November, 1905. On dry grassy
slopes in thin pine forest above 2,000 m. A second Philippine Species is repre-
sented by No. 6592 Elmer and No. 4414 Merrill from Benguet Province. From
the descriptions available, and without access to authentic material, T am unable
to separate these two numbers from the Australian Galiwm gaudichaudii DC.,
as determined by Mr. Elmer.

HEDYOTIS Linn.

Hedyotis microphylla Merrill sp. nov.

An erect glabrous shrub or undershrub 1 to 2 m. high, with small.
obseurely nerved, usually ovate-lanceolate leaves 1 to 3 em. long, ana
axillary and terminal peduncled, usually 3-flowered cymes, the capsules
2-celled, septicidal. Branches slender light gray, quadrangular. Leaves
submembranous dull or slightly shining, somewhat paler beneath, 5 to
8 mm. wide, the base acute, the apex acute or slightly acuminate, often
blunt; nerves very obscure, nearly obsolete, about 3 on each side of the
midrib; stipules short, 3-partite, the lobes narrow. Cymes 1 to 2 cm.
long, the bracts foliaceous often 3 mm. long, the pedicels about 5 mm.
long. Flowers white. Calyx 3 mm. long, the tube ovoid 1.5 mm. long,
the lobes 4, persistent, oblong ovate, obtuse, about 1.5 mm. long. Corolla
5 mm. long, glabrous outside, the throat and base of the lobes villous
inside, the tube broad, the lobes recurved, oblong ovate or ovate, acute,
about 2 mm. long. Filaments nearly 2 mm. long; anthers narrowly
oblong, 1.5 mm. Ovary 2-celled, each few ovuled; style 2.9 mm. long ;
stigma 2-cleft. Capsule elliptical-ovoid about 4 mm. long crowned by
the calyx lobes, 2-celled, each cell with 2 or 3 flattened elliptical seeds
about 1.5 mm. long, at length septicidally dehiscent from the apex.

Luzon, Province of Benguet, Pauai to Baguio (4693 Merrill) November, 1905;
Suyoe to Pauai (4736 Merrill) November, 1905. On high ridges in the mossy
forest 2,000 to 2,300 m.

PAVETTA Linn.

Pavetta dolichostyla Merrill sp. nov.

A small shrub about 2 m. high with membranous slightly pubescent
or nearly glabrous leaves and peduncled terminal cymes, the corolla tubes
3 cm. long, the exserted portion of the styles 4 to 4.5 em. long. Branches
greenish, glabrous, the stipules glabrous, narrowly ovate, 6 to 7 mm. long,
acuminate, deciduous. Leaves 20 to 25 em. long, 7 to 9 cm. wide, cilona
oblanceolate to broadly oblong-lanceolate, gradually narrowed below to
the acute or acuminate base, and above to the acuminate apex, shining
above, beneath paler and puberulent on the midrib and lateral nerves ;
nerves 12 to 14 on each side of the midrib, curved, anastomosing, the
reticulations lax; petioles 1 to 2.5 cm. long, pubescent: Cymes terminal,
pubescent, about 3 from each branchlet, few flowered, the peduncles 1.5
em. long, subtended by broad bracts, the upper bracts and bracteoles
linear to linear-lanceolate. Pedicels pubescent, 3 to 4 mm. long. Calyx

240

cup-shaped, about 2.5 mm. long, pubescent with short spreading hairs,
4-toothed, the teeth acute, 0.5 mm. long. Corolla slender, glabrous,
white, 3 cm. long, the lobes spreading, oblong-lanceolate to oblong-
oblanceolate, obtuse, 10 mm. long, 3 to 3.5 mm. wide. Anthers linear,
5 to 6 mm. long, spirally twisted when dry. Exserted portion of the
style slender, glabrous 4 to 4.5 cm. long.

Mrnporo, Bongabong River (3714 Merritt) March 21, 1906. In forests, river
valley near sea level. A species well characterized by its’ long corolla tubes and
very long exserted styles.

PSYCHOTRIA Linn.

Psychotria crispipila Merrill sp. nov.

A shrub 3 to 4 m. high with elliptical-ovate leaves which are promi-
nently neryed and rather densely fulyous pubescent beneath with crisped
hairs, the flowers crowded in dense subcapitate cymes on 2.5 to 4 cm.
long terminal peduncles. Branches, petioles, and inflorescence also rather
densely fulvous pubescent with crisped hairs. Leaves 7 to 10 cm. long,
3 to 5 cm. wide about equally narrowed to both the acute apex and base,
coriaceous, glabrous above; nerves 9 to 11 on each side of the midrib,
obscure above, prominent beneath, parallel, anastomosing near the margin,
the reticulations obscure; petioles 1 to 1.5 cm. long; stipules caducous.
Peduncles 2 or 3 from each branchlet. Flowers white, sessile or short
pediceled. Calyx 3 to 4 mm. long obscurely 4-toothed or subtruncate,
fulvous pubescent with crisped hairs especially on the margin. Corolla
5 mm. long glabrous outside, the throat villous within, 4-cleft to or
below the middle, the lobes oblong-ovate, obtuse, 2 to 3 mm. long. Fila-
ments and anthers each about 1 mm. long, the latter elliptical-oblong.
Ovary 2-celled, each cell with one ascending ovule; style about 2 mm.
long. Fruit unknown.

Luzon, District of Lepanto, Mount Data (4490 Merrill) November, 1905. In
the mossy forest at 2,200 m.

Related to Psychotria bataanensis Elmer, differing from that species in its
elongated peduncles, densely crisped fulvous pubescent branches, petioles, under
surface of its leaves and inflorescence. The leaves in the present species are
always acute at the base, while in P. bataanensis they are narrowly cordate or
auriculate.

CAPRIFOLIACE.
LONICERA Linn.

Lonicera philippinensis Merrill sp. noy.

A scandent glabrous or nearly glabrous shrub 2 to 3m. high, with 2-
lipped corollas about 1.5 cm. long. Branches dark brown, glabrous,
shining obscurely’ angled, slender, the young parts sparingly hirsute.
Leaves narrowly oblong-ovate, entire, acute, the base broad, truncate or
subcordate, 2.5 to 4 em. long, 1 to 2 em. wide, coriaceous, shining above,
quite glabrous or sparingly hirsute on the midrib on both surfaces:
nerves about 5 on each side of the midrib, the reticulations distinct ;

241

petioles about 5 mm. long, rugose, sparingly hirsute, becoming glabrous.
Flowers white, turning yellowish in age, faintly odorous, in pairs in
the upper axils, nearly sessile. Calyx 4 mm. long, ovoid, glabrous, the
teeth ovate, acute, 5 mm. long, shghtly hirsute-ciliate, the bract linear,
glabrous or nearly so, about 3 mm. long. Corolla glabrous outside, hir-
sute inside, the tube about 7 mm. long. Anthers oblong, 4 mm. long;
filaments hirsute. Ovary 3-celled; style about 13 mm. long, slender,
hirsute except just below the stigma. Fruit subglobose or ovoid, glabrous,
black when mature, 5 to 6 mm. in diameter; seeds few, 2 or 3, about
4 mm. long.

Luzon, Province of Benguet, Pauai (4775 Merrill) November 8, 1905. In
thickets, border of mossy forest at about 2,200 m.

A species of the § Nintooa, subsect. Breviflorae, apparently most closely related
to Lonicera glabrata Wall., of the Himalayan region, and quite distinct from
the only other known Philippine species of the genus, L. rehderi Merr., which
is also known only from the same province as is the present species.

CUCURBITACE.
ZANONIA Linn.

Zanonia philippinensis Merrill sp. noy.

Scandent in large trees reaching a height of from 30 to 50 m.
Branches rather slender, striate, glabrous, grayish brown. ‘'Tendrils gla-
brous 8 to 10 cm. long or more. Leaves ovate, the base broad, somewhat
cordate, coriaceous, glabrous, shining on both surfaces, paler beneath,
the apex acute, 15 to 24 cm. long, 10 to 12 em. wide, 3-nerved from the
base, the lateral nerves prominent, few, distant; petioles 2 to 3.5 em.
long. Flowers unknown. Fruit subglobose, brown, glabrous, somewhat
shining, the pericarp brittle when dry, about 20 em. in diameter. Seeds
indefinite, elliptical, flattened, 2.5 cm. long, 1.5 em. wide, wings mem-
branous, surrounding the seed, each about 6 cm. long, 4 em. wide, the base
entire, curved, the apex irregularly lobed or toothed.

Minpanao, Lake Lanao, Camp Keithley (324 Mrs. Clemens) February, 1905.
A species evidently related to the Malayan Zanonia macrocarpa Blume, differing
from the latter in its cordate leaves, smaller seeds which have much wider and
somewhat longer wings than in Blume’s species. Of this genus, Zanonia indica
Linn., has been reported from the Philippines by F.-Villar, but his record has
never been verified.

CAMPANULACE®.
PRATIA Gaud.

Pratia begonifolia Lind]. Bot. Reg. ¢. 1373; Clarke in Hook. f. Fl. Brit. Ind. 3
(1881) 422; Forbes & Hemsl. Journ. Linn. Soe. Bot. 26 (1889) 2.

Luzon, Province of Benguet, Sablan (6201 Elmer) April, 1904; Baguio to
Ambuklao (4369 Merrill) October 24, 1905. Growing on steep damp banks in
ravines. Japan to Central and Southern China, Formosa, Eastern India, and
Malaya.

No species of the genus previously reported from the Philippines.

242

WAHLENBERGIA Schrad.

Wahlenbergia bivalvis Merrill sp. nov.

An erect, glabrous, lax, branched, annual herb 20 to 40 em. high with
linear or linear-lanceolate entire leaves and solitary, long peduncled,
blue flowers, the capsule 2-celled, 2-valved at the apex inside the persistent
calyx teeth. Branches slender, terete. Leaves 1.5 to 3 em. long, 1 to
L.5 mm. wide, acuminate, sessile or the basal ones petioled and sometimes
sparingly pilose, the margins thickened, the lateral nerves obsolete.
Peduncles solitary, slender, 10 cm. long or less. Calyx glabrous, 5 mm.
long, the tube 3 mm. long, oblong-ovoid, the lobes 5, erect, linear-lan-
ceolate, acute, 2 mm. long. Corolla blue, campanulate, about 7 mim.
long, 5-lobed, the tube nearly 3 mm. long, the lobes oblong, acute, about
2mm. wide. Stamens free; anthers narrow, 2 mm. long. Ovary infe-
rior, 2-celled; style nearly 4 mm. long; stigma broadly lobed. Capsule
membranous, oblong, 6 mm. long, 3 mm. in diameter, 2-celled, 2-valved
at the apex inside the persistent erect calyx teeth, the apex convex, sub-
conical. Seeds indefinite, elliptical-oblong, glabrous, 0.5 mm. long.

Luzon, Province of Benguet, Mount Tonglon (Santo Tomas) (4811 Merrill)
November 12, 1905; Baguio to Ambuklao (4361 Merrill) October 24, 1905:
District of Lepanto, Mount Data (4559 Merrill) November 4, 1905.

A species apparently distinct from Wahlenbergia gracilis A. DC., especially in
its 2-celled, 2-valved capsules, growing on dry open grassy slopes in thin pine
forests 1,600 to 2,100 m., widely distributed but not abundant.

COMPOSITE.
AINSLIAEA DC.

Ainsliaea reflexa Merrill sp. noy.

Erect, simple, the leaves mostly radical, long petioled, the petioles
winged-margined, the inflorescence long, racemose, the heads 3-flowered,
about 12 mm. long, solitary or in pairs. Stems glabrous, terete, simple,
erect 20 to 60 em. tall with few reduced linear-lanceolate leaves. Radical
leaves membranous, glabrous or with few long hairs beneath and on the
midrib above, the blade elliptical-oblong to oblong-ovate, 2.5 to 6 cm.
long, 1 to 2 em. wide, acute, with distant, short spinulose teeth the
nerves 2 to 3 on each side of the midrib, rather abruptly contracted at
the base; petioles narrowly winged, much exceeding the lamina 6 to 8
cm. long. Heads narrow, pedicellate, the pedicels 2 to 3 mm. long,
pubescent, mostly reflexed, at least in anthesis. Involucral bracts spar-
ingly pilose with scattered hairs, the lower ones 1.5 mm. long, ovate to
lanceolate, acuminate, the inner ones linear-lanceolate to lnear-oblan-
ceolate, 12 mm. long, about 1.8 mm. wide. Achenes 4.5 to 5 mm. long,
10 ribbed, somewhat compressed, about 1 mm. thick above, pilose with
scattered spreading white hairs; pappus brownish, the bristles about 30
on each achene, 6 to 7 mm. long, plumose-pilose with 1 mm. long hairs.

24

CO

Luzon, District of Lepanto, Mount Data (4571 Merrill) November 4, 1905.
Abundant in the mossy forests at about 2,200 m.

Another form of this species is apparently represented by No. 4824 Merrill,
Mount Tonglon, Province of Benguet, Luzon, November 12, 1905, this number
growing under similar conditions as the preceding, but with the petioles scarcely
exceeding the leaf blade in length. No species of the genus has preyiously been
reported from the Philippines, the present species being apparently related to
Ainsliaea pteropoda DC.

CARPESIUM Linn.

Carpesium cernuum Linn. Sp. Pl. (1753) 859; Clarke in Hook. f. FI. Brit.
Ind. 3 (1881) 3800; Forbes & Hemsl. Journ. Linn. Soe. Bot. 23 (1888) 430.
Luzon, Province of Benguet, Mount Tonglon (4832 Merrill) November 12,
1905; Suyoe to Pauai (4688 Merrill) November 7, 1905: District of Lepanto,
Mount Data (4512 Merrill) November, 1905.

A decidedly interesting addition to our knowledge of the northern element in
the Philippine flora, growing in the mossy forests on the higher mountains and
ridges above 2,000 m., no species of the genus having been previously reported
frbm the Philippines. The form here referred to Carpesiwm cernwum Linn. is
apparently closest to the var. glandulosum Clarke in Hook. f. Fl. Brit. Ind. 3
(1881) 301.

This exceedingly variable species extends from France to the Caucasus, the
temperate Himalayan region, Khasia and Nilgherry Mountains, China, and Japan,
also according to Clarke to Java, I can find no record of its having been found
in southern China or Formosa although another species, C. abrotanoides Linn.,
has been encountered in the latter place.

DICHROCEPHALA DC.

Dichrocephala chrysanthemifolia DC. Prodr. 5 (1836) 372; Hook. Fl. Brit.
Ind. 3 (1881) 245; Forbes & Hemsl. Journ. Linn. Soc. Bot. 23 (1888) 406.

Luzon, District of Lepanto, Mount Data (4533 Merrill) November, 1905. On
dry grassy slopes in thin pine forests at about 2,000 m., the second species of
the genus to be found in the Philippines. China to British India and tropical
Africa.

EMILIA Cass.
Emilia pinnatifida Merrill sp. nov.

An erect, simple or shghtly branched herb 15 to 35 em. high with
glabrous or more or less pubescent usually finely lyrately lobed leaves,
the involueral bracts much shorter than the flowers. Stems glabrous,
finely channeled. Leaves various, the radical ones sessile or short
petioled, 4 to 5 em. long or less, 1 to 1.5 cm. wide, glabrous or with
scattered weak hairs beneath, the lobes irregular, 2 to 4 mm. wide, often
extending nearly to the midrib, their margins irregularly crenate, the
terminal lobe usually larger than the lateral ones; cauline leaves similar
to the radical, sessile, clasping, few, the lower ones as long as the radical
leaves, the upper one much reduced. Peduncles 2 to 7 cm. long. Heads
1 cm. long, the flowers pink. Involucral bracts glabrous, 1-seriate, 7
to 8 mm. long, about 5 mm. wide, oblong-lanceolate, acute or slightly
acute, about 5-nerved. Achenes when mature nearly 3 mm. long, ob-
securely angled, finely pubescent ; pappus copious, soft, white, about 6 mm.
long, minutely scabrid. Corollas slender, 7 to 8 mm. long, 5-lobed, the

: 244

lobes lanceolate, acute or obtuse, about 2.5 mm. long. Style arms nearly

?

1.5 mm. long. Anthers slender, 3 mm. long or less.

Luzon, Province of Benguet, Bugias (4664 Merrill) October 28, 1905. The fol-
lowing specimens are apparently referable here: Benguet, Baguio (6599, 6606
Bilmer) June, 1904; (4265 Merrill) October, 1905.

A species characteristic of the open, dry, grassy slopes in thin pine forests from
1.500 to 2,100 m., widely distributed in Benguet and Lepanto; abundant. The
species here proposed differs from Emilia sonchifolia DC., in the involucral bracts
being constantly much shorter than the flowers, in this character approaching
Hmilia flammea Cass. As in Emilia sonchifolia, the species is exceedingly
variable, but is apparently distinct from all other previously described forms.
Emilia sonchifolia Elmer, in herb.

GYNURA Cass.

Gynura clementis Merrill sp. nov.

Seandent, more or less pubescent with crisped hairs throughout.
Branches dark brown, obscurely angled, ferruginous crisped pubescent
with scattered hairs. Leaves alternate, elliptical-oblong to narrowly
oblong-obovate, 5 to 8 em. long, 1.5 to 3.5 cm. wide, coriaceous, acute
at both ends, coarsely irregularly toothed, the teeth acute, the upper
surface subscabrid, the lower rather densely ferruginous pubescent;
nerves 5 to 6 on each side of the midrib; petioles 0.8 to 1.5 cm. long,
pubescent, the stipules orbicular, often 1 cm. in diameter. Heads
purplish, about 25-flowered, 1.5 cm. long or less, in few flowered axillary
and terminal corymbs 10 em. long or less. Involucre cylindrical or
subcampanulate, the bracts 8, oblong-lanceolate, acute, 7 to 8 mm. long,
1.5 to 2.5 mm. wide, somewhat pubescent with erisped hairs, and with
several short linear bracteoles at the base. Flowers homogamous, pur-
plish, slender, about 12 mm. long including the achene, the tube 7 mm.
long, the upper slightly expanded portion about 3 mm. long, the teeth 5,
lanceolate, acute, 1.5 mm. long. Stamens 5; anthers 2 mm. long,
lanceolate, the cells not produced at the base. Style arms 3 mm. long,
slightly pubescent. Achenes oblong, 3 mm. long, brown, about 10-ribbed,
sparingly hispid. Disk glabrous, pitted.

MinpAnao, Lake Lanao, Camp Keithley (49 Mrs. Clemens) January, 1906.

MYRIACTIS Less.
Myriactis humilis Merrill sp. nov.

An herb 20 em. high or less, sparingly pilose or nearly glabrous, with
long petioled deeply and coarsely pinnately lobed leaves and_ solitary
long peduncled heads about 7 mm. in diameter. Branches few, terete,
with few scattered weak hairs. Leaves membranous, 3 to 5 em. long,
1.5 to 3 cm. wide, coarsely lyrately lobed, the lobes entire or coarsely
toothed, apiculate, both surfaces with few scattered weak hairs, the upper
leaves much smaller, the uppermost entire and bract like; petioles 1 to
3 em. long. Peduncles slender, 5 em. long or less, solitary. Heads

5 mm. long, the ray flowers short, white, the disk flowers greenish yellow ;

245

involucral bracts 2-seriate, narrowly oblong, obtuse or acute, sparingly
pilose on the median portion, about 4 mm. long, 1 mm. wide. Ray
flowers pistillate, ligulate, 2 or more seriate, the ligule oblong about
1.5 mm. long or less; style short, 2-cleft at the apex. Disk flowers
greenish yellow, tubular, hermaphrodite. Corolla 1.5 mm. long, regularly
5-cleft, the lobes 1 mm. long, acute. Anthers oblong-ovate, acute, the
base obtuse. Style equaling the corolla, 2-cleft at the apex. Achenes
compressed, glabrous, oblong, the margins thickened, 3 mm. long, 1.2 mm.
wide, tipped with a short somewhat glandular ring. Pappus 0.

Luzon, District of Lepanto, Mount Data (4530 Merrill) November, 1905. Com-
mon in the damp, mossy forests at 2,100 m. and above. The first species of the
genus to be reported from the Philippines.

SENECIO Linn.

Senecio luzoniensis Merrill sp. noy.

An erect, stout, glabrous, simple or sparingly branched herb about
1m. high, with oblong-lanceolate, coriaceous, rather obscurely irregularly
crenate leaves and terminal corymbose many flowered panicles, the heads
about 1 to 1.3 cm. long, 1.5 cm. or more in diameter, including the rays.
Stems terete, firm, glabrous, reddish brown. Leaves 10 to 15 em. long,
1.5 to 2.5 cm. wide, glabrous, strongly reticulate veined, the upper ones
sessile or subsessile, the lower ones petiolate, the petiole 1 em. long, base
acute, apex usually long acuminate. Panicles about 15 em. long, nearly
as wide, the branches ascending, the bracts and bracteoles linear, the
primary branches subtended by linear 3 to 4 cm. long leaf-like bracts, the
bracteoles usually about 2 or 3 mm. long. Heads about 25-flowered ;
involucral bracts about 13, 1-seriate, imbricate, free, oblong-lanceolate,
7 to 8 mm. long, 1.5 to 2 mm. wide, acute glabrous, membranous
inargmed, a few short outer bracts at the base. Ray flowers about 6
in each head, yellow, the tube slender, 4 to 5 mm. long, the ligule oblong,
10 to 11 mm. long, 3.5 mm. wide, strongly 4-nerved, obtuse, the apex
with three small blunt teeth. Disk flowers yellow, the corolla tubular,
inflated above, about 8 min. long, 5-lobed, the lobes about 1.3 mim. long,
oblong, blunt. Anthers slender, about 3 mm. long, blunt at the base.
Style 2-cleft, the arms about 1.5 mm. long, truncate. Achenes about 4
mm. long, less than 1 mm. thick obscurely 10-striate, glabrous, narrowly
oblong. Disk paleaceous.

Luzon, Province of Benguet, Suyoe to Pauai (4791 Merrill) November 7, 1905.
A stout erect herb growing in grass lands on high ridges near the lower limits
of the mossy forest at about 2,150 m.

SOLIDAGO Linn.

Solidago virgaurea Linn. Sp. Pl. (1753), 880.

Luzon, Province of Benguet, Baguio to Ambuklao (4864 Merrill) October 24,
1905. Widely distributed in the region of Pinus insularis Endl., 1.500 to 2,000 m.
North America, Europe, and temperate Asia south to Formosa.

246
SPILANTHES Linn.

Spilanthes ovata Merrill sp. noy.

An erect or spreading slightly branched, nearly glabrous herb with
opposite, short, broadly-ovate to triangular-ovate, irregularly and rather
coarsely crenate-dentate acute leaves and solitary long peduncled radiate
heads about 1.5 em. in diameter. Branches purplish, somewhat angled or
channeled, glabrous or sparingly pubescent. Leaves membranous or sub-
membranous, 2.5 cm. long or less, 1.2 to 1.8 em.. wide, the base broad,
truncate or slightly cordate, 3-nerved, paler beneath, the margins thick-
ened, usually somewhat ciliate, the nerves rather prominent beneath, often
with few scattered hairs, the reticulations distinct; petioles 3 to 4 mm.
long. Peduncles about 7 em. long. Heads yellow; involucral *bracts
sub 2-seriate, oblong-lanceolate, acute or acuminate, about 5 mm. long,
the margins ciliate. Ray flowers about 10, the tube cilate, about 2 mm.
long, the ligule oblong-ovate, 10 mm. long, 4 mm. wide, 10-nerved,
truncate, 3-toothed at the apex, the teeth short, rounded. Disk flowers
tubular, the corolla 5-toothed, the teeth about 0.8 mm. long, acute.
Anthers about 1 mm. long, the cells acute at the base. Style arms nearly
1 mm. long, truncate. Achenes 2 mm. long, flattened, 2-keeled, ciliate
on the keels; pappus of 2 slender, scabrid, 1 mm. long bristles, the achenes
of the ray flowers triquetrous. Pales 4 to 5 mm. long, keeled, or the
inner ones flat or nearly so, 3-nerved, obtuse, irregularly minutely toothed
at the apex, the keel minutely scabrid.

Luzon, District of Lepanto, Mount Data (4573 Merrill) November 4, 1905.
Rather common on dry grassy slopes in thin pine forests at about 2,100 m.

A species apparently closely related to Spilanthes grandiflora Turez., differing
from that species in its much shorter, very differently shaped leaves.

fe 5 RY , An tly iy tab aM ‘ ye HR Ge

‘NOTEWORTHY PHILIPPINE.

i

Ra aR tS By Exumer D. MERRILL
” : A (Fred the Botanical Section of the Biological Laboratory, Bureau of Science, Manila, P. I.)

REPRINTED FROM iy
THE PHILIPPINE JOURNAL OF SCIENCE
S opablanaas by the Bureau of Science of the Philippine Government, Manila, P. 1
Vor. III, No. 4, SECTION C, Botany, Aucust, 1908

MANILA hi" iit
BUREAU OF PRINTING ik

PREVIOUS PUBLICATIONS OF THE BUREAU OF COVERNMENT AE
LABORATORIES, _ ‘

1No. 1, 1902, Biological Laboratory.—Preliminary Report of. the Appearance in the dy

sth ache i ‘Islands of a Disease Clinically Resembling Glanders. By R. P; Strong, M. D.
0
Its Use as an Intestinal Antiseptic in Cholera and Dysentery. Preliminary Notes. By
Paul C. Freer, M. D., Ph. D.

1No. 3, 1908, Biolog gical Laboratory.—A Preliminary Report on Trypanosomiasis of

Horses in ihe Philippine Islands. By W. E. Musgrave, M. D., and Norman E. Williamson. .

1No. 4, 1908, Serum Laboratory.—Preliminary Report on the Study of Rinderpest of
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Reference to Surra in the Philippine Islands. By W. E. Musgave, M. D., and Moses

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1 No. 6; 1903.—New and Noteworthy -Plants, I. The American eet in the Philip-
pine Flora. By Elmer D. Merrill, Botanist. (Issued January 20, 1904

1No. 7, 1903, Chemical Laboratory. Pipe Gutta Percha and Rubber of the Philippine
Islands. By Penoyer L. Sherman, jr., Ph.

1No. 8, 1903.—A Dictionary of the Plant Names of the Philippine Islands. By Elmer
D. Merrill, Botanist.

1No. 9, 1908, Biological and Serum Laboratories—A Report on Hemorrhagic Septi-
cemia in ” Animals in the Philippine Islands. By Paul G. Woolley, M. D., and J. W.
Jobling, M. D.

1 No. 10, 1908, Biological Laboratory.—Two Cases of a Peculiar Form of Hand Infection
(Due to an Organism Resembling the Koch- “Weeks Bacillus). By John R. McDill, M. D.,
and Wm. B. Wherry, M. D.

1 No. 11, 1903, Biological Laboratory.—Entomological Division, Bulletin No. 1: ‘Prelimi-
nary Bulletin on "Insects of the Cacao. (Prepared Especially for the Benefit of Harmers:)
By Charles §. Banks, Entomologist.

1 No. 12, 1903, Biological Laboratory.—Report on Some Pulmonary Lesions Produced by
the Bacillus of Hemorrhagic Septicemia of Carabaos. By Paul G. Woolley, M.

No. 13, 1904, Biological Laboratory—A Fatal Infection by a Hitherto Undescribed
Chromogenic Bacterium: Bacillus Aureus Fetidus. By Maximilian Herzog, M. D.

1No. 14, 1904.—Serum Laboratory: Texas Fever in the Philippine Islands and the Far
East. By J. W. Jobling, M. D., and Paul G. Woolley, M. D. Biological Laboratory:
Entomological Division, Bulletin No. 2: The Australian Tick (Boophilus Australis Fuller)
in the Philippine Islands. By Charles §. Banks, Entomologist.

No. 15, 1904, Biological and Serum Laboratories.—Report on Bacillus Violaceus Ma-
nile: A Pathogenic Micro-Organism. By Paul G. Woolley, M. D.

1 No. 16, 1904, Biological Laboratory.—Protective Inoculation Against Asiatic Cholera:
An Experimental Study. By Richard P. Strong, M. D.

No. 17, 1904.—New or Noteworthy Philippine Plants, II. By Elmer D. Merrill, Botanist.

1 No. 18, 1904, Biological Laboratory.—I. Amebas: Their Cultivation and Etiologie
Significance. By W. E. Musgrave, M. D., and Moses T, Clegg. Il. The Treatment of
Intestinal Amebiasis (Amcebic Dysentery) in the Tropics. By W. E. Musgrave, M. D.

No. 19, 1904, Biological Laboratory.—Some Observations on the Biology of the Cholera
Spirilum. By W. B. Wherry, M. D.

No, 20, 1904.—Biological Laboratory: I. Does Latent or Dormant Plague Exist Where
the Disease is Endemic? By Maximilian Herzog, M. D., and Charles B. Hare. Serum
Laboratory: Il. Broncho-Pneumonia of Cattle: Its Association with B. Bovisepticus.
By Paul G. Woolley, M. D., and Walter Sorrell, D. V. S. III. Pinto (Pafio Blanco). By
Paul G. Woolley, M. D. Chemical Laboratory: IV. Notes on Analysis of the Water from
the Manila Water Supply. By Charles L. Bliss, M. 8S. Serum Laboratory: V. Frambeesia:
Its Occurrence in Natives in the Philippine Islands. By Paul G. Woolley, M. D.

No. 21, 1904, Biological Laboratory.—Some Questions Relating to the Virulence of
cam Bal Organisms with Particular Reference to Their Immunizing Powers. By Richard

rong
No. 23, 1904, Bureau of Government Laboratories.—I1. A Description of the New Build-
ings of the Bureau of Government Laboratories. By Paul C. Freer, M. D., Ph. D. II. A
Pere nae of the Library of the Bureau of Government Laboratories. By Mary Polk,
ibrarian. ,
1No. 23, 1904, Biological Laboratory.—Plague: Bacteriology, Morbid Anatomy, and
datas etd (Including a Consideration of Insects as Plague Carriers). By Maximilian
erzog, I

No. 24, 1904, Biological Laboratory.—Glanders: Its Diagnosis and Prevention (Together
with a Report on Two Cases of Human Glanders Occurring in Manila and Some Notes on the
Bacteriology and Polymorphism of Bacterium Mallei). By William B. Wherry, M. D.

No. 25, 1904.2—Birds from the Islands of Romblon, Sibuyan, and Cresta de Gallo. By
Richard “C. McCregor.

No. 26, 1904, Biological Laboratory.—The Clinical and Pathological Significance of
Balantidium Coli. By Richard P. Strong, M. D.

No. 27, 1904.—A Review of the Identification of the Species Described in Blanco’s
Flora de Filipinas. By Elmer D, Merrill, Botanist.

No. 28, 1904.—I. The Polypodiacez of the Philippine Islands. II. Edible Philippine
Fungi. By Edwin B. Copeland, Ph. D.

No. 29, 1904.—I. New or Noteworthy Philippine Plants, III. II..The Source of Manila
Elemi. By Elmer D. Merrill, Botanist.

No. 80, 1905, Chemical ‘Laboratory.—I. Autocalytic Decomposition of Silver Oxide.
Il. Hydration in Solution. By Gilbert N. Lewis, Ph. D.

No. 81, 1905, Biological Laboratory.—I. Notes on a Case of Hwmatochyluria (Together
with Some Observations on the Morphology of the Embryo Nematode, si a Nocturna).
By William B. Wherry, M. D., and John R. McDill, M. D., Manila, wT A Search
Into the Nitrate and Nitrite Content of Witte’s “Peptone,” with Special Batarenes to Its
pecan a Demonstration of the Indol and Cholera-Red Reactions. By William B.

erry, }

1 Out of print.

2 The first four bulletins in the ornithological series were published by the Bthnological
Survey under the title ‘Bulletins of the Philippine Museum.” Later ornithological
publications of the Government appeared as publications of the Bureau of Government
Laboratories,

(Concluded on third page of cover.)

Chemical Laboratory.—The Preparation of Benzoyl-Acetyl Peroxide and ~ Hf

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI.

By Evmer D. Merrit.

(From the Botanical Section of the Biological Laboratory, Bureau of Science,
+ Manila, P. 1.)

During the past five years approximately 1,000 new species have been
_____ deseribed from the Philippines, the material on which they were based

oe being for most part of recent collection. In spite of the large amount
# z of work accomplished, the number of new forms that are constantly
og being found is surprising, and our herbarium to-day contains several
as hundred undescribed species, while additional ones are found in nearly
every collection that is made, especially in those from previously unex-
By? plored regions.

In the present paper representatives of six genera previously not
known from the Archipelago are recorded. Hight species, first described
from extra-Philippine material, are also here listed for the first time
from the Philippines, and about sixty species are described as new. One
- new genus, Sagittipetalum, of the Rhizophoraceae, is described.

a | GRAMINEZ.

4 | ORYZA L.

Bee Oryza manilensis sp. nov.

Annua, erecta, 1 ad 1.3 m alta; foliis flaccidis, 10 ad 25 cm longis,
——S« 6 ad % mm Matis; paniculis laxis, 10 ad 15 cm longis, flaccidis, pauci-
be floris; spiculis 4 mm longis, arista 8 ad 10 mm longa.

An annual 1 to 1.3 m high. Culms slender, glabrous, the nodes 6
or 7, glabrous. Leaves flaccid, minutely scabrid when dry, 10 to 25
em long, 6 to 7 mm wide; sheaths rather loose, shorter than the inter-
nodes, the ligules about 2.5 mm long. Panicles lax, 10 to 15 cm long,
the branches slender, scabrid, usually drooping, few-flowered. Spikelets
4 mm long, the empty glumes narrowly lanceolate, acuminate, 1.5 mm
long. Flowering glume oblong, granulate, spinescent-ciliate on the keel
and ribs, its awn slender, straight, 8 to 10 mm long. Palea slightly
exceeding the flowering glume, short-acuminate, spinescent on the keels.

Luzon, Province of Rizal, Antipolo, Bur. Sci. 2194 Ramos, March, 1907, grow-
ing in open wet places.

A species allied to Oryza ridleyi Hook. £., of the Malayan Peninsula and

Borneo, differing from that species in its smaller spikelets, fide Hackel in lit.
219

220) MERRILL.

CHLORIS §&w.

Chloris mearnsii sp. nov.

Caespitosa, perennis, usque ad 40 cm alta; culmis gracilibus, genicula-
tis; spicis 3, digitatis, gracilibus, longe exsertis; foliis linearibus, acumi-
natis, 5 ad 8 cm longis; spiculis imbricatis, atropurpureis, 3.5 mm
longis, arista 1 cm longa.

A tufted perennial 40 cm high or less. Culms very slender, glabrous,
geniculate below, the inflorescence long-exserted. Leaves 5 to 8 cm long,
2 to 4 mm wide, minutely seabrid, the sheaths longer than the inter-
nodes, the ligule consisting of a few long white hairs. ‘Spikes 3, digitate,
slender, 5 to 7% em long, the spikelets imbricate, dark-purple. Empty
glumes hyaline, 1-nerved, scabrid on the nerves, the first 1.2 mm long, ~
linear, the second 3 mm long, linear-lanceolate, mucronate-acuminate.
Flowering glume lanceolate, dark-purple, 3.5 mm long, scabrid, the tip-
with a slender straight awn 1 cm long. Palea equaling the glume,
scabrid! Seed narrowly oblong, 2.5 mm long. Callus pilose.. Rachilla
produced 1 to 1.2 mm and bearing a minute, 0.5 mm long, glume, the
awn slender, scabrid, 3 to + mm long.

Luzon. Province of Ilocos Norte, Bur. Sci. 2294 Mearns, January, 1907.

A species allied to Chloris cynodontoides Bal., fide Hackel in lit., but that
species has blunt leaves, while in the present species they are long-acuminate.

CYPERACEZ.
CAREX Linn.

Carex cryptostachys Brongn. in Duperr. Voy. Bot. (1829) 152; Clarke in
Hook. f. Fl. Brit. Ind. 6 (1894) 714.

Luzon, Province of Rizal, Bur. Sci. 1760 Ramos, January, 1907: Province of
Sorsogon, Sorsogon, Hliner 7306, November, 1905.

Malay Peninsula to southern China, south to New Guinea; new to the Philip-
pines.

The former number cited was compared by me with authentic material in Herb.
Kew, and the identification has also been confirmed by Herr G. Kiikenthal.
Elmer’s specimen is manifestly the same.

ARACE/#.
ALOCASIA Neck.

Alocasia heterophylla (Presl) comb. nov.

Caladium heterophyllum Presl Rel. Haenk. 1 (1827) 148.
Alocasia warburgii Engl. Bot. Jahrb. 25 (1898) 25; Merr. in Philip. Journ.

Sci. 1 (1906) Suppl. 33.

Presl’s type, preserved in the Prague herbarium, is identical with the type of
Alocasia warburgii, direct comparison having been made by me in the Berlin
herbarium, Dr. Engler concurring in the identity of the two species. Presl’s

name being the older is here retained.

bo
es

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. Z

MORACE.AA.
ARTOCARPUS Forst.

Artocarpus woodii sp. nov.

Arbor ca. 10 m alta, glabra; foliis anguste oblongo-obovatis, nitidis, 8
ad 15 em longis, abrupte late acuminatis, basi cuneatis, integris, nervis
utrinque 5 vel 6, prominentibus, anastomosantibus; fruetibus axillaribus,
pedunculatis, irregulariter 7—12-lobatis, 6 ad 9 em diam., rugosis.

A tree about 10 m high, glabrous. Branches light-gray, the tips
brown and with few appressed hairs. Leaves narrowly oblong-obovate,
glabrous, shining, submembranaceous, entire, 8 to 15 em long, 5 to 7%
cm wide, the apex broadly and abruptly acuminate, the acumen retuse,
narrowed below to the cuneate base; nerves 5 or 6 on each side of the
midrib, prominent beneath, anastomosing near the margins, the reticula-
tions lax; petioles 3 to 5 cm long, dark colored, pruinose. Fruit pe-
duncted, the peduncles 5 to 7 em long, solitary in the upper axils, sub-
globose, 6 to 9 cm in diameter, irregularly obscurely lobed, the lobes
rounded, rugose, gray, the anthocarps truncate or depressed, 2 to 3 mm
in diameter. Ripe seeds 6 to 12, about 2 em long.

Bucas (off the northeast coast of Mindanao), Merrill 5259, October 4, 1906.

Growing in ravines along the forest border at an altitude of about 10 m above
the sea, well characterized by its shining narrowly oblong-obovate leaves, prom-
inent nerves, lax reticulations, and solitary, peduneled, rugose fruits. Dedicated
to Major-General Leonard Wood, to whom [ am indebted for the opportunity
to explore botanically many of the remote islands in the southern Philippines.

ANONACEAE.
GONIOTHALAMUS Blume.

Goniothalamus dolichopetalus sp. noy.

Arbuscula glabra; foliis oblongo-lanceolatis vel lanceolatis, usque ad
20 cm longis, subcoriaceis, acuminatis, basi acutis, nervis obscuris ; flo-
ribus axillaribus, solitariis, 10 ad 12 em longis; pedicillis ca. + em longis.

A shrub about 1 m high, glabrous throughout, the branches terete,
slender, light-gray. Leaves lanceolate to oblong-lanceolate, rarely oblan-
ceolate, 11 to 20 em long, 2.5 to 5 em wide, subcoriaceous, dull, apex
acuminate, the acumen blunt, base acute; nerves about 12 on each side of
the midrib, obscure, distant, irregular, anastomosing, the reticulations
nearly obsolete; petioles about 5 mm long. Flowers solitary, axillary,
their pedicels slender, about 4 cm long, slightly thickened upwards.
Sepals 3, triangular-ovate, acuminate, shghtly pubescent, about 3.5 mm
long, 3 mm wide. Outer petals linear-lanceolate, 10 to 12 em long, 1
em wide, somewhat narrowed towards the base and gradually narrowed
upwards, submembranaceous, glabrous, flat ; inner petals oblong to oblong-
lanceolate, about 13 mm long, 5 to 6 mm wide, connivent, acuminate,

292, MERRILL.

glabrous, slightly clawed. Stamens many, 2.4 mm _ long, somewhat
curved, their connectives short, triangular. Carpels indefinite, oblong,
slightly pubescent, 1-ovuled; styles slender, 1.5 mm long.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9268, 9261 Whitford
& Hutchinson, January, 1908.

An undershrub in dipterocarp forests 30 to 60 m above sea level, a species well

characterized by its very long flowers.
POLYALTHIA Blume.

Polyalthia venosa sp. nov.

Foliis oblongo-ellipticis, acuminatis, valde reticulato venosis, nervis
utrinque ca. 9; floribus axillaribus et caulinis solitariis, pedicellatis ;
petals elliptico-ovatis vel lanceolato-ovatis ca. 2 cm longis, pilosis.

A shrub or small tree 4 m high or less, the branches slender, terete,
glabrous, brown, the branchlets densely ferruginous-pubescent. Leaves
oblong-elliptical, 10 to 16 em long, 3 to 6 em wide, papyraceous, glabrous
and shining on both surfaces except the midrib which is shghtly pubes-
cent, paler beneath, the base acute or rounded, apex sharply acuminate ;
nerves about 9 on each side of the midrib, distant, very prominent beneath,
anastomosing, the reticulations very prominent; petioles about 3 mm
long, densely pubescent. Flowers solitary, from the leaf-axils on the
ultimate branchlets and from tubercles on the trunk, their pedicels slen-
der, 1.5 to 3.5 em long, densely ferruginous-pilose. Sepals 3, free, ovate,
about 8 mm long, 6 mm wide, slightly narrowed below, apex acute, 7-
nerved, pilose outside, glabrous within. Petals 6, elliptical-ovate to
lanceolate-ovate, about 2 em long, 6 to 11 mm wide, the inner three
slightly narrower than the outer ones, somewhat narrowed below, apex
acute, glabrous inside, rather densely pilose or pilose-hirsute outside.
Stamens indefinite, 2 mm long, the anthers concealed by the truncate
overlapping connectives. Carpels many, oblong, appressed-hirsute, 2
mim long; stigmas ellipsoid, pubescent; styles very short or none; ovules
2, subbasal. Fruit ellipsoid or ovoid, about 1 em long, apiculate, slightly
pubescent, each with two seeds.

MINDANAO, District of Zamboanga, Port Banga, Por. Bur. 9108, 9232 Whitford
& Hutchinson, December, 1907, in dipterocarp forests 15 to 30 m above sea level.

A species well characterized by its pilose flowers which are both axillary and
cauline, and by its strongly nerved leaves.

Polyalthia elongata sp. noy.

Arbor glabra: foliis oblongis vel oblongo-lanceolatis, coriaceis, 30 ad
35 cm longis, basi cordatis, nervis utrinque ca. 16; petalis 4 ad 4.5 em
longis, oblongo-lanceolatis, glabris vel sparse pubescentibus.

A tree 16 m high, glabrous or nearly so throughout, the branches
grayish-brown, rather slender, terete. Leaves 30 to 35 em long, about
11 em wide, coriaceous, shining, the base broad, cordate, the apex
acuminate; nerves about 16 on each side of the midrib, very prominent

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 223

beneath, the reticulations fine, indistinct ; petioles very stout, 5 mm long.
Flowers lateral (?), nearly glabrous, pedicels about 4 cm long. Sepals
3, broadly ovate, united below, acute, slightly pubescent, about 3 mm
long. Petals 6, flat, glabrous, or with few scattered hairs, oblong-lan-
ceolate, acute, the three outer ones 4 to 4.5 cm long, 11 to 13 mm wide,
the three inner ones slightly shorter and narrower. Stamens indefinite,
1.1 mm long, connectives truncate, overlapping. Carpels indefinite,
somewhat pubescent.

Mrnpanao, District of Zamboanga, Port Banga, For. Bur. 9456 Whitford &
Hutchinson, February, 1908, in forests at 50 m above the sea.

A species well characterized by its very long, strongly nerved and cordate
leaves, the attachment of the inflorescence not being indicated and the specimens
not showing the same. A very similar species, in fruit, from the same locality
is represented by For. Bur. 9113 Whitford & Hutchinson, but the leaves are
relatively narrower, not cordate at the base and have about 25 pairs of lateral

nerves.
MELODORUM Dunal.

Melodorum rufum (Presl) comb. nov.

Anona rufa Presl Rel. Haenk. 2 (1835) 75.

Luzon, without locality, Haenke in Herb. Prague, native name Apnit, ex Presl.

After examining Presl’s type in Herb. Prague I do not hesitate to refer this
species to Melodorum, it apparently being allied to M. latifoliwm, differing from
that species in its glabrous, acuminate leaves. Index Kewensis gives the locality
of the species as “Am. trop.” but on what authority is not known, probably,
however, on the assumption that Presl was correct in describing the species as
Anona, the genus being largely developed in tropical America, and represented
in the Malayan region only by introduced and cultivated species.

OROPHEA Blume.

Orophea bracteolata sp. nov.

Arbor parva: foliis oblongis, usque ad 12 cm longis, breviter acumi-
natis, nervis utrinque ca. 9, valde obliquis; floribus in cymis axillaribus
congestis, petalis interioribus valde unguiculatis, laminis rhomboideis.

A small tree, about 9 m high, the branches terete, dark-reddish-brown,
glabrous, lenticellate, the ultimate branchlets somewhat ferruginous-
pubescent. Leaves oblong, subcoriaceous, 9 to 12 em long, 3 to 4 em
wide, base acute or rounded, often slightly oblique, the apex short-
acuminate, somewhat shining, glabrous on both sides, except on the
midrib and nerves beneath which are appressed-pubescent ; nerves about
9 on each side of the midrib, oblique, curved-ascending, very prominent
beneath; petioles 2 to 3 mm long, pubescent. Cymes axillary, densely
ferruginous-pubescent, densely flowered, 1.5 cm long or less, the pedicels
about 5 mm long, pubescent, with a large ovate, strongly acuminate,
pubescent bracteole about 3.5 mm long, at about the middle. Sepals 3,
ovate, acute, pubescent, about 3 mm long. Outer three petals broadly
ovate, acute or slightly acuminate, ferruginous-pubescent, 4 to 5 mm long,

722994

224 MERRILL.

5- to T-nerved; inner three petals about 7 mm long, the claw 3 mm long,
the blade rhomboidal, 4 mm long, free. Stamens about 12, 1 mm long,
miliusoid. Carpels many, pubescent, 1 mm long, the style very short.
Fruit unknown. ;

Luzon, Province of Cagayan, San Vicente, For. Bur. 7055 Klemme, May, 1907,
in dense flat forests slightly above sea level, Cag., Panganauin.

A species resembling Orophea cumingiana and O. enterocarpoidea in gross char-

acters but readily distinguished by its large bracteoles.

Orophea luzoniensis nom. noy.

Orophea maculata Merr. in Govt. Lab. Publ. (Philip.) 35 (1906) 11; Philip.
Journ. Sci. 1 (1906) Suppl. 54, non Scort.

In publishing the species under the specific name maculata, the fact was
overlooked that the same name had previously been used for a different species
of the same genus.

UNONA Linn. f.

Unona rubra sp. nov.

Arbuscula 1.5 m alta; foliis lanceolatis vel late oblongo-oblanceolatis,
submembranaceis, acuminatis, basi inaequilateralibus, leviter auriculato-
_cordatis, subtus pallidis, glabris; floribus rubris, axillaribus, solitariis, ca.
2 cm longis.

A small shrub about 1.5 m high, the branches dark-gray, terete, slender,
rather densely pubescent with short spreading hairs, especially the
younger ones. Leaves lanceolate to broadly oblanceolate, about 20 em
long, 4 to 6 em wide, pale, especially beneath, glabrous, submembranaceous,
apex shortly acuminate, somewhat narrowed below to the mequilateral
base which is auriculate-cordate; nerves about 15 on each side of the
midrib, curved-ascending, anastomosing, distinct beneath, the reticula-
tions obsolete or nearly so; petioles densely pubescent, about 3 mm
long. Flowers few, red, solitary, in the upper axils, their pedicels, about
5 mm long, glabrous. Sepals 3, free, ovate, 7 mm long, 5 mm wide,
acute, glabrous or with few hairs outside. Petals fleshy, coriaceous,
oblong, obtuse or blunt, about 18 mm long, 4 to 5 mm wide, the sides
parallel, nearly glabrous. Stamens indefinite, 2 mm long, connectives
truncate, overlapping. Carpels about 15, oblong, 1.5 mm long, hirsute,
each with 4 parietal ovules; styles ovoid, the stigmas minutely papillate.

3ALABAC, Bur, Sci. 480 Mangubat, March 1, 1908, in forests.

A characteristic species, distinguishable by its glabrous pale leaves and solitary
axillary red flowers.

POPOWIA Endl.

Popowia polyandra (Presl) comb. nov.

Bocagea polyandra Presl Rel. Haenk. 2 (1835) 77.

Luzon, Province of Sorsogon, Haenke, in Herb. Prague: Province of Batangas,
7724 Curran & Merrill, November, 1907. Nucros, For. Bur. 4302, 5229, 52381,
7289 Bverett, June, 1906, and September, 1907; Gimagaan River, Whitford 1561,
May, 1906. MrInpANAO, District of Zamboanga, For. Bur. 9275 Whitford &
Hutchinson, January, 1908; Lake Lanao, Mrs. Clemens 1007, April, 1907, and
without number July, 1907.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 220

A widely distributed species in the Philippines, extending from central Luzor
to southern Mindanao, the only representative of the genus known from the
Archipelago. After an examination of Presl’s type in Herb. Prague, I am able to
affirm the identity of the above-cited specimens, and accordingly transfer the
species to the genus in which it properly belongs.

PHAEANTHUS Hook. f. et Thoms.

Phaeanthus ebracteolatus (Presl) comb. nov.

Uvaria ebracteolata Presl Rel. Haenk. 2 (1835) 77.

Uvaria tripetala Blanco Fl. Filip. (1837) 465, non Roxb.

Unona tripetala Blanco 1. ¢. ed. 2 (1845) 324; ed. 3, 2: 236.

Phaeanthus cumingii Miq. Fl. Ind. Bat. 2 (1859) 51; Vidal Phan. Cuming.
Philip. (1885) 93; Rev. Pl. Vase. Filip. (1886) 42; Ceron Cat. Pl. Herb. (1892)
11; Merr. in Forest. Bur. Bull. 1 (1903) 20; Philip. Journ. Sei. 1 (1906)
Suppl. 54.

Phaeanthus malabaricus Vid. Cat. Pl. Prov. Manila (1880) 16; F.-Vill. Nov.
App. (1880) 7, non Bedd.

Phaeanthus nutans F.-Vill. Nov. App. (1880) 6, non Hook. f. et Thoms.

A very common and widely distributed endemic species for which Presl’s
specific name is here adopted, being the oldest valid one. In January, 1908,
I had opportunity to examine Presl’s type, preserved in the Prague Herbarium,
and consider it to be identical with Cuming’s plant on which Miquel based his
Phacanthus cuningir.

MONIMIACEZE.

KIBARA Endl.

Kibara mollis sp. nov.

Omnibus partibus dense et molliter stramineo-fulvo-pilosis vel pubes-
centibus; foliis ellipticis vel elliptico-ovatis, membranaceis, 12 ad 16 cm
longis, apice breviter acuminatis, nervis utrinque 5 vel 6; inflorescentiis
paniculatis vel racemoso-paniculatis, paucifloris, solitariis, axillaribus, 6
ad 10 em longis, ramis patentibus, inferioribus ca. 3 cm longis; re-
ceptaculis elliptico-ovatis, chartaceis, extus dense pilosis; carpellis ca.
20, pubescentibus; tepalis ovatis, glabris, acutis, ca. 0.5 mm longis.

A shrub or small tree very densely and softly pilose or pubescent
throughout. Branches terete, slender. Leaves pale, elliptical or ellip-
tical-ovate, densely pilose on both surfaces, membranaceous, 12 to 16 cm
long, 4 to 8 em wide, base acute, apex short-acuminate, the margins
above unequally undulate-denticulate; nerves 5 or 6 on each side of the
midrib; petiole 1.5 to 2 cm long. Inflorescence axillary, solitary, panic-
ulate or racemose-paniculate, 6 to 10 cm long, densely pilose, the
branches spreading, the lower ones about 3 cm long, the upper gradually
shorter, each bearing 1 to 3 flowers. Flowers 3 to 4 mm long, their
pedicels 1 to 2 cm long, the receptacle elliptical-ovate, chartaceous, exter-
nally densely pilose; carpels about 20, 2 mm long, pubescent ; tepals ovate,
glabrous, acute, about 0.5 mm long.

Luzon, Province of Rizal, Bosoboso, Bur. Sci. 1070 Ramos, July, 1906.
A very characteristic species, readily recognizable by its soft, dense, rather
pale pubescence, which extends to all parts of the plant.

226 MERRILL.
ROSAGE AS.
PYGEUM Gaertn.

Pygeum glandulosum sp. nov.

Arbor usque ad 18 m alta; foliis oblongo-lanceolatis vel ovato-lanceola-
tis, acuminatis, coriaceis, subtus utrinque ad basin glandulis prominenti-
bus impressis; racemis axillaribus, dense ferrugineo-pubescentibus ; flori-
bus subsessilibus, 5-meris.

A tree 6 to 18 m high. Branches glabrous, dark-reddish-brown,
lenticellate, terete, the branchlets slender, the younger ones usually more
or less pubescent. Leaves 9 to 16 ecm long, 3 to 6 em wide, oblong-
lanceolate or ovate-lanceolate, coriaceous, glabrous and shining above
except the midrib which is somewhat pubescent, the lateral nerves some-
times pubescent also, on the lower surface glabrous or slightly pubescent,
rather slenderly acuminate, the acumen blunt, often apiculate, base
acute or cuneate, with two very prominent basal glands, one on each side
of the midrib, these glands deeply impressed and protruding on the
upper surface of the leaf: nerves 5 or 6 on each side of the midrib, prom-
inent beneath, anastomosing; petioles pubescent, becoming glabrous or
nearly so, 5 mm long. Racemes densely ferruginous-pubescent, axillary,
solitary, 9 cm long or less. Flowers white, fragrant, subsessile or very
shortly pedicellate, each subtended by a small densely villous bracteole
about 2mm long. Calyx funnel-shaped, about 3.5 mm long, ferruginous-
pubescent, the lobes 5, oblong, obtuse, villous, 1.5 to 2 mm long. Petals
5, resembling the sepals but somewhat oblong-rhomboid. Stamens about
20, unequal, their filaments glabrous, 2 to 4 mm long; anthers 0.3 mm
long. Ovary ovoid, villous; style glabrous, 4 mm long; stigma disciform.
Fruit glabrous or with few scattered hairs, red, about 1 em wide, slightly
compressed, about 8 mm long; seeds with pronounced flavor like those of
Prunus.

Luzon, Province of Rizal, Bosoboso, For. Bur. 3092 Ahern’s collector, May,
1905, also from the same province, For. Bur. 1134, 3342 Ahern’s collector; For.
Bur, 10007, 10037 Curran, February, 1908; Merrill 1707, March, 1903; Loher
2223, November, 1892: Province of Zambales, Bur. Sci. 5029 Ramos, December,
1907; Hallier, January, 1904; For. Bur. 381 Maule, March, 1904: Province of
Bataan, Lamao River, Williams 529, 642, January, Mareh, 1904; For. Bur. 6317
Curran, February, 1907; Whitford 8, 84, April, 1904; For. Bur. 49 Barnes,
August, 1903; For. Bur. 2064, 2366 Borden, 1905; For. Bur. 2629 Meyer, February,
1905; Blmer 6697, November, 1904; Dinalupijan, Merrill 1537, January, 1903:
Province of Laguna, Santa Maria Mavitac, For. Bur. 10063 Ourran, February,
1908: Province of Benguet, Sablan, Blmer 6110, April, 1904. Minporo, For, Bur.
38719, G8I8 Merritt, March, 1906, 1907. Without locality, Cuming 797, 1836-40.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 227

A very common and widely distributed species in Luzon, confused with
Pygeum latifolium Migq., and distributed as such, readily recognized, however, by
its subsessile flowers and very prominent glands at the base of the leaf, which
strongly protrude on the upper surface. 'T., Lago, Gupit.

Pygeum preslii nom. nov.

Germaria latifolia Pres] Epim. Bot. (1849) 221; Miq. Fl. Ind. Bat. 1* (1855)
401.

Pygeum latifolium Vidal Phan. Cuming. Philip. (1885) 111, Rev. Pl. Vase.
Filip. (1886) 122; Merr. in Philip. Journ. Sei. 1 (1906) Suppl. 60 (in part),
non Pygeum latifolium Mig. 1. ¢. 361.

Pygeum arboreum F.-Vill. Nov. App. (1883) 76, excel. syn.; Vidal Sinopsis
Atlas (1885) ¢. 46, f. B, non Endl.

This species is common and widely distributed in the Philippines, and is allied
to the preceding, readily distinguished, however, by its rather long-pedicelled
flowers and basal leaf glands which are not prominent and not impressed, being
scarcely visible on the upper surface of the leaf and not at all protruding as
in Pygeum glandulosum Merr. It has been considered as Pygeum latifoliwm Miq.,
but judging from the description of that species, the Philippine plant is different.
Unfortunately both Presl and Miquel used the same specific name for what are
apparently two distinct species, Miquel publishing his Pygeum latifoliwm without
any reference to the earlier Germaria latifolia Presl, and not recognizing that
Presl’s species was in reality a true Pygeum. The use of the specific name
latifoliwm under Pygewm, precludes the possibility of transferring Presl’s earlier
name, and accordingly the above new name for the Philippine plant is here
proposed.

Pygeum clementis sp. nov.

Arbor glabra; foliis oblongis, acuminatis, usque ad 20 cm longis, nervis
utrinque ca. 10, subtus utrinque ad basin glandulis concavis impressis ;
fructibus subcompresso-globosis, 1.5 ad 2 cm diametro.

A tree, glabrous or nearly so throughout, 10 to 15 m high. Branches
dark-reddish-brown, glabrous, terete, lenticellate. Leaves oblong, coria-
ceous, dull or shghtly shining above, 12 to 20 cm long, 4.5 to 8 em wide,
entire, acuminate, base rounded or subacute, with a small but prominent
gland on each side of the midrib at the base, protruding on the upper
surface ; nerves about 10 on each side of the midrib, prominent beneath,
anastomosing; petioles 5 to 7 mm long. Flowers not seen. Racemes
axillary, solitary, in fruit 4 to 5 em long, glabrous. Fruit glabrous,
subcompressed-globose, 1.5 to 2 em wide, 1 to 1.3 em long, the pedicels
7 to 10 mm long.

Mrinpanao, Lake Lanao, Camp Keithley, Mrs. Clemens 760, 966, September,
1906, March, 1907, both specimens in fruit.

A species allied to Pygeum preslii Merr., and P. glandulosum Meryr., differing
from both in having twice as many lateral nerves, and in its much larger fruit.

DIBe a: MERRILL.
LEGUMINOSA¢.
ADENANTHERA Linn.

Adenanthera intermedia nom. noy.

Mimosa virgata Blanco Fl. Filip. (1837) 7387, non Linn.

Mimosa punctata Blanco 1. ¢. ed. 2 (1845) 508; ed. 3, 3: 139, non Linn.

Adenanthera pavonina ¥.-Vill. Nov. App. (1883) 73; Vidal Sinopsis Atlas
(1883) 24, t. 44, f. B; Rev. Pl. Vase. Filip. (1886) 119; Phan. Cuming. Philip.
(1885) 110; Perk. Frag. Fl. Philip. (1904) 7; Merr. in Philip. Journ. Sci. 1
(1906) Suppl. 62, non Linn.

Luzon, Province of Isabela, for. Bur. 6645 Klemme, March, 1907: Province of
Zambales, Subic, Merrill 1977, April, 1903; For. Bur. 6086 Aguilar, January, 1907:
Province of Pangasinan, Libtong, Alberto 51, March, 1904: Province .of Bataan,
Lamao River, Merrill 38863, August, 1904; For. Bur. 1496 Ahern’s collector, Au-
gust, 1904; For. Bur. 1599, 2345 Borden, August, 1904, January, 1905: Bur. Sci.
1617 Foaworthy, October, 1906; Williams 230, November, 1903; Whitford 1256,
1286, May, 1905: Province of Rizal, For. Bur. 3164 Ahern’s collector, June, 1905;
Province of Sorsogon, Elmer 7315, November, 1905. Minpanao, Lake Lanao,
Camp Keithley, Mrs. Clemens 336, March, 1906.

A species widely distributed in the Philippines, previously confused with
Adenanthera pavonina Linn., to which it is closely allied, differing constantly in
its seeds being about half jet-black and half bright red as in A. bicolor Moon.
The leaflets are quite different from those of A. bicolor Moon, obtuse, elliptical or
oblong-elliptical, as in A. pavonina Linn. The species seems to have the vegetative
characters of Adenanthera pavonina Linn., and the seed characters of A. bicolor
Moon.

PITHECOLOBIUM Mart.

Pithecolobium scutiferum (Blanco) Benth. in Lond. Journ. Bot. 3 (1844)
211; Miq. Fl. Ind. Bat. 17 (1855) 39.

Mimosa scutifera Blanco Fl. Filip. (1837) 735; ed. 2 (1845) 507; ed. 3, .
3.) 138: :

Pithecolobium lobatum ¥.-Vill. Nov. App. (1883) 75; Merrill in For. Bur. Bull.
1 (1903) 23; Philip. Journ. Sci. 1 (1906) Suppl. 62; Perk. Frag. Fl. Philip.
(1904) 4; Vidal Phan. Cuming. Philip. (1885) 111; Rev. Pl. Vase. Filip. (1886)
120; Ceron Cat. Pl. Herb. (1892) 72, in part, non Benth.

PHILIPPINES, without locality, Cuming 475. Luzon, Province of Cagayan, For.
Bur. 4294 Klemme, June, 1906: Province of Bataan, Lamao River, Whitford 1257,
May, 1905; For. Bur. 726, 1687, 1933 Borden, For. Bur. 1441 Ahern’s collector,
For Bur. 6366 Curran: Province of Rizal, Bosoboso, For. Bur. 2007 Ahern’s col-
lector, November, 1904; San Mateo, no. 117, Decades Philip. For. Fl.: Province of
Laguna, Los Bafios, Williams 2053, January, 1905; Hlmer 8310, April, 1906: Prov-
ince of Tayabas, For. Bur. 6069 Kobbe, December, 1906; Guinayangan, Merrill 2013,
April, 1903: Province of Albay, I’rancisco s. n., 1901. Minporo, For. Bur. 4073,
4099, 6211, 8570 Merritt; Pinamalayan, Merrill 2153, May, 1903; Puerto Galera,
Merrill 3321, October, 1903; Pola, Merrill 2212, 2376, 2474, May-June, 1903;
Bongabon River, Whitford 1422, February, 1906. Trcao, For. Bur. 1067 Clark,
May, 1904. Maspnare, Whitford 1685, October, 1906. GuIMARAS, Por, Bur, 233
Gammill, January, 1904,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 229

A widely distributed endemic species, correctly identified by Bentham with
Blanco’s Mimosa scutifera and transferred by him to Pithecolobium, but later *
considered by him to be the same as Pithecolobium lobatum Benth., and reduced
to that species. It is, however, very distinct from Pithecolobium lobatum Benth.,
both in its very deeply lobed pods and in the venation of the leaflets. Univer-
sally known in the Philippines as Anagap.

Pithecolobium pauciflorum Benth. in Lond. Journ. Bot. 3 (1844) 212.

This species was based on flowering specimens, Cuming 1854, collected in the
Philippines, without locality given, and like the preceding species later reduced
by Bentham to Pithecolobium lobatum Benth. It does not, however, appear to be
the same as that species. Cuming’s specimen is well matched by Bolster 286,
Surigao, Mindanao, April, 1906, in fruit. I have seen no typical Pithecolobium
lobatum Benth. from the Philippines.

ENTADA Adans.

Entada parvifolia sp. nov.

Scandens, subglabra; foliis bipinnatis, pinnis 2-jugatis; foliolis utrin-
que 8, oblongis, mucronatis, ca. 1.5 cm longis, 3 ad 5 mm latis; spicis
ca. 15 cm longis, plus minus hirsutis; leguminibus glabris, 10 ad 25 em
longis, 4 ad 5.5 cm latis, seminibus nitidis, 2 em diam.

Scandent, the branches slender, terete, brownish, glabrous. Leaves
bipinnate, the rachis about 6 cm long, slightly pubescent when young,
the terminal tendril 6 cm long or more; pinnae 5 to 8 cm long, the
rachis slightly pubescent; leaflets oblong, chartaceous, about 8 pairs,
about 1.5 em long, 3 to 5 mm wide, the apex obtuse, strongly mucronate,
margins revolute, nerves obscure; petiolules very short. Spikes about
15 cm long, many flowered, the rachis somewhat ferruginous-hirsute, the
bracteoles lanceolate, acuminate, about 1.5 mm long. Flowers subsessile,
the calyx glabrous, small, campanulate, about 1.8 mm in diameter,
obscurely 5-toothed. Petals 5, glabrous, oblong, equal, 3.5 mm long, 1
mm wide. Stamens 10; filaments about 5 mm long; anthers 0.8 mm
long. Ovary glabrous, narrowly oblong, 2 mm long; style about 4 mm
long. Pod narrowly oblong, compressed, 10 to 25 cm long, 4 to 5.5 em
wide, somewhat acuminate at both ends, the valves shining, firmly coria-
ceous, dark-brown, glabrous; seeds shining, round, compressed, about
2 cm in diameter.

Luzon, Province of Zambales, Bur. Sct. 4810, 5067 Ramos, December, 1907;
Haller, January, 1904.

A species well characterized by its small leaflets and comparatively small
pods; very distinet from the widely distributed Hntada scandens Benth. Like

Entada scandens Benth., the stem of this species yields a substitute for soap
which is used by the natives of Zambales. Z., Hinagui.

1Trans. Linn. Soc. 30 (1875) 575.

230 MERRILL.
BAUHINIA Linn.

Bauhinia copelandii sp. noy. § Phanera.

Scandens; ramulis, subtus foliis, racemis, calycibusque plus minus pu-
bescentibus; foliis ovatis vel oblongo-ovatis, integris, 5-nerviis, 7 ad 9
cm longis, basi cordatis, apice breviter acuminatis, acuminibus retusis ;
floribus racemosis ; staminibus fertilibus 3, sterilibus 3.

Scandent, the branches and branchlets terete, slightly pubescent.
Leaves ovate to oblong-ovate, 7 to 10 cm long, 4 to 6 cm wide, entire,
not cleft, subcoriaceous, shining on both surfaces, glabrous above, beneath
with few appressed hairs, the base broad, cordate, the apex shortly and
broadly acuminate, the acumen retuse; nerves 5, prominent; petioles 1
em long or less, glabrous or slightly pubescent. Racemes terminal,
somewhat pubescent, the rachis 1.5 em long or less, the pedicels slender,
about 2.5 em long. Calyx-tube cylindrical, about 7 mm long, 3 mm in
diameter, pubescent, the lobes equal, lanceolate or oblong-lanceolate,
shortly acuminate, pubescent outside, about 1 cm long, 3 mm wide.
Petals obovate to oblong-obovate, slightly appressed-pubescent externally,
about 1.5 cm long, 1.3 em wide, short-clawed. Stamens 6, three fertile,
three sterile.: Ovary and style somewhat pubescent; stigma capitate.
Fruit unknown.

MINDANAO, District of Davao, Todaya, Copeland 1429, October, 1904.

Among the Philippine species most closely allied to Bauhinia leptopus Perk.,
but with much larger flowers; apparently most closely allied to Bauhinia corni-
folia Baker, of the Malayan Peninsula, but sufficiently distinct, ex deser.

Bauhinia subglabra sp. nov. § Phanera.

Scandens, glabra; ramulis pruinosis, nigricantibus nitidis; foliis sub-
coriaceis, glabris, 5 ad 7 longis, usque ad medium lobatis, lobis rotunda-
tis; nervis ca. 11; racemis glabris; petalis oblongo-lanceolatis vel elliptico-
lanceolatis, unguiculatis; staminibus fertilibus 3; ovario tomentoso,

Scandent, 3 to 5 m high, glabrous throughout except the tomentose
ovary and style, and the petals which are slightly pubescent outside.
Branches terete, lenticellate, gray, the branchlets terete, slender, black-
ish, somewhat shining. Leaves subcoriaceous, slightly shining, 5 to
7 em long, 5.5 to % em wide, suborbicular, the base cordate, cleft
nearly to the middle, the sinus very narrow, the lobes rounded;
nerves about 11, prominent; petioles slender, 4 to 5 cm long. Racemes
terminal, glabrous, the rachis 3 to 6 em long, black; pedicels 2 to 3 em
long; bracteoles none or caducous. Calyx cylindrical, about 1.4 em long,
3 mm thick, the lobes glabrous, reflexed, narrowly oblong-lanceolate,
about 2 em long, 3 mm wide. Petals oblong-lanceolate to elliptical-
lanceolate, nearly 3 em long, long-clawed, slightly pubescent on the

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. Qai

back, acuminate. Stamens 3, fertile; anthers 12 mm long. Ovary and
style densely brown-tomentose; stigma capitate. Fruit unknown.

PALAWAN, Iwahig, Bur, Sci. 821 Foxworthy, April 30, 1906.

A species well characterized by being entirely glabrous, except the ovary, styles
and petals.

Bauhinia dolichocalyx sp. nov. § Lysiphyllum.

Arbor ca. 10 m alta; foliis ovatis, integris, basi cordatis, apice bre-
viter acuteque acuminatis, 10 ad 14 em longis, nervis 7; inflorescentiis
terminalibus, densissime fusco-tomentosis ; calycis segmentis anguste lan-
ceolatis, 4 ad 5 cm longis; staminibus fertilibus 10; lezuminibus ca. 20
em longis, 7 cm latis, glabris; seminibus 2, compressis, 3.5 em diam.

A tree about 10.m high. Branchlets terete, lenticellate, somewhat
brown-pubescent. Leaves subcoriaceous, ovate, 10 to 14 em long, 11 em
wide or less, entire, the base broad, usually prominently cordate, the apex
shortly and sharply acuminate, glabrous and shining above, somewhat
pubescent along the nerves beneath; nerves 7, prominent; petioles 2 to
3 cm long, pubescent. Inflorescence terminal, short, the flowers race-
mosely disposed, densely brown-appressed-tomentose. Calyx tube and
pedicels not differentiated, 7 to 8 em long, gradually thickened upwards,
densely brown-tomentose, the calyx lobes narrowly lanceolate or linear-
lanceolate, 4 to 5 cm long, about 5 mm wide, densely brown-pubescent
outside, subequal. Petals about 5 em long, glabrous or nearly so, long-
clawed. Fertile stamens 10. Ovary glabrous. Pods woody, about 20
cm long, 7 cm wide, glabrous, each with two rounded compressed shining
seeds about 3.5 cm in diameter.

Luzon, Province of Batangas, for. Bur. 7756 Curran & Merritt, October 28,
1907.

A striking species, well characterized by its entire leaves, undifferentiated
calyx tube and pedicel, very long calyx lobes, 10 fertile stamens, and rather

large pods.
MUCUNA Adans.

Mucuna mindorensis nom. nov.
Mucuna acuminata Mery. Philip. Journ. Sei. 1 (1906) Suppl. 196, non Grah.
In describing the above species the fact was overlooked that the specific name
acuminata had already been used. Accordingly the above new name is here
proposed for the species.
GLYCINE Linn.

Glycine warburgii (Perk.) comb. nov.

Pueraria warburgii Perk. Frag. Fl. Philip. (1904) 87.

MInpANAO, Taumo, Warburg 14664, in Herb. Berol. (type); District of
Davao, DeVore & Hoover 368, May, 1903; Santa Cruz, Williams 2953, June, 1905:
Province of Surigao, Baganga, Merrill 5430, October 6, 1906.

The plant is apparently Glycine and not Pueraria, and seems to be allied to
G. javanica Linn. I have examined the type of Puweraria warburgii in Herb.
Berol., and find that the other specimens here cited closely match it. According
to Perkins 1. ¢., a closely allied, if not identical form, is found in Celebes.

232 MERRILL.
MEZONEURUM Dest. ‘

Mezoneurum mindorense sp. noy.

Scandens; foliis bipinnatis, usque ad 35 cm longis, rhachidibus
postice aculeis geminis ternisve recurvis armatis; pinnis ca. 12, 8 ad
10 em longis; foliolis 8- ad 12-jugatis, oblongo-ellipticis, glabris, 1 ad
1.5 cm longis; paniculis terminalibus; leguminibus brunneis, nitidis,
ca. 7 em longis, monospermis, alis 5 mm latis.

Seandent, nearly glabrous throughout, the branches terete, dark-
reddish-brown, with scattered stout recurved spines about 2.5 mm long.
Leaves 35 em long or less, bipinnate, the rachis with short but stout
recurved spines in pairs, rarely in threes at the insertion of the pinnae;
pinnae 12 or less, 8 to 10 em long, the rhachis shghtly pubescent; leaf-
lets 8 to 12 pairs, oblong-elliptical, glabrous, submembranaceous, 1 to 1.5
em long, 5 to 7 mm wide, paler beneath, the apex rounded, minutely
apiculate, rarely shghtly emarginate, the base acute or obtuse; nerves
obscure; petiolules very short. Panicles terminal, in fruit 30 cm long
or more, the branches somewhat pubescent. Flowers unknown. Pods
glabrous, shining, about 7 cm long, about 2.2 em wide in the middle,
coriaceous, brown, somewhat semilunar in outline, the winged margin
nearly straight, the other regularly curved, giving nearly the outline
of a sector of a circle, the wing 5 mm wide, base and apex acute; seed
solitary, in the middle of the pod, flat, circular in outline, 7 mm in
diameter (immature).

Minporo, Pinamalayan, For. Bur. 5383 Merritt, October, 1906, in thickets
near sea level.

Var. inerme var. nov.

Differt a typo foliorum rhachidibus inermibus.

Mrnporo, Bulalacao, Bur. Sci. 1514 Bermejos, August 30, 1906.

ERYTHROXYLACE/D.
ERYTHROXYLUM P. Browne.

Erythroxylum cuneatum (Wall.) Kurz in Journ. As. Soc. Beng. 43° (1874)
35; Schulz in Pflanzenreich 29 (1907) 146.

Erythroaylum burmanicum Griff. Not. Pl. Asiat. 4 (1854) 468. ¢. 5871, f. 3;
Merrill in Forestry Bur. Bull. 1 (1903) 25.

Luzon, Province of Rizal, Loher s. n., May, 1904; Bosoboso, Bur. Set. 1023
Ramos, June, 1906: Provinee of Bulacan, Fors Bur. 7158 Curran, June, 1907:
Provinee of Camarines Sur, Pasacao, Ahern 292.

Tenasserim to the Malay Peninsula, Sumatra and Java. T., Saling, Manambo.

Erythroxylum platyphyllum sp. nov. § Coelocarpus.
Differt H. cuneato pedicellis brevioribus, ca. 2 mm longis, foliis ma-
joribus, usque ad 11 em longis, 6 em latis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 233

A small tree, the branches terete, reddish-brown, glabrous. Leaves
elliptical, submembranaceous, 6 to 11 cm long, 3 to 6 cm wide, the apex
broad, rounded, the base acute, shining above, paler beneath, glabrous ;
nerves about 10 on each side of the midrib, not prominent; petioles 5
mm long or less; stipules-7 mm long, lanceolate, acuminate, two-keeled,
puberulent on the keels. Flowers axillary, solitary or in pairs, their
pedicels 2 mm long. Calyx lobes 1.2 mm long, ovate, acuminate.
Petals oblong, obtuse, 3.5 mm long, 1.5 mm wide, the ligule nearly one-
half as long as the petal. Staminal tube 1.5 mm long, the stamens 10,
the filaments unequal, about 1.5 mm long; anthers 0.4 mm long. Ovary
glabrous, the style about 3 mm long, cleft into three parts above (short
styled flowers not seen). Fruit oblong, 8 mm long, 4.5 mm wide.

Mrinpanao, District of Zamboanga, Port Banga, Por. Bur. 9285 Whitford &
Hutchinson, January 2, 1908, growing along the margins of mangrove swamps.
The second species of the family for the Philippines.

RUTACEZE.
LUVUNGA Ham.

Luvunga philippinensis sp. nov.

Scandens, inermis ; foliolis oblongis vel oblongo-ellipticis, subcoriaceis,
usque ad 25 cm longis, 10 em latis; floribus in cymis racemosis paucifioris
lateralibus dispositis; calycibus cupulatis, truncatis; petalis 3 vel 4,
oblongis, ca. 1 cm longis, ovario 4-loculare.

A scandent shrub without spines, glabrous throughout, the branches
gray or brownish. Petioles terete, 9 to 13 cm long, greenish ; leaflets
3, their petiolules stout, 5 mm long or less, 15 to 25 cm long, 6 to 10
cm wide, subcoriaceous, shining, paler beneath, oblong to oblong-elliptical,
the apex acuminate, the acumen blunt or retuse, base acute, margins
entire; nerves 8 to 10 on each side of the midrib, irregular, somewhat
prominent beneath, anastomosing. Inflorescence of small, usually 3-
flowered, racemose cymes, borne on the branches below the leaves or in
the leaf-axils, 1.5 cm long or less; pedicels 4 to 5 mm long. Flowers
white, fragrant. Calyx cup-shaped, about 4 mm in diameter, truncate.
or very obscurely 5-toothed, its stipe 2 mm long. Petals 3 or 4, oblong,
obtuse, imbricate, 9 to 10 mm long, 3.5 to 4.5 mm wide. Stamens
10, free; filaments stout, swollen, about 6 mm long; anthers erect, oblong,
3 mm long. Ovary oblong, 4-celled, quadrangular in cross section,
about 3 mm long, 1.2 mm thick, scarcely narrowed into the stout style
which is about as long as the ovary; stigma capitate. Fruit unknown.

MinpDANAO, District of Zamboanga, Port Banga, For. Bur. 9104, 9267 Whitford
& Hutchinson, December, 1907, in forests 30 to 40 m above sea level. The first
representative of the genus to be found in the Philippines.

234 MERRILL.
MELICOPE Forst.

Melicope curranii sp. nov. § Hntoganum.

Arbuscula glabra; foliis oblongo-obovatis vel oblongo-lanceolatis, acu-
minatis, sunplicibus; paniculis axillaribus, usque ad 4 cm longis, pauci-
floris; floribus 4-meris, 2 ad 2.5 mm longis.

A shrub, glabrous throughout. Branches slender, brown, terete, the
branchlets greenish. Leaves simple, membranaceous, oblong-obovate to
oblong-lanceolate, apex acuminate, base cuneate, 11 to 17 em long, 4
to 6 cm wide, shining; nerves 8 or 9 on each side of the midrib, irregular,
somewhat prominent beneath, distant, anastomosing; petiolule about 1
em long, the petiole 2 to 5 cm long, geniculate at the joint with the
petiolule. Panicles small, axillary, peduncled, 4 cm long or less, the
branches short, spreading. Flowers 4-merous, greenish-white, their ped-
icels 2 mm long. Calyx short, 4-toothed, the teeth acute. Petals
oblong, acute, 2 to 2.5 mm long, 1 mm wide, valvate and with an ap-
pendage at the apex inside. Stamens 8, about 1 mm long. Ovary
4-lobed; style short; stigma 4-lobed.

Luzon, Province of Tayabas, For. Bur. 9663 Curran, March, 1908.

In general appearance resembling Melicope luzoniensis Engl., but characterized
at once by its simple leaves. The same form has also been collected by Vidal
(Herb. Kew).

CHISOCHETON Blume.

Chisocheton curranii sp. nov. § Huchisocheton, Paniculati.

Foliis modice petiolatis, 2- vel 3-jugatis, petiolis rhachide petiolulisque
fulyo-olivaceo-pubescentibus ; foliolis oppositis, elliptico-oblongis vel ellip-
tico-ovatis, basi acutis, apice breviter acuminatis, costa nervisque utrinque
hirsutis: paniculis axillaribus, angustis, 9 ad 15 em longis, pauce breviter
ramosis, dense pubescentibus; floribus breviter pedicellatis; petalis 5,
pubescentibus, ca. 16 mm longis; tubo cylindrico, 5-laciniato; ovario
2-loculare.

A small tree about 5 m high, the branches appressed-brownish-pubes-
cent. Leaves about 20 cm long, the rachis, petiolules, midribs, and
nerves on both sides of the leaflets densely olivaceous-brownish-pubes-
cent or hirsute; leaflets subcoriaceous, opposite, 2 or 3 pairs, 8 to
11 cm long, 3 to 4.5 em wide, apex short acuminate, base acute, the
nerves and midrib on both surfaces pubescent or hirsute; nerves about 7
on each side of the midrib, prominent, the reticulations distinct; petio-
lules 5 to 7 mm long. Panicles pubescent, axillary, narrow, 9 to 15 em
long, the branches very short, spreading or ascending, few, 1 to 2 em
long. Calyx pubescent, cup-shaped, 4 to 5 mm long, the teeth short,
rounded. Petals yellowish-white, about 16 mm long, 2 mm wide, pubes-
cent outside, glabrous within. Staminal tube 13 to 14 mm long, eylin-
drical, free, appressed-hirsute on both sides, the apex 5-lobed, the lobes

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 235

narrowly oblong, obtuse, 2.5 mm long, 0.7 mm wide. Stamens 5, the
anthers sessile, 2.5 mm long, alternating with the teeth. Disk none.
Ovary hirsute, narrow, 2-celled; style hirsute, about 10 mm long. Fruit
globose, brown, 5 to 6 cm in diameter, the seeds 2.5 to 3 cm long.

Luzon, Province of Benguet, Baguio, For. Bur. 4865, 4923 Curran, August,
1906, in ravines in the limestone region at an altitude of about 1,500 m.

AGLAIA Lour.

Aglaia palawanensis sp. nov. § Hearma.

Arbuscula ca. 5 m alta, ramis petiolis rhachide paniculisque dense fer-
rugineo-stellato-pubescentibus ; foliis imparipinnatis, 2-jugatis, usque ad
20 em longis; foliolis elliptico-ovatis, elliptico-lanceolatis vel ovato-lan-
ceolatis, acuminatis, subcoriaceis, pallidis, utrinque glabris, costa subtus
stellato-pubescente excepta; paniculis pyramidato-ramosis, multifloris,
foliis subaequantibus; floribus pedicellatis; calycibus dense stellato-pu-
bescentibus, profunde 5-dentatis; petalis 5.

A shrub or small tree about 5 m high, the branches, branchlets, petioles,
rachis, petiolules, midrib of the leaflets beneath and the inflorescence
rather densely stellate-ferruginous-pubescent. Branches slender, terete,
brownish. Leaves 20 cm long or less, usually 2-jugate, the rachis, in-
cluding the petiole, about 6 cm long; leaflets subcoriaceous, pale when
dry, somewhat shining, opposite, the terminal one slightly larger than the
lateral ones, elliptical-ovate to ovate-lanceolate, the apex acuminate, the
base acute, 7 to 11 em long, 2 to 3.5 em wide; nerves about 9 on each
side of the midrib, rather distinct beneath, the reticulations faint,
petiolules 5 to 7 mm long, that of the terminal leaflet about 1 cm long.
Panicles about as long as the leaves, the lower branches 5 to 8 cm long,
the upper ones gradually shorter. Flowers white or pale-yellow, pedicel-
late. Calyx densely ferruginous-stellate-pubescent, 1 mm long, deeply
5-cleft, the lobes about 0.7 mm. long, narrowly ovate, acute. Petals 5,
free, concave, glabrous, ovate, obtuse, about 1.2 mm long. Staminal
tube truncate, broad, 0.5 mm long, glabrous. Anthers 5, inserted on
the margin of the tube, exserted, broadly triangular-ovate, 0.5 mm long.

PALAWAN, Victoria Peak, Bur. Sci. 689 Foxworthy, March, 1906, on steep
forested slopes along streams at an altitude of about 1,000 m.

Aglaia affinis sp. nov. § Hearnia.

Foliis breviter petiolatis, 2-jugatis ; foliolis oppositis, elliptico-oblongis,
pallidis, subcoriaceis, supra glabris, subtus leviter ferrugineo-lepidotis,
apice acuminatis, nervis utrinque ca. 10; paniculis foliis subaequantibus,
pyramidato-ramosis, dense ferrugineo-lepidotis; floribus subsessilibus ;
ealycibus dense lepidotis.

A tree about 15 m high, the branches brownish, glabrous, striate, not
lenticellate, the younger parts somewhat lepidote. Leaves 12 to 20 cm

236 MERRILL.

long, the rachis and petiolules rather densely ferruginous-lepidote; leaf-
lets subcoriaceous, 2-jugate, 7 to 10 em long, 2.4 to 5 em wide, elliptical-
oblong, pale, glabrous above, beneath somewhat ferruginous-lepidote, base
acute, equal, apex acuminate ; nerves about 10 on each side of the midrib;
petiolules 2 to 5 mm long. Panicles terminal, pyramidal, about as long
as the leaves, densely ferruginous-lepidote throughout, branched from
near the base, the lower branches about 7 cm long, the upper ones grad-
ually shorter, the branchlets densely many-flowered. Flowers yellowish,
fragrant, about 1 mm long, the buds globose, subsessile. Calyx 5-toothed,
lepidote. Petals 5, glabrous, about 1 mm long. Staminal tube about
0.8 mm long, crenate, glabrous, bearing on its margin 5 sessile anthers.

BALaBac, Bur. Sci. 446 Mangubat, March, 1906.

A species apparently allied to Aglaia cumingiana Turez., and to A. harmsiana
Perk., differing from the former in its densely lepidote inflorescence, nearly sessile
flowers which are densely disposed, and its acuminate leaflets, and from the
latter by having only two pairs of shorter leaflets, shorter panicles, and dif-
ferent indumentum, the branches and leaves in the species above described not
being at all stellate-tomentose-lepidote, the scales being appressed and entire or
only very minutely ciliate.

EKUPHORBIACEZ.
OMPHALEA Linn.

Omphalea philippinensis sp. noy.

Scandens, inflorescentiis exceptis glabra; foliis coriaceis oblongo-lan-
ceolatis, acuminatis, nitidis, basi supra 2-glandulosis; floribus pedicel-
latis, minutis; fructibus dehiscentibus, 2.5 ad 3 em longis.

A large woody vine, glabrous except the inflorescence. Branches
terete, somewhat rugose-striate when dry, glabrous, light-gray or yellowish.
Leaves alternate, oblong-lanceolate, 12 to 22 em long, 3 to 7 em wide,
coriaceous, glabrous, shining, pale in drying, the margins entire, some-
what recurved, apex rather abruptly acuminate, the acumen 1 em long
or less, blunt, the base acute or obtuse; nerves about 9 on each side of
the midrib, prominent beneath, curved, the reticulations distinct on
both surfaces; petioles stout, 1.5 to 3.5 em long, with two prominent
glands on the upper surface at the juncture with the leaf. Inflorescence
axillary, paniculate, about 40 em long (or longer?), somewhat ferrugi-
nous-hirsute, the branches stout, spreading, the flowers many, pedicelled,
fasciculate, the pedicels slender, thickened upwards, 6 to 7 mm long.
Buds globose, 1.5 to 2 mm in diameter. Sepals 5, imbricate, glabrous,
obovate or orbicular, about 2mm long. Petals wanting. Anthers sessile.
Pistillate flowers not seen. Fruit dehiscent, 2.5 to 3 em long, the valves
twisting in dehiscence, firmly coriaceous, glabrous, gray, the seeds

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. Zod

glabrous, shining, oyoid, 1.5 to 2 cm long, somewhat flattened on the two
inner sides, apparently three in each fruit.

Luzon, Province of Rizal, Antipolo, Merrill 1716, March, 1903; Santander,
Bur. Sci. 3270 Ramos, June, 1907: Province of Tayabas (Infanta), Whitford
821, September, 1904. The same species is apparently represented by the follow-
ing specimens in Herb. Kew: Cuming 1468; Vidal 1712, 2380, 3875, and Loher
5218, all from the Philippines.

The only representative of the genus known from the Philippines, about 12
species being known from tropical America, one from Queensland and one from
Madagasear. The material available is not sufficient to draw up a complete
diagnosis, flowers being immature and fruit opened, so that it is impossible to
describe the entire fruit.

AQUIFOLIACEE.

ILEX Linn.

Ilex gracilipes sp. nov.

Arbor parva, 3 ad 4 m alta, glabra; foliis oblongo-ovatis vel elliptico-
ovatis, membranaceis, acuminatis, margine apiculato-denticulatis ; floribus
3- ad 5-meris, solitariis vel fasciculatis, axillaribus, longe pedunculatis ;
pedunculis ca. 1 em longis, fructiferis longioribus.

A small tree 3 to 4 m high, glabrous, the branches and branchlets
slender, terete, reddish-brown, lenticellate. Leaves oblong-ovate to ellip-
tical-ovate, membranaceous, 2.5 to 5 em long, 1 to 3 em wide, opaque or
obscurely minutely punctate, shining, base acute, the apex rather long-
acuminate, the acumen apiculate, the margins regularly apiculate-den-
ticulate; nerves 4 or 5 on each side of the midrib, not prominent,
anastomosing ; petioles slender, 5 mm long or less. Flowers solitary or
in 2- to 4-flowered fascicles in the axils of the leaves, long-pedicelled,
the pedicels about 1 cm long, much elongated in fruit. Calyx about 2
mm in diameter, 3- to 5-lobed, the lobes orbicular-reniform, 0.5 mm
long the margins shortly obscurely ciliate. Corolla lobes 3 to 5, im-
bricate, orbicular, rounded, about 1.5 mm long, the tube 0.5 mm long.
Stamens 3 to 5, inserted on the corolla and alternating with the lobes,
the filaments and anthers each 0.8 mm long. Ovary usually 5-celled, with
a single pendulous ovule in each cell; stigma capitate. Fruit subglobose
or ovoid, 5 to 6 mm long, with about 15 longitudinal ridges, usually
with 5 nutlets, the peduncles 2 to 3 em long. :

Luzon, Province of Benguet, Williams 1047, May, 1904; Bur. Sci. 2839 Mearns,
April, 1907; Elmer ; Loher 5129: Province of Zambales, Mount Tapulao, For, Bur.
8060 Curran & Merritt, December, 1907: Province of Bataan, Mount Mariveles,
Whitford 414: Province of Rizal, Bosoboso, Bur. Sci. 2115, 2677 Ramos, February,
May, 1907.

A very characteristic species readily recognizable by its solitary or fascicled,
long-pedicelled flowers, very long-pedicelled fruits and membranaceous leaves which
are apiculate-denticulate. The flowers are sometimes 3- to 5-merous on the
same plants.

238 MERRILL.
CELASTRACE/®.
MICROTROPIS Wall.

Microtropis curranii sp. nov.

Arbor parva, glabra, 4 ad 9 m alta; foliis coriaceis, nitidis, subtus
pallidis, elliptico-ovatis, 4 ad 7 em longis, breviter obtuseque acuminatis,
basi acutis, margine revolutis ; cymis axillaribus, solitariis vel fasciculatis,
ca. 1.5 cm longis, 3-floris; floribus 4-meris; fructibus cylindraceis,
elongatis, 1 ad 1.5 cm longis.

A small tree, glabrous throughout, 4 to 9 m high, the branches and
branchlets terete, dark-reddish-brown or nearly black when dry. Leaves
coriaceous, shining, pale beneath, elliptical-ovate, 4 to 7 em long, 1.5
to 4.5 cm wide, the apex short and bluntly acuminate, the base acute,
margins entire, revolute; nerves 5 or 6 on each side of the midrib, not
prominent, the reticulations indistinct; petioles 1 em long or less.
Cymes axillary, solitary or two or three in each axil, 1.5 em long or
less, the peduncles about 1 cm long, each cyme usually 3-flowered.
Flowers 4-merous, the sepals orbicular, rounded, the two outer ones 2
mm long, the two inner ones somewhat petaloid, 3 mm long. Corolla
lobes 4, obovate or elliptical-obovate, rounded, about 3 mm long, united
below. Stamens 4, attached to the corolla and alternating with its
lobes, the filaments short, the anthers 0.5 mm long, broader than long.
Ovary ovoid, glabrous. Fruit cylindrical, elongated, yellow when fresh,
dark-reddish-brown when dry, short-apiculate, 1 to 1.5 cm long, about
6 mm thick, 1-celled, dehiscent, the pericarp coriaceous.

Luzon, Province of Benguet, Mount Tonglon, For. Bur. 4966, 4970 Curran,
August, 1906; Williams 1300, October, 1904: Provinee of Zambales, Mount Ta-
pulao, For. Bur. 8071 Curran & Merritt, December, 1907; Bur. Sci. 4698, 5026
Ramos, December, 1907.

The first species of the genus to be found in the Philippines, about 9 species
being previously known, extending from British India to Ceylon, Cochin China and
Java. The species here described differs from all the others in the genus by its
4-merous flowers.

EUONYMUS Linn.

Euonymus philippinensis sp. noy.

Buonymus timorensis Turez. in Bull. Soc. Nat. Mose. 31 (1858) 447; Lawson
in Hook. f. Fl. Brit. Ind. 1 (1875) 610, saltem pro parte; F.-Vill. Nov. App.
880) 46; Vidal Phan. Cuming. Philip. (1885) 103; Ceron Cat. Pl. Herb. (1892)

(1
47, non Zipp.

Arbor parva, glabra, 3 ad 8 m alta; foliis subcoriaceis, nitidis, ellip-
ticis, oblongo-ellipticis, vel anguste obovato-oblongis, basi acutis, apice
acuminatis, integris vel versus apicem minute obscureque denticulatis ;
eymis axillaribus, laxe dichotomis, diffusis, 8 ad 10 em longis; floribus
5-meris; sepalis petalisque fimbriatis.

A small tree, 3 to 8 m high, the branches reddish-brown or greenish,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. OE

slender, terete. Leaves subcoriaceous, shining, elliptical, oblong-ellip-
tical or narrowly oboyate-oblong, 7 to 13 cm long, 3 to 6 em wide, entire
or slightly and obscurely denticulate towards their apices, the base
acute, the apex short acuminate, the acumen acute or blunt, rarely
retuse; nerves 5 to 7 on each side of the midrib, not prominent, distant,
irregular, anastomosing, the secondary nerves and lax reticulations nearly
as prominent as the primary veins; petioles 5 to 7 mm long. Cymes

9

axillary, dichotomous, diffuse, 8 to 10 em long, the peduncles 3 to 5
cm long, the flowers numerous, white, 8 to 10 mm in diameter, the
bracts and bracteoles linear or acicular, the former about 3 mm, the
latter 1 mm long. Sepals 5, orbicular to reniform, the margins shortly
fimbriate. Petals obovate, narrowed below, not clawed, 5 mm long,
fimbriate. Filaments 2 mm long; anthers 0.5 mm long. Fruit glab-
rous, red, broadly obovoid or turbinate, nearly 1 em long, 1.5 cm in
diameter, 5-lobed, the lobes rounded.

Mrinporo, Cuming 1552. Luzon, Province of Benguet, Hlimer 6462, June, 1904;
Bur. Sci. 3551 Mearns, July, 1907; Williams 1024, Oetober, 1904: Provinee of
Rizal, Bur. Sci. 1037, 1443, 4612 Ramos, July, August, 1906; August, 1907:
Province of Tayabas, Lagumanoc, Merrill 3356, November, 1903. MaAssBartr,
Merrill 3064, August, 1903.

A species previously confused with Huonymus timorensis Zipp., which is how-
ever a synonym of Huonymus javanicus Blume. Most closely allied to H. attenwa-
tus Wall., of British India and to EH. gibber Hance, of Hongkong, differing from
the former in its shorter cymes, bracts and bracteoles and in the shape of its
leaves, and from the latter in its longer leaves and cymes, the leaves of Hance’s
species being rounded and retuse at the apex. Lawson* admits Huonymus timo-
rensis Zipp., as a distinct species, giving its range as from Tenasserim or the
Andaman Islands to Pegu, Timor, and the Philippines, the Philippine distribution
being undoubtedly based on Cuming’s number cited above. The Andaman Island
‘reference is probably erroneous, as King* does not admit the species in his
“Materials for a Flora of the Malayan Peninsula.” Not having seen the other
specimens examined by Lawson, I am unable to state whether or not they are
identical with the Philippine plant, but it seems probable that they represent a
different species, or that the Indian specimens are really the same as Huonymus
alttenuatus Wall.

SIPHONODON Griff.

Siphonodon celastrineus Griff. in Cale> Journ. Nat. Hist. 4 (1844) 247, ¢.
14; Laws. in Hook. f. Fl. Brit. Ind. 1 (1875) 629; Mig. Fl.Ind. Bat. 1? (1859)
629; Koord. & Valet. Bijdr. Boomsoort. Java 7 (1900) 105.

Luzon, Province of Bataan, Lamao River, Whitford 1284, May, 1905: Province
of Tarlac, Garcia, August, 1903: Province of Rizal, Bosoboso, Bur. Sci. 1466
Ramos, September, 1906. Minporo, Bongabong River, Whitford 1423, February,
1906; For. Bur. 4053 Merritt, April, 1906. Ticao, For. Bur. 1072 Clark, June,
1904.

British India and Java.

ZEHOOK tile | brite. Inds en cls ta)mmG us
3’ Journ. As. Soc. Beng. 65° (1896) 343.

72299—_5

240 MERRILL.

A peculiar genus, anomalous in the family, here reported from the Philippines
for the first time; apparently widely distributed in the Archipelago, but variable.
The genus contains the above species, Siphonodon australe Benth., and S. pen-
dulum Bailey, of Australia, and the following species, which is evidently
undescribed.

Siphonodon pyriformis sp. nov.

Arbor parva, glabra, 5 ad 6 m alta; foliis coriaceis, nitidis, oblongo-

Nn

ellipticis vel oblongo-lanceolatis, 7 ad 15 em longis, basi acutis, margine
obscure crenulatis; nervis utrinque 7 ad 10; fructibus pyriformibus, ea.
3 cm longis, crustaceo-carnosis, glabris.

A small tree, 5 to 6 m high, glabrous throughout. Branches and
branchlets terete, dark-reddish-brown, lenticellate. Leaves alternate,
coriaceous, glabrous, elliptical-oblong to oblong-lanceolate, rarely ellip-
tical-ovate, 7 to 15 em long, 3 to 6 em wide, the base acute, apex shortly
and obtusely acuminate or rather long acuminate, the margins obscurely
crenulate; lateral nerves 7 to 10 on each side of the midrib, irregular,
anastomosing, the secondary ones nearly as prominent; petioles 4 mm
long or less. Fruit axillary, solitary, the peduncles 5 to 8 mm long,
crustaceous-fleshy, glabrous, pyriform, about 3 em long, 2 em in diam-
eter, shining; seeds scattered.

Luzon, Province of Benguet, Elmer 5985, March, 1904; For. Bur. 5141 Curran,
August, 1906; Bur. Sci. 2875 Mearns, April, 1907.

A species allied to the preceding, readily distinguishable however by its
pyriform fruits.

ICACINACE 8.
STEMONURUS Blume.

Stemonurus laxiflorus (Miers) comb. nov.

Platea laxiflora Miers in Ann. & Mag. Nat. Hist. If 10 (1852) 111.

Gomphandra laxiflora Vidal Rey. Pl. Vase. Filip. (1885) 108.

Cissus flecwosa Turez. in Bull. Soc. Nat. Mose. 31 (1858) 415.

The species is enumerated here to call attention to the reduction of Cissus
flecuosa Turez., it being based on the same number of Cuming’s collection as was
Platea laxiflora. ‘Turezaninow’s description also applies to the specimens of
Cuming 891 that we have seen. Miers’ name having priority, is here retained,
but is transferred to Stenomurus, where the species apparently belongs. Plan-
chon in his monograph of the Ampelideae* remarks concerning Cissus flexuosa
“Est-ce bien un Ampélidée? Plus que douteux.”

Stemonurus merrittii sp. noy.

Arbor 10 ad 12 m alta, inflorescentiis ramulis petiolisque puberulis ;
folis papyraceis, oboyvato-ellipticis vel obovato-oblongis, acuminatis, ca.
20 em longis; cymis axillaribus, 1 ad 2 em longis; floribus sessilibus,
fasciculatis; filamentis brevibus, glabris; fructibus anguste oblongis,
2 ad 2.5 em longis, plus minus triangulari-compressis, suleatis.

‘DC. Monog. Phan. 5 (1887) 424.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 241

A tree 10 to 12 m Imgh, the branches terete, nearly black, the
branchlets somewhat compressed, yellowish-gray, puberulent. Leaves
obovate-elliptical to oblong-elliptical, papyraceous, about 20 em long,
S to 10 em wide, slightly shining, glabrous, paler beneath, the apex
short-acuminate, the base rounded or subacute, margins entire; nerves
8 to 10 on each side of the midrib, prominent beneath, somewhat ascend-
ing, parallel, the reticulations obscure, very lax; petioles puberulent,
about 1 cm long. Cymes axillary, solitary, 1 to 2 em long, puberulent,
rather congested, sometimes branched from the base, but more often
branched above only, the branches short. Flowers yellowish or cream-
colored, sessile, fascicled at the ends of the branchlets. Calyx puberu-
lent, shallow, about 2 mm in diameter, truncate or very obscurely toothed.
Petals 5, free, valvate, oblong, about 2 mm long, 1 mm wide, acute,
puberulent outside, glabrous and with an obscure keel within. Stamens
5; filaments very short, about 0.5 mm long; anthers 1.2. mm long,
deeply cleft at the base. Ovary glabrous, oblong. Fruit narrowly oblong,
glabrous, 2 to 2.5 em long, about 7 mm thick, compressed, triangular
in cross-section, sulcate on one side, the other two sides with a rib
or keel.

Minporo, Pinamalayan, in the mountains, altitude 100 m, For. Bur. 9916, 9915
Merritt, April 2, 1908.

IODES Blume.

lodes philippinensis sp. nov.

Iodes ovalis Vidal Phan. Cuming. Philip. (1885) 103; Rev. Pl. Vase. Filip.
(1886) 85, non Blume.

Ramis ramulis inflorescentiis folusque plus minus ferrugineo-pubes-
centibus ; foliis ovatis vel oblong-ovatis, membranaceis, usque ad 12 cm
longis, basi late cordatis, apice acuminatis; floribus breviter pedicellatis,
ca. 8 mm longis, plus minus hirsutis.

Scandent, the branches, branchlets, leaves, and inflorescence more or
less softly ferruginous-pubescent, the stems yellowish- to reddish-brown,
the ultimate branchlets very densely pubescent. Leaves opposite, ovate
to oblong-ovate, rather strongly acuminate, base broad, cordate, mem-
branaceous, pubescent on both surfaces, more densely so on the nerves and
especially on the under surface, 5 to 12 cm long, 3 to 8 em wide; nerves
about 6 on each side of the midrib, very prominent beneath, the reticula-
tions lax, distinct; petioles densely pubescent, 1 cm long or less. Cymes
axillary and terminating the short lateral branches, pubescent, few-
flowered, the peduncles short or elongated. Pedicels about 1 mm long,
the calyx ovoid, 2 mm long, more or less hirsute, deeply 5-lobed, the
lobes oblong-lanceolate, 1 mm long, acuminate. Corolla-tube cylin-
drical, more or less hirsute, about 7 mm long, the lobes 5, ovate-lanceolate,
strongly acuminate, reflexed, about 3 mm long, 1 mm wide. Anthers
about 1 mm long. Ovary narrowly obovoid, glabrous, about 1 mm long.

24? MERRILL.

Fruit broadly oblong, somewhat compressed, 1.5 em long, about 1 em
wide, somewhat hirsute, strongly reticulate.

PHILIPPINES, without locality, Cuming 1532. Minporo, Calapan, Merrill 987,
1287, January, April, 1903; Pinamalayan, Por. Bur. 5407 Merritt, October, 1906.
toMBLON, Hallier, January, 1904. Masparr, Merrill 3383, November, 1903.
CreBu, Bur. Sci. 1709 McGregor, September, 1906 (type). Lryrre, Palo, Blmer
T108, January, 1906. PALAWAN, Bur. Sci. 729 Foaworthy, March, 1906.

The specimen collected by Cuming, cited above, has been referred by Baillon ®
to Jodes ovalis Blume, but the Philippine plants appear to differ constantly from
typical Javan material (Pl. Bogor. Exsice. no. 27) in their differently shaped,
more cordate and much more acuminate, thinner leaves, less dense pubescence,
and very different inflorescence, in Jodes ovalis the inflorescence being about 20 em
long, and in J. philippinensis never more than 7 em in length and usually much
shorter,

URANDRA Thiwaites.

Urandra luzoniensis sp. nov.

Arbor glabra, 12 ad 29 m alta; foliis alternis, ellipticis vel obovato-
ellipticis, subcoriaceis, nitidis, usque ad 15 em longis, breviter obtuseque
acuminatis, basi attenuatis; cymis termininalibus 3 ad 6 cm longis;
floribus 5-meris, ca. 5 mm longis; fructibus oblongo-ellipsoideis, 2 ad
2.5 cm longis.

A tree, glabrous throughout, except the inflorescence, 12 to 29 m
high. Branches terete, light-gray, the ultimate branchlets usually
reddish-brown. Leaves alternate, elliptical to obovate-elliptical, sub-
coriaceous, shining, paler beneath, glabrous, 7 to 15 cm long, 4 to 7.5
cm wide, the apex shortly and obtusely acuminate, sometimes obtuse, the
base gradually narrowed and somewhat decurrent-acuminate; nerves
about 5 on each side of the midrib, slender, distinct beneath, ascending,
the reticulations very lax, nearly obsolete; petioles about 1 em long.
Cymes terminal, 3 to 6 em long, dichotomous, the peduncles 1 to 3
cm long, the ultimate branches and pedicels more or less ferruginous-
hirsute. Calyx short, cup-shaped, truncate or obscurely toothed, about
1.5 mm long and 1.7 mm in diameter. Petals 5, oblong-oblanceolate,
acute, glabrous, 5 mm long, 1.5 mm wide, crested at the apex inside.
Stamens 5; filaments 5 mm long, glabrous below, above on the inner
side below the anther, and on the back opposite the anther covered with
long weak hairs; anthers ovoid, 1.2 mm Jong. Ovary ovoid, glabrous ;
style conical. Fruit elliptical-oblong, glabrous, black when dry, 2 to
2.5 em long, about 1 em in diameter, slightly striate, the exocarp coria-
ceous, the seed similar in shape to the fruit, 1.5 em long or less.

Luzon, Province of Bataan, Lamao River, lor. Bur. 711, 1926, 2949 Borden,
May, September, and March, 1904-05; Mor. Bur. 562, 574 Barnes, March, 1904:
Province of Rizal, Bosoboso, Merrill 2669, June, 1903; For. Bur. 2445, 2989 Ahern’s
collector: Province of Cagayan, For. Bur. T077 Klemme, May, 1907. Mtnporo,
For. Bur. 6214 Merritt, January, 1907.

' DOC, Prodr. 17. GLsie) 20:

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 243

Urandra pauciflora sp. nov.

Arbor parva, glabra; ramulis tenuibus, glabris, teretibus; foliis ovatis
vel oblongo-oyatis, membranaceis, valde acuminatis, alternis, basi acutis
vel rotundatis; cymis axillaribus, paucifloris, ca. 4 cm longis; floribus
ca. 3, longe pedicellatis, 4-meris, 4 mm longis.

A small tree, glabrous throughout except the slightly puberulent
inflorescence. Branches and branchlets slender, terete, gray or brown-
ish. Leaves alternate, ovate to oblong-ovate, membranaceous, glabrous,
somewhat shining, entire, base acute or somewhat rounded, apex slen-
derly long-acuminate; nerves 5 or 6 on each side of the. midrib, not
prominent, the reticulations very lax, nearly obsolete; petioles 5 to 8
mm long, slender. Cymes axillary, slender, 3-flowered, obscurely
puberulent, the peduncles about 2 em long, the pedicels 1 to 1.5 em
long. Calyx short, obscurely 4-toothed, less than 1 mm long. — Petals
4, about 4 mm long, free, or united in bud. Stamens 4; filaments as
long as the petals, flattened, with a few long hairs on the back opposite the
anthers and along the margins just below the anthers; anthers 0.6
mm long. Ovary oblong, truncate, glabrous. Fruit unknown.

Mrinporo, near Lake Naujan, and Mount Haleon, For. Bur. 6770, 4327 Merritt,
April, 1907, and June, 1906.

A species characterized by its few-flowered very slender inflorescence and
4-merous flowers, the anthers with but few hairs.

GONOCARYUM Mig.

Gonocaryum calleryanum (Baill.) Becee. Malesia 1 (1877) 123.

Phlebocalymna calleryana Baill. in Adansonia 9 (1869) 147.

Gonocaryum tarlacense Vidal Sinopsis Atlas (1883) 20, ¢. 30, f. C.; Rev. Pl.
Vase. Filip. (1886) 86; Ceron Cat. Pl. Herb. (1892) 46; Merr. in Philip. Journ.
Sci. 1 (1906) Suppl. 86.

Gonocaryum sp. Mery. in For. Bur. Bull. 1 (1892) 34.

CAMIGUIN, (Babuyanes Islands), Bur. Sci. 2987 Fenix, June, 1907. Luzon,
Province of Zambales, Botolan, Merrill 2983, June, 1903: Province of Pampanga,
Arayat, Merrill 1406, March, 1903: Province of Principe, Baler, Merrill 1036,
August, 1902: Province of Laguna, Los Banos, Hliner 8317, April, 1906: Province
of Rizal, Bosoboso, Merrill 1862, 2816, April, July, 1903; For. Bur. 1989 Ahern’s
collector, November, 1904; Bur. Sci. 4601 Ramos, August, 1907; Decades Philip.
Forest Fl. no. 38, Mareh, 1904: Province of Bataan, Lamao River, For. Bur.
636, 831, 1366, 1805, 2106, 2744 Borden, April-December, 1904; Hlmer 6884, 6887,
November, 1904; For. Bur. 2199, 2645 Meyer, December, 1904; Whitford 475,
1212, April, July, 1904-05; Williams 493, January, 1904; For.» Bur. 6355 Curran,
March, 1907: Province of Tayabas, Mor. Bur. 7853 Merritt & Curran, November,
1907; Atimonan, Whitford 709, August, 1904; Lagumanoc, Merrill 3360, Novem-
ber, 1903; Pagbilao, Merrill 1928, April, 1903: Province of Camarines Sur,
Ahern 86, March, 1902.

A species common and widely distributed in Luzon, the oldest specifie name
being here adopted. From the description given by Baillon, Phelobocalymna cal-
leryana and Gonocaryum tarlacense are conspecific, but Platea laxiflora Miers,
is a quite different species. The species seems to be very closely allied to the
Malayan Gonocaryum teysmannianum Scheff.

244 MERRILL.
ELABHOCARPACEZ®.
ELAEOCARPUS Linn.

Elaeocarpus foxworthyi sp. nov.

Arbor ca. 15 m alta; foliis elliptico-ovatis, ca 18 cm longis, coriaceis,
obtusis, subtus ferrugineo-pubescentibus, nervis utrinque ca. 12, pro-
minentibus, margine breviter obscureque denticulatis ; fructibus ovoideis
vel ellipsoideis, dense ferrugineo-pubescentibus, ca. 3.5 cm longis.

A tree about 15 m high, the branches, branchlets, petioles, leaves
beneath, and above when young, panicles and fruits rather densely
ferruginous-pubescent. Leaves elliptical-ovate, about 18 cm long, 10
to 13 em wide, coriaceous, obtuse, rarely slightly and broadly acute,
the base rounded to subacute, glabrous above when mature, except on
the somewhat pubescent midrib and nerves, the margins minutely and
distantly denticulate; nerves about 12 on each side of the midrib, very
prominent, parallel, the reticulations very distinct beneath, subparallel ;
petioles 5 to 6 em long. Flowers unknown. Infrutescence axillary,
pubescent. Fruits ovoid or ellipsoid, about 3.5 cm long, 2.5 em thick,
very hard, obtuse, densely ferruginous-pubescent, with one mature seed.

PALAWAN, Iwahig, Bur. Sci. 858 Foxworthy, May, 1906; For. Bur. 4158 Curran,
May, 1906. .

A tree growing along the river, well characterized by its rather large, ellip-
tical-ovate leaves, and hard, ferruginous, ellipsoid, rather large, fruits.

MALVACE A.
HIBISCUS Linn.

Hibiscus cannabinus Linn. Syst. ed. 10 (1759) 1149; Mast. in Hook. f. Fl.
Brit. Ind. 1 (1874) 339; Hochr. in Ann. Consery. Jard. Bot. Genev. 4 (1900) 114.

Luzon, Manila, Merrill 38801, December, 1903: Province of Pangasinan, Bur.
Sci. 4904 Ramos, December, 1907. MINDANAO, Province of Surigao, Baganga,
Merrill 5434, October, 1906.

Widely distributed in the tropics, cultivated; not previously reported from the
Philippines.

FLACOURTIACEAS.
HOMALIUM Jacq.

Several species of /Tomalium occur in the Philippines, all endemic,
so far as is known at the present time. Some of the species are im-
portant timber trees, but as there has been considerable confusion in
the group, due to misinterpretation of some of the previously deseribed
species, the following key to the Philippine forms has been made. The
types of F.-Villar’s species are no longer extant, but those of Vidal are
preserved in the Kew Herbarium, and these have been examined. ‘There
are some manifest errors in the descriptions of the former author, which

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 245

have given rise to much of the confusion in the Philippine repre-
sentatives of the genus, and it is hoped that the following enumeration
will clear up the doubtful points:
Stamens 1 opposite each petal (§- BLACKWELLIA).

en ves;sotukys pubescent, beneath. _...-.--.0 4. ue eek ee 1. H. barandae Vid.

Leaves entirely glabrous.
Flowers 5—6-merous; petals subspatulate, somewhat broader than the sepals,

MITA UGC he Lea yeraCLeNa uekons se cnn. ene cae ee) eames 2. H. loheri Merr.
Flowers 6—7-merous; petals and sepals linear, equal, covered with very lone
’ b] 7 * 8

spreading hairs; leaves entire .......................-.. 3. H. panayanum F.-Vill.

Stamens 2 or more opposite each petal (§ MyYRIANTHEIA).
Petals spatulate, much exceeding the ovate-lanceolate sepals; flowers 4-5-
THELEN ON DIST SU Seach Same he We Tad, pate ae Lees Bee eee aN 4. H. bracteatum Benth.
Petals and sepals subspatulate, equal or subequal; flowers 5—8-merous.
Flowers 5-6-merous; lateral nerves of the leaves about 6
5. H. villarianum Vid.
Flowers 7-8-merous; lateral nerves of the leaves 11 to 13.
6. H. luzoniense ¥.-Vill.

(1) Homalium barandae Vidal Cat. Pl. Prov. Manila (1880) 32; Sinopsis
Atlas, (1883) 27, ¢. 53, f. A; Rev. Pl. Vase. Filip. (1886) 141; Ceron Cat. PI.
Herb. (1892) 87; F.-Vill. Nov. App. (1883) 94, in part, excluding the description
of the flowers.

Luzon, Province of Rizal, Bosoboso, For. Bur. 2959, 3377 Ahern’s collector,
April, September, 1905: Province of Batangas, For. Bur. 7749 Curran & Merritt,
November, 1907.

The type of this species was from the Province of Manila (Rizal) ; and Vidal,
in the original description, which is very short, speaks especially of the dense
pubescence on the lower surface of the leaves, which is also very characteristic
of the specimens above cited. It is the only Philippine species that has densely
pubescent leaves. F.-Villar’s description applies, at least in part, to an entirely
different species, as he describes the flowers with fourteen stamens, which applies
to the species of the section Myriantheia. The specimen cited by Vidal in his
Revision can not be considered as the type. T., Laing.

(2) Homalium loheri sp. nov.

Arbor inflorescentiis exceptis glabra; foliis oblongo-ellipticis, sub-
coriaceis, breviter obtuseque acuminatis, 12 ad 17 em longis, margine
crenato-serratis, nervis utrinque 8 ad 10; inflorescentiis terminalibus,
paniculatis, villosis, ramis elongatis; floribus fasciculatis, ebracteolatis,
5- vel 6-meris, pedicellatis; petalis subspatulatis, quam sepalis paullo
longioribus; staminibus 5 vel 6; ovario villoso.

A tree, apparently of medium size. Branches terete, light-gray,
lenticellate. Leaves alternate, subcoriaceous, shining, glabrous, oblong-
elliptical, 12 to 17 cm long, 5 to 8 cm wide, the apex shortly and
obtusely acuminate, the base acute, the margins crenate-serrate .through-
out; nerves 8 to 10 on each side of the midrib, prominent, anastomosing,
the reticulations distinct; petioles about 5 mm long. — Inflorescence
terminal, villous, paniculate, the branches few, elongated, 12 to 20 cm
long. Flowers in fascicles, the bracteoles, if any, very deciduous, the

246 MERRILL.

pedicels pubescent, 3 to 4 mm long, jointed to the calyx. Calyx tube
narrowly funnel-shaped, pubescent, about 2 mm long, the lobes 5 or 6,
oblong, 2 mm long, pubescent. Petals 5 or 6, slightly exceeding the
calyx lobes in length, subspatulate, pubescent. Stamens one opposite
each petal. Ovary villous; styles 4 or 5.

Luzon, Province of Rizal, Loher 2210; Bosoboso, Decades Philip. Forest Fl.
no. 251; For. Bur. 1975 Ahern’s collector, November, 1905.

A species closely allied to Homalium barandae Vid., the type, Loher 2210,
having been identified at Kew with Vidal’s species. The plant above described is
at once distinguished by its entirely glabrous leaves, even in young specimens. T.,
Laing.

(3) Homalium panayanum F.-Vill. Nov.. App. (1883) 94; Vidal Rev. PI.
Vase. Filip. (1886) 141; Ceron Cat. Pl. Herb. (1892). 87.

Homalium grandiflorum Naves in Blanco Fl. Filip. ed. 3, pl. 443, non Benth.

Guimaras, For. Bur. 262 Gammill, January, 1904; For. Bur. 4589 P. del Villar,
June, 1906. Luzon, Province of Zambales, for. Bur. 8111, 8419 Curran & Merritt,
December, 1907; Bur. Sci. 5051 Ramos, December, 1907.

F.-Villar describes the species as having petals and sepals 10, stamens 10,
rarely 20 or 30, while Vidal states that the petals do not appear to exceed 7 in
number, the stamens two to three times as many as the petals. Our specimens
show 6 to 7 petals and sepals, and an equal number of stamens, and I have
accordingly included the species in the section Blackwellia. Vidal’s statement,
however, would place it in the section Myriantheia. ‘The plate cited by F.-Villar,
shows 7-merous flowers, and apparently 7 stamens, and our specimens agree per-
fectly with the figure. A species well characterized by its entire leaves and linear
petals and sepals which are covered with long ciliate hairs. V., Puyot.

(4) Homalium bracteatum Benth. in Journ. Linn. Soc. Bot. 4 (1860) 37;
Rolfe in Journ. Bot. 23 (1885) 213; Vidal Phan. Cuming. Philip. (1885) 115;
Rev. Pl. Vase. Filip. (1886) 142.

Homalium luzoniense Merr. in Philip. Journ. Sei. 1 (1906) Suppl. 99, non
F.-Vill.

PHILIPPINES, without locality, Cuming 1109 (cotype). Luzon, Province of Ba-
taan, Lamao, Mor. Bur. 2071 Borden, October, 1904, For. Bur. 5473 Curran, Novem-
ber, 1906: Province of Bulacan, Angat, For. Bur. 11165 Agwilar, April, 1908:
Province of Pampanga, Mount Arayat, Merrill 4210, September, 1905: Province
of Camarines, Pasacao, Ahern 24, 72, January, March, 1902.

A species well characterized by its 4- or 5-merous flowers, and dissimilar
sepals and petals, the former being ovate-lanceolate, reflexed and much shorter
than the latter, which are spatulate and erect or spreading. T., Aranga.

(5) Homalium villarianum Vidal Rev. Pl. Vase. Filip. (1886) 142; Ceron
Cat. Pl. Herb. (1892) 87.

Homalium sorsogonense Elmer Leatl. Philip. Bot. 1 (1908) 325.

Luzon, Province of Camarines Norte, Vidal 791, type in Herb. Kew: Province
of Sorsogon, Hlmer 7311, November, 1905, type of H. sorsogonense Elm.

After a careful examination of the specimens and descriptions of these two
species, I can find no characters by which they can be separated, and accordingly
EImer’s recently described species is here reduced. On Vidal’s specimen the
flowers are 6-merous, and on Elmer’s specimen they are 5-merous, but this char-
acter alone is not sufficient to distinguish them, as in many species of Homalium
the floral parts vary in number even on the same plant.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 247

(6) Homalium luzoniense F.-Vill. Nov. App. (1883) 94; Vidal Rev. Pl. Filip.
(1886) 141; Sinopsis Atlas (1883) 27, ¢. 453, f. B.; Ceron Cat. Pl. Herb.
(1892) 87.

Homalium aranga Vidal 1. ¢., as syn.

Luzon, Province of Tayabas, For. Bur. 19, 24, Ware, September, 1903, For. Bur.
6037 Kobbe, December, 1906. ALABar, Vidal 354.

A species well characterized by its 7- or 8-merous flowers, narrow sepals and
spatulate petals. T., Aranga. ;

Homalium foetidum Benth., has been reported from the Philippines by F.-Villar,
Noy. App. 94, and by Ceron, Cat. Pl. Herb. 87, but both are probably erroneous
identifications. I .-Villar reduces Gordonia polysperma Blanco, to Homalium
foetidum Benth., but this is certainly an error. I can not identify Blanco’s
species with any known form of Homalium, and it may apply to some plant
of a very different genus.

RHIZOPHORACEAE.
SAGITTIPETALUM gen. nov.

Calyx basi bracteolis liberis; tubus brevis, ovarii basi adnatus; limbus
6-partitus, lobis lanceolatis, acuminatis, coriaceis, valvatis. Petala 6,
basi disci carnosi inserta, unguiculata, laminis oblongo-ovatis, margine
irregulariter lacerato-fimbriatis, basi valde sagittatis. Stamina 12, al-
terna breviora; filamentis capillaribus. Ovarium inferum, 1-loculare :
ovulis 12, collateralibus, in axillis interioribus superioribus — affixis,
pendulis. Stylus filiformis, stigma simplex, disciforme.

Sagittipetalum mindanaense sp. nov.

Arbor parva, glabra; ramulis teretibus, junioribus compressis; foliis
oppositis, petiolatis, subcoriaceis, nitidis, ellipticis vel oblongo-ellipticis,
acuminatis, basi acutis; cymis in avillis superioribus, ca. 3-floris, brevi-
bus; floribus mediocris, 6-meris.

A small tree, about 10 m high, glabrous throughout. Branches slender,
terete, the younger ones somewhat compressed, reddish-brown, the tips
resinous. Leaves elliptical to oblong-elliptical, 7 to 10 cm long, 3. to
5.5 em wide, the apex acuminate, the base acute, entire, shining, sub-
coriaceous; nerves 6 or 7 on each side of the midrib, irregular, inter-
arching near the margin, not prominent, the reticulation rather lax,
petioles 1 cm long or less; stipules caducous, lanceolate, 1.2 to 1.5 em
long. Cymes solitary in the upper axils only, short, usually 3-flowered,
the peduncle about 5 mm long, compressed, the bracts ovate, acuminate,
about 3 mm long; pedicels very short, the bracteoles two, similar to the
bracts, not united. Flowers greenish, 1 cm long. Calyx ovoid, cleft
to the middle, the lobes 6, lanceolate, acuminate, erect, 2 mm wide at the
base. Petals 6, clawed, alternate with the sepals, 5.5 mm long, the
claw 2 mm long, the blade oblong-ovate, strongly sagittate atthe base,
the apex acuminate, the margins irregularly lacerate-fimbriate, 3 mm
wide below. Stamens 12, 6 opposite the petals with slender filaments

248 MERRILL.

4.5 mm long, 6 opposite the sepals with filaments 3.5 mm long; anthers
0.7 mm long. -Ovary inferior, 1-celled; ovules 12 on a central placenta,
pendulous from the inner upper angle; style slender, 7 mm long;
stigma small, disciform. Fruit unknown, but the calyx accrescent, in the
immature specimens 1.7 cm long, when mature probably much longer.

MINDANAO, District of Zamboanga, Port Banga, lor. Bur. 9171 Whitford &
Hutchinson, December 9, 1907, in the dipterocarp forest at an altitude of about
20 m above the sea.

A curious genus well characterized by its strongly sagittate petals, from which
the generic name is taken, these being long-clawed and irregularly lacerate-fim-
briate, its 6-merous flowers, 12 stamens, of which the 6 opposite the petals are
longer than the 6 opposite the sepals, and its l-celled ovary with 12 pendulous
ovules. It is apparently closely allied to Caralliia but seems to be generically
distinct.

MELASTOMATACE/5.
MEMECYLON Linn.

Memecylon densiflorum sp. nov.

Ramulis acute tetragonis, non alatis; foliis ovato-oblongis, coriaceis,
nitidis, 4 ad 5.5 em longis, 1-nerviis, nervulis transversalibus obsoletis,
apice breviter obscure acuminatis, acuminibus obtusis, basi cuneatis;
cymis axillaribus, subsessilibus vel breviter pedunculatis, fasciculatis,
subcapitatis, densis, ca. 1.5 cm diam.

Erect, glabrous, the branches brown or grayish, slender, terete, the
branchlets 4-angled. Leaves coriaceous, shining, ovate-oblong, 4 to 5.5
em long, 1.5 to 2.5 em wide, base acute, apex shortly obscurely acuminate,
the acumen blunt, sometimes subobtuse; midrib prominent, the lateral
nerves obsolete; petioles slender, 5 mm long or less. Cymes axillary,
fasciculate, subsessile or with peduncles 2 to 3 mm long densely many-
flowered, forming subeapitate heads about 1.3 cm in diameter, the
pedicels ebracteolate, about 2 mm long. Calyx funnel-shaped, 1.8 mm
long and wide, truncate. Petals ovate, acuminate, about 2 mm long,
1.2. mm wide. Filaments 3 mm long; anthers 1.2 mm long.

MINDANAO, Province of Surigao, Bolster 368, October, 1906; Ahern 515,
February—May, 1901, V., Babagion.

A species allied to Memecylon cumingianum Presl, but readily distinguished
by its dense capitate inflorescence; -apparently also allied to M. pauciflorum
Blume.

MEDINILLA Blume.

Medinilla philippensis (Cham. & Schlecht.) comb. nov.

Avanthes philippensis Cham. & Schlecht, in Linnaea 4 (1829) 193.

LuzON, without locality given, but from Cavite Province between the town of
Cavite and Taal Lake, Chamisso, in Herb. Berol. (type): Province of Cavite,
Mendez Nunez, Bur. Sci. 1341 Mangubat, August, 1906.

This interesting species was based on very fragmentary material, the type,
which | have examined in the Berlin Herbarium, consisting of a single detached

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 249

leaf and a single detached fruit. It is exactly matched by the specimen collected
by Mangubat, cited above, which came from the same general region as_ the
original. Unfortunately the specimens collected by Mangubat are poorly prepared,
but I am able to add the following to the original imperfect description :
Branches terete, light-gray, densely tomentose. Leaves 8 to 12 em
long, 4 to 8 em wide, the nerves 3 or 4 on each side of the midrib,
curyed-ascending. Cymes lateral, 4 cm long or less, few-flowered,
densely tomentose; bracts narrowly obovate, about 1 cm long. Calyx
obseurely 4-toothed, the teeth short, broad. Petals 4, about 9 mm long.
Stamens 8, subequal; anthers 5 mm long.

The species is most closely allied to Medinilla lagunae Vidal but has quite
differently shaped leaves. It differs from J/. halconensis Mery., in its 4-merous
flowers and more numerously nerved leaves, which are pubescent beneath only,
while in MW. halconensis they are pubescent on both surfaces.

Medinilla cogniauxii sp. nov.

Medinilla bracteata Cogn. in DC. Monog. Phan. 8 (1891) 601, in part (Cuming
1335, 1487; Vidal).

Differt M. bracteatae Blume foliis longioribus, inflorescentiis termina-
libus, racemosis, multo longioribus, usque ad 20 cm longis, simplicibus.

A scandent or decumbent shrub 2 to 6 m high, the branches terete,
slender, shining, glabrous, the branchlets densely stellate-plumose-to-
mentose. Leaves membranaceous, elliptical-lanceolate, acuminate, base
narrowed, abruptly rounded and narrowly cordate, 7 to 12 em long, 2 to 5
em wide, glabrous above when mature, more or less stellate-tomentose
on the nerves when young, beneath densely stellate-plumose-tomentose
on the nerves and with scattered hairs on the lamina; nerves prominent
beneath, two on each side of the midrib, the outer pair subbasal
and extending to the middle of the leaf or above, the inner pair leaving
the midrib some distance above the base and extending to the apex, rarely
a third pair of short basal nerves present; petioles densely tomentose,
1 to 3 mm long. Racemes from the terminal axils only, 7 to 20 em
long, pendulous, slender, densely stellate-tomentose, the flowers usually
arranged in whorls of threes, each node with three membranaceous, ovate,
5 to 8 mm long, 5-nerved, persistent bracts, which are somewhat stellate-
tomentose; pedicels about 4 mm long. Calyx 4 to 5 mm long, truncate,
ovoid, densely hirsute, the stiff hairs ciliate. Petals 5, narrowly obovate,
obtuse, about 9 mm long, 4.5 mm wide. Stamens 10, subequal; anthers
3 mm long. Style 8 mm long. Each flower subtended by two per-
sistent bracteoles, which are orbicular-ovate, obtuse, 7-nerved, mem-
pranaceous, about 12 mm long, somewhat tomentose. Fruit subglobose,
setose, about 5 mm in diameter, enclosed by the persistent bracteoles.

Luzon, Province of Batangas, Cuming 1487, in Herb. Bur. Sci. M1inporo,
Baco River, Merrill 4035, March, 1905; McGregor 183, 200, April, 1905.

A very characteristic species, confused by Cogniaux with Medinilla bracteata
Blume, which is a quite different species. In January of the present year I

250 MERRILL.

examined Blume’s type in Herb. Leiden, and compared with it a specimen of
Cuming 1487. Blume’s specimen has an axillary and terminal somewhat branched
inflorescence, much shorter than in the Philippine specimens referred here, shorter
leaves and much smaller bracts. Medinilla bracteata Blume must be excluded
from the known Philippine flora, as Cuming 1335, and Vidal 777, 778, 1390, are
undoubtedly referable to the species above described, and not to IM. bracteata
Blume.

Var. angustifolia var. nov.

Differt typo foliis multo minoribus, lanceolatis, basi vix angustatis.

Stellate-plumose-tomentose as in the type. Leaves 6 em long or less

« 5 2

8 to 14 mm wide, not gradually narrowed below, the base rather abruptly
broad-cordate. Racemes axillary, shorter than in the type, the bracts,

bracteoles and flowers as in J/. cogniaurt.

MINDANAO, Province of Misamis, Mount Malindang, Mor. Bur. 4567 Mearns &
Hutchinson, May, 1906.

Medinilla malindangensis sp. noy.

Frutex glabra; ramis teretibus, ramulis tetragonis; foliis verticillatis,
quaternis, elliptico-ovatis, acuminatis, 5-nerviis, usque ad 5 em longis;
eymis axillaribus, paucifloris; floribus 5-meris.

A glabrous shrub. Branches terete, light-gray, the branchlets 4-angled,
not winged, slender. Leaves whorled, 4 rarely 3 at each node, coria-
ceous, elliptical-ovate, 2.5 to 5 em long, 1.5 to 2.5 em wide, acuminate,
base cuneate; nerves 5, rather prominent beneath, reticulations obso-
lete; petioles 2 to 3 mm long. Cymes lateral, 3 em long or less, few-
flowered, the rachis 1.5 em long or less. Flowers unknown. Calyx
in fruit cup-shaped, about 6 mm long, 5 mm in diameter, truncate,
the limb somewhat produced, and with 5 very obscure teeth.

MINDANAO, Province of Misamis, Mount Malindang, Mor. Bur. 4557 Mearns &
Hutchinson, May, 1906.

A species apparently allied to Medinilla crassinervia Blume, but with very
differently shaped leaves which are much smaller. Well characterized by its
small, acuminate, 5-nerved, verticillate leaves.

Medinilla cephalophora sp. noy.

Glabra: ramis teretibus; foliis oppositis, papyraceis, oblongis, acumi-
natis, ca. 20 em longis, 5-neryiis; inflorescentiis terminalibus (?) ; floribus
5-meris, in capitulis cylindraceis, 8 ad 12 cm longis congestis; bracteolis
spatulatis.

Scandent, 6 m high, glabrous. Branches terete, glabrous, gray.
Leayes opposite, papyraceous, oblong, acuminate, about 20 em long, 7
em wide, 5-nerved from the base, the base subacute or somewhat rounded ;
petioles 1.5 to 2.5 em long, rather stout. Inflorescence terminal (?),
the flowers racemosely disposed on the thickened rachis forming a rather

)

dense cylindrical head 8 to 12 em long, about 8 em in diameter; rachis

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VL 201

5 to 7 mm thick, densely covered with pedicel- and bract-scars, setose ;
pedicels about 5 mm long, the bracteoles numerous, exceeding the flowers,
about 18 mm long, spatulate, the limb elliptical-ovate, obtuse, 6 mm
long, 5 mm wide. Calyx 4 mm in diameter, truncate. Petals 5, in-
equilaterally obovate, apex oblique-subtruncate, retuse-apiculate, 7 to
8 mm long, 5 mm wide. Stamens 10, subequal; filaments and anthers 4
mm long. Fruit ovoid, black, glabrous, about 1 cm in diameter, crowned
hy the persistent calyx.

Necros, Cadiz Nuevo, For. Bur. 4223 Hverett, February, 1906. MInpbANnao,
Province of Surigao, Bolster 290, April, 1906.

A species quite different from any in the genus, characterized by its congested,
cylindrical inflorescence. According to Everett the inflorescence is red and the
flowers have an offensive odor when fresh.

Medinilla congesta sp. nov.

Ramis teretibus, glabris; ramulis, foliis subtus, paniculisque plus
minus dense setosis; foliis oppositis 3-nerviis, oblongo-ellipticis, acutis
vel breviter acuminatis, coriaceis, ca. 12 cm longis, sessilibus; inflores-
centiis ca. 18 em longis, longe pedunculatis, pedunculis gracilibus, den-
sissime setosis ; floribus 4-meris, congestis, valde bracteatis, prope ad apices
ramulorum umbellatis.

A shrup, the branches light-gray, terete, glabrous, the branchlets
slightly setose, the nodes very densely setose with subulate chaffy scales
1 cm long or less. Leaves opposite, coriaceous, pale, shining, oblong-
elliptical, 10 to 14 cm long, 4 to 6.5 em wide, margins somewhat revolute,
sessile or subsessile, the base somewhat clasping, strongly 3-nerved, rarely
with a supplementary pair of faint marginal nerves, the apex acute or
short-acuminate, glabrous above, beneath with numerous long setose hairs,
especially on the nerves. Inflorescence axillary (?), the peduncle very
densely setose, about 12 cm long, slender, the flowers crowded at the
apex, forming an oblong, rather dense head, 6 cm long and 3 em thick,
its branches about 1 em long, densely setose, each bearing at its apex two
or three, obovoid, slightly setose bracts about 12 mm long, and 5 to 7
short-pedicelled flowers, each flower subtended by three narrowly obo-
void 1 em long bracteoles, the pedicels densely setose, about 3 mm long.
Calyx urceolate, the tube ovoid, 2 mm long, the limb produced, 2 mm
long, somewhat spreading, truncate, with 4 nerves corresponding to teeth.
Petals 4, about 10 mm long, 5.5 mm wide, narrowly inequilaterally obo-
void, apex obtuse, oblique. Stamens 4; anthers 5 mm long.

Mrnporo, Mount Sablayan, For. Bur. 9760 Merritt, March, 1908, in forests
on exposed ridges at 1,000 m alt.

A species well characterized by its long-pedunculate, dense inflorescence which
is densely setose, 4-merous, 3-bracteolate flowers, and sessile, strongly 3-nerved,
opposite leaves.

252 MERRILL.
ARALIACEZ#.
BOERLAGIODENDRON Harms.

Boerlagiodendron luzoniense sp. nov.

Arbuscula 1 ad 5 m alta; foliis palmatim 5- ad 7-lobatis, lobis ob-
longis vel obovato-oblongis, dentatis et grosse irregulariter sinuatis ; um-
bellis plus minus dense castaneo-paleaceis; floribus 4-meris.

An erect shrub 1 to 3 m high. Branches somewhat thickened, light-
gray, glabrous. Leaves submembranaceous, glabrous, suborbicular in out-
line, 15 to 25 cm long, palmately 5- to 7-lobed, truncate or cordate at
the base, the lobes oblong to obovate-oblong, reaching to within 3 or 4
cm of the base, 5 to 7 cm wide, much narrowed below, apex short-
acuminate, margins irregularly dentate and each lobe with two or three
lobules, the lobules- ovate or oblong-ovate, acuminate, 2 to 5 em long,
their sinuses round; petioles 10 to 30 em long, rather slender, the basal
portion with from 3 to 7 strong crests, their margins rarely slightly
pectinate. Umbels terminal, compound, the peduncles 20 to 30, the
whole inflorescence beset with numerous, dark-brown, narrow, paleaceous
scales, the peduncles 2 to 3 cm long, subtended by numerous 1 em long
basal bracts, these bracts oblong, their margins somewhat fimbriate and
their backs covered with narrow, brown, paleaceous scales, each peduncle
bearing three branches, the middle one very short, the lateral ones 2.5 to
3 cm long, the bracts at the apex of the common peduncle and at the upper
two-thirds of the lateral branches similar to the basal ones but smaller.
Lateral heads about 1 cm in diameter, densely many-flowered, the flowers
perfect, yellow, sessile, and subtended by numerous, dark-brown, chaffy
bracteoles which are fimbriate and covered with setose scales. Calyx
glabrous, obovoid, truncate, 2.5 to 3 mm long, 1.5 to 2 mm thick. Petals
4, ovate-elliptical, glabrous, united in bud, about 3 mm long, 2 mm
wide. Stamens 4; anthers 1.5 mm long. Ovary 4-celled. Middle heads
subsessile, the flowers sterile, pedicellate, the pedicels 5 to 6 mm long,
the flowers glabrous, purplish, globose, 5 to 6 mm in diameter, 2-celled,
rarely 3-celled.

Luzon, Province of Benguet, Baguio, Blmer 5928, March, 1904; Williams
1123, July, 1904; For. Bur. 5088 Curran, August, 1906; Bur. Sei. 2729 Mearns,
April, 1907.

A species recognizable by its 4-merous flowers and dark-brown, densely palea-
ceous inflorescence, the specimens cited distributed as B. pulcherrimum (Vid.)

Harms., to which species they do not seem to be closely allied.
Boerlagiodendron camiguinense sp. noy.
Differt a praecedente foliis 3—5-lobatis, lobis non sinuatis, usque
ad dimidian partem laminae attingentibus, basi vix angustatis; floribus
5-meris.

Very similar to the preceding, the inflorescence with the same dense

ig

-. teow

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 253

covering of brown, paleaceous scales. Leaves paliately 3- to 5-lobed,
15 to 20 em long, subcoriaceous, glabrous, the base truncate or rounded,
the lobes oblong, acuminate, coarsely apiculate-dentate but not sinuate
and but slightly or not at all narrowed below, reaching to about the
middle of the leaf. Inflorescence similar to that of Boerlagiodendron
luzoniense, the perfect and sterile flowers much the same but the former
with only three petals and three stamens, the ovary 3-celled.

CAMIGUIN, (Babuyanes Islands) Bur. Sci. 4135 Fenix, July, 1907, a shrub 1
to 15 m high, on slopes along trails.

Boerlagiodendron pectinatum sp. nov.

Arbor parva, glabra; foliis coriaceis, circiter 25 cm longis, usque ad
medium palmatim 5-lobatis, petiolis ad basin seriatim manicato-cristatis,
cristulis margine valde pectinatis; inflorescentiis glabris, ramulis omni-
bus aequalibus; floribus 5-meris.

A tree 7 to 8 m high, the branches glabrous. Leaves coriaceous,
suborbicular in outline, about 25 em long, base subtruncate, palmately
5-lobed, lobes reaching to about the middle of the lamina, oblong-ovate,
acuminate, margins coarsely dentate, the smuses narrow, obtuse at the
base; petioles 13 to 15 em long the base with a series of 3 or 4 crests
which are strongly pectinate, their divisions being 6 to 10 mm long.
Umbels compound, glabrous, primary branches about 25, subtended by
numerous, coriaceous, ovate bracts about 3 mm long, the peduncles 1.5
to 2 em long, each peduncle with a pair of narrowly ovate, acuminate
bracts at the apex, each peduncle tripartite at the apex, its branches
equal, or subequal, 1 to 1.5 cm long, the two lateral branches bearing
heads of perfect flowers, the middle one with sterile flowers only; lateral
heads 6 to 8 mm in diameter, densely many-flowered, flowers sessile,
the basal bracteoles ovate, 1 to 1.5 mm long, their margins shghtly
fimbriate. Calyx oblong, truncate, 2 mm long, 1 mm thick, glabrous,
slightly angular. Petals 5, oblong-ovate, in bud 2 mm long. Stamens
5; anthers 1 mm long. Ovary 5-celled. Flowers in the middle heads
all pedicelled, pedicels 5 mm long, the flowers 3 mm long, their ovaries
3-celled.

BATAN (Batanes Islands), Mount Iraya, Bur. Seti. 3775 Fenix, June, 1907.

A species readily recognizable by its coriaceous leaves which are palmately
lobed to the middle, the base of the petioles furnished with strongly pectinate
crests, its 5-merous flowers and the secondary branches of the umbels being of
the same length in both the lateral and middle heads.

Boerlagiodendron lineare sp. noy.

9

Arbuscula ca. 3 m alta; foliis usque ad basin palmatim 4—6-divisis,
lobis hnearibus, 15 ad 20 em longis, 1 ad 1.5 cm latis, basi attenuatis;
floribus 5-meris.

An erect shrub about 3 m high, the stems light-gray, glabrous,
shining, somewhat scurfy at the tips. Leaves crowded at the apices of

254 MERRILL.

the branches, palmately cut into 4 to 6 linear segments which are free
to the base, these segments glabrous, submembranaceous, 15 to 20 em long,
1 to 1.5 em wide, narrowed above to the acuminate apex, the base
narrowly decurrent, the decurrent parts so narrow that the segments
appear to be petiolulate, this portion 1 to 1.5 em long, margins irregu-
larly and slightly repand, the teeth minute, distant; nerves numerous,
spreading; petioles: 4 to 6 em long, the base with 2 or 3 prominent
crests. Umbels compound, with few, dark-brown, fimbriate scales, the
peduncles about 10, 2 cm long, bracteate at the base and apex, each bearing
at its apex a short-peduncled central head and two lateral branches, the
central head composed of numerous, dark-brown, fimbriate-lacerate
bracteoles (sterile flowers fallen), the lateral branches about 3 cm long,
with a pair of bracts at about the middle, each bearing a terminal head
of perfect flowers 1 cm in diameter or less, these heads also with
numerous dark-brown lacerate-fimbriate bracteoles. Flowers _ sessile.
Calyx more or less funnel-shaped, truncate, about 2 mm long, 1 mm in
diameter. Petals and stamens not seen. Ovary 5-celled. Fruit ovoid,
5-ridged, 5-celled, about 3 mm long.

Luzon, Province of Pangasinan, Bur. Sci. 4953 Ramos, December, 1907.

A most characteristic species, at once recognizable by its 4 to 6 linear segments
which are almost distinct enough to be considered as leaflets.

Boerlagiodendron pulcherrimum (Vidal) Harms in Engl. & Prantl Nat.
Pflanzenfam. 3° (1904) 32.

Osmoxylon pulcherrimum Vidal Sinopsis Atlas 18 (1883) ¢. 55, f. B.

This is an imperfectly known species, the type being no longer extant. The
material on which it was based was from Binangonan de Lampon, Province of
Principe, Luzon, and the description states that it is a plant with digitately
7- to 9-lobed leaves, the lamina being 1 m long and the petioles 40 to 50 em long.
The figure shows 5-merous flowers, with the central head of sterile flowers
strongly peduncled. It is doubtless allied to Boerlagiodendron mindanaense above
described, but is quite different from that species both in its leaves and in its
peduncled central heads. A full description of the species can not be written

until it is again discovered.
SCHEFFLERA Forst.

Schefflera foxworthyi sp. nov.

Glabra: foliis 7-foliolatis, foliolis ovato-lanceolatis vel oblongo-lan-
ceolatis, apice sensim acuminatis, basi Cuneatis, margine integris,
reyolutis: inflorescentiis terminalibus, ramis elongatis, rhachidibus bre-
vibus; floribus 5-meris in imbellulis paucifloris dispositis.

Scandent, glabrous throughout, except some parts of the inflorescence,
branches light-gray, somewhat thickened. Leaves 7-foliolate, the peti-
oles 10 em long, the stipule clasping; leaflets ovate-lanceolate to oblong-
lanceolate, subcoriaceous, dull, 9 to 14 cm long, 3 to 5 em wide, apex
gradually acuminate, acumen sharp, base cuneate, margins entire, revo-
lute; primary lateral nerves 5 or 6 on each side of the midrib, irregular,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 1539)

distant, the secondary ones nearly as prominent, anastomosing and
forming looped submarginal nerves; petiolules 1 to 2.5 em long. In-
florescence terminal, the rachis 2 or 3 em long, rather stout, the branches
4 to 6, crowded, ascending, 20 to 30 em long, glabrous, each subtended
by a persistent, coriaceous, lanceolate, acuminate bract 2 to 3 cm long,
the bracts when young densely flocculose. Flowers numerous, 5-merous,
borne in 3- to 6-flowered umbels which are arranged along the primary
branches, the peduncles 3 to 4 mm long, each subtended by a small lan-
ceolate bracteole, the pedicels about as long as the peduncles. Calyx
disciform, about 1.5 mm in diameter. Ovary 5-celled. Fruit 3.5 mm
long, oblong-ovoid, the ridges 5, not prominent.

PALAWAN, Iwahig, Bur. Sci. 796, 915 Foxworthy, April and May, 1906.

A species allied to Schefflera caudata (Vid.) Merr., S. clementis Merr., and WN.
longifrutescens Elm., but readily distinguishable from all by the shape and venation
of its leaflets, and various other characters.

ERICACEAS.
RHODODENDRON Linn.

Rhododendron curranii sp. noy.

~

Arbuscula 2 ad 2.5 m alta, ramis ramulis foliisque glabris; foliis
coriaceis oblongo-obovatis vel oblongo-oblanceolatis, acutis vel obtusis,
basi sensim angustatis; pedicellis dense hirsuto-pilosis; floribus 2.5 ad
3 cm longis, purpureo-coccineis; staminibus 10, filamentis in parte infe-
riore plus minus hirsutis; ovario dense piloso.

A shrub 2 to 2.5 m high, the branches light-gray or brownish, glab-
rous, slender, terete, the ultimate ones 1.5 to 2 mm in diameter. Leaves
whorled, 4 to 6 or 7 at each node, coriaceous, oblong-obovate to oblong-
oblanceolate, glabrous and shining on both surfaces, paler beneath and
with scattered small glands, 2.5 to 5.5 em long, 0.5 to 2 cm wide, apex
blunt or acute, gradually narrowed towards the cuneate or somewhat
decurrent base, the margins slightly revolute; nerves obscure, about 4+ on
each side of the midrib; petioles 2 to 4 mm long. Flowers crimson-
purplish, in terminal sessile fascicles, three or four flowers at the apex
of each branchlet, the bracts smooth, imbricate, deciduous; pedicels
densely hirsute-pilose, 1.5 to 2 em long. Calyx an obscurely toothed ring
about 4mm in diameter. Corolla 2.5 to 3 em long, shghtly pubescent on
the outside, the tube rather broad, the lobes orbicular-obovoid, rounded
or retuse, 1.3 mm long, 1.5 mm wide. Stamens 10; filaments 1.5 to
1.8 cm long, 5-celled, slightly enlarged and hirsute below, glabrous above.
Ovary oblong, 5-celled, densely pilose, 5 mm long; style glabrous, about
9 mm long.

Luzon, Province of Zambales, Mount Tapulao, For. Bur. 8061 Curran, Decem-

ber, 1907, in thickets on ridges at 2,000 m alt., also from the same locality
Bur. Sci. 4988 Ramos, December, 1907.

72299——_6

256 MERRILL.

A species closely allied to Rhododendron lussoniense Rendle, differing somewhat
in the shape of the leaves, shorter corolla-tube and lobes and different color of
the flowers. Rendle speaks of the flowers of RK. lussoniense as having been pink
or tinged with pink, but Whitehead’s note on the type in the Herbarium of the
British Museum says “flowers pure white.”

Rhododendron malindangense sp. noy.

Arbor parva, ramis glabris, griseis, ramulis junioribus brunneis, pu-
berulis; foliis coriaceis, pallidis, oblongo-ovatis, apice rotundatis vel emar-
ginatis, basi cuneatis, utrinque squamulis glandulosis paucis notatis;
bracteis ovatis, coriaceis, acutis, margine breviter ciliato excepto glabris;
floribus solitariis, 2 em longis, tubo cylindraceo; staminibus 10, inaequa-
libus, glabris; ovario 5-loculare, dense lepidoto.

A small tree about 7 m high, the branches terete, grayish, glabrous,
the younger branchlets reddish-brown, puberulent. Leaves coriaceous,
pale, oblong-ovate, the apex rounded or emarginate, the base cuneate,
alternate or subverticillately arranged at the apices of the branchlets,
glabrous except for the few scattered glandular scales on both surfaces,
1.5 to 3 cm long, 7 to 10 mm wide; nerves nearly obsolete; petioles 2 to
3 mm long. Bracts ovate, brown, rather thin, 6 mm long or less.
Flowers red, solitary, few, the pedicels slender, 6 to 7 mm long, pu-
berulent or pubescent. Calyx a small disk about 1.5 mm in diameter.
Corolla 2 cm long, 4 to 5 mm in diameter, cylindrical, slightly lepidote,
the lobes 5, ovate, rounded, 5 to 6 mm long, 4 to 5 mm wide, somewhat
spreading. Stamens 10, unequal; filaments glabrous; anthers 1.4 mm
long. Ovary oblong, 5-celled, about 3.5 mm long, densely lepidote ;
style glabrous, about 18 mm long.

MINDANAO, Province of Misamis, Mount Malindang, for. Bur. 4705 Mearns &
Hutchinson, May, 1906. Common in forests at about 1,800 m. alt.

SAPOTACE®.
PALAQUIUM Blanco.

Palaquium retusum sp. noy.

Arbor ca. 10 m alta; foliis anguste oblongo-obovatis vel oblanceolatis,
coriaceis, glabris, 20 ad 35 em longis, 7 ad 12 em latis, apice rotundatis,
retusis, basi longe sensim angustatis; nervis utrinque 13, prominentibus,
ascendentibus; stipulis oblongo-ovatis, coriaceis, 1 ad 1.5 em longis,
persistentibus; floribus pedicellatis, dense ferrugineo-pubescentibus ;
ovario 9- vel 10-loculare.

A tree about 10 m high. Branches thickened, rugose, glabrous.
Leaves crowded at the apices of the branches, narrowly oblong-obovate or
oblanceolate, 20 to 35 em long, 7 to 12 em wide, apex broad, rounded,
retuse, gradually narrowed from the upper third to the acute base, coria-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 257

ceous, glabrous, shining, the midrib very stout, the lateral nerves 13 on
each side of the midrib, very prominent, ascending, the reticulations not
prominent; petioles stout, 1 to 3 cm long; stipules persistent, covering
the apices of the branchlets, grayish, glabrous, coriaceous, oblong-ovate,
acute, or obtuse, strongly keeled, 1 to 1.5 em long. Flowers in few-
flowered fascicles on the branches below the leaves; pedicels about 1.5
em long, densely ferruginous-pubescent. Outer three calyx lobes valvate,
densely ferruginous-pubescent, broadly ovate, acute or obtuse, coriace-
ous, about 6 mm long, the inner three somewhat petaloid, imbricate,
nearly glabrous, or pubescent only on the median portion of the back,
suborbicular, rounded or retuse. Stamens about 20. Ovary glabrous,
9- or 10-celled ; styles nearly 2 cm long.

Luzon, Province of Benguet, Baguio, For. Bur. 5095 Curran, August, 1906;
Elmer 8523, March, 1907.

A species remarkable for its elongated retuse leaves, prominent, persistent
stipules and 9- to 10-celled ovaries. In most species of Palaquium the ovary is
6-celled, so that the present species is anomalous in this respect as well as in its
persistent stipules. In all other characters it is a true Palaquiwm, and is ac-
cordingly described as such.

Palaquium elongatum sp. nov.

Arbor ca. 20 m alta; ramulis glabris; foliis lanceolatis vel anguste
elongato-lanceolatis, acuminatis, coriaceis, supra glabris, subtus dense
ferrugineo-pubescentibus nitidisque, 20 ad 25 cm longis, 4 ad 8 cm
latis; nervis utrinque 14 ad 16, prominentibus; floribus axillaribus,
solitariis, longe pedicellatis, dense ferrugineo-pubescentibus.

A tree about 20 m high, the branches thickened, gray, glabrous, the
apices of the branchlets with numerous, narrowly lanceolate, about 1 cm
long, coriaceous, glabrous stipules. Leavés lanceolate to narrowly oblong-
lanceolate, coriaceous, 20 to 25 em long, 4 to 8 em wide, glabrous on the
upper surface, beneath densely ferruginous-pubescent and shining, the
midrib and lateral nerves glabrous or nearly so, apex short-acuminate,
gradually narrowed below to the acute or slightly acuminate base;
nerves on each side of the midrib 14 to 16, prominent, somewhat ascend-
ing, reticulations nearly obsolete; petioles about 4 cm long, glabrous or
nearly so. Flowers in the leaf-axils, solitary, few, the pedicels ferrugi-
nous-pubescent, 5 cm long or less. Outer three calyx lobes ferruginous-
pubescent, valvate, broadly triangular-ovate, acute, about 6 mm long,
the inner three thinner, ovate, appressed-pubescent on the back. Corolla
appressed-pubescent on the outside. Stamens about 20. Ovary ferru-
ginous-pubescent, 6-celled; the style, in bud, glabrous, 4 mm long.

Luzon, Province of Tayabas, Mauban, For. Bur. 6620 Reyes, March, 1907.

A species well characterized by its narrow elongated leaves, and few, long-

pedicelled flowers.

258 MERRILL.

SIDEROXYLON Linn.

Sideroxylon stenophyllum sp. nov.

Arbor glabra, ca. 25 m alta; foliis anguste oblongo-lanceolatis, basi
apiceque acuminatis, subcoriaceis, nitidis, usque ad 11 cm longis, 1.5
ad 2.5 em latis ; nervis utrinque 8 vel 9, prominentibus ; fructibus ovoideis,
glabris, ca. 5 cm longis; seminibus nitidis, 3.2 em longis.

A tree about 25 m high, the branches terete, gray, glabrous, the
terminal buds slightly pubescent. Leaves narrowly oblong-lanceolate,

subcoriaceous, shining, glabrous, 7 to 11 cm long, 1.5 to 2.5 cm wide,
the apex acuminate, the acumen blunt, the base decurrent-acuminate ;
nerves 8 or 9 on each side of the midrib, prominent, the reticulations
obscure; petioles slender, 1 to 2 cm long. Fruit ovoid, glabrous, 5 em
long, apparently somewhat fleshy when fresh, but the pericarp brittle
when dry, gray, shining, 5-celled, usually with but two matured seeds
which are shining, brown, pointed at both ends, 3.2 em long.

Minporo, For. Bur. 8626 Merritt, January, 1908.

A very characteristic species, readily recognizable by its narrow few-nerved
leaves and very large fruits; apparently allied to Sideroxrylon macranthum Merr.,
but very different from that species.

OLEACE ZS.
JASMINUM Linn.

Jasminum macrocarpum sp. nov. § Unifoliolata.

Frutex scandens; ramis ramulisque brunneis, teretibus vel leviter com-
pressis, gracilibus; foliis ovatis, simplicibus, subcoriaceis, supra nitidis,
8 to 10 em longis, usque ad 7’ cm latis, apice tenuiter acuminatis; pe-
tiolis articulatis; nervis utrinque ca. 9; paniculis axillaribus terminali-
busque; fructibus nitidis, nigris, ca. 2 em longis.

A scandent shrub, nearly glabrous throughout, the branches and
branchlets terete or slightly compressed, slender, brownish,’ glabrous,
Leaves simple, ovate, subcoriaceous, glabrous, shining on the upper
surface, 8 to 10 em long, 4 to 7 cm wide, the base rounded or acute, the
apex slenderly and sharply acuminate; lateral primary nerves about
9 on each side of the midrib, not prominent, anastomosing, irregular ;
petioles 1 to 2 em long, jointed at the middle or at the lower two-thirds.
Flowers unknown. Panicles in fruit axillary and terminal, 10 em
long or less, slightly pubescent. Pedicels about 5 mm long. Calyx
somewhat urceolate, 2 to 3 mm long, slightly pubescent, obscurely 6-
toothed, in fruit much enlarged and thickened. Fruit ellipsoid or

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 259

obovoid, glabrous, fleshy, shining, black when dry, about 2 em long, with
a single large seed.

Luzon, Province of Rizal, Bosoboso, For. Bur. 3091 Ahern’s collector, May,
1905, Bur. Sci. 1073 Ramos, July, 1906; Tanay, Merrill 2326, May, 1903: Province
of Bataan, Lamao River, Whitford 394, June, 1904.

A very characteristic species, readily distinguished by its subcoriaceous,
sharply acuminate and shining leaves, and very large fruits.

Jasminum truncatum sp. noy. § Unifoliolata.

Subscandens, glabrum; foliis simpheibus, calycis limbo truneato,
corollae tubo usque ad 1 cm longo, laciniis 7 ad 9, lanceolatis.

A subseandent shrub 2 to 3 m high, glabrous throughout. Branches
slender, reddish-brown, terete. Leaves opposite, simple, ovate to oblong-
ovate, the base broad, rounded, gradually narrowed above to the acum-
inate apex, 6 to 11 em long, 3 to 6 cm wide; nerves about 11 on each
side of the midrib; petioles 1 to 2 em long, jointed at the lower fifth.
Inflorescence terminal, lax, each ultimate branch bearing three flowers,
the bracts subulate, about 2 mm long, the bracteoles smaller. Flowers
white, fragrant. Calyx cup-shaped, 1.5 to 2 mm long, truncate, or
with very minute obscure teeth. Corolla tube 1 cm long, the lobes 7
to 9, lanceolate, acute, nearly 1 cm long.

Sisuru (Sulu Archipelago), Merrill 5286, October, 1906, in thickets along

the seashore. .
A species characterized by its truncate calyx.

LOGANIACEZE.
GENIOSTOMA Forst.

Geniostoma philippinense sp. nov.

Arbuseula 2 ad 4 m alta; ramulis, subtus foliis, petiohs pedicellisque
plus minus pubescentibus vel puberulis; foliis oblongo-lanceolatis, ellip-
tico-lanceolatis vel ovato-lanceolatis, submembranaceis, nitidis, acum-
inatis, 9 ad 13 cm longis; nervis utrinque 5 ad 7, subtus prominentibus ;
floribus axillaribus, fasciculatis; ovario glabro.

A shrub 2 to 4m high. Branches terete, slender, light-gray or brown,
ultimately glabrous, the branchlets rather densely pubescent or puberulent.
Leaves oblong-lanceolate, elliptical-lanceolate or ovate-lanceolate, sub-
membranaceous, 9 to 13 cm long, 3.5 to 5.5 em wide, base rounded or acute,
apex acuminate, the upper surface glabrous and shining, usually blackish
in drying, beneath, especially on the midrib and nerves, more or less
pubescent with very short rather stiff hairs; nerves rather prominent
beneath, 5 to 7 on each side of the midrib, anastomosing, the reticulations
lax; petioles 4 to 7 mm long, usually rather densely pubescent. Flowers
in 5- to 10-flowered axillary fascicles, the pedicels slender, pubescent,

260 MERRILL.

about 5 mm long. Calyx teeth triangular-ovate, acute. Corolla about
3 mm long, the lobes oblong-ovate, acute or acuminate, reflexed, about
1.5 mm long. Filaments short; anthers minutely puberulent, 1 mm
long. Ovary glabrous. Fruit whitish, about 7 mm long; seeds numer-
ous, 1.5 to 2 mm long. ;

Luzon, Province of Rizal, Bosoboso, Merrill 1833, April, 1903; Bur. Sci. 2667
Ramos, May, 1907; Bur. Sci. 60 Foxworthy, January, 1906; Montalban, Loher
4105: Province of Benguet, Twin Peaks, Elmer 6451, June, 1904, a more pubes-
cent form, but apparently the same species.

The second species of the genus to be found in the Philippines, readily
recognizable by its pubescence, its fascicled flowers and glabrous or nearly glabrous

corolla-throat.
FAGRAEA Thunb.

Fagraea longiflora sp. nov.

Frutex scandens, glabra, pseudo-epiphytica; ramulis teretibus, cras-
siusculis; foliis ovato-ellipticis, subcoriaceis, 30 ad 40 cm longis, breviter
acuminatis; nervis utrinque 8; floribus fasciculatis, terminalibus, ses-
silibus, congestis ; corollae tubo cylindraceo, 10 cm longo.

A scandent pseudo-epiphytic shrub, glabrous throughout. Branches
terete, thickened, 1.5 to 2 em thick. Leaves crowded at the ends of
the branches, elliptical-ovate, 30 to 40 cm long, 20 cm wide or less, sub-
coriaceous, tough in texture and not brittle when dry, the apex short-
acuminate, the base decurrent-acuminate; nerves 8 on each side of the
midrib, prominent; petioles about 4 cm long; stipules large. Flowers
large, crowded in terminal fascicles or in depauperate sessile cymes. Ca-
lyx about 4 cm long, 5-lobed, the lobes about 2 cm long, imbricate, oblong-
ovate, blunt or subacute, the bracteoles 2.5 to 3 cm long. Corolla white,
the tube 10 cm long, cylindrical, slightly enlarged upwards, somewhat
villous within, the lobes 5, elliptical-ovate, about 4 em long. Anthers
about 7 mm long. Fruit unknown.

LUZON, Province of Laguna, Mount Banajao, lor. Bur. 8026, 8028 Curran &
Merritt, November, 1907, in forests at an altitude of about 700 m.

A very characteristic species, well distinguished by its long sessile or subsessile
flowers which are densely crowded into terminal fascicles or depauperated cymes.

Fagraea auriculata Jack in Malay Miscel. 2 (1822) 82; Blume Rumphia 2
(1836) 26, t. 72; Clarke in Hook. f. Fl. Brit. Ind. 4 (1883) 83.

PALAWAN, Mount Victoria, Bur. Sci. 642 Foaworthy, March, 1906.

British India to the Malay Peninsula and Archipelago; not previously reported -
from the Philippines.

MITREOLA Linn.

Mitreola oldenlandioides Wall. Cat. (1828) no. 4350; DC. Prodr. 9 (1845) 9;
Clarke in Hook. f. Fl. Brit. Ind. 4 (1883) 79.

Luzon, Province of Pangasinan, Bur. Sci. 4852 Ramos, December, 1907:
Province of Bulacan, near Norzagaray, Yoder 126, December, 1906.

British India to the Malay Archipelago, Australia and the Caroline Islands.
The first representative of the genus to be found in the Philippines.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 261
APOCYNACEZ%.
CARRUTHERSIA Seemann.

Carruthersia macgregori comb. nov.

Hilertonia macgregort Merrill in Govt. Lab. Publ. (Philip.) 35 (1906) 59.

A reéxamination of the type material, and study of additional specimens
collected by Mrs. Clemens in Mindanao, no. 756, September, 1906, and without
numbers, April and June, 1907, leads me to refer this species to Carruthersia,
the second species of the genus to be found in the Philippines. It is closely
allied to Carruthersia pilosa (A. DC.) F.-Vill., but is readily distinguished by
its glabrous leaves. The stamens are very slightly united and the two lobes of
the disk alternating with the carpels are present. The genus Hllertonia is thus
to be excluded from the Philippine flora.

STROPHANTHUS DC.

Strophanthus erectus sp. nov.

Frutex erecta, 1 ad 2 m alta; foliis submembranaceis, elliptico-ovatis
vel oblongo-ovatis, breviter acuminatis, nervis utrinque 5 vel 6; cymis
paucifloris, bracteis et lobis calycinis lanceolatis, acuminatis, staminum
aristis antheris 5-plo longioribus; corollae lobis ca. 13 cm longis.

A glabrous erect shrub 1 to 2 m high, the branches brown or reddish-
brown, terete, lenticellate, the branchlets shghtly compressed. Leaves
elliptical-ovate to oblong-ovate, 8 to 13 cm long, 3 to 6 cm wide, submem-
branaceous, glabrous, dull or slightly shining, paler beneath, the apex
short-acuminate, the base rounded to acute or somewhat acuminate, some-
times slightly inequilateral ; nerves 5 or 6 on each side of the midrib, very
prominent, anastomosing, the reticulations lax, distinct; petioles 1 cm
long or less. Cymes terminal, glabrous, few-flowered, the bracts and calyx
lobes lanceolate, long-acuminate, the latter 7 to 9 mm long. Corolla
tube white, slightly yellow within, 2 cm long, about 8 mm in diameter,
slightly constricted at the middle, enlarged above, the lobes about 8 mm
wide at the base, attenuate into long, slender, drooping, yellowish tips,
about 13 cm long; appendages about 1 cm long, cleft into two long,
slender, acuminate parts. Anthers 3 mm long, sagittate at the base, the
filaments very short, somewhat hirsute, the apical awn very slender,
about 15 mm long. Ovary glabrous; style cylindrical, about 12 mm
long. Follicle woody, oblong-ovoid, about 16 cm long, 7 mm wide;
seeds lanceolate, 1.5 to 2 cm long.

PALAWAN, Puerto Princesa, Merrill 695, February, 1903; also specimens culti-
vated in Manila from seeds taken from this specimen, Merrill 5176, March, 1906,
and Cuzner, February, 1908.

This species differs from Strophanthus cumingii DC., in its much longer petals
and appendages to the anthers and in other characters; it appears to be more
closely allied to 8. dichotomus DC., than to S. eumingti DC.

262 MERRILL.

VERBENACE®.
CALLICARPA Linn.

Callicarpa surigaensis sp. nov.

Arbor parva, ca. 8 m alta; ramis ramulis inflorescentiis foliisque
plus minus dense ferrugineo-stellato-villosis hirsutisque; foliis oblongo-
ovatis vel lanceolato-ovatis, caudato-acuminatis, basi acutis, 10 ad 15 em
longis, nervis utrinque ca. 7; cymis axillaribus, pedunculatis, usque ad
4 em longis, dense ferrugineo-hirsutis; floribus ca. 5 mm longis.

A small tree, about 8 m high, the branches, branchlets, inflorescence
and petioles densely ferruginous-stellate-tomentose and with numerous,
long hirsute hairs. Leaves oblong-ovate to lanceolate-ovate, 10 to 15
em long, 4 to 7 em wide, subcoriaceous, base acute, apex gradually
narrowed into a long, slender, caudate acumen, the upper surface rather
densely hirsute on the midrib and nerves, with scattered hairs on the
lamina, the lower surface paler, rather densely ferruginous-stellate-
villous, and with scattered long hairs on the nerves and midrib; margins
minutely denticulate; nerves about 7 on each side of the midrib, promi-
nent beneath, the reticulations distinct; petioles densely hirsute, about
1 em long. Cymes axillary, penducled, 4 cm long or less, densely
ferruginous-hirsute. Flowers red. Calyx about 3 mm long, densely
villous, 4-toothed. Corolla glabrous, 5 mm long, 4-lobed, the lobes
2 mm long, obtuse. Stamens 4; filaments subequal, 4 mm long the
anthers 2mm long. Ovary glandular; style 7 mm long; stigma capitate.

MINDANAO, Province of Surigao, W. B. Allen 168, July, 1907; Ahern 318,
May, 1901.

A species well characterized by its dense stellate-tomentose and_ hirsute,
ferruginous indumentum,

Callicarpa ramiflora sp. nov.

Arbor parva, ca. 5 m alta; ramulis petiolisque densissime fulvo- vel
ferrugineo-hirsutis; foliis elliptico-ovatis vel oblongo-ellipticis, usque
ad 30 em longis, subtus plus minus stellato-tomentosis; cymis brevibus,
congestis, fasciculatis, in ramis vetustioribus; floribus ca. 5 mm longis,
calyce plus minus stellato-tomentoso.

A small tree about 5m high. Branches stout, terete, gray, glabrous,
the branchlets densely ferruginous-hirsute. Leaves opposite, elliptical-
ovate to oblong-elliptical, 30 em long and 15 cm wide or less, sub-
coriaceous, apex short-acuminate, base acute, margins obscurely dentic-
ulate, glabrous on the upper surface except the ferruginous-pubescent
midrib and nerves, beneath rather densely stellate-tomentose ; nerves 12
to 14 on each side of the midrib, prominent beneath, the reticulations
very distinct; petioles very densely ferruginous-hirsute, 1 to 2 em long.
Cymes about 1 em long, fascicled, densely hirsute, congested, in the
axils of branches or in the axils of fallen leaves on the older branches.
Calyx slightly stellate-villous, 4 mm long, 4-toothed. Corolla 5 mm

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 263

long, the lobes 4, oblong, obtuse, 2 mm long. Stamens 4; filaments 4
mm long; anthers glandular, 2 mm long. Style 6 mm long; stigma
capitate,
Minpanao, Lake Lanao, Camp Keithley, Mrs. Clemens 1167, September, 1907,
and without number, July, 1907.
. A species well characterized by its large leaves, and fascicled, congested, short
eymes which are from the branches below the leaves.

Callicarpa basilanensis sp. nov.

Arbor parva, ca. 6 m alta; ramulis petiolisque densissime. ferrugineo-
hirsutis; foliis oblongo-oyatis vel oblongo-ellipticis, caudato-acuminatis,
usque ad 25 cm longis, subtus stellato-tomentosis; cymis 2 ad 2.5 cm
longis, fasciculatis, plus minus stellato-tomentosis, e axillis foliorum
delapsorum ; floribus glabris, ca. 8 mm longis, 4- rariter 5-meris,

A small tree about 6 m high, the branches stout, gray or brownish,
glabrous, the branchlets densely ferruginous-hirsute. Leaves opposite,
oblong-ovate to elliptical-oblong, 25 cm long and 12 cm wide or less,
subcoriaceous, the apex rather slenderly caudate-acuminate, the base
acute or rounded, the margins entire, the upper surface hirsute on the
midrib and nerves, and more or less papillate-hirsute on the lamina,
beneath rather densely stellate-villous and, along the nerves and midrib,
somewhat hirsute; nerves 9 to 12 on each side of the midrib, prominent
beneath ; petioles densely hirsute, 1 to 2 cm long. Cymes 2 to 2.5 cm
long, from the axils of fallen leaves, fascicled, more or less stellate-~
villous. Flowers fragrant, purplish, glabrous. Calyx 3 mm long, 2.5
mm in diameter, 4-toothed. Corolla 7 to 8 mm long, the lobes 4,
rarely 5, oblong, obtuse, about 3 mm long. Stamens 4, rarely 5, sub-
equal; filaments 5 mm long; anthers 3 to 3.5 mm long. Stigma ob-
scurely 4-lobed. Pyrenes four, the fruit depressed-globose, 5 mm in
diameter.

BASILAN, For. Bur. 3974, 6124 Hutchinson, January and July, 1906; Hallier,
January, 1904.

A species well characterized by its large, acuminate, entire leaves, fascicled
eymes from the axils of fallen leaves, and rather large glabrous flowers, which are
rarely 5-merous, hence intermediate between Callicarpa and Geunsia.

RUBIACEZ..
WENDLANDIA Bartl.

Wendlandia nervosa sp. nov.

Arbuscula 2 m alta: ramulis, subtus foliis, paniculisque hirsutis; folis
subsessilibus, elliptico-ovatis vel oblongo-ellipticis, coriaceis, mitidis, 3
ad 6 cm longis, breviter acuminatis; nervis utrinque 6 ad 8, supra valde
impressis, subtus prominentibus; paniculis terminalibus, 3 ad 4 cm
longis, congestis, densissime ferrugineo-hirsutis; calycis lobis erectis, 2
mm longis.

264 MERRILL.

A shrub about 2 m high, the branches rather stout, terete, brown,
glabrous, the branchlets rather densely brown- or ferruginous-hirsute.
Leaves opposite, coriaceous, elliptical-ovate or oblong-elliptieal, 3 to 6
cm long, 1.5 to 2.5 em wide, the upper surface shining, glabrous except
the somewhat pubescent midrib, the nerves strongly impressed, the
lower surface with scattered short hairs, the midrib and nerves hirsute,
the apex shortly acuminate, the base acute or somewhat rounded ; nerves
6 to 8 on each side of the midrib, very prominent; petioles stout, densely
ferruginous-hirsute, 1 mm long or less; stipules ovate, hirsute, 2 to 3
mm long. Panicles terminal, congested, 3 to 4 em long, very densely
ferruginous-hirsute, the bracts narrowly oblong-lanceolate, hirsute, 4 mm
long, the bracteoles similar but smaller. Calyx hirsute, 3 mm long, the
tube 1 mm, the lobes narrowly lanceolate, erect, hirsute, 2 mm long.
Fruit ovoid, about 2 mm in diameter. :

Luzon, Province of Zambales, Mount Tapulao, Bur. Sci. 5007 Ramos, December,
1907.

A very characteristic species, readily recognizable by its small, subsessile,
very strongly nerved leaves, congested inflorescence, and rather long erect calyx-
teeth.

MUSSAENDA Linn.

Mussaenda philippinensis sp. noy.

Arbuscula 1 ad 3 m alta; ramis, ramulis, subtus foliis, paniculisque
plus minus hirsutis; folis membranaceis, oblongo-ellipticis, leviter falea-
tis, usque ad 25 cm longis, apice acuminatis, basi longe decurrentibus ;
nervis utrinque ca. 10; calycis segmentibus anguste lanceolatis, hirsutis,
1.5 em longis, persistentibus, sepala foliacea alba.

A shrub 1 to 3 m high, more or less hirsute. Branches reddish-brown,
terete, the younger ones grayish-brown, hirsute. Leaves membranaceous,
oblong-elliptical, slightly falcate, 15 to 25 cm long, 6 to 10 em wide,
nearly glabrous above, except the slightly hirsute midrib and nerves,
somewhat shining, the lower surface paler, hirsute on the midrib and
nerves and with scattered hairs on the lamina, the apex rather strongly
acuminate, the base long and narrowly decurrent-acuminate, equilateral ;
nerves about 10 on each side of the midrib, distinct; petioles 2.5 em long
or less; stipules 1 em long, ovate, long-acuminate. Panicles terminal,
hirsute, the bracts 1.5 to 1.8 em long, hirsute, cleft into three long,
acuminate, narrow segments. Calyx segments persistent, narrowly lan-
ceolate, about 1.5 em long, 2 to 3 mm wide, with long spreading hairs,
one segment sometimes produced, foliaceous, white, its lamina narrowly
elliptical-ovate, acuminate, 7 em long. Corolla-tube slender, yellow, 2.5
em long, hirsute with long hairs, its lobes ovate, acuminate, about 3
mm long. Fruit obovoid, 1 to 1.5 cm long, when young with few
scattered long hairs.

SeEMERARA, Merrill 4139, July, 1905, common in thickets at about 6 m above

sea level.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 265

A species well characterized by its elongated, narrow, persistent calyx-lobes;
apparently allied to Mussaenda wrayii King, of the Malay Peninsula, but very
different from that species.

LASIANTHUS Jack.

Lasianthus everettii sp. noy.

Arbor parva vel arbuscula; ramulis foliis stipulis bracteisque plus
minus fusco- vel flavescenti-pilosis vel villosis; foliis lanceolatis vel oblan-
ceolatis, ca. 20 cm longis, longe tenuiter acuminatis, basi acutis; nervis
utringque 9 vel 10; stipulis ovato-lanceolatis, 3 cm longis, dense villosis,
Jonge caudato-acuminatis, submembranaceis; floribus axillaribus, solita-
riis vel fasciculatis, 5-meris; bracteis anguste lanceolatis, numérosis.

A small tree or shrub, the branches, leaves and stipules more or less
densely covered with the rather long brown to yellowish-green hairs,
the branchlets nearly black when dry. Leaves lanceolate or oblanceolate,
about 20 cm long, 3 to 4.5 em wide, submembranaceous, the apex long and
slenderly acuminate, base acute, both surfaces with numerous long yellow-
ish or brownish hairs ; nerves 9 or 10 on each side of the midrib, prominent
beneath, curved-ascending, anastomosing, the reticulations lax; petioles
1 cm long, densely villous; stipules prominent, ovate-lanceolate, long: and
slenderly acuminate, 3 cm long, 8 mm- wide, densely yellowish-villous
narrowed below towards the base, submembranaceous. Flowers axillary,
two or three or more in each axil, sessile, the bracts many, lanceolate,
slenderly acuminate, densely villous, about 2 em long. Fruit about 5 mm
long sparingly hirsute, crowned with a tuft of long brown hairs, 5-celled.

NeEeRos, near Cadiz Nuevo, For. Bur. 5591 Everett, November, 1906.

A very characteristic species, distinguishable by its yellowish hairs, lanceolate
long-acuminate leaves and very prominent stipules and bracts which are covered
with long yellowish hairs.

RANDIA Foust.

Randia racemosa (Cav.) F.-Vill. Nov. App. (1883) 108; Maxim. in Bull.
Acad. Pétersb. 29 (1884) 167; Forbes & Hemsl. in Journ. Linn. Soc. Bot. 23
(1888) 382. :

Stylocoryna racemosa Cay. Icon. 4 (1797) 45, t. 368; DC. Prodr. 4 (1830)
377; Hook. & Arn. Bot. Beech. Voy. (1841) 264; Mig. Fl. Ind. Bat. 2 (1856) 205.

Remijia odorata Blanco FI. Filip. ed. 2 (1845) 115; ed. 3 1: 205.

Randia densiflora Elm. Leafl. Philip. Bot. 1 (1906) 32; Merr. in Philip. Journ.
Sei. 1 (1906) Suppl. 130, non (7?) Benth.

Luzon, Province of Bataan, Lamao River, Mor. Bur. 1472 Ahern’s collector ; For.
Bur. 2047 Borden; Elmer 6873: Province of Batangas, For. Bur. 7631 Curran &
Merritt, October, 1907: Province of Pangasinan, For. Bur. 8390 Curran & Merritt,
December, 1907: Province of Rizal, Bosoboso, Merrill 2789, July, 1903; Bur. Sci.
1054 Ramos, July, 1906; San Mateo, For. Bur. 1844, 3166 Ahern’s collector.
September, 1904, June, 1905. Mrinporo, Pola, Merrill 2219, May, 1903. Mas-
BATE, Merrill 3054, August, 1903. MinpANAo, Lake Lanao, Camp Keithley,
Mrs. Clemens 1193, September, 1907.

Widely distributed in the Philippines, and extending to the Riu Kiu Islands.

266 MERRILI.

A species closely allied to and possibly identical with Randia densiflora ( Wall.)
Benth., the type of the genus Stylocoryna Cay. (not Stylocoryne Wight &
Arn.). The transfer to Randia is usually credited to Maximowiez, but F.-Villar’s
combination has priority. The type was from Cavite Province, collected by Née
in 1793. The same form was also collected by Haenke, a colleague of Née, speci-
mens of whose exist in the Prague Herbarium and in the Berlin Herbarium,
both of which I have examined.

Randia wallichii Hook. f. Fl. Brit. Ind. 3-(1880) 113; Forbes & Hemsl.
in Journ. Linn. Soc. Bot. 23 (1888) 382; Ceron Cat. Pl. Herb. (1892) 95. ;

Randia fitzalani Elm. Leafl. Philip. Bot. 1 (1906) 31; Merr. in Philip. Journ.
Sci. 1 (1906) Suppl. 130, non F.-Muell.

This species was reported from the Philippines by Ceron, |. ¢., the identifica-
tion of Vidal’s specimen having been made by Rolfe. After an examination of the
material representing this species in the Kew Herbarium, I can see no reason
for distinguishing the Philippine form from the species described by Hooker f.
The Australian Randia fitzalani F. Muell., is different, and the Philippine form
can not be referred to it. Im addition to the numerous specimens from the
Province of Bataan, cited by myself 1. ¢., under Randia fitzalani, and which must
all be referred to R. wallichii, | have examined the following specimens:

LuzON, Province of Benguet, Baguio, Hlmer 5927, 5994, March, 1904: Province
of Llocos Sur, For. Bur. 5664 Klemme, November, 1906: Province of Zambales,
Subie, Hallier, January, 1904; Botolan, Merrill 2988, June, 1903: Province of
Principe, Baler, Werrill 1103, September, 1902: Province of Batangas, For. Bur.
7635, T751 Curran & Merritt, October, November, 1907: Province of Pangasinan,
Bur. Sci. 4834 Ramos, December, 1907; For. Bur. 8327 Curran & Merritt,
December, 1907: Province of Rizal, Bosoboso, Merrill 2632, June 1903; For. Bur.
3095 Aherns collector, May, 1904.

sritish India to Yunnan and Java.

UNCARIA Schreb.

Uncaria velutina Havil. in Journ. Linn. Soe. Bot. 33 (1897) $4.

Nauclea canescens Bart]. in DC. Prodr. 4 (1830) 346, non Unearia canescens
Korth.

Nauclea clavisepala Elm. Leafl. Philip. Bot. 1 (1908) 350.

Luzon, without locality, Haenke, in Herb. Prague: Province of Camarines Sur,
Cuming 1470; without locality, Cuming 1503: Province of Laguna, Los Banos,
Elmer 8262, April, 1906 (type of Uncaria clavisepala Elm.!). MaAsparr, Vidal
2952.

The type of this species is Cuming 1503, given by Haviland through error as
no. 503. I have examined the specimens collected by Cuming and Vidal in Herb.
Kew, and the specimen collected by Haenke, type of Nauclea canescens Bartl.,
in Herb. Prague, as well as Hlmer 8262, a specimen of which is in our Herbarium,
as is fragment of Cuming 1503. Elmer’s specimen exactly matches Cuming’s,
except that the former is in flower and the later in fruit. Haviland states 1. ¢.,
that he had not seen the type of Nauclea canescens, but that judging from the
description, it appeared to be the same as Unearia velutina. After an examina-
tion of Haenke’s specimen I am able to aflirm the identity of the two species.
Bartling’s specific name, although the earliest one, is untenable in Uncaria on
account of the later use of the same name by Korthals, for a different species.

Kndemic,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 267
CUCURBITACEAE.
GYNOSTEMMA Blume.

Gynostemma elongatum sp. nov.

Foliis pedatim 5-foliolatis; foliolis submembranaceis, integris, glabris,
acuminatis, basi acutis inaequilateralibus ; floribus albis, dioicis, femineis
8 ad 10 mm longis.

Scandent, glabrous throughout, except the inflorescence. Stems
slender, brownish. Leaves pedately 5-foliolate, the petioles 2 cm long,
petiolules about 5 mm long; leaflets elliptical-ovate, 4 to 6 cm long, 2 to
3.5 cm wide, submembranaceous, glabrous, entire, base inequilateral, acute,
apex short acuminate and sharply apiculate; nerves about 4 on each
side of the midrib, distant, curved, reticulations very few; tendrils simple,
10 cm long or more. Panicles about 15 em long, narrow, the branchlets
slightly pubescent. Pistillate flowers white, 8 to 10 mm long, the calyx
tube narrowly oblong, densely ferruginous-puberulent, 2 mm thick, the
lobes spreading, 1.5 mm long. Corolla rotate, the lobes ovate, acute,
2.5 mm long, veined. Ovary 3-celled; styles 3, free, cleft. Immature
fruit oblong, 2 cm long, 4 mm thick, truncate. Staminate flowers
not seen.

Luzon, Province of Bataan, near Lamao, Mor. Bur. 5474 Curran, November 23,
1906, climbing on trees in clearings.

A curious species, the leaves nearly identical with those of Gynostemma
integrifoliolum Cogn., but well characterized by its elongated ovary; possibly a
new genus, but difficult to determine this with certainty without staminate
flowers and mature fruit.

Gynostemma pedatum Blume Bijdr. (1825) 23; Cogn. in DC. Monog. Phan.
3 (1881) 913.

Luzon, Province of Benguet, Baguio, Hlmer 6033, March, 1904: Province of
Bataan, Lamao River, Whitford s. n., June, 1904. Mrxpanao, Lake Lanao, Camp
Keithley, Mrs. Clemens s. n., November, 1906, and March, 1907.

British India to Japan to Sumatra, Borneo and Java; new to the Philippines.

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PREVIOUS PUBLICATIONS OF THE BUREAU OF GOVERNMENT
LABORATORIES—Continued.

(Concluded from second page of cover.)

No. $2, 1905.—Biological Laboratory: I. Intestinal Hemorrhage as a Fatal Com-
plication in Amebie Dysentery and Its Association with Liver Abscess. By Richard
P. Strong, M. D. II. The Action of Various \Chemical Substances upon Cultures of
Amebz. By J. B. Thomas, M. D., Baguio, Benguet. Biological and Serum Laboratories:
III. The Pathology of Intestinal Amebiasis. By Paul G. Woolley, M. D., and W. E.
Musgrave, M. D.

No. 33, 1905, Biological Laboratory.—Further Observations on Fibrin Thrombosis
in the Glomerular and in Other Renal Vessels in Bubonic Plague. By Maximilian
Herzog, M. D.

1No. 34; 1905.—I. Birds from Mindoro and Small Adjacent Islands. II. Notes on
Three Rare Luzon Birds. By Richard C. McGregor. ,

No. 35, 1905.—I. New or Noteworthy Philippine Plants, IV. Il. Notes on Cuming’s
Philippine Plants in the Herbarium of the Bureau of Government Laboratories. IIT.
Hackel, ‘“‘Notes on Philippine Grasses.” IV. Ridley, ‘‘Scitiminez: Philippinenses.” V.
Clarke, “Philippine Acanthacee.’”’ By Elmer D. Merrill, Botanist.

No. 36, 1905.—A Hand-List of the Birds of the Philippine Islands. By Richard C.
McGregor and Dean C, Worcester.

LIST OF PREVIOUS PUBLICATIONS OF THE MINING BUREAU (NOW DIVISION OF
MINES OF THE BUREAU OF SCIENCE).

1890.—Descripeién fisica, geol6gica y minera en bosquejo de la Isla de Panay por
D. Enrique Abella y Casariego, Inspector General de Minas del Archipiélago.

1890.—Memoria descriptiva de los manantiales minero-medicinales de la Isla de Luzon,
estudiados por la comisién compuesta de los Sefiores D. José Centeno, Ingeniero de Minas
y Vocal Presidente, D. Anacleto del Rosario y Sales, Vocal Farmacéutico, y D. José de
Vera y Gomez, Vocal Médico.

1893.—Estudio descriptivo de algunos manantiales minerales de Filipinas ejecutado
por la comisién formada por D. Enrique Abella y Casariego, Inspector Genera] de Minas
D. José de Vera y Gomez, Médico, y D. Anacleto del Rosario y Sales, Farmacéutico ;
precedido de un prologo escrito por el Excmo. Sr. D. Angel de Avilés, Director General
de Administracién Civil.

1893.—Terremotos experimentados en la Isla de Luzén durante los meses de Marzo y
Abril de 1892, especialmente desastrosos en Pangasinan, Unién y Benguet. Hstudio ejecu-
tado por D. Enrique Abella y Casariego, Inspector General de Minas del Archipiélago.

1901.—The Coal Measures of the Philippines. Charles H. Burritt.
is EG Tate aelinieg of the Mining Laws (in force in the Philippines, 1902). ‘Charles H.

urritt.

1902, Bulletin No. 1.—Platinum and Associated Rare Metals in Placer Formations.
H. D. McCaskey, B. S.
es Pac aT oak of the Chief of the Mining Bureau of the Philippine Islands. Charles

. Burritt.

1903, Bulletin No. 2.—Complete List of Spanish Mining Claims Recorded in the Mining
Bureau. Charles H. Burritt. :

1903, Bulletin No. 3.—Report on a Geological Reconnoissance of the Iron Region of
Angat, Bulacan. H. D. McCaskey, B. S&S.

1904.—Fifth Annual Report of the Mining Bureau. H. D. McCaskey.

1905.—Sixth Annual Report of the Chief of the Mining Bureau. H. D. McCaskey.

1905, Bulletin No. 4.—A Preliminary Reconnoissance of the Mancayan-Suyoc Mineral
Region, Lepanto, P. I. A. J. Eveland, Geologist.

1905, Bulletin No, 5.—The Coal Deposits of Batan Island. Warren D. Smith, B. &.,

M. A., Geologist.
DIVISION OF MINES.

1908.—The Mineral Resources of the Philippine Islands, with a Statement of the
Production of Commercial Mineral Products during the year 1907, issued by Warren D.
Smith, Chief of the Division of Mines. \

LIST OF PREVIOUS PUBLICATIONS OF THE ETHNOLOGICAL SURVEY (NOW DIVI-
SION OF ETHNOLOGY, BUREAU OF SCIENCE).

Vol. I.—The Bontoc Igorot, by Albert Ernest Jenks. Paper, ®2; half morocco, P3.

Vol. II, Part 1.—Negritos of Zambales, by William Allen Reed. Paper, #0.50; half
morocco, 1.50.

Vol. II, Part 2 and Part 3.—The Nabaloi Dialect, by Otto Scheerer. The Bataks of
Palawan, by Edward Y. Miller. (Bound also in one yolume with Part 1, Negritos of
Zambales.) Paper, P0.50; half morocco, ®1.50. Combined, half morocco, P3.

1 Vol. I1I.—Relaciones Agustinianas de las razas del Norte de Luzon, by Pérez. Paper,
PO.75 ; half morocco, P2.

Vol. IV, Part 1.—Studies in Moro History, Law, and Religion, by Najeeb M. Saleeby.
Paper, ®0.50; half morocco, #1.50.

DIVISION OF ETHNOLOGY.

Vol. IV, Part 2.—The History of Sulu, by Najeeb M. Saleeby. Paper, #1.50.

Vol. V, Part 1 and Part 2.—The Batan Dialect as a Member of the Philippine Group

of Malayan Languages, by Otto Scheerer. The F and V in the Philippine Languages, by
C. E. Conant. Paper, 1.60.
Directions for Ethnographic Observations and Collections, 1908. For free distribution.
_ _AIl the above publications which are for sale may be obtained from the Director of
Printing, Manila, P. I. All publications for free distribution may be obtained from the
Librarian, Bureau of Science, Manila, P. I. Correspondents will confer a favor by
returning to the Library of the Bureau of Science any bulletins which they may have
dn duplicate, as there is a considerable demand for bulletins out of print.

me

1Qut of print.

f

The Philippine Journal of Science

Edited by
PAUL C. FREER, M. D., Ph. D.

The ‘‘Philippine Journal of Science’’ is issued as follows:

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A limited number of complete copies of Volume I and Supplement to
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(Copyrighted in the Philippine Islands, September, 1907. Entered at the post-office
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he
"4 eT if
JL i eae

By E. D. Merrit ia gt
m the B Botanical Section of the Biological Laboratory, Bureau of Science, Manila, P. I oe

Ge akan! FROM

"THE, PHILIPPINE JOURNAL OF ‘SCIENCE. pee oy aes:
"Published a the Bureau of Science ¢ of the Philippine Government, Manila, P, I

i

L. IV, No. 3, SECTION c, Beranr, Aususr, 1909

“A

na

~ MANILA
REAU OF PRINTIN ic

ORDER
No,

15.

17.

19.

20.

21.

32.

PREVIOUS PUBLICATIONS,
Bureau of Government Laboratories,
* No. 1, 1902, to No. 14, 1904.

No. 15, 1904, Biological and Serum Laboratories.—Report on Bacillus Violacevus Mas.

nile: A Pathogenic Micro-Organism. By Paul G. Woolley, M.

* No, 16, 1904, Biological Laboratory.—Protective Inoculation against Asiatic Cholera:
An Experimental Study. By Richard P. Strong, M. D. ;

abs 17, 1904.—New or Noteworthy Philippine Plants, II. By Elmer D, Merrill,

otanist.

* No. 18, 1904, Biological Laboratory.—l. Amebas: Their Cultivation and Etiologie
Significance. By W. E. Musgrave, M. ID., and Moses T. Clegg. Il. The Treatment
of alge Amebiasis (Amcebic Dysentery) in the Tropics. By W. E. Mus-
grave, D

No. 19, 1904, Biological Laboratory.—Some Observations on the Biology of the Chol-
era Spirillum. By W. B. Wherry, M. D.

No. 20, 1904.—Biological Laboratory: I. Does Latent or Dormant Plague Bxist
Where the Disease is Endemic? By Maximilian Herzog, M. D., and Charles B.
Hare. Serum Laboratory: Il. Broncho-Pneumonia of Cattle: Its Association with
B. Bovisepticus. By Paul G. Woolley, M. D., and Walter Sorrell, D. V. S. ‘
Pinto (Pano Blanco). By Paul G. Woolley, M. D. Chemical Laboratory:
IV. Notes on Analysis of the Water from the Manila Water Supply. By Charles
L. Bliss, M. S. Serum Laboratory: V. Frambesia: Its Occurrence in. Natives in
the Philippine Islands. By Paul G. Woolley, M. D.

No, 21, 1904, Biological Laboratory.—Some Questions Relating to the Virulence of
Micro-Organisms with Particular Reference to Their Immunizing Powers. By
Richard P. Strong, M. D. f

. No. 22, 1904, Bureau of Government Laboratories.—I. A Description of the New

Buildings of the Bureau of Government Laboratories. By Paul C. Freer, M. D.,
Ph, D. II. A Catalogue of the Library of the Bureau of Government Laboratories,
By Mary Polk, Librarian.

*No. 23, 1904, Biological Laboratory.—Plague: Bacteriology, Morbid Anatomy, and
Histopathology (Including a Consideration of Insects as Plague Carriers). By
Maximilian Herzog, M. D.

No. 24, 1904, Biological Laboratory,—Glanders: Its Diagnosis and Prevention (To-
gether with a Report on Two Cases of Human Glanders Occurring in Manila and
Some Notes on the BaUreriolony and Polymorphism of Bacterium Mallei). By
William B. Wherry, M.

. No. 25, 1904.—Birds she ‘the Islands of Romblon, Sibuyan, and Cresta de Gallo.

By Richard C. McGregor. _ (For first four bulletins of the ornithological series, see
Philippine Museum below.)

. No. 26, 1904, Biological Laboratory.—The Clinical and Pathological Significance of

Balantidium Coli. By Richard P. Strong, M. D.

. No. 27, 1904.—A Review of the Identification of the Species Described in Blaneo’s ©

Flora de Filipinas. By Elmer D. Merrill, Botanist.

. No. 28, 1904.—I. The Polypodiacee of the Philippine Islands. II. Edible Philip- -

pine Fungi. By Edwin B. Copeland, Ph. D.

. No. 29, 1904.—I1. New or Noteworthy Philippine Plants, III. II. The Source of

Manila Elemi. By Elmer D. Merrill, Botanist.

. No. 80, 1905, Chemical Laboratory.—I. Autocatalytic Decomposition of Silver Oxide.

II. Hydration in Solution. By Gilbert N. Lewis, Ph. D

. No. 31, 1905, Biological Laboratory.—I. Notes on a Case of Hematochyluria (To-

gether with Some Observations on the Morphology of the Embryo Nematode, Filaria
Nocturna). By William B. Wherry, M. D., and John R. McDill, M. D., Manila, P. I.
TI. A Search into the Nitrate and Nitrite Content of Witte’s ““Peptone,” with
Special Reference to Its Influence on the Demonstration of the Indol and Cholera-
Red Reactions. By William B. Wherry M. D.

No. 22, 1905.—Biological Laboratory: I. Intestinal Hemorrhage as a Fatal Com-
plication in Ameebic Dysentery and Its Association with Liver Abscess. By Richard
P. Strong, M. D. II. The Action of Various Chemical Substances upon Cultures
of Amebe. By J. B. Thomas, M. D., Baguio, Benguet. Biological and Seruit
Laboratories: I1Il. The Pathology of Intestinal Amebiasis. By Paul G. Woolley,
M. D., and W. E. Musgrave, M. D.

No. 38, 1905, Biological Laboratory.—Further Observations on Fibrin Thrombosis in
the Glomerular and in Other Renal Vessels in Bubonie Plague. By Maximilian
Herzog, M..D. )

* No. 84, 1905.—I. Birds from Mindoro and Small Adjacent Islands. II. Notes on
Three Rare Luzon Birds. By Richard C. McGregor.

. No. 85, 1905.—I. New or Noteworthy Philippine Plants, IV. II. Notes on Cuming’s

Philippine Plants in the Herbarium of the Bureau of Government Laboratories.
III. Hackel, ‘“‘Notes on Philippine Grasses.” IV. Ridley, “Scitaminem Philippinex-
ses. V. Clarke, “Philippine Acanthaceer.”” By Elmer D. Merrill, Botanist.

. No. 36, 1905.—A Hand-List of the Birds of the Philippine Islands. By Richard C:

McGregor and Dean C. Worcester.

*Report of the Superintendent of Government Laboratories for the Year Ending
August 31, 1902. (Appendix M of Third Annual Report of the ae ah Se
Commission.)

*Report of the Sunerintendent of Government Laboratories in the Philippine Telands
for the Year Ending September 1, 1903. (Appendix G of the Fourth Annual Report
of the Philippine Commission.)

Third Annual Report of the Superintendent of the Bureau of Government Laboratories
for the Year Ending August 31, 1904.

. Fourth Annual Report of the Sunerintendent of the Bureau of Government Laboratories

for the Year Ending August 31, 1905.
Bureau of Science—Publications,

. Price and Exchange List of Philinpine Bird Skins in the Collection of the Bureau of

Science, Manila, P. I. (Effective January 1, 1908.)

2. No. 1, 1909.—A Check-List of Philippine Fishes. By David Starr Jordan and Robert re

Earl Richardson. In press.

. No. 2, 1909.—A Manual of Philippine Birds. By Richard C, McGregor, Parts E.

and Il. A systematic index to the orders, families, and genera, Paper, P8 for the
two parts.

* Out of print.
(Concluded on third page of cover.)

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII.

By E. D. MErrRILt.

(From the Botanical Section of the Biological Laboratory, Bureau of Science,
Manila, P. I.)

The present contribution is, like the preceding papers of the series,
composed of miscellaneous diagnoses of apparently previously undescribed
species in various families, and records of various species previously
described and now found for the first time in the Philippines. A number
of nomenclatural changes are included, in accordance with the rules of
priority, and notes are added on some previously obscure or little-known
species. Four genera, Hmbolanthera, Everettiodendron, Greeniopsis, and
Ahernia are proposed as new, while the representatives of no less than
twenty-one genera, previously unrecorded from the Philippines are enu-
merated, Boottia, Anacolosa, Illicium, Erythrophloeum, Kingiodendron,
Azadirachta, Heynea, Blachia, Galearia, Microdesmis, Ostodes, Stro-
phioblachia, Pleiogynium, Leptonychia, Trichadenia, Ochrosia, Willough-
bya, Coptosapelta, Damnacanthus, Greenea, and Litosanthes. 'Twenty-
five species previously described by various authors from extra-Philippine
material are here recorded from the Archipelago for the first time, while
eighty-five species are described as new. On account of the rule adopted
by the Vienna Botanical Congress, requiring Latin diagnoses of all
species proposed after January 1, 1908, short diagnoses have been
added, and in cases where it has been necessary to consider species
proposed since that date, but which were without the required diagnosis,
this has been added.

ALISMACE/.

BOOTTIA Wallich.

Boottia renifolia sp. nov.

Planta aquatica, glabra; foliis reniformibus, longe petiolatis, apice
late rotundatis, basi cordatis, circiter 8 cm latis, nervis 7 ad 9; spathis
unisexualibus, tubulosis ; floribus masculinis vulgo binis, exsertis, antheris
9, pistilli rudimentiis pulviformibus; floribus femineis circiter 2.5 em
diametro, solitariis, pedicellatis; stylis 6, bifidis; staminodiis 9 quorum
6 linearibus, 3 capitatis.

248 MERRILL.

An aquatic glabrous plant. Leaves tufted, their petioles 10 to 25 em
long, the blades reniform, 4.5 to 6 cm long, about 8 em wide, papyraceous,
the apex broadly rounded, the base cordate; primary nerves 7 to 9, the
cross reticulations lax, indistinct. Male flowers one, or commonly two,
from tubular, linear spathes which are very slightly cleft at the apex
and not keeled or ribbed, exserted in anthesis, the pedicels very slender ;
sepals and petals as in the pistillate flowers; stamens 9, the filaments
flattened, 1.5 to 2 mm long; anthers basifixed, oblong, 1.5 mm long;
rudimentary pistil represented by a sessile, 1.5 mm long, cushion-like
body, the styles none. Pistillate flowers white, solitary, their pedicels
5 em long or less; sepals 3, elliptic-oblong, about 7 mm long, 3.5 to 4
mm wide, about 12-nerved; petals obovate, 12 mm long; styles 6, cleft
two-thirds to the base, the arms glandular-hairy; staminodes of two
kinds, 6 linear, 2.5 to 3 mm long, and 3 subsessile, capitate, 1 mm in
diameter. Ovary several-celled, cylindric, smooth. Fruit unknown.

Luzon, Province of Camarines, Bicol River, Bula, For. Bur. 12274 Curran,
June 5, 1908.

An interesting addition to our knowledge of the Philippine flora, the first
representative of the genus to be found in the Philippines. It is allied to Boottia
cordata Wall., of British India, but differs in many characters, notably in its
reniform leaves, linear, tubular spathes which contain but one or two flowers, the
staminate flowers without rudimentary styles, and the pistillate flowers with two
kinds of staminodes, six linear and three capitate. The genus contains about 15
species, in tropical Africa and Asia.

GRAMINE/#.
PANICUM Linn.

Panicum malabaricum (Linn.) comb. nov.

Poa malabarica Linn. Sp. Pl. (1753) 69.

Panicum arnottianum Nees in Steud. Syn. Pl. Gram. (1854) 59.

Panicum nodosum Hook. f. Fl. Brit. Ind. 7 (1897) 48, exel. syn., non Kunth.

Hooker f.* referred Poa malabarica Linn. to Centotheca lappacea (Linn.) Desv.,
but gave no reason for this. Considering that he was correct I transferred the
name to Centotheca as C. malabarica (Linn.) Merr. It seems, however, that
Hooker f. was in error in citing Poa malabarica Linn. as a synonym of Cento-
theca lappacea. Munro in his paper on the grasses of the Linnean herbarium
states regarding Poa malabarica “P(oa) malabarica is Panicum arnottianum
Nees. The reference to Rheede is correct; it is a very fair drawing.” * Linnaeus’
species is manifestly based on Tsjama-pullu of Rheede, Hortus Malabaricus 12:83,
t. 45. Through the kindness of Mrs. Agnes Chase of the United States Depart-
ment of Agriculture, I have received a tracing of the above plate, and although
the plate is crude, it seems to be well matched by several specimens in our het-
barium determined as Pamcum nodosum Kunth. Three specimens from Singa-
pore, collected by Ridley resemble it very closely.

“ay Brit. Ind; wale ooesoes * Journ. Linn. Soe. Bot. 6 (1862) 43.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 249

Panicum nodosum Kunth, which is closely allied, differs mainly in being much
smaller, with shorter, narrower leaves, and much smaller panicles. The type of
Kunth’s species was from the Philippines, P. nodosum being based on P. multinode
Presl, non Lam. Balansa, apparently assuming the Indian form to be typical
Panicum nodosum Iunth, has distinguished the smaller form, apparently typical
P. nodosum Wunth, as P. owonbiense.

I am disposed to refer to Panicum malabaricum (Linn.) Merr., two specimens
from the Philippines, Palmas Island, Merrill 5367, and For. Bur. 5349 Merritt,
from Mindoro. The former was previously referred by me to P. nodosum Kunth.

In a broad sense Panicum malabaricum might include P. nodosum Kunth, but
I am disposed to consider the small form described by Kunth as worthy of specific
rank.

Panicum cordatum Biise in Miq. Pl. Jungh. (1854) 376.

P. luxurians Merr. in Philip. Journ. Sei. 1 (1906) Suppl. .359, cum syn., non
Willd. :

Luzon, Province of Union, Bauang, Elmer 5595: Province of Bataan, Lamao
River, Williams 134, Elmer 6735: Province of Bulacan, Norzagaray, Yoder 1, 163.
CuLIon, Merrill 554.

This species was previously determined by me as Panicum luxurians Willd.,
but is apparently not that species. I have examined the material in the Will-
denow Herbarium so named, there being two specimens, one marked “Klein Ind.
1799” which is apparently a form of Panicum prostratum Lam.=P. reptans Linn.,
the other from St. Vincent, collected by Bory, which may be the same as P.
caespitosum Sw. Willdenow, Enum. 1 (1833) 109, gives as the only locality for
the species “Luzonia,” an apparent error on his part.

The specimens cited above closely resemble Panicum montanum Roxb., but can
be readily distinguished from that species by the first glume being as long as the
spikelet. The Philippine .material differs from Javan cordatum in its relatively
narrower and less prominently cordate leaves, but does not seem to be specifically
distinct from that species.

Panicum reptans Linn. Syst. Nat. ed. 10, 2 (1759) 871; Hitche. in Contr.
U. 8S. Nat. Herb. 12 (1908). 119.

Panicum prostratum Lam., Ill. 1 (1791) 171; Kunth Enum. 1 (1833) 89;
Steud. Syn. 1 (1854) 61; Hook. f. Fl. Brit. Ind. 7 (1897) 33; Merr. in Philip.
Journ. Sci. 1 (1906) Suppl. 355.

The oldest name for this common and widely distributed species is Panicum
reptans Linn., as shown by A. 8. Hitchcock 1. ¢., who has examined the type in
the Linnean herbarium.

Panicum paludosum Roxb. Hort. Beng. (1814) 6, nomen; Fl. Ind. 1 (1820)
307.

Panicum proliferuwm Hook. f. Fl. Brit. Ind. 7 (1897) 50; Merr. in Philip.
Journ. Sci. 1 (1906) Suppl. 359, non Lam.

Mr. A. 8. Hitchcock has examined the type of Lamarck’s Panicum proliferum
in the Paris Herbarium, and finds it to be identical with Panicum miliare Lam.,’
and has accepted P. dichotomiflorum Michx. as the earliest valid name for the
species, commonly identified by American authors as P. prostratum Lam. I am
now of the opinion that the species of tropical Asia and Malaya is distinct from
the American form, and have here adopted Roxburgh’s specific name for the
oriental form, as it seems to be the earliest valid one.

*Contr. U. 8. Nat. Herb. 12 (1908) 147.
85754——-4

250 MERRILL.

NEYRAUDIA Hook. f.

NEYRAUDIA MADAGASCARENSIS (Kunth) Hook. f.; Merr. in Philip. Journ. Sei.
1 (1906) Suppl. 381.

This species was admitted on the strength of identifications made by Vidal,
and Ceron, based on Cuming 623, and Vidal 4017. I have examined both in the
Kew Herbarium, and find them to be referable to Phragmites karka (Retz.) Trin.
Neyraudia madagascarensis must therefore be excluded from the Philippine flora.

ANDROPOGON Linn.

v-4

Andropogon leptos Steud. Syn. 1 (1855) 397; Merr. in Philip. Journ. Sci. 1
(1906) Suppl. 337.

Garnotiella philippinensis Stapf in Hook. Ic. Pl. LV 5 (1896) pl. 2494; Merr.
ures

Here enumerated to all attention to the reduction of the monotypic genus
Garnotiella. The type of Garnotiella philippinensis in the Kew Herbarium, is
identical with Andropogon leptos, and is a true Andropogon. Dr. Stapf, to whose
attention I called the matter, concurs in this reduction. To be sure of the identity
of the Philippine plant with Andropogon leptos Steud., I later examined the type
of Steudel’s species in the Berlin Herbarium, and found the determination to be
correct.

CYPERACEA.
FIMBRISTYLIS Vahl.

FIMBRISTYLIS CYMOSA R. Br.; Clarke in Philip. Journ. Sci. 2 (1907) Bot. 97.
This species must be excluded from the Philippine flora, as the specimen in
the Kew Herbarium, on which the record was based, bears a printed label with
the date 1831, and is from Cuming’s first collection from the southern Pacific,
and not from the Philippines. Cuming’s Philippine plants were distributed ten
years later.
BURMANNIACE.
BURMANNIA Linn.
B. coelestis D. Don Prodr. Fl. Nepal. (1802-03) 44; Ridley Materials Fl.
Malay. Penin. 2 (1907) 71.
B. azurea Griff. Notul. 3 (1851) 236; Ic. Pl. As. t. 272, f. 1; Beec. Malesia
2 (1878) 242, #. 15, f. 1-8.
Luzon, Province of Pangasinan, Infanta, Bur. Sci. 4913 Ramos, December, 1907.
The fourth species of the genus to be found in the Philippines. Widely
distributed from India to the Malay Peninsula, Borneo, Labuan and the Caroline
Islands.
FAGACEAS.
QUERCUS Linn.

Quercus obliquinervia sp. noy. § Cyclobalanus.

Arbor 20 m alta, cupulis exceptis glabra; ramis ramulisque griseis:
foliis oblongo-lanceolatis vel late lanceolatis, integris, crasse coriaceis,
usque ad 11 em longis, valde acuminatis, nervis utrinque 8, valde obliquis,
reticulis obscuris; cupulis 2 em diametro, intus glabris, extus pubescen-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 251

tibus ; glandibus oblongo-ovoideis, 2.3 cm longis apice rotundatis, breviter
apiculatis.

A tree about 20 m high, glabrous throughout, except the cups (in-
florescence unknown), branches terete, lenticellate, light-gray. Leaves
thickly coriaceous, oblong-lanceolate or broadly lanceolate, 7 to 11 cm
long, 2.5 to 4 cm wide, entire, the apex strongly acuminate, acumen
blunt, often slightly falcate, base decurrent-acuminate, both surfaces shin-
ing, the lower slightly paler than the upper; nerves 8 on each side of
the midrib, oblique, rather distinct beneath, nearly straight, more or
less curved near the margins, not anastomosing, the reticulations fine,
indistinct ; petioles 1.5 to 2 em long. Flowers unknown. Cup including
the stout stipe 1.5 cm high, inclosing only about the lower one-fourth
of the glans, 2 cm in diameter, glabrous and shining inside, outside
cinereous-pubescent, lamellae about 5 the lower ones distant, the upper
close, denticulate. Glans oblong-ovoid, 2.3 em long, about 1.8 cm in
diameter, glabrous, apex rounded, apiculate.

Luzon, Province of Benguet, Pauai, For. Bur. 18380 Alvarez, January, 1909:
Province of Abra, For. Bur. 14605 Darling, February, 1909, altitude 1,400 to 2,000
m. in forests; known in Benguet as Viclick, and in Abra as Bultioc.

A species well characterized by its very obliquely nerved leaves, probably most

closely allied to Quercus merrittii Merr., among the Philippine forms, but very
different from that.

ULMACE 2.
GIRONNIERA Gaudich.

Gironniera curranii sp. nov.

Arbor circiter 15 m alta, glabra; foliis ovato-lanceolatis vel elliptico-
lanceolatis, glabris, nitidis, 8 ad 12 cm longis, basi rotundatis, apice
caudato-acuminatis, margine integris vel superne pauce irregulariter
serratis; nervis utrinque 13 ad 15, prominentibus; floribus femineis
axillaribus, solitariis, longe pedicellatis, 5-meris.

A tree about 15 m high, glabrous throughout. Branches slender,
terete, gray or brownish, the branchlets somewhat angled, sometimes with
very few appressed hairs. Leaves ovate-lanceolate to elliptic-lanceolate,
subcoriaceous, shining, 8 to 12 cm long, 3.5 to 5 cm wide, paler beneath,
the base rounded, the apex caudate-acuminate, the acumen 2 em long or
less, 3 to 4 mm wide, abruptly acute or apiculate, margins entire, or
with few prominent, irregular teeth at the apex just below the acumen;
nerves 15 to 15 on each side of the midrib, prominent, curved-ascending ;
petioles 6 to 8 mm long. Pistillate flowers axillary, solitary, the calyx
lobes (in fruit) 5, ovate, acute or obtuse, about 2 mm long, the margins
somewhat ciliate, the fruit (immature) ovoid or elliptic-ovoid, 1.3 cm
long, 9 mm thick, glabrous, shining; style arms 2, stout, about 3 mm
long; pedicels about 1.5 em long, slender.

252 MERRILL.

Luzon, Province of Camarines, Mount Isarog, For. Bur. 10498 Curran, May 25,
1908, in forests on the lower slopes at an altitude of about 100 m.

A species well characterized by being glabrous throughout, and its smooth
shining leaves which are entire or with but few irregular teeth at the apex only.

MORACEZ#.
FICUS Linn.

Ficus merrittii sp. nov. § Covellia.

Arbor usque ad 12 m alta, ramulis petiolis, foliis subtus et inflores-
centiis plus minus ferrugineo-hirsutis ; foliis obovato-oblongis vel obovatis,
chartaceis, 18 ad 25 cm longis, integris vel obscure dentatis, apice breviter
acuminatis, basi inaequilateraliter cordatis; nervis utrinque 8 ad 10,
prominentibus ; stipulis caducis, lanceolatis, acuminatis, brunneis, 2.5 ad
3 cm longis; inflorescentiis caulinis, 10 ad 25 em longis, ramosis, plus
minus ferrugineo-hirsutis; receptaculis obovoideis, 1.5 ad 2 em longis,
plus minus ferrugineo-hirsutis, basi 3—bracteolatis; pedunculis hirsutis,
usque ad 2 cm longis; ovario oblongo-ovoideo; perigonibus nullis.

A tree 12 m high or less. Branches reddish-brown, terete, more or
less ferruginous-hirsute. Leaves opposite, obovate to oblong-obovate,
18 to 25 em long, 9 to 13 cm wide, chartaceous, somewhat shining, the
midrib and nerves on both surfaces with few or many, long, pale or
ferruginous hairs, the upper surface ultimately glabrous, margins entire
or obscurely dentate, the apex shortly and abruptly acuminate, somewhat
narrowed below to the slightly inequilateral and cordate base; the lower
surface minutely punctate; nerves 8 to 10 on each side of the midrib,
prominent, anastomosing, the reticulations lax, distinct; petioles ferru-
ginous-hirsute, 6 cm long or less; stipules caducous, lanceolate, acuminate,
brown, ferruginous-hirsute at the base only, otherwise glabrous, 2.5 to
3 cm long. Inflorescence from the trunk, few or many branched, 10
to 25 em long, reddish-brown, more or less ferrugineous-hirsute at the
nodes and on the bracts and bracteoles, the bracts ovate, acuminate, 8
mm long or less, deciduous. Receptacles usually at the ends of the
branchlets, few or many, obovoid, 1.5 to 2 em long, brown, purplish or
greenish, with few or many pale or ferruginous hairs, the base with three
small narrowly-ovate bracteoles 2 to 3 mm long; peduncles more or less
ferruginous-hirsute, 2 cm long or less. Staminate flowers not seen.
Fertile female flowers pedicellate, glabrous, the ovaries ovoid to oblong-
ovoid, about 1.5 mm long; styles slender, about 2 mm long. Perianth
none, or present in very young flowers as minute scales.

Minporo, For. Bur. 11466 Merritt, May, 1908 (type). Also represented by the
two following specimens from the same Island For, Bur, 6852, 11477 Merritt,
April, 1907 and May, 1908. N..v., Tibig.

A species closely allied to Ficus nota (Blanco) Merr., differing especially in
the long ferruginous hairs on the leaves, petioles, inflorescence and receptacles.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 253

SANTALACEZE.

EXOCARPUS Lab.

Exocarpus rolfeanus (O. Kuntze) comb. nov.

Xylophyllos rolfeana O. Kuntze Rev. Gen. Pl. 1 (1891) 589, (rolfsiana).

PALAWAN, Vidal 3647, type in Herb. Kew.; Victoria Peak, For. Bur. 3852
Curran, March, 1906, altitude 300 m.

A curious endemic species, known only from the above two collections. Curran’s
specimen is sterile, and is described by him as a 3 m high shrub.

OLACACEZ.
ANACOLOSA Blume.

Anacolosa luzoniensis sp. nov.

Arbor glabra, inflorescentiis exceptis, usque ad 15 m alta; foliis
chartaceis vel subcoriaceis, ovato-oblongis, elliptico-oblongis, vel oblongis,
8 ad 12 em longis, basi acutis, apice obtusis vel obscure acuminatis,
nervis utrinque circiter 5; floribus axillaribus, congestis, griseo-puberulis,
stipitatis, 6-meris ; petalis intus in partibus inferioribus glabris, carinatis,
supra crassis, subtrigonis, dense pubescentibus; filamentis latis, glabris,
connectivo indumento denso coronato; ovario 2-loculare; fructibus ellip-
soideis, 2 cm longis, disco accrescenti.

A tree reaching a height of 15 m, glabrous except the inflorescence.
Branches slender, terete, reddish-brown. Leaves alternate, oblong, ovate-
oblong, or elliptic-oblong, 8 to 12 em long, 3 to 5 em wide, chartaceous
to subcoriaceous, dark or pale when dry, sligtly shining, the base acute,
the apex obtuse, rarely obscurely and broadly acuminate; nerves:about 5
on each side of the midrib, anastomosing, the reticulations lax; petioles
5 to 8mm long. Flowers pale-green, gray-puberulent, densely congested,
axillary, the stipes 2 to 3 mm long. Calyx 3 mm in diameter, sub-
truncate and with 6 minute teeth. Petals 6, valvate, pubescent, lanceo-
late or oblong-lanceolate, 4 mm long, 1.8 mm wide, acute or somewhat
acuminate, the lower half, inside, glabrous and keeled, the upper portion
thickened, 3-sided, and densely pubescent in the central portion. Stamens
6, opposite the petals; filaments broad, about 1 mm long and broad,
flattened, glabrous, the antheriferous portion as broad as the filament and
crowned with copious hairs. Ovary superior, 2-celled, each cell with
a single pendulous ovule; style narrowly conical, 1 to 1.5 mm long.
Fruit fleshy, ellipsoid, about 2 cm long, 1-celled, entirely inclosed in the
accrescent disk.

Luzon, Province of Bataan, Mount Mariveles, For. Bur. 2830 Meyer, March,
1905 (type), For. Bur. 1199 Borden, June, 1904, For. Bur. 7222 Curran, June,
1907, Bur. Sci. 5165 Ramos, February, 1908. Minporo, For. Bur. 8650, 11363
Merritt, January, April, 1908. Masparte, For. Bur. 12601 Rosenbluth, January,
1909.

254 MERRILL.

The first representative of the genus to be found in the Philippines and ap-
parently Uistinet from the few other species of the genus. On Mount Mariveles
it is found at altitudes of from 700 to 850 m, but in Mindoro it occurs at lower
elevations.

MAGNOLIACE#.
ILLICIUM Linn.

Illicium philippinense sp. nov.

Arbor parva vel arbuscula, glabra; foliis congestis, alternis vel sub-
verticillatis, coriaceis, integris, acuminatis, oblongo-ellipticis vel elliptico-
lanceolatis, 5 ad 8 cm longis, nervis lateralibus obsoletis; floribus axilla-
ribus, solitariis, albis; staminibus circiter 15; carpellis circiter 8, 3 mm
longis.

A small tree or a shrub 4 m high or less, glabrous. Branches rather
stout, dark-colored, wrinkled when-dry, the branchlets somewhat oliva-
ceous. Leaves crowded near the tips of the branchlets, alternate, sub-
verticillate, coriaceous, oblong-elliptic or elliptic-lanceolate, 5 to 8 cm
long, 1.5 to 3.5 em wide, coriaceous, the upper surface shining, the
lower dull, acuminate at both ends; lateral nerves obsolete, the midrib
prominent; petioles 5 to 10 mm long. Flowers solitary, axillary, white,
fragrant, about 1 cm long. Sepals four or five, elliptic-oblong, 10 mm
long, 5.5 mm wide, obtuse, nerved. Petals 2-seriate, the inner ones
narrower than the outer, as long as the sepals but much narrower,
narrowly oblong, obtuse, 2.5 to 3mm wide. Stamens about 15; filaments
stout, swollen, 2 mm long; anthers about 1 mm long. Carpels about 8,
glabrous; 3 mm long including the recurved style. Mature fruit about
2 cm in diameter, consisting of from 5 to 8 carpels which are narrowly
ovate, rostrate, wrinkled when dry. Seed pale, shining.

Luzon, Province of Zambales, Mount Tapulao, Por. Bur. 9515 Curran & Merritt
(type); Bur. Sci. 5078 Ramos, December, 1907, on exposed mountain top in the
elfinwood, altitude about 2,100 m. Mrnporo, Mount Halecon, For. Bur. 4411
Merritt, June, 1906.

The only representative of the genus known from the Philippines, apparently
well characterized by its small leaves which are without evident lateral nerves.

It has previously been recorded by me from Mindoro as Jllicium sp.* It is
probably most closely allied to J. evenium King, of the Malay Peninsula.

>
LAURACEZE.
CRYPTOCARYA R. Br.

Cryptocarya lauriflora (Blanco) comb. nov.

Salgada lauriflora Blaneo Fi. Filip. ed. 2 (1845) 221, ed. 3, 2:50; Merr. in
Govt. Lab. Publ. (Philip.) 27 (1905) 73.

Busideroxylon borneense ¥.-Vill. Nov. App. (1883) 179, non T. & B.

Cryptocarya luzoniensis Vidal Rev. Pl. Vase. Filip. (1886) 222; Ceron Cat.
Pl. Herb. (Manila) (1892) 141; Merr. in Philip. Journ. Sei. 1 (1906) Suppl. 58.

‘Philip. Journ. Sci. 2 (1907) Bot. 272.

yt
* Sa
at?

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 255

Luzon, Province of Rizal, Cardona, Vidal 1680 (type, of C. luzoniensis, in
Herb. Kew.) ; Antipolo and Bosoboso, For. Bur. 401, 2663 Ahern’s collector, Feb-
ruary, 1904, and January, 1905: Province of Pampanga, Mount Arayat, Vidal
1686, in Herb. Kew.: Province of Bataan, Mount Mariveles Whitford 1050, 1043,
January, 1905, December, 1904, For. Bur. 6264, 6274, 7372 Curran, February and
July, 1907: Province of Camarines, Pasacao, Ahern 32, 183, January, February,

~ 1902. Mrnvoro, For. Bur. 8609 Merritt, January, 1908.

Blanco’s monotypic genus Salgada, has not previously been satisfactorily
identified, although it was referred by Fernandez-Villar® without question and
without discussion, to the Bornean genus Eusiderorylon Teysm. & Binn., but a
cursory examination of Blanco’s description shows at once that his species can not
be referred to Eusiderowylon. It does, however, apply closely to Cryptocarya,
and morover his specific description applies to Cryptocarya luzoniensis Vidal.
I have accordingly here reduced Salgada to Cryptocarya, and in making this
reduction, have adopted the earliest specific name for the species, reducing Vidal’s
Cryptocarya luzoniensis. Blanco’s description very clearly states that his genus
has nine fertile stamens, of which the outer six have introrse anthers, and the
inner three have extrorse ones; moreover he describes the anthers as 1-celled, his
expression “cuerpecillos laterales” certainly applying to the valves of the anther-
cells. Blanco describes the ovary as inferior, probably because it is immersed in
the calyx-tube, and later states that the fruit is inferior, the latter being ap-
parently an error in his description, or due to a mixture of material. There seems
to be absolutely no doubt as to the identity of Salgada with Cryptocarya, in spite
of the above discrepancies in Blanco’s description.

Cryptocarya bicolor sp. nov.

Arbor circiter 25 m alta; foliis alternis, elliptico-oblongis vel late
oblongo-lanceolatis, subcoriaceis, supra glabris, nitidis, subtus albo-glau-
cescentibus, 10 ad 20 cm longis, usque ad 7 cm latis, apice breviter
obtuseque acuminatis, basi acutis; nervis utrinque circiter 10, subtus
prominentibus, ferrugineo-pubescentibus, tenuiter anastomosantibus,
reticulis densis, obscuris; paniculis pubescentibus, axillaribus terminali-
busque, quam folia brevioribus; floribus sessilibus vel breviter pedicellatis,
circiter 3 mm longis.

A tree about 25 m high, more or less pubescent. Branches slender,
brown, terete, somewhat pubescent, the young branchlets rather densely
pubescent. Leaves alternate, elliptic-oblong or broadly oblong-lance-
olate, 10 to 20 cm long, 4 to 7 cm wide, subcoriaceous, the apex shortly
acuminate, the acumen obtuse, the base acute, the upper surface glabrous,
shining, somewhat olivaceous, the midrib sometimes pubescent, the lower
surface very pale, glaucous, nearly white, sparingly pubescent, the midrib
and lateral nerves ferruginous ; nerves about 10 on each side of the midrib,
obscure on the upper surface, very distinct on the lower, slightly curved,
spreading-ascending, attenuate and obscurely anastomosing near the
margins, the reticulations rather dense, not distinct; petioles pubescent,
about 8 mm long. Panicles axillary and terminal, 10 cm long or less,
pubescent, many-flowered, somewhat diffuse. Flowers yellow, fragrant,

5Nov. App. (1880) 179.

256 MERRILL.

about 3 mm long, sessile or short pedicellate, the bracteoles small. Calyx-
tube about 1.5 mm long, pubescent, cylindric; perianth-lobes pubescent
on the outside, subequal, oblong, about 2 mm long, acute or blunt.
Stamens as in other species of the genus, the filaments short, glabrous.
Staminodes 1.5 mm long, stipitate, acuminate. Fruit unknown.

Minpanao, District of Zamboanga(?), For. Bur. 6548 Hutchinson, March, 1907.
N. y., Ducatan, Masagcunadug.

A species well characterized by its leaves which are shining and glabrous above,
and nearly white beneath, the midrib and lateral veins ferruginous, the reticula-
tions not distinct, and its panicles which are about one-half as long as the leaves.
The timber is used for house-posts.

Cryptocarya palawanensis sp. nov.

Arbuscula cireiter 3 m alta; foliis chartaceis, oblongis, usque ad 35
em longis, 10 cm latis, apice acuminatis, basi acutis, utrinque glabris,
costa nervisque exceptis ; nervis utrinque circiter 15; paniculis axillaribus,
foliis multo brevioribus, pubescentibus ; perianthiis segmentis subaequali-
bus, pubescentibus, 2.5 mm longis; fructibus ovoideis, 1.5 cm longis,
nigris, nitidis, glabris.

A shrub about 3 m high. Branchlets ferruginous-pubescent. Leaves
chartaceous, oblong, up to 35 em long, 10 cm wide, the apex rather
strongly acuminate, the base acute, somewhat shining and glabrous on
both surfaces except the midrib and lateral nerves which are somewhat
pubescent, the upper surface olivaceous, the lower brownish; nerves about
15 on each side of the midrib, beneath very distinct, obscurely anastomos-
ing, the primary reticulations lax, the ultimate ones dense; petioles
pubescent, about 1 em long. Panicles axillary, pubescent, 8 to 10 em
long, the branches 3 em long or less. Flowers pedicellate or sessile,
pubescent, the tube cylindric, quickly inflated, somewhat longitudinally
striate, the perianth-lobes 6, subequal, pubescent, 2.5 mm long. Fruit
ovoid, black, shining, glabrous, 1.3 cm long.

PALAWAN, near the Iwahig Penal Settlement, For. Bur. 4141 Curran, April,
1906, along the banks of tidal streams.

A species characterized by its rather large leaves and comparatively short
panicles,

Cryptocarya pallida sp. noy.

Arbor vel arbuscula; ramis teretibus nigricantibus, glabris, ramulis
plus minus pubescentibus ; foliis ovatis vel elliptico-ovatis, chartaceis, 8 ad
11 em longis, 3 ad 5 em Jatis, apice acuminatis, basi acutis vel rotundatis,
supra olivaceis, glabris, nitidis, subtus subalbidis, parce pubescentibus,
costa nervis reticulisque brunneis; paniculis axillaribus, plus minus pubes-
centibus; fructibus ellipsoideis, 2.5 em longis, 1.5 cm diametro, glabris,
nitidis, utrinque apiculatis.

A shrub or tree, nearly glabrous. Branches terete, dark-colored, glab-
rous, the branchlets somewhat pubescent. Leaves alternate, chartaceous,
ovate to elliptic-ovate, 8 to 11 cm long, 3 to 5 em wide, the apex rather

a.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 257

sharply acuminate, the base acute or rounded, the upper surface oliva-
ceous, glabrous, shining, the lower surface nearly white, slightly pubescent,
the midrib nerves and primary reticulations brown; nerves 5 on each side
of the midrib, distinct beneath, anastomosing, the reticulations lax;
petioles about 5 mm long, slightly pubescent. Panicles axillary, in fruit
10 cm long or less, slightly pubescent. Flowers unknown. Fruits ellip-
soid about 1.5 cm in diameter, 2.5 cm long, dark-colored, glabrous,
shining, apiculate at both ends, obscurely longitudinally striate.

MINDANAO, District of Zamboanga, Tetuan, Ahern 594, 566, February to June,
1901, N. v., Ponit.

Characterized by its relatively large fruits which are apiculate at both ends,
and by its leaves which are nearly white beneath.

Cryptocarya everettii sp. nov.

Arbor, 10 ad 18 m alta; foliis alternis, oblongo-ellipticis, ellipticis,
vel oblongo-lanceolatis, subcoriaceis, 10 ad 20 cm longis, acuminatis,
supra nitidis, glabris, brunneis, subtus glaucescentibus, valde pulcher-
rime reticulato-venosis; paniculis pubescentibus, 8 ad 10 cm longis,
axillaribus terminalibusque; floribus albis, circiter 4 mm longis, pedicel-
latis vel subsessilibus ; fructibus oblongis, glabris, circiter 3 em longis.

A tree 10 to 18 m high, nearly glabrous throughout except the in-
florescence. Branches slender, light-gray, glabrous, the young branchlets
somewhat pubescent. Leaves alternate, oblong-elliptic, elliptic or oblong-
lanceolate, 10 to 20 em long, 4 to 7 cm, rarely 9 cm wide, subcoriaceous,
the upper surface glabrous, or sometimes slightly pubescent along the
midrib, shining, brown, the lower surface glaucous, the nerves and
reticulations prominent, brown, slightly pubescent, the apex abruptly
and sharply acuminate, the base acute; nerves about 10 on each side of
the midrib, beneath very prominent, anastomosing, the reticulations
rather lax, very distinct, the brown midrib, nerves and reticulations in
strong contrast to the glaucous under surface of the leaf ; petioles 5 to
10 mm long, somewhat pubescent. Panicles axillary and terminal, 8 to
10 cm long, pubescent, the flowers white, pedicellate or subsessile, about
4 mm long, the bracteoles deciduous, about 2.5 mm long. Calyx-tube’
densely pubescent, 2 mm long, the perianth segments subequal, 2.5 mm
long, elliptic-ovate, obtuse, pubescent. Fertile stamens 9, the filaments
somewhat pubescent, the anthers of the outer two series introrse, those
of the inner series extrorse; staminodes ovate, short-stipitate, somewhat
pubescent, acute. Ovary narrowly oblong, glabrous; style 1.5 mm long.
Fruit black when dry, glabrous, smooth, oblong or narrowly oblong,
about 3 cm long, 1 cm thick, narrowed upwards, not striate.

Necros, For. Bur. 4236, 4311, 7288, 7311 Everett, April—June, 1906-07, For.
Bur. 7409 Danao, June, 1907, Whitford 1614, May, 1906.

A species common in the dipterocarp forests in northern Negros, well charac-
terized by its leaves being glaucous beneath, and strongly reticulate, the midrib,

nerves, and reticulations on the lower surface of the leaf being brown, in sharp
contrast to the glacous surface. V., Balit, Putian,

258 MERRILL.

Cryptocarya ampla sp. nov.

Arbor alta, inflorescentiis exceptis glabra; foliis oblongis, coriaceis,
nitidis, 18 ad 25 em longis, 5 ad 9 em latis, apice late acuminatis, basi
cuneatis vel rotundatis, concoloribus vel subtus brunneis; nervis utrinque
8 ad 10, prominentibus, obscure anastomosantibus, reticulis validis, densis ;
paniculis axillaribus terminalibusque, foliis subaequalibus, pubescentibus ;
floribus numerosis, pedicellatis, 4 mm longis; fructibus ellipsoideis vel
obovoideis, parce pubescentibus, longitudinaliter striatis, circiter 12 mm
longis.

A tall tree, glabrous throughout except the infloresence. Branches
terete, lenticellate, brownish. Leaves alternate, oblong, 18 to 25 em
long, 5 to 9 em wide, coriaceous, shining, the apex rather broadly
acuminate, the base acute or rounded, about the same color on both
surfaces; nerves 8 to 10 on each side of the midrib, rather prominent,
curved-ascending, obscurely anastomosing, the reticulations beneath dense,
distinct; petioles 1 to 1.5 cm long. Panicles axillary and terminal,
about as long as the leaves, many flowered, the rachis and branches
somewhat pubescent. Flowers white, 4 mm long, pedicellate, the pedicels
pubescent, 1 to 2 mm long, with three small bracteoles at the apex.
Calyx-tube cylindric, sulcate, pubescent, 1.5 to 2 mm long, the perianth-
lobes 6, subequal, pubescent, oblong-obovate, acute or obtuse, 2 mm long.
Fertile stamens 9; anthers all 2-celled, those of the two outer series
introrse, those of the inner series extrorse, the filaments short ; staminodes
acuminate, about 1 mm long. Style 1 mm-long. Fruit ellipsoid or
obovoid, obtuse, somewhat narrowed at both ends, slightly pubescent,
longitudinally striate, about 12 mm long.

Luzon, Province of Rizal, Antipolo and Montalban, For Bur. 463, 397, 2438
Ahern’s collector, February and April, 1904, and January, 1905. T., Bagarilao.
A species well characterized by its large leaves and panicles.

Cryptocarya glauca sp. nov.

Arbor; foliis alternis, coriaceis, oblongo-ovatis vel elliptico-lanceolatis,
usque ad 13 cm longis, 5 em latis, supra brunneis, glabris, nitidis, subtus
plus minus pubescentibus, glaucis ; nervis subtus prominentibus, brunneis,
circiter 8 utrinque; paniculis axillaribus terminalibusque, ferrugineo-
pubescentibus, foliis aequalibus vel longioribus vel interdum brevioribus.

A tree, more or less pubescent. Branches terete, reddish-brown
glabrous, lenticellate, the branchlets ferruginous-pubescent. Leaves
alternate, coriaceous, oblong-ovate or elliptic-lanceolate, the apex shortly
and obtusely acuminate, the base acute, 7 to 13 cm long, 2 to 5 em wide,
the upper surface dark- or pale-brown, glabrous, shining, the lower surface
elaucous, somewhat ferruginous-pubescent, ultimately glabrous or nearly
so; nerves prominent on the lower surface, brown, about 8 on each side
of the midrib, obscurely anastomosing, the reticulations not very distinct,
rather lax; petioles pubescent, 1 to 1.5 em long. Panicles axillary and

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VI. 259

terminal, ferruginous-pubescent, about as long as the leaves, sometimes
longer, sometimes shorter. Flowers subsessile or shortly pedicellate,
pubescent, 3 to 3.5 mm long, the bracteoles small, pubescent. Calyx-tube
cylindric, 1.5 mm long. Perianth-lobes 6, equal, pubescent outside,
elliptic-ovate, obtuse or acute, 2 to 2.5 mm long, 1 to 1.3 mm wide.
Stamens as in other species of the genus.

Luzon, Province of Rizal, Bosoboso, For. Bur. 2981 Ahern’s collector, April,
1905 (type). Apparently the same species, but with small panicles, is repre-
sented by Merrill 2337, from Tanay, the same Province.

Allied to Cryptocarya lauriflora (Blanco) Merr., but differing in many char-
acters, notably in its leaves being very glacous beneath.

LITSEA Lam.

Litsea bicolor sp. nov.

Arbor circiter 8 m alta; ramis griseis, teretibus, glabris, ramuls junior-
ibus dense ferrugineo-pubescentibus ; foliis verticillatis, elliptico-obovatis,
usque ad 35 cm longis, coriaceis, supra glabris, brunneis, nitidis, subtus
albo-glaucescentibus, costa nervisque ferrugineis, plus minus tomentosis,
apice breviter abrupteque acuminatis, basi acutis; fructibus fasciculatis,
ellipsoideis, nigris, nitidis, circiter 2 em longis, obtusis; calycibus auctis,
disciformibus, circiter 1 cm diametro.

A tree about 8 m high. Branches terete, gray, glabrous, the young
branchlets densely ferruginous-pubescent. Leaves verticillate, usually 5
in each whorl, elliptic-obovate, 25 to 35 cm long, 12 to 17 em wide, the
apex shortly and abruptly acuminate, the base acute, coriaceous, the upper
surface glabrous, shining, dark-brown when dry, the lower surface pale,
glaucous, nearly white, somewhat deciduous-tomentose, the nerves and
midrib ferruginous-pubescent ; nerves about 10 on each side of the midrib,
very prominent beneath, curved-ascending, obscurely or not anastomos-
ing, the reticulations lax; petioles ferruginous-pubescent, 3.5 cm long,
stout. Flowers unknown. Fruit fascicled on the branches below the
leaves, the pedicels stout, short, the calyx accrescent, disciform, about 1
em in diameter, glabrous, the fruit ellipsoid, black, shining, glabrous,
obtuse, about 2 cm long.

MrnpaNao, Province of Surigao, Surigao, Bolster 318, April, 1906. N. v.,
Hindang.

A species will characterized by its large, verticillate leaves, which are very
pale beneath.

Litsea hutchinsonii sp. nov.

Arbor circiter 10 m alta; ramis teretibus, glabris, ramulis junioribus
dense ferrugineo-tomentosis; foliis oppositis vel suboppositis, oblongis,
coriaceis, 10 ad 20 cm longis, 5 ad 8 cm latis, concoloribus vel subtus
brunneis, supra glabris, nitidis, subtus parce pubescentibus, nervis utrin-
que circiter 12; inflorescentiis axillaribus, fasciculatis vel glomeratis,
floribus umbellulatis, involucro biseriali, 4-phyllo; staminibus fertilibus
9, antheris omnibus introrsis, 4-locellatis.

260 MERRILL.

A tree about 10 m high, nearly glabrous except the young branchlets
and inflorescence. Branches terete, reddish-brown, glabrous, the young
branchlets rather densely ferruginous-tomentose. Leaves opposite or sub-
opposite, oblong, 10 to 20 cm long, 5 to 8 em wide, coriaceous, of the
same color on both surfaces, or the lower surface brownish, the upper
surface glabrous, shining, the lower very sparingly pubescent, the apex
acute, obtuse, or very obscurely acuminate, the base acute; nerves about
12 on each side of the midrib, beneath very prominent, obscurely anasto-
mosing ; petioles subglabrous, about 1 cm long. Flowers umbellate, the
umbels involucrate, crowded in the leaf-axils, sessile or short-pedunculate,
the inyolucral scales 4, concave, orbicular or obovoid, 4 to 5 mm long,
outside ferruginous-pubescent. Staminate flowers 3 in each umbel, sessile
or subsessile, the tube very short, ferruginous-pubescent outside, the
segments 6, oblong, obtuse, subequal, 3 to 3.5 mm long, 1.5 mm wide,
glabrous or nearly so. Fertile stamens 9, the longer filaments 2.5 mm
long, glabrous or nearly so; anthers all introrse, all 4-locellate.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9148 Whitford &
Hutchinson, January, 1908.

A species apparently allied to Litsea luzonica (Bl.) F.-Vill., but with much
larger, differently shaped leaves. As the genera in this group are defined in the

Nat. Pflanzenfamilien, this species, having 9 fertile stamens, might be referred to
Actinodaphne.

Litsea philippinensis sp. nov.

Arbor usque ad 20 m alta; foliis alternis, coriaceis, nitidis, glabris vel
subglabris, oblongo-ellipticis, 10 ad 20 cm longis, apice rotundatis vel
acutis, basi acutis; nervis utrinque 12 ad 15, prominentibus, reticulis
validis subparallelis; petiolis 1 ad 2.5 cm longis; floribus capitulatis,
capitulis globosis, fasciculatis vel in racemis brevibus axillaribus disposi-
tis; calycis lobis 6, pubescentibus; staminibus fertilibus 12, filamentis
villosis ; fructibus oblongo-ovoideis, glabris, obtusis, circiter 3.5 cm longis,
tubis auctis, cupulatis, circiter 2 em diametro.

A tree 15 to 20 m high. Branches terete, stout, brown, glabrous, the
branchlets somewhat ferruginous-pubescent. Leaves alternate, elliptic-
oblong, 10 to 20 em long, 5 to 10 em wide, coriaceous, somewhat shining,
usually brownish when dry, at least the nerves, not at all glaucous, the
upper surface glabrous, or the midrib sometimes pubescent, the lower
surface glabrous or the midrib and nerves usually more or less pubescent,
the apex rounded or acute, the base usually acute; nerves 12 to 15 on
each side of the midrib, very prominent, parallel, obscurely anastomosing
near the margin of the leaf, the reticulations very distinct, subparallel ;
petioles pubescent, 1 to 2.5 cm long. Flowers in heads, axillary, faseicled
or in short racemes, mostly from the branches below the leaves, in the
axils of fallen leaves, the rachis, when present, stout, not exceeding 1

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 261

em in length. Involucral bracts 4 or 5, pubescent, concave, orbicular or
obovoid, the inner ones thinner, their margins ciliate, strongly imbricate,
6 to 8 flowers in each head. Calyx-tube slender, tubular, 2 mm long,
cylindric, the lobes 6, 3 mm long, pubescent. Fertile stamens 12, their
filaments slender, 3 mm long, clothed with long weak hairs. Fruit oblong-
ovoid, glabrous, obtuse, about 3.5 mm long, the calyx-tube accrescent,
persistent, cup-shaped, about 2 cm in diameter.

Luzon, Province of Bataan, Lamao River, For. Bur. 7503 Curran, September,
1907 (type), For. Bur. 1363, 1794 Borden, July, September, 1904, Whitford 472,
July, 1904, Williams 619, February, 1904: Province of Rizal, Bosoboso, For.
Bur. 3171 Ahern’s collector, July, 1905. Mindoro, Bongabong River, Whitford
1413, April, 1906; Balete, For. Bur. 6161 Merritt, January, 1907. MINDANAO,
Lake Lanao, Camp Keithley, Mrs. Clemens 298, February, 1906.

This species is well represented in our herbarium, especially in material from
the Mount Mariveles region, but the specimen collected by Curran in September,
1907, is the first one secured with mature flowers, thus allowing accurate generic
identification. The specimen from Mindanao, Clemens 298, is immature, being
much more pubescent than the type, and may possibly represent a different species.
It is well characterized by its oblong-elliptic, usually obtuse, strongly nerved
leaves. Apparently allied to Litsea grandis Hook. f.

NEOLITSEA (Benth.) Merr.

Neolitsea villosa (Blume) comb. noy.

Litsea villosa Blume Mus. Bot. Lugd. Bat. 1 (1851) 349; Vidal Rey. Pl. Vase.
Filip. (1886) 226; Ceron Cat. Pl. Herb. (Manila) (1892) 143; Rendle in Journ.
Bot. 34 (1896) 355. Meissn. in DC. Prodr. 15 (1864) 221.

Neolitsea zeylanica Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 57; non
Litsea zeylanica C. & Fr. Nees,

Luzon, Province of Benguet, Mount Santo Tomas, Williams 1986, November,
1904; Loher 4521; Suyoce to Panai, Merrill 4761, November, 1905: Province of
Rizal, Bosoboso, For. Bur. 2674 Ahern’s collector, January, 1905: Province of
Bataan, Mount Mariveles, Whitford 119, 458, May, July, 1904, Elmer 6806
November, 1904, Merrill 3204, October, 1903, For. Bur. 1337, 1586 Borden, 1503
Ahern’s collector, 2617 Meyer, 6262 Curran: Province of Pampanga, Mount Abu.
Bur. Sci. 1958, 1959 Foxworthy, December, 1906: Province of Zambales, Mount
Tapulao, For. Bur, 80738 Curran; Candelaria, For. Bur. 8250 Curran & Merritt,
December, 1907. Minporo, Mount Halecon, For. Bur. 4369 Merritt, Merrill 5666 ;
Mount Malasumbu, For. Bur. 8747 Merritt. MinpANnao, Province of Misamis,
Mount Malindang, For. Bur. 4681, 4695 Mearns & Hutchinson, May, 1906.

This species is widely distributed in the Philippines, but is never found at low
altitudes. It was previously confused by me with Litsea zeylanica C. & Fr. Nees,
but appears not to be that species, but rather Blume’s L. villosa, as identified by
Vidal and Rendle. The specimens cited above show considerable variation, some
of them having the leaves decidedly ferruginous-pilose beneath, but this indument
is deciduous; pubescent and glabrous leaves can frequently be found on the same
specimen. This densely villous form has been identified by Vidal as Litsea
cinnamomea Blume, but whether correctly so or not, the Luzon form so named
appears to me to be the same as L. villosa=Neolitsea villosa. This is doubtless
the form approximately identified by Stepf, in his paper on the flora of Mount
Kinabalu, as Litsea zeylanica, so far as his Philippine reference goes.

262 MERRILL.

MACHILUS Nees.

Machilus nervosa sp. nov.

Arbor glabra circiter 6 m alta; foliis ovato-ellipticis, usque ad 12 em
longis, coriaceis, nitidis, subtus subglaucescentibus, grosse et pulcherrime
reticulatis, nervis urtinque 6 vel 7, prominentibus, elevatis, anastomo-
santibus; paniculis axillaribus terminalibusque, circiter 7 cm longis;
fructibus obovoideis, in sicco nigris, leviter angulatis, circiter 12 mm
longis, perianthii lobis late ovatis, acutis, persistentibus, patulis.

A glabrous tree about 6 m high. Branches terete, lenticellate,
erayish- or reddish-brown. Leaves ovate-elliptic, coriaceous, 7 to 12
em long, 2.5 to 5 em wide, base and apex acute, the upper surface
shining, the lower, at least when young, subglaucous, nerves 6 or: 7
on each side of the midrib, beneath very prominent, elevated, darker-
colored than the surface of the leaf, curved-ascending, anastomosing, the
primary reticulations lax, distinct; petioles 2 to 3 em long. Panicles
axillary and glabrous in fruit, about 7 cm long. Flowers unknown. Fruit
obovoid, red when fresh, black and shining when dry, about 12 mm long,
somewhat keeled or angled and obscurely reticulate, the persistent
perianth-lobes broadly ovate, acute, 1.5 mm long, glabrous or their
margins slightly pubescent, spreading.

Luzon, Province of Benguet Mount Ugo, For. Bur. 10846 Curran, For. Bur.
18009 Merritt, December, 1908, in hardwood forests in ravines at an altitude of
about 2,000 m.

PHOEBE Nees.

Phoebe sterculioides (Elmer) comb. nov. § Huphoebe.

Persea sterculioides Elmer Leafl. Philip. Bot. 1 (1908) 295.

Arbor 7 ad 18 m alta; foliis subcoriaceis, obovatis vel oblanceolatis,
circiter 18 cm longis, 8 cm latis, integris, apice breviter acuminatis,
rariter acutis, basi sensim angustatis, supra glabris, subtus plus minus
glaucescentibus, ferrugineo-pubescentibus ; nervis utrinque 9 ad 11; pani-
culis axillaribus, ferrugineo-pubescentibus, angustis, usque ad 20 em
longis ; calycis lobis 6, subaequalibus vel exterioribus admodum minoribus,
pubescentibus, plus minus acerescentibus, persistentibus, erectis ; stami-
nibus fertilibus 9, 3-seriatis, antheris 4-locellatis, ordinis primi et
secundi introrsis, ordinis tertii extrorsis.

Luzon, Province of Tayabas, Lucban, Hlmer 9121 (cotype), May 1907. Mrn-
poro, near Lake Naujan, For. Br. 6793a, 6824, 6859 Merritt, March-April, 1907.
Necros, 7273, 7287, 7327 Everett, May, 1907. MinpAnao, Lake Lanao, Camp
Keithley Mrs. Clemens 518, April-May, 1906, and six sheets without numbers.

This species appears to be not uncommon in the Philippines, and extends from
central Luzon to Mindanao. In Mindoro it is know as Baticulin, a name applied
to many different species of Lauraceae, in various genera. Specimens of this
plant with young fruit show accrescent, persistent, and appressed calyx segments,
which throws the species into Phoebe, as defined by Pax in Engler & Prantl’s

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 263

Natiirlichen Pflanzenfamilien, although Bentham & Hooker reduce Phoebe to
Persea. In the original description of the species the outer series of stamens are
described as having extrorse anthers, and the other series to have introrse anthers.
I have reéxamined the type number and the reverse seems to be the case, the
outer two series having introrse anthers, and the inner series having extrorse
anthers. Apparently allied to the Malayan Phoebe opaca Blume.

DROSERACEE.
DROSERA Linn.

Drosera burmanni Vahl Symb. 3 (1794) 50; Clarke in Hook. f. Fl. Brit. Ind.
2 (1878) 424; F.-Vill. Nov. App. (1880) 78, Diels in Pflanzenreich 26 (1306) 75.

Luzon, District of Bontoe, For. Bur. 10995 Curran, January, 1909, altitude
about 700 m.

The fourth species of the genus for the Philippines, previously recorded from
Panay by F.-Villar, but his record as to its occurring in the Archipelago not
before verified.

India and Ceylon to southern Japan, south to Malaya and northern Australia.

HAMAMELIDACE 4b.
EMBOLANTHERA gen. noy.

Calycis tubus ovario adnatus, limbus irregulariter 2- vel 3-fidus, denum
circumscisso-deciduus. Petala 5, lineari-elongata, basi utrinque valde
auriculata vel alata. Stamina 5, petalis alternantia, filamentis brevis-
simis; antherae basifixae, loculis in valvis 2 verticaliter dehiscentibus,
connectivo longe producto. Ovarium inferum, 2-loculare; styli 2, sub-
ulati, stigmatibus simplicibus; ovula in loculis solitaria, pendula. Arbor.
Folia alterna, integra vel subintegra, persistentia, ovata vel oblonga,
acuminata. Stipulae lanceolatae, membranaceae, deciduae, pectintae.
Flores in spicas terminales densissime dispositi, calyce stellato-pubescente.

Embolanthera spicata sp. noy.

Arbor circiter 10 m alta, glabra, inflorescentiis exceptis. Ramis
ramulisque tenuibus, griseis vel brunneis; foliis alternis firmiter char-
taceis, ovatis vel oblongis, usque ad 10 em longis, apice breviter acuminatis,
basi rotundatis, acutis vel acuminatis, leviter inaequilateralibus, nitidis.
Spicis foliis aequantibus vel subaequantibus, densis; floribus albis,
5-meris; petalis circiter 2 em longis, 2 mm latis, basi utrinque valde
auriculatis vel alatis; staminibus 5, connectivo longe producto.

A tree about 10 m high, glabrous or ne&rly so except the infloresence.
Branches and branchlets terete, slender, gray or brown, slightly lenti-
cellate. Leaves alternate, ovate to oblong, firmly chartaceous, shining,
7% to 10 em long, 2 to 5 em wide, gradually narrowed upward to the
short-acuminate apex, the base rounded, acute, or slightly acuminate,
usually somewhat inequilateral, the margins entire, recurved, some-
times slightly aculeate-denticulate near the apex; nerves 7 or 8 on

264 MERRILL.

each side of the midrib, prominent beneath, anastomosing, the retic-
wations lax; petioles 5 to 10 mm long. Stipules deciduous, mem-
branaceous, usually lanceolate, acuminate, 5 mm long or less, somewhat
appressed-pubescent on the back, the margins below prominently pectinate,
the tips of the teeth glandular. Spikes as long as the leaves or some-
times shorter, about 2.5 cm in diameter, densely many-flowered, the
rachis somewhat pubescent, the bracteoles linear, pubescent, 1.5 mm long.
Flowers white, sessile. Calyx-tube short, adnate to the ovary, pale-
stellate-pubescent outside, the limb splitting irregularly into two or
three ovate, acute or acuminate lobes, 4 to 5 mm long, membranaceous,
somewhat reticulate-veined, stellate-pubescent outside, finally circumscis-
sile and deciduous. Petals 5, about 2 cm long, 1.8 to 2 mm wide, the basal
3 mm strongly auricled, including the auricles 3 mm wide, narrowed
above the auricles to 1 mm, incurved, and then linear-elongate, mem-
branous, nerved, about 2 mm wide, acuminate. Staminodes none, unless
represented by the auricles to the petals. Stamens 5, alternate with the
petals, and inserted with them; filaments adnate to the corolla and falling
with it, stout, about 1 mm long; anthers ovoid, about 1 mm long, the
cells opening laterally each by two valves, the connective produced as a
straight, 2 mm long, awn. Ovary inferior, 2-celled, each cell with a
single pendulous ovule, the top of the ovary pubescent; styles 2, less
than 1 mm long, slender; stigmas minute. Fruit unknown.

PALAWAN, Victoria Peak, Bur. Sci. 739 Foxworthy, March 24, 1906. A spread-
ing tree about 10 m high, with a trunk 15 to 20 cm in diameter growing on the
river bank at an altitude of about 250 m above the sea.

The genus above proposed is allied to Maingaya Oliver, of Penang and Perak,
and to Loropetalum R. Br., of India and China, differing from both in its spicate
infloresence, and in the strongly auriculate bases of the petals, from the former
also in its different calyx and absence of staminodes, and from the latter also
in its 4-merous flowers. It is apparently more closely allied to Maingaya than
to Loropetalum, but seems to be distinct from both, and from all other hitherto
described genera. But one genus of the family, Sycopsis, was previously known
from the Philippines.

ROSACE AS.
PARINARIUM <Aubl.

Parinarium curranii nom. nov.

Parinarium racemosum Merr. in Govt. Lab. Publ. (Philip.) 17 (1904) 19;
non Vidal Cat. Pl. Prov. Manila)(1880) 29.

Maspate, Merrill 2614, May, 1903 (type). Luzon, Provinee of Albay, For.
Bur, 10575 Curran, June, 1908.

The original name of this species is invalidated by Parinarium racemosum
Vidal, and accordingly the above new name is proposed for it. Vidal’s species
is not enumerated in Index Kewensis.

So bw kek

NEW OR: NOTEWORTHY PHILIPPINE PLANTS, VIL. 265

LEGUMINOSE.
ALBIZZIA Duvrazz.

Albizzia scandens sp. nov.

Frutex scandens, subglabra, usque ad 10 m alta; foliis bipinnatis,
circiter 20 cm longis 4- vel 5-jugatis; foliolis oblongis vel elliptico-
oblongis, 1 ad 2 cm longis, acutis vel apiculatis, leviter inequilateralibus ;
paniculis circiter 20 cm longis, parce pubescentibus; floribus sessilibus,
capitulatis, extus parce pubescentibus; staminibus circiter 20, filamentis
hberis.

A scandent shrub about 10 m high, subglabrous. Branches dark-
colored when dry, terete, lenticellate. Leaves alternate, about 20 em
long, evenly 4- or 5-jugate, the common rachis slightly pubescent, 12 to
15 cm long, with a single large gland on the upper surface near the base,
the stipules, if any, caducous, the base of the petiole subtended by a stout,
somewhat recurved, thickened puvinus about 4 mm long; leaflets 8 to 12
pairs, crowded, oblong or elliptic-oblong, 1 to 2 cm long, 5 to 7 mm
wide, chartaceous, somewhat inequilateral, the base broad, acute, the apex
acute or apiculate, the lower surface paler than the upper, slightly pubes-
cent. Panicles axillary and terminal, 20 cm long or less, slightly pubes-
cent. Flowers white, sessile, in small capitate heads at the ends of the
ultimate branchlets. Calyx about 1.5 mm long, slightly cinereous-pubes-
cent outside, obscurely 5-toothed. Corolla shghtly cinereous-pubescent
outside, 4.5 mm long, the tube very slender below, widened above, the
lobes oblong-ovate, acute, about 1.5 mm long, 1.2 mm wide. Stamens 20
or more; filaments about 12 mm long, free; anthers 0.2 mm long.

PALAWAN, Iwahig, Bur. Sci. 829 Foxworthy, May, 1906.

A species erowing near the sea, climbing over Nylocarpus, allied to Albizzia
myriophylla Roxb., of the Malay Peninsula, but with much fewer jugate leaves,
less numerous and differently shaped leaflets, and free filaments.

BAUHINIA Linn.

Bauhinia monandra Kurz in Journ. As. Soc. Beng. 42° (1873) 73; Forest
Fl. Brit. Burma 1 (1877) 395.

Bauhinia richardiana Wall. in Voigt. Hort. Suburb. Caleutt. (1845) 255, non
DC. fide Prain.

Phanera maculata Rich. ex Teysm. & Binn. Cat. Hort. Bog. (1866) 268, non
Bauhinia maculata Tenore. a

Bauhinia krugii Urban in Ber. Deutsch. Bot. Ges. 3 (1885) 83.

Bauhinia kappleri Sagot in Ann. Sci. Nat. VI 13 (1882) 317; Urban Symb.
Antill. 1 (1899) 83; Perk. Frag. Fl. Philip. (1904) 13; Merr. in Govt. Lab.
Publ. (Philip.) 29 (1905) 17.

Luzon, Manila, Werrill s. n. November, 1903: Province of Pampanga, Arayat,
Warburg, in Herb. Berol., Werrill 3934, October, 1904, Bolster 51, May, 1905:

85754——_5

266 ) MERRILL. ue oe

Province of Rizal, For. Bur. 3248 Ahern’s collector, August, 1905: Province
Bataan, Lamao, For. Bur. 7350 Curran, June, 1907. Bounor, Bur. Sci. 123
Gregor, May, 1906.
There is no doubt but that Bauhinia monandra Kurz is the oldest talid na
for this widely distributed species, and it is accordingly here adopted. Materi =
supplied me by Mr. Craib of the Royal Botanic Garden, Calcutta, as representing le
Kurz’s species is unquestionably the same as our Philippine specimens. In order —
to verify this I wrote to Dr. Prain, Director of the Royal Botanic Gardens, Er
Kew, who has kindly examined all the material in the Kew herbarium, and
who informs me that Bauhinia krugti, and B. kappleri are unquestionably iden- A
tical with B. monandra of Kurz, the reductions being also verified by Mr. W. B.
Hemsley, and Mr. N. E. Brown. My reduction of Phanera maculata Rich., is — ’ ;
based on a specimen so named received from Buitenzorg, taken from a tree in Ey
the botanic garden so labelled, and said to have been received from Bourbon —
under that name. TI suspected that it might be the same as B. maculata Tenore,
but this is not the case, as Tenore’s description does not at all apply to the
present species, although Dr. R. Pampanini of Florence, who kindly supplied *
me with a copy of the original description, informs me that Tenore’s type does
not appear to be extant. The species is now widely distributed in the tropics
of the World, and is probably a native of ezOpical America; it is certainly an
introduced species in the Philippines.

CYNOMETRA Linn.

Cynometra luzoniensis sp. nov.

Arbor glabra, circiter 15 m alta; foliis unifoliolatis, foliolis coriaceis |
vel subcoriaceis, in sicco brunneis, nitidis, oblongis, usque ad 12 cm longis,
reticulatis, apice acute acuminatis, basi late rotundatis vel subcordatis;
fructibus axillaribus, racemosis, compressis, brunneis, circiter 2 cm longis,
verrucoso-lenticellatis, obtusis.

A glabrous tree about 15 m high. Branches light-brown, terete, lenti-
cellate. Leaves alternate, unifoliolate, the petiole stout, rugose, 3 to 4
mm long, the single leaflet sessile, oblong, coriaceous or subcoriaceous,
brown and shining when dry, 5 to 12 em long, 2 to 4.5 em wide, the apex
sharply acuminate, the base rather broad, rounded or slightly cordate;
primary nerves about 8 on each side of the midrib, anastomosing, the
secondary ones and reticulations nearly as distinct, the latter netted,
rather close, distinct on both surfaces. Flowers unknown. Infrutescence
of axillary, solitary racemes, the rachis about 1 cm long, marked by
numerous pedicel-scars, the pedicels about 8 mm in length. Fruits
(immature) compressed, a) SR the dorsal suture nearly straight,
the ventral semicircular, about 2 cm long, 11 to 13 mm wide, brown, by
lenticellate-verrucose, apex and base rounded. 3

Luzon, Province of Tayabas, Laguimanoe, Merrill 2128, April, 1903. 4
A species closely allied to Cynometra simplicifolia Harms, but with larger 4
much more sharply acuminate leaves which are broad and rounded or subcordate
at the base and with quite different, much more prominent venation and retic- ‘af
ulation. The inflorescence of C. simplicifolia is fascieulate and the fruits are —
usually somewhat faleate, the dorsal suture being frequently ineurved.

.<. 38 Tey

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 267

FLEMINGIA Roxb.

Flemingia lineata (Linn.) Roxb. Hort. Beng. (1814) 56; Fl. Ind. 3:341;
Baker in Hook. f. Fl. Brit. Ind. 2 (1876) 228; F.-Vill. Nov. App. (1880) 67.

Hedysarum lineatum Linn. Sp. Pl. (1753) 1054.

Flemingia blancoana Llanos Fragm. (1851) 80; Blanco FI. Filip. ed. 3, 4:62.

Luzon, Province of Bulacan, near Malolos, Mrs. Templeton, February, 1909.

A species previously known from the Philippines only by F.-Villar’s record,
and to which he reduced, and apparently correctly so, Flemingia blancoana Llanos.

India to Siam, the Malay Archipelago and northern Australia.

DESMODIUM Desy.

Desmodium retroflexum (Linn.) DC. Prodr. 2 (1825) 336; Baker in Hook.
f. Fl. Brit. Ind. 2 (1876) 170; Forbes & Hemsl. in Journ. Linn. Soc. Bot. 23
(1887) 176.

Luzon, Province of Nueva Ecija, Cabanatuan, Bur. Sci. 5278 McGregor, Sep-
tember, 1908. :

Himalayan region to Tenasserim and southern China; new to the Philippines.

ERYTHROPHLOEUM Afzel.

Erythrophloeum densiflorum (Elm.) comb. nov.

Cynometra densiflora Elm. Leafl. Philip. Bot. 1 (1907) 222.

Arbor glabra, inflorescentiis exceptis, usque ad 25 m alta; foliis alternis
abrupte bipinnatis, pinnis 1- vel 2-jugatis; foliolis oppositis, abrupte
pinnatis, 3- ad 5-jugatis, subcoriaceis, nitidis, in sicco brunneis vel
pallidis, usque ad 15 cm longis, valde acuminatis, basi rotundatis vel
acutis, leviter inaequalibus; floribus parvis, spicatis, spicis pubescentibus,
in paniculis terminalibus vel subterminalibus dispositis; petalis 5, im-
bricatis, circiter 4 mm longis; ovarium stipitatum, 4-ovulatum ; legumi-
nibus ligneis, oblongis vel anguste oblongo-obovatis, obtusis, compressis,
basi angustatis, longitudinaliter laxe reticulato-striatis, rectis vel leviter
falcatis, dehiscentibus, usque ad 18 cm longis, 4.5 cm latis; seminibus 1
ad 4, compressis, orbicularibus, circiter 3 cm diametro.

The type of this species was collected by-Mr. Elmer No. 9013, near Lucban,
Province of Tayabas, Luzon, in flower, and described by him as Cynometra densi-
flora. It is represented by the following additional material; Luzon, Province of
Tayabas, Bulin, For. Bur, 10272 Curran, with nearly mature fruit; Guinayangan,
For. Bur. 12507 Rosenbluth; Dugatan, For. Bur. 10215 Curran; Apad, For. Bur.
11513 Whitford. A sterile specimen from Mindanao, For. Bur. 9163 Whitford &
Hutchinson, may be referable here. Locally known to the Tagalogs as Camatog
or Calamantao.

A most interesting addition to our knowledge of the Philippine flora, the genus,
up to the present time consisting of about six species, mostly in Africa and
Madagascar, one in Australia and one in China.

KINGIODENDRON Harms.

Kingiodendron alternifolium (Elmer) Merrill & Rolfe, comb. nov.
Cynometra alternifolia Elmer Leafl. Philip. Bot. 1 (1907) 223.
Hardwickia alternifolia Elmer 1. c. (1908) 362.

Arbor magna, glabra: foliis imparipinnatis, alternis, rariter oppositis,

268 MERRILL.

foliolis 3 ad 5, coriaceis, elliptico-ovatis vel oblongo-ellipticis, alternis,
valde glanduloso-punctatis, plus minus inaequilateralibus, apice acu-
minatis, basi acutis vel obtusis; paniculis axillaribus; calycibus 5-
vel 4-meris, lobis valde imbricatis, glanduloso-punctatis ; ovario sessile,
dense villoso; fructibus suborbicularibus, ellipticis vel obovoideis, crassis,
leviter compressis, byunneis, ligneis, 3 ad + em longis, circiter 3 em latis,

2.5 em erassis, Indehiscentibus; seminibus solitariis, albumine ruminato.

The type of this species is Elmer 7366 from Leyte, the specimens with im-
mature fruits and without flowers, which perhaps accounts for its originally being
described as Cynometra. Later Mr. Elmer transferred it to Hardwickia, follow-
ing the conception of that genus as defined by Bentham and Hooker. Harms has
however proposed the new genus HKingiodendron, based on Hardiwickia pinnata

Roxb., to which the present species is allied. As we consider Aingiodendron

to be a valid genus, the Philippine species is here transferred as the second species’
g PI ]

of the genus.

In addition to the type, cited above, the species is represented by the following
specimens: Luzon, Province of Tayabas, Guinayangan, Bath s. n., May, 1904
(fruit), Mulanaw and San Narciso, For. Bur. 10327, 10354 Curran, April, 1908
(fruit): Province of Camarines, For. Bur. 10671 Curran (sterile): Province of
Sorsogon, For. Bur. 10624 Curran (sterile). MaAspatre, Merrill 2761, June, 1903
(fruit), Whitford 1679, October, 1906 (sterile). Ticao, For. Bur. 1084 Clark,
May, 1904 (fruit). PANAy, Miagao, Vidal 2468 in Herb. Kew. MinpaAnao, Dis-
trict of Zamboanga, For. Bur. 6567 Hutchinson, March, 1907 (immature fruit) ;
Port Banga, For. Bur. 9301, 9007, 11036 Whitford & Hutchinson (in flower).

The present species is a timber tree of considerable importance, being com-
mercially known as Batete. The wood structure has been considered by Fox-
worthy ° who states that it is much Hke that of Sindora supa Merr., in structure,
but is darker and with a greater amount of oil. It is widely distributed in the
central and southern Philippines, and is known by the following native names
V., Batete; T.. Dangay; B., Salalangin; in Zamboanga as Palo ‘Maria and
Bitanhol, but these two names belong properly to Calophyllum.

MEZONEURUM Desf.

Mezoneurum latisiliquum (Cav.) comb. nov.

Bauhinia % latisliqua Cav. leon. 5 (1799) 5, t. 408, in part, excluding deserip-
tion and figure of leaves.

Vezoneurum glabrum Desf. in Mém. Mus. Paris 4 (1818) 245, ¢#. 10; DC.
Prodr.. 2 (1825) 484; Baker in Hook. f. Fl. Brit. Ind. 2 (1878) 258; F.-Vill.
Nov. App. (1880) 70; Miq. Fl. Ind. Bat. 1* (1855) 103; Vidal Phan. Cuming.
Philip. (1885) 110, Rev. Pl. Vase. Filip. (1886) 114; Merr. in Philip. Journ. Sei.
1 (1906) Suppl. 64.

Caesalpinia torquata Blanco Fl. Filip. (1837) 336.

Vezoneurum procumbens Blanco |. c. ed. 2 (1845) 235, ed. 3, 2:73.

Luzon, Province of Pampanga, Arayat, Merrill 1438: Province of Rizal,
Merrill 1711: Province of Bataan, For. Bur. 5958 Curran; Whitford s. n.; Wil-
liams 701: Provinee of Laguna, Los Baiios, Elmer; Bur. Sei. 6095 Robinson.
MINDORO, Paluan, Merrill 956; For. Bur. 9748 Merritt. Neeros, For. Bur, 7815
Hverett. BAsiran, For. Bur. 3978 Hutchinson; Hallier. MINDANAO, Province of

“This Journal, 2 (1907) Botany 376.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 269

Surigao, Bolster 366; Lake Lanao, Camp Keithley, Mrs. Clemens 262; District of
Davao, Copeland 470.

A common and widely distributed species in the Philippines, for which the
earliest specific name is here adopted. Cavanilles’ Bauhinia ? latisiliqua is a
mixture, the leaves being those of a true Bauhinia, but the fruit manifestly a
Mezoneurum. It was based on material collected in the Philippines by Née, the
fruit, at least, coming from the town of Cavinti in Laguna Province, Luzon. As
the specific name was taken from fruit characters, and as the fruit as figured and
described is manifestly Mezoneurum, I consider that Bauhinia ?
typified by the fruit, and that it should be considered as Mezoneurum. Loeal
names, T., Camut pusa, Camut cabag, Cabit cabag, Sagnit, Sapnit; V., Tugabang,
Ugabang, Sampinit; in Basilan Sokit, Sampinit.

Timor, and according to Baker, in Tenasserim.

latisiliqua is

SESBANIA Pers.

Sesbania roxburghii nom. noy.

Aeschynomene paludosa Roxb. Hort. Beng. (1814) 56, nomen, FI, Ind. 3 (1832)
333,

Sesbania paludosa Prain in Journ. As. Soc. Beng. 6° (1897) 82, non Jacq.,
1825.

Sesbania grandiflora Miq. Fl. Ind. Bat. 11 (1855) 333, non Pers.

Sesbania cochinchinensis Kurz in Journ. As. Soe. Beng. 45? (1876) 271, non DC.

Sesbania aculeata var. paludosa Baker in Hook. f. Fl. Brit. Ind. 2 (1876) 115;
in part, and excluding the synonym Aeschynomene uliginosa.

Luzon, Province of Laguna, Siniloan, Bur. Sci. 6530 Robinson; Santa Maria
Mavitae, For. Bur. 10098 Curran, in shallow water in Lake Bay, T., Balakla.

Bengal to Burma, southern China and Java.

As there appears to be no valid specific name for this species, the above is
here proposed.

RUTACEZ.
EVODIA Forst.

Evodia monophylla sp. nov.

Arbuscula glabra circiter 3 m alta; foliis oppositis, unifoliolatis,
fololis 6 ad 13 cm longis, elliptico-lanceolatis, firmiter chartaceis,
utrinque pallidis nitidisque, apice, acuminatis, basi acutis; cymis pedun-
culatis, circiter 1 cm longis; floribus 4-meris, ovario obtuse 4-angulato.

A glabrous shrub about 3 m high. Branches terete, slender, grayish
or grayish-brown, shining, striate when dry. Leaves opposite, unifolio-
late, the petiole 0.8 to 3 em long, the petiolule short; leaflet firmly
chartaceous, elliptic-lanceolate, pale and shining on both surfaces, 6 to
13 em long, 2 to 4.5 em wide, entire, glabrous, the apex acuminate,
acumen blunt, the base acute; primary nerves 10 to 13 on each side of
the midrib, rather distant, anastomosing, the secondary nerves and rather
lax reticulations almost as prominent. Cymes axillary, solitary, ped-
uncled, about 1 cm long, when young slightly puberulent. Pedicels
about 2 mm long. Calyx-lobes 4, ovate, 1 mm long. Petals 4, oblong-
elliptic, acute, punctate, 2.5 mm long, 1.3 mm wide. Stamens 4, al-

nN

ternating with the petals; filaments 1.7 mm long; anthers 0.8 mm long.

270 MERRILL.

Disk prominent, obtusely 4-angled, somewhat undulate. Ovary with
four obtuse lobes, sunk in the disk; style very short, simple. Immature
carples glabrous.

Minporo, Mount Haleon, Merrill 6169, November 9, 1906, in forests at about
750 m altitude; flowers pink. 7
A species readily recognizable by its unifoliolate leaves, pale, shining, elliptic- : o
lanceolate, acuminate leaflets, and very short axillary cymes. a

is

MELIACEZ.
AGLAIA Lour. ic.

Aglaia badia sp. nov. § Huaglaia. 5

Arbor 20 ad 25 m alta, ramulis paniculisque densissime badio-lepidotis, |
subnitentibus ; foliis alternis, imparipinnatis, 4- vel 5-jugatis, 20 ad 25
cm longis; foliolis oppositis, in sicco brunneis, utrinque decidue brunneo-
lepidotis; paniculis axillaribus, folia subaequantibus, pedunculatis; flor-
ibus racemoso-dispositis, minutis, 5-meris. .

A tree 20 to 25 m high the branches, petioles and infloresence
somewhat shining, minutely and very densely brown-lepidote. Leaves
alternate, odd-pinnate, 4- or 5-jugate, 20 to 25 cm long, the rachis and Pa
petiolules minutely and densely lepidote; leaflets opposite, brown when
dry, chartaceous, oblong to elliptic-oblong, somewhat shining, deciduously
lepidote on both surfaces, 5 to 10 cm long, 2.5 to 4.5 em wide, the
terminal one sometimes larger, equilateral, the lateral ones somewhat
inequilateral ; nerves 10 to 12 on each side of the midrib, not prominent, 5
the reticulations fine, rather lax; petiolules 8 mm long or less. Panicles
axillary, peduncled, nearly as long as the leaves, branched in the upper 7
two-thirds, the rachis, branches, branchlets and calyces minutely and
densely brown-lepidote, shining, the lower primary branches 7 cm long,
the upper ones shorter. Flowers minute, racemosely arranged on the
ultimate branchlets, their pedicels 1 to 2 mm long. Calyx about 1 mm |
long, densely lepidote, its teeth broad, rounded. Petals about 1 mm long, “
glabrous. Staminal tube 0.5 mm long, truncate or obscurely toothed
free, glabrous ; anthers 5, borne on the upper part of the tube, included.

Luzon, Province of Cagayan, San Vicente, For. Bur. 7082, 11305 Klemme, May,

1907, April, 1908, in dense flat- and hill-forests, altitude 5 to 30 m. Cag., Salotot;
Neg., Masaleng.

A species in the group with Aglaia hexandra Turez., but not closely allied to
it, well characterized by its brownish color and densely disposed, minute, some-
what shining, lepidote scales. .

Aglaia bicolor sp. noy. § Huaglaia.

Arbor 15 ad 25 m alta, ramulis foliolis subtus inflorescentiisque den-
sissime cupreo-lepidotis, nitidis; foliis alternis, imparipinnatis, 30 ad 45
cm longis, foliolis 9 ad 11, alternis firmiter chartaceis vel subcoriaceis, .
elliptico-oblongis vel oblongo-lanceolatis, utrinque acuminatis, base valde

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. Ate

inaequaliteralibus, supra glabris; paniculis folia subaequantibus, multi-
floris; floribus minutis, 5—meris, pedicellatis, in ranvulis ultimis racemoso-
dispositis; fructibus ellipsoideis vel anguste obovoideis, circiter 1.5 cm
longis, 1—locellatis.

A tree 15 to 25 m high, the branchlets, inflorescence, rachis of the
leaves, petiolules and under surfaces of the leaflets very densely cupreous-
lepidote, shining. Branches rather stout, lepidote, brownish, striate,
glabrous. Leaves alternate, odd-pinnate, 30 to 45 cm long, the petiole
and rachis cupreous-lepidote, terete; leaflets 9 to 11, alternate, petiolate,
firmly chartaceous or subcoriaceous, elliptic-oblong to oblong-lanceolate,
8 to 15 cm long, 3 to 6 em wide, the apex rather abruptly acuminate, the
base very strongly inequilateral, acuminate, the upper surface glabrous,
rarely with very few scattered scales, shining, grayish-green when dry,
the lower very densely cupreous-lepidote, also shining; nerves 10 to 12
on each side of the midrib, straight, rather distinct, obscurely anastomos-
ing, the reticulations obsolete; petiolules 1 to 1.5 cm long. Panicles
axillary, about as long as the leaves in anthesis, shorter in bud, all parts
very densely cupreous-lepidote, shining, many-flowered, the lower primary
branches often 20 cm in length. Flowers minute, pedicellate, race-
mosely disposed on the ultimate branchlets, their pedicels 1 mm long
or less. Calyx obscurely toothed, densely lepidote. Petals 5, orbicular,
glabrous, 0.6 mm in diameter or less. Staminal tube obovoid, 0.7 mm
long, glabrous, free from the petals, shghtly toothed. Anthers 5, in-
serted near the apex of the tube, sessile, inflexed, 0.2 mm long. Fruits
ellipsoid or narrowly obovoid, 1.5 cm long, densely cupreous-lepidote,
shining, 1-celled, 1—seeded.

Luzon, Province of Cagayan, San Vicente, For. Bur. 4288 Klemme, June,
1906, (type): Province of Ilocos Sur, San Quintin, For. Bur. 7117 Klemme,
April, 1907: Province of Rizal, Antipolo, Merrill 1659, March, 1903, For. Bur.
413 Ahern’s collector, February, 1904.

A species manifestly allied to Aglaia denticulata Turcz., but distinguished by
its more numerous leaflets, which are very much more densely lepidote beneath,
longer panicles and much smaller flowers. Tagalog (Rizal) Salamongay; Ne-
grito (Cagayan) Matanaota; Hocano, Sal-lapugud.

Aglaia everettii sp. nov. § Hearnia.

Arbor 10 ad 20 m alta; foliis alternis, circiter 30 cm longis, im-
paripinnatis, 2—jugatis, glabris; foliolis oppositis, chartaceis vel sub-
membranaceis, in sicco pallidis, nitidis, late ellipticis vel elliptico-obovatis,
usque ad 20 cm longis, apice breviter acuminatis, basi rotundatis vel
acutis, nervis utrinque 10 ad 14; paniculis axillaribus, foliis longioribus,
multifloris, plus minus ferrugineo-lepidotis, pubescentibus; floribus 5-
meris, pedicellatis, in ramulis ultimis dense racemose dispositis.

A tree 10 to 20 m high, glabrous except the inflorescence. Branches
terete, light-gray, shining, lenticellate. Leaves alternate, about 30 cm

212 MERRILL. ~ at)

long, odd-pinnate, 2-jugate ; leaflets opposite, chartaceous to submet
naceous, when dry pale and shining, glabrous, broadly elliptic to ellipt
obovate, 12 to 20 cm long, 7 to 10 em wide; nerves 10 to 14 on ea ch
side of the midrib, distinct, anastomosing near the margin, the reticule ere
tions fine, distinct ; petiolules stout, 1 to 1.4 em long. Panicles axilla y =
equaling or longer than the leaves, 50 cm long or less, ferruginous-lepidote |
and somewhat ciliate-pubescent, the lower branches frequently 20 em —
in length. Flowers rather densely racemosely arranged on the ultimate
branchlets, the pedicels about 1 mm long. Sepals 5, free or nearly
so, orbicular-ovate, obtuse, 1 mm long, their margins ciliate. Petals —
5, ovate or orbicular-ovate, glabrous, obtuse, 2 mm long. Staminal tube |
depressed-globose, glabrous, free from the petals, truncate. Stamens 5, —
inserted on the edge of the tube, inflexed. Fruit narrowly ellipsoid,
rounded at the apex, brown, 3 to 4 em long, 1- or 2-celled, each cell
with a single seed.
Necros, Province of Negros Occidental, Painguion River, For. Bur. 7319
Everett, March, 1907 (type); Cadiz, For. Bur. 15035 Danao, March, 1908. CrsBu,
Mount Licos, For. Bur. 6452 Everett, February, 1907. Visayan Bubua, Bunguas.

AZADIRACHTA A. Juss.

Azadirachta integrifoliola sp. noy.

Arbor glabra, circiter 30 m alta; foliolis 12 ad 16, inferioribus alternis,
superioribus suboppositis, ovato-lanceolatis, integris, membranaceis, acu-
minatis vel acutis, basi inaequilateralibus, rotundatis; paniculis foliis
longioribus, anguste pyramidatis, circiter 45 em longis; floribus pedicel-
latis, circiter 7 mm longis; ovario glabro, 3-loculari. se

A tree about 30 m high, glabrous. Leaves equally pinnate, about ,
30 cm long, glabrous: leaflets 12 to 16, ovate-lanceolate, the lower ones _
alternate, the upper opposite or subopposite, membranaceous, glabrous,
shining above, 6 to 9 em long, 2.5 to 3.5 em wide, entire, the apex acute
or somewhat acuminate, the base rather strongly inequilateral, broad and
rounded on one side of the midrib, narrower and usually acute on the
other: nerves 6 to 8 on each side of the midrib, distinct beneath, ob-
scurely anastomosing, the reticulations lax, obscure; petiolules very
short, the leaflets almost sessile. Panicles longer than the leaves, about
45 cm long, glabrous, narrowly pyramidal, many flowered. Sepals ovate,
acuminate, about 1 mm long, the margins slightly ciliate. Petals free,
oblong-lanceolate, acute or obtuse, about 7 mm long, 1.8 to 2 mm wide,
recurved, nearly glabrous outside, inside puberulent, the margins slightly —
puberulent-ciliate. Staminal tube cylindric, 6 mm long, glabrous out- _
side, slightly narrowed in the middle, somewhat hairy in the upper
half inside, the apex with 10 small teeth. Stamens 10, inserted opposite
the teeth, the anthers sessile, 0.8 mm long, their apices slightly exserted.
Disk none. Ovary ovoid, glabrous, 3-celled, each cell with two collateral”
ovules; style 4 mm long; stigma slightly 3-lobed.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 213

PALAWAN, Maleampo, For. Bur. 11248 Manalo, March, 1908, said by the col-
lector to be common in old clearings, and locally known as Marango. The height
is given as 30 m and the diameter of the trunk as 90 em, while the timber is
said to be used for construction purposes.

Azadirachta A. Juss. has previously been a monotypic genus, consisting of
the one species A. indica A. Juss. extending from India to Java. The Species
above described can at once be distinguished by its entire leaflets, its panicles
much longer than the leaves, not shorter as in A. indica, and its longer flowers.

HEYNEA Roxb.

Heynea sumatrana Mig. Ann. Mus. Bot. 4 (1868) 60; C. DC. Monog. Phan,
1 (1878) 714.

Scutinanthe engleri Elm. Leafl. Philip. Bot. 1 (1908) 298.

Luzon, Province of Tayabas, Lueban, Elmer 9179; Atimonan, For. Bur. 6702
Kobbe. Basiuan, Hallier s. n.

The first representative of the genus to be found in the Philippines, for
Trichilia rimosa Blanco, although reduced to Heynea trijuga Roxb., by C. De-
Candole, is, I feel certain, not a Heynea. The type of Scutinanthe engleri Elm.,
is a specimen with very young fruits, mature fruits or flowers not present, and
is manifestly referable to Heynea and not a Burseraceous plant, where it was
placed by Mr. Elmer. The Philippine specimens agree closely with those of
H. sumatrana Miq., taken from cultivated trees in the Buitenzorge Botanical
Garden, and there seems to be little doubt as to their specific identity. Miquel’s
species has, however, been reduced by King to Heynea trijuga var. multijuga
C. DC., but a specimen of Wallich 1259 from Penang, the type number of this
variety, is in our herbarium and does not agree with our material of H. swmatrana.
nor with Miquel’s description, and it seems that while Heynea sumatrana Miq.,
is not the same as H. trijuga var. multijuga, still it may not be specifically distinet
from H. trijuga Roxb.

Sumatra.

DYSOXYLUM Blume.

Dysoxylum klemmei sp. nov. § Eudysoxylum.

Arbor subglabra, circiter 16 m alta; foliis alternis, imparipinnatis, 40
ad 45 cm longis, rhachidibus densissime ferrugineo-puberulis; foliolis
cireiter 15 cm longis, alternis vel suboppositis, circiter 17, glabris, nitidis,
brunneis, apice acuminatis, basi acutis, valde inaequilateralibus, reticulis
obsoletis ; paniculis axillaribus folia subaequantibus, densissime ferrugi-
neo-puberulis ; floribus 4-meris, circiter 5 mm longis.

.A tree, nearly glabrous except the petioles, young branches and in-
floresence. Leaves alternate, 40 to 50 cm long, odd-pinnate, the leaflets
about 17, the rachis densely ferruginous-puberulent ; leaflets alternate or
subopposite, oblong, about 15 cm long, 4 to 5 em wide, subcoriaceous,
shining, dark-brown when dry, glabrous, the apex acuminate, the base
very strongly inequilateral, one side much narrower than the other and
always acute or acuminate, the other side often rounded, sometimes acute
and extending farther down the midrib; nerves 8 to 13 on each side of
the midrib, not prominent, the reticulations obsolete; petiolules about 1
cm long, glabrous or somewhat puberulent. Panicles axillary, about as

274 MERRILL.

long as the leaves, densely ferruginous-puberulent, peduncled, the lower

branches about 7 cm long, the upper ones shorter. Flowers pedicelled, —

glabrous. Calyx about 2.5 mm in diameter, short, obscurely 4-toothed.
Petals 4, oblong, 4 to 5 mm long, about 1.4 mm wide, glabrous. Staminal
tube cyclindric, 3 mm long, irregularly toothed at the apex; anthers 8,
oblong, 1 mm long, borne at the top of the tube, included. Disk eylindrie,
1 mm long, free, the apex densely ferruginous-hirsute. Ovary somewhat
ferruginous-hirsute ; style 2.5 mm long, glabrous.

Luzon, Province of Cagayan, San Vicente, For. Bur. 7079 Klemme, May, 1907,
in dense hill-forests at an altitude of about 30 m. Neg., Tibungao.

A species in the group with Dysoxylum thyrsoideum Hiern., but very different
from that species and apparently also from all other allied forms.

EUPHORBIACEZE.
AGROSTISTACHYS Dalz.

Agrostistachys pubescens sp. nov.

Arbor parva, ramulis, subtus foliis, inflorescentiisque plus minus pubes-
centibus; foliis oblongo-obovatis, membranaceis vel chartaceis, circiter
22 em longis, apice acuminatis, basi angustatis, cuneatis, minute irre-
gulariter glanduloso-serratis vel subintegris, supra glabris, subtus ad
costam nervosque plus minus villosis; inflorescentiis foliis longioribus ;
fructibus dense pubescentibus.

A small tree, the branchlets, inflorescence and the under surface of the
leaves more or less pubescent. Branches terete, grayish-brown, pubescent,
sparingly lenticellate. Leaves membranaceous or chartaceous, 18 to 25
em long, 9 to 11 em wide, oblong-obovate, the apex rather abruptly
acuminate, gradually narrowed from just below the middle to the cuneate
base, subentire or slightly and distantly irregularly glandular-serrate, the
upper surface glabrous, shining, the lower more or less villous on the
midrib and lateral nerves; nerves prominent, parallel, 25 to 30 on each
side of the midrib, the reticulations distinct, parallel; petioles stout,
pubescent, less than 5 mm long. Inflorescence axillary, solitary, exceed-
ing the leaves, incipient spikes very short, composed of numerous imbri-
cate, ovate, acuminate, more or less pubescent bracts mostly less than 1
mm in length, not distichous; mature inflorescence slightly pubescent,
the rachis 40 to 60 cm long. Flowers unknown. Fruits dry, subglobose,
dehiscent, 3-celled, about 8 or 9 mm in diameter, outside densely oliva-
ceous- or yellowish-pubescent ; seeds subglobose, about 5 mm in diameter,
shining.

Luzon, Province of Bataan, Mabayo, For. Bur. 5940 Curran, January, 1907,
in bamboo thickets along streams.

A species readily recognizable in the genus by its pubescence, the second one to
be found in the Philippines.

ae

‘

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 275

ANTIDESMA Burm.

Antidesma cordato-stipulaceum sp. nov.

Arbuscula glabra 2 ad 3 m alta; foliis oblongis, chartaceis vel sub-
coriaceis, nitidis, prominente acuminatis, apiculatis, usque ad 20 cm
longis, nervis utrinque 12 ad 14; stipulis persistentibus, foliaceis, late
ovatis vel suborbicularibus, basi cordatis, 1 ad 2.5 em longis; floribus
masculinis spicatis, 4- vel 3-meris, calycibus subtruncatis; staminibus 3,
rariter 4; floribus femineis racemosis, calycibus breviter obscureque
5-dentatis, stigmatibus subterminalibus; fructibus in sicco compressis,
1 em longis, basi rotundatis, apice obscure acuminatis.

A glabrous shrub 2 to 3 m high. Branches slender, terete, glabrous,
light-gray. Leaves oblong, chartaceous or subcoriaceous, shining, 12 to
20 em long, 4 to 8 cm wide, entire, the apex rather prominently acuminate,
the acumen blunt, apiculate, base rounded or acute; nerves 12 to 14 on
each side of the midrib, prominent beneath, curved-spreading, anas-
tomosing ; petioles about 1 cm long; stipules foliaceous, broadly ovate to
suborbicular, 1 to 2.5 cm long, 1.5 em wide or less, apex acute, rounded,
or sometimes somewhat acuminate, base broad, cordate, usually clasping
the stem. Staminate inflorescence very slender, of 5 to 10 em long more
or less fascicled spikes from the leaf axils, or from axils below the leaves ;
flowers minute, about 1.2 mm in diameter, glabrous, sessile, solitary,
more or less scattered, the calyx somewhat saucer-shaped, subtruncate or
very obscurely toothed; stamens usually 3, rarely 4, their filaments 0.5
mm long. Pistillate inflorescence of slender, axillary, solitary racemes,
15 cm long or less; flowers more or less scattered in the lower portion,
somewhat crowded above, their pedicels 1 mm long or less, each subtended
by a minute, oblong, 0.5 mm long bracteole; calyx somewhat cupshaped,
obscurely 5-toothed, the margins minutely pubescent; ovary glabrous,
ovoid; styles terminal or subterminal. Fruit, when dry, compressed,
narrowly ovate, somewhat inequilateral, base rounded, apex somewhat
acuminate, 1 cm long, reticulate, reddish, crowned by the somewhat
falcate styles.

Mrinporo, Baco River, Merrill 1807, 4048, the former with pistillate flowers,
the latter with staminate ones, McGregor 179, 311, the former with pistillate
flowers, the latter with fruits; Subaan, Yor. Bur. 6794 Merritt, with staminate
flowers.

A very characteristic species, readily recognized by its comparatively large,
persistent, ovate, cordate, foliaceous stipules, as well as by its very slender
inflorescence, the staminate being spicate, and the pistillate racemose.

Antidesma subcordatum sp. nov.

Arbor 10 ad 15 m alta, ramulis, inflorescentiis, foliisque plus minus
pubescentibus ; foliis oblongis vel ovato-oblongis, usque ad 12 em longis,
membranaceis vel chartaceis, plus minus nitidis, utrinque dense molli-
terque pubescentibus, apice acuminatis, basi cordatis, subcordatis vel

276 MERRILL. eae

rariter rotundatis, nervis utrinque circiter 8; inflorescentiis masculi
spicatis, spicis densis, densissime pubescentibus, solitariis vel fasciculat:
terminalibus, 3 ad 7 cm longis, circiter 3 mm diametro, floribus 5- rariter
G-meris, calycis lobis brevibus, densissime pubescentibus; inflorescentiis—
femineis spicatis, spicis terminalibus, solitariis ; stigmatibus terminalibus.
A tree more or less pubescent in all parts, 10 to 15 m high. Branches _
terete, reddish- or grayish-brown, nearly glabrous, lenticellate, the younger |
branchlets densely and softly pubescent. Leaves oblong to ovate- oblonga
6 to 12 cm long, 3 to 6 cm wide, membranaceous or subchartaceous, more
or less shining, in the typical form densely and softly pubescent on both
surfaces, entire, the apex rather abruptly and somewhat broadly acu- —
minate, the acumen apiculate, base rather broad, cordate or subcordate, ba
sometimes rounded; neryes about 8 on each side of the midrib, distinct,
somewhat ascending, laxly anastomosing; petioles softly pubescent, 5 to
10 mm long. Staminate inflorescence spicate, spikes solitary or fascicled,
usually terminating the short lateral branchlets, very dense, cylindric,
very densely pubescent, 3 to 7 cm long, about 3 mm in diameter; calyx
lobes 5, short, narrowly ovate, acute or obtuse, about 0.6 mm long, very
densely pubescent. Stamens 4 to 6; filaments glabrous, 1.5 to 2 mm
long. Rudimentary ovary densely lanate. Pistillate inflorescence race-
mose, the racemes terminating the short lateral branches, solitary, 4 to 6
em long, the rachis, calyces and bracteoles densely pubescent, the pedicels —
nearly glabrous; bracteoles narrowly ovate, more or less acuminate, 1 mm
long; pedicels glabrous or nearly so, 1 to 1.5mm long. Calyx cylindric-
cupshaped, 1 mm in diameter, pubescent, obscurely 5-toothed. Ovary _
narrowly elliptic, equilateral, with few scattered hairs; styles terminal, —
each cleft, and the arms also cleft.

Luzon, Province of Rizal, Bosoboso, Bur. Sci. 1114, 4564 Ramos, the former
with staminate, the latter with pistillate flowers; from the same province also a
Merrill 2813, and For. Bur. 3160 Ahern’s collector. *

A species well characterized by its very dense cylindric staminate spikes
which are very densely pubescent, its more lax racemose pistillate spikes, and
its softly pubescent leaves which are acuminate, the base cordate or subcordate
and usually oblong or ovate-oblong. Probably as closely allied to A. ghaesembilla
Gaertn., as to any other species, but very different in vegetative characters.

A

a
>

ACTEPHILA Blume.

Actephila dispersa (Elmer) comb. nov.

Pimelodendron dispersum Elmer Leafl. Philip. Bot. 1 (1908) 308.

Arbuscula glabra, circiter 3 m alta; foliis submembranaceis, oblongis ‘a
vel obovatis, integris, nitidis, 15 ad 30 cm longis, apice acutis, vel obscure
acuminatis, vel obtusis, basi subacutis vel abrupte obtusis, nervis utrinque — .
9 ad 13; petiolo 2 ad 4 em longo; floribus ignotis ; fenetibys axillaribus, —
longe meduni ulatis, pendulis, depresao-dlobosia, circiter 2 cm diene
obscure trigonis, trilocularibus, loculidice dehiscentibus ; seminibus in,
loculis 2

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. at

Leyte, Palo, Elmer 7245, January, 1906, type number,

This species was originally described by Mr. Elmer as a Pimelodendron, but the
presence of two seeds in each cell of the mature fruit excludes it at once from
that genus and all allied ones. It seems, however, to be referable to Actephiia,
and to be closely allied to Actephila gigantifolia Koorders of Celebes, but the
latter species is described as having the leaves abruptly long-acuminate. The
five small calyx-segments described by Mr. Elmer are the squamiform petals, the
sepals and petals persisting on the mature fruit.

BACCAUREA Lour.

B. lanceolata (Miq.) Muell. Arg. in DC. Prodr. 15’ (1862) 457,

Hedycarpus lanceolatus Miq. Fl. Ind. Bat. 1° (1859) 359.

PALAWAN, San Antonio Bay, Merrill 853, February, 1903.

Java and Borneo.

The above specimen has somewhat larger leaves than the plants described by
Muller-Arg., but seems to be the same species. New to the Philippines.

BLACHIA Baill.

Blachia philippinensis sp. nov.

Arbor glabra, monoica, circiter 5 m alta; foliis alternis, chartaceis vel
submembranaceis, oblongo-ovatis, usque ad 13 cm longis, acuminatis, basi
acutis vel obtusis, nervis utrinque circiter 6; floribus umbellatis, umbellis
axillaribus terminalibusque ; petalis obovatis, hyalinis et glandulis aequi-
longis ; staminibus 20, liberis; stylis revolutis.

A glabrous monoecious tree about 5 m high. Branches slender, terete,
brownish, the branchlets reddish-brown. Leaves alternate, chartaceous
or submembranaceous, oblong-ovate, 7 to 13 em long, 2.5 to 4.5 cm wide,
brown or greenish when dry, shining, the apex rather prominently
acuminate, the base acute or obtuse; nerves about 6 on each side of the
midrib, the reticulations lax; petioles 5 mm long or less. Inflorescence
axillary and terminal, the staminate umbels long-peduncled, the peduncles
slender, 2 to 3 em long, each bearing about 12 flowers, the pedicels about 8
mm long. Sepals 5, obovate to elliptic-obovate, 2 mm long, thin, obtuse,
imbricate. Petals 5, hyaline, obovate, 1 mm long, broadly rounded at
the apex or slightly retuse, not longer than the disk-glands. Stamens
20; filaments 2 mm long, slender; anthers obovoid, 0.3 mm long. Disk-
glands quadrangular, truncate. Pistillate flowers in subsessile umbels at
the base of the staminate inflorescence. Sepals 5, free, lanceolate, some-
what acuminate, 2 mm long, more or less accrescent. Ovary narrow,
subglabrous, 3-celled, each cell 1-ovuled ; styles three, elongated, cleft, the
style-arms revolute.

PALAWAN, near Puerto Princesa, For. Bur. 4128 Curran, April, 1906. Luzon,
Province of Pangasinan, For. Bur. 8369 Curran & Merritt, December, 1907:
Province of Ilocos Norte, Cape Bojeador, For. Bur. 13829 Merritt & Darling,
November, 1908.

The first representative of the genus to be found in the Philippines, and ap-
parently distinct from all previously described forms.

278 MERRILL. ae

CROTON Linn.

Croton argyratus Blume Bidjdr. (1825) 602; Muell. Arg. in DC. ‘Prod.
15? (1862) 526. He
Minpanao, District of Zamboanga, Port Banga, For. Bur. 9421 Wiis
Hutchinson, February, 1908, in dipterocarp forests at 300 m altitude,
Burma to the Malay Peninsula, Java, Sumatra, and Borneo; new to the Philip
pines. ,

*

ath

Croton ardisioides Hook. f. FJ]. Brit. Ind. 5 (1887) 393. oe
PALAWAN, For. Bur. 3851, 4159 Curran, Bur. Sci. 857 Foxworthy, March and —
May, 1906. a
Malay Peninsula and Borneo; new to the Philippines, ee.

DIMORPHOCALYX Thwaites.

Dimorphocalyx denticulatus sp. nov.

Arbor parva, monoica, glabra; foliis oblongo-lanceolatis vel lanceolatis,
utrinque angustatis, apice tenuiter gradatim acuminatis, usque ad 15 cm
longis, margine glanduloso-denticulatis; inflorescentiis axillaribus, mas-
culinis cymosis, cymis brevibus, congestis, floribus femineis solitarius vel
fasciculatis, petalis sepalis brevioribus. ,

A small glabrous monoecious tree. Branches grayish-brown, terete, 3g
slender. Leaves alternate, firmly chartaceous or subcoriaceous, oblong-
lanceolate to lanceolate, narrowed at both ends, 7 to 15 cm long, 2 to4
em wide, the apex gradually and slenderly acuminate, the base acute, the
margins rather distantly and irregularly glandular-denticulate, grayish-
brown when dry, shining; nerves 10 or 11 on each side of the midrib,
spreading, anastomosing and forming a distinct, submarginal nerve, the Br
secondary nerves rather distinct, the reticulations lax, obscure; petioles —
2 to 3 mm long. Staminate inflorescence axillary, of short, dense, 1 to i
2 cm long cymes, each subtended by two ovate-lanceolate, acuminate, 3.5
mm long bracts. Male flowers short-pedicelled. Calyx-lobes 5, elliptic,
obtuse, imbricate, about 1.5 mm long. Petals elliptic, obtuse, glabrous, ay
3 mm long. Stamens about 15, the outer 5 larger than the others, the
filaments short, the inner ones more or less united. Pistillate flowers on |
the same plant, axillary, solitary or fascicled, stipitate, and subtended by _
several, imbricate, ovate-lanceolate, acute or acuminate bracteoles. Calyx yi
somewhat urceolate, the lobes elliptic, 6 mm long, 3 mm wide, obtuse, in
bud much exceeding the petals. Petals, in bud, imbricate, glabrous, iy
elliptic, 3 mm long, obtuse. Ovary ellipsoid or ovoid, obscurely 3-suleate, —
glabrous, 3-celled, each cell 1-ovuled; styles 3, more or less united below, |
cleft, in bud recurved. :

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9033 Whitford & 7
Hutchinson, December, 1907, in dipterocarp forests at an altitude of about 30 mH /
A species well characterized by its glandular-denticulate leaves.

Se

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 279

EVERETTIODENDRON gen. nov.

Flores dioici, apetai. Discus nullus. Fl. ¢: Sepala 4, valvata
Petala O. Stamina 4, filamentia subnullis; antherae erectae, basifixae,
minutae, subglobosae, loculis longitudinaliter dehiscentibus. Ovarii rudi-
mentum O. Fl. @: Ovarium 3-loculare, ovula in loculis geminata,
angulo centrali loculorum affixa, collateralia, raphe ventrali; styli 3, erecti
patentes vel curvati, incrassati, integri, a basi papilloso-stigmatosi. Fruc-
tus ovoideus vel depresso-globosus, exocarpio suberoso, endocarpio duro,
fere osseo, 3-loculari, loculicide 3-valvato. Arbor ramulis inflorescentiis
ovarioque dense ferrugineo-pubescentibus. Folia alterna, longe petiolata,
penninervia, integra, coriacea. Inflorescentia axillaria, flores 2 numerosi,
paniculati, bracteolati, ¢ solitarii vel (?) racemosi, pauci.

Everettiodendron philippinense sp. nov.

Arbor 15 ad 20 m alta; foliis alternis, coriaceis, oblong-ovatis vel
oblongo-lanceolatis, integris, coriaceis, usque ad 18 cm longis, in sicco
brunneis, nitidis, glabris vel subtus ad nervos parce pubescentibus, nervis
utrinque 5 ad 7, distinctis, ascendentibus, anastomosantibus, reticulis
laxis; petiolo 2 ad 5 cm longo; floribus masculinis in paniculis angustis
axillaribus dispositis, minutis, 4-meris, bracteolatis, congestis, dense ferru-
gineo-pubescentibus; floribus femineis ignotis, ut videtur solitariis vel
breviter racemosis, ex fructibus immaturis ovario dense ferrugineo-pubes-
centi, 3-loculari, loculis 2-ovulatis; fructibus axillaribus, solitariis, ovoi-
deis vel depresso-globosis, circiter 3 cm diametro.

A tree 15 to 20 m high, the young branches and inflorescence densely
ferruginous-pubescent. Branches terete, brown or reddish-brown, glab-
rous. Leaves alternate, coriaceous, oblong-ovate to oblong-lanceolate,
entire, coriaceous, 9 to 18 cm long, 4.5 to 8 em wide, brown and shining
when dry, entire, glabrous or somewhat pubescent on the midrib and
lateral nerves beneath, the apex usually prominently and sharply acumi-
nate, the acumen sometimes subfaleate, the base acute, rarely somewhat
rounded; nerves 5 to 7 on each side of the midrib, prominent beneath,
ascending, slightly curved, anastomosing near the margin, the reticula-
tions lax; petioles often ferruginous-pubescent ultimately subglabrous,
3 to 5 em long, slightly thickened at the apex and sometimes a little
geniculate. Staminate inflorescence in the upper axils, of narrow, 3 to
5 cm long, racemose panicles, all parts of it densely ferruginous-pubescent,
the branches short, each one subtended by an ovate bract about 3 mm
long, the flowers minute, sessile, congested, subtended by bracteoles similar
to but smaller than the bracts. Sepals 4, free, valvate, acute or slightly
acuminate, ovate, pubescent, 2 mm long. Petals none. Stamens 4, in
the center of the flower, with no staminodes and no disk; filaments very
short or subobsolete; anthers globose, basifixed, minute, about 0.2 mm in

280 MERRILL.

diameter, 2-celled, apparently longitudinally dehiscent. Rudimentary
ovary none. Pistillate flowers unknown, but from the fruiting specimens
axillary, solitary or (7) shortly racemose, the sepals from immature
fruits lanceolate, acuminate, densely pubescent, 7 mm long, deciduous.
Ovary ovoid, densely pubescent, 3-celled, each cell with two pendulous
ovules in the upper inner angle; styles 3, simple, free or slightly united
at the base, erect, spreading or incurved, thickened, their backs ferrugi-
nous-pubescent, their inner surfaces papillose-stigmatose from base to
apex. Fruit, when young, ovoid, densely pubescent, when nearly mature
depressed-globose, glabrous or nearly so, 3 cm in diameter, the exocarp
corky, the endocarp hard, almost bone-like in texture, 3-celled, loculicid-
ally 3-valved; seeds (immature) ellipsoid-ovoid, glabrous.

NeGrRos, Himugaan River, For. Bur. 7282 Everett, May 22, 1907, with stami-
nate flowers, in dense forests at 60 m altitude; same locality, For. Bur. 7316
Everett, March, 1907, sterile. Luzon, Province of Pangasinan, Salasa, For. Bur.
9633 Zschokke, December, 1907, in forested stream-depressions, sterile: Province
of Zambales, Bolet River, near Santa Cruz, For. Bur. 8230 Curran & Merritt,
December 4, 1907, with immature fruits, on forested slopes at an altitude of
270 m: Province of Cagayan, Calamaniugan, For. Bur. 11311 Klemme, November
14, 1907, with nearly mature fruits, in forests at 15 m altitude. Local names,
Pangasinan Hbnel; Cagayan Maraculilem.

Mr. Zschokke notes that the tree is cut for lumber; Messrs. Curran & Merritt
that the tree has a very thin brick-red bark which is red inside, and that the
tree is subject to heart-decay, while the native ranger accompanying Mr. Klemme
states that the fruit is used as a condiment in the preparation of food.

The affinities of this new genus are not clear to me, although following
Bentham and Hooker, and Pax in Engler and Prantl, it apparently falls in the
Phyllantheae of the former, and in the Platylobeae-Phyllanthoideae-Brideliae ot
the latter, except in the latter case the petals are wanting, and moreover the
present genus does not resemble any of those placed here by Pax. The sepals
are not in the least imbricate, so far as I can determine, but assuming that
they are slightly so, or that the above form is anomalous in this respect, it
would then fall into the Platylobeae-Phyllanthoideae-Phyllanthineae, and under
this into the Drypetinae, near Putranjiva Wall., and Petalostigma F. Miill.; it
is however very different from both these genera, although its affinity may be
here. There is a possibility that it does not really belong in the Huphorbiaceae,
but I have been unable to place it elsewhere.

The above new genus is dedicated to Mr. H. D. Everett, one of the collectors,
and formerly a forester in the Philippine Forestry Bureau, who lost his life at
the hands of members of the wild tribe inhabiting the interior of southern Negros,
while prosecuting field work there in May, 1908.

GALEARIA Zoll. & Mor.

Galearia filiformis (Blume) Boerl. Hand]. Fl. Nederl. Ind. 3' (1899) 282.
Antidesma filiforme Blume Bijdr. (1826) 1124.

Bennettia filiformis Muell. Arg. in DC. Prodr, 15? (1862) 1038.
Bennettia javanica R. Br. Pl. Jav. Rar. (1852) 249, pl. 50.

MINDANAO, Lake Lanao, Wres. Clemens s.n. May, June, 1906-7.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 281

The Mindanao specimens do not agree perfectly with Galearia filiformis as
deseribed by Robert Brown and Mueller-Arg., but in the absenee of authentic
' material for comparison, I can do no better, at the present time, than refer them
here. The chief point of difference seems to be that the petals of the Philippine
specimens are not strongly cucullate. Hooker f." states that the sections founded
by Mueller-Arg. on the hooded petals are not tenable. The genus is new to the
Philippines.

Java.

GELONIUM Roxb.

Gelonium racemulosum sp. noy.

Arbuscula glabra circiter 1 m alta; foliis oblongis vel lanceolate-
oblongis, pallidis, nitidis, glanduloso-punctatis, usque ad 20 cm longis,
versus apicem grosse irregulariter repando-dentatis, basi acutis vel acu-
minatis; inflorescentiis masculinis oppositifoliis, racemulis binis, pauci-
floris, brevibus, petiolo vix longioribus; sepalis obtusis, plus minus
cucullatis; staminibus circiter 16; fructibus depresso-globosis, circiter
1 cm diametro, plus minus trigonis, loculicide dehiscentibus.

A shrub or undershrub about 1 m high, glabrous throughout. Branches
grayish, slender, the ultimate ones slightly angled. Leaves alternate,
glabrous, pale when dry, glandular-punctate, shining, oblong to. lanceo-
late-oblong, 9 to 20 em long, 2.5 to 4 em wide, chartaceous, the apex
acuminate, the apical portion very coarsely and irregularly repand-
dentate, otherwise entire, the base acute or slightly acuminate; nerves
about 12 on each side of the midrib, anastomosing, the reticulations
distinct; petioles stout, 3 to 4 mm long. Staminate flowers racemose,
leaf-opposed, the racemes in pairs, scarcely as long as the petioles, few-
flowered, flowering from the base to the apex, the lower flowers opening
first and falling, so that there is usually but two or three flowers on a
raceme, the rachis slender, the pedicel-scars prominent. Sepals elliptic
or elliptic-ovate, obtuse, 3 mm long, 2 mm wide, more or less cucullate,
imbricate. Stamens about 16; filaments free, 2 mm long; anthers oblong,
0.8mm long. Disk-glands small. Rudimentary ovary none. Pistillate
flowers unknown, but the calyx, from the fruit, as in the staminate ones.
Fruit depressed-globose, glabrous, about 1 cm in diameter, obtusely
trigonous, 3-celled, loculicidally 3-valved, each cell 1-seeded; styles per-
sistent, free, each shortly cleft, the arms very short, recurved.

Mrnpanao, District of Zamboanga, Port Banga, For. Bur. 9260, 9349 Whitford
& Hutchinson, January, 1908, in dipterocarp forests 20 to 30 m above sea level.

A species well characterized by its leaves being coarsely repand in the apical
portion, and especially by its paired, short racemes, the staminate flowers of most
species of the genus being glomerate.

7El. Brit. Ind..5) (188i jy oar.
85754——-6

282 MURRILL.

HOMALANTHUS A. Juss.

Homalanthus bicolor sp. nov.

Arbuseula glabra, circiter 2 m alta; foliis longissime petiolatis, cordato-
ovatis, breviter acuminatis, basi latis, leviter cordatis, interdum leviter
peltatis, glabris, chartaceis, supra olivaceis, subtus glauco-pallidis, nervis
utringue 11 ad 15, petiolo 8 ad 20 em longo; floribus ignotis; fructibus —
in racemis congestis, obovoideis vel obovoideo-orbicularibus, compressis, —
griseo-brunneis, loculicide 2-valvatis.

A glabrous shrub about 2 m high. Young branches dark-brown,
glabrous. Leaves alternate, very long-petioled, cordate-ovate or subor- es
bicular-ovate, the base broad, cordate, the apex shortly acuminate, entire, *
7 to 13 em long and nearly as wide, chartaceous, glabrous, the upper z
surface olivaceous, the lower very pale, subglaucescent; petiole 8 to 20
cm long, glabrous, with two more or less prominent glands at the apex,
sometimes peltately inserted about 5 mm from the margin of the leaf,
more often marginally inserted; stipules caducous, thin, brown, lanceo- ‘7
late, sharply acuminate, about 7 cm long. Flowers unknown. Fruit *
crowded, racemose, the racemes about 4 cm long; pedicels spreading or
somewhat reflexed, about 8 mm long; fruit obovoid or orbicular-obovoid, 2
compressed, about 8 mm long, grayish-brown, 2-celled compressed at ~
right angles to the dehiscence, loculicidally 2-valved, crowned by the style a
which is nearly as long, and which is cleft nearly to the base,

MinpAnao, Province of Misamis, Mount Malindang, For. Bur. 4722 Mearns ae.
¢& Hutchinson, altitude about 350 m. Moro Topi. “a

A species allied to Homalanthus populneus Pax, and to H. fastwosus F.-Vil.,
and somewhat intermediate between the two, distinguishable from the former by
its much larger leaves and larger fruits, and from the latter by its non- or but

slightly peltate leaves,
MACARANGA Thouars.

Macaranga congestiflora sp. noy. § Pachystemon.

Arbor parva circiter 4 m alta; foliis suborbicularibus, subintegris vel — :
leviter trilobatis, apice breviter acuminatis, basi late peltatis, supra nitidis,
subtus parce hirsutis, glandulosis, radiato 8-nerviis; petiolo 10 ad 15 cm
longo; inflorescentiis axillaribus caulinisque, 1 ad 3 cm longis, femineis
racemosis, masculinis breyiter racemoso-paniculatis, congestis, bracteis
pectinato-laciniatis, ceraceo-glandulosis, pubescentibus; ovario 2—4-loc-
ulare.

A small tree about 4m high. Branches dark-brown, glabrous or more
or less pubescent, the young branchlets slightly brownish-pubescent.
Leaves alternate, long-petiolate, broadly peltate, suborbicular, subentire
or slightly and broadly 3-lobed, the apex slightly acuminate, base rounded,
10 to 20 cm in diameter, subcoriaceous, rather pale when dry, the upper
surface glabrous, shining, the lower somewhat hirsute, especially on the
nerves, and with numerous scattered waxy glands; nerves radiate, the

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 283

basal ones 8, prominent; nerves leaving the midrib above the base about |
6 on each side, the reticulations distinct; petioles 10 to 15 cm long,
terete, pubescent. Inflorescence axillary and from the stems below the
leaves, the staminate fasciculate, dense, of very short racemose panicles
* cm long or less, their branches. 1 cm long or less, gray-pubescent ;
bracteoles cuneate, pubescent, yellow-glandular, about 5 mm long, lacerate-
pectinate; stamens few: pistillate inflorescence axillary and from the
axils below the leaves, racemose, usually solitary 3 cm long or less,
pubescent, each raceme few-flowered ; pedicels stout, pubescent, 2 mm
long; calyx pubescent, cup-shaped, truncate, about 2.5 mm long; ovary
globose, densely yellowish waxy-glandular, 2- to 4-celled ; styles recurved,
more or less pectinate ; bracts ovate or ovate-lanceolate, lacerate-pectinate,
more or less pubescent, 6 to 8 mm long.

PALAWAN, near Puerto Princesa, For. Bur. 3590, 3591 Curran, January, 1906,
in old clearings at about 20 m above sea level, the former with staminate flowers,
the latter with pistillate ones.

This species probably belongs in the section Pachystemon, although 2-, 3-, and
4-celled ovaries were found on the same plant. It is well characterized by its
suborbicular, nearly entire or slightly 3-lobed leaves, and especially by its short,
congested, axillary and cauline inflorescences, the staminate being paniculate, and
the pistillate racemose.

MALLOTUS Lour.

Mallotus korthalsii Muell. Arg. in DC. Prodr. 152 (1862) 976.

Luzon, Province of Cagayan, Aparri, For. Bur. 17072 Curran, February, 1909.
Minpanao, District of Zamboanga, Port Banga, For. Bur. 9017 Whitford &
Hutchinson, November, 1907.

New to the Philippines; previously known only from Borneo and Celebes.

MICRODESMIS Planch.

Microdesmis caseariaefolia Planch. in Hook. Ic. Pl. (1844) sub. t. 758;

Muell. Arg. in DC. Prodr. 15? (1862) 1041.

PALAWAN, near Puerto Princesa, Bur. Sci. 249 Bermejos, December, 1905;
Mount Victoria, For. Bur. 4138 Curran, Bur. Sci. 732 Foxworthy, March, 1906.
BaLaBac, Merrill 5380, October, 1906.

Burma to southern China, the Malay Peninsula, and Borneo. The genus is
new to the Philippines except for a single specimen collected in Palawan by Vidal
and reported without specific name by Ceron;® Vidal’s specimen in the Kew
herbarium, is apparently identical with the material here referred to Planchon’s
species,

OSTODES Blume.

Ostodes serrato-crenata sp. nov.

Arbor subglabra 8 ad 12 m alta; foliis chartaceis vel subcoriaceis, in
sicco brunneis, nitidis, oblongis vel elliptico-oblongis, usque ad 30 cm
longis, basi acutis vel acuminatis, apice breviter acuminatis, margine
gross irregulariter crenato-serratis, dentibus glanduliferis ; nervis utrinque

*Cat. Pl. Herb. (Manila) (1892) 151.

= * 5 OT
PRS a 7.
- aoe
: ae
” < a el
»
I84 MERRILL.

circiter 17, prominentibus ; inflorescentiis masculinis axillaribus vel pse
oterminalibus, elongatis, floribus in glomerulis congestis, plus mim
stellato-ferrugineo-pubescentibus ; staminibus circiter 12.
A subglabrous tree 8 to 12 m high. Branches light-gray or brownish-
gray, glabrous, striate, lenticellate, the young branchlets ferruginous- —
stellate-pubescent. Leaves alternate, oblong or elliptic-oblong, 15 to 30
em long, 6 to 13 em wide, brownish and shining when dry, glabrous or
with very few seattered stellate hairs beneath, the base acute or some-
what acuminate, the apex short-acuminate, margins coarsely and irreg-
ularly crenate-serrate, the teeth bearing glands at their apices; nerves —
about 17 on each side of the midrib, prominent, the primary reticulations |
rather distinct, the ultimate ones faint; petioles 4 to 7 em long, glabrous
or with few, scattered, stellate hairs, frequently shghtly geniculate at
the apex. Staminate inflorescence axillary and pseudoterminal, slender,
elongated, 20 to 40 em long, the branches few, short, the flowers densely
crowded in reduced, scattered cymes, glomerate, the rachis and branches
ultimately glabrous or nearly so, the younger parts more or less stellate-
or furfuraceous-pubescent. Buds more or less trigonous; sepals (in bud)
orbicular-ovate, obtuse, 3 mm long, strongly imbricate, the exposed por-_
tions densely stellate-pubescent. Petals similar but glabrous, and in
bud somewhat smaller. Stamens about 12. Pistillate flowers and fruit
unknown. : %
Luzon, Province of Cagayan, Aparri, For. Bur. 11271, April 1908; San Vicente,
For. Bur. TOTS Klemme, May, 1907. In dense forests 20 to 65 m above sea-level,
known to the Negritos as Tagalipa and Aguindulong.
The genus is new to the Philippines.

STROPHIOBLACHIA bBoerl.

Strophioblachia fimbricalyx Boer]. Mandl. Fl. Nederl. Ind. 3? (1900) 236.
PALAWAN, For, Bur. 4504, 7458 Curran, June, 1905, 1906, Bur. Sei. 241 Ber-
mejos, near the seashore. MINDANAO, district of Zamboanga, Mearns 114, Jan-
uary, 1904.
An interesting addition to the Philippine flora, and especially to the long
list of species known only from the Philippines and Celebes. A monotypic genus
known previously only from Celebes.

ANACARDIACEAS.
PLEIOGYNIUM Engl.

P. solanderi ( Benth.) Kngl. in DC. Monog. Phan. 4 (1883) 255.

Npondias solanderi Benth. Fl. Austral. 1 (1863) 492.
Npondias pleiogyna F. Muell. Fragm. 4:78. '
Luzon, Province of Rizal, For. Bur. 2166, 3163 Ahern’s collector, December, —
1904, and June, 1905, the former, with mature fruits, from Tenay, the latter,
with staminate flowers from Antipolo. “4
A monotypic genus, previously known only from northern Australia, and its

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 285

discovery in Luzon is a striking addition to the list of Australian types in the
Philippine flora. The Philippine form was at first considered by me to be an
undescribed species, but after a careful examination of the description, and

comparison with material from the Port Jackson District, Australia, coll. J. H.

Camfield, December, 1907, I am at loss to discover any character by which the
Philippine plant can be distinguished from the Australian except the unscientific
one of geographical distribution. It is possible that a revision of the genus will
lead to the characterization of more than one species, for Bentham, in the
original description, states that it is quite glabrous in all of its parts, while
Engler describes the young branches and leaves as densely pilose, the Philippine
specimens and the one Australian specimen before me agreeing with the latter.

ACERACE®.
ACER Linn.

Acer curranii sp. nov. § Jntegrifolia.

Arbor glabra usque ad 25 m alta; foliis concoloribus, nitidis, glabris,
integris, coriaceis, reticulatis, ovatis vel elliptico-ovatis, breviter obtuse
acuminatis, basi rotundatis vel obtusis, 3- vel 5-nerviis, usque ad 13
em longis; floribus masculinis corymbosis, corymbis axillaribus, brevibus,
glabris: fructibus 4 cm longis, alis angulo acuto divergentibus, paullo
introssum falcatis.

A glabrous tree 25 m high or less, the trunk reaching a diameter of

(110 em. Branches terete, smooth, glabrous, reddish-brown, with few

scattered lenticels. Leaves ovate to elliptic-ovate, coriaceous or sub-
coriaceous, shining, glabrous, of the same color on both surfaces, distinctly
reticulate, 9 to 13 em long, 5 to 6.5 em wide, entire, the apex shortly
and obtusely acuminate, the base usually broad and rounded, sometimes
blunt, rarely acute; basal nerves one or two pairs, the outer pair, when
present, short, the lateral nerves above the basal ones usually 4 on each
side of the midrib, distant, very prominent, the primary reticulations
very prominent, rather lax, the ultimate ones fine; petioles 2 to 5 em
long. Male inflorescence axillary, corymbose, about 2 cm long, glabrous,
the branches few, short; pedicels 2 to 5 mm long. Sepals 4, free, oblong-
ovate, 2 to 3.5 mm long, about 2 mm wide. Petals 4, similar to the
sepals but narrower. Stamens 4 or 5, inserted in the disk on its outer
side; filaments 2 to 3 mm long; anthers about 1 mm in length. Disk
lobed, fleshy, glabrous, very thick: bracts subtending the inflorescence
numerous, ovate, coriaceous, closely imbricated, 3 mm long, their upper
margins pubescent, deciduous, leaving a thickened, rough base to the
inflorescence 2 to 3 mm long and 2 thick, strongly and densely marked
by the bract-sears. Pistillate flowers unkown. Infrutescence corymbose,
10 cm long or less, glabrous, with few branches, the fruit, including the
wing, 4 em long, the wings diverging at an acute angle, slightly falcate,
10 to 13 mm wide.

286 MERRILL.

Luzon, Province of Bataan, Mount Mariveles, For, Bur. 6254 Curran, Feb- _
ruary, 1907, with nearly mature fruit, in forests at an altitude of about 550 m A
(type): Province of Benguet, Mount Ugo, Bur. Sci. 5708 Ramos, December, 1908,
staminate flowers: Province of Abra, Tue, For. Bur. 14582 Darling, February,
1909, sterile, altitude 1,000 m. Known to the Igorots in Abra as Baleag.

A species manifestly allied to the Asiatic Acer laevigatum Wall., but with
different leaves which are usually broad and rounded at the base, and with 4-
merous staminate flowers. It is quite different from the only other species of the
genus known from the Philippines, A. philippinuwm Merr., that species having the
leaves very glaucous beneath, and the staminate inflorescence racemose. The
latter species is a shrub or small tree, while the present one is a large tree.

A third species is possibly represented by sterile material, For. Bur. 10948 aot
Curran, Mount Data, District of Lepanto, Luzon, the young leaves being densely +3
tomentose beneath. ’ . a

SABIACE/.
MELIOSMA Blume.

Meliosma monophylla sp. nov.

Arbor parva; foliis simplicibus, alternis, oblanceolatis, integris, brun- .
neis, nitidis, usque ad 35 cm longis, 7 cm latis, apice acuminatis, basi <-
sensim decurrento-acuminatis ; nervis utrinque circiter 20, prominentibus,
ascendentibus, anastomosantibus; paniculis terminalibus axillaribusque,
dense ferrugineo-pubescentibus, 15 ad 20 em longis; floribus numerosis,
minutis, ad apices ramulorum congestis.

A small tree, more or less ferruginous-pubescent. Branches terete,
lenticellate, somewhat ferrugineous-pubescent. Leaves simple, oblanceo-
late, 20 to 35 m long, 5 to 7 em wide, subcoriaceous, brownish when dry,
shining, entire or with few irregular and obscure teeth above, the apex
acuminate, the base long and slenderly decurrent-acuminate, glabrous
on the upper surface, except on the somewhat pubescent midrib, beneath
with scattered hairs especially on the midrib and lateral nerves; nerves
about 20, prominent, ascending, anastomosing, the reticulations distinct ; |
petioles pubescent, 2 to 2.5 em long. Panicles terminal and in the upper
axils, 15 to 20 cm long, densely ferruginous-pubescent, the lower branches
spreading. Flowers small, crowded on the ultimate branchlets, sessile,
the bracts more or less pubescent. Outer three petals orbicular, 1.5 to
1.8 mm long, the inner two linear, cleft, about 1 mm long. Stamens 2,
about 1 mm long. Ovary compressed, glabrous, 2-celled, the style short, ‘
simple. ;

Luzon, Province of Rizal, Tanay, Merrill 2339, May, 1903; Antipolo, For. Bur.
}31 Ahern’s collector, February, 1904; Montalban, For. Bur. 3406 Ahern’s col-
lector, November, 1905, distributed as Semecarpus perrottetii March, which they
remotely resemble. ‘T., Malaligas.

The first simple leaved form to be found in the Philippines, manifestly allied

to Meliosma lancifolia Hook. f., of the Malay Peninsula, but apparently sufficiently
distinct.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 287

RHAMNACE.
VENTILAGO Gaertn.

Ventilago dichotoma (Blanco) Merr. in Govt. Lab. Publ. (Philip.) 27
(1905) 32, excl. syn. V. luzontensis Vid.

Enrila dichotoma Blanco Fl. Filip. (1837) 709.

Ventilago monoica Blanco |. ec. ed. 2 (1845) 124; ed. 3, 1:223.

Ventilago maderaspatana F.-Vill. Nov. App. (1880) 48, ex syn. Blanco, non
Gaertn.

Kurrimia gracilis Vid. Rey. Pl. Vase. Filip. (1886) 89.

Ventilago gracilis Merr. & Rolfe in Philip. Journ. Sci. 3 (1908) Bot. 110.

Luzon, Province of Rizal, San Mateo, Vidal 1122 in“Herb. Kew., type of
Kurrimia gracilis Vid.; Bosoboso, For. Bur. 3073 Ahern’s collector, May, 1905,
in flower; Pilea, Bur. Sci. 3303 Ramos, June, 1907, in fruit; Montalban, Loher
4685, 4686, in Herb. Kew. sub Galearia.

Endemic.

The type of Blanco’s new genus and species, Enrila dichotoma, was from San
Mateo, Province of Rizal, Luzon, and all the above specimens are from the same
province, and agree with his description. The genus Hnrila was placed by
Bentham in the Anacardiaceae, as a doubtful one, but Blanco properly localized
it, in the second edition of his Flora de Filipinas, although in reducing Hnrila to
Ventilago, he changed the specific name. It was reduced by F.-Villar to Ven-
tilago maderaspatana Gaertn., which is certainly an error. Having only flowering
specimens, Vidal redescribed the species as Kurrimia gracilis, of the Celastraceae,
but failed to connect Blanco’s species with it, and later in looking over Vidal’s
specimens in the Kew Herbarium, Mr. Rolfe and myself found Vidal’s type to
be a Ventilago, rather than a Kurrimia, and accordingly transferred the species
to the former genus. In making the original transfer of Ventilago dichotoma, -
I cited as a synonym, Ventilago luzoniensis Vid., but this is an error, as an exami-
nation of Vidal’s type shows that this species is quite distinct from the one here
considered, and one to which Blanco’s deseription does not apply.

Ventilago oblongifolia Blume Bijdr. (1826) 1144; Miq. Fl. Ind. Bat. 1%
(1855) 640.

PALAWAN, Puerto Princesa, Bur. Sci. 264 Bermejos, December, 1905. Luzon,
Province of Bulacan, near Norzagaray, Yoder 105, December, 1906.

New to the Philippines; previously reported only from Java. :

Ventilago luzoniensis Vid. Rev. Pl. Vase. Filip. (1886) 90.

V. maderaspatana Vid. Sinopsis Atlas (1883) ¢t. 32, f. D., non Gaertn.

Luzon, Province of Tarlac, La Paz, Vidal 198, in Herb. Kew.

This endemic species somewhat resembles the preceding one, but is distinguish-
able by its much smaller leaves.

Ventilago lucens Miq. FI. Ind. Bat. Suppl. (1860) 330.

Luzon, Province of Tayabas (Principe), Baler, Merrill 1105, August, 1902.

This species has previously not been reported from the Philippines, but so far
as I can determine at present the specimens well represent Miquel’s species,
although I have not seen the type. King?’ states that Ventilago lucens Miq.
must be very near, if not identical with V. leiocarpa Benth., but the specimen
above referred to V. lucens Miq., is quite distinct from material in our herbarium
from Hongkong and from Singapore, supposedly representing Bentham’s species.

Sumatra.

* Journ. As. Soc. Beng. 65? (1896) 380.

288 MERRILL.

ZIZYPHUS Juss.

Zizyphus hutchinsonii sp. nov.

Arbor glabra vel subglabra, inermis; foliis coriaceis vel subcoriaceis,
ovatis vel oblongo-ovatis, acuminatis, minute crenulatis, basi valde ima-
equilateralibus, supra nitidis, subtus plus minus glaucis, trinerviis, reti-
culis tenuibus, obscuris: fructibus globosis, circiter 1.5 cm diametro,
carnosis, glabris, in sicco nigris.

A tree 15 to 25 m high, glabrous or nearly so throughout, spineless.
Branches terete, slender, minutely lenticellate, black or nearly so when
dry, glabrous, the growing parts sometimes slightly pubescent. Leaves
ovate to oblong-ovate, coriaceous or subcoriaceous, 7 to 11 cm long, 3 to
5 em wide, glabrous, the upper surface greenish or brownish when dry,
shining, the lower surface more or less glaucous, the margins minutely
crenulate, the apex slightly acuminate, acumen obtuse or acute, the base
strongly inequilateral, rounded on both sides of the midrib, or rounded
on one side and acute on the other; nerves three, prominent, the nervules
and reticulations very fine, obscure, more prominent on the upper surface
than on the lower; petioles 6 to 8 mm long, sometimes slightly pubescent.
Flowers unknown. Fruits globose, glabrous, black and shining when dry,
about 1.5 em in diameter, the pericarp fleshy, the stone bony, two- or
three-celled ; seeds brown, shining, compressed, 5 mm long.

BaAsILAN, Matangal Point, For. Bur. 3444 Hutchinson, December, 1905, in
forests at about 10 m altitude, said to be abundant locally and known to the
Yacans as Toncud langit. It is also represented by For. Bur. 9265 Whitford &
Hutchinson, from Port Banga, District of Zamboanga, Mindanao, January, 1908.

ELAEOCARPACEA).
ELAEOCARPUS Linn.

Elaeocarpus curranii sp. nov. § Dicera.

Arbor 15 ad 30 m alta, ramulis inflorescentiisque exceptis, glabra;
foliis anguste obovatis vel oblongo-obovatis, usque ad 10 cm longis, apice
obtuse acuminatis apiculatisque, basi cuneatis, margine valde crenato-
serratis; racemis numerosis e axillis defoliatis, folia subaequantibus vel
brevioribus; petalis laceratis, basi agustatis, subglabris: staminibus cir-
citer 12, cellulis antherae aequalibus obtusis; ovario glabro; fructibus
ellipsoideis, circiter 2 cm longis, 1-locellatis.

A tree 15 to 30 m high glabrous or nearly so except the young branch-
lets and inflorescence. Branches terete, gray or brown, lenticellate, the
young branchlets, as well as the petioles and sometimes the young leaves
appressed-pubescent, obovate, rarely oblanceolate, 5 to 10 em long, 1.5
to 4 em wide, firmly chartaceous to subcoriaceous, usually brownish and
somewhat shining when dry, the apex acuminate, acumen usually short,
blunt, apiculate, sometimes minutely retuse, the base narrowed, cuneate,
the margins strongly crenate-serrate, when young sometimes. slightly

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 289

hairy, ultimately glabrous; nerves 6 on each side of the midrib, distinct,
spreading-ascending, shghtly curved, their axils not glandular, the re-
ticulations indistinct; petioles 1 to 2 cm long, when young pubescent,
ultimately glabrous. Racemes numerous, 5 to 7 em long, on the branches
below the leaves, the rachis and pedicels more or less pubescent, in fruit
becoming quite glabrous; pedicels about 5 mm long. Sepals lanceolate,
acuminate, 5 mm long, 1.2 mm wide, with very few scattered hairs on
the back or glabrous, the margins densely pubescent. Petals as long as
the sepals, the upper one-third or one-half lacerate, base narrowed,
glabrous or with few scattered hairs on the back and especially at the
base. Stamens about 12; anthers 1.8 mm long, cells equal, blunt, not
apiculate or ciliate. Ovary ovoid, glabrous, 3-celled; style 2 mm long,
glabrous. Disk pubescent. Fruit ellipsoid about 2 em long, 1-celled,
the endocarp osseous, rugose.

Luzon, Province of Cagayan, Aparri, Mor. Bur. 11314 Klemme, in flower,
April, 1908; Province of Zambales, Subic, For. Bur. 847,918 Maule, April, June,
1904, in fruit: Province of Bataan, Lamao, For. Bur. 7506 Curran, September,
1907, in flower: Province of Tayabas, Lagumanoc, For. Bur. 4 Ware, in flower,
September, 1903, (type): Province of Camarines, Sipocot, For. Bur. 705 Van
Wickle, May, 1904, in fruit. Local names, Cagayan Cumao; Zambales Tagatoy,
Malacadios ; Tayabas Camaysahan; Camarines Maloc-maloc.

A species allied to Hlaeocarpus cumingii Turze. but readily distinguished by its
glabrous ovaries and vegetative characters.

Elaeocarpus luzonicus sp. nov. § JMonocera.

Arbor circiter 15 m alta, glabra, racemis exceptis; foliis subcoriaceis,
oblongo-lanceolatis, usque ad 7 cm longis, nitidis, apice acuminatis, basi
cuneatis, margine crenatis, subtus in axillis glandulosis; racemis axil-
laribus, pubescentibus, folia aequantibus vel longioribus; petalis oblongo-
obovatis, supra simpliciter fimbriatis, extus leviter argenteo-pubescentibus ;
staminibus circiter 24, cellulis antherae inaequalibus, posticis apiculatis,
vix ciliatis; ovarium 2-loculare.

A tree about 15 m high, glabrous throughout except the inflorescence.
Branches terete, brown or grayish, not or but obscurely lenticellate.
Leaves subcoriaceous oblong-lanceolate, or sometimes elliptic-lanceolate,
5 to 7 em long, 1.5 to 3 em wide, shining, beneath sometimes subglaucous,
the apex shortly, or sometimes rather long-acuminate, acumen obtuse,
base cuneate, margins crenate ; nerves about 6 on each side of the midrib,
distinct, the axils beneath mostly prominently glandular; petioles 1 to
1.4 cm long. Racemes axillary, solitary, 6 to 8 em long, the rachis and
pedicels appressed-pubescent with pale, often shining hairs, the pedicels
5 to 6 mm long. Sepals lanceolate, 5 to 6 mm long, 1.5 mm wide, thin,
sparingly appressed-silvery-pubescent outside, the keel within, and the
margins white-villous. Petals thin, oblong-oboyate the upper one-third
simply fimbriate, 6 to 7 mm long, the back appressed-silvery-pubescent,
the margins and lower part of the inner surface white-villous. Stamens
about 24; anthers scabrid, 2 to 2.5 mm long, one cell slightly longer than

6?

290 MERRILL. Fae

the other, apiculate, not ciliate. Ovary pubescent, ovoid, 2-celled; sty!
glabrous, about 5 mm long. Disk pubescent.

Luzon, Province of Camarines, Mount Isarog, For. Bur, 10488 Curran, May
1908, in forests at about 1,000 m altitude. A specimen collected by Rosenbluth, —
For. Bur. 12220, from either Mindoro or Lubang is possibly referable here, but— it -
has more strongly acuminate leaves, and no axillary glands. :

Probably most closely allied to Elaeocarpus multiflorus (Turez.) F.-Vill., but "
quite distinct from that species. 3

Elaeocarpus subglobosus sp. nov. § Ganitrus.

Arbor 10 ad 20 m alta glabra, partibus junioribus inflorescentiisque
exceptis ; foliis oblongis, firmiter chartaceis, usque ad 12 em longis, apice
breviter obtuseque acuminatis, basi acutis, margine leviter crenatis, nervis
utrinque 10 ad 12; racemis axillaribus, pubescentibus, foliis multo
brevioribus; floribus 5-meris, petalis laciniatis; staminibus circiter 25;
ovario 5-loculare; fructibus subglobosis 5- vel 4-locellatis, circiter 12 mm
diametro.

A tree 10 to 20 m high. Branches terete, dark-grayish-brown, gla-
brous, the younger branchlets, petioles and young leaves rather densely
pubescent with appressed, pale, often shining hairs. Leaves oblong,
firmly chartaceous, 7 to 12 cm long, 3 to 4.5 em wide, not or but
slightly shining, glabrous except the very youngest ones, apex shortly
and obtusely acuminate, base acute, margins slightly crenate; nerves, 10
to 12 on each side of the midrib, distinct, forked, obscurely anastomosing,
the axils of the primary nerves and of the larger reticulations glandular ;
petioles 1 to 2 cm long, at first appressed-pubescent, ultimately glabrous.
Racemes axillary, solitary, 2 to 4 em long, appressed-pubescent, the
pedicels about 5 mm. long. Sepals 5, lanceolate, acuminate, 6 mm. long,
1.5 mm wide, pubescent outside, keeled within. Petals as long as the
sepals, lacinate to the middle, slightly hairy on the margins below.
Stamens about 25; anthers linear, scabrid, one cell slightly longer than
the other and ciliate at its apex. Ovary ovoid, pubescent, 5-celled ; style
5 mm long, glabrous. Fruits purple, globose to obscurely globose-ellip-
soid, about 12 mm in diameter, 5-rarely 4-celled, the endocarp bony.

Luzon, Province of Benguet, Baguio, For. Bur. 48317 Alvarez, December, 1908;

Williams 1036, October, 1904.
Allied to Elaeocarpus sphaericus (Gaertn.) K. Sch., but quite distinct.

MALVACEZE.
KOSTELETZKYA Presl.

Kosteletzkya batacensis (Blanco) F.-Vill. Nov. App. (1880) 24.

Hibiscus batacensis Blanco FI. Filip. (1837) 544, ed. 2 (1845) 380, ed. 3,
2:334. ;
Luzon, Province of Ilocos Norte, Bur. Sci. 3315 Mearns, January, 1907: Prov-
ince of Rizal, Pasay, F. de la Llana, February, 1908, Normal School distribu- —
tion 327. $3

The rediscovery of Hibiscus batacensis is considered worthy of record, as the
species has not otherwise been collected since Blanco’s time. F.-Villar «gives a

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. ZO

description in Latin, taken entirely from Blanco’s original description, and states
that he had seen no specimens. An examination of Blanco’s description shows
that it applies unmistakably to Kosteletzkya, and F.-Villar was quite correct in
making the transfer to the latter genus. On studying the above specimens, I
found that they agreed with none of the descriptions of the various species of
Kosteletzkya, other than Blanco’s but thinking that the species might be an
introduced one here, I sent specimens to Kew for comparison, and received the
following communication: “The specimen of Kosteletekya batacensis does not
match any of the species in the Kew Herbarium, from America or elsewhere,
and in view of the wide distribution of the genus, America, Mediterranean region,
Tropical Africa and Madagascar, the existence of an endemic Philippine species
is not theoretically improbable.” Blanco’s type was from the town of Batac, Prov-
ince of Ilocos Norte, Luzon, and the species is probably endemic in the Philip-
pines.

STERCULIACEA.
LEPTONYCHIA Turez.

Leptonychia banahaensis (Elmer) comb. nov.

Grewia banahaensis Elmer Leafl. Philip. Bot. 1 (1908) 319.

Arbuscula glabra vel subglabra, circiter 3 m alta; foliis alternis,
glabris vel subglabris, lanceolatis vel oblongo-lanceolatis, 7 ad 13 cm
longis, nitidis, membranaceis vel submembranaceis, apice caudato-acu-
minatis, basi obtusis vel subrotundatis, minute glanduloso-punctatis,
subtus in axillis glandulosis; nervis utrinque 5, ascendentibus, prominen-
tibus; floribus axillaribus, solitariis, pedicellatis; petalis orbicularibus
vel ovato-orbicularibus, pubescentibus, circiter 2 mm _ longis, liberis;
staminibus 10, 5-adelphis, basi in annulo connatis; staminoideis ex-
terioribus nullis, interioribus 5, brevibus, phalangibus alternantibus;
ovario 3-loculare ; fructibus ovoideis vel obovoideis, glabris, rugosis, 3- vel
abortu 2- vel 1-locularibus, loculicide dehiscentibus, loculis 1-spermis ;
seminibus ovoideis, glabris, leviter compressis, nitidis, circiter 1.3 em
longis, arillatis.

Luzon, Province of Tayabas, Lucban, Hlmer 7569, May, 1906, type number:
Province of Laguna, Santa Maria Mavitac, For. Bur. 10069 Curran, February,
1908; Mabaluebalue Pass, Bur. Sci. 6044 Robinson, March, 1908.

The first representative of the gerfus to be found in the Philippines, and
erroneously described by Mr. Elmer as a Grewia, of the Tilaceae. The species
strongly resembles the Malayan Leptonychia heteroclita (Roxb.) Kurz., but differs
in the total absence of the external row of staminodes.

‘ GUTTIFERZ.
CALOPHYLLUM Linn.

Calophyllum auriculatum sp. nov. § Apetalum.

Arbor circiter 20 m alta, glabra, gemmis ferrugineo-puberulis exceptis,
ramis teretibus, griseis, ramulis crassiusculis, in sicco valde angulatis sul-
catisque; foliis subsessilibus vel breviter petiolatis, coriaceis, nitidis,
oblongo-ellipticis vel oblongo-obovatis, apice obtusis vel rotundatis, rariter
subacutis, basi distincte auriculatis. Inflorescentiis axillaribus, solitariis

a a rag So ea ee ey ee

292 . MERRILL. a r

vel binis, glabris, folia multo brevioribus, floribus paucis, winbellaee
racemosis, longe pedicellatis; fructibus ellipsoideis vel ovoideis, cireit
2 em longis, in sicco rugosis.

A tree about 20 m high, glabrous throughout except the terminal ti s
which are densely ferruginous-puberulent. Branches terete, light-gray,
the branchlets stout, gray or yellowish, strongly angled and suleate. —
Leaves subsessile or very shortly petioled, coriaceous; shining, oblong-
elliptic to oblong-obovate, 14 to 18 em long, 4 to 7 em wide, the apex
obtuse, rounded, or rarely subacute, the base distinctly auricled, the lower
surface paler than the upper, the midrib impressed on the upper surface,
very prominent beneath, the lateral nerves very numerous, more distinct Bs
on the upper surface than on the lower; petioles none, or very stout and :
less than 3 mm long. Inflorescence axillary, solitary or in pairs, rarely
more than 3-flowered, the flowers umbellately disposed at the apex of the
peduncle which is 1 to 1.5 em long or less, the pedicels slender, 1 to
2 em long, elongated in fruit. Buds globose. Sepals 4. Petals none.
Stamens indefinite. Fruit ovoid or ellipsoid, about 2 cm long, rugose
when dry, the pedicels in fruit frequently 4 cm in length.

MINDANAO, District of Zamboanga, Sax River, Williams 2339 (type), February
20, 1905, Copeland 1617, February, 1905: Lake Lanao, Camp Keithley, Mrs.
Clemens 1019, May, 1907. Basttan, Por. Bur. 6126 Hutchinson, July, 1906.

A species much resembling Calophyllum venulosum Zoll. of Java, differing
especially in its shorter petioles, the auriculate bases of the leaves, and the
terminal buds only puberulent, not sericeous.

CRATOXYLON Blume.

ree tals inappendiculate § ANCISTROLOBUS.
. Flowers in axillary, short, few-flowered cymes...............:------c0-r0-- 1. C. chinense ©
. Flowers in terminal, often leafy panicles.
3. Leaves distinetly petioled, lanceolate or rarely oblong-lanceolate, gradually
. narrowed at both ends; panicles ample................---------------- 2. C. celebicum

3. Leaves shortly petioled or subsessile; oval to elliptic, or oblong-ovate,
base usually broad, rounded, often subcordate; panicles usually small.

3. C. blancot

1. Petals with a basal squamule; flowers in few-flowered, axillary cymes. § TRI-
DESMIG = (2. os cdesenteseaceeratek once rte noc oo Sere, oe epee dae Saco asa MUS aces aan 4. C. formosum

to

te

1. Cratoxylon chinense ( Retz.) comb. noy.

Hypericum chinense Retz. Obs. 5 (1789) 27.

Hypericum cochinchinense Lour. Fl. Coechineh. (1790) 472.

Hypericum biflorum Lam. Eneyel. 4 (1797) 170.

Ancistrolobus ligistrinus Spach Suit. Buff. 5 (1836) 358.

Cratorylon polyanthum Worth. Verhandl, Nat. Geschied. Bot. (1839-42) 175,
t. 36; Dyer in Hook. f. Fl. Brit. Ind. 1 (1874) 257; Miq. Fl. Ind. Bat. 1° (1859)
516; Forbes & Hemsl. in Journ. Linn. Soc. Bot. 23 (1886) 74. ae

Cratorylon ligustrinum Blume Mus. Bot. Lugd.-Bat. 2 (1852) 16, f

Cratorylon biflorum Turez. in Bull. Soe. Nat. Mose. 36' (1863) 580.

CuLion, Merrill 454.

Southern China to Burma, the Malay Peninsula and Archipelago.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 293

This species is rather variable, and is apparently rare in the Philippines,
although it has previously been reported from the Archipelago by both Vidal and
F.-Villar. What is apparently the earliest valid name is here adopted, as Britten ”
shows that the part of the volume of Lamack’s work containing the description of
Hypericum biflorum really appeared in the year 1797, and not in 1789. the date
given on the title page.

2. Cratoxylon celebicum Blume Mus. Bot. Lugd.-Bat. 2 (1852) 16; Merr.
in For. Bur. (Philip.) Bull. 1 (1903) 39.

Ancistrolobus floribundus Turez. in Bull. Soc. Nat. Mose. 31! (1858) 382.

Cratoxylon floribundum F.-Vill. Nov. App. (1880) 16; Vid. Phan. Cuming.
Philip. (1885) 95, Rev. Pl. Vase. Filip. (1886) 92; Merr. in Philip. Journ. Sci.
1 (1906) Suppl. 96.

Cratoxylon polyanthum F.-Vill. Nov. App. (1880) 15, non Korth.

Cratoxylon sumatranum Naves in Blanco FI. Filip. ed. 3, pl. 308, non Blume.

Luzon, Province of Cagayan, For. Bur. 5246, 11316, 11312, 12270 Klemme:
Province of Ilocos Norte, Cuming 1221, type number of Ancistrolobus floribundus
Turez.: Province of Benguet, Elmer 6061: Province of Pangasinan, Cuming 965,
Merrill s. n.: Province of Nueva Ecija, For. Bur. 8434 Curran: Province of Zam-
bales, Merrill 2711, 2928, Bur. Sci. 2593 Foxworthy, For. Bur. 6922 Curran,
Por. Bur, 857 Maule: Province of Laguna, Hallier, Elmer: Province of Rizal,
For. Bur. 1157 Ahern’s collector, Merrill 1689, 2823: Province of Bataan, For.
Bur. 6292 Curran, For. Bur. 1601, 2713, 3035 Borden, Whitford 27, Merrill 3152,
Williams 102: Province of Bulacan, For. Bur. 7183 Curran, Yoder 209, For. Bur.
11173 Aguilar: Manila, Ahern 728: Province of Tayabas, For. Bur. 6039 Kobbe.
Merrill 2609, For. Bur. 10744 Curran: Province of Albay, For. Bur. 10568 Curran.
Minporo, For. Bur. 4071, 11390 Merritt. Minpanao, District of Davao, Williams
2790, DeVore & Hoover 165, 169, Copeland 883: Province of Surigao, Ahern 358:
Agusan Valley, For. Bur. 7594 Hutchinson.

A species widely distributed in the Philippines, and somewhat variable, in
some forms approaching Cratowylon blancoi Blume, but usually readily distin-
guished from that species by its much narrower, differently shaped leaves, and
larger panicles. It is well represented by Naves’ plate, cited above. Cratoxrylon
floribundum (Turez.) F.-Vill., appears to be in all respects identical with (.
celebicum Blume, and is accordingly here reduced. The wood is considerably
utilized in the manufacture of charcoal, the common name for charcoal. uling,
appearing in most of the native names. Common names: T., Guyong-guyong;
Cag., Uttw; Zamb., Panagulingon; V. (Surigao), Ulingon; Manobo (Agusan
Valley), Ulingun.

Celebes.

3. Cratoxylon blancoi Blume Mus. Bat. Ludg.-Bat. 2 (1852) 17; Vid. Phan.
Cuming. Philip. (1885) 95, Rev. Pl. Vase. Filip. (1886) 51; Merr. in Philip.
Journ. Sei. 1 (1906) Suppl. 96.

Hypericum olympicum Blanco Fl. Filip. (1837) 613; ed. 2 (1845) 429: ed. 3,
2:416, non Linn.

Ancistrolobus micradenius Turez. in Bull. Soc. Nat. Mose. 31' (1858) 382.

Cratoxylon micradenium F.-Vill. Nov. App. (1880) 16.

Cratoxylon arborescens Vid. Cat. Pl. Prov. Manila (1880) 17, Sinopsis Atlas
(1883) ¢. 10, f. A.; F.-Vill. Nov. App. (1880) 16, non Blume.

Cratoxylon sumatranum F.-Vill. Nov. App. (1880) 16, non Blume.

Cratoxylon hornschuchii Naves in Blanco FI. Filip. ed. 3, pl. 254, non Korth.

" Journ. Bot. 44 (1906) 319.

294 MERRILL.

Luzon, Province of Ilocos Sur, For. Bur. 7107 Klemme: Province of Benguet,
Elmer 6455: Province of Pangasinan, Merrill 2866, small form: Province of Bu- _

lacan, For. Bur. 7165 Curran: Province of Rizal, For. Bur. 5192 Curran: Proy-
ince of Bataan, For. Bur, 1478 Ahern’s collector: Province of Tayabas, Cuming
700: Province of Camarines, Ahern 31, 828, For. Bur. 10422: Province of Albay,
For. Bur. 10582 Curran. Bonor, Cuming 1822, type number of Ancistrolobus
micradenius Turez. MAspBatTr, For. Bur. 991 Clark. Necros, For. Bur. 12339
Everett. MinpdANnao, District of Zamboanga, Ahern 655; Lake Lanao, Mrs. Clem-
ens 464. BAsILan, Hallier s. n.

Like the preceding species widely distributed in the Philippines, and even _

more variable than that, approaching it in some forms. The extreme form can,
however, be readily distinguished by its broader, differently shaped leaves, which
are broad, rounded, and biauriculate at the base, and very short petioled. It
is unquestionably the form described by Blanco as Hypericum olympicum, on
which Blume based his Cratoxylon blancoi, although it is probable that Blanco
included both this, and the more common C. celebicwm in his species. Cratoxylon
blancoi Blume is undoubtedly very closely allied to OC. hornschuchii Blume.
Common names: T., Guyong-guyong, Cansilay; U., Pang., Baringcocoron; B.,
Saling-gogon; V., Oringon, Pagalingan.
Endemic.

Var. apiculatum var. nov.

Differt a typo foliis minoribus, 4 ad 6 cm longis, anguste obovatis, basi
angustatis, apice latis,-abrupte breviter apiculato-acuminatis. —

GUIMARAS, Buena Vista, For. Bur. 31 Gammill, August, 1903, V., Cansilay.

4. Cratoxylon formosum (Jack) Dyer in Hook. f. Fl. Brit. Ind. 1 (1874)
258; Vidal Phan. Cuming. Philip. (1885) 95, Rev. Pl. Vase. Filip. (1886) 52,
Sinopsis Atlas (1883) ¢. 10, f. B; F.-Vill. Nov. App. (1880) 16.

Blodea formosa Jack in Hook. Journ. Bot. 1 (1834) 374.

Tridesmis ochnoides Spach Suit. Buff. 5 (1836) 358.

Hypericum aegyptium Blanco Fl. Filip. (1837) 615; ed. 2 (1845) 430; ed.
3, 2:418, non Linn.

Luzon, Province of Zambales, For. Bur. 6959 Curran, Merrill 2115, 2950:
Province of Bulacan, For. Bur. 11169 Aguilar: Province of Laguna, Bur. Sei.
2385 Foxworthy: Province of Rizal, For. Bur. 481, 2879 Ahern’s collector,
Merrill 1770, 2638: Province of Tayabas, Llmer 9109, For. Bur. 10276 Curran,
Merrill 3997. Cunton, Merrill 584. Guimaras, For. Bur. 283 Gammill. LEYTE,
Cuming 1754. MiInDANAO, District of Zamboanga, For. Bur. 9339 Whitford &
Hutchinson.

Common names: T., Aligogon, Banga, Apang; Zamb., Cayantol; V., Camon-
tayo.

Siam to the Malay Peninsula and Archipelago.

HYPERICUM Linn.

Hypericum loheri sp. noy.

Abuscula glabra circiter 1 m alta; foliis oblongis vel elliptico-oblongis,
chartaceis vel subcoriaceis, circiter 3 cm longis, glanduloso-punctatis,
subtus subglaucis, subsessilibus, apice acutis vel brevissime acuminatis,
nervis tenuibus, utrinque 5 ad 7; floribus axillaribus solitariis, longe
pedicellatis, circiter 2 cm diametro ; stylis coalitis ; ovario 5-loculare.

A glabrous shrub about 2 m high. Branches slender terete, reddish-—

NEW .OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 295

brown rarely somewhat grayish. Leaves opposite, oblong or elliptic-
oblong, 2.5 to 3 em long, 7 to 12 mm wide, firmly chartaceous or sub-
coriaceous, shining, narrowed and acute at both ends or the apex very
shortly apiculate-acuminate, glandular-punctate, beneath slightly glau-
cous; nerves 5 to 7 on each side of the midrib, slender, obscure, the
reticulations obsolete; petioles very short or none. Flowers white, axi-
llary, solitary, about 2 cm in diameter, their pedicels about 1 cm long,
with three or four pairs of short imbricate bracteoles at the base. Sepals
ovate, acute or obscurely acuminate, 1.5 mm long. Petals oblong-obovate,
inequilateral, 10 to 12 mm long, 6 to 7 mm wide, apex more or less
obliquely subtruncate or rounded. Stamens 5-adelphous; filaments 5 to
9 mm long. Ovary and style 11 mm long, the former narrowly oblong-
ovoid, 5-celled, the latter consisting of five entirely united styles. Cap-
sules about 10 mm long, 3 mm in diameter, crowned by the style, septici-
dally 5-valved ; seeds 1 mm long, thick-spindle-shaped.

Luzon, Province of Benguet, Ambuklao, Loher 66 in Herb. Kew.; Mount Ugo,
Bur, Sci. 5716 Ramos, December, 1908: Province of Zambales, Mount Pinatubo,
Bur, Sci. 2562 Foxworthy, April, 1907.

A species apparently most closely allied to Hypericum formosanum Maxim.,
and H. giraldii Keller.

FLACOURTIACEZE.
AHERNIA gen. nov.

Flores hermaphroditi. Sepala 4 vel 5, imbricata, in petala transeuntes.
Petala 10 ad 15, sepalis consimilia sed interiora gradatim angustiora.
Stamina indefinita, epipetala vel perigyna, filamentis filiformibus, elonga-
tis; antherae parvae, abbreviatae, loculis longitudinaliter dehiscentibus.
Ovarium ovoideum vel ellipsoideum, 1-loculare, placentae 5, co-ovulatae ;
stylus simplex; stigmate minuto, obscure 3-lobato vel subdisciforme.
Fructus ovoideus vel ellipsoideus, co-spermus, pericarpio crustaceo,
obscure longitudinaliter sulecato, indehiscente. Semina obovoidea, plus
minus compressa. Arbor subglabra. Folia alterna, chartacea vel sub-
membranacea, acuminata, integra vel supra obscure repando-crenata, basi
5-nervia vel 5-plinervia, 2-glandulosa. Flores mediocres in racemis sim-
plicibus axillaribus dispositi.

Ahernia glandulosa sp. nov.

Arbor subglabra 8 ad 15 m alta; foliis alternis, ovatis vel oblongo-
ovatis, usque ad 15 cm longis, chartaceis vel submembranaceis, acuminatis,
basi obtusis, rotundatis vel subtruncatis, 2-glandulosis, 5-nerviis vel 5-
plinerviis; floribus circiter 1 em longis, in racemis axillaribus solitariis
dispositis, hermaphroditis.

A tree 8 to 15 m high, glabrous except the infloresence and fruits.
Branches terete, brownish-gray, glabrous, obscurely lenticellate. Leaves
alternate, estipulate, ovate or oblong-ovate, chartaceous or submembra-
naceous, 9 to 15 em long, 4 to 8 cm wide, glabrous, shining and nearly

296 MERRILL.

the same color on both surfaces, entire or obscurely repand above, the
apex rather abruptly acuminate, the base usually broad, rounded or sub-
truncate, and with two glands either on the margins at the juncture with
the petiole, or on the petiole itself: basal nerves 5, the outer shorter pair
from the very base of the leaf, the inner and longer more prominent pair
leaving the midrib a short distance above the base and extending to
beyond the middle, the lateral nerves above the basal ones two or three
on each side of the midrib, ascending, prominent, anastomosing, the
reticulations distinct, rather lax; petioles 2.5 to 4.5 em long. Racemes
in the upper axils, solitary, in flower 2.5 to 3.5 em long, few-flowered, m
fruit somewhat longer, densely gray-pubescent, the pedicels 5 to 8 mm
long, longer in fruit. Petals and sepals scarcely distinct, and irregularly
arranged, the sepals + or 5, ovate, acute or slightly acuminate, 10 mm
long, 6 mm wide, densely pubescent outside, strongly imbricate, the outer
two or three entirely free, the inner two or three more or less united
with the petals and usually staminiferous; petals 10 to 15, similar to the
sepals, and like them densely gray-pubescent, of equal length but narrower,
gradually narrower inwards, the innermost ones linear and only 1 to 1.5
mm wide, all more or less imbricate and all connate below. Stamens
indefinite, inserted on the base of the petals and on the inner sepals;
filaments very slender, elongated, very slightly united at the base, 6 to
10 mm long: anthers minute, longitudinally dehiscing, less.than 0.5 mm
long. Ovary free, densely gray-pubescent, slightly stipitate, ovoid or
ellipsoid, narrowed at both ends, 1-celled, with 5 parietal placentae, each
bearing numerous ovules; style simple, terminal, glabrous or slightly
pubescent at the base, 2 to 4 mm long; stigma minute, subdisciform or
obscurely 3-lobed, not larger than the style. Fruit crustaceous, indehis-
cent, ovoid or ellipsoid, 1.5 to 2 cm long, apiculate, densely gray-pubes-
cent, obscurely longitudinally ribbed; seeds many, obovoid, often irreg-
ularly compressed, black, shining, the testa crustaceous, the albumen
fleshy. Rarely flowers are found with an imperfect ovary which is nearly
glabrous, and contains no ovules.

Luzon, Province of Tayabas, Mount Banajao, Mor. Bur. 8039 Curran & Merritt,
November, 1907, in forests at an altitude of about 600 m, in flower: Provinee
of Rizal, Bosoboso, For. Bur. 2005 Ahern’s collector, November, 1904, with nearly
mature fruits: Province of Bataan, Lamao River, Williams 537, January, 1903,
with mature fruits, in forests at an altitude of about 110 m.

This new genus is somewhat anomalous, but seems to be most closely allied
to Oncoba Forsk., differing in its hermaphrodite flowers, its inner perianth lobes
smaller than the outer ones, the stamens slightly united and inserted on the
petals and sepals, its racemose inflorescence, ete. In its perianth characters
it seems also to approach Pyramidocarpus Oliver, of Africa, but is quite different
from that genus in other floral characters. ; f

The above new genus is dedicated to Major George P. Ahern, Director of
Forestry for the Philippine Islands, in commemoration of his active interest in,
and strong support of botanical investigations in the Archipelago during the

past nine years.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 297

HOMALIUM Jacq.

Homalium curranii sp. nov. § Myriantheia.

Arbor cireiter 16 m alta, glabra, inflorescentiis exceptis ; foliis oblongo-
ellipticis, coriaceis, nitidis, usque ad 20 cm longis, acutis, margine leviter
crenatis, nervis utrinque 8; inflorescentiis axillaribus terminalibusque,
paniculatis, folia aequantibus, dense pubescentibus; floribus 4- vel 5-
meris; sepalis anguste oblongis, 5 mm longis, pubescentibus; petalis
sepalis aequilongis, oblongo-spatulatis ; staminibus brevibus, 12 vel 15.

A tree about 16 m high, glabrous except the inflorescence. Branches
brown, terete, lenticellate. Leaves oblong-elliptic, 18 to 20 em long,
7 to 9 em wide, coriaceous, shining, acute at both ends, margins slightly
crenate; nerves 8 on each side of the midrib; rather distinct beneath,
the reticulations rather close; petioles stout, rugose, 1 cm long, Panicles
terminal and axillary, as long as the leaves, densely pale-pubescent, the
branches scattered, spreading, few, the lowest ones often 10 to 12 cm
long, bracts ovate to ovate-lanceolate, pubescent 4 to 5 mm long, the
bracteoles similar but much smaller. Flowers greenish-white, very
shortly pedicelled, racemosely disposed, the buds densely pale-pubescent.
Sepals four or five, narrowly oblong, obtuse, densely pubescent, 5 mm
long, 1.5 mm wide. Petals as long as the sepals, less pubescent, oblongo-
spatulate, obtuse, 1.8 mm wide, base narrowed. Stamens in groups of
threes opposite each petal; filaments glabrous, about 1 mm long. Ovary
villous; styles usually four, glabrous, short.

Luzon, Province of Benguet, near Twin Peaks, For. Bur. 10811 Curran, De-
cember, 1908, on river banks.

A species allied to Homalium luzoniense F.-Vill., and to #. villarianum Vid.,
but quite distinct from both in its floral characters.

HYDNOCARPUS Gaertn.

Hydnocarpus subfalcata sp. nov.

Arbor subglabra 5 ad 15 m alta, dioica; foliis subcoriaceis vel coriaceis,
oblongo-ovatis vel oblongo-lanceolatis rectis vel leviter subfalcatis, glabris,
nitidis, valde caudato-acuminatis, basi acutis vel rariter obtusis, integris,
nervis utrinque 5, obliquis, prominentibus; floribus axillaribus, femineis
solitarilis, pedicellatis, masculinis paniculato-racemosis, sepalis petalis
staminibusque 5; fructibus pyriformibus, usque ad 6 cm longis, dense
ferrugineo-pubescentibus, basi valde angustatis.

A subglabrous dioecious tree 5 to 15 m high. Branches terete, slender,
gray, the younger branchlets often brown, the growing tips more or less
ferruginous-pubescent. Leaves alternate, oblong-ovate to oblong-lan-
ceolate or lanceolate, 8 to 15 cm long, 3 to 6 cm wide, coriaceous or
subcoriaceous, straight or slightly subfaleate, shining, usually brownish
when dry and paler beneath, the margins entire, the apex strongly
caudate-acuminate usually faleate, the acumen blunt, the base acute, rarely

857547

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298 MERRILL.

obtuse; nerves 5 on each side of the midrib, prominent, sharply as
ing, slightly curved, obscurely anastomosing, usually brown when d
contrast to the pale lower surface of the leaf, the reticulations fi
distinct; petioles 5 to 10 mm long, often slightly pubescent. Flower
small, 5-merous, the pistillate ones solitary, axillary, on 3 to 4 mm long ©
pedicels; sepals free, imbricate, ovate, 2 to 4.5 mm long, the inner two -
much larger than the outer ones, pubescent; petals suborbicular, 3 mm _
in diameter, membranaceous, rounded, ciliate ; ovary ovoid, 2 mm in ~
diameter, densely pubescent, 1-celled with three parietal placentae, the
ovules numerous ; stigmas 3, spreading, flattened, 2 mm long. Staminate a
flowers small, in very short paniculate racemes, the inflorescence slightly — 5
pubescent, axillary, solitary, about as long as the petioles; sepals 5, es
free, ovate, acute or acuminate, the two inner ones larger than the outer,
about 3 mm long in bud, outside ferruginous-pubescent; petals 5, ovate
or rounded, membranaceous, ciliate, each with a large orbicular, ciliate — -
scale at the base, about 1 mm in diameter; rudimentary ovary none; —
stamens 5; filaments stout, tapering upwards, 1 mm long, glabrous;
anthers 1 mm long, about 0.6 mm long, the connective rather broad.
Fruit pyriform, indehiscent, the pericarp crustaceous, rather brittle when | é
dry, densely brown- or ferruginous-pubescent, when mature about 65
em long, the base much narrowed; seeds about 8 in each fruit, ellipsoid 4
to narrowly ovoid, often irregularly compressed, 1.5 to 2 cm long. a
Luzon, Province of Zambales, For. Bur. 5906 (type), 5994 Curran, January,
1907, with staminate flowers and fruit, Por. Bur. 379, 917 Maule, March, June,
1904, the former with pistillate flowers, the latter with fruit; Merrill 2934, _
Hallier s. n.: Province of Tayabas (Principe), Baler, Merrill 1006. Common ‘oe
names given in Zambales are Mala usa, Dalinias, Binting dalaga, and Putian; |
in Baler Ngeret.
This species is closely allied to H. venenata Gaertn. of Ceylon, but differs in —
its less numerously nerved and entire leaves, and in its fruit being strongly
narrowed at the base, and pyriform in shape.
Although the Ceylon flora is not especially closely allied to that of the Philip-
pines, and although the Flacourtiaceae is not strongly represented in either region,
this family contains a striking series of species peculiar to the two, including
the above which is manifestly closely allied to Hydnocarpus venenata Gaertn., a
species confined to Ceylon; the genus Osmelia with but four known species, two
closely allied ones confined to the Philippines, one in Celebes, and one in Ceylon;
and the genus J'richadenia, previously a monotypic one and confined to Ceylon,
but of which a second species has now been found in Luzon.

TRICHADENIA Thwaites.

Trichadenia philippinensis sp. noy. F

Arbor dioica, subglabra, 15 ad 20 m alta; foliis longe petiolatis, coria-
ceis vel subcoriaceis, elliptico-oblongis vel oblongis, nitidis, glabris, 14 —
ad 30 cm longis, integris vel supra obscure repandis, acuminatis, basi
rotundatis; nervis utrinque 10 ad 12, prominentibus; racemis axilla-
ribus, solitariis, usque ad 10 em longis, ferrugineo-puberulis, calycibus:
irregulariter 2- vel 3-lobatis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. Zza9

A dioecious subglabrous tree 15 to 20 m high. Branches stout, brown
or grayish, glabrous, terete, with very large leaf-scars, the ultimate
branchlets densely ferruginous-pubescent ; stipules linear-lanceolate, pubes-
cent, decidous, about 6 mm long. Leaves alternate, coriaceous or sub-
coriaceous, shining, elliptic-oblong or oblong, 14 to 30 em long, 6 to
15 em wide, glabrous, entire, or the margins in the upper portion slightly
and obscurely repand, the apex rather abruptly and sharply acuminate,
the base broad, rounded; nerves 10 to 12 on each side of the midrib,
prominent, anastomosing, the reticulations distinct, rather lax; petioles
3 to 8 cm long, when young ferruginous-puberulent, ultimately glabrous.
Flowers in axillary, solitary, ferruginous-puberulent racemes which are
10 cm long or less: pistillate flowers pedicillate, the calyx glabrous,
splitting from the top into two or three irregular lobes; petals 5, free,
ovate to elliptic-ovate, obtuse, about 4 mm long, 2.5 to 3 mm wide,
glabrous except the oblong-lanceolate, densely pubescent scale on the
inside, which is nearly 3 mm long; ovary ovoid, pubescent, 1-celled, with
three parietal placentae, each with one ovule; styles three, short ; stigmas
three, thick, large, subpeltate, irregularly lobed: staminate flowers similar
to the pistillate, the irregular calyx divisions reflexed in anthesis; stamens
5; filaments 2.6 mm long; anthers 2 mm long, 2-locellate. Fruit
(immature) subglobose, glabrous, 2 cm in diameter, 1-seeded, the pericarp
crustaceous when dry. s

Luzon, Province of Rizal, Tanay, Merrill 2280 pistillate flowers; Bosoboso,
Por. Bur. 2982 Ahern’s collector, staminate flowers, Bur. Sci. 2649 Ramos,
staminate flowers: Province of Laguna, Santa Maria Mavitac, For. Bur. 10065
Curran, February, 1908, sterile: Province of Tayabas, Lagumanoec, For. Bur. 9
Ware, immature fruit. Minpanao, Lake Lanao, Camp Keithley, Mrs. Clemens
695, with detached fruit.

A species closely allied to Trichadenia zeylanica Thwaites, the type of the
genus, but is readily distinguished by its more numerously nerved, entire or
nearly entire, glabrous leaves, as well as by its smaller flowers, the calyx splitting
irregularly from the top. A most interesting discovery, the genus previously
consisting of the single species 7’. zeylanica Thwaites, confined to Ceylon. Local
Tagalog names are given by Ramos as Tadong or Tandong, and by Ware as
Malapingan.

LECYTHIDACE2.
BARRINGTONIA Forst.

Barringtonia balabacensis sp. nov. § Stravidium.

Arbor circiter 10 m alta, glabra; foliis chartaceis, oblongis, nitidis,
*5 ad 40 cm longis, basi acuminatis, apice breviter abrupteque acumi-
natis, margine minute crenato-serratis, nervis utrinque 16 ad 18, pro-
minentibus ; spicis axillaribus, usque ad 40 cm longis; fructibus anguste
oblongis, circiter 10 cm longis, 2.5 cm diametro, plus minus quadran-
gulatis.

A tree about 10 m high, glabrous throughout. Leaves oblong, charta-
ceous, shining, 25 to 40 cm long, 8 to 12 em wide, the base somewhat

e + sy 88 - a

. af “ +) a
300 MERRILL. oe
mt

acuminate, the apex sharply and abruptly crureinale the acumen ab
1 cm long, the margins rather finely crenate-serrate or in the lower part a
subentire; petioles 3 to 6 cm long; nerves 16 to 18 on each side of the |
midrib, prominent beneath, anastomosing, the reticulations distinct. %
Spikes axillary, many-flowered, in anthesis 10 to 15 cm long, in fruit
up to 40 cm in length and somewhat thickened. Flower-buds sessile,
each subtended by three bracteoles, a lanceolate, acuminate middle one
about 5 mm long, and two lateral, much smaller ones. Calyx-tube 5
mim long, somewhat quadrangular, the lobes three, one of which is some-
times split at the apex, ovate, obtuse, about 6 mm long. Petals 4,
elliptic, 1.8 cm long. Mature flowers not seen. Fruit narrowly oblong,
about 10 em long, 2.5 em thick, somewhat quadrangular, the angles
rounded, the style persistent im young fruits, slender, 5 cm long.
Bavapac, Bur. Sei. 422 Mangubat, March, 1906.

A species said by the collector to be rare, growing in forests, and used by
the natives to poison fish. N. v. (Moro), Ulam.

COMBRETACE.
COMBRETUM Linn.

Combretum extensum Roxb. Hort. Beng. (1814) 28; Fl. Ind. 2 (1824) 229;
King in Journ. As. Soc. beng. 66* (1897) 337.
PALAWAN, Mount Pulgar, Bur. Sci. 547 Foxworthy, March, 1906. M1Nporo,
Madrugo River, For. Bur. 4074 Merritt, April, 1906.
British India to the Malay Peninsula, Andaman Islands, and Java; new to the

Philippines.
TERMINALIA Linn.

Terminalia comintana (Blanco) comb. nov.
Bucida comintana Blanco FI. Filip. (1837) 856, ed. 2 (1845) 265, ed. 3, 2:48.
Terminalia chebula F.-Vill. Nov. App. (1880) 80, non Retz.

Terminalia multiflora Merr. in Govt. Lab. Publ. (Philip.) 17 (1904) 34.

Terminalia polyantha Pres] Epim. Bot. (1851) 213?

Luzon, Province of Zambales, Canad, For. Bur. 5897 Curran: Province of
Pangasinan, Cape Bolinao, For. Bur. 8381 Curran & Merritt: Provinee of Rizal,
Bosoboso, Merrill 2796 (type of Terminalia multiflora Merr.), 2647, For. Bur. -
2984, 2009 Ahern’s collector, For. Bur. 10085 Curran, Bur. Sci. 3265 Ramos:
Province of Batangas, San Jose, Guerrero s. n. (topotype of Bucida comintana
Blanco): Province of Bataan, Mount Mariveles, Elmer 6990, Whitford 1257:
Province of Tayabas (Infanta), Tinuan River, Whitford 767: Province of Cama-
rines, Mount Isarog, For. Bur. 10448 Curran. Minpvoro, For. Bur. 6193, 8708
Merritt, For. Bur, 12211 Rosenbluth. Trcao, For, Bur. 1030 Clark. MtnpdANAo,
District of Zamboanga, For. Bur. 9063 Whitford & Hutchinson. Local names;
Zambales Bingas, Batitinan; Pangasinan Magtalopoi; Rizal Naghubo, Palauag,
Saplungan; Batangas Dinglas; Tayabas (Infanta) Bangias; Camarines Tiroon; *
Mindoro Bangias; Tieao Batitinan-babaye; Zamboanga Malatagum, Batitinan.

Blanco’s Bucida comintana has previously been considered as a doubtful species, |
and I have expressed the opinion” that it was referable to the genus Calycopteris,
and that F.-Villar was wrong in transferring it to the genus Terminalia. While —

* Govt. Pab. Publ. (Philip.) 17 (1904) 34.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 301

F.-Villar was certainly in error in transferring Blanco’s species to Terminalia
chebula Retz., | am now of the opinion that he was correct as to the genus, and
that Bucida comintana Blanco is identical with the species previously described
by me as Terminalia multiflora. Through the kindness of Dr. Leon Guerrero,
member of the Philippine Assembly, we have been able to secure specimens of the
tree locally known as Dinglas in the town of San Jose, Province of Batangas,
which native name was cited by Blanco in his original description and from
which town Blanco secured his specimens. A similar name occurs on_ speci-
mens from Zambales, Bingas, while Bingias is applied to the same species in the
Province of Tayabas, and in Mindoro. Blanco did not have mature fruits when
he described the species, and apparently assumed that the ealyx was persistent
in ripe fruits; the calyx-rim is, however, very early deciduous. The specific
name is from an old name of the Province of Batangas, according to Blanco.

Terminalia polyantha Presl is probably not specifically distinct from the above
species, but there are slight differences in the leaves. I previously reduced Presl’s
species to Terminalia catappa Linn., from the abridged description given by
Miquel, to which species it is not at all allied; a specimen of Cuming 1516, on
which the species was based, is now in our herbarium.

A specimen from Celebes in the Kew Herbarium, collected by Beceari, is
probably referable to Terminalia comintana (Blanco) Merr., which species is
otherwise not known from outside of the Philippines.

Terminalia quadrialata sp. noy.

Arbor glabra, 15 ad 35 m alta; foliis oblongo-obovatis vel elliptico-

- obovatis, subcoriaceis, nitidis, usque ad 20 cm longis, apice breviter acu-

minatis, basi sensim angustatis, decurrento-alatis, nervis utrinque 15 ad
20, distinctis, reticulis obscuris; paniculis terminalibus, ramis patulis;
fructibus 2 ad 3 cm longis, apice retusis, regulariter 4-alatis, alis tenuiter
coriaceis vel submembranaceis, 1 ad 1.3 cm latis.

A tree, glabrous throughout, 15 to 35 m high. Branches rather stout,
grayish, or reddish-brown, lenticellate. Leaves somewhat crowded to-
wards the apices of the branches, oblong-obovate to elliptic-obovate, 15
to 25 cm long, 4 to 10 cm wide, subcoriaceous, shining, glabrous, of about
the same color on both surfaces when dry, the apex shortly acuminate,
rarely subobtuse, gradually narrowed towards the base which is somewhat
decurrent; nerves 15 to 20 on each side of the midrib, distinct, parallel,
anastomosing, the reticulations lax, obscure; petioles stout, usually 5
mm long or less, but the lamina sometimes decurrent to the branch, so
that the leaves often appear to be sessile or subsessile. Flowers unknown.
Fruiting panicles terminal, 20 cm long or less, the branches spreading,
sometimes reflexed, the lower ones sometimes 8 cm long, with few:
secondary branches, or unbranched, the upper ones gradually shorter,
scattered, more or less thickened towards their apices, and in the apical
portions bearing many pedicel-scars. Fruit, including the wings, ellip-
soid or suborbicular in outline, 2 to 3 cm long, nearly as wide, retuse at
the apex, rounded at the base, the seed-bearing portion very narrow,
thin-walled, usually but 5 mm thick; wings four, equal, thinly coriaceous
or submembranaceous, 1 to 1.3 cm wide, transversely nerved; seeils
oblong, 6 to 7 mm long.

B02 MERRILL.

Luzon, Province of Sorsogon, Sua, Mor. Bur. 4526 Zschokke. MASBATE, For.
Bur. 12562, 12558 (type), 12583, 12597, 12814 Rosenbluth. Samar, For. Bur.
12617, 12874 Rosenbluth. In forests up to 100 m altitude.

A species well characterized by its four-winged fruits, the wings all the same
width, very thin, and the seed-bearing portion very narrow. All the specimens
cited bear the native name T'oog.

MELASTOMATACE/®.
MEDINILLA Gaudich.

Medinilla curranii sp. nov.

Frutex scandens, glabra ; ramis ramulisque teretibus ; foliis verticillatis,
quaternis, elliptico-ovatis, acutis vel breviter acuminatis, circiter 14 cm
longis, chartaceis vel subcoriaceis, 5-plinerviis,. petiolatis; cymis axilla-
ribus, brevibus, paucifloris; floribus 4-meris, circiter 3 cm longis.

A seandent shrub, glabrous throughout. Branches and_ branchlets
terete, gray. Leaves verticillate, quaternate, elliptic-ovate, about 14 cm
long, 6 to 7 em wide, chartaceous, or subcoriaceous, shining, the base
acute, the apex acute or very shortly acuminate; nerves 5, the middle
one prominent, the lowest pair leaving. the midrib near the base, the
upper pair at 2 or 3 cm above the base, reticulations obsolete; petioles
about 1.5 em long. Cymes from the branches, in the axils of fallen
leaves, usually solitary, few-flowered, the peduncles about 1 cm long.
Flowers white and pink, nearly 3 cm long. Calyx cup-shaped, about
11 cm long, truncate. Petals 4, about 20 mm long, 9 mm wide above,
narrowly obovate, narrowed below. Stamens 8, unequal; filaments about
7 mm long; four anthers about 18 mm long, and four 10 to 12 mm
long, the anterior lobes two, about 1.5 mm long, the posterior one
club-shaped, 2 to 3 mm long, the short anthers relatively stouter than
the long ones. Style 11 mm long.

Luzon, Province of Camarines, Caramoan, For. Bur. 12289 Curran, June 27,
1908.

A species allied to Medinilla verticillata Merr., differing in its much larger
and 4-merous flowers.

Medinilla mindorensis sp. noy.

Frutex scandens, glabra; ramis ramulisque tenuibus, teretibus; foliis
oppositis, petiolatis, ovatis, ellipticis, vel obovatis, acuminatis, subcoriaceis,
7 ad 11 em longis, 3- vel 5-plinerviis, reticulis obsoletis; paniculis
terminalibus, 15 ad 20 em longis, diffusis, pedunculatis ; floribus 4-meris ;
bracteis membranaceis, reticulatis, obovatis, 8 ad 10 mm longis, per-
sistentibus.

A scandent shrub glabrous throughout. Branches and_ branchlets
slender, terete, light-gray. Leaves opposite, ovate to elliptic, rarely
obovate, 7 to 11 em long, 3.5 to 6 em wide, subcoriaceous, shining, the
base acute or acuminate, the apex rather strongly acuminate ; nerves three,
sometimes five, the additional pair, if present, faint, the lateral ones

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 308

leaving the midrib slightly above the base of the leaf, reaching the apex,
or evanescent, the reticulations obsolete; petioles 0.5 to 2 ecm long.
Panicles terminal, 15 to 20 cm long, rather diffuse, the peduncles 7 to
10 cm long, the lower branches 5 to 6 em long, spreading, opposite,
few-flowered. Pedicels slender, 3 to 10 mm long, the bracts persistent,
obovate, about 1 cm long. Flowers 4-merous. Calyx urceolate, 3 mm
long, truncate. Petals 4, broadly obovate, 4 mm long. Stamens 8,
their anthers about 4 mm long, the filaments of four about 4 mm long,
and of the other four 3 mm long, the spur and appendages small.
Bracteoles persistent, obovate, reticulate, 8 to 10 mm long, white or
pink, two for each flower.

Minporo, Ibalo River, For. Bur. 11489 Merritt, May, 1908; Mount Halcon,
For. Bur. 4868 Merritt, June, 1906.

Well characterized by its diffuse panicles, rather small flowers and prominent,
persistent, reticulate bracts and bracteoles.

Medinilla ovalis sp. noy.

Frutex erecta vel scandens, subglabra; ramis ramulisque teretibus,
minute stellato-lepidotis; foliis oppositis, petiolatis, ovalibus, 5 ad 8 em
longis, subcoriaceis, apice breviter abrupteque acuminatis, basi obscure
5- vel 7-plinerviis; inflorescentiis terminalibus, paniculatis, pedunculatis,
10 ad 12 em longis, minute stellato-lepidotis; floribus +-meris, circiter
1 cm longis.

An erect or scandent shrub, the younger branchlets, petioles, midrib of
the leaves on the lower surface, and panicles minutely brown-stellate-
lepidote, the indumentum scarcely visible without a lens. Branches
grayish-brown, terete, nearly glabrous, the branchlets sometimes slightly
angled; nodes smooth. Leaves opposite, oval, subcoriaceous, 5 to 8 em
long, 3.5 to 6 cm wide, the base broad, rounded or subtruncate, the apex
shortly and abruptly acuminate, shining on both surfaces, the lower one
paler than the upper, the midrib prominent beneath, minutely lepidote,
the surface with scattered, minute, obscure glands; nerves from near the
base 5 or 7, obscure, except the midrib, reticulations obsolete; petioles 1
to 2 em long. Panicles terminal, 10 to 12 cm long, branched above the
middle, the branches verticillate, the lower ones about 3 cm long,
the bracts oblong, about 6 mm long, the bracteoles minute. Flowers
purplish, about 1 em long, the pedicels 4 mm long or less. Calyx some-
what urceolate, truncate, 4 mm long. Petals 4,5 mm long. Stamens 8,
subequal, the filaments 4 mm long; anthers lanceolate, acuminate, 5
mm long.

Luzon, Province of Albay or Sorsogon, Adlumoy Hills, For. Bur. 12387 Curran,
June 17, 1908.

A very characteristic species, readily distinguishable by its opposite, oval,

obscurely nerved leaves, peduncled terminal panicles, and minute lepidote-stellate
indumentum.

A
a
304 MERRILL. 2 oR
: ; ‘bY “
MEMECYLON Linn. a

‘
Memecylon oligoneuron Blume Mus. Bot. 1 (1851) 353; Mig. Fl. Ind. Bat.
1* (1855) 574; Cogn. in eo Monog. Phan. 7 (1891) 1132; King. in Journ. As.
Soe. Beng. 69° (1900) 7
Rhodamnia glabra ihe Rev. Pl. Vase. Filip. (1886) 129; Ceron Cat. PL fea
(1892) 79. &
Luzon, Province of Tayabas, Guinayangan, Vidal 782, in Herb. Kew (type of | p
Rhodamnia glabra Vid.) : Province of Rizal, Bur. Sci. 3314 Ramos, June, 1907: 5
Province of Laguna, For. Bur. 10111 Curran, February, 1908. .
Rhodamnia glabra Vid., was deseribed from fruiting specimens, and in the
absence of flowers was placed by Vidal in the Myrtaceae. Additional material
shows it to be Memecylon, and identical with M. oligoneuron Blume. The genus —
Rhodamnia must therefore be excluded from the Philippines.
Perak, Penang, Java and Borneo; not previously reported from the Philippines
under its correct name. :

ERICACEZ.
VACCINIUM Linn.

Vaccinium alvarezii sp. noy.

Abuscula glabra circiter 3 m alta; foliis Baeaae: obovatis vel elliptico-
oblongis, coriaceis nitidis, integris, usque ad 10 em longis, breviter obtuse
acuminatis, nervis utrinque 5 vel 6, ascendentibus, tenuibus; racemis —
axillaribus,’ solitariis vel binis, foliis bevioribus; corolla cylindracea, 1
leviter inflata, circiter 1 cm longa; staminibus 10, antheris scaberulis, |
dorso vix aristatis, apice breviter productis appendicibus infundibuli-
formibus, divaricatis, poris apicaliter dehiscentibus.

A glabrous shrub about 3 m high. Branches reddish-brown, some-
what mottled with gray, terete. Leaves oblong-ovate to elliptic-oblong,
6 to 10 cm long, 2.5 to 5.5 em wide, entire, the apex shortly and obtusely
acuminate, the base acute, margins slightly recurved, shining on both —
surfaces ; nerves 5 or 6 on each side of the midrib, slender, not prominent,
ascending, anastomosing, the reticulations lax; petioles 4 to 6 mm long.
Racemes axillary, solitary or in pairs, 3 to 5 em long, the pedicels about
1.5 cm long, articulated with the calyx. Calyx-tube short, the lobes
broadly ovate, obtuse, 1.5 to 2 mm long, each lobe gland-tipped. Corolla
pink, cylindric, 10 to 11 mm long, about 5 mm. in diameter, slightly
inflated in the midle, the lobes erect, ovate, obtuse, 1.5 to 2 mm long.
Stamens 10; filaments 4 to 5 mm long, white-villous; anthers 3 mm long,
scaberulous, not awned, the apex produced into two short, broad, funnel-
shaped divergent tubes, opening by terminal, orbicular pores. Disk ir
prominent erenate-undulate ; style 1 em long.

Luzon, Province of Cagayan, Dalisay River, For. Bur. 18466 Alvarez, March,
1909, in forests, altitude about 650 m.

Probably most closely allied to Vaccinium barandanum Vid, and V. benguetense —
Vid., but quite distinet from both. .

| ie
’

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 305

KBENACE®.

DIOSPYROS Linn.

Diospyros ahernii sp. noy.

Arbor glabra inflorescentis fructibusque exceptis; ramis pallidis,
eriseis vel brunneis; foliis subcoriaceis, oblongis, 16 ad 20 em longis,
apice acuminatis, basi acutis vel leviter acuminatis, in sicco brunneis,
utrinque nitidis, nervis utrinque circiter 8, prominentibus, anastomosan-
tibus, nervis reticulisque densis, validis; inflorescentiis femineis axilla-
ribus, racemosis, pubescentibus ; fructibus ut videtur globosis, circiter 5 em
diametro, obtusis vel apiculatis, extus densissime ferrugineo-velutinis vel
pubescentibus, calycis lobis 4, accrescentibus, plus minus connatis, dense
ferrugineo-pubescentibus, patulis vel reflexis, vix imbricatis, acuminatis,
1 ad 1.5 cm longis.

A tree, glabrous except the infloresence. Branches usually pale, lght-
gray or brownish, glabrous, the ultimate branchlets sometimes somewhat
ferruginous-pubescent. Leaves subcoriaceous, oblong, 16 to 20 em long,
4.5 to 7 em wide, glabrous and shining on both surfaces, usually brown
when dry, the apex distinctly acuminate, base acute or somewhat acu-
minate; nerves about 8 on each side of the midrib, distinct, anastomosing,
the secondary nerves and reticulations distinct, close. Flowers unknown,
but the pistillate inflorescence axillary, racemose, densely pubescent, the
racemes in fruit 1.5 to 3 cm long, usually bearing but one or two fruits,
the pedicels stout, about 5 mm long. Fruit apparently globose, about
5 cm in diameter, rounded or apiculate, outside densely ferruginous-
velvety when young, pubescent when old, about 5-celled. Calyx per-
sistent, accrescent, spreading in young fruits, reflexed in mature ones,
3 to 4 cm in diameter, 4-lobed, the lobes united for the lower third, ovate,
coriaceous, acuminate, 1 to 1.5 cm long, densely ferruginous-pubescent.

Luzon, Province of Rizal, Bosoboso, For. Bur. 3071 Ahern’s collector, May,,
1905; Pilea, Bur. Sct. 3298 Ramos, June, 1907, both with immature fruits:
Province of Tayabas, Lucban, Elmer 9110, May, 1907, distributed as Diospyros
discolor Willd.

A very distinct species, possibly allied to D. discolor Willd., but at once distin-
guished by its glabrous leaves and its connate acuminate calyx-lobes which are
not at all imbricate but spreading or reflexed, while its general appearance is
quite different from Wildenow’s species. The specimen collected by Elmer prob-
ably had nearly mature fruits, but the sheet before me has only some fragments

of the fruit, so that it is impossible to give a full description at the present
time. It is known in Rizal Province as Talong-gubat.

Diospyros buxifolia (Blume) Hiern Monog. Eben. (1873) 218.

Leucoxylum busxifolium Blame Bijdr. (1826) 1169; Choisy Mém. Ternstr.
(1855) 43, t. 2; Miq. Fl. Ind. Bat. 2 (1858) 1050.

Diospyros microphylla Bedd. Ie. Pl. Ind. Or. (1871) 27, t. 133; Clarke in Hook.
f. Fl. Brit. Ind. 3 (1882) 559; King & Gamble in Journ. As. Soc. Beng. 74°
(1905) 210.

306 .~ MERRILL.

MinpANAO, District of Zamboanga, Port Banga, For. Bur. 9435 Whitford &
Hutchinson, February, 1908.

The specimen is sterile but agrees closely with the description, with material |

from Java, and fairly well with specimens from Singapore. New to the Philip-
pines.

Southern India to the Malay Peninsula, Andaman Islands, Sumatra, Java,
and Borneo.

Diospyros curranii nom. noy. § Paralea.

Diospyros reticulata Elm. Leafl. Philip. Bot. 2 (1908) 506, non Willd. nec.
Sieber, nec Wall.

Arbor glabra vel subglabra, 7 ad 20 m alta; ramis teretibus, griseis
vel brunneis, glabris, vel junioribus plus minus pubescentibus; foliis
glabris, lanceolatis, elliptico-lanceolatis vel oblongo-lanceolatis, 10 ad 18
cm longis, 2.5 ad 5 cm latis, basi acutis vel acuminatis, apice acuminatis,
subcoriaceis, nervis utrinque 9 ad 15, anastomosantibus, nervulis reticulis-
que prominentibus; floribus masculinis 4- rariter 5-meris, in cymulis
brevibus axillaribus dispositis, staminibus circiter 18; floribus femineis
axillaribus, solitariis vel fasciculatis, 4- meris, pubescentibus, calycis lobis
ovatis, reflexis, accrescentibus ; ovario 3- vel 4-loculare ; fructibus ovoideis,
1.5 cm longis.

A tree 7 to 20 m high, glabrous or nearly so. Branches glabrous,
terete, brownish or grayish, the branchlets sometimes somewhat ferrugi-
nous-pubescent. Leaves lanceolate to elliptic-lanceolate or oblong-lance-
olate, 10 to 18 cm long, 2.5 to 5 cm wide, chartaceous to subcoriaceous,
shining, pale, greenish or yellowish when dry, entire, the base acute or
somewhat acuminate, eglandular, the apex acuminate; nerves 9 to 15
on each side of the midrib, anastomosing, rather distinct but scarcely
more prominent than are the secondary nerves and rather dense reticula-
tions; petioles 1.5 cm long or less, sometimes as short as 0.5 em. Stami-
nate flowers in short, axillary, solitary or fascicled, 1 to 2 em long cymes,
which are slightly pubescent. Calyx lobes 4, rarely 5, ovate to broadly
ovate, acute or acuminate, about 1.5 mm long, with short, appressed,
scattered, black hairs; stamens about 18, 2-seriate, inserted on the base
of the corolla; filaments 1 to 1.5 mm long; anthers lanceolate, acuminate,
glabrous, nearly 2 mm long; corolla (immature) 5 mm long, the outside
with black appressed hairs. Pistillate flowers axillary, solitary or fasci-
cled, shortly pedicellate, the pedicels and calyx somewhat ferruginous-
pubescent: calyx-lobes 4, nearly free, ovate, about 7 mm long, 5 mm wide,

obtuse, subfoliaceous, the margins reflexed, the base subcordate, somewhat -

accrescent and persistent in fruit; corolla about 6 m long, the lobes ovate,
3 mm wide, slightly united below; staminodes about 10; ovary narrowly
ovoid, ferruginous-pubescent; style short, 2-cleft. Fruit ovoid, 1.5 em
long, when young more or less ferruginous-pubescent, ultimately glabrous,
the calyx-lobes somewhat accrescent, their margins reflexed.

Luzon, Province of Rizal, Bosoboso, For. Bur. 100384 Ourran, February, 1908;
Antipolo, Merrill 1654, 2675 March, June, 1903, Por. Bur. 448 Ahern’s collector,

4
;
‘
&

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 307

April, 1904, Bur. Sci. 2192 Ramos, March, 1907: Province of Tayabas, Guina-
yangan, Merrill 2029; Lueban, Elmer 9241 (type); Atimonan, For. Bur. 6623
Reyes, For. Bur. 10294, 10662 Curran: Province of Camarines, Mount Isarog,
For. Bur. 10452 Curran: Province of Sorsogon, For. Bur. 5165 Bridges. MAaARIn-
DUQUE, For. Bur. 12182 Rosenbluth. Minpanao, Lake Lanao, Camp Keithley,
Mrs. Clemens, June, 1906: District of Davao, Santa Cruz, Williams 2861.

In the above description the chief additions to that given by Mr. Elmer are
of the staminate and pistillate flowers, as the type of the species, Elmer 9241,
was with fruit only, although nearly all the above specimens have long been
available for description. Many of the specimens cited above have abnormal,
much-branched, often leafy inflorescences, probably due to the work of some
insect: these abnormal inflorescences apparently never bear normal flowers.
Similar ones are found in Canariwm, the form of C. villosum described by Engler
as CO. luawrians var. monstroswm, and in some species of Hugenia.

Local names given for this species are Malagaitmon, Alinao, Anang, Bolongeta,
Panigilman, and Bagnito, many of which are also applied to other species of the
genus, although of the above, Malagaitmon seems to be rather consistently applied
to the present species.

Diospyros everettii sp. nov.

Arbor glabra vel subglabra; foliis oblongis vel oblongo-lanceolatis, sub-
coriaceis, in sicco brunneis, usque ad 15 em longis, apice acuminatis, basi
late rotundatis subcordatisque, nervis utrinque circiter 10, ascendentibus,
anastomosantibus, reticulis laxis; floribus femineis fasciculatis, axillari-
bus, sessilibus, 4-meris, circiter 22 mm longis, calycis lobis lanceolatis ;
staminodeis glabris, 4; ovario dense hirsuto.

A tree glabrous or subglabrous. Branches dark-colored, glabrous.
Leaves alternate, oblong or oblong-lanceolate, subcoriaceous, brownish
when dry especially beneath, the upper surface dull, the lower slightly
shining, glabrous, the apex acuminate, the base rather broad, rounded
and subcordate, 8 to 15 em long, 2.5 to 4.5 em wide; nerves about 10 on
each side of the midrib, ascending, attenuate-anastomosing, the reticula-
tions lax; petioles stout, about 3 mm long. Pistillate flowers axillary,
sessile, fascicled, just before opening 22 mm long. Calyx cleft nearly to
the base into four, lanceolate, spreading, acute or acuminate lobes 7 to 8
mm long, 2.5 mm wide, their margins slightly ciliate. Corolla salver-
shaped, the tube cylindric, 9 mm long, 4 mm thick below and slightly
contracted above, inside glabrous, outside with very few long hairs; lobes
4, spreading, elliptic-oblong, 12 mm long, 6 mm wide. Staminodes 4,
their filaments 2.5 mm long, the anther-like portion 1.5 mm long,
glabrous. Ovary densely covered with very stiff, 2 to 3 mm long, brown-
ish hairs; style stout, cylindric, 6 mm long, pubescent, 4-cleft at the apex.
The ovary is apparently 8- or 10-celled.

Necros, Mount Silay, For. Bur. 7261 Everett, May 9, 1907, in ridge forests at
an altitude of about 650 m.

A species apparently belonging to the section Hrmellinus, and is well charac-
terized by its leaves being rounded and subcordate at the base, and by its

comparatively large, axillary, fascicled, 4-merous flowers, and the very numerous,
stiff, brown hairs that entirely cover the ovary.

308 MERRILL.

Diospyros foveo-reticulata sp. nov. uke

Arbor inflorescentiis fructibusque exceptis glabra, circiter 20 na
foliis subcoriaceis vel coriaceis, oblongis, elliptico-oblongis vel oblo
lanceolatis, usque ad 30 cm longis, utrinque nitidis, dense foveo-retic
latis, subtus pallidioribus, -basi 2-glandulosis, acuminatis, basi acutis,
rariter rotundatis; fructibus axillaribus, solitariis, pedunculatis, ovoideis s
vel subglobosis, 8- vel 10-locellatis junioribus 2 ad 2.5 em diametro, palli- h
dis, ferrugineo-pubescentibus, pericarpio coriaceo ; calycis lobis 4, reflexis, —
ovatis, crassis, acutis vel obtusis, utrinque ferrugineo-pubescentibus, —
circiter 8 mm longis.

A glabrous tree, except the inflorescence and fruits, about 20 m high.
Branches dark-colored, terete, the branchlets paler, sometimes yellowish,
glabrous or minutely puberulent. Leaves alternate, subcoriaceous or’ —
coriaceous, oblong, elliptic-oblong, or oblong-lanceolate, rather pale when
dry, frequently yellowish, 15 to 30 cm long, 4 to 9 em wide, glabrous,
shining on both surfaces and densely foveolate-reticulate, the apex short-
acuminate, the base acute, sometimes rounded, the lower surface near
the base with a rather prominent gland on each side of the midrib;
primary lateral nerves 10 to 12 on each side of the midrib, indistinct,
faintly anastomosing; petioles 1 to 1.5 em long. Fruit (immature)
axillary, solitary, ovoid or subglobose, 2 to 2.5 cm in diameter, pale, the
pericarp crustaceous, thin, deciduously ferruginous-pubescent, 8- or 10-
celled, the calyx persistent, accrescent, 4-lobed, the tube very short, the
lobes very thick, crustaceo-coriaceous, ovate, reflexed, about 8 mm long |
and nearly as wide, acute or obtuse, densely ferruginous-pubescent on
both sides.

Luzon, Province of Camarines, Pasacao, Ahern 29, 278 (type), 790, January
to June, 1902, the last two with immature fruits; Lupi, Por. Bur. 10780 Curran,
July, 1998, sterile. Minpanao, District of Zamboanga, Port Banga, For, Bur,
9062 Whitford & Hutchinson, December, 1907; San Ramon, Hallier s, n., Feb-
ruary, 1904, with immature fruits.

This species is well characterized by its densely foveolate-reticulate leaves
which are prominently 2-glandular on the lower surface near the base, and by
its thickned, ovate, reflexed, 4-lobed calyx which is densely ferruginous-pubescent,
as are the young fruits. The plants under Ahern’s name, cited above, were
erroneously distributed as Diospyros discolor, to which the present species is not
closely allied. It is known in the Camarines as Alahan, and at Port Banga as
Palo negro. The section is undeterminable from the material available.

Diospyros inclusa sp. nov.

Arbor glabra, circiter 15 m alta; foliis coriaceis vel subcoriaceis, —
oblongis vel elliptico-oblongis, glabris, nitidis, apice breviter obtuse acumi-_
natis, basi acutis, circiter 9 cm longis, eglandulosis, nervis utrinque —
circiter 7, vix distinetis, reticulato-anastomosantibus; fructibus axillari-_
bus, solitariis, pedunculatis, depresso-globosis, nigris, apiculatis, glabris
vel parce adpresso-hirsutis, 1 cm diametro, circiter 8-locellatis, calyce eu
accrescente, ellipsoideo, crasso, lenticellato, nigro, glabro, breviter 4- v ol
5-lobato inclusis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 309

A glabrous tree about 15 m high. Branches terete, light-gray, glab-
rous, shining, somewhat striate when dry, the branchlets dark-colored,
their tips slightly pubescent. Leaves coriaceous or subcoriaceous, oblong
or elliptic-oblong, glabrous, shining on both surfaces, 7 to 10 cm long,
3 to 4.5 em wide, the apex shortly and obtusely acuminate, the base acute,
eglandular; primary lateral nerves about 7 on each side of the midrib,
not very distinct, irregular, anastomosing, the secondary ones and reticula-
tions nearly as prominent; petioles black, glabrous, about 6 mm long.
Flowers unknown. Fruits axillary, solitary, on stout, 3 to 4 mm long
peduncles which are usually spreading, sometimes recurved, quite inclosed
by the accrescent calyx, including the calyx ellipsoid or ovoid, 1.5 to 2
em long. Calyx black, glabrous, shining, lenticellate, thickly crustaceous,
inside pubescent, the apex shortly and obscurely 4- or 5-lobed, the opening
at the top about 6 mm in diameter. Fruit free from the calyx except at
the base, depressed-globose, black, shining, glabrous or with few appressed
hairs, about 1 cm in diameter, apex apiculate, usually 8-celled.

Maspate, Aroroy, Whitford 1682, October, 1906, on hillsides at an altitude of
about 240 m. Locally known as Bantolinao.

A most characteristic species, recognizable by its accrescent, ellipsoid or
ovoid, crustaceous, lenticellate calyx which entirely incloses the fruit, except for
the small opening at the apex. The section is uncertain, as flowers are necessary
to determine it.

Diospyros mindanaensis sp. noy. § JMJelonia.

Arbor glabra 15 ad 25 m alta; foliis oblongis vel oblongo-lanceolatis,
coriaceis vel subcoriaceis, glabris, nitidis, apice acuminatis, basi rotun-
datis, acutis vel leviter acuminatis, usque ad 25 cm longis; floribus
femineis racemosis, racemis axillaribus, paucifloris, calycibus breviter
4-dentatis ; fructibus globosis, vel depresso-globosis, circiter 4 cm diametro,
4-locellatis, glabris, nitidis, calycibus accrescentibus persistentibus, crasse
coriaceis, reflexis, tubo acuminate 4-angulato, breviter 4-dentato ; semini-
bus osseis, albumine ruminato.

A glabrous or subglabrous tree, 15 to 25 m high. Branches terete,
glabrous, dark-gray. Leaves alternate, coriaceous or subcoriaceous, oblong
or oblong-lanceolate, 10 to 25 cm long, 4 to 8 em wide, glabrous, the
upper surface shining, the lower somewhat paler, dull, apex acuminate,
base rounded, acute or sometimes slightly acuminate, eglandular; nerves
10 to 13 on each side of the midrib, rather distinct beneath, somewhat
arched-ascending, anastomosing, the secondary nerves distinct, the re-
ticulations fairly dense; petioles stout, 1 to 1.5 em long. Pistillate
flowers 4-merous, in axillary, solitary, few-flowered racemes, the racemes
(when young), about 2 em long, glabrous or with few, scattered, appressed
hairs. Calyx cup-shaped, shortly 4-toothed. Fruit globose or depressed-
globose, 3 to 4.5 cm in diameter, glabrous, shining, yellowish when
mature, the pericarp crustaceous, thick, 8-celled, 8-seeded, the seeds
ellipsoid, somewhat compressed, 2 cm long, 1.5 em wide, brown outside,

310 MERRILL.

the albumen bony, ruminate. The persistent calyx is accrescent, thickly |

coriaceous, glabrous, reflexed, forming an acuminately 4-angled tube 1

to 1.5 cm long, and diagonally 2 to 2.5 em in diameter, with four short, —

acute or acuminate teeth, one at each corner. The side-walls of the
calyx-tube are concave, so that the apex of the reflexed calyx-tube is
formed of four lobes radiating from the center to the corners of the
tube. The peduncle is about 2 cm long in mature fruits.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9453 (type), 9175
Whitford & Hutchinson, January, February, 1908, with mature fruits. BASILAN,
San Rafael, For. Bur. 6098 Hutchinson, January, 1907, with mature fruits. Nr-
GRos, Gimagaan River, For. Bur. 4243, 4269 Everett, April, May, 1906, the
former with abnormal inflorescence, the latter with immature flowers; Cadiz,
For, Bur. 4309 Everett, For. Bur. 7407 Danao, June 1906, 1907, the former with
abnormal inflorescence, the latter sterile. Luzon, Province of Tayabas, Guina-
yangan, Merrill 2007 March, 1903, with immature fruits.

This species is well characterized by its comparatively large, 8-celled, 8-seeded
fruits, its ruminated albumen, and especially by its persistent calyx which forms
an acuminately 4-angled, reflexed tube. It is apparently allied to the Bornean
Diospyros korthalsiana Hiern, but has much larger leaves, and a quite different
calyx-tube. In some respects it resembles D. subrigida Hochr., but is quite
distinct from that species. Like D. cwrranii Merr. this species frequently has an
abnormal, densely much branched inflorescence, apparently due to whe work of
insects. It is known in Negros as Ata-ata, in Tayabas as Anang, and in Basilan
as Bolongita.

Diospyros montana Roxb. Pl. Coromend. 1 (1795) 37, t. 48; Hiern Monog.
Eben. (1873) 220.

Luzon, Province of Nueva Ecija, near Cabanatuan, Bur, Sci. 5266 McGregor,
September, 1908.

A species not previously found in the Philippines, the above specimen agreeing
rather closely with the form considered by Hiern as the variety cordifolia, exeept
that the leaves are not cordate.

India to the Moluceas and northern Australia.

Var. parva var. noy.

Differt a typo foliis multo minoribus, 1 ad 2.5 em longis, subtus dense
molliter pubescentibus; fructibus globosis, Junioribus 8 mm diametro,
glabris, nigris, apiculatis, 8-locellatis; calycis lobis accrescentibus, re-
trorsis.

Luzon, Province of Ilocos Norte, Badoe, For. Bur. 18953 Merritt & Darling,
November, 1908, on rocky hillsides at an altitude of about 65 m. Il., Antinagam.

Well characterized by its small densely pubescent leaves; additional material
may show it to be worthy of specific rank.

Diospyros phanerophlebia sp. nov. '

Arbor 5 ad 10-m alta, glabra, inflorescentiis exceptiis; foliis crasse
coriaceis, oblongis vel oblongo-lanceolatis, nitidis, basi acutis, subtus
2-glandulosis, apice obtusis vel leviter obtuso-acuminatis, usque ad 20
em longis; nervis utrinque 8 ad 10, subtus prominentibus, elevatis,

ae

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 311

curyato-ascendentibus, anastomosantibus, reticulis laxis, prominentibus,
elevatis ; petiolo crasso, 1 ad 1.5 em longo; fructibus ovoideis vel depresso-
globosis, solitariis vel binis in axillis defoliatis, in sicco nigris, glabris,
nitidis, 1.5 ad 1.8 cm diametro, 8-locellatis, locellis monospermis.

A tree 5 to 10 m high, glabrous except the inflorescence. Branches
terete, grayish, the younger ones slender, black, dark-brown, or oliva-
ceous, smooth. Leaves thickly coriaceous, oblong, oblong-lanceolate,
rarely lanceolate, 15 to 20 cm long, 4 to 8 cm wide, of about the same
color on both surfaces or the lower slightly paler, the wpper shining, the
apex obtuse or slightly and obtusely acuminate, the base acute, margins
often slightly revolute, the lower surface with a distinct gland at the
base on each side of the petiole; primary nerves 8 to 10 on each side of
the midrib, very prominent on the lower surface, elevated, curved-
ascending, anastomosing, the primary reticulations lax, nearly as prom-
inent as the nerves, the ultimate reticulations less distinct ; petioles
stout, black when dry, 1 to 1.5 cm long. Flowers unknown. Fruit
solitary or in pairs in the axils of fallen leaves, subsessile, numerous,
ovoid or depressed-globose, 1.5 to 1.8 em in diameter, black and shining
when dry, glabrous, the apex slightly apiculate, 8-celled, each cell with
a single seed. Calyx accrescent, closely appressed to the base of the
fruit, more or less pubescent outside, densely so within, nearly square
and subtruncate, or with four short, broad teeth, 1.2 to 1.5 em wide at
the top. Seeds about 8 mm long, 5 mm wide, compressed, brown, shining,
smooth, the albumen not ruminate.

Type specimen collected by H. M. Ourran, For. Bur. 10998, District of Bontoe,
Luzon, January, 1909, locally known as Aliuac. I am disposed to refer to it
also the following specimens, all from Luzon: Province of Bulacan, For. Bur. 7.172
Curran, June, 1907: Province of Rizal, San Mateo, For. Bur. 1847 Ahern’s col-
lector; Merrill 236, Decades Philippine Forest Flora: Province of Laguna, Santa
Maria Mavitac, For. Bur. 10052 Curran, locally known in the last three provinces
as Canomoi.

A species well characterized by its very prominent veins and reticulations,
manifestly allied to Diospyros maritima Blume, and to D. canomoi <A. DC.,
differing especially from the former in its much thicker leaves and very prominent
nerves, and from the latter in being nearly glabrous throughout.

Diospyros whitfordii sp. nov.

Arbor circiter 18: m alta; ramulis junioribus densissime ferrugineo-
pubescentibus; foliis coriaceis vel subcoriaceis, oblongis vel elliptico-
oblongis, 8 ad 18 cm longis, breviter acuminatis, basi rotundatis vel
obtusis, utrinque nitidis, supra glabris, subtus glabris vel ad costa pubes-
centibus ; fructibus globoso-ovoideis, 5 cm diametro, extus nigris, plus
minus pruinosis, glabris, in sicco plus minus reticulato-rugosis, pericarpio
osseo vel ligneo; calycibus persistentibus, accrescentibus, 5-lobatis, 4.5
ad 5 cm diametro, crasse coriaceis, lobis triangulari-ovatis, acutis, vix
reflexis, utrinque densissime ferrugineo-pubescentibus.

of .o, --e- ° ee ee

312 : MERRILL. ee

A tree about 18 m high. Branches glabrous, grayish-brown, $01
times pruinose, the younger branchlets densely ferruginous-pubescent 7
Leaves coriaceous or subcoriaceous, oblong or elliptic-oblong, 8 to 18 cm
long, 3 to 6 em wide, the apex shortly acuminate, the base rounded ore
obtuse, both surfaces shining, the upper one glabrous, the lower glabro
or more or less pubescent on the midrib; nerves 6 to 8 on each side of a
the midrib, not very prominent, anastomosing, the secondary ones and — :
the reticulations distinct on the lower surface, rather close; petioles 8
to 10 em long, more or less pubescent. Fruit ovoid-globose, about 5 em
in diameter, the pericarp bony or woody, outside black, more or less
pruinose, reticulate-rugose when dry, glabrous, the persistent, 5-lobed
calyx appressed to the base of the fruit, 4.5 to 5 em in diameter, the
lobes united for half their length, triangular-ovate, spreading, acute,
thickly coriaceous or almost woody, both sides densely ferruginous-
pubescent. The central portion of the calyx is depressed, the depression
extending along the median portions of the lobes, while there are five
prominent protuberances at the juncture of the lobes.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9019 Whitford,
November, 1907, in fruit; Avala, Por. Bur. 4811 Hutchinson, July, 1906, sterile.
On forested ridges at an altitude of about 30 m, locally known as Camagon. The
fruit is immature and is described by Whitford as green. :

OLACACEA®.
LINOCIERA Swartz.
Linociera acuminatissima sp. nov.

Arbor glabra, circiter 9 m alta, ramis ramulisque teretibus, pallidis
foliis coriaceis, nitidis, oblongo-ellipticis, basi acuminatis, apice valde
caudato-acuminatis, margine plus minus recuryatis; inflorescentiis race-_
mosis, axillaribus, racemis solitariis, paueifloris, 2 ad 3 em longis; floribus—
circiter 4 mm longis.

A tree, glabrous throughout, about 9m high. Branches and branchlets
slender, terete, pale-gray, sparingly lenticellate. Leaves opposite, coria-
ceous, somewhat shining, oblong-elliptic, 8 to 12 em long, 2.5 to 4 em
wide, olivaceous above when dry, brownish and paler beneath, the base
somewhat acuminate, the apex rather strongly caudate-acuminate, the
acumen 1.5 to 2 cm long, slender, narrowed upward, blunt, the margins
somewhat recurved ; nerves about 7 on each side of the midrib, indistinct,
distant, obscurely anastomosing, the reticulations obsolete; petioles 1 to
1.5 em long. Racemes axillary, solitary, 2 to 3 em long, few-flowered,
the pedicels stout, 1 to 4 mm long, the bracteoles narrowly oblong or
lanceolate, obtuse or rarely acuminate, about 2 mim long. Calyx-lobes
broadly ovate, acute, 1 mm long. Corolla-lobes lanceolate, obtuse, 4 mm
long, somewhat connate below. Stamens 2, the filaments broad, short; 3
anthers elliptic, 1 mm long. a

PALAWAN, Malampaya Bay, For. Bur. 4506, 7454 Curran, June 21, 1906.
Well characterized by its caudate-acuminate leaves.

NEW OR NOTEWORTHY* PHILIPPINE PLANTS, VII. 313

Linociera philippinensis nom. nov.

Mayepea pallida Merr. in Govt. Lab. Publ. (Philip.) 35 (1906) 58.

Linociera pallida Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 116, non Kx. Sch.

Olea sp. Vidal Phan. Cuming. Philip. (1885) 125; Rev. Pl. Vase. Filip. (1886)
181.

The above change of name is necessitated because of the previous use of the
name pallida for a different species of the same genus, by K. Schumann. [I also
include under this species the specimens referred by Vidal to Olea sp. in Phan.
Cuming. Philip. (1885) 125; Rev. Pl. Vase. Filip. (1886) 181, as the specimens
cited appear to me to be Linociera and not Olea. Linociera philippinensis is
represented by the following specimens:

Luzon, Province of Bataan, Mount Mariveles, For. Bur. 2792 Meyer, For. Bur.
2939 Borden, Whitford 1142, Williams 569: Province of Rizal, Merrill 1835,
2670, For. Bur. 2874 Ahern’s collector, Bur. Sci. 1457, 3356 Ramos: Province of
Camarines Sur, Pasacao, Ahern 44: Province of Zambales, For. Bur. 6942 Curran.
Mrnporo, Pola, Merrill 2255. Guimaras, For. Bur. 267, 280, 309 Gammill.
Native names, T., Anatao, Malabocboc, Pulat; V., Magubay, Cabating.

APOCYNACEZE.
ALIXIA Banks.

Alyxia luzoniensis sp. nov.

Frutex scandens, glabra; foliis oppositis, ternis, vel rariter quarternis,
oblongis, ovato-oblongis, vel elliptico-oblongis, coriaceis, nitidis, usque ad
5 cm longis, apice breviter obtuse acuminatis, basi acutis, nervis laterali-
bus subobsoletis; cymis axillaribus, paucifloris, pedunculatis; fructibus
ellipsoideis, obtusis vel apiculatis, 1 ad 1.4 cm longis.

A scandent shrub, glabrous throughout. Branches terete or slightly
angled, brownish, rather slender. Leaves opposite, mostly ternate, or
rarely quaternate, oblong, ovate-oblong, or elliptic-oblong, coriaceous, shin-
ing, 2 to 5 cm long, 1 to 2 cm wide, the apex shortly and obscurely blunt-
acuminate, the base acute, margins slightly recurved; midrib prominent,
the lateral nerves nearly obsolete, very slender; petioles 1 to 2 mm long.
Cymes axillary, solitary, few-flowered, the peduncles 1 to 1.5 cm long,
the buds congested, calyx-segments about 1 mm long. Flowers unknown.
Fruits ellipsoid, glabrous, obtuse or apiculate, 1 to 1.4 cm long, about
8 mm thick, very rarely the carpel is constricted in a moniliform manner,
bearing above the constriction a second seed-bearing portion.

Luzon, Province of Ilocos Norte, Mount Piao, For. Bur. 13977, 13988 Merritt
& Darling, altitude 1,000 m; Vintar, Por. Bur. 13945 Merritt & Darling: District
of Bontoc, Bur. Sci. 7007 Ramos (type): Province of Zambales, Mount Tapulao,
For. Bur. 8069 Curran & Merritt, Bur. Sci. 5107 Ramos, altitude 2,000 m.

This species is apparently most closely allied to Alyxia sinensis Champ., of
southern China, but seems to be quite distinct from that, as well as from the
previously described Philippine forms. It is apparently also closely allied to
Alyxia parvifolia (Gynopogon parvifolia Merr.), but has much larger and
differently shaped leaves. It is at once distinguishable from A. monilifera Vidal
by its very different fruit, and practically nerveless leaves. Like the other
Philippine species of the genus dried specimens have a strong odor of cumarin.

857548

314 MERRILL.

CHONEMORPHA G. Don.

Chonemorpha elastica sp. noy.

Frutex alte scandens, ramis ramulisque rubro-brunneis, glabris; foliis
membranaceis vel submembranaceis ovato-ellipticis vel obovato-ellipticis,
15 ad 20 em longis, breviter acuminatis, basi rotundatis, subtus plus
minus pilosis; floribus albis, 2.5 ad 3 em diametro; calycibus 5-partitis.

A scandent shrub of large size. Branches and branchlets reddish-

brown, glabrous, usually shining. Leaves opposite, membranaceous or 5
submembranaceous, ovate-elliptic or obovate-elliptic, 15 to 20 cm long, 8 ;

to 15 cm wide, entire, the base rather broad, rounded, the apex shortly
acuminate, the upper surface glabrous, shghtly shining, the lower surface
slightly paler and somewhat pilose; petioles rather slender, 2.5 to 4 em
long ; nerves about 10 on each side of the midrib, distinct on both surfaces,
the reticulations very distinct beneath. Cymes terminal, few-flowered,
about 6 cm long, glabrous or nearly so, the bracteoles more or less pubes-
cent, about 3 mm long. Flowers white, fragrant, their pedicels slender,
1 cm long or less. Calyx 5-partite, the lobes oblong, somewhat acuminate,
the margins above somewhat ciliate, about 5 mm long, 2 mm wide, united
below. Corolla-tube 9 to 10 mm long, swollen and about 3 mm in diam-
eter in the lower third, contracted above, glabrous outside, inside with
reflexed hairs below the insertion of the stamens; lobes 5, obliquely
obovate, spreading, about 15 em long, 12 mm wide. Stamens 5, the fila-
ments short, inserted at about the lower one-fourth of the tube; anthers
narrowly-lanceolate, acuminate, 4 mm long, sagittate at the base. Disk
entire, truncate, 1 mm high. Ovaries and style 4 mm long, the ovaries
two, free, united by the style; ovules many. Fruit unknown.

MINDANAO, District of Zamboanga, Dumanquilas Bay, Tigbalubu, For. Bur. ,
12351 Hutchinson, May, 1908, in forests at an altitude of about 40 m: District
of Davao, H. S. Peabody, July, 1906, leaves only. Basiran, P. L. Sherman, July,
1903, leaves only; also reported from Mindanao and from Tawi-Tawi by Sherman.

So far as is known at present this is the most important rubber producing
vine in the Philippines. It is one of the two species considered by P. L. Sherman
in his paper on “Gutta-Percha and Rubber in the Philippine Islands,” * and an
illustration is given by him, |. ¢. fig. 33, of leaf specimens collected on Basilan
Island. This specimen, as well as the one collected by Mr. Peabody, cited above,
were previously provisionally identified as Chonemorpha, but the determination
could not be verified until flowering specimens were received. Mr. Sherman
reports this vine from Mindanao, Basilan, and Tawi-Tawi, but states that only
the natives of Tawi-Tawi understand the commercial value of the product, and
that they gather the latex and mix it with gutta-percha. No information is at
hand as to the amount exported, but it is probably very small. According to Mr, — a
Sherman’s notes, the vine attains a length of from 150 to 200 feet, and a diameter
of from 6 to 8 inches.

Apparently allied to Chonemorpha griffithii Hook. f., of British India, especially
in its deeply cleft calyx, but the branches not hispid. Without comparison with
the type of Hooker’s species, it is impossible to give other differential characters,
as Hooker's deseription is too short and imperfect.

“ Publications of the Bureau of Government Laboratories, 7 (1903), 36-389.

NEW OR NOTEWORTHY: PHILIPPINE PLANTS, VII. ote

KICKXIA Blume.

Kickxia merrittii sp. nov.

Arbor glabra, 18 ad 20 m alta; foliis oppositis, subcoriaceis, oblongo-
vel elliptico-lanceolatis, utrinque acuminatis, 7 ad 12 cm longis, usque
ad 3.5 em latis; floribus albis, axillaribus, solitariis vel binis, pedicellatis,
7 em longis, corollae tubo 2.5 em longo.

A tree 18 to 20 m ligh, glabrous throughout. Branches terete, lenti-
cellate, dark-reddish-brown, the branchlets slightly compressed. Leaves
opposite, subcoriaceous, oblong-lanceolate or elliptic-lanceolate, 7 to 12 cm
long, 2 to 3.5 cm wide, entire, shining, the apex acuminate, acumen
blunt, the base acuminate; nerves about 7 on each side of the midrib,
beneath rather distinct and often reddish-brown; petioles 1 cm long or
less. Flowers axillary, solitary or in pairs, white, about 7 cm long, the
pedicels 2 cm long or less. Calyx-lobes oblong-ovate, acuminate, keeled,
5 mm long, 3 mm wide. Corolla-tube 2.5 cm long, narrow below, some-
what inflated above, the lobes 5, about 4 em long, 1.3 em wide. Anthers
about 7 mm long. Disk 2 mm long, truncate. Ovaries elongated, united
by the style, the latter 8 mm long. Follicles (immature) cylindric, 8
em long, about 2 em thick, seeds spindle-shaped, long acuminate, about
4 em long, the hairs brownish, 4 cm long.

Mrnporo, Ibalo River, For. Bur. 11488 Merritt, May 9, 1908, in mountain
forests at 600 m altitude. Mang., Ayete.

The second species of the genus to be found in the Philippines, distinguished
from Kickxia blancoi Rolfe by its elongated corolla tube, and more acuminate

leaves.
OCHROSIA Juss.

Ochrosia littoralis sp. nov.

Arbor circiter 10 m alta, glabra; foliis ternis vel quaternis, membrana-
ceis, late oblanceolatis, breviter obtuseque acuminatis, basi cuneatis, usque
ad 8 cm longis; cymis axillaribus terminalibusque, 2 ad 35 cm longis;
floribus circiter 1 cm longis; drupis 2.5 ad 4 em longis, divergentibus,
acuminatis, subeylindraceis, basi connatis.

A tree about 10 m high, glabrous throughout. Branches slender,
terete, reddish-brown. Leaves usually ternate, rarely quaternate, mem-
branaceous, broadly oblanceolate, 6 to 8 em long, 2 to 2.5 cm wide,
shining, the apex shortly and obtusely acuminate, the base cuneate ;
nerves very numerous, parallel, not prominent; petioles 4 to 5 mm long,
sometimes shorter. Cymes axillary and terminal, 2 to 3 cm long, the
bracts and bracteoles ovate, 1 to 1.5 mm long. Flowers mostly at the
apices of the branchlets, usually crowded. Calyx-lobes ovate to oblong-
ovate, obtuse or acute, imbricate, about 2 mm long. Corolla-tube cyln-
dric, slender, about 6 mm long, 1.5 mm in diameter, glabrous throughout,
the lobes 5, narrow, about 4 mm long, overlapping to the right. Stamens
inserted just above the middle of the tube; anthers lanceolate, acuminate,
1.3 mm long. Ovaries two, united at the base, and at the apex by the 2

316 MERRILL.

e
mm long style; ovules 6 to 8 in each. Drupes united at the base, divari-
cate, the pericarp bony, the mesocarp corky, subcylindric, 2.5 to 4 em
long, 1 to 1.5 cm in diameter, somewhat acuminate; seeds 6 to 8 in each
drupe, 3 or 4 on each side of the placenta, compressed, orbicular, 4 to 6
mm in diameter.

Luzon, Province of Batangas, Malabrigo, for. Bur. 7754 Curran & Merritt,
November, 1907, along the seashore.

A species well characterized by its ternate leaves, short cymes, and subeylindrie,
somewhat acuminate drupes, which are not flattened on the inner surface, and
by its unusually large number of seeds. The first species of the genus to be
found in the Philippines.

RAUWOLFIA Linn.

Rauwolfia samarensis sp. nov.

Arbor glabra, circiter 8 m alta; foliis’ quarternis, oblongo-ellipticis,
breviter acuteque acuminatis vel acutis, chartaceis, circiter 20 cm longis,
6 cm latis, nervis utringue 30 ad 35; pedunculis e axillis terminalibus,
superne umbellatis; baccis ellipsoideis, atropurpureis, circiter 1.5 cm
longis, pericarpio carnoso.

A tree about 8 m high, glabrous throughout. Branches light-gray,
stout, the leaf-scars very prominent. Leaves in whorls of four, oblong-
elliptic, 13 to 20 em long, 4 to 7 em wide, the apex shortly and sharply
acuminate or acute, the base acute or slightly decurrent-acuminate, char-
taceous, shining, the lower surface paler than the upper; primary nerves
30 to 35 on each side of the midrib, distinct, the intermediate secondary
ones evident; petioles 1.5 to 3 cm long. Inflorescence from the terminal
axils, the peduncles in anthesis about 5 cm long, in fruit elongated and
the panicle ultimately about as long as the leaves, the branches umbellately
arranged at the apex of the peduncle. Flowers white, umbellately dis-
posed, the pedicles 1 to 2 mm long. Calyx 3 mm long, the lobes obtuse. —
Corolla 5 mm long, cylindric, the lobes orbicular-ovate, 2 mm long, the
throat hirsute. Anthers lanceolate, 1.3 mm long. Style about 1.5 mm
long. Fruit ellipsoid, dark-purple when mature, the pericarp fleshy.

SAMAR, Lanang, Merrill 5233, October 3, 1906, in thickets on bluffs along the
seashore at an altitude of about 10 m.

A species allied to Rawwolfia spectabilis (Miq.) Boerl., R. javanica Koord. &

Valet., and to R. sumatrana Jack., of Malaya, but apparently distinct from all.
Well characterized by its many-nerved leaves and rather fleshy fruits.

TABERNAEMONTANA Linn.

Tabernaemontana caudata sp. nov.

Arbuscula circiter 2 m alta, glabra; ramis ramulisque gracilibus,
pallidis; foliis ovato-lanceolatis vel elliptico-ovatis, usque ad 4 cm longis,
basi acutis vel acuminatis, apice valde caudato-acuminatis; floribus paucis
axillaribus, cymosis, albis, circiter 12 mm longis, calycis lobis ovatis,
0.5 mm longis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 317

A shrub about 2 m high, glabrous throughout. Branches and
branchlets slender, terete, pale-gray, somewhat shining. Leaves ovate-
lanceolate to elliptic-lanceolate, 2.5 to 4 cm long, 1 to 1.5 em wide,
chartaceous or submembranaceous, shining, the base acute or acuminate,
sometimes slightly inequilateral, the apex strongly caudate-acuminate,
the acumen about one-third as long as the blade, blunt; nerves 6 to 8
on each side of the midrib, faint, spreading, anastomosing, the reticula-
tions very obscure; petioles very slender, about 5 mm long. Flowers in
few-flowered, axillary, solitary cymes, but two or three flowers in a
cyme, the peduncles very short, the pedicels slender, 1 em long. Calyx-
teeth ovate, blunt, 0.5 mm long. Corolla-tube 12 mm long, slender,
the lobes narrowly oblong, 7 mm long, 3 mm wide. Anthers 2 mm long.
Carpels narrow, about 2 mm long; styles 10 mm in length. Fruit
unknown.

aa

7

Luzon, Province of Ilocos Sur, Mount Bulangao, Por. Bur. 14025 Merritt &
Darling, November, 1908, in forests at an altitude of about 1,150 m.

A species well characterized by its small, very strongly caudate-acuminate
leaves, which do not exceed 4 em in length, and its few-flowered cymes.

Se tei TS ict taint fs Ay
< ny 7

Tabernaemontana linearifolia sp. noy.

Arbuscula glabra, circiter 4 m alta; foliis chartaceis, lnearibus vel
anguste lanceolato-linearibus, utrinque angustatis, apice gradatim acum-
inatis, usque ad 7 cm longis, 5 ad 8 mm latis; floribus axillaribus,
solitariis, longe pedicellatis ; folliculis oblongis, utrinque angustatis acum-
inatisque, circiter 2.5 cm longis.

A glabrous shrub about 4 m high. Branches and branchlets slender,
terete, gray, shining. Leaves linear or narrowly lanceolate-lnear, 3 to
7 em long, 5 to 8 mm wide, chartaceous, somewhat shining, olivaceous
above when dry, the lower surface paler and somewhat brownish, nar-
rowed at both ends, the apex gradually acuminate, the base acute ; lateral
nerves distant, indistinct, spreading, the reticulations obsolete; petioles
2to4mm long. Flowers axillary, solitary, white, their peduncles often
3 em long, frequently much shorter. Calyx-teeth ovate, acute or obtuse,
1.5 mm long. Corolla-tube 13 mm long, slender, the lobes oblique, acute,
about 7 mm long, 3 mm wide. Carpels lanceolate, narrowed above,
3 mm long; styles 6 to 7 mm long. Follicle oblong, orange-yellow,
nearly 2.5 em long, 8 mm in diameter in the middle, acuminately
narrowed at both ends, with three faint ridges extending the whole length,
the apical portion with five faint ridges: seed solitary, narrowly ellipsoid,
8 mm long, the groove not prominent.

3
)

Luzon, District of Lepanto, Mancayan, For. Bur. 10945 Curran, January, 1909,
in open grass-lands in thin pine forests, altitude 1,600 m.

A species well characterized by its very arrow, elongated, linear or linear-
lanceolate leaves, its solitary, frequently long-pediceled flowers, and its one-
seeded follicles which are acuminately narrowed at both ends.

318 MERRILL.

Tabernaemontana megacarpa sp. nov. .

Arbor circiter 10 m alta, glabra; foliis ellipticis, oblongo-elliptie
vel obovato-ellipticis, apice rotundatis, basi acutis vel obtusis, 15 ad 3
cm longis, nervis utrinque 20 ad 24, wapininennibags paniculis termina- -
libus, circiter 20 cm longis; fioribus circiter 3.5 em diametro; fructibus —
carnosis, laevibus, rubris, in sicco rugosis, griseis, circiter 8 em ional ;
+ em latis, vix paciaine . ee |

A tree about 10 m high, glabrous throughout. Branches and branch- — P
lets light-gray, glabrous, stout. Leaves opposite, firmly chartaceous to |
subcoriaceous, somewhat shining above, dull beneath, elliptic, oblong-
elliptic or obovate-elliptic, 15 to 30 cm long, 9 to 15 em wide, the apex
broad, rounded, the base acute or obtuse; nerves 20 to 24 on each side
of the midrib, very prominent, the reticulations obsolete or nearly so;
petioles about 2 cm long, the base inflated and somewhat clasping the
branches. Panicles terminal, about 20 cm long. Calyx-lobes broadly
ovate, about 5 mm long. Corolla-tube 2 cm long, the hmb spreading, 3
to 3.5 cm in diameter. Follicles smooth and red when fresh, fleshy,
when dry strongly wrinkled, gray or brownish, somewhat curved, not
costate, about 8 cm long, 4 to 5 em wide and 3 cm ES Seeds nu-
merous, 10 to 12 mm long. .

Minpanao, Lake Lanao, Camp Keithley, Mrs. Clemens s. n., June, April, and
September, 1906: Province of Misamis, Mount Malindang, For. Bur. 4781 Mearns —
& Hutchinson, May, 1906: District of Zamboanga, Copeland s. n., April, 1905;
Sax River, Williams 2179, February, 1905. Bastian, For. Bur. 3964 Hutchinson,
January, 1906; Hallier s. n., January, 1904.

This species is allied to Vocanga plumeriaefolia Elmer Leaftl. Philip. Bot. 1
(1908) 333, but has relatively broader leaves, and more numerous nerves. The
type of Voacanga plumeriaefolia Elm., is a fruiting specimen, and so far as I
can determine from the material before me, is a Tabernaemontana, but flowers are
necessary to determine exactly to which genus it really belongs.

Tabernaemontana mucronata sp. nov.

Arbor parva, circiter 6 m alta, glabra; ramis griseis, teretibus, ramulis ‘ig
plus minus angulatis; foliis membranaceis, oblongo-ellipticis, apice =
acuminatis, acuminibus apiculatis, 6 ad 10 cm longis, 2 ad 4 em latis, -
nervis utringue 9 yel 10; cymes axillaribus, brevibus, solitariis vel fas-
ciculatis, circiter 2 em longis; floribus gracilibus, circiter 12 mm longis.

A small tree about 6 m high, glabrous thoughout. Branches terete,
gray, the branchlets somewhat angled. Leaves membranaceous, oblong-
elliptic, 6 to 10 em. long, 2 to 4 cm wide, the apex rather prominently
acuminate, the acumen apiculate, the base somewhat decurrent-acuminate,
the upper surface brownish when dry, dull, the lower slightly paler;
neryes 9 or 10 on each side of the midrib, not prominent, the reticula-
tions lax, obscure; petioles about 3 mm long. Cymes axillary, solitary —
or several in each axil, short, rather congested, including the flowers — ot]
2 em long or less, the peduncles very short, the pedicles about 5 mm
long. Calyx-teeth acute, short. Corolla-tube cylindric, very slender, a

as

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 319

1 mm or less in diameter, the lobes spreading, 3 or 4 mm long. Fruit
unknown.

GUIMARAS, Nagaba, For. Bur. 304 Gammill, February, 1904 altitude about
80 m, in thickets &nd open places. V., Alibutbut.

A species with somewhat the appearance of Tabernaemontana LN es re Poir.,
but with quite different inflorescence. Readily recognzable by its very short

~ eymes and slender flowers.

Tabernaemontana puberula sp. nov.

Arbuscula vel arbor parva, ramulis petiolis foliis fructibusque plus
minus dense puberulis vel pubescentibus; foliis 6 ad 9 em longis, mem-
branaceis, acuminatis, oblongo-ellipticis; folliculis rubris, ovoideis vel
oblongis, 1 ad 2 cm longis, 3- ad 6-spermis.

A shrub or small tree, rather uniformly and softly puberulent or
pubescent. Branches slender, terete, grayish-brown, glabrous, the
branchlets puberulent. Leaves opposite, oblong-elliptic, puberulent on
both surfaces, especially beneath, 6 to 9 em long, 2.5 to 4 em wide,
membranaceous, the apex shortly acuminate, the base acute; nerves about
10 on each side of the midrib, distinct beneath, the reticulations lax;

6

petioles pubescent, 5 mm long or less. Follicles red, ovoid or oblong,
the ane acute, longitudinally 3-ribbed, 1 to 2 em long, 1 em wide or
less, the pericarp coriaceous, somewhat pubescent. Seeds 3 to 6.

Dace Province of Rizal, Morong, Bur. Sci. 1440 Ramos, August, 1906, in
thickets and open places; Malapadnabato, Merrill 2725, 2746, June, 1903.

A species with the general appearance of Tabernaemontana pandacaqui Poivr.,
but readily recognizable by its uniform soft pubescence.

VOACANGA Thou.

Voacanga globosa (Blanco) comb. nov.

Tabernaemontana globosa Blanco FI. Filip. (1837) 116, ed. 2 (1845) 83, ed. 3,
teeloo = Migiehike Inds Bot. 22 (1856) 424.

Orchipeda foetida Vidal Sinopsis Atlas (1883) 32, t. 66, f. D; F.-Vill. Nov.
App. (1880) 130, non Blume.

Voacanga cumingiana Rolfe in Journ. Linn. Soe. Bot. 21 (1884) 313; Vidal
Phan. Cuming. Philip. (1885) 126; Rev. Pl. Vase. Filip. (1886) 184; Merr. in
For. Bur. Bull. 1 (1903) 49; Philip. Journ. Sci. 1 (1906) Suppl. 117.

Luzon, Province of Benguet, Sablan, Elmer 6165, April, 1904: Province of
Bataan, Lamao River, Mount Mariveles, Werrill 3798, January, 1904, Whitford
1079, January, 1905, For. Bur. 684, 1521, 1755, 2331, Borden, May to December,
1904, For. Bur. 2282, 3014 Meyer, December, May, For. Bur. 5467 Curran, No-
vember, 1906, Williams 509, January, 1904: Province of Cavite, Mendez Nuiiez,
Bur, Sci. 1356 Mangubat, August, 1906: Province of Batangas, Lipa, Marave
16: Province of Laguna, Calauan, Cuming 476; Los Bafios, Elmer, April,
1906, Hallier, December, 1903; Pagsanjan, Merrill 2186, May 1903: Prov-
ince of Tayabas, Lucena, Merrill 2887, June, 1903; Atimonan, For. Bur. 6704
Kobbe, April, 1907; Gregory 108, August, 1904; Mount Banajao, Whitford 992,
October, 1904; Gumaca, Whitford 878, September, 1904; Binangonan, Whitford
829, September, 1904; Mauban, For. Bur. 10200 Curran, March, 1908. Mrnvoro,
Cuming 1500; Bongabong River, For. Bur. 4075 Merritt. Massatr, Merrill 3030.
Cresu, For. Bur. 6415 Espinosa, September, 1906. Lreyrr, Elmer 7103, January,

320 MERRILL.

1906. MINDANAO, Province of Surigao, Ahern 330; District of Davao, Williams
2941.

A widely distributed endemic species, for which the earliest valid specific
name is here adopted. The type of Voacanga cumingiana Rolfg was Cuming 1806
from Negros, and while it has somewhat smaller leaves than most of the specimens
cited above, I consider it to be identical with Blanco’s species.

WILLOUGHBYA Roxb.

Willoughbya luzoniensis sp. nov.

Frutex alte scandens, glabra, ecirrhosa; foliis oppositis, firmiter mem-
branaceis vel papyraceis, oblongo-ellipticis, nitidis, utrinque acuminatis,
usque ad 9 cm longis, nervis primariis utrinque circiter 12; fructibus
ovoideis vel ovoideo-ellipsoideis, glabris, nitidis, circiter 2.5 em longis;
seminibus circiter 1 cm longis.

A scandent shrub without tendrils, glabrous throughout, about 15 m
high. Branches slender, terete, grayish-brown, lenticellate, the branchlets
slender, somewhat olivaceous. Leaves opposite, oblong-elliptic, 6 to 9
cm long, 2 to 3.5 em wide, the apex rather prominently acuminate, the
base somewhat decurrent-acuminate, shining, firmly membranaceous or
papyraceous, pale or brownish when dry; primary nerves about 12 on
each side of the midrib, scarcely more distinct than are the secondary
ones and the reticulations ; petioles 5 mm long or less. Flowers unknown.
Berry ovoid or ovoid-elliptical, about 2.5 cm long, and 2 cm in diameter,
the pericarp crustaceous, pale, smooth, shining, the peduncle 1 to 1.5 em
long. Seeds irregular, 8 to 10 mm long, strongly and irregularly pitted,
in this character sponge-like.

Luzon, Province of Camarines, Caramoan, For. Bur. 12277 Curran, June, 1908.

The first species of the genus to be found in the Philippines, characterized by
its rather small fruits and its seeds which are sponge-like in appearance, but not
in texture,

CONVOLVULACE.
IPOMOEA Linn.

Ipomoea congesta R. Br. Prodr. (1810) 485; Choisy in DC. Prodr. 9 (1845)
369; Benth. Fl. Austr. 4 (1869) 417.

Luzon, Province of Cagayan, For. Bur. 16622 Curren, February, 1909. PAtMas

(southeast of Mindanao), Merrill 5353 October, 1906.
Queensland to Polynesia; new to the Philippines.

VERBENACEZ.

VITEX Linn.
Vitex pentaphylla sp. nov.
Arbor 10 ad 20 m alta; foliis 5-foliolatis, foliolis chartaceis vel mem-
branaceis, ovatis vel elliptico-ovatis, acuminatis, subtus plus minus pubes-
centibus, vix glandulosis; cymis axillaribus, solitariis, pedunculatis ; flori-

bus pubescentibus, calyce regulariter 5-dentato; fructibus nigris, glabris,
oblongo-ovoideis, circiter 1 em longis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. 321

A tree 10 to 20 m high. Branches brownish, glabrous, the young
parts sometimes slightly pubescent. Leaves alternate, 5-foliolate, the
common petiole 6 to 15 cm long, glabrous or somewhat pubescent: leaf-
lets chartaceous or membranaceous, ovate to elliptic-ovate, petiolulate, 7
to 16 cm long, 4 to 8 em wide, entire, base acute, apex acuminate, the
upper surface glabrous and shining, or the midrib and nerves slightly
pubescent, the lower surface slightly paler, more or less pubescent, or
ultimately glabrous or nearly so, not at all glandular; nerves 15 to 20 on
each side of the midrib, prominent, anastomosing; petiolules 5 to 15 mm
long, those of the lateral leaflets the shortest. Cymes axillary, solitary,
pedunculate, 10 to 15 em long, in anthesis more or less pubescent, in
fruit nearly glabrous. Flowers purplish, tinged with white, the bracteoles
very small. Calyx cup-shaped, somewhat pubescent, about 4 mm long,
regularly 5-toothed, the teeth less than 1 mm long, broadly ovate, acute
or acuminate. Corolla about 11 mm long, somewhat pubescent outside,
the tube 5 to 6 mm long, inflated upwards; middle lobe of the lower lip
narrowly obovoid, rounded, crisped, 6 to 7 mm long, densely villous on
the median portion inside, the lateral lobes and the upper lip 3 to 4 mm
long. Filaments shghtly villous below. Ovary ovoid, villous at the
apex ; style glabrous, 9mm long. Fruit black when dry, glabrous, oblong-
ovoid, rounded at the apex, about 1 cm long, the calyx persistent, accres-
cent, somewhat saucer-shaped, 6 to 7 mm in diameter.

Minpanao, District of Zamboanga, Siocon River, For. Bur. 9490 Whitford &
Hutchinson, February, 1908; For. Bur. 11245 Hutchinson, March, 1908. <A speci-
men collected by R. 8. Williams, Vo. 2949, in the District of Davao, Mindanao,
is probably referable here; it is with mature fruits, and is entirely glabrous.

This species is most closely allied to Vitex littoralis Dene.; but is at once
distinguishable by its five leaflets which are not at all glandular beneath. It is
distinguished from V. turczaninowii Merr. by its purplish, not yellow flowers,
non-glandular leaves and inflorescence, and other characters. It is locally known
as Malaun aso, and to the Moros as Calipapa aso.

ACANTHACE 2.

JUSTICIA Linn.
. Justicia tenuis sp. nov.

Herba annua, suberecta, diffusa, vel procumbens, usque ad 20 cm alta;
foliis membranaceis, ovatis, 1 ad 2 cm longis, acutis vel leviter acuminatis,
ramis ramulisque angulatis, plus minus hirsutis; inflorescentiis spicatis,
terminalibus axillaribusque, tenuibus; bracteis lanceolatis vel ovatis, acu-
minatis; calycis segmentis 5, anguste lanceolatis, acuminatis, hyalinis ;
corolla 2-lobata, lobis imbricatis ; seminibus 4, verrucosis.

An annual, suberect, diffuse, or procumbent, rather weak herb, 20 cm
high or less. Stems and branches slender, angled, more or less covered
with weak white hairs, the leaves also with a few similar ones.. Leaves
ovate, 1 to 2 cm long, membranaceous, acute or slightly acuminate, the
base broad, rounded; petioles very short. Spikes many, slender, short,

es MERRILL.

axillary, and terminating the branches, 1 to 3 em long, few-flowered, the
flowers distant or close, Flowers blue, the bracteoles paired, the lower pair
2.5 mm long, lanceolate, acuminate, shghtly hairy, the upper pair larger,
ovate to elliptic-ovate, abruptly acuminate, green with hyaline margins,
3 mm long, 2 mm wide. Sepals 5, free or nearly so, hyaline, lanceolate
or narrowly lanceolate, narrowed at both ends, apex slenderly acuminate,

more or less ciliate-hairy. Corolla about 5 mm long, blue, the tube short, =
1 mm long, glabrous, 2-lobed, lobes imbricate, the larger, outer lobe F
inclosing the other in bud, 4 mm long, obovate, 3-lobed, the terminal

lobe broad, retuse, the lateral ones ovate, blunt; the smaller, inner corolla-
lobe ovate, entire, 3 mm long, 2.5 mm wide, blunt-acuminate. Stamens
2; filaments about 2 mm long; anthers 2-celled, one cell above the other,
the lower cell spurred at the base, the spur rounded at its tip; pollen .
elliptic. Ovary glabrous, ovoid; style simple, 2.5 to 3 mm long. Fruit n
(immature) oblong, compressed, 3 mm long, 1.2 mm wide, hyaline, a
apiculate, containing four rounded, compressed, verrucose seeds.

Luzon, Province of Ilocos Norte, Bur. Sci. 2312 Mearns, February, 1907; Vin-
tar, For. Bur. 15509 Merritt & Darling; Pasuquin, For. Bur. 15536 Merritt &
Darling, November, 1908, in open grass-lands, ete., 100 to 400 m altitude.

Possibly most closely allied to Justicia procumbens Linn., which it resembles
in habit, but is much more slender than any forms of that species known to me,
differing in its very slender, continuous or interrupted spikes, five, instead of
four sepals, its blue corolla, and many other characters.

STROBILANTHES Blume.

Strobilanthes mearnsii sp. nov.

Herba erecta, omnibus partibus plus minus hirsutis; foliis oblongo-
lanceolatis, vel late ovato-lanceolatis, usque ad 15 cm longis, longe tenuiter
acuminatis, denticulatis, in paribus inaequalibus; inflorescentiis laxis,
paucifloris, pedunculis longis, hirsutis; floribus congestis, paucis; sepalis
circiter 6 mm longis, anguste lanceolatis, hirsutis; corolla alba, usque ad
2.7 em longa; staminibus 4; filamentis villosis. -

An erect, branched herb, more or less villous or hirsute throughout,
exceeding 60 cm in height. Branches terete, slender, dark-green or
brownish, hirsute. Leaves opposite, in unequal pairs, oblong-lanceolate
to broadly ovate-lanceolate, hirsute, membranaceous, the larger ones 8 to
15 cm long, 3 to 5 em wide, the smaller ones of each pair half as large
or smaller, acuminate at both ends, the apex slenderly so, dentate, the
upper surface with numerous small rhaphides; nerves about 8 on each f
side of the midrib, distinct; petioles densely hirsute, 1 to 2 em long.
Inflorescence axillary, 7 to 10 em long, few-flowered, hirsute, the flowers am
capitate at the ends of the-few branches, 3 to 5 flowers in a head, the
bracts foliaceous, deciduous. Calyx about 8 mm long, the lobes 6 mm
long, narrowly lanceolate. Corolla white, about 2.7 em long, the tube
slender below, widened above, slightly hirsute outside, subequally 5-lobed.
Stamens 4; filaments villous; anthers 1.3 mm long.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. O20

A
Luzon, Province of Benguet, Mount Tonglon, Mearns s. n., December, 1906;
Mount Ugo, For. Bur. 15920 Bacani, Bur, Sci. 5727 Ramos, December, 1908.
A species allied to Strobilanthes pluriformis Clarke, but distinguished from

that and all other known Philippine forms in being strongly hirsute in all

parts.
RUBIACE.

COPTOSAPELTA Korth.

C. flavescens Korth. Ned. Kruid. Arch. 2 (1851) 113; Mig. Fl. Ind. Bat. 2
(1857) 155; Hook. f. Fl. Brit. Ind. 3 (1880) 35; King & Gamble in Journ. As.
Soc. Beng. 72? (1903) 138.

Randia olaciformis Mery. in Philip. Journ. Sci. 3 (1908) Bot. 163.

Mindanao, Lake Lanao, Camp Keithley, Mrs. Clemens 1220.

Randia olaciformis Merr. must be reduced to Coptosapelta flavescens Iorth.,
a species extending from Burma to the Malay Peninsula, Sumatra, Java, and
Borneo, and the first representative of the genus to be found in the Philippines.
Hooker f. gives Webera macrophylla Roxb. as a synonym of the above, which, if
correctly reduced, would be the earliest specific name for the species. Roxburgh’s
description, however, does not seem to me to apply to Coptosapelta flavescens
Korth.

DAMNACANTHUS Gaertn.

Damnacanthus indicus Gaertn. f. Fruct. 3 (1805) 18, ¢. 182; DC. Prodr. 4
(1830) 478; Hook. f. Fl. Brit. Ind. 3 (1880) 158; Forbes & Hemsl. in Journ.
Linn. Soe. Bot. 23 (1888) 386.

Luzon, Province of Laguna, Mount Banajao, Por. Bur. 7856 Curran & Merritt,
November, 1907, Bur. Sci. 2432 Foxworthy, March, 1907, altitude 1,400-2,000 m.

The first representative of the genus for the Philippines, an interesting
addition to the continental element in the Philippine flora; eastern Himalayan
region to southern China and Japan.

GREENEA Wight & Arnott.

Greenea longiflora sp. nov.

Arbor parva, circiter 8 m alta, ramulis petiolis foliis cymisque hirsutis ;
foliis oblongo-ellipticis, papyraceis, 20 ad 30.cm longis, apice acutis vel
obscure acuminatis, nervis utrinque circiter 15; cymis axillaribus, subscor-
poideis, pauciramosis, usque ad 9 cm longis; floribus sessilibus, circiter
15 em longis, 4-meris. .

A small tree about 8 m high. Branches densely hirsute with long
brownish hairs. Leaves papyraceous, oblong-elliptic, 20 to 30 em long,
10 to 12 em wide, very sligthly shining, the lower surface somewhat paler
than the upper, both surfaces with numerous spreading or appressed
scattered hairs, those on the lower surface softer and paler than those
on the upper; nerves about 15 on each side of the midrib, distinct, the
reticulations rather lax; petioles densely hirsute, 2 to 4 cm long; stipules
membranaceous, caducous, oblong, about 4 em long and 1.5 cm wide,
rounded or obtuse at the apex, hirsute. Cymes axillary, peduncled, 9
em long or less, hirsute, the branches few. Flowers sessile, arranged

“ae eo ees ae
+ ~<a el 4 y k r 1)
) Ie. fee eee
P ‘oe

324 MERRILL. beat
along one side of the branches. Calyx-tube very short, the limb mem-
branaceous, 1 mm long, obscurely 4-toothed. Corolla-tube about 15 mm
long, cylindric, 8 mm in diameter, the outside with very few short
appressed hairs; lobes 4, spreading or recurved, imbricate, broadly elliptic. ;
ovate, acute or blunt, 4 to 4.5 mm long. Stamens 4, the filaments very
short, inserted above the middle of the tube; anthers linear, 6 mm lone
included. Style in the male flowers about 4 mm long, 4-cleft, obscurely j
pubescent.

MINDANAO, District of Zamboanga, Tibucuy, For. Bur. 9430 Whitford & a
Hutchinson, February, 1908, in forests at an altitude of about 20 m. ‘

The first species of the genus to be reported from the Philippines, allied to
Greenea corymbosa (Jack) K. Sch., of Burma and the Malay Peninsula, but
abundantly distinct.

GREENIOPSIS gen. nov.

Calycis tubus turbinatus; limbi lobi 5, subaequales, breves, sub-
rotundati, corolla breviter infundibularis, fauce villosa; limbi lobi 5,
breves, rotundati, in alabastro sinistrorsum contorto-imbricati. Stamina
5, fauci corollae inserta, inclusa, filamentis brevibus ; antherae dorso supra
basin affixae, ellipticae. Ovarium 2-loculare; stylus gracilis, lobis stig-
matis brevibus, crassis; ovula in loculis plurima, in placentis septo
longitudinaliter adnatis conferta. Capsula oblonga vel ovoidea, 2-locu-
laris, septicide 2-valvis, coriacea. Semina numerosissima, elongata
praecipue in nucleo reticulato-areolata. Arbores. Folia opposita, char-
tacea vel subcoriacea, oblongo-obovata vel elliptico-obovata, petiolata.
Stipulae amplae, deciduae, interpetiolares, acuminatae. Flores _parvi.
Inflorescentia terminalis, multiflora, cymis amplis, paniculatis, ramis
ramulisque patulis.

This proposed new genus is allied to Emmenopterys of southern China and to
Mussaendiopsis of the Malay Peninsula, Sumatra, and Borneo, but differs from
both in none of its calyx lobes being acerescent, and from the latter also in its
stamens inserted at the throat of the corolla-tube. It consists of two closely
allied Philippine species, the following, the type of the genus, which has been
previously referred to Greenea, species undetermined, and the plant described
by Mr. Elmer as Mussaendiopsis multiflora,

Greeniopsis philippininensis sp. nov.

Arbor circiter 10 m alta; foliis oblongo-obovatis vel elliptico-obovatis,
subchartaceis, nitidis, 20 ad 30 em longis, apice breviter acuminatis, basi
augustatis, decurrento-acuminatis, nervis utrinque 15 ad 18, prominen-
tibus, glabris vel junioribus subtus minus hirsutis ; paniculis terminalibus,
amplis, ramis ramulisque patentibus, circiter 20 cm longis; floribus
breviter pedicellatis, albis, 4 ad 5 mm longis; capsulis ellipsoideis, vel
ovoideis, 4 mm longis, extus griseo-pubescentibus, septicide 2-valvatis,
apice limbo persistente coronatis.

A tree about 10 m high. Branches terete, reddish-brown, rugose, the
younger branches reddish-brown, smooth, glabrous or sometimes hirsute.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. a25

Leaves opposite, oblong-obovate to elliptic-obovate, 20 to 30 cm long, 7
to 12 em wide, shining, glabrous, or when young somewhat hirsute
beneath, subchartaceous, the apex rather abruptly short-acuminate ; nerves
15 to 18 on each side of the midrib, prominent on both surfaces, the
reticulations subparallel; petioles 1.5 to 2 cm long, glabrous, or when
young somewhat hirsute; stipules interpetiolar, oblong-lanceolate, brown,
glabrous or sometimes hirsute, 1 to 3 cm long, deciduous, acuminate,
sometimes cleft. Panicles terminal, grayish-appressed-pubescent, ample,
the branches and branchlets spreading, many flowered, the flowers mostly
on the upper half of the branches alternate, not or rarely secund. Flowers
white, short-pedicelled. Calyx turbinate, the tube short, about 1 mm
long and nearly 2 mm wide, appressed-gray-pubescent outside, the limb
1.5 mm long, with 5 broad, short, rounded teeth which are slightly ciliate
on the margins, none of them accrescent. Corolla-tube about 3 mm
long, somewhat widened above, the limb with 5 rounded lobes about 1
mm long, in bud somewhat overlapping to the left, the throat densely
villous within below the anthers. Stamens 5, alternate with the corolla-
lobes, the filaments short, inserted on the throat of the tube; anthers
elliptic, attached at the back above the base, about 1 mm long, obtuse,
‘longitudinally 2-celled, included. Ovary 2-celled; ovules indefinite, on
the central placenta which is adnate to the disseptment ; style 5 mm long,
entire, crowned by the subcapitate or ellipsoid stigma which is about
1.2 mm long and somewhat cleft at the apex. Capsule coriaceous, ap-
pressed-gray-pubescent, ellipsoid or ovoid, about 4 mm long, 3 to 3.5
mm thick, crowned by the persistent calyx-lmb, septicidal, 2-valved,
2-celled; seeds numerous, elongated, rather pointed at both ends, some-
times angular, 1 mm long, externally reticulate-areolate.

Samar, Borongan, Merrill 5216 (type of the genus and species), October, 1906,
in flower and fruit, growing along the border of a Nipa swamp; without definite
locality, Cuming 1713, in fruit, in Herb. Kew. sub Greenea. Luzon, Province of
Albay, Cabit, Bur. Sci. 6402 Robinson, August 31, 1908, near hot springs along
the seashore, with flowers and fruits; Manito, Por. Bur. 10580 Curran, June, 1908,
along the seashore, in fruit.

In young specimens, such as the type, the branchlets, inflorescence, and the
midrib and nerves on the under surface of the leaves are supplied with numerous

long, brownish hairs, but these are deciduous, the mature specimens being glabrous
or subglabrous with the exception of the more or less pubescent inflorescence.

Greeniopsis multiflora (Elmer) comb. nov.

Mussaendiopsis multiflora Elm. Leafl. Philip. Bot. 1 (1906) 14.

This differs from the above in having leaves with from 9 to 12 pairs of lateral
nerves only, the fruits relatively longer and narrower (5 mm long, 2-2.5 mm
thick), and which are secund on the ultimate branchlets of the infrutesence.
In habit and general appearance it strongly resembles the preceding species, but
can be distinguished by the above characters. The flowers are unknown.

Dinacat, Ahern 490. (type) N. v., Hamagos.

326 MERRILL. Jp

LITOSANTHES Blume.

L. biflora Blume Cat. Gew. Buitenzorg (1823) 21; Bijdr. (1826) 994;
Fl. Ind. Bat. 2 (1857) 314; DC. Prodr. 4 (1830) 465.
Luzon, Province of Sorsogon, Adlumay Hills, For. Bur. 12383 Curran, Fe a
1908. Muinporo, Mount Halcon, Merrill 6172, November, 1906; Mount Sablanga, —
For. Bur. 11002 Merritt, March, 1908; Mount Teluti, For. Bur. 11478 Sern, c
May, 1908. Nercros, Elmer. >

The first representative of this small genus to be found in the Philippines; oe
Java and Celebes.

Two species have been described from New Guinea.

NAUCLEA Linn.

Nauclea jagori sp. nov.

Arbor vel arbuscula glabra; foliis lanceolatis vel anguste lanceolatis,
longe sensim acuminatis, 8 ad 20 em longis, 9 ad 18 mm latis, basi
angustatis, subcoriaceis, nitidis, nervis utrinque circiter 15; capitulis
solitariis, circiter 3 cm diametro stylis exclusis; corolla glabra, 1 cm
longa.

A shrub or tree, glabrous. Branches grayish-brown, terete, the branch-
lets somewhat compressed. Leaves lanceolate or narrowly lanceolate,
8 to 20 cm long, 9 to 18 mm wide, subcoriaceous, shining, brownish when
dry, the base narrowed, acute, the apex long and gradually acuminate; —
nerves about 15 on each side of the midrib, not prominent, obscurely
anastomosing, the reticulations indistinct; petioles 3 to 7 mm long;
stipules oblong, obtuse, 12 to 15 mm long, 3 mm wide, slightly hirsute
below, caducous. Heads solitary, the peduncles about 3 cm long, the |
bracts, if any, caducous, the heads, excluding the long exserted styles,
about 3 cm in diameter. Corolla 1 em long, glabrous, widened upward,
the lobes oblong, obtuse, 2.5 mm long, 1mm wide. Anthers 1.7 mm long.
Style about 1.5 cm long; stigma about 2.5 mm long, swollen in the
middle. Cocci 8 mm long; seeds, including the narrow, acuminate wings,
4 mm long, slightly pubescent.

Samar, /’. Jagor 987, in Herb. Berol.
A very characteristic species, at once distinguished from most hitherto described
ones in the genus by its very narrow leaves. It is apparently allied to Nauclea
angustifolia Havil., of Borneo, and N. chalmersii F. Muell., of New Guinea, both
of which have very narrow leaves, but the present species differs from these in
its more numerously nerved leaves and larger flowers.

PLECTRONIA Linn.

Plectronia megacarpa sp. nov.

Arbor glabra circiter 15 m alta; foliis coriaceis, elliptico-ovatis vel
obovato-ellipticis, circiter 20 cm longis, brunneis, nitidis, apice rotundatis
vel late obscure acuminatis, basi cuneatis, neryis utrinque circiter Be
prominentibus, nervulis obscuris; fructibus axillaribus, pedicellatis, obo-

voideis, compressis vel trigonis, 2- vel 3-locellatis, 15 em longis, apice
truncatis vel retusis, basi acutis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VII. . 32

~I

A glabrous tree about 15 m high. Branches brownish, terete or
slightly compressed. Leaves coriaceous, elliptic-ovate or obovate-elliptic,
about 20 cm long, 9 to 11 em wide, the upper surface brown, shining,
the lower paler, dull, margins somewhat recurved, apex broad, rounded,
or broadly and obscurely blunt-acuminate, base cuneate, sometimes slightly
acuminate; nerves about 8’on each side of the midrib, prominent, the
nervules and reticulations obscure; petioles stout, 2 to 2.5 em long;
stipules deciduous, ovate, acuminate, less than 5 mm long. Flowers
unknown. Infrutescence axillary, umbellate, the peduncle stout, about 5
mm long, the pedicels 1.5 to 3 em long. Fruit ‘yellow when fresh, 1.5
em long, obovoid, trigonous and three-celled, or somewhat compressed and
two-celled, nearly 1.5 cm thick above the middle, the apex truncate or
retuse, the base acute.

Luzon, Province of Zambales, near Santa Cruz, altitude 100 m, For. Bur. 8232
Curran & Merritt, December, 1907.

A species apparently allied to Plectronia glabra (Blume) Kurz, but with

different fruits and leaves.
TIMONIUS Rumph.

Timonius appendiculatus sp. nov.

Arbor parva, circiter 8 m alta, glabra; foliis coriaceis oblongis vel
anguste elliptico-oblongis, usque ad 10 cm longis, basi plus minus decur-
rento-acuminatis, apice acutis vel obtusis, nervis lateralibus obscuris;
cymis axillaribus, pauciramosis, foliis multo brevioribus; floribus 4-meris,
unilateraliter spicatis, lobis corollae recurvatis, 3-appendiculatis; drupis
globosis, obscure 12-sulcatis, polypyrenis.

A small tree glabrous throughout. Branches reddish-brown or grayish,
terete, wrinkled when dry. Leaves coriaceous, oblong to narrowly ellip-
tic-oblong, rather pale when dry, slightly shining, the apex acute or
blunt, the base somewhat decurrent-acuminate, 6 to 10 cm long, 2 to 3
em wide; lateral nerves few, obscure, and, with the reticulations, nearly
obsolete; petioles less than 1 cm long; stipules annular, truncate, very
short. Cymes in the upper axils, about 3 em long, peduncled, with
usually but two branches, the flowers spicately arranged on one side of
the branches only, sessile, articulated with the rachis. Calyx cupular,
about 3 mm long, 1.5 mm wide, truncate. Corolla about 9 mm long,
tubular, the lobes 4, reflexed, narrowly ovate, acute, about 2 mm long,
and with three retrorse appendages near the apex, the two lateral ones
about 1 mm long, the terminal one shorter. Filaments short, inserted
at the throat or just below it; anthers linear, nearly 5 mm long, their
tips slightly exserted; style about 3 mm long. Fruit globose, shining,
obscurely about 12-suleate, crowned by the truncate calyx, about 5 mm in
diameter, about 12-celled, and containing about 25 pyrenes.

Luzon, Province of Rizal, Bosoboso, Bur. Sci. 2164, 2651 Ramos, January, May,
1907, the former with staminate flowers, the latter with fruit. M1rnporo, Mount
Agong, For. Bur, 9867 Merritt.

ne Ps é »
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die ta ie , ote ) pas a
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328 MERRILL. gi eee

A very characteristic species, differing from typical Timonius in its appen¢
corolla-lobes, but in other characters typical, readily recognized by the above
character, its few-branched cymes, 4-merous, sessile flowers in unilateral spikes,
and its nearly nerveless leaves.

XANTHOPHYTUM Reinw.

Xanthophytum fruticulosum Reinw. ex Blume Bidjr. (1826) 839; Mig. Fl. —
Ind. Bat. 2 (1857) 175. $ ee
Metabolus ferrugineus DC. Prodr. 4 (1830) 436. 5"
Sclerococcus Bart. 1. ¢., as syn.
Xanthophytum villarii Vidal Rev. Pl. Vase. Filip. (1886) 150.
Lasianthus pilosus F.-Vill. Noy. App. (1880) 112, non Wight.
Luzon, without locality, Haenke in Herb. Prague, type of Metabolus ferru-
gineus DC.; Province of Rizal, San Mateo, Vidal 392, in Herb. Kew, type of
X, villarii Vidal.
I have examined both of the above specimens, and can see no reason for |
distinguishing one from the other, or either from Reinwardt’s species. Vidal’s
specimen has smaller leaves than Haenke’s, but in all essential characters they
appear to be the same, and after a cursory examination of Javan specimens and
a careful comparison of the Philippine material with the descriptions of Y.
fruticulosum I do not hesitate to reduce both DeCandolle’s and Vidal’s species
to Reinwardt’s. Miquel |. ¢., had already reduced Metabolus ferrugineus to
Nanthophytum fruticulosum, without however giving the range of the species as
extending to the Philippines. An interesting species known only from Java,
Borneo, and Luzon.

CAPRIFOLIACEZ.
VIBURNUM Linn.

Viburnum floribundum sp. nov. § Luviburnum, Opulus.

Frutex circiter 2 m alta, subglabra vel parce pubescenti; foliis ovatis,
usque ad 7 em longis, chartaceis, nitidis, supra glabris, subtus in axillis
barbatis, apice acutis vel acuminatis, basi inequilateraliter rotundatis,
margine serrato-dentatis, nervis utrinque 5, prominentibus; cymis termi-
nalibus, umbellato-paniculatis, dense multifloris; corolla rotata.

A shrub about 2m high. Branches and branchlets terete, grayish- or |
reddish-brown, rather stout, glabrous, the branchlets minutely puberulent
and with few scattered long hairs. Leaves ovate, 4 to 7 em long, 2 to 4
em wide, chartaceous, shining on both surfaces, the apex acute or acumi-
nate, the base somewhat narrowed, rounded, inequilateral, the margins i *
especially in the upper half, rather strongly serrate-dentate, the upper
surface glabrous, or the midrib sometimes slightly puberulent, the lower
surface glabrous except the midrib and primary nerves which are often
slightly pubescent, and the axils of the primary nerves which are barbate; _
nerves 5 on each side of the midrib, ascending, very prominent, the —
reticulations distinct; petioles about 5 mm long, somewhat pubescent. —
Inflorescence ‘eeaciaeate arp $i umbellate, intermixed frequently with: ;
small leaves, rather dense, 5 to 7 em in diameter, very many flowered, —

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIL. 329

the rachis, branches and pedicels puberulent and with intermixed longer
hairs. Flowers small. . Calyx-lobes narrowly ovate, less than 0.5 mm
long, pubescent. Corolla rotate, 1.8 mm long, the tube short, the lobes
elliptic, rounded. Ovary depressed-globose, glabrous. Fruit ovate, com-
pressed, 5 mm long, 4 mm wide, not ribbed, or with an obscure rib on
one side.

Luzon, Province of Abra, Mount Paraga, Bur. Sci. T074 Ramos, February,
1909, the altitude given as about 600 m.

A species manifestly allied to Viburnum luzonicum Rolfe, but distinguished
from that species by its nearly glabrous leaves which are less acuminate, more
strongly toothed, rounded and inequilateral at the base, as well as by its dense
inflorescence. :

Viburnum glaberrimum sp. nov. § Huviburnum, Lantana ?

Arbor parva, glabra, ramis ramulisque crassis, valde lenticellatis; foliis
coriaceis, glabris, nitidis, obovatis, integris, apice rotundatis, obtusis, vel
late obtuse acuminatis, basi obtusis vel leviter decurrenti-acuminatis,
nervis utrinque 6 ad 8, prominentibus; inflorescentiis terminalibus, umbe-
llato-corymbosis ; corollae tubo cylindraceo, lobis erectis.

A small tree, glabrous throughout. Branches and branchlets stout,
terete, dark-reddish-brown, strongly lenticellate. Leaves coriaceous, shin-
ing, obovate, entire, the apex broad, rounded, obtuse, or sometimes broadly
and obtusely acuminate, the base obtuse or slightly decurrent-acuminate,
8 to 14 cm long, 6 to 8 cm wide; nerves 6 to 8 on each side of the midrib,
very prominent beneath, the reticulations not distinct; petioles stout, 3
to 4 cm long. Inflorescence terminal, about 5 peduncles at the tip of
each branch, forming an umbellate corymb, the peduncles 4 to 5 cm long.
Calyx-tube short, the teeth very small, rounded, about 0.4 mm long.
Corolla cyclindric, 5 mm long, thick, the tube 3 mm long, the erect,
elliptic, obtuse lobes 2 mm long. Filaments about 5 mm long; anthers
exserted, 2 mm in length. Ovary 1-celled. Fruit (immature) elliptic,
compressed.

Luzon, Province of Pangasinan, near Imogen, For. Bur. 15848 Merritt, Decem-
ber, 1908, in forests 300 to 450 m altitude.

A species manifestly allied to Viburnum coriacewm Blume of the Indo-Malayan
region, but at the same time very distinct, especially in its vegetative characters.

CUCURBITACE.
GYNOSTEMMA Blume.

Gynostemma simplicifolia Blume Bijdr. (1825) 23; Cogn. in DC. Monog,
Phan. 3 (1881) 915.
MINDANAO, District of Zamboanga, Sax River, Williams 2119, February, 1905.
New to the Philippines and the fourth species of the genus for the Archipelago;
previously known only from: Java.
85754——-9

Ca MERRILL. a

TRICHOSANTHES Linn.

Trichosanthes villosa Blume Bijdr. (1826) 934; Cogn. in DC. Monog. Phang
3 (1881) 366. ee:
Luzon, Province of Rizal, Bosoboso, Bur. Sci. 1027 Ramos, July, 1906, with
staminate flowers. MINDANAO, District of Davao, Todaya, Copeland 1301, April, ‘
1904, with mature fruit.

Previously known only from Java.
MELOTHRIA Linn.

Melothria scaberrima sq. nov. § Humelothria.

Planta monoica, floribus masculinis solitariis vel binis, breviter pedi-
cellatis; foliis lanceolatis vel oblongo-lanceolatis, chartaceis, apice acu-
minatis, basi hastatis, margine integris vel leviter denticulatis, utrinque
scaberrimis, subtus ad nervos hispido-aculeatis, usque ad 9 cm longis;
fructibus ovoideis vel oblongo-ovoideis, glabris, basi acutis, apice acu-
minato-rostratis.

Monoecious. Branches slender, striate-sulcate, glabrous or nearly so.
Leaves lanceolate or oblong-lanceolate, 4 to 9 cm long, 1 to 3.5 em wide,
chartaceous, the base hastate, truncate, the basal lobes short, spreading
or slightly reflexed, the margins aculeate, entire, or in the lower part
denticulate, gradually narrowed upward to the sharply acuminate apex,
green or olivaceous when dry, of about the same color on both surfaces,
the upper surface strongly white-glandular-scabrous, the midrib setose,
the lower surface less scabrid than the upper, the midrib and nerves
aculeate-hispid ; basal nerves 5 to 7, the inner longer pair reaching at
least to the middle of the leaf, ascending, anastomosing with the primary
lateral nerves, the reticulations very lax, obscure; petioles 0.5 to 2 cm
long, tendrils simple, leaf-opposed or from the axils below the leaves.
Flowers white, solitary or two in an axil: staminate short-pedicelled,
solitary or paired; calyx-lobes linear, 1 mm long; corolla lobes ovate,
shorter; stamens 3, subsessile: pistillate flowers long-pedicelled; calyx
teeth minute; corolla-lobes ovate, acuminate 3.5 mm long; stigma 2 mm
in diameter. Fruit ovoid or oblong-ovoid, terete, glabrous, about 3 cm
long, the base acute, the apex strongly acuminate-rostrate ; seeds narrowly
elliptic, compressed, 5 mm long, the base appendiculate-winged.

Luzon, Province of Benguet, Elmer 5862; Williams 1055; Merrill 4657; Topp-
ing 83; Pond s. n.; For. Bur. 15979 Bacani.

A species widely distributed in the pine region of Benguet Province, well
characterized by its lanceolate, hastate, very scabrous leaves, and ovoid or oblong-

ovoid rostrate. fruits. Probably most closely allied to Melothria zeylanica Clarke,
but very different from that species.

PREVIOUS PUBLICATIONS—Continued.
eg Bureau of Science—Annual Reports.

201. Fifth Annual Report of the Director of the Bureau of Science for the Year Ending
August 1, 1906.

202. Sixth Annual Report of the Director of the Bureau of Science for the Year Ending

August 1, 1907. .
203. Seventh Annual Report of the Director of the Bureau of Science for the Year Ending
August 1, 1908.

Philippine Museum, ’
(Now Section of Ornithology.)

* No. 1, 1903.—On Birds from Luzon, Mindoro, Masbate, Ticao, Cuyo, Culion, Caga-
yan Sulu and Palawan. By Richard C. McGregor, :
* No, 2, 1903.—List of Bird Skins Offered in Exchange.
43. No, 3, 1904.—Birds from Benguet Province, Luzon, and from the Islands of Lubang,
Mindoro, Cuyo, and Cagayancillo. By Richard C. McGregor.
*No. 4, 1904.—The Birds of Calayan and Fuga, Babuyan Group. By Richard C.
McGregor.

Mining Bureau.
(Now Division of Mines.)

45. 1890.—Descripcién fisica, geol6gica y minera en bosquejo de ia Isla de Panay por
D. Enrique Abella y Casariego, Inspector General de Minas del Archipiélago.

* 1890.—Memoria descriptiva de los manantiales minero-medicinales de la Isla de
Luzon, estudiados por la comisi6n compuesta de los Sefiores D. José Centeno, Inge-
niero de Minas y Vocal Presidente, D. Anacleto del Rosario y Sales, Vocal Far-
macéutico, y D. José de Vera y G6mez, Vocal Médico.

47. 1893.—Estudio descriptivo de algunos manantiales minerales de Filipinas ejecutado
por la comisién formada por D. Enrique Abella y Casariego, Inspector General de
Minas, D. José de Vera y Gémez, Médico, y D. Anacleto del Rosario y Sales, Far-
macéutico ; precedido de un prélogo escrito por el Excmo. Sr. D. Angel de Avilés,
Director General de Administracién Civil. ,

48. 1895.—Terremotos experimentados en la Isla de Luzén durante los meses de Marzo y
Abril de 1892, especialmente desastrosos en Pangasinan, Unién y Benguet. Estudio
ejecutado por’ D. Enrique Abella y Casariego, Inspector General d= Minas del
Archipiélago.

49. 1901.—The Coal Measures of the Philippines. Charles H. FPurritt,

50. 1902.—Abstract of the Mining Laws (in force in the Philippines, 1902). Charles H.
Burritt.

51. 1902., Bulletin No. 1—Platinum and Associated Rare Metals in Placer Formations,
H. D. McCaskey, B: 8.

52. 1993.—Report of the Chief of the Mining Bureau of the Philippine Islands. Charles
H. Burritt.

53. 1903, Bulletin No. 2.—Complete List of Spanish Mining Claims Recorded in the
Mining Bureau. Charles H. Burritt. ; -

54. 1903, Bulletin No. 3.—Report on a Geological Reconnoissance of the Iron Region of
Angat, Bulacan. H. D. McCaskey, B. S.

55. 1904.—Fifth Annual Report of the Mining Bureau. H. D. McCaskey.

56. 1905.—Sixth Annual Report of the Chief of the Mining Bureau. H. D. McCaskey.

57. 1905, Bulletin No, 4.—A Preliminary Reconnoissance of the Mancayan-Suyoc Mineral
Region, Lepanto, P. I. A. J. Eveland, Geologist.

58, 1905, Bulletin No. 5.—The Coal Deposits of Batan Island. Warren D. Smith, B. 8,
M. A., Geologist.

Division of Mines.

301. 1908.—The Mineral Resources of the Philippine Islands, with a Statement of the
Production of Commercial Mineral Products during the year 1907, issued by
Warren D. Smith, Chief of the Division of Mines.

Ethnological Survey,
(Now Division of Ethnology.)
401. Vol. I, 1905.—The Bontoc Igorot, by Albert Ernest Jenks. Paper, ®2; half morocco,
F3

402. Vol. II, Part 1, 1904.—Negritos of Zambales, by William Allen Reed. Paper, #0.50;
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403. Vol. II, Part 2 and Part 3, 1905.—The Nabaloi Dialect, by Otto Scheerer. The Bataks
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404. Vol. IIIT, 1904.—Relaciones Agustinianas de las razas. del Norte de Luzon, by Perez.
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405. Vol. IV, Part 1, 1905.—Studies in Moro History, Law, and Religion, by Najeeb M.
Saleeby. Paper, #0.50; half morocco, #1.50.

Division of Ethnology.

406. Vol. IV, Part 2, 1908.—History of Sulu, by Najeeb M. Saleeby. Paper, P1.50.

407. Vol. V, Part 1-and Part 2, 1908.—The Batan Dialect as a Member of the Philippine
Group of Languages, by Otto Scheerer. “F’’ and ‘“‘V” in Philippine Languages, by
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408. Vol. V, Part 3.—A Vocabulary of the Igorot Language as spoken by. the Bontok
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409. Directions for Ethnographic Observations and Collections, 1908. For free dis-
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C. BOTANY
Vou. V AUGUST, 1910 No. 3

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII.

By E. D. MERRILL.

(From the Botanical Section of the Biological Laboratory, Bureau of Science,
Manila, P. I.)

The following paper is largely composed of the descriptions of about
100 new species of Philippine plants, that have been worked out from

time to time during the past year. In the paper will also be found the

descriptions of four proposed new genera, Astrocalyx and Cephalomedi-
nilla of the Melastomataceae, Curraniodendron of the Saxifragaceae, and
Pygmaeopremna of the Verbenaceae. A number of species previously de-
scribed by various authors are here recorded from the Philippines for
the first time, while several previously considered Philippine forms are

‘ admitted for the purpose of discussion, additional data being available.

Following the rules of priority in nomenclature, a few new combinations
have been made.

GRAMINEZ.
ANDROPOGON Linn.

Andropogon citratus DC. Cat. Hort. Monsp. (1813) 78.

Cymbopogon citratus Stapf in Kew Bull. (1906) 322, 357, cum lamina.

Andropogon schoenanthus Blanco Fl. Filip, (1837) 39, ed. 2 (1845) 27, ed. 3,
1 (1877) 50; Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 339, non Linn.

This species is commonly cultivated in the Philippines, although not on a
commercial scale. I have never seen it in flower, but chemical analysis of the
oil extracted from it shows the species to be Andropogon citratus DC., and not
A. schoenanthus Linn.

95948 ; 167

168 MERRILL.

DIGITARIA Scop.

Digitaria ciliaris (Retz.) Pers. Syn. 1 (1805) 85.

Panicum ciliare Retz. Obs. 4 (1786) 16.

Paspalum sangwinale var. ciliare Hook. f. Fl. Brit. Ind. 7 (1897) 15.

Luzon, Province of Ilocos Norte, Bambam, Bur. Sci. 7661 Ramos, March, 1909:
Province of Rizal, Phil. Pl. 147 Merrill.

Not previously reported from the Philippines; widely distributed in the Indo-
Malayan region, extending to Polynesia. .

PANICUM Linn.

Panicum psilopodium Trin. Gram. Panic. (1826) 217; Hook. f. Fl. Brit.
Ind. 7 (1897) 46.

Luzon, Province of Benguet, Baguio, Williams 1182, Elmer 6589.

This species has previously been reported from the Philippines, but apparently
on a wrong identification. The specimens here referred to Trinius’ species are
considerably smaller than the typical form.

India to Ceylon, Burma, and Malacca.

ISACHNE R. Br.

Isachne incrassata (Hack.) comb. nov.

Isachne debilis Rendle var. incrassata Hack. in Philip. Journ. Sci. 1 (1906)
Suppl. 268; Merr. 1. c. 350.

Tufted, erect, rather stiff, the culms simple or shghtly branched, their
lower nodes sometimes decumbent, glabrous. Leaves rigid, mostly spread-
ing, lanceolate, sharply acuminate, 2.5 to 5 em long, 2 to 4.5 mm wide,
the lower surface puberulent or pubescent, the nerves obscure, the upper
surface glabrous, minutely scabrid, the nerves distinct, close, about 20,
margins thickened and cartilaginous, scabrid; sheaths subglabrous, or
with few to rather many, long, white hairs, especially on the margins.
Panicles exserted, peduncled, subpyramidal, 5 cm long or less, the
branches spreading or ascending, slender, glabrous, 2 em long or less, each
branch with 8 spikelets or less, the pedicels 1 to 3.5 mm long, glabrous.
Spikelets oblong-obovoid, usually dark-purple, sometimes pale, 1.2 to 1.5
mm long. Empty glumes glabrous, obscurely 5- to 7-nerved. First flow-
ering glume elliptic-oblong, rounded, glabrous, sessile, inclosing a ¢
flower, the second minutely pedicelled, similar to the first but slightly
smaller, inclosing a @ flower.

The type of Doctor Hackel’s variety was a rather poor specimen collected in
Mindanao, Mount Apo, DeVore & Hoover 358. Better material has been recently
collected in Negros, Merrill 6977, Canlaon Voleano, on ledges in the Nahalin
River, altitude about 1,260 m, and after studying this specimen I have concluded

that the form is not closely allied to Jsachne debilis Rendle, but that it is worthy
of specific rank.

Isachne micrantha sp. nov.

Annua, debilis, parva, vix 10 cm alta, caulibus tenuibus, ramosis, gla-
bris; foliis oblongis vel oblongo-lanceolatis, circiter 1 cm longis, leviter
pilosis, vaginis quam internodia breyioribus ; paniculis depauperatis, stric-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 169

tis, compressis, paucifloris ; spiculis obovoideis, 1 ad 1.2 mm longis, glumis
sterilibus obscurissime 5-nerviis, subglabris, fertilibus densissime et brev-
iter pubescentibus.

An annual, weak, ascending, loosely tufted plant, 10 cm or less in
height. Stems very slender, glabrous, branched, the lower parts decum-
bent and often rooting at the nodes. Leaves oblong to oblong-lanceolate,
about 1 em long, often shorter, rarely 1.5 cm in length, 2 to 3,5 mm
wide, the base rather broad, apex acuminate, both surfaces with scattered,
white, rather soft hairs about 1 mm long, the nerves about 10, the margins
minutely scabrid; sheaths shorter than the internodes, less than 1 cm
long, with few, scattered, long, white hairs, especially on the margins ;
ligule of few white hairs. Panicles strict, narrow, glabrous, less than 1
cm long, the branches few, 4 mm long or less, each with from 1 to 3 or 4
spikelets, apparently never spreading, the pedicels 1 to 2 mm long, gla-
brous: Spikelets obovoid, pale or slightly purplish, 1 to 1.2 mm long,
the empty glumes subglabrous, or at least with only a few scattered hairs
in the upper part, very obscurely 5-nerved or nerveless. Flowering
glumes densely and apparently softly pubescent with short hairs, 1 mm
long, the second very minutely pedicelled; paleas slightly pubescent.

Luzon, Province of Cagayan, Mount Cueva, For. Bur. 16837, 16841 (type)
Curran, March 9, 1909, altitude about 300 m.

A species well characterized by its small size, small leaves, contracted panicles,
and densely pubescent flowering glumes. It is probably more closely allied to
Tsachne debilis Rendle, of Formosa, than any other Philippine form at present
known, but seems to be distinguished by its contracted panicles, smaller leaves and

smaller spikelets. Among the Philippine species it is perhaps most closely allied
to Lsachne myosotis Nees.

Isachne vulcanica sp. nov.

Densissime caespitosa, perennis, rigida; culmis inferne ramosis, rigi-
dis, vix 10 cm altis; foliis rigidis, lanceolatis, acuminatis, 1 ad 2.5 cm
longis, 2 ad 5 mm latis, scabridis, margine incrassatis, vaginis quam
internodia longioribus; paniculis oblongis, congestis, 1 ad 2 cm longis,
usque ad 1 cm diametro; spiculis atropurpureis vel rariter pallidis, densis-
sime dispositis, 2 mm longis; glumis exterioribus 7-nerviis, subglabris,
fertilibus obscure sparseque pubescentibus.

A densely caespitose, rigid perennial, forming tufts a few centimeters
in diameter or mats which are often 0.5 m in diameter. Culms rigid,
much branched below, not exceeding 10 cm in height, often much shorter,
the internodes short. Leaves rigid, lanceolate, acuminate, 1 to 2.5 em
long, 2 to 5 mm wide, the margins thickened, cartilaginous, scabrid, the
upper surface prominently ribbed with about 18 nerves, scabrid, beneath
slightly hispidulous; sheaths exceeding the internodes, imbricate, with
scattered, long white hairs, the marginal hairs more numerous; ligule
of few stiff, white hairs 2 to 3 mm long. Panicles slightly or not at
all exserted, oblong, very dense, purple, 1 to 2 cm long, 0.5 to 1 em in

170 MERRILL.

diameter, the lower branches often 1 cm long, closely appressed, the
rachis and branches glabrous, the pedicels 0.3 to 1.5 mm long. Spikelets
numerous, ovate to oblong-oyate, 2 mm long, usually dark-purple, some-
times greenish. Empty glumes subequal, 7-nerved, glabrous except for
few long hairs on the apical parts. Third glume elliptic-oblong, 1.8
mm long, with very few, obscure, short hairs, the margins obscurely
ciliate, the palea as long as the glume, inclosing a ¢ flower. Fourth
glume similar to the third, slightly shorter, containing a ? flower.

NecGros, Canlaon Volcano, Merrill 6975, April, 1910.

Abundant in the old crater, on bare slopes and on debris washed down from
the new cone, forming dense tufts or mats, altitude 1,800 to 2,100 m. A species
well characterized by its dense panicles, short, rigid, densely caespitose habit,
rigid, crowded leaves, and dark-purple spikelets.

MISCANTHUS Anderss.

Miscanthus depauperatus sp. noy.

Culmis erectis vel suberectis, usque ad 60 cm altis, densissime caes-
pitosis, vix 3 mm diametro; foliis 3 ad 6 mm latis, margine scabridis;
spiculis purpureis, 6 mm longis; paniculis laxis, racemis paucis, race-
moso-dispositis, solitariis vel inferioribus binis, usque ad 15 cm longis.

A densely caespitose perennial, forming tufts up to 1 m in diameter,
the culms erect or ascending, usually about 40 em high, often shorter,
rarely 60 cm in height, simple or very rarely branched, terete, 2 to 2.5
mm in diameter. Leaves numerous, those of the culm up to 30 em long,
6 mm wide, the basal ones much shorter and usually narrower, scabrid
on the margins, long- and slenderly acuminate; sheaths exceeding the
internodes, the lower ones usually purplish; ligules broad, about 2 mm
long, margins ciliate. Panicles exserted or not, glabrous, the rachis and
branches angled, purple, minutely scabrid on the angles, the rachis always
less than 5 em in length, the branches rather slender, erect or somewhat
spreading, solitary, or the lower ones in pairs, 3 or 4 to about 9 in
number, 4+ to 15 cm in length. Spikelets purple, lanceolate, acuminate,
about 6 mm long, the shorter pedicels about 2 mm long, the longer ones
5 to 6 mm long, the involucral hairs slender, about as long as the spike-
lets, usually purplish, unequal in length, numerous. First two glumes
lanceolate, sharply acuminate, equal, about 6 mm long, when spread 2.3
mm wide, purple, shining, glabrous, or the margins with very few ciliate
hairs, the first obscurely 5-nerved, the second obscurely 3-nerved. Third
glume oblong-lanceolate, hyaline, 5 mm long, 2 mm wide, acuminate,
l-nerved, margins slightly ciliate, empty. Fourth glume hyaline, lanceo-
late, acuminate, 4 mm long, 1 mm wide, apex cleft into two slender
teeth and bearing between them a slender, scabrid, twisted awn about 1
em in length. Palea ovate, hyaline, 1 mm long. Lodicules truneate,
about 0.8 mm long. Anthers 3, 2.5 to 3 mm long.

Neoros, Canlaon Voleano, common in open places in the old crater, altitude

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. sg |

about 1,800 m, ascending the new cone to an altitude of about 2,000 m, Merrill,
April 12, 1910. :

This species is manifestly allied to Miscanthus sinensis Andr., differing in
its densely tufted habit of growth, very much smaller size, narrower leaves, rather
lax, depauperate panicle, solitary or at most paired panicle-branches, and larger
spikelets.

SPOROBOLUS R. Br.

Sporobolus virginicus (L.) Kunth Rev. Gram. (1829) 67; Enum. 1 (1833)
210; Hook. f. Fl. Brit. Ind. 7 (1897) 247.

Agrostis virginica Linn. Sp. Pl. (1753) 63. :

Luzon, Province of Cagayan, Bur. Sci. 7881 Ramos, April, 1909; also collected
by Loher at Navotas, Province of Rizal, nos. 1785, 1786 in Herb. Kew; Manila,
Merrill, June, 1910.

Not previously reported from the Philippines; widely distributed in tem-
perate and tropical parts of the world.

CYPERACEZ..
CLADIUM Schrad.

Cladium philippinense sp. nov. § Hucladium.

Dense caespitosum, culmis gracilibus, teretibus, usque ad 2 m altis, 2
ad 3 mm diametro; foliis radicalibus nullis, caulinis 2 vel 3, brevibus,
verticaliter compressis, 1 ad 5 cm longis, vix 3 mm latis; inflorescentis
laxis, 10 ad 15 cm longis, “zigzag”; spiculis brunneis, circiter 2 mm
longis; fructibus ellipsoideis, teretibus, admodum nitidis, rugosis, apice
truncatis.

A densly caespitose plant from creeping rhizomes, the culms rush-like,
often nearly 2 m high, sometimes shorter, terete, glabrous, 2 to 3 mm
in diameter, their bases covered with several short, imbricated bracts,
leafless except for the 2 or 3 culm leaves which are much reduced or
sometimes represented only by sheaths, or sometimes 5 cm long, always
less than 3 mm wide, vertically compressed, glabrous. Panicles 10 to
15 em long, interrupted, thyrsoid, lax, both the primary and secondary
rachises strongly zigzag, on the ultimate branches each group of two or
three spikelets subtended by an ovate, keeled, brown, prominently acu-
minate bract about 3 mm long, the basal portion broad, 7- to 9-nerved,
the spikelets sessile or shortly pedicelled, crowded. Spikelets brown,
glabrous, 2 to 2.5 mm long, each bearing a single perfect flower, the glumes
few, about four, ovate to oblong-ovate, somewhat acuminate, keeled, 2
mm long. Nutlet ellipsoid, terete, straw-colored, somewhat shining,
about 1.8 mm long, wrinkled when dry, the base somewhat acute, the
apex minutely puberulent, truncate or rounded ; style filiform, elongated,
the arms three.

Mrnporo, southwest of Lake Naujan, altitude about 120 m,. For. Bur. 6724
Merritt, April, 1907 (type). Luzon, Province of Zambales, along streams near

Candelaria, Bur. Sci. 4729 Ramos, December, 1907. PALnAwaNn, Mount Victoria,
on rocks at base of waterfall, Bur. Sci. 718 Foxworthy, March, 1906, altitude

142 MERRILL.

about 600 m; Iwahig, in bed of mountain stream, altitude about 300 m, Merrill
758, February, 1903, specimen very young.

This species is probably most closely allied to Cladiwm riparium Benth., of
Australia, and to the var. crassum (Thwaites) Clarke of India and Ceylon; it is,
apparently, distinct from both forms. Among the Philippine species it is prob-
ably most closely allied to Cladiwm distichum Clarke but lacks the numerous
imbricated glumes of that species.

Cladium filiforme sp. nov. § Hucladium.

Caespitosum, tenerum, circiter 40 cm altum; foliis angustis, planis,
elongatis, in sicco plus minus plicatis, margine scabridis; paniculis laxis,
angustis, thyrsoideis; spiculis paucis, lanceolatis, brunneis, circiter 5.5
mm longis, 1-floris; fructibus nitidis, ovoideis vel ellipticis, albidis, ses-
silibus, vix rostratis.

A caespitose perennial about 40 cm high, slender. Stems terete, about
1.5 mm in diameter. Leaves mostly basal, slender, about 20 cm long,
1.5 mm wide, scabrid, apparently plane when fresh, more or less folded
when dry, the culm leaves two or three, similar to the basal ones. Pani-
cles slender, thyrsoid, comparatively few-flowered, narrow, rather lax.
Spikelets lanceolate; brown, 5 to 6 mm long, their pedicels slender, 2 to
8 mm in length, each group of three or four spikelets subtended by a
setaceous, scabrid, leaf-hke bract 1 cm long or less. First two glumes
empty, lanceolate-ovate, acuminate, about 3 mm long. Third glume in-
closing a perfect flower, narrowly oblong, acuminate, 3.5 mm long, thicker
than the empty glumes. Stamens three; anthers linear, 2 mm long.
Nutlet ovoid or ellipsoid, white and shining, sessile, not beaked, gla-
brous; style 5 mm long, divided to the middle into three arms. Fourth
glume similar to the third, empty, the fifth smaller and thinner, also
empty.

PALAWAN, Mount Victoria, Bur. Sci. 717 Foxworthy, March 24, 1906, on rocks
at base of a waterfall, altitude about 600 m.

A species characterized by its slender habit, slender leaves and panicles, some-
what resembling Cladium undulatum Thwaites but smaller and with no traces of

hypogynous bristles.
SCIRPUS Linn.

Scirpus lacustris Linn. Sp. Pl. (1753) 48; Clarke in Hook. f. Fl. Brit. Ind.
6 (1893) 658.

Luzon, Province of Cagayan, Buguey, For. Bur. 17290 Curran, March, 1909,
a common and conspicuous plant in lagoons back of the town.

Frequent in fresh water nearly throughout the world, except South America
and Malaya. Not previously reported from the Philippines.

XYRIDACEA.
XYRIS Linn,

Xyris anceps Lam. Ill. 1 (1791) 1382; Hook. f. Fl. Brit. Ind. 6 (1892) 364.
Luzon, Province of Isabela, Carig, Bur. Sci. 8065 Ramos, May, 1909. SEME-
RARA, Merrill 4151, June, 1905.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 173

Not previously reported from the Philippines, and the second species definitely
known to occur in the Archipelago; India to Burma, the Malay Peninsula and
Archipelago.

LILIACEZ.

SMILAX Linn.

Smilax verruculosa sp. nov. § Eusmilax.

Species S. bracteatae Presl valde affinis, differt ramis ramulisque dense
verruculosis et spinis plus minus numerosis armatis.

Scandent, the branches and branchlets yellowish or brownish, rather
slender, terete, or the latter slightly sulcate, densely verruculose and with
numerous, straight or slightly curved, sharp spines 1 to 3 mm long.
Leaves broadly ovate to elliptic-ovate, coriaceous, shining, 6 to 10 cm
long, 4 to 7 em wide, the base rounded or acute, the apex shortly and
abruptly apiculate-acuminate, the acumen thickened ; nerves 5, the outer
pair faint, submarginal, the inner three stout, prominent, the inner pair
leaving the middle one just above the base, the reticulations lax,
prominent; petioles about 1 cm long, the lower half inflated, somewhat
clasping the stem, some tendril-bearing at about the middle, others simply
auriculate. Inflorescence axillary, solitary, the rachis emerging from
between two, ovate, coriaceous, 4 to 5 mm long bracts, each inflorescence
consisting of from 1 to 4 racemosely disposed, peduncled umbels, the
peduncles to the umbels subtended by small bracts, solitary. Flowers
10 to 20 in each umbel, 3.5 to 4 mm long, the perianth-segments reflexed.
Ovary-cells 1-ovuled. Fruit ovoid, about 3.5 long, 1-seeded.

Luzon, Province of Benguet, Baguio, Topping 13 (type), Hlmer 5820, 8572,
Williams 1046, Bur. Sci. 2810, 3878 Mearns, For. Bur. 15617 Curran.

Most of the above specimens have been distributed as Smilax bracteata Presl,
from which the present species is at once distinguished by its densely verruculose
and more or less spiny branches and branchlets. The terminal undeveloped bud is

sometimes present on the racemes, but more often absent, or developed into an
umbel.

Smilax williamsii sp. nov. § Husmilaw.

Frutex alte scandens, ramis ramulisque teretibus vel leviter striatis,
verruculosis, vix spinosis; foliis amplis, late elliptico-ovatis, chartaceis vel
subcoriaceis, basi acutis vel subcordato-rotundatis, apice abrupte apiculatis,
nervis 5 vel 7; inflorescentiis axillaribus, solitariis, umbellis 1 vel 2,
racemoso-dispositis. |

A scandent shrub, the branches and branchlets terete, or slightly
striate, pale-brown, densely verruculose, not spiny. Leaves alternate,
ample, broadly elliptic-ovate, chartaceous or subcoriaceous, 9 to 15 cm
long, 5 to 11 cm wide, shining, base acute or subcordate-rounded, the
apex shortly and abruptly apiculate; primary nerves 5, basal, prominent,
with an additional pair of fainter submarginal nerves, the reticulations
distinct; petiole 1.5 to 2 cm long, the lower half inflated, clasping the
stem, auriculate or tendril-bearing at about the middle, curved. Racemes

174 MERRILL.

axillary, solitary, the rachis emerging from between the petiole and an
ovate, coriaceous, 5 to 7 mm long bract, the umbels 1 or 2, the terminal
bud wanting, the peduncles to the umbels solitary, subtended by bracts,
slender, 2.5 em long. Staminate flowers 20 to 25 in each umbel, about
7 mm long, the perianth segments reflexed; stamens 6. Fruit globose,
much wrinkled when dry, with from 1 to 3, more or less compressed, 4
mm long seeds.

MINDANAO, Lake Lanao, Mrs. Clemens 751, near streams, September, 1906:
District of Davao, Williams 2519, March, 1905 (type).

A species manifestly allied to Smilax bracteata Presl, but distinguished at once
by its verruculose, but not spiny stems, larger leaves with more numerous veins,
and larger flowers.

ULMACE.
CELTIS Linn.

Celtis crenato-serrata sp. nov.

Arbor circiter 30 m alta, glabra, vel ramulis ultimis parce pubes-
centibus; foliis subcoriaceis, ovatis vel elliptico-ovatis, acuminatis, basi
rotundatis, leviter inaequilateralibus, 3-nerviis, margine in tertia inferiore
parte integris, supra valde crenato-serratis ; fructibus ovatis, leviter com-
pressis, circiter 1 cm longis. |

A glabrous tree about 30 m high. Branches slender, terete, reddish-
brown, distinctly lenticellate with small lenticels, the ultimate branchlets
slightly pubescent. Leaves ovate to elliptic-ovate, 7 to 10 em long, 3
to 5 cm wide, subcoriaceous, slightly shining, of the same color on both
surfaces or slightly paler beneath, the base broad, somewhat inequilateral,
rounded on one side of the midrib, subacute on the other side, the apex
prominently acuminate, the acumen 1 to 1.5 cm long, apiculate, the
margins in the lower one-third entire, above prominently crenate-serrate ;
basal nerves three, prominent, the two lateral ones extending nearly to
the apex and above somewhat looped at the anastomoses of the lateral
veins, the lateral ones slender, horizontal, about 10 to 12 on each side
of the midrib, the reticulations lax, indistinct ; petioles 5 to 7 mm long.
Flowers unknown. Fruits ovate, slightly compressed, about 1 cm long,
the pericarp thin, fleshy.

Luzon, Province of Bataan, Duale, Mor. Bur. 20043 Topacio, October 2, 1909,

in forests along streams, altitude about 100 m, locally known as malabatulan.
A species well characterized by its prominently crenate-serrate leaves.

ARISTOLOCHIACEH.
ARISTOLOCHIA Linn.

Aristolochia macgregorii sp. noy. § Diplolobus.

Folis subtus ad costam, ramulis inflorescentiis petiolisque plus minus
breviter hirsuto-pubescentibus; foliis chartaceis, oblongis, basi subsagit-
tato-cordatis, apice breviter acute acuminatis, petiolo vix 5 mm longo;
racemis axillaribus, solitariis, floribus cireiter 4 cm longis.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. ivi

Apparently scandent. Branches terete, grayish, slightly striate, slightly
zigzag, the leaf-bearing branchlets more or less densely pubescent with
short brownish hairs. Leaves oblong, 11 to 17 cm long, 4.5 to 6.5 cm
wide, chartaceous, shining when dry, above entirely glabrous, beneath
somewhat hirsute-pubescent with short hairs on the midrib and primary
nerves, the apex shortly and sharply acuminate or merely acute, the base
sagittate-cordate, the auricles broad, rounded, the sinus somewhat obtuse,
about 1 em deep, the auricles somewhat surrounding the stems but free
from them; basal nerves two or three pairs, the lower pair or pairs short,
the upper pair reaching to about the middle of the leaf, the primary
nerves above the basal ones 3 or 4 on each side of the midrib, anasto-
mosing, the reticulations lax; petioles pubescent, less than 5 mm long.
Inflorescence axillary, solitary, simply racemose, the rachis 1 to 1.5 em
long, pubescent, the pedicels about 3 mm in length, each opposed by an
ovate-lanceolate, acuminate bract, the lower ones 6 mm long, the upper
gradually shorter. Flowers 4 cm long, the basal 4 mm ovoid, narrowed
and tubular above, the tube about 16 mm long, 2 to 2.5 mm in diameter,
the upper portion expanded, the lp pubescent, lanceolate, acuminate,
about 2 em long, 3 mm wide. Column very obscurely lobed. Anthers
6, 1 mm long. Fruit (immature) obovoid, 1.5 cm long.

BABUYANES ISLANDS, Dalupiri, Bur. Sct. 10656 McGregor, August 20, 1909.

A species manifestly allied to Aristolochia tagala Cham., but at once distin-
guished by its differently shaped leaves, which are pubescent on the nerves beneath,
very short petioles, dense racemes, and quite different flowers.

CHENOPODIACEA.
CHENOPODIUM Linn.

Chenopodium polyspermum Linn. Sp. Pl. (1753) 220.

Luzon, Province of Benguet, Baguio, Leon Guerrero, March, 1910.

In waste places, apparently of recent introduction; a widely dispersed European
weed, introduced and now widely distributed in eastern North America.

NYCTAGINACEA.
PISONIA Linn.

Pisonia gammillii sp. noy.

Arbor glabra, inflorescentiis exceptis, circiter 10 m alta; foliis oblongo-
ellipticis, in sicco chartaceis, nitidis, breviter acuminatis, basi inaequi-
lateralibus, acuminatis, usque ad 20 em longis; inflorescentiis laxis, ter-
minalibus axillaribusque; floribus hermaphroditis; staminibus 12 vel 13,
breviter exsertis.

A glabrous tree, except the inflorescence, unarmed, about 10 m high,
the trunk 40 cm in diameter. Leaves mostly opposite, oblong-elliptic,
ample, 17 to 20 cm long, 8 to 10.5 cm wide, when dry chartaceous and
somewhat shining, apparently somewhat fleshy when fresh, entire, the
apex shortly acuminate, the base acuminate-decurrent, inequilateral ;

176 . MERRILL.

lateral nerves 8 or 9 on each side of the midrib, distant, rather distinct,
anastomosing, the reticulations obsolete; petioles 1 to 2 em long. In-
florescence axillary and terminal, the branches and branchlets umbellately
arranged, ample, lax, nearly as long as the leaves, the axillary peduncles
1 or 2, the terminal ones about 5, the younger parts ferruginous-pubes-
cent, the peduncles 9 to 11 cm long, with or without a single node;
primary branches umbellately disposed, 4 or 5, 1.5 to 3 em long, spread-
ing, each bearing from 2 to 5 umbellately disposed secondary branches
6 to 12 mm in length. Flowers white, fragrant, 2 to 6 at the tips of
the ultimate branchlets, the perianth 6 to 7 mm long, the pedicels 2 to 3
mm long, puberulent. Perianth urceolate, the throat up to 5 mm in
diameter, the lobes 5, spreading or somewhat reflexed, 2 to 2.5 mm
broad, about 1 mm long, apiculate. Ovary and style about 5 mm long;
stigma fimbriate, about 2 mm in diameter. Stamens 12 or 13; filaments
slender, glabrous, somewhat united below, unequal, 4 to 6 mm long, some-
what exserted; anthers 0.8 mm. long. Fruit unknown. .

GUIMARAS, Nagaba, For. Bur. 288 Gammill, February 22, 1904, in upland
valleys, altitude about 50 m, locally known as anuring.

A species well characterized by its ample leaves and very lax inflorescence. Its
flowers are apparently all hermaphrodite. Manifestly allied to Pisonia wmbellifera

(Forst.) Seem. (P. excelsa Bl.), but with more numerous stamens, larger flowers,
and quite different inflorescence.

MAGNOLIACEE.

KADSURA Juss.

Kadsura paucidenticulata sp. nov.

Frutex scandens, glaber; foliis ellipticis vel anguste obovato-ellipticis,
acuminatis, chartaceis; floribus masculinis terminalibus, solitariis, circi-
ter 2 cm diametro, pedicellis ebracteolatis, sepalis petalisque vix vel
‘obscure puncticulatis.

A scandent glabrous shrub. Branches terete, with scattered large lenti-
cels, dark-colored when dry. Leaves mostly on short lateral branchlets,
elliptic or narrowly obovate-elliptic, chartaceous, 4 to 7 em long, 2.5 to
3.5 em wide, minutely glandular-puncticulate, somewhat shining, the
apex abruptly short-acuminate, the base acute, the margins in the upper
half with few, scattered, small teeth; lateral nerves 5 to 7 on each side
of the midrib, not prominent, not much more distinct than are the
primary reticulations; petioles 0.4 to 0.7 mm long. Staminate flowers
solitary, about 2 cm in diameter, terminating the stems and the short
lateral branches, the pedicels about 12 mm long, ebracteolate. Sepals
about 5, ovate to elliptic-ovate, obtuse or rounded, the outermost one
3 mm long and 2 mm wide, the inner ones gradually larger, the largest
about 8 mm long, 6 mm wide, very obscurely or not glandular-punctate,
margins minutely ciliate. Petals about 7, the outer four narrowly obovate

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. Lz

or oblong-obovate, thick, 10 mm long, 6 mm wide, broad and rounded at
the apex, the margins glabrous, the inner three similar but smaller, 8
mm long or less. Stamens many, united into a rather dense, depressed-
globose head, the connectives very broad, the anthers 0.8 mm long.

Luzon, Province of Benguet, Pauai, Bur. Sci. 8498 McGregor, June, 1909, alti-
tude about 2,100 m.

A species allied to Kadsura philippinensis Elmer, differing especially in its
terminal, comparatively short-pedicelled flowers and other minor characters.

Kadsura macgregorii sp. noy.

Species praecedenti simillima et ut videtur valde affinis, sed differt
floribus masculinis axillaribus, breviter pedicellatis, pedicellis basi bract-
eolis imbricatis praeditis.

Scandent, glabrous, the branches dark-colored, lenticellate. Leaves
similar in shape and size to those of the preceding species, membran-
aceous, rather densely and minutely glandular-punctate, the margins in
the upper half with scattered, small teeth. Staminate flowers solitary,
in the axils of leaves or of fallen leaves, their pedicels about 5 mm long,
each subtended by several, densely imbricated, brown bracteoles about 1
mm long and longer than wide, and with a similar one at about the
middle of the pedicel. Sepals about 5, orbicular or broadly orbicular-
ovate, minutely and densely glandular-punctate, rounded, the outer one
about 2 mm long, the inner gradually larger, the innermost about 8 mm
long, their margins minutely ciliate. Petals about 5, somewhat larger
than the inner sepals, distinctly and densely glandular-punctate. Sta-
mens united in a globose mass, the connectives very broad.

Luzon, Province of Benguet, Pauai, Bur. Sci. 8340, McGregor, June, 1909,
altitude about 2,100 m.

A species in general appearance quite similar to Kadsura paucidenticulata,
but at once distinguished by its axillary, short-pedicelled staminate flowers, and
by its pedicels subtended by several, small, imbricated bracts. It is apparently
more closely allied to Kadsura philippinensis Elmer, than is the preceding

species, judging from the attachment of the flowers, but its short-pedicelled
flowers and denticulate leaves are sufficient to distinguish the two species.

SAXIFRAGACEAE.
CURRANIODENDRON gen. nov.

Genus Dedeae Baill. valde affine, sed differt floribus 4-meris, ovulis
numerosis, usque ad 16.

Curraniodendron dedeaeoides sp. nov.

Arbuscula glabra, dioica, 2 ad 3 m alta, ramulis foliisque junioribus
plus minus resinosis ; foliis alternis, oblongis vel oblongo-ellipticis, leviter
acuminatis, chartaceis vel subcoriaceis, nitidis, subtus plus minus nigro-
glandulosis; racemis axillaribus, solitariis, floribus femineis parvis, 4-
meris.

178 MERRILL.

A glabrous dioecious shrub 2 to 3 m high, glabrous, the young branches
and leaves more or less resinous, shining. Branches terete, slender,
dark-colored and longitudinally striate when dry, with scattered lenticels,
the young branchlets somewhat compressed and angular. Leaves alter-
nate, oblong to oblong-elliptic, chartaceous or subcoriaceous, 6 to 11 em
long, 2 to 3.2 em wide, entire, the apex shortly and not prominently
acuminate, the base acute or somewhat decurrent-acuminate, shining,
somewhat paler beneath when dry, and at least the older leaves with
numerous, small, black glands on the under surface at the intersections
of the ultimate reticulations; primary lateral nerves about 12 on each
side of the midrib, not prominent, spreading, obscurely anastomosing,
the secondary alternating ones often nearly as prominent, the ultimate
reticulations rather dense, fine, the basal pair of nerves ascending, anas-
tomosing with the other lateral nerves shortly above the base of the leaf;
petioles 1 to 2 em long; stipules none. Racemes axillary, solitary, 5 to
7 cm long, more or less resinous, as are the buds and young flowers.
Pistillate flowers alternate, 4-merous, whitish, rather scattered, solitary
or sometimes two in the axil of each bracteole, the bracteoles oblong-
ovate, about 1.5 mm long, 1 mm wide, deciduous, the pedicels 2 to 3
mm long. Calyx-tube funnel-shaped, about 2 mm long and wide, bear-
ing four broadly triangular-ovate, 0.5 mm long lobes. Petals 4, alter-
nating with the calyx-lobes, oblong-ovate, obtuse, about 2 mm long, 1.3
mm wide, attached by a broad base, spreading or reflexed, in bud dis-
tinctly imbricate. Imperfect stamens 4, alternating with the petals, the
filaments about 1 mm long; anthers 0.8 mm long, oblong-ovate, basifixed,
bearing no pollen. Ovary half inferior, the free portion broadly conical,
somewhat sulcate, glabrous, 1-celled, with 4 or 3 prominent parietal
placentae; ovules 16 or 12, attached to the introflexed margins of the
placentae, ascending, imbricate; styles 4 or 3, entirely connate into a
sulcate, 1 mm long column, bearing 4 or 3 minutely papillate, depressed-
hemispheric stigmas. Staminate flowers and fruits unknown.

Necros, Mount Marapara, For. Bur, 13634 Curran & Foxworthy, September 8,
1909, in the mossy forest of the summit, altitude about 1,300 m.

This proposed new genus is manifestly closely allied to Dedea Baill., a genus
of two or possibly three species confined to New Caledonia. It differs from that
genus in its 4-merous flowers and in its more numerous ovules, as well as in
some other minor characters. It agrees with it not only in gross characters
and general appearance, but especially in its 1-celled ovary, which is unusual in the
family.

We are fortunate in having in this herbarium cotypes of the three species of
Dedea proposed by Baillon, and the present species in facies is very similar to
). minor Baill., and D. media Baill. Distinguishing characters that at once
strike the eye are the somewhat resinous younger parts of the Philippine plant
and its older leaves distinetly glandular beneath with numerous, small, black
glands, while all of Baillon’s species are prominently lepidote, this character
being absent in the form above described. In general appearance, however,
Curraniodendron dedeaeoides is exceedingly similar to Dedea minor and D. media:

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 179

an examination of the pistillate flowers, however, shows sufficiently important
differential characters to warrant the characterization of the Philippine plant
as a distinct genus.

PITTOSPORACE AH:
PITTOSPORUM Banks.

Pittosporum littorale sp. nov.

Arbor glabra usque ad 6 m alta; foliis anguste oblongo-obovatis, sub-
coriaceis, apice rotundatis, basi angustatis, decurrento-acuminatis vel
acutis; fructibus aurantiacis, ovoideis, circiter 2 cm longis, 2-valvatis,
apiculatis, in sicco rugosis; seminibus circiter 20, nigris.

A glabrous tree about 6 m high. Branches terete, light-gray, smooth.
Leaves somewhat crowded toward the apices of the branchlets, subcoria-
ceous, narrowly oblong-obovate, 9 to 17 em long, 3 to 5 cm wide, when
dry somewhat shining, paler beneath, the apex rounded, broad, rarely
somewhat acute, the base gradually narrowed, acute or decurrent-acum-
inate, the margins often somewhat recurved; nerves about 15 on each
side of the midrib, not prominent; petioles 2 to 2.5 cm long. Flowers
unknown. Fruiting racemes 2 to 3 cm long, in the upper axils. Fruits
ovoid, about 2 cm long, 2-valved, valves ultimately recurved, orange-yellow
when fresh, wrinkled when dry, the pericarp rather thick. Seeds about
20, black, shining. :

Minporo, For. Bur. 9845 Merritt, March, 1908, along the seashore. SIQuIJoR,
For. Bur. 16999 Everett, December, 1907, rocky point at Liloan, locally known
as ticala.

‘A species quite different from any, of the other Philippine form, readily
distinguishable by the shape of its leaves.

Pittosporum megacarpum sp. nov.

Arbor vel arbuscula glabra, usque ad 8 m alta, ramis pallide griseis,
teretibus; foliis chartaceis, oblongo-ellipticis vel obovato-ellipticis, basi
acutis, apice abrupte acuminatis; nervis utrinque 8 ad 10, subtus pro-
minentibus ; fructibus ovoideis, in sicco yalde rugosis, 4 cm longis.

A shrub or tree 3 to 8 m high, glabrous throughout. Branches terete,
light-gray. Leaves chartaceous, oblong-elliptic to obovate-elliptic, 10 to
18 cm long, 4 to 7 cm wide, shining when dry, the apex abruptly and
sharply acuminate, the acumen 1 cm long or less, the base acute; nerves
8 to 10 on each side of the midrib, prominent beneath, impressed on
the upper surface, anastomosing; petioles 1 to 2 cm long. Flowers un-
known. Fruits ovoid, 2-valved, yellow, densely wrinkled when dry, about
4 cm long, shortly apiculate, the pericarp thick. Seeds many, irregular,
about 6 mm long, black, minutely wrinkled when dry.

MassBatTeE, Bulo River, For. Bur. 12557 Rosenbluth, January, 1909. MrNporo,
Balete River, For. Bur. 5892 Merritt, October, 1906.

A species with larger fruits than any other known Philippine form, Among
our species most closely allied to Pittosporum odoratum Merr.

180 MERRILL.

Pittosporum ramosii sp. noy.

Arbuscula vel arbor glabra, 3 ad 6 m alta; foliis elliptico-ovatis vel
oblongo-ovatis, usque ad 6 cm longis, utrinque acuminatis, nervis tenui-
bus, vix distinctis; fructibus lateralibus, ovoideis, apiculatis, 1 ad 1.2
em longis, 2-valvatis.

A shrub or tree 3 to 6 m high, glabrous. Branches terete, dark-gray
or brownish. Leayes somewhat crowded at the apices of the branchlets,
subcoriaceous, shining when dry, elliptic-ovate to oblong-ovate, 4 to 6
em long, 2 to 2.5 em wide, the apex sharply subcaudate-acuminate, the
base decurrent-acuminate; nerves about 8 on each side of the midrib,
slender, indistinct, the reticulations fine, close; petioles slender, about 1
em long. Flowers unknown. Fruit from the branches below the leaves,
axillary, solitary (the inflorescence apparently a short raceme), 2-valved,
ovoid, smooth, yellow, apiculate, 1 to 1.2 em long, the peduncles about
1 cm long; seeds few, 10 or less, black, shining, about 3 mm long.

Luzon, District of Lepanto, Balbalasan, For. Bur. 5683 Klemme, altitude 1,500 m:
Province of Abra, Mount Bawagan, Bur. Sci. 7211 Ramos (type), February, 1909.

The fruits are very oily and with a strong odor of turpentine. Those that
have been in the herbarium three years, when opened, were still wet with the

aromatic oil characteristic of the genus. Similar in some respects to P. pen-
tandrum Merr., but distinguished by its lateral inflorescence and by its leaves.

Pittosporum ramiflorum Zoll. ex Miq. Fl. Ind. Bat. 1? (1858) 122.

Glyaspermum ramiflorum Zoll. & Mor. in Nat. Gen. Arch. Neerl. Ind. 2
(1845) 11.

Pittosporum clementis Merr. in Philip. Journ. Sci. 3 (1908) Bot. 137.

Additional study of the type material of Pittosporwm clementis and comparison
of the same with Javan material representing Pittosporum ramiflorum Zoll. has
convinced me that the two species are identical and that P. clementis should be
reduced to Zollinger’s species. It is known from Java, Amboina, Celebes, Min-
danao, and Negros (Canlaon Volcano, Phil. Pl. 228 Merrill, April, 1910). The
name Pittosporwm ramiflorum Zoll. is not listed in Index Kewensis or in any of
the supplements of that work.

ROSACEAE.
PRUNUS Linn.

Prunus junghuhnianus Miq. Il. Ind. Bat. 1* (1855) 366.

PALAWAN, Mount Victoria, Bur. Sci. 731 Foxworthy, March 24, 1906, in stream
depressions, altitude about 250 m.

The specimen agrees well with Miquel’s description, except that the racemes
are nearly or quite glabrous, and also agrees perfectly in twig and leaf characters
with sterile material received under the above name from Jaya.

New to the Philippines; previously definitely recorded only from Java.

RUTACEZ.
CLAUSENA Burm.

Clausena worcesteri sp. nov.
Arbor vel arbuscula glabra; foliis alternis, foliolis 2—3-jugatis, ovatis,
nitidis, apice abrupte obtuse acuminatis; paniculis terminalibus, pauci-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 181

floris, corymbosis; floribus 5-meris, petalis basi acutis ; fructibus junior-
ibus Genta stipitatis.

An erect shrub or small tree glabrous throughout. Branches slender,
terete, shining, nearly black when dry. Leaves simply pinnate, less than
20 cm long, 2- or 3-jugate, the petiole and rachis terete, slender. Leaflets
ovate, firmly chartaceous, shining 5 to 7 cm long, 2.5 to 4 em wide, entire,
the base acute or fmsded sometimes inequilateral, apex rather prom-
inently acuminate, acumen blunt or retuse, prominently glandular-punc-
tate; nerves about 5 on each side of the midrib, distant, anastomosing,
the reticulations lax; petiolules 3 to 5 mm long. Panicles terminal,
comparatively few-flowered, corymbose or subcorymbose, the branches few.
Flowers 5-merous, white. Sepals broadly ovate, acute or obtuse, about
1mm long. Petals 5, oblong-oblanceolate or oblanceolate, 5 to 5.5 mm
long, 1.5 mm wide, the apex acute or slightly acuminate, narrowed below
to the acute base, imbricate, somewhat coherent in the upper part. Sta-
mens 10, the longer filaments 4 mm in length, abruptly narrowed from
1 mm below the anther, the alternating shorter filaments 3 mm long,
abruptly narrowed just below the anther; anthers 1 mm long. Ovary
oblong, cylindric, glabrous, about 2 mm long, 5-celled; styles thick, 2
mm long, slightly sulcate. Young fruits ovoid or ellipsoid, with a
distinct, stout, 1 mm long stipe.

Luzon, Province of Cagayan, Apiao, near Tauit, Bur. Sci. 10743 Worcester,
August, 1909.

This species is distinguished from all known Philippine forms by being quite
glabrous. It is well characterized by its few leaflets, corymbose or subcorymbose,
few-flowered panicles, its cylindric, glabrous ovary, and stipitate fruits. It is
named in honor of its collector, Hon. Dean C. Worcester, Secretary of the Interior
of the Philippine Government. When fresh the leaves are very aromatic.

Sterile material of what is manifestly the same species has been collected at
the Mission River and on Mount Aluntang, both in the Province of Cagayan,
For. Bur. 17165, 17347 Curran. Mr. Curran notes that the plant is used by the
Negritos for ornamental purposes and for its odor.

Clausena mollis sp. noy.

a

Arbuscula circiter 5 m alta, omnibus partibus plus minus dense mol-
liter pubescens ; foliis 20 ad 30 em longis, foliolis alternis vel subalternis,
5 ad 8 utrinque, integris, valde inaequilateralibus ; paniculis terminalibus,
anguste pyramidatis, floribus 5-meris, sessilibus vel subsessilibus, subglom-
eratis ; fructibus globosis, albidis vel albido-viridibus, 1-spermis.

A slender shrub about 5 m high. Branches and branchlets terete,
grayish or brownish, the younger ones often greenish, and with the
petioles densely and softly pubescent with short spreading hairs. Leaves
alternate, 20 to 30 cm long, the leaflets alternate 5 to 8 on each side of
the pubescent rachis, chartaceous, somewhat shining, rather pale when
dry, oblong-ovate, entire, 5 to 9 cm long, 2.5 to 3.5 cm wide, the base
rounded or acute, strongly inequilateral, the apex shortly acuminate or
nearly blunt, prominently glandular-punctate, both surfaces softly pubes-

182 MERRILL.

cent, especially on the nerves, or the upper surface pubescent only on
the midrib; nerves about 8 on each side of the midrib; petiolules densely
pubescent, about 2 mm long. Panicles terminal, narrowly pyramidal,
about 20 cm long, the rachis, branches and branchlets densely and softly
pubescent with pale spreading hairs, the lower branches 7 cm long or
less, the upper ones gradually shorter, spreading or ascending. Flowers
5-merous, sessile or shortly pedicelled, subglomerate on the ultimate
branchlets, the buds globose or obovoid. Sepals broadly ovate, 1.2 mm
long, free, densely pubescent outside. Petals elliptic or broadly elliptie-
oblong, concave, about 4 mm long, 2.5 mm wide, acute at both ends,
imbricate, with few, rather large glands, the back in the upper third
slightly pubescent. Stamens 10, the filaments broad, abruptly narrowed
just below the insertion of the anthers, about 1 mm long; anthers 2 mm
long. Ovary ovoid or ellipsoid, about 1.5 mm long, glabrous, prominently
5-sulcate, 5-celled, each cell with two superimposed ovules; style 1 mm
long and thick, 5-suleate. Fruit globose, white or greenish-white, 6 to
7 mm in diameter, the pericarp rather thick, glandular-punctate, con-
taining a single somewhat compressed seed about. 5 mm in diameter,
surrounded by a gelatinous pulp, with a strong odor and taste of pine
pitch; cotyledons flat, plano-convex.

Luzon, District of Bontoc, near Bontoe, For. Bur. 16530 Curran (type), Jan-
uary 21, 1909, altitude above 1,000 m: Province of Benguet, Twin Peaks, Hlmer
6352, May, 1904. Bur, Sei. 7863 Ramos from Cagayan Province, Luzon, with
immature fruits, is possibly referable here, while a specimen from Zamboanga,
Mindanao, Hallier s. n., has much the facies of the present species, but has

4-merous flowers and 2-seeded fruits. Clausena mollis is well characterized by its
soft, rather dense pubescence.

MELICOPE Forst.

Melicope densiflora sp. nov.

Arbuscula glabra 3 ad 5 m alta; foliis trifoholatis, foliolis chartaceis
vel submembranaceis, oblongo-obovatis, apice late breviter acuminatis,
acuminibus retusis, nervis utrinque circiter 8; inflorescentiis axillaribus,
petiolo aequalibus vel longioribus; floribus 4-meris, filamentis pilosis.

A shrub 3 to 5 m high, glabrous throughout. Branches terete, light-
gray, the branchlets pale-reddish-brown, rather stout. Leaves 3-foliolate,
opposite, their petioles 2 to 5 em long; leaflets oblong-obovate to elliptic-
obovate, chartaceous or submembranaceous, somewhat shining, 7 to 11
em long, 3 to 5 em wide, the apex shortly and broadly acuminate, the
acumen retuse, narrowed from about the middle to the cuneate base, the
terminal leaflet equilateral, the lateral ones somewhat inequilateral ;
petioles 3 to 10 mm long; lateral nerves about 8 on each side of the
midrib, anastomosing, the secondary ones often nearly as prominent.
Inflorescence axillary, solitary, 5 to 6 em long, narrowly pyramidal, the
lower branches often 3.5 em in length, rather densely flowered. Stamin-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 183

ate flowers somewhat fascicled on the ultimate branchlets, their pedicels
1.5 to 2 mm long, glabrous. Sepals 4, ovate, acute, 0.8 mm long.
Petals 4, oblong, 3 mm long, 1 to 1.3 mm wide, thin, obtuse, the apex
appendiculate inside. Stamens 8, the filaments rather densely clothed
with spreading hairs in the lower half, the longer four 3 mm, the shorter
four 2 mm in length; anthers 0.8 mm long. Pistillate flowers similar
to the staminate ones, the staminodes slightly pubescent. Ovary ovoid,
glabrous, 1.2 mm long, glabrous, deeply longitudinally 4-suleate, 4-celled,
each cell 2-ovuled; style very short (less than 0.5 mm) ; stigma radiately
4-lobed.

BATANES ISLANDS, Batan, Santo Domingo de Basco, Bur. Sci. 3235 Mearns,
- Bur. “Sci. 3603 Fénix, Bur. Sci. 10682 McGregor, locally known as idacacayo:
Sabtan, Bur. Sci. 10676 McGregor.

Some of these specimens were previously referred by me* to Melicope luzonensis
Engl., but they are sufficiently distinct to warrant description as a separate
species. M. densiflora is distinguished from M. luzonensis Engl. by its differently
shaped and fewer nerved leaves, its dense inflorescence, and especially by its pilose
filaments and staminodes. Bur. Sci. 3215 Mearns is possibly referable here, but
its filaments seem to be quite glabrous.

EVODIA Forst.

Evodia acuminata sp. nov.

Arbor inflorescentiis exceptis glabra, circiter 10 m alta; foliis trifo-
liolatis vel aliis unifoliatis, foliolis subcoriaceis, oblongo-obovatis, nitidis,
basi acuminatis, apice abrupte subcaudato-acuminatis, acuminibus circiter
1 cm longis; inflorescentiis axillaribus, pedunculatis, 6 ad 8 cm longis;
floribus 4-meris, sepalis petalisque glanduloso-punctatis.

A tree about 10 m high, glabrous except the inflorescence, the ultimate
branchlets, petioles and under surfaces of the leaves distinctly glandular-
punctate, the branches terete, smooth, pale-brownish, the ultimate branch-
lets somewhat compressed. Leaves opposite, the petioles 6 to 9 cm long,
or those of unifoliolate leaves onky about 2 cm long; leaflets usually
three, sometimes solitary, oblong-obovate, subcoriaceous, shining, paler
beneath, 9 to 14 cm long, 4 to 6 cm wide, entire, the apex broad,
abruptly swbhcaudate-acuminate, the acumen about 1 cm long, the base
decarrent-acuminate, the lateral leaflets somewhat inequilateral; petio-
lules 0.5 to 1.5 em long; primary lateral nerves 8 or 9 on each side of
the midrib, distant, irregular, anastomosing, the secondary ones often
nearly as prominent. Cymes axillary, peduncled, the peduncles 3.5 to
5 em long, solitary, glabrous, each cyme about 3 cm wide, the branches
and branchlets cinerous-puberulent. Flowers white, somewhat crowded,
their pedicels 3 to 4 mm long, puberulent, each subtended by two or
three, ovate, 1 mim long bracteoles. Sepals 4, orbicular, imbricate, about
3 mm in diameter, rounded, glandular-punctate, margins minutely ciliate.

1This Journal 3 (1908) Botany 411.
95948—_—2

184 MERRILL.

Petals 4, oblong or oblong-ovate, about 5 mm long, 2.2 to 2.5 mm wide,
apex acute, minutely appendaged inside. Stamens 4, the filaments stout,
2mm long, bearing imperfect, oblong, 1 mm long anthers. Ovary pubes-
cent, deeply 4-lobed, 4-celled, each cell 2-ovuled; style stout, 3 mm long,
pubescent; stigma subcapitate. Staminate flowers and fruits unknown.

Luzon, Province of Sorsogon, Sorsogon, For. Bur. 10520 Curran, June 10, 1908,
near abaca (Musa textilis) plantations, altitude about 200 m.

A species well characterized by its abruptly acuminate leaflets, 3-foliolate and
l-foliolate leaves occurring on the same branches, its rather small, rather long-
peduncled cymes, and comparatively large flowers. It is apparently as closely
allied to Hvodia glabra Bl., as to any other species, but is quite different
from that.

MELIACEZ.
AGLAIA Lour.

Aglaia lanceolata sp. nov. § Huaglaia.

Arbor parva vel arbuscula, ramulis foliis junioribus inflorescentiisque
densissime brunneo-lepidotis; foliis alternis, imparipinnatis, foliolis 7
ad 11, anguste lanceolatis, membranaceis vel chartaceis, apice sensim
acuminatis, basi inaequilateralibus, acutis vel acuminatis, in sicco pallidis,
nitidis, utrinque praesertim subtus plus minus brunneo-lepidotis; pani-
culis axillaribus terminalibusque, foliis subaequalibus vel brevioribus,
diffusis, multifloris ; floribus minutis, 5-meris, racemose dispositis, breviter
pedicellatis, tubo stamineo libero.

A small tree or shrub (2 m high according to the collector), all parts
more or less brown-lepidote, the branchlets, inflorescence and young
leaves densely so. Branches terete, gray or brownish, ultimately gla-
brous. Leaves alternate, 20 to 30 cm long, the rachis at first lepidote,
ultimately glabrous or nearly so. Leaflets 7 to 11, alternate, or the
upper ones opposite, narrowly lanceolate, 8 to 12 em long, 1.5 to 2 cm
wide, rather pale and somewhat shining when dry, the young ones
densely brown-lepidote on both surfaces, the mature ones ultimately nearly
glabrous, the apex narrowly and gradually acuminate, the base inequi-
lateral, acute or acuminate; nerves 15 to 20 on each side of the midrib,
indistinct; petiolules about 2 mm long. Panicles axillary and terminal,
about 15 em long, pyramidal, diffuse, branched from the base, all parts
densely brown-lepidote. Flowers 5-merous, small, raeemosely disposed
on the ultimate branchlets, very numerous, their pedicels about 1 mm
long. Calyx-lobes lepidote, rounded, 0.5 mm long. Petals glabrous,
orbicular or orbicular-elliptic, about 1 mm long. Staminal-tube globose,
glabrous, free from the petals, crenate at the apex; stamens 5, inserted
at about the middle of the tube, included-

Luzon, Province of Nueva Vizeaya, Amucuean, near Bayombong, Bur. Sci. 8141
Ramos, May 13, 1909, in forests along streams.

A species manifestly allied to Aglaia angustifolia (Miq.) C. DC., but abundantly
distinet, well characterized by its narrowly lanceolate leaves.

»

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 185

DYSOXYLUM Blume.

Dysoxylum venosum sp. nov. § Hudysoxylum.

Arbor, partibus junioribus subtus foliis inflorescentiisque molliter
puberulis; foliis alternis, imparipinnatis, foliolis 11, oblongis vel ellip-
. tico-oblongis, acuminatis, nitidis, supra glabris, in sicco pallidis, nervis
utrinque circiter 10, supra impressis, subtus prominentibus ; inflorescentits
axillaribus, brevibus, spiciformibus ; floribus 4-meris, tubo stamineo libero,
ovario pubescente.

A tree of medium size, the branches terete, grayish, glabrous, the
branchlets, inflorescence, rachis and under surface of the leaflets softly
pale-olivaceous-puberulent. Leaves about 45 cm long, alternate; leaflets
11, the lower ones alternate, the upper opposite, oblong or elliptic-oblong,
“10 to 17 em long, 4.5 to 7 em wide, rather pale when dry, shining, the
upper surface glabrous, or the midrib often puberulent, apex acuminate,
base acute; nerves about 10 on each side of the midrib, impressed above,
beneath prominent, anastomosing, the reticulations lax, obscure; petio-
lules puberulent, about 5 mm long. Inflorescences in the upper axils,
and in the axils of fallen leaves, solitary, spiciform, unbranched, 4 cm
long or less, the pedicels very short. Flower-buds globose, 4 mm in
diameter, the calyx shortly 4-toothed, puberulent outside, the teeth
triangular-ovate, 1 mm long or less. Petals 4, densely gray-puberulent
outside, oblong or oblong-ovate, 4 mm long. Staminal-tube cylindric,
free, glabrous, 3 mm long, minutely crenate. Anthers 8, included. Disk
tubular, 1 mm long, obscurely denticulate, glabrous outside, pubescent
within. Ovary ovoid, densely pubescent, 4-celled; style, including the
stigma, about 2 mm long.

Luzon, Province of Cagayan, Mount Cura, For. Bur. 16839 Curran, March,
1909, altitude about 200 m.

A species much resembling D. turczaninowti C. DC., but distinguished by its

very strongly veined leaves, most parts densely puberulent, and many other
characters.

Dysoxylum biflorum sp. nov. § Hudysoxylum.

Arbor glabra vel subglabra, partibus junioribus exceptis; foliis al-
ternis, abrupte pinnatis, 3-jugatis, foliolis elliptico-ovatis, subcaudato-
acuminatis, basi acuminatis, subtus in venarum axillis glandulosis bar-
batisque; inflorescentiis Pei depauperato-paniculatis, pedunculis
bifloris; floribus longe pedicellatis, 4-meris; calycibus pyriformibus, bre-
viter obscure crenatis; petalis 4, glabris, tubo stamineo libero; ovario
glabro vel subglabro, 4-loculare.

A tree, nearly glabrous except the innovations which are somewhat
pubescent. Branches terete, lenticellate, slender, brown. Leaves al-
ternate, 20 em long, equally pinnate, 3-jugate; leaflets opposite, elliptic-
ovate to oblong-ovate, chartaceous or somewhat coriaceous, glabrous and
shining on the upper surface, 6 to 10 cm long, 2.5 to 3.5 em wide, the

186 ‘MERRILL.

apex rather abruptly subcaudate-acuminate, acumen abgut 1 cm long,
blunt, base acuminate; nerves about 8 on each side of the midrib, prom-
inent beneath, impressed above, anastomosing, the reticulations lax,
obscure, with a barbate gland in the axil of each nerve where it leaves
the midrib; petiolules 3 to 5 mm long. Inflorescences few, axillary,
each consisting of a 5 to 6 em long peduncle, bearing at its apex two
long-pedicelled flowers, each pedicel subtended by a small bract about 2
mm long, and each calyx subtended by a smaller, but similar bracteole ;
pedicels 1 cm long. Calyx pear-shaped, 4 mm long, 3.5 mm in diameter
above, the mouth with four broad, obscure, rounded teeth. Petals 4, in
bud elliptic, 3 mm long. Staminal-tube free, about 2 mm long, cylindric,
denticulate; stamens 8, the anthers 1 mm long, included. Disk tubular,
1 mm long, free. Ovary ovoid, 4-celled, each cell 1-ovuled, glabrous or
with few scattered hairs, including the short style 2 mm in length.

Luzon, Province of Isabela, Cabagan River, For. Bur. 18563 Alvarez, April 22,
1909, altitude about 100 m.
A species well characterized by its two-flowered inflorescences.

MALPIGHIACEZ.
HIPTAGE Gaertn.

Leaves not gradually narrowed upward, obtuse or rounded; uniformly and dis-
tinctly reticulate on both surfaces; flowers small.............-....-- l. H. reticulata
Leaves more or less gradually narrowed upward to the acuminate or acute apex.
Leaves pubescent, often densely so.
Carpels small, less than 1.5 em long, including the wings; leaves less than
2.5 em in width.
An erect tree or shrub; leaves glabrous on the upper surface, densely
pubescent with pale hairs beneath; racemes many flowered.
2. H. pubescens
Scandent; leaves pubescent on both surfaces; racemes few-flowered.
3. H. curranit
Carpels large, including the wings reaching 4.5 em in length; leaves 3.5 to
0) 6 a2 (0 (- eee ee RON Pree E: Semen So ee ol oe 4. H. tetraptera
Leaves glabrous.
Carpels large, including the wings reaching a length of 8 em.
5. H. macroptera
Carpels small, including the wings not exceeding 2.5 em in length.
Leaves broad, rather abruptly acuminate, the lateral nerves 4 or 5; central
wing of the carpels 1 to 1.4 em broad.............-------00----+--- 6. H. cumingit
Leaves relatively narrow, gradually narrowed upward, the lateral nerves 6
or 7; central wing of the carpels less than 1 em broad.

Leaves ample, usually exceeding 10 em in length...........-... 7. H. javanica
Leaves small, Gem) long or Lessis5:2c.tses eeemeeeentee st ean e enc 8. H. luzonica

1. Hiptage reticulata sp. noy. ;

Frutex scandens (?); foliis elliptico-oblongis, coriaceis, utrinque gla-
bris, nitidis, reticulatis, obtusis vel rotundatis, nervis utrinque cireiter
8: racemis circiter 8 cm longis, leviter pubescentibus, compositis ; floribus

vix 1 em diametro.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 187

An erect or scandent shrub or a tree. Leaves elliptic-oblong, coria-
ceous, glabrous, shining, about 6 em long, 2.5 cm wide, abruptly narrowed

at both base and apex which are obtuse or rounded; nerves about 8 on
each side of the midrib, anastomosing, the reticulations rather close,
distinct on both surfaces. Racemes compound, about 8 cm long, shghtly
pubescent, the pedicels about 10 mm long, scattered along the rachis, each
subtended by a 1 mm long bracteole, and bearing-at about the middle
an additional bracteole subtending a sessile or shortly pedicelled bud.
Calyx-gland very prominent, 1.5 to 2 mm long. Sepals 1.5 to 2 mm
obtuse. Petals 3 to 4 mm long, rounded.

long,

Luzon, Province of Zambales, Vidal 2243 in Herb. Kew.

This form has as yet not been rediscovered in the Philippines, and is described
from the single specimen preserved in the Kew herbarium. It is well charac-
terized by its elliptic-oblong, glabrous, shining, reticulate, blunt leaves, and by its
comparatively small flowers.

2. Hiptage pubescens sp. nov.

Arbor parva usque ad 5 m alta, ramulis, subtus foliis, inflorescentiisque
dense pallide adpresse sericeo-pubescentibus; foliis coriaceis, elliptico-
oblongis vel oblongo-lanceolatis, acuminatis, basi rotundatis vel acutis;
racemis axillaribus foliis subaequalibus vel longioribus: carpellis vix 1.5
cnr longis.

A small tree 4 to 5 m high (9 m according to Ramos). Branches
brown or gray, terete, lenticellate, glabrous, the young branchlets densely
and pale-silky-pubescent. Leaves opposite, coriaceous, elliptic-oblong to
oblong-lanceolate, 5 to 8 cm long, 1.5 to 2.5 em wide, glabrous and
shining above, on the lower surface densely covered with pale, appressed,
silky hairs; the base rounded or sometimes somewhat acute, the apex
distinctly, often strongly and gradually acuminate; nerves 7 or 8 on each
side of the midrib, not prominent, obscurely anastomosing, obscured on
the lower surface by the pubescence, but the hairs sometimes rubbing
off, the veins then appearing brown in contrast to the pale surface of
the leaf; petioles 2 to 3 mm long, pubescent. Racemes axillary, solitary,
about as long as the leaves, sometimes crowded in. the upper axils and
simulating a terminal inflorescence, many-flowered, densely silky-pubescent
with pale appressed hairs, the pedicels 5 to 7 mm long. Flowers yellow
or red, about 1 em in diameter when open. Sepals obtuse, the gland
prominent. Petals 5 to 6 mm long, obtuse, pubescent. Fruit of two
carpels, somewhat pubescent, the central wing of each carpel 10 to 12
mm long, about 5 mm wide, rounded or obtuse, the two lateral ones
similar but less than one-half as long, and truncate or rounded.

Luzon, Province of Abra, Mount Paraga, Bur. Sci. 7257 Ramos, February,
1909 (type): Lepanto-Bontoe, For. Bur. 11263 Klemme, February, 1908, altitude
about 1,200 m: Province of Ilocos Norte, Mount Piao, lor. Bur. 13979 Merritt &

Darling, altitude about 1,000 m.
This species is readily recognizable by its comparatively small leaves and small

188 MERRILL.

fruits, but especially by the dense, pale, appressed, silky pubescence on the
inflorescence, branchlets and lower surfaces of the leaves. It is remarkable in the
genus in that it is erect and arborescent, not scandent. The trunk-diameter is
_ given by the various collectors as from 8 to 30 cm.

Var. lanceolata var. nov.
A typo differt foliis angustioribus, lanceolatis, circiter 1 cm latis.

Luzon, Province of Ilocos Norte, Badoc, For. Bur. 13955 Merritt & Darling,
altitude about 65 m, locally known as pangardisin; near Vintar, altitude 700 m,
For. Bur. 13943 Merritt & Darling: Province of Ilocos Sur, For. Bur. 5632
Klemme. ? :

In general appearance, pubescence, flowers, etc., quite the same as the species,
differing only in its narrower and lanceolate leaves. The fruits are unknown.

3. Hiptage curranii sp. nov.

Frutex scandens, omnibus partibus pubescens ; foliis coriaceis, elliptico-
oblongis vel late oblongo-lanceolatis, acuminatis, nervis utrinque circiter
5; racemis axillaribus, brevibus, paucifloris; carpellis circiter 1 cm longis.

A scandent shrub, reaching a height of 4 m, in vegetative characters
similar to Hiptage pubescens Merr. Branches terete, slender, becoming
glabrous, dark-colored, often nearly black, scarcely lenticellate, the branch-
lets densely pale-pubescent. Leaves opposite, oblong-elliptic to broadly
oblong-lanceolate, coriaceous, 4 to 8 em long, 1.5 to 2.5 em wide, the
base acute, rarely obtuse, the apex gradually and distinctly: acuminate,
the upper surface covered with short, yellowish-brown hairs, the lower
surface very densely pubescent with pale appressed hairs; nerves about
5 on each side of the midrib, curved-ascending, anastomosing, not prom-
inent; petioles densely pubescent, 3 to 4 mm long. Flowers unknown.
Racemes in fruit 2.5 em long or less, densely pubescent with pale hairs,
few-flowered, often only three flowers in a raceme, or reduced to a single
flower. Fruit of two carpels, more or less pubescent, the central wing
of each carpel rather thin, 1 cm long, about 5 mm wide, the lateral ones
similar but less than one-half as long.

Luzon, Province of Zambales, Baquilis River, Mor. Bur. 6951 Curran, May 9,
1907, in the dry river: bed.

Similar in most respects to H. pubescens, differing in being scandent instead of

erect, in its short, few-flowered racemes, and by its leaves being pubescent on both
surfaces, mostly acute at the base, and with fewer lateral nerves.

4. Hiptage tetraptera sp. nov.

Frutex suberectus, vix scandens, omnibus partibus plus minus adpresse
pubescens; foliis coriaceis, ovatis vel anguste ovatis, obscure late acum-
inatis, nervis utrinque 4 vel 5; racemis axillaribus, solitariis, simplicibus ;
carpellis 1 vel 2, alato-cristatis, crista 1 ad 1.5 em longa.

A suberect shrub about 2 m high, scarcely scandent. Branches terete,
pale, densely appressed-pubescent with short, pale hairs. Leaves opposite,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 189

coriaceous, 6 to 8 cm long, 3.5 to 5 cm wide, the upper surface somewhat
appressed-pubescent, especially on the nerves, later becoming subglabrous,
the lower surface rather densely pubescent with short, pale, appressed
hairs, the base rounded or subacute, the apex obscurely and broadly
acuminate; neryes 4 or 5 on each side of the midrib, distinct, curved-
ascending, the reticulations obscure ; petioles pubescent, the apical glands
prominent. Racemes axillary, solitary, 8 to 10 cm long, pubescent, the
pedicels 1 to 2 cm long. Sepals pubescent, obtuse. Petals unknown.
Carpels one or two, somewhat appressed-pubescent, about 8 mm long and
wide, the crest prolonged into a narrow, oblong or oblong-lanceolate
wing, 1 to 1.5 cm long and 3 to 4 mm wide; central wing 3.5 to 4.5
em long, about 1.5 cm wide, rounded, the lateral ones similar and about
one-half as large.

PALAWAN, Separation Point, Merrill 1791, February 18, 1903.

A species recognizable by its pubescent leaves, and especially by its carpel-
crests being prolonged into a manifest wing, making the carpels appear as though
they were four-winged, whence the specific name.

5. Hiptage macroptera sp. nov.

Frutex scandens, inflorescentiis exceptis glaber vel subglaber; foliis
elliptico-oblongis vel ovato-ellipticis, acuminatis, nervis utrinque circiter
6; racemis densis, axillaribus; carpellis 1 vel 2, obscure late carinatis,
vix cristatis, ala media usque ad 7 cm longa.

A scandent shrub, glabrous except the inflorescence. Branches terete,
reddish-brown or grayish, somewhat lenticellate. Leaves chartaceous or
subcoriaceous, elliptic-oblong to ovate-elliptic, 8 to 12 cm long, 3 to 5.5
em wide, glabrous, shining, the base rounded, rarely subacute, the apex
prominently and usually abruptly acuminate; nerves about 6 on each
side of the midrib, prominent, curved upward, obscurely anastomosing,
the reticulations not prominent; petioles about 7 mm long, the leaf-base
with usually two distinct glands at the junction with the petiole.
Racemes axillary, solitary, densely rather many-flowered, pubescent, in
anthesis 4 to 5 cm long, longer in fruit. Flowers pinkish-white, their
pedicels 10 to 12 mm long, longer in fruit. Sepals elliptic, rounded,
about 4 mm long, 2.5 mm wide, pubescent. Petals 10 to 12 mm long,
prominently fimbriate. Carpels one or two, slightly pubescent, with a
broad low ridge along the top but scarcely crested, the central wing 6
to 7 cm long, usually about 1.5 cm wide, somewhat narrowed at both
ends, apex obtuse, the lateral wings similar and about one-half as long.

Mindanao, Lake Lanao, Mrs. Clemens s. n., May, 1907, the specimen in fruit
(type), and also no. 1056, same date, in flower:

This species grows in thickets and forests along the margin of the lake, the
young leaves and rather prominent fruits being red in color. It is manifestly
allied to H. benghalensis (L.) O. Ktze., differing especially in its much larger
wings.

190 MERRILL.

6. Hiptage cumingjii sp. nov.

Hiptage madablota Vid. Phan. Cum. Philip. (1885) 99, non Gaertn.

Scandens, inflorescentiis exceptis glabra vel subglabra: foliis coriaceis,
ovato-ellipticis vel oblongo-ovatis, usque ad 9 em longis, basi acutis vel
rotundatis, apice acuminatis, nervis utrinque 4 vel 5; racemis simplicibus,
foliis subaequilongis; carpellis 2 vel 3, vix vel obscure cristatis, ala media
1.5 ad 2 em longa.

A scandent shrub, glabrous or nearly glabrous except the inflorescence.
Branches terete, lenticellate, rather slender, usually reddish-brown, the
branchlets more or less pubescent soon becoming glabrous. Leaves coria-
ceous, ovate-elliptic to oblong-ovate, 5.5 to 9 em long, 3 to 5 em wide,
shining above, the base acute or rounded, the apex distinctly and often
abruptly acuminate; lateral nerves 4 or 5 on each side of the midrib,
anastomosing, the reticulations not distinct; petioles 5 to 7 mm _ long.
Racemes axillary, solitary, mostly in the upper axils and simulating a
terminal inflorescence, 5 to 8 em long, pubescent, the flowers numerous.
the pedicels about 1 cm long, somewhat elongated in fruit, the bracteoles
near the middle about 2 mm in length. Sepals oblong, rounded, about 3
mm long. Petals 6 to 7 mm long, pubescent. Carpels 2 or 3, somewhat
appressed-pubescent, not or very obscurely crested, the central wing
broadly oblong-elliptic or obovate-elliptic, 1.5 to 2 em long, often nearly
1.4 cm wide, the lateral ones about one-half as long.

Luzon, Province of Pangasinan, Cuming 971 (type).

I am also disposed to refer here Por. Bur. 6732 Merritt, from near Pinamalayan,
Mindoro, which differs from Cuming’s specimen in having the leaves somewhat
pubescent beneath, and in its very slightly smaller fruits, and Bur. Sci. 753
Foxworthy from Mount Victoria, Palawan, the latter very closely matching the
type.

Hiptage cumingii is manifestly allied to H. benghalensis (1.) O. Ktze. (H.
madablota Gaertn.), but differs especially in its smaller, fewer-nerved leaves,
smaller flowers, its crestless carpels, and much smaller wings.

7. Hiptage javanica Bl. Bijdr. (1825) 224; Mig. Fl. Ind. Bat. 17 (1858)
586; Hochr. Pl. Bogor. Exsice. no. 32. ;

MINDANAO, District of Cotabato, near Fort Reina Regente, For. Bur. 3944
Hutchinson.

This specimen closely matches a very full series of specimens representing
Blume’s species, received from the Botanic Garden at Buitenzorg, differing in
having some of the leaves slightly wider. The species has not previously been
recorded otherwise than from Java.

There are in this herbarium four specimens, all with flowers, from the Prov-
ince of Rizal, Luzon, that previously have been referred to H. madablota Gaertn.,
and the duplicates distributed under that name. This material is manifestly not
specifically the same as Gaertner’s species, and may possibly be referable to
H. javanica Bl., although there are some manifest differences in vegetative char-
acters, especially in the much more obscure reticulations. Otherwise the speci-
mens very closely resemble HH. javanica Bl., but in the absence of fruiting material,
they are not at present definitely referred to that species. The specimens are

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 191

Merrill 1704, 5046, and For. Bur. 420, 2660 Ahern’s collector. H. madablota Vid.
Sinopsis Atlas (1883) @. 22, f. A. (non Gaertn.) manifestly represents the same
form as the four specimens above mentioned.

8. Hiptage luzonica Merr. in Govt. Lab. Publ. ( Philip.) 35 (1905) 33, Philip.
Journ. Sei. 1 (1906) Suppl. 74.

Luzon, Province of Bataan, Mount Mariveles, Whitford 1148.

This species is known only from the original collection, and the type is possibly
only a dwarfed state of the Rizal form discussed above under H. javanica BI.
A full series of specimens will be necessary definitely to settle this point. So
far as our material goes, H. luzonica is distinguishable by its small leaves.

HIpTAGE MADA'BLOTA Gaertn. (=H. benghalensis (L.) O. Ktze.) has been cred-
ited to the Archipelago by various authors, but I have seen no Philippine material
that I consider to be referable to that species. The plant so figured by Vidal
in his ‘“‘Sinopsis Atlas” unquestionably represents the Luzon form discussed under
H. javanica, while the plant so identified by him in his ‘“Phanerogamae Cumin-
gianae Philippinarum” has above been made the type of a new species, H. cu mingti.
The form so credited to the Philippines by F.-Villar in the “Novissima Appendix”
is doubtless, for most part, the same as that figured by Vidal, as the specimens
F.-Villar examined came from the Province of Manila (=Rizal).

Triopteris jamaicensis Blanco Fl. Filip. (1837) 379, ed. 2 (1845) 207, non
Linn., is manifestly Hiptage, although not H. madablota Gaertn., where it was
referred by F.-Villar. It is probably the form figured by Vidal, mentioned above,
as this is apparently the only species of the genus that is at all common in the
region from which Blanco secured most of his material.

EUPHORBIACE.
ACALYPHA Linn.

Acalypha grandibracteata sp. nov.

Species A. stipulaceae valde affinis, differt foliis latioribus, basi cordatis
vel subcordatis, bracteis multo majoribus, usque ad 1 ad 2 cm longis.

A shrub or small tree, slightly puberulent or pubescent. Branches
pale or reddish-brown, puberulent, sometimes stout and thickened.
Leaves broadly ovate to oblong-ovate, chartaceous or submembranaceous,
12 to 20 em long, 7 to 15 em wide, with minute, scattered, white pustules
on both surfaces, and with very few, scattered, long hairs, the margins
regularly and rather finely crenate-serrate, the apex acuminate, the base
broad and cordate or subcordate, palmately 7- or 9-nerved from the base ;
petioles 20 cm in length or less ; stipules linear-lanceolate, long-acuminate,
1.5 to 2 cm long. Staminate and pistillate spikes on the same plant, or
apparently more often on separate plants, the staminate ones dense, eylin-
dric, pubescent, often 20 cm long, about 3 mm in diameter, the flowers
3. or 4-merous. Pistillate spikes peduncled, stout, 20 cm long or less,
about 2 cm in diameter, the bracts broadly ovate, acuminate, toothed,
about 1 cm long, the lowermost ones sometimes 2 cm in length, more
or less appressed-hirsute on the back, the pistillate flowers solitary in
the axil of each bract. Ovary hirsute; styles nearly 3 mm long, split

192 MERRILL.

into several, filiform, elongated lobes. Capsule about 2 mm long, hirsute,
the seeds elliptic-oblong, 1.2 mm long, smooth and glabrous.

BATANES ISLANDS, Batan, Santo Domingo de Basco, Bur. Sci. 3607 Fénia
(type), with staminate and pistillate spikes on the same branch, Bur. Sci. 3206
Mearns, May, 1907, with pistillate spikes. A narrower-leaved form is apparently
represented by Bur. Sci. 4084 Fénix, from Camiguin Island, Babuyanes, locally
known as ajas.

These specimens were previously referred by me to Acalypha stipulacea Klotz.,
which they closely resemble in many respects, differing especially in the points
noted in the diagnosis above.

Acalypha australis Linn. Sp. Pl. (1753) 1004; Hemsl. in Journ. Linn. Soe.
Bot. 26 (1894) 437.

Luzon, Province of Cagayan, Bur. Sci. 7800, 7869 Ramos, April, 1909.

Not previously reported from the Philippines; Manchuria and Japan to

southern China.
ALCHORNEA Muell.-Arg.

Alchornea sicca (Blanco) comb. noy.

Excoecaria sicca Blanco Fl. Filip. (1837) 787, ed. 2 (1845) 542, ed. 3, 3:94;
Naves ll. ¢.\ ed. 3; pl. S07.

Stipellaria parviflora Benth. in Hook. Journ. Bot. & Kew. Miscel. 6 (1854) 4.

Alchornea parviflora Muell.-Arg. in Linnaea 34 (1865) 168, DC. Prodr.
15 *:902; Vid. Phan. Cuming. Philip. (1885) 144, Rev. Pl. Vase. Filip. (1886)
244; F.-Vill. Nov. App. (1880) 194. :

Alchornea mollis F.-Vill. 1. ¢., non Muell.-Arg.

“Acalypha tiliaefolia Muell.-Arg.”; Vid. Rev. Pl. Vase. Filip. (1886) 244.

Luzon, Province of Rizal, Bur. Sci. 3334, 5220 Ramos, Vidal 592, 1710 (herb.
Kew.) , Loher 4667: Province of Laguna, Elmer: Province of Pampanga, For. Bur.
18314 Curram. Necros, Cuming 1800.

Blanco’s description unmistakably applies to this species, and his name being
the earliest one is here adopted. Cuming’s specimen was from Negros according
to his own list of localities. The enumeration of “Acalypha tiliaefolia Muell.-
Arg.” by Vidal seems to have been a slip for “Alchornea tiliaefolia Muell.-Arg.”’
There is no such species as “Acalypha tiliaefolia Muell.-Arg.”

DIMORPHOCALYX Thwaites.

Dimorphocalyx luzoniensis sp. nov.

Arbor glabra, dioica, circiter 12 m alta; foliis alternis, chartaceis,
elliptico-ovatis, acuminatis, integris vel distanter obscureque denticulatis,
nervis utrinque 10 ad 12; inflorescentiis axillaribus, racemoso-cymosis,
quam folia brevioribus ; floribus masculinis circiter 8 mm longis, stamini-
bus 15, filamentis in columna connatis sed exterioribus elongatis, fere
liberis; floribus femineis albis, 2.5 em diametro, sepalis liberis, accres-
centibus.

A glabrous dioecious tree about 12 m high. Branches slender, terete,
grayish-brown, somewhat lenticellate. Leaves alternate, elliptic-ovate,
chartaceous, grayish or brownish and somewhat shining when dry, 10 to
15 cm long, 4 to 7 em wide, the base rounded or somewhat acute, the
apex sharply acuminate, the margins entire or distantly denticulate with

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 193

very small subglandular teeth ; petioles 1 to 3 em long; lateral nerves 10
to 12 on each side of the midrib, rather distinct, the reticulations lax.
Inflorescence axillary, shorter than the leaves, of racemosely arranged
cymes, the staminate and pistillate similar. Pistillate flowers white,
ample, their pedicels 7 to 10 mm long. Calyx of five free sepals, im-
bricate, elliptic-ovate, rounded, often slightly retuse, glabrous, 1.2 to 2
cm long, 6 to 12 mm wide, reticulate, apparently persistent. Petals 5,
imbricate, broadly elliptic-ovate, about 10 mm long, 8 mm wide, apex
rounded, not reticulate or nerved. Disk small. Staminodes none.
Ovary glabrous, longitudinally 3-suleate, the lobes rounded, 3-celled, each
cell with a single ovule; styles 3, free, about 5 mm long, each cleft
half way to the base into two divergent arms. Staminate flowers smaller
than the pistillate ones. Calyx 5 to 6 mm long, cleft into five elliptic-
oblong, imbricate, obtuse lobes about 3 mm long and 2 to 2.5 mm wide.
Petals 5, free, imbricate, oblong-elliptic, rounded, 7 to 8 mm long, 4 mm
wide. Disk-glands 5, prominent, white, 1.5 to 2 mm long. Stamens
15, the filaments all more or less united into a column, the interior ones
very short, the outer five longer and nearly free, these outer ones about
3.5 mm long, the free portions of the inner ones progressively shorter ;
anthers introrse. Rudimentary ovary none.

Luzon, Province of Laguna, Los Baiios, For, Bur. 11907 Tamesis, January 22,

1910, pistillate flowers; same locality, Bautista s. n., February 11, 1904, staminate
flowers. Borders of clearings, altitude about 100 m.

AQUIFOLIACE.
ILEX Linn.

Ilex foxworthyi sp. nov. § Thyrsoprinus, Indico-Malaicae.

Arbor inflorescentiis exceptis glabra, circiter 8 m alta; foliis ovatis
vel elliptico-ovatis, coriaceis, usque ad 9 cm longis, nitidis, subtus obscure
minutissime dense puncticulatis et punctis majoribus sparsis intermixtis,
nervis utrinque circiter 7; racemis axillaribus, solitariis, in alabastro
circiter 3 cm longis, puberulis, floribus femineis 5- vel 6-meris.

A tree about 8 m high, glabrous except the inflorescence. Branches
gray, terete, wrinkled when .dry, not lenticellate. Leaves ovate to
elliptic-ovate, coriaceous, shining, brown or olivaceous when dry, and
slightly paler beneath, the margins recurved, entire, 4 to 9 em long,
2.5 to 5 em wide, the lower surface very minutely, obscurely, and densely
puncticulate, and with scattered larger points, the latter not prominent,
the apex shortly blunt-acuminate, base rather broad, acute; nerves
about 7 on each side of the midrib, not prominent, slender, obscurely
anastomosing, the reticulations lax, about as distinct on the upper sur-
face as on the lower; petioles 1 cm long or somewhat less on the smaller
leaves. Pistillate flowers racemose, the racemes axillary, sqlitary, simple,
in bud 3 em long or less, puberulent, some flowers also occurring solitary

194 MERRILL.

or in pairs on the growing branchlets, the pedicels puberulent, 3 to 4
mm. long, each subtended by a triangular-ovate, acute or acuminate,
puberulent, 1 mm long bracteole. Pistillate flowers (in bud), 5- or
6-merous, the buds globose, the calyx-segments elliptic, rounded, puberu-
lent, about 2 mm long, 1.2 mm wide, imbricate. Corolla (immature)
nearly 3 mm long. Staminodes 1 mm long, bearing imperfect anthers.
Ovary not compressed, subglobose, 5- or 6-celled, the stigma nearly as
broad as the ovary, somewhat sulcate. Staminate flowers and fruits
unknown.

Luzon, Province of Tayabas, Mount Banajao, Bur. Sci. 2416 Foxworthy,
Mareh 24, 1907, altitude about 1,450 m.

This species is apparently allied to /lex malaccensis Loesener, of the Malay
Peninsula and Borneo, but its leaves with the lateral nerves distinctly visible
on the upper surface, acute at the base, and its 5- or 6-celled ovaries are ap-
parently sufficiently distinctive to warrant the description of the present form
as new.

It is also allied to Jleax haleonensis Merr., but is distinguished at once by its
puberulent racemes.

ICACINACE.
URANDRA Thwaites.

Urandra hallieri sp. noy.

Arbor glabra vel subglabra, circiter 20 m alta; foliis alternis, coriaceis,
oblongo-ellipticis vel oblongis, breviter obtuse acuminatis, in sicco pallidis,
nitidis, nervis utrinque circiter 15, tenuibus, obscuris; cymis peduncu-
latis, axillaribus, solitariis, quam petiolus paullo longioribus, floribus
5-meris, calyce truncato, connectivo antherarum longe barbato.

A tree about 20 m high, glabrous, or the younger branchlets and
inflorescence more or less puberulent. Branches terete, brownish or
olivaceous, smooth or slightly wrinkled when dry. Leaves oblong-elliptic
to elliptic, coriaceous, 10 to 16 em long, 5 to 7 em wide, pale and
somewhat shining when dry, the apex shortly and abruptly blunt-acum-
inate, the base acute or slightly acuminate, the margins entire, often
slightly revolute; midrib prominent, the lateral nerves about 15 on each
side of the midrib, slender, obscure on both surfaces, the primary ones
hardly more prominent than are the secondary ones, the reticulations
faint on the upper surface, obsolete or subobsolete beneath; petioles
about 2 cm long. Cymes axillary, solitary, peduncled, about 3 em in
diameter, the peduncles about as long as the petioles. Flowers sessile in
fascicles of three each at the tips of the umbellately disposed primary
branches, 4-merous. Calyx shallowly cup-shaped, slightly puberulent,
truncate, about 2 mm long, 3 mm in diameter, slightly puberulent
outside and on the margins, strongly wrinkled when dry. Petals 4,
narrowly oblong, 5.5 mm long, 1.8 mm wide, valvate, near the base
the margins slightly coherent, the apex acute or obtuse, appendaged

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 195

inside. Stamens 4, the filaments about 4.5 mm long, flattened, 1 mm
wide and of about the same width throughout, the anterior face at the
base of the anther bearded with long hairs; anthers about 1 mm long,
the back and apex of the connective very densely bearded with numerous,
pale, rather stout, soft hairs about + mm long, the tips of the hairs
somewhat club-shaped. Disk prominent, truncate. Ovary conical, gla-
brous, strongly wrinkled, tipped by the very short style.

BASILAN, Hallier s. n., January, 1904 (type). Mrinpanao, District of Zam-
boanga, Port Banga, For. Bur. 9176 Whitford & Hutchinson, December 9, 1907,
in dipterocarp forests at an altitude of about 20 m. :

A species in gross characters very similar to Urandra apicalis Thwaites, of
Ceylon, but distinguished at once by its truncate calyx.

Urandra elliptica sp. nov.

Arbor glabra circiter 30 m alta; foliis coriaceis, ellipticis vel late
ellipticis, utrinque late rotundatis vel apice abrupte brevissime acumi-
natis, coriaceis, nitidis, circiter 10 cm longis, nervis utrinque 4 vel 5;
fructibus globosis, 1.5 cm diametro.

A glabrous tree about 30m high, the trunk 90 em in diameter. Branches
terete, smooth, brown-olivaceous or reddish-brown. Leaves alternate,
coriaceous, shining and of about the same color on both surfaces, elliptic
or broadly elliptic, 8 to 11 cm long, 6 to 9 em wide, equally and very
broadly rounded at both ends, or the apex very abruptly and_ shortly
acuminate; nerves + or 5 on each side of the midrib, distant, curved-
ascending and faintly anastomosing, the reticulations lax, not prominent ;
petioles 1 to 1.5 em long. Inflorescence (young) axillary, peduncled,
of three or four spicately arranged, short branches, each branch with
about 6 pairs of densely imbricated, distichous, broadly ovate, brown, 1 mm
long bracteoles. Flowers unknown. Fruit globose, yellow and somewhat
fleshy when fresh, about 1.5 cm in diameter, one or two developing from
each branch of the inflorescence, the rachis slightly elongated, thickened,
the peduncles of the fruits stout, 1 to 1.5 em long.

Luzon, Province of Bataan, Duale, For. Bur. 20003 Topacio, September 14,

1909, in semi-open flat country, altitude about 100 m.
Well characterized by its broadly elliptic leaves.

SABIACE.
MELIOSMA Blume.

Meliosma reticulata sp. nov.

Arbor parva, circiter + m alta, ramulis, foliolis ad nervos, petiolis,
paniculisque dense ferrugineo-villosis; foliis imparipinnatis, foliolis 5
ad 11, ellipticis vel oblongo-ellipticis, coriaceis, utrinque valde reticulatis,
apice rotundatis vel abrupte brevissime acuminatis, margine distanter
denticulatis, nervis utrinque 8 ad 10, prominentibus; paniculis termina-
libus, foliis subaequalibus ; floribus numerosis, subsessilibus.

196 MERRILL.

A small tree about 4 m high. Ultimate branches rather stout, dark-
brown, glabrous or nearly so, nearly 1 cm in diameter, the growing parts
smaller and densely ferruginous-villous. Leaves alternate, 40 cm long
or less, the petiole, rachis, petiolules, and midrib and nerves on both
surfaces of the leaflets densely ferruginous-villous; leaflets 5 to 11,
elliptic to oblong-elliptic, coriaceous, 6 to 11 cm long, 3 to 6.5 cm wide,
the base broad, rounded, the apex also broad, rounded or very abruptly
and shortly acuminate, the margins in the lower half entire, above with
few, small, scattered teeth; nerves 8 to 10 on each side of the midrib,
prominent, anastomosing, and with the rather lax primary reticulations
impressed on the upper surface, prominent beneath; petiolules 12 mm
long or less. Panicles terminal, as long as the leaves, the branches few,
the lower ones often 15 to 20 em long, all parts densely ferruginous-
villous. Flowers white, subsessile, the bracteoles about 2 mm long.
Calyx-segments 4 or 5, ovate to suborbicular, the outer ones smaller than
the inner and more or less ferruginous-villous. Three larger petals
orbicular, about 3 mm in diameter, the two smaller ones reduced to mere
scales less than 1 mm long and adnate to the filaments. Fertile stamens
2, the filaments less than 1.3 mm long.

Luzon, Province of Benguet, Losod, Bur. Sci. 5594 Ramos, December, 1908.
I am inclined to refer here also For. Bur. 15803 Curran, from the Kuyapa District
in Benguet, but the specimen is with nearly mature fruit, and differs from the
type in being very much less pubescent, possibly due to development. The
fruits are narrowly obovoid, slighly compressed, and about 6 mm long. The
two specimens in other characters than the pubescence are very similar.

The species is well characterized by its dense ferruginous pubescence which

extends even to the nerves on both sides of the leaflets, and by its very strongly
reticulate leaflets.

VITACEAE.

LEEA Linn.

Leea quadrifida sp. nov.

Arbuscula circiter 1 m alta; foliis pinnatis, foliolis circiter 10, oblongis,
acuminatis, nervis utrinque circiter 12, subtus glandulis numerosis, brun-
neis, parvis sed prominentibus, conspersis; cymis brevibus, floribus con-
gestis, 4-meris.

A shrub about 1 m high. Branches brown, somewhat pubescent.
Leaves alternate, simply pinnate, 40 to 50 em long, the rachis prominently
longitudinally suleate, the petiole not dilated at the base, when very
young brown-puberulent. Leaflets oblong, firmly chartaceous, 14 to 20
em long, 5 to 6 em wide, the apex rather prominently acuminate, the
base rounded, very slightly inequilateral, the margins obscurely and
distantly crenulate or denticulate, the upper surface smooth, glabrous,
grayish and somewhat shining when dry, the lower surface brown, pubes-
cent or puberulent on the nerves and midrib, the whole surface with
numerous, small, elevated, brown glands distinctly visible to the naked

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 197

eye; lateral nerves about 12 on each side of the midrib, prominent, some-
what ascending, nearly straight, anastomosing near the margin, the reticu-
lations distinct on the lower surface; petiolules about 1 cm long.
Inflorescence axillary, brown-pubescent, the stipe about 1 cm long, very
stout, bearing about 4 primary branches 5 to 6 cm long, branched near
the apex, and bearing numerous, subsessile, congested, white flowers.
Calyx 4 to 5 mm long, slightly pubescent, brown when dry, 4-toothed,
the teeth broadly ovate, 1.5 to 2 mm long, with few scattered glands.
Corolla 6 mm long, the lobes 4, reflexed in anthesis, oblong, 3 mm long.
Anthers 4, 2 mm long, connate, inflexed and included in the tube.

Luzon, Province of Laguna, Mount Maquiling, For. Bur. 13309 Tamesis, Sep-
tember 25, 1909, in forests, altitude about 700 m.

A most distinct species, well characterized by its prominently glandular

leaflets, but especially by its 4-merous flowers, in the latter character differing
from all species of the genus known to me.

MALVACEA,
ABUTILON Tourn.

Abutilon hirtum (Lam.) G. Don Gen. Syst. 1 (1831) 503.

Sida hirta Lam. Encycl. 1 (1783) 7.

Abutilon graveolens W. & A. var. hirtum Mast. in Hook. f. Fl. Brit. Ind. 1
(1874) 327.

MINDANAO, District of Zamboanga, Mrs. Clemens 675, Hallier s. n.

Widely distributed in the Tropics.

This is probably the form credited to the Philippines by F.-Villar? as A.
graveolens W. & A. If the forms described as Abutilon graveolens W. & A., and
A. hirtum (Lam.) G. Don, are varietally distinct, as several authors consider
them, it is believed that the specific name should be that of the one first described.

GUTTIFERAE.
CALOPHYLLUM Linn.
Calophyllum gracilipes sp. nov. § Microphyllum.

Arbor glabra circiter 12 m alta, ramis tenuibus, teretibus, ramulis
valde quadrangulatis, circiter 1.5 mm diametro; foliis oblongo-ellipticis,
firmiter membranaceis vel subcoriaceis, usque ad 9 cm longis, nitidis,
subtus pallidis, basi acutis, apice breviter obtuse acuminatis; inflores-
centiis axillaribus, solitariis, racemis 3-floris, pedicellis tenuibus, 2 ad 3
em longis, floribus 4-meris, circiter 1.5 em diametro.

A tree about 12 m high, glabrous throughout except the short, brown-
puberulent terminal buds. Branches slender, terete, brown or olivaceous,
the branchlets strongly 4-angled, slender, about 1.5 mm in diameter.
Leaves oblong-elliptic, 5 to 9 em long, 2 to 3.5 cm wide, firmly mem-
branaceous or subcoriaceous, shining when dry, the lower surface paler

Nov. App. (1880) 23.

198 MERRILL.

than the upper, the base acute, the apex shortly and bluntly acuminate ;
nerves very numerous, densely disposed, about as prominent on one sur-
face as on the other; petioles 5 to 10 mm long. Racemes axillary,
solitary, 3-flowered, the peduncles slender, about 1 cm long, the pedicels
very slender, 2 to 3 em long, umbellately arranged at the apex of the
peduncle. Flowers white, the outer two sepals broadly ovate, obtuse,
about 5 mm long, the inner two similar but petaloid. Petals 4, elliptic
or elliptic-ovate, about 8 mm long. Stamens indefinite. Ovary glabrous,
globose or oyoid; style 4 mm long.

MinpANAO, District of Zamboanga, Port Banga, For. Bur. 9405 Whitford &
Hutchinson, February 3, 1908, in ridge forests at an altitude of about 600 m;
also represented by Williams 2193, from the Sax River, same district, February
28, 1905.

A species similar in vegetative characters to Calophyllum whitfordii Merr.,
at once recognizable by its 3-flowered, solitary, axillary racemes, and very slender,

elongated pedicels.

Calophyllum racemosum sp. noy. § /nophyllum.

Arbor circiter 11 m alta, glabra, gemmis ferrugineo-puberulis ex-
ceptis; foliis coriaceis, oblongo-ellipticis, usque ad 25 cm longis, nitidis,
basi acutis, apice acuminatis, margine distincte revolutis ; racemis simpli-
cibus, axillaribus, solitariis, usque ad 10 cm longis, floribus circiter 2 em
diametro.

A tree about 11 m high, glabrous except the ferruginous-puberulent
terminal buds which are lanceolate and 1 to 1.5 em long. Branches
stout, terete, somewhat rugose, brown to olivaceous, the branchlets some-
what angled, sulcate. Leaves coriaceous, oblong-elliptic, 15 to 25 em
long, 4 to 7 em wide, shining, when dry about the same color on both
surfaces, or somewhat paler beneath, the apex distinctly and rather
abruptly acuminate, the base acute, the margins rather strongly revolute ;
nerves very numerous, close, about equally distinct on both surfaces ;
petioles rather stout, 1 to 2 cm long. Racemes axillary, solitary, 10
em long or less, simple, each with from 6 to 10 flowers, the pedicels 1
to 2 em long, those of the lower flowers the longer. Sepals orbicular-
ovate, 8 to 10 mm long. Petals 4. Stamens indefinite. Fruit (im-
mature) ovoid, glaucous, smooth, 1 to 1.5 cm long, apiculate.

Leyte, between Dolores and Ormoc, For. Bur. 12620 Rosenbluth (type), Feb-
ruary 26, 1909, in forests, altitude about 100 m. I am disposed to refer here
also the following specimens from Mindanao: Lake Lanao, Camp Keithley, Mrs.
Clemens 1009, a luxuriant form, the racemes forming almost leafless, terminal
panicles, and Williams 2124, 2346, with immature fruits, from the Sax River,
District of Zamboanga.

A species well characterized by being entirely glabrous, except the terminal
buds, the margins of the leaves distinetly revolute, and the flowers arranged in
simple racemes. It is probably as closely allied to Calophyllum blancoi Pl. & Tr.,
as to any other species but is distinguished by the above characters.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 199

Calophyllum amplexicaule Choisy ex Planch. & Triana in Ann. Sci. Nat.
1V 15 (1861) 281; Vesque in DC. Monog. Phan. 8 (1893) 564; Vidal Phan.
Cuming. Philip. (1885) 96, Rev. Pl. Vase. Filip. (1886) 54.

Tovomita pentapetala Blanco Fl. Filip. (1837) 432, ed. 2 (1845) 301, ed. 3,
2:194.

Ochrocarpus pentapetalus F.-Vill. Nov. App. (1880) 17.

Luzon, Province of Ilocos Norte, Cuming 1077 (type number): Province of
Pangasinan, Salasa, For. Bur. 9625 Zschokke: Province of Zambales, For. Bur.
§226, 8229, 8236 Curran & Merritt, Bur. Sci. 4755, 5100 Ramos.

This species is here enumerated chiefly to call attention to the reduction of
Blanco’s Tovomita pentapetala, which was referred by F.-Villar to Ochrocarpus.
Blanco’s material was from the Provinces of Ilocos Norte and Sur, and he speaks
of the plant as being common near the seashore, flowering in December, and
locally known as Pamitlain and Pamitlatin. The specimen collected by Cuming,
on which Calophyllum ampleaxicaule Choisy was based, was also from Ilocos Norte,
according to Cuming’s own list of localities, and not from Albay, as stated by
Vesque; that the former is probably correct is borne out by the fact that the
species is only known from northern Luzon, and has so far never been found
in the south. Blanco’s description applies absolutely to Calophyllum ampleaxicaule
with the exception of the flowers, which he describes as having two sepals, and
five petals. This was undoubtedly an error on his part, due to misconception
of the parts of the flower. This is the only plant known to me that agrees with
Blanco’s description as to its resinous properties, sessile leaves, ete., and even
to the axillary glands mentioned by him, these glands being really the axillary
buds. The specimen from Pangasinan was received under the native name
“Pamitaoyon.”

Blanco’s specific name is hence the oldest available one for the species, but
I am loath to transfer it to Calophyllum, as it was apparently selected by him
on a misconception of the floral parts, and in no way applies to any species of
the genus, all the species of Calophyllum having 4-merous flowers.

GARCINIA Linn.

Garcinia cordata sp. noy. § Hugarcinia.

Arbor glabra circiter 8 m alta; foliis sessilibus vel subsessilibus, ovato-
lanceolatis vel late lanceolatis, chartaceis vel subcoriaceis, nitidis, apice
acutis vel obscure acuminatis, basi late rotundatis distincte cordatis,
nervis utrinque 20 ad 25; floribus axillaribus, fasciculatis, pedicellatis,
4-meris, staminibus numerosis, in phalangibus 4 brevissime stipitatis vel
subsessilibus dense congestis; pistill rudimento fungiforme.

A glabrous tree about 8 m high. Branches brown, terete, the branch-
lets distinctly 4-angled, frequently olivaceous. Leaves opposite, sessile
or subsessile, ovate-lanceolate to broadly ovate-lanceolate, 10 to 16 cm
long, 3.5 to 6 em wide, broadest in the lower part, chartaceous or sub-
coriaceous, shining, the apex acute or obscurely acuminate, the base
broad, rounded, distinctly cordate; lateral nerves 20 to 25 on each side of
the midrib, slender, irregular, the reticulations nearly obsolete. Flowers
axillary, fascicled, white, 4-merous, 4 to 8 in each axil, their pedicels
about 7 mm long. Outer two sepals elliptic-ovate, obtuse, 4 mm long,
smaller than the inner two which are petaloid. Petals membranaceous,

959483

200 MERRILL.

elliptic-ovate, rounded, concave, 5 to 6 mm long. Stamens indefinite,
arranged on both sides of four very slightly stipitate or subsessile, nar-
rowly obovoid phalanges. Rudimentary ovary fungiform, the stipe about

3 mm long, the stigma circular, 2 mm in diameter.

Luzon, Province of Cagayan, San Vicente, For. Bur. 17236 Ourran, March 8,
1909, on river banks near sea level. .

A species well characterized by its ovate-lanceolate or broadly lanceolate,
sessile leaves which are broadest in the lower part and distinctly cordate at the
base. Among the Philippine species apparently allied to Garcinia dives Pierre,
and G@. eugeniaefolia Wall., but very different from both.

KAYEA Wall.

Kayea brevipes sp. nov. § BPulkayea.

Arbor glabra circiter 10 m alta; foliis anguste oblongo-ellipticis vel
anguste oblongis, usque ad 23 cm longis, subcoriaceis, nitidis, basi acutis,
apice acuminatis, nervis utrinque circiter 30, petiolo crasso, furfuraceo,
3 ad 4 mm longo; floribus axillaribus, solitariis, sessilibus vel brevissime
pedicellatis, circiter 2.6 cm diametro.

A glabrous tree about 10 m high. Branches and branchlets terete,
grayish-brown, rather slender. Leaves narrowly oblong-elliptic or nar-
rowly oblong, 15 to 23 cm long, 2 to 4 cm wide, narrowed towards both
ends, the base acute, the apex sharply acuminate, subcoriaceous, when
dry shining and somewhat pale; primary nerves about 30 on each side
of the midrib, not prominent, anastomosing, the alternating secondary
nerves frequently nearly as distinct; petioles stout, furfuraceous, 3 to 4
mm long. Flowers yellow, solitary, axillary, sessile or subsessile, about
2.6 cm in diameter. Outer two sepals orbicular-ovate, 7 to 8 mm long,
coriaceous, concave, obtuse, the inner two similar, thinner. Petals oblong-
obovate, about 13 mm long, the apex broad, retuse. Stamens indefinite.
Ovary glabrous, 1-celled, 8-ovulate.

Luzon, Province of Nueva Vizeaya, Amucucan, For. Bur. 14846 Darling, May
13, 1909, along streams at an altitude of about 500 m, locally known as babac.

A species allied to Kayea nawesii (¥.-Vill.) Vesque, differing especially in its
much shorter petioles and larger flowers. It is probably the species identified

by Vidal as “Ochrocarpus longifolius Thouars 7? ,” and of which he figures a leaf
only (Sinopsis Atlas (1883) t. 12, f. E.).

VIOLACEA..

VIOLA Linn.

Viola patrinii Ging. in DC. Prodr. 1 (1824) 293; Hook. f. & Thoms. in Hook.
f. Fl. Brit. Ind. 1 (1872) 183; Forbes & Hemsl. in Journ. Linn. Soe. Bot. 23
(1886) 53.

Luzon, Province of Cagayan, For, Bur. 17095 Curran: District of Lepanto,
Merrill 4452, 4499: Province of Benguet, Bur. Sci. 5899, 5999 Ramos, For. Bur.
5097, 10940 Curran, Bur. Sci. 2472, 2767, 3554 Mearns. Mtnvoro, For, Bur.
9771 Merritt. Mitnpanao, Lake Lanao, Mrs, Clemens 21, and several unnumbered
specimens: Bokidnon, Worcester.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 201

Not previously reported from the Philippines; India to Amur and Manchuria,
Japan, China and Formosa: reported also from Timor. As in China, the species
is very variable in the Philippines.

Viola diffusa Ging in DC. Prodr. 1 (1824) 293; Forbes & Hemsl. 1. ¢. 52.

Luzon, District of Lepanto, Balili, Merrill 4612, November, 1905, growing in
crevices of rocks on terraces of coffee plantations.

Himalaya and Khasia Mountains to China and Formosa; not previously re-
ported from the Philippines.

Viola mearnsii sp. nov.

Planta nana, glabra, acaulis, vix stolomifera, 2 ad 3 em alta; foliis
ovatis vel late cordato-ovatis, acutis vel obtusis, crenatis, membranaceis,
1 ad 1.5 cm longis, basi cordatis; stipulis lineari-lanceolatis, acuminatis,
4 ad 5 mm longis, parce setaceo-dentatis ; bracteolis 2, lnearibus, circiter
4 mm longis; floribus albis, circiter 6.5 mm longis, calcare saccato.

A small, glabrous, acaulescent, non-stoloniferous plant 2 to 3 em high
or less. Leaves membranaceous, ovate or broadly cordate-ovate, obtuse or
acute, base strongly cordate, the lobes and sinus rounded, margins cre-
nate, 1 to 1.5 cm long, nearly as wide; petioles slender 1 to 1.5 em long;
stipules linear-lanceolate, acuminate, 4 to 5 mm long, slightly setaceous-
dentate. Peduncles about 3 em long, often shorter, slender, glabrous,
bearing above the middle a pair of linear bracts about 4 mm long.
Flowers white, solitary, about 6.5 mm long. Sepals ovate-lanceolate,
acuminate, glabrous, 3 mm long, 3-nerved. Spur ovoid, saccate, 2 mm
long or less.

MiInpDANAO, Province of Misamis, Mount Malindang, Mor. Bur. 4625 Mearns &
Hutchinson (type), May, 1906. Two specimens from Luzon are apparently
referable here, Copeland s. n., from Mount Banajao, Province of Laguna, and
Bur. Sci. 4310 Mearns from Pauai, Province of Benguet.

RINOREA Aubl.

Rinorea acuminata sp. nov. § Prothesia.

Arbuscula circiter 2.5 m alta subglabra, ramulis junioribus inflores-
centiisque exceptis; foliis membranaceis vel chartaceis, nitidis, oblongis
vel anguste oblongo-ellipticis, apice valde tenuiter acuminatis, basi acutis,
nervis utrinque circiter 18, prominentibus; cymis axillaribus, brevibus,
ferrugineo-pubescentibus ; staminibus inclusis, antheris liberis.

A shrub about 2.5 m high, the branchlets and inflorescence rather
densely ferruginous-pubescent, the branches gray, ultimately glabrous,
lenticellate. Leaves membranaceous or chartaceous, oblong or narrowly
oblong-elliptic, about 20 cm long, 7 to 8 em wide, shining, glabrous, or
when young with very few hairs beneath, especially on the midrib, the
apex strongly and slenderly sharp-acuminate, base acute, margins sub-
entire, very obscurely glandular-crenate, or very obscurely glandular-
undulate; nerves about 18 on each side of the midrib, prominent, the
reticulations subparallel; petioles pubescent or glabrous, 1 to 1.5 cm
long. Cymes axillary, few-flowered, about as long as the petioles, fer-

202 MERRILL.

ruginous-pubescent. Sepals orbicular or orbicular-ovate, rounded, pubes-

cent externally, 5 mm in diameter. Petals glabrous, elliptic-oblong,
glabrous, obtuse, 4.5 mm long, 3 mm wide. Disk 1 mm long, the stamens
inserted on it, included, their filaments 1 mm long; appendage to the
connective orbicular, rounded, 1 mm in diameter, membranaceous, the
anther-cells tipped with a narrow appendage. Ovary pubescent; style
glabrous, 2 mm long.

Crsu, Mount Licos, For. Bur. 6455 Everett, February, 1907, on steep rocky
slopes, altitude 400 m. V., Maupao.

A species very closely allied to Rinorea copelandii Merr., differing especially
in its nearly glabrous, very strongly acuminate leaves, fewer-flowered cymes, and
somewhat larger flowers.

A specimen from the Catanduanes Islands, For. Bur. 6682 Pray is probably
referable here, differing from the type in its nearly glabrous branchlets.

COMBRETACEZ.
TERMINALIA Linn.

Terminalia darlingii sp. nov. § Diptera.

Arbor subglabra circiter 10 m alta; foliis coriaceis, ad apices ramulo-
rum densissime dispositis, anguste oblongo-obovatis, nitidis, apice abrupte
subtruncato-rotundatis, late breviter acuminatis, basi sensim angustatis,
nervis utrinque 12 ad 14, petiolo 1 ad 1.5 em longo; fructibus in spicis
patulis vel reflexis dense dispositis, ellipticis, compressis, 3 cm longis,
2-alatis.

A tree about 10 m high, nearly glabrous throughout. Branches stout,
the ultimate branchlets much thickened at the ends for the upper 6 to 8
cm and there up to 1.5 cm in diameter, strongly marked with the scars
of fallen petioles. Leaves densely crowded at the apices of the branch-
lets, coriaceous, yellowish-brown and shining when dry, glabrous, nar-
rowly obovate-oblong, about 20 em long, about 7 cm wide near the apex,
the tip abruptly subtruncate-rounded and shortly, broadly acuminate,
gradually narrowed from the upper one-fourth or one-fifth to the narrow,
cuneate base; midrib very prominent, the lateral nerves 12 to 14 on each
side of the midrib, prominent, anastomosing; petioles stout, slightly
pubescent with appressed hairs, 1 to 1.5 cm long. Flowers unknown.
Fruiting spikes in the axils of the upper leaves, 3 to 5 or more on each
branchlet, spreading or recurved, about 15 em long, the peduncles 5 to 7
cm long, terete. Fruits very densely disposed, elliptic, about 3 em long,
2 cm wide, strongly compressed, flattened or rounded on one side, trian-
gular-keeled on the other, surrounded by a thin, straw-colored wing nearly
| cm wide, rounded or acute at the base, apex more or less retuse.

Luzon, Province of Camarines, Mambulao, in forests, altitude about 100
m, For. Bur, 18735 Darling, April 6, 1910.

A very characteristic species, readily recognizable by its crowded leaves which
are narrowly oblong-ovate and gradually narrowed from about the upper one-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 203

fourth to the base, and especially by its strongly compressed, 2-winged fruits
which are borne in dense, peduncled, spreading or reflexed spikes. Locally known
to the Negritos as pagatpagat, and to the Tagalogs as malaputat.

MELASTOMATACE#.
ASTROCALYX gen. nov.

Calyx dense molliter furfuraceo-setaceus,. ttbus infundibuliformis ;
limbus 5-lobatus. Petala 5, elliptico-ovata, acuminata, imbricata. Sta-
mina circiter 65, aequalia, filamentis filiformibus, elongatis; antherae
anguste lineari-oblongae, teretae, rectae, basi angustatae, apice suboblique
truneatae, 2-rimosae, connectivo basi nec elongato nec incrassato ecalcarato.
Ovarium calyci adhaerens, 5-loculare; ovula plurima, placentis incras-
satis angulo inferiore loculorum affixa; stylus elongatus, stigmate puncti-
formi. Arbor, ramulis, petiolis, subtus foliis ad nervos, inflorescentiisque
dense molliter brunneo-furfuraceo-setaceis. Folia opposita, petiolata, in-
tegra, elliptica vel oblongo-elliptica, basi 5-plinervia, nervulis trans-
versalibus numerosis, distinctis. Flores in paniculis terminalibus dis-
positi, mediocres, minute bibracteolati.

Astrocalyx pleiosandra sp. nov.

Arbor circiter 25 m alta; foliis subcoriaceis, 10 ad 20 em longis, 3.5
ad 8 em latis, breviter acuminatis, basi acutis.

A tree about 25 m high. Branches rather slender, terete, glabrous,
light-gray or brownish, the growing parts denscly covered with brown,
rather soft, furfuraceous-setaceous indumentum, as are the petioles, inflo-
rescences, and nerves on the under surfaces of the leaves. Leaves opposite,
elliptic to oblong-elliptic, 10 to 20 em long, 3.5 to 8 cm wide, sub-
coriaceous, entire, the base acute, the apex shortly and rather broadly
blunt-acuminate, the upper surface green or olivaceous when dry, glabrous,
dull or slightly shining, the lower surface of about the same color; lon-
gitudinal nerves 5, the two inner ones leaving the midrib at from 0.5 to
2 cm above the base, extending to the apex, nearly as prominent as the
midrib, the outer pair near the margin, more slender, leaving the midrib
almost at the base of the lamina, scarcely reaching the apex of the leaf,
more or less looped by the anastomoses of the transverse veins ; transverse
veins about 20 between the midrib and the first longitudinal pair of
nerves, distinct, parallel, spreading, alternating with similar nerves be-
tween the first and second pairs of longitudinal nerves; petioles 1 to 3
em long. Inflorescence terminal, 8 to 15 em long, branched at or above
the base, the primary branches 3 to 5 cm long, the branches flower-
bearing above the middle. Flowers red, borne in threes on the ultimate
branchlets, the branchlets subtended by a pair of narrowly oblong, 4
mm long bracts, the flowers subtended by a pair of similar but much
smaller bracteoles ; pedicels 3 to4 mm long. Calyx broadly funnel-shaped,
the tube in anthesis 4 mm long, about 7 mm wide, the limb with 5,

204 MERRILL.

lanceolate, thickened, acuminate, 3 mm long lobes or teeth, these teeth
narrow and keeled on the inside. Petals 5, glabrous, in bud strongly
rostrate, imbricate, elliptic-ovate to elliptic-oblong, slightly inequilateral,
acuminate, about 9 mm long, 4 to 6 mm wide. Stamens about 65,
1-seriate, subequal, the filaments slender, more or less coherent below
in five phalanges, some or all ultimately free or nearly so, 7 to 8 mm
long, glabrous; anthers ‘narrowly linear-oblong, terete, straight or nearly
so, erect, in bud inflexed, gradually narrowed to the base, about 5 mm
long, 0.5 mm in diameter, the connective not produced, appendages none,
opening by two terminal slits, each cell prolonged into a 0.5 mm long,
compressed tube, slightly obliquely truncate. Ovary adherent to the
calyx, 5-celled, the ovules indefinite, on all sides of the thickened
placenta which is attached in the lower inner angle; style elongated,
rather stout, about 12 mm long; stigma 0.5 mm in diameter, punctiform.
Fruit unknown.

Luzon, Province of Camarines, near Daet, For. Bur. 14349 bis Aguilar, July,
1909, in forests near the Maniba River (type): Province of Laguna, Dajican,
Bur. Sci. 8983 Foxworthy, July, 1909, altitude about 300 m.

Aguilar states that the flowers are red, and his specimens bear open flowers;
Foxworthy states that they are greenish-white, but on his specimen the flowers
are not quite mature. The diameter of the trunk is given by Aguilar as 27 em,
and by Foxworthy as 25 em.

This new genus belongs in the Astronieae, and is perhaps most closely allied
to the Bornean monotypic genus Plethiandra. It is, however, very different from
that genus and from all others in the tribe and family. Characteristic features are
its prominently 5-lobed star-shaped calyx, and especially its very numerous
stamens, the anthers being slender and gradually narrowed to the base, opening
by two terminal slits, the cells being produced into very short, compressed tubes,
the connectives not produced and in no way appendiculate.

In the entire family the only genera previously known in which numerous
stamens are found are the Bornean Plethiandra, mentioned above, and the Amer-
ican ones Calyptrella and Miconia.

CEPHALOMEDINILLA gen. nov.

Flores 4-meri. Calycis tubus ovoideus, limbus valde 4-lobatus. Petala
anguste oblongo-obovata, leviter inaequiliteralia, rotundata vel subacuta.
Stamina petalorum numero dupla, aequalia; antherae lineari-lanceolatae,
elongatae, apice 1-porosae, connectivo basi non producto, antice bilobo,
postice minute 1-calcarato. Ovarium calycis tubo adhaerens, 4-loculare,
vertice dense pilosum. Ovula in loculis numerosa, placentis prominulis
angulo interiore loculi affixis; stylus elongatus,, stigmate punctiformi.
Bacca ignota. Frutex scandens, ramulis foliis junioribus inflorescentiis-
que plus minus dense simpliciter pilosis. Folia opposita, sessilia, valde
inaequalia, integerrima, penninervia.. Flores in capitulis axillaribus,
sessilibus, multifloris dispositi, rosei; alabastro in bractea clausa incluso.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 205

Cephalomedinilla anisophylla sp. nov.

Frutex scandens circiter 2 m altus; ramis teretibus, griseis, ramulis
foliis junioribus inflorescentiisque plus minus dense pilosis, pilis simpli-
cibus, albis; foliis oppositis, elliptico-oblongis, chartaceis vel submem-
branaceis, acuminatis, sessilibus, valde inaequalibus, majoribus usque ad
15 cm longis, minoribus.vix 2.5 cm longis; nervis utrinque 4, curvato-
adscendentibus; inflorescentiis capitatis, axillaribus, solitariis, 1.5 ad
2 em diametro, dense multifloris, bracteis numerosis late ovatis involu-
cratis; floribus 4-meris, subsessilibus vel brevissime pedicellatis.

A scandent shrub about 2 m high. Branches terete, rather slender,
glabrous, light-gray, the younger branchlets densely pilose with long,
simple, pale or brownish hairs. Leaves opposite, sessile, elliptic-oblong,
chartaceous or submembranaceous, very unequal, the larger ones of each
pair 12 to 15 em long, 5 to 6 cm wide, the smaller ones less than 2.5 cm
long and 1 cm wide, rather prominently acuminate, the base gradually
narrowed, acute or obtuse, the younger ones more or less densely covered
with pilose hairs, becoming quite glabrous; midrib prominent, the four
pairs of lateral nerves leaving the midrib in the lower one-half of the
leaf, the innermost two pairs reaching to the apex, curved-ascending,
the reticulations transverse, distinct, subparallel. Heads solitary, ses-
sile, in the leaf-axils or in the axils of fallen leaves, hemispheric, 1.5 to
2 cm in diameter, each with from 12 to 20 densely arranged subsessile
flowers, each head subtended by about 10 broadly ovate, membranaceous,
more or less pilose, somewhat acuminate, imbricate, pink or reddish
bracts, 10 to 11 mm long, 8 to 9 mm wide. Flowers 4-merous, pink,
each subtended by two bracteoles, one elliptic-ovate, flat, the other entirely
inclosing the bud, at length splitting down one side and when spread
suborbicular-ovate, about 9 mm in diameter, cleft to the middle into two
elliptic-ovate lobes, the sinus acute, the lobes faintly 3- to 5-nerved,
more or less pilose. Calyx 6 to 7 mm long, ovoid, somewhat narrowed
to the base, very densely pilose with long, soft, simple, white hairs, the
limb 4 mm long, cleft into four narrowly ovate, acute or acuminate, 2.5
to 3 mm long lobes, pilose on both sides. Petals 4, imbricate, thin,
glabrous, 8 to 9 mm long, narrowly oblong-obovate or obovate-sub-
spatulate, much narrowed in the lower one-half, about 4 mm wide, the
apex somewhat inequilateral, rounded or subacute. Stamens 8, equal;
filaments slender, 5 mm long; anthers lanceolate, 4 mm long, somewhat
curved, acuminate,’ opening by a terminal pore, the connective not at
all produced, the base in front with two short, more or less connate,
somewhat curved, thick lobes less than 1 mm long, and with a small,
0.3 mm long spur behind. Ovary adherent to the calyx, 4-celled, the
top densely pilose with long white hairs; ovules many, the placenta

206 MERRILL.

attached to the lower inner angle of each cell; style slender, 9 mm long;
stigma punctiform.

Luzon, Province of Laguna, Dajican, near Paete, Bur. Sci. 8986 Foxworthy,
July 25, 1909, in forests, altitude not given.

This proposed new genus is manifestly closely allied to Medinilla, differing
especially in its produced and prominently 4-lobed calyx-tube and densely pilose
top of the ovary. It differs also from that genus in its dense, hemispherical,
sessile, involucrate heads, and in its buds being entirely inclosed within one of
the bracteoles, the bracteole later splitting and becoming 2-lobed; it also differs
from most of the known species of Medinilla in its very unequal leaves.

MEDINILLA Gaudich.

Medinilla cardiophylla sp. nov.

Species M. myrtiformi simillima et valde affinis, differt foliis paulo
majoribus, basi late rotundatis valde cordatis.

An epiphytic shrub about 1.5 m. high, glabrous throughout. Branches
slender, terete, reddish-brown or grayish. Leaves ovate to ovate-lan-
ceolate, opposite, subcoriaceous, 7 to 10 cm long, 3 to 4.5 cm wide,
the base broad and rounded, rather strongly cordate, the apex long and
rather slenderly acuminate, the acumen blunt, 5-plinerved, the interior
pair prominent and reaching the apex of the leaf, the outside pair much
fainter and reaching to about the middle of the leaf, the reticulations
very faint or subobsolete; petiole about 1 mm long or almost wanting.
Cymes axillary, solitary, slender, few-flowered, 3 to 5 em long, the
peduncles 3.5 cm long or less, the pedicels slender. Flowers 4-merous.
Calyx somewhat campanulate, 3 to 3.5 mm long, base narrowed, limb
produced about 1 mm and with four small, distant teeth. Petals 4,
oblong-lanceolate, somewhat acuminate, about 6 mm long, 2.2 mm wide.
Stamens 8, the four longer ones about 6 mm long, the four shorter ones
5 mm long; anthers 3 to 3.5 mm long, lanceolate, straight, the connective
not produced, with a dorsal, stout, broad, 0.5 mm long spur, the front
with two broad, rather obscure auricles. Ovary 4-celled; style about 6
mm long; stigma punctiform. Fruit scarlet, globose, 6 mm in diameter,
crowned by the minute calyx-teeth.

MINDANAO, Lake Lanao, Camp Keithley, Mrs. Clemens 861, December, 1906,
in fruit, s.., September, 1907, in flower (type). Luzon, Province of Albay,
Mount Mayon, Bur. Sci. 6509 Robinson, September 6,°1908, on trees in stream
depressions, altitude 400 m.,

A species manifestly very closely allied to Medinilla myrtiformis Triana,
differing in the points stated in the diagnosis. The présent species with M.
myrtiformis Triana and M. ramiflora Mery. form a group of closely allied forms
which perhaps have as much the characters of Anplectrum as of Medinilla. Medi-

nilla ramiflora Merr. may not prove to be specifically distinct from the Bornean
Anplectrum homoeandrum Stapf.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 207

Medinilla cauliflora sp. nov.

Arbuscula circiter 3 m alta, foliis junioribus subtus ad_ nervos,
ramulisque plus minus plumoso-stellato-tomentosis; foliis oppositis,
membranaceis, acuminatis, elliptico-ovatis, basi angustatis, 5-plinerviis,
breviter petiolatis ; inflorescentiis caulifloris, densissime confertis ; floribus
4-meris, bracteatis, calycibus obscure 4-angulatis, 4-lobatis, lobis 5-7-
nerviis; staminibus 8, admodum inaequalibus, antheris postice longe
calcaratis.

A shrub about 3 m high, the branchlets and the younger leaves on the
midrib and nerves beneath more or less covered with soft, stellate-plumose
hairs. Branches rather slender, grayish, terete or obscurely 4-angled, the
upper axils more or less bearded. Leaves opposite, membranaceous, ellip-
tic-ovate, 9 to 15 cm long, 4 to 7.5 em wide, the apex shortly acuminate,
the base narrowed and acute or slightly acuminate; nerves three on each
side of the midrib, leaving it in the lower one-third, curved-ascending, the
innermost pair reaching to the apex, distinct, the transverse reticulations
slender, distant, not prominent; petioles 5 mm long or less, sometimes
subobsolete. Inflorescence on the trunk below the leaves, of very short,
congested branches, forming a compact mass 8 cm in diameter or less, the
branches stout, each bearing many, subdistichous, oblong-ovate, crowded, -
9 to 10 mm long bracts which are acute or obtuse, some empty, some
subtending flowers. Flowers pink or red, 4-merous, glabrous, the pedicels
stout, 4 mm long. Calyx obovoid, becoming more or less urceolate,
obscurely 4-angled, 6 mm long, constricted below the mb, the limb
produced, 2.5 to 3 mm long, cleft nearly to its base into four, broadly
ovate, 5- to 7-nerved, apiculate lobes. Petals obovate, 7 to 8 mm long,
4 to 5 mm wide, the apex strongly imequilateral, obliquely truncate,
base narrowed. Stamens 8, the filaments of four about 5 mm long, of
the other four 5.5 mm in length; anthers equal, 3.5 mm long, oblong-
lanceolate, slightly curved, opening by a single terminal pore, the base
with a slender, curved, dorsal spur nearly 2 mm in length, in front with
two stouter, curved auricles less than 1 mm long. Ovary 4-celled, the
lower one-half adherent to the calyx, free above, the top more or less
conical, glabrous ; style joined with the ovary, at least 5 mm long; stigma
punctiform. Berry globose, fleshy, pink, 7 to 8 mm in diameter ; seeds in-
definite, 0.7 to 0.8 mm long.

Neeros, Canlaon Voleano, For. Bur. 17397 Curran, September, 1909, in forests
at an altitude of about 1,200 m.

A species well characterized by its dense, cauline inflorescence and its 4-lobed
calyx-limb, the lobes distinctly nerved. It is apparently not very closely allied to
any described species.

208 MERRILL.

Medinilla clementis sp. noy.

Frutex glaber ut videtur scandens, ramulis angulatis, valde quadria-
latis; foliis subcoriaceis, oppositis, petiolatis, oblongis, acuminatis, basi
rotundatis vel leviter subcordatis, usque ad 28 cm longis, 7- vel 9-pliner-
viis; inflorescentiis terminalibus, elongatis, bracteis parvis; floribus
5-meris.

A shrub, apparently scandent, glabrous throughout. Branches stout,
4-angled, the,angles winged, the wings 2 to 3 mm wide, the nodes setose.
Leaves opposite, oblong, subcoriaceous, somewhat shining, 20 to 28 cm
long, 7 to 11 cm wide, the apex acuminate, the base rather broad, rounded,
often somewhat subcordate; nerves 7 or 9, the outer ones basal, the
interior one or two pairs leaving the midrib shortly above the base and
reaching the apex, the reticulations obsolete or nearly so; petioles 2.5
to 3 cm long. Inflorescence terminal, long-peduncled, slender, 25 to
40 cm long, the branches few, short, the bracts 5 to 7 mm long. Pedicels
about 1 em long. Calyx cup-shaped, about 4 mm long, the limb somewhat
produced, truncate. Petals 5, pink, narrowly obovate, inequilateral,
about 10 mm long, 5 mm wide. Stamens 10, equal, the filaments 6 mm
long; anthers narrowly lanceolate, somewat curyed, 6 mm long, the basal
dorsal spur slender, less than 5.5 mm long, the anterior two auricles
stout, about 1 mm long. Style slender, 13 mm long.

MINDANAO, Lake Lanao, Sacred Mountain near Camp Keithley, Mrs. Clemens
s. n., July, 1907 (type), and between Camp Keithley and Malabang, Mrs. Clemens
s. n., November, 1906.

A species well characterized by the details given in the diagnosis; it is perhaps
as closely allied to Medinilla teysmanni Miq. as to any other described form, but
is quite different from Miquel’s species.

Medinilla obovata sp. nov.

Frutex epiphyticus, glaber, vel ramulis junioribus plus minus brunneo-
furfuraceis; foliis parvis, verticillatis, ternis vel quarternis, coriaceis,
obovatis, petiolatis, apice late rotundatis vel leviter retusis, vix 3 em longis,
triplinerviis, reticulis obsoletis; floribus ignotis, ut videtur 6-meris, longe
pedicellatis, solitariis, vel in cymis paucifloris dispositis, axillaribus;
fructibus urceolatis, limbo calycis producto truncato.

A glabrous epiphytic shrub, or the ultimate branchlets more or less
brown-furfuraceous. Branches stout, grayish, terete, the branchlets some-
what quadrangular, the internodes short, mostly less than 1 cm in length,
the nodes not barbellate. Leaves whorled, in threes or fours, coriaceous,
obovate, less than 3 cm long, and less than 1 cm in width, the apex
broadly rounded or somewhat retuse, narrowed below to the cuneate base,
the margins sometimes recurved; nerves.3 only, the lateral pair leaving
the midrib shortly above the base, the reticulations obsolete ; petioles about
5 mm long. Flowers unknown. Infrutescence axillary, the peduncles
solitary, axillary, about 1 cm long, each bearing a single pedicel as long

os Beh
- —_—— —_ )l

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 209

or longer than the peduncle, the fruits urceolate, about 8 mm long, 6 mm
in diameter, the calyx-limb persistent, truncate, produced about 4 mm,
6-celled.

Necros, Mount Marapara, For. Bur. 17353 Curran, September 11, 1909, epi-
phytic on trees in forests, altitude about 500 m. é

Although the specimen is without flowers I have no doubt but-that it is
referable to Medinilla, even though the fruits are distinctly 6-celled, indicating
a 6-merous flower, a character uncommon in the genus. It is well characterized
by its comparatively very small leaves which are whorled, petioled, and broadly
obovate, the veins three only, and the reticulations obsolete. The fruits are
solitary, but at the junction of the pedicels with the peduncles are found some
minute scars indicating a few-flowered, probably cymose, or possibly umbellate
inflorescence. Medinilla obovata has much smaller leaves than most of the other
Philippine species of the genus.

Medinilla whitfordii Merr. in Govt. Lab. Publ. (Philip.) 29 (1905) 37.

Carioma triplinervia Rolfe in Journ. Linn. Soe. Bot. 21 (1884) 310; Vid.
Rev. Pl. Vasc. Filip. (1886) 136; Cogn. in DC. Monog. Phan. 7 (1891) 571, non
Medinilla triplinervia Cogn. ;

Luzon, District of Lepanto, Mount Data, For. Bur. 10956 Curran: Province of
Benguet, Pauai, Bur. Sci. 4478 Mearns; Pauai to Baguio, Merrill 4779: Province
of Zambales, Mount Pinatubo, Bur. Sci. 2556 Foawworthy, Provinces of Laguna
and Tayabas, Mount Banajao, Whitford 961 (type of M. whitfordii), Bur. Sci.
6062 Robinson, Bur. Sci. 2388 Foxworthy, For. Bur. 7899 Curran & Merritt:
Province of Albay, Mount Mayon, Vidal 779 in herb. Kew (type of Carionia trip-
linervia Rolfe), Bur. Sci. 6504 Robinson.

Reéxamination of the type of Medinilla whitfordii Merr. shows that its flowers
are 6-merous instead of 5-merous as originally described, and comparison of
our recently collected Philippine material with the type of Carionia triplinervia
Rolfe at Kew has shown that Medinilla whitfordii Merr. is identical. The
species is to me a Medinilla rather than a Carionia, in spite of its 6-merous
flowers. Carionia is distinguished from Medinilla by its 6-merous flowers and
its spreading 4 to 5 mm long calyx-teeth, but Carionia triplinervia has a truncate
calyx with only very minute teeth, and in all other respects is a typical Medinilla ;
either it must be transferred to Medinilla, or the few species with 6-merous
flowers now placed in Medinilla must be transferred to Carionia. Medinilla has
4- to 6-merous flowers, and it would be just as logical to segregate those species
with 4-merous flowers in one genus, and those with 5-merous flowers into another,
as it would be to refer species like the present with 6-merous flowers but with a
truncate calyx to Carionia. The specific name selected by Mr. Rolfe is invalidated
in Medinilla.

MEMECYLON BI.

Memecylon sessilifolium sp. nov.

Arbor glabra circiter 16 m alta; ramulis acute tetragonis, angustissime
alatis, crassis; foliis sessilibus, coriaceis, nitidis, usque ad 12 cm longis,
late obtuseque acuminatis, penninerviis, nervis vix distinctis. Fructibus
fasciculatis vel solitariis, pedicellatis, 1 ad 1.5 cm diametro.

A glabrous tree about 16 m high. Branches stout, subterete, covered
with a thin, grayish bark, the branchlets sharply 4-angled, very narrowly
winged on the angles, the wings less than 0.5 mm wide. Leaves sessile,

210 MERRILL.

oblong or narrowly oboyate-oblong, firmly coriaceous, yellowish and
shining when dry, 8 to 12 cm long, 3 to 5 cm wide, the apex broadly
and obtusely acuminate, the base subacute, margins recurved; lateral
nerves not distinct, about 25 on each side of the midrib, nearly obsolete
beneath. Flowers unknown. Fruit axillary, solitary or fascicled, sub-
globose, 1 to 1.5 cm in diameter, their peduncles 5 to 4 mm long, 2-celled,
2-seeded.

Luzon, Province of Camarines, Mambulao, For, Bur. 18734 Darling, April 6,
1910, in forests, altitude about 100 m, locally known as madadignay.

A strongly characterized species, probably most closely allied to Memeeylon
wightii Thwaites of India and Ceylon.

ARALIACEA.
SCHEFFLERA Forst.

Schefflera brevipes sp. nov.

Glabra,. inflorescentiis sparse puberulis exceptis; foliolis circiter 7, vel
foliorum superiorum 3, oblongis, coriaceis, integris, apice subrotundatis,
usque ad 20 longis; petiolo vix 1 cm longo; paniculis terminalibus, ramis
racemoso-dispositis inferioribus usque ad 30 cm longis; umbellulis race-
moso-dispositis, breviter pedunculatis, 8- ad 12-floris; floribus breviter
pedicellatis, 5-meris.

A glabrous, erect or subscandent shrub. Branches rather stout, the
ultimate ones about 1 cm in diameter, glabrous. Leaves alternate, the
petioles stout, very short, on the material available none exceeding 1 cm
in length, inflated and clasping at the base; leaflets about 7, in the
uppermost leaves sometimes only 3, oblong, coriaceous, 15 to 20 em long,
6.5 to 8 cm wide, entire, upper surface shining, the lower dull, the apex
mostly broad and rounded, rarely broadly and obtusely acuminate, the
base acute or somewhat decurrent-acuminate; lateral nerves about 20,
distinct, anastomosing, not much more prominent than are the secondary
ones and the primary reticulations; petiolules 5 to 8 em long. Panicles
terminal, the common rachis about 30 cm long, all parts slightly puber-
ulent, becoming glabrous or nearly so, the branches alternate, spreading
or ascending, the lower ones 30 cm long. Umbels numerous, mostly
scattered, racemosely arranged along the primary branches, each with
from 8 to 12 flowers, the peduncles 3 to 4 mm long, puberulent, the
pedicels usually about 2 mm long. Flowers 5-merous. Calyx obeconic,
truncate, about 2 mm long and the same diameter at the apex. Petals
5.2 mm long, base 2 mm wide, cohering by their apices and falling as a
whole. Stamens 5; filaments 2.5 mm long; anthers 1 mm long. Ovary
5-celled.

Luzon, Province of Isabela, Cabojan River, For. Bur. 18545 Alvarez, March 21,
1909.

A species well characterized by its oblong, coriaceous, ample, entire leaflets,

NEW OR NOTEWORTHY -PHILIPPINE PLANTS, VIII. Ay i

and especially by its very short petioles. Similar in some respects to Schefflera
clementis Merr., of Mindanao, but with more slender branches which are not.
covered with bracts, differently shaped leaflets, and short petioles.

Schefflera leytensis sp. nov.

Glabra; foliis longe petiolatis, foliolis circiter 8, ellipticis vel oblongo-
ellipticis, usque ad 14 cm longis, basi rotundatis, apice abrupte subcaudato-
acuminatis, margine integris; floribus 5-meris, in umbellulis longe
pedunculatis dispositis, umbellulis racemoso-dispositis ; ovario 10-loculare.

Glabrous throughout. Stems unknown. Leaves long-petioled, the
petioles at least 20 cm in length; leaflets about 8, elliptic or oblong-
elliptic, coriaceous, glabrous, shining on both surfaces when dry, 10 to
14 cm long, 4.5 to 7 em wide, entire, the margins reflexed, the apex
abruptly subcaudate-acuminate, acumen 2 cm long or less, base rounded ;
nerves 10 to 12 on each side of the midrib, spreading, anastomosing, the
reticulations lax; petiolules 4 to 5 em long. Inflorescence (or partial
inflorescence?) about 40 cm long, the rachis stout, dark-colored when
dry, the umbels racemosely disposed, their peduncles about 4 em long,
about 30 on each imflorescence or branch. Flowers 5-merous, about
25 in each umbel, their pedicels 3 to 5 mm long. Calyx cup-shaped, 3
mm long, truncate. Petals 5, oblong-ovate, about 4 mm long, coherent
by their apices and falling as a whole. Stamens 5; filaments 4 mm long;
anthers 2 to 2.2 mm long. Ovary 10-celled; stigma conic, less than 1
mm long.

LEYTE, central divide, altitude about 1,150 m, For. Bur. 16904 Rosenbluth,
February, 1909.

HRICACEA.

DIPLYCOSIA Blume.

Diplycosia parvifolia sp. nov.

Frutex parvus epiphyticus vix 1m altus, ramulis junioribus plus minus
setosis ; foliis coriaceis, nitidis, ellipticis vel elliptico-oblongis, basi acutis,
apice acutis vel leviter acuminatis, 1 ad 2.5 em longis, subtus glanduloso-
punctatis, nervis lateralibus obsoletis, basi interdum obscure triplinerviis ;
floribus paucis, axillaribus, solitariis, pedicellatis, calycis segmentis minute
ciliatis.

A small, epiphytic, erect shrub less than 1 m high, diffusely branched,
the branches slender, glabrous, brown, slightly striate, terete, the branch-
lets with scattered slender, brown, more or less appressed, setose hairs.
Leaves coriaceous, elliptic to elliptic-oblong, 1 to 2.5 cm long, 1 cm wide
or less, entire, the base acute, the apex acute or slightly acuminate,
glabrous and shining when dry, but the margins of the younger leaves
more or less ciliate-setose, the lower surface with scattered, dark-colored,
small glands ; midrib distinct, the lateral nerves and reticulations obsolete,
the base sometimes very obscurely triplinerved ; petioles about 2 mm long,

2T2 MERRILL.

glabrous. Flowers axillary, solitary, few, their pedicels setose, up to 7
mm in length, the corolla unknown, basal bract minute, less than 1 mm
long, the apex of the pedicel with two orbicular-ovate bracteoles about 1.5
mm in diameter. Calyx glabrous except the slightly ciliate margins of the
lobes, acerescent, the lobes just after anthesis ovate, acuminate, about 2
mm long; style persistent, 2 mm long.

Neeros, Canlaon Voleano, on mossy trunks in forests at an altitude of about
1,500 m, Merrill 6995, April, 1910.

A species well characterized by its comparatively small leaves, the lateral
nerves obsolete except sometimes the very faint subbasal pair.

MYRSINACEZE.
ARDISIA Sw.

Ardisia biflora sp. nov. § Akosmos.

Arbuscula glabra usque ad 3 m alta; foliis petiolatis, oblongis vel
lanceolato-oblongis, integris, chartaceis, valde acuminatis, subtus valde
glanduloso-punctatis, in sicco nitidis, pallidis; inflorescentiis axillaribus,
solitariis, tenuibus, bifloris, quam folia multo brevioribus; sepalis petalis-
que valde glanduloso-punctatis.

A shrub 1 to 3 m high, glabrous; branches and branchlets slender,
terete, gray or brownish. Leaves alternate, oblong to oblong-lanceolate,
chartaceous, pale and shining when dry, entire, 4 to 8 cm long, 1 to 2.5
em wide, the apex strongly acuminate, the base cuneate, the lower surface
very strongly and densely glandular-punctate; nerves about 10 on each
side of the midrib, not prominent, slender, anastomosing, the secondary
ones nearly as prominent; petioles slender, about 5 mm long. Inflores-
cence axillary, solitary, slender, the peduncle 6 to 10 mm long, bearing
at its apex two flowers, their pedicels 6 to 8 mm in length. Flowers
greenish-white, tinged with pink, 5-merous, hermaphrodite. Sepals
reniform-ovate, less than 1mm long, united for the lower third, strongly
glandular-punctate, scarcely overlapping, spreading, rounded, the margins
minutely ciliate. Petals elliptic-ovate, acute, 3.5 to 4 mm long, prom-
inently glandular-punctate throughout, the tube about 0.5 mm_ long.
Anthers 2 mm long, oblong-ovoid, minutely apiculate, glandular on the
back, the filaments very short. Ovary ovoid, apparently with few (one?)
ovules; style 2 mm long. Fruit globose, glandular, about 5 mm in
diameter.

Luzon, Province of Zambales, Mount Tapulao, in exposed ridge-forests, altitude
about 1,400 m, Bur. Sci. 5073 Ramos (type), For. Bur, 8110 Curran & Merritt,
December, 1907.

A species apparently belonging in the section Akosmos, strongly characterized
by its slender, axillary, 2-flowered inflorescences; the apparently l-oyuled ovary

is suggestive of Discocalyx, but in all other characters the species is unmistakably
an Ardisia.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 213

Ardisia clementis Elm. Leafl. Philip. Bot. 2 (1910) 665. § Tinopsis.

Arbor vel arbuscula, inflorescentiis exceptis glabra; foliis elliptico-
oblongis vel elliptico-ovatis, chartaceis vel subcoriaceisy acuminatis, glan-
dulis manifestis destitutis, nervis utrinque circiter 14, subtus prominen-
tibus; paniculis terminalibus quam folia brevioribus, bipinnatim com-
positis ; floribus brevissime racemosis vel subumbellatis ; sepalis imbricatis,
integris, rotundatis, margine cillatis.

A tree or shrub, glabrous except the inflorescence which is somewhat
brown-pubescent or puberulent. Branches terete, gray or brownish.
Leaves alternate, elliptic-oblong to elliptic-ovate, chartaceous or sub-
coriaceous, 11 to 25 em long, 4 to 9 cm wide, entire, the apex acute or
acuminate, the base cuneate, dull or slightly shining when dry; nerves
about 14 on each side of the midrib, beneath distinct, anastomosing,
the reticulations -rather fine; petioles stout, about 1 cm long. Panicles
terminal, about 10 cm long, the lower branches about 4 cm long, usually
spreading, the upper ones shorter, each branch bearing from 5 to 7
flowers near the apex, subumbellately arranged or in a short raceme
the pedicels about’ 1 cm long, slightly accrescent in fruit. Calyx about
5 mm in diameter, the lobes broadly ovate, rounded, overlapping to the
right, glandular-punctate, margins ciliate, united for the lower one-third.
Petals elliptic-ovate, obtuse, about 7 mm long, 5 mm wide, slightly
glandular-punctate in the upper one-half, the tube about 1 mm long.
Anthers narrowly ovoid, slightly apiculate, 4 mm long, the median
portion of the back with few, rather large glands. Ovary glabrous;
style about 4.5 mm long, not exserted in bud. Fruit globose, slightly
longitudinally striate when dry, about 6 mm in diameter.

Mrnpanao, Lake Lanao, Camp Keithley, Mrs. Clemens 231, 889, and s. n.
(type), February, 1906, March, 1907, and January, 1907.

Ardisia confertiflora sp. nov. § Tinus.

Arbor parva, usque ad 5 m alta, glabra; foliis ut videtur plus minus
succulentis, in sicco chartaceis vel submembranaceis, oblongo-obovatis,
vel obovatis, apice rotundatis, basi cuneatis, nervis lateralibus obscuris;
inflorescentiis axillaribus, racemosis, floribus in tertia superiore parte
plus minus dense dispositis; sepalis glandulosis, emarginatis, margine
minute ciliatis.

A small tree reaching a height of 5 m, glabrous. Branches and
branchlets rather stout, terete, brownish. Leaves somewhat crowded
- on the younger branchlets, when fresh apparently somewhat thick and
succulent, when dry chartaceous or submembranaceous, dull or slightly
shining, obovate or narrowly obovate, 6 to 10 cm long, 2.5 to 5 em long,
entire, the apex broad, rounded, narrowed from the wpper one-third or
one-half to the cuneate base, the lower surface minutely puncticulate ;
nerves obscure, about 13 on each side of the midrib, very slender, some-

214 MERRILL.

times nearly obsolete; petioles 5 mm long or less. Racemes in the upper
axils, solitary, stout, 5 to 10 cm long, only the upper third flower-bearing,
this portion of tke rachis densely covered with scars and pedicels, the
latter 1 to 2 cm long. Flowers pink or purplish, 5-merous, comparatively
large. Calyx about 8 mm in diameter, the lobes united for the lower
one-third or one-half, about 3 mm wide, elliptic-ovate or ovate, apex
slightly emarginate, all parts densely glandular, the margins minutely
ciliate. Corolla-tube about 2 mm long, the lobes ovate or broadly
oblong-ovate, about 13 mm long, 7 mm wide, somewhat acuminate, the
acumen obtuse, glandular-punctate. Anthers about 6.5 mm long, slightly
apiculate-acuminate, with few large glands on the back, the cells not
septate or rugose, the filaments about 2 mm long. Ovary ovoid; style
about 1 em long, obscurely glandular. Fruit globose or depressed-
globose, about 8 mm in diameter, violet when mature, said to be edible.

BaTaNnes Istanps, Batan, Santo Domingo de Basco, Bur. Sci. 3589 Fénix
(type), Bur. Sci. 3214, 8216 Mearns. BapuyANES ISLANDS, Fuga, Bur. Sci. 3245
Mearns.

This species is manifestly allied to Ardisia humilis Vahl, A. boissieri A. DC.,
and A. littoralis Andr., but is at once distinguishable from all by its racemosely
disposed flowers, these being crowded along the upper one-third of the racemes
and not umbellately arranged at the ends of the branches. Other distinguishing
characters are its comparatively large flowers and its emarginate sepals. The

material on which the above species is based, was previously referred by me to
Ardisia humilis Vahl2 Local name pain.

Ardisia curranii sp. nov. § Tinopsis.
Arbor glabra, circiter 15 m alta; foliis pseudoverticillatis, coriaceis,
oblongis vel late oblongo-lanceolatis, breviter acuminatis, nervis num-

crosis, patulis, vix distinctis; paniculis terminalibus, quam folia multo—

longioribus; sepalis pubescentibus, punctatis, margine ciliatis; ovario
puberulo.

A glabrous tree about 15 m high. Branches terete, striate, light-gray,
rather stout. Leaves alternate, somewhat pseudo-verticillately crowded
at the apices of the branchlets, oblong to broadly oblong-lanceolate,
coriaceous, brown when dry, entire, 15 to 20 em long, 5 to 7 cm wide,
dull or slightly shining, the apex shortly and’ broadly acuminate, the
base somewhat decurrent-acuminate, scarcely glandular-punctate, or very
minutely and obscurely so; nerves 25 to 30 on each side of the midrib,
obscure, spreading, scarcely anastomosing, the reticulations obsolete;

~

petioles 2.5 to 3 em long. Panicles terminal, about 30 cm long, somewhat |

narrowly pyramidal, the lower branches 15 em in length, the flowers
racemosely disposed on the ultimate branchlets, their pedicels 1 to 2 em
long. Calyx about 4.5 mm in diameter, ferruginous-pubescent outside,

®This Journal 3 (1908) Botany 425.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 215

glabrous within, the lobes ovate, rounded, united for about one-half
their length, their margins ciliate, glandular-punctate. Petals ovate,
7 to 8 mm long, about 4 mm wide, glabrous, glandular-punctate, acu-
minate, the tube short. Anthers lanceolate-ovoid, 5 mm long, the
apex long-apiculate-acuminate, the median portion of the back dis-
tinctly glandular, the filaments very short. Ovary ovoid, minutely
ferruginous-puberulent ; style about 4 mm long. Fruit globose or de-
pressed-globose, 6 to 7 mm in diameter, glabrous, minutely apiculate.

Luzon,. Province of Camarines, near Lupi, in forests, altitude about 100 m,
For. Bur. 10760 Curran, July 11, 1908.

Ardisia darlingii sp. nov. § Tinopsis.

Arbor glabra circiter 20 m alta; foliis oblongis vel anguste oblongo-
obovatis, chartaceis vel subcoriaceis, nitidis, minutissime obscureque
puncticulatis, apice rotundatis, obtusis vel obscure late acuminatis, basi
cuneatis, margine recurvatis; inflorescentiis terminalibus axillaribusque,
corymboso-paniculatis ; sepalis glandulosis, margine ciliato-pubescentibus.

A tree about 20 m high, glabrous. Branches terete, grayish-brown,
rather stout. Leaves alternate, somewhat crowded on the younger branch-
lets, oblong or narrowly oblong-obovate, entire, 7 to 11 cm long, 1.5
to 3.5 em wide, shining, chartaceous or subcoriaceous, the apex rounded,
obtuse, or broadly and obscurely acuminate, narrowed from about the
middle to the cuneate base, the margins recurved, the lower surface very
minutely and obscurely glandular-puncticulate ; nerves about 15 on each
side of the midrib, not prominent, obscurely anastomosing, the reticula-
tions lax, obscure or subobsolete ;-petioles about 1 cm long. Inflorescence
terminal and in the upper axils, corymbose-paniculate 10 cm long or less,
the lower branches of the terminal panicles about 5 cm long, the upper ones
shorter, each branch bearing at its apex a congested raceme or umbel of
from 5 to 10 flowers, the pedicels unequal in length, each subtended by an
oblong or oblong-ovate, obtuse bracteole about 3.5 mm long, 2 mm wide, the
pedicels 3 to 12 mm long. Sepals elliptic-ovate, about 3 mm long, 2.5
mm wide, rounded, entire, slightly united at the base, overlapping, dis-
tinctly glandular-punctate with small reddish glands, the margins ciliate-
pubescent. Petals about 4.5 mm long, 3 mm wide, free or nearly so, acute
or obscurely acuminate, obscurely glandular-punctate. Anthers oblong-
ovate, apiculate-acuminate, 2.5 mm long, obscurely glandular on the back,
the filaments very short. Ovary glabrous; style 2 mm long.

Luzon, Province of Abra, near Bacog, in mountain forests, altitude about
1,800 m, For. Bur. 14611 Darling, February 9, 1909 (type); Tue, altitude about
1,300 m, For. Bur. 14601 Darling.

Somewhat remotely allied to Ardisia lanceolata Roxb., but very different from
that species.

95948——-4

216 MERRILL.

Ardisia diffusa sp. nov. § Acrardisia.

Arbuscula vel arbor glabra; foliis chartaceis vel subcoriaceis, acumin-
atis, brevissime petiolatis, integris, subtus prominente verrucoso-glandu-
losis; paniculis terminalibus, amplis, diffusis, quam folia multo longio-
ribus, pyramidatis; floribus umbellulatis, longissime pedicellatis, 5-meris,
sepalis petalisque glandulosis.

An erect, glabrous shrub or tree, the branches terete, reddish-brown.
Leaves alternate, scattered, elliptic-oblong, chartaceous or subcoriaceous,
brown and somewhat shining when dry, entire, 7 to 9 cm long, 3 to 5 em
wide, the apex shortly and sharply acuminate, the base acute, the lower
surface with numerous, scattered, verrucose glands; nerves 15 to 20 on
each side of the midrib, not prominent, anastomosing, the reticulations
lax; petiole stout, 2 to 3 mm long. Panicles terminal, pyramidal,
reaching a length of 20 cm, the branches alternate, spreading, the lower
ones 9 cm long, the upper ones gradually shorter, the secondary branches
_ Slender, 1.5 to 2.5 em long, racemosely arranged on the primary ones,
each bearing at its apex from 3 to 7 long-pedicelled, umbellately arranged
flowers, the pedicels slender, 1 to 2 em long. Calyx about 4 mm in
diameter, the lobes 5, oblong or oblong-ovate, obtuse, entire, about 1.6
mm long, somewhat united at the base, spreading in anthesis, not at all
imbricate, prominently glandular-punctate with large glands, the margins
ciliate. Petals oblong-ovate, 6 to 7 mm long, about 3 mm wide, united
for about the lower 1.5 mm, the apex prominently acuminate, with few
large, scattered glands in the upper one-half. Anthers 4.5 mm long,
prominently acuminate, cordate at the base, with few glands along the
median portion of the back, the filaments 1.5 to 2 mm long. Ovary
ovoid, glabrous; style about 5 mm long.

MINDANAO, Province of Misamis, Calagan, on the route to Mount Malindang,
For. Bur, 4759 Mearns & Hutchinson, May, 1906.

_ A species most closely allied to Ardisia gardneri, A. divergens, and A. ceyla-
nica, but abundantly distinct from all. It is well characterized by its ample,
pyramidal, rather lax panicles, long-pedunecled umbels and long-pedicelled flowers.

Ardisia lanceolata Roxb. Hort. Beng. (1814) 85, nomen nudum, Fl. Ind. ed.
Carey & Wall. 2 (1824) 275, ed. Carey 1 (1832) 583; Mez in Engl. Pflanzenreich
9 (1902) 122.

Ardisia purpurea Reinw. in Bl. Bijdr. (1826) 684.

MINDANAO, District of Zamboanga, Tetuan, Ahern 596.

Not previously reported from the Philippines; Malay Peninsula, Java, Suma-
tra, Borneo, and Celebes.

Ardisia littoralis Andr. Repos. 10 (1811) ¢. 630; Gamble in Journ. As. Soe.
Beng. 747% (1905), Extra Number, 135.

Ardisia humilis Mez in Engl. Pflanzenreich 9 (1902) 127, fig. 20, A—B, not of
Vahl, or in part only.

LUBANG, Merrill 968. SrmeraARA, Merrill 4160.

This species is apparently a coast shrub only, from 1 to 3 or 4 m in height.
Mr. Gamble has kindly supplied me the following additional information regard-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. Pa Fi

ing the separation of this form from Ardisia humilis: “Ardisia humilis Vahl is
a Ceylon coast shrub which is endemic there and does not extend to the Malay
Peninsula. The figures A-E on page 128 of Mez’s monograph are Climacandra
obovata=Ardisia littoralis Andr. which has septate anthers, which A. humilis has
not. The authors of the “Flora of British India” have mixed up A. humilis and
A, solanacea Roxb., which is quite a different plant, almost a tree and of inland
forests, and Mez has correctly separated them, but he has incorrectly put together
A. humilis.Vahl and A. littoralis Andr.”

Ardisia littoralis Andy. is apparently rare in the Philippines, but we have
several very closely allied forms, at least one of which is abundant and widely
distributed in the Archipelago.

Malay Peninsula and Archipelago, southern China, and the Philippines.

Ardisia boissieri A. DC. in DC. Prodr. 8 (1844) 129; Mez in Engl. Pflanzen-
reich 9 (1902) 129.

This species is very common and widely distributed in the Philippines, extend-
ing from northern Luzon to southern Mindanao, a tree of the hill forests at low
and medium altitudes ascending to at least 600 m in some localities;: it reaches
a height of 15 m in some regions, and is not a seacoast plant. It is very similar
in all superficial characters to A. littoralis Andr., but can usually be at once
distinguished by its anthers being prominently glandular on the back, and not
transversely septate. What I take to represent this species comprises about 80
‘specimens in this herbarium, from all parts of the Philippines, which have, for
most part, been identified as Ardisia humilis Vahl, many of them so named by
Doctor Mez. It is very probable that some of the extra-Philippine specimens
cited by Mez under Ardisia humilis, should be referred to A. boissieri, and it is

likewise very probable that this name will not prove to be the oldest one.
°

Ardisia pirifolia Mez 1. c. 129.

This species, the type of which I have examined in the Berlin Herbarium, is
distinguishable from A. boissieri only by the most trivial characters; in all
respects except in having the sepals minutely emarginate, it is quite the same
as A. boissieri. The type was from Polillo, not from Luzon, and the species is
represented by Bur. Sci. 9292 Robinson, from the same island, Merrill 1101, from
Baler, Province of Tayabas, Luzon, and apparently also by Elmer 5645 from the
Province of Union, Luzon, the latter so identified by Doctor Mez. The sepals are
not always glabrous, but are usually more or less ciliate on the margins; the
only character left for specific separation of this form from A. boissieri is the
very trivial one of the emarginate (very slightly) sepals.

Ardisia verrucosa Pres] Rel. Haenk. 2 (1835) 65; Mez 1. c. 134.

This species is also manifestly closely allied to, and perhaps not specifically
distinct from Ardisia boissieri A. DC. Mez distinguishes it especially by its 2-
flowered umbels, but Pres] describes it as having from 2- to 5-flowered umbels,
and one of the original specimens, in the Prague Herbarium, which I have
examined, shows at least 5 flowers. Doctor Mez examined the specimen of the
original collection preserved in the Vienna Herbarium.

Ardisia macgregorii sp. nov. § Z%inus.

Arbuscula glabra, circiter 1 m alta; foliis lanceolatis, obtusis, coriaceis,
subtus minute dense puncticulatis, nervis reticulisque densis, obscuris ;
inflorescentiis axillaribus, solitariis, simplicibus, fructibus subumbellatim
dispositis.

218 MERRILL.

A glabrous shrub about 1 m high. Branches terete, brown, glabrous.
Leaves coriaceous, brown and somewhat shining when dry, lanceolate,
8 to 10 cm long, 1 to 1.5 em wide, gradually narrowed at both ends,
the apex blunt, the base cuneate, the margins entire, somewhat recurved,
the lower surface minutely and densely puncticulate; nerves numerous,
obscure, densely disposed, the reticulations also obscure; petioles 1 cm
long or less. Inflorescence axillary, solitary, much shorter than the
leaves, the peduncles about 2 cm long. Flowers unknown. Fruits
umbellately disposed at the apices of the peduncles, their pedicels 1 to 1.5
em long, 4 or 5 at the apex of each peduncle. Persistent calyx about 5
mm in diameter, the lobes ovate, rounded, glabrous or nearly so, glandular-
punctate, united for about one-half their length. Fruit globose, about 5
mm in diameter.

CreBu, near Toledo, Bur. Sci. 1722 McGregor, October 28, 1906.

A species manifestly allied to Ardisia humilis Vahl, and A. boissieri A. DC.
distinguishable by its narrow, lanceolate leaves.

Ardisia mindorensis sp. noy. § Pyrgus.

Species A. grandidenti Mez similis, sed differt petiolo multo breviori,,
dentibus minoribus, a A. serrata (Cav.) A. DC. differt inflorescentiis
plus minus dense ferrugineo-tomentosis, foliis subtus parce pubescentibus.

A shrub 3 to 5 m high. Branches terete, brown or grayish, striate,
usually rather thick, glabrous, the branchlets often ferruginous-pubescent.
Leaves pseudo-verticillately crowded at the apices of the branchlets, sub-
tending the terminal panicles, elliptic-ovate to elliptic-lanceolate, char-
taceous, 9 to 18 cm long, 3 to 6 em wide, gradually narrowed towards
both ends, the base acute, rarely somewhat obtuse, the apex rather
prominently acuminate, margins distinctly and irregularly serrate-dentate,
the teeth rather small, the upper surface glabrous and shining, or the
midrib and nerves slightly puberulent, beneath also shining and distinctly
ferruginous-tomentose on the midrib and primary nerves, obscurely
glandular-punctate; nerves 12 to 15 on each side of the midrib, elevated
and very prominent on the lower surface, curved-ascending, anastomosing,
the reticulations rather distinct; petioles 3 to 7 mm long, ferruginous-
tomentose, ultimately nearly glabrous. Panicles terminal, pyramidal,
6 to 13 em long, the rachis, branches and branchlets densely ferruginous-
tomentose, the primary branches spreading, the lower ones often 5 em
long, the upper ones shorter, the secondary branches mostly in the upper
half of the primary ones, each bearing from 5 to 7 flowers arranged in a
condensed raceme or subumbellate, the pedicels ferruginous-tomentose,
9 to 12 mm long. Calyx-lobes broadly triangular-ovate, acute, about 1.5
mim long and wide, imbricate, pubescent, the margins prominently ciliate,
glandular-punctate. Petals ovate, 5 mm long, 3 mm wide, acute or

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 219

obscurely acuminate, with few, scattered, comparatively large glands.
Anthers 2.5 mm long, apiculate, not glandular. Ovary ovoid, ferru-
ginous-puberulent; style about 5 mm long. Fruit globose, black-purple
when mature, somewhat fleshy, about 7 mm in diameter.

Minporo, Mount Halcon, in forests, altitude about 1,800 m, Merrill 5675, 5782,
6145 (type), November, 1906, For. Bur. 4342 Merritt; Mount Irauan, For. Bur.
8728 Merritt, January, 1908, altitude about 1,300 m; Mount Sablayan, For. Bur.
11012 Merritt, March, 1908, altitude about 970 m; mountains back of Abra de:
llog, For. Bur. 8793 Merritt, January, 1908, altitude 500 m.

As noted above, this: species is closely allied to Ardisia grossedentata Mez,
differing in its much shorter petioles. It bears much the same relationship to
that species as does A. curtipes Merr. to A. serrata (Cayv.) A. DC. It is distin-
guished from H. serrata var. brevipetiolata Merr. by its ferruginous tomentum and
differently shaped leaf-bases.

Ardisia oblongifolia sp. nov. § Stylardisia.

Arbor vel frutex erecta, ramulis, foliis junioribus subtus, inflorescen-
tiisque minute brunneo-puberulis; foliis oblongis vel oblongo-lanceolatis,
chartaceis, acuminatis, petiolatis, alternis; paniculis terminalibus, quam
folia brevioribus, ramis divaricatis, paucis.

An erect shrub or tree. Branches terete, light-gray or brownish, the
younger ones brown-puberulent. Leaves alternate, scattered, oblong to
oblong-lanceolate, 11 to 18 em long, 2.5 to 5 em wide, chartaceous, entire,
dull or shghtly shining when dry, the apex rather gradually and sharply
acuminate, the base acute, beneath, when young, somewhat ferruginous-
puberulent becoming glabrous, not manifestly glandular-punctate ; nerves
up to 18 on each side of the midrib, not very prominent, often obscure ;
petioles 5 to 7 mm long. Panicles terminal, pyramidal, 10 em long
or less, the branches few, alternate, divaricately spreading, the lower
ones 3 to 4 cm long, the rachis, branches and branchlets puberulent
with dark-brown hairs. Flowers subumbellately disposed, the umbels
peduncled and racemosely arranged on the primary panicle-branches, the
peduncles about 5 mm long, sometimes less, the pedicels slender, 2 to 3
mm long. Calyx about 3 mm in diameter, the five lobes ovate, obtuse,
about 1.5 mm long, glandular-punctate, the margins minutely puberulent.
Petals ovate, nearly free, acute, 3.5 mm long, 2.5 mm wide, not at all
. glandular-punctate.- Anthers 3 mm long, not glandular. Ovary gla-
brous; style 3.5 to 5 mm long, often exserted before anthesis.

MiInDANAO, Lake Lanao, Mrs. Clemens 779 (type), from between Malabang and
Camp Keithley, November, 1906, and three sheets without number from Camp
Keithley.

Most of the flowers do not have the styles exserted, but a few of them on
the type specimen have the styles decidedly exserted, hence the species is placed
in the § Stylardisia. In might, with almost equal propriety, be placed in the
§ Akosmos.

220 MERRILL. .

Ardisia palawanensis sp. nov. § Pyrgus.

Arbuscula circiter 1.5 m alta; foliis petiolatis, oblongo-oblanceolatis
vel late oblongo-lanceolatis, chartaceis, irregulariter serratis, subtus ad
costa plus minus pubescentibus, valde glanduloso-punctatis; floribus
5-meris, sepalis petalisque ferrugineo-villosis, plus minus punctatis ; ovario
villoso.

A shrub about 1.5 m high. Branches terete, brownish, pubescent.
Leaves pseudo-verticillate at the nodes and subtending the inflorescence,
normal leaves also subtending each, or most of the panicle-branches,
oblong-oblanceolate or broadly oblong-lanceolate, chartaceous, shining
when dry, 14 to 20 em long, 3.5 to 7 em wide, somewhat pubescent on the
midrib on both surfaces, the apex obscurely and bluntly acuminate, the
base gradually narrowed, cuneate, margins entire near the base, in the
upper one-half or two-thirds irregularly serrate, the under surface dis-
tinctly and rather densely glandular-punctate; nerves 9 to 12 on each
side of the midrib, beneath very distinct, anastomosing ; petioles pubescent,
1 em long or less. Inflorescence terminal, subtended by a whorl of leaves,
the rachis about 15 cm long, and with the branches and pedicels more or
less densely brown-pubescent, the branches alternate, spreading, simple,
most of them subtended by a normal leaf, the lower ones 6 cm long, the
upper gradually shorter, each bearing at the somewhat swollen apex from
2 to 5 subumbellately disposed flowers, and with numerous scars of fallen
pedicels, each flower subtended by an oblong, pubescent bracteole about 5
mm in length. Pedicels about 1.5 cm long, ferruginous-villous. Sepals
5, broadly ovate, acute or shortly acuminate, about 5 mm long, 3.5 mm
wide, accrescent and 6 to 7 mm in length, densely ferruginous-villous,
margins strongly villous-ciliate, glandular-punctate. Petals nearly free,
elliptic-ovate, about 6 mm long, 4 mm wide, obtuse, slightly pubescent,
glandular only in the median part of the upper one-half. Anthers 3.5
mm long, not glandular, apiculate-acuminate, their filaments nearly 2
mm long. Ovary globose, ferruginous-pubescent; style glabrous, 5 mm
long. Immature fruits globose, somewhat ferruginous-pubescent, inclosed
by the somewhat accrescent calyx-lobes.

PALAWAN, about 3 miles northeast of Puerto Princesa, Por. Bur. 3518 Curran,
January 19, 1906, an undershrub in flat forests. 3

As species undoubtedly belonging in the section Pyrgus in spite of the normal
leaves subtending the panicle-branches, the whole inflorescence subtended by a
whorl of leaves. It is apparently most closely allied to Ardisia grandidens Mez,

but is very different from that species. The ferruginous-pubescent or villous
panicles, pedicels, sepals, petals, and immature fruits are characteristic.

Ardisia reptans sp. nov. § Bladhia.

Suffruticosa, caulis reptans, ramulis foliiferis erectis vel ascendentibus,
brevibus, densissime ferrugineo-tomentosis; foliis ternato-pseudoverticil-
latis, ellipticis vel ovato-ellipticis, membranaceis, acutis vel obtusis, basi
rotundatis, obscure punctatis, margine prominente distanter serrulatis,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 2

nervis utrinque circiter 5, subtus prominentibus ; inflorescentiis axillaribus,
solitariis, tenuibus, paucifloris, foliis subaequalibus vel brevioribus.

A suffrutescent Abas the stems creeping or prostrate, rooting, striate,
glabrous or nearly so, slender, brown, the erect leaf- bearing branches
densely pubescent with dark-brown, crisped hairs, these branches less
than 20 ecm high, often some roots appearing above the lowermost leaves.
Leaves pseudo-verticillate, in threes, their petioles 5 mm long or less,
densely crisped-tomentose, the lamina membranaceous, elliptic or elliptic-
ovate, 3 to 4 cm long, 1.5 to 2.5 em wide, with scattered, crisped, brown
hairs on both surfaces, especially on the midrib and nerves, obscurely
glandular, the apex acute or obtuse, base rounded, margins prominently
and rather distantly denticulate; nerves about 5 on each side of the
midrib, prominent beneath, obscurely anastomosing, the reticulations lax,
obscure. Flowers unknown. Infrutescence axillary, solitary, very slender,
brown-puberulent, with intermixed longer crisped hairs, the rachis 2 to 2.5
em long, bearing near its apex few long-pedicelled fruits, the pedicels
about 1 em long, puberulent, each subtended by a narrowly lanceolate
or linear, acuminate, 2.5 mm long bracteole. Sepals persistent, reflexed
in fruit, lanceolate, gradually narrowed upward to the acuminate apex,
about 3 mm long, united for the lower 0.5 mm, more or less brown-
puberulent, margins obscurely ciliate, very obscurely glandular-punctate.
Fruit fleshy, globose, 5 to 6 mm in diameter, red, glabrous, tipped with
the slender, 3 mm long, persistent style; seed globose, 4 mm in diameter.

Luzon, Province of Pampanga, Mount Abu, Bur. Sct. 1933 Foxworthy, Decem-
ber 31, 1906, in forested ravines, altitude about 1,360 m.

A species of the section Bladhia, and apparently closely allied to Ardisia
pusilla A. DC. (A. villosa Mez, non Roxb.) of Japan, and to A. faberi Hemsl. of

China. All other species of the section, with one exception, A. metallica N. E.
Br., of Sumatra, are confined to the Himalayan region, China, and Japan.

DISCOCALYX Mez.

Discocalyx insignis sp. nov.

Frutex glaber circiter 3 m altus; foliis alternis, vix pseudo-verticillatis,
elliptico-oblongis, usque ad 40 cm longis, margine dense denticulatis, basi
longe decurrento-acuminatis; petiolo 6 ad 9 cm longo; floribus dioicis,
5-meris, glabris.

An erect glabrous shrub about 3 m high, the ultimate branchlets
stout, about 1 cm in diameter. Leaves alternate, not pseudo-verticillate,
elliptic-oblong, 35 to 40 em long, 10 to 14 cm wide, chartaceous or sub-
coriaceous, usually grayish when dry, shining, not glandular-punctate,
the apex shortly and obscurely blunt-acuminate, the base long-decurrent-
acuminate, the margins entire in the lower part of the leaf, but above
the lower one-fourth densely denticulate; nerves about 20 on each side
of the midrib, prominent, anastomosing, the reticulations distinct, rather
lax; petiole stout, 6 to 9 cm long. Panicles fascicled at the ends of

222 MERRILL.

special branches, these branches simple or ‘branched near their apices,
up to 18 em long, the apical portions thickened, cylindric, marked by
numerous scars, the panicles numerous, entirely glabrous, slender, 5 to 8
em long, all parts marked with linear or punctate glands, the panicles
2-pinnate, the flowers racemosely arranged on the ultimate branchlets.
Staminate flowers 5-merous, their pedicels 2 to 3 mm long. Calyx 1.6
mm in diameter, glandular-punctate, glabrous, the lobes ovate, obtuse,
united for one-half their length. Corolla 3 mm in diameter, glandular-
punctate, the lobes ovate, obtuse, united for one-half their length.
Anthers less than 0.5 mm long, sessile. Rudimentary ovary wanting.
Pistillate flowers and fruits unknown.

MINDANAO, Province of Surigao, in the valley of the Agusan River near Amparo,
in forests at an altitude of about 130 m, For. Bur. 7616 Hutchinson, August 26,
1907 (type). Nrcros, Mount Marapara, Por. Bur. 13688 Curran, with immature
flowers. .

A species similar in vegetative characters to Discocalyx effusa Mez, but with
very much larger leaves and much longer petioles, the inflorescence also entirely
different. It is apparently most closely allied to D. montana Elm., but is quite
distinct from that species.

Discocalyx macrophylla sp. noy.

Frutex erectus, glaber, circiter 2 m altus; foliis alternis, elliptico-ovatis
vel elliptico-oblongis, subcoriaceis, in sicco nitidis, usque ad 31 cm
longis, valde denticulatis, basi rotundatis, petiolo usque ad 15 cm longo;
floribus in paniculis brevibus congestis, 5-meris, glabris.

An erect shrub about 2 m high, the ultimate branches very stout,
brown, 1.5 to 2 cm in diameter. Leaves alternate, or somewhat crowded
at the apices of the branchlets, elliptic-ovate to elliptic-oblong, about 30
em long, 12 to 16 cm wide, glabrous, subcoriaceous, shining, the lower
surface obscurely and minutely glandular-punctate, somewhat paler than
the upper one, the apex very shortly and obscurely blunt-acuminate, the
base broad, rounded, the margins strongly and densely denticulate except
at the very base where they are entire; nerves 25 to 30 on each side of
the midrib, very prominent, anastomosing, the reticulations distinct on
both surfaces; petioles stout, dark-brown, about 15 cm long. Inflores-
cence on special, leafless (or with one very much reduced leaf) branches,
40 em long or less, from just below the leaves, the apex thickened and
bearing one or several short, dense panicles which doubtless become more
or less diffuse in anthesis. Flowers (in bud, and immature), 5-merous,

all parts glandular-punctate, glabrous, the anthers sessile. Immature

fruit globose, 3 mm in diameter.

Luzon, Province of Cagayan, near San Vicente, in forests, at sea level, lor.
Bur. 17287 Curran, March 8, 1909.

A very characteristic species, distinguishable by its very large and long-petigled
leaves, as well as by the very long, specialized branches that bear the inflorescences,
It is unquestionably allied to Discocalya effusa Mez, although quite different from

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 223

that species, and even more closely allied to D. insignis Merr., differing from
the latter especially in its differently shaped leaves, which are broad and rounded
at the base, not decurrent-acuminate, its much longer petioles, and more numerous
leaf-veins.

EMBELIA Burm.

1. Embelia coriacea Wall. Cat. (1829) no. 2314; A. DC. Prodr. 8 (1844) 87;
Mez in Engl. Pflanzenreich 9 (1902) Silos

MinpANaAo, Lake Lanao, Camp Keithley, Mrs. Clemens 671, 1069, and three
sheets without number, September-October, 1906, 1907. The specimens agree per-
fectly with Maingay 1018, and with 5056 Dr. King’s Collector, from the. Malay
Peninsula, in our herbarium.

Penang and Malacca, Sumatra, Java, and Borneo; not previously reported from
the Philippines.

OLEACEZ.

JASMINUM Linn.

Jasminum cumingjii sp. nov.

Frutex ut videtur scandens, glaber, inflorescentiis obscure puberulis
exceptis; foliis oppositis unifoliolatis, ovatis, acuminatis, nitidis, basi late
-rotundatis, nervis utrinque 5 vel 6; inflorescentiis axillaribus, longe
pedunculatis, 5-floris; calycis lobis 4 vel 5, subulatis, corollae lobis 6,
oblongo-lanceolatis, acuminatis, quam tubus paullo brevioribus.

A shrub, apparently scandent, glabrous throughout except the obscurely
puberulent inflorescence. Branches terete, brown, smooth. Leaves sim-
ple, opposite, ovate, 7 to 11 cm long, 4 to 7 cm. wide, membranaceous,
brown and shining when dry, the base broad, rounded, the apex rather
sharply acuminate; nerves 5 or 6 on each side of the midrib, distinct,
anastomosing, the reticulations very lax; petiole about 1.5 em long,
jointed. Cymes axillary, solitary, the peduncles rather slender, 3 to 7
em long, each with 5 flowers near the apex, rarely with an additional
pair of flowers at about the middle of the peduncle, the bracteoles
setaceous, 3 to 5 mm long, the pedicels about 3 mm long. Calyx cup-
shaped, about 4 mm long, 2.5 mm in diameter, slightly puberulent, with
4 or 5 setaceous, slender, 0.5 to 0.8 mm long teeth. Corolla-tube 1.5 cm
long, the lobes 6, narrowly oblong or oblong-lanceolate, acuminate, about
12, mm long, 2.5 to 2.8 mm wide. Anthers 4 mm long. Style 11 mm
long.

PHILIPPINES, without definite locality, Cuming 1565.

A species well characterized by its long-peduncled, axillary, solitary, few-

flowered, slightly puberulent inflorescence, the corolla-lobes nearly as long as the
eorolla-tube. The form has not been rediscovered as yet in the Philippines.

Jasminum triplinervium sp. nov.

Frutex scandens glaber; foliis oppositis, simplicibus, chartaceis,
oblongo-lanceolatis vel anguste oblongo-ovatis, usqte ad 15 em longis,
apice rotundatis vel obtusis, rariter acutis, basi tenuiter triplinervlis ;

224 . MERRILL.

inflorescentiis axillaribus, solitarius, pedunculatis, multifloris, foliis sub-
aequilongis; calycis lobis 4 vel 5, setaceis, brevibus, corollae laciniis 5,
ellipticis, rotundatis, quam tubus dimidio breyioribus.

A scandent glabrous shrub. Branches slender, terete, gray. Leaves
opposite, simple, chartaceous, shining when dry and paler on the under
surface, oblong-lanceolate to narrowly oblong-ovate, 12 to 15 cm long,
+ to 6 cm wide, the base acute, the apex narrowed, rounded, obtuse, or
rarely acute, the base with a pair of slender lateral nerves leaving the
midrib shortly above the insertion of the petiole, reaching ‘at least to
the middle of the leaf and anastomosing with the lateral nerves, these
primary lateral nerves usually 5 or 6 pairs, distant, not prominent,
scarcely more distinct than are the secondary ones; petioles jointed, 1.5
cm long. Cymes in the upper axils, solitary, about as long as the leaves,
many-flowered, the peduncles about 5 cm long, the branches bearing from
two to five flowers, the bracts and bracteoles minute, setaceous, 1 to 2
mm long. Pedicels 8 to 10 mm long. Calyx cup-shaped, glabrous, the
tube 3 to 4 mm long, the teeth 4 or 5, unequal, 1 to 2.5 mm long, linear-
lanceolate, acuminate, minutely scaberulous. Corolla white, the tube 2.4
em long, the lobes 5, elliptic or oblong-elliptic, about 12 mm long, 8 mm
wide, rounded at the apex, the base broad and minutely biauriculate.
Anthers 6 to 7 mm long, acuminate.

Neoros, Faraon, For. Bur, 13557 Meyer & Foaworthy, September 10, 1909,
in forests, altitude not given.

A species well characterized by its-slenderly triplinerved leaves, and its com-
paratively large flowers. There are from 20 to 25 flowers in each inflorescence.

APOCYNACEA.
ALSTONIA R. Br.

Alstonia paucinervia sp. noy.

Arbor glabra circiter 15 m alta; foliis chartaceis vel subcoriaceis,
anguste oblongo-obovatis, usque ad 13 cm longis, basi angustatis, acutis,
apice breviter latissime acuminatis vel obtusis, nervis utrinque circiter 15,
patulis, conspicuis, distantibus; folliculis 20 ad 40 em longis.

A glabrous tree about 15 m-high. Ultimate branches terete, or the
tips somewhat 4-angled, smooth, olivaceous. Leaves usually in whofls
of 4, narrowly oblong-obovate, chartaceous or subcoriaceous, 9 to 13 cm
long, 3.5 to 5 em wide, the upper surface smooth and shining, the lower
surface of about the same color but dull or very slightly shining, the
apex very broadly and shortly acuminate or obtuse, gradually narrowed
from about the middle to the acute base; nerves about 15 on each side
of the midrib, spreading, distant, prominent, anastomosing close to the
margin and forming a faint, somewhat looped, submarginal nerve, the
reticulations lax, rather indistinct; petioles 1 to 1.5 cm long. Flowers
unknown. Fruiting peduncles from the apical axils, 3 to 4 em long,

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 225

simple or dichotomously branched at the apex, each bearing from 2 to
6 follicles. Follicles cylindric, glabrous, longitudinally striate, about 3
mm in diameter, 20 to 40 em long, pendulous. Seeds flattened, oblong,
6 mm long, 1.5 mm wide, covered with short, brown hairs, both ends with
numerous, long, brown hairs 7 mm in length or less.

Luzon, Province of Camarines, Paracale, For. Bur. 18726 Darling, March 19,
1910, in forests at an altitude of about 70 m, locally known as batino.

A species closely allied to Alstonia macrophylla Wall., but with smaller, some-
what differently shaped leaves which have fewer lateral nerves.

‘ CONVOLVULACEZ.
DICHONDRA Forst.

Dichondra repens Forst. Char. Gen. Pl. (1776) 39, ¢. 20; DC. Prodr. 9
(1855) 451.

Sibthorpia evolvulacea Linn. f. Suppl. (1781) 288.

Dichondra evolwulacea Britton in Mem. Torr. Bot. Club. 5 (1894) 268.

Luzon, District of Bontoc, Bauco, dry hillsides, altitude about 1,300 m,
Father M. Vanoverbergh 19, January, 1910, locally known to the Ilocanos as
napalapayag.

The genus is new to the Philippines, the present species being widely distributed
in tropical, subtropical and subtemperate regions of the world.

VERBENACE.
PYGMAEOPREMNA gen. noy.

Calyx parvus, anguste campanulatus sub anthesi regulariter 5-dentatus,
sub fructu auctus, 2-lobatus, lobo altero 2-dentato; altero 3-dentato.
Corollae tubus brevis, cylindraceus, rectus, breviter exsertus; lmbus
distincte 2-labiatus, lobo minore exteriore, integro, lobo majore interiore,
3-lobato, fauce intus parce villoso. Stamina 4, subaequalia, tubo affixa,
quam corolla breviora ; antherae ovatae, loculis parallelis vix divergentibus.
Ovarium 2-loculare, loculis 2-ovulatis; stylus apice brevissime 2-fidus.
Drupa parva, calyce insidens, obovoidea, excentrica, exocarpio carnoso,
tenui, endocarpio duro, indiviso, saepissime 1-loculare vel rarius obscure
2-loculare. Semina oblonga, compressa, exalbuminosa. Suffruticosa,
parva, erecta, parce ramosa, sparse pubescens. Folia opposita, simplicia,
integra. Cymae parvae, breves, terminales vel in axillis superioribus
pedunculatae. Flores parvi, albi vel virido-albi.

Pygmaeopremna humilis sp. nov.

Suffrutex erectus, usque ad 15 cm altus; foliis obovato-oblongis, opposi-
tis, breviter petiolatis, membranaceis vel subchartaceis, acutis vel brevissime
acuminatis, integris, basi acutis, subtus pallidioribus, nervis utrinque
circiter 6, distantibus, distinctis; inflorescentiis terminalibus axillari-
busque, usque ad 2 cm longis, puberulis; floribus parvis, circiter 4.5 mm
longis.

2296 MERRILL.

An erect somewhat woody plant about 15 cm high, from stout, elongated,
woody roots, the stems simple or dichotomously once branched, terete,
slender, the branchlets pale or dark in color, puberulent. Leaves obovate-
oblong, 8 to 12 em long, 3 to 5 em wide, membranaceous or subchartaceous,
entire, the apex acute or very shortly acuminate, rather gradually narrowed
from about the upper third to the acute base, the upper surface shining,
with very few, scattered, short hairs, becoming glabrous or nearly so, the
lower surface paler, eglandular, often shining, sometimes somewhat
puberulent on the nerves; nerves about 6 on each side of the midrib,
distant, distinct, anastomosing, the reticulations lax; petioles 2 to 4 mm
Jong, puberulent. Inflorescence cymose, rather densely puberulent, termi-
nating the stems and branches, or in the dichotomously branched plants
terminating the main stem between the branches and also frequently
terminating the branches, pedunculate, the peduncles about 1 cm long,
the cymes in anthesis about 1 em long and wide, somewhat larger in fruit,
rather densely flowered, the pedicels about 2 mm long, subtended by small
bracteoles. Calyx in anthesis about 2 mm long, puberulent outside,
narrowly funnel-shaped, equally 5-toothed, the teeth obtuse, 0.5 mm long,
as wide as long, accrescent and persistent, 2-lpped in fruit. Corolla
white or greenish-white, slightly exserted, 4.5 mm long, the tube broad,
cylindric, straight, glabrous outside, inside somewhat villous, about 1.5
mm long, the limb prominently 2-lipped, the smaller lip rounded, about
2 mm in diameter, entire, imbricately covering the larger lip in bud, the
larger lip prominently 3-lobed, the middle lobe rounded, 1.6 mm in
diameter, entire, the lateral lobes about 1 mm long and 1.5 mm wide, all
lobes reflexed, or the middle lobe of the upper lp suberect. Stamens 4,
inserted in the tube, the filaments 0.5 mm long; anthers ovate or broadly
elliptic, 0.6 mm long, the cells parallel, not divergent. Ovary ovoid,
glabrous, 2-celled, each cell 2-ovuled ; style less than 1 mm long, minutely
cleft at the apex. Fruit black, broadly obovoid, tipped by the remains
of the style, about 5 mm long, 4 mm in diameter, somewhat inequilateral,
the pericarp thin, fleshy, the endocarp bony, longitudinally rugose, 1-celled,
containing a single seed, but frejuently with indications of an additional
cell, the second cell rarely developing a seed. Calyx accrescent, persistent,
in fruit nearly 4 mm in diameter, distinctly 2-lipped, the larger lp with
three, 0.5 mm long teeth, the smaller lobe with two similar but distant
teeth. Seeds oblong, compressed, exalbuminous.

Luzon, Province of Cagayan, Piat, Bur. Sci. 7841 Ramos, April 2, 1909:
Province of Isabela, Ilagan, Bur. Sci. 8124 Ramos, April 29, 1909, in open grassy
plains.

This curious little plant apparently represents the type of a new genus allied
to Premna and Vitex, but more especially to the former. It differs remarkably
from all species of both genera in its habit and in its small size as well as
in floral and fruit characters indicated in the diagnosis. The stout roots, much
larger then the stems, reach a diameter of 5 mm. Striking characters of the
genus, aside from the small size and habit of the plant are its 2-lipped corollas.

en ae

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. De

the smaller, exterior lip entire, the upper and inner lip strongly 3-lobed, and its -
ealyx, which in anthesis is equally 5-toothed, but in fruit is accrescent and dis-
tinetly 2-lipped, one lip 3-toothed, the other 2-toothed.

VITEX Linn.

Vitex longifolia sp. noy.

Arbor inflorescentiis exceptis glabra, circiter 12 m alta; foliis trifo-
holatis, petiolo 5 ad 9 cm longo, foliolis oblongo-lanceolatis vel anguste
oblongo-ovatis, longe subcaudato-acuminatis, basi acutis vel acuminatis,
plus minus inaequilateralibus, usque ad 25 cm longis, subcoriaceis, nervis
utrinque circiter 10, subtus prominentibus ; paniculis terminalibus, pedun-
culatis, amplis, plus minusve pubescentibus; floribus fasciculatis.

A tree about 12 m high, glabrous except the inflorescence. Branches
grayish, the ultimate ones somewhat compressed. Leaves 3-foliolate, the
petioles terete, 5 to 9 cm long. Leaflets oblong-lanceolate to broadly
ovate-lanceolate, subcoriaceous, somewhat shining when dry, brownish,
paler beneath, 20 to 25 em long, 5 to 7 em wide, entire, the apex with
a long, slender acumen, the base somewhat inequilateral, acute or acu-
minate; nerves about 10 on each side of the midrib, beneath prominent,
curved-ascending, rather distant, anastomosing, the primary reticulations
lax; petiolules of the lateral leaflets less than 1 cm long, of the middle
leaflet nearly 2 cm in length. Panicles terminal, solitary, equaling the
leaves, the peduncle 15 cm long, sometimes with a single branch from
the base, most of the primary branches from above the middle, usually
about four at each node, the ultimate branches and branchlets more or
less brown-pubescent. Flowers in fascicles on the ultimate branchlets,
usually in groups of one central slightly pedicelled flower and two lateral
shortly peduncled groups of three flowers each, the bracts small, 2 mm
long or less. Calyx pubescent, cup-shaped, 2 mm long and wide, with
five, short, acute teeth less than 0.5 mm long. Corolla lilac, pubescent
inside and outside, the tube cylindric, 5 to 6 mm long, the upper lip
bifid, 5 to 6 mm long, the lobes narrowly obovate, rounded, the lower lip
3-cleft, the lobes oblong, acute or obtuse, 3 mm long. Filaments some-
what pubescent, the longer two about 8 mm, the shorter ones about 6 mm
in length. Fruit unknown.

MINDANAO, Province of Surigao, in well-drained flat forests on the Gibon River,
altitude about 55 m, For. Bur. 7574 Hutchinson, June, 1907, locally known to the
Manobos as manamu and to the Visayans as aticéco.

A species in the group with Vitew parviflora Juss. (V. littoralis Dene.), but
distinguishable by its quite different and much larger leaflets. ;

TECTONA Linn. f.

Tectona philippinensis Benth. & Hook. f. Gen. Pl. 2 (1876) 1152; F.-Vill.
Nov. App. (1880) 158; Vidal Phan. Cuming. Philip. (1885) 134, Rev. Pl. Vase.
Filip. (1886) 209, (nomen nudum in all cases).

Tectona hamiltoniana Wall.; Schauer in DC. Prodr. 11 (1847) 629, pro parte
(quoad pl. Philip.) ; F.-Vill. 1. e.

228 . MERRILL.

Arbor usque ad 15 m alta; foliis elliptico-ovatis ad ovato-lanceolatis,
acuminatis, 8 ad 15 cm longis, supra glabris vel subglabris, albido-
verruculosis, subtus dense pallide stellato-puberulis; cymis terminalibus,
densis ; floribus circiter 8 mm longis; fructibus circiter 13 mm diametro,
calycibus persistentibus vix inflatis.

A tree reaching a height of 15 m. Leaves elliptic-ovate to ovate-
lanceolate, acuminate, 8 to 15 em long, 3 to 6 cm wide, subcoriaceous,
subentire or the margins above obscurely undulate-crenate, the upper
surface glabrous or nearly so, rather densely white-verrucose, beneath
paler and densely stellate-puberulent; nerves 5 to 7 on each side of the
midrib, distinct beneath, the reticulations dense; petioles densely puber-
ulent, 5 to 7 mm long. Cymes terminal, sometimes in the upper axils, in
anthesis rather dense, becoming rather diffuse in fruit, densely puberulent.
Flowers nearly 8 mm long and 10 mm in diameter. Calyx densely
puberulent, funnel-shaped, 5 mm long, equally 5-toothed, the teeth tri-
angular-ovate, 2 mm long. Corolla-tube for the lower 1 to 1.5 mm
cylindric, about 5 mm in diameter, then abruptly enlarged, the lobes
elliptic-ovate, obtuse, about 4 mm long, the throat villous inside. Fila-
ments about 8 mm Jong, slender, somewhat exserted. Fruit about 13 mm
long, 5 to 6 mm in diameter, the persistent calyx enclosing the drupe but
not inflated, densely puberulent with pale-brownish indumentum, the
drupe about 8 mm long.

Luzon, Province of Batangas, Cuming 1432 (type number), For. Bur. 7746
Curran & Merritt, November, 1907, the latter growing in rather open brush lands
at an altitude of about 50 m, locally known as malapangit.

As no description of the above species has ever been published, a short one
has been given above. Cuming’s specimen was referred by Schauer to Tectona
hamiltoniana Wall., but Bentham & Hooker f. were undoubtedly right in specifi-
cally separating the Philippine plant from the Asiatic one. It is manifestly closely
allied to Wallich’s species, but differs remarkably in the nature of the indumen-
tum, which in 7. hamiltoniana Wall. is tomentose or stellately wooly, and in the
present species minutely and very densely puberulent.

Cuming’s plant has been localized from his own list of localities preserved

in his correspondence with Sir William Hooker at Kew, and is undoubtedly

correct. !

LABIATAE.
SALVIA Linn.

Salvia scaphiformis Hance in Journ. Bot. 23 (1885) 368: Forbes & Hemsl.
in Journ, Linn. Soe. Bot. 26 (1890) 287.

Luzon, Province of Nueva Vizcaya, Merrill 174: Province of Abra, Bur. Sci.
7206 Ramos: Province of Benguet, Blmer 5834, 8687.

Not previously reported from the Philippines; Formosa, and Szechuen, China.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 229

SCROPHULARIACEZ.
BYTHOPHYTON Hook. f.

Bythophyton indicum (Hk. f. & Th.) Hk. f. Fl. Brit. Ind. 4 (1884) 286.

Micranthemum indicum Hk. f. & Th. in Journ. Bot. 9 (1857) 245, t. 7 (excel.
fig. of anthers, fide Hooker f.)

Luzon, District of Lepanto, Mount Data, Merrill 4519, November, 1905, in
shallow water of a small pond at the summit, altitude about 2,250 m.

A monotypic genus previously recorded only from the Nonkreem marshes,
Khasia Mountains, India, altitude about 1,400 m. I am indebted to the director
of the Royal Gardens, Kew, for the identification of the above specimen.

ACANTHACEA.
PERISTROPHE Nees.

Peristrophe lancifolia sp. nov.

Herba erecta vix vel parce ramosa, sparse strigosa; foliis membran-
aceis, anguste lanceolatis, acuminatis, usque ad 20 em longis, 2.5 em
latis ; foliis floralibus oblongis vel oblongo-lanceolatis, acuminatis ; corolla
alba, 5 cm longa.

An erect herb about 50 cm high or less, usually unbranched. Stems
green, longitudinally sulcate, about 2 mm in diameter, glabrous, or the
younger parts slightly appressed-strigose. Leaves lanceolate or narrowly
lanceolate, 12 to 20 em long, 1 to 2.5 cm wide, entire or very slightly
undulate, gradually narrowed upward to the long, slender, acuminate
apex, the base also narrowed, acute or acuminate, somewhat shining
when dry, the upper surface dark-green, the lower surface pale-green,
both surfaces with numerous, scattered, oblong cystoliths, the lower also
minutely white-puncticulate and slightly appressed-strigose on the midrib
and nerves ; lateral nerves about 9 on each side of the midrib, ascending,
anastomosing and forming a nearly straight submarginal nerve, the
secondary nerves and reticulations lax; petioles 5 mm long or less, stri-
gose. Peduncles few, 2.5 cm long or less, terminal and in the upper
axils, strigose, few-flowered. Floral leaves 2, unequal, oblong to oblong-
lanceolate, acuminate, 2.5 to 3.5 cm long, more or less strigose, especially
on the margins. Flowers white, congested, each subtended by one or
two linear-lanceolate, more or less hispid, long-acuminate bracts about 2
cm long, 2 mm wide, and by two or three similar but smaller bracteoles.
Calyx-lobes similar to the bracteoles, subequal, about 8 mm long, hispid.
Corolla 5 cm long, the tube slender, 2.5 cm in length, one lip entire, 1.5
em broad, the other somewhat narrower and shortly 2-toothed at the
apex. Filaments slightly hirsute: anthers 2-celled, the upper cell about
3.5 mm long, the lower 2 mm long, muticous. Style minutely bifid.

230 MERRILL.

Pod about 1.5 cm long, hirsute, long-stalked; seeds 4. Placenta per-
sistent, not separating from the capsule in dehiscence.

Luzon, Province of Bataan, Balanga Mountains, For. Bur. 19258 Curran, Jan-
uary, 1910.

A species well characterized by its narrowly lanceolate, long leaves and by its
large flowers.

RUBIACEA.
GREENIOPSIS Merr.

Greeniopsis pubescens sp. nov.

Arbor parva vel mediocris, omnibus partibus plus minus dense brunneo-
pubescens; foliis oblongo-lanceolatis vel oblongo-oblanceolatis, usque ad
40 em longis, longe acuminatis, basi angustatis, nervis utrinque circiter
20, stipulis 3 cm longis; capsulis 4 mm longis.

A small or medium-sized tree pubescent throughout. Branches stout,
the ultimate ones about 8 mm in diameter, pubescent with short brownish
hairs. Leaves oblong-lanceolate or oblong-oblanceolate, 20 to 40 cm
long, 6 to 11 cm wide, subcoriaceous, shining when dry, both surfaces
pubescent with rather short hairs, or the upper subglabrous, entire, the
apex rather slenderly long-acuminate, narrowed to the base, the lamina
decurrent practically to the base of the short petiole, so that the
leaves are subsessile; nerves about 20 on each side of the midrib, prom-
inent, curved, anastomosing, the ultimate reticulations distinct ; stipules
oblong-lanceolate, chartaceous, 3 cm long, 1 em wide at the base, acu-
minate, at first pubescent, becoming nearly glabrous, deciduous. Panicles
terminal and in the upper axils, peduncled, nearly as long as the leaves,
branched at or above the middle, the rachis, branches and branchlets
densely brown-pubescent with short hairs, the branches flower-bearing
only above the middle. Flowers white, pedicelled, the calyx pubescent,
about 3 mm long, narrowly funnel-shaped, becoming somewhat inflated,
the teeth 5, short, truncate, imbricate, broader then long. Corolla rather
densely pubescent outside, the throat villous within, the tube about 2 mm
long, rather abruptly enlarged above, the lobes 5, rounded, imbricate,
about 1.5 mm long, 2.5 mm wide. Anthers 1.4 mm long. Capsules
subsecund on the ultimate branchlets, oblong or oblong-ovoid, densely
pubescent, 4 mm long, the persistent calyx-teeth oblong, pubescent on
both surfaces.

Luzon, Province of Isabela, Bicobian Bay, Bur. Sci. 10659 McGregor, August,
1909: Province of Cagayan, Bur. Sci. 7407 Ramos, March, 1909 (type).

In general appearance very similar to the other two species of the genus, but

well characterized by its rather uniformly distributed pubescence, which consists»
of short, usually spreading and brownish hairs. The third species for the genus.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 23

HEDYOTIS Linn.

Hedyotis cagayanensis sp. noy.

Frutex erectus, simplex, vix 1 m altus, glaber, partibus junioribus plus
minus puberulis exceptis; caulibus teretibus, partibus junioribus quad-
rangulatis, pulcherrime undulato-crenato-alatis ; foliis membranaceis, am-
pls, petiolatis, usque ad 18 cm longis, oblongo-obovatis vel elliptico-
obovatis, nervis utrinque circiter 8; cymis axillaribus, in capitulis sub-
globosis dense congestis.

An erect undershrub about 60 cm high, unbranched, glabrous, or the
younger parts more or less puberulent. Stem stout, brownish, about 6
mim in diameter, the basal part terete, the younger parts above 4-angled,
each angle with a striking undulate-crenate wing 1 to 2 mm in width.
Leaves oblong-obovate to elliptic-obovate, membranaceous, glabrous, some-
what shining, 13 to 18 cm long, 5 to 8 em wide, the apex blunt, acute,
or broadly acuminate, the base acute; nerves about 8 on each side of the
midrib, distinct, ascending, the reticulations very obscure, lax; petioles
1.5 to 2 cm long; stipules ovate or oblong, about 12 mm long, cut into
about 12 laciniae. Cymes axillary, glabrous, densely many-flowered,
forming subglobose, axillary heads about 2 cm in diameter, the flowers 4-
merous; pedicels 2 mm long or less, the bracts oblong, 2 to 3 mm long.
Calyx-tube somewhat 4-angled, glabrous, 1.5 mim long, the lobes 1.8 mm
in length. Capsule obovoid, 3 mm long.

Luzon, Province of Cagayan, Pamplona, Bur. Sci. 7502 Ramos, March 16, 1909,
in forests along streams.

A species manifestly allied to H. pilosissima Merr., but in adult stage quite

glabrous, but more especially characterized by its square, prominently 4-winged
stems, the wings undulate-crenate.

Hedyotis macgregorii sp. nov.

Frutex erectus, ramulis foliis inflorescentiisque plus minus strigoso-
pubescentibus ; foliis elliptico-oblongis vel elliptico-lanceolatis, usque ad
5.9 cm longis; cymis parvis, axillaribus terminalibusque, subsessilibus
vel breviter pedunculatis, densis, paucifloris; corolla circiter 6 mm longa.

An erect much-branched shrub. Branches terete, glabrous, grayish,
the younger ones prominently 4-angled and rather densely pubescent.
Leaves elliptic-oblong to elliptic-lanceolate, chartaceous or subcoriaceous,
2 to 5.5 cm long, 1 to 2 cm wide, the base acute, the apex blunt, acute
or even slightly acuminate, the lower surface more or less strigose-pubes-
cent with short hairs, especially on the midrib and nerves, the upper
surface also more or less pubescent, ultimately becoming glabrous or
nearly so; nerves about 5 on each side of the midrib, curved-ascending,
anastomosing, the reticulations obsolete or nearly so; petioles densely

959485

229 MERRILL.

rat

pubescent, 2 to 4 mm long; stipules pubescent, ovate, persistent, 3 to 4
mm long, cut into six or more linear lacinizw. Cymes terminal and in
the upper axils, small, dense, comparatively few-flowered, subsessile or
the axillary ones sometimes with peduncles up to 1.5 em in length, all
parts more or less pubescent, the bracts foliaceous, spatulate, 5 to 6 mm
long, the pedicels usually about 1 mm in length. Calyx-tube ovoid,
pubescent, 2 mm long, the lobes usually 4, lanceolate, acuminate, pubes-
cent, 3 mm long. Corolla apparently white, 6 mm long, the tube 4 mm
in length, cylindric, glabrous outside, villous within, usually with 4 lobes,
rarely with 3 or 5, the lobes oblong, obtuse or acute, 2 mm long, reflexed
in anthesis, shghtly strigose on the back. Anthers 1.2 mm long. Style
3.5 mm long, cleft at the apex. Capsule ovoid, 3.5 mm long, somewhat
strigose-pubescent, the seeds black, compressed, angular, irregular, 1.2 to
1.5 mm long.

Luzon, Province of Benguet, Pauai, Bur. Sci. 8490 McGregor, June, 1909
(type), Bur. Sct. 4473 Mearns, August, 1907.

Apparently a species of the mossy forest, allied to Hedyotis benguetensis Elm.,
and other species of that group, but well characterized by its strigose pubescence.

Hedyotis pilosissima sp. noy.

Frutex vel suffrutex erectus, simplex, vix 1 m altus, omnibus partibus
plus minus dense pilosus; foliis chartaceis vel submembranaceis, oblongo-
-lanceolatis vel oblongo-ellipticis, usque ad 16 cm longis, acuminatis, basi
acutis, nervis 6 vel 7 utrinque, adscendentibus ; stipulis amplis, laciniatis ;
cymis in axillis dense congestis, subcapitatis, dense multifloris.

An erect unbranched suffrutescent or decidedly woody undershrub less
than 1 m high, the stems terete, about 6 mm in diameter, brownish or
grayish, densely pilose. Leaves oblong-lanceolate to oblong-elliptic, 12
to 16 cm long, 3 to 5.5 cm wide, chartaceous or submembranaceous,
greenish when dry, somewhat shining, the apex acuminate, base acute,
both surfaces with numerous, long, white hairs, especially so on the
midrib and nerves on the under surface; nerves 6 or 7 on each side of
the midrib, distinct, ascending, not or obscurely anastomosing, the sec-
ondary nerves and the reticulations obsolete; petioles pilose, 1.5 to 2 em
long; stipules ample, green, somewhat pilose, up to 1.5 cm long, 1.2
em wide, cut into about 12, narrowly lanceolate, acuminate laciniae 3 to
5 mm in length, the middle one longer. Cymes axillary, more or less
pilose, congested, forming subglobose or hemispherical inflorescences 1 to
2 cm in diameter. Flowers 4-merous, their pedicels up to 2 mm in
length, sometimes wanting, the bracteoles lanceolate, acuminate, 2 mm
long. Calyx-tube ovoid, glabrous, about 1.2.mm long, the lobes green,
lanceolate, acuminate, 2 mm long, the margins ciliate with long white
hairs. Corolla-tube 2 mm long, the lobes about the same length, oblong,
recurved, usually acute. Anthers 1 mm long. Capsule globose or ovoid,
glabrous except for the few hairs on the persistent calyx-lobes, about 2

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 233

mm in diameter; seeds numerous, angular, black, about 0.3 mm in
diameter. ;

PANAY, Dumarao, in damp, shaded ravines along streams, at an altitude of
about 100 m, Merrill 6704, March 25, 1910.

A species well characterized by its ample leaves and dense pubescence. A
specimen from Maagnas, Province of Camarines, Luzon, Bur. Sci. 6326 Robinson,
August, 1908, may be referable here, but in this plant the capsules are pilose, not
glabrous.

IXORA Linn.

Ixora capitulifera sp. nov.

Arbor glabra circiter 10 m alta; foliis subcoriaceis, in sicco brunneis,
nitidis, oblongis vel oblongo-obovatis, acuminatis, basi leviter rotundatis,
breviter petiolatis; floribus circiter 1.5 cm longis, subsessilibus, in capit-
ulis parvis, densis, longe pedunculatis dispositis; calycis dentibus quam
tubus brevioribus.

A glabrous tree about 10 m high. Branches terete, stout, grayish.
Leaves oblong or oblong-obovate, 4 to 11 cm long, 1.5 to 5 em wide,
subcoriaceous, brown and shining on both surfaces when dry, the apex
shortly and usually bluntly acuminate, rarely nearly acute, the base
narrowed, somewhat rounded, rarely subacute; nerves about 12 on each
side of the midrib, slender, brown, distinct, anastomosing; petioles less
than 2 mm long. Inflorescence terminal and axillary, the peduncles
slender, 2 to 4 cm long, each subtended by from 2 to 4, distichous, broadly
ovate, acuminate bracts about 2.5 mm long, 2 mm wide, usually with
a pair of smaller, narrower bracts above the middle. Flowers 5 to 8 at
the end of each peduncle, sessile, congested, the calyces forming a rather
dense head less than 6 mm in diameter. Calyx 2 mm long, the teeth 4,
triangular-ovate, acute or somewhat obtuse, small, the bracteoles 2, linear,
about 1 mm long. Corolla-tube 13 mm long, 1 mm in diameter, the
lobes four, elliptic, rounded or obtuse, about 4 mm long, 2.5 mm wide.

PaLAwANn, Mount Victoria, in forests along streams at an altitude of about
1,050 m, Bur. Sci. 686 Foxworthy, March 23, 1906, the flowers said to be pinkish
or whitish, with a faint wintergreen odor.

The species is a very characteristic one, readily recognizable by its slenderly
peduncled capitate inflorescence.

Ixora crassifolia sp. nov.

Arbor parva usque ad 9 m alta, inflorescentiis puberulis exceptis glabra,
foliis crassissime coriaceis, ellipticis vel oblongo-ellipticis, apice rotundatis
vel leviter retusis, usque ad 34 cm longis, nitidis, utrinque in sicco dense
minute rugosis, nervis utrinque circiter 10; inflorescentiis terminalibus,
puberulis, circiter 9 cm longis, dense multifloris; floribus circiter 2.5 em
longis.

A small tree 9 m high or less, glabrous except the somewhat cinereous-
puberulent inflorescence. Branches rather stout, brown or grayish.
Leaves elliptic, broadly elliptic or oblong-elliptic, 14 to 24 em long, 7

924 MERRILL.

to 18 em wide, very thickly coriaceous, somewhat shining, when dry
minutely and densely rugose on both surfaces, the apex rather broadly
rounded, sometimes slightly retuse, the base acute or somewhat acuminate,
rarely broadly rounded; nerves about 10 on each side of the midrib,
distinct, anastomosing, the reticulations lax ; petioles stout, 1 to 2 em long ;
stipules very broadly ovate, abruptly acuminate, 5 to 7 mm long, deciduous.
Inflorescence terminal, puberulent, corymbose, subtended by one or two
pairs of broadly ovate, abruptly acuminate bracts 8 mm long or less, with
two lateral basal branches, the rachis short, trichotomously branched, the
primary branches stout, 3 to 5 cm long, all subtrichotomously branched at
their apices, forming a rather dense inflorescence about 9 cm long, and
as wide or wider than long. Flowers white or greenish-white, rather
densely crowded at the ends of the ultimate branchlets, their pedicels 1
to 2 mm long, ebracteolate. Calyx ovoid, puberulent, about 2 mm long,
with 4 short, broadly ovate, acute teeth about 0.3 mm long. Corolla-tube
rather slender, 2.4 cm long, 2 mm wide when more or less flattened out,
the lobes 4, spreading or reflexed, oblong, rounded, 7 mm long, 4 mm
wide. Anthers linear, 5 mm long, the filaments exserted about 3 mm.
Style slender, exserted about 6 mm, the arms thickened, more or less
flattened, about 2 mm long.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9039, 9070, 9439
(type), 9479 Whitford & Hutchinson, December, 1907, and February, 1908, in
dipterocarp forests at from 30 to 50 m above the sea.

A species well characterized by its unusually large, very coriaceous leaves,
which, when dry, are rather pale and minutely, densely rugose on both surfaces.

Ixora ebracteolata sp. nov.

Iazora amboinica Elm. Leafi. Philip. Bot. 1 (1906) 9, non DC.

Arbuscula vel arbor parva, 3 ad 8 m alta, inflorescentiis exceptis glabra ;
foliis petiolatis, coriaceis vel subcoriaceis, in sicco brunneis, nitidis,
oblongis vel oblongo-ellipticis, obtusis vel late brevissime acuminatis,
basi acutis, nervis utrinque circiter 10, distinctis; cymis terminalibus,
puberulis vel subglabris, pedunculatis, multifloris; floribus 9 ad 12 mm
longis, ebracteolatis.

An erect shrub or tree 3 to 8 m high, glabrous except the inflorescence
which is usually puberulent. Branches terete, gray, the younger ones
usually reddish-brown. Leaves oblong to oblong-elliptic, 6 to 12 cm long,
2.5 to 5 em wide, the base acute or decurrent-acuminate, the apex obtuse,
rounded, or broadly and obtusely short-acuminate, brown and shining
when dry, the lower surface paler than the upper; primary nerves about
10 on each side of the midrib, anastomosing, brown, distinct, the reticula-
tions rather lax, distinct; petioles 5 to 10 mm long; stipules lanceolate-
acuminate from an ovate base, 5 mm long or less. Inflorescence terminal,
rather dense, 5 to 7 cm in diameter, usually puberulent, sometimes
glabrous peduncled, many-flowered, the peduncles 3 to 5 cm long, the lower
branches spreading, about 2 em long. Flowers white, mostly in triads on

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 235

the ultimate branchlets, the middle one of each triad sessile or subsessile,
the two lateral ones with pedicels 3 to 5 mm in length, the bracts and
bracteoles wanting. Calyx glabrous, ovoid, about 3 mm long, the teeth
broadly triangular-ovate, acute, 0.5 mm long. Corolla-tube 6 to 9 mm
long, about 2 mm in diameter, the lobes 4, elliptic-oblong, acute or slightly
acuminate, 6 mm long, 2.5 to 3 mm wide. Anthers 3.5 mm long, lanceo-
late, acuminate. Style slightly exserted ; stigma cleft, 2 mm long. Fruit
ovoid, smooth, somewhat fleshy when fresh, 8 mm long or less, dark-
colored when dry.

Luzon, Province of Zambales, Mahumaling, For. Bur. 5845 Curran, January,
1907 (type), on dry cogon-covered slopes; other specimens from the same province
are: Bur. Sci. 4798, 5038 Ramos, For. Bur. 375 Maule, Merrill 2953, 2985, 2080.
Various local names are pamutim, pilis, lumboy-manoc, talab, and tatanie.

The species is entirely different from Jxora amboinica DC., to which Mr. Elmer

referred several of the specimens above cited; it is distinguished from the
majority of the species in the genus by the entire absence of bracts and bracteoles.

Ixora longissima sp. nov.

Arbuscula erecta, glabra; foliis petiolatis, lanceolatis, usque ad 40 cm
longis, crasse membranaceis, sensim longe acuminatis, margine minute
crenato-undulatis; cymis terminalibus, diffusis, amplis, multifloris;
floribus circiter 5 em longis; calycis dentibus acutis, quam tubus brey-
loribus. :

An erect glabrous shrub. Branches terete, pale-brown, smooth and
shining. Leaves lanceolate, about 40 cm long, 5.5 cm wide, thickly
chartaceous, shining when dry, the base acute or somewhat decurrent-
acuminate, gradually narrowed upward into the long, slender, acuminate
apex, the margins minutely crenate-undulate; nervés about 23 on each
side of the midrib, slender, not very prominent, anastomosing, the reti-
culations lax ; petioles stout, 1 to 1.5 em long; stipules very broad, connate,
abruptly acuminate, about 5 mm long. Cymes terminal, very large,
trichotomously branched and rebranched, the peduncle stout, about 5 mm
long, the primary branches about as long as the peduncle, spreading, the
whole inflorescence, including the corollas, about 20 cm wide. Flowers
apparently pink or reddish, numerous, in triads on the ultimate branch-
lets, the middle one of each triad sessile, the two lateral ones with
pedicels 2 to 3 mm long, the bracts oblong-ovate, acuminate, 2 to 6 mm
long, the bracteoles similar but smaller, 1 mm long. Calyx 2 mm long,
the teeth ovate, acute, about 0.5 mm long. Corolla-tube slender, about
4.5 cm long, the lobes broadly ovate-lanceolate or elliptic-lanceolate,
acuminate, thin, reticulate, about 10 mm long, 4 mm wide.

LEYTE, without definite locality, For. Bur. 16975 Rosenbluth, March, 1909.

A species well characterized by its very long leaves which are long and slenderly
acuminate, as well as by its ample, diffuse panicles and very long flowers. It is
probably most closely allied to Ixora salicifolia DC., but seems to be sufficiently
distinct.

236 MERRILL.

Ixora longistipula sp. nov.

Frutex glaber 1.5 ad 3,m altus; foliis petiolatis, oblongo-lanceolatis,
chartaceis, acuminatis, basi acutis; stipulis setaceis, 1 ad 1.8 cm longis;
inflorescentiis terminalibus longe pedunculatis, floribus circiter 2 cm
longis in capitulis parvis dense confertis; calycis dentibus acutis, quam
tubus brevioribus.

A glabrous shrub 1.5 to 3 m high. Branches terete, slender, reddish-
brown. Leaves oblong-lanceolate, 10 to 20 em long, 3 to 4.5 em wide,
chartaceous, usually firmly so, shining, narrowed at both ends, the base
acute, the apex sharply: acuminate; lateral nerves about 13 on each side
of the midrib, distinct, anastomosing, the reticulations lax; petioles
about 1 cm long; stipules setaceous, about 1 cm long, broadened at the
base. Inflorescence terminal, solitary, the peduncles slender, 9 to 15
em long, the flowers subsessile or shortly pedicelled, disposed in a ter-
minal, simple, dense head, this head, excluding the corollas, less than 1 cm
in diameter, usually about 25 flowers in each. Bracteoles narrowly lan-
ceolate, acuminate, 1 to 2 mm long. Calyx narrowly campanulate, 2
to 2.5 mm long, the teeth ovate, acute, about 1 mm in length. Corolla
slender, pink, the tube about 2 cm long, 1 mm in diameter, the lobes
4, broadly elliptic, spreading, rounded, 4 to 4.5 mm long, 3 to 3.6 mm
in diameter. Anthers 8 mm long. Style slightly exserted, the arms
flattened, 1.5 mm long. Fruit globose, fleshy, white to pink, about 1
cm in diameter, the seeds elliptic in outline, 7 mm long, 5 mm wide.

Necros, Mount Marapara, For. Bur. 13625 Curran & Foxworthy, September,
1909 (type), For. Bur. 13694 Curran; near Cadiz, Bur. Sci. 7327 Celestino, March,
1909. Mrnporo, Mount Halcon, Merrill 5569, November, 1906.

A sylvan species ranging from 500 to 700 m above the sea, well characterized
by its elongated, setaceous stipules, and its long-peduncled, capitulate inflorescence.

Ixora mearnsii sp. noy.

Arbuscula erecta glabra; foliis oblongis vel late oblongo-lanceolatis,
chartaceis vel submembranaceis, acuminatis, basi acutis vel leviter rotun-
datis, petiolatis, usque ad 18 cm longis, in sicco nitidis, nervis utrinque
circiter 11, distinctis; cymis terminalibus, multifloris, densis; floribus
circiter 3 cm longis, calycis segmentis acutis, quam tubus brevioribus.

An erect glabrous shrub. Branches terete, or the younger ones ob-
scurely angled, dark-reddish-brown, shining. Leaves oblong to broadly
oblong-lanceolate, 12 to 18 cm long, 3 to 7% em ‘wide, chartaceous or
submembranaceous, shining on both surfaces, when dry olivaceous above,
paler beneath, the apex very sharply acuminate, the base acute or nar-
rowed and slightly abruptly rounded; nerves about 11 on each side of
the midrib, distinct on the lower surface, anastomosing, the reticulations
lax; petioles about 1 em long; stipules connate, subtruncate, abruptly
and shortly apiculate-acuminate, the margins slightly ciliate, 2 to 3 mm
long. Cymes terminal, shortly peduncled, including the flowers about 7

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. woe

em long, 10 em wide, dense, many-flowered, the branches trichotomously
branched. Flowers apparently pink, mostly in triads on the ultimate
branchlets, the middle one of each triad sessile, the two lateral ones with
pedicels 3 mm long or less; bracts subtending the branches small, ovate,
acuminate, the bracteoles similar, ovate, acuminate, 1 to 1.2 mm long.
Calyx 2 to 3 mm long, the teeth ovate, acuminate or acute, 1 mm long.
Corolla-tube 26 mm long, 1 mm in diameter, the lobes elliptic to elliptic-
oblong, reticulate, membranaceous, acute or minutely acuminate, 7 to 8
mm long, 4 to 5 mm wide. Anthers 3 mm long, abruptly caudate-
apiculate. Stigma about 1 mm long, shghtly exserted.

Luzon, Province of Tayabas, Casiguran, Bur. Sci. 2999 (type), 2976 Mearns,
June 1, 1907. A specimen from Baler, Province of Tayabas, Luzon, Bur. Sov.
10672 McGregor, August, 1909, is similar but has a more lax inflorescence,
' longer flowers, the corolla-tube 3 em in length, and the calyx-teeth are obtuse
or rounded.

A species allied to Iaora congesta Roxb., but with thinner, fewer-nerved leaves,
and more lax inflorescence.

{xora mindanaensis sp. noy.

Arbuscula 2 ad 3 m alta, cymis parce puberulis exceptis glabra;
foliis lanceolatis, oblongo-lanceolatis, vel oblongo-oblanceolatis, coriaceis
vel subcoriaceis, petiolatis, usque ad 16 cm longis, basi acutis vel acu-
minatis, apice acuminatis, nervis utrinque 8 ad 10, distinctis, laxissime
reticulatis vel reticulis subobsoletis ; cymis terminalibus, e basi 3-ramosis ;
floribus circiter 11 mm longis, calycis dentibus parvis, obtusis, quam
tubus brevioribus.

A shrub 2 to 3 m high, erect, branched, glabrous except the inflores-
cence. Branches terete, rather slender, lght-gray. Leaves lanceolate,
oblong-lanceolate, or oblong-oblanceolate, 8 to 16 cm long, 3 to 6 cm
wide, coriaceous or subcoriaceous, when dry shining on both surfaces,
brown, paler beneath, the apex rather sharply acuminate, the base grad-
ually narrowed, acute or somewhat acuminate; nerves 8 to 10 on each
side of the midrib, beneath distinct, usually brown, anastomosing, the
reticulations very lax, often nearly obsolete; petioles about 5 mm long;
stipules ovate to oblong-ovate, acuminate, about 5 mm long, deciduous.
Cymes terminal, somewhat puberulent, branched from the base, the
branches three, 2 cm long or less, each bearing at the apex from three to
five short secondary branches, the flowers all sessile or subsessile, in groups
of threes on the ultimate branchlets, densely disposed, the cymes 5 cm
long or less; bracts very small, obscure, the bracteoles similar, minute,
linear, 0.5 mm long. Calyx 2 to 2.5 mm long, puberulent, the teeth
ovate, obtuse, 0.6 mm long. Corolla-tube white or pinkish, about.9 mm
long, 1 mm in diameter, the lobes elliptic-oblong, 3.6 mm long, 1.5 to
1.8 mm wide, obtuse, 6-nerved, nerves mostly anastomosing. Anthers 3
mm long, acute or obtuse. Stigma slightly exserted, 2 mm long, cleft.

238 MERRILL.

_ ~

Fruit red, depressed-globose, somewhat compressed, about 1 cm wide,
nearly as long, glabrous, smooth, somewhat longitudinally depressed be-
tween the seeds, crowned by the short calyx-rim.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9010 (type), 9034

Whitford & Hutchinson, November 29 and December 2, 1907; Sax River, Williams
2192, February 4, 1905: Province of Misamis, Malabug River, trail to Mount

Iny

Malindang, For. Bur. 4773 Mearns & Hutchinson: Lake Lanao, Camp Keithley,

Mrs. Clemens s. n., May and November, 1906.

Ixora palawanensis sp. nov.

Frutex erectus 2 ad 3 m altus, glaber; foliis subcoriaceis, nitidis, acu-
minatis, lanceolatis vel oblongo-lanceolatis, usque ad 15 em longis, nervis
utrinque circiter 14; cymis terminalibus, dense multifloris; floribus 3.3
ad 3.8 em longis, bibracteolatis, bracteolis parvis; calycis segmentis
ovatis, acutis, quam tubus brevioribus.

An erect shrub 2 to 3 m high, glabrous throughout. Branches terete
or somewhat compressed, smooth, somewhat shining, reddish-brown.
Leaves lanceolate to oblong-lanceolate, rarely oblanceolate, 7 to 15 cm
long, 1.5 to 4.5 em wide, subcoriaceous, somewhat pale when dry,
shining on both surfaces, the base acute or acuminate, apex sharply
acuminate; primary nerves 12 to 15 on each side of the midrib, not
prominent, anastomosing, scarcely more distinct than are the alternating
secondary ones, the reticulations obscure, ‘lax: petioles 0.3 to 0.8 em
long; stipules 5 mm long or less, base broad, apex abruptly contracted,
prominently acuminate. Cymes terminal, dense, 4 to 8 cm wide, the
peduncles 1 em long or less, the lower branches up to 5 em in length,
trichotomously branched, the lower bracts lanceolate, acuminate, about 1
em long, the bracts of the secondary branches much smaller, ovate-oblong,
acuminate, 2.2 mm long: bracteoles 2 at the base of each flower, similar
to the upper bracts but only 1.5 mm long. Flowers salmon- to orange-
colored, numerous, mostly in triads at the ends of the ultimate branch-
lets, the middle one of each triad sessile, the two lateral ones with
pedicels 8 to 8 mm in length. Calyx 3 to 3.3 mm long, the lobes 4,
ovate, acute, 1.5 mm in length. Corolla-tube 3 to 3.5 em long, less
than 1.5 mm in diameter, the lobes oblong-lanceolate, acuminate, reticu-
late, rather thin, about 11 mm long, 3.5 mm wide. Anthers 3 mm long.
Style slightly exserted; stigma cleft. Fruits ovoid, about 1 cm long,
dark-reddish-brown when dry, obtuse or slightly beaked.

PALAWAN, in forests about 14 miles northwest of Iwahig, Bur. Sci. 793 Pox-
worthy, April 22, 1906, in forested ravines.

A species manifestly allied to Jaora congesta Roxb., but with relatively
narrower, smaller leaves which are sharply acuminate, the veins not prominent, ete.

Ixora philippinensis sp, nov.

Arbuscula vel arbor parva, 2 ad 7 m alta, inflorescentiis exceptis glabra ;
foliis subcoriaceis, subsessilibus, oblongo-ovatis, elliptico-ovatis, vel ovatis,

-

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 239

usque ad 15 em longis, in sicco nitidis, plerumque brunneis, breviter
acuminatis vel acutis, basi late rotundatis plerumque distincte cordatis ;
eymis puberulis, terminalibus, paucifloris, breviter pedunculatis, densis ;
floribus 1.8 ad 2.2 cm longis, calycis dentibus minutis, apiculato-acum-
inatis, quam tubus brevioribus.

A shrub or small tree 2 to 7 m high, glabrous except the usually puber-
ulent inflorescence. Branches grayish to reddish-brown, terete. Leaves
oblong-ovate to elliptic-ovate or ovate, shining when dry, usually brownish,
paler beneath, subcoriaceous, 6 to 15 cm long, % to 6 cm wide, rarely
wider, the apex shortly and broadly acuminate, acute, or rarely obtuse,
usually minutely apiculate by the excurrent midrib, the base broadly
rounded, usually distinctly cordate, the uppermost leaves sometimes some-
what surrounding the stems; petioles none or very short, rarely reaching
a length of 3 mm; primary nerves about 11 on each side of the midrib,
slender, usually brown and distinct beneath, anastomosing, the reticula-
tions brown, slender, distinct; stipules lanceolate or ovate-lanceolate,
prominently acuminate, 3 to 5 mm long. Cymes terminal, solitary,
usually somewhat puberulent, their peduncles mostly 1 to 1.5 cm long,
often subtended by a pair of reduced, ovate or suborbicular, cordate leaves
1 to 3 em in length; bracts ovate-lanceolate, acuminate, 1.5 to 1.8 mm
long. Flowers white to pale-pink, shortly pedicelled or sessile, crowded,
forming a dense inflorescence which, excluding the corollas, is less than
1 cm in diameter, 10 or usually less flowers in each cyme, the branches
very short. Calyx 2 mm long, slightly puberulent, the teeth very broad,
minute, abruptly apiculate-acuminate, less than 0.3 mm long, the brac-
teoles lanceolate, acuminate, less than 1 mm long. Corolla-tube 1.8 to
2 em long, usually slightly hairy outside, the lobes coriaceous, elliptic-
oblong, rounded or obtuse, about 7 mm long, 3.5 mm wide. Anthers
3.5 mm long, apiculate. Stigma 2 mm long, cleft, slightly exserted.
Fruit ovoid, apparently red when dry, the pericarp slightly fleshy, glabrous,
slightly or distinctly beaked, nearly 1 cm long when mature, usually
distinctly longitudinally depressed hetween the seeds.

The type of this species is For. Bur. 2299 Meyer, from the Lamao River,
Province of Bataan, Luzon; other specimens from the same locality are Whitford
1270, Leiberg 6118, Williams 378, For. Bur. 2024 Borden, and For. Bur. 1463
Ahern’s collector.

Ixora philippinensis includes most of the Philippine specimens that recently
have been identified as Jvora coccinea Linn., and it appears to be one of the
most widely distributed and abundant species in the Archipelago. Of our
abundant material, for the most part distributed as Jwora coccinea Linn., I
would refer to [xora philippinénsis specimens from the Batanes and Babuyanes
Islands, from the Provinces of Ilocos Norte, Zambales, Pangasinan, Nueva. Keija,
Bulacan, Rizal, Tayabas, and Camarines in Luzon, and from the Islands of Min-
doro, Lubang, Guimaras, Negros, Panay, Palawan, Balabae, and Mindanao.

Ixora philippinensis is manifestly closely allied to /rora coccinea Linn., espe-
cially in its sessile or subsessile, cordate leaves and other vegetative characters,

24() MERRILL.

but differs in its very short calyx-teeth, shorter and usually more slender corolla,
which is white or pale-pink, and in its very small, congested, few-flowered cymes.
In spite of its wide distribution in the Philippines I have been unable to refer
it with any degree of satisfaction to any previously described species.

Var. brevituba var. noy.

A typo differt cymis sessilibus vel brevissime pedunculatis, corollae
tubo vix 1 cm longo.

Luzon, Province of Ilocos Norte, Pasuquin, Por. Bur. 15527 Merritt & Darling,
November 3, 1908, on limestone formation, hillsides, altitude about 100 m.

Ixora coccinea Linn. apparently does not occur in the Philippines except as an
introduced and cultivated plant.

Ixora ‘littoralis sp. noy.

Arbuscula glabra, erecta ; foliis coriaceis, breviter petiolatis, basi leviter
cordatis, apice acutis vel acuminatis, ovatis vel oblongo-ovatis, usque ad
6 cm longis; cymis depauperatis, congestis, paucifloris, breviter pedun-
culatis, axillaribus terminalibusque ; floribus sessilibus vel brevissime pedi-
cellatis, 1.6 ad 1.8 cm longis, calycis lobis acutis, quam tubus brevioribus.

A glabrous shrub. Branches stout, terete, dark-gray. Leaves ovate
to oblong-ovate, 2.5 to 6 cm long, 1 to 3 em wide, subcoriaceous, brown.
and shining when dry, paler beneath, the base rather broad, rounded-
subcordate, apex acute or obscurely acuminate, apiculate; nerves about
8 on each side of the midrib, slender, brown, distinct beneath, the
reticulations rather lax, distinct; petioles about 1 mm long; stipules
lanceolate-acuminate, 3 mm long or less. Cymes axillary and termi-
nating short lateral branches, the peduncles 5 to 6 mm long, subtended
by about three pairs of imbricated, distichous bracts about 2.5 mm
long, their bases broad, abruptly caudate-acuminate, the bracts sub-
tending the few branches ovate-lanceolate, acuminate, 2 mm long, the
bracteoles linear, 1 mm long or less. Cymes few-flowered, congested, the

branches only about 2mm long. Flowers in triads, the middle one sessile

or nearly so, the pedicels of the lateral ones 1 to 2 mm long. Calyx 3
mim long, the teeth broadly triangular-ovate, acute, 0.5 mm long. Corolla-
tube 14 to 16 cm long, 1 mm in diameter, the lobes coriaceous, not
reticulated, mottled, elliptic-ovate, 5 mm long, 3 mm wide, acute or obtuse.
Anthers 3.5 mm long, apiculate. Style slightly exserted ; stigma cleft.

Bono, Tagbilaran, on beach cliffs, Bur. Sci. 1274 McGregor, July, 1906;
flowers white.

A species closely allied to Ixora philippinensis Merr., differing in its smaller
leaves and flowers and frequently axillary inflorescence.

LASIANTHUS Jack.

Lasianthus cyanocarpus Jack in Trans. Linn. Soe. 14 (1823) 125; Hook. f.
rl. Brit. Ind. 3 (1880) 179; F.-Vill. Nov. App. (1880) 112; King in Journ. As.
Soc. Beng. 73? (1904) 113.

Luzon, Province of Cagayan, Bur. Sci. 7406 Ramos, March, 1909.

The specimen agrees closely with the description and with the single specimen

4

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 241

available here for comparison; not previously reported from the Philippines
except for .the unverified record of F.-Villar; India to the Malay Peninsula and
Archipelago.

LUCINAEA DC.

Lucinaea monocephala sp. nov.

Frutex vel arbor glaber; foliis oblongo-ellipticis, brunneis, nitidis,
acuminatis, usque ad 8 cm longis, nervis utrinque circiter 10, tenuibus ;
capitulis axillaribus, solitariis, pedunculatis, circiter 2 cm diametro.

A shrub or tree, glabrous throughout. Branches terete, rugose, gray
or brown. Leaves oblong-elliptic, brown and shining when dry, paler
beneath, subcoriaceous or thickly chartaceous, 5 to 8 cm long, 1.5 to 3.5
em wide, the base acute, the apex shortly acuminate, the lower surface
covered with minute, obscure, whitish, lepidote scales; nerves about 10
on each side of the midrib, slender, not prominent, the reticulations nearly
obsolete; petioles 1 to 1.5 em long. Heads axillary, solitary, brown, the
peduncles stout, 3 to 4 cm long, the heads globose, about 2 cm in diameter.
Corella at least 1 cm long, the petals valvate. Calyx-rim truncate or
subtruncate.

Luzon, Province of Laguna, near Dahican, Bur. Sci. 10034 Ramos, July, 1909
(type). NeEcros, Faraon, For. Bur. 19074 Curran.

Apparently most closely allied to Lucinaea ridleyi King, of the Malay Penin-
sula and Borneo, but differing, according to the description of that species, in
a number of characters.

MUSSAENDA Linn.

Mussaenda albiflora sp. nov.

Frutex erectus 2 ad 5 m altus, omnibus partibus plus minus dense
hirsutis; foliis chartaceis, usque ad 30 em longis, breviter petiolatis,
acuminatis, nervis utrinque 11 ad 15; paniculis terminalibus, floribus
ad apices ramulorum congestis, corollae tubo albo, circiter 2.5 cm longo.

An erect shrub 2 to 5 m high, all parts more or less densely hirsute
with mostly long, spreading, pale or brownish hairs. Branches terete,
reddish-brown or grayish, ultimately glabrous, the young branchlets
densely hirsute. Leaves chartaceous, ovate to oblong-ovate, 11 to 30 cm
long, 6 to 9 cm wide, the apex rather slenderly acuminate, the base
more or less decurrent-acuminate and usually slightly inequilateral, both
surfaces with scattered, spreading, long hairs, either pale or brownish,
and especially dense on the midrib and primary nerves; petioles 1 to 1.5
em long, densely hirsute; stipules lanceolate, acuminate, about 1 cm long,
usually ultimately cleft. Panicles terminal, all parts more or less clothed
with long or short, mostly spreading hairs, the flowers. congested at the
apices of the branchlets, the bracts acuminate, about 7 mm long, the
bracteoles similar, more or less hirsute. Pedicels short, gradually merging
into the slender, slightly hirsute calyx, the calyx-tube about 2 mm in
diameter, the lobes linear-lanceolate, acuminate, hirsute, 6 to 7 mm long,
about 1 mm wide at the base, gradually narrowed upward to the acuminate

242 MERRILL.

apex. Corolla-tube very slender, 2.5 cm long, about 1 mm in diameter,
the antheriferous portion near the apex slightly inflated and about 2 mm
in diameter, this portion also densely bearded inside, the outside with
scattered, short hairs, the lobes linear-lanceolate, 9 to 10 mm long, 2 mm
wide at the base, gradually narrowed upward to the slenderly acuminate
apex, more or less pubescent. Anthers 4 to 5 mm long. Style 4 to 5 mm
long, cleft one-third to one-half into two arms. The persistent leaf-like,
accrescent calyx-lobe is white, elliptic-ovate, ovate, or oblong-ovate,
membranaceous, 6 to 9 em long, 3 to 5.5 cm wide, 5- to 7-nerved from
the base, the nerves more or less hirsute, the reticulations lax, the apex
shortly and sharply acuminate, the base acute or rounded, the stipe about
2 em long. Fruit fleshy, narrowly obovoid, about 1.3 mm long, black
when dry, with few, long, scattered hairs, the calyx-lobes, other than the
acerescent one, early deciduous.

Necros, Faraon, lor. Bur. 17358 (type), 17359 Curran, September, 1909, and
near the same locality, For. Bur. 5539 Hverett, September, 1906, For. Bur. 5218
Danao & Aspillera, June, 1906, in thickets, stream depressions, ete., at low
altitudes, locally known as agboy.

A species well characterized by its long, slender, corolla-tubes which are white
instead of yellow, differing in these characters from all Philippine species known
to me.

Mussaenda philippica A. Rich. in Mém. Soc. Hist. Nat. Paris 5 (1834) 245.

Calycophyllum grandiflorum Meyen Reise 2: 234; Walp. in Nov. Act. Acad. Nat. .

Cur. 19 (1843) Suppl. 1: 356.

Mussaenda grandiflora Rolfe in Journ. Linn. Soe. Bot. 21 (1884) 311; Vid.
Rev. Pl. Vase. Filip. (1886) 152, non Benth. (1849).

Mussaenda frondosa Auct. Philip., non Linn.

There are at present about 60 sheets in the herbarium of the Bureau of Science
that are apparently referable to a single variable species, or perhaps to several

closely allied ones. The material has been identified at various times, some ,

specimens as Mussaenda frondosa Linn., some as MV. grandiflora Rolfe, and some as
M. glabra Vahl. Rolfe in 1884 stated that he had seen no Mussaenda frondosa
Linn., from the Philippines, and I can only agree with him in considering that
the typical form of Linneus’ species does not extend to the Archipelago. The
type of Mussaenda frondosa Linn., was from Ceylon, and the Director of the
Botanic Garden at Peradeniya has kindly supplied me with a full series of
specimens representing the Ceylon plant, presumably some of which are typical
Mussaenda frondosa Linn. None of this Ceylon material matches any of our
Philippine specimens. Mr. Rolfe transferred Calycophyllum grandiflorum Meyen
to Mussaenda, to supply a specific name for the Philippine plant, but overlooked
the fact that the above specific name was invalidated in Mussaenda by the earlier
M. grandiflora Benth. I have accordingly adopted for the Philippine species the
name Mussaenda philippica A. Rich., which was based on material secured in the
Philippines by Perrottet. Whether or not it is the oldest valid name for the
species, I am unable to determine at the present time; some of our Philippine
specimens apparently closely match some Javan and Caroline ‘Islands material
distributed as M. frondosa Linn., and M. glabra Vahl. Mussaenda philippica
A. Rich. is an erect shrub or small tree, not at all scandent, and as at present
interpreted, extends from northern Luzon to southern Mindanao, and will probably
be found to extend to other parts of Malaya.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 243

Mussaenda villosa Wall. Cat. (1832) no. 6254; Hook. f. Fl. Brit. Ind. 3
(1880) 91; King in Journ. As. Soc. Beng. 72? (1903) 184.

What is apparently this species has been collected at Camp Keithley, Lake
Lanao, Mindanao, by Mrs. Clemens, no. 562, and two specimens without number.
The material agrees with Wallich’s species as interpreted by Sir George King l. c.,
except that the pubescence is pale rather than rusty. It is the only scandent
species at present known from the Philippines; widely distributed in’ the Malay
Peninsula.

OLDENLANDIA Linn.

Oldenlandia pterita (Bl.) Miq. Fl. Ind. Bat. 2 (1857) 193.

Hedyotis pterita Bl. Bijdr. (1826) 972.

Gonotheca blumei DC. Prodr. 4 (1830) 429.

Oldenlandia alata Hook. f. Fl. Brit. Ind. 3 (1880) 70; F.-Vill. Nov. App. (1880)
107, non Koenig ex Roxb.

Luzon, Province of Cagayan, Bur. Sci. 7824 Ramos, April, 1909. NEGROS,
Cabancalan, Merrill 6430, March, 1910. PALAWAN, Puerto Princesa, Bur. Sct.
213 Bermejos, December, 1905. MinpANnao, District of Davao, DeVore & Hoover
124, April, 1903, Copeland 596, March, 1904: District of Zamboanga, Hallier
s. n., February, 1904.

Previously credited to the Philippines by Hooker f., and by F.-Villar. What
is apparently the oldest valid specific name is adopted, for according to Hooker
f., the species originally published by Roxburgh, ascribed to Koenig, is a synonym
of Oldenlandia paniculata Linn.

Widely distributed, India to China, and Malaya.

PRISMATOMERIS Thwaites.

Prismatomeris tetrandra (Roxb.) K. Sch. in Engl. & Prantl. Nat. Pflanzen-
fam. 4* (1891) 138.

Coffea tetrandra Roxb. Fl. Ind. 1 (1832) 538.

Prismatomeris albidiflora Thwaites in Hook. Kew Journ. 7 (1855) 268, ¢. 7;
Hook. f. Fl. Brit. Ind. 3 (1880) 159; Trimen Fl. Ceyl. 2 (1894) 355: King in
Journ. As. Soc. Beng. 737 (1904) 90; Valeton in Bull. Inst. Bot. Buitenzorg 8
(1901) 5.

Luzon, Province of Cagayan, Bur. Sci. 7365 Ramos, For. Bur. 13409 Bernardo,
For. Bur. 16592, 17166, 17260 Curran, For. Bur. 14721, 14747 Darling, For. Bur,
18452 Alvarez, February, March, 1909, in forests at from 30 to 200 m altitude.

Some of the specimens cited above differ from the species as described in
having somewhat larger flowers and larger leaves, while others are almost an
exact match for a specimen of Thwaites’ Ceylon plants no. 728 in our herbarium.
All the Luzon material has 5-merous flowers and is much closer to the Ceylon
plant than to several sheets in our herbarium from Perak, Penang, Singapore, and
Java.

No representative of the genus has previously been reported from the Philip-
pines; Ceylon, Burma, Khasia Mountains, Malay Peninsula and Archipelago.

PSYCHOTRIA Linn.

Psychotria phanerophlebia sp. noy.

Frutex erectus, ramulis, inflorescentiis, subtus foliis, plus minus dense
castaneo- vel subrubiginoso-pubescentibus ; foliis coriaceis, late oblongo-
oblanceolatis, usque ad 29 cm longis, nervis utrinque cireciter 22, pro-
minentibus ; cymis dense congestis, pedunculatis.

244 MERRILL.

An erect shrub. Branches terete, stout, very densely dark-brown-
pubescent. Leaves broadly oblong-oblanceolate, 23 to 29 cm long, 7 to
8 em wide, the apex acute or very obscurely acuminate, gradually nar-
rowed from about the middle to the acute base, margins recurved, coria-
ceous, reddish-brown when dry, glabrous and shining on the upper surface,
beneath, especially on the midrib and lateral nerves, rather densely
pubescent with short, dark-brown hairs; nerves about 22 on each side
of the midrib, very prominent beneath, anastomosing very close to the
margin, somewhat ascending, the primary reticulations subparallel ;
petioles densely pubescent, stout, about 3 cm long. Cymes in the upper
axils, densely dark-brown-pubescent, three or more on each branchlet,
their peduncles stout, about as long as the petioles, the branches short,
crowded, the primary ones not exceeding 1 cm in length, the flowers
densely congested, the inflorescence 3 cm or less in diameter. Flowers
5-merous, their pedicels stout, 1 to 3 mm long, densely pubescent, the
bracts few, linear-lanceolate, acuminate, about 5 mm long. Calyx pubes-
cent, the tube 4-angled, 4-suleate, the rim above the ovary somewhat
spreading, 3.5 to 4 mm long, the lobes 5, rarely 6, lanceolate, acuminate,
2 to 2.5 mm long. Corolla unknown.

Luzon, Province of Cagayan, near Pamplona, Bur. Sci. 7404 Ramos, March 17,
1909. ;

A species well characterized by its dark-brown pubescence, its congested cymes,

and by its very prominently nerved, elongated leaves. It is quite different from
all other Philippine forms known to me.

Psychotria ramosii sp. nov.

Arbuscula circiter 2 m alta, glabra, partibus junioribus inflorescen-
tiisque molliter sublanato-pubescentibus exceptis; foliis membranaceis,
oblongo-ellipticis vel oblongo-obovatis, in sicco nitidis, pallidis, apice
acuminatis, basi angustatis, nervis utrinque circiter 12; cymis axillaribus,
pedunculatis, paucifloris, plus minus dense pallide sublanato-pubescenti-
bus.

A shrub about 2 m high. Branches terete, glabrous, smooth, dark-
colored. Leaves membranaceous, oblong-elliptic to oblong-obovate, 10
to 12 cm long, 4 to 5 cm wide, pale and shining when dry, the apex
shortly and obtusely acuminate, the base gradually narrowed, acute or
acuminate; nerves about 12 on each side of the midrib, prominent
beneath, the midrib and lateral nerves with weak, scattered, crisped
hairs; petioles about 2 cm long; stipules deciduous, broadly ovate, pubes-
cent, about 5 mm long. Cymes in the upper axils, several from each
branchlet, the peduncles slender, 2 to 3 em long, all parts more or less
densely covered with weak, soft, pale, somewhat woolly hairs. Flowers
white, their pedicels:1 to 2 mm long, densely pubescent. Calyx 4 to 5
mm long, densely pubescent, the lobes. 5, ovate, acute or acuminate, 2 mm

te

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 245

Jong. Corolla glabrous, the tube cylindric, 3 mm long, the lobes 5, more
or less spreading, coriaceous, oblong-ovate, 3 mm long, acute or obtuse.
Anthers about 1 mm long. Style and stigma together 2.5 mm long.
Ovary 2-celled, each cell with a single ovule.

Luzon, Province of Cagayan, near Pamplona, Bur. Sci. 7499 Ramos, March 17,
1909.

A species differing from all Philippine forms known to me in its sublanate

inflorescence.
RANDIA Linn.

Randia ticaensis sp. nov.

Arbor cireiter 6 m alta, glabra; ramulis teretibus, foliis breviter petiol-
atis, oblongis vel oblongo-ovatis, chartaceis, acuminatis, basi angustatis,
leviter cordatis, nervis utrinque circiter 9; floribus axillaribus, solitariis
vel fasciculatis, calycis lobis sub fructu lanceolato-acuminatis, circiter 4
mm. longis.

A glabrous tree about 6 m high. Branches terete, grayish, slender.
Leaves oblong or oblong-ovate, chartaceous, brown and only slightly
shining when dry, 10 to 13 cm long, 3 to 5.5 em wide, the apex acu-
minate, the base somewhat narrowed and then abruptly rounded-sub-
cordate; nerves about 9 on each side of the midrib, distinct, somewhat
ascending, obscurely anastomosing, the reticulations faint, lax; petioles
about 2 mm long; stipules setaceous or lanceolate-acuminate, 4 to 4.5
mm long. Flowers unknown, but the fruits axillary, solitary or two in
an axil, very shortly pedicelled, black when dry, globose, about 11 mm
in diameter (not quite mature), glabrous, the calyx-tube subpersistent,
cylindric, slightly puberulent, 3 mm long, with five spreading, lanceolate-
acuminate, 4 mm long, 3-nerved lobes, the tube ultimately deciduous.

Ticao, Linadlaran Point, For. Bur, 12547 Rosenbluth, December 24, 1908, on

steep hillsides at an altitude of about 30 m, locally known as turutulang. Allied
to R. cumingiana Vid., but quite distinct.

Randia stenophylla sp. noy.

Frutex vel arbor parvus, plus minus puberulus; foliis lanceolatis vel
anguste lanceolatis, subcoriaceis, basi acutis, apice acuminatis, usque ad
8 cm longis, 1.2 cm latis; cymis axillaribus, pedunculatis, vel floribus
solitariis; floribus circiter 5 mm longis, 5-meris; fructibus carnosis, glo-
bosis vel ovoideis, circiter 1 cm diametro.

An erect shrub or small tree, the branches, leaves and inflorescence
more or less grayish-puberulent. Branches slender, terete. Leaves lance-
olate or narrowly lanceolate, 3.5 to 8 em long, 0.5 to 1.2 em wide,
subcoriaceous, scarcely shining when dry, brown, ultimately glabrous or
nearly so on the upper surface, base acute, apex acuminate ; lateral nerves
faint, about 13 on each side of the midrib, sometimes nearly obsolete ;
petioles puberulent, 1 to 2 mm long; stipules ovate, acuminate, puberu-

246 ‘ MERRILL.

lent, 2 to 3 mm long. Flowers axillary, solitary, or two on a more or
less elongated peduncle, the peduncle, when present, 12 mm long or less,
puberulent, with 2, opposite, lanceolate-acuminate, puberulent, 3 mm
long bracts at the apex, subtending two flowers. Flowers sessile or
shortly pedicelled. Calyx puberulent, the tube funnel-shaped, about 2
mm long, the lobes more or less spreading, 5, ovate-oblong, acuminate,
2.5 mm long. Corolla-tube 3 mm long, villous within, the lobes oblong,
3 to 3.5 mm long, acute or acuminate, recurved or spreading. Anthers
2.5 mm long, scarcely exserted. Style 2 mm long, glabrous; stigma
cleft, 2.5 to 3 mm long, densely villous. Fruit red, soft and fleshy,
apparently globose or ovoid and about 1 cm in diameter, the seeds
numerous, ovoid, about 3 mm long.

Luzon, Province of Bulacan, Norzagaray, on rocky river banks, For. Bur. 7170
Curran, June 16, 1907.

A species well characterized by its very narrow leaves and small, axillary,
solitary or paired flowers, which may be sessile, or pedunculate; probably most
closely allied to Randia angatensis ¥.-Vill., but quite different from that species.

TIMONIUS DC.

Timonius macrophyllus sp. nov.

Arbor circiter 10 m alta, partibus junioribus, subtus foliis, ramulis
inflorescentiisque plus minus fulvo-hirsuto-villosis; foliis coriaceis, ob-
oyatis, usque ad 23 cm longis, apice rotundatis, basi acutis, nervis utrinque
8 vel 9, prominentibus; inflorescentiis brevibus, dense pubescentibus,
floribus 4-meris, secundis, spicatis; fructibus subovoideis, circiter 7 mm
longis, leviter longitudinaliter 4-sulcatis, pyrenis 25 ad 30.

A tree about 10 m high. Branches light-gray, subterete, rather stout,
apical portions more or less compressed and pubescent. Leaves obovate,
17 to 23 em long, 12 to 14 cm wide, the apex rounded, the base acute,
coriaceous, brown and shining when dry, glabrous on the upper surface,
beneath sparingly hirsute-villous, especially on the midrib and lateral
nerves; nerves 8 or 9 on each side of the midrib, prominent, curved-
anastomosing, the primary reticulations curyed, subparallel, prominent ;
petioles stout, about 1 cm long. Inflorescence in the upper axils, at
most 7 em long, consisting of two or three branches 2 to 3 em in length
or less, densely fulvous-villous, the branches bearing numerous, crowded,
sessile flowers along one side. Flowers 4-merous, cylindric. Calyx 3
mm long, the rim produced about 1.5 mm above the ovary, truncate,
densely fulvous-villous outside. Corolla (in bud) 3.5 mm long, the four
lobes oblong, obtuse, 3 mm long. Anthers 4, alternating with the corolla-
lobes, 3 mm long. Style 4-angled, cleft into four arms about 2 mm in
length, each arm again minutely cleft at the apex. Ovary 4-celled, each
cell with from 5 to 7, rarely more locelli. Fruit subovoid, about 7 mm
long, 6 mm in diameter, slightly hairy, with four rounded angles and

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 247

somewhat longitudinally 4-sulcate, 4-celled, each cell with from 5 to 7
or more pyrenes. ; ,

Samar, near Catbalogan, For. Bur. 12856 Rosenbluth, February 7, 1909,
hillsides along streams at an altitude of about 200 m, locally known as canilan.

A species well characterized by its ample leaves, its secund spicate flowers and
its pubescent younger parts, inflorescence, and leaves. In its foliage it is some-
what suggestive of Timonius stipulosus Val., but is very distinct from that
species.

UROPHYLLUM Wall.

Urophylilum elliptifolium sp. nov.

Arbuscula erecta circiter 3 m alta, partibus junioribus plus minus
furfuraceis exceptis glabra ; foliis ellipticis vel oblongo-ellipticis, coriaceis,
nitidis, basi acutis vel acuminatis, apice rotundatis vel breviter late
acuminatis, nervis utrinque circiter 12, subtus prominentibus; umbellis
axillaribus, solitariis, breviter pedunculatis; fructibus paucis, carnosis,
ovoideis, circiter 7 mm longis.

An erect shrub about 3 m high. Branches brownish or grayish, terete,
or the younger ones somewhat compressed, the tips of the branches, leaf-
axils, stipules, and petioles of the younger leaves with few to many, thin,
small, ultimately deciduous, appressed, pale scales. Leaves elliptie to
oblong-elliptic, coriaceous, shining, rather pale when dry, 8 to 12 em long,
3.5 to 6 cm wide, the base acute or somewhat acuminate, the apex rounded .
or shortly and broadly acuminate; primary nerves about 12, prominent
beneath, spreading, curved, anastomosing, the alternating secondary ones
also prominent, the reticulations distinct; petioles 1.5 to 3 em long;
stipules oblong, rounded, 1.5 cm long, furfuraceous, deciduous. Flowers
unknown. Fruit in solitary, axillary umbels, the peduncles about 8 mm
long, with few small bracts at the apex, each peduncle bearing from
three to five ovoid fruits about 7 mm long, 5 mm in diameter, glabrous,
their ‘pedicels about as long as the peduncles. Seeds very numerous,
pale-brown, 0.5 mm long, densely and finely foveolate.

PALAWAN, Mount Pulgar, For. Bur. 3871 Curran, February, 1906, on forested
slopes, altitude 700 to 1,300 m.

Possibly as closely allied to the Bornean Urophyllum subaneurum Stapf as
to any other species, but quite distinct from that.

Urophyllum negrosense sp. nov.

Arbuscula erecta, ramulis foliis subtus ad nervos, stipulis bracteisque ©
plus minus ciliato-hirsutis; foliis late oblongo-lanceolatis, submembrana-
ceils, usque ad 18 cm longis, apice sensim subcaudato-acuminatis, basi
acutis, nervis utrinque circiter 16; stipulis oblongis, membranaceis, usque
ad 2.5 cm longis; floribus axillaribus, sessilibus, fasciculatis.

An erect shrub (fide Everett). Branches terete, brownish, glabrous or
nearly so. Leaves broadly oblong-lanceolate, submembranaceous, 15 to

95948 6

248 MERRILL.

18 cm long, 4 to 5 em wide, the apex gradually narrowed to the slender,
subcaudate acumen, the base acute, rather pale when dry, shining on both
surfaces, glabrous on the upper surface, beneath with numerous, pale,
spreading or somewhat appressed ciliate hairs on the midrib and nerves,
otherwise glabrous ; nerves about 16 on each side of the midrib, prominent,
curved-ascending, anastomosing near the margin, the primary reticula-
tions distinct, subparallel; petioles 1 to 1.5 em ice glabrous or slightly
ciliate; stipules membranaceous, oblong, subpersistent, 2 to 2.5 cm long,
rather densely ciliate on the back. Flowers few, axillary, fascicled,
apparently sessile. Fruit fleshy, ovoid, about 1.3 cm long, somewhat
villous,, crowned by the ovate, obtuse calyx-lobes which are more or less
villous and 5 to 6 mm long.

Necros, Himugaan River, in ravines at an altitude of about 50 m, For. Bur.
5550 Everett (type), October 25, 1906, the fruit green when collected; said to be
abundant locally; Faraon, For. Bur. 13574 Meyer & Foxworthy, August, 1909.

A species probably closely allied to Urophyllum streptopodium Wall., but with
quite different leaves and much larger persistent calyx-lobes.

VILLARIA ‘Rolfe.

Villaria acutifolia (Elmer) comb. nov.

Gardenia acutifolia Elmer Leafl. Philip. Bot. 1 (

MINDANAO, District of Davao, Davao, Copeland 43
2975.

The original description of this species was based on two specimens, the first
one cited being Copeland 437, which I assume to be the type of the species. The
second specimen cited, Ahern 457, “457, Forestry Bureau, collected by J. F. Qua-
dras,” from Dinagat Island, is also a Villaria, and is probably referable to
V. philippinensis Rolfe. The original description must be emended as follows:
Leaves chartaceous, apex acuminate, base rounded or acute. Flowers mostly
solitary, axillary, rarely in short, 3-flowered, cymes, the inflorescence and calyx
puberulent (not glabrous), the peduncles in fruit not exceeding 2 em in length,
each subtended by two lanceolate, acuminate, puberulent bracts, no bracts or
bracteoles above the base. Calyx puberulent, up to 1.5 em long, sometimes
4-merous. The flowers were apparently described from immature buds, but the
dissected material was not preserved, and there are no buds or open flowers left
on the type sheet. Open flowers on Mr. Williams’ specimen are white, the corolla
tube cylindric, 4 mm long, the lobes 4 or 5, elliptic, rounded, 4 mm long, 2.5 mm
wide, the throat pubescent. Anthers 4 mm long, inserted on the throat, not
exserted; style 2 mm long, glabrous; stigma oblong, 4 mm long, 1.5 mm wide,
felted-pubescent. The description of the fruit must be excluded as it was based
on Ahern 457, and refers to Villaria philippinensis. —

The species is well characterized by its usually solitary flowers, the pedicels
bibracteate at the base, the inflorescence more or less puberulent, and especially by
the calyx-lobes exceeding the corolla in length.

The form described by Mr. Elmer, 1. ¢., as Gardenia elliptica is exactly the
same as Villaria littoralis Vidal.

1906) 6.
7 (type); Padada, Williams

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 249

GOODENOVIACE.
‘SCAEVOLA Linn.

Scaevola. acuminatissima sp. nov. § Hnantiophyllum.,

Scandens, glabra, vel inflorescentiis plus minus pubescentibus; foliis
oppositis, ovato-lanceolatis vel late oblongo-lanceolatis, membranaceis,
nitidis, leviter distanter denticulatis, basi acutis, apice longissime caudato-
acuminatis, usque ad 11 cm longis; pedunculis axillaribus, brevibus,
saepe trifloris ; floribus aurantiacis, 5-meris, circiter 2.5 em longis.

A scandent herbaceous vine, nearly glabrous throughout, the stems
brownish, slightly striate, up to 3.5 mm in diameter, the branches and
leaves opposite. Leaves ovate-lanceolate to broadly oblong-lanceolate,
membranaceous, shining, entirely glabrous, 6 to 11 cm long, 2 to 3 em
wide, the base acute, the apex gradually narrowed into a long, slender,
straight or somewhat falcate, caudate acumen, the margins entire or
distantly and slightly denticulate; nerves about 6 on each side of the
midrib, slender, anastomosing, the reticulations lax; petioles 5 mm long
or less. Peduncles axillary, solitary, 5 to 10 mm long, more or less
appressed-pubescent, the axils of the stems at the insertion of the pedun-
cles also usually pubescent, each peduncle bearing 3, rarely 5 flowers, the
pedicels 5 mm long or less, the bracts at the apex of the peduncles narrow,
up to 4 mm in length. Flowers yellow. Calyx-tube oblong, glabrous or
nearly so, in anthesis 2 to 3 mm long, the lobes 5, lanceolate, acuminate,
about 3.5mm long, 1mm wide. Corolla 2 to 2.2 em long, nearly glabrous
outside or with very few, scattered, appressed hairs, villous within, the
lobes 5, lanceolate or oblong-lanceolate, acuminate, 6 to 9 mm long, 1.8
to 2 mm wide, 3-nerved, the corolla-tube 15-nerved. Filaments slender,
glabrous, 8 to 9 mm long; anthers about 2 mm in length. Style glabrous,
13 mm long, the stigma flattened, 2 to 2.5 mm wide, slightly horned at
the upper corners, surrounded by a cup 2 to 2.5 mm in diameter which is
densely ciliate on the margins. Fruits oblong-ellipsoid, about 9 mm long,
3.5 mm in diameter, somewhat longitudinally sulcate, black when dry,
glabrous.

Mrnporo, Arunay River, For. Bur. 12125 (type), 12133 Merritt, May 8, 1908,
in forests at an altitude of about 600 m.

A species allied to the Philippine Scaevola dajocnsis Merr., of Jolo, S. mina-
hassae Koord., of Mindanao and Celebes, 8. similis Hemsl., of Celebes, S. novo-
guineensis K. Schum., of New Guinea, S. oppositifolia Roxb., of Ternate, and
S. amboinensis Miq., of Amboina. Specimens sent to Kew for comparison were
reported as “nearest 8. novo-guineensis, differing from it in having larger flowers

and an almost glabrous corolla.” Among the Philippine species it is manifestly
allied to 8. dajoensis Merr., but differs in its much larger flowers.

250 MERRILL.

Scaevola mindorensis sp. nov. § Hnantiophyllum.

Species praecedenti valde affinis, differt foliis pro rata latioribus, mar-
gine distincte sinuato-dentatis, subtus plus minus pubescentibus, ramulis
leviter pilosis, inflorescentiis floribusque dense pubescentibus.

A scandent herbaceous vine similar to Scaevola acuminatissima, differ-
ing in the points above indicated. Branches brown or grayish, striate,
pilose, the younger ones rather densely so. Leaves ovate-oblong, mem-
branaceous, 5 to 7 em long, 2 to 3 em wide, base acute, apex slenderly
subcaudate-acuminate, margins distinctly sinuate-dentate, the lower sur-
face with numerous, short, scattered, spreading hairs, the upper surface
glabrous or with very few hairs. Peduncles axillary, solitary, 1 to 2 cm
long, densely pubescent, each usually bearing three flowers, and also two
much reduced leaves at the apex, the pedicels about 5 mm in length, the
bracteoles 1.5 mm long. Flowers yellow, 5-merous. Calyx-tube 4 mm
long, rather densely pubescent, the lobes 5, lanceolate, about 5 mm long,
1.2 to 1.4 mm wide, acuminate. Corolla outside rather strongly pubes-
cent, villous within, 2.2 em long, the lobes 8 mm long, 2 to 2.5 mm wide,
3-nerved, the tube 15-nerved. Filamants glabrous. Style glabrous, 13
mm long, the cup surrounding the stigma not only densely ciliate on the
margins, but also with numerous long white cilia on the outside.

Minporo, Mount Sablayan, For. Bur. 9756 Merritt, March 2, 1908, on the
exposed cleared summit at an altitude of about 1,000 m.

Specimens sent to Kew for comparison were reported as “nearest S. similis

Hemsl., but with longer peduncles.” It differs also from that species, as described,
in its 5-merous, not 4-merous, much larger flowers and its different calyx-teeth.

Scaevola sericea Forst. Prodr. (1786) 89; Pres] Rel. Haenk. 2 (1830) 57;
DC. Prodr. 7 (1839) 506.

Luzon, Haenke in Herb. Mus. Kénigr. Béhmen, Prague: Province of Ilocos
Sur, Salomague, Merrill 339.

This species is apparently much less common in the Philippines than is
S. koenigii Vahl, and is to me sufficiently distinct from Vahl’s species to warrant
being given specific rank, although recent authors have treated it as a synonym
of S. koenigii Vahl. It differs from S. koenigii in being pubescent throughout, the
inflorescence very densely so. I have what is apparently the same form from
Java, and from the Caroline Islands, Yap, Volkens 133, distributed as S. koenigit
Vahl.

Scaevola micrantha Presl Rel. Haenk. 2 (1830) 59; Miq. Fl. Ind. Bat. 2
(1856) 582; F.-Vill: Nov. App. (1880) 121.

Luzon, Province of Albay, on barren rocky hills, altitude about 120 m, near
Calanaga, Batan Island, Bur. Sci. 6289 Robinson, August 23, 1908. I have also
examined the type in the herbarium of the Museum des Kénigreichs Béhmen,
Prague, and find it to be quite the same as the specimen collected by Doctor
Robinson. Haenke’s specimen probably came from what is now the Province
of Albay, or from Sorsogon.

The species is a very distinct one, as indicated by Presl. It is, perhaps, most
closely allied to S. plumieri Vahl, but its flowers are less than | em in length.

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 251

Scaevola pedunculata sp. nov.

Species 8. micranthae affinis, sed differt ramulis foliis inflorescentiisque
glabris, axillis barbatis exceptis, foliis longioribus, cymis longe peduncu-
latis, floribus paulo longioribus.

A shrub 2 to 4 m high, erect, nearly glabrous, axils excepted. Branches
terete, smooth, olivaceous, glabrous, the axils of the leaves and pedun-
cles densely bearded with long white hairs. Leaves distant, scat-
tered, chartaceous or submembranaceous, narrowly oblong-obovate or
oblong-oblanceolate, glabrous and shining on both surfaces, 6 to 10 cm
long, 1.5 to 3 em wide, the apex broad, rounded or very slightly and
obscurely acuminate, gradually narrowed from the upper third to the
base, the petiolar part about 1 cm long; nerves about 8 on each side of
the midrib, indistinct. Cymes axillary, solitary, as long as the leaves,
glabrous except the densely bearded axils of the branches and bracts,
the peduncles about 5 cm long, each bearing at its apex two linear-lance-
olate to subspatulate 1 to 1.5 cm long bracts, and four primary branches
about 1.5 em long, the branches in turn bracteate at their apices and
bearing usually four shorter branchlets, the ultimate ones bearing usually
three flowers, a central sessile one, and two lateral pedicellate ones, the
bracteoles about 5 mm long. Flowers white. Calyx 2.5 mm long, gla-
brous, the five teeth broadly ovate, acute, about 1 mm long. Corolla 1
em long, slightly curved, somewhat pubescent outside but not densely so,
pilose inside, the lobes about 4 mm long, with broad, thin, infolded
margins. Style slightly silky, the indusium surrounding the stigma
densely ciliate. Drupe glabrous, obscurely costate, about 3 mm long.

PALAWAN, on rocky river banks, altitude about 175 m, Mount Victoria, Bur.
Sci. 744 Foxworthy, March 25, 1906.

A species manifestly allied to S. micrantha Presl, but apparently sufficiently
distinct, recognizable by its somewhat larger leaves, long peduncled cymes, the

axils of the peduncles, leaves, bracts and bracteoles densely bearded with long
white hairs, the plant otherwise glabrous or nearly so.

Scaevola pedunculata var. mollis var. noy.

A typo differt omnibus partibus dense breviter pubescentibus.

PALAWAN, Mount Victoria, altitude about 1,000 m, Bur. Sci. 700 Foxworthy,
March 23, 1906.

The specimen on which the above variety is based in all essential characters
is the same as the type, differing in being softly and rather densely pubescent
throughout with short grayish hairs. The flowers appear to be quite the same
as in the species, but the fruits are slightly pubescent. Additional material may
show this form to be worthy of specific rank, but it is considered best for the
present to consider it as a variety only. It is apparently more closely allied to
8. micrantha Pres] than is the species, but differs from Presl’s species in all the
characters indicated for 8. pedunculata except in its pubescence, and in this
it is very decidedly more pubescent than is S. micrantha Presl. Logically, if
Scaevola sericea Forst. is to be considered specifically distinct from 8. koenigti

252 MERRILL.

Vahl, then perhaps the present variety should be given specific rank. However,
the material on which the species and the variety are based, came from the
same region, although at different altitudes, and was collected on the same date,
and the pubescence of the latter may be a character largely due to altitude.

COMPOSITE.
VERNONIA Schreb.

Vernonia elmeri sp. nov.

Gynura angulosa Elmer Leafl. Philip. Bot. 1 (1906) 146, excel. syn., non DC.

Herba scandens, usque ad 3 m alta, foliis subtus, ramulis, inflorescen-
tiisque leviter pubescentibus; foliis alternis, petiolatis, oblongo-ovatis vel
lanceolato-ovatis, valde acuminatis, margine distanter denticulatis; in-
florescentiis terminalibus, corymboso-paniculatis, capitulis pedunculatis,
cireiter 1 cm longis; floribus purpureis.

A scandent herbaceous plant reaching a height of at least 3m. Stems
and branches terete; striate, appressed-pubescent with grayish hairs.
Leaves alternate, oblong-ovate to lanceolate-ovate, or the upper ones
nearly lanceolate, 4 to 8 em long, 1 to 3 cm wide, the upper ones often
smaller, chartaceous or submembranaceous, gradually narrowed into the
sharply acuminate apex, the base acute or broad and rounded, the mar-
gins with distant, small teeth, somewhat pubescent with scattered hairs
on both surfaces, or nearly glabrous above, glandular-punctate beneath ;
petioles 3 to 4 mm long, pubescent. Panicles terminal, corymbose,
slightly pubescent. Heads comparatively few, about 1 cm long. Involu-
eral bracts 5- or 6-seriate, the outer ones gradually smaller, and the outer-
most almost linear, 1 to 1.5 mm long, all pubescent, the innermost ones
about 5 mm long, 1 to 1.8 mm wide, apiculate-acuminate, slightly keeled.
Disk at first paleaceous with short scales, ultimately quite glabrous.
Flowers all hermaphrodite, homogamous, tubular, purple. Achenes about
1 mm long, slightly pubescent, obscurely ribbed; pappus white, copious,
about 6 mm long, with a few very short hairs in the outer series. Corolla
about 9 mm long, cleft at the apex into 5, oblong-lanceolate, about 3 mm
long lobes; style exserted, the arms 2 mm long; anthers 2.5 mm long,
apex blunt or acute, hyaline, base shortly cleft.

PALAWAN, Separation Point, Merrill 793, February, 1903; San Antonio Bay,
Merrill 5256, October, 1906; Mount Victoria, Bur. Sci. 703 Foxworthy, March,
1905. Luzon, Province of Nueva Vizeaya, Bur. Sct. 8196 Ramos, May, 1909,
MrnpANAo, Lake Lanao, Mrs. Clemens 915, January, 1907.

This species is apparently closely allied to Vernonia cinerea (L.) Less., but is
at once distinguishable by its much larger size, scandent habit, and larger leaves
and heads. In floral structure it is exceedingly similar to that species. It was
referred by Mr. Elmer to Gynura angulosa DC., but does not remotely resemble
that species, and is, moreover, not a member of the Senecioneae. The structure of
the involuere at once distinguishes it from Gynuwra and allied genera. More ma-
terial is necessary to dispose of Gynura sarmentosa Elm., l. c. (non DC.!), as the
specimen referred to it by that author, Copeland 1258, is very similar to some

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 253

forms of the species above described, and is certainly cogeneric, if not cospecific
with it; the specimen is, however, young, and additional material may show more
marked points of differentiation.

Vernonia acrophila sp. nov. § Strobocalyx.

Arbor parva, circiter 5 m alta, subglabra; foliis coriaceis, oblongo-
ovatis, acuminatis, circiter 3.5 cm longis; corymbis terminalibus, plus
minus congestis, capitulis 3- vel 4-floris, squamis imbricatis, plus minus
puberulis, margine obscure ciliatis; achenio 2 mm longo, glanduloso.

A small tree about 5 m high, the trunk 15 cm in diameter, subglabrous.
Branches short, stiff, the ultimate ones more or less crowded, black or
grayish, lenticellate, glabrous or nearly so. Leaves oblong-ovate, coria-
ceous, about 3.5 cm long, 1.2 to 1.5 cm wide, the apex shortly acuminate,
acumen blunt, base acute or acuminate, the margins somewhat reflexed,
glabrous, dark-colored when dry ; nerves 5 or 6 on each side of the midrib,
distant, anastomosing, distinct beneath ; petioles slender, 4 to 6 mm long.
Inflorescence terminal, somewhat congested, subglabrous. Involucral
bracts several-seriate, the outer ones ovate, obtuse, 1 mm long or less, the
inner ones gradually longer, the innermost oblong, 2.5 to 3 mm long and
deciduous, all slightly puberulent or nearly glabrous, the margins ob-
scurely ciliate. Achenes 3 or 4 in each involucre, 2 mm long, irregularly
and obscurely angled, and with numerous, scattered, waxy glands; pappus-
hairs stiff, scabrid, about 24, 4 mm long, with a number of shorter sup-
plementary ones intermixed, these 1 mm long or less.

Luzon, Province of Zambales, Mount Tapulao, For. Bur. 8057 Curran &
Merritt, October 13, 1907, in the elfinwood, exposed peaks, above an altitude of
2,000 m.

As Vernonia arborea Ham. is interpreted by Hooker f., perhaps the present
form would be included, possibly as a variety or as a reduced form. It differs so
strongly in its very much reduced leaves and in being nearly glabrous throughout,
that it has been considered to be worthy of specific rank.

Vernonia lancifolia sp. nov. § Strobocalya.

Vernonia arborea Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 138, non Ham.

Arbor subglabra 4 ad 5 m alta; foliis lanceolatis, valde acuminatis,
usque ad 12 cm longis, 2.5 cm latis, coriaceis vel subcoriaceis, glabris,
subtus glandulosis, nervis utrinque 9 ad 12, distinctis; squamis puber-
ulis; achenio 2 ad 2.4 mm longo.

A tree 4 to 5 m high, nearly glabrous throughout. Branches grayish
to nearly black, terete, glabrous, the ultimate branchlets sometimes pubes-
cent. Leaves coriaceous, lanceolate, 5 to 12 em long, 1.5 to 2.5 cm wide,
glabrous, somewhat shining above when dry, the apex sharply subcaudate-
acuminate, the base acute or acuminate, the lower surface with numerous,
minute, yellow, shining, waxy glands; nerves 9 to 12 on each side of the
midrib, beneath distinct, anastomosing, the reticulations rather lax ; peti-
oles slender, 3 to 10 mm long. Inflorescence terminal, 6 to 12 cm in
diameter, glabrous or nearly so, or the branchlets somewhat pubescent.

254 MERRILL.

Outer involucral scales suborbicular, less than 1 mm long, pubescent, the
inner ones gradually longer, the innermost less pubescent, deciduous, 2 to
2.5 mm long, obtuse. Achenes 3 or 4 in each involucre, when mature
nearly terete, when young more distinctly angled, 2 to 2.4 mm long,
sometimes distinctly waxy-glandular, at other times with very few,
scattered, short hairs; pappus bristles stiff, about 30, 4 to 4.5 mm long,
with numerous shorter ones less than 1 mm long intermixed.

Luzon, Province of Bataan, Mount Mariveles, Merrill 3200, Williams 420
(type), Elmer 6695, Whitford s. n., summit of the mountain, altitude about
1,400 m: Province of Tayabas, Mount Malaraya, For. Bur. 7841, 7844 Curran &
Merritt, November, 1907, altitude about 1,000 m.

A species like the preceding allied to Vernonia arborea Ham., differing in being
nearly glabrous and especially in its much narrower, smaller, lanceolate leaves

which are glandular beneath. It is manifestly allied to the preceding species but
has very much Jarger leaves.

BLUMEA DC.

Blumea confertiflora sp. nov.

Herba erecta, robusta, circiter 1 m alta; foliis sessilibus, basi angus-
tatis, plus minus decurrentibus, oblongo-ellipticis vel oblongo-obovatis,
usque ad 12 cm longis, subcoriaceis, breviter acuminatis, margine irreg-
ulariter sinuato-denticulatis ; capitulis circiter 1 em diametro, in ramulis
ultimis densissime congestis, sessilibus vel breviter pedunculatis.

An erect, robust herb about 1 m high. Stems stout, dark-brown or
purplish, longitudinally striate, more or less pubescent, apparently un-
branched, except the inflorescence. Leaves sessile, oblong-elliptic to
oblong-obovate, the apex shortly acuminate, the base gradually narrowed,
the margins irregularly sinuate-denticulate with small teeth, not at all
lobed, 6 to 12 cm long, 2.5 to 4 em wide, coriaceous or subcoriaceous,
slightly shining when dry, somewhat pubescent with short hairs on both
surfaces, paler beneath; nerves about 9 on each side of the midrib; leaf-
bases sometimes decurrent along the stem for 2 to 3 em. Panicle-branches
rather few, the lower ones up to 40 em long and subtended by leaves,
ascending, the secondary ones very short, mostly less than 1 cm long,
pubescent, each bearing numerous, densely disposed heads which are ses-
sile or shortly peduncled. Heads 8 to 9 mm long, about 1 em in diameter.
Involucral bracts many-seriate, the outer ones oblong-ovate to oblong-
lanceolate, about 2 mm long, 1.2 mm wide, pubescent, the inner ones
gradually longer, the innermost about 6 mm long, 0.5 to 1 mm wide,
slightly hairy, acuminate, the margins toward the apex more or less
ciliate. Flowers yellow, heterogamous, the outer ones indefinite (more
than 100), pistillate; achenes 1 to 1.2 mm long, somewhat angled, slightly
hairy ; pappus hairs about 20, white, slender, scabrid, 5 mm long; corolla
cylindric, slender, 5 mm long, minutely 3-toothed; style-arms slender,
exserted, less than 1 mm long. Disk-flowers perfect, about 14 in each

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 250

head; corolla 5 mm long, enlarged above, the teeth 5, ovate, acute or
obtuse, less than 0.5 mm long; anthers 2 mm long, the tails very slender,
less than 0.55 mm in length; style-arms stout, blunt, 0.5 mm long. Disk
pitted, glabrous or with very few, short, scattered hairs.

Minporo, For. Bur. 11027 Merritt, March, 1908 (type), the specimen from
the southwestern part of the island; also For. Bur. 11016 Merritt from Mount
Sablayan, exposed summit of the mountain at edge of the forest, altitude about
970 m, March 2, 1908.

A species characterized by its comparatively large heads which are glomerate
on the ultimate panicle-branches. Among the Philippine species it is probably
most closely allied to Blumea mindanaensis Merr.

Blumea longipes sp. nov.

Herba erecta ramosa, usque ad 80 cm alta; foliis oblongo-obovatis vel
oblongo-oblanceolatis, submembranaceis, usque ad 10 cm longis, petiolatis,
acutis vel rotundatis, basi sensim angustatis, plus minus repando-dentatis,
vix lobatis; paniculis diffusis, ramis ramulisque glandulosis, pedunculis
solitariis, gracilibus, usque ad 2.5 cm longis; capitulis 0.8 cm longis.

An erect, much branched, aromatic herb about 80 cm in height. Stems
stout, brownish or purplish, slightly striate, more or less puberulent,
ultimately nearly glabrous, 5 mm in diameter or less. Leaves oblong-
obovate to oblong-oblanceolate, submembranaceous, the apex rounded or
acute, the base gradually narrowed, the margins somewhat repand-dentate
but not lobed, those of the stem 7 to 10 cm long, 1.5 to 3 cm wide, those
of the branches often much smaller and usually more pubescent. Pani-
cles rather diffuse, the branches and branchlets slender, mostly densely
glandular and often also somewhat pubescent, the peduncles solitary, 0.5
to 2.5 em long, slender, glandular. Heads 7 to 8 mm long, about 1 cm
in diameter. Involucral bracts many-seriate, the outer ones about 2 mm
long, 0.5 mm wide, the inner gradually longer, the innermost 5 mm long
and 0.5 mm wide, the outer ones usually prominently glandular, the
innermost ones ciliate above and slightly glandular in the median por-
tion. Flowers yellow, heterogamous, the outer ones pistillate, indefi-
nite, many-seriate; achenes minute, 1 mm long, shghtly angled, very
obscurely pubescent with few, short, scattered hairs; pappus hairs slender,
white, minutely scabrid, 3 mm long; corolla slender, cylindric; 4 mm
long, very obscurely 3-toothed ; style exserted, the arms filiform, less than
1 mm long. Disk-flowers perfect, about 14; achene like that of the
pistillate flowers but stouter ; corolla enlarged above, 5-toothed, the teeth
0.5 mm long, acute, slightly glandular; stamens 2 mm long, the tails
minute, less than 0.5 mm long; style-arms about 0.8 mm long, slender.
Disk glabrous, pitted.

Mrnpanao, Lake Lanao, Camp Keithley, Mrs. Clemens 893 (type), January,
1907; also unnumbered specimens collected in May and June.

A species growing in the open grass lands about Lake Lanao, characterized
to] Ss 5)
by its very glandular inflorescence and by its long-peduncled heads.

256 MERRILL.

Blumea mindanaensis sp. nov.

Herba erecta vix vel parce ramosa, stricta, usque ad 1 m alta; foliis
sessilibus, oblongis vel oblongo-obovatis, subcoriaceis, scabridis, plus minus
pubescentibus, margine irregulariter denticulatis ; capitulis in ramuis plus
minus congestis, circiter 1.7 em diametro.

An erect stout herb about 1 m high, slightly or not at all branched,
except the inflorescence. Stems stout, brown or purplish, striate, more
or less pubescent, 5 to 7 mm in diameter. Leaves sessile, oblong to
oblong-obovate, 6 to 12 em long, 1.5 to 3.5 em wide, subcoriaceous, some-
what shining when dry, scabrid, the upper surface often supplied with
numerous, small, white dots, beneath somewhat pubescent, the apex acute
or slightly acuminate, the base acute, the margins distantly and irreg-
ularly denticulate, not at all lobed or sinuate. Panicles about 40 cm
long, the lower branches sometimes 15 cm in length, often much smaller,
all parts rather densely brown-pubescent. Heads somewhat crowded,
shortly peduncled, 10 to 12 mm long, 15 to 18 mm in diameter, the
involucral bracts often purplish. Bracts several-seriate, the outer ones
ovate-lanceolate, acuminate, about 2 mm long, rather strongly pubescent,
the inner gradually larger, the innermost 8 mm long, 1 mm wide, ciliate
on the margins toward the apex. Flowers yellow, heterogamous, the outer
ones indefinite (more than 100), pistillate; achenes slender, 1 mm long,
glabrous or with very few scattered hairs; pappus-hairs about 20, white,
slender, minutely scabrid, 6 mm long; corolla cylindric, slender, 7 mm
long, obscurely 3-toothed ; style-arms exserted, slender, 1 mm long. Disk-
flowers perfect, about 11; achenes like those of the pistillate flowers but
stouter; corolla 6 mm long, enlarged above, 5-toothed, the teeth oblong-
ovate, acute, 0.6 mm long; stamens 2 mm long, minutely tailed at the
hase; style slightly exserted, the arms 0.5 mm long, slender. Disk glab-
rous or with very few, short, scattered hairs, pitted.

Minpanao, Lake Lanao, Camp Keithley, Mrs. Clemens 736 (type), November,
1906, and also unnumbered specimens collected in March, 1907, and in September,
1907.

A species well characterized by its comparatively large heads which are densely
disposed, its erect, strict habit, and sessile, scabrid leaves. It is probably allied

to Blumea chinensis Less., and among the Philippine species most closely allied
to B. incisa (Elm.) Merr., differing from the latter notably in its leaf characters.

Blumea ramosii sp. nov. :

Species B. sericanti Hook. f. affinis, sed differt foliis multo majoribus,
usque ad 16 cm longis, late oblongo-oblanceolatis, capitulis pedunculatis.

An erect, unbranched, suffrutescent plant at least 80 cm high, the
stem stout, terete, pubescent, mostly covered with the persistent, densely
arranged, petiole-bases, this part of the stem about 20 cm long, the leaves
crowded above this naked portion, subtending the inflorescence. Leaves
subcoriaceous, sessile, crowded, broadly oblong-oblanceolate, 11 to 16 em

NEW OR NOTEWORTHY PHILIPPINE PLANTS, VIII. 257

long, 3.5 to 5 cm wide, the apex acute, the base gradually narrowed,
slightly clasping, the margins rather finely denticulate, the upper surface
with numerous, somewhat stiff hairs more or less thickened at the base,
the lower surface very densely covered with long, grayish, silky hairs;
nerves about 12 on each side of the midrib. Panicles terminal, about 40
em long, the lower branches 10 em long or more, often subtended by
reduced leaves, the rachis and branches pilose. Heads somewhat racem-
osely arranged, 9 to 10 mm long, about 12 mm in diameter, their peduncles
3 to 10 mm long. Involucral bracts many-seriate, the outer ones ovate to
oblong, acuminate, 1 to 2 mm long, the inner gradually longer, the
innermost ones linear-oblong, 6 to 7 mm long, 1 mm wide, the interme-
diate ones somewhat wider, all somewhat pubescent on the back, more
or less 1-nerved, the margins, especially above, more or less ciliate.
Receptacle glabrous, somewhat pitted. Flowers heterogamous, the outer
ones pistillate, many-seriate, about 40 in each head, yellow, the achenes
slender oblong, pubescent, obscurely angled, 1 to 1.2 mm long; pappus-
hairs about 30, white, slender, scabrid, 4.5 mm long; corolla-tube slender,
cylindric, 4 mm long, obscurely 3-toothed; style-arms exserted, slender,
1 to 1.5 mm long.. Disk-flowers perfect, about 11 in each head; achenes
like those of the pistillate flowers but stouter; corolla 5.5 mm _ long,
enlarged above, 5-toothed, the teeth ovate, acute, nearly 1 mm long;
anthers 2 mm long, apex appendiculate, base with two minute, slender,
0.5 mm long tails; style exserted, the arms 0.5 mm long, stout, truncate.

Luzon, Province of Zambales, Mount Tapulao, Bur. Sci. 5089 Ramos, Decem-
ber 14, 1907.

A species apparently closely allied to Blwmea sericans Hook. f., and to
B. hieracifolia DC., but so far as I can determine from the material and descrip-
tions available here, distinct from both. It is characterized by its minutely
toothed and very densely silky leaves, suffrutescent stems, the lower portion
leafless and more or less covered with persistent petiole-bases. In many respects
it appears to agree with Inula § Cappa as closely as with Blumea, and may
eventually have to be referred to the former genus. The characters by which
Inula § Cappa is separated from Blumea do not appear to me to be well defined.

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THE PHILIPPINE

JOURNAL OF SCIENCE

C. BOTANY

VoL. VII NOVEMBER, 1912 No. 5
NEW OR NOTEWORTHY PHILIPPINE PLANTS, IX.

By E. D. MERRILL.

(From the Botanical Section of the Biological Laboratory, Bureau of Science,
and the Department of Botany, University of the Philippines,
Manila, P. I.)

The last paper published under this title appeared in the
year 1910, and the present contribution, like the preceding
numbers of the series, is composed of descriptions of presumably
new species, records of species previously described which have
recently been discovered in the Philippines, notes on nomencla-
ture, Synonymy, etc. Two new genera are proposed, Freeria,
of the Icacinaceae, and Macgregorianthus, of the Thymelaeaceae.
Four genera, previously not reported from the Archipelago, are
included, Parishia, Suriana, Cansjera, and Vallaris. One hun-
dred presumably new species are described, all dicotyledons,
while about 10 species, previously described from extra-Phil-
ippine material, are recorded from the Archipelago for the first
time. As is the case with the preceding numbers of the series,
all the material on which the present paper is based is preserved
in the Herbarium of the Bureau of Science.

CHLORANTHACEAE.

CHLORANTHUS L.

CHLORANTHUS PHILIPPINENSIS sp. nov.
Species C. henryi Hemsl. ut videtur valde affinis, differt flor-
ibus multo minoribus, circiter 2.5 mm longis.

An erect, simple, glabrous herb about 40 cm high, from aro-

* This Journal, 5 (1910) Botany 167-257.
113111 259

260 MERRILL.

matic, woody rootstocks, the stems brown, angled and sulcate
when dry, with about 5 nodes. Leaves 4, whorled at the apex
of the stem, membranaceous, broadly obovate, 12 to 15 em long, 8
to 9 cm wide, slightly shining, the lower surface paler than the
upper one, the apex prominently and sharply acuminate, the base
acute, margins sharply and rather finely serrate, the teeth more
distant below, the basal margins quite entire; nerves about 9 on
each side of the midrib; petioles about 1 em long. Inflorescence
terminal, about 15 cm long, slender, the peduncle about 7 cm long,
the branches opposite, usually 4 in two pairs, the lower ones about
6 cm long, the internode 2 cm long or less. Anther-scale about
2.5 mm long, divided nearly to the base into three lobes, the
lateral lobes a little shorter than the middle one, oblong, obtuse,
the middle one with a 4-locellate anther, the laterial ones with
2-locellate anthers; anthers about 1 mm long, half as long as the
lobes or less. Fruit unknown.

LUZON, Province of Cagayan, Abulug River, Weber 1582, January, 1912,
growing in forests, altitude about 250 m, the roots said to be fragrant, and
the fruits white, although none of the latter were preserved.

A species manifestly closely allied to Chloranthus henryi Hemsl., of
China, apparently differing chiefly in its smaller flowers, which are about
one-half as large as in Hemsley’s species. In habit, general appearance,
vegetative characters, etc., the Philippine plant very closely matches a
specimen in the Herbarium of the Bureau of Science, representing
Chloranthus henryi Hemsl., collected by Farges in Su-tchuen; ,the only dif-
ference appears to be in the flowers. I had at first determined the Luzon
plant to be the same as Chloranthus oldhami Solms-Laub., a species known
only from Formosa, and of which the flowers are unknown. That species,
however, is described as having its leaves subsessile, which does not apply
to our plant, nor are the leaves of our species crenate-dentate throughout,
the lower 2 to 4 em of the margins being quite entire.

ULMACEAE.
TREMA Lour.

TREMA VULCANICA sp. nov.

Frutex circiter 2 m altus subtus foliis ramulisque villosis;
foliis numerosis, confertis, oblongo-ovatis vel oblongis, usque ad
5 cm longis, acutis vel acuminatis, basi leviter cordatis, supra
scabridis, petiolo circiter 2 mm longo; cymis brevissimis, vix
7 mm longis, floribus 5-meris.

A shrub about 2 m high, the branchlets, petioles, and the lower
surface of the leaves, especially on the nerves, rather promi-
nently villous with rather short, pale, spreading or somewhat
appressed hairs, the older branches terete, glabrous or nearly

PHILIPPINE PLANTS, IX. 261

so, reddish-brown; branchlets somewhat crowded, leafless in
their lower parts, the leaves rather crowded above the middle.
Leaves subcoriaceous, brittle when dry, brownish, oblong-ovate
or oblong, 2.5 to 5 em long, 1 to 2 cm wide, acute or somewhat
acuminate, base somewhat cordate, equilateral or subequilateral,
the upper surface scabrid, usually with few hairs along the
midrib, the lower surface of about the same color as the upper,
prominently pubescent with pale hairs on the midrib, lateral
nerves, and primary reticulations, the ultimate reticulations close,
evident on both surfaces, margins closely and finely coriaceous-
serrulate; lateral nerves about 5 on each side of the midrib, prom-
inent, ascending, impressed on the upper surface; petioles
densely pale-villous, about 2 mm long. Cymes axillary, few-
flowered, pubescent, 4 to 7mm long. Flowers 5-merous. Male
flowers: Sepals 5, elliptic to elliptic-ovate, concave, about 1.8 mm
long, imbricate, glabrous except the slightly ciliate margins.
Stamens 5. Pistillode cylindric, truncate, 1 mm long, glabrous,
except the distinctly ciliate base. Female flowers similar to the
males, the style-arms 1 mm long. Fruit reddish, 2.5 mm long,
ovoid, fleshy, glabrous, more or less wrinkled when dry, the seed
subglobose, about 1.2 mm in diameter.

CAMIGUIN DE MINDANAO, in thickets on slopes of the old volcano, Bur. Sci.
14600 Ramos, April 27, 1912.

A specieg well characterized by its depauperate size, small crowded
leaves, short petioles and cymes, and other characters. It resembles

somewhat Trema amboinensis Blume, and is undoubtedly allied to that
species, but is quite distinct in many essential characters.

LORANTHACEAE.
LORANTHUS L.

LORANTHUS CONFUSUS sp. nov. § Cichlanthus.

Frutex parasiticus, ramis elongatis, tenuibus, ramulis subtus
foliis inflorescentiisque indumento stellato-leproso pallido dense
obtecto; foliis oppositis, breviter petiolatis, oblongis ad anguste
elliptico-oblongis, rotundatis vel acutis, usque ad 5.5 cm longis;
racemis brevibus, paucifloris; floribus 4-meris, corolla 12 mm
longa.

A slender parasitic shrub, the branches elongated, the branch-
lets terete, about 1 mm thick, these, the lower surfaces of the
leaves and the inflorescence densely covered with a pale, stellate-
leprose indumentum, the tips of the branchlets sometimes fer-
rugineous. Leaves opposite, coriaceous, oblong to narrowly
elliptic-oblong, 2 to 5.5 cm long, 8 to 20 mm wide, apex rounded

262 MERRILL.

or acute, base narrowed, acute, the upper surface, when young,
stellate-leprose, soon becoming quite glabrous; nerves 3 or 4 on
each side of the midrib, slender, not prominent; petioles 2 mm
long. Racemes axillary, short, solitary or fascicled, 2- to 5- -
flowered, the rachis about as long as the petiole. Flowers 4-
merous, somewhat curved. Pedicels about 3 mm long, the brac-
teole ovate, concave, 0.5 mm long. Calyx 3.5 to 4 mm long,
narrowed below into a pseudo-stalk, narrowly obovoid, truncate.
Corolla 12 mm long, tube slightly inflated. Reflexed parts of
the lobes above the insertion of the stamens 4 mm long, narrowly
oblong to linear-oblong. Anther sessile, erect, 1 mmlong. Fruit
(immature) narrowly obovoid, 5 mm long, shortly stipitate
densely stellate-leprose with a pale indumentum.

LUZON, without definite locality, Cuming 1959 (type): Province of Union,
Elmer 5711, February, 1904, Bur. Sci. 12951 Fénix, December, 1910:
Province of Pangasinan, Bur. Sci. 4977 Ramos, December, 1907, Alberto 35,
May, 1904: Province of Tarlac, Merrill s. n., July, 1903.

This form was previously considered by me’ to be referable to Loranthus
sphenoideus Blume, but I am now of the opinion that it is specifically dis-
tinct. It differs from the Philippine material that has been referred to
Loranthus sphenoideus in its pale, not ferruginous indumentum, its rela-
tively much narrower, differently shaped leaves, and its somewhat smaller
flowers. It is manifestly very closely allied to Blume’s species, but less
closely allied to Loranthus estipitatus Stapf.

LORANTHUS SIMILIS sp. nov. § Cichlanthus. ao

Species praecedente affinis, differt foliis majoribus, petiolo
longioribus, indumento ferrugineo, floribusque longioribus, 1.8
cm longis.

A slender parasitic shrub, the branches terete, brown, lenti-
cellate, the branchlets densely brown-stellate-leprose, as are the
petioles, lower surfaces of the leaves, and the inflorescence.
Leaves elliptic to oblong-elliptic, coriaceous, 5 to 7 cm long, 2 to
3 cm wide, rounded, base somewhat narrowed, rounded or sub-
acute, brown when dry, the upper surface glabrous, somewhat
shining, the lower surface densely brown-stellate-leprose; nerves
5 or 6 on each side of the midrib, rather distinct beneath, anas-
tomosing; petioles 8 to 10 mm long. Racemes axillary, solitary
or several at each node, few-flowered, the rachis about as long as
the petioles, the pedicels 2 to 3 mm long, the bracteole concave,
ovate, 1.3 mm long, all parts, including the outside of the flowers,
densely stellate-leprose. Calyx subcylindric, 3 mm long, slightly
or not narrowed toward the base. Corolla 18 mm long, slender,

? This Journal 4 (1909) Bot. 139.

PHILIPPINE PLANTS, IX. 263

slightly curved, very slightly inflated in the lower one-half, the
lobes 4.

Luzon, Benguet Subprovince, Baguio, Williams 984, July, 1904.

A species allied to Loranthus confusus Merr., and to L. sphenoideus,
differing from both in its longer flowers, and from the former also in its
ferruginous indumentum, larger leaves, and longer petioles. Like the
preceding species, the type was previously referred by me to L. sphenoideus
Blume.

LORANTHUS SUBSESSILIS sp. nov. § Dendrophthoé.

Glaber, ramulis teretibus; foliis oppositis, oblongo-ovatis,
acuminatis, petiolatis, coriaceis, usque ad 15 cm longis; floribus
sessilibus vel subsessilibus, e axillis defoliatis, solitariis vel binis,
6-meris, magnis, miniatis, circiter 7 cm longis, corolla falcata.

A glabrous parasitic shrub, the branches up to 1 m in length,
the branchlets terete, slender, grayish or reddish-brown, the
ultimate ones 1 to 1.5 mm in diameter. Leaves opposite, the
internodes 3 to 9 cm long, coriaceous, oblong-lanceolate, 8 to 15
cm long, 3 to 6 cm wide, acuminate, base narrowed, acute, rarely
somewhat rounded, rather pale and somewhat shining when dry;
nerves 6 to 8 on each side of the midrib, obscure, slender, often
nearly or quite obsolete; petioles about 5 mm long. Flowers in
the axils of fallen leaves, large, red, sessile or subsessile, solitary
or in pairs, each subtended by two bracteoles, the outer one
broadly ovate, slightly apiculate, about 5 mm long, the inner one
similar but truncate. Calyx about 7 mm long, 4 mm thick,
slightly enlarged upward, truncate. Corolla 6.5 to 7 cm long,
distinctly curved, the tube 4 to 4.5 em long, about 3 mm in di-
ameter at the base, gradually enlarged upward and 7 mm in
diameter at the throat, the lobes 6, 4 mm wide at the base, soon
narrowed, the reflexed part above the insertion of the stamens
2 cm long, lanceolate, acuminate, about 3 mm wide, much thick-
ened. Filaments 10 to 11 mm long, erect; anthers linear-lan-
ceolate, continuous, about 6 mm long.

MINDANAO, District of Zamboanga, Sax River, Merrill 8315, (type),
November 28, 1911, altitude about 900 m; Williams 2425, March 5, 1905,
altitude about 600 m.

A species characterized by its large flowers which are sessile or subsessile
in the axils of fallen leaves, solitary or in pairs. When in full flower the

plant is a very showy and striking one in the forests where it occurs. Its
flowers are larger than in any other known Philippine species.

LORANTHUS PUBIFLORUS sp. nov. § Dendrophthoé.
Frutex inflorescentiis exceptis glaber, ramis ramulisque tere-

tibus, crassis; foliis crasse coriaceis, oblongis vel anguste oblongo-
ovatis, usque ad 15 cm longis, verticillatis, petiolatis, acutis vel

264 MERRILL.

acuminatis; floribus in triadibus fasciculatis in radicibus vel in
ramulis specialibus, inflorescentiis brevibus, puberulis; floribus
4-meris, circiter 3 cm longis, tenuibus, extus puberulis.

A parasitic shrub glabrous except the inflorescence, the
branches and branchlets smooth, grayish or brownish, terete,
stout, smooth, the apparently specialized flower-bearing branches
(roots?) with a brown scaly bark. Leaves in whorls of three
or four, the internodes 10 to 12 cm long, petioled, usually brown
when dry, scarcely shining, thickly coriaceous, brittle, oblong to
narrowly oblong-ovate, acute or somewhat acuminate, 12 to 15
cm long, 4 to 8 cm wide; petioles stout, 1 to 1.5 em long; lateral
nerves 4 to 6 on each side of the midrib, not prominent, the
reticulations obsolete. Flowers in fascicled triads, the fascicles
scattered, brown-puberulent, the inflorescences, excluding the
corollas, less than 1 cm long. Bracteoles oblong-ovate, obtuse,
puberulent, about 1.5 mm long. Calyx urceolate-campanulate,
brown-puberulent, about 3 mm long, the limb somewhat spread-
ing, about 1 mm long, truncate. Corolla slender, about 28 mm
long, outside slightly puberulent, the lobes united for the lower
2 to 3 mm into a short tube, the free parts slender, about 1.5
mm wide, the reflexed part above the insertion of the stamen
narrowly oblong, about 5 mm long, 1.3 mm wide, acute or obtuse.
Anthers erect, continuous, narrowly oblong, about 3 mm long,
the filaments 2 mm long.

LUZON, Province of Laguna, near Paete, For. Bur. 13086 Curran, De-
cember 17, 1911, in forests.

A very characteristic species, distinguishable by its whorled, thickly
coriaceous, petioled leaves, and its flowers puberulent externally and borne
in fascicled short triads on the roots or on special leafless branches. In
the scattered character of its inflorescences it resembles Loranthus mira-
bilis Muell.-Arg. & Van Huerck, but is not, however, at all allied to that
species.

LORANTHUS CAPITULIFERUS sp. nov. § Lepiostegeres.

Ramulis brunneo-furfuraceis exceptis glaber; foliis oppositis
vel suboppositis, oblongo-ellipticis ad oblongo-ovatis, coriaceis,
utrinque angustatis, petiolatis, usque ad 12 cm longis; capitulis
axillaribus, solitariis, sessilibus, paucifloris; floribus circiter 1.8
cm longis, 6-meris.

A glabrous parasitic shrub, the branches and branchlets stout,
brown, the younger parts furfuraceous, more or less rugose
when dry. Leaves opposite or subopposite, coriaceous, oblong-
elliptic to oblong-ovate, olivaceous and slightly shining when
dry, 7 to 12 cm long, 2.5 to 5 em wide, obtuse, base narrowed,

PHILIPPINE PLANTS, IX. 265

acute or cuneate; lateral nerves 4 or 5 on each side of the mid-
rib, indistinct, slender; petioles about 1 cm long. Heads axil-
lary, solitary, sessile, but the flowers quite enclosed by imbricated
bracts, forming a globose head. Flowers 6-merous, about 5
in each head, sessile. Calyx 2 mm long, truncate. Corolla 1.6
em long, the lobes free nearly to the base, the reflexed part
above the insertion of the stamens 4 mm long. Anther erect,
sessile, 4 mm long.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8270, November 28, 1911, parasitic on Englehardtia, altitude

about 1100 m.
Allied to Loranthus williamsii Merr., but with much shorter flowers.

OLACACEAE.
CANSJERA Juss.

CANSJERA RHEEDII Gmel. Syst. Nat. 1 (1791) 220; Mast. in Hook. f.
Fl. Brit. Ind. 1 (1875) 582.
Cansjera scandens Roxb. Pl. Coromandel 1 (1795) 582.
Cansjera malabarica Lam. Encyel. 3 (1791) 438.

Sisutu IsLAND, Sulu Archipelago, Merrill 5291, October 13, 1906, in
thickets along the seashore.

India to southern China, through Malaya to northern Australia; the
genus new to the Philippines.

Gmelin’s specific name is here retained as being the one in common use.
Lamarck’s Cansjera malabarica was, however, published during the same
year, and it will be difficult if not impossible to determine which has
priority.

The species has previously been reported from the Philippines by
Meisner,’ but solely on Blanco’s error in interpreting the species. The
species Blanco referred here is an Antidesma.

MENISPERMACEAE.
STEPHANIA Lour.

STEPHANIA RAMOSII Diels sp. nov.

Caulis scandens glaber. Foliorum petiolus 4.5 ad 5 cm longus,
lamina peltata, papyracea, supra fere lucida, subtus paulo pal-
lidior, triangulari-ovata, apice acuta, acuminato-mucronulata, 8
ad 9 cm longa, 4 ad 5.5 cm lata. Inflorescentiae ¢ pseudo-
racemosae, 5 ad 7 cm longae; pedicelli 1.5 ad 2 mm longi. Sep-
ala 6, 3 exteriora anguste spathulato-oblonga, 1.5 mm longa,
0.4 mm lata, 3 interiora latiora, elliptico-ovata, concava, cir-
citer 1.2 mm longa, 0.8 mm lata. Petala 5 conchiformia vel

* DC. Prodr. 14 (1857) 519.

266 MERRILL.

fere orbicularia basi intus glandulosa, 0.7 ad 1 mm longa et
lata. Synandrium 1 mm diametro.

Luzon, Benguet Subprovince, near Baguio, Bur. Sci. 18487 Ramos,
May, 1911.

Sect. Thamnothyrsa Diels. Species nova foliis triangulari-ovatis S. cato-
sepalae Diels eiusdem provinciae indigenae haud absimilis sepalis petalisque
duplo minoribus haud deflexis conspicue differt.

PYCNARRHENA Miers.

PYCNARRHENA CELEBICA (Boerl.) Diels in Engl. Pflanzenreich 46
(1910) 53.
Cocculus celebicus Boerl. Cat. Pl. Bogor. (1899) 40.

LuzoN, Province of Nueva Ecija, For. Bur. 22157 Alvarez, December,
1910. MINDANAO, District of Davao, Mount Apo, Elmer 12000, September,
1909.

Previously known only from Celebes, and from plants cultivated in the
Botanical Garden at Buitenzorg; the Philippine specimens agree closely
with specimens in our herbarium from Buitenzorg, and have, morever, been
determined by Doctor Diels.

LIMACIA Lour.

LIMACIA BLUMEI (Boerl.) Diels in Engl. Pflanzenreich 46 (1910) 215.
Cocculus blumei Boerl. Cat. Pl. Bogor. (1899) 40.

MINDANAO, District of Davao, Mati, C. V. Piper 427, May 15, 1911.

The specimen closely matches typical material in our herbarium taken
from cultivated plants in the Botanical Garden at Buitenzorg, and has,
moreover, be2n determined by Doctor Diels. Previously known only from
specimens cultivated at Buitenzorg which originated in some part of the
Malay Archipelago; new to the Philippines.

ANONACEAE.
MITREPHORA Hook. f. & Thoms.

MITREPHORA WEBERI sp. nov.

Arbor circiter 8 m alta partibus junioribus inflorescentiisque
exceptis glabra; foliis ovatis vel oblongo-ovatis, subcoriaceis,
nitidis, usque ad 18 cm longis, acuminatis, basi late rotundatis,
nervis utrinque 8 ad 10; floribus hermaphroditis, circiter 4 cm
diametro, petalis interioribus oblongis, acutis, extus pubescen-
tibus, intus glabris, interioribus brevioribus, arcuatis, basi longe
angustatis, glabris, laminibus densissime ciliato-villosis; ovario
glabro, ovulis circiter 12.

A tree about 8 m high, glabrous except for the younger parts
and the inflorescence. Branches slender, terete, grayish- or red-
dish-brown, lenticellate, glabrous, the buds and very young leaves
densely ferruginous-pubescent. Leaves ovate to oblong-ovate,

PHILIPPINE PLANTS, IX. 267

subcoriaceous, shining, green and of about the same color on
both surfaces, 10 to 18 cm long, 4 to 8 cm wide; lateral nerves 8
to 10 on each side of the midrib, prominent, curved upward,
anastomising, the reticulations lax; petioles 7 to 10 mm long.
Flowers perfect, yellow and purple, extra-axillary or leaf-opposed,
solitary, or at least but one flower opening at a time, the short
rachis, the bracts, and the calyx externally densely ferruginous-
pubescent. Sepals broadly ovate, acute or slightly acuminate,
about 6 mm long. Outer three petals oblong, acute or somewhat
obtuse, about 2 cm long, 8 mm wide, externally somewhat ap-
pressed-pubescent with brown hairs, inside glabrous, the inner
three petals about 14 mm long, vaulted, conniving, their claws
slender, glabrous, about 9 mm long, their limbs about 5 mm long,
subrhombic, but when spread much wider than long, the inner
surface very densely ciliate-villous with long, dark-brown hairs.
Stamens indefinite, narrowly oblong, 1 to 1.2 mm long, their
connectives truncate, overlapping. Ovaries 12 to 15, glabrous,
1 to 1.2 mm long, oblong; ovules about 12, 2-seriate; styles
clavate-oblong. Fruits unknown.

BUSUANGA (Calamianes Islands), C. M. Weber 1550bis, February 25,
1911, in forests, limestone region, altitude about 30 meters.

Apparently a very characteristic species, distinguishable by the inner
faces of the limbs of the interior petals being very densely ciliate-villous
with long, dark-brown hairs, and by its outer petals being quite glabrous
inside.

OXYMITRA Hook. f. & Thoms.
OXYMITRA PUBESCENS sp. nov.

Frutex scandens omnibus partibus plus minusve ferrugineo-
pilosis, ramulis densissime pilosis; foliis oblongis vel anguste
oblongo-obovatis, abrupte brevissime acuminatis, basi leviter
cordatis, subcoriaceis vel chartaceis, usque ad 24 cm longis, nervis
utrinque circiter 10, distantibus, distinctis; petiolo crasso, vix
6 mm longo; floribus ignotis; fructibus ovoideis vel ellipsoideis,
circiter 8 cm longis, aurantiacis, ferrugineo-pilosis.

A scandent shrub, all parts more or less ferruginous-pilose,
the branchlets very densely so, the older branches bearly black,
lenticellate, becoming glabrous. Leaves subcoriaceous or firmly
chartaceous, oblong or narrowly oblong-ovate, 14 to 24 cm long,
5.5 to 8.5 cm wide, usually widest above the middle, the apex
shortly and abruptly acuminate, below usually more or less nar-
rowed, the base rounded, somewhat cordate, both surfaces more
or less pilose with ferrugenous hairs, the upper surface in age
becoming more or less glabrous, the lower surface slightly paler,

268 MERRILL.

the nerves and reticulations especially pilose; lateral nerves 10
on each side of the midrib, prominent, somewhat ascending,
anastomosing, the reticulations subparallel, distinct; petioles
densely ferruginous-pilose, thickened, 6 mm long, or less. Flow-
ers axillary, sessile or very shortly pedicelled, apparently solitary
(not seen). Fruits umbellately arranged on stout, axillary,
pubescent tubercles, 6 or more in each umbel, their pedicels
pubescent, 5 to 8 mm long, the fruits yellow when fresh, dark-
brown when dry, ovoid or ellipsoid, about 8 mm long, more
or less ferruginous-pilose, the apex minutely apiculate.

LuzON, Province of Cagayan, Abulug River, Bur. Sci. 138870 Ramos,
January 26, 1912, on trees in forests.

A species well characterized by its pilose, ferruginous indumentum and
comparatively few, distant, lateral nerves.

POLYALTHIA Blume.

POLYALTHIA LOHERI sp. nov.

Species P. lanceolatae Vid. similis et affinis, differt foliis ma-
joribus, tenuiter acuminatis, floribus majoribus, circiter 2 cm
longis.

A tree 5 to 10 m high, nearly glabrous, the axils bearded with
short, stiff, ferruginous hairs, the branchlets slender, terete,
dark-colored when dry, usually lenticellate. Leaves chartaceous
or subcoriaceous, glabrous, rather pale when dry, shining and
of the same color on both surfaces, lanceolate to oblong-lanceo-
late, 15 to 25 cm long, 2 to 5 cm wide, gradually narrowed into
a long and rather slender acumen, base rounded or narrowed,
minutely cordate, often very slightly inequilateral; primary
nerves about 19 on each side of the midrib, spreading, anas-
tomosing, the secondary ones often nearly as prominent, the
reticulations rather lax, evident; petioles 3 mm long or less.
Flowers axillary, solitary, perfect, white, cream-colored, or
yellow, the pedicels slender, somewhat thickened upward, 3
to 4 cm long. Sepals triangular-ovate, acute or slightly acu-
minate, 3 to 4 mm long. Petals 6, subequal, oblong to oblong-
obovate, thick, coriaceous, somewhat pubescent especially along
the margins toward their tips, obtuse or acute, somewhat united
below, 2 to 2.5 cm long, 4 to 7 mm wide. Anthers numerous,
the connectives truncate. Ovaries villous. Fruit ellipsoid,
apparently red, about 2 em long, glabrous or nearly so.

LUZON, Province of Cagayan, Cuming 1846; Culasing Mountain, For.

Bur. 18487 Alvarez (type), February 22, 1909, in dipterocarp forests;
Pamplona, For. Bur. 14780 Darling, March, 1909: Province of Rizal, San

PHILIPPINE PLANTS, IX. 2969

Mateo, For Bur. 1103 Ahern’s collector, May, 1904, in fruit; Montalban,
Loher 5531, January, 1906, with immature flowers: Province of Tayabas,
Mauban, For. Bur. 10191 Curran, March, 1908.

The alliance of this species is manifestly with Polyalthia lanceolata Vid.,
and some of the material cited above has been distributed under that name.
P. loheri, however, has much larger, differently shaped leaves, and much
larger flowers, and is abundantly distinct.

Var. CAGAYANENSIS var. nov.

A typo differt ramulis pallidis, foliis majoribus, usque, ad 35
cm longis, venis magis distinctis.

LUZON, Province of Cagayan, Abulug River, near Dabba, Bur. Sci. 13948
Ramos, February 2, 1912, in forests, a tree about 5 m high with yellow
flowers.

Additional material may show that this form is specifically distinct, or
may merge it with Polyalthia loheri.

POLYALTHIA TENUIPES sp. nov.

Ut videtur arbor parva, ramulis junioribus exceptis glabra;
foliis lanceolatis, sensim angustatis, acuminatis, basi angustatis,
obtusis, minute cordatis, usque ad 12 cm longis, nervis utrinque
10 ad 12; floribus in axillis superioribus, solitariis, tenuiter
pedicellatis, circiter 8 mm longis, pedicellis 2 ad 3 cm longis;
petalis exterioribus triangulari-ovatis, interioribus lanceolatis
quam exterioribus duplo longioribus.

Apparently a small tree, nearly glabrous, the growing shoots
densely appressed ferruginous-pubescent, the branches slender,
glabrous, dark-colored, terete. Leaves lanceolate, chartaceous,
9 to 12 cm long, 1.5 to 2.5 cm wide, gradually narrowed above
to the slenderly acuminate apex, the base somewhat narrowed,
obtuse, slightly cordate, of nearly the same color and shining
on both surfaces when dry; primary nerves 10 to 12 on each
side of the midrib, spreading, anastomosing, irregular, the
secondary ones often nearly as prominent as the primary ones,
the reticulations lax; petioles 2 mm long or less. Flowers soli-
tary, perfect, axillary, in the upper axils, about 8 mm long,
their pedicels very slender, 2 to 3 cm long, with a small oblong-
lanceolate bracteole near the base. Sepals slightly pubescent,
triangular-ovate, acute, about 2 mm long, united. Petals 6, in
two series, dissimilar, the outer ones triangular-ovate, acute or
somewhat acuminate, 5 mm long or less, the inner three lanceo-
late, somewhat acuminate, about 8 mm long.

Luzon, Province of Tayabas, Kabibihan, Bur. Sci. 13472 Foxworthy
& Ramos, March, 1911.

A species with vegetative characters quite similar to those of Polyalthia
lanceolata Vid., but sharply differentiated by its dissimilar petals.

270 MERRILL.

CAPPARIDACEAE.
CAPPARIS L.

CAPPARIS LOHERI sp. nov.

Frutex scandens, armatus, subtus foliis ramulisque pubescen-
tibus; foliis papyraceis, concoloribus, lanceolatis ad late lanceo-
latis, brevissime petiolatis, basi rotundatis, apice sensim angus-
tatis, acuminatis, nervis utrinque circiter 11, distinctis, anas-
tomosantibus; floribus longe pedicellatis, 4 ad 6 in umbellis
axillaribus.

A scandent shrub, the nodes armed with short, sharp, straight
spines 2 mm long or less, the branchlets terete, brownish,
pubescent. Leaves lanceolate to broadly lanceolate, rarely ovate-
lanceolate, chartaceous, 9 to 19 em long, 2 to 4.5 em wide,
green and of about the same color on both surfaces when dry,
slightly shining, the upper surface quite glabrous or the midrib
slightly pubescent, the lower surface softly pubescent at least on
the midrib and nerves, base rounded, gradually narrowed up-
ward from about the middle to the long and slender acumen,
sometimes slightly falcate; lateral nerves about 11 on each side of
the midrib, spreading, prominent, looped-anastomosing, the
recticulations lax, distinct; petioles pubescent, about 2 mm long.
Umbels axillary, solitary, the peduncles pubescent, slender, 1
to 1.5 cm long, 4- to 6-flowered, the pedicels 2 to 3 cm in length,
slender. Sepals 5 to 6 mm long, concave, orbicular-elliptic,
rounded. Petals hyaline, thin, oblong, rounded, about 3.5 mm
long, base truncate, margins ciliate. Stamens numerous, their
filaments about 2 cm long; anthers narrowly oblong, 2 mm long.
Gynophore slender, 3 cm in length in anthesis, elongated and
3.5 to 5 cm in length in fruit; ovary narrowly ovoid. Fruit
(immature) ellipsoid, fleshy, 1.8 cm long.

LUZON, Province of Rizal, Montalban, Loher 2016 August, 1890, in herb.
Kew., 6787 (type), November, 1905.

A species well characterized by its long and slenderly acuminate
pubescent leaves, its axillary, solitary, few-flowered umbels, its long-
pedicelled flowers, and greatly elongated gynophore. It is allied to Capparis

lobbiana Turcz., but is very different from that species in vegetative and
many other characters.

CAPPARIS LITTORALIS sp. nov.
Species C. lobbianae Turcz., similis et affinis, differt omnibus

partibus glabris, vix molliter pubescentibus.
An erect (or somewhat scandent?) shrub about 2 m high,

PHILIPPINE PLANTS, IX. yeti

the branchlets terete, slightly pubescent, the nodes with very
short sharp spines 1.5 mm long or less. Leaves ovate to oblong-
ovate, 1.5 to 5 cm long, 1 to 2.5 cm wide, coriaceous, shining,
glabrous, acute or very slightly apiculate-acuminate, base broad,
cordate; nerves about 8 on each side of the midrib, very slender,
not at all prominent; petioles 2 mm long or less. Flowers (not
seen) in axillary, solitary, apparently few-flowered umbels, the
peduncle in fruit 1.5 to 2 cm long, slender, glabrous, the pedicels
(gynophore) 1.5 to 2.5 cm in length. Fruit ovoid or subglobose,
about 8 mm in diameter, white when fresh, brown when dry,
apparently fleshy, 2- or 3- seeded, the seeds 3 to 4 mm in length.

LUZON, Province of Tayabas, For. Bur. 11111 Curran (type), April, 1908,
on sand dunes along the seashore: Province of Ilocos Norte, Bur. Sci. 7602
Ramos, March, 1909, in sandy places near the sea.

A species manifestly very closely allied to Capparis lobbiana Turcz.,
which it closely resembles, differing in having entirely glabrous, shining,
coriaceous or subcoriaceous, not softly pubescent leaves.

SAXIFRAGACEAE.
POLYOSMA Blume.

POLYOSMA PIPERI sp. nov.

Frutex vel arbor parva; foliis anguste oblongo-obovatis vel late
oblanceolatis, usque ad 16 cm longis, integris, apice acuminatis,
basi angustatis, acutis, oppositis, chartaceis, subtus ad costa ner-
visque plus minusve hirsutis, nervis utrinque circiter 12, promi-
nentibus; racemis terminalibus, pubescentibus, quam folia lon-
gioribus; floribus 4-meris, circiter 12 mm longis.

A shrub or a small tree, the branches terete, glabrous, the
younger ones nearly black when dry and more or less pubescent
with short, spreading, often fulvous hairs. Leaves opposite,
narrowly oblong-obovate to broadly oblong-oblanceolate, charta-
ceous, 12 to 16 em long, 3.5 to 5 cm wide, entire, the apex rather
prominently acuminate, gradually narrowed from about the
middle to the acute base, the upper surface glabrous or nearly
so, the lower of about the same color, more or less fulvous or
pale-pubescent along the midrib and lateral nerves; lateral
nerves about 12 on each side of the midrib, prominent, curved,
anastomosing, the reticulations rather lax; petioles 1.5 to 2.5
cm long, pubescent. Racemes terminal, more or less hirsute-
pubescent with short, pale or grayish hairs, a little longer than
the leaves, erect. Flowers numerous, about 12 mm long, their

272 MERRILL.

pedicels 2 to 2.5 mm long, the bracteoles lanceolate, acuminate,
pubescent, about 3 mm long. Calyx somewhat urceolate, about
2.5 mm long, slightly constricted at the apex, the 4 teeth spread-
ing, triangular-ovate, acuminate, short. Petals 4, linear, obtuse,
10 to 11 mm long, to 1.5 mm wide, longitudinally 5- or 6-nerved,
the nerves distinct, stout, dark-colored, outside more or less
appressed-pubescent with short hairs, inside ciliate-pilose with
white hairs. Filaments 6 mm long, ciliate-pilose; anther linear,
4mm long. Fruit (rather immature) ovoid-ellipsoid, glabrous,
somewhat longitudinally suleate when dry, about 12 mm long,
6 to 7mm wide, somewhat narrowed at both ends.

MINDANAO, Province of Surigao, Hinatuan, C. V. Piper 519, May 16, 1911.

A species distinguishable by its leaves being somewhat pubescent beneath,
its long racemes, and relatively short flowers.

POLYOSMA VERTICILLATA sp. nov.

Arbor parva 4 ad 5 m alta, partibus junioribus inflorescen-
tiisque parce pubescentibus; foliis oblongis, coriaceis, ternis,
verticillatis vel subverticillatis, nitidis, acuminatis, 4 ad 10 cm
longis, irregulariter glanduloso-dentatis; racemis axillaribus,
multifloris; floribus purpureis, vix 1 ecm longis.

A small tree 4 to 5 m high, nearly or quite glabrous except
the sparingly pubescent inflorescence and younger parts.
Leaves in whorls of threes, .coriaceous, when young somewhat
pubescent, when mature quite glabrous, shining, usually oblong,
4 to 10 cm long, 1.5 to 3.5 cm wide, rather coarsely and irregularly
glandular-dentate, sometimes subentire, acuminate, base acute;
nerves 15 to 20 on each side of the midrib, spreading at nearly
right angles, slender, anastomosing; petioles 1 to 2.5 cm long.
Racemes axillary, slightly pubescent, peduncled, 4 to 5 em long.
Flowers numerous, crowded in the upper one-half of the ra-
ceme, their pedicels 2 mm long, pubescent, with 3 oblong, acute,
1 mm long bracts at the apex subtending the calyx. Calyx
pubescent, somewhat urceolate, 2.5 mm long, the lobes 4, trian-
gular-ovate, acute, 1 mm long. Petals 4, narrowly oblong, 8
mm long, 1.5 mm wide, slightly appressed-pubescent with very
short, scattered hairs. Fruit subglobose to ovoid, black when
mature, about 8 mm long.

LUZON, Benguet Subprovince, Mount Tonglon (Santo Tomas), Phil. Pl.
772 Merrill, May, 1911 (type), Williams 1526, November, 1904, Bur. Sci.
5411 Ramos, December, 1908; Mount Ugo, Bur. Sci. 5593 Ramos; Mount

Pulog, For. Bur. 18148 Curran, Merritt, & Zschokke; Baguio, Elmer 8798.
A very characteristic species, readily recognizable by its leaves being

PHILIPPINE PLANTS, IX. 2le

in whorls of threes, not opposite as in most species of the genus; it differs
from Polyosma philippinensis also in its considerably shorter and less
pubescent flowers, as well as in its usually more prominently dentate leaves.

LEGUMINOSAE.
DESMODIUM Desy.

DESMODIUM TRIFOLIASTRUM Mig. Fl. Ind. Bat. 1* (1855) 248.

MINDANAO, Province of Surigao, Hinatuan, C. V. Piper 549, May 16, 1911.
This species has not been reported from the Philippines previously, and,
in fact, is definitely recorded only from Java. The species has been deter-
mined through the kindness of Sir D. Prain, director of the Royal Gardens,
Kew, to whom a specimen was sent for comparison with the material in the
Kew Herbarium.
DERRIS Lour.

DERRIS CEBUENSIS sp. nov. § Dipteroderris.

Frutex scandens, glaber; foliis 13 ad 20 cm longis, foliolis 7,
nitidis, oblongis vel oblongo-ellipticis, in siccitate pallidis, utrin-
que concoloribus, 4 ad 8 cm longis, breviter obtuse acuminatis;
floribus ignotis; leguminibus oblongis vel oblongo-lanceolatis,
6 ad 10 em longis, planis, 2.5 ad 3 em latis; seminibus 2 vel 38.

A scandent vine, glabrous throughout (flowers unknown), the
stems brownish-olivaceous, prominently lenticellate, terete, the
young branchlets smooth. Leaves odd-pinnate, 13 to 20 cm long,
the common petiole and rachis 7 to 11 em long. Leaflets 7,
coriaceous or subcoriaceous, pale, shining and of the same color
on both surfaces when dry, oblong to oblong-elliptic, 4 to 8 cm
long, 2.5 to 3.5 em wide, the base obtuse or rounded, apex
shortly and obscurely blunt-acuminate, or sometimes obtuse;
nerves slender, obscure, anastomosing, 7 to 10 on each side of
the midrib; petiolules 3 to 8 mm long. Flowers unknown. In-
frutescence axillary, paniculate, up to 25 cm long, entirely
glabrous, the lower branches up to 8 cm long, the upper ones
shorter. Persistent calyces shallowly cup-shaped, truncate, 5
mm in diameter. Pods thin, flat, oblong to oblong-lanceolate,
pale when dry, 6 to 10 cm long, 2.5 to 3 cm wide, the base
acute or acuminate, apex shortly and slightly falcately apiculate-
acuminate, the wings thin, 3 to 7 mm wide, of the same width
on both sides of the pod. Seeds 2 or 3, rarely only 1, thin,
flat, 2 mm long.

CEBU, Bucacao, Bur. Sci. 11014 Ramos, March 19, 1912, on dry hills.

A species apparently most closely allied to Derris mindorensis Perk.,

and to D. micans Perk., but distinguishable by being entirely glabrous
throughout.

274 MERRILL.

MEDICAGO L.

MEDICAGO LUPULINA L. Sp. Pl. (1753) 779; Baker in Hook. f. Fl. Brit.
Ind. 2 (1876) 90.

LUZON, Benguet Subprovince, Baguio, Merrill 7762, May, 1911, in waste
places, open grassy slopes, altitude about 1500 m. Introduced and appar-
ently established, as is Medicago denticulata Willd.

Widely distributed in Europe and Asia, introduced in North America.

SIMARUBACEAE.
SURIANA L.

SURIANA MARITIMA L. Sp. Pl. (1758) 284; A. Gray Bot. Wilkes U. S.
Explor. Exped. (1854) 307.

LUMBUCAN ISLAND, Sulu Sea, Phil. Pl. 403 Merrill, September, 1910.
MANGSEE ISLAND, Wilkes Expedition.

This species has already been reported from the Philippines by A. Gray,
specimens having been collected by the Wilkes Expedition on Mangsee
Island, a short distance south of Lumbucan this record having been
overlooked by me in my paper on the Philippine plants collected by the
Wilkes Expedition. It is widely distributed in the tropics of the World,
but its occurrence in the Old World is rather peculiar and apparently
restricted. I have found no published record of its occurrence on the
Asiatic continent, nor in Formosa. I have specimens collected on Pratas
Island, in the China Sea between Hongkong and Luzon, sent to me by
Mr. Kawakami of Taihoku, Formosa. Both Miquel and Boerlage record
it as Malayan, but without citing any locality or localities for it. It is
found in the Laccadive Islands, in German New Guinea, and on the
islands off the north-east coast of Australia, while it seems to be widely
distributed in southern Polynesia.

MELIACEAE.
AGLAIA Lour.

AGLAIA BRACHYBOTRYS sp. nov. § Euaglaia.

Arbor circiter 8 m alta, ramilis subtus foliis inflorescentiisque
minute subferrugineo-ciliato-lepidotis; foliis alternis, circiter 40
em longis, foliolis membranaceis, alternis, circiter 7, oblongo-
ovatis vel oblongo-ellipticis, usque ad 17 cm longis; paniculis
lateralibus, circiter 9 cm longis; floribus racemose dispositis,
brevissime pedicellatis, 4-meris, tubo stamineo cum petalis di-
stincte coalito.

A tree about 8 m high. Branches pale-brown, terete, densely
covered with minute ciliate-lepidote scales. Leaves alternate,
unequally pinnate, about 40 cm long, the petiole, rachis, and

‘This Journal 3 (1908) Bot. 73-84.

PHILIPPINE PLANTS, IX. 275

petiolules densely covered with pale-brown, lepidote, ciliate scales.
Leaflets alternate, usually 7, membranaceous, ovate-oblong to
oblong-elliptic, 12 to 17 cm long, 5.5 to 8 em wide, slightly
shining when dry, glabrous above, beneath of the same color
and ciliate-lepidote on the nerves and midrib, and with scattered,
small scales on the surface, the base of the lateral ones inequi-
lateral, somewhat acuminate, of the terminal one equilateral, the
apices somewhat acuminate; lateral nerves 8 to 12 on each side
of the midrib, prominent; petiolules of the lateral leaflets 1 cm
long, of the terminal one about twice as long. Panicles lateral,
about 9 cm long, the peduncle less than 1 cm long, somewhat
dichotomously branched, the branches spreading, all parts covered
with small, pale-brown, ciliate-lepidote scales. Flowers about 3
mm long, racemosely rather densely arranged on the ultimate
branchlets, their pedicel$ 0.5 mm long or less. Calyx ciliate-
lepidote, when spread nearly 2 mm square, the teeth 4, rarely
5, broadly triangular, acute, less than 0.5 mm long. Petals 4,
glabrous, elliptic-oblong or oblong-obovate, obtuse, about 3 mm
long, 2 mm wide, adherent to the staminal-tube in the lower
one-third. Staminal-tube 2.5 mm long, truncate, glabrous, bear-
ing 7 or 8 stamens attached at or just above the middle, the
anthers about 1 mm long, included. Ovary sessile, ovoid, minute,
pubescent. Fruit unknown.

LUZON, Province of Cagayan, San Vicente, in dense forests at an altitude
of about 15 m, For. Bur. 15497 Bernardo, April 15, 1910.

A species with somewhat the general appearance of Aglaia hexandra
Turcz., but with fewer leaflets, much shorter panicles, and quite different
flowers. According to DeCandolle’s scheme of classification it is much
more closely allied to Aglaia turczaninowii A. DC., but is entirely different
from that species.

AGLAIA CAGAYANENSIS sp. nov. § Euaglaia.

Arbor circiter 7 m alta, partibus junioribus inflorescentiisque
lepidotis, vetustioribus glabris; foliis usque ad 14 cm longis,
3- ad 5-foliolatis, foliolis anguste oblongo-ellipticis vel oblongo-
ovatis, caudato-acuminatis; paniculis axillaribus, quam folia
brevioribus, paucifloris; floribus racemose dispositis, longe pedi-
cellatis, tubo stamineo liber.

A tree about 7 m high, the branchlets and inflorescence rather
densely lepidote, the older parts entirely glabrous. Branches
slender, terete, dark-colored when dry, the branchlets pale.
Leaves 14 cm long or less, alternate, the rachis and petiole 3
to 5 cm long; leaflets subcoriaceous, brittle when dry, 3 to 5,

113111——2

276 MERRILL.

narrowly oblong-elliptic to narrowly oblong-ovate, 6 to 10 cm
long, 1.5 to 3 cm wide, narrowed at both ends, the apex somewhat
caudate-acuminate, base acute, both surfaces shining, the lower
a little paler than the upper; nerves 6 to 8 on each side of
the midrib, not prominent; petiolules 2 to 3 mm long. Panicles
in the upper axils, 10 cm long or less, peduncled, with few
branches, each branch with from 1 to 3 racemosely disposed
flowers, the pedicels 3 to 10 mm long. Calyx cup-shaped, about
2 mm in diameter, shallowly 5-toothed. Petals glabrous, orbi-
cular, concave, 2.5 to 3 mm in diameter. Staminal-tube globose,
free from the petals, about 2mm long. Anthers 5, 1.5 mm long,
included.

LuZzON, Province of Cagayan, Abulug River, Bur. Sci. 13801 Ramos,
January, 1912.

A species well characterized by its lax, very few-flowered panicles, the
flowers long-pedicelled.

AGLAIA CURRANII sp. nov. § Huaglaia.

Arbor circiter 8 m alta, ramulis petiolis inflorescentiisque
plus minusve ciliato-lepidotis ; foliis alternis, circiter 30 cm longis,
imparipinnatis, foliolis utrinque 5, lanceolatis, subcoriaceis, us-
que ad 11 cm longis, acuminatis, basi valde inaequilateralibus,
subtus ad costa plus minusve ciliato-lepidotis; paniculis axilla-
ribus, circiter 12 cm longis; floribus 5-meris, sessilibus vel sub-
sessilibus, spicatim dispositis; petalis liberis.

A tree about 8m high. Branches terete, glabrous, lenticellate,
brownish, the younger parts more or less covered with minute,
ciliate, lepidote scales, pale-brownish in color. Leaves alternate,
about 30 cm long, the petioles, rachis, and petiolules densely
covered with small, brownish, lepidote-ciliate scales. Leaflets
usually 11, lanceolate, subcoriaceous, rather pale and of about
the same color on both surfaces when dry, slightly shining above,
9 to 11 cm long, 2 to 3 cm wide, the apex prominently acuminate,
the acumen blunt, the base strongly inequilateral, acuminate;
nerves about 12 on each side of the midrib, slender, not prom-
inent; petiolules about 1 ecm long, that of the terminal leaflet
about 1.5 cm long. Panicles axillary, ciliate-lepidote, about 12
cm long, branched from just above the base, the primary branches
up to 7 cm in length. Flowers spicately disposed on the ultimate
branchlets, sessile or subsessile, not at all crowded. Calyx
densely ciliate-lepidote, the teeth 5, ovate, obtuse, ciliate, less
than 0.5 mm long. Petals 5, glabrous, free from the staminal
tube, oblong, obtuse, 1.5 to 1.8 mm long (just before anthesis).
Staminal tube very short, glabrous, 1 mm long or less, truncate.

PHILIPPINE PLANTS, IX. Dun

Stamens 5, inserted on the base of the tube, included, the anthers
0.5 mm long. Fruit unknown.

LUZON, Province of Bataan, Limay Peak, Mariveles Mountain, For. Bur.
17580 Curran, November, 1909, on dry forested ridges at an altitude of
about 700 m.

A species well characterized by its lanceolate leaflets, its small ciliate-
lepidote scales on the inflorescence, petioles, etc., its spicately arranged,
sessile or subsessile flowers, and very short staminal-tube. According to De-
Candolle’s scheme of classification it seems to be allied to Aglaia llanosiana
C. DC., but in all respects is quite different from that species.

AGLAIA DIFFUSA sp. nov. § Hearnia.

Arbor circiter 25 m alta, inflorescentiis exceptis glabra; foliis
alternis, imparipinnatis, 3-jugatis, vix 30 cm longis; foliolis
lanceolatis vel elliptico-lanceolatis, utrinque acuminatis aequi-
lateralibusque, usque ad 44 cm longis, nervis utrinque circiter
10; paniculis axillaribus, folia aequantibus, diffusis, multifloris,
ramis primariis patulis, usque ad 15 cm longis, plus minusve
lepidotis vel puberulis; floribus 5-meris, pedicellatis, racemose
dispositis.

A tree about 25 m high, glabrous except the somewhat lepidote
and puberulent inflorescence. Branches terete, brownish, gla-
brous, the ultimate ones 4 mm in diameter or less. Leaves
alternate, less than 30 cm long, the petiole and rachis 10 to 18
em long. Leaflets 7, the lateral ones opposite, lanceolate or
elliptic-lanceolate, 9 to 14 cm long, 2 to 4 cm wide, narrowed
and acuminate at both ends, rather pale and somewhat shining
when dry, of about the same color on both surfaces, glabrous,
or the midrib beneath with few, small scales; nerves about 10
on each side of the midrib, slender, not very prominent, the
reticulations lax, subobsolete; petiolules 4 to 6 mm long. Pani-
cles in the upper axils, usually but one on a branchlet, diffuse,
about 30 cm long, the lower branches spreading, 15 cm long,
very many flowered, the rachis somewhat brown-lepidote, the
ultimate branchlets somewhat brown-puberulent. Flowers yel-
low, racemosely arranged on the ultimate branchlets, scattered,
their pedicels 1 to 1.5 mm long. Calyx-teeth 5, broadly ovate,
obtuse, 0.3 mm long, ciliate. Petals 5, glabrous, oblong-obovate
to obovate, obtuse, 1.6 mm long, free from the staminal-tube.
Staminal-tube broadly obovoid, truncate, about 1 mm long and
1.2 mm in diameter. Stamens 5, inserted on the edge of the
staminal-tube, slightly inflexed, 0.25 mm long.

LuzON, Province of Tayabas, Guinayangan, For. Bur. 18684 Darling,
November 21, 1909. In forests at an altitude of about 50 m, locally known

as malasaguin.
A species of the section Hearnia, well characterized by being nearly

278 MERRILL.

glabrous throughout, and by its very diffuse, ample, many-flowered panicles
which are more or less lepidote and puberulent. It does not seem to me
to be closely allied to any of the previously described species of the genus,
although in many respects it resembles Aglaia harmsiana Perk.

DYSOXYLUM Blume.

DYSOXYLUM LAXUM sp. nov. § Eudysoxylum.

Arbor circiter 10 m alta, partibus junioribus puberulis ex-
ceptis glabra; foliis alternis, abrupte pinnatis, circiter 30 cm
longis, 4- vel 5-jugis, foliolis oblongo-ovatis ad late oblongo-
lanceolatis, valde acuminatis, in siccitate pallidis; paniculis axil-
laribus, folia aequantibus, parce ramosis, paucifloribus; floribus
4-meris, circiter 1 cm longis, petalis supra puberulis.

A tree about 10 m high, glabrous except the somewhat pu-
berulent younger parts. Branches terete, brownish, lenticellate,
the growing parts grayish and somewhat puberulent. Leaves
alternate, evenly pinnate, about 30 cm long, the rachis and petiole
slender, more or less puberulent, about 15 cm long; leaflets 4 or
5 pairs, the uppermost ones opposite, the others subopposite or
alternate, thinly chartaceous or membranaceous, pale when dry,
concolorous, slightly shining, oblong-ovate to broadly oblong-
lanceolate or somewhat elliptic-oblanceolate, the lower ones in-
equilateral at the base, 4 to 5 cm long, the upper ones equilateral,
gradually larger and up to 15 cm long and 4.5 cm wide, base
acute, apex rather slenderly acuminate; lateral nerves about
10 pairs, slender; petiolules 2 to 5 mm long. Panicles axillary,
slender, as long as the leaves, lax, each with very few spreading
branches, the lower branches 6 cm long or less and each bearing
2 or 3 flowers, the slender pedicels up to 1.5 cm long. Flowers
pinkish, 4-merous. Calyx narrowly funnel-shaped, narrowed
below, about 5 mm long, puberulent, obscurely toothed. Petals
narrowly oblong, obtuse, about 8 mm long, glabrous on the back
except at the apex where they are somewhat puberulent. Sta-
minal tube cylindric, free or nearly so, 6 mm long, somewhat
toothed at the apex. Stamens 8, inserted at the top of the
tube, included. Disk cylindric, 4-toothed, about 2.5 mm long,
glabrous. Ovary hirsute; style 5 mm long, slender, glabrous.
Young fruits obovoid or somewhat ellipsoid, 1.5 em long, gla-
brous, apiculate at the apex.

CAMIGUIN DE MINDANAO, Bur. Sci. 14694 Ramos, March 28, 1912, in
forests at Panaytayum.

A very distinct and characteristic species recognizable by its lax, very
few-flowered panicles, long-pedicelled flowers, etc. It is probably most

closely allied to Dysoxylum pauciflorum Merr., but is entirely distinct from
that species.

PHILIPPINE PLANTS, IX. 279

TOONA Roemer.

TOONA PAUCIJUGA sp. nov.

Species ut videtur 7. calantas Merr. & Rolfe valde affinis,
differt foliolis paucioribus, 7 vel 9, glabris, subtus in axillis
venarum prominente glandulosis.

A tree reaching a height of about 10 m, glabrous throughout
(inflorescence unknown). Branches brownish, glabrous, some-
what striate when dry. Leaves alternate, odd-pinnate, 30 to
35 cm long. Leaflets 7 or 9, oblong-ovate to elliptic-ovate, firmly
chartaceous, brownish and of nearly the same color on both
surfaces when dry, shining, the median ones 10 to 13 cm long,
5 to 6 cm wide, the lower ones somewhat smaller, opposite or
subopposite, entire, the base inequilateral, rounded, or on one
side of the lamina sometimes acute or acuminate, the apex dis-
tinctly acuminate, the acumen usually somewhat falcate, blunt;
lateral nerves 10 to 12 on each side of the midrib, distinct, their
axils on the lower surface with distinct glands; petiolules 1 em
long or less. Flowers unknown. Panicles terminal, in fruit
about 12 cm long. Mature fruits quite similar to those of Toona
calantas, about 3.5 cm long.

LEYTE, between Dolores and Ormoc, in forests at an altitude of about
100 m, For. Bur. 12618 Rosenbluth, February, 109, locally known as calantas.

A species well characterized by its few leaflets which are prominently
glandular in the axils of the veins on the lower surface.

DICHAPETALACEAE.
DICHAPETALUM Thou.

DICHAPETALUM CILIATUM sp. nov.

Frutex scandens, ramulis petiolis foliisque utrinque ad costa
nervisque insigniter ciliatis; foliis oblongis vel oblongo-ellipticis,
usque ad 14 cm longis, chartaceis, apice acuminatis, basi acutis
vel subrotundatis, nervis utrinque circiter 8, prominentibus;
cymis axillaribus, brevibus, petiolo subaequantibus; fructibus
dense griseo-villosis, 14 mm longis, 2-locellatis.

A scandent shrub, prominently ciliate with long, spreading
hairs. Branches slender, terete, reddish-brown when dry, the
older ones nearly glabrous, the younger ones prominently ciliate
with rather pale hairs. Leaves oblong to elliptic-oblong, 8 to
14 cm long, 3 to 6 cm wide, chartaceous, somewhat narrowed
to the acute or subrounded base, the apex rather slenderly and
sharply acuminate, the acumen 0.5 to 1.5 em long, the margins
and the midrib and nerves on both surfaces prominently ciliate
with spreading, pale or somewhat brownish hairs, otherwise

280 MERRILL.

glabrous, both surfaces shining, the lower of about the same
color as, or a little paler than, the upper; lateral nerves about
8, very prominent on the lower surface, curved-ascending, anas-
tomosing, the primary reticulations rather lax, prominent; pe-
tioles 5 to 10 mm long, prominently ciliate. Cymes axillary,
small, sessile or subsessile, dense, not longer than the petioles,
ciliate, many-flowered, the bracteoles lanceolate, 2 mm long or
less. Sepals oblong, obtuse, 2 mm long, outside densely pubes-
cent. Petals narrowly oblong-spatulate, the apex broad, retuse
or slightly cleft, the cleft less than 0.5 mm deep, the base nar-
rowed. Filaments about 2mm long; anthers 1mm long. Ovary
pubescent. Fruit broadly ellipsoid, more or less compressed,
densely gray-pubescent, about 1.5 cm long, divided longitudinally
by deep lateral grooves and slightly retuse at the apex, 2-celled.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9293 (type),
9294, 92384 Whitford & Hutchinson, December, 1907 and January, 1908,
Bur. Sci. 11819 Robinson, July, 1910, in forests at low altitudes.

A species well caracterized by its prominently ciliate branchlets, in-
florescence, and leaves, the latter glabrous except for the ciliate midrib
and lateral nerves on both surfaces and usually also the margins.

DICHAPETALUM ROBINSONII sp. nov.

Arbor parva, circiter 8 m alta, subtus foliis ramulis inflores-
centiisque dense sericeo-villosis; foliis subcoriaceis, late ellipticis
vel ovato-ellipticis, usque ad 15 cm longis, supra nitidis, parce
pubescens, basi rotundatis, apice acutis, nervis utrinque circiter
9, prominentibus; petiolo crasso, circiter 4 mm longo; inflores-
centiis in axillis superioribus vel subterminalibus, breviter pe-
dunculatis, floribus congestis ; fructibus obovoideis, dense breviter
fulvo-villosis.

A small tree, reaching a height of 8 m, the branches terete,
reddish-brown, somewhat pubescent, the younger branchlets
densely covered with short, villous, fulvous hairs, as are the
petioles, inflorescence, and lower surfaces of the leaves. Leaves
subcoriaceous, broadly elliptic to ovate-elliptic, 10 to 14 cm long,
6 to 8 cm wide, the upper surface brownish-olivaceous when
dry, shining, sparingly pubescent, the hairs more numerous on
the midrib and nerves, the lower surface densely fulvous-pubes-
cent with short hairs; petioles about 4 mm long; lateral nerves
about 9, prominent, the primary reticulations rather lax, distinct.
Inflorescence in the upper axils and subterminal, shortly pe-
duncled, up to 4 cm long, all parts fulvous-villous, the bracteoles
linear, 1 to 1.5 mm long. Sepals ovate-lanceolate, acute or
somewhat acuminate, about 2 mm long, externally densely pubes-

PHILIPPINE PLANTS, IX. 281

cent. Petals glabrous, elliptic to obovate, cleft to the middle.
Filaments as long as the sepals; anthers about 0.5 mm long.
Fruit apparently obovoid (immature), 1 to 1.5 em long, externally
densely fulvous-pubescent, apparently 1-seeded.

MINDANAO, District of Zamboanga, Port Banga, Bur. Sci. 11771 Robinson,
July 5, 1910, growing at the edge of the forest.

A species very similar in appearance Dichapetalum luzoniense Merr. &
Rolfe, and distinguishable only by some minor characters. The leaves
are quite different in shape, not narrowed to the base, acute, not at all
acuminate, with a few more nerves and more densely pubescent on the
lower surface than in the Luzon species. D. luzoniense Merr. & Rolfe is
manifestly a woody vine, while D. robinsonii is described as a tree, reclining,
and reaching a height of 8 m. The inflorescence is also subterminal, not
strictly axillary, and not long-peduncled.

ANACARDIACEAE.
PARISHIA Hook. f.

PARISHIA MALABOG sp. nov.

Arbor alta, glabra, vel partibus junioribus inflorescentiisque
minute puberulis; foliis 30 ad 40 em longis, imparipinnatis;
foliolis 11 ad 15, nitidis, crasse coriaceis, oblongis vel oblongo-
ovatis, acuminatis, basi obliquis, valde inaequilateralibus, sub-
acutis vel rotundatis, usque ad 14 cm longis; paniculis folia
aequantibus, glabris vel obscure puberulis; floribus breviter
pedicellatis, petalis quam sepalis vix longioribus, segmentis
calycinis auctis, anguste oblongis, obtusis, usque ad 10 cm longis.

A tall tree, reaching a height of 25 m, glabrous or nearly so.
Ultimate branches thickened, 1 to 1.5 cm thick, grayish or
brownish, marked with large petiolar scars, glabrous, or the
growing tip somewhat puberulent. Leaves somewhat crowded
at the ends of the branchlets, 30 to 40 cm long, their petioles
stout. Leaflets 11 to 15, oblong to oblong-ovate, thickly coria-
ceous, 8 to 15 cm long, 4 to 6 cm wide, shining, inequilateral,
entire, the apex acuminate, the base oblique, one side of the
lamina much broader than the other, usually broadly rounded,
the narrower side usually acute; nerves about 14 on each side
of the midrib, prominent, anastomosing, the reticulations distinct;
petiolules about 8 mm long. Panicles in the upper axils, as
long as the leaves, glabrous or very slightly puberulent, their
rachises stout, straight, the branches rather few, scattered, the
lower ones 10 cm long or less, the upper ones shorter. Flowers
4-merous, short-pedicelled. Calyx at anthesis slightly puberu-
lent, 4-lobed, the lobes about 6 mm long, obtuse, somewhat
spreading. Petals 4, triangular-ovate, obtuse, glabrous, reti-

282 MERRILL.

culate, imbricate, about 4 mm long. Ovary ovoid, pubescent,
1-celled, with a single ovule pendulous from near the apex,
tapering into the short style; style trifid, sometimes bifid; stigmas
capitate; rudimentary stamens 4, inserted outside of the disk,
the filaments 1 mm long, the anthers as long as the filaments.
Male flowers not seen. Fruit densely villous, about 2 cm long,
the accrescent calyx-tube about 2.5 cm in diameter, 1.5 cm long,
the lobes when young reddish or reddish-brown, when mature
brown, narrowly oblong, rounded, up to 10 cm long, 1 to 1.2 em
wide.

CEBU, Buacao, Bur. Sci. 11109 Ramos, March 19, 1912 (type), with
leaves, immature fruits, and female flowers. Ticao, For. Bur. 125380, 12540
Rosenbluth, December, 1908, sterile specimens. MASBATE, For. Bur. 21030
Darling, May, 1910, sterile, Foxworthy. Nercros, For. Bur. 17356 Curran,
September, 1909, seedlings and fallen fruits. TasBuas, For. Bur. 19526 José.
MINDoRO, For. Bur. 9831 Merritt.

This very characteristic species is most closely allied to Parishia insignis
Hook. f. of Tenasserim and the Andaman Islands, and Penang, although
quite different from that species. It is the first representative of the genus
to be found in the Philippines.

Rosenbluth notes that it is the most common tree species on the Island
of Ticao, especially abundant on dry rocky hills, and that the trunk has
a clear length seldom exceeding 15 meters. He notes also that it has
abundant white juice in the bark and leaves. In Ticao the lumber is not
used by the natives, but in Masbate it is used for making bancas, that is,
dugout canoes. Mr. Elmer has noted on two of the sheets of sterile
specimens “Anacardiaceae fide Radlkofer, see Elmer 12164.” The inference
is that Elmer 12164, a number apparently undistributed as yet, represents
the same species. Our material shows conclusively that the plant is refer-
able to Parishia, and the specific name is taken from its current Visayan

name, maldbog.
SEMECARPUS L.

SEMECARPUS ACUMINATISSIMA sp. nov.

Arbor parva (vel frutex?) inflorescentis exceptis glabra vel
subglabra; foliis anguste oblongo-ellipticis ad oblongo-oblanceo-
latis, chartaceis vel subcoriaceis, usque ad 20 cm longis, basi
angustatis, acutis, apice subcaudato-acuminatis, subtus pallidio-
ribus haud glaucescentibus, nervis prominentibus, utrinque
circiter 16; paniculis terminalibus, brevibus, angustis, quam
folia multo brevioribus, parce pubescentibus vel puberulis,
floribus masculinis parvis, 5-meris.

A small tree or a shrub, except the inflorescence nearly gla-
brous. Branches terete, slender, grayish. Leaves oblong-
elliptic to oblong-oblanceolate or narrowly oblong-ovate, 13 to
20 cm long, 4.5 to 6 em wide, chartaceous or subcoriaceous,
narrowed from about the middle or below to the acute base,

PHILIPPINE PLANTS, IX. 283

the apex prominently and rather slenderly subcaudate-acumi-
nate, the acumen straight or somewhat curved, 1 to 2 em long,
the upper surface glabrous, when dry somewhat olivaceous,
shining, the lower surface pale but not glaucous, slightly shin-
ing, very minutely puberulent or nearly glabrous; lateral nerves
about 16 on each side of the midrib, prominent, spreading, anas-
tomosing, the reticulations distinct, the primary ones slender,
leaving the nerves at about right angles and anastomosing with
the secondary lateral nerves, the ultimate free ends of the
nervules rather distinct; petioles 1 to 2 em long. Panicles ter-
minal, slender, narrowly pyramidal, about 7 cm long, the lower
branches about 2 cm in length, puberulent or somewhat pubes-
cent with grayish hairs, the branchlets more or less ferruginous.
Male flowers somewhat crowded on the ultimate branchlets, their
pedicels 1 to 1.5 mm long, the bracteoles lanceolate, acuminate,
pubescent, 1 to 1.4 mm long. Calyx pubescent, 2.5 mm in diam-
eter, 5-lobed, the lobes broadly ovate, obtuse, puberulent. Pet-
als prominently punctate-striate, oblong-ovate, obtuse, 2 mm
long. Disk 1 mm in diameter, ferruginous-pubescent. Anthers
1 mm long. Immature fruits with puberulent pedicels 5 to 8
mm long, the hypocarpium puberulent, when dry about 4 mm
long and wide, the drupe obliquely ovoid, somewhat compressed,
5 to 6 mm long, very slightly pubescent, apparently becoming
quite glabrous.

LUZON, Province of Tayabas, Kabibihan, Bur. Sci. 13241 Ramos, February
27, 1911, in forests along streams, ¢ flowers: Province of Zambales, Bur.
Sci. 5058 Ramos, December, 1907, with immature fruits.

A species manifestly allied to Semecarpus philippinensis Engl., but
distinguished by its short and slender panicles, and its differently shaped
leaves which are prominently subcaudate-acuminate and usually obscurely
puberulent beneath.

SEMECARPUS EUPHLEBIA sp. nov.
Arbor circiter 18 m alta, inflorescentiis puberulis exceptis
glabra; foliis crassissime coriaceis, oblongis vel anguste oblongo-
ovatis, apice obtusis, basi acutis, usque ad 20 cm longis, supra
nitidis, subtus glaucescentibus, nervis utrinque circiter 20, valde
prominentibus, petiolo 4 ad 5 cm longo; paniculis folia subae-
quantibus, anguste pyramidatis, puberulis, floribus 5-meris.

A tree about 18 m high, glabrous except the puberulent inflor-
escence. Branches stout, brownish, the ultimate ones about 8
mm in diameter. Leaves very thickly coriaceous, oblong to
narrowly oblong-ovate, 16 to 20 cm long, about 6 cm wide,
narrowed upward to the obtuse apex, and below to the acute

284 MERRILL.

base, of about the same color on both surfaces when dry, rather
pale, the upper one shining, the lower more or less glaucous,
dull; nerves about 20 on each side of the midrib, very promi-
nent on the lower surface, spreading, parallel, anastomosing
near the margins, the primary reticulations leaving the veins
at right angles and anastomosing with intermediate secondary
veins, the ultimate reticulations not prominent; petioles stout,
4to5cm long. Panicles terminal, about as long as the leaves,
uniformly and somewhat olivaceous-puberulent, narrowly pyra-
midal, the lower branches 5 to 6 cm long. Flowers unknown,
but the persistent sepals 5. Hypocarpium 5 to 6 mm long, 2
mm wide at the base, 4 mm wide at the apex, longitudinally
obscurely sulcate, slightly puberulent. Drupe (very immature)
7 mm long, 5 mm in diameter, oblong-ovoid, slightly compressed,
sparingly pubescent at the apex, apparently becoming entirely
glabrous.

LuzoN, Province of Tayabas, Quinatacutan River, Bur. Sci. 13163 Fozx-
worthy & Ramos, March, 1911, in forests along the river.

Apparently a very characteristic species, its distinguishing features being
its very coriaceous, glabrous, comparatively long-petioled, prominently nerved
leaves, its narrowly pyramidal puberulent panicles, its hypocarpium much
longer than wide, and its oblong-ovoid drupes.

SEMECARPUS LANCEOLATA gp. nov.

Arbor parva, glabra; foliis crasse coriaceis, lanceolatis, usque
ad 30 cm longis, utrinque angustatis, apice obtusis, basi acutis,
nervis utrinque circiter 22, haud prominentibus, in siccitate
supra nitidis, subtus pallidioribus, vix glaucescentibus, petiolo
5 ad 7 cm longo; paniculis glabris, ramis inferioribus circiter
13 cm longis, drupis reniformibus, compressis, leviter carinatis,
2 cm latis.

A small tree, apparently glabrous throughout (inflorescence
and flowers not seen.) Branches stout, the ultimate ones 1.5
cm in diameter, lenticellate, dark-brown, shining. Leaves lan-
ceolate, 20 to 30 cm long, 4 to 7 cm wide, narrowed at both
ends, the apex obtuse, base acute, thickly coriaceous, when dry
shining, the upper surface rather pale, the lower one paler than
the upper but not glaucous; lateral nerves about 22 on each side
of the midrib, not prominent, spreading, faintly anastomosing,
the reticulations very irregular, the ultimate ones dense, indis-
tinct; petioles 5 to 7 cm long. Panicles terminal, in fruit quite
glabrous, the lower branches about 12 cm long. Flowers un-
known. Hypocarpium, when dry, about 1 cm long, 8 mm wide
at the tip, narrowed below, apparently very much larger when
fresh, the peduncle usually about 1 cm long. Drupe reniform,

PHILIPPINE PLANTS, IX. 285

compressed, about 2 cm wide, 1.3 cm long, somewhat keeled,
nearly or quite smooth when dry.

Luzon, Abra Subprovince, Dolores, Bur Sci. 7083 Ramos, February, 1909,
in forests, fruit red.

A very characteristic species, distinguishable by being entirely glabrous,
by its lanceolate, elongated, coriaceous, long-petioled leaves, and by its
comparatively large drupes which are reniform and distinctly keeled.

SEMECARPUS MAGABOTRYS sp. nov.

Arbor dioica ramulis subtus foliis inflorescentiisque ferrugi-
neo- vel pallide fulvo-villosis; foliis anguste oblongo-obovatis ad
obovato-lanceolatis, crasse coriaceis, usque ad 22 cm longis, apice
obtusis, acutis, vel rotundatis, basi angustatis, abrupte obtusis
vel rotundatis, supra glabris, in siccitate nitidis, subtus pallidis;
nervis primariis utrinque circiter 22, prominentibus; paniculis
terminalibus, 40 ad 50 cm longis.

A dioecious tree probably of small or medium size. Branches
stout, brownish, the younger ones rather prominently pubescent.
Leaves narrowly oblong-obovate to obovate-lanceolate, thickly
coriaceous, the apex acute, blunt, or rounded, the base narrowed
and abruptly obtuse or rounded, 15 to 22 em long, 5 to 8 cm
wide, when dry the upper surface rather pale, shining, glabrous,
the lower surface much paler than the upper one, more or less
glaucous, and rather densely villous with pale-brownish hairs;
lateral nerves about 22 on each side of the midrib, very promi-
nent on the lower surface, spreading or somewhat ascending,
the secondary ones subparallel,- prominent, the ultimate reticu-
lations distinct; petioles stout, pubescent, 1 to 1.5 cm long.
Panicles pyramidal, terminal, 40 to 50 em in length, ferruginous-
or fulvous-villous, the flowers very numerous. Male flowers
fascicled or glomerate at the nodes, the calyx 2 mm in diameter,
pubescent, 5-toothed, the teeth short, triangular. Petals 5,
oblong-ovate, acute,2 mm long. Disk about 1.3 mm in diameter.
Female flowers: Calyx pubescent, 3 mm in diameter. Petals 5,
oblong-ovate, acute, 3 mm long. Ovary depressed, densely fer-
_ ruginous-villous, about 2 mm in diameter. Young fruit with
pedicels 1.5 to 2 cm in length, the hypocarpium rather densely
ferruginous-villous, 4 mm long, about 5 mm wide when dry,
the drupe somewhat compressed (immature), obovoid, about 7
mm long, with scattered, apparently deciduous, ferruginous hairs,
apparently ultimately glabrous.

Luzon, Province of Nueva Vizcaya, For. Bur. 22359 Alvarez, January,
1911, one specimen with male flowers, one with female flowers (type), and
one with immature fruits.

286 MERRILL.

Probably as closely allied to Semecarous perrottetti March., as to any
other species, but at once distinguished by its indumentum, its more or
less pubescent fruits (at least the hypocarpium), and its very large panicles.

SEMECARPUS OBTUSIFOLIA sp. nov.

Arbor circiter 20 m alta, inflorescentiis exceptis glabra; foliis
ellipticis, usque ad 13 cm longis, crasse coriaceis, apice late
rotundatis, basi acutis, in siccitate supra subolivaceis, nitidis,
subtus pallidis, glaucescentibus, nervis utrinque circiter 10, valde
prominentibus; paniculis terminalibus pyramidatis, quam folia
multo longioribus, plus minusve ferrugineo-hirsutis; floribus 5-
meris. ‘

A tree about 20 m high, the branches slender, terete, grayish,
sparingly lenticellate. Leaves thickly coriaceous, elliptic or sub-
elliptic, 7 to 13 cm long, 3.5 to 7 cm wide, the apex very broadly
rounded, the base abruptly acute, glabrous, when dry the upper
surface olivaceous, shining, the lower surface pale, glaucous;
lateral nerves about 10 on each side of the midrib, prominent
on the lower surface, spreading and somewhat ascending, curved,
anastomosing, the primary reticulations lax, distinct, leaving the
nerves at right angles, the ultimate ones indistinct; petioles 8
to15 mm long. Panicles terminal, pyramidal, about 20 cm long,
the branches scattered, the lower ones up to 12 cm in length,
the younger parts rather prominently ferruginous-pubescent or
hirsute. Flowers subglomerately arranged on the ultimate
branchlets, sessile, 5-merous, numerous. Male flowers: Calyx
2 mm in diameter, 5-lobed, the lobes obtuse or rounded. Petals
oblong-ovate, obtuse, 1.5 mm long. Disk ferruginous-pubescent.
Anthers 0.8 mm long.

BALABAG, in forests, Bur. Sci. 512 Mangubat, March 4, 1906.

A species distinguishable at once by its elliptic, broadly rounded leaves
which are olivaceous above and glaucous beneath.

SEMECARPUS PAUCINERVIA sp. nov.

Arbor parva circiter 10 m alta, inflorescentiis exceptis glabra;
foliis anguste oblongo-obovatis vel oblongo-ellipticis, coriaceis,
nitidis, apice late rotundatis, basi acutis, usque ad 9 cm longis,
nervis utrinque circiter 8, arcuatis, subtus distinctis; paniculis
leviter hirsutis, laxis, quam folia multo longioribus, multifloris,
floribus ¢ in fasciculis racemosis, distincte pedicellatis, 5-meris.

A small tree about 10 m high. Branches terete, grayish,
slightly lenticellate, shining, slender, glabrous, the growing parts
slightly pubescent. Leaves narrowly oblong-obovate to oblong-
elliptic, 5 to 9 cm long, 2 to 3 em wide, coriaceous, the apex

PHILIPPINE PLANTS, IX. 287

broadly rounded, the base usually narrowed, acute, glabrous and
shining on both surfaces, when dry the upper surface rather
pale, the lower one much paler and somewhat glaucous; nerves
about 8 on each side of the midrib, curved-ascending, anastomos-
ing, distinct on the lower surface, the primary reticulations lax,
leaving the nerves at right angles, the ultimate reticulations
obscure; petioles 5 to 7 mm long. Panicles terminal and in the
upper axils, sparingly hirsute with short, scattered, pale-brownish
hairs, lax, pyramidal, 12 to 20 cm in length, the lower branches
up to 10 cm in length. Male flowers in scattered fascicles which
are racemosely arranged on the ultimate branches, their pedicels
slender, 1.5 to 2 mm long, the bracteoles ovate-lanceolate, 0.5
mm long. Calyx 1.5 mm in diameter, 5-lobed, the lobes ovate,
obtuse, 0.5 mm long. Petals oblong-ovate, obtuse or acute, 1.8
mm long. Disk somewhat ferruginous-pubescent. Anthers 1
mm long.

PALAWAN, Mount Victoria, Bur. Sci. 570 Foxworthy March 25, 1906, on
river banks, altitude about 180 meters.

A very distinct species, characterized by its small, few-nerved leaves
which are broadly rounded at their apices, and by its lax panicles, the
flowers being distinctly and slenderly pedicelled.

SEMECARPUS PILOSA sp. nov.

Arbor parva ut videtur S. cuneiformis affinis, differt foliis sub-
tus densissime pilosis, paniculis ferrugineo-pilosis.

A tree apparently of small size, the branches stout, pubescent.
Leaves more or less crowded at the ends of the branches, thickly
coriaceous, oblong-obovate, 9 to 20 cm long, 4 to 8 em wide, the
apex broadly rounded, sometimes retuse, narrowed below to the
abruptly obtuse, not at all decurrent base, the upper surface
somewhat grayish when dry, shining, quite glabrous except for
the more or less pubescent midrib, the lower surface densely
pilose, paler than the upper one; lateral nerves 13 to 16 on
each side of the midrib, very prominent, spreading, somewhat
curved and anastomosing, the secondary nerves leaving them
at right angles, subparallel, distinct; petioles stout, pubescent,
8 to 10 mm long. Panicles terminal, pyramidal, ferruginous-
pilose, about 15 cm long, the branches spreading, the lower ones
about 8 cm in length. Petals 5, ovate to oblong-ovate, somewhat
acuminate, 2.5 mm long. Ovary ovoid, densely fulvous-villous.
Hypocarpium, when fresh, orange-yellow, when dry about 4
mm long nearly as thick as the drupe, more or less ferruginous-
pubescent, the drupe slightly compressed, obliquely ovoid, obtuse,

288 MERRILL.

smooth, black, about 8 mm long, slightly pubescent externally,
ultimately becoming glabrous or nearly so, the pericarp leathery.
Luzon, Province of Nueva Vizcaya, Dupax, Bur. Sci. 11435 McGregor,
March, 1912, locally known as baliunasoy.
A species well characterized by its leaves being densely pilose on the

lower surface. Like most or all the species in the genus, the pericarp
of the fruit contains a black resin.

SEMECARPUS WHITFORDII sp. nov.

Species S. cuneiformis Blanco similis et ut videtur valde
affinis, differt foliis distincte acuminatis, paniculis dense pallide
fulvo-pilosis.

A small tree, the branches terete, brownish, slightly sulcate
when dry, more or less pubescent. Leaves oblong-oblanceolate,
thinly coriaceous, 14 to 24 cm long, 3 to 7 cm wide, narrowed
from above the base to the acute, and usually distinctly inequi-
lateral base, the apex distinctly acuminate, the acumen rather
sharp, less than 1 cm long, the upper surface pale and shining
when dry, glabrous or very slightly pubescent along the midrib,
the lower surface rather pale-fulvous-pilose, the hairs rather
short, spreading, more dense on the midrib and lateral nerves;
lateral nerves 15 to 17 on each side of the midrib, prominent,
spreading, anastomosing, the reticulations rather close, distinct;
petioles 1 to 2.5 cm long, somewhat pubescent. Panicles ter-
minal, ample, diffuse, about 25 cm long, the lower branches up
to 12 em long, the upper ones shorter, spreading, all parts rather
densely pilose with pale-fulvous, spreading hairs. Flowers very
numerous, 5-merous, more or less glomerate on the ultimate
branchlets, sessile or subsessile. Male flowers 4 mm in diameter
in anthesis. Calyx villous externally, shallowly cup-shaped, the
lobes orbicular-ovate, obtuse, 0.5 to 0.8 mm long, prominently
striate-reticulate, the reticulations dark-colored. Petals oblong-
ovate, acute, pubescent externally, reticulate-striate like the
calyx. Disk glabrous, shallowly 5-crenate. Rudimentary ovary
very small, densely fulvous-villous. Filaments filiform, 3 mm
long; anthers ovoid, 0.8 mm long. Female flowers and fruits
not seen.

MINDANAO, District of Cotabato, Lebak, For. Bur. 11774 Whitford, March
4, 1912, in dipterocarp forests at low altitudes.

A species similar to, and certainly closely allied to Semecarpus cuneti-
formis Blanco, from the typical form of which it differs in its distinctly
acuminate leaves which are pale beneath but scarcely glaucous, and by its
densely pilose panicles, the indumentum of a pale-fulvous color. It is

apparently more closely allied to Blanco’s species than to any of the
numerous forms recently described by Doctor Perkins.

PHILIPPINE PLANTS, IX. ; 289

SEMECARPUS CUNEIFORMIS Blanco FI. Filip. (1837) 220, ed. 2 (1845)

155, ed. 8, 1: 276, Naves l. c. ed. 3, pl. 75.

Semecarpus perrottetii March. Rév. Anac. (1869) 169; Engl. in DC.
Monog. Phan. 4 (1883) 380; Vid. Rev. Pl. Vase. Filip. (1886) 101;
Perk. Frag. Fl. Philip. (1904) 28.

Semecarpus anacardium Blanco FI. Filip. (1837) 217, ed. 2 (1845)
152, non L. f.

Semecarpus microcarpa F.-Vill. Novis. App. (1880) 55, non Wall.

Semecarpus pubescens F.-Vill. 1. ¢., non Thwaites.

Semecarpus siderowyloides Perk. Frag. Fl. Philip. (1904) 28.

This species is common and widely distributed in the Philippines, and
is the most abundant one of the genus found in the Archipelago. There
is no valid reason why Blanco’s specific name, cuneiformis, should not be
adopted for the species, although like many of Blanco’s descriptions, that
of Semecarpus cuneiformis is short and imperfect. The one distinctive
character that he gives is that the leaves are retuse at the apex, which
is true of a number of specimens manifestly referable to Semecarpus
perrottetii March.

The type of Semecarpus perrottetii March. was collected by Perrottet
in Luzon, and is preserved in the Herbarium of the Paris Museum of
Natural History. Two sheets are so named by Marchand, one of which
bears Perrottet’s note “trés comun .... 4 Manille,” indicating that the type
was collected in Manila. The species is still very common in and about
the city, and in essential characters is rather constant. There is considerable
variation in the shape of the leaves, their apices varying from broadly
rounded to retuse on the one hand, and to acute or shortly and broadly
acuminate on the other. Perrottet’s type has broadly rounded leaves,
according to carbon rubbings kindly made for me by Dr. C. B. Robinson,
and according to a carbon rubbing made by myself of the single leaf of
Perrottet’s plant preserved in the Berlin Herbarium.

Semecarpus anacardium Blanco, non L. f., is manifestly the same
species, although, of course, Blanco may have included in it more than the
common form. Semecarpus anacardium L. f. certainly does not extend
to the Philippines.

Semecarpus microcarpa F.-Vill. is only a mis-identification of Blanco’s
S. cuneiformis on the part of F.-Villar., while S. pubescens F.-Vill., is a
manifest mis-identification of Blanco’s S. anacardium.

Semecarpus sideroxyloides Perk., is typical S. perrottetii March., = S.
cuneiformis Blanco, and is accordingly reduced. A number of specimens
referred by Doctor Perkins to Semecarpus perrottetii March.’ are not
properly referable to that species.

I consider Semecarpus cuneiformis Blanco (S. perrottetii March.) to be
well represented by the following specimens:

LuzON, Province of Cagayan, For. Bur. 16983 Bacani, For. Bur. 18€05
Klemme: Province of Ilocos Norte, Bur. Sci. 7676 Ramos: Bontoe Sub-
province, For. Bur. 10981 Curran: Province of Union, Elmer 5562, 5637:
Province of Pangasinan, Alberto 83: Province of Zambales, For. Bur.
6977 Curran, Merrill 3010, Sankuhl s. n., Hallier s. n.: Province of Bulacan,
Mrs. Templeton s. n. MANILA, Perrottet (type of S. perrotteti March.,

Frag. Fl. Philip. (1904) 28.

290) MERRILL.

in herb. Paris.), Malvar 349, Cuzner 55, Marave 30, Ahern 716: Province of
Rizal, Pasay, For. Bur. 15385 Curran & Schneider; Antipolo, Merrill 1720;
Montalban, Loher 5850: Province of Bataan, Merrill 1484, For. Bur. 1305,
2720 Borden, For. Bur. 2513 Meyer, Ahern 769, Williams 646, 632: Province
of Laguna, Hallier s. n. Minporo, For. Bur. 8813 Merritt, Merrill 2160
(cotype of Semecarpus sideroxyloides Perk.). GUIMARAS, For. Bur. 6477
Everett. Leyte, Elmer 7159. NeEcGros, Bur. Sci. 7348 Celestino, For. Bur.
15159 Tarrosa.

The common Tagalog name is ligas, the Visayan agas and anagas, the
Ilocano camiring, the Pampangan caming, and the Bontoe pacan.

This species is the most common and the best known contact-poison in
the Philippines, its noxious properties being well known to the natives. As
with the well known contact-poisons in the United States, such as Rhus
toxicodendron L., and R. venenata DC. some persons are immune while
others are very subject to it. The symptoms caused by contact with fresh
leaves of Semecarpus cuneiformis Blanco, especially when the leaves are
wet, are quite like those caused by the species of Rhus mentioned above,
and the eruption is amenable to the same treatment.

SEMICARPUS MICRANTHA Perk. Frag. Fi. Philip. (1904) 27.

Semecarpus taftiana Perk. 1. c. 28.

But a single species is represented, for which the name micrantha is
retained, this having page priority. The type of Semecarpus micrantha
Perk., is a specimen with staminate flowers, while the type of S. taftiana
Perk., is a specimen with pistillate flowers. They are from the same locality,
with the same habitat, and were collected at the same time, February 15,
1903. In inflorescence, twigs, indumentum, color, shape, and venation of
the leaves the specimens are quite the same. The only evident difference
is that the leaves of Semecarpus taftiana are somewhat smaller and
relatively a little narrower than are those of S. micrantha.

PALAWAN, Iwahig River, Merrill 709, 3 flowers, cotype of Semecarpus
micrantha Perk., 734, 2 flowers, cotype of Semecarpus taftiana Perk., both
collected February 15, 1903.

SEMECARPUS PHILIPPINENS!S Engl. in DC. Monog. Phan. 4 (1883)
481.

Semecarpus perrottetii March. var. glabra March. Rev. Anac. (1869)
170.

This species is rather common and of wide distribution in the Philippines,
and is decidedly variable in the shape of its leaves, although in essential
characters quite constant. The type of the species and of Semecarpus
perrottetii var. glabra March. is Cuming 1146, from the Province of Ilocos
Sur, Luzon, two sheets of which are in the Herbarium of the Bureau of
Science. Engler’ has referred to another Philippine specimen Cuming
1776, which was collected in Cebu, as apparently being referable to Seme-
carpus albescens Kurz. Specimens of this number are in the Herbarium
of the Bureau of Science, and while they are not identical with the type
number of Semecarpus philippinensis, I can, after examining a large series
of specimens, showing numerous intermediate forms, detect no characters

6L. ¢. 489.

PHILIPPINE PLANTS, IX. 991

that warrant the recognition of two distinct species, and accordingly refer
Cuming 1776 to Semecarpus philippinensis Engl. In degree of distinctness
the ultimate reticulations in the two specimens differ considerably, and by
Engler are accordingly placed in different sections in his key to the species.
This character, however, is not at all constant, and all intergrades occur.
I refer to Semecarpus philippinensis Engler the following material:

LuZzoON, Province of Abra, Bur. Sci. 7209 Ramos ¢ flowers: Province of
Ilocos Sur, Cuming 1146 3 flowers (cotype): Benguet Subprovince, Elmer
6447, in fruit: Province of Pampanga, For. Bur. 17669 Curran, & flowers:
Province of Bataan, Lamao River, Williams 905 ¢ flowers, For. Bur. 337
Barnes ¢ flowers, For. Bur. 674, 708, 1193 Borden, all in fruit, For. Bur.
2777 Meyer @ flowers: Province of Tayabas, For. Bur. 10382 Curran. LEYTE,
Elmer 7117, Piper 601. CEBU, Cuming 1776.

In this material the leaves vary in shape and size, the aa in the type
being rounded or obtuse, in some leaves subacute, but in other specimens
it varies to acute or acuminate.

HIPPOCRATEACEAE.
SALACIA L.

SALACIA PHILIPPINENSIS sp. nov.

Frutex scandens, glaber; foliis oppositis, oblongis, coriaceis,
nitidis, usque ad 15 cm longis, basi acutis, apice obtusis vel late
obscure acuminatis, margine subintegris vel leviter crenatis;
inflorescentiis cymosis, dichotomis, pedunculis quam petiolo long-
ioribus; floribus circiter 8 mm diametro.

A scandent shrub, glabrous, the branches slender, terete, dark-
colored when dry, with numerous small lenticels, the younger
ones somewhat compressed. Leaves opposite, oblong, coriaceous
or subcoriaceous, 12 to 15 cm long, 5 to 6 cm wide, shining,
when dry the lower surface a little paler than the upper one,
the apex obtuse or shortly and broadly acuminate, the base
acute, the margins entire or very obscurely and distantly crenate;
lateral nerves 7 or 8 on each side of the midrib, curved-ascending,
anastomosing, distinct; petioles jointed near the base, 5 to 8
mm long. Inflorescence axillary, solitary, cymose, dichotomous,
the peduncles a little longer than the petioles, the cymes about
3 cm long and wide, rather many flowered. Flowers greenish-
yellow, their pedicels about 4 mm long, subtended by very short,
obscure bracteoles, apparently jointed to the inflorescence. Calyx
about 3 mm in diameter, the lobes 5, rarely 4, orbicular-ovate,
usually rounded, minutely ciliate on the margins. Petals 5,
rarely 4, oblong, spreading, broadly rounded at the apex, about
4 mm long, 2 mm wide. Stamens 3; filaments about 1.2 mm
long; anthers 0.5 mmin diameter. Ovary 3-celled; cells 2-ovuled,

113111——3

292 MERRILL.

narrowed above into the short narrowly conical style, the disk
cylindric, prominent. Fruit unknown.

LUBANG ISLAND, Merrill 977, April 8, 1903, a shrub about 3 m high
sprawling or climbing in thickets near sea-level. CEBU, Bur. Sci. 11095
Ramos, March, 1912, in thickets.

This apparently undescribed species seems to be allied to Salacia perak-
ensis King, and to S. griffithii Laws., but is entirely different from both.
Mature fruits have not been seen, but they were sufficiently developed to
show that the plant is a Salacia and not a Hippocratea.

ICACINACEAE.
FREERIA gen. nov.

Flores dioici, spicati. Perianthium 4-merum, simplex, lobi
valvati, tubo brevi. Fl. 4 -........ Fl. 2 : Ovarium sessile, ]-locul-
are; stigma sessile, crassum, truncatum; ovula 2, ex apice
loculi pendula. Drupa oblonga leviter compressa, epicarpio
carnoso, putamine crustaceo extus punctato, intus papillato.
Semen pendulum; embryo albumine carnoso valde ruminato
parum brevior, cotyledonibus planis, tenuiter foliaceis. Frutex
alte scandens, ramis ramulisque tenuibus. Folia alterna, oblonga
vel oblongo-lanceolota, glabra, repanda, glanduloso-dentata, acu-
minata, basi leviter cordata. Racemi supra-axillares, simplices.
Flores pauci, parvi, sessiles, bracteolati.

FREERIA REPANDA sp. nov.

Frutex scandens subglaber, ramulis teretibus, tenuibus, minut-
issime verruculosis; foliis oblongis vel oblongo-lanceolatis, sub-
coriaceis, usque ad 14 cm longis, in siccitate pallidis, nitidis,
subtus leviter asperulis, nervis utrinque 5 vel 6, prominentibus,
distantibus, anastomosantibus, reticulis laxis, distinctis; spicis
2 ad 4 em longis, pedunculatis, tenuibus; floribus paucis, circiter
2 mm longis, bracteolatis; fructibus oblongis, leviter compressis,
circiter 1.8 cm longis.

A woody vine reaching a height of about 8 m (fide Ramos),
nearly glabrous, the branches and branchlets slender, terete, 1
to 2 mm in diameter, very minutely verruculose, the growing
parts minutely pubescent. Leaves alternate, subcoriaceous,
oblong to oblong-lanceolate, 6 to 14 cm long, 1.5 to 4 cm wide,
rather slenderly acuminate, base slightly narrowed, distinctly
cordate, the margins more or less repand and with a short glan-
dular tooth opposite the excurrent end of each primary lateral
nerve, rather pale, shining, and of about the same color on both
surfaces when dry, the upper surface smooth and glabrous, the

PHILIPPINE PLANTS, IX. 298

lower one minutely scabrid; lateral nerves 5 or 6 on each side
of the midrib, prominent, distant, somewhat curved, anasto-
mosing, the reticulations lax, distinct; petioles 0.5 to 1.5 cm long,
minutely pubescent, becoming glabrous, usually strongly curved.
Spikes supra-axillary, slender, minutely pubescent, 2 to: 4.:e¢m
long, flower-bearing only in the upper one-fourth or one-fifth.
Male flowers unknown. Female flowers few, 10 or less on each
spike, sessile, scattered, each subtended by an oblong-lanceolate,
acuminate, 0.4 mm long bracteole. Perianth simple, 1.8 to 2
mm long, the calyx represented by a very obscure disk, the tube
very short, the lobes 4, valvate, oblong, obtuse, about 0.7 mm
wide, pubescent externally. Staminodes very minute, alternat-
ing with the perianth-lobes. Ovary ellipsoid, densely pubescent,
about 1 mm long, 1-celled; ovules 2, pendulous; stigma sessile,
truncate, round, minutely papillate. Drupe oblong, slightly com-
pressed, glabrous, red or yellowish when mature, about 1.8 cm
long, 1 cm wide, the pulp rather thin, fleshy. Seed about 1.5
long, 8 mm wide, about equally narrowed and acute at both ends,
slightly compressed, the testa crustaceous, externally somewhat
punctate, internally papillose, the papillae projecting into the
interstices of the ruminate albumen. Albumen firm, deeply
ruminated, yellow, oily, externally deeply pitted.

Luzon, Province of Tayabas, Kabibihan, climbing in tall trees in forests
along the river, Bur. Sci. 13289 Ramos, March 38, 1911.

Apparently a very distinct generic type, possibly most closely allied to
Sarcostigma Wight & Arnott, although differing from that genus in many
essential characters, such as the absence of the calyx, or at least the
calyx represented by a very obscure disk, the scattered, not fascicled,
spicate flowers, the repand and glandular-toothed, not entire leaves, and in
{ts albuminous seeds, the albumen being deeply pitted and ruminate.

Dedicated to the late Doctor Paul Caspar Freer, founder and director of

the Bureau of Science from its inception as the Bureau of Government
Laboratories in the year 1901, until his death in April, 1912.

PHYTOCRENE Wall.

PHYTOCRENE LOHERI sp. nov.

Frutex scandens, ramulis foliisque subglaber; ramulis teret-
ibus, 3 ad 4 mm diametro, leviter longitudinaliter striatis, brun-
neis, parce breviter pubescentibus; foliis ovatis, coriaceis, integris,
acuminatis, basi late rotundato-truncatis vel subcordatis, 5-
nerviis, reticulis obscuris; capitulis ut videtur globosis, drupis
numerosis, oblongis, 7 ad 8 cm longis, dense adpresse villosis.

A scandent shrub, nearly glabrous, the branchlets terete, dark-
brown when dry, dull, slightly longitudinally striate, slightly pu-

294 MERRILL.

bescent with scattered hairs. Leaves alternate or subopposite,
ovate, coriaceous, brown and slightly shining when dry, entire,
8 to 10 cm long, 4 to 7 em wide, apex shortly acuminate and
minutely apiculate, base broadly rounded-truncate or subcordate,
the upper surface glabrous, the lower surface at first sparingly
pubescent especially along the nerves, ultimately becoming gla-
brous; basal nerves 5, the outer pair short, the inner pair curved-
ascending, reaching nearly to the middle of the leaf, evanescent,
the lateral nerves above the basal ones usually two on each side
of the midrib, curved-ascending, evanescent or very obscurely
anastomosing, the primary reticulations very lax, not prominent,
the ultimate ones rather close but indistinct; petioles about 2
em long. Heads apparently globose, of many drupes. Individ-
ual drupes oblong or narrowly oblong, 7 to 8 cm long, about
2 cm thick, longitudinally 5-ridged, more or less narrowed at both
ends, densely covered with yellowish-brown or tawny, stiff,
appressed, villous hairs.

LuZzON, Province of Rizal, Montalban, Loher 5769, November, 1905.

A very distinct species, characterized by its comparatively small leaves,

with few, distant nerves, and its obscure reticulations; in this latter charac-
ter it is very different from the other Philippine forms.

SABIACEAE.
MELIOSMA Blume.

MELIOSMA MAGCROPHYLLA sp. nov.

Arbor 12 ad 15 m alta, subglabra, partibus junioribus plus
minusve pubescentibus; foliis imparipinnatis, elongatis, usque
ad 1 m longis; foliolis oppositis vel suboppositis, 15 ad 17, ovatis
ad oblongis vel oblongo-ovatis, integris, acuminatis, basi inaequi-
lateralibus, rotundatis, superioribus usque ad 27 cm longis,
inferioribus minoribus; paniculis amplis, terminalibus axillari-
busque; fructibus circiter 5 mm longis.

A tree 12 to 15 m high, ultimately glabrous or nearly so, the
younger parts more or less ferruginous-pubescent. Leaves
elongated, up to 1 m in length, odd-pinnate, the leaflets 15 to 17,
opposite or the lower ones subopposite, ovate to oblong-ovate or
oblong, coriaceous, puberulent on the midrib and nerves beneath,
the upper ones up to 27 cm in length and 10 cm wide, entire,
acuminate, somewhat falcate, base inequilateral, of the upper
leaves acute or acuminate, of the lower ones broad and rounded,
the lower leaves gradually smaller than the upper ones; nerves 9
to 13 on each side of the midrib. Panicles axillary and terminal,
ample, diffuse, the axillary ones as long as the leaves or shorter.

PHILIPPINE PLANTS, IX. 295

Flowers unknown. Fruit ovoid to globose-obovoid, 5 to 6 mm
long, glabrous.

LuzON, Benguet Subprovince, Sablan, Phil. Pl. 483 Fénix, November,
1910, on slopes along trails, locally known to the Igorots as arocong.

A species distinguishable from the other Philippine ones by its com-
paratively very large leaves, entire leaflets, and in being nearly glabrous,
at least when mature.

ELAEKOCARPACEAE.
ELAEOCARPUS L.

ELAEOCARPUS GRANDIFLORUS Sm. in Rees Cyclop. 12 (1802-20) no. 5;
Pierre in Fl. Forest. Cochinch. pl. 142; Gagnepain in Lecomte
Fl. Gén. Indo-Chine 1 (1910) 568.

BUSUANGA, Bintuan, C. M. Weber 1554 bis, March 3, 1912, common along
streams in forests.

Not previously reported from the Philippines; Burma, Indo-China, and
Java.

The Busuanga specimens appear to be quite typical, at least of the
species as interpreted by Pierre and by Gagnepain. It agrees with
Gagnepain’s description in all essential details, and, so far as the specimens
are comparable, with a Cochinchina collection by Thorel, in the Herbarium
of the Bureau of Science. The Philippine material is in full flower and
with a single nearly mature fruit, while our specimen of Thorel’s collection
has very immature fruits. A specimen from Java, VI, C, 178 of the
Botanical Garden at Buitenzorg, probably typical Elaeocarpus lanceolatus
Blume, which is generally considered to be a synonym of FE. grandiflorus Sm.,
seems to differ from the Philippine material more than does the Cochinchina
material. Although the three specimens before me are not directly com-
parable, all being in different stages of development, they apparently
represent a single species.

ELAEOCARPUS PUSTULATUS sp. nov. § Huelaeocarpus.

Arbor, foliis utrinque sepalisque extus pustulis minutis in-
structis; foliis lanceolatis vel anguste ovato-lanceolatis, acumi-
natis, usque ad 8 ecm longis, glabris, longe acuminatis; racemis
numerosis, e axillis defoliatis; floribus 5-meris, circiter 5 mm
longis.

A tree, glabrous except the sparingly appressed-pubescent in-
florescence, the branches smooth, terete, the branchlets usually
reddish-brown. Leaves lanceolate to narrowly ovate-lanceolate,
5 to 8 cm long, 1.5 to 2.5 em wide, subcoriaceous, the base acute,
the apex slenderly acuminate, margins distantly and somewhat
obscurely toothed, both surfaces with numerous small pustules,
the lower one a little paler than the upper and with glands in
the axils of the lateral nerves; nerves about 6 on each side of
the midrib, not prominent; petioles about 7 mm long, often with
a pair of small lateral glands at the apex. Racemes in the axils

296 MERRILL.

of fallen leaves, numerous, solitary, rather slender, 5 to 7 cm
long, sparingly appressed-pubescent, about 15-flowered. Flowers
5-merous, about 5 mm long, their pedicels 5 to 7 mm in length.
Sepals externally sparingly appressed-pubescent, distinctly pus-
tulate, lanceolate, narrowed upward to the acute or somewhat
acuminate apex, 5 mm long, 1.2 mm wide, inside glabrous or
nearly so. Petals equaling or a little longer than the sepals,
up to 1.8 mm wide, externally glabrous except for very few
hairs at the base, the margins densely villous-ciliate, inside
prominently villous-ciliate in the lower one-half, the apical one-
third split into 9 to 11 linear-filiform segments about 2 mm in
length. Stamens about 20, their filaments 1 mm long or less;
anthers linear, 2 mm long, cleft at the apex, the cells scabrid,
one a little longer than the other and with a tuft of few short
hairs at the tip. Ovary densely villous, 3-celled.

CAMIGUIN DE MINDANAO, in forests, Mount Mahinog, Bur. Sci. 14635
Ramos, April 11, 1912.

A species manifestly allied to Hlaeocarpus verruculosus Aug. DC., which
it strongly resembles, differing especially in its much longer racemes,
long and slender pedicels, and in its much shorter petioles.

TILIACEAE.
TRIUMFETTA L.

TRIUMFETTA PROCUMBENS Forst. f. Prodr. (1786) 35; Hemsl. in
Journ. Bot. 28 (1890) 1, fig. 1; Sprague & Hutchinson in Journ.
Linn. Soc. Bot. 39 (1909) 246; Gagnepain in Not. Syst. 1 (1910)
170, cum descr.
Triumfetta fabreana Gaudich. Voy. (1826) 478, t. 102.

COMIRAN ISLAND, Sulu Sea, Phil. Pl. 410 Merrill, distributed as “Trium-
fetta repens Forst.,”’ September, 1910, sandy seashore just above the limits
of high tides, extending inland only a short distance.

The specimens previously reported from the Philippines by me as this
species ‘ were later found to represent the allied but quite distinct Triumfetta
repens (Bl.) Merr. & Rolfe.*

Triumfetta procumbens Forst. is widely distributed in Polynesia, extend-
ing eastward to the islands off the north-east coast of Australia, Purdy
Island, north of New Guinea, to the small islands in the Indian Ocean off the
east coast of Africa, and the Keeling Islands. Triwmfetta repens (Blume)
Merr. & Rolfe, for which Gagnepain prefers the later name T. radicans
Bojer (1843), extends from Madagascar and the Seychelles to the Keeling
Islands, Java, Borneo, Indo-China, the islands in the Gulf of Siam, and to
the small islands off the north-east coast of Australia.

"Govt. Lab. Publ. (Philip.) 6 (1904) 17.
*This Journal 3 (1908) Bot. 111.

PHILIPPINE PLANTS, IX. 297

COLUMBIA Pers.

COLUMBIA MACGREGORII sp. nov.

Arbor, ramulis petiolis inflorescentiisque fulvo-ciliatis; foliis
oblongis vel ovato-oblongis, acuminatis, basi rotundatis vel ob-
tusis, subaequilateralibus, 5-nerviis, margine irregulariter den-
ticulatis; paniculis terminalibus, bracteolis trifidis, circiter 8
mm longis, dense ciliatis; sepalis extus dense stellato-pubescen-
tibus, petalis, basi stellato-puberulis exceptis, glabris.

A large tree, fide McGregor, the younger parts and the inflo-
rescence prominently fulvous-ciliate. Leaves chartaceous, oblong
to oblong-ovate, shining, somewhat brownish when dry, of nearly
the same color on both surfaces or the lower one a little paler,
14 to 20 cm long, 4.5 to 7.5 cm wide, the base rounded or obtuse,
subequilateral, the apex acuminate, the margins irregularly den-
ticulate, upper surface with few, scattered, ciliate hairs, the lower
one very minutely stellate-puberulent; basal nerves 5, the lateral
ones above the base about 7 on each side of the midrib, promi-
nent, ascending; petioles prominently fulvous-ciliate, 1.5 to 2
em long; stipules oblong, stellate-pubescent and ciliate, about 6
mm long. Panicles terminal, about 15 cm long, prominently
fulvous-ciliate with spreading hairs and also fulvous-stellate-
pubescent. Flowers numerous, yellowish with reddish spots,
about 1.5 cm in diameter. Sepals narrowly oblong or oblong-
lanceolate, about 8 mm long, 3 mm wide, narrowed to the acute
base and the acuminate apex, externally densely fulvous-stellate-
pubescent. Petals narrowly oblong-obovate or subspatulate, 6
to 7 mm long, 2.5 mm wide, the apex broadly rounded, narrowed
below, glabrous except the basal part, which is stellate pubescent,
and the slightly setose back, the basal gland about 1.5 mm in
diameter. Ovary subglobose, 2 mm in diameter, 5-sulcate, 5-
ridged, densely stellate-pubescent with fulvous hairs. Style 4
mm long, narrowed above and glabrous except near the base.
Bracteoles involucrate, 8 to 10 mm long, usually 3-cleft or
divided into three, lanceolate, acuminate lobes, their backs and
margins ciliate with 1 to 1.5 mm long fulvous hairs.

LUZON, Province of Nueva Vizcaya, Dupax, Bur. Sci. 11492 McGregor,
April 12, 1912.

A species distinguishable by its fulvous indumentum composed in part of
short stellate hairs and in part of long slender, ciliate ones. It belongs in
the group with the nearly equilateral leaves, allied to Columbia subaequalis

Planch., but differs from Planchon’s species, as well as from the numerous
forms described by Warburg, in the character of its indumentum.

298 MERRILL.

COLUMBIA MEGACARPA sp. nov.

Arbor (22 m alta fide Manalo) ; foliis oblongis, coriaceis, cir-
citer 20 cm longis, rectis vel leviter falcatis, basi valde oblique
cordatis, apice longe acuminatis, margine irregulariter repando-
dentatis, supra, nerviis exceptis, glabris vel subglabris, subtus
dense molliter cinereo-pilosis; fructibus dense pubescentibus, 2
ad 3 cm longis, 3-alatis.

A tree, 10 m high or more. Branches terete, rather stout,
brownish, strongly pubescent with long, spreading, yellowish-
brown hairs. Leaves alternate, oblong, coriaceous, about 20 cm
long, 5 to 8 cm wide, the apex rather long acuminate, the base
strongly and obliquely cordate, both lobes rounded, but one much
broader than the other, the margins irregularly repand-dentate,
the upper surface brown, shining when dry, glabrous or nearly
so except the somewhat pubescent nerves, the lower surface
softly and densely stellate-pilose with cinereous hairs; petioles
stout, densely pilose with spreading hairs; basal nerves reaching
to or above the middle of the leaf. Flowers unknown. Panicles
in fruit ample, terminal and in the upper axils, pilose with long,
soft, spreading, yellowish-brown hairs. Fruits oblong-ovoid in
outline, 3-winged, 2 to 3 cm long, nearly 2 cm wide, densely
subcinereous-pubescent or puberulent throughout, the portions
opposite the seed-bearing parts also densely pilose with long,
spreading hairs.

PALAWAN, about one mile north-east of Tanabag, For. Bur. 7416 Manalo,
December, 1906, in flat forests near sealevel, locally known as anilao.

A species manifestly allied to Columbia serratifolia (Cav.) DC., but
distinguished at once by its very much larger fruits. The fruits are
considerably larger than those of any other species of the genus known to
me, and apparently dehisce tardily.

Imperfect material, possibly a mixture, collected in Palawan by Mr.
Curran (For. Bur. 3587), may also be referable here.

GREWIA L.

GREWIA EDULIS sp. nov. § Omphacarpus.

Arbor parva, ramulis subtus foliis inflorescentiisque simpli-
citer pilosis; foliis breviter petiolatis, oblongo-ovatis, subcoria-
ceis, usque ad 18 cm longis, acuminatis, aequilateralibus, integris;
fructibus pyriformibus, carnosis, 2.5 ad 3 cm longis, puberulis.

A small tree, the branchlets, petioles, inflorescence, and lower
surface of the leaves more or less pubescent with simple, not
stellate, hairs. Branches terete, grayish-brown, glabrous, the
branchlets rather densely pubescent with short, brownish, spread-
ing hairs. Leaves oblong-ovate, equilateral, 15 to 18 em long, 5

PHILIPPINE PLANTS, IX. 299

to 7 cm wide, rather pale when dry, shining, the base rounded,
3-nerved, the apex shortly acuminate, the acumen blunt, the
margins entire, or very slightly undulate above, the upper surface
pubescent only on the midrib and primary nerves, the lower sur-
face pubescent also on the reticulations; basal nerves not extend-
ing above the middle of the leaf, the lateral ones above the basal
pair 6 or 7, anastomosing, prominent, the reticulations prominent,
lax; petioles very pubescent, stout, about 5 mm long. Inflores-
cence axillary, solitary, umbellate or subracemose, pubescent,
the peduncles about 1 cm long, the pedicels in fruit about the
same length. Flowers unknown. Fruit pyriform, obovoid,
fleshy, 2.5 to 3 em long, yellowish-green, the pericarp thin, slightly
puberulent, the mesocarp fleshy, containing numerous fibers, the
seed apparently solitary.

LUZON, Province of Nueva Ecija, Mataas na Cahoy, near Cabanatuan,
Bur. Sci. 5292 McGregor, September, 1908.

A species manifestly allied to Grewia stylocarpa Warb., but well dis-
tinguished by its pubescence, and by its differently shaped fruits. The

fruits are fleshy, yellowish-green when nearly mature, and have a pleasant
acid flavor.

GREWIA OVATA sp. nov. § Eugrewia.

Frutex scandens, omnibus partibus plus minusve stellato-
pubescentibus; foliis breviter petiolatis, ovatis, subcoriaceis,
breviter acuminatis, basi cordatis, margine crenato-denticulatis,
supra scabridis, plus minusve pubescentibus, subtus molliter stel-
lato-pubescentibus; inflorescentiis terminalibus axillaribusque,
umbellatis; floribus 2 cm longis; fructibus 4-lobatis, circiter 1.5
cm diametro, extus dense stellato-hirsutis, endocarpio osseo, 4-
lobato, lobis 2-locellatis.

A scandent shrub. Branches terete, brownish, and with the
branchlets densely covered with short, stellate, brownish-oliva-
ceous hairs. Leaves ovate, subcoriaceous, 7 to 12 cm long, 4
to 8 cm wide, the upper surface scabrid when dry, brownish,
distinctly stellate-pubescent, and on the nerves densely so, the
lower surface paler, rather densely and uniformly stellate-pubes-
cent, somewhat paler than the upper surface, the apex shortly
acuminate, the base broad, rounded, distinctly cordate, the mar-
gins finely and rather uniformly crenate-denticulate; basal nerves
reaching to or above the middle, the lateral ones above the basal
pair 5 or 6 on each side of the midrib, prominent; petioles densely
pubescent, less than 5 mm long; stipules lanceolate-acuminate,
pubescent, 2 mm long. Inflorescence axillary and terminal,
umbellate, densely stellate-pubescent with ferruginous hairs, the

300 MERRILL.

peduncles 1 to 1.5 cm long, each with 4 to 6 flowers, subtended
by a whorl of small, oblong-ovate to oblong-lanceolate, pubescent,
4 to 5 mm long bracteoles. Flowers yellow and white. Sepals
2 cm long, oblong-lanceolate, 3.5 mm wide at the base, somewhat
narrowed upward, very densely ferruginous-stellate-pubescent
outside, obtuse or subacute. Petals 5, thin, oblong, obtuse, 6
to 7 mm long, 2.5 mm wide, the basal gland large, prominent,
ciliate-pubescent, coriaceous, truncate, about 3 mm long and wide.
Stamens indefinite; filaments 6 to 11 mm long; anthers sub-
globose, 0.7 mm in diameter. Ovary densely villous with fer-
ruginous hairs; style glabrous, 10 to 12 mm long; stigma 1.5
mm wide. Fruit depressed, 4-lobed, the lobes rounded, densely
stellate-hirsute outside, about 1.5 cm in diameter, the endocarp
bony, each lobe 2-celled, each cell with a single seed.

Luzon, Province of Rizal, San Mateo, For. Bur. 1121 Ahern’s collector,
May, 1904, Decades Philip. Forest Flora 287 Ahern’s collector: Province of
Bataan, Lamao River, For. Bur. 1483 Ahern’s collector, July, 1904 (type).

A species manifestly allied to Grewia acuminata Juss. (G. umbellata
Roxb.), characterized by its very different and more dense pubescence, its
distinctly cordate leaves, etc.

GREWIA PALAWANENSIS sp. nov. § Hugrewia.

Frutex 2 ad 3 m altus, erectus vel subscandens; foliis char-
taceis vel subcoriaceis, oblongis, acuminatis, basi rotundatis,
margine denticulatis, usque ad 12 cm longis, utrinque, praesertim
ad nervos, plus minusve stellato-pubescentibus; inflorescentiis
axillaribus, umbellatis, umbellis solitariis vel fasciculatis, pauci-
floris; floribus circiter 5 mm longis; fructibus obovoideis, circiter
1 cm diametro, rugosis, parce hirsutis, 4-locellatis, vix lobatis.

An erect subscandent shrub 2 to 3 m high. Branches terete,
brownish or reddish-brown, lenticillate, ultimately glabrous, the
younger ones rather densely stellate-puberulent. Leaves char-
taceous or subcoriaceous, oblong, 6 to 12 cm long, 2 to 5 cm wide,
the base equilateral, rounded, the apex shortly acuminate, the
margins denticulate, the upper surface somewhat stellate-pu-
bescent on the nerves and midrib, the lower surface with short,
scattered, stellate hairs, more dense on the nerves; basal nerves
3, prominent, extending above the middle of the leaf, the lateral
ones on each side of the midrib about 4, prominent; stipules not
seen, apparently very early deciduous; petioles densely stellate-
pubescent, 5 to 10 mm long. Umbels axillary, solitary or more
usually fascicled, numerous, those in the uppermost axils some-
times simulating a terminal inflorescence, densely stellate-pube-

PHILIPPINE PLANTS, IX. 801

rulent, their peduncles 1 cm long or less. Flowers usually 3 in
each umbel, subtended by a whorl of very early deciduous, linear-
oblong, pubescent, 2 to 2.5 mm long bracteoles. Sepals 5,
stellate-puberulent with grayish hairs, obtuse or acute, 5 mm
long, 2 mm wide. Petals oblong, 2 mm long, obtuse, thin, gla-
brous except the large, basal gland which is 1 mm long and
prominently ciliate-pubescent. Stamens indefinite; filaments 2
to 2.5 mm long. Ovary densely villous. Style 2 mm long,
glabrous; stigma somewhat lobed, about 1 mm wide. Fruit de-
pressed-obovoid, about 1 cm in diameter, prominently wrinkled
when dry, brown, when immature prominently hirsute, when
mature with few, scattered, long hairs, not at all lobed, the
endocarp bony, 4-celled, each cell with a single seed.

PALAWAN, near Iwahig, For. Bur. 3522 Curran, January, 1906 (type),
Bur. Sci. 794, 864, 888 Foxworthy, April to May, 1906.

A species with much the aspect of Grewia acuminata Juss., and manifestly
allied to that species, differing especially in its very much smaller flowers.

GREWIA PARVA sp. nov. § Eugrewia.

Arbor parva circiter 4 m alta, ramulis subtus foliis inflores-
centiisque parce stellato-pubescentibus; foliis oblongis, acumi-
natis, 3.5 ad 4.5 cm longis, aequilateralibus, apice acuminatis, basi
acutis vel obtusis; inflorescentiis axillaribus, solitariis, umbel-
latis; fructibus subglabris, 2-lobatis, lobis 1- vel 2-locellatis,
circiter 5 mm longis.

A small tree or a shrub about 4 m high. Branches slender,
terete, brownish, prominently lenticellate, glabrous, the branch-
lets slightly pubescent with somewhat appressed, short, simple
or stellate hairs. Leaves oblong, chartaceous, 3.5 to 4.5 em long,
1.2 to 1.6 cm wide, somewhat brownish when dry, the upper
surface shining, slightly pubescent on the midrib and lateral
nerves, the lower surface slightly paler, rather uniformly pub-
escent with scattered, short, stellate hairs, the apex acuminate,
the base obtuse or acute, the margins distinctly and uniformly
crenate-serrate; base 3-nerved, the nerves not prominent, reach-
ing the middle of the leaf or slightly above, the lateral nerves
above the basal pair 3 or 4 on each side of the midrib, as-
cending, curved, slender; petioles pubescent, 2 to 3 mm long;
stipules not seen, apparently very early deciduous. Flowers
unknown. Infrutescence axillary, solitary, umbellate, somewhat
pubescent, the peduncles less than 1 cm long, each bearing from
1 to 3 fruits. Fruits black when dry, nearly glabrous, or with

302 MERRILL.

few, scattered, short hairs, 2-lobed, the lobes 5 mm long or less,
1- or 2-celled, each cell with a single seed.

LuzoN, Province of Ilocos Sur, San Quintin, For. Bur. 5627 Klemme,
October 18, 1906, on steep rocky slopes at an altitude of about 90 m. locally
known to the Ilocanos as duraring.

A species manifestly allied to Grewia laevigata Vahl, G. multiflora Juss.,
and related forms, characterized, however, by its small, pubescent leaves.

GREWIA ROLFEI sp. nov. § Hugrewia.
Grewia tiliaefolia Rolfe in Journ. Bot. 23 (1885) 211, non Vahl.

Arbor parva 10 ad 15 m alta, ramulis inflorescentiis foliisque '
utrinque, praesertim ad nervos, griseo-stellato-puberulis; foliis
chartaceis, oblongo-ovatis, acuminatis, basi rotundatis, leviter
inaequilateralibus, margine denticulatis; stipulis linearis, acu-
minatis; umbellis axillaribus, solitariis vel binis, 3-floris; floribus
parvis, circiter 7 mm longis.

A small tree 10 to 15 m high, sometimes less. Branches
slender, terete, glabrous, reddish-brown to nearly black when
dry, lenticellate, the branchlets rather densely stellate-puberulent
with grayish or sometimes brownish hairs. Leaves oblong-
ovate, chartaceous, 5 to 12 cm long, 3 to 5 cm wide, of nearly
the same color on both surfaces when dry, slightly shining, the
base rounded, usually somewhat inequilateral, the margins den-
ticulate except at the base which is entire, the apex rather
slenderly acuminate, the acumen 1 to 2 cm long, denticulate,
both surfaces rather densely gray-stellate-puberulent on the
midrib and lateral nerves, and with scattered, minute, stellate
hairs on the surface, the veins pale in contrast to the darker color
of the leaf; basal nerves reaching above the middle of the leaf, the
lateral ones above the basal pair 3 on each side of the midrib;
petioles 5 to 10 mm long, rather densely stellate-puberulent;
stipules linear, acuminate, pubescent, nearly 1 cm long. Umbels
3-flowered, numerous, axillary, solitary or paired, puberulent,
their peduncles 5 to 10 mm long, the pedicels shorter, subtended
by a whorl of three or four, oblong, obtuse or acute, 5 to 6 mm
long and 2 mm wide, pubescent bracteoles. Sepals narrowly
oblong or narrowly oblong-obovate, acute or obtuse, pubescent,
spreading or reflexed in anthesis, 7 mm long, 2 to 2.5 mm wide.
Petals 5, membranaceous, glabrous, narrowly oblong-obovate,
apex rounded or retuse, about 3 mm long, 1 mm wide, the basal
glands very minute or wanting. Stamens indefinite; filaments
3 to 4 mm long; anthers 0.25 mm in diameter. Ovary densely

PHILIPPINE PLANTS, IX. 303

villous; style glabrous, 3 mm long; stigma broad, somewhat
cleft. Very young fruit obovoid, densely pale-puberulent and
with longer pale hairs intermixed.

LuZzON, Province of Cagayan, Cuming 1319 (type): Province of Bataan,
Lamao River, For. Bur. 8048 Borden, May, 1905, For. Bur. 1489 Ahern’s
collector, July, 1904, Whitford 1288, May, 1905: Province of Pampanga,
Turo, For. Bur. 19290 Curran, March, 1910; Mount Arayat, For. Bur. 9616
Zschokke, October, 1907: Province of Tayabas, San Narciso, For. Bur.
14970 Darling, October, 1909.

This form has been referred by recent authors, following Rolfe, to the
- Asiatic Grewia tiliafolia Vahl. It does not appear to me to be referable to
Vahl’s species, and is accordingly here described as new. Hooker f. states
that Grewia tiliaefolia Vahl is best distinguished by its auricled, falcate
stipules, a character that does not at all apply to the specimens cited above.

GREWIA RIZALENSIS sp. nov. § Hugrewia.

Species G. rolfei simillima et valde affinis, differt foliis subtus
densissime ubique griseo-stellato-puberulis.

A small tree. Branches terete, slender, black when dry,
glabrous, slightly lenticellate, the branchlets densely and uni-
formly stellate-puberulent with cinereous or sometimes ferrugi-
nous hairs. Leaves ovate to oblong-ovate, chartaceous, 5 to 10
cm long, 2 to 4 cm wide, base rounded, slightly inequilateral, apex
acuminate, margins denticulate except in the basal portions
which are entire, the upper surface dark-colored when dry,
slightly stellate-puberulent, especially on the nerves, the lower
surface densely and uniformly covered with grayish, stellate,
very short hairs; petioles stellate-puberulent, 1 cm long or less;
stipules lanceolate, acuminate, pubescent, deciduous, about 5
mm long. Umbels numerous, axillary, solitary or fascicled,
similar to those of G. rolfei. Sepals lanceolate, acuminate, or
acute, pubescent, 7 mm long, 2 mm wide. Petals membrana-
ceous, glabrous, oblong, 2.5 to 3 mm long, 1 mm wide, obtuse or
retuse. Ovary densely villous; style glabrous, 3 mm long. Very
young fruits densely cinereous-pubescent.

LuZON, Province of Rizal, San Mateo, For. Bur. 1132 Ahern’s collector,
May, 1904 (type); Antipolo, For. Bur. 3139 Ahern’s collector, June, 1905,
Phil. Pl. 265 Ramos, June, 1910; Bosoboso, Merrill 2679, June, 1903, small
leaved form.

A species manifestly very closely allied to Grewia rolfei Merr., and per-
haps not specifically distinct, although differing in its uniform, dense, grayish
indumentum on the lower surfaces of the leaves, while G. rolfei is densely

puberulent on the nerves only, and with scattered hairs on the surface other-
wise, that do not conceal the epidermis of the leaf.

204 MERRILL.

STERCULIACEAE.
PTEROSPERMUM Schreb.

PTEROSPERMUM ELMERI sp. nov.

Arbor plus minusve stellato-pubescens; foliis obliquis, ovatis
vel oblongo-ovatis, subcoriaceis, 7 ad 10 cm longis, basi rotun-
datis, subpeltatis, apice breviter acute acuminatis, in siccitate
supra nigricantibus, glabrescentibus, subtus densissime fusco-
stellato-pubescentibus; floribus racemosis, bracteolis profunde
tripartitis, segmentis lineari-lanceolatis; sepalis circiter 3 cm
longis.

A tree, all parts more or less stellate-pubescent, the branchlets,
inflorescence, sepals, and lower surface of the leaves very densely
so. Branches reddish-brown, terete, ultimately glabrous, the
younger ones densely brown-stellate-pubescent. Leaves subco-
riaceous, entire, strongly oblique, ovate to oblong-ovate, 7 to 10
cm long, 3.5 to 4.5 cm. wide, the wider side of the leaf more than
twice wider than the narrower one, the base rounded, subpeltate,
the petiole inserted 2 to 4 mm from the margin, the apex shortly
and sharply acuminate, upper surface, when dry, black or nearly
so, somewhat covered with deciduous, brown, stellate hairs, ulti-
mately glabrous or nearly so, the lower surface very densely
brown-stellate-pubescent; lateral nerves 4 or 5 on each side of the
midrib; petioles densely pubescent, about 3 mm long. Racemes
terminal and in the upper axils, few-flowered, short; pedicels 2.5
cm long or less, each with several bracteoles which are cleft
nearly to the base into three linear, acuminate, 8 to 12 mm long
lobes. Sepals 3 cm long, about 3 mm wide, externally densely
stellate-pubescent with brown hairs, internally densely grayish-
pubescent. Petals about 3 cm long, 8 mm wide. Sterile sta-
mens 2 cm long, the fertile ones, including the linear 7 mm long
anthers, 2cminlength. Ovary densely brown-stellate-pubescent.

MINDANAO, District of Davao, Todaya (Mount Apo) Elmer 11928, Oc-
tober, 1909, distributed as Pterospermum obliquum Blanco.

A species manifestly very closely allied to P. subpeltatum Merr., differing
in its somewhat smaller, fewer nerved leaves and in its shorter sepals.

PTEROSPERMUM LONGIPES sp. nov.

Arbor circiter 10 m alta, ramulis subtus foliis inflorescentiisque
dense albido- vel subfusco-tomentosis; foliis oblongo-ovatis,
subcoriaceis, leviter obliquis, usque ad 8 cm longis, integris,
acuminatis, basi rotundatis vel subtruncatis vix cordatis, supra
glabris, subtus albido- vel subfusco-tomentosis; floribus panicu-
latis, long pedicellatis, sepalis circiter 5 cm longis.

PHILIPPINE PLANTS, IX. ~—68B05

A tree about 10 m high, more or less densely tomentose, the
branches terete, reddish-brown, the younger ones covered with
a somewhat deciduous, white or somewhat brownish tomentum.
Leaves oblong-ovate, often subrhomboidal, subcoriaceous, 5 to
8 cm long, 3 to 5 cm long, entire, somewhat inequilateral, the base
broad, rounded or subtruncate, not cordate or auriculate, the
apex shortly and sharply acuminate, the upper surface glabrous,
shining, the lower surface densely white-tomentose, or the
tomentum somewhat brownish; lateral nerves distinct, ascending,
3 or 4 on each side of the midrib; petioles 5 mm long or less,
tomentose. Flowers terminal and in the upper axils, forming a
distinct and rather many-flowered panicle, their pedicels, or
pedicels and the ultimate branchlets, 2 to 5 cm long, each with
several scattered bracteoles which are lanceolate, acuminate,
about 6 mm long, and somewhat sagittate at the base by a pair of
prominent, rounded auricles. Buds densely brown-stellate-
pubescent externally, cylindric, oblong. Sepals about 5.5 cm
long, 5 mm wide, acuminate, inside appressed pubescent with
grayish hairs. Petals thin, about 4cm long, 7mm wide. Sterile
stamens five, 3.5 to 4 cm long, the fertile ones 15, including the
8 mm long linear anthers, 2.5 cm in length. Ovary pubescent;
style 3.5 cm long. Immature capsules cylindric, glabrous, nar-
rowly oblong, about 4 cm long.

CAMIGUIN DE MINDANAO, Bur. Sci. 14632 Ramos, April 9, 1912, in forests,
Mount Mahinog, flowers yellow, fragrant.

A species manifestly closely allied to Pterospermum niveum Vid., which
it greatly resembles, differing in its somewhat smaller leaves which are
not at all cordate or auriculate at the base, but are rounded or subtruncate
and much less inequilateral than in Vidal’s species, and in its paniculate,
long-pedicelled flowers.

DILLENIACEAE.
DILLENIA L.

DILLENIA BOLSTERI sp. nov. § Wormia ?

Arbor circiter 10 m alta, foliis junioribus ramulis inflores-
centiisque plus minusve pilosis; foliis ellipticis vel oblongo-ellip-
ticis, acute acuminatis, basi acutis vel rotundatis, margine
serratis, uSsque ad 25 em longis, nervis utrinque circiter 17;
inflorescentiis in axillis superioribus terminalibusque, 2 ad 5-
floris; floribus albidis, circiter 6 cm diametro; stylis 8 vel 10.

A tree about 10 m high. Branches terete, reddish-brown,
glabrous, the branchlets appressed-pilose, rarely nearly glabrous.
Leaves alternate, elliptic to oblong-elliptic, subcoriaceous, 10 to
25 cm long, 4 to 10 cm wide, more or less pilose when young,

306 MERRILL.

becoming nearly glabrous, the apex sharply and rather abruptly
acuminate, the acumen about 1 cm long, the base acute or some-
what rounded; nerves about 17 on each side of the midrib, prom-
inent; petioles 2 to 3 cm long, when young somewhat pilose,
becoming glabrous, at first with the membranaceous stipules
adherent to and bordering the petioles, these soon deciduous.
Inflorescence more or less pilose, in the upper axils and terminal,
solitary, 2- to 5-flowered. Flowers white, about 6 cm in diameter.
Petals ovate, obtuse, about 1.5 cm long. Petals elliptic, obtuse,
3cm long. Stamens very numerous. Styles 8 or 10, spreading.
Fruit unknown.

MINDANAO, Province of Surigao, Surigao, Bolster 311, April, 1906 (type),
in forests, altitude about 65 m, Piper 224, April, 1911; Hinatuan, Piper 515,
April 1911.

This species was indicated in the herbarium as new in 1906, but was not
described on account of paucity of material. The additional specimens col-
lected by Mr. Piper show that the characters of the species are constant
and that it is apparently distinct. Its venation and shape of the leaves
distinguish it from other Philippine forms. Specimens from Leyte dis-
tributed by Mr. Elmer (No. 7149) under this name certainly do not represent
the species, but are probably referable to Dillenia philippinensis Rolfe.

SAURAUIA Willd.

SAURAUIA MACGREGORII sp. nov.

Arbor circiter 10 m alta, novellis setae 1.5 ad 3.5 mm longae
dense obtectis; foliis oblongis, coriaceis, usque ad 35 cm longis,
subtus parce setosis; inflorescentiis caulifloris, fasciculatis, vix
4 cm longis, setosis, paniculatis; floribus circiter 1.5 cm diametro,
ovario 5-loculare.

A tree about 10 m high, the young parts very densely covered
with brownish, slender, spreading, 1.5 to 3.5 mm long setae,
these setae less numerous on the petioles and midribs of the
leaves and on the older parts. Leaves oblong, coriaceous, shin-
ing, the lower surface considerably paler than the upper, 15
to 35 cm long, 5 to 10 cm wide, the apex shortly acuminate, the
base obtuse or rounded, the margins distantly glandular-denti-
culate, each small tooth bearing a long ciliate seta; lateral nerves
11 to 18, prominent on the lower surface, anastomosing, the
reticulations prominent, the midrib, nerves, and reticulations with
scattered setae; petioles 2.5 to 3.5 em long, setiferous. Inflores-
cence fascicled on the trunk, paniculate, less than 4 cm long,
rather densely brownish pubescent and with scattered short
setae, the bracts narrowly oblong, 4 to 5 mm long, thick. Flowers
white, 5-merous, about 1.5 cm in diameter. Outer two sepals
elliptic-ovate, rounded, concave, slightly setose externally, 5 to

PHILIPPINE PLANTS, IX. 307

6 mm long, thicker than the three somewhat petaloid inner ones
which are broadly ovate, rounded or retuse, somewhat longer
than outer two and with thin margins. Petals 8 mm long, about
6 mm wide, glabrous, elliptic-obovate, more or less retuse.
Stamens about 25; filaments stout, 3 mm long; anthers narrowly
oblong, as long as the filaments. Ovary depressed, glabrous,
somewhat 5-sulcate, 5-celled. Style 2 mm long, divided into 5,
slender, 4 mm long arms.

Luzon, Province of Nueva Vizcaya, Campote, Bur. Sci. 11407 McGregor,
March, 1912.

A species at once distinguishable by its dense indumentum on the younger
parts, composed of long, slender, spreading, setae-like hairs. In this char-
acter it differs from all the known Philippine forms. Another distinctive
character is its cauliflory, although this is by no means uncommon in the
genus.

GUTTIFORAE.
CALOPHYLLUM L.

CALOPHYLLUM KUNSTLERI King. in Journ. As. Soc. Beng. 59° (1890)
174; Vesque in DC. Monog. Phan. 8 (1893) 607.

LuzON, Province of Nueva Ecija, For. Bur. 22182 Alvarez, December,
1910: Province of Laguna, San Antonio, For. Bur. 13197 Curran, March,
1912.

Not previously reported from the Philippines; Malay Peninsula and
Borneo.

My conception of Calophyllum kunstleri King is based on the original de-
scription by King, and the later one by Vesque, together with a cotype
(King’s collector 5328) preserved in the herbarium of the Bureau of Science.
While the Philippine form shows some very slight differences, it appears
to me to agree with King’s species in all essential characters. A specimen
from Borneo, Sarawak, Foxworthy 302, appears also to be referable to the
same species.

GARCINIA L.

GARCINIA RAMOSII sp. nov. § Cambogia.

Arbor parva, ramis teretibus, ramulis tenuibus, obscure qua-
drangularis; foliis subcoriaceis, anguste oblongo-obovatis, obtusis
vel acutis, basi angustatis usque ad 6 cm longis, nervis obscuris,
utrinque circiter 10, subtus lineis longitudinalibus undulatis
tenuibus nigris vel purpureis notatis; floribus axillaribus, sessili-
bus, fasciculatis ad nodos incrassatis, 4-meris; staminibus 14;
ovario rudimento nullo.

A small tree, quite glabrous, the branches terete, dark-colored,
the branchlets obscurely 4-angled, slender, often olivaceous.
Leaves opposite, small, subcoriaceous, when dry rather pale,
shining and of the same color on both surfaces, 4 to 6 cm long,
1 to 2 cm wide, narrowly oblong-obovate or oblong, apex acute

1131114

308 MERRILL.

or obtuse, base narrowed, acute; lateral nerves very obscure,
about 10 on each side of the midrib, the lower surface marked
with slender, more or less undulate, not prominent, longitudinal,
purplish lines; petioles 2 to3 mm long. Flowers fascicled at the
thickened nodes, axillary, sessile, mostly in the axils of fallen
leaves, 4-merous. Male flowers: Sepals 4, orbicular, rounded, 2
mm in diameter. Petals oblong, in mature bud 3 to 3.5 mm long,
thick. Stamens 14, obscurely 2-seriate, covering the slightly
raised torus, oblong-obovate, about 1.2 mm long, the anthers
opening by introrse slits. Rudimentary ovary none.

LuZON, Province of Cagayan, Abulug River, Bur. Sci. 13919 Ramos, Jan-
uary, 1912, in forests on slopes.

A species well characterized by its small leaves with the faint, longitu-
dinal, purplish lines on the under surface.

BEGONIACEAE.
BEGONIA L.

BEGONIA AFFINIS sp. nov. § Petermannia.

Species B. agusanensis valde affinis differt ramulis dense longe
ciliato-setosis, capsulis majoribus, circiter 1.5 cm longis.

An erect, somewhat branched, suffrutescent herb 50 cm high
or less, the stems sparingly long-ciliate, the branchlets densely
covered with long, brown, ciliate hairs. Leaves glabrous, mem-
branaceous, inequilaterally oblong-obovate to oblong-oblanceolate,
rarely merely oblong, 9 to 15 cm long, 1.5 to 3.5 em wide, apex
slenderly acuminate, gradually narrowed below the middle to the
rather narrow, laterally cordate base, the margins irregularly
and rather coarsely toothed above the middle, the lower one-
half entire or subentire, the base usually less than 1 cm wide,
one side rounded, the narrower side acute; petioles prominently
ciliate, about 3 mm long; stipules oblong-lanceolate, with a long
and slender, ciliate, flagellate acumen. Panicles terminal, gla-
brous, 4 to 7 cm long, the peduncles slender, the branches
dichotomous, few, slender; bracts lanceolate, acuminate, 5 to
7 mm long, the bracteoles similar, smaller. Flowers pink. Male
flowers: Pedicels 4 to 5 mm long. Sepals 2, elliptic, obtuse,
about 6 mm long, 3 to 3.5 mm wide, 6-nerved. Petals 2, oblong,
obtuse, 4 to 5 mm long, 1.5 to 2 mm wide. Stamens about 25;
anthers oblong-obovoid, retuse, 1.5 mm long, the filaments
shorter. Female flowers solitary at the base of the male panicle,
their pedicels slender, 1 to 1.5 em long. Perianth segments 5,
the outer two oblong-elliptic, somewhat narrowed to the obtuse
apex, about 12 mm long, 5 to 5.5 mm wide, slenderly 9-nerved,

PHILIPPINE PLANTS, IX. 309

the inner three as long as the outer ones, about 4 mm wide,
slenderly 5-nerved. Styles 3, forked, 4 mm long; stigmas spirally
arranged. Capsules about 1.5 cm long, 1.5 to 2 cm wide, equally
3-winged, somewhat depressed-rhomboid in outline, the wings
subequally rounded at their tips.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8251 (type), 8248, November 27, 1911, in damp ravines,
altitude about 900 m.

A species manifestly closely allied to Begonia agusanensis Merr., dis-
tinguished chiefly by its larger capsules and its densely ciliate branchlets.

BEGONIA ELATOSTEMATOIDES sp. nov. § Petermannia.

Species B. ciliiferae affinis, differt foliis minoribus, integris vel
subintegris.

An erect or ascending, simple or slightly branched suffrutes-
cent herb, the stems often prostrate below, 20 to 40 cm high,
the stems, petioles, and lower surfaces of the leaves, at least on
the veins, prominently ciliate with brown hairs. Leaves green
above, red beneath when fresh, membranaceous, oblong to nar-
rowly oblong-obovate, often subequally narrowed at both ends,
3.5 to 7 cm long, 1 to 2.5 cm wide, slightly acuminate, entire,
base slightly inequilateral and slightly cordate, the upper surface
with very few, short, scattered, brown hairs or nearly glabrous,
the lower surface prominently ciliate with brown hairs on the
nerves, and with scattered shorter hairs on the surface; nerves
pinnately arranged, 3 or 4 on each side of the midrib, ascending ;
stipules lanceolate, acuminate, brown-ciliate, about 1 mm long;
petioles 1 to 3 mm long. Flowers pink and white, in axillary
fascicles, but one or two opening at a time. Male flowers: Ped-
icels slender, 4 to 5 mm long, subtended by several, lanceolate,
long-acuminate, brown-ciliate bracts about 2 mm wide, 5 to 7
mm long, their margins prominently fimbriate-ciliate. Sepals 2,
broadly ovate, acute, 8 to 9 mm long, 5 to 6 mm wide, slenderly
7-nerved, outside with few, scattered, short hairs. Petals 2,
oblong, obtuse, about 6 mm long, 2 mm wide. Stamens about
30, the torus somewhat elongated; filaments about 0.5 mm long;
anthers oblong-obovate, retuse, about 1 mm long. Capsules
broadly subrhomboid-obovate, truncate, about 8 mm long, 1 cm
wide, equally 3-winged, the outer upper angles of the wings
subacute, base narrowed and acute.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8232, November 27, 1911, in damp shaded ravines, and on
forested slopes, altitude 800 to 1000 m.

A species manifestly closely allied to Begonia ciliifera Merr., a species
also known only from the Zamboanga Peninsula. The present species

310 MERRILL.

differs especially in its smaller size, and in its smaller entire leaves which
are nearly glabrous on the upper surface, not prominently ciliate with
long hairs. In habit and general appearance it approximates some species
of Elatostema, from which the specific name.

BEGONIA OBLONGATA sp. nov. § Petermannia.

Species B. contractae Warb. affinis, differt foliis angustioribus,
valde oblique cordatis, capsulis majoribus, circiter 15 mm longis.

An erect, glabrous, sparingly branched, succulent herb up to
60 cm in height, the stems often somewhat prostrate below, the
nodes sometimes emitting rootlets. Leaves membranaceous,
green, oblong to oblong-lanceolate, 8 to 12 cm long, 1.2 to 3 cm
wide, often somewhat falcate, gradually narrowed above and
slenderly acuminate, the base very strongly inequilateral, cordate,
the sinus lateral, the lower lobe very much larger than the upper
one, usually about 2 cm wide, rounded-angular, the upper lobe
acute or slightly rounded, about 5 mm wide, the sinus narrow,
acute, the lobes not overlapping; basal nerves 1 or 2 in the narrow
lobe, 4 or 5 in the broader one, radiate, the lateral ones above
the base 2 to 4, ascending, anastomosing, the margins rather
prominently but irregularly toothed; petioles 1.5 to 3 cm long;
stipules membranaceous, oblong-lanceolate, acuminate, about 1
em long. Inflorescence terminal, slender, 6 cm long or less,
divaricately branched, few-flowered, the basal flower female,
the upper ones male. Male flowers: Sepals 2, suborbicular, pink,
rounded, about 4 mm in diameter, slenderly 7-nerved. Petals
none. Stamens 30 to 35; anthers obovoid, slightly retuse, 0.8 to
1 mm long, the filaments less than 1 mm in length. Female
flowers not seen. Capsules ovoid, truncate, about 15 mm long,
15 to 17 mm wide, equally 3-winged, the outer angles of the wings
subacute, the base broadly rounded.

MINDANAO, District of Zamboanga, mountains back of Zamboanga, Mer-
rill 8166 (type) 8175, December 6, 1911, in damp shaded ravines, altitude
100 m and above.

A species manifestly allied to Begonia contracta Warb., but differing in
a number of characters, especially in its relatively narrow leaves and its
much larger capsules.

BEGONIA MACGREGORII sp. nov. § Petermannia.

Species ut videtur B. cumingii A. Gray affinis, differt floribus
masculinis multo minoribus, partibus junioribus ramisque longe
ciliato-setosis.

An erect or ascending sparingly branched plant 20 to 100 cm
high, the stems with numerous, long, slender, brownish, ciliate-

PHILIPPINE PLANTS, IX. 311

setose, spreading hairs 5 mm long or less. Leaves oblong, when
fresh dark-green on the upper surface and often blotched with
gray, the lower surface deep red, chartaceous or membranaceous,
15 to 20 cm long, 6 to 7 cm wide, not narrowed at the base but
very strongly obliquely cordate, the broad lobe rounded, the
narrow one subacute or obtuse, the apex rather slenderly acumi-
nate, margins shallowly, distantly, and irregularly sinuate-toothed
or lobed, ciliate-denticulate, the upper surface glabrous, the
lower one brownish-pubescent and ciliate-setose on the nerves;
basal nerves radiate, about 8, prominent; petioles prominently
ciliate-setose with long hairs, 1 to 3 cm long; inflorescence ax-
illary, few-flowered, 4 cm long or less, sparingly branched. Male
flowers: Sepals 2, ovate, obtuse, about 8 mm long. Petals none.
Stamens about 30; anthers narrowly oblong-obovate, obtuse, 2
mm long, the filaments very short or none. Bracteoles lanceolate,
acuminate, 3 mm long, deciduous. Female flowers on separate
plants. Sepals 5, thin, reticulate, broadly lanceolate to ovate-
lanceolate, acuminate, at anthesis about 8 mm long, 4 to 5 mm
wide, accrescent and up to 13 mm long before falling. Styles
2 mm long; stigmas forked, spirally twisted, about 2 mm long.
Mature capsules 2 to 2.5 cm long, about 2 cm wide, truncate at
the apex, the outer upper angles acute, subequally 3-winged, one
wing rounded and broader at its base than are the other two
which are narrowed and acute at the base.

LuZON, Province of Nueva Vizcaya, Dupax, Bur. Sci. 113834 McGregor,
March, 1912, in forests along small streams.

This distinct species is probably most closely allied to Begonia cumingti
A. Gray, from which it is at once distinguished by its long and slender
ciliate-setose hairs and by its small male flowers. It somewhat resembles

Begonia crispipila Elmer, but lacks the prominently bracteate inflorescence
of that species, and has very different capsules.

BEGONIA QUERCIFOLIA A. DC. in Ann. Sci. Nat. IV 11 (1859) 129,
Prodr. 15* (1864) 320; F.-Vill. Novis. App. (1880) 99; Vid. Phan.
Cuming. Philip. (1885) 116, Rev. Pl. Vasc. Filip. (1886) 148.

Begonia leytensis Elm. Leafl. Philip. Bot. 2 (1910) 739; Merr. in
Philip. Journ. Sci. 6 (1911) Bot. 384.

This species was wrongly interpreted by me in my recent paper of Philip-
pine Begonia’*, for all the recently collected material I rather tentatively
referred to DeCandolle’s species, represents a quite different form. After
a careful study of the available material and the descriptions I am confident
that Begonia leytensis Elm. is an exact synonym of B. quercifolia A. DC.,
and it is accordingly here reduced. So far as known B. quercifolia A. DC.

* This Journal 6 (1911) Bot. 387.

312 MERRILL.

is represented only by the type collection from Samar, Cuming 1696, of
which I have examined a specimen preserved in the Kew Herbarium, and of
which I have a carbon rubbing representing the leaf-outline and venation,
and Elmer 7255 from the neighboring island of Leyte, type collection of
Begonia leytensis Elm. The material I referred to B. quercifolia may prove
to be only a luxuriant form of B. contracta, but it seems advisable to wait
for more material before definitely deciding this point.

THYMELAEACEAE.
MACGREGORIANTHUS gen. nov.

Flores hermaphroditi. Perianthii tubus elongatus, cylindricus,
lobi 5, patentes, fauce intus nuda. Stamina 10, tubo 2-seriatim
inclusa, filamentis brevissimis; antherae oblongae, parvae.
Discus annularis, brevissimis, obscurissimis, lobatis. Ovarium
sessile, villosissimum, 1-loculare, in stylum desinens, stigmate
obovoideo, papilloso. Fructus ignotus. Frutex ut videtur scan-
dens. Folia opposita, plana, margine cartilaginacea. Flores in
paniculis axillaribus terminalibusque laxis elongatis dispositis.
Perianthia extus puberula.

MACGREGORIANTHUS PANICULATUS sp. nov.

Frutex inflorescentiis exceptis glaber; foliis coriaceis, ovato-
ellipticis ad oblongo-ellipticis, usque ad 9 cm longis, obtusis vel
obscure brevissime late acuminatis, nervis utrinque circiter 13;
paniculis elongatis, paucifloris, griseo-puberulis, 12 at 25 cm
longis; floribus ad apices ramulorum fasciculatis, circiter 8 mm
longis.

A shrub, climbing (fide McGregor), glabrous except the inflo-
rescence. Branches reddish-brown, terete, slender, the younger
ones often somewhat puberulent, soon becoming quite glabrous,
wrinkled when dry. Leaves opposite, coriaceous, olivaceous and
slightly shining when dry, the lower surface somewhat paler
than the upper, ovate-elliptic to oblong-elliptic, 6 to 9 cm long,
3 to 5.5 em wide, base rounded or subacute, apex obtuse or very
broadly, shortly, and bluntly acuminate, the margins entire,
cartilaginous; nerves about 13 on each side of the midrib, spread-
ing-ascending, anastomosing directly with the cartilaginous leaf-
margin, the reticulations obscure; petioles reddish-brown when
dry, about 1 cm long. Panicles terminal and in the upper axils,
elongated, lax, few-flowered, all parts densely and uniformly gray-
puberulent, and with scattered, reduced, linear bracts 5 mm long
or less. Flowers pale-green, 5-merous, 9 mm long, fascicled at the
ends of the ultimate branchlets, their pedicels 5 mm long or less.

PHILIPPINE PLANTS, IX. o13

Perianth-tube cylindric, about 1.5 mm in diameter, inside gla-
brous, outside gray-puberulent, the lobes 5, elliptic, spreading,
2.5 to 3 mm long, rounded, imbricate, densely gray-pubescent on
the inner surface. Stamens 10, alternate in two series, inserted
just at the apex of the perianth-tube but included, the anthers
oblong, about 1 mm long; filaments of the upper row about 1 mm
long, those of the lower row shorter. Ovary sessile, densely
villous, oblong-ovoid, about 3 mm long, 1-celled, narrowed above;
style slender, glabrous, about 1.5 mm long; stigma obovoid, pap-
illose, about 0.5 mm long. Fruit unknown.

Luzon, Province of Laguna, Calauan, Bur. Sci. 12360 McGregor, Decem-
ber 12, 1910, a vine in hill-side forests.

The genus here proposed is dedicated to Mr. R. C. McGregor of the
Bureau of Science by whom it was collected, and who has made extensive
botanical collections in the Philippines and in Guam during the past eight

ears.

" Macgregorianthus is manifestly allied to Wikstroemia Endl., differing
especially in its 5-merous flowers, as well as in its habit. All known species
of Wikstroemia have a 4-parted perianth, and 8 stamens. Mr. McGregor
notes that the plant is a vine which is presumably correct, although this
is a most unusual character in the Thymelaeaceae. The cartilaginous leaf-
margin, with which the lateral nerves anastomose direct, is a striking
character of the proposed genus.

GYRINOPSIS Decne.

GYRINOPSIS BRACHYANTHA sp. nov.

Arbor parva, glabra; foliis chartaceis vel subcoriaceis, nitidis,
oblongis, usque ad 16 cm longis, acuminatis, basi acutis vel
obtusis; floribus axillaribus, solitariis vel fasciculatis, 4 mm
longis, perianthio campanulato, capsulis anguste obovoideis, levi-
ter compressis, 1.2 ad 1.5 cm longis.

A small tree, or according to Ramos a shrub 2 m high, glabrous,
the branches slender, terete, brownish when dry, the growing
tips usually pubescent. Leaves oblong, chartaceous or subcor-
iaceous, oblong, 8 to 16 cm long, 2 to 4.5 cm wide, shining on both
surfaces, the base acute or obtuse, the apex acuminate, margins
recurved; midrib very prominent, the nerves exceedingly nu-
merous, spreading, parallel, very dense, slender; petioles about
5 mm long. Flowers greenish, axillary, solitary or several in a
fascicle on a short axis, the pedicels about 3 mm long. Perianth
campanulate, 4 mm long, in bud distinctly pubescent externally,
becoming nearly glabrous, the lobes 5, oblong, or ovate-oblong, -
pubescent internally, as long as the perianth-tube. Stamens 10,
five opposite the perianth-lobes, their anthers about 0.7 mm long,

314 MERRILL.

with longer filaments than the five alternate with the lobes, all
inserted with the scales, not below them, the scales 10, oblong,
ciliate, alternating with the filaments, about 1 mm long. Ovary
ovoid, 2-celled, slightly pubescent; style 1 mm long. Capsule
narrowly obovoid, 2-valved, more or less compressed, 1.2 to 1.5
long, the seed pubescent, including the projecting ends, about 1
cm long.

LUZON, Province of Cagayan, Abulug River, Bur. Sci. 13862 Ramos
(type), For. Bur. 19562 Curran, January, 1912; Pamplona, Bur. Sci. 7489
Ramos, March, 1909, distributed as G. cumingiana Decne.

A species in vegetative characters closely resembling Gyrinopsis cuming-
tana Decne., but with entirely different flowers. In floral characters it
approaches the genus Aquilaria, and appears to be more or less inter-
mediate between Gyrinopsis and Aquilaria. In consideration of its axillary
inflorescence, and in its having 10 scales at the throat instead of five, I
have referred it to the former genus. In structure the flower appears to
differ a little from both genera cited above in that the stamens are inserted
with and alternate with the scales, not below them.

MYRTACE.
EUGENIA L.

EUGENIA CAMIGUINENSIS sp. nov. § Syzygium.

Species E. densinerviae Merr. simillima et valde affinis, differt
foliis minoribus, floribus brevioribus.

A glabrous tree about 8 m high, the branches terete, light-gray,
the ultimate ones 3 to 4 mm thick. Leaves elliptic to elliptic-
obovate, thickly coriaceous, 8 to 11 cm long, 5 to 7 cm wide, some-
what shining when dry, the lower surface glandular-punctate,
a little paler than the upper one, the apex broad, rounded or very
broadly and obtusely acuminate, the base acute; nerves dense,
slender, indistinct, parallel, 25 to 30 on each side of the midrib,
anastomosing into a very obscure submarginal nerve; petioles
stout, 5 to 7 mm long. Inflorescence terminal, corymbose, many
flowered, peduncled, or branched from the base, the branches
trichotomous, the whole inflorescence up to 8 cm long, 7 to 13
cm in diameter. Ultimate branchlets short, densely arranged,
the flowers in threes, subsessile, each subtended by a pair
of ovate, concave, stiff, rounded bracteoles 1.5 mm long, the
bracts similar, a little larger. Buds 6 to 7 mm long, narrowly
obovoid to broadly club-shaped. Calyx about 5 mm long, 3 mm
wide at the apex, obscurely 4-toothed. Petals calyptrate.

CAMIGUIN DE MINDANAO, in forests near the summit of Mount Mahinog,
Phil. Pl. 1175 Ramos, April, 1912.

PHILIPPINE PLANTS, IX. 315

EUGENIA CILIATO-SETOSA sp. nov. § Syzygium ?

Arbor parva 2 ad 5 m alta, ramulis foliis subtus ad nervos
inflorescentiisque insigniter ciliato-setosis; foliis oblongo-lanceo-
latis, usque ad 20 cm longis, acuminatis, basi cordatis, brevissime
petiolatis, nervis lateralibus marginalibusque subtus prominen-
tibus; inflorescentiis terminalibus, paniculatis, ut videtur pauci-
floris, fructibus inaequilateraliter obovoideis, 1.5 ad 2 cm longis.

A small tree or a shrub 2 to 5 m high, the branches, nerves
on the lower surfaces of the leaves, especially the midribs, and
the inflorescence prominently ciliate-setose. Branches terete,
grayish, slender. Leaves opposite, oblong-lanceolate, 14 to 20
cm long, 3 to 6 cm wide, chartaceous, the upper surface glabrous,
olivaceous, shining, the lower paler, the midrib prominently, the
lateral nerves sparingly ciliate-setose, apex sharply and slenderly
acuminate, the base rather abruptly rounded-cordate; nerves
very prominent on the lower surface, 20 to 25 on each side of the
midrib, anastomosing in nearly straight marginal nerves as
prominent as the lateral ones; petioles ciliate-setose, about 2 mm
long. Inflorescence terminal, ciliate-setose, paniculate, 10 to
15 cm long including the elongated peduncle, the branches few,
spreading, apparently few-flowered. Immature buds obovoid,
about 4 mm long. Fruits inequilaterally obovoid, 1.5 to 2 em
long, nearly smooth, green when fresh.

Luzon, Province of Cagayan, Abulug River, Bur. Sci. 18974 Ramos
(type), with immature buds and fruits, For. Bur. 11590, 13974 Curran,
with fruits, all collected in January, 1912, in dipterocarp forests 30 to 50
meters above sea-level.

A most striking species, differing from all the Philippine forms, and
from most or quite all of the previously described species in the genus by
its prominently ciliate-setose branches, leaves, and inflorescence.

EUGENIA PROPINQUA sp. nov. § Syzygium.

Species E. pauciveniae C. B. Rob., simillima et affinis, differt
foliis minoribus, distincte petiolatis, floribus ut videtur mino-
ribus.

A tree (fide Ramos), 10 m high, glabrous, the branches terete,
somewhat grayish and reddish-brown, the young branchlets
slightly compressed, slender. Leaves numerous, crowded, coria-
ceous, shining on both surfaces, oblong-elliptic to obovate, the
base acute, the apex rounded, usually broadly so, 3 to 5 cm long,
1.5 to 38 cm wide; nerves about 10 on each side of the midrib,
indistinct, irregular, very obscurely anastomosing; petioles 3 to
4 mm long. Inflorescence terminal, many-flowered, 3 to 4 cm

316 MERRILL.

long, 5 to 6 em wide, corymbose, 3-branched from the base, the
branches trichotomous, the ultimate branchlets bearing from 3
to 5 sessile flowers, the nearly mature buds oblong-ellipsoid or
narrowly obovoid-ellipsoid, about 5 mm long.

Luzon, Province of Cagayan, Abulug River, Bur. Sci. 13911 Ramos,
January, 1912, in forests.

A species manifestly very closely allied to Hugenia paucivenia C. B. Rob.,
differing in its smaller, differently shaped, distinctly petioled leaves which
are acute at the base, its more densely flowered inflorescence, and its
apparently somewhat smaller flowers.

EUGENIA TENUIPES sp. nov. § Jambosa.

Species E. bataanensis simillima et valde affinis, differt pedi-
cellis tenuibus, valde elongatis, usque ad 12 cm longis.

A small tree, quite glabrous, the branches terete, grayish or
reddish, slender, the ultimate branchlets 1 to 1.5 mm in diameter.
Leaves opposite, shortly petioled, chartaceous, oblong-ovate to
ovate-lanceolate, 5 to 12 cm long, 1.5 to 3.5 em wide, rather pale
and somewhat shining when dry, the lower surface paler than
the upper, narrowed to the acute or rarely somewhat rounded
base, the apex slenderly and sharply acuminate, the acumen
usually about 1 cm long; nerves irregular, distant, about 6 on
each side of the midrib, anastomosing and forming an undulate
submarginal nerve about as prominent as the primary lateral
ones; petioles about 2 mm long. Flowers axillary and terminal,
solitary, white, their pedicels very slender, elongated, 4 to 12 cm
long, with 2 minute bracteoles in the lower one-fifth or one-
sixth. Calyx jointed to the pedicel, funnel-shaped in anthesis,
about 2 cm long, narrowed below to the slender stalk, about 1.8
em wide, the lobes distinct, rounded, in fruit becoming ovoid
and about 2 cm long and wide. Petals free. Stamens very
numerous.

LuZON, Province of Cagayan, Abulug River, For. Bur. 19608 Curran
(type), January, 1912, Bur. Sci. 18963 Ramos. I refer here also Bur. Sci.
7867 Ramos, from Claveria, Cagayan Province, previously considered by
Doctor Robinson in his treatment of the Philippine Myrtaceae” as Eugenia
bataanensis Merr. Doctor Robinson notes that this Cagayan specimen differs
from the typical form of Eugenia bataanensis Merr., in its leaves being
acute at the base. Additional material shows that the peculiar character

of the inflorescence, the very long and slender pedicel, is constant, and
hence the form is described as new. Unquestionably closely allied to Euge-

“This Journal 4 (1909) Bot. 348.

PHILIPPINE PLANTS, IX. aL

nia bataanensis Merr., differing especially in its leaves being usually acute
at the base, and in its flowers always being long and slenderly pedicelled.

TRISTANIA R. Br.

TRISTANIA LITTORALIS sp. nov. § Hutristania.

Species T. obovatae R. Br. similis et affinis, differt foliis brevis-
Sime petiolatis vel subsessilis, basi distincte auriculatis, petalis
integris, vix denticulatis.

A tree with hard wood, quite glabrous except the puberulent
inflorescence. Branches terete, light-gray, slender, the branch-
lets reddish-brown as are the panicles and midribs of the leaves
when dry. Leaves alternate, coriaceous, narrowly obovate-oblong
to obovate-elliptic, 9 to 22 cm long, 4 to 10 em wide, shining when
dry, the lower surface distinctly paler than the upper one, the
apex obtuse, acute, or distinctly and shortly acuminate, narrowed
below to the distinctly auriculate base, the margins recurved;
lateral nerves slender, 8 to 10 on the smaller leaves, up to 30 on
the larger ones, usually 5 to 8 mm apart, spreading, straight or
nearly so, anastomosing and forming a continuous, slender, sub-
marginal nerve; petioles stout, 2 to 3 mm long, wider than long,
or wanting and the leaves sessile. Inflorescence axillary, pe-
duncled, cymose, somewhat puberulent, 2 to 9 cm long. Flowers
sessile or nearly so, the calyx funnel-shaped, about 3 mm long,
3 mm wide at the throat, with 5 broadly triangular teeth which
are acute or obtuse. , Petals 5, glabrous, rounded, obovate, 1.2
mm long. Stamens 15, in five groups of three each opposite
the petals, the middle filament of each group 1 to 1.2 mm long,
the two lateral ones a little shorter, the anthers very small.
Ovary globose, slightly gray-pubescent; style slender, 1 mm long.
Capsules obovoid-ellipsoid, 4 mm long, the three valves coria-
ceous, slightly pubescent, ultimately glabrous or nearly so. - Seeds
thin, flat, narrowly obovoid, including the wings about 4 mm
long.

MINDANAO, District of Zamboanga, Taligbao River, For. Bur. 13534 Foa-
worthy, De Mesa, & Villamil, June 17, 1912, common here, occurring just
back of the mangrove on dry land, and on a low ridge running back from
the edge of the swamp. The same species is represented by a sterile
specimen, For. Bur. 11246 Hutchinson, from Santa Maria, Zamboanga. It
is known to the Moros as taba.

The wood is hard and is said to be very durable. The species is the
second one of the genus to be recorded from the Archipelago, but appa-

rently one or two additional distinct forms are represented by sterile ma-
terial from different parts of the Philippines.

318 MERRILL.

ARALIACEAE.
SCHEFFLERA Forst.

SCHEFFLERA STELLULATA sp. nov. § Heptapleurum.

Arbor parva, inflorescentiis stellato-furfuraceis exceptis gla-
bra; foliis longe petiolatis, palmatim 6- ad 9-foliolatis, foliolis
subcoriaceis, oblongis, brevissime abrupte acuminatis, usque ad
25 em longis; inflorescentiis terminalibus, ramis paucis, elongatis,
racemosis; floribus 5-meris, capitato-umbellatis, umbellis pedun-
culatis, racemosis.

A small tree (fide McGregor), glabrous except the inflorescence
which is minutely stellate-furfuraceous throughout with a some-
what brownish or grayish indumentum. Branches terete, gray,
marked with the annular scars of fallen petioles. Leaves pal-
mately 6- to 9-foliolate, their petioles about 28 cm long. Leaflets
oblong, subcoriaceous, somewhat shining, of about the same color
on both surfaces when dry, 15 to 25 cm long, 6 to 7 cm wide,
the apex very shortly and abruptly acuminate or merely acute,
the base usually rounded, sometimes subacute; primary nerves
10 to 12 on each side of the midrib, rather distinct, anastomos-
ing; petioles 5 to 8 cm long. Inflorescence terminal, paniculate,
peduncled, the peduncle and rachis about 30 cm long, each
panicle with about 6, slender, elongated, racemosely arranged
primary branches 25 to 35 cm in length, all parts stellate-fur-
furaceous, each primary branch with about 30 peduncled, race-
mosely arranged subcapitate umbels, the umbels with from 15
to 20 flowers, the lower umbels with peduncles 10 to 12 mm in
length, the upper ones gradually shorter, the umbels 1 cm in
diameter or less. Flowers greenish-yellow, 5-merous, their
pedicels 1 to 1.5 mm long, in fruit up to 4 mm long. Calyx
funnel-shaped, subtruncate, about 2 mm in diameter. Petals 5,
ovate, free, acute or obtuse, 2 mm long, marked with small
brownish-yellow dots and dashes. Stamens 5; filaments slender,
3 to 4 mm long; anthers about 1 mm long. Ovary 5-celled.
Fruit turbinate, 2.5 mm in diameter, the apex convex, grayish,
rugose.

Luzon, Province of Nueva Vizcaya, Dupax, Bur. Sci. 11251 McGregor,
April 15, 1912.

A characteristic species, distinguishable by its stellate-furfuraceous in-
florescence, its capitate-umbellate flowers, the umbels being racemosely dis-
posed on the elongated branches of the inflorescence, and by its oblong
leaflets.

PHILIPPINE PLANTS, IX. 319

ALANGIACEAE.
ALANGIUM Lam.

ALANGIUM BRACHYANTHUM sp. nov. § Eualangium.

Arbor circiter 7 m alta; floribus in ramis defoliatis, axillaribus,
fasciculatis, brevissime pedicellatis, petalis 6, 1.5 cm longis,
pubescentibus; staminibus 15, basi vix curvato-geniculatis.

A tree about 7 m high. Branches and branchlets light-gray
or somewhat reddish, slender, glabrous. Flowers yellowish-
white, very fragrant, numerous, appearing before the leaves,
axillary, fasciculate, several at each node, the fascicles rather
congested; pedicels very short, pubescent. Calyx 2 mm long,
3 mm in diameter, pubescent, obscurely 6-toothed. Petals 6, 1.5
em long, about 1 mm wide, appressed-pubescent externally.
Stamens 15; filaments slender, 7 to 8 mm long, not inflexed or
geniculate at the base, somewhat villous at about the middle;
anthers 6 to 7 mm long; style as long as the petals; stigma
globose.

LUZON, Province of Tayabas, Pagsalam, For. Bur. 10341 Cur-
ran, April 9, 1908, in beach forests, altitude about 5 m, locally
known as malatapay.

I have refrained from describing the leaves of this apparently very dis-
tinct species, as Mr. Curran notes that they were picked up on the ground
from under the tree, there being no leaves on the tree at the time it was
found in flower. The species is manifestly allied to Alangium salviifolium
(L. f.) Wangerin, but differs remarkably in its much smaller flowers and in
its fewer stamens. The detached leaves with the specimen, which however

may belong to a different plant, are penninerved, not at all 3-nerved from
the base.

ALANGIUM LONGIFLORUM sp. nov. § Hualangiwm.

Species A. salviifolio (L.) Wangerin affinis, differt floribus
5-meris, multo longioribus, staminibus paucioribus, 16 vel 17,
petalis usque ad 4.5 cm longis.

A deciduous tree 7 to 10 m high. Branches terete, grayish-
brown, glabrous, the ultimate ones softly pubescent with short,
pale-brown or olivaceous hairs. Leaves appearing after the
flowers, oblong to obovate-oblong, membranaceous or chartaceous,
ultimately glabrous on both surfaces, when young: slightly
pubescent on the nerves beneath, 10 to 15 cm long, 4 to 7 cm
wide, entire, equilateral, the base rather abruptly rounded or
obtuse, distinctly 3-nerved, the apex prominently acuminate;
lateral nerves above the basal pair 4 or 5 on each side of the

320 MERRILL.

midrib, as prominent as the basal ones, somewhat ascending,
dnastomosing, the reticulations distinct; petioles 5 to 8 mm long,
pubescent when young, ultimately glabrous. Flowers on defol-
iated branches, axillary, several in each axil, white, the pedicels
densely pubescent, stout, 3 to 4 mm long. Calyx obconic, about
5 mm long, 4 mm in diameter, the limb produced 1.5 to 2 mm
above the ovary, very broadly and obscurely 5-toothed. Petals
5 (rarely 4), on the type 4.5 cm long (on other specimens referred
here 3.5 to 4 cm in length), 2 mm wide below, free, narrowed
above, cinereous-pubescent. Stamens 16 or 17, as long as the
petals; filaments villous in the lower one-half or one-third, not
curved-geniculate; anthers 10 mm long. Style glabrous, as long
as the petals; stigma globose, 2 mm in diameter. Fruit ovoid or
ellipsoid, cinereous-puberulent, nearly 2 cm long.

LuzON, Province of Cagayan, Malueg, For. Bur. 14773 Darling, April 16,
1909, on slopes along streams, altitude about 60 m (type); Casambalangan,
For. Bur. 13112 Bernardo, May, 1909: Province of Laguna, Los Banos,
Elmer, April, 1906, Alberto, May, 1905; Santa Maria Mavitac, For. Bur.
10084 Curran, February, 1908: Province of Camarines, Caramoan, For. Bur.
10576 Curran, June, 1908; Mount Isarog, For. Bur. 10501 Curran, May,
1908: Province of Tayabas, For. Bur. 10388 Curran.

A species manifestly allied to Alangium salviifolium (L. f.) Wangerin,
differing in its very much longer, 5-merous flowers, fewer stamens, and in
its filaments which are not curved-geniculate near the base, but which are
more or less pilose for the lower one-third or one-half, and very much
longer than in A. salviifolium. ’

I do not hesitate to refer here Cuming 1716 from Samar, which is cited
by Wangerin™ under Alangium salviifolium (L. f.) Wangerin, subsp.
hexapetalum (Lam.) Wangerin. Our specimen of Cuming’s plant has 5-
merous flowers with the petals 4 cm in length, characters by which I have
distinguished the species above described. Alangium salviifolium var. hex-
apetalum, as described by Wangerin, has 6-merous flowers and petals 2 to
2.5 cm in length.

In Cagayan Province the species is known as apitan; in Camarines as
bunglas and malatapay.

This form has previously been considered by Philippine botanists to
represent Alangium octopetalum Llanos, F.-Villar and Vidal reducing
that species to Alangium lamarckii Thwaites. In working over the iden-
tifications of the species described by Blanco, I followed F.-Villar in re-
ferring A. octopetalum to A. lamarckii. The description of Alangiwm octo-
petalum Llanos, for Blanco definitely states that it was described by Llanos,
is very short and incomplete, but a careful examination of it shows con-
clusively that it can not be the same as Alangium salviifolium, or A. lon-
giflorum. The description calls for a plant with 8 petals and from 9 to
11 stamens, with no style or stigma; if Llanos correctly described his plant,
then the last character would exclude Alangium octopetalum from the genus.

“ Engl. Pflanzenreich 41° (1910) 11.

PHILIPPINE PLANTS, IX. 321

ALANGIUM SALVIIFOLIUM (L. f.) Wangerin, subsp. HEXAPETALUM
(Lam.) Wangerin in Engl. Pflanzenreich 41° (1910) 9.

MINDANAO, Lake Lanao, Camp Keithley, Mrs. Clemens 594, and several
sheets without number.

The above specimens seem to agree fairly well with Wangerin’s concep-
tion of this species, as expressed in his description. The flowers have 6
petals, which vary from 2.5 to 3.3 cm in length, hence averaging considerably
longer than indicated for Alangium salviifolium. The stamens are usually
about 22, and as long as the petals, glabrous below, then villous on the
inside for from 5 to 7 mm, and glabrous above.

A specimen from Balabac Island, Bur. Sci. 400 Mangubat, may represent
still another form. In vegetative characters it is very similar to A. salvii-
folium var. hexapetalum, the petals are 6 in number, about 2.2 cm long,
while the stamens vary from 15 to 17, the filaments near the base supplied
with a prominent, internal, densely villous appendage.

ALANGIUM MEYERI Merr. in Govt. Lab. Publ. (Philip.) 35 (1905) 54;
Wangerin l. c. 15.

Additional material of this endemic species is as follows: Luzon, Prov-
ince of Cagayan, San Vicente, For. Bur. 11301 Klemme: Province of Laguna,
Los Banos, Elmer 8308: Province of Camarines, Mount Isarog, For. Bur.
104387 Curran; Dalupaon, Ahern 120. PouiLLo, Bur. Sci. 10413 McGregor.
MINDoRO, For. Bur. 12902 Rosenbluth, Whitford 1444, For. Bur. 3643,
8682, 6186 Merritt. NeEGROS, For. Bur. 4334 Everett, For. Bur. 15016 Danao.

Native names: Angatuan (Cagayan: Negrito); putian (Negros, Min-
doro: Visayan); malatapay, paang daraga (Camarines: Bicol); maraga-
bulo (Tayabas: Tagalog).

ERICACEAE.
VACCINIUM L.

VACCINIUM CAMIGUINENSE sp. nov.

Species V. jagori Warb. affinis et similimis, differt foliis
obtusis vel rotundatis, vix acuminatis vel apiculatis, calycis lobis
ciliatis.

A species very similar to and apparently closely allied to
Vaccinium jagori Warb. A small glabrous tree, the branches
terete, olivaceous or brownish, smooth. Leaves thickly coria-
ceous, subsessile, oblong-obovate to narrowly oblong-obovate, 3.5
to 8 cm long, 1.5 to 3 cm wide, somewhat brownish when dry,
shining, the lower surface a little paler than the upper one,
somewhat glandular, the apex rounded or obtuse, base narrowed
to the very short petiole, subacute or abruptly obtuse, margins
slightly recurved; midrib prominent, the lateral nerves sharply
ascending, slender, obscure, 2 or 3 basal pairs and one or two
‘additional pairs leaving the midrib below the middle; petioles
stout, very short ornone. Flowers unknown. Racemes axillary,
solitary, 5 to 6 cm long, glabrous. Nearly mature fruits sub-

399 MERRILL.

globose, glabrous, about 5 mm in diameter, their pedicels 5 to
10 mm long, 10 to 20 in a raceme. Persistent calyx lobes
triangular-ovate, acute or somewhat acuminate, 2 mm long, their
margins above distinctly ciliate.

CAMIGUIN DE MINDANAO, Bur. Sci. 14622 (type), 14680 Ramos, April 9,
1912, in forests near the summit of Mount Mahinog.

A species similar to and manifestly allied to Vaccinium jagori Warb.,
from which it differs chiefly in its obtuse or rounded, not acuminate or apic-
ulate leaves. Among the extra-Philippine species it appears to be allied
to Vaccinium bancanum Miq., from which it also differs in its vegetative
characters.

VACCINIUM EPIPHYTICUM sp. nov.

Frutex epiphyticus, 2 ad 3 m altus, glaber; foliis ovatis vel
oblongo-ovatis, coriaceis, nitidis, acute acuminatis, basi rotun-
datis vel acutis, usque ad 12 em longis, 7-plinerviis; petiolo 5 mm
longo; racemis axillaribus, solitariis, quam folia brevioribus;
floribus circiter 14 mm longis, cylindraceo-campanulatis, fauce
vix contractis.

An epiphytic shrub, manifestly closely allied to Vaccinium
barandanum Vid., glabrous or nearly so,2 to 3m high. Branches
terete, smooth, grayish, the younger ones somewhat wrinkled or
angular when dry, slender. Leaves alternate, ovate to oblong-
ovate, coriaceous, shining, 8 to 12 em long, 3.5 to 6.5 em wide,
the base broad and rounded or acute, 7-plinerved, the apex
sharply and rather slenderly acuminate, margin entire; outer
two pairs of nerves leaving the base of the leaf, the outermost
ones not reaching the middle of the leaf, the next inner pair
extending above the middle, the innermost pair leaving the
midrib 5 to 10 mm above the base, more prominent than the
others, extending nearly to the apex, the primary lateral nerves
above these basal ones about 5 on each side of the midrib, ascend-
ing, not prominent, reticulations distinct; petioles 5 mm long.
Racemes axillary, solitary, about 5 cm long, comparatively few-
flowered, glabrous or very slightly pubescent. Bracteoles
lanceolate-oblong, acuminate, membranaceous, about 11 mm long,
4 mm wide; pedicels about 12 mm long. Flowers pink. Calyx-
tube globose-ovoid, about 2 mm long, not contracted, the teeth 5,
oblong-ovate, acute, about 2 mm long, margins minutely pubes-
cent. Corolla about 12 mm long (10 mm wide when flattened),
glabrous, or with very few scattered hairs, tubular, not con-
tracted above, the lobes broadly ovate, obtuse, reflexed, about 2
mm long. Stamens 10; filaments 5 mm long, sparingly ciliate;

PHILIPPINE PLANTS, IX. 323

anthers 4 to 4.5 mm long, the terminal tubes 2 mm long, opening
by pores. Style 12 mm long, pubescent.

MINDANAO, District of Zamboanga, Sax River Mountains, back of San
Ramon, Merrill 8087, November 30, 1911, altitude about 900 m.

A species closely allied to Vaccinium barandanum Vidal, of Luzon and
Mindoro (V. hutchinsonii Merr.), which it closely resembles, differing in

its somewhat smaller flowers, shorter petioles, its epiphytic habit, and its
pubescent, not glabrous styles.

VACCINIUM LOHERI sp. nov.

Frutex parvus, epiphyticus, glaber; foliis crasse coriaceis,
ellipticis, in siccitate pallidis, nitidis, 1.5 ad 2 cm longis, integris,
apice rotundatis, basi obscure triplinerviis, acutis, 2-glandulosis,
petiolo vix 2 mm longo; floribus solitariis, axillaribus, pedicellatis.

A small, epiphytic, glabrous shrub, the branches terete, grayish,
the younger ones reddish-brown, wrinkled when dry. Leaves
numerous, crowded, elliptic, 1.5 to 2 cm long, thickly coriaceous,
when dry rather pale and shining, the apex rounded, the base
acute, with two prominent glands near the juncture with the
petiole, the margins thickened, somewhat recurved, base obscurely
3-plinerved, the lateral nerves nearly as prominent as the midrib,
nearly or quite reaching the apex, the reticulations obsolete or
nearly so; petioles 1 to 2 mm long. Flowers white, according
to the collector (corolla not seen). Pedicles solitary, axillary,
slender, about 6 mm long, somewhat curved, the basal bracteoles
ovate, about 0.5 mm long. Calyx 2 mm long, broadly urceolate,
the tube ovoid, globose, about 1.5 mm in diameter, the limb
somewhat spreading, 5-toothed, the teeth broadly triangular-
ovate, acute or slightly acuminate, 0.5 mm long.

LUZON, Province of Rizal, Montalban; that is, in the mountains back of
Montalban, Loher 6187, April 18, 1905.

Although my specimen is imperfect, the corolla being lacking, I have not
hesitated to describe this distinct form. It is well characterized by its
small, crowded, entire, obtuse, obscurely nerved leaves which are rounded at
the apex, acute and 2-glandular at the base, and by its solitary, axillary
flowers. Among the Philippine species it is most closely allied to Vacci-

nium microphyllum Blume (V. mindorense Rendle), but is very different
from that species.

MYRSINACEAE.
ARDISIA Sw.

ARDISIA CAGAYANENSIS sp. nov. § Acrardisia.

Species A. scabridae Mez affinis, differt foliis majoribus, usque
ad 16 cm longis, obscure crenato-dentatis, nervis magis num-
erosis, 25 ad 30 utrinque.

113111—_5

324 MERRILL.

A shrub or small tree, the branches brown, smooth, somewhat
shining, the branchlets distinctly ferruginous-furfuraceous,
slender. Leaves oblong-elliptic to broadly oblong-oblanceolate,
chartaceous, 10 to 16 cm long, 3 to 4.5 cm wide, subequally
narrowed at both ends, the base acute or decurrent-acuminate,
the apex sharply acuminate, the margins subentire or obscurely
serrate-crenate, the upper surface slightly shining, the lower of
about the same color, with very numerous, dark-brown glands;
nerves slender, 25 to 30 on each side of the midrib; petioles
about 1 cm long. Inflorescence terminal or subterminal, much
shorter than the leaves, usually leaf-opposed with the uppermost
leaf on the branchlet, more or less glandular-furfuraceous with
brown glands, the branches few, the flowers umbellately
arranged on the ultimate branchlets, about 6 in an umbel, the
pedicels glandular, 5 to 7 mm long. Flowers about 8 mm in
diameter. Calyx about 3.5 mm in diameter, the lobes spreading,
triangular-ovate, glandular, acute, about 1 mm long. Corolla-
lobes ovate, imbricate, acuminate, about 3.5 mm long, gland-
ular-punctate. Anthers narrowly ovoid, 2 mm long, sharply
acuminate, somewhat glandular in the median part of the back,
subsessile. Ovary ovoid, glandular, style 4 mm long. Fruit
globose, apparently fleshy when fresh, black when dry, about
7 mm in diameter.

LuZON, Province of Cagayan, Abulug River, For. Bur. 19620 Curran,
January, 1912.

A species of the section Acrardisia, manifestly allied to Ardisia scabrida
Mez, but differing from that species in very many characters.

ARDISIA RAMOSII sp. nov. § Crispardisia.

Frutex circiter 1 m altus, glaber; foliis anguste oblong-ellip-
ticis, membranaceis, utrinque subaequaliter angustatis, acute
acuminatis, subtus pallidioribus, minute vel haud puncticulatis,
subintegris vel obscurissime undulatis, usque ad 20 em longis,
nervis utrinque 15 ad 20; floribus ignotis; fructibus globosis,
valde glandulosis, glabris, sepalis oblongis, obtusis, glandulosis,
vix ciliatis, circiter 2.5 mm longis.

A shrub about 1 m high, quite glabrous, the branches pale-
brownish. Leaves narrowly elliptic-oblong, membranaceous, 15
to 20 ecm long, 4 to 7 cm wide, subequally narrowed at both
ends, the base acute or cuneate, the apex slenderly and sharply
acuminate, the margin subentire or very obscurely undulate, with
distant glands, when dry slightly shining or dull, the lower
surface a little paler than the upper, beneath eglandular or with
small, scattered glands which are not more numerous toward

PHILIPPINE PLANTS, IX. 325

the margins; nerves 15 to 20 on each side of the midrib, rather
distinct on the lower surface, anastomosing and forming sub-
marginal nerves very close to the edges of the leaf; petioles 8
to 12 mm long. Flowers unknown. Infrutescence leaf-opposed
or subterminal, always at the tips of the branchlets, the peduncle
5 mm long or less, bearing usually about 4 or 5 umbellately
arranged primary branchlets 1 to 1.5 cm in length, the branchlets
in turn bearing usually two umbellately arranged fruits, the
pedicels as long as the peduncles. Fruit globose, about 8 mm
in diameter, very prominently marked with round and elongated
glands, the persistent calyx about 7 mm in diameter, promi-
nently glandular-punctate, the lobes oblong, obtuse, about 2.6
mm long, not at all ciliate.

Luzon, Province of Cagayan, Abulug River, For. Bur. 19613 Curran,
Bur. Sci. 18968 Ramos (type), January, 1912, in forests.

A species probably most closely allied to Ardisia jagori Mez, but very
distinct in vegetative and other characters.

DISCOCALYX Mez.

DISCOCALYX LONGIFOLIA sp. nov.

Frutex glaber 2.5 m altus; foliis oblanceolatis, circiter 55 cm -
longis, coriaceis, acutis vel acuminatis, basi angustatis, acutis,
margine supra distincte dentatis; inflorescentiis bipinnatim
paniculatis, tenuibus, paniculis fasciculatis, in ramis specialibus
dispositis; floribus 0.8 mm diametro, prominente glanduloso-
punctatis.

A glabrous, dioecious shrub about 2.5 m high. Leaves appa-
rently crowded at the apices of the branches, coriaceous, oblan-
ceolate, about 55 cm long, 11 em wide at the upper two-thirds,
from there gradually narrowed to the acute or cuneate base, the
apex acute or acuminate, the margins in the upper two-thirds
distinctly and regularly dentate, the lower surface obscurely
glandular-punctate; nerves 40 or more on each side of the
midrib, rather distinct, anastomosing, the ultimate reticulations
indistinct, petioles stout, about 4 cm long. Inflorescence of
slender, bipinnately paniculate panicles 10 cm long or less, which
are fascicled near the apices of stout, specialized branches, these
branches about 4 mm thick, more or less thickened at their
apices and marked with scars of fallen bracts. Male flowers
crimson, about 3 mm in diameter, the calyx rotate, very prom-
inently glandular-punctate, the glands black, the lobes oblong,
obtuse, about 1.3 mm long, their margins thin, subhyaline. Co-
rolla of the same diameter as the calyx, prominently and densely
glandular-punctate with black glands, the lobes oblong-ovate,

326 MERRILL.

obtuse. Rudimentary ovary narrowly club-shaped, 1 mm long.
Anthers broadly ovoid, 0.5 mm long.

LUZON, Province of Tayabas, Quinacatucan, Bur. Sci. 18181 Foxworthy
& Ramos, March, 1911.

A characteristic and strongly marked species, distinguished by its
unusually long, oblanceolate, denticulate leaves, and by its rotate flowers,
the calyx of the same size as the corolla, and both calyx and corolla very
densely glandular-punctate with black glands. Its allies are Discocalyx
insignis Merr., and D. macrophylla Merr.

DISCOCALYX MACULATA sp. nov.

Frutex parvus, circiter 4 m altus, glaber; foliis oblongis,
coriaceis, utrinque angustatis, acutis vel acuminatis, nitidis,
usque ad 25 cm longis, subtus parce glanduloso-punctatis et
maculis purpureis notatis, nervis utrinque 12 ad 15, distinctis;
inflorescentiis bipinnatim paniculatis, paniculis tenuibus, glandu-
loso-punctatis, in ramis specialibus dispositis; floribus ¢ minutis,
5-meris.

A small tree or a shrub, 4 m high (fide Ramos), dioecious, gla-
brous. Leaves oblong, coriaceous, entire, 15 to 25 em long, 4
to 8 cm wide, subequally narrowed at both ends, shining, the
apex acuminate, base acute, the lower surface somewhat paler
than the upper, with minute, scattered glands and with nu-
merous, scattered, purplish spots 1 to 1.3 mm in diameter; pri-
mary lateral nerves 12 to 15 on each side of the midrib, distinct,
anastomosing, the secondary ones sometimes nearly as distinct,
the reticulations rather lax; petioles 2 to 3 cm long. Special
branches bearing the inflorescence up to 18 cm long, slightly
branched, the upper one-half often much thickened and marked
with the scars of fallen bracts, the panicles more or less fascicled
at the ends of the branches, slender, bipinnately paniculate, more
or less glandular-punctate, 3 to 8 em or more in length. Male
flowers 5-merous, small, nearly mature buds about 1 mm in
diameter, the calyx and corolla glandular-punctate.

Luzon, Province of Cagayan, Abulug River, Bur. Sci. 18967 Ramos,
February, 1912, in forests.

A species apparently most closely allied to Discocalyx montana Elm.,

which it rather closely resembles, distinguished, however, by its small
flowers and by its leaves being prominently maculate on the lower surface.

EMBELIA Burm. f.

EMBELIA NIGRO-PUNCTATA sp. nov. § Choripetalum.

Species ut videtur E. viridiflorae (A. DC.) Scheff., affinis,
differt floribus majoribus, sessilibus vel subsessilibus, petalis
acutis vel acuminatis.

An erect (or scandent?) shrub, glabrous throughout, the

PHILIPPINE PLANTS, IX. 397

branches terete, smooth, gray. Leaves coriaceous, oblong to
oblong-obovate, usually brown when dry, somewhat shining, 8 to
16 cm long, 3.5 to 7 cm wide, the base acute, apex obtuse, acute,
or very obscurely and broadly acuminate, both surfaces, but espe-
cially the lower one, with rather large, scattered, black glands;
primary lateral nerves 8 to 10 on each side of the midrib, not
very prominent, the secondary ones nearly as distinct, the retic-
ulations rather close; petioles 1.5 to 3 cm long. Inflorescence
spicate or subracemose, from the axils of fallen leaves, solitary,
2.5 to 4 cm long, the base subtended by numerous, small, im-
bricate bracts. Flowers 4-merous, sessile or subsessile, whitish,
the bracteoles lanceolate, acute or acuminate, 2 mm long, 0.6
mm wide, with black glands. Calyx 2.5 mm long, black-
glandular, the lobes ovate, acuminate, nearly 2mm long. Petals
4, free, oblong-ovate, acute or acuminate, 3.5 mm long, 1.7
mm wide, black-glandular. Filaments slender, 3 mm long;
anthers oblong, 1.5 mm long, the connective black-glandular.
Ovary ovoid, tapering into the style. Fruit ovoid, apiculate,
8 to 10 mm long, smooth.

MINDORO, near Calapan, Bur. Sci. 938 Mangubat, June, 1906, said by the
collector to be a common shrub in thickets. Probably referable here is Bur.
Sci. 1068 Ramos (distributed as Antidesma), from Bosoboso, Province of
Rizal, Luzon, the specimen in fruit.

The species is manifestly allied to Hmbelia viridiflora (A. DC.) Scheff.,
but differs in the size of its flowers which are sessile or subsessile, acute

or acuminate petals, and in its leaves being punctate with decidedly scat-
tered but prominent black glands.

MAESA Forsk.

MAESA FERRUGINEA sp. nov. § Humaesa.

Arbuscula circiter 1 m alta, ramulis foliis inflorescentiisque
plus minusve ferrugineo-pilosis; foliis oblongo-ovatis, crenato-
dentatis, usque ad 12 cm longis, nervis utrinque circiter 6; ra-
cemis axillaribus, solitariis, densis, petiolo subaequantibus vel
brevioribus; sepalis extus dense tomentosis.

A shrub about 1 m high, all parts rather densely and softly
ferruginous-pilose. Branches dark-brown, pilose, with small
scattered lenticels. Leaves chartaceous, oblong-ovate, 7 to 12
cm long, 2.5 to 5.5 ecm wide, both surfaces pilose with soft
scattered hairs, base rounded or subacute, apex acute or slightly
acuminate, margins distinctly crenate-serrate; petioles very
densely ferruginous-pilose, about 2 cm long; nerves 5 to 7 pairs,
prominent, reticulations indistinct. Inflorescence racemose,
densely pilose, very rarely in depauperate panicles, the racemes
dense, many-flowered, usually about as long as the petioles; pedi-

298 MERRILL.

cels 1 to 1.5 mm long; bracts lanceolate, acuminate, 1 mm long.
Calyx about 2 mm long, the lobes ovate, obtuse or acute, about
8 mm long, marked with longitudinal lines, obscurely glandular,
the tube and lobes densely pilose. Corolla about 1.8 mm long, the
lobes suborbicular, rounded, slightly glandular-punctate, about
one-half as long as the corolla. Anthers 0.5 mm long, exceeding
the filaments. Ovary wholly inferior, the apex quite glabrous.
Fruit unknown.

Luzon, Province of Abra, Mount Paraga, Bur. Sci. 7109 Ramos, February
7, 1909, said to grow in forests at an altitude of about 300 m.

A species allied to Maesa manillensis Mez, but very distinct from that

species; it is well characterized by its rather dense, soft, ferruginous
tomentum.

MAESA PACHYPHYLLA sp. nov. § Humaesa.

Species ut videtur M. ramentaceae affinis, differt foliis mino-
ribus, crasse coriaceis, inflorescentiis axillaribus terminalibusque
puberulis.

Apparently a scandent shrub, glabrous except the inflores-
cence, the branches terete, lenticellate, the reddish-brown slender
branchlets terete, with few scattered lenticels. Leaves ovate to
elliptic-ovate, thickly coriaceous, pale and shining when dry, of
about the same color on both surfaces, 5 to 8 cm long, 2 to 4
em wide, the base rounded or acute, the apex obtuse or very
broadly and shortly acuminate, margins entire; lateral nerves
5 or 6 on each side of the midrib, curved-ascending, the retic-
ulations obscure; petioles 1 to 1.5 cm long. Inflorescence puber-
ulent, axillary, solitary, narrow, bipinnately-paniculate, by the
reduction of the upper leaves forming terminal panicles, the
lower ones 10 em long or less, their branches less than 1 em in
length, the upper ones much shortened, the upper reduced leaves
about 1 em long, narrowly oblong. Flowers unknown, race-
mosely arranged. Bracts oblong-lanceolate, puberulent, acumi-
nate, about 1.5 mm long, the pedicels about 1.2 mm long, with
two, ovate, acuminate, 1 mm long bracteoles immediately below
the fruit. Fruit globose, glabrous, about 2 mm in diameter,
the persistent calyx-lobes reniform-ovate, about 0.5 mm long,
not punctate, their margins minutely ciliate.

CEBU, hills back of the City of Cebu, Bur. Sci. 11080 Ramos, March, 1912.

A species apparently closely allied to Measa ramentacea Wall., from
which it differs in its smaller, much thicker leaves, and its puberulent
inflorescence. It is distinguishable among the Philippine species by its very
thick, entire, glabrous leaves in conjunction with is puberulent axillary and
pseudo-terminal inflorescence.

PHILIPPINE PLANTS, IX. 329

OLEACEAE.
LIGUSTRUM Linn.

LIGUSTRUM PUBINERVE Blume Mus. Bot. Lugd. Bat. 1 (1850) 314.
Visiania pubinervis Miq. Fl. Ind. Bat. 2 (1856) 548.
Ligustrum cumingianum Decne. in Nouv. Arch. Mus. Paris II 2 (1879)
28; Rolfe in Journ. Bot. 23 (1885) 214; Vid. Phan. Cuming. Philip.
(1885) 125; Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 116.

There seems to be but a single species represented in the Philippines,
although there is some variation in the material before me. The type
of Blume’s species was from the Philippines, collector not indicated, and
the leaves are described as “in nervo medio subtus pubescentibus.” Most
of our material has the leaves entirely glabrous, but a few specimens from
northern Luzon agree with Blume’s description as to the pubescent
midrib: Hlmer 6031, Vanoverbergh 78, Bur. Sci. 5998 Ramos, and For.
Bur. 15993 Bacani. Eighteen other specimens from northern and central
Luzon, and from Mindoro, agree with the above specimens in all essential
characters, but have entirely glabrous leaves. Unless distinctions are
exceedingly finely drawn, but a single species is represented. It is with
no hesitation that I reduce Ligustrum cumingianum Decne. (1879), to
L. pubinerve Blume (1851). It is very closely allied to some species of
southern China (Henry 7879 EF), and of India; in fact Miquel has queried
if it is not the same as Ligustrum pubescens DC.

Doctor H. Hallier has kindly examined the type of Blume’s species in
the Rijks Herbarium, Leiden, which is indicated as “ex herb. Delessert,”
no collector given, and considers that Ligustrum cumingianum Decne. is
only a more glabrous variety of L. pubinerve, the type of the latter and
cotype of the former differing only in the pubescent midribs of the leaves
of L. pubinerve.

LOGANIACEAE.
GENIOSTOMA Forst.

GENIOSTOMA STENOPHYLLUM sp. nov.

Frutex vel arbor parva usque ad 4 m altus, glaber vel ramulis
junioribus petiolisque minute pubescentibus; foliis anguste
elliptico-oblongis vel lanceolatis, chartaceis, acuminatis, basi
acutis, 2 ad 6 cm longis, nervis lateralibus utrinque 5 vel 6, subtus
prominentibus, anastomosantibus; floribus axillaribus, solitariis,
breviter pedicellatis; fructibus ovoideo-ellipticis, circiter 8 mm
longis.

A shrub or small tree reaching a height of 4m, glabrous except
the slightly pubescent young branchlets and petioles. Branches
pale-gray, the branchlets slender. Leaves chartaceous, narrowly
elliptic-oblong to lanceolate, 2 to 6 cm long, 0.5 to 2 em wide,
slightly falcate, acuminate, base acute, pale when dry, not turning
blackish, shining; lateral nerves 5 or 6 on each side of the midrib,
prominent on the lower surface, anastomosing in a distinct, looped,

330 MERRILL.

submarginal vein; petioles 1.38 to3 mm long. Flowers greenish-
white, axillary, solitary, rarely 2 in an axil, the pedicels about 2
mm long, the base with two or three, ovate, 0.5 mm long bracts,
a similar bracteole subtending the flower. Calyx-lobes ovate,
acute, 1 mm long. Corolla broadly campanulate, about 3 mm
long, 4mm in diameter, the throat hairy, the lobes reflexed, ovate,
obtuse, about 2 mm long. Anthers nearly 1 mm long. Ovary
subglobose, 1 mm in diameter; style stout, 2 mm long; stigma
globose. Fruit ovoid-elliptic, about 8 mm long, red when mature.
Seeds numerous, oblong-ellipsoid, brown, about 8 mm long.

Luzon, Benguet Subprovince, Mount Santo Tomas, Williams 1127 (type),
1068, 923, June and September, 1904; San Fernando Trail, Phil. Pl.
725 Merrill, May, 1911.

A species allied to Geniostoma cumingianum Benth., differing in its
relatively much narrower leaves which are pale, not at all blackish when

dry, its more prominent nerves which anastomose to form distinct sub-
marginal veins, and in its solitary, short-pedicelled flowers.

APOCYNACEAE.

ALYXIA Banks.

ALYXIA BLANCOI sp. nov.
Brabejum lucidum Blanco FI. Filip. ed. 2 (1845) 40, non Alyxia lucida
Wall.

Frutex scandens, glaber, ramis teretibus, ramulis 4-angulatis;
foliis lanceolatis vel oblanceolatis, coriaceis, nitidis, 3.5 ad 5 cm
longis, verticillatis, obtusis, breviter petiolatis; fructibus ovoideis
vel subglobosis, 1 cm longis, crasse apiculatis.

A scandent glabrous shrub, aromatic in drying. Branches
slender, terete, grayish or brownish, the branchlets distinctly
4-angled, the internodes on the branchlets 5 to 12 mm long.
Leaves whorled, 3, rarely 4 at each node, lanceolate or obanceo-
late, coriaceous, shining, 3.5 to 5 cm long, 6 to 10 mm wide, the
margins recurved, base acute, apex obtuse or subacute; lateral
nerves very obscure; petioles 1.5 to 2.5 mm long. Flowers
unknown. Fruit ovoid or subglobose, not moniliform, about
1 cm long, with a thick apiculate point, yellow when fresh.

CEBU, in thickets on dry hills back of the City of Cebu, Bur. Sci. 11118
Ramos, March, 1912.

F. Villar reduced Blanco’s species to Alywis stellata R. & S., which was
undoubtedly an error. Vidal referred Alyxia stellata F.-Vill. to his A. mon-

ilifera, but correctly excluded the reference to Blanco. In my paper on
the determination of Blanco’s species * I followed F.-Villar in his reduction

% Govt. Lab. Publ. (Philip.) 27 (1905) 58.

PHILIPPINE PLANTS, IX. 331

of Brabejum lucidum, but erroneously carried the reduction still further to
Alyxia monilifera Vid. Now that I have specimens from Cebu, from which
island Blanco received his material, which agree perfectly with his short
description, I have no hesitation in redescribing it under the above specific
name, for Blanco’s name, lucida, is invalid in Alywia.

Alyxia blancoi is allied to A. parvifolia Merr., and to A. luzoniensis
Merr., but its leaves are shaped very differently from those of either of the
above species.

ANODENDRON A. DC.

ANODENDRON AXILLARE sp. nov.

Frutex scandens subglaber; foliis oblongis, nitidis, usque ad
15 cm longis, apice breviter acuminatis, basi acutis vel subrotun-
datis, nervis patulis, tenuibus, utrinque circiter 13; inflorescentiis
axillaribus, cymosis, vix 2 cm longis, brevissime pedunculatis,
alabastro 8 mm longo.

A scandent woody vine, the branches, when dry, reddish-brown,
slender, minutely and densely brownish-lepidote or punctate.
Leaves opposite, oblong, subcoriaceous or firmly chartaceous, 12
to 15 cm long, 5 to 6 cm wide, shining and of about the same color
on both surfaces when dry, the upper surface entirely glabrous,
the lower one with minute, scattered, brown, lepidote spots, base
subacute or rounded, apex abruptly short-acuminate; nerves
distant, spreading, not prominent, anastomosing near the margin,
about 13 on each side of the midrib, the secondary ones nearly as
distinct as the primary ones; petioles 8 to10 mm long. Inflores-
cence axillary, cymose, short, many-flowered, the peduncle 2 to
3 mm long. Flowers greenish to yellowish-white, in nearly
mature bud about 8 mm long, the pedicels 2 to 3 mm long, each
subtended by two or three, ovate, 1 mm long bracteoles. Sepals
5, ovate, acute, imbricate, 2 mm long, eglandular. Corolla
apparently salver-shaped, the tube cylindric, not inflated, about
5 mm long, 1.5 mm in diameter, the lobes twisted to the left,
oblong-ovate, acute, somewhat falcate, about 6 mm long, 2.5 mm
wide. Stamens inserted at the base of the tube, conniving in a
cone around the stigma, united to it by projections from the
connective; anthers oblong-ovate, sessile or subsessile, base
sagittate by prolongation of the cells, apex acuminate or acute,
about 1.5 mm long. Disk cup-shaped, about 0.5 mm high,
crenate. Carpels 2, free, united by their backs to the disk,
enclosed, that is, not exceeding the disk; ovules many. Stigma
narrowly top-shaped, 1 mm long.

NecRos, Faraon, For. Bur. 18572 Meyer & Foxworthy, August 25, 1909.

A species apparently allied to Anodendron pauciflorum Hook. f., of
the Malay Peninsula, but judging from the description quite distinct.

332 MERRILL.

ANODENDRON LOHERI sp. nov.

Frutex alte scandens glaber; foliis lanceolatis ad anguste
elliptico-lanceolatis, utrinque acutis vel leviter acuminatis, usque
ad 10 cm longis, nervis utrinque circiter 9, obscuris; paniculis
terminalibus, parvis, pedunculatis, multifloris; corollae tubo intus
villosus, 4 ad 5 mm longus.

A glabrous scandent shrub reaching a height of 10 m or more,
the branches slender, reddish-brown when dry. Leaves lanceo-
late to narrowly elliptic-lanceolate, narrowed about equally at
both ends and acute or slightly acuminate, 5 to 10 cm long, 1.5
to 3 cm wide, shining when dry, the lower surface paler than the
upper; nerves about 9 on each side of the midrib, slender, in-
distinct, distant, faintly anastomosing, the reticulations obsolete
or nearly so; petioles 1 to 1.5 ecm long. Panicles terminal,
including the peduncle 4 to 9 cm long, usually many-flowered,
trichotomous, the peduncles 2 to 5 cm long. Flowers numerous,
cream-colored or yellowish, fragrant, somewhat crowded on the
ultimate branchlets of the inflorescence, the bracts ovate, 1.3 to
1.7 mm long. Sepals oblong-ovate, acute, imbricate, 2 to 2.5
mm long. Corolla-tube cylindric, 4 to 5 mm long, about 2 mm
in diameter, villous inside, the lobes oblong-lanceolate, obtuse,
falcate, about 6 mm long, 1.5 mm wide, villous on the inner
surface, spreading, in bud twisted to the left. Anthers sessile on
the bottom of the tube, lanceolate, acuminate, 2 mm long, sur-
rounding the stigma and adherent to it. Stigma about 1 mm
long, top-shaped. Disk cupular, about 0.5 mm long. Follicles
unknown.

LuzoN, Province of Zambales, Mount Pinatubo, Loher 6494 (type), Feb-
ruary, 1906: Province of Bataan, Lamao River, Mount Mariveles, For.
Bur. 2607 Meyer, February, 1905, For. Bur. 6256 Curran, February, 1907,
Whitford s. n., 1905, month not indicated.

This new species is allied to Anodendron paniculatum A. DC., and to
A. candolleanum Wight, and one of the specimens from Bataan Province
was identified as the former species “, all duplicates having been distributed
as A. paniculatum A. DC. Anodendron loheri differs from A. paniculatum
especially in its very much smaller leaves and fewer nerves; it is appa-
rently much more closely allied to A. candolleanum Wight, differing not
only in its smaller leaves but in the corolle-tube and lobes being pro-
minently villous.

A specimen from Rizal Province, Montalban, For. Bur. 2452 Ahern’s
collector, distributed as Anodendron paniculatum A. DC., is probably refer-
able to A. loheri, but the leaves are a little larger than in the type, and
the nerves are decidedly prominent, in fact nearly as prominent as in
typical A. paniculatum.

“This Journal 1 (1906) Suppl. 117.

PHILIPPINE PLANTS, IX. 383

ANODENDRON MANUBRIATUM (Wall.) comb. nov.

Echites manubriata Wall. Cat. (1829) No. 1663.

Echites paniculata Roxb. Fl. Ind. ed. 2, 2 (1832) 17, non Poir.

Anodendron paniculatum A. DC. Prodr. 8 (1844) 444; King & Gamble
in Journ. As. Soc. Beng. 74° (1907) extra number 486; Rolfe in
Journ. Bot. 23 (1885) 214; Vid. Rev. Pl. Vasc. Filip. (1886) 187.

Echites coriacea Wall. 1. c. no. 1586, non Blume.

This species was first reported from the Philippines by Rolfe, Cuming
1176, from the Province of Ilocos Sur, Luzon. It is apparently also
represented by the following specimens: LUZON, Province of Union, Mount
Tonglon, Loher 6482, April, 1906. PALAWAN, Puerto Princesa, Bur. Sci.
352 Bermejos, January, 1906. The specimens agree closely with the de-
scriptions, and with a specimen in our herbarium from a plant cultivated
in the Royal Botanic Garden at Calcutta.

Echites paniculata Roxb., being invalidated by the earlier EF. paniculata
Poir., it is not clear how the specific name can be retained under Ano-
dendron. Echites manubriata Wall. apparently supplies the earliest name
that is tenable, although originally only a nomen nudum.

India to Ceylon, through the Malay Peninsula to the Malay Archi-
pelago and the Philippines.

ICHNOCARPUS R. Br.

ICHNOCARPUS OVATIFOLIUS A. DC. Prodr. 8 (1844) 485; Vid. Rev. Pl.

Vasc. Filip. (1886) 116; Hook. f. Fl. Brit. Ind. 3 (1882) 670; King
& Gamble in Journ. As. Soc. Beng. 74’ (1907) extra number 494.

Echites caudata Blanco FI. Filip. (1887) 106, ed. 2 (1845) 77, non
Burm.

Springia indica Muell.-Arg. & Van Huerck Plant. Nov. (1871) 143.

Ichnocarpus frutescens F.-Vill. Novis. App. (1880) 131; Vid. Sinop-
sis Atlas (1888) t. 67, f. H.: Naves in Blanco FI. Filip. ed. 3,
pl. 97, non R. Br.

Ichnocarpus navesti Rolfe in Journ. Linn. Soc. Bot. 21 (1884) 313.

BATANES ISLANDS, Dalupiri, Bur. Sci. 10647 McGregor. Luzon, Province
of Ilocos Norte, For. Bur. 13892 Merritt & Darling: Benguet Subprovince,
Elmer 6610: Province of Nueva Ecija, For. Bur. 8499 Curran, For. Bur.
22131 Alvarez: Province of Pangasinan, Merrill 2871: Province of Pam-
panga, Merrill 4214: Province of Bataan, For. Bur. 1464 Ahern’s collector,
Leiberg 6013: Manila, Didrichsen, Galathea Expedition 3192, Loher 3884:
Province of Bulacan, Bur. Sci. 6118 Robinson & Merritt: Province of Rizal,
Guerrero 36, Bur. Sci. 13536 Ramos, For. Bur. 3125, Merrill 2629, Loher
8889, 6527: Province of Cavite, Bur. Sci. 1820 Mangubat: Province of La-
guna, Hallier, Elmer. MINbDoRO, Cuming 1547, 1572, Bur. Sci. 10869 Celes-
tino, For. Bur. 5385 Merritt. PALAWAN, Bur. Sci. 206 Bermejos, Bur.
Sci. 690 Foxworthy. MASBATE, Merrill 3080. CrBU, Cuming 1770. NEGROS
Bur. Sci. 9924 Robinson, Cuming 1809 (cotype). MINDANAO, Lake Lanao,
Mrs. Clemens 665: District of Davao, Piper 476. BASILAN, For. Bur. 6569
Hutchinson.

India, Malay Peninsula, Sumatra, Borneo, Celebes, and Timor.

This species is common and widely distributed at low altitudes in the
Philippines, but I can see no reason for considering that more than

334 MERRILL.

one species is represented. There is some variation in the shape of the
leaves, and in the pubescence, sometimes the leaves being quite glabrous,
sometimes villous on the midrib beneath.

Echites affinis R. & S. Syst. 4 (1819) 393 (E. caryophyllata Roth Nov.
Sp. Pl. (1821) 133, non Roxb.) would supply the oldest specific name for
this species were Hooker f. correct in the reduction of Echites affinis to
Ichnocarpus ovatifolius A. DC.“ Finding, however, that the description
given by Roemer & Schultes did not agree at all well with specimens of
De Candolle’s species, I asked Herr J. Martin, director of the Natural
History Museum at Oldenburg, Germany, to examine Roth’s specimen,
the type both of Echites afinis R. & S., and of E. caryophyllata Roth.
This he has kindly done and informs me that Ichnocarpus ovatifolius
A. DC. is certainly not the same as Es caryophyllata Roth (E. affinis
R. & S.) in Roth’s herbarium. The sketch he has kindly supplied me
shows Echites affinis to have broadly ovate or ovate-elliptic leaves with
but two pairs of lateral nerves, the latter character entirely at variance
with Ichnocarpus ovatifolius A. DC.

Ichnocarpus frutescens R. Br., has been reported from the Philippines
by several authors, but I have seen no material from the Archipelago
that I would refer to that species. Ichnocarpus navesii Rolfe is typified
by plate 97 of the third edition of Blanco’s “Flora de Filipinas;” in fact
the species was based solely on the plate. The figure is manifestly a
very poor representation of J. ovatifolius, and hence J. navesii Rolfe is
reduced.

Three species enumerated by F.-Villar*” under this genus are not species
of Ichnocarpus, but I. velutinus F.-Vill. is Aganosma velutina A. DC., I.
acuminatus F.-Vill. is Aganosma acuminata G. Don, and J. macrocarpus
F.-Vill. is a synonym of A. acuminata G. Don.

MICRECHITES Mig.

MICRECHITES SCHRIECKI! (Huerck & Muell.-Arg.) Rolfe in Journ. Bot.
23 (1885) 214; Vid. Phan. Cuming. Philip. (1885) 126.
Ecdysanthera schrieckii Huerck & Muell.-Arg. Obs. Bot. 191.
Trachelospermum sp. Merrill in Forestry Bureau (Philip.) Bull. 1
(1903) 49.
Trachelospermum philippinense Elm. Leafi. Philip. Bot. 2 (1908) 488.

LUZON, Province of Tayabas, Lucban, Elmer 9185 (cotype of Trachelo-
spermum philippinense Elm.), in flower, May, 1907: Province of Camarines
Sur, Pasacao, Ahern 300, January, 1902, in fruit: Province of Albay,
Cuming 910 (cotype of Ecdysanthera schrieckii Huerck & Muell.-Arg.).
NEGROS, Gimagaan River, For. Bur. 4250 Everett, April, 1906, in fruit.
MINDANAO, District of Zamboanga, Dumanquilas Bay, For. Bur. 12360 Hut-
chinson, previously determined as Ichnocarpus ovatifolius A. DC. Locally
known in Camarines as bayacto.

The cotypes of Trachelospermum philippinense Elm., and Micrechites
schrieckii Rolfe, both in flower, are identical in all respects, hence the
necessity of reducing the former species.

Endemic.

“Fl, Brit. Ind. 3 (1882) 670. “Novis. App. (1880) 181.

PHILIPPINE PLANTS, IX. 8385

PARSONSIA R. Br.

PARSONSIA OBLANCIFOLIA sp. nov.

Frutex scandens, glaber; foliis oblanceolatis ad anguste
oblongo-obovatis, usque ad 12 cm longis, apice acuminatis, basi
angustatis, acutis, nervis utrinque 8 ad 10; cymis terminalibus,
pedunculatis, multifloris, floribus circiter 1 cm longis, filamentis
inter se spiraliter dextrorsum contortis.

A scandent, glabrous, woody vine, the branches pale-brown or
somewhat grayish, very minutely pustulate, the younger ones
dark-brown. Leaves chartaceous, oblanceolate to narrowly
oblong-obovate, 7 to 12 cm long, 2 to 3.5 cm wide, slightly
shining when dry, the lower surface paler than the upper one,
apex shortly but sharply acuminate, base narrowed, acute; nerves
8 to 10 on each side of the midrib, distant, anastomosing, the
reticulations very lax, faint or often nearly obsolete; petioles
1 to 1.5 cm long. Cymes terminal, many-flowered, 5 to 6 cm
long, sometimes slightly pubescent, usually trichotomous, the
peduncle 1.5 cm long or less, the branches again trichotomous,
the flowers crowded on the ultimate branchlets. Bracteoles
oblong-ovate, acute, 1 to 1.5 mm long, the pedicels 5 to 8 mm
long. Sepals oblong, acute or obtuse, about 3 mm long, slightly
imbricate in bud. Corolla-tube about 3.5 mm long and 3 mm
in diameter in the middle, more or less inflated, glabrous on
both sides, the throat glabrous, not bearded, the lobes oblong-
lanceolate, obtuse, 6 to 7 mm long, about 1.5 mm wide, base
a little wider. Filaments about 5 mm long, somewhat villous,
strongly twisted about the style; anthers lanceolate, 4 mm long.

LuzoN, Province of Zambales, Candelaria, Bur. Sci. 4723 Ramos,
December 7%, 1907.

A species manifestly allied to Parsonsia confusa Merr., and P. cumin-
giana A. DC., of the Philippines, and P. spiralis Wall., of India and
Malaya, but quite different in vegetative characters. It is probably most
closely allied to P. cumingiana A. DC., but is at once distinguished from
that species by its oblong-lanceolate to oblong-ovate, more numerously

veined leaves; from P. confusa it differs in its leaf characters and its
glabrous or nearly glabrous inflorescence.

VALLARIS Burm.

VALLARIS ANGUSTIFOLIA sp. nov.
Arbor parva, 6 ad 10 m alta, glabra; foliis lanceolatis, coriaceis
vel subcoriaceis, utrinque angustatis acuminatisque, usque ad 8
cm longis, 0.7 ad 1.5 em latis; floribus axillaribus, fasciculatis,
circiter 13 mm diametro, corollae tubo apice constricto.

336 MERRILL.

A small glabrous tree 6 to 10 m high, the branches slender,
terete, dark-reddish-brown, often nearly black when dry.
Leaves lanceolate, coriaceous or subcoriaceous, narrowed and
more or less acuminate at both ends, shining when dry, the
lower surface paler than the upper one, 5 to 8 em long, 0.7 to
1.5 cm wide; nerves 6 or 7 on each side of the midrib, slender,
distant, spreading, anastomosing, not glandular in the axils, the
reticulations obsolete or nearly so; petioles 5 to 10 mm long.
Flowers white or yellowish, fragrant, axillary, fascicled, few
to many in an axil, or sometimes in a very dense, fascicle-
like, reduced and very shortly peduncled cyme, the pedicels
slender, 5 to 8 mm long. Sepals slightly pubescent, triangular-
ovate, acute, about 1.5 mm long, imbricate. Corolla-tube 5 mm
long, inflated below, narrowed above and constricted at the apex,
the lobes spreading, oblong-ovate, obtuse, falcate, 6 to 7 mm long,
about 3.5 mm wide, somewhat villous on the inner surface.
Stamens lanceolate, acuminate, slightly hairy on the back, the
base of the connective with a rather large gland. Style slender,
5 mm long.

LUZON, Province of Sorsogon, Sorsogon, For. Bur. 10507 Curran, June,
1908, in thickets, altitude about 300 m; near Pilar, For. Bur. 15077
Rosenbluth (type), April 15, 1909, in flat lands not far back of the limits
of high tide, locally known as lanete.

A species well characterized by its small lanceolate leaves, probably
most closely allied to Vallaris lancifolia Hook. f., of the Malay Peninsula,
but very different from that species. The first representative of the
genus to be reported from the Philippines other than an undetermined
form, Vidal 3278, from the Province of Albay, reported as Vallaris sp. by
Ceron.”

VOACANGA Thouars.

VOACANGA MEGACARPA sp. nov.

Arbor glabra usque ad 10 m alta; foliis oblongis vel anguste
oblongo-obovatis, breviter acuminatis, basi acutis, petiolatis, 15
ad 30 cm longis, nervis utrinque 10 ad 14; paniculis axillaribus,
longe pedunculatis, dichotomis, paucifloris; calycibus 2 ad 3 cm
longis, profunde 5-lobatis; corolla alba, tubo circiter 3 em longo,
contorto, lobis late oblique ovatis, usque ad 4 cm longis; fruc-
tibus globosis, usque ad 10 cm diametro.

A glabrous tree 5 to 10 m high, the branches pale-gray.
Leaves oblong to narrowly oblong-obovate, 15 to 30 cm long,
6 to 12 cm wide, entire, apex shortly and rather abruptly acum-
inate, base narrowed, cuneate; nerves 10 to 14 on each side

* Cat. Pl. Herb. (Manila) (1892) 115.

PHILIPPINE PLANTS, IX. 337

of the midrib, distant, prominent, the reticulations very faint,
lax; petioles 1 to 2.5 cm long, the base somewhat inflated and
clasping the stems. Inflorescence paniculate, axillary, lenticel-
late, the peduncles about 15 cm long, dichotomously branched,
the flowers somewhat crowded at the ends of the branches, the
two elongated primary branches frequently equaling the pe-
duncles in length, marked with numerous scars of fallen flowers
or branchlets. Flowers large, white, fragrant. Calyx 2 to 3
em long, split nearly to the base into 5, narrowly oblong, erect,
obtuse lobes. Corolla-tube about 3 cm long, somewhat con-
tracted above, distinctly twisted at and above the insertion of
the anthers; lobes spreading, obtuse, obliquely and broadly ovate,
up to 3.5 cm long and 4 cm wide. Anthers narrowly lanceolate,
acuminate, base sagittate, about 7 mm long. Ovary of 2 dis-
tinct ecarpels united by the style, surrounded by the fleshy disk.
Fruit in pairs, sessile, globose, when fresh smooth, up to 10 cm °
in diameter, when dry much smaller, wrinkled. Seeds very
numerous, rugose, 8 to 10 mm long.

Luzon, Province of Tayabas, Tununi River, along streams, Bur. Sci.
12324 Foxworthy, February, 1911 (type); Binangonan, Whitford 829,
September, 1904; Baler, Bur. Sci. 10668 McGregor, August, 1909: Province
of Camarines, Tinambuc, For. Bur. 14256 Aguilar, April, 1909.

A very characteristic species, readily distinguished from the others in

the genus by its large flowers and fruits, and especially by its calyx
being split nearly to the base into 5, long, narrow lobes.

TABERNAEMONTANA L.

TABERNAEMONTANA CORDATA sp. nov.

Frutex circiter 2 m altus, fructibus immaturis exceptis glaber ;
foliis oblongis, usque ad 14 cm longis, acute acuminatis, sessi-
libus, basi late cordatis; cymis axillaribus, paucifloris; folliculis
maturis glabris, junioribus griseo-puberulis, rubris, oblongis,
falcatis, longitudinaliter 5-carinatis; seminibus circiter 15.

An erect shrub about 2 m high. Branches slender, terete,
light-gray. Leaves oblong, sessile, base broad, cordate, apex
sharply acuminate, 2 to 4 cm wide, 6 to 14 cm long, chartaceous,
glabrous, one of each pair somewhat smaller than the other;
nerves 9 or 10 on each side of the midrib. Cymes in the upper
axils, short, few-flowered, the flowers often only 2 or 3, ap-
parently white, the pedicels slender, about 1.5 cm long. Calyx-
tube short, the lobes ovate, acute, about 1.5 mm long. Corolla-
tube about 1.4 em long, 1.5 mm in diameter in the middle,
the lobes narrowly oblong, falcate, rounded, about 9 mm long,
2.5 to 3 mm wide. Calyx-teeth oblong-ovate, acute. Follices

338 MERRILL.

red, oblong, when young gray-puberulent, soon becoming quite
glabrous, 2 to 3.5 cm long, 1.2 to 1.5 em wide, curved, slightly
keeled down the back, rather distinctly winged or broadly keeled
down the suture, with two intermediate keels along the sides.
Seeds about 15, irregularly triangular-ovoid, not or but very
slightly sulcate.

MINDANAO, Butuan Subprovince, Cabadbaran, Weber, April, 1911 (type),
in clearings, altitude about 15 m, Piper 287, 345, April, 1911.

A species strongly characterized by its sessile leaves which are broad
and cordate at the base.

VERBENACEAE.
CALLICARPA L.

CALLICARPA ELEGANS Hayek in Just’s Bot. Jahresb. 33 (1905) 8.

This species was based on Cuming 1460, a specimen of which is in
the herbarium of the Bureau of Science; according to Cuming’s own list
of localities, the above number was collected in the Province of Camarines
Sur, Luzon. The species appears to be quite widely distributed in the
northern Philippines, and is also represented by the following specimens:

LuZON, Province of Abra, Mount Paraga, Bur. Sci. 7054 Ramos, Feb-
ruary, 1909: Province of Ilocos Norte, Mount Piao, For. Bur. 13994 Merritt
& Darling: Province of Zambales, Subic, Hallier, January, 1904; Cande-
laria, Bur. Sci. 4815 Ramos, December, 1907; Cabiluagan, For. Bur. 7018
Curran, May, 1907: Province of Bataan, Lamao River, Williams 340, De-
cember, 1903. MINDORO, south-west of Lake Naujan, For. Bur. 6725 Mer-
ritt, April, 1907.

Endemic.

CALLICARPA CAULIFLORA sp. nov.

Arbor parva, ramulis quadrangularibus stellato-plumoso-
pilosis glandulosisque; foliis amplis, chartaceis, oblongo-ellipticis,
acuminatis, usque ad 35 em longis, leviter irregulariter denticu-
latis vel subintegris, supra glabris, subtus glandulosis, ad costa
nervosque plus minusve stellato-pilosis; inflorescentiis fascicu-
latis, caulifloris; floribus 4-meris, pedicellatis; corolla circiter
7 mm longa.

A small tree, the branchlets quadrangular, rather stout, rather
densely covered with brownish plumose-stellate hairs and also
yellow-glandular. Leaves chartaceous, oblong-elliptic, 30 to 35
cm long, 9 to 13 em wide, the apex slenderly and sharply acu-
minate, the base gradually narrowed, margins distantly and
irregularly denticulate or subentire, the upper surface glabrous,
the lower one somewhat paler, yellow-glandular, sparingly
stellate-pilose on the midrib, nerves, and reticulations, the hairs
often plumose; primary nerves about 14 on each side of the
midrib, prominent beneath, anastomosing, the reticulations prom-

PHILIPPINE PLANTS, IX. 339

inent, rather lax; petioles stout, densely stellate-pubescent, 1
to 2 cm long. Inflorescences on the trunk, of sessile, hemi-
spherical, rather dense fascicles 2 to 3 cm in diameter, slightly
hirsute, the pedicels 3 to 5 mm long, the bracteoles linear-
lanceolate, 1 to 2 mm long. Calyx oblong-ovoid, about 3 mm
long, 1.8 mm diameter, equally 4-toothed, the teeth narrowly
ovate, acute, 0.5 mm long. Corolla dark-red, about 7 mm long,
cylindric, 4-lobed, slightly glandular and pubescent externally.
Anthers oblong, 3 mm long, glandular on the back. Fruit glo-
bose, the lower one-half enclosed in the calyx, about 4 mm in
diameter, containing 4 pyrenes.

MINDANAO, District of Zamboanga, Port Banga, in canyons in forests,
altitude about 20 m, For. Bur. 9321 Whitford & Hutchinson, January 13,
1908.

A species well characterized by its cauline, fascicled inflorescence, in
this character differing from all the other species of the genus known

to me. Allied to C. ramiflora Merr., but with a quite different indu-
mentum.

CALLICARPA DOLICHOPHYLLA sp. nov.

Arbor parva, glabra, vel partibus junioribus pilis simplicibus,
non stellatis, munita; foliis lanceolatis, chartaceis, 15 ad 30 cm
longis, 2.5 ad 7 cm latis, subtus glandulosis, apice tenuiter acu-
minatis, basi acutis vel acuminatis; nervis utrinque 13 ad 16;
cymis axillaribus, solitariis, laxis, usque ad 6 cm longis, pedun-
culis quam petioli longioribus.

A small tree, entirely glabrous, or the younger parts more or
less pubescent with simple, not stellately arranged hairs, a few
hairs sometimes persistent on the leaves, especially beneath.
Branches terete or somewhat compressed, smooth, brown, gla-
brous or somewhat pubescent when young. Leaves lanceolate,
chartaceous, 15 to 30 cm long, 2.5 to 7 cm wide, mostly entirely
glabrous except the younger ones which are sometimes puberulent
or pubescent, the base acute or acuminate, the apex rather slen-
derly acuminate, the margins crenate-denticulate or subentire, the
lower surface distinctly yellow-glandular, the glands in minute
pits; petioles 1 to 2.5 em long; nerves 13 to 16, distinct, curved-
ascending, anastomosing, the reticulations subparallel. Cymes
axillary, solitary, lax, glabrous or somewhat pubescent with
simple hairs, 6 cm long or less, often as wide as long, the ped-
uncles longer than the petioles. Flowers apparently reddish.
Calyx cup-shaped or somewhat funnel-shaped, 1 to 1.5 mm long,
with 4 obscure, short teeth. Corolla somewhat exserted, 2.5 to

3 mm long, glabrous, the tube somewhat enlarged upward.
113111——6

340 MERRILL.

Stamens exserted; anthers 0.6 mm long. Fruit (immature),
globose, small, containing 4 pyrenes.

LUZON, Province of Cagayan, Cuming 1880 (type): Province of Nueva
Vizcaya, Mount Umugum, Bur. Sci. 8268 Ramos, May, 1909: Province of
Rizal, Bosoboso, Bur. Sci. 1063 Ramos, July, 1906.

Cuming’s specimen has been referred by Schauer to Callicarpa longi-
folia Lam., var. subglabra Schauer, in conjunction with other specimens
from India, Java, and Japan. The description does not apply particularly
well to the Philippine plant, which to me does not appear to be very
closely allied to Lamarck’s species. It is well characterized by lax cymes
and its entirely glabrous vegetative parts, or if pubescent at all, then
with simple, not stellate hairs, in this last character differing from most,
if not all, other species of the genus. Two specimens of Cuming’s No.
1330 are in the Herbarium of the Bureau of Science, one with rather
narrow, somewhat pubescent leaves, and with pubescent stems and inflo-
rescence, quite manifestly a young stage of the plant, the other with
larger, glabrous leaves and inflorescence; this latter I have made the type
sheet.

CALLICARPA RIVULARIS sp. nov.

Species C. angustae Schauer simillima et valde affinis, differt
foliis paulo majoribus, nervis lateralibus paucioribus, indumento
plus minusve stellato-plumosis, antheris majoribus.

A shrub, apparently sometimes subscandent, 2 to 5 m high.
Branches terete, or the ultimate ones somewhat compressed, very
densely covered with a whitish or yellowish-white indumentum
composed of short, stellate hairs, with some stellate-plumose ones
intermixed, the inflorescences, petioles, and lower surface of
the leaves with a similar indumentum. Leaves narrowly lan-
ceolate or narrowly oblanceolate, chartaceous, 10 to 18 cm long,
1.5 to 3.5 em wide, the upper surface dark-colored when dry,
quite glabrous, or with few stellate hairs along the midrib, the
lower surface with a dense, whitish or yellowish-white indu-
mentum, about equally narrowed at both ends, the apex slenderly
acuminate, the margins in the lower one-half entire, above slightly
and irregularly denticulate; petioles about 1 cm long; nerves
about 9 on each side of the midrib, curved-ascending, anasto-
mosing, prominent beneath. Cymes axillary, solitary, the pe-
duncles about as long as the petioles, dichotomously branched,
the cymes rather lax, comparatively few-flowered, 3 cm in diam-
eter or less. Calyx somewhat funnel-shaped, about 2 mm long,
slightly 4-toothed, externally densely white- or grayish-puberu-
lent. Corolla-tube scarcely exserted, the corolla white, about
3 mm long, 4-lobed, slightly pubescent externally. Anthers 1.3
mm long, glandular on the back. Fruit globose, glabrous, about
38 mm in diameter, containing 4 pyrenes.

PHILIPPINE PLANTS, IX. 341

PALAWAN, Mount Victoria, on rocky river banks at an altitude of about
1,150 m, Bur. Sci. 660 Foxworthy (type), March 23, 1906, also from the
same locality, at an altitude of 60 m, on large boulders in the river bed,
Bur. Sci. 719 Foxworthy, March 24, 1906.

A species manifestly very closely allied to Callicarpa angusta Schauer,
differing especially in its indumentum.
CALLICARPA ANGUSTA Schauer in DC. Prodr. 11 (1847) 642.

LuZON, Province of Nueva Ecija, Cuming 1425 (cotype): Province of
Zambales, Subic, Hallier s. n.: Province of Rizal, Bosoboso, Bur. Sci. 1030
Ramos, For. Bur. 1888 Ahern’s collector. MiINnporo, Tubili, For. Bur. 8878
Merritt, January, 1908.

Endemic.

CLERODENDRON L.

CLERODENDRON ELLIPTIFOLIUM sp. nov.

Arbor parva, circiter 3 m alta, inflorescentiis exceptis glabra;
foliis ellipticis vel ovato-ellipticis, chartaceis, usque ad 15 cm
longis, basi rotundatis, apice breviter acuminatis, margine leviter
subundulatis, nervis utrinque circiter 6, distinctis; inflorescentiis
terminalibus, subsessilibus, densis, multifloris, circiter 10 cm
diametro, bracteolis aciculatis; floribus circiter 2.5 cm longis.

A small tree or a shrub about 3 m high, glabrous except the
inflorescence. Branches terete or obscurely angled, the younger
ones olivaceous, lenticellate. Leaves elliptic to elliptic-ovate,
chartaceous or submembranaceous, olivaceous when dry, the
upper surface shining, the lower one of about the same color,
dull, 12 to 15 cm long, 7 to 11 cm wide, the base rather broadly
rounded, sometimes very obscurely cordate, the apex shortly
and stoutly acuminate, the margins obscurely subundulate, the
incipient teeth 1 to 1.5 cm apart; lateral nerves 5 or 6 on each
side of the midrib, prominent, curved, anastomosing, the retic-
ulations lax; petioles 3 to 3.5 cm long. Inflorescence terminal,
dense, many-flowered, subsessile or at least branched from the
base, cymose, sparingly pubescent with short, scattered hairs,
the bracteoles acicular, pubescent, 1 to 2.8 mm long, the inflor-
escence about 10 cm in diameter. Flowers numerous, white
or nearly so. Calyx narrowly funnel-shaped or somewhat cup-
shaped, about 6 mm long, the apex truncate, obscurely 5-toothed,
3.5 mm in diameter, narrowed below to the acute base, externally
sparingly pubescent with very short hairs. Corolla-tube slender,
cylindric, glabrous, about 18 mm long, the limb spreading, about
12 mm in diameter, the lobes ovate or broadly oblong-ovate,
rounded. Ovary glabrous.

CAMIGUIN DE MINDANAO, Panatayum, in thickets, Bur. Sci. 14463 Ramos,
March 20, 1912.

342 MERRILL.

A characteristic species not, apparently, very closely allied to the other
Philippine forms, although manifestly in the same group with Clerodendron
quadriloculare Merr., C. mindorensis Merr., and C. klemmei Elm. It is
distinguishable by its elliptic, obscurely undulate leaves, and its dense,
terminal, many-flowered, cymose panicles.

CLERODENDRON VILLOSUM Blume Bijdr. (1826) 811; Schauer in DC.
Prodr. 11 (1847) 667; Clarke in Hook. f. Fl. Brit. Ind. 4 (1885)
589; F.-Vill. Novis. App. (1880) 161; Gamble in Journ. As. Soe.
Beng. 74 (1909) Extra Number 836.

CULION, Halsey Harbor, Merrill 761, February, 1903.

Burma to the Malay Peninsula, Java, and Sumatra.

The specimen agrees closely with the description and with Singapore
specimens in our herbarium.

CLERODENDRON MINDORENSE nom. nov.
Clerodendron simile Merr. in Govt. Lab. Publ. (Philip.) 35 (1906)
64, non Pearson (1901).

A new name is necessary for the Philippine plant, as the one previously
selected by me was preoccupied.

GEUNSIA Blume.

GEUNSIA HOOKERI sp. nov.
Callicarpa pentandra Schauer in DC. Prodr. 11 (1847) pro parte,
quoad Cuming 1778.

Species G. farinosae Bl., et G. cumingianae (Schauer) Rolfe
similis et affinis, differt foliis subtus parce pubescentibus, vetus-
tioribus subglabrescentibus.

A shrub or a small tree, the young branches, petioles, leaves,
and inflorescence more or less stellate-pubescent with brown
hairs, the older parts becoming glabrous or nearly so. Branches
terete, or the younger ones somewhat compressed, ultimately
glabrous. Leaves opposite and alternate, ovate to oblong-ovate,
chartaceous, entire, the base abruptly acuminate, the apex long
and slenderly subcaudate-acuminate, 9 to 17 cm long, 5 to 9
cm wide, brownish when dry, of about the same color on both
surfaces, the upper surface slightly stellate-pubescent, the hairs
more numerous on the midrib and lateral nerves, ultimately
nearly glabrous, the lower surface slightly pubescent, and with
numerous, small, yellow, glands; nerves about 8 on each side
of the midrib, the reticulations subparallel, distinct; petioles 2
to 4 cm long. Cymes axillary, solitary, 10 cm long or less,
rather densely brown-stellate-pubescent, the peduncles about
as long as the petioles, mostly dichotomously branched, the cymes
10 cm wide or less. Pedicels 1 to 2 mm long. Calyx cup-
shaped, 2 mm long, somewhat pubescent externally, with 5, short,
broad, rather sharp teeth. Corolla 5 mm long, glandular exter-

PHILIPPINE PLANTS, IX. 343

nally, the tube somewhat enlarged upward, the lobes 5, oblong,
blunt, about 2 mm long. Filaments exserted; anthers 2 mm
Jong, more or less glandular.

CEBU, Cuming 1778.

Duplicates of the above number were referred by Schauer to Callicarpa
pentandra Roxb.—Geunsia farinosa Blume, but Sir Joseph Hooker in his
“Flora of British India” under Geunsia farimosa Blume, says “Cuming’s No.
1773, reduced to G. farinosa by Schauer, is probably, as stated in Gen. Pl.
2, p. 1150, a good species.”

It seems to me to be much more distinct from Blume’s species than is
Geunsia cumingiana (Schauer) Rolfe, which Hooker thinks is perhaps not
distinct from G. farinosa Blume. In this genus, as in Callicarpa, the amount
of pubescence on different forms seems to vary considerably, but Geunsia
hookeri, above described, is distinguished from the previously described
forms especially by its very scanty pubescence, which by no means covers the
lower surface of the leaf, as in G. farimosa, and in G. cuwmingiana.

GEUNSIA CUMINGIANA (Schauer) Rolfe in Journ. Linn. Soc. Bot. 21
(1884) 315.

SAMAR, Cuming 1707 (cotype). NEGROS, Cadiz, For. Bur. 12444 Danao,
March, 1908. LEYTE, Palo, Hliner 7368, January, 1906. MINDANAO, District
of Zamboanga, Tetuan, Ahern 378: Province of Misamis, Mount Malindang,
For. Bur. 4712 Mearns & Hutchinson, May, 1906.

Endemic.

VITEX L.
VITEX NITIDA sp. nov.

Arbor circiter 5 m alta, partibus junioribus floribusque plus
minusve pubescentibus; foliis tri- vel quinquefoliolatis, foliolis
nitidis, elliptico-ovatis, obtusis vel obscure acuminatis, usque
ad 10 cm longis, 2 basilaribus multo minoribus; cymis pedun-
culatis, axillaribus, folia subaequantibus; floribus ut videtur
albidis, calycibus truncatis, subglabris, corolla extus dense pu-
bescente.

A tree about 5 m high, the branches light-gray, smooth, some-
what 4-angled, the growing parts black when dry, more or less
fulvous-pubescent, becoming glabrous or nearly so. Leaves 3-
and 5-foliolate on the same branches, their petioles 5 to 7 cm
long. Leaflets elliptic-ovate, dark-colored when dry, shining,
glabrous or nearly so, entire, base acute or acuminate, apex
obtuse or shortly and broadly acuminate, the central one 7 to
10 cm long, the two next below somewhat smaller, the basal pair,
when present, much reduced, 2 to 3.5 em long, all petiolulate.

Cymes axillary, solitary, in the upper axils, forming a terminal
leafy inflorescence, a little shorter than the leaves, their pedun-

cles 2 to 6 cm long, very slightly pubescent or glabrous, dicho-
tomous, eventually rather lax, comparatively few-flowered.
Flowers white, according to the collector, in triads on the ulti-

344 MERRILL.

mate branchlets, the middle one of each triad sessile or sub-
sessile, the two lateral ones pedicelled; bracts triangular-ovate,
acute, 1.5 mm long, deciduous. Calyx cup-shaped, about 2.5 mm
long, very slightly appressed-pubescent, truncate or very ob-
securely 5-toothed. Corolla-tube about 6 mm long, villous within,
outside densely brown-pubescent above, the upper lip 2-lobed, the
lobes oblong-ovate, 3.5 to 4 mm long, the lower lip much larger,
3-lobed, the middle lobe orbicular, 5 mm in diameter, the two
lateral ones oblong-ovate, obtuse, 3 mm long, both lips densely
brown-pubescent on both surfaces. Filaments villous below.
Style 10 mm long.

MINDANAO, Province of Misamis, Tangob, For. Bur. 19546 Klemme, May
7, 1911, growing near the mountains, locally known to the Visayans as
tugaspan or sasalit.

A species well characterized by its 3- and 5-foliolate leaves, the lower
two leaflets, when present, much reduced, its axillary, peduncled cymes, trun-
cate calyx, and densely pubescent corolla. It is most closely allied to
Vitex pentaphylla Merr., but is apparently sufficiently distinct from that
species.

LABIATAE.

COLEUS Lour.

COLEUS AMBOINICUS Lour. FJ. Cochinch. (1790) 372.

Marrubium album Amboinicum Rumph. Herb. Amboin. 5 (1747) 294,
£102, Ff. 2.

Coleus aromaticus Benth. in Wall. Pl. As. Rar. 2 (1831) 15, Lab. Gen.
Sp. (1832-36) 51, DC. Prodr. 12 (1848) 72; Hook. f. Fl. Brit.
Ind. 4 (1885) 635; F.-Vill. Novis. App. (1880) 163.

Coleus suganda Blanco FI. Filip. (1887) 483, ed. 2 (1845) 337; Mig.
Fl. Ind. Bat. 2 (1856) 948.

Luzon, Manila, Merrill 7568, sterile specimen.

This species is found in the Philippines only in cultivation, and is locally
known by the Tagalog name suganda, and the Spanish name orégano. Its
probable origin is the Malayan region, and it is certainly of prehistoric
introduction in the Philippines. The species very rarely produces flowers
in the Philippines, a fact already noted by Blanco. Although this species
has been known to me many years I have never seen a flowering specimen
and hence have hesitated in considering it. Material of the above number
was sent to Kew for comparison, eliciting the statement that the leaf
specimens supplied agree with Ceylon material of Coleus amboinicus, and
with the figure given by Rumphius. Although no Philippine flowering
material is available I consider the specimens certainly to represent Blanco’s
Coleus suganda, which species is identical with Coleus aromaticus Benth.=
Coleus amboinicus Lour., a reduction already made by F.-Villar. Coleus
amboinicus Lour. was based in part on botanical material from plants
cultivated in Cochichina, and on several references to pre-Linnean liter-
ature including the description and plate given by Rumphius, which he
erroneously cites as I. 8 c. 75. tab. 72, the reference apparently taken from

PHILIPPINE PLANTS, IX. 845

the index in the last volume of Rumphius’ work. So far as sterile material
is comparable with the figure, I consider Rumphius’ delineation to be an
excellent representation of the Philippine plant. Coleus amboinicus Lour.
is the type of the genus.

MESONA Blume.
MESONA CLAUSA sp. nov.

Herba erecta, stricta, vix ramosa, circiter 30 cm alta, caulibus
foliis inflorescentiisque plus minusve dense pallide fulvo-villosis ;
foliis petiolatis, oblongis, usque ad 6 cm longis, obtusis vel
acutis; inflorescentiis interruptis circiter 15 cm longis, verticillis
densis, subglobosis; calycibus sub fructu plus minusve inflatis,
8 mm longis, 2-labiatis, labiis integris, inferioribus orbicula-
ribus os calycis claudentibus, superioribus oblongis.

An erect unbranched herb about 30 cm high. Stems rather
stout, obscurely angled, rather densely covered with pale-fulvous,
villous hairs. Leaves oblong or narrowly oblong, 5.5 to 6 cm
long, 1 to 1.5 em wide, apex acute or obtuse, base acute, the
margins somewhat crenulate, both surfaces more or less villous,
especially on the nerves and midrib; nerves ascending, about
8 on each side of the midrib; petioles 5 to 10 mm long, those
of the basal leaves sometimes 2 cm in length. Inflorescence
interrupted, about 15 cm long, the whorls dense, globose, 2 to
2.5 cm apart, the upper ones closer, about 1.5 cm in diameter,
the rachis densely villous, the bracts subtending the whorls
oblong-ovate, brown, reflexed, more or less villous, acuminate,
0.8 to 1.5 cm long, the lowermost ones subfoliaceous. Calyx just
after anthesis 3 mm long, densely villous, in fruit accrescent, 8
mm long, slightly inflated in the middle, the mouth oblique, closed
by the orbicular lower lip which is 2 mm in diameter and
villous on the outside, entire, the upper lip oblong, 1.5 mm long,
obtuse, not at all lohed or toothed, villous on the outside. Co-
rolla unknown.

CULION, in damp, open grass lands, Merrill 460, December 13, 1902.

A species well characterized by its entire calyx-lips, the lower one
orbicular, appressed to and quite closing the mouth of the calyx-tube. I
have found on the specimen a single badly withered corolla showing the four
stamens, the upper two filaments appedanged at the base, characteristic of
the genus.

POGOSTEMON Desf.

POGOSTEMON CABLIN (Blanco) Benth. in DC. Prodr. 12 (1848) 156;
Mip. Fl. Ind. Bat. 2 (1856) 964; F.-Vill. Novis. App. (1880) 164;
Vid. Rev. Pl. Vase. Filip. (1886) 213; Prain in Journ. As. Soc.
Beng. 74? (1907) Extra Number 708, Kew Bull. (1908) 78.
Mentha cablin Blanco FI. Filip. (1837) 472.

346 MERRILL.

Pogostemon patchouly Pellet. in Mém. Soc. Se. Orléans 5 (1845) 277,
boas

Mentha auricularia Blanco 1. c. ed. 2 (1845) 329, non Linn.

Pogostemon suavis Ten. in Giorn. Bot. Ital. 2 (1847) 56.

Pogostemon patchouli Hook. Kew Journ. Bot. 1 (1849) 328, t. 11.

Pogostemon patchouli var. suavis Hook. f. Fl. Brit. Ind. 4 (1885) 6384.

Pogostemon heyneanus Merr. in Philip. Journ. Sci. 2 (1907) Bot. 426,
non Benth.

The above is the nomenclatural history of this interesting plant, which
is represented by the following Philippine specimens: LuzoN, Province of
Cagayan, For. Bur. 16470 Bacani, January, 1909: Bontoc Subprovince,
Bauco, Vanoverbergh 1057, December, 1910, in gardens, Ilocano “cablin”:
Province of Pampanga, Mount Arayat, Merrill 5025, on dry, steep, talus
slopes, altitude about 400 m, February, 1906: Province of Laguna, Vidal
505 in Herb. Kew.: Province of Rizal, Bosoboso, Bur. Sci. 2060 Ramos,
February, 1907; Montalban, For. Bur. 2442 Ahern’s collector, January, 1905;
Antipolo, Bur. Sci. 5199 Ramos, June, 1908, in gardens, Tagalog “cadling’’:
Manila, Bur. Sci. 299 Merrill, February, 1910, in gardens: without definite
locality, Loher 4211.

Doctor Prain has recently shown ™ that the patchouli of commerce is the
product of Pogostemon cablin (Blanco) Benth. (P. patchouli Pellet., non
P. patchowli Dalz. & Gibs), and that the specific name given to the plant by
Blanco has priority over the other proposed ones, and is hence the proper one
for the species. I had previously confused it with Pogostemon heyneanus
Benth., and recorded it from the Philippines under the latter name."

Pogostemon cablin (Blanco) Benth., is of wide distribution in the
Philippines, more commonly in cultivation only, but occasionally found
growing wild (Mount Arayat, Bosoboso, and Montalban). There is no
evidence that it is a native of the Philippines, but it has unquestionably
been introduced, although probably in prehistoric times. Whatever its
origin it has found favorable conditions for growth and reproduction in
the Philippines, for here it flowers freely and has become spontaneous in
some regions. In contrast to this, in India and in the Malay Peninsula,
where it is extensively cultivated, it very rarely produces flowers. In the
Philippines the plant is not now, and apparently never has been, cultivated
on a commercial scale, but as a cultivated plant one finds only scatered
individuals in native gardens. Dr. Leon Guerrero informs me that it is
claimed that if a few leaves of this plant be enclosed in a book, the small
beetles whose larvae are frequently so destructive to bindings in tropical
countries will not attact bindings of volumes so protected.

POGOSTEMON HEYNEANUS Benth. in Wall. Pl. As. Rar. 2 (1831) 16;
DC. Prodr. 12 (1848) 153; Wight Icon. t. 1440; Prain in Journ.
As. Soc. Beng. 74? (1907) Extra Number 707.
Pogostemon patchowli Dalz. & Gibs. Bomb. Fl. Addend. (1861) 66,
non patchouly Pellet.
Pogostemon patchouli Hook. f. Fl. Brit. Ind. 4 (1885) 633, non P.
patchouly Pellet.

* Journ. As. Soc. Beng. 74? (1907) Extra Number 708, Kew Bull. (1908)
78.
“This Journal 2 (1907) Bot. 426.

PHILIPPINE PLANTS, IX. 347

MINDANAO, District of Zamboanga, near Zamboanga, Merrill 5467, October
1906, in coconut groves along small streams; Basagan, Merrill 8238, De-
cember, 1911, in waste places near houses. PALAWAN, near Puerto Princesa,
Bur. Sci. 287 Bermejos, January, 1905, locally known as cadlum.

This species, like the preceding, is certainly not a native of the Philip-
pines, but has been introduced from some other part of Malaya. It appears
to be less common than Pogostemon cablin, or at any rate more restricted
in its distribution. It is found in waste places in and near settlements,
and is thoroughly naturalized; it is doubtless also cultivated, but I have
seen no cultivated specimens.

POGOSTEMON NEPETOIDES Stapf in Kew Bull. (1908) 116.

This species was described from a specimen collected by Micholitz in
the Philippines, no definite locality being given. Dr. C. B. Robinson, who
has kindly examined the type in the Kew Herbarium informs me that it
seems to be well matched by a specimen collected in Ilocos Norte, Luzon, by
Merritt and Darling, For. Bur. 12479, although the specimens were not quite
comparable, the type being more mature. So far as the specimen goes,
For. Bur. 12479 Merritt & Darling agrees with Stapf’s description, and
doubtless represents his species. It has much the appearance of Pogos-
temon cablin Benth., but differs in being less pubescent, its leaves scarcely
lobed, and its panicled spikes much more numerous and smaller than in
Bentham’s species.

Var. GLANDULOSUS var. nov.

A typo differt calycibus minoribus, 3.5 ad 4 mm longis,
glandulosis.

Luzon, Province of Rizal, Bosoboso (San Isidro), Bur. Sci. 1849, 2130,
18406 (type) Ramos, January, 1907, February, 1911, the specimens in-
dicated as growing in forests.

This form may eventually prove to be a distinct species, but it appears
to be closely allied to Stapf’s Pogostemon nepetoides, which it closely re-
sembles. It differs at least in that its calyces are shorter, 3.5 to 4 mm
long, and distinctly glandular with waxy glands. The plant is aromatic,
with about the odor of Pogostemon cablin Benth., but much weaker. No
native names are recorded.

POGOSTEMON MEMBRANACEUS sp. nov.

Herba erecta, ramosa, erecta, 50 cm alta vel ultra, subglabra
vel plus minusve puberula; foliis membranaceis, ovatis, nitidis,
basi rotundatis, apice acuminatis, margine distincte crenato-
serratis, usque ad 9 cm longis, subtus glandulosis, ad costa ner-
visque puberulis; inflorescentiis terminalibus, solitariis, cylin-
draceis, usque ad 15 cm longis, 1.5 ad 2 cm diametro, anguste
paniculatis, spiciformibus; verticillastris subapproximatis, in-
ternodiis 5 ad 10 mm longis; floribus 5-meris, calycibus circiter
6.5 mm longis.

An erect branched herb 50 cm high or more, subglabrous,
or more or less puberulent. Branches subterete or slightly

A

348 MERRILL.

angled, brownish, slightly pubescent or puberulent. Leaves
ovate, membranaceous, 5 to 9 em long, 2.5 to 5 cm wide, shining,
nearly glabrous, the base rounded or subacute, the apex acu-
minate, margins distinctly and usually doubly crenate-serrate,
the lower surface distinctly glandular-punctate, somewhat pube-
rulent on the midrib and nerves, the nerves distinct, the reti-
culations lax, not prominent; petioles 1 to 3 cm long, puberulent.
Inflorescence a terminal, solitary, cylindric, spiciform panicle
6 to 15 cm long, 1.5 to 2 cm in diameter, somewhat pubescent
with short hairs, the verticels subapproximate, 5 to 10 mm
apart, each with from 12 to 25 flowers racemosely arranged on
the short spreading branches, the bracts none or very early de-
ciduous. Calyx shortly pedicelled, oblong, acute at both ends,
5-angled, sparingly pubescent, glandular-punctate, 6 to 7 cm long,
the teeth 5, lanceolate, slenderly acuminate, 1.5 mm long. Co-
rolla slender, glabrous, 10 mm long, the tube very slender below,
6 to 7 mm long, more or less funnel-shaped above; lower lip
lanceolate, acuminate, entire, about 3 mm long, 1 mm wide,
3-nerved; upped lip much larger, 3 mm wide, 3-lobed, the middle
lobe oblong, obtuse, about 1.8 mm long, 0.8 mm wide, 3-nerved,
the two lateral ones broadly ovate, rounded, about 1.2 mm long
and wide, usually 4-nerved. Filaments slender, exserted, bearded
below the middle. Nutlets oblong-ovoid, black, shining, smooth,
obscurely 3-angled, 0.8 mm long.

Luzon, Province of Laguna, Capuan, Bur. Sci 16419 Ramos, August 20,
1910, near streams, flowers pink.

Distinct from all other known Philippine forms, characterized by its
thin, nearly glabrous leaves and its spiciform narrow panicles.

POGOSTEMON RETICULATUS sp. nov.

Herba erecta, aromatica, ramosa, 50 cm alta vel ultra, ciliata;
foliis membranaceis, late ovatis, usque ad 12 cm longis, obtusis
vel late breviter obtuse acuminatis, basi distincte cordatis, mar-
gine grosse irregulariter crenatis, subtus nervis reticulisque
laxis prominentibus; inflorescentiis terminalibus, spiciformibus,
verticillastris inferioribus remotis, superioribus approximatis;
calycibus 6 mm longis, 5-dentatis, minute puberulis, parcissime
ciliatis, bracteis filiformibus.

An erect, branched, apparently annual herb 50 cm or more
in height, all parts distinctly ciliate with long, weak, scattered,
spreading, white hairs, and also more or less puberulent.
Branches obscurely angled, dark-colored when dry. Leaves op-
posite, long-petioled, broadly ovate, membranaceous, 5 to 12 cm
long, 3.5 to 9 em wide, obtuse or very shortly and obtusely

PHILIPPINE PLANTS, IX. 349

acuminate, the base broadly rounded, prominently cordate, the
sinus 1 to 1.5 cm deep, the margins coarsely and irregularly
crenate, both surfaces ciliate, the hairs on the upper surface
scattered, those on the lower surfaces mostly on the midrib and
nerves; nerves and reticulations lax, prominent, dark-colored in
contrast to the grayish lower surface of the leaf; petioles 5
to 8emlong. Inflorescence of terminal, solitary, long-peduncled,
spiciform racemes, these, when young, and including the pe-
duncles, 6 to 14 cm long, apparently much longer when mature,
the verticels with from 6 to 8 flowers, the lower ones distant,
the internodes up to 2 cm in length, the upper ones approximate,
the bracteoles filiform, 2 mm long, puberulent, very sparingly
ciliate. Flowers 5-merous, subsessile or shortly pedicelled.
Calyx 6 mm long, 5-angled, minutely puberulent and very spar-
ingly ciliate toward the upper end, narrowed at both ends,
5-toothed, the teeth oblong-lanceolate, acuminate, margins mi-
nutely ciliate. Mature corollas not seen, but from buds with
the characteristic lobing and with the characteristic stamens of
the genus.

LuZON, Province of Rizal, Montalban, For. Bur. 3395 Ahern’s collector
(type), November, 1905. Apparently also represented by Loher 3395 from
Angat, Province of Bulacan, Luzon, in the Kew Herbarium.

This species has the characteristic odor of Pogostemon cablin Benth.,
and bears the same native name as that species. It is, however, not at all
closely allied, being entirely different in its vegetative characters, its
indumentum, and in its inflorescence. It is well characterized by its thin,
coarsely crenate, broadly ovate leaves which are prominently and laxly
reticulate, and by all parts being sparingly ciliate and more or less minutely
puberulent.

SCUTELLARIA L.

SCUTELLARIA COPELANDII sp. nov. § Stachymachris.

Herba usque ad 30 cm alta, plus minusve puberula vel leviter
pubescens; foliis ovatis, obtusis, usque ad 4 cm longis, basi
late rotundatis, truncatis, vel leviter cordatis, margine crenatis;
racemis terminalibus, brevibus; floribus oppositis, circiter 13
mm longis.

A perennial herb, slightly branched, the stems slender, terete
or obscurely angled, often prostrate below, puberulent or slightly
pubescent, pale when dry. Leaves ovate, 1 to 4 cm long, 1 to
2.7 cm wide, membranaceous, of about the same color on both
surfaces when dry, the apex obtuse, base broadly rounded,
subtruncate, or somewhat cordate, both surfaces minutely pube-
rulent or the lower one somewhat ciliate-pubescent on the nerves,
the lower surface minutely glandular, the margins crenate; pe-

350 MERRILL.

tioles puberulent, slender, 1 to 2 cm long. Racemes terminal,
puberulent, comparatively few-flowered, 2to5cmlong. Flowers
opposite, pale-blue; pedicels puberulent, 2 to 3 mm long; brac-
teoles oblong-spatulate, obtuse, about 2mm long. Calyx slightly
puberulent, the lips rounded, about 2.2 mm long, the appendage
convex, nearly 2 mm in diameter, accrescent and in fruit 5 to 6
mm in diameter. Corolla 12 to 13 mm long, enlarged upward,
slightly curved, slightly ciliate on the outside. Nutlets about 1.6
mm long, minutely verruculose.

MINDANAO, District of Cotabato, Weber 1512 (type), August, 1911, on
damp rocky banks of streams, altitude about 135 m: District of Zamboanga,
Sax River, Copeland 1597, February, 1905, Merrill 8210, November, 1911,
on cliffs and boulders along streams, altitude 100 to 150 m.

A species manifestly allied to Scutellaria indica Linn., differing especially
in its stems, leaves, etc. being slightly puberulent, not villous, and its
somewhat smaller flowers.

SOLANACEAE.
SOLANUM L.

SOLANUM EPIPHYTICUM sp. nov.

Frutex epiphyticus, glaber vel subglaber, simplex vel leviter
ramosis, ramis griseis, nitidis, teretibus, in siccitate plus minusve
suleatis vel angulatis; foliis membranaceis, geminis, altero
oblongo , acuminato, usque ad 17 cm longo, altero ovato, haud 5
cm longo; floribus 5-meris, axillaribus, 1 ad 5 fasciculatis, pedi-
cellis 1 ad 2 cm longis; calycis truncatis; fructibus globosis,
seminibus 2 vel 3, obovatis, minute rugosis.

An epiphytic glabrous: or subglabrous, simple or slightly
branched shrub usually less than 1 min length. Stem pale-gray,
somewhat shining, stout, when dry somewhat striate or angled,
often a little zigzag, in life somewhat fleshy. Leaves membra-
naceous, shining, in pairs at each node, one very much larger and
of different shape than the other: Larger leaves of each pair
oblong, 10 to 17 cm long, often slightly faleate, acuminate, base
strongly inequilateral, acute or acuminate; nerves 5 to 8 on each
side of the midrib, curved-ascending, anastomosing, the reticula-
tions indistinct; petioles 1 to 1.5 emlong. Smaller leaves of each
pair ovate, 2 to 5 em long, shortly and abruptly acuminate, acute,
or obtuse, base usually rounded, subequilateral; petioles 2 to 5
mm long. Flowers axillary, solitary, or in fascicles of from 2 to
5, pedicelled, the pedicels 1 to 2 cm long, slender, slightly thick-
ened upward. Calyx cup-shaped, truncate, not at all toothed,
about 2mm long. Corolla white, 6 to 7 mm long, the tube short,

PHILIPPINE PLANTS, IX. 351

the lobes oblong-lanceolate, acute or somewhat obtuse, about 4
mm long. Stamens 5, equal; anthers 2 mm long. Style 5 mm
long. Berry globose, fleshy, at first white, becoming orange or
red when mature, 5 to 6 mm in diameter, containing but 2, rarely
3 seeds, the seeds obovate, somewhat compressed, about 5 mm
long, pale, minutely pitted-rugose.

LuzON, Province of Albay, Cuming 873 (type): Province of Cagayan,
Bur. Sci. 7482 Ramos: Province of Tayabas, Lucban, Elmer 7494. MIN-
poro, Alag River, Merrill 6157; South of Lake Naujan, For. Bur. 6895
Merritt. MINDANAO, Butuan Subprovince, Merrill 7307, Weber 1207: Dis-
trict of Davao, Copeland 329: Lake Lanao, Mrs. Clemens 428, s. n.

This species is well characterized by its epiphytic habit, its pale-gray,
more or less thickened stems, its leaves in very unequal pairs, its solitary
or fascicled, pedicelled, 5-merous flowers with truncate calyces, and its 2-
or 3-seeded berries. It has been determined at times as Solanum blumei
Nees, and at other times as S. parasiticum Blume. It is, from its habit,
a very different plant from Solanum blumei Nees, and is at once dis-
tinguished from S. parasiticum Bl. by its geminate, not solitary leaves.
SOLANUM PHILIPPINENSE sp. nov.

Herba annua, erecta, glabra, dichotoma, inermis, ramis tere-
tibus, in siccitate plus minusve sulcatis; foliis geminis, membra-
naceis, acuminatis, basi decurrento-acuminatis, ovatis vel oblong-
ovatis, altero subdimidio minore; pedunculis filiformibus, uni-
floris, axillaribus, solitariis geminis ternisve; calycis truncatis,
integerrimis.

An annual, erect, glabrous, dichotomously branched herb 1 m
high or less, the branches terete, more or less sulcate when dry.
Leaves membranaceous, entire, in pairs at each node, the smaller
of each pair half as large as the other or smaller, of the same
shape, acuminate, base decurrent-acuminate, the larger ones 7 to
12 cm long, 3.5 to 6 cm wide, their petioles up to 2 cm in length,
the smaller ones of each pair 2 to 5 cm long, short-petioled.
Pedicels axillary, slender, solitary, in pairs, or in threes, 1.5 to 3
cm long, somewhat thickened upward, 1-flowered. Flowers pale-
yellowish. Calyx cup-shaped, about 3 mm long, 3 mm in diame-
ter, truncate, not at all toothed. Corolla 6 to 7 mm long, the lobes
ovate, 2 to 3 mm long, acute or somewhat obtuse. Stamens
equal; anthers broadly ovoid, about 1.2 mm long. Fruit globose,
fleshy, red when mature, about 7 mm in diameter. Seeds num-
erous, pale, somewhat translucent, orbicular-elliptic, flattened,
smooth, about 1.2 mm long.

LuZON, Benguet Subprovince, Mount Tonglon (Santo Tomas), Elmer
6561, June, 1904, Williams 1275 (type) July 1, 1904, Bur. Sci. 5406 Ramos,
December, 1908, Merrill 8003, May, 1911. MINDANAO, Sax River Mountains
back of San Ramon, Merrill 8164, November, 1911.

352 MERRILL.

This species is common along the trail through the mossy forest on
Mount Tonglon, above an altitude of 1,700 m; in Mindanao it grows along
small streams in very damp shaded ravines at an altitude of about 800 m.,

It is allied to Solanum blumei Nees, and some of the specimens cited above
have been determined as that species, and the duplicates so distributed.
Among other characters it differs in its habit, being an erect, dichotomously
branched herb, not a suffrutescent or woody plant, with longer petioles
than in Nees’ species. It is not certain that the plant described as Solanum
blumei by Prain in Journ. As. Soc. Beng. 747 (1905), Extra Number 330,
is the same as S. blumei Nees, as Nees describes the calyx as entire and
truncate, while Prain describes it as subtruncate, with 5 subulate and 5
intervening minute teeth.

BIGNONIACEAKE.

RADERMACHERA Hassk.
RADERMACHERA WHITFORDII sp. nov.

Arbor parva, glabra, novellis resinosis; foliis pinnatis, usque
ad 35 ecm longis, foliolis 5 vel 7, subcoriaceis, oblongo-ellipticis,
acuminatis, basi acutis, 11 ad 15 ecm longis; paniculis quam folia
longioribus, diffusis, floribus 2.5 cm longis.

A small tree, glabrous throughout, the young parts more or
less resinous. Ultimate branches somewhat compressed, with
few, large, scattered lenticels. Leaves simply pinnate, 25 to 35
em long, the basal part of the petiole more or less lenticellate.
Leaflets 5 to 7, oblong-elliptic, subcoriaceous, subequally narrowed
at both ends, the base acute, apex acuminate, 11 to 15 cm long,
3.5 to 6 cm wide; lateral nerves spreading, about 12 on each side,
the reticulations lax; petiolules of the lower leaflets 1.5 to 2 cm
long, of the upper ones half as long or shorter. Panicles longer
than the leaves, up to 40 cm in length, rather lax, open, the
branches distant, spreading, the lower ones up to 15 cm long.
Flowers rather few, 2.5 em long, the corolla slightly pubescent
externally in the upper part. Calyx not ribbed. Immature
follicles 25 em long, 4mm wide, somewhat compressed.

MINDANAO, District of Cotabato, Lebak, For. Bur. 11817 Whitford, March
7, 1912, in dry river bottoms at low altitudes.

Well characterized by its simply pinnate leaves; among the Philippine

species most closely allied to Radermachera elliptica Merr., but with very
different leaves and inflorescence, and smaller flowers.

CUCURBITACEAE.
ALSOMITRA Roem.

ALSOMITRA SARCOPHYLLA (Wall.) Roem. Syn. 2 (1846) 118; Cogn.
in DC. Monog. Phan. 3 (1881) 929.
Zanonia sarcophylla Wall. Cat. (1831) no. 3724, nomen, Pl. As. Rar.
2 (1831) 28, t. 138.
LUZON, Province of Bataan, Lamao River, Copeland 255, January, 1904.

PHILIPPINE PLANTS, IX. 353

Not previously reported from the Philippines: Siam, Burma, and Timor.
The specimen collected by Doctor Copeland has staminate flowers, and
while it agrees with the description of Alsomitra sarcophylla Roem., very
closely, still with the lack of pistillate flowers and fruits the identity of the
Philippine form with the above species can not be considered to be certain.

ALSOMITRA INTEGRIFOLIOLA (Cogn.) Hayata in Journ. Col. Sci. Tokyo
Sor Genre
Gynostemma integrifoliolum Cogn. in DC. Monog. Phan. 3 (1881) 916.
Alsomitra clavigera F.-Vill. Novis. App. (1880) 98 (probably), non
Hook. f.
Gynostemma elongatum Merr. in Philip. Journ. Sci. 3 (1908) Bot. 267.

LUZON, Benguet Subprovince, Bued River, Merrill 4312, with staminate
flowers; Sablan, Bur. Sci. 12675 Fénix, with staminate flowers, Phil. Pl.
458 Fenix, with mature fruits: Province of Rizal, Bosoboso and Montalban,
For. Bur. 1897, 3409 Ahern’s collector, with staminate flowers: Province
of Bataan, Elmer 6694, Merrill 1527, For. Bur. 2425 Meyer, all with
staminate flowers, For. Bur. 5474 Curran (type of Gynostemma elongatum
Merr.), with pistillate flowers: Province of Laguna, Calauan, Bur. Sci.
12460 McGregor, with staminate flowers: Province of Tayabas, Cuming
767 (cotype of Gynostemma integrifoliolum Cogn.), with staminate flowers.

I concur with Hayata in the transfer of Gynostemma integrifoliolum Cogn.
to Alsomitra, for the mature fruits of Gynostemma elongatum Merr., which
is certainly conspecific with G. integrifoliolum Cogn., are typical of Alsomi-
tra, not of Gynostemma.

Cogniaux, as noted by Hayata, has described the fruits of Gynostemma
integrifoliolum as brown, glabrous or slightly puberulent, 7 to 8 mm thick.
On several of our specimens are what I had previously taken to represent
immature fruits of Cogniaux’s species. These organs are ovoid or sub-
globose, 1 cm long or less, and closely simulate very immature fruits. An
examination of them, however, shows that they are globose or ovoid galls,
crowned by the calyx, and usually also the stamens of the pistillate flowers;
they usually contain numerous larvae. It is strongly suspected that Cog-
niaux saw similar galls on one of Cuming’s specimens examined by him,
and mistook them for immature fruits.

Alsomitra integrifoliola Hayata is closely allied to A. clavigera Hook. f.,
but has smaller fruits, in our specimens 4 to 5 cm long and about 1 cm in
diameter, and the seeds are quite glabrous, not at all muricate.

A species known only from Luzon and Formosa.

GOODENOVIACEAKE.
SCAEVOLA L.

SCAEVOLA FRUTESCENS (Mill.) Krause in Engl. Pflanzenreich 54
(1912) 125.

Lobelia frutescens Mill. Gard. Dict. ed. 8 (1768) No. 1, p. p.
Whether or not Doctor Krause is correct in his interpretation of Lobelia
frutescens Mill., it is probably best to follow him and accept the above
specific name for this common and widely distributed species. Miller’s
Lobelia frutescens was a mixture, based on two references, Fl. zeyl. 313,
which is the same as Scaevola koenigii Vahl”, and Lobelia frutescens por-

*Trimen Fl. Ceyl. 3 (1895) 54.

354 MERRILL.

tulacae folio Plum. Nov. Gen. 21, which is manifestly Scaevola plumieri
(L.) Vahl, where it is referred by Krause. The seeds mentioned by Miller
as having been sent to England from the Bahamas by Mr. Catesby must
have been of the latter species, S. plumieri, not of S. koenigii Vahl, for
S. koenigti Vahl is not found in the western hemisphere. If the first ref-
erence be taken as the type, then Krause is correct in adopting Miller’s
specific name for the present species; if on the other hand the first ref-
erence to a figure be taken as the type, then Lobelia frutescens Mill., is a
synonym of Scaevola plumieri Vahl.

W. F. Wight” in applying the principle of generic types has decided that
the plants referred by botanists generally to Scaevola should be referred to
Lobelia, as the type of the genus Lobelia is a Scaevola. While this may
be the logical outcome of the application of the principle in this case, yet
the nomenclatural confusion that this course of procedure entails is so
great that it is not considered probable that Mr. Wight’s proposition will
receive the support of any considerable number of botanists.

Scaevola frutescens (Mill.) Krause is represented by the following Phil-
ippine material:

BABUYANES ISLANDS, Camaguin, Bur. Sci. 4100 Fénix; BATANES ISLANDS,
Batan, Bur. Sci. 3643 Fénix. LuzoN, Province of Zambales, Merrill 2095:
Province of Pangasinan, For. Bur. 8864 Curran & Merritt: Province of
Tayabas, Whitford 897, For. Bur. 3208 Hagger: Province of Camarines,
Ahern 229. Apo ISLAND, Mindoro Strait, Merrill 416. Busuanca, Phil.
Pl. 405 Merrill. PowuitLto, Bur. Sci. 6950 Robinson. PALAWAN, Bur. Sci.
617 Foxworthy. BAuaBAc, Bur. Sci. 384 Mangubat. MASBATE, Merrill
3043. BASILAN, For. Bur. 3997 Hutchinson. MINDANAO, District of Davao,
Elmer 12083.

The above form is the most common one in the Philippines and has
glabrous or only slightly pubescent leaves. The corolla externally and
the fruits are quite glabrous.

A form with nearly or quite glabrous leaves, the corolla, externally,
and the fruits more or less hirsute is less common and is represented
by the following specimens:

Luzon, Province of Zambales, Sandkuhl, November, 1909: Province of
Pangasinan, Bur. Sci. 4897 Ramos. PALAWAN, For. Bur. 3774 Curran.
MINDANAO, District of Davao, Piper 405, Copeland 674.

This form appears to be intermediate between the glabrous plant,
which I take to be typical Scaevola frutescens as interpreted by Krause,
that is, the Ceylon form, and Scaevola sericea Forst., which Krause
has sunk with numerous other synonyms under Scaevola frutescens. I
consider that Forster’s plant is at least worthy of varietal rank.

Var. SERICEA (Forst) comb. nov.

Scaevola sericea Forst. Prodr. (1786) 504.

LUZON, without definite locality, Haenke in Herb. Prague: Province of
Ilocos Sur, Merrill 339, June, 1912. Volkens 133, from the Island of Yap,
Carolines, is identical.

This variety is distinguished from the more common and_ nearly
glabrous form by its leaves being uniformly and rather softly pubescent
on both surfaces, the inflorescences, flowers externally, and the fruits
also pubescent.

* Contr. U. S. Nat. Herb. 9 (1905) 310.

PHILIPPINE PLANTS, IX. 355

COMPOSITAE.

GYNURA Cass.
GYNURA PIPERI sp. nov.

Species G. sarmentosae DC. simillima et ut videtur valde affi-
nis, differt foliis utrinque ramulis inflorescentiisque plus minusve
dense crispato-pubescentibus.

A scandent herbaceous vine the ultimate branches 2 to 3 mm
in diameter, reddish-brown, slightly striate, glabrous, the younger
parts sparingly crisped-pubescent with short, weak hairs.
Leaves chartaceous or subcoriaceous, oblong to oblong-lanceolate,
entire or rarely distantly and irregularly toothed, oblong to
oblong-lanceolate, acuminate, base obtuse or acute, 3 to 9 cm long,
1.5 to 2.5 cm wide, the upper ones much reduced, the uppermost
scarcely 2 cm in length and bract-like, both surfaces rather
densely and uniformly pubescent with short, rather pale or
brownish, weak, crisped hairs; lateral nerves about 3 on each
side of the midrib, obscure; petioles 3 to 6 mm long, densely pubes-
cent. Inflorescence terminal, peduncled, lax, crisped-pubescent,
the heads 7 to 10, slenderly peduncled, oblong, in anthesis about
14mm long. Involucral bracts linear, about 10 mm long, acute,
sparingly pubescent on the back.

Siquisgor, C. V. Piper 384, May 9, 1911.

A species manifestly closely allied to Gynura sarmentosa DC., but
distinguished at once by its entire or subentire leaves and by its charac-

teristic indumentum.
EUPATORIUM L.

EUPATORIUM JAPONICUM Thunb. FI. Jap. (1784) 308; Forbes & Hemsl.
in Journ. Linn. Soc. Bot. 23 (1888) 403.

BATANES ISLANDS, Batan, Santo Domingo de Basco, Bur. Sci. 3177
Mearns, May 27, 1907.

The specimen here referred to Hupatorium japonicum Thunb. has
leaves up to 12 cm in length and divided quite to the base into three
segments, even the upper leaves subtending the branches of the inflo-
rescence being deeply divided. It may be referabie to some other species,
but from the material available here for comparison, and from the de-
scriptions, I can see no reason for separating it from Thunberg’s species.

Japan to Formosa and southern China. A distinct northern type in
the Philippine flora.

EUPATORIUM CAMIGUINENSE sp. nov. § Eximbricata.

Herba scandens, caulibus teretibus, leviter pubescentibus;
foliis ovatis vel late oblong-ovatis, submembranaceis, usque ad 13
cm longis, basi late rotundatis, apice longe tenuiter acuminatis,
margine argute serrato-dentatis, utrinque leviter glandulosis, ad

costa nervisque pubescentibus; inflorescentiis terminalibus,
118111——-7

356 MERRILL.

corymboso-paniculatis; capitulis numerosis, 5-floris, squamis cir-
citer 12, 6 interioribus 6 mm longis, acuminatis, glabris, exterior-
ibus brevioribus, extus leviter pubescentibus.

A scandent herb, the stems brownish when dry, striate, terete,
slightly brownish-pubescent with short hairs, the younger parts
more densely so. Leaves opposite, ovate to broadly oblong-ovate,
submembranaceous, 9 to 13 cm long, 4 to 6 cm wide, the base
broadly rounded, the apex long and slenderly subcaudate-
acuminate, the margins rather coarsely serrate-dentate except
at the base and along the sides of the acumen which are entire,
both surfaces somewhat shining when dry, the upper one oliva-
ceous, the lower somewhat paler, both with scattered minute
glands, and shortly pubescent on the midrib and nerves; lateral
nerves about 5 on each side of the midrib, slender, distinct,
anastomosing; petioles densely pubescent, 1.5 to 2 cm long.
Inflorescence terminal, corymbosely paniculate, about 15 cm long,
the branches few, distant, spreading, opposite, the lower ones up
to 12 cm long, all parts densely brown-pubescent with short hairs.
Heads rather crowded on the ultimate branchlets, greenish-
white, their pedicels 1 to 3 mm long. Involucral bracts about
12, the outer 6 slightly pubescent externally, oblong-ovate to
oblong, obtuse, the outermost ones 1.5 to 2 mm long, the others
longer, the inner six oblong-linear, 6 mm long, 1 mm wide,
glabrous, acuminate, 3-nerved. Flowers 5 in each head. Corolla
white, 3.6 mm long, the teeth 5, oblong-ovate, acute, 0.5 mm long.
Style-arms exserted,3 mm long. Achenes 1.8mm long. Pappus
white, about as long as the corolla.

CAMIGUIN DE MINDANAO, Mount Mahinog, in forests, Bur. Sci. 14688
Ramos, April 11, 1912.

A species manifestly allied to the Luzon forms, Eupatorium sambuci-
folium Elm., E. toppingianum Elm., and FE. benguetense C. B. Rob., and
nearer the first than to either of the others. It is distinguished from
all by its much larger leaves which are differently shaped and with a
different venation.

BLUMEA DC.
BLUMEA BICOLOR sp. nov.

Herba erecta, simplex vel parce ramosa, omnibus partibus
leviter adpresse pilosis; foliis alternis, usque ad 16 cm longis,
chartaceis vel membranaceis, oblongo-ellipticis, apice acute acumi-
natis, margine irregulariter denticulatis, basi longe decurrente
acuminatis, supra subglabra, subtus leviter adpresse pilosis, pur-
purascens; paniculis circiter 15 cm longis, pedunculatis; capitulis
circiter 8 mm longis.

PHILIPPINE PLANTS, IX. 357

An erect, simple or sparingly branched herb 80 cm high or less,
the lower part of the stem naked, leafless. Stems terete,
brownish when dry, sparingly appressed-pilose. Leaves mostly
in the middle of the stem, somewhat crowded, alternate, oblong-
elliptic, 8 to 16 cm long, 3 to 6 cm wide, chrtaceous or membra-
naceous, the apex sharply acuminate, the base long-decurrent-
acuminate, margins irregularly and sharply denticulate, not at
all lobed; nerves 8 to 11 on each side of the midrib, curved-
ascending; upper surface of the leaf green, sparingly pubescent,
becoming glabrous or nearly so, the lower surface uniformly and
rather dark-purple, sparingly appressed-pilose. Panicles pedun-
cled, about 15 cm long, rather lax, pilose, the branches few, the
lower ones 7 cm long or less, the upper ones gradually shorter.
Heads rather few, somewhat scattered, peduncled, about 8 mm
long; peduncles slender, pubescent, 5 to 8 mm long, the bracteoles
few, scattered, linear. Involucral-bracts appressed-pubescent,
the outer few linear-lanceolate, acuminate, 1.5 to 2 mm long, the
inner ones gradually longer, the innermost up to 7.5 mm long,
about 0.5 mm wide. Disk glabrous, pitted. Outer female
flowers numerous, yellow, their achenes slender, about 1 mm long,
slightly ciliate-hirsute; pappus copious, white, minutely scabrid,
about 5 mm long; corolla-tube very slender, cylindric, about 5 mm
long. Perfect flowers few, usually 5 or 6 in each head, their
achenes stouter than those of the female flowers, 0.8 to 1 mm
long, sparingly ciliate. Corolla 5.5 to 6 mm long, slightly en-
larged upward, equally 5-toothed. Anthers 2.5 mm long.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8080, November 28, 1911, on boulders and banks in dense
damp forests, altitude 900 to 1,000 m, very rare.

A species characterized by its simple stems, its leaves mostly in the
median portion, not at all lobed but sharply and irregularly denticulate,

sharply acuminate, base decurrent and their lower surfaces uniformly
purple.

REVIEW.

Koorders, S. H. Exkursionsflora von Java umfassend die Bliitenpflanzen
mit besonderer Berticksichtigung der im hochgebirge wildwachsende
Arten. 1 (1911), pp. XXIV + 413 (Monocotyledonen); 2 (1911), pp.
742 (Dicotyledenen Archichlamydeae); 3 (1912), pp. VIII + 498
(Dicotyledonen Metachlamydeae). Jena: Gustav Fischer.

Java is botanically better known than is any other island of
the Malay Archipelago, and its rich and varied flora has long been
studied and by many botanists. Up to the present time, on
account of the very numerous articles dealing with the subject,
it has been quite impossible for the working botanist to gain any
comprehensive idea of the flora of the island as a whole, the total
number of genera and species, habitats and ranges of the latter,
and other data of interest. Doctor Koorders has undertaken to
compile a complete list of Javan flowering plants, but his work
is immensely more valuable than a mere enumeration of genera
and species, on account of the included keys, references, and
other data. With his extensive knowledge of the subject, based
on long residence in Java, very comprenhensive botanical explora-
tion, and critical study of all available material, Doctor Koorders
has succeeded in completing an exceedingly valuable and appar-
ently eminently usable work. He has included analytical keys
to the families, genera, and species, the key to families alone
occupying 48 pages of text. This extensive treatment is largely
due to the necessity of frequently including the same family
under two or more heads on account of exceptional characters
in various genera or species. An excellent glossary of technical
terms is included.

The arrangement of families and genera is that of Engler and
Prantl’s “Natiirlichen Pflanzenfamilien,” and it is the first work
of great importance on the Malayan flora that has been published
after this system.- As practically all recent works on the floras
of tropical Asia and Malaya have followed the sequence of
Bentham & Hooker’s “Genera Plantarum,” an alphabetical list
of families has been given with their equivalents in the latter

359

360 REVIEW.

system. In nomenclature the Vienna Code is followed rather
closely, but, as to accepted specific names, not invariably. No
attempt seems to have been made critically to study the various
cases of synonymy, and few new combinations are made. A
strict interpretation of the Vienna Code would necessitate the
acceptance of different specific names in an appreciable percent-
age of cases. Very few new forms are described.

The work is unequal in its treatment of various groups. In
some cases the species are simply enumerated under their proper
genera in the form of analytical keys, while in others more
critical enumerations are included, with literature references,
synonyms, native names, and other data. Where recent mono-
graphs were available these have been followed closely as to the
Javan forms. In most families, however, no recent monographs
are available, and the data given under these of necessity have
been compiled by the author. However, he has had the assist-
ance of many eminent European specialists in working up the
material in various families. Special attention is given to the
mountain flora throughout the work.

The “Exkursionsflora von Java” should prove to be of the
very greatest value to residents of Java who are at all interested
in the rich vegetation of that island, but it has a much wider
application. It will, in fact, prove to be of great assistance to
students of the entire Malayan-Polynesian flora, in the vast area
extending from the Malay Peninsula to Polynesia and including
the Philippines. As to the Philippines practically every family
found in the Archipelago should be determinable by the use of the
analytical key given by Doctor Koorders, as well as approximately
90 per cent of the genera found in the Islands. As to-species,
with an endemic element in the Philippines averaging about 40
per cent, comparatively few of our forms can be determined to the
species, by this work, excepting in the case of the low country
flora. In the settled areas the great majority of the species are
of very wide distribution, and of these at least 50 per cent should
be determinable by the keys given by Doctor Koorders. Ap-
proximately the same figures will apply to the Malay Peninsula
and to most of the islands in the Malay Archipelago.

Including introduced and cultivated forms, a total of about
4,930 species are considered, distributed into about 1,560 genera,
in 194 families, which gives botanists additional definite data
for comparisons of temperate and tropical floras of restricted
areas. In this connection it is of interest to note that in the

REVIEW. 361

entire Philippine Archipelago, a region much larger than the
Island of Java, we now know approximately 5,500 species, in
about 1,350 genera, representing 183 families.

The work is illustrated by a total of 19 plates, 139 text figures,
most of the former and some of the latter original, and 4 charts.
It must be considered to be one of the most important publications
dealing with the Malayan flora that has been issued in recent
years and the most important single publication dealing with the
flora of the Netherlands East Indies that has appeared since the
publication of Miquel’s “‘Florae Indiae Batavae”’ over fifty years
ago.

EK. D. MERRILL.

Vol. VII, No. 4, including pages 209 to 258, was issued September 30, 1912.

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mer

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_ THE PHILIPPINE JOURNAL OF SCIENCE, C. BOTANY. .

Vou. IX, No. 3, JUNE, 1914.

NEW OR NOTEWORTHY PHILIPPINE PLANTS
By E. D. MERRILL *

(From the Botanical Section of the Biological Laboratory,
Bureau of Science, Manila, P. I.)

The tenth paper under the above title consists of the descrip-
tions of new species in the following families: Gramineae, Cy-
peraceae, Moraceae, Loranthaceae, Olacaceae, Hernandiaceae,
Rutaceae, Meliaceae, Buxaceae, Celastraceae, Icacinaceae, Ster-
culiaceae, Theaceae, Dilleniaceae, Lecythidaceae, Flacourtiaceae,
Araliaceae, and E’'benaceae. The genus Worcesterianthus of the
Olacaceae, is proposed as new, while the genera Buetineria and
Firmiana are new to the Philippine flora. A few species of
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first time and a few changes in nomenclature are proposed.
A total of eighty-six species are described as new. The last
number of the series appeared in the year 1912.?

GRAMINEAE

ISACHNE R. Brown

ISACHNE CONFERTA sp. nov.

Planta parva, prostrata, ramosa, ramis floriferis erectis vel
suberectis, 6 ad 8 cm longis; foliis numerosis, lanceolatis, acumi- -
natis, circiter 2 cm longis, subtus leviter pilosis, supra scabridis;
paniculis exsertis, angustis, 1 ad 2 cm longis, vix 5 mm latis,
ramis paucis, erectis, 4—1-floris; spiculis confertis, circiter 1.8
mm longis, plus minusve purpureis; glumis sterilibus tenuiter
7- vel 9-nerviis, supra leviter hispidis; glumis fertilibus ellip-
soideis vel elliptico-oblongis, inferioribus glabris, 1.6 mm longis,
superioribus 4 brevioribus, apice minute ciliatis.

A small, slender, prostrate, branched plant, rooting at the
nodes, the flowering branches erect or suberect, 6 to 8 cm long.
Sheaths rather lax, longer than the internodes, glabrous; ligule
a ring of short white hairs. Leaves lanceolate, about 2 cm long,
2.5 to 3 mm wide, narrowed to the acute base and the acuminate

* Associate Professor of Botany, University of the Philippines.

? Philip. Journ. Sci. 7 (1912) Bot. 259-357.
261

962 The Philippine Journal of Science agi

apex, the lower surface somewhat pilose with scattered white
hairs, the hairs arising from minute papillae, the upper surface at
first papillate and sparingly pilose, becoming scabrid. Panicles
exserted above the upper leaves, the peduncles 1.5 cm long or less,
narrow, 1 to 2 cm long, less than 5 mm wide, the branches ap-
pressed, the lower ones 6 mm long or less and usually 4-flowered,
the upper ones shorter, the uppermost usually bearing solitary
spikelets. Spikelets crowded, about 1.8 mm long, more or less
tinged with purple. Empty glumes about 1.8 mm long, slenderly
7- or 9-nerved, slightly hispid externally in the upper part, obtuse.
Flowering glumes two, the lower one elliptic-oblong, obtuse,
quite glabrous, 1.6 mm long, the upper one ellipsoid, slightly
ciliate near the apex, usually about one-third shorter than the
lower one.

LuZON, Province of Laguna, Dahican River back of San Antonio, Bur.
Sci. 14914 Ramos, June, 1912, on stones along the river.

A rather characteristic species, distinguishable by its very narrow,

dense panicles. It is perhaps most closely allied to Jsachne pauciflora Hack.,
but is distinguished by its narrow leaves and glabrous flowering glumes.

DIMERIA R. Brown

DIMERIA CILIATA sp. nov.

Caespitosa, erecta, vaginis foliisque pilis longis albis mollibus
instructis; racemis digitatis vel subfasciculatis, 4 ad 6, tenuibus,
usque ad 12 em longis, rhachibus circiter 0.7 mm latis, triangu-
laribus, glabris vel leviter pubescentibus; spiculis 4 mm longis,
glumis I et II carinatis, carinis longe ciliato-pilosis.

A densely tufted, erect, perennial grass 50 to 80 cm high, the
culms slender, the sheaths and leaves rather densely clothed with
long, soft, white hairs, some longer (5 mm) and somewhat stiffer

ones intermixed with the shorter and softer ones, the nodes—

bearded. Leaves linear-lanceolate, acuminate, kase slightly or
not narrowed, 8 to 15 cm long, 3 to 4 mm wide, softly ciliate-
pilose on both surfaces; ligule about 1 mm long, truncate. In-
florescence long-exserted, of 4 to 6 digitate or fasciculately
arranged racemes, the racemes slender, 7 to 12 cm long, at first
pale, soon turning pale-brownish, the rachis 3-angled and some-
what flattened, about 0.7 mm wide, glabrous or slightly pubescen#,
the internodes 2 to 4mm long. Spikelets 4 mm long; first glume
somewhat boat-shaped, keeled, 4 mm long, acute or obtuse, prom-
inently ciliate-pilose with long white hairs on the keel; second
glume similar but lanceolate, and also prominently ciliate-pilose
on the keel; third glume shorter, hyaline, the awn slender, about
10 mm long.

+)
a
iy

1X, 6,3 Merrill: Noteworthy Philippine Plants 263

PALAWA®, Taytay, in open swampy places at sea level, Merrill 9320, May
28, 1913.

A species allied both to Dimeria chloridiformis K. Schum. & Lauterb.,
of the Marianne Islands, and to the Indian Dimeria fuscescens Trin., differ-
ing from the former in its smaller spikelets and narrower, not ciliate
rachises, and from the latter in its prominently ciliate-pilose sheaths and
leaves and its prominently ciliate-pilose keels of the empty glumes.

ISCHAEMUM Linnaeus

ISCHAEMUM GLAUCESCENS sp. nov. § Euischaemum.

Species J. aristato affinis, differt planta majoribus, 1 ad 3 m
longis, plus minusve glaucescens, racemis usque ad 15 cm longis,
spiculis majoribus, 8 mm longis, glumis I spiculae sessilis in 3
inferiore prominente transverse rugosis, rugis 4 vel 5, elevatis.

A rather coarse, wiry, perennial grass reaching a height of
from 1 to 1.5 m, the stems long-prostrate, scarcely or only
slightly branched, reaching a total length of from 2 to 3 m,
about 5 mm in diameter, hard, more or less glaucous under the
sheaths. Leaves narrowly lanceolate to linear-lanceolate, glab-
rous, 15 to 20 cm long, about 1 cm wide, acuminate, base grad-
ually narrowed, acute or obtuse, not at all cordate; sheaths about
as long as the internodes, rather loose; ligule truncate, 2 to 3
mm long. Racemes two, closely appressed, 12 to 15 cm long,
long-exserted, the joints of the rachis about 6 mm long, 3-angled,
bearded on the outer angle. Sessile spikelets about 8 mm long,
lanceolate; first glume coriaceous, 8 mm long, 2 mm wide,
glabrous, somewhat glaucous, acuminate, margins incurved
throughout, borders in upper part minutely scabrid, the dorsal
part in the lower one-half with 4 or 5 prominent transverse
ridges; second glume coriaceous, lanceolate, acuminate, keeled,
7 mm long; third glume hyaline, lanceolate, acuminate, 7 mm
long; fourth glume similar to the third, a little shorter, cleft
to the middle, bearing in the cleft a twisted, geniculate, 12 to 13
mm long awn. Anthers 3 mm long. Pedicellate spikelets
dimidiate, their pedicels 2.5 mm long, stout, 3-angled, bearded
on the outer angle, the first glume lanceolate, 8 mm long, 3
mm wide, acute or acuminate, coriaceous, one side rather broadly
winged, the wing short ciliate-toothed; second glume somewhat
lanceolate, 6 to 7 mm long, acuminate, the remaining ones
somewhat shorter, hyaline.

PALAWAN, Lake Manguao, Merrill 9453, April 23, 1913, gregarious along
the more or less swampy grassy borders of the lake in the openings of
narrow valleys, with or without streams of water, areas submerged during

periods of high water in the lake, altitude about 50 meters. The grass
is abundant in suitable habitats, and is utilized by the crocodiles in build-

264 The Philippine Journal of Science 1914

ing their nests. One nest examined by members of our party® containing
numerous eggs, consisted of a large mound made entirely of sand and this
grass very closely packed.

The species is manifestly allied to Ischaemum aristatum Linn., but differs
in s0 many characters that it has been considered advisable to treat it as
a distinct form.

ISCHAEMUM PUBESCENS sp. nov. § Huischaemum.

Species J, aristato affinis, differt foliis pilosis, spiculis paullo
minoribus, glumis I spiculae sessilibus utrinque nodulis 2 obs-
curis praeditis, dorso parce piloso.

An erect perennial grass about 1 m high, the sheaths and
leaves rather softly pilose with long white or pale hairs, the
nodes rather densely ciliate-bearded, the internodes, under the
sheaths, somewhat glaucous, about 3 mm in diameter. Leaves
narrowly lanceolate, 10 to 18 cm long, 8 to 12 mm wide, acu-
minate, base of the upper ones truncate, of the lower ones
narrowed, margins scabrid, both surfaces softly pilose with
scattered, long hairs; sheaths rather lax, pilose; ligule brown,
cleft, somewhat pubescent, 2 to 3 mm long. Racemes binate,
closely appressed, 8 to 10 cm long, long-exserted, rather promin-
ently white-ciliate, the joints 3-angled, bearded on all angles, 4
mm long. Sessile spikelets oblong, obtuse, about 5 mm long, 1.8
mm wide, the callus bearded; first glume coriaceous, obtuse, mar-
gins inflexed throughout, with two, broad, obscure undulations
(scarcely nodules) near each side in the lower part, the back
usually with very few, scattered, ciliate hairs; second glume
lanceolate, acuminate, somewhat keeled, 5 mm long; third and
fourth glumes hyaline, the fourth cleft to the middle, bearing in
the cleft a geniculate, somewhat twisted, scabrid awn about
13 mm long. Pedicellate spikelets dimidiate, 5 to 6 mm long,
the pedicels stout, 2 mm long, bearded; first glume widely winged
on one side, springly ciliate-pilose on the back, smooth, the
wing obscurely denticulate.

DuMARAN, Bur. Sci. 21639 Escritor, August, 1913.

Similar to Jschaemum aristatum Linn., and manifestly allied to that
species. It is characterized, however, by its prominently pilose leaves and
sheaths, its white-ciliate racemes, and the first glume of the sessile spikelets
with very obscure marginal undulations which can scarcely be called

nodules; the very short rachis-joints, and the bearded callus is also charac-
teristic.

CYPERACEAE

ELEOCHARIS R. Br.

ELEOCHARIS ACICULARIS (L.) R. & S. Syst. 2 (1817) 154; Clarke in
Hook. f. Fl. Brit. Ind. 6 (1893) 628, et in Journ. Linn. Soc. Bot. 36
(1903) 225.

IX, C, 3 Merrill: Noteworthy Philippine Plants 265

Luzon, Renguet Subprovince, Baguio, Merrill 7665, May, 1909, on seepage
slopes about rice paddies, altitude about 1,450 m.

Not previously reported from the Philippines, an additional distinct
northern type in the flora of northern Luzon. Widely distributed in North
America, throughout Europe and northern Asia, extending southward only
in China.

FIMBRISTYLIS Vahl

FIMBRISTYLIS CAPITULIFERA sp. nov. § Trichelostylis.

Planta perennis glabra, subrigida, usque ad 40 cm alta; foliis
numerosis, angustis, culmo brevioribus; spiculis omnibus ses-
silibus, capitato-congestis, capitulis umbellato-dispositis; glumis
numerosis, 5-nerviis, oblongis, subobtusis, marginibus latis, sca-
riosis; nucibus compressis vel plano-convexis, minutissime stria-
tis, circiter 0.7 mm longis; stylo trifido.

A glabrous, somewhat rigid, apparently densely caespitose,
glabrous, perennial plant 20 to 40 cm high. Leaves very nu-
merous, sheathing the bases of the stems, subrigid, flat or nearly
so, 1 to 2 mm wide, 8 to 18 cm long. Culms terete, striate,
exserted, rather slender. Inflorescence umbellate, consisting of
a central sessile head and from 3 to 6 peduncled ones, each head
with from 10 to 20 sessile, densely disposed spikelets 3 to 6 mm
in length. Glumes numerous, oblong, about 2 mm long, some-
what keeled, closely 5-nerved in the median portion, with broad,
somewhat scarious and thin margins, apex somewhat obtuse,
usually apiculate. Nut obovate, brown, about 0.7 mm long,
somewhat compressed or plano-convex, not trigonous, minutely
striate; style glabrous, slender, about 2 mm long, 3-fid.

BATANES ISLANDS, Batan, Bur. Sci. 3575 Fénix, Bur. Sci. 10203 McGregor,
Bur. Sci. 3171, 3172, 3173 Mearns. BABUYANES ISLANDS, Babuyan Islands,
Bur. Sci. 3926 Fénix. Camiguin Island, Bur. Sci. 4042 Fénix (type).

This species was previously recorded by me under the name of Fimbris-
tylis spathacea Vahl, to which it does not seem to be closely allied. It is
well characterized by its spikelets being all sessile and disposed in dense
heads, the central head sessile, the others peduncled; the rays of the umbel
vary from 1 to 4 cm in length. The species belongs in the section Triche-
lostylis, and is probably as closely allied to Fimbristylis junciformis Kunth
as to any other species. It differs from Kunth’s species in its much smaller
umbels, the spikelets much more numerous in each head, none of them

being solitary, and in the style being quite glabrous, not villous below the
fork.

FIMBRISTYLIS PALUDOSA sp. nov. § Trichelostylis.

Densissime caespitosa, erecta, glabra, efoliosa, 60 ad 100 cm
alta, culmis distincte 3- vel 4-angularibus; umbellis decompositis,
2 ad 4 cm longis; spiculis numerosis, ovoideis, 2.5 ad 3.5 mm
longis, brunneis, glumis usque ad 10, ovatis, acutis vel obtusis,

266 The Philippine Journal of Science 1914

obscure carinatis; nucibus obovoideis, triangularibus, laevibus
vel obscurissime transverse lineatis; stylo 3-fido.

A densely caespitose, leafless, perennial plant 0.6 to 1 m high
from stout rhizomes, the lower parts of the culms with 3 or 4
rather loose, imbricate sheaths, the lower ones ovate to oblong-
ovate, 1 to 2 cm long, the upper one up to 10 cm long, the mouth
oblique, with a broad, brown, membranaceous margin. Culms
prominently 3- or 4-angled, rather stiff. Inflorescence termi-
nal, 2 to 4 cm long, open, umbellately decompound, subtended
by one or two, lanceolate, acuminate, leaf-like, 1 cm long bracts.
Spikelets numerous, not fascicled, usually one sessile median one
and two lateral pedicelled ones on each ultimate branchlet, 2.5
to 3.5 mm long, brown, ovoid, the pedicels scabrid; glumes 10
or fewer, brown, ovate, acute or obtuse, obscurely keeled.

Stamens 3. Nut obovoid, 3-angled, white, 0.8 to 1 mm long,
smooth or very obscurely transversely lineate; style-arms 3.
Luzon, Benguet Subprovince, Baguio, Williams 1239 (type), May, 1904,
Elmer 6497, June, 1904, Phil. Pl. 551 Merrill, June, 1911.
Apparently a very characteristic species allied to Fimbristylis miliacea
Vahl, F. quinquangularis Kunth, etc., but differing in its leafless stems,
large rhizomes, smooth or nearly smooth nuts, and other characters.

FIMBRISTYLIS PINETORUM sp. nov. § Trichelostylis.

Erecta, glabra, culmis e rhizomate crasso, solitariis, gracilibus,
20 ad 40 em altis, basi foliosis; foliis paucis, subsetaceis, usque ad
13 em longis; inflorescentiis simpliciter umbellatis; spiculis pau-
cis, 3 ad 7, brunneis, oblongo-ovoideis, 6 ad 11 mm. longis; nucibus
obovoideis, verruculosis, albidis, 1 ad 1.2 mm longis, obscure
triangularibus; stylo trifido.

A solitary, erect, slender, perennial plant from stout, woody —

rhizomes, the rhizomes clothed with membranaceous, lanceolate,
usually brownish scales or sheaths, usually about 1 cm long.
Culms striate, 20 to 40 cm high, the base with several imbricate
leafless sheaths, and at or near the base 2 to 4 leaf-bearing
sheaths, the sheaths oblique, their margins membranaceous, the
leaves linear or setaceous, involute when dry, 1 to 1.5 mm wide,
4 to 13 em long. Inflorescence usually a simple umbel, rarely
compound, 2 to 3 cm long, the subtending bract lanceolate, acu-
minate, usually less than 1 cm long. Spikelets oblong-ovoid,
brown, many-flowered, 6 to 11 mm long, 3 to 7, one sessile, the
others mostly slenderly pedicelled, the pedicels 1 to 2 em long.
Glumes ovate, concave, keeled, glabrous, acute to obtuse, often
retuse, the margins thinner and paler. Stamens 3. Nut white,
obovoid, 1 to 1.2 mm long, obscurely 3-angled, apex broadly

1X, 6,8 Merrill: Noteworthy Philippine Plants 267

rounded or subtruncate, verruculose, not at all reticulate. Style-
arms 3.

Luzon, Benguet Subprovince, Baguio, Merrill 7664 (type), Phil. Pl.,
558 Merrill, June, 1911, scattered on slopes among various grasses in thin
pine forests, altitude about 1,550 m.

A species apparently well characterized by its solitary, not at all caes-
pitose stems, woody rhizomes, narrow leaves, usually simple inflorescence
of few spikelets, and verrucose, not at all reticulate, white, obscurely trian-
gular nuts. It is allied to Fimbristylis monticola Steud., and to F. pierotii
Miq., of India, the latter extending to Japan.

MAPANIA Aublet

MAPANIA PALUSTRIS (Hassk.) F.-Vill. Novis. App. (1882) 309; C. B.
Clarke in Hook. f. Fl. Brit. Ind. 6 (1894) 681.
Pandanophyllum palustre Hassk. in Tijdschr. Nat. Vereen. Ned. Ind. 10
(1843) 119; Boeck. in Linnaea 37: 138; Kurz in Journ. As. Soc. Beng
38:? 78.

Lepironia palustris Migq. Ill. Fl. Archip. Ind. (1871) 63, t. 25.

MINDANAO, Agusan Subprovince, Waloe, in muddy places along small
streams in forests, Merrill 7298, October, 1910.

Except for F.-Villar’s previously unverified record, not before reported
from the Philippines; Malay Peninsula and Archipelago.

There is some doubt as to the proper specific name for this plant, for
Clarke, l. c., states that Hasskarl’s description calls for a bifid style and
8 to 5 spikelets, which does not apply to Mapania palustris; however, these
statements may have been based in part on erroneous observations. Miquel
definitely states that the number of spikelets in a head is variable, and his
drawing shows a 2-cleft style. It is not clear, that in case Mapania palus-
tris is distinct from Pandanophyllum palustre Hassk., why the specific name
is retained, for if not based on Hasskarl’s description, then it was based
on a later one under the same name by Boeckler or Kurz. It seems only
reasonable to suppose that Boeckler and Miquel were familiar with type or
typical material of Hasskarl’s Pandanophyllum palustre, and under the
circumstances it seems best to retain the specific name palustris, at least
until opportunity is had to examine Hasskarl’s type. Incidentally Clarke
credits Bentham with authorship of the combination Mapania palustris;
this is incorrect, for Bentham did not make the actual transfer, but only
indicated Pandanophyllum to be a synonym of Mapania. F.-Villar appears
to be the first to make the actual transfer of the species.

MAPANIA GRACILLIMA Kiikenthal & Merrill sp. nov.

Rhizoma breve, lignosum, crassum. Culmi laterales scapi-
formes, 5-10 em alti, gracillimi, obtusanguli, striati, asperi, basi
vaginis nonnullis lanceolatis fuscis nervosis tecti et in medio
vagina unica aequali obsiti. Folia culmos longe superantia, 4—5
mm lata, plane utrinque attenuata, marginibus aculeato-scabra,
coriacea, glaucescentia. Spica parva, primo lineari-ellipsoidea
demum ovata, densa, 5 mm longa, 4 mm lata, bracteae squami-
formes. Spiculae haud numerosae oblongae. Squamae late

968 The Philippine Journal of Science 1914

ovatae, apice rotundatae, rufae. Squamellae 4 squama longiores,
2 exteriores navicularis in carina valde setulosae. Nux ellip-
soideo-lageniformis, 2 mm longa, medio turgidula, straminea,
laevis, sessilis, erostrata. Stylus brevissimus. Stigmata 3 vel 4.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8205.

Omnium Mapaniarum gracillima, foliis pro ratione angustissimis spicaque
minima ab omnibus affinibus bene distinguenda. Squamellas non nisi 4
observari, sed forsan 6 adsunt, quarum duae jam elapsae.

SCHOENUS Linnaeus

SCHOENUS FALCATUS R. Br. Prodr. (1810) 232; Benth. Fl. Austral. 7
(1878) 372; Clarke ex Hemsl. in Journ Linn. Soc. Bot. 36 (1903)
261.

LuzON, Province of Zambales, Bur. Sci. 5036 Ramos, December, 1907.

Not previously reported from the Philippines; tropical Australia and
Queensland, Borneo, and Formosa.

The specimens differ from the species, as described by Bentham, in having
somewhat smaller panicles, and in some of the flowering-glumes being dis-
tinctly retuse. A distinct Australian type, the fifth species of the genus
to be found in the Philippines.

SCIRPIODENDRON Zippel

SCIRPIODENDRON GHAERI (Gaertn. f.) comb. nov.
Chionanthus ghaeri Gaertn. f. Fruct. 1 (1788) 190, t. 29, fig. a-e; Boerl.
in Journ. Linn. Soc. Bot. 31 (1896) 246.
Scirpiodendron costatum Kurz in Journ. As. Soc. Beng. 38° (1869) 85;
Clarke in Hook. f. Fl. Brit. Ind. 6 (1904) 684; Merr. in Philip. Journ.
Sci. 2 (9107) Bot. 422.

Scirpiodendron sulcatum Migq. Ill. Fl. Archip. Ind. (1871) 65, pl. 28.

LUZON, Province of Cagayan, Abulug River, For. Bur. 19612 Curran,
January, 1911. PoLiLLo, Bur. Sci. 10249 McGregor, October, 1909. Pa-
LAWAN, San Antonio Bay, Merrill 5257, October, 1906; Taytay, Phil. Pl.
1295 Merrill. MINDANAO, District of Zamboanga, Malasugat, Merrill 8114,
December, 1911.

Chionanthus ghaeri Gaertn. f., which was described and figured from
detached fruits alone, remained a doubtful species until the year 1896, when
Boerlage was able to determine its identity from examination of the orig-
inal material used by Gaertner in describing and figuring it. Gaertner’s
name is by far the earliest one for the species, and there is no reason why
it should not be adopted.

MORACEAE

ARTOCARPUS Forster

ARTOCARPUS OVATIFOLIA sp. nov.

Arbor circiter 40 m alta, stipulis parce brevissime pubescen-
tibus exceptis glabra; foliis coriaceis, ovatis, integris, usque ad
30 em longis, acutis vel breviter acute acuminatis, basi acutis

IX, C, 8 Merrill: Noteworthy Philippine Plants 269

vel subrotundatis, nervis prominentibus, utrinque circiter 8;
stipulis membranaceis, caducis, lanceolatis, 10 cm longis; fruc-
tibus globosis vel subglobosis, circiter 6 em diametro (imma-
turis), anthocarpiis numerosissimis, apicibus subovoideis, obtusis,
vix 2 mm longis, dense ceraceo-furfuraceis.

A tree about 40 m high, glabrous or nearly so. Branches dark
reddish-brown, smooth, the ultimate ones nearly 1 cm in diame-
ter, marked with slender annular rings less than 1 cm apart.
Leaves ovate, coriaceous, glabrous, 10 to 30 cm long, 7 to 20 em
wide, entire, the apex acute or very shortly and acutely acum-
inate, or apiculate-acuminate, base acute or somewhat rounded,
both surfaces slightly shining, of about the same color and
somewhat brownish when dry; lateral nerves about 8 on each side
of the midrib, prominent, spreading, curved only near their tips,
the ultimate reticulations slender, subparallel; petioles 3 to 5
em long; stipules caducous, lanceolate, acuminate, membrana-
ceous, 10 cm long, outside sparingly pubescent with very short,
somewhat appressed hairs. Flowers unknown. Immature
fruits globose or subglobose, about 6 cm in diameter (immature),
composed of very numerous, slender anthocarps, the projecting
tips of the anthocarps ovoid or subovoid, about 2 mm long,
rounded or obtuse, about 2 mm in diameter, densely covered with
waxy-furfuraceous scales.

LUZON, Province of Laguna, San Antonio, in forests, Bur. Sci. 15040
(type), 20530 Ramos, June, 1912, February, 1913, fruits yellow.

A very characteristic species, manifestly, however, in the same group
as Artocarpus communis Forst. In texture and venation the leaves re-
semble those of Forster’s species, but are always quite entire. In being

almost glabrous the present species differs also from Artocarpus communis
Forst. The waxy-furfuraceous tips of the anthocarps is characteristic.

FICUS Linnaeus

FICUS CAMARINENSIS sp. nov. § Urostigma.

Arbor alta, glabra, ramis ramulisque rugosis; foliis alternis,
oblongo-ellipticis, subcoriaceis, nitidis, laevis, usque ad 23 cm
longis, integris, apice abrupte tenuiter acute acuminatis, basi
rotundatis, 3-plinerviis, nervis lateralibus utrinque 10 ad
12, subpatulis, prominentibus, anastomosantibus; petiolo 2.5 cm
longo; receptaculis axillaribus, solitariis, sessilibus, ellipsoideis,
in siccitate brunneis, 3 ad 3.5 cm longis, bracteis 3, reniformibus,
3 mm longis, 5 mm latis.

A tall tree, quite glabrous except the bud-scales which are
appressed-hirsute. Branches and branchlets prominently rugose
when dry, somewhat lenticellate, with distinct stipular scars, the
latter 5 to 6 mm in diameter. Leaves alternate, subcoriaceous,

126079——6

270 The Philippine Journal of Science 1914

brown and shining when dry, smooth, oblong-elliptic, 14 to 23
cm long, 7 to 11 cm wide, entire, 3-plinerved, base rounded, apex
abruptly and usually slenderly acutely acuminate, the acumen
1.5 to 2 cm long; primary lateral nerves prominent, spreading,
straight, 10 to 12 on each side of the midrib, anastomosing near
the margin, the reticulations rather lax, anastomosing between
each two lateral nerves into less distinct secondary nervs; petioles
2.5 ecm long; bud-scales lanceolate, acuminate, 1.5 cm long,
appressed-hirsute. Receptacles axillary, solitary, sessile, ellip-
soid, brown when dry, smooth or somewhat warted, 3 to 3.5
cm long, 2 to 2.5 cm in diameter, the apex rounded, the umbil-
icus distinct. Basal bracts 3, reniform, brown, coriaceous, 3
mm long, 5 mm wide. Gall flowers numerous, their perianth
segments usually 3, about 2 mm long. Male flowers few and
chiefly near the orifice, anthers 1 mm long.

LUZON, Province of Camarines, Niog, Phil. Pl. 1547 Ramos, December
17, 19138, locally known as balete.

A species well characterized by its large, ellipsoid, sessile, solitary, axil-
lary receptacles. Its alliance is apparently with Ficus forstenii Miq. (F.
vidaliana Warb.) from which it differs in its much larger receptacles and
larger leaves.

FICUS PRODUCTA sp. nov. § Sycidium.

Frutex erectus, circiter 2 m altus, foliis utrinque ramulisque
scaberulis; foliis alternis, subcoriaceis, usque ad 18 cm longis,
oblongis ad oblongo-obovatis vel ovato-ellipticis, aequilateralibus,
basi 3-nerviis, acutis vel obtusis, apice longe tenuiter caudato-
acuminatis, margine irregulariter grosse dentatis vel dentato-
serratis, haud lobatis, in siccitate subviridis, nervis utrinque
circiter 8, subtus cum reticulis valde prominentibus; receptaculis
axillaribus, solitariis vel in paribus dispositis, subglobosis vel
leviter obovoideis, scabridis, pedunculatis, 8 ad 10 mm diametro.

An erect shrub about 2 m high, scabrid. Branches terete,
slender, pale-brownish when dry, glabrous, the branchlets
scabrid. Leaves alternate, rather prominently scabrid in both
surfaces, otherwise glabrous, oblong to oblong-obovate or even
ovate-elliptic, 12 to 18 cm long, 4 to 7 cm wide, equilateral, the
base shortly 3-nerved, acute or obtuse, the apex rather abruptly
long and slenderly caudate-acuminate, the acumen 2 to 3.5 cm
long, narrow at the base, entire, acute or apiculate, the margins
rather coarsely dentate or dentate-serrate, not lobed, the teeth
irregular, mostly 5 mm apart or more, both surfaces when dry
somewhat greenish, shining, scabrid, the lower very slightly paler
than the upper; lateral nerves about 8 on each side of the midrib,
very prominent on the lower surface, anastomosing, the reticula-

Tm ¢,3 Merrill: Noteworthy Philippine Plants Hit

tions very prominent, the lower surface more or less punctic-
ulate; petioles about 1 cm long, scabrid. Receptacles orange-red,
seabrid, solitary or in pairs, axillary, subglobose to somewhat
obovoid, 8 to 10 mm in diameter, their peduncles 5 to 7 mm long,
scabrid, the apical bracts small, ovate, about 1 mm long, the
wall of the receptacle ciliate-hispid inside. Perfect female
flowers numerous, the perianth-segments narrowly lanceolate,
obtuse or acute, membranaceous, about 2 mm long, prominently
ciliate-hispid in the upper part, one about twice as broad as the
other three and cleft to about the middle, the narrower three
often somewhat spatulate. Ovary ovoid-ellipsoid, obtuse, sube-
quilateral, about 1.2 mm long, the styles distinctly lateral, 1.2
to 1.8 mm long.

MINDANAO, Subprovince of Butuan, Veruela, C. M. Weber 1182 (type),
June 19, 1911, in thickets; Talacogon, Bur. Sci. 1131 Weber, July 20, 1911,
among abacd plants.

A species in the ulmifolia group, but with equilateral leaves which are
abruptly, long and slenderly caudate-acuminate, and with somewhat smaller
receptacles than in F. ulmifolia Lam., under which name the specimens
were distributed. Its closest ally is Ficus euphlebia Merr., which has less
prominently acuminate, more distinctly and closely toothed leaves which
are acute or merely obtuse at the base, not rounded; in the present species
the interior of the receptacle and the perianth-segments are also promi-
nently ciliate-hispid with stiff white hairs.

FICUS GRANDIDENS sp. nov. § Covellia.

Arbor circiter 6 m alta, ramulis adpresso-hirsutis exceptis
glabra; foliis oblongis vel late oblongis, subcoriaceis vel charta-
ceis, usque ad 23 cm longis, glabris, laevis, nitidis, acuminatis,
basi acutis, margine grosse irregulariter sinuato-lobato-dentatis,
dentibus vel lobis 1 vel 2 utrinque; receptaculis obovoideis,
pedunculatis, in fasciculis densis hemisphericis caulinis dispo-
sitis.

A tree 5 to 6 m high, quite glabrous except the appressed-
hirsute younger branchlets. Branches terete, smooth, grayish-
brown, the ultimate ones about 4 mm in diameter. Leaves
oblong or broadly oblong, subcoriaceous or chartaceous, glabrous,
smooth, 14 to 23 em long, 5 to 11 cm wide, narrowed more or less
at both ends, apex acuminate, base acute, the margins very coarse-
ly sinuate-lobed or -toothed, the lobes or teeth 1 or 2 on each side,
obtuse or acute, the upper surface shining, dark-colored when
dry, the lower surface slightly shining, paler; nerves 5 or 6 on
each side of the midrib, prominent, usually brownish when dry,
anastomosing, the reticulations distinct, rather slender; petioles
1 to 2 cm long; stipules lanceolate, acuminate, 10 to 12 mm long,
subpersistent. Fruits fascicled on stout protuberances on the

272 The Philippine Journal of Science 1914

trunk, forming very dense, hemispheric masses 8 to 10 cm in
diameter. Receptacles obovoid, often more or less compressed-
angular by pressure of contiguous ones, 10 to 12 mm in diameter,
green, glabrous or nearly so, the umbilicus about 3 mm in
diameter; peduncles slender, 1.5 to 2 cm long, with three small
bracteoles at the apex. Fertile female flowers numerous, their
pedicels up to 1.5 mm in length; perianth short, sheathing the
pedicel below, truncate, about 1 mm long. Ovary ovoid, 1.5
mm in diameter; style lateral.

MINDANAO, District of Zamboanga, Sax River Mountains back of San
Ramon, Merrill 8089, November 28, 1911, in forests, altitude about 800 m.

A species manifestly closely allied to Ficus rubrovenia Merr., and very
similar to that species, differing in its somewhat appressed-hirsute, not
glabrous, branchlets, and its very coarsely lobed-toothed, not merely undu-
late leaves.

FICUS RIVULARIS sp. nov. § Eusyce.

Frutex circiter 4 m altus ramulis minute puberulis exceptis
glaber; foliis alternis, lanceolatis vel anguste lanceolatis, leviter
faleatis, usque ad 25 cm longis, utrinque angustatis, apice longe
tenuiter caudato-acuminatis, basi acutis, integris, subcoriaceis,
laevis, subtus albido-punctatis, nervis utrinque circiter 20; recep-
taculis axillaribus, solitariis, obovoideis, 1 ad 1.5 cm diametro,
pedunculatis, bracteis circiter 2 mm longis.

An erect, nearly glabrous shrub about 4 m high. Branches
terete, slender, reddish-brown, glabrous, the younger branchlets
somewhat puberulent. Leaves alternate, subcoriaceous, lanceo-
late to narrowly lanceolate, 12 to 25 cm long, 1.5 to 3 cm wide,
somewhat falcate, narrowed at both ends, the apex very slenderly
caudate-acuminate, the acumen up to 4 cm in length, minutely
apiculate, the base equilateral, usually acute, rarely slightly
obtuse, minutely cucullate on the upper surface, the margins
quite entire, somewhat revolute, the upper surface smooth, gla-
brous, shining, rather pale or somewhat olivaceous when dry,
the lower surface somewhat paler, or brownish, distinctly white-
puncticulate under a lens; lateral nerves about 20 on each side
of the midrib, rather distinct, anastomosing, the ultimate retic-
ulations distinct, angular, about 1 mm in diameter; petioles 5
to 10 mm long; membranaceous, deciduous, linear-lanceolate or
narrowly lanceolate, acuminate, 1.5 to 2 cm long. Receptacles
axillary, solitary, obovoid, 1 to 1.5 cm in diameter, glabrous,
somewhat narrowed at the base into a very short pseudo-stalk,
their peduncles slender, 5 to 10 mm long, with three, ovate,
obtuse, 2 mm long bracts at the summit. Gall flowers very num-
erous, in general obovoid, sessile or shortly pedicelled, about 2.5

FX, 'C, 8 Merrill: Noteworthy Philippine Plants vied

mm long, the perianth brown, chartaceous or coriaceous, irreg-
ularly 3-lobed, the lobes short, the perianth itself glabrous, about
2 mm in diameter.

LUZON, Province of Cagayan, For. Bur. 17806 Curran (type), January,
1912, Bur. Sci. 7899 Ramos, March, 1909: Province of Tayabas, Piapi,
For. Bur. 10134 Curran, March, 1908, all the specimens indicated as grow-
ing on the banks of rivers, and from their “stenophyllous” leaves, appar-
ently in places subject to submergence in times of floods or high water.

The specimens have been previously referred to Ficus linearifolia Elmer,
with which they have very little relationship, except some resemblance in
leaf-form, and manifestly the present species belongs in an entirely different
section from Elmer’s species. The very narrow, entire, slenderly caudate-
acuminate, somewhat falcate leaves which are white-puncticulate beneath
are characteristic.

FICUS LAGUNENSIS sp. nov. § Eusyce.

Frutex scandens, glaber, vel ramulis parcissime ciliato-hirsu-
tus; foliis alternis, ovatis vel oblongo-ovatis, coriaceis, in siccitate
plus minusve brunneis, usque ad 20 cm longis, acuminatis, basi
late cordatis, integris; nervis utrinque circiter 7, subtus cum
reticulisque valde prominentibus; receptaculis axillaribus, soli-
tariis vel fasciculatis, obovoideis, glabris, pedunculatis, circiter
7 mm diametro.

A scandent shrub, the young branchlets sometimes emitting
rootlets, glabrous or nearly so. Branches terete, brown, wrin-
kled when dry, glabrous or with few scattered long hairs. Leaves
alternate, coriaceous, ovate to oblong-ovate, 10 to 20 cm long,
5 to 10 cm wide, entire, the apex acuminate, the base broadly
rounded, cordate, the upper surface glabrous, shining, the
nerves impressed, the lower surface very prominently reticulate,
glabrous or with very scattered hairs along the midrib and pri-
mary nerves; lateral nerves 7 on each side of the midrib, very
prominent, the anastomosing, primary reticulations subparallel;
petioles 1.5 to 3 cm long, somewhat ciliate-pubescent or glabrous;
stipules lanceolate, acuminate, deciduous, about 1.5 cm long.
Receptacles axillary, solitary or fascicled, obovoid, glabrous,
smooth, about 7 mm in diameter, their peduncles 5 to 8 mm in
length.

Luzon, Province of Laguna, Dahican River, back of San Antonio, Phil.
Pl. 1128 Ramos, September 17, 1912, in forests.

A species very similar to and manifestly closely allied to Ficus pro-
pingua Merr., and to Ficus villosa Bl., perhaps even too closely allied to
the former to be distinguished as a true species. Its glabrous receptacles
and leaves distinguish it from Blume’s species, while from the Philippine
F. propinqua it is distinguishable by its longer peduncled, obovoid, not glo-
bose receptacles, its glabrous or nearly glabrous leaves, and its only slightly
hirsute branchlets.

274 The Philippine Journal of Science 1914

FICUS WEBERI sp. nov. § Sycidium.

Frutex circiter 4 m altus, ramulis stipulis et subtus foliis ad
nervos plus minusve brunneo-hirsutis; foliis alternis oppositis-
que, oblongo-ovatis ad obovato-oblongis, chartaceis, 10 ad 20 cm
longis, acuminatis, basi acutis, truncatis, vel leviter cordatis,
subaequilateralibus vel leviter inaequilateralibus, nervis utrinque
circiter 10; stipulis anguste lanceolatis, acuminatis, usque ad
3 cm longis; receptaculis axillaribus, sessilibus, ovoideis, sub-
glabris, circiter 1.5 cm longis, basi 3-bracteatis.

An erect shrub about 4 m high. Branches terete, brown,
hirsute with spreading brownish hairs. Leaves alternate, a
few also opposite, chartaceous, brown and slightly shining when
dry, paler beneath, oblong-ovate to obovate-oblong, 10 to 20
cm long, 6 to 10 cm wide, the upper surface smooth and glabrous,
the lower hirsute with spreading, scattered, brown or pale hairs
along the midrib and lateral nerves, apex rather slenderly and
abruptly acuminate, base acute but more often truncate or
slightly cordate, usually somewhat inequilateral, one side some-
times acute and the other rounded, margins slightly crenulate;
nerves about 10 on each side of the midrib, prominent, the
reticulations rather lax; petioles hirsute, 1 to 3 cm long; stipules
narrowly lanceolate, long-acuminate, 2 to 3 cm long, hirsute on
the back, subpersistent. Receptacles green, axillary, sessile,
ovoid, about 1.5 cm long, glabrous or nearly so, the base with
three, triangular-ovate to ovate-lanceolate, acuminate, 3 to 5 mm
long bracts. Male flowers not seen.

MINDANAO, Butuan Subprovince, Veruela, Weber 1002, in second growth
forests, June 19, 1911.

A species probably most closely allied to Ficus carpenteriana Elm., but
differing from that species in its much broader leaves, nearly glabrous
receptacles, and other characters.

FICUS WORCESTERI sp. nov. § Sycidiwm.

Arbor parva, erecta, glabra, circiter 4 m alta, ramulis teretibus ;
foliis alternis, circiter 20 cm longis, chartaceis, in siccitate palli-
dis, utrinque leviter scabridis vel laevis, concoloribus, equilate-
ralibus, apice abrupte subcaudato-acuminatis, basi acutis vel
obtusis, 3-nerviis, margine integris vel leviter obscure undulatis,
nervis utrinque circiter 10, distinctis; petiolo 1.5 ad 4 cm longo;
receptaculis axillaribus, solitariis vel fasciculatis, pedunculatis,
subglobosis, aurantiacis, 1 cm diametro.

A small, erect, glabrous tree about 4 m high, the branches
terete, smooth, somewhat straw-colored. Leaves alternate, char-
taceous, very slightly scabrid on both surfaces or smooth, slightly

THC; 3 Merrill: Noteworthy Philippine Plants 275

shining, of the same color on both surfaces when dry, and rather
pale-yellowish-green, about 20 cm long, 8 to 10 cm wide, oblong,
the apex rather abruptly subcaudate-acuminate, the acumen 1 cm
long, the base equilateral, acute or obtuse, shortly 3-nerved, the
margins entire or slightly and obscurely undulate; lateral nerves
about 10 on each side of the midrib, prominent, straight, some-
what anastomosing, the reticulations subparallel, distinct, rather
lax; petioles 1.5 to 4 cm long. Receptacles axillary, solitary or
somewhat fascicled, orange-red when mature, globose, about 1
cm in diameter, their peduncles 8 to 10 mm long, ebracteolate
or with small bracteoles near the base.

CAVILLI ISLAND, Sulu Sea, Merrill 7178, September 24, 1911.

Common in thickets near the sea on both Cavilli Island, and the neigh-
boring smaller islet known as Arena Island. Apparently allied to Ficus
ulmifolia Lam., F. blepharostoma Warb., and similar forms, distinguishable
at once by its smooth or nearly smooth leaves. Dedicated to the Honorable
Dean C. Worcester, formerly Secretary of the Interior of the Philippine
Government, to whom I am indebted for opportunity botanically to explore
Cavilli Island.

FICUS HEMICARDIA sp. nov. § Sycidium.

Arbor parva, erecta; foliis oblongis, brevissime petiolatis,
scabris, inaequilateralibus, 10 ad 30 cm longis, valde oblique
cordatis, uno latere angustatis altero latissime rotundatis; re-
ceptaculis ovoideis vel subglobosis, circiter 11 mm diametro,
hispidis, pedunculatis, solitariis vel binis in axillis foliorum vel
in ramis vel caulis fasciculatis.

An erect tree, apparently of small size. Branches terete,
light-gray, glabrous. Leaves varying greatly in size, 10 to 30
cm long, 3 to 11 cm wide, oblong, very harsh and scabrous on
both surfaces, not pubescent, somewhat shining when dry, oblong,
strongly inequilateral, margins coarsely dentate, apex rather
prominently acuminate, base very strongly obliquely cordate,
one side very much narrower than the other, narrowly rounded,
the other broadly rounded, the sinus narrow; petioles stout, 5
mm long or less; lateral nerves very prominent as are the coarse
reticulations, 6 or 7 on each side of the midrib above the base,
the basal nerves radiating, the narrow side of the leaf usually
with one, the broad side with four. Receptacles ovoid or sub-
globose, about 11 mm in diameter, hispid with scattered, spread-
ing, white, bristle-like hairs, solitary or in pairs in the axils of
the leaves, or solitary or fascicled on the larger branches and
on the trunk near its base, th peduncles hispid, about 4 mm
long, with 2 or 3, ovate, acute, 1.5 mm long bracts near the apex.
Male flowers not seen. Fertile female flowers pedicelled, the

276 The Philippine Journal of Science 1914

perianth lobes hyaline, equaling or a little longer than the ovary
which is about 1.1 mm in length; style sublateral, 1.5 mm long.

MINDANAO, Butuan Subprovince, Mount Hilong-Hilong, Weber 1001,
March 11, 1911, altitude about 130 m, on rocky slopes near streams.

The species is allied to Ficus fiskei Elm., differing in its short-petioled,
differently shaped leaves and hispid fruits, and also to F. odorata Merr.,
differing from that species especially in its smaller fruits that are not
densely hirsute. It is one of the most strongly marked species of the
genus known from the Philippines, well characterized by its harsh, inequi-
lateral, prominently and obliquely cordate, subsessile leaves. The recep-
tacles are borne in the axils of the leaves on the ultimate branches, on
the large branches below the leaves, and on the trunk of the tree near its
base, a varied distribution that is quite unusual in the genus, although
occasionally one finds the allied Ficus odorata Merr. with most of its
receptacles in the leaf-axils, and a few supplementary ones near the base
of the trunk.

FICUS CAMIGUINENSIS sp. nov.

Species F.. paucinerviae Merr. similis et ut videtur affinis,
differt foliis majoribus, receptaculis minoribus glabris.

A small tree about 6 m high. Branches terete, wrinkled and
brownish when dry, the ultimate ones about 5 mm thick. Leaves
alternate, somewhat crowded at the ends of the branchlets,
chartaceous or thinly coriaceous, subelliptic or elliptic-ovate,
greenish and shining when dry, narrowed to the acute or obtuse
base, the apex shortly and slenderly caudate-acuminate, the
acumen 1 cm long or less, 14 to 22 cm long, 8 to 13 cm wide,
entire, the uper surface glabrous, the lower one more or less
white-ciliate on the nerves and reticulations; basal nerves a
single pair, extending to about the middle of the leaf, straight,
prominent; lateral nerves above the basal pair 4 or 5 on each
side of the midrib, ascending, straight, prominent, the reticu-
lations distinct, subparallel; petioles ciliate-pubescent, 3 to 4.5
em long; stipules caducous, ovate or ovate-lanceolate, 5 to 8 mm
long, densely appressed-pubescent, the stipular scars prominent.
Receptacles mostly in pairs in the axils of fallen leaves, sometimes
solitary, sometimes somewhat fasciculate, numerous, their pe-
duncles 4 to 8 mm long, with three prominent bracts at the apex
or somewhat below the apex, glabrous, yellow, globose, about
8 mm in diameter.

CAMIGUIN DE MINDANAO, in mossy forests probably above an altitude of
1,200 m, Phil. Pl. 1197 Ramos, March 24, 1912.

Apparently a distinct species, at least not matched in our extensive
collections. I have also been unable to refer it to any of the numerous
Malayan species of which the descriptions are available here. It is similar

to Ficus paucinervia Merr. (F. integrifolia Elm.), apparently -closely allied
to it, and certainly belongs to the same section of the genus.

y

1X, ©, 3 Merrill: Noteworthy Philippine Plants 277
LORANTHACEAE

LORANTHIS Linnaeus

LORANTHUS LUCIDUS sp. nov. § Dendrophthoé.

Frutex parasiticus, subscandens, glaber, ramis ramulisque te-
retibus; foliis oppositis vel suboppositis, usque ad 12 em longis,
breviter petiolatis, oblongo-ovatis ad lanceolatis, utrinque valde
nitidis, longe acute acuminatis, basi late rotundatis cordatisque
rariter subobtusis; inflorescentiis terminalibus et in axillis supe-
rioribus, 5 ad 10 cm longis, floribus 6-meris, circiter 1.6 cm longis,
in triadibus dispositis, lateralibus pedicellatis, intermedio sessile,
triadibus racemose dispositis.

A parasitic shrub, apparently somewhat scandent along its
host, glabrous, the branches stout, terete, gray, slightly lenti-
cellate, the younger ones somewhat reddish-brown, smooth. |
Leaves opposite or subopposite, coriaceous, oblong-ovate to lan-
ceolate, normally 8 to 12 cm long, 3.5 to 5 em wide, broadly
rounded and prominently cordate at the base, exceptionally lan-
ceolate, 1.5 to 3 cm wide, and subobtuse at the base, prominently
shining on both surfaces, gradually narrowed upward to the
long and sharply acuminate apex; lateral nerves 6 to 8 on each
side of the midrib, slender, indistinct; petioles 3 to 7 mm long.
Inflorescence terminal and in the uppermost axils, 5 to 10 cm
long, the axis and lateral branches gray, obscurely furfuraceous,
composed of racemosely disposed triads. Lateral branches of
the panicles (peduncles of the triads), spreading, 1 cm long or
less, each bearing a central sessile fertile flower, and two lateral
flowers, each flower subtended by a broadly ovate, acute or acu-
minate bracteole as long as the calyx, the pedicels of the lateral
flowers 4to5 mm long. Flowers 6-merous. Calyx subcylindric,
3 mm long, truncate, the rim not produced, very minutely and
obscurely 6-denticulate. Corolla about 1.8 cm long, the basal
part somewhat inflated, about 3 mm in diameter, the lobes united
for the lower 2 mm, narrow, very slender upward, the reflexed
part above the insertion of the stamen spatulate, 5 mm long.
Filament very slender, 3 mm long; anther narrowly oblong,
continuous, 2mm long. Fruit very soft and fleshy, ovoid, abou
8 mm long.

LuZzON, Province of Laguna, San Antonio (Dahican River), Bur. Sci.
16647 Ramos, September 16, 1912, parasitic on Worcesteranthus, in forests.

A species in the same group with Loranthus ahernianus, L. acuminatis-
simus, L. saccatus, and L. ovatifolius, differing from the first three in its

very much smaller fiowers, and other characters, and from the latter in its
very different vegetative and floral characters.

278 The Philippine Journal of Science 1914

LORANTHUS FRAGILIS sp. nov. § Dendropthoé.

Frutex parasiticus, glaber, ramis ramulisque crassis, teretibus,
fragilis; foliis late ovatis, crasse coriaceis, nitidis, usque ad 15
em longis, ovatis, sessilibus vel subsessilibus, obtusis, basi late
rotundatis vel leviter cordatis; inflorescentiis in axillis superio-
ribus, floribus omnibus sessilibus, 6-meris, pallide flavidis, circiter
2.4 cm longis, in triadibus brevissime pedunculatis racemose
dispositis.

A species in the alliance with Loranthus secundiflorus Merr.,
and L. mindanaensis Merr., distinguished at once by its sessile
or subsessile, broadly rounded or subcordate leaves. Branches
up to 1 m in length, stout, terete, smooth and shining when dry,
grayish-brown, very brittle when fresh, branches 1 cm in diam-
eter breaking very readily. Leaves ovate, subopposite, thickly
coriaceous, pale greenish-olivaceous when dry, shining, 9 to 15
em long, 5.5 to 9 cm wide, apex rounded or obtuse, base broadly
rounded or somewhat cordate, sessile or subsessile, the lateral
nerves not prominent, 6 or 7 on each side of the midrib. Inflores-
cence in the upper axils, usually fascicled, about 4 cm long, the
flowers 6-merous, all sessile in triads on very short peduncles
which are racemosely disposed. Flowers pale-yellow, their sub-
tending bracts broadly ovate, obtuse, 1 mm long, the peduncles
bearing the triads of flowers very stout, about 2mm long. Calyx
oblong-ovoid, truncate, 3.5 to 4 mm long. Corolla about 2 cm
long, the petals 6, united for the lower 2 to 3 mm, the reflexed
parts of the petals above the insertion of the anthers 5 mm long.
Filaments 1.5 long; anthers continuous, linear, 2.5 mm long.

PALAWAN, Taytay, on Vernonia in forests at sea level, May 31, 1913,
Merrill 9248.

A species well characterized by its brittle stems, its thickly coriaceous
leaves, which are almost fleshy and very brittle when fresh, and which
are sessile or subsessile, broadly rounded or somewhat cordate at the base.
Its inflorescence is of the type found in Loranthus secundiflorus Merr., and

L. mindanaensis Merr., that is, the flowers sessile in triads, the triads
shortly peduncled and racemosely disposed.

LORANTHUS LEYTENSIS sp. nov. § Dendrophthoé.

Frutex parasiticus, inflorescentiis puberulis exceptis glaber;
foliis oppositis, alternis, vel subverticillatis, petiolatis, crassissime
coriaceis, oblongis, usque ad 6 cm longis, acutis, nervis lateralibus
obsoletis. Inflorescentiis axillaribus, pedunculatis, floribus 5-
meris, circiter 3.3 cm longis, in triadibus umbellatim dispositis.

A stiff, parasitic shrub, glabrous except the inflorescence, the
branches stout, reddish-brown or grayish, terete, the ultimate
branchlets about 3 mm in diameter, the nodes often thickened,

IX, C, 3 Merrill: Noteworthy Philippine Piants 279

the internodes not elongated. Leaves opposite, alternate, or
somewhat whorled at the lower nodes, thickly coriaceous, opaque,
oblong, 4 to 6 cm long, 1.5 to 3 em wide, acute, the base rounded,
obtuse, or acute, usually brownish when dry, slightly or not at
all shining, the lateral nerves obsolete; petioles about 1 cm long.
Inflorescence axillary, solitary, umbellate, the peduncles about
2 cm long, all parts more or less gray-puberulent, each peduncle
bearing about 8, umbellately arranged, 5 mm long, primary
branches, each branch bearing at its apex a triad of one middle
sessile flower and two lateral, very shortly pedicelled flowers,
each flower subtended by a broadly ovate, concave, obtuse, 1 to
1.5 mm long bracteole, the pedicels of the lateral flowers 2 mm
long or less. Flowers 5-merous, reddish or yellowish, about
3.8 cm long. Calyx 3 mm long, puberulent, truncate, the limb
scarcely produced. Corolla cylindric in bud, the lobes quite
united below, forming a 4 mm long tube, the reflexed parts of
the lobes above the insertion of the stamens narrowly oblong,
obtuse, 5 to 6 mm long, 1 mm wide. Filaments 4 mm long;
anthers continuous, oblong, 1.5 mm long.

LEYTE, Mount Ibuni back of Dagami, Bur. Sci. 15243 Ramos, August 22,
1912, growing in the tops of trees.

A species in the same group with Loranthus haenkeanus Presl, and mani-
festly closely allied to that species. It is distinguishable by its very much
smaller, differently shaped leaves.

LORANTHUS HOPEAE sp. nov. § Dendrophthoé.

Frutex inflorescentiis minute puberulis exceptis glaber; ramis
ramulisque teretibus, bi- vel trichotomis, rigidis; foliis oppositis,
crasse coriaceis, sessilibus, ovatis, in siccitate pallidis, nitidis,
usque ad 9 cm longis, apice rotundatis, basi late cordatis; inflores-
centiis axillaribus, solitariis, brevibus, paucifloris; floribus extus
puberulis, 5-meris, circiter 17 mm longis.

A parasitic shrub usually less than 50 cm in length, glabrous
except the minutely grayish-puberulent inflorescence. Branches
_and branchlets terete, stiff, brownish or grayish, smooth, di- or
trichotomously branched, more or less divaricate, the ultimate
branchlets about 2 mm in diameter. Leaves opposite, sessile,
ovate, thickly coriaceous, pale and equally shining on both sur-
faces when dry, smooth, 5 to 9 cm long, 4 to 6 cm wide, apex
broadly rounded, base broadly cordate, often prominently so;
lateral nerves 5 or 6 on each side of the midrib, very obscure
or nearly obsolete. Inflorescence solitary, axillary and at the
nodes, minutely puberulent, 2.5 cm long or less, with few, usually
4, lateral branches which are less than 1 cm in length, each
bearing at its apex 2 or 3 sessile flowers, the bracteoles puber-

280 The Philippine Journal of Science 1914

ulent, reniform, about 2.5 mm wide, 1.5 mm long, rounded.
Flowers 5-merous, orange, the tips of the petals glaucous. Calyx
cylindric, 5 mm long, puberulent, the limb slightly produced,
very obscurely crenulate or entire. Corolla 12 mm long, puber-
ulent externally, slightly inflated, cylindric, the lobes united for
the basal 3 to 4 mm, forming a tube, the reflexed part above
the insertion of the anthers oblong, obtuse, 2.5 mm long.
Anther lanceolate, sessile, 2 mm long.

MINDANAO, District of Zamboanga, coast opposite Olutanga Island, For.
Bur. 18297 Foxworthy, De Mesa, & Villamil, May 18, 1912, on Hopea sp.,
altitude about 20 meters.

A very characteristic species, recognizable by its opposite, sessile, coria-
ceous leaves which are rounded at the apex and cordate at the base, its
few-flowered, axillary, puberulent inflorescences, and puberulent flowers.
In leaf-form it is quite similar to Loranthus merrillii Elm., but is entirely
different in its inflorescence.

LORANTHUS DEMESAE sp. nov. § Dendrophthoé.

Frutex glaber, ramis ramulisque teretibus; foliis oppositis,
sessilibus, coriaceis, nitidis, usque ad 10 cm longis, ovatis
vel late elliptico-ovatis, apice acutis vel leviter acuminatis, basi
rotundatis, reticulatis; nervis utrinque circiter 10, tenuibus; in-
florescentiis axillaribus, solitariis, brevibus, paucifioris; floribus
circiter 2 cm longis 6-meris; corolla inflata, infra cylindrica,
supra acute 6-angulata.

A glabrous parasitic shrub the branches and branchlets terete,
brownish or somewhat grayish, smooth. Leaves opposite, ses-
sile, coriaceous, shining when dry, brownish-olivaceous, of about
the same color on both surfaces, 6 to 10 cm long, 4 to 6 cm wide,
the apex acute or somewhat acuminate, the base rounded; lateral
nerves about 10 on each side of the midrib, slender, not promi-
nent, the reticulations rather fine, the lateral nerves anastomosing
near the margin. Inflorescence axillary, solitary, few-flowered,
the rachis 5 mm long or less, the flowers 2 or 3 (rarely more?).
Calyx oblong-cylindric, truncate, 3 mm long, 1.8 mm in diameter.
Corolla red, about 18 mm long, much inflated, 6 mm in diameter,
the lower one-half cylindric, the upper part sharply and promi-
nently 6-angled, the tube two-thirds to three-fourths as long as
the corolla, the lobes about 4 mm wide at the base, narrowed to
the insertion of the anthers and there 2 mm wide, thence
gradually narrowed to the acute or obtuse apex, the reflexed
portion above the insertion of the anthers 5 to 6 mm long,
lanceolate. Anthers oblong, 2 mm long, much more slender than
are the thick, cylindric, 2.5 mm long filaments.

TICKS Merrill: Noteworthy Philippine Plants Q81

MINDANAO, District of Zamboanga, Talisay, on tall trees, altitude 40 to
50 meters, For. Bur. 13788 Foxworthy, DeMesa, & Villamil, June 19, 1912.

A species well characterized by its ovate, sessile leaves which are acute
or acuminate at the apex and rounded at the base, by its axillary, very
short, few-flowered inflorescences, and its much inflated corollas which are
cylindric below and sharply and prominently 6-angled above.

LORANTHUS LAGUNENSIS sp. nov. § Dendrophthoé.

Frutex glaber; foliis coriaceis, ovatis vel oblongo-ovatis,
brunneis, oppositis, breviter petiolatis, usque ad 11 cm longis;
floribus 6-meris, sessilibus, circiter 1.7 cm longis, in fasciculis
axillaribus vel lateralibus dense confertis, fasciculis vix involu-
cratis, circiter 8-floris.

A glabrous shrub, the branches very stout, rough, brownish
or grayish, the ultimate ones about 5 mm in diameter, lenticel-
late. Leaves opposite, thickly coriaceous, ovate to oblong-ovate,
6 to 11 cm long, 3.5 to 6.5 cm wide, brown when dry and some-
what shining on both surfaces, the lower a little paler than the
upper, the apex obtuse, the base usually acute; lateral nerves
about 7, very obscure or subobsolete, the reticulations entirely
obsolete; petioles stout, about 5 mm long. Inflorescence of
solitary, sessile, axillary or at the nodes, very dense, non-in-
volucrate fascicles, about 8 flowers in each fascicle. Flowers
sessile, subtended by a solitary bracteole which is reniform,
rounded, about 2 mm wide and 1.5 to2 mm long. Calyx globose
or ovoid, 2 mm long, truncate. Corolla about 1.5 cm long, the
tube 1.5 mm long, the lobes 6, linear, narrow, the reflexed part
above the insertion of the stamens thick, obtuse, 4 mm long.
Anthers linear-lanceolate, acuminate, sessile, 3.5 mm long.

LuzON, Province of Laguna, Dahican River back of San Antonio, Bur.
Sci. 15064 Ramos, June, 1912, host not indicated.

A species apparently in the group with Loranthus haenkeanus Presl,
in spite of the differences in the characters of the inflorescence in the
species. If the fascicles were involucrate the species might well be placed
in the section Lepiostegeres, but there is quite no indication of an involucre
in the material examined.

LORANTHUS FENICIS sp. nov. § Dendrophthoé.

Frutex parasiticus, inflorescentiis exceptis glaber, ramis ramu-
lisque teretibus; foliis alternis, ovatis, opacis, crasse coriaceis,
usque ad 10 cm longis, acutis vel acuminatis, basi late rotundatis,
nervis utrinque circiter 5, obscuris vel obsoletis; pedunculis
solitariis, axillaribus, floribus umbellatim dispositis; floribus
5-meris, circiter 3.3 cm longis, extus puberulis, in triadibus
dispositis, omnibus sessilibus.

A species manifestly allied to Loranthus haenkeanus, differing

289 The Philippine Journal of Science 1914

in its few-flowered umbels and in its flowers all being sessile, the
Jateral ones of each triad not pedicelled. Branches stout, terete,
grayish or reddish-brown, the branchlets grayish-brown, terete,
rather slender, the internodes 1 to 3 cm long. Leaves alternate,
thickly coriaceous, opaque and rather pale or brownish when dry,
ovate, 5 to 10 cm long, 3.5 to 6 cm wide, the apex acute or acumi-
nate, the base broadly rounded; lateral nerves usually about 5
on each side of the midrib, obscure, sometimes obsolete; petioles
about 8 mm long. Umbels solitary, in the axils of leaves on the
larger branches, gray-puberulent, the peduncles about 1.5 cm long,
the primary umbellately arranged branches few, about 6 mm
long, each bearing at its apex a triad of three sessile flowers, the
subtending bracteoles triangular-ovate, acute, 2.56 mm long,
puberulent. Calyx pubescent, narrowly obovoid, 4 mm long, the
limb slightly produced, obscurely and broadly 5-toothed. Corolla
puberulent, in bud cylindric, the tube about 6 mm long, the lobes
5, about 1.5 mm wide, the part above the insertion of the stamens
narrowly oblong, 6 to 7 mm long, 1.5 mm wide, acute. Fila-
ments 4 to 5 mm long; anthers continuous, oblong, obtuse, 2 to
2.5 mm in length.

MINDANAO, District of Davao, Baganga, Bur. Sci. 15852 Fénix, August
20, 1912, the flowers green and purplish.

LORANTHUS MARITIMUS sp. nov. § Dendrophthoé.

Frutex epiphyticus inflorescentiis exceptis glaber; ramis
ramulisque teretibus; foliis alternis oppositisque, oblongis, in
siccitate pallidis, utrinque concoloribus, opacis, usque ad 8 cm
longis, petiolatis, nervis obsoletis vel subobsoletis; inflorescentiis
e ramis_ vetustioribus, pedunculatis, umbellato-subcapitatis,
pedunculatis; floribus circiter 3.3 cm longis, 5-meris.

A parasitic shrub glabrous except the inflorescence. Branches
stout, terete, reddish-brown, the branchlets terete, slender, gray-
ish, the ultimate internodes 2 mm in diameter or less, up to 3
cm in length, frequently much shorter. Leaves oblong, alternate,
opposite or subopposite, coriaceous, uniformly pale when dry,
opaque, 5 to 8 cm long, 1.5 to 3 cm wide, the apex rounded or
obtuse, the base acute; midrib prominent below, frequently
evanescent below the apex, the lateral nerves entirely obsolete
or very faint and about 3 on each side of the midrib, ascending;
petioles about 5 mm long. Inflorescence from the larger
branches, lateral, not axillary, scattered along the internodes,
the peduncles 1.5 to 2 em long, all parts gray-puberulent. Pri-
mary branches umbellately arranged at the apex of the pedun-

ex

1X, C3 Merrill: Noteworthy Philippine Plants 983

cle, 3 mm long or less, stout, rather numerous, each bearing a
terminal triad of 3 sessile flowers, the three bracteoles broadly
ovate, obtuse, 1.5 mm long, connate below, the persistent calyces
after the fall of the corollas appearing like a rather lax globose
head. Calyx puberulent, narrowly obovoid, truncate, 3.5 mm
long, the limb very slightly produced. Corolla cylindric in bud,
3 cm long, purplish and green, puberulent externally, the lobes
5, united below and forming a cylindric tube about 6 mm long,
the lobes about 1 mm wide below, the reflexed part above the
insertion of the stamens linear-oblong, obtuse, 7 mm long, 1
mm wide. Filaments 6 mm long; anthers continuous, oblong,
obtuse, 2.4 mm long.

MINDANAO, Province of Surigao, Tamano, Bur. Sci. 15824 Fénix, August
16, 1912, on trees along the seashore.

A species manifestly in the same group with Loranthus haenkeanus Presl,
but only distantly allied to that species. It is characterized by its inflores-
cences being scattered along the older branches, not axillary, rather dense,
and its flowers all sessile, the two lateral ones not at all pedicelled as in most
of the forms in this group. ‘i

LORANTHUS ALTERNIFOLIUS sp. nov. § Lepiostegeres?

Frutex glaber, ramis ramulisque teretibus, tenuibus, brunneis,
internodiis elongatis; foliis alternis, usque ad 11 cm longis,
anguste oblongis ad oblongo-lanceolatis, subcoriaceis, nitidis,
petiolatis, apice obtusis, basi acutis; floribus 6-meris, axillaribus,
fasciculatis, sessilibus, 2 cm longis, fasciculis in alabastro in
bractea solitaria inclusis.

A glabrous shrub, the branches elongated, slender, terete,
smooth or somewhat striate, dark-brown when dry, the inter-
nodes 8 to 9 cm long, the ultimate branchlets 1.5 mm in diameter.
Leaves subcoriaceous, brittle when dry, olivaceous, of about the
same color and shining on both surfaces, 7 to 11 cm long, 2 to
3 cm wide, narrowly oblong to oblong-lanceolate, the apex
rounded, the base gradually narrowed, acute; lateral nerves very
slender, obscure or subobsolete, 6 or 7 on each side of the midrib;
petioles 1 to1.5cmlong. Flowers in sessile, axillary, solitary fas-
cicles at the nodes, 9 sessile flowers in each fascicle, in bud entirely
enveloped by a single, brown, coriaceous or subcoriaceous bract,
which bursts irregularly and falls at anthesis. Bracteoles sub-
tending the fiowers one to each calyx, orbicular or reniform,
about 2 mm long. Calyx 2 mm long and about 2.5 mm in
diameter, truncate. Corolla yellow, 6-merous, 18 mm long,
cylindric, slightly gibbous, more or less inflated, the lobes united
below forming a 5 mm long tube, the free portions 2 mm wide

284 ' The Philippine Journal of Science 1914

below, gradually narrowed upwards to the acute apex, the parts
above the insertion of the anthers lanceolate, 5 mm long.
Anther sessile, linear-lanceolate, acuminate, 4 mm long.

MINDANAO, District of Zamboanga opposite the Island of Olutanga, in
forests, altitude 30 meters, For. Bur. 13295 Foxworthy, DeMesa, & Villamil,
May 17, 1912.

A characteristic species probably belonging in the section Lepiostegeres,
although the involucre enclosing the buds is composed of a single bract.
Its comparatively narrow leaves which are rounded or obtuse at the apex
and acute at the base, as well as its 9-flowered, axillary, sessile fascicles
are its distinguishing features.

LORANTHUS WORCESTERI sp. nov. § Macrosolen.

Glaber, ramis ramulisque teretibus, griseis; foliis oppositis,
sessilibus, anguste oblongis ad late ovatis, nitidis, obtusis, basi
obtusis, late rotundatis, vel distincte late cordatis, nervis utrin-
que 4 ad 8, laxis, irregularis, tenuibus; inflorescentiis axillaribus,
pedunculis solitariis (vel fasciculatis), 3 ad 5 mm longis, 2-floris,
floribus sessilibus, 6-meris, circiter 2 cm longis.

A glabrous, parasitic shrub, the branches and branchlets terete,
glabrous, light-gray, the bark slightly wrinkled when dry.
Leaves opposite, sessile, exceedingly variable in size, narrowly
oblong to broadly ovate, 6 to 11 cm long, 2 to 7 cm wide, pale
when dry, shining on both surfaces, the apex obtuse or rounded,
the base obtuse, broadly rounded, or broadly and distinctly cor-
date; nerves 4 to 8 on each side of the midrib, slender, irregular,
anastomosing, the reticulations lax. Peduncles axillary, solitary
or few in each axil, 3 to 5 mm long, each bearing at its apex
two sessile flowers, each flower subtended by an ovate, obtuse,
3 mm long bract and two similar but smaller, free, orbicular,
1.5 mm long bracteoles. Calyx cylindric, about 7 mm long, the
limb produced about 1.5 mm, slightly spreading, thin, truncate.
Corolla pink and yellow, the lobes 6, very slightly united below,
the buds cylindric, in anthesis more or less swollen above the
base, lobes 6, about 1.5 cm long, 2.5 mm wide above the base,
then narrowed to 1.5 mm at the insertion of the anthers, the
produced part above the anthers narrowly oblong, obtuse or
acute, very thick, 6 to 7 mm long, about 1.8 mm wide and thick.
Anthers sessile, linear, 5 mm long.

MINDANAO, Bukidnon Subprovince, near Sumilao, Bur. Sci. 15673 Fénix,
August, 1912.

A species with much the floral and inflorescence characters of Loranthus
geminatus Merr., but not closely allied to that species, being at once distin-
guishable by its very diversely shaped sessile leaves. Named in honor of the
Honorable Dean C. Worcester, formerly Secretary of the Interior of the Phil-

1X: G, 3 Merrill: Noteworthy Philippine Plants 285

ippine Government, through whose invitation I was able to send a collector
with him on his southern trip of inspection in 1912.

LORANTHUS ELMERI sp. nov. § Macrosolen.

Frutex parasiticus, glaber, ramis ramulisque teretibus; foliis
oblongo-ovatis, crassissime coriaceis, oppositis, breviter petiolatis,
usque ad 14 em longis, sursum angustatis, obtusis vel obscure
acuminatis, basi acutis, nervis lateralibus obscuris, utrinque cir-
citer 7; racemis brevissimis, paucifloris, axillaribus, solitariis
vel fasciculatis, haud 1 cm longis; floribus 6-meris, circiter 1.8
em longis, basi bracteis bracteolisque 2 ornatis, pedicellis circiter
1.5 mm longis.

A parasitic shrub, quite glabrous, the branches and branchlets
gray or grayish-brown, stout, terete. Leaves opposite, very
thickly coriaceous, oblong-ovate, 8 to 15 em long, 3.5 to 5 cm
wide, brownish-olivaceous when dry, the upper surface shining,
narrowed upward to the acute or obscurely acuminate apex, the
base acute; lateral nerves very faint, about 6 on each side of the
midrib; petioles stout, 4 to 8 mm long. Inflorescence of axillary,
solitary and fascicled, very short, few-flowered racemes, the
racemes 1 cm long or less. Flowers 6-merous, their pedicels 1
to 1.5 mm long, each bearing at its apex one, broadly ovate, acute,
1.5 mm long bract and two smaller but similar bracteoles which
are usually more or less connate. Calyx ovoid, about 3 mm
long, truncate. Corolla 1.5 em long, the tube about 6 mm
long, 3.5 mm in diameter, somewhat inflated and obscurely
angled, the lobes 6, rather abruptly narrowed, about 1 mm wide
below the insertion of the stamen, the refilexed parts thickened
upward, 7 mm long, acute or obtuse; filament about 3.5 mm
long; anther continuous, 2 mm long.

PALAWAN, Mount Pulgar, Elmer 12749 (type), 13138, March and May,
1911, both distributed as Loranthus mirabilis Huerk & Muell.-Arg., a
species that is quite different and which belongs in an entirely different
section.

The alliance of Loranthus elmeri is with L. ampullaceus Roxb. It is
readily distinguished, however, by its very short racemes.

LORANTHUS SERIATUS sp. nov. § Heteranthus.

Frutex parasiticus glaber, ramis ramulisque teretibus, inter-
nodiis elongatis; foliis oppositis, breviter petiolatis, oblongo-
ovatis, coriaceis, opacis, usque ad 12 cm longis, apice breviter
obtuseque acuminatis, basi acutis, nervis lateralibus obscuris,
circiter 6 utrinque, subobsoletis; floribus 5-meris, circiter 2.5
em longis, in triadibus breviter pedunculatis dispositis, triadibus

126079——-7

286 The Philippine Journal of Science 1914

in ramis seriatim dispositis et ad nodos fasciculatis; petalis
intus ad basim ligulatis.

A parasitic glabrous shrub, the branches and branchlets
terete, reddish-brown, smooth except where the flowers are
borne, the internodes 10 to 15 cm long, the ultimate branchlets
about 2 mm in diameter. Leaves opposite, coriaceous, oblong-
ovate, 8 to 12 cm long, about 5 em wide, greenish-olivaceous on
both surfaces when dry, not shining, the apex shortly and obtusely
acuminate, the base acute, sometime a little decurrent; lateral
nerves slender, very obscure, nearly obsolete, about 6 on each
side of the midrib; petioles 3 to 5 mm long. Flowers red,
5-merous, the peduncles of the double triads stout, 2 mm long,
fascicled at the nodes and also numerous ones seriately arranged,
along one side of the branches along the internodes. Flowers
6 on each peduncle, sessile in two triads, each flower subtended
by a reniform-orbicular, rounded, obscurely pubescent, 2 mm
long bracteole. Calyx cylindric, 3.5 to 4 mm long, the limb
produced about 1 mm, truncate, minutely ciliate-pubescent.
Buds cylindric. Petals 5, free, 1.8 mm wide below, 1.5 mm
wide above, 2 to 2.2 cm long, the reflexed part above the insertion
of the stamens 6 to 7 mm long, each petal with a membrana-
ceous, reflexed, ovate, obtuse, 1 mm long, ligule-like organ on the
inner side attached about 3 mm above the base. Filaments
about 2 mm long; anthers continuous, linear, 3 to 4 mm long.

MINDANAO, District of Zamboanga, Mount Pulongbato, Bur. Sci. 16424
Reillo, September 28, 1912.

A very characteristic species, distinguishable by its peculiarly arranged
flowers, the flowers sessile in double triads on very short peduncles which
are fascicled at the nodes and serially arranged along one side of the
internodes. The peculiar ligule-like growth on the inner surface of the
petals shortly above the base is characteristic, both of the present species
and of the very similar and closely allied Loranthus cauliflorus Merr.
Loranthus seriatus is distinguished from L. cauliflorus by its very obscurely
veined leaves and its flowers in double triads, that is, 6 flowers in two
triads sessile at the apex of each peduncle, not with three flowers only as
in the latter species.

LORANTHUS FALCATIFOLIUS sp. nov. § Heteranthus.

‘e

Frutex scandens, glaber, ramulis teretibus; foliis oppositis, —

usque ad 16 cm longis, brevissime petiolatis, oblongo-lanceolatis,
leviter falcatis, coriaceis, opacis, acuminatis, basi acutis vel obtu-
sis, nervis utrinque circiter 8, obscuris, subobsoletis; floribus
5-meris, ad nodos fasciculatis, sessilibus, circiter 2.5 em longis,
alabastro cylindraceo.

A scandent, parasitic, glabrous shrub, the stems long-climbing
on the host, the branches and branchlets terete, the latter reddish-

ae

Lx; 0, 8 Merrill: Noteworthy Philippine Plants 987

brown or grayish, 3 mm in diameter or less, the internodes 5 to
8 cm long. Leaves opposite, subsessile or very shortly petioled,
oblong-lanceolate, 10 to 16 cm long, 3 to 6 cm wide, somewhat
falcate, not shining, coriaceous, the apex acuminate, the base
obtuse or acute, brownish or greenish when dry; lateral nerves
subobsolete, about 8 on each side of the midrib, very obscure;
petioles 2 mm long or less. Flowers 5-merous, axillary, sessile,
fascicled, 12 or less at each node, red below, yellow above, the
buds rather slender, cylindric, outside very obscurely pubescent
with very short scattered hairs. Calyx 3.5 to 4 mm long, straight
or somewhat curved, the limb produced about 2 mm, membrana-
ceous, somewhat spreading, truncate, margins minutely ciliate-
pubescent. Petals 5, quite free, 1.55 mm wide below, very
slightly narrowed upward, the apex obtuse, the reflexed portion
above the insertion on the stamens linear-oblong, about 8 mm
long, 1 mm wide. Filaments 4 mm long; anthers continuous,
linear, 2 mm long.

MINDANAO, Bukidnon Subprovince, Sumilao, on trees in forests, Bur. Sci.
15746 Fénix, August 3, 1912.

A species well characterized by its scandent habit, its opposite, subsessile,
subfalcate, obscurely nerved, coriaceous leaves, and its quite segsile, axillary,

fascicled flowers. It is perhaps as closely allied to Loranthus cuernosensis
Elm. as to any other species, but is very different from that form.

LORANTHUS MEDINILLICOLA sp. nov. § Heteranthus.

Frutex scandens, parasiticus, inflorescentiis exceptis glaber;
ramulis teretibus, tenuibus; foliis verticillatis, lanceolatis vel
ovato-lanceolatis, coriaceis, acuminatis, usque ad 10 cm longis,
nervis utrinque 4 vel 5, obscuris; inflorescentiis axillaribus, soli-
tariis, pedunculatis, subumbellatis, paucifloris; floribus 4-meris,
circiter 3 cm longis.

A slender, nearly glabrous, scandent, parasitic shrub, the
branches and branchlets rather slender, terete, light-gray or
brownish, mostly smooth, the ultimate branchlets 2 mm in
diameter or less. Leaves whorled, usually 4 at each node, the
internodes 4 to 8 cm long, the leaf-blades coriaceous, lanceolate
or oblong-lanceolate, 6 to 10 cm long, 1 to 3.4 em wide, narrowed
below to the acute base and above to the rather slenderly acumi-
nate apex, rather dull when dry; lateral nerves 4 or 5 on each
side of the midrib, slender, obscure, ascending; petioles 4 to 8
mm long. Inflorescence axillary, solitary, the peduncle slender,
1.5 to 2 cm long, sparingly pubescent, bearing at its apex 4 or
more subumbellately arranged branchlets, each branchlet bear-
ing two, spicately arranged, sessile flowers, the branchlets pubes-
cent, about 4mm long. Flowers sessile, 4-merous, bright yellow

288 The Philippine Journal of Science 1914

above, reddish toward the base, about 3 cm long, the basal
bracteole narrowly ovate, 1 mm long or less. Calyx narrowly
funnel-shaped, pubescent, about 3 mm long, the throat about
2.5 mm wide, the limb produced, truncate, obscurely 4-toothed.
Petals 4, entirely free, about 2.8 cm long, 2 mm wide, pubescent
externally. Filaments 5 to 6 mm long; anthers continuous,
narrowly oblong, obtuse, 3.5 mm long. Fruit not seen, said by
the collector to be white.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19842 McGregor,
February 10, 1913, on Medinilla.

Apparently most closely allied to Loranthus acutus Engl., but with much
larger flowers than that species. It also resembles L. polillensis C. B. Rob.,
in some respects, but has smaller leaves, and more numerous, pubescent
flowers.

OLACACEAE

WORCESTERIANTHUS genus novum

Flores unisexuales. Calyx parvus, 5-dentatus, vix auctus.
Petala 5, angusta, puberula. Fl. ¢: Petala valvata. Stamina
petala duplopluria, alternatim inaequalia, filamentis gracilis;
antherae orbiculari-ovoidae, rimis intus dehiscentibus. Ovarium
rudimentum cylindricum, elongatum, puberulum. FI. ¢@: Petala
imbricata. Stamina vel staminodia nulla. Ovarium ovoideum,
glabrum vel basi pubescens, 2-loculare, loculis 1-ovulatis, ovula
pendula. Stigma sessile, breviter 2-lobata. Drupa ovoidea vel
subovoidea, 2-locellata, carne crasse coriacea vel sublignosa, pu-
tamine crustaceo. Arbor dioica, parva, glabra vel subglabra.
Folia alterna, ovata vel oblongo-ovata, penninervia, integerrima.
Flores parvi, axillari, breviter pedicellati, ¢ numerosi, @ subso-
litari.

WORCESTERIANTHUS CASEARIOIDES sp. nov.

Arbor parva, usque ad 12 m alta, partibus junioribus floribus-
que exceptis glabra; foliis alternis, glabris, ovatis ad oblongo-
ovatis, chartaceis, in siccitate nitidis, pallidis, usque ad 15 cm
longis, basi acutis vel subrotundatis, apice acuminatis, nervis
utrinque circiter 5, distantibus, laxe anastomosantibus, promi-
nentibus; floribus parvis, 5-meris, ¢ circiter 3 mm longis ?¢
paullo longioribus; fructibus solitariis, glabris, circiter 1.5 cm
longis, acutis.

A small tree reaching a height of about 12 m, glabrous except
the young branchlets and the flowers, dioecious. Branches gray,
terete, slender, the young ones very obscurely angled, the growing
tips finely grayish-puberulent. Leaves alternate, exstipulate,

1X, C, 8 Merrill: Noteworthy Philippine Plants 289

ovate to oblong-ovate, chartaceous, 9 to 15 cm long, 4 to 7 cm
wide, entire, the base slightly inequilateral, acute or somewhat
rounded, the apex blunt-acuminate, the acumen broad, short,
both surfaces shining when dry, rather pale and of about the
same color, or the upper surface somewhat olivaceous; lateral
nerves about 5 on each side of the midrib, very prominent on
the lower surface, curved or curved-ascending, distant, promi-
nently looped-anastomosing, the reticulations fine, distinct; peti-
oles about 5 mm long. Flowers greenish-white, small, axillary,
shortly pedicelled, the males numerous fascicled, the females
solitary or subsolitary. Male flowers: Calyx shallowly cup-
shaped, about 1.3 mm long, somewhat pubescent, shallowly and
acutely 5-toothed, the teeth broad. Petals 5, free, valvate,
alternating with the calyx-teeth, narrowly oblong, puberulent on
both surfaces, acute or obtuse, about 3 mm long, 1.2 mm wide.
Stamens 10, all fertile, 5 long ones alternating with 5 short ones,
inserted in a single row on the rather obscure, ferruginous-
pubescent disk; longer filaments 2.3 mm long, the shorter ones
1.5 mm long, glabrous, slender; anthers broadly ovoid or orbic-
ular-ovoid, 0.5 mm long. Rudimentary ovary stout, cylindric,
puberulent, 2.5 mm long, truncate. Female flowers 5-merous,
solitary or subsolitary, their pedicels 2 mm long. Calyx about
as in the males. Petals lanceolate, puberulent, imbricate, acute
or somewhat acuminate, about 4 mm long, 1.6 mm wide. Rudi-
mentary stamens or staminodes wanting. Ovary ovoid or
narrowly ovoid, glabrous except the ferruginous-pubescent base,
narrowed above to the sessile stigma, 2-celled, each cell with a
solitary apical ovule. Stigma 2-lobed, curved or spreading, ses-
sile, about 2 mm long. Fruit ovoid, acute, about 1.5 cm long,
glabrous, the pericarp thick, very coriaceous or almost woody, the
putamen crustaceous, 2-celled, cells 1-seeded. Mature seeds not
seen.

Luzon, Province of Laguna, between San Antonio and Paete, Bur. Sct.
14943 (type), 16541 Ramos, the former with male flowers, June, 1912, the
latter with nearly mature fruits and few female flowers, September, 1912:
Province of Cagayan, For. Bur. 18436 Alvarez, February, 1909, in fruit.

This apparently represents a very distinct generic type, and one pre-
viously not described. It belongs in the Tribe Olaceae, and apparently
near Ximenia Plum., and Scorodocarpus Becc., although not at all closely
allied to either. The alternate leaves, dioecious flowers, the males fasci-
cled and the females solitary, the 10 stamens, 5 short ones alternating with
5 longer ones, and the absence of staminodes or reduced stamens in the
female flowers are distinguishing characters.

The first specimen received, in fruit, was tentatively referred by me

to the Olacaceae, but later was transferred to the Flacourtaceae, but with
no further attempt to determine it, on account of the similarity of its

290) The Philippine Journal of Science 1914 a

leaves to a specimen named Erythrospermum phytolaccoides Gardn., in
the Herbarium of the Bureau of Science. The next collection received,
with male flowers, that I have made the type, was hurridly determined and
erroneously referred to Casearia solida Merr., which in leaf-characters and
in the position and size of its’flowers it rather closely resembles; the dupli-
cates of this number were distributed as Casearia solida Merr. The next
collection yielded female flowers and nearly full grown, but immature,
fruits, and a study of all the available specimens shows that it is well
worthy of being described as a new genus. There appears to be nothing
in the Olacaceae that very closely approaches it, although there is no
doubt whatever in my mind but that it properly belongs in this group.

The genus is dedicated to the Honorable Dean C. Worcester, for many
years Secretary of the Interior of the Philippine Government, in recogni-
tion of his interest in and great services to the advancement of the scientific
work in the Philippines.

HERNANDIACEAE

HERNANDIA Linnaeus

HERNANDIA OVIGERA Linn. Amoen. Acad. 4 (1757) 125; Meissn. in DC.
Prodr. 15° (1864) 262; F.-Vill. Novis. App. (1880) 182.

MINDANAO, District of Zamboanga, Mount Pulongbato, Bur. Sci. 16442
Reillo, Sept. 20, 1912. CAMIGUIN DE MINDANAO, Bur. Sci. 17424 Ramos,
April, 1912.

This species was based on Arbor ovigera Rumph. Herb. Amboin. 3: 103,
pl. 123, and is characterized by its leaves not being peltate. In other
respects it is very similar indeed to Hernandia peltata Meissn., and it
seems to be doubtful whether or not the two are really distinct. In the
Philippines Hernandia peltata Meissn., is common and widely distributed
along the seashore, and this seashore form apparently always has peltate
leaves, although sometimes but slightly so, as in Copeland 1331 from the
strand at Tagalaya, Davao, Mindanao. At any rate, if the type of Her-
nandia sonora L. be interpreted as the American species, which seems to
be the logical course to follow, H. ovigera L. is the oldest name for the
oriental form, should H. peltata Meissn. and H. ovigera L. be united.
Hernandia ovigera L. has already been reported from the Philippines by
F.-Villar, but the record has not previously been verified.

Both specimens cited above are from the interior forests, not from the
coast; Hernandia peltata Meissn, is confined to the beach forests.

Malay Archipelago, Java to Amboina.

ILLIGERA Blume

ILLIGERA MEGAPTERA sp. nov.

Frutex scandens, inflorescentiis exceptis glaber; foliis 3-folio-
latis, foliolis oblongo-ovatis ad ovato-ellipticis, coriaceis, nitidis,
usque ad 15 cm longis, breviter acuminatis, basi late rotundatis,
integris, nervis utrinque 5 vel 6, prominentibus; fructibus 3.5
em longis, cum alis 10 ad 12 em latis.

Scandent, glabrous except the inflorescence, the stems
terete, at least 5 mm diameter, dark-colored when dry, striate.

eeamear

is; C,'s Merrill: Noteworthy Philippine Plants 291

Leaves 3-foliolate, their petioles 9 to 12 ecm long. Leaflets
oblong-ovate to elliptic-ovate, coriaceous, entire, 13 to 15 cm
long, 8 to 9 cm wide, apex shortly acuminate, base broadly
rounded, pale-olivaceous when dry, both surfaces equally shining ;
lateral nerves 5 or 6 on each side of the midrib, prominent, the
reticulations distinct; petiolules 1.5 to 2 em long. Flowers not
seen, the infructescence at least 20 cm long, subferruginous-
pubescent, the mature fruits 3.5 cm long, including the wings 10
to 12 cm wide, the wings brown, shining, elliptic-ovate, broadly
rounded, firmly chartaceous to subcoriaceous, 3.5 to 4 em wide.

MINDANAO, Butuan Subprovince, Bur. Sci. 21511 Escritor, August, 1913,
locally known as cabacaba.

Very characteristic on account of its relatively large leaflets and its
very large fruits, the broad long wings being rather remarkable.

ILLIGERA RETICULATA sp. nov.

Frutex scandens, subglaber; foliis 3-foliolatis, foliolis coria-
ceis, late ovatis, acuminatis, basi rotundatis vel leviter cordatis,
usque ad 11 cm longis, nervis utrinque 5 vel 6, subtus cum
reticulis prominentibus; fructibus 3.5 cm longis, 3-alatis, alis
lateralibus coriaceis, nitidis, late ovatis, rotundatis, circiter 4
cm longis, altero 1 ad 1.5 em longis latisque.

A seandent shrub, nearly glabrous (flowers unknown).
Leaves 3-foliolate, the petioles about 8 cm long; leaflets broadly
ovate, coriaceous, about 11 cm long, slightly acuminate, entire,
base rounded or somewhat cordate, pale when dry, the lower
surface bearded in the axils; lateral nerves 5 or 6 on each side
of the midrib, prominent, the reticulations lax, also prominent;
petiolules 2 cm long. Panicles ample, in fruit glabrous. Fruits
about 3.5 em long, 3-winged, the lateral wings broadly ovate,
coriaceous, rounded, shining, about 3.5 cm long and 3 cm wide,
the other one much reduced, 1 to 1.5 ecm long and wide and
confined to the lower one-half of the fruit, the opposite side
with a mere keel, not at all winged.

LuzON, Province of Laguna, Mount Maquiling, For. Bur. 7761 Curran
& Merritt, October 24, 1907, on trees, open cogon slopes, altitude 160 meters.

A species well characterized by its prominently reticulate leaves and its
comparatively large, 3-winged fruits, the lateral wings very large, the

other very much reduced, and the fourth represented by a mere ridge or
keel.

ILLIGERA ELLIPTIFOLIA sp. nov.
Frutex scandens inflorescentiis exceptis glaber; foliis trifolio-
latis, foliolis ellipticis, 7 ad 138 cm longis, utrinque rotundatis,
in siccitate pallidis, nitidis, nervis utrinque circiter 6; infruc-
tescentiis laxis, ampli, circiter 20 cm longis, dense olivaceo- vel

292 The Philippine Journal of Science 1914

subferrugineo-pubescentibus; fructibus junioribus circiter 3 em
longis, bialatis, alis chartaceis, circiter 2 cm longis, ovatis,
rotundatis.

A scandent shrub, glabrous except the inflorescence. Stems
terete, about 5 mm in diameter, nearly black when dry. Leaves
3-foliolate, the petioles up to 12 cm long; leaflets entire, sub-
coriaceous, elliptic, 7 to 13 em long, 4 to 7.5 em wide, rounded at
both ends, pale and somewhat shining when dry, the petiolules
1 to 2.5 cm long; lateral nerves about 6 on each side of the midrib.
Inflorescence lax, subpyramidal, about 20 cm long, the branches
few, scattered, the lower ones up to 9 em long, all parts rather
densely olivaceous- or subferruginous-pubescent with short hairs.
Flowers unknown. Fruits (immature) about 3 cm _ long,
2-winged, the wings rather thin, ovate, rounded, about 2 cm
long.

MINDANAO, Agusan Subprovince, in forests along streams southeast of
Nasipit, altitud about 20 meters, For. Bur. 20571 Miranda, September, 1918.

Well characterized by its lax, rather densely pubescent panicles, its two-
winged fruits, and its elliptic leaflets.

ILLIGERA CARDIOPHYLLA sp. nov.
Frutex scandens, glaber (floribus ignotis) ; foliis 3-foliolatis,
foliolis late ovatis, coriaceis, 9 ad 12 ecm longis, base late
rotundatis cordatisque, apice late, abrupte, obtuse acuminatis;
fructibus 3 cm longis, 4-alatis, alis coriaceis, late rotundatis,
glabris, lateralibus 3.5 cm longis, ceteris triplo brevioribus.

A seandent glabrous shrub (flowers unknown). Branches
terete, black when dry, 4 to 5 mm in diameter. Leaves 3-folio-
late, the petioles 8 to 10 cm long; leaflets broadly ovate, coria-
ceous, 9 to 12 em long, nearly or quite as wide, olivaceous and
somewhat shining when dry, base broadly rounded, rather
prominently cordate, apex abruptly, shortly, and obtusely acu-
minate; lateral nerves 5 to 6 on each side of the midrib, the
reticulations lax, not prominent; petiolules 1.5 to 2 em long.
Panicles in fruit 15 cm long, glabrous or nearly so; fruits
glabrous, 4-winged, about 3 em long, the lateral wings coria-
ceous, rounded, broadly ovate, about 3.5 cm long, the other two
about one-third as long as the lateral, ones.

BABUYANES ISLANDS, Camiguin, Bur. Sci. 4082 Fenix, June 28, 1907, in

thickets near old clearings.
The characteristic features of this species are its broadly ovate, rather
prominently cordate leaves ard its comparatively large, 4-winged fruits.

(To be concluded)
[Vol. 1X, No. 2, including pages 97 to 189, was issued June 23, 1914]

THE PHILIPPINE

JOURNAL OF SCIENCE

C. BOTANY

VoL. IX AUGUST, 1914 No. 4

NEW OR NOTEWORTHY PHILIPPINE PLANTS, X

By E. D. MERRILL *

(From the Botanical Section of the Biological Laboratory, Bureau of
Science, Manila, P. I.)

(Concluded )
RUTACEAE

ATALANTIA Correa

ATALANTIA MARITIMA sp. nov.

Species A. distichae (Blanco) Merr. affinis, differt foliis ellip-
ticis haud prominente acuminatis, margine leviter crenulatis
vix integris.

An unarmed shrub or small tree 2 to 6 m in height, glabrous
except the inflorescence, or the younger branchlets sometimes
slightly pubescent. Branches terete, slender, usually reddish-
brown when dry, the branchlets greenish or straw-colored and
more or less angled or compressed near the nodes. Leaves
alternate, subcoriaceous or coriaceous, elliptic to ovate-elliptic,
shining, rather pale when dry and of nearly the same color
on both surfaces, 5 to 10 cm long, 2.5 to 6 cm wide, subequally
narrowed at both ends, the base acute, the apex obtuse and dis-
tinctly retuse, not gradually narrowed to an acumen, the margins
above distinctly but distantly crenulate, not entire, glandular-
punctate beneath; lateral nerves very numerous, close; petioles
5to10mm long. Panicles axillary and terminal, 3 to 6 cm long,
pubescent, many-flowered, narrow, the lower branches 2 cm long
or less, the panicles usually larger in fruit and up to 11 cm in
length. Flowers 5-merous. Calyx pubescent, cup-shaped, about

* Associate Professor of Botany, University of the Philippines, Manila,
: ee
129553 293

294 The Philippine Journal of Science 1914

3 mm in diameter, 5-lobed, the lobes orbicular-reniform, rounded,

about 2 mm long and somewhat wider than long, pubescent.

Petals 5, about 6 mm long, glandular-punctate, oblong-obovate,

rounded. Stamens 10, 2-seriate; filaments flattened, free, the

longer ones about 5 mm long, the shorter 4 mm in length; anthers
ovoid, rounded at both ends, 1.3 mm long. Disk annular,
surrounding the base of the ovary. Ovary glabrous, ovoid or
globose; style thick, cylindric. Fruit globose, yellow, glabrous,
about 8 mm in diameter, fleshy, with a single seed.

Apo ISLAND, Mindoro Strait, Bur. Sci. 178 Bermejos (type), November,
1905. Ticao, For. Bur. 1060 Clark, May, 1904. PANay, Capiz, Copeland
107, January, 1904. BoHoL, Tagbilaran, Bur. Sci. 1278 McGregor, July,
1906. MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9469, 9270
Withford & Hutchinson, February, 1908. PALAWAN, For. Bur. 3796 Curran,
February, 1906, Bur. Sci. 628 Foxworthy, March, 1906.

The species is manifestly closely allied to Atalantia disticha (Blanco)
Merr., from which it may be easily distinguished by its very differently
shaped leaves which are not gradually narrowed upward to an acuminate
apex and which are distinctly crenulate, not entire. All the specimens are
indicated as growing in thickets or forests at the edge of the beach, on
beach-cliffs, or near mangrove swamps. The Visayan name in Ticao is
indicated by Clark as carucabagao.

CLAUSENA Burmann

CLAUSENA GRANDIFOLIA sp. nov.

Frutex 2 ad 3 m altus, vix aromaticus, glaber vel subglaber;
foliis usque ad 40 cm longis, foliolis 11 ad 15, chartaceis vel
subcoriaceis, 9 ad 15 cm longis, oblongo-ovatis ad oblongis,
inaequilateralibus, acuminatis; paniculis circiter 20 cm longis
angustis, leviter pubescentibus; floribus parvis, 5-meris; fruc-
tibus globosis, carnosis, 7 ad 10 mm diametro.

A shrub 2 to 3 m high, glabrous except the slightly puberulent
younger parts and the inflorescence, not aromatic. Branches
terete, stout, brownish-gray. Leaves alternate, 25 to 40 cm long,
the petioles and rachis minutely puberulent, becoming glabrous;
leaflets 11 to 15, alternate, oblong-ovate to oblong, the larger
ones up to 15 cm long and 6 cm wide, the smaller ones about

10 cm long and 3 to 4 ecm wide, those in the upper part of the ©

rachis longer than the lower ones, chartaceous to subcoriaceous,
strongly inequilateral, entire, acuminate, base rounded to acute,
rather pale, shining, and of about the same color on both sur-
faces when dry; lateral nerves 7 to 9 on each side of the midrib,
prominent, distant, irregular, anastomosing, the reticulations
lax; petiolules about 8 mm long. Panicle terminal, narrowly
pyramidal, about 20 cm long, somewhat puberulent, the lower

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Tx. 'c, £ Merrill: Philippine Plants, X 295

branches 7 cm long or less, the upper gradually shorter. Flowers
‘apparently numerous, somewhat crowded at the apices of the
branchlets, 5-merous, the pedicels very short. Calyx shallow,
1.5 mm in diameter, 5-lobed, lobes broadly ovate, acute. Petals
5, imbricate, elliptic-ovate, 2.5 mm long, glandular, glabrous.
Stamens 10, somewhat 2-seriate; anthers oblong, 1 to 1.2 mm
long, much longer than the filaments which are somewhat
enlarged below. Ovary cylindric, prominently rugose, glabrous,
5-celled. Fruit globose, fleshy, up to 1 cm in diameter,
with from 1 to 3 seeds; seeds about 7 mm long, the cotyledons
thick, plano-convex.

PALAWAN, Mount Capoas, Merrill 9544, April 21, 1918, on talus slopes,

steep forested ridge, altitude about 800 meters.
A species well characterized by its unusually large leaves and leaflets.

EVODIA Forster

EVODIA LAXIRETA sp. nov.

Frutex vel arbor parva, glabra; foliis 3-foliolatis, foliolis
coriaceis vel subcoriaceis, nitidis, anguste oblongo-obovatis, usque
ad 12 cm longis, 3 ad 5 cm latis, obtusis, basi sensim angustatis,
cuneatis; nervis lateralibus utrinque circiter 10, prominentibus,
reticulis laxis, distinctis; inflorescentiis in axillis superioribus,
anguste paniculatis, circiter 5 cm longis; coccis 1 vel 2, ovoideis
vel ellipsoideis, circiter 4 mm longis.

A shrub or small tree, quite glabrous (flowers not seen) ;
branches terete, pale-brownish, shining, rather stout, wrinkled
when dry. Leaves 3-foliolate, the petioles 2.5 to 5 cm long;
leaflets narrowly oblong-obovate, coriaceous or subcoriaceous,
when dry prominently shining and of about the same color on both
surfaces, 9 to 12 cm long, 3 to 5 ecm wide, entire, the apex obtuse,
below gradually narrowed to the acute or cuneate base; lateral
nerves about 10 on each side of the midrib, prominent, anas-
tomosing, the reticulations lax, the nerves and reticulations
about equally prominent on both surfaces; petioles about 1.5 cm
long. Panicles solitary, in the upper axils, narrowly pyramidal,
about 5 cm long, shortly peduncled, the branches few, the lower
ones 1.5 cm long or less. Fruit of 1 or 2 cocci, the cocci ovoid to
ellipsoid, rounded, about 4 mm long.

- Minpanao, Bukidnon Subprovince, Bur. Sci. 21407 Escritor, July 27,
1913, locally known as pamintang gubat.

Thus species may prove to belong to the genus Melicope when the flowers

are known. In general it resembles Melicope triphylla (Lam.) Merr., but

differs in its thicker leaves which are prominently and laxly reticulate
on both surfaces.

296 The Philippine Journal of Science 1914

EVODIA CAMIGUINENSIS sp. nov.

Arbor parva partibus junioribus inflorescentiisque villosis;~
foliis trifoliolatis, foliolis anguste oblongis, anguste oblongo-
obovatis, vel oblongo-oblanceolatis, utrinque angustatis, apice
acuminatis, usque ad 9 cm longis, nitidis, glabris, vel subtus ad
costa parce villosis; inflorescentiis axillaribus, paniculatis,
pallide subdense villosis, quam folia multo brevioribus.

A small tree, the branches light-gray, shining, glabrous, sub-
terete or very obscurely 4-angled, the younger parts slightly
villous. Leaves 3-foliolate, their petioles 1 to 3.5 ecm long,
sparingly villous, becoming nearly glabrous, glandular; leaflets
narrowly oblong, narrowly oblong-obovate or oblong-oblanceo-
late, chartaceous or submembranaceous, green and shining
when dry, of nearly the same color on both surfaces, 6 to 9
em long, 1.5 to 3 cm wide, narrowed at both ends, the apex
rather distinctly acuminate, the base acute, the upper surface
quite glabrous, the lower one glabrous or slightly villous along
the midrib, prominently glandular-punctate with very numerous,
small, dark-colored glands; petiolules 3 to 5 mm long; lateral
nerves about 12 on each side of the midrib, very slender, anas-
tomosing, distinct but not prominent. Panicles axillary, nar-
rowly pyramidal, rather prominently villous-pubescent with
pale hairs, 5 to 7 cm long, open, rather lax, the branches spread-
ing, the lower ones 2 cm long or less, the upper ones gradually
shorter. Female flowers: Pedicels 1.5 to 2 mm long, pubescent.
Sepals 4, oblong-ovate obtuse, pubescent, about 1.2 mm long.
Petals 4, oblong, obtuse, about 2.4 mm long, 1.2 wide. Stami-
nodes 4, very slender, 1 mm long. Ovary densely villous; style
slender, 1.5 mm long, villous below. Male flowers and fruits
not seen.

CAMIGUIN DE MINDANAO, Bur. Sci. 14664 Ramos, April, 1912.

A species distinguishable by its rather narrow leaflets, but more espe-
cially by its open, narrowly pyramidal panicles which are prominently

villous with pale hairs. It has somewhat the appearance of the Asiatic
Evodia pteleaefolia (Champ.) Merr., but is quite distinct from that species.

EVODIA VILLAMILII sp. nov.

Arbor alta, glabra, gemmis inflorescentiisque exceptis glabra;
foliis 3-foliolatis, foliolis subcoriaceis, oblongis ad oblongo-
ellipticis, usque ad 22 cm longis, in siccitate pallidis, breviter
acuminatis, basi angustatis, plus minusve decurrento-acuminatis,
subsessilibus vel brevissimme petiolulatis, nervis utrinque cir-
citer 20, prominentibus; inflorescentiis axillaribus, 5 ad 8 cm
longis, breviter pedunculatis, dense multifloris, floribus rosaceis,
4-meris, circiter 5 mm longis.

IX, C, 4 Merrill: Philippine Plants, X 297

A tree about 20 m high, glabrous except the buds and the
inflorescence. Branches stout, olivaceous. Leaves opposite,
their petioles 5 to 10 cm long, the leaflets 3, oblong to oblong-
elliptic, or sometimes narrowly elliptic-obovate, subcoriaceous,
pale and somewhat shining when dry, 15 to 22 cm long, 6 to 12
ecm wide, the apex somewhat acuminate, the acumen short, the
base narrowed, usually somewhat decurrent-acuminate, sessile or
the petiolules very short; lateral nerves about 20 on each side of
the midrib, prominent. Inflorescence of solitary, axillary,
densely many flowered, peduncled, somewhat pubescent cymes 5
to 8 em long, 6 to 11 cm in diameter, all parts more or less gray-
pubescent. Flowers pink, their pedicels pubescent, 5 mm long.
Sepals 4, orbicular-reniform, rounded, 1.5 mm in diameter.
Petals 4, pink, ovate to ovate-elliptic, acute, 5 mm long, 3 mm
wide, appressed-pubescent inside. Stamens 4; filaments 6 to
7 mm long, glabrous, the upper 1 mm much narrowed and
abruptly inflexed; anthers versatile, oblong, 2 mm long. Ovary
deeply 4-lobed, densely villous, the cells 2-ovuled; style glabrous,
6 mm long; stigma punctiform. Cymes in fruit about 7 cm in
diameter, rather dense, the cocci cartilaginous, about 5 mm long,
the seeds jet black, shining, ellipsoid, about 3 mm long.

Luzon, Province of Laguna, Mount Maquiling, For. Bur. 20653 (type),
20880 Villamil, October, 1918, February, 1914, in forests, altitude 300 to
350 meters, the former in flower, the latter in fruit and from the same
tree; also C. F. Baker 449, November, 1912, detached inflorescence only.

A very striking species not closely allied to any other known Philippine
form, readily recognizable by its short, dense, hemispheric, many flowered,
short-peduncled cymes, the flowers retaining their pink color long after
being dried.

EVODIA TERNATA (Blanco) comb. nov.

Orixa ternata Blanco FI. Filip. (1837) 62, ed. 2 (1845) 45, ed. 3, 1: 84.

Evodia robusta F.-Vill. Novis. App. (1880) 34, non Hook. f.

Evodia triphylla Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 68, non DC.

For this Philippine form that has been confused with two different
species, Evodia robusta Hook. f. and LE. triphylla DC., it appears necessary
to establish a new name, for it is apparently a distinct and valid species.
I have rather arbitrarily typified the species by Blanco’s Orixa ternata,
transferring his specific name to the proper genus. It is, however, impos-
sible definitely to determine whether or not the material referred here,
and below described, is really the species Blanco described, due to his im-
perfect description. So far as his description goes, it applies, not only in
the characters ascribed by him to the species, but also in its time of
flowering. It is, moreover, the only form known to me to which Blanco’s
Orixa ternata can be referred. F.-Villar’s reduction of Orixa ternata to
Evodia robusta Hook. f. is manifestly incorrect, for the latter species does
not occur in the Philippines. The following description is typified by Lev-
berg 6133.

298 The Philippine Journal of Science 1914

A shrub or small tree 2 to 4 m high, glabrous except the in-
florescence and the younger parts. Branches rather stout,
smooth, more or less compressed, usually brownish when dry,
the growing tips more or less pubescent. Leaves opposite,
3-foliolate, their petioles 4 to 9 em long; leaflets chartaceous, 10
to 18 em long, 5 to 9 cm wide, elliptic, oblong, or oblong-obovate,
of about the same color on both surfaces, shining when dry, oli-
vaceous or yellowish-brown, the apex distinctly acuminate, the
base acute; lateral nerves about 15 on each side of the midrib,
rather slender, distinct, anastomosing; petiolules 3 to 8 mm long.
Panicles axillary, narrowly pyramidal, slightly pubescent,
about 9 cm long, the branches distant, spreading, the lower
ones about 2 cm long on staminate inflorescences, on female
ones, in fruit, the panicles up to 15 cm long, and the lower
branches 5 cm in length. Male flowers: Pedicels short. Sepals
4, slightly pubescent, orbicular-ovate, obtuse, about 1 mm long.
Petals 4, elliptic-oblong, acute, 2.5 mm long, about 1.5 mm wide,
glabrous. Stamens 4; filaments 2.5 to 3 mm long; anthers
elliptic-oblong, 1.2mm long. Rudimentary ovary densely villous.
Female flowers not seen. Cocci subellipsoid, 5 mm long, the
shining black seeds 2.5 to 3 mm in diameter.

Luzon, Province of Bataan, Mount Mariveles, Leiberg 6133, July, 1904,
with ¢ flowers, For. Bur. 2055 Borden, For. Bur. 1474 Ahern’s collector,
both in fruit, August, 1904: Province of Rizal, Bur. Sci. 13601 Ramos,
August, 1911, in fruit: Province of Cagayan, Claveria, Bur. Sci. 107382
McGregor, August, 1909, with immature ¢ flowers.

The species is perhaps as closely allied to Evodia glabra Blume as to
any other, but has quite different leaves and venation. It resembles quite
closely the Asiatic material referred to Hvodia pteleaefolia (Champ.) Merr.,
but is specifically distinct.

EVODIA SUBCAUDATA sp. nov.

Species quoad foliis inflorescentiisque EH. ternatae (Blanco)
Merr. simillima differt foliolis subcaudato-acuminatis, nervis
lateralibus magis numerosis, floribus hermaphroditis.

A shrub or small tree the branches, branchlets, inflorescences,
petioles and lower surfaces of the leaflets on the midribs and
lateral nerves rather softly pubescent with short grayish hairs.
Branches and branchlets terete, or the tips of the latter a little
compressed. Leaves opposite, 3-foliolate, their petioles 4 to 9 em.
long, those of the same pair of leaves unequal in length. Leaflets
membranaceous, oblong to oblong-elliptic, or obovate-elliptic,
subequally narrowed to the acute base and to the slenderly sub-
caudate-acuminate apex, the acumen 1 to 1.5 cm long, the termi-
nal leaflet up to 15 cm long and 6 cm wide, the lateral ones

IX, C, 4 Merrill: Philippine Plants, X 299

somewhat smaller, all entire, the upper surfaces smooth, glabrous,
shining, the lower slightly paler, pubescent on the midrib and
lateral nerves, minutely glandular-punctate; lateral nerves 17 to
20 on each side of the midrib, prominent, anastomosing, the retic-
ulations slender, rather lax; petiolules 2 to 3 mm long. Panicles
axillary, pubescent, solitary, about 8 cm long, narrowly pyramidal,
the lower branches 2.5 cm long or less, densely many flowered.
Flowers 4-merous, perfect, their pedicels 1 to 1.2 mm long, the
bracteoles less than 0.6 mm in length. Sepals ovate, acute, pubes-
cent, about 0.8 mm long. Petals oblong, obtuse or acute by the
inflexed tips, 2 mm long. Stamens 4; filaments 2 mm long, the
oblong anthers 1 mm in length. Ovary depressed-globose,
4- lobed, minutely pubescent, the cells 1-ovuled; style 2 mm long.

MINDANAO, Butuan Subprovince, Bur. Sci. 15922 Fenix, August, 1912.

A species quite similar in general appearance to the Philippine Hvodia
ternata (Blanco) Merr., redescribed above, and probably most closely
allied to that form, differing in some vegetative details, much more pubes-
cent, more slenderly acuminate leaflets with more numerous nerves, and
especially in its perfect, not dioecious flowers. As to the Asiatic forms
it is apparently most closely allied to Evodia pteleaefolia (Champ.) Merr.,
but is abundantly distinct.

LUNASIA Blanco

LUNASIA MOLLIS sp. nov.

Species L. amarae affinis, differt partibus junioribus inflores-
centiis foliisque subtus molliter stellato-pubescentibus.

A shrub or small tree, the branchlets stout, yellowish-brown,
densely stellate-pubescent with short, yellowish-brown hairs, the
same type of indumentum on the inflorescence, petioles, and lower
surfaces of the leaves. Leaves oblong-obovate, subcoriaceous, 18
to 24 cm long, 6 to 10 cm wide, entire, the upper surface green,
shining, glabrous except for the more or less stellate-pubescent
midrib, the lower surface pale, rather densely stellate-pubescent,
prominently glandular-punctate, the apex rounded, obtuse, or
very obscurely, broadly, and obtusely acuminate, narrowed below
to the abruptly rounded base; lateral nerves 15 to 17 on each side
of the midrib, prominent, spreading, anastomosing, the reticula-
tions rather lax; petioles 4 to 5 cm long; inflorescence a very
narrow panicle, solitary, axillary, densely stellate-pubescent, 7 to
- 15 cm long, none of the branches (in young bud) exceeding 1 cm
in length, but perhaps longer in age. Young buds densely stel-
late-pubescent, globose. Open flowers and fruits not seen.

CEBU, Limusan, Bur. Sci. 11026 Ramos, on dry hills, March 14, 1912.

This proposed new species, although represented by immature material,
certainly belongs in the genus Lunasia, and is so different from our common

300 The Philippine Journal of Science 1914

and variable Lunasia amara Blanco, that I have no hesitation whatever
in describing it as new. Lunasia amara Blanco is distinctly lepidote, not
at all stellate-pubescent. JL. mollis may be more closely allied to L. babu-
yanica than to L. amara, for L. babuyanica is more or less stellate-pubescent,

although not all to the degree of L. mollis; there are also vegetative —

differences.

LUNASIA OBTUSIFOLIA sp. nov.

Species L. amarae affinis differt foliis apice late rotundatis vel
obtusis, vix acuminatis, baseque distinct cordatis vix acutis.

A shrub, the young branches, petioles, and inflorescence densely
pale-lepidote. Leaves oblong-obovate, membranaceous, shining,
of nearly the same color on both surfaces or the lower one a
little paler than the upper when dry, 8 to 17 cm long, 4 to 8 em
wide, entire or the upper part very obscurely undulate, the apex
broadly rounded or obtuse, narrowed from about the middle or
somewhat above to the distinctly cordate base, the base 1 to
1.5 cm wide, the lobes rounded, the sinus narrow, shallow, both
surfaces with few, scattered, pale, lepidote scales when young,
in age glabrous or nearly so, glandular-punctate; lateral nerves
up to 15 on each side of the midrib, spreading, prominent,
slenderly anastomosing, the reticulations slender, lax, not prom-
inent; petioles 3 to 6 cm long. Male panicles up to 30 cm in
length, narrow, their branches few, the lower ones up to 3 cm
in length, the flowers straw-yellow, in scattered, small, globose
heads. Sepals about 0.5 mm long. Petals oblong-ovate, acute
or acuminate, about 2 mm long. Filaments about 0.5 mm long.
Female flowers and fruits not seen.

BOHOL, Tagbilaran, on beach cliffs, Bur. Sci. 1278 McGregor, July 12,
1906.

A species decidedly similar to and manifestly very closely allied to
Lunasia amara Blanco, differing in its broadly rounded or obtuse, not
acuminate leaves which are also distinctly cordate and not acute at the
base; another distinguishing character is its elongated male panicles, some
of which greatly exceed the leaves in length, while in Lunasia amara
Blanco the panicles are shorter than the leaves.

LUNASIA MACROPHYLLA sp. nov.

Species L. amarae affinis, differt foliis multo majoribus, usque
ad 45 cm longis, coriaceis, integerrimis, nervis usque ad 45
utrinque.

A shrub about 2 m high, the younger parts, petioles, and in-
florescence densely pale-lepidote. Leaves pale when dry, coria-
ceous, shining and of about the same color on both surfaces, the
upper surface glabrous, minutely and obscurely beaded along the
‘reticulations, the lower surface very sparingly lepidote, the apex

1X, C, 4 Merrill: Philippine Plants, X 301

prominently acuminate, narrowed from the upper two-thirds to
the acute or abruptly obtuse base, the margins quite entire;
lateral nerves, at least on larger leaves, up to 45 on each side
of the midrib, very prominent, faintly anastomosing near the
margins, the reticulations slender, not very prominent; petioles
stout, thickened at their apices, about 10 cm long. Male pani- °
cles axillary, in young bud narrow, spike-like, the linear bracts
about 5 mm long. Open flowers and fruits not seen.

MINDANAO, District of Zamboanga, Port Banga, For. Bur. 9299 Whitford
& Hutchinson, January 7, 1908, in dipterocarp forests, altitude about 50
meters.

A species distinguishable by its very large, quite entire, coriaceous leaves
which have about 45 pairs of very prominent lateral nerves.

LUNASIA NIGROPUNCTATA sp. nov.

Species L. amarae simillima et affinis, differt foliis subtus valde
nigro-punctatis, capsulis vix rostratis.

A shrub about 4 m high, the branches, branchlets, petioles,
and inflorescences densely pale-lepidote, somewhat shining.
Leaves subcoriaceous, oblong-oblanceolate to narrowly obovate-
oblancolate, 22 to 25 cm long, 5.5 to 6.5 cm wide, rather gradually
narrowed from above the middle to the acute or somewhat obtuse
base, the apex broadly blunt-acuminate, the margins irregularly
undulate or slightly repand, the upper surface dark-olivaceous
when dry, shining, glabrous, or with scattered lepidote scales ~
along the midrib, the lower surface much paler, with scattered
lepidote scales, and with numerous, black, shining, round glands
which are distinct to the naked eye, 1 to 4 glands to each ultimate
reticulation; lateral nerves about 25 on each side of the midrib,
prominent, anastomosing; petioles 5to 6cm long. Inflorescences
axillary, solitary, apparently narrowly paniculate, in fruit 6
em long or less. Capsules usually of three cocci, by abortion
sometimes reduced to two, rarely to one, densely lepidote, pale,
shining, the individual cocci obovoid, base rounded, apex trun-
cate, about 12 mm long, 9 to 10 mm wide across the top, concen-
trically wrinkled when dry, the outer corner scarcely rostrate, or
very shortly so.

Luzon, Province of Tayabas, Baler, Bur. Sci. 21188 Escritor, June, 1913.

A species manifestly closely allied to Lunasia amara Blanco which it
strongly resembles, but from which it differs in its rather remarkable black
glandular-punctate leaves and in its scarcely rostrate capsules. In Lunasia
amara, while the leaves are glandular-punctate, the glands are never black

and shining as in the present species, while the capsules are very promi-
nently rostrate on the upper outer angle of each coccus.

302 The Philippine Journal of Science 1914

LUNASIA AMARA Blanco Fl. Filip. (1837) 783.

This species is common in forests and of very wide distribution in the
Philippines. It is exceedingly variable in its vegetative characters, but
seems to be decidedly constant in floral and fruit characters. The normal
form has nearly entire leaves, usually distinctly, often prominently, acumi-
nate. In his original description Blanco describes the leaves as “serpen-
teadas,” which is best translated as undulate, and many of our specimens
show this character. The extreme repand form, however, is so distinct,
that were no intergrades present most botanists would not hesitate to
describe it as new. I had even written up a diagnosis of the form as a
new species, but distinct intergrades being present, and no characters other
than vegetative ones being detected by which to distinguish the forms, I
have considered it best to indicate the repand-leaved type as a variety.

Var. REPANDA var. nov.

A typo differt foliis prominente undulato-repandis vel repandis.

Luzon, Province of Cagayan, Bur. Sci. 7828 Ramos: Province of Nueva
Vizcaya, Bur. Sci. 11264, 11184 McGregor, For. Bur. 14876 Darling, For.
Bur. 15825 Curran & Merritt. MINDANAO, District of Davao, Bur. Sci.
15842 Fénix, August, 1912 (type).

The first two specimens cited have very large leaves, up to 45 cm in
length, but the others have medium-sized or small leaves, those on the
type being less than 15 em in length.

LUVUNGA Hamilton

LUVUNGA SCANDENS (Roxb.) Hamilt. in Wall. Cat. (1832) no. 6382.
Limonia scandens Roxb. FI. Ind. 2 (1832) 380.

PALAWAN, Malampaya Bay, Binaloan, Merrill 9405, May, 1913, forested
slopes, altitude 20 meters.
India to the Malay Peninsula and Indo-China; new to the Philippines.

MELIACEAE
AGLAIA Loureiro

AGLAIA BERNARDOI sp. nov. § Huaglaia.

Arbor alta partibus junioribus subtus foliolis inflorescentiis-
que dense stellato-pubescentibus; foliis circiter 70 cm longis;
foliolis circiter 15, oblongis, membranaceis vel chartaceis,
acuminatis, basi cordatis, usque ad 15 cm longis, supra glabris,
subtus densissime brunneo-stellato-pubescentibus; paniculis ju-
venilibus quam folia multo brevioribus, floribus numerosis,
sessilibus, in ramulis glomeratim dispositis.

A tree reaching a height of 20 meters, the younger parts,
inflorescence, petioles, rachis, and lower surfaces of the leaves
densely stellate-pubescent with pale-brownish hairs, or some-
times ferruginous in color. Ultimate branches terete, 8 to 10
mm in diameter. Leaves alternate, about 70 cm long, the leaflets
about 15, oblong, membranaceous or chartaceous, 10 to 15 cm

IX, C, 4 Merrill: Philippine Plants, X 308

long, 3.5 to 5 em wide, acuminate, base cordate, the upper
surface brown when dry, strongly shining, glabrous or slightly
stellate-pubescent along the midrib; lateral nerves straight,
parallel, distinct, 20 to 27 on each side of the midrib; petiolules
pubescent, about 3 mm long. Panicles axillary, when young
less than one-half as long as the leaves, peduncled, stellate-
pubescent, about 25 cm long, the primary branches less than 4 cm
long, densely many flowered, the flowers glomerate. Flowers
5-merous, sessile, the buds globose, 1.5 mm in diameter, the calyx
densely stellate-pubescent externally. Petals 5, orbicular, gla-
brous, rounded, about 1 mm long. Staminal tube depressed-
globose, free from the petals, truncate, the anthers inserted at
the base, reaching the orifice but hardly protruding, 0.6 mm
long.

LUZON, Province of Cagayan, Gattaran, in open forests, altitude about
20 meters, For. Bur. 15205 Bernardo, May 27, 1912.

A species manifestly belonging in the group with Aglaia argentea Blume,
but entirely distinct from that species in its floral, vegetative, and indu-
mentum characters.

AGLAIA TRUNCIFLORA sp. nov. § Euaglaia?

Arbor circiter 18 m alta ramulis inflorescentiis petiolis subtus
foliisque ad costa dense stellato-tomentosis; foliis imparipinnatis,
foliolis 7, ellipticis vel oblongo-ellipticis, usque ad 25 cm longis,
subcoriaceis, basi obtusis vel subrotundatis, apice abrupte sub-
caudato-acuminatis, nervis utrinque circiter 17; paniculis cauli-
nis, circiter 25 cm longis, fructibus ellipsoideis, circiter 2 cm
longis.

A tree about 18 m high, the branches, branchlets, inflorescence,
petioles, rachis, petiolules, and midribs on the lower surfaces of
the leaflets densely stellate-tomentose with brown hairs, not at
all lepidote. Branches terete, the ultimate ones about 4 mm in
diameter. Leaves alternate, rather distant, about 45 cm long;
leafiets 7, the lower pair less than one-half as large as the
upper ones, ovate, the others elliptic to oblong-elliptic, up to 25 cm
long, and 10 cm wide, rather pale-brownish when dry, the upper
surface glabrous and slightly shining, the lower surface of the
same color, nearly glabrous except for the stellate-tomentose
midribs, the base obtuse or rounded, the apex abruptly subcau-
date-acuminate, the acumen narrow, blunt, about 1.5 em long;
lateral nerves about 17 on each side of the midrib, very prominent
on the lower surface, obscurely anastomosing, the reticulations
lax, indistinct; petiolules stout, about 8 mm long. Panicles
from the trunk, stellate-tomentose, 20 to 25 cm long. Flowers
not seen (5-merous). Fruit ellipsoid, brown when dry, minutely

+

804 The Philippine Journal of Science 1914 |

_Stellate-tomentose with brown hairs, rounded at both ends,
about 2 cm long, the persistent calyx with 5 short teeth.

LEYTE, Dagami, in forests along streams, Mount Ibuni, Bur. Sci. 15282
Ramos, August 21, 1912.

A species probably allied to Aglaia cauliflora Koord., of Celebes, which
is inadequately described. The Philippine form agrees with Koorders’s
species in the peculiar character of its cauline inflorescence, which although
not uncommon in other genera of the Meliaceae, is exceedingly rare in
Aglaia. It differs in its stellate-tomentose, not lepidote, indumentum, its
smaller leaves, 7 instead of 5 leaflets, and slightly larger fruits.

DYSOXYLUM Blume

DYSOXYLUM ROSTRATUM sp. nov. § Hudysoxylum.

Arbor alta, partibus junioribus inflorescentiisque brunneo-pube-
rulis exceptis glabra; foliis alternis, circiter 40 cm longis; foliolis
10, alternis, oblongis, coriaceis, nitidis, rectis, subaequilateralibus,
acuminatis, in siccitate brunneis, nervis utrinque 8 ad 11, subtus
valde prominentibus, reticulis obscuris, tenuibus, laxis; inflores-
centiis brunneo-puberulis, paniculatis, multifloris, in axillis
superioribus, circiter 20 cm longis; floribus 4-meris, circiter 7
mm longis, petalis glabris, liberis; tubo utrinque villoso; ovario
pubescente; fructibus 1-locularibus, ad 4 cm longis, valde inaequi-
lateralibus, apice lateraliter rostratis.

A tall tree, the younger parts and the panicles appressed
brown-puberulent, otherwise glabrous. Branchlets less than 1
cm in diameter, brown, wrinkled, the older parts glabrous.
Leaves alternate, about 40 cm long, the rachis at first puberulent,
soon entirely glabrous; leaflets 10, alternate or subalternate,
oblong, coriaceous, brown when dry, straight, subequilateral or
entirely equilateral, the apex rather prominently acuminate, the
acumen blunt, the base rounded to acute or somewhat decurrent,
the upper surface shining when dry, the lower slightly paler, also
shining; lateral nerves 8 to 11 on each side of the midrib, promi-
nent on the lower surface, impressed on the upper surface, not
anastomosing, the reticulations very slender, lax, obscure,
often nearly obsolete; petiolules 1.5 to 2 em long, when young
grayish-puberulent. Panicles in the upper axils, forming a sub-
terminal inflorescence, all parts brown-puberulent, about 20 em
long, oblong in outline, the lower branches 5 to 7 cm in length,
the flowers numerous, white, racemosely arranged on the ultimate
branchlets, 4-merous, their pedicels 2.5 to 4 mm long. Calyx
disk-shaped or shallowly saucer-shaped, about 3 mm in diameter,
irregularly but often rather prominently 4-toothed, some of the
teeth often apiculate-acuminate. Petals 4, entirely free, glab-
rous, about 7 mm long, 2.5 mm wide, obtuse. Staminal-tube

Pe

“,

;
4
os
,
y
2
f

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r.

5
Ae
‘

IX, ©, 4 Merrill: Philippine Plants, X 805

somewhat villous on both surfaces, cylindric, 6 mm long, 10-
toothed, the teeth distinct, short. Anthers 10, sessile, included,
about 1 mm long. Disk cup-shaped, villous, about 1.5 mm high.
Ovary ovoid, somewhat pubescent, the style glabrous, about 2.5
mm long. Fruit in general obovoid, falcate, 3 to 4 cm long,
2 to 2.5 cm in diameter, 1-celled, with a single large seed, the tip
of the fruit projecting laterally as a stout, more or less strongly
recurved beak 1 to 1.5 cm long.

LuzON, Province of Laguna, Papot, near San Antonio, Phil. Pl. 1473
Ramos (type), February 26, 1918, in forests; Dahican River, Bur. Sci.
16552 Ramos, September, 1912, in fruit.

Apparently in the same group with Dysoxylum alliaceum Blume. The
inequilateral, 1-seeded, rostrate fruit is very characteristic.

DYSOXYLUM EUPHLEBIUM sp. nov. § Hudysoxylum.

Arbor alta, glabra, ramulis crassis, 1 ad 1.5 em diametro;
foliis 20 ad 40 cm longis, circiter 7-jugis, alternis, foliolis alter-
nis vel supoppositis, coriaceis, nitidis, oblongo-ovatis, late obtuse-
que acuminatis, inaequilateralibus, plus minusve falcatis, usque
ad 12 cm longis, nervis utrinque circiter 9, valde prominentibus,
reticulis obsoletis; paniculis ramosis, anguste pyramidatis, dense
multifioris, folia subaequantibus, terminalibus; floribus 4-meris,
circiter 8 mm longis, petalis glabris, tubo libero, utrinque villoso,
ovario pubescente.

A tall tree, entirely glabrous except the staminal tube and
the ovary. Branchlets stout, terete, brownish, much wrinkled,
1 to 1.5 cm in diameter, marked with very large petiolar scars.
Leaves alternate, 20 to 40 cm long, about 7-jugate, the rachis
and petiole brown and somewhat longitudinally rugose when dry.
Leaflets thickly coriaceous, oblong-ovate, more or less falcate,
inequilateral, 8 to 12 cm long, 3 to 6 cm wide, apex shortly and
bluntly broad-acuminate, base rounded to acute, brownish-oliva-
ceous when dry, the upper surface very strongly shining, the
lower of about the same color but dull; lateral nerves about 9
on each side of the midrib, very prominent on the lower surface,
not anastomosing, the reticulations obsolete; petiolules about 5
mm long.. Panicles in the uppermost axils forming a terminal
inflorescence, the individual ones narrowly pyramidal, often as
long as the leaves, the branches spreading, the lower ones about
8 cm long, densely many-flowered, the flowers crowded, shortly
pedicelled or subsessile, 4-merous. Calyx disk-like or shallowly
saucer-shaped, obscurely toothed, about 3 mm in diameter.
Petals 4, oblong, obtuse, 8 mm long, 3 mm wide, glabrous, quite
free. Staminal-tube cylindric, 7 to 8 mm long, somewhat villous
inside and outside, truncate. Anthers 10, sessile, 0.8 mm long,

306 The Philippine Journal of Science 1914

included. Disk cylindric, truncate, about 1.5 mm high, villous.
Ovary sparingly appressed-pubescent; style glabrous, 7 mm long.
Fruit (immature) globose or obovoid, about 3 cm in diameter.

LuzOoN, Province of Laguna, Gumihan, near San Antonio, Phil. Pl. 1404
Ramos (type), February 27, 1913, in forests; between San Antonio and
Paete, Bur. Sci. 15102 Ramos, June, 1912.

A species manifestly in the group with the Malayan Dysoxylum allia-
ceum Blume, but differing from that and allied forms in many characters.
The strongly shining leaves, with prominent primary nerves and entirely
obsolete reticulations, and the densely flowered panicles are characteristic.
The dried flowering specimens have a strong, very disagreeable odor that
is characteristic of various parts of many species in the genus.

TURRAEA Linnaeus

TURRAEA MEMBRANACEA sp. nov. § Huturraea.

Arbor parva, circiter 5 m alta; foliis membranaceis, ovatis vel
oblongo-ovatis, utrinque angustatis, acuminatis, usque ad 9 cm
longis, nervis utrinque circiter 7, vetustioribus glabris vel sub-
glabris; racemis axillaribus, brevissimis, paucifioris; floribus 5-
meris, circiter 2.5 cm longis, tubo apice 10-laciniato, ovario glabro,
5-loculare; fructibus 5-locellatis, valvis valde recurvatis, crassis-
sime coriaceis, 12 mm longis.

A small tree 4 to 5 m high, deciduous, the branchlets, young
leaves, and inflorescence more or less pubescent, in age nearly
glabrous. Branches slender, terete, glabrous, brown or reddish-
brown, the branchlets gray-pubescent. Leaves simple, alternate,
membranaceous, ovate to oblong-ovate, subequally narrowed to
the acute base and to the acuminate apex, 5 to 10 cm long, 2 to
4 cm wide, when young slightly pubescent on both surfaces, at
least on the midrib and nerves, in age nearly or quite glabrous,
slightly shining; lateral nerves about 7 on each side of the midrib,
prominent; petioles about 1 cm long, pubescent. Racemes axil-
lary, very short, pubescent, few-flowered, the flowers appearing
with the new leaves, the rachis of the racemes 5 mm long or
less. Flowers yellowish-white, their pedicels slender, 2 cm long.
Calyx cup-shaped, about 3 mm long, pubescent, 5-toothed, the
teeth ovate-acute, about 1.5 mm long. Petals 5, free to the
base, 2 to 2.3 cm long, linear, above somewhat narrowly spatulate,
below 1 mm wide, near the apex 3 to 3.5 mm wide, glabrous.
Staminal-tube glabrous, slender, 2.5 em long, laciniate-lobed at
the apex, the lobes thin, linear, acuminate, about 2 mm long,
alternating with the anthers; anthers subsessile, attached appar-
ently at the very apex of the tube, 1.8 mm long. Ovary ovoid,
5-celled ; cells 2-ovuled; style slender, exserted 6 to 7 mm beyond
the end of the staminal-tube; stigma about 2 mm in diameter.

1. C, 4 Merrill: Philippine Plants, X 307

Fruit before dehiscence apparently ovoid, glabrous, splitting into
5 valves, the valves strongly recurved, very thickly coriaceous,
almost woody, ovate to oblong-ovate, acute or acuminate, about
1.2 em long, longtudinally keeled along the inside and grooved
along the back. Seeds obovoid, black and shining when dry,
about 6 mm long.

Luzon, Province of Rizal, Bosoboso, Bur. Sci. 977 Ramos (type), June,
1906, in flower: Province of Pampanga, Mount Arayat, Merrill 3913, Oc-
tober, 1904, For. Bur. 9619 Zschokke, October, 1907, both in fruit.

This is in all probability the form that has been credited to the Phil-
ippines by several authors as Turraea pubescens Hellen. The Philippine
record was based on Vidal 1654 from Marinduque, which I have examined
in the Kew Herbarium, and which I have noted as being matched by two
of the specimens cited above as well as by Loher 4648 from Arayat. At
the time the Kew material was examined it was noted that the Philippine
material differed from 7. pubescens Hellen., in its differently shaped leaves,
glabrous ovary, and larger, quite different fruit. I now have a specimen
of Turraea pubescens in the herbarium of the Bureau of Science, Cochin-
china, Pierre 2772, which confirms the above notes. The fruits are remark-
ably different both in texture and in shape, the valves being merely coria-
ceous and not reflexed. It is quite evident from the material at present
available that the Philippine form is specifically very distinct from the
one found in southeastern Asia, and that Turraea pubescens Hellen. does not
extend to the Philippines.

TURRAEA PALAWANENSIS sp. nov. § Euturraea.

Suffrutex parvus, circiter 20 cm altus, erectus, partibus junior-
ibus pubescentibus; foliis membranaceis, oblongis, usque ad 10
em longis, irregulariter lobatis vel undulato-lobatis, costa nervis-
que subtus pubescentibus; racemis paucifloris; floribus 4 ad 4.5
cm longis.

A species similar and allied to Turraea humilis (Blanco)
Merr., and T. pumila Benn. An undershrub about 20 cm high,
the woody parts slender, reddish-brown, glabrous or slightly
pubescent, not or but sparingly branched, the growing parts
gray-pubescent. Leaves alternate, oblong in general outline,
membranaceous, 5 to 10 cm long, 2 to 4.5 cm wide, the apex
obtuse, the base decurrent-acuminate, the margins irregularly
lobed or undulate lobed, the lobes usually 3 on each side, the
upper surface glabrous or nearly so, the lower one very slightly
paler then the upper and more or less pubescent on the midrib
and lateral nerves; lateral nerves 4 or 5 on each side of the
midrib, prominent, usually forked, the reticulations very lax;
petioles 2 cm long or less, pubescent. Racemes axillary, solitary,
3-flowered or less, pubescent. Flowers white, 4 to 4.5 cm long,
the bracts linear, 2 to 3 mm long. Calyx pubescent, the lobes
5, linear, acuminate, pubescent, 6 to 8 mm long, about 1 mm

308 The Philippine Journal of Science 1914

wide. Corolla slender and tube-like below, the tube-like portion
about 3 cm long and 2 mm in diameter, the lobes 5, oblong,
narrowed at both ends, about 1.5 cm long, 6 mm wide, acute,
narrowed below into the long and slender claws that make
up the tube-like part. Staminal tube slender, glabrous, ex-
serted from the tube-like part of the corolla about 1.5 cm,
the reflexed laciniae at its apex 9 or 10, linear, acuminate, 4
mm long. Filaments filiform, inserted inside the tube, the
oblong anthers sparingly hispid, just exserted, about 1.5 mm
long, blunt at both ends, the apex obliquely apiculate. Ovary
ovoid, pubescent, 5-celled; ovules 2 in each cell; stigma depressed-
globose, 0.5 mm in diameter.

PALAWAN, San Miguel, near the seashore, Bur. Sci. 15563 Fénix, July
10, 1912.

A species very similar to and manifestly allied to both Turraea humilis
(Blanco) Merr., and to T. pumila Benn., differing from both in its larger
flowers. The latter species was described from Javan specimens, and is at
present unknown outside of that island unless the Philippine T. humilis
proves to be identical. Both F.-Villar and myself have considered the
Philippine and Javan plants to be identical, the former reducing Blanco’s
Plagianthus humilis to Turraea pumila Benn., while I retained Blanco’s
specific name as the valid one, it being the older. It seems, judging from
Bennett’s description of T. pwmila, that his species is not the same as the
Philippine one described by Blanco as Plagianthus humilis and which I
have transferred to Turraea as a valid species. Turraea humilis (Blanco)
Merr. is known only from Luzon, and it is exceedingly local and rare. It
is represented by the following specimens:

Luzon, Province of Rizal, San Juan del Monte, near Manila, Merrill
6282, June, 1908: Province of Laguna, San Antonio, Bur. Sci. 14994 Ramos,
June, 1912.

The leaves are 3 to 6 cm long, ovate or elliptic-ovate, undulate but not
lobed, and the flowers are 3 to 3.5 cm in length.

WALSURA Roxburgh

WALSURA VILLAMILII sp. nov.

Species distinctissima ap omnibus adhuc cognitarum differt
filamentis haud } connatis, foliis 7-foliolatis.

A tree about 13 m in height, nearly glabrous, or the inflores-
cence and younger parts very sparingly pubescent, the growing
parts distinctly brownish-puberulent. Branches glabrous,
brownish-olivaceous, lenticellate. Leaves 7-foliolate, about 50
ecm long, the petiole 10 to 15 cm in length, brown when dry.
Leaflets oblong to oblong-lanceolate, coriaceous, entire, glabrous,
15 to 20 em long, 3 to 5.5 em wide, the apex slenderly subcau-
date-acuminate, the base acute, the upper surface, when dry,
pale-greenish-olivaceous, shining, the lower one glaucescent, the

IX, ©, 4 Merrill: Philippine Plants, X 309

prominent nerves and slender reticulations brown; lateral nerves
about 16 on each side of the midrib, very prominent on the
lower surface, somewhat curved, anastomosing near the mar-
gins; petiolules 2 to 5 mm long. Panicles terminal and in
the upper axils, brown when dry, very slightly pubescent,
in fruit up to 40 cm long. Immature fruits obovoid, 1 to
1.5 em long, externally densely covered with a short, dense,
brownish-purple indumentum, the persistent petals oblong,
pubescent, 4 mm long. Staminal tube very short, 0.5 mm long
or less, the filaments flattened, ciliate-pubescent, about 1.5 mm
long, obtuse, the anthers 0.5 mm in length.

MINDANAO, District of Zamboanga, near Margosatubig, in hill forests,
For. Bur. 18764 Foxworthy, DeMesa, & Villamil, May 11, 1912, altitude
about 120 meters, and locally known to the Moros as sasd.

A very distinct and characteristic species, readily distinguished from
all previously described ones by its 7-foliolate leaves and its short staminal
tubes. ;

BUXACEAE
BUXUS Linnaeus

BUXUS RIVULARIS sp. nov. § Hubuxus.

Frutex circiter 1 m altus, glaber; foliis coriaceis vel sub-
coriaceis, lanceolatis, usque ad 5 cm longis, utrinque subaequaliter
angustatis, apice acute acuminatis, basi acutis; racemis axillari-
bus, solitariis, 1 ad 2 cm longis.

A shrub about 1 m high, glabrous, the branches slender, light-
gray, terete, the branchlets distinctly angled. Leaves lanceolate,
2.5 to 5 cm long, 5 to 12 mm wide, rather pale when dry, of
about the same color and slightly shining on both surfaces,
subequally narrowed to the acute base and to the sharply acum-
inate apex, subsessile or very shortly petioled, often a little fal-
cate; nerves very faint, anastomosing in a distinct marginal
nerve. Racemes axillary, solitary, 1 to 2 cm long, glabrous, male
flowers below, the terminal flower usually female, the pedicels
about 3 mm long, the bracteoles ovate, acute, 1 mm long. Male
flowers: Outer two sepals lanceolate, acuminate, about 2.2 mm
long, 1 mm wide, the inner two as long but ovate or oblong-ovate,
acute, 1.5 mm wide. Filaments about 2.5 mm long. Rudimen-
tary ovary depressed-globose, entire. Female flowers at the
end of the raceme bearing the male flowers. Bracteoles linear-
lanceolate, the outer 2 sepals ovate, a little longer than the inner
four which are broadly ovate, acute, 2.2 mm long, slightly ciliate-
pubescent, all slightly accrescent in fruit, persistent, and about
3 mm long. Ovary glabrous. Young fruit ovoid, glabrous, 5

129553

2
a

310 The Philippine Journal of Science 1914

to 6 mm long, crowned by the prominent styles and recurved
stigmas.

Luzon, Province of Tayabas, Guinatacutan, Bur. Sci. 18169 Foxworthy
& Ramos, March, 1911, on rocks along the river, altitude 75 to 100 meters,
the flowers white and greenish.

A most characteristic species, recognizable by its small size and its
lanceolate, sharply acuminate leaves. The narrow leaves have doubtless
been developed to meet the exigencies of its habitat, for the plant undoubt-
edly grows in situations subject to overflow during heavy rains. It has
almost exactly the habit and appearance of Atalantia linearis (Blanco)
Merr., and Eugenia mimica Merr., which grow in similar habitats, and in
fact in making the preliminary identifications of the Guinatacutan collec-
tion, the specimens were referred to Atalantia linearis.

Stenophylly, due to habitat, is not as highly developed in the Philippines
as in the neighboring Island of Borneo, and aside from the widely distrib-
uted Homonia riparia Lour., we have the endemic species Atalantia linearis
Merr., Eugenia mimica Merr., Buxus rivularis Merr., and Ficys rivularis
Merr., and doubtless others remain to be noted. The rather numerous
Bornean forms have been considered by Beccari.*

BUXUS PACHYPHYLLA sp. nov.

Arbor glabra; foliis crassissime coriaceis, nitidis, oblongis,
usque ad 11 cm longis, 3 ad 5.5 cm latis, basi acutis vel acumina-
tis, apice acute acuminatis, margine valde revolutis; capsulis
axillaribus, solitariis, breviter pedunculatis, ovoideis, circiter 1
cm longis.

A glabrous tree, size not indicated. Branches terete, pale-
olivaceous, the branchlets slender, somewhat sulcate on two sides.
Leaves oblong, very thickly coriaceous, 9 to 11 cm long,
3 to 5.5 cm wide, narrowed below to the acute or somewhat
acuminate base and above to the sharply acuminate apex, the
margins strongly recurved, both surfaces shining, the lower
usually a little paler than the upper; lateral nerves slender, up
to 30 on each side of the midrib, irregular, sometimes rather
indistinct and confused with the secondary ones and the reticula-
tions; petioles 5 to 7 mm long. Peduncles solitary, axillary, in
fruit 5 mm long or less, with numerous, spreading bracteoles,
the lower ones smaller than the upper. Flowers not seen.
Capsules ovoid, about 1 cm long, solitary, smooth, somewhat
glaucous-purple when dry.

LUZON, Province of Tayabas, Mount Cadig near Guinayangan, Bur. Sei.
20828 Escritor, March 9, 1918.

Well characterized by its very thickly coriaceous leaves, and its solitary,
few, short-peduncled fruits.

* Nelle Foreste di Borneo (1902) 524.

1% 6, 4 Merrill: Philippine Plants, X 311

BUXUS LOHERI sp. nov.

Arbor glabra; foliis lanceolatis, coriaceis, usque ad 6 cm longis,
utrinque angustatis acuminatisque, in siccitate brunneis, nitidis,
margine revolutis, nervis primariis utrinque 15 ad 20, tenuibus,
obscuris; capsulis terminalis axillaribusque, solitariis vel binis,
circiter 1 cm longis; pedunculo circiter 1 cm longo.

A glabrous tree, size not indicated. Branches terete, pale-
gray or somewhat brownish, the younger branchlets slender, dark
reddish-brown, somewhat angled or sulcate. Leaves lanceolate,
thickly coriaceous, 4.5 to 6 cm long, 1.5 to 2 em wide, when dry
brownish and shining on both surfaces, the lower surface usually
a little paler than the upper, about equally narrowed and acumi-
nate at both ends, the margins strongly revolute; lateral nerves
slender, not prominent, in fact often indistinct, 15 to 20 on each
side of the midrib, irregular, usually more or less confused by
the secondary ones. Flowers unknown. Capsules ovoid, about
1 cm long, solitary or in pairs, terminal and in the leaf-axils,
the peduncles about 1 cm long, each with several scattered brac-
teoles, the calyx, in fruit, about 7 mm in diameter, the lobes 5,
broadly ovate.

Luzon, Province of Rizal, Montalban, Loher 6857, February, 1905.

A species probably as closely allied to Buxus rolfei Vid. as to any other
species, but distinguished by its narrow, brown, lanceolate, much smaller,
obscurely nerved leaves.

CELASTRACEAE
GYMNOSPORIA Bentham & Hooker

GYMNOSPORIA NITIDA sp. nov.

Frutex vel arbor parva, glabra, inermis; foliis ellipticis ad
oblongo-obovatis, coriaceis, supra valde nitidis, usque ad 7 cm
longis, apice obtusis vel rotundatis, basi acutis, margine crenu-
latis; capsulis anguste obovoideis vel oblongo-ellipsoideis, circiter
1.5 cm longis, valvis crassissimis.

A shrub or small tree, quite glabrous, unarmed. Branches
pale-gray, slender, somewhat wrinkled and shining when dry.
Leaves rather thickly coriaceous, elliptic to oblong-obovate, 4 to
7 cm long, 2 to 3.5 em wide, the upper surface strongly shining
when dry, the lower much duller, apex obtuse or rounded, base
acute, margins crenulate; lateral nerves about 8 on each side of
the midrib, slender, not prominent, rather laxly anastomosing;
petioles 8 to 10 mm long. Inflorescence axillary, apparently
cymose, in fruit up to 3 cm long. Flowers unknown. Capsules

312 The Philippine Journal of Science 1914 .

narrowly obovoid to oblong-ellipsoid, about 1.5 cm long, 8 to 10
mm in diameter, rounded at the apex, the valves 3, very thick.
Luzon, Province of Pangasinan, Salasa, For. Bur. 11836 Domingo,
November 20, 1912.
Well characterized by its strongly shining leaves and its very thick
capsule-valves. Not closely allied to Gymnosporia spinosa Merr. & Rolfe,
the only other known Philippine species.

EUONYMUS Linnaeus

EUONYMUS VIBURNIFOLIUS (Juss.) comb. nov.
Aegiphila viburnifolia Juss. in Ann. Mus. Paris 7 (1806) 76; Walp.
Repert. 4 (1844-48) 124; Schauer in DC. Prodr. 11 (1847) 655.
Euonymus philippinensis Merr. in Philip. Journ. Sci. 3 (1908) Bot. 238.

Jussieu’s species has long been a doubtful one, for manifestly it could
not belong in the genus Aegiphila which is confined to tropical America.
The type, Commerson, in Herb. Mus. Paris, was from the Philippines. The
original specimen was examined and photographed by the late Dr. C. B.
Robinson in November, 1911, and from an examination of the photograph
and notes I am now able definitely to refer the species to the Celastraceae,
and very definitely to the species I described a few years ago as Euonymus

philippinensis.
ICACINACEAE
MIQUELIA Meissner

MIQUELIA RETICULATA sp. nov.

Scandens, partibus junioribus subtus foliis inflorescentiisque
leviter breviter hirsutis; foliis oblongis, chartaceis, usque ad 17
em longis, in siccitate pallidis, nitidis, apice tenuiter acuminatis,
basi rotundatis leviter cordatisque, nervis lateralibus circiter 6,
subtus cum reticulis laxis valde prominentibus; floribus 9?
umbellatis, 4-meris, 3.5 mm longis.

A scandent, apparently woody plant, the branchlets terete,
pale when dry, slender, sparingly hirsute with short hairs.
Leaves oblong, entire, 14 to 17 em long, 4 to 6 cm wide, pale and
somewhat shining when dry, the upper surface smooth and
glabrous, the lower prominently reticulate and sparingly hirsute,
the apex slenderly and prominently acuminate, somewhat
narrowed below to the rounded and slightly cordate base; basal
nerves 3 pairs, the lower two pairs very short, the lateral nerves
above the base 5 or 6 on each side of the midrib, anastomosing,
very prominent on the lower surface, the reticulations lax,
prominent; petioles pale, 1.5 to 8 cm long, very sparingly hirsute.
Female flowers umbellate, on slender, axillary, simple peduncles
at anthesis about 3 cm long, elongated in fruit, about 10 in
each umbel, the pedicels sparingly pubescent, slender, 6 to 8
mm long. Calyx very minute or nearly obsolete. Petals 4,

IX, ©, 4 Merrill: Philippine Plants, X 313

oblong, 3.5 mm long, 1.2 mm wide, slightly pubescent outside,
apex acute or obtuse, more or less inflexed. Staminodes none.
Ovary oblong-ovoid, somewhat hirsute, 2 mm long; stigma shal-
lowly cup-shaped, about 1.3 mm in diameter. Fruit narrowly
ovoid, somewhat compressed, about 2 cm long, 1.3 cm wide,
wrinkled when dry, the pseudostipe about 8 mm long, the per-
sistent petals recurved, pulp scanty, the endocarp crustaceous,
coarsely foveolate.

CAMIGUIN DE MINDANAO, Panatayuan, Bur. Sci. 14674 Ramos, March
28, 1912, in forests, the flowers greenish-yellow.

Quite distinct from our other Philippine species, Miquelia cumingii Baill.,
and readily recognizable by its prominently reticulate leaves and its sparse
pubescence of short, pale, hirsute hairs.

PLATEA Blume

PLATEA PHILIPPINENSIS sp. nov.

Platea latifolia Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 58, non

Blume.

Species P. latifoliae Bl. affinis, differt foliis minoribus, basi
acutis, partibus junioribus inflorescentiisque ferrugineo-lepidotis,
vix tomentosis, fructibus minoribus.

A dioecious tree 8 to 20 m high, the branches terete, nearly
black when dry, glabrous, the branchlets densely lepidote, the
scales minute, older ones pale, the younger ones ferrugineous.
Leaves oblong-ovate, coriaceous 7 to 12 cm long, 2.5 to 6.5 cm
wide, the apex rather prominently acuminate, the base acute,
the upper surface dark-colored when dry, glabrous, somewhat
shining, the lower pale, densely lepidote; lateral nerves 10 to
12 on each side of the midrib, prominent on the lower surface,
darker colored than the surface itself, usually anastomosing;
petioles 2 to 3 cm long, more or less lepidote. Male panicles 3
to 6 cm long, narrow, interrupted, the rachis, branches, and
calyces ferruginous-lepidote, the scales somewhat ciliate, not
tomentose, the lower branches 2 cm long or less, the upper ones
reduced to fascicles of flowers. Flowers sessile, glomerate,
numerous, the buds globose or depressed-globose. Calyx about
2 mm in diameter, the sepals nearly free, ovate, acute, about 1
mm long. Petals narrowly ovate or ovate-lanceolate, slightly
united at the base, glabrous, 2.5 to3 mm long. Anthers broadly
elliptic, about 1.2 mm long. Male racemes axillary, solitary,
about 2 cm long, 3- to 5-flowered, ferruginous-lepidote, the
pedicels very stout, 2 to 3 mm long. Calyx somewhat cup-
shaped, the lobes very broadly ovate or orbicular-ovate, about
2.5 mm long, 3 mm wide. Ovary densely stellate-lepidote, the

314 The Philippine Journal of Science 1914

thick and short style as broad as the ovary, glabrous. Fruit
narrowly ovoid, gradually narrowed upward to the acute or
obtuse apex, 2.5 to 3 cm long, apparently black when mature,
the pulp scanty, the endocarp hard, deeply longitudinally pitted
and sulcate.

LuzON, Province of Bataan, Mount Mariveles, Hlmer 6835, November,
1904, in fruit; For. Bur. 2098 Borden, November, 1904, in fruit; Whitford
1202 (type), March, 1905, with pistillate flowers. LryYTE, Dagami, Bur.
Sci. 15242 Ramos, August, 1912, in fruit. CAMIGUIN DE MINDANAO, Bur.
Sci. 14651 Ramos, March, 1912, with staminate flowers. NEGROS, Cuernos
Mountains, Elmer 9777, March, 1908, with staminate flowers.

The material from Mount Mariveles, consisting of specimens with pistil-
late flowers and fruits, was originally referred by me to the Javan Platea
latifolia Blume, to which the Philippine form is manifestly closely allied.
On receipt of staminate material all the specimens were reéxamined with
the result that it has been considered advisable to describe our local form
as a distinct species. The original description of Platea latifolia Blume
Bijdr. (1826) 647 is quite inadequate, and the later one by Miquel FI.
Ind. Bat. 1:* 793 is also too short. Koorders,* however, gives an excellent
and detailed description of the Javan species of the genus, on which my
conception of Platea latifolia Blume is based. Sterile specimens from Java
are also in the herbarium of the Bureau of Science. Platea philippinensis
grows in forests at and above an altitude of 900 meters.

STERCULIACEAE
FIRMIANA Marsigli

FIRMIANA MERRITTII sp. nov.

Arbor alta; foliis orbiculari-ovatis vel late ovatis, acuminatis,
integris, basi latissime rotundatis, profunde et anguste cordatis,
vetustioribus glabris, usque ad 18 cm longis; folliculis 8 ad 9
em longis, circiter 4 cm latis, inflatis, chartaceis, laxe reticulatis,
tarde dehiscentibus.

A large tree, reaching a height of 30 m and a trunk diameter
of 90 em, deciduous. Leaves broadly ovate to orbicular-ovate,
coriaceous or thickly chartaceous, in maturity glabrous, up to 18
em long and about as wide, entire, the apex acuminate, the base
very broadly rounded, deeply and narrowly cordate, palmately
7- or 9-nerved, the reticulations distinct; petioles 20 cm long.
Follicles inflated, tardily dehiscent, narrowly oblong-ovate, base
acute, apex obtuse, 8 to 9 cm long, 4 cm wide, the valves laxly
reticulate, glabrous or nearly so, thickly chartaceous, not mem-
branaceous or coriaceous, each containing 1 or 2 seeds.

MINDORO, Igsoro River, west coast, For. Bur. 8555 Merritt, February 4,
1908, in forested river flats, altitude about 10 meters.

*Meded. ’s Lands Plantent. 33 (1900) 156.

IX, ©, 4 Merrill: Philippine Plants, X 315

The first species of the genus to be found in the Philippines, and remark-
able among the few members of the genus in its very tardily dehiscent
follicles. In my material, which consists of fallen leaves and follicles, the
seeds are nearly mature, yet the follicles scarcely show a sign of opening.

MELOCHIA Dillenius

MELOCHIA UMBELLATA (Houtt.) comb. nov.

Visenia umbellata Houtt. Handl. 8 (1777) 309.

Wisenia indica Gmel. Syst. 2 (1791) 515.

Melochia arborea Blanco FI. Filip. (1837) 524.

Melochia indica A. Gray ex F.-Vill. Novis. App. (1880) 29; K. Sch. in
Engl. Bot. Jahrb. 9 (1887) 209.

This widely distributed and much-named plant has a peculiarly compli-
cated synonymy, and for the last twenty years has been considered by many
botanists, after K. Schumann, under a specific name that was neither pub-
lished by the original author Houttuyn under Visenia, as V. indica, and
was never transferred to Melochia, as M. indica, by A. Gray until the
transfer was made by F.-Villar and K. Schumann and wrongly credited to
Gray. Houttuyn in 1777 published the species as Visenia umbellata, and
this is apparently the oldest valid specific name. Gmelin seems to have
made the first use of the specific name indica, for he publishes it as Wisenia
indica with a reference to Christmann and Panzer’s German edition of
Houttuyn’s work Vol. 6 (1780), where, however, the species appears as
Visenia umbellata. Gmelin, then, simply proposed a new specific name,
indica, to replace that proposed by Houttuyn. Hasskarl® seems to have
been the first author to credit the combination Visenia indica to Houttuyn,
whieh he later repeated in his Platae Javanicae Rariores, from whence it
passed into Miquel’s Flora Indiae Batavae and other works. K. Schumann
manifestly took up the specific name from Miquel.

Asa Gray never published the combination “Melochia indica (Houtt.)
A. Gray” in the Botany of the Wilkes Expedition as credited to him by
K. Schumann° but simply indicates that: “Visenia cannot be generically
distinguished from Melochia.” K. Schumann was, hence, in error both in
taking up the specific name indica, and in crediting its transfer to Melochia
to A. Gray. Visenia wmbellata Houtt. seems to supply the correct specific
name, under Melochia, for this very common and widely distributed species,
and is accordingly here taken up.

Fernandez-Villar is the first author actually to make the combination
Melochia indica, but his publication of the combination has been entirely
overlooked by later authors, and does not appear in Index Kewensis. In
the Novissima Appendix to the third edition of Blanco’s Flora de Filipinas
(1880) 29 the name Melochia indica appears, but is erroneously credited
to A. Gray on the authority of Bentham & Hooker f. Gen. Pl. 1 (1862)
224. Bentham & Hooker f., however, do not make the transfer, but simply
state: “Cetera omnia Riedleiae ceonveninunt et monente Grayo Viseniam
pro sectione Melochiae potius quam genere proprio habemus.”

Other synonyms of this species are: Visenia tomentosa Miq., Riedleia
tiliaefolia DC., R. velutina DC., Glossospermum velutinum Wall., G. cor-

* Tijdschr. Nat. Gesch. 12 (1845) 122.
“Engl. Bot. Jahrb. 9 (1887) 209.

316 The Philippine Journal of Science 1914

datum Wall., Aleurodendron album Reinw., Melochia velutina Bedd., and
Hypericum pentandrum Blanco.

I am indebted to Sir D. Prain, director, Royal Gardens, Kew, for refer-
ences from the publications of Houttuyn and Gmelin, which are not avail-
able in Manila, and for a memorandum covering the case as to the oldest
specific name for the species.

BUETTNERIA Linnaeus

BUETTNERIA ECHINATA Wall. Cat. (1829) no. 1149; Gagnep. in Le-
comte Fl. Gén. Indo-China 1 (1911) 520.

MINDORO, near Calapan, Bur. Sci. 21268 Escritor, July, 1913.

The identification has been made after Gagnepain, the Mindoro specimen
agreeing well with Pierre 3746 from Tay-ninh, Cochin China, so named by
Gagnepain, and with his description of the species. Masters refers the
Wallichian species with doubt to Buettneria crenulata Wall., while Gagne-
pain cites the latter as a synonym of B. echinata. The genus is new to the
Philippines.

Nepal and Burma to Indo-China.

THEACEAE
ADINANDRA Jack

ADINANDRA ROSTRATA sp. nov.

Arbor circiter 30 m alta partibus junioribus exceptis glabra;
foliis subcoriaceis, nitidis, oblongo-ovatis vel oblongo-ellipticis,
integris, usque ad 14 cm longis, utrinque angustatis, apice acumi-
natis, basi acutis vel decurrento-acuminatis, nervis utrinque
circiter 14, tenuibus; fructibus axillaribus, solitariis, longe
pedunculatis, ovoideis, 1.5 em longis, longe caudato-rostratis.

A tree about 30 m high, glabrous except the branchlets (flowers
not seen). Branches terete, brownish, glabrous, the branchlets
sparingly pilose, the tips rather densely so. Leaves entire,
subcoriaceous, the younger ones thinner, oblong-ovate to oblong-
elliptic, 8 to 14 cm long, 3.5 to 6.5 cm wide, narrowed at both
ends, the apex acuminate, the base acute or decurrent-acuminate,
shining, of about the same color on both surfaces and somewhat
yellowish when dry; lateral primary nerves about 14 on each side
of the midrib, slender, anastomosing, the reticulations rather lax;
petioles 5 to 10 mm long. Flowers not seen. Fruits axillary,
solitary, their peduncles 4 cm long, the persistent sepals subor-
bicular to subreniform, coriaceous, glabrous, rounded, about
8 mm wide, the outer ones somewhat smaller than the inner,
the fruit ovoid, about 1.5 cm long, 1.2 to 1.4 mm thick, glabrous,
prominently caudate-rostrate, the beak 1 to 1.4 cm long, 3-celled,
the pericarp thick. Seeds two or three in each cell, about 6

Tm, C, 4 Merrill: Philippine Plants, X 317

mm long, flattened, reddish-brown, obovate, shining, the cotyle-
dons hippocrepiform.

Luzon, Province of Laguna, Dahican River, Phil. 1567 Ramos, September
18, 1912, in forests.’

A species characterized by its few, comparatively large seeds, in which
it differs from most species in the genus. It is possible that when flowers
are known it will be necessary to transfer the species to some other genus,
but in general appearance and in all characters, perhaps other than the
seed, it is apparently an Adinandra.

ADINANDRA MAQUILINGENSIS sp. nov.

Arbor parva, ramulis foliisque junioribus pilosis, vetustioribus
glabris vel subglabris; foliis coriaceis, 3.5 ad 6 cm longis, nitidis,
oblongo-ellipticis ad oblongo-obovatis breviter acuminatis, basi
acutis, margine in 4} superiore crenulato-denticulatis, nervis
utrinque circiter 12, tenuibus; sepalis 5, subaequalibus, oblongo-
ovatis, obtusis, 8 mm longis; ovario leviter piloso, 5-loculare.

A tree about 12 m high, the very young branchlets and young
leaves rather softly pilose with appressed hairs, the older parts
glabrous or nearly so. Branches terete, very dark-gray or nearly
black, rather smooth, the branchlets brownish. Leaves coria-
ceous, 3.5 to 6 cm long, 1.5 to 2.5 em wide, oblong-elliptic to
oblong-obovate, the base acute, the apex shortly acuminate, the
margins in the upper one-half crenulate-denticulate, below entire,
both surfaces shining, the lower a little paler than the upper,
yellowish-green when dry; lateral nerves about 12 on each side
of the midrib, slender, distinct but not prominent, about equally
evident on both surfaces, anastomosing; petioles 3 to 5 mm long.
Flowers axillary, solitary, the pedicels stout, curved, glabrous,
about 2 cm long. Sepals subequal, oblong-ovate, 8 mm long,
5 mm wide, obtuse, somewhat appressed-pubescent. Petals and
stamens not seen. Very immature fruit ovoid, 7 mm long, 5-
celled, sparingly pilose, the style 8 mm long. Ovules very
numerous in each cell, apparently but few developing into seeds.

Luzon, Province of Laguna, Mount Maquiling, in forests on the upper
slopes, altitude probably about 1,000 m, Bur. Sci. 13650 Ramos, September,

1911, from the Batangas side of the mountain.
A species well characterized by its comparatively small leaves.

ADINANDRA CORIACEA sp. nov.

Arbor glabra; foliis integris, oblongo-ellipticis, coriaceis, 12
ad 15 cm longis, basi acutis, apice latissime breviter obtuse
acuminatis, nervis utrinque circiter 15, distinctis, tenuibus, pe-
tiolo circiter 1 cm longo; fructibus brunneis, ovoideis, laevis,
apiculatis, circiter 1.5 em longis, 2-locellatis; seminibus 10 ad

318 The Philippine Journal of Science 1914

12, planis, anguste ovoideis, utrinque sulcatis, nitidis, 6 mm
longis; sepalis persistentibus, margine leviter ciliatis.

A glabrous tree, size not indicated. Branches terete, reddish-
brown or grayish, smooth. Leaves coriaceous, entire oblong-
elliptic, 12 to 15 cm long, 4 to 6.5 em wide, subequally narrowed
at both ends, the base acute, the apex very broadly, shortly, and
bluntly acuminate, the upper surface pale-olivaceous when dry,
the lower brownish, paler, both slightly shining and very minutely
puncticulate, glabrous; lateral nerves about 15 on each side of
the midrib, slender, rather distinct, about equally prominent on
both surfaces, anastomosing, forming a double row of arches,
the reticulations lax, indistinct; petioles stout, about 1 cm long.
Flowers not seen, the pedicels in fruit stout, 2 to 2.5 cm long,
the persistent calyx lobes coriaceous, at first apparently some-
what pubescent, becoming quite or nearly glabrous, the outer
two smaller than the inner three which are more or less reni-
form, entire or nearly so, somewhat reniform or very broadly
rounded-ovate, about 8 mm wide, margins deciduously ciliate, in
age glabrous or nearly so. Fruit ovoid, about 1.5 cm long,
smooth, brown, somewhat shining, entirely glabrous, 2-celled, the
pericarp rather thick, dry and brittle in texture. Seeds 10 to
12, brown, shining, flattened, narrowly obovoid, rounded at the
apex, about 6 mm long, longitudinally grooved along the middle
of both faces.

Luzon, Province of Laguna, Cavinti, For. Bur. 19667 Racelis, August,
1912.

A species manifestly closely allied to the Malayan Adinandra acuminata
Korth., from which it differs in its differently shaped, very broadly and
obtusely acuminate, more numerously nerved leaves, longer petioles, and
other characters.

ADINANDRA LOHERI sp. nov.

Arbor, gemmis petalis staminibusque exceptis glabra; foliis
ellipticis, coriaceis, usque ad 10 cm longis, basi decurrento-acumi-
natis, apice late rotundatis ad brevissime late acuminatis, mar-
gine crenulatis vel denticulato-crenulatis, subtus obscurissime
glanduloso-maculatis, nervis utrinque tenuibus, distinctis, circiter
15; floribus 5-meris; sepalis glabris; petalis extus in partibus
medianus villosis, 1.8 em longis; staminibus circiter 50, leviter
hirsutis; ovario 5-loculare, glaber.

A tree, quite glabrous except the vegetative buds and some
parts of the flowers. Branches terete, grayish or dark-colored,
the young branchlets reddish-brown, the vegetative buds pale-
villous. Leaves coriaceous, elliptic, 6 to 10 cm long, 3.5 to 5

we C74 Merrill: Philippine Plants, X 319

em wide, the apex broadly rounded to broadly and shortly obtuse-
acuminate, base somewhat decurrent-acuminate, margins crenu-
late or crenulate-denticulate, the upper surface smooth and
shining, dark-olivaceous when dry, the lower surface paler,
brown, obscurely glandular-maculate; lateral nerves about 15
on each side of the midrib, slender, distinct, anastomosing, more
prominent on the lower than on the upper surface; petioles 5 to 7
mmlong. Flowers 5-merous, axillary, solitary, the peduncles gla-
brous, curved, stout, about 2 cm long. Calyx glabrous, the lobes
orbicular to orbicular-reniform, broadly rounded, coriaceous,
concave, about 7 mm long, 6 to 8 mm wide, margins sometimes
slightly ciliate, not glandular. Petals obovate, broadly rounded,
about 1.8 cm long, 1.3 cm wide, the exposed median portions of
the back rather densely appressed-pubescent with pale hairs
otherwise glabrous. Stamens about 50; filaments 4 to 8 mm
long, slightly hirsute; anthers oblong-lanceolate, acute or acumi-
nate, slightly hirsute, 4 mm long. Ovary ovoid, glabrous,
narrowed upward into the glabrous style, 5-celled, the ovules
indefinite.

Luzon, Province of Rizal, Oriud, Loher 5604, December, 1905.

A characteristic species, apparently allied to Adinandra lamponga Miq.,
but differing in many details. It is well characterized, among the Phil-
ippine forms, by being nearly glabrous, the only pubescent parts being the
very small vegetative buds, the median parts of the petals, outside, and the
stamens.

ADINANDRA MACGREGORII sp. nov.

Arbor circiter 15 m alta, plus minusve villosa; foliis coriaceis,
ellipticis ad oblongo-ellipticis, usque ad 10 cm longis, supra
nitidis, gabris, subtus adpresse villosis, basi acutis ad subro-
tundatis, apice breviter acuminatis, margine distincte minute-
que glanduloso-denticulatis, nervis utrinque 12 ad 15, distinctis;
sepalis extus villosis, petalis late ellipticis, rotundatis, circiter
1.4 em longis; staminibus 35, antheris parce villosis; ovario
4-loculare, dense pallide villoso.

A tree about 15 m in height, the buds and growing branchlets
very densely appressed-villous with pale hairs, the petioles, lower
surfaces of the leaves, and flowers less densely villous. Branches
terete, grayish, glabrous. Leaves coriaceous, elliptic to oblong-
elliptic, 5 to 10 cm long, 2.5 to 4 cm wide, apex shortly acuminate,
base somewhat rounded to somewhat acute, margins very
minutely glandular-denticulate, the upper surface olivaceous
when dry, glabrous and shining, the lower much paler, with
scattered, more or less appressed, pale hairs; lateral nerves

320 The Philippine Journal of Science 1914

12 to 15 on each side of the midrib, slender, distinct, anasto-
mosing, about equally evident on both surfaces; petioles densely
villous, 3 mm long. Flowers axillary, solitary, 5-merous, their
peduncles stout, curved, somewhat villous, about 2.5 cm long.
Sepals about 1 cm long, 7 mm wide, coriaceous, ovate, acute or
slightly acuminate, glabrous inside, outside somewhat densely
appressed-villous with pale, short hairs, the margins minutely
glandular-denticulate. Petals broadly elliptic to elliptic-ovate,
rounded, not at all retuse, 1.4 cm long, 1 cm wide, externally
appressed-villous in the exposed median parts, inside and the
broad margins externally glabrous. Stamens 35; filaments
glabrous, 4 to 5 mm long; anthers lanceolate, acuminate, 5 mm
long, very sparingly villous with long stiff hairs on the back.
Ovary ovoid, densely villous, 4-celled, the ovules indefinite; style
stout, about 8 mm long, widened below to the ovary, prominently
villous except near the glabrous apex, cleft into four 1.5 to 2 mm
long arms, the stigmas capitate.

Luzon, Benguet Subprovince, Pauai, Bur. Sci. 8425 McGregor, June,
1909, in forests, altitude above 2,100 m.

A species manifestly allied to Adinandra elliptica C. B. Rob., but distin-

guishable at once by its much larger flowers, as well as by numerous details
in vegetative and floral characters.

ADINANDRA NIGRO-PUNCTATA sp. nov.

Arbor circiter 10 m alta, subglabra;-foliis coriaceis, ellipticis
ad oblongo-obovatis, 4 ad 7 cm longis, nitidis, basi acutis, apice
late acuminatis vel obtusis, minute retusis, margine minute
glanduloso-denticulatis, subtus glandulis minutis sparsis nigris
instructis, nervis utrinque circiter 7; sepalis 5, subaequalibus,
ellipticis, glabris; staminibus circiter 50, densissime hirsutis;
ovario glabro, 5-loculare.

A tree about 12 m in height, glabrous except the flowers and
the growing tips of the branchlets. Branches stout, terete,
reddish-brown, glabrous, the terminal buds pubescent. Leaves
coriaceous, elliptic to oblong-obovate, 4 to 7 cm long, 1.5 to 3
em wide, the base acute, the apex very broadly and shortly
blunt-acuminate, sometimes merely obtuse, usually minutely
retuse, margins very obscurely and minutely glandular-denticu-
late, in the lower part quite entire, the upper surface somewhat
olivaceous when dry, strongly shining, usually minutely ver-
ruculose, the lower surface a little paler, with small, scattered,
black glands; lateral nerves about 7 on each side of the
midrib, slender, not prominent, about equally evident on
both surfaces; petioles 5 mm long. Flowers axillary, soli-

IX, C, 4 Merrill: Philippine Plants, X 821

tary, white, their pedicels glabrous, stout, about 2 cm long.
Sepals subequal, glabrous, elliptic, broadly rounded, usually
somewhat retuse, 10 mm long, 8 mm wide. Petals broadly ellip-
tic, about 1.8 cm long, 1 to 1.3 ecm wide, coriaceous broadly
rounded, slightly retuse, the median exposed parts very densely
pale-hirsute, otherwise glabrous. Stamens about 50; filaments
5 mm long, densely hirsute; anthers ovate-lanceolate, somewhat
acuminate, 4 mm long, densely hirsute. Ovary glabrous, ovoid,
5-celled, ovules very numerous in each cell; style glabrous, 12
mm long, the stigma minute, subcapitate.

LEYTE, Dagami, in forests, Bur. Sci. 15355 Ramos, August 13, 1912.

A characteristic species, distinguishable by its rather large flowers which
are glabrous except for the densely hirsute median parts of the petals exter-
nally and the stamens, subequal broadly elliptic petals, subequal broadly
elliptic sepals, and its vegetative characters; the scattered, minute, black
glands on the lower surface are characteristic.

DILLENIACEAE
DILLENIA Linnaeus

DILLENIA MONANTHA sp. nov. § Wormia.

Arbor glabra, usque ad 12 m alta; foliis coriaceis, ellipticis ad
obovato-ellipticis, usque ad 12 cm longis, obtusis, acutis, vel
latissime breviter acuminatis, basi acutis, decurrentibus, margine
subintegris vel distanter irregulariter dentatis, nervis utringque
6 ad 8; floribus flavidis, in axillis superioribus, solitariis, circiter
10 cm diametro, sepalis extus cinereo-pubescentibus; carpellis 5,
leviter hirsutis; staminibus interioribus quam exterioribus mul-
to longioribus, antheris poro terminalibus dehiscentibus.

A glabrous tree reaching a height of 12 m, the branches terete,
rugose, reddish-brown or grayish. Leaves alternate, coriaceous,
rather pale when dry, shining, elliptic to elliptic-obovate, 8 to 12
em long, 5 to 8 cm wide, the apex subacute, obtuse, or broadly
and shortly acuminate, the base acute, the lamina decurrent on
the petiole as narrow wings, the margins subentire to distantly
dentate; lateral nerves 6 to 8 on each side of midrib, curved,
anastomosing, distinct; petioles stout, narrowly decurrent-
winged, 1 to 1.5 cm long. Flowers solitary in the upper axils,
yellow, about 10 cm in diameter, their peduncles 2 to 4 cm long.
Sepals 5, elliptic to elliptic-obovate, rounded, concave, coriaceous,
1.5 to 2 cm long, 1 to 1.4 cm wide, outside more or less cinerous-
pubescent. Petals yellow, obovate, membranaceous, about 5 cm
long, 3.5 cm wide. Stamens very numerous, the interior ones up
to 1.8 cm long, the exterior ones gradually shorter, the outermost

322 The Philippine Journal of Science 1914

8 to 10 mm long, the anthers opening by terminal pores. Car-
pels 5, slightly hirsute, 7 mm long, oblong, subcylindric; styles
5, about 1 cm long; ovules about 16, 2-seriate. Fruit not seen.

PALAWAN, Taytay, Merrill 9287 (type), flowering from April 10 to June
and probably later, 1913; Malampaya Bay, Bur. Sci. 21555 Escritor, August,
1913, For. Bur. 4518 Curran, June 21, 1918. DUMARAN, Bur. Sct. 21642
Escritor, August, 1918. CuLion, Bur. Sci. 15647 Fénix, July 11, 1918.

This species is common at low altitudes in northern Palawan, growing
in thin, second growth forests, and in and along the borders of open cogo-
nales, that is, areas occupied by the cogon or lalang grass (Imperata cylin-

drica). It is, perhaps, most closely allied to the Malayan Dillenia pulchella —

(Jack) Gilg, and among the Philippine species to Dillenia sibuyanensis
(Elm.) (Wormia sibuyanensis Elm.). Its solitary flowers, together with
its somewhat decurrent laminae and its few carpels are striking differential
characters.

LECYTHIDACEAE
BARRINGTONIA Forster

BARRINGTONIA PTERITA sp. nov.

Arbor parva, glabra; foliis ad apices ramulorum plus minusve
confertis, oblanceolatis, sessilibus vel subsessilibus, usque ad 40
cm longis, acuminatis, basi sensim angustatis; racemis circiter
70 cm longis, pendulis; fructibus oblongis, 6 cm longis, longi-
tudinaliter 4-alatis.

A small glabrous tree, 10 m high fide Ramos. Branches terete,
the ultimate ones about 6 mm in diameter, above with rather
prominent petiolar scars. Leaves somewhat crowded at the ends
of the branchlets, oblanceolate, 20 to 40 cm long, 4 to 8 em
wide, entire, rather pale when dry, somewhat shining, charta-
ceous, the apex rather sharply acuminate, gradually narrowed
from above the middle to the base, the base 1 cm wide or less,
then abruptly acute or rounded, the petiole none, or stout and
4 mm long or less; lateral nerves about 20 on each side of the
midrib, rather prominent on the lower surface. Racemes pend-
ulous, about 70 cm long, the flowers unknown. Fruit oblong
or narrowly oblong, 6 cm long, 1.5 to 2.5 cm wide, the base acute
the pedicels about 1 cm long, the persistent sepals crowning the
apex about 1 cm in length, the four angles longitudinally winged,
the wings subcoriaceous, about 5 mm wide.

LUZON, Province of Laguna, Dahican River back of San Antonio, Bur.
Sci. 15121 Ramos, June, 1912.

A species manifestly allied to Barringtonia racemosa Blume, but at once
distinguishable by its prominently longitudinally 4-winged fruits, in this
character differing from all the previously known Philippine species. Elmer

9168 from Lucban, Tayabas Province, is undoubtedly a small leaved form
of the same species; it was distributed as Barringtonia racemosa Blume.

cow one Merrill: Philippine Plants, X 323
FLACOURTIACEAE

HYDNOCARPUS Gaertner

HYDNOCARPUS CAULIFLORA sp. nov. § Huhydnocarpus, Oliganthera.

Arbor parva inflorescentiis exceptis glabra; foliis late oblongis,
chartaceis vel tenuiter coriaceis, usque ad 30 cm longis, nitidis,
acuminatis, basi late cordato-rotundatis, nervis utrinque circiter
15, valde prominentibus, petiolo vix 1 cm longo; inflorescentiis
fulvo-villosis, usque ad 15 cm longis, anguste paniculatis vel
racemosis, caulinis.

A small tree, glabrous except the inflorescence. Branches
slender, terete. Leaves alternate, broadly oblong, chartaceous
or thinly coriaceous, of the same color and shining on both sur-
faces when dry, 23 to 30 cm long, 9 to 12 cm wide, the base
broad, rounded, somewhat cordate, the apex shortly acuminate;
lateral nerves about 15 on each side of the midrib, very
prominent, somewhat curved, anastomosing near the margin,
the primary reticulations rathers lax, the ultimate ones
slender, rather dense; petioles stout, 5 to 8 mm _ long.
Inflorescence of very narrow raceme-like panicles or of simple
racemes fascicled on the trunk, 5 to 15 cm long, densely fulvous-
villous. Male flowers: Pedicels stout, villous, 3 mm long, the
subtending bracteole densely villous, narrowly oblong, 2 mm
long. Sepals 5, free, concave, villous, rounded, about 5 mm long.
Petals 5, free, imbricate, thinner than the sepals, elliptic,
rounded, 3 mm long, somewhat appressed-pilose on the back,
the basal scale cleft, the lobes reflexed, about 1 mm long. Fila-
ments villous-bearded, 1.5 mm long. Anthers ovoid, 1 to 1.2
mm long. Rudimentary ovary villous, small. Female flowers
and fruits not seen.

MINDANAO, District of Cotabato, Lebak, For. Bur. 11799 Whitford, March,
1912, in dipterocarp forests at low altitudes.

A very characteristic species, its inflorescence looking more like that of
Ryparosa than of Hydnocarpus. The structure of its flowers, however,
place it in Hydnocarpus. It is strongly characterized by its broadly oblong
leaves which are prominently nerved and broadly rounded-cordate at the
base, and especially by its cauline inflorescence.

XYLOSMA Forster f.

XYLOSMA LUZONENSIS (Presl) comb. nov.

Prockia luzonensis Pres] Rel. Haenk. 2 (1835) 94.

Xylosma cumingii Clos. in Ann. Sci. Nat. IV. 8 (1857) 252; F.-Vill.
Novis. App. (1880) 18; Vid. Phan. Cuming. Philip. (1885) 94, Rev.
P]. Vasc. Pilip. (1886) 49.

LUZON, without definite locality, Haenke in Herb. Prague (type): Prov-
{nce of Cagayan, For. Bur. 17114 Curran, For. Bur. 18506 Alvarez, For.

324 The Philippine Journal of Science 1914

Bur. 14758, 14748 Darling: Province of Ilocos Norte, Cuming 1250: Prov-
ince of Ilocos Sur, Cuming 1128: Province of Nueva Vizcaya, For. Bur.
15844 Curran & Merritt: Benguet Subprovince, Elmer 6423: Province of
Pangasinan, For. Bur. 8376 Curran & Merritt: Province of Zambales, Bur.
Set. 5042 Ramos, Merrill 2916: Province of Rizal, For. Bur. 1893 Ahern’s
collector: Province of Tayabas (Principe), Merrill 1018.

This species appears to be common and widely distributed in northern
Luzon. The specimens cited above show considerable variation but are
all apparently referable to a single species. Presl’s description was based
on at least two specimens, one in flower, and one in fruit. The type in the
herbarium of the Museum des Ké6nigreichs Bohmen at Prague, consists of a ~
fruiting and a flowering specimen mounted on the same sheet, and is, at
least in part, identical with the later Xylosma cumingii Clos. The earliest
name is here adopted.

XYLOSMA SULUENSIS sp. nov.

Frutex glaber circiter 5 m altus; ramulis pallidis, foliis oblongo-
ovatis vel oblongo-lanceolatis, chartaceis, nitidis, basi acutis,
apice acuminatis, margine integris, usque ad 18 cm longis; race-
mis axillaribus, solitariis, glabris, quam petiolo paullo longiori-
bus, paucifloris; fructibus globosis, 6 ad 8 mm diametro.

A glabrous shrub about 5 m high, the branches and branchlets
slender, terete, very pale, with few, scattered lenticels. Leaves
oblong-ovate to oblong-lanceolate, entire, 9 to 18 cm long, 3.5 to
6 cm wide, chartaceous, shining and of about the same color on
both surfaces, when dry brownish-olivaceous, the base acute, the
apex acuminate, gradually narrowed upward from below the
middle, the base often with one or two small glands on the
margins near the insertion of the petiole; lateral nerves slender,
ascending, about 10 on each side of the midrib; petioles 8 to 10
mm long. Racemes axillary, solitary, glabrous, 2 em long or
less, few-flowered. Flowers unknown. Fruits usually about
five to each raceme, globose, 6 to 8 mm in diameter, crustaceous
when dry, smooth, crowned by the very short style, their pedicels
3 to 4 mm long, the subtending bracteoles ovate-lanceolate, 1.5
mm long or less. Seeds 4, black, concave, about 4 mm wide.

UBIAN ISLAND, Sulu Archipelago, Merrill 5398, October 12, 1906, in
thickets back of the beach.

A species well characterized by its entire leaves with are considerably
larger than are those of our other species, Xylosma luzonensis (Presl)
Merr.

FLACOURTIA Commerson

FLACOURTIA EUPHLEBIA sp. nov.

Frutex spinosus circiter 4 mm altus subtus foliis ad nervos
dense pubescentibus; foliis oblongis, subcoriaceis, usque ad 18
cm longis, basi acutis vel rotundatis, apice breviter acuminatis,
margine distincte crenato-serratis; nervis utrinque circiter 10,

IX, C, 4 Merrill: Philippine Plants, X 325

valde prominentibis, anastomosantibus; fructibus subovoideis,
circiter 1 cm longis, in siccitate circiter 7-sulcatis.

A spiny shrub about 4 m high, the younger branchlets, petioles,
and nerves on the lower surfaces of the leaves more or less
pubescent. Branches terete, reddish-brown or brown, glabrous,
lenticellate, the younger ones with solitary or paired, sharp,
nearly straight, 5 to 10 mm long spines subtending each leaf, or
the spines often scattered along the branchlet. Leaves oblong,
subcoriaceous, brownish when dry and of about the same color on
both surfaces, 12 to 18 cm long, 5 to 8 cm wide, the base acute or
somewhat rounded, usually with a pair of glands near the in-
sertion of the petiole, the apex rather abruptly and shortly
acuminate, margins, except near the base, distinctly crenate-
serrate, the midrib and lateral nerves on the lower surface, and
in younger leaves on the upper surface, rather densely brown-
pubescent; lateral nerves about 10 on each side of the midrib,
very prominent on the lower surface, straight, looped-anas-
tomosing near the margin; petioles pubescent, becoming nearly
glabrous in age, stout, 4 to 12 mm long. Flowers unknown.
Fruit ovoid, fleshy, edible, acid, about 1 cm long, black, when
dry, and rather distinctly 7-sulcate.

MINDANAO, District of Davao, Todaya, Williams 2572, April 4, 1905.

The specimen has been identified as Flacourtia montana Grah., which
species, to a certain degree, it resembles. It is, however, entirely different
in its leaf-venation, and does not agree with any previously described
species of the genus so far as I am able to determine. It is remarkable
for its very prominently veined leaves.

RYPAROSA Blume

RYPAROSA CAULIFLORA sp. nov.

Species R. longipedunculatae Boerl. similis et ut videtur affinis,
differt foliis majoribus, usque ad 35 em longis, late oblongo-
obovatis, apice late rotundatis, nervis utrinque circiter 9.

A shrub or small tree, apparently glabrous except the more
or less puberulent inflorescence. Branches terete, brownish
when dry. Leaves alternate firmly chartaceous, broadly oblong-
obovate, up to 35 cm long and 17 cm wide, shining when dry,
the upper surface brownish-olivaceous, the lower surface pale
and somewhat glaucous, the base acute, the apex broadly
rounded; nerves 9 on each side of the midrib, curved-ascending,
prominent, the reticulations distinct, rather lax; petioles about
9 cm long, geniculate above. Inflorescence apparently racemose,
the racemes solitary, springing from the trunk or from the
larger branches, up to 35 cm in length, cinereous-puberulent.

129553——3

326 The Philippine Journal of Science 1914

Flowers unknown. Immature fruits globose, wrinkled when dry,
subglobose, 1 to 1.5 cm in diameter, gray-puberulent externally.

TINAGO ISLAND, Ahern 416, collected by Quadras between February and
May, 1901, locally known as bunganon.

The species above described was previously determined by me as Rypa-
rosa longipedunculata Boerl., but comparison with authentic material from
specimens cultivated in the Botanical Garden at Buitenzorg, show that it
is a species quite different from that, although probably closely allied to it.

CASEARIA Jacquin

CASEARIA BREVIPES sp. nov. § Pitumba.

Arbor parva, glabra; foliis oblongis vel late oblongo-lanceo-
latis, chartaceis vel subcoriaceis, in siccitate brunneis, usque ad 20
cm longis, subintegris vel distanter minuteque glanduloso-dentic-
ulatis, utrinque angustatis, apice longe acuminatis, basi acutis
vel subrotundatis, brevissime petiolatis, nervis utrinque 8 ad
10; fioribus axillaribus, fasciculatis, 5-meris, pedicellatis. Stami-
nibus 10, staminodeis oblongis, apice capitatis, dense ciliatis.

A small tree, quite glabrous, the branches prominently zigzag,
terete, grayish, rather slender. Leaves oblong to broadly oblong-
lanceolate, 14 to 20 cm long, 4 to 7 cm wide, subentire or with
widely scattered, minute, gland-like, obscure teeth, chartaceous
to subcoriaceous, when dry somewhat brownish, the upper sur-
face slightly shining, the lower a little paler, narrowed at both
ends, the base acute or somewhat obtuse, rarely somewhat
rounded, equilateral, the apex rather long-acuminate, the acumen
blunt; lateral nerves 8 to 10 on each side of the midrib, rather
prominent on the lower surface, somewhat curved-ascending,
anastomosing near the margin, the reticulations distinct; peti-
oles stout, 2 to 3 mm long. Flowers axillary, few, fascicled, on
thickened, prominent, axillary tubercles, 5-merous. Pedicels
glabrous, up to 3 mm long, jointed, each subtended by two
broadly ovate bracteoles about 1 mm long. Sepals 5, broadly
elliptic, rounded, concave, glabrous, membranaceous, 3 to 4 mm
long. Stamens ten, 3 mm long, the tubular part about 1 mm
long, glabrous, the free parts of the filaments and the anthers
each about 1 mm long, the ten, alternating, free staminodes
oblong, shorter than the stamens, the apical parts suborbicular-
capitate, densely ciliate. Ovary narrowly ovoid, about 2 mm
long, narrowed upward to the sessile or subsessile capitate
stigma, glabrous, or with very few scattered hairs. Fruit
unknown,

BASILAN, Bur. Sci. 16118 Reillo, August 31, 1912, along streams.
The species is well characterized by its leaves being narrowed at both

=

IX, C, 4 Merrill: Philippine Plants, X 327

ends, prominently acuminate at the apex and usually acute at the base, the
distant, minute, gland-like marginal teeth, the very short petioles, and the
subcapitate tips of the staminodes.

CASEARIA LOHERI sp. nov.

Arbor parva, 4 ad 6 m alta, partibus junioribus floribusque
exceptis glabra; foliis chartaceis, oblongo-ovatis ad obovatis,
usque ad 18 cm longis, breviter petiolatis, in siccitate brunneis,
integris, basi rotundatis, apice late acuminatis, nervis utrinque
5 vel 6; floribus axillaribus, fasciculatis, paucis, 5-meris, extus
leviter hirsutis. Staminibus 10.

A small tree 4 to 6 m high, quite glabrous except the flowers
and the growing tips of the branchlets which are sparingly
hirsute with short, appressed hairs. Branches slender, terete,
smooth, grayish. Leaves chartaceous, oblong-ovate to obovate,
8 to 18 cm long, 4 to 8 cm wide, entire, the upper surface rather
dark-brown or olivaceous-brown when dry, slightly shining, the
lower somewhat paler, the base usually broad, rounded, some-
times subacute, equilateral or nearly so, the apex with a short,
broad acumen; lateral nerves 5 or 6 on each side of the midrib,
distant, rather prominent on the lower surface, curved or curved-
ascending, anastomosing, the reticulations distinct; petioles 4 to 7
mm long. Flowers 5-merous, axillary, fascicled and few or sub-
solitary, their pedicels very short, jointed, the subtending brac-
teoles 2, reniform or very broadly ovate, somewhat connate,
broadly acuminate, externally somewhat pubescent with short
hairs. Sepals 5, elliptic, rounded, slightly pubescent externally,
concave, 3.5 to 4mm long. Stamens 10, their filaments united
below with the alternating staminodes, the tubular part glabrous,
0.8 mm long, the free parts of the filaments slender, glabrous,
1.4 mm long; anthers oblong-ovate 0.8 mm long. Alternating
staminodes shorter than the stamens, oblong, obtuse, 1 to 1.2
mm long, glabrous externally, prominently ferruginous-hirsute
at the apex. Ovary ovoid, 1.2 mm long, ferruginous-hirsute in
the upper one-half, narrowed to the subsessile or sessile capitate
stigma. Fruit ovoid or ellipsoid, glabrous, fleshy, reddish when
mature, 2 cm long, the valves coriaceous. Seeds about 15,
obliquely obovoid, about 5 mm long, externally densely covered
with roundish, distinct, pale dots. Aril thin, submembrana-
ceous, 8 to 10 mm long, enveloping the seed, the upper part more
or less lacerate.

Luzon, Province of Rizal, San Isidro, Bur. Sci. 18525 Ramos (type),

August, 1910, in forests, Bur. Sci. 1851 Ramos, January, 1907, in fruit;
Oriud, Loher 6215.

328 The Philippine Journal of Science 1914

A species manifestly allied to Casearia fuliginosa Blanco, but readily ©
distinguishable by its fewer-nerved leaves. In C. fuliginosa Blanco the
nerves are usually about 10 on each side of the midrib, in C. loheri 5 or 6
on each side. The type was from Rizal Province, not from Laguna Province
as distributed.

CASEARIA SUBCORDATA sp. nov. § Pitumba.

Arbor parva partibus junioribus floribusque exceptis glabra;
foliis coriaceis, oblongis, nitidis, in siccitate brunneis, usque ad
15 cm longis, breviter acute acuminatis, basi abrupte subtrun-
cato-cordatis, leviter inaequalibus, margine serratis, nervis
utrinque circiter 12; floribus axillaribus, fasciculatis, ut videtur
numerosis, 5-meris, tenuiter pedicellatis; fructibus anguste
ovoideis vel ellipsoideo-ovoidis, 1 ad 1.4 cm longis, acutis, in
siccitate leviter 6-sulcatis.

A small tree, nearly glabrous except the younger parts.
Branches terete, lenticellate, grayish-brown or reddish-brown,
glabrous, the younger branchlets distinctly hirsute with short
hairs. Leaves oblong, coriaceous, 8 to 15 cm long, 3.5 to 5.5 em
wide, the base abruptly and somewhat obliquely subtruncate-
cordate, the sinus shallow, the basal lobes broadly rounded, the
apex shortly but rather sharply acuminate, the margins regularly .
and rather finely serrate throughout, brown when dry, the upper
surface strongly shining, the lower a little paler, shining, glabrous
or when young with few short hairs on the lower surface; lateral
nerves about 12 on each side of the midrib, prominent, some-
what curved, anastomosing, the reticulations fine, distinct;
petioles 5 to 8 mm long, the younger ones slightly pubescent.
Flowers 5-merous, axillary, fascicled, their pedicels slender, 3
to 6 mm long, slightly pubescent. Sepals 5, oblong-ovate, glab-
rous or nearly so, coriaceous, obtuse, 3 mm long slightly accres-
cent and persistent in fruit, somewhat hirsute on the median
portion inside. Stamens apparently 10; filaments 0.5 mm long;
anthers a little longer; staminodes shorter than the stamens,
densely ciliate-hirsute. Ovary narrowly ovoid, glabrous, nar-
rowed upward to the short style, the stigma capitate. Fruit
narrowly ovoid to ovoid-ellipsoid, 1 to 1.4 cm long, glabrous,
numerous, 2 to 6 or more in each axil, shallowly longitudinally
6-suleate when dry, acute; valves thickly coriaceous. Seeds
about 12 in each fruit, smooth, brown, somewhat inequilaterally
obovoid or oblong-obovoid, 4 mm long; aril fleshy, about 5 mm
long, quite inclosing the seeds, irregularly lacerate and divided.
Peduncles of the fruits about 1 cm long. ;

MINDANAO, Bukidnon Subprovince, Bur. Sci. 15698 Fénix, August, 1912.
A species manifestly very closely allied to Casearia polyantha Merr.,

1X, C, 4 Merrill: Philippine Plants, X 329

which it greatly resembles, but distinguishable by its leaves being truncate-
cordate at the base. Its leaf-form is somewhat similar to that of C. cinerea
Turcz., and C. grewiaefolia Vent., but these two species are at once distin-
guished by their pubescence. G. truncata BIl., is also apparently closely
allied.

ARALIACEAE
BOERLAGIODENDRON Harms

BOERLAGIODENDRON HETEROPHYLLUM sp. nov.

Arbor erecta, ramosa, circiter 5 m alta, glabra; foliis palma-
tim 3- ad 7-foliolatis, foliolis valde inaequalis, oblongo-obovatis
vel late oblongo-lanceolatis, acuminatis, inferioribus vix 12 cm
longis, superioribus usque ad 85 ecm longis, acuminatis, lobatis
dentatisque, basi sensim angustatis; umbellis densis, floribus sub-
capitato-dispositis, 4-meris.

An erect, branched tree about 5 m high, glabrous, the trunk
about 8 cm in diameter. Leaves chartaceous, palmately 3- to
7-foliolate, or sometimes some of the leaves very deeply 3-parted,
the lobes extending almost to the base; leaflets very unequal in
size, when 5 or 7 are present the lower two usually 12 cm long
or less, the upper ones larger, the middle one up to 35 cm in
length, oblong-obovate to broadly oblong-oblanceolate, acuminate,
usually lyrately lobed, dentate, the lobes broadly ovate, acumi-
nate, the base gradually narrowed, the petiolules of the middle
leaflets up to 3 cm in length, of the others gradually shorter;
petioles 10 to 25 cm long, the basal part with four or more
somewhat pectinate crests. Umbels terminal, about 30 termi-
nating each branch, forming a dense cauliflower-like inflores-
cence, the peduncles 10 to 15 mm long, each with two lanceolate,
8 to 10 mm long bracts at the apex subtending the two lateral
branches. Central head of each peduncle almost sessile, sub-
capitate, of numerous sterile flowers, their pedicels 3 to 4 mm
long, the ovaries ellipsoid, about 3 mm long. Lateral two um-
bels about 1 cm in diameter, dense, subcapitate, their peduncles
8 to 10 mm long. Flowers numerous, sessile or subsessile,
4-merous. Petals 2 mm long. Anthers about 1 mm long.

MINDANAO, Butuan Subprovince, Mount Hilong-Hilong, Weber 1126,
March 29, 1911, in forests, altitude about 550 m.

A species most closely allied to Boerlagiodendron clementis Merr., of
Mindanao, but at once distinguished by its leaves being palmately compound.

SCHEFFLERA Forster

SCHEFFLERA OBOVATA sp. nov. § Euschefflera, Heptapleurum.
Frutex epiphyticus, inflorescentiis exceptis glaber; foliis pal-
matim 5- ad 7-foliolatis, foliolis subcoriaceis, nitidis, obovatis,

why

330 The Philippine Journal of Science 1914 |

integris, basi angustatis, acutis, apice abrupte caudatis, nervis

primariis utrinque 5 vel 6, quam secondariis reticulisque haud
magis distinctis; inflorescentiis terminalibus, racemis circiter 8,
racemose dispositis, circiter 15 cm longis, fructibus 5-locellatis,

An epiphytic shrub, glabrous except the _ inflorescence.

ranches terete, grayish, striate when dry. Leaves alternate,
their petioles 6 to 7 cm long, inflated and clasping at the base,
the leaflets 5 to 7, palmately arranged; leaflets obovate, subcori-
aceous, 5 to 8 em long, 2 to 4 em wide, entire, narrowed gradually
to the acute or cuneate base, the apex abruptly and prominently
acuminate, the acumen stout, 5 to 8 mm long, acute, the upper
surface shining, the lower dull, of about the same color; lateral
nerves 5 or 6 on each side of the midrib, the basal ones sharply
ascending, distinct but not more prominent than are the second-
ary ones and the reticulations, all equally prominent on both
surfaces; petiolules 1.5 to 2.5 em long, those of the outer leaflets
somewhat shorter than of the middle ones. Inflorescence ter-
minal, sparingly furfuraceous-pubescent with short pale hairs,
the rachis short, about 2 cm long, stout. Racemes usually about
8, 12 to 15 em long, bearing numerous fascicled fruits, the pedi-
cels slender, 3 to 4 mm long, usually 2 or 3 fruits in each fascicle.
Fruits reddish-yellow when fresh, glabrous, ellipsoid, when dry
prominently longitudinally 5-sulcate, 5-celled, crowned by the 5,
round, sessile stigmas.

Luzon, Province of Laguna, Dahican, Phil. Pl. 1108 Ramos, September
Bie i912:

A species manifestly allied to Schefflera caudata Vid. (S. acwminatissima
Merr.) from which it differs especially in its quite differentially shaped,
much smaller leaves.

Additional material from the same province (Bur. Sci. 10001, 10973
Ramos), the specimens in flower, is very similar to the species above de-
scribed, but in both the flowers are in racemosely disposed umbels, while

in the type of S. obovata the flowers are merely fascicled, not at all
umbellate.

SCHEFFLERA DEMESAE sp. nov. § Euschefflera.

Frutex scandens, glaber; foliis alternis, palmatim 5-foliolatis,
foliolis chartaceis, usque ad 24 cm longis, integris, nitidis, apice
breviter acuminatis, basi rotundatis, nervis tenuibus, utrinque
usque ad 25, petiolulis valde inaequalibus; inflorescentiis termi-
nalibus, pedunculatis, floribus 5-meris, racemosis, racemis elon-
gatis, subumbellatim dispositis.

A scandent glabrous shrub, the stem reaching a diameter of
about 3 cm, purplish, the ultimate branches terete, about 5 mm
in diameter, striate when dry, grayish or brownish. Leaves

IX, C, 4 Merrill: Philippine Plants, X 831

palmately 5-foliolate, their petioles inflated at the base, about
5 em long. Leaflets oblong to elliptic-oblong, chartaceous, shin-
ing, 15 to 24 em long, 7.5 to 9.5 em wide, entire, the apex shortly
acuminate, the base rounded; lateral nerves slender, about 25
on each side of the midrib, the reticulations distinct; petiolule of
the central leaflet 7 em long, of the next two inner leaflets 2.5
to 3 cm long, of the outer leaflets about 1 cm long. Inflorescence
terminal, peduncled, the peduncle up to 15 cm long, usually with
two distant, ovate bracts, the lower one 5 mm long, the upper
one twice as long. Branches of the inflorescence subumbellately
disposed near the apex of the peduncles, usually 6, slender, about
30cm long, many-flowered. Flowers racemosely disposed, usually
fascicled at the nodes, few in each fascicle, pale-green, their pedi-
cels about 6 mm long. Calyx short, truncate, somewhat disk-like
Petals 5, oblong-ovate or ovate, acute, appendaged at the apex
inside, 3 mm long, 3-nerved. Filaments 2 mm long; anthers
broad, about 1.3 mm long. Fruit not seen.

MINDANAO, District of Zamboanga, Siay River, For. Bur. 18896 Fox-
worthy, DeMesa, & Villamil, May 29, 1912, on river banks, known to the
Moros as canonucan.

Apparently a very distinct species, characterized by being quite glabrous;
by its peduncled inflorescence, with its elongated subumbellately disposed

primary branches; and its 5-foliolate, entire, ample leaves with their very
unequal petiolules.

SCHEFFLERA CAUDATIFOLIA sp. nov. § Euschefflera.

Frutex scandens, glaber, ramis ramulisque teretibus; foliis
palmatim 5-foliolatis, foliolis subcoriaceis, nitidis, oblongis ad
oblongo-lanceolatis, usque ad 10 cm longis, utrinque angustatis,
basi acutis, apice longe tenuiter caudato-acuminatis; floribus
racemosis, 5-meris, racemis subterminalis, solitariis vel binis,
tenuibus, usque ad 13 cm longis.

A scandent shrub, quite glabrous, the branches and branchlets
terete, when dry light-gray, wrinkled. Leaves scattered, pal-
mately 5-foliolate, their petioles 4 to 8 cm long, inflated at the
base; leaflets oblong to oblong-lanceolate, 5 to 10 cm long, 2 to
3.4 em wide, subequally narrowed to the acute base and to the
slenderly caudate-acuminate apex, entire, the acumen straight
or falcate, up to 2 cm long, the upper surface subolivaceous,
shining, the lower shining, paler than the upper; lateral nerves
5 or 6 on each side of the midrib, rather distinct, anastomosing;
petiolules 1 to 2 cm long. Inflorescence terminal, of one or two
rather slender racemes, the racemes up to 13 cm long. Flowers
5-merous, solitary, in pairs, or somewhat fascicled, their pedicels

332 The Philippine Journal of Science 1914

about 4 mm long. Calyx small, less than 2 mm in diameter.
Petals 5, oblong-ovate, about 2.5 mm long, 1.5 mm wide, blunt,
inflexed-keeled at the apex inside. Anthers about 1.5 mm long.
Fruit 4 to 5 mm long, about 2.5 mm in diameter, oblong, longi-
tudinally 5-suleate, apparently somewhat fleshy when fresh,
brown when dry.

Luzon, Province of Camarines, Mount Isarog, Bur. Sci. 20062 Ramos,
November 8, 1913, on forested slopes.

Probably most closely allied to Schefflera obovata Merr., but with quite
different leaflets.

SCHEFFLERA CRASSIFOLIA sp. nov. § Euschefflera.

Frutex scandens, glaber; foliis palmatis, foliolis 6, oblongis,
coriaceis, nitidis, acuminatis, usque ad 18 cm longis, subtus palli-
dis, nervis reticulisque valde prominentibus; racemis 3 ad 6 in
ramulis junioribus plus minusve confertis, usque ad 35 cm longis;
floribus fasciculatis vel solitariis, pedicellatis, 5-meris; fructibus
ellipsoideis, longitudinaliter 5-sulcatis, 5 ad 6 mm longis.

A scandent shrub, quite glabrous, or the very young parts
sometimes slightly furfuraceous. Branches terete, lightly-gray,
striate or wrinkled when dry. Leaves palmately compound,
scattered, their petioles 5 to 10 cm long, inflated at the base,
clasping the stem. Leaflets 6, oblong, thickly coriaceous, 10 to
18 cm long, 3.5 to 7 cm long, entire, the base rounded to subacute,
apex rather abruptly and prominently acuminate, the acumen
straight or somewhat falcate blunt, up to 2 cm long; upper sur-
face olivaceous, prominently shining, the veins and reticulations
distinct, the lower surface pale, grayish-yellow, the veins and
reticulations very prominent, the lateral nerves about 10 on each
side of the midrib, spreading, anastomosing; petiolules about 3
em long. Inflorescence terminal, the racemes 3 to 6,. arranged
near the apices of the branchlets, the individual racemes up to
35 em long, dark-brown when dry. Flowers numerous, fascicled
and solitary along the racemes, their pedicels 3 to 4 mm long.
Fruits apparently somewhat fleshy, when dry brown, ellipsoid,
5 to 6 mm long, about 4.5 mm in diameter, longitudinally
5-celled.

Luzon, Province of Camarines, Sagnay, on trees in damp forests, Bur.
Sci. 22152 Ramos, December 15, 1913 (type): Province of Albay, Adumoy
hills, For. Bur. 12884 Curran, June, 1908, locally known as caranglang.

A species well characterized by its palmately 6-foliolate leaves, the leaflets
very thick and with prominent nerves and reticulations, and its elongated
racemes which are somewhat crowded at the apices of the branchlets. Its

alliance seems to be with Schefflera clementis Merr., but the flowers are
all in strict racemes, not in racemosely arranged umbels.

ee, 4 Merrill: Philippine Plants, X 333
EBENACEAE

MABA Forster
MABA EUPHLEBIA sp. nov.

Frutex vel arbor parva (ex Ramos 3 m alta) ; foliis alternis,
breviter petiolatis, oblongis, usque ad 33 cm longis, acuminatis,
basi rotundatis, supra glabris subtus parce pubescentibus; nervis
utrinque circiter 12, distantibus, valde prominentibus; floribus
paucis, sessilibus, bracteolatis, fasciculatis, axillaribus; stamini-
bus 12; corolla circiter 1.5 cm longa, extus dense fulvo-hirsuta.

A shrub or small tree (8 m high ex Ramos), the branchlets and
lower surfaces of the leaves somewhat pubescent, the flowers
densely so, otherwise glabrous. Branches slender, terete, brown
or grayish, Leaves subcoriaceous or thickly chartaceous, oblong,
20 to 33 cm long, 7 to 9 cm wide, somewhat shining when dry,
the upper surface olivaceous, glabrous, the lower pale-brownish,
distinctly pubescent on the midrib and nerves, the base somewhat
narrowed, rounded, the apex acute or acuminate; lateral nerves
about 12 on each side of the midrib, very prominent, curved,
distinct, anastomosing, the reticulations lax, prominent; petioles
stout, pubescent, about 3 mm long. Flowers few, 2 or 3, rarely
more, in axillary, sessile fascicles, each flower subtended by three
ovate to ovate-lanceolate, acuminate, hirsute, about 3 mm long
bracteoles. Calyx about 5 mm long, slightly hirsute, cup-shaped,
3-lobed, the lobes broadly ovate, acute or acuminate, one about
as long as the calyx-tube. Corolla in nearly mature bud about
1.5 cm long, cylindric, 3 mm in diameter below, narrowed above,
outside very densely hirsute-pubescent with appressed fulvous
hairs. Stamens 12, in three series, the anthers linear-oblong,
2 to 3 mm long, the filaments of the inner series about 2 mm
long, of the outer series about 6 mm long, flattened. Female
flowers and fruits not seen.

Luzon, Province of Tayabas, Tagcauayan, in forests, altitude about
90 meters, Bur. Sci. 18885 Ramos, March 15, 1911.

A very characteristic species, distinguishable by its comparatively large,
prominently nerved leaves which are pubescent beneath, and its few, axil-
lary, fascicled, sessile flowers. It is, perhaps, most closely allied to Maba
venosa King & Gamble, of Singapore, but is entirely distinct from that

species as described.
DIOSPYROS Linnaeus

DIOSPYROS TRIFLORA sp. nov. —

Arbor parva, floribus exceptis glabra, ramis ramulisque tenu-
ibus, teretibus; foliis oblongis, usque ad 23 cm longis, subcoria-
ceis, supra nitidis, acuminatis, basi acutis, leviter decurrento-

834 The Philippine Journal of Science 1914

acuminatis, vel subrotundatis, eglandulosis, nervis utrinque
circiter 9, tenuibus, haud prominentibus; inflorescentiis 4
axillaribus, solitariis, pedunculatis, circiter 2 cm longis, 3-floris;
floribus 5-meris, calycibus cylindraceis, 6 mm longis, leviter
5-dentatis; corolla extus dense pubescens; staminibus 10.

A small tree, glabrous except the flowers. Branches and
branchlets slender, terete, smooth, the latter black, the former
black when dry. Leaves oblong, subcoriaceous, 12 to 23 cm long,
4 to 9 em wide, the apex distinctly acuminate, the acumen blunt,
the base acute, somewhat decurrent-acuminate, or somewhat
rounded, eglandular, the upper surface very smooth and strongly
shining when dry, the lower surface of nearly the same color
but duller; lateral nerves 8 to 10 on each side of the midrib,
slender, not prominent, loosely anastomosing, the reticulations
not prominent, lax; petioles about 8 mm long. Male inflores-
cences axillary, solitary, about 2 cm long, the peduncles 5 to 10
mm long, each bearing three, subumbellately arranged, pedicelled
flowers, ebracteolate. Male flowers 5-merous, their pedicels
pubescent, 2 to 3 mm long, the calyx, in bud, cylindric, black
when dry, slightly pubescent, about 6 mm long, 2 to 2.5 mm in
diameter, the base obtuse, somewhat narrowed, the apex trun-
cate and slightly 5-toothed, the teeth acute, 0.5 mm long or less.
Corolla (in young bud) narrower than the calyx, externally
densely gray-pubescent, the tube apparently short, the lobes
elongated, imbricate. Stamens 10, in two series, the anthers
lanceolate, acuminate, up to 3 mm long (immature). Female
flowers and fruit not seen.

BALABAC, Cape Melville, Bur. Sci. 15652 Fénix, July 18, 1912, along trails
at low altitudes.

A species well characterized by its 3-flowered, peduncled, solitary inflores-
cences, and its cylindric and truncate, obscurely 5-toothed calyces. It is

apparently allied to the Malayan Diospyros truncata Zoll. & Mor., but has
a peduncled, not fasciculate inflorescence, and quite differently shaped leaves.

DIOSPYROS FASCICULIFLORA sp. nov.

Arbor parva subtus foliis junioribus ramulis floribusque ex-
ceptis glabra, ramis ramulisque teretibus; foliis chartaceis vel
subcoriaceis, oblongis vel oblongo-lanceolatis, usque ad 20 cm
longis, utrinque angustatis, acuminatis; nervis lateralibus cir-
citer 12, subtus valde prominentibus, arcuato-anastamosantibus,
reticulis laxis, distinctis; floribus 4-meris, ferrugineo-pubescen-
tibus, solitariis vel fasciculatis in axillis defoliatis vel in ramis
infra foliis, calycibus post anthesis brevibus, rotatis, circiter 12
mm diametro; ovario 4-loculare, dense hirsuto; fructibus 4-locel-

IX, ©, 4 Merrill: Philippine Plants, X 335

latis, globosis vel globoso-obovoideis, circiter 2 cm diametro,
seminibus 4, albumine vix ruminato.

A small tree, glabrous except for the flowers, the younger
branchlets, and the lower surfaces of young leaves. Branches
and branchlets terete, slender, dark-gray or brownish-black
when dry, wrinkled, lenticellate, the younger branchlets somewhat
ferruginous-pubescent. Leaves oblong, chartaceous or subcor-
iaceous, 12 to 20 cm long, 3.5 to 8 cm wide, narrowed at both
ends, the apex rather sharply acuminate, the base acute, eglandu-
lar, the upper surface smooth and shining when dry, brownish
or olivaceous, the lower surface usually much paler, slightly
shining, when young rather softly pubescent or puberulent,
often becoming glabrous or nearly so; lateral nerves about 12
on each side of the midrib, very prominent, arched-anastomosing,
the reticulations lax, prominent; petioles up to 1 cm in length.
Female flowers solitary or fascicled in the axils of fallen leaves,
or fascicled on rather prominent protuberances on the branchlets
below the leaves, 4-merous, sessile or very shortly and stoutly
pedicelled. Calyx densely ferruginous-pubescent, the tube very
short, broad, the lobes immediately after flowering orbicular-
reniform, rounded, densely ferruginous-pubescent on both sur-
faces, about 3 mm in diameter, soon accrescent, spreading,
reniform, 5 mm long and 8 mm in diameter, often recurved,
thickly coriaceous. Corolla and stamens not seen. Ovary ovoid,
densely ferruginous-pubescent, 4-celled. Fruit apparently fleshy,
globose or obovoid-globose, when dry about 2 cm in diameter and
often grayish, the pericarp glabrous, wrinkled, the coriaceous
calyx-lobes persistent at the base of the fruit. Seeds 4, one in
each cell, 12 to 15 mm long, about 1 cm wide and thick, the
albumen hard but scarcely bony, smooth, whitish or yellowish
when dry, not at all ruminate.

BASILAN, Bur. Sci. 16101 Reillo, August, 1912 (type).

I refer also to this species the following material: LUZON, without definite
locality, Loher 6562, in fruit: Province of Tayabas, Bur. Sci. 10283 Curran,
in fruit. CreBu, Asturias, For. Bur. 6450 Everett, in fruit. MINDANAO,

District of Zamboanga, San Ramon, Hallier, in fruit. The only recorded
native name is the Cebuano panangtilong.

DIOSPYROS MIRANDAE sp. nov.

Arbor circiter 30 m alta inflorescentiis exceptis glabra; foliis
crasse coriaceis, oblongis, nitidis, utrinque concoloribus, usque
ad 12 cm longis, acuminatis, nervis utrinque circiter 10, tenuibus,
vix prominentibus, reticulis densis; floribus 9 5-meris, axilla-
ribus, in racemis brevibus paucifloris dispositis; calycibus in

336 The Philippine Journal of Science 1914

alabastra obovoideis, 6 mm diametro, crassissime coriaceis,
longitudinaliter plicatis, lobis reniformibus; ovario 10-locellato.

A dioecious tree about 30 m high, glabrous except the in-
florescence. Branches terete, grayish, lenticellate. Leaves
alternate, thickly coriaceous, oblong, 8 to 12 cm long, 3 to 5
cm wide, shining and of about the same color on both surfaces,
base rounded or subacute, apex shortly acuminate; lateral nerves
about 10 on each side of the midrib, slender, not prominent,
anastomosing, the reticulations slender, dense; petioles about 5
mm long. Female flowers in short, axillary, solitary, few-
flowered, pubescent, spike-like racemes, the whole about 2 cm
long. Flowers 5 or less in each raceme, their pedicels very stout,
jointed, pubescent, about 2 mm long. Calyx in bud, just before
anthesis, obovoid, pubescent, about 6 mm long and wide, with
5 reniform, rounded lobes, very thickly coriaceous, 3 mm long
and 6 mm wide, the lobed portion longitudinally plicate or
folded-plicate, star-shaped in cross-section, pubescent inside.
Corolla-tube 6 mm long, 2.5 mm wide, cylindric, basal part gla-
brous, pubescent above, the lobes broadly ovate, spreading in
anthesis, 5, pubescent on both surfaces, about 5 mm long, acute.
Ovary ovoid, densely appressed-pubescent, narrowed upward
into the thick style, ovary and style about 5 mm long, 10-celled.
Staminodes 5, inserted at the base of the corolla, the sterile
anthers linear, about 2 mm long.

MINDANAO, District of Cotabato, southeastern ridge of Mount Glan, For.
Bur. 18752 Miranda (type), For. Bur. 14248 Tarrosa, June 6 and May 25,
1912, in forests, altitude 90 to 125 meters, locally known to the Moros as
bantulinay.

A species well characterized by its peculiar calyx which is somewhat

of the type of that of Diospyros affinis Thw. The flowers are said to be
white and fragrant, and the heart-wood is described as black.

DIOSPYROS PLICATA sp. nov.

Arbor dioica, 8 ad 25 m alta, partibus junioribus inflorescent-
iisque exceptis glabra; foliis junioribus pubescentibus, vetus-
tioribus glabris, oblongis, usque ad 18 cm longis, coriaceis, nitidis,
subtus pallidioribus, acuminatis, nervis utrinque circiter 12,
distantibus, patulis, distinctis, anastomosantibus; floribus @
axillaribus, fasciculatis, dense fulvo-pubescentibus, 5-meris,
sepalis in anthesis breviter connatis, longitudinaliter plicatis et
horizontaliter undulato-plicatis, accrescentibus et 2.5 cm longis.

A tree 8 to 25m high. Branches terete, slender, nearly black
when dry, the growing parts, the young leaves, and the flowers
appressed-pubescent. Leaves oblong, coriaceous, 12 to 18 cm

IX, C, 4 Merrill: Philippine Plants, X 837

long, 2.5 to 8 cm wide, shining on both surfaces, the lower
somewhat paler than the upper, the apex blunt-acuminate, the
base acute or rounded; nerves about 12 on each side of the midrib,
spreading, distant, prominent on the lower surface, looped-
anastomosing, the reticulations very lax, distinct; petioles 3 to
8 mm long. Female flowers 5-merous, axillary, in few-flowered
fascicles, densely fulvous-pubescent. Calyx in bud densely
pubescent, the sepals oblong-lanceolate, longitudinally folded or
plicate, the reflexed margins very prominently horizontally un-
dulate-plicate, free nearly to the base. Corolla cylindric, 6 mm
long in bud, densely pubescent. Ovary densely appressed-hir-
sute, narrowly ovoid, 5- or 6-celled. Young fruit narrowly
ovoid, obtuse, pubescent, about 1 cm long (very young) quite
free from the accrescent, involucre-like calyx. Accrescent
calyx-lobes ovate, somewhat foliaceous, coriaceous, slightly
pubescent, 2.5 cm long, free except at the very base, folded back
longitudinally and also prominently horizontally undulate-
plaited, forming an ovoid involucre surrounding but free from
the fruit, this accrescent calyx 5 cm in diameter when spread.

MINDANAO, District of Zamboanga, near Margosatubig, For. Bur. 18281
Foxworthy, DeMesa, & Villamil, May 11, 1912 (type), in hill forests, altitude
about 120 meters, specimens with young fruit: District of Cotabato, Cabla-
can, For. Bur. 14901 Tarrosa, May, 26, 1912, near the seashore; Buluan Point,
For. Bur. 18284 Miranda, May 27, 1912, the last two with young flowers.

A very striking species, well characterized by its accrescent calyx which
is quite free from the fruit, and which is longitudinally folded or plicate
and at the same time very prominently horizontally undulate-plicate. Its

leaves are very similar to those of our common Diospyros pilosanthera, to
which, however, it is not otherwise at all closely allied.

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s rae’ WARS Ps

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THE PHILIPPINE

JOURNAL OF SCIENCE

C. BoTANY
Vou. X JANUARY, 1915 No. 1

NEW OR NOTEWORTHY PHILIPPINE PLANTS, XI

By E. D. MERRILL *

(From the Botanical Section of the Biological Laboratory, Bureau of
Science, Manila, P. I.)

The eleventh paper of this series is essentially like its pred-
ecessors.” It consists of the descriptions of 90 new species of
Philippine plants in the families Chloranthaceae, Aristolochia-
ceae, Saxifragaceae, Cunoniaceae, Leguminosae, Simarubaceae,
Burseraceae, Anacardiaceae, Sabiaceae, Rhamnaceae, Elaeo-
carpaceae, Begoniaceae, Ericaceae, Sapotaceae, Apocynaceae,
Verbenaceae, and Gesnertaceae. About 12 previously described
species are for the first time credited to the Archipelago, making
the total number of additions to the Philippine flora, recorded
in this paper, approximately 100. Genera recorded from the
Archipelago for the first time are Aquilaria, Koompassia, Meli-
lotus, Neptunia, Cymodocea, Diplanthera, Hanguana, Urceola,
Vallaris, and Protium. A few reductions are made and a few
changes in nomenclature are recorded as such changes have
been indicated by the rules of the international code of botanical
nomenclature.

* Associate professor of botany, University of the Philippines.

*New or Noteworthy Philippine Plants. Govt. Lab. (Philip.) Publ. 6
(1904) 5-18; II, 1}. c. 17 (1904) 5-47; ITI, 1. c. 29 (1905) 5-50; IV, }. c.
35 (1905) 5-68; V, Philip. Journ. Sci. 1 (1906) Suppl. 169-246; VI, 1. c.
3 (1908) Bot. 219-267; VII, 1. c. 4 (1909) 247-330; VIII, 1. c. 5 (1910)
167-257; IX, 1. c. 7 (1912) 259-357; X, 1. ec. 9 (1914) 261-337.

181281

2 The Philippine Journal of Science 1915

HYDROCHARITACEAE
THALASSIA Solander

THALASSIA HEMPRICHII (Ehrenb.) Aschers. in Engl. & Prantl Nat.
Pflanzenfam. 2* (1889) 254.

Schizotheca hemprichii Ehrenb. in Abh. Akad. Berlin 1 (1832) 429.

CAVILLI ISLAND, Sulu Sea, Merrill 7180, September, 1910, on coral sand,
submerged at low tide; sterile specimens, det. C. H. Ostenfeld. Luzon,
Manila Bay, Merrill, August, 1911, washed up on Pasay beach.

Not previously recorded from the Philippines; widely distributed along
the tropical shores of the Indian and Pacific Oceans.

HALOPHILA Thouars

HALOPHILA OVALIS (R. Br.) Hook. f. Fl. Tasm. 2 (1860) 45.

Caulinia ovalis R. Br. Prodr. (1810) 339.

BANCORAN ISLAND, Sulu Sea, Merrill 7155, September, 1910, sterile
specimens growing on coral sand in shallow water, 0.4 to 1.8 m deep at
low tide, det. C. H. Ostenfeld.

This species has previously been recorded from the Philippines by
Naves,’ but the record has not before been verified. It is to be noted that
Halophila ovalis Hook. f. is quite distinct from H. ovata Gaudich. For
a consideration of the latter see Ostenfeld’s paper.‘

Widely distributed along tropical shores of the Indian and Pacific
Oceans.

HALOPHILA SPINULOSA (R. Br.) Aschers. in Neumayer Anleit. Wis-
sensch. Beob. 368, ed. 3 (1905) 396; Benth. Fl. Austral. 7 (1878)
183.
Caulinia spinulosa R. Br. Prodr. (1810) 339.
LUZON, Province of Camarines, For. Bur. 18875 Curran, June, 1908,
sterile specimens thrown up by the waves, det. C. H. Ostenfeld.
This species has previously been reported from Mindanao by Aschegrson,
l. c. ed. 38, as indicated to my by Dr. Ostenfeld.
Eastern and northern coasts of Australia, the Philippines, and Java.

POTAMOGETONACEAE
CYMODOCEA Koenig

CYMODOCEA ROTUNDATA (Ehrb. & Hempr.) Asch. & Schweinf. in
Sitzber. Ges. Freunde Berlin (1870) 84; Graebner in Engl. Pflan-
zenreich 31 (1907) 147.

Phucagrostis rotundata Ehrb. & Hempr. Symb. Phys. Bot. t. 17.

CAVILLI ISLAND, Sulu Sea, Merrill 7180A, September, 1910, sterile spec-
imens mixed with Thalassia hemprichii Aschers., det. C. H. Ostenfeld.

This species has already been reported from Mindanao by Graebner,

l. c. It extends from the shores of the Red Sea southward to Madagascar

and eastward to Mindanao, Timor, and Australia.

*Novis. App. (1880) 298.
‘Philip. Journ. Sci. 4 (1910) Bot. 68.

x,c,.1 Merrill: New or Noteworthy Philippine Plants, XI 8

DIPLANTHERA Thouars

DIPLANTHERA UNINERVIS (Forst.) Aschers. in Engl. & Prantl Nat.
Pflanzenfam. 2* (1897) 37; Graebner in Engl. Pflenzenreich 31 (1907)
152.

Zostera uninervis Forsk. Fl. Aeg.-Arab. (1775) 159.

CAVILLI ISLAND, Sulu Sea, Merrill 7179, September, 1910, on coral sand
in shallow water, usually exposed at low tide; sterile specimens det. C. H.
Ostenfeld. Luzon, Manila Bay, on the Bataan coast, Shaw.

Not previously reported from the Philippines; tropical shores from the
Red Sea and the east coast of Africa to Malaya, Australia, and Polynesia.

FLAGELLARIACEAE
HANGUANA Blume (Susum Blume)

HANGUANA MALAYANA (Jack) comb. nov.

Veratrum ? malayanum Jack in Malay Miscel. 1 (1820) 25; Hook. Bot.
Miscel. 2 (1831) 74.

Hanguana kassintu Blume Enum. Pl. Jav. (1827) 15, ed. 2 (1830) 15;
Mig. Fl. Ind. Bat. 3 (1857) 248.

Susum malayanum Planch ex Hook. f. Fl. Brit. Ind. 6 (1892) 391; Ridl.
Mater. Fl. Malay. Penin. 2 (1907) 1382.

Veratonia malayana Miq. Fl. Ind. Bat. 3 (1857) 553.

PALAWAN, Malampaya Bay, Merrill 7221, September, 1910. MINDANAO,
Province of Surigao, Hinatuan, Piper 529, May, 1911.

There seems to be very little doubt as to the identity of Veratrum sic
yanum Jack and Hanguana kassintu Blume, and as Blume’s generic name
Hanguana has three years priority over his generic name Susum, it is
here adopted. The case is not covered by the lists of nomina conservanda
adopted by the Vienna or the Brussels botanical congress.

The genus has previously not been reported from the Philippines.
Malay Peninsula and Archipelago.

CHLORANTHACEAE
CHLORANTHUS Linnaeus

CHLORANTHUS VERTICILLATUS sp. nov.

Herba erecta, ramulis subtus foliisque ad nervos plus minusve
furfuraceo-pubescentibus; foliis membranaceis, breviter petio-
latis, verticillatis, ovatis ad elliptico-ovatis vel elliptico-obovatis,
acuminatis, usque ad 10 cm longis, utrinque angustatis, margine
distincte acute apiculato-serrulatis; spicis 1 vel 2, 2 ad 3 cm
longis, pedunculo elongato; squamis antheriferis 3-partitis, lobis
inaequalibus, intermedia 4 mm longa.

An erect, apparently branched herb, the younger branchlets,
the lower surfaces of the leaves on the veins and reticulations,
and to a lesser degree the inflorescence more or less furfuraceous-
pubescent. Leaves verticillate at the apices of the branchlets,
in whorls of 4, membranaceous, ovate to elliptic-ovate or elliptic-

4 The Philippine Journal of Science 1915"

obovate, about equally narrowed to the slenderly acuminate apex
and to the acute base, 8 to 10 cm long, 3.5 to 5 em wide, the
margins prominently and acutely apiculate-serrate except in the
lower 1 cm and the margins of the acumen which are entire;
nerves 7 or 8 on each side of the midrib, distinct, anastomosing,
the reticulations lax; petioles furfuraceous-pubescent, about 2
mm long. Inflorescence terminal, slightly pubescent, solitary,
including the peduncle up to 13 cm in length, each bearing one
or two, rather many fiowered, 2 to 3 cm long spikes. Flowers
white, the bract orbicular-ovate, irregularly toothed, 1.7 to 2
mm long, rounded. Pistillate flower: Ovary narrowly ovoid,
somewhat narrowed at both ends, about 2 mm long, glabrous.
Staminate flowers: Antheriferous scale 3-partite, the lobes ob-
long, rounded, the central one about 4 mm long, 1.2 mm wide,
bearing an oblong, 1.2 mm long, 2-celled anther, the lateral lobes
about as wide, one-half as long, each bearing a 1-celled anther.

LuzON, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19757 McGregor,
February, 1913.

Probably as closely allied to Chloranthus henryi Hemsl. as to any other
species. It is distinguished, however, by its pubescent branchlets and
lower surfaces of its smaller leaves, its inflorescence reduced to 1 or 2
spikes, and its smaller flowers. From its closest Philippine ally, C. philip-
pinensis Merr., it differs in its much smaller, pubescent, shortly petioled
leaves, and its larger flowers.

ARISTOLOCHIACEAE
ARISTOLOCHIA Linnaeus

ARISTOLOCHIA LEYTENSIS sp. nov.

Species A. tagala et A. mindanaensis similis et affinis, differt
Horibus multo majoribus, usque ad 8.5 cm longis.

A slender vine, nearly glabrous, the branches brownish when
dry, glabrous, deeply sulcate. Leaves ovate, membranaceous,
glabrous, of about the same color on both surfaces, somewhat
shining, 14 to 17 cm long, 8 to 10 cm wide, entire, the apex
acuminate, the base deeply and prominently cordate, the sinus
about 2 cm deep, 1.5 to 2 em wide, often obovate, the basal lobes
rounded; basal nerves 3, the lateral pair soon forked, and
its lower branches soon forked again, the lateral nerves above
the base about 3 on each side of the midrib, prominent, the re-
ticulations very lax; petioles 4 to 7 cm long, glabrous. Racemes
axillary, solitary, the rachis slender, slightly pubescent, about
3 cm long, each raceme bearing about 6 flowers, not all de-
veloping at one time, each flower subtended by an ovate-lanceo-
late, acuminate, slightly pubescent bract, the pedicels minutely

x%c1 0 ©Merrill: New or Noteworthy Philippine Plants, XI 5

pubescent, 7 to 10 mm long. Ovary somewhat clavate, about
8 mm long, the tube constricted for 4 mm above the ovary, then
inflated, the larger portion of the tube about 3 cm long, swollen
at the base, then constricted, and again enlarged and somewhat
funnel-shaped, the lip narrowly lanceolate, glabrous, gradually
narrowed upward to the slenderly acuminate apex, 5 to 6 cm
long, about 5 mm wide.

LEYTE, near Dagami, Bur. Sci. 15201 Ramos, August, 1912, in thickets
along streams.

A species with much the same vegetative characters as Aristolochia
tagala Cham., and A. mindanaensis Warb., the leaves being very similar
in all three species. Aristolochia leytensis is, however, entirely distinct
from both the above in its much larger flowers.

SAXIFRAGACEAE
CURRANIODENDRON Merrill

CURRANIODENDRON APOENSE (Elm.) comb. nov.
Dedea apoensis Elm. Leafl. Philip. Bot. 3 (1910) 682 (August 8).
Curraniodendron dedeaeoides Merr. in Philip. Journ. Sci. 5 (1910) Bot.
177 (August 19).

This characteristic species was approximately simultaneously described
by Mr. Elmer and by myself, but Mr. Elmer’s publication of the species
under Dedea antedates my own publication of the new genus and species
Curraniodendron dedeaeoides by about ten days. I have reéxamined the
material and am still of the opinion that the plant is worthy of being
considered the type of a distinct genus, although as previously indicated,
very similar to, and manifestly closely allied to the New Caledonian genus
Dedea Baill. The question for the future monographer to decide is whether
or not the resinous, not lepidote vegetative parts, the difference in number
in floral parts, and the very decided difference in the number of ovules
constitute sufficiently valid generic differences.

The species is represented by the following material: NrEGrRos, Mount
Marapara, For. Bur. 18684 Curran & Foxworthy. LEYTE, Dagami, Bur.
Sci. 15288 Ramos. MINDANAO, Mount Apo, Elmer 11524, 10627.

POLYOSMA Blume

POLYOSMA LAGUNENSIS sp. nov.

Species P. philippinensis affinis, differt foliis subtus ad nervos
adpresse hirsutis, nervis magis numerosis, floribus minoribus.

A small tree 5 to 8 m high, the older branches terete, light-
gray or pale-brownish, the younger branchlets and inflorescences
rather densely pubescent with pale-fulvous, short, appressed
hairs. Leaves opposite, mostly oblong to obovate-oblong, char-
taceous, 8 to 14 cm long, 3 to 6 cm wide, narrowed above to
the abruptly, slenderly, and sharply acuminate apex, the acumen
somewhat apiculate, and below to the cuneate base, entire or

6 The Philippine Journal of Science 1915"

with minute, scattered teeth opposite the ends of some of the
nerves, the upper surface, when dry, dark brownish-olivaceous,
slightly shining, smooth, glabrous, the lower surface paler, ap-
pressed hirsute with fulvous hairs on the midrib and lateral
nerves; lateral nerves 12 to 15 on each side of the midrib, prom-
inent on the lower surface, anastomosing, the reticulations
lax; petioles fulvous-pubescent, 2 to 2.5 cm long, in age be-
coming nearly glabrous. Racemes terminal, many flowered, soli-
tary, 10 to 14 em long, densely hirsute with short, fulvous hairs.
Flowers white, somewhat crowded, their pedicels 3 mm long,
the bracteoles usually 3, lanceolate, acuminate, 2 to 2.3 mm long.
Calyx hirsute, 2 to 2.5 mm long, the lobes lanceolate-ovate,
acuminate, 0.8 mm long. Petals 4, linear-lanceolate, 8 to 9 mm
long, about 2 mm wide at the base, narrowed upward to the
acute apex, prominently 5-nerved, the nerves dark-colored, out-
side hirsute. Filaments flattened, hirsute inside, about 4 mm
long; anthers linear, narrower than the filaments, 3 mm long.
Style glabrous, 8 mm long. Fruit subglobose, not rugose,
sparingly pubescent, 7 to 8 mm in diameter.

LuZON, Province of Laguna, near San Antonio and Paete, Bur. Set.
2893 (type), 14960, 10056 Ramos, For. Bur. 13185 Curran, flowering in
February and March, fruiting June and July.

A species manifestly closely allied to Polyosma philippinensis Merr.,
from which it differs in its smaller flowers, and its more numerously
nerved leaves which are appressed-pubescent on the midrib and nerves on
the lower surface, not glabrous. Another close ally appears to be the.

recently described Polyosma gitingensis Elm., which, however, has gla-
brous leaves, even smaller flowers, and much larger bracteoles.

POLYOSMA LINEARIBRACTEA sp. nov.

Arbor, omnibus partibus plus minusve pubescentibus vel hir-
sutis; foliis crasse coriaceis, oppositis, oblongis, usque ad 17 cm
longis, utrinque acutis, apice mucronatis, margine grosse dis-
tanter serratis; nervis utrinque circiter 12, subtus valde prom-
inentibus; racemis terminalibus, solitariis, circiter 15 cm longis;
fioribus circiter 2.4 cm longis, bracteolis linearis, elongatis, 8
ad 12 mm longis.

A tree said by the collector to be 20 m high, the branches
terete, dark-brown, glabrous, the younger branchlets, the leaves,
and the inflorescence more or less densely hirsute with grayish-
brown hairs. Leaves thickly coriaceous, oblong, 12 to 17 cm
long, 4 to 5 cm wide, narrowed about equally to the acute base and
apex, the apex also mucronate, the upper surface, when young,
appressed-hirsute, in age becoming glabrous, shining, greenish-
olivaceous, the lower surface paler, rather prominently hirsute

-x,c1 Merrill: New or Noteworthy Philippine Plants, XI 7

on the midrib and nerves, becoming nearly glabrous, the margins
rather coarsely and distantly apiculate or glandular-serrate, the
teeth corresponding to the lateral nerves; lateral nerves about 12
on each side of the midrib, curved, anastomosing, very prom-
inent on the lower surface, the reticulations lax, prominent;
petioles 2 to 3 cm long, hirsute, becoming nearly glabrous. Ra-
cemes terminal, solitary, about 15 cm long, densely hirsute, the
rachis rather stout. Flowers pale-lavender, about 30 in each
raceme, their densely hirsute pedicels about 4 mm long, the
bracteoles 3 at the apices of the pedicels, linear, hirsute, the
two lateral ones 8 to 9 mm long, 0.5 mm wide or less, the middle
one up to 12 mm long and a little wider than the lateral ones.
Calyx somewhat hirsute, including the lanceolate, acuminate
calyx-teeth about 6 mm long. Petals 4, about 2.5 cm long,
2.2 mm wide, scarcely narrowed upward, with about 4, dark-
colored nerves, hirsute, the apex thickened. Filaments slender,
scarcely flattened, 8 to 9 mm long, sparingly pubescent; anthers
linear, about as wide as the filament, obtuse, 6 to 7 mm long.
Style about 1.7 cm long, slender.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19656 McGregor,
February, 1913.

A most striking species, at once distinguished from all known Philip-

pine forms by its large flowers and very prominent, linear, elongated
bracteoles.

CUNONIACEAE
WEINMANNIA Linnaeus

WEINMANNIA LUCIDA sp. nov. § Leiospermum.

Arbor circiter 10 m alta inflorescentiis exceptis glabra; foliis
3- vel 5-foliolatis, usque ad 15 cm longis, foliolis subcoriaceis,
lanceolatis vel oblongo-lanceolatis, in siccitate brunneis, utrinque
valde nitidis, longe acuminatis, basi acutis, margine distanter
crenato-serratis, reticulis laxis, haud prominentibus; racemis
multifloris, folia subaequantibus, pubescentibus; floribus 4-meris,
sepalis caducis. —

A tree about 10 m high, glabrous except the inflorescence.
Branches grayish, terete, the younger parts brownish. Leaves
opposite, 9 to 15 cm long, the rachis and petiole quite glabrous;
leaflets 3 or 5, lanceolate or oblong-lanceolate, subcoriaceous,
strongly shining, brownish and of the same color on both sur-
faces when dry, 5 to 10 cm long, 1.5 to 4 cm wide, the apex
slenderly acuminate, the acumen blunt, the base acute, the
margins slightly and distantly crenate-serrate; lateral nerves
about 10 on each side of the midrib, slender, not prominent,

g The Philippine Journal of Science 1915

the reticulations lax, slender, rather obscure; petiolules of the
lateral leaflets 2 to 3 mm long, that of the terminal one much
longer. Inflorescence in the upper axils and subterminal, uni-
formly pubescent with short, rather closely arranged hairs, the
common peduncles thick, flattened, about 5 mm long, each bear-
ing at its apex 4 elongated racemes arranged in one plane;
racemes up to 10 cm long, many-flowered. Flowers 4-merous,
their pedicels slender, pubescent, 2.5 to 3 mm long, solitary
or somewhat fascicled along the rachis. Sepals oblong, obtuse
or subacute, 0.8 mm long, deciduous, leaving in fruit a disk-
like base nearly 1 mm in diameter and bearing 8 marginal
glands. Petals elliptic, rounded, about 1.2 mm long. Stamens
8; filaments 1 to 1.2 mm long; anthers heart-shaped, 0.8 mm
long. Capsules oblong, narrowed at both ends, the base acuté,
apex acuminate, 4 to 5 mm long including the two styles, 1 to
2 mm in diameter, uniformly pubescent with scattered, short
hairs, the styles 1 to 1.5 mm long.

Luzon, Province of Laguna, Dahican, in forests along the river, Phil.
Pl. 1109 Ramos, September 17, 1912.

A species manifestly allied to Weinmannia luzoniensis Vid., from which
it differs in its entirely glabrous leaves, but more especially in its much
narrower, differently shaped leaflets which are only slightly toothed,
strongly shining on both surfaces, and with slender, laxly arranged veins
and reticulations. Several of the Philippine representatives of this genus
find their closest allies (§ Leiospermum) in Polynesia, not in the Malayan
region.

LEGUMINOSAE
ALBIZZIA Durazzini

ALBIZZIA MYRIANTHA sp. nov.

Arbor circiter 20 m alta, inflorescentiis exceptis glabra; foliis
bipinnatis, circiter 35 cm longis, petiolo basi supra 1-guanduloso,
rhachibus glandulis 2 vel 3 instructis; pinnis circiter 10 utrinque,
10 ad 14 em longis; foliolis circiter 30 utrinque, oblongis, equi-
lateralibus, basi obtusis, apice rotundatis, circiter 1.5 cm longis,
costa centralis; inflorescentiis axillaribus folia aequantibus,
racemoso-paniculatis; floribus numerosis, albido-viridis, pedicel-
latis, ad apices ramulorum racemoso-capitatis.

A tree about 20 m high, glabrous except the inflorescence.
Branches subterete, very dark-colored when dry, slightly lenti-
cellate. Leaves bipinnate, about 35 cm long, the petiole with
a large gland near the base, the rachis with 2 or 3 smaller
glands in the upper part; pinnae about 10 on each side of the
rachis, opposite, the secondary rachises with glands in the upper
one-half, 10 to 14 cm long; leaflets opposite, about 30 pairs on

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI e)

-each pinna, oblong, chartaceous, 1.4 to 1.8 cm long, 5 to 7 mm
wide, equilateral, the midrib central, apex rounded, base obtuse
or rounded the lateral nerves about 6 on each side of the midrib,
indistinct, the upper surface olivaceous, shining when dry, the
lower paler. Panicles in the upper axils, usually one in each
axil, as long as the leaves, narrow, the whole forming a ter-
minal leafy inflorescence, sparingly fulvous-pubescent; primary
branches confined to the upper 10 cm, spreading, solitary or
fascicled, scattered below, rather crowded toward the apices,
5 em long or less, each branch bearing near its apex numerous,
racemose-capitate, greenish-white flowers, the heads, including
the anthers, about 3 cm in diameter. Pedicels about 2 mm
long, appressed-pubescent with short fulvous hairs. Calyx nar-
rowly funnel-shaped, about 4 mm long, sharply 5-toothed, densely
appressed fulvous pubescent. Corolla narrowly funnel-shaped,
twice as long as the calyx, 5-lobed, the lobes 2 mm long or less,
oblong, acute; stamens long-exserted.

PALAWAN, Taytay, Merrill 9259, May 15, 1913, in forests near the sea
and slightly above sea level.

A very characteristic species apparently most closely allied to Albizzia
pedicellata Baker, of the Malay Peninsula.

PARKIA R. Brown

PARKIA SHERFESEEI sp. nov.

Arbor alta, plus minusve ferrugineo-pubesecentibus; foliis cir-
citer 50 cm longis; pinnis 8- ad 12-jugatis, inferioribus bre-
vioribus; foliolis 11- ad 16-jugatis, oblongis, coriaceis, usque
ad 2.5 cm longis, 1 cm latis, sessilibus, basi subtruncatis, apice
rotundatis vel leviter inaequilateraliter retusis; pedunculis usque
ad 30 em longis, inflorescentiis junioribus obovoideis, dense fer-
rugineo-pubescentibus.

A tree reaching a height of 27 meters, the trunk 1.2 m in
diameter, the young branchlets, rachis, secondary rachises, pe-
duncles and inflorescence rather prominently ferruginous-pubes-
cent with short hairs. Leaves up to 50 cm long, the pinnae
8 to 12 pairs, the lower ones about 10 cm long, the intermediate
and upper ones up to 16 cm long; leaflets coriaceous, rather
pale when dry, oblong, up to 2.5 cm long, and 1 cm wide, sessile,
base rounded-truncate, inequilateral, apex rounded or somewhat
inequilaterally retuse, the lateral nerves slender, anastomosing,
indistinct. Peduncles up to 30 cm long, the inflorescence (in
bud) obovoid or pyriform, rather abruptly contracted below
the middle, 4 to 5 cm long, about 2.5 cm in diameter, the bract-
eoles. densely imbricate, spatulate, about 7 mm long, glabrous

10 The Philippine Journal of Science 1915

except the upper exposed portion which is densely ferruginous-
pubescent. Calyx about 7 mm long, the tube glabrous, the lobes
densely pubescent.

MINDANAO, Agusan Subprovince, near Butuan, For. Bur. 21962 Sher-
fesee, Cenabre, & Ponce, May 22, 1914, along nipa swamps at sea level,
locally known as cunding.

A very characteristic species, recognizable at once by its ferruginous
indumentum, its very dense inflorescences, and its unusually large leaflets.

PITHECOLOBIUM Martius

PITHECOLOBIUM CAULOSTACHYUM sap. nov.

Arbor 5 ad 10 m alta, subglabra; foliis bijugatis, circiter
20 cm longis, pinnis 6-foliolatis, foliolis chartaceis, lanceolatis
vel oblongo-lanceolatis, usque ad 11 cm longis, acuminatis, basi
acutis, nervis utrinque circiter 9; paniculis parce breviter hir-
sutis, paucifloris, usque ad 5 cm longis, plerumque fasciculatis,
caulinis et in ramis vetustioribus, corolla circiter 5 mm longa.

A tree 5 to 10 m high, the panicles mostly fascicled on the
trunk and larger branches. Branchlets slender, grayish-brown,
glabrous, the younger ones somewhat puberulent. Leaves 1-
jugate, about 20 cm long, the rachis 2 to 3 cm long, slightly
puberulent, usually with two prominent glands, one between
the pinnae, and one at the upper two-thirds; pinnae with 6-
leaflets opposite, the lower subalternate, rachis puberulent, with
one gland between each of the two upper pairs of petiolules
and usually one gland below the lowermost leaflet. Leaflets
chartaceous, lanceolate or oblong-lanceolate, 7 to 11 cm long,
2 to 3 cm wide, dark-brownish when dry, shining, glabrous,
the apex rather slenderly and sharply acuminate, base acute;
lateral nerves about 9 on each side of the midrib, slender, anas-
tomosing; petiolules 2 mm long or less, puberulent, becoming
glabrous. Panicles fascicled, mostly on the trunk and larger
branches, occasionally a few on the ultimate branchlets, 5 cm
long or less, few-flowered, sparingly hirsute with short hairs.
Flowers white, two or three subcapitately disposed at the tip
of each ultimate branchlet, the subtending bracteoles narrow,
about 1 mm long. Calyx cup-shaped, about 1.5 mm long, gla-
brous or nearly so, the teeth short, acute. Corolla campanulate,
about 5 mm long, the tube 2.5 mm long, the lobes as long as
the tube, recurved, oblong, obtuse or subacute. Stamens about
25, the free portions of the filaments about 1 cm long, the
lower 2 to 3 mm united into a tube. Ovary glabrous, oblong-
cylindric, 2 mm long; style slender, about 13 mm long.

x%,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 11

SAMAR, Mount Cauayan, Phil. Pl. 1637 Ramos, April, 1914, on forested
slopes.

The most striking character of this species is its cauline inflorescence.
It is manifestly allied to Pithecolobium pauciflorum Benth., but differs
not only in the disposition of its inflorescences, but in its very differently
shaped, more numerously nerved leaflets. In vegetative characters it
closely resembles Pithecolobium bigeminum Benth. as represented by
Thwaites C. P. 644.

PITHECOLOBIUM MULTIFLORUM sp. nov.

Arbor 6 or 8 m alta partibus junioribus inflorescentiisque
plus minusve ferrugineo-pubescentibus; foliis circiter 40 cm
longis, bijugatis, pinnis inferioribus 4-foliolatis, superioribus
8-foliolatis, foliolis subcoriaceis, oblongis vel oblongo-lanceolatis,
breviter obtuse acuminatis, basi acutis ad subrotundatis, usque
ad 14 cm longis, nervis utrinque circiter 9, subtus prominentibus;
paniculis terminalibus, quam folia longioribus, e basi ramosis,
multifloris; floribus subcapitatis, capitulis 4- vel 5-floris, in
ramulis plus minusve confertis, petalis fere liberis.

A tree 6 to 8 m high, closely allied to Pithecolobium ellipticum
Hassk., the branches dark-colored when dry, glabrous or nearly
so, the branchlets sparingly brown-pubescent with short hairs,
the primary and secondary rachises of the leaves and the in-
florescence with similar hairs, the younger parts of the inflores-
cence densely ferruginous-pubescent. Leaves about 40 cm long,
evenly 2-jugate, the rachis about 10 cm long, with a single very
large and prominent gland about 1.5 cm above the base, none
between the pinnae; lower pinnae 2-jugate, the rachis about
3.5 cm long, with a gland between the petioles of the distal
leaflets; upper pinnae 4-jugate, about 30 cm long, with a large
gland between the upper two pairs of petiolules, the lower two
leaflets alternate, the others opposite. Leaflets subcoriaceous,
oblong to rather broadly oblong-lanceolate, entirely glabrous,
shining, brownish-olivaceous, the lateral ones somewhat falcate
or inequilateral, shortly and bluntly acuminate, base acute to
somewhat rounded, or at least rounded on one side of the petio-
lule, the lower ones smaller than the upper, 8 to 14 cm long,
3.5 to 5 ecm wide; lateral nerves about 9 on each side of the
midrib, prominent; petiolules about 5 mm long. Panicles term-
inal, ample, many-flowered, branched from or near the base,
as long as or longer than the leaves, the lower branches up to
40 cm in length, the secondary branches in the upper two-thirds.
Flowers in small heads on the ultimate branchlets, the heads
4- or 5-flowered, often crowded, the subtending bracteoles ovate,
about 1 mm long. Calyx brown, about 3 mm long, rather

2 The Philippine Journal of Science 1915

densely appressed-pubescent with short, shining, tawny hairs,
the teeth broad, subacute, 0.5 mm long. Corolla and stamens
straw-colored, the petals free quite to the base or merely slightly
connate below, about 6 mm long, 2 mm wide, acute or obtuse,
pubescent externally. Stamens indefinite, the filaments at least
1 cm long, the lower 2 mm united into a tube. Pods unknown.

LuzON, Benguet Subprovince, railroad grade west of Baguio, Phil. Pl.
1779 Merrill May, 1914, in ravines along small streams, altitude about
1,300 meters.

A species manifestly closely allied to Pithecolobium ellipticum Hassk.,
from which its differs in its very differently shaped, smaller, more numer-
ously nerved leaflets, its petals free or nearly so, and other characters;
the duplicates were erroneously distributed as Pithecolobium platycarpum
Merr., to which species the present form is not closely allied.

NEPTUNIA Loureiro

NEPTUNIA OLERACEA Lour. Fl. Cochinch. (1790) 654.

MINDANAO, Butuan Subprovince, Bunauan, E. H. Taylor, September,
1913.

Widely distributed in the tropics of both hemispheres.

This species was credited to the Philippines by F.-Villar,’ but was
excluded by me° as at the time I revised the Philippine Leguminosae as
F.-Villar’s record was based on Cuming 2352, which was from Malacca,
not from the Philippines. Taylor’s specimen is the first collection of the
species in the Philippines.

KOOMPASSIA Maingay

KOOMPASSIA EXCELSA (Becc.) Taubert in Engl. & Prantl Nat. Pflan-
zenfam. 3° (1891) 156; Bericht Deutsch. Bot. Ges. 10 (1892) 641,
bina) of As:

Abauria excelsa Becc. Malesia 1 (1877) 169; Nelle Foreste di Borneo
(1902) 172, f. $4.

PALAWAN, Alphonso III, For. Bur. 21580 Danao, May 10, 1914, in
forests, altitude about 20 meters, flowers fragrant, light-yellow, locally
known by the Tagbanuas as manggis.

Beccari proposed the genus Abauria for this plant, but Taubert has
reduced it to Koompassia, although the fruits are not definitely known.
The Palawan specimen is manifestly identical with the Bornean species,
agreeing in all essentials with Beccari’s description and figure, and with
a Sarawak specimen, Foxworthy 334, collected under the native name
tapang, the same native name cited by Beccari. Dr. Foxworthy’s spec-
imen is sterile, but with it is a detached fruit, picked up from the ground,
which may or may not belong to the species; this fruit is the characteristic
winged one of Koompassia, strongly resembling that of Koompassia bec-
cariana Taubert. A very interesting addition to the few known species
confined to the Philippines and Borneo, most of which are not found in
the Philippines proper but in Palawan and in the Sulu Archipelago.

* Novis. App. (1880) 73.
‘Philip. Journ. Sci. 5 (1910) Bot. 136.

: Ds

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 13

CYNOMETRA Linnaeus

CYNOMETRA COPELANDI!I (Elm.) comb. nov.

Gleditschia copelandii Elm. Leafl. Philip. Bot. 2 (1910) 695.

Srpvyan, Elmer 12234, April, 1910 (type number).

The species is in all essential characters a member of the Cynometreae-
Caesalpinioideae, not of the Eucaesalpineae. It is anomalous in Cyno-
metra in that the ovules are solitary. The species is an exceedingly
characteristic one, notably in its leaf-characters.

BAUHINIA Linnaeus
BAUHINIA PAUCIFLORA sp. nov. § Phanera.

Frutex scandens inflorescentiis exceptis glaber vel subtus foliis
ad nervos parce pubescentibus; foliis oblongo-ovatis, usque ad
10 cm longis, 9- vel 11-nerviis, basi cordatis, 2-lobatis, lobis vix
ad medium coalitis, oblongis, obtusis, racemis terminalibus, pauci-
floris, floribus magnis, calycis tubo elongatis, angustis, 2 ad 2.5
em longis, petalis oblongo-obovatis, 4 ad 4.5 cm longis.

A scandent shrub, nearly glabrous except the brown-pubescent
inflorescence, the branches terete, brownish, glabrous. Leaves
alternate, oblong-ovate, 8 to 10 cm long, 4.5 to 6 cm wide, char-
taceous, the upper surface olivaceous, shining, the lower a little
paler, shining, often with few short hairs on the nerves, the
base distinctly cordate, 2-lobed, the lobes oblong, obtuse, extend-
ing about two-thirds to the base, the midrib of the leaf excurrent
as a slender, acicular, 4 mm long awn; nerves 9 to 11, prominent;
petioles slender, 3 to 4 cm long. Racemes terminal, few
flowered, pubescent with short brown hairs, the rachis and pe-
duncle 10 cm long or less. Flowers on the type two or three
in each raceme, their pedicels pubescent, 2 cm long, the
subtending bracteoles ovate, pubescent, about 5 mm long. Calyx-
tube cylindric, 2 to 2.5 cm long, 3 mm in diameter, brown-
pubescent, the lobes lanceolate, acuminate, pubescent externally,
2.5 to 3 cm long, 0.5 cm wide, the limb, in bud, ovoid-oblong,
about 2 cm in length. Petals oblong-obovate, obtuse, glabrous,
white or somewhat yellowish, narrowed to the short claws. Fer-
tile stamens 3, the anthers oblong, 1 cm in length. Ovary pubes-
cent with appressed, brown, deciduous hairs; style glabrous.

Luzon, Province of Tayabas, Dap-Dap Point, Bur. Sci. 18113 Fox-
worthy & Ramos, March, 1911, in forests.

A well marked species manifestly allied to Bauhinia warburgii Perk.,
but with very differently shaped leaves, the lobes much narrower.

CANAVALIA DeCandolle
CANAVALIA MACROBOTRYS sp. nov.

Secandens, racemis obscure pubescentibus exceptis glabra;
foliolis chartaceis, ovatis, prominente acute acuminatis, basi

14 The Philippine Journal of Science 1915

rotundatis, usque ad 14 cm longis, nervis utrinque 4 vel 5, sub-
tus prominentibus; racemis axillaribus, circiter 70 cm longis;
floribus roseis, 3 cm longis, breviter pedicellatis.

A scandent apparently herbaceous plant, glabrous except the
slightly pubescent inflorescence, the branches slender, terete,
greenish-straw-colored when dry. Petioles about 7 cm long,
the rachis extended 3 to 4 cm above the insertion of the lateral
leaflets. Leaflets ovate, chartaceous, 12 to 14 cm long, about
7.5 cm wide, olivaceous and shining when dry, apex rather
slenderly and prominently acuminate, the acumen apiculate, base
rounded, the central leaflet equilateral, the lateral ones somewhat
inequilateral; lateral nerves 4 or 5 on each side of the midrib,
prominent on the lower surface; petiolules about 7 mm long.
Racemes axillary, solitary, about 70 cm long, the lower 25 to
30 cm without flowers, the rest with prominent alternate nodes,
flowering in sequence from the base upward, the flowers below
falling, persisting only in the apical part. Flowers pink, about
3 cm long, short-pedicelled, the calyx about 1.5 cm long, the upper
lip cleft, very much larger than the lower lip. Standard about
2.5 cm wide, retuse.

SAMAR, Cauayan Valley, Bur. Sci. 17517 Ramos, March 14, 1914.

A species strongly characterized by its rather slenderly acuminate,

prominently nerved leaflets and especially by its greatly elongated racemes
which reach a length of at least 70 cm.

KUNSTLERIA Prain

KUNSTLERIA ATRO-VIOLACEA (Elm.) comb. nov.

Derris atro-violacea Elm. Leafi. Philip. Bot. 5 (1913) 1798.

The type is Elmer 18105, from Palawan, in flower, April, 1911. Al-
though the fruits are unknown J do not hesitate to transfer the species to
Kunstleria.

DALBERGIA Linnaeus f.

DALBERGIA SUBALTERNIFOLIA (Elm.) comb. nov.

Derris subalternifolia Elm. Leafi. Philip. Bot. 5 (1913) 1801.

The type is Elmer 12965 from Palawan, in flower, April, 1911, and is
in all respects a Dalbergia, not a Derris. It seems to be allied to Dal-
bergia densa Benth. which extends from Amboina and New Guinea to
Australia, and may not prove to be distinct from Bentham’s species.
Elmer’s specimen has 3-foliolate leaflets which are distinctly coriaceous;
Merrill 9345, from Palawan, which is probably referable to the same
species, however, has much thinner leaflets varying in number from 2
to 5.

DALBERGIA RETICULATA sp. nov. § Sissoa, Podiopetalum.

Frutex scandens, subglaber; foliis circiter 10 cm longis, 5- ad
7-foliolatis, foliolis chartaceis, late elliptico-ovatis, vel late ellip-

dt

—

x¢1 Merrill: New or Noteworthy Philippine Plants, XI 15

ticis, rotundatis vel leviter retusis, usque ad 4.5 cm longis;
inflorescentiis axillaribus, racemosis, brevibus, paucifloris; stami-
nibus monadelphis; leguminibus anguste oblongis, membrana-
ceis, samaroideis, circiter 7 cm longis, 1.3 ad 2 cm latis, omnibus
partibus valde reticulatis; seminibus 1 vel 2.

A scandent shrub, glabrous except the somewhat pubescent
infiorescence and the leaflets which are very sparingly pubescent
on the lower surface. Branches brownish, wrinkled when dry,
lenticellate. Leaves alternate, about 10 cm long, each with from
5 to 7 leaflets, the leaflets opposite or alternate, membranaceous
or chartaceous, broadly elliptic-ovate to broadly elliptic, some-
what brownish when dry, lower surface a little paler than the
upper, rounded at both ends or the apex slightly retuse, the
lower surface with few, scattered, brownish, very short hairs,
the upper surface glabrous; lateral nerves about 8 on each side
of the midrib, slender, not prominent; petiolules 2 to 3 mm
long. Inflorescence racemose, axillary, solitary, few-flowered,
the racemes sparingly pubescent, 2.5 to 4 cm long. Calyx spar-
ingly pubescent, about 3.5 mm long. Wings about 6 mm long,

the lamina about 3 mm long, 1.6 mm wide, obtuse or rounded,

base acute on one side, rounded on the other, the claw very
slender, as long as the lamina; standard and keel not seen.
Stamens all united into a sheath split down one side to the base.
Pods narrowly oblong, membranaceous, about 7 cm long, 1.3
to 2 cm wide, apex rounded and minutely apiculate, base some-
what decurrent and with a slender stalk about 8 mm long, all
parts of the valves prominently and rather laxly reticulate; seeds
one or two, central, immature.

Luzon, Province of Laguna, Mount Maquiling, For. Bur. 21406 Vellamil,
April 26, 1914, in forests along the Molauin trail beyond the hot springs,
altitude about 500 meters.

A very characteristic species, anomalous in the genus in its racemose
inflorescence, and further characterized by its very thin, prominently
reticulate pods. The specimen is with immature fruits, but portions of
the flower were found on one raceme, leaving very little doubt as to the
correctness of the genus and section.

MELILOTUS Tournefort

MELILOTUS INDICA (Linn.) All. Fl. Ped. 1 (1785) 308.
Trifolium melilotus-indica Linn. Sp. Pl. (1753) 765.

Luzon, Benguet Subprovince, Baguio, Merrill 9700, May, 1914, dis-
turbed soil along the railroad grade, also as a weed in the Forestry
nursery.

A plant of wide distribution, recently introduced into the Philippines.

16 The Philippine Journal of Science 1915

STRONGYLODON Vogel

STRONGYLODON PAUCINERVIS sp. nov.

Frutex scandens glaber; foliis 3-foliolatis, foliolis chartaceis,
ovatis, obscure latissime obtuse acuminatis, usque ad 12 cm
longis, nervis utrinque circiter 5; racemis ut videtur caulinis,
fasciculatis, circiter 20 cm longis, multifloris, floribus azureo-
violaceis ad nodos leviter elongatis subracemose dispositis, cur-
vatis, circiter 3 cm longis.

A scandent glabrous vine of large size, the branchlets terete,
brownish, 3 to 4 mm in diameter. Leaves 3-foliolate, their pe-
tioles 6 to 9 cm long, the stipules oblong, obtuse, strongly nerved,

about 4 mm long, deciduous, the stipels linear, about 3 mm long;

leaflets chartaceous, the terminal one equilateral, the lateral ones
strongly inequilateral, ovate, 10 to 12 cm long, 5 t0 6 cm wide,
obscurely, very broadly, and bluntly acuminate, base rounded,
shining, subolivaceous; lateral nerves about 5 on each side of the
midrib, prominent; petiolules about 8 mm long. Racemes ap-
parently from the trunk, at least three in a fascicle, about 20 em
long, many flowered, the peduncles 4 to 6 cm long. Flowers
bluish-violet, about 3 cm long, corolla upcurved at nearly right
angles at about the middle, in bud nearly straight or only
slightly faleate, the nodes produced as short, about 5 mm long,
branchlets, the flowers racemosely arranged on the produced
nodes. Pedicels 1 cm long or less. Calyx cup-shaped, about
6 mm long, margins wavy. Standard about 3 cm long, when
spread nearly 2 cm wide, curved upward at about the middle,
rostrate-acuminate. Wings slightly falcate, the claw about 8
mm long, the lamina oblong, rounded, 1.5 cm long, 6 mm wide.
Keel strongly curved upward, 4 cm long, when spread 2 cm wide,
rostrate-acuminate. Ovary long stipitate, the ovules about 5.

BiuiRAN, Bur. Sci. 18858 McGregor, June 22, 1914, a very large vine
in forests, altitude about 300 meters.

Among the Philippine species with short racemes well characterized

by its fascicled racemes, its produced nodes, its numerous flowers, and its
few-nerved leaflets.

STRONGYLODON MEGAPHYLLUS sp. nov.

Scandens, ramulis junioribus exceptis glaber; foliolis oblongo-
ovatis, coriaceis, in siccitate brunneis, usque ad 28 cm longis,
basi subacutis, apice obtusis vel retusis, nervis utrinque circiter
8, subtus prominentibus; inflorescentiis angustissime paniculatis,
30 ad 40 cm longis, ut videtur multifloris; floribus azureo-
violaceis, 5 vel 6 cm longis.

A scandent plant, apparently of large size, quite glabrous

~x,G,1 Merrill: New or Noteworthy Philippine Plants, XI 17

except the young branchlets which are more or less appressed-
pubescent, the branches brown, terete, 4 to 5 mm in diameter.

Leaves 3-foliolate, the leaflets oblong-ovate, coriaceous, brown

and shining when dry, glabrous, 20 to 28 cm long, 9 to 16 cm
wide, subequally narrowed to the acute or subacute base and
to the blunt or somewhat retuse apex, equilateral, the lateral
nerves about 8 on each side of the midrib, prominent on the
lower surface, the petioles about 1 cm long. Racemes 30 to
40 em long, apparently many-flowered, narrowly paniculate,
flower bearing in the upper two-thirds, the branchlets from the
nodes 1 to 1.5 cm long, each bearing several flowers, judging
from the scars of fallen pedicels. Pedicels 1.5 to 2 cm long.
Calyx about 1 cm long, nearly 1 cm in diameter, truncate, the
upper side somewhat gibbous. Corolla blue-violet in color, very
strongly recurved. Standard about 5 cm long, 2.5 cm wide,
very sharply bent upward and backward from the mouth of
the calyx, acuminate, pubescent inside in the median portion
below, auriculate at the sharp curve inside; wings strongly
faleate-curved, their claws 1.5 cm long, the lamina. 2.5 cm long,
1 em wide, obtuse, one side auriculate at the base. Keel very
strongly recurved, when straightened out about 7 cm long, long-
acuminate, rostate. Fruit apparently fleshy, when dry falcate-
ovoid, about 5 cm long, 2.5 cm in diameter, not compressed,
beaked at the apex, contracted at the base into a short stipe.

Luzon, Province of Rizal, Montalban, Loher 5935, April 15, 1906.

The most strongly marked species known from the Philippines, at once
recognizable by its unusually large leaflets, its large flowers, and the nodes
of the racemes produced as slender branchlets up to 1.5 cm in length,
making the inflorescence a narrow panicle rather than a true raceme.

LUZONIA Elmer

LUZONIA PURPUREA Elm. Leafl. Philip. Bot. 1 (1907) 220.

This genus and species were described from flowering specimens. The
fruits are fleshy, oblong to oblong-obovoid in outline, when fresh some-
what fleshy, cylindric, not at all compressed, about 12 cm long, 5.5 cm
thick, smooth, shining, purple, short-apiculate at the apex, the valves very
thick, apparently tardily dehiscent, firm, fleshy, greenish-white in color, 1
to 1.5 cm thick; sutures not prominent. Seeds usually 4 in each pod,
ellipsoid, about 3 cm long and 2 cm thick.

Luzon, Benguet Subprovince, Pacdal, Merrill 9685, May, 1914.

MILLETTIA Wight & Arnott
MILLETTIA BRACHYCARPA sp. nov.

Arbor circiter 7 m alta, inflorescentiis exceptis glabra; foliis
circiter 20 cm longis, imparipinnatis, foliolis circiter 7, oblongo-
ovatis, subcoriaceis, breviter acuminatis, usque ad 13 cm longis;

131281-—_2

18 The Philippine Journal of Science 1915

floribus ignotis, racemis axillaribus, solitariis, usque ad 10 cm
longis; leguminibus 1-spermis, oblongo-ellipticis, 6 cm longis, 3
cm latis, 1 cm crassis, utrinque angustatis, basi acutis, apice
rostrato-acuminatis, valvis lignosis.

A tree about 7 m high, glabrous except the inflorescence.
Branches terete, lenticellate, grayish or reddish-brown. Leaves
alternate, odd-pinnate, the petiole and rachis 10 to 12 em long;
leaflets usually 7, subcoriaceous, ovate to oblong-ovate, up to
13 cm long and 4.5 cm wide, the apex shortly and obtusely
acuminate, the base usually rounded, equilateral or somewhat
inequilateral, the upper surface brownish-olivaceous when dry,
shining, the lower paler; lateral nerves about 9 on each side
of the midrib, prominent; petiolules 5 to 8 mm long. Flowers
not known, the persistent rachises of the racemes axillary, soli-
tary, thickened, somewhat pubescent, with numerous thickened
protuberances. Pods brown when dry, oblong-elliptic, 1-seeded,
woody, about 6 cm long, 3 cm wide, and at least 1 cm thick,
narrowed below to the acute base and above to the rostrate-
acuminate apex, glabrous, the corners rounded, not angled, the
valves woody, shining, smooth. Seed solitary, subelliptic, some-
what narrowed to the rounded ends, about 3.5 em long, 2.38 em
wide, and nearly 1 cm thick.

PALAWAN, Silanga, in deserted clearings on slopes, Merrill 9578, May

24, 1913.
A species well characterized by its very thick, woody, 1-seeded pods.

CROTALARIA Linnaeus

CROTALARIA ORIXENSIS Willd in Ges. Naturf. Fr. Neue Schr. 4
(1803) 217; Baker in Hook. f. Fl. Brit. Ind. 2 (1876) 83.

LuZON, Manila, Bur. Sci. 19145 Guerrero, December, 1912, in waste
places, certainly of recent introduction.

For the identification of the above specimen I am indebted to Sir D.
Prain, director of the Royal Gardens, Kew, England, to whom a specimen
was sent. The species is of wide distribution in British India (Western
Peninsula), in Abyssinia, and is reported from tropical Africa. It is
undoubtedly a recent introduction in the Philippines.

SIMARUBACEAE
BRUCEA J. S. Miller

BRUCEA AMARISSIMA (Lour.) comb. nov.

Gonus amarissimus Lour. Fl. Cochinch. (1790) 658.
Brucea sumatrana Roxb. Hort. Beng. (1814) 12, Fl. Ind., ed. Carey,
1 (1832) 449; Hook. f. Fl. Brit. Ind. 1 (1875) 521; Lecomte Fl.
Gén. Indo-Chine 1 (1911) 698.
This widely distributed Indo-Malayan species is apparently quite common
in the southern Philippines, and is represented in our collections by spec-
imens from Negros, Leyte, Camiguin, Cebu, Palawan, Basilan, and Min-

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 19

danao, but is not reported from Luzon unless Cuming 987 from the Prov-
ince of Pangasinan is correctly referred here. In our extra-Philippine
material it is represented by specimens from Cochin China, southern China,
Malay Peninsula, Singapore, and Queensland. Loureiro’s specific name,
being much the older, is here adopted.

BRUCEA MACROBOTRYS sp. nov.

Frutex 1 ad 2 m altus partibus junioribus inflorescentiisque
minute adpresse pubescentibus; foliis usque ad 60 cm longis,
9-foliolatis, foliolis ovatis ad oblongo-ovatis, membranaceis, basi
late acutis ad rotundatis, apice tenuiter acuminatis, margine
integris vel leviter undulatis, distanter glandulosis, subtus par-
cissime pubescentibus, nervis utrinque 7 vel 8; paniculis angustis,
quam folia longioribus, cymis distantibus, circiter 1.5 cm longis;
floribus circiter 2 mm diametro.

An erect shrub 1 to 2 m high, the branches terete, pale-
brownish, lenticellate, glabrous, the younger ones somewhat
pubescent with short pale hairs as are the inflorescences and
parts of the leaves. Leaves up to 60 cm long, the rachis greenish
when dry, slightly pubescent; leaflets 9, ovate to oblong-ovate,
membranaceous, 12 to 17 cm long, 5 to 6 cm wide, olivaceous
and shining when dry, base broadly acute to rounded, apex rather
slenderly and sharply acuminate, margins entire or slightly. un-
dulate, the incipient teeth indicated by distant marginal glands;
lateral nerves 7 or 8 on each side of the midrib; petiolules
puberulent, about 6 mm long. Panicles axillary, up to 80 cm
long, very narrow, many flowered, the flowers in distant, pubes-
cent, rather many-flowered cymes about 1.5 cm in length, few
flowers opening at one time, their pedicels pubescent, slender,
4 to 5 mm long, the staminate ones about 2.2 mm in diameter.
Sepals 3, pubescent, oblong, acute, 1 mm long. Petals slightly
longer than the sepals, oblong-ovate, obtuse, very slightly pubes-
cent. Stamens about as long as the petals. Fruit ovoid, black
when dry, about 12 mm long, distinctly reticulate-rugose.

SamaR, Cauayan Valley, Phil. Pl. 1626 Ramos, April, 1914, in forests
along small streams.

A species manifestly allied to Brucea mollis Wall. and B. luzoniensis
Vid., closely approaching some forms of the latter. It differs in its larger
leaflets, longer leaves, much longer inflorescences, and larger fruits, but

in spite of these different characters may be only a variety of Wallich’s
species unusually large in size.

CANARIUM Linnaeus

CANARIUM HETEROPHYLLUM sp. nov. § Choriandra.
Arbor circiter 9 m alta, floribus exceptis glabra; ramulis
tenuibus, teretibus; foliis alternis, 1- ad 3-foliolatis, foliolis

20 The Philippine Journal of Science 1915

subcoriaceis, 7 ad 16 cm longis, integris, utrinque subaequaliter
angustatis, basi acutis, apice acuminatis, nervis utrinque 8 ad
10, subtus prominentibus; infructescentibus axillaribus, soli-
tariis, circiter 10 cm longis, racemosis; fructibus oblongis, cir-
citer 1.5 cm longis, obscure 3-angulatis, apiculatis, breviter pedi-
cellatis, pedicellis calycibusque breviter cinereo-pubescentibus.

A tree about 9 m high, glabrous except the flowers. Branches
slender, terete, grayish, the very slender branchlets reddish-
brown as are the petioles and rachises. Leaves alternate, the
rachis and petiole 2 to 6 cm long, slender; leaflets 1 to 3,
subcoriaceous, elliptic-oblong, olivaceous, somewhat shining
when dry, entire, of about the same color on both surfaces, 6
to 17 cm long, 3 to 7 cm wide, subequally narrowed to the
acute base and to the rather prominently acuminate apex;
lateral nerves 8 to 10 on each side of the midrib, prominent
on the lower surface, anastomosing, the reticulations promi-
nent; petiolules of the lateral leaflets, when present, about 8
mm long, the petioles of the simple leaves up to 3 cm long.
Infructescence simple, racemose, axillary, slender, up to 10 cm
long, apparently flower bearing only in the upper part.
Flowers not seen. Calyx, in fruit, cinereous-pubescent with
short hairs, the lobes 3, ovate, acute or obtuse, about 2.5 mm
long, the pedicels stout, pubescent, 1.3 mm long. Mature fruits
oblong, about 1.5 cm long, about 7 mm in diameter, apiculate,
indistinctly, or the upper part distinctly 3-angled, the pericarp
1 mm thick or less, brownish, somewhat wrinkled when dry.

Luzon, Province of Cagayan (or Subprovince of Apayao), Tamoc, Bur.
Sci. 18878 Ramos, January, 1908.

A species well characterized by its 1- and 3-foliolate leaves, both types
occurring on the same branches. In this character it differs from all
known species of the genus. On account of its fruit characters and its
general aspect I have placed it in the same section with Canarium villosum
F.-Vill. (C. cumingii Eng].).

CANARIUM DOLICHOPHYLLUM sp. nov.

Arbor circiter 18 m alta plus minusve pubescentibus; ramulis
inecrassatis circiter 1 cm diametro; foliis circiter 70 cm longis,
foliolis 13, lanceolatis vel oblongo-lanceolatis, coriaceis, in sic-
citate pallidis, nitidis, integris, usque ad 25 cm longis et 5 cm
latis, apice tenuiter caudato-acuminatis, basi subrotundatis,
supra glabris, subtus pubescentibus; nervis utrinque 18 ad 20,
subtus valde prominentibus, reticulis primariis subparallelis,
prominentibus; infructescentibus axillaribus, racemosis, circiter
20 cm longis, ferrugineo-pubescentibus; fructibus ovoideis ad
subellipsoideis, obtusis, glabris vel leviter hirsutis, pallidis,

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI VAI

obscurissime 3-angulatis, circiter 2 cm longis, in siccitate valde
rugosis.

A tree about 18 m high. Branchlets thickened, about 1 em
in diameter, brown when dry, pubescent with short, reddish-
brown hairs. Leaves up to 70 cm in length, the rachis and pe-
tioles glabrous, pale-brown when dry. Leaflets lanceolate to
oblong-lanceolate, coriaceous, entire, 20 to 25 cm long, 4 to 5
cm wide, the upper surface pale, shining, the lower ferruginous-
pubescent with short hairs on the midrib, nerves, and very prom-
inent reticulations, the base rounded to subacute, nearly equi-
lateral, the apex slenderly caudate-acuminate, the acumen up to
2 cm long, blunt; lateral nerves 18 to 20 on each side of the
midrib, very prominent on the lower surface, curved-anastomos-
ing, the reticulations very prominent; petiolules glabrous or
slightly pubescent, rugose, 3 to8 mm long. Infructescence race-
mose, axillary, solitary, rather stout, about 20 cm long, rather
densely ferruginous-pubescent with short hairs, the rachis 4
to 5 mm in diameter. Fruits numerous, on stout peduncles 8
mm long or less, the lower peduncles rarely branched, the per-
sistent calyx-lobes 3, coriaceous, ovate, obtuse, about 4 mm long,
the fruits rather pale when dry, glabrous, or with few scattered
hairs, ovoid to ellipsoid, about 2 cm long, 1.3 cm in diameter,
obtuse, obscurely 3-angled, the pericarp very prominently wrin-
kled when dry.

MINDANAO, Butuan Subprovince, near Amparo, Agusan Valley, For.
Bur. 20749 Rafael & Ponce, October 10, 1913, in forests slightly above
sea level.

A species well characterized by its long leaves, and especially by its

elongated, relatively narrow, caudate-acuminate leaflets which are very
prominently reticulate beneath.

CANARIUM LAGUNENSE sp. nov. § Choriandra ?

Arbor 12 ad 20 m alta plus minusve breviter brunneo-pubes-
centibus vel floribus ferrugineo-pilosis; foliis 15 ad 25 cm longis,
foliolis 5 vel 6, coriaceis, ellipticis ad oblongo-ellipticis, in sic-
citate brunneis, nitidis, integris, acuminatis, usque ad 12 em
longis, supra glabris, subtus ad costa nervisque breviter brunneo-
puberulis, nervis utrinque circiter 12, subtus valde prominen-
tibus; inflorescentiis axillaribus, folia aequantibus vel paullo
longioribus, floribus ¢ numerosis, sessilibus, fasciculatis, 6 ad
7 mm longis, 2° angustissime paniculatis; fructibus ovoideis,
circiter 12 mm longis, apiculatis, extus in siccitate rugosis,
glabris, endocarpio osseo, circiter 2 mm crasso.

A tree 12 to 20 m high, branchlets, petioles, inflorescences
and leafiets on the nerves and reticulations of the lower surface

2? The Philippine Journal of Science 1915

more or less covered with very short dark-brown hairs.
Branches terete, brownish or grayish, glabrous, lenticellate, 5
to 7 mm in diameter. Leaves 15 to 25 cm long, alternate, the
leafiets 5 or 6, coriaceous, brown and shining when dry, entire,
5 to 12 cm long, 4 to 6 em wide, the apex acuminate, the acumen
usually abrupt, short, blunt, the base acute to rounded, mostly
nearly equilateral; lateral nerves about 12 on each side of the
midrib, very prominent on the lower surface, anastomosing,
the reticulations prominent, the nerves, midrib, and reticulations
with very short, stout, short, glandlike hairs, the upper surface
entirely glabrous; peticlules 5 to 10 mm long; stipules linear-
lanceolate, thick, stiff, straight or curved, brown-pubescent,
about 5 mm long. Inflorescences axillary, solitary, numerous,
equalling or a little longer than the leaves, brown-pubescent, or
the flower clusters ferruginous-pubescent. Male flowers numer-
ous, fascicled, the fascicles widely separated, each with numerous
ovate, hirsute, 1.2 to 1.8 mm long bracteoles, the flowers sessile.
Calyx cylindric, 4 mm long, hirsute, the lobes 3, broadly ovate,
obtuse, about 2 mm long. Petals 3, oblong, 6 mm long, 2 mm
wide, obtuse, the upper two-thirds externally pubescent. Fila-
ments six, 3 to 3.5 mm long, flattened below and slightly united
for the lower 0.5 mm forming an obscure disk; anthers oblong,
1.8 mm long. Rudimentary ovary obovoid, glabrous, 1.5 mm
in diameter, more or less evidently 3-cleft and readily separat-
ing into three parts. Female or perfect flowers apparently sim-
ilar to the male, the infructescence narrowly paniculate, the
lower branches up to 3 cm in length, each branch with from 1
to 3 fruits, the pedicels stout, 5 to 10 mm long. Mature fruits
ovoid, not at all 3-angled, apiculate, about 1.2 cm long, 7 to
8 mm in diameter, the pericarp thin, wrinkled when dry, gla-
brous, the endocarp 1-celled, bony, about 2 mm thick.

LuZzON, Province of Laguna, San Antonio, Bur. Sci. 14945 (type), 20468,
20579 Ramos; Mount Banajao, For. Bur. 19722 Barber, in flower in Feb-
ruary, in fruit in April and May, growing in forests up to an altitude of
500 meters.

A species in many characters closely resembling Canarium villosum
F.-Vill. (C. cumingti Engl.), and probably as closely allied to that species
as any other. It is distinguished by its very short, dark-brown indumentum
which appears on the younger parts of the inflorescence, and on the lower

surface of the leaflets. The male flowers are disposed in sessile, dense,
distant fascicles and are ferruginous-pubescent.

CANARIUM BARNESII sp. nov. § Choriandra.

Arbor circiter 20 m alta a C. lagunense differt foliis longio-
ribus, foliolis magis numerosis, 6 vel 7, inflorescentiis multo

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 23

brevioribus, 5 ad 10 ecm longis, floribus @ solitariis, haud
fasciculatis.

A tree about 20 m high, most parts except the upper surfaces
of the leaflets more or less covered with very short brown hairs.
Leaves about 35 cm long, the leaflets 6 or 7, coriaceous, oblong-
elliptic to oblong-obovate, 7 to 12 cm long, 3.5 to 6 em wide,
rather dark-brown when dry, the upper surface glabrous, shin-
ing, base acute to rounded, apex prominently and abruptly
acuminate, the acumen blunt, 1 cm long or less; lateral nerves
12 to 14 on each side of the midrib, prominent on the lower sur-
face, anastomosing, the reticulations rather close, prominent;
petiolules 1 to 1.5 cm long; stipules stout, stiff, pubescent, curved,
about 5 mm long. Spikes 5 to 10 cm long, simple, the flowers
scattered below, solitary, above rather close, sessile. Calyx
oblong-ovate, about 4 mm long, pubescent, the teeth 3, broadly
ovate, obtuse, 1 mm long. Petals 3, narrowly oblong, acute to
obtuse, about 5 mm long, 1.5 mm wide, pubescent externally in
the upper part. Disk annular, glabrous, rather thick, more or
less lobed, 1 mm high or less; filaments 6, slender, 1.5 mm long,
attached to the outer upper part of the disk; anthers 1 mm long.
Ovary globose-obovoid, pubescent, 1.5 mm in diameter; style
stout, pubescent, about 1 mm long, cylindric; stigma capitate,
obscurely sulcate.

MASBATE, Marintoc River, collected by P. T. Barnes, May, 1903, distrib-
uted as Merrill 2617. In forests, locally known as mili-pili.

A species in facies much resembling Canarium lagunense Merr., but
with longer leaves, more numerous leaflets, very much shorter inflores-
cences, the flowers not at all fascicled, and the flowers in structure quite
different from those of C. lagunense. The indumentum on the younger

parts, inflorescence, and lower surfaces of the leaflets is quite the same in
both forms.

CANARIUM OLIGANTHUM sp. nov. § Crassipyrena.

Arbor, ut videtur alta, partibus junioribus plus minusve pube-
rulis; foliis circiter 70 cm longis, foliolis circiter 9, amplis, in-
tegris, chartaceis vel subcoriaceis, in siccitate brunneis, usque
ad 25 cm longis, et 11 cm latis, acuminatis, nervis utrinque 15
ad 17, prominentibus; inflorescentiis axillaribus, solitariis, cir-
citer 15 cm longis, anguste paniculatis, paucifloris; floribus longe
pedicellatis, ebracteolatis, circiter 8 mm longis.

A tree, apparently tall, the trunk about 50 cm in diameter,
the branchlets, petioles, midribs on the lower surface of the
leaflets, and inflorescence more or less puberulent with pale-
brownish hairs. Branches terete, lenticellate, stout, striate, the
ultimate branchlets about 1 cm in diameter, pale-brownish.

94 The Philippine Journal of Science 1915

Leaves ample, up to 70 cm long, the petiole long, about 5 mm
in diameter, and with the rachis and petiolules puberulent; leaf-
lets ovate to oblong, 15 to 25 cm long, 8 to 11 cm wide, chartace-
ous to subcoriaceous, brown and shining when dry, the lower
surface a little paler than the upper, entire, apex acuminate,
base rounded, subtruncate, or slightly cordate, broad, equilateral,
the upper surface quite glabrous, the lower pubescent on the
midrib; petiolules 1 to 2 cm long; stipules not seen, apparently
early deciduous. Panicles axillary, solitary, 15 cm long or less,
narrowly pyramidal, branched at or above the middle, the
branches few, scattered, the lower ones 2.5 cm long or less, each
branch with one or two long-pedicelled, ebracteolate flowers, or
if the bracteoles are present then small and early deciduous;
pedicels 1 cm long or less. Flowers 3-merous, about 8 mm long,
the calyx cylindric, 6 to 7 mm long, puberulent, the teeth three,
broadly triangular-ovate, acute, 2 mm long. Petals 3, oblong,
obtuse, 6 to 7 mm long, 3 mm wide, the upper one-third spread-
ing. Stamens 6; filaments 2 mm long, nearly free, flattened and
slightly enlarged below, apparently connate into a short obscure
disk. Ovary ellipsoid, glabrous, about 4 mm long, narrowed
upward into the short, cylindric, stout, 1 mm long style; stigma
capitate.

MASBATE, Marintoc River, P. T. Barnes, May, 1903, distributed as
Merrill 2618. In forests along the river, altitude about 16 m, locally
known as pili and yielding a white resin.

A species manifestly allied to Canarium luzonicum A. Gray, but well
characterized by its large leaves, ample leaflets, and especially by its

narrowly pyramidal, few-flowered panicles, the flowers ebracteolate, their
pedicels about 1 cm long.

CANARIUM NITENS sp. nov.

Arbor circiter 15 m alta, infiorescentiis puberulis exceptis
glabra; foliis usque ad 35 cm longis, 7- vel 9-foliolatis, foliolis
crasse coriaceis, utrinque nitidis, integris, oblongis, acuminatis,
8 ad 13 cm longis, nervis utrinque circiter 10, subtus promi-
nentibus; paniculis obscure brunneo-puberulis, terminalibus,
quam folia brevioribus; fructibus anguste ovoideis.

A tree about 15 m high, glabrous except the inflorescence.
Branches brown, terete, glabrous, wrinkled when dry. Leaves
alternate, up to 35 cm in length. Leaflets oblong, thickly coria-
ceous, prominently shining on both surfaces, the lower a little
paler then the upper, 7 or 9 to each leaf, 8 to 13 em long, 2.5
to 3.5 cm wide, entire, rather prominently acuminate, base acute,
often somewhat inequilateral and sometimes rounded on one
side and acute on the other; lateral nerves about 10, very

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 25

prominent on the lower surface, strongly curved, obscurely anas-
tomosing near the margins, the reticulations not at all prom-
inent; petiolules of the lateral leaflets 0.6 to 1.5 cm long, of the
terminal one up to 4 cm in length. Panicles terminal, 13 cm
long in fruit, brown-puberulent, branched from the base. Per-
sistent calyces about 3 mm in diameter, puberulent. Immature
fruits narrowly ovoid, somewhat inequilateral or somewhat gib-
bous, about 2 cm long, glabrous, not at all angled, the pericarp
wrinkled when dry.

MINDANAO, District of Zamboanga, Taglibao River, For. Bur. 18772
Foxworthy, DeMesa, & Villamil, June 17, 1912, growing at sea level
immediately back of the mangrove swamp, known to the Moros as batli.

The species is apparently a rather characteristic one, and is perhaps
as closely allied to Canarium calophyllum Perk., as to any other. It
differs from Canarium calophyllum, however, in its fewer nerved leaves
and in its much smaller persistent calyces. .

CANARIUM STENOPHYLLUM sp. nov. § Eucanarium, Monadelpha.

Arbor alta partibus junioribus exceptis glabra; foliis circiter
20 cm longis, foliolis 13 ad 15, lanceolatis, integris, usque ad 11
em longis, inaequilateralibus, leviter falcatis vel subrectis, basi
acutis, apice longe acuminatis; racemis ¢ axillaribus, solitariis,
8 ad 10 cm longis; floribus circiter 1.3 cm longis, cylindraceis,
filamentis deorsum connatis, disco libero.

A tree reaching a height of. 40 meters, glabrous except the
younger parts and the inflorescence. Branches terete, glabrous,
lenticellate, grayish-brown, about 5 mm in diameter, the branch-
lets more or less appressed-villous or hirsute, especially the
growing parts. Leaves alternate, about 20 cm long. Leaflets
18 to 15, lanceolate, inequilateral, straight or somewhat falcate,
chartaceous, entire, 1 to 2.4 cm wide, 6 to 11 cm long, narrowed
below to the acute or obtuse base, gradually narrowed upward
to the long and rather slenderly acuminate apex, when dry oli-
vaceous or subolivaceous, shining; lateral nerves about 9 on
each side of the midrib, slender, curved, anastomosing; petiolules
about 2 mm long; stipules not seen, if present very early deci-
duous. Male racemes axillary, solitary, 8 to 10 cm long, slightly
pubescent, the flowers somewhat crowded in the upper one-half,
white, their pedicels about 2 mm long. Calyx about 4 mm long,
the lobes 3, broadly ovate, 2 mm long. Petals 3, oblong, im-
bricate, glabrous, about 13 mm long, 5 mm wide. Stamens 6,
the filaments united below into a 2 mm high disk, the free parts
6 to 7 mm long; anthers oblong, 3 mm long. Disk free, oblong-
cylindric, hirsute, somewhat fluted, about 4 mm long, 2.5 mm
in diameter.

26 The Philippine Journal of Science 1916

LuZzON, Province of Camarines, Mount Isarog, Phil. Pl. 1550 Ramos,
November 20, 1913, in forests.

This species is probably as closely allied to Canarium perkinsiae Merr.
as to any other, differing in its much smaller, lanceolate leaves and in
its short pedicels. It is well characterized by its narrowly lanceolate
leafiets.

CANARIUM ELLIPSOIDEUM sp. nov. § Fucanarium.

Arbor alta, glabra (floribus ignotis) ; ramis incrassatis, gla-
bris, circiter 2 cm diametro, brunneis, nitidis; foliis alternis,
usque ad 55 em longis, foliolis circiter 9-jugis, coriaceis, nitidis,
ovatus ad oblongis, usque ad 16 cm longis, basi cordatis, apice
distincte acuminatis; inflorescentiis ut videtur paniculatis, axil-
laribus, solitariis, infructescentibus 10 ad 30 cm longis; fruc-
tibus ellipsoideis, circiter 3 cm longis, utrinque rotundatis, laevis
vel obscure rugosis, in siccitate castaneis, endocarpio longitu-
dinaliter 6-carinato.

A tall tree, nearly glabrous, the ultimate branches much thick-
ened, about 2 cm in diameter, dark-brown, shining, more or less
striate or wrinkled when dry, marked with scattered large scars
of fallen petioles, the very tip, above the ultimate leaves, densely
ferruginous-pubescent. Leaves more or less crowded toward
the ends of the branchlets, up to 55 em long, each with about
19 leaflets, odd-pinnate, the rachis and petiole rather stout, shin-
ing, striate; stipules none or very early deciduous. Leaflets op-
posite, the lower ones ovate, 8 to 10 cm long, the intermediate
and upper ones oblong, up to 16 cm long, 4 to 6 cm wide, shin-
ing on both surfaces, glabrous, rather pale when dry, the base
prominently and broadly cordate, apex acuminate, the acumen
blunt, rather stout; lateral nerves of the median leaflets about
16 on each side of the midrib, of the lower leaves about 10,
distinct, anastomosing; petiolules stout, about 5 mm long.
Flowers not seen. Infructescence axillary, solitary, 10 to 30
em long, dark-brown, glabrous, shining, somewhat paniculate.
Persistent calyx 3-lobed, very coriaceous, about 8 mm in diam-
eter, slightly pubescent externally, inside densely appressed
cinerous-pubescent, the lobes broadly ovate; pedicels stout, very
short. Fruit ellipsoid, about 3 cm long, rounded at both ends,
not at all 3-angled, the pericarp apparently fleshy, when dry
castaneous, slightly wrinkled, glabrous, or the younger fruits
with few, scattered, appressed, stiff, ferruginous hairs, the
endocarp bony, 3-celled, only a single cell developing a seed,
rounded at both ends, not at all 3-angled, but with three very
prominent primary longitudinal keels or ridges that unite with
each other at the base, and three intermediate keels or ridges,

x.c,1 Merrill: New or Noteworthy Philippine Plants, XI 27

nearly as prominent as the primary ones, which unite at the
apex but are evanescent near the base.

LUZON, Province of Bulacan, Angat, Bur. Sci. 21968 Ramos, September
22, 1913, in forests.

A striking species on account of its thickened stems, its long leaves,
and the almost entire absence of indumentum of any kind. The strictly
ellipsoid, very dark-brown, shining, nearly smooth fruits are exceedingly
characteristic, as is also the longitudinally 6-ridged endocarp, the fruit
not being at all triangular.

CANARIUM SANCHEZII sp. nov. § Monadelpha.

Arbor, omnibus partibus, alabastris exceptis, glabra; foliis
trifoliolatis, vix 20 ecm longis, foliolis elliptico-ovatis, subco-
riaceis, apice abrupte subcaudato-acuminatis; inflorescentiis
axillaribus, spicatis, quam petioli brevioribus; floribus 3-meris,
filamentis basi breviter connatis.

A tree, quite glabrous except the buds. Branches terete,
light-gray, rather slender. Leaves 3-foliolate, 14 to 18 cm long,
the petioles 2 to 3.5 cm long; leaflets elliptic-ovate, or the ter-
minal one somewhat obovate-elliptic, subcoriaceous, somewhat
shining, brown beneath when dry, the terminal ones up to 11
em long and 5.5 cm wide, the lateral ones somewhat smaller,
the base acute, the apex rather abruptly and slenderly subcau-
date-acuminate, the acumen 1 cm long or less. Spikes axillary,
solitary, 2.5 cm long or less, the bracteoles small, oblong-ovate,
acute, 1 mm long or less. Calyx 2.5 mm long, 3-lobed, the lobes
ovate, obtuse or subacute, about 1.5 mm long, pubescent exter-
nally. Petals 3, oblong, obtuse, glabrous, about 4 mm long, 2
mm wide. Stamens 6; filaments somewhat dilated below, united
for the lower 0.5 mm, free from the disk; anthers about 1 mm
long. Fruit unknown.

MINDANAO, Butuan Subprovince, Butuan, J. F.. Quadras 172, from the
herbarium of the Ateneo de Manila, locally known as sauangan.

Named in honor of Father F. Sanchez, for many years connected with

the Museum of the Ateneo de Manila. A species well characterized by
its trifoliolate leaves and short spikes.

GARUGA Roxb.

GARUGA LITTORALIS sp. nov.

Arbor 15 ad 20 m alta, inflorescentiis plus minusve dense
cinereo-puberulis exceptis glabra vel subglabra; foliis 30 ad 45
cm longis, 9- vel 10-jugatis; foliolis oblongis, ovato-oblongis vel
lanceolato-oblongis, acuminatis, basi late rotundatis, leviter cor-
datis, valde inaequilateralibus, integris, rariter distanter irreg-
ulariter leviter crenulatis; paniculis terminalibus, in anthesis

98 The Philippine Journal of Science 1916

circiter 8 cm longis, dense cinereo-puberulis; pedicellis quam ala-
bastra ovoidea brevioribus; petalis quam laciniae calyciniae
duplo longioribus.

A tree 15 to 20 m high, nearly or quite glabrous except the
rather densely cinereous-puberulent inflorescence. Branches
reddish-brown, glabrous, lenticellate, much thickened, 1 to 1.5
cm in diameter. Leaves alternate, crowded at the apices of the
branchlets, 30 to 45 cm long, deciduous before anthesis, the rachis
often somewhat puberulent, becoming glabrous. Leaflets straight
or somewhat falcate, 9 or 10 on each side of the rachis, opposite or
nearly so, chartaceous or thinly coriaceous, oblong, ovate-oblong,
or lanceolate-oblong, glabrous and somewhat shining when
dry, of the same color on both surfaces, the lower ones usually
smaller than the median ones, 5 to 12 cm long, 2 to 4.5 cm wide,
the margins entire or rarely distantly and irregulary crenulate,
the apex rather prominently acuminate, the base usually broad,
rounded or sometimes subtruncate, usually somewhat cordate,
strongly inequilateral, never with an additional pinnule at the
base as in Garuga floribunda Dene. Panicles appearing before
the new leaves, terminal, usually many from the apices of the
branchlets, about 8 cm long, all parts densely cinereous-puberu-
lent. Flowers yellow, rather congested on the ultimate branch-
lets of the inflorescence, puberulent, their pedicels 1 to 1.5
mm long, shorter than the ovoid buds. Calyx 3 mm long, the
lobes oblong-ovate, acute, 1.5 mm long. Petals puberulent,
oblong-oblanceolate or narrowly oblong, usually acute, reflexed,
about twice as long as the calyx-lobes. Filaments 2.5 mm long,
pubescent, much thickened below; anthers 1 mm long. Ovary
ovoid, sessile, pubescent, 2 mm long, 4-celled, each cell 2-ovuled,
style stout, puberulent, 2.5 mm long; stigma obscurely 4-lobed.
Fruit black when ripe, obovoid, much wrinkled when dry, gla-
brous, 8 to 10 mm in diameter, each containing one or two sub-
globose, prominently rugose seeds about 5 mm in diameter.

LuZON, Province of Tayabas, For. Bur. 12342 Curran (type), April,
1908, near the seashore. MASBATE, For. Bur. 12572, 12587 Rosenbluth,
January, 1909, both specimens sterile. CEBU, Talisay, For. Bur. 6481

Espinosa, September, 1906. Minporo, For. Bur. 8705 Merritt, January,
1908.

A tree, usually growing near the seashore, apparently rather widely
distributed in the Philippines. It is manifestly allied to Garuga flori-
bunda Dene., of Timor and northern Australia, but appears to be distin-
guished from that species by its entire, much larger, leaflets, absence of
the supplementary reduced pinnules at the base of the leaflets, and its
short-pedicelled flowers. The inflorescence is not strictly terminal, but
the peduncles are in the crowded axils of the fallen leaves at the apices

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 29

of the branchlets. The leaves appear after the flowers, and with the
growth of the branchlet the panicles in fruiting stage become lateral,
being situated in the axils of fallen leaves below the terminal crown of
leaves.

The species is known in Tayabas as libas (Tagalog); in Masbate and
Cebu as bogo (Visayan), and in Mindoro as abilo (Tagalog), the last
name more usually applied to Garuga abilo (Blanco) Merr.

Var. PAUCIJUGA var. nov.
A typo differt foliis 5- vel 6-jugatis.

CAGAYAN DE SULU, Merrill 5301, October 15, 1906.

The specimen is in fruit, but in all essential characteris it appears to
be like the species, differing only in its fewer leaflets. A specimen from
Balabac Island, with flowers only, no leaves, may be referable here, Bur.
Sci. $98 Mangubat, March, 1906, locally known as baro.

GARUGA CLARKII sp. nov.

Arbor circiter 30 m alta, subglabra; foliis alternis, usque ad
40 cm longis, imparipinnatis, 11- vel 12-jugatis, rhachibus plus
minusve puberulis; foliolis inaequilateralibus, oblongo-lanceo-
latis, usque ad 10 cm longis, 3 cm latis distincte acuminatis,
basi oblique rotundatis, margine distincte regulariterque crenato-
dentatis; floribus ignotis; paniculis sub fructu axillaribus,
pedunculatis, leviter puberulis, circiter 16 cm longis; fructibus
obovoideis, nigris, circiter 1 cm diametro.

A tree about 30 m high, subglabrous, or somewhat gray-puber-
ulent. Branches reddish-brown, glabrous, about 5 mm in dia-
meter. Leaves crowded at the apices of the branchlets, 35 to
40 cm long, the rachis usually black when dry, somewhat puber-
ulent. Leaflets opposite or subopposite, lanceolate or oblong-
lanceolate, chartaceous, straight or slightly falcate, the median
ones 8 to 10 cm long, 2.5 to 3 cm wide, the upper and lower ones
somewhat smaller, the base strongly inequilateral, rounded on
one side of the midrib, acute on the other, the apex prominently
acuminate, the acumen about 1 cm long, blunt, the margins
distinctly and regularly crenate-dentate, the upper surface quite
glabrous, shining, the lower very slightly paler, glabrous or
with few soft hairs; nerves about 13 on each side of the midrib,
distinct; petiolules about 2 mm long, usually puberulent.
Flowers unknown. Fruiting panicles axillary, peduncled, about
16 cm long, gray-puberulent. Fruits black and wrinkled when
dry, obovoid, glabrous, 1 cm in diameter or less, each containing
one or two rugose, subglobose seeds.

MASBATE, For. Bur. 1700 Clark, July 23, 1904, in forests at an altitude

of about 150 m, on moderate slopes, locally known as bugo (Visayan), and
said to yield a valuable timber.

30 The Philippine Journal of Science 1915

This species is manifestly allied to the preceding one, but differs in its
longer leaves and more numerous leaflets which are regularly crenate-
dentate and although rounded, are never cordate at the base.

PROTIUM Burman

PROTIUM CONNARIFOLIUM (Perk.) comb. nov.
Canarium connarifolium Perk. Frag. Fl. Philip. (1904) 92.

This species, of which Dr. Perkins had no fruiting material, and of
which she described only the male flowers is shown to be a representative
of the genus Protiwm by additional material. It is not closely allied to
Protium javanicum Burm., from which it differs in vegetative characters
and its very short inflorescence. The fruits are apparently subglobose,
when mature about 1.5 cm in diameter, ultimately 2- or 3-valved, and with
1 to 3 pyrenes, the pyrenes hard, about 8 mm long. It is further distin-
guished from Canarium by its 4-merous flowers. The species is repre-
sented by the following material, all from Palawan: Merrill 787 (cotype),
Bur. Sci. 758, 754, 787, 866 Foxworthy, For. Bur. 21604 Agama, from
Iwahig, Alphonso XIII, and Mount Victoria, flowering February to May.

The genus is new to the Philippines.

SANTIRIA Blume

SANTIRIA CAUDATA sp. nov. § Fusantiria.

Arbor glaberrima circiter 15 m alta; foliis 25 ad 35 cm longis,
foliolis 7, chartaceis vel subcoriaceis, in siccitate pallidis, nitidis,
oblongo-ellipticis ad oblongo-lanceolatis, usque ad 18 cm longis,
apice tenuiter caudato-acuminatis, basi acutis, nervis utrinque
12 ad 14, subtus prominentibus; paniculis axillaribus, 10 ad 15
cm longis, e basi ramosis; floribus 3 mm diametro.

A tree about 15 m high, entirely glabrous. Branches terete,
light-gray, somewhat striate when dry. Leaves 25 to 35 cm
long, the leaflets 7, oblong-elliptic to oblong-lanceolate, 12 to 18
cm long, 3.5 to 6 cm wide, chartaceous or subcoriaceous, pale
and shining when dry, entire, the apex slenderly caudate-acumi-
nate, acumen blunt, 1.5 to 2.5 cm long, the base acute, equilateral
or nearly so; lateral nerves 12 to 14 on each side of the midrib,
prominent, anastomosing, the reticulations lax, rather distinct;
petiolules 1 to 2 cm long. Panicles axillary, pyramidal, up to
15 cm long, branched at or from near the base, the branches
few, spreading, the lower ones 5 to 7 cm long. Flowers about
3 mm in diameter, greenish-yellow, their pedicels 2 to 3 mm
long. Calyx shallow, 2.5 mm in diameter, the lobes rounded,
about 1 mm long, 2 to 2.56 mm wide. Petals 3, broadly ovate.
2.5 mm wide and about 2.2 mm long, acute or obtuse, glabrous.
Disk annular, thick, glabrous, obscurely undulate; stamens 6,
the filaments broadened below, 0.5 mm long, attached to the outer
part of the disk; anthers 0.5 mm long. Ovary ovoid, glabrous,

Xe

a a ee SS
ey Z

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 81

1.5 mm in diameter; style very short, stout; stigma depressed-
globose or subdisciform.

Luzon, Province of Camarines, Tamban River, For. Bur. 21211 (type),
21217 Alvarez, April 1, 1914, forested slopes along streams, altitude 20
to 100 meters.

A species apparently belonging in the group with Santiria beccarii
Engl., but well characterized by being entirely glabrous. It is probably
most closely allied to the Philippine S. glabra Merr., from which it is
readily distinguished by its larger, more numerously nerved, caudate-
acuminate leaves and its longer panicles.

SANTIRIA LAGUNENSIS sp. nov. § Husantiria ?

Arbor alta partibus junioribus inflorescentiisque leviter fur-
furaceis exceptis glabra; foliis usque ad 20 cm longis, foliolis
5 vel 6, oblongo-ovatis, integris, in siccitate pallidis, nitidis,
usque ad 11 cm longis, subcoriaceis, acuminatis, basi acutis ad
rotundatis, nervis utrinque circiter 8; paniculis axillaribus, usque
ad 7 cm longis, leviter ferrugineo-furfuraceis, e basi ramosis.

A tree, apparently tall, glabrous except the rather densely
ferruginous-furfuraceous tips of the branchlets and the slightly
furfuraceous inflorescence. Branches pale-gray, terete, some-
what striate, glabrous, the tips somewhat brownish. Leaves
about 20 cm long, the leaflets usually 6, sometimes 5, oblong-
ovate, subcoriaceous, pale and shining when dry, 8 to 11 cm
long, 2.5 to 5 cm wide, entire, apex narrowed, rather prominently
acuminate, base acute to rounded; lateral nerves about 8 on each
side of the midrib, slender but distinct, anastomosing, the reti-
culations lax; petiolules 7 to 10 mm long. Panicles axillary,
about 7 cm long, branched at or from near the base, the branches
few, the lower ones up to 2 cm long, the younger parts distinctly
but minutely ferruginous-furfuraceous. Flowers 3-merous, the
young buds globose, rather numerous. Young fruits globose,
black when dry, 8 mm in diameter, the stigma lateral.

LUZON, Province of Laguna, San Antonio, Bur. Sci. 20505 Ramos, Feb-
ruary, 1913, forested slopes.

Somewhat resembling Santiria nitida Merr., but the inflorescence entirely
different. The species is characterized by its minutely furfuraceous, more

or less ferruginous indumentum on the growing tips of the branchlets
and on the inflorescence, otherwise quite glabrous.

SANTIRIA SAMARENSIS sp. nov. § Eusantiria.

Arbor 20 ad 30 m alta ramulis inflorescentiisque exceptis
glabra; ramulis crassis, in siccitate brunneis, partibus junioribus
plus minusve breviter ferrugineo-pubescentibus; foliis usque ad
80 cm longis, foliolis 9 vel 11, coriaceis, nitidis, oblongis ad
oblongo-ellipticis, usque ad 20 cm longis, basi acutis ad subro-

39 The Philippine Journal of Science 1915

tundatis, leviter inaequilateralibus, spice caudato-acuminatis,
nervis utrinque 12 ad 15, subtus prominentibus; paniculis axil-
laribus, 30 ad 40 cm longis, amplis, multifloris, ab infima tertia
parte ramosis; floribus pubescentibus, circiter 6 mm longis.

A tree 20 to 30 m high, glabrous except the branchlets and
infiorescence. Branchlets thickened, reddish-brown or brown
when dry, about 1 cm in diameter, terete, lenticellate, the
younger parts more or less ferruginous-pubescent with short
hairs. Leaves alternate, up to 80 cm in length, the rachis and
petioles reddish-brown when dry; leaflets 9 or 11, coriaceous,
rather pale when dry, somewhat shining, oblong to oblong-
elliptic, entire, 12 to 20 cm long, 5 to 9 cm wide, the base acute
to subrounded, somewhat inequilateral, the apex slenderly and
rather abruptly caudate-acuminate, the acumen up to 2.5 cm
long, blunt; lateral nerves 12 to 15 on each side of the midrib,
very prominent on the lower surface, curved-anastomosing, the
reticulations distinct; petiolules 2 to 3 cm long, reddish-brown
when dry. Panicles axillary, solitary, many on each branchlet,
30 to 40 cm long, the lower one-third without branches, the
primary branches spreading or ascending, the lower ones up to
10 cm in length, sparingly pubescent, reddish-brown, the ulti-
mate branchlets and pedicels rather densely pale-ferrugineous
pubescent with short hairs. Flowers yellowish, about 6 mm
long, the pedicels 3 to 4 mm long, the bracts lanceolate, acu-
minate, deciduous, about 3 mm long. Male flowers: Calyx some-
what cup-shaped, pubescent, about 3 mm long and wide, with
3 broadly triangular-ovate, acute teeth about 1 mm long. Petals
3, free, pubescent on both surfaces, coriaceous, broadly ovate,
obtuse, 4 to 4.5 mm long. Disk prominent, thickened, annular,
glabrous, about 3 mm in diameter. Stamens 6; filaments at-
tached to the outer margin of the disk, flattened below, narrowed
above, about 1.5 mm long; anthers oblong, dorsifixed, 1 to 1.2
mm long. Rudimentary ovary very small, glabrous. Female
flowers in general similar to the males. Staminodes 1.5 mm
long. Ovary ovoid, glabrous, 3-celled; stigma capitate, sessile,
about 1 mm in diameter.

SAMAR, Mount Cauayan, Phil. Pl. 1641 Ramos (type), April, 1914,
distributed sub Canarium. Luzon, Province of Camarines, Mount Balu,
For. Bur. 21136 Miranda, April, 1914; Tarangtong, For. Bur. 21098 Abe-
llanosa, March 31, 1914, in forests, altitude about 160 meters, Bicol name
palaspas.

A species belonging in the same group with Santiria laevigata Bl. and
S. maingayi Benn., characterized among the few Philippine forms by its

large leaflets which are prominently caudate-acuminate, and by its elon-
gated, many flowered panicles.

oo

x.c,1 Merrill: New or Noteworthy Philippine Plants, XI 33

ANACARDIACEAE
DRACONTOMELUM Blume

DRACONTOMELUM EDULE sp. nov.

Arbor alta, ramis incrassatis, ramulis junioribus dense fer-
rugineo-puberulis; foliis circiter 40 cm longis, rhachibus pube-
rulis, foliolis alternis subtus, axillis barbatis exceptis, glabris,
chartaceis oblongis, nitidis, acuminatis, 12 ad 17 cm longis, ner-
vis utrinque circiter 12, prominentibus; fructibus globosis, car-
nosis, edulis, glabris, circiter 2.5 cm diametro, putamine depres-
so, 1.8 cm diametro.

A tree reaching a height of 20 m, similar and closely allied
to Dracontomelum cumingianum Baill., differing especially in its
glabrous leaflets. Branches thickened, terete, about 1 cm in
diameter, grayish-brown, marked with large petiolar scars, the
tips densely ferruginous-puberulent. Leaves crowded at the
apices of the branchlets, about 40 cm long, the petiole and rachis
somewhat cinereous-puberulent, becoming glabrous; leaflets
about 11, chartaceous, oblong, entire, rather sharply acuminate,
base acute to rounded, somewhat inequilateral, mostly 12 to 17
em long, 4 to 5.5 em wide, shining on both surfaces, the lower
slightly paler than the upper, quite glabrous except for the
bearded glands in the axils on the lower surface; lateral nerves
about 12 on each side of the midrib, prominent, curved upward,
distant from the margin, anastomosing, the reticulations dis-
tinct; petiolules about 2 mm long. Panicles in the upper axils,
puberulent, flowers not seen. Fruit globose, green, turning
yellowish when ripe, about 2.5 cm in diameter, glabrous, smooth,
the mesocarp fleshy, acid, edible; stone depressed, about 1.8 cm
in diameter, 8 mm high, obscurely undulate-ridged.

PALAWAN, Taytay, Merrill 9391, May 14, 1913, in forests immediately
back of the mangrove and on steep slopes, ascending to at least 40 meters
altitude.

A species manifestly closely allied to Dracontomelum cumingianum Baill.,
from which it differs in its leaflets being quite glabrous except for the

bearded axils. It might be considered merely as a variety or form of
Baillon’s species by some authors.

BUCHANANIA Sprengel

BUCHANANIA PLATYPHYLLA sp. nov. § Sagittatae.

Arbor 10 ad 20 m alta, ramulis junioribus inflorescentiisque
parce adpresse ferrugineo-hirsutis; foliis coriaceis, oblongis,
nitidis, usque ad 35 cm longis, apice breviter, abrupte, obtuseque
acuminatis, basi cuneatis vel leviter decurrento-acuminatis,
nervis utrinque circiter 20; paniculis circiter 20 cm longis,

181281 3

34 The Philippine Journal of Science 1915

multifloris, floribus 5-meris; fructibus parcissime hirsutis vel
vetustioribus glabris, apiculo centrico.

A tree 10 to 20 m high, the growing tips of the branchlets
rather densely appressed ferruginous-hirsute. Branches terete,
brown, glabrous. Leaves oblong, coriaceous, somewhat shining,
20 to 35 cm long, 8 to 12 cm wide, pale or brownish-olivaceous
when dry, of about the same color on both surfaces, the apex
shortly, abruptly, and obtusely acuminate, base narrowed in the
lower one-fourth, cuneate or somewhat decurrent-acuminate;
primary nerves 17 to 22 on each side of the midrib, prominent,
the secondary nerves and reticulations netted, not parallel, dis-
tinct; petioles 1 to 2 cm long. Panicles numerous, in the upper
axils, usually about 20 cm long, many-flowered, all parts with
few, scattered, usually appressed, pale or ferruginous, hirsute
hairs, in fruit sometimes becoming nearly or quite glabrous.
Flowers 5-merous, white, the pedicels 1 mm long or less. Calyx-
lobes oblong, obtuse, about 1 mm long. Petals oblong, obtuse,
about 2.6 mm long, 1 mm wide, reflexed. Disk glabrous. Fila-
ments about 1.3 mm long, much narrowed at the apex; anthers
about 1 mm long, prominently sagittate. Carpels rather densely
pubescent. Mature fruits red, orbicular to broadly obovoid,
somewhat compressed, about 1 cm long, the apiculus central,
when immature with few, scattered, pale or ferruginous hairs,
in age glabrous or with very few hairs.

This species is widely distributed in the Philippines and is manifestly
allied to the common Buchanania arborescens Blume, differing notably in
its much larger, more numerously nerved leaves. The type is For. Bur.
20274 Miranda, from the Province of Misamis, Mindanao, March, 1913,
and I refer here also the following specimens:

Luzon, Province of Cagayan, For. Bur. 17154, 17808 Curran, For. Bur.
18459, 18468, 18481 Alvarez, For. Bur. 14767 Darling. MINDANAO, District
of Zamboanga, For. Bur. 21894 Villamil, Williams 2863, 2879: District of
Cotabato, For. Bur. 11772 Whitford. BAsILaN, Bur. Sci. 16356 Reillo, For.
Bur. 17896 Rafael, For. Bur. 8983 Hutchinson: District of Lanao, For.
Bur. 22082 Sherfesee, Cenabre, & Ponce. It is known in Cagayan Prov-
ince, Luzon, as paleng, arangas, sambrit, and pappagan; in Mindanao as
manguian; and in Basilan as lingabunu.

BUCHANANIA ACUMINATISSIMA sp. nov. § Sagittatae.

Arbor glabra circiter 10 m alta; foliis coriaceis, nitidis, in sic-
citate brunneis, oblongo-oblanceolatis, tenuiter acuminatis, usque
ad 17 cm longis, longe petiolatis, basi longe anguste decurrento-
acuminatis, nervis utrinque 20 ad 25; paniculis folia subae-
quantibus, glabris; floribus 4-meris, longe pedicellatis, pedicellis
articulatis; carpellis glabris; antheris sagittatis.

A glabrous tree about 10 m high, or the very tips of the

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 35

growing branchlets somewhat pubescent. Branches terete, dark
reddish-brown. Leaves brown when dry, shining, coriaceous,
oblong-oblanceolate 15 to 17 cm long, 3 to 4 cm wide, the apex
slenderly subcaudate acuminate, the acumen up to 2 cm long,
blunt, the base narrowed, long and slenderly decurrent-acumi-
nate; lateral nerves 20 to 25 on each side of the midrib, slender,
distinct on both surfaces, the secondary veins not parallel, re-
ticulations distinct; petioles 3 to 3.5 cm long. Panicles quite
glabrous, slender, rather few-flowered, about as long as the leaves,
branched from the middle and above. Flowers white, 4-merous,
their pedicels somewhat 4-angled, distinctly jointed, gradually
thickened upward, 2 to 3 mm long. Calyx lobes 0.5 mm long
or less. Petals oblong, obtuse, 2.5 mm long, about 1 mm wide.
Filaments 1 mm long, the apical part very slender; anthers
sagittate, about 1 mm long. Disk and carpels glabrous.

LuZON, Province of Laguna, San Antonio, Bur. Sci. 10932 Ramos,
August, 1910, in forests, Dahican River.

A species similar to Buchanania sessilifolia Blume (B. acuminata
Turcz.), from which it differs in its much more numerous lateral nerves,
and entirely glabrous leaves and panicles. The material was distributed

as Buchanania arborescens Blume, but it is quite different from that
species and not closely allied to it.

BUCHANANIA RETICULATA Elm. Leafi. Philip. Bot. 4 (1912) 1499.
This is typical Mangifera altissima Blanco, and must be reduced to
Blanco’s species. The type is Elmer 12834 from the Island of Sibuyan.
PRUNUS ? LAURIFOLIA Dene. in Nouv. Ann. Mus. Paris 3 (1834)
458; Miq. Fl. Ind. Bat. 1° (1855) 866=Buchanania arborescens Blume.

This species was considered by Miquel as a true Prunus, but he mani-
festly saw no specimens. A cotype of the species, received from the
Museum of Natural History, Paris, is in the Herbarium of the Bureau
of Science. The specimen is with fruits, and is typical Buchanania arbo-
rescens Blume.

SWINTONIA Griffith

SWINTONIA ACUMINATA sp. nov.

Arbor glabra, 15 ad 20 m alta; foliis coriaceis vel subcoriaceis,
usque ad 15 cm longis, oblongo-ellipticis, utrinque subaequaliter
angustatis, apice acuminatis, basi leviter decurrentibus, subtus
pallidis, nervis utrinque circiter 15; paniculis folia aequantibus
vel longioribus, multifloris, pedicellis glabris vel minute pubes-
centibus, petalis intus puberulis, sub anthesin circiter 2.5 mm
longis, obtusis, elliptico-ovatis.

A tree 15 to 20 m high, entirely glabrous except the short
pedicels and slightly puberulent petals. Branches terete, dark
reddish-brown when dry. Leaves alternate, coriaceous or sub-

36 The Philippine Journal of Science 1915

coriaceous, oblong-elliptic, 8 to 15 cm long, 3 to 6 em wide,
pale and shining when dry, the lower surface somewhat glaucous,
subequally narrowed to the acuminate apex and to the acute or
acuminate and very slightly decurrent base, the acumen up to
1 cm in length, blunt; lateral nerves about 15 on each side of
the midrib, mostly spreading, slightly curved, prominent, the
reticulations distinct on both surfaces; petioles slender, semi-

terete, not sulcate, reddish-brown when dry, 4 to 5 em long. .

Panicles axillary, slender, numerous, many flowered, up to 20
cm in length, the lower branches up to 12 cm in length. Flowers
cream-color or white, numerous, somewhat crowded on the ulti-
mate short branchlets, the pedicels 2 mm long or less. Calyx
campanulate, about 2 mm long, the lobes broad, rounded, im-
bricate. Petals in anthesis elliptic-ovate, obtuse, about 2.5 mm
long, glabrous outside, somewhat puberulent inside. Stamens
5. Ovary ovoid, glabrous, inequilateral, about 1 mm long; style
short; stigma capitate.

LuzON, Province of Camarines, Pinamuhagan, For. Bur. 21749 Fischer |

(type), April 16, 1914; same province and month of collection, For. Bur.
21729 Penas, Soriano, & Abellanosa, For. Bur. 21232 Alvarez. In forests,
altitude 10 to.80 meters.

Apparently allied to the Bornean Swintonia glauca Engl., but the
branchlets entirely glabrous, leaves not dark-colored on the upper surface
when dry, and with more prominent and more numerous lateral nerves,
relatively shorter and glabrous panicles, and glabrous ovaries. It is quite
’ different from the other known Philippine species.

SABIACEAE

MELIOSMA Blume
MELIOSMA ACUMINATISSIMA sp. nov.

Arbor circiter 5 m alta, glabra vel subglabra; foliis pinnatis,
usque ad 50 cm longis, foliolis subcoriaceis, usque ad 13, oblongo-
ovatis ad lanceolatis, tenuiter acute acuminatis, basi acutis,
margine integris, usque ad 11 cm longis, nitidis, nervis utrinque
6 vel 7; infructescentibus folia aequantibus, terminalibus, fruc-
tibus obovoideis, circiter 6 mm diametro, leviter compressis,
obscure reticulatis.

A tree about 5 m high, glabrous or nearly so. Branches terete,
pale brownish, the very tips ferruginous-hirsute, the plant (in
fruit) otherwise glabrous. Leaves up to 50 cm long, pinnate,
the leaflets about 13, oblong-ovate to lanceolate, entire, sub-
coriaceous, pale-olivaceous, shining, 7 to 11 em long, 3 to 4 em
wide, apex rather slenderly and sharply acuminate, base acute,
often a little inequilateral; lateral nerves about 6 on each side
of the midrib, prominent, curved-ascending, prominently anas-

pe a se, ee

~~

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 37

tomosing; petiolules about 4 mm long. Panicles (in fruit) ter-
minal, glabrous, as long as the leaves, the fruits nearly black
when dry, obovoid, about 6 mm in diameter, slightly compressed,
obscurely reticulate.

Luzon, Province of Tayabas, Mount Pular, Bur. Sci. 19417 Escritor,
January, 1913, in forests near the summit of the mountain.

A species well distinguished by being glabrous or nearly so, and by its
entire leaflets. It is probably most closely allied to Meliosma sylvatica
Elm., but that species is prominently pubescent and has denticulate
leaflets.

MELIOSMA MACGREGORII sp. nov.

Arbor erecta vel scandens (?) prominente ferrugineo-hirsutis;
foliis pinnatis, usque ad 40 cm longis, foliolis 9, oblongis ad
oblongo-oblanceolatis, coriaceis, usque ad 12 cm longis, acumi-
natis, basi acutis, nervis utrinque circiter 10, subtus valde prom-
inentibus, margine integris ad leviter distanter denticulatis;
paniculis terminalibus, amplis, multifloris, folia subaequantibus;
floribus racemose dispositis, circiter 3 mm longis.

An erect tree of small size (or scandent ?), the branches,
inflorescence, petioles and rachis, and the leaflets beneath and
on the midrib and lateral nerves on the upper surface prom-
inently ferruginous-hirsute. Leaves pinnate, up to 40 cm long,
the leaflets 9, coriaceous, oblong to oblong-oblanceolate, 7 to 12
cm long, 2.5 to 4 cm wide, the apex sharply acuminate, base
acute, margins entire to distantly and irregularly denticulate,
the upper surface grayish to brownish when dry, somewhat shin-
ing, the midrib, nerves, and primary reticulations more or less
impressed, the former two ferruginous-hirsute, the lower surface
brown, paler than the upper surface, prominently hirsute on the
midrib and lateral nerves, with few hairs on the reticulations;
lateral nerves about 10 on each side of the midrib, very prom-
inent, curved-anastomosing, the reticulations prominent; pet-
iolules 6 to 10 mm long and with the rachis and petioles
prominently ferruginous-hirsute. Panicles terminal, about as
long as the leaves, prominently ferruginous-hirsute, the primary
branches distant, spreading, up to 16 cm long, the lower second-
ary ones of each branch up to 7 cm long, many flowered, the
flowers white, racemosely disposed on the ultimate branchlets,
not crowded, their pedicels 1 mm long or less, the bracts and
bracteoles oblong, pubescent, 1 to 2 mm long. Sepals ovate to
elliptic-ovate, about 1.8 mm long, margins ciliate. Outer three
petals orbicular-ovate, rounded, 2.5 to 3 mm long, the inner
two reduced to a bifid membranaceous scale about 1 mm long,
cleft into two oblong lobes. Stamens about 1.8 mm long, the

38 The Philippine Journal of Science 1915 a

imperfect ones membranaceous, retuse, oblong, about 0.8 mm
long. Ovary ovoid, glabrous. Fruit ovoid, slightly compressed,
black, about 7 mm in diameter.

Luzon, Ifugao Subprovince, Mount Polis, Bur. Sci. 19749 McGregor,
February, 1913, in flower (type): Lepanto Subprovince, Mount Data,
Merrill 4600, November, 1905, in fruit, distributed as M. pendula.

A species manifestly allied to Meliosma pendula Merr., from which it
differs in its much narrower, very differently shaped leaflets.

MELIOSMA LOHERI sp. nov.

Arbor parva circiter 5 m alta partibus junioribus inflorescen-
tiisque dense ferrugineo-pubescentibus; foliis simplicibus, an-
guste oblongo-obovatis ad oblongo-oblanceolatis, coriaceis, usque
ad 20 cm longis, acuminatis, basi angustatis, cuneatis, margine
sursum irregulariter grosse serratis, nervis utrinque circiter 15,
subtus valde prominentibus; inflorescentiis terminalibus, pani-
culatis, folia subaequantibus, multifloris; floribus plus minusve
confertis, brevissime pedicellatis, circiter 2.5 mm longis.

A small tree, about 5 m high according to Vanoverbergh,
the younger parts and inflorescence densely ferruginous-pubes-
cent. Branches terete, dark-brown, pubescent, the younger
parts very densely so. Leaves simple, coriaceous, narrowly
oblong-obovate to oblong-oblanceolate, 10 to 20 cm long, 3 to 6

cm wide, apex prominently acuminate, acumen apiculate, base »

gradually narrowed, cuneate, margins in the lower part entire,
in the upper one-half usually with few, irregular, rather coarse,
sharp teeth, the upper surface olivaceous, glabrous, shining, the
ultimate reticulations distinct, minute, subfoveolate, the lower
surface more or less pubescent, somewhat paler than the upper,
the nerves and reticulations very distinct; lateral nerves about
15 on each side of the midrib, prominent, anastomosing, the
primary reticulations very prominent; petioles 2 to 3.5 cm long,
pubescent. Panicles terminal, about as long as the leaves, fer-
ruginous-pubescent, many-flowered, the flowers white, rather
crowded on the ultimate branchlets, their pedicels very short,
the bracts pubescent, oblong-lanceolate, acuminate, about 1.2 mm
long. Sepals ovate, rather thin, obtuse, about 1.2 mm long,
pubescent. Outer three petals orbicular-ovate, rounded, about
2.5 mm long, the inner two adnate to the filaments, 1 mm long or
less, cleft nearly to the base into two linear-oblong lobes. Fila-
ments about 1 mm long, the anthers 1 mm wide, one-half as long;
imperfect stamens strongly incurved, channeled, retuse, broadly
ovate. Ovary compressed, glabrous, ovoid, about 1 mm long.

Luzon, Bontoc Subprovince, Malamey, Vanoverbergh 1181 (type),
March 7, 1914, in forests, altitude about 1,600 meters; Ifugao Subprov-

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 39

ince, Mount Polis, Bur. Sci. 196830 McGregor, February, 1913. Here
probably should be referred Loher 268 (herb. Kew.) from Atoc, Benguet
Subprovince, a mere fragment of which is in the Herbarium of the Bureau
of Science.

Closely allied to Meliosma monophylla Merr. of central Luzon, but at
once distinguishable by its fewer nerved leaves, about 15 pairs in the
present species, about 20 pairs in M. monophylla.

MELIOSMA PAUCINERVIA sp. nov.

Arbor, partibus junioribus inflorescentiisque plus minusve fer-

rugineo-hirsutis; foliis pinnatis, usque ad 35 cm longis, foliolis
7 vel 8, subcoriaceis, oblongis, acute acuminatis, basi acutis ad
subrotundatis, usque ad 12 cm longis, nitidis, margine distanter
apiculato-serratis, nervis utrinque 4 vel 5, prominente anasto-
mosantibus; paniculis terminalibus, laxis, circiter 25 cm longis;
floribus circiter 3 mm longis; ovario hirsuto.
_ A tree, size not indicated, the younger parts and inflorescence
more or less ferruginous-hirsute. Branches reddish-brown when
dry, nearly glabrous, the younger ones sparingly hirsute. Leaves
pinnate, up to 35 cm in length, the rachis reddish-brown, sparing-
ly hirsute, leaflets 7 or 8, 8 to 12 cm long, 3 to 4.5 cm wide, sub-
coriaceous, oblong, sharply acuminate, base acute to somewhat
rounded, margins of the larger leaflets with from 4 to 6, distant,
small, apiculate, serrate teeth, the upper surface olivaceous when
dry, shining, glabrous or the midrib sparingly hirsute, the lower
surface paler, sparingly hirsute on the midrib and lateral nerves
which are brown in contrast to the paler surface; lateral nerves
4 or 5 on each side of the midrib, very prominent, curved,
prominently anastomosing, the reticulations lax; petiolules spar-
ingly hirsute, 5 mm long or less. Panicles terminal, ample, lax,
up to 25 cm in length, rather prominently ferruginous-hirsute,
the branches distant, the lower ones up to 16 cm long, spreading
or curved-ascending. Flowers white, racemosely and laxly dis-
posed on the ultimate branchlets, their pedicels hirsute, 1 to 2
mm long, the bracts oblong-lanceolate, acuminate, about 2 mm
long. Sepals pubescent, oblong to ovate, about 2 mm long.
Outer three petals orbicular-ovate, rounded, about 3 mm in
diameter, with a short ligulelike appendage near the apex inside;
two inner petals reduced to mere rudiments, each consisting
of two, narrowly linear lobes about 1.2 mm long, adnate to the
stamens. Fertile stamens 1.5 mm long, the imperfect ones broad,
incurved, oblong-ovate, about 1.5 mm long (spread) and 1 mm
wide. Ovary ovoid, hirsute.

MINDANAO, District of Zamboanga, Santa Maria, Bur. Sci. 16436 Reillo,
October, 1912.

cy
40 The Philippine Journal of Science 1915

A species well characterized by its distantly and sharply toothed leaflets,
few but prominent and prominently anastomosing lateral nerves, lax pani-
cles with laxly arranged flowers, and hirsute ovaries.

RHAMNACEAE
VENTILAGO Gaertner
VENTILAGO LANCEOLATA sp. nov.

Frutex scandens, partibus junioribus exceptis glaber; ramulis
tenuibus, teretibus; foliis lanceolatis, chartaceis, integris, usque
ad 8 cm longis, brevissime petiolatis, sursum angustatis, obtusis,
basi subacutis ad subrotundatis, nervis utrinque circiter 9, tenui-
bus; racemis axillaribus, tenuibus, sub fructu usque ad 6 cm
longis, solitariis; fructibus 4 ad 5 cm longis.

A scandent shrub, glabrous except the younger parts.
Branches slender, terete, the branchlets glabrous or slightly and
obscurely pubescent. Leaves lanceolate, chartaceous, 5 to 8 cm
long, 1 to 2.5 cm wide, rather pale when dry, shining on both
surfaces, entire, gradually narrowed upward to the obtuse apex,
the base subacute to subobtuse; lateral nerves about 9 on each
side of the midrib, very slender, curved-ascending, obscurely
anastomosing very close to the margins, the reticulations fine,
close, evident on both surfaces; petioles glabrous, 2 mm long
or less. Racemes axillary, solitary, slender, in fruit up to 6
cm long, slightly pubescent. Fruits, including the wing, 4 to 5
cm long, the basal part nearly 5 mm in diameter, quite glabrous,
the wing 7.5 to 10 mm wide, reticulate, somewhat rounded at
the apex, shortly apiculate.

BILIRAN, Bur. Sci. 18646 McGregor, June 14, 1914, in thickets near
the seashore.

A species characterized by its lanceolate, slenderly nerved leaves, and its
inflorescence reduced to a simple axillary raceme.

ZIZYPHUS Linnaeus
ZIZYPHUS OTANESII sp. nov.

Arbor circiter 5 m alta, ramulis pubescentibus, spinosis; foliis
parvis, suboblique ellipticis vel elliptico-ovatis, chartaceis, usque
ad 3 em longis, obtusis, 3-nerviis, subtus parce pubescentibus;
fructibus globosis, glabris, 1.5 cm diametro.

A tree about 5 m high, the branches terete, reddish-brown,
glabrous, the branchlets with very short sharp spines at the
nodes 1 to 2 mm in length, the younger branchlets ferruginous-
pubescent. Leaves small, somewhat inequilateral, chartaceous,
1.5 to 3 cm long, 1 to 2 em wide, obtuse, base rounded or sub-
acute on one side and rounded on the other, 3-nerved, the upper
surface olivaceous, glabrous or slightly pubescent on the nerves,

G
x.c,.1 Merrill: New or Noteworthy Philippine Plants, XI 41

the lower surface slightly paler, sparingly appressed-pubescent,
the hairs on the nerves somewhat ferruginous; petioles ferrugi-
nous-pubescent, 2to 3mm long. Fruits globose, hard, glabrous,
smooth, about 1.5 cm in diameter.

Luzon, Province of Pangasinan, Umingan, Bur. Sci. 17964 Otanes,
April 16, 1914, in forests east of the town, known to the Ilocanos as
dilap.

Manifestly allied to Zizyphus trinervia Poir., but at once distinguished
by its much smaller, but slightly pubescent leaves.

ELAEOCARPACEAE
ELAEOCARPUS Linnaeus

ELAEOCARPUS BATAANENSIS sp. nov. § Dicera.

Arbor circiter 12 m alta, novellis inflorescentiisque parcissime
pilosis exceptis glabra; foliis oblongo-ellipticis, chartaceis, usque
ad 9 cm longis, basi acutis, apice acuminatis, nitidis, subtus
eglandulosis, nervis utrinque circiter 6, tenuibus; racemis axi-
llaribus, 3 ad 4 cm longis, paucifloris; floribus 5-meris, sepalis
lanceolatis, acuminatis, circiter 5 mm longis, petalis laciniatis,
basi cuneatis, margine dense villosis; staminibus 25, antheris
apice ciliatis; fructibus 1-locellatis, ellipsoideis, circiter 2.5 cm
longis.

A tree about 12 m high, quite glabrous except the slightly ap-
pressed-pilose younger branchlets and inflorescence. Branches
slender, reddish-brown. Leaves oblong-elliptic, chartaceous, 6
to 9 cm long, 2 to 3.5 cm wide, subequally narrowed to the
acute base and the somewhat acuminate apex, margins serrate-
crenulate, somewhat olivaceous when dry, shining; lateral
nerves slender, about 6 on each side of the midrib, anasto-
mosing, the reticulations slender, axils of the leaves eglandular;
petioles slender, 1 to 1.5 cm long. Racemes. axillary, solitary,
few, 3 to 4 cm long, few-flowered, the pedicels about 5 mm long.
Flowers white, 5-merous. Sepals lanceolate, acuminate, about
5 mm long, 1.2 mm wide, externally very slightly puberulent.
Petals 5.5 to 6 mm long, base cuneate, margins in the lower
part densely villous with long hairs, the apical 2 mm cut into
about 15, very slender, subequal divisions. Stamens about 25,
3 mm long, the anthers narrow, scabrid, cleft at the apex, one
cell very slightly exceeding the other and ciliate-bearded at the
tip. Ovary pubescent; style slender, lower part somewhat pu-
bescent. Fruit ellipsoid or narrowly ellipsoid, about 2.5 cm
long, 1.5 cm in diameter, rounded at both ends, the endocarp
hard, bony, 1-celled.

Luzon, Province of Bataan, forests back of Limay, For. Bur. 21908
Medina, July 2, 1914, on forested slopes, altitude about 300 meters.

4? The Philippine Journal of Science 1915

This species is characterized by its few flowers, equally and slenderly
divided petals which are densely villous on the margins of the lower
one-half, and its eglandular leaves. It is an ally of EHlaeocarpus isotri-
chus F.-Vill., but has much smaller, quite different leaves.

ELAEOCARPUS OLIGANTHUS sp. nov. § Dicera.

Arbor parva, plus minusve pubescens; foliis oblongis, char-
taceis, usque ad 8 cm longis, basi acutis, apice acuminatis, mar-
gine leviter crenatis, dentibus tenuiter aristatis, nervis utrinque 5
vel 6; racemis, axillaribus, solitariis, paucifloris; floribus 5-meris,
5 mm longis, petalis obovatis, 4 laciniatis, laciniis 12, tenuibus,
intus dense hirsutis; staminibus 15; ovario 2-loculare, dense
villoso; fructibus 2.5 cm longis, vix 1 cm diametro, utrinque
angustatis, acutis.

A tree about 8 m high, the branchlets, petioles, lower surfaces
of the leaves on the midribs, and inflorescence rather densely
pubescent. Branches terete, reddish-brown, glabrous, smooth,
very slightly lenticellate. Leaves oblong to broadly oblong-lan-
ceolate, 5 to 8 cm long, 1.8 to 3 em wide, the base acute, the apex
acuminate, the margins crenulate, each tooth tipped with a
short, slender, awn or mucro, the upper surface shining, the
lower a little paler, chartaceous; lateral nerves 5 or 6 on each
side of the midrib, slender; petioles 3 to 4 mm long. Racemes
axillary, solitary, few-flowered, pubescent, 4 to 5 cm long.
Flowers 5-merous, their pedicels slender, about 8 mm long,
pubescent. Sepals lanceolate, acute, 5 mm long, 1.5 mm wide,
slightly pubescent externally, margins densely puberulent, inside
glabrous. Petals obovate in outline, 4 mm long, cut one-half to
the base into 12, slender, subequal, segments, the lower one-half
inside densely hirsute, margins hirsute, back glabrous except
near the base. Stamens 15, the anthers narrowly oblong, 2 mm
long, minutely scabrid, one cell slightly longer than the other
and shortly ciliate at the apex, both obtuse; filaments 0.56 mm
long or less. Ovary densely villous, 2-celled, narrowed upward
to the style, densely pale-villous; disk-glands prominent, pubes-
cent. Fruit narrowly oblong, about equally narrowed and acute
at both ends, smooth, about 2.5 mm long, 0.8 mm in diameter.

MINDANAO, District of Zamboanga, Siay River, For. Bur. 18398 Fox-
worthy, DeMesa, & Villamil, May 29, 1912, along the river, near sea level.

A species characterized by its pubescence, its rather small leaves, its
few-flowered racemes, and its petal-characters. It somewhat resembles
and probably is closely allied to Hlaeocarpus villosiusculus Warb., from
which, however, it differs in many characters.

*
x,¢C,1 Merrill: New or Noteworthy Philippine Plants, XI 48

ELAEOCARPUS RAMIFLORUS sp. nov. § Ganitrus.

Arbor alta partibus junioribus inflorescentiisque leviter pu-
berulis vel pubescentibus, vetustioribus glabris; foliis lanceolatis,
membranaceis vel chartaceis, usque ad 12 cm longis, nitidis,
leviter acuminatis, basi acutis, margine obscure crenulatis;
racemis numerosis, e axillis defoliatis, circiter 6 cm longis;
floribus 5-meris, 6 mm longis, ovario 5-loculare.

A tall tree, nearly glabrous except the younger parts and the
inflorescence. Branches terete, dark-colored when dry, gla-
brous, the growing parts slightly gray-puberulent. Leaves lan-
ceolate, membranaceous or somewhat chartaceous, 9 to 12 cm
long, 2 to 3.5 cm wide, subequally narrowed at both ends, the
upper surface shining and olivaceous when dry, the lower a
little paler and when young a little pubescent along the midrib,
becoming quite glabrous, smooth, the apex acuminate, base acute,
margins obscurely crenulate; lateral nerves about 12 on each
side of the midrib, slender, rather distinct, usually with glands
in the axils; petioles 1 to 1.5 cm long, somewhat appressed-
pubescent. Racemes numerous, about 6 cm long, from the
branches below the leaves in the axils of fallen leaves, one
raceme from each axil, distinctly pubescent with appressed
grayish hairs. Pedicels about 6 mm long, appressed-pubescent.
Sepals lanceolate, somewhat acuminate, 4.5 to 5 mm long,
externally sparingly appressed-pubescent with grayish hairs,
internally slightly puberulent, the margins rather densely so.
Petals 5 to 6 mm long, cut one-half way to the base into 9 to
11 slender segments, margins rather prominently pubescent in
the lower one-half, the back slightly pubescent in the median
part, otherwise glabrous. Stamens many; filaments very short,
hispid; anthers linear-oblong, about 2 mm long, minutely hispid,
one cell slightly longer than the other and with a tuft of few
short cilia at the apex. Ovary globose, villous, 5-celled; style
slightly pubescent, 2 mm long. Fruits not seen.

LuZON, Province of Laguna, San Antonio, trail between San Antonio
and Piape, in forests, Bur. Sci. 13115 Ramos, June, 1912.

One of the very few species of the section at present known from the
Philippines, characterized by its lanceolate shining leaves and its numer-

ous racemes which are borne on the ultimate branches but entirely below
the leaves.

ELAEOCARPUS CALOMALA (Blanco) comb. nov.

Vallea calomala Blanco FI. Filip. (1837) 439.
Ageracium (Ageratium) calomala Blanco 1. c. ed. 2 (1845) 306, ed. 3,
2: 204.

44 The Philippine Journal of Science 1918

Monocera isotricha Turcz. in Bull. Soc. Nat. Mose. 19° (1846) 494; Walp.
Ann. 1 (1848) 112.

Elaeocarpus lancaefolius F.-Vill. Novis. App. (1880) 31, non Roxb.

Elaeocarpus oblongus F.-Vill. 1. c.; Vid. Sinopsis Atlas (1883) ¢. 21, f.
A. (poor), Rev. Pl. Vasc. Filip. (1886) 72; Perk. Frag. Fl. Philip.
(1904) 100, p. p.; Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 90;
Aug. DC. in Elm. Leafl. Philip. Bot. 2 (1909) 636, non Gaertn.

Elaeocarpus isotrichus F.-Vill. 1. ¢.; Aug. DC. 1. ec.

Elaeocarpus philippinensis Warb. in Perk. Frag. Philip. (1904) 100;
Aug. DC. 1. c. 636.

This much named endemic species is of wide distribution in Luzon and
Mindoro, its oldest valid specific name being here adopted. Blanco’s descrip-
tion is quite good and agrees perfectly with the material cited below. It
is, moreover, the only form known to me from Luzon to which his descrip-
tion can be referred. His material was from the Province of Batangas,
and his specific name is from its Tagalog name calomala.

Elaeocarpus isotrichug (Turcz.) F.-Vill. has been considered by several
authors as a species of doubtful status. It was based on Cuming 886 in
Turczaninow’s herbarium, but this number in other herbaria is Mediniilla
cumingii Naud. Through the kindness of the Director of the Botanical
Institute of the University of Kharkoff I have been able to examine Turcza-
ninow’s type, and find it to be the common Luzon form, identical in all
respects with the species recently described by Warburg as Elaeocarpus
philippinensis; of the later I have examined the type in the Berlin Herba-
rium, an unnumbered specimen of Cuming’s collection.

Elaeocarpus oblongus Gaertn. apparently does not occur in the Philip-
pines, the numerous specimens so named having been determined as such
through error. The exact status of Gaertner’s species is doubtful, as it
was based primarily on Ganitrum oblongum Rumph. Herb. Amb. 3: 161,
t. 102, from which the specific name was taken. The only description
given by Gaertner is that of the fruit, taken from a specimen in the Leiden
Herbarium, its origin not indicated. Rumphius’s figure represents a plant
with entire leaves, a form entirely different from the Elaeocarpus oblongus
of all modern authors except Gaertner and DeCandolle.

LUZON, without definite locality, Cuming 836 (type of Monocera isotricha
Turcz.), Cuming s. n. (type of Elaeocarpus philippinensis Warb.): Sub-
province of Benguet, Elmer 6084: Province of Bataan, For. Bur. 2987
Borden, For. Bur. 2852 Meyer, For. Bur. 17602 Curran: Province of
Rizal, Phil. Pl. 1065 Ramos, Bur. Sci. 2665 Ramos, Merrill 1719, For.
Bur. 454, 2906, 3201 Ahern’s collector: Province of Tayabas, For. Bur.
6693 Kobbe. Muinporo, Merrill 1174, 2875, Whitford 1892, For. Bur. $727,
4059 Merritt. ;

THYMELAEACEAE

AQUILARIA Lamark

AQUILARIA MALACCENSIS Lam. Encycl. 1 (1783) 49; Gamble in Journ.
As. Soc. Beng. 75° (1912) 264.
Luzon, Province of Camarines, Salauigan, For. Bur. 21452 Alvarez,
May, 1914.
The identification was originally made from Gamble’s description, Alva-
rez’s specimens being in fruit, no flowers available. Through the kindness
of Mr. I. H. Burkill, director of the Botanic Garden, Singapore, I have

x,G,1 Merrill: New or Noteworthy Philippine Plants, XI 45

recently received a fruiting specimen of Lamark’s species from Malacca,
and consider the Philippine plant to be specifically identical with the
Malacca one. The genus is new to the Philippines.

BEGONIACEAE

BEGONIA Linnaeus
BEGONIA ALBA sp. nov. § Diploclinium.

Herba parce bruneo-setosa praesertim partibus junioribus;
foliis longe petiolatis, subreniformibus vel subrotundatis, inte-
gris vel leviter undulatis, basi profunde subaequaliter cordatis,
palmatim 7- vel 9-nerviis; floribus masculinis paucis, albis, cir-
citer 2.5 cm diametro, bracteis orbicularibus, deciduis; stami-
nibus circiter 20.

A succulent herb, the rhizome creeping, rather slender, gla-
brous or nearly so, the stipules persistent only at the growing tip,
sparingly brown-setose, ovate, acuminate, submembranaceous,
about 1.5 cm long. Petioles up to 30 cm in length, glabrous, or
when young with few, long, scattered, brown, setose hairs.
Leaves chartaceous when dry, pale-brownish, orbicular or sub-
reniform, up to 15 cm long and 17 cm wide, entire or the
margins slightly undulate, very broadly rounded, subequilateral,
the base deeply and subequally cordate, the sinus very narrow,
up to 4 cm deep, the basal lobes very broadly rounded, the upper
surface glabrous, the lower with long scattered, brown hairs
when young, becoming glabrous or nearly so, the margins dis-
tinctly brown-setose; nerves 7 or 9, radiating from the base,
prominent, forked, the lateral ones above the base 2, rarely 3
pairs; petioles up to 30 cm in length, brown-setose when young,
becoming glabrous. Scape rather slender, 20 cm long or less,
the staminate flowers few, subumbellately cymose at the apex of
the peduncle, the primary branches 2 or 3, 1 cm long or less.
Bracts prominent, orbicular or subreniform, membranaceous,
deciduous, white, about 6 mm in diameter, two subtending each
flower. Staminate flowers: Sepals 2, elliptic to elliptic-orbi-
cular, broadly rounded, about 12 mm long. Petals 2, oblong-
oblanceolate, about 10 mm long, narrowed below. Stamens
about 20; filaments 1 mm long; anthers narrowly obovoid, as
long as the filaments. Pistillate flowers and fruits not seen.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19963 McGregor,
February, 1913.

A very characteristic species, at once distinguishable by its nearly
equilateral, entire, deeply and subequally cordate leaves, and by its few,
comparatively large, white flowers. In vegetative characters it resembles

Begonia suborbiculata Merr., of Palawan, but is entirely different from
that species in its floral characters.

46 The Philippine Journal of Science 1915

BEGONIA BILIRANENSIS sp. nov. § Diploclinium.

Herba subtus foliis ad nervos petiolis pedunculisque brunneo-
setosis, caulibus prostratis; foliis confertis, longe petiolatis,
usque ad 22 ecm longis, oblique ovatis, acuminatis, valde inae-
quilateraliter cordatis, margine undulatis vel parce undulato-
lobatis; inflorescentiis folia subaequantibus, multifloris, floribus
masculinis circiter 11 mm diametro, sepalis orbiculari-ovatis,
quam petalis multo latioribus; staminibus circiter 60; capsulis
eirciter 1 cm longis, 1.5 cm latis, subaequaliter 3-alatis, alis
rotundatis.

Stems creeping, rather stout, more or less brown-setose and
with brown, ovate, acuminate stipules about 1 cm long, the
leaves crowded, the peduncles, leaves on the nerves beneath, and
petioles prominently brown-setose, the petioles 11 to 18 em long.
Leaf-blades obliquely ovate, strongly inequilaterally cordate,
submembranaceous when dry, 15 to 22 cm long, 10 to 15 cm
wide, acuminate, margins undulate or somewhat undulate-lobed,
the upper surface somewhat olivaceous when dry, slightly shin-
ing, minutely puncticulate, glabrous, the lower surface greenish-
or brownish-olivaceous, the nerves in strong contrast, brown,
brown-setose on the nerves and primary reticulations, every-
where white-puncticulate, the nerves about 10. Inflorescence
about as long as the leaves, long-peduncled, ample, with both
male and female flowers. Male flowers white or very pale-pink,
about 11 mm in diameter. Sepals orbicular-ovate, rounded,
about 5.5 mm long, 5 mm wide. Petals oblong-obovate, rounded,
narrowed below, as long as the sepals, about 2 mm wide. Sta-
mens about 60; anthers 0.6 mm long, obovoid, as long as the
filaments. Female flowers: Sepals 2, broadly elliptic-ovate,
rounded, about 6 mm long, 5 mm wide. Petals 2, as long as
the sepals, oblong-obovate, 2.5 mm wide. Styles 3, about 3 mm
long, forked above, the stigmas spiral. Capsules about 1 cm
long, 1.5 cm wide, including the wings, the wings 3, subequal,
rounded.

BILIRAN, Bur. Sci. 18822 (type), 18760, 18544 McGregor, June, 1914,
on rocks in forests, altitude 100 to 800 meters.

A species allied to Begonia mindorensis Merr., but with larger leaves,
smaller flowers, and decidedly larger fruits.

BEGONIA PLATYPHYLLA sp. nov. § Petermannia.

Herba erecta, ramosa, B. cumingii affinis, differt foliis latio-
ribus, usque ad 9 cm latis, minus pubescentibus, floribus mas-
culinis multo minoribus, petalis circiter 5 mm longis.

An erect branched herb, nearly glabrous. Leaves inequilat-
eral, broadly oblong to oblong-ovate, 11 to 18 cm long, 7 to 9

x%,¢.1 Merrill: New or Noteworthy Philippine Plants, XI ny

cm wide, chartaceous, the apex abruptly short-acuminate, the
base scarcely narrowed, one side much narrower then the other,
acute, the broader side very broadly rounded, forming a shallow,
wide sinus with the petiole, the margins irregularly and rather
coarsely toothed, not at all lobed, the primary teeth very broad,
short, acute, the shallow sinuses denticulate; upper surface glab-
rous, subolivaceous when dry, slightly shining, the lower surface
much paler, glabrous except for the slightly subfurfuraceous
nerves; nerves prominent, nearly straight, forked, the basal ones
usually 9, of which 2, sometimes 3, are on the narrower side of
the lamina, the lateral nerves above the base 2 on the narrower
side, 3 on the broader side, ascending, the reticulations lax, in-
distinct; petioles 3.5 to 5.5 cm long, when young somewhat
brown-furfuraceous; stipules 2 cm long. Staminate and pis-
tillate flowers in different inflorescences on the same plant. Sta-
minate inflorescence terminal, lax, divaricately branched, up to
10 cm long, few-flowered. Flowers pink or white. Sepals 2,
orbicular, about 5mm in diameter. Petalsnone. Stamens about
30; anthers ellipsoid, about 1 mm long, equaling the filaments.
Pistillate inflorescence axillary and terminal, few-flowered, 5 cm
long or less, each flower subtended by 2, deciduous, oblong-
elliptic, acute or obtuse, membranaceous bracts. Sepals 5, nar-
rowly elliptic, narrowed at both ends, acute, about 1.3 em long.
Mature capsules, including the wings, turbinate, about 2 cm long,
2.5 cm wide, subequally 3-winged, the apex truncate, the upper
outer corners of the wings rounded, the base broadly rounded.

LUZON, Province of Nueva Vizcaya, near Imugan, Bur. Sci. 20074
McGregor, January, 1912.

In comparison with Begonia cumingii A. Gray (B. philippinensis A.
DC.), to which the species is manifestly allied, the striking differences
are in the small staminate flowers and the relatively broad, nearly glabrous
leaves of the present species.

BEGONIA MEGALANTHA sp. nov. § Petermannia.

Species B. merrittii affinis, differt ramulis et subtus foliis ad
nervos prominente pubescentibus, foliis subintegris vel leviter
dentatis, vix lobatis, petiolo brevioribus, floribus majoribus, pe-
talis usque ad 2.5 cm longis.

An erect, branched shrub 1 to 2 m high, the stems very woody,
stout. Branches and branchlets more or less zig-zag, prom-
inently pubescent with short, more or less crisped, pale-brown
hairs. Leaves inequilateral, oblong, chartaceous, 5 to 10 cm long,
1.8 to 4 cm wide, the upper surface brownish or olivaceous,
slightly shining, the lower surface paler, densely puncticulate,
glabrous except the midrib and nerves which are prominently

48 The Philippine Journal of Science 1915

brown-pubescent, the hairs similar to those on the branchlets, the
apex slenderly acuminate, somewhat falcate, the base not at all or
but slightly narrowed on the broadly rounded wider side, slightly
narrowed and acute or somewhat cordate on the narrower side,
the margins distantly and slightly toothed, not at all lobed, the
base 6 or 7-nerved, the primary lateral nerves above the base
2 on the narrower side, usually 3 on the broader side, ascending,
forked, prominent on the lower surface, the reticulations lax,
indistinct; petioles brown-pubescent, 2 to 4 mm long; stipules
oblong, submembranaceous, about 7 mm long, their midribs
excurrent as long and slender apiculi. Staminate and pis-
tillate flowers on separate inflorescences on the same plant, in
rather lax, few-flowered, terminal cymes, sometimes reduced to
two or three, long-pedicelled flowers. Staminate flowers: Sepals
2, pure white to somewhat pink, broadly ovate, membranaceous,
glabrous, 2 to 2.5 cm long, 2 to 2.8 em wide, the base broadly
cordate, the apex broadly rounded. Petals none. Stamens about
50, inserted on the slightly elongated torus, the filaments 2.5
mm long, the anthers narrowly oblong-obovoid, about 1.5 mm
long. Bract’y membranaceous, apiculate, about 7 mm long.
Pistillate flowers: Bracts elliptic-ovate, membranaceous, 1 cm
long, apiculate, two subtending each flower, deciduous; pedicels
slender, pubescent, about 2.5 cm long. Sepals 5, narrowly ovate,
membranaceous, acute or obtuse, about 2.3 cm long. Stigmas
spirally arranged. Capsules turbinate, including the wings
about 1.5 cm long and wide, subequally 3-winged, the apex trun-
cate or somewhat cordate, the tips of the wings broadly rounded,

the base acute or subtruncate, broadly rounded. ‘

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19857, 19858,
19859, 19860 (type) McGregor, February, 1913.

A striking species, manifestly closely allied to Begonia merrittii Merr.,
from which it is distinguished by the numerous characters indicated in
the diagnosis. It is remarkable for its large size, the development of
woody tissue in the stems, and its very large flowers.

BEGONIA LANCIFOLIA sp. nov. § Petermannia.

Erecta vel suberecta, ramosa, usque ad 60 cm alta, glabra,
ramis ramulisque tenuibus; foliis lanceolatis, in siccitate mem-
branaceis, viridis, usque ad 12 cm longis, deorsum angustatis, basi
acutis vel obtusis, subaequilateralibus, apice longissime caudato-
acuminatis, margine grosse irregulariter sinuato-lobatis ; inflores-
centiis terminalibus, paniculatis, tenuibus; floribus ¢ parvis,
sepalis 2, petalis 0; capsulis 8 mm longis, aequaliter 3-alatis.

An erect or suberect branched glabrous herb 60 cm high or

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 49

more, the stems and branches slender, pale-brown when dry,
sometimes geniculate and emitting small roots at the nodes.
Leaves lanceolate, when dry membranaceous, green, of nearly
the same color on both surfaces, glabrous, 8 to 12 cm long, 1.5
to 2.5 cm wide, narrowed below to the acute or obtuse, nearly
equal or slightly inequilateral base, not at all cordate, the apex
very long and slenderly caudato-acuminate, the acumen sharply
toothed, the margins in the basal part entire, in the median por-
tions coarsely and irregularly sinuate-lobed, about 3 lobes on
each side of the midrib, the lobes acute, broad, the sinuses mostly
rather shallow, rounded; stipules lanceolate, long and slenderly
aristate-acuminate, about 10 mm long, 2 mm wide at the base;
petioles slender, 1 to 2 cm long. Inflorescence terminal, 9 cm
long or less, the branches and branchlets rather few, very slender,
one or more female flowers at the base, the male flowers above,
apparently rather numerous but early deciduous. Male flowers:
Sepals 2, orbicular, broadly rounded, in nearly mature bud 4
mm in diameter. Petals none. Stamens about 25, the anthers
oblong-obovoid, nearly 1 mm long. Capsules about 8 mm long,
including the 3 subequal wings about 1.4 cm wide, the base trun-
cate, the apex broadly rounded or subtruncate, the wings thin,
rounded.

BASILAN, Bur. Sci. 16162 Reillo, August 28, 1912, in forests, Cuma-
larang River.

Most closely allied to Begonia littleri Merr., also known only from Basi-
lan, but distinguished by its entirely different leaves, and its equally
_ 3-winged eapsules. From B. longistipula Merr. and B. palawanensis it
not only differs in its much narrower, differently shaped leaves, but also
in its very much smaller capsules.

BEGONIA LACERA sp. nov. § Petermannia.

Species praecedente affinis, differt foliis brevioribus angus-
tioribusque, usque ad 7 cm longis, 1 cm latis, irregulariter lace-
rato-lobatis, stipulis brevioribus, circiter 6 mm longis, sepalis
(fl. ¢) late ovatis, subacutis.

An erect, branched, slender herb about 40 cm high, quite
glabrous, the branches and branchlets reddish-brown when dry,
slender. Leaves lanceolate, thinly membranaceous when dry,
green, 3 to 7 cm long, about 1 cm wide, narrowed to the sub-
equilateral acute or obtuse base, the apex very long and slenderly
caudate-acuminate, the margins irregularly lacerate-lobed in the
median portion, the basal portion entire, the long and slender
acumen sharply and rather coarsely toothed, the lobes few,
rather narrow, acute or acuminate; petioles 5 mm long or less;

131281——-4

50 The Philippine Journal of Science 1915

stipules lanceolate, 6 mm long, 1.8 mm wide below, long and
slenderly aristate-acuminate. Inflorescence paniculate, terminal
or in the uppermost axil, lax, 4 cm long or less, the branches
few, slender. Flowers few, the staminate ones with 2, broadly
ovate, subacute or obtuse, not rounded, sepals 3 mm in length.
Petals none. Stamens about 20. Anthers broadly obovoid,
rounded, 0.6 mm long. Female flowers apparently solitary at
the base of the staminate panicle, the sepals lanceolate, acumi-
nate or acute, 4 mm long, 1.2 to 1.5 mm wide. Stigmas 3, about
1 mm long, spirally twisted. Capsules not seen, but from the
female flowers (in anthesis) apparently equally 3-winged.

BASILAN, Cumalarang River, in forests, Bur. Sci. 16111 Reillo, August
30, 1912.

A rather striking species, distinguishable by its lanceolate, very slen-
derly acuminate, more or less lacerate-lobed leaves which are narrowed
to the subequilateral acute or obtuse bases. It is manifestly allied to
Begonia lancifolia Merr., but can be readily distinguished by its smaller,
more narrowly lobed leaves, shorter petioles and stipules, subacute or
obtuse sepals of the male flowers and fewer shorter anthers.

BEGONIA OLIGANTHA sp. nov. § Petermannia.

Herba erecta, parce ramosa, glabra, circiter 30 cm alta; foliis
oblongis, vel anguste oblongo-obovatis, membranaceis, usque ad
7 cm longis, sinuato-dentatis, acuminatis, deorsum angustatis,
leviter inequilateralibus, basi minute oblique subauriculatis; in-
florescentiis terminalibus, paucifloris, floribus ¢ circiter 18 mm
diametro, sepalis 2, late ovatis, petalis 2, oblanceolatis; capsulis
1.3 cm longis, aequaliter 3-alatis, basi rotundatis, apice truncatis.

An erect herb, glabrous throughout, about 30 cm high, spar-
ingly branched, the stems and branches reddish-brown when
dry. Leaves membranaceous when dry, green, oblong to nar-
rowly oblong-obovate, 5 to 7 cm long, 1 to 2 cm wide, slightly
faleate, narrowed below to the slightly inequilateral, obtuse,
obliquely subauriculate base, and above to the slenderly acumi-
nate apex, the margins distantly and irregularly sinuate-
toothed; petioles 2 to 7 mm long; stipules narrowly lanceolate,
long and slenderly acuminate, about 12 mm long, 1.5 mm wide.
Inflorescence terminal or in the uppermost axil, paniculate, 5 to
7 cm long, the branches very few, few-flowered, slender. Male
flowers: Sepals 2, broadly ovate, 9 to 10 mm long, about 6 mm
wide, obtuse or narrowly rounded. Petals 2, oblanceolate, ob-
tuse, 8 mm long, 2 mm wide. Stamens about 20, inserted on
the elongated torus; anthers narrowly obovoid, rounded, base
acute, 1.5 mm long. Pistillate flowers apparently solitary at
the base of each panicle. Capsules 1.8 ecm long, 1 to 1.5 cm

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 51

wide, equally 3-winged, broadly rounded at the base, the apex
truncate, the upper outer angles sharp.

*' MINDANAO, Bukidnon Subprovince, Sumilao, Bur. Sci. 157383 Feénix,
August 4, 1912, on wet rocky slopes, locally known as tabaring.

A species with the vegetative characters of Begonia littleri Merr., in
general similar to that species, and to the allied forms B. brevipes Merr.,
B. longistipula Merr., and B. palawanensis Merr. It is at once distin-
guished by its large male flowers, which in this species are about 2
em in diameter, while in the above species they do not exeeed 1 cm
in diameter.

BEGONIA LATISTIPULA sp. nov. § Petermannia.

Herba erecta, ramosa, ramis ramulisque crassis, teretibus,
parce setoso-ciliatis; foliis usque ad 13 cm longis, valde inaequi-
lateralibus, oblongis vel ovato-oblongis, acuminatis, irregulariter
lobatis denticulatisque, lateraliter cordatis, basi haud angustatis;
stipulis 1.5 ad 2 em longis, ovatis, cilato-setosis; floribus paucis,
axillaribus, fasciculatis, ¢ 1 vel 2, longe pedicellatis, circiter 3
em diametro, ¢ solitariis, subsessilibus; capsulis aequaliter 3-
alatis, apice truncatis, basi acutis, circiter 1.5 cm longis, 2 cm
latis.

An erect, sparingly branched, rather stout herb at least 50
em in height, the stems and branches, stipules, petioles, and
young leaves on the lower surface more or less setose-ciliate.
Branches terete, reddish-brown when dry. Leaves thinly mem-
branaceous when dry, strongly inequilateral, oblong to ovate-
oblong, somewhat falcate, 9 to 138 cm long, 3 to 5 cm wide,
when young ciliate-setose with brown hairs on the nerves of
the lower surface, becoming glabrous or nearly so, the margins
irregularly lobed, the broader side with one or two rather coarse,
acute lobes and 5 or 6 mueh smaller ones more like coarse teeth,
irregularly ciliate-denticulate, the apex rather slenderly acu-
minate, the base scarcely narrowed, prominently laterally cord-
ate, the lobes rounded, that of the broader side very much wider
than that of the narrow side; petioles 8 to 15 mm long, rather
densely setose-ciliate with brown hairs; stipules ovate, some-
what inequilateral and slightly falcate, membranaceous, 1.5 to
2 cm long, 1 to 1.2 cm wide, prominently ciliate-setose on the
margins, the midrib excurrent as a short mucro. Flowers ax-
illary, fascicled, one male and one female, or two males and
one female in the same axil. Male flowers: Pedicels slender, up
to 2 cm long. Sepals 2, broadly ovate, rounded, 1.5 cm long.
Petals none. Stamens about 20, inserted on the elongated torus;
anthers narrowly ovoid, 2 mm long, rounded. Capsule subses-
sile or shortly pedicelled, triangular, the base acute, the apex

52 The Philippine Journal of Science 1915

truncate, the outer upper angles sharp, about 1.5 cm long and 2
em wide across the apex.

LEYTE, Dagami, Bur. Sci. 15367 Ramos, August, 1912, in forests near
the summit of Mount Buraui.

A characteristic species readily distinguishable by its broad stipules
and its rather large axillary fascicled flowers, two or three flowers in
a fascicle, one of which is pistillate and one or two staminate, the former
shortly pedicelled or subsessile, the latter long and slenderly pedicelled.
Its alliance seems to be with Begonia robinsonii Merr., which, however,
it does not greatly resemble.

ERICACEAE
DIPLYCOSIA Blume

DIPLYCOSIA FASCICULIFLORA sp. nov.

Species D. parvifoliae affinis, differt floribus fasciculatis, pedi-
cellis brevioribus, ramis ramulisque glabris, vix setosis.

Apparently scandent, probably epiphytic, the branches terete,
glabrous, dark reddish-brown, the branchlets paler, glabrous,
not at all setose. Leaves coriaceous, elliptic to elliptic-oblong,
dull, 2 to 4 cm long, 1 to 1.5 cm wide, when young with few,
long, slender, brown, setose hairs on the margins, and scattered
ones on both surfaces, when mature quite glabrous, the midrib
prominent, lateral nerves obsolete, base and apex acute; petioles
2 to 3 mm long, slightly setose. Flowers small, pedicelled, in
axillary fascicles, up to 8 in each fascicle, the pedicels slender,
glabrous, 5 mm long or less, the basal bracts small, the apical
bracteoles very broadly ovate, obscurely acuminate, puberulent.
Calyx glabrous, somewhat accrescent and 2.5 mm long, the lobes
triangular, acute, about 1 mm long. Corolla and stamens not
seen. '

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19758 McGregor,
February 1, 1913.

A species very similar and manifestly closely allied to Diplycosia parvi-
folia Merr., of Negros, strongly resembling that species in vegetative
characters, especially in the obsolete lateral nerves and reticulations. It
differs from that species, however, in its flowers being fascicled, not
solitary, its longer pedicels, and somewhat larger leaves.

VACCINIUM Linnaeus

VACCINIUM IRIGAENSE sp. nov.

Frutex glaber, circiter 5 m altus; foliis usque ad 6 cm longis,
coriaceis, oblongo-obovatis, petiolatis, utrinque angustatis, basi
acutis, tenuiter 5- vel 7-plinerviis, apice subrostrato-acuminatis;
racemis axillaribus, solitariis, usque ad 6 cm longis, multifloris;
floribus 1 cm longis, oblongis, sursum angustatis, ebracteolatis;
filamentis pilosis.

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 53

A glabrous shrub about 5 m high, the branches dark reddish-
brown, terete, shining, the branchlets brownish. Leaves coria-
ceous, rather narrowly oblong-obovate, 4.5 to 6 cm long, 1.5 to
2.5 em wide, shining, the lower surface paler than the upper,
narrowed below to the acute and slenderly 5- or 7-plinerved base,
and above to the somewhat rostrate-acuminate apex, the acumen
blunt, less than 1 cm long; reticulations slender, lax; petioles
5 mm long or less. Racemes axillary, solitary up to 6 cm long,
rather many flowered. Flowers red, 1 cm long, their pedicels
6 to 7 mm long, ebracteate. Calyx glabrous, about 1.5 mm long,
3 to 3.5 mm in diameter, the lobes broadly ovate, blunt, 1 mm
long or less. Corolla oblong, 3 mm in diameter below, narrowed
above and 1 mm in diameter under the orifice, glabrous, the
lobes slightly spreading, oblong, obtuse, 1 mm long. Stamens
10; filaments flattened below, somewhat pilose, 4 mm long;
anthers oblong, truncate, straight or slightly curved, 1.5 mm
long. Style stout, glabrous, 8 mm long.

Luzon, Province of Camarines, Mount Iriga, Phil. Pl. 1549 Ramos,
December 3, 1913, in the mossy forest.

A species manifestly allied to Vaccinium caudatum Warb., from which
it differs in its quite differently shaped leaves and larger flowers.

VACCINIUM MACGREGORII sp. nov.

Frutex erectus (vel scandens ?), inflorescentiis parce pubes-
centibus exceptis glaber; foliis magnis, coriaceis, usque ad 18
cm longis, petiolatis, basi acutis, apice subcaudato-acuminatis,
anguste ovatis ad oblongo-ovatis, basi prominente 7-plinerviis;
racemis paucifloris, pubescentibus, axillaribus, solitariis, usque
ad 9 cm longis, bracteolis oblongo-lanceolatis, acuminatis, 2 cm
longis, deciduis; corolla rubra, 1.6 cm longa, supra leviter am-
pliata.

‘An erect (or scandent ?) shrub, the branches slender, brown-
ish, striate, somewhat zig-zag between the distant leaves,
glabrous. Leaves narrowly ovate to oblong-ovate, coriaceous,
shining on both surfaces, 14 to 18 cm long, 6 to 7 cm wide,
entire, the base acute, the apex slenderly subcaudate-acuminate,
the base prominently 7-plinerved, sometimes with an additional
outer fainter pair, the inner one or two pairs reaching the apex,
the primary reticulations lax, the ultimate ones rather dense;
petioles stout, 8 mm long. Racemes axillary, solitary, up to 8
cm in length, each about 10-flowered, all parts sparingly pubes-
cent with short, white, scattered hairs, the bracteoles oblong-
lanceolate, sharply acuminate, about 2 cm long, deciduous, the
pedicels 1 to 1.5 cm long. Calyx-tube cup-shaped, slightly con-
stricted above, the limb somewhat spreading, making the whole

54 The Philippine Journal of Science 1915

calyx somewhat urceolate, the tube 2.5 mm long, about 3 mm
in diameter, the lobes 5, broadly triangular, acute or acuminate,
1.8 mm long, margins distinctly ciliate. Corolla red, slightly
funnel-shaped, rather broad, 1.6 cm long, sparingly pubescent
outside, the lobes 5, short, acute or rounded, about 4 mm wide
and 2 mm long. Stamens 10; filaments 9 mm long, more or
less pilose with long white hairs; anthers 8.5 mm long, narrowly
oblong, base slightly curved and apiculate, the apical tubes
laterally compressed, 5 mm long, the slit slightly oblique. Top
of the ovary somewhat white-hispid; style 1.5 cm long, glabrous.
Fruit unknown.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19846 McGregor,
February 11, 1913, indicated by the collector, with query, as a vine.

A species manifestly allied to Vaccinium indutum Vidal from which
it is distinguished by its larger, more prominently nerved, glabrous or
nearly glabrous leaves; from Vacciniwm barandanum Vid., which it also

resembles, it is distinguished by the vegetative characters just indicated
and its pubescent inflorescence.

VACCINIUM TURBINATUM sp. nov.

Frutex epiphyticus, glaber; foliis crasse coriaceis, obovatis ad
anguste obovatis, usque ad 9 cm longis, obtusis vel abrupte bre-
viter obtuseque acuminatis, basi acutis, integris, margine re-
volutis, nervis utrinque 6, tenuibus, adscendentibus; floribus
ignotis; racemis in axillis superioribus vel terminalibus, fruc-
tibus longe pedicellatis, turbinatis, circiter 4 superioribus, 7 mm
diametro.

An epiphytic glabrous shrub (flowers unknown), the branches
olivaceous or brownish and more or less sulcate when dry, the
growing parts reddish-brown. Leaves thickly coriaceous, ob-
ovate to narrowly obovate, 5 to 9 cm long, 2.5 to 5 em wide,
apex obtuse or abruptly, broadly, shortly, and obtusely acumin-
ate, the base acute, entire, the margins revolute, with usually one
or two pairs of prominent glands above the insertion of the
petiole, shining on both surfaces, the upper surface pale, the
lower one brownish when dry; lateral nerves about 6 on each
side of the midrib, ascending, slender, not prominent, anastomos-
ing; petioles stout, 5 to 7 mm long; bracteoles deciduous (not
seen). Flowers unkown. Racemes, in fruit, about 5 cm long,
in the uppermost axils or terminal, solitary, few, the pedicels
about 1.5 em long, thickened upward and about 2 mm thick at
the apex, distinctly jointed to the calyx. Fruit turbinate, about
5 mm in diameter, one-third superior, the persistent calyx-teeth
broadly triangular, acute or obtuse, about 2 mm long.

LuzON, Province of Laguna, San Antonio, in forests on trees, Bur. Sei.
15068 Ramos, June, 1912.

a

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 55

A characteristic species distinguishable by its fruit being one-third
superior, the rounded upper part of the fruit as wide as the calyx in its
thickest part, protruding above the persistent calyx-teeth. The plant
somewhat resembles Vaccinium jagori Warb. in vegetative characters, but
has larger, longer petioled leaves.

RHODODENDRON Linnaeus

RHODODENDRON LEYTENSE sp. nov. § Vireya ?

Frutex epiphyticus, subglaber, ramis ramulisque teretibus;
foliis coriaceis, alternis vel subverticillatis, oblongis vel oblongo-
ellipticis, usque ad 7 cm longis, utrinque angustatis acutisque,
supra nitidis, subtus lepidotis, nervis lateralibus utrinque cir-
citer 8, tenuibus, obscuris; floribus terminalibus, in umbellis
sessilibus dispositis, bracteis involucrantibus caducis, oblongis,
acuminatis, circiter 2.5 cm longis; corolla flava, 4 cm longa, late
tubuloso-campanulata.

An epiphytic, nearly glabrous shrub, the branches and branch-
lets terete, grayish or reddish-brown, smooth, the ultimate ones
about 2.5 mm in diameter. Leaves alternate or subverticillate,
coriaceous, oblong to oblong-elliptic, 4 to 7 cm long, 1.5 to 3
cm wide, subequally narrowed to both the acute base and apex,
or the base sometimes a little decurrent-acuminate, brownish
when dry, the upper surface glabrous, shining, the lower some-
what paler, and with numerous, but not densely arranged, brown
lepidote scales; lateral nerves slender, obscure, about 8 on each
side of the midrib; petioles 1 to 1.5 cm long. Inflorescence ter-
minal, the flowers in sessile umbels, in bud quite enclosed by im-
bricate bracts, the bracts caducous, brown when dry, glabrous,
shining, oblong, acuminate, about 2.5 cm long, the bracteoles
narrow. Flowers yellow, usually 4 or 5 in each umbel, their
pedicels pubescent, about 1 cm long in anthesis, twice as long
in young fruit. Calyx obsolete, represented by a mere thicken-
ing of the apex of the pedicel. Corolla broadly tubular-cam-
panulate, yellow, about 4 cm long, the tube broad, about 2 cm
long, the lobes broadly elliptic to obovate, rounded, 1.3 to 1.5
cm wide. Stamens 9 or 10, the filaments slender, a little un-
equal; anthers oblong, obtuse, 3 mm long. Ovary rather densely
pubescent, cylindric, elongated, narrowed upward into the style
which is pubescent below and glabrous above; stigma with 5,
stout, broad, obtuse lobes.

LEYTE, Mount Ibuni, back of Dagami, Bur. Sci. 15252 Ramos, August
23, 1912, growing in the tops of trees.

A rather characteristic species, among the Philippine forms perhaps
most closely allied to Rhododendron kochii Stein, although entirely different

from that species in many details of its leaves, and in the color and
character of its flowers.

56 The Philippine Journal of Science 1915

SAPOTACEAE
BASSIA Koenig

There appears to be no reason whatever for substituting the generic
designation Jillipe for Bassia, as Baillon and Engler have done. This
conclusion has been reached by Trimen, Cooke, and Gamble,’ and at my
request Mr. F. V. Coville has examined the original publication of the
two names, expressing the opinion that Illipe, as published in Linn.
Mantissa 2 (1771) 563, has no standing whatever as a published generic
name. It is manifest that Koenig merely intended Illipe to represent one
of the native names of the original species, Bassia longifolia Linn. This
status of the names Bassia and Illipe is confirmed by Richter."

Following Engler and Prantl several Philippine species have been
referred to the genus I/lipe, but are now transferred to the genus Bassia.
It is possible that more complete material will necessitate the transfer
of some to the genus Payena.

BASSIA BETIS (Blanco) comb. nov.

Azaola betis Blanco Fl. Filip. (1837) 402.
Payena betis F.-Vill. Novis. App. (1880) 125.
Illipe betis Merr. in Bull. Bureau of Forestry (Philip.) 1 (1903) 46.

A large tree of wide distribution in the Philippines, yielding a valuable
timber commercially known as betis.
BASSIA CORIACEA (Merr.) comb. nov.

Illipe coriacea Merr. in Govt. Lab. Publ. (Philip.) 17 (1904) 41.

BASSIA MULTIFLORA (Merr.) comb. nov.
Illipe multiflora Merr. 1. c.

BASSIA RAMIFLORA (Merr.) comb. nov.
Illipe ramiflora Merr. 1. c. 42.

BASSIA MONTICOLA sp. nov.

Arbor circiter 8 m alta, glabra vel subglabra; foliis crasse
coriaceis, oblongis ad oblongo-oblanceolatis, pallidis, nitidis,
usque ad 14 cm longis, obtusis, basi cuneatis, nervis utrinque
circiter 15; fructibus fasciculatis, e ramis defoliatis, ovoideis
ad oblongo-ovoideis, in siccitate brunneis, glabris; sepalis late
ovatis, extus parce pubescentibus glabrescentibus, circiter 6 mm
longis.

A tree up to 8 m in height, nearly glabrous. Branches stout,
terete, brownish, brown-pubescent at the attachment of the
pedicels, otherwise glabrous, the branchlets with numerous peti-
olar scars. Leaves crowded near the apices of the branchlets,
oblong to somewhat oblong-oblanceolate, thickly coriaceous,
when dry pale and shining on both surfaces, 9 to 14 cm long,

‘Trimen Fl. Ceyl. 2:79; Cooke Bombay Flora (cited by Gamble);
Gamble in Journ. As. Soc. Beng. 72? (1905) 176.
*Codex Botanicus Linnaeanus (1840) 455.

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 57

8 to 4 cm wide, narrowed upward to the obtuse apex and below
to the cuneate base; lateral nerves about 15 on each side of the
midrib, distinct on the lower surface, the reticulations lax, not
prominent; petioles stout, 1 to 1.5 cm long. Flowers not seen.
Fruits numerous, fascicled on the branches in the axils of fallen
leaves, their pedicels brown, 1.2 to 2.2 cm long, glabrous, the
points of attachment densely brown furfuraceous-pubescent.
Young fruits brown when dry, ovoid to oblong-ovoid, glabrous,
about 1.5 cm long, tipped by the rather stout 1 cm long style,
the persistent calyx-lobes four, 2-seriate, broadly ovate, coria-
ceous, obtuse, the outer two about 6 mm long, 5 mm wide, the
inner two somewhat thinner and a little broader, outside with
few, appressed, dark-brown, shining hairs, becoming nearly or
quite glabrous.

PALAWAN, Silanga Peak, Merrill 9622, May 30, 1913, in forests on the
exposed peak, summit, altitude about 530 meters, distributed as Payena.

Apparently allied to Bassia multiflora Merr., but with relatively much
narrower, less prominently nerved leaves.

BASSIA OBOVATIFOLIA sp. nov.

Arbor glabra circiter 20 m alta; foliis crasse coriaceis, obo-
vatis, usque ad 12 cm longis, in siccitate nitidis, pallidis, apice
obtusis, rotundatis vel retusis, basi cuneatis, nervis utrinque
circiter 12; fructibus e axillis defoliatis, ellipsoideis, circiter 2.5
em longis, pedicellis circiter 2 cm longis, sursum incrassatis;
sepalis persistentibus, circiter 5 mm latis, irregulariter lacerato-
retusis.

A tree about 20 m in height, entirely glabrous, the branches
reddish-brown, terete, somewhat wrinkled when dry. Leaves
thickly coriaceous, obovate to oblong-obovate, 7 to 12 cm long,
4 to 6.5 cm wide, pale and shining when dry, apex obtuse, broadly
rounded or retuse, base narrowed, cuneate, margins recurved;
lateral nerves about 12 on each side of the midrib, distinct, the
reticulations lax, not prominent; petioles stout, about 1.5 cm
long. Fruits solitary or in pairs in the axils of fallen leaves on
the ultimate branchlets, in general ellipsoid, coarsely wrinkled
when dry, pale-brownish, about 2.5 cm long, the style persistent
as a slender straight apiculus up to 1 cm in length. Persistent
calyx lobes 4, in two pairs, suborbicular, concave, coriaceous,
about 5 mm wide, irregularly lacerate-retuse; pedicels about 2
em long, rather prominently thickened upward.

Luzon, Province of Camarines, near Daet, For. Bur. 21426 (type), 21454
Alvarez, May, 1914, on forested slopes, 20 to 50 meters altitude.

A species similar to and manifestly closely allied to Bassia coriacea

Merr., from which it is readily distinguished by its obovate leaves and
fewer lateral nerves; distributed as Palaquium.

58 The Philippine Journal of Science 1915

BASSIA MINDANAENSIS sp. nov.

Arbor circiter 15 m alta, floribus exceptis glabra; foliis
oblongis, usque ad 23 cm longis, subcoriaceis, in siccitate brun-
neis, nitidis, apice breviter acuminatis, basi acutis, nervis
utrinque 15 ad 17, subtus prominentibus; filoribus axillaribus,
fasciculatis, sepalis 2-seriatis, extus pubescentibus, circiter 7.5
mm longis; staminibus circiter 30; ovario glabro, 8-loculare.

A tree about 15 m high, glabrous except the flowers. Ultimate
branches subterete, brown, smooth. Leaves oblong, subcoria-
ceous, brown and shining when dry, 12 to 23 cm long, 5 to 8
em wide, base shortly but rather prominently acuminate, the
acumen blunt, base acute, sometimes a little inequilateral; lateral
nerves 15 to 17 on each side of the midrib, very prominent on
the lower surface, the reticulations rather fine, distinct; petioles
1.5 to 2 cm long. Flowers white, fragrant, axillary, about 3
in each fascicle, their pedicels somewhat appressed-pubescent,
10 to 12 mm long. Sepals four, 2-seriate, the outer two broadly
ovate, obtuse, coriaceous, appressed-pubescent, about 7.5 mm
long, 6.5 mm wide, the inner two thinner, somewhat narrower,
their margins ciliate. Corolla tube about 3.5 mm long, enlarged
upward, villous inside, the lobes 8, erect, oblong-lanceolate, ob-
tuse, 6 to 7 mm long, 1.5 to 2 mm wide. Stamens about 30, the
filaments very short; anthers lanceolate, slenderly acuminate,
about 4 mm long, slightly pubescent. Ovary glabrous, 8-celled;
style about 1 cm long.

MINDANAO, Province of Misamis, east of Mount Catmon, For. Bur. 17977
Miranda, February 21, 1913, in forests along streams, altitude about 120
meters.

A species recognizable by its leaves turning brown in drying, its
prominent nerves, and its glabrous ovaries.

BASSIA PLATYPHYLLA sp. nov.

Arbor alta, floribus exceptis glabra; foliis oblongo-ellipticis,
usque ad 30 em longis, coriaceis, pallidis, nitidis, apice rotundatis
vel brevissime obtuse abrupteque acuminatis, basi acutis, nervis
utrinque circiter 23, subtus valde prominentibus; floribus axil-
laribus, longe pedicellatis, sepalis coriaceis, extus leviter pubes-
centibus, late ovatis, obtusis, circiter 8 mm longis.

A tree reaching a height of about 20 meters, glabrous except
the flowers. Ultimate branchlets terete, brown, 4 to 5 mm in
diameter. Leaves oblong-elliptic, coriaceous, 23 to 30 cm)\long,
10 to 13 em wide, pale and shining when dry, the apex rounded
or very shortly, broadly, obtusely, and abruptly acuminate, base
acute; lateral nerves about 23 on each side of the midrib, very
prominent on the lower surface, curved-spreading, scarcely anas-

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 59

tomosing, the reticulations distinct; petioles 4.5 to 5.5 cm long,
much thickened and rugose at the base. Flowers axillary, about
5 in each axil, the pedicels glabrous or very slightly pubescent,
nearly 3 cm long. Calyx-lobes four, 2-seriate, coriaceous,
broadly ovate, obtuse, about 8 mm long, 7 mm wide. Corolla
and stamens not seen. Fruit “green, globose” (not seen) ; seeds
brown, shining, about 18 mm long, 10 mm wide, slightly com-
pressed, acute.

MINDANAO, District of Zamboanga, Margosatubig, For. Bur. 21873 Villa-
mil, June 9, 1914, forested slopes, altitude about 70 meters.

A species characterized by its large leaves with numerous, very prom-
inent nerves; flowering specimens may show it to belong in some other
genus.

SIDEROXYLON Linnaeus

SIDEROXYLON VILLAMILII sp. nov.

Arbor alta partibus junioribus floribusque parce pubescentibus
exceptis glabra; foliis oblongis, chartaceis vel subcoriaceis, usque
ad 20 cm longis, nitidis, sursum angustatis, obtusis, basi cuneatis
vel decurrento-acuminatis, nervis utrinque circiter 8, subtug
prominentibus; floribus parvis, axillaribus, fasciculatis, breviter
pedicellatis; sepalis 6, circiter 2 mm longis, extus parce pubes-
centibus; staminoideis oblongis, obtusis, integris, 0.7 mm longis;
ovario dense hirsuto.

A tree reaching a height of 30 meters, glabrous except the
slightly cinereous-pubescent flowers and pedicels and the growing
tips of the branchlets. Branches terete, dark reddish-brown
when dry, glabrous. Leaves oblong, chartaceous to subcoria-
ceous, 10 to 20 cm long, 3.5 to 7 em wide, about equally narrowed
to the obtuse tip and to the cuneate or decurrent-acuminate base,
dark-olivaceous and shining when dry; lateral nerves about 8
on each side of the midrib, slender but prominent, curved, anas-
tomosing, the reticulations lax, distinct; petioles 1 to 3 em long.
Flowers small, axillary, fascicled, up to 7 in each fascicle, the
pedicels sparingly cinereous-pubescent, 3 to 5 mm long. Sepals
6, outside sparingly cinereous-pubescent, about 2 mm long, 1.5
to 2.5 mm wide, concave, rounded, the inner ones larger than
the outer ones, varying from oblong-ovate to subreniform-ovate.
Corolla tubular, about 2.7 mm long, glabrous, the lobes 5, orbi-
cular-ovate, rounded, about 1 mm in diameter. Stamens 5, the
filaments very short; anthers ovoid, acute, 0.6 mm long. Sta-
minodes alternating with the corolla lobes, oblong, obtuse, entire,
0.7 mm long, glabrous. Ovary densely hirsute; style stout,
glabrous, 2 mm long. Fruit globose or depressed-globose, gla-
brous, fleshy, when dry up to 4 cm in diameter, the seeds paie-

60 The Philippine Journal of Science 1918

brown, shining, hard, somewhat compressed, about 1.8 cm long
and 1.2 cm wide.

Luzon, Province of Laguna, Mount Maquiling, For. Bur. 19762 (type),
20604 Villamil, June and September, 1913, and from the same tree, For.
Bur. 18154 Curran, November, 1911, with mature fruits, For. Bur. 11927
Tamesis, April, 1910, locally known as dolitan.

A species resembling in vegetative characters Sideroxylon macranthum
Merr., but quite different in floral characters; it is readily distinguished
by its very much smaller flowers which are but slightly pubescent.

SAPOTACEAE
PALAQUIUM Blanco

PALAQUIUM FOXWORTHY'I sp. nov.

Arbor, partibus junioribus floribusque exceptis glabra; foliis
coriaceis, oblongo-obovatis ad obovatis, usque ad 12 cm longis,
in siccitate brunneis, nitidis, nervis utrinque 12 ad 14, apice
breviter obtuseque acuminatis, basi cuneatis; floribus axillaribus,
fasciculatis, sepalis extus pubescentibus; staminibus 16; ovario
6-loculare, pubescente; pedicellis circiter 1.5 cm longis, leviter
pubescentibus.

A tree, apparently of large size, glabrous except the very tips
of the branchlets, pedicels, and parts of the flowers. Branches
terete, grayish-brown, glabrous. Leaves coriaceous, oblong-
obovate to obovate, 8 to 12 cm long, 3 to 5 cm wide, dark-brown
and shining when dry, the lower surface a little paler than the
upper, the apex very shortly and broadly blunt-acuminate, base
gradually narrowed, cuneate; lateral nerves 12 to 14 on each
side of the midrib, prominent, the reticulations very fine; petioles
5 to 10 mm long. Flowers axillary and in the axils of fallen
leaves, fascicled, 2 to 5 in each axil, their pedicels slightly pu-
bescent, about 1.5 cm long. Sepals broadly ovate, about 3 mm
long, the outer three thickly coriaceous, acute or slightly acu-
minate, the inner three rounded or obtuse. Corolla tube about
3 mm long, the lobes 6, reflexed, lanceolate, about 6 mm long,
2.5 mm wide. Stamens 16; filaments 3 mm long; anthers lan-
ceolate, acuminate, 2.8 mm long. Ovary appressed-pubescent,
6-celled; style about 12 mm long.

Luzon, Province of Tayabas, Apad, Bur. Sci. 18112 Foxworthy &
Ramos, March 8, 1911, in forested valleys, altitude about 30 meters.

A species similar to Palaquium lanceolatum Blanco, differing in its
somewhat smaller leaves, much shorter, less pubescent pedicels, smaller
flowers, and pubescent ovaries. To this species I tentatively refer the

following sterile material: Luzon, Province of Laguna, For. Bur. 22328
Mariano: Province of Pangasinan, For. Bur. 19441 Agama.

x,C,1 Merrill: New or Noteworthy Philippine Plants, XI 61

PALAQUIUM GLABRUM sp. nov.

Arbor alta, sepalis interioribus exceptis glabra; foliis sub-
coriaceis, usque ad 16 cm longis, obovatis ad oblongo-obovatis,
in siccitate brunneis, nitidis, apice obtusis ad rotundatis, basi
cuneatis, nervis utrinque 8 ad 10, subtus prominentibus; flori-
bus numerosis, axillaribus, fasciculatis, longe pedicellatis, sepalis
exterioribus glabris, interioribus leviter pubescentibus; stami-
nibus 18; ovario glabro, 6-loculare.

A tree about 20 m high, quite glabrous except the inner sepals.
Branches terete, grayish-brown, rugose when dry. Leaves
rather crowded on the ultimate branchlets, obovate to oblong-
obovate, 11 to 16 cm long, 5 to 7 cm wide, subcoriaceous, brown
and shining when dry, of about the same color on both surfaces,
the apex obtuse to somewhat rounded, base gradually narrowed,
cuneate; lateral nerves 8 to 10 on each side of the midrib, prom-
inent on the lower surface, the reticulations very slender, not
prominent; petioles 1 to 1.5 cm long. Flowers very numerous,
in the axils of leaves and of fallen leaves, several in each axil,
their pedicels slender, glabrous, about 3 cm long. Sepals 6,
2-seriate, broadly ovate, coriaceous, 3.5 to 4 mm long, the outer
three quite glabrous, the inner three sparingly pubescent. Co-
rolla lobes (in bud) 6, oblong, obtuse, glabrous, 3.5 mm long.
Stamens 18; anthers lanceolate, acuminate, 2.5 mm long. Ovary
glabrous, 6-celled; style stout, about 2.5 mm long.

Luzon, Province of Rizal, back of Bosoboso, Bur. Sci. 13582 Ramos,
August 15, 1911, in forests along streams, locally known as alacaac na
putt.

A very characteristic species, readily recognized by its long slender
pedicels, and in being quite glabrous in all parts except the inner sepals.

PALAQUIUM HETEROSEPALUM sp. nov.

Arbor circiter 15 m alta, ramulis junioribus floribusque ex-
ceptis glabra; foliis crasse coriaceis, obovatis, apice rotundatis,
basi cuneatis, usque ad 4 cm longis, nervis utrinque circiter 10,
obscuris vel subobsoletis; floribus breviter pedicellatis, circiter
6 mm longis, sepalis 4 vel 6, staminibus 12.

A tree about 15 m high, quite glabrous except the growing tips
of the branchlets and the flowers. Branches and branchlets
rather stout, dark-colored when dry, glabrous. Leaves crowded
toward the apices of the branchlets, thickly coriaceous, obovate,
2 to 4 cm long, 1.5 to 2.5 cm wide, brownish when dry, shining,
apex broadly rounded, base cuneate; lateral nerves about 10 on
each side of the midrib, very slender, obscure or nearly obsolete;
petioles 5 mm long or less. Flowers in the axils of fallen leaves

62 The Philippine Journal of Science 1916

on the ultimate branchlets, rather numerous, white, their pedicels
slightly ferruginous-pubescent with short appressed hairs as
are the sepals externally. Sepals 4 or 6, orbicular-ovate, coria-
ceous, rounded, 2 to 3 mm long, the inner two or three a little
larger than the outer two or three. Corolla about 6 mm long,
the tube about 1.5 mm, the lobes 6, oblong-elliptic, rounded or
obtuse, about 4.5 mm long, 2.5 to 3 mm wide, externally very
slightly pubescent. Stamens 12; anthers oblong, acute or acu-
minate, about 2.5 mm long. Ovary ovoid, pubescent, 5- to 7-
celled, narrowed upward into the rather stout style, the ovary
and style about 6 mm long.

LuZON, Province of Rizal, Mount Susong Dalaga, Bur. Sci. 19160 Reillo,
December 6, 1914, altitude not indicated, but from the mossy forest.

The species somewhat resembles a small leaved form of Palaquium
cuneatum Vidal, and is characterized not only by its unusually small
leaves, but by its very obscure, often nearly obsolete nerves, very short
pedicels, and the variable number of sepals, either 4 or 6; in those flowers
with 4 sepals, however, there are always 6 corolla lobes and 12 stamens.

I know of no other species of Palaquium where the number of calyx lobes
is variable.

PALAQUIUM LANCEOLATUM Blanco FI. Filip. (1837) 403, ed. 2 (1845)
282, ed. 8, 2 (1878) 159.

This species is of importance as it is the type of the genus Palaquium
Blanco, and no material previously referred to the species, unless it be
Vidal’s specimen cited by Dubard, has agreed with Blanco’s description
in essential characters; specimens referred by me at various times to the
species have correctly been indicated by Dubard as a distinct species,
Palaquium merrillii Dubard,’ as Blanco’s species has 18 stamens, and the
material Dubard has referred to P. merrillii has but 12 stamens. Speci-
mens in flower, recently collected, agree in practically all details with
Blanco’s description, and I feel perfectly confident that the present inter-
pretation of the species is the correct one, and an amplification of Blanco’s
description follows:

A tall tree, quite glabrous except the younger parts and the
flowers. Branchlets and young petioles ferruginous-pubescent.
Leaves oblong to oblong-oblanceolate or obovate, rarely ap-
proaching the lanceolate type, coriaceous or subcoriaceous, brown
and shining when dry, glabrous on both surfaces, shortly and
obtusely acuminate to obtuse or even rounded at the apex, base
narrowed, cuneate; lateral nerves about 11 on each side of the
midrib, prominent, the reticulations fine; petioles about 1 mm
long. Flowers numerous, axillary, in the axils of leaves and of
fallen leaves on the ultimate branchlets, fascicled, their pedicels
ferruginous-pubescent, 2.5 to 4cm long. Outer 3 sepals densely
ferruginous-pubescent, ovate, acute to somewhat acuminate, cori-

* Bull. Mus. Hist. Nat. Paris 15 (1909) 381.

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 63

aceous, about 4.5 mm long, 3 to 3.5 mm wide, the inner three
thinner, oblong to oblong-elliptic, obtuse, pubescent, 4 to 4.5
mm long, 2.5 to 3 mm wide. Corolla glabrous, the tube about
_3 mm long, the lobes lanceolate, acuminate, reflexed, about 8
mm long, 2 mm wide. Stamens 18; filaments 4 to 5 mm long,
3 to 3.5 mm wide. Ovary glabrous, 6-celled; style 12 to 15 mm
long.

Luzon, Province of Tayabas, Mauban, Bur. Sci. 19483 Ramos, January,
1913: Province of Camarines, Sagnay, Bur. Sci. 22135 Ramos, December,
1913.

As to Blanco’s description, the above specimens agree with it absolutely
in all respects except that the leaves are not “lanceoladas,” but while some
approach the lanceolate type, most of them are oblong-obovate to oblong-
oblanceolate; they are glabrous and the petioles are short. The flowers
agree perfectly especially in the corolla lobes “estrechas, aguzadas, y muy
revueltas hacia abajo.” The stamens are 18 (in Blanco’s description 18,
19, or fewer); ovary (fruit) 6-celled; style longer than the stamens.
Blanco gives the date of flowering as March, but saw only old dried
flowers; his specimens were from the Province of Batangas which may
account for the difference in the time of flowering, which at most would
be about one month, as the specimens from Camarines were collected on
January 25. The native (Tagalog) name bagalangit does not appear on
any of our specimens of Palaquiwm.

Whether or not “Vidal 7” referred to Blanco’s species by Dubard is the
same as my interpretation of Blanco’s species I cannot say. I have a
copy of Pierre’s detailed drawing of the flower, apparently taken from
young buds, and it does not appear to be the same as the specimens cited
above; the differences, however, may be due to the stage of development.
At any rate, Pierre’s drawing does not agree with Blanco’s description in
the corolla lobes which in Vidal’s specimen are rounded and merely elliptic,
not “estrechas, aguzadas.”

PALAQUIUM NEGROSENSE sp. nov. § Palaquoides.

Arbor circiter 20 m alta, partibus junioribus inflorescentiisque
exceptis glabra; foliis crasse coriaceis, obovatis, usque ad 22 cm
longis, in siccitate brunneis, nitidis, apice late rotundatis, basi
angustatis, cuneatis, nervis utrinque circiter 12, subtus prom-
inentibus; floribus axillaribus, paucis, sepalis extus pubescen-
tibus; staminibus 18; ovario parce pubescente, 6-loculare.

A tree about 20 m high, glabrous except the younger parts and
the flowers. Branches rather stout, terete, brown, the growing
tips somewhat ferruginous-pubescent, as are the very young
petioles, the pedicels, and the calyx externally. Leaves thickly
coriaceous, obovate, 14 to 22 cm long, 6 to 12 cm wide, apex
broadly rounded, base gradually narrowed, cuneate, when dry
brown and shining on both surfaces; lateral nerves about 12 on
each side of the midrib, prominent, the reticulations very fine;
petioles 1.5 to 2.5 cm long. Flowers few, axillary and in the

64 The Philippine Journal of Science 1915

axils of fallen leaves, but one or two in each axil, the pedicels
ferruginous-pubescent, up to 3 cm long. Young sepals broadly
ovate, obtuse, ferruginous-pubescent, coriaceous, about 4 mm
long. Very young corolla-lobes elliptic-ovate, rounded or slightly |
retuse, 3 mm long. Stamens 18, the young anthers 2 mm long.
Ovary very slightly pubescent, 6-celled. Fruit oblong, inequi-
lateral, up to at least 3 cm in length (immature) tipped by the
style which is up to 14 mm in length, glabrous.

NEGROS, Cadiz, For. Bur. 15029 Danao, March, 1908; Paniquon River,
For. Bur. 7325 Everett, March, 1907; near San Carlos, For. Bur. 20696
Tamesis (type), September, 1913, locally known as nato and bayog.

This is the species that Dubard has referred, with doubt, to Palaquium
obovatum King & Gamble,” from fruiting specimens; a more recent collec-
tion, however, presents young flowers, and an examination of these shows
18, not 12 stamens, so that the specimens must be referred to the section
Palaquoides, not to Eupalaquium, to which P. obovatum belongs. It is
closely allied to Palaquiwm lanceolatum Blanco, differing in its larger
leaves which are broadly rounded at the apex, not at all acuminate, its
somewhat shorter pedicels, its much fewer flowers, but one or two in each
axil, and its slightly pubescent ovary.

APOCYNACEAE
ALYXIA R. Brown
ALYXIA CONFERTIFLORA sp. nov.

Species ut videtur A. lucidae Wall. affinis, differt inflorescentiis
valde congestis, haud pedunculatis, quam petiolo multo brevio-
ribus, floribus longioribus, circiter 1 cm longis.

A scandent shurb, quite glabrous except the inflorescence.
Branches smooth, reddish-brown, distinctly 4-angled. Leaves
fragrant in drying, whorled, 4 in each whorl, oblong, chartaceous
to subcoriaceous, 6 to 10 em long, 2 to 3.5 cm wide, about equally
narrowed at both ends, the apex blunt-acuminate, the base de-
current-acuminate, both surfaces slightly shining when dry, the
lower a little paler than the upper, the margins recurved; lateral
nerves very slender, indistinct, about 50 on each side of the
midrib, anastomosing with the faint submarginal nerve; petioles
1 to 1.5 ecm long. Cymes axillary, sessile or very shortly pe-
duncled, congested, about 0.8 mm long (excluding the corollas),
pubescent, the flowers sessile, 5-merous, each subtended by an
ovate, pubescent, acute or acuminate bract about 3 mm long and
several similar but much smaller bracteoles. Calyx 3 mm long,
the lobes ovate, acute, about 2 mm long, ciliate-pubescent or
puberulent. Corolla-tube cylindric, externally glabrous, 10 mm
long, swollen above opposite the anthers, contracted at the

* Bull. Mus. Hist. Nat. 15 (1909) 280.

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 65

throat, the tube prominently pubescent inside; lobes spreading-
recurved, orbicular, 3 mm in diameter. Anthers narrowly ovate-
lanceolate, 1.2 mm long. Ovary short, pubescent. Fruit not
seen.

BASILAN, Bur. Sci. 16143 Reillo, September, 1912.

A species manifestly allied to the Malayan Alyzxia lucida Wall., which
it greatly resembles, but from which it is distingushed by its Geobeestad
inflorescence which is much shorter than the petioles, its longer flowers,
and more numerously nerved leaves. It is also closely allied to Alyzxia
odorata Wall., perhaps closer to this than to A. lucida, but is distinguish-
able by its acuminate, not acute or obtuse leaves, shorter cymes, sessile
flowers, and orbicular, not ovate, corolla-lobes.

ALSTONIA R. Brown

ALSTONIA OBLONGIFOLIA sp. nov, § Dissuraspermum.

Arbor circiter 10 m alta, ramulis subtus foliis inflorescen-
tiisque leviter pubescentibus; foliis ternis vel quaternis, char-
taceis vel subcoriaceis, subtus pallidis, oblongis ad oblongo-
lanceolatis, usque ad 14 cm longis, nervis distantibus, utrinque
14 ad 18, distinctis; inflorescentiis terminalibus, floribus circiter
8 mm longis.

A tree about 10 m high. Branches obscurely angled, slightly
pubescent. Leaves whorled, 3 or 4 at each node, oblong to
oblong-lanceolate, narrowed and somewhat acuminate at both
ends, chartaceous or subcoriaceous, 6 to 14 cm long, 2 to 4 cm
wide, the upper surface glabrous, shining, brown when dry, the
lower surface much paler, slightly pubescent, especially on the
midrib and nerves; nerves distant, spreading, distinct, brown,
14 to 18 on each side of the midrib, the reticulations slender,
not prominent; petioles somewhat pubescent, less than 1 cm
long. Inflorescence terminal, sessile, that is numerous peduncles
springing from the apex of the branchlet, the peduncles slender,
slightly pubescent, 2 to 7 cm long, subumbellately or corymbosely .
branched at their apices, the branches gray-villous but not
densely so. Flowers numerous, white, their pedicels 1 to 2 mm
long. Calyx cup-shaped, slightly gray-pubescent, about 1.5 mm
long, the lobes very short. Corolla-tube about 5 mm long, cylin-
dric, slightly pubescent on the outside, somewhat inflated above
the middle opposite the insertion of the stamens, inside and
throat villous, the lobes oblong, obtuse, 3.5 to 4 mm long, over-
lapping to the right, their margins ciliate-villous. Anthers in-
serted above the middle, about 1 mm long. Ovary glabrous,
narrowed into the slender, 2 mm long style.

PALAWAN, in old clearings near Puerto Princesa, For. Bur. 4495 Curran,
June 5, 1906.

66 The Philippine Journal of Science 1915

A species apparently allied to Alstonia angustifolia Wall., of the Malay
Peninsula, but differing in many characters, notably, in its quite different
flowers.

KICKXIA Blume

KICKXIA BLANCO! Rolfe in Journ. Linn. Soc. Bot. 21 (1884) 313.

Kickxia arborea Vid. Sinopsis Atlas (1883) t. 67, f. G; F.-Vill. Novis.

App. (1880) 132; Naves in Blanco FI. Filip. ed. 3, t. 428 bis, non
Blume.

Kickxia merrittti Merr. in Philip. Journ. Sci. 4 (1910) Bot. 355.

Kickxia macgregorti Elm. Leafl. Philip. Bot. 4 (1912) 1457.

Rolfe’s species is typified only by the plate in the third edition of
Blanco’s Flora de Filipinas, no description having been published; the
plate well represents the species as here interpreted. There is some
variation in vegetative characters, but in essentials the species is quite
constant. A careful examination of all available material has convinced
me that the two recently described species, above cited as synonyms, are
not distinct from Kickxia blancoi Rolfe. The species is sometimes known
as lanete, as laneting gubat, and as ayate, and probably represents the
form mentioned by Blanco Fl. Filip. (1887) 114, ed. 2 (1845) 81 after
the deecaption of Anasser laniti as “Hai otra especie con las flores axilares
solitarias.” It is-represented by the following material:

Luzon, Province of Batangas, Looc, For. Bur. 7649 Curran & Merritt;
Santo Tomas, For. Bur. 21545 Tamesis: Province of Laguna, Calamba,
For. Bur. 20964 Villamil. Muinporo, For. Bur. 11488 Merritt (type of
K. merrittii Merr.). GuimarAs, For. Bur. 862 Gammill. SiBuyaNn, Elmer
12378 (type number of K. macgregorii Elm.). NecGros, For. Bur. 15112
Tabat. LEYTE, Wenzel 330, July, 1918. Without definite locality, Loher
6829.

By description alone Kickxia macgregorii Elm. appears to be a remark-
ably distinct form, but this is due to errors in the description. “Calyx
1.5 to 2.25 cm long” is the description of the pedicel, the calyx being very
short; the pedicel is subtended by a very few, small, obscure bracteoles.
The flowers are about 6 cm long as in the other specimens referred to Kichkwia
blancoi Rolfe and as represented by Naves’ plate.

TABERNAEMONTANA Linnaeus

TABERNAEMONTANA HEXAGONA sp. nov.

Frutex glaber, circiter 2 m altus; foliis chartaceis vel sub-
membranaceis, oblongo-ellipticis, usque ad 17 cm longis, acutis
ad abrupte obtuseque acuminatis, nervis utrinque circiter 14,
patentibus; pedunculis axillaribus, ut videtur paucifloris; fruc-
tibus oblongo-lanceolatis, rectis, acuminatis, basi acutis, 6-an-
gulatis vel angulis propriis (3) anguste 3-alatis, alternantibus
vix alatis, circiter 6 cm longis, in siccitate minute verruculosis.

A glabrous shrub, according to the collector about 2 m
high. Branches terete, light-gray, the younger ones somewhat
compressed when dry. Leaves membranaceous to somewhat
chartaceous, oblong-elliptic, the larger ones 18 to 17 cm long, 4
to 7 em wide, base acute or decurrent-acuminate, apex acute to

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 67

shortly, broadly, and obtusely acuminate, when dry shining, the
upper surface pale-olivaceous, the lower very much paler; lateral
nerves about 14 on each side of the midrib, spreading, evanes-
cent or subevanescent near the margins, sometimes obscurely
anastomosing, the reticulations subobsolete; petioles 1 cm long
or less, inflated at the base on the upper side. Peduncles axil-
lary, apparently few-flowered (1-flowered ?), in fruit 3 to 4 cm
long. Flowers not seen. Follicles two, free to the very base,
oblong-lanceolate in outline, straight, not at all curved, acute
at the base, acute or somewhat rostrate-acuminate at the apex,
about 6 cm long, 1.3 to 1.5 cm in diameter, 6-angled, or better
with 3 longitudinal, narrow wings, and 3 alternating, less
prominent ridges, the pericarp coriaceous, greenish-olivaceous
when dry, shining, minutely verruculose. Seeds about 24 in
each follicle, irregularly oblong-obovoid, about 7 mm long.

PANAY, Capiz, Bur. Sci. 21241 Escritor, June, 1918.

The species presents no particularly striking vegetative characters by
which it may be distinguished from some other species. Its follicles,
however, are very distinct in form, being quite different from those of
any other species known to me except the following one. They are straight,
not at all falcate as is usually the case in this genus, free to the very base,
oblong-lanceolate in outline, acuminate, and with 3 longitudinal, narrow,
thick wings, and 3 alternating, less prominent ridges.

TABERNAEMONTANA MINDANAENSIS sp. nov.

Frutex glaber 2 ad 4 m altus; foliis chartaceis, oblongo-lanceo-
altis ad late oblongo-oblanceolatis, usque ad 12 cm longis, utrin-
que angustatis, apice acuminatis, basi acutis, nervis utrinque
circiter 9; cymis axillaribus, brevibus, paucifloris; folliculis rec-
tis vel leviter curvatis, oblongis vel oblongo-lanceolatis, acumi-
natis, 4. ad 5 cm longis, circiter 1 cm diametro.

A glabrous shrub about 4 m high. Branches terete, slender,
gray, not lenticellate, the younger ones sometimes slightly com-
pressed. Leaves chartaceous, oblong-lanceolate to broadly
oblong-oblanceolate, 7 to 12 cm long, 2 to 4.5 cm wide, narrowed
upward to the sharply acuminate apex, the acumen straight or
' somewhat falcate, somewhat apiculate, and to the cuneate base;
lateral nerves about 9 on each side of the midrib, slender but
prominent on the lower surface, the reticulations fine; petioles
4 to 8 mm long. Cymes axillary, few-flowered, in anthesis 4
cm long or less, somewhat elongated in fruit, the immature buds
up to 1.5 cm long, the pedicels less than 1 cm in length. Follicles
usually in pairs, oblong to lanceolate, 4 to 5 em long, about 1
cm in diameter, straight or somewhat curved, with 4 rather
prominert ridges and 2 Jess prominent ones, red, the base

68 The Philippine Journal of Science 1915

acute, the apex acuminate, the pericarp coriaceous. Seeds about
20 in each follicle, irregular, about 6 mm long.

MINDANAO, District of Cotabato, Craan, For. Bur. 14930 Tarrosa &
Almagro, April, 1912 (type); Reina Regente, Bur. Sci. 11653 Robinson,
June, 1910, in thickets and open places at low altitudes.

A species in vegetative characters somewhat resembling the common
Tabernaemontana pandacaqui Poir., but with entirely different fruits,
which resemble those of Tabernaemontana hexagona Merr.; it differs from
T. hexagona Merr. in its smaller, much fewer nerved leaves.

TABERNAEMONTANA OLIGANTHA sp. nov.

Frutex glaber circiter 2 m altus; foliis membranaceis vel
chartaceis, oblongo-ellipticis, usque ad 15 em longis, utrinque
subaequaliter angustatis, apice acuminatis apiculatisque, basi
acutis, nervis utrinque circiter 9, subtus prominentibus; cymis
axillaribus, laxis, circiter 10 cm longis, paucifloris; floribus albis,
longe pedicellatis, circiter 2.5 cm diametro, corollae tubo circiter
1.7 em longo.

A glabrous shrub about 2 m high with very lax, few-flowered,
axillary cymes, the flowers about 5 to each cyme, long pedicelled.
Branches terete, slender, grayish-brown, lenticellate. Leaves
membranaceous to chartaceous, oblong-elliptic, 12 to 15 cm long,
5 to 7 cm wide, olivaceous and somewhat shining when dry,
subequally narrowed to the acute base and to the distinctly acu-
minate and slightly apiculate apex; lateral nerves about 9 on
each side of the midrib, prominent on the lower surface, curved,
obscurely anastomosing, the reticulations fine; petioles 5 mm
long or less. Cymes axillary, solitary, about 10 cm long, each
about 5-flowered, the branches divaricate, lax. Flowers white,
their pedicels 1.5 to 3.5 cm long. Calyx 4 to 5 mm long, the
lobes elliptic-ovate, obtuse or acute, 2.5 to 3 mm long. Corolla
tube cylindric, 1.5 to 1.8 cm long, the lobes spreading, 12 to 13
em long, 4 to 5 mm wide, oblong, falcate, rounded. Anthers 2
mm long. Ovary glabrous.

SAMAR, Cauayan Valley, forested slopes, Phil. Pl. 1621 Ramos, April,
1914,

A species well characterized by its long, very lax, few-flowered cymes,
the flowers about 5 in each cyme, long pedicelled.

URCEOLA Roxburgh

URCEOLA IMBERBIS (Elmer) comb. nov.
Carruthersia imberbis Elmer Leafl. Philip. Bot. 2 (1909) 588.

LuZON, Province of Tayabas, Lucban, Elmer 9239 (type number): Prov-
ince of Laguna, San Antonio, Bur. Sci. 12057 Ramos, August, 1910, in
fruit.

x,c,1 Merrill: New or Noteworthy Philippine Plants, XI 69

The species is in all respects a typical Urceola as evidenced by its very
short flowers and valvate corolla lobes. It belongs in the group with
Urceola brachysepala Hook. f. of the Malay Peninsula. The follicles are
cylindric and brown when dry, not at all torulose, straight or slightly
curved, about 13 cm long, 8 to 4 mm in diameter. The genus, which is
new to the Philippines, is also represented by the following allied
species:

URCEOLA PHILIPPINENSIS sp. nov.

Species praecedente similis et affinis, differt foliis chartaceis,
haud coriaceis, calycis lobis longioribus prope ad basin porrectis.

A scandent shrub, glabrous except the inflorescence, the
branches slender, terete, reddish-brown. Leaves chartaceous,
oblong-ovate, brown or brownish-olivaceous when dry, slightly
shining, 7 to 11 cm long, 3 to 5.5 cm wide, apex prominently
acuminate, base rounded to subacute; lateral nerves about 7 on
each side of the midrib, rather prominent, curved-anastomosing
near the margin, the reticulations distinct; petioles 1 to 1.8 cm
long, reddish-brown. Inflorescence corymbose, terminal and in
the upper axils, frequently forming an ample, somewhat leafy
inflorescence, 7 to 25 cm in width, the younger parts more or
less villous with pale or brownish hairs. Flowers very numerous,
crowded on the ultimate branchlets, the bracts subtending the
primary branches linear, 5 to 15 mm long. Calyx pubescent,
cleft: nearly to the base, the lobes narrowly ovate, 1 to 1.3 mm
long, acute. Corolla about 3 mm long, somewhat pubescent
externally, the lobes 5, narrowly ovate, acute, about 1 mm long,
valvate. Anthers about 1.3 mm long, base sagittate. Disk
prominent, somewhat 5-costate. Ovary glabrous.

MINDANAO, District of Lanao, Camp Keithley, Mrs. Clemens 1098, June,
1907 (type): Province of Surigao, Piper 219, 512, 545, May, 1911. LEYTE,
Jaro, Wenzel 689, May, 1914.

A species manifestly closely allied to Urceola imberbis (Elm.) Merr.,
to which most of the specimens have previously been referred, differing
in its thinner leaves and manifestly much more deeply cleft calyx. In
U. imberbis the leaves, when mature, are rather thickly coriaceous, and the
calyx lobes extend less than half way to the base.

URCEOLA LAEVIS (Elmer) comb. nov.
Carruthersia laevis Elmer Leafl.. Philip. Bot. 4 (1912) 1449 (leavis).

Like Carruthersia imberis Elm., this species has the characteristic
flowers of Urceola and must be transferred to that genus. It is readily
distinguished from the two other Philippine species by its oblong-obovate
bracts and bracteoles, fewer nerves, less prominently acuminate coriaceous
leaves, and especially by its calyx lobes extending quite to the base.

PALAWAN, Puerto Princesa, Elmer 12837, March, 1911, type number.

70 The Philippine Journal of Science 1915

VALLARIS Burman f.

VALLARIS DARONENSIS (Elmer) comb. nov.

Holarrhena daronensis Elmer Leafl. Philip. Bot. 4 (1912) 1455.

MINDANAO, District of Davao, Elmer 11912 (type number), in fruit,
October, 1909: District of Lanao, Camp Keithley, Mrs. Clemens s. n., May,
1907, with young flowers: Province of Misamis, Quadras 100, from Tali-
sayan, locally known as pamacoton.

This species is in all respects a Vallaris, not a Holarrhena, and is
accordingly transferred to the former genus; it is closely allied to Val-
laris gitingensis (Elmer) Merr., but has broader, more prominently nerved
leaves and apparently somewhat larger flowers.

VALLARIS GITINGENSIS (Elmer) comb. nov.
Kickxia gitingensis Elmer Leafl. Philip. Bot. 4 (1912, March 18) 1455.
Vallaris angustifolia Merr. in Philip. Journ. Sci. 7 (1912, November
15) Bot. 335.

The name proposed by Mr. Elmer, sub Kickwia, has priority over that
used by me, sub Vallaris, the latter being the proper genus as the anthers
are exserted, not included in the corolla tube. Mr. Elmer’s specific name
is accordingly adopted, transferred to Kickwia, and K. angustifolia Merr.
reduced as a synonym. The two are identical in all respects. Vallaris
gitingensis is represented by the following material:

Luzon, Province of Laguna, Paete, Holman 8, April, 1910; Nangaiby,
For. Bur. 20175 Manuel, September, 1913 (sterile): Province of Tayabas,
Lucban, Bur. Sci. 19587 Ramos, January, 1913, in fruit: Province of Sor-
sogon, For. Bur. 10507 Curran, June, 1908, For. Bur. 15077 Rosenbluth,
April, 1909 (type of V. angustifolia Merr.). SipuyAN, Elmer 12208, April,
1910 (type number of Kickxia gitingensis Elmer). LEYTE, Wenzel 652,
678, May, 1914. Some of the material has been distributed as Vallaris
holmanii Merr., but I consider now that but a single species is represented.

The follicles are in pairs, about 20 cm long, 5 mm in diameter, nar-
rowed at both ends, straight or slightly curved; seeds linear-lanceolate,
about 2 cm long, the coma plumose, the hairs very soft, pale-brownish,
abundant, 5 to 6 cm long.

VERBENACEAE
CALLICARPA Linnaeus

CALLICARPA LANCIFOLIA sp. nov.

Frutex 1 ad 4 m altus plus minusve stellato-pubescentibus
glandulosisque; foliis lanceolatis vel anguste oblongo-lanceolatis,
chartaceis, usque ad 20 cm longis, basi leviter inaequilateralibus,
obtusis, apice longe caudato-acuminatis, supra parce simpliciter
pubescentibus, eglandulosis, subtus leviter stellato-pubescentibus,
glandulosis, nervis utrinque circiter 11; cymis axillaribus soli-
tariis, laxis, 2 ad 4 em longis, dichotomis; floribus 4-meris, cir-
citer 3 mm longis, corolla glabra vel subglabra.

A shrub 1 to 4m high. Branches terete, slender, subglabrous
or more or less ferruginous-stellate-pubescent, the younger ones

x%c,1 Merrill: New or Noteworthy Philippine Plants, XI Fail

densely stellate-pubescent and with scattered, longer, sparingly
branched hairs intermixed. Leaves chartaceous, lanceolate to
narrowly oblong-lanceolate, 15 to 20 cm long, 3 to 5 cm wide,
narrowed above to the long and slender, often subfalcate, caudate-
acuminate apex and below to the obtuse, usually slightly in-
equilateral base, the margins serrate-dentate, the teeth distinct,
gland-tipped, the upper surface usually olivaceous when dry,
eglandular, with scattered, short simple hairs, the lower surface
usually somewhat paler, sparingly stellate-tomentose, minutely
glandular; lateral nerves about 10 on each side of the midrib, dis-
tinct, curved-ascending, anastomosing; petioles densely stellate-
tomentose, 5 to 8 mm long. Cymes axillary, solitary, peduncled,
dichotomous, 2 to 4 cm long, rather lax and open, many-flowered,
the branches divaricate, rather densely pubescent with simple
and stellate hairs intermixed. Flowers lilac, 4-merous, small,
the bracteoles linear, pubescent, small. Calyx about 1 mm long,
sparingly hirsute-pubescent with short, straight, simple hairs,
obscurely and subequally 4-toothed. Corolla-tube about 2 mm
long, glabrous, the lobes 4, orbicular-ovate, rounded, about 1
mm long. Filaments 4 mm long; anthers 0.5 mm long. Style
slender, 5.5 mm long, slightly thickened near the apex.

MINbDOoRO, Mount Halcon, Merrill 5556. Ticao, For. Bur. 2534 Clark.
Cresu, Bur. Sci. 11078 Ramos, March, 1912 (type). MINDANAO, District
of Lanao, Camp Keithley, Mrs. Clemens s. n.: District of Zamboanga, Bur.
Sci. 13802 Fénix, Merrill 8115, Bur. Sci. 11802 Robinson, Williams 2807.
BASILAN, Hallier s. n., DeVore & Hoover 41.

This species has been confused with Callicarpa caudata Maxim., and
C. longifolia Lam., and is manifestly allied to the former, differing in its
very different indumentum. It is apparently more closely allied to C. steno-
phylla Merr., than to C. caudata, but is distinguished from the former by
its broader leaves. Among the extra-Philippine forms it is apparently
most closely allied to Callicarpa longifolia Lam., differing in its indu-
mentum, shape of its leaves, and in details of its flowers.

CALLICARPA MEGALANTHA sp. nov.

Arbor circiter 10 m alta, omnibus partibus plus minusve dense
glandulosis et stellato-plumoso-pubescentibus, indumento obscure
fusco vel griseo-fusco; foliis oppositis, oblongo-ovatis, subcoria-
ceis, acuminatis, integris, basi acutis, usque ad 16 cm longis,
nervis utrinque circiter 9, subtus prominentibus; cymis axillari-
bus, longe pedunculatis, dense multifioris; floribus albis, 6 ad
7 mm longis.

A tree about 10 m high, most parts, except the upper surfaces
of adult leaves more or less yellow-glandular and stellate-plumose
pubescent, the indumentum dark-brown or dark grayish-brown
in color. Branches terete, the yeunger ones more or less com-

72 The Philippine Journal of Science 1915

pressed, yellow-glandular, and the younger parts densely stellate-
plumose pubescent. Leaves oblong to oblong-ovate, subcoria-
ceous, entire, 12 to 16 cm long, 5 to 6 cm wide, about equally
narrowed to the acute base and to the acuminate apex, the
upper surface when young more or less stellate-pubescent, be-
coming nearly glabrous, brownish-olivaceous, slightly shining,
the lower surface paler, with numerous, scattered, pale-yellow,
shining glands, densely stellate-pubescent on the midrib and
nerves, with scattered stellate hairs on the lamina; lateral nerves
about 9 on each side of the midrib, prominent on the lower
surface, anastomosing, curved upward; petioles very densely
stellate-pubescent, 2 to 2.5 cm long. Cymes in the upper axils,
solitary, densely many-flowered, dichotomously branched, 7 to &
em in diameter, the peduncles stout, about 8 cm long, the pe-
duncles, branches, bracts, bracteoles, and calyces densely stellate-
plumose pubescent, the indumentum almost obscuring the
scattered, shining, pale-yellow glands. Flowers white, 6 to 7mm
long, the bracts oblanceolate to spatulate, 6 to 8 mm long, the
bracteoles similar but much smaller. Calyx somewhat funnel-
shaped, equally 4-lobed, 3 mm long, the lobes short, acute. Co-
rolla 6 to 7 mm long, outside sparingly supplied with small,
yellow, shining glands, the lobes 4, subequal, oblong-ovate,
broadly rounded, 3 to 3.5 mm long, externally in the median
portion sparingly stellate-pubescent as well as glandular. Fila-
ments 7 and 8 mm long; anthers ovoid, 1.2 mm long, somewhat
glandular on the back. Ovary ovoid, very densely covered with
small, shining, pale-yellow glands.

LUZON, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19687 McGregor,
February, 1913.

Probably most closely allied to Callicarpa subglandulosa Elm., but
differing from that species in many characters. Callicarpa megalantha
is remarkable for its comparatively large flowers which are indicated by
the collector as being white, a color otherwise unknown or at least very
rare in the genus, its long-peduncled cymes, and its dark-brown or dark
grayish-brown indumentum.

CLERODENDRON Linnaeus

CLERODENDRON PUBERULUM sp. nov.

Species C. intermedio valde affinis, differt omnibus partibus
plus minusve puberulis vel pubescentibus.

A suffrutescent, erect, sparingly branched plant 2 m high or —
less, very similar in appearance to Clerodentron intermedium
Cham. Older stems glabrous or nearly so, 4-angled, the younger
ones distinctly puberulent. Leaves broadly ovate, chartaceous,
10 to 20 cm long, 9 to 17 em wide, acuminate, base with a

x.C.1 Merrill: New or Noteworthy Philippine Plants, XI 73

deep, wide sinus, the basal lobes broadly rounded, margins re-
pand-toothed, the upper surface hispid-hirsute with scattered,
appressed, stiff hairs, the lower surface densely and softly puber-
ulent, the indumentum nearly obscuring the glands; petioles
up to 15 em in length, puberulent. Panicles terminal, narrowly
pyramidal, about 20 cm long, the lower branches about 7 cm
long, all parts puberulent, the bracts narrowly spatulate, 1 to 1.5
em long, the bracteoles linear or subfiliform, short. Flowers
numerons, red, their pedicels slender, 1 to 1.5 em long. Calyx-
lobes pubescent, oblong, obtuse, somewhat unequal, about 3 mm
long, accrescent in fruit and 5 to 6 mm long. Corolla-tube
slender, about 12 mm long, 1 mm in diameter, slightly enlarged
above, sparingly pubescent externally, the lobes somewhat un-
equal, oblong to oblong-obovate, obtuse or rounded, 5 to 7 mm
long. Filaments slender, the exserted parts about 2 cm in length.
Fruit about 5 mm long, 8 mm wide, depressed-globose, of 2 to 4,
1-seeded pyrenes.

MINDANAO, District of Davao, Davao, in open wet places, Copeland, 404
(type), March 9, 1904, DeVore & Hoover 177, April, 1903.

A species manifestly very closely allied to Clerodendron intermedium
Cham., which it greatly resembles in aspect. The chief differences appear
to be in the indumentum, C. intermedium Cham. being entirely glabrous,
the present species rather densely puberulent and pubescent.

PREMNA Linnaeus

PREMNA AREOLATA sp. nov. § Gumira.

Frutex scandens, inflorescentiis exceptis glaber ; foliis oblongis,
usque ad 14 cm longis, chartaceis ad subcoriaceis, nitidis, in-
tegris, breviter acuminatis, basi acutis ad rotundatis, subtus
dense areolatis, nervis utrinque circiter 6; inflorescentiis ter-
minalibus, brunneo-puberulis, multifloris; floribus 4-meris, par-
vis, calycibus obsolete 4-dentatis, corollae lobis late rotundatis,
subaequalibus.

A scandent shrub, glabrous except the inflorescence, the
branches terete, grayish, somewhat lenticellate, the younger ones
dark-colored when dry. Leaves chartaceous to submembrana-
ceous, oblong, entire, 8 to 14 cm long, 3 to 5 cm wide, the
apex shortly acuminate, the base acute to rounded, olivaceous
or somewhat brownish when dry, shining, of about the same
color on both surfaces or the lower one a little paler, the lower
surface very densely areolate, the areolae small, 1 mm or less
in diameter, each enclosed by the ultimate reticulations; lateral
nerves about 6 on each side of the midrib, slender, rather distinct,
somewhat ascending, anastomosing; petioles 1 to 2 cm long.
Inflorescence terminal, many flowered, usually peduncled, up to

74 The Philippine Journal of Science 1915

10 em long and wide, the rachis, branches, branchlets, and bracts
rather densely brown-puberulent, the bracts lanceolate, the
largest ones 5 mm long or less. Flowers 4-merous, white or
nearly so, cymose, sessile. Calyx somewhat puberulent, 2 mm .
long, truncate, obsoletely 2-toothed. Corolla-tube glabrous ex-
ternally, about 2.5 mm long, throat and upper part of the
tube inside densely villous, the lobes 4, subequal, broadly ovate,
rounded, about 1.4 mm long. Filaments slender, 2 to 2.6 mm
long. Style slender, glabrous, 3 mm long, the arms slender,
nearly 1 mm in length.

BASILAN, Bur. Sci. 16179 Reillo, September 6, 1912 (type). MINDANAO,
District of Lanao, Camp Keithley, Mrs. Clemens 604, September, 1906.
Said by Mrs. Clemens to be a vine reaching a length of from 75 to 100
feet, growing in large Ficus trees.

The species is a very characteristic one, distinguishable by its oblong
leaves which are densely areolate on the lower surface, its scandent habit,
and its brown-puberulent inflorescence.

PREMNA WILLIAMSII sp. nov. § Gumira.

Species P. areolatae affinis, differt foliis subtus haud areo-
latis, basi semper acutis, apice acute acuminatis, nervis utrinque
5, petiolo longioribus, floribus paullo majoribus.

A scandent shrub, glabrous except the inflorescence. Branches
terete, gray or brownish, somewhat lenticellate, the younger ones
scarcely darker in color than the mature ones. Leaves oblong,
chartaceous, entire, shining, of the same color on both surfaces,
8 to 14 cm long, 3 to 5 cm wide, narrowed below to the acute
base, the apex sharply acuminate; lateral nerves 5 on each side
of the midrib, curved-ascending, distinct; petioles up to 3 cm
in length. Inflorescence terminal, cymose, brown-puberulent up
to 8 em long and wide, many-flowered. Flowers 4-merous.
Calyx cup-shaped, very obscurely puberulent or nearly glabrous,
somewhat 4-angled, 2.2 mm long, truncate, obscurely and broadly
4-toothed. Corolla-tube 3 mm long, glabrous externally, inside
in the upper part and the throat densely villous, subequally
4-lobed, the lobes broadly ovate, rounded, about 1.5 mm long.
Style slender, glabrous, 5 mm long, the arms about 1 mm in
length.

MINDANAO, District of Zamboanga, Sax River, Williams 2330, the stems
trailing, reaching a diameter of 5 cm, flowers white.

Manifestly very closely allied to the preceding species which it greatly
resembles. The leaves, however, are not at all areolate on the lower
surface, the petioles are longer, nerves fewer, and the flowers slightly
larger.

x61 Merriil: New or Noteworthy Philippine Plants, XI 75

GESNERIACEAE
CYRTANDRA Forster

CYRTANDRA ATROPURPUREA sp. nov. § Aureae.

Frutex erectus, simplex, circiter 2 m altus; foliis confertis,

oblongi-obovatis, usque ad 30 cm longis, sessilibus, coriaceis,
grosse irregulariter serratis, acuminatis, basi valde angustatis,
in siccitate atropurpureis, subtus ad costa nervisque adpresse
pubescentibus, nervis utrinque circiter 13, prominentibus; in-
florescentiis axillaribus, pedunculatis, bracteis numerosis, ob-
longo-ellipticis, usque ad 5 cm longis.
. An erect, unbranched shrub about 2 m high, the stems stout,
the leaves more or less crowded near the summit. Leaves oblong-
obovate, 25 to 30 cm long, about 10 cm wide, sessile, coriaceous,
the apex shortly and rather sharply acuminate, the base much
narrowed, the lower 2 to 4 cm 2 cm wide or less, the margins
coarsely serrate, the larger teeth corresponding to the ends of
the lateral nerves and up to 6 mm long, with intermediate similar
but smaller teeth, the upper surface dark-colored when dry,
smooth, somewhat shining, the lower surface when fresh red,
when dry dark-purple, the midrib, lateral nerves, and primary
reticulations on the lower surface rather densely ferruginous-
pubescent with appressed hairs; lateral nerves about 13 on each
side of the midrib, anastomosing, curved-ascending, very prom-
inent on the lower surface, the reticulations lax. Inflorescence
axillary, the peduncles stout, up to 2 cm long, furfuraceous-
pubescent, bearing at their apices numerous, crowded, large
bracts and apparently numerous, crowded flowers, the bracts
mostly elliptic-oblong to ovate-elliptic, narrowed at both ends,
membranaceous or chartaceous 2.5 to 5 cm long, rather promi-
nently nerved, acuminate, coarsely toothed, the nerves more or
less pubescent. Flowers not seen. Capsules narrowly ellipsoid,
about 12 mm long, narrowed above, glabrous, their pedicels
about 5 mm long.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19852 McGregor,
February 2, 1913.

A species manifestly allied to Cyrtandra rex Kranzl., from which it is
at once distinguished by its colored leaves; from other Philippine species

in this group such as Cyrtandra cumingii Clarke, C. oblongata Merr., C.
alnifolia Kranzl., etc., it differs in its sessile leaves.

CYRTANDRA FERRUGINEA sp. nov. § Cuneatae?
Frutex erectus, omnibus partibus plus minusve dense ferrugi-
neo-pubescentibus; foliis oppositis, longe petiolatis, oblongis ad

76 The Philippine Journal of Science 1915

oblongo-ellipticis, usque ad 20 cm longis, inaequilateralibus, plus
minusve falcatis, acutis vel leviter acuminatis, margine denti-
culatis, basi acutis vel subrotundatis, nervis utrinque circiter
10; inflorescentiis axillaribus, 1- ad 3-floris, usque ad 5 cm longis,
bracteis lanceolatis, liberis; floribus 4 cm longis, calycibus per-
sistentibus, lobis lanceolatis, acuminatis, quam tubo paullo
brevioribus.

An erect shrub or suffrutescent plant apparently sparingly
branched, all parts more or less densely covered with pale-
ferruginous, weak, shining hairs, the younger parts very densely
pubescent. Branches terete, or somewhat compressed when dry.
Leaves opposite, in somewhat unequal pairs, chartaceous, oblong
to oblong-elliptic, 12 to 20 cm long, 2 to 12 cm wide, inequi-
lateral, somewhat falcate, apex acute or slightly acuminate,
margins distantly denticulate, base acute or rounded, very in-
equilateral, the lamina blunt or usually rounded and extending
farther on one side than on the other which is acute; lateral
nerves about 10 on each side of the midrib; petioles 4 to 7 cm
long, those of the smaller leaves of each pair shorter than those
of the larger ones. Inflorescence axillary, solitary, few-flowered,
5 cm long or less, the flowers 1 to 3; bracts few, lanceolate,
acuminate, free, 1 cm long. Calyx in anthesis 1.5 cm long, in
fruit 2 cm long, persistent, the lobes 5, lanceolate, acuminate,
somewhat shorter than the tube, the pedicels 1 to 1.5 em long.
Corolla white, 4 cm long, the tube narrow, the lobes rounded.
Fruit lanceolate, including the persistent slender style about
3 cm long, slightly pubescent, the fruit proper about as long
as the calyx, the style exserted.

Luzon, Province of Camarines, Mount Cauayan, Phil. Pl. 1548 Ramos,
December 9, 19138.

Not closely allied to any Philippine form known to me, well character-

ized by its rather uniformly dense ferruginous pubescence, its opposite
inequilateral leaves, few flowers, narrow free bracts, and persistent calyces.

CYRTANDRA KRANZLINII sp. nov. § Campanulaceae ?

Suffrutex erectus vel adscendentibus, simplex, usque ad 50
em altus, plus minusve ferrugineo-villosus; foliis oppositis, cre-
natis, in paribus valde inaequalibus, majoribus oblongo-obovatis,
apice rotundatis, usque ad 18 cm longis, minoribus subellipticis,
1.5 ad 4 em longis; floribus axillaribus, solitariis, longe pedicel-
latis, albis, circiter 5 cm longis, anguste campanulatis.

An erect or ascending undershrub, simple, 30 to 50 cm high,
the stems, petioles, lower surfaces of the leaves on the midrib
and lateral nerves densely covered with long, weak, ferruginous,
jointed hairs. Leaves opposite, firmly chartaceous, slightly cre-

x%,¢,1 Merrill: New or Noteworthy Philippine Plants, XI ae

nate, the upper surface glabrous, dark-olivaceous, the lower much
paler, in very unequal pairs; larger leaf of each pair oblong-
obovate, 12 to 18 cm long, 5 to 8 cm wide, the apex usually
broadly rounded, narrowed below to the acute or obtuse, usually
strongly inequilateral base; primary lateral nerves about 7 on
each side of the midrib, prominent on the lower surface, irreg-
ular, the reticulations lax; petioles densely villous, 12 mm long
or less: smaller leaves of each pair mostly elliptic, 1.5 to 4 cm
long, 0.8 to 2 em wide, shortly petioled, slightly acuminate.
Flowers white, axillary, solitary, ebracteolate, the pedicels
densely villous, slender, about 2 cm long. Calyx 2 cm long, sub-
persistent, narrowly campanulate, villous-pubescent with scat-
tered, long, ferruginous, jointed hairs, the lobes ovate-lanceolate,
slenderly caudate-acuminate, about 8 mm long. Corolla nar-
rowly tubular-campanulate, white, about 5 cm long, slightly
pubescent externally, the tube (flattened) about 4 mm wide for
the lower 1.5 cm, then gradually widened and about 2 cm wide
at the throat, the lobes broadly rounded, comparatively short.
Disk glabrous, truncate, 2 mm high. Stamens included, the fila-
ments of both the fertile stamens and the staminodes somewhat
spirally twisted; anthers narrowly oblong, 3 mm long. Ovary
puberulent; style somewhat hirsute; stigma obliquely oblong, 3
mm long. Capsule (immature) cylindric, about 3 cm long, 3.5
mm in diameter, rostrate, rugose, surrounded at the base by the
persistent calyx.

PALAWAN, Malampaya Bay, Binaloan, Merrill 7247, September, 1910,
in forests at sea level, on damp shaded banks.

A very characteristic species, distinguished by its very unequal leaves,
its villous, ferruginous indumentum, and its solitary, axillary, long-pedi-

celled, comparatively large flowers. The flowers are frequently borne in
the axils of fallen leaves slightly above the surface of the ground.

CYRTANDRA LONGIPEDUNCULATA sp. nov. § Polynesiae.

Frutex subglaber, ramulis junioribus inflorescentiisque parce
pubescentibus; foliis oppositis, lanceolatis ad oblongo-oblanceo-
latis, in paribus leviter inaequalibus, usque ad 10 cm longis,
glabris, acuminatis, basi acutis, margine distanter serratis vel
denticulatis, nervis utrinque 5 vel 6, prominentibus; inflorescen-
tiis axillaribus, solitariis, umbellatis, paucifloris, laxis, quam folia
brevioribus, longe graciliterque pedunculatis; floribus albis, cir-
citer 1.8 cm longis.

An erect shrub, glabrous except the younger branchlets and
the inflorescence. Branches and branchlets slender, light-gray,
terete, or when dry more or less ridged or striate, the younger
parts minutely pubescent with short, brown, appressed hairs.

78 The Philippine Journal of Science 1915

Leaves opposite, in somewhat unequal pairs, chartaceous or sub-
coriaceous, lanceolate to oblong-oblanceolate, 4 to 10 em long,
1 to 3 cm wide, narrowed above to the rather long-acuminate
apex and below to the acute or cuneate base, the margins dis-
tantly and coarsely serrate to merely denticulate, the upper sur-
face smooth, shining, brownish-olivaceous, the lower paler, often
brownish, when young somewhat pubescent, becoming entirely
glabrous; lateral nerves 5 or 6 on each side of the midrib, prom-
inent on the lower surface, anastomosing, the reticulations lax,
prominent; petioles of the larger leaves about 5 mm long, of
the smaller ones somewhat shorter. Inflorescence very slender,
axillary, solitary, few-flowered, shorter than the leaves, the
flowers umbellately disposed at the apices of the simple peduncles.
Peduncles very slender, 3.5 to 5.5 cm long, the bracts lanceolate,
4 mm long, the pedicels 1 to 1.5 cm long. Flowers few, white,
but one or two opening at the same time in each umbel. Calyx
6 mm long, glabrous or nearly so, ovoid, the tube 3 mm long,
the lobes ovate-lanceolate, 3-nerved, long-acuminate, more or less
spreading, about 3 mm long. Corolla glabrous, narrowly cam-
panulate, about 1.8 cm long, slightly widened upward, 2-lipped,
the upper lip with 2, broadly ovate, rounded lobes about 3 mm
long, the lower lip with 3 lobes, similar to those of the upper
lip but slightly larger. Disk cupular, truncate, glabrous, 2 mm
high. Stamens 2; filaments slightly pubescent; anthers 1 mm
long, connate. Ovary narrowly ovoid, glabrous, tapering up-
ward into the style which is glabrous below, and pubescent in
the upper one-half or two-thirds.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19664 (type),
19796 McGregor, February 8, 1913.

A species similar to and manifestly closely allied to Cyrtandra plectran-

thiflora Kranzl., from which it differs in its differently shaped, glabrous,
smaller, fewer-nerved leaves and its larger flowers.

CYRTANDRA OBLONGATA sp. nov. § Aureae.

Frutex erectus, 2 ad 3 m altus, plus minusve pubescentibus;
foliis oblongis, coriaceis, usque ad 25 cm longis, acuminatis, base
longe angusteque decurrentibus, margine irregulariter serratis,
supra glabris, subtus dense ferrugineo-pubescentibus; nervis
utrinque 14 ad 16, adscendentibus, subtus valde prominentibus;
inflorescentiis axillaribus, sessilibus, floribus confertis, circiter
3 cm longis, bracteis involucrantibus albis, usque ad 3 em longis,
anguste obovatis vel elliptico-ovatis.

An erect, more or less branched shrub 2 to 3 m high, the
branches stout, terete, or the younger ones more or less angled.

x%,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 79

Leaves somewhat crowded toward the ends of the branchlets,
oblong, coriaceous, 18 to 25 cm long, 4 to 8 cm wide, the upper
surface glabrous, dark-olivaceous, smooth, the lower much paler,
the apex shortly but sharply acuminate, the base long and
narrowly decurrent, winging the stout, 3 to 6 cm long petiole,
the margins rather coarsely and irregularly serrate; nerves 14
to 16 on each side of the midrib, curved-ascending, very prom-
inent on the lower surface, and with the midrib and the primary
reticulations densely lanate-pubescent with brown hairs. Inflo-
rescence axillary, sessile, the flowers crowded, shortly pedicelled,
subtended by an involucre of large bracts. Bracts numerous,
white, narrowly obovate to ovate-elliptic, membranaceous, up to
3 cm long and 2 cm wide, acuminate, coarsely toothed, sessile,
prominently veined, the reticulations lax, slightly pubescent.
Calyx about 18 cm long, outside appressed ferruginous-lanate,
the upper 1 cm divided into 5, ovate-lanceolate, long and slender-
ly acuminate lobes. Corolla white, narrowly campanulate, 3
cm long, the two lobes of the upper lip about 6 mm long,
orbicular-ovate, rounded, the cleft between them narrow, 5 mm
deep, the three lobes of the lower lip broadly ovate, rounded,
about 10 mm long. Stamens 4; anthers about 2.5 mm long.
Ovary narrowly ovoid, glabrous, 4 to 5 mm long; style pubescent
below the stigma. Capsule narrowly ellipsoid, about 12 mm
long.

LUZON, Provinces of Tayabas and Laguna, Mount Banajao, Merrill 7515
(type), February, 1911, Whitford 931, 1008, October, 1904: without definite
locality, Loher 6650. Merrill 5578, from Mount Halcon, Mindoro, a very
poor specimen, is also apparently referable here.

This species has previously been confused with Cyrtandra cumingii
Clarke, to which it is allied, but from which it differs in many charaeters,
the shape and pubescense of its leaves, and the elongated, narrowly winged
petioles. It is apparently closer to Cyrtandra rex Kranz). than to C. cum-
ingu Clarke, but is very different from Kranzlin’s species.

CYRTANDRA NANA gp. nov. § Brevicaules.

Planta parva, glabra, erecta, simplex, circiter 4 cm alta; foliis
oppositis et ternis, sessilibus, oblongis, chartaceis, acutis, serratis,
aequalibus, circiter 2 cm longis, 4 mm latis; floribus axillaribus,
solitariis, circiter 1 cm longis, calycibus alte 5-partitis.

A small, glabrous, erect, unbranched plant about 4 cm high.
Leaves rather crowded, the internodes short, mostly ternate or
some opposite, equal, sessile, chartaceous, oblong, about 2 cm
long, 4 mm wide, acute, base narrowed-decurrent, margins
sharply serrate, the nerves obscure. Flowers apparently pur-

80 The Philippine Journal of Science 1915

plish, solitary, axillary, their pedicels about 7 mm long, the
bracts small or wanting. Calyx divided nearly to the base into
5, linear-oblong, obtuse lobes, the lobes about 6 mm long, 0.7
mm wide. Corolla narrowly funnel-shaped, somewhat enlarged
upward, obscurely 2-lipped, the lobes broadly rounded, quite
glabrous, 1 cm long. Ovary narrowly oblong, glabrous, 1.5
mm long, narrowed upward into the 5 mm long style. Stamens
included, the staminodes inserted higher up than the stamens.

MINDANAO, Bukidnon Subprovince, near Tangcolan, Bur. Sci. 21462
Escritor, August 2, 1918.

A species well characterized by its very small size, the entire plant
not exceeding 4 cm in height. Other striking characters are its solitary,
pedicelled flowers, and its sessile ternate and opposite sessile leaves.

DICHROTRICHUM Reinwardt

DICHROTRICHUM PAUCIFLORUM sp. nov.

Suffrutex epiphyticus, scandens, plus minusve setoso-hirsutus;
foliis longe petiolatis, ellipticis, usque ad 18 cm longis, 7 cm
latis, basi acutis, apice breviter obtuse acuminatis, nervis utrin-
que circiter 7; inflorescentiis axillaribus, brevibus, paucifloris;
floribus circiter 3 cm longis, extus setoso-hirsutis.

An epiphytic, sparingly branched suffrutescent vine, the stems
terete, somewhat fleshy when fresh, glabrous, emitting numerous
roots along the side next to the supporting plant, glabrous,
light-gray when dry, the apical parts more or less setose-hirsute.
Leaves elliptic, 10 to 13 cm long, 5.5 to 7 cm wide, chartaceous,
olivaceous, the base acute, the apex shortly and obtusely acum-
inate, margins irregularly and doubly toothed, the teeth small,
both surfaces with more or less scattered, long, stiff, white,
jointed, setose-hirsute hairs especially along the midrib and
lateral nerves, similar hairs also on the petioles and the in-
florescence; lateral nerves about 7 on each side of the mid-
rib, rather prominent on the lower surface, curved-ascending ;
petioles 4 to 6 cm long, when young densely setose-hirsute,
less so when old. Inflorescence axillary, few-flowered, setose-
hirsute, including the prominent flowers 4 cm long or less, the
flowers subfascicled or arranged on a very short peduncle.
Flowers up to 5 in each axil, but one or two opening at the same
time, red, their pedicels 3 to 4 mm long, setose. Sepals 5,
oblong-obovate to oblong-oblanceolate, 1.2 to 1.5 cm long, 3 to 4
mm wide, obtuse or acute, irregularly and rather prominently
toothed, dorsally setose-hirsute with long stiff hairs. Corolla
about 3 cm long, somewhat curved, externally somewhat setose-
hirsute, the mouth very oblique, lobes broadly rounded. Fila-

rs c,1 Merrill: New or Noteworthy Philippine Plants, XI 81

ments exserted 6 to 10 mm. Ovary glabrous; style spatulate.
Disk prominent, glabrous, about 1.5 mm in diameter.

PALAWAN, Mount Capoas, Malampaya Bay, Merrill 9497, April 21, 1918,
epiphytic in the mossy forest, altitude about 950 meters.

A species readily distinguished by its very short inflotescence, in this
character approaching D. asperifolium Benth. & Hook. f., and D. brevipes
Clarke, from both of which, however, it is very distinct.

TRICHOSPORUM D. Don

TRICHOSPORUM BRACHYSEPALUM sp. nov.

Frutex epiphyticus, scandens, glaber, tenuibus; foliis oppos-
itis, coriaceis, oblongo-lanceolatis, usque ad 8 cm longis, acum-
inatis, nervis lateralibus obsoletis; floribus axillaribus, solitariis,
pedicellatis, calycis usque ad basin divisi, segmentis 2 mm
longis; corolla 2.2 em longa, curvata, staminibus exsertis, fila-
mentis pubescentibus. .

An epiphytic, glabrous, slender, branched vine, the stems and
branches rather slender, terete, light-gray, the branchlets often
longitudinally sulcate or striate when dry. Leaves opposite,
oblong-lanceolate, fleshy when fresh, coriaceous and wrinkled
when dry, 6 to 8 cm long, 1.5 to 2 em wide, narrowed below to
the acute base, and above to the sharply and usually slenderly
acuminate apex, the lateral nerves obsolete or nearly so; petioles
2 to 5 mm long. Flowers axillary, solitary, their pedicels
slender, glabrous, about 1 cm long. Calyx consisting of 5, small,
ovate-lanceolate, acuminate, 2 mm long, glabrous lobes. Corolla
tubular, curved, base broadly rounded, glabrous, 2 to 2.2 cm
long, about 4.5 mm in diameter, the throat somewhat oblique,
limb 2-lipped, 2 lobes somewhat larger than the other 3, the lobes
orbicular, rounded, about 4 mm in diameter, their margins with
few, scattered, stiff hairs. Disk glabrous, truncate, 0.5 mm in
diameter. Ovary glabrous; style pubescent. Stamens exserted
about 1 cm; filaments pubescent; anthers connate, about 2 mm
long.

BASILAN, Cumalarang, Bur. Sci. 16163 Reillo, August 25, 1912.

A species well characterized by its greatly reduced calyx which consists
merely of 5, ovate-lanceolate, acuminate, 2 mm long lobes.

TRICHOSPORUM MACGREGORII sp. nov. § Haplotrichium.

Species 7. rizalense affinis, differt floribus majoribus, 3.5 em
longis, corolla deorsum haud angustata, lobis majoribus (circiter
6 mm diametro), foliis tenuiter acuminatis. ;

An epiphytic vine, glabrous except the inflorescence, the
branches light-gray, terete, the branchlets slender. Leaves

1312816

82 The Philippine Journal of Science 1915

oblong-ovate to ovate-lanceolate, chartaceous or subcoriaceous
when dry, usually olivaceous, 3.5 to 5 cm long, 1.3 to 1.6 cm
wide, the base acute, the apex slenderly and sharply acuminate;
lateral nerves 3 or 4 on éach side of the midrib, ascending, very
slender, often obscure; petioles about 2 mm long. Flowers
solitary, axillary, their pedicels rather densely hirsute, about 6
mm long. Calyx cleft to the base into 5, narrowly lanceolate,
acuminate or blunt lobes which are somewhat hirsute, about 7
mm long, 1.2 mm wide, 3-nerved. Corolla red, somewhat curved
and ventricose, 3.5 cm long, hirsute outside, the mouth somewhat
oblique, the lower 3 lobes broadly ovate, rounded, about 6 mm
in diameter, the upper two about 4 mm in diameter. Disk
glabrous, truncate, about 2 mm long. Ovary glabrous; style
hirsute. Anthers 2 mm long, connate, included, the filaments as
long as the corolla-tube.

LuzOoN, Province of Nueva Vizcaya, near Imugan, Bur. Sci. 20075
McGregor, January, 1913.

Manifestly closely allied to Trichosporum rizalense, which it closely
resembles, differing in the indicated floral and vegetative characters.
TRICHOSPORUM RIZALENSE sp. nov. § Haplotrichium.

Species JT. philippinense simillima et affinis, differt floribus
majoribus, circiter 3 cm longis, extus parce hirsutis, nervis
lateralibus vix obsoletis, tenuibus, adscendentibus.

A scandent, slender, epiphyte, glabrous except the inflores-
cence. Branches and branchlets slender, terete, mostly light
gray. Leaves opposite, when fresh fleshy, when dry coriaceous,
elliptic-ovate, 3.5 to 4 cm long, 1.5 to 2 cm wide, the base acute,
the apex acute or slightly acuminate, rather pale when dry;
lateral nerves slender, ascending, about 4 on each side of the
midrib, sometimes indistinct; petioles about 3 mm _ long.
Flowers solitary, axillary, their pedicels slender, hirsute, 5 to
6 mm long. Calyx cleft to the base into 5, lanceolate, obtuse,
3-nerved lobes 5 to 6 mm long, about 1.4 mm wide, sparingly
hirsute. Corolla red, somewhat curved, 3 cm long, sparingly
hirsute outside, narrowed below, the mouth somewhat oblique,
2-lipped, lobes of the upper lip broadly rounded-ovate, about 4
mm in diameter, of the lower lip about 5 mm in diameter. Disk
glabrous, truncate, 1.5 mm long. Ovary glabrous; style hirsute.
Anthers included, connate in pairs, 2 mm long, the filaments as
long as the corolla.

Luzon, Province of Rizal, Mount Canumay, Bur. Sci. 18775 Ramosp
October, 1911.

In general appearance greatly resembling Trichosporum philippinense,
differing however, among other characters, in its larger hirsute flowers.

_

x,¢,1 Merrill: New or Noteworthy Philippine Plants, XI 83

TRICHOSPORUM MALINDANGENSE sp. nov. § Haplotrichium ?

Frutex epiphyticus, scandens, gracile, inflorescentiis exceptis
glaber ; foliis coriaceis, anguste lanceolatis vel oblongis, glabris,
circiter 7 cm longis, integris, nervis lateralibus obsoletis; caly-
cibus tubulosis, breviter dentatis, extus parce capitellato-hirsutis,
2 ad 2.5 cm longis; corolla 4 cm longa, cylindrica, extus parce
hirsuta.

An epiphytic, scandent shrub, the branches rather stout,
terete, pale-brown when dry, very slightly pubescent or subglab-
rous. Leaves opposite, lanceolate to narrowly oblong, thickly
coriaceous, pale and more or less wrinkled when dry, 6 to 7 cm
long, 1.5 to 2 cm wide, narrowed above to the blunt apex, the
base acute, somewhat shining, the midrib prominent, the nerves
and reticulations obsolete; petioles 5 to 7 mm long. Flowers
axillary, the pedicels 6 to 8 mm long, solitary or 2 to 4 fascicled
in the upper axils. Calyx tubular, 2 to 2.5 cm long, 6 to 8 mm
in diameter, slightly inflated in the middle, 5-toothed, the teeth
short, acute, outside with scattered, short, white, capitate hairs.
Corolla red, about 4 cm long, narrowly tubular, the limb some-
what spreading, irregularly 5-lobed, the lobes ovate, rounded,
about 6 mm in diameter. Stamens 4, included or slightly ex-
serted; anthers about 3mm long. Disk glabrous. Ovary nearly
as long as the corolla, rather densely capitellate-hirsute.

MINDANAO, Province of Misamis, Mount Malindang, For. Bur. 4660
Mearns & Hutchinson, May, 1906, in forests, altitude about 1,000 m.

TRICHOSPORUM MINDORENSE sp. nov. § Haplotrichium.

Species 7. philippinense affinis, differt pedunculis unifloris,
floribus brevioribus latioribusque, circiter 2 cm longis,

An epiphytic, scandent shrub, quite glabrous, the branches
slender, terete, light-gray, 2 to 3 mm in diameter. Leaves fleshy
when fresh, when dry coriaceous, pale-brownish, dull or slightly
shining, oblong-ovate to oblong-lanceolate, 4 to 6 cm long, 1.5 to
2 em wide, entire, the base acute, the apex long and slenderly
subcaudate-acuminate, the midrib distinct, the lateral nerves and
reticulations obsolete; petioles about 2 mm long. Flowers scar-
let, axillary, solitary, their pedicels about 1 cm long, jointed in
the middle, with a pair of linear, obtuse, 4 mm long bracts near
the joint. Calyx cleft to its base into linear, obtuse, 4 to 5
mm lobes. Corolla scarlet, 2 cm long, glabrous, slightly curved,
' when flattened about 1 cm wide at the throat, somewhat ven-
tricose, the mouth somewhat oblique, not contracted, the three
lower lobes rounded-reniform, about 7 mm in diameter. Stamens

2

@..F

84. The Philippine Journal of Science»

included; anthers connate in pairs, 2’mm long. Disk 2 mm
high, truncate, about 1.5 mm in diameter.

MINDoRO, Mount Halcon, Merrill 6141, November, 1906 (type), For.
Bur. 4879 Merritt, June, 1906, altitude about 1,300 m; Mount Sablayan,
For. Bur. 9761 Merritt, March, 1908, altitude about 1,000 m.

A species manifestly allied to Trichosporum philippinense 0. Ktze.,
with which the three specimens above cited have previously been confused,
and which they greatly resemble, especially in vegetative characters. The
flowers, however, are very different, relatively much shorter and broader,
while the peduncles bear but a single flower, both the peduncles and
pedicels being longer in the present species than in T. philippinense O. Ktze.

TRICHOSPORUM IRIGAENSE sp. nov.

Frutex epiphyticus, glaber, ramulis teretibus; foliis oppositis,
petiolatis, coriaceis, oblongo-lanceolatis vel ovato-lanceolatis,
nitidis, usque ad 9 cm longis, basi acutis, apice longe tenuiterque
acuminatis, nervis lateralibus numerosis, densis, tenuibus, ad-
scendentibus; floribus axillaribus, solitariis, pedicellatis, circiter
3 cm longis; sepalis oblongis, acutis, circiter 1 cm longis, 3 ad
4 mm latis; corolla glabra, leviter inflata, plus minusve curvata.

An epiphytic plant, sparingly branched, quite glabrous, the
branches terete, apparently somewhat fleshy, when dry -2 to 3
mm in diameter. Leaves opposite, coriaceous, oblong-lanceolate
to ovate-lanceolate, 6 to 9 cm long, 2 to 3 cm wide, base acute
or somewhat rounded, apex long and slenderly acuminate,
straight or slightly falcate, the upper surface olivaceous, shining,
the lower paler; lateral nerves 10 or more on each side of the
midrib, slender, densely arranged, sharply ascending, about
equally evident on both surfaces; petioles 10 to 12 mm long.
Flowers axillary, solitary, apparently red, quite glabrous, about
3 cm long, their pedicels about as long as the petioles. Calyx
lobes nearly free, oblong, acute or obtuse, about 1 cm long,
3 to 4 mm wide, narrowed at both ends. Corolla somewhat
inflated, slightly curved; when flattened out about 7 mm in
diameter, enlarged upward.

Luzon, Province of Camarines, Mount Iriga, on trees at the summit,
altitude about 1,200 m, Bur. Sci. 22211 Ramos, December 3, 1913.

The alliance of this species seems to be with TRICHOSPORUM FOX-

WORTHY! (Krinzl.) Aeschynanthus foxworthyi (Kranzl.), but is suf-
ficiently distinct. The slender ascending nerves are characteristic.

[Vol. IX, Sec. C, No. 6, including pages 495 to 563, was issued March
20, 1915.]

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C. BOTANY
VoL. X SEPTEMBER, 1915 No. 5

NEW OR NOTEWORTHY PHILIPPINE PLANTS, XIl

By E. D. MERRILL *

(From the Botanical Section of the Biological Laboratory, Bureau of
Science, Manila, P. I.)

The preceding number of this series was published in the early
part of the present year,? and the present contribution is essen-
tially like its predecessors. Seven genera are for the first time
credited to the Archipelago, Avena, Polytoca, Angelesia, Glypto-
petalum, Ochrocarpus, Asystasia, and Polytrema. Sixty-two
new species are described in various families, while 18 species
previously described from extra-Philippine material are for
the first time credited to the Archipelago. The total additions
to the Philippine flora in the present paper are eighty species.
Some reductions are made, and the synonymy of some species is
discussed, resulting in a few cases in the substitution of older
specific names for those in more general use. All proposed
changes in nomenclature are in accordance with the rules of the
International Botanical Congress.

GRAMINEAE
AVENA Linnaeus

AVENA FATUA L. Sp. Pl. (1753) 80; F.-Vill. Novis. App. (1880) 319;
Hook. f. Fl. Brit. Ind. 7 (1897) 275.

LUZON, Benguet Subprovince, Bur. Sci. 2812 Mearns, April, 1907.

This European species is now widely distributed in Asia, northern Africa,
and North America, a weed of cultivation. It has previously been recorded
from the Philippines by F.-Villar, but his record has not before been verified.
It is apparently a casual plant here and may or may not persist.

* Associate professor of botany, University of the Philippines.
* Philip. Journ. Sci. 10 (1915) Bot. 1-84.
133910 : 287

288 The Philippine Journal of Science 1915

CHLORIS Swartz

CHLORIS INCOMPLETA Roth. Nov. Sp. Pl. (1821) 60; Hook. f. Fl. Brit.
Ind. 7 (1897) 290.

MINDANAO, District of Zamboanga, Merrill 8279, December 6, 1911,
along roadsides near Calarian, and on dry hills back of Zamboanga, ascend-
ing to at least 200 meters altitude. PALAWAN, Taytay, Merrill 9275, sandy
seashore; Silanga, Merrill 9616, Phil. Pl. 1308 Merrill, open places near the
seashore. LUZON, Province of Zambales, Gates.

Not previously reported from the Philippines, India, Formosa, and
Malaya. In addition to the Philippine specimens, cited above, I have
before me also Henry 1151 from Formosa, and Koorders 21287 from Java.
Chloris mearnsii Merr., from Luzon, is allied, but is much smaller, with
much narrower leaves, fewer, slenderer, and shorter spikes, and smaller
spikelets.

POLYTOCA R. Brown

POLYTOCA HETEROCLITA (Roxb.) comb. nov.

Coix heteroclita Roxb. Fl. Ind. ed. Carey 3 (1882) 572.
Polytoca bracteata R. Br. in Benn. Pl. Jav. Rar. (1838) 20, t. 5; Hook.
f. Fl. Brit. Ind. 7 (1897) 101.

MINDANAO, Bukidnon Subprovince, Tongkulan, C. F. Baker 3616, June,
1914.

The first representative of the genus to be found in the Philippines, here
recorded under its oldest specific name; India to Burma, Tonkin, and Java.

CYPERACEAE
CLADIUM Schrader

CLADIUM RIPARIUM (Nees) Benth. Fl. Austral. 7 (1878) 405; C.
B. Clarke Ill. Cyp. (1909) t. 85, fig. 5-8.

Chapelliera riparia Nees in Lehm. PI. Preiss. 2 (1846-48) 76.
Baumea riparia Boeckl. in Linnaea 38 (1874) 246.

MINDANAO, Lake Lanao, Camp Keithley, Mrs. Clemens 916, January,
1907, in swamps.

In attempting to identify the above specimen, it was suspected that it
might represent the species described by Gaudichaud as Baumea glomerata,
and accordingly a duplicate was sent to the Museum of Natural History,
Paris, for comparison with Gaudichaud’s type. Dr. E. G. Camus, who
kindly made the comparison, reports that it is not Gaudichaud’s species,
but is manifestly the same as Cladium riparium Benth., in which identifica-
tion I concur after studying the description of Bentham’s species and
fragments supplied me by Doctor Camus for comparison. A _ distinct
Australian type.

ARACEAE

ACORUS Linnaeus

ACORUS GRAMINEUS Soland. in Ait. Hort. Kew 1 (1789) 474; Engl.
Pflanzenreich 21 (1905) 312.

LuzON, Province of Laguna, near San Antonio, on bowlders in stream
beds, Bur. Sci. 14949 Ramos, June 20, 1912.

Bie, 6 Merrill: Philippine Plants, XII 289

This species has not previously been reported from the Philippines.
Widely distributed in central and in temperate eastern Asia, extending from
Sikkim and Tibet to Japan, southward to Burma, southern China, and
Formosa. It is known from no part of Malaya and must be considered a
northern type in the Philippine flora.

The commoner and widely distributed Acorus calamus L. occurs in
the Philippines as an introduced and cultivated plant, but is thoroughly
spontaneous in some localities in Mountain Province, Luzon.

AMORPHOPHALLUS Blume

AMORPHOPHALLUS LUZONIENSIS sp. nov. § Conophallus.

Tuber parvum, depresso-globosum, in siccitate circiter 2 cm
diametro; foliis umbraculiformibus, tripartitis, segmentis cir-
citer 20 cm longis, pinnatisectis, segmentis utilimis oblongis vel
oblongo-ovatis, caudato-acuminatis, 5 ad 9 cm longis; petiolo
circiter 50 cm longo; pedunculis tenuibus, 10 cm longis; spathis
circiter 15 cm longis, 6 cm latis, purpureis; spadicis quam spatha
paulo brevioribus, cylindraceis, supra sensim caudato-angustatis;
stilus quam ovario triplo longior.

Tubers depressed-globose, when dry about 2 cm in diameter,
emitting many roots. Petioles slender, about 50 cm high; lamina
umbraculiform, 3-partite, the segments pinnatisect, about 20 cm
long, membranaceous, the ultimate segments oblong to oblong-
ovate, caudate-acuminate, 5 to 9 cm long, nearly the same color
on both surfaces or slightly paler beneath, the sinuses acute,
extending almost or quite to the rachis. Sheaths of the peduncles
membranaceous, brown when dry, the outer ones oblong-lanceo-
late, 2 to 3 cm long, the inner ones lanceolate up to 10 cm in
length; peduncles slender, up to 10 cmin length. Spathes purple,
about 15 cm long, thin, the tube broad, about 6 cm long, the lamina
ovate, slightly acute-acuminate or merely acute, about 6 cm wide.
Spadix slender, cylindric, a little shorter than the spathe, the
pistillate part about 2.5 cm long, 1 cm wide, the staminate part
contiguous, about 3 cm long and 8 mm wide, the appendage
slender, continuous, verruculose, caudate, at the base 5 mm in
diameter or less, gradually narrowed upward, the entire spadix
about 14 cm long. Pistillate flowers: Ovary depressed, about
1 mm long, 1.5 mm wide, 1-celled, 1-ovuled; style slender, about
3 mm long; stigma ovoid, entire, somewhat oblique, about 1 mm
long. Staminate flowers crowded, mostly 2-androus, the anthers
subglobose, 1 mm long, 1.5 mm wide, opening by two terminal
slits.

Luzon, Province of Cagayan, Abulug River, For. Bur. 19560 Curran
(type), January, 1912, Weber s. n., same locality and date.

A species growing in crevices of limestone ledges, quite different from
any other known Philippine form. Apparently through its elongated styles

290 The Philippine Journal of Science 1915

allied to both Amorphophallus longistilus Kurz, of the Andaman Islands,
and to A. harmandii Engl. & Gehrm., of Cambodia, but differing from
both in many characters. It is at once distinguished from the former by
its much smaller spathes, and from the latter by its larger spathes and
relatively much longer spadices.

AMORPHOPHALLUS DECURRENS (Blanco) Kunth Enum. 3 (1841) 581;
Engl. in DC. Monog. Phan. 2 (1879) 319, Pflanzenreich 48 (1911)
108 = AMORPHOPHALLUS CAMPANULATUS (Roxb.) Blume!

Kunth’s Amorphophallus decurrens was based on Arum decurrens Blanco
Fl. Filip. (1837) 656, and is retained by Engler, in his recent monograph
of the group, among the doubtful and imperfectly known species. In all
characters Blanco’s description applies very closely to Blume’s species,
which is very common and widely distributed in the settled areas throughout
the Archipelago. There is no reason whatever for doubting that Arum
decurrens Blanco, that is, Amorphophallus decurrens Kunth, is identical
with the older Amorphophallus campanulatus (Roxb.) Blume.

ERIOCAULONACEAE
ERIOCAULON Linnaeus

ERIOCAULON NIGRICEPS sp. nov.

Planta dense casepitosa, inflorescentiis exceptis glabra; caulis
brevibus, crassis; foliis numerosis, lanceolatis, usque ad 6 cm
longis, basi brevissime vaginantibus; pedunculis numerosis, 5
ad 18 cm longis, gracilis, leviter tortis, 5-costatis; capitulis sub-
globosis, circiter 5 mm diametro, in siccitate nigris, bracteis al-
bido-ciliatis; floribus ¢ sepalis 3, connatis; petalis nullis vel 1
ciliato-fissis; antheris nigris; floribus @ exacte trimeris, sepalis
oblongis vel oblanceolatis, breviter acuminatis, apice ciliatis.

A densely czespitose plant, glabrous except the inflorescence.
Stems stout, short. Leaves numerous, densely crowded, lanceo-
late or narrowly lanceolate, 2 to 6 cm long, 5 to 8 mm wide, about
10-nerved, much wider at the base than above, gradually nar-
rowed and usually obtuse. Peduncles numerous, slender, up to
18 em in length, 5-ribbed, slightly twisted. Heads subglobose,
4 to 5 mm in diameter, nearly black or somewhat grayish when
dry, dense, the involucral bracts pale, oblong to obovate, truncate,
about 2 mm long. Receptacle glabrous. Floral bracts dark-
colored, oblong-oblanceolate to oblong-obovate, shortly acum-
inate, distinctly white-ciliate toward the apex on the back. Male
flowers: Calyx about 1.5 em long, dark-colored, spathelike, at
first 3-toothed, the lobes eventually becoming free nearly or quite
to the base, eglandular. Petals none, or one only and ciliate-cleft
into several hyaline-jointed segments. Stamens 6; anthers black.
Center of the flower with 3 minute, dark-colored sessile glands.
Female flowers: Sepals 3, subequal, oblanceolate to oblong, 1.8 to

X, C, 5 Merrill: Philippine Plants, XII 291

2 mm long, obtuse or acute, slightly ciliate at the apex, dark-
colored, not glandular. Petals 3, hyaline, linear-oblanceolate,
about 1.5 mm long, slightly ciliate at the apex, not glandular.
Ovary 3-celled; style-arms 3. Seeds minutely striate.

LuzON, Benguet Subprovince, Trinidad River, Bur. Sci. 5544 Ramos
(type), December, 1908; Baguio, Merrill 7748, May, 1911, Williams 1004,
October, 1904; Lepanto Subprovince, Balili, Merrill 4646, November, 1905.

This species is apparently distinct from all described ones, characterized
especially by the corolla of the male fiowers being reduced to a single
ciliate-cleft petal, consisting of several jointed filiform lobes. It appears
to fall in the section defined by Ruhland under section 5 of his key.

ERIOCAULON ALPESTRE Hook. f. & Th. ex Koern. in Mig. Ann. Mus.
Lugd. Bat. 3 (1867) 163; Ruhl. in Engl. Pflanzenreich 13 (1903) 95.

MINDANAO, District of Davao, Mount Apo, Copeland 1431, October, 1904,
altitude about 1,800 meters, seen in only one place.

Mountains of India to Indo-China, China, and Japan; not previously
reported from the Philippines.

ERIOCAULON CINEREUM R. Br. Prodr. (1810) 254; Benth. Fl. Austral.
7 (1878) 193.

Eriocaulon sieboldianum Sieb. & Zucc. ex Steud. Syn. 2 (1855) 272;
Ruhl. in Engl. Pflanzenreich 13 (1903) 111.

LUZON, Subprovince of Abra, in rice paddies, For. Bur. 16458 Bacani:
Subprovince of Bontoc, in rice paddies, Vanoverbergh 602: Province of
Rizal, Caloocan, Phil. Pl. 298 Merrill (distributed as E. merrillii), Bur. Sci.
. 9501 Robinson (pro parte).

This very widely distributed species does not appear to be common in
the Philippines; at least it has been collected but a few times. I can see
no valid reason for ignoring Robert Brown’s specific name, as it is certainly
valid and antedates the one accepted by Ruhland (E£. seiboldianum) by
forty-five years.

ERIOCAULON MERRILLI! Ruhl. in Perk. Frag. Fl. Philip. (1904) 136.

This species was based on Merrill 572, collected in the Island of Culion.
As described by Ruhland, one of the special characters by which the species
is distinguished from allied forms is the absence of sepals in the female
flowers. This character alone does not appear to be a valid one, for in
the original collection, two sheets of which are in our herbarium, most of
the heads have female flowers with two sepals, while other flowers are
without them. The species, as interpreted by me, is the commonest and
most widely distributed one in the Philippines, and appears to me to be very
closely allied to, if not identical with, Eriocaulon truncatum Ham. I would
refer to it the following specimens:

LUZON, Province of Tarlac, Merrill 3624: Province of Zambales, For.
Bur. 8158 Curran & Merritt: Province of Nueva Ecija, Bur. Sct. 5277
McGregor: Province of Rizal, vicinity of Manila, Merrill 7122, Bur. Sci.
9500, 9501 (pro parte) Robinson, McGregor s. n. POLILLO, Bur. Sci. 9031
Robinson. CUuLIoNn, Merrill 572 (cotype). MINDANAO, Province of Surigao,
Piper 534: District of Lanao, Camp Keithley, Mrs. Clemens 909 (pro parte),
8. Nn.

292 The Philippine Journal of Science 1915

Among the species of this genus credited to the Philippines by various
authors, of which I have seen no specimens originating in the Archipelago,
are the following:

Eriocaulon “longissimum” Nees; Usteri Beitr. Ken. Phil. Veg. (1905)
131, manifestly an error for E. longifolium Nees, as there is no such species
as E. longissimum. I have specimens of E. longifoliwm Nees from Singapore
(coll. Ridley), Labuan (coll. Merrill), Cochinchine (coll. Pierre), and the
Caroline Islands, Yap (coll. Volkens 406). The species is certainly to be
expected in the Philippines.

Eriocaulon sexangulare L. Reported from the Philippines, but without
the citation of specimens, by Miquel Fl. Ind. Bat. 3 (1859) 523, Naves
Novis. App. (1882) 299, and Ruhland in Engl. Pflanzenreich 13 (1903)
110. I have seen no Philippine specimens.

Eriocaulon truncatum Ham.; Naves Novis. App. (1882) 299; Ruhl. in
Engl. Pflanzenreich 13 (1903) 107. The specimen cited by the latter author
(Cuming 2862) was not from the Philippines, but from the Malay Peninsula.
See #. merrillii above.

Eriocaulon setaceum L.; Naves Novis. App. (1882) 299. Probably admit-
ted on an erroneous determination.

COMMELINACEAE
ANEILEMA R. Brown

ANEILEMA AZUREUM sp. nov. Tricarpellaria, Euaneilema.

Caespitosum, plus minusve pubescens praesertim inflorescen-
tiis; foliis numerosis, anguste lanceolatis vel elongato-lanceolatis,
usque ad 25 cm longis, 5 ad 10 mm latis, in siccitate dense minu-
teque puncticulatis, laevis, acutis vel apiculato-acuminatis;
floribus azureis, cymoso-paniculatis, inflorescentiis angustis, con-
fertis, bracteis magnis, spathulatis, pubescentibus, plus minusve
imbricatis; capsulis oblongo-ellipsoideis, trigonis, circiter 6 mm
longis; seminibus angulatis, in quisque loculo 4, uniseriatis.

A tufted, distinctly pubescent plant 20 to 40 cm high, the
roots fibrous, no tuberous ones present on the material ex-
amined. Leaves numerous, narrowly lanceolate to elongate-
lanceolate, chartaceous, 12 to 25 cm long, 5 to 10 mm wide,
when dry minutely and distinctly puncticulate, smooth, not scab-
rous, sparingly pubescent or nearly glabrous, base not narrowed,
somewhat sheathing, apex acute or apiculate-acuminate. Scapes
up to 40 cm in length, simple, leafless, more or less pubescent, the
flower-bearing parts rather densely pubescent. Flowers deep
blue, cymose, the inflorescence narrow. Bract subtending the
first branch about 4 cm long, somewhat sheathing, leaflike, the
upper ones smaller, more or less imbricate, pubescent, about 1.5
cm long, 7 mm wide, acuminate, sheathing, elliptic to oblong-
elliptic; bracteoles sheathing, obovate when spread out, 3 to 4

SIG; 5 Merrill: Philippine Plants, XII 993

mm long. Flowers numerous. Sepals oblong to elliptic-oblong,
obtuse, 3-nerved, not glandular, externally pubescent, rather thin,
4 to 5 mm long, 1.5 to2 mm wide. Petals thinly membranaceous,
about as long as the sepals, finely nerved. Stamens 6, their fila-
ments all bearded, three sterile and three perfect, bearing oblong,
1.5 mm long anthers. Capsule 3-angled, oblong-ellipsoid, about
6 mm long, 3 mm in diameter, 3-celled, shortly pointed. Seeds
1-seriate, 4 in each cell, angular, brown, 1 to 1.3 mm. long.

SEMERARA, Merrill 4134, July, 1905, common in open, wet grasslands
a few meters above sea level.

A species very closely allied to the Indian Aneilema scapiflorum Wight,
differing in its relatively longer, narrower leaves which are not at all

scabrous, its distinctly pubescent inflorescence, and its more or less imbricate
bracts.

ANEILEMA PLATYPHYLLUM sp. nov. Tricarpellaria, Huaneilema.

Species a praecedente differt planta minoribus, foliis brevio-
ribus latioribusque, usque ad 10 cm longis et 2.5 cm latis,
inflorescentiis glabris, bracteis minoribus, non imbricatis, sepalis
glanduloso-puncticulatis.

Like Aneilema azureum closely allied to the Indian Aneilema
scapiflorum Wight, from which it differs in many characters.
Tufted, the leaves comparatively few, from fibrous and tuberous
roots, the tubers more or less fusiform, when dry up to 3 cm long
and 6 mm in diameter. Leaves smooth, membranaceous or char-
taceous, 4 to 8 in each rosette, oblong-ovate to ovate-lanceolate,
4 to 10 cm long, 1.5 to 2.5 em wide, base not or but slightly
narrowed, sheathing, apex acute or somewhat acuminate. Scapes
2 to 5 from each rosette, leafless, or with one much-reduced leaf,
12 to 20 em high, the flowers cymosely arranged, pink or nearly
white, the cymes narrow, comparatively few-flowered. Primary
bracts oblong-ovate, sheathing, less than 1 cm long, acute or
acuminate, the bracteoles triangular-ovate, 2 to 3 mm long.
Sepals oblong-elliptic, 5 mm long, 2.2 mm wide, glandular-punc-
tate with yellowish dots and short dashes, 3-nerved, obtuse.
Petals thinly membranaceous, finely nerved, elliptic-obovate,
rounded, about 5 mm long, 3.5 mm wide. Stamens 6, three
sterile, three fertile, all bearded; anthers broadly elliptic, 1.5 mm
long. Capsule elliptic, 3-angled, 3-celled, acute, 6 mm long, 3.5
mm in diameter. Seeds 1-seriate, 3 in each cell, superposed,
brownish, angular, irregular, about 1.5 mm long.

LUZON, Province of Rizal, Antipolo, Bur. Sci. 20985 Ramos (type), June,

1913; Tanay, Bur. Sci. 3342 Ramos, June, 1907; Bosoboso, Bur. Sci. 985
Ramos, June, 1906. |

294 The Philippine Journal of Science 1915

Not as closely allied to Aneilema scapiflorum as is the preceding species,
differing from both in its few, relatively much shorter and broader leaves,
in its glandular-punctate sepals, and in its small bracts. Loher 1340,
1939, in the Kew Herbarium under Aneilema scapiflorum Wight, are both
probably referable to A. platyphyllum.

ANEILEMA GIGANTEUM (Vahl) R. Br. Prodr. (1810) 271; Naves in
Novis. App. (1880) 271; C. B. Clarke in DC. Monog. Phan. 2
(1881) 212.

Commelina gigantea Vahl Enum 2 (1806) 177.

SEMERARA, Merrill 4182, July, 1905. MINDANAO, Bukidnon Subprovince,
Bur. Sci. 21885 Escritor, July, 1913.

Naves’s record for this species as a Philippine one has not previously
been verified. Tropical Africa and Asia through Malaya to tropical
Australia.

JUNCACEAE
JUNCUS Linnaeus

JUNCUS BUFONIUS L. Sp. Pl. (1753) 328; Buchenau in Engl. Pflanzen-
reich 25 (1906) 105.

LUZON, Benguet Subprovince, Mount Tonglon, Merrill 7794, May, 1911.

Not previously reported from the Philippines; warmer parts of the
entire world.

This species was found scattered along the recently constructed bridle
trail to Mount Tonglon, in the mossy forest, at an altitude of about 1,900
meters. It is undoubtedly an introduced plant here.

LILIACEAE

SMILAX Linnaeus

SMILAX LOHERI sp. nov. § Eusmilaz.

Frutex scandens glaber, ramulis teretibus, aculeis sparsis te-
nuibus nectis vel curvatis 2 ad 5 mm longis armatis; foliis crasse
coriaceis, late ovatis, 5-nervis, basi, late cordatis, apice abrupte
acuminatis apiculatisque, 8 ad 12 cm longis, subtus glaucescen-
tibus; racemis 1-umbellatis, pedunculis 4 ad 5 cm longis; floribus
numerosis, sepalis 5 mm longis.

A scandent, glabrous vine, the branches slender, terete, armed
with scattered, slender, straight or somewhat curved spines 2
to 5 mm in length. Leaves very broadly ovate, thickly coria-
ceous, 8 to 12 cm long, 7 to 10 cm wide, when dry slightly
shining, the upper surface brownish, the lower glaucous or sub-
glaucous, the base very broad, shallowly cordate, distinctly 5-
nerved, the inner pair of nerves reaching the apex of the leaf,
much more prominent than the outer pair, the reticulations
rather distinct, the apex abruptly acuminate, the acumen about
8 mm long, apiculate; petiole about 1 cm long, inflated, the

Bic, 6 Merrill: Philippine Plants, XII 295

tendrils slender, up to 15 cm in length. Inflorescence axillary,
solitary, consisting of a single umbel, the floral branch less than
1.5 em long, subtended by a coriaceous, ovate, acuminate bract
about 8 mm in length which is split down the back, the floral
branch bearing at its apex a very much reduced or sometimes
aborted bud, and opposite to it a single oblong, obtuse, or retuse
and apiculate bract about 5 mm in length, this bract subtending
the solitary, slender, 4 cm long peduncle. Male flowers 40 to 50
in each umbel, their pedicels about 8 mm long, the subtending
bracteoles ovate, 2 mm long or less. Sepals about 5 mm long,
1.5 mm wide, recurved, obtuse, cucullate. Petals as long as the
sepals, 0.56 mm wide. Stamens 6, all fertile, their filaments
about 3.5 mm long, the anthers 1.5 mm long. Female flowers
and fruits not known.

LUZON, Province of Rizal, Angilog, Loher 6986, April 15, 1906.

A characteristic species, distinguishable by its short-petioled, thickly
coriaceous, broadly ovate leaves which are abruptly acuminate and apic-
ulate, the base very broad, cordate, and 5-nerved, the lower surface
glaucous. Among the Philippine species it is probably most closely allied
to Smilax bracteata Pres], but that species has an entirely different inflores-
cence and different leaves.

SMILAX LUZONENSIS Presl Rel. Haenk. 1 (1827) 131; Mig. Fl. Ind.
Bat. 3 (1859) 566.

Luzon, Haenke (type in herb. Prag.). MINDANAO, Lake Lanao, Camp
Keithley, Mrs. Clemens 835, 905 bis, s. n.

This species was described by Pres] from a sterile specimen, no flowers
or fruits being known. I have seen the type in the Prague Herbarium, and
it seems to be matched by the specimens cited above, two of which are with
male flowers, the other with very immature fruits. The specimens agree
with Presl’s description, so far as it goes, and with a carbon rubbing of
typical leaves of the type, preserved in the herbarium of the Bureau of
Science.

C. DeCandolle* reduces Smilax luzonensis Presl (1827), with doubt, to
Smilax helferi C. DC. While the two species may be identical, Presl’s
specific name has a priority by many years, and should be adopted, at least
for the Philippine form. Only by a careful comparison between Philippine
specimens of Smilax luzonensis Presl, and authentic specimens of S. helferi
C. DC., will it be possible to determine whether one or two species are
represented.

Smilax luzonensis Pres] has solitary male umbels which are 2 to 2.5 cm
in diameter, each with from 20 to 30 flowers, the peduncles slender, axillary,
2.5 to 3 em long, bracteate at the base, and with a small ovate bracteole
at the lower one-fourth or below. The pedicels are about 8 mm long;
the sepals are narrowly oblong, obtuse, 6 mm long, the petals as long as the
sepals but much narrower. Anthers about one-fifth as long as the filaments.
Female flowers and mature fruits unknown.

*Monog. Phan. 1 (1878) 176.

996 The Philippine Journal of Science 1915
ZINGIBERACEAE

ALPINIA Linnaeus

ALPINIA BRACHYANTHA sp. nov. § Hellenia.

Species A. pubiflorae K. Schum. ut videtur affinis, differt
floribus multo minoribus vix 1 cm longis, connectivo inappen-
diculato.

Simple, erect, nearly glabrous, except the pubescent inflores-
cence, apparently less than 1 m high. Sheaths rather loose,
ciliate on their margins; ligule 8 to 10 mm long, oblong-ovoid,
obtuse, margins ciliate. Leaves lanceolate or narrowly oblong-
lanceolate, 22 to 30 cm long, 2.5 to 3.5 cm wide, subequally
narrowed at both ends, subcoriaceous, glabrous, base cuneate,
apex sharply acuminate. Panicle erect, about 20 cm long, the
branches two or three, spreading-ascending, about 7 cm long,
many-flowered, the rachis and pedicels softly pubescent with
short, spreading hairs, the pedicels 2 to 3 mm long; bracteoles
apparently small, very caducous (not seen). Ovary globose to
ovoid-globose, 1 to 1.5 mm in diameter, purple when fresh, very
densely ciliate-pilose. Calyx-tube about 7 mm long, subequally
3-toothed, glabrous, the teeth broadly ovate, obtuse, 1.5 mm long
or less, ciliate at their apices. Corolla-tube 6 to 7 mm long,
about 1 mm in diameter, light green, glabrous, the lobes oblong,
somewhat cucullate, 4 to 5 mm long, 2.5 to 3 mm wide. Lip
3 mm long, glandular-punctate, twice cleft, the lateral lobes
rounded, obovoid, twice as wide as the oblong, obtuse inner
lobes, white marked with purple. Anther 3 mm long, the con-
nective inappendiculate.

Luzon, Province of Cagayan, Abulug River, Weber 1571 (type), Feb-
ruary, 1912, altitude about 60 meters; same locality, Bur. Sci. 13920 Ramos,
For. Bur. 19617 Curran.

A species allied to Alpinia patiiare K. Schum., but at once distinguished
by its much smaller flowers and its inappendiculate anther-connective.

KAEMPFERIA Linnaeus

KAEMPFERIA PHILIPPINENSIS sp. nov. § Monolophus.

Planta erecta, saltem 50 cm alta, leviter hirsuta; foliis 9, lan-
ceolatis, subcaudato-acuminatis, basi angustatis, acutis, vix vel
breviter petiolatis, circiter 20 cm longis; spicis solitariis, pauci-
floris, in vaginis superioribus, corollae lobis inaequalibus, supe-
rioribus valde tubuloso-cucullatis.

A rather slender, erect plant at least 50 cm high, the stems
3 to 4 mm in diameter, the sheaths, leaves, and inflorescence
with scattered, spreading, pale brown, ciliate-hirsute hairs.

X, C, 5 Merrill: Philippine Plants, XII 2907

Leaves 9 (or more), lanceolate, membranaceous, about 20 cm
long, 3 to 3.5 em wide, subequally narrowed to the slender,
caudate-acuminate apex and to the acute base; sheaths rather
loose or slightly inflated above, rather prominently ciliate-hirsute ;
petioles none or up to 5 mm in length. Spikes in the uppermost
sheath, more or less included, short, few-flowered, only one or
two flowers opening at a time, the maximum number apparently
4 or 5 in each spike, the bracts lanceolate, acuminate, about
2 mm long, 5 mm wide, hirsute. Calyx cylindric, about 12 mm
long, punctate-glandular, as are the other parts of the flower,
hirsute, very broadly and _ shortly 3-toothed. Corolla-tube
slender, slightly enlarged upward, 16 mm long, sparingly hirsute,
the lobes unequal, the upper one about 11 mm long, 4 mm wide
in the flattened portion, slenderly 9-nerved, the apical 3 mm a
cylindric, obtuse hood, the two lateral ones oblong, 7 mm long,
3 mm wide, 5-nerved, slightly cucullate at the apex; staminodes
slender, lateral, attached to the margins of the lip near its base,
about 2 mm long, 0.6 mm wide; lip not seen. Filament flat,
glandular-punctate, 5 to 6 mm long, about 1 mm wide; anther-
cells parallel, 3 mm long, the connective appendiculate, the ap-
pendage ovate, obtuse, more or less inflexed or curled when
dry. Ovary villous, 3-celled, the cells with few axile ovules;
style slender; stigma obconic, ciliate. Fruit not seen.

LUZON, Province of Laguna, Dahican River back of San Antonio, Bur.
Sci. 14952 Ramos, June, 1912, in forests, flowers white.

The second species of the genus to be found in the Philippines, the only
other one, K. galanga L., manifestly an introduced plant in the Archipelago.
Kaempferia philippinensis Merr. manifestly belongs in the section Mono-
lophus, and is very distinct from the other known species, although probably
as closely allied to the Bornean K. gracillima K. Schum. as to any other

species. The lip on the flowers available for study has been destroyed, and
hence cannot be described until the plant is again collected.

FAGACEAE
QUERCUS Linnaeus

QUERCUS ROBINSONII sp. nov. § Cyclobalanus.

Arbor circiter 20 m alta, subtus foliis ramulisque dense ferru-
gineo-furfuraceis; foliis oblongo-ovatis ad oblongo-lanceolatis,
coriaceis, integris, usque ad 12 cm longis, basi acutis, apice
tenuiter subcaudato-acuminatis, nervis utrinque circiter 8, pro-
minentibus; glandibus late ovoideis, glabris, circiter 1.3 cm longis
latisque, apiculatis, cupulis circiter 8 mm altis, 1.6 cm diametro,
ferrugineo-pubescentibus, truncatis, laminis numerosis, indis-
tinctis, denticulis numerosis.

298 The Philippine Journal of Science 1915

A tree about 20 m high, the younger parts and the leaves on
the lower surface densely ferruginous-furfuraceous. Branches
terete, dark-colored when dry, glabrous, sparingly lenticellate,
the younger ones densely ferruginous-furfuraceous. Leaves al-
ternate, oblong-ovate to oblong-lanceolate, coriaceous, 7 to 12
cm long, 2.5 to 5 cm wide, subequally narrowed to the acute
base and to the slender subcaudate-acuminate apex, the acumen
usually about 1 cm long, blunt, the upper surface furfuraceous
in very young leaves, soon becoming quite glabrous, brownish-
olivaceous when dry, shining, the lower surface paler than the
upper, in extreme age glabrous or nearly so; lateral nerves
about 8 on each side of the midrib, prominent on the lower
surface; petioles 5 to 8 mm long. Male spikes up to 14 cm long,
the flowers scattered below, rather crowded above. Glands
broadly ovoid or conic-ovoid, glabrous, about 1.3 cm long and
wide, apiculate. Cup thick, ferruginous-pubescent, broadly
saucer-shaped, truncate, base thickened, about 8 mm high and
1.6 cm in diameter, the laminz numerous, indistinct, below scat-
tered, above approximate, the teeth numerous, distinct, less than
1 mm long.

LUZON, Province of Laguna, Mount Maquiling, Bur. Sci. 17911 Brown
(type), May, 1913, Bur. Sci. 17148 Robinson, December, 1912, in forests,
altitude about 700 meters. Possibly referable here are Loher 6968, from
Rizal Province (specimens with male flowers), and Bur. Sci. 9462 Robinson,
from Infanta, Province of Tayabas, Luzon (glands pubescent).

The alliance of this species seems to be with Quercus caudatifolia Merr.,
from which it is at once distinguished by its differently shaped gland which
is as wide as long. Among the extra-Philippine species it approaches Quer-
cus sundaica Blume, but has much smaller fruits than our Philippine mate-
rial so referred.

URTICACEAE

LAPORTEA Gaudichaud
LAPORTEA BRUNNEA sp. nov.

Frutex, inflorescentiis exceptis glaber; foliis in siccitate utrin-
que brunneis, nitidis, oblongis, usque ad 22 cm longis, subcoriaceis
vel chartaceis, acute acuminatis, basi acutis, leviter 3- vel obscure
5-plinerviis, longe petiolatis, nervis lateralibus utrinque circiter
15; inflorescentiis pilis paucis urentibus instructis, axillaribus,
usque ad 5 cm longis, floribus ¢ flabellatim dispositis.

A shrub 1 to 2 m high, glabrous except the slightly hairy in-
florescence. Branches light gray, terete, wrinkled when dry.
Leaves uniformly brown, of the same color on both surfaces and
slightly shining when dry, the upper surface minutely verrucu-
lose, the lower smooth, chartaceous to subcoriaceous, brittle, ob-
long, 12 to 22 cm long, 4 to 7.5 em wide, subequally narrowed to

X, 5 Merrill: Philippine Plants, XII 299

the usually acute base and to the sharply acuminate apex, entire,
the base somewhat 3- or obscurely 5-plinerved; lateral nerves
about 15 on each side of the midrib, rather prominent, somewhat
curved; petioles 3 to 6.5 cm long. Pistillate inflorescences axil-
lary, with few short, stinging hairs, up’ to 5 cm in length, the
branches few. Flowers flabellately disposed on the ultimate
branchlets, the ultimate branchlets usually dichotomous, short,
the achenes deflexed, ovoid, compressed, about 3 mm long and
2 mm wide, the styles reflexed, very slender, about 2 mm long.

SAMAR, Cauayan Valley, Bur. Sci. 17559 Ramos, March, 1914, on forested
slopes, the flowers blue and white.

A species well characterized by its long-petioled leaves which are uni-
formly brown when dry, and which are quite glabrous, with no vestiges of
stinging hairs; the only part of the dried plant that presents stinging hairs
is the inflorescence. The alliance of the species is manifestly with Laportea
luzonensis Warb., on account of its flabellately arranged pistillate flowers,
although in all details it is entirely different from Warburg’s species.

LAPORTEA MONTICOLA sp. nov.

Frutex vel arbor parva, glabra vel subglabra; foliis oblongis
ad oblongo-lanceolatis, chartaceis vel subcoriaceis, integris, te-
nuiter caudato-acuminatis, usque ad 10 cm longis; infructes-
centiis axillaribus, 6 ad 8 cm longis, fructibus circiter 4 mm
diametro haud flabellatis.

A shrub or small tree, glabrous, or with very few persistent
hairs when dry, the branches slender, grayish, wrinkled, the
internodes short, especially on the ultimate branchlets where
they are usually less than 5 mm long. Leaves oblong to oblong-
lanceolate, dark olivaceous when dry, slightly shining, thickly
chartaceous to subcoriaceous, entire, 7 to 10 cm long, 2 to 3.5 cm
wide, base obtuse to rounded, 3-nerved, apex very slenderly cau-
date-acuminate, the acumen up to 2 cm in length, sharp, the upper
surface rather densely white-puncticulate; lateral nerves about
6 on each side of the midrib, rather distinct, curved, anasto-
mosing, the reticulations lax; petioles 1.5 to 2.4 cm long. In-
fructescences 6 to 8 cm long, axillary, the branches few, spread-
ing, 2.5 cm long or less. Fruits white, fleshy, about 4 mm in
diameter, the receptacle enlarged in fruit, the achenes inequi-
lateral, compressed, about 2.5 mm long and 2 mm wide, but one
or two on each ultimate branchlet.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19765 McGregor,
February 1, 1913, altitude not indicated, but apparently from the mossy
forest.

A species somewhat resembling a small-leaved form of Laportea luzonensis
Warb., but its fruits not flabellate. It is readily distinguished by its small
leaves which are slenderly and prominently caudate-acuminate.

300 The Philippine Journal of Science 1915

LAPORTEA TRIPLINERVIA sp. nov.

Arbor parva, subtus foliis pilis longis sparsis urentibus in-
structis; foliis oblongis, chartaceis vel membranaceis, usque ad
18 em longis, in siccitate viridis, utrinque subaequaliter angus-
tatis, basi acutis, triplinerviis, apice acuminatis, nervis utrinque
circiter 6; infructescentiis axillaribus, usque ad 6 cm longis, fruc-
tibus in siccitate nigris, capitulis globosis, 8 ad 10 mm diametro,
acheniis numerosis.

A tree up to 13 m in height, the leaves with persistent, long,
stiff, scattered, stinging hairs on the lower surface. Branches
pale gray, terete, 4 to 6 mm in diameter. Leaves green when
dry, in general oblong, chartaceous or membranaceous, 10 to
18 cm long, 3 to 6.5 cm wide, entire, subequally narrowed
to the acute, 3-plinerved base and to the acuminate apex, the
upper surface rather densely white-puncticulate; lateral nerves
about 6 on each side of the midrib, rather prominent, curved;
petioles 2 to 4 cm long. Infructescences axillary, up to 6 cm
long, with few branches, the achenes arranged in dense globose
heads at the tips of the branchlets, up to 15 or more achenes in
each head, the heads black when dry, 8 to 10 mm in diameter.

Luzon, Province of Tayabas, Mount Pular, Bur. Sci. 19462 Ramos,
January 15, 1913, on forested slopes.

A species somewhat resembling Laportea luzonensis Warb., but with the
leaves constantly acute and 3-plinerved at the base, the stinging hairs
persistent on the lower surface, which sting even when dry, and with

an entirely different arrangement of its fruits, the achenes crowded into
dense, globose heads which are black when dry, not flabellately arranged.

PROTEACEAE
HELICIA Loureiro

HELICIA LONGIFLORA sp. nov.

Arbor parva, circiter 8 m alta, glabra; foliis late oblongo-
oblanceolatis, integris, usque ad 15 cm longis, breviter obtuse
acuminatis, basi acutis; racemis folia subaequantibus, paucifloris,
pedicellis in paribus connatis; floribus 2.5 cm longis.

A small tree, quite glabrous, or the racemes with few, scattered,
deciduous hairs, the branches rather stout, terete, brownish.
Leaves subcoriaceous, broadly oblong-oblanceolate, 11 to 15 cm
long, 3.5 to 5.5 em wide, entire, the apex rather shortly and
obtusely acuminate, sometimes falcate, the base gradually nar-
rowed to the short stout petiole, cuneate and sometimes a little
decurrent, upper surface olivaceous when dry, somewhat shin-
ing, the lower brownish, dull; lateral nerves prominent, about
10 on each side of the midrib, curved-ascending; petioles 5 to 8

X, C, 5 Merrill: Philippine Plants, XII 301

mm long. Racemes about as long as the leaves, solitary, axil-
lary, comparatively few-flowered. Pedicels in pairs, 4 mm long,
united nearly to the middle; bracteoles deciduous (not seen).
Perianth 2.5 cm long, rather slender. Anthers, including the
prominent connective, 2 mm long. Hypogynous scales free or
nearly so, coriaceous, ovate, 1 to 1.3 mm long.

LUZON, Province of Tayabas, Lucban (Mount Banajao), Whitford 982,

October, 1904, margins of forests, altitude about 750 meters.
Characterized by its entire leaves and comparatively long flowers.

HELICIA INTEGRA sp. nov.

Arbor parva, glabra; foliis oblongo-lanceolatis vel oblongo-
oblanceolatis, usque ad 14 cm longis, integerrimis, utrinque an-
gustatis, apice acuminatis, basi cuneatis, brevissime petiolatis,
chartaceis vel subcoriaceis; racemis folia subaequantibus, mul-
tifloris, axillaribus; floribus tenuibus, circiter 1.5 cm longis.

A small glabrous tree, or the racemes with very few, scattered,
deciduous hairs. Branches terete, grayish-brown, smooth.
Leaves oblong-lanceolate to oblong-oblanceolate, entire, 10 to 14
cm long, 2 to 5 cm wide, chartaceous or subcoriaceous, somewhat
yellowish when dry, slightly shining, glabrous, narrowed at
both ends, the apex shortly and usually sharply acuminate or
acute, the base cuneate, narrowed into the very short petiole;
lateral nerves 10 to 12 pairs, slender, distinct, the reticulations
very slender; petioles stout, less than 3 mm long. Racemes
solitary, axillary, about as long as the leaves, many-flowered.
Flowers yellowish, in pairs, their pedicels in pairs, free to the
rachis, about 3 mm long; bracteoles ovate, subpersistent, 1 mm
long. Perianth very slender, about 0.5 mm in diameter, a little
thicker at the base and apex, 1.6 cm long, the segments linear,
spirally twisted in anthesis. Anthers, including the prominent
connective, 1.2 mm long. Hypogynous scales free, oblong-ovate,
obtuse, coriaceous, 1 mm long.

Luzon, Province of Laguna, San Antonio, Bur. Sci. 15127 Ramos, June,
1912, in forests along streams.

The species is characterized by its entire, nearly sessile leaves, and ap-
pears to be most closely allied to Helicia moluccana Blume.

POLYGONACEAE
POLYGONUM Linnaeus

POLYGONUM BENGUETENSE sp. nov. § Persicaria, Trigyna.
Eglandulosum, caulibus tenuibus, quadrangularibus, simplici-
bus vel supra leviter ramosis; ochreis cylindraceis, circiter 1 cm
longis, truncatis, apice rigide ciliatis, ciliis 2 ad 3 mm longis;
foliis oblongo- ad elliptico-ovatis, 1 ad 2.5 cm longis, acuminatis,

302 The Philippine Journal of Science 1915

basi rotundatis vel subcordatis; racemis pedunculatis, oblongis
vel cylindraceis, densis, usque ad 1 cm longis, pedunculis leviter
capitato-glandulosis.

A slender, ascending, simple or very sparingly branched plant,
nearly glabrous, the stems often decumbent below, 4-angled, 1
mm in diameter or less. Nodes elongated. Leaves scattered,
oblong- to elliptic-ovate, chartaceous, entire, apex somewhat
acuminate, base rounded or subcordate, 1 to 2.5 em long, 7 to
12 mm wide; petioles 5 mm long or less, inserted at or near
the base of the sheath, often scabrid; sheaths cylindric, split
down one side, brown, membranaceous, about 1 cm long, apex
truncate, the apical margin with erect, rigid, 2 to 3 mm long
cilia, the base of the sheath often with few reflexed, spinelike
cilia. Racemes few, short, dense, 1 cm long or less, oblong or
cylindric, very loosely corymbose, the peduncles slender, spar-
ingly capitate-glandular. Bracts oblong-ovate, margins and apex
prominently ciliate, about 3 mm long. Calyx pink, 3 to 3.5 mm
long, the pedicels short, the sepals oblong-obovate, obtuse.
Stamens 6, with 6 alternating, small, perigynous scales. Style
3-cleft, short. Achene oblong-ovate, sharply 3-angled, about 3
mm long.

LuzON, Benguet Subprovince, Baguio, scattered among grasses and sedges
on wet seepage slopes about springs, altitude about 1,550 meters, Phil. Pl. 768
Merrill, May, 1911, Merrill 9682, May, 1914, Bur. Sci. 18483 Ramos, Bur.
Sct. 14120 Robinson.

A species apparently belonging in the section Persicaria, but very different
from the other species placed here.

POLYGONUM CONVOLVULUS Linn. Sp. Pl. (1753) 364.

LUZON, Benguet Subprovince, Baguio, Merrill 9698, May, 1914, a weed in
gardens, recently introduced, altitude about 1,400 meters.

Widely distributed in the north temperate zone, a native of Europe or
Asia. It has not before been reported from the Philippines.

CARYOPHYLLACEAE
POLYCARPON Linnaeus

POLYCARPON INDICUM (Retz.) comb. nov.
Loeflingia indica Retz. Obs. 4 (1786) 38.
Pharnaceum depressum Linn. Mant. 2 (1771) 564, non Polycarpon
depressum Nutt.
Polycarpaea depressa DC. Prodr. 3 (1828) 357.
Polycarpon loeflingiae Benth. & Hook. f. ex Edgew. in Hook. f. Brit.
Ind. 1 (1874) 245.
Polycarpon polyphyllum Blanco FI. Filip. (1837) 53, ed. 2 (1845) 36,
ed. 3, 1 (1877) 66.
This species has been credited to the Philippines by F.-Villar, who
correctly, I think, reduced to Polycarpon loeflingiae Blanco’s P. polyphyllum.

%, 0, 5 Merrill: Philippine Plants, XII 303

The species is apparently very rare in the Philippines, but although Blanco’s
specimens were from Pasig, near Manila, I have seen but a single Philippine
specimen that I consider referable to the genus and species: LUZON, Province
of Pampanga, Arayat, Merrill 1462, March, 1903, erroneously placed under
Mollugo.

RANUNCULACEAE

CLEMATIS Linnaeus

CLEMATIS LESCHENAULTIANA DC. var. SUBGLABRIFOLIA var. nov.
A typo differt foliolis glabris vel subglabris, subtus parcissime
breviter hirsutis.

Luzon, Subprovince of Ifugao, Mount Polis, Bur. Sci. 19813 McGregor,
February, 1913: Subprovince of Lepanto, Mount Malaya, For. Bur. 14496
Darling, January, 1909.

The typical form of the species, which is well represented in the Bureau
of Science herbarium by a large series of specimens from Mountain
Province, Luzon, has rather densely ciliate-hirsute leaves. In the variety
above proposed the leaves are nearly glabrous; otherwise the form is
very similar to typical Clematis leschenaultiana DC.

MENISPERMACEAE
HYPSERPA Miers

ZANTHOXYLUM ? TRIPLINERVE Turcz. in Bull. Soc. Nat. Mose. 36°
(1863) 597; F.-Vill. Novis. App. (1880) 35; Vid. Rev. Pl. Vasc.
Filip. (1886) 74; Perk. Frag. Fl. Philip. (1905) 161 = HYPSERPA
CUSPIDATA (Wall.) Miers in Ann. Nat. Hist. II 7 (1851) 40;
Diels in Engl. Pflanzenreich 46 (1910) 206.

The status of Turczaninow’s species has been doubtful, as from his
description it is at once evident that the plant he described, having simple
leaves, could not be a Zanthoxylum, as indicated by Perkins |. c. Doctor
Perkins indicates that such a Province as “North Hows,” cited by Turc-
zaninow, does not exist in Luzon, and suggested that the plant may have
come from Lord Howe’s Island. Through the kindness of the director of the
Botanical Institute of the University of Kharkoff, I have been able to
examine Turczaninow’s type which is preserved in the herbarium of that
institution. The original label reads ‘“Runner No. 2, Ilocos Norte, Island
of Luzon, April, 1839, H. C.,’’ with notes on the dissection of the flowers by
Turczaninow. The plant is identical in all respects with Cuming 1252,
localized by Cuming himself as from the Province of North Ilocos (= Ilocos
Norte), which is Hypserpa cuspidata Miers; the specimen in Turczaninow’s
herbarium, type of Zanthoxylum ? triplinerve Turez., is unquestionably
a part of the same collection.

CAPPARIDACEAE
CAPPARIS Linnaeus
CAPPARIS AFFINIS sp. nov.
Species C. sepiariae affinis, differt foliis majoribus, usque ad
9 cm longis, coriaceis, nervis utrinque magis numerosis, utrinque

circiter 8.
133910——2

304 The Philippine Journal of Science 1915

A scandent, armed shrub, apparently entirely glabrous.
Branches pale greenish-yellow when dry, terete, stout, slightly
zigzag, glabrous, or the younger ones very slightly pubescent.
Leaves alternate, oblong, coriaceous, 8 to 9 em long, 3 to 4 cm
wide, pale yellowish-green when dry, brittle, the upper surface
very smooth and strongly shining, the lower paler, dull, base
rounded, apex retuse; lateral nerves about 8 on each side of
the midrib, slender, indistinct, scarcely anastomosing, the retic-
ulations obsolete or nearly so; petioles 5 to 7 mm long; spines
stout, recurved, about 2 mm long. Inflorescence fascicled or
umbellate, axillary, apparently few-flowered. Flowers not
seen. Peduncles in fruit 3 mm long or less, each bearing from
2 to 4, long-pedicelled fruits, the pedicels about 1 cm long, the
androgynophore as long as the peduncles. Fruit globose, about
8 mm in diameter when dry, the pericarp somewhat wrinkled,
glabrous, pale, shining.

PALAWAN (Arricife Island, near the southern end of Palawan), Bur. Sci.
21583 Escritor, August 14, 1913.

A species with much the appearance of Capparis sepiaria Linn., but

with much larger, quite glabrous, coriaceous, more numerously nerved
leaves.

CAPPARIS PALAWANENSIS sp. nov.

Frutex suberectus, 2 ad 3 m altus, partibus junioribus plus
minusve pubescens, ramis ramulisque inermis vel subinermis;
foliis oblongo-ovatis, usque ad 11 cm longis, acuminatis, basi
rotundatis, nervis utrinque 10 ad 12; floribus subterminalibus,
solitariis vel binis; fructibus longissime pedunculatis, ellipsoideis
ad globoso-obovoideis, circiter 1.5 cm diametro, extus pube-
scentibus.

A suberect shrub, 2 to 3 m high, the branches somewhat droop-
ing or subscandent, unarmed, or with only few, scattered, short
spines. Branches terete, glabrous, dark-colored, the branchlets
yellowish-olivaceous, terete, slender, rather densely pubescent
with short hairs. Leaves alternate, oblong-ovate, 9 to 11 em
long, 3.5 to 4.5 cm wide, chartaceous, base broadly rounded,
narrowed in the upper one-third to the rather sharply acuminate
apex, pale olivaceous, rather dull, and glabrous or very slightly
pubescent on the upper surface, the lower surface much paler,
slightly pubescent on the midrib and nerves; lateral nerves 10
to 12 on each side of the midrib, spreading, anastomosing, the
reticulations lax; petioles densely pubescent, 2 to 4 mm long.
Flowers subterminal, in the uppermost axils, solitary or in pairs,

x, ¢, 5 Merrill: Philippine Plants, XII 305

not superposed (not seen). Peduncles in fruit about 4 cm long,
slender, the androgynophore about as long as the peduncle,
somewhat thicker and slightly thickened upward, the fruit el-
lipsoid to globose-obovoid, about 1.5 cm in diameter, nearly black
when ripe, the pericarp distinctly pubescent.

PALAWAN, between Taytay and Lake Manguao, Merrill 9459, April 23,
1913, on forested dry slopes, altitude about 60 meters.

Probably as closely allied to Capparis lobbiana Turcz. as to any other

species, but with much larger, differently shaped leaves, larger fruits, and
unarmed or nearly unarmed branches.

CAPPARIS VENOSA sp. nov.

Frutex scandens, floribus exceptis glaber, spinis brevissimis,
vix 1 mm longis; foliis oblongis vel elliptico-oblongis, chartaceis,
usque ad 18 cm longis, acuminatis, basi acutis vel rotundatis,
nervis utrinque circiter 7, prominentibus, anastomosantibus;
floribus paucis, in racemis terminalibus dispositis, pedicellis in
paribus seriatim dispositis supra nodis aculeatis sed defoliatis;
petalis dense pubescentibus.

A scandent shrub, glabrous except the flowers, the branches
terete, straw-colored when dry, the ultimate ones 2 to 3 mm
in diameter, armed with very short spines at the nodes, the
spines less than 1 mm in length. Leaves oblong to elliptic-oblong,
15 to 18 cm long, 5.5 to 7.5 cm wide, chartaceous or subcoriaceous,
pale when dry, shining, the lower surface decidedly paler than
the upper, the apex acuminate, the base acute or rounded; lateral
nerves prominent, distinctly impressed on the upper surface,
projecting on the lower, about 7 on each side of the midrib, curved
and distinctly anastomosing, the reticulations rather lax; petioles
5 to 8 mm long. Racemes terminal, few-flowered, the flowers
in pairs, seriate above the leafless nodes, but each node with a
pair of minute spines, scattered, the pedicels about 2 cm long.
Sepals 8 mm long, oblong-elliptic or elliptic, concave, acute, glab-
rous or nearly so. Petals obovate to broadly oblong-obovate,
rather densely pubescent, rounded, about 12 mm long, 8 mm wide.
Stamens numerous, their filaments up to 2.5 cm long; anthers
oblong, 1.3 mm long. Gynophore glabrous, 1.5 cm long, bearing
the glabrous, narrowly ovoid, 3 to 4 mm long ovary.

MINDANAO, District of Cotabato, near Lebak, For. Bur. 14216 Whitford,
April, 1912, in dipterocarp forests at low altitudes.

A species probably as closely allied to Capparis micracantha DC. as to

any other, but very different in many characters. The venation of the
leaves and the inflorescence are characteristic.

306 The Philippine Journal of Science 1915

NEPENTHACEAE
NEPENTHES Linnaeus
NEPENTHES BRACHYCARPA sp. nov.

Alte scandens, partibus junioribus inflorescentiisque exceptis
glabra; inflorescentiis racemosis, pedicellis brevibus, 1-floris;
operculum ascidii intus planum; foliis petiolatis, nervis longitu-
dinalis circiter 2 utrinque, tenuibus, obscurissimis, basi angus-
tatis, longe in petiolo decurrento-alatis, leviter amplexicaulibus,
haud decurrentibus; capsulis brevibus, 6 ad 15 mm longis.

Scandent, 4 to 5 m high or more, glabrous except the dark
brown pubescent very young parts and the racemes, the stems
terete, brown, about 5 mm in diameter. Leaves alternate, spir-
ally arranged, the lamina coriaceous, 12 to 14 cm long, 2 to 3 cm
wide, brownish or olivaceous when dry, slightly shining, nar-
rowed below and winging the 4 to 6 cm long petiole, the wings
about 2 mm wide, about one-third amplexicaul at the base, apex
acute or acuminate and extended as a long, slender tendril, the
lateral longitudinal nerves at most 2 on each side of the midrib,
very slender, obscure, or obsolete. Pitchers reddish, glabrous,
or when young somewhat pubescent, about 9 cm long, 3 to 3.5
cm in diameter, cylindric-ventricose, dimorphous, those on young
plants with two ventral wings which are about 5 mm wide and
prominently lacerate-toothed, those on mature plants with the
wings reduced to merely low ridges which are not all toothed,
0.5 mm wide and high, the base gradually narrowed into the
tendril, the mouth oblique, 3 to 4 cm in diameter, ovate, dorsally
produced into a short stalk, the peristome cylindric, equal, 2 to 3
mm in diameter, densely striate; operculum elliptic to elliptic-
ovate, broadly rounded, base rounded or somewhat cordate, about
3.5 em long, 2.5 cm wide, plane, not at all keeled or cristate,
inside with numerous small, round glands. Racemes dark brown
pubescent, the female ones in flower about 9 cm long and 1.5 cm
wide, in fruit very long-peduncled, about 13 cm long and 3 cm
wide, the peduncles up to 35 cm long. Pedicels 3 to 5 mm long,
pubescent. Sepals elliptic, obtuse, 3 mm long, somewhat pubes-
cent. Ovary narrowly ellipsoid, 3 mm long, densely ferruginous-
pubescent. Capsules glabrous, oblong to lanceolate, 6 to 1.5
mm long.

PALAWAN, Silanga Peak, back of Silanga, Merrill 9588 (type), 9615, May
30, 19138.

The first specimen cited is with female flowers and mature, but old,
fruits, while the second is a juvenile stage of what is apparently the same
species, both being collected in the same place, on steep, dry, forested
ridges at an altitude of from 250 to 400 meters.

3, C, 5 Merrill: Philippine Plants, XII 307

The species is apparently most closely allied to Nepenthes copelandii
Merr., of Mindanao, differing, however, in many characters. It resembles
Nepenthes alata Blanco, at least the form I take to be a juvenile stage,
but the type number has quite wingless pitchers. Good differential charac-
ters are its comparatively short capsules and its short-petioled flowers,
together with its very obscurely nerved leaves, the lateral nerves, at times,
being quite obsolete.

ROSACEAE
ANGELESIA Korthals

ANGELESIA SPLENDENS Korth. in Ned. Kruidk. Arch. 3 (1854) 384;
Mig. Ann. Mus. Bot. Lugd. Bat. 3 (1867) 236; Boerl. & Koord. in
Ic. Bogor. 1 (1901) t. 96.

Licania angelesia Blume Mél. Bot. (1855) no. 2, Flora 41 (1858) 256.

Trichocarya splendens Miq. FI]. Ind. Bat. 1* (1855) 358.

Chrysobalanus splendens Mia. 1. ¢. in syn.

Parinarium nitidum Hook. f. Fl. Brit. Ind. 2 (1878) 310; King in
Journ. As. Soc. Beng. 667 (1897) 284.

MINDANAO, Province of Surigao, Piper 249, April, 1911. Samar, Bur.
Sci. 17481 Ramos, April, 1914, Phil. Pl. 1696 Ramos. PALAWAN, Phil.
Pl. 1224 Merrill, Elmer. SIBUYAN, Elmer 12180, 12214, distributed as a
new species of Parinarium.

I have examined the following extra-Philippine material: PENANG,
Curtis 14a, 147, Ridley s. n. MALAY PENINSULA, Setul, Ridley 1508.
BoRNEO, Sarawak, Tabuan, Native collector 784 Bureau of Science.

I think that there can be no doubt as to the validity of Korthals’s genus,
which is well figured by Boerlage and Koorders 1. c. The species and
genus were based on material collected in Borneo by Korthals, and as
Boerlage and Koorders explain, were joined by Miquel* with Diemenia
racemosa Korth. to form the genus Trichorcarya Mig. Later Miquel
himself* separated the two genera, which, as Boerlage and Koorders note,
has been overlooked by later authors. Bentham and Hooker f.’ retain
Miquel’s Trichocarya with two species, and Focke‘ maintains the genus
Angelesia with 2 or 3 species. Trichocarya splendens Miq., the type of
the genus Trichocarya, is identical with Angelesia splendens Korth., but
Miquel’s second species, Trichocarya ? racemosa Mig. (Diemenia racemosa
Korth.) is, according to Boerlage and Koorders, identical with the earlier
Parastemon urophyllus A. DC.

The only synonym I have added to the list given by Boerlage and
Koorders is Parinarium nitidum Hook. f., which is manifestly identical
with Korthals’s species. Hooker cites in the original description “Malacca,
Griffith, Maingay (Herb. Kew 619).-Distrib. Borneo;” King, among other
specimens, cites “Griffith 2047/1 ;” while Boerlage and Koorders cite in their
discussion Griffith 2047, and a part of their plate, the fruiting specimen,
is drawn from Griffith’s specimen. Hooker himself indicated Parinarium
nitidum as the type of an unnamed subgenus. King includes the species

‘Fl. Ind. Bat. 1* (1855) 358.

* Ann. Mus. Bot. Lugd. Bat. 3 (1867) 236.

°Gen. Plant. 1: 607.

‘Engl. & Prantl Nat. Pflanzenfam. 3° (1888) 60.

308 The Philippine Journal of Science 1915

in Parinarium with doubt, and expresses the opinion that, when mature
fruits are known, a separate genus might be formed with advantage for
the species, apparently not realizing that two generic names, Angelesia
Korth. and Trichocarya Migq., had already been proposed for an identical
species.

Angelesia splendens Korth. is represented by the following material:
MALAY PENINSULA, Setul, Ridley 15043: Malacca, Griffith 2047, Maingay
619, Derry 189, 1180: Perak, King’s collector 8599, 8680, 8711. PENANG,
Curtis 14a, 147, 858, King’s collector 1274, 1472, Ridley s. n. BORNEO,
Sarawak, Native collector 784, Bureau of Science; southern Borneo,
Korthals. SUMATRA, Koorders, and the Philippine material cited above.

PARINARIUM Aublet

PARINARIUM VILLAMILII sp. nov.

Arbor alta partibus junioribus inflorescentiisque exceptis
glabra; foliis coriaceis, oblongis, acuminatis, usque ad 16 cm
longis, nervis utrinque circiter 12, basi subtus 2-glandulosis,
nervis reticulisque utrinque minute scaberulis, ramulis junioribus
stipulisque longe adpresse ciliato-hirsutis, inflorescentiis axillar-
ibus, racemosis (vel paniculatis?), adpresse pubescentibus; fruc-
tibus ellipsoideis, 2.5 ad 3 cm longis.

A tree reaching a height of 27 m, glabrous except the in-
florescence and the younger parts. Branchlets slender, terete,
dark reddish-brown and somewhat glaucous, the very young ones
appressed-ciliate-hirsute with long, rather pale hairs, similar
hairs on the stipules, younger petioles, and near the base of the
leaf on the midrib beneath in young leaves. Leaves oblong, coria-
ceous, pale when dry, 12 to 16 cm long, 4 to 8 cm wide, rather
prominently acuminate, the acumen obtuse, base acute to nearly
rounded, with two small glands on the lower surface near the
junction with the petiole; lateral nerves about 12 on each side
of the midrib, prominent on the lower surface, curved, anastomos-
ing, these and the reticulations minutely scaberulous on both
surfaces; petioles 5 mm long or less, at maturity quite glabrous;
stipules oblong-lanceolate, acute or obtuse, about 1 cm long, with
scattered, appressed, long, pale hairs. Racemes (or panicles?)
axillary, appressed-pubescent, in fruit up to 9 cm in length.
Fruit ellipsoid, brown, 2.5 to 3 cm long (immature), prominently
verruculose, the pedicels stout, pubescent, about 5 mm long, the
persistent sepals pubescent on both surfaces, the tube villous
within, stamens apparently at least 30.

MINDANAO, District of Zamboanga, Margosatubig, For. Bur. 21863
Villamil, June 9, 1914, forested slopes, altitude about 80 meters.

Among the Philippine species most closely allied to Parinarium laurinum
A. Gray, but entirely different in vegetative characters.

¥ G5 Merrill: Philippine Plants, XII 309

PARINARIUM BICOLOR sp. nov.

Arbor circiter 10 m alta, subtus foliis ramulis inflorescen-
tiisque dense albido- vel ferrugineo-villosis; foliis coriaceis, usque
ad 7 cm longis, oblongo-ellipticis, basi acutis ad subrotundatis,
apice breviter obtuse acuminatis, supra castaneis, nitidis, glabris,
subtus albidis vel subferrugineis, densissime tomentosis villo-
sisque, nervis utrinque circiter 15, prominentibus; inflorescentiis
terminalibus confertis, dense multifloris, usque ad 4 cm diametro,
bracteis bracteolisque numerosis; floribus circiter 4 mm longis,
petalis sepalisque subaequilongis.

A tree about 10 m high, the younger parts and inflorescence
densely ferruginous-villous, the lower surface of the leaves very
densely white- or subferruginous-tomentose and more or less
villous. Branches dark-colored, terete, glabrous, the younger
ones ferruginous. Leaves alternate, coriaceous, oblong-elliptic,
4 to 7 cm long, 2 to 3 cm wide, base rounded to subacute, apex
shortly and obscurely blunt-acuminate, the upper surface cas-
taneous, glabrous and shining when dry, when young somewhat
ferruginous-tomentose, the lower surface white or subferrugi-
nous; lateral nerves about 15 on each side of the midrib, very
prominent on the lower surface, nearly straight; petioles densely
ferruginous-villous, 3 mm long or less; stipules inequilateral, de-
ciduous, oblong acuminate, up to 7 mm long, somewhat pubescent.
Panicles terminal, dense, many-flowered, densely ferruginous-
tomentose, up to 4 mm in diameter. Flowers yellowish-brown,
their pedicels short, densely villous, the bracteoles oblong-ovate,
acuminate, pubescent, concave, about 4 mm long. Calyx about
4 mm long, densely ferruginous-villous, the lobes 5, oblong,
acuminate, 1.2 to 2 mm long. Petals membranaceous, glabrous,
about 2 mm long, oblong-obovate. Calyx-tube densely villous
within. Style densely villous below, glabrous or nearly so above.
Fruit hard, irregular, dark brown, somewhat ovoid, about 3
em long and 2.3 cm in diameter, apex rounded.

MINDANAO, Province of Surigao (Ginituan Island), For. Bur. 28022
Razon, October 6, 1914, on slopes, altitude about 30 meters.

A species entirely different from the other Philippine representatives

of the genus, well characterized by its small leaves and its indumentum.
Its alliance is with the Malayan Parinarium costatum Blume.

PARINARIUM CORYMBOSUM (Blume) Mia. Fl. Ind. Bat. 1* (1855) 356.

Maranthes corymbosa Blume Bijdr. (1825) 89.

Maranthes multiflora Korth. Verh. Nat. Gesch. Bot. (1839-42) 259,
t. 70, fide Blume Mus. Bot. Lugd. Bat. 2: 98.

Parinarium griffithianum Benth. in Hook. Niger Fl. (1849) 334;
Mig. 1. c. 356; F.-Vill. Novis. App. (1880) 76; Vid. Sinopsis Atlas

310 The Philippine Journal of Science 1915

(1883) 25, t. 46, f. A; Koord. & Valeton Bijdr. Boomsoort. Java
5 (1900) 334; Perk. Frag. Fl. Philip. (1904) 118; Whitford in
Philip. For. Bureau Bull. 10* (1911) 34, pl. 14, 15.

Grymania salicifolia Pres] Epim. Bot. (1849) 193.

Parinarium salicifolium Miq. Fl. Ind. Bat. 1* (1855) 357; F.-Vill.
]. c. 76; Vid. Rev. Pl. Vase. Filip. (1886) 121.

Parinarium multiflorum Migq. 1. ¢c. 356.

Parinarium racemosum Vid. Cat. Pl. Prov. Manila (1880) 29.

This species is very widely distributed in the Philippines, and is re-
presented in the herbarium of the Bureau of Science by about 60 speci-
mens, from all parts of the Archipelago from northern Luzon to southern
Mindanao. What is manifestly the oldest specific name is here adopted,
for Blume’s species was reduced to Parinarium griffithianum Benth. by
Koorders and Valeton, which reduction is apparently correct. Blume’s
name, being by far the oldest one, must be adopted. The species is a
widely distributed one, extending from the Andaman Islands through
Sumatra, Borneo, and Java to New Guinea and northern Australia.

PARINARIUM LAURINUM A. Gray Bot. Wilkes U. S. Explor. Exped.
(1854) 490, pl. 55; K. Schum. & Lauterb. Fl. Deutsch. Schutzegeb.
Siidsee (1901) 341.

Parinarium scabrum Merr. in Philip. For. Bureau Bull. 1 (1903)
22, non Hassk.

Parinarium mindanaense Perk. Frag. Fl. Philip. (June, 1904) 119.

Parinarium racemosum Merr. in Govt. Lab. Publ. (Philip.) 17
(October, 1904) 19, non Vidal.

Parinarium curranii Merr. in Philip. Journ. Sci. 4 (1910) Bot. 264.

Luzon, Province of Albay, For. Bur. 10575 Curran, June, 1908.
MassaTE, Merrill 2614, May, 1903 (type of P. racemosum Merr.-P. currant
Merr.). PALAWAN, Malampaya Bay, For. Bur. 11253 Manalo, January,
1908, Merrill 7255, September, 1910. MINDANAO, Province of Surigao,
Ahern 375: Butuan Subprovince, La Paz, Miller s. n., July, 1910: District
of Davao, Warburg 14484, in herb. Berlin (type of P. mindanaense Perk.),
Elmer 11237, 12388.

There appears to be no doubt as to the specific identity of all the
Philippine material cited above, and equally as little doubt as to the
identity of the Philippine form with Parinarium lawrinum A. Gray. The
type of Parinarium lawrinum was from the Samoan Islands, and is well
figured and illustrated by Gray. The species is also represented in the
herbarium of the Bureau of Science by Vaupel 237 from Samoa (dis-
tributed as Parinarium insularum A. Gray), a specimen that agrees
perfectly with the original description and plate of Parinariwm lawrinum
A. Gray, but not with P. insularum A. Gray. This specimen in all essential
characters also agrees with the Philippine material.

Parinarium laurinum A. Gray is known from the Samoan, Fiji, Solomon,
and Admiralty Islands, and in consideration of the general character of
the Polynesian flora, its discovery in the Philippines is not at all surprising.
A considerable number of species are known only from the Philippines and
Polynesia, and a still greater number are represented in the two regions
by very closely allied forms. In a collection of plants made in Samoa
by Vaupel, which I have examined carefully, the similarity between the

K€. 5 Merrill: Philippine Plants, XII 311

constituents of the Samoan and Philippine flora is so very striking that
Vaupel’s entire collection might almost as well have been made at some
point in the Philippines; the percentage of difference as to species is no
greater between Vaupel’s Samoan collection and the general run of the
Philippine flora than is noted in current collections made in botanically
unexplored parts of the Philippines as compared with the known flora of
the Archipelago.

K. Schumann and Lauterbach note that oil produced by the fruits is
used in the Solomon Islands for caulking the seams of boats; in the
Philippines it is used by the natives of Agusan Valley, Mindanao, for
water-proofing bamboo and rattan baskets. The following note was sup-
plied by Dr. M. L. Miller, of the division of ethnology, Bureau of
Science, and communicated by him with botanical specimens: “The fruit
of the tabon-tabon tree, when mature, is full of a yellowish-white pulp,
that has about the hardness of a camote (sweet potato). On being rubbed
over a rough surface, such as a rattan plaiting, it fills the interstices,
assuming a chocolate color and drying within an hour to a hardness that
does not crack under a torrid sun. The coating of tabon-tabon on baskets,
etc., is frequently darkened in color with charcoal.”

Native names: tambon-tambon (Masbate); batobon (Palawan); tabun-
tabun (Albay) ; tabon-tabon (Surigao, Agusan).

As to nomenclature, the oldest valid specific name applied to the Philip-
pine form is Parinarium mindanaense Perk., the type of which I have
examined in the Berlin herbarium. Parinarium racemosum Merr., published
a few months later, and during the same year, is invalidated by P. race-
mosum Vid. (1880); P. curranii Merr. was proposed as a new name for
P. racemosum Merr., non Vidal.

PYGEUM Gaertner

PYGEUM EUPHLEBIUM sp. nov.

Arbor 10 m alta, subtus foliis ad costa ramulis inflorescen-
tiisque ferrugineo-villosis; foliis oblongis, coriaceis, usque ad
10 cm longis, integris, acuminatis, basi subacutis ad rotundatis,
nervis utrinque 8 ad 10, supra impressis, subtus valde prominen-
tibus, reticulis laxis, obscuris; racemis spiciformibus, brevibus,
fasciculatis, 1 ad 1.5 cm longis, bracteolis deciduis, fructibus
junioribus anguste ovoideis, plus minusve ferrugineo-villosis.

A tree about 10 m high, the young branches, inflorescence,
petioles, and the leaves along the midrib on the lower surface
rather prominently ferruginous-villous. Branches slender, te-
rete, dark brownish-purple, glabrous. Leaves oblong, coria-
ceous, 7 to 10 cm long, 2 to 4 cm wide, the upper surface shining,
brownish-olivaceous, the lower paler, brownish, the apex shortly
and sharply acuminate, the base subacute to rounded, with one
or two plane, dark-colored, small glands evident on the lower
surface near the base; lateral nerves impressed on the upper
surface, very prominent on the lower surface, 8 to 10 on each

312 The Philippine Journal of Science 1915

side of the midrib, curved upward near the margins, scarcely
anastomosing, the reticulations lax, obscure; petioles 7 to 10
mm long. Racemes short, ferruginous-villous, fascicled, spike-
like, 1 to 1.5 cm long, usually two or three in a fascicle in the
axils of leaves or of fallen leaves, the bracteoles small, deciduous;
young fruit narrowly ovoid, 5 mm long, ferruginous-villous with
scattered hairs, the rather slender style persistent.

MINDANAO, District of Lanao, Momungan, For. Bur. 20619 Miranda,
May 20, 1913, along Agus River, sea level to 80 meters in altitude.

A species allied to Pygeum merrillianum Koehne, but entirely different
from that form, well characterized by its ferruginous-villous, short, fas-,

cicled, spikelike racemes, its pubescent young fruit, indicating a pubescent
ovary, and its very prominently nerved leaves.

PYGEUM MONTICOLUM sp. nov.

Arbor circiter 10 m alta, partibus junioribus ferrugineo-
pubescentibus; foliis crasse coriaceis, oblongis, usque ad 10 cm
longis, utrinque subaequaliter angustatis, apice acuminatis, basi
acutis, eglandulosis; nervis utrinque circiter 8, subtus prominen-
tibus; racemis axillaribus, solitariis, usque ad 5 cm longis; fruc-
tibus globosis, glabris vel subglabris, 2 cm diametro.

A tree about 10 m high, the younger parts, branchlets, young
leaves, etc. rather densely ferruginous-villous. Branches terete,
glabrous. Leaves thickly coriaceous, oblong or somewhat oblong-
lanceolate, 8 to 10 cm long, 2.5 to 4 cm wide, entire, about equally
narrowed to the distinctly acuminate apex and to the acute base,
the basal glands none, the upper surface dark olivaceous, prom-
inently shining, the lower paler, dull, very slightly pubescent
along the midrib at maturity; lateral nerves about 8 on each
side of the midrib, very prominent on the lower surface, some-
what ascending, distinctly anastomosing, the reticulations rather
fine, distinct; petioles 1 to 1.5 cm long. Racemes axillary, soli-
tary, up to 5 cm long, ferruginous-pubescent. Fruits globose,
glabrous or nearly so, 2 cm in diameter.

LuZON, Province of Bataan, Mount Mariveles, Whitford 12038, April
7, 1905, forested slopes, altitude about 1,060 meters.

A species best characterized by its thickly coriaceous, eglandular leaves
and its very large fruits.

PYGEUM MEGAPHYLLUM sp. nov.

Arbor parva, racemis exceptis glabra; foliis ellipticis vel ovato-
ellipticis, chartaceis, circiter 20 cm longis, breviter acuminatis,
basi subrotundatis vel subacutis, glandulis 2 magnis instructis;
racemis solitariis, axillaribus vel in axillis defoliatis, 6 ad 9

mC, 5 Merrill: Philippine Plants, XII 313

cm longis, dense fulvo-pilosis; floribus in anthesis 1 ad 1.5 em
diametro.

A small tree about 8 m high, glabrous except the inflorescence.
Branches brown, smooth, slender, terete, the ultimate ones 2 to
3mm in diameter. Leaves elliptic or elliptic-ovate, chartaceous,
brown when dry, the lower surface paler than the upper, some-
what shining, about 20 cm long, 10 cm wide, the apex rather
broadly acuminate, the acumen blunt, the base rounded or sub-
acute, sometimes a little decurrent, supplied at the junction with
the petiole with a pair of very prominent oblong glands which
are open on the lower surface, slightly projecting on the upper
one; lateral nerves about 10 pairs, rather prominent; petioles
2 cm long. Racemes axillary, 6 to 9 cm long, rather densely
pubescent with pale fulvous hairs; bracteoles oblong-lanceolate,
acuminate, pubescent, 3.5 mm long, deciduous; pedicels about
2mm long. Flowers white, in anthesis 1 to 1.5 cm in diameter.
Calyx-tube funnel-shaped, 3 mm long, 4 mm in diameter at the
throat, the lobes 6, ovate-lanceolate, somewhat acuminate, pu-
bescent, 2 to 2.5 mm long. Petals 6, suborbicular to orbicular-
ovate, 2 mm long, very densely pubescent on both surfaces.
Stamens indefinite, the filaments 2 to 6 mm long. Ovary gla-
brous, ovoid; style 5 mm long.

LUZON, Province of Laguna, Dahican River back of San Antonio, Bur.
Sci. 14923 Ramos, June, 1912.

A distinct species, well characterized by its unusually large leaves with

their very prominent basal glands, and by its flowers which are con-
siderably larger than those of any other known Philippine species.

LEGUMINOSAE
ORMOSIA Jack

ORMOSIA VILLAMILII sp. nov. § Amacrotropis.

Arbor circiter 25 m alta, glabra, inflorescentiis ignotis; foliis
imparipinnatis, circiter 25 cm longis, foliolis circiter 7, alternis
vel inferioribus suboppositis, chartaceis, nitidis, oblongis, usque
ad 11 cm longis, obtuse acuminatis, basi rotundatis; leguminibus
anguste oblongis, tenuibus, 11 cm longis, 2.5 ad 3 cm latis, semi-
nibus 3 vel 4.

A tree about 25 m in height, quite glabrous so far as the ma-
terial shows (inflorescence not seen). Branches slender, brown-
ish, shining, terete, lenticellate. Leaves alternate, about 25 cm
long, the rachis and petiole dark brown or nearly black when dry ;
leaflets usually 7, alternate, or the lower ones subopposite,

314 The Philippine Journal of Science 1915

shining, mostly oblong, the lower surface a little paler than the
upper one, 6 to 11 cm long, 3 to 4.5 cm wide, the base rounded, the
apex blunt-acuminate, the acumen distinctly retuse; lateral nerves
5 to 7 on each side of the midrib, slender, not prominent; peti-
olules 4 to 5 mm long, the rachis bearing the terminal leaflet
produced. Inflorescence and flowers not seen. Pods narrowly
oblong, thin, pale brown or grayish, smooth and shining, about
11 cm long, 2.5 to 3 em wide, acute or acuminate, base decurrent
into a short stipe, the margins narrowly keeled, apparently
tardily dehiscent, containing three or four seeds.

MINDANAO, District of Zamboanga, opposite Olutanga Island, For. Bur.
18290 Foxworthy, DeMesa, & Villamil, May 16, 1912, on forested slopes
near the sea, altitude about 20 meters.

There is some doubt as to the correctness of referring the species to
Ormosia in the absence of flowers, but it is probably correct, and probably
referable to the section Amacrotropis. Its Moro name, bahay, is also sig-
nificant, for in other parts of the Archipelago this is the common name of
Ormosia calavensis Azaola. The seeds, in the specimens available, were
destroyed by insects before the time of collection.

SINDORA Miquel

SINDORA INERMIS sp. nov.

Arbor alta, partibus junioribus leviter pubescentibus; foliolis
6, chartaceis vel subcoriaceis, usque ad 12 cm longis, subellipticis,
breviter obtuse acuminatis, basi subacutis ad rotundatis aliquis
inaequilateralibus; stipulis late acinaciformibus, 2 ad 2.5 cm
longis; leguminibus compressis, inermis, inaequilateraliter subel-
lipticis, circiter 7 cm longis et 5 cm latis, brevissime apiculatis.

A tall tree, the younger parts sparingly pubescent, otherwise
glabrous (inflorescence not seen). Branches slender, terete,
brown. Leaves about 25 cm long, each with 6 leaflets, the leaflets
chartaceous to subcoriaceous, subelliptic, somewhat shining,
brownish-olivaceous, 8 to 12 cm long, 5.5 to 6.5 em wide, some of
them more or less inequilateral, apex shortly and obtusely
acuminate, base subacute to rounded; lateral nerves slender, 15 or
more on each side of the midrib, both surfaces rather distinctly
and densely subfoveolate-reticulate; stipules broadly scimitar-
shaped, falcate, coriaceous, 2 to 2.5 cm long, shortly acuminate.
Flowers not seen. Fruits compressed, somewhat inequilateral,
about 7 cm long, 5 cm wide, brown, glabrous, with a short excen-
tric apiculus at the apex, the valves unarmed, thickly coriaceous.

MINDANAO, District of Cotabato, Sapu, Sarangani Bay, For. Bur. 238054

Ferraris, October 3, 1914, growing at sea level near the mangrove, locally
known as gayugalo, oil-producing like Sindora supa. A sterile specimen,

x, C, 5 Merrill: Philippine Plants, XII 315

For. Bur. 23039 Ferraris from the same locality, dry slopes, altitude about
50 meters, probably represents the same species.

The species is decidedly similar to Sindora supa Merr. in its vegetative
characters, but differs in having its pods entirely unarmed.

BURSERACEAE
CANARIUM Linnaeus

CANARIUM CAUDATIFOLIUM sp. nov.

Arbor glabra (floribus ignotis), circiter 10 m alta; foliis
circiter 70 cm longis, foliolis 11, subcoriaceis, oblongis ad oblongo-
lanceolatis, integris, usque ad 20 cm longis, apice caudato-acumi-
natis, basi acutis, plus minusve inaequilateralibus, in siccitate
pallidis nitidis, nervis utrinque circiter 15, patulis, prominen-
tibus; infructescentiis 22 cm longis, cymosis, longe pedunculatis;
fructibus oblongo-ovoideis vel oblongis, haud angulatis, inaequila-
teralibus, 4 cm longis, obtusis, 1-locellatis, pericarpio crustaceo,
in siccitate valde rugoso.

A glabrous tree (flowers unknown), about 10 m high, the
branches and branchlets, petioles, petiolules, axis, and branches
of the infructescence reddish-brown, with numerous small len-
ticels. Ultimate branches subterete, about 1 cm in diameter,
longitudinally wrinkled when dry. Leaves alternate, about 70
em long; leaflets 11, oblong to oblong-lanceolate, 15 to 22 cm long,
5.5 to 7 em wide, entire, subcoriaceous, the apex prominently
caudate-acuminate, the acumen 1.5 to 2 cm long, slender, blunt,
the base acute or subacute, somewhat inequilateral, the upper
surface pale and shining when dry, the lower a little browner
than the upper; lateral nerves about 15 on each side of the midrib,
prominent, spreading, somewhat curved upward, anastomosing,
the reticulations lax; petiolules about 2 cm long. Infructescence
axillary, about 22 cm long, cymose, the peduncles at least 12 cm
long, above somewhat dichotomously branched, the pedicels about
1.5 ecm long. Fruits oblong to oblong-ovoid, subterete, not at
all angled, inequilateral, about 4 cm long, 1.6 cm wide, base
rounded, apex obtuse, 1-celled, the pericarp crustaceous, yellowish,
and prominently wrinkled when dry.

MINDANAO, Province of Surigao, Ilaya Mainit, For. Bur. 22827 Ponce,
August 16, 1914, on semiopen slopes, altitude about 15 meters, locally
known as pilihanay.

A species well characterized by its rather large, prominently caudate-
acuminate, entire leafiets, its cymose infructescence, and its subterete,
inequilateral, rather large fruits, the pericarp thin, crustaceous, and prom-

inently wrinkled when dry. Its alliance is apparently with Canariwm
nitens Merr., although it is entirely different from that species.

316 The Philippine Journal of Science 1915

POLYGALACEAE
XANTHOPHYLLUM Roxburgh
XANTHOPHYLLUM PSEUDOSTIPULACEUM sp. nov.

Arbor circiter 10 m alta, inflorescentiis exceptis glabra; foliis
coriaceis, oblongis vel oblongo-ovatis, usque ad 7 cm longis, basi
acutis, apice obtusis ad obtuse acuminatis, subtus pallidis, nervis
vix prominentibus, axillis petiolorum squamis binis oblongis
coriaceis 5 ad 7 mm longis instructis; racemis puberulis, nodis
valde incrassatis; fructibus globosis, 1.5 cm diametro.

A tree about 10 m high, glabrous except the inflorescence.
Branches terete, smooth, brown or yellowish-brown, quite glab-
rous. Leaves rather thickly coriaceous, oblong to oblong-ovate,
5 to 7 cm long, 1.5 to 2.5 cm wide, the base acute, apex obtuse
to obtusely acuminate, the upper surface somewhat yellowish
when dry, shining, the lower much paler but scarcely glaucous;
lateral nerves about 7 on each side of the midrib, not prominent,
anastomosing, the reticulations rather close; petioles 5 to 8 mm
long; axillary scales stipulelike, oblong, coriaceous, closely ap-
pressed in pairs, 2 scales in each axil, oblong, obtuse, persistent,
5 to 7 mm long. Racemes terminal, simple, rarely panicled,
puberulent, in fruit up to 6 cm long, the internodes very short,
the nodes strongly thickened, each with a prominent scar of the
fallen pedicel. Flowers not seen. Fruit globose, smooth, about
1.5 ecm in diameter.

Luzon, Province of Isabela, Palanan Bay, Bur. Sci. 21185 Escritor
(type), June, 1913: Province of Nueva Ecija, Mount Macasantol, For.
Bur. 22408 Alvarez, February, 1911.

The species is strongly characterized by its unusually large, coriaceous,
persistent, oblong, 5 to 7 mm long axillary scales and its comparatively

small leaves. It does not appear to be closely allied to any previously
described form, at least from the Philippines.

AQUIFOLIACEAE
ILEX Linnaeus

ILEX GUERREROII sp. nov. § Aquifolium, Lemurenses.

Frutex vel arbor parvus, glaber; foliis coriaceis, integris vel
subintegris, oblongis vel oblongo-ellipticis, usque ad 7 cm longis,
acutis, obscure acuminatis vel obtusis, nervis utrinque circiter 6,
supra obsoletis; floribus ¢ axillaribus, fasciculatis, 4-meris,
pedicellis circiter 4 mm longis, bibracteolatis; ovario 4-loculare.

A shrub or small tree, glabrous throughout. Branches terete,
rather stout, somewhat wrinkled when dry, pale gray. Leaves

#20: 5 Merrill: Philippine Plants, XII 317

coriaceous, oblong to narrowly oblong-elliptic, shining when
dry, 5 to 7 cm long, 1.5 to 2.8 cm wide, entire or sometimes
obscurely and distantly subcrenulate, the apex acute, slightly
acuminate, or blunt, the base usually somewhat decurrent-acu-
minate; nerves about 6 on each side of the midrib, obsolete above,
spreading, anastomosing, the reticulations very few, lax; petioles
5 to 10 mm long. Pistillate flowers axillary, fascicled, two to
six in each axil, the pedicels about 4 mm long, each with a pair
of ovate, acuminate bracteoles at the lower one-fourth or one-
third. Flowers 4-merous. Calyx-lobes about 1.5 mm wide, 1
mm long, rounded. Petals free, 3 to 3.5 mm long, about 2 mm
wide, oblong-ovate, rounded. Staminodes about 2 mm long.
Ovary rhomboid-ovoid, 4-celled; stigma large, sessile, 1.5 mm in
diameter. Very immature fruit somewhat angled, wrinkled, 4
mm long, 4-celled.

LuzoN, Province of Rizal, mountains back of San Mateo, comm. Leon
Guerrero, 1910.

A species apparently allied to Ilex swmatrana Loesener, but quite dif-
ferent from that form.

ILEX SUBCAUDATA sp. nov. § Thyrsoprinus, Indico-Malaicae.

Frutex vel arbor parvus, glaber; foliis ovato-ellipticis, usque
ad 7 cm longis, integris, subcoriaceis, siccitate nitidis, breviter
petiolatis, utrinque acuminatis, apice subcaudatis, acuminibus
obtusis, circiter 1 cm longis; racemis ¢ axillaribus, solitariis
vel binis, quam folia paulo brevioribus; floribus 4-, rariter 5-
meris.

An erect, glabrous shrub or small tree. Branches rather slen-
der, terete, wrinkled when dry, reddish-brown or grayish, not
lenticellate. Leaves scattered, the internodes 1 to 3.5 cm long,
ovate-elliptic, subcoriaceous, shining when dry, not punctate or
glandular, somewhat pale, 5 to 7 cm long, 2 to 3.5 cm wide,
entire, the apex rather abruptly subcaudate-acuminate, the acu-
men about 1 cm long, blunt, straight or somewhat falcate, the
base somewhat acuminate, the margins not, or very slightly, re-
curved; nerves about 6 on each side of the midrib, slender, not
prominent, ascending, anastomosing, the reticulations lax; peti-
oles 3 to 4 mm long; stipules, if present, very early deciduous.
Staminate racemes solitary or in pairs in the leaf-axils, 3 to 5 cm
long, each with from 10 to 20, slenderly pedicelled flowers, the
pedicels 4 to 5 cm long, each subtended by a small, ovate, obtuse
bracteole less than 1 mm long. Flowers 4-, rarely 5-merous.
Calyx-lobes ovate, obtuse, 0.5 to 0.8 mm long. Corolla-tube 1
mm long, the lobes oblong-ovate, obtuse, 1.5 mm long. Anthers

318 The Philippine Journal of Science 1915

0.8mm long. Ovary rudimentary. Pistillate flowers and fruits
unknown.

LuzoN, Province of Rizal, mountains back of San Mateo, comm. Leon
Guerrero, 1910.

A species probably as closely allied to Jlex spicata Bl. as to any other
species, but abundantly distinct.

ILEX PACHYPHYLLA sp. nov. § Byronia, Eubyronia.

Species J. cymosae affinis, differt foliis crassissime coriaceis,
apice late rotundatis vel obtusis retusisque, vix acuminatis, flo-
ribus majoribus, ovario 4- vel 5-locellato.

A tree about 10 m high, glabrous except the somewhat pubes-
cent inflorescence. Branches terete, stout, dark-colored. Leaves
very thickly coriaceous, elliptic to obovate-elliptic, 5 to 10 cm
long, 2.5 to 6 cm wide, entire, the apex broadly rounded to obtuse,
usually more or less retuse, never acuminate, the base somewhat
narrowed, acute or a little decurrent-acuminate, the margins
strongly revolute, the upper surface olivaceous when dry, some-
what shining, the lower shining, much paler; lateral nerves 7
or 8 on each side of the midrib, slender, not prominent, the
reticulations obsolete or nearly so; petioles 1 to 1.5 cm long,
dark reddish-brown. Inflorescence cymose, sparingly cinereous-
pubescent, the cymes in the upper axils pseudo-terminal, many-
flowered, 2 to 5 em long and wide. Flowers white, 5-merous,
the pedicels and calyx somewhat puberulent, the calyx-lobes
orbiculate-ovate, about 1.5 mm long, somewhat serrate-lacerate.
Petals 5, oblong to oblong-ovate, rounded, about 3 mm long.
Ovary 4- or 5-celled, glabrous, oblong, narrowed upward. Young
fruit narrowly ovoid.

Luzon, Province of Tayabas, Mount Binuang, back of Infanta, Bur. Sci.

9400 Robinson, August 28, 1909, in the mossy forest, altitude about 980
meters.

ILEX BRUNNEA sp. nov. § Thyrsoprinus, Indico-Malaicae.

Frutex ut videtur epiphyticus, glaber, ramis ramulisque pal-
lidis; foliis alternis, integris, coriaceis, oblongo-ovatis ad ellip-
tico-lanceolatis, utrinque acuminatis vel basi acutis, usque ad 5
cm longis, petiolatis, in siccitate nitidis, brunneis, nervis latera-
libus utrinque circiter 6, tenuibus, indistinctis; inflorescentiis ¢
axillaribus, solitariis, racemosis, racemis numerosis, multifloris,
2 ad 4 cm longis; floribus 4- vel 5-meris, petalis 2 mm longis.

Apparently an epiphytic shrub, quite glabrous or the racemes
very slightly and obscurely pubescent. Branches pale gray, te-
rete, slightly wrinkled when dry, the branchlets similar, some-
what angled, 1 to 1.2 mm thick. Leaves alternate, 3 to 5 cm

X, C, 5 Merrill: Philippine Plants, XII 319

long, 1.4 to 3 cm wide, oblong-ovate to elliptic-lanceolate, about
equally narrowed to the somewhat blunt-acuminate apex and to
the acute or somewhat acuminate base, entire, coriaceous, when
dry strongly shining on the upper surface, a little paler on the
lower surface, uniformly dark brown, the lower surface very
minutely, densely, and obscurely puncticulate, and with few, scat-
tered, larger, dark glands; lateral nerves about 6 on each side
of the midrib, very slender, obscure, anastomosing, the reticu-
lations obsolete or nearly so; petioles about 5 mm long. Male
racemes axillary, solitary, rarely 2 in an axil, numerous, 2 to
4 ecm long, many-flowered, glabrous or very obscurely and spar-
ingly pubescent, the flowers sometimes solitary at the nodes of
the racemes, more often in fascicles of from 2 to 6, their pedicels
slender, up to 5 mm long, the bracteoles narrowly ovate, 0.5
mm long. Flowers white, 4- and 5-merous in the same raceme.
Calyx glabrous, about 2.5 mm in diameter, the lobes broadly
elliptic-ovate to suborbicular, broadly rounded, 0.5 mm long.
Petals narrowly oblong, 2 mm long, rounded or obtuse, glabrous.
Filaments 2 mm long; anthers somewhat ovoid, 0.8 mm long.

LuzON, Province of Tayabas, Mount Pular, Bur. Sci. 19461 Ramos,
January 15, 1913.

Said by the collector to grow on trees at the summit of the mountain.
Most closely allied to Ilex halconensis Merr., from which it differs in its
smaller, obscurely nerved, brown leaves, more numerous racemes, and
smaller flowers.

CELASTRACEAE
MICROTROPIS Wallich

MICROTROPIS PLATYPHYLLA sp. nov.

Arbor glabra; foliis coriaceis, in siccitate pallidis, usque ad
20 cm longis, petiolatis, breviter acuminatis, basi decurrento-
acuminatis, nervis utrinque 10 ad 12; cymis axillaribus, extra-
axillaribusque, pedunculatis, usque ad 2.5 cm longis, multifloris,
floribus 5-meris, sessilibus, capitato confertis; capsulis ellipsoi-
deis, acutis vel apiculato-acuminatis, circiter 1.5 cm longis.

Apparently a tree, quite glabrous. Branches terete, reddish-
brown or pale when dry, the tips of the branchlets somewhat
compressed. Leaves large, coriaceous, pale, and somewhat shin-
ing when dry, the lower surface somewhat brownish, 12 to 20
em long, 5.5 to 12 cm wide, subequally narrowed at both ends,
elliptic-ovate, apex shortly and sharply acuminate, base decur-
rent-acuminate; lateral nerves 10 to 12 on each side of the mid-
rib, slender, anastomosing, the reticulations rather close, fine,
distinct ; petioles stout, 1.5 to 2.5 cm long. Cymes axillary and

133910——3

320 The Philippine Journal of Science 1915

extra-axillary, peduncled, mostly 3-branched, up to 2.5 cm long,
each subtended by several, coriaceous, linear bracts 2 to 5 mm
in length. Flowers numerous, 5-merous, capitate at the tips of
the branches, the heads up to 7 mm in diameter. Sepals obovate-
orbicular, concave, externally prominently wrinkled-reticulate,
about 2 mm in diameter, glabrous except the somewhat furfu-
raceous-incised margins. Corolla about 3.5 mm long, the tube
short, the lobes elliptic-obovate, rounded, about 2 mm long, glab-
rous; filaments 1 mm long; anthers about 0.5 mm long. Ovary
brown, glabrous, 2 mm in length. Capsule apparently some-
what fleshy when fresh, ellipsoid, about 1.5 cm long, 2-valved,
the pericarp thickly coriaceous, apex acute or apiculate-acumi-
nate; seed solitary, about 1 cm long, brown, and much wrinkled
when dry.

Luzon, Province of Rizal, Montalban, Loher 5779 (type), in flower,

March, 1906, 5774, in fruit, July, 1905.
A species readily recognizable by its unusually large leaves.

EUONYMUS Linnaeus

EUONYMUS OLIGANTHUS sp. nov.

Arbor glabra circiter 8 m alta; foliis chartaceis ad subcoria-
ceis, oblongo-oblanceolatis vel oblongo-ellipticis ad obovatis, usque
ad 10 cm longis, pallidis, nitidis, sursum plus minusve serrato-
crenulatis, utrinque subaequaliter angustatis, basi cuneatis, apice
breviter obtuse acuminatis, nervis utrinque circiter 6, tenuibus;
cymis axillaribus, tenuibus, longe pedunculatis, usque ad 9 cm
longis, dichotomis, paucifloris; fructibus obovoideis, circiter 6
mm longis, vix sulcatis, laevis.

A glabrous tree about 8 m high, the branches pale yellowish-
green, slender, terete. Leaves opposite, chartaceous to subcoria-
ceous, pale yellowish-green on both surfaces when dry, shining,
oblong-elliptic to oblong-oblanceolate or obovate, 6 to 10 em long,
2 to 4.5 cm wide, subequally narrowed to the cuneate base and
to the shortly broad-acuminate apex, the upper part distinctly
serrate-crenulate with small distant teeth, below entire; lateral
nerves about 6 on each side of the midrib, slender, ascending,
anastomosing, the reticulations lax, obscure; petioles 4 to 7 mm
long. Cymes axillary, slender, few-flowered, dichotomous, in
fruit up to 9 cm long. Flowers mostly in groups of threes at
the tips of the branchlets, the very young buds obovoid, sepals
and petals more or less ciliate, the latter at maturity possibly
lacerate. Mature fruits crimson, smooth, obovoid when dry,
about 6 mm long, somewhat apiculate by the persistent style,
not or very obscurely longitudinally suleate when dry, when

AG, 5 Merrill: Philippine Plants, XII 321

fresh not at all so, the persistent calyx thickly coriaceous, 4 to 5
mm in diameter, the lobes reniform, 2 mm long, about 3 mm
wide, glabrous.

Luzon, Benguet Subprovince, Baguio, Merrill 9644, May, 1914, in
thickets, limestone region, Sablan trail, altitude about 1,300 meters.

A species allied to Euonymus viburnifolius (Juss.) Merr. (E. philip-
pinensis Merr.), differing in its smaller leaves, slender, elongated, very
few-flowered cymes, and smaller fruits which are not sulcate or lobed.

GLYPTOPETALUM Thwaites
GLYPTOPETALUM LOHERI sp. nov.

Frutex vel arbor glabra; foliis subcoriaceis, ellipticis ad ellip-
tico-ovatis, integris, breviter petiolatis, usque ad 10 cm longis, in
siccitate pallidis, acuminatis, basi acutis, nervis utrinque 5 vel
6, tenuibus, obscuris, reticulis subobsoletis; cymis axillaribus,
brevibus, paucifloris ; fructibus globosis, 6 ad 10 mm diametro, 1-
ad 4-locellatis.

A glabrous shrub or small tree, the branches terete, reddish-
brown, the younger ones pale straw-colored. Leaves rather un-
equal in size, opposite, subcoriaceous, pale when dry, elliptic to
ovate-elliptic, 6 to 10 cm long, 3 to 6 cm wide, subequally nar-
rowed to the acute base and to the sharply acuminate apex, shin-
ing, entire; lateral nerves 5 or 6 on each side of the midrib,
slender, obscure, the reticulations nearly obsolete; petioles 3 to 4
mm long. Cymes axillary, short, the peduncles 1 cm long or less,
unbranched or with two very short branchlets. Flowers ap-
parently sessile, crowded, few. Fruits globose, 6 to 10 mm in
diameter, not sulcate, 1- to 4-celled, 1- to 4-seeded, the pericarp
pale when dry, thickly coriaceous; seeds brown, about 7 mm
long, the apical one-third surrounded by a thin, membranaceous,
brittle, brown aril. Persistent sepals coriaceous, reniform, 4,
entire, 2 to 3 mm wide.

LuzOoN, Province of Rizal, Montalban, Loher 5765 (type), 5796, February
and December, 1904.

A species characterized by its pale, obscurely nerved leaves, its nearly
obsolete reticulations, and its globose, not sulcate fruits.

GLYPTOPETALUM MARIVELENSE sp. nov.

Arbor parva, glabra, 3 ad 7 m alta; foliis oblongis ad oblongo-
lanceolatis, chartaceis ad subcoriaceis, usque ad 13 cm longis,
in siccitate pallidis, utrinque aequaliter angustatis, basi acutis,
apice acuminatis, margine crenulato-serrulatis, nervis utrinque
6 vel 7, tenuibus, laxe anastomosantibus, reticulis distinctis,
laxis; inflorescentiis axillaribus, tenuibus, usque ad 10 cm longis,
plerumque breviter furcatis, ut videtur paucifloris; fructibus de-

322 The Philippine Journal of Science 1915

presso-globosis, circiter 1.5 cm diametro, in siccitate brunneis,
rugosis, 3- vel 4-locellatis, sepalis 4, persistentibus integris.

A shrub or small tree 3 to 7 m high, quite glabrous. Branches
terete, slender, straw-colored or yellowish-green. Leaves op-
posite, chartaceous to subcoriaceous, oblong to oblong-lanceolate,
pale and shining when dry, 7 to 13 cm long, 2.5 to 4.5 cm wide,
subequally narrowed to the acute base and the shortly acuminate
apex, the margins distinctly crenulate-serrate, the midrib very
prominent; lateral nerves 6 or 7 on each side of the midrib,
slender, laxly anastomosing, the reticulations slender, lax; peti-
oles about 5 mm long. Inflorescence axillary, solitary, slender,
usually forked near the apex, apparently few-flowered, up to
10 cm long in fruit, the bracts in pairs, lanceolate, acuminate,
about 1.5 mm long. Flowers 4-merous. Fruits depressed-glo-
bose, brown when mature, about 1.5 cm in diameter, 3- or 4-celled,
the pericarp coriaceous, wrinkled. Seeds 1 in each cell, attached
near the apex of the cell, brown, about 8 mm long, the upper
one-third surrounded by a thin brown aril.

Luzon, Province of Bataan, Mount Mariveles, Elmer 6644 (type),
November, 1904, Whitford s. n., For. Bur. 2689 Meyer, February, 1905
(the two latter with mature fruits): Province of Tayabas, For. Bur. 10306
Curran, April, 1908. From this can scarcely be distinguished For. Bur.
6457 Everett, from Cebu, February, 1907, and this specimen probably
represents a form of the same species. In forests, altitude about 800
meters.

Duplicates of some of the above specimens were studied by me in the
Kew Herbarium in December, 1907, and the memorandum then made was
attached to one of the sheets in the herbarium of the Bureau of Science
to the effect that the material represented a new species of Huonymus; later
Mr. Elmer added the specific name marivelensis (sub. Huonymus) to our
specimen of his No. 6644. A further study of the material leads me to
consider that the species should be treated under the genus Glyptopetalum.

Var. EUPHLEBIUM var. nov.
A typo differt foliis valde nervosis, nervis, lateralibus reti-
culisque primariis subtus admodum prominentibus.

Luzon, Province of Zambales, Mount Tapulao, For. Bur. 8108 Curran
& Merritt, December 15, 1907, in forests, altitude 100 to 1,400 meters.

ICACINACEAE
PHYTOCRENE Wallich

PHYTOCRENE OBOVOIDEA sp. nov.

Frutex alte scandens; foliis ovatis, coriaceis, glabris, in siccitate
pallidis, usque ad 16 cm longis, acuminatis, integris, basi cordatis,
5-nervis, nervis lateralibus utrinque 3 vel 4, subtus valde prom-
inentibus; capitulis 2 binis, pedunculatis, globosis, circiter 2 cm

iC, 6 Merrill: Philippine Plants, XII 323

diametro, floribus 4-meris, sub fructu globosis, circiter 10 cm
diametro, fructibus obovoideis vel oblongo-obovoideis, circiter 3.5
cm longis, apice abrupte rotundatis, depressis, abrupte brevissime
apiculatis, densissime retrorse hirsutis.

A scandent shrub apparently of large size, the fruit-bearing
branches up to 1.5 cm in diameter, pale brownish, prominently
wrinkled when dry. Leaves ovate, coriaceous, pale, 12 to 16 cm
long, 7 to 10 cm wide, entire, the upper surface glabrous, shining,
the lower surface paler, glabrous, apex acuminate, base prom-
inently cordate; basal nerves 5, the lateral ones about 4 pairs
(or 3 above the base), very prominent, looped-anastomosing near
the margins, the reticulations very distinct; petioles 5 to 7 em
long, very slightly hirsute. Female heads, just after anthesis,
in pairs, apparently from the axils of fallen leaves, brown,
globose, about 2 cm in diameter, their peduncles brown-hirsute,
up to 4 cm in length. Flowers 4-merous, the calyx 4 to 5 mm
long, lobes broadly ovate, acute or somewhat acuminate, up to 1.5
mm long, externally densely appressed-hirsute. Ovaries densely
hirsute, when slightly developed (6 to 8 mm long) distinctly 3-
or 4-toothed at the apex, the base narrowed. Heads in fruit (im-
mature) globose, 8 to 10 cm in diameter, brown, the drupes very
numerous, obovoid to oblong-obovoid, about 3.5 cm long, the apex
up to 2 cm in diameter, angular by mutual pressure, gradually
narrowed from the apex to the base, the apex itself abruptly
rounded, usually distinctly depressed, and with a short, stout,
blunt apiculus 2 mm long or less, all parts very densely hirsute
with stiff, brown, appressed, reflexed hairs.

BILIRAN, Bur. Sci. 18882 McGregor, June 28, 1914, in forests.

This species is manifestly allied to Phytocrene blancoi (Azaola) Merr.,
which in turn is doubtfully distinct from Phytocrene macrophylla Blume;
it differs, however, in its leaves being glabrous beneath, and in its entirely
differently shaped drupes which are not gradually narrowed upward to
the tip, but are obovoid or oblong-obovoid, abruptly rounded and depressed,

gradually narrowed from the apex to the base. In the shape of its drupes
it is equally distinct from Phytocrene dasycarpa Miq.

TILIACEAE
COLUMBIA Persoon
COLUMBIA SUBINTEGRA sp. nov.

Arbor circiter 35 m alta, inflorescentiis exceptis glabra; foliis
ovatis ad oblongo-ovatis, coriaceis, usque ad 17 cm longis, utrin-
que glabris, nitidis, concoloribusque, integris vel leviter undulatis,
aequilateralibus, basi late rotundatis, apice acuminatis; paniculis
terminalibus, pyramidatis, pubescentibus; fructibus 2 cm dia-
metro, suborbicularis, 5-alatis.

324 The Philippine Journal of Science 1915

A tree about 35 m high, glabrous except the inflorescence.
Branches slender, terete, brownish. Leaves coriaceous, equilat-
eral, ovate to oblong-ovate, 14 to 17 cm long, 7 to 10 cm wide,
of the same color, brownish and shining on both surfaces, en-
tirely glabrous, the base broadly rounded, the apex acuminate,
the margins entire or somewhat undulate, or even obscurely un-
dulate-toothed in the upper part; nerves 6 on each side of the
midrib, prominent, the reticulations slender, but distinct; petioles
2to2.5cemlong. Panicles terminal, pyramidal, uniformly pubes-
cent, about 25 cm long, the branches scattered, the lower ones
10 cm long. Flowers not seen. Fruit orbicular in outline (lat-
eral view), 2 cm in diameter, slightly pubescent or becoming
glabrous, somewhat retuse at both base and apex, equally 5-
winged, the wings about 8 mm wide.

MINDANAO, District of Zamboanga, Talisay, Yor. Bur. 13776 Foxworthy,
DeMesa, & Villamil, June 18, 1912, in forests, altitude about 20 meters.

The species is locally known as malacayan a name applied also to other
unrelated species, and is allied to Columbia longipetiolata Merr. of Luzon,
from which it differs in its entire or nearly entire leaves. The fruit is red

when fresh, the bark very tough and stringy as in other species of the
genus, and the wood has a slightly pungent odor.

THEACEAE
TERNSTROEMIA Mutis

TERNSTROEMIA JAPONICA Thunb. in Trans. Linn. Soc. 2 (1794) 335;
Dyer in Hook. f. Fl. Brit. Ind. 1 (1874) 280.
Adinandra coriacea Elm. Leafi. Philip. Bot. 1 (1908) 322.

LUZON, Province of Tayabas, Lucban (Mount Banajao), Elmer 7438,
9214: Province of Laguna, Mount San Cristobal, Copeland, May, 1908:
Province of Zambales, Mount Tapulao, For. Bur. 8075 Curran & Merritt,
Bur. Sct. 4696, 5077 Ramos. MINporo, Mount Haleon, For. Bur. 4473
Merritt, June, 1906. PALAWAN, Mount Victoria, Bur. Sci. 646 Foxworthy,
March, 1906.

This species is widely distributed, extending from Japan and China to
the mountains of India, Ceylon, and Sumatra. The Philippine material,
while variable, agrees rather closely in all essential characters with speci-
mens from Japan, Ceylon, and China. The species described by Mr. Elmer
as Adinandra coriacea is a typical Ternstroemia in all respects.

GUTTIFERAE
CALOPHYLLUM Linnaeus
CALOPHYLLUM FERRUGINEUM sp. nov. § Inophyllum.
Arbor alta, ramis ramulisque crassis, partibus junioribus in-
florescentiisque dense ferrugineo-pubescentibus; foliis crassissi-

me coriaceis, oblongis, usque ad 10 em longis, in siccitate pallidis,
nitidis, basi acutis, apice obtusis ad obscure acuminatis, subtus

x, C,5 Merrill: Philippine Plants, XII 825

leviter ferrugineo-pubescentibus; inflorescentiis terminalibus,
paniculatis, circiter 15 cm longis, bracteolis oblongo-obovatis ad
lanceolatis, 1.5 ad 2 cm longis, deciduis; sepalis 4; petalis 6.

A tall tree with stout branches, the ultimate branchlets stout,
ferruginous-pubescent, 4 to 5 mm in diameter, reddish-brown.
Leaves thickly coriaceous, mostly oblong, about 10 cm long, 4 to
5 em wide, base acute, the apex obscurely acuminate, obtuse or
sometimes rounded, pale when dry, the upper surface shining, the
lower more brownish-yellow and, at least when young, sparingly
ferruginous-pubescent; nerves spreading, rather distinct, about
16 to a centimeter ; petioles dark brown, pubescent, about 1.5 cm
long. Inflorescence a terminal panicle, all parts very densely
ferruginous-pubescent, in nearly mature bud up to 15 cm long,
many-flowered, bipinnately paniculate, each flower subtended by
a very thick, densely pubescent, deciduous, oblong-oblanceolate
to oblong-obovate bracteole 1.5 to 2 cm in length, the pedicels
about 1 cm long. Sepals 4, densely pubescent, concave, the two
outer ones about 10 mm long and 8 mm wide, the two inner
ones orbicular, about 1 cm in diameter. Petals 6, in bud subor-
bicular, very slightly pubescent. Stamens very numerous.
Ovary glabrous.

Luzon, Province of Laguna, in forests back of San Antonio, Bur. Sci.
20545 Ramos, February, 1918.

For the genus a rather characteristic species. It is apparently allied
to Calophyllum blancoanum Pl. & Tr., from which it differs in its very
densely ferruginous-pubescent inflorescence, younger branches, and to a
less degree the lower surfaces of its leaves, in its prominent bracteoles

which are up to 2 cm in length, and in its thickly coriaceous, more coarsely
nerved, rather differently shaped leaves.

GARCINIA Linnaeus

GARCINIA NERVOSA Miq. Ann. Mus. Lugd. Bat. 1 (1863-64) 208; King
in Journ. As. Soc. Beng. 59” (1890) 169.
Stalagmites ? nervosa Miq. Fl. Ind. Bat. Suppl. (1861) 496.

PALAWAN, near Taytay, Merrill 9387, May, 1913, in bamboo thickets
at about sea level. Probably referable here is a single specimen from the
Province of Camarines, Luzon, For. Bur. 21750 Fischer, but this specimen
has leaves 65 cm long and 25 cm wide, the base broadly rounded and dis-
tinectly cordate.

The species is new to the Philippines. Perak, Singapore (Ridley 5966!),
Malacca, and Sumatra.

OCHROCARPUS Thouars
OCHROCARPUS RAMIFLORUS sp. nov.

Arbor parva, glabra; foliis oppositis, oblongis, subcoriaceis,
usque ad 25 cm longis, in siccitate pallidis, basi acutis ad rotun-

326 The Philippine Journal of Science 1915

datis, apice acutis vel breviter acuminatis, utrinque obscure
foveolatis, subtus plus minusve glandulosis, nervis primariis
utrinque 16 ad 20, tenuibus, quam secundariis reticulisque vix
magis distinctioribus; floribus in ramis fasciculatis, pedunculis
sub fructu 3 ad 4 cm longis.

A small tree about 5 m high, quite glabrous. Branches terete,
pale, the younger ones brownish-yellow or yellow, somewhat
compressed or angled, wrinkled when dry. Leaves opposite,
oblong, subcoriaceous, 15 to 25 cm long, 6 to 9 cm wide, base
rounded to acute, apex acute or shortly acuminate, margins re-
curved, both surfaces pale when dry, shallowly foveolate, the
foveole on the lower surface often glandular; primary lateral
nerves very slender, obscure, 16 to 20 on each side of the mid-
rib, scarcely more distinct than are the secondary nerves and
the reticulations; petioles wrinkled, 1.5 to 2 em long. Flowers
fascicled on the branches below the leaves, the subpersistent
petals oblong, about 5 mm long, the stamens very numerous, about
6mm long. Young fruits subglobose, about 1.5 cm in diameter,
much wrinkled when dry, brownish, apparently when mature 1-
celled and 1-seeded, when young at least 2-celled.

Luzon, Province of Camarines, Madadagat River, For. Bur. 22795
Tabat, August 5, 1913, on the banks of the river.

The first representative of the genus to be found in the Philippines,
Ochrocarpus pentapetalus F.-Vill., based on Tovomita pentapetala Blanco
being a Calophyllum and a synonym of C. amplexicaule Choisy. Ochro-
carpus ramiflorus appears to be most closely allied to O. siamensis T.

Anders. from which it differs in its acute or acuminate, fewer nerved
leaves and longer pedicels.

FLACOURTIACEAE
HOMALIUM Jacquin

HOMALIUM OBLONGIFOLIUM sp. nov. § Myriantheia.

Arbor circiter 35 m alta, inflorescentiis exceptis glabra; foliis
oblongis, usque ad 30 cm longis, coriaceis, nitidis, breviter
acuminatis, nervis utrinque circiter 15; paniculis amplis, folia
subaequantibus, axillaribus subterminalibusque, pubescentibus;
floribus 7-meris, sepalis quam petalis subduplo longioribus;
staminibus 21.

A tree 35 to 40 m high, glabrous except the inflorescence.
Branches terete, lenticellate, dark-colored when dry. Leaves
oblong, 20 to 30 cm long, 7 to 11 em wide, entire, coriaceous,
shining, the lower surface a little paler than the upper when
dry, entire, shortly acuminate, base acute; lateral nerves about

x, C, 5 Merrill: Philippine Plants, XII 327

15 on each side of the midrib, slender, distinct, anastomosing ;
petioles 1.5 to 2 cm long. Panicles axillary and subterminal,
ample, many-flowered, open, about as long as the leaves, branched
from the base, the lower branches sometimes 25 cm in length,
all parts rather densely gray-pubescent. Perianth-tube funnel-
shaped, 4 mm long, somewhat hirsute. Sepals 7, when fully
grown 6 to 7 mm long, about 1.8 mm wide, obtuse, pubescent,
oblong-oblanceolate. Petals 7, oblong, obtuse, pubescent, about
4 mm long. Stamens 21, in groups of three each opposite the
petals; filaments filiform, 2.5 to 3 mm long, very slightly pubes-
cent; anthers 0.8 mm long. Ovary densely hirsute, the styles 1
to 1.5 mm long.

MINDANAO, District of Zamboanga, Talisay, For. Bur. 18781 Foxworthy,
DeMesa, & Villamil (type), June 18, 1912, in forests, altitude about 30
meters; near Port Banga, For. Bur. 15216 Klemme, July, 1910, sterile
specimen.

A species most closely allied to Homalium luzoniense F.-Vill., but well
characterized by its oblong entire leaves.

The specimen collected by Klemme consists of leaves only, taken from
sprouts. One leaf is entire, 40 cm long and 14 cm wide, while the other,
a more juvenile form, is 40 cm long and 20 cm wide, the margins being
distinctly undulate-toothed.

SCOLOPIA Schreb.

Most of the recently collected Philippine material of the genus Scolopia
has been referred to Scolopia crenata Clos, and to S. luzonensis (Presl)
Warb. While Briquet’s paper on the genus has been known to me for
some years, it has only recently become available.’ Following Briquet in
his interpretation of the group of species closely allied to Scolopia crenata
Clos, in which he is doubtless correct, the known Philippine forms may
be disposed of as follows:

SCOLOPIA LUZONENSIS (Presl) Warb. in Engl. & Prantl Nat. Pflan-
zenfam. 3 (1893) 30, f. 11; Briq. in Ann. Conserv. Jard. Bot.
Genéve 2 (1908) 45; Merr. in Philip. Journ. Sci. 1 (1906) Suppl.
98.

Dasianthera luzonensis Pres] Rel. Haenk. 2 (1835) 90, t. 66.

Phoberos dasyanthera Benn. Pl. Jav. Rar. (1844) 192; F.-Vill. Novis.
App. (1880) 12.

Banara racemosa Blanco FI. Filip. (1837) 425.

Flacourtia corollata Blanco 1. c. ed. 2 (1845) 559.

Flacourtia crenata F.-Vill. Novis. App. (1880) 12; Vid. Rev. Pl. Vasc.
Filip. (1886) 48; Sinopsis Atlas (1883) 13, t. 7, f. B, non Clos.

Phoberos sp. Turcz. in Bull. Soc. Nat. Mose. 27 (1854) 333.

Scolopia rhinanthera F.-Vill. Novis. App. (1880) 12, non Clos.

* Briquet, J. Remarques sur les espéces asiatiques du genre Scolopia
Schreb., Ann. Conserv. Jard. Bot. Genéve 2 (1908) 41-47.

328 The Philippine Journal of Science 1915

This species is common and widely distributed in the Philippines, and
to it should be referred most of the Philippine material distributed from
the Bureau of Science as Scolopia crenata Clos and as S. luzonensis Warb.
It is well characterized by its ciliate anther-appendages.

I think it probable that Banara brevifolia Blanco FI. Filip. (1837)
426 = Flacourtia parvifolia Blanco FI. Filip. ed. 2 (1845) 560 should also be
referred here. Blanco’s description is altogether too short and incomplete
properly to determine the plant he had in mind, and of which he saw no
flowers. F.-Villar has reduced it to Scolopia dasyanthera Benn., which, if
correct, would place it under Scolopia luzonensis Warb., for Bennett’s
designation is only a new name for the plant originally described by Presl
as Desianthera luzonensis. This fact I overlooked at the time I worked
over the determinations of Blanco’s species, which accounts for my state-
ment® that Scolopia dasyanthera Benn. was a species-unknown from the
Philippines.

Just how constant are the characters selected by Briquet in distinguish-
ing the closely allied forms that have, for the most part, been reduced at
one time or another to form a comprehensive species, Scolopia crenata Clos,
remains to be seen. In our rich Philippine collections evidences of inter-
grading forms occur, that to a greater or less degree invalidate the key
characters adopted by Warburg and by Briquet in distinguishing the sec-
tions Adenoscolopia and Sphenoscolopia. Elmer 5625, 6363, from Benguet
Subprovince, Luzon, and Bur. Sci. 10376 McGregor, from Polillo, are in all
essential characters typical Scolopia luzonensis Warb., and have the peculiar
ciliate anthers of that species; yet all three specimens have glands at
the apices of their petioles, or on the leaf margins near the insertion of the
petioles which would place them in the section Adenoscolopia, and as a
result necessitate the description of the form as a new species. In my
mind there is absolutely no doubt but that all three sheets are fairly typical
Scolopia luzonensis Warb., a species manifestly belonging in the section
Sphenoscolopia.

SCOLOPIA SAEVA (Hance) Hance in Ann. Sci. Nat. IV 28 (1862) 217;
Briq. in Ann. Conserv. Jard. Bot. Genéve 2 (1898) 46.

Phoberos saevus Hance in Walp. Ann. 3 (1853) 825.
Scolopia lanceolata Clos, in Ann. Sci. Nat. IV 8 (1857) 252, p. p.,
quoad pl. Philip.; Vid. Rev. Pl. Vase. Filip. (1886) 49.

This species is very similar to Scolopia luzonensis Warb., but is dis-
tinguished by its entirely glabrous anthers. The following material is ap-
parently referable to it:

LuzON, Benguet Subprovince, For. Bur. 10921 Curran: Province of
Pangasinan, For. Bur. 9682 Zschokke: Province of Zambales, Merrill 2949,
For. Bur. 908 Maule: Province of Rizal, For. Bur. 428, 2680 Ahern’s
collector, Merrill 1658.

I have seen no specimen of Cuming 1061, cited by Clos, but the above
specimens agree with Hongkong material and with the descriptions avail-
able. Scolopia lanceolata Clos is based on Phoberos lanceolata W. & A.
Prodr. (1834) 380, and is an older name than that proposed by Hance and
accepted by Briquet. The type of Phoberos lanceolata W. & A. was from
India, and the species is, presumably, different from the Philippine form.

Govt. Lab. Publ. (Philip.) 27 (1905) 18.

xc, 5 Merrill: Philippine Plants, XII 329

FLACOURTIA Commerson

FLACOURTIA RUKAM Zoll. & Mor. Syst. Verz. (1854) 33; King, in Journ.
As. Soc. Beng. 59? (1890) 117; Vid. Phan. Cuming. Philip. (1885)
94; F.-Vill. Novis. App. (1880) 112.

Hisingera grandifolia Turez. in Bull. Soc. Nat. Mosc. 27* (1854) 332.
Flacourtia inermis Merr. in Philip. Journ. Sci. 1 (1906) Suppl. 99,
non Roxb.

This species is widely distributed in the Philippines, and is somewhat
variable in its vegetative characters. In general the leaves of the Philip-
pine form average considerably larger than in our rather comprehensive
set of Malayan forms, but there appears to be no specific difference. The
species is very similar in vegetative characters to Flacourtia inermis Roxb.,
with which the Philippine specimens have been confused. As indicated
in the original description, as later noted by King, and as shown in our
Malayan specimens, Flacourtia inermis has perfect flowers, while fF’. rukam
has dicecious flowers. I refer to F'lacourtia rukam the following Philippine
material:

LuZON, Province of Tayabas, Cuming 771 (cotype of Hisingera grandi-
folia Turez.): Benguet Subprovince, Elmer 8890: Province of Pangasinan,
For. Bur. 19455 Agama: Province of Zambales, For. Bur. 5901, 5999 Curran:
Province of Laguna, Bur. Sci. 6041 Robinson: Province of Rizal, Loher
6772: Province of Bataan, For. Bur. 1744 Borden, Whitford 1252. POLILLO,
Bur. Sci. 9125 Robinson, Bur. Sci. 10410 McGregor. MINDboRO, For. Bur.
5413 Merritt. CrBu, For. Bur. 15258 Cenabre. DINAGAT, Ahern 482. MIN-
DANAO, District of Davao, Copelend 534.

CASEARIA Jacquin
CASEARIA PHILIPPINENSIS sp. nov.

Frutex circiter 3 m altus, subtus foliis ramulis floribusque mol-
liter pubescentibus; foliis integris, oblongis ad oblongo-lanceo-
latis, usque ad 29 cm longis, in siccitate brunneis, acuminatis,
basi acutis, breviter petiolatis, nervis utrinque circiter 12; flo-
ribus paucis, fasciculatis, sepalis circiter 4.5 mm longis; stamino-
deis planis, sursum dense villosis.

A shrub about. 3 m high, the branchlets, flowers, and lower
surface of the leaves rather densely and softly pubescent;
branches rather slender, terete, brown. Leaves alternate, entire,
oblong to oblong-lanceolate, firmly chartaceous or subcoriaceous,
20 to 29 cm long, 7 to 10 em wide, brown when dry, the upper
surface glabrous, shining, the lower softly brown-pubescent, the
apex acuminate, base acute; lateral nerves about 12 on each
side of the midrib, somewhat curved-ascending, prominent; pe-
tioles stout, 5 mm long or less. Flowers yellowish, axillary, few,
subsolitary or somewhat fascicled, shortly pedicelled. Sepals
broadly ovate to elliptic-ovate, rounded, pubescent, 4 to 5 mm
long. Stamens 10, the free parts of the filaments glabrous, about
1 mm long, slender, the lower 1.5 mm, entirely united with the

330 The Philippine Journal of Science 1915

staminodes, glabrous; anthers about 1 mm long; staminodes
flattened, about 1.4 mm long, the upper part densely villous.
Ovary ovoid, about 3 mm long, with very few, scattered, short
hairs; style short; stigma capitate.

LUZON, Province of Cagayan, Claveria, Bur. Sci. 7364 Ramos (type),
March, 1909, in forests. SAMAR, Phil. Pl. 1644 Ramos, April, 1914, as
Rinorea.

A species well characterized by its comparatively large, entire leaves,
which are brown when dry, glabrous and shining on the upper surface and
softly pubescent on the lower surface; it does not appear to be closely allied
to any other Philippine species.

PASSIFLORACEAE
ADENIA Forskal

ADENIA LONGIFOLIA sp. nov.

Scandens, glabra; foliis oblongis, membranaceis, 20 ad 25 cm
longis, basi acutis vel leviter acuminatis, biglandulosis, vix auri-
culatis, apice breviter acute acuminatis, margine distanter
repando-denticulatis, nervis utrinque circiter 10; cymis peduncu-
latis, paucifloris, floribus, ¢ 4-meris, alabastro clavato, circiter
1 cm longo.

A scandent, herbaceous, glabrous vine, or the stems somewhat
woody, firm, striate and pale when dry, 3 mm in diameter.
Leaves oblong, membranaceous, 20 to 25 em long, 8 to 10 cm
wide, shining, somewhat olivaceous on the upper surface, the
lower surface a little paler, base acute or somewhat decurrent-
acuminate, 2-glandular, the glands prominent on the lower sur-
face, not auricled, apex shortly and sharply acuminate, margin
distantly repand-denticulate; lateral nerves about 10 on each side
of the midrib, distinct, anastomosing, the reticulations lax; pe-
tioles about 4 cm long. Peduncles solitary, axillary, up to 6
cm long, the cymes few-flowered. Staminate flowers greenish
or nearly white, the fully mature buds clavate, about 1 cm long,
the lobes 4, ovate, short. Stamens 4, narrowly oblong, about 3.5
mm long. Pistillate flowers and fruits not seen.

BASILAN, Bur. Sci. 15495 Reillo, August, 1912.

A species well characterized by its oblong, thin, repand-denticulate leaves,
which are acute or decurrent-acuminate at the base but not auriculate.

ADENIA PALMATIFOLIA sp. nov.

Scandens, glabra; foliis profunde palmato-5-lobatis, basi pro-
minente subauriculato-glandulosis, lobis lanceolatis, rectis vel
leviter falcatis, acuminatis, integris, usque ad 20 cm longis, 1.5
ad 2.5 cm latis; pedunculo elongato; fructibus .(immaturis)
circiter 6 cm longis, nitidis, ut videtur obovoideis.

6; 6 Merrill: Philippine Plants, XII 331

A scandent, glabrous, herbaceous vine, the branches pale,
striate, terete, 2 to 3 mm thick. Leaves chartaceous or sub-
coriaceous, pale when dry, slightly shining and of the same color
on both surfaces, the base very broadly truncate or abruptly
acute, subauriculate-glandular, 3- or obscurely 5-plinerved, the
lamina up to 23 cm long, subreniform in outline, very deeply
palmately 5-lobed, the lobes lanceolate, straight or somewhat
faleate, entire, sharply acuminate, 8 to 20 cm long, 1.5 to 2.5
cm wide, the sinuses obtuse or acute, each with a prominent
gland on the lower surface of the leaf, each lobe with a prom-
inent midrib, the nerves slender, distinct, spreading at right
angles, anastomosing; petioles 2.5 to 4 cm long. Peduncles up
to 15 cm long, the tendril part short. Flowers not seen. Im-
mature fruits apparently obovoid, about 6 cm long, shining,
yellowish-brown when dry, the pericarp thin, brittle.

Luzon, Subprovince of Benguet, Sablan, Elmer 6262, April, 1904 (type),
Bur. Sci. 12577 Fénix, December, 1910, in forests and thickets.

The specimens, or at least those of the last number cited, were distrib-
uted as Modecca parviflora Blanco, but that species is described as having
3-lobed leaves, while the present form has constantly 5-lobed leaves. It is
suspected that Modecca triloba Blanco and M. parviflora Blanco are the
same species, and moreover that they are only forms of Adenia coccinea
(Blanco) Merr. (Modecca coccinea Blanco). There are several specimens
in the herbarium of the Bureau of Science that present entire ovate leaves
on the same branches with prominently 3-lobed ones. Modecca saponaria
Blanco is a species of Momordica, and does not belong to the Passifloraceae.
Adenia palmatifolia is well characterized by its deeply palmately 5-lobed
leaves and its narrow, straight or falcate lobes.

ADENIA CRASSA sp. nov.

Scandens, glabra; foliis integris, ovatis, circiter 10 cm longis,
basi late rotundatis, cordatis, auriculatis, apice acute acuminatis,
chartaceis vel submembranaceis; fructibus ellipsoideis vel ovoi-
deis, circiter 8 cm longis, pericarpio crassissimo; seminibus inae-
quilateraliter obovatis, compressis, foveolatis, 8 mm longis.

A scandent, glabrous, apparently herbaceous vine, the stems
slender, pale, striate, terete. Leaves ovate, chartaceous or sub-
membranaceous, 8 to 10 cm long, 5 to 7 cm wide, entire, base
broadly rounded, prominently cordate, and biauriculate, the
apex rather slenderly and sharply acuminate, both surfaces some-
what shining, the lower a little paler then the upper; base 5-
nerved or somewhat 5-plinerved, the lateral nerves above the
base one or two pairs, distinct, reticulations lax; petioles 1.5
to 2.5 em long; peduncles axillary, solitary, slender, about 3 cm
long, the cymes apparently few-flowered, the tendril-parts short.
Flowers unknown. Fruit when fully mature about 8 cm long,

332 The Philippine Journal of Science 19165

ellipsoid or ovoid, smooth, pale, shining, splitting into 3 valves,
the pericarp very firm, about 2 mm thick; seeds numerous, com-
pressed, inequilateral, about 8 mm long, pale, prominently pitted,
base acute, apex somewhat oblique, their funicles up to 2 cm in
length.

BASILAN, Bur. Sci. 15419 Reillo, August, 1912.

In vegetative characters somewhat similar to our common Adenia coccinea
(Blanco) Merr., but in this species the pericarp of the mature fruit is
very thin, almost membranaceous.

THYMELAEACEAE
WIKSTROEMIA Endlicher

WIKSTROEMIA POLYANTHA sp. nov.

Frutex vel arbor parva, ramulis puberulis vel leviter pubes-
centibus; foliis brevissime petiolatis, ovatis ad oblongo-ovatis,
coriaceis, nitidis, usque ad 6 cm longis, acutis vel obscure
acuminatis, basi late rotundatis vel obscure cordatis, nervis
utrinque circiter 8, tenuibus; inflorescentiis racemosis, floribus
numerosis, brevissime pedicellatis, circiter 1 cm longis, extus
glabris vel parcissime pubescentibus.

A shrub or small tree, nearly glabrous, the branches terete,
reddish-brown, glabrous, the branchlets obscurely puberulent, the
younger ones with very short, appressed, cinereous hairs, some-
what angled or compressed. Leaves ovate to oblong-ovate,
glabrous, brownish when dry, shining on both surfaces, the lowen
surface a little paler than the upper, the apex acute or very ob-
scurely acuminate, the base broadly rounded, often obscurely
cordate; nerves slender, about 8 on each side of the midrib,
obscurely anastomosing; petioles reddish-brown, obscurely pu-
berulent, 2 mm long or less. Racemes terminal or in the up-
permost axils, appressed-hirsute, 2 to 3 cm long, many-flowered,
the entire lower part thickly covered with the persistent pedicels
of fallen flowers, the buds and open flowers crowded at the apices,
the pedicels sparingly appressed-hirsute, stout, less than 1 mm
long. Perianth cylindric, yellow, 10 to 11 mm long, about 1.5
mm in diameter, glabrous outside, or with very few scattered
hairs in the upper part, the two outer lobes orbicular-elliptic,
broadly rounded, 2.2 mm long, the two inner ones somewhat
shorter and narrower. Stamens 8, 4 inserted at about the mid-
dle of the tube, 4 inserted near the apex, included; anthers
narrowly oblong, 1.5 mm long. Ovary narrowly oblong, cylin-
dric, 2.5 mm long, villous at the apex, glabrous below; style
slender, about 1 mm long; stigma globose, 0.5 mm in diameter.

Rie, 5 Merrill: Philippine Plants, XII 833

Fruit narrowly ovoid, fleshy, red, about 8 mm long, sparingly
villous near the apex, otherwise entirely glabrous.

LuzON, Province of Rizal, near the summit of Mount Susung Dalaga,
Bur. Sci. 19248 Reillo, December 9, 1913.

A very characteristic species, quite distinct from the other Philippine
forms in its many-flowered, somewhat elongated racemes, the rachises of
which are rather densely covered with the short, persistent, stout pedicels.

RHIZOPHORACEAE
GY NOTROCHES Blume

GYNOTROCHES PUBERULA pp. nov..

Arbor parva, ramulis junioribus, subtus foliis stipulisque plus
minusve puberulis vel breviter pubescentibus; foliis oblongis, sub-
coriaceis, integris, acuminatis, basi acutis, usque ad 15 cm longis,
nervis utrinque circiter 7, curvato-adscendentibus ; fructibus axil-
laribus, fasciculatis, pedicellatis, ovoideis, circiter 4 mm longis.

A species very similar to Gynotroches axillaris Blume, differing
chiefly in its indumentum. Branches slender, terete, brownish-
red, somewhat compressed at the nodes, glabrous, the young
branchlets puberulent. Leaves subcoriaceous, oblong, 10 to 15
em long, 3 to 6 cm wide, pale or brownish when dry, shining, the
base acute, the apex rather prominently acuminate, the upper
surface quite glabrous, the lower distinctly puberulent or short-
pubescent especially on the midrib and lateral nerves; lateral
nerves about 7 on each side of the midrib, rather prominent,
curved-ascending; petioles about 8 mm long, puberulent; stipules
linear-lanceolate, 1.2 cm long, puberulent, deciduous. Fruits
numerous, axillary, fascicled, red when mature, ovoid, about
4 mm long, the pedicels jointed at about the middle, about as
long as the fruits, the persistent sepals ovate, acute, 1.5 mm long.

SaMAR, Catbalogan, Bur. Sci. 17396 Ramos (type); Cauayan Valley,
Phil. Pl. 1622 Ramos, April, 1914, in forests.

Very similar to and manifestly closely allied to Gynotroches axillaris

Blume, from which it differs chiefly in its puberulent indumentum on the
young branchlets, petioles, stipules, and lower surface of its leaves.

ARALIACEAE
BOERLAGIODENDRON Harms

BOERLAGIODENDRON DIVERSIFOLIUM sp. nov.

Frutex 2 ad 3 m altus, inflorescentiis exceptis glaber; foliis
coriaceis, valde diversis, majoribus usque ad 35 cm longis, usque
ad 4 vel 2 palmatim 3- vel 5-lobatis,.superioribus minoribus,
ellipticis ad oblongo-ovatis, 15 ad 30 cm longis, margine grosse

834 The Philippine Journal of Science 1915

irregulariter serratis; inflorescentiis terminalibus, plus minusve
ferrugineo-furfuraceo-villosis vel hirsutis; floribus numerosis, 5-
meris.

A shrub 2 to 3 m high, glabrous except the inflorescence, the
branches terete, gray, 1 to 1.5 cm in diameter. Leaves crowded
at the apices of the branches or of the simple (?) stem, coria-
ceous, exceedingly variable in form; larger leaves palmately 3-
or 5-lobed, up to 35 cm long, the lobes reaching to the middle or
lower two-thirds of the leaf, 5 to 11 cm wide, acuminate, coarsely
and irregularly serrate, the sinuses rounded; other leaves on the
same branchlets not at all lobed, elliptic to oblong-ovate, acumi-
nate, coarsely and irregularly serrate, 15 to 30 cm long, 7 to 17 cm
wide, the nerves prominent, more or less ascending; petioles 5
to 25 cm long, base with several crestlike ridges. Inflorescence
terminal, crowded, furfuraceous-villous or hirsute with few to
many short, brown hairs, the primary branches numerous, 2 to 3
cm long, each subtended by lanceolate, furfuraceous, and some-
what pectinate-ciliate bracts 2 to 2.5 cm long; central sterile head
1 to 2 cm in diameter, sessile or very shortly peduncled, the sterile
fruits globose, fleshy, glabrous, purple when fresh, black when
dry, 3 to 5 mm in diameter, the pedicels about as long as the
fruits; each primary branch with two lateral branches bearing
globose heads of perfect flowers, these lateral branches 2 to 3 cm
long, the heads globose, dense, about 1.5 cm in diameter.
Flowers many, densely disposed, 5-merous, yellowish, their ped-
icels stout, somewhat pubescent, 2 mm long. Calyx 2 mm long
and wide, glabrous, truncate. Corolla 3.5 to 4 mm long, with 5,
ovate, acute or obtuse, 1 to 1.2 mm long teeth; filaments 4 to 5
mm long; anthers about 1.7 mm long. Ovary 5-celled. Bracteoles
numerous, brown-hirsute, broadly ovate, obtuse, 2 to 2.5 mm long.
Entire inflorescence up to 15 cm in diameter.

MINDANAO, Bukidnon Subprovince, Sumilao, Bur. Sci. 15727 Fénix
(type), August 2, 1913, steep slopes near streams; Mount Dilirig, Bur.
Sci. 21447 Escritor, July 30, 1913: Butuan Subprovince, Weber 1099, July
14, 1911, at Talacogon, distributed as B. serratifolium Elm.

A species characterized by its exceedingly variable leaves, these small
to large, and entire to deeply 5-lobed. The variation somewhat parallels
that of Boerlagiodendron heterophyllum Merr., but the species are very
different from each other.

MYRSINACEAE
ARDISIA Swartz
ARDISIA ZAMBALENSIS nom. nov.

Ardisia biflora Merr. in Philip. Journ. Sci. 5 (1910) Bot. 212, non
King & Gamble, 1905.

X, C,5 Merrill: Philippine Plants, XII 335

The above new name is necessary for this species as the one previously
selected was preoccupied.

ARDISIA KEITHLEYI nom. nov.

Ardisia oblongifolia Merr. in Philip. Journ. Sci. 5 (1910) Bot. 219,
non King & Gamble, 1905.

Dedicated to Lieutenant Keithley in whose honor Camp Keithley was
named. The new name is necessary as the original specific name was
preoccupied.

EBENACEAE
DIOSPYROS Linnaeus

DIOSPYROS ALVAREZII sp. nov.

Arbor circiter 15 m alta, glabra vel subglabra, ramulis tenui-
bus, leviter puberulis vel subglabris; foliis chartaceis, oblongo-
ellipticis ad oblongis, usque ad 16 cm longis, utrinque subaequal-
iter angustatis, basi acutis, eglandulosis, apice acutis vel obscure
acuminatis, nitidis, pallidis, nervis utrinque 5 vel 6, tenuibus,
prominentibus, reticulis laxis; floribus axillaribus, ut videtur soli-
tariis vel binis, pedicellatis 4-meris; fructibus depresso-globosis,
junioribus 1.8 cm diametro, pubescentibus, 8-locellatis, calycibus
persistentibus, coriaceis, tubo circiter 1.5 cm diametro, lobis
late ovatis, 1.5 em longis, reflexis.

A tree about 15 m high, nearly glabrous except the fruits
(flowers unknown). Branches terete, very slender, the branch-
lets sometimes minutely puberulent. Leaves chartaceous, pale
when dry, oblong-elliptic to oblong, 10 to 16 cm long, 4 to 7 cm
wide, shining, subequally narrowed to the acute, eglandular base
and to the acute or somewhat acuminate apex; lateral nerves 5
or 6 on each side of the midrib, distinct, slender, rather prom-
inent, anastomosing, the reticulations lax; petioles 6 to 8 mm
long. Flowers 4-merous, axillary, apparently solitary or in pairs,
the pedicels, in fruit, minutely puberulent, 1.5 cm long or less.
Young fruits depressed-globose, about 1.8 cm in diameter,
grayish-pubescent with short, appressed hairs, apiculate; per-
sistent calyx large and prominent, thickly chartaceous, externally
more or less puberulent, the tube very shallow, up to 1.5 cm in
diameter, 4-angled, the angles rather sharp, the lobes sharply
reflexed, broadly ovate, obtuse, up to 1.5 cm long, more or less
plicate when dry; fruit 8-celled.

LuzON, Province of Camarines, Lanot, For. Bur. 21417 Alvarez, May
12, 1914, forested slopes at about sea level, locally known as bantolinao.

A species well characterized by its chartaceous, distantly but rather
prominently nerved leaves, and especially by its prominent, accrescent,

persistent, reflexed calyx which in young fruit is sharply 4-angled.
133910-—4

336 The Philippine Journal of Science 1915

DIOSPYROS PONCEI sp. nov.

Arbor circiter 9 m alta, ramulis junioribus dense adpresse
subferrugineo-hirsutis, foliis junioribus subtus obscure adpresse
pubescentibus, fructibus subglobosis, extus densissime ferrugi-
neo-hirsutis, junioribus circiter 2 cm diametro; foliis oblongis ad
oblongo-lanceolatis, coriaceis, usque ad 9 cm longis, in siccitate
pallidis, nitidis, utrinque subaequaliter angustatis, basi acutis,
apice acutis vel breviter acuminatis, nervis lateralibus obscuris;
sepalis persistentibus, orbiculari-ovatis, circiter 6 mm diametro,
adpresse hirsutis.

A tree about 9 m high, the branches terete, glabrous, wrinkled,
dark gray, the younger ones reddish-brown, smooth, the growing
parts densely appressed ferruginous-hirsute. Leaves coriaceous,
oblong to oblong-lanceolate, 6 to 9 cm long, 1.4 to 2.8 em wide,
subequally narrowed to the acute base and to the acute or
somewhat acuminate apex, eglandular, the upper surface pale
olivaceous when dry, shining, glabrous, or the midrib in the
lower part somewhat appressed-pubescent with pale, shining
hairs, the lower surface paler than the upper, when young spar-
ingly appressed-pubescent with pale hairs, when mature quite
glabrous; lateral nerves slender, obscure, ascending, the primary
ones about 12 on each side of the midrib, scarcely more distinct
than are the secondary ones and the reticulations; petioles 5 mm
long or less, appressed-hirsute when young, becoming glabrous
in age. Flowers not seen. Fruits globose, the young ones about
2 cm in diameter, axillary, sessile or subsessile, solitary, the
pericarp very densely hirsute with ferruginous or fulvous hairs.
Sepals persistent, plane, coriaceous, orbicular-ovate, about 6 mm
in diameter, rounded, outside rather densely appressed-hirsute
with pale or ferruginous hairs. :

MINDANAO, Province of Surigao, Combot, For. Bur. 22842 Ponce, August
28, 1914, on semiopen slopes, altitude about 15 meters, locally known as
ituman.

A species in many characters resembling the common Diospyros discolor
Willd., and probably belonging in the same section as Willdenow’s species.

It differs remarkably, however, in its very much smaller, entirely differently
shaped leaves which are quite glabrous on both surfaces when mature.

DIOSPYROS CAMARINENSIS sp. nov.

Arbor circiter 25 m alta, floribus exceptis glabra; foliis char-
taceis, oblongis, usque ad 27 cm longis, breviter acuminatis,
basi acutis vel subacutis, in siccitate nitidis, nigricantibus, nervis
utrinque 12 ad 14, subtus distinctis, reticulis laxis; floribus axil-
laribus, fasciculatis, 4-meris, extus pubescentibus, calycis lobis
erectis, acuminatis, corolla inflata; ovario 8-loculare; fructibus

RC, 5 Merrill: Philippine Plants, XII 337

globosis vel depresso-globosis, circiter 3 cm diametro, in siccitate
nigris, nitidis, glabris, Sseminibus 7 vel 8.

A tree about 25 m in height, quite glabrous except the flowers,
the branches slender, terete, dark-colored, the branchlets and
petioles black when dry. Leaves chartaceous, dark-colored or
somewhat blackish when dry, shining, 14 to 27 cm long, 5 to 9
cm wide, of about the same color on both surfaces, the apex
shortly acuminate, base acute to subacute, usually with a pair
of distinct glands near the junction with the petiole; lateral
nerves 12 to 14 on each side of the midrib, slender, distinct, anas-
tomosing, the reticulations rather lax, distinct; petioles 1 to 1.5
em long. Flowers axillary, fascicled, 4-merous, few in each fasci-
cle, some in the axils of leaves, some in the axils of fallen leaves,
each subtended by one or two pairs of broadly ovate, pubescent
bracts 2 mm long or less. Calyx about 8 mm long and 7 mm in
diameter, pubescent externally, black when dry, the lobes broadly
ovate, acuminate, erect, about 4 mm long, 4 to 4.5 mm wide at
the base. Corolla-tube 8 to 9 mm long, inflated, about 7 mm in
diameter, much narrowed toward the throat, rather densely
pubescent with short, grayish-brown hairs, the lobes 4, spread-
ing, narrowly oblong, somewhat falcate, acute, thickly coria-
ceous, about 9 mm long, 5 mm wide. Stamens about 20, mostly
in pairs, 2-seriate, the filaments of each pair more or less united;
anthers narrowly lanceolate, acuminate, 3.5 to 5.5 mm long.
Ovary ovoid, pubescent, about 4 mm in diameter, 8-celled; style
stout, pubescent, widened below, about as long as the ovary, the
stigma irregularly lacerate. Fruit globose or depressed-globose,
yellow to red when mature, black and shining when dry, about
3 cm in diameter, the persistent calyx black, coriaceous, glabrous
or nearly so, nearly square in outline, about 1.5 cm broad, shallow.
Seeds 7 or 9, somewhat flattened, about 13 cm long, 7 mm wide,
rounded at both ends, smooth, black, shining, the albumen smooth.

LuzoN, Province of Camarines, For. Bur. 21448 (type) 21705, 22631
Alvarez, April and May, 1914, March, 1912, from Mount Calinigan, Mount
Labo, and Lanot River, in forests, 100 to 500 meters.

The species is perhaps as closely allied to Diospyros maritima Blume
as any other species, but with entirely different, much thinner, fewer nerves
and different flowers and fruits.

MIMUSOPS Linnaeus

MIMUSOPS CALOPHYLLOIDES sp. nov.

Arbor circiter 18 m alta, floribus exceptis glaberrima; foliis
oblongo-obovatis, subcoriaceis, usque ad 12 cm longis, obtusis ad
rotundatis, basi acutis, nitidis, nervis lateralibus numerosissimis,
densis, distinctis, margine recurvatis, venis marginalibus dis-

338 The Philippine Journal of Science 1915

tinctis; floribus 6-meris, sepalis puberulis circiter 5 mm longis,
exterioribus quam interioribus bis latioribus; staminibus 6;
ovario glabro, 6-loculare; fructibus globosis, 2 ad 2.5 cm diametro,
breviter apiculatis, glabris, pericarpio fragile.

A tree about 18 m high, entirely glabrous except the flowers.
Branches dark-colored when dry, the younger ones marked with
numerous, crowded, petiolar scars. Leaves subcoriaceous,
oblong-obovate, 9 to 12 cm long, 2.5 to 5 cm wide, entirely gla-
brous on both surfaces, shining, subolivaceous when dry, the
base acute, the apex obtuse to rounded, margins distinctly re-
curved; lateral nerves very numerous, slender but rather dis-
tinct, the primary ones not more prominent than the secondary
ones and the reticulations, crowded, about 20 to a centimeter,
uniting to form a distinct submarginal nerve about 1 mm from
the edge of the leaf; petioles 2 to 2.5 cm long. Flowers axillary,
6-merous, their pedicels puberulent, in fruit thickened upward
and about 2.5 cm long. Sepals 6, 2-seriate, puberulent, the outer
three about 5 mm long, triangular-oblong, acute, the inner three
as long as the outer ones but one-half as wide. Stamens 6.
Ovary glabrous, 6-celled. Fruit globose, 2 to 2.5 cm in diameter,
brown when dry, tipped by the very short style, glabrous, the
pericarp brittle. Seeds 1 or 2 in each fruit, brown and shining,
obtuse, about 1.5 cm long and 1 cm wide, slightly compressed.

MINDANAO, Province of Surigao, Cadyangao, For. Bur. 22834 Ponce,
August 20, 1914, in rather dense forests at about sea level, locally known
as duyuk-duyuk.

The alliance of this species is with the Malayan Mimusops kauki Linn.
from which it differs in its leaves being entirely glabrous, even when very
young, not at all pubescent on the lower surface, its much more distinct

lateral nerves, giving the leaves quite the appearance of those of some
species of Calophyllum, and the distinct marginal veins.

OLEACEAE
LINOCIERA Swartz

LINOCIERA OBOVATA sp. nov.

Arbor parva, ramulis junioribus inflorescentiisque minute
puberulis exceptis glabra; foliis crassissime coriaceis, obovatis,
4 ad 7 em longis, apice late rotundatis vel late breviter acumi-
natis, basi angustatis, decurrento-acuminatis, nervis lateralibus
obscuris, circiter 8 utrinque. Cymis axillaribus, solitariis, an-
guste pyramidatis, dense multifloris, sessilibus.

A small tree, glabrous except the minutely puberulent branch-
lets and the inflorescence. Branches gray, stout, terete, gla-
brous. Leaves very thickly coriaceous, obovate, 4 to 7 cm long,
2 to 4.5 cm wide, pale when dry, the upper surface shining, the

%, 6, 5 Merrill: Philippine Plants, XII 339

lower dull, apex broadly rounded, sometimes very shortly and
broadly acuminate, sometimes even slightly retuse, the base nar-
rowed and more or less decurrent on the petioles, the margins
recurved; lateral nerves indistinct, often subobsolete, about 8 on
each side of the midrib; petioles 1 to 1.5 cm long. Cymes pan-
icled, axillary, sessile, solitary, densely many-flowered, branched
from the base, slightly puberulent, brown when dry, 2 to 4.5
cm long, 2 to 3 cm wide, the branches spreading. Flowers 4-
merous, the buds ellipsoid or obovoid, about 2 mm long, their
pedicels 1 to 3 mm long. Calyx-teeth broadly triangular, acute,
short. Mature flowers and fruits not seen.

LUZON, Province of Laguna, San Antonio, in forests, Bur. Sci. 15014
Ramos (type), Phil. Pl. 1161 Ramos, June, 1912.

A very characteristic species, recognizable by its very thick, obovate,
obscurely nerved leaves, and by its dense pyramidal inflorescence.

LINOCIERA NITIDA sp. nov.

Ut videtur arbor parva, glabra; foliis oblongis, in siccitate
nitidis, subcoriaceis, oblongis vel late oblongis, usque ad 18 cm
longis, basi rotundatis vel subacutis, apice brevissime acumina-
tis, nervis utrinque circiter 8, patulis, prominentibus, anastomo-
santibus, reticulis obsoletis vel subobsoletis; racemis brevibus,
1 ad 1.5 cm longis, solitariis vel fasciculatis, axillaribus, densi-
floris; floribus 4-meris, 5 mm longis.

Apparently a small tree, glabrous, the branches terete, smooth,
light gray. Leaves oblong or broadly oblong, subcoriaceous, 12
to 18 cm long, 5 to 7 cm wide, entire, brown when dry, or the
upper surface somewhat olivaceous and the lower surface brown,
shining on both surfaces, smooth, the base broadly rounded or
somewhat acute, the apex very shortly and somewhat abruptly
acuminate; lateral nerves 7 or 8 on each side of the midrib,
prominent, spreading at nearly right angles, somewhat curved,
anastomosing, the reticulations obsolete or nearly so; petioles
dark brown, 2 to 3 mm long. Racemes short, axillary or in
the axils of fallen leaves, solitary or somewhat fascicled, 1 to
1.5 cm long, densely flowered, quite glabrous or very obscurely
and sparsely puberulent, the bracts broadly ovate, acute or acu-
minate, 1.5 mm long. Sepals ovate, acute, about 1 mm long.
Petals 4, narrowly oblong, thick, 5 mm long, about 1 mm wide,
slightly narrowed upward, the apex obtuse, somewhat cucullate,
the margins somewhat inflexed. Anthers about 1 mm long, the
connective very broad.

BASILAN, Bur. Sci. 15406 Reillo, August 11, 1912, in forests.

A species well characterized by its vegetative characters, especially by
its leaves being brown when dry and with obsolete or nearly obsolete

340 The Philippine Journal of Science 1915

reticulations, and its short, many-flowered, mostly fascicled, racemose
inflorescence.

LINOCIERA PALUDOSA King & Gamble in Journ. As. Soc. Beng. 74?
(1905) 268.

PALAWAN, Taytay, Merrill 9177, 9335, Phil. Pl. 1373 (as L. euphlebia
Merr.), April, May, 1913.

The specimens are not quite identical with King’s collector 6476 from
Larut, Malay Peninsula, a duplicate of one of the specimens cited in the
original description. I can, however, detect no essential differences and
prefer to consider the Philippine form under King & Gamble’s specific name.

Previously known only from the Malay Peninsula.

BORAGINACEAE
HELIOTROPIUM Linnaeus

HELIOTROPIUM OVALIFOLIUM Forsk. var. DEPRESSUM (Cham.)
Merr. in Philip. Journ. Sci. 9 (1914) Bot. 134.
Heliotropium gracile R. Br. var. depressum Cham. in Linnaea (1829)
457.
Heliotropium coromandelinum Retz. var. depressum A. DC. Prodr. 9
(1845) 542.

LuzON, Province of Laguna, Calamba, along the shores of Lake Bay,
F. C. Gates 6651, July 27, 1913.

This species, other than the record of F.-Villar, sub H. coromandelinum
Retz., is new to the Philippines. The specimen cited above agrees in all
essentials with our Guam material, the type of the variety depressum being
from Guam. It differs from the Indian material notably in its relatively
longer and much narrower leaves. Forskal’s specific name dates from the
year 1775, Retzius’s name from the year 1781.

India, Tropical Africa, and Australia, the variety in Luzon and the
Mariana Islands (Guam).

GESNERIACEAE
TRICHOSPORUM D. Don

TRICHOSPORUM BAKERI sp. nov. § Holocalyx.

Scandens, ramulis junioribus plus minusve pilosus; foliis coria-
ceis, in siccitate pallidis, ovatis ad elliptico-ovatis, usque ad 4.5
cm longis, obtusis vel late obscure obtuseque acuminatis, basi
rotundatis, nervis lateralibus circiter 3, obscurissimis vel obso-
letis; floribus glaberrimis, 5.5 em longis, corolla quam calycis
duplo longioribus.

Scandent, slender, elongated, the stems terete, grayish,
wrinkled, 3 mm in diameter or less, rooting at the nodes, spar-
ingly branched, the younger branchlets rather densely villous.
Leaves numerous, when fresh apparently fleshy, when dry co-
riaceous or subcoriaceous, pale, dull or slightly shining, the

MHC, 6 Merrill: Philippine Piants, XII 341

nerves and reticulations obsolete, or sometimes about 3 pairs
of lateral nerves evident but obscure, 2 to 4.5 em long, 1.5 to
3.8 cm wide, base rounded to obscurely cordate, apex obtuse to
broadly and obscurely blunt-acuminate, the younger ones often
slightly villous on the margins; petioles 2 to 4 mm long, when
young more or less villous, when old quite glabrous. Flowers
apparently dark red, on short, 1- or 2-flowered peduncles in the
uppermost axils. Calyx cylindric, quite glabrous, about 2.5 cm
long, very shallowly lobed, slightly enlarged upward. Corolla
about 5.5 cm long, entirely glabrous, somewhat curved, contract-
ed above the base, then expanded, the lobes broad, subequal;
stamens slightly exserted; style pubescent in the lower one-half.

LUZON, Province of Laguna, hills back of Paete, C. F. Baker 3714 (type),
3719, December 28, 1914; Mount Maquiling, Bur. Sci. 16889 Servinas,
November, 1912, trailing over bowlders or ledgers.

A species similar to and manifestly closely allied to TRICHOSPORUM
LOHERI (Kranzl.) [Aeschynanthus loheri Kranzl. in Philip. Journ. Sci.
8 (1918) Bot. 165], from which it differs in its much slenderer, entirely
glabrous flowers. Kranzlin describes the flowers of Trichosporum loheri
as glabrous, but both the calyx and corolla are distinctly but minutely
pubescent externally.

ACANTHACEAE
ASYSTASIA Blume

ASYSTASIA GANGETICA (Linn.) T. And. in Thwaites Enum. Pl. Zeyl.
(1859-64) 235.

Justicia gangetica Linn. Amoen. Acad. 4 (1759) 299.
Asystasia coromandelica Nees in Wall. Pl. As. Rar. 3 (1832) 89,
F.-Vill. Nov. App. (1880) 154.

LuZzON, Province of Isabela, Ilagan, Bur. Sci. 7981 Ramos: Province of
Laguna, Los Banos, Merrill 8613, For. Bur. 20858 Villamil, Bur. Sci. 17282
Robinson & Foxworthy, all from living plants secured in Cagayan Province
by Mr. Curran.

The above is the first verification of F.-Villar’s record for this genus and
species as a Philippine plant. Widely distributed in tropical Asia, Africa,
and Malaya.

POLYTREMA C. B. Clarke

POLYTREMA ADDISONIENSE (Elmer) comb. nov.
Hypoestes addisoniensis Elmer Leafl. Philip. Bot. 5 (1913) 1697.

The type number of this is Elmer 12715, from Addison Peak, Palawan;
the same species is apparently represented by Merrill 7235 from Ulugan
Bay, Palawan, September, 1910. It is not a Hypoestes but is congeneric with
the three species of the Malay Peninsula for which C. B. Clarke has pro-
posed the generic name Polytrema. The Palawan species is apparently
closely allied to Polytrema vulgare C. B. Clarke. The genus appears to
be very closely allied to Hallieracantha Stapf. Here I also refer the
following species:

342 The Philippine Journal of Science 1915

POLYTREMA PULGARENSE (Elmer) comb. nov.
Hypoestes pulgarensis Elmer Leafl. Philip. Bot. 5 (1913) 1698.

The type is Elmer 12785, from Mount Pulgar, Palawan, and is not
matched by any other collection so far received.

POLYTREMA AEQUIFOLIUM C. B. Clarke sp. nov.

Procumbens, radicans, parum pilosa, foliis oppositis, aequa-
libus, lamina 6 ad 7 cm longa, elliptica aut lanceolata, petiolo 5
ad 10 mm longo; corolla 11 ad 12 mm longa; capsula 15 mm
longa, glabra; seminibus 4, minute verrucosis.

Resembling a weak axillary-flowered Justicia, the corolla and
capsule much as in that genus. Stamens 2; anther-cells paral-
lel, oblong, at equal height, muticous. Pollen globose, minutely
granular, without bands, with 4 (or more) stopples. Cymes
5- to 1-flowered, small, axillary. Calyx-segments 5, linear, 6 mm
long; bract lanceolate, shorter than the calyx.

PALAWAN, Puerto Princesa, Bur. Sci. 857 Bermejos, January, 1906.

The above description was supplied to me by the late C. B. Clarke some

time before his death, but was not printed then as the description of the
genus had not been published at that time.

LEPIDAGATHIS Willdenow

LEPIDAGATHIS PALAWANENSIS sp. nov.

Herba erecta usque ad 60 cm alta, haud vel parce ramosa,
subtus foliis ad nervis ramulisque prominente subfurfuraceo-
pubescentibus, infiorescentiis dense albido-ciliato-pilosis; foliis
chartaceis, oblongo-ovatis, usque ad 12 cm longis, integris vel
obscure undulatis, obtusis vel subacutis, base longe decurrento-
acuminatis, nervis utrinque 8 vel 9, subtus prominentibus; spi-
cis axillaribus terminalibusque, fasciculatis, densis, usque ad 5
cm longis, calycis lobis bracteisque dense ciliato-pilosis; capsulis
5 mm longis, 4-angulatis.

An erect, unbranched or sparingly branched herb up to 60 cm
in height, the stems terete, about 4 mm in diameter, the younger
parts obscurely 4-angled, rather prominently subfurfuraceous-
pubescent as are the nerves on the lower surface of the leaves
and petioles. Leaves in equal pairs, oblong-ovate, chartaceous,
9 to 12 cm long, 3.5 to 6 cm wide, the upper surface olivaceous,
shining, glabrous, with scattered, short cystoliths, the lower sur-
face a little paler, apex obtuse or subacute, base long-decurrent-
acuminate, winging the petiole nearly to its base, the petioles 3
to 4 cm long; lateral nerves 8 or 9 on each side of the midrib,
prominent on the lower surface. Spikes axillary and terminal,

x, C5 Merrill: Philippine Plants, XII 3438

numerous, fascicled, up to 5 cm in length, dense, many-flowered,
ciliate-pilose with white hairs. Bracts oblong-lanceolate, acumi-
nate, about 4.5 mm long, 1.5 mm wide, imbricate, prominently
ciliate-pilose with white hairs as are the sepals. Sepals 5, four
narrowly lanceolate, acuminate, 4 mm long, 0.8 mm wide, the
other as long but twice as wide. Corolla at least 5 mm long,
the lobes subequal, rounded. Capsule oblong-lanceolate, nar-
rowed upward, 4-angled, about 5 mm long, slightly puberulent
toward the apex; seeds 4.1 mm in diameter, the margins rather
prominently ciliate.

PALAWAN, Taytay, Merrill 9760, April, 1913, in dry thickets along trails
at sea level.

Perhaps as closely allied to Lepidagathis mindorensis Merr. as any other
species, but the indumentum not at all capitate-glandular and the leaves
entirely different in shape, not rounded or cordate at the base and with
short petioles, but long decurrent-acuminate and long-petioled.

LEPIDAGATHIS MINDORENSIS sp. nov.

Herba erecta circiter 20 cm alta; foliis oblongo-lanceolatis,
subcoriaceis, glabris, usque ad 9 cm longis, basi late rotundatis
vel subcordatis, supra sensim angustatis, obtusis vel obscure ob-
tuse acuminatis, breviter petiolatis; spicis terminalibus dense
fasciculatis, 2 ad 4 cm longis, angustis, densis, calycis segmentis
bracteisque capitato-glandulosis, plus minusve dense ciliato-pilo-
Sis; capsulis dense cinereo-pubescentibus.

An erect herb about 20 cm high, somewhat branched, the basal
parts of the stems somewhat woody, terete, glabrous, grayish-
brown, the younger parts distinctly 4-angled, brown, more or
less densely covered with weak, crisp, brownish hairs. Leaves
in equal pairs, oblong-lanceolate, subcoriaceous, 5 to 9 cm long,
2 to 3 cm wide, widest in the lower part, gradually narrowed
upward to the blunt, or slightly blunt-acuminate apex, the base
broad, rounded or subcordate, margins slightly undulate, the
upper surface glabrous and shining, the lower when young some-
what puberulent on the midrib and nerves; nerves slender, dis-
tinct beneath, anastomosing and forming a submarginal nerve;
petioles pubescent, 2 to 4mm long. Spikes terminal, fasciculate,
many at the apex of each branch, crowded, 2 to 5 cm long, less
than 5 mm in diameter, the bracts and calyx-segments rather
densely ciliate-pilose, with numerous glandular-capitate hairs
intermixed. Bracts narrowly lanceolate, slenderly long-acumi-
nate, about 4 mm long, less than 1 mm wide, 1-nerved. Calyx-seg-
ments all 1-nerved, slenderly acuminate, narrowly lanceolate, one

344 The Philippine Journal of Science 1915

about 1.2 mm wide, two slightly less than 1 mm wide, the fourth
cleft to within about 1 mm of the base. Corolla pink, pubescent
outside, 5 mm long, the tube slightly narrowed upward, not
abruptly. contracted, the upper lip about 1.3 mm wide, rounded
or very obscurely retuse, the lower one cleft into three obtuse
lobes about 1.2 mm long and 1mm wide. Anthers 0.5 mm long,
the cells parallel. Capsule narrowly oblong-ovate, densely cine-
reous-pubescent with short hairs.

MINDORO, Bulalacao, in rocky soils along streams, altitude about 50
meters, Bur. Sci. 6702 Robinson, March 17, 1909.

A species well characterized by its firm, short-petioled leaves which are
rounded or subcordate at the base, and by its narrow, dense, fasciculate
spikes, the bracts, bracteoles, and calyx-segments being rather densely
ciliate-pilose and capitate-glandular, and in its capsules being densely
cinereous-pubescent.

LEPIDAGATHIS CLARKEI sp. nov.

Species L. incurvae Don affinis differt spicis oblongis, solitariis
vel subsolitariis, bracteis majoribus, margine parce ciliatis ex-
ceptis glabris, in siccitate brunneis vel brunneo-purpureis.

An erect herbaceous plant 20 to 30 cm high, not or but slightly
branched, the basal part rarely decumbent. Stems brown or
olivaceous, glabrous, slender, terete or slightly angled. Leaves
in equal pairs, ovate to oblong-ovate, membranaceous, greenish
when dry, slightly shining and somewhat paler on the lower sur-
face than on the upper, 5 to 10 cm long, 2.5 to 4.5 em wide, the
base usually long and slenderly decurrent-acuminate, the apex
acute or obscurely acuminate, margins entire, both surfaces with
scattered, short cystoliths, glabrous; lateral nerves about 6 on
each side of the midrib, slender; petioles 1 to 2.5 emlong. Spikes
solitary, terminating the stem and the short, axillary branches,
the whole inflorescence appearing compound, the individual
spikes oblong, 2 to 5 cm long, about 1.5 ecm in diameter, the
bracts lanceolate, long and slenderly acuminate, about 11 mm
long, 2 mm wide, brown or brown-purple when dry, 3-nerved,
with distinct transverse reticulations in the upper part, glabrous
except the slightly ciliate margins. Calyx-lobes all acuminate,
one lanceolate, 3-nerved, with transverse reticulations in the up-
per half, about 10 mm long, 2.1 mm wide, two linear, 9 mm long,
the fourth one cleft to within 4 mm of the base, the lobes about
2 mm wide, with a prominent midnerve and each with a pair of
marginal nerves, all the segments slightly ciliate on the margins,
otherwise glabrous. Corolla 8 mm long, white, glabrous, the tube
slightly constricted, the lobes four, subequal, oblong, obtuse,

XC, 5 Merrill: Philippine Plants, XII 345

about 2.5 mm long, 1 to 1.2 mm wide. Capsule oblong, 6 mm
long, 2 mm wide, glabrous.

LuzON, Province of Bataan, Lamao River, Williams 37, October, 1903
(type), Merrill 3119, October, 19038, Whitford 493, July, 1904, on. damp
shaded banks along the stream at an altitude of 150 meters or less.

One of the specimens (Whitford 493) was identified by the late C. B.
Clarke as Lepidagathis incurva Don (L. hyalina Nees), but it seems to me
to be specifically distinct from that species. The brown or brown-purplish
color of the spikes is characteristic. It is the form previously reported
by me as Lepidagathis incurva Don.”

LEPIDAGATHIS SUBINTERRUPTA sp. nov.

Herba parva, prostrata, ramis floriferis suberectis, vix 10 cm
altis, inflorescentiis exceptis glabra; foliis parvis, oblongis, usque
ad 2.5 cm longis, breviter petiolatis, obtusis vel obscure acumi-
natis; spicis terminalibus, gracilibus, plus minusve interrup-
tis, solitariis vel subfasciculatis, terminalibus, usque ad 5 cm
longis; calycis segmentis bracteisque leviter ciliatis capitato-
glandulosisque.

A small prostrate herb, glabrous except the inflorescence, the
stems 4-angled, slender, rooting at the nodes, the flowering
branches erect or suberect, 10 cm high or less. Leaves in equal
pairs, mostly oblong, chartaceous, 1 to 2.5 cm long, 4 to 8 mm
wide, glabrous, somewhat shining, paler on the lower surface
than on the upper, both surfaces with small scattered cystoliths,
the base acute or somewhat decurrent-acuminate, the apex obtuse
or obscurely acuminate; petioles 1 to 3 mm long. Spikes term-
inal, solitary or at most three together, slender, interrupted, 1 to
5 em long, about 5 mm in diameter, the flowers somewhat scat-
tered. Bracts and bracteoles similar, lanceolate, acuminate, 3
to 4 mm long, less than 1 mm wide, 1-nerved, slightly ciliate and
capitate-glandular. Calyx-segments 5, 3 to 4 mm long, ob-
scurely ciliate and slightly capitate-glandular, one about 1.2 mm
wide, the other four similar but less than 1 mm wide. Corolla
white, 4 mm long, the tube not contracted, the upper lip 1.5 mm
long, 1.1 mm wide, rounded or very obscurely retuse, the
lower one 3-lobed, the lobes about 1 mm long, 0.8 mm wide,
obtuse. Capsule glabrous, 4 mm long, the seeds brown, about
1 mm in diameter.

LuzoN, Province of Isabela, San Luis, Bur. Sci. 8024 Ramos, May, 1909,

on powlders along shaded streams.
A species probably most closely allied to Lepidagathis laxa Nees, differing
especially in its much smaller leaves.

* Philip. Journ. Sci. 1 (1906) Suppl. 125.

346 The Philippine Journal of Science 1915

LEPIDAGATHIS MACGREGORII sp. nov.

Species L. laxae Nees ut videtur affinis, differt foliis oblongo-
ovatis, multo majoribus, usque ad 12 cm longis, 5 cm latis, floribus
4-meris, spicis longioribus, calycis segmentis bracteisque leviter
ciliatis, vix glandulosis.

Erect, nearly glabrous, the branches quadrangular, slender,
often somewhat puberulent at the nodes, reddish-brown. Leaves
in equal pairs, membranaceous, glabrous, oblong-ovate, 10 to 12
cm long, 4.5 to 5 em wide, somewhat shining when dry, slightly
paler on the lower surface than on the upper, the base rather
abruptly and shortly decurrent-acuminate, the apex acute or
slightly acuminate, margins somewhat undulate, both surfaces
with small, scattered cystoliths, the lower one sometimes slightly
puberulent along the midrib; lateral nerves 7 or 8 on each side
of the midrib, curved, anastomosing; petioles 1 cm long or less.
Spikes fasciculate, from one to three on each peduncle, the
peduncles very short or up to 4 cm in length, crowded in the
upper axils, the spikes slender, 2 to 6 cm long, 5 to 6 mm in
diameter. Bracts and bracteoles similar, lanceolate, acuminate,
1-nerved, 4 to 5 mm long, 1.2 mm wide, margins slightly ciliate
with short hairs, otherwise glabrous, not at all glandular. Calyx-
segments all acuminate, slightly ciliate on the margins, not gland-
ular, one lanceolate, 5 mm long, 1.8 mm wide, very obscurely
3-nerved, not reticulate, two linear-lanceolate, less than 1 mm
wide, the fourth cleft to within 2.5 mm of the base, the lobes lan-
ceolate, about 1 mm wide. Corolla apparently white, 5 mm long,
glabrous, the tube not or very slightly contracted, the upper lip
elliptic-ovate, slightly retuse, 1.5 mm wide, the lower one cleft into
three, oblong, obtuse lobes, about 2 mm long, 1 mm wide.
Anthers about 0.8 mm long. Capsule 4.5 mm long, glabrous
except the puberulent apex.

MINDoRO, Baco River, McGregor 128, March 15, 1905.

A species well characterized by its slender, nearly glabrous spikes; it is
similar to Lepidagathis laxa Nees in some respects, but differs from that
form in the points indicated in the diagnosis; from L. incurva Ham., to
which it is also manifestly allied, it differs in its slender, nearly glabrous
spikes.

LEPIDAGATHIS HUMILIS sp. noy.
Herba parva, plus minusve prostrata, ramis floriferis erectis
vix 10 em altis; foliis in paribus inaequalibus, breviter petiolatis,
ovatis vel late ovatis, usque ad 2 cm longis, basi late truncatis,
rariter leviter cordatis vel decurrentibus; spicis axillaribus termi-
nalibusque, solitariis, laxis, angustis, usque ad 5 cm longis, vix 5
mm latis, calycis lobis bracteisque vix ciliatis, leviter hispidis.

¥, °C, 6 Merrill: Philippine Plants, XII 347

A small, nearly glabrous herb, the stems prostrate and rooting
at the nodes, 4-angled, obscurely puberulent or glabrous, the
flower-bearing branches erect, slightly branched, less than 10
em tall. Leaves in unequal pairs, the larger ones of each pair
ovate to broadly ovate, 1.5 to 2 cm long, about 1.5 cm wide, the
base broad, truncate or sometimes subcordate, rarely somewhat
decurrent, the apex acute or blunt, margins slightly undulate,
the smaller leaves of each pair similar but less than one-half as
large, both surfaces with small, scattered cystoliths; petioles 3
mm long or less. Spikes terminal and axillary, solitary, slender,
1 to 5 cm long, less than 0.5 cm in diameter, somewhat inter-
rupted, the bracts and bracteoles similar, broadly lanceolate, acu-
minate, 1-nerved, about 3 mm long, 0.8 mm wide, very slightly
hispid, not ciliate nor glandular. Calyx-segments all acuminate,
obscurely hispid with short hairs, not ciliate, 3 to 4 mm long,
one 1.2 mm wide, very obscurely 3-nerved, very obscurely retic-
ulate above, two narrowly lanceolate, 0.7 mm long, 1-nerved, the
fourth cleft to within 1.5 mm of the base, the lobes lanceolate,
1-nerved. Corolla 4 mm long, pale pink, glabrous, the tube not
constricted, the upper lip 1.5 mm long, 1.1 mm wide, rounded,
the lower one 3-cleft, the lobes elliptic-ovate, obtuse, nearly 1 mm
wide. Anthers 0.6 mm long, the cells parallel.

MINDANAO, District of Davao, Santa Cruz, Williams 2955, June 19, 1905.

A species perhaps as closely allied to Lepidagathis tenuis C. B. Clarke
as to any other Philippine species, but quite distinct from that form. Well
characterized by its small size, small, unequal, broadly ovate, short-petioled
leaves and its slender, nearly glabrous spikes.

RUELLIA Linnaeus

RUELLIA PANAYENSIS sp. nov.

Species R. nudispicae C. B. Cl. affinis, differt brevissime petio-
latis, inflorescentiis fasciculatis haud solitariis tenuioribus,
interdum ramosis.

An erect herb about 20 cm in height, simple or sparingly
branched, the stems terete, 2.5 mm in diameter or less, ciliate-
hirsute, the younger parts rather densely so. Leaves oblong-
ovate, chartaceous, 4 to 9 em long, 2 to 4 cm wide, somewhat
olivaceous, shining, apex rounded, base acute or subacute, the
upper surface glabrous, the lower more or less hirsute with
short hairs especially on the midrib and nerves, margins entire
or obscurely undulate; lateral nerves about 8 on each side of the
midrib, slender; petioles pubescent, 3 to 5 mm long. Inflores-
cence mostly spikelike, apical, fascicled, 3 or 4 from each stem,
8 to 12 cm long, slender, glabrous or very slightly pubescent,

3848 The Philippine Journal of Science 1915

occasionally branched, mostly simple. Bracts linear, 2, acumi-
nate, 2.5mm long. Sepals 5, equal, linear-lanceolate, acuminate,
about 8.5 mm long, 1 mm wide, slightly pubescent, free to the
base. Corolla (very young) apparently about 1 cm long; sta-
mens 4, all fertile, the anthers 2-celled, cells contiguous. Ovary
oblong, glabrous; style-arms one only, short. Capsule linear,
slightly pubescent, 8 to 9 mm long, 1.5 mm wide, the sides
parallel, seed-bearing to the base. Seeds about 12, on retinacula,
somewhat compressed, about 1 mm in diameter.

PANAY, Capiz, Bur. Sci. 21236 Escritor, June, 1913.

It is not entirely clear that the specimen came from Panay, as from the
field label it seems probable that it was collected on the neighboring Island
of Romblon; the locality is given simply as “Capiz, Mount Romblon.” The
species is manifestly allied to Ruellia nudispica C. B. Clarke (Gymnosta-
chyum nudispicum Elm.), which is a true Ruellia in the sense that Clarke
and Lindau interpret the genus; it differs notably in its inflorescence which
consists of fascicled spikes, rarely branches, instead of solitary spikes.

HEMIGRAPHIS Nees

HEMIGRAPHIS BAKERI sp. nov.

Planta parva, glabra vel subglabra, circiter 20 cm alta, parce
ramosa, ramulis plus minusve geniculatis, tenuibus; foliis in
paribus aequalibus, obovato-oblongis, obtusis, integris vel leviter
undulatis, basi angustatis, cuneatis, usque ad 3 cm longis; spicis
tenuibus, paucifloris, bracteolis angustis, quam sepalis haud
latioribus, 4 ad 5 mm longis; calycis lobis lineari-lanceolatis, acu-
minatis, circiter 5 mm longis, glabris; corolla 11 mm longa;
capsulis lineari-oblongis, 7 mm longis, 1.7 mm diametro, semi-
nibus circiter 12.

A small, nearly glabrous, sparingly branched plant 20 em high
or less, the branches slender, more or less geniculate in the lower
part, the very young ones slightly pubescent, soon quite glabrous.
Leaves opposite, those of each pair equal, oblong-obovate; char-
taceous greenish-olivaceous when dry, with prominent cystoliths
on the upper surface, apex rounded to obtuse, base gradually nar-
rowed, cuneate, margins entire or obscurely undulate, 1 to 2.5 cm
long, 4 to 8 mm wide, the lateral nerves about 4 on each side of
the midrib, slender; petioles 2 mm long or less. Spikes terminal,
slender, few-flowered, up to 5 em long, the flowers distant, brac-
teoles not imbricate except toward the tips of the spikes. Calyx
about 8 mm long, glabrous, the tube 3 mm long, the lobes 5,
linear-lanceolate, acuminate, equal, about 0.6 mm wide below.
Corolla about 11 mm long. Anthers 1.3 mm long. Capsule
linear-oblong, about 7 mm long, 1.7 mm wide, apiculate, glabrous,

XC, 5 Merrill: Philippine Plants, XII 349 *

not at all clavate. Seeds about 12, brown, slightly compressed,
obtuse, base inequilaterally cordate, somewhat shaggy-ciliate,
about 1.5 mm long. Bracteoles linear, 4 to 5 mm long, green,
1 mm wide or less, the lower ones slightly spatulate, glabrous.

LUZON, Province of Tayabas, hills near Malimao, C. F. Baker 3272
(type), May 7, 1914. Samar, For. Bur. 21081 Sherfesee, Cenabre, & Cortes,
April, 1914. .

A species probably belonging in the group with Hemigraphis fruticulosa
C. B. Clarke, but nearly or quite glabrous, with entirely differently shaped
leaves and relatively very narrow bracteoles.

[Vol. X, Sec. C, Nos. 3 and 4, including pages 159 to 285, were issued
July 13 and August 9, respectively.]

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