Contributions to Canadian
Palaeontology
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OTTAWA
F. A, ACLAND
PRINTER TO THE KING’S MOST EXCELLENT MAJESTY
1930
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CANADA
DEPARTMENT OF MINES
Hon. Charles Stewart, Minister; Charles Camsell, Deputy Minister
NATIONAL MUSEUM OF CANADA
W. H, Collins, Acting Director
BULLETIN No. 63
Geological Series No. 51
Contributions to Canadian
Palaeontology
OTTAWA
F. A. ACLAND
PRINTER TO THE KINO’S MOST EXCELLENT MAJESTY
1930
CONTENTS
Page
New Species of Marine Invertebrate Fossils from the Bearpaw Formation of
Southern Alberta: M. Y. Williams, University of British Columbia 1
New Species of Invertebrate Fossils from the Non-marine Formations of Southern
Alberta: W. S. Dyer, Department of Mines, Toronto 7
Fossil Plants from the Cypress Hills of Alberta and Saskatchewan: Edward W.
Berry, Johns Hopkins University 15
A New Specimen of Eodelphia cutleri from the Belly River Formation of Alberta:
George Gaylord Simpson, American Museum of Natural History 29
Fossils from Harrison Lake Area, British Columbia: C. H. Crickmay, University
of California at Los Angeles 33
83259-H
NEW SPECIES OF MARINE INVERTEBRATE FOSSILS FROM
THE BEARPAW FORMATION OF SOUTHERN ALBERTA
By M. Y. Williams, University of British Columbia
Illustrations
Plates I and II. Illustrations of fossils
Page
68-71
INTRODUCTION
In a forthcoming memoir by W. S. Dyer and the writer of this article,
the general geology of southern Alberta is dealt with; a chapter on strati-
graphy includes lists of fossils found in the different formations of the
region. Descriptions of new species of fossils have, however, been left
for publication in this bulletin. The marine faunas, studied by the author,
belong to the Pierre sea of Upper Cretaceous age. The marine formations
represented are, from older to younger, Pakowki shale, Bearpaw shale,
and a lower, marine shale division of the Fox Hills formation.
The author wishes to acknowledge his indebtedness to Dr. C. H.
Crickmay, who made a preliminary study of the ammonites, and called
attention to the new species herewith described. The ammonites were
later studied in detail by Louis G. Millward, under the author’s super-
vision, at the University of British Columbia.
Of twelve species of pelecypods from the Pakowki shale, thirty-four
species from the Bearpaw shale, and thirty-one species from the Fox
Hills formation, as identified by the author, only one new variety was
discovered. It is described below.
Of the ammonites, one new species is described, one form is referred
to a western rather than to an eastern species, and two species are referred
to different genera from those in which they were originally placed. The
account of the ammonites is taken, with modifications, from a thesis by
Millward.
DESCRIPTIONS
Class, PELECYPODA
Order, Teleodesmacea
Veniella subtrapeziformis (Whiteaves) var. dyeri var. nov.
Plate II, figures 4-7
Small, sub-trigonal, gibbous, especially below and backward from the
umbo, from which a rounded slope emends to the posterior bf»Ral extremity.
2
Beaks prominent, incurved, directed anteriorly, and overhanging the
anterior dorsal margin. Basal margin gently arcuate, posterior dorsal
margin strongly curved to the short hinge-line.
Shell thin, smooth closed, marked by varying lines of growth; lunule
absent, escutcheon short and narrow.
Pallial line entire, muscular scars as in Arctica but not well defined in
the specimens studied.
Teeth, as seen in sectioned specimens, consist of 3 cardinals and one
lateral in each valve, generally similar to those of the V. conradi (Morton),
This form corresponds in general characters with “C^/prmo" suh-
trapeziformis Whiteaves^ Whiteaves did not see the hinge of his species
and placed it doubtfully in “Cyprina” stating that it might belong instead
to Cypricardia or Veniella.
y. suhtrapeziformis var. dyeri differs from Whiteaves' form in being
smaller, more gibbous, having more overarching beaks and a more arcuate
basal margin. The following measurements illustrate the differences.
V
Measurements in mm.
Whiteaves’
species
Typical specimens of
new variety
1
2
3
4
5
Height
15
12
14
13
13
14
Rroadth at iimho
10
11
12
11
11'7
12
18
Length
23-5
16
17
16
17
Variety dyeri appears to belong to Stoliczka's genus Veniella, but
differs from the general form in lacking a lunule. Its subquadrate or
sub-trigonal shape separates it from Arctica. Within the genus Veniella,
it is somewhat similar to V. suhtumida Meek and Hayden, from the Fox
Hills and Fort Pierre groups of Yellowstone river, but differs from that
species in somewhat smaller size and less rounded and prominent posterior
umbonal shape. The depression in front of the slope in suhtumida is
entirely lacking in internal casts of the present species, which have been
used in comparison with the figures of internal casts of suhtumida in United
States Geol. Survey of the Territories, vol. IX, PI. 17, figs. 5a, b.
Occurrence. In the Baculites compressus zone 450 feet above the
base of the Bearpaw formation, as exposed on Gap creek, sec. 24, tp. 9,
range 27, W. 3rd mer. Collector, W. S. Dyer. Two lots of fossils, one
marked SW. \ sec. 24; the other marked sec. 24, presumably came from
nearly the same place. Together they contain upwards of eighty separate
specimens, in fair state of preservation, and several pieces of rock packed
full of specimens. Strangely enough the form has not been noted else-
where in the region.
With this species are found Chlamys nehrascensis M. and H., BacuUtes
compressus Say, and Dentalium gradle H. and M.
>Geol. Surv,, Canada, Cont. to Can. Pal., vol. I, p. 29, FI. XXIV, figs. 2, 2a, 2b.
3
Class, CEPHALOPODA
Sub-class, TETRABRANCHIATA
Order, Ammonoidea
Ammonites were collected from the three formations laid down in
the western Canadian extension of the Pierre sea. These formations are,
from oldest to youngest, the Pakowki shale ( = Claggett of United States),
the Bearpaw shale, and the lower marine shale member of the Fox Hills
“sandstone’* formation. The species found are listed below.
Species
Pakowki
shale
Lower
beds,
Bearpaw
Upper
beds,
Bearpaw
Lower
Fox Hills
shale
Baculites compressus Say
?
z
X
r
Baculites graiidis H. and M
X
Baculites crickmayi n.s
X
Placenticeras intercalare Meek
X
X
Placenticeras meeki Boehm
X
X
X
Rha^oceras haUi Meek
X
Acanthoscaphites nodosus (Owen) var. brevis Meek
7
X
X
Baculites compressus probably occurs in the Pakowki shale as reported
by Dowling^ B, asper Morton has not been recognized, however, nor
has B. ovatus Say.
Baculites crickmayi n.sp,
Plate I, figure 1; Plate II, figures 1, 2, 3
Fossil in the form of an internal cast. Elongated; of fairly large
size, and rather rapidly tapering compared with other Baculites. Cross-
section ovate, tapering toward the siphonal side. Surface of cast marked
by broad, rounded, arcuate undulations, which commence at the dorsal
and pass obliquely down and backward in a broad curve, terminating
abruptly on the ventral-lateral region.
Septa not crowded, lobes and sinuses deep and divided into somewhat
divergent, digitate branches. Siphonal lobe about as long as wide and
provided with two terminal, more or less spreading, branches, each of
which has generally three, but in some cases two, nearly equal, digitate
branchlets at the end, and two similar lateral ones on the outer side.
First lateral saddle two-thirds as wide as long, much narrower than the
siphonal lobe, and divided at the anterior end into two nearly equal branches
each of which is subdivided into three or four spreading, digitate branch-
lets. First lateral lobe nearly twice as long as wide, and divided at its
end into two nearly equal parts, each with three spreading and digitate
subdivisions. Second lateral saddle very similar in branches and sub-
divisions to the first and only a little larger in size. Second lateral lobe
about as broad, but shorter than the first, and bearing two large, equal,
iQeol. Surv., Canada, Mem. 93, “Southern Plains of Alberta”, p. 47.
4
bipartite, digitate terminal branches, and small digitate and simple lateral
branches. Third lateral saddle much smaller than either of the others,
with two unequal, short, digitate terminal divisions, and a few short,
irregular, smaller lateral branchlets. Dorsal or antisiphonal lobe very-
small, much longer than wide, with one or two small lateral branches,
and a trifid extremity.
As there are only two incomplete specimens, exact measurements can-
not be obtained, but the larger specimen in hand has a greater diameter
of 46 mm. and a lesser diameter of 36 mm. From the average taper of
the cast the original shell must have been over 700 mm. in length when
complete.
In cross-section the specimen rather resembles the broad variety of
B. ovatus figured by Meek^, but it differs from that species in the well-
marked ridges on the shell and by the much more regular suture with its
deeper lobes and saddles. Indeed the suture is much more deeply incised
than any of those figured in the publications available to the author. The
ornamentation resembles that of B. aquilaensiSy but it differs in the cross-
section and suture.
The smaller specimen shows that these sutural characters are distinct
from near the larval condition. The lip of the living chamber is, appar-
ently, like that of BacuUtes compressus.
The specimen does not resemble any previously described BacuUtes
even closely enough to be referred to it as a variety. Its closest relatives
appear to be B. ovatus and B, compressus, being as much like one as the
other. For these reasons the author feels justified in referring it to a
new species.
To designate this species the writer proposes the name BacuUtes
crickmayi, after Dr. Colin H. Crickmay, who first examined the collection.
Occurrences. (1) Mid-line, between SE. and SW. \ sec. 25, lot 9,
range 6, W. 4th mer. (2) Coulee, west of road 3| miles south of
Irvine, Alberta. Collector, W. S. Dyer.
Genus, Rhaehoceras Meek^
Meek described the only known species of this genus as Phylloceras ?
halU, expressing a strong doubt regarding its position in Suess’ genus
which was founded on Jurassic forms. Continuing, Meek cites the following
variations from the generic characters of Phylloceras Suess:
“Its septa differ in some details, such, for instance, as the proportionally smaller
size of its first lateral lobe as compared with tne siphonal lobe, and the more nearly bipartite
termination of the former: also in the less obtusely-rounded terminations of the sub-
divisions of the lateral sinuses."
Of a large distorted specimen he says:
“ I am strongly impressed with the belief that its outer volution naturally made two
deflexions from the regular curve of those within." ..... “This deflexion, or
departure, from the regular curve, makes the outer volution much less deeply embracing
than the inner, and the umbilicus consequently much larger proportionally in the adult,
than in the young and medium-sized specunens.”
*U.S. Geol. Surv., Terr., vol. IX, PL 20.
>Ueek, F. B.: U.S. Geol. Surv., Terr., vol. IX, pp. 468-462 (1876).
5
III conclusion Meek says:
“Should complete undistorted specimens show as I think very probably will be the
case, that it differs generically or subgenerically from Phylloceras proper, I would propose
for the group into which it would in that case naturally fall, the name Rhaeboceras.”
Rhaeboceras halli Meek
Ammonites halli
F. B. Meek and F. V. Hayden: Acad. Nat. Sci., Phil. Proc., vol. VIII
(1856), p. 7; vol. XII (1860), p. 420.
F. B. Meek: Smith. Check-List of North American Cret. Fossils
(1864), p. 24, Fig. 64.
Phylloceras f halli
F. B. Meek: U.S. Geol, Surv., Terr., vol. IX (1876), pp. 458-462,
PI. 24, figs. 3a, b, c.
Description from Meek:
“Shell attaining a large size, moderately compressed discoidal; volutions with their
convexity about equalling two-thirds their diameter from the ventral side to the rather
narrowly rounded periphery, in young and medium sized examples, each embracing nearly
the entire breadth of the next within, but the last one in the adult becoming proportionally
less deeply embracing; umbilicus very narrow, and rather deep in the young but propor-
tionally wider in the adult; surface ornamented with numerous small bifurcating, sligMly
flexuous costae, that are larger near the umbiUcal side, and on the la.st turn of medium
and large sized specimens become proportionally somewhat more prominent, more curved,
and suddenly bifurcate near the umbilicus, and again divide and subdivide into num-
erous smaller ones, so that their number, including others intercalated betweenj amounts
to from five to seven times as many where they pass straightly over the periphery, as
near the umbilical side; body-chamber forming at least the entire outer volution.
Locality and Position. One hundred and fifty miles above the mouth of Milk riverj
on the Missouri, in Montana territory; from the Fort Pierre group of the Upper Missouri
Cretaceous series.”
Two specimens are found in the present collection, one being only
15 mm. in diameter and probably far from complete, and the other 11 cm.
across and possessing more than 6 cm. of living chamber, measured along
its central axis. This larger specimen is well preserved, and has been
dissected so as to show surface markings, suture lines, umbilicus, etc.,
of both young and old stages of development. The two flexures of curva-
ture are well shown, one occurring at the last septum and the other about
4 cm. back along the periphery. The expansion of the umbilicus is also
well shown.
Thus in an undistorted specimen, the characters which Meek noted
as separating this form from the genus Phylloceras are verified. Crickmay
concluded after examining this specimen that Rhaeboceras was established
as a separate genus, which is not related to Phylloceras as Meek supposed,
but is probably descended from the Hoplitidae.
Occurrences. In the Baculites compressus zone of the Bearpaw for-
mation, on the south branch of Box Elder creek, sec. 30, lot 10, range
29, W. 3rd mer. (Sask.). Collector, W. S. Dyer.
6
Family, placenticepatidae
Genus, Placenticeras Meek
Placenticeras meeki J. Boehm
Placenticeras placenta (deKay) (part.)
F. B. Meek: U.S. Geol. Surv., Terr., vol. IX (1876), pp, 465-468,
PI. 24, fig. 2.
Geol. Surv., Canada, numerous reports on western Great Plains.
Placenticeras meeki J. Boehm
J. Boehm: Deutsche Geol. Gesell. Zeitsche., vol. L (1898), p. 200
(footnote) .
J. B. Reeside, jun.: U.S. Geol. Surv., Prof. Paper 151 (1927), pp. 29-30,
PI. 22, figs. 5-7; Pis. 23-24; PI. 25, figs. 1-2.
Placenticeras whitfieldi Hyatt
A. Hyatt: U.S. Geol. Surv., Mon. 44, pp. 221-232, PI. 45, figs. 3-16;
PI. 46; PL 47, figs. 1-4 (1903).
A. W. Grabau and H. W. Shimer: North American Index Fossils,
vol. II (1910), p. 218, figs. 1493-1494.
In some specimens, on being broken down, lines of tubercles may
be seen on the young shell. These are all small and are generally absent
on the larger volutions. Hyatt, in one place^ regards them as merely
individual reversions to the primitive type, but in the specific description*
refers them to a variety ‘‘tuberculatum” j stating them to be intermediate
between this species and P. intercalare.
The species can be distinguished from P. placenta of the Atlantic
Cretaceous by certain characters which, however, can be observed only
in comparing the two species. The venter is narrower throughout life
and becomes less completely rounded in the gerontic stage. This rounding
also comes at a much larger size than in P. placenta. Tubercles are lacking
in the typical forms, but when present are small, whereas those of P.
placenta are large and elongated, much coarser, and less numerous. The
suture is also more complex and sinuous than that of the eastern form.
Of the specimens preserved in the Bearpaw collection, the largest
specimen, when complete, had a greater diameter of 365 mm. and a width
of 75 mm.
Occurrences, (1) East side of Oldman river, sec. 20, tp. 8, range 22,
W. 4th mer. (2) Immediately north of the old Canadian Pacific Railway
bridge over St. Mary river. Fox Hills, lower shalo — Cypress lake.
Collector, M. Y. Williams.
(3) Coulee, west of road 3| miles south of Irvine, Alberta.
(4) Sec. 22, lot 9, range 23, W. 4th mer., Oldman river. (5) Sec. 1, lot 11,
range 3, W. 4th mer., Ross creek. (6) Mid-line between SE. and SW. |
sec. 25, lot 9, range 6, W. 4th mer. Collector, W. S. Dyer.
*Hyatt, A.: U.S. Geol. Surv., Mon. 44, p. 190 (1903).
*Hyatt, A.: U.S. Geol. Surv., Mon. 44, pp. 211, 232 (1903).
7
NEW SPECIES OF INVERTEBRATE FOSSILS FROM
THE NON-MARINE FORMATIONS OF
SOUTHERN ALBERTA
By W. S. DyeTf Department of Mines, Toronto
Page
72-75
Illustrations
Plates III and IV. Illustrations of fossils
The invertebrate fossils of the non-marine formations of southern
Alberta will be listed and commented upon by the writer in a forthcoming
memoir by Williams and the present writer, on the geology of southern
Alberta and southwestern Saskatchewan. The non-marine formations
include the Belly River, Edmonton, and St. Mary River of Upper Creta-
ceous age; the Paskapoo, Willow Creek, and Porcupine Hills of Eocene
age; and the Ravenscrag, probably also of Eocene age. In this paper,
which should be regarded as complementary to the memoir, the new species
and varieties are described and figured. The writer is indebted to Frank
H. McLearn of the Geological Survey, Canada, for advice and assistance
during the progress of the work, and to Dr. T. W. Stanton of the United
States Geological Survey for allowing free access to the extensive fossil
collections in the United States National Museum in Washington.
PELECYPODA
Unio suhprimaevis sp. nov.
Plate III, figure 2
This form is similar to U. primaevis White in possessing peculiar,
posterior, radiating ridges; it also has anterior, radiating ridges of the
same order. These ridges are not mentioned in White’s description of
the type of U, primaevis, but are present on forms referred to it in the
United States National Museum in Washington. U. suhprimaevis also
has the umbonal ridge characteristic of U. primaevis and the furrow in
front of it. The new species differs from White’s form in being considerably
smaller, and in having the umbonal ridge less prominent and the furrow
correspondingly more shallow.
Dimensions of Type. Length, 26 mm.; height, 14 mm.
Horizon and Locality of Type, Foremost beds, South Saskatchewan
river one mile below the mouth of Bow river.
Unio rncJearni sp. nov.
Plate III, figure 15
Several specimens of a highly ornamental Unio were collected from
the Foremost beds on South Saskatchewan river. They are similar to
Unio holmesianus White, redescribed and figured by Stanton^ but differ
>Stanton, T. W.: U.S. Geol. Surv., Prof. Paper 98.
8
in several details of ornamentation. The new species lacks the short,
transverse ridges which in U. holmesianus are “nearly at right angles
to the lines on the front of the shell”. It also lacks the thread-like, raised
lines which in White's species run from the beak down the crest of the
umbonal ridge. In U. mclearni the V-shaped sinuses of ornamentation
reach to the beak and are not preceded by the sharp, slightly undulating,
concentric ridges nearly parallel with the growth lines. Still another
difference is that in U. holmesianus the apex of the deepest sinus of orna-
mentation is in the broad umbonal furrow, whereas in the new species
it lies in front of the furrow.
Dimensions of Type (a slightly distorted form). Length, 34 mm.;
height, 29 mm,; thickness, about 22 mm.
Unio humei sp. nov.
Plate IV, figures 1 and 4
In the collections made by T. C. Weston in 1888 from Fossil coulee
(about secs. 7 and 8, tp. 4, range 19, W. 4th mer.) there are certain specimens
of Unio which in form resemble the marine shell Arctica ovata.
The shell is transversely ovate, moderately convex, and the valves
quite thick; the anterior extremity evenly rounded, but the posterior
extremity narrowly rounded; basal margin semi-oval; the posterior dorsal
part of the type is partly broken away, but other specimens show it to be gently
convex, the anterior dorsal margin is somewhat more abruptly rounded;
beaks moderately elevated and located midway in the anterior third of
the shell. Internally the shell is quite thick and heavy; the lateral teeth
have not been seen and the smaller of the cardinal teeth is not well pre-
served; the cardinal tooth which is preserved is large.
It is rather close to the form, referred doubtfully to Unio pyramida-
toides by Stanton^, from the Fruitland formation of New Mexico, but
is longer in proportion to height and the posterior margin is not truncated.
Dimensions of Type. Length, 78 mm.; height, 64 mm.
Horizon and Locality of Type. Pale beds; Fossil coul4e (about sec. 17,
tp. 4, range 19, W. 4th mer.).
GASTEROPODA
Viviparus crickmayi sp. nov.
Plate III, figure 3
This species differs from V. tasgina, the most closely related form,
chiefly in the greater prominence of the shoulder on the upper part of the
whorl.
The lower two-thirds of the body whorl is evenly rounded, but on
its upper part there is a shallow, revolving depression surmounted by a
moderately prominent, rounded ridge. Above the ridge the outline of
the whorl turns in abruptly almost at right angles to the axis of the shell
to form the shoulder. In many cases the shoulder is 3 mm. wide. It is
usually quite flat, but occasionally slightly concave. Proceeding toward
‘Stanton, T. W.: U.S. Geol. Surv., Prof. Paper 98, p. 309 et seq. (1916).
9
the apex of the shell the ridge and the shoulder become less prominent
and the angle between the shoulder and the axis acute as in V. tasgina.
Finally, near the apex, the ridge and the shoulder fade out entirely. The
character of the apex and the first few whorls leaves no doubt of the close
relationship of the species with V. nidaga. The apex is obtuse and the
first few whorls are quite angular at the base.
Horizon and Locality. It occurs abundantly at one locality in the
Willow Creek formation on Willow creek, i.e., centre of sec. 12, tp. 10,
range 27, W. 4th mer. All the specimens at this locality were from a
drift flock, but judging from the conditions in the field the writer believes
that it could not have travelled far from its original position. A few
specimens were also found in the Edmonton formation on Bow river.
Dimensions of Type. Length, 27 mm.; width, 20 mm.; apical angle,
55 degrees.
Viviparus nidaga sp. nov.
Plate III, figure 1
Some of the forms which Meek^ regarded as V. conradi depart rather
widely from the typical form of the species. They are larger, more elongate,
and have more rounded whorls. It is possible that they are simply speci-
mens more advanced in size or age as Meek suggests, but they approach
V. nidaga, a new species from the Belly Eiver formation, rather closely.
In Alberta specimens have been found which definitely link the two species.
The more typical form of V. conradi is represented by Meek’s figures 15a
and 15b, and the divergent form by figures 15c and 15d. V. conradi is
rare in the Pale beds, but more common in the Foremost member of the
Belly River formation.
Viviparus nidaga differs from V. conradi, the only species from the
Belly River formation for which it might be mistaken, in the greater
convexity of the whorls, in the absence of the angle at the base of the body
whorl, and in the more elongate shape of the shell. The spiral whorls
of the new species, however, are angular at the base. Certain species
are clearly intermediate between V. conradi and V. nidaga, leaving no
doubt of the close relationship of the two species.
V. nidaga is close to V, leai, which occurs in several post-Bearpaw
formations, differing chiefly in being larger and more slender in shape
and in having more angular spiral whorls. It is rare in the Pale beds
and in the Foremost member of the Belly River formation. The specific
name means “prairie chicken” in Sarcee Indian.
Dimensions of Type. Length, 27 mm. ; breadth, 22 mm. ; apical angle,
60 degrees.
Several well-preserved specimens of Viviparus prudentius White^
were collected on Pincher creek by Weston in 1883, and by Dawson from
Gooseberry canyon, St. Mary river, in 1881. Both localities are in the
St. Mary River formation. V. prudentius also occurs in the Willow
Creek and Paskapoo formations. This species is the culmination of the
V. nidagor-V. leai-V. prudentius line of development, in which is shown a
progressive rounding of the whorls and flattening of the spire.
iMeek: U.S. Geol. Surv., vol. 9, p. 679, PI. 42, figs. 15a, b, c, d (1876).
2U.S. Geol Surv., 3rd Ann. Kept., p. 467, Pi. 25, figs. 17, 18 (1883).
«
10
Viviparus tasgina sp. nov.
Plate III, figure 5
This species is midway in development between V. nidaga of the
Belly River formation and V. crickmayi of the Willow Creek and Edmonton
formations. This is well shown in the character of the shoulder on the
upper part of the whorls. This shoulder, which is lacking in V. nidaga^
is always present in V, tasgina, but never so well defined as in V. crickmayi.
The three species are similar in the general shape of the shell and in the
character of the apex and spiral whorls.
V. tasgina sometimes attains large sizes, the largest specimen found
measuring 51 mm. in length, and others reach lengths of 33 and 40 mm.
The increase of the apical angle with age, which was seen many times
during the course of the present study, is excellently illustrated by V.
tasgina, and it was noticed that the coarseness of the growth lines also
increased with age. In the larger specimens there is a tendency for the
whorl above the body whorl to be more elongated in the general direction
of the axis of the shell than the younger specimens.
The species is rather rare in the Edmonton formation and one specimen
from the St. Mary River formation was doubtfully referred to it. The
specific name means “policeman'’ in Sarcee Indian.
Dimensions of Type. Length, 51 mm.; breadth, 42 mm.; apical angle,
53 degrees.
Campeloma veivla tenuis var. nov.
Plate III, figure 4
The forms found in Alberta referable to Campeloma veiula Meek^
are highly variable, ranging from short, stout forms with apical angles
of 50 degrees to forms with elevated spires and apical angles of 30 degrees.
They are abundant in the Foremost member of the Belly River formation.
The more slender forms have been separated in the paper under a new
varietal name C. tenuis. Several of the specimens show distinctly the
revolving strije characteristic of C. multilineata and there is no doubt
that the two species are very closely related. These stria? are very often
present on the upper and lower parts of the whorl, but absent from the
middle of the whorl. Occasionally, however, they are present over the
whole whorl. The forms identified as C. multilineata by Whiteaves from
the Belly River formation should be referred to this species.
It was found on examining the collection of C. vetula in the museum
of the Geological Survey in Washington that the forms were stouter than
the major part of the Canadian individuals. These latter slender forms
have hence been separated, but as a new variety.
By a gradual flattening of the whorls and by the gradual appearance
of an angulation at the base of the body whorl, C. vetula tenuis passes
into C. praecursa.
Dimensions of Type. Length, 18 mm.; width, 11 mm.; apical angle,
47 degrees.
iMeek: U.S. Geol. Surv., Terr., vol. 9, p. 678, PI. 42, figs, 14a, b (1876).
11
Horizon and Locality of Type. Foremost beds of the Belly River
formation; northwest of centre of sec. 1, tp. 3, range 12, W. 4th mer.,
southern Alberta.
Campeloma praecursa sp. nov.
Plate III, figure 6
Shell of moderate size, conical in outline; the volutions, which number
five or six in those specimens having the apex preserved, flat-sided; body
whorl slightly angulated at the base; surface marked by fine lines of growth,
which in a few specimens are crossed by obscure revolving strise, suture
very slightly impressed; aperture narrow, ovate; apparently imperforate.
This species is distinguished from C. veiula tenuis by the flatness
of the whorls, and, in consequence, the conical outline of the shell. It is
quite similar to C. producta, but differs in the flatness of the whorls, there
being no hint of a revolving ridge as in that species. C. praecursa is the
forerunner of C. producta and allied species so common in the post-Bearpaw
formations, just as C. vetula is the forerunner of C. multilineata.
It has been found in both the Pale beds and Foremost members of
the Belly River formation, but is more abundant in the lower member.
Dimensions of Type. Length (one whorl missing), 18 mm.; breadth,
10 mm. ; apical angle, 35 degrees.
Campeloma cypressensis sp. nov.
Plate III, figure 7
This species has developed directly from C. producta^ there being
many specimens intermediate in form between the two. C. cypressensis
is distinguished by the very prominent shoulder in the upper part of the
body whorl; this shoulder is so prominent in some specimens as to form
a swollen ring which is much the widest part of the shell. The body whorl
is much more robust porportionately than in C. producta, as a consequence
of which the apical angle becomes considerably larger. The most extreme
specimen measured 18 mm. in length and had an apical angle of 70 degrees.
It occurs only in the Ravenscrag formation in Cypress hills in sec. 29,
tp. 8, range 4, W. 4th mer.
Dimensions of Type. Length, 14 mm.; breadth, 11 mm.; apical angle,
70 degrees.
Goniohasis svhtortuosa mut. tenuis n, mut.
Plate III, figures 9, 10
The remains of a gasteropod species closely resembling G. suhtortuosa
of the Foremost beds was found in the Willow Creek formation in the bed
of Willow creek a few miles west of the village of Granum (sec. 12, tp. 16,
range 27, W. 4th mer.). The Willow Creek form, however, differs in
being somewhat thicker shelled, more slender, and in having coarser growth
lines. Owing to these differences and to its much higher geological horizon,
it is herein regarded as a mutation of G. suhtortuosa.
Dimensions of Type. Length, 12 mm.; breadth, 7 mm.; apical angle,
40 degrees.
I
12
Goniobasis williamsi sp. nov.
Plate III, figure 8
This species resembles G. tenuicarinata of the Willow Creek and
Paskapoo formations in its ornamentation, but differs in its greater com-
parative breadth, more widely divergent apical angle, and in the fact that
the upper carina is always larger in comparison with the second carina,
and always coincides with the widest part of the whorl. It also occurs at
a considerably lower horizon. G. williamsi also closely resembles G. sub-
tortuosa, differing only in the additional ornamentation. In cases of poor
preservation it is difficult to ascribe a specimen to its proper place. The
species is named after Dr. M. Y. Williams who collected the type speci-
mens. It occurs only in the Foremost member of the Belly River for-
mation.
Dimensions of Type. Length, 17 mm.; breadth, 10 mm.; apical angle,
45 degrees.
Goniobasis judithensis minimus var. nov.
Plate III, figure 12
This variety differs from the type species only in its smaller size.
Whereas Stanton^ states that G. judithensis is lai^er than G. tenuicarinata
and averages 22 mm. in length by 13 mm. in breadth, our forms do not
average more than 10 mm. in length by 7 mm. in breadth, and the largest
specimen is only 16 mm. long. It is abundant at two localities in the Pale
beds and other specimens were collected from beds of doubtful ages in
Oyster creek one-half mile from Oldman river.
Dimensions of Type. Length, 9 mm.; breadth (flattened), 8 mm.;
apical angle, 60 degrees.
Goniobasis whittakeri sp. nov.
Plate III, figure 11
Shell variable in size, moderate to small; whorls seven, moderately
convex, those near the apex slender and uniformly rounded, the lower
whorls angulated at the middle, the body whorl almost flat above the
angle and convex below, whorls immediately above the body whorl flat
both above and below the angle; surface marked by numerous (about
forty in each whorl), evenly spaced, fine, revolving strife, an equal number
above and below the angle; strise slightly waving where crossed by faint
irregular lines of growth, presenting a silky appearance to the surface of
the shell; aperture ovate.
Comparatively abundant in the St. Mary River formation and a few
specimens have also been found in the Edmonton formation. The forms
from the St. Mary River formation described by Whiteaves^ as a new
variety of G. tenuicarinata are undoubtedly poorly preserved examples of
this species, and the specimens referred by him (op. cit., p. 21, PI. 3, figs.
4, 4a) to G. nebrascensis also belong to it.
iStanton, T. W.: U.S. Geol. Surv., Bull. 257, p. 117, PL 13, fig. 4 (1906).
‘Gcol. Surv., Canada, Cont. Can. Pal., vol. 1, p. 22, PL 3, figs. 6, 6a.
13
The possession of the silky ornamentation makes the species easily
distinguishable from any other species in Alberta.
Dimensions of Type. Length, 17 mm.; breadth, 9 mm.; apical angle,
35 degrees.
Goniobasis wehbi sp. nov.
Plate III, figure 14
This species differs from Goniobasis convexa (Meek and Hayden^) the
most closely related form in its perfectly smooth surface: G. convexa has
quite well defined revolving ridges which give it the appearance oia Melania.
The new species is also somewhat smaller in size, and the whorls, especially
the body whorl, are somewhat more angular than in G. convexa. Several
samples of the young forms were found along with the adults. In these
the apical angle is larger and the angle at the base of the body whorl sharper
than in the adults. The species is named after Mr. John B. Webb who so
ably assisted the author for two seasons in southern Alberta. Abundant
in the Willow Creek and Edmonton formations, and one specimen was
found in the St. Mary River formation.
Dimensions of Type: Length, 27 mm.; breadth, 10 mm.; apical angle,
25 degrees.
Velatella rectistriata sp. nov.
Plate IV, figures 2 and 3
This species differs from V. haptista White of the Fort Union of Wyo-
ming and Colorado in the character of the colour bands. In V. haptista
the colour bands are irregularly radiating or vein-like, whereas in the new
species they radiate from the beak in straight lines. Very few specimens
of the new species show the character of the spire, but from the material
at hand it would appe'ar that the spire is not so slender as in V. haptista
and not so much coiled. Whiteaves (op. cit., p. 73) records V. haptista
from the Belly River, but all his specimens were found to be referable to
the new species.
Dimensions of Type. Length, 11 mm.; width, 9 mm.; height, 6 mm.
Horizon and Localities. This species occurs only in the Foremost
beds, but examples were found at numerous localities. The best speci-
mens were found on the north side of Oldman river in SW. | sec. 23,
tp. 11, range 18, W. 4th mer. It is always indicative of brackish water
conditions and occurs with Melania, Corhula, and Ostrea.
Melania whiteavesi nodosa var. nov.
Plate III, figure 16
A few specimens from the Brosseau formation in North Saskatchewan
river differ so decidedly from the regular form of M. whiteavesi that it is
herein described as a new variety. The geology of North Saskatchewan
‘G. convexa Meek, Kept. U.S. Geol. Surv., Terr., vol. 9, p. 592, PL 42, figs. 2a, b, and test figure 71 (1876).
93259-2
14
river does not come within the scope of this report, but the new variety
was worked out while studying the genus Melania, and it was thought best
not to leave it undescribed.
The upper whorls of the variety have the ordinary appearance of the
type, but in the lower whorls the transverse ribs have become almost
obsolete. The revolving ridges are also not continuous, but well-developed
nodes are present where the discontinuous ridges and the nearly obsolete
transverse ribs cross. The result is a regular pattern of nodes over the
whole whorl; these nodes are not circular or spiny, but are comparatively
low and noticeably drawn out in the direction of the spiral ridge. In
size and shape it is typical of M. whiteavesi.
Horizon and Locality. Brosseau (Ribstone Creek) formation, North
Saskatchewan river above Saddle Lake creek, sec. 8, tp. 37, range 12, W.
4th mer.
Pupa sp. indet.
Plate III, figure 13
At Nobleford, Alberta, in the St. Mary River formation, one specimen
of a gasteropod was found which should, undoubtedly, be referred to the
air-breathing genus Pupa. It will very likely prove to be a new species,
but as it is incomplete, it was thought best to leave the detailed description
until more specimens are found. It is somewhat similar to Pupa ata-
vuncula White^ from the Upper Green River formation (Eocene), but is
much larger and shows many minor differences.
‘White, C. A.: U.S. Geol. Surv., Terr., Wyoming and Idaho, Con. Invert. Pal., pp. 46-47, PI. 19, fig. 9 (1878).
15
FOSSIL PLANTS FROM THE CYPRESS HILLS
OF ALBERTA AND SASKATCHEWAN
By Edward W, Berry, Johns Hopkins University
Illustrations
Plates V and VI. Illustrations of fossils
Page
7&-79
I recently examined a collection of fossil plants made by Professor
M. Y. Williams in Cypress Hills district of Alberta and Saskatchewan,
which contains several interesting things and which is, so far as I know,
the first formal record of the occurrence of fossil plants in that region.
Cypress hills constitute an upland in the plains country and are
situated a few miles south of the Canadian Pacific railway and about 40
miles north of the International Boundary, between 108| and 110| degrees
west longitude.
The general section is as follows:
Oligoceue conglomerate
Unconformity
Raven scrag beds
Whitemud
Estevan sandstone
Fox Kills
It is no part of my purpose to attempt a summary of the geology.
The district was surveyed by McConnelP who reported on it in 1886.
The present collections of plants come from the Estevan and Ravenscrag.
Mammals, reptilia, and fishes to the extent of some forty species are
known from the Oligocene, having been noticed first by Cope in 1891
and later by Lambe in the strata unconformably overlying the Ravenscrag,
and originally considered by McConnell to be of Miocene age. More
recently dinosaurs have been discovered in both the Estevan sandstone
and Whitemud, which are, therefore, correlated with the Lance formation
as developed in Montana. This leaves the Ravenscrag as the representa-
tive of the Fort Union of Montana, although all three were originally
referred to the Laramie in its original indefinite sense.
The present collections, which are the property of the Geological
Survey, Canada, are from ten localities, although some of these are repre-
sented by material which is not determinable. None is sufficiently large
or varied to afford data for the discrimination of the three horizons men-
tioned, as may be seen from the accompanying table of distribution of
the species identified, so that the conclusions here presented are of
palaeobotanical rather than stratigraphic interest.
Eight species are recorded from the Estevan and of these only two,
represented by rather indefinite material, have been recognized in the
Ravenscrag. This, at first sight, might be thought to indicate a floral
difference, but all of the Estevan plants except the Marchantites and
Trochodendroides sp., both of which are obscure, are found in the Lance
iGcol. Surv., Canada, N.S., voL 1, pt. C (1886).
93259-2)
16
and Fort Union of the United States, where also both genera are repre-
sented by characteristic material, so that the Estevan plants thus far
discovered lack precise stratigraphic significance.
Nineteen species are recorded from the Ravenscrag beds. Two of
these have been discovered in the Estevan and all except the Ginkgo (?)
stones, the fruit tentatively referred to Paliurus (?), and the new species
of Cercocarpus and Leguminosites represent rather widespread types common
in the Lance (ten of the forms), Fort Union (thirteen of the forms), Paskapoo
(seven of the forms), or equivalent horizons. It would appear that the
Ravenscrag beds are properly correlated with the Fort Union as it is now
delimited in Montana and elsewhere in the United States. Evidently
further collecting from the Ravenscrag beds will yield a large flora.
Botanically it may be noted that tiiere are nineteen genera and fifteen
families in thirteen or fourteen orders contained in the present collections
from Cypress hills, including representatives of the Bryophyta, ArthnophyUi,
Ginkgophyta, Coniferophyta, and Angiospermophyta. No Monocotyledons
have been discovered.
So far as they go they have the same ecological significance as the
corresponding larger floras of the same age to the southward, in indicating
a distinctly temperate flora of probably northern origin and possibly
Holarctic in distribution, certainly at least common to North America
and northeastern Asia.
Table of Distribution
Estevan
Ravenscrag
Lance
Ft. Union
Paskapoo
Pre-Lance
Post-Ft. Union
Wilcox
Raton
Post- Laramie
Black Buttes
Hanna
formation
Edmonton
AI archantitea sp
X
Equiaetum ap
X
Equiaetum rhizome
?
X
Ginkgo adiantoidea
X
X
X
X
X
X
X
Ginkgo ? stones
X
X
Taxodium dubium
X
X
X
X
X
X
X
X
Sequoia nordenakioldi
?
X
X
X
X
X
X
Juglans nigella
X
X
X
X
X
Ficus mburnifolia
X
X
Ariatolochia crassifolia
X
X
X
X
Trochodendroides cuneata
X
X
X
X
“ speciosa
X
X
X
X
" sp
X
Cercocarpus ravenscragensis
X
Leguminositea williamsi
X
Celaatrus wardii
X
X
X
** tanrinensia
X
X
X
X
Enonymua splendens
X
X
X
Rhamnites knowltoni
X
X
X
Paliurus (?) sp
X
Trapa microphylla
X
X
X
X
X
X
Viburnum finale
X
X
“ limpidum
X
X
A peibopsia discolor
X
X
X
X
Phyllites aquatictta
X
17
Phylum, BRYOPHYTA
Class, HEPATIC AE
Order, Marchantiales
Genus, Marchantites Brongniart
Marchantites sp.
A small fragment of a narrowly linear, forked, thallose liverwort occurs
in the Estevan beds along a road up a hill in SE. ^ sec. 28, tp. 7, range 3,
W. 4th mer. It is too incomplete for description. Similar but larger
forms occur in the Lower Eocene of France, the Mississippi Gulf embayment,
and in the Lance formation of Montana.
Phylum, ARTHROPHYTA
Order, Equisetales
Genus, Equisetum Linn^
Equisetum sp.
The original and counterpart of the upper part of a long internode
with sheath at the node. Internode preserved for a length of 3-5 cm.
and 8 mm. in maximum width, showing twelve wide, rather flat ridges.
The sheath is about 1 cm. long and is crowned with sharp, linear, lanceolate
teeth free for 6 to 7 mm. and connate for their basal 3 or 4 mm.
The specimen is somewhat similar to the Green River Equisetum
wyomingense Lesquereux, but differs in the greater length of both internode
and sheath. It is distinct from any Tertiary species known to m*e. A
nominal species of Equisetum has been recorded^ by Knowlton from the
Lance formation, and there are three nominal species known from the
Fort Union. These are all poorly known — E. canaliculatum Knowlton
is a robust form with more numerous stem ridges and short sheaths;
E. decidium Knowlton is based upon a large, detached entire sheath; and
E. globulosum Lesquereux is based on a very slender rhizome with tubers.
The present specimens, although probably representing a new species,
are too incomplete to serve as types. Ravenscrag, above a coal seam
below road bridge on central branch of Fairwell creek, sec. 28, tp. 7, range
24, W. 3rd mer.
Equisetum rhizome
A Cctst of a rather stout Equisetum rhizome showing three somewhat
inflated internodes each about 2*5 cm. in length comes from the Estevan,
Eagle Butte near the centre of sec. 35, tp. 7, range 4, W. 4th mer. It is
much like the specimens from the Lance formation of Rocky creek, Sas-
katchewan, which I referred to Equisetum articum Heer^.
^Knowlton, F. H.: Proc. Wash. Acad. Sci., vol. 11, p. 198 (1909).
®Berry, Edward W.: Canadian Field Nat., vol. 38, pp. 131-132, Fig. 1 (1924).
18
Phylum, GINKGOPHYTA
Order, Ginkgoales
Genus, Ginkgo Linn 6
Ginkgo adiantoides (Unger) Heer
Salishuria adiantoides Unger, Syn. PI. Foss., p. 211, 1845.
Ginkgo adiantoides Heer, FI. Foss. Arct., vol. 5, Abt. 3, 1878, p. 21, PI. ii,
f. 7-10; Ward, U.S. Geol. Surv., Sixth Ann. Kept., 1884-85 [1886],
p. 549, PI. xxxi, figs. 5, 6; idem, Bull. 37, 1887, p. 15, PI. i, figs. 5, 6;
Lesquereux, U.S. Nat. Mus., Proc., vol. 10, 1887, p. 35; Knowlton,
idem, vol. 17, 1894, p. 215; Geol. Soc. Am., Bull., vol. 5, 1893, p. 579;
Washington Acad. Sci., Proc., vol. 11, 1909, pp. 197, 198, 204, 213;
Jour. Geol., vol. 19, 1911, p. 370; Penhallow, Rept. Tert. PI. Brit.
Col., 1908, p. 57, text fig. 12.
This species had a Holarctic distribution during the Tertiary and is
recorded from a large number of horizons. It is common in both the
Lance and Fort Union formations of the United States. A single distal
fragment of a leaf occurs in a lignitic clay in the Estevan on Sexton Hill
creek near the headwaters of Medicine Lodge creek, NE. I sec. 21, tp. 7,
range 3, W. 4th mer. Associated with this leaf fragment and also present
at other localities are an abundance of lenticular circular fruits that may
be of the same species.
Ginkgo (?) stones
Poorly preserved fruits of some uncertain plant are associated with
the fragment of a Ginkgo leaf in the Estevan on Sexton Hill creek. They
also occur in the Estevan in SW. i sec. 28, tp. 7, range 3, W. 4th mer.
and in the Estevan above a coal bed in SW. | sec. 6, tp. 8, range 3, W.
4th mer.
These fruits very probably represent the stones of the drupaceous
fruits of Ginkgo, although this is by no means conclusive. They are
approximately circular in profile, lenticular in form, and about 1 cm.
in diameter. There is an equatorial border nearly a millimetre in width —
all features comparable to those of the stones of the existing Ginkgo.
Phylum, CONIFEROPHYTA
Order, Coniferales
Family, pinaceae
Genus, Taxodium L. C. Richard
Taxodium duhium (Sternberg) Heer^
There is little point to discussing this composite species. It has an
incredible range, both geologic and geographic, and is recorded as common
in the Lance formation, although the equally common occurrences in the
Fort Union are generally referred to Taxodium ocddentale Newberry.
Some of each probably represent the other.
^The very extensive synonymy is given in Berry, TJ.S. Geol. Surv., Prof. Paper 01, p, 171 (1016).
19
Poorly preserved material occurs in the Ravenscrag beds associated
with what I have called Sequoia nordenskioldi but lacking the decurrence
which serves to distinguish the latter.
Occurrence, Ravenscrag beds, Ravenscrag butte, near centre of sec,
27, tp. 6, range 23, W. 3rd mer.
Phylum, ANGIOSPERMOPHYTA
Class, DICOTYLEDON AE
Order, Juglandales
Family, juglandaceae
Genus, Juglans Linn4
Juglans nigella Heer
Juglans nigella Heer, FI. Foss. Arct., vol. 2, Abt. 2, 1869, p. 38, PL ix,
figs. 2-4; Dawson, Geol. Surv., Canada, 1875-76, p. 57; Lesquereux,
Rept. U.S. Geol. Surv., Terr., vol. 8 (Cret. and Tert. FL), 1883, p. 235,
PI. xlvi-A, fig. 11; Khowlton, U.S. Nat. Mus., Proc., vol. 17j 1894,
p. 222; Geol. Soc. Am., Bull., vol. 5, 1893, p. 583; Ward, U.S. Geol.
Surv., Sixth Ann. Rept., 1884-85 [1886], p. 551, PI. xl, fig. 6; idem.,
Bull. 37, 1887, p. 33, PL xv, fig. 1; Newberry, U.S. Geol. Surv., Mon.
35, 1898, p. 33, PL li, fig. 2 (in part), 4; Penhallow, Rept. Tert. PL
Brit. Col., 1908, p. 60; Knowlton, U.S. Geol. Surv., Prof. Paper 101,
1918, p. 292, PL Iv, fig. 2; PL Ixiii, fig. 2.
The type locality for this species was in the Kenai formation of Alaska,
and it has subsequently been recorded from the Raton, Fort Union, and
Paskapoo formations. There are several specimens in the Ravenscrag
beds at Ravenscrag butte, near centre of sec. 27, tp. 6, range 23, W. 3rd
mer.
Order, Urticales
Family, mobaceae
Genus, Ficus Linn6
Ficus viburnifolia Ward
Ficus viburnifolia Ward, U.S. Geol. Surv., Sixth Ann. Rept., 1884-85 [1886],
p. 552, PL xlv, figs. 5-9; U.S. Geol. Surv., Bull. 37, 1887, p. 42, PI.
xxii, figs. 4-8.
This species is represented by a single small specimen, which because
of its small size is less cordate than the type. It agrees with the latter in
venation and has a faintly dentate margin. Length 4-25 cm. Maximum
width 3 cm.
Occurrence. Estevan beds, on a hill, SW. ^ sec. 16, tp. 7, range 29
W. 3rd mer.
20
Order, Aristolochiales
Family, Aristolochiaceae
Genus, Aristolochia Linn^
Aristolochia crassifolia Cockerell
Plate VI, figures 1-3 (| natural size)
Caialpa crassifolia Newberry, N.Y. Lyc. Nat. Hist., Ann., vol. 9, 1868’
p. 56; [Lesquereux], U.S. Geol. and Geog. Surv., Terr., 111., Cret. and
Tert. PL, 1878, PI. xxii; Penhallow, Kept. Tert. PI. Brit. Col., 1908,
p. 44.
Aristolochia cordifolia Newberry, N.Y. Lyc. Nat. Hist., Ann., vol. 9, 1868,
p. 74; [Lesquereux], U.S. Geol. and Geog. Surv., Terr., Hi., Cret. and
Tert. PL, 1878, PL xxv, fig. 7; Newberry, U.S. Geol. Surv., Mon. 35,
1898, p. 90, PL xxxix; PL xl, -fig. 7; PL lx, fig. 4.
Aristolochia crassifolia Cockerell, Am. Mus. Nat. Hist., Bull., voL 24,
1908, p. 90.
Cocculus haydenianus Ward, U.S. Geol. Surv., Sixth Ann. Kept., p. 556,
PL 59, figs. 1-5, 1886; Bull. 37, p. 100, PL 47, figs. 1-4; PL 48, fig. 1,
1887; Knowlton, Wash. Acad. Sci., Proc., vol. 11, pp. 189, 198, 200,
213, 215, 1909.
Cehatha haydenianus Knowlton, U.S. Geol. Surv., Bull. 152, p. 62, 1898.
Whatever may be thought of the generic relations of this plant there
is no doubt that the material which Ward referred to Cocculus is identical
with what Newberry referred successively to Catalpa and Aristolochia.
It is obviously not a Catalpa and after comparisons I believe it to represent
Aristolochia. This was Ward’s original idea, subsequently abandoned in
favour of Cocculus. A feature not shown in Newberry’s types but empha-
sized by Ward in connexion with what he called Cocculus haydenianus
is the “marginal vein or hem” for which Ward coined the term parypho-
drome. This is present in the Canadian material and is nothing but the
flattened arches of the ultimate areolation. The species occur abundantly
in the Lance, Fort Union, and Paskapoo formations and highly character-
istic, but usually much broken remains are not uncommon in the Ravens-
crag beds at Kavenscrag butte, near centre of sec. 27, tp. 6, range 23,
W. 3rd mer.
Order, Ranales
Family, trochodendeaceae
Genus, Trochodendroides Berry
This term is used as a form genus for fossil representatives of the
family Trochodendraceae and not as implying any direct relationship to
the genus Trochodendron.
Trochodendroides cuneata (Newberry)
Plate V, figures 2, 3
Populus cuneata Newberry, N.Y. Lyc. Nat. Hist., Ann., vol. 9, 1868, p, 64;
Ward, U.S. Geol. Surv., Mon, 35, 1898, p. 41, PL xxviii, figs. 2-4;
PL xxix, fig. 7; U.S. Geol. Surv., Sixth Ann. Kept.; 1884-85 [1886],
21
p. 550, PL xxxiii, figs. 5-11; U.S. Geol. Surv., Bull. 37, 1887, p. 19,
PI. Iv., figs. 5-8; PL V, figs. 1-3; Penhallow, Kept. Tert. PL Brit. CoL,
'1908, p. 77; Knowlton, Wash. Acad. Sci., Proc., vol. 11, 1909, pp.
185-215.
“Populus nervosa var. 0 elongata Ny.” [Lesquereux], U.S. Geol. and Geog.
Surv., Terr., 111., Cret. and Tert. PL, 1878, PL xiii, figs. 2-4.
^‘Populus nebrascensis Ny.” [Lesquereux], op. cit,, PL xiv, fig. 7.
Populus amblyrhyncha Ward, U.S. Geol. Surv., Sixth Ann. Kept., 1884-85
[1886], p. 550, PL xxxiv, figs. 5-9; PL xxxv, figs. 1-6; idem. Bull. 37,
1887, p. 20, PL vi, figs. 1-8; PL vii, figs. 1-3; Knowlton, Wash. Acad.
Sci., Proc., vol. 11, 1909, pp. 188, 189, 194, 195, 198, 201, 202; Jour.
Geol., vol. 19, 1911, p. 361; in Calvert, U.S. Geol. Surv., Bull. 471,
1912, p. 16.
Populus cyclomorpha Knowlton and Cockerell. U.S. Geol. Surv., Bull,
696, p. 487, 1919.
Populus roiundifolia Newberry, U.S. Nat. Mus,, Proc., vol. 5, 1882 [1883],
p. 506; U.S. Geol. Surv., Mon. 35, 1898, p. 51, PL xxix, figs. 1-4; Pen-
hallow, Kept. Tert. PL Brit. CoL, 1908, p. 79; Knowlton, Wash. Acad.
Sci., Proc., vol. 11, 1909, p. 189. [Homonym, Griffith, 1847.]
Populus cuneata Newberry [Lesquereux], U.S. Geol. and Geog. Surv.,
Terr., 111., Cret, and Tert. PL, 1878, PL xiv, figs. 1-4.
In Ward’s account of the Fort Union flora he recorded thirteen species
of Populus, although Newberry had previously recorded eight species,
only one of which Ward found represented in his extensive collections.
In Knowlton’s list of the Fort Union flora (1919, page 771) twenty-five
species of Populus are listed. It seems obvious a priori that there were
not twenty-five species of this genus in the Fort Union; and when one
examines critically the published illustrations or the named specimens in
the United States National Museum collections, it becomes equally obvious
that a considerable number of these so-called species are without any basis
and were frequently not recognized by their founders. Moreover, one
becomes impressed with the unlikeness of many of them to Populus, a
subject remarked on by Ward as far back as 1887.
Newberry described the present species in 1868 and its chief feature
was its more or less cuneate base. It is hardly necessary to give a detailed
description of my conception of the species, suffice to say that these leaves
are variable in size, and to a less degree in form, orbicular in general outline,
bluntly pointed or rounded at the tip, and cuneate to rounded truncate at
the base with long petioles. The margin may be entire or variably toothed.
There are three basal or sub-basal primaries and the lowermost lateral
secondaries may be sub-primary in character. The main lateral primaries
are ascending and variably subacrodrome in character.
Two extremes of this species are figured from the Ravenscrag beds at
Ravenscrag butte (near the centre of sec. 27, tp. 6, range 23, W. 3rd mer.)
They agree in all features except marginal character and are intimately
associated in the deposit. As can be readily seen they are unlike any
existing species of Populus, particularly in their venation, whereas, on the
22
other hand, they have every feature of the existing Cercidophyllum japoni-
cum, which may be crenate, dentate, or entire, or Tetracentron sinensis.
I hope to discuss the geological history of the family Trochodendraceae
with necessary illustrations in a more appropriate place.
This species is abundant and widely distributed in the Lance and Fort
Union of Dakota, Montana, and Wyoming, and has also been recorded
from Red Deer river, Alberta.
Trochodendroides speciosa (Ward)
Plate V, figure 8
Populus speciosa Ward, U.S. Geol. Surv., Sixth Ann. Kept., 1884-85 [1886]f
p. 550, PI. xxxiv, figs. 1-4; U.S. Geol. Surv., Bull. 37, 1887, p. 20>
PI. V, figs. 4-7; Penhallow, Kept. Tert. PI. Brit. Col., 1908, p. 79;
Knowlton, U.S. Geol. Surv., Mon. 32, pt. 2, 1899, p. 694, PI. Ixxxiv,
fig. 3; Wash. Acad. Sci., Proc., vol. 11, 1909, p. 189; Jour. Geol.,
vol. 19, 1911, p. 369.
Populus xantholitherisis Knowlton, Mon. U.S. Geol. Surv., vol. 32, pt. 2,
p. 695, PI. 85, figs. 1, 2, 1899.
This is a widespread and variable form in the Lance and Fort Union
formations, recorded from South Dakota, Montana, Yellowstone National
park, and British Columbia (Paskapoo formation).
Several leaves from the Ravenscrag may be assigned to this species.
They are from 3-2 to 7-5 cm. in width, and from 4-5 to probably 8-0
cm. in length, with rounded and occasionally markedly cordate base and
rather blunt apex. The margins are crenate-dentate, commencing a
short distance from the top of the petiole and frequently double. The
petiole is long, the preserved part being from 2-0 to 7-5 cm., with a width
of about 1 mm. The venation is palmate, the middle primary compara-
tively strong, with two or three pairs of lateral primaries diverging from
the summit of the petiole, though the principal pair are sometimes
more or less fused with the middle one near the petiole. The basal pair
are much finer and delicate. The first and second pairs adjacent to the
middle one have stronger secondary veins on the outer side than on the
inner side. The secondaries branch, reach the margins of the teeth, and
form series of polygonal loops with tertiaries. The characteristic features
are the rounded teeth and palmate nervation with subacrodrome main
primaries from the top of the petiole.
Occurrence. In Ravenscrag beds above a coal bed, sec. 28, tp. 7,
range 24, W. 3rd mer., below the road bridge on central branch of Fairwell
creek.
Trochodendroides sp.
Two specimens which fail to show details, but apparently with entire
margins, possibly the same as Trochodendroides cuneata (Newberry) or T.
speciosa (Ward) of the Ravenscrag beds.
Occurrence. Estevan beds, SW. \ sec. 16, tp. 7, range 29, W. 3rd mer.
23
Order, Rosales
Family, rosaceae
Genus, Cercocarpus H.B.K.
Cercocarpus ravenscragensis Berry n. sp.
Plate V, figure 6
Leaves small, somewhat inequilateral, obovate or spatulate; margin
prominently dentate distad; teeth diminishing proximad and basal one-
third of margins entire. Apex broadly rounded. Base cuneate. Length
3-5 cm. Maximum width, above middle, about 2 cm. Petiole short,
stout, curved, expanded proximad, about 4 mm. in length. Mid vein
relatively very stout and prominent. Secondaries stout, about seven
fairly regularly spaced pairs; they diverge from the midvein at acute
angles, are relatively straight, and subparallel and camptodrome. Ter-
tiaries relatively stout, not well preserved, forming a rosaceous areolation.
This form is clearly rosaceous in its venation and agrees rather well
with the existing Cercocarpus parvifoUus Nuttall of the somewhat arid
uplands of western North America. The genus comprises arborescent
and shrubby species and is confined to North America in the region between
Idaho and Mexico. Several fossil species have been recorded, mostly
from the later Tertiary, but one species has been described by Knowlton
from the Raton formation of New Mexico.
Occurrence. Ravenscrag beds, north branch of Frenchman river,
SE. I sec. 22, tp. 7, range 22, W. 3rd mer.
Family, leguminosae (Incertae sedis)
Leguminosites williamsi Berry n. sp.
Plate V, figure 1
Leaflet small, obovate in outline, markedly inequilateral, with a
broadly rounded apex and an abruptly cuneate base. Margin entire.
Texture coriaceous. Length about 2 cm. Maximum width about 1 cm.
Midvein stout, curved. Secondaries relatively stout, about five ascending,
camptodrome pairs.
This leaflet is new and represents a type met with in a great variety
of leguminous genera. In the absence of more information than is afforded
by the present material it is impossible to determine its generic affinity,
although it probably represents Cassia or Sophora. It is named for the
collector, Professor M. Y. Williams.
Occurrence. Ravenscrag beds, north branch of Frenchman river, SE. |
sec. 22, tp. 7, range 22, W. 3rd mer.
24
Order, Sapindales
Family, celastraceae
Genus, Celastrus Linn4
Celastrus wardii Knowlton and Cockerell
Celastrus ovatus Ward, U.S. Geol. Surv., Sixth Ann. Rept., 1884-85, p. 555,
PL liii, fig. 7, 1886; U.S. Geol. Surv., Bull. 37, 1887, p. 71, PI. xxvi,
fig. 1; Knowlton, Washington Acad. Sci., Proc., vol. 11, p. 190, 1909.
[Homonym, Hill, 1865.]
Celastrus wardii Knowlton and Cockerell, U.S. Geol. Surv., Bull. 696,
p. 160, 1919.
The type of this species came from the Fort Union formation near
Glendive, Montana. It has been recorded from other localities in the
Fort Union as well as in the underlying Lance formation. A single frag-
mentary specimen was collected from the Ravenscrag beds on north
branch of Frenchman river, SE. | sec. 22, tp. 7, range 22, W. 3rd mer.
Celastrus taurinensis Ward
Celastrus taurinensis Ward, U.S. Geol. Surv., Sixth Ann. Kept., 1884-85,
p. 555, PI. lii, figs. 15, 16, 1886; U.S. Geol. Surv., Bull. 37, p. 79,
PI. xxxiv, figs. 5, 6, 1887; ? Hollick, Geol. Surv., La., Special Rept. 5,
p. 285, PI. xlvi, fig. 1, 1899; Knowlton, Washington Acad. Sci., Proc.,
vol. 11, p. 213, 1909; Berry, U.S. Geol. Surv., Prof. Paper 91, p. 267,
PI. lx, figs. 1-3, 1916.
The type of this species came from the Fort Union near Glendive,
Montana. It is apparently common in the Fort Union and underlying
Lance formation, and has also been reported from the Wilcox Eocene of
the Mississippi Gulf embayment. I believe that Celastrus curvinervis Ward
probably represents the same botanical species.
In the present collections there are several excellent specimens from
the Ravenscrag beds at Ravenscrag butte, near centre of sec. 27, tp. 6,
range 23, W. 3rd mer.
Genus, Euonymus Linn 6
Euonymus splendens Berry
Euonymus splendens Berry, U.S. Geol. Surv., Prof. Paper 91, 1916, p. 267,
PI. Ixii, figs. 1-5; Knowlton, U.S. Geol. Surv., Prof. Paper 101, 1917,
p. 329, PI. Ixix, fig. 1.
This species is one of the commonest and most widespread plants in
the Wilcox of the Mississippi Gulf embayment. What appears to be
the same species has been identified by Knowlton from the Raton formation
of New Mexico. A single large but incomplete specimen in the present
collection is highly characteristic, but lacks the extended tip.
Occurrence. Ravenscrag beds, Ravenscrag butte, near centre of
sec. 27, tp. 6, range 23, W. 3rd mer.
25
Order, Rhamnales
Family, rhamnaceae
Genus, Rhamnites Forbes
Rhamnites knowltoni Berry
Cornus studeri Lesquereux, Rept., U.S. Geol. Surv., Terr., voL 7, 1878,
p. 244, PI. 42, figs. 4 and 5 (not Heer 1859); Ward, U.S. Geol. Surv.,
Bull. 37, 1887, p. 55, PI. 26, fig. 1; Hollick, Rept., La. Geol. Surv.,
1899, p. 286, PI. 45, fig. 2; Berry, U.S. Geol. Surv., Prof. Paper 91,
1916, p. 331, PI. 68, fig. 3.
Rhamnites knowltoni Berry, Idem, 131, p. 16, PI. 12, fig. 7, 1922.
Small leaves represent this species. Unfortunately they are incom-
plete, lacking the upper part. The leaves are large in size, but variable,
with entire margins. The petiole is long and thick, with a width of about
2 mm.
The species was long confused with the European Miocene Cornus
studeri Heer, a quite different form. Rhamnites knowltoni occurs in the
Denver, Raton, and Wilcox.
Occurrence. Ravenscrag beds, north branch Frenchman river, SE. I
sec. 22, tp. 7, range 22, W. 3rd mer.
Genus, Paliurus Jussieu
Paliurus (?) sp.
Plate V, figure 7
A single specimen shows what appears to be some winged fruit resemb-
ling that of Paliurus. The fruit is circular and disk-like, about 2-6 cm.
in diameter but not complete, with the wing showing radial striations.
For an account of a well-preserved Paliurus fruit and a discussion of
the geograpliical and geological range of the genus See Berry, Am. Jour.
Sci., vol. 16, pp. 39-44, 1928.
Occurrence, Estevan, above coal bed. Burnt shale associated only
with abundant circular lenticular fruits not determinable but possibly
representing Ginkgo. SW. | sec. 6, tp. 8, range 3, W. 4th mer.
Order, Myrtales
Family, hydrocaryaceae (?)
Genus, Trapa Linn6 (?)
Trapa (?) microphylla Lesquereux
Trapa microphylla Lesquereux, Rept. U.S. Geol, Surv., Terr. (Tertiary
Flora), vol. 7, p. 295, PI. 61, figs. 16-17a, 1878.
Trapa microphylla Ward, U.S. Geol. Surv., Bull. 37, p. 64, PI. 28, figs. 2-5,
1887; Knowlton, U.S. Geol. Surv., Mon. 32, pt. 2, p. 761, PI. 77, figs.
3, 4, 1898; U.S. Geol. Surv., Bull. 163, p. 62, PI. 5, fig. 7, 1900.
26
Only detached leaves are scattered on several specimens, and some
are much like pinnules of some fern. The leaves are small, varying in size
from 7 mm. to more than 13 mm. in length and from 5 mm. to 10 mm. in
maximum width; orbicular or broadly ovate, rounded or more or less
truncate at the base or at the apex, some a little narrowed to the base and
some narrowed to the apex; entire but slightly undulate on upper margins.
No petiole is preserved. The nervation is not quite distinct, some show a
comparatively thick midrib with 6 or 7 secondary nerves on each side; the
midrib being more or less curved and centric; the secondaries diverging
from the midrib at angles of more than 30 degrees in the lower part and less
than 30 degrees in the upper part, bifurcating below the middle and reach-
ing the margins, the basal ones joining the midrib at the base and likely to
be opposite, the upper ones mostly alternate. The nervilles are not
clearly seen.
There can be no question of the identification of the present specimens,
but most students regard their reference to the genus Trapa as highly
problematical. The present species has been recorded from a variety of
horizons from the Belly River Cretaceous to the Fort Union. Whether
these records represent a single botanical species or not I am not prepared
to say, but whatever it may be or how widely it may range Trapa (?)
microphylla is especially abundant in the Lance and Fort Union horizons.
Occurrence. Estevan beds, along road up a hill SW. J sec. 28, tp. 7,
range 3, W. 4th mer.
Order, Rubiales
Family, caprifoliaceae
Genus, Viburnum Linn6
Viburnum limpidum Ward
Viburnum limpidum Ward, U.S. Geol. Surv., Sixth Ann. Rept., p. 556,
PI. 63, figs. 1-4, 1886; Bull. 37, p. 110, PI. 53, figs. 3-6, 1887.
The leaf is not completely preserved, but the venation is distinct.
It is ovate, about 70 mm. long and about 52 mm. in maximum width;
narrowed and rather rounded at the base, likely to be bluntly pointed at
the apex ; serrate on the upper margins and entire at the base. The venation
is ascending pinnate, with the mid vein more or less curved in somie parts;
the secondary veins about seven on each side, diverging from the midvein
at angles of from 35 degrees to 60 degrees, mostly curving upward, branching
near and above the middle, and craspedodrome ; the nervilles distinct and
simple, mostly parallel each other and percurrent.
It very much resembles Ward’s figures in outline, size, and especially
venation. The type is from the Fort Union of Montana.
Occurrence. Ravenscrag beds, north branch, Frenchman river, SE. j
sec. 22, tp. 7, range 22, W, 3rd mer.
27
Viburnum finale Ward
Plate V, figure 4
V iburnum finale Ward, U.S. Geol. Surv., Sixth Ann. Kept., 1884-85 [1886],
p. 557, PI. Ixv, fig. 8; U.S. Geol. Surv., Bull. 37, 1887, p. 115, PI. Ivii,
fig. 5.
Some of the specimens from the Ravenscrag which I have referred to
this species are relatively wider and with more ascending secondaries than
the type, and I have figured one of these. It show's the same slight in-
equality of the two sides of the lamina, and differences in the distal branch-
ing of the secondaries on the two sides, exactly as in the type. Associated
with it are leaves which agree more exactly with Ward’s type, which latter
came from the Fort Union at Iron Bluff, Glendive, Montana. The speci-
men from Ravenscrag butte, figured, has the complete petiole, hitherto
not known. It is stout and 1*5 cm. long.
(Incertae sedis)
Genus, Apeibopsis Heer
Apeibopsis discolor Lesquereux
Rhamnus discolor Lesquereux, U.S. Geol. and Geog. Surv., Terr., Ann.
Rept., p. 398, 1872.
Apeibopsis discolor Lesquereux, Rept. U.S. Geol. Surv,, Terr. (Tertiary
Flora), vol. 7, p. 259, Pi. 46, figs. 4-7, 1878.
An original and counterpart represent this species. They are ovate,
rounded, and slightly cordate at the base, likely to be acutely pointed at
the apex, and entire; from 5-5 to 7-0 cm. long and about 5-5 cm. in maxi-
mum width. The venation is pinnate, with rather strong midvein more or
less wavy in traversing the whole length and much diminishing toward
the apex; the secondary veins seven or eight on each side, diverging from
the midvein at angles of from 30 degrees to 90 degrees (the more distal
the larger the angles), opposite in the lower part and sub-opposite or
alternate in the upper part, sub-parallel, curving upward, camptodrome,
branching mostly near the margins but som.e near the midrib; the nervilles
connecting the secondaries and tertiaries at various angles, some being
nearly parallel.
The genus w'as proposed by Heer in 1859 for leaves from the Miocene
of central Europe which he supposed were related to the existing genus
Apeiba Aublet, a South American tropical genus of the family Malvaceae.
The correctness of this identification is supported in the case of some of
the fossil species by their association with characteristic fossil fruits, but
this is not the case with any North American species referred to Apeibopsis.
Apeibopsis discolor is not especially close to the leaves of the existing
South American species of Apeiba and though the ancestors of the latter
might be expected to be represented in North America if they occur in
Europe, there is no sound basis for such a conclusion and I have, therefore,
listed Apeibopsis discolor under Incertae sedis.
28
The previous records are in the much disputed Black Buttes beds of
Wyoming, of post-Laramie age, and in the Hanna formation of Wyoming.
Occurrence, Ravenscrag beds, Ravenscrag butte, near centre of
sec. 27, tp. 6, range 23, W. 3rd mer.
Phyllites aquaticus Berry n. sp.
Plate V, figure 5
Several specimens of a delicate plant are contained in the collection
from the Ravenscrag beds. They represent filiform stems with whorls of
slender linear leaves at the nodes. These are lax, with one to three longi-
tudinal veins, and are about 0-5 mm. in width and preserved for lengths
up to 2 cm. None has been observed to branch but whether their distal
parts were branched as in Ceratophyllum is not known. One slab of
material shows a whorl on each side and appears to have been a part of the
same stem. If this is so then the plant was rapidly buried in place and the
internodes were relatively short. The material is evidently not algal, but
a vascular plant.
29
A NEW SPECIMEN OF EODELPHIS CUTLERI
FROM THE BELLY RIVER FORMATION
OF ALBERTA
By George Gaylord Simpson,
American Museum of Natural History
Page
80
Illustration
Plate VII. Illustration of fossil.
INTRODUCTION
On May 30, 1916, Doctor (now Sir) Arthur Smith Woodward announced
the discovery of a lower jaw of a mammal in the Belly River formation
of Albertah A few months later a more detailed description and a figure
of this important specimen, to which he gave the name Cimolestes cutleri,
new species, were published^. In the meantime, without knowledge of
Smith Woodward’s work. Dr. W. D. Matthew described a similar specimen,
naming it Eodelphis hrowni, new genus and species’.
The type of Cimolestes” cutleri was found by William E. Cutler in
1914 on Sand creek, Red Deer river, Alberta, and consists of part of the
right ramus with P 3 , Mi-g, and roots or alveoli of other teeth. That of
E. hrowni was found by Barnum Brown in 1915, also on Sand creek, 15
miles below Steve ville, and consists of the left lower jaw nearly complete
but with the teeth very much worn, a small fragment of the right lower
jaw, and two skull fragments. The importance of these two discoveries
lay chiefly in the fact that they were, at that time, the most complete known
specimens of Cretaceous mammals and that they could be definitely
shown to be marsupials, confirming previous belief that this group was
largely represented in the American Upper Cretaceous. Eodelphis is still
the oldest known marsupial.
The type of “Cimolestes” cutleri has been redescribed by the present
writer,^ who referred it to Matthew’s genus as Eodelphis cutleri (Smith
Woodward). The distinctions between the two specimens, aside from
verbal discrepancies in the original descriptions of the molars, were con-
sidered as probably too slight to warrant recognition of both trivial names,
but these were retained pending some discovery which might provide
definitive knowledge of molar structure in this genus.
This discovery has now been made. During the past summer (1928),
C. M. Sternberg found a third Eodelphis jaw that clearly reveals the molar
structure, permits better correlation of the two previous finds, and adds
another to the four or five really adequate mammalian specimens so far
found in the Cretaceous of this continent. For the privilege of studying
this specimen and of preparing the present remarks, the writer is much
indebted to Mr. Sternberg.
>Zool. Boc. London, Abstract No. 158 (May 30, 1916).
*Proc. Zool. Soc. London, 1916, pp. 525^28 (Sept., 1910).
•Bull. Am. Mufl. Nat. Hist., XXXV, pp. 477^ (July 24, 1916).
•Catalogue of the Mesozoic Mammalia, etc., British Museum (Nat. Hist.), pp. 147-149 (Mar. 24, 1928).
93259-3
30
DESCRIPTION
Eodelphis cutleri (Smith Woodward), May 30, 1916
Synonym: E. browni Matthew, July 24, 1916
Plate VII
New Specimen. Cat. No. 8356, Geological Survey, Canada. Part
of right lower jaw with Mi_ 4 , alveoli of P 3 , and part of jaw posterior to
teeth.
Horizon and Locality. About 100 feet below top of Belly River
formation (Pale Beds), about 2 miles southwest of Steveville, Alberta.
Collected by C. M. Sternberg, July, 1928.
Dentition. The molars increase much m*ore markedly in size from
front to back than in any Tertiary or Recent didelphid. M 4 is over half
again as large as Mi. M 4 is slightly larger than M 3 and its heel is somewhat
wider and longer than that of any of the preceding teeth. Of all the
molars the trigonid is very slightly wider basally than the talonid, of about
equal length, and somewhat higher. On Mi the trigonid is longer than
wide, on M2 length and width are about equal, and on M3-4 the trigonid
is sharply compressed antero-posteriorly, ’width considerably greater than
length. On Mi _2 the trigonids are evenly truncated by wear, but their
bases indicate the protoconid and paraconid as subequal, the metaconid
as smaller. On M3-4 the cusps are relatively well preserved and the
trigonid consists essentially of the sharp, erect, subequal protoconid and
paraconid, united by a shearing edge, while the metaconid is much reduced,
almost vestigial. There is an anterior cingulum, very definite but not
distinctly basined, rising toward the paraconid. The talonids are of the
almost invariable didelphid type, basined, 'with three marginal cusps,
the hypoconulid and entoconid closely approximated. In this case the
latter are almost connate and on M3-4 are slightly higher than the hypoco-
nid, although not so much so as indicated by Matthew, whose specimen
has the hypoconids eroded.
Mandible. The mandible agrees very closely with the E. cutleri type,
the only definite distinction, and that slight, being that the lower border
is apparently more swollen beneath and just behind M4.
Dimensions
Mm.
Length Mu 17*0
“ Ml 3-3
“ Ms 4-1
“ Mb 4-6
“ Mi 5-1
Maximum depth of ramus below alveolar border internally
(beneath Mi) 12 -5
31
DISCUSSION
Taxonomy of Eodelpkis. In no way does this new specimen differ
sufficiently from the type of E. cutleri to permit any taxonomic distinction.
Matthew’s description of E. browni differs from the details given above
chiefly as follows:
( 1 ) He states that second and third molars are similar in size (both
are worn and corroded).
(2) The paraconid and metaconid of Ms are said to be apparently
of equal height and the protoconid lower than either (inferred from bases
of broken cusps).
(3) On M 4 the metaconid is recognized as lower than the paraconid,
but the protoconid (which is broken) is said to be certainly lower than the
paraconid and apparently also lower than the metaconid. The hypoconid
(corroded) is said to be decidedly, rather than slightly, lower than the
hypoconulid and entoconid.
After careful direct comparison it is clear that Matthew’s specimen
must originally have been exactly like the present specimen in all these
particulars, although its true nature in these details could not have been
established until this less worn specimen was at hand. This is insisted
upon only because, as previously suspected, it causes all discrepancy in
molar structure between E. browni and E. cutleri to disappear, confirms
their reference to the same genus, and, as other differences are quite un-
important, necessitates their union as a single species: Eodelpkis cutleri.
Like the type of E. cutleri, the present jaw is somewhat more robust
than Matthew’s specimen (although the teeth are not larger), and the
positions of the mental foramina, though similar, are not exactly the same
in any two of the three specimens. These differences, however, are cer-
tainly well within the range of individual variation.
Relationships to Later Forms. Both Smith Woodward and Matthew
recognized the similarity of this species to “Cimolestes” curtus Marsh
from the Lance. The latter does not belong in Cimolestes and has been
placed by the writer in a new genus, Diaphorodon^. Smith Woodward
believed the trigonid to be more elevated in the earlier form. Matthew
stated that “the lower molar figured by Marsh as Cimolestes curtus is
somewhat similar to the posterior molars of Eodelpkis but the metaconid
is more reduced, heel wider and shorter, its marginal cusps less differentiated,
and the postero-internal cusps not so high or backwardly prominent. The
tooth is considerably larger, agreeing more nearly with in size.”
Although it can hardly be doubted that Eodelpkis and Diaphorodon are
distinct genera, if only because of their wide geologic separation (no genus
of dinosaurs is common to Lance and Belly River), the differences, in at
least the posterior molars, are very slight. The difference in elevation
of the trigonid is doubtful, and slight at best. The supposed differences
in the heel are largely or wholly due to the corrosion, in different ways,
lAm. Jour. Sei., (5) XIV, p. 127 (1927).
93259— 3i
32
of both types. The metaconid has nearly the same relative size in both.
In fact the agreement is so striking and the molar type so peculiar that
it seems probable that Diaphorodon and Eodelphis were closely related,
the latter perhaps ancestral to the former.
Eodelphis is clearly a marsupial, as shown by Matthew, and, in common
with most Lance marsupials, its resemblance to the later didelphids
is so striking that the writer prefers to place it in the Didelphiidae, at least
until it is much better known. Within the family, however, it occupies
an isolated position, certainly not ancestral to any known Tertiary or
Recent phylum. Aberrant specialization is seen in the stout jaw, the
three incisors with the median one enlarged, the crowded nature of the
premolars and enlargement of Pa,^ the relative enlargement of the paraconid,
and reduction of the metaconid. It is one of the aberrant members of
the varied Cretaceous didelphid group. More generalized forms are
common in the Lance, but have not yet been found in the Belly River.
iPi larger than Pt is a primitive character seen in most fossil didelphids. but in Eodelphis Pi is onsually large
and heavy, although less so than in Thlaeodon, for instance.
33
FOSSILS FROM HARRISON LAKE AREA,
BRITISH COLUMBIA
By C. H. Crickmay, University of California at Los Angeles
Illustrations
Page
Plates VIII-XXIII. Illustrations of fossils 82-113
Figure 1. Index map showing positions of fossiliferous localities in the
vicinity of Harnson lake, B.C 36
INTRODUCTION
This report contains a brief account of the palaeontological materials
acquired during a geological study of the country adjacent to Harrison
lake, British Columbia; made for the Geological Survey, by the writer,
during the summers of the years 1924 and 1926. The writer had no
assistant in 1924. In 1926 he was ably assisted by James R. Pollock.
The writer is indebted to Mr. J. Forsyth of the Provincial Library at
Victoria, to Mr. C. Brakenridge, City Engineer of Vancouver, and to Mr.
L. A. Agassiz, of Agassiz, for courtesies extended to him and information
freely given during the course of the work. He is especially indebted to
Professor J. P. Smith for invaluable advice freely given on many occasions.
The area under consideration lies in the heart of the southern extremity
of the Coast range of British Columbia. Its southern boundary is Fraser
river. The other boundaries are controlled by the distance possible to
cover in one-day trips from the shores of the lake and river. Harrison
lake lies in a large, fiord-like valley adjoining the Fraser on its north side.
It is 1 to 4 miles wide and 38 miles long; its general trend is south-south-
east. Its mean surface level is 30 feet above mean sea-level. It is fed at
its north end by the turbulent waters of Lillooet river, and drained from its
west side near the south end by the slow-moving Harrison river. Lillooet
river and Harrison lake occupy one large valley which joins the Fraser
valley at right angles. South of the lake the drainage is obstructed by
moraines, so that the lake is drained through a narrow side valley which
has been so modified that it now carries the entire overflow of the lake.
Numerous streams, of the size referred to in the west as “creeks,’^ enter the
lake along its shores, at steep gradients.
The topography is entirely mountainous except for the small areas of
flat land forming the bottoms of the larger valleys. In the southern half
of the district the relief averages 4,000 feet (maximum 5,100), and though
the slopes are steep the peaks are rounded. In the northern, the relief is
about 5,500 feet (maximum 7,500); the slopes are steep; the peaks, sharp;
mountain glaciers abound. A thick forest composed mainly of Douglas
fir, western hemlock, and western red cedar covers all the land, except
excessively steep slopes and “cleared” areas, up to an altitude of 4,000
feet. Above this the trees are much dwarfed, and few survive above 5,000
feet.
The shores of Harrison lake are principally low, rocky cliffs. Beaches
form a very small proportion of the whole. The lake is deep, averaging
over 100 fathoms throughout most of its length; however, at the south end
34
it has been silted up so that very slight depths (such as 3 fathoms) are
encountered nearly midway across. The bottom is covered with fine silt
containing some vegetable matter. In late winter and early spring the
water is clear; during the rest of the year it is turbid, though never so foul
as to be unfit to drink. The surface level has a maximum range of variation
of about 7 feet. It is lowest in late winter and early spring, highest about
midsummer, falls all summer, attaining a second minimum about the end
of August, but rises to a second though inferior maximum in the autumn.
The first minimum is due to the fact that at this time of the year precipi-
tation takes the form of snow and so does not reach the lake; the first
maximum is caused by the sudden melting of this snow in the early sum-
mer. The second minimum is due to depletion of the supply of snow
water and lack of rain at this season, and the second maximum is the result
of autumn rains.
The southernmost part of the district is traversed by the Canadian
Pacific railway. A good road provides communication with Harrison
Hotsprings at the south end of Harrison lake. Communication up and
down the lake is effected entirely by boats.
Very little was known of the geology of this district at the time the
investigation was commenced. No areal mapping had been done except
around Agassiz, where the geology was known only in a very general way.
The first reference to the geology of the district was made in the Director’s
Report,^ Geological Survey, Canada, in 1888. It stated simply that the
older Cretaceous rocks were extensively developed in Harrison Lake dis-
trict. In the following year proof of this statement appeared, in the
announcement that Amos Bowman had in 1882 collected Aucella mosquensis
var. concentrica Fischer from several localities on Harrison lake.^ In the
sam*e year Dawson® listed the localities of the hot springs — St. Alice’ well at
the south end of the lake, and St. Agnes’ well 5 miles beyond the north end.
The following year Hoffmann^ gave analyses of the waters from the two
springs, which compose St. Alice’ well. In 1895 Stanton® reported that the
Ancellae from Harrison lake belonged to the species A. crassicollis Key-
serling.
Since these early contributions Harrison lake has scarcely been men-
tioned in the literature. There is no information of any importance except
Bowen’s reports on his reconnaissance of 1912 in which he barely touched
the south end of Harrison lake, but provided a route map of the country
about Agassiz.® Bow’en discerned a fossiliferous sedimentary series, of
which he gives a section, underlying the country around Agassiz. In his
“Columnar Sections”^ he called this the Agassiz series of Palaeozoic age,
but in his summ.ary report® he used no name for it, but said it was of Lower
Cretaceous or Jurassic age. He noticed that these rocks are intruded by a
iSelwyn, A. R. C.: Geol. Surv., Canada, Ann. Kept., vol. Ill (1888).
*Whiteave8, J. F.: Geol. Surv., Canada, Cont. to Can. Pal., vol. I, pt. 2 , No. 4 (1889).
•Dawson, G. M.: “The Mineral Wealth of British Columbia"; Geol. Surv., Canada, Ann. Rept., vol. Ill (1889).
‘Hoffmann, G. C.: "Chemical Contributio.ns to the Geology of Canada"; Geol. Surv., Canada, Ann. Rept.,
vol. IV (1890).
•Stanton, T. W.: "The Fauna of the Knoxville Beds"; U.S. Geol. Surv., Bull. 133 (1895).
•Routo-map between Lytton and Agassiz; Route-map between Agassiz and Vancouver; Geol. Surv., Canada,
Guide Book No. 8, pt. II (1913).
'Columnar Sections. N. L. Bowen, In the Coast Range. Camscll, C.: Geol. Surv., Canada, Guide Book
No. 8, pt. II (1913).
•Bowen, N. L.: "A Geological Reconnaissance of the Fraser River Valley from Lytton to Vancouver, Britiah
Ckdumbia”; Geol. Surv., Canada, Bum. Rept. 1912 (1914).
35
large mass of granite; he mentions the quartz-porphyry on the railway
west of Agassiz, which he regarded as of Lower Cretaceous (?) age, and
with an unconformable relation toward the Agassiz series. The age of
the Agassiz series is Upper Jurassic. The quartz porphyry flows, mentioned
by Bowen, are part of the Porphyrite series, and are of Middle Jurassic age.
Harrison Lake valley has been produced by erosion of disordered
sedimentary series of great thickness accompanied by several masses of
plu tonic rocks. The long axis of the lake is located mainly on what is
believed to be the site of a great overthrust. This fault dips steeply to the
east; Carboniferous and Triassic rocks to the east of it are thrust over
Jurassic and Cretaceous rocks to the west. This great fault, which extends
beyond the limits of the area, is single throughout most of its length, but
in the middle part of Harrison lake it is divided into several subparallel
and oblique faults between which are wedges of rock of the nature of
schuppen. These are well illustrated by the structure of the peninsula,
and that of Long island. In these two areas the surface is composed of
narrow selvages of Jurassic and Cretaceous sediments, bearing their char-
acteristic fossils, and all dipping to the east yet alternating with one
another in a perfectly bewildering fashion. Such an arrangement could
not possibly be the result of peculiarities of deposition, and could have been
formed only by a succession of thrust faults. Throughout most of the
zone the rocks have been considerably crushed.
The oldest rocks lying to the east of the thrust fault are dark grey
argillites with massive limestone lenses bearing Pennsylvanian fossils.
These crop out over the west flank of Bear mountain, at the south end of
the area. They dip steeply to the east, being bounded on the west by the
thrust fault, and to the east by an overlying thick accumulation of sedi-
ments, mostly argillites but with some beds of pyroclastics. This series
borders the thrust fault on the east from Trout Lake creek to Fifteenmile
creek and makes up all the mountains between these creeks. The sedi-
ments dip uniformly steeply to the east (or a few degrees north of east).
They are considerably crushed, in some places even to the state of foliation.
So far they have yielded no fossils, but it is suspected that they are of
Triassic age. The total thickness has not been determined, but is esti-
mated at 14,000 or 15,000 feet.
Overlying the supposedly Triassic series and, therefore, succeeding
them to the north and east, is a thick formation of crushed greenstones of
volcanic origin. This formation occupies most of the country east of
Harrison lake. These rocks have been considerably crushed and are more
or less schistose throughout, but apart from this there is a great resemblance
as regards thicknesses and details of lithology and succession between these
rocks and Jurassic rocks lying to the west of Harrison lake, but no fossils
have been obtained from the greenstones lying east of the lake.
The oldest rocks west of the great thrust occur in a small patch on the
west shore of the lake immediately south of Camp cove. They consist of
some 3,000 feet of sediments, mostly argillites with some tuffs and a con-
glomerate bed. The conglomerate contains besides other things, pebbles
of the fossiliferous Pennsylvanian limestones. The series has yielded no
fossils of its own date; it belongs betw'een the top of the Pennsylvanian and
the base of the Middle Jurassic. These rocks lie in the centre of a short
36
Figure 1. Index map shomng positions of fossiliferous localities in the vicinity of Harrison
lake, sourfiwest British Columbia. Numbers correspond with those used in text.
37
anticline, and are surrounded by a thick accumulation of tuffs, agglomer-
ates, and flows, which cover a very considerable area west of Harrison lake
and occupy most of Echo island. The total thickness is about 9,000 feet.
Fossils were obtained from a band of aqueous sediments in the lower part
of the formation (locality No. 3, See Figure 1) and also from higher hori-
zons. These show the volcanic assemblage to be of Middle Jurassic age.
To the north, near Deer creek, to the east on Echo island, and to the south
on Harrison river, the volcanics are overlain conformably by well-stratified
tuffs. These have an average thickness of 2,700 feet. Only a few fossils were
found in them and none of these is indicative of exact dates. However, the
formation is overlain conformably by dark grey argillites, the upper part of
which yielded excellent fossils of early Upper Jurassic age. This deposit of
argillite, which is about 2,400 feet thick, is well exposed from the mouth of
Deer creek across the hills to the northwest, through the valley of Myster-
ious creek, and into the Chehalis Creek basin; also in another strip on the
south side of Harrison river. In both sections the argillites are overlain
by an accumulation of well-stratified tuffs that on Mysterious creek are
about 1,800 feet thick. They have yielded early Upper Jurassic fossils.
The early Upper Jurassic argillites and tuffs on the south side of Har-
rison river are overlain unconformably by the rocks to which Bowen applied
the name Agassiz series. The lower part of this series consists of about
3.000 feet of unfossiliferous conglomerates. These are overlain by about
5.000 feet of black argillites which contain poorly preserved, unidentifiable
fossils. At this point the sequence is broken by the thrust faults. How-
ever, presumably the same formation occurs as narrow strips on the
west shore of the peninsula where it yields the Argovian ammonite. Ana-
cardiocerds.
The argillites that on the peninsula are regarded as part of the Agassiz
series and the stratified Upper Jurassic tuffs near Deer creek, are overlain
unconformably by early Cretaceous deposits bearing abundant fossils,
mostly of the genus Aucella. The lower part of the Cretaceous series
consists of a basal conglomerate overlain by grey sandstones and totalling
1,260 feet in thickness. The upper part is made up of 2,200 feet of pyro-
clastics overlain by about 1,500 feet of sandstones composed of waste
from the volcanics. These Cretaceous beds occur in a strip running north-
west from the peninsula, along the west side of Long island, to Broken-
back hill and beyond. This strip is cut off along its east side by the thrust
fault.
The sedimentary rocks of the district are cut by plutonics thought to
be of two ages. Those of one age are mostly altered quartz monzonite
and occur in masses of various sizes, the largest of which is a batholith
lying west of Chehalis creek. It intrudes the Middle Jurassic rocks and
appears to have supplied pebbles to the basal conglomerate of the early
Cretaceous. It is placed tentatively as late Jurassic.
The other plutonic intrusive is a fresh quartz diorite. It occurs in
stocks in various parts of the area, notably between Agassiz and Harrison
Hotsprings, and at Doctor point, also in a batholith lying mostly in the
upper part of the Chehalis drainage basin. These quartz diorites cut
both Jurassic and Cretaceous strata and intersect the great overthrusts
38
and the high-angled structures of the district. They are, therefore, sub-
sequent to the main orogeny of the region which is thought to be Lara-
mide.
A system of dykes of basalt cuts all the other rocks of the district.
These are thought to be of fairly late date.
FOSSIL LOCALITIES AND FAUNAL LISTS
The localities, and the fossils found there, are here listed according to
the known or presumed ages of the strata. Individual localities are dis-
tinguished by numbers and these numbers on the accompanying index
map (Figure 1) indicate the positions of the various fossil-bearing locali-
ties.
PENNSYLVANIAN
(1) West side of Bear mountain, at an elevation of 2,000 feet, Ij miles
due east of the Harrison Hotsprings hotel.
Large crinoid stalks — Melocrinidae?
Small crinoid stalks — Batocrinidae?
(2) West shore of Harrison lake, 2j miles from Harrison River point.
Conglomerates. The fossils occur in pebbles of limestone derived from a
Pennsylvanian formation.
Producliis cf. clarkei Ischemychew
Crinoidea, several species
JURASSIC
(3) West shore of Harrison lake, 1,820 yards north of Harrison River
point.
Rhynchonella sp.
Enlolium volcanicum Crickmay
Cylindroteuthis themis Crickmay
(4) East shore of Echo island, 1,860 yards north of the southeast
comer.
BeUmnites sp.
(5) South end of the first small islet southeast of Echo island.
Belemnites sp.
Pelecypoda, various unidentifiable forms.
(6) Northeast side of the second small islet southeast of Echo island.
Belemnites sp.
(7) West shore of Harrison lake, 600 yards south of the mouth of
Deer creek.
Inoceramus ? sp.
(8) South shore of Harrison river on the west side of a small point
150 yards east of V. Macdonald's house, or 2\ miles from Harrison River
point.
LiUoettia sp. — crushed specimens of slightly earlier date than the described species
39
(9) Deer creek at 1,450 feet altitude, If miles from the mouth.
Lilloettia sp.
(10) Deer creek at 1,275 feet altitude, 1| miles from the mouth.
Lilloeilia sp.
(11) Deer creek at 300 feet altitude, 700 yards from the mouth.
Pai acadoceras sp.
(12) Deer creek, f mile from the mouth.
Paracadoceras haroeyi Crickmay
Cadoceras catostoma Pompeckj
“ schmidti “
“ brooksi Crickmay
“Belemnites'^ sp.
(13) On the small tributary which joins Deer creek from the north
at 1,700 feet elevation or 2| miles from the mouth.
Parallelodon sp.
Entolium hertleini Crickmay
Ammonoid fragment, cadoceratoid
“Belemnites" sp. indet.
(14) South side of Mysterious Creek valley 2| miles from the mouth
and at 1,400 feet elevation.
Parnllelodon sp.
“Belemnites” sp.
(15) Billhook creek, a tributary to Mysterious creek, at 3,100 feet
altitude, and 4 miles in a direct line from the mouth of Mysterious creek.
Callovian (Proplanulitan)
Anomia columbiana Crickmay
Lilloettia lilloetensis Crickmay
“ mertonyarwoodi “
Buckmaniceras buckmani “
(16) One hundred yards from shore and 600 yards north of the mouth
of Deer creek.
*‘Belemnites’' sp.
(17) West shore of Harrison lake, 720 yards north of the mouth of
Deer creek.
Pelecypoda, various, imidentifiable
(18) Southwest slope of Fossil hill at 1,850 feet.
Haidaia aflf. dawsoni Whiteaves
^‘Belemnites” sp.
(19) Small creek 1 mile east of Billhook creek at 1,500 feet.
Astarle harrismierms Crickmay
(in talus)
(20) Billhook creek at 3,270 feet elevation and slightly over 4 miles
from the mouth of Mysterious creek.
Cylindroteuthis sp.
(21) Billhook creek at 3,400 feet elevation.
Haidaia aff. dawsoni Whiteaves
“ billhookensis Crickmay (talus)
“ packardi “ (talus)
Ammonoid, a young cadoceratoid form
40
(22) Billhook creek at 3,600 feet elevation.
Pelecypoda, various, unidentifiable
Haidaia stcUltiensis Crickmay
Ammonoid, a young cadoceratoid fonn
“Belemnites" sp.
(23) Southernmost tip of the more westerly of the two small hills
lying west of Agassiz.
Ammonoid, indeterminable
(24) Southwest side of a narrow point which bounds on the west the
small bay on the southwest shore of the peninsula, and 1,850 yards from
the southeast point of the peninsula.
Parallelodon cardiocercUanum Crickmay
Anacardioceras perrini Crickmay
Phylloceras columbianum Crickmay
(25) East shore of Long island, miles from its south end.
Ammonoid, a young cadoceratoid form
“Belemnites” sp.
(26) A little islet off the middle of the three points at the north end
of the peninsula.
‘‘Belemnites” sp.
(27) At 250 feet altitude on hillside half a mile south-southwest from
Doctor point.
Ammonoid, indeterminable
CRETACEOUS
(28) West shore of a little bay on the southwest shore of the peninsula.
AuceUa acutistriata Crickmay
catamorpha
spasskensoides
cascadensis
canadiana
Eniolium auceUarum
ti
U
(29) On the hillside one-quarter mile due north of the last locality.
Aitcella catamorpha Crickmay
spasskensoides “
sp.
4 <
u
(30) One-third mile south of the highest peak on the peninsula.
Aucella spasskensoides Crickmay
“ canadiana “
** sp.
(31) West shore of peninsula 3| miles in a straight line from its
southern tip.
Aucella acutistriata Crickmay
catamorpha **
spasskensoides “
canadiana
‘^Belemnites" sp.
ii
€i
a
u
41
(32) West shore of Harrison lake, 900 yards north of the mouth of
Deer creek.
Aucella acutistriata Crickmay
“ spasskemoidea “
“ canadiana **
CylindroteiUhu bacidm Crickmay
PachyleiUhis eocretacicua “
(33) Three hundred and fifty yards west of the shore of the lake and
1,200 yards north of the mouth of Deer creek.
AuceUa acutistriata Crickmay
“ catamorpha “
“ spasskensoides “
“ cascadensis "
" canadiana “
U
U
u
(34) Three hundred and fifty yards from shore and 1,400 yards
north of the mouth of Deer creek.
AuceUa acutistriata Crickmay
“ spasskensoides
“ cascadensis
Entolium aucellarum
Mcleamia mcleami
A starts Barbara
Phylloceras aff. knoxvillense Stanton
“ sp. indet.
Cylindroteuthis hacuhis Crickmay
Pachyteuthis eocretacicus Crickmay
(35) One and two-fifths miles northwest of the mouth of Deer creek,
at an elevation of 1,000 feet.
Aucella acutistriata Crickmay
caiamorpfia “
canadiana “
spasskensoides **
U
€4
U
(36) Right bank of Mysterious creek at 1,000 feet altitude.
Aucella acutistriata Crickmay
** spasskensoides “
(37) North slope of Fossil hill, 250 yards southwest of a point on
Mysterious creek of 1,000 feet altitude.
Aucella acutistriata Crickmay
“ spasskensoides '■
(38) Two hundred and fifty yards west of the last locality.
AuMlla acutistriata Crickmay
spasskensoides “
i(
(39) Two hundred and thirty-five yards from the shore, on the left
bank of a little brook that enters Harrison lake from the west, 1,500
yards north of the mouth of Deer creek.
Aucella kwoiekensis Crickmay
Quoiecchia aliciae
(40) Lonetree island, off south tip of peninsula.
Aucella crassicollis Keyserling
Aucella solida Lahusen
42
(41) East shore of peninsula, 200 yards north of Lonetree island.
Aucella crassicollis Keyserling
“ solida Lahusen
“ harrisonensis Crickmay
“ gigas “
Plexiromya harrisonensis “
Homolsomites poecilochotomus Crickmay
(42) East shore of peninsula 400 yards north of Lonetree island.
Aucella crassicollis Keyserling
“ solida Lahusen
“ harrisonensis Crickmay
Pleuromya harrisonensis “
Homolsomites poedlochotcrmus Crickmay
(43) Half a mile southeast of the highest ‘peak on the peninsula.
Pelecypod, imidentifiable
(44) East shore of peninsula, 2^ miles north of Lonetree island.
Aucella crassicollis Keyserling
(45) West shore of Long island 400 yards north of its southern tip.
Aucella crassicollis Keyserling
“ solida Lahusen
(46) Little islet on south side of south entrance to Roberts bay, west
side of Long island.
Aucella crassicollis Keyserling
solida Lahusen
(47) South shore of a little point on the west side of Cascade bay,
1,090 yards south of the north end of that bay; or 4,660 yards north of
Lonetree island.
Aucella teutoburgensis Weerth
Yaadia leurisagassizi Crickmay
(48) Forty feet east of the Yaadia lewisagassizi locality on south
shore of a little point on west side of Cascade bay 1,090 yards due south
of the north end of that bay.
Aucella somewhat crushed and not identifiable with certainty, but probably the
sharp ribbed species of the canadiana zone
(49) Northeast side of the little point on west side of Cascade bay,
1,090 yards due south of the north end of that bay.
Aucella somewhat sheared and so unidentifiable, but probably the sharp ribbed
species of the canadiana zone
(50) East side of Cascade bay 2f miles from the north end of that
bay, also
(51) East side of Cascade bay 3| miles from the north end of that
bay, also
(52) East side of Cascade bay 4| miles from the north end of that
bay.
Aucellae^ greatly sheared and so unidentifiable, but probably the sharp ribbed
species of the canadiana zone
(53) Northwest corner of Roberts bay, west side of Long island.
Aucellae, somewhat crushed and so unidentifiable, but probably the sharp ribbed
species of the canadiana zone
43
(54) East side of Roberts bay 650 yards from the north end, also
(55) Islet, 100 yards west of the last locality, also
(56) West shore of Long island, 2f miles south of the north end of
that island, also
(57) East shore of Long island, 3| miles in a straight line from the
north end of that island.
Aucelhs, QiTon^y sheared and so unidentifiable, but probably the sharp ribbed
species of the canadiana zone
(58) West side of Long island, barely 2 miles south of the north end
of that island.
Aucellae, considerably crushed and so unidentifiable with certainty, but never-
theless very probably Ancella crassicollis Keyserling
(59) North side of Twentymile point, one-quarter mile from the
point, also
(60) West shore of Harrison lake, one-half mile south of Rock point.
Pelecypoda, unidentifiable fragments
"Belemnites” sp. indet.
(61) Cut bank on south side of main line of Canadian Pacific railway
838 yards west of the west end of the Harrison River bridge.
Strongylocenlrolus drobachiensis Muller
M ytilus edulis Linnaeus
Terebratalia tramversa Sowerby?
Balanus crenatus Bruguiere
(62) Cut bank on south side of main line of Canadian Pacific railway
750 yards west of the west end of the Harrison River bridge.
Saxicava rugosa Linnaeus
Other pelecypoda, indeterminable fragments
DESCRIPTION OF FOSSILS
Phylum, ECHINODERMATA
Class, CRINOIDEA
Order, Camerata
Family, mblocrinidae?
Genus, (?)
Fragments of large stalks, 10 to 27 mm. in diameter, the joints of
which exhibit the structure and intercolumnal spaces of those of this
family. They are imperfectly preserved and do not admit of complete
study.
Locality. West side of Bear mountain at 2,000 feet elevation, 1^
miles due east of Harrison Hotsprings hotel. Similar crinoid stalks are
common at the type locality of the Chilliwack series on Chilliwack river,
also in the Pennsylvanian rocks of Kamloops district, British Columbia.
44
Family, batocrinidae?
Fragments of smaller stalks which closely resemble in a superficial
way those of this family. The reference is, of course, far from certain.
There are probably several forms: the shapes of the small columnals differ
somewhat. Diameters from 3 to 8 mm.
Locality. West side of Bear mountain at 2,000 feet, IJ miles due east
of the Harrison Hotsprings hotel.
Class, ECHINOIDEA
Order, Centrechinoidea
Family, strongylocentrotidae
Genus, Strongylocentrotus Brandt
Strongylocentrotus drobachiensis Mtiller
Fragments of corona and spines belonging to this species.
Locality. Main line of Canadian Pacific railway 838 yards west of
the west end of Harrison River bridge.
Phylum, MOLLUSCOIDEA
Class, BRACHIOPODA
Order, Protremata
Family, productidae
Genus, Productus
Productus cf. darkei Tschernychew
Specimens comparable with this species from limestone pebbles in
conglomerate.
Locality. West shore of Harrison lake 2J miles from Harrison River
point.
Order, Telotremata
Family, ehynchonellidae
Genus, Rhynchonella
Rhynchonella sp.
Indeterminable fragments belonging to this genus.
Locality. West shore of Harrison lake 1,820 yards from Harrison
River point.
45
Family, tebebbatellidae
Genus, Terebratalia
Terehraialia transversa Sowerby?
Small, decrepitated fragments that have the structure of this species.
Occurs with Mytilus edulis and Strongylocentrotus drobachiensis.
Locality, On main line of Canadian Pacific railway 838 yards west
of the west end of Harrison River bridge.
Phylum, MOLLUSCA
Order, Prionodesmacea
Family, paballelodontidae
Genus, Parallelodon Meek
Parallelodon cardioceraianum sp. nov.
Plate VIII, figures 1, 2, 3
Surface marked only by excessively delicate, concentric lines.
Dimensions. Holotype, a left valve — length, 22 mm.; height, 14;
semidiameter (diameter of one valve, or distance from lateral convexity
to median plane), 5.
Locality. Southwest shore of peninsula, on west side of point that
bounds the small bay.
Family, pebnidae
Genus, Inoceramus Sowerby
Indeterminable fragments belonging to this genus.
Locality. West shore of Harrison lake, 600 yards south of Deer creek.
Family, ptebiidae
Genus, Mclearnia gen. nov.
A giant pteriid without sculpture. Right valve flattish with pec-
tiniform byssal notch. Left valve, somewhat more convex. Ears, sub-
equal, all well developed. Chondrophore, small, central, and vertical.
Mclearnia mclearni sp. nov.
Plate VIII, figure 4; Plate IX, figure 1
Umbo lies at centre of hinge-line, somewhat posterior to centre of
disk, giving the shell a peculiar outline. Shell is much thickened in the
umbonal region. Surface smooth except for occasional growth-lines.
When split the shell shows traces of fine, radial ribbing. Interior smooth
except for the single, large, bilobate muscle-scar; and two curved lines of
*‘diraples", one on each side of the umbo. These appear in the figures,
as does also the central chondrophore.
9325ft-4
46
Dimensions. Holotype, a right valve seen from interior aspect —
length, 115 mm.; height, 112; semidiameter, 11; hinge-line, 71. Paratype,
a left valve, cast of interior — height, 118; semidiameter, 25; hinge-line, 78.
Locality. Zone of aucella canadiana, west side of Harrison lake 350
yards from shore and 1,400 yards north of Deer creek.
Name. Generic and specific name after F. H. McLearn, palaeontolo-
gist.
Family, myalinidae
Genus, Aucella Keyserling
The species of Aucella from Harrison lake form a considerable and
an important assemblage belonging to four separate hemerse. This was
obvious at the beginning of this study, but at that time it was thought
that the earliest Harrison forms corresponded to the earliest California
Cretaceous species — Aucella piochii, and, in consequence, some mistaken
identifications were made. Subsequently, larger collections and a more
exhaustive study have shown that the species of the A. canadiana zone
are later than A. piochii. Moreover, they are not related to that species
but rather to the group of A. hononiensis. They seem to represent large
shelled derivatives of this group evolved during the great expan five evolu-
tion of Aucellae which occurred at the end of the Jurassic period. They
represent one of the earliest Cretaceous hemerffi, probably that of Cras~
pedites stenomphalus. The other Harrison Aucellae, except A. kwoiekensis,
belong to the period of restrictive and degenerative evolution of Polypty-
ehites polyptychus and later time.
Aucella acutistriata sp. nov.
Plate IX, figure 2
Resembles A. andersoni somewhat but differs in its much larger size
and greater diameter. The ornament consists of concentric costae that
are as sharp as knife-blades and stand 0-5 to 1 mm. above the general
surface. They are spaced 8 or 9 in a cm. of radius. Become obsolete
on anterior and posterior ends of shell which are striate concentrically.
Dimensions. Holotype, a left valve — long axis, 78 mm,; short axis,
that is the greatest line at right angles to the long axis, 42; semidiameter,
28; apical angle, 69 degrees. Paratype, both valves in contact: left^ — long
axis, 42; short axis, 25; semidiameter, 13; apical angle, 65 degrees; right-
long axis, 39; short axis, 25; semidiameter, 11; apical angle, 80 degrees.
Locality. Zone of Aucella canadiana, west side of Harrison lake, 350
yards from shore and 1,200 yards north of Deer creek.
Aucella catamorpha sp. nov.
Plate IX, figure 5
Very similar to the last except that the shell is unornamented. It is
marked only by growth lines. Also the right valve has a more acute
apical angle, and the diameter is slightly less.
47
Dimensions. Holotype, right valve — long axis, 37 mm.; short axis,
25; semidiameter, 7; apical angle, 60 degrees.
Locality. Zone of Aucella canadiana, west side of a small bay on
southwest shore of the peninsula.
Aucella spasskensoides
Plate IX, figures 3, 4
Resembles A. spasskensis Pavlow, but the diameter is notably greater
and the right valve has a greater apical angle. Strong concentric costae,
sharp as in .4. acutistriata, spaced 4 to 5 in a cm. of radius on the mature
shell, though closer in the umbonal region.
Dimensions. Holotype, right valve — long axis, 38 mm.; short axis,
29; semidiameter, . . ; apical angle, 80 degrees; right valve — long axis, 36;
short axis, 29 ; semidiameter, 7 ; apical angle, 90 degrees. Paratype, left
valve — long axis, 55; short axis, 39; semidiameter, 16; apical angle, 67
degrees. The largest specimen found, a left valve, had a long axis measuring
60 mm.
Locality. Zone of Aucella canadiana, west side of small bay on south-
west shore of peninsula.
Aucella cascadensis sp. nov.
Plate X, figures 1, 2
Having the same general aspect of the fischeriana stock which the
last species shows. The costation is wider spaced; 3 coatse in a centimetre
of radius on the mature shell. Also the costse are sharp like those of
A. acutistriata.
Dimensions. Holotype, left valve — long axis, 57 mm.; short axis, 43;
semidiameter, 15; apical angle, 80 degrees. Paratype, right valve — long
axis, 63; short axis, 54; semidiameter, 15; apical angle, 105 degrees.
Locality. Zone of A. canadiana, west side of Harrison lake, 350 yards
from shore and 1,200 yards north of Deer creek.
Name. Cascade mountains.
Aucella canadiana sp. nov.
Plate X, figures 3, 4, 5
This form has departed farther from the fischeriana ancestry than the
preceding species. These costas are wide spaced: 1-5 to each centimetre
of radius, and are sharp like those of A. acutistriata.
Dimensions. Holotype, a left valve — long axis, 54 mm.; short axis,
43; semidiameter, 22; apical angle, 75 degrees. Paratype, right valve —
long axis, 60; short axis, 49; semidiameter, 12; apical angle, 93 degrees.
Locality. Zone of Aucella canadiana, west side of Harrison lake 350
yards from shore and 1,200 yards north of Deer creek.
Name. A Canadian species, and peculiarly so, because it is strikingly
different from any of the foreign forms.
9325fr-4i
48
Aucella kwoiekensis sp. nov.
Plate X, figures 6, 7
Resembles members of the inflata group, but differs markedly from all
in proportions and in the fine, closely spaced, concentric striae. Byssal
sinus unusually long.
Although Pavlow suggests a very different history, yet it is quite
possible that this form is ancestral to the crassicollis group.
Dimensions. Holotype, a right valve — long axis, 42 mm.; short axis,
38; semidiameter, 15; apical angle, 90 degrees.
Locality. Zone of Quoiecchia aliciae, 235 yards from shore, on the
left bank of a little brook which enters Harrison lake from the west, 1,450
yards north of the mouth of Deer creek.
Name. Kwoiek, Indian geographical name.
Aucella crassicollis Keyserling 1846
1846. Keyserling: Wissenschaftliche. Beobachtungen auf einer Reise
in dar Petschoraland.
The left valve is very strongly convex, and the left umbo is strongly
incurved. Right valve, less so. The long axis of the left valve is about
twice the short axis. The form is rather irregular. The external surface
is marked by fine growth lines and concentric varices. The latter appear
with maturity, but at slightly different stages in different specimens.
Locality. Zone of Homolsomites poecilochotomus, Lonetree island;
also on the southeastern shore of the peninsula, etc.
Aucella solida Lahusen
Aucella crassicollis var. solida Lahusen 1888: Mem. du Comite g^ol.,
St. Petersburg, vol. VIII, No. 1.
This species somewhat resembles the last, but differs in that the long
axis is not much greater than the short.
Locality. Occurs associated with the last at all localities.
Aucella harrisonensis sp. nov.
Plate XI, figures 1, 2, 3
Left valve, somewhat convex. Left umbo, slightly incurved. Right
valve, flattish. Right umbo, short, obtuse, and almost ecurved. Shell,
very thin, and, consequently, there is no ornament except joint growth
lines. Concentric varices appear beyond 25 mm. from the umbo in some
specimens.
Dimensions. Holotype, a right valve — long axis, 32 mm.; short
axis, 30; semidiameter, 8; apical angle, 95 degrees. Paratype, a left valve —
long axis, 30 mm.; short axis, 20; semidiameter, 9; apical angle, 73 degrees.
Locality. Zone of Homolsomites poecilochotomus, southeast shore of
the peninsula, 200 yards from the south end.
49
Aucella gigas sp. nov.
Plate XI, figures 4, 5, 6
Valves resemble each other strongly in shape. Both are broad,
flattish, and rounded ventrally. Umbones, short, obtuse, very slightly
incurved. Ornament of obtuse, concentric costse that have a very char-
acteristic curve. Ornament degenerates beyond 35 mm. from the umbo.
Dimensions. Holotype, left valve — long axis, 100 mm.; short axis,
78; semidiameter, 20; apical angle, 80 degrees; right valve — long axis, 83
mm,; short axis, 72; semidiameter, 16; apical angle, 90 degrees.
Locality. Zone of Homolsomites poecilochotomus, southeast shore of
the peninsula 200 yards from the south end.
Name. From the fact that it is the largest Aucella known.
Aucella teutoburgensis Weerth
Avicula (?) teutoburgensis Weerth 1884: Palaeont. abhandl. Bd. II.
Both valves notably convex, left valve somewhat more so than right.
Umbones incurved. The outline is characteristic. Ornament, striate.
Locality. Zone of Yaadia lewisagassizi, on south shore of a little point
on the west side of Cascade bay 1,090 yards south of the north end of that
bay.
Family, trigoniidae
Genus, Haidaia Crickmay
Of the Harrison Lake species referred to this genus only one is typical.
This form, related to H. dawsoni Whiteaves, is unluckily not represented
by material good enough for specific description. The other three are
new species which are referred to this genus for convenience, as it seems
undesirable to make any further division until more forms are known.
Haidaia sp. nov. aff. H. dawsoni Whiteaves
Fragments of a new form, similar to dawsoni in the main but larger,
longer in proportion to height, and having a narrower area.
Locality. Southwest slope of Fossil hill at 1,850 feet; and on Billhook
creek at 3,400 feet.
Haidaia billhookensis sp. nov.
Plate XII, figure 1
Costae, numerous for a member of this genus. Tubercles, numerous,
arranged in concentric series, degenerating toward basal margin. Costellae
become irregular at maturity, rather resembling coarse growth lines.
Marginal carina at maturity is a line of weak, irregularly spaced bullae.
Median carina, lacking. Inner carina, not seen. Median furrow, shallow.
Dimensions. Length, 77 mm.; height, 70; semidiameter, 25.
Locality. Talus at 3,400 feet on Billhook creek.
Name. After the locality.
50
Haidaia packardi sp. nov.
Plate XII, figures 2, 3
Radial costsD, strong. Tubercles, strongest on a concentric zone
midway between umbo and base; that is, degenerating before maturity —
a sign of catamorphism. Tubercles are arranged in concentric series and
are joined by weak, concentric costae. Costellae, very fine, degenerating
with maturity to coarse, irregular striae resembling rugose growth lines.
Inner and marginal carinae strong, coarsely tuberculate. Median carina,
absent. Furrow, weak. Posterior basal margin has three or four crenu-
lations on the inside.
Dimensions. Length, 70 mm.; height, 58; semidiameter, 20.
Locality. Talus at 3,400 feet on Billhook creek.
Name. After E. L. Packard, in recognition of his work on west
American Trigoniae.
Haidaia statluensis sp. nov.
Plate XII, figures 4, 5, 6
Costae weak, though marked by strong, obtuse tubercles that are
arranged in concentric series and joined by concentric costae. Two figures
of specially prepared artificial casts are introduced to show this peculiarity,
which does not appear very plainly on the holotype. Both costae and
tubercles are small and weak on the umbonal region and anterior end.
Area marked by faint, transverse striae which become with maturity coarse
costellae. Carinae not specially marked. Furrow, shallow, becoming
obsolete with maturity.
Dimensions. Length, 65 mm.; height, 54; semidiameter, 15.
Locality. Billhook creek at 3,600 feet.
Name. Geographical — Statlu creek.
Genus, Yaadia gen. nov.
Genotype, Yaadia lewisagassizi sp. nov.
This genus is instituted for a branch of the pseudoquadrate Trigoniae
that has a discrepant ornament on the anterior end of the disk, and so can
not well be included in the true pseudoquadrates, genus Steinmanella nov.
genotype of which is S. holuhi Kitchin. The discrepant anterior ornament
of Yaadia is not necessarily any indication of relationship with Scaphi-
trigon, because this kind of thing might well be developed independently.
Steinmanella is an Indo-Pacific genus of the early Cretaceous. Yaadia
is presumably a north Pacific group of the same time.
Name. From Yaada, Indian legendary name.
Yaadia lewisagassizi sp. nov.
Plate XIII, figures 1, 2
Near the umbo, the two sets of ornament are united into one resemb-
ling that of Trigonia s.s. The two sets of ornament on the disk are separ-
ated by a perfectly smooth area. Carinae marked only by rows of coarse
51
tubercles (bullse), which are widely spaced and have a peculiar oblique
elongation. Area, otherwise smooth. Escutcheon, marked by transverse
lines of small tubercles continuous with tubercles of inner carina.
Dimensions. Length, 77 mm.; height, 63; semidiameter, 22. This
length and height may be slightly vitiated by diastrophic distortion.
Locality. Zone of Yaadia lewisagassizi, on the south shore of a little
point on the west side of Cascade bay, 1,090 yards south of the north end
of that bay. Occurs with Aucella teutoburgensis.
Name. In honour of Captain Lewis N. Agassiz, Royal Welsh Fusiliers,
one of the pioneers of British Columbia.
Genus, Quoiecchia gen. nov.
Genotype, Quoiecchia aliciae sp. nov.
The genus is distinguished by the small-sized shell and the peculiar
arrangement of the ornament. It is unlike any other group of Trigoniae,
though there is a suggestion of this style of ornament in Haidaia statluensis.
Name. From Kwoiek, Indian geographical name.
Quoiecchia aliciae sp. nov.
Plate XIII, figures 3-8
According to the Agassiz classification this species would be referred
to “les lisses” (glabrae of Lycett), a group of composite origin, being smooth
derivatives of various stocks. The form is probably an aberrant offshoot
of the great, protean Haidaia. The ornament of the disk is formed by
two sets of furrows. One, a radial set, is strong near the umbo but becomes
obsolete near the middle of the disk. The other, a concentric set, appears
first at 10 mm. from the umbo and becomes increasingly stronger toward
the basal margin. The spaces between furrows stand up as low, obtuse
costse; between two intersecting sets of furrows, as low, squarish tubercles.
The carinae are obsolete, the area being ornamented only by the posterior
continuation of the concentric furrows. The interior surface of the pos-
terior half of the basal margin is crenulated, forming four or five broad,
tooth-like processes which interlock in opposite valves when the shell
is closed.
Dimensions. Holotype, a right valve — length, 21 mm.; height, 19;
semidiameter, 6. Paratype, a right valve — length, 30; height, 34; semi-
diameter, 15. This shows that as it grows the shell becomes dispro-
portionately high and robust.
Locality. Zone of Quoiecchia aliciae, 235 yards from lake shore, on
the left bank of a little brook that enters Harrison lake from the west
1,450 yards north of the mouth of Deer creek.
Name. Geographical — ^St. Alice’ well.
52
Family, pectinidab
Genus, Entolium Meek
Entolium hertleini sp. nov.
Plate XIV, figure 1
Shell, very flat. Ears, small. Upper margins of the ears make a
shallow re-entrant. External surface appears smooth, but the X 10 power
of magnification shows it to be marked by even, continuous, concentric
striae. These have a characteristic curve which is shown in the plate
by a black line. Lateral margin of disk, also ears, are marked by faint,
microscopic radial striae as well. The species is very similar to the wide
variety of Entolium leachii McLearn, but is separable by its smaller ears,
concentric curve.
Dimensions, Holotype — length, 27 mm.; height, 27; semidiameter, 2;
base of anterior ear, 5*5; base of posterior ear, 7 ; upper margins of ears,
total 6*5; re-entrant angle, 165 degrees; apical angle, 115 degrees. Para-
type — length, 57; height, 56; apical angle, 125 degrees. Both types are
right valves.
Locality, Above zone of Cadoceras brooksi, at an altitude of 2,100 feet
on a small brook which enters Deer creek from the north at 1,700 feet.
Name. After Mr. Leo G. Hertlein.
Entolium vulmnicmn sp. nov.
Plate XIV, figure 2
Shell, very flat, large. Ears, small, with a notable re-entrant above.
Next below the dorsal margin of the shell, and bounded below by a radial
line, there is an area where the shell is thickened. These are plainly
visible in the figure. Exterior, smooth or nearly so. Some specimens
have an appearance of faint, concentric striae.
Dimensions. Holotype, a left valve — length, 68 mm.; height, 62;
semidiameter, 4; base of anterior ear, 12; base of posterior ear, 17; upper
margins of ears, total, 17; re-entrant angle, 140 degrees; apical angle, 128
degrees.
Locality. Associated with Cylindroteuthis ihemis, on the west shore
of Harrison lake, 1,820 yards north of Harrison River point.
Name. In allusion to habitat. The species was an inhabitant of the
powerfully volcanic British Columbia of Bajocian times.
Entolium aucellarum sp. nov.
Plate XIV, figure 3
Shell, flat, smootfi. The ears are larger than those of E. hertleini
and E. vulcanicum, and are of a different shape from those of either, or of
E. leachii McLearn. As is usual with this genus the apical angle increases
with age.
Dimensions. Holotype, a right valve — length, 27 mm.; height, 29;
midiameter, 2; base of anterior ear, 7; base of posterior ear, 8; upper
53
margins of ears, total, 9; re-entrant angle, 160 degrees; apical angle, 113
degrees.
Locality. Zone of Aucella canadiana, west side of a small bay on the
southwest shore of the peninsula.
Name. The species is a syntopite with the Aucellae of the canadiana
zone.
Family, anomiidae
Genus, Anomia Linn^
Anomia columhiana sp. nov.
Plate XIV, figures 4, 5, 6
Differs from A. albertensis McLearn in its larger size and in having
a much finer ornament of radial stri®. The stri® become slightly stronger
as they strike the margin of the shell. The outline of the shell is rather
variable. The shell is marked also by concentric irregularities of surface
comparable with varices.
Dimensions. Holotype — equidimensional, though somewhat crushed;
width, 57 mm. Para type — width, 55; height, 60.
Locality, Zone of Lilloettia lilloetensis, on Billhook creek at 3,100
feet altitude.
Name. After British Columbia.
Family, mytilidae
Genus, Mytilus Linnaeus
Mytilus edulis Linnaeus
Fragments of this common living species, very typical as regards
form and dimensions.
Locality. Main line of Canadian Pacific railway, 838 yards west of
the west end of the Harrison River bridge.
Order, Anomalodesmacea
Family, pleuromyacidae
Genus, Pleuromya Agassiz
Pleuromya harrisonensis sp. nov.
Plate XV, figures 1, 2, 3
Shell, pleuromyiform, small, robust, not perceptibly gaping, having
a shallow sulcus in the antero-lateral area running radially from umbo
to basal margin. Cardinal area obscure. Post-umbonal slope, concave.
Posterior dorsal margin, expanded postero-dorsaily. No cardinal groove.
Maximum diameter is vertically below the umbones. Ornament of con-
centric wrinkles, which become weak on the posterior end of the shell.
They are obscured by matrix on the anterior end of the only specimen.
54
Dimensions. Holotype — length, 42 mm.; height, 35*5; semidiameter,
13 ; horizontal distance from anterior extremity to umbo, 15.
Locality. Zone of Homolsomites poecilochotomuSj on the southeast
shore of the peninsula.
Order, Teleodesmacea
Family, astartidae
Genus, Astarte Sowerby
A starts harrisonensis sp. nov.
Plate XV, figures 4, 5
Lunule and escutcheon, deeply impressed. Ornament of high, con-
centric costae, which are somewhat irregular as to size and continuity.
On a shell of 21 mm. height there are 85 costae in 10 mm. of radius, counted
about the centre of the disk. Superimposed upon this ornament are fine
growth lines visible only with a magnifier. The whole basal margin from
lunule to escutcheon is finely denticulate inside. The species might be
confused with A. dacotensis Whitfield and Hovey, but it is distinguished by
its much less diameter.
Dimensions. Holotype — length, 21 mm.; height, 21; semidiameter,
4*3; apical angle, *108 degrees.
Locality. Talus on small creek one mile east of Billhook creek at
1,500 feet altitude.
Astarte barbara sp. nov.
Plate XV, figure 6
Subquadrate, with the umbo extremely anterior. Ornament of weak,
concentric costae. Basal margin, denticulate. Somewhat resembling
A. calif ornica Stanton, but differing in the more anterior and less prom-
inent umbo.
Dimensions. Holotype — length, 38 mm.; height, 30; semidiameter,
5 '5; apical angle, 116 degrees.
Locality. Zone of Aucella canadiana, west side of Harrison lake 350
yards from shore and 1,400 yards north of the mouth of Deer creek.
Name. The word barbarus denoted to the Homans that which was
not Homan, that is to say, foreign. So this Astarte, a lonely foreigner in
the great community of Aucellae of the canadiana zone, may well be called
barbara.
Family, saxicavidab
Genus, Saxicava Fleurian
Saxicava rugosa Linnaeus
Two well preserved specimens from the clay of the Squawkum for-
mation.
Locality. Main line of Canadian Pacific railway, 750 yards west of
the Harrison Hiver bridge.
65
Class, CEPHALOPODA
Order, Ammonoidea
Family, cardioceratidae
The family name was published first by Hyatt who included Cardio-
eeras, CadoceraSf Quenstedioceras (Hyatt^ always spelled it this way so we
can not accept Quenstedioceras as Buckman insists nor Quenstedticeras as
Reeside demands), and N eumayria. H. Douville’s supposed publication
was not distributed. In 1900 Hvatt attempted to change the name to
Cadoceratidae, but this is not valia. Also some later writers have included
a lot of utterly unrelated genera, which has caused some confusion. How-
ever, excellent summaries of the family have been given by Buckman^
whose interpretation of the group is clear and connected. Many genera
occur in North America as yet unreported. For instance, among the species
described by Reeside® under “Quenstedticeras^^ and Cardioceras there are
a dozen or so generic groups; some named, some unnamed. Two known
genera and one new genus occur in the Upper Jurassic deposits of the
Harrison Lake country. The occurrence of species of Cadoceras in the
Mysterious Creek formation establishes the general age of that deposit.
Also it establishes the exact date of Paracadoceras gen. nov. which is asso-
ciated with Cadoceras. Also it establishes the approximate date of Lillo-
ettia and Buckmaniceras, family Macrocephalitidae, which occur a short
distance above Cadoceras and associated with cadoceratoid forms close to
Cadoceras, but not sufficiently fully grown to be generically identified.
Genus, Paracadoceras nov.
A cadoceratoid serpenticone. The young is like that of Cadoceras,
but inflation that supersedes the elevation of the whorl is so moderate that
the mature form is only a robust serpenticone. Ribbing notably prorsira-
diate. However, the genus differs strongly from Prorsiceras Buckman in
that the number of secondary ribs is less than double that of the primaries,
and in its narrow umbilicus and depressed whorls. Also in the ventral
ribbing.
Paracadoceras harveyi sp. nov.
Plate XVI, figures 1, 2; Figure 2
Ornament. In the young that part of the rib within the umbilicus is
reclined as in the Canadian and Alaskan Cadocerata, but at 27 mm. dia-
meter this part of the rib becomes versiradiate. The extrumbilicate portion
of the rib is prorsiradiate at all stages. Ribs cross venter with only a
slight forward bend. Venter is almost smooth on last part of last whorl.
Table showing number of ribs in a quadrant at various stages.
Diameter
18 mm 10 primaries 14 secondaries
34 “ 8^ “ 14
53 “ 8 “ 15
JHyatt: Bull. Geol. Soc, Am., vol. 3, p. 410 (1892).
*Type Ammonites, vois. II and III (1913-1921).
*Reeflide: U.S. Geol. Suxv., Prof. Paper 118 (1919).
56
OfSmetGn
tr> m m.
Geological Survey, Canada.
Figure 2. Paracadoceras harveyi sp. nov,; width of umbilicus and thickness of whores
expressed as fractions of the diameter. Umbilicus of holotype shown by a
broken line; whorls of holotype by a solid line; continuations of curves, based
on topotypes, by a dotted line.
Locality. Zone of Cadoceras hrooksi, on Deer creek one-quarter mile
from its mouth, west side of Harrison lake.
Name. After Eobert Valentine Harvey.
Genus, Cadoceras Fischer 1882
The North American forms of this genus differ somewhat from the
European, but not enough to warrant separation. The young are very
similar to those of Paracadoceras and other genera from North America
and the Arctic regions as yet unnamed. It is, therefore, quite unsafe to
name a form of less than 45 or 50 mm. diameter unless the development
curves of several features show that the specimen is mature. Much
revision of the Cadoceratoids is necessary, and it is among Arctic faunas
that this can be done. Probably in the Arctic Jurassic is to be found the
early history, as yet unknown, of the great Cardioceratid family.
Two Alaskan and one new species of Cadoceras were found at Harrison
lake.
Cadoceras caiostoma Pompeckj 1900
Figure 3
1900. Pompeckj : Verhandl. Kais. Euss. Min. Gesell. St. Petersburg, 2 te
Ser., Bd XXXVIII.
57
Costae, strong, strongest on venter, bent forward, but not strongly, in
crossing venter, strongly reflexed in crossing umbilical border. At 30
mm. diameter there are eight primaries and twelve secondaries in a quad-
rant. Resembles C. grewingki in form and rib-curve, but differs in its
more closely crowded ribs. Only immature specimens were found.
Dimensions. See graph.
Locality. Zone of Cadoceras hrooksi, on Deer creek one-quarter mile
from its mouth; also other places.
Cadoceras schmidti Pompeckj 1900
Figure 3
1900. Pompeckj : Verhandl. Kais. Russ. Min. Gesell. St. Petersburg,
2 te Ser., Bd. XXXVIII.
Somewhat similar to the last, but having a much narrower umbilicus
and costae bent strongly forward in crossing the venter. At 30 mm. dia-
meter there are eight primaries and fourteen secondaries in a quadrant.
This species like the last has the costae strongly reflexed in crossing the
umbilical border.
Dimensions. See graph.
Locality. Zone of Cadoceras hrooksi, on Deer creek, one-quarter mile
from the mouth; also other places.
Cadoceras hrooksi sp. nov.
Plate XVI, figures 3, 4, 5; Figure 3
Inner whorls rounder in cross-section than is usual in this genus.
Cadicone is attained at about 38 mm. diameter, so possibly a large size is
never reached. Costae, obtuse, very slightly bent forward in crossing
venter, and slightly reflexed in crossing umbilical border. Table showing
number of ribs in a quadrant of whorl at various stages.
Diameter
30 mm. 8 primaries 13 secondaries
50 “ primaries obsolete, 13 “
8 butlse on umbilical
“keel”
Locality. Zone of Cadoceras hrooksi, on Deer creek, one-quarter mile
from its mouth, west side of Harrison lake.
Name. After Mr. Allan Brooks, in recognition of his inimitable pic-
torial contributions to natural history.
Genus, Anacardioceras Buckman 1923
This stock has not previously been reported from North America, but
it seems likely that it is represented by some of the species of ‘‘Cardio-
ceras** described from Wyoming and Alaska. The form from Harrison
lake, described in the sequel, differs from the typical British species in its
perfect smoothness, venter of persistently knife-edge sharpness, and
58
Figures. Cadoceraa brooksi sp. nov.; C. caiostoma; and C. sc/iwidfi; widths of umbilicus
and thicknesses of whorls expressed as fractions of the diameter. Umbilicus
of C. brooksi, holotjT>e, shown by a solid line marked U; thickness of C. brooksi,
holotype, by a solid line marked T; umbilicus of Harrison Lake specimens oi
C. colostoma, by a broken line marked U ; thickness of specimens of C. colostoma
by a broken line marked T; umbilicus of Harrison Lake specimens of C. sch~
midti by a dotted line marked U; thickness of specimens of G. sckmidti by a
dotted fine marked T.
slightly ellipticoDic periphery. However, these differences are acquired
only with late maturity so are not to be taken as a basis for separation.
The genus indicates a late Cardioceratan date : most of the European species
being in the excavatum zone.
Anacardioceras perrini sp. nov.
Plate XVII, figures 1, 2, 3; Plate XXII, figure 1; Figures 4, 5, 6
Living chamber, one-half whorl. Ornament, striate in the young.
Becomes heavy enough to be called costate at about 20 mm. diameter.
59
Ornament reaches acme at 45 mm., then declines. Smooth from 90 mm.
on. Table showing number of ribs in a quadrant of whorl at various
stages.
Diameter
16 mm. 8 primaries 13 secondaries
30 “ 6 “ 16
62 « 7 “ 15
and 17 peripheral knots
Venter rounded up to 5 mm. at which it becomes fastigate. It develops a
knotted keel at about 15 mm. Loses its knots about 85 mm. becoming
more acute. A true knife-edge from 90 mm. onward. Mouth border
Figure 4. Anacardioceras perrini sp. nov.; correlation of diameter and number of whorls.
inclined and rostrate. Conch form, complicated — an oxygastric contracti-
ellipti-oxycone. The species differs from the genotype in having the
primary rib furcation nearer the umbilicus: about one-third way across the
60
flank. The ribs are more crowded. Also the persistently acute venter,
smoothness, and catagenetic peripheral spiral. The last is not uncommon
in various large Cardioceratoids.
hjL = L2 Li
Locality. On the west side of the point that bounds on the west the
small bay on the southwest shore of the peninsula.
Name. After my teacher, Professor James Perrin Smith.
v\/hc>r'ls
V
\
\
/
/
/
^ /
-
-
M
\
\
\\
\ \
1 \
-
1 '
1 \
1
y
/
1 .
-
O O.P 0.-4 0.6
Geological Survey, Canada.
Figure 5. Anacardioceras perrini sp. nov.; width of umbilicus (shown by a broken line) and
thickness of whorls (by a solid line) of holotype correlated with number of
whorls.
Family, macrocephalitidae
Genus, Lilloettia nov.
The family is represented by two new genera of slightly later date than
most of the group and corresponding in age to Catacephalites. Both are
widespread in deposits of the same age in southern Alaska where they are
associated with Cadoceras sensu lato.
The young of Lilloettia have the ornament and something of the form
of DolikephaliteSj but a smaller umbilicus. The ornament degenerates
during late adolescence. Smoothness supervenes first round the umbilicus.
61
then spreading across the flank it finally extinguishes the ventral orna-
ment. Conch form, a compressed sphaerocone from youth on. Flanks
become convergent with maturity. Mouth border, plain, i.e., without
lappets, strongly inclined, slightly swollen. A helically twisted umbilical
Figure 6. Anacardioceras perrini sp. nov.; width of umbilicus (shown by a broken line)
and thickness of whorls (by a solid line) of holotype expressed as fractions of
the diameter.
columella is present. Differs from other macrocephalitids in its early
smoothness and narrow umbilicus. Differs from Buckmaniceras in septal
line: L1>EL, LI has longer though less divergent branches; and in conch
form. Septal line rather simple as late as early maturity. Resembles
Buckmaniceras in its obtuse ornament.
93289—5
62
Lilloettia lilloetensis sp. nov.
Plate XVIII, figures 1-4; Figure 7
Body chamber of holotype seven-eighths whorl.
Table showing number of ribs in a quadrant of whorl at various stages.
Diameter
27 mm. 6 primaries 16 secondaries
55 “ primaries obsolete 17 “
70 “ 19
85 “ complete smootlmess
Dimensions. See graph under Buckmaniceras.
Locality. Zone of Lilloettia lilloetensis, on Billhook creek at 3,100 feet
altitude.
Name. Of both genus and species Indian geographical name Lillooet.
Lilloettia mertonyarwoodi sp. nov.
Plate XIX, figures 1, 2; Figure 7
Body chamber of holotype slightly more than two-thirds whorl.
Table showing number of ribs in a quadrant of whorl at various stages.
Diameter
55 mm . 8 primaries 25 secondaries
60 “ primaries obsolete 25 “
80 “
87 complete smoothness
Dimensions. See graph under Buckmaniceras.
Locality. Zone of Lilloettia lilloetensis, on Billhook creek at 3,100 feet
altitude.
Name. After my former teacher, Professor Merton Yarwood W il-
liams.
Genus, Buckmaniceras nov.
Robust, rounded, sphaerocones with the ornament of Lilloettia, from
vouth on. Differs from Lilloettia in a few important respects, such as a
total lack of compression and in septal line. EL = L1, LI branches are
short.
Buckmaniceras buckmani sp. nov.
Plate XX, figures 1-4; Figure 7
Table showing number of ribs in a quadrant of whorl at various stages.
Diameter
23 mm. 6 primaries 16 secondaries
55 “ primaries obsolete 17 “
90 “ complete smoothness
63
D/^metjen
O 0.5 /.b
G.S.C.
Figure 7. Lilloettia liUoeiensis sp. nov. shown by a solid line; L. merionyarwoodi sp. nov.
by a dotted line; and Buckmaniceras buckmani sp. nov. by a broken line; thick-
ness of whorls of holotypes expressed as fractions of the diameter.
Locality. Zone of Lilloettia lilloetensis, on Billhook creek at 3,100 feet
altitude.
Name. Of genus and species after Mr. S. S. Buckman, the great
master of ammonitology.
Family, virgatitidae
Genus, Homolsomites nov.
To be included in this family is the great series of forms of late Jurassic
and early Cretaceous showing relationship to the planulates with virgatome
ornament typified by Virgatites. The genus Homolsomites includes platy-
cones with narrow umbilicus and narrow though rounded venter. Costse,
delicate, branching according to various plans, but mostly virgatome,
and crossing centre unbroken and with a strong forward bend. In maturity
all but the ventral ornament become obsolete. Septum, strongly reclined.
Stems of lobes narrower than is usual with this family.
Name. Indian geographical — Homolsom.
Homolsomites poecilochotomus sp. nov.
Plate XXI, figures 1-4
Ornament is striate up to early maturity. It branches according to
several plans and combinations thereof, including dichotomous, bidicho-
tomous, and virgatome (3-branch). The plans of branching become more
64
complex, thereby increasing the number of secondary ribs. After 35 mm.
diameter the costae on the umbilical half of the flank become obsolete,
after which the remaining ornamented area narrows so that at 100 mm.
only the centre and one-sixth of the flank are costate.
Table showing number of ribs in a quadrant of whorl at various stages.
Diameter
20 mm. 9 primaries 21 secondaries
35 “ 8 “ 33 “
90 “ primaries obsolete 25 "
Dimensions. Holotype has a maximum diameter of about 105 mm.
Thickness at this diameter is 18 mm. Width of umbilicus, 18 mm.
Name. In allusion to the various plans of branching of the costae.
Locality. Zone of Homolsomites poecilochotomus, on the southeast
shore of the peninsula.
Family, phylloceratidae
Genus, Phylloceras Suess 1865
Phylloceras columbianum sp. nov.
Plate XXII, figures 1, 2, 3
Shell, quite smooth. Septal line with about seven lobes, exclusive
of the external lobe. Mouth-border unknown.
Dimensions. Holotype — maximum diameter, 65 mm.; major radius,
40-5; minor radius, 24*5; thickness, 21-5; thickness at minor radius, 16;
umbilicus, 4 • 5 per cent of diameter.
Locality. On the southwest shore of a small point on the southwest
side of the small bay on the southwest shore of the peninsula, where it is
associated with Anacardioceras perrini.
Name. Geographical from British Columbia.
Phylloceras'^ aff. knoxvillense Stanton
Stanton: Bull. 133, U.S. Geol. Surv.
Fragments of a very large form highly similar in ornament and septal
line to Stanton’s species. It may differ in proportions. Not suflficiently
complete for naming or special description.
Locality. Zone of Aucella canadiana, 350 yards from shore and 1,400
yards north of the mouth of Deer creek.
^^Phylloceras" sp. indet.
Fragments of a smooth, latumbilicate ‘‘Phylloceras", having a complex
septal line. Quite distinct from Phylloceras s.s., but not sufficiently
complete for further description.
C5
Order, Belemnoidea
Family, belemnitidae
Genus, Cylindroteuthis Bayle 1879
Cylindroteuthis themis sp. nov.
Plate XXIII, figures 1, 2
Rostrum, cylindroid, very blunt pointed. Dorsum and flanks,
smooth, rounded. Venter, straight, flattish, bearing a broad, shallow
canal which is deepest near the apex and becomes so shallow as to disappear
15 mm. from the apex. Taper, very slight, except in the apical region.
Axis, slightly eccentric. Phragmacone, very nearly right conical. Septa,
alveolus, and proostracum, not preserved.
Dimensions
Vertical diam., at apex of phragmacone 10 mm.
Ventral radius “ “ 3-5
I^ateral diam. “ ** 10-5
Axis, apex of phragmacone to apex of rostrum 40
Apical angle of phragmacone, lateral 21 degrees
Locality. Associated with Entolium vulcanicum, on the west shore
of Harrison lake, 1,820 yards north of Harrison River point.
Name. Themis, of Greek mythology.
Cylindroteuthis haculus sp. nov.
Plate XXIII, figures 3, 4
Rostrum, very long, almost perfectly cylindrical through most of its
length. Resembling ‘^Belemnites” tehamaensis Stanton and certain Russian
species of similar date, such as ^‘Belemnites” obeliscoides Pavlow and
Lamplugh, but differs from some of these in its very gentle taper, from
others in its almost perfectly circular cross-section. The holotype has a
nearly right conical phragmacone, containing thirty-three chambers and
occupying two-fifths of the length of the alveolus. The alveolus is slightly
more than one-fifth the entire length of the rostrum.
Dimensions
Vertical diam, at apex of phragmacone 14 mm.
Ventral radius ** “ 6*5
Lateral diam. “ “ 15
Axis, from apex of phragmacone to apex of rostrum, from
a composite specimen 230
Phragmacone, length 20
Alveolus, length 50
Apical angle of phragmacone. 15 degrees
Locality. Zone of Aucella canadiana, 350 yards from shore and 1,400
yards north of the mouth of Deer creek.
Name. In allusion to external form.
66
Genus, Pachyteuthis Bayle 1879
Pachyieuthis eocretacicus sp. nov.
Plate XXII, figure 4; Plate XXIII, figure 5
nostrum, very short and thick, resembling the Russian species,
“Belemniies” subquadratus Pavlow and Lamplugh, but differing slightly
though distinctly in outlines and cross-section. Surface, perfectly smooth.
The type specimens are moulds.
Dimensions
Vertical diam. at apex of phragmacone 31 mm.
Ventral radius ** " 15
Lateral diam. “ 31
Alveolus, length 65
Axis, apex of pliragmacone to apex of rostrum 59
Apical angle of phragmacone 21 degrees
Locality. Zone of Aucella canadiana, 350 yards from shore and 1,400
yards north of the mouth of Deer creek.
Name. In allusion to the date of existence of the species — the dawn
of the Cretaceous.
Phylum, ARTHROPODA
Class, CRUSTACEA
Order, Cirripedia
Family, balanidae
Genus, Balanus da Costa
Balanus crenatus Brugui^re
Several well-preserved fragments of this species. This may well be
the form referred to by Lamplugh as ‘‘Balanus sp.”
Locality. Main line of Canadian Pacific railway 838 yards west of
the west end of the Harrison River bridge.
68
Plate I
Baadites crickmayi n. sp. (Page 3.)
Plate I
70
I’late 11
I’igiires 1, 2, 3. Baculites crickmayi n. sp. (Page 3.)
Figures 4, 5, t), 7. Veniella subtrapeziformis (\\ hiteaves) var. dyen var. aov. (1 a;e 1.)
Plate II
7
72
Plate III
Figure 1. Vwi'parus nidaga sp. nov. (Page 9.)
Figure 2. Unto mhprimaens sp. nov. (Page 7.)
Figure 3. Viviparits crickmayi sp. nov. (Page 8.)
Figure 4. Campeloma vetula tenuis var. nov. (Page 10.)
Figure 5. Vitfiparus tasgina sp. nov. (Page 10.)
Figure 6. Campeloma praecursa sp. nov. (Page 11.)
Figiu*e 7. Campeloma cy^essensis sp. nov. (Page 11.)
Figure 8. Goniobasis williamsi sp. nov. (Page 12.)
Figures 9, 10. Goniobasis subtortuosa mut. tenuis n. mut. (Page 11.)
Figure 11. Goniobasis whiUakeri sp. nov. (Page 12.)
Figure 12. Goniobasis judithensis minimus var. nov. (Page 12.)
Figure 13. Pupa sp. indet. (Page 14.)
Figure 14. Goniobasis webbi sp. nov. (Page 14.)
Figiu'e 15. Unio mcleami sp. nov. (Page 7.)
Figure 16. Melania whiteavesi nodosa var. nov. (Page 13.)
Plate III
16
74
Plate IV
I'igure 1. Unio humei sp. nov. (Page 8.)
Figures 2, 3. Velatella rectistriata sp. nov.
Figure 4. Unio humei sp. nov. (Page 8.)
(Page 13.)
Plate IV
76
Plate V
Figure 1. Legwnmosiles williamsi Berry n. sp. (Page 23.)
I'igures 2, 3. Trochodendroides cuneata (Newberry). (Page 20.)
Figure 4. Viburnum finale Ward. (Page 27.)
Figure 5. Phylliies aquaticm Berry n. sp. (Page 28.)
Figure 6. Cercocarpus ravenscragensis Berry n. sp. (Page 23.)
Figure 7. Paliurus (?) sp. (Page 25.)
Figure 8. Trochodendroides speeiosa (Ward). (Page 22.)
:PLAT£; Y
78
Plate VI
Figures 1-3. Arisiolochia crassifolia Cockerell (drawing now ^ natural size). (Page 20.)
Plate VI
80
Plate VII
Eodelphis cuileri (Smith Woodward) Cat. 8536, Geol. Surv., Canada.
size. (Page 30.)
Figure 1. Internal view.
Figure 2, Superior view.
Figure 3. External view.
2-28 times natural
Plate VII
£
3
82
Plate VllI
Figure 1.
Figure 2.
Figure 3,
Figure 4.
Parallelodon cardioceratanum Crickmay sp. nov., holotype, X 3-28. Locality
No. 24, Upper Jurassic. (Page 45.) • . r tLo
Parallelodon cardioceratanum Crickmay sp. nov., mk-prmt of the t^th of the
holotype (gelatine process), X 3-6. The black areas correspond to the sockets
of the right valve. (Page 45.) , . x i •
Parallelodon canlioceraianum Crickmay sp. nov., holotype about natural size.
Mckarnia wcZearm ^Crickmay sp. nov., holotype, about natural size. A right
valve showing byssal sinus, small central chondroplmre, and single, large
muscle- scar. Locality No. 34. Lower Cretaceous. (Page 45.)
Plate VIII
4
84
Figure 1.
Figure 2.
Figure 3.
Figure 4.
Figure 5,
I
Plate IX
Mcleornia mcleunii Grickmay sp. nov., paratype, about natural size. A left
valve, natural cast of interior of shell. 8ho\v.s a peculiar sinuous senes of
small muscle scars quite separate and distinct from the large adductor scar.
Locality No. 34. Lower Cretaceous. (Page 45.)
Aucellu oculislriata Crickmay sp. iiov., holotyjoe, about natural size. A leit
valve. Locality No. 33. Lower Cretaceous. (Page 46.)
Aucella spasskejisoides Crickmay sp. nov., holotype, about natural size. A
right valve. Locality No. 28. Lower Cretaceous. (Page 47.)
Aticella spasskensoides Crickmay sp. nov., paratype, about natural .size. A lett
valve. Locality No. 28. Lower Cretaceous. (Page 47.) \ i i.
AiiccUa eatamorpha Crickmay sp. nov., holotype, about natural size. A light
valve. Locality No. 28. Lower Cretaceous. (Page 46.)
Plate IX
86
Plate X
Figure 1,
Figure 2.
Figure 3.
I'igure 4.
I- igure 5.
Figure 6.
Figure 7.
A ucelki
valve.
Aiicelki
valve.
A ucelki
valves
cascadattiis Crickinay sp. iiov., liolotyiie, about natural size. A left
Locality No. 33. Lower Cretaceous. (Page 47.) . ■
cascadevsis Crickmay sp. rov., paralype, about natural size. A right
Locality No. 33. Lower Cretaceous. (Page 47.)
canadiana Crickir.ay sp. nov., paratjTie, about natural size, two
in contact. Eight aspect. LocaUty No. 33. Lower Cretaceous.
iucella canadiana Cricknr.ay sp. nov., liolot 3 'pe, about natural size,
valve. Locality No. 33. Lower Cretaceous. (Page 47.)
Aucella canadiava Crickir.ay sp. nov., holotjTe, antenor aspect. (1 age 47.)
^ucella kuoiekcnsis Crickmay sp. rov., wax cast of holotype. (Holotyiie con-
sists of natural cast of interior of shell, and a small imipression of part of the
external surface.) A right valve. Locality No. 39. Lower Cretaceous.
(I age 48.) ^ ^ Anterior aspect of the specim.en of
A left
AuccUu ku'oickeiisis Crickmaj’’
figure 0. (Page 48.)
sp.
Plate X
88
Plate XI
Figure 1.
Figure 2.
Figure 3.
Figure 4.
Figure 5.
Figure 6.
Avcella harrisonensis Crickmay sp. nov^ paratype, X l-S. Anterior aspect of
left valve. Locality No. 41. Lower Cretaceous. (Page 48.)
A ucella harrisonensis Crickmay sp. nov., paratype, X 1 • 8. Left valve. Locality
No. 41. Lower Cretaceous. (Page 48.) ,,, r i ^ •
A ucella hanisonensis Crickmay sp. nov., holotype, X 1 - 5. Kight valve, anterior
aspect. Locality No. 41. Lower Cretaceous. (Page 48.) . ^
A ucella gigas Crickmay sp. nov., holotype, X *53. Two valves in contact, left
aspect. Locality No. 41. Lower Cretaceous. (Pa^ 49.)
Aucella gigas Crickmay sp. nov., holotype, X -53. Right aspect. (^S6 49.)
Aucella gigas Crickmay sp. nov., holotype, X -49. Anterior aspect. (Page 49.)
Plate XI
90
Plate XII
Figure 1.
Figure 2.
Figure 3.
Figure 4.
Figure 5.
Figure 6.
Haidaia billhookensis Crickmay sp. nov., plaster cast of holotype, an external
mould of the left valve. Anterior and dorsal outline, partly restored. Locality
No. 21. Upper Jurassic. (Page 49.)
Haidaia packardi Crickmay sp. nov., plaster cast of holotype, part of which
is an external mould of the right valve, -73 of natural size. Locality No. 21.
Upper Jurassic. (Page 50.)
Haidaia packardi Crickmay sp. nov., internal natural cast of holotype, with
some shell attached, -73 of natural size. Locality No. 21. Upper Jurassic.
(Page 50.)
Haidaia statlmnsis Crickmay sp. nov., plaster cast of holotype, an external
mould of two valves in contact, left aspect, -73 of natural size. Shows outline
and conch contour well, but gives poor idea of ornament. Locality No. 22.
Upper Jurassic. (Page 50.)
Haidaia statluensis Crickmay sp. nov., wax cast of first paratype, -73 of natural
size, to show ornament. Locality No. 22. Upper Jurassic. (Page 50.)
Haidaia statluensis Crickmay sp. nov., wax cast of second paratype. (Page 50.)
Plate XII
92
Plate XIH
Figure 1. Yaadia lewisagassizi Crickmay sp. nov., wax cast of holotype, an external
mould of left valve, *87 of natural size. Locality No. 39. Lower Cretaceous.
(Page 50.)
Figure 2. Yaadia lewisagassizi Crickmay sp. nov., wax cast of holotype, *87 of natural
size, postero-dorsal aspect. Shows ornament of area and escutcheon. (Page 50.)
Figure 3. Quoiecchia aliciae Crickmay sp. nov., holotype, -87 of natural size, posterior
aspect. Locality No. 39. Lower Cretaceous. (Page 51.)
h^gure 4. Quoiecchia aliciae Crickmay sp, nov., holotype, a right valve, -87 of natural
size. (Page 51.)
Figure 5. Quoiecchia aliciae Crickmay sp. nov., holotype, anterior aspect, (Page 51.)
Figure 6. Quoiecchia aliciae Crickmay sp. nov., paratype. Juvenile, right valve. Locality
No. 39. Lower Cretaceous. (Page 51.)
Figure 7. Quoiecchia aliciae Crickmay sp. nov., paratype, same specimen as figure 6,
dorsal aspect, -87 of natural size. (Page 51.)
Figure 8. Quoiecchia aliciae Crickmay sp. nov., paratype, internal mould of right valve,
X 1*9. Shows crenulation of ventral margin. Locality No. 39. Lower
Cretaceous. (Page 51.)
Plate XIII
94
Plate XIV
Figure 1. Entolium hertleini Crickmay sp. nov., holotype, a right valve, external aspect,
X 3*19. Shows the microscopic ornament. Dark line shows curve of con-
centric striae. Locality No. 13. Upper Jurassic. (Page 52.)
Figure 2. Entolium vulcanicum Crickmay sp. nov., holotype, an internal mould of left
valve, X -66. Locality No. 3. Middle Jurassic. (Page 52.)
Figure 3. Entolium aucellarum Crickmay sp. nov., holotyp>e, an external mould of right
valve, A natural size. Locahty No. 28. Lower Cretaceous. (Page 52.)
Figure 4. Anomia colurrMana Crickmay sp. nov.^ holotype, external aspect, natural
size. Locality No. 15. Upper Jurassic. (Page 53.)
Figure 5. Anomia colunwiana Crickmay sp, nov., holotype, X 1*9. (Page 53.)
Figure 6. Anomia Columbiana Crickmay sp. nov,, paratype, external mould with some
shell attached. Locality No. 15. Upper Jurassic. (Page 53.)
Plate XIV
5
6
96
Plate XV
Figure 1. PUuromya harriso^ierisis CrickiLay Bp. nov., holotype, two valves together,
right aspect, X 1*3. Locality Ko. 41. Lower Cretaceous. (Page 53.)
f'igure 2. Phnroniya /,ormoT<tt?sts Crick n ay Bp. r.ov., holot> 7 )e, dorsal aspect, X 1-18.
(Page 53.)
Figure 3. Pleuromya harrisonensis Crickniay sj>. nov., holotype, anterior aspect, ttht
natural size. (Page 53.)
Figure 4. A&tarte harrisomnsis Crickmay sp. nov., artificial cast in clay of part of holo-
type, mould of two valves in contact, X 1-5. Left aspect, showing ornament.
Locality No. 19. Uimer Jurassic. (Page 54.)
Figure 5. Astarte harriwnensis Crickmay sp. nov., artificial cast of holotype, X 1'5.
Left aspect, showing outline, etc. (Page 54.)
Figure 6. Asiurte harhara Crickmay sp. nov., holotype, a left valve with shell mostly
decorticated, jVtj ii^fural size. Locality No. 34. Lower Cretaceous. (Page 54.)
Plate XV
1
3
98
Plate XVI
Figure 1.
Figure 2.
Figure 3.
Figure 4.
Figure 5.
Paracadocerm harveyi Crickmay sp. nov., holotype, about natural size. An-
terior aspect, showing mouth-border, which is somewhat distorted, and ventral
portion of septal lines. Locality No. 12. Upper Jurassic. (Page 55.)
Paracadoceras harveyi Crickmay sp. nov., holotype. Lateral aspect. Shows
living chamber, f whorl. (Page 55.)
Cadoceras brooksi Crickmay sp. nov., holotype, about natural size. Ventro-
lateral aspect. Locality No. 12. Upper Jurassic. (Page 57.)
Cadoceras brooksi Crickmay sp. nov., holotype, about natural size. Lateral
aspect. (Page 57.)
Cadoceras brooksi Crickmay sp. nov., holotype, with some of the missing parts
restored in clay, X 1 *6. (Page 57.)
Plate XVI
5
100
Plate XVII
Figure 1. AriacardiocercLs perrini Crickmay sp. nov., cross-sectional outlines of holotype,
taken at 90 degrees behind the mouth border, A natural size. From external
mould of holotype. (Page 58.)
Figure 2. Anacardioceras perrini Crickmay sp. nov., holotype, lateral aspect, -h natural
size. The reconstructed peripheral curve and mouth-border were obtained
from part of the holotype not included in the photograph, namely an external
mould of the full-grown shell. At 5 and 5i whorls the ribbing and venter
are well shown. At 61 whorls a small fragment shows the ornament of the
umbilical border. The entire specimen consists of 7i whorls. Locality No.
24. Upper Jurassic. (Page 58.)
Figure 3. Anacardioceras perrini Crickmay sj). nov., parat}™, X 5-2 cross-sectional
outlines at 3f whorls. Protoconch is marked by a line of dashes which repre-
sents the central axis of the shell. Shows elevation of whorl between 2f and
3i whorls, and appearance of fastigate venter between 31 and 31. Locality
No. 24. Upper Jurassic. (Page 58.)
Plate XVIT
3
102
Plate XVIII
Figure 1. lAlloetlia Ulloeiensis Crickmay sp. nov., holotype, genotype, apertural aspect,
about natural size. Last whorl slightly distorted by diastrophic forces. Mouth
border outlined in black. Lost parts of shell partly restored in outline from
measurements of the rest of the shell. Locality No. 15. Upper Jurassic.
(Page 62.)
Figure 2. Ldlloettia lilloetensis Crickmay sp. nov., holotype, genotype, lateral aspect,
about natural size. Mouth border outlined in black. Periphery partly
restored in outline by proportional method (vide Crickmay: Am. Jour. Sci.,
5th ser., XIII, 1927). (Page 62.)
Figure 3. Lilloettia lilloetensis Crickmay sp. nov., holotype, about natural size, lateral
aspect. Shows ornament of fiftn whorl. (Page 62.)
Figure 4. lAUoettia lilloetensis Crickmay sp. nov., holotype, peripheral aspect, about
natural size. Same specimen as shown in figure 3 — the internal whorls of
the holotype. (Page 62.)
104
Plate XIX
Mgure 1. Lilloeltia nierlonyarwoodi Crickmay sp. nov., holotype, lateral aspect, 1 natural
size. Mouth border restored in outline from about one-third of it preserved
and by the analogy of the other species of this genus. Periphery restored
purely on basis of proportional method. This is unreliable when used for
extrapolation, especially on the ultimate whorl of ammonites. So this outline
is intended only to complete the picture and so elucidate the interpretation
of the specimen. Ornament is seen 1 and IJ whorls behind mouth border.
Locality No. 15. Upper Jurassic. (Page 62.)
Figure 2. Lilloeltia inert onyarwoodi Crickmay .sp. nov., holotype, peripheral aspect, 1
natural size. (Page 62.)
Plate XIX
106
Plate XX
Figure 1.
Figure 2.
Figure 3,
Figure 4.
Buckvianiceras huckmani Crickmay sp. nov., holotype, lateral aspect, about
natural size. Small black line shows position of emergence of penultimate
whorl. Mouth border not preserved. Dotted lines show the great thickness
of shell substance around the umbilicus. Locality No. 15. I pper Jurassic.
eSfcwaSras huckmnni Crickmay sp. nov., holotj^e, peripheral aspect, about
natural size. This, as well as figure 1, shows the disappearance of ornament on
the last whorl. (Page 62.) , , , i . i + f
Buckvianiceras huck 7 nani Crickmay sp. nov., holotype, last quadrant oi penul-
timate whorl, lateral aspect, about natural size. (Page 62.) , , r i
Buckvianiceras huckmani Crickmay sp. nov., holotype, last quadrant ot penul-
timate whorl, ventro-lateral aspect, about natural size. (Page 62.)
Plate XX
2
108
i’LATE XXI
Figure 1.
Figure 2.
Figure 3.
Figure 4.
Homolsondtes poedlochotomus Crickmay ap. nov., paratype, X l-o. Locality
No. 41. Lower Cretaceous. (Page 63.)
Homolsomitcs poedlochotomus Crickmay sp. nov., holotype, • 77 ot natwaj
size. Restoration of spiral by proportion. Locality Xo. 41. Lower Lre-
taceous. (Page 63.) -r- r * i ■
H ornolsomites poedlochoto 7 nu^ Crickmay sp. nov., paratype, -7/ oi natural size.
Locality No. 41. Lower Cretaceous. (Page 63.) v q no
H ornolsomites poedlochotomus Crickmay sp. nov., mratype, A o-us.
nrimary ribs are outlined with their secondanes. They show two styles of
furcation common with this species, the bidichotomous and the virgatome.
Locality No. 41. Lower Cretaceous. (Page 63.)
Plate XXI
/ %
■- « J
n^'
1 1
i J
, iv A
110
Plate XXII
Figure 1. Phylloceras colunibianum Crickmay sp. nov., holotype, natural size. A young
specimen, and several impressions of Anacardioceras perrini are to lie seen on
this specimen. Locality No. 24. Upper Jurassic, (Pages 58, 64.)
Figure 2. Phylloceras colunibianum Crickmay sp. nov., holotype. (Page 64.)
Figure 3. Phylloceras columbianum Crickmay sp. nov., holotype. (Page 64.)
Figure 4. Pachyteuthis eocretacicus Crickmay sp. nov., cross-sectional outhne of paratype,
natural size. Locality No. 34. Lower Cretaceous. (Page 66.)
Plate XXII
112
Plate XXI II
Figure 1.
Figure 2.
Figure 3.
I igure 4.
Figure 5.
::ylindroteuthu themis Crickmay sp. nov., holotype, lateral aspect, -66 of natural
size. Locality No. 3. Middle Jurassic. (Page 65.) * c
Cylindroteuthis theniis Crickmay sp. nov., holotype, ventral aspect, -bb ot
natural size. (Page 65.) * c-a „r
Cylindroteuthis baculus Crickmay sp. nov., holotype, lateral aspect, -66 of natural
size. Locality No. 34. Lower Cretaceous. (Page 65.) . i .
Cylindroteuthis baculus Crickmay sp. nov., outhnes restored from the holotype
and several paratypes, -66 of natural size. (Page 65.) u i + i
Pachyteulhis eocretacicus Crickmay sp. nov^ holotj^pe, an external mould, lateral
view X -44 . Locality No. 34. Lower Cretaceous. (Page 66.)
Plate XXIII
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J
J
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