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MIDDLE ALBIAN STRATIGRAPHY
LIN. EEE
ANGLO-PARIS BASIN

BY

HUGH GWYN OWEN

3 Plates, 52 Text-figures

BULLETIN OF
THE BRITISH MUSEUM (NATURAL HISTORY)
GEOLOGY Supplement 8
LONDON : 1971

THE BULLETIN OF THE BRITISH MUSEUM
(NATURAL HISTORY), instituted im 1949, 1s
issued in five series corresponding to the Departments
of the Museum, and an Historical series.

Parts will appear at irregular intervals as they become
veady. Volumes will contain about three or four
hundred pages, and will not necessarily be completed
within one calendar year.

In 1965 a separate supplementary series of longer
papers was instituted, numbered serially for each
Department.

This paper is Supplement No. 8 of the Geological
(Palaeontological) series. The abbreviated titles of
periodicals cited follow those of the World List of
Scientific Periodicals.

© Trustees of the British Museum (Natural History), 1971

World List abbreviation
Bull. Br. Mus. nat. Hist. (Geol.).

TRUSTEES OF
THE BRITISH MUSEUM (NATURAL HISTORY)

Issued 19 January, 1971 Price £6

MIDDLE ALBIAN STRATIGRAPHY

IN THE
ANGLO-PARIS BASIN

By H. G. OWEN

CONTENTS

I. INTRODUCTION
II, AcKNOWLEDGEMENTS
III. DrEscRIPTION AND CORRELATION OF SECTIONS
A. WEALD
(i) FoLKESTONE :
(ii) FOLKESTONE TO THE Mrpway
(iii) MEDWay TO TROTTISCLIFFE
(a) Paddlesworth
(b) Trottiscliffe
(iv) SEVENOAKS AREA
(a) Sevenoaks Brick Works Ltd
(b) St. John’s Brickyard
(c) Dunton Green
(d) Brasted ‘
(v) BrasTtED TO BUCKLAND
(a) Westerham
(b) Tandridge .
(c) Buckland ‘ :
(vi) BucKLAND TO UPPER BEEDING :
(vii) UPPER BEEDING
(viii) Hassocks To EASTBOURNE
B. IsLe or WicutT, Dorset Coast
(i) IsLE oF WIGHT
(a) Redcliff
(b) Rookley
(c) Compton Bay
(d) Ventnor to Niton
(e) Blackgang . .
(ii) BALLARD CLIFF TO OsmincTon ;
(iii) THORNCOMBE BEACON TO BLACK VEN
(iv) CoNCLUSIONS
C. THE OUTCROP FROM DEvon TO BEDFORDSHIRE.
(i) OKEFORD FITzPAINE (DORSET)
(ii) VALE oF WARDOUR TO DEVIZES (WiLtsHIRE)
(a) Vale of Wardour 3
(b) Maiden Bradley to Devizes
(c) Caen Hill, Devizes
(iii) Devizes TO THAME (Oxon)
(a) Badbury Wick (Wiltshire)
(b) Badbury to Thame

MIDDLE ALBIAN STRATIGRAPHY

(iv) THAME TO LEIGHTON-BUZZARD (BEDFORDSHIRE) .
(a) Long Crendon (Bucks)
(b) Haddenham (Bucks)
(c) Aylesbury (Bucks) ;
(d) Aylesbury to Leighton Buzzard
E. BOREHOLE EVIDENCE : :
(i) HAMPSHIRE BASIN
(a) Winchester district :
(b) Relationship of Winchester to Portsdown, the Weald,
and the Isle of Wight : :
(ii) Lonpon Basin, East ANGLIA, AND KENT
(a) London Basin, S. Essex, AND N. Kent
(b) The area of the Kent Coalfield
F. SELECTION OF SECTIONS IN FRANCE
(i) COMPARISON BETWEEN WISSANT AND FOLKESTONE
(ii) THE OUTCROP FROM THE RIVER ORNAIN (MEUSE)
To THE RIVER ARMANCE (YONNE)
(a) Revigny-sur-Ornain (Meuse)
(b) Pargny-sur-Saulx (Marne)
(c) Les Cétes Noires (Haute Marne)
(d) Courcelles prés Clérey (Aube)
(e) La Vendue Mignot (Aube)
(f) St. Florentin area (Yonne)
(g) Summary
(iii) PAys DE Bray
(a) Villers St. Barthélemy . :
(b) Forges-Les-Eaux, and St. Martin
(iv) COMPARISON BETWEEN THE Pays DE CAUX AND THE
IsLE-OF-WIGHT .
(a) St. Jouin
(b) Cauville
(c) Octeville
(d) Cap de la Héve
(e) Summary, and comparison with the Isle- -of- -Wight
IV. DEFINITION OF THE MIDDLE ALBIAN SUBSTAGE AND ITS ZONAL
SCHEME IN THE ANGLO-PARIS BasIN
A. HISTORICAL BACKGROUND
B. THE ZONAL SCHEME OF THE MippLE ALBIAN IN THE ANGLo-
Paris BASIN
(1) DEFINITION OF THE BASE OF THE , MippLe ALBIAN
(ii) THE SUBZONAL SEQUENCE
(a) Subzone of Hoplites (I sohoplites) codentatus
(b) Subzone of Lyelliceras lyelii . : 3
(c) Subzone of Hoplites (Hoplites) spathi
(d) Subzone of Anahoplites intermedius
(e) Subzone of Dimorphoplites niobe
(f) Subzone of Mojsisovicsia subdelaruet
(g) Subzone of Euhoplites meandrinus
(h) Subzone of Euhoplites nitidus
(i) Subzone of Anahoplites daviesi
(iii) THE POSITION OF THE SUBZONE OF Dipoloceras cristatum
(iv) THE ZONAL GROUPING :
(a) The Zone of Hoplites (H.) dentatus
(b) The Zone of Euhoplites loricatus

IN THE ANGLO-PARIS BASIN 5

(c) The Zone of Euhoplites lautus : 4 : ‘ 129

V. LINKS WITH OTHER FAUNAL PROVINCES . 4 ; 130

A. THE BOUNDARIES OF THE HOPLITINID PROVINCE : : 130

B. LINKS WITH SEQUENCES OF OTHER COUNTRIES Z . 133

(i) Wrest PAKISTAN ; ; j ; : ; : 133

(ii) TETHYAN BELT ? : : : ‘ 5 : 133

(iii) ALGERIA : : : : . . ‘ ‘ 134

(iv) SOMALIA . : : 2 ; 5 : 2 : 134

(v) MapaGascaR . : F ; ! : 5 : 134

(vi) SoutH AMERICA : : : : : : : 134

(a) Colombia . : : : : : : : 134

(b) Peru . ; : : : - . ; : 135

(vii) Texas. : : : ; ‘ : j ‘ 135

(viii) CANADA . : : 3 ‘ ; Z , ‘ 137

(ix) GREENLAND. ; i : ; ; 5 : 138

(x) CONCLUSION . ; ; : ; : 138

VI. CONDITIONS OF DEPOSITION IN “ENGLAND : 139
(a) The Margins of the depositional basin in England and

Northern France : 140

(b) The Structural Controls on Depositioni in Southern England 143

(c) Source of Middle Albian Sediments E : ‘ : 147

(d) The cristatum Subzone disturbance : : . : 148

VII. REVIEW OF THE AMMONITE FAUNA . : : ; : : 149

A. Description of new species. 5 : : : F 149

B. Stratigraphical list ‘ : : ; : : ‘ 152

VIII. REFERENCES ‘ : ‘ s : : : : 155

SYNOPSIS

Much new information on the stratigraphy of the Middle Albian substage gained from sections
in England and France is recorded, and where possible previous knowledge is revised, sufficient
to stabilize the Ammonite Zonal scheme in the Anglo-Paris Basin ; d’Orbigny’s type area of the
Stage. The succession in England is compared in detail with that of northern France by refer-
ence to sections in the Pas de Calais, Meuse, Marne, Haute Marne, Aube, Yonne, Pays de Bray,
and Pays de Caux, several misconceptions being eliminated. The ammonite zonal scheme is
discussed in detail to provide the basis for international agreement of a zonal scheme for the
faunal province, here termed the hoplitinid ammonite faunal province, of which the Anglo-
Paris Basin is but a part. A brief review is made of the faunal links between the hoplitinid
province, covering much of Europe, and neighbouring ammonite faunal provinces. These links
occur essentially in the /yelli Subzone near the base of the Middle Albian, and in the cristatum
Subzone at the base of the Upper Albian. The palaeogeographical implications of the distribu-
tion of the faunal provinces are mentioned briefly. A study of the conditions of Middle Albian
deposition in England shows that it tends to follow the pattern of Lower Albian and Aptian
sedimentation in direct contrast to that of the Upper Albian where a different pattern pre-
vailed. A review of the ammonite fauna terminates the work.

RESUME

L’auteur présente des nouvelles informations sur la stratigraphie de l’Albien moyen de
l’Angleterre et de la France, afin d’établir le plan zonale des ammonites dans le bassin anglo-
parisien, qui est la région typique de l’étage de d’Orbigny. Les couches de 1’Angleterre sont
comparés en détail avec ceux-la de la France septentrionale. L’auteur donne tous les détail du
plan zonale des ammonites, afin d’établir une province faunistique des ammonites hoplitinidés.
L’auteur discute les liens faunistique entre la province des hoplitinidés et les provinces faunisti-
que avoisinantes des ammonites. Ces liens ont bien surtout dans la sous-zone de /yelli auprés

6 MIDDLE ALBIAN STRATIGRAPHY

de la base de |’Albien moyen, et la sous-zone de cristatum a la base de 1’Albien supérieur. Une
étude de la méde de déposition de l’Albien moyen en Angleterre montre qu’elle imite la déposition
de l|’Albien inférieur et Aptien, non pas celle-la de l’Albien superieur. L’auteur finit en passant
en revue la faune des ammonites.

I. INTRODUCTION

TuE Albian Stage terminating the Lower Cretaceous has been divided into Lower,
Middle, and Upper Substages. In England, the Lower Albian is represented within
the top beds of the Lower Greensand and its junction with the overlying clays of the
Gault ; the Middle Albian within the Gault, and by the Lower Gault where this
division is recognisable ; and the Upper Albian by the Upper Gault and the contigu-
ous Upper Greensand. Both the Middle and Upper Albian are represented within
the Red Chalk facies. In northern France and the Paris Basin there is a similar
lithological sequence, excluding a Red Chalk facies but including local lithological
units in various areas such as the Sables de Puisaye flanking the northern area of the
Massif of Morvan. This essentially common sequence reflects the closely comparable
depositional environment which existed in the area of the Anglo-Paris Basin during
Albian times.

This Basin extended from the area of the Massif Central and the Vosges in the
south to the English Midlands (text-fig. 51). It was flanked on the west by the
Variscan massifs of Armorica and Cornubia, and on the east and north east by those
of the Rhine State Mountains and the Ardennes. Late Jurassic—early Cretaceous
deformation provided the basic structural pattern for Lower Cretaceous sedi-
mentation which achieved its greatest geographical extent during Albian times.
Middle Albian sedimentation followed this earlier pattern, but this changed in the
Upper Albian due to tectonic disturbances at its start. The Basin was linked with
the surrounding shelf seas north of the Ardennes, and with Tethys by means of the
Morvano-Vosges strait. These sea-ways provided important migration routes for
the fauna.

In terms of absolute radiometric dates, the Albian is taken by Casey (1964 ; I99)
to commence at 106 my B.P., and is considered to have had a duration of 6 million
years. However, as that author clearly states, this is purely an arbitrary figure, the
Cretaceous being divided into twelve equal time units corresponding to the twelve
Stages. At present, therefore, there is no alternative but to use the relative units of
time implied in the zonal schemes based on the faunal succession. Of these schemes,
that based on the ammonites is the one which has been most thoroughly investigated,
and the one that has led to much difference of opinion both national and international.
A good deal of this disagreement is due to insufficient knowledge of the succession,
despite the considerable number of papers which have been published.

Within the lengthy bibliography on the English Albian two major works stand
out ; the Memoir by Jukes-Browne (1900), and the Monograph by Spath on the
Albian Ammonoidea (1923-43). The work carried out before 1900 in certain cases
was truly excellent. For example, that of De Rance (1868) and Price (1879, 1880)
at Folkestone, Newton (1897) at Okeford Fitzpaine, Dorset, and Keeping (1868) at
Upware, Cambridgeshire. All this earlier work is ably summarised by Price and

IN THE ANGLO-PARIS BASIN |

Jukes-Browne and it is unnecessary to repeat it here. Jukes-Browne’s memoir
included almost all the stratigraphical information then available obtained either by
earlier workers, or by himself and William Hill for the Geological Survey. He
provided a good picture of the stratigraphy of the Albian in England, and extended
his study to make a brief examination of the Paris Basin. The accuracy of some of
his and Hill’s field observations can still be demonstrated, and their reading of
certain sections now long since vanished can be interpreted in the light of recent
information with reasonable confidence.

The first part of Spath’s Monograph appeared twenty-three years after the publica-
tion of Jukes-Browne’s memoir, and it was completed over a period of twenty years.
Its coverage was not complete, however, a fact which it is important to bear in mind.
It consists of the description of ammonite faunas from the comparatively few localities
that Spath himself collected from, or that were represented in museum collections
upon which he largely worked. Other sections, although available during this
period, were not apparently collected from by him and so, important parts of the
ammonite fauna were not described. Nonetheless, this very important work pro-
vided the basis and stimulus for the detailed stratigraphic work that was carried out
during that period largely by officers of the Geological Survey, and the work that
has been carried out since. The final part, published in 1943, contains a useful
review of the stratigraphic work carried out in the period since I9g00. Since the
completion of his Monograph, a great deal of new information has been obtained
about Albian sediments in England. Contributions to our knowledge have been
made by a number of authors, particularly by Casey and C. W. & E. V. Wright.
Their papers and many others are listed in the bibliography and are discussed in the
appropriate part of the text.

In France, before Jukes-Browne’s Memoir was published, Charles Barrois wrote
three important papers on the French Albian (1875a, 1875b, 1878). These, in associa-
tion with d’Orbigny’s Paléontologie Francaise, provided a good picture of the strati-
graphy and fauna of the Albian deposits of the Paris Basin. Barrois followed
Hébert (e.g. 1875a) and others in including sediments, now classified as Upper Albian,
within the Cenomanian. It is significant to note that Barrois already appreciated
the magnitude of the break in the succession between what we now consider to be
Middle and Upper Albian deposits in areas of the Paris Basin some 50 years before
Kitchin & Pringle (1922a) recognised it in England. In his papers Barrois incorpor-
ated the results gained by earlier workers such as Michelin, Leymerie, Raulin, Buvig-
nier, Cornuel, Ebray, Hébert, Delatour, and others, and a comprehensive biblio-
graphy is given by him (1878 ; 230-238). English workers such as Hopkins (1845),
Topley (1868), Jukes-Browne & Hill (1896) and Jukes-Browne (1900) carried their
researches into France, the latter authors providing new information particularly
about the sequence in Normandy described previously by Lennier (1867).

Much useful information was published in the period of 30 years which separated
Barrois’ work from the publication of an important thesis by Jacob (1908). This
thesis greatly increased our knowledge of the Aptian and Albian stratigraphy of the
French Alpine area and adjacent Switzerland, the site of the Morvano-Vosges strait.
In the following two decades, however, only a few papers were published. Of these,

8 MIDDLE ALBIAN STRATIGRAPHY

Lemoine (1910) on the Yonne, Aube, and Haute-Marne, Ciry (1927) on the Céte d’Or,
and Stieler (1922) on the coastal sections at Petit Blanc Nez, may be mentioned. In
the period from 1930 until the war interrupted work, much new information was
contributed by authors such as Breistroffer working in the French Alps (1931, 1933,
1936, 1940), P. & J-P. Destombes at Wissant (1938a) and in the Pays de Bray (1938b),
Larcher (1937), Houdard (1933, 1940) and Marie (1939, 1941a) working in the Aube,
Yonne, Marne, and Haute-Marne. Marie’s work on the Albian foraminifera and
their zonal value included studies of sequences in the Pays de Bray (1941b) and
Wissant (I94Ic).

After the war, and with Spath’s Monograph completed, Breistroffer (1947)
presented an important discussion of the ammonite zones of the Albian in France
and England. Since then papers on various aspects of French Albian strati-
graphy have been published. This more recent work is summarised, and much new
information added, in the report of the Colloque sur le Crétacé inférieur held at
Lyon in 1963 (1965).

Of necessity the foregoing review is very brief, and only the more important works
have been mentioned including those which give comprehensive bibliographies of
earlier work. However, the relevant papers on the Albian of the Anglo-Paris Basin
are discussed at the appropriate place in the text. Recent papers published in
France and England show that disagreement exists on various aspects of the zonal
scheme, and even the litho-stratigraphy. This disagreement is both national and
international ; a serious state of affairs rendering distant correlation difficult (e.g.
Young 1966). It is even more serious when one realises that here we are dealing with
d’Orbigny’s ‘“‘ type area ”’ for the Albian Stage.

An agreed zonal scheme can only be based on a detailed accurate account of the
succession throughout a whole province, and inter-provincial correlation can only
be accurately made once the stratigraphical successions are fully known. The
object of the present work is to attempt to give this detailed information. Its
presentation is the more urgent now that recent geophysical work indicates the
closer proximity of Greenland, North America, and Europe in Albian times (e.g.
Carey 1955, 1958, 1963, Bullard e¢ al. 1965), and the need to be able to compare in
detail sequences in these areas with that of our own.

The first part of this work, therefore, consists of the description and correlation of
sections in England and France, and this contains much new information. The
sections are shown graphically for easy reference. This descriptive part is sub-
divided into convenient geographical areas. The second part consists of a detailed
discussion of the ammonite zonal scheme given in Table 1 (p. 10), preceded by an
historical introduction. This is followed by a review of the links between the Euro-
pean ammonite faunal province and other areas, and in turn by a discussion of the
conditions of depositionin England. The work is terminated by a brief review of the
ammonite fauna, with descriptions of three new species of stratigraphic utility.

II. ACKNOWLEDGEMENTS

This paper is an abridged version of the thesis accepted for the Ph.D. degree of the
University of London. It is a very great pleasure indeed to acknowledge the help

IN THE ANGLO-PARIS BASIN 9

and support which has made the present work possible. In particular, I wish to
thank the late Professor J. H. Taylor, and my supervisor Dr. J. M. Hancock for the
considerable help and hospitality afforded to me as a research student at King’s
College, University of London. Without the facilities provided by the various brick,
tile, and cement companies, by national, regional and local authorities, and by
private landowners both in England and France, all mentioned in the text, it would
have been impossible to carry out the work at all.

The following organisations and members of them have aided the work and to these
my thanks are due: my colleagues in the Department of Palaeontology, British
Museum (Nat. Hist.), in particular, Dr. H. W. Ball (Keeper), Dr. E. I. White (former
Keeper), Dr. M. K. Howarth and Mr. D. Phillips, Dr. R. P. S. Jefferies, Mr. H. A.
Toombs ; the Director of the Institute of Geological Sciences Dr. K. C. Dunham, Dr. V.
Wilson (Assistant Director), Mr. S. C. A. Holmes (District Geologist S.E. District),
Mr. R. V. Melville (Chief Palaeontologist) and Dr. F. W. Anderson, Dr. D. A. Gray
(Water Department), Mr. L. S. O. Morris (Boring Department), Dr. R. Casey, and the
members of their staff for making available outcrop and borehole material ; the Gas
Council, Mr. P. Hinde (Head Geologist), and the late Professor V. C. Illing for per-
mission to examine the Winchester and Cliffe groups of borings ; the Board of
Directors of British Petroleum Ltd., Dr. P. E. Kent (Chief Geologist) and Dr. A. J.
Martin, for making available borehole material and information ; Mr. S. B. Thomas
(Ministry of Transport, Channel Tunnel Division), M. R. Malcor (Channel Tunnel
Study Group), Mr. E. W. Jaccomb Hood (Channel Tunnel Site Investigation), and
Mr. D. J. Carter (Imperial College) ; Mr. A. G. Brighton, Sedgwick Museum ; Brig-
adier G. Bomford, Mr. F. H. Edmunds, and Dr. W. J. Kennedy, Oxford University ;
Miss J. Royston, Buckingham County Museum ; and Mr. H. J. Bick (Central Electric-
ity Generating Board). It is a pleasure to thank Mr. C. W. Wright for the loan of
specimens and information.

My special thanks are due to Dr. P. Destombes for his hospitality, and introduction
to sections in the Pays de Bray and in the Yonneand Aube. M. J. Fourcher (Rouen)
provided access at very short notice to the Bucaille Collection. I am indebted also
to Dr. J. A. Jeletsky of the Canadian Geological Survey and Dr. Keith Young of the
University of Texas.

I am particularly grateful for financial aid from the following authorities : the
Godman Fund, the Educational Authority of the London Borough of Bromley, the
Central Research Fund of the University of London, the G. W. Young Fund of the
Geologists’ Association, and my parents. I wish to thank my wife also for her help
in various ways.

III. DESCRIPTION AND CORRELATION OF SECTIONS

A good deal of stratigraphic information can be displayed to advantage in the form
of diagrams, and so all the lithological sections are presented as text-figures.
Correlation charts, however, cannot provide the necessary accuracy when used alone,
and require an accompanying text. Only the critical ammonites are mentioned in
this section as a more detailed list and its stratigraphical distribution is given later.
For ease of description the account is divided into convenient geographical areas to

10 MIDDLE ALBIAN STRATIGRAPHY

which no structural significance should be attached : depositional controls on sedi-
mentation will also be discussed later. The sections in the Weald are described first,
then the Isle of Wight, Dorset coast, and the outcrop from Devon to the Leighton
Buzzard area of Bedfordshire. The Albian sediments of the Leighton Buzzard area
and of East Anglia are to be described elsewhere, but in no way do they affect the
stabilising of the Albian Zonal scheme. Borehole evidence in southern England is

TABLE I

Ammonite zonal scheme of the Middle Albian adopted here and discussed in detail on pages
118-130.

Substage Zone Subzone
Upper
Albian Mortoniceras inflatum (part) Hysteroceras orbignyt
(part) Dipoloceras cristatum”
Anahoplites daviesi
Euhoplites lautus Euhoplites nitidus
Euhoplites meandrinus
Middle Euhoplites loricatus™ Mojsisovicsia subdelaruei
Albian Dimorphoplites niobe
Anahoplites intermedius
Hoplites (Hoplites) spathi
Hoplites (H.) dentatus Lyelliceras lyelli®
Hoplites (‘ Isohoplites’)
eodentatus
Lower Douvilleiceras Protohoplites puzosianus
Albian mammillatum Otohoplites raulinianus
(part) (part)

(1) Formerly included in the /autus Zone sensu Spath (e.g. 1941 ; 668).

(2) First recognised by Owen (1958 ; 162).

(3) Approximately equivalent to the benettianus Subzone of Spath non De Rance (1868).

(4) In better developed sequences in France (Pays de Bray, Aube), this index fossil does not range up
to the base of the eodentatus Subzone.

then considered before attention is turned to the Paris Basin. A detailed compar-
ison is made between thesections at Folkestone and Wissant (Pas de Calais). Selected
sections in the Meuse, Marne, Haute Marne, Aube, Yonne, and in the Pays de Bray,
are described followed by a comparison of sections in the Pays de Caux with the Isle
of Wight. The ammonite zonal and subzonal scheme employed in this account is
given in Table 1 above.

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30

Fic. 1. Sketch map of the Weald and adjacent areas showing positions of sections and boreholes discussed in the text.

IN THE ANGLO-PARIS BASIN GE

A. Weald

The Gault describes a narrow outcrop at the foot of the Downs from Folkestone in
the east, around the northern, western and southern borders of the Weald to reach
the sea again at Eastbourne (text-fig. 1). Deposits of Middle Albian age are present
throughout and reach their greatest known thickness in Sussex. No section has yet
provided a complete and relatively uncondensed sequence. Although the greatest
degree of representation is to be found in the Folkestone area of Kent and between
Steyning and Ringmer in Sussex, even at these localities condensation at certain
horizons is very marked. Fortunately, further west in the northern Weald some
condensed horizons are greatly expanded, but this is offset by truncation westwards
of deposits of the higher subzones of the Middle Albian due to tectonic movements and
associated erosion within the cristatum Subzone.

(i) FOLKESTONE

In response to the general north-easterly dip, the Gault appears above the Folke-
stone Beds at the top of East Cliff, where it overlooks the harbour, and declines to the
shore in East Wear Bay. Copt Point is the promontary at the NE. end of East Cliff
at the entrance to East Wear Bay, and it was shortly after a substantial cliff-fall in
1959 that the section given in text-fig. 2 was measured. By far the bulk of the
fossils, for which the Folkestone Gault is famous, were collected from the foreshore
exposures in East Wear Bay (East Weir Bay of early authors), but due to cliff and
shore stabilising work carried out by British Railways, these sections are now hidden
beneath beach-sand (see Bisson in Smart ef al. 1966 ; 293-296). It is worth recording
that in the period between 1948 and 1956 when remedial work obscured the sections,
a large slice of Gault extending from the top of Bed III to the base of Bed X could be
seen from the area now covered by the western half of the Toe-weighting to about 100
yds west ofit. This slice was not greatly disturbed within itself, but at the line shown
by Bisson (Smart et al., 1966 ; fig. 17, p. 294) the dip was in the order of 70°—80°
landwards.

The Folkestone Gault has attracted the attention of many workers because of the
intrinsic beauty of its fossils. The early history of research is ably summarised by
Price (1879, 1880) whose bed notation is still broadly used today. De Rance (1868),
however, was the first to divide the Gault into Lower and Upper Divisions and to
subdivide them further into eleven Beds. Price (1874, 1875) accepted De Rance’s
Upper and Lower Gault and also recognised eleven Beds but these did not coincide
with those of De Rance. De Rance later (im Topley 1875 ; 146) accepted the bed
notation of Price but they do not coincide lithologically. Jukes-Browne subse-
quently modified Price’s account, and it is his reading of the section which has been
accepted by subsequent workers (1900 ; 69-83). Spath (1923-43) drew heavily on
the well preserved ammonite fauna of this locality, and indeed, the degree of repre-
sentation in his Monograph is largely a reflection of the high degree of representation
within the section itself. Spath (1923a, b, c ; 1926b), like Jukes-Browne (1900 ; 45),
expressed his zonal scheme for the Middle and Upper Albian essentially in terms of
the lithological sequence at Folkestone.

N

MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ft Ins M
§
(iii) Seam of phosphatic nodules and many partly phosphatised fossils in grey clay = 34
with crushed shells. ~
VIII[G) Grey clay with shells mainty of Inoceramus sulcatus. 2 10-11
(i) Seam of black phosphatic nodules in grey clay. i 2 10
a
2
>
J
»
9
VII | Grey clay with shell seams containing pyritised fossils and scattered °
phosphatic nodules.
4
3 8
=
=
c
VI Mottled lignt-grey clay with numerous dark grey burrows; occuring as fo)
irregular tenticles in dark grey clay.
Vv Midgrey burrowed clay with occasional scattered brown phosphatic nodules, 67 7
iii) Abraded biack phosphatic nodules, often fragments of ammonites,in grey clay. 1-2
IV } fui) Bark grey shelly clay. 4
Gi) Seam of bivalves, part-phosphatised fossils,and buff phosphatic nodules. va 1
6
a fe)
| II Fawn-grey clay with shell seams and partly phosphatised fossils, Occasional &
lenticles of ferruginous merlstone occur, s
5

(iv) Clay like II(ii) passing at the top by lenses into Bed ill. 11

(iii) Seam of blackisnh phosphatic nodules in dark grey clay.

(ii) Dark-grey clay, black when wet,with seams of crushed shells and scattered
osphatic nodules.
crushed & part

(vii) Dark-grey slightly silty shelly clay.

(vi)Gritty grey clay becoming more glauconitic and gritty downwards;
sparsely tossiliferous and with occasional thin seams of phosphatic ‘clots”

spathi

(v) Large phosphatic nodules mainly casts of ammonites in shelly glauconitic
loam — ‘dentotus nodule bed?

(iv) Highly glauconitic loam with a few blackish atic nodules.

(ii) Scattered spherical septarian phosphatic nodules.

lauconitic loam with scattered lumps of blackish phosphate and a few
osphatic nodul h

<- 3 ord
intermedius
|
n nw =
ny w +

a lyelli

ent.

.

10

Fic. 2. Cliff section of Lower Gault a few yards NE of the sewer, Copt Point, Folkestone,
Kent.

IN THE ANGLO-PARIS BASIN 13

Since Spath completed his Monograph, our knowledge of the Lower Gault at
Folkestone has been increased by Casey (1950) and the author (Owen 1958 ; 1963a).
Bisson (in Smart, Bisson & Worssam 1966 ; 56-58) has given an account of the
section, but this is based essentially on Jukes-Browne. The revised section of the
Lower Gault, given in part by me in 1963 (1963a ; 36-38) and in full in text-fig. 2, is
the first new account since Price (1880). It is slightly thicker than that of Price, but
this is probably due to his section being further out from the modern cliff-line, the
seaward slope of the land at Copt Point producing a slightly thinner sequence due to
creep at the cliff of Price’s day.

Casey (1950 ; 272-3) divided Bed I into four sub-divisions corresponding to (i) the
‘Sulphur Band’, (ii) the ‘Greensand Seam’, (iiia) the dentatws nodule bed, and
(iiib) the remainder of the clays. He classified the whole of the ‘ Greensand Seam ’
with the inaequinodum Subzone, later (e.g. in Worssam et al. 1963 ; 59) replaced by
the index Hoplites (Isohoplites) eodentatus. The dentatus nodule bed and the lower
part of the overlying clays were, following Spath, classified with the spathi Subzone,
and the upper part of Bed I with the intermedius Subzone. He concluded that the
benettianus Subzone (i.e. the lyella Subzone) was probably absent (see also 7» Worssam
et al., 1963 ; 59).

Subsequent work has shown that Casey’s subdivisions are too broad and a more
detailed notation is given by me (Owen 1963a ; 36-38, 49) and in text-fig. 2.

Fossils are rare in the ‘Greensand Seam’. The specimens of Hoplites (Isohoplites)
in the Institute of Geological Sciences (GSM 83165-6 Spath Coll.) are angular frag-
ments like the nodules of I(ii) which is here classified with the eodentatus Subzone.
Bed I(iii) has not yielded fossils to me, but in I(iv) at a depth of 2 inches (0-05 m.)
beneath the dentatus nodule bed, I have collected, apart from the specimen of
Hoplites (H.) cf. bayle: Spath figured by me (1963a ; 37, pl. 3, fig. 3), a fragment of
Beudanticeras sp. indet. together with one specimen of Neohibolites minimus (Miller)
and a few indigenous and very delicate bivalves. The association of Hoplites
(Hoplites) and Beudanticeras indicates the lyell1 Subzone. From a knowledge of the
lyella Subzone successions at Small Dole (p. 35), Swindon (p. 61) and the Aube (p. 91),
the writer is firmly convinced that the specimens of Hoplites (H.) aff. benettianus
which occur rarely in the spherical phosphatic nodules immediately beneath the
dentatus nodule bed (Owen 19632 ; 37, pl. 3, figs. 1a, b ; 2a, b) are of late lyelli Sub-
zone age. The presence of this Subzone at Folkestone is not surprising because it is
well represented in the Gault of the Guilford (Waldeshire) Colliery shaft (p. 76) about
7 miles (11-25 km.) from Copt Point, and in the Aycliff boring (p. 78).

The classification of the remainder of Bed I has been discussed by the writer
already (Owen 1963a ; 37-38, 49), and it is only necessary to question one statement
made by Casey (tn Hancock 1965 ; 247) that Anahoplites intermedius comes in a foot
or two above the spathi nodule bed as a minority element in a fauna still dominated
by Hoplites. If this were the case, then the Folkestone section would be unique.
Species of Hoplites (H.) occur crushed in Bed I (vi) up to a height of 3 feet 2 inches
(0-96m.). Higher up, ammonites are absent or very rare until one reaches the base of
I (vii), at which level Anahoplites intermedius appears. At Folkestone, as at most
other localities in the Weald, there is a gap in the ammonite fauna between un-

I4 MIDDLE ALBIAN STRATIGRAPHY

doubted deposits of spathi and intermedius Subzone age. There is no break apparent
in sedimentation, but the only common fossils are large Inoceramus concentricus. At
localities such as Petersfield (Sussex p. 34), Osmington (Dorset p. 51), Devizes (Wilt-
shire p.60),andtoapointalsoin thesouthern part of the Paris Basin (pp. 88,93, 97) this
gap in the Weald sequence is filled. There is indeed the association of Anahoplites
and Hoplites (H.) but the forms of Anahoplites are very distinct and include A. grims-
dalei sp. nov. and A. osmingtonensis sp. nov. ; forerunners of the evolutus-intermedius-
praecox group. Hoplites (H.) continues on into the intermedius Subzone but is a very
subordinate element in the fauna.

The classification of the remainder of the Lower Gault at Folkestone by Spath has
seen only a few modifications (Owen 1958 : 1960 ; 376-7). Spath pointed out (1926b;
421) that the divisions between Subzones do not always coincide with the junction
between beds. The details of Bed I (vi—vii) given by the writer in 1958 were taken
from what are now obviously disturbed sections, but were then the best available,
and should be discounted. The account given in 1963 was taken from a perfect
section. Bed I (vii) and the whole of Bed II up to a level 3 inches (0-70 m.) below
its top, are classified with the intermedius Subzone. The bulk of the ammonites are
crushed and consist essentially of species of Anahoplites such as A. intermedius,
A. praecox,and A. mantelli. Partly phosphatised ammonites occur in Bed II (i) and
pyritic specimens occur very sparingly among the numerous crushed fossils in II (ii).
The fauna of these three subdivisions is very uniform, but in II (iii) we see the intro-
duction of a particularly coarse development of A. praecox, and in II (iv), where
‘solid’ pyritic fossils are more common, Ewhoplites pricet Spath is a very character-
istic form.

Dimorphoplites niobe Spath, already present in the imtermedius Subzone, becomes
common in the shell seam about 3 inches (0-076 m.) below the base of Bed III at which
level Anahoplites of the praecox-intermedius group quite suddenly ceases to be im-
portant. D. mobe, together with A. planus and A. splendens then characterises the
whole of Bed III, within the upper 2 to 3 inches (0:051-0-076 m.) of which, part or
wholly phosphatised fossils with the nacreous shell occur sparingly. Bed IV (i) does
not mark a great break in the sequence although semi-derived phosphatic fragments
of fossils do occur. D. niobe, A. planus and A. splendens still occur but the fauna
becomes more diversified and Mojsisovicsia subdelaruei M. remota and M. spinulosa
(Spath) appear as infrequent but highly characteristic species. These species of
Mojsisovicsia occur indigenous in both IV (i) and (ii) which are classified with the
subdelaruet Subzone (Owen 1960 ; 376). Bed IV (iii) marks a greater period of
erosion and its characteristic ammonite fauna was listed by Owen (1958 ; 157). This
bed, together with the basal 2 inches (0-051 m.) of Bed V which contains the same
fauna is classified with the meandvinus Subzone. Casey (in Smart, Bisson & Wor-
ssam 1966 ; 109) and in effect Milbourne (in Hancock 1965 ; 247) state that Euhoplites
meandrinus Spath occurs outside Bed IV (iii) and the basal part of Bed V. However,
the author has never seen a specimen, either in the field or in the collections, from any
other bed. Spath’s original (BMNH, C 32306) was a pyritic specimen, now decom-
posed, which with very little doubt came from the basal part of Bed V, as Spath
thought likely (1930b ; 271).

SUBZONES

E STREET WYE BRABOURNE — SELLINDGE SANDLING FOLKESTONE) cristatum
PADDLESWORTH A249 CHRISMILL = EYHORN : Par fieseaies
STED DUNTON GREEN ST JOHN’S SEVENOAKS — FORD PLACE DLE: A249 SSI ss
ts > TANDRIDGE SQUERRYES BRASTED senna 6 peace, cooeest] 6 leegsees @ 5 ta ee aan ica =daviesi a 7——? 2
2——— fesrea, 6 v= pease v Ren PE ni. Ar ieee : ‘ jesi
Bell I eee 5") —} O00 ie (apa feliii) Fe ci) ——— i) 2 2 daviesi
F “|s@ [onan sa ———

" j ?7—___, a
—— —————t (J
ee + eet C(ii) : ? ————— ——? —————?
‘soa: fracaand 1 eee)

nitidus

? a2]
4 : rp
V 4+5 MSS 50) hus 5 (i)
aN oe OD) A
cu

?
a ov ? > 4 i
P ; ; [oe = —-, ? ?7——__2
| / PS ee Ly ae ee ran EH aes foe ? ? ? 2 ? ?
L faeaeaeal hess —~ SS a 3 ee B
| P ING SE

intermedius

Subzonal correlation lines

Lithological correlation lines

Fic. 3. Correlation of sections in Kent and east Surrey.

IN THE ANGLO-PARIS BASIN 15

The sudden change from a deeply sulcate to a channelled venter in Euhoplites which
occurs at a height of 2 inches (0-051 m.) up from the base of Bed V was taken by the
writer to mark the base of the mitidus Subzone and the lautus Zone (Owen 1958 ;
157-8, 162). There are a few scattered phosphatic nodules at this level and a small
break in deposition is further suggested by the sequence at Ford Place, Trottiscliffe
(p. 22). The remainder of Bed V together with Bed VI are classified with the mitidus
Subzone and yield a typical fauna. Apart from the highly burrowed nature of the
clay, Bed VI is also characterised by particularly tuberculate examples of Dimor-
phoplites of the parkinsoni type. The lower part of Bed VII, previously classified
with the daviesi Subzone, in fact still contains a nitidus Subzone fauna and must be
classified with that Subzone (Owen 1960 ; 376). Anahoplites daviesi does not appear
in the sequence until a height of about 5 feet (1-524 m.) is reached, and this and
closely related forms characterise the remainder of these clays of Bed VII which are
classified with the davies: Subzone.

Bed VIII at the summit of the Lower Gault was classified by Spath and subsequent
authors with the cristatum Subzone here included in the Upper Albian (inflatwm
Zone). The depositional history of Bed VIII is complex. The lower nodule bed
VIII (i) represents a moderate break in deposition, and its fauna includes the bivalves
Inoceramus concentricus, I. sulcatus subsulcatus and I. sulcatus, and the ammonites
Dipoloceras bouchardianum (d’Orbigny) and Beudanticeras beudantt (Brongniart).
At Wissant (p. 85) on the French coast 22 miles from Folkestone, the equivalent of
Bed VIII (i) is represented by the clays of Bed 12 (v) in which a coarse form of J.
concentricus at the base soon passes into the subsulcatus stage to achieve the sulcatus
form at the top. This bed also yielded the holotype of D. bouchardianum and B.
beudanti also occurs. It is, however, apparent that some material of late daviesi
Subzone age is also present in the remanié fauna of Bed VIII (i) at Folkestone. This
element can be demonstrated by the occurrence of very coarse developments of
Anahoplites of the davies: group and typical I. concentricus.

A detailed discussion of Bed VIII and its fauna is out of place here but it is
essentially of cristatwm Subzone age. The uncondensed sequence at Wissant indicates
that the incoming of the typical fauna of the cristatwm Subzone was quite rapid. This
fauna continues on into the basal few feet of Bed IX which will also have to be
classified with the crvistatum Subzone. Bed VIII is in all truth a junction bed as the
early workers recognised.

(u) FOLKESTONE TO THE MEDWAY

No complete sequence in the Lower Gault has been seen between Folkestone and
Chrismill Lane, Thurnham, on the Maidstone By-Pass, a distance of 29 miles (46-67
km.). What little information is available is ably presented by Smart, Bisson &
Worssam 1966 (Folkestone to Westwell), Worssam ¢é¢ al., 1963 (to a few miles W. of
Maidstone), and Dines, Holmes & Robbie 1954. A little additional information is
given here, and the correlation of the sections is shown in text-fig. 3.

It is apparent that the sequence seen at Copt Point has changed already by
Elenden Gardens, Cheriton (Spath 1923c ; 141-2) but unfortunately no precise

16 MIDDLE ALBIAN STRATIGRAPHY

details of this section were recorded. Bisson im Smart e¢ al. (1966 ; 100) has described
the Folkestone Brickworks section at Cheriton (TR 205376) in which I suspect that
the lowest 2 feet 6 inches (0-762 m.) recorded as ‘ Dark-grey slightly micaceous
blocky clay with occasional phosphatic nodules’, underlying the basal cristatum
nodule bed, is of daviest Subzone age as at Copt Point, and at Wye near Ashford.
However, I did not see this part of the succession.

At Sandling Junction (text-fig. 4) the fauna of Bed 9 classified with the spathi
Subzone is identical to that of Division A in the Maidstone By-Pass (Owen 1960 ;
372), and so, the age of the dentatus nodule bed has already changed at the outcrop
within 5 miles (8-05 km.) of Copt Point (Owen 1963a; 49-50). No fossils have yet
been found in the underlying glauconitic loams of Beds 4-8. Bed 8 (of Worssam
1966 ; 99) at File’s pit in Swan Lane, Sellindge (TR 11853915) is the equivalent of
Bed 9 at Sandling, and Bed 6 yielded the specimen of Hoplites ? recorded by Worssam.

Bed Lithology

Weathered grey clay. ' |

Phosphatic nodules mainly fragmentary casts of Hoplites(H.),

Grey slightly glauconitic clay.

Scattered phosphatic nodules in grey slightly glauconitic clay. —_—

|
Mottled yellow, green, grey, glauconitic loam becoming more !
argillaceous upwards.

?eodentatus—lyelli spathi

Shattered ferruginous phosphatic nodules in loam.

Mottled glauconitic loam with occasional phosphatic nodules.
Shattered fterruginous phosphatjc nodules in grey loam.
Dense band of phosphatic nodules in lenticular cemented grit, |

puzosianus

Yellowish sand with small scattered phosphatic nodules

Fic. 4. Section in Gault-Lower Greensand junction beds at Folkestone Quarries Ltd’s
Sandling Sandpit, c. 150 yds. NW. of Sandling Junction railway station, Saltwood, Kent
(TR 14703690).

This specimen (GSM., CW 855) is a definite Hoplites (H.) but there is no other
indication of the age of the sediments overlying the puzosianus Subzone sediments.
The principle bed of phosphatic nodules in the spathi Subzone sediments at the
Granary Court Sand pit at Brabourne (TR 09004005) is also on the same horizon as
that of Bed 9 at Sandling. At Brabourne, in the area where Hill (in Jukes-Browne
1900 ; 84) records a section, the former Ashford & Naccolt Brick, Tile & Potteries
Ltd., dug shallow pits in higher beds of the Lower Gault about 800 yards (731 m.) a
little north of west from Park Farm (TR 08904065). About 8 feet (2-43 m.) of
weathered brownish clay streaked with ferruginous matter was to be seen beneath
superficial deposits. Towards the top of the clays a disturbed seam containing
phosphatic nodules yielded fragments of the following :—

Anahoplites planus (Mantell), A. pleurophorus Spath, Dimorphoplites aff. niobe,
D. cf. doris Spath, Euhoplites cf. subtuberculatus Spath late mutation, E. microceras

IN THE ANGLO-PARIS BASIN 17

Spath late mutation, Hamites tenuicostatus Spath, Inoceramus concentricus Parkin-
son.

The assemblage indicates the subdelaruei Subzone as restricted here and by the
writer in 1960, which coincides with the term ‘ lower subdelaruet Subzone’ given by
Smart (im Smart, Bisson & Worssam 1966 ; 98). The age of the underlying clay was
not determinable, but the whole sequence is similar to that of the Maidstone By-Pass
(text-fig. 3). Smart records cristatwm Subzone fossils from a similar shallow working
300 yds (274 m.) to the NNW.

In the Ashford Brickworks Ltd pit (formerly the Ashford & Naccolt Brick, Tile &
Potteries Ltd) situated 700 yds NW. of Sillibourne Farm and about 110 yds ENE. of
Blackwall Farm, Wye (TR 04954445) the upper beds of the Lower Gault have
recently been exposed (text-fig. 5). This is the pit recorded by Cornes (7m Dewey
et al., 1925 ; 263-4) as “ New Nackholt’, and by Smart at TR 049445 (1966 ; 98).

Bed Lithology Ft Ins M

Dense seam of phosphatic nodules and phosphatic tragments of fossils.

(iv) Dark grey clay shelly witn part phosphatised fossils particularly inthe
basal inch.

(iii) Dark grey clay with few fossils and occasional scattered phosphatic
nodules,

Gi) Bed of scattered brownish phosphatic nodules in dark grey clay.
(i) Dark grey clay.

?nitidus daviesi cristatum

~~ — => <— ¢—_—

seen to

Fic. 5. Upper beds of Lower Gault at Naccolt Brick Works (Ashford Brickworks Ltd),
700 yds NW. of Sillibourne Farm and c. 1100 yds ENE. of Blackwall Farm, Wye, Kent

(TR 04954445).

Anahoplites daviesi and A. daviesi ornata appear at a depth of ro inches (0-254 m.)
below bed 2, the cristatum nodule bed, and the remainder of the clay up to bed 2 is
classified with the daviest Subzone. Whether the lower part of the exposed clays
of bed 1 is also of daviesi Subzone age is not certain. Only a single nodule bed
(bed 2) here represents Bed VIII at Folkestone and its character and fauna is almost
identical to that of Division D in the Maidstone By-Pass (Owen 1960 ; 374). The
section recorded by Hill (7m Jukes-Browne 1900 ; 84) at Kennington, the ammonites
from which were re-determined by Casey (im Smart et al., 1966 ; 97), apparently
showed some Lower Gault, but it is at present impossible to interpret the sequence.
Unfortunately the sections recorded between Wye and Hollingbourne are in-
sufficient to determine the succession. The preservation of the ammonites from
Westwell Leacon and Kennington preserved in the Institute of Geological Sciences
suggests nodule beds about the same horizon as beds C (ii) and D in the Maidstone
By-Pass. The pit at Eyhorne Street, Hollingbourne, described by Hill (im Jukes-
Browne 1900 ; 85) was alleged to show beds spanning the Lower Gault-Upper Gault
junction. However, Casey (in Worssam et al., 1963 ; 59) has re-determined the fossils
collected from this pit and demonstrated that the section was wholly in the Lower
Gault and that Hill’s reading of it was incorrect. If one compares the section given

B

18 MIDDLE ALBIAN STRATIGRAPHY

Div. | Lithology § Ft Ins
So
Dense seam of blackish phosphatic nodules and fossils in 4
bluish grey clay. Nodules tend to divide Into two seams. S 3-4
a
3
(iil) Bluish-grey clay becoming darker upwards, with brownish |=
partings and pockets of limonite after pyritised fossils. =
Occasional phosphatised fossils occur with the shell WY 4 °
especially ata level 4inches below the top. A scattered =
line of small buff phosphatic nodules occurs Sinches above b
the base. 2
€ e . . . .
. e — —'
(il) Bed of chocolate-brown phosphatic nodules in dark fawnis Sud ae are a tee
jet grey: clay. ° 3
(1) Darkish, rather fawn-grey clay about 1 foot in thickness
with shell seams and scattered phosphatic nodules, 6 0

underlain by dark-grey shelly clay passing down into
light speckled fawn-blue-grey clay with sporadic
lenticles of ferruginous marly clay and marlstone near
and at the base.

niobe

(ii) Dark blue-grey clay with shell seams, lightening In 2 2
colour towards the top.

ee ses - @ea+ Pe ae — 2
(it) Seam of irregular—shaped phosphatic nodules.
”
3
v
a
€
x
a
&
(i) Grey slightly fawn clay with many shell seams and

bands of burrowed clay passing down into three feet

three inches of greenish grey clay becoming progressiv-

ely darker and more glauconitic downwards The clay

is blocky with yellowish partings and contains seams

of scattered incipient buff phosphatic nodules at °

heights of 2feet Binches, 3 feet Qinches, 4 feet 6

inches, 6feet 3inches and 8 feet 8 inches.
<
J
Q
”

Bed of blackish pho gnats nodules, some septarian,!n highlyX\
glauconitic clay with some crushed fossils.

Glauconitic sandy clay, becoming sandier below.

Piyelli’ :

Fic. 6. Lower Gault sections at NW. quadrant of A 249 clover leaf, Maidstone By-Pass
(East Section) M 20, extending from a point 585 yds NE. of the Chiltern Hundreds public
house to the A 249, Boxley, Kent (TQ 77805745).

IN THE ANGLO-PARIS BASIN 19

by Hill with that of the Maidstone By-Pass (Owen 1960 ; 369-371 and text-fig. 6
herein) it can be seen that the sequence exposed at Eyhorne Street included parts of
the equivalent of Divisions B and C which appear to be identical to that of the By-
Pass.

There is nothing further to add to my account of the sections exposed during the
construction of the Maidstone By-Pass motorway (M20) (Owen 1960), except for an
unfortunate omission of a complete sentence in my reply to the discussion of my
paper by Gray (1962 ; 469). The missing sentence read, ‘ As seen in the sections, the
Tertiary faulting affects the beds [in the A 249 clover-leaf] in the following manner.’
Without this sentence there is the implication that the faulting is intra-Albian which
it certainly isnot. The correlation of the sections is shown in text-fig. 3, and it can
be seen that the stratigraphical succession is very uniform.

(ii) MEDWAY TO TROTTISCLIFFE
(a) Paddlesworth

West of the Medway, the outcrop changes direction through an arc of about 12° but
no complete sequence is seen until one reaches the Ford Place clay-pit, Trottiscliffe,
where the Middle Albian sequence is thicker than at the Maidstone By-Pass. How-
ever, in the early months of 1968 the Associated Portland Cement Manufacturers
clay pit at Paddlesworth, which has for many years shown an important Upper
Gault sequence, was cut further south exposing the higher part of the Lower Gault.
The Middle Albian sequence is shown in text-fig. 7, and its correlation with the
Maidstone By-Pass and Ford Place in text-fig. 3. The basic similarity with the
sequence at Ford Place from the middle of Division 4 permits the use of the same
divisional enumeration.

By analogy with Ford Place, the 3 feet 6 inches (1-067 m.) of clays at the top of
Division 4 at Paddlesworth are almost certainly of swbdelaruet Subzone age. They
contain numerous Inoceramus concentricus and Hamites tenuicostatus as at Ford
Place, but no specimens of Mojsisovicsia are yet to hand. These clays represent an
expansion of nodule bed C (ii) at the Maidstone By-Pass. Bed 5 (i) contains a
typical meandrinus Subzone fauna and the sediments up to the top of 5 (vii) are also
classified with this Subzone. 5 (i) corresponds exactly to 5 (i) at Ford Place but does
not contain phosphatised fossils with the shell preserved as at the latter locality.
5 (ii-vi) correspond to 5 (ii-iv) at Ford Place and 5 (vii) is the direct equivalent of
5 (v). In general the sequence is slightly thinner at Paddlesworth than at Ford
Place, but it is much thicker than that seen at the Maidstone By-Pass where the
equivalent sediments are but 2 feet 7 inches (0-787 m.) thick.

The sediments of Jautus-Zone age at Paddlesworth 5 (viii) are 2 inches (0-051 m.)
thicker than at Ford Place 5 (vi), and 1 inch (0-025 m.) thinner than at the Maidstone
By-Pass. Part phosphatised fossils occur at the base of 5 (viii) and the species of
Euhoplites indicate the nitidus Subzone, although the commonest ammonite is
Dimorphoplites biplicatus (Mantell). Crushed Anahoplites planus occur in the top
few inches, and there are a few uncrushed Euhoplites truncatus in the uppermost inch.

20 MIDDLE ALBIAN STRATIGRAPHY

Whether the daviesi Subzone is represented at this height is uncertain. Sediments
of daviesi Subzone age are 4 inches (0-101 m.) thick in the Maidstone By-Pass.

The nodule bed Division 6 is of cristatum Subzone age, and it is interesting to note
that there is a tendency for this bed to divide a little into two, a feature seen in
Division D in Sections 2 and 4 of the Maidstone By-Pass (Owen 1960 ; 371). There
is no question of there being two distinct nodule beds, but the feature is of interest in
connection with the tectonic disturbance and associated erosion of the upper surface
of the Lower Gault in cristatum Subzone times (p. 72).

—
Div. Lithology 2 Ins M.
6 Brownlsh phosphatic nodules tending to divide into two concentrationg 2 4-9 3
all in shelly medium-grey burrowed clay. 2
(vill) Mid-grey shelly clay with fawn patches- some extensively 3
| burrowed; scattered phosphatic nodules and part phosphatised BS 6
fossils at base, tc
Wii) Highly burrowed light fawn-grey clay, shelly and with lenticles 1
of maristone at the base.
5 3 2-5
vi Mid-grey burrowed shelly clay. ey lI ~ 2 =
Vv eam of scattered phosphatic nodules. Sea |
iv. G mid-grey clay. os is
i) Buffish phosphatic nodules in mid-grey clay. c 1
o
e 8
(li) Shelly mid grey clay with lighter patches. E
(i) Brown phosphatic nodules, mainly casts of fossils in shelly clay~ : 1-1
‘o
3
i
4 Lightish fawn grey clay, shelly, becoming darker below and burrowed] © S
«
»
a2
J
“”
0

Fic. 7. Section in Lower Gault at southern side of the Associated Portland Cement
Manufacturers’ Holborough clay pit, 880 yds SE. of Paddlesworth, Snodland, Kent

(TQ 69156165).

(b) Trottiscliffe

The sequence exposed in the Rugby Portland Cement Co’s Ford Place Clay pit
extends from the middle of the Folkestone Beds up to the varicosum Subzone of the
Upper Gault. The Middle Albian sediments are shown graphically in text-fig. 8.
Casey (1959) has given a brief account of the whole section, and a detailed account of
the Gault-Lower Greensand junction beds (1961a ; 545). The upper part of the
junction beds and the Gault have been described by Milbourne (1963) but his account
is inaccurate both in lithological detail and in its subzonal classification. The writer
has described the sequence in the spathi Subzone (1963a ; 38), but in view of Mil-
bourne’s account of this section it is necessary to redescribe the sequence in the
Lower Gault (text-fig. 8).

Bed 9 of Casey (1961a ; 545) was stated to have a thickness of 3 feet (0-914 m.) and
by Milbourne (1963 ; 58) as 5 feet (1-524 m.) : it is capable of subdivision (text-fig. 8).
Bed 9 (ii) has yielded pyritic Hoplites (H.) spp. and Beudanticeras sp. and I follow
Milbourne in classifying this horizon with the lyelli Subzone. Whether the eodentatus

IN THE ANGLO-PARIS BASIN

Div. Lithology
[6 | Dense seam of phosphatic nodules and fossils in shelly grey clay. |
(vl) Dark grey shelly clay with partly phosphatised and partly pyritised fossils. i
(v) Mottled Bed. Light fawn grey clay with numerous burrows of dark grey clay.
A shell seam with scattered phosphatic nodules marks the base, Scattered
|_smal! phosphatic nodules occur at top. ——
5
(iv) Dark grey shelly clay.
(iii) Shell seam with scattered smal! phosphatic nodules,
(ii) Dark grey shelly clay, a little burrowed, with a few scattered small
phosphatic nodules and some partly phosphatised fossils,
(i) Bed of large Irregular phosphatic nodules and occasional part-phosphatise: n
in dark grey clay.
Fawn grey clay with shell seams and partly phosphatised fossils more
A ferruginous and marly at the base which is marked by lenticular developments
of ferruginous maristone. ‘i
uN

v

nitidus cristatum

meandrinus

subdelarue/s

niobe

10

or

into Division 4,

—

(v)Irregular line of phosphatic nodules in dark grey clay.

(iv)Dark grey shelly clay.

(vi)Lighter grey clay than below, and more marly, passing in the top few inches

(iii) Seam of scattered phosphatised ammonite body chambers,

(ii) Darkish grey clay becoming increasingly shelly upwards, and lighter
in colour.

(i) Dark gritty grey clay with seams of whitish phosphate incipiently
odular,

Seam of black phosphatic nodules and mainly fragmentary casts o
and other fossils in gritty dark grey clay.

Gii)Dark grey gritty clay with numerous crushed fossils

f ammonited.

) Dark grey gritty clay.
(ili) Highly glauconitic green loam with scattered septarian phosphatic nodules.

(ii) Highly glauconitic dark grey-green clay with a few pyritic ammonites.

(i) Highly glauconitic dark grey-green loam with patches of grey clay

septarian phosphatic nodules occur at 2and10 inches above the base,

puzosianus Subzone sediments

1 (ii) Bed of pale soft phosphatic nodules and partly phosphatised Hoplites (H.) LG:

ill-graded and pebbly in basal 6 inches. Thin seams of scattered spherical

intermedius

spathi

lyelli

Peodentatus

10

3
Se Me Oe
es
i

e
ry alla
.

2

° . .

rites

10—

=|

8-4

34

o—

21

Fic. 8. Section in Lower Gault at the Rugby Portland Cement Co. Ltd’s Ford Road clay
pit, 450 yds N. of Ford Place House, and c. 1100 yds SSW. of Trottiscliffe on W. side of

Ford Road, Trottiscliffe, Kent (TQ 63605910).

22 MIDDLE ALBIAN STRATIGRAPHY

Subzone is represented in g (i), or whether 9g (ili) is of lyelli Subzone age, in the
absence of fossils cannot be determined at present. The Lower Gault can be divided
into six broad lithological divisions corresponding to those recognised by the
author in the Sevenoaks area (1958 ; 152 and text-figs. 3, and 9g herein), and the
correlation of these with Paddlesworth and the four divisions seen in the Maidstone
By-Pass is shown in text-fig. 3. Division 1 (i) has not yielded fossils, but 1 (ii) to the
top of 3 (i) are classified with the spathi Subzone. Division 1 (ii—iii) contain the same
species of Hoplites (H.) that occur in the dentatus nodule bed at Folkestone. Division
2 is the ‘ upper dentatus-spathi nodule bed’ and contains the same ammonites that
were recorded by the writer from bed 4 in the Lower Gault of the Buckland Sand &
Silica Co. pit (Owen 1958 ; 151).

No ammonites have been obtained from Division 3 (i), but at the base of 3 (ii)
crushed Anahoplites intermedius and A. praecox appear in the succession, marking the
base of the intermedius Subzone sediments, and range up through the remainder of
Division 3. The coarse development of A. praecox known to occur in Bed II (iii) at
Folkestone is found in the condensed bed 3 (iii), and Euhoplites price: ranges through
3 (iv) to the base of Division 4 (Milbourne 1963 table 1). The lower 4 feet 2 inches
(1-321 m.) of Division 4 contains crushed examples of Dimorphoplites niobe, and these
sediments are classified with the nzobe Subzone. As Milbourne has demonstrated,
Mojstsovicsia subdelaruer and its contemporaries range throughout the remainder of
Division 4. The meandrinus Subzone is represented within the sequence from
Division 5 (i) to the top of 5 (v). The clays of 5 (1) to 5 (iv) contain the same fauna as
Bed IV (iii) and the basal 2 inches (0-151 m.) of Bed V at Folkestone (Owen 1958 ;
157). However, 5 (v) was probably deposited at Trottiscliffe during a minor phase
of non-deposition at Folkestone. The marked change in the ventral aspect of
species of Ewhoplites which occurs suddenly 2 inches (0-051 m.) above the base of
Bed V at Folkestone is not so abrupt at Trottiscliffe. In Division 5 (v) the peripheral
aspect is transitional from the sulcate to the channelled state. Nonetheless, the
characteristic species are more closely allied to the meandrinus Subzone rather than
the nitidus Subzone.

Division 5 (vi) contains a typical nitidus Subzone fauna but at the top, immediately
beneath the cristatum nodule bed (Division 6), there are to be found the occasional
part-phosphatised examples of Anahoplites daviest. Division 6 contains numerous
usually fragmentary fossils in a matrix containing crushed or partly phosphatised
fossils of cristatum Subzone age. There are well preserved phosphatised ammonites
with the shell which are of daviesi or nitidus Subzone age which have just been caught
up into the phosphatic debris of the nodule bed from the clays beneath. Nonetheless,
the clay sediment of Division 6 is of cvistatum Subzone age. The phosphatic debris
contains ammonites indicating the daviest Subzone, but essentially the cristatum
Subzone.

At Trottiscliffe, therefore, as at Paddlesworth at least in part, we see an expansion
of the sequence found at the Maidstone By-Pass, but virtually all sediments of
daviest Subzone age have been removed by the cvistatum Subzone transgression. This
expansion of the sequence reaches its known maximum in this area at the Sevenoaks
Brick Works Ltd’s pit at Otford.

IN THE ANGLO-PARIS BASIN 23

Part of Division 5 and Division 6 were exposed during the excavation of the
reservoir for the Mid Kent Water Board some 600 yds ENE. of the centre of the Ford
Place Clay pit (TQ 64055920). The lower part of the Gault was also exposed in the
now long-abandoned Pascall’s pit at Wrotham (TQ 62155780) about 2100 yds SW.
of the Ford Place Clay pit and was described very briefly by H. J. W. Brown (1924 ;
79, 81). Nothing can be concluded from his account, but it is unlikely that the
sequence has changed much from that at Trottiscliffe.

(iv) SEVENOAKS AREA

This area has been an important centre for the manufacture of bricks and tiles for
overacentury. Pits have been opened near Kemsing Station (H.J.W. Brown 1924 ;
80), Greatness Lane, Otford (Austin Browne 1949 : Khan 1952 : Casey 1954a :
Milbourne 1956, 1962, 1963 : Owen 1958, 1963a, b), St. John’s, Sevenoaks (Jukes-
Browne 1900 : H. J. W. Brown 1924), Dunton Green (C. W. Wright 1947 : Khan
1952 : Casey 1954a : Owen 1958) and Chevening (Lobley 1880). These sections, with
the exception of the last, together with borehole evidence has given a very good
picture of the Lower Gault in this area. Brown (1924 ; 80) records the occurrence of
the nodule bed (Division 6) at the top of the Lower Gault just N. of the railway line
at Kemsing Station, but there is no other information available about the succession
between Wrotham and Otford, a distance of 64 miles. Today, the only section
available is that exposed in the Sevenoaks Brick Works Ltd., pit at Otford.

(a) Sevenoaks Brick Works Ltd., Greatness

The sequence in the Lower Gault at the Sevenoaks Brick Works Ltd pit at Great-
ness Lane, Otford, is shown in text-fig. 9. It was first described by Khan who dis-
cussed the foraminiferal sequence (1952), then in part by Casey (1954a) when
describing the distribution of Falciferella. Milbourne has described the succession
(1956, 1962) but his reading of it was questioned by the writer (1958, 1963a). The
six broad lithological divisions seen at Ford Place reach their maximum development
here (text-figs. 3 & 9).

A combination of evidence provided by several boreholes together with surface
mapping shows that the Gault-Lower Greensand Junction beds are over 13 feet 6
inches (4-11 m.) thick at Greatness. Division 1 (i) apparently is the transitional bed
linking the Junction beds with the Gault. The writer reported (1963a ; 39) that the
clays of i (ii) to (iv) probably represent the upper part of the benettianus (i.e. Lyellt)
Subzone, and so the bulk of the Subzone is probably present in I (i) and, together with
the eodentatus Subzone, in the sediments below. The species of Protanisoceras (P.)
in I (ii-iv) include P. (P.) barrense (Buvignier) a characteristic lyelli Subzone am-
monite. In comparison with Ford Place, it can be seen that clay sedimentation
commenced earlier at Greatness. The spathi Subzone is represented by Divisions
I (v—vi), 2, and 3 (i-ii). Division 2 is the ‘ Upper dentatus-spati nodule bed ’ as at
Ford Place, and so, the sfathi sediments of Division 1 are slightly thicker at Great-

24 MIDDLE ALBIAN STRATIGRAPHY

Olv. Lithology

Dense band of phosphatic nodules containing semi-derived and indigenous fossils
ng shelly clay (‘lautus Zone nodule bed’). $
| (x) Mo db gre awnish clay with some phosphatic nodules and ammonites. $nitiaus
Line of partly phosphatised ammonites. = :
| (vil) Blue-grey and fownish mottled clay with yellow ochreous balls after pyritic

fossifs and occasional part-phosphatised fossils.

(Wi) Seam of scattered part-phosphatised ammonites.

|
|
| (v) Mottled blue-grey fawnish clay with yellow ochreous balls afterpyrite, and
|
|
|
|
|
|
|

ristatum

5 scattered phosphatic nodules

(lv) Seam of dark-brown phosphatic nodules in clay of Stil). Bee I
(ill) Mottled blue-grey and fawnish clay.
(ii) Persistant seam of scattered part-phosphatised ammonites. »

meandrinus

(1) Mottled blue-gqrey and fawnish clay becoming more fawn-grey towards the
base which Is marked by small lenticular developments of highly ferruginous 2 9
marly clay. Impersistant shell seams occur throughout,

= =
“ aie |
>
(ill) Dark slightly brownish-grey clay. = 1 9
i] °
= = ——— Pe Ce i 1
(ii) Seam of scattered dark brown phosphatic nodules, E Fa =
: v a 2
>
”
3 104
(i)Broad banded clays; alternating dark-grey clay and lighter grey marly,
with lanticular developments ot ferruginous maristone especially inthe 6 2

lower band which varies between 10 and 18 inches in thickness.

SSS
niobe
I

ba are
(vi)Dark grey clay with scattered chocolate coloured phosphatic nodules witha Aine Oe RUE sole 9
zinch thick line of them at the base. stele -o ie, i alate AY, pe e@aesecse o
(v) Dark grey shelly clay. 2 Bi
a—
— = 1-
(iv) Seam of shells with small buff phosphatic nodules and phosphatised ammonite Sete SSS 2
body chambers. ohh
cy 7]
>
>
| .
] iS
&
.
3 (iii) Dark grey shelly clay with lighter grey bands alternating: scattered buft = 12 °
c
phosphatic nodules. ~ os
|
1

<

= 4—s
(il) Greenish glauconitic clay, = 9

(i) Dark grey shelly clay,

- 2 Lo}
2 Dense seam of black phosphatic nodules and casts of ammonites, often with the =
shell, in dark grey shelly clay. ° eae “G 1-2
mae oe ba’
(vi)Shelly dark grey clay with part phospatised fossils in a shell seam 10 inches ~ 3
”

from the base, 1 5

{v) Shelly seam with phosphatic nodules and fossils, Vv eRe dees 1
= eee of—
(tv) Dark grey shelly clay, * — 9
| (ill) Shel! seam with phosphatic nodules, ue cine meer (S ;
1 "
(li) Dark grey shelly clay. 2
1
.
(i) Dork grey sparsely fossiliferous clay becoming more glauconitic downwards ~
with patches of glauconite in the lowest part exposed, = 5 9
1
|
|
| | |
{e)

Fic. 9. Lower Gault section at the Sevenoaks Brick Works Ltd’s pit, 1300 yds NNE. of
Bat & Ball railway station, 150 yds S. of the Otford-Kemsing railway line, and 150 yds W.
of Greatness Lane, Otford, Kent (TQ 53605780).

IN THE ANGLO-PARIS BASIN 25

ness. Division 3 at both localities is closely comparable in thickness and character
except at its base, where 3 (i) at Ford Place has become finer in character and shelly
by Greatness and is represented in 3 (i-1). Ammonites that occur in 3 (i) at Great-
ness consist almost exclusively of species of Hoplites (H.), the fauna being identical to
that of the indigenous element of Division 2 itself which contains rarities such as
Oxytropidoceras but no examples of Anahoplites were found by the writer. At Great-
ness, 3 (ii) contains a few large Inoceramus concentricus but no ammonites.

At the base of 3 (ii), Anahoplites intermedius and A. praecox appear in the sequence
and this is taken by the author to mark the base of the intermedius Subzone. They
range up to the top of Division 3 as at Ford Place. The thin seam of ‘ solid’ phos-
phatised ammonite body chamber fragments containing the coarse form of A. praecox
is also present at Greatness (3 iv), and above this Ewhoplites priced and its close
relatives are characteristic as at Ford Place. Division 4 is thinner than at Ford
Place, and only the basal 2 feet 2 inches (0-66 m.) can be classified with the niobe
Subzone in contrast to the 4 feet 2 inches (1:27 m.) at Ford Place. At the level 2 feet
2 inches (0-66 m.) above the base of Division 4 at Greatness there is a thin bedding
plane (bed 10 of Milbourne 1956 ; 236) which contains species of Mojsisovicsia
including M. subdelarue: (Spath). This marks the base of the swbdelaruei Subzone
which is here 8 feet 7 inches (2-616 m.) thick, and is, therefore, thicker than at Ford
Place. M. remota (Spath) has been obtained from 4 (iii) ; and 5 (i) has yielded
species of Dimorphoplites which indicate the swbdelarwei Subzone rather than the
meandrinus Subzone although Mojsisovicsia has not yet been found init. It is
important to note that lithologically the base of Division 5 at Ford Place does not
correspond to the base of the same lithological Division at Greatness. The meandrinus
Subzone commences with 5 (ii) which has yielded Euhoplites meandrinus Spath, E.
aff. asbasia Spath, Dimorphoplites spp. and large partly crushed Anahoplites planus
(Mantell). No ammonites were recovered from 5 (iii) but the nodules of 5 (iv) have
yielded fragmentary abraded Dimorphoplites and Euhoplites. At Greatness 5 (ii) to
(iv) can be correlated with 5 (i) at Ford Place. Division 5 (v) to (vi) at Greatness
also contains a meandrinus Subzone ammonite fauna, but 5 (vii) did not yield ammon-
ites to me and probably represents 5 (v) at Ford Place which contains an ammonite
fauna which shows some affinity with the mztidus Subzone above.

The mtidus Subzone without doubt commences at the base of 5 (viii) at Greatness
and includes 5 (ix). Together they have yielded a typical mitidus Subzone fauna and
measure 4 inches (0-102 m.) in thickness in comparison with the 1 foot 4 inches
(0-406 m.) of 5 (vi) at Ford Place. Division 6, the ‘lautuws Zone nodule bed’, as the
author demonstrated in 1958 contains a fauna including elements of the mitzdus and
daviest Subzones as well as of cristatwm Subzone age. Here, as at Ford Place, the age
of the clay in which the phosphatic debris is embedded is of cvistatum Subzone age.

It is worth revising here the stratigraphical positions of the samples studied by
Khan (1952) from the Sevenoaks Brick Works.

Sample Sr 5 ft 6 ins down from the top of Division 3 ;
intermedius Subzone as Khan recorded.

Sample S2 6 ins down from the top of Division 3 ;
intermedius Subzone as Khan recorded.

26 MIDDLE ALBIAN STRATIGRAPHY

Sample $3 1 ft 6 ins below the top of Division 4 ;
subdelarueit Subzone not niobe as recorded.

Sample $4 _ 6 ft below the top of Division 5 ;
subdelaruet Subzone not niobe as recorded.

Sample $5 4 ft below the top of Division 5 ;
subdelaruet Subzone.

Sample S6 2 ft below the top of Division 5 ;
meandrinus Subzone.

Sample $7 2 ins below the top of Division 5 ;
nitidus Subzone not subdelaruei Subzone as recorded.

Sample S8 ‘ Jautus-Zone nodule bed’ Division 6 ;
cristatum Subzone.

(b) St. John’s Brickyard

The brickpit owned by Durtnell and known in the literature as St. John’s Brick-
yard, or the ‘Bat & Ball’ pit, is now overgrown. It was situated about 900 yards
NNW. of the Bat & Ball railway station, 100 yds W. of Otford Road, Otford (TO
52805755), and goo yds towards the WSW. of the Greatness pit. The section was
first described by Hill (tn Jukes-Browne 1900 ; 85) and additional information was
given by H. J. W. Browne (1924) and by Spath (e.g. 1925 ; pl. XII, fig. 4). It
showed a sequence (text-fig. 3) intermediate in thickness between that of the Great-
ness pit and the Dunton Green section described next.

(c) Dunton Green

The Dunton Green Brick, Tile and Pottery Works Ltd pit was situated about 1650
yds WSW. from the St. John’s Brickyards. The section is now obliterated. It was
first described in detail surprisingly late in its history by C. W. Wright (1947 ; 315-318)
although it had been mentioned in the literature as early as 1880 (Lobley 1880).
Spath recorded ammonites from it (1923-43) and Khan (1952) and I (1958) have
referred to it briefly. Unfortunately, I saw only the sequence up to the basal part
of Division 4, and the dotted portion in text-fig. 10 is taken from the account by
Wright.

Text-fig. 3 shows the extent to which the sequence is attenuated in comparison
with that at Greatness to the ENE. and the Brasted borehole to the WSW. (not ESE.
as given in Casey 1954 ; 266) discussed below.

The exposed portion of Division 1 at Dunton Green was seen to be identical to the
corresponding portion of Division 1 at Greatness. Division 2 contains the same
fauna as at Greatness but the phosphatised material is devoid of the shell. Division 3
(i) contains ‘solid’ pyritised Hoplites (H.) with the nacreous shell, including H. (H.)
dentatus densicostata Spath and H. (H.) escragnollensis Spath. This bed corresponds
to 3 (i) at Greatness. The basal 2 feet 3 inches (0-686 m.) of 3 (ii) at Dunton Green
consists of a glauconitic clay, immediately above which Anahoplites intermedius
appears in the sequence and ranges up through the remainder of the Division.
Although the sequence is much more condensed, the thin bed of phosphatised body

IN THE ANGLO-PARIS BASIN 27

—
=
Div. Lithology Ft Ins M.
[eT Bade doth town bresphetle nadulen OIIIILILII“AS eye 4
!
1 ! l
1 I |
! ! |
i. i
] | | “
3
1 5 , | =
; a Lightish grey clay, slightly mottled, darkening below. 1s
nes sy Wright 1947;318] | | 0]
\ I 1s
6
! I 1
c
1 1 1
i} ! |
! 1 '
J !
1 ' !
| ! | 5
Lightish-somewhat fawn-grey clay. seen to fe)
(lv) Dark-grey clay with small dark-brown phosphatic nodules, °
(iil) Shell seam with phosphatised ammonite body chambers. { 4
“”
=
vv
+7
iS
= 3
=
(ii) Dark grey clay with crushed fossils andscattered small
phosphatic nodules passing down at the base into 10 6
glauconitic gritty clay.
2
1
(i) Dark grey clay, shelly with some pyritised ammonites. ~ 6
Dense seam of black phosphatic fragments of ammonites etc. = 12
Dark grey clay with crushed fossils chiefly ammonites. 6
ce)

Fic. 10. Section in Lower Gault at the Dunton Green Brick, Tile & Pottery Works Ltd’s
pit, c. 550 yds S. of Dunton Green railway station on E. side of railway line, Longford,
Dunton Green, Kent (TQ 515570). The dotted portion is completed from the account
by Wright (1947).

28 MIDDLE ALBIAN STRATIGRAPHY

chambers of ammonites 3 (iii) occurs, and may be correlated with 3 (iv) at Greatness.
3 (ili) at Dunton Green has yielded the coarse form of A. praecox figured by Spath
(1925 ; 132, text-fig. 35e) according to its preservation. Only a foot of Division 4
was seen clear by the writer, and for the remainder of the sequence Wright’s account
is available.

(d) Brasted

The Metropolitan Water Board well at Brasted has been described by Casey
(1954a ; 266 : 1961a ; 544-5) and is displayed in text-fig. 11. Its relationship to the
sections at Dunton Green and Squerryes is shown in text-fig. 3. The sequence in the
higher part of the well is similar to that of Dunton Green and Greatness. Un-
fortunately, the top of the Lower Gault bisects the ground surface further to the north
of the site of the well, and the boring commenced at an unknown depth below the
base of the Upper Gault.

At a depth of 12 feet 6 inches (3:81 m.) the reddish burrowed marl suggests a
correlation with the base of 5 (i) at Greatness. The only two ammonites preserved
in the collection of the Institute of Geological Sciences (GSM) are Ca 339, Ewhoplites
sp. from a depth of about 8 feet (2-438 m.), of either swbdelarue: or meandrinus Sub-
zone age, in the preservation typical of the lower part of Division 5 at Greatness. The
other, Ca 340 from a depth of 10 feet (3-048 m.), is of no diagnostic value. Together
they neither confirm nor deny a subdelaruei Subzone age. The dividing line between
Divisions 4 and 3 occurs between a depth of about 22 feet and 25 feet (6-706—7-62 m.)
where the lithological change from fawn-grey clay to mid-grey clay occurs. The
equivalent of 3 (iii) at Dunton Green occurs at a depth of 29 feet (8-839 m.) and
yielded Anahoplites praecox (GSM Ca 364-6) and Hoplites (H.) sp. (GSM Ca 367). At
a height of 1 foot 6 inches (0-457 m.) above Division 2 a phosphatised fragment of a
coarsely ribbed Anahoplites with the nacreous shell preserved was recovered.

Division 2, the ‘ upper dentatus-spathi nodule bed ’ was struck at a depth of 38 feet
6 inches (9:296 m.). Below this level the sequence departs markedly from that of
Dunton Green and Greatness. The 20 feet 6 inches (6-248 m.) of mid-grey clay below
Division 2 contains Hoplites (H.) spp. throughout. No definite indication of the
lyelli Subzone is seen, but a boring encompasses only a very small lateral area and the
possibility that part of this succession is of lyelli Subzone age should not be dis-
counted. There was also no evidence of the eodentatus Subzone although this may
be present below 59 feet (17-983 m.) depth.

(v) BRASTED TO BUCKLAND
(a) Westerham

The northern face of the Squerryes Estate Sand pit situated 24 miles SW. of the
Brasted Well was cut back in 1964 and provided the section given in text-fig.12. The
sequence of well-marked lithological divisions seen in the lower part of the Lower
Gault in the Sevenoaks area, already indistinct at Brasted, becomes even less dis-
tinct at Squerryes. Here, two clear-cut divisions are immediately apparent and

IN THE ANGLO-PARIS BASIN

Olv. Lithology

Light grey weathered clay marked at the base by burrowed
reddish marl,

Vs
c
le
Ie
le
,&
1
le
a
LS
le
Fawn grey clay with phosphatic nodules. |
1
|
|
/
(ill) Mid grey shelly clay.
“
=
v
ev
(li) Shell seam with phosphatised ammonite body chambers. E
£
=
(1)Mid grey shelly clay.
yN
Ur
te
Blackish phosphatic nodules and fragmentary fossils with the shell ie
Qa
”

(v)Mid grey shelly clay.

(iv)Phosphatic fragments of fossils with the shell.

(iii)Mid grey shelly clay with part-phosphatised, part-pyritised
fossils.

(ii) Phosphatic fragments of fossils with the shel! in mid-grey clay.

(i) Grey shelly clay.

Black sandy clay with phosphatic nodules andiron pyrites.

Remainder of Bed 8 of Casey 1961a;545.

Ft

— i —

22 at 4% see a —

29

Ins M.

=

=

=

Fic. 11. Vertical section of Middle Albian sediments in the Metropolitan Water Board
well, 1000 feet NE. of St. Martins Church, to the E. of Station Road Brasted, Kent

(TQ 47095574).

30 MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology

(vil) Mottled blue-grey-fawn clay with crushed /Inoceramus
concentricus and Hoplites(H.) spp. with the shell. Seen to

(vii) Blackish phosphatic nodules mainly casts of Hoplites (H.)with
portions of shells preserved, in shelly mottled clay.

(vl) Clay as in 2 Wiii) with partly phosphatised Hoplites (H.)

spathi

(v) Clay asin 2 Wiii) but much less fossiliferous.

(iv) Shelly grey-blue-fawn clay with part-phosphatised part
pyritised Hoplites (H.)

iii) /.concentricus shell seam with part-phosphatised fossils
| (il) 2’ mottled Dlue-grey clay with crushed fossils
i) Dense seam of small phosphatic nodules and phosphatised fosstts

Mottled blue-grey-fawn clay with lenticles of ferruginous marl.

Clay as in 1(v) but without phosphatic nodules, and with
pockets of glauconitic quartz sand, and ferruginous
lenticles. Crushed part-phosphatised Hoplites (H.)occur
without the shell.

(iii) Darker grey clay with ferruginous lenticles with
indigenous part—phosphatised Hoplites (H,).

(ii) Lighter grey clay with many ferruginous lenticles.
Small phosphatic pieces with Hoplites (H.).

tyelli

(i) Dark grey shaly mottled clay with reddish ferruginous
lenticles.

Dark grey gritty clay.

eign. pyritic dark grey clay with greyish septarian phosphatic

no jes and .part-phosphatised fossils wit PY rite replaced shells;
s| c Dn oO Cg no © a 0 en

Dark grey pyritic clay, gritty, and weathering ferruginous,

Dark grey highly pyritic clay with septarian phosphatic nodules
weathering ferruginous.

Dark gritty grey clay with scattered septarian phosphatic nodules.
Seam of large spherical blackish phosphatic nodules in glauconitic lob

Black gritty blocky weathering clay with glauconite, When
weathered clay partings are coated with pyrite decompos-
ition products,

eodentatus

_ = =
ale
ele

Bed 12 of Casey 19610;543 puzosianus Subzone

Fic. 12. Middle Albian sediments at the Squerryes Estates Sand Pits’ pit, c. 150 yds N. of
Covers Farm, Westerham, Kent (TQ 43305395).

IN THE ANGLO-PARIS BASIN 31

their correlation is shown in text-fig. 3. The sequence is, however, thinner than at
Brasted.

The g feet 2 inches (2-794 m.) of Division 2, excluding bed (i), contains a typical
spatht Subzone fauna, the ‘ upper dentatus-spathi nodule bed’ being represented in
2 (vii). Bed (i) is the highest level in which Protanisoceras (P.) moreanum (Buvignier)
occurs, and this is taken to mark the top of the /yelli Subzone sediments. The under-
lying 11 feet (3-353 m.) of Division 1 represents the uppermost part of the lyelli
Subzone as developed at Horton Hall, Sussex (p. 35). However, the typical develop-
ment of this Subzone occurs within beds 13-15 of the Gault-Lower Greensand
Junction. Bed 15 has yielded the following ammonites:

Protanmisoceras (P.) moreanum (Buvignier), Beudanticeras laevigatum (J. de C.
Sowerby), B. sanctaecrucis (Bonarelli), B. albense Breistroffer, Hoplites (H.) dentatus
(J. Sowerby), H. (H.) bullatus Spath, H. (H.) baylet Spath, Lyelliceras lyelli (d’Or-
bigny), Brancoceras versicostatum (Michelin non d’Orbigny, nec Douvillé), ‘ Oxytropi-
doceras ’ cf. evansi (Spath).

Bed 13 has yielded Beudanticeras laevigatum, Hoplites (H.) spp., ? Otohoplites sp.
ind., Lyelliceras sp. Beds 10-12 contain Hoplites (Isohoplites) spp., including H. (J.)
eodentatus (Casey 1961a ; 543).

(b) Tandridge

The Coney Hill Sand-pit, Barrow Green (TQ 37755250), situated about 3} miles
WSW. from Squerryes, has already been described by the author (1963a ; 39). It is
apparent that the sequence has become thinner (text-fig. 3). Between Tandridge
and Buckland, a distance of 9 miles, there is no information concerning the sequence
at the outcrop.

(c) Buckland

The Buckland Sand & Silica Co’s pit at Buckland (TQ 231512) was described by
the author in 1958 (1958 ; 149-152), however, a certain amount of revision is now
necessary. The sequence (text-fig. 13) is not unlike that of Ford Place, a fact already
recognised in the Gault-Lower Greensand Junction Beds by Casey (1961a ; 552).
Whether the eodentatus and lyelli Subzones are present in the upper part of the
Gault-Lower Greensand Junction as at Ford Place is at present unknown (p. 20).
The spathi Subzone is represented within beds 2-6. Bed 2, tentatively classified
with the benettianus Subzone in 1958, was later included in the spathi Subzone (Owen
1963a ; 47-48). It contains the same ammonites as Division I (ii) at Ford Place.
Bed 3 is the obvious correlative of 1 (iii), and Bed 4 is the equivalent of Division 2,
the ‘ upper dentatus-spathi nodule bed’. Bed 4 has yielded the best preserved fauna
yet known from this horizon, and the effect of strong erosive currents on the sea-
bottom is demonstrated by the effaced nature of the upper surface of the nodules.

Beds 5-11 probably represent Division 3 at Ford Place but the character of the clay
is very different. In general the sediments are gritty clays with intercalated beds of
ferruginous marlstone, and the fossils have shells replaced by pyrite. Beds 5 and 6

32 MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology

subdelaruei

Mottled brown and blue-grey clay with selenite. A seam of crushed 2 {e)
bivalves occurs at the base with poorly preserved part-phosphatised
| ammonites.
8
Light-grey weathered clay with much selenite. S 3 3
°
c
7-
11. | Very dark brown clay. 4 ie)
64
r_ “
=
v
vo
10 | Dark blue-black blocky clay with yellowish patches mainly on g 3 fe)
partings. ©
=
Highly ferruginous yellowclay with stony lenticles of reddish marlst— = 7 5
2h one containing disc-shaped pebbles of micaceous maristone,
4 (e)
8 Dark grey clay with pyritised and partly-phosphatised ammonites
but usually crushed flat, 4
"GM ID NEGLI LD) — 4

Impersistent but well developed lenticles of hard ferruginous
marlstone with lateral interstitial gritty clay-marl. PL |

fo)
6 Hard brownish marly clay with marly seams passing down into
the bed below.
A 2
5 Dark grey slightly mottled clay. = Oo
[ 3
Greyish-white to black phosphatic nodules, brownish when a
4) weathered, mainly ammonite casts with abraded upper surfaces ™4
3 Dark grey clay with pockets of glauconite, some oxidised.
Phosphatic nodules occur scattered at about the middle of the 34
bed,
Scattered subangular phosphatic nodules in gritty glauconiticclay. | 2
2
Gritty glauconitic clay with patches of glauconite sand. ° 6
= (@)

Gault—Lower Greensand junction beds

Fic. 13. Section of Lower Gault in the Buckland Sand & Silica Co’s sandpit extending
from a point 2100 yds WNW. of Reigate railway station to a position c. 250 yds SW. of
Dowdes Farm, and 50 yds S. and roughly parallel to the Dorking-Redhill railway line,
Buckland, Surrey (TQ 232512).

M Ft HASSOCKS HORTON HALL STORRINGTON PITSHAM NYEWCOD SELBORNE WRECCLESHAM SHERE BUCKLAND TANDRIDGE
140 oS ubdelarvei{_ 13
niobe 12
ee
a0 M1
130 ile)
fee 9
8
intermedius a7
? ? 2 ? 6
7120 L,
354 — >] —— 4
6 9 3
P2Zyeee| 5 — a
4
[110 |
3 spothi 7
>
30400 =
SS
NSN
0 ON 6
. S
= 4
}90 — 3
? 2) ————e ?
255 L
Broad eodentatus
60
? ?
770
205
+60
15450
Subzonal correlation lines
—_- — — ——Lithological correlation lines
L40
104
30
20
5:
10

Fic. 14. Correlation of sections in Surrey, Hampshire, and Sussex, at the outcrop.

ar

}
2

Sacre. 0
- £ :

7
-

IN THE ANGLO-PARIS BASIN 33

are classified with the spathi Subzone, Beds 7-11 with the intermedius Subzone. The
siltier nature of the sediments of the zmtermedius Subzone heralds the sequence seen
further west at such localities as Winchester (p.69), Didcot (p.63), and Devizes (p.60).
The mobe Subzone is probably present in the top of these sediments and possibly in
the light grey clay of Bed 12 which can be correlated with the base of Division 4 at
Ford Place. A re-examination of the ammonites from Bed 13 indicates that it
should be classified with the subdelaruet Subzone as the writer considered possible in
1958.

The presence of higher Middle Albian subzones in the sequence is at present un-
certain. Gossling (1929 ; 251) reported, from Spath’s determinations of the am-
monites found in the Merstham trench, that here the Upper Gault rests directly upon
the dentatus Zone. However, Gossling did not give a detailed account of the succes-
sion and, moreover, the trench was only 4 feet (1-219 m.) deep. In the Buckland
Sand & Silica Co. pit large distorted blocks of pale-grey clay were seen in the super-
ficial deposits. These blocks contain orbignyi Subzone fossils and the bed from which
they were torn cannot be far to the northward. However, in the foundations of
Wray Common County Primary School at the N. end of Kendal Close, Reigate
(TQ 26975092), the ‘ lautus Zone nodule bed’ was located. The fauna obtained
indicated that its character was closely comparable to that of Division 6 in the
Sevenoaks area.

(vi) BUCKLAND TO UPPER BEEDING (text-fig. 14)

There is very little information available at the outcrop between Buckland and
Shere, a distance of 104 miles, except that given by Dines & Edmunds (1933). The
sections exposed during the cutting of the Shere By-Pass road have been described
by the writer (Owen 1963a). There is very little doubt that the outcrop here is
affected by a strike fault with a substantial northerly throw as Kirkaldy concluded
(1958 ; 18). The structure illustrated by the author (1963a ; 41, text-fig. 1), the
greatly attenuated width of the outcrop, the far greater thickness of the Gault in the
West Clandon Waterworks boring, and the sequence at Albury (Edmunds im Dines &
Edmunds 1929 ; 41-2) all support the presence of such a fault. The true degree of
representation in the Lower Gault in this area is, therefore, uncertain, and will not
be determinable until a cored borehole is drilled to the north of the disturbed ground.

The notable feature in the lower part of the Lower Gault between Shere and
Shalford is the development of the eodentatus and lyelli Subzones (Owen 19638,
Wright & Wright 1948). Whether this development continues west to Whiteacre
Copse on the Guildford-Godalming By-Pass road is unknown (Lea 1932 ; 320-1 ;
Owen 1963a ; 50). There is a similar dearth of information further west. However
in the Institute of Geological Sciences there is a small suite of specimens from the
Gault exposed in a gravel pit off the Farnham to Runfold road at about 1 mile ENE.
of Farnham and immediately W. of the railway bridge (SU 85854755) GSM.
He 2337-45, which indicates the presence of Jawtus Zone sediments in this area.

The sections at Wrecclesham, Selborne, and Nyewood, have all been described by
the writer (Owen 1963a). Text-fig. 14 shows that the sequence in the spathi Subzone

Cc

34 MIDDLE ALBIAN STRATIGRAPHY

expands south of Wrecclesham to reach its greatest known thickness in this area at
Selborne. It then thins southwards towards Nyewood, and between this locality
and Storrington the Gault rests directly upon the oxidised indurated ‘ iron grit’ at
the eroded top of the Folkestone Beds. How much more of the Middle Albian is
represented in the higher beds of the Gault is uncertain. I expressed the qualified
opinion (1963a ; 51) that at Selborne it appeared that the cristatum-orbignyi Subzone
transgressions had cut down to the spathi Subzone, a view disputed by Milbourne (in
Hancock 1965 ; 250).

Osborne White (1910 ; 17) followed Jukes-Browne’s interpretation of the ‘ inter-
ruptus ’ Zone (see p. 111). He records at Bradshott Hall (1910 ; 20) 2 feet (0-609 m.)
of clay with phosphatised ‘ Hoplites interruptus’ separated by 3 feet (0-914 m.) of
deposits from sediments containing Inoceramus sulcatus indicative of the lower part
of the Upper Albian. Unfortunately, this material has not been traced. The lowest
part of the intermedius Subzone with Anahoplites osmingtonensis nov. is present at
the outcrop in the Petersfield area (e.g. BMNH, C 35482-3). Further information
has been provided by borings carried out for the Gas Council by the British Petro-
leum Co. in the Winchester area in the deeper part of the Wessex basin (p. 69).
There, sediments of the intermedius Subzone are present above the spathi Subzone,
and the orbignyi Subzone is also represented within the silty clay facies.

Between Petersfield and Storrington only three sections are available, all in the
spatht Subzone ; Nyewood, Pitsham, and Sullington (Owen 1963a). Recently,
Mr. C. J. Wood of the Institute of Geological Sciences recognised in the Brydone
Collection the material recorded by Jukes-Browne (1900 ; 112) from a well to the N.
of Graffham village. The specimen recorded as Hamites punctatus ? is here referred
to Protanisoceras (P.) barrense (Buvignier) and is preserved in lilac-grey silty clay,
the shell originally having been replaced by pyrite. It is accompanied by Inoceramus
concentricus in the same preservation. The ammonite indicates the lyelli Subzone.
The specimen of Hamutes attenuatus is indeed of that species, but is preserved in
weathered yellowish pale-grey tough silt. It is of either Jovicatus or possibly lautus
Zone age.

Osborne White (1924 ; 26) described a well section examined by Templeman
situated on the W. side of the lane to Barns Farm, 300 yds S. of its junction with the
Washington-Storrington A283 road (TQ 10501345). Below 10 feet (3-048 m.) of
clay containing an Upper Gault fauna, about 40 feet (12-192 m.) of Lower Gault was
proved including the cristatwm and the upper part of intermedius Subzones. Ina well
bored at Wiston Hall between Washington and Steyning 176 feet (53-644 m.) of
Gault was proved (Osborne White 1924 ; 25). It seems likely, therefore, that in this
area the succession in the post spathi Subzone sediments is closely comparable to that
described below exposed in the Horton Clay pit.

(vii) UPPER BEEDING

The Lower Gault section exposed in the British Portland Cement Manufacturers
Ltd’s Horton Clay pit is shown in text-fig. 15, and its relationship to sections west and
east is shown in text-fig.14. The pre-war workings were described by Osborne White

IN THE ANGLO-PARIS BASIN 35

(1924 ; 27-8), and the current sections have been described in part by Milbourne
(1961 ; 135-7), Casey (I961a ; 558), and Owen (1963a ; 46). It shows the thickest
sequence yet known in the Lower Gault of England, and is undoubtedly the most
important. The clays contain finely disseminated pyrite, and their silty nature
allows the deep penetration of weathering agents ; this obscures the bedding for at
least 12-14 feet (3-658-4:267 m.) from the surface, but fortunately there is a dip of
8°S. present. In the fresh condition, the clays are seen to be fairly uniform through-
out, and the Divisions adopted here are based upon cycles of sedimentation. Each
cycle commences with relatively rapid sedimentation and terminates with partly
arrested deposition indicated by marly seams and cementstone nodules. Individual
units display an alternation of dark-grey and more fawn-grey bands. That this is an
original feature is occasionally shown by the fauna ; there being often a more diverse
benthos in the fawn-grey bands.

Casey has described the boreholes drilled over the area of the northern field
situated to the E. of Horton Wood (1961a ; 558). The subsequent excavation of this
field commenced in 1964 at the northern boundary, and the sequence is now being cut
down dip. There is unfortunately a small gap in the observed sequence as shown in
text-fig. 14, between this new field and the older workings.

Eodentatus & basal lyella Subzones

About 13 feet (3-962 m.) of glauconitic sandy clay and loam (rz (i) & (ii) ) classified
with these Subzones occurs below the lowest level seen in the excavations (Casey
rg6ra ; 558).

lyella Subzone

Divisions I (ii) to the top of 2 (vi) are classified with this Subzone. As this is the
first time that such a faunal sequence has been described in England, it is discussed
in detail. The top 3 feet 8 inches (1-117 m.) of I (ii) is seen in the excavations, but
only the upper foot is fossiliferous and this has yielded Hoplites (H.) spp. and the
ubiquitous Inoceramus concentricus, among other fossils. The shell seam that marks
the base of 2 (i) has yielded the following :—

Protanisoceras (Protanisoceras) nodonewm (Buvignier). P. (P.) barrense (Buvignier),
P. (P.) alternotuberculatum (Leymerie), Beudanticeras laevigatum (J. de C. Sowerby),
B. albense Breistroffer, Hoplites (H.) spp., Lyelliceras aff. lyell (d’Orbigny), Branco-
cevas (Brancoceras) sp., Neohibolites minimus (Miller), I. concentricus Parkinson,
Acila (Truncacila) bivirgata (J. de C. Sowerby), Natica sp.

A comparable fauna occurs throughout the remainder of 2 (i). In 2 (ii) the
benthonic element of the fauna increases in importance and one of the characteristic
fossils is a solitary caryophyllid coral. The ammonites consist essentially of species
of Hoplites (H.) and Protanisoceras (P. barrense & P.nodonewm) ; Lyelliceras is not
common. The benthonic element is much reduced and the nekton to a lesser extent
in 2 (iii). The ammonites include the following :

H. (H.) dentatus (J. Sowerby), H. (H.) spp. common ; P. (P.) barrense, Lyelliceras
Wells (d’Orbigny) the lowest definite occurrence of the typical form, Beudanticeras
sp., accompanied by I. concentricus.

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37

IN THE ANGLO-PARIS BASIN

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38 MIDDLE ALBIAN STRATIGRAPHY

A much more diverse fauna appears in 2 (iv). A thin bedding plane 4 inches
(0-I0or m.) above the base, sometimes washed out, has yielded one of the best pre-
served ammonite faunas yet known from the lyelli Subzone. The fossils are part-
phosphatised and part pyritised. Some ammonites are in a ‘ death rest’ position ;
that is, they are vertically orientated resting on the venter. The amount of distortion
permits the compaction ratio of the clay to be deduced. The fauna includes :

P. (P.) nodoneum, P. (P.) barrense, Pseudhelicoceras argonnensis (Buvignier), P. sp.,
Beudanticeras laevigatum, B. albense Breistroffer, B. sanctaecrucis Bonarelli, Hoplites
(H.) baylet Spath, Hoplites (H.) spp. Douvilleiceras sp. juv., Oxytropidoceras evansi
(Spath), O. sp., Lyelliceras lyelli (d’Orbigny), L. gevreyi (Jacob), L. sp., Brancoceras
senequiert (d’Orbigny), B. versicostatwm (Michelin non d’Orbigny nec Douville) B.
spp., Inoceramus concentricus, Semisolarium moniliferum (Michelin). ‘ Auricula’
acuminata Deshayes. Hemuaster sp.

This list shows a preponderance of lyelliceratid ammonites over the hoplitids which
are greatly subordinate in actualnumbers. The fossils are mainly crushed flat in the
remainder of 2 (iv) and (v) but there is no significant difference in the fauna. Large
uncrushed or partly crushed fossils occur in 2 (vi) and include :

P. (P.) barrense, Beudanticeras sanctaecrucis, H. (H.) benettianus (J. Sowerby), H.
(H.) spp., Lyelliceras lyell, Brancoceras spp. Eutrephoceras sp.

At this level Hoplites occurs in roughly equal numbers to the non-hoplitid genera
combined. Some of the ammonites are also in a vertical ‘death rest’ position and
these, like those in 2 (iv), have a common orientation indicating a current direction
coming from what is now 210° Magnetic.

spatht Subzone

Divisions 3 and 4 are classified with this Subzone. There is a sharp change in the
ammonite fauna at the base of Division 3, the ammonites consisting essentially of
Hoplites (H.) spp. At the commencement of the spathi Subzone, deposition of
sediments increased rapidly and the bulk of the remainder of the Lower Gault,
although very fossiliferous, yields little but crushed material except at a few horizons.
Sedimentation during the spatht Subzone was particularly thick. At least 56 feet 9
inches (19:812 m.) of clays have been observed and the gap in the sequence between
the northern field and the main workings is probably only a few feet.

Division 3 (i) contains numerous J. concentricus and crushed Hoplites (H.) the
bulk of which possess dentatus-like ribbing : a few pyritic nuclei occur. Protanisoceras
(P.) spp. have been obtained from Division 4 (i) but otherwise the heteromorph
ammonites are almost exclusively species of Hamites. A beautifully-preserved
fauna occurs in 3 (ii)-(iv) consisting mainly of ammonites and almost entirely of
species of Hoplites (H.) identical to that of the well preserved element in the dentatus
nodule bed at Folkestone (p. 13). Some specimens reach a diameter of 9 inches
(0-228 m.). A loose block certainly from either 3 (ii) or (iv) contains an example of
Mojsisovicsia delaruei (d’Orbigny). Division 4 (i) has a fauna closely comparable to
that of 3 below. However 4 (ii) contains species of Hoplites (H). transitional from
those seen below to those characteristic of the upper part of the spathi Subzone.
They are of the grade well represented condensed in Division A of the Maidstone

IN THE ANGLO-PARIS BASIN

Fic. 16. Correlation of Lower Gault sections at Horton Hall and Folkestone.

M Ft pclae : Pel |
{ a oe
Gy Ps
ii 6) Em
4
4 <a ad
j440 6(iv)
“a G(iii)
SD TIN.
+130 :
6(i)
t 5(iv)
ed Siiii) |.
a ace aiid
5(ii) — me :
5(i)
- tis ale. De > 4
it Aliv) face ed "ie \ \._ Folkestone
i 3 “s VINKi)
Bie
t 4 (iii) +e i ey
+90 -
| ith
25.
+ 80 Alii) i
af | (vid
+70 li)
204 Ai) = |(v)
7
+60
| Ali)
L
157 50 3liv
3iiii)
nf 3 ii)
4
3 40 3(i)
4
10-
130 2(vi)
2(iv)
r 2(ii)
+20 2(i)
5
10i)
10
4
I; 10)
‘o-—o0

39

40 MIDDLE ALBIAN STRATIGRAPHY

By-Pass. Division 4 (iii) contains species of Hoplites (H.) that are found in the upper
part of the spathi Subzone, and condensed in the ‘ upper dentatus-spathi nodule bed ’
in the northern Weald.

In the most northern cut of the old W. field, situated S. of Horton Wood, the top
of the spathi Subzone was seen in 4 (ili). This consists of 1 foot 6 inches of shelly clay
containing numerous crushed Hoplites (H.) some with well developed lautiform
ribbing. The remainder of 4 (iii) in this cut contains numerous Inoceramus con-
centricus but no ammonites were found by the author either in these clays or in 4 (iv).
Anahoplites intermedius appears in 5 (i) and this level is taken to mark the base of the
intermedius Subzone.

Before describing the remainder of the Lower Gault here, the correlation of the
spatht Subzone sediments with the sequence at Selborne (Owen 1963a ; 43-4) will
first be considered (text-fig. 14). Division 3 (i) at the Horton Clay pit contains the
same fauna as Beds 1-3 at Selborne. On faunal and lithological grounds, 3 (ii) may
be correlated with Bed 4, 3 (iii) with Bed 5, and 3 (iv) with Bed 6 at Selborne.
Division 4 at the Horton Clay pit represents Beds 7-9 and the remainder of the
sediments of the spathi Subzone not yet exposed at Selborne. The correlation of the
Folkestone and Horton Clay pit sections is shown in text-fig. 16.

intermedius Subzone

The sediments from the base of 5 (i) to approximately 1 foot (0-305 m.) below the
top of 5 (iii), a total thickness of 20 feet (6-096 m.), contain a typical intermedius Sub-
zone fauna. The ammonites are quite often of good size (up to 4—5 inches (0-127 m.)
in diameter) but crushed flat. Anahoplites intermedius, A. praecox, A. mantelli and
A. planus are common to within 6 feet (1-829 m.) of the top of 5 (iii) but then decline in
numbers above, with Inoceramus concentricus becoming the dominant fossil.

niobe Subzone

At a level about 1 foot (0-305 m.) below the top of 5 (iti), Dimorphoplites niobe
appears sparingly together with Anahoplites planus, Hamuites tenutcostatus and
numerous J. concentricus. In 5 (iv) partly crushed ammonites and bivalves occur in the
cement-stone nodules, and crushed fossils occur in the interstitial clays ; the stony
lenticles are original sedimentary features. The ammonites include Anahoplites
planus, Dimorphoplites niobe, D. spp. The same fauna occurs in 6 (1) and (ii). The
niobe Subzone is represented, therefore, by 9 feet (2-743 m.) of sediments.

subdelaruet Subzone
This subzone appears to be represented only within 6 (iii) and has yielded species
of Mojsisovicsia including M. subdelaruet and M. remota.

meandrinus Subzone

Division 6 (iv) contains shell seams which, near the top, yield pyritised fossils. The
fauna is typically that which occurs in Bed IV and the basal few inches of Bed V at
Folkestone classified with the meandrinus Subzone. Three shell seams at depths of
approximately 2 feet 4 inches (0-711 m.), 3 feet 4 inches (1-016 m.), and 4 feet 4 inches

IN THE ANGLO-PARIS BASIN 41

(1-321 m.), from the top of 6 (iv) have yielded ammonites suggesting a correlation
with the basal 2 inches (0-051 m.) of Bed V at Folkestone.

nitidus Subzone

A typical nitidus Subzone fauna has been obtained from 6 (v) (Owen 1963a, 46)
preserved in a manner identical to that of Bed V at Folkestone. No diagnostic
ammonites were found by the author in 6 (vi) or (vii) and it is still uncertain whether
these sediments are of nitidus or daviest Subzone age.

? daviest Subzone

In 1963, the author stated that the davies: Subzone was absent. However, a few
phosphatic nodules from the top of 6 (vii) have yielded ammonites including Ana-
hoplites planus and Euhoplites truncatus together with I. concentricus. These occur
immediately below the basal cristatum nodule bed at the base of Division 7. In the
absence of Anahoplites davies: it is not yet possible to determine whether these
deposits represent this Subzone or not.

(viii) HASSOCKS TO EASTBOURNE

The only section in the Lower Gault now available east of Small Dole is to be seen
above the Folkestone Beds in Messrs Hudsons Ltd’s pit at Hassocks (text-fig. 17).

Bed Lithology Ft Ins M.

White to buff spherical septarian phosphatic nodules in weathered 2 ie)

grey clay. A scattered line occurs at the base anda scattered
! 9 Inches above it.
ES a
Dark grey clay with patches of glauconitic sand which become less be 24
marked upwards. The clay Is well bedded and slightly shaly. > 8 fo)
=
c
x
>»
°
«
| 15
|
ee | 7
Loam streaked orange and greylsh with grey clay content ae" 89
increasing upwards and passing down into browntsh loam which fills eres a
ows in the bed below. 9
3 10
oH

Folkestone Beds

Fic. 17. Basal Gault and Gault-Lower Greensand junction beds at Hudsons Ltd’s Hudsons
Red Sand pit, 1400 yds W. of Hassocks railway station and 350 yds S. of the B 2116 road,
Hurstpierpoint, Sussex (TQ 29121552).

42 MIDDLE ALBIAN STRATIGRAPHY

This section has been discussed by Osborne White (1924 ; 29) Kirkaldy (1935 ; 526)
and Casey (1961a ; 559-560). The eodentatus Subzone is present within Bed 4, and
the lyelli Subzone within beds 4 and ?5.

Further east in Sussex there is very little information about the Lower Gault other
than that recorded by Clement Reid (1898), Jukes-Browne (1900) and Osborne
White (1924 & 1926). Spath demonstrated that at Ringmer the daviesi Subzone is
represented (1926a ; 154) and it seems from the well records (Edmunds 1928) that the
Middle Albian sediments probably maintain the thickness seen at Small Dole and may
well thicken a little. The major increase in thickness of the Gault eastward is mainly
explained by the change in facies from Upper Greensand to Gault. This is certainly
the case in the Lewes area where the proven Upper Gault is very thick.

The discovery of a nodule bed of spathi Subzone age in the sea-bed ESE. of Beachy
Head was used by the writer as evidence of an attenuation of deposits of this age in
that area (Owen 1963a ; 46, 48, text-fig. 2). However, phosphatised Hoplites (H.)
occur in Division 4 at the Horton Clay Pit where the sequence is very thick, and the
record of the Eastbourne Waterworks well given by Jukes-Browne (1900, 118) is
probably misleading.

B. Isle of Wight, and Dorset Coast

This section deals with the outcrops in the Isle of Wight, from Punfield to Osming-
ton, and from near Seatown to the Devon border (text-fig. 18). Of necessity the
account is very incomplete for the exposures are seldom good and the facies is such
that ammonites are uncommon except at a few horizons in the various localities.
For this reason no correlation diagram is given on this occasion. Despite the
difficulties, useful results have been obtained.

(i) ISLE OF WIGHT

The Carstone, and the overlying Gault (aptly named the ‘ Blue Slipper ’) describe,
except in the centre of the Island, narrow outcrops from Redcliff near Culver in the
east to Compton Bay in the west. This is in response to the high dip on the northern
limb of the Sandown and Brighstone anticlines respectively. | Where the two axes
meet in the centre of the Island there occurs a structural ‘ no-man’s land’ with com-
paratively gentle northerly dips and thus a broader outcrop. An outlier of Chalk
and Upper Greensand in the southern part of the island is fringed by outcrops of
Carstone, and Gault dipping gently south. The Carstone and Gault have been
described in stratigraphical detail notably by Bristow et al. (1889), Jukes-Browne
(1900 ; 126-130), Osborne-White (1921), Kitchin & Pringle (1922a ; 160-161), Spath
(1943 ; 741-743), and Casey (1961a ; 512-515), but there are many other references to
them in the literature. The Gault is responsible for the major landslips on the
southern coastline, and much of the outcrop is obscured by slipping, sludging, and
deep weathering. The sections that are available are not always easy to work and
fossils are far from plentiful.

Fic. 18. Sketch map showing positions of sections in South West England discussed in the text.

CARDIFF

he

a,

BEER HEAD

@ TAUNTON

@ NEwPoRT

eaqinroRt
20 6 s

stonedarra
GoldenCop & .
Therneombe Beacon

BUDLEIGH

SALTERTON
¢ = = up z miles go
0 10 20 km 30

@ BRISTOL

@ YEOVIL

@ SHERBORNE

@ MAIDEN BRADLEY

ia a
1 Fitzpane,

@ DORCHESTER

SQ Hoiworth House

Black Head
IEY MOUTH

PORTLAND

WAREHAM
.

PQOLE

nes!
UMtHAgton

i”

@ MARLBOROUGH

@ SALISBURY

CHRISTCHURCH

BOURNEMOUTH ee

\STUDLAND

aa Cove
SWANAGE

P PORTLAND BEDS

K KIMMERIDGE CLAY
C CORALLIAN

OC OXFORD CLAY

CB CORNBRASH

GO GREAT OOUTE

10 INFERIOR OOLITE
UL UPPER LIAS

ML MIDDLE LIAS

LL LOWER LIAS

T TRIAS

CTS KINGSCLERE

@ ANDOVER

@ WINCHESTER

SOUTHAMPTON
@

Outcrop symbols

Upper Greensand

0” <_/| Lower Greensand
I

Purbeck Beds

IN THE ANGLO-PARIS BASIN 43

Bristow e¢ al. (1889), Jukes-Browne (1900), and Osborne-White (1921) all give a
roughly uniform thickness of 100 feet (30-48 m.) for the Gault throughout the Island.
However, this is certainly not correct for although the thickness is about 100 feet
(30-48 m.) at Redcliff, it is reduced to about 65 feet (19-81 m.) at Compton Bay, and
may be over 100 feet (30-48 m.) in the southern part of the Island. Kitchin &
Pringle (1922 ; 160-161) recognised that Middle Albian sediments of the ‘ interruptus
Zone ’ were present at Culver and in the south of the Island, but they considered that
the Gault in Compton Bay was wholly Upper Gault. They based their conclusion
on the old record of Inoceramus sulcatus (Norman 1887 ; 70), but it will be shown
below that this argument together with their view that the ‘ —so-called ‘‘ Carstone ”’
of this locality— ’ did not represent the true top of the Lower Greensand, is com-
pletely fallacious. Spath (1926b ; 422, and im Jackson 1939 ; 74) considered that the
Gault represented only the dentatus Zone and that the ‘ lower benettianus (=inaequin-
odum) zone ’ passed into the Carstone beneath. This latter conclusion was confirmed
by Casey (1961a ; 515) based on material collected by C. W. Wright and the author.

(a) Redcliff

The Gault in the hollow between Redcliff and the Upper Greensand face at the W.
end of Culver Down (SZ 62758550) is badly slumped and overgrown. However,
from time to time exposures of a few feet of clay dipping steeply NE. have been seen
near the top and the base of the Formation. The Gault is here about 100 feet
(30-48 m.) thick but it is impossible at this time to obtain an accurate measurement.
From sections exposed near the base it can be seen that there is a fawn band within
six feet (1-828 m.) of the junction with the underlying pebbly Carstone. This fawn
band contains crushed Hoplites (H.) spp. with pyritic films replacing the shells,
together with a few part-phosphatised specimens. On the basis of the ammonites
so far seen, a basal spathi Subzone age is indicated. However, a similar unit at Bon-
church has yielded rare but definite lyelli Subzone fossils.

Mr. J. McA. Hart collected a pyritised Euhoplites of Upper Gault aspect from a
small exposure near the top of the Gault and below Jukes-Browne’s Division A. This
indicates that at Redcliff the lower part of the Upper Albian is within the clay facies.

(b) Rookley

Perhaps the most important section available in the Isle of Wight at this time is
exposed in the extensive workings of Island Bricks Ltd., at Rookley. The sequence
extends from well down in the Carstone up into the Gault and is shown graphically
in text-fig. 19. It has never been described in detail but has been mentioned by
Pritchett & Jackson (1941). The pit is cut by an E.-W. fault down-throwing to the
south, and the southern part of the section is tectonically disturbed. The sequence
north of the fault dips NW. at 6° and shows variations in the thickness of certain
beds which cannot be ascribed to a later tectonic cause. These are here considered
to be due to slumping before consolidation of the sediment.

MIDDLE ALBIAN STRATIGRAPHY

Lithology Et Ins M

Weathered mottled grey clay,

oO
©
a

Ferruginous marl band with phosphatic nodules,

12 Dark grey clay streaked with ochreous seams, 3-4 re
8
4 Ochreous clay with occasional lenticles of ferruginous 1
marl & large spherical or elongated septarian phosphatic nodules ie}
10 Ochreous mottied clay passing down into Bed 9 below. 3 2 7
=
io]
Qa
“”
Lo | Highly ferruginous marl. = 9-10
6
8 Dark grey clay with partly phosphatised fossils the shells 2 3
being replaced by films of pyrite.
Fawn-grey clay with occasional scattered phosphatic
Zz nodules, At base part-phosphatised fossils occur with pyritic shells) — 10-12
ls | Dark grey clay. Ae a
awn grey clay,
5 F grey y. — 10-19
4
3 2
4 Dark gritty glauconitic clay,
7S 3
Fawn grey silty clay with burrows of dark grey clay and =
3 crushed pyrite replaced shells of /noceramus concentricus. S 1 3
2 Glauconitic dark grey micaceous clay with streaks of 3 2
glauconite sand, 2
- (il) Streaked grey gritty clay with pockets of sand & glauconite, = 5
(i) Lenticles of false-bedded glauconitic pebbly loam. RQ SS] — 0-6
TOP OF CARSTONE rn
Variegated dark and light brown ferruginous grit separated | > 3 8!
from Bed1 by a thin seam of iron pan. Top few inches with b Ae
5 many small pebbles and thin seams of pan. Gritty pale =
phosphatic nodules occur scattered at about middle. Nine 5 4 6
inches from base there occur crushed ammonites, rarely iy
part—phosphatised, with the shell, «
°

Fic. 19. Middle Albian sediments at Island Bricks Ltd’s Rookley Brickworks, situated
600 yds ESE. of the school, Rookley, and 150 yds NW. of the A 3020 road, South Arreton,
Isle of Wight (SZ 51338395).

IN THE ANGLO-PARIS BASIN 45

Bed 5 of the Carstone contains crushed filmy shells of ribbed ammonites. Very
rarely these are partly phosphatised and can be identified as Hoplites (Isohoplites)
spp. including H. (I.) eodentatus (e.g. BMNH., C 73358 author’s colln., figured Casey
1965 ; 538, text-fig. 202 g.h.) indicating an eodentatus Subzone age. Beds 1-6 of the
Gault have not yielded fossils and may represent the /yellc Subzone known to be
present in the Ventnor area to the south. Beds 7-11 can definitely be classified with
the spathi Subzone. Bed 7 contains crushed pyritic ammonites, some partly phos-
phatised, including the typical H. (H.) spathi and H. (H.) dentatus marking the base
of the spathi Subzone. Bed 8 also contains crushed Hoplites (H.) spp. with pyritic
tests which decompose very rapidly on exposure. Beds g—10 have not yielded fossils,
but Bed 11 has yielded a few fragments of large Hoplites (H.) which still suggest the
lower part of the spatht Subzone. No fossils were found in Beds 12-14.

(c) Compton Bay

In view of Kitchen & Pringle’s statement that no Lower Gault is present in Comp-
ton Bay (1922 ; 161) it was particularly fortunate that a good section has been ex-
posed during recent years. The true Gault here was stated by Strahan (7m Bristow
et al. 1889 ; 63) to be 95 feet (28-956 m.) thick excluding the passage beds to the
Upper Greensand but this is far in excess of the true figure. The very high angle of
true dip seen in the cliff, levels off sharply at no great depth below beach level due to
a slight flexure and change in direction of apparent dip. On a very accurate
measurement based on the detailed sequence given in text-fig. 20, the thickness
is little more than 65 feet (19-812 m.).

The phosphatic nodules at the top of Bed 8 of the Carstone have yielded Hoplites
(Isohoplites) eodentatus indicating that Subzone. No ammonites have been found in
either Beds 1 or 2 of the Gault and their exact Subzonal age is unknown. However,
the species of Hoplites (H.) in Bed 3 indicate the basal part of the spatht Subzone and
it is possible that both Beds 1 and 2 are of lyelli Subzone age. Apart from Bed 3, the
only other ammonites found were crushed Hoplites (H.) sp. at the base of Bed 10, at
which point a shelly facies appears. One is tempted to compare this junction be-
tween the pyritic facies below and the shelly facies above, with a similar junction in
the spathi Subzone sequence in the Nyewood-Wrecclesham area of the outcrop in the
Weald. However, such a correlation may well be more apparent than real. Beds 3
to the base of Bed 10 can, therefore, be classified with the sbatht Subzone. Howmuch
of the overlying sediments belong to the spathi Subzone is not yet known, as also
whether any other Middle Albian Subzones are represented. However, there is no
doubt that there is a substantial thickness of ‘ Lower’ Gault present at Compton
Bay.

(d) Ventnor to Niton

A complete section of the Carstone is exposed in the sea-cliff extending from
Dunnose south-westwards to the esplanade at Bonchurch ; the stretch of coast named

46

MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ins M
20
Deeply weathered grey clay the detailed lithology of which is
12 obscure. In the upper 1 foot the sediment consists of dark grey 6
clay This is followed by Division A of the Upper Greensand.
c
3
°
c
x
c
11 | Very glauconitic silty weathered clay. > fo)
10 Dark silty, micaceous, glauconitic, sparsely shelly clay becoming 0 15
more glauconitic inthe upper 3 feet.
We
Nn
Darkish fine-grained micaceous clay becoming darker upwards 0 1
2 with increase of the silt and very fine-grained glauconite fraction.
No fossils seen.
8 Fawnish silty clay containing plant remains at top. oss
BI
Bed 6 becomes tougher and more silty with scattered pockets
of glauconitic silt. Bed 7 in turn passes by wisps into Bed 8, 6
<
o
Q
“”
6 Darkish grey micaceous silty clay with obscure pyritic shell °
fragments.
5 Fawn gritty clay. fe)
5
Dark grey micaceous gritty clay. o
6
Hard blocky silty dark grey clay micaceous and weathering
battleship grey. . 2
rs
yy
Dark grey slightly mottled clayey fine grained micaceous | oam, =
weathering light grey. 5 :
BASE _ OF THE GAULT !
THE GAUL “
Glauconitic micaceous fine soft silty sandstone with pipes of dark grey clay; £3
bed weathe ferruginous. Small pebbies in clayey bottom 6-8inches, Sandy a) 9
9 of top. oc Oo

Fic. 20. Section of the Gault exposed in the sea-cliff about 200 yds NW. of Compton Chine,
Compton Bay, Freshwater, Isle of Wight (SZ 36708524).

IN THE ANGLO-PARIS BASIN 47

Monks Bay. Above this, the basal beds of the Gault are occasionally well exposed
at the top of the cliff, but the remainder of the Gault is badly slumped in the Under-
cliff itself. Other sections are known from Ventnor, Steephill, and from Reeth Bay,
Niton and round St. Catherine’s Point and are mentioned below. An ammonite
described by Spath as Anahoplites mimeticus (1925 ; 131, pl. X, fig. 7a, b) was alleged
to have come from the ‘Carstone’ of Niton, Isle of Wight. However, the preserva-
tion of the holotype BMNH., C 30535 indicates that it is definitely not from the
Carstone but from an horizon in the Gault possibly above the spathi Subzone sedi-
ments ; it is possibly a species of Hoplites (H.) with a smooth outer whorl. The
specimen figured by Casey (1966 ; 547, text-fig. 207a, b, C. W. Wright Colln., 9983)
as Anahoplitoides mimeticus Spath which he states came from the top of the Carstone
(eodentatus Subzone), of Bonchurch, is preserved in a manner quite unlike any of the
material from the top of the Carstone and C. W. Wright has informed me that it was
picked up loose. It must come from a higher horizon in the Gault than the proved
spatt Subzone sediments. However, Hoplites (Isohoplites) eodentatus does occur in
the top part of the Carstone at Bonchurch but the preservation is the typical sandy
phosphate. It might be mentioned here that the specimen of Hoplites vectensis from
the Carstone near Niton mentioned by Spath (1925a ; 128 L.F. Spath Colln. No. 238
= BMNH., C 30555) is indeed from the Carstone and is an external mould without
the peripheral area. Nonetheless, it is almost without doubt a specimen of H.
(Isohoplhites).

The few feet of Gault immediately overlying the Carstone at Monks Bay has
yielded, from a fawn band, crushed and partly phosphatised Hoplites (H.) spp.,
specimens of which are in the British Museum (Nat. Hist.) L. F. Spath Colln. An
external mould of a Beudanticeras sp. in an identical matrix is preserved in the
Institute of Geological Sciences (GSM., Zn 1483 also L. F. Spath Colln.) and indicates
the presence of the /yelli Subzone. A pyritic specimen of Lyelliceras is in fact known
from this locality and was figured by Spath (1931 ; 320 pl. XXXIII, fig. 15a-c,
BMNH., C 32845). Further evidence of the presence of sediments of lyell1 Subzone
age is provided by ammonites from the basal Gault near the Gas House, Gas House
Hill, Ventnor (SZ 56857747) preserved in the Sedwick Museum. These consist of
black brittle phosphate steinkerns without the shell, typical of the lower fawn band,
and include Beudanticeras sanctaecrucis and Hoplites (H.) spp., one of them (B 42586)
with pyritic inner whorls like that of the Lyelliceras mentioned above. C. W., &
E. V. Wright have recorded Protanisoceras moreanum from west of Luccomb Chine
(1942 ; 286).

More definite information about the higher part of the Gault is provided by the
section in the cliff SW. of Steephill Cascade, Ventnor (SZ 55467707). Here the
Gault dips seaward becoming much steeper in the foreshore landward of an old
slipped mass. Dark grey gritty pyritic clay with rolled pieces of phosphatised
Hoplites (H.) spp. occurs along the axis of the ‘ fold ’ and is overlain by sparsely shelly
clay in which ammonites have yet to be found. The cliff behind these foreshore
exposures is deeply weathered at this time.

MIDDLE ALBIAN STRATIGRAPHY

Lithology [ae Ins M.

= Weathered blocky marly clay with ferruginous streaks. 12 o 16

? intermedius
=e
w

Homogeneous light grey clay tending to be hard and blocky 16 6
with occasional streaks of silt. Pyritic shell seams occur
at 4.and11feet above the base and contain /noceramus

concentricus

Grey blocky micaceous silty clay with streaks of white silt at
base, A pyritic shell seam occurs 2inches from base.

Homogeneous grey silty clay.

Tough grey silt = 10 ©
4 {e)
Light fawny grey silty clay with pyritic fossils. 7

Grey silt,

Fawn clay with dark burrows,

spathi

Soft dark grey silty micaceous clay with some fine

_
a = wo ~
N SS %
a
> —
aa
=a
© oO

glauconite. 3 6 5
Tough silty homogeneous fawnish clay. 1 6
Bedded pyritic silty clay; the top 4inches being more silty a3 a 4
and glauconitic. 1 8
Blocky grey silty micaceous clay. NV, 1 6
= \ | 3
Obscured interval | | | 3 fo)
—--—- - - H-e-F Hee KH — —- ~~ ~— -- |], |! I
Mottled pyritic clay, = (= Scape 1 02
— -— = — ae. SS ss — =. foes Si Be Se ee eae a
esas '
Obscured interval | o. | I
with Was ie)
| i]
mottled weathered greyclay at the base wi! | :
—— 2
Dark grey-brown blocky loam with grey clay streaks, we
increasing in number upwards,and some black phosphatic nodules, es 2 os

Remainder of CARSTONE

Fic. 21. Gault section in cliff below old coast road 50 yds NW. of Cliff Cottage and
360 yds SSE. of Blackgang Hotel, Blackgang, Chale, Isle-of-Wight (SZ 48877644).

IN THE ANGLO-PARIS BASIN 49

(ec) Blackgang

The section in the Carstone and Gault at Blackgang shows that the sequence has
changed fairly considerably in the mere 44 miles from Ventnor. The section given
in text-fig. 21 exposed below Cliff Cottage, Blackgang, shows that the sediments are
generally coarser in grade. It is exposed at the summit of a cliff of Sandrock Series
and in wet weather is dangerous.

Bed 8 of the Carstone is probably of eodentatus Subzone age, but Hoplites (Iso-
hoplites) has not yet been found here. Unfortunately, the lower 8 feet (2-438 m.) of
the Gault is not clearly exposed but it certainly contains at least one fawn pyritic clay
band which has not yet yielded fossils. The only ammonite seen in Bed 3 was a
crushed Hamuites (H.) sp. 4 inches (0-101 m.) below the top which indicates the spathi
rather than the /yelli Subzone. Bed 12 has also yielded crushed Hoflites (H.) spp.
which indicate the spatht Subzone. No other ammonites have been found higher in
the section but J. concentricus still occurs 11 feet (3-352 m.) up in Bed 13.

A section below Gore Cliff about 825 yards SE. of Cliff Cottage shows higher Gault
(text-fig. 22). Unfortunately the two sections probably do not overlap although
there is an obscured interval of 12 feet (3°657 m.) at the base of the Gore Cliff sequence.
I. concentricus is present in the top 1 foot 6 inches (0-457 m.) of ‘ Bed 1’ showing it to
be still of Middle Albian age, but no other age indicative fossils are yet known from
this section.

Certainly some of the lower part of the Gault here is of spbathi Subzone age. How-
ever, the specimen of Hoplites aff. vectensis recorded by Spath (1925 ; 128, BMNH.,
C 890) and said to be from Blackgang is in fact identical in preservation to specimens
from the top of the spathi Subzone at Osmington, Dorset, from whence it undoubtedly
came.

(i) BALLARD CLIFF TO OSMINGTON

The early accounts of the Gault in this coastal area of Dorset were given by Strahan
(1898) and Jukes-Browne (1900). This work was revised by Wright (im Arkell
1947b ; 178-194), and there is no new information to add to his lithological account
for the sections have deteriorated considerably. However, his Subzonal classifica-
tion of the basal beds requires revision. Wright redescribes, as far as it is possible,
the sections at Punfield Cove, Swanage (SZ 03878110), Flower’s Barrow, Worbarrow
Bay (SY 86388045), Lulworth Cove (East SY 82867988, West SY 82427988),
Durdle Cove (SY 80578028), White Nothe, and Black Head, Osmington, covering in
all a distance of 35 miles (text-fig. 18).

Beds 1 and 2 at Flower’s Barrow, Worbarrow Bay, are apparently of the same age
as the ferruginous clay with concretions at Black Head, Osmington, as Wright stated.
However, the fauna from these sediments at Black Head, first recorded by Cunning-
ton (1929) is not of ‘ benettianus ’ Subzone age as Spath originally thought (Wright
im Arkell 1947b ; 181) for all the species of Hoplites (H.) which occur, such as H. (H.)
dorsetensis and H. (H.) vectensis, can be matched in the highest part of the spathi
Subzone in the Weald and elsewhere. Even Spath was to change his mind about the
Subzonal age of these two species, placing them in the imtermedius Subzone (1942 ;

’

D

50

MIDDLE ALBIAN STRATIGRAPHY
Bed Lithology Ins M
9
: Dark grey silty clay with wisps of glauconite and sand, shelly
with scattered pyrite nodules. o
8
z
5) Homogeneous dark grey clay with pyrite nodules and crushed
fossils with the shell preserved, 6
‘4 Blocky homogeneous dark grey clay with pyrite nodules and 4 6
occasional bivalves with the shell preserved.
Passing down at the base into;
Glauconitic silty clay with rafts of glauconite sand, 595
| Fawn clay with pyritic nodules, and phosphatic nodules at the top, 5
4
4p Dark grey glauconitic silty clay with Inoceramus concentricus 6
_ preserved with the shell. ean eed at
3
Sequence obscured by scree, 0 2
1
°

ExGs22;

Cliff section of Gault at top of undercliff below Gore Cliff, 480 yds ESE. of South
View House, Blackgang, Chale, Isle of Wight (SZ 49437590).

IN THE ANGLO-PARIS BASIN 51

675). In this, he was very nearly right, for the concretions at Black Head yield,
albeit very rarely, early forms of Anahoplites of the intermedius group. These
include the specimen figured by Spath as A. mimeticus (1927 ; 188, pl. XVII, fig. 8a, b)
which belongs to A. osmingtonensis sp. nov., and A. grimsdalei sp. nov. which is the
direct forerunner of A. evolutus a form found at the base of the intermedius Subzone.
These early Anahoplites are associated with the usual species of Hoplites (H.) of this
bed, however, it is important to note Cunnington’s remark (1929 ; 126) : ‘ Ana-
hoplites—are scarcer, but wherever one is found there are almost always others or
fragments of others in the same block’. Ihave not yet found Anahoplites in this line
of concretions containing an upper spathi Subzone fauna. This might be purely a
collecting error, for the matrix and mode of preservation of the specimens of Ana-
hoplites is identical to that of the other fossils of this bed and this is very distinctive.
From a morphological point of view these are unquestionably earlier than the basal
intermedius Subzone species of this genus, e.g. A. evolutus (p. 151). Equally certain,
they are not earlier than the high spathi Subzone concretions ; that is, they are not of
lyelli Subzone age. The sections in this area are shown in Arkell (1951 ; Fig. 4).

Deposits of Jautus Zone age are also present in this area (Cunnington 1929 ; 129 :
Spath 1943 ; 743 : Wright 7m Arkell 1947 ; 193) but the position in the section of the
material picked up loose has yet to be determined.

(iii) THORNCOMBE BEACON TO BLACK VEN

The account of the sections on Golden Cap, and Black Ven, given by Jukes-Browne
(1900 ; 182-189) was followed fourteen years later by the very important detailed
account by Lang (1914) who also discovered the formation on Stonebarrow. To this,
additional information was added in Lang & Thomas (1936), and these sections to-
gether with that on Thorncombe Beacon east of Seatown have more recently been
described by Welch (im Wilson, Welch, Robbie & Green 1958 ; 139-150). Here
again, there is nothing to add to the lithological account but a re-examination of the
ammonites has provided a little, but important, additional information. The
sediments classified with the Gault are much thinner in this area of the coast.

Bed 1 of Lang (1914) yielded Anahoplites praecox both at Black Ven and Stone-
barrow (1936 ; 310). Bed 2 on Stonebarrow has also yielded A. praecox including
one specimen BMNH., C 15661 which is very close indeed to the neotype of ‘ Dimor-
phoplites’ alternatus, which is in reality a very coarse development of A. praecox
(p. 153). A. praecox also occurs at Charton Goyle in a matrix which is almost cer-
tainly the unweathered representative of Bed 2 (BMNH., C 68394-6). Beds 1 and 2
definitely belong to the intermedius Subzone. According to Spath (e.g. 1943 ; 744),
Bed 3 contains both intermedius Subzone and varicosum Subzone ammonites. This
is untrue. BMNH., C 41035 from Bed 3, recorded by Spath as Epihoplites aff.
trifidus, is in reality an early transition between Hoplites (H.) and Dimorphoplites
consistent with an intermedius Subzone age. The ‘ Idiohamites of the turgidus
group’ (BMNH., C 41038) is a Protanisoceras (H.) cf. nodosum, also indicating an
intermedius Subzone age. Another specimen BMNH., C 41035 from Bed 3 is here
identified as Anahoplites cf. intermedius.

52 MIDDLE ALBIAN STRATIGRAPHY

There is, therefore, no evidence of the presence of Upper Albian sediments in
Lang’s Bed 3 and this is consistent with the distribution of Inoceramus concentricus
and J. sulcatus. I. concentricus is known from Beds 1, 2, and 3 (e.g. Lang Colln.,
BMNH., L 55076 from Black Ven) both on Black Ven and Stonebarrow. Specimens
of I. sulcatus are known from Black Ven (e.g. BMNH., L 55368-71 Grimsdale Colln.)
preserved without the shell in glauconitic sandstone which suggests Lang’s Bed Io,
certainly no lower horizon. The record of Anahoplites planus from Lang’s Bed 10 by
Spath (1943 ; 744) isanerror. The Middle-Upper Albian boundary falls somewhere
between the top of Bed 3 and ?the base of Bed Io.

The above indicates that Kitchin & Pringle’s statement that no Lower Gault
exists at Black Ven is quite fallacious (1922 ; 163). This statement is the more
incredible when one realises that they had never examined the section and apparently
had overlooked Lang’s paper of 1914. There may well be a non-sequence in these
sections but at this time there is no evidence to indicate its extent.

No sections further west on the Devon Coast have been examined by the writer.

(iv) CONCLUSION

The poor state of the sections in the Isle of Wight and on the Dorset Coast are
particularly tantalising, nonetheless, some results can be drawn from this incomplete
information. It is apparent that in the Isle of Wight the eodentatus Subzone is
represented at the summit of the Carstone ; the /yells Subzone is represented in the
lower part of the Gault at least in the southern part of the island ; and the spathi
Subzone sediments are well developed. The intermedius Subzone is probably repre-
sented but the conclusive proof is not yet to hand.

By Worbarrow Bay and Osmington the fossiliferous concretions very near the
base of the Formation are of uppermost spathi Subzone age, and there is evidence of
lautus Zone sediments near the top of the Gault in the Osmington-White Nothe area.
In the sea-truncated outliers between Seatown and Lyme Regis the lowest fossilifer-
ous sediments are of intermedius Subzone age. As one proceeds westwards from the
Isle of Wight, therefore, the lowest fossiliferous sediments become later in age (text-
fig. 23). The intermedius Subzone is widely represented by sediments in the deeper
parts of the Wessex Basin ; for example in the Petersfield and Winchester districts,
Hampshire (p. 69), at Didcot, Berkshire (p. 63), Devizes, Wiltshire (p. 60), and
possibly at Okeford Fitzpaine, Dorset (p. 56). Its presence in the Isle of Wight, and
in the Dorset coast sections as far west as Osmington should not be. discounted.
Across the Channel at Cauville on the French coast NE. of Le Havre, there occurs,
above a vemanzé bed of basal dentatus Zone age, even later sediments of niobe Subzone

age (p. 107).

C. The outcrop from Devon to Bedfordshire

There is no information available about Middle Albian sediments inland in the
area W. of the River Axe. Neither is there any further stratigraphical information
available at the outcrop from the valley of the Axe to Okeford Fitzpaine, N. Dorset,

a

IN THE ANGLO-PARIS BASIN 53

other than that recorded by Jukes-Browne (1900 ; 163-4), Welch and Robbie (in
Wilson, Welch, Robbie & Green 1958 ; 148-152)', Reid (1903 ; 34-35), Osborne-
White (1923 ; 49-50) and Smart (1955 ; 43-4). It is apparent that Middle Albian
sediments occur north-eastwards from the area of Beaminster for although am-
monites have not been found, IJnoceramus concentricus has been recorded from a
number of sections and in this context definitely indicate a Middle Albian age. From
the coast between Thorncombe Beacon and Black Ven (p. 51), Middle Albian
sediments thin markedly inland in places to less than Io feet (3-04 m.) to thicken
again in the area of Toller Porcorum (SY 562974) S. of Evershot, and also in the out-
crop N. of Evershot eastwards from West Chelborough (Welch and Robbie 7m Wilson
et al., 1958). The Gault continues to thicken eastwards and in the region between
Alton Pancras and Ansty, Smart records a thickness ranging from 25-35 feet (1955 ;
42-4). The increasing thickness is maintained up to Okeford Fitzpaine.

Ww B

Black Ven Osmington Isle of Wight Sussex

intermedius

(garly)

Fic. 23. Sketch section, from W. to E. across the Late Jurassic-early Cretaceous modified
Wessex basin in the area of the south coast, demonstrating the transgressive nature of the
Middle Albian sediments and the diachronous nature of the base.

It is very regrettable that no good sections are now available throughout this long
strip of outcrop (text-fig. 18). The sequence seen on the coast between Thorncombe
Beacon and Black Ven, with its development of the intermedius Subzone of the
loricatus Zone has passed in the area of Okeford Fitzpaine to a sequence showing the
development of the eodentatus, lyelli & spathi Subzones of the dentatus Zone. In this
latter area it is underlain by deposits of mammullatuwm Zone age absent at the coastal
section to the SW. This difference in sequence is made somewhat significant by a
comparison with the sections in the Pays de Caux in northern France (p. Ior), and is
discussed later in the section dealing with the conditions of deposition (p. 142).

1 It is worth stating here that the stream and river sections described by the Survey officers are only
well visible in periods of drought (cf. Dewey 1934 ; 42).

54 MIDDLE ALBIAN STRATIGRAPHY

(i) OKEFORD FITZPAINE (DORSET)

The section at the Okeford Brick & Tile Works situated about 4 mile E. of the
village of Okeford Fitzpaine on the road to Shillingstone is no longer visible. It was
first described by Newton (1896 ; 198 : 1897 ; 66-68) from notes and material provided
by the Misses Forbes and Lowndes, then by Jukes-Browne (1900 ; 162), Reid 1903 ;
34-35), and Osborne White (1923 ; 48). Text-fig. 24 is taken from the account given
by Newton (1897, 67-8) with additions from Jukes-Browne. Spath (1925 ; 73
pl. V, fig. 6) figured one of Newton’s specimens of ‘ Acanthoceras mammillatum’ as
Douvilleiceras inaequinodum (Quenstedt) so demonstrating the presence of the
inaequinodum Subzone to which Casey assigns the base of the Gault here (1961a ;
565) ; now included in the eodentatus Subzone. Spath also demonstrated the
presence of the benettianus Subzone (i.e. lyelli Subzone) by the occurrence of Hoplites
(H.) ‘ pseudodeluci’ Spath (type locality, 1925a ; 120) and forms close to H. (H.)
benettianus (J. de C. Sowerby) (1925a ; 117-8), together with Beudanticeras probably
laevigatum and ‘Anahoplites of mimeticus type’ (1926a ; 147).

The material described by Newton is in the British Museum (Nat. Hist.). A re-
examination of the fossils and a careful reading of Newton’s account (1897) has
provided the following important stratigraphical information. All the specimens
were undoubtedly indigenous and not semi-derived. The three fragments (BMNH..,
C 6856-8) recorded by Newton as Acanthoceras mammuillatum described by Spath as
Douvilleiceras inaequinodum could well belong to one partly phosphatised individual.
The nacreous shell was clearly preserved and the matrix adhering to it consists of the
bluish grey micaceous clayey sand with glauconite of Bed3. Thespecimen could not,
therefore, have come from Bed 2. The specimens of Ostrea leymeriet (BMNH.,
L 11579 figured specimen, L 11591) have traces of the same sediment adhering to
them as that of the specimen of D. inaequinodum. Moreover, internally there is the
same blackish phosphate. It would seem also that these come from Bed 3 and not
Bed 2 (cf. Newton 1897 ; 68).

If this is the case then Bed 2, which Jukes-Browne (1900 ; 163) considered to be a
separate lithological unit and from which he records no fossils, together with Bed 1
may correspond to sediments at Dinton in the Vale of Wardour which have yielded a
kitchint Subzone fauna (Casey 1956 ; 231, 196Ia ; 564). Jukes-Browne considered
that these two beds should possibly be grouped with the Lower Greensand. My
reading differs from that of Casey (1961a ; 565) who states that the Gault here rests
directly upon the Kimmeridge Clay.

Two fragments of large specimens of Hoplites (H.) spp. (BMNH., C 6859-60)
identified by Newton as Hoplites benettianus (1897 ; 70), one of which he figured, are
preserved one with and the other without the nacreous shell in a ferruginous brown
(weathered) and grey micaceous clayey sandstone. Bearing in mind Newton’s
remarks (1897 ; 67-8) this lithology indicates the upper part of Bed 4. The specimen
figured by Newton (1897 ; 70, pl. 2, fig. 1, BMNH., C 6860) was subsequently made
the holotype of Hoplites pseudodeluci by Spath (1925a ; 120-123) but due to its
crushed state and lack of inner whorls the true nature of it is impossible to determine.
Moreover, a specimen from Bed 5 at Badbury Wick shows a closely comparable outer

IN THE ANGLO-PARIS BASIN 55

Lithology

(li) Dark-grey micaceous silty clay with phosphatic nodules.

?spathi

(1) Shelly dark-grey silty clay with glauconite.

lyelli

Brown sandy rock with fossils inthe upper part.

Brown, green grey, and yellowish clayey micaceous loam, with
part phosphatised fossils with the shell preserved, and small
pebbles particularly In the lower part.

Brown sandy Ironstone breaking Into small lumps,

? kitchini

Clean green sand,

D
i)
a

€ — — — — ee ee - - - = EF - - - - -
eodentatus

KIMMERIDGE CLAY

Fic. 24. Section in Albian sediments (after Newton 1897, Jukes Browne 1900, & Osborne
White 1923) formerly exposed at the old Okeford Brick & Tile Works, immediately S. of
the Okeford Fitzpaine to Shillingstone road, 800 yds E. of St. Andrew’s Church, Okeford
Fitzpaine, Dorset (ST 81501080).

56 MIDDLE ALBIAN STRATIGRAPHY

whorl but the inner whorls are those of H. (H.) bullatus Spath. Bed 4 can definitely
be classified with the lyelli Subzone even if no specimens of Lyelliceras, Beudanticeras
or Protanisoceras have been preserved. A more typical lyelli Subzone fauna was
recovered from the lower part of Bed 5. The ammonites described by Newton are
preserved in two distinct lithologies, although they all possess remains of the nacreous
shell and, except for one, are preserved as partly crushed clay steinkerns. They were
re-identified by Spath as follows, to which my own comments are added.

Hoplites interruptus Bruguiére

(1) Hoplites pseudodeluct Spath (BMNH., C 6864) figured by Spath (1g925a ; 121,
pl. X, fig. 6) which is preserved in an identical manner to those of Bed 4 from which it
probably came.

(2) Hoplites sp. transitional between benettianus & paronar (BMNH., C 6863)
according to Spath (1925a ; 115, 118). This specimen which has pyritic inner whorls
is crushed ventrally, having come to rest on the sea floor on its venter. It is pre-
served in fawn clay with glauconitic loam filled burrows. It is specifically indeter-
minate.

(3) Hoplites sp. benettianus, baylet group (BMNH., C 6862) (1925a ; 118) probably
does not belong to either of these two species.

(4) Newton’s figured specimen (BMNH., C 6861), identified by Spath as Hoplites
dentatus (J. Sowerby) (1925a ; 118), is preserved crushed in glauconitic sandy dark
grey clay.

Hoplites splendens J. Sowerby

(5) The two small specimens (BMNH., C 6866-7) were identified by Spath as
Beudanticeras probably laevigatum. They are preserved in exactly the same type of
matrix as (2) above.

(6) The larger specimen (BMNH., C 6865) was identified by Spath (1926a ; 147)
as Anahoplites resembling A. mimeticus. It certainly is an Anahoplites, preserved in
the same type of matrix as (4) above, and its occurrence is discussed below.

Hamites sp.

(7) In his description of Hamites attenuatus Spath (1941 ; 611 footnote) referred to
the two specimens indicating that they might have been tuberculate and, therefore,
generically distinct. A close examination shows that they are both Protanisoceras
sensu-stricto, one of them (BMNH., C 6868) being P. (P.) barrense, the other (BMNH.,
C 6869) closely comparable to that species. They are both preserved in the same
lithology as (2).

The specimens (Nos. 2, 5, & 7 above) preserved in fawn clay with darker glauconitic
burrows are certainly a lyelli Subzone assemblage. The two specimens (Nos. 4 & 6)
preserved in glauconitic sandy dark grey clay suggest, however, a high spatht Subzone
age. It has been suggested above (p. 47) that the type of Anahoplites mimeticus is a
spathi Subzone species of Hoplites (H.) and certainly did not come from the Carstone,
of the Isleof Wight. The association of Anahoplites of the osmingtonensis-grimsdalet
group with species of Hoplites (H.) in the Osmington area of Dorset which Spath

|
|

IN THE ANGLO-PARIS BASIN 57

(1926b ; 422) considered to be of benettianus Subzone age in fact marks the extreme
summit of the spathi Subzone (p. 51). This association is also to be seen at the
summit of the spathi Subzone at Caen Hill, Devizes (p. 60) and probably also at
Dilton Marsh.

(i) VALE OF WARDOUR TO DEVIZES (WILTSHIRE)

(a) Vale of Wardour

No sections in Middle Albian sediments are now to be seen in the Vale of Wardour.
A brickyard was worked throughout much of the 19th Century near Ridge and was
first described by Fitton (1836 ; 247) and was listed by d’Orbigny (7m Geinitz 1849) as
‘Rudge’. Jukes-Browne (1900 ; 230), Reid (1903 ; 32, 39), and Andrews (in Andrews
et al., 1903 ; 158) provided further information about this section. Fitton’s ‘ Am-
monites rhotomagensis ’ suggests Lyelliceras lyelli indicating the lyell1 Subzone but
the specimen has not beentraced. Both Fitton and Jukes-Browne record Inoceramus
sulcatus as well as I. concentricus from this area and the varicosum Subzone is certainly
present in marls at the Watercress beds Fovant (Mottram 1957 ; 166, 1961).

The only other sections are a well at Dinton described by Jukes-Browne & Andrews
(1891 ; 292 & 1900 ; 228 : and mm Reid 1903 ; 31, 38), and an exposure of the basal
beds in Wardour Park (Reid 1903 ; 34, Mottram 1957 ; 161). Casey demonstrated
(1956 ; 231, 1961a ; 565) that the lower part of the sequence in the Dinton well was of
kitchini Subzone (Lower Albian) age and was overlain non-sequentially by clays of
dentatus Zone age. However, no fossils have been preserved from the clays and it is
impossible to determine their subzonal position. At the present time it is not possi-
ble to correlate these three sections with each other, or any section south or north of
the Vale.

(b) Maiden Bradley to Devizes

No sections now exist in this area of the outcrop, although brick pits formerly
existed at Redford Water, Flintford, Crockerton, and Westbury, all described by
Jukes-Browne (1900 ; 235-6). From his account of them it is possible to gain some
idea of the lithological sequence in the lower part of the Gault in this area (text-fig. 25).

The Crockerton section was worked in the early part of the rgth Century and
yielded to Miss Benett the holotypes of Ammonites benettianus and Ammonites

_ laevigatus described by J. de. C. Sowerby. It also probably yielded the specimen of

Ammonites monile mentioned by Fitton (1836 ; 258) and a good deal of the English
lyella Subzone ammonites in the various collections used by Spath in his Monograph
of the Ammonoidea of the Gault. Until the late 1930’s it was the only section known
in England to have exposed sediments definitely containing Lyelliceras lyelli. There
is now a factory on the site.

Unfortunately, there are no detailed accounts either of this section or the others
mentioned above. However, Jukes-Browne’s account (text-fig. 25) suggests that
the sequence is fairly uniform. It appears that ‘ Division’ 3 definitely yielded lyelli

58 MIDDLE ALBIAN STRATIGRAPHY

Subzone fossils and by comparison with Caen Hill, Devizes (p. 60), probably was the
source of some spathi Subzone ammonites also known from Crockerton.

In the Westbury area, the Eden Vale Brickyard described by Jukes-Browne (1900 ;
236) is no longer exposed, but the pit worked by the Westbury Potteries Ltd., has
shown sections of the basal beds of the Gault from time to time. Casey (1956 ; 233 :
1961a ; 564) has referred to this section as the Bremeridge pit demonstrating the
presence of the kitchini Subzone overlain non-sequentially by the basal beds of the
Gault of dentatus Zone age. This pit is presumably the source of the specimen of
‘ Anahoplitoides’ from Dilton (Ponsford Colln.) illustrated by Casey (1966 ; 547
text-fig. 207c).

REDFORD WATER FLINTFORD CROCKERTON EDEN VALE

M Ft
9-730

25

3 3 3

6720

15

?

ate Za Lk 4 ee

: 1 1
o+0 er

Fic. 25. Possible correlation of sections in the Warminster area of Wiltshire described by
Jukes-Browne (1900 ; 235-6).

(c) Caen Hill, Devizes

The area W. of Devizes has been a centre of brick and tile production since the
latter half of the last century. The sections then exposed at Caen Hill and Dunkirk
were described by Jukes-Browne (1892 : Ig00 ; 249-250, 252: 1905 ; 15-16) and
Osborne-White (1925 ; 39). Old collections in the British Museum (Nat. Hist.) and
Institute of Geological Sciences indicate the presence of the ?eodentatus, lyelli &
spathi Subzones in this area, and some of the ammonites were described by Spath

(1923-1925).

IN THE ANGLO-PARIS BASIN

Bed Lithology

Light grey weathered clay with limonitic concretions after pyritic ?fossils,

Weathered grey clay.

Pyritic dark grey clay weathering ferruginous.

Weathered blocky silty micaceous dark grey clay with bleached areas at
parting planes.

Poorly bedded massive micaceous silty clay, rather more silty in the lower
1foot Sinches, with many pyritic filaments. A snell seam occurs 3 feet
from the base and contains brown phosphatic concretions. The clay, darkish
grey in colour, contains scattered shells with a few large crushed ammon-
ites in the sediments above the 3 feet level.

grey clay.

(iii) Tough silty glauconitic dark grey clay with a few scattered shells.

(ii) Part-phosphatised ammonites &cementstone nodules in burrowed clay.
(i) Passage bed between lithology of Beds 4and S({iii),

Fawn silty clay with glauconite-sand infilled burrows.

Finely silty micaceous clay, coppery brown in colour when wet and fresh,
with patches of dark grey clay. Both are cut by fawn coloured burrow
infillings 1m top Binches. Pyritic filaments occur throughout; shelly
especially in the upper part.

Prominent and persistant seam of sandy ferruginous marlistone.

Medium grey clay with some phosphatic nodules.

\
(v) Blocky massive grey clayey silt, weathering white. The top 12inches
contains more clay.

(iv) Dark grey silty clay with filaments.

(Gi)Grey clayey silt with pyritic filaments; passing at the base into 8(ii),

(ii) Sparsely shelly glauconitic grey clay with pyritic filaments. in
>

(i) Very silty micaceous clay with pyrite and lenses of siltstone. =
Ww
=
&
wu
~
=

Brownish fawn silty clay, extensively burrowed, with infillings of dark Wy

SPATHI

42

10
os

14—

134

12

|

10

244

Fic. 26. Section in Gault at Messrs Hills of Swindon Ltd’s Caen Hill brickyard, c. 180
yds ENE. of the Olive Branch Inn, in the W. side of Caen Hill, Rowde, Wiltshire (ST

98246135).

60 MIDDLE ALBIAN STRATIGRAPHY

The presence of the eodentatus Subzone is suggested by Cleontceras? devisense
Spath (19238 ; pl. IV, fig. 7a, b) and Hoplites cunningtoni Spath (1923< ; 109, pl. VIII,
fig. 8a, b). Specimens of Cleoniceras (C.) are known from the eodentatus Subzone in
France but they are very rare and Mrs. P. Jennings has obtained one undoubted
lyelli Subzone example from Badbury Wick. Hoplites cunningtont isa late Otohobplites,
a genus also known to occur in the eodentatus Subzone elsewhere in southern England
and in France. The /yelli Subzone is indicated by ammonites such as Beudanticeras
laevigatum and Hoplites (H.) baylet, and the spatht Subzone sediments are still ex-
posed. The most important result of the present study is the discovery of a
reasonably thick sequence of sediments of intermedius Subzone age at Caen Hill. This
occurrence and that at Didcot (p. 63) represents the first time that the subzone has
been recorded in this area of the outcrop in Wiltshire and Berkshire.

Spath (1943 ; 745) followed Osborne-White (1925 ; 39) in considering that the
lautus Zone was present in the Gault of the Devizes area. This opinion was based
on material obtained by Cunnington from the long since vanished brickpit at Dun-
kirk. However, Jukes-Browne (1900 ; 252, 1905 ; 16) lists Inoceramus sulcatus as
well as J. concentricus and it is quite possible that this indicates the orbignyi Subzone.
This particular record of the Jautus Zone should be treated at this time with caution.

The brick pit at Caen Hill is still in work. It was formerly owned by the Devizes
Brick & Tile Co. Ltd., but now by Messrs Hills of Swindon Ltd. It is situated on the
W. side of Caen Hill and has been worked eastwards into the hill exposing the section
given in text-fig. 26. A dip of 3° towards the E. is present. The lyelli Subzone
sediments are seldom exposed now, and no satisfactory section has been seen by the
writer. It is possible that ‘ Division’ 4 (text-fig. 25) of the Warminster-Devizes area
is the same as Bed 2 of my section. If this is the case then the pre-spathi Subzone
sediments are thick at Caen Hill.

Beds 1 to 5 contain Hoplites (H.) spp., such as H. (H.) dentatus and H. (H.)
maritimus, indicating the spatht Subzone together with a good benthonic fauna of
bivalves and gastropods. A shell seam 3 feet (0-914 m.) above the base of Bed 7
contains crushed evolute Anahoplhites of grimsdalet type and ribbed forms comparable
to A. evolutus and A. osmingtonensis together with occasional specimens of Hoplites
(H.). This association continues through part of the remainder of Bed 7, and these
sediments are here considered to be a little later than the concretions in the Osming-
ton area, Dorset (p. 51) which are classified with the uppermost part of the spat
Subzone and may be represented at Devizes by Bed 6 and the basal 3 feet of Bed 7.
However, at Devizes in Bed 7 the situation is reversed, Hoplites (H.) being here sub-
ordinate to Anahoplites, and so these clays are classified with the basal part of the
intermedius Subzone. This level is, therefore, closely comparable to the basal part
of the intermedius Subzone in the Départements of the Meuse (p. 88) and Aube (p. 93).
In the Weald, this interval does not contain ammonites.

The remainder of Bed 7, and particularly 8, contains a typical intermedius Subzone
fauna with crushed Anahoplites praecox and A. intermedius. No fossils were ob-
tained from Beds 9-11 and their subzonal classification is, therefore, unknown at this
time.

ee

(2 — &- = eae a> a0

IN THE ANGLO-PARIS BASIN 61

(i) DEVIZES TO THAME (OXON)
(a) Badbury Wick (Wiltshire)

There is no information available at the outcrop between Devizes and the Swindon
area, a distance of about 18 miles (text-fig. 18). Messrs Hill’s of Swindon Ltd’s pit at
Badbury Wick has shown a section in spatht Subzone sediments for some years, but
in 1967 the pit was deepened and exposed sediments of lyelli Subzone age. The
sequence now exposed is shown in text-fig. 27. Although this is the first time in this
century that a lyell1 Subzone sequence has been well exposed in Wiltshire, it is
apparently different from that of Caen Hill and further south-west (text-figs. 26 & 25).
A working near the present pit was mentioned by Ramsey, Aveline & Hull (1858 ;
33).
Beds 1 to 6 contain a typical lyelli Subzone fauna but in contrast to Small Dole
(p. 38) Lyelliceras lyelli and Brancoceras spp. occur only infrequently and the bulk
of the fossils are crushed. The commonest ammonites are the heteromorphs such as
Protanisoceras (P.) barrense and P. (P.) alternotuberculatum together with Beudanticeras
laevigatum, and Hoplites (H.) spp. The facies is a shelly one, albeit sparsely in places,
with in general a better developed benthos than that seen in Sussex. Ammonites are
apparently rare in Bed 6 which otherwise contains very well preserved but fragile
bivalves and gastropods. That it still belongs to the /yelli Subzone is indicated by
the occurrence of Beuwdanticeras spp. as well as Hoplites (H.) spp.

Beds 7 to 11 are classified with the spathit Subzone. Bed 7 shows the major change
in the ammonite fauna which marks the base of the sfathi Subzone, and these now
consist of species of Hoplites (H.) such as H. (H.) dentatus and H. (H.) maritimus sp.
nov. associated with the bivalve Inoceramus concentricus in shell seams. The
benthonic fauna is very reduced in comparison with the lyelli Subzone sediments
below. Bed & contains the same fauna with individuals partly phosphatised with the
shell, while in Bed 9 the fossils are again crushed flat. In Beds 10 and 11 the shells
are replaced by pyrite and the non-ammonite element of the fauna becomes un-
common. No fossils have been found in Bed 12 and its age is uncertain.

The lithological sequence in the spathi Subzone is quite different from that of Caen
Hill, Devizes (text-fig. 26), where there is no pyritic facies in any part of the spathi
Subzone sequence and a good benthos is present throughout. The sequence at
Badbury in this Subzone is surprisingly reminiscent of that exposed in the Nyewood-
Selborne area of the western margin of the Weald (Owen 1963a). However, there,
the situation is somewhat reversed, the pyritic facies encompassing the sediments up
to and including the two ferruginous marly bands, the shelly facies prevailing in the
higher beds.

(b) Badbury to Thame

The Gault outcrop in the Vale of White Horse (Berkshire) has been discussed
principally by Hull & Whitaker (1861), Jukes-Browne (1900 ; 268) and Arkell (19472 ;
167-9). No sections now exist either at Uffington or in the area N. of Childrey.
Arkell recorded the discovery of specimens of Dimorphoplites by Mr. C. W. Wright in

62 MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology

Ins M.

Micaceous, glauconitic, very clayey burrowed silt.

Weathered light-grey pyritic clay with a few shell seams,
the shells being replaced by pyritic films.

Ferruginous pyritic marl, weathering rusty, with phosphatic nods.

spathi

Fawnish-grey slightly micaceous shelly clay.

Fawn marly clay weathering ferruginous with part-phosphat ised
fossils and a few phosphatic nodules, The bed is shelly.

Shelly highly glauconitic dark grey clay with more
glauconitic silty bands, The clay is micaceous, and pyritic
filaments occur scattered throughout. Pockets of glauconite
occur also, together with some burrows infilled with grey
7 silt. Although still shelly, the basal 18 inches does not
contain ammonites and is immediately overlain by a 12inch
seam of highly glauconitic silty clay. A bedding plane 4
inches below the top contains pyritised ammonites with their
shells preserved,

Dark grey gritty micaceous shelly clay becoming more plastic
downwards,

Fawn grey plastic clay, shelly, with a few burrows, Scattered
black septarian phosphatic nodules at top, anda few similar
ones occur to 1 foot below the top. A few pyritised ammon-
ites occur scattered throughout, together with large part
phosphatised ammonites in gritty concretions 4-Sinches from base

Tough dark grey gritty clay shelly with large black phosphatic
nodules at top, & scattered septarian phosphatic nodules at base!

tyelli

Fawn grey plastic, silty, micaceous, sparsely shelly clay, with
3 numerous pyritic filaments and part-phosphatised fossils
at base.

Brownish fawn shelly clay, burrowed, with dark fawn
infillings; passing down into Bed 1.
eas)

Dark fawn-grey clay.

Fic. 27. Section in Gault at Messrs Hills of Swindon Ltd’s Badbury Brickworks, W. of Day
House Lane, 200 yds SW. of Badbury Wick House, Chiseldon, Wiltshire (SU 18828160).

IN THE ANGLO-PARIS BASIN 63

the Childrey section (1947a ; 169) and I have been kindly allowed to re-examine them.
They are, as identified by Mr. Wright, a specimen of Dimorphoplites biplicatus
(C.W.W., 9762) and a specimen of D. glaber (C.W.W., 9761). Together they
indicate an horizon somewhere between the base of the meandrinus and the top of the
daviesi Subzones.

Nine miles ENE. of the Childrey brickyard is the famous section at Culham (Oxon).
This pit situated approximately 200 yds. N. of the River Thames, 425 yds. E. of the
road bridge over Culham Cut and 700 yds. SSW. of Culham College, Culham, Oxon
(SU 51159487) was in work from the middle of the last century until the late 1940’s.
It has been described by Phillips (1860 ; 548-550, 1871 ; 426-428), Jukes-Browne
(tg00 ; 268-9, 1908 ; 13-14), Osborne-White (1904 ; 300-304), Treacher (1908 ;
548-550) Pringle (1926 ; 101-2), and Arkell (1947a ; 169-170). Osborne-White’s
account paved the way for the more detailed description given by Pringle, but the
section is now badly degraded and overgrown and it is not possible to confirm or deny
Pringle’s subzonal grouping repeated by Spath (1943 ; 745-6). Neither is it possible
to make a direct comparison with the sequence at Badbury Wick. From the list of
fossils collected by Osborne-White (Jukes-Browne & Osborne-White 1908 ; 14) from
Bed 3 (= Bed 1 of Pringle 1926) it seemed possible that either the mammillatum Zone
was present or possibly the eodentatus Subzone. Douvilleiceras does range up into the
lyella Subzone but is rare in that Subzone in England except at Shere. Casey,
who has revised the identification of the ammonites, is in favour of a basal dentatus
Zone age (19g61a ; 565), and it is significant that Osborne-White records ‘ Hoplites
interruptus ’ just above Bed 2 (1904 ; 304). Beds 2 and 3 of Pringle were classified
by him with the ‘ benettianus Subzone ’, and Beds 4 and 5 with the spathi Subzone.
From the ammonites preserved in the various collections it is apparent that both the
lyella & spathi Subzones are present.

It is interesting to note that in the remnant of Gault formerly exposed at the
abandoned Chawley Brickyard, Hurst Hill (SP 47550420), Cumnor, Pringle (1926 ;
g8, 103) recorded ammonites which he considered to be characteristic of Bed 4 at
Culham. This section was situated 7 miles NNW. of Culham and has also been
discussed by Arkell (in Richardson, Arkell & Dines 1946 ; 104). If the equivalents of
Beds 1 to 3 are truly absent in this outlier, this is of some palaeogeographic significance.

The only other useful information yielded by this area is provided by two borings,
Nos 6 and 21, drilled for the new Central Electricity Generating Board Didcot Power
Station (SU 51289174 & 51339194 respectively). Together, the fragments of these
two cores show the following features in the sequence. The orbignyi Subzone is in a
Gault facies, indicated in Boring No. 21 by a fragment of core containing Inoceramus
sulcatus preserved in mid-grey micaceous silty clay with lighter coloured burrows
from a height of 122 feet 6 inches (37:34 m.) above the base of the Gault. The
intermedius Subzone is also present at a height of 68 feet (20-73 m.) above the base of
the Gault in Boring No. 6 where the core yielded a crushed specimen of Anahoplites
praecox preserved with a pyrite-replaced shell in silty brownish grey clay (cf. Bed
8 ii at Caen Hill, Devizes, p. 60). Boring No. 21 shows that the spathi Subzone with
Hoplites (H.) spp. is certainly present in the sequence between 42 feet 6 inches and
17 feet 6 inches (12:95-5:33 m.) above the base of the Gault. The pieces of core

64 MIDDLE ALBIAN STRATIGRAPHY

preserved are from 42 feet 6inches, 35 feet, 32 feet 6 inches, 30 feet and 22 feet 6 inches
above the base of the Gault and the lithology consists essentially of dark fawn-grey
silty clay with shelly fossils. There is no need to emphasise that the cores are very
incomplete and that no Subzonal limits can be deduced.

Sections in the Gault were formerly exposed in the area of Thame (Oxon) and
mentioned by Davies (1899b ; 160). The only information about the zonal strati-
graphy is the comment by Spath that at Priestend (SP 691055) the lower Gault,
which was exposed for at least 40 feet (12-16 m.), contained plentiful impressions of
ammonites of the dentatus Zone often of unusually large size (1943 ; 746).

(iv) THAME TO LEIGHTON BUZZARD (BEDFORDSHIRE)

The published information on the stratigraphy of the Gault between Thame and
Leighton Buzzard, a distance of about 19 miles is again not particularly satisfactory.
This is due mainly to the paucity of good exposures and, to a certain extent, to the
controversies which have tended to colour the accounts. Itis apparent that deposits
of Middle Albian age are present throughout the area, although they are greatly
reduced in thickness in comparison with the alleged sequence at Thame. The
Gault rests in this area either upon the Kimmeridge Clay, Portland, Purbeck, or
Lower Greensand deposits.

(a) Long Crendon (Bucks.)

No section in the Gault now exists at Long Crendon, which is situated about 24
miles towards the NW. of Thame, but sections in this outlier were described by Jukes-
Browne (1900 ; 277), Davies (1899a ; 22), Lamplugh (1922 ; 40-44), and Kitchin &
Pringle (1922 ; 164-5). The sequence has also been discussed by Kitchin & Pringle
(Ig21a ; 62 : 1922 ; 284-5), Spath (1943 ; 746) and Casey (1961a ; 569). Kitchin &
Pringle (Ig21a ; 62 : 1g2Ib ; 174 see also Spath 1943 ; 746) considered that the Upper
Gault rested directly upon Purbeck Beds here but the detailed evidence to sub-
stantiate this conclusion was not given.

On three counts it appears certain that Lower Gault is present. It is important
to note that Davies (1899a ; 22 : 1899b ; 161) recorded I. concentricus from the 8 feet
of Gault then exposed, but no ammonites were discovered and, therefore, the exact
age still remains uncertain. In the main outcrop to the S., the lower part of the
Upper Albian is quite fossiliferous with orbignyi Subzone ammonites and the ubiquit-
ous Inoceramus sulcatus. Also Lamplugh (1922 ; 40-44) demonstrated that a thin
development of Shenley Limestone was present below the Gault (see also Casey
1961a ; 569). It seems probable, therefore, that Davies’ record of I. concentricus is
correct, that these clays are of Middle Albian age, and that there is no overlap of
Upper Gault in this area as Kitchin & Pringle held.

(b) Haddenham (Bucks)

Although no section exists in the main outcrop between Thame and Aylesbury,
two ammonites are preserved in the Buckinghamshire Country Museum, Aylesbury,

IN THE ANGLO-PARIS BASIN 65

which are labelled Haddenham. Although the circumstantial evidence indicates that
this is the Buckinghamshire Haddenham, this locality is not on the Gault whereas the
Cambridgeshire locality is. With this reservation in mind these ammonites (W. J.
Welford Colln., accession No. 176-24) are here identified as Euhoplites aff. meandrinus
and Dimorphoplites aff. niobe (the late mutation known from the upper nodule bed
of Bed IV at Folkestone). Both are preserved as incomplete blackish phosphatic
casts without the shell, and indicate a distinct nodule bed or clays with scattered
nodules of late lovicatus Zone age within the Gault of this area.

Davies (1899a ; 55-56) disputes that the Gault was exploited S. of Haddenham
Low situated about 14 miles NE. of Haddenham (See also Balance 1964 ; Map 2).
The main outcrop, however, is at no great distance to the E. of Haddenham. No
information has been published about Middle Albian sediments in the 6 mile tract
of country between Haddenham and Aylesbury, although Balance (1964 ; 396) has
reported the Lower Gault to be present throughout the area.

(c) Aylesbury (Bucks)

The basal part of the Gault was formerly exposed in the Walton Cutting on the
Metropolitan railway line (L.T.E.). The cutting extends SSE. from the bridge
(SP 823130) carrying the B 4443 (Stoke Road) over the railway line, Walton,
Aylesbury. It was described by Pringle & Chatwin (im Sherlock 1922 ; 9) who
reported that the basal bed of the Gault rests directly upon the Portland Beds. No

Bed Lithology Ft Ins M
Fi Mottled fawn shelly clay marked by a shell seam at the 3
base with part phosphatised fossils. < 4 10
3
a
a)
| 2
Dark grey shelly clay with scattered phosphatic nodules 2 2 north
becoming glauconitic and gritty downwards In basal 10 inches | to
the sediment has become very glauconitic and gritty in current | 3 6south
swirls of glauconite sand.
“.
a5. | Scattered black septarian phosphatic nodules in glauconitic clay. wy enLoms eulatiel = aac
v
rae | Weathered grey glauconitic clay, l BN ele 4-5
| 3 | whitish phosphatic nodules with a few black septarian phosphates — Bre gee te 2-3
Ee
Glauconitic gritty loam brownish in basal 1foot with SERIE! 3 2 0
brown phosphatic nodules. \z
S|
at | Ferruginous yellow gritty clay resting on guttered surface of:- | e! |= 7 a

Portland Beds

Fic. 28. Section exposed in trench dug by Messrs S. A. Leach & Co. along the field bound-
ary extending for about 150 yds from SP 8235012425 to SP 8245012333, approximately
midway between the B 4443 Stoke Mandeville road and the L.T.E. Metropolitan railway
line from Aylesbury to Stoke Mandeville, Aylesbury, Buckinghamshire.

E

66 MIDDLE ALBIAN STRATIGRAPHY

fossils were found, and indeed, the only fossils recorded from the lower part of the
Gault in this area were obtained from a nodule bed stated to occur at about 10 feet
(3:04 m.) above the base of the formation (Pringle & Chatwin im Sherlock 1922 ; 9).
This bed has yielded an upper Albian fauna (Jukes-Browne 1900 ; 278 whose inclusion
of this horizon in the Lower Gault was an error : C. W., & E. V. Wright 1939 ; 115-
116 : Spath 1943 ; 746). Kitchin & Pringle took this fact to confirm an extreme
conclusion, stating that the Lower Gault had been overstepped by the Upper Gault
in this area (1922 ; 164-5). Pringle & Chatwin (7m Sherlock 1922 ; 8) were more
objective in that they pointed out that there was no evidence at that time for the
presence of the Lower Gault. Spath (1943 ; 746) although vague was a little more
cautious.

Important new information, demonstrating the presence of the spathi Subzone in
the lower beds of the Gault, was obtained by the writer from a trench dug by Messrs.
S. A. Leach & Co., during the laying of sewer pipes in January 1967. This very
temporary section was situated only a few hundred yards S. of the Walton Cutting
and the sequence is shown graphically in text-fig. 28.

No fossils were obtained from Beds I-5 and their exact age is uncertain. Bed 6
yielded a small bivalve fauna consisting mainly of a small Ostvea and a few specimens
of Nucula together with a single example of Neohibolites minimus. The shell seam
at the base of bed 7 contains numerous Inoceramus concentricus and some partly-
phosphatised Hoplites (H.) with simple dentatus ribbing, all with the nacreous shell.
The species include H. (H.) dentatus and H. (H.) cf. maritimus some being quite large
—6 to 8 inches in diameter, and the specimens of J. concentricus are also of large size.
A few specimens of Dentalium and Nucula also occur. This same faunal assemblage
occurs in the remainder of Bed 7 but it is crushed flat. The ammonites indicate the
lower part of the spathi Subzone, and it is unfortunate that no higher levels were
exposed at this locality.

Another ‘ cut and fill’ trench exposing higher beds was excavated in February
1967 in the area approximately 1475 yds a little S. of E. of the trench described
above. This trench was dug to a depth of 6 feet and extended from a point about
600 yds NNE. of Stoke Grange to a point about 400 yds. ENE. of Stoke Grange
(SP 8357512340 to SP 83801190), to the NE. of the A 413 (Wendover Road), Ayles-
bury. The trench was apparently cut in the direction of strike and exposed weathered
dark blue-grey clay with small buffish phosphatic nodules and patches. No
determinable fossils were found except at the field boundary (SP 83621225) where
two very badly preserved fragments of ammonites were seen. One was comparable
to Dimorphoplites, the other an equally poor fragment of Euhoplites. Together they
suggest an horizon below that exposed in the trenches along the A 41 in the Aston
Clinton area described by Wright & Wright (1939) which at certain points yielded an
orbignyt Subzone fauna.

There is no doubt that the thickness of sediments below the orbignyi Subzone
nodule bed in the Aylesbury area exceeds Io feet (3-04 m.) and it is likely that there
is a good deal more.

IN THE ANGLO-PARIS BASIN 67

LITTLEWORTH

5 AYLESBURY

15

BILLINGTON
CROSSING

SOUTHCOTT

spathi

eodentatus

condensed
floridum,
kitchini &
regularis

regularis

Woburn
Sands

Portland Kimmeridge
Beds Clay

Fic. 29. Correlation of sections between Aylesbury and Leighton Buzzard.

68 MIDDLE ALBIAN STRATIGRAPHY

(d) Aylesbury to Leighton Buzzard (Beds.)

At the outcrop between Aylesbury and Leighton Buzzard, a distance of about 10
miles, there is very little information about the stratigraphy of the Gault except for
borehole records e.g. Jukes-Browne (1900). A brickyard at Littleworth (SP 881233),
Wing, Buckinghamshire, described by Jukes-Browne (1900 ; 278), Davies (Igor ; 140,
1915 ; 92) and Lamplugh (1922 ; 89-90), is now badly degraded, Lamplugh (1922 ; 40)
records Inoceramus concentricus from the ‘10 to 15 feet of shattery dark-blue
Gault . . . ’ which was estimated to lie at about 10-12 feet (3:04-3:65 m.) above
the Gault basement bed. Specimens from the Lamplugh Collection are preserved
in the British Museum (Nat. Hist.) (BMNH L 59863-8) and indicate that these
sediments are of Middle Albian age. There is, therefore, over 27 feet (8-22 m.) of
Middle Albian sediments in this area resting upon a thin development of Shenley
Limestone which in turn rests upon Kimmeridge Clay. It is significant that Neo-
hibolites minimus is apparently plentiful here, in contrast to the Aylesbury area.

In the short distance (14 miles) between Littleworth and Southcott, Buckingham-
shire (SP 90052452), the Woburn (or Leighton) Sands intervenes below a similar
development of Shenley Limestone (Lamplugh 1922 ; 38). Moreover, the Shenley
Limestone lenticle forms the base of a 2 to 3 foot bed of loam with phosphatic nodules
below the Gault. These phosphatic nodule beds form an important feature at the
base of the Gault at Leighton Buzzard, Bedfordshire, where they are essentially of
late tardefurcata and early mammullatum Zone age, with the eodentatus Subzone of
the Middle Albian at the top. The equivalent of the Lower Gault represents the
spathi, intermedius, and niobe Subzones. A description of the Albian sediments in
the Leighton Buzzard area is to be presented elsewhere, and at a later date, an account
of the Middle and Upper Albian sediments of East Anglia will also be given. This
region flanking the London platform and the North Sea area, although of considerable
stratigraphical interest, does not contribute any fundamentally new knowledge to the
ammonite zonal sequence of the Anglo-Paris Basin, the stabilization of which is the
main purpose of this paper.

E. Borehole Evidence

There are now a great number of deep boreholes in southern and eastern England
drilled principally in the search for water, oil and gas, and in Kent, for coal. To this
number can be added a few purely exploratory borings. These have provided a good
picture of the post Tertiary configuration of the Palaeozoic surface, and of the strati-
graphy of the Mesozoic sediments which have buried it (e.g. Kent 1949, Falcon &
Kent 1960). It is now apparent that Albian sediments underlie the Chalk through-
out the area covered by that formation in England. In this work, the Wessex basin,
and the area of the London Basin, Essex, and East Kent only will be considered.
Borings in these areas have yielded important new information, including strong
indications of post Jurassic to basal Upper Albian faulting along part of the Thames
axes, certainly E. of London (Owen, in press).

IN THE ANGLO-PARIS BASIN 69
(i) HAMPSHIRE BASIN

(a) Winchester District

The only borings through the Gault which have yielded information of zonal value
are situated in the Winchester area and were drilled for the Gas Council by the British
Petroleum Co.

Winchester No. r Chilcomb, Hants. (SU 50172830)

The division between the Upper Greensand and Gault is taken at a depth of
310 feet (94.48 m.) and the Gault is 250 feet (76-2 m.) thick. Coring was not con-
tinuous and neither the thickness of Upper Albian sediments nor the exact Subzonal
boundaries in the Middle Albian sediments could be determined. Nonetheless, the
fragments of core show that between 426 feet 6 inches and 438 feet (128-16 m.—
133°50 m.) the essentially dark-grey silty and shelly micaceous clays contain
Inoceramus sulcatus indicating a lower Upper Albian age (cristatum or orbignyi Sub-
zones). There is then a gap of 41 feet (12-49 m.) in cored samples.

At 479 feet (146 m.) depth, the mid-grey shelly silty micaceous clays are of Middle
Albian, intermedius Subzone age, and the sequence contains Anahoplites of the
intermedius group to a depth of 490 feet 2 inches (149-40 m.), a level 69 feet 10 inches
(21-28 m.) above the base of the Gault. The spathi Subzone with crushed Hoblites
(H.) spp. is certainly present at a depth of 501 feet 1 inch (152-73 m.), and the sedi-
ments from this level to a depth of at least 513 feet 3 inches (156-43 m.) consist of
alternating fawn and mid-grey, silty micaceous shelly clay bands. At 521 feet
(158-80 m.) the nacreous shells are found to be completely replaced by pyritic films,
and this pyritic facies is a feature of the sequence down to the base of the Gault at
560 feet (160-68 m.). Crushed Hoflites (H.) spp., are present in the predominantly
fawn-grey silty micaceous clay of the lower part of the Gault to a depth of 526 feet
(160-32 m.) but no ammonites have been found in the remaining 34 feet (10-36 m.).
The basal 5 feet (1-52 m.) of the Gault is very pebbly, becoming a pebbly loam at
559 feet (170-38 m.) depth. The Lower greensand underlies the Gault.

Winchester No. 4 Itchen Valley, Hants. (SU 51133007)

This boring produced only poor chip and core returns during its traverse of the
Gault. Hoplites (H.) sp. with dentatus ribbing was found at 11 feet 6 inches (3-50 m.)
above the base of the Gault, at a depth of 1258 feet 6 inches (383-43 m.). This was
preserved in an identical manner and lithology to that seen at the same level in
Winchester No. r.

Winchester No. 5 Twyford, Hants. (SU 50252712)

Only a few cored samples were recovered from the chipped sequence. At a depth
of 963 feet 8 inches (293-52 m.) only 7 feet 4 inches (2-235 m.) above the base of the
Gault, a crushed heteromorph ammonite with indications of lateral spines was found.
This is almost certainly a Protanisoceras (P.) and indicates the lyelli Subzone.

70 MIDDLE ALBIAN STRATIGRAPHY

(b) Relationship of Winchester to Portsdown, the Weald, and the Isle
of Wight

The correlation of the Gault sequence in the Winchester No. 1 boring with those of
Selborne, Nyewood, and Compton Bay, is shown as far as is possible in text-fig. 30.
The lithological succession shown by the Winchester borings is comparable to that of
the outcrop at Selborne on the western border of the Weald. They both show in the
spathi Subzone sediments a lower pyritic facies overlain by clays in which the shells
are preserved and in which a good benthos is present (Owen 1963a ; 43-44). More-
over, at Bradshott Hall clays of basal Upper Albian age were seen by Osborne-White
(IgIO ; 20) to overlie clays classified with the ‘ interruptus Zone’ in Jukes-Browne’s
sense. The lower clays could be of intermedius Subzone age (Owen 1963a ; 51).

From the map given in text-fig. 18, it is apparent that the borings at Winchester
and Portsdown, and the natural exposure at Culver Cliff in the Isle of Wight all lie
roughly on the same NNW-SSE. line. The total thickness of the Gault at Win-
chester No. I is 250 feet (76-2 m.) compared with 163 feet (49:68 m.) at Portsdown
situated 15 miles to the SSE., and just over 100 feet (30-4 m.) at Culver Cliff. No
information on the degree of ammonite subzonal representation in the chipped
sequence of the British Petroleum Portsdown boring in known (Taitt & Kent 1958),
but, in Hampshire, West Sussex, and in the Isle of Wight, there is a similar change in
facies within the spathti Subzone from pyritic clays below to shelly clays above.
Superficially it seems that as one proceeds SSE. from Winchester the decreasing
thickness of the sediments suggests the shallowing of a basin in this direction. How-
ever, at Winchester the lower part of the Upper Albian is represented by at least
128 feet (39-01 m.) of silty Gault, more than the total thickness of the Gault at
Culver Cliff in which the lower part of the Upper Albian is also represented (p. 43).
The decrease in thickness of Middle Albian sediments from Winchester to Culver
Cliff is, therefore, not particularly well marked. It is also apparent that the detailed
lithological sequence in the spathi Subzone sediments which is recognisable for a
distance of over 35 miles at the outcrop in the south western part of the Weald is
totally different from that of the Isle of Wight (text-fig. 30). The common distri-
bution of the pyritic facies reflects the presence of a common sea environment
which affected different depositional areas, and it is apparent from the pre-Albian
sequence at Portsdown that this area formed a ridge separating the area of the south
western Weald from that of the Isle of Wight.

The Lower Greensand in the south-western area of the Weald thins rapidly towards
Portsdown (Falcon & Kent 1960). Continuing SW. into the Isle of Wight the Lower
Greensand as a whole thickens from Redcliff NE. of Sandown to reach a known
maximum in the southern part of the Island and this increase in thickness can be
correlated with an increase in finer grade sediments. It is apparent from the dis-
tribution of lyell1 and spathi Subzone sediments that the Portsdown ridge affected
Middle Albian sedimentation. By Middle Albian times this submarine feature con-
sisted of an elongated swell (text-fig. 52), and its extent can be determined by the
presence of the ‘Iron Grit’ beneath the Gault and the absence of tardefurcata and
mammillatum Zone phosphatic nodule beds. The Lyelli Subzone sediments are

IN THE ANGLO-PARIS BASIN

COMPTON WINCHESTER SELBORNE NYEWOOD
BAY NO. 1

Ft

intermedius

Subzonal correlation lines
—— ——Lithological correlation lines

Provisional comparison of the Winchester No. 1 boring with the outcrop to the
south and east.

72 MIDDLE ALBIAN STRATIGRAPHY

probably present at Winchester, and may well be present in the unexposed lowest
part of the Gault at Selborne in the western Weald in view of the close similarity
which exists in the detailed lithological sequence between this locality and the Horton
Clay pit, Upper Beeding (text-fig. 14). Sediments of this Subzone are known to
intervene between the ‘ Iron Grit’ and the spathi Subzone sediments along the out-
crop between Nyewood and Storrington, but they are thin and gritty (p. 34). Inthe
southern part of the Isle of Wight, however, sediments of this age are again compar-
atively well developed. The spathi Subzone sequence recognised at Selborne thins
southwards to Nyewood but lithologically it remains the same (Owen 1963a & text-
fig. 30). In the Isle of Wight, however, the sediment sequence is quite different. In
the absence of sections it is not possible at this time to determine whether the Ports-
down swell affected deposition during later Middle Albian Subzones.

It is becoming apparent that the thinning of the Lower Greensand and the Gault
at Compton Bay relates to yet another ‘swell’, probably that indicated by the
Ringwood gravity high, evidence of which is provided by the British Petroleum Co.,
borings at Fordingbridge, Hants (Falcon & Kent 1960 ; 48-49), and Bere Regis,
Dorset (Falcon & Kent 1960 ; 7), in which the Gault was found to rest directly upon
the Kimmeridge and Oxford Clays respectively.

(i) LONDON BASIN, EAST ANGLIA, AND KENT
(a) London Basin, S. Essex & N. Kent

Borings north of the Thames at Canvey Island (Smart, Sabine & Bullerwell e ai.,
1964) Fobbing (Dewey eé al., 1925) Beckton Gasworks No. 4 (Barrow & Wills 1973),
Essex ; Tottenham Court Road (Prestwich 1878, Judd 1884), Willesden No. 1 (Falcon
& Kent 1960 ; 15), London ; and Bushey, Hertfordshire, provide evidence that the
Upper Gault rests directly upon either the Palaeozoic rocks of the Mesozoic floor or
upon a thin development of Jurassic or Lower Greensand sediments. However, in
the Gas Council Cliffe group of borings encompassing the area of the East Tilbury
Marshes, Essex, and across the Thames in the Cliffe and Higham parishes of Kent,
Middle Albian sediments are represented in a sequence closely comparable to that
seen in the Lower Gault of the Maidstone By-Pass (p. 18), and, moreover, they show
no sign of either a land area or submarine cliff only a few miles to the north.

The Lower Gault of the Cliffe group of borings rests upon a thin development of
Lower Greensand which in turn rests upon Oxford Clay preserved in a late Jurassic-
early Cretaceous graben structure, or in the case of the two most southerly borings
upon Devonian sediments as at Canvey Island and Fobbing situated to the north of
the graben. The stratigraphy of these borings, the structure of the area, and its
tectonic history, are discussed more fully elsewhere (Owen in press). In this work it
is concluded that the absence of Lower Gault over much of the area north of the
Thames in London and South Essex, is due to a further movement of the northern
fault of this graben in early Upper Albian times.

Further west in northern Surrey, three borings have yielded information about
Middle Albian sediments. These are located at Addington, Richmond and Egham

IN THE ANGLO-PARIS BASIN 73

(Virginia Water), but the information is very incomplete, and in the case of the last
boring mentioned it is highly suspect. The location of the borings is shown in
text-fig. I.

Addington

The boring at the Croydon Waterworks, Addington Pumping Station on the E.
side of Featherbed Lane (TQ 371628) yielded a core, parts of which are preserved in
the Institute of Geological Sciences. The Upper Gault-Lower Gault junction con-
sists of a phosphatic nodule bed with cristatwm Subzone fossils and was reached at
about 879 feet (267-91 m.) depth. This is underlain by grey shelly clay represented
by fragments of core from between 880 and 882 feet (268-22—268-83 m.) and which
yield Inoceramus concentricus and poorly preserved ammonites which could indicate
either a loricatus or lautus Zone age. No further core fragments have survived from
the remainder of the Lower Gault sequence which has a total thickness of 22 feet
(6-70 m.). This boring is situated 9} miles WNW. of the section at Dunton Green at
the outcrop (p. 26) and indicates that the thin development of Lower Gault there
continues along the WNW. direction.

Richmond

The boring at the old Richmond Vestry Waterworks, Water Lane, Richmond
(TQ 17657470) was first described by Judd and Homersham (in Judd 1884) and
subsequently by Whitaker (1889 ; 214-217). Spath (1926a ; I51, 1930a ; 294)
demonstrated the presence of the Ewhoplites inornatus band at the base of the orbignyz
Subzone (Upper Gault) and that the Lower Gault is also present.

The Gault core was stored in the British Museum (Nat. Hist.) for many years before
being transferred to the Institute of Geological Sciences and is, unfortunately, in a
dirty state. J. sulcatus is still present at a depth of 1116 feet (340-15 m.), and by
I11g feet (341-07 m.) J. concentricus is present. The Upper Gault-Lower Gault
junction occurs, therefore, between these two depths. The base of the Gault was
located at a depth of 1139 feet 6 inches (347-16 m.) and so the Lower Gault is between
20 feet 6 inches (6:25 m.) and 23 feet 6 inches (7:16 m.) thick. No subzonally
diagnostic ammonites are present in the Lower Gault core. The base of the Gault
rests upon I0 feet (3:04 m.) of sediments tentatively classified with the Lower Green-
sand which in turn rests upon Jurassic sediments. In the Griffin Brewery boring at
The Mall, Chiswick, 34 miles to the NE. of the Richmond boring, the Gault rests
directly upon Devonian sediments of the London Platform. The age of the base of
the Gault at Chiswick is, however, unknown.

Egham (Virginia Water)

This boring situated at the Holloway Sanatorium, Egham (TQ 002685), was
described by Dewey (im Dewey e¢ al., 1925 ; 128). He records a band crowded with
I. sulcatus between depths of 1358-1360 feet (415-9I-414:52m.) some 67 feet (20-42 m.)
above the base of the Gault. From the basal nodule beds, Templeman collected
ammonites which led Chatwin to conclude (im Dewey e¢ al., 1925 ; 130, 132) that the
base of the Gault was of Upper Gault age providing another example of overlap.

74 MIDDLE ALBIAN STRATIGRAPHY

Now, Inoceramus sulcatus is characteristic of and restricted to the orbignyi and
cristatum Subzones and yet the ammonites recorded from the nodule bed at 1424
feet (434:03 m.) depth include material of varicosum Subzone age.

A re-examination of the material stated to have come from this nodule bed shows
that it includes ammonites from various Upper Gault horizons. The specimen of
Prohysteroceras (GSM. AT 3787) is indeed correctly identified. It is, however, a form
of the varicosum Subzone preserved in an identical manner to those of the basal
varicosum nodule bed in the Leighton Buzzard area indicating a southerly extension
of that bed. As it was found below the lowest recorded occurrence of I. sulcatus at
1400 feet (426-72 m.) depth, the only possible explanation is that it must have fallen
from the side of the hole together with the other phosphatic fragments during the
collapse of the hole reported by Dewey (in Treacher & Dewey 1925 ; 450).

Material acquired later from Templeman is preserved in the Palaeontology Depart-
ment of the Institute of Geological Sciences and shows that the Lower Gault under-
lain by mammuillatum Zone sediments was in fact traversed by this boring. These
specimens, unfortunately, have no depth measurements recorded against them, but
include Euhoplites cf. opalinus (GSM. AT 4800) indicating the lautus Zone ; Euhoplites
of the meandrinus group (GSM. AT 4799) indicating the upper part of the loricatus
Zone ; and spathit Subzone Hoplites (H.) spp. (GSM. AT 4801-4) preserved in pebbly
gritty greyish phosphate.

Unfortunately, this boring is now stratigraphically suspect, but if one disregards
the so-called phosphatic nodule bed at 1424 feet depth then it is possible to reinterpret
the lower part of the hole. The last record of Inoceramus sulcatus was at 1400 feet
(426-72 m.) depth about 27 feet (8-23 m.) above the base of the Gault. This figure of
27 feet is not an unreasonable one for the Lower Gault when one considers the
geographical position of the boring. The highest record of J. sulcatus is at 1358 feet
(413°9I m.) boring depth which indicates that the combined thickness of the cristatum
and orbignyt Subzone sediments is at least 42 feet (12-80 m.) thick. This is a thick,
but not impossibly thick, sequence.

(b) The area of the Kent Coalfield

Despite the large number of borings and various colliery shafts which penetrated
through the Gault in the search for Coal Measures in Kent, only a small fraction has
yielded information on the stratigraphy of the Gault. Financial costs dictated that
boring through the Mesozoic rocks should be as rapid as possible and the sequence
was often chipped. However, at the following seven localities shown on text-fig. 31,
useful information has come to light and it is apparent that eodentatus and lyelli
Subzones sediments are of widespread occurrence.

Chislet Colliery

In the downcast shaft of the Chislet Colliery situated 3020 yds N. 54° 30’E. of the
North Shaft (TR 232657) an exposure of approximately 12 feet (3-65 m.) was seen
of Lower Gault resting on the basal conglomerate of mammillatwm Zone age which in
turn rests unconformably on Coal Measures (Casey 1961a ; 535). A phosphatised

IN THE ANGLO-PARIS BASIN 75

fragment of Hoplites (Isohoplites) sp. (GSM. Ca 1416) was obtained by Dr. R. Casey
from 1 foot 6 inches (0-457 m.) above the basal conglomerate indicating the eodentatus
Subzone. From 2-4 feet (0-60-1-21 m.) above the conglomerate there occur glau-
conitic gritty darkish grey clays with fossils in which the shells have been replaced
by pyrite. Ammonites from this bed collected by Dr. Casey include Lyelliceras cf.
lyelu (GSM. Ca 1423-4) and Hoplites (H.) spp. including H. (H.) baylet (GSM. Ca
1426), and Beudanticeras cf. albense (GSM. Ca 1431). A specimen of Protanisoceras
(P.) cf. barrense (GSM. Ca 1437) preserved in the same manner was picked up from
the tip. This assemblage indicates the lyelli Subzone. One specimen (GSM. Zn
2472) is a Hoplites (H.) sp. preserved partly phosphatised in mid-grey shelly clay and
is stated to have come from a height of about 12 feet (3:65 m.) above the basal con-
glomerate ; it indicates the spathi Subzone.

| ? Fa 1D miles
fo) 5 10 15km

@ Bobbing Chisigt downcast

Ebb sfleete

Benet

Tilmanst one
°

Ringwould
Guilford
°
St Margarets
Bay
KEY (outcrop)
Bi co:
DOVER
Aycliff ©
FOLKESTONE

pay

Fic. 31. Locality map of borings yielding subzonal information in the Kent coalfield.

Ebbsfleet

Fragments of the core preserved in the Institute of Geological Sciences from this
boring, situated 495 yds S. 5°E. of Ebbsfleet House, Eastry (TR 337619), about 7?
miles E. of the Chislet Colliery, shows that the Lower Gault was entered at between
depths of 977 and 978 feet (297°79-298-I m.) and is about 27 feet thick (8-23 m.)
(Lamplugh, Kitchin & Pringle 1923 ; 178). Lovricatus Zone sediments are definitely
present at 12 feet 6 inches (3:81 m.) above the base of the Gault (GSM., Pl. 3854), and

76 MIDDLE ALBIAN STRATIGRAPHY

at 7 feet 6 inches to 7 feet (2-286 to 2-133 m.) above the base crushed Hoplites (H.) spp.,
occur with the shell, indicating the spathi Subzone.

Lilmanstone

The original manuscript accounts of the succession shown in the Shafts Nos. 1, 2
and 3 of the Tilmanstone Colliery are preserved in the Institute of Geological
Sciences. The collecting was carried out by Burr and Griffiths. Griffiths, then
employed as the Survey fossil collector, knew the stratigraphical value of Inoceramus
sulcatus and I. concentricus and it is possible to state with some confidence from the
Log that the junction between the Upper and Lower Gault occurs at a depth of 870
feet (265-17 m.) in the No. 1 Shaft (TR 288505) and that the Lower Gault is 62 feet
4 inches (18-99 m.) thick. From the account of the No. 2 Shaft (TR 288504), how-
ever, the Lower Gault appears to be only about 52 feet (15:85 m.) thick. The
possible explanation of this difference in thickness is provided by the No. 3 Shaft
(TR 288505) which showed the Gault to be affected by faulting. Whether this
faulting is a posthumous movement of the Tilmanstone Fault which affects the
Palaeozoic rocks and earlier Mesozoic rocks is not clear. Nonetheless, it is readily
apparent that the Lower Gault at the Tilmanstone Colliery is very thick. Very little
material has been preserved from these shafts but one specimen (GSM. Zm 5153) is of
considerable interest. It is an early form of Lyelliceras known to occur in the
eodentatus Subzone in France, but unfortunately no depth has been recorded against
the specimen. It is interesting to note that in No. 2 shaft at a height of 21 feet 8 inches
(6-60 m.) from the basal conglomerate a ‘ 3 inch band of Ammonites interruptus ’ was
recorded. This might indicate a considerable expansion of the dentatus Zone sediments
in this area. Further evidence of this is provided by the material from the Shaft of
the old Guilford Colliery (TR 281469) 24 miles SSW. of Tilmanstone.

Guilford Colliery

The shaft of the Guilford Colliery, situated near the south-western end of Walder-
share Park, Coldred (TR 281469) is now disused. Fossils collected from the Gault
during the sinking of the shaft are preserved in the Institute of Geological Sciences
(presented by the Kent County Education Authority), and in the collection of
Brigadier G. Bomford to whom I am particularly indebted for permitting me to
examine his material. Unfortunately, the depths indicated for the individual
specimens presented by the Kent County Education Authority is suspect. It seems,
however, that at about 851 feet (259-38 m.) depth, the /autus Zone nodule bed at the
top of the Lower Gault was reached. This contains material of daviest Subzone age
as wellas of crvistatum Subzone age, and is, therefore, comparable to the nodule bed
at the outcrop to the west. If this depth of 851 feet (259-38 m.) is correct then the
Lower Gault is 59 feet (17.98 m.) thick, a little thinner than the No. 1 shaft at
Tilmanstone.

The only definite information about the Lower Gault here is provided by Brigadier
Bomford’s collection made from the tip heap of the shaft. This includes material
from a nodule bed of lyelli Subzone age which yielded Lyelliceras lyelli (GB. 5443,
5447, 5446), L. radenaci (Pervinquiere) (GB. 5445), Protanisoceras (P.) buvigniert (GB.

IN THE ANGLO-PARIS BASIN 77

5465), together with species of Hoplites (H.) of both the lyelli and spathi Subzones.
This dentatus Zone sequence in the central area of the Kent Coalfield thins consider-
ably in a south westerly direction towards the outcrop, and there is evidence to
suggest that it thins also eastwards towards the Kent coast.

Ringwould

The Mesozoic rocks traversed by the boring for the National Coal Board made in
1955 and situated 760 yds W. 14°S. of St. Nicholas’s Church, Ringwould (TR
35294812), has been described by Bisson (7m Bisson ef al., 1967 ; 111-114), and fossils
from the Gault were identified by Casey. The Upper-Lower Gault junction consists
of a phosphatic nodule bed and was met at a depth of 839 feet 4 inches (255-83 m.).
The top of the tough phosphatic rock bed of the type seen elsewhere at the base of the
Gault was reached at a depth of 876 feet 11 inches (267-28 m.). The Lower Gault,
therefore, has thinned to 37 feet 3 inches (11-35 m.) and the basal few inches in fact
may well be of mammuillatum Zone age. A re-examination of the fragments of the
core preserved in the Institute of Geological Sciences has yielded the following
additional information.

Crushed Dimorphoplites comparable to D. tethydis Spath non Bayle occur at depths
of 842 feet 3 inches (256-71 m.) and 845 feet I inch (257-58 m.) and indicate either the
top of the meandrinus Subzone, or the lautus Zone. At 860 feet 5 inches (262-25 m.),
a specimen of Hamites tenwicostatus together with a juvenile ?Dimorphoplites niobe
at 862 feet 2 inches (262-79 m.) suggest the presence of the mzobe Subzone. The
intermedius Subzone is certainly represented at 866 feet 8 inches (264-16 m.) and
867 feet g inches (264-49 m.) by crushed examples of Anahoplites intermedius. The
presence of the spatii Subzone is indicated by crushed specimens of Hoplites (H.) at
871 feet 11 inches (265-76 m.) to 872 feet I inch (265-82 m.) in dark grey clay. At
873 feet (265-25 m.) the clays become glauconitic, and at 875 feet 10 inches (266-95 m.)
they become sandy for the remaining 1 foot I inch (0:33 m.) before the basal rock bed
is reached.

The sequence in the lower part of the Middle Albian sediments is demonstrably
thinner than at Tilmanstone where ‘ dark sandy Gault’ commences some 21 feet
(6-4 m.) above the base. An even thinner sequence may be present in the next
boring mentioned here.

St. Margaret's Bay.

The National Coal Board boring at St. Margaret’s Bay situated 1030 yds E. 30°N.
of St. Margaret’s Church, St. Margaret’s at Cliffe (TR 36654533), has been described
by Bisson and Melville (im Bisson et al., 1967 ; 105-110). The bulk of the sequence
was chipped, but at a depth of 800 feet (243-84 m.) cores were taken for 2 feet
(0-61 m.), at 830 feet (252-98 m.) a I foot (0-30 m.) core was taken, and a 3 feet 8 inch
(I-12 m.) core from 840 feet (256-03 m.) and the base of the Gault at 843 feet 8 inches
(25715 m.). The only ammonite recorded was a specimen of a mortoniceratid
ammonite, probably Prohysteroceras, said to have come from a depth of 830 feet
(252-98 m.). What is probably a portion of the same ammonite is stated to have
come from a depth of 830 feet 3 inches (253-05 m.). Casey considers that its position

78 MIDDLE ALBIAN STRATIGRAPHY

only 13 feet 8 inches (4-16 m.) above the base of the Gault is anomalous and should
not be accepted. However, the portion from 830 feet 3 inches (253-05 m.) is not
apparently derived and even if the specimen had come from the core between 800-802
feet (243-84 m.) this still suggests a thinner Lower Gault sequence than at Ringwould.
These records need to be verified in any future boring in this area, but the possibility
of early Upper Albian faulting here of the type seen in the region of the Thames E. of
London should not be excluded (Owen in press). The Gault as a whole on the
E. coast of Kent thins considerably northwards. Inthe Segas Deal Gas Works boring
(TR 374533) it is 86 feet (26-21 m.) thick and in the Thanet Water Board Well,
Margate (TR 365701), it is only 67 feet 6 inches (20°57 m.) thick.

Aycliff

The increased thickness of the Lower Gault seen in the Tilmanstone and Guilford
Collieries is maintained in the Dover area. Lamplugh & Kitchin (1911 ; 8) considered
from an examination of the Dover Colliery shafts that this was due to an expansion
of the higher beds of the Lower Gault, but the exploratory borings in the Dover area
for the Channel Tunnel show in fact that the reverse is the case. There are phosphatic
nodule beds in the cristatum Subzone comparable to those at Folkestone within
Bed VIII. The Gault in one of these borings, Dover No. 1 (Aycliff) (TR 294395), has
been described lithologically by Bisson (in Smart, Bisson & Worssam 1966 ; ror), and
the Lower Gault sequence is shown in text-fig. 32.

The lautus Zone is indicated in Bed 11 by the presence of a Dimorphoplites sp. of
the chloris-biplicatus group, and in another boring by Euhoplites opalinus. It is
considerably attenuated in comparison with Beds V-VII at Folkestone. Bed 10 has
yielded a crushed Dimorphoplites niobe which might indicate either the meandrinus,
subdelaruei, or niobe Subzones. Neither Bed g nor the bulk of Bed 8 yielded any
zonally significant ammonites but crushed Falciferella occurs in the lower 6 inches
(1-828 m.) of Bed 8 which suggest the zutermedius Subzone or possibly the miobe
Subzone. Anahoplites of the intermedius group occur from I foot (0-304 m.) above
the base of Bed 7, and in Bed 6, definitely indicating the presence of the intermedius
Subzone. No subzonally diagnostic ammonites are known from Beds 5 and 4, but
Bed 3 contains phosphatised fragments of H. (H.) persulcatus and H. (H.) of the
paronai group. This is the direct equivalent of Bed I (v) at Folkestone, the dentatus
nodule bed, classified with the spatht Subzone. It is highly probable that Bed 4
above is the equivalent of Bed I (vi) at Folkestone (p. 12).

The particularly interesting feature of the sequence occurs in Bed 2. This bed is
classified with the lyelli Subzone, and contains species of Protanisoceras (P.) at only
12 and 15 inches (0:304—0-381 m.) below Bed 3, and species of Hoplites (H.) occur
throughout. This sequence bears comparison with the lower part of the Gault in the
Guilford, and Chislet Collieries where the lyelli Subzone is also well developed. At
Folkestone, the lyelli Subzone is very condensed and is represented within Bed I (iv).
Whether the eodentatus Subzone is represented within the higher part of Bed 1 is
uncertain in the absence of ammonites but it is highly likely when one considers the
development of the lyelli Subzone here. The lower part of Bed 1 is probably equiv-
alent to the ‘ Sulphur’ Band at Folkestone.

IN THE ANGLO-PARIS BASIN
Bed Utho! Ft Ins M.
olo
No. ad ¢ristatum Subzone eee 2

14

Mid-grey shelly clay with Inoceramus concentricus

flautus

Light fawn shelly grey clay with dark burrows

——

late loricatus

Shelly battleship grey clay with small brown phosphatic
nodules in the basal 1inch

Light fawn tough shelly clay with Falciferella

Medium grey shelly clay with crushed Anahoplites
intermedius

intermedius
@

Pie A alee eae See fe S|
a»
w
:
o

Gritty grey burrowed shelly clay 1 8
6
ae
5 Mid grey shelly clay becoming paler below with
scattered pale buff phosphatic clots I
'
5
'
Gritty burrowed greyclay sparsely shelly with = 3
glauconite in lower half & +
2
| 3 | Blackish phosphatic nodules in gritty glauconitic grey clay
)
Mid grey fine clay shelly and burrowed. In lowest 2ft = 42
lin clay becomes gritty. In lowest 4ins glauconite .
and black phosphatic fragments occur
1
“
>
Is
5
1 Grey silty clay with light burrows and phosphatic nodules 3
pa an

Fic. 32. Lower Gault sequence in the Dover No. 1 (Aycliff) boring, Kent (TR 294395).

79

8o MIDDLE ALBIAN STRATIGRAPHY

These borings in the area of the Kent Coalfield show that the sequence in the Lower
Gault expands considerably eastwards from the outcrop between Folkestone and
Maidstone. In this trough area the eodentatus and lyelli Subzones are well developed.
The whole sequence thins towards the coastal margin of Kent from the Isle of Thanet
to St. Margaret’s Bay northeast of Dover.

F. Selection of sections in France

It is not possible here to describe the Middle Albian stratigraphy of northern and
central France in the same detail as the English sections. The French sections
require just as long and careful study, and must include temporary sections seldom
available on a chance visit and which are thus the prerogative of our colleagues in
France. The purpose of this portion of the work is to make a comparison of the
English sequence with a representative selection of sections of four regions ; (i) the
Boulonnais ; (ii) the outcrop extending from the River Ornain (Meuse) to the River
Armance (Yonne) which includes also parts of the Départements of Meuse, Marne,
Haute-Marne, Aube and Yonne ; (iti) the Pays de Bray ; and (iv) the Pays de Caux
(text-fig. 33).

(i) COMPARISON BETWEEN WISSANT & FOLKESTONE

The Albian deposits of the Boulonnais describe a narrow outcrop at the foot of the
Chalk escarpment extending from the coast at Petit Blanc Nez south-eastwards to
Lottinghen and then roughly westwards towards the coast to disappear beneath the
dunes at Hardelot Plage. The deposits rest upon Aptian sediments at Wissant, but
inland they may rest directly upon Aptian, ‘ Wealden’, Jurassic, and, near Caffiers,
on Palaeozoic rocks. The exposures in the shore and in the cliffs between Wissant
and Petit Blanc Nez are indicated in text-fig. 34.

The Gault of the Wissant area was briefly described by Barrois (1873, 1875, 1878,
but particularly 1879 ; 27-28), Price (1879, 1880 ; 34 etc.) and Jukes-Browne (1900 ;
378-381), but it was not until 1938 (a 98-121) that a good detailed description was
given by J.-P. & P. Destombes. Barrois had considered that the sequences at
Folkestone and Wissant were not comparable in detail, a conclusion with which the
present writer agrees. Destombes & Destombes, however, followed Price in consider-
ing that the Wissant succession is comparable to that of Folkestone, although reduced
in thickness ; a view accepted by Spath (1943 ; 721).

P. & P.-J. Destombes have written an emended account of the Wissant sequence
(1965 ; 257-260), the lithological accuracy of which can be confirmed by the writer’s
own examination of the section. However, their subzonal classification of 1938 and
the implied classification of 1965 requires some revision at certain levels as also does
the account by Marie (1965 ; 280-284, table 1). In April 1967 the writer observed
good clean sections in the Lower Gault both in the cliffs and in the foreshore, and
from the study of these the graphical section (text-fig. 35) has been drawn. The
primary bed numbers employed are those used by Destombes & Destombes (1965 ;
258), and the correlation with Folkestone is shown in text-fig. 36.

‘St Jouln
= rst
cauvitic &

seville
pdeic Héeve

O TOURS

St Martin.
I'Hortien

oForgug les Eaux

© ROUEN

Villars St.
Barthélemy

© EVREUX

O CHATEAUDUN

© VENDOME

60 70 80 90 100km
. 1
10 20 30 40 50 60 miles
© ARRAS
ae O CAMBRAI
SS r O AVESNES
sec Wiis
a 2.

© AMIENS

© BEAUVAIS

PARIS

JOIGNYO

GIEN
fe}

© SANCERRE

© BOURGES

© MEZIERES
© SEDAN

RETHEL
°

VOUZIERS

OREIMS

CHALONS sur
MARNE

Revigny

Pargny s. Saulx
& Maurupt

Le Gaty_
TROYES Dienville

Q

‘Courcelles prés Clérey
La Vendue Mignot:
Montiéhu &

StFlorentin Bois de Perchois

“Mont St. Sulpice
Garey

© CHATILLON sur SEINE
& Beaumont

© AUXERRE

© CLAMECY

Fic. 33. Sketch map of the outcrop of Albian sediments in the Paris Basin showing the positions of the sections mentioned in the text.
The pecked line represents the approximate margin of Albian sediments buried beneath overlapping Upper Cretaceous sediments.

IN THE ANGLO-PARIS BASIN 81

Cap Blanc
Nez

---

CHANNEL

as

Bowen “Chal®

~

-
a ORS

Escalles

Fic. 34. Sketch map of the coast between Wissant and Cap Blanc Nez, Pas de Calais
(modified from J-P. & P Destombes 1963).

F

82 MIDDLE ALBIAN STRATIGRAPHY

At the point marked X on the sketch-map (text-fig. 34) a vertical section normally
buried within the dune sands showed a sequence from the Argiles d’Ostrea Leymerii
(Upper Aptian) up into the lower part of the Lower Gault!. Price (1879, 1880 ; 34)

Bed Lithology g Ins M
Blackish phosphatic nodules, mainly casts of fossils with portions of the shell ~ 2-4
13 preserved, in shelly grey clay. v 0
(v) Burrowed grey clay, the burrows having pale infillings, $
(iv) Very dark grey clay with a few scattered part-phosphatised fossils 5
with the nacreous shell preserved. A line of scattered phosphatic 7
nodules occurs 2 inches from the base.
lii)Part phosphatised fossils in extensively burrowed Clay. = 1
>
= 9
(i1)Grey burrowed $shelly clay. =
4
(i) Dark grey shelly clay. fe)
[11 | Black angular phosphatic nodules in qrey clay. v 2
10 Darkish grey clay with scattered shells, and with a scactered & 1
line of phosphatic nodules buff in colour. a 8
°o
9 | Grey shelly clay. re 4
3
“
2
(il) Sparsely shelly well-bedded dark grey clay, 2 fe)
— 2
x
a3
8 s
1
(i) Glauconitic blocky dark grey clay, with a thin bedded junction p
with 8(li) above,and becoming progress ively more < 8
glauconitic downwards. o
o
7 Large brown grey phosphatic nodules with abraded upper surtaces, some compound, 6
with smaller black phosphatic nodulesand fragments of fossils,in glauconitic loam, 2 0

puzosianus

Fic. 35. Section in Lower Gault in the sea-cliff extending from point X on the sketch map
(Fic. 34) up to the old german block-house 0.4 km SW. of Strouanne, Wissant, Pas de
Calais.

had stated that at Wissant the ‘ Ammonites-mammillaris Zone and the A.-interruptus
zone (Bed I) are mixed together, so much so that it is difficult to divide them; but the
fossils from the former have the greensand matrix. ’. Now, in Bed 7 in the section
mentioned above, a magnificent fauna of puzosianus Subzone age was collected by
the writer preserved in exactly the manner described by Price and associated with a
spathi Subzone fauna. The large puzosianus Subzone nodules had obviously stood

1 It appears to be the first time this century that this section has beenseen. It was described by Gaudry
(1860). Le Hon (1864 ; 14-16), and Barrois (1879). The section examined by Dutertre, in company
with Kirkaldy (1938 ; 121-2), was situated in the cliffs near the farm of Saint Pé. I can confirm the
accuracy of Barrois’ section from a point about 4ft. (1°21 m.) below the top of the Argiles d’Ostrea
Leymerii up to the top of the Sables Vert.

IN THE ANGLO-PARIS BASIN 83

up as a hard-ground on the spathi Subzone sea floor for their upper surfaces are
strongly eroded (text-fig. 35). The currents had scoured out the sandy matrix from
around the nodules, and, subsequently, phosphatic fragments from a later spathi
Subzone period of erosion accompanied by a gritty clay sediment were forced into the
crevices between and even underneath the nodules of puzosianus age. J.-P. & P.
Destombes record ‘ Protohoplites raulinianus ’ from this bed (1938a ; 102).

In the foreshore to the NE. up to Strouanne, these puzostanus Subzone phosphatic
nodules are still present but are scattered and much more rolled : they have not
yielded fossils. In the reef on the foreshore in front of Petit Blanc Nez a few putty-
coloured gritty well-rolled smaller phosphatic nodules of puzostzanus Subzone type
still occur mixed in with the predominantly spathi Subzone debris. It can be seen,
therefore, that the degree of reworking increases north-eastwards from the cliff
section near Wissant. In this respect it is important to note that the Palaeozoic
floor rises sharply off-shore (J.-P. & P. Destombes 1963 ; 53 text-fig. 3).

There is no evidence of the presence of an eodentatus or lyelli Subzone element in
Bed 7, and the spathi element is strongly reminiscent of the fauna in Division A in
the Maidstone By-Pass (p. 38) which indicates that the earliest part of the spathi
Subzone may not be represented either. The equivalent of the Greensand Seam and
the basal spatht Subzone element of the dentatus nodule bed at Folkestone is, there-
fore, absent at Wissant.

It is not easy to make out the complete sequence between Bed 7 and Bed 11 and it
is possible that these clays might be somewhat thicker than has been previously
recorded. Nonetheless, for the purpose of this account, the general sequence given
by P. & J.-P. Destombes will be the one considered. Hoflites (H.) occurs crushed in
the matrix of Bed 7 and in the lower 3 inches of Bed 8 (i), and these sediments are
classified with the spbatht Subzone. The base of the intermedius Subzone has not been
satisfactorily defined but probably commences at the base of 8 (ii). Scattered
crushed examples of Anahoplites intermedius & A. praecox occur throughout the bulk
of 8 (ii) and P. Destombes records Falciferella in the uppermost 8 inches (20 cms)
(1962 ; 196-7). Bed 8 (ii) is, therefore, classified with the intermedius Subzone,
however, as can be seen from the section and the general nature of the fauna it differs
greatly from the upper part of Bed I and Bed II at Folkestone. P. Destombes (1962)
classifies the bed later numbered 9 with the niobe Subzone and this must also include
Bed Io.

Bed 11 has yielded a remanié fauna of ammonites including Mojsisovicsia sub-
delaruet & M. remotum indicating the subdelaruei Subzone, Euhoplites of the mean-
drinus group indicating that Subzone, and also fragments of Ewhoplites lautus and
E. mitidus indicating the nitidus Subzone. The degree of condensation at Wissant is,
therefore, greater than that represented by Bed IV at Folkestone and includes mater-
ial also found in the nitidus Subzone sediments of Bed V.

The tidus Subzone is well developed in Bed 12 (i-iv) and the preservation of the
fossils particularly in 12 (iii) is identical to that of Bed V—VI at Folkestone. It is
possible that 12 (iv) may represent the equivalent of the lower part of Bed VII below
the base of the daviest Subzone but there is no certain evidence for this. However,
it is certain that there is an important break in the sequence between 12 (iv) and (v),

84 MIDDLE ALBIAN STRATIGRAPHY
FOLKESTONE
M Ft E
ZIPS oe SSSeVGii
a
ic
10 © SINVIIIC)
3
:
30
9 ir)
Vil
WISSANT
Pa
2
8 = 0 ae*. © o*e*/13
25 5 2(v)
42 (iv)
ooo 2 (iii)
7
meand, 12 (ii)
RE Fe (FC a LR tle 2 ee
42(i)
20 ote Sek feted 1
6 10
IF 7
2
© 9
r c
SL
Las B(ii)
4
ee 6 oo oe alll (iii)
Wii)
= Gee Wd)
° B(i)
110) “
3 E
c
©
© 1 (vii) 7
2
5
1 <
°
Qa
“

fo) ° eodent

Fic. 36.

yelli

I(ii)

Correlation of Lower Gault sections at Folkestone and Wissant.

IN THE ANGLO-PARIS BASIN 85

and that the daviest Subzone is absent at Wissant. At the base of Bed 12 (v)
Inoceramus concentricus is present but as one works up through the bed it passes
through a subsulcatus stage to achieve the form of J. sulcatus just below the cristatum
nodule bed 13 (cf. P. & J.-P. Destombes 1965 ; 260). Beudanticeras beudanti also
occurs partly crushed with its shell, and it should be borne in mind that d’Orbigny’s
holotype of Dipoloceras bouchardianum is from Wissant and is pyritic with the shell ;
it almost certainly came from 12 (v). Anahoplites daviesi and its close relatives are
absent and in fact the fauna is that which occurs in the lower nodule bed of Bed VIII
at Folkestone minus the late davies: Subzone element (p. 15) absent at Wissant.
Therefore, the lower part of the cvistatwm Subzone is represented at Wissant by these
clays of 12 (v) and this is the only proven section known to the writer where the basal
part of the Upper Albian is represented by an uncondensed sequence. Bed 12 (v)
certainly does not belong to the daviest Subzone as Marie (1965 ; 279) has indicated.

It is worth recording here that Bed 13 at Wissant contains a remanié ammonite
fauna which indicates that it represents Bed VIII (ii & iii) at Folkestone together
with the clays of Bed IX up to the level at which Hysteroceras orbignyi becomes
common. The Euhophites inornatus level, which provides a useful indicator horizon
in the lower part of the Upper Albian, is caught up within Bed 13. This bed re-
presents, therefore, the bulk of the cristatuwm Subzone together with what has been
considered previously to be the lower part of the orbignyi Subzone (see also p. 126).

The original account by Barrois (1875a) of the succession between the Wissant area
of the Boulonnais and the Département of the Meuse has had very little added to it.
This area includes the Ardennes where Barrois demonstrated the major stratigraphic
break which exists between the spathi Subzone and the sediments of Upper Albian age
classified by him with his ‘ Zone of Ammonites inflatus ’ which he included in the
Cenomanian. The sequence in the dentatus Zone is itself incomplete, reflecting the
proximity of the area to the Variscan massifs to theeast. Inthe Département of the
Meuse, the Middle Albian sediments begin to thicken and it is at Revigny-sur-Ornain
that the more detailed account of the sequence in the southern part of the Paris
Basin commences.

(i) THE OUTCROP FROM THE RIVER ORNAIN (MEUSE)
TO THE RIVER ARMANCE (YONNE)

The outcrop of the Albian sediments in the southern part of the Paris Basin is
shown in text-fig. 33. This strip of country is the classic area for the study of the
French Albian ; the name Albian stems from the Roman province of Alba, now the
Département of the Aube. It includes the portions of the Départements of the
Meuse, Marne, Haute Marne, Aube and Yonne, divided into the old regions of the
Argonne (part), then Perthois, and part of the Puisaye. The succession and its
facies changes at the outcrop can be demonstrated by brief descriptions of the follow-
ing seven sections (a) Revigny-sur-Ornain (Meuse) ; (b) Pargny-sur-Saulx (Marne) ;
(c) Les Cétes-Noires prés de Moéslain (Haute-Marne) ; (d) Courcelles prés Clérey
(Aube) ; (e) La Vendue Mignot (Aube) ; (f) St. Florentin area (Yonne).

These sections demonstrate the important development of the dentatus Zone and

86 MIDDLE ALBIAN STRATIGRAPHY

in particular of the eodentatus and lyelli Subzones in this area. The clay facies of the
dentatus Zone passes rapidly in the St. Florentin area to a predominately sandy facies
characteristic of the succession in the Puisaye-Yonne, Niévre, Cher, indicating the
proximity of the Variscan massif of Morvan. In the whole area under consideration
the proved sediments of lovicatus Zone age represent only the intermedius Subzone
and in the Puisaye these are largely remanié. Sediments of lautus Zone age have
not yet been detected.

This area has been studied by many French Cretaceous workers. One of the
earliest papers written was by Michelin (1838) on the sequence at Gaty, prés Geraudot
(Aube). Leymerie (1841, 1842) then described the Gault in the Aube but, unfortun-
ately, d’Orbigny (1841) just antedated Leymerie’s description of Ammonites lyelli
one of the most characteristic fossils. This was followed by a similar description
of the Gault in the Département of the Meuse by Buvignier (1852). Various papers
on individual localities were then published but the next important work before
Barrois was that of Ebray (1863) who attempted to coordinate the sequence in the
various Départements.

Barrois made the first attempt to tie in the apparently different sequences of the
Boulonnais, Ardennes, and the strip of country from the Meuse to the Niévre (1875).
Unfortunately, two very inaccurate attempts were made to correlate the succession
in the Aube with that of Folkestone (Price & Delatour in Price 1879 ; 1880 ; 37-40,
Jukes-Browne 1900 ; 388-390). The result completely obliterated Barrois’s work in
English minds, and eventually led to a great deal of uncertainty as to the strati-
graphical position of the clays containing Ammonites lyelli in relation to the sequence
known in England. This uncertainty was not completely settled even by Spath
(e.g. 1926b ; 1943, 722). It was not until Wright & Wright demonstrated the
occurrence of Lyelliceras in the ‘ benettianus’ Subzone in Surrey (1948), and the
stratigraphical position more definitely indicated by Casey (1961a) that the question
was put beyond doubt in English minds.

The first general account of the Albian in this area of France to appear after
Barrois was a paper by Lemoine (1910). Larcher (1937) subsequently produced a
very interesting paper in which the fauna of the broad lithological units were listed
accurately for the first time. However, it was not until 1965 that a more detailed
picture of the sequence and its facies changes could be obtained. Four very im-
portant papers were presented to the Colloque sur le Crétacé inférieur held in Lyon in
1963. These were published in 1965 and written by :— Larcher, Rat, & Malapris ;
P. & J.-P. Destombes ; Marie ; and Ciry, Rat, Malapris & Nicolas. Of these, the
paper by P. & J.-P. Destombes is of paramount importance. Recently, Lauverjat
(1969) has described the broad lithological sequence and facies changes shown by
deep borings through the Chalk along two lines, parallel to the Albian sediment out-
crop, from the area of Troyes (Aube) south west to the river Loing (Yonne).

(a) Revigny-sur-Ornain (Meuse)

Barrois demonstrated (1878), that in the northern part of the Argonne (part of the
Départements of the Ardennes, Meuse, and Marne) sediments now included in the

IN THE ANGLO-PARIS BASIN

Div. Lithology

22

Ww
Rather fawn blocky marly clay somewhat ferruginous; >
shelly with numerous /noceramus concentricus. The clay mH
passes near the base into soft maristone. =
S
=
15
(ill) Blocky dark fawn grey shelly clay in lower part
becoming more massive bedded upwards-—the fossils
losing the shell until near the top where shells are
preserved again.
10
(il) Phosphatic nodules and part-phosphatised and pyritic fossils
In fawn grey slity clay.
x
~
x
a
wn

(1) Fawn grey slightly silty shelly clay, weathering to a
reddish colour, with scattered pyritic ammonites.

°

Fic. 37. Section in Gault at the claypit of the Société B.H.T.P. on the SE. side of the
Marne-Rhine canal, about 2 km S. of the centre of Revigny-sur-Ornain, Meuse.

87

88 MIDDLE ALBIAN STRATIGRAPHY

Middle Albian appear and thicken southwards. Deposits of lyelli Subzone age,
although no longer exposed, are present in this area. Buvignier (1852 ; 525-6 &
pl. explanation p. 45) lists lyelli Subzone fossils from as far N. as Clermont-en-
Argonne, 31 kms NNE. of Revigny-sur-Ornain. These include Pseudhelicoceras
argonnenis (type locality), Brancoceras versicostatum and Lyelliceras lyelli (although
he states on p. 521 that the only locality at which this species had been found was
at Senard about 17 kms a little E. of N. of Revigny-sur-Ornain). From Revigny-
sur-Ornain itself, and, in the case of the last-named species below, between here and
Mussey (presumably from the excavation for the Marne-Rhine Canal) he records
Nucula bivirgata, Protanisoceras (P.) alternotuberculatum, P. (P.) moreanum (type
locality), P. (P.) nodoneum (type locality) and P. (P.) barrense (type locality).

Approximately 2 kms S. of the centre of Revigny-sur-Ornain, and to the SE. of the
N 395, the Société B.H.T.P. have a large brick and tile works with an extensive
clay pit on the SE. side of the Marne-Rhine canal. The pit is worked in two stages by
multi-bucket excavators and a sketch section is given in text-fig. 37. Deposition of
sediment here was apparently fairly constant and there is an almost perfect transition
from the lithology seen at the base to that seen at the top. The only sign of slight
condensation occurs approximately 30 feet (9:14 m.) above the base of the section
where part-phosphatised and pyritic fossils are more common.

The fossils are essentially crushed flat, and at the base of the sequence the ammon-
ites consist of Hoplites (H.) spp. including H. (H.) dentatus and Metahamites sabliert
(d’Orbigny) indicating the spatht Subzone. A higher spatht Subzone fauna ranges
up into the top few feet of Division I, and the band of phosphatised and pyritised
fossils about 30 feet (9:14 m.) from the base has yielded ammonites including H. (H.)
aff. dorsetensis Spath, H. (H.) pretethydis Spath, and H. (H.) canavariformis com-
parable to those found in the upper part of the spathi Subzone in the Weald.

Anahoplites praecox and A. intermedius appear at the base of Division 2 and range
up through the remainder of the measured sequence. In Dr. P. Destombes’ collection
there is a single example of the sulcate form of A. praecox which indicates the lower
part of the intermedius Subzone and was probably derived from the top few feet of
Division I.

The spathi Subzone is represented in this section, therefore, by about 50 feet
(15:24 m.) of sediments, and the base of the Subzone has not yet been reached. The
top of the intermedius Subzone has not been reached either, and the exposed portion
of sediments belonging to this Subzone is about 15 feet (4:57 m.) thick.

(b) Pargny-sur-Saulx (Marne)

In the area of Pargny-sur-Saulx and Maurupt, situated approximately 12 kms SW.
of the section at Revigny-sur-Ornain, there is an important centre of brick and tile
production. Houdard (1940 ; 625-636) has recorded the distribution of the fauna
collected from the Gault (Argiles Tégulines) in this area but there are no details of the
sections. From his lists it is apparent that the lyelli and spathi Subzones are definitely
present, and probably the intermedius Subzones as at Revigny. His record of
‘ Acanthoceras’ camatteanum and ‘ Parahoplites’ steinmanni from Pargny suggested

IN THE ANGLO-PARIS BASIN 89

that the eodentatus Subzone might also be present. The section between Pargny-sur-
Saulx and Maurupt described below provides new information on the basal part of
the Middle Albian in this area.

The extensive clay pit belonging to the brick and tile works of the Hugenot Freres,
situated a few hundred yards E. of the D6o1, 1km NNW. of Maurupt, has been deepened
and shows the lithological sequence given in text-fig. 38. Bed 1 did not yield
ammonites to the writer but bivalves are common. It is possible, although by no
means certain, that it is of uppermost mammuillatum Zone age. Bed 2 has yielded
phosphatised or pyritised Hoplites (Isohoplites) eodentatus, H. (I.) sp., Beudanticeras
albense, B. sanctaecrucis, Otohoplites sp., Lyelliceras camatteanum (d’Orbigny),
Brancoceras sp., indicating the eodentatus Subzone, and its top is an erosion surface.
The sediments of Bed 3 contain crushed fossils including ammonites, some quite large,
such as Hoplites (H.) spp., Lyelliceras of lyelli Subzone appearance, and Douvilleiceras
sp., and can be classified with the basal part of the /yelli Subzone.

Bed Lithology Ins M
pi
Fawn-grey slightly silty clay, shelly, with iron-stained = 6
partings. Some fossils are partly phosphatised. =
1
(11) Black phosphatic nodules scattered in fawn grey silty clay. ~ 344
”
2
(i) Fawn-grey shelly clay, silty with iron stained partings. 3 =
Part-phosphatised and part-pyritic ammonites occur & ae 4 5
scattered throughout. aS
“
——) 2
[——
/ Lew eel
|
ee | |
Silty fawn-grey shelly clay with some pyritic fossils | 5 Gi
| “.
| = ete o4

Fic. 38. Section in basal Gault exposed in the claypit of the Hugenot Freres, situated a
few hundred yards E. of the D 61 road, 1 km NNW. of Maurupt, and 1-4 km S. of Pargny-
sur-Saulx, Marne.

(c) Les Cétes Noires (Haute Marne)

Approximately 15 kms SSE. of the section described above is the natural river cliff
of Les Cotes Noires situated on the W. bank of the River Marne 1 km to the W. of
Moéslain, near St. Dizier. This magnificent natural section can only be safely
worked in reasonably dry weather and is approached by way of the summit of the

go MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ft Ins M

9 Yellow grey clay. = 15 850
.
a
”
8 More calcareous clay with pyrito phosphatic ammonites. 1 7
4
cl
ap Blue black micaceous clay with gypsum crystals and
scattered phosphatic nodules. - 31 ie)
<
=
10
|
|
in
|
6 Grey micaceous clay. | 15 6
5
|
| s.
|
”
2
~~
Black micaceous clay. a 9 3
©
v»
°
“
le}

Fic. 39. Section in Middle Albian sediments (largely after P. & J-P. Destombes 1965)
at Les Cétes Noires, 1 km W. of Moeslain on W. bank of the river Marne (Haute Marne).

IN THE ANGLO-PARIS BASIN gi

cliff from the D196 between St. Aubin and Laneuville-au-Pont. The section has
recently been described by P. & J.-P. Destombes and is quoted in text-fig. 39. It
forms one of the original localities cited by d’Orbigny when defining the stage (1842 ;
404) and is doubly important because it shows a complete sequence from the mam-
millatum Zone to the spathi Subzone of the dentatus Zone.

The outline classification of the sequence is as follows. The eodentatus Subzone is
definitely represented within Bed 5 and may include 6 and the basal part of 7. The
remainder of Bed 7 together with 8 contains a lyelli Subzone fauna, Bed 8 lithologic-
ally represents the NE. extension of a marker horizon recognisable at the top of the
lyeli Subzone in the Gault of the Aube and in the St. Florentin area of the Yonne
(text-fig. 43). Bed 9 contains a spathi Subzone fauna.

(d) Courcelles prés Clérey (Aube)

This clay pit, No. 3 of the Tuileries de St. Parres les Vaudes, is situated to the E. of
the river Seine on the eastern side of the D 49 about 2:5 kms SE. of Clérey. It now
forms the most important section available in the Middle Albian sediments in the
Aube, and is typical of the sequence formerly exposed at such famous localities as
Dienville on the River Aube, and Gaty-prés Géraudot in the Forét d’Orient (Larcher,
Rat & Malapris 1965 ; 246). It is very close to the old section at Courcelles figured
by Leymerie (1846 ; pl. 3, fig. 4). The section is given in text-fig. 40, and its correla-
tion with sections to the NE. and SW. in text-fig. 43. It has been described briefly
by P. & J.-P. Destombes (1965 ; 262), and is particularly important in that it permits a
direct comparison to be made with the sequence in the Horton Clay pit, Small Dole,
Sussex (p. 35).

Bed I to a metre above the base of Bed 4, are classified with the /yelli Subzone, and
the following list of ammonites is certainly not exhaustive and represents material
collected strictly im situ. Bed 1 is not very fossiliferous but has yielded Hoplites (H.)
bullatus, H. (H.) dentatus group, Beudanticeras albense, Lyelliceras pseudolyelli
(Parona & Bonarelli), L. aff. gevreyi. Bed 2 (i) Beudanticeras santaecrucis, Brancoceras
sp., and on a bedding plane 1 foot (0-3 m.) from the base, Desmoceras latidorsatum is
not uncommon and this horizon has also yielded a single example of Hypophylloceras.
Bed 2 (ii) Brancoceras sp., Pseudhelicoceras argonnense. Bed 2 (iii) Lyelliceras lyelli,
Brancoceras sp., Eubrancoceras aegoceratoides (Steinmann), ‘ Oxytropidoceras’ evanst.
Bed 2 (iv) Desmoceras latidorsatum, Beudanticeras laevigatum, B. sanctaecrucis, B.
albense, Hoplites (H.) sp. dentatus group, Lyelliceras gevreyi, Brancoceras spp.,
Protanisoceras (P.) alternotuberculatum, P. (P.) barrense P. (P.) nodoneum. Bed 2 (v)
Beudanticeras laevigatum, Hoplites (H.) baylei, H. (H.) sp. dentatus group, P. (P.)
alternotuberculatum, Pseudhelicoceras argonnense. Bed 3 Douvilleiceras clementinum,
Hoplites (H.) dentatus, H. (H.) baylei, H.(H.) spp. Bed 4 basal x metre is character-
ised by Hoplites (H.) spp. but P. & J.-P. Destombes record Dowvilleiceras up to this
height, and it is here included in the lyelli Subzone.

In comparison with the Horton Clay pit, Small Dole, the fauna listed above shows
the following important features. Bed zr must be close to the underlying eodentatus
Subzone for Lyelliceras pseudolyelli is directly transitional from Lyelliceras camatteanum

MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ft Ins M.
in 16
=
ry

6 Weathered grey silty clay somewhat marly near the base. = 6 °
&

5 15
=
Vv oOo On tate se}— 2
5 Blackish phosphatic nodules in weathered grey silty clay. K
144
13
12
11
10
4 Dark grey slightly silty clay with crushed shells and part pyritised = 34 Te)
fossils. Brownish phosphatic nodules are scattered throughout. At 8 =
feet trom the base there is a seam of ‘solid’ pyritic ammonites, a
Approximately 2inches above the base thereis a similar bedding ? 2)
plane with well preserved fossils.
8
| 7.
|
= =| = =
6
| V 2
|
— 4
3 Marly silty light grey clay with large concretions of silty shelly marlistone. 3
(oe ee
| (v) Darkish fawn-grey silty clay, shelly with scattered ‘solid’ pyritic 4
fossils
3
(iv) Mottled fawn-grey clay, silty with more numerous ‘solid’ pyritic ammonites. 4
2 (ill) Darkish fawn-grey shelly silty clay with scattered pyritic fossils. A =
distinct line of brown cavernous phosphatic nodules occurs 4 inches above x 8
the base, below which the clays become much siltier. ~ E
x
(ii) Mottled fawn-grey shelly clay silty with ‘solid’ fossils, 8
—
(1) Darkish silty fawn-grey shelly clay with a bedding plane of scattered 1
pyritic fossils at the middle, ©
1 Oark grey silty clay with scattered line of phosphatic nodules 9inches from (e}
the top.
°

Fic. 40. Gault section at the No. 3 claypit of the Tuileries de St. Parres les Vaudes at
Courcelles prés Clérey on E. side of the river Seine, and to the E. of the D 49 road 2°5 km
SE. of Clérey, Aube,

IN THE ANGLO-PARIS BASIN 93

of the eodentatus Subzone to Lyelliceras lyelli of the typical development of the lyelli
Subzone. Beds 2 and 3 contain a closely comparable fauna to that of the English
lyella Subzone except that in England the Tethyan element represented by Desmoceras
and Hypophylloceras is absent. Douvilleiceras clementinum is common in Bed 3 at
Courcelles but this genus is very uncommon in the lyelli Subzone in England. The
species of Brancoceras are somewhat different to those found in England and they
frequently show a tendency to a zig-zag arrangement of the ribs as they sweep across
the venter. The occurrence of Eubrancoceras aegoceratoides in Bed 2 (iii) is very
important for long-range correlation (p. 135).

The beautifully preserved fauna of Bed 3 includes also bivalves, gastropods, and
corals, and it is this horizon that has yielded many of the fine specimens of Hoplites
(H.) spp., and Dowvilleiceras clementinum from such localities as Dienville, Gaty, or
just Aube, found in museum collections.

The remainder of Bed 4 contains a typical spatht Subzone fauna consisting essenti-
ally of species of Hoplites (H.) together with Metahamites sablieri and Inoceramus
concentricus. Ata height of 8 feet (2-438 m.) from the base of Bed 4 there is a small
thickness of clay with scattered pyritised ammonites and this has yielded to the
author a single example of Mojsisovicsia delaruet compressa (Spath).

Bed 5 probably represents much of the higher part of the spathi Subzone sediments
seen at Revigny-sur-Ornain (text-fig. 37), for the species of Hoplites at the top of Bed
4 are not particularly high forms. Bed 6 above contains Anahoplites sp. indicating
the extreme base of the iutermedius Subzone.

Larcher, Rat & Malapris (1965 ; 246) considered that the section at Villemoyenne
(No. 1 of the Tuileries of St. Parres-lés-Vaudes) showed a mammillatum Zone sequence.
However, P. & J.-P. Destombes (1965 ; 263) and Marie (1965 ; table 1) indicated that
there was overlap between the sequence exposed at Villemoyenne and that of
Courcelles. The pit is extensive and the dip appears to be negligible. It is situated
less than 2 kms SE. of the section at Courcelles and 0-5 km along the road from
Villemoyenne to Le Ht. Villeneuve. An examination of the sequence has convinced
the writer that it is wholly of high mammullatum Zone age, a conclusion which Dr.
P. Destombes has also arrived at (personal communication).

(ec) La Vendue Mignot (Aube)

The Tuilerie Le Clerc, situated about 200 yds W. of the D1 and a few hundred
yards S. of the D108 roads at La Vendue Mignot, has been nominated as the type
section of the Subzone of Lyelliceras lyelli & Hoplites benettianus by P. & J.-P.
Destombes (1965 ; 262, 266). The section is very shallow, like many of the smaller
terriers in this region of France, but with the combination of a northerly 3° dip and
the slope of the ground surface, approximately 16 feet of weathered clays are exposed
(text-fig. 41). The section is situated about 9 kms WSW. of that of Courcelles prés
Clérey, and the sequence although well weathered in its upper part is apparently the
same.

Bed I was seen to a depth of 7 feet (2-133 m.) but yielded no ammonites to the writer.
At 2 inches (0-050 m.) above the base of 2 (i) a single example of Desmoceras latidor-

94 MIDDLE ALBIAN STRATIGRAPHY

satum (Michelin) was found im situ enabling a direct correlation to be made with
Courcelles. Bed 2 (ii) at La Vendue Mignot corresponds to 2 (ii) at Courcelles, but
yields a somewhat higher percentage of part-phosphatised fossils. The remainder of
Bed 2 is too deeply weathered to permit the recognition of the remaining subdivisions
seen at Courcelles, although its fauna grosso modo is the same as that listed above. In
the soil at about the middle of the northern face of the pit, there are pieces of well
weathered sandy marlstone yielding the fauna of Bed 3 at Courcelles.

Although this is the nominated type section of the /yellc Subzone proposed by P. &
J.-P. Destombes it shows neither the relationship with the eodentatus Subzone below
or the spatht Subzone above. Neither does it show a complete sequence in the lyelli
Subzone itself.

Bed Lithology

efi: Ins M.

Fragments of gritty maristone in subsoil.
Very weathed buffish silty clay with some limonitic fossils 4
in the lower part. 6 4

= 3
(ii) Slightly lighter clay with part-phosphatised part-limonitic fossils. 8

~
(i) Weathered buffish silty clay with a few limonitic fossils. 2 fe)

2

Weathered gritty glauconitic buffish clay. 7 le)

(0)

Fic. 41. Section in Gault at the Tuilerie Le Clerc, c. 200 yds W. of the D 1 road and a few
hundred yards S. of the D 108 road, at La Vendue Mignot, on the N. side of the Foret
d’Aumont, Aube.

(f) St. Florentin area (Yonne)

The early work of Ebray (1863) and Hébert (1863) did not present a true picture
of the Albian succession in the area of St. Florentin, and Lambert (1894, I913) was
the first worker to give a more correct sequence. Lambert’s sequence was the one

IN THE ANGLO-PARIS BASIN 95

quoted subsequently by Lemoine (1910). Houdard (1933) confirmed Lambert’s
observations and presented accurate and important new information. More recently
P. & J.-P. Destombes (1965 ; 264-265) have reinterpreted the sequence indicating a
facies change certainly within the spathi Subzone between St. Florentin and Montléhu
a distance of barely 1 km. In this area the clay facies of the Perthois (the strip of
country flanking the Chalk and including portions of the Départements of the Haute
Marne, Aube, and Yonne) gives place to the predominantly sandy facies of the
Puisaye (the similar strip of country stretching from the River Armance to the Loire).

No sediments of eodentatus Subzone age have been proved in this area. Although
known to occur, lyelli Subzone sediments are no longer exposed, but from sections
which existed formerly at St. Florentin, and to the SW. near Mont St. Sulpice,
Seignalay and Beaumont, it is readily apparent from the old collections that the
equivalent of Bed 3 at Courcelles is present. The fauna is the same but it is pre-
served in a much grittier and pebbly matrix. Two sections in sediments of spathi
Subzone age exist in this area today.

Montléhu

The Tuilerie Montléhu is situated immediately S. of the N77 at the village of that
name. It exposes about 18 feet (5-48 m.) of weathered grey clays with crushed
Hoplites (H.) spp., and Inoceramus concentricus, and is classified without doubt with
the spatht Subzone.

As one proceeds SW. the topography changes quickly, and St. Florentin is built
upon the high ground formed by the essentially sandy deposits of both Middle and
lower-Upper Albian age.

Sabliére Binot

This disused sandpit in the Sables de Frecambault at the SE. end of the town of
St. Florentin is situated on the same quarried escarpment as that described by
Houdard (1933 ; 47). The pit, which is now being filled in, shows sediments of Middle
& Upper Albian age (text-fig. 42). It has been mentioned by P. & J.-P. Destombes
(1965 ; 264), and Marie (1965 ; table opp. p. 286) who includes additional information
on the other sections in the area.

Bed 1 has not yielded fossils but it lies above the equivalent of Bed 3 at Courcelles
(Lyell Subzone), known to be present in this area. It is almost certainly of spathi
Subzone age and is considered to be the equivalent of the clays exposed at Montléhu.
Bed 2 shows the incoming of clay sediment and it seems to the writer that this part
of the sequence is more likely to have been deposited at the same time as the clays at
Montléhu. However, the only fossils found are phosphatised bivalves. Dr. P.
Destombes has informed me that large Hoplites (H.) were obtained from Bed 3 during
the quarrying operation.

Bed 4, the Bed VII Graviers 4 Opis glareosa of Lambert, is of considerable interest.
It can be divided into two very irregular subdivisions. A lower dark grey sub-
division with Hoplites (H.) spp., derived from the spathi Subzone, and an upper

96 MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ft Ins M,
a aa a . a - |
|
7 Soft maim with buff phosphates and pyritic concretions. | 3 ° 16
Blocky sandy yellowish maim. Z 1 fe)
SSS
1%
he 15
_<
le
Wi
jo
a
5
| 14
= Alternating broad seams of gritty clay and sandy beds Incompletely | 10 6
seen. |
|
| 13
Ig
Is 12
ee ieee ee —_— — —- |:
Grey gritty clay seen clear for: re
4 Coarse pebbly sandy loam with admixed ape) clay. Phosphatic nodules occur 5 o
LE in roughly two concentrations (see text). = 7
3 Coarse hard phosphatised pebbly Stone=pand: dark-grey in top 4 inches with
L phosphatic nodules. jel Ser : we 8-16 4,
2 Coarse pebbly loamy sand with streaks of grey clay. 1 6
104
(v) Coarse yellowish pebbly sand. 6 fo)
9
(iv) Lenticles of sandstone. 1 °
=a
(lil) Coarse yellow pebbly sand. 5 8
= 7
=
x
a
n
a Lenticles of sandstone. 1 te)
1 6
|
| S
4
(i) Coarse yellow pebbly sand. 20 fo}
3
2
1
= = = °

Fic. 42. Section in the Sables de Frecambault at the Carriere Binot, on the escarpment
at the SW. end of St. Florentin, a few hundred yards N. of the lane leading to Crécy,
Yonne.

IN THE ANGLO-PARIS BASIN 97

lighter coloured subdivision which contains some material scoured out from the
spathi Subzone sediments, and a predominant element derived from sediments of
intermedius Subzone age. Indigenous Anahoplites intermedius and Inoceramus
concentricus occur in the upper subdivision indicating that it is undoubtedly of
intermedius Subzone age. The derived intermedius Subzone material includes
species of Anahoplites which occur at the extreme top of the sfathi Subzone and basal
part of the intermedius Subzone in England.

Bed 5 is still pebbly in its lower 1 foot (0-30 m.) but it has not yielded fossils to the
writer. Marie indicates that at about the middle of Bed 5 at the base of an argil-
laceous member the Upper Albian commences.

(g) Summary

The sections described briefly above provide a picture, albeit very imperfect, of
Middle Albian sedimentation across the basin of deposition extending from the
Kimmerian modified Palaeozoic massifs of the Ardennes on the NE. side to those of
Morvan to the SW. (text-fig. 43).

The only information available about the sediments of eodentatus Subzone age come
from the sections near Maurupt and Les Cotes Noires, inthe NE. It is acurious fact
that as yet it has not been detected in the Aube or Yonne. This must be due to a
lack of exposures for the sequence in the mammuillatum Zone in the area of the Bois
de Perchois (Aube) is very thick and the Middle Albian sediments here are also
apparently of considerable thickness. In England deposits of eodentatus Subzone
age are in general very condensed and ammonites are not common except at a very
few localities. Species of Hoplites (Isohoplites) form the majority of the ammonite
fauna, and its usual associate in France Lyelliceras of the camatteanum group, is
exceedingly rare in England.

Deposits of lyelli Subzones age already proven at Clermont-en-Argonne are also
apparently thicker at Les Cétes Noires than in the Aube, but the lithological sequence
is comparable from Les Cétes Noires to the St. Florentin area (Yonne).

At Revigny-sur-Ornain it is obvious that the spathi Subzone is represented by
thick sediments especially the upper part. Unfortunately, the top of the Subzone
has not been exposed at Les Cétes Noires, but at Courcelles the upper part of the
Subzone is represented by a single nodule bed. No information is available about the
sequence until the St. Florentin area is reached where, between Montléhu and St.
Florentin itself, there is a change from the argillaceous facies of the Aube to the sandy
facies of the Puisaye.

The intermedius Subzone is now known from five localities. It is quite thick at
Revigny-sur-Ornain and the top of the Subzone was not determined in the sequence.
| Intermedius Subzone ammonites were determined by Breistroffer at Montierender

(Haute Marne) and Le Plessis (Aube) (P. & J.-P. Destombes 1965 ; 262). Only the
basal part of the Subzone has as yet been determined at Courcelles. At St. Florentin
and at other localities in the Puisaye, the Subzone is represented within condensed
deposits. No other Middle Albian Subzones are as yet known in this area.

Very little information is available about the stratigraphy of the Gault between

i@ c

i

98 MIDDLE ALBIAN STRATIGRAPHY
ST FLORENTIN MONTLEHU LA VENDUE COURCELLES Les COTES MAURUPT
M. Ft. MIGNOT NOIRES
30-7109
a ? ?
2 ? ? ? ? ae
os nw
mt 4 INTERMEDIUS
2'
? ? ? 2
80
(
SK “Po eny
i
we
N
70 .
»
N
\
eaAEHy SPATHI we
‘\
(lower) ty \
Oy Ne
e
60 )
)
15.50 ? 2 ? ? 2
faa oes oo el
(fea == SSH a at |e, age)
LYELLI
40
7—
10 7
30
S
2
EODENTATUS ™
aw
=
? ?
20 a7 7 ? ? 7
5
|——-
10
6
oto

subzonal correlation lines

{}thological correlation lines

REVIGNY
sur
ORNAIN

1c)

Fic. 43. Correlation of Middle Albian sections from the Meuse to the Yonne.

4

IN THE ANGLO-PARIS BASIN 99

this area and the Pays de Bray. However, a comparison between the succession
shown by the La Chapelle boring at St. Denis, Paris (Jukes-Browne 1900 ; 397) and
that of St. Florentin (Yonne) and Villers St. Barthélemy in the Pays de Bray below
shows that the predominantly sandy beds in the Yonne, fringing the massif of Morvan,
give way to clays under Paris, but the sequence is thinner. The sequence thickens
again northwards from Paris, and in the Pays de Bray the Middle Albian is represented
by clays, and the Upper Albian by an Upper Greensand facies. The clay facies
extends down to include at least the top of the mammillatum Zone towards the NW.
end of the Bray.

(iii) PAYS DE BRAY

The Pays de Bray both geologically and scenically resembles the Weald (text-fig.
33). The NE. side of the Bray dips very steeply beneath the Chalk and there is no
information on the sequence on this side. However, at the NW. end and along the
whole of the south western side the dip is much more gentle and brick-pits in the
Gault have been opened at a number of places. Usable information has been ob-
tained from only four of these : Briqueterie Ledoict, St. Martin, at the NW. end of the
Bray (P. Destombes 1970 and im Pomerol & Feugueur 1968) ; a section near Forges-
les-Eaux (P. Destombes 1958) ; and two sections in the area of Villers St. Barthélemy
(J.-P. & P. Destombes 1938b). The position of these sections is indicated on
text-fig. 33.

The most detailed information on the Middle Albian sequence in this region yet
published is contained in the two papers already cited by J.-P. & P. Destombes
(1938b) and P. Destombes (1958). They demonstrate the presence of the eodentatus,
lyel, spathi and intermedius Subzones all in a clay facies. The niobe Subzone is
indicated as being represented by sandy deposits (P. Destombes 1958) but no definite
evidence has been published to support this. This is followed by a break in the
sequence involving the remainder of the Middle Albian.

(a) Villers St. Barthélemy

Two sections were described by J.-P., & P. Destombes in this area (1938b ; 122,
123). Their section 1 is no longer exposed but showed deposits of both the spathi and
intermedius Subzones which together are over 30 feet thick (9:15 m.). Section 2 is
now well exposed and shows the sequence given in text-fig. 44.

Bed 1 (ii) contains Otohoplites spp. including O. destombesi, Beudanticeras spp. and
_ Douvilleiceras spp., indicating the uppermost part of the mammullatum Zone. No
_ fossils were seen in the lower part of 1 (iii) and it is not possible to say whether it is of

eodentatus or basal lyelli age: the top of I (iii), however, contains crushed Hoplites

(H.) spp. Bed 2 contains a good bivalve fauna but only a few crushed specimens of

Beudanticeras cf. laevigatum, Protanisoceras (P.) sp. cf. barrense, and in Dr. P.

Destombes’ collection, a few examples of Lyelliceras lyelli. This bed definitely can

be classified with the lyelli Subzone, as also can Bed 3 on the occurrence of a few

crushed Beudanticeras at about the middle of the sequence. The basal part of Bed 4

|

100 MIDDLE ALBIAN STRATIGRAPHY

however, contains large crushed Hoplites (H.) including H. (H.) cf. dentatus and H. (H.
cf. maritimus sp. nov. indicating the lower part of the spatht Subzone. How much
of this sequence overlaps that of section I, if at all, cannot be determined. Un-
fortunately, during 1967 this section was rapidly expanded, and the higher part of
the sequence seen in a rise has now been quarried away.

Bed Lithology Ft Ins M

Weathered grey silty clay with large partly phosphatised
ammonites. Shells are replaced by pyrite.

4
Dark grey somewhat silty glauconitic clay with occasional . 8 0 3
phosphatic nodules.
2
Fawnish grey clay burrowed by dark grey clay witha few 4 °
crushed fossils with pyrite replaced shells
oN
|
I
|
|
I
|
W

spathi

lyelli

(il) Sandy glauconitic green grey clay with sandy pockets
particularly in the lower part.

t PREOe = Z 0

Fic. 44. Section in Gault in a claypit 0'8 km N. of the village of Villers St. Barthélemy,
c. 150 yds E. of the D2 road, Pays de Bray.

(b) Forges-Les-Eaux, & St. Martin

Further north in the Pays de Bray, the eodentatus and lyell1 Subzones sediments
expand considerably and consist of shelly clays. The section in the Briqueterie
Ledoict near St. Martin l’Hortien situated at the NW. end of the Pays de Bray has
been described by P. Destombes, who introduced the writer to it. It shows a good
development of clays of eodentatus Subzone age overlying high mammullatum zone
clays (see P. Destombes 1970 & in Pomerol & Feugueur 1968 ; 129-130, where the
locality is given as Bully).

The lyelli Subzone is known from a section west of Forges-Les-Eaux where Fortin
collected phosphatic and pyritic Lyelliceras with the shell, now in the Muséum
d’Histoire Naturelle, Rouen, which were recorded by P. Destombes (1958 ; 309).

IN THE ANGLO-PARIS BASIN 101

However, no detailed information on the sedimentary sequence in this Subzone in
the northern area of the Bray has as yet been recorded.

(iv) COMPARISON BETWEEN THE PAYS DE CAUX AND
THE ISLE OF WIGHT

The expansion of the sequence towards the NW. end of the Pays de Bray apparently
increases further to the NW., for the deep boring at Puys, near Dieppe, showed a very
considerable thickness of clay (Jukes-Browne 1900 ; 398). However, an unknown
thickness of this clay must be of Upper Albian age, representing a facies change from
the Upper Greensand sequence seen in the Pays de Bray. Jukes-Browne is probably
wrong (1900 ; 398) in classifying only the lowest 6-7 feet (2 m.) of sandy black clay of
the core with the Lower Gault, but in the absence of palaeontological evidence, the
boring cannot be interpreted. In the Pays de Caux a somewhat different facies is
seen reflecting the proximity of a marginal area of Middle Albian deposition.

The Albian sediments of the Pays de Caux (Seine Maritime) are well exposed in the
cliffs between St. Jouin and Le Havre. The Upper Albian Gaize rises from sea-level
at the base of the Chalk cliffs to the N. of St. Jouin followed quickly by the Gault,
and the top of the underlying Poudingue ferrugineux. Below these pebbly beds are
sands of Upper Aptian age. In the area between Octeville and St. Adresse these
sands are seen to rest upon Kimeridgian sediments, and in this area the whole
sequence of Lower Cretaceous sediments can be seen sandwiched between the Chalk
and the Kimmeridge Clay. However, along the entire coast from St. Jouin to Cap
de la Héve a large number of rock falls tend to obscure the lower part of the Albian
sequence in particular, nonetheless, it is possible to make out the succession at many
points.

Lennier (1867) first described the succession and provided the foundation upon
which subsequent stratigraphic work has been based. His is still the only published
section of the sequence at Cauville (1867 ; plate 4). The early history of research was
summarised by Jukes-Browne & Hill (1896) who made the first major attempt to
correlate Upper Albian and Cenomanian sediments in the Pays de Caux with those of
southern England. Hill also provided some information on earlier Albian sediments
and gave the first detailed accounts of the Albian sequence seen between Octeville
and Ste. Adresse. Jukes-Browne (1900 ; 395-401) reviewed the Albian sediments in
this area, presenting a useful picture but without any real detail. Subsequent
stratigraphic work has tended to concentrate on the sequence at Cap de la Héve, as
for example the recent important studies by P. Destombes (1958), Cayeux (1960),
and Rioult (1962). Destombes (1958 ; 306-308) sets out to describe the Albian
sediments between Le Havre and St. Jouin but, and this is important, he bases his
account of the stratigraphy on the sequence seen at Cap de la Héve and Octeville.
His classification of these sediments gives a good picture of the zonal sequence in this
area. Cayeux (1960 ; 21-25) quotes large extracts from Destombes’ paper, but adds
to this important new information pointing out that the sequence in the Poudingue
at Cauville shows marked lithological variation to those seen elsewhere. The
account by Rioult (1962 ; 39-42) of the section at Cap de la Héve is very useful and he

102 MIDDLE ALBIAN STRATIGRAPHY

also presents a comparison with the Albian sequence in the Isle of Wight and the
Dorset and Devon coastal area. All three papers ably summarise the earlier
literature.

As Cayeux (1962 ; 2) has pointed out, the stratigraphy of the Albian sediments
between Cap de la Héve and St. Jouin needs revision. Such a revision requires
patient study because of the rarity of age diagnostic fossils. The following account
is quite obviously incomplete but it adds some new information on the succession and
provides a much more accurate foundation for the correlation of the sediments seen
in the Pays de Caux with those of the Isle of Wight. In this section of the work the
full exposed sequence of the Aptian and Albian sediments up to the base of the Gaize
is recorded. The terms ‘ Poudingue ferrugineux ’, & ‘ Argiles du Gault ’ of Lennier
(1867) cannot be accurately delimited in all the sections.

It is essential that these sections are worked with the greatest care particularly in
the early spring months. Winter frosts and the general freezing of ground water
cause the shattering of the Chalk in the cliffs and large blocks can be dislodged merely
by the ringing note of a hammer or by the slight vibration of a heavy surf. Major
cliff falls are not infrequent during the early months of the year, but the collecting is
far better at these times !

The sections described below indicate that the cliffs between St. Jouin and Cap de
la Héve present a cross section through a depositional trough. The Lower Albian
sediments are slightly comparable to those of the Isle of Wight, but those of the
Middle Albian are quite different.

(a) St. Jouin

The section described in text-fig. 45 is exposed about 300 yds S. of the cliff-top car
park west of St. Jouin. It has not previously been described, and includes Lower,
Middle and Upper Albian sediments up to the base of the Gaize. Unfortunately no
age diagnostic Middle Albian fossils have been found here by the writer and the
correlation of this section with that of Cauville (text-fig. 49) is based purely on the
lithology, and is, therefore, suspect in detail.

(b) Cauville

From approximately 50 yds SW. of the waterfall to about 300 yds NE. of it there
are good sections interrupted by cliff falls (text-fig. 46). As Cayeux has indicated
(r960 ; 23), there are striking variations in the Poudingue, the sediments of the
mammullatum and ?tardefurcata Zones, in this area. This is well shown in the four
sections described here for the first time, however, the Middle and Upper Albian
sediments remain reasonably constant.

Beds 1-9 are of Lower Albian, essentially mammillatum Zone, age. In the Bucaille
collection in the Muséum d’Histoire Naturelle, Rouen, there are three specimens of
Hoplites (Isohoplites) and one specimen of Hoplites (H.) which come from Cauville.
They are preserved in blackish phosphate with traces of the inner nacreous layer of
the shell preserved, and with evidence of pyritic inner whorls and glauconitic loamy

ite]
a
a

14

Ww

— N

FIG. 45.

IN THE ANGLO-PARIS BASIN

Lithology
First concretionary stone band of the Gaize.

Tough blocky grey sandy malm with scattered pale phosph-
atic nodules.

Argillaceous dark grey silty glauconitic sediment resting
on the guttered upper surface of Bed 13 below, becoming
more of a clay upwards, then passing into about 6 inches
of hard shaley grey shelly clay.

Highly glauconitic blackish loam resting ona guttered
surface of Bed 12 below, Pyritic concretions occur
throughout.

Highly glauconitic very dark grey loam a bit pebby
passing down by rafts into sparsely shelly loam resting
on guttered surface of Bed 11 below.

ough tawn sandy clay with brown phosphatic nodules
with dark interiors, septarian at the top.

Very pebbly grey loam with crushed bivalves,

Dark grey less pebbly clay than Bed 8 with intercalations of
dirty yellowish sand. Scattered nacreous shells. occur,

Pebbly dark grey ill-graded fossiliferous sandy clay with
nacreous shells preserved.

Hard gritty concretions in tough grey pebbly clayey matrix.
Tough ill graded ferruginous loam with phosphates & grit.

Ferruginous pebbly sand with some streaks of grey clay.
Lenticles of grit occur near the top.

Scattered sandy phosphatic nodules in brown pebbly sandy loam.

Rather pebbly streaked grey loamy clay and brown sand,

Mottled grey sandy clay-loam, and brown sand less pebbly
than Bed 1 into which it passes,

Pebbly dark grey green loam with streaks of brown sand
becoming fewer downwards. In the top of the bed there
occur pebbly concretionary sandstone lenticles 4-Sinches thick.

103

fe}

Sequence in the Poudingue and Gault in the sea-cliff, c. 150 yds SW. of the cliff
path exit, St. Jouin Bruneval, Pays de Caux.

Lithology

Bed
| 16 | Hard light-grey concretionary sandstone, Basal bed of the Gaize.
| 15 | Light-grey cross-bedded sandy marly clay with a few concretions.

Grey glauconitic claytoam with black, buff-rinded, phosphatic nodules at top and. bottom

(v) Dark grey glauconitic slightly pebbly loam.

13 (iv) Thin seam of irregular blackish light-rinded phosphatic nodules in loam. /

(iii) Lighter grey glauconitic sparsely shelly loam,

(ii) Thin seam of scattered blackish light-rinded phosphatic nodules in loam.

(i) Darkish grey glauconitic shelly loam,

|
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|
i
|
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|
|
|
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|
|
pares | Streaked dark green glauconitic sandy pebbly loam, with a few phosphatic nodules, }

Very shelly gritty glauconitic fawny pebbly loamy clay with rolled fragments of fawn clay in the basal few a
12 inches, pebbly patches at the top, and a few concretions. S
a
Black clay channelled into the b low. Wwe >
9 Irregular bed of pebbly ferruginous sand with large quartzite pebbles and phosphatic nodules, fp

Grey greenish glauconitic loam with a 4inch thick oxydised zone at the top, In the two end sections there
is a single bed of grey pebbly sandy concretions, very hard, with phosphatic nodules in the lower part, |
8 Between the two end sections, an upper bed of less pebbly grey green sandstone lenticles appearsin the
sequence. The lower lenticles are underlain by dark grey pebbly glauconitic loam with gritty and pebbly
phosphatic nodules at the base.

190 yds NE of waterfall Between path and waterfall

(vi) Tough mottled brown-dark grey loam becoming
coarser and darker upwards with pebbles at top.

(v) Grey green pebbly glauconitic loam with patches of
pure quartz sand, the whole extensively burrowed

(v) Dark grey sandy clay and yellowish sand with line
of irregular ey phosphatic nodules at top and
Gv)iEuwinish) pebblyniicam small Hard placquettes at the base.

(iv) Variegated greenish grey-brownish loam,

(iii) Very pebbly grey-green loam,
(iii) Coarse glauconitic loam.

(ii) Bed of pebbles with a few gritty phosphatic (ii) Thin ferruginous pale-hearted fine-grained
nodules. concretions,

a few pyrite nodules and wisps of white sand, (i) Coarse glauconitic loam,
together with brown phosphatic nodules.

KITCHINI

(ii) Mixture of ferruginous sand and blackish clay in streaks.

|

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(i) Dark grey gritty glauconitic micaceous clay with |
|

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(i) Coarse dirty yellow sand. |
|

|

Grey sandy clay and dirty yellow sand mixed together in a mottled loam,

Yellowish bedded sand with glauconite: some thin—bedded and cross-bedded units. Nodules of phosphatised
grit occur scattered throughout,

Irregular shaped masses of ferruginous pebbly grit.

Dirty yellow sand with glauconite,

(v) Concretions of massive ferruginous pebbly grit, with interstitial yellow sand.
(iv) Yellowish sand.

(iii) Slightly cemented ferruginous pebbly grit passing into 5 (iv) above.

(ii) Coarse yellowish sand grading into 5 (iii).

(i) Ferruginous pebbly cemented grit grading up into 5 (ii).

300 yds NEof the 270 yds NE of the 190 yds NE of the Between path and

waterfall waterfall waterfall waterfall

o "1
°
oJ
3
6
3 o 74
-. 6
r* ae : 0 64
Peg ate
; “lat a 54
Bie “a fe)
6 7 all
°
)
°
10 2
2
9
1
6
Fic. 46. Section in Poudingue and Gault in the sea-cliff at Cauville, :

Pays de Caux (Seine Maritime).

106 MIDDLE ALBIAN STRATIGRAPHY

Bed Lithology Ft Ins M

First concretionary stone band of the Gaize. pit) cI=F] 12
Silty glauconitic mid-grey clay at base becoming siltier and 6 ce)
lighter in colour upwards as clay content decreases. 11

Phosphatic and pyritic nodules occur scattered throughout.

Definite break in sequence at the base.

10

Tough mid-grey, sparsely shelly slightly glauconitic clay,
silty at base, becoming progressively less silty upwards.
Scattered shells occur throughout.

ao
°o

W
oS

Dark grey-greenish glauconitic loam with scattered shells.
Thin seams of light-rinded brown phosphatic nodules occur
as shown. The basal line contains gritty phosphatic nodules,
the uppermost line contains septarian phosphatic nodules.

I> TU

Dark grey—black greenish tinged glauconitic loam; a little
7 pebbly with very scattered gritty phosphatic nodules, and
shells,

Definite break in sequence at the base,
6 Dark grey—green glauconitic loam with small phosphatic
nodules, and shelfs.

Transitional bed with
upwards. Dark brown

=
=

dark gre sand lay content increasi
hosphdtic. tg d bottom. 9 10 5

ic nodules top an

Brownish sand with a 1-2inch bed of pure running grit at the

A
ro) =
oa

3 5
4 base. The sand is slightly glauconitic in the lower half, but
upper half is seamed with glauconitic wisps. 4
=
3 Hard dark brown very pebbly indurated grit with a few 2 ie)
LL phosphatic nodules.
3
Mottled brown and dirty yellowish sand with a few brownish
2 clay wisps, becoming ferruginous in the upper foot or so. 4 2
= 2
Heavily bioturbated mottled loam consisting of a churned-up
1 mass of grey sandy clay brown and dirty white sand in 6 4
wisps and pockets. Bed becomes more sandy downwards, 1
and contains scattered soft buff phosphatic nodules,
ie)

Fic. 47. Section in Poudingue and Gault in the sea-cliff immediately to the NE. and below
the cliff-top car-park, Octeville, Pays de Caux.

IN THE ANGLO-PARIS BASIN 107

matrix. This matrix is very similar to Bed 11. Bed 12 at the base shows angular
pieces of clay indicating heavy erosion of previously deposited clay sediment. The
bed itself is sandy and contains very fragile but well preserved Inoceramus concen-
tricus, Anahoplites planus, A. splendens, and Dimorphoplites miobe, indicating the
niobe Subzone not previously recognised in the Pays de Caux. The loams of Bed 13
also contain J. concentricus in the lower 1 foot 3 inches (0-38 m.) but apparently no
ammonites. Post eodentatus Subzone Middle Albian sediments are, therefore,
present in this area. The Upper Albian sediments probably commence at some level
within Bed 13.

(c) Octeville

This section (text-fig. 47) also has not yielded age diagnostic Middle Albian fossils
to the writer. It is situated in the cliff immediately to the NE. of the cliff-top car
park, and again has not previously been described in detail. The lithological correla-
tion with the sequence at Cap de la Héve and Cauville is, however, more definite than
that of St. Jouin and Cauville (text-fig. 49).

(d) Cap de la Héve

This famous section can be seen high in the cliff on the northern side of the Cap
(text-fig. 48). It has been studied by a number of workers, but the best recent
description of the sequence is that given by Rioult (1962 ; 39-42). Cayeux (1960 ;
25) has recorded Goodhallites goodhalli from the base of the bed here numbered 6, and
Bed 4 is of mammullatum Zone age. In the Bucaille collection at Rouen, there is a
single example of Hoplites (H.) preserved in blackish, gritty phosphate stated to have
come from Cap dela Héve. The preservation is identical to fossils found occasionally
in Bed 5 which, therefore, probably contains eodentatus and possibly basal lyelli Sub-
zone fossils in remanié.

(e) Summary, and comparison with the Isle of Wight

The lithological correlation of the sections described above is shown in text-fig. 49.
It can be seen that Lower and Middle Albian sediments reach their maximum develop-
ment in the area of Cauville. Eodentatus Subzone sediments are present at all
localities with the possible exception of St. Jouin where it has not been proved. At
Cauville, the eodentatus Subzone is apparently less condensed, and is followed by
sediments of the mio0be Subzone. To what extent higher Middle Albian sediments are
present, if at all, is unknown at this time. At Cap dela Héve it appears that Upper
Albian sediments rest directly upon the residue of basal dentatus Zone age.

Even if the classification of these sections is unsatisfactory, it is immediately
apparent that all previous correlations of the ‘ Argiles du Gault ’ of the Pays de Caux
with the Gault of the Isle of Wight are no longer tenable(p.42). It appears that the
sequences in the Isle of Wight and the Pays de Caux are truly out of phase. In the
Isle of Wight the kitchint Subzone of the mammillatum Zone (Lower Albian) is

108 MIDDLE ALBIAN STRATIGRAPHY

represented vemamnzé in the basal pebble bed of the Carstone ; in the Pays de Caux it
is well developed at Cauville. The eodentatus Subzone is represented at the top of the
Carstone by sediments containing indigenous Hoplites (Isohoplites) ; in the Pays de
Caux it is present but very condensed even at Cauville. The major difference, how-
ever, occurs in the Gault. In the Isle of Wight the clays are of lyellt, spatht, and
probably intermedius Subzones age, followed possibly directly by some orbignyi
Subzone sediments (Upper Albian) ; the bulk of the loricatus and the whole of the
lautus Zone being absent. In the Pays de Caux no sediments of any of the Subzones
mentioned above have yet been detected. However, the niobe Subzone is present at
Cauville, although absent in the Isle of Wight, and the lowest Upper Albian Subzone
yet proved is varicosum.

Bed Lithology

8 Basal Ienticular marly sandstone concretions of the Gaize,

7 Light grey micaceous silty malm.
6 Mid grey silty micaceous clay becoming a highly glauconitic

Ins M

varicosum

loam by the middle of the bed.

5 Very pebbly ill-graded greenish loam with pebbly phosphatic nodules. \[eod. re peers

Massive indurated blocks of pebbly pale-brown grit in a matrix
of loose similar material.

Darker brown ill-graded pebbly grit with streaks of dark grey
clay. Lenticles of indurated material occur,

Loose pebbly brown grit with little clay, A few scattered phosphates)

Very coarse brown ill-graded pebbly grit, loamy with streaks of
grey brown clay in the middle. Large blocks of brown hard
sandstone, as shown, inlower mottled yellow sand & fawn loam.

mammillatum

Fic. 48. Section in Poudingue and Gault in the sea-cliff on the N. side of Cap de la Heéve,
100 yds N. of the lighthouse, Ste Adresse, Le Havre, Pays de Caux.

It is probable that the Isle of Wight and the Pays de Caux belong to two separate
depositional troughs (p. 142) and do not form one area of deposition as implied by
previous workers. The sediment of the ‘ Argiles du Gault’ is a sandy loam rather
than a clay and small pebbles are present. This together with the Poudingue below
indicates the close proximity of a shoreline but outcrop and borehole information
indicate that this could hardly be to the south (see p. 142).

10g

IN THE ANGLO-PARIS BASIN

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110 MIDDLE ALBIAN STRATIGRAPHY

With the completion of the description of the individual sections in England to-
gether with a brief review of those in France, it is now possible to consider in detail
the Zonal and Subzonal scheme of the Middle Albian. It is instructive and sobering
to examine also the history of the development of this particular scheme, not atypical
of many zonal schemes. There are workers who would insist that zonal schemes
should be fixed for all time with little regard to future detailed work, or whether the
scheme is based upon firm foundations.

IV. DEFINITION OF THE MIDDLE ALBIAN SUBSTAGE AND ITS
ZONAL SCHEME IN THE ANGLO-PARIS BASIN

A. Historical background

The Formation name Gault was accepted in much the same sense by English and
French geologists within the first half of the 19th Century. The history of its use in
England was given by Jukes-Browne (1900 ; 14-31). D’Orbigny took William
Smith’s concept, that individual formations could be determined by their fossil con-
tent, a major step forward when he recognised that fossils characteristic of one litho-
logical unit occurred in different lithologies and that these, although deposited at
different localities, were formed at the same time. He erected, therefore, a series of
chronostratigraphic stages to include these diverse lithologies. Apart from the
localities mentioned by d’Orbigny (1842 ; 404-5) in his definition of the Albien stage
(latinized to Albian), he recorded others in 1849 (¢m Geinitz 1849 ; 6-7). Pictet &
Campiche adopted d’Orbigny’s stage name when they commenced their description
of the Cretaceous fauna of Ste Croix, Vaud, Switzerland, subdividing it into Albien
infériur, moyen and supérieur (1858 ; table facing p. 27).

De Rance (1868 ; 163-171) was the first worker to describe the Gault section at
Folkestone in detail. He accepted d’Orbigny’s term Albian and divided it into lower
and upper divisions drawing the boundary between them at what is now known as
the junction between Beds VIII and IX. The lower division corresponds approx-
imately, therefore, to Pictet & Campiche’s Albien moyen. He recognised eleven beds
in the Folkestone Gault and referred each to a zone based on its characteristic fossil ;
employing essentially the characteristic ammonite. Later Price (1874 ; 342-368)
revised De Rance’s description but neither of these workers on these occasions
attempted to apply their zonal scheme to sections other than at Folkestone.

Barrois (1875b ; 707~714) was the first geologist to formally define a zonal scheme
for the Albian of the Anglo-Paris Basin in the sense that we use today. This idea
of the application of an index fossil denoting a segment of time and represented by
different types of sediment, or none at all, differed from that of De Rance & Price who
used them merely in a local sense for an actual lithological unit. However, the idea
crystallised by Barrois for the Albian is implicit in the writings of earlier French
workers including d’Orbigny. Barrois recognised a tripartite division into :—

Zone a Ammonites inflatus
Zone a Ammonites interruptus
Zone a Ammonites mammillare

Of these only the lower two zones were included in the Albian, the Zone 4 Ammon-

IN THE ANGLO-PARIS BASIN IIt

ites inflatus being included by him in the Cenomanian. Nonetheless, the terms ‘ Zone
of Ammonites mammilare ’ and ‘ Zone of Ammonites interruptus ’ were first used by
De Rance (1868) at Folkestone. Barrois stated that the type area of the interruptus
Zone was the Aube but he recognised that in this area there was a mixture of what he
thought to be the fauna of the mammillare Zone (e.g. Dowvilleiceras of the clemen-
tinum group) and that of the interruptus Zone in the clays classified with the latter
Zone. In 1878 (265, footnote) it was obvious that Barrois was worried by this
admixture for he states that possibly Ammonites lyelli might have been preferable as
the index of this zone ; this species being characteristic of the clays of the Gault
classified with the interruptus Zone from the Aisne to the Yonne.

Price & Delatour (1879 ; 38-42) concluded that all the beds of the Lower Gault at
Folkestone had their representatives in the Gault of the Aube, but neither of these
workers had seen the sections on the opposite side of the Channel. Price and Barrois
knew each other, and it is significant that Price also refers to a Zone of Ammonites
lyelli and places it above the Zone of Ammonites mammillaris. Barrois, like many
of his contemporaries in France, when considering the zonal scheme, was strongly
influenced by the very full development of the mammillatum Zone, and the eodentatus
and /yell1 Subzones in the Gault of the southern part of the Paris Basin. Although
Barrois knew the sections at Wissant and Folkestone, and realised that there was a
break in the succession below the sediments of the inflatus Zone, he does not appear to
have fully grasped the extent of this gap in the observed sequence in the Aube and,
therefore, the reason why the sequence in the Aube appeared so different from those
on each side of the Channel. The observed gap in the southeastern area of the Paris
Basin involves the equivalent of the greater part of the Lower Gault at Wissant and
Folkestone, and includes all from the top of the intermedius Subzone to the base of
the cristatum Subzone. Whether there is a total absence of deposits of this age in
this area is uncertain (p. 97).

Jukes-Browne (1900 ; 45) adopted the Zone of Ammonites interruptus but
restricted it to beds of equivalent age to that of Bed I at Folkestone, the interruptus
Zone of De Rance in part and Price. This reading, excluding the equivalent of the
“Sulphur Band’ brought the English interpretation more into line with the known
sequence in the Aube. He realised that the remainder of the Lower Gault at Folke-
stone could not be classified with the interruptus Zone, and he proposed that Beds II
to VII and their lateral equivalents be included in a Zone of Ammonites lautus. He
excluded Bed VIII from both the lautus and rostratus zones (the latter being the
equivalent of Barrois ’ Zone a Ammonites inflatus) treating it as a junction bed.

Because Barrois had classified his Zone 4 Ammonites inflatus with the Cenomanian,
Jukes-Browne felt that he could not accept d’Orbigny’s name Albien for the stage.
He had proposed the name Devisian to encompass the Gault and Upper Greensand of
England (1892 ; 266), but this name clashed with the earlier Oxfordian substage
name Divesian and the term was replaced by Selbornian (1900 ; 30-31). In reality
there was no need for a new name and the terms Devisian and Selbornian are
synonyms of d’Orbigny’s Albian. Jukes-Browne’s zonal scheme and stage name
(1900) were employed in England until Spath commenced work on the nomenclature
of the Albian (1921).

Er2 MIDDLE ALBIAN STRATIGRAPHY

Kilian (1907 ; 62, 67) presented the following scheme influenced probably by the
work of his pupil Jacob.
4 Zone der Schloenbachia (Mortoniceras) inflata Sow. sp.
(mit zwei Subzonen)
3 Zone des Hoplites dentatus Sow. sp. und Acanthoceras
Gault stufe lyelli Leym. sp.
(Albien D’ORB.) 2 Zone des Hoblites tardefurcatus Leym. sp. und Hoplites
vegularis Brongn. sp.
I Zone des Parahoplites Nolani Seunes sp. (sog.
Milletianusschichten) und Douvilleiceras nodosoco-
statum D’ORB. sp., D. Bigoureti Seunes sp.

For the first time Lyelliceras lyelli is used formally in a zonal scheme. Jacob’s
thesis was completed in 1907 but not published until the following year (1908 ; 208—
590, pls. I-VI) when he presented the stratigraphical results of his study of the
middle part of the Cretaceous of the French Alps and adjoining regions. He divided
the Albian into the four zones which appear in Kilian’s work but in a simplified form.
His scheme is as follows (1908 ; 296).

VIb_ Sous-Zone a Mortoniceras inflatum Sow. sp.
et Turrilites bergert Brong.
VIa Sous-Zone a Mortoniceras hugardianum d’Orb. sp.
V_= Zone a Hoplites dentatus Sow. sp.
IV Zone a Hoplites (Leymeriella) tardefurcatus Leym. sp.
III Zone a Douvilleiceras nodosocostatum d’Orb. sp. et
Douv. Bigoureti Seunes sp.

The Clansayes horizon (Kilian’s Zone 1, Jacob’s Zone III) is, therefore, added to the
Albian. Jacob could not accept Barrois’ zone a Ammonites mammillaris because he
considered that this form was equally abundant in the following horizon (i.e dentatus
= interruptus Zone), mistaking forms such as Douvilleiceras clementinwm (d’Orbigny)
for ‘Ammonites mammillaris’. So, he renamed the zone ‘Zone IV a Hoplites
(Leymeriella) tardefurcatus’ (= Zone of Hoplites tardefurcatus and Hoplites regularis
in Kilian). Kilian and Jacob correct the specific name of the interruptus zone to that
of dentatus, but Jacob does not adopt the index Acanthoceras lyelli. When revising
the section at Sainte Croix, Vaud, Switzerland, Jacob indicates (1908 ; 291) that
Zone IV = I’ Albien inférieur, V = |’Albien moyen, and VIb = 1’Albien supérieur of
Pictet & Campiche (1858).

Jacob seems to have misread Jukes-Browne (1900) for it was not the term Gault
that Jukes-Browne could not accept, but the term Albian in the sense of Barrois
(Jacob 1908 ; 584). The re-inclusion of the Zone 4 Ammonites inflatus of Barrois
(Kilian’s Zone 4, Jacob’s Zone VI) in the Albian, a return to d’Orbigny’s original
definition in terms of lithology, removed the cause of Jukes-Browne’s objection to the
term Albian. Neither Jacob nor Kilian accepted or even mentioned Jukes-Browne’s
Zone of Ammonites lautus. This is understandable when one considers that they
were strongly influenced by the succession in the Aube (1908 ; 547), with which Price
& Delatour had given a completely inaccurate correlation of the beds at Folkestone

IN THE ANGLO-PARIS BASIN 113

(1879 : 1880 ; 38-42). Moreover, the time span represented by Beds II to VII at
Folkestone are represented in the Alpine area of France and adjoining Switzerland by
very condensed deposits in which many horizons are not represented at all. At that
time the only description of any detail of the section near Wissant was a poor one by
Barrois (1879 ; 27-28) an abridged version of which was given by Jukes-Browne
(1900 ; 378-381). Although Barrois had been unable to recognise exactly at Wissant
the equivalents of the beds numbered by Price at Folkestone, Price stated definitely
(1879 : 1880 ; 34) that they held good at Wissant. So, according to Price, the
numbered sequence at Folkestone could be correlated not only with the Aube but
also with Wissant. The gravity of this error can be judged from the stratigraphical
account of the French sections given above (p. 80) where it is apparent that a major
part of the Middle Albian sequence in the Aube is not represented at Wissant and
vice versa. Although Jukes-Browne agreed with Barrois that the exact equivalents
of the Beds at Folkestone could not be recognised at Wissant, he perpetuates Price &
Delatour’s erroneous correlation between Folkestone and the Aube (1900 ; 388-9).
There is no doubt that this strongly influenced subsequent work on both sides of the
Channel, and this is the reason why French geologists of that period considered that
the beds at Folkestone included by Jukes-Browne in his Zone of Ammonites lautus
were of the same age as part of the clays in the Aube included by them correctly in
the interruptus (dentatus) zone. This led to the time spans, which I now label
mammuillatum, loricatus and lautus Zones, being not recognised at all in Kilian or
Jacob’s zonal schemes. The historical background to the zonation of the Middle
Albian only will now be considered below. That of the Lower Albian has been dis-
cussed by Casey (1961a ; 492-499) and the Upper Albian will be discussed elsewhere.

In Germany the term ‘ Gault ’ included clays of Aptian as well as Albian age (see
for example Kilian 1907-13). It had been divided into lower, middle and upper
Gault within the first half of the 19th Century and the question of nomenclatorial
priorities has been discussed by Spath (1942 ; 670-671). Stolley (1908 ; 246-7)
recognised in the ‘Oberen Gault ’ of north Germany the zones already well established
in France. The ‘Zone des Hoplites interruptus’ No. 6 is represented by the
Minimus Tone and is followed above by the ‘ Zone der Schloenbachia inflata und
Puzosia planulata’ No. 7 to include the Flammenmergel. Jacob’s scheme con-
tinued to be used in France (e.g. Tomitch 1918, Ciry 1927, Houdard 1940).

Spath’s work on the classification of the Albian commenced with the following
arrangement (19214, 32).

Gault Up. & Mid. Albian Zones 7 & 6 of Stolley (1908)
Mammillatus Bed Lower Albian eS tt se (LQOS)
Later in the same year, Spath amplified this classification (I92Ib ; 311).

Upper Albian Hor. IX—XIII (Folkestone) Hor. 7 (Stolley)
(Upper Gault) VI (Jacob)

Middle Albian Hor. I-VIII (Folkestone) Hor. 6 (Stolley)
(Lower Gault) V (Jacob)

mammillatum bed

The mammuillatum Zone is, therefore, now included in the Middle Albian. In 1922
H

114 MIDDLE ALBIAN STRATIGRAPHY

Spath produced a scheme of ammonite horizons which foreshadowed the zonal scheme
which appeared the following year (1922 ; 96). He was far too critical of the previous
work on the Albian and certainly misread Jukes-Browne, and so in 1923 he dis-
carded the broad zonal classification of previous workers in favour of the scheme
shown in Table 2. This zonal scheme was strongly influenced by the sequence at
Folkestone which Spath had examined in detail (1923a ; 73 : 1923b ; 4 : and see also
1923c). Unfortunately, in England the sequence between the ‘ mammillatus Zone’
and the ‘dentatus Zone’ was at that time very imperfectly known. The ‘ inaequin-
odus Zone’ formed a very uncertain taxon, and the ‘ benettianus Zone’ was par-
ticularly ill-founded for Spath had no idea of the exact stratigraphical position of the
lyellu fauna (1923c : 142, see 1926b ; 422), and he was later to classify even high
spatht Subzone sediments with that Subzone. De Rance had already used the term
‘benettianus Zone ’ in a different sense, and it is unfortunate that Spath ignored the
sequence in the southern part of the Paris Basin. In effect he restricted the dentatus
Zone to exclude much of what the French workers understood by this term. He also
discarded Jukes-Browne’s lautus zone replacing it with zones of intermedius, delaruet
and cornutus, but added a cristatus zone for Bed VIII and its lateral equivalents.
It was unfortunate that this scheme should have been presented before Spath had
fully studied the species he had used as indices. In the following two years he had
to alter this zonal scheme (1924 ; 505 : 1925b ; 31-36), and the year after (1926b ;
421-2, 425) saw the extensive modification shown in Table 2 (p. 116).

In 1926 Spath first renamed the lower part of his cornutwm Zone, the Zone of
Euhoplites alphalautus (1926a ; 154 footnote 1), and then later recognised that this
species was a form found in the varicoswm Zone. The major change in the scheme
given by him later that year (1926b ; 421-2, 425) was the relegation of his zones to
the rank of subzones in the Table on p. 421. However, the presentation shows
several inconsistencies for he refers to these subzones in the text as zones. In the
table he groups the subzones into the ‘ old zones’ which are in fact those used by
Jukes-Browne, the names being corrected where necessary. Spath included the
intermedius Subzone in the dentatus [olim interruptus] Zone, but Jukes-Browne had
in fact included Bed II at Folkestone in his lautus zone. Spath recognised, however,
that the zonal schemes which had been used in Europe were of provincial value only,
and this was a significant step forward.

The zonal scheme was further modified during the course of publication of successive
parts of Spath’s Monograph, and one saw the firm readoption of the broader zones.
He formally presented the various emendations to his earlier scheme in I94I (1941 ;
668) and discussed them briefly the following year (1942 ; 671-673) : these are given
in Table 2. His ‘ zones’ of 1923 are now emended and reduced to the status of
Subzones which are grouped into three Zones. The mammillatum Zone is much the
same as that of Barrois and Jukes-Browne, however, the dentatus and lautus Zones
do not correspond with the views of earlier workers despite Spath’s comment (1942 ;
672 footnote 3). Is the arrangement given by Spath an improvement ? The junction
between the dentatus and lautus Zones was placed by Spath at a level where there is
no significant change in the fauna and the arrangement is quite arbitrary.

The older zonal grouping of French and English workers such as Barrois and Jukes-

IN THE ANGLO-PARIS BASIN II5

Browne was based on a comprehensive knowledge of all the sections then available.
Even if they did not consider the detail essential to modern work to be of great
importance, they possessed a broader picture than many later workers, some of whom
had never examined the sections in the country separated from them by the Channel.
Superficially Spath’s early stratigraphic work appears to have given far greater
precision to the zonation of the Albian, but this was not so. His zonal scheme was
introduced without sufficient initial research and suffered greatly by the early need
for radical alteration, and in the end his zonal boundaries were ill-chosen. He
appears not to have examined the French sections (1943 ; 722-3), which is most
unfortunate as it is absolutely essential to have some first-hand knowledge of them.
There is no question, however, of the immense value of his contribution. Without
his work on the Albian ammonites the progress made in the study of the stratigraphy
during the last 25 years would have been very slow indeed.

Breistroffer (1947) made the first revision to Spath’s zonal scheme in an important
paper comparing essentially the French with the English succession. In this work
he includes the mammullatum Zone in the Lower Albian and the cvistatum Subzone in
the Upper Albian (1947 ; and Table 2 herein). Casey (1950 ; 270) noted Breistroffer’s
reading of the mammillatum Zone but followed Spath in including it in the Middle
Albian. Khan (1952 ; 73) produced a useful emendation when he included the
subdelaruet Subzone in Spath’s sense, in the dentatus Zone ; thus placing the junction
between the dentatus and lautus Zones at a point where there is some significant
change in the ammonite funa. Casey, however, again followed Spath in terminating
the dentatus Zone at the top of the niobe Subzone (1954a ; 264). Milbourne (1956 ;
241) could not accept a separate subdelaruei Subzone in Spath’s sense. He included
the lower part of Bed IV and its lateral equivalents in the miobe Subzone, and the
upper nodule bed of Bed IV and its lateral equivalents in the lautus-nitidus Subzone
recognising that these probably fell within a distinct Subzone.

The writer in 1958 reviewed briefly the zonal scheme of the Middle Albian, placing
the zonal boundaries at levels where significant changes in the ammonite fauna
occurred (1958 ; 160-164). At the same time I drew attention to the difficulties that
existed in accepting Breistroffer’s emendations of Spath’s zonal scheme for the
dentatus and lautus Zones ; except for the position of the cristatwm Subzone, these
have not changed. Subsequently, I proposed formally that the time span repre-
sented at Folkestone by the upper nodule bed of Bed IV and the basal few inches of
Bed V and their lateral equivalents be recognised as the Subzone of Euhoplites
meandrinus (1960 ; 373, 376). The zonal grouping suggested by the author in these
two papers is shown in Table I, p. Io.

In 1961 Casey produced his important revision of the zonal scheme for the Aptian
and Lower Albian (1961a ; 492-499). He now follows Breistroffer in including the
mammillatum Zone in the Lower Albian with the exception of Spath’s inaequinodum
Subzone. For this Subzone he proposes a new index, Hoplites (Isohoplites) eodentatus,
and includes it in the dentatus Zone of the Middle Albian, pointing out that this
species is the most characteristic ammonite at this horizon in England and France.
The division between the mammillatum and dentatus Zones and thus the Lower and
Middle Albian now falls at a distinct change in the ammonite fauna.

MIDDLE ALBIAN STRATIGRAPHY

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IN THE ANGLO-PARIS BASIN 117

Milbourne (1963 ; 58) places the division between the dentatus and lautus Zones
above his niobe Subzone (which also includes the swbdelarue: Subzone of Spath in
part). He finds my Subzone of Euhoplites meandrinus unacceptable and substitutes
for it a Subzone of Dimorphoplites doris and Euhoplites neglectus. He places the
subzone in the /autus Zone and so once more the boundary between the two zones is
placed at a level where there is no significant change in the ammonite fauna. The
differences of opinion between Milbourne and myself led to a skilful review of the
zonal scheme by Hancock (1965). Unfortunately two errors were overlooked when
preparing Table I of Hancock’s paper (1965 ; 245). The eodentatus Subzone was not
recognised by me in 1958 but by Casey (1961), and Spath’s lautus-nitidus Subzone
did not include the time span referred by me to the meandrinus Subzone. The
meandrinus Subzone formed part of the swbdelaruet Subzone in Spath’s sense and it is
important to make this point absolutely clear.

In 1963 a colloquium on the Lower Cretaceous was held in France ; two very
important papers being contributed by P. & J.-P. Destombes, and Breistroffer, on
the zonal scheme of the Albian. These were published in 1965. P., & J.-P. Des-
tombes propose the following arrangement (1965 ; 266).

Zone Sous-zone Localité-type
2—Lyelliceras lyelli et La Vendue-Mignot
Hoplites benettianus
LYELLICERATIEN
1—Tegoceras camatteanum Cotes Noires de Moeslains
Isohoplites eodentatus niv. 5

This proposition is interesting as it shows in the case of Lyelliceras lyelli an
independent return to the older view expressed by Barrois, Price, and Kilian, al-
though it should be noted that Collignon has also used this index (1963 ; 2). P.&
J.-P. Destombes use a hemeral name for the zone. The use of these terms particu-
larly by Spath and Breistroffer, is not supported here for they are far too nebulous to
have any really precise application. Breistroffer’s paper (1965 ; 311 & table)
presents an emendation of his zonal scheme of 1947 and he accepts P. & J.-P.
Destombes subzonal arrangement for the basal part of the Middle Albian. The
scheme given by Collignon (1965b) is quite unacceptable.

The history of this zonal scheme, like that of any other, has been one of progressive
refinement as knowledge of the succession has improved. Unfortunately, the Albian
in particular has suffered greatly because of arbitrary decisions concerning the fixing
of ammonite zonal and subzonal boundaries. These boundaries have been placed
sometimes without sufficient initial research and have been upheld later purely on
rather dubious ‘ historical’ grounds ignoring whether or not there is a significant
change in the ammonite fauna. A zonal scheme must remain sufficiently flexible to
take account of new discoveries and better developed sequences. To bang in a
‘golden stake’ at a convenient level in a so-called permanent type-section might
help the theorist but in reality it only hinders progress towards accurate international
correlation, and the knowledge of events in the evolution of the Earth that will stem
from it.

118 MIDDLE ALBIAN STRATIGRAPHY

(B) The Zonal Scheme of the Middle Albian in the Anglo-Paris Basin
(i) DEFINITION OF THE BASE OF THE MIDDLE ALBIAN

Towards the top of the mammillatum Zone (puzosianus Subzone) in the Anglo-
Paris Basin the ammonite genus Pseudosonneratia (s.s. non Casey!) develops as a
minority element subordinate to such hoplitid genera as Protohoplites s.s., P. (Hem1-
sonneratia), Otohoplites, and Sonneratia as well as the common element consisting of
Douvilleiceras, Beudanticeras etc. The lyelliceratids are represented by very rare
specimens of Tegoceras.

Pseudosonneratia of the pusosianus Subzone is the direct forerunner of the simple-
ribbed non-lautiform species of Hoplites of the dentatus group. Casey (1954b ; 112,
1g61a ; 599) has separated off those species which are transitional between these two
genera as a Subgenus Hoplites (Isohoplites). They are characterised by a partial
interruption in the strength of each rib along the siphonal line as it sweeps across the
venter, but no general en echelon offsetting of the ventro-lateral rib terminations
occurs although the tendency towards this is often apparent. Casey has demon-
strated that Hoplites (Isohoplites) characterises an horizon above the development of
the typical mammuillatwm Zone fauna and corresponds approximately to Spath’s
inaequinodum Subzone (Casey 1961a ; 498 : Spath 1923b ; 73). He included the
Subzone in the dentatus Zone and subsequent work in France (P., & J.-P. Destombes
1965 : and herein) and England support this reading.

The junction between the mammuillatum Zone and the eodentatus Subzone has not
yet been seen in an uncondensed sequence in England. Sediments spanning the zonal
boundary have, however, been exposed in France at St. Martin in the Pays de Bray
(p. 100) andat Les Cotes Noires in the Haute Marne (p. 91). At both localities there is
a marked change in the ammonite fauna with the virtual disappearance of Otohoplites
at the top of the mammullatum Zone, and the appearance in bulk of Hoplites (Iso-
hoplites) at the base of the eodentatus Subzone above. In England the available
exposures of the sequence at this level show only condensed phosphatic nodule beds
in which the sudden change in the character of the ammonite fauna is probably
accentuated. The base of the Middle Albian in the Anglo-Paris basin is marked,
therefore, by the appearance of the genus Hoflites, and as is shown below by the
appearance of Lyelliceras.

1 Dr. P. Destombes has collected a very fine ammonite fauna from two localities in the Bois de Perchois,
Aube, which I have visited in company with him. It is apparent from the material he has shown me
that Pseudosonneratia iserensis figured by Casey (1965 ; 541 text-fig. 203e, f) is associated in a very little
condensed sequence—the ‘ Red Bed ’—at Perchois Ouest with species of Cleoniceras (Neosaynella) and
Cleoniceras s.s., indicating a flovridum Subzone age. It contains species of Otohoplites earlier than any yet
known in England, showing morphological gradations to the Pseudosonneratia-like forms which accom-
pany them in the same bed. The collection also contains species of Sonneratia with distinct hitchini
Subzone affinities, although higher in the succession, as well as Douvilleiceras, Beudanticeras and Pro-
tanisocevas. The fauna of Perchois Est, although preserved in a similar manner, occurs higher in the
mammillatum Zone sequence. The very important fauna obtained from these two sections is to be
described in due course by Dr. P. Destombes (see also Destombes 1965). The area is also of interest in
that at Perchois Ouest the grey clays above the ‘ Red Bed’ contains an ammonite fauna of distinct
tethyan aspect.

IN THE ANGLO-PARIS BASIN 119

(ii) THE SUBZONAL SEQUENCE (Table 1, p. ro)

(a) Subzone of Hoplites (Isohoplites) eodentatus

Recognised by Casey (1961a ; 498), although few details were given on that
occasion, this index replaces the Subzone of Douvilleiceras inaequinodum of Spath
(1923a ; 4 : 1923b ; 73 : 1941 ; 668). In England, the Subzone is represented in a
fossiliferous uncondensed state in the Isle of Wight (p. 52) and at Okeford Fitzpaine,
Dorset (54). Elsewhere, the known sections exhibit either unfossiliferous sediments
which probably represent this time span, or rvemanié phosphatic nodule beds. Future
deepening of the section at Small Dole, Sussex (p. 35) and at Badbury Wick, Wilt-
shire (p. 61) may provide a fossiliferous little-condensed sequence from which a direct
comparison can be made with the succession at St. Martin (p. 100). Maurupt (p. 89)
and Les Cétes Noires (p. 91) in France. Nowhere in England are the sections at the
outcrop in this subzone particularly fossiliferous.

In England, the ammonite fauna consists essentially of species of Hoplites (Iso-
hoplites) of which H. (I.) eodentatus is typical. It is necessary to bear in mind that
the tendency to produce the peripheral rib pattern of a typical Hoplites (H.) men-
tioned already, can be so well advanced that a fragment or even a complete individual
would have to be referred to Hoplites (H.). This fact alone shows the artificial nature
of the subgenus Isohoplites. However, these are associated with species in which
the Pseudosonneratia ventral aspect is still well developed. Apart from the forms
derived from Pseudosonneratia, there are other Hoplites derived from Otohoplites and
possibly Protohoplites, and in fact rare specimens of Otohoplites still occur. The
associated ammonites include Beudanticeras such as B. laevigatum, B. albense, and B.
santaecrucis, together with Douvilleiceras spp. including D. inaequinodum.

Lyelliceras of the camatteanum type, which stands rather in the same relationship
to Tegoceras on the one hand and Lyelliceras of the lyelli type on the other, as does
Isohoplites in the hoplitinids, is a great rarity in England. In France it is a reason-
ably constant and characteristic companion of Hoplites (Isohoplites) and the other
ammonites mentioned above, and this has led P. & J.-P. Destombes to recommend
that the Subzone be defined as that of Tegoceras camatteanum and Isohoplites
eodentatus (1965 ; 265-6). They indicate a type locality, Les Cétes-Noires prés de
Moeslain, where the subzone is represented by clays in an apparently unbroken
sequence from the top of the mammillatum Zone to the base of the lyelli Subzone.
In the lower clays classified with the eodentatus Subzone there is the association of
Hoplites (Isohoplites) eodentatus and Lyelliceras camatteanum as there is also at
Maurupt and St. Martin. If one wishes to use a double index then this emendation
proposed by P. & J.-P. Destombes should be adopted. However, as H. (I.) eodentatus
is common to this time span throughout the Anglo-Paris basin, the writer employs
this index only.

(b) Subzone of Lyelliceras lyelli

P., & J.-P. Destombes (1965 ; 265-7) have formally proposed that the benettianus
Subzone in Spath’s sense should be redefined as the Subzone of Lyelliceras lyelli &
Hoplites benettianus. So, the tentative suggestion of Barrois, the bald statement of

120 MIDDLE ALBIAN STRATIGRAPHY

Price, the formal proposal of Kilian and the employment of the name by Collignon
(1963 ; 2) given precision by P., & J.-P. Destombes, has at last taken root. H. (H.)
benettianus is not a satisfactory subzonal index for in the strict interpretation it may
be restricted to only a comparatively narrow horizon within the Subzone that it is
supposed to represent. They have proposed a type locality for this Subzone, the
Tuilerie Clerc at La Vendue-Mignot, Aube (p. 95), where the Lyelliceras fauna is con-
sidered to be in the pure state without risk of contamination (in terms of collecting
fossils that is). In fact the section does not show the relationship between either the
eodentatus Subzone below or the spatht Subzone above, and there are other objections
already mentioned.

The benettianus Subzone was defined without precision by Spath (1923a ; 4, 1923b ;
73, 1926b ; 422), and was used by him for a range of sediments some of which are
truly lyelli in age but others have subsequently proved to be of spathi age. It would
be far more satisfactory if the proposal of P., & J.-P. Destombes were adopted
modified to a single index of Lyelliceras lyellt.

In France, the finest section now available in sediments of this Subzone is certainly
that of Courcelles prés Clérey, Aube (p. 91), although that at Les Cétes Noires is
probably the most completely displayed in terms of sediments. The best develop-
ment seen in England was exposed at the Horton Clay pit near Small Dole, Sussex
(p. 35). A comparison of these two sections brings out the nature of the ammonite
fauna and the differences to be found between them. Lyelliceras camatteanum and
its related species of the eodentatus Subzone is connected with Lyelliceras lyelli by a
series of morphological transitions. Intermediate forms such as Lyelliceras pseud-
olyelli (Parona & Bonarelli non Spath), Lyelliceras huberianum (Pictet) and L.
hirsutum (Parona & Bonarelli) form such transitional species. These show the
transition from the extreme en-echelon arrangement of the ventro-lateral rib termin-
ations and non-tuberculate siphonal line characteristic of Tegoceras, to the single ribs
commencing at the umbilical margin and sweeping without break straight across the
periphery and bearing mid-lateral, ventro-lateral and siphonal clavi typical of
Lyelliceras lyelli. These transitional forms occur right at the base of the lyelli Sub-
zone both at Courcelles and to a limited extent at Small Dole, and on balance it seems
that the time span in which they existed was of comparatively short duration. Un-
condensed basal /yelli Subzone sediments are also apparently well developed in the
Céte d’Or (e.g. Ciry 1927 : Ciry, Rat, Malapris & Nicolas 1965).

It is apparent that ecological conditions have a marked effect upon the ammonite
fauna. In deposits of the lyelli Subzone containing a benthonic fauna, the hetero-
morph ammonites Protanisoceras (P.) barrense and P. (P.) alternotuberculatum are
common (e.g. Small Dole, and Courcelles) and are as equally characteristic of this
time span as is Lyelliceras lyelli. Where no benthos is present, or it is very reduced
in numbers, the heteromorph ammonites are absent or very rare (e.g. the Isle of
Wight). Lyelliceras lyelli is also affected by ecological conditions in that in beds
containing abundant Hoplites (H.), Lyelliceras is not common. Casey has already
noted the tendency to mutual exclusiveness between the earlier members of these
two families (1957 ; 43-44), although this could in part be due to the hoplitids being
able to withstand more adverse conditions in the seas of that time. However, the

IN THE ANGLO-PARIS BASIN 121

distribution of Beudanticeras laevigatum, B. sanctaecrucis and B. albensis is not
affected by either of these two factors. In fact the lyelli Subzone may be recognised
in areas in which sea-bottom conditions produced a sulphide facies in the sediments,
by the association of Hoplites (H.) spp. and these three species of Beudanticeras. The
distribution of Douvilleiceras is also not constant. It is rare at Small Dole, although
very common at the top of the lyelli Subzone at Courcelles. It was also not un-
common in Bed 14 at Shere (Owen 1963< ; 42).

The uppermost part of the lyelli Subzone and its junction with the overlying spathi
Subzone cannot alone be determined by the abrupt disappearance of Lyelliceras lyellv.
In England, at Small Dole, Sevenoaks (p. 23), and Westerham (p. 31), and at Courcel-
les (Aube), there are clays just below the spathi Subzone which still contain lyelli
Subzone species of Beudanticeras, Douvilleiceras (Courcelles only), and Protanisoceras
(P.), but Lyelliceras is absent. It is recommended that these sediments be included
in the lyelli Subzone for the significant change in the ammonite fauna occurs at the
top of them where all the non-Hoplites (H.), lyelli Subzone ammonite species vanish
abruptly from the sequence. In the English and French sections mentioned here
there is no apparent break in deposition and no change in facies. Pvrotanisoceras (P.)
still occurs in the spathi Subzone but it is very rare, and, in effect, the heteromorphs
are almost exclusively species of Hamites (H.) with subordinate Metahamites. There
is no major change in the species of Hoplites (H.) in the transitional period between
the lyelli and spatht Subzones. The transitional sediments at the top and bottom
of the lyell1 Subzone are of no great thickness in well developed sequences. Even
slight condensation produces an apparent sharp change in the ammonite fauna at the
subzonal boundaries.

(c) Subzone of Hoplites (Hoplites) spathi

H. (H.) spathi in its typical form occurs in this subzone but it is not very common,
although there is no dearth of closely related forms. It is possible that Spath
selected this species (1941 ; 668 : 1942 ; 672, under the preoccupied name bonarellit)
because in terms of ornament it stands mid-way between the more discoidal finer
ribbed species of the dentatus type and the inflated coarsely ornamented forms of the
maritimus-rudis group. The name dentatus-spathi Subzone is in any case too well
established in the recent literature both sides of the Channel to justify altering it,
except to reduce it to a single index of H. (H.) spathi.

The Subzone is well developed in England and the section at Small Dole is par-
ticularly important as it shows in a very fossiliferous little condensed sequence the
junction with the lyelli Subzone below and, albeit imperfectly, the intermedius Sub-
zone above. In France, the junction with the lyelli Subzone is seen in an uncon-
densed sequence at Courcelles, and the junction with the intermedius Subzone again
in an uncondensed sequence at Revigny-sur-Ornain (p. 88). The ammonite fauna
consists very largely of species of Hoplites (H.) of which the general evolutionary
characteristics of stratigraphical value have been given by me (Owen 1963a ; 49).
The very small percentage minority element in the fauna provides a good list of
genera and species (p. 152) some of which are of value in correlation with other faunal
provinces.

122 MIDDLE ALBIAN STRATIGRAPHY

The division between the /yelli and spathi Subzones has been discussed above. The
sediments representing the transitional period between the spathi and intermedius
Subzones do not contain ammonites in the area of the Weald except at Petersfield
(p. 34). At present it cannot be demonstrated in any one section in England, how-
ever, the section at Osmington (p. 51), Dorset, and Caen Hill, Devizes (p. 60) show
the extreme top of the spathi Subzone and the extreme base of the intermedius Sub-
zone respectively. Bed 2 at Osmington contains abundant high spathi Subzone
Hoplites (H.) together with rare early forms of Anahoplites of the intermedius group
such as A. osmingtonensis and A. grimsdalei spp. nov. In Bed 7 at Caen Hill,
Hoplites (H.) has become very subordinate to slightly later forms of Anahoplites such
as A. evolutus which are in turn earlier than Anahoplites intermedius and its con-
temporaries. A. evolutusis also known from Bed C at Hunstanton. These sediments
containing A. evolutus are here considered to mark the base of the intermedius
Subzone.

In France, fragments of Anahoplites osmingtonensis and A. evolutus occur in Bed 4
at St. Florentin (p. 97), a phosphatic nodule bed containing en melée material derived
from both the spathi and intermedius Subzones. At Revigny-sur-Ornain (p. 88)
these transitional sediments are uncondensed but the ammonites are mainly crushed
flat. At Wissant sediments deposited during this period have not yielded ammonites.

There is no difference between this interpretation of the spathi Subzone and the
views of French geologists (e.g. Breistroffer 1965 ; 313). From this account it is now
obvious that the parochial view expressed by Milbourne (see Hancock 1965 ; 246-7),
fixing the top of the spathi Subzone at an horizon of condensation in a comparatively
small area of the outcrop in the northern Weald, is totally unacceptable. The record
of Anahoplites intermedius a foot or two above the spathi nodule bed at Folkestone
(Bed I (vi) ) by Casey (in Hancock 1965 ; 247) is here considered to be a misidentifica-
tion of the finely-ribbed H. (H.) dentatus densicostata Spath which is just as common
in the higher part of the Subzone.

(d) Subzone of Anahoplites intermedius

Although there is some condensation at Folkestone, this section (p.14) provides the
best sequence yet known in this Subzone recognised by Spath in 1923. The sequence
at Small Dole is the least condensed and also shows, albeit imperfectly, the junction
with the spathi Subzone below, and also the junction with the nzobe Subzone above ;
however, the fauna is crushed flat. The junction with the sfatht Subzone has been
discussed above. The lowest part of the Subzone with Anahoplites evolutus has not
yet been discovered in the Weald and so at Folkestone the earliest intermedius Sub-
zone sediments containing ammonites yield Anahoplites intermedius and A. praecox.
In France the section at Wissant shows an imperfect development of sediments
representing this Subzone, but at Revigny-sur-Ornain, and at Courcelles the lower
part is well developed. .

The characteristic ammonites of this time span are the group of Anahoplites
typified by A. intermedius. The ammonite fauna of the Subzone as a whole is more
diverse than that of the spathi Subzone below. Hoplites (H.) is greatly subordinate
to the other genera, but the group which had produced rare Euhoplites by the top of

eS

IN THE ANGLO-PARIS BASIN 1736)

the spathi Subzone is well represented by forms such as E. loricatus, E. microceras, E.
subtabulatus and E. priceit. Early forms of Dimorphoplites including D. niobe
(praemutation) occur sparingly, and some of these show a tendency to the lautiform
ribbing not generally developed again in the genus until the meandrinus Subzone.
Heteromorphs are not uncommon and consist not only of Hamites (H.) spp., but also
of species of Protanisoceras (Heteroclinus). The early binneyitid Falciferella mil-
bourne: can be abundant at certain horizons. Apart from the endemic forms, there
is a much rarer element represented by specimens of Uhligella, Tetragonites, Des-
moceras, Eubrancoceras and Pseudhelicoceras (e.g. P. subcatenatum Spath.)

The upper limit of the Subzone is marked by the quite sudden decline of Ana-
hophites of the intermedius group, which do not extend into the miobe Subzone above.

(e) Subzone of Dimorphoplites niobe

The Subzone was recognised by Spath (1924 ; 505), it having previously formed
part of his intermedius Zone (1923a, b). It has been considered to be of local value
(Spath 1942 ; 672) but in fact it is of widespread occurrence. Sediments of this time
span are relatively uncondensed at Folkestone and to a lesser extent at Small Dole.
Elsewhere in the Weald the sequence is somewhat condensed, where indeed it has
escaped basal Upper Albian planation. Outside the Weald, it is developed in the
Leighton Buzzard district. In France, the Subzone may be developed at Wissant
(p. 83), but the only locality at which it can be proved with certainty is Cauville,
Seine Maritime (p.107). The restricted development of the miobe Subzone led Breis-
troffer in effect to include it in the imtermedius Subzone (1947 ; 44, 1965 ; table
opposite p. 312) but his view was probably also influenced by Spath (1942 ; 672).

The ammonite fauna is a curious one in that it has no species restricted to it, yet
it is distinctive. Anahoplites intermedius and its allies have gone, and Dimorphoplites
niobe, A. planus and A. splendens are the characteristic species. The upper limit
of the Subzone occurs immediately below the appearance of Mojsisovicsia in the
sequence.

(f) Subzone of Mojsisovicsia subdelaruei

Originally indexed by Spath (1923a ; 4, b ; 73) using ‘ Dipoloceras’ delaruei. He
realised later that this species did not occur in this time span, and it has been found
subsequently in the spathti Subzone. The Subzone as originally defined included
sediments now classified with the meandrinus Subzone. The Subzone is represented
by uncondensed deposits at Ford Place, Wrotham (p. 22) and at Sevenoaks (p. 25),
but elsewhere in England where deposits of this age are preserved they are condensed.
In France, they are known only in a condensed state (e.g. Bed 11 at Wissant).

Apart from the species of Mojsisovicsia, a genus which last made its appearance
in the Anglo-Paris basin in the spathi Subzone, and which in the swbdelaruet Subzone
shows marked evolutionary changes, the hoplitinid fauna shows more diversity of
form than in the niobe Subzone below. Mojsisovicsia subdelarue: appears at the
base of the Subzone and evolves to M. remota at the top. The occurrence of Dimor-
phoplites mobe led Milbourne to unite this time span with the miobe Subzone (1956 ;
241, 1963 ; 64), but on balance this incursion of keeled ammonites into the Anglo-

124 MIDDLE ALBIAN STRATIGRAPHY

Paris basin dictates that it be kept separate. The species of Ewhoplites show little
difference to those of the niobe Subzone below, but Dimorphoplites commences to
differentiate towards the forms seen in the meandrinus Subzone above.

The upper limit of the Subzone coincides with the last appearance of M. remota.
However, Mojstsovicsia is not at all common at this height and it is necessary to use
the grade of development shown by the species of Euhoplites and Dimorphoplites at
the base of the meandrinus Subzone, discussed below.

(g) Subzone of Euhoplites meandrinus

This time span, included by Spath in his subdelaruei Subzone, was indexed by
the writer (Owen 1960 ; 373, 376). Its separate nature was recognised by Milbourne
(1956 ; 241), who included it provisionally in the mit#idus Subzone, and by the author
(Owen 1958 ; 162). Unfortunately, there is an error in Hancock (1965 ; 245 Table I)
for this Subzone was not included by me in the mtidus Subzone, that is, in the basal
Subzone of the Jautus Zone as restricted by me in 1958. Milbourne (1963 ; 65)
indexed this Subzone with his ‘ species’ Euhoplites neglectus (which on examination
of the type material proves to be a synonym of EF. meandrinus) together with Dimor-
phoplites doris. My objections to this emendation were set out in Hancock (1965 ;
247) and this clash of opinion led to a mistake in the zonal scheme given by Kaye
(1965 ; 220).

The Subzone is characterised by EF. meandrinus and closely related forms such as
E. cantianus, E. loricatus (late mutation), and E. beaneyi which still possess the
deeply sulcate venter characteristic of the genus in the preceding subzones. The
late mutation of E. subtuberculatus tends to show the development of the channelled
venter characteristic in the lautus Zone on its outer whorl, but the inner whorls are
still sulcate. The pattern of ribbing on most of these species is transitional to that
of the typical Jautus Zone species. Dimorphoplites has become more diverse in form
and apart from the typical meandrinus Subzone species such as D. doris and D. pinax,
there are the early mutations present of the species which occur commonly in the
lautus Zone above but they are greatly subordinate in numbers. Mojsisovicsia is
absent from this time span.

In England, the meandrinus Subzone is present in an uncondensed sequence at the
Horton Clay Pit, Small Dole (p. 40). At the Sevenoaks Brick Works (p. 25) and
further east in Kent, the sediments are condensed to a variable extent. Its known
representation is confined to eastern England. In France, the only locality at which
the Subzone is known to be represented at this time is at Wissant in Bed 11 which
also contains material derived from subdelaruei Subzone sediments and mitidus Sub-
zone sediments as well.

At Small Dole, and at Ford Place, in particular, the sudden change from a sulcate
to a channelled venter in the species of Euhoplites can be well seen in sediments which
are uncondensed. This marks the base of the overlying mitidus Subzone.

(h) Subzone of Euhoplites nitidus
Spath originally included Beds V—VII at Folkestone in a zone of Dipoloceras
cornutum (1923a ; 4, b ; 73) but having realised the rarity of that species he divided

IN THE ANGLO-PARIS BASIN 125

this zone into two ; the zone of Ewhoplites alphalautus (1926a ; 154 footnote I) to
include Beds V and VI, and that of Anahoplites daviesi to include Bed VII (see also
1925b ; 34-35). Within a few months he realised that E. alphalautus was a form of
the varicosum Subzone, and substituted for it Ewhoplites lautus and E. nitidus (1926b ;
425). Breistroffer (1965 ; table opp. 312) has grouped the mtidus and daviest Sub-
zones as Subzone of mitidus + cornutum + daviesi. This view is probably influenced
by the section near Wissant where the daviest Subzone was thought to be present, but
where it is now known to be absent (p. 85).

In England, the best known development is at Folkestone where the relationship
with the meandrinus Subzone below and the daviesi Subzone above is well seen (p. 15).
In France, the Subzone is well represented at Wissant but the sediments there are
marked by erosion levels at the base and the top (p. 84). The Subzone is not known
to be represented elsewhere in France at this time. The base of the Subzone is
marked by the general adoption of a channelled venter by the various species of the
genus Euhoplites. The ammonite fauna of the Subzone has been discussed by Owen
(1958 ; 154) and Hancock (1965 ; 246) and there is nothing further to add except that
the top of the Subzone is now drawn at Folkestone at the level within Bed VII
immediately below the first appearance of Anahoplites daviesi in the sequence.

(i) Subzone of Anahoplites daviesi

This Subzone recognised by Spath (1925b ; 35, 1926a ; 153-4) is characterised by
Anahoplites of the daviesi group. There is virtually no difference in the accompany-
ing ammonite fauna in England, but these very characteristic species of Anahoplites
have a wide geographical range in this time span, and on this point alone the Subzone
should remain separate from that of FE. mitidus. Breistroffer’s grouping (1965 ; table
opp. 312) is not acceptable, and the British reading should be adhered to (e.g. Owen
1958 and Hancock 1965 ; 245, 246).

The Subzone is best developed at Folkestone although it is present in uncondensed
sediments elsewhere in Kent and Sussex (Ringmer). However, at Folkestone there
are clays representing horizons higher than any known elsewhere outside the U.S.S.R.
Even at Folkestone the top of the clays of Bed VII representing this Subzone are
planed-off and the basal nodule bed of Bed VIII contains material of late daviesi
Subzone age. At Wissant, the lower part of the cristatum Subzone (the partial
equivalent of Bed VIII (i) at Folkestone) is represented by clays. Unfortunately these
rest non-sequentially upon sediments of mitidus Subzone age (p. 85) and neither at
Wissant nor elsewhere in France at this time have sediments of daviest Subzone age
been proved. The development of the Subzone in Russia is mentioned later (p. 132).

The upper limit of the Subzone can, however, be determined by reference to Bed 12
(v) at Wissant which contains no Anahoplites of daviesi type but in which Dipoloceras
bouchardianum and Beudanticeras beudanti make their appearance marking the base
of the cristatwm Subzone. In this bed also there occurs the morphological transition
from Inoceramus concentricus to the shell form I. sulcatus (p. 85).

126 MIDDLE ALBIAN STRATIGRAPHY

(iu) THE POSITION OF THE SUBZONE OF DIPOLOCERAS
CRISTATUM

The junction of the Lower with the Upper Gault at Folkestone is marked by two
seams of phosphatic nodules separated by a few inches of clay (p. 15). This junction
bed (Bed IV of De Rance 1868, and Bed VIII of Price 1879, 1880, Jukes-Browne 1900
and subsequent workers) has always been included in the Lower Gault. However, it
has long been recognised that the fauna is a transitional one containing elements
characteristic of the beds below and above. The bed formed the zone of Ammonites
beudantii of De Rance, and that of Ammonites cristatus of Price. Jukes-Browne
(1900 ; 45) did not include the time span represented by Bed VIII in his zone of
Ammonites lautus. Spath, however, placed his cristatwm ‘zone’ in the Middle
Albian (1923a, b) and later relegated it to subzonal rank and included it in the lautus
Zone in his sense (1926b ; 425). This classification has been followed by later Eng-
lish workers (e.g. in Hancock 1965 ; 245, 246). Breistroffer (1947 ; 48, 68) included
the Subzone in the Upper Albian where it stood in isolation. The author stated that
more research would have to be carried out before a final decision could be made on
the position of the cristatum Subzone (1958 ; 164). However, in the meantime,
Breistroffer’s recommendation (see also 1965 ; table) has been accepted by other
workers such as Collignon (1963 ; 2) for the sequence in the Malagasy Republic, and
Young (1966 ; 15) for the succession in Texas, and in England by Melville (in Smart
et al., 1964 ; 7).

Although Bed VIII contains a fauna which on balance links it with the Upper
Albian, its lower nodule bed includes an important Middle Albian element derived
from sediments of daviest Subzone age. One of the principle objections to placing
the cvistatum Subzone in the Upper Albian is that hitherto it has meant placing the
Middle-Upper Albian junction at a level of erosion. However, it is now known that
the basal part of the cristatum Subzone is represented in an uncondensed sequence in
Bed 12 (v) near Wissant. This fact removes any objection that the writer might
formerly have held against placing the Subzone in the Upper Albian, and I now
recommend that it be included in the Upper Albian to form the basal Subzone of the
inflatum Zone.

In Bed 12 (v) at Wissant we can see the incoming of a basal Upper Albian fauna in
an uncondensed sequence (p. 85). The change in Inoceramus from a concentricus to a
sulcatus form has already been mentioned. Beudanticeras beudanti appears together
with D. bouchardianum. These forms are all known from Bed VIII (i) at Folkestone.
Bed 13 at Wissant contains en melée material derived from the equivalent of Bed VIII
(ii) & (iii) at Folkestone together with the lower part of Bed IX up to and including
the horizon of Ewhoplites inornatus. In truth Bed 13 at Wissant forms the base of
the Upper Gault in our sense as well as that of our French colleagues. However, they
consider it to be the direct equivalent of Bed VIII, whereas it is in fact the product
of one period of erosion which occurred at Wissant later in the crvistatwm Subzone,
while at Folkestone there were essentially two phases of erosion at earlier dates
within this Subzone. It has been, up to now, the view of our French colleagues that

IN THE ANGLO-PARIS BASIN 127

all the Gault below Bed 13 of Destombes was of Middle Albian age, but it must now be
recognised that Bed 12 (v) below is in fact of basal cristatum Subzone age.

The fauna of the cristatum Subzone on balance, bearing in mind that a late daviesi
Subzone element is present in Bed VIII (i), has its most important faunal link with
the beds above rather than those beneath. The very characteristic lower Upper
Albian bivalve form I. sulcatus develops in the basal part of the cvistatwm Subzone and
ranges up to the top of the orbignyt Subzone. Towards the top of the Middle Albian
daviest Subzone, extreme forms of Anahoplites of the daviest group occur which are
close to, but still generically distinct from Epihoplites (including Metaclavites). At
the same point in time species of Ewhoplites and Dimorphoplites modify towards those
of the cristatum Subzone. The hoplitinids are almost completely dominant, the only
Mojsisovicsiinid being Dipoloceras cornutum which is very rare. In the cristatum
Subzone above we see the introduction of an important non-hoplitinid element in the
ammonite fauna. In fact many of the subfamilies last well-represented in the Anglo-
Paris basin in lyelli Subzone times appear once again in consort, and as before are
subordinate to the hoplitinids.

Euhoplites is well represented but the species are essentially different to those of
the Middle Albian below, and are, moreover, more closely linked to those of the
orbignyt Subzone. This genus survives until the awritus Subzone and it is interesting
to find that a largely unfigured varicoswm Subzone fauna in the Leighton Buzzard
area contains forms which are morphologically closer to those species of Bed V—VII
at Folkestone than to those of Bed VIII. Dimorphoplites in the strict sense does not
survive the cristatwm Subzone, its nitch being filled by Epihoplites (including Meta-
clavites Casey). Itis quite possible that the ecologic factors which caused Inoceramus
concentricus to develop the strengthened shell from of J. sulcatus, also brought about
changes in the morphology of the endemic hoplitinids during the clearly unsettled
physical conditions in the cristatwm Subzone sea.

The non-hoplitinid element in the ammonite fauna is subordinate but highly
characteristic of this Subzone and foreshadows the major development of the branco-
ceratinid and mortoniceratinid ammonites which occurs at the base of the orbignyi
Subzone in the sense used here. Hysteroceras is represented by H. pseudocornutum,
H. capricornu, and H. simplicicosta, forms which are transitional from the earlier
Eubrancoceras. An early mutation of H. orbignyz does in fact occur in the cristatum
Subzone, but the only species of Mortoniceras known is M. rigidum although the
generic position given it by Spath is uncertain. Dzpoloceras itself, the characteristic
genus of this time span, probably gave rise to Mortoniceras but the picture is not
clear at this time. The Lyelliceratidae is represented by Neophlycticeras which also
ranges into the orbignyi Subzone but it is rare. Beudanticeras represented in the
cristatum Subzone by the type-species B. beudanti and forms such as B. subparandieri,
also continued on into the orbignyi and higher Upper Albian Subzones.

The cristatum Subzone must also include the time span represented within the
sediments of the basal part of Bed IX at Folkestone up to the level at which Hyster-
oceras and Mortoniceras characteristic of the orbigny: Subzone suddenly became
dominant. This just excludes the horizon of Euhoplites inornatus which is of wide-
spread occurrence in England and forms a good marker for the base of the orbignyi

128 MIDDLE ALBIAN STRATIGRAPHY

Subzone. The cristatwm Subzone as here defined, therefore, includes sediments
grouped with the Upper Gault.

As Breistroffer has pointed out (1947 ; 48-50), Dipoloceras cristatum and the other
contemporary species of this genus are of widespread occurrence. They are known
in sequences as far removed from each other as the Anglo-Paris basin, the Malagasy
Republic and Zululand, Russia and Texas, that is, in more than one ammonite
faunal province. The well-marked period of erosion of Middle Albian sediments in
the Anglo-Paris basin which occurred during the crvistatum Subzone, is just as well
marked in the Malagasy Republic (Besairie & Collignon 1956). Although not
specifically stated by Young (1966) there are signs of a similar break in Texas. It is
also possible that there is a break in the sequence in Peru (Benavides-Caceras 1956).
In an entirely different faunal province still, the occurrence of Gastroplites cantianus
in Bed VIII at Folkestone and in the lower part of the Gastroplites Zone in Canada is
extremely important (e.g. Jeletsky 1964 ; Table 1, 1968). In Canada there is a
definite break in the ammonite sequence although apparently not in the sedimentary
sequence (Jeletsky 7m litt.), for the mcconnela Zone which contains genera such as
Arcthoplites, Cymahoplites and Cleoniceras, both known from the mammuillatum Zone
of the old world (Casey 1961c ; 167) is definitely of Lower Albian age. A similar
ammonite faunal gap involving the whole of the Middle Albian appears to exist in
Alaska (Imlay 1960 : 1961) and California. Imlay’s Cleoniceras (Grycia) presents no
difficulty here because it does not possess umbilical bullae and almost certainly
belongs to Beudanticeratinae. In effect, by taking the base of the Upper Albian to
coincide with the base of the cvistatuwm Subzone and its provincial equivalents, it is
possible to arrive at a common point of division between the two substages capable of
recognition in the local successions of each country.

(iv) THE ZONAL GROUPING (Table 1, p. 10)

(a) The Zone of Hoplites (H.) dentatus

As defined here this Zone comprises the Subzones of H. (I.) eodentatus, L. lyelli and
H. (H.) spathi. This represents virtually the total range of the closely related
morphological group of Hoplites represented by the index species H. (H.) dentatus.
In the eodentatus Subzone, this species is not typically developed, but in fact H. (J.)
eodentatus is a direct transition between the earlier Psewdosonneratia and H. (H.) of
the dentatus group. This group dies out at the top of the spathi Subzone, although
Hoplites (H.) continues onwards into the loricatus Zone above. With some modifi-
cation, this is almost a return to the old concept of the interruptus Zone in Barrois’
and Jukes-Browne’s sense.

The Zone is geographically widespread represented in sediments both condensed
and uncondensed. The hoplitinid faunal province by the spathi Subzone can be
shown to have extended from the western border of asiatic Russia, south to the
northern margin of Tethys now represented in the Caucasus mountains. The
boundary then runs along the northern side of Tethys westwards to France. The
land area flanking the east side of the Atlantic rift system which existed at this
time (cf. Carey 1958) apparently formed the western boundary, although the situation

IN THE ANGLO-PARIS BASIN 129

in Greenland is not yet clear. Ina similar manner the primitive Arctic Ocean may
have formed the northern boundary, for Hoplites is not yet known from Canada,
Alaska or Japan. The province, therefore, consists of the shelf seas of Europe. In
the lyelli Subzone especially, there are important links with adjoining ammonite
faunal provinces in which Hoplites (H.) is as yet unknown. These links are discussed
in greater detail below.

(b) The Zone of Euhoplites loricatus

This Zone comprises the Subzones of A. intermedius, D. niobe, M. subdelaruei and
E. meandrinus. It is almost the total range of E. lovicatus which is typical of the
group of Ewhoplites with sulcate rather than channelled venters seen late in the Jautus
Zone. Deposits of this Zone, with the exception of the basal part of the intermedius
Subzone, are of very limited occurrence in the European province. This is probably
due to erosion in the early part of the Upper Albian which apparently produced a
fairly general hiatus involving the lovicatus and lautus Zones throughout much of
Europe except in the deeper basins. It could be argued that there is no point in
dividing Jukes-Browne’s original /autws Zone into two parts, but the morphological
change in Ewhoplites is quite striking at the level at which the writer has drawn the
base of the Jautus Zone. Although the change in the remainder of the ammonite
fauna is not quite so clear cut, nonetheless it does occur at about the same point in
time. The ammonite fauna of the lovicatus Zone is grosso modo sufficiently distinct
for it to be capable of definite recognition at zonal rank. Spath placed the division
between the dentatus and Jautus Zones in his sense at the top of the niobe Subzone
where in fact there is no significant change in the ammonite fauna. This quite
arbitrary and meaningless arrangement is firmly rejected here.

The intermedius Subzone has as great a geographical range as the spatht Subzone
of the dentatus Zone but known sediments of the niobe, subdelaruet and meandrinus
Subzones are preserved in only a very limited area, that of eastern England and
northern France. Links with other ammonite faunal provinces are very few and
mainly with the tethyan belt.

(c) The Zone of Euhoplites lautus

The Jautus Zone as now defined consists of two Subzones ; that of Euhoplites nitidus
and that of Anahoplites daviesi. It is essentially the total range of Euhoplites lautus,
and the contemporary species of this genus with their well-marked clean cut ventral
channel. The base of the Zone as indicated above is defined by the quite sudden
change in the peripheral aspect of Ewhoplites. The top is defined by the appearance
of Dipoloceras bouchardianum and Beudanticeras beudanti indicating the base of the
cristatum Subzone.

Deposits of the nitidus Subzone are of very limited known geographical extent.
They are known from eastern England and northern France, and apparently occur
also in Poland, but these are probably only remnants which survived the basal Upper
Albian erosional movements. The daviest Subzone can definitely be identified in
Russia (Mangyshlak Peninsula) but this is the only area outside the eastern Weald of
England that deposits of this Subzone have as yet been recognised. Russian

I

130 MIDDLE ALBIAN STRATIGRAPHY

workers (e.g. Glasunova 1953a ; 18) have recognised above the dentatus Zone, which
in effect is equivalent only to the spatht Subzone, a Subzone of Anahoplites asiaticus.
It is apparent that this is, in part, of entermedius (e.g. Kopet Dagh, Mangyshlak), and
in part of daviest Subzone age (e.g. Mangyshlak). Anahoplites asiaticus and A.
tvanscaspius Glasunova (1953a) look like early intermedius Subzone Anahoplites com-
parable to, although perhaps not specifically identical with species known in the
uppermost spathi and basal intermedius Subzones of England and France. A some-
what different scheme has been proposed by Sokolov (1966) and this is discussed
below (p. 132).

V. LINKS WITH OTHER FAUNAL PROVINCES

The following is a brief review of the known links between the hoplitinid and other
faunal provinces at certain levels in the Middle Albian. A vast amount of work still
remains to be done to determine the degree of representation of Middle Albian
sediments throughout the whole surface of the Earth and so to determine accurately
the boundaries of the various faunal provinces. It is becoming apparent, however,
that sediments of Middle Albian age are far more restricted in occurrence than those
of the Upper Albian, and in many areas where they both occur, there is often evidence
of a hiatus between them. Basal Upper Albian sedimentation in widely scattered
places on the Earth commenced after a period of erosion (e.g. Europe, and the
Malagasy Republic), or they transgress onto very much older surfaces (e.g. Africa).
The fauna itself shows evidence of unstable conditions at the base of the Upper
Albian (p.127). Whether Middle Albian sediments were deposited over a widespread
area of the Earth only to be removed largely by subsequent erosion is a matter of
conjecture. Here, however, we are concerned only with an outline sketch of the
boundaries of our province and at what definite levels it is possible to correlate with
adjoining provinces.

A. The boundaries of the hoplitinid province (text-fig. 50)

The westward boundary of the hoplitinid province during the Middle Albian was
formed by the massifs of Morvan and Armorica. These separate the European shelf-
sea from the narrow sea-way connected with the Tethyan belt which was then the
‘Atlantic’ (Carey 1958 ; Bullard et al., 1965). No species of Hoplites, Euhoplites,
Anahoplites or Dimorphoplites have yet been found in the United States of America
and Canada. However, there are tantalising references to a typical hoplitinid fauna
in argillaceous sediments in the coastal area N. of Scoresby Sound, E. Greenland
(Spath 1946 : Donovan 1949, 1953) which demonstrate that this area also is to be
included in the province. If E. Greenland is brought back to its apparent middle
Cretaceous position, the coast N. of Scoresby Sound is approximately opposite the
northern outlet of the present day North Sea basin.

At the present time no information is available about Middle Albian sediments in
the North Sea Basin, but the evidence from the eastern margin of England and from
deep borings in Holland suggest that sediments of Middle Albian age thicken towards
at least the south central part of the Basin. The Middle Albian sediments of Ger-

PARIS BASIN

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132 MIDDLE ALBIAN STRATIGRAPHY

many belong to the hoplitinid province and these extend through Poland (e.g.
Cieslinski 1959) into Russia, although they are largely concealed by later rocks. In
this respect it is interesting to note that Ravn (1925) has figured ammonites from
the Cenomanian basal conglomerate on the Baltic Island of Bornholm. These indi-
cate not only that elements derived from the tardefurcata and mammillatum Zones are
present en melée with Cenomanian ammonites but also his Sonneratia Baylei (pl. III,
fig. 6a, b) looks like an Anahoplites suggesting the possibility of Middle Albian
material also being represented in the deposit. This paper seems to have escaped
the attention of previous English workers (see also @dum 1928 ; 44-5), and throws
light on the provenance of the glacial erratics described by Skeat & Madsen (1808)
from the Jutland drift.

The hoplitinid province in the Middle Albian may not include all of Russia in
Europe, but it certainly extends as far east as the southern Urals and the eastern
border of the Caspian Sea south to the Kopet Dagh in the border region with Iran.
The area of Daghestan flanking the western side of the Caspian, and the area between
the Mangyschlak Peninsula and the Kopet Dagh is of considerable interest. From
the ammonites figured by Semenov (1899), Sinzov (1909 : I9I5) and Glazunova
(1953a, b) together with the recent stratigraphical work carried out by Sokolov (1966)
it is apparent that Middle Albian sediments are well developed in this area. Sedi-
ments of lyelli Subzone age appear to be absent, but there is a good fauna of Hoplites
(H.) spp. indicating the spathi Subzone (the dentatus Zone of Sokolov). The inter-
medius Subzone is also definitely represented including the basal part with Ana-
hoplites of evolutus type. The next horizon which can definitely be correlated with
the sequence in the Anglo-Paris basin are Sokolov’s zones of ‘ Anahoplites ’ rossicus
(Sinzov) and ‘ A.’ whligi (Semenov). These forms can be matched in the cristatum
Subzone of Kent, and are descended directly from Anahoplites of the daviesi group,
which together with its ‘variety’ orvnata, are also known from this region of the
U.S.S.R. Spath (1943 ; 732) was quite incorrect in comparing ‘ A.’ uhligi with
A. daviest ; the two are quite distinct although closely related.

Between the Zone of ‘ Anahoplites’ rossicus and that of A. intermedius Sokolov
recognises in ascending order a Zone of Daghestanites daghestanensis Glasunova and a
Zone of Anahoplites kelendensts sp. nov. (of Sokolov which appears to be undescribed).
Daghestamtes daghestanensis Glasunova has not yet been recognised outside the
Soviet Union, and Anahoplites kelendensis in the absence of figures cannot be com-
pared. Undoubtedly the most striking feature of the Middle Albian sequence in this
area is the apparent total absence of Euhoplites so prolific in the area of Western
Europe and present also in Greenland. In Siberia the Albian is represented in a
continental facies.

The spatht and intermedius Subzones are certainly represented on the northern
margin of the tethyan belt in northern Bulgaria (Zakharieva-Kovatcheva 1957,
Nikolov 1965, 1970). Seitz (1930) has figured a definite spathi Subzone fauna on the
same margin in the Rhaetic Alps at Vorarlberg, Austria. From there westward the
southern boundary of the province follows the margin through Switzerland and into
France. The Albian ammonite fauna of the tethyan belt is quite distinct in char-
acter from that of the European shelf seas and the hoplitinids are absent from Albian

IN THE ANGLO-PARIS BASIN 133

sequences south of it. The alleged occurrence of Hoplites (H.) in Mexico has not yet
been substantiated by illustrations.

B. Links with the sequences of other countries

There are two rather fortuitous major links between the hoplitinid faunal province
and other faunal provinces. The first of these comes in the lyelli Subzone near the
base of the Middle Albian, the other in the cvistatum Subzone at the base of the Upper
Albian. Between the two there are very few links known.

(i) WEST PAKISTAN

Spath (1930b) described an Albian fauna from the area of Hazara, now Abbotabad,
in W. Pakistan. The ammonites come from a condensed deposit which yielded
possibly mammullatum Zone, and definitely lyelli Subzone fossils. The lyelli Subzone
is indicated by the occurrence of Lyelliceras lyelli, L. cotteri, species of Oxytropidoceras
(sensu lato) and Brancoceras. However, no species of Hoplites (H.) have been re-
corded. No direct comparison can yet be made with the sequence in the U.S.S.R.
where this Subzone has not yet been detected, and in fact in the hoplitinid province
the Subzone is not represented E. of the western area of Switzerland. No other
Middle Albian links are known from Pakistan, although the cristatum Subzone is
definitely represented. The two specimens of Oxytropidoceras figured by Spath
(1934b ; 18-21, 30, pl. vi, figs. 1, 2) from the Attock district are of Middle Albian age,
but cannot be assigned to a subzone at this time.

(ii) TETHYAN BELT

As yet very little knowledge exists on the degree of Zonal and Subzonal repre-
sentation in the Middle Albian sediments of the part of the tethyan belt stretching
westwards from W. Pakistan through Iran and Asia Minor, the Mediterranean
countries to reach the proto-Atlantic on the W. coast of Spain. It is clear that the
phylloceratids, some lytoceratids, and desmoceratids, where they occur, are too long
time-ranging to be of even zonal value. Desmoceras latidorsatum, for example, occurs
in the mammullatum Zone, as well as in the lyelli and intermedius Subzones in the
Middle Albian hoplitinid province. From the discussion of the American sequences
it seems that Oxytropidoceras sensu lato may also include long time-ranging species.
However, the mojsisovicsiinids, and in particular the engonoceratids may eventually
aid correlation with the provinces to the north and south of Tethys. In this respect
the distribution of Platiknemiceras is of significance (Casey 1961b). The strati-
graphic range of this genus as indicated by Casey can in two instances be narrowed.
It is associated with an example of Lyelliceras gevreyi at Hamiran, Iran (BMNH.,
C 68410) cited by Spath (1931 ; 315) and recorded as Prolyelliceras by Casey (1961b ;
354). This specimen shows the extra-intercalated siphonal crenules and tendency to
en-echelon ventro-lateral crenules characteristic of forms which occur at the base of
the lyelli Subzone. L. flandrini crenulata discussed below is of the same age and is
also associated with Platyknemiceras.

134 MIDDLE ALBIAN STRATIGRAPHY

(iii) ALGERIA

Although it is in need of revision, the work of Dubourdieu (1953 : 1956 ; 185-228)
indicates that at least the equivalent of the lower part of the /yelli Subzone is well
represented by sediments in the Monts du Mellégue, Djebel Ouenza, Djebel Def, the
environs of the Djebel Hameima, and the Djebel Bou Khadra. The ammonites
include Lyelliceras flandrini Dubourdieu and L. radenaci (Pervinquiére). L.flandrini
has extra-intercalated siphonal crenules indicating the lower part of the /yelli Subzone,
and this is confirmed by L. flandrini crenulata (1953 ; pl. III, figs. 25-35) which is
closely comparable to the inner whorls of the grade reached by L. gevrey (Jacob) in
Bed 1 at Courcelles (Aube). JL. vadenaci occurs also in the lyell1 Subzone of the
Guilford Colliery, Kent (p. 76).

(iv) SOMALIA

Tavani (1949) has described and figured what appears to be a lyelliceratid under
the name Somalites vertebralis. This is associated with Brancoceras and they may
well indicate the equivalent of the /yelli Subzone. However, Somalites has not been
found outside its type locality.

(v) MADAGASCAR

Collignon has demonstrated recently that the lyelli Subzone is represented without
question in the Middle Albian sediments of the Malagasy Republic (1963, 1965a) and
he employs the term Zone a Lyelliceras lyelli. Lvyelliceras lyelli and close relatives
were found by him at Khomihevitra (1963 ; pl. 315, figs. 1333-5). It should also
be noted that the fauna of brancoceratids described by him (1949) from d’Ambara-
maninga could well indicate a basal Middle Albian age for the sediments which con-
tain them. The species of Pseudosonneratia from the same bed certainly do not
belong to that genus. Dzpoloceras cristatum, and contemporary species of this genus,
marking the unconformable base of the Upper Albian sediments, occur at Andrano-
fotsy (e.g. Collignon 1963 ; 2). At present it is not possible to correlate the inter-
vening Middle Albian sediments, referred to a Zone a Oxytropidoceras acutocarinatum
and Manuaniceras jacobt in the Malagasy Republic, with those of Europe or America.

(vi) SOUTH AMERICA

Ammonites of Middle Albian age have been well illustrated from Colombia and in
particular Peru, and for the purpose of the present comparison these only will be
considered. Important Middle Albian faunas are also known from other South
American countries such as Venezuela and Brazil but these have not yet been figured
in full.

(a) Colombia

In Colombia, Gerhardt (1897 ; 168-170, pl. IV, figs. 8a, b) described an ammonite
which he named Acanthoceras prorsocurvatum from Ubaque (Cundinamarca) and

IN THE ANGLO-PARIS BASIN 135

considered it to be of Aptian age. Douvillé (1906) recognised it to be of Albian age
and figured a fragment (1906 ; pl. II, figs. 1, 1a) which Spath (1930b ; 65 footnote)
later renamed Prolyelliceras peruvianum, the type species of his ‘genus’. Riedel
(1934 ; pl. 9, figs. 9-11) figured another specimen as Prolyelliceras? lobatum confirm-
ing a Middle Albian age for these forms, because they are associated with Lyelliceras
of the lyelli type also figured by him. From specimens in the British Museum (Nat.
Hist.) e.g. BMNH C 74786 J. V. Harrison Colln., from the road between Viola and
Portillo, it is apparent that these so-called ‘ Prolyelliceras ’ are in fact large specimens
of Lyelliceras of lyelli-ulrichi type, the ornament of which modifies at diameters which
are as yet unknown in the old world. Pvolyelliceras is considered here to be a junior
subjective synonym of Lyelliceras, and the lyelli Subzone is apparently well repre-
sented in Colombia.

(b) Peru

In the last twenty years our knowledge of the Middle Albian zonal stratigraphy of
Peru has been greatly increased by the work of Knetchel (in Knetchel et al., 1947)
and Benavides Caceras (1956). It is apparent that the lyelli Subzone is well repre-
sented in the Pariatambo, Crisnejas and Chulec formations classified by Benavides
Caceras with the Zone of Oxytropidoceras carbonarium. These sediments, between
100 and 200 metres thick, have yielded ammonites such as Desmoceras latidorsatum,
Lyelliceras lyell, L. ulricht and Eubrancoceras aegoceratoides, all known from localities
in the Anglo-Paris basin, especially at Courcelles (Aube). It is interesting to see that
Benavides-Caceras records Lyelliceras pseudolyeli from Bed 20 of the Crisnejas
locality classified by him with the underlying Knemiceras raimondii Zone. It is also
instructive to compare the sequence of Zones recognised in Peru with that of Algeria.
As yet Mojstsovicsia has not been found in the lyelli Subzone in the Anglo-Paris
Basin but it does occur in the overlying spathi Subzone where it is represented by
M. delaruei and subspecies. The genus is represented in Peru by the type species of
Mojstsovicsia, M. ventallinensis (Gabb) whose exact Subzonal age is uncertain
(Douglas 1921). It is also uncertain at present whether there are any higher Middle
Albian sediments than those of the lyelli Subzone preserved in Peru, but it appears
that the Subzonal sequence is incomplete reflecting the break in sedimentation seen
elsewhere in the World.

(vii) TEXAS

Young (1966) has presented an extremely important work on the ammonite
zonation of the Fredericksburg Division of the Texas Albian and on the mojsisovic-
siinid ammonites contained in it. He presents a correlation of the Texas zonal
sequence with that of Europe. Following a discussion of the faunal sequence both
in Texas and elsewhere outside the United States, he examines four possible con-
clusions raised because of an apparent anomaly which exists between the carbon-
avium Zone of Peru and that of Texas. In Peru the carbonariwm Zone of Benavides
Caceras contains Lyelliceras and it is here correlated with the lyelli Subzone of the

136 MIDDLE ALBIAN STRATIGRAPHY

European Subzonal sequence. In Texas, Young’s carbonarium Zone does not con-
tain Lyelliceras and the possible conclusions that he poses are as follows (1966 ; 18).

‘

1. We do not understand yet the taxonomy and ranges of species of
Lyelliceras. After all, the entire family Lyelliceratidae is still little known.

2. The rocks between the dentatus and mitidus subzones are greatly con-
densed in Peru, Venezuela, and Colombia.

3. The beds in Texas without diagnostic ammonites (Upper Glen Rose and
lower Walnut Formations) are to be correlated with the dentatus and benettianus
subzones, and the zones of salasi and carbonarium are condensed and represent
all of the Folkestone section between the dentatus and cristatum subzones
(= beds II-VII, inclusive). ‘‘ 0.” carbonarium 1s interpreted with such latitude
that it could include forms of the salasi zone of Texas.

4. The ammonite zones are migrating against each other. In other words,
Lyelliceras is younger in South America than in Europe, and Manuaniceras
carbonarium is younger in Texas than in South America. This possibility will
be distasteful to orthochronologists, but it is a factor that cannot be overlooked
and is supported by the occurrence of Inoceramus concentricus with Lyelliceras
(Benavides, 1956, p. 414).

I am inclined to look with favor on the first explanation, but it is a personal
preference. In the final analyses, some combination of two or more of these
explanations may seem preferable. I must point out, however, that Benavides’
(1956) section descriptions do not indicate condensation.’

Unfortunately it is necessary to point out that the evidence for his correlation of
the Texas Middle Albian zonal sequence with that of the Anglo-Paris basin is non-
existent. The only direct correlation that can be made between the two areas is in
the cristatum Subzone. Dzipoloceras fredericksburgense and D. cristatum are known
from the upper part of the Goodland Limestone in Tarrant Co. Texas. Inoceramus
subsulcatus was figured by Bose (1927; 189-193, pl. XVIII, figs. 1-5) from the
Edwards Limestone which is of the same age. But perhaps the most significant
link is provided by Oxytropidoceras cantianum Spath (1931 ; 350-1, pl. 32, fig. 5)
which is almost identical to the specimen of ‘ Manuaniceras carbonarium transitional
to M. peruvianum multifidum (Steinmann) ’, figured by Young (1966 ; pl. 17, fig. 6)
and which is also from the upper part of the Goodland Limestone (= cristatwm Sub-
zone). The only other possible link is provided by Difoloceras of the cornutum group
also known from the Goodland Limestone (e.g. Spath 1931 ; 363 text-fig. 118).
Now we have the measure of the problem. Oxytvopidoceras in the wide sense occurs
in the Anglo-Paris basin in the mammullatum Zone (Lower Albian), in the lyelli and
spathi Subzones (Middle Albian), and in the cvistatwm Subzone (Upper Albian) ; it is,
therefore, a very long-ranging group. The development of the genus Lyelliceras is
now well known both in the hoplitinid province and outside it, and the evidence
indicates that the beds containing it in Pakistan, Madagascar, Algeria, Colombia,
and Peru are of the same age as those containing Lyelliceras in the Anglo-Paris basin.
Therefore, the specimens of ‘ Oxytropidoceras’ carbonarium from the lyell1 Subzone
sediments of the Pariatambo Formation of Peru cannot be of the same age as those

1 It is now known from the Loricatus Zone of north Kent (Owen in press).

———

IN THE ANGLO-PARIS BASIN S77

occurring in the Goodland Limestone of Texas where they are associated with
Dipoloceras of the cristatwm Subzone. It now becomes apparent that certain species
of the group typified by Oxytropidoceras, like other tethyan genera, may be very
long time-ranged. Unlike the hoplitinids which show a great deal of morphological
differentiation, Oxytropidoceras and its closely allied genera show comparatively
little such differentiation. On their own, it might prove very difficult, except in a
few cases, to use them for the fine subdivision of which the hoplitinids have in
particular proved capable.

As yet no attempt has been made to compare the species of Oxytropidoceras (s.lat.)
which are known from the lyelli and spatht Subzones of the Anglo-Paris basin with
those occurring outside the hoplitinid province. It might well prove possible
eventually to correlate the spatht Subzone with the tethyan succession (including
Texas) on the basis of certain species of Oxytropidoceras (s.lat.). At the moment the
intermedius and mobe Subzones have no exact zonal links with Tethys although long
ranged tethyan forms are known from both time spans. Mojsisovicsia in the sub-
delavuet Subzone should be a renewed tethyan incursion, but none of the species from
this Subzone or the lautus Zone have yet been recognised elsewhere. So, at this time
it is not possible to correlate the Zones recognised in Texas with those of the Anglo-
Paris basin. It is however, apparent from Texas and Madagascar that sediments
characterised by the group typified by Oxytropidoceras occur between the /yelli and
cristatum Subzones. Although sedimentary breaks can be difficult to detect in
carbonate-marl sequences it seems to the writer that Young’s salasi and carbonarium
Zones are high Middle Albian as he states.

(vii) CANADA

The zonal scheme of the Canadian Albian has recently been ably reviewed by
Jeletzky (1968), but although this represents a refinement on his earlier review of
1964 it is still open to criticism. In 1964 Jeletzky indicated that the Zone of
Arcthoplites mcconnelli contained Cleoniceras and Cymahoplites. If these ammonites
are correctly assigned generically then together they indicate a Lower Albian, mam-
millatum Zone age in the sense of Casey (1961a ; Table 1) and herein. Cymahoplites
occurs in the mammillatum Zone of Europe (Casey 1961c ; 165-169, Pl. XXIX, figs.
ta-d), and although Cleoniceras (C.) ranges from the tardefurcata Zone (regularis
Subzone) to the basal Middle Albian eodentatus Subzone, the weight of evidence, from
the associated ammonites indicates a mammillatum Zone age. There is no evidence
at this time to indicate that the Subzones of Lemuroceras irenense and L. mcconnelli
are the correlatives of the European Subzone of Douvilleiceras inaequinodum (Jeletzky
1968 ; Fig. 1) now the basal Middle Albian Subzone of Hoplites (Isohoplites) eodentatus.

The Gastroplites Zone has been equated by Jeletzky (1968 ; Fig. 1) with both the
daviesit Subzone (Middle Albian) and the cristatum Subzone (Upper Albian). How-
ever, G. cantianus Spath which occurs in the Gastroplites Zone in Canada occurs also
in Bed VIII at Folkestone its type locality. It is undoubtedly of cristatwm Subzone
age, and there is no evidence for correlating the daviest Subzone with any part of the
Gastroplites Zone at this time.

138 MIDDLE ALBIAN STRATIGRAPHY

Between the Gastroplites Zone marking the base of the Upper Albian and the
meconnelli Subzone is a thick interval of shales which have not yet yielded ammonites.
Jeletzky has classified these sediments with a Zone F which he considers to include
part or all of the time span between the mammillatum ‘Subzone’ and the cristatum
Subzone (1968 ; 17-18, Fig. 1). If there are any Middle Albian sediments in Canada
then they are contained in these shales of Zone F. At the moment, however, no
Middle Albian ammonites are known from Canada. It is worth noting that the
alleged Cleoniceras associated with a gastroplitinid ammonite fauna in Alaska (Imlay
1961) does not stand up after examination. Cleoniceras (Grycia) sablei Imlay which
is known from crushed material does not appear to possess umbilical bullae and is
here considered to be a member of Beudanticeratinae. The gastroplitinid Upper
Albian province included the area that is now the western cordillera of North
America stretching from Alaska to California.

(ix) GREENLAND

Our knowledge of the Albian sediments of the area between Traill O. and the
Wollaston Foreland on the E. coast of Greenland N. of Scoresby Sound, is due to the
work of Spath (e.g. 1946 ; 8-10) and Donovan (1949 ; 6-7 : 1953, 35-37, 50-51). It
is tantalising in its incompleteness for it seems that in this area we have the boundary
between the European Albian province and that of the area of Canada and the western
cordillera of N. America. Both Spath and Donovan record Lower Albian ammonites
which occur in both provinces ; Middle Albian ammonites such as Hoplites and
Euhoplites which are characteristic of the European hoplitinid province ; and Upper
Albian ammonites which may also link the two provinces.

(x) CONCLUSION

The foregoing very brief review indicates that a considerable amount of work now
requires to be done on Middle Albian sequences outside the Anglo-Paris Basin along
the lines attempted here. The object of this review is to stimulate such research.
Superficially, it seems that there is in many parts of the Earth a major break in
sedimentation, particularly at the top of the Middle Albian. General sedimentation
occurred once again in early Upper Albian times. There is evidence of this even in
the area of Kent and Sussex, wherein the Jovicatus and lautus Zones are apparently
the most completely represented by sediments. Breaks of such an extensive nature
in the Cretaceous, must be due to major events in the Earth’s crustal development,
for there is no evidence of unusual climatic conditions of sufficient magnitude to
reduce sea-level by a significant amount.

There has always been a tendency to equate movements on a regional scale in
Europe with various periods of deformation in the Tethyan belt. Until recently the
development of the Atlantic, Arctic and Pacific ocean basins has been largely ignored
and yet with recent geophysical work we are now beginning to see the important
effect that initial faulting must have had even on Cretaceous sedimentation. The

—————————

IN THE ANGLO-PARIS BASIN 139

Royal Society’s Symposium on Continental Drift posed many new questions for the
stratigrapher and challenged many long held concepts. The important reviews by
Wilson (1965 ; 228-251), and Maack (1969) on the formation of the ocean basins drew
particular attention to the role and effect of initial major rift, block and transcurrent
faulting in the Jurassic and Cretaceous before the pulling apart of the old continents
to the positions of the new within the Tertiary. The evidence of faulting and igneous
activity indicate that the pulling apart of the continental masses bordering the
present day Atlantic commenced earlier in the southern Atlantic (e.g. Maack 1969).
In the northern Atlantic the vulcanism commenced in the Tertiary.

The major faulting must at times have had a profound effect on the distribution
and depth of water, and thus the depositional and erosional conditions, in the
continental shelf seas. This would produce characteristics quite distinct from the
pressure of the African continent against Europe which in the Tertiary culminated
in the Alpine ‘storm’. During the Albian there is good evidence of the effect of
this faulting, associated with the separation of Africa and South America, in and
around Africa (e.g. Furon 1963). Even in southern England faulting occurred
during cristatum Subzone times associated with marked erosion of Middle Albian
sediments. This is but one symptom of one short period of crustal instability which
is apparent in both the sediments and fauna in a number of areas in the World and
points to a significant event in the development of the Earth.

The old idea of a Jurassic and Cretaceous North Atlantis continent now foundered
below the N. Atlantic was not so far off the mark. However, this ancient land mass
was the continental area which is now Greenland and Canada, long before it was
broken up and pulled away from Europe since the Tertiary to the present day. It is
apparent that the boundary between the Albian ammonite provinces of the deposi-
tional areas that is now Europe, and that of Canada and the Western Cordillera of N.
America, occurred in the sea-way represented by the sediments in E. Greenland.

VI. CONDITIONS OF DEPOSITION IN ENGLAND

From the stratigraphical account of the Middle Albian sediments given above it is
possible to obtain some idea of the conditions which influenced their deposition in
England. Before commencing the discussion of these it is essential to consider first
two factors which provide a key to the interpretation of the field evidence.

It is apparent that the early Upper Albian (cristatum and orbignyi Subzones)
tectonic movements caused the planing-off of the upper surface of the Middle Albian
sediments throughout England. This period of erosion, although not as great as
Kitchin & Pringle held (1922a), removed a considerable amount of sediment,
including marginal deposits. Within a reasonably narrow limit the resulting surface
was probably plane, and for the purpose of this study it is taken so to be. From this
datum level, by comparing both the surviving thickness and the lithological sequence
from place to place, it is possible to make out the configuration of the surface upon
which the Middle Albian sediments were deposited.

It appears that there were only comparatively minor regional tectonic movements
within Middle Albian times. These seem to have consisted of minor shifting of the
axes of older folds indicated below, causing condensation or increased sedimentation.

140 MIDDLE ALBIAN STRATIGRAPHY

The main factor governing sedimentation appears to have been the pattern of
parallel ridges and troughs produced initially in late tardefurcata Zone times by a
comparatively mild folding phase. Casey from the field evidence afforded by largely
vemamé mammullatum Zone deposits interpreted these troughs as ‘ dimples ’ (196ra),
but the Middle Albian sediments have provided far more definite information on the
trend of these structures (text-fig. 52).

Although the ridges were not obviously active structures the sediments thin across
them due to water-current activity, and probably to some gravitational movement
of clay particles down-slope. During minor periods of current erosion the degree of
condensation is greater in the area of the ridges as one would expect.

(a) The Margins of the depositional basin
in England and Northern France (text-fig. 51)

To what extent the Brabant massif and the London Platform acted as positive
areas in Middle Albian times is far from certain. Nothing is yet known of the Gault
sequence in the area of the North Sea adjacent to the shores of Kent, Essex, Suffolk
and Norfolk. There certainly is no evidence made available at this time of the land
area suggested in this region by Jukes-Browne’s frontispiece map (1900). It is also
apparent from the borings in the Cliffe area of Kent (Owen in press), that renewed
movements along late Jurassic or early Cretaceous faults in the area of what is now
the Thames estuary, caused strong current action which removed Middle Albian
sediments over the southern part of Essex. This disturbance contributed to, and
was associated with other movements which planed-off the upper surface of the Lower
Gault throughout the Anglo-Paris Basin. How much sediment and the areal extent
that has been removed is at present unknown, but in Kent, and in Cambridgeshire
and Norfolk, where Middle Albian sediments are preserved, there is no evidence in
the sequence of an area of Palaeozoic rocks actively undergoing erosion to the east.

A shoal area existed in north-west Norfolk and in the area of the Lincolnshire and
Yorkshire Wolds and to an unknown extent in the adjacent area of the North Sea.
Its position is indicated by the pebbly development of Bed C of the Hunstanton Red
Rock (Wiltshire 1869 ; 185-188) of Middle Albian age, and its lateral equivalents
in the Red Chalk with its shallow-water fauna. To the south and east in Norfolk,
Bed C is replaced by clays of the contiguous Lower Gault. This shoal area probably
flanked the Palaeozoic massif of the Pennines and its southerly extension of the Peak
District and a possible positive area in the adjacent North Sea (Collette 1968 ; 20).
Gault clay probably existed south of the Pennine massif because derived Albian
fossils are known from the glacial boulder clay as far north as Chellaston, Derbyshire,
as well as elsewhere in the Midlands.

On the balance of evidence the writer is inclined to doubt that the Middle Albian
sea extended into the Cheshire lowlands but this could prove to be incorrect. The
clays of the lyelli and spathi Subzones at Swindon (Badbury Wick) and at Devizes are
very silty. Those of the intermedius Subzone at Devizes and Didcot are even coarser
in grade. These examples do not necessarily indicate the proximity of a marginal
area but equally they do not suggest an extensive basin area to the north west.

NORTH SEA
BASIN

> ARDENNES
mS
xX
NY

\

|

|
oe
| 3

xX MORVANO-VOSGES \
STRAIT

MORVAN

/
yi

Fic. 51. Palaeogeographic map of the Anglo-Paris Basin in the Middle Albian showing
links with the North Sea Basin, and Tethys via the Morvano-Vosges Strait. The
Aquitaine Basin is linked with Tethys via the proto-Atlantic.

142 MIDDLE ALBIAN STRATIGRAPHY

Nonetheless, the outcrop from Aylesbury (Bucks.) to Okeford Fitzpaine (Dorset)
shows a section across a basin in which the /yelli Subzone in particular is well devel-
oped, and apparently the zmtermedius Subzone as well. Whether the margin of the
Middle Albian depositional area ever reached Palaeozoic rocks in the Welsh
borders is unknown. However, it seems more probable that the margin flanked a
land area of Jurassic rocks from the nature of the sediments of the Gault in this area
(compare the relationship seen in the Ardennes). There is no doubt that a con-
siderable volume of clay sediment was carried eastwards into the depositional area
throughout Middle Albian times (p. 147) (cf. Jones 1955).

It has been shown (p. 53) that as one proceeds westwards along the south coast
from the Isle of Wight to the Devon border the basal fossiliferous bed and presumably
the underlying pebble beds become later in age. In the deeper part of the Wessex
Basin in the Isle of Wight, fine grade lyelli Subzone sediments rest upon coarsely-
graded marginal mammullatum Zone Carstone. By Lyme Regis, the lowest fossili-
ferous sediments are of intermedius Subzone age consisting of very gritty clays, and
rest upon a pebble bed which is of uncertain Subzonal age but probably not older than
spathi. The evidence indicates marked diachronism of the base of the Middle
Albian sediments, and the progressive transgression of the sea westwards across
Jurassic sediments particularly during the spathti Subzone. The furthest margin in
the extreme west probably followed the eastern boundary of the Mendip Hills and
across to the area of what is now the Blackdown Hills. There is no evidence of any
Middle Albian sediments in the western half of the English Channel.

On the northern coast of France, Middle Albian marginal sediments probably
flanked the Palaeozoic rocks north of the Cotentin Peninsula. They are certainly
present in the Pays de Caux where sediments of the eodentatus and niobe Subzones
consist of very coarse pebbly loams with a high clay content. However, the available
evidence from the outcrop and boreholes indicates that the land margin was not to
the south-west, south, or south-east, and it is necessary to look for a ridge which
underwent active erosion during the Middle Albian in the Baie de la Seine. There
seems no doubt that the western margin of the Middle Albian Paris Basin was flanked
by Jurassic sediments fringing the Armoricain Massif (text-fig. 33).

In the Artois, the eastern margin of the Middle Albian sea was formed by the
Brabant Massif. In the Boulonnais at Caffiers, and in borings in the area of the
Franco-Belgian border, Albian sediments rest upon Palaeozoic rocks on the fringe
of the Massif. Of particular interest here, especially in connection with the thinning
of Middle Albian sediments in the extreme east of Kent, is the marked rise in the
Palaeozoic floor in the region of the Quenoc off-shore from Cap Blanc Nez (p. 83).

In general, therefore, it is apparent that in England and in France a progressive
transgression over earlier Cretaceous and on to wide areas of Jurassic sediments
occurred between the eodentatus and niobe Subzones. Whether this continued later
into the loricatus and lautus Zones is uncertain because early Upper Albian erosion has
removed the evidence. From this definition of the margin of the northern part of
the Anglo-Paris Basin, it is now possible to look at the structures within the de-
positional area in England.

IN THE ANGLO-PARIS BASIN 143

(b) The Structural Controls on Deposition
in southern England (text-fig. 52)

The Variscan Wessex Basin of Kent (1949 ; 99) and its continuations into the
Weald and across the Channel into France subsided fairly steadily throughout the
Jurassic roughly in pace with the sediments which infilled it. At the end of the
Jurassic and at the opening of the Cretaceous Period, a strong phase of folding
and faulting occurred which in Dorset must have included fold amplitudes of well
over 1000 feet (>305 m.). The resulting basin of sedimentation was very greatly
reduced in area in comparison with that of the Jurassic. The history of Lower
Cretaceous sedimentation in southern England is the progressive erosion of the
resulting land area of Jurassic sediments and their redeposition within the basin.
This basin was by now restricted to the area of the Weald proper and eastern Hamp-
shire, and the English Channel flanking the Isle of Wight and Sussex. That the
deformation was essentially early Cretaceous is indicated by the distribution and
character of the Purbeck Beds and contiguous deposits. The marine Cinder Bed
within the Purbeck Beds of Dorset and equivalent horizons elsewhere are considered
by Casey to mark a marine incursion from the direction of the North Sea Basin (1963).
The sea did not invade southern England again until the early Aptian. It is
also important to note that in the Speeton area it is the top of the Kimmeridge
Clay which is eroded and that heavy clay sedimentation did not commence again
until after the start of the Neocomian.

The depositional history of the Lower Greensand has been discussed by Casey
(1961a ; 499-501). At the end of the tardefurcata Zone (Lower Albian) there occurred
the last of a number of minor folding phases which Casey has demonstrated affected
sedimentation during the formation of the Lower Greensand. This last phase
produced a number of parallel ridges and troughs trending between NW. and SE. to
WNW.-ESE., the axes of which were slightly modified during mammullatum Zone times
and again later in Joricatus Zone times. These, together with a general subsidence
of the whole Basin both in England and France oy a rise in sea-level, set the stage for
Middle Albian sedimentation. The positions of the axes of the structures are shown
diagrammatically in text-fig. 52.

The Middle Albian sequence under Dover shows a good development of the lyelli
Subzone overlain by a nodule bed of lower spathi Subzone age. By Folkestone the
lyella Subzone sediments are greatly reduced in thickness but the overlying dentatus
nodule bed is of exactly the same age as at Dover. The rest of the Lower Gault
sequence also is thinner at Folkestone, where it rests upon a mammillatum Zone
sequence in which all four Subzones are represented.

At Sandling Junction, the basal Gault rests upon puzostanus Subzone sediments as
at Folkestone, but these in turn rest directly on an eroded surface of early tardefurcata
Zone sediments. Here, the dentatus nodule bed contains species of Hoplites (H.)
transitional between those of the dentatus nodule bed at Folkestone and Dover, and
those which occur in the ‘ upper dentatus-spathi nodule bed’ in the northern Weald.
From Sandling to Maidstone, puzosianus Subzone sediments rest upon either tarde-
furcata or jacobt Zone Folkestone Beds, and the dentatus nodule bed in the basal Gault

MIDDLE ALBIAN STRATIGRAPHY

144

‘UOI}EJUIUIIPIs ULIQTY S[PPIN SUlOUeNpUr soxe [eInjonI}s 9y} SULMOYsS pue[sUY ysey YNOS Jo deur yo}eHS

WDYUS2/2234M-25M
WDUOIM-4IM
JSIPSPUDUIM-IM

UOID27 112M} S2M-2M
24M-M

2uopPSudWYIy-1y
2Bpispundy—1
u0}Bu1ss04S—-35S
s2a2Assuanbs—bs

Avg sz2406bsu0w 3S-EBWS

‘2S “OI

242uS-uS
2U4OQ 125-125
SYDOUDA2ZS—-as
Buijpuns-0s
umMOopuDS-Ss
A2| HOO Y-OY

pinomBuiy-y

UMOPS }JO0g—Og
UVJOMS2IPPDd-—d
POOMAN-AN

ssog Ag 2u0}spiow-gaw

S420SSDH-DH
11DH YORJOH-HH

AsPi1OD Psojyting-ng

2u0}S32y/04-04
Buiqqo4-—4
322435 2usOYAR-Sa
3221399q93-3
uz2uy vO.UNG-5g
Avg vO}dwo5-05
2ysD-1D

puojs} A2Au0D-[D
Asdipiod 32IS!UD-uD
pud|x2ng-ng
Buv6y201g-19
ausnoqgnsg-g
ys11pAy-Ay
uo}buippy-y

IN THE ANGLO-PARIS BASIN 145

has exactly the same degree of representation as at Sandling. There is strong
evidence to indicate that in lithological sequence and Subzonal representation the
remainder of the Gault sequence is uniform between Brabourne and the A 249 Clover
Leaf on the Maidstone By-Pass, and they are certainly identical from Hollingbourne
to the A 249.

In the area of the Kent Coalfield to the east, the pattern is not so clear but the
eodentatus and lyell1 Subzones sediments are well developed to the N. and NW. of
Dover in the Guilford, Tilmanstone, and Chislet collieries. The Lower Gault as a
whole is much thicker here than at the outcrop, but further east, on the Thanet and
E. Kent coast, the Gault in its entirety thins rapidly.

At the valley of the Medway north of Maidstone, the outcrop swings westwards
through an arc of about 12°. The change in the lithological sequence is striking
(text-fig. 3, p. 14) as the outcrop turns away from the line parallel to an axes to one
that cuts across the structural trend. From the four-division sequence in the Maid-
stone By-Pass, the Lower Gault expands into the six-division sequence recognisable
from Paddleworth near Snodland westwards at least as far as Dunton Green. The
central area of this Middle Albian trough seen in section at the outcrop occurs
roughly in the position of the Sevenoaks Brick Works but it migrated slightly NE.
during Middle Albian times. At Sevenoaks, the top of the Folkestone Beds contain
developments of soft sandstone in regular beds. This is overlain by a comparatively
thick development of the mammullatum Zone sediments. The top of lyelli Subzone
and the lower part of the spathi Subzone are represented by fine clays in contrast to
glauconitic clays or loam at Ford Place on the one side and at Dunton Green on the
other. The overlying ‘ upper dentatus-spathi nodule bed’ contains an ammonite
assemblage identical to that seen in all sections from Parsons Corner, Snodland,
to the Shere By-Pass. The remainder of the Lower Gault Divisions at Seven-
oaks when compared with Ford Place show the offset of the trough axes towards
the NE.

The whole Middle Albian sequence thins from the Sevenoaks Brick Works towards
Dunton Green. Further west, the eodentatus, lyelli, and spathi Subzones at least,
expand and are well developed in the area between Brasted and Covers Farm west
of Westerham. Unfortunately the rest of the Lower Gault is not exposed although
from the Brasted well it also is expanding west of Dunton Green. This boring also
demonstrates that the mammullatum Zone troughs do not always correspond to those
of the Middle Albian sediments, for here the mammuillatum Zone sediments are much
thicker than those to the east and west and are little condensed. In the Addington
Pumping Station situated 94 miles WNW. of Dunton Green the Lower Gault is of
much the same thickness and probably lies near a common ridge axis, running NW.
from Dunton Green.

At the Buckland Sand & Silica Co’s pit together with Wray Common, at Reigate,
the sequence although in general thicker and the sediments coarser, shows a similar
succession to that seen at Ford Place. At Shere, however, the eodentatus, lyelli and
basal spatht Subzones are again relatively well developed, but from here westwards
to Farnham the information is either poor or of uncertain value. Moreover, the
outcrop is faulted over much of this stretch of country.

K

146 MIDDLE ALBIAN STRATIGRAPHY

From the foregoing brief description, the evidence for NW. to SE. trending ridges
and troughs is not conclusive, but is very strong. However, in the southern Weald
and the Isle of Wight three pieces of evidence, in the writer’s opinion, tip the balance
strongly in favour of the interpretation here given.

The spatht Subzone sequence expands southwards from Wrecclesham to reach
a known maximum thickness at Selborne. From there it thins southwards towards
Nyewood, but the lithological sequence in detail remains the same. Although there
is a partial facies change and the sediments are coarser in E. Hampshire and W.
Sussex, the sequence in the spatii Subzone at Selborne lithologically is remarkably
close to that of the Horton Clay Pit, Upper Beeding (text-fig. 14, opp. p. 42) where it is
underlain by a thick development of the /yelli Subzone. Yet this sequence at Horton
Hall is quite different to that seen at Storrington where the lyelli Subzone sediments
have not been proved and if present at all are very thin and pebbly. In the opposite
direction, at Hassocks, the eodentatus and lyelli Subzones sediments are still well
developed but are grittier and more glauconitic and here as at Storrington the Gault
rests upon an ‘ [ron-grit’ which forms the indurated top of the sands of the Folkestone
Beds. At Horton Hall, however, the tardefurcata Zone is represented within clays
and loams, totally different to that of Hassocks and Storrington.

Along the WNW.-ESE. trending outcrop at the base of the South Downs from
Storrington to Petersfield, the Gault rests upon the ‘ Iron-grit’ (Kirkaldy 1935), below
which are normal loose sands of the Folkestone Beds. This sequence is seen at
Portsdown where the pre-Gault Lower Cretaceous sediments are greatly attenuated.
This area in which the ‘Iron-grit’ is present at the base of the Gault marks a long
swell on the pre-Middle Albian sea-floor which apparently increased in amplitude
towards the ESE. (p. 34). The sequence on the other side of the trough at Hassocks
has already been mentioned, and from what little is known of the Lower Albian
sequence near Eastbourne, and the Middle Albian sequence at Ringmer, the trough
extended ESE. from Upper Beeding towards Eastbourne. The sedimentation
remains very thick along this axis and if one projects the line through Selborne
into Wiltshire it again coincides with a broader area of thick dentatus Zone
sedimentation.

On the other, southern, side of the Storrington-Portsdown swell the lithological
sequence in the eodentatus, lyelli, and spatht Subzones in the Isle of Wight is totally
different from that of the Ringmer-Selborne trough. It is possible therefore, that
yet another WNW.-ESE. trending trough exists in the English Channel and which
includes the Isle of Wight. Both the mammillatum Zone sequence and the Lower
Gault increase in degree of representation towards the southern part of the Island.
The diachronous base of the Gault along the Dorset Coast can be explained if one
considers this line to be a diagonal section across the trough, the iutermedius Sub-
zone sediments in the Charmouth area being near the southern bounding ridge which
may have flanked a positive area in view of the sequence in the Pays de Caux.

In the E. and NE. part of the Weald the axes of the parallel ridges and troughs
trend as far as it is possible to judge in a NW. to SE. direction and they are with the
possible exception of the Kent Coalfield fairly closely set and linear (text-fig. 52).
The apparent opening-out of the troughs in the northern Weald W. of Dunton Green

IN THE ANGLO-PARIS BASIN 147

is due partly to the fact that the outcrop tends to swing more parallel to the axes.
In the southern Weald the axes have swung WNW.-ESE. and the structures are
more open. These structures have what is normally considered to be an Armorican
trend, and the Middle Albian is the last time that such closely lineated structures are
fully identifiable in the depositional environment.

(c) Source of the Middle Albian Sediments

Middle Albian sediments ranging in age from the eodentatus Subzone to the
intermedius Subzone rest, outside the area of the Weald and the eastern part of
Hampshire, the Isle of Wight and part of Purbeck, and the area extending NE. of
Aylesbury, directly upon Jurassic rocks. These Jurassic rocks had been folded and
faulted in the late Jurassic-early Cretaceous to becomeland. Below the London area
and N. Kent the Lower Gault, when present, rests upon thin Lower Greensand which
is underlain by either Jurassic or Palaeozoic rocks. The Middle Albian sea, therefore,
was clearly transgressive far outstripping the depositional area of the Lower Green-
sand which itself oversteps the Wealden and both of which derived their sediments
from the Jurassic land area. It is evident from the cobbles and blocks included
in the mammillatum Zone sediments of Kent that Palaeozoic rocks of the London
Platform were by then undergoing active erosion. From the borings in N. and
E. Kent there is no evidence, however, that the London Platform contributed
any large quantity of sediment during the deposition of the Lower Gault. This
probably consisted only of the silty fraction which is mixed with a fine clay
fraction.

In general as one moves west from the Kent coast the Middle Albian clays coarsen
in particle size and increase in the quantity of admixed silt andsand. This is readily
apparent if one compares for example in succession the sediments of the intermedius
Subzones at Folkestone, Buckland, in the Winchester borings, Devizes, and on the
Dorset coast in the Charmouth area. This suggests a main sediment source from
the western and north western margins of the sea, and possibly also from the south
in the area of the English Channel.

On the coast in the Isle of Purbeck and towards Weymouth and in the Charmouth
area the diachronous base of the Gault can be seen to rest directly upon extensive areas
of Jurassic clays such as the Kimmeridge Clay, Oxford Clay, and the Lias (text-fig.
23). In the area of the Hampshire Basin, British Petroleum Co., borings at Bere
Regis near Wareham (Dorset) and at Fordingbridge (Hants) show the Gault to rest
directly upon Oxford Clay and Kimmeridge Clay respectively. Along the northern out-
crop the position is much the same (text-fig. 18), with Middle Albian sediments resting
upon the Lias in the far west and then eastwards upon an eroded surface of folded
Jurassic sediments in which the Oxford and Kimmeridge Clays bulk large. Now this
is the state of affairs within the area of the depositional basin itself already having
undergone erosion since early Cretaceous times. Moreover, this is the depositional
basin which extended rapidly during spathi Subzone times to its greatest known
Middle Albian extent in the intermedius Subzone. Without any question the
originally far greater depositional area of the Jurassic clays must have undergone

K*

148 MIDDLE ALBIAN STRATIGRAPHY

active erosion during this period, and the Middle Albian sea may well have extended
further during later Subzones. In the writer’s opinion all the evidence points to a
source in the clays of the Jurassic, west of the London Platform, for the sediment
which was redeposited as the Lower Gault.

A Jurassic source to the west in Wales is suggested by the work of Jones (1955 ;
348-50), and by the borings at Port More, Antrim (Robbie & Manning 1966), and
Mochras, Merioneth (Wood & Woodland 1969) in which Lias is preserved. At
Port More, the incomplete remnant of Lower Lias is overlain by Upper Chalk
indicating a major intra-Mesozoic hiatus, the exact nature and extent of which is
uncertain at present.

Much the same state of affairs existed in the Paris Basin and it is an interesting
fact that here also the Middle Albian sediments are coarser in the west and finer in
the eastern areas of the Basin.

(d) The cristatum Subzone disturbance

After the commencement of the cristatwm Subzone, a major disturbance affected
the whole of the Anglo-Paris Basin and adjoining areas. In fact a break in sedimenta-
tion associated with erosion occurs widely throughout the Earth at about this time,
and is sometimes accompanied by folding. The disturbance caused the partial
planing-off of Middle Albian sediments over the whole area of the Anglo-Paris Basin.

The writer considered (1960 ; 377) that the planing-off of the upper surface of the
Lower Gault in southern England was due to a tilting movement up towards the west.
This may be true for the eastern half of England where definite early Upper Albian
faulting has been proved (Owen im press), but it is not necessarily the explanation
for the southern part of the country as a whole. In France, there is some evidence
of a similar tilting movement towards Morvan and Armorica. Although the effects
on the sediments is readily apparent, the main cause is much more obscure and may
be connected with faulting at the margin of Europe and America before the later de-
velopment of the Atlantic Ocean (p. 138).

Tectonic features are few in number, and there is certainly no evidence of anything
but a slight broad warping of the Basin as a whole, except for the faulting mentioned
above which removed Lower Gault sediments at least from the southern part of
Essex. The turbulent water conditions are reflected in the nektonic fauna ; for
example Inoceramus concentricus quite rapidly develops the far stronger sulcatus
form, and does not revert back to a concentricus form until the varicosum Subzone
when thick, little condensed sequences are seen again.

On the resulting planed-off Middle Albian surface Upper Albian sediments were
laid down in an entirely different pattern to that seen in the Middle Albian. More-
over, there are two intergrading facies ; the Upper Greensand and the Upper Gault.
This change in pattern renders meaningless isopachyte maps based on the sediments
of the whole Stage (Wooldridge & Linton 1938). It appears at present that the
Upper Albian depositional pattern is more closely related to that of the Upper
Cretaceous.

IN THE ANGLO-PARIS BASIN 149

VII. REVIEW OF THE AMMONITE FAUNA

The foregoing stratigraphical account has drawn heavily upon the evidence of
relative ages provided by the ammonites. For the purpose of this Bulletin the
subzonal distribution of the ammonite fauna of the Middle Albian, will be considered
only. Of the other stratigraphically useful fossils, some have been mentioned in the
text, but the foraminifera and ostracods require careful revision based on accurate
collecting. The two plates of zonal ammonites should be used in conjunction with
Spath’s Monograph (1923-43).

A. Description of new species

In order to stabilize the new taxa used in this Bulletin, brief descriptions are given
of one species of Hoplites (H.) and two species of Anahobplites.

Family HOPLITIDAE Douvillé 1890
Subfamily HOPLITINAE Douvillé 1890
Genus HOPLITES Neumayr 1875

Subgenus HOPLITES Neumayr 1875

Hoplites (Hoplites) maritimus sp. nov.
(PE, ayes. ga, “b)

1925a Hoplites rudis Parona & Bonarelli (pars) ; Spath : 108, pl. 8, fig. roc, d.

DERIVATION OF NAME. Hopflites :—heavily armoured soldier, maritimus : — of the
sea.

Diacnosis. Hoplites (Hoplites) with stout well-rounded whorl section bearing
coarse projecting tuberculate bullae about 10 per whorl, each buttressed by short
umbilical rib stemming from umbilical suture. Each bulla gives rise to two short
coarse ribs terminating above ventrolateral margin in coarse projecting clavi arranged
en-echelon each side of venter : intercostal areas merging onto venter. Ribs simple to
about 40 mm. diameter, thereafter there is tendency to develop occasional lautiform
ribs. Above 100 mm. ornament decreases in strength. Septal suture similar to
H. (H.) dentatus.

TYPE MATERIAL. Holotype BMNH. C 862a (J. S. Gardner Coll.) from Bed I (v) at
Folkestone.

DIMENSIONS. 53 °42 ‘52 °32.

Remarks. Like all species of this genus, H. (H.) maritimus shows variation
among individuals. Specimens of this form are, however, common in the lower part
of the spathi Subzone taking descent from the lyelli Subzone H. (H.) of the baylei-
benettianus group. At the base of the Subzone the ribbing is usually simple, but
higher up occasional intercalated ribs occur which produce a lautiform effect. Later
still, lautiform ribbing becomes well developed producing direct transitions to H. (H.)
canavarit Parona & Bonarelli of the upper part of the spathi Subzone.

150 MIDDLE ALBIAN STRATIGRAPHY

When found in condensed phosphatic nodule beds, this species shows a bewildering
series of morphological transitions to other coarsely ornamented species of Hoplites.
Most of these transitions are more apparent than real, in that coarse members of a
number of indirectly related offshoots which developed at slightly different times
have been collected together en melée in the same bed. To Spath it appeared that
this species belonged to Parona & Bonarelli’s Hoplites rudis but was not typical of it
(1925a ; 108). H. (H.) rudis does occur in England in the upper dentatus-spathi
nodule bed in the northern Weald, however, in the writer’s opinion it represents a
éoarse end member of a different development of Hoplites (H.).

Horizon & LocaLities. The species occurs throughout the spathi Subzone and is
ubiquitous in the Anglo-Paris basin.

Genus ANAHOPLITES Hyatt 1900
Anahoplites osmingtonensis sp. nov.
(Pl. 1, figs. ra, b)

1925a Anahoplites mimeticus Spath (pars) ; Spath : 142.
1927. Anahoplites mimeticus Spath (pars) ; Spath : 188, pl. 17, figs. 8a, b.

DERIVATION OF NAME. From Osmington, Dorset, the type locality.

Diacnosis. Anahoplites of the intermedius group : discoidal, compressed, evolute,
with excentric umbilicus. Umbilical wall steep in early whorls becoming rounded in
outer whorl. Umbilical margin marked by faint comma-shaped bullae giving rise to
faint striate ribs on gently curving whorl flank, terminating on ventro-lateral shoulders
at faint clavi absent on body chamber. Venter subtabulate slightly sulcate.
Suture line like A. planus, but symmetrical across venter and highly interlocked.

TYPE MATERIAL. Holotype BMNH., C 68385 (T. F. Grimsdale Coll.). Uppermost
spathi Subzone between Osmington Mills and Black Head, Dorset. Paratypes
BMNH., C 26595 same collection, horizon, and locality as Holotype. BMNH.,
37604 (Astier Coll.) condensed Middle Albian phosphatic bed, Escragnolles, Alpes
Maritimes, France.

DIMENSIONS.
C 68385 [107] -39 25 +33 (Holotype is slightly crushed)
37004 107 -40 <28 +30

REMARKS. This species differs from Anahoplites planus (Mantell) in its marked
excentric umbilicus, slightly sulcate venter which becomes broadly rounded on the
body chamber, and the symmetrical arrangement of the suture line each side of the
venter. The elements of the suture line are closely interlocked resembling specimens
of A. planus from the cristatum Subzone. It is, however, probably not directly
related to A. planus, an early mutation of which occurs in the same bed (e.g. Spath
1927 ; 188, pl. 18, figs. 7a, b), but is very near the stem from which A. grimsdalet,
A. evolutus, A. intermedius, A. mantelli, A. praecox, and A. alternatus sprang.

IN THE ANGLO-PARIS BASIN I51

Spath referred the paratypes of A. osmingtonensis to A. mimeticus. However,
the type of A. mimeticus (BMNH. C 30535, 1925 ; 131, pl. 11, figs. 7a, b) shows that
the inner whorls are strongly costate with well developed umbilical bullae quite
unlike A. osmingtonensts and in fact probably does not belong to Anahoplites at all.
The stratigraphical horizon of the Holotype of A. mimeticus has been discussed
above (p. 47), and it is not considered here to belong to the mammullatum Zone genus
Anahoplitoides (Casey 1966 ; 547-8).

HORIZON & LOCALITIES. A. osmingtonensis occurs at the extreme top of the spathi
Subzone both in the Osmington area of Dorset, and the Petersfield area, Hampshire
(BMNH. C 35483). It is also present in the condensed spathi, intermedius Subzones
assemblage of Bed 4 at the Carriére Binot, St. Florentin (Yonne), and also at
Escragnolles (Alpes Maritimes).

Anahoplites grimsdalei sp. nov.
(Pl. 1, figs; 2a, b)

DERIVATION OF NAME. After Mr. T. F. Grimsdale.

Diacnosis. <Anahoplites of the intermedius group ; discoidal, compressed, evolute.
Umbilical wall rounded, marked at margin by 20 comma-shaped bullae on outer
complete whorl. Each bulla gives rise to primary rib which bifurcates either just
above it or near middle of whorl flank. Ribs sickle-shaped, faint on inner whorls
terminating at ventro-lateral margins in distinct clavi arranged en-echelon each side of
tabulate slightly sulcate venter. Occasional short intercalated ribs stem from
primary ribs, one for each primary, on ventro-lateral shoulder to end at ventro-
lateral margin in intercalated clavi. Suture line as in A. intermedius.

TYPE MATERIAL. Holotype BMNH. C 31702 (Lt. Col. R. H. Cunnington Coll.)
Uppermost spathi Subzone, between Osmington Mills and Black Head, Dorset.

DIMENSIONS. 93 -40 -20 -30 (slightly crushed laterally)

REMARKS. This species differs from A. evolutus of the basal part of the intermedius
Subzone, to which it is closely related, by possessing fainter and more distant ribbing
but which are more regularly bifurcating. Although the Holotype is slightly crushed
it is probably more compressed than A. evolutus. The short intercalated ribs on the
ventro-lateral shoulders, absent in A. evolutus, are reminiscent of those seen in the
much later Semenovites gracilis (Spath) of the varicosum Subzone (Upper Albian).
These are taken to an extreme development in a form which occurs in the basal part
of the intermedius Subzone at Caen Hill, Devizes (Bed 7). Nonetheless, A. grimsdalei
in fact links A. osmingtonensis and A. evolutus.

HORIZON AND LOCALITY. Extreme top of the spathi Subzone in the Osmington
area of Dorset, and at Petersfield, Hampshire (BMNH. C. 35482).

152 MIDDLE ALBIAN STRATIGRAPHY

B. Stratigraphical List

To conserve space, Subzones are numbered consecutively. (1) eodentatus, (2) lyelli, (3) spatht ;
(4) intermedius, (5) niobe, (6) subdelaruet, (7) meandrinus ; (8) nitidus, (9) daviesi. p= praemuta-

tion,
Sa Se
x 351k Sat Cee
Phylloceratidae
Hypophylloceras sp. . . ° ° ° x
Lytoceratidae
Pictetia astieriana (d’Orbigny) ° ‘ . x
Tetragonitidae
Tetragonites kitchint (Krenkel) . . . x
Hamitidae
Hamites (Hamites) attenuatus J. Sowerby ee | Oe Oe, |e oe
3 a votundus J. Sowerby . DS POS WPS OES I OS |] xs
9 Ap tenuicostatus Spath . c DOS OS IL
55 re compressus J. Sowerby . DE) Rh Re |] DS eae
o Fr is gracilis Spath . PP Se Pe PSS, 5
as 7 gibbosus J. Sowerby . : > eP:GPsGaesS |] Os || 2<
a5 i subrotundus Spath : Se [| ee
ss Ee maximus J. Sowerby ~ 2 |) 8
ep oF tenuis J. Sowerby A Xen | nexe
55 33 3» subacuaria Spath . Slt DS
Anisoceratidae
Protanisoceras (Protanisoceras) alternotuberculatum
(Leymerie) x
a An barrense
(Buvignier) x
ae Ay nodoneum
(Buvignier) x
as a3 moreanum
(Buvignier) xX
of spp. DS ||) Oe, |] O5€
‘ (Heteroclinus) nodosum (J. Sowerby) x
F flexuosum (v’ ae x
M stahamites sablieri (d’ fener, ‘ ; x
spp. 4 : ‘ xX |X
Turrilitidae |
Proturvilitoides densicostatus (Passendorfer) . x
Pseudhelicoceras argonnensis (Buvignier) 3 x
3 subcatenatum Spath . . x
Binneyitidae ?
Falciferella nulbournet Casey - . : x |X
Desmoceratidae
Puzosia (Anapuzosia) provincialis (Parona &
Bonarelli) x
Uhligella derancet Casey : . : 5 x
5 evugata Casey . : xX
Beudanticeras laevigatum (J. de C. Sowerby) : xX] X
yy sanctaecrucis Bonarelli . Exe | a
HA albense Breistroffer . BS || e.<
Desmoceras (Desmoceras) latidorsatum (Michelin) xX x

IN THE ANGLO-PARIS BASIN

Douvilleiceratidae
Douvilleiceras inaequinodum (Quenstedt)

clementinum (d’Orbigny)

Engonoceratidae
Engonoceras tvis Spath a : :

Hoplitidae

Cleoniceras (Cleoniceras) devisense Spath
Otohoplites spp.

3?

2 cunningtoni Spath

Hoplites (Isohoplites) steinmann (Jacob)

= eodentatus Casey

» Sppsnetr : é
(Hoplites) dentatus (J. Sowerby)
robusta Spath
Bs », adensicostata Spath
», sulcata Seitz
5 baylet Spath
ap bullatus Spath .
benettianus (J. de C. Sowerby)
spathi Breistroffer :
persulcatus Spath 4
a paronai Spath . -
maritimus Owen nov.
a mirvabiliformis Spath . -
Aa obtusus Spath . :
+ pringlet Spath . > 5
os similis Spath 4
mivabilis Parona & Bonarelli
vectensis Spath . . :
BS latesulcatus Spath : 4
escragnollensis Spath
yudis Parona & Bonarelli
BA dorsetensis Spath
canavarit Parona & Bonarelli
uf canavartiformis Spath
45 pretethydis Spath
Fe dentatifornris ekg

SPp-

” Pel

Anahoplites osmingtonensis Owen noy.

grvimsdalet Owen nov. .
evolutus Spath
mimeticus Spath
intermedius Spath
mantellt Spath
praecox Spath :
alternatus (Woodward)
planus (Mantel!)
» compressa Spath
» ~mflata Spath p
x discoidea Spath ;
» sulcata Spath . C 5
» &vacilis Spath . 5 F

PA A WM OM

A A
A

AA

“Anan Analanalala’

VA A A PM MOM
an naw
A Po 4 A OA

A WA'PA p4
PA PA

vA bd bd bd

al

354 MIDDLE ALBIAN STRATIGRAPHY

oo a 3 dS Oe ae
Anahoplites splendens (J. Sowerby) : : OG DS 12S Ee@ HOS, || OS
“e pleurophorus Spath , : é X
35 daviest Spath 4 x
55 » ovnata Spath dS
a » elegans Spath , x
ce Sph 4. a : : 5 : x
Dimorphoplites niobe Spath . P ‘ : oD I-45) | 9.<
a doris Spath PXS a | ex
Ce pinax Spath . Xp| X
a », elegans Spath xX
a biplicatus (Mantell) Xp) | See ee
“8 hilli Spath x
a perelegans Spath xX
ee crassa Spath , ipae
80 parkinsont Spath DS 126 || ES
aS tethydis (Bayle) . xX |X
is », Spath non Bayle D1] D8
re glaber Spath. 4 Dall ee
chloris Spath GRE

Euhoplites subtabulatus Spath

55 pricei Spath xX |X
oe lovicatus Spath aX 2S [2S |] 2s
Ss subtuberculatus Spath SHES EXC |e
5 aspasia Spath =, - XS) EXE |/PXE 5 Xe
A microcevas Spath , 2) Sail 3S, |] BS
Ag meandrinus Spath x
sf cantianus Spath xX
55 bilobus Spath x
oF beaneyi Milbourne de
oF truncatus Spath xX |X
» a5 quadrata Spath S| OS
de lautus (J. Sowerby) DG Ds€
NB », duntonensis Spath PSE 2S
ie nitidus Spath XK |X
Ap opalinus Spath . xX |X
5 proboscideus (J. Sowerby) SSeS
” intermedia Spath xX |X
” bucklandi Spath ee
Lyelliceratidae

Lyelliceras camatieanum (d’Orbigny) ‘ , x
fs pseudolyelli (Parona & Bonarelli) xX
a5 hirsutum (Parona & Bonarelli) x
Be hubevianum (Pictet) x
a lelli (d’Orbigny) : x
” vadenaci (Pervinquiere) x ;
a cotteri Spath x ;

gevreyt (Jacob) xX

Brancoceras (Brancoceras) senequieri (V’ Orbigny) x -
7 55 versicostatum (Michelin) x
” rr spp. ; . , x | xX

IN THE ANGLO-PARIS BASIN 155

Eubrancoceras (Eubrancoceras) aegoceratoides

(Steinmann) x
ts fs crickt Spath SP 2:€
a5 3 spp. . . EXE ilies lhe Ne Nees
Mojsisovicsiidae

Oxytropidoceras evanst Spath . ‘ ‘ x

Bs voissyanum (d’Orbigny) . De

33 mivapeiianum (d’Orbigny) oe

5 cf. carbonarium (Gabb) . x

3 Soh e P 5 DPSS
Mojsisovicsia delaruet (d’Orbigny) ‘ x

ff 56 compressa Spath xX

5 subdelaruei Spath OS

i vemota Spath xX

A spinulosa Spath 5 : x

os equicostata Spath : : : x
Dipoloceras cornutum (Pictet) “ - . x |X

A

Falloticeras proteum (d’Orbigny) . 4 :

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A ee OE. ya

— ra

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H. G. OwEn B.Sc., Ph.D., F.G.S.
Department of Palaeontology

British Museum (NATURAL History)
CROMWELL Roap

Lonpon S.W.7

>)

PLATE 1

Anahoplites osmingtonensis sp. nov.
Fics. 1a, b. Lateral and peripheral views of holotype (BMNH. C 68385) X 1. Uppermost
spathi Subzone, between Osmington Mills and Black Head, Dorset.

Anahoplites grimsdalei sp. nov.
Fics. 2 a, b. Lateral and peripheral views of holotype (BMNH. C 31702) X 1. Uppermost
spatht Subzone, between Osmington Mills and Black Head, Dorset.

Hoplites (Hoplites) maritimus sp. nov.
Fics. 3 a, b. Lateral and peripheral views of holotype (BMNH. C 862a) X 1. Spathi Sub-
zone, Lower Gault Bed I (v), Folkestone, Kent.

ee

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 8 PLATE 1

PLATE 2

Dentatus Zone Indices
Hoplites (Isohoplites) eodentatus Casey
Fics.1a,b. Lateral and peripheral views X 1 of an example from Bed 1, eodentatus Subzone,
Coney Hill Sand pit, Tandridge, Surrey. (BMNH. C 76480)

Lyelliceras lyelli (d’Orbigny)
Fics. 2a,b. Lateral and peripheral views X 1 of an example from Division 2 (iv), lyelli Sub-
zone, Horton Clay Pit, Upper Beeding, Sussex. (BMNH. C 76481)

Hoplites (Hoplites) spathi Breistroffer
Fics. 3 a,b. Lateral and peripheral views X 1 of a late mutation from Bed 4, upper part of
the spatht Subzone, Buckland Sand & Silica Co. pit, Reigate, Surrey. (BMNH. C 76483)

Hoplites (Hoplites) dentatus (J. Sowerby)
Fics. 4a, b. Lateral and peripheral views X 1 of a late mutation from the same bed and
locality as Fics. 3 a, b. (BNMH. C 76482)

(All specimens author’s coll.)

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 8 PLATE 2

PLATE 3

Loricatus and Lautus Zone Indices
Euhoplites loricatus Spath
Fics. 1a,b. Lateral and peripheral views X 1 of a late form from Division 5 (ii), meandvinus
Subzone, Sevenoaks Brick Works, Otford, Kent. (BMNH. C 76484)

Anahoplites intermedius Spath
Fics. 2 a,b. Lateral and peripheral views X 1 of a body chamber fragment from Division 3
(iv), intermedius Subzone, same locality as Fics. 1 a,b. (BMNH. C 76485)

Dimorphoplites niobe Spath
Fics. 3 a,b. Lateral and peripheral views X 1 of an example from 4 inches below the top of
Bed III, niobe Subzone. Folkestone, Kent. (BMNH. C 76488)

Mojsisovicsia subdelaruei Spath
Fic. 4. Two immature crushed individuals in a block of clay from 2 feet 2 inches above the
base of Division 4, subdelaruei Subzone, Sevenoaks Brick Works, Otford, Kent. (BMNH.
C 76486)

Euhoplites meandrinus Spath
Fics. 5 a, b. Lateral and peripheral views X 1 of a typical fragment from Bed IV (iii),
meandrinus Subzone, Folkestone, Kent. (BMNH. C 76480)

Euhoplites nitidus Spath
Fics. 6 a,b. Lateral and peripheral views X 1 of an example from Bed V, nitzdus Subzone,
Folkestone, Kent. (BMNH. C 76490)

Euhoplites lautus (J. Sowerby)
Fics. 7a,b. Lateral and peripheral views X 1 of a wholly septate phosphatic steinkern from
the condensed ‘ /autus Zone nodule bed’ Division 6, Sevenoaks Brick Works, Otford, Kent.
(BMNH. C 76487)

Anahoplites daviesi Spath
Fics. 8a,b. Lateral and peripheral views X 1 of an involute form from the upper part of Bed
VII daviesi Subzone, Folkestone, Kent. (BMNH. C 76491)

(All specimens author’s coll.)

PLATE 3

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 8

—_

‘4

+4

A LIST OF SUPPLEMENTS
TO THE GEOLOGICAL SERIES Pane
OF THE BULLETIN OF a,
THE BRITISH MUSEUM (NATURAL HISTORY) =

1. Cox, L. E, Jurassic Bivalvia and Gastropoda from Tanganyika and Kenya. ae
Pp. 213; 30 Plates; 2 Text-figures. 1965. £6. ne

2. Et-Nacecar, Z. R. Stratigraphy and Planktonic Foraminifera of the Ue >
Cretaceous—Lower Tertiary Succession in the Esna-Idfu Region, Nile Yay
Egypt, U.A.R. Pp. 291; 23 Plates; 18 Text-figures. 1966. {10.

3. DAveEy, R. J., Downtz, C., SARGEANT, W. A. S. & WiLtiams, G. L. Studies. on
Mesozoic and Cainozoic Dinoflagellate Cysts. Pp. 248; 28 Plates, 64 Text R
figures. 1966. {7.

3. APPENDIX. DAVEY, R. J., DowniE, C., SARGEANT, W. A. S. & Wittams, G L
Appendix to Studies on Mesozoic and Cainozoic Dinoflagellate Cysts. Fp. 24.
1969. 16s.

4. E.tiott, G. F. Permian to Palaeocene Calcareous Algae (Dasycladaceae) of the
Middle East. Pp. 111; 24 Plates, 17 Text-figures. 1968. £5 2s. 6d. “e

5. Ruopves, F.H.T., Austin, R.L.& Druce,E.C. British Avonian (Carboniferous)
Conodont faunas, and their value in local and continental correlation, et. 31
31 Plates, 92 Text-figures. 1969. 11. —

6. Cuitps, A. Upper Jurassic Rhynchonellid Brachiopods from Northwestern
Europe. Pp. 119; 12 Plates, 40 Text-figures. 1969. {£4 I5s. iS

7. Goopy,P.C. Therelationships of certain Upper Cretaceous Teleosts with special as
reference to the Myctophoids Pp. 255; 102 Text-figures. 1969. {610s. x

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BULLETIN OF
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Parts willappear atirregular intervals as they become
veady. Volumes will contain about three or four
hundred pages, and will not necessarily be completed
within one calendar year.

In 1965 a@ separate supplementary series of longer
papers was instituted, numbered serially for each
Department.

This paper is Supplement 9 of the Geological
(Palacontological) series. The abbreviated titles of
periodicals cited follow those of the World List of
Scientific Periodicals.

World List abbreviation
Bull. Br. Mus. nat. Hist. (Geol.) Suppl.

© Trustees of the British Museum (Natural History) 1971

TRUSTEES OF
THE BRITISH MUSEUM (NATURAL HISTORY)

Issued 23 February, 1971 Price £8

EARLY TERTIARY OSTRACODA OF THE
FAMILY TRACHYLEBERIDIDAE FROM
WEST PAKISTAN

By Q. A. SIDDIQUI

CONTENTS
Page
I. INTRODUCTION é - - : : : : : 3 5
II. ACKNOWLEDGMENTS . - ; : - : : - - 7
III. LiTHOLoGIcaL UNITS : : : : - : : : 8
IV. SYSTEMATIC DESCRIPTIONS : : : : - : : Io
Subclass Ostracoda Latreille . : 5 : ; - - 10
Order Podocopida Miiller . : : : ° : 10
Suborder Podocopina Sars : : ‘ : ; : Io
Superfamily Cytheracea Baird : : : Io
Family Trachyleberididae Sylvester Bradley : : 10
Genus Actinocythereis Puri. : ; Io
Actinocythereis ? quasibathonica sp. nov. : : Io
Genus Alocopocythere nov. : : : : 13
Alocopocythere tvanscendens sp.nov. . : ; 14
Alocopocythere rupina sp.nov. . : : : 16
Alocopocythere abstvacta sp. nov. . : : : 17
Alocopocythere coarctata sp. nov. . ‘ : c 18
Alocopocythere longilinea sp. nov. : : : 18
Alocopocythere transversa sp. nov. : : : 19
Alocopocythere vadiata sp.nov. . : : : 22
Genus “ Anommatocythere’’ Sohn . : ‘ : 23
““ Anommatocythere ’’ laqueta sp. nov. : ; 23
“ Anommatocythere’’ confirmata sp.nov. . - 24
Genus Bradleya Hornibrook . : : : 5 26
Bradleya ? voraginosa sp. nov. . ; : : 26
Genus Buntonia Howe : : : : 5 27
Buntonia devexa sp.nov. . : ‘ : : 27
Buntonia sp. A , : : ‘ ‘ ‘ 28
Genus Costa Neviani_. : ; ; : : 28
Subgenus Paracosta nov. . : : : 28
Costa (Paracosta) declivis sp.nov. . : é 29
Costa (Paracosta) compitalis sp. nov. : : 30
Costa (Paracosta) disintegrata sp. nov. : : 31
Genus Echinocythereis Puri. é é - 31
Subgenus Echinocythereis sensu stricto : : 32
Echinocytherets (Echinocythereis) contexta sp. nov. 32
Echinocythereis (Echinocythereis) elongata sp. nov. 33

Subgenus Scelidocythereis nov. . ‘ F : 33

EARLY TERTIARY OSTRACODA

Echinocythereis (Scelidocythereis) multibullata sp.
nov. . .
Echinocythereis (Scelidocytheveis) sp. A é
Echinocythereis (Scelidocythereis) vasilis sp. nov. .
Echinocythereis (Scelidocythereis) sparsa sp.nov. .
Genus Gyrocythere nov. : ; .
Gyrocythere exaggerata sp. nov.
Gyrocythere parvicarinata sp. nov.
Gyrocythere grandilaevis sp. nov.
Gyrocythere mitigata sp. nov.
Gyrocythere perfecta sp. nov.
Genus Hermanites Puri
Hermanites cracens sp. nov.
Hermanites scopus sp. nov.
Hermanites palmatus sp. nov.
Genus Occultocythereis Howe .
Occultocythereis interrupta sp. nov.
Occultocythereis sp.A
Occultocythereis spilota sp. nov.
Occultocythereis peristicta sp. nov.
Occultocythereis indistincta sp. nov.
Genus Patagonacythere Hartmann
Patagonacythere ? nidulus sp. nov.
Genus Phalcocythere nov. -
Phalcocythere horrescens (Bosquet)
Phalcocythevre improcera sp. nov.
Phalcocythere rete sp. nov.
Phalcocythere vetispinata sp. nov.
Phalcocythere sentosa sp. nov.
Phalcocythere dissenta sp. nov.
Phalcocythere spinosa sp. nov.
Phalcocythere sp. cf. P. spinosa
Genus Quadvacythere Hornibrook
Subgenus Hornibrookella Moos
Quadracythere (Hornibrookella) platybomus sp. nov.
Quadracythere (Hornibrookella) directa sp. nov.
Quadracythere (Hornibrookella) arcana aera
and Guha)
Quadracythere (Hornibrookella) subquadva sp. nov.
Quadracythere (H. the ae sp.A
Genus Stigmatocythere nov.
Stigmatocythere obliqua sp. nov.
Stigmatocythere portentum sp. nov.
Stigmatocythere calia sp. nov.
Stigmatocythere delineata sp. nov.
Stigmatocythere lumaria sp. nov.
Genus Tvachyleberis Brady 5
Subgenus Tvachyleberis sensu stricto
Trachyleberis (Tvachyleberis) lobuculus sp. nov.
Trachyleberis (Tvachyleberis) bimammillata sp. nov.
Subgenus Acanthocythereis Howe
Tvachyleberis (Acanthocythereis) procapsus sp. nov.
Trachyleberis (Acanthocythereis) usitata sp. nov.
Trachyleberis (Acanthocythereis) pedigaster sp. nov.

FROM WEST PAKISTAN 5

Trachyleberis (Acanthocythereis) postcornis sp. nov. 82
Trachyleberis (Acanthocythereis) decoris sp. nov. . 83

V. OsSTRACODA AND EARLY TERTIARY CORRELATION IN THE SULAIMAN
RANGE ; é ; : 5 . 2 , 85
(a) Biostratigraphic Units ; 85

(b) Statistical Correlation of ranges of ostracod species common to
the Rakhi Nala and Zao River sections : A : F 89
(c) Conclusions ‘ : i 0 - . : : : gI
VI. APPENDICES : : 5 : ; ‘ j 5 ‘ 93
VII. REFERENCES . : , 95
TABLES, AND RAKHI Naa AND Zao RIVER “SECTIONS In pocket on
Cover Pp. ili

SYNOPSIS

Ostracoda from the Palaeocene of the Sor Range and from the Palaeocene and Eocene of the
Rakhi Nala, Zao River and Shpalai Khwara sections, Sulaiman Range, West Pakistan, have
been examined. The family TRACHYLEBERIDIDAE has been studied in detail. It is
represented by fourteen genera, four subgenera and fifty-nine species. Four new genera
(Alocopocythere, Gyrocythere, Phalcocythere and Stigmatocythere) and two new subgenera (Para-
costa and Scelidocytheveis) are proposed. Out of the fifty-nine species described, fifty-four are
new. Two species belonging to the genus Phalcocythere one from the Paris Basin and the other
from Tanzania are also described.

The Palaeocene and Eocene of the Rakhi Nala section are divided into five ostracod bio-
stratigraphic units. The biostratigraphic units IV and V of the Rakhi Nala are represented in
the Zao River section and have almost identical ostracod faunas. The biostratigraphic unit
IV of the Rakhi Nala is also represented in the Shpalai Khwara section. The Equations of
Correlation between the Rakhi Nala and Zao River sections for biostratigraphic unit V (i.e.
Middle-Upper Eocene) have been calculated by means of ranges of ostracod species common to
the two sections. The standard errors of estimate for the Equations of Correlation have also
been calculated. The boundaries between the Palaeocene-Lower Eocene, Lower-Middle
Eocene and Middle-Upper Eocene in the Sulaiman Range are discussed.

t. INTRODUCTION

THE most comprehensive work so far published on the area is that of Eames (1952 ab).
Most of his lithological subdivisions for the Eocene succession of the Rakhi Nala and
Zinda Pir areas occur in the northern Sulaiman Range, i.e. in the Zao River and
Shpalai Khwara sections. These can easily be distinguished on the basis of lithology
and microfauna. Eames’ terminology of the rock units is therefore adopted here.
Bayliss (1961) and Latif (1961 and 1964) are other recent workers who have con-
tributed to our knowledge of the Palaeocene and Eocene in the Rakhi Nala section.
However, they used a different terminology for the rock units to that used by
Eames, and Fig. 2 shows the correlation between these workers along the Rakhi Nala
section.

The samples from the Rakhi Nala section examined for ostracods were the same as
used by Bayliss and Latif, who worked on larger and pelagic foraminifera respect-
ively. These samples were collected by Bayliss. The sample numbers as given by
the collector are used in this paper. Latif altered the sample numbers after 3200 by
subtracting two hundred, i.e, his sample no. 3201 is the same as collector’s no. 3401,
and so on,

EARLY TERTIARY OSTRACODA
°
. Zao River Section 68
. Shpalai Khwara Section

Rakhi Nala Section
G. Sor Range Section

Eocene Outcrops

32°

Kandahar

Y Fpl sii
KARACHI lie :

Outcrops of Eocene rocks of part of West Pakistan.
(After Eames 1952.) Sections described are indexed.
FIG.1

Correlation between Eames, Bayliss and Latif along the Rakhi Nala

Eames (1952, p.162-163 ) Bayliss & Samanta (in press) Latif (1961, p. 33-36, 1964, p. 31)

Thickness [Sample no. Thickness | Thickness
i 6 jas used in i i j Zones Succession
Succession in feet [present Succession in feet & | in feet &
paper sample nos.}/samplenos.
: 60 657 No

==] = |Pellatispira Beds EEE S 3666 se, |GlObigerina cf. trilocula ; BE
as | ¢ 425-495 a9 [zy ——"seso__[ Chiloguembeling _victoriana] 4 | Tapti a §
80 E Set 0 FS a 3b20 eee aS 1425 34 | Hastigerina_micra Sg
4 ye = 5 Chiloguembelina aff_ martini Go =
CAV-D | at Ais H70 $85 | Gatapsydrax unicavu ie =e

7 ? a2 3606 : : ; v
Kirthar os (|| 2212 160 “f° |Globigerina yequaensis =e

34!
3478

No Pelagic foraminifera

ig

[100-4484 «| Globorotalia sp. 3

3459

1160 Globorotalia sp.4

(Ghazij ) **
Lower Eocene

Lower Eocene

Upper Rakhi Gaj Shales

Laki

Lower Rakhi Gaj Shales. (topmost
portion only = Irregularis Limestone.

+3667-3672 Dunghan
Lower Rakhi Gaj Shales (Max pars) 775 t
3116

fo | Globigerina sp.5

S [100 38s | Globorotalia rex
ealee 5 | 759 roo 38 a Globorotalia (Tr) crater S
S) = Oo Globorotalia angulata 4 9
Cc 3116
8 e Gorge Beds ae 8 3115 Rani kot 9
G loro = ; at Ranikot © [553 310 |850 No Pelagic foraminifera s
a mi! Venericardia Shales 95 e is

Pab

UPPER
CRETACEOUS

» Probable equivalents of Eames, 1952.
**Eames, personal communication.

+ Danian according to Nagappa, 1959, which he regards as basal Paleocene. FIG. 2
* 2 Paleocene.

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FROM WEST PAKISTAN 7

Samples from the Zao River and Shpalai Khwara sections were taken by S. M.
Ahmed and W. A. Zuberi and those from the Sor Range section by J. A. Reinemund.

All specimens with the prefix Io. are in the Department of Palaeontology, British
Museum (Natural History). Those with the prefix GSP BM are in the Museum of the
Geological Survey of Pakistan, Quetta.

II. ACKNOWLEDGMENTS

I would like to express my sincere gratitude to Professor P. C. Sylvester-Bradley
for his supervision, encouragement, constant help throughout this work, and for the
use of the Department facilities at Leicester University. I am greatly indebted to
Dr. F. E. Eames, lately Chief Palaeontologist of the British Petroleum Company Ltd.,
for his help and advice on the stratigraphy, particularly that of the Sulaiman Range,
and for allowing me to examine East African Eocene ostracod collections at the BP
Research Centre. Thanks are due especially to Dr. E. Triebel of the Senckenberg
Museum, Frankfurt a.M., for his instructions in photomicrography and for kindly
permitting me to study the ostracod collection at the Museum. During my several
visits to the British Museum (Natural History), I have been received with courtesy
by Dr. J. P. Harding and Dr. R. H. Bate, who gave me free access to the ostracod
collections under their care. In addition, Dr. R. H. Bate read the manuscript
critically. I have profited from useful discussions with Professor R. A. Reyment
which I had while on a study tour to Stockholm. He kindly allowed me to see his
West African ostracod collection. Dr. P. Marks helped me during my stay at
Utrecht and gave me free access to the van den Bold, Kingma and Keij Collections
housed in the Geologisch Instituut.

I should like to thank Dr. F. T. Banner (University College, Swansea) for examin-
ing some of the smaller foraminifera from the Rakhi Nala and Sor Range sections ;
Dr. C. G. Adams (British Museum, Natural History) for his help in identifying the
genus Pellatispiva from the Zao River ; and Mr. J. A. Reinemund (U.S. Geological
Survey) for the information on the Sor Range locality.

For the loan of samples, I am indebted to the following : Standard Vacuum Oil
Company, Karachi ; The Director, Geological Survey of Pakistan, Quetta ; Dr. I.
Strachan, Birmingham University ; Dr. D. D. Bayliss, Robertson Research Ltd.
I would like to acknowledge the following persons for comparative material : Mr. E. S.
Pinfold, Geological Adviser of the Attock Oil Co. Ltd. ; Dr. F. E. Eames, lately Chief
Palaeontologist of the British Petroleum Co. Ltd. ; Mr. I. G. Sohn, U.S.A. ; Pro-
fessor A. Wood, Aberystwyth ; The Director, Oil and Gas Commission, India ; Dr.
W. A. van den Bold, U.S.A. ; Dr. W. D. I. Rolfe, of the Hunterian Museum, Glasgow ;
Dr. N. Grekoff, France ; Dr. R. C. Whatley, Aberystwyth ; Dr. J. E. van Hinte,
Holland ; and Professor G. Ruggieri, Italy.

I would like to thank the departmental technical staff at Leicester University,
particularly Mr. M. Barker and Mr. G. McTurk for their assistance in photography.
I am very grateful to Mrs. N. Farquharson for making the diagrams and charts.

This work has been done during the tenure of a Leicester University Research
Scholarship. The study tours to Frankfurt, Utrecht and Stockholm were made
possible by two travelling grants from the Leicester University Research Board.

8 EARLY TERTIARY OSTRACODA

III. LITHOLOGICAL UNITS

Sulaiman Range. The lithological units of the Rakhi Nala section have been
described in detail by Eames (1952, pp. 162-165), Bayliss (1961) and Latif (196r,
p-. 32). Fig. 2 shows the succession, and the formation names give some idea of the
lithology ; for a fuller description, see the authors mentioned above. The Eocene
succession in the Zao River and Shpalai Khwara sections (Fig. 3) is very similar to
that of the Rakhi Nala section. A detailed lithological description of rocks exposed
along the Rakhi Nala, Zao River and Shpalai Khwara sections is given by means of
two charts.

ZAO RIVER - |SHPALAI KHWARA

; Thickness | Sample] Thickness} Sample

24210
Upper Chocolate Clays (Upper part)}| 1796
24161

24160
Upper Chocolate Clays (Lower part) |
24138

[White Mart Band —Ss=~—~—Ss—sSCiSC*dS

24133
Lower Chocolate Clays 754 |
24120
178

Platy Limestone Ce ae er 594
ee ; at least 1 at least | 24693
Shales with Alabaster 24113 pe

Part_of the Eocene Succession in the Northern Sulaiman Range.

FIG. 3

Tapti of Eames)

Upper Kirthar

(

Lower Kirthar
(Kirthar ss. of Eames)

! NiO! 7, BEsx. ‘P/O S*EeD

Middle Eocene Upper Eocene

Eoc

Lr

Sor Range. Samples were collected from the “‘ Claystones ”’ which are overlain by
fifty feet of conglomerates. The Ghazij Shales overlie the conglomerates. A chart
showing these lithological units is given (Fig. 6), and a detailed succession is given in
Appendix 1. “ The locality is in Lease 58 on the north slope of the Sor Range, about
eight miles by road east of Quetta (Survey of Pakistan Topo.Sheet No. 34 N/4, co-
ordinates 30° 11'20” N., 67° 10’ E, grid reference P 125210). Samples were collected
from a road cut along the main access road that crosses the lease approximately
parallel with the outcrop and along the contour of the slope ; structurally the locality

FROM WEST PAKISTAN

7 ial

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SHOYND
“—s

10 EARLY TERTIARY OSTRACODA

is near the northern end of the Sor Range syncline, which is the major structural
feature of the Sor Range-Danghari coalfield.”” (Reinemund, personal communi-
ication 1966).

Dr. F. T. Banner of University College, Swansea, was kind enough to examine
smaller foraminifera from sample 460-i. He has dated this horizon as the Upper
Palaeocene (pseudomenardi Zone).

IV. SYSTEMATIC DESCRIPTIONS
Subclass OSTRACODA Latreille 1806
Order PODOCOPIDA Miiller 1896
Suborder PODOCOPINA Sars 1866
Superfamily CYTHERACEA Baird 1850
Family TRACHYLEBERIDIDAE Sylvester-Bradley 1948

Diacnosis. Cytheracea with heavily calcified carapace, often highly ornamented
with more or less conspicuous eye-tubercle. Muscle scar pattern basically consisting
of four adductor scars (one or more vertically divided in some genera) with a
frontal scar which may be simple, V-Shaped, U-shaped or multiple. Posterior
characteristically sub-triangular or auricular, but in some genera produced to form a
caudal process. Subcentral-tubercle present or absent.

REMARKS. The classification adopted here is to retain the trachyleberids and the
hemicytherids in the family Trachyleberididae. Although in general trachyleberids
possess a subcentral-tubercle and a V-shaped frontal scar whilst hemicytherids
possess divided frontal and adductor scars, an auricular posterior end but lack the
subcentral-tubercle there still remain a large number of genera which tend to overlap,
thus making it impossible to clearly define the groups at the present time. Hazel
(1967) identified two families, the Trachyleberididae having six podomeres in the
antennule and the Hemicytheridae with five podomeres. This morphological
character is useless palaeontologically and considering the number of trachyleberids
which share hemicytherid characters (e.g. divided frontal scars) and hemicytherids
having a subcentral-tubercle it would appear optimistic to expect the number of
podomeres in the antennule to be so unique as to be restricted to only one group when
other, equally good morphological characters obviously are not.

Pokorny (1964) consideredgthe Hemicytherinae to be a group having a horizontal
classification, and the situation at the present time has not been effectively clarified.

Genus ACTINOCYTHEREIS Puri 1953
TYPE SPECIES. Cythere exanthemata Ulrich and Bassler 1904

Actinocythereis ? quasibathonica sp. nov.
(Plate 1, figs. 1-3, 6, 7, 10-13)

DERIVATION OF NAME. Latin quasibathonica, “‘ simulating Bathonian”’ ; with
reference to the resemblance to the Middle Jurassic (Bathonian) genus Oligocytherets.

30 10

6S
& Kawal
B Nawangaon 4

Q UETTA

4 ichari
OD B Nic

KASI

Traced from topo sheet No, 34/4,

Spin Lake

be Ush Bul N

Scale 1:50,000

1 2

Furlongs 8 6 4 2 t) 3 Miles

0 1 2 3 Kilometres

MAP OF PART OF SOR RANGE SHOWING FOSSIL LOCALITY, QUETTA DIVISION,

FIG.5

Spin Ghund 338

Wy See Aa
SS
SS
Sau
\ IN
\\ Sy
NY ’ =
Ne =
»)
Ny
\
\,
\
\
SO
Sy
Nh
1
Ny
HI
«

BIGUR Shin Maghzai

WEST PAKISTAN.

Road

Rcilway

Nala

Fossil

locality

7 aes

4
~~
» i ‘

——— =
ee
ors
ie
=
Oe
——

‘
= — a nn a ae ty a om - ~
: ibe J0at oh ets pao? aunt osoet
‘ 2
- ) — - _ —_- — ed

'.
i

A lie fours ’

4

- é .
P b Pa - ) pi N : j sii ; OG mi, 1 Lf Q JETTA
Py,

FROM WEST PAKISTAN
SOR RANGE SECTION
Measured by John A. Reinemund

Middle Eocene

Kirthar formation

") 1600 (approx )

Ghazij formation Lower Eocene

°o
25-0 °°
Ilo 0? 20075 ?

50 Ft

res Limestone
Claystone’
Conglomerate

Sandstone

Upper Palaeocene

Base not exposed
FIG. 6

12 EARLY TERTIARY OSTRACODA

DiaGnosis. Medium size, thick shelled. High and distinct anterior marginal
rim with posterior ornamentation. Surface sparsely punctate. Subcentral tubercle
prominent and rounded.

HootyPe. Io. 4311, a female carapace (PI. 1, Figs. 2, 3, 11, 12).
PARATYPES. Io. 4260 + Io. 3100-1.

MATERIAL. 29 specimens from the Rakhi Nala section from 8 horizons (sample
nos. 3610, to 3615, 3617, 3618 and 3620). 10 specimens from the Zao River section
from two horizons (sample nos. 24150 and 24154). GSP BM 2506-7.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 3611.

DESCRIPTION. Sexual dimorphy moderate, the females are higher and wider than
the males. Carapace sub-rectangular, medium size and thick shelled. Anterior
margin broadly rounded, postero-dorsal margin slightly concave, posterior extremity
and postero-ventral margin somewhat rounded. Dorsal and ventral margins almost
straight, slightly tapering towards the posterior. In lateral view the dorsal orna-
mentation over-reaches the dorsal margin. Valves almost equal. In dorsal view
the greatest width passes through the sub-central node. Fye-tubercle rounded,
prominent and situated below and slightly anterior to a well-developed anterior
cardinal angle. Anterior marginal rim high. Ventral and posterior marginal rims
less high. Swub-central tubercle prominent, rounded and distinct. Surface sparsely
punctate (punctation is not distinct in some specimens), with an alate ventral ridge
which slightly slopes upwards towards the posterior. The postero-dorsal process is
a blade-like projecting ridge over-reaching the dorsal margin and extending vertically
below for a short distance. A prominent mid-dorsal tubercle with a small tubercle
present in front. There is also a small tubercle posterior and at some distance from
the sub-central node. About 12 short marginal spines anteriorly, partly concealed
in external lateral view by the anterior marginal rim, and 5-6 spines posteriorly.
Radial pore canals few, simple and straight. Inner margin and line of concrescence
coincide. Duplicature moderately wide. Selvage well marked lying sub-peripheral
in left valve, but at some distance from the outer margin in right valve. There is a
fairly well-developed flange groove in right valve. Muscle scars unknown. Hinge
holamphidont with the details given below :

Hinge element Left Valve Right Valve
Anterior Socket Conical projecting tooth
Anteromedian Subconical tooth Deep socket

having a straight

anterior and a convex
posterior in dorsal

view
Posteromedian Denticulate bar Shallow locellate groove
Posterior Slightly elongate Pessular tooth

socket, open on
venter,

FROM WEST PAKISTAN 13

DIMENSIONS (mm).

iD H Ww
Io. 4260 Carapace male 0-51 0:29 0:27
Io. 4311 Carapace female (holotype) 0°52 0-32 0:29
Io.3101 Left valve male 0-51 0:29 —
Io.3100 Right valve male 0-51 0:29 —

REMARKS. This species is tentatively assigned to the genus Actinocythereis. It
differs from the type species of the genus in having a continuous rather than a
broken ventral ridge. In addition, the present species is much smaller, has a pitted
surface and fewer radial pore canals.

Genus ALOCOPOCYTHERE nov.

DERIVATION OF NAME. Greek alokos, = furrow, opos = eye ; with reference to the
furrow behind the eye-tubercle + cythere.

Diacnosis. Trachyleberididae in which the eye-tubercle is confluent with both
the elevated marginal rim and a short almost vertical ridge, delimited posteriorly by
a deep furrow. Anterior and posterior cardinal angles protruding in left valve, only
anterior cardinal angle protruding in right valve. Posterior cardinal angle of right
valve over-reached by protruding cardinal angle of left valve. Dorsal margin
humped.

TYPE SPECIES. Alocopocythere transcendens sp. nov.

DESCRIPTION. Dimorphic, the males are proportionally longer than the females.
Carapace sub-rectangular to sub-quadrate in shape. Dorsal margin in lateral view
sinuous, dominated by protruding anterior and posterior cardinal angles, with a
hump between, ventral margin evenly curved or almost straight. Anterior margin
broadly rounded, posterior straight or very slightly concave in postero-dorsal margin
(between posterior cardinal angle and posterior extremity) ; posterior extremity
rounded, postero-ventral region rounded or straight. Valves almost equal in size.
Sub-central tubercle and eye-tubercle more or less distinct. Surface ornamentation
either reticulate (with or without superimposed lineations or with superimposed
papillae) or papillose. A marginal rim always present, usually upstanding anteriorly,
less high along venter and posterior. Anterior and posterior margins ornamented
with small spines or denticles. Normal pores simple, widely spaced. Radial pore
canals simple, almost straight, often slightly inflated towards the middle, tending to
occur in groups of two or three, often apparently crossing one another, about 32-35
anteriorly and 18-20 posteriorly. Inner margin and line of concrescence coincide.
Duplicature of moderate width. Selvage well-marked—sub-peripheral in left valve
but at some distance from the outer margin in the right valve. Right valve with a
deep flange groove on the centre and anterior. Muscle scar pattern consists of four
adductors in a vertical row situated on the posterior margin of the muscle scar pit
and an oval frontal scar with two more or less rounded mandibular scars below.
Hinge holamphidont. Right valve with highly projecting, stirpate anterior tooth,

14 EARLY TERTIARY OSTRACODA

postjacent socket, posteromedian locellate groove and a pessular posterior tooth ;
left valve with anterior socket, anteromedian sub-conical tooth, postero-median
denticulate bar and a deep posterior socket.

CoMPARISON. This genus differs from Echinocythereis in having a short vertical
ridge below and a furrow behind the eye-tubercle, also there are two frontal scars in
Echinocythereis, but only one in Alocopocythere. Stigmatocythere has a curved ridge
joining the eye-tubercle and the sub-central tubercle, whereas in Alocopocythere a
short, almost vertical ridge joins the eye-tubercle and is delimited posteriorly by a
furrow. Henryhowella has three longitudinal plications in the posterior half of the
valve and an anterior vestibule, not present in Alocopocythere. Moreover, the
frontal scar in Henryhowella is V-shaped, while Alocopocythere has an oval frontal
scar.

RemARKS. In addition to the species described here, the Miocene species Tvachy-
leberis fossularis Lubimova and Guha (1960, p. 40, pl. 3, fig. 7), which Guha in 1961
(p. 4, figs. 5, 9) transferred to the genus Echinocythereis should be ascribed to Alo-
copocythere.

Alocopocythere transcendens sp. nov.
(Plate 1, figs. 4, 5, 8, 9 ; Plate 2, figs. 1-4, 6, 7)

DERIVATION OF NAME. Latin, transcendens, rising above ; with reference to the
stratigraphic position in relation to A. abstracta.

Diacnosis. Strongly reticulate Alocopocythere with rounded postero-ventral
margin, sub-central tubercle more or less distinct, eye-tubercle distinct, marginal
rim well marked.

HototyPe. Io. 4315, a female left valve (PI. 2, figs. 1, 6).
PARATYPES. Io. 4261 + Io. 3104-6.

MATERIAL. 263 specimens from the Zao River section from 7 horizons (sample
NOs. 24127, 24131, 24132, 24145, 24147, 24148 and 24151). Approximately 600
specimens from the Rakhi Nala section from 46 horizons (sample nos. 3168, 3198,
to 3200, 3401 to 3405, 3407, 3409, 3410, 3418, to 3422, 3424, 3426, 3428, 3429, 3432,
3434, 3435, 3438, 3457 to 3459, 3498, 3499, 3607, 3614, 3615, 3617 and 3618). GSP.
BM. 2508.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 24148.

DESCRIPTION. Carapace sub-rectangular to sub-quadrate in lateral outline.
Sexual dimorphism rather marked, the females being shorter, higher and wider than
the males. Dorsal margin sinuous with protruding anterior and posterior cardinal
angles, ventral margin almost straight in the right valve but evenly curved in the
left valve. Anterior margin broadly rounded, postero-dorsal margin very slightly
concave particularly in the right valve, posterior extremity rounded, postero-
ventral marginrounded. Valvesalmost equalinsize. Eye-tubercle distinct, rounded
and polished. Swb-central tubercle more or less distinct. Shell surface strongly

FROM WEST PAKISTAN 15

reticulate. Antero-dorsal furrow deep, bounded anteriorly by a short almost
vertical ridge joining the eye-tubercle. Anterior marginal rim high, continuing as a
less high rim round the venter and posterior. Anterior margin ornamented with
8-10 short spines, posterior with a postero-ventral spine, although these are pre-
served in a few specimens only. Duplicature moderately wide, 0-073 mm. anteriorly
in right valve female. Selvage prominent in both valves, situated in the outer third
of the duplicature in right valve but sub-peripheral in left valve. Along the venter
it is markedly concave antero-medially. Right valve has well-developed ventral
and anterior flange grooves. Normal pore canals simple, small. Radial pore canals
more or less straight, simple, some in groups of two or three, frequently crossing each
other. There are approximately 35 radial pore canals in the anterior and 20 in the
posterior. Line of concrescence and inner margin coincide throughout. Muscle
scars consist of sub-vertical row of four adductors, situated on the posterior margin
of the muscle scar pit, with an oval frontal scar and two somewhat rounded mandi-

bular scars below. Hinge holamphidont with the following details :

Element Left valve Right valve

Anterior Deep socket confluent Highly projecting
with ocular sinus, stirpate tooth, ocular
bounded on all sides. sinus lies below it.

Anteromedian Subconicaltoothwith Deep rounded socket
straight anterior and opening into postero-
convex posterior in median groove
dorsal outline.

Posteromedian Denticulate bar Locellate groove

Posterior Deep slightly elongate _—_ Pessular tooth, high

DIMENSIONS (mm).

socket open on ventral
side.

on posterior tending
towards reniform.

ie H W
Io. 3104 Carapace male 0-72 0°43 0-42
Io. 4315 Left valve female (holotype) 0:63 0-44 —
Io. 3105 Right valve female 0:64 0-39 —
Io. 4261 Left valve female 0:59 0-39 --
To. 3106 Right valve male 0:63 0:38 =
CoMPARISON. Alocopocythere abstracta sp. nov. is a very closely related species,

but is more elongate, and has a straight rather than rounded posteroventral margin
and less deep reticulations. Alocopocythere transcendens is perhaps ancestral to
Alocopocythere transversa sp. nov. but is smaller, and lacks the posterior concentric
ridges and a short ridge in the anteroventral area. Alocopocythere fossularis (Lubi-
mova and Guha) (1960) from the Miocene of Kutch is a similar species but which
differs however, in the lateral outline of the carapace.

REMARKS. Specimens of A. fossularis (Lubimova and Guha) from the type
locality in Kutch were not available for comparison.

16 EARLY TERTIARY OSTRACODA

Alocopocythere rupina sp. nov.
(Plate 2, figs. 5, 8-10 ; Plate 3, figs. 1-4)

ce

DERIVATION OF NAME. Latin, rupina,
dorsal furrow and associated ridges.

chasm ”’ ; with reference to the antero-

DriaGnosis. Alocopocythere in which anterodorsal furrow is delimited anteriorly
by a short almost vertical ridge and posteriorly by the anterior part of the dorsal
ridge. Surface reticulate with seven longitudinal ridges. Anterior and posterior
plains almost smooth.

HoLotyPe. Io. 4314, a male carapace (PI. 2, figs. 5, 8-10).
PARATYPE. Io. 4262.

MATERIAL. 4I specimens from the locality below from one horizon (sample no.
3111). GSP BM 2509-10.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Gorge Beds, sample no. 31II.

DESCRIPTION. Sexual dimorphism strong, the males are longer than the females.
Carapace subrectangular in lateral view. Dorsal margin sinuous, ventral margin
slightly concave in front of the middle, anterior margin broadly rounded, posterior
narrowly rounded. Anterior cardinal angle protruding particularly in right valve,
posterior cardinal angle less well-developed. Left valve slightly over-reaches right
valve at anterior cardinal angle and in the region of posterodorsal corner. Evye-
tubercle rounded and distinct. Swubcentral-tubercle well-developed. Surface orna-
mentation with seven longitudinal ridges ; the dorsal ridge begins above and very
slightly to the anterior of the subcentral-tubercle and is convex in the middle
culminating in the posterior quarter. The four ridges below the dorsal ridge are
almost confined posterior to the subcentral-tubercle, the second ridge from the centre
is the longest and is slightly curved ; it commences above the anteroventral corner
and slopes obliquely upwards towards the posterior ending in the posterior quarter.
The ventral ridge is confined in the posterior part of the carapace and is intercalated
between the ventral margin and the second ventral ridge, to which it is almost
parallel. Anterodorsal furrow well-developed, delimited on the anterior by a short
almost vertical ridge, and on the posterior by the anterior portion of the dorsal ridge.
Anterior and posterior platforms almost smooth, compressed. Anterior marginal
rim elevated, ventral and posterior marginal rims less high. Radial pore canals not
detectable. Duplicature moderate. Selvage well-marked ; it is submarginal in left
valve but in the outer third of the duplicature in right valve, which also has well-
developed anterior and ventral flange grooves. Hinge holamphidont with stirpate
anterior tooth in right valve.

DIMENSIONS (mm).

L H WwW
Io. 4314 Carapace male (holotype) 0-68 0°37 0°34
Io. 4262 Carapace female 0°59 0°37 0°34

CoMPARISON. A. vupina can easily be differentiated from other known species of

FROM WEST PAKISTAN 17

Alocopocythere by its anterodorsal groove, which is not only delimited by an anterior
ridge but by a posterior ridge as well.

REMARKS. This is so far the oldest known species of the genus Alocopocythere.
It occurs abundantly in one horizon (sample no. 3111) of the Gorge Beds of the Rakhi
Nala section, the male to female ratio being I : 3.

Alocopocythere abstracta sp. nov.
(Plate 3, figs. 5-11 ; Plate 4, fig. 1)

DERIVATION OF NAME. Latin abstvactus, separated, referring to the difficulty in
separating this species from A. tvanscendens because of the many intermediate
forms.

Dracnosis. A reticulate Alocopocythere with straight postero-ventral margin in
lateral outline. Subcentral-tubercle present but not prominent.

Hototyre. lo. 4312, a female carapace (PI. 3, figs. g-11) ; (Pl. 4, fig. 1).
PARATYPE. lo. 4263.

MATERIAL. Over 2600 specimens (including adults and juveniles) from the Rakhi
Nala section from 69 horizons (sample nos. 3147, 3152, 3153, 3157 to 3180, 3183,
3184, 3186 to 3191, 3193 to 3194, 3197 to 3200, 3401 to 3405, 3407, 409 J48Os S425
to 3424, 3426, 3428, 3429, 3432, 3434, 3435, 3438, 3443 and 3445). 5 specimens from
the Zao River section from one horizon (sample no. 24127). GSP BM 2511-2512.

TYPE Locality. Rakhi Nala section.

TYPE HORIZON. Upper Rakhi Gaj Shales, sample no. 3163.

DESCRIPTION. Carapace subrectangular in side view. Sexual dimorphism rather
pronounced ; the males are longer than the females. Dorsal margin sinuous, ventral
margin nearly straight. Anterior margin broadly and evenly rounded, postero-
dorsal margin very slightly concave ; posterior extremity rounded ; postero-ventral
margin straight. Anterior and posterior cardinal angles protruding. Valves more
orless equal. Eye-tubercle distinct. Subcentral-tubercle present but not pronounced.
Surface reticulate. Anterodorsal furrow deep, bounded anteriorly by a short
almost vertical ridge diagnostic of the genus. Anterior and posterior margins
denticulate, although the denticles are only present in a few specimens. Internal
details not known.

DIMENSIONS (mm).

iB H Ww
Io. 4263 Carapace male 0:66 0:38 0:34
To. 4312 Carapace female (holotype) 0°63 0:39 0°35

CoMPARISON. Alocopocythere coarctata sp. nov. is smaller than the present species
and has a combination of reticulations and weak ridges and a more sinuous dorsal
margin. Alocopocythere radiata sp. nov., however, is larger, has deeper reticulations
and a better developed subcentral-tubercle having posterior radial ridges.

A. abstracta has already been compared with Alocopocythere transcendens sp. nov.

B

18 EARLY TERTIARY OSTRACODA

Alocopocythere coarctata sp. nov.
(Plate 4, figs. 2-9)

DERIVATION OF NAME. Latin coarctatus, ‘‘ pressed together ’’ ; with reference to
the carapace.

Diacnosis. Alocopocythere in which carapace in lateral outline appears to be
compressed ; dorsal and ventral margins tapering towards the posterior end, sub-
central-tubercle distinct, anterior marginal rim high, surface finely reticulate (with
superimposed weak longitudinal ridges).

Ho.otyPe. Io. 4313, a female carapace (PI. 4, figs. 6-9).
PARATYPE. Io. 4264.

MATERIAL. 49 specimens from the below locality from five horizons (sample
NOS. 3432, 3434, 3435, 3458 and 3459). GSP BM 2513-4.
TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Shales with Alabaster, sample no. 3458.

DESCRIPTION. Carapace subrectangular to subquadrate in lateral view. Sexual
dimorphism strong ; the females are shorter than the males. Anterior margin
broadly and evenly rounded, posterodorsal margin straight, particularly in the left
valve, posterior extremity rounded, posteroventral margin rounded. Dorsal margin
sinuous with a hump between the protruding anterior and posterior cardinal angles,
ventral margin slightly concave in the middle. Both dorsal and ventral margins
taper towards the posterior. Valves almost equal. Surface finely reticulate with
superimposed weak longitudinal ridges. Swubcentral-tubercle distinct, eye-tubercle
more or less distinct. Marginal rim present, elevated in the anterior, less elevated
round the venter and posterior. Anterodorsal furrow fairly distinct and is bounded
anteriorly by a short almost vertical ridge. Anterior and posterior margins denticu-
late. Internal characters not known.

DIMENSIONS (mm).

ib H Ww
Io. 4264 Carapace male 0-51 0:27 0-27
Io. 4313 Carapace female (holotype) 0°50 0-32 0:29

CoMPARISON. Unlike A. coarctata, Alocopocythere rupina sp. nov. has better
developed longitudinal ridges and coarser reticulation. In addition, these two
species differ in lateral outline, particularly the male dimorphs. Alocopocythere
tvanscendens, sp. nov. is larger, has a less well-developed hump between the protruding
anterior and posterior cardinal angles and lacks longitudinal ridges.

Alocopocythere longilinea sp. nov.
(Plate 4, figs. 10-13 ; Plate 5, figs. 1-3, 6)

DERIVATION OF NAME. Latin longi, longitudinal + linea, line.
Dracnosis. A small Alocopocythere in which surface ornamentation is reticulate,

FROM WEST PAKISTAN 19

the reticulae being arranged in longitudinal lines with weak ridges in between,
subcentral-tubercle indistinct, marginal rim low, anterior marginal area compressed.

Hototype. Io. 4318, a male carapace (Pl. 4, figs. 10-13).
PARATYPE. Io. 4265.

MATERIAL. Nearly 670 specimens (including adults and juveniles) from the Rakhi
Nala section from 10 horizons (sample nos. 3438, 3440, 3443 to 3445, 3448, 3450, 3451,
3457 and 3458). One specimen from the Shpalai Khwara section from one horizon
(sample no. 24683). GSP BM 2515-6.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Shales with Alabaster, sample no. 3443.

DESCRIPTION. Carapace ovate in lateral outline and slightly tapering towards the
posterior. Sexual dimorphism marked ; the males are longer in proportion than the
females. Anterior margin broadly and obliquely rounded, somewhat compressed,
posterior almost straight, posteroventral margin slightly curved. Dorsal margin
sinuous, ventral margin evenly curved. Valves nearly equal. Swbcentral-tubercle
indistinct. Evye-tubercle low. Surface ornamentation consists of a combination of
reticulations and weak ridges. Anterodorsal furrow deep with a short more or less
vertical anterior ridge characteristic of the genus. Marginal rim low. Internal
characters unknown.

DIMENSIONS (mm).

L H WwW
Io. 4318 Carapace male (holotype) 0°54 0°32 0°25
Io. 4265 Carapace female 0-46 0-30 0-24

CoMPARISON. The present species differs from Alocopocythere abstracta sp. nov.
and Alocopocythere transcendens sp. nov. in being smaller and having weak longitudinal
ridges. Moreover, A. longilinea has a low marginal rim and an indistinct sub-
central-tubercle. Alocopocythere coarctata sp. nov. is about the same size but has a
high marginal rim, well-developed subcentral-tubercle and more sinuous dorsal
margin.

REMARKS. A. longilinea occurs in the lower part of the Shales with Alabaster
of the Rakhi Nala section and at several horizons it is very abundant. It is very rare
in the Shpalai Khwara section.

Alocopocythere transversa sp. nov.
(Plate 5, figs. 4, 5, 7-10 ; Plates 6-8 ; Plate 9, figs. 1-5)

DERIVATION OF NAME. Latin transversus, transverse ; with reference to the
posterior ridges.

Dracnosis. A species of the genus Alocopocythere with three posterior transverse
concentric ridges. A short ridge in the anteroventral area runs obliquely from the
anterior towards the venter, a shallow groove on the dorsal side of the ridge. Surface
reticulate (with or without superimposed papillae) or papillose.

HototyPe. Io. 4316, a female carapace (PI. 5, figs. 8, 10) ; (PI. 6, figs. 1, 2).

20 EARLY TERTIARY OSTRACODA

PARATYPES. Io. 4266-9 + Io. 3107-12.

MATERIAL. Over 800 specimens from the Zao River section from 20 horizons
(sample nos. 24131, 24155, 24157, 24159, 24170, 24173 to 24178, 24180, 24181, 24183
to 24188 and 24195). Approximately 300 specimens from the Rakhi Nala section
from 20 horizons (sample nos. 3624 to 3626, 3630, 3631, 3634, 3640 to 3642, 3645,
3646, 3648 to 3653, 3658 and 3660). GSP BM 2157-8.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24155.

DESCRIPTION. Sexual dimorphism rather marked, the males are more elongate
than the females. Cavapace subrectangular in lateral view. Dorsal margin sinuous,
ventral margin straight or evenly curved. Anterior margin broadly rounded,
posterodorsal margin very slightly concave, posterior extremity rounded, postero-
ventral margin rounded or almost straight. Anterior and posterior cardinal angles
well-developed and protruding. Valves nearly equal. Eye-tubercle distinct,
polished and rounded. Swbcentral-tubercle distinct. Surface either reticulate (with
or without superimposed papillae) or papillose. There are three posterior transverse
concentric ridges approximately parallel to the posterior margin with grooves in
between. There is a short ridge in the anteroventral area running obliquely from
the anterior towards the venter, with a groove on the dorsal side. Anterodorsal
groove fairly deep and bounded on the anterior by a short almost vertical ridge
running from the eye-tubercle. The marginal rim is high in the anterior but less high
on the venter and posterior. Anterior and posterior margins ornamented with short
spines, only present in some specimens and approximately 20 anteriorly and 10
posteriorly. Normal pore canals simple, small and numerous. Radial pore canals
numerous, simple, nearly straight, some in groups of two or three, often apparently
crossing one another. Duplicature moderately wide. Selvage pronounced in the
outer third of the duplicature in the right valve but sub-marginal in the left valve.
Right valve with deep ventral and anterior flange grooves. Line of concrescence and
inner margin coincide. Muscle scars consist of four adductors in a vertical row with
an oval frontal scar and two almost rounded mandibular scars below. Hinge

holamphidont :
Element Left valve Right valve
Anterior Deep rounded socket — Strongly projecting stirpate
bounded on all sides, —_ tooth (ocular sinus situated
confluent with ocular below and slightly anterior
sinus. to it).

Anteromedian Subconical projecting Deep socket.
tooth with a straight
anterior and convex
posterior in dorsal view.
Posteromedian Denticulate bar Locellate groove
Posterior Deep elongate socket Pessular tooth with a
unbounded on venter —__ tendency towards reniform,
higher on posterior.

FROM WEST PAKISTAN 21

CoMPARISON. Alocopocythere radiata sp. nov. is similar and perhaps related but
lacks the posterior concentric ridges. Moreover, A. radiata has ventral inflation
culminating in a ventral ridge and the longitudinal ridges radiate from the posterior
of a subcentral-tubercle.

The present species has already been compared with Alocopocythere transcendens
sp. nov.

REMARKS. This species may be divided into the following morphotypes, which
may represent the chronological subspecies of A. transversa. However, because of
the difficulty in separating these from one another and also owing to the fact that the
reticulate forms recur in the Upper Eocene succession of the Rakhi Nala and Zao
River sections, these are here considered as morphotypes.

MORPHOTYPE A
(Pl. 5, figs. 4, 5, 7-10 ; Pl. 6, figs. 1-4)
This has a reticulate surface. The reticulae are usually without any super-
imposed papillae but in some specimens a few small papillae at the junction of

reticulae are present. The posteroventral margin in lateral outline is curved in the
male but straight in the female.

DIMENSIONS (mm).

1g H WwW
Io. 4266 Carapace male 0-76 0°44 0°44
To. 4316 Carapace female (holotype) 0-71 0°45 0-44
Io.3107 Right valve male 0°85 0-46 —

MORPHOTYPE B

This comprises the transitional forms which fall between Morphotype A and
Morphotype C. It has slightly papillose reticulae. Specimens Io. 5004-5 from
sample 24155.

MORPHOTYPE C
(Plate 6, figs. 5-8 ; Plate 7, figs. 1-4 ; Plate 8, fig. 4)
This is similar to Morphotype B, but has a combination of reticulations and
papillae and a curved posteroventral margin in both male and female.
DIMENSIONS (mm).

L H WwW
Io. 4267 Carapace male 0-78 0-46 0°44
Io. 4268 Carapace female 0-76 0-46 0°44
To. 4269 Right valve male (broken) 0-80 — —

MorRPHOTYPE D

This includes the intermediate forms between Morphotype C and Morphotype E.
Specimens Io. 5006-7 from sample 24175.

22 EARLY TERTIARY OSTRACODA

MORPHOTYPE E
(Plate 7, figs. 5-8 ; Plate 8, figs. 1-3, 5)

This is similar in all characters to Morphotype A and Morphotype C but has a
papillose surface. It has a curved posteroventral margin in the male and female
dimorphs asin Morphotype C. There is a smooth, shallow groove on the dorsal side
of the ventral ridge.

DIMENSIONS (mm).

L H W
Io.3110 Carapace male 0-80 0-46 0-46

Io. 3111 Carapace female 0°74 0-46 0°45

MORPHOTYPE F
(Plate 8, figs. 6-9 ; Plate 9, figs. 1-5)

This has a small carapace. The surface is ornamented with slightly papillose
reticulae. There is a rim behind the anterior marginal rim and almost parallel to it
with reticulations in between. It originates from the eye-tubercle and fuses ventrally
with a short, oblique ventral ridge. It is likely that these forms may be juveniles of
Morphotype A or Morphotype C or may even belong to a distinct species.

DIMENSIONS (mm).

L H WwW
Io. 3109 Carapace male 0-68 0°39 0°39
Io. 3108 Carapace female 0:64 0°39 0:38
Io. 3112 Right valve female 0-59 0:37 —

Alocopocythere radiata sp. nov.
(Plate 9, figs. 6-9 ; Plate 10, figs. 1-4)
DERIVATION OF NAME. Latin vadiatus, rayed ; with reference to the ridges
radiating from the subcentral-tubercle.

Diacnosis. A coarsely reticulate Alocopocythere with longitudinal ridges radia-
ting from the posterior of a well-developed subcentral-tubercle. Eye-tubercle

distinct, marginal rim high, ventral inflation ends in a marked ridge, almost parallel
to the ventral marginal rim.

HoLotyre. Io. 4317, a male carapace (Plate 9, figs. 6, 8 ; Plate 10, figs. 1, 2)
PARATYPE. Io. 4270.

MATERIAL. 14 specimens from the locality below from one horizon (sample
no. 3652). 8 specimens from the Zao River section from one horizon (sample no.
24173). GSP BM 2519-20.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 3652.
DESCRIPTION. Sexual dimorphism distinct ; the females are shorter than the

FROM WEST PAKISTAN 23

males. Carapace subrectangular in lateral view. Dorsal margin sinuous with
protruding anterior and posterior cardinal angles ; ventral margin almost straight.
Anterior margin broadly and evenly rounded, posterior extremity rounded, postero-
dorsal margin very slightly concave, posteroventral margin curved in the male
dimorph but almost straight in the female. Valves more or less equal. Evye-
tubercle rounded and distinct. Surface ornamentation consists of coarse reticula-
tions with superimposed ridges radiating from the posterior of a well-developed
subcentral-tubercle. The ventral inflation culminates in a marked ventral ridge
almost parallel to the ventral marginal rim. There are two ridges which join the
eye-tubercle, one a short more or less vertical ridge bounded posteriorly by a deep
anterodorsal furrow which is better seen in dorsal view, and the other a high anterior
marginal rim which continues along the venter and around the posterior as a less high
rim. Anterior and posterior margins decorated with numerous very short and
delicate spines.

DIMENSIONS (mm).

L H WwW
Io. 4317. Carapace male (holotype) 0-72 0-42 0-42
Io. 4270 Carapace female 0-68 0°42 0-40

CoMPARISON. Alocopocythere transcendens sp. nov. shows some resemblance and
is perhaps ancestral to the present species ; but A. tvanscendens has a less well-
developed subcentral-tubercle without radial ridges and lacks a ventral inflation
ending inaridge. Alocopocythere coarctata sp. nov. is much smaller, has a carapace
which tapers towards the posterior end and has a less deep surface reticulation.

Genus “ ANOMMATOCYTHERE ” Sohn

TYPE SPECIES. “‘ Anommatocythere microreticulata ’’ Sohn.

REMARKS. This is a new genus erected by Sohn whose paper is in press. The
two species described below are provisionally assigned to the genus but their final
designation will depend on the publication of Sohn’s paper.

“ Anommatocythere ” laqueata sp. nov.
(Plate ro, figs. 5-10)
DERIVATION OF NAME. Latin laqueatus, fluted ; with reference to the ornamenta-
tion of the anterior rim.

Diacnosis. Anterior rim ornamented with seven small more or less rectangular
depressions. Carapace subtriangular with a gently convex dorsal margin.

HOLOTYPE. Io. 4320, a female carapace (Plate 10, figs. 8-10).
PARATYPE. lo. 4271.

MATERIAL. 32 specimens from the locality below from three horizons (sample
nos. 3403, 3405 and 3466). Two specimens from the Zao River section from one
horizon (sample no. 24107). GSP BM 2521-2.

24 EARLY TERTIARY OSTRACODA

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Green and Nodular Shales, sample no. 3403.

DESCRIPTION. Sexual dimorphy rather apparent ; the males are longer in pro-
portion than the females. Carvapace subtriangular in lateral view. Dorsal margin
gently convex, ventral margin almost straight, anterior margin broadly rounded,
posterior with a caudal process. Greatest length lies below mid-point, greatest
height at anterior cardinal angle. Anterior and posterior cardinal angles more or
less rounded, somewhat better developed in the left valve. Left valve very slightly
larger than the right valve, over-reaching it in the anterodorsal corner and postero-
dorsal slope. Swubcentral-tubercle indistinct. Eye-tubercle distinct but low. Surface
reticulate, the reticulae being arranged in lines separated by longitudinal ribs.
Anterior marginal rim ornamented with seven small rectangular depressions like a
scallop or bivalve mollusc. Internal characters not observed.

DIMENSIONS (mm).

L H W
To. 4271 Carapace male 0:66 0:37 0°32
Io. 4320 Carapace female (holotype) 0:66 0°39 0°34
COMPARISON. “‘ Anommatocythere’’ confirmata sp. nov. differs from the present

species by having a thick-shelled and ventrally inflated carapace, a convex rather
than straight ventral margin in lateral view, and no ornamentation of the anterior
rim.

REMARKS. Specimens from the Shales with Alabaster show fainter longitudinal
ribs. This is perhaps due to the form of preservation.

“ Anommatocythere ” confirmata sp. nov.
(Plate ro; figs. nxyn2/; Plater. ;¢Plate, aayifigs, 1, 2)
DERIVATION OF NAME. Latin confirmatus, “strengthened ’’ ; with reference to
the variation in the strength of the longitudinal ribs.

DiaGnosiIs. Carapace ventrally inflated and with a short caudal process. Ventral
longitudinal ribs are curved and better developed. Anterior and posterior cardinal
angles well-marked.

HototypPe. Io. 4319, a male carapace (PI. ro, figs. 11, 12) ; (Pl. 11, figs. 1, 2)

PARATYPE. Io. 4272 + Io. 3102-3.

MATERIAL. 70 specimens from the Rakhi Nala section from five horizons (sample
nos. 3499, 3611, 3613-3615). 53 specimens from the Zao River section from six
horizons (sample nos. 24145, 24147, 24148, 24150 to 24152). GSP BM 2523-4.

TYPE LocALiITy. Rakhi Nala section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 3611.

DESCRIPTION. Sexual dimorphism rather marked, the females are shorter and
wider in proportion than the males. Carapace plump, thick-shelled and with ventral
inflation. Dorsal margin slightly convex particularly in the right valve, ventral

FROM WEST PAKISTAN 25

margin anteromedially concave but is convex in lateral outline due to the ventral
inflation ; anterior margin broadly rounded, posterior with a short caudal process.
Anterior and posterior cardinal angles well-developed particularly in the left valve.
Left valve slightly over-reaches the right at the anterodorsal and posterodorsal
corners. Swbcentral-tubercle present but not distinct. Evye-twbercle rounded, shiny
and distinct and lies below and slightly anterior to cardinal angle. Surface ornamen-
tation consists of reticulations and longitudinal ribs. There is a variation in the
strength of the longitudinal ribs, those on the ventral surface are stronger and curved
convexly downwards in the middle. Marginal rim narrow and low. Anterior and
posterior margins denticulate. Valves deep in internal view. Normal pore canals
fairly numerous and perhaps each reticule has one normal pore canal. Radial pore
canals simple, straight, sparse, irregularly spaced, few crossing one another, approxi-
mately 20 anteriorly and 8 posteriorly. Line of concrescence and inner margin
coincide—no vestibule. Duplicature fairly wide—o-073 mm. anteriorly, 0-055 mm. on
the posterior extremity. Selvage distinct and subperipheral ; situated in the right
valve on the outer sixth of the anterior margin. Right valve with a ventral flange
groove between selvage and flange. Muscle scar pattern with four adductor scars in
an almost vertical superposition at the posterior margin of the muscle scar pit and
two more or less rounded frontal scars. Hinge holamphidont with the following
details of the hinge elements :

Element Left valve Right valve

Anterior Deep socket bounded Strongly projecting
on all sides, eye- stirpate tooth.
socket lies almost in
the middle of it.

Anteromedian Conical tooth, Deep socket narrowing
projecting slightly posteriorly into a long
towards anterior. groove.

Posteromedian Denticulate bar Locellate groove, the
anterior part deeper than
posterior.

Posterior Deep and elongate Large bilobate tooth, the

socket unbounded on
ventral side.

anterior lobe lower than
the posterior.

DIMENSIONS (mm).

L H WwW
Io. 4319 Carapace male (holotype) 0-66 0-39 0°39
To. 4272 Carapace female 0:63 0°42 0-44
Io. 3102 Left valve male 0-64 0°39 =
To. 3103 Right valve male 0:63 0°37 —
COMPARISON. This species has already been compared with “ Anommatocythere”’

laqueata sp. nov.

REMARKS. The vertical range of the present species in the Rakhi Nala and Zao

26 EARLY TERTIARY OSTRACODA

River sections is 286 ft. and 378 ft. respectively. Hence, it is a very useful species
as a horizon marker in the region.

Adult specimens in the two sections vary in size and in the strength of
ornamentation.

Genus BRADLEYA Hornibrook 1952

TYPE SPECIES. Cythere arata Brady 1880.

Bradleya ? voraginosa sp. nov.
(Plate 12, figs. 3-9)

DERIVATION OF NAME. Latin voraginosus, full of pits.

DiaGnosis. A species provisionally placed in the genus Bradleya with sub-
parallel dorsal and ventral margins, projecting anterior cardinal angle, truncated
posterior, coarsely and deeply reticulate surface.

HoLotyPe. Io. 4321, a male carapace (PI. 12, figs. 3, 5, 7, 8).
PARATYPE. Io. 3115.

MATERIAL. 10 specimens from the locality below from two horizons (sample
nos. 24159 and 24161). GSP BM 2525.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24161.

DESCRIPTION. Cavapace subrectangular in lateral outline. Valves ventrally
inflated. Dorsal and ventral margins almost straight and subparallel, anterior
margin broadly rounded, posterior truncated, posterodorsal slope very slightly
concave particularly in the right valve. Anterior cardinal angle projecting, posterior
cardinal angle rather prominent (approximately 110°). Valves almost equal. Evye-
tubercle rounded and distinct and situated just below the anterior cardinal angle.
Subcentral-tubercle more or less distinct. A marginal rim runs around the anterior,
ventral and posterior margins. It is fairly well-developed anteriorly and posteriorly
but is not so prominent along the venter. Surface ornamentation consists of
coarse, deep reticulations and dorsal and ventral ridges.

The dorsal ridge is ill-defined in the anterior half, slightly arched upward in the
posterior third culminating in a short horn-like posterodorsal process. The ventral
ridge is better developed and slightly alate posteriorly. Anterior margin finely
denticulate, posteroventral margin ornamented with 4-5 short spines. Internal
details not very well displayed. Duplicature fairly wide. Selvage in the left valve
is subperipheral and less well-developed than in the right valve where it is at some
distance from the outer margin. It has a deep flange groove, particularly in the
venter. Hinge holamphidont ; left valve with a deep almost rounded anterior
socket which is bounded on all sides, a conical projecting anteromedian tooth, an
apparently denticulate bar and a deep elongate posterior socket which is bounded
on the venter. Hinge of right valve not clearly seen.

FROM WEST PAKISTAN 27

DIMENSIONS (mm).

L H WwW
Io. 4321 Carapace male (holotype) 0-76 0-42 0-42
Io. 3115 Carapace female 0°73 0°40 0-42

CoMPARISON. Bradleya ? cornuelina (Bosquet) Keij (1957) is similar to B?
voraginosa in lateral view but has three rather than two longitudinal ridges. Further,
it has a less well-developed anterior cardinal angle. Bradleya approximata (Bosquet)
Keij (1957) has a different posterior and larger posteroventral spines.

Genus BUNTONIA Howe 1935

TYPE SPECIES. Buntonia shubutaensis Howe 1935.

Buntonia devexa sp. nov.
(Plate 13, figs. 1-5)

DERIVATION OF NAME. Latin devexus, sloping ; with reference to the tapering
lateral outline.

Diacnosis. A species of Buntonia in which carapace is elongate, subrectangular
in lateral view ; surface ornamented with 9-11 longitudinal ribs in posterior three-
fifths of carapace.

Hototyre. lo. 4322, a female carapace (PI. 13, figs. 2, 4, 5).

PARATYPE. lo. 3113.

MATERIAL. 8 specimens from the locality and horizon below.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Gorge Beds, sample no. 311T.

DESCRIPTION. Sexual dimorphism rather pronounced ; the males are longer and
less wide than the females. Carvapace sub-triangular in lateral view, tapering
towards the posterior. Anterior margin broadly and obliquely rounded, posterior
narrowly rounded, dorsal and ventral margins almost straight, dorsal margin slopes
downwards towards posterior. Greatest length passes through the midpoint,
greatest height in the anterior third and greatest width in the posterior two-fifths.
Anterior cardinal angle rounded. Left valve slightly larger than the right valve.
Surface ornamentation consists of 9-11 longitudinal ridges, which are more or less
confined in the posterior three-fifths. Anterior marginal rim distinct.

DIMENSIONS (mm).

L H WwW
Io. 3113. Carapace male 0-80 0:39 0:24
Io. 4322 Carapace female (holotype) 0°73 0°35 0:27

COMPARISON. Buntonia virgulata Apostolescu (1961) has punctae between
longitudinal ridges and a less elongate carapace. Cythere cf. costellata (Roemer)
Latham (1938) is similar and may even be conspecific. However, her figure, which

28 EARLY TERTIARY OSTRACODA

appears to be drawn upside down, shows longitudinal ridges continuing in the
anterior part of the carapace.

REMARKS. Cythere costellata (Roemer) is now regarded as a species of the genus
Cytheretta. Because of the imperfect preservation, it has not been possible to observe
whether the present species has any eye-tubercles.

Buntonia Sp.A
(Plate 13, figs. 6, 7, 9)

FIGURED SPECIMEN. lo. 3114.

MATERIAL. Two specimens from the locality and horizon below.

LocaLity. Rakhi Nala section.

Horizon. Lower Rakhi Gaj Shales, sample no. 3133.

DESCRIPTION. Carapace small, almost triangular in lateral outline. Greatest
length lies below mid-point, greatest height in anterior third. In dorsal view the
carapace is widest just posterior to the middle and tapers towards anterior and
posterior ends ; posterior pointed. Anterior end broadly and obliquely rounded,
posterior narrow and somewhat rounded ; dorsal and ventral margins taper towards
the posterior. Anterior cardinal angle rounded and well-developed. Left valve
larger than right valve. Surface ornamented with some ten longitudinal ridges.

DIMENSIONS (mm).

L H Ww
Io. 3114 Carapace 0:50 0:27 0-32

CoMPARISON. Buntonsa devexa sp. nov. (Pl 13, figs. I-5) is much larger, has a less
triangular carapace with a gentle slope on the dorsal margin. Further, B. devexa has
a rounded rather than pointed posterior in dorsal view and has longitudinal ridges
which do not continue towards the anterior.

Genus COSTA Neviani 1928

DiaGnosis. Trachyleberididae in which ornamentation is dominated by three or
four longitudinal ridges, the median or second ridge running back from the sub-
central-tubercle towards posterodorsal corner in anterior two-thirds of length, then
curving sharply down towards posteroventral corner in posterior third of length.

TYPE SPECIES. Cytherina edwards Roemer 1838.

Subgenus COSTA sensu stricto

DiaGnosis. Costa with three longitudinal ridges.

Subgenus PARACOSTA nov.

DERIVATION OF NAME. Greek fara, near ; with reference to the strong resem-
blance to the subgenus Costa.

FROM WEST PAKISTAN 29

Diacnosis. Costa with a fourth ventral ridge intercalated between third ridge
and ventral margin.

TYPE SPECIES. Costa (Paracosta) dechivis sp. nov.

ReMARKS. The subgenus Paracosta is so far only known from the Rakhi Nala
section. It is represented by two species in the Upper Chocolate Clays and one
species in the Pellatispira Beds.

Costa (Paracosta) declivis sp. nov.
(Plate 13, figs. 8, 10-14 ; Plate 14, figs. 1, 2)

DERIVATION OF NAME. Latin declivis, sloping downward ; referring to the
direction of the ridge running anteroventrally from the subcentral-tubercle.

Diacnosis. A small species of Pavacosta in which longitudinal ridges are well-
developed, median or second ridge runs anteroventrally from subcentral-tubercle.

HoLotype. lo. 4325, a male carapace (pl. 13, figs. 8, 10-12).
PARATYPES. Io. 4273—Io. 3116.

MATERIAL. 34 specimens from the Rakhi Nala section from four horizons (sample
nos. 3661 to 3664). GSP BM 2526-7.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Pellatispira Beds, sample no. 3662.

DESCRIPTION. Sexual dimorphism marked, the males are longer than the females.
Carapace elongate, subrectangular in lateral view with greatest height at anterior
cardinal angle. Dorsal and ventral margins almost straight, subparallel, anterior
end broadly rounded, posterior subtriangular. Valves almost equal. Anterior
cardinal angle rounded, posterior cardinal angle obtuse. Greatest width in the
posterior third. Swbcentral-tubercle distinct. Evye-tubercle rounded and distinct.
Ornamentation consists of reticulations dominated by four longitudinal ridges.
The dorsal ridge commences just above the subcentral-tubercle and is slightly
arched upward (in lateral view over-reaching dorsal margin), the median or second
ridge runs almost diagonally from anteroventral margin towards posterodorsal
corner, then bending down towards posteroventral margin ; the ventral or fourth
ridge (better seen in ventral view) lies between the third ridge and the ventral
margin and is not as well-developed as the other three. Anterior and posterior
marginal rims high. Anterior margin ornamented with 15-18 small spines, postero-
ventral margin with 5-6 relatively large spines. Internal characters unknown.

DIMENSIONS (mm).

Iv, H W
Io. 4325 Carapace male (holotype) 0:83 0:39 0:37
Io. 4273, Carapace female O77 0°39 0:37
Io. 3116 Carapace female 0°77 0°39 0:37

CoMPARISON. This species differs from Costa (Paracosta) disintegrata sp. nov. in
having well-developed longitudinal ridges and a different outline. Costa (Paracosta)

30 EARLY TERTIARY OSTRACODA

compitalis sp. nov. is larger, lacks well-developed longitudinal ridges and has a ridge
running from the eye-tubercle to the subcentral-tubercle.

Costa (Paracosta) compitalis sp. nov.
(Plate 14, figs. 3-10)

DERIVATION OF NAME. Latin compitalis, pertaining to a cross-roads ; referring
to the nexus of the ridges running from the subcentral-tubercle.

DiaGnosis. A large, strongly reticulate species of the subgenus Paracosta in
which longitudinal ridges are moderately developed, subcentral-tubercle prominent,
joined by three ridges—dorsal, median and a ridge running from eye-tubercle.

HototyPe. Io. 4323, a female carapace (PI. 14, figs. 5, 6, 9, 10).
PARATYPE. Io. 4274.

MATERIAL. 14 specimens from the locality below from one horizon (sample
no. 3604). GSP BM 2528-9.

TYPE LOCALITY. Rakhi Nala section.
Type Horizon. Upper Chocolate Clays, sample no. 3604.

DESCRIPTION. Carapace subrectangular in lateral outline. Sexual dimorphism
rather pronounced ; the females are shorter and higher than the males. Dorsal
margin slightly curved in lateral view because of over-reaching by the dorsal ridge ;
ventral margin straight, anterior broadly and evenly rounded, posterodorsal margin
very slightly concave, posterior extremity slightly subtriangular, posteroventral
margin rounded. Greatest height at anterior cardinal angle, greatest length
through the mid-point and greatest width in posterior third. Anterior cardinal
angle well-developed with a concavity behind. Left valve slightly larger than right
valve, over-reaching at posterodorsal margin and in the region of anterior cardinal
angle. Eye-tubercle rounded and distinct and confluent with the anterior marginal
rim and a ridge running from the subcentral-tubercle. The prominent subcentral-
tubercle lies more or less in the anterior third. Surface coarsely reticulate. There
are four longitudinal ridges ; the dorsal ridge commences at the subcentral-tubercle
and is curved convexly upward, the median or second ridge stretches back from the
subcentral-tubercle towards posteroventral margin, the third ridge slightly slopes
upward towards the posterior end and the ventral ridge is almost parallel to the
ventral margin and is intercalated between the third ridge and the ventral margin.
Anterior and posterior marginal rims distinct. Anterior and posterior margins
ornamented with numerous small spines. Internal details not known.

DIMENSIONS (mm).

L H WwW
lo. 4274 Carapace male 0:98 0-51 0:46
Io. 4323 Carapace female (holotype) 0:93 0°51 0°44

COMPARISON. Costa (Paracosta) disintegrata sp. nov. is smaller than the present
species, has ill-defined longitudinal ridges and a carapace tapering towards the
posterior end.

FROM WEST PAKISTAN 31

Costa (Paracosta) disintegrata sp. nov.
(Plate 14, figs. 11 ; Plate 15, figs. 1-6)

,

DERIVATION OF NAME. Latin disintegratus, ‘‘ broken down”? ; referring to the

relict nature of the ridges characteristic of Costa.

Dracnosis. Paracosta of medium size with weakly developed longitudinal ridges.
Carapace tapering towards posterior end in lateral view.

Ho.otyPe. Io. 4324, a male carapace (PI. 14, figs. 11 ; Pl. 15, figs. 3, 4).

PARATYPE. Io. 4275.

MATERIAL. Four specimens from the locality below from two horizons (samples
no. 3621 and 3622). GSP BM 2530-31.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 3622.

DESCRIPTION. Sexual dimorphy moderate ; the males are longer in proportion
than females. Carapace subrectangular, tapering to posterior in side view. Dorsal
and ventral margins almost straight, anterior margin broadly and evenly rounded,
posterior subtriangular. Anterior cardinal angle rounded, posterior cardinal angle
obtuse—well-developed in left valve. Left valve over-reaches the right slightly at
the anterior cardinal angle and in the region of the posterodorsal slope. A distinct
eye-tubercle lies just below the anterior cardinal angle. Swbcentral-tubercle well-
developed. Anterior and posterior marginal rims high. Surface coarsely reticulate
(some reticulae being slightly papillose). There are four ill-defined longitudinal
ridges ; the dorsal ridge commences just behind the subcentral tubercle and is curved
convexly upward ; the second ridge stretching back from the subcentral-tubercle
towards the posterodorsal corner in the anterior two-thirds and then bends sharply
round towards the posteroventral corner ; the third ridge commences below the
subcentral-tubercle and slopes upward towards the posterior end ; the fourth ridge
(better seen in ventral view) is more or less parallel to the third ridge and lies between
the ventral margin and the third ridge. Anterior and posterior margins spinose.

DIMENSIONS (mm).

L H W
Io. 4275 Carapace male 0°83 0:42 0:32
Io. 4324 Carapace female (holotype) 0°85 0-44 0:37

ComPARISON. This species falls between Costa (Paracosta) compitalis sp. nov. and
Costa (Paracosta) declivis sp. nov. in size and stratigraphical position and has already
been compared with these species.

Genus ECHINOCYTHEREIS Puri 1954

Diacnosis. Trachyleberididae with or without ventral ridges. Carapace often
inflated and with curved posteroventral margin, particularly in right valve. Surface
ornamented with papillae, nodes, reticulations (or combination of these—con-

32 EARLY TERTIARY OSTRACODA

centrically arranged in some species) or almost smooth. Muscle scars are in a
vertical column of four adductors with two frontal scars.

TYPE SPECIES. Cythereis garettt Howe and McGuirt 1935.

Subgenus ECHINOCYTHEREIS sensu stricto

Diacnosis. Echinocythereis without ventral ridges.

Echinocythereis (Echinocythereis) contexta sp. nov.
(Plate 15, figs. 7, 8, 10, 13)

DERIVATION OF NAME. Latin contextus, joined together ; from the ornamentation
of the papillae joined by the walls of the reticulae.

Diacnosis. A species of the subgenus Echinocythereis in which posterior end is
obliquely rounded towards posterodorsal corner, eye-tubercle prominent, surface
reticulate with superimposed papillae.

HootyPe. Io. 4326, a female carapace (Pl. 15, figs. 8, 13).

PARATYPE. Io. 4276.

MATERIAL. Five specimens from the Sor Range section from four horizons
(sample nos. 460-f, 460-1, 460-j and 460-0). GSP BM 2532-3.

TYPE LOCALITY. Sor Range section.

TYPE HORIZON. Upper Palaeocene, sample no. 460-1.

DESCRIPTION. Sexual dimorphism rather strong ; the carapace is subrectangular in
the male and subquadrate in the female. Dorsal margin in lateral outline un-
dulating because of ornamentation, ventral margin almost straight, anterior broadly
and evenly rounded. Greatest length passes above the mid-point, greatest height
in the anterior fourth and greatest width behind the middle. Anterior and posterior
cardinal angles well-developed. Left valve larger than the right, over-reaching
it at the anterior, ventral and posterodorsal margins. Eye-tubercle rounded and
prominent, standing out from the shell surface in lateral and dorsal views. Sub-
central-tubercle more or less distinct. Surface ornamentation consists of slightly
papillose reticulae which are concentrically arranged near the margins. Anterior
and posterior margins are set with a double row of papillae ; those on the posterior
are larger, and in some specimens become short spines. Internal details not seen.

DIMENSIONS (mm).

L H WwW
Io. 4276 Carapace male 0-78 0°44 0°37
Io. 4326 Carapace female (holotype) 0-71 0-46 0:42

CoMPARISON. Unlike E. (E.) contexta sp. nov. Echinocythereis (Scelidocytherets)
sp.A has a straight rather than curved posterodorsal margin and less prominent
eye-tubercles. Moreover, it has a weak ventral ridge and a short horn-like postero-
dorsal process. Echinocythereis (Echinocythereis) elongata sp. nov. also differs by
possessing a very elongate carapace and a better developed subcentral-tubercle.

FROM WEST PAKISTAN 33

Echinocythereis (Echinocythereis) elongata sp. nov.

(Plate 15, figs. 6, 11, 12, 14.; Plate 16, figs. 1, 2)

DERIVATION OF NAME. Latin elongatus, elongate ; with reference to the carapace.

Diacnosis. An elongate species of the subgenus Echinocythereis in which posterior
end is rounded towards posterodorsal corner, subcentral-tubercle distinct, surface
ornamented with reticulae and papillae.

HototyrPe. Io. 4327, a female carapace (Pl. 15, figs. 12, 14 ; Pl. 16. fig. 1).
PARATYPE. lo. 3130.

MATERIAL. Nine specimens from the Rakhi Nala section from three horizons
(sample nos. 3404, 3409 and 3416). GSP BM 2534.

TYPE LocALiTy. Rakhi Nala section.
TYPE HORIZON. Rubbly Limestones, sample no. 3416.

DESCRIPTION. Carapace elongate, subrectangular in lateral outline. Sexual
dimorphism rather prominent in dorsal view ; males longer and less wide than females.
Anterior margin broadly rounded, posterior narrowly rounded towards postero-
dorsal corner particularly in the left valve, dorsal margin irregular due to ornamenta-
tion, ventral margin more or less straight. Greatest length lies above the middle,
greatest height at the anterior cardinal angle and greatest width in front of the
middle. Valves almost equal. Swbcentral-tubercle distinct. LEvye-tubercle fairly
distinct, but worn in some specimens. Surface ornamentation a combination of
reticulations and papillae. The reticulae are in the anterior and ventral regions and
the papillae in the middle and posterior. The papillae are perhaps revealed by the
removal of an upper layer of reticulae. The decorated papillae show normal pore
canals and nexus of reticulae, which are smaller than the anterior and ventral ones.
Normal pore canals are situated between the papillae. Anterior and posterior
margins denticulate, although the denticles are not preserved in some specimens.
Internal characters unknown.

DIMENSIONS (mm).

IC, H W
Io. 3130 Carapace male 0:73 0°39 0:29
Io. 4327. Carapace female (holotype) O-71 0:38 0°34

COMPARISON. This species can easily be separated from the other known species
of the subgenus Echinocythereis by its much more elongate carapace:

Subgenus SCELIDOCYTHEREIS nov.

DERIVATION OF NAME. Greek skelis, rib ; with reference to the development of
the ventral ridges.

Diacnosis. Echinocythereis with ventral ridges.

Cc

34 EARLY TERTIARY OSTRACODA

TYPE SPECIES. Echinocythereis (Scelidocythereis) multibullata. sp. nov.

Echinocythereis (Scelidocythereis) multibullata sp. nov.
(Plate 16, figs. 3-9 ; Plate 17, figs. 1, 2, 7)

DERIVATION OF NAME. Latin multus, much + bullatus, knobbed ; with reference
to the ornamentation.

Diacnosis. A species of the subgenus Scelidocythereis with a prominent sub-
central-tubercle consisting of 4-5 small nodes. Surface nodose or tuberculate.
Right valve over-reaches left valve anteriorly but is over-reached by the latter at
anterior and posterior cardinal angles.

HoLotyPe. Io. 4328, a male carapace (PI. 16, figs. 3, 5, 6 ; Pl. 17, fig. 7).
PARATYPES. GSP BM 2558-Io. 3133-4 + Io. 4277.

MATERIAL. 76 specimens from the Zao River section from five horizons (sample
NOS. 24154, 24156, 24159, 21461 and 24183). 28 specimens from the Rakhi Nala
section from three horizons (sample nos. 3621, 2624 and 2625). GSP BM 2535-6.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24161.

DESCRIPTION. Sexual dimorphism rather marked ; presumed females shorter,
higher and wider than presumed males. Cavapace subrectangular in lateral view.
Anterior margin broadly and evenly rounded in both valves. In the right valve the
posterodorsal corner is very slightly concave, whilst the posterior extremity and the
posteroventral margins are rounded ; in the left valve the posterior end is truncated.
Dorsal margin intricate in lateral view because of ornamentation ; ventral margin
slightly incurved anterior to the middle in the right valve but curved convexly down-
ward in the left valve. Greatest length passes through the middle, greatest height
at the anterior cardinal angle and greatest width in front of the middle (i.e. at the
subcentral-tubercle). Anterior cardinal angle protruding. Right valve over-
reaches the left at the anterior margin and posteroventral margin ; but left valve
over-reaches the right in the region of the anterior and posterior cardinal angle.
Subcentral-tubercle prominent and is composed of 4-5 small nodes. Evye-tubercle
rounded and distinct. Surface ornamented with nodes, or tubercles, those nearest
the ventral margin being the larger. There are three small ventral ridges, the two
near the ventral margin are smaller and almost confined to the anteroventral and
mid-ventral regions. Anterior and posterior margins are denticulate. Viewed
internally the valves are deep. Duplicature fairly wide, 0-11 mm. at the posterior
extremity (Pl. 16, fig. 9). Seluage well-developed, subperipheral in the left valve but
almost at the outer third in the right valve. A deep flange groove, better developed
at the venter, lies between the selvage and flange in the right valve. Radial pore
canals fairly numerous, simple, almost straight, few occurring in groups of two or
three. Inner margin and line of concrescence coincide. Muscle scars (best seen in
weathered specimens from the outside) are in an almost vertical column of four
adductors and two more or less rounded frontal scars. Hinge holamphidont:

FROM WEST PAKISTAN 35

Element Left valve Right valve

Anterior Deep, almost rounded Highly projecting
socket bounded on all __ pessular tooth.
sides.

Anteromedian Projecting subconical Socket opening into
tooth. posteromedian groove.

Posteromedian Slightly projecting Locellate groove.

denticulate ridge (den-
ticles are seen only in
nicely preserved
specimens).
Posterior Deep elongate socket Large subpessular tooth,
unbounded on venter. less high on anterior.

DIMENSIONS (mm).

L H WwW
Io. 4328 Carapace male (holotype) 0°85 0-50 0°45
Io. 3134 Carapace female 0:83 0-51 0:48
Io. 3133 Left valve female 0-84 0°54 —
Io. 4277. Right valve female 0°83 0:50 —

COMPARISON. LEchinocythereis (Scelidocythereis) sparsa sp. nov. is smaller than the
present species, has a different lateral outline, distinct marginal rims and scattered
tubercles as surface ornamentation. In addition to this, it has an indistinct rather
than prominent subcentral-tubercle.

REMARKS. This species occurs in the Upper Chocolate Clays of the Zao River and
Rakhi Nala sections. It has a short vertical range and hence can be used as an index
marker.

Echinocythereis (Scelidocythereis) sp.A
(Plate 17, figs. 3, 4, 8, 9)
FIGURED SPECIMEN. Io. 3129.
MATERIAL. Two specimens from the locality and horizon below.
Locatity. Sor Range section.
Horizon. Upper Palaeocene, sample no. 460-i.

DESCRIPTION. Carapace short, subquadrate in lateral view. Dorsal margin
slightly irregular due to surface ornamentation, ventral margin almost straight,
anterior broadly and evenly rounded, posterodorsal margin straight, posterior
extremity somewhat rounded, posteroventral margin curved. Greatest length lies
below mid-point, greatest height in the anterior third and greatest width behind the
middle. Anterior and posterior cardinal angles well-developed. Valves almost
equal. Evye-tubercle rounded, polished and distinct. Swbcentral-tubercle present but
not well-developed. Surface reticulate with superimposed papillae. A weak ventral
ridge at some distance from the ventral margin slopes obliquely upwards towards the

36 EARLY TERTIARY OSTRACODA

posterior end. The posterodorsal process is a short horn-like ridge slightly anterior
to the posterior cardinal angle. A marginal rim runs along anterior, venter and
posterior margins.
DIMENSIONS (mm).
L H WwW
Io. 3129 Carapace 0°59 0:39 0°34
CoMPARISON. This species has already been compared with Echinocythereis
(Echinocythereis) contexta sp. nov.

Echinocythereis (Scelidocythereis) rasilis sp. nov.
(Plate 17, figs. 5, 6, 10; Plate 18, figs. 1-3, 5, 7)

DERIVATION OF NAME. Latin vasilis, smoothed ; with reference to the carapace.

DraGnosis. Carapace subreniform. Dorsal margin arched with a slight con-
cavity behind the protruding anterior cardinal angle. Surface smooth with two
ventral ridges.

HoLotyPe. Io. 4329, a female carapace (Pl. 17, figs. 6 ; Pl. 18, figs. 2, 3).
PARATYPES. Io. 4278 + Io. 3131-2.

MATERIAL. 17 specimens from the Rakhi Nala section from three horizons
(sample nos. 3499, 3614 and 3617). 41 specimens from the Zao River section from
seven horizons (sample nos. 24145, 24147, 24148, 24150, 24152 and 24157). GSP
BM 2537-8.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Lower Chocolate Clays, sample no. 3499.

DESCRIPTION. Carapace subreniform in lateral outline, with the greatest height
at the anterior cardinal angle. Dorsal margin arched with a slight concavity behind
the anterior cardinal angle, ventral margin incurved anterior of the middle, particu-
larly in the right valve ; anterior margin broadly rounded, posterodorsal slope very
slightly concave, posterior extremity rounded, posteroventral margin curved or
straight. In dorsal view the greatest width lies almost at the middle. Anterior and
posterior marginal areas compressed. Anterior cardinal angle protruding, Right
valve over-reaches left valve along the anterior and posteroventral margins. Left
valve over-reaches right valve slightly in the regions of the anterior cardinal angle
and posterodorsal slope. Surface smooth. There are two ventral ridges, the one
nearest the ventral margin being smaller. Eye-tubercle more or less distinct and situ-
ated below the anterior cardinal angle. Anterior margin finely denticulate (20-25
small denticles), posterior with 6-8 larger denticles. Duplicature fairly wide,
0-073 mm. anteriorly. In the right valve the selvage and flange groove are well-
developed particularly in the anteroventral and ventral regions. In the left valve
the selvage is well-marked but the flange groove is somewhat less well-developed.
Radial pore canals not clearly visible but would seem to be simple, straight and
numerous. No vestibule. Hinge not determinable.

FROM WEST PAKISTAN 37

DIMENSIONS (mm).

Ib H W
Io. 4278 Carapace male 0°59 0:46 0°37
Io. 4329 Carapace female (holotype) 0:56 0°45 0°37
Io. 3131 Carapace male 0-76 0-49 0°37
Io. 3132 Carapace female 0-78 0-49 0:42

CoMPARISON. Hemucythere sahnit Tewari and Tandon (1960) appears to be a
closely related species. Specimens of this were not available for comparison but
from the description and figures given by these authors it does not seem to have the
concavity behind the anterior cardinal angle which is present in E(S.) vasilis, sp. nov.

REMARKS. The marginal denticles are not preserved in all specimens.

Echinocythereis (Scelidocythereis) sparsa sp. nov.
(Plate 18, figs. 4, 6, 8, 9)

DERIVATION OF NAME. Latin sparsus, scattered ; with reference to the papillae.

Diacnosis. A species of Scelidocythereis with subrectangular carapace, dorsal
margin slightly arched, ventral margin incurved in front of the middle. Surface
ornamented with scattered papillae and two ventral ridges. Anterior and posterior
marginal rims distinct. Left valve larger than right.

HoLotyPe. Io. 4330, a female carapace (Pl. 18, figs. 8, 9).
PARATYPE. Io. 4279.

MATERIAL. 43 specimens from the locality below from three horizons (sample
NOS. 24159, 24181, and 24183). GSP BM 2539-40.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24159.

DESCRIPTION. Sexual dimorphy moderate, the males are longer in proportion
than the females. Cavapace subrectangular in side view with greatest height at the
anterior cardinal angle. Dorsal margin slightly arched, ventral margin sinuated
anterior to the middle. Anterior margin broadly rounded, posterior somewhat
rounded. Right valve larger than left valve, which it over-reaches along the
anterior and ventral margin. Anterior cardinal angle rounded. Swubcentral-tubercle
indistinct, eye-tubercle rounded and distinct. Surface ornamentation consists of
sparsely distributed papillae. There are two ventral ridges ; the top ridge bifurcates
posteriorly but the bottom ridge is shorter. Anterior and posterior marginal
rim fairly well-developed. Anterior margin ornamented with small and numerous
denticles, posterior with 6-8 larger denticles. Duflicature moderately wide
with a prominent selvage and flange-groove in the right valve particularly in the
ventral and anteroventral regions. Radial pore canals simple, almost straight,
irregularly spaced, 25-30 anteriorly and 12-15 posteriorly. Hinge holamphidont :
right valve with anterior tooth—conical and projecting, followed by postjacent
socket, shallow posteromedian groove and posterior reniform tooth. Muscle scar

38 EARLY TERTIARY OSTRACODA

pattern consists of four adductors in a vertical superposition and two more or less
rounded frontal scars.

DIMENSIONS (mm).

L H W
Io. 4279 Carapace male 0-78 0°45 0°37
Io. 4330 Carapace female (holotype) 0-76 0:49 0°39

CoMPARISON. LEchinocythereis (Scelidocytherets) rasilia sp. nov. although smaller
than the present species may be ancestral but has a smooth rather than a papillose
surface. Moreover, it has a concavity behind the anterior cardinal angle and lacks
the distinct marginal rims.

REMARKS. Echinocytherets (Scelidocytherets) sparsa has so far only been found in
the Upper Chocolate Clays of the Zao River area.

Genus GYROCYTHERE nov.

DERIVATION OF NAME. Greek gyros, circle ; with reference to the concentric
arrangement of the ornamentation + cythere.

DiacGnosis. Reticulate Trachyleberididae with three or four longitudinal ridges,
the dorsal ridge distinct from the eye-tubercle, arcuate, sloping down towards
anterior and terminating below the eye-tubercle ; the third ridge more or less distinct
in different species.

TYPE SPECIES. Gyrocythere exaggerata sp. nov.

DESCRIPTION. Sexual dimorphism rather pronounced ; the females are shorter,
higher and wider than the males. Carapace subrectangular to subquadrate in
lateral view. Valves almost equal. Eye-tubercle and subcentral tubercle present,
more or less pronounced. Surface reticulate. Three to four longitudinal ridges
present ; the dorsal ridge commences anteriorly below the eye-tubercle and is arched
convexly upwards ; the second ridge stretches backwards from the subcentral-
tubercle and is also arched convexly upwards, its continuation in front of the sub-
central-tubercle being less pronounced ; the third ridge situated below the sub-
central-tubercle slopes obliquely upwards towards the posterior and is curved con-
vexly downward towards the anterior and the ventral ridge is confined to the
posterior two-thirds of the carapace and culminates in a slight alar expansion in the
posterior, almost obsolete or absent in some species. Normal pores simple, fairly
numerous. Radial pore canals simple, irregularly spaced, more or less straight, a few
seem to bifurcate, approximately 25 anteriorly. Inner margin and line of concresc-
ence coincide. Duplicature moderately wide. Selvage well-marked, submarginal
in left valve but at some distance from the outer margin in right valve. Ventral
and anterior flange grooves well-developed in right valve. Hinge holamphidont
with stirpate anterior tooth in right valve. Muscle scar pattern consists of four
adductor scars in an almost vertical row and a U-shaped frontal scar, which opens to
the anterodorsal angle.

CoMPARISON. This genus differs from the genus Costa in having an arcuate dorsal

FROM WEST PAKISTAN 39

ridge and in the less evident anterior marginal rim. Further, the subcentral-
tubercle in Costa lies more towards the anterior. Hermanites has only two longitu-
dinal ridges and has a concave posterodorsal slope. Gyrocythere lacks the very wide
duplicature seen in Paracytheretta.

Gyrocythere exaggerata sp. nov.
(Plate 18, figs. 10-14 ; Plate 19, Plate 20, fig. 5)

DERIVATION OF NAME. Latin exaggeratus, exaggerated ; with reference to the well-
developed longitudinal ridges.

Diacnosis. A species of the genus Gyrocythere with prominent eye-tubercle,
bilobate subcentral-tubercle and well-developed longitudinal ridges.

HototyPe. lo. 4331, a female carapace (PI. 19, figs. I-4).
PARATYPES. Io. 4280 + 3122-3128

MATERIAL. 39 specimens from the Zao River section from six horizons (sample
NOS. 24145, 24147, 24148, 24150 24152). Eight specimens from the Rakhi Nala
section from two horizons (sample nos. 3613 and 3614). GSP BM 2541-2.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24151.

DESCRIPTION. Carapace subrectangular in the male dimorph and subquadrate in
the female. Anterior margin broadly and evenly rounded, posterior narrow, almost
rounded in left valve but slightly subangular in the right valve. Dorsal and ventral
margins almost concealed in lateral outline by the dorsal and ventral ridges. Great-
est height in the ocular region, greatest length passes through the mid-point. Anterior
cardinal angle prominent with a concavity behind in lateral view. Valves almost
equal. LEvye-tubercle prominent, rounded and polished, stands out in lateral view.
Subcentral-tubercle distinct and bilobate. Surface coarsely reticulate. Reticulae
are slightly papillose in some specimens. There are four well-developed longitudinal
ridges ; the dorsal ridge begins below the eye-tubercle anteriorly and is convex up-
wards, whilst the median or second ridge runs from the subcentral-tubercle pos-
teriorly and is convex upwards, its extension anterior to the subcentral-tubercle is
less well-marked. A third ridge is intercalated between the median and the ventral
ridges. It slopes obliquely upwards towards the posterior and is convex downwards
inits anterior part. The ventral ridge is restricted to the posterior two-thirds of the
carapace. It ends in a slight alar expansion in the posterior third of the carapace.
Anterior margin denticulate, posteroventral margin with short spines present in some
specimens. Normal pore canals simple, numerous (PI. 19, figs. 6, 7). Radial pore
canals not very well-displayed due to the form of preservation, but appear to be
simple, almost straight, irregularly spaced (few seem to bifurcate), with some 25 at
the anterior margin. Line of concrescence and inner margin coincide. Duplicature
of moderate width, 0-07 mm. anteriorly. Selvage pronounced in both valves ; it is in
the outer third of the duplicature in right valve but submarginalin left valve. Right
valve with well-developed flange groove, particularly on the venter. Muscle scars are

40 EARLY TERTIARY OSTRACODA

in a vertical row of four adductors and a U-shaped frontal scar opening towards the
anterodorsal corner. Hinge holamphidont :

Element Left valve Right valve
Anterior Rounded socket, confluent Projecting stirpate
with ocular sinus, seen tooth.

in a few specimens.
Anteromedian Subconical tooth which has Deep rounded socket
straight anterior but convex opening into postero-

posterior in dorsal view. median groove.
Posteromedian Denticulate bar. Locellate groove.
Posterior Deep, slightly elongate Pessular tooth, but sub-
socket. rectangular in lateral view.
DIMENSIONS (mm).
L H W
Io. 3125 Left valve male 0-81 0-49 —
Io. 3127. Right valve male 0:83 0°49 —
Io. 4331 Carapace female (holotype) 0-78 0°49 0-46
Io. 3126 Left valve male 0-76 0-46 —
Io. 3124 Right valve female O-7I 0°44 —_-
Io. 3128 Right valve female o-71 0°44 —
Io. 4280 Left valve male 0-77 0-46 —
Io. 3120 Right valve female 0-72 0°44 —
Io. 3122 Right valve male 0-79 0-44 —

COMPARISON. This species resembles Gyrocythere perfecta sp. nov. (Pl. 22, figs. I-10)
but differs from it in being larger and having more prominent longitudinal ridges and
an eye-tubercle. Moreover, the subcentral tubercle in G. exaggerata is distinctly
bilobate.

REMARKS. The occurrence of this species ranges through 390 ft. in the Zao River
section and 15 ft. in the Rakhi Nala section. It seems likely that it will prove a
useful horizon marker.

Gyrocythere parvicarinata sp. nov.
(Plate 20, figs. 1-4, 6-8, 12)
DERIVATION OF NAME. Latin parvus, little + carinatus, ridged; with reference to
the longitudinal ridges.

Diacnosis. A strongly reticulate species of the genus Gyrocythere with three
longitudinal ridges, median ridge ill-defined, posterior subtriangular, eye-tubercle
distinct, subcentral-tubercle well-developed.

HoLotyPe. Io. 4334, a male carapace (PI. 20, figs. 1, 2, 6, 7).

PARATYPE. Io. 4281.

MATERIAL. Over 100 specimens from the Rakhi Nala section from 25 horizons

(sample nos. 3153, 3168 to 3172, 3179, 3180, 3185, 3192, 3193, 3199, 3200, 3401 to
3405, 3407, 3409, 3410, 3415 and 3417). GSP BM 2543-4.

FROM WEST PAKISTAN 41

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Green and Nodular Shales, sample no. 3407.

DESCRIPTION. Sexual dimorphism distinct ; the males are more elongate than the
females. Cavapace subrectangular in side view. Anterior margin broadly rounded,
posterior subtriangular. Dorsal margin straight but appears slightly convex in
lateral view due to the over-reaching of the dorsal ridge, ventral margin slightly
concave in front of the middle. Anterior cardinal angle distinct with a concavity
behind in lateral view. Left valve slightly over-reaches right valve at anterior
cardinal angle and posterodorsal slope. In dorsal view the greatest width lies in the
anterior two-fifths. Swubcentral-tubercle well developed. Evye-tubercle distinct. Surface
strongly reticulate with three longitudinal ridges ; the dorsal ridge is curved con-
vexly upwards ; the median ridge is more or less ill-defined in many specimens ; the
third or ventral ridge is curved convexly downward anteriorly. Anterior and
posterior marginal rims present but not high. Both anterior and posterior margins
are denticulate. Dwuplicature of medium width. Selvage distinct and at some
distance from the outer margin in right valve. Anterior and ventral flange grooves
well-developed in right valve. Radial pore canals not clearly seen due to mineraliz-
ation. Hinge as for the genus.

DIMENSIONS (mm).

L H W
Io. 4334 Carapace male (holotype) 0:68 0°37 0°34
Io. 4281 Carapace female 0:67 0°42 0°37

COMPARISON. This species is smaller than Gyrocythere grandilaevis sp. nov.
Although the longitudinal ridges are no better developed than G. grandilaevis the
eye-tubercle and subcentral-tubercle are more prominent, the reticulation is deeper
and wider in proportion.

Gyrocythere grandilaevis sp. nov.
(Plate 20, figs. 9-11, 13 ; Plate 21, figs. 1-4)
DERIVATION OF NAME. Latin grandis, large + laevis, smooth ; with reference to
the carapace.

Diacnosis. A species of Gyrocythere with large, reticulate, smooth carapace.
Three longitudinal ridges, including median ridge which is not well-developed.
Anterior and posterior marginal rims distinct.

HOLotyPe. Io. 4332, a female carapace (PI. 20, figs. 11, 13 ; Pl. 21, figs. 3, 4).

PARATYPE. lo. 4282.

MATERIAL. 16 specimens from the locality below from four horizons (sample

nos. 3463 to 3466). Two specimens from the Shpalai Khwara section from one
horizon (sample no. 24692). GSP BM 2545-6.

TYPE LOCALITY. Rakhi Nala section.
TyprE Horizon. Shales with Alabaster, sample no. 3463.

42 EARLY TERTIARY OSTRACODA

DESCRIPTION. Sexual dimorphism moderate ; the males are proportionally longer
than the females. Carapace subrectangular in lateral outline. Anterior margin
broadly rounded, posterior margin almost rounded in left valve but with a slight
concavity in the posterodorsal slope of the right valve. Dorsal margin almost
straight but appears slightly convex in lateral view due to the dorsal ridge, which
slightly over-reaches it ; ventral margin slightly concave in front of the middle.
Valves almost equal. Swbcentral-tubercle and eye-tubercle present but not pro-
nounced. Surface reticulate ; the reticulae concentrically arranged around the
subcentral-tubercle. There are three longitudinal ridges, a dorsal ridge curved
convexly upward, a median or second less well-developed ridge and a ventral or
third ridge which runs obliquely from above the anteroventral corner towards the
posterior and is curved convexly downward in its anterior portion. Anterior and
posterior marginal rims distinct. Anterior and posterior margins denticulate.
Internal features not seen.

DIMENSIONS (mm).

L H W
Io. 4332 Carapace male (holotype) 0°85 0-46 0°44
Io. 4282 Carapace female 0°83 0:46 0:46

COMPARISON. This species resembles Gyrocythere parvicarinata sp. nov. but differs
from it in being larger. Moreover, the posterior in G. parvicarinata is subacuminate.
G. grandilaevis is perhaps ancestral to Gyrocythere perfecta sp. nov. which is smaller
and has stronger ornamentation.

Gyrocythere mitigata sp. nov.
(Plate 21, figs. 5-11)

DERIVATION OF NAME. Latin mitigatus, mellowed ; with reference to the orna-
mentation, less emphatic than in the typical species of G. exaggerata.

Diacnosis. A large, strongly reticulate species of the genus Gyrocythere with
three longitudinal ridges, the median ridge almost ill-defined.

HoLotyPe. Io. 4333, a male carapace (P. 21, figs. 5-8).

PARATYPES. Io. 4283 + Io. 3119.

MATERIAL. 9g specimens from the locality below from two horizons (sample nos.
24131 and 24132). lo. 4283. GSP BM 2547-8.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Lower Chocolate Clays, sample no. 24131.

DESCRIPTION. Sexually dimorphic ; the males are longer than the females.
Carapace subrectangular in lateral view. Anterior margin broadly and evenly
rounded, dorsal margin almost straight in reality but appears slightly convex in
lateral view because of the over-reaching of the dorsal ridge ; posterodorsal slope
very slightly concave particularly in the right valve, posterior extremity rounded,
posteroventral margin curved, ventral margin almost straight. Greatest height in
the anterior quarter, greatest length almost in the middle. Anterior and posterior

FROM WEST PAKISTAN 43

cardinal angles well-marked, particularly in left valve. Left valve slightly over-
reaches the right valve in the anterodorsal corner and at the posterodorsal slope.
Greatest width in dorsal or ventral view lies in the posterior third. Swubcentral-
tubercle prominent, eye-tubercle rounded and distinct. Surface strongly reticulate
with three longitudinal ridges : the dorsal ridge is curved convexly upwards in the
middle and starts anteriorly below the eye-tubercle ; the median ridge is less well-
developed, almost ill-defined in most specimens ; it runs posteriorly from the sub-
central-tubercle and is curved convexly upwards. The ventral ridge commences
anteriorly above the anteroventral corner and runs obliquely upwards towards the
posterior and culminates in the posterior third. Marginal rim is distinct at the
anterior and posterior but less distinct along the venter. Anterior margin denti-
culate, posterior extremity and posteroventral margin ornamented with about six
short spines or papillae. Duplicature of moderate width. Selvage well-developed,
subperipheral in the left valve but situated at some distance from the outer margin
in the right valve. The right valve has a fairly deep flange groove along the venter
and around the anterior margin. Hinge as for the genus.
DIMENSIONS (mm).

L H W
Io. 4333 Carapace male (holotype) 0-88 0°49 0°49
Io. 4283 Carapace female 0°83 0°49 —
To. 3119 Left valve female 0-80 0°47 —

COMPARISON. Gyrocythere grandilaevis sp. nov. is somewhat similar to the present
species and might even be ancestral although G. grandilaevis is smaller and has a less
well-developed subcentral-tubercle. The dorsal and ventral ridges in G. grandilaevis
are also less well-marked. G. mitigata differs from Gyrocythere exaggerata sp. nov. in
being larger, having a different lateral outline and less emphatic ornamentation.
Further, G. mitigata has three, rather than four longitudinal ridges and lacks a
bilobate subcentral-tubercle.

REMARKS. G. mitigata has so far only been found in the Zao River section, where
it occurs at two horizons.

Gyrocythere perfecta sp. nov.
(Plate 22, figs. 1-10)

DERIVATION OF NAME. Latin perfectus, perfect ; with reference to the beauty of
the material.

Diacnosis. Gyrocythere with strongly reticulate, concentrically arranged orna-
mentation. Eye-tubercle, subcentral-tubercle and longitudinal ridges distinct.

HoLotyPe. Io. 4335, a female carapace (Pl 22, figs. 3, 4, 7, 8).
PARATYPES. Io. 4284 + Io. 3120-1.

MATERIAL. 20 specimens from the locality below from two horizons (sample
nos. 3498 and 3499). GSP BM 2548-50.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Lower Chocolate Clays, sample no. 3499.

44 EARLY TERTIARY OSTRACODA

DESCRIPTION. Carapace subrectangular in lateral view, arrow-shaped in ventral
view. Sexual dimorphism rather pronounced ; the females are higher and wider
than the males. Anterior margin broadly rounded, posterior slightly subangular,
particularly in the right valve. Dorsal margin straight but appears to be convex
due to the over-reaching of the dorsal ridge ; ventral margin slightly concave
anterior to the middle. Anterior cardinal angle distinct and rounded. Valves
almost equal. Fye-tubercle distinct but not high. Swubcentral-tubercle distinct,
slightly lobate. Surface strongly and deeply reticulate, the reticulation being
concentric around the subcentral-tubercle. Four longitudinal ridges occur : the
dorsal ridge is convex upwards ; it commences below the eye-tubercle and culminates
in the posterodorsal region ; the median or second ridge is also convex upwards ; it
runs from the subcentral-tubercle towards the posterior but its continuation anterior
to the subcentral-tubercle is not distinct ; the third ridge is convex downwards in its
anterior half and slopes obliquely upwards towards the posterior ; the ventral ridge
is better seen in ventral view ; it is confined in the posterior three-quarters. Anterior
margin denticulate. There isa short posteroventral spine present in most specimens.
Radial pore canals simple, more or less straight, irregularly spaced, few seem to
bifurcate, about 25-28 in the anterior. Inner margin and line of concrescence
coincide throughout. Dwuplicature moderately wide with a distinct selvage. In the
right valve the selvage lies at some distance from the outer margin and the anterior
and ventral flange grooves are distinct. Adductor scars in a vertical column of four
and with a U-shaped frontal scar. Hinge holamphidont.

DIMENSIONS (mm).

L H W
Io. 4281 Carapace male 0-71 0°39 0:38
Io. 4335 Carapace female (holotype) 0-71 0-42 0°42
Io. 3121 Right valve male 0-76 0°42 0°42
Io. 3121 Right valve female 0-76 0-42 —
Io. 3120 Right valve female 0-76 0°44 a

CoMPARISON. The present species is similar in all characters to Gyrocythere
exaggerata sp. nov. but is smaller and with a less well-marked ornamentation. On
the other hand, the ornamentation is stronger than in Gyrocythere grandilaevis sp. nov.
In both the morphological development and stratigraphical position, G. perfecta falls
between G. grandilaevis and G. exaggerata.

REMARKS. G. ferfecta has so far only been found in the type locality.

Genus HERMANITES Puri 1955
TYPE SPECIES. Hermania reticulata Puri 1954.
Hermanites cracens sp. nov.
(Plate 22, figs. 11 ; Plate 23, figs. 1-3)

DERIVATION OF NAME. Latin cracens, graceful ; with reference to the pleasing
curve of the ridges.

FROM WEST PAKISTAN 45

Diacnosis. A large Hermanites with well-developed slightly curved dorsal and
ventral ridges. Subcentral-tubercle prominent with three small curved longitudinal
ridges behind.

Ho.otyPe. lo. 4336, a carapace.

MATERIAL. Only one specimen from the locality and horizon below.
TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Gorge Beds, sample no. 3I1I1T.

DESCRIPTION. Carapace large, massive, subrectangular in lateral view. Greatest
length through the mid-point, greatest height through the anterior cardinal angle and
greatest width in the posterior third. Dorsal and ventral margins almost straight,
anterior broadly and evenly rounded, posterodorsal margin very slightly concave,
posteroventral margin and posterior extremity rounded. Anterior cardinal angle
well-developed particularly in the left valve. Left valve slightly over-reaches the
right at the anterior cardinal angle and posterodorsal slope. Swbcentral tubercle
prominent. Evye-tubercle rounded and distinct and with a rounded groove in front,
particularly in the left valve. Marginal rim high in the anterior but somewhat less
high in the posterior and venter. Surface coarsely reticulate with well-marked,
dorsal and ventral ridges ; the dorsal ridge commences above the subcentral-tubercle
and is slightly curved convexly upwards ; the ventral ridge slopes obliquely upward
towards posterior and then curves sharply round towards posteroventral margin
in posterior quarter. There are three short curved longitudinal ridges behind the
subcentral-tubercle, the bottom one being the shortest. Anterior and posterior
margins spinose—anterior with numerous small spines but posterior with few
larger spines.

DIMENSIONS (mm).
iv H WwW

Io. 4336 Carapace 0:95 0°51 0°51

CoMPARISON. Hermanites palmatus sp. nov. is much smaller, has the dorsal and
ventral ridges joined posteriorly by a transverse ridge and the ridges in front of the
subcentral-tubercle have a palmate appearance.

REMARKS. So far only one specimen of this species has been recovered from the
Gorge Beds of the Rakhi Nala section, where it occurs in association with Alocopocy-
there rupina sp. nov. and Buntonia devexa sp. nov.

Hermanites scopus sp. nov.
(Plate 23, figs. 4-10)
DERIVATION OF NAME. Latin scopus, target ; in allusion to the fancied resem-
blance of the ornamentation to a bull’s-eye.

Diacnosis. A species of the genus Hermanites in which ventral ridge curves down-
ward in the middle and is joined by a short vertical ridge at its posterior end ; surface

46 EARLY TERTIARY OSTRACODA

coarsely reticulate with prominent subcentral-tubercle from which a ridge runs
towards anterior margin.

Ho.otyPe. Io. 4338, a male carapace (Pl. 23, figs. 4-7).
PARATYPE. Io. 4285.

MATERIAL. 13 specimens from the Rakhi Nala section from six horizons (sample
Nos. 3499, 3610, 3613 to 3615 and 3618). Two specimens (including holotype) from
the Zao River section from two horizons (sample nos. 24148 and 24150). GSP BM
2552-3.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 24148.

DESCRIPTION. Sexual dimorphy observed ; the males are longer in proportion
than the females. Carapace thick-shelled, sub-rectangular in lateral view. Anterior
margin broadly and evenly rounded, posterior slightly subtriangular. Dorsal and
ventral margins almost straight. Valves more or less equal. Anterior cardinal
angle well-developed. In dorsal view, greatest width lies in the posterior third.
Evye-tubercle rounded and prominent with rounded deep groove in front (better seen
in dorsal view). Swubcentral-tubercle prominent with a ridge running towards the
anterior margin. Surface strongly reticulate (occasionally with superimposed
rounded spines particularly at the junction where two reticulae meet). Dorsal and
ventral ridges well-developed : the dorsal ridge is almost straight while the ventral
ridge is curved convexly downward in the middle culminating in a short vertical
ridge in the posterior third. Anterior and posterior marginal rims elevated, ventral
marginal rim somewhat less elevated. Anterior and posterior margins ornamented
with double row of spines (not preserved in all specimens)—one row of spines lies on
the anterior and posterior marginal rims and the second row below these rims.
Duplicature of moderate width with subperipheral selvage. Radial pore canals and
muscle scars not known. Hinge holamphidont : left valve hinge consists of two
terminal sockets, median element subdivided into anteromedian subconical pro-
jecting tooth and denticulate bar ; right valve hinge (seen only in a broken valve)
consists of a stirpate, projecting anterior tooth, deep anteromedian socket, locellate
posteromedian groove and posterior tooth (broken).

DIMENSIONS (mm).

L H WwW
Io. 4338 Carapace male (holotype) 0-78 0°44 0°44
Io. 4285 Carapace female 0-81 0:46 0-49

COMPARISON. This species shows some resemblance to Hermanites cracens sp. nov.
but is smaller, has a more curved ventral ridge culminating in a short vertical ridge
in the posterior third and lacks the three small, curved longitudinal ridges behind the
subcentral-tubercle.

REMARKS. Hermanites scopus rarely occurs in the Lower and Upper Chocolate
Clays of the Rakhi Nala and Zao River sections and can easily be recognized by its
characteristic ventral ridge.

EEE eee

FROM WEST PAKISTAN 47

Hermanites palmatus sp. nov.
(Plate 24, figs. I-9, II, 12)

DERIVATION OF NAME. Latin palmatus, palmate ; with reference to the palmate
appearance of the ridges in front of the subcentral-tubercle.

Diacnosis. Hermanites in which dorsal and ventral ridges are alate and joined
posteriorly by a transverse ridge which is slightly concave towards posterior. Sub-
central-tubercle prominent with palmate appearance of ridges in front.

Hototyre. lo. 4337, a female left valve (Pl. 24, figs. 6, 8, 9, I1).

PARATYPES. Io. 4286 + Io. 3117-8.

MATERIAL. 9g specimens from the Rakhi Nala section from five horizons (sample
nos. 3613 to 3615, 3617 and 3618). 17 specimens from the Zao River section from
four horizons (sample nos. 24131, 24150, 24152 and 24156). GSP BM 2554-5.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 24152.

DESCRIPTION. Sexual dimorphism apparent ; the females being shorter than the
males. Cavapace subrectangular in lateral outline with greatest height in the region
of the anterior cardinal angle. Dorsal and ventral margins almost straight, anterior
margin broadly rounded, posterodorsal corner very slightly concave, posterior
extremity and posteroventral margin rounded. Valves almost equal. Swubcentral-
tubercle prominent. Evye-tubercle rounded and distinct. Surface recticulate with
alate dorsal and ventral ridges which are joined posteriorly by a transverse ridge
which is slightly concave towards the posterior. Anterior and posterior marginal
platforms compressed. Anterior and posterior marginal rims distinct. Anterior
margin ornamented with 20-25 small spines, posterior with 6-8 larger spines.
Duplicature of moderate width, 0-050 mm. anteriorly. Both valves have a distinct
seluage. Hinge holamphidont : left valve hinge with fairly deep anterior socket,
anteromedian tooth rounded, subconical (slightly projecting towards the anterior in
dorsal view), posteromedian ridge denticulate and deep, slightly elongate posterior
socket ; right valve hinge with anterior subconical, projecting tooth followed by deep
postjacent socket, posteromedian shallow groove and an almost rounded, posterior
tooth.

DIMENSIONS (mm).

L H WwW
Io. 4286 Carapace male 0:69 0°37 0°37
Io. 3117 Carapace female 0:63 0:37 0:37
Io. 4337. Left valve female (holotype) 0-73 0-40 —
Io. 3118 Left valve female 0-73 0°44 _-

CoMPARISON. Hermanites scopus sp. nov. is larger, has a curved ventral ridge and
subtriangular posterior margin. Hermanites indicus Tewari and Tandon (1960) has
a subtriangular posterior end and lacks the transverse ridge which posteriorly joins
the dorsal and ventral ridges. The Miocene species Hermanites purist Tewari and
Tandon (1960) has somewhat similar lateral outline, but lacks both the transverse

48 EARLY TERTIARY OSTRACODA

ridge which posteriorly joins the dorsal and ventral ridges and the palmate appear-
ance of ridges in front of the subcentral-tubercle. Moreover, the greatest width in
Hermanites palmatus is in the posterior third, but in Hermanites puri it is a little to
the anterior of the middle (see Tewari and Tandon, p. 158).

REMARKS. The true relationship of the present species with Hermanites indicus
and Hermanites purti cannot be determined until topotype material from Kutch is
available for comparison.

Genus OCCULTOCYTHEREIS Howe 1951

TYPE SPECIES. Occultocythereis delumbata Howe 1951.

Occultocythereis interrupta sp. nov.
(Plate 24, figs. 10, 13-18)

DERIVATION OF NAME. Latin interruptus, broken apart ; with reference to the
break in the dorsal ridge.

Diacnosis. A small Occultocythereis with well-marked subangular postero-
dorsal process which is confluent with four ridges including an oblique ridge running
towards posteroventral region and then extending towards anteroventral corner as
an oblique ventral ridge.

Ho.otyPe. Io 4339, a female carapace (Pl. 24, figs. 14, 17, 18).

PARATYPE. Io. 4287.

MATERIAL. 28 specimens from the locality below from three horizons (sample
nos. 460-f, 460-i and 460-j). GSP BM 2556-7.

TYPE LOCALITY. Sor Range section.

TYPE HORIZON. Upper Palaeocene, sample no. 460-1.

DESCRIPTION. Sexual dimorphism marked ; the females are shorter and wider
than the males. Cavapace subrectangular in side view, tapering towards the
posterior. Dorsal and ventral margins almost straight ; the dorsal margin appears
slightly irregular in lateral outline because of the over-reaching of the dorsal ridge ;
anterior margin broadly and obliquely rounded, posterior narrow, slightly concave
in posterodorsal slope but more or less rounded in posteroventral margin. Greatest
length lies through the mid-point and greatest height at the anterior cardinal angle.
Anterior and posterior cardinal angles well-marked particularly in the right valve.
Valves almost equal. Eye-tubercle distinct, situated on the anterior marginal rim.
Shell surface undulating with compressed anterior and posterior platforms. A well-
marked, subangular posterodorsal process is confluent with four ridges : (1) a short
ridge extends vertically towards the posterodorsal corner ; (2) a dorsal ridge runs
towards the anterior, culminating in the anterior third of the dorsal margin—it is
slightly convex upwards ; (3) a short ridge extends vertically below, terminating
before reaching the mid-line, whilst the fourth ridge runs obliquely towards the pos-
teroventral region where it is joined by an oblique ventral ridge running towards

FROM WEST PAKISTAN 49

the anteroventral corner. Anteromedian swelling well-developed. A marginal rim
extends around the anterior, ventral and posterior margins, elevated on the anterior
margin but less elevated around the venter and posterior. Small ridges run be-
tween the anterior marginal rim and the anterior margin (these are better seen in
ventral view). Four to five short spines ornament the posteroventral margin.
Internal details not known.

DIMENSIONS (mm.)

L H W
Io. 4287 Carapace male 0°39 0-18 O-II
Io. 4339 Carapace female (holotype) 0:38 0-20 0-13

CoMPARISON. Occultocythereis indistincta sp. nov. is much larger, has a continuous
rather than a broken dorsal ridge and lacks the oblique posterior ridge which joins
the posterodorsal process and the ventral ridge.

REMARKS. O. interrupta is so far only known from the Upper Palaeocene of the
Sor Range section.

Occultocythereis Sp.A
(Plate 25, figs. 1, 2, 5, 12)

FIGURED SPECIMEN. Io. 3136.

MATERIAL. Only one specimen from the locality and horizon below.

Locatity. Rakhi Nala section.

Horizon. Lower Rakhi Gaj Shales, sample no. 3672.

DESCRIPTION. Carapace subrectangular with ventral inflation. Dorsal and
ventral margins almost straight, tapering towards the posterior ; anterior margin
broadly and obliquely rounded, posterior narrowly rounded with a slight concavity
in the posterodorsal corner. Greatest length runs through the middle, greatest
height in the anterior two-fifths and greatest width in the posterior third. Anterior
and posterior cardinal angles rounded. Valves almost equal. FE ye-tubercle distinct
but low. Swbcentral-tubercle weak. Surface reticulate. Posterodorsal process
consists of a more or less rounded tubercle which extends anteriorly in a weak dorsal
ridge. Anterior marginal rim well-marked, ventral and posterior marginal rims less
elevated. Posterior ornamented with 4-5 short spines.

DIMENSIONS (mm).

L FA Ww
To. 3136 Carapace 0°42 0:22 Or

REMARKS. This species differs from other known species of the genus Occulto-

cytherets in its obliquely rounded anterior margin.

Occultocythereis spilota sp. nov.
(Plate 25, figs. 3, 4, 6-11)

DERIVATION OF NAME. Greek sfilotos, spotted ; with reference to the largish
puncta.

D

50 EARLY TERTIARY OSTRACODA

DiaGnosis. A species of Occultocythereis in which surface is ornamented with
largish puncta. Posteroventral margin rounded, posteroventral process a short
slightly oblique ridge well-developed in female, ill-defined in male, anteroventral
swelling small.

HoLotyPe. Io. 4342, a female carapace (Pl. 25, figs. 6, 7, 10, 1).
PARATYPE. Io. 4288.

MATERIAL. Four specimens from the locality below from three horizons (sample
nos. 3173, 3174 and 3177). GSP BM 2558-9.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Green and Nodular Shales, sample no. 3177.

DESCRIPTION. Dimorphic ; the females are higher and wider than the males.
Carapace subrectangular in lateral view with greatest length in the middle and
greatest height in the anterior third. Anterior margin broadly rounded, postero-
dorsal slope very slightly concave, posteroventral margin rounded, dorsal and ventral
margins almost straight, very slightly converging towards the posterior. Anterior
cardinal angle rounded, posterior cardinal angle distinct about 110°. Valves almost
equal. In dorsal view the greatest width lies in the posterior third. Eye-tubercle
more or less distinct. Swbcentral-tubercle present but not pronounced with a small
swelling below and slightly anterior toit. Surface ornamented by large puncta. The
posterodorsal process is a short projecting ridge extending vertically towards the
mid-line but not reaching it. It extends anteriorly in a short dorsal ridge termina-
ting in the anterior third of the dorsal margin. The posteroventral process is a short
slightly oblique ridge lying in the posteroventral swelling. In the male dimorph
the posteroventral swelling is less well-developed and the posteroventral ridge ill-
defined. Anterior marginal rim prominent, ventral and posterior marginal rims less
prominent. Anterior marginal area ornamented by numerous, short ridges lying
between the rim and the margin. Posteroventral margin decorated by four to five
short spines.

DIMENSIONS (mm).

L H W
Io. 4288 Carapace male 0:38 0-21 O13
Io. 4342 Carapace female (holotype) 0-38 0-22 0-16

CoMPARISON. Occultocythereis peristicta sp. nov. Morphotype C is larger, has a
vertical rather than an oblique posteroventral ridge joining the second posterior
tubercle and the ventral ridge. Further, it lacks the dorsal ridge and the short
ridges between the anterior rim and the anterior margin.

Occultocythereis peristicta sp. nov.
(Plate 25, figs. 13-17 ; Plate 26 ; Plate 27, figs. 1-2)

DERIVATION OF NAME. Greek pferistiktos, punctate or dappled.
Dracnosis, A punctate group of morphotypes of the genus Occultocythereis with-

FROM WEST PAKISTAN 51

out a dorsal ridge, ventral ridge well-marked, anteromedian swelling distinct,
posterodorsal tubercle present.

Ho.otyPe. Io. 4341, a female carapace (PI. 25, figs. 15) ; (Pl. 26, figs. 2, 3).
PaRATYPES. Io. 4289-93 + Io. 3137-40.

MATERIAL. Approximately 800 specimens from the Rakhi Nala section from 42
horizons (sample nos, 3160, 3163, 3167, 3170, 3171, 3173, 3174, 3177, 3179, 3180,

3186-3194, 3197 to 3200, 3401 to 3405, 3407, 3409, 3410, 3415, 3418 to 3423, 3428,
3432, 3434 and 3435). GSP BM 2560-65.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Upper Rakhi Gaj Shales, sample no. 3167.

DESCRIPTION. Sexual dimorphism rather pronounced, the males are longer in
proportion than the females. Cavapace subrectangular or wedge-shaped in lateral
outline. Dorsal and ventral margins almost straight, tapering towards the posterior,
anterior margin broadly and evenly rounded, posterior narrow, slightly subangular
in the middle or almost rounded. Greatest length passes through the middle,
greatest height lies at the anterior cardinal angle which is fairly well-developed. Left
valve slightly over-reaches the right valve at the anterior cardinal angle and in the
region of the posterodorsal slope. LEvye-tubercle rounded and distinct, lying on the
anterior marginal rim. Subcentral or anteromedian swelling distinct, either elongate
or almost rounded (elongate in most morphotypes). Thereisnodorsalridge. Ventral
ridge fairly well-developed, short in most morphotypes, either almost straight or runs
slightly obliquely towards the posterior end where it may be connected to the second
posterior tubercle or to the posterodorsal tubercle by means of a short vertical ridge.
A dorsal ridge runs between the second posterior tubercle and the anteromedian
swelling in some morphotypes. Anterior marginal rim high, ventral marginal rim
less distinct, posterior marginal rim distinct. Short spines decorate the anterior and
posterior margins.

Trends of variants :—

I. Become sparsely punctate.

2. Gain second posterior tubercle, which may join either the posterodorsal

tubercle or the ventral ridge.

3. Become wedge-shaped.

REMARKS. O. feristicta commonly occurs in the Upper Rakhi Gaj Shales, Green
and Nodular Shales and Rubbly Limestones of the Rakhi Nala section.
This species may be divided into the following morphotypes :

MORPHOTYPE A
(Pl. 25, figs. 13-17; Pl. 25, figs. 1-3).

This has a well-delimited vertical posterodorsal tubercle. There is no second
posterior tubercle. The anteromedian swelling is less well-developed. The ventral

52 EARLY TERTIARY OSTRACODA

ridge is almost straight, short, confined in the mid-ventral region. The surface is
densely punctate.

DIMENSIONS (mm).

L H WwW
Io. 4292 Carapace male 0°43 0:22 O-II
Io. 4341 Carapace female (holotype) 0°42 0:23 0-16

MORPHOTYPE B
(Pl. 26, figs. 4-9)
This is close to Morphotype A but has a more sparsely punctate surface and a

second posterior tubercle. In addition, the present morphotype has a well-developed,
somewhat elongate anteromedian swelling.

DIMENSIONS (mm).

L H WwW
Io. 4293 Carapace male 0°44 0:22 O15
Io. 4291 Carapace female 0-40 0-22 0-16

MORPHOTYPE C
(Pl. 26, figs. 10-15)
This has a higher carapace than the other morphotypes. The ventral ridge runs
slightly obliquely towards the posterior. It is joined posteriorly to the second

posterior tubercle by a short vertical ridge. The anteromedian swelling and the
second posterior tubercle form a broken diagonal ridge.

DIMENSIONS (mm).

L H W
Io. 4289 Carapace male 0°45 0:26 0-17
Io. 4290 Carapace female 0°44 0°25 0-18

MoRPHOTYPE D
(Pl. 27, figs. 1-6)

The carapace is much more elongate than Morphotype A, B,C and E. The dorsal
and ventral margins taper very slightly towards the posterior. An oblique posterior
ridge joins the posterodorsal tubercle, the second posterior tubercle and the posterior
end of the ventral ridge. The ventral ridge commences above the anteroventral
corner, slopes obliquely upwards towards the posterior and meets the oblique
posterior ridge in the posterior quarter. A diagonal ridge which may or may not be
continuous passes through the second posterior tubercle and the anteromedian
swelling.

DIMENSIONS (mm).

L H WwW
Io. 3146 Carapace male 0°51 0:24 0-16
Io. 3139 Carapace female 0-51 0-24 0-17

FROM WEST PAKISTAN 53

MorRPHOTYPE E
(Pl. 27, figs. 7-12)
This is very similar to Morphotype B, but has a wedge-shaped carapace.

DIMENSIONS (mm).

L H WwW
Io. 3137. Carapace male 0°39 0°20 0-13
Io. 3138 Carapace female 0°37 0:20 0-13

Occultocythereis indistincta sp. nov.

(Plate 27, figs. 13-15 ; Plate 28, figs. 1-4)

DERIVATION OF NAME. Latin indistinctus, dim or obscure ; named from the
absence of well-marked diagnostic characters.

Diacnosis. A species of the genus Occultocythereis with a well-developed dorsal
ridge ending posteriorly in a large subangular posterodorsal process, ventral ridge
oblique running from anteroventral corner towards posterior, surface ornamentation
consists of indistinct puncta.

HototyPe. Io. 4340, a female carapace (PI. 27, figs. 15 ; Pl. 28, figs. 1, 3, 4).
PARATYPES. Io. 4294 + Io. 3135.

MATERIAL. 44 specimens from the locality below from seven horizons (sample
nos. 3499, 3614, 3615, 3621, 3625, 3648 and 3649). GSP BM 2566-7.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Lower Chocolate Clays, sample no. 3499.

DESCRIPTION. Sexual dimorphism apparent ; the males are more elongate, less
high and less wide than the females. Cavapace subrectangular, slightly tapering
towards the posterior. Dorsal margin straight but irregular in side view due to the
over-reaching of the dorsal ridge, ventral margin almost straight, anterior margin
broadly and evenly rounded, posterior narrow, subangular in the middle with
slightly concave posterodorsal slope. Greatest length lies in the middle, greatest
height in the anterior third. Anterior cardinal angle well-developed particularly in
left valve, posterior cardinal angle rounded in the right valve but pointed in left
valve. Left valve over-reaches right valve in the region of anterior cardinal angle
and posterodorsal slope. LEvye-tubercle distinct. Surface ornamented with small,
indistinct puncta. Anteromedian swelling (which perhaps represents a subcentral-
tubercle) distinct. A well-marked dorsal ridge commences behind the eye-tubercle
and is slightly convex upward in the middle terminating in a large subangular
posterodorsal process. In most specimens it is ornamented with three small tubercles
which lie at some distance from one another. Ventral ridge runs obliquely from the
anteroventral corner towards the posterior, culminating in the posterior third. (In
some specimens it is not well-developed). A small tubercle is present in the postero-

54 EARLY TERTIARY OSTRACODA

median part of the carapace (halfway between the dorsal and ventral ridges) in a few
specimens. A high marginal rim runs round the anterior extending along the venter
and posterior as a less high rim. Anterior and posterior margins ornamented with
short spines. Duplicature wide in anterior and posteroventral regions. Selvage
distinct and lies at some distance from the outer margin. Radial pore-canals and
muscle scars not determinable. Hinge as for the genus.

COMPARISON. Occultocythereis mutabilis abducta Triebel (1961) is very similar, but is
larger, has a different posterior end, less well-developed subcentral-swelling and
posterodorsal tubercle. Occultocythereis mutabilis mutabilis Tiebel (1961) has a
vertical posteroventral ridge in the right valve of the male and in both valves of the
females.

DIMENSIONS (mm).

L H W
Io. 4294 Right valve male 0°47 0°24 —
To. 3135 Carapace male 0°43 0:22 0-13
Io. 4340 Carapace female (holotype) 0°43 0-24 0-16

REMARKS. O. indistincta has so far been found in the Lower and Upper Chocolate
Clays of the Rakhi Nala section.

Genus PATAGONACYTHERE Hartmann 1962

TYPE SPECIES. Patagonacythere tricostata Hartmann 1962.

REMARKS. The species here assigned with query to Patagonacythere differs in
details of muscle scar pattern both from the type species described by Hartmann and
from the two species described by Benson (1964) from the Antarctic. In common
with the described species it shows the three longitudinal ridges in which a characteris-
tic posterodorsal loop joins the upper two.

Patagonacythere ? nidulus sp. nov.
(Plate 28, figs. 5-12 ; Plate 29, figs. 1-4)
DERIVATION OF NAME. Latin nidulus, small bird’s nest ; with reference to the
reticulate complex of the subcentral node.

DiacGnosis. Carapace highly reticulate in which ventral ridge culminates ab-
ruptly in posterior third, subcentral-tubercle prominent.

HoLotyPe. Io. 4349, a female carapace (PI. 28, figs. gQ—12).

PARATYPES. lo. 4295 + Io. 3096-8.

MATERIAL. Over 400 specimens from the Zao River section from 16 horizons
(sample nos. 24155 to 24157, 24159, 24166, 24170, 24173, 24175 to 24178, 24180,
24183, 24185, 24187 and 24193). Approximately 60 specimens from the Rakhi
Nala section from 13 horizons (sample nos. 3624, 3625, 3628, 3631, 3634, 3640 to 3642,
3645, 3646, 3649, 3658 and 3662). GSP BM 2568-9.

FROM WEST PAKISTAN 55

TYPE Locality. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24173.

DESCRIPTION. Carapace subrectangular in side view with dorsal and ventral
margins almost straight, subparallel ; anterior broadly rounded, posterodorsal margin
very slightly concave (particularly in the right valve), posterior extremity and
posteroventral margin rounded. Anterior cardinal angle well-developed in both
valves ; posterior cardinal angle in the left valve obtuse and well-marked but in the
right valve it is not very well developed. Left valve over-reaches the right valve
slightly at the anterior cardinal angle and at the posterodorsal slope, otherwise
the two valves are equal in size. In the dorsal view the greatest width passes
either through the subcentral-tubercle or through the posterior third with a
sulcus in between. Evye-tubercle rounded, polished and distinct ; swbcentral-tubercle
prominent, composed of reticulate complex. Shell surface strongly reticulate with
three longitudinal ridges : the ventral ridge slopes obliquely upward towards the
posterior, ending abruptly just before reaching the compressed posterior platform;
the median ridge springs from the subcentral node, stretching backward to join the
ventral ridge and forms a posterodorsal loop. (In some specimens at certain
horizons longitudinal ridges are not well-developed.) A marginal rim runs round the
anterior, ventral and posterior margins—it is upraised around the anterior margin.
Anterior margin ornamented with approximately 20 small spines, posterior margin
with 6-8 spines. Normal pore canals fairly numerous (these become exaggerated in
specimens cleaned in ultrasonic vibrator). Radial pore canals simple, nearly
straight, irregularly spaced ; few cross one another. There are approximately 40
anterior radial pore canals and about 20 posterior canals. Inner margin and line of
concrescence coincide. Dwuplicature of moderate width (0-073 mm. at anterior,
0:06 mm. at posterior extremity). Selvage prominent, submarginal in left valve,
at some distance from the margin in the right valve. Flange groove well-developed
(particularly in the right valve) on the venter and around the anterior margin.
Adductor muscle scars are in a vertical column of four elongate scars, the second from
the top being longest and the bottom one the shortest. There are two frontal scars,
the top one is smaller and almost rounded, but the bottom one is ovate. Hinge
holamphidont :

Element Left valve Right valve

Anterior Deep socket bounded on Strongly projecting
venter, confluent with slightly stirpate tooth
ocular sinus. with a concavity on

anterior in dorsal view.
Ocular sinus opens below
and slightly anterior to it.

Anteromedian Conical tooth with Deep rounded socket.
straight anterior and
convex posterior in
dorsal outline.

56 EARLY TERTIARY OSTRACODA

Element Left valve Right valve

Posteromedian Denticulate bar. Locellate groove.

Posterior Deep socket bounded Tooth which in dorsal view
on venter. appears pessular but in

reality is semicircular or
slightly trilobate. In
oblique view it can be
seen that the line of
concrescence deviates in
the neighbourhood of the
posterior tooth so that
only the outside of the
semi-circular tooth is
bilamellar thus enclosing
a monolamellar core formed
by the invagination of the

line of concrescence.
DIMENSIONS (mm).

EL H WwW
Io. 3096 Carapace male 0-80 0°44 0-41
Io. 4349 Carapace female (holotype) 0-74 0-44 0°39
Io. 3097 Left valve female 0-75 0°45 —
Io. 3098 Right valve female 0-76 0°44 —
Io. 4295 Right valve male 0-80 0°45 —-

CoMPARISON. The presence of a reduced ventral ridge, which does not reach the
posterior marginal rim, separates this species from Patagonacythere tricostata Hart-
mann, Patagonacythere devexa (Miiller) and Patagonacythere longiducta antarctica
Benson. Patagonacythere tricostata Hartmann (1962) has a smaller and more
elongate carapace and three rather than two frontal scars. Patagonacythere devexa
(Miller) Benson (1964) is larger, has narrow anterior and posterior vestibules, split
adductor scars and ill-defined median and dorsal ridges including the posterodorsal
loop. (Comparative material of this species from the British Antarctic region was
obtained through the courtesy of Dr. R. C. Whately of Aberystwyth.) Patagona-
cythere longiducta antarctica Benson (1964) is about the same size, has a more concave
posterodorsal slope and the two median adductor scars are split.

REMARKS. The present species commonly occurs in the Zao River and Rakhi
Nala sections, where it has a short vertical range. It is thus a very useful species for
correlation in this region. Patagonacythere has so far only been described from cold
water regions, but P ? indulus occurs here in a warm water environment. A new
generic assignment will therefore probably be necessary at a later time.

Genus PHALCOCYTHERE nov.

DERIVATION OF NAME. Greek phalkes, beam or rib of a ship ; with reference to
the ventral ridge + cythere.

FROM WEST PAKISTAN 57

Diacnosis. Reticulate Trachyleberididae with a ventral ridge ; with or without
spines or papillae ; mostly with pronounced posterodorsal process.

TYPE SPECIES. Cythere horrescens Bosquet 1852.

DESCRIPTION. Sexual dimorphism present in most of the species. Carapace
subrectangular to subquadrate in lateral outline. Anterior margin broadly rounded,
posterodorsal margin very slightly concave, posterodorsal margin either curved or
almost straight, dorsal margin almost straight or slightly convex (but appears
irregular in lateral outline in many species due to surface ornamentation), ventral
margin slightly concave in front of the middle or nearly straight (over-reached by a
ventral ridge in lateral view in some species). Valves almost equal in size although
the right valve over-reaches the left at the anterior margin. Subcentral-tubercle
more or less well-developed. Eye-tubercle distinct. Surface reticulate with or
without superimposed papillae or spines. A posterodorsal process is generally
present. A ventral ridge more or less prominent is always present ; it is either
straight or slightly curved convexly downward in the middle culminating in the
posterior fourth usually in a spine or an ala. Anterior and posterior marginal rims
always present, more or less distinct. Radial pore canals simple, almost straight,
irregularly spaced, sometimes crossing one another, fairly numerous (approximately
30 anteriorly in the type species). Line of concrescence and inner margin coincide.
Duplicature fairly wide. Selvage more or less pronounced, submarginal in left valve
but at some distance in the right valve. Right valve with a deep and well-developed
anterior and ventral flange groove. Adductor scars in a vertical column of four
elongate scars with two almost rounded frontal scars (see description of P. horrescens).
Hinge holamphidont.

COMPARISON. Hirsutocythere Howe 1951 has a wider duplicature and lacks a
ventral ridge. Avustralicythere Benson 1964 is a much larger genus in which fine
pittings occur within the reticulae. The two median adductor scars are also divided
into two. Moreover, Australicythere has a posterior vertical ridge and a less promin-
ent ventral ridge not ending in a spine posteriorly. Bradleya Hornibrook 1952 has
both dorsal and ventral ridges, in this respect it differs from Phalcocythere which has
only a ventral ridge.

REMARKS. This genus is so far known from the Eocene of the Paris Basin, West
Pakistan, Tanzania and an undescribed species from the Aquitaine Basin.

Phalcocythere horrescens (Bosquet)
(Plate 29, fig. 5 ; Plate 30, figs. 1-6 ; Plate 33, figs. 12, 13)

1852 Cythere horvescens Bosquet, p. 119, pl. 6, fig. 5.

1852 Cythere thierensiana Bosquet (pars), p. 98.

1852 Cythere nebulosa Bosquet, p. 105, pl. 5. fig. 8.

1955 Tvachyleberis horrescens (Bosquet), Apostolescu, p. 272, pl. 8, figs.125—-126.
1957 Hirsutocythere horrescens (Bosquet), Keij, p. 101, pl. 15, fig. 4 ; pl. 17, figs.6—7.

Diacnosis. Phalcocythere in which posteroventral margin is straight in left valve
but curved in the right with five or six large spines, shell surface ornamented by well-

58 EARLY TERTIARY OSTRACODA

developed spines superimposed on reticulations ; ventral ridge and posterodorsal
process well-marked.

FIGURED SPECIMENS. Io. 4253-7.

MATERIAL. 8 specimens from Grignon, Paris Basin, from the Lutetian IV (sample
no. CAB 1002, Keij 1957, p. 19). 5 specimens from Villiers-St.-Frederic, Paris
Basin, from the same horizon.

TYPE LOcALITy. Grignon, Paris Basin.
TYPE HORIZON. Lutetian.
DIMENSIONS (mm).

L H
Io. 4253 Left valve 0:60 0°33
Io. 4257. Left valve 0°59 0°33
Io. 4254 Left valve 0-72 0°37
Io. 4255 Right valve 0°59 0°33
Io. 4256 Right valve 0°63 0:37

CoMPARISON. This species shows some affinity to Phalcocythere retispinata sp. nov.
but has a more spinose surface, less tapering carapace and a better developed
subcentral-tubercle. Further, P. horrescens has straight posteroventral margin in
the right valve, different posterodorsal process and less prominent eye-tubercle which
lies below and slightly posterior to the anterior cardinal angle.

ReEMARKS. The present species has been redescribed by Keij (1957) in detail,
where, although he ascribed it to the genus Hirsutocythere, he noted that it lacked the
very wide duplicature of that genus.

Adult specimens vary in size. According to Apostolescu (1955, p. 272), they
range from 0:52 mm. to 0-70 mm. in length.

Phalcocythere improcera sp. nov.
(Plate 30, figs. 7-12 ; Plate 31, figs. 1-4)
DERIVATION OF NAME. Latin, «mprocerus, short ; with reference to the carapace.

Diacnosis. A small Phalcocythere in which the prominent ventral ridge possesses
an alar expansion ; posteroventral margin slightly protracted towards the venter,
subcentral-tubercle prominent.

Hootype. Io. 4344, a male carapace (PI. 31, figs. 1, 2 ; Pl. 30, figs. 8, 9).

PARATYPES. Io. 4296 + Io. 4258-9.

MATERIAL. 68 specimens from the Sor Range Section from four horizons (sample
nos. 460-1, 460-j, 460-1 and 460-n). GSP BM 2570-1.

TYPE LOCALITY. Sor Range section.

TYPE HORIZON. Upper Palaeocene, sample no. 460-i.

DESCRIPTION. Carvapace subrectangular in the male dimorph and sub-quadrate
inthe female. Sexual dimorphy is rather marked ; the males are longer in proportion

FROM WEST PAKISTAN 59

than the females. Anterior margin broadly rounded, posterodorsal margin very
slightly concave, posteroventral margin rounded and slightly protracted towards the
venter. Dorsal margin straight but appears irregular in lateral view because of the
surface ornamentation ; ventral margin nearly straight (concealed in side view by
the ventral ridge). Anterior and posterior cardinal angles well-developed in the right
valve but more or less rounded in the left valve. Right valve slightly over-reaches
the left at the anterior margin but the left valve over-reaches the right in the region
of the posterodorsal slope. Swbcentral-tubercle prominent. Fye-tubercle distinct,
rounded and glassy. Shell surface deeply reticulate with superimposed papillae or
spines, the ornamentation extending onto the prominent ventral ridge (over-reaching
the ventral margin in lateral view) which also develops an alar expansion. The
posterodorsal process is developed as a short curved horn-like ridge in specimens with
papillose ornamentation but is a projecting, blade-like process in specimens having a
spinose surface. Anterior and posterior marginal rims distinct. Anterior margin
finely denticulate with 20-22 denticles, posteroventral margin with 7-8 short spines.
Normal pore canals fairly numerous, one to each reticule. (These become exagger-
ated in specimens cleaned in the ultrasonic vibrator). Radial pore canals not very
well preserved but appear to be simple, more or less straight, irregularly spaced,
some crossing one another, approximately 35 anteriorly. Line of concrescence and
imner margin coincide. Duplicature moderately wide. Selvage prominent—sub-
marginal in the left valve but situated in the outer two-fifths of the duplicature in
right valve which also has a deep and well marked anterior and ventral flange groove.
Hinge not clearly seen but appears to be holamphidont.

DIMENSIONS (mm).

L H W
Io. 4344 Carapace male (holotype) 0-49 0:29 0-27
Io. 4258 Carapace female 0°49 0°30 0:29
Io. 4295 Left valve female 0-46 0-28 —
Io. 4296 Right valve female 0:46 0:27 —

COMPARISON. Distinguished from the other known species of the genus Phalco-
cythere by its small size and the posteroventral margin slightly drawn out towards
the venter particularly in the right valve.

REMARKS. The surface ornamentation is variable. Most of the specimens
examined have a combination of reticulations and papillae but in a few specimens
spines are superimposed on reticulations.

Phalcocythere rete sp. nov.
(Plate 31, figs. 5-12)
DERIVATION OF NAME. Latin rete, net ; with reference to the surface ornament-
ation.

Diacnosis. Reticulate Phalcocythere in which eye-tubercle is prominent, ventral
ridge present but not prominent, dorsal margin slightly convex particularly in
female.

60 EARLY TERTIARY OSTRACODA

HouotyPe. Io. 4348, a female right valve (PI. 31, figs. 11).
PARATYPES. Io. 4297 + Io. 3099 + Io. 3141.

MATERIAL. 14 specimens including the holotype from the Sor Range section from
one horizon (sample no. 460-1). GSP BM 2572-3.

TYPE LOCALITY. Sor Range section.
TYPE HORIZON. Upper Palaeocene, sample no. 460-1.

DESCRIPTION. Sexual dimorphy rather strong ; the males are longer, less high and
less wide than the females. Cavapace subrectangular to subquadrate in lateral view
with a slight taper towards the posterior. Dorsal margin slightly convex particularly
in the female, ventral margin almost straight, anterior broadly and evenly rounded,
posterodorsal slope very slightly concave, posteroventral margin rounded. Left
valve slightly over-reaches the right at the anterior margin. Swbcentral-tubercle
distinct, eye-tubercle prominent, rounded and polished. Shell surface reticulate, the
reticulae are slightly papillose. The ventral ridge is present but not prominent,
sloping obliquely upward towards the posterior and culminating in the posterior
third. Marginal rim distinct. Anterior and posterior margins denticulate. Radial
pore canals simple, straight, slightly thicker on the proximal end, irregularly spaced,
about 30 anteriorly. Inner margin and line of concresence coincide. Duplicature
fairly wide, 0-060 mm. anteroventrally. Selvage well-developed—submarginal in
left valve but almost in the middle in the right valve. There is a deep anterior and
ventral flange groove in the right valve. Hinge not clearly distinguished due to
mineralization but presumably holamphidont.

DIMENSIONS (mm).

L H W
Io. 3099 Carapace male 0-65 0°35 0°24
Io. 4297 Carapace female 0°47 0°35 0:27
Io. 3141 Left valve female 0-46 0°35 a
Io. 4348 Right valve female (holotype) 0-46 0°35 —

CoMPARISON. Phalcocythere retispinata sp. nov. is a closely related species but has
a reticulate and spinose surface, a more elevated ventral ridge and a well-developed
posterodorsal process.

Phalcocythere retispinata sp. nov.
(Plate 31, figs. 13-17 ; Plate 32, figs. 1-3)
DERIVATION OF NAME. Latin rete, net + spinatus, spined ; with reference to the
surface ornamentation.

Diacnosis. Phalcocythere with a prominent ventral ridge with alar expansion,
surface ornamentation a combination of reticulations and spines, subcentral-tubercle
present but not pronounced, eye-tubercle prominent, posterodorsal process well-
marked.

Ho.otyPe. Io. 4345, a female carapace (PI. 31, figs. 15, 16 ; Pl. 32, figs. 2, 3).

PARATYPE. Io. 3165.

FROM WEST PAKISTAN 61

MATERIAL. Six specimens from the below locality from three horizons (sample
nos. 460-1, 460-j and 460-0). GSP BM 2574.

TYPE LOCALITY. Sor Range section.

TYPE HORIZON. Upper Palaeocene, sample no. 460-1.

DESCRIPTION. Sexual dimorphism rather pronounced ; the females are higher and
wider than the males. Cavrapace tapering towards the posterior, subrectangular in
side view. Dorsal margin nearly straight in the male dimorph but very slightly
convex in the female (appears to be irregular in dorsal view due to ornamentation) ;
ventral margin almost straight but concealed by the ventral ridge in side view ;
anterior broadly rounded, posterodorsal margin very slightly concave ; postero-
ventral margin rounded. Greatest length lies below mid-point, greatest height in
the anterior third. Anterior and posterior cardinal angles well-developed. Right
valve very slightly over-reaches the left at the anterior margin. Swubcentral-tubercle
present but not pronounced. LEvye-tubercle prominent, rounded and polished. Orna-
mentation consists of reticulations and spines. The spines are of variable size. A
posterodorsal process consists usually of a large spine, which stands out in lateral
view. The ventral ridge is high and with an alar expansion, slightly concave in the
middle culminating in the posterior third with a pointed end. Anterior and posterior
marginal rims more or less distinct and decorated with a row of spines. Anterior
and posterior margins denticulate. Internal characters not seen.

DIMENSIONS (mm).

IG; H Ww
Io. 3165 Carapace male 0-64 0°37 0°27
Io. 4345 Carapace female (holotype) 0°64 0°39 0°32

CoMPARISON. Phalcocythere tmprocera sp. nov. is much smaller, has deeper
reticulations and a posteroventral margin slightly drawn out towards the venter.

Phalcocythere sentosa sp. nov.
(Plate 32, figs. 4-10)
DERIVATION OF NAME. Latin sentosus, rough ; with reference to the surface
ornamentation.

Diacnosis. A species of the genus Phalcocythere in which ventral ridge is present
but not high ; surface ornamentation consists of combination of reticulations and
papillae ; posterodorsal process a small tubercle or short spine. Subcentral-tubercle
distinct, eye-tubercle prominent.

HoLotyPe. Io. 4346, a male carapace (PI. 32, figs. 4, 5, 8, 10).
PARATYPE. Io. 4208.

MATERIAL. 67 specimens from the Rakhi Nala section from 11 horizons (sample
NOS. 3153, 3165, 3167, 3169, 3170, 3173 to 3177 and 3180). GSP BM 2575-6.

TYPE LocALITy. Rakhi Nala section.
TYPE HORIZON. Upper Rakhi Gaj Shales, sample no. 3167.

62 EARLY TERTIARY OSTRACODA

DESCRIPTION. Strongly dimorphic, the females are less elongate than the males.
Carapace subrectangular in lateral view. Anterior broadly rounded, posterodorsal
slope very slightly concave, posteroventral margin rounded, dorsal margin almost
straight, appearing irregular in lateral view, ventral margin slightly concave anterior
to the middle. Right valve slightly over-reaches left valve at the anterior margin
but is over-reached by the latter in the region of the posterodorsal slope. Anterior
and posterior cardinal angles well-developed, particularly in the right valve. Evye-
tubercle rounded and prominent, projecting out from the eye-socket. Swbcentral-
tubercle distinct. Surface reticulate with superimposed papillae, posterodorsal
process either a small more or less rounded tubercle or a cluster of short spines ; ventral
ridge present but not elevated. Anterior and posterior marginal rims distinct.
Anterior margin denticulate, posteroventral margin papillose. Internal details not
observed.

DIMENSIONS (mm).

L H W
Io. 4346 Carapace male (holotype) 0:56 0-32 0°25
Io. 4298 Carapace female 0°55 0°32 0-29

CoMPARISON. -Phalcocythere rete sp. nov. is larger than the present species, has a
less papillose surface and slightly convex dorsal margin in the female.

REMARKS. So far only known from the Rakhi Nala section. The posterodorsal
process varies; in some specimens it is almost a rounded tubercle but in others it is a
short spine.

Phalcocythere dissenta sp. nov.
(Plate 32, figs. 11-18)

DERIVATION OF NAME. Latin dis, not + sentus, spiny.

Diacnosis. A reticulate species of the genus Phalcocythere with dorsal and ventral
margins sub-parallel, anterior rim ornamented like a scallop or flute, subcentral-
tubercle prominent, eye-tubercle and ventral ridge distinct.

HoLotyPe. Io. 4343, a male carapace (PI. 32, figs. 11, 14, 18).

PARATYPE. Io. 4299.

MATERIAL. Approximately 400 specimens from the locality above from six
horizons (sample nos. 3454, 3456, 3460 to 3462 and 3464) and 6 specimens from the
Zao River section from one horizon (sample no. 24107). GSP BM 2577-8.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Shales with Alabaster, sample no. 3456.

DESCRIPTION. Sexual dimorphism marked, the presumed males are more
elongate, less high and less wide than the females. Carapace sub-rectangular in lateral
outline with sub-parallel dorsal and ventral margins. Anterior margin broadly
rounded ; posterodorsal slope more or less straight ; posteroventral margin rounded ;
dorsal margin slightly convex particularly in the female ; ventral margin almost
straight, although partly hidden by the ventral ridge in lateral view. Greatest

FROM WEST PAKISTAN 63

length passes through mid-point, greatest height in the anterior third and greatest
width in the anterior two-fifths. Anterior and posterior cardinal angles protruding.
Valves almost equal. Swbcentral-tubercle prominent, eye-tubercle distinct and
situated below the cardinal angle. Anterior marginal rim distinct, and ornamented
with seven very short ridges with small depressions in between (like flutes or scallops),
posterior rim more or less distinct. Surface reticulate with a distinct ventral ridge
which slopes obliquely upwards posteriorly ending in the posterior third. In many
specimens the posterior ending is pointed or spinose. Internal details not determin-
able, all specimens being complete carapaces.

DIMENSIONS (mm).

Ie H WwW
Io. 4343 Carapace male (holotype) 0-60 0-44 0°32
Io. 4299 Carapace female 0°56 0°44 0°44

CoMPARISON. This species shows some resemblance to Phalcocythere rete sp. nov.
but is smaller and has subparallel rather than tapering dorsal and ventral margins.
It differs from Phalcocythere improcera sp. nov. and Phalcocythere sentosa sp. nov. in
shape and surface ornamentation.

REMARKS. P. dissenta seems to be restricted to the Shales with Alabaster only
and has been found in the Rakhi Nala and Zao River sections. It is abundant in the
Rakhi Nala section but rare in the Zao River area.

Phalcocythere spinosa sp. nov.
(Plate 33, figs. 1, 2, 7, 8)

DERIVATION OF NAME. Latin spinosus, spiny.

Diacnosis. A species belonging to the genus Phalcocythere with short spines
and/or papillae superimposed on reticulations ; ventral ridge distinct and terminating
in a spine in posterior third ; posterodorsal process well-marked and blade-like.

HootyPe. lo. 4347, a carapace.

MATERIAL. 16 specimens from the Zao River section from one horizon (sample
no. 24161). GSP BM 2579.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24161.

DESCRIPTION. Carvapace subrectangular in side view. Dorsal margin almost
straight, appearing to be irregular in lateral view due to ornamentation ; ventral
margin more or less straight but hidden by the ventral ridge in side view ; anterior
margin broadly rounded ; posterodorsal margin slightly concave ; posteroventral
margin rounded. Greatest length passes through the mid-point and greatest height
passes through the anterior cardinal angle. Valves nearly equal. Swubcentral-
tubercle well-developed, eye-tubercle rounded, distinct and lies below the anterior
cardinal angle. Surface ornamentation consists of short spines and/or papillae
superimposed on reticulations. A distinct ventral ridge, diagnostic of the genus,

64 EARLY TERTIARY OSTRACODA

over-reaches the ventral margin in lateral view and is spinose posteriorly. It
extends from the anteroventral corner to the posterior third. The posterodorsal
process is projecting and blade-like (present in most specimens). Anterior and
posterior marginal rims distinct. Anterior margin finely denticulate, posteroventral
margin ornamented with 6-7 short spines or papillae. Internal features not seen.
DIMENSIONS (mm).
L H WwW
Io. 4347 Carapace (holotype) 0°54 0°33 0°44
CompaRIson. -Phalcocythere sentosa sp. nov. is similar and perhaps ancestral to the
present species. These two, however, can be separated easily due to the fact that
P. sentosa is strongly dimorphic and has a less well-developed ventral ridge and a
posterodorsal process.

REMARKS. This species has so far only been recorded from one horizon of the
Upper Chocolate Clays of the Zao River section. Sexual dimorphism has not been
observed in this species.

Phalcocythere sp., cf. P. spinosa
(Plate 33, figs. 3-6, 9-II)

FIGURED SPECIMENS. lo. 4230—lo. 4232.

MATERIAL. Five specimens from the locality and horizon below (other specimens
in the collections of the British Petroleum Co. Ltd., under registration no. FCRM
1648). GSP BM 2580.

Locatity. Lindi survey, 10-50 ft. above shore at Kitunga, Tanzania.
Horizon. Upper Eocene.

DESCRIPTION. Sexual dimorphism rather marked ; the females are higher and
wider than the males. Carapace subrectangular to subquadrate in lateral outline.
Dorsal margin irregular in lateral view due to ornamentation ; ventral margin almost
straight ; anterior margin broadly rounded ; posterodorsal margin very slightly
concave ; posterior extremity rounded. Right valve over-reaches the left slightly
at the anterior margin. Anterior and posterior cardinal angles well-developed
particularly in right valve. Greatest length below mid-point, greatest height at
anterior cardinal angle and greatest width in the posterior third. Swubcentral-tubercle
distinct. Evye-tubercle rounded and prominent. Anterior and posterior marginal
rims distinct. Surface ornamentation consists of reticulations with superimposed
spines ; ventral ridge prominent, posteriorly alate, ending abruptly in the posterior
third ; posterodorsal process prominent and blade-like, standing out in lateral and
dorsal views. Anterior margin ornamented with numerous very short spines ;
posterior with six larger spines. Radial pore canals not discernible. Duplicature
moderately wide. Selvage strong—marginal in left valve but in right valve it is in
the outer third. Right valve with a deep flange groove. Muscle scars are in a
vertical row of four adductors, and frontal scars not seen. Hinge holamphidont
with the details of each element as follows:

FROM WEST PAKISTAN 65

Element Left valve Right valve

Anterior Deep rounded socket. Projecting conical tooth.
Eye-socket opens below
and slightly anterior to

this tooth.
Anteromedian Conical projecting tooth. Socket opening into groove.
Posteromedian Denticulate bar. Shallow locellate groove
narrowing towards posterior.
Posterior Elongate groove, Subpessular tooth, higher
presumably deep (filled on the posterior side.
in with matrix).
DIMENSIONS (mm).
L H WwW
Io. 4230 Carapace male 0°55 0°32 0°30
Io. 4231 Carapace female 0°56 0°34 0°34

Io. 4232 Right valve female (broken)—4w, 0-17

Comparison. Phalcocythere spinosa sp. nov. closely approaches the present form
but is smaller, has a less reticulate and spinose surface and a more concave postero-
dorsal slope. Phalcocythere retispinata sp. nov. is much larger, has a slightly convex
dorsal margin particularly in the female dimorph, and the carapace tapers towards
the posterior.

REMARKS. The specimens studied were made available through the kindness of
Dr. F. E. Eames, lately Chief Palaeontologist of the British Petroleum Co. Ltd.
These specimens may represent a distinct sub-species of P. spinosa.

Genus QUADRACYTHERE Hornibrook 1952

TYPE SPECIES. Cythere truncula Brady 1808.

Subgenus HORNIBROOKELLA Moos 1965

TYPE SPECIES. Cythere anna Lienenklaus 1894.

Quadracythere (Hornibrookella) platybomus sp. nov.
(Plate 33, figs. 14, 15, 18, 19).
DERIVATION OF NAME. Greek flatys, broad + bomos, bottom ; with reference to
the expanded venter.

Diacnosis. Carapace with expanded venter, subrectangular to subquadrate in
lateral outline.

Houotype. Io. 4351, a male carapace (PI. 33, figs. 14, 18).

PARATYPE. Io. 4300.

MATERIAL. Nine specimens from the locality below from two horizons (sample
nos. 460-i and 460-0). GSP BM 2581-2.

E

66 EARLY TERTIARY OSTRACODA

TYPE LOCALITY. Sor Range section.
TYPE HORIZON. Upper Palaeocene, sample no. 460-1.

DESCRIPTION. Cavrapace subrectangular in the male dimorph and subquadrate in
the female. Sexual dimorphism rather apparent ; the females are higher and wider
than the males. Carapace compressed in the posterior region. Anterior margin
broadly rounded, posterodorsal slope very slightly curved, posterior almost rounded,
posteroventral margin slightly curved. The ornamentation over-reaches the dorsal
margin giving a jagged appearance, ventral margin almost straight in the male but
slightly curved in the female. Left valve very slightly over-reaches the right at the
posterodorsal slope and at the anterior cardinal angle. In dorsal view the greatest
width is situated anterior to the middle. Swbcentral-tubercle prominent, eye-tubercle
distinct. A marginal rim runs around the anterior, ventral and posterior margins.
It is upraised around the anterior but less elevated along the venter and around the
posterior end. Surface reticulate with a well-marked ventral ridge giving rise to an
expanded venter. At the posterodorsal corner a small horn-like projecting ridge is
present. Anterior margin ornamented by small, numerous denticles (20-25 in
number) but the posterior has only a few denticles.

DIMENSIONS (mm).

L H W
Io. 4351 Carapace male (holotype) 0°57 0°32 0°24
Io. 4300 Carapace female 0°57 0°34 0°29

CoMPARISON. Quadracythere (Hormbrookella) directa sp. nov. is larger, has a less
well-developed subcentral-tubercle and lacks an expanded venter.

REMARKS. The preservation of the material prevents a description of the internal
characters. So far this species is only known from the Upper Palaeocene of the Sor
Range section.

Quadracythere (Hornibrookella) directa sp. nov.
(Plate 33, figs. 16, 17 ; Plate 34, figs. I, 2)
DERIVATION OF NAME. Latin divectus, rectangular ; with reference to the outline
in lateral view.

Diacnosis. In lateral view carapace subrectangular with protruding anterior and
posterior cardinal angles. Surface ornamentation consists of reticulation with an
oblique ventral ridge sloping upward towards posterior and a short horn-like ridge at
posterodorsal corner. Sexual dimorphism pronounced.

Ho.otype. Io. 4350, a female carapace (PI. 33, figs. 17 ; Pl. 34, fig.2).
PARATYPE. Io. 4301.

MATERIAL. 96 specimens from the type locality from four horizons (sample
nos. 3184, 3192, 3402 and 3403). GSP BM 2583-4.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Green and Nodular Shales, sample no. 3403.

FROM WEST PAKISTAN 67

DESCRIPTION. Sexual dimorphy rather apparent ; the females are shorter, higher
and wider than the males. Cavapace subrectangular in side view with the greatest
height at the anterior cardinal angle and the greatest length below the middle.
Anterior margin broadly and evenly rounded, posterodorsal slope very slightly
concave, posterior extremity subtriangular, posteroventral margin almost straight.
Dorsal margin very slightly undulating, venter slightly incurved in front of the middle.
Posterior portion of carapace compressed. Anterior marginal rim high continuing
along the venter and around the posterior end as a somewhat less elevated rim.
Anterior margin set with numerous small and delicate denticles, posterior with a few
denticles. Anterior and posterior cardinal angles protruding. Left valve over-
reaches the right valve slightly at the anterior cardinal angle and posterodorsal
slope. In dorsal view the greatest width is situated almost in the middle of the
carapace. Evye-tubercle distinct. Swubcentral-tubercle more or less distinct. Surface
reticulate with a ventral ridge, which slopes slightly upwards towards the posterior.
A short curved, hornlike ridge at the posterodorsal corner is present (better seen in
dorsal view). Internal details not known.

DIMENSIONS (mm).

L H WwW
Io. 4301 Carapace male 0:68 0°35 0:27
Io. 4350 Carapace female (holotype) 0°63 0°37 0:29

COMPARISON. Quadracythere (Hornibrookella) subquadra sp. nov. is subquadrate
in lateral outline, has deeper reticulations and a better developed subcentral-tubercle.

Quadracythere (Hornibrookella) arcana (Lubimova and Guha)
(Plate 34, figs. 3-5)
1960 Cythereis avcanus (sic., recte Cythereis arcana) Lubimova and Guha, p. 33, pl. 3, fig. 1a—b.

D1AGNOsIs. Carapace with distinct caudal process. Surface coarsely reticulate
with superimposed longitudinal lineations in posterior half of carapace.

FIGURED SPECIMEN. Io. 3142.

MATERIAL. Two specimens from the locality and horizon below.
Locatity. Rakhi Nala section.

Horizon. Lower Chocolate Clays, sample no. 3499.

DESCRIPTION. Carapace thick-shelled, subquadrate in lateral view. Anterior
margin broadly rounded, posterior with a pronounced caudal process. Dorsal margin
slightly concave behind the round and protruding anterior cardinal angle, particularly
in the right valve. Ventral margin almost straight. Greatest height at the anterior
cardinal angle and greatest length below the middle. Left valve slightly over-
reaches the right in the region of the anterior cardinal angle and posterodorsal slope.
Evye-tubercle rounded and distinct, swbcentral-tubercle well-developed. Surface
ornamentation consists of coarse reticulations with superimposed longitudinal
lineations in the posterior half of the carapace. The ventral ridge is slightly concave
downwards culminating in an ala posteriorly. A short ridge at the posterodorsal

68 EARLY TERTIARY OSTRACODA

corner meets the dorsal margin at an angle (better seen in dorsal view) and ends as an
ala at the posterior. A marginal rim runs round the anterior, along the venter and
round the posterior margin.

DIMENSIONS (mm).

IL, H WwW
Io. 3142 Carapace 0°54 0°32 0°32

REMARKS. Topotype material was not available for study, and it is, therefore,
difficult to determine whether or not the Rakhi Nala specimens are conspecific with
those from the type locality in Kutch.

Quadracythere (Hornibrookella) subquadra sp. nov.
(Plate 34, figs. 6-11)

DERIVATION OF NAME. Latin subquadrus, almost square ; with reference to the
outline in lateral view.

Diacnosis. Carapace subquadrate with dorsal and ventral margins almost
straight and subparallel. Surface strongly and coarsely reticulate. Sexual dimorphy
moderate.

HoLotype. Io. 4352, a female carapace (PI. 34, figs, 7, 10, II).
PARATYPE. Io. 4302.

MATERIAL. 41 specimens from the locality below from one horizon (sample
no. 24161). GSP BM 2585-6.

TYPE LOCALITY. Zao River section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 24161.

DESCRIPTION. Carapace subquadrate in lateral outline with a short caudal process.
Anterior margin broadly rounded, posterodorsal slope slightly concave, posterior
extremity almost straight, posteroventral margin curved. Dorsal and ventral
margins nearly straight and sub-parallel. Greatest height at the anterior cardinal
angle, greatest length below the mid-point. In dorsal view the greatest width is
situated in front of the middle. Anterior cardinal angle rounded, posterior cardinal
angle well-developed. Left valve over-reaches the right valve slightly in the region
of the posterodorsal slope and the anterior cardinal angle. Swbcentral-tubercle
prominent, eye-tubercle rounded and distinct. Surface coarsely and deeply reticulate.
There is a distinct ventral ridge at some distance from the venter and a short
curved hornlike ridge at the posterodorsal corner. Anterior marginal rim prom-
inent continuing along the venter and around the posterior margin as a less
prominent rim. Anterior margin ornamented with numerous denticles.

Sexual dimorphism moderate ; the presumed females are wider than the males.

DIMENSIONS (mm).

L H WwW
Io. 4302 Carapace male 0:66 0:40 0°32
Io. 4352 Carapace female (holotype) 0-67 0°42 0:34

FROM WEST PAKISTAN 69

CoMPARISON. Quadracythere (Hornibrookella) arcana (1960) is smaller, has a well-
developed caudal process, superimposed longitudinal lineations and a ventral ridge
slightly curved downward. Quadracythere (Hornibrookella) platybomus sp. nov. is
also smaller, has a different lateral outline, less deep reticulations and an expanded
venter.

REMARKS. Quadracythere (Hormbrookella) subquadra sp. nov. commonly occurs
at one horizon (sample no. 24161) of the Upper Chocolate Clays in the Zao River
section associated with Echinocythereis multibullata sp. nov. and Phalcocythere spinosa
sp. nov.

Quadracythere (Hornibrookella) sp.A
(Plate 34, figs. 12-14)

FIGURED SPECIMEN. lo. 3143.

MATERIAL. Only one specimen from the locality and horizon below.
Locatity. Zao River section.

Horizon. Upper Chocolate Clays, sample no. 24148.

DESCRIPTION. Carapace subquadrate in side view. Dorsal and ventral margins
almost straight, anterior margin broadly and obliquely rounded, posterodorsal
margin slightly concave, posteroventral margin rounded. Greatest length lies below
the middle, greatest height at the anterior cardinal angle. In dorsal view the great-
est width lies anterior to the middle. Valves almost equal. LEye-tubercle distinct,
rounded. Swbcentral-tubercle distinct. Surface coarsely and deeply reticulate.
Reticulae are somewhat concentrically arranged around the subcentral-tubercle.
Dorsal ridge present but not well-defined, ventral ridge more or less distinct. A low
marginal rim runs around the anterior and posterior margins. Anterior and postero-
ventral margins denticulate. Internal characters not observed.

DIMENSIONS (mm).

L H W
To. 3143 Carapace 0:93 0°54 0:56

ComPARISON. This is similar to Quadracythere (Hornibrookella) subquadra sp. nov.
but is much larger, has coarser and deeper reticulations more or less arranged in a
concentric pattern around the subcentral-tubercle. In addition, these two species
differ markedly in their posterior outline.

Genus STIGMATOCYTHERE nov.
DERIVATION OF NAME. Greek stigma, mark ; with reference to the ornamentation
+cythere.

Dracnosis. Highly ornamented Trachyleberididae in which two ridges spring
from the eye-tubercle, one to form a high anterior marginal rim, the other curving
sharply round to join the subcentral-tubercle.

TYPE SPECIES. Stigmatocythere obliqua sp. nov.

70 EARLY TERTIARY OSTRACODA

DESCRIPTION. Carapace subrectangular in lateral outline. Sexual dimorphy
apparent ; the males are longer, less high and less wide than the females. Left valve
slightly over-reaches the right in the region of the anterior cardinal angle and at the
posterodorsal slope. Subcentral node and eye-tubercle distinct. Surface reticu-
late, spiny, or a combination of reticulations and spines or with only one to three
longitudinal ridges or lines of ornamentation. The dorsal ridge or line when
developed may be straight or arched convexly upwards. A strongly curved ridge,
diagnostic of the genus, runs from the eye-tubercle to the anterodorsal corner of the
subcentral complex and this may be continued posteriorly either as a ridge or a line
of tubercles. The anterior marginal rim also springs from the eye-tubercle and is
more or less elevated in the anterior region, continuing as a less elevated rim round
the venter and posterior margins. A ventral ridge or line of tubercles diverges
posteriorly from the ventral marginal rim. Anterior and posterior margins spinose.
Normal pore canals simple, medium, some 60 in the female left valve. Radial pore
canals simple, straight, irregularly spaced, some crossing one another, 24—26 anteriorly.
Inner margin and line of concrescence coincide. Anterior duplicature moderately
wide. Selvage well-marked in both valves, submarginal in the left valve but almost
in the outer third of the duplicature in right valve. In right valve a well-developed
flange groove is present along the ventral margin and around the anterior margin.
Muscle scars consist of four adductor scars in an almost vertical column with an oval
frontal scar. Hinge holamphidont ; right valve hinge with a strongly projecting
anterior tooth followed by anteromedian socket, posteromedian locellate groove
present or reduced to a narrow shelf, a posterior tooth, projecting reniform or pessular.
Left valve hinge with anterior and posterior sockets, a conical anteromedian tooth and
a posteromedian denticulate or almost smooth bar.

COMPARISON. Stigmatocythere differs from the genus Gyrocythere in the arrange-
ment of the longitudinal ridges and by having a strongly curved ridge connecting the
eye-tubercle and the subcentral-tubercle. The anterior marginal rim is less evident
in Gyrocythere while it is well developed in Stigmatocythere. Costa has three or four
continuous longitudinal ridges, the median or second of which has a characteristic
posterior termination absent in Stigmatocythere and lacks the anterior connection
to the eye-tubercle present in Stigmatocythere. Bradleya has only dorsal and ventral
ridges. Carinocythereis Ruggieri 1956 has a V-shaped frontal scar and small vesti-
bules, characters not found in Stigmatocythere.

REMARKS. Stigmatocythere is so far only known from the Middle and Upper
Eocene of the Sulaiman Range.

Stigmatocythere obliqua sp. nov.
(Plate 35, figs. 1-10 ; Plate 36, figs. 1-2)
DERIVATION OF NAME. Latin obliqua, oblique ; with reference to the ventral
ridge.
Dracnosis. A strongly reticulate species of Stigmatocythere with three well-
developed longitudinal ridges including an oblique ventral ridge.

FROM WEST PAKISTAN 71

Ho.otypPe. Io. 4355, a female carapace (Plate 35, figs. 2, 5, 6 ; Plate 36, fig. 2).
PARATYPES. Io. 4303 + Io. 3147 + Io. 3148 + Io. 3149.

MATERIAL. Over 1400 specimens from the Rakhi Nala section from 17 horizons
(sample nos. 3448, 3451 to 3454, 3456, 3457, 3460 to 3467, 3470 and 3473). 470
specimens from the Zao River section from two horizons (sample nos. 24107 and
24110). Approximately 600 specimens from the Shpalai Khwara section from three
horizons (sample nos. 24681, 24683 and 24686). GSP BM 2587-8.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Shales with Alabaster, sample no. 24173.

DESCRIPTION. Carapace subrectangular in lateral view. Sexual dimorphism
rather pronounced ; the females are shorter, higher and wider than the males.
Anterior margin broadly and obliquely rounded, posterior nearly straight. Dorsal
margin almost intricate because of the over-reaching of the dorsal ridge, venter
slightly incurved in front of the middle. Valves almost equal except that the left
valve over-reaches the right in the region of the anterior cardinal angle and along the
posterodorsal corner. Evye-tubercle and subcentral-tubercle distinct. Shell surface
strongly reticulate. Three longitudinal ridges present ; the dorsal ridge starts
almost above the subcentral node and is arched convexly upwards ; the median ridge
commences from the eye-tubercle, curves sharply round to join anterodorsal corner
of the subcentral complex and continues posteriorly ; the ventral ridge slopes
obliquely upwards towards the posterior. The reticulations in the mid-posterior
region of the carapace show a concentric pattern, although it is not always present.
A high anterior marginal rim commences from the eye-tubercle continuing as a less
high rim round the venter and posterior. Anterior and posterior short marginal
spines present, 18-20 anteriorly. Normal pore canals simple, some 60 in the female
left valve. 24-25 anterior radial pore canals, simple straight, irregularly spaced, few
crossing one another, mostly terminating in marginal spines. Inner-margin and line
of concrescence coincide. Anterior duplicature 0-050 mm. wide in the female left valve.
Selvage strong in both valves, subperipheral in left valve but in right valve it lies
in the outer third of the duplicature. Right valve with deep anterior and ventral
flange grooves. Muscle scar pattern consists of four adductor scars in a vertical row
with an oval frontal scar. Hinge holamphidont, the details are as follows :

Element Left valve Right valve
Anterior Socket confluent with Strongly projecting sub-
ocular sinus. conical or subpessular tooth
with a tendency for the

anterior profile in dorsal
view to appear concave.
The ocular sinus lies
distally beyond this and
opens to the interior below
and in front of it.

72 EARLY TERTIARY OSTRACODA

Anteromedian Conical tooth with Deep socket.
straight anterior in
dorsal view.

Posteromedian Denticulate bar. Locellate shelf, only
detectable in best pre-
served specimens.

Posterior Deep socket, slightly Pessular tooth, tending

elongate, open in centre. towards reniform in some
specimens.

DIMENSIONS (mm).
1g H Ww

Io. 4303 Carapace male 0-61 0°30 0:29
Io. 4355 Carapace female (holotype) 0°54 0°33 0°29
Io. 3149 Left valve female 0°52 0°34 —
Io. 3148 Right valve female 0°54 0°32 —
Io. 3149 Left valve female 0°54 0°34 —
Io. 3146 Right valve female 0-51 0:29 _

CoMPARISON. This species is perhaps ancestral to Stigmatocythere portentum sp.
nov. which it resembles very closely, but differs in being smaller, having deeper
reticulations and less prominent subcentral-tubercle.

REMARKS. The present species occurs abundantly in the Shales with Alabaster
of the Rakhi Nala, Zao River and Shpalai Khwara sections and seems to be restricted

to this formation. It is very likely that this species will prove to be a valuable
horizon marker in the region.

The longitudinal ridges in some of the specimens from the upper few horizons of the
Rakhi Nala section are exaggerated and in some the dorsal and ventral ridges
posteriorly terminate in spines. This is regarded here as specific variation.

Stigmatocythere portentum sp. nov.
' (Plate 36, figs. 3-6, 10)
DERIVATION OF NAME. Latin fortentwm, omen or sign ; with reference to the
diagnostic ornamentation.

Diacnosis. A large, reticulate species of the genus Stigmatocythere with three
distinct longitudinal ridges, prominent subcentral-tubercle.

HoLotyPe. Io. 4357, a male carapace (PI. 36, figs. 3-6).
PARATYPES. Io. 3144 + Io. 3145.

MATERIAL. Eight specimens from the locality below from two horizons (sample
nos. 3498 and 3499). GSP BM 2551.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Lower Chocolate Clays, sample no. 3498.

DESCRIPTION. Carapace subrectangular in side view with the greatest height at
the anterior cardinal angle and the greatest length above mid-point. Anterior

FROM WEST PAKISTAN 73

margin broadly rounded, posterior truncated. Dorsal margin straight, but looks
irregular in lateral view because of over-reaching of the dorsal ridge ; ventral margin
almost straight. Anterior and posterior cardinal angles well-developed particularly
in the left valve. Left valve slightly over-reaches the right at the anterior cardinal
angle and along the posterodorsal slope. Eye-tubercle distinct and rounded. Sub-
central-tubercle prominent. Surface reticulate with three longitudinal ridges. Dorsal
ridge wavy and convex upwards, it begins above the subcentral-tubercle and in the
posterodorsal region, curves sharply down to meet the median ridge. A strongly
curved ridge runs from the eye-tubercle to the subcentral-tubercle, continuing
posteriorly as a median ridge. Ventral ridge commences above the anteroventral
corner and is slightly convex downwards. There is a short, curved ridge on the
ventral side of the subcentral-tubercle running towards the anteriorend. A marginal
rim runs along the anterior, ventral and posterior margins. It is high on the anterior
but less high along the venter and posterior. Anterior ornamented with short,
numerous spines. Thereisalarge posteroventral spine. Radial pore canals simple.
more or less straight, some crossing one another, few seem to bifurcate, some 25
anteriorly. Inner margin and line of concrescence coincide ; no vestibule. Duplicature
of moderate width with a well-marked selvage. Hinge holamphidont with a pro-
jecting conical anterior tooth in the right valve.

DIMENSIONS (mm).

L H W
Io. 4357 Carapace male (holotype) O-71 0°37 0°32
Io. 3144 Right valve (broken) — 0°37 —-
Io. 3145 Carapace (juvenile) 0-59 0°35 0°29

CoMPARISON. This species has already been compared with Stigmatocythere
obliqua sp. nov.

REMARKS. S. portentum is a very rare ostracod and has so far only been found in
the uppermost beds of the Lower Chocolate Clays of the Rakhi Nala section.

Stigmatocythere calia sp. nov.
(Plate 36, figs. 7-9 ; Plate 37, figs. 1, 3)
DERIVATION OF NAME. Greek, kalia, bird’s nest ; from a fancied appearance of
the ornamentation in lateral view.

Diacnosis. A _ non-reticulate Stigmatocythere with straight posterior, high
anterior marginal rim, prominent and projecting subcentral-tubercle, dorsal and
ventral lines of ornamentation, posteroventral ridge, almost straight.

HoLotyPe. Io. 4353, a female carapace (PI. 36, figs, 8, 9 ; Pl. 37, figs. 1, 3).
PARATYPE. Io. 4304.

MATERIAL. 15 specimens from the locality below from five horizons (sample
nos. 24148, 24150 to 24153). GSP. BM. 2589.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 24151.

74 EARLY TERTIARY OSTRACODA

DESCRIPTION. Sexual dimorphism distinct ; the males are more elongate than
the females. Cavapace subrectangular in lateral outline. Dorsal margin almost
straight but appears undulating in side view due to the over-reaching of the dorsal
line of ornamentation, ventral margin nearly straight. Anterior margin broadly
and evenly rounded, posterior straight, posterodorsal slope almost straight. Left
valve slightly over-reaches the right valve in the anterodorsal and posterodorsal
corners ; otherwise the two valves are equal. LEye-tubercle distinct, rounded and
polished. Swbcentral-tubercle prominent and projecting particularly in dorsal and
ventral views. Dorsal line of ornamentation consists of three nodes including a large
subangular posterodorsal node which extends vertically down for a short distance.
Ventral line of ornamentation consists of two nodes but in some specimens it is just a
short ridge. A projecting, almost vertical posteroventral ridge runs from behind the
ventral line of ornamentation to a point which is slightly above the mid-line. A
curved ridge diagnostic of the genus runs from the eye-tubercle to the subcentral-
tubercle ; it is not well-marked. The subcentral-tubercle has two faint, short,
curved ridges on its ventral side, one to the anterior and one to the posterior. Anterior
marginal rim high, extending along the ventral and posterior margins as a less high
rim. Anterior margin set with numerous short spines, concealed in lateral view by
the elevated anterior marginal rim. There is one short posterodorsal marginal spine
and one short posteroventral marginal spine. Internal details not determinable.

DIMENSIONS (mm).

L H W
To. 4304 Right valve male 0-61 0°34 —
Io. 4353 Carapace female (holotype) 0:56 0:36 0°32

Stigmatocythere delineata sp. nov.
(Plate 37, figs. 2, 4-10)
DERIVATION OF NAME. Latin delineata, outlined ; from the resemblance of the
ornamentation to sketch map.

Diacnosis. A species of the genus Stigmatocythere with a large hexagon formed of
ridges in the posteromedian region, dorsal ridge broken in the middle and extending
vertically below in the posterodorsal region.

Ho.LotyPe. Io. 4356, a female carapace (PI. 37, figs. 7-10).

PARATYPE. Io. 4305.

MATERIAL. Six specimens from the type locality from two horizons (sample
nos. 24154 and 24155). GSP BM 2590-01.

TYPE LOCALITY. Zao River section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 24154.

DESCRIPTION. Sexual dimorphism apparent ; the males are proportionally
longer than the females. Cavapace subrectangular in side view. Dorsal margin
slightly concave, ventral margin almost straight, anterior margin broadly and evenly
rounded, posterodorsal slope and posteroventral margin nearly straight. Greatest

FROM WEST PAKISTAN 75

height lies at the anterior cardinal angle, greatest length above the mid-point and
greatest width in the posterior third. Anterior and posterior cardinal angles pro-
jecting particularly in the left valve. Left valve slightly over-reaches the right valve
in the region of the anterodorsal corner and posterodorsal slope. Swubcentral-tubercle
and eye-tubercle distinct. The most prominent part of the ornamentation is a large
slightly irregular hexagon formed of ridges just to the posterior of centre. Other
short ridges, mostly running outwards, join this hexagon at its corners. A sharply
curved ridge characteristic of the genus connects the eye-tubercle and the subcentral
tubercle. Dorsal ridge is broken in the middle and in the posterodorsal region it
extends vertically below to a point slightly above mid-line. The ventral ridge runs
at a slightly oblique angle towards the posterior end, its posterior portion forming the
ventral part of the hexagon. A high marginal rim runs on around the anterior margin
and continues along the ventral and posterior margins as a less high rim. Anterior
margin denticulate, posteroventral corner ornamented with a short spine. Dup-
licature of moderate width with a strong seluage. Hinge holamphidont.

DIMENSIONS (mm).

Lb H W
Io. 4305 Carapace male 0-61 0°34 0-29
Io. 4356 Carapace female (holotype) 0:56 0:34 0°29

COMPARISON. It is easy to separate S. delineata from other described species of
the genus Stigmatocythere due to its characteristic surface ornamentation particularly
the large hexagon formed of ridges just to the posterior of centre.

Stigmatocythere lumaria sp. nov.
(Plate 37, figs. 11 ; Plate 38, figs. 1-10 ; Plate 39, figs. 1-8, 11)
DERIVATION OF NAME. Latin lumarius, thorny.

Diacnosis. A species of Stigmatocythere with a prominent and bilobate subcentral-
tubercle. Surface tuberculate or combination of reticulations and tubercles. Three
large, projecting tubercles in the mid-dorsal region.

HoLotyPe. Io. 4354, a male carapace (PI. 38, figs. 1, 5, 8).

PARATYPES. Io. 4306-7 + Io. 3150-4.

MATERIAL. Approximately 340 specimens from the Rakhi Nala section from 21
horizons (sample nos. 3621, 3624 to 3628, 3630, 3640 to 3642, 3645 to 3652, 3658,
3662 and 3663). 86 specimens from the Zao River section from 11 horizons (sample
NOS. 24156, 24157, 24159, 24170, 24173 to 24176, 24180, 24187 and 24193). GSP BM
2592-94.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Upper Chocolate Clays, sample no. 3642.

DESCRIPTION. Sexual dimorphism rather pronounced ; the females are shorter
and higher than the males. Cavapace subrectangular in lateral view. Anterior
margin broadly rounded, posteroventral margin and posterior extremity more or

76 EARLY TERTIARY OSTRACODA

less rounded, posterodorsal slope very slightly concave. Dorsal margin straight but
appears intricate due to surface ornamentation ; ventral margin slightly concave in
front of the middle, particularly in right valve. Greatest height lies in the anterior
third, greatest length passes above mid-line. Anterior and posterior cardinal angles
well-marked. Valvesalmost equal. Evye-tubercle distinct androunded. Swubcentral-
tubercle prominent and bilobate. Surface ornamentation consists of either tubercles
or a combination of reticulations and tubercles. In some cases tubercles become
almost spinose. There are three tubercles in the mid-dorsal region and in most
specimens these project beyond the dorsal margin in lateral outline. The eye-
tubercle is joined to the subcentral-tubercle, by a sharply curved ridge, diagnostic of
the genus. The ventral side of the subcentral-tubercle has two weak, short, curved
ridges ; one extends towards the anterior and the other towards the posterior end.
Anterior marginal rim well-marked, ventral and posterior marginal rims less well-
marked. Anterior and posterior margins decorated by short, numerous spines.
Posterior has several short spines and two large, somewhat blunt spines, one in the
posteroventral corner and the other in the posterodorsal corner. Radial pore canals,
simple, almost straight, irregularly spaced, some crossing one another, 25-26
anteriorly. Line of concrescence and inner margin coincide. Anterior duplicature
moderately wide, one-twelfth of the entire length of the valve. Selvage pronounced,
in right valve it lies in the outer third of the duplicature but in left valve it is sub-
marginal. Right valve with deep ventral and anterior flange grooves. Adductor
scars in a vertical column of four. Fvontal scar not clearly seen but appears to be
ovalin shape. Hinge holamphidont with the following details :

Element Left valve Right valve

Anterior Socket bounded on all Strongly projecting conical
sides, ocular sinus tooth. Ocular sinus lies
opening into it. below and slightly

anterior to it.

Anteromedian Subconical tooth with Deep socket bounded on
straight anterior and venter and opening into
convex posterior in posteromedian groove.
dorsal view.

Posteromedian Denticulate bar. Locellate groove.

Posterior Deep socket open in Tooth more or less rounded
venter. in lateral view but

pessular in dorsal view.

COMPARISON. Stigmatocythere portentum sp. nov. is larger, than the present
species, has three distinct longitudinal ridges, lacks a tuberculate surface and bilobate
subcentral-tubercle. Stigmatocythere calia sp. nov. is probably ancestral to
S. lumaria but has a vertical posteroventral ridge and more elevated anterior
marginal rim. Further, it lacks a tuberculate surface and bilobate subcentral-
tubercle.

This species can be separated into two morphotypes, although it is rather difficult
to maintain a distinction between them because of many intermediate forms :

FROM WEST PAKISTAN a7,

MORPHOTYPE A
(Plate 37, figs. 11 ; Plate 38, figs. 1-10 ; Plate 39, fig. 11)
This has a tuberculate surface. Tubercles vary in size and number. Some have

few large tubercles with a tendency to become spinose and in others tubercles are
small and rounded.

DIMENSIONS (mm).

L H WwW
Io. 4354 Carapace male (holotype) 0-67 0°37 0°34
Io. 4307. Right valve female 0:63 0°37 —
Io. 4306 Left valve male (juvenile) 0°59 0°32 -—
Io. 3151 Right valve male 0-68 0°37
Io. 3152 Left valve female 0°59 0:40 i
Io. 3154 Right valve female 0-60 0°37 —

MoRPHOTYPE B
(Plate 39, figs. 1-8)
This is very similar to Morphotype A, but has a surface ornamentation which is a
combination of reticulations and tubercles.
DIMENSIONS (mm).

IL, H WwW
Io. 3150 Carapace male 0-66 0-37 0°32
Io. 3153 Carapace female 0-62 0°37 0-31

REMARKS. This species has been described as Genus and sp. indet.G. by I. G.
Sohn in his paper on Lower Tertiary ostracods from Western Pakistan, still in press.

Genus TRACHYLEBERIS Brady 1808

TYPE SPECIES. Cythere scabrocuneata Brady 1880.

Subgenus TRACHYLEBERIS sensu stricto

Trachyleberis (Trachyleberis) lobuculus sp. nov.
(Plate 39, figs. 9, 10 ; Plate 40, figs. 1, 3)
DERIVATION OF NAME. Latin lobus, lobe + oculus, eye ; with reference to the
lobate eye-tubercle.

Diacnosis. A species of the subgenus Tvachyleberis in which eye-tubercle is
lobate, surface ornamented with tubercles, posterior cardinal angle well-marked in
left valve.

HoLotyPe. lo. 4364, a female carapace (PI. 40, figs. 1, 3).
PARATYPE. Io. 4308.

78 EARLY TERTIARY OSTRACODA

MATERIAL. 287 specimens from the locality below from 49 horizons (sample
nos. 3147, 3160, 3162, 3163, 3166, 3167, 3169 to 3171, 3173 to 3175, 3177 to 3180,
3183 to 3193, 3197 to 3200, 3401 to 3404, 3407, 3409, 3410, 3415, 3417 to 3422, 3428,
3429, 3434 and 3435). GSP BM 2595-6.

TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Upper Rakhi Gaj Shales, sample no. 3166.

DESCRIPTION. Carapace subrectangular in the male dimorph and sub-quadrate in
the female. Sexual dimorphy apparent, the males being larger in proportion to the
females. Dorsal and ventral margins subparallel (undulating in lateral view because
of surface ornamentation). Anterior margin broadly rounded, posterodorsal margin
almost straight, posterior extremity somewhat rounded, posterodorsal margin
curved. Greatest height at anterior cardinal angle and greatest length in the
middle. Valves more or less equal. Eye-tubercle lobate and prominent situated just
below a well-developed anterior cardinal angle. Posterior cardinal angle well-
marked in the left valve, armed with a node or short spine pointing upwards. Swub-
central-tubercle distinct. Both anterior and posterior margins ornamented with a
double row of tubercles or very short spines. Surface tuberculate or nodose (oc-
casionally tubercles or nodes develop into spines). Anterior and posterior marginal
rim more or less distinct. Hinge holamphidont : left valve with terminal sockets,
postjacent conical tooth and median denticulate bar ; right valve hinge compliment-
ary (anterior tooth being conical). Duplicature of moderate width with a submarginal
selvage. Other internal details not determinable.

DIMENSIONS (mm).

L H W
Io. 4308 Carapace male 0-61 0°34 0:24
Io. 4364 Carapace female (holotype) 0°59 0°35 0:28

ComMPARISON. T. lobuculus is probably related to Cythereis spinellosa Lubimova
and Guha (1960) but differs in having a lobate eye-tubercle and a different lateral
outline and surface ornamentation.

REMARKS. Specimens of Cythereis spinellosa Lubimova and Guha were not
available for comparison, but from the description and figure given by these authors
it appears that the eye-tubercle in that particular species is not lobate.

Trachyleberis (Trachyleberis) bimammillata sp. nov.
(Plate 40, figs. 2, 4-11)
DERIVATION OF NAME. Latin bimammiullata, two-breasted ; with reference to the
split subcentral-tubercle.

Diacnosis. A small species of the subgenus Tvachyleberis in which subcentral-
tubercle is divided into two horizontally disposed nodes and posterodorsal process
consisting of two vertically arranged nodes.

Ho.otyPe. Io. 4363, a male carapace (PI. 40, figs. 2, 8, 10).
PARATYPES. Io. 3155-9.

FROM WEST PAKISTAN 79

MATERIAL. 42 specimens from the Rakhi Nala section from 5 horizons (sample
nos. 3610, 3613 to 3615 and 3617). 7 specimens from the Zao River section from
two horizons (sample nos. 24150 and 24152). GSP BM 2597.

TYPE LOCALITY. Rakhi Nala section.
TypEe Horizon. Upper Chocolate Clays, sample no. 3613.

DESCRIPTION. Carapace subrectangular to subquadrate in lateral outline.
Sexual dimorphism moderate ; the presumed males are longer and less high than the
females. Dorsal and ventral margins straight and tapering towards the posterior.
Anterior margin broadly rounded, posterodorsal slope very slightly concave, posterior
extremity rounded, posterodorsal margin somewhat rounded. Anterior, posterior
and ventral margins decorated with a double row of short spines, but dorsal margin
with only one row of very short spines (these in some specimens almost look like
pustules). Greatest height at the anterior cardinal angle (which is obtuse and
angular) and greatest length through the mid-point. In dorsal view the greatest
width lies at the anterior node of the subcentral-tubercle. Valves almost equal.
Eye-tubercle rounded and distinct. Swubcentral-tubercle divided into two nodes,
horizontally arranged, the anterior one being larger (spinose in some specimens).
The posterodorsal process consists of two nodes (spines in some specimens), which
are vertically disposed. In a few specimens a posteroventral node is also present.
Surface ornamented with scattered tubercles and spines. Duplicature fairly wide.
The selvage is subperipheral and well-developed in both valves. Radial pore canals
not seen because of mineralization. The adductor muscle scars are in an oblique row
of four at the posterior margin of the subcentral pit. The frontal scar is large and
U-shaped and opens towards the anterodorsal corner. Hinge holamphidont with
the following details :—

Element Left valve Right valve
Anterior Socket. Projecting subconical tooth.
Anteromedian Subconical tooth. Socket.
Posteromedian Denticulate ridge. Locellate groove.
Posterior Fairly deep socket. Tooth, subpessular in dorsal
view.
DIMENSIONS (mm).
L H W
Io. 4363 Carapace male (holotype) 0°52 0:29 0:22
Io. 3159 Carapace female 0-50 0:29 0°24
To. 3158 Left valve male (broken) -= 0°29 —
To. 3156 Left valve female 0°50 0-29 —
Io. 3155 Right valve female 0°49 0-29 —
Io. 3160 Carapace male 0°54 0-29 0:22
Io. 3157 Carapace female 0-49 0°29 0:24

CoMPARISON. This species can easily be distinguished from Tvachyleberis (Trachy-
leberis) lobuculus sp. nov. by its smaller size, slightly concave posterodorsal margin
and split subcentral-tubercle. Further, Tvachyleberis (Tvachyleberis) bimammuillata

80 EARLY TERTIARY OSTRACODA

has a posterodorsal process consisting of two nodes in a vertical row and lacks a
lobate eye-tubercle.

Subgenus ACANTHOCYTHEREIS Howe 1963

TYPE SPECIES. Acanthocythereis araneosa Howe 1963.

Trachyleberis (Acanthocythereis) procapsus sp. nov.
(Plate 40, figs. 12, 13 ; Plate 41, figs. 1, 3, 4)

DERIVATION OF NAME. Latin frocapsus, anterior cage ; with reference to the
smooth walled area enclosed behind the anterior marginal rim.

Diacnosis. Acanthocytheris in which a smooth walled area lies behind the anterior
marginal rim, anterior and posterior platforms compressed.

HototyPe. Io. 4360, a male carapace (PI. 40, fig. 12 ; Pl. 41, figs. 1, 3).

PARATYPE. Io. 3164.

MATERIAL. Six specimens from the locality below from two horizons (sample
nos. 460-j and 460-1). GSP BM 2508.

TYPE LOCALITY. Sor Range section.

TYPE HORIZON. Upper Palaeocene, sample no. 460-}.

DESCRIPTION. Sexual dimorphism apparent ; the males are longer in proportion
to the females. Carapace elongate, subrectangular in lateral outline with dorsal and
ventral margins almost straight, tapering towards the posterior. Anterior margin
broadly rounded, posterior subtriangular. Anterior cardinal angle rounded. Left
valve over-reaches the right very slightly at the posterodorsal margin. Greatest
height through the anterior cardinal angle and greatest length through the mid-
point. In dorsal view the greatest width is situated at the anterior third. Swub-
central-tubercle distinct. Evye-tubercle rounded and distinct and lies at the anterior
cardinal angle. Surface reticulate (reticulae joined by walls or pustules or papillae).
Posterodorsal process consists of small more or less rounded protuberances. Anterior
and posterior margins ornamented by a double row of short spines. Anterior and
posterior marginal rims high with a smooth walled area behind. Internal details not
known.

DIMENSIONS (mm).

L H W
Io. 4360 Carapace male (holotype) 0-68 0°32 0°20
Io. 3164 Carapace female 0°59 0:30 0:20

CoMPARISON. This species is distinguishable from Tvachyleberis (Acanthocytherets)
usitata sp. nov. by its deeper reticulation, more elevated marginal rims and spinose
anterior and posterior margins.

The present species has already been compared with Trachyleberis (Acanthocythereis)
postcornis sp. nov. and Trachyleberis (Acanthocytherets) decoris sp. nov.

FROM WEST PAKISTAN 81

Remarks. T. (A.) procapsus has so far only been recovered from the Upper
Palaeocene of the Sor Range section.

Trachyleberis (Acanthocythereis) usitata sp. nov.
(Plate 41, figs. 2, 5, 7)
DERIVATION OF NAME. Latin usitatus, usual.

Diacnosis. Carapace tapering towards posterior. Subcentral-tubercle distinct.
Surface reticulate with superimposed pustules and a posterodorsal process.

Hototyre. lo. 4362, a male carapace (PI. 41, fig. 2).
PARATYPE. lo. 3161.

MATERIAL. Five specimens from the Rakhi Nala section from four horizons
(sample nos. 3111, 3130, 3132 and 3133). GSP BM 2599.

TYPE LOCALITY. Rakhi Nala section.
TYPE HORIZON. Gorge Beds, sample no. 31IT.

DESCRIPTION. Sexual dimorphy present ; the males are longer in proportion to
the females. Carapace elongate, subrectangular, tapering towards the posterior.
Anterior margin broadly rounded, posterior narrowly rounded. Dorsal and ventral
margins almost straight. Greatest height at the anterior cardinal angle which is
well-developed in the left valve. Greatest length passes through mid-point. Valves
almost equal. In dorsal view greatest width lies in the anterior third (in the region
of the subcentral-tubercle). Eye-twbercle rounded and distinct. Swubcentral-tubercle
fairly distinct. Anterior and posterior marginal rims sharply defined. Surface
reticulate with pustules at reticulae intersections. A posterodorsal process in the
form of more or less rounded tubercle of medium size present. A double row of
pustules decorates anterior and posterior margins. Internal characters not known.

DIMENSIONS (mm).

L H WwW
To. 4362 Carapace male (holotype) 0-63 0°32 —
Io. 3161 Carapace female 0:59 0°32 0°22

COMPARISON. This species shows some resemblance to Trachyleberis (Acantho-
cythereis) decoris sp. nov. but is smaller, has marginal pustules rather than spines and
carapace more tapering towards the posterior. Tvachyleberis (Acanthocytherets)
postcornis has a characteristic posterodorsal process, more high marginal rims and
spinose anterior and posterior margins.

REMARKS. In some specimens the posterodorsal process is not well-developed.

Trachyleberis (Acanthocythereis) pedigaster sp. nov.
(Plate 41, figs. 6, 8)

DERIVATION OF NAME. Greek pedigaster, flat belly ; with reference to the ventral
inflation.

Dracnosis. A large species of the subgenus with ventral inflation. Carapace
tapering towards posterior. Posterior margin subtriangular.

F

82 EARLY TERTIARY OSTRACODA

Ho.otype. Io. 4358, a carapace.

MATERIAL. Only one specimen from the locality and horizon below.
TYPE LOCALITY. Rakhi Nala section.

TYPE HORIZON. Lower Rakhi Gaj Shales, sample no. 3671.

DESCRIPTION. Cavapace large, elongate, tapering towards posterior and with
ventral inflation. Anterior and posterior marginal platforms compressed. Dorsal
and ventral margins almost straight, anterior margin broadly rounded, posterior
subtriangular. Anterior and posterior cardinal angles well-developed particularly
in the left valve. Left valve slightly larger than the right, which it over-reaches in
the region of the anterodorsal corner and posterodorsal slope. Evye-tubercle rounded
and distinct and lies just below the anterior cardinal angle. Swubcentral-tubercle more
or less distinct. Greatest height through the anterior cardinal angle, greatest length
below mid-point and greatest width a little posterior to the middle. Surface
ornamentation consists of reticulations with superimposed papillae. The postero-
dorsal process consists of two almost rounded small tubercles (or papillae) joined
together in the left valve and one small rounded tubercle in the right. Anterior and
posterior marginal rims fairly distinct. Anterior and posterior margins ornamented
with a double row of papillae. Internal details unknown.

DIMENSIONS (mm).

L H WwW
Io. 4358 Carapace (holotype) I-02 0-51 -

CoMPARISON. There is no difficulty in separating T. (A.) pedigaster sp. nov. from
other described species of the subgenus Acanthocythereis by its large carapace and
subtriangular posterior.

Trachyleberis (Acanthocythereis) postcornis sp. nov.
(Plate.4z, figs. 9,10; Pl.-42; figs. 2, 2;'7,.10)
DERIVATION OF NAME. Latin post, posterior mie cornis, horned ; with reference to
the posterodorsal process.

Diacnosis. A species of the subgenus Acanthocythereis with distinct subcentral-
tubercle and eye-tubercle. Surface reticulate with small superimposed spines.
Posterodorsal process divided into two spines.

HototyPe. Io. 4361, a male carapace (PI. 42, figs. I, 2, 7, IO).
PARATYPES. Io. 4309-+3162—4.

MATERIAL. 45 specimens from the locality below from two horizons (sample
nos. 3498 and 3499) and 3 specimens from the Zao River section from two horizons
(sample nos. 24131 and 24148). GSP BM 2600.

TYPE LOCALITY. Rakhi Nala section.
Type Horizon. Lower Chocolate Clays, sample no. 3499.

DESCRIPTION. Carapace elongate, subrectangular in lateral outline with dorsal
and ventral margins straight, tapering towards the posterior. Anterior margin

FROM WEST PAKISTAN 83

broadly and evenly rounded, posterior slightly sub-triangular in right valve but
almost rounded in the left, posterodorsal margin very slightly concave. Anterior
and posterior cardinal angles well-developed. Sexual dimorphism rather strong ;
the presumed males are longer, less high and less wide than the females. Valves
almost equal. In dorsal view greatest width lies at the anterior third (in the region
of the subcentral-tubercle). Eye-tubercle rounded, polished and prominent (standing
out from the carapace). Swubcentral-tubercle distinct. Surface ornamentation consists
of combination of reticulations and small spines. The posterodorsal process is
divided into two spines (although in some specimens this division is not detectable).
In a few specimens a posteromedian process is also developed. The anterior and
posterior margins are decorated with a double row of spines ; the second row lies on
high anterior and posterior marginal rims. The posterior marginal spines are
larger and less in number. Duplicature fairly wide. Selvage prominent and sub-
marginal. Radial pore canals not clearly displayed because of mineralization, but
appear to be simple, more or less straight with median swellings, 30-35 anteriorly.

Hinge holamphidont :
Element Left valve Right valve
Anterior Socket. Stirpate tooth.
Anteromedian Subconical tooth. Deep socket.
Posteromedian Locellate shallow groove. Denticulate bar.
Posterior Deep socket. Tooth, almost rounded in

lateral view.

DIMENSIONS (mm).

L H W
Io. 4361 Carapace male (holotype) 0-62 0-30 0:22
To. 3162 Carapace female 0°52 0:29 0-21
To. 3164 Right valve male 0-61 0:29 —
To. 3163 Right valve female 0°50 0-28 —

COMPARISON. The present species shows some affinity to Tvachyleberis (Acantho-
cytherets) decoris sp. nov. but is shorter, less high and less wide. These two species
also differ in surface ornamentation. T. (A.) postcornis has a combination of
reticulations and small spines, while T. (A.) decoris is reticulate with superimposed
pustules. Further, JT. (A.) postcornis has a well-developed posterodorsal process
divided into two spines and a distinct subcentral-tubercle. This species may also be
distinguished from T. (A.) procapsus sp. nov. in being smaller and lacking a smooth
walled area behind the anterior marginal rim. These two species also differ in dorsal
outline.

Trachyleberis (Acanthocythereis) decoris sp. nov.
(Plate 42, figs. 3-6, 8, 9)

DERIVATION OF NAME. Latin decoris, beautiful, adorned ; with reference to the
bejewelled appearance of the pustules and reticulae.

Diacnosis. Acanthocythereis in which surface ornamentation consists of reticula-

F*

84 EARLY TERTIARY OSTRACODA

tions with superimposed pustules. Carapace subrectangular with dorsal and ventral
margins almost straight and subparallel.

Ho.otyPe. Io. 4359, a male carapace (PI. 42, figs. 3, 4, 5).
PARATYPE. Io. 4310.

MATERIAL. Over 250 specimens from the type locality from 18 horizons (sample
nos. 3604, 3607, 3610, 3613 to 3615, 3629, 3640, 3642, 3645, 3648 to 3650, 3661 to
3664). 7 specimens from the Zao River section from 3 horizons (sample nos. 24154,
24173 and 24193). GSP BM 2601-2.

TYPE SECTION. Rakhi Nala section.
TYPE HORIZON. Upper Chocolate Clays, sample no. 3640.

DESCRIPTION. Sexual dimorphism rather marked ; the males are longer in pro-
portion than the females. Cavapace subrectangular in lateral view with dorsal and
ventral margins almost straight and subparallel. Anterior margin broadly and evenly
rounded, posterior slightly subtriangular. Both anterior and posterior margins
ornamented by a double row of short spines, the posterior ones being larger. Anterior
cardinal angle rounded, posterior cardinal angle well-marked. Greatest height at the
anterior cardinal angle and greatest length in the middle. In dorsal view greatest
width lies in the anterior third. Valvesalmost equal. Surface reticulate with super-
imposed pustules. Evye-tubercle rounded and distinct. Swubcentral-tubercle more or
less distinct (better seen in slightly worn specimens). A large number of specimens
(particularly the females) show development of posteroventral prominence. Ina few
specimens a small posterodorsal process also develops, but these characters here are
regarded as variations within the species. Dwplicature fairly wide with a submarginal
seluvage. Radial pore canals not seen. Muscle scary pattern consists of four adductors
in a vertical superposition at the posterior margin of the muscle scar pit with a
U-shaped frontal scar opening towards the anterodorsal corner. Hinge holamphidont :

Element Left valve Right valve

Anterior Socket. Slightly stirpate tooth.
Anteromedian Subconical tooth. Denticulate ridge.
Posterior Deep socket. Somewhat rounded tooth

in lateral view.

DIMENSIONS (mm).

L H W
Io. 4359 Carapace male (holotype) 0:67 0:32 0:24
Io. 4310 Carapace female 0°59 0:32 0:23

CoMPARISON. In some respects this species resembles Tvachyleberis (Acantho-
cythereis) procapsus sp. nov., but differs in lacking the smooth walled area enclosed
by the anterior marginal rim and compressed anterior and posterior marginal
platforms. These two species also differ in size, T. (A.) decoris being smaller, higher
and wider in proportion than T. (A.) procapsus.

REMARKS. This species commonly occurs in the Upper Chocolate Clays of the
Rakhi Nala section but it is very rare in the Zao River section.

FROM WEST PAKISTAN 85

V. OSTRACODA AND EARLY TERTIARY CORRELATION
IN THE SULAIMAN RANGE
(a) BIOSTRATIGRAPHIC UNITS

Throughout the succession the ostracods have revealed a fairly shallow-water
marine environment. Although Eames (1952a) has recorded small freshwater
gastropods in the lower part of the Lower Chocolate Clays (in his local zones 8 & 9) of
the Rakhi Nala section, he believes they were carried down from a closely neigh-
bouring source and deposited under estuarine conditions. No freshwater ostracods
have been found, however, and the presence of Neocyprideis sp. in the Shales with
Alabaster could represent either estuarine or super saline conditions.

Except for a few gaps, ostracods occur throughout almost the whole succession.
At many horizons, particularly in the Eocene, samples are completely crowded with
ostracods. The diversity of the fauna being suggestive of ideal conditions. They
usually occur in association with larger and smaller benthonic Foraminifera, but
are very rare or almost absent in samples with rich pelagic Foraminifera. The most
conspicuous gap in the Eocene succession of the Rakhi Nala and Zao River sections
which has not yielded any ostracods is the Platy Limestone and the lower part of the
Lower Chocolate Clays. In the Zao River section the top 600 ft. of the Upper
Chocolate Clays are devoid of any recognizable ostracods, although at a few horizons
some Nummulites have been found.

RAKHI NALA SECTION

The following ostracod biostratigraphic units in the Rakhi Nala section have been
recognized (see Table 4). Each unit is identified by a distinct ostracod fauna, a
change of faunal suite marking the base.

Ostracod Biostratigraphic Unit I, Palaeocene (lower part)

The first Tertiary ostracod assemblage is encountered in the lower part of the
Gorge Beds (samples from the Venericardia Shales were not available for study).
Seven out of the eight species recorded are restricted to the unit. The species which
ranges up into Unit II is Tvachyleberis (Acanthocytherets) usitata sp. nov. Alocopo-
cythere rupina sp. nov., Neocyprideis ? sp.A and Bairdia sp.A are abundant and make
up over 80% of the ostracod fauna.

Ostracod Biostratigraphic Umit II, Palaeocene (upper part)

This Unit contains the Lower Rakhi Gaj Shales. Ostracods are very rare and
have only been found in the upper part which is very rich in pelagic Foraminifera.
The ostracods, although very rare, are easily distinguishable from the assemblages
below and above. Of the eight species found, all are restricted to the present Unit,
with the exception of Tvachyleberis (Acanthocythereis) usitata sp. nov. which is also
present in the underlying Unit.

Ostracod Biostratigraphic Unit III, Lower Eocene (lower part)
This includes the Upper Rakhi Gaj Shales, Green and Nodular Shales and Rubbly
Limestones. Eames’ local zones 3, 4, 5 and 6 lie in this Unit.

86 EARLY TERTIARY OSTRACODA

This is the first Eocene ostracod assemblage. It is fairly rich and at several
horizons the ostracods are very abundant. None of the Palaeocene species survive
and a completely new fauna evolves. The ostracod fauna is of changing suite ;
species appear and disappear in the unit, but there seems to be no major break of
any kind in the fauna. Tvachyleberis ([rachyleberis) lobuculus sp. nov., Gyrocythere
parvicarinata sp. nov., Occultocythereis peristicta sp. nov. (with five morphotypes),
Schizocythere sp.A and Pontocythere sp.A are the most important members restricted
to the unit. Approximately 50% of the species range up into the overlying Unit IV.

Ostracod Biostratigraphic Unit IV, Lower Eocene (upper part)

This consists of the Shales with Alabaster and includes Eames’ local zone 7. It has
a very rich ostracod faunal assemblage. Most of the samples studied were extremely
rich in ostracods, which are mostly complete carapaces. The most typical species
confined to the unit are Stigmatocythere obliqua sp. nov., Phalcocythere dissenta sp. nov.
Genus C sp. rand Genus C sp. 2. More than 50% of the species are restricted to the
unit, although approximately 44° are common to Unit III, and only one species
ranges up into Unit V.

Ostracod Biostratigraphic Unit V, Middle—Upper Eocene

This comprises the Platy Limestone, Lower Chocolate Clays, Upper Chocolate
Clays and Pellatispira Beds. Eames’ local zones 8 to 15 and Latif’s top six pelagic
foraminiferal zones occur in this unit. The lowest 730 ft., which form the Platy
Limestone and most of the Lower Chocolate Clays, excluding the top 30 ft., are
devoid of any recognizable ostracods and are provisionally included in the unit.
There are 200 ft. of covered sediments in the Lower Chocolate Clays below sample
3494.

The Unit is very rich in very well-preserved ostracods. It differs markedly from
the underlying Unit. All the species except Alocopocythere transcendens sp. nov.,
which survives from the Unit below, appear for the first time, although a few have their
ancestors in the Unit IV. The first appearance of ostracods in the Unit is in the
uppermost part of the Lower Chocolate Clays (sample nos. 3498 and 3499), which lies
at the base of the Globigerina yeguaensis zone of Latif. The ostracod fauna is varied
and of changing suite. 12% of the species are restricted to the Lower Chocolate
Clays (topmost portion) ; 25°% are confined to the Upper Chocolate Clays (lower
part) ; and only 8% have been recorded from the Pellatispira Beds. 33% of the
species are shared between the Lower Chocolate Clays (topmost portion) and Upper
Chocolate Clays (lower part) ; 17% range from the Lower Chocolate Clays (upper-
most part) to the Upper Chocolate Clays (upper part) ; 37% are shared between the
Upper Chocolate Clays (lower part) and Upper Chocolate Clays (upper part) ; 8%
range from the Upper Chocolate Clays (lower part) to the Pellatispiva Beds ; and 12%
are found in both the Upper Chocolate Clays (upper part) and Pellatispiva Beds.
(The percentages are approximate and are based on the entire ostracod fauna of the
Unit).

The genus Alocopocythere nov. occurs abundantly almost throughout the Unit.

FROM WEST PAKISTAN 87

Alocopocythere transcendens sp. nov., which ranges up from the underlying Units ITI
and IV, is replaced by A. tvansversa sp. nov. just above the middle of the lower part
of the Upper Chocolate Clays. This last species has several morphotypes ; in the
upper part of the Upper Chocolate Clays the papillose form becomes more common
and in the Pellatispiva Beds this is the only morphotype present. Stigmatocythere
obliqua sp. nov., which was very abundant in the underlying biostratigraphic Unit
IV, is replaced by the larger Stigmatocythere portentum sp. nov., which has only been
found in the uppermost part of the Lower Chocolate Clays. Higher up in the
succession, i.e. in the lower part of the Upper Chocolate Clays the place of S. porten-
tum is taken by Stigmatocythere lumaria sp.nov. which ranges up into the Pellatispira
Beds. The genera Cytherella, Cytherelloidea, Krithe, and Patjenborchella are repre-
sented by several species. The genus Gyrocythere nov. has two species in the Unit.
Gyrocythere perfecta sp. nov. occurs in the uppermost part of the Lower Chocolate
Clays but in the lower part of the Upper Chocolate Clays it is replaced by the larger
Gyrocythere exaggerata sp. nov. The subgenera Scelidocythereis nov. and Paracosta
nov. are represented by two and three species respectively. Paracosta is known so far
only from this Unit. The following are some of the most important species of the
Unit : Bairdoppilata sp.A, Cytherelloidea cf. C. costatruncata Lubimova and Mohan,
Cytheromorpha sp.A, Cytheropteron sp.D, Alocopocythere transversa sp. nov. (with six
morphotypes), Patagonacythere ? nidulus sp. nov., Stigmatocythere lumaria sp. nov.
(with two morphotypes) and Tvachyleberis (Acanthocytherets) decoris sp. nov.

ZAO RIVER SECTION

The two biostratigraphic Units IV and V of the Rakhi Nala section are found in
the Zao River section (see Table 5).

Ostracod Biostratigraphic Unit IV, Lower Eocene (upper part)

This is very similar to biostratigraphic Unit IV of the Rakhi Nala section. The
base of the Unit has been taken arbitrarily at the base of the four foot limestone,
which lies 332 ft. below the base of the Platy Limestone. The actual base of the Unit,
or the Shales with Alabaster, probably lies much lower in the succession, but sediments
below the 4 ft. limestone have not been analysed. These have been recorded as the
undifferentiated Ghazij by the collectors. No megafossils have so far been recorded
from these sediments and it is unlikely that these would yield any smaller foramini-
fera or ostracods because of their lithology—mostly silty shales.

Ostracods have been found in the upper part of the Unit at two horizons (samples
24107 and 24110). They are extremely abundant in 24107. Approximately half
of the Rakhi Nala species of the Unit are found in the Zao River section. None of
the species range up into Unit V. Stigmatocythere obliqua sp. nov. is the most
dominant species and makes about one-third of the ostracod fauna. Neocypridets ?
sp.B, Neocyprideis sp.C, Pontocyprella sp.B, Pontocyprella sp.C, Xestoleberis sp.C,
Xestoleberis sp.D, Xestoleberis sp.E, Genus C sp.1 and Genus C. sp.2 are some other
important species,

88 EARLY TERTIARY OSTRACODA

Ostracod Biostratigraphic Unit V, Middle-Upper Eocene

The biostratigraphic Unit V of the Rakhi Nala section occurs in the Zao River and
is 3743 ft. thick. The Unit includes the Platy Limestone, Lower Chocolate Clays
and both lower and upper parts of the Upper Chocolate Clays. The Pellatispira
Beds have not been recorded in the Zao River section. The bottom 460 ft. of the
Unit comprising the Platy Limestone and the lower part of the Lower Chococlate
Clays and the top 600 ft. of the upper part of the Upper Chocolate Clays have not
yielded any ostracods and are only provisionally included in the Unit.

The Unit is very rich in well-preserved ostracods. The ostracod fauna is com-
pletely new since none of the species from the Unit below survive. There are 7%
of the species which are restricted to the upper part of the Lower Chocolate Clays ;
26% have only been found in the lower part of the Upper Chocolate Clays ; and 17%
have been recorded from the upper part of the Upper Chocolate Clays only. 24% of
the species occur in the upper part of the Lower Chocolate Clays and the lower part
of the Upper Chocolate Clays ; 13% range from the upper part of the Lower Choco-
late Clays to the upper part of the Upper Chocolate Clays and 41% are shared be-
tween the lower and upper parts of the Upper Chocolate Clays.

The ostracod fauna of the Unit in the Zao River section is very similar to that of
the Rakhi Nala section. It has about 74% of its species in common with the Rakhi
Nala. These are shown in Appendix 2. As in the Rakhi Nala, Alocopocythere is
one of the commonest genera and it occurs in great abundance at several horizons.
It is represented by three related species ; A. tvanscendens sp. nov., A. transversa
sp. nov. (with six morphotypes), and A. radiata sp. nov. Stigamatocythere is another
common genus and has three species in the Unit : S. calia sp. nov., S. delineata
sp. nov. and S. lumarvia sp. nov. (with two morphotypes). Among these, S. lwmaria
is the commonest, occurring in the lower and upper parts of the Upper Chocolate
Clays. S.caliaand S. delineata have so far not been found in the Rakhi Nala section.
Bairdoppilata sp.A, Pterygocythereis (Pterygocythere) sp.A are more common in the Zao
River section. Tvachyleberis (Acanthocythereis) decoris sp. nov. and Cytheromorpha
sp.A, which were very common in the Rakhi Nala are rare in the Zao River. The
subgenus Paracosta nov. which is represented by three species in the Rakhi Nala has
not so far been recorded from the Zoa River. Phalcocythere spinosa sp. nov. and
Quadracythere (Hornibrookella) subquadra sp. nov. have only been found in sample
24161 of the Zao River section, where they occur in association with the larger
foraminifera Pellatispiva orbitoidea. Some of the more important members of the
unit are : Alocopocythere transcendens sp. nov., Alocopocythere transversa sp. NOov.,
Bairdoppilata sp.A, Cytherelloidea cf. C. costatruncata Lubimova and Mohan,
Patagonacythere ? nidulus, sp. nov. and Stimatocythere lumaria sp. nov.

SHPALAI KHWARA SECTION

The ostracod biostratigraphic Unit IV of the Rakhi Nala and Zao River sections
is also represented in the Shpalai Khwara section.

FROM WEST PAKISTAN 89

Ostracod Biostratigraphic Unit IV, Lower Eocene (upper part)

Like the Zao River section, the base of the Unit is taken arbitrarily at the base of
the 4 ft. limestone, which is 320 ft. below the base of the Platy Limestone. The
Platy Limestone is only 40 ft. thick in this section. The 12,450 ft. thick sediments
below the 4 ft. limestone are barren except for a few horizons which contain poorly
preserved pelagic Foraminifera. These probably represent the following lithological
units in ascending order : Upper Rakhi Gaj Shales, Green and Nodular Shales,
Rubbly Limestones and lower part of the Shales with Alabaster.

Only the upper part of the Unit has yielded ostracods and the fauna is similar to
that of the Zao River and Rakhi Nala sections. About 70% of its species are in
common with the Zao River section. Over 40% of the species of the Unit in the
Rakhi Nala have been recorded from the Shpalai Khwara section. Stigmatocythere
obliqua sp. nov. is very abundant, particularly in sample 24686, which is absolutely
crowded with this species. Some other common species are : Neocyprideis ? sp.B,
Pontocyprella sp.B, Cytherella sp.B, Cytherella sp. and Genus C sp.2.

(b) STATISTICAL CORRELATION OF RANGES OF OSTRACOD SPECIES COMMON TO THE
RAKHI NALA AND ZAO RIVER SECTIONS.

The tops and bases of ostracod species common to the Rakhi Nala and Zao River
sections have been plotted on a graph (Fig. 6). These fall into two rectilinear
patterns, one in biostratigraphic Unit IV and the other in biostratigraphic Unit V.

The tops and bases of ostracod species in biostratigraphic Unit IV (i.e in the
Shales with Alabaster) lie almost in a straight line on the graph ; this, however, is
because ostracods have only been found at two horizons in the Zao River section.
The Equations of Correlation for the array of biostratigraphic Unit V (i.e. above the
Platy Limestone) can be computed from the data given in Appendix 2. This
method has been discussed in detail by A. B. Shaw in his book ‘Time in Stratigraphy’,
published in 1964. The points marked ‘+’ in Appendix 2 have been omitted because
they fall outside the main array. Eighty-one points have been considered. The
Equations of Correlation between the Rakhi Nala and Zao River sections can be
calculated as follows :

A = RN—RN 7 2
RN = RN + eee NS (ZR—ZR), where RN = Rakhi Nala and
x(ZR—ZR)?
ZR = Zao River.
21,675228
48,697965

= 0°4451 ZR + 5635-2 (I)

= 6638 + (ZR—2253)

and
x(RN—RN) (ZR—ZR)

(RN—RN)
=(RN—RN)?

ATR 4

90 EARLY TERTIARY OSTRACODA

21,675228
TI,153941
= 1:9433 RN—10646 «5 (2)

= 2283 + (RN—6638)

Any point in the Zao River section in biostratigraphic Unit V can be correlated
with the corresponding point in the Rakhi Nala section by means of Equation (I).
Similarly any point in the Rakhi Nala section can be correlated with the correspond-
ing point in the Zao River section by using Equation (2).

The Coefficient of Correlation is expressed by the formula :

t = /b, x by
By substituting the values b, and b,, we get
5 = Jouase x £9433
= ,/0°8649
==07959
This high value of r is above the 99% confidence level.

The standard error of estimate for RN (Sey)

me BUSSE
¥ N

where RN—RN is the difference between each observed point and its computed
equivalent and N is the number of entries.

Hence, Sky = ; ee
/18324'554
135°4 ft.
The standard error of estimate for ZR (Sx)
= | (ZR—ZR)?
N

__ /(6464244-61
ol 8x

= /79805°489

= 282°5 ft.

When the two straight lines given by Equations (1) and (2) are drawn on a graph,
they intersect one another at an angle of 3°. Since the Rakhi Nala section has been
regarded as the standard section, therefore, for practical purposes only one straight

gI

aph. This is the
iver with parallel

divided into five
these Units, I and
ycene. Biostrati-
Zao River section
‘tions have almost

ikhi Nala is also

n the Rakhi Nala
‘pecies common to
tion can easily be
vice versa. (see
§ also been calcu-
in the Zao River
rrelated with any

hi Nala and Zinda
River and Shpalai
es with Alabaster
ys (upper Eocene).
e Shpalai Khwara
poorly preserved
bly equivalent to
ly Limestones and
1 sediments below
and are probably
in the northern
able for abundant

cide with Eames’
shales, Green and
yd fauna and are
>is in the Kirthar
y. This occurs in
w the White Marl
2 Lower Chocolate
White Marl Band

section has been
nes and Nagappa.

ZAO RIVER

—L.EOCENE—>|.____________—-MIDDLE EOCENE
Lee

S.WA: PL: ae
S
is}

a

7G Slee
| | 4009
! I
| |
|
| {3500
2
z
if
FS
Ss
oO
(Ss)
=)
WW
& | 3000 R.Lst
1S)
gS SwaA
a PL
a LCC
D>
ucc
WMB
PB.
2500 Data

<=

2000

U.C.C. Lower Parr

1500|-

Graphic representation of range
the Rakhi Nala and Zao River sections.

s of Ostracod species common to
Regression and standard

error of Rakhi Nala on Zao River indicated by lines.

INDEX

+ Top of range

e Base of range

@ Coincident base and top

Opoint not used
G.&NS. Green and nodular shales

Rubbly limestones
Shales with alabaster
Platy limestone
Lower chocolate clays
Upper chocolate clays
White marl band
Pellatispira beds

from Appendix 2.

te 3814
fe 3546, 44 13 5345 gcl3*5 52 48 "36
a” 50 a5!

a

46
5

LOWER EOCENE

36 49

2

7 Ey) 7
i
h /
V ms) ee /
U
40 /
4 20+ +e
1 /
/q By 7
2 A wes 385-27
j ae
/ /
/ fo26 /
/
Y i}
f +2 /
} /
7 /
/+3¢
fi oy)
n=) 7 15>
Y 1 *0%
/ /
/ /
i J
/ ff
if 8 Sea
/ /
/
/ /
/ /
/ /
/ /
i /
19 he aa /
° + /2
/ j=
1 /
/
1 Nigofqu2 / a
#22 “2 He
18 / 844!
Q fo yt
aie iey/2 #
7 216 +n 4
as js
17 a f Z
tn or /
47 393 32 n
oo RN = 0-4451ZR + 5635-2
/ /
U /
ts # 09 /
/ i
/
/
/
/ /
/
i :
/ i
/ /
/ d
i /
/ /
7 /
¢ hs 20 |
/ /
2A3 10 23
2h 0”
/ 38, i 08
/
/
/
39/—20
/
/

Feet above the base of the Venericardia Shales.

6000 WI 6500 7000 Pp
LEG Ks |. C.C. LOWER PART UCC. uprer rt

MIDDLE EOCENE —————>}=-UPPER EOCENE——>|

7500

FROM WEST PAKISTAN gI

line (Le. RN = 0:4451ZR + 5635-2) has been drawn on the graph. This is the
Correlation or Regression Line of the Rakhi Nala on the Zao River with parallel
dotted lines showing the standard error of estimate.

(c) CONCLUSIONS

The Palaeocene and Eocene of the Rakhi Nala section can be divided into five
distinct biostratigraphic units on the basis of Ostracoda. Two of these Units, I and
II, occur in the Palaeocene and three, III, IV and V, in the Eocene. Biostrati-
graphic Unit IV of the Rakhi Nala section is represented in the Zao River section
by at least 332 ft., and Unit V by 3743 ft. The Units in the two sections have almost
identical ostracod faunas. Biostratigraphic Unit IV of the Rakhi Nala is also
represented in the Shpalai Khwara section by at least 320 ft.

The Equations of Correlation of biostratigraphic Unit V between the Rakhi Nala
and Zao River sections have been calculated by means of ostracod species common to
the two sections. From these two equations any point in one section can easily be
correlated with the corresponding point in the other section or vice versa. (see
Fig. 6). The standard error of estimate for the two equations has also been calcu-
lated. Since only the upper part of biostratigraphic Unit IV in the Zao River
section has yielded ostracods, only this part of the unit can be correlated with any
certainty.

Eames’ lithological units for the southern Sulaiman Range (Rakhi Nala and Zinda
Pir) extend into the northern part of the Sulaiman Range (Zao River and Shpalai
Khwara). This is particularly true for sediments from the Shales with Alabaster
(upper Lower Eocene) to the upper part of the Upper Chocolate Clays (upper Eocene).
Sediments below the upper part of the Shales with Alabaster of the Shpalai Khwara
section are unfossiliferous except for a few horizons containing poorly preserved
pelagic foraminifera. These are 12,450 ft. thick and are probably equivalent to
Eames’ Upper Rakhi Gaj Shales, Green and Nodular Shales, Rubbly Limestones and
lower part of the Shales with Alabaster. In the Zao River section sediments below
the upper part of the Shales with Alabaster are undifferentiated and are probably
unfossiliferous. This suggests that environmental conditions in the northern
Sulaiman Range during most of the Early Eocene were not suitable for abundant
marine life.

The faunal breaks in the sections studied do not always coincide with Eames’
lithological subdivisions. For example, the Upper Rakhi Gaj Shales, Green and
Nodular Shales and Rubbly Limestones have a similar ostracod fauna and are
regarded as one ostracod biostratigraphic unit. Another example is in the Kirthar
Formation where a new fauna appears before a change in lithology. This occurs in
the uppermost part of the Lower Chocolate Clays (i.e. just below the White Marl
Band). Most of the species range from the uppermost part of the Lower Chocolate
Clays to the lower part of the Upper Chocolate Clays, although the White Marl Band
lies in between.

The Palaeocene/Lower Eocene boundary in the Rakhi Nala section has been
drawn at the base of the Nummulites ivregularis Limestone of Eames and Nagappa.

92 EARLY TERTIARY OSTRACODA

(Bayliss, however, identified this as Nummulites crasseornata (Henrici) form B.)
This is in agreement with Eames, Bayliss and Latif. The ostracod faunal assemblages
below and above the irvregularis Limestone are completely different and have no
species in common. These assemblages have been included in ostracod biostrati-
graphic Units II and III respectively. The Palaeocene and Lower Eocene boundary
in fact has been placed between these two biostratigraphic units. The ranges of
ostracod species found in the two biostratigraphic units are shown in Appendix 2.

The Shales below the ivregularis Limestone have a very rich assemblage of pelagic
Foraminifera and have been assigned to the Globrotalia rex Zone by Latif. Dr.
Banner, lately of the British Petroleum Co. Ltd., who very kindly examined these
samples, considers them to be of the high Globorotalia rex Zone with derived Lower/
Middle Palaeocene pelagic Foraminifera.

The Lower/Middle Eocene boundary in the Rakhi Nala, Zao River and Shpalai
Khwara sections has been placed at the base of the Platy Limestone. This is in
conformity with Eames, who examined the Rakhi Nala and Zinda Pir sections of the
Sulaiman Range. Bayliss and Latif, who worked on the Rakhi Nala section, how-
ever, have drawn the boundary in the uppermost part of the Lower Chocolate Clays
(i.e. below sample 3498). The Platy Limestone serves as an important horizon
marker in the region. The Lower/Middle Eocene boundary lies between ostracod
biostratigraphic Units IV and V, which have very different ostracod assemblages.
Except for Alocopocythere transcendens sp. nov., none of the Lower Eocene ostracod
species survive into the Middle Eocene.

The Middle/Upper Eocene boundary in the Rakhi Nala and Zao River sections has
been placed between the lower and upper parts of the Upper Chocolate Clays. The
upper part of the Upper Chocolate Clays contains the genus Pellatispira, which is of
Upper Eocene age. In the Rakhi Nala section the boundary is taken arbitrarily
between samples 3627 and 3628. This is approximately the same level as drawn by
Eames (1952), who recorded the first appearance of Pellatispira just above this
horizon in the section. Bayliss, however, recorded Pellatispira only from one
horizon (sample 3657) in the Pellatispira Beds. Latif has placed the boundary in the
middle of his Chiloguembelina aff. martini Zone (i.e. above sample 3618). In the
Zao River section sample 24161 contains specimens of Pellatispira in abundance.
These have been assigned to Pellatispira orbitoidea (Povale) sensu Rao 1941 by Dr.
C. G. Adams of the British Museum (personal communication), who very kindly
examined these specimens. According to Dr. Adams these fall midway between
P. orbitoidea and P. madaraszi var. indica. The Middle/Upper Eocene boundary in
the Zao River section can, therefore, safely be placed below sample 24161.

The Middle and Upper Eocene ostracod fauna of the Rakhi Nala and Zao River
sections is of changing suite and has been included in ostracod biostratigraphic
Unit V. It does not show any sharp break between the Middle and the Upper
Eocene. Some species are restricted to the Middle Eocene, but others range from
the Middle to the Upper Eocene. Some of the important species restricted to the
Middle Eocene are : Actinocythereis ? quasibathonica sp. nov., ““ Anommatocythere ”’
confirmata sp. nov., Echinocythereis (Scelidocythereis) rasilis sp. nov., Cytheropteron
sp.C, Gyrocythere exaggerata sp. nov., Trachyleberis (Trachyleberis) bimammillata

FROM WEST PAKISTAN 93

sp. nov. Some of the common species which range from the Middle to the Upper
Eocene are : Alocopocythere transversa sp. nov., Bairdoppilata sp.A, Cytherelloidea
cf. C. costatruncata Lubimova and Mohan, Cytheromorpha sp.A, Cytheropteron sp.D,
Pavenborchella sp.C, Patagonacythere ? nidulus sp. nov., Pterygocythereis (Pterygocy-
there) sp.A and Stigmatocythere lumaria sp. nov. (A complete list of these species is
given in Tables 4 & 5).

VI. APPENDICES
APPENDIX I

SOR RANGE LEASE 58, MEASURED SECTION AT LOCALITY 460.
SECTION MEASURED BY J. A. REINEMUND.

Thickness (feet)

10+ Claystone, grey

42 Conglomerate containing limestone and chert
pebbles, cobbles, boulders as much as 8 in.
across ; matrix of medium grained, yellowish
brown sandstone forms about 20% of rock.

42 Concealed by talus

18 Claystone, medium grey, not fissile, semiplastic,
containing scattered fossils. Calcareous nodules
at top ; silty and carbonaceous in lower part.

2-3 Sandstone, very fine grained to silty brownish-

grey, imperfect and irregular bedding, contains
carbonized plant fragments and vertical root
moulds.

Description of Unit Sample No.

460a (near top)

24

N
bol

iS)

Claystone, dark olive grey, not fissile, silty in
upper few feet and lower few feet ; contains
irregular coal layers as much as 4 in. thick in
lower 2 ft.

Sandstone, fine to very fine grained, light brown-
ish grey, mostly even beds 2-8 in. thick, locally
cross-bedded.

Siltstone, poorly bedded, carbonaceous, contain-
ing very carbonaceous layers as much as }¢ in.
thick in top 2 in.

Claystone, silty, olive grey, not fissile, containing
carbonised plant chips.

Siltstone, brownish grey, imperfect beds, cross
laminated.

Claystone, silty at top, olive grey not fissile.

460b (6-8 above
bottom)

460c (3-4 feet
above bottom)

460d

460e (5-6 feet
above bottom)

46of (1-2 feet
above bottom)

94

Thickness (feet)

EARLY TERTIARY OSTRACODA

Description of Unit

8 Claystone, containing profuse white calcareous
concretions.
2 Claystone, olive grey, fissile, grading down into

siltstone, yellowish-brown, hard, fossiliferous.

50=-

Claystone, slightly fissile, olive grey.

APPENDIX 2

Sample No.

460g (Channel
sample)
460h—o

LIST OF OSTRACOD SPECIES COMMON TO THE RAKHI
NALA AND ZAO RIVER SECTIONS

Species

“ Anommatocythere’”’ laqueta sp. nov.
Phalcocythere dissenta sp. nov.
Stigmatocythere obliqua sp. nov.
Cytherella sp.C

Neocypridets ? sp.G
Neocypridets sp.C

Xestoleberis sp.C

Xestoleberis sp.D

Xestoleberis sp.E

Genus C sp.1

Genus C sp.2

Bairdia sp.C

Bairdia sp.D

Cytherella sp.B

Pontocyprella sp.B
Pontocyprella sp.C

Actinocytherets ? quasibathonica sp. nov.

Alocopocythere transcendens sp. nov.

. transversa sp. nov. Morphotype A

. transversa sp. nov. Morphotype B

. transversa sp. nov. Morphotype C

. transversa sp. nov. Morphotype D

. ransversa sp. nov. Morphotype E

. transversa sp. nov. Morphotype F

. radiata sp. nov.

“ Anommatocythere’’ confirmata sp. nov.

Echinocytherets (Scelidocytherets) rasilis
Sp. nov.

E.(S.) multibullata sp. nov.

Gyrocythere exaggerata sp. nov.

Hermanites palmatus sp. nov.

my AR AAR AR AS

Rakhi Nala
Base Top
4045 5223
4968 = 5195
4815 5373
5112 5195
4815 5373

2373
4919 5208
5112 5195
4815 5208
4687 5304
4687 5373
3682+ 4815
3995 5195
2687+ 5223
5023 5304
4687 5208
6369 6524
3215+ 6488
6575 6706
6589+ 6690
6764 7014
68391-87537
6839 = 7107
6943 7037

7029
6138 6424
6138 6463
6531 = 6589+
6401 6515
6401 6488

Zao River

Base Top

252

252
252 294

252
252 294
252 294
252 294

252

252

252

252

252

252

252

252

252

1872
650+
2032

1994
1948
2190
3384+
3230
3230

2190
2504.
2086 3186
2504 2808
2712
1570 1948

1986
3095
1948
2060

1570
1994
1570

994

FROM WEST PAKISTAN

No. Species

It Hermanites scopus sp. nov.

12 Patagonacythere ? nidulus sp. nov.

15a Stigmatocythere lumaria sp. nov.
Morphotype A

15b S.lumariasp. nov. Morphotype B

16 Tvrachyleberis (Trachyleberis) bimammuillata

Sp. nov.
17 T.(Acanthocytherets) postcornis sp. nov.
18 T.(Acanthocytherets) decoris sp. nov.
196 - Aglatocypris sp.B
20 Bairdoppilatasp.A
22 Cytherellasp.E
23. Cytherellasp.F
24 Cytherellasp.G
25 Cytherelloidea cf. C.
costatruncata Lubimova and Mohan
26 = Cy therelloidea sp.E
27 Cytherelloidea sp.F
28 Cytheromorpha sp.A
29 Cytheropteron sp.C
30 ©. Cytheropteron sp.D
31 Knithe sp.C
32 ~@©Krithesp.D
33 Krithesp.E
34 Knrithesp.F
37. + Neocyprideis ? sp.D
38 Parenborchella sp.C
39 ~3=—©- Patjenborchella sp.E
40 Pazenborchella sp.F
41 Propontocypris sp.A
42 Pterygocythereis (Pterygocythere) sp.A
43 Schizocythere sp.B
47 Xestoleberis sp.G

+ Points omitted from computation.

Rakhi Nala
Base Top
6138 6488
6575 7218
6531 Fon
6839 7029
6369 6463
6121 6138
6200 7124

6415
6138 7014
6121 6401
6424 6985
6121 6985
6121 7001
6985 7029
6121 7124
6138 6974
6121 6606
6121+ 7029
6121 7096
6311 7107
6138 7014
6931 6985
7096 7124
6121 7096
6311 7096
7114 7124
6266 6564
6575 6985
6138 6524
6138 6931

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Cahiers géol., Paris, 28/29 : 241-279, pls. 1-8.

Zao River
Base Top
1766 1872
2032 3384
2060 3384
2712
1872 1948
994 1766
1994 3384
2190 3040+
794 3186
994 2032
994 3040
994 3095
994 3095
3000 3095
994 3095
994 2904
994 1872
2086+ 3095
1766 3095

1710 3140
1710 3140

2504 2760
3344 3450
794 3095
1900 2712
3186 = 3450
1766 2266
2060 3130
994 1994
1766 3130

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Eames, F. E., BANNER, F. T., Blow, W. H. & CLARKE, W. J. 1962. Fundamentals of Mid-
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FROM WEST PAKISTAN 97

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HornrBrook, N. DE B. 1952. Tertiary and Recent marine Ostracoda of New Zealand.
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Surv. Fla., 34 : 1-43.

1962. Ostyvacod Taxonomy. Louisiana Univ. Press. 366 pp.

Howe, H. V. & CHAMBERS, J. 1935. Louisiana Jackson Eocene Ostracoda. Bull. geol. Surv.
La., Baton Rouge, 5 : 1-65.

Howe, H. V. & Garrett, J. B. 1934. Louisiana Sabine Eocene Ostracoda. Dept. Cons.
Bureau Sci. Res. Sta. Mineral Section, geol. Bull., 4 : 1-64.

Howe, R.C. 1963. Type Saline Bayou Ostracoda of Louisiana. Bull. geol. Surv. La., Baton
Rouge, 40 : 1-62, pls. 1-4.

IMBRIE, J. 1956. Biometrical methods in the study of invertebrate fossils. Bull. Amer. Mus.
Nat. Hist., New York, 108 (2) : 215-252.

Ker, A. J. 1957. Eocene and Oligocene Ostracoda of Belgium. Inst. voy. Sci. Nat. Belgique,
Brussels, Mem. 136 : 1-210.

Kinema, J.T. 1948. Contributions to the Knowledge of the Young-Caenozoic Ostracoda from
the Malayan Region. Thesis Univ. Utrecht, 119 pp.

KrIsHNAN, M. S. 1960. Geology of India and Burma. Edition 4, Higginbothams Ltd.,
Madras, 604 pp.

LatHam, M. H. 1938. Some Eocene Ostracoda from North-West India. Pyvoc. R. Soc.
Edinb. 59 (1) : 38-48, text figs. 1-8.

Latir, M. A. 1961. The use of pelagic Foraminifera in the sub-division of the Palaeocene-
Eocene of the Rakhi Nala, West Pakistan. Geol. Bull. Panjab Univ., Lahore, 1: 31-46,
pls. 1-5.

1963. Some related groups of pelagic Foraminifera in the Palaeocene-Eocene of the

Rakhi Nala, West Pakistan. Geol. Bull. Panjab Univ., Lahore, 3 : 19-24, pls. 1-3.

1964. Variation in abundance and morphology of pelagic Foraminifera in the Palaeocene-
Eocene of the Rakhi Nala, West Pakistan. Geol. Bull. Panjab Univ., Lahore, 4 : 29-100,
pls. 1-11.

Lusimova, P. S., Guua, D. K. & Mowan, H. 1960. Ostracoda of Jurassic and Tertiary
deposits from Kutch and Rajasthan (Jaisalmer), India. Bull. geol. Min. Met. Soc. India,
Calcutta, 22 : 1-60, pls. 1-4.

Moors, R. C. (Editor). 1961. Tveatise on Invertebrate Paleontology. Part Q, Arthropoda 3.
Geological Society of America and University of Kansas Press.

Moos, B. 1963. Uber einige der ‘‘ Cythere macropora’’ Bosquet 1852 (Ostr.) ahnliche Arten

aus verschiedenen Tertiarstufen. Geol. Jb., Hannover, 82 : 21-42, pls. 1-2.

1965. Die ostraceoden-Fauna des Unteroligozans von Biinde (Bl. Kerford-West 3817)

und einige verwandte jiingere Arten (Ostr., Crust.). Geol. Jb. Hannover, 82 : 593—630,
pls. 34-39.

NaGcappa, Y. 1959. Foraminiferal biostratigraphy of the Cretaceous-Eocene succession in the
India-Pakistan-Burma region. Micropaleontology, New York, 5 (2) : 145-192.

Neviani, A. 1928. Ostracodi fossili d'Italia. 1. Vallebiaja (Calabriano). Memorie Accad.
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OeErRTLI, H. J. 1956. Ostracoden aus der Oligozanen und Miozanen Molasse der Schweiz.
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Rec. geol. Surv. India, Calcutta, 49 (3) : 137-160.

98 EARLY TERTIARY OSTRACODA

PINFOLD, E. S. 1939. The Dunghan Limestone and the Cretaceous-Eocene unconformity in

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1954. Contribution to the study of the Miocene of the Florida Panhandle. Bull. geol.

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Soun, I. G. Early Tertiary ostracodes from West Pakistan. Mem. geol. Surv. Pakist. 3: 4 pls.
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SYLVESTER-BRADLEY, P.C. 1941. The shell structure of the ostracoda and its application to

their palaeontological investigation. Ann. Mag. nat. Hist., London, Ser. 11, 8 : I-33.

1948. The ostracode genus Cythereis. J. Paleont., Tulsa, 22 (6) : 792-797.

1951. The Subspecies in Palaeontology. Geol. Mag. London, 88 (2) : 88-102.

—— 1956. The structure, evolution and nomenclature of the ostracod hinge. Bull. Br. Mus.
nat. Hist. (Geol.), London, 3 (1) : 1-21.

SYLVESTER-BRADLEY, P. C. & Harpinec, J. P. 1954. Postscript notes of the ostracode
Trachyleberis. J. Paleont., Tulsa, 28 (5) : 560-562.

Tewarl, B. S. & Tanpon, K. K. 1960. Kutch Microfauna—Lower Tertiary Ostracoda.
Proc. natn. Inst. Sci. India Calcutta, 26B, 4 : 147-167, pls. 1-6.

TRIEBEL, E. 1958. Zwei neue Ostracoden-Gattungen aus dem Lutet des Pariser Beckens.
Senckenberg. leth., Frankfurt a.M. 39 (1-2) : 105-117, 3 pls.

1961. Geschlechts-Dimorphismes und Asymmetric der Klappen bei der Ostracoden-
gattung Occultocythereis. Senckenberg. leth., Frankfurt a.M. 42 (3-4) : 205-225, 5 pls.
VAN HinTE, J.E. 1962. Ostracoden aus dem Alttertiar des Sonnberges, Karnten, Oesterreich.

Proc. K. ned. Akad. Wet., Amsterdam, Ser. B 65 (2) : 166-189.

1964. A new Occultocythereis species of the Austrian Eocene. Proc. K. ned. Akad. Wet.,
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Van MorkHoven, F. P.C. M. 1962. Post-Palaeozoic Ostracoda. 1, 204 pp. Elsevier Pub-

lishing Company, Amsterdam.

1963. Post-Palaeozoic Ostracoda. 2,478 pp. Elsevier Publishing Company, Amsterdam.
Wanpia, D.N. 1953. Geology of India. MacMillan and Co. Ltd.

WittiaMs, M.D. 1959. Stratigraphy of the Lower Indus Basin, West Pakistan. Fifth world
Petrol. Cong., New York, section 1, paper 19, pp. 377-394, text-figs. 1-5.

QADEER AHMAD Sipp1QuI, M.Sc., Ph.D.,
Dept. of Geology,

St. Mary’s UNIVERSITY,

Ha.iFax, Nova Scotia,

CANADA.

4

PLATE 1

Actinocythereis ? quasibathonica sp. nov.

Fics. 1, 10. Dorsal and right views, carapace male, xX 70. Paratype, Io. 4260. Upper
Chocolate Clays (lower part), sample 3613, Rakhi Nala.

Fics. 2, 3, 11, 12. Dorsal, ventral, left and right views, carapace female, x 70. Holotype,
Io. 4311. Upper Chocolate Clays (lower part), sample 3611, Rakhi Nala.

Fic. 6. Dorsal view of hinge, left valve male, x 230. Paratype, Io. 3101. Upper Choco-
late Clays (lower part), sample 3613, Rakhi Nala.

Fics. 7, 13. 7, dorsal view of hinge x 230 ; 13, anterior radial pore canals x 230. Right
valve male. Paratype, Io. 3100. Upper Chocolate Clays, sample 3611, Rakhi Nala.

Alocopocythere transcendens gen. et sp. nov.
Fics. 4, 5, 8, 9. Dorsal, ventral, right and left views, carapace male, x 90. Paratype,
Io. 3104. Upper Chocolate Clays (lower part), sample 3607, Rakhi Nala.

PLATE 1

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w
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9

PLATE 2
Alocopocythere transcendens gen. et sp. nov.

Fics. 1,6. External and dorsal views, left valve female, x 90. Holotype, lo. 4315. Upper
Chocolate Clays (lower part), sample 24148, Zao River.

Fic. 2. Muscle scars ( 200) showing four adductors and an oval frontal scar. Right valve
male (broken), x 90. Paratype, lo. 3106. Upper Chocolate Clays, sample 24151, Zao River.

Fic. 3. Anterior radial pore canals x 128. Left valve female. Paratype, lo. 4261. Upper
Chocolate Clays (lower part), sample 24148, Zao River.

Fics. 4, 7. External and dorsal views, right valve female, x 90. Paratype, lo. 3105.
Upper Chocolate Clays (lower part), sample 24148, Zao River.

Alocopocythere rupina sp. nov.
Fics. 5, 8-10. Right, left, dorsal and ventral views, carapace male x 90. Holotype,
Io. 4314. Gorge Beds, sample 3111, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PATE 2

PLATE 3

Alocopocythere rupina sp. nov.
Fics. 1-4. Left, mght, dorsal and ventral views, carapace female, x 90. Paratype,
Io. 4262. Gorge Beds, sample 3111, Rakhi Nala.

Alocopocythere abstracta sp. nov.
Fics. 5-8. Left, dorsal, ventral and right views, carapace male, x 90. Paratype, lo. 4263.
Upper Rakhi Gaj Shales, sample 3163, Rakhi Nala.
Fics. g-11. Right, dorsal and ventral views, carapace female <x 90. Holotype, Io. 4312.
Upper Rakhi Gaj Shales, sample 3163, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 3

PLATE 4
Alocopocythere abstracta sp. nov.

Fic. 1. Left view, carapace female, x 90. Holotype, Io. 4312. Upper Rakhi Gaj Shales,

sample 3163, Rakhi Nala.

Alocopocythere coarctata sp. nov.

Fics. 2-5. Dorsal, ventral, left and right views, carapace male, x 90.

Shales with Alabaster, sample 3448, Rakhi Nala.

Fics. 6-9. Dorsal, ventral, left and right views, carapace female, x go.

Shales with Alabaster, sample 3458, Rakhi Nala.

Alocopocythere longilinea sp. nov.

Fics. 10-13. Dorsal, right, ventral and left views, carapace male, x go.

Shales with Alabaster, sample 3443, Rakhi Nala.

Paratype, lo. 4264.

Holotype, Io. 4313.

Holotype, Io. 4318

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9

PLATE 5

Alocopocythere longilinea sp. nov.
Fics. 1-3, 6. Left, right, dorsal and ventral views, carapace female, x 90. Paratype,
Io. 4265. Shales with Alabaster, samples 3443, Rakhi Nala.

Alocopocythere transversa sp. nov.
Morphotype A
Fics. 4, 5, 7, 9. Dorsal, ventral, left and right views, carapace male, x go. Paratype,
Io. 4266. Upper Chocolate Clays (lower part), sample 24155, Zao River.
Fics. 8, 10. Left and right views, carapace female, x 90. Holotype, lo.4316. Upper
Chocolate Clays (lower part), sample 24155, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 5

PLATE 6
Alocopocythere transversa sp. nov.
Morphotype A

Fics. 1, 2. Dorsal and ventral views, carapace female, x 90. Holotype, lo. 43106.

Chocolate Clays (lower part), sample 24155, Zao River.

Fics. 3, 4. Dorsal and internal views, right valve male, x 90. Paratype Io. 3107.

Chocolate Clays (lower part), sample 3625, Rakhi Nala.

Morphotype C

Fics. 5, 7. Left and right views, carapace male, x 90. Paratype, Io. 4267.

Chocolate Clays (upper part), sample 24183, Zao River.

Fics. 6, 8. Left and right views, carapace female, x 90. Paratype, lo. 4268.

Chocolate Clays (upper part), sample 24183, Zao River.

Upper

Upper

Upper

Upper

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 6

PLATE 7

Alocopocythere transversa sp. nov.
Morphotype C
Fics. 1, 2. Dorsal and ventral views, carapace male, x 90. Paratype, Io. 42607. Upper
Chocolate Clays (upper part), sample 24183, Zao River.
Fics. 3, 4. Dorsal and ventral views, carapace female, x 90. Paratype, lo. 4268. Upper
Chocolate Clays (upper part), sample 24183, Zao River.

Morphotype E
Fics. 5-7. Dorsal, ventral and left views, carapace male, x 90. Paratype, lo. 3110. Upper
Chocolate Clays (upper part), sample 24175, Zao River.
Fic. 8. Left view, carapace female, x 90. Paratype, lo. 3111. Upper Chocolate Clays
(upper part), sample 24175, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 7

4"

. 3

PLATE 8
Alocopocythere transversa sp. nov.
Morphotype E
lic. 1. Right view, carapace male, x 90. Paratype, Io. 3110. Upper Chocolate Clays
(Upper part), sample 24175. Zao River.
Fics. 2, 3, 5. Right, dorsal and ventral views, carapace female, x 90. Paratype, Io. 3111
Upper Chocolate Clays (upper part), sample 24175, Zao River.

Morphotype C
Fic. 4. Muscle scars (x 200) showing four adductor, an oval frontal and two mandibular
scars. Right valve male (broken). Paratype, Io. 4269. Upper Chocolate Clays (upper part),
sample 24174, Zao River.

Morphotype F
Fics. 6, 8. Left and right views, carapace male, x 90. Paratype, Ilo. 3109. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.
Fics. 7, 9. Left and right views, carapace female, x 90. Paratype, Io. 3108. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.

00
ey
Gal
a

Suppl. 9

-)

l

Mus. nat. Hist. (Geo

Bull. Br.

PLATE 9

Alocopocythere transversa sp. nov.
Morphotype F

Fics. 1, 2. Dorsal and ventral views, carapace male, x 90. Paratype, Io. 3109. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.

Fics. 3, 5. Dorsal and ventral views, carapace female, x 90. Paratype, Io. 3108. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.

Fic. 4. Internal view to show radial pore canals, right valve female, * 108. Paratype,
Io. 3112. Upper Chocolate Clays (upper part), sample 24174, Zao River.

Alocopocythere radiata sp. nov.
Fics. 6, 8. Left and right views, carapace male, x 90. Holotype, lo. 4317. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.
Fics. 7, 9. Left and right views, carapace female, x 90. Paratype, lo. 4270. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.

PLATE 9

ey)
a
Qy
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id p)
S
(
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S
Ss
oO
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=~
isa]

PLATE to

Alocopocythere radiata sp. nov.
Fies. 1, 2. Dorsal and ventral views, carapace male, x 90. Holotype, lo. 4317. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.
Fics. 3, 4. Ventral and dorsal views, carapace female, x 90. Paratype, lo. 4270. Upper
Chocolate Clays (upper part), sample 3652, Rakhi Nala.

“ Anommatocythere ” laqueta sp. nov.
Fics. 5-7. Right, left and dorsal views, carapace male, x 70. Paratype, lo. 4271. Green
and Nodular Shales, sample 3403, Rakhi Nala.
Fics. 8-10. Left, right and dorsal views, carapace female, x 70. Holotype, lo. 4320.
Green and Nodular Shales, sample 3403, Rakhi Nala.

“ Anommatocythere ” confirmata sp. nov.
Fics. 11, 12. Right and left views, carapace male, x 70. Holotype, lo. 4319. Upper
Chocolate Clays (lower part), sample 3611, Rakhi Nala.

fo)
a
a

Mus. nat. Hist. (Geol.)

Bull. Br.

PLATE 11

“ Anommiatocythere ” confirmata sp. nov.

Fics. 1, 2. Dorsal and ventral views, carapace male, x 70. Holotype, Io. 4319 Upper
Chocolate Clays (lower part), sample 3611, Rakhi Nala.

Fic. 3. Anterior radial pore canals x 232, left valve male. Paratype, Io. 3102. Upper
Chocolate Clays (lower part), sample 24151, Zao River.

Fics. 4, 5, 8, 9. Dorsal, ventral, right and left views, carapace female, x 70. Paratype,
lo. 4272. Upper Chocolate Clays (lower part), sample 24148, Zao River.

Fics. 6, 7. Anterior and posterior radial pore canals x 232, right valve male. Paratype,
Io. 3103. Upper Chocolate Clays (lower part), sample 24151, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9

PLATE 11

PVA Er 2
“ Anommiatocythere ”’ confirmata sp. nov.
Fic. 1. Dorsal view of hinge x 183, left valve male. Paratype, lo. 3102. Upper Chocolate

Clays (lower part), sample 24151, Zao River.
Fic. 2. Dorsal view of hinge x 183, right valve male. Paratype, lo. 3103. Upper Choco-

late Clays (lower part), sample 24151, Zao River.

Bradleya ? voraginosa sp. nov.
Fics. 3, 5, 7, 8. Dorsal left, right and ventral views, carapace male, x 70. Holotype,

lo. 4321. Upper Chocolate Clays (upper part), sample 24161, Zao River.
Fics. 4, 6. 9. Dorsal, right and left views, carapace female, x 70. Paratype, lo. 3115.

Upper Chocolate Clays (upper part), sample 24161, Zao River.

PLATE 12

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9

PIL INANIS, 403)

Buntonia devexa sp. nov.
Fics. 1, 3. Left and right views, carapace male, x 70. Paratype, Io. 3113. Gorge Beds,
sample 3111, Rakhi Nala.
Fics. 2, 4, 5. Left, dorsal and right views, carapace female, x 70. Holotype, Io. 4322.
Gorge Beds, sample 3111, Rakhi Nala.

Buntonia sp.A
Fics. 6, 7, 9. Left, right and dorsal views, carapace, x 70. Paratype, lo. 3114. Lower
Rakhi Gaj Shales, sample 3133, Rakhi Nala.

Costa (Paracosta) declivis subgen. et sp. nov.
Fics. 8, 10-12. Left, right, dorsal and ventral views, carapace male, x 68. Holotype,
lo. 4325. Pellatispira Beds, sample 3662, Rakhi Nala.
Fic. 13. Right view, carapace female, x 68. Paratype, lo. 4273. Pellatispiva Beds,
sample 3662, Rakhi Nala.

Fic. 14. Left view, carapace female, x 68. Paratype, Ilo. 3116. Pellatispiva Beds, sample
36062, Rakhi Nala.

—
fal
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Ay
5)

Nn

x

(CG

Hist.

nat.

Mus.

Bull. Br.

PLATE 14

Costa (Paracosta) declivis subgen. et sp. nov.
Fic. 1. Dorsal view, carapace female, x 68. Paratype, Io. 4273. Pellatispiva Beds,
sample 3662, Rakhi Nala.
Fic. 2. Ventral view, carapace female, x 68. Paratype, lo. 3116. Pellatispiva Beds,
sample 3662, Rakhi Nala.

Costa (Paracosta) compitalis sp. nov.
Fics. 3, 4, 7, 8. Dorsal, ventral, right and left views, carapace male, x 68. Paratype,
Io. 4274. Upper Chocolate Clays (lower part), sample 3604, Rakhi Nala.
Fics. 5, 6, 9, 10. Dorsal, ventral, right and left views, carapace female, x 68. Holotype,
Io. 4323. Upper Chocolate Clays (lower part), sample 3604, Rakhi Nala.

Costa (Paracosta) disintegrata sp. nov.
Fic. 11. Left view, carapace male, x 68. Paratype, Ilo. 4275. Upper Chocolate Clays
(lower part), sample 3622, Rakhi Nala.

PLATE 14

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ai
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o
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8
4
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%

PLATE 15

Costa (Paracosta) disintegrata sp. nov.
Fics. 1, 2, 5, 6. Left, right, dorsal and ventral views, carapace female, x 68. Holotype,
lo. 4324. Upper Chocolate Clays (lower part), sample 3621, Rakhi Nala.
Fics. 3, 4. Dorsal and ventral views, carapace male, x 68. Paratype, lo. 4275. Upper
Chocolate Clays (lower part), sample 3622, Rakhi Nala.

Echinocythereis (Echinocythereis) contexta sp. nov.
Fics. 7, 10. Left and dorsal views, carapace male, x 68. Paratype, Ilo. 4276. Upper
Palaeocene, sample 460-j], Sor Range, 8 miles east of Quetta.
Fics. 8, 13. Left and dorsal views, carapace female, x 68. Holotype, lo. 4326. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Echinocythereis (Echinocythereis) elongata sp. nov.
Fics. 9, 11. Left and right views, carapace male, x 68. Paratype, lo. 3130. MRubbly
Limestones, sample 3416, Rakhi Nala.
Fics. 12, 14. Left and right views, carapace female, x 68. Holotype, lo. 4327. Rubbly
Limestones, sample 3416, Rakhi Nala.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 15

PLATE 16

Echinocythereis (Echinocythereis) elongata sp. nov.
Fic. 1. Dorsal view, carapace female, x 68. Holotype, Io. 4327. Kubbly Limestones,
sample 3416, Rakhi Nala.
Fic. 2. Dorsal view, carapace male, x 68. Paratype, Io. 3130. KRubbly Limestones,
sample 3416, Rakhi Nala.

Echinocythereis (Scelidocythereis) multibullata subgen. et sp. nov.
Fics. 3,5, 6. Dorsal, left and right views, carapace male, x 68. Holotype, lo. 4328. Upper
Chocolate Clays (upper part), sample 24161, Zao River.
Fics. 4, 7, 8. Dorsal, right and left views, carapace female, x 68. Paratype, lo. 3134.
Upper Chocolate Clays (upper part), sample 24161, Zao River.
Fic. 9. Internal view to show radial pore canals, right valve female, « 132. Paratype,
Io. 4277. Upper Chocolate Clays (lower part), sample 24159, Zao River.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 16

PLATE 17
Echinocythereis (Scelidocythereis) multibullata sp. nov.

Fic. 1. Dorsal view of hinge, left valve female, x 146. Holotype, Io. 4327. Upper
Chocolate Clays (lower part), sample 24159, Zao River.

Fic. 2. Dorsal view of hinge, right valve female, x 146. Paratype, lo. 4277. Upper
Chocolate Clays (lower part), sample 24159, Zao River.

Fic. 7. Ventral view, carapace male, x 68. Paratype, lo. 3133. Upper Chocolate Clays
(upper part), sample 24161, Zao River.

Echinocythereis (Scelidocythereis) sp.A
Fics. 3, 4, 8, 9. Left, right, dorsal and ventral views, carapace, x 68. Io. 3129. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Echinocythereis (Scelidocythereis) rasilis sp. nov.
I'tcs. 5, 10. Left and ventral views, carapace male, x 68. Paratype, Ilo. 4278. Lower
Chocolate Clays, sample 3499, Rakhi Nala.
Fic. 6. Left view, carapace female, x 68. Holotype, lo. 4329. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 17

PLATE 18

Echinocythereis (Scelidocythereis) rasilis sp. nov.

Fic. 1. Dorsal view, carapace male, x 68. Paratype, lo. 4278. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Fics. 2, 3. Dorsal and ventral views, carapace female, x 68. Holotype, lo. 4329. Lower
Chocolate Clays, sample 3499, Rakhi Nala.

Fic. 5. Right view, carapace male, x 68. Paratype, Io. 3131. Upper Chocolate Clays
(lower part), sample 24157, Zao River.

Fic. 7. Right view, carapace female, x 68. Paratype, lo. 3132. Upper Chocolate Clays
(lower part), sample 24145, Zao River.

Echinocythereis (Scelidocythereis) sparsa sp. nov.
Fics. 4, 6. Dorsal and left views, carapace male, x 68. Paratype, Io. 4279. Upper
Chocolate Clays (lower part), sample 24159, Zao River.
Fics. 8, 9. Dorsal and left views, carapace female, x 68. Holotype, lo. 4330. Upper
Chocolate Clays (lower part), sample 24159, Zao River.

Gyrocythere exaggerata gen. et sp. nov.
Fics. to, 12. External and internal views, left valve male, x 68. Paratype, lo. 3125.
Upper Chocolate Clays (lower part), sample 24151, Zao River.
Fics. 11, 14. External and internal views, right valve male, x 68. Paratype, lo. 3127.
Upper Chocolate Clays (lower part), sample 24151, Zao River.
Fic. 13. Dorsal view, carapace male (specimen now split giving separate valves Io. 3125 +
Io. 3127 above).

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 18

PLATE 19

Gyrocythere exaggerata gen. et sp. nov.

Fics. 1-4. Left, right, dorsal and ventral views, carapace female, x 68. Holotype, lo. 4331.
Upper Chocolate Clays (lower part), sample 24151, Zao River.

Fic. 5. Muscle scars x 120 showing four adductors and a U-shaped frontal scar, right valve
female. Paratype, lo. 4280. Upper Chocolate Clays (lower part), sample 24148, Zao River.

Fic. 6. Internal view to show radial pore canals, left valve male, x 134. Paratype, lo. 3126.
Upper Chocolate Clays (lower part), sample 24151, Zao River.

Fic. 7. Internal view to show radial pore canals, right valve female, x 134. Paratype,
Io. 3124. Upper Chocolate Clays (lower part), sample 24148, Zao River.

Fic. 8. Dorsal view of hinge x 145, left valve male. Paratype, Io. 3120. Upper Chocolate
Clays (lower part), sample 24151, Zao River.

Fic. 9. Dorsal view of hinge x 145, right valve male. Paratype, lo. 3122. Upper Choco-
late Clays (lower part), sample 24151, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 19

PLATE 20
Gyrocythere exaggerata gen. et sp. nov.
Paratype, lo. 3128. Upper

Fic. 5. Dorsal view of hinge, right valve female, x 150.
Chocolate Clays (lower part), sample 24148, Zao River.
Gyrocythere parvicarinata sp. nov.
Fics. 1, 2, 6, 7. Right, left, dorsal and ventral views, carapace male, x 68. Holotype,
Io. 4334. Green and Nodular Shales, sample 3407, Rakhi Nala.
Right, left, dorsal and ventral views, carapace female, x 68. Paratype,

FIGs. 3, 4, 8, 12.
Io. 4281. Green and Nodular Shales, sample 3407, Rakhi Nala.

Gyrocythere grandilaevis sp. nov.
Shales with

Fics. 9, 10. Left and right views, carapace male, x 68. Holotype, Io. 4332.
Shales

Alabaster, sample 3463, Rakhi Nala.
Fics. 11, 12. Left and right views, carapace female, x 68.

with Alabaster, sample 3463, Rakhi Nala.

Paratype, Io. 4282.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9

PLATE 20

PLATE 2t1r

Gyrocythere grandilaevis sp. nov.
Fics. 1, 2. Dorsal and ventral views, carapace male, x 68. Holotype, Io. 4332. Shales
with Alabaster, sample 3463, Rakhi Nala.
Fics. 3, 4. Dorsal and ventral views, carapace female, x 68. Paratype, lo. 4282. Shales
with Alabaster, sample 3463, Rakhi Nala.

Gyrocythere mitigata sp. nov.
Fics. 5-8. Left, right, dorsal and ventral views, carapace male, x 70. Holotype, lo. 4333.
Lower Chocolate Clays, sample 24131, Zao River.
Fics. 9, 11. Dorsal and external views, left valve female, x 70. Paratype, lo. 3119.
Lower Chocolate Clays, sample 24131, Zao River.
Fic. 10. Right view, carapace female, x 70. Paratype, lo. 4283. Lower Chocolate Clays,
sample 34131, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 21

PLATE 22

Gyrocythere perfecta sp. nov.

Fics. 1, 2, 5, 9. Left, right, dorsal and ventral views, carapace male, x 68. Paratype,
Io. 4281. Lower Chocolate Clays, sample 3499, Rakhi Nala.

Fics. 3, 4, 7, 8. Left, right, dorsal and ventral views, carapace female, x 68. Holotype,
Io. 4335. Lower Chocolate Clays, sample 3499, Rakhi Nala.

Fic. 6. Muscle scars x 140, right valve male. Paratype, lo. 3121. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Fic. 10. Internal view to show radial pore canals, right valve female, x 134. Paratype,
Io. 3120. Lower Chocolate Clays, sample 3498, Rakhi Nala.

Hermanites cracens sp. nov.
Fic. 11. Ventral view, carapace, x 70. Holotype, Io. 4336. Gorge Beds, sample 3111,
Rakhi Nala.

Oo)

Mus. nat. Hist. (Geol.) Suppl.

Bull. Br.

PLATE 23

Hermanites cracens sp. nov.
Fics. 1-3. Left, right and dorsal views, carapace, x 70. Holotype, lo. 4336. Gorge Beds,
sample 3111, Rakhi Nala.

Hermanites scopus sp. nov.
Fics. 4-7. Left, dorsal, ventral and right views, carapace male, x 70. Holotype, Io. 4338.
Upper Chocolate Clays (lower part), sample 24148, Zao River.
Fics. 8-1o. Right, dorsal and ventral views, carapace female, x 70. Paratype, lo. 4285.
Lower Chocolate Clays, sample 3499, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 23

PLATE 24

Hermanites palmatus sp. nov.

Fics. 1, 2, 5. Dorsal, ventral and left views, carapace male, x 70. Paratype, lo. 4286.
Upper Chocolate Clays (lower part), sample 24156, Zao River.

Fics. 3, 4, 7. Dorsal, ventral and right views, carapace female, x 70. Paratype, lo. 3117.
Upper Chocolate Clays (lower part), sample 24156, Zao River.

Fics. 6, 8, 9, 11. Dorsal, external, internal and ventral views, left valve female, x 7o.
Holotype, lo. 4337. Upper Chocolate Clays (lower part), sample 24152, Zao River.

Fic. 12. External view, left valve female, x 70. Paratype, Io. 3118. Upper Chocolate
Clays (lower part), sample 3613, Rakhi Nala.

Occultocythereis interrupta sp. nov.
Fics. 10, 13, 15,16. Left, right, dorsal and ventral views, carapace male, x 116. Paratype,
lo. 4287. Upper Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.
Fics. 14, 17, 18. Right, dorsal and ventral views, carapace female, x 116. Holotype,
lo. 4339. Upper Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 24

PLATE 25
Occultocythereis sp.A
Fics. 1, 2,5. Left, right and ventral views, carapace, x 116. Io. 3136. Lower Rakhi Gaj
Shales, sample 3672, Rakhi Nala.

Occultocythereis spilota sp. nov.
Fics. 3, 4, 8, 9. Left, right, dorsal and ventral views, carapace male, x 120. Paratype,

Io. 4288. Green and Nodular Shales, sample 3177, Rakhi Nala.
Fics. 6,7, 10,11. Left, right, dorsal and ventral views, carapace female, x 120. Holotype,

Io. 4342. Green and Nodular Shales, sample 3177, Rakhi Nala.

Occultocythereis peristicta sp. nov.
Morphotype A
Fics. 13, 14,17. Left, right and ventral views, carapace male, x 118. Paratype, Io. 4292.
Upper Rakhi Gaj Shales, sample 3167, Rakhi Nala.
Fics. 15, 16. Left and right views, carapace female, x 118. Holotype, Io. 4341. Upper
Rakhi Gaj Shales, sample 3167, Rakhi Nala.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 25

PICA E26

Occultocythereis peristicta sp. noy.
Morphotype A
Fic. 1. Dorsal view, carapace male, x 118. Paratype, lo. 4292. Upper Rakhi Gaj Shales,
sample 3167, Rakhi Nala.
Fics. 2,3. Dorsal and ventral views, carapace female, x 118. Holotype, lo. 4341. Upper
Rakhi Gaj Shales, sample 3167, Rakhi Nala.

Morphotype B
Fics. 4, 6, 7. Ventral, left and right views, carapace male, x 118. Paratype, Io. 4293.
Green and Nodular Shales, sample 3193, Rakhi Nala.
Fics. 5, 8, 9. Ventral, left and right views, carapace female, x 118. Paratype, Io. 4291.
Green and Nodular Shales, sample 3193, Rakhi Nala.

Morphotype C
Fics. 10-12. Ventral, left and right views, carapace male, x 118. Paratype, lo. 4289.
Green and Nodular Shales, sample 3191, Rakhi Nala.
Fics. 13-15. Left, right and ventral views, carapace female, x 118. Paratype, lo. 4290.
Green and Nodular Shales, sample 3191, Rakhi Nala.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9

PLATE 27

Occultocythereis peristicta sp. nov.
Morphotype D
Fics. 1, 2, 5. Left, right and ventral views, carapace male, x 118. Paratype, Io. 3146.
Green and Nodular Shales, sample 3191, Rakhi Nala.
Fics. 3, 4, 6. Left, right and ventral views, carapace female, x 118. Paratype, Io. 3139.
Green and Nodular Shales, sample 3191, Rakhi Nala.

Morphotype E
Fics. 7, 9, 10. Ventral, right and left views, carapace male, x 118. Paratype, lo. 3137.
Rubbly Limestones, sample 3418, Rakhi Nala.
Fics. 8, 11, 12. Ventral, left and right views, carapace female, x 118. Paratype, lo. 3138.
Rubbly Limestones, sample 3418, Rakhi Nala.

Occultocythereis indistincta sp. nov.
Fics. 13, 14. Ventral and right views, carapace male, x 120. Paratype, lo. 3135. Lower
Chocolate Clays, sample 3499, Rakhi Nala.

Fic. 15. Right view, carapace female, x 120. Holotype, lo. 4340. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 27

IPL /NABIE: A213}

Occultocythereis indistincta sp. nov.
Fics. 1, 3, 4. Left, dorsal and ventral views, carapace female, x 120. Holotype, lo. 4340.
Lower Chocolate Clays, sample 3499, Rakhi Nala.
Fic. 2. Dorsal view, carapace male, x 120. Paratype, lo. 3135. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Patagonacythere ? nidulus sp. nov.
Fics. 5-8. Left, right, dorsal and ventral views, carapace male, x 70. Paratype, Io. 3096.
Upper Chocolate Clays (upper part), sample 24173, Zao River.
Fics. 9-12. Left, right, dorsal and ventral views, carapace female, x 70. Holotype, lo. 4349
Upper Chocolate Clays (lower part), sample 24173, Zao River.

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jor
io
3
nN
mH
O
+
”
+
8
a)
=~
S

Bull. Br.

PLATE 29

Patagonacythere ? nidulus sp. nov.
Specimens showing exaggerated normal pores after being cleaned in the
ultrasonic vibrator.

Fics. 1, 2. 1, Internal view to show radial pore canals, x 160 ; 2, Dorsal view of hinge,
x 150. Left valve female. Paratypes, Ilo. 3097. Upper Chocolate Clays (upper part), sample
24170, Zao River.

Fic. 3. Dorsal view of hinge, right valve female, x 150. Paratype, lo. 3098. Upper
Chocolate Clays (upper part), sample 24170, Zao River.

Fic. 4. Muscle scars < 260. Right valve male. Paratype, lo. 4295. Upper Chocolate
Clays (upper part), sample 34170, Zao River.

Phalcocythere horrescens (Bosquet) gen. nov.
Fic. 5. Anterior radial pore canals, * 212. Left valve. Io. 4257. Lutetian IV (sample
CAB 1002, Keij 1957, p. 19), Grignon, Paris Basin.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9

PLATE 29

PLATE 30
Phalcocythere horrescens (Bosquet) gen. nov.

Fics. 1, 2. External and internal views, left valve, x 68. Io. 4254. Lutetian IV (sample
CAB 1002, Keij 1957, p. 19), Grignon, Paris Basin.

Fics. 3, 4. Dorsal and external views, left valve, « 68. Io. 4253. Lutetian IV (sample
CAB 1002, Keij 1957, p. 19), Grignon, Paris Basin.

Fic. 5. Muscle scars x 195. Right valve. Io. 4255. Lutetian IV (sample CAB 1002,
Keij 1957, p. 19), Grignon, Paris Basin.

Fic. 6. Internal view, right valve, x 132. Io. 4256. lLutetian, Villiers-St.-Frederic, Paris
Basin.

Phalcocythere improcera sp. nov.
(See Pl. 33, figs. 12, 13 for hinge).

Fic. 7. Anterior radial pore canals x 224. Left valvefemale. Paratype, lo. 4295. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Fics. 8, 9. Left and right views, carapace male, x 68. Holotype, lo. 4344. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Fics. 10, 11. Left and right views, carapace female, x 68. Paratype, lo. 4258. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Fic. 12. Anterior radial pore canals x 224. Right valve female. Paratype, lo. 4296.
Upper Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 30

PLATE 31
Phalcocythere improcera sp. nov.
Fics. 1, 2. Dorsal and ventral views, carapace male, x 68. Holotype, lo. 4344. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.
Fics. 3, 4. Dorsal and ventral views, carapace female, x 68. Paratype, lo. 4258. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Phalcocythere rete sp. nov.

Fics. 5-8. Left, right, dorsal and ventral views, carapace male, x 68. Paratype, lo. 3099.
Upper Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.

Fics. g-10. Dorsal and left views, carapace female, x 68. Paratype, lo. 4297. Upper
Palaeocene, sample 460-i, Sor Range, 8 miles east of Quetta.

Fic. 11. External view, right valve female, x 68. Holotype, lo. 4348. Upper Palaeocene,
sample 460-1, Sor Range, 8 miles east of Quetta.

Fic. 12. External view of valve female, x 68. Paratype, lo. 3141. Upper Palaeocene,
sample 460-1, Sor Range, 8 miles east of Quetta.

Phalcocythere retispinata sp. nov.
Fics. 13, 14, 17. Left, right and dorsal views, carapace male, x 68. Paratype, lo. 3165.
Upper Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.
Fics. 15, 16. Left and right views, carapace female, x 68. Holotype, lo. 4345. Upper
Palaeocene, sample 460-i, Sor Range, 8 miles east of Quetta.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 31

PLATE 32
Phalcocythere retispinata sp. nov.
lic. 1. Ventral view, carapace male, x 68. Paratype, Io. 3165. Upper Palaeocene,
sample 460-i, Sor Range, 8 miles east of Quetta.
Fics. 2, 3. Ventral and dorsal views, carapace female, x 68. Holotype, lo. 4345. Upper
Palaeocene, sample 460-i, Sor Range, 8 miles east of Quetta.

Phalcocythere sentosa sp. nov.
Fics. 4, 5, 8, 10. Dorsal, left, right and ventral views, carapace male, x 68. Holotype,
lo. 4346. Upper Rakhi Gaj Shales, sample 3167, Rakhi Nala.
Fics. 6, 7, 9. Left, dorsal and right views, carapace female, x 68. Paratype, lo. 4208.
Upper Rakhi Gaj Shales, sample 3167, Rakhi Nala.

Phalcocythere dissenta sp. nov.
Fics. 11, 13, 14, 18. Left, right, dorsal and ventral views, carapace male, x 68. Holotype,
lo. 4343. Shales with Alabaster, sample 3456, Rakhi Nala.
Fries. 12, 15-17. Left, right, dorsal and ventral views, carapace female, x 68. Paratype,
Io. 4299. Shales with Alabaster, sample 3456, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 32

PEATE 33
Phalcocythere spinosa sp. nov.

Fics. 1, 2, 7, 8. Left, right, dorsal and ventral views, carapace, x 68. Holotype, Io. 4347.
Upper Chocolate Clays (upper part), sample 24161, Zao River.

Phalcocythere sp., cf. P. spinosa
lics. 3,4,9. Left, right and ventral views, carapace male, x 68. Io. 4230. Upper Eocene,
Lindi survey, 10-50 ft. above shore at Kitunga, Tanzania.
Fics. 5, 6, 10. Left, right and dorsal views, carapace female, x 68. Io. 4231. Upper
Eocene, Lindi survey, 10-50 ft. above shore at Kitunga, Tanzania.
Fic. 11. Muscle scars x 210, fragment of right valve female. Io. 4232. Upper Eocene,
Lindi survey, 10-50 ft. above shore at Kitunga, Tanzania.

Phalcocythere horrescens (Bosquet) gen. nov.
Fic. 12. Dorsal view of hinge x 183, left valve. Io. 4257. Lutetian IV (sample CAB 1002,
Keij 1957, p. 19), Grignon, Paris Basin.
Fic. 13. Dorsal view of hinge x 183, right valve. Io. 4255. Lutetian IV (sample CAB
1002, Keij 1957, p. 19), Grignon, Paris Basin.

Quadracythere (Hornibrookella) platybomus sp. nov.
Fics. 14, 18. Right and dorsal view, carapace male, x 70. Holotype, lo. 4351. Upper
Palaeocene, sample 460-1, Sor Range, 8 miles east of Quetta.
Fics. 15, 19. Right and dorsal views, carapace female, x 70. Paratype, lo. 4300. Upper
Palaeocene, sample 460-i, Sor Range, 8 miles east of Quetta.

Quadracythere (Hornibrookella) directa sp. nov.
Fic. 16. Left view, carapace male, x 70. Io. 4301. Green and Nodular Shales, sample
3403, Rakhi Nala.
Fic. 17. Left view, carapace female, x 70. Holotype, Io. 4350. Green and Nodular Shales,
sample 3403, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 33

PLATE 34
Quadracythere (Hornibrookella) directa sp. nov.
Fic. 1. Dorsal view, carapace male, x 70. Paratype, Ilo. 4301. Green and Nodular

Shales, sample 3403, Rakhi Nala.
Fic. 2. Dorsal view, carapace female, x 70. Holotype, lo. 4350. Green and Nodular

Shales, sample 3403, Rakhi Nala.

Quadracythere (Hornibrookella) arcana (Lubimova and Guha)
Fics. 3-5. Right, left and dorsal views, carapace, x 70. Io. 3142. Lower Chocolate Clays,
sample 3499, Ralkhi Nala.

Quadracythere (Hornibrookella) subquadra sp. nov.
Fics. 6, 8, 9. Left, dorsal and ventral views, carapace male, x 70. Paratype, lo. 4302.
Upper Chocolate Clays (upper part), sample 24161, Zao River.
Fics. 7, 10, 11. Left, right and dorsal views, carapace female, x 70. Holotype, Io. 4352.
Upper Chocolate Clays (upper part), sample 24161, Zao River.

Quadracythere (Hornibrookella) sp.A
Fics. 12-14. Left, right and dorsal views, carapace, x 70. Io. 3143. Upper Chocolate
Clays (lower part), sample 24148, Zao River.

PLATE 34

or)
=

or
=]
Y
~
DZ
~~
in

Bull. Br. Mu

PLATE 35

Stigmatocythere obliqua gen. et sp. nov.
Fic. 1. Dorsal view, carapace male, X 92. Specimen lost. Shales with Alabaster, sample

24173, Rakhi Nala.
Fics. 3, 4. Left and right views, carapace male, x 92. Paratype, Io. 4303.

Alabaster, sample 24173, Rakhi Nala.
Fics. 2, 5, 6. Dorsal, left and right views, carapace female, x 92. Holotype, Io. 4355.

Shales with Alabaster, sample 24173, Rakhi Nala.
Fic. 7. Dorsal view, left valve female, x 92.
sample 24173, Rakhi Nala.
Fic. 8. Dorsal view, right valve female, x 92.
sample 24173, Rakhi Nala.

Shales with

Paratype, lo. 3147. Shales with Alabaster,

Paratype, lo. 3148. Shales with Alabaster,

Fic. 9. Internal view to show radial pore canals, right valve female, x 160. Paratype,
Io. 3146. Shales with Alabaster, sample 24173, Rakhi Nala.
Fic. to. Internal view to show radial pore canals, left valve female, x 160. Paratype,

Io. 3149. Shales with Alabaster, sample 24173, Rakhi Nala.

Bull, Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 35

PLATE 36

Stigmatocythere obliqua gen. et sp. nov.
Fic. 1. Ventral view, carapace male, x 92. Paratype, lo. 4303. Shales with Alabaster,
sample 24173, Rakhi Nala.
Fic. 2. Ventral view, carapace male, * 92. Holotype, Io. 4355. Shales with Alabaster,
sample 24173, Rakhi Nala.

Stigmatocythere portentum sp. nov.
Fics. 3-6. Dorsal, ventral, left and right views, carapace male, x 92. Holotype, lo. 4357.
Lower Chocolate Clays, sample 3499, Rakhi Nala.
Fic. 10. Anterior radial pore canals x 216, fragment of right valve. Paratype, Io. 3144.

Stigmatocythere calia sp. nov.
Fic. 7. External view, right valve male, x 92. Paratype, lo. 3404. Upper Chocolate
Clays (lower part), sample 24152, Zao River.
Fics. 8, 9. Right and dorsal views, carapace female, x 92. Holotype, lo. 4353. Upper
Chocolate Clays (lower part), sample 24151, Zao River.

30

ATE

ale

I

=
oy
ral
ov
=
ar
O
~
D
Pa
eS
~

Mus.

Bull. Br.

PA 3 \7

Stigmatocythere calia sp. nov.
Fics. 1, 3. Left and ventral views, carapace female, x 92.
Chocolate Clays (lower part), sample 24151, Zao River.

Holotype, Io. 4353. Upper
Stigmatocythere delineata sp. nov.

Left dorsal, right and ventral views, carapace male, x 92. Paratype,

Io. 4305. Upper Chocolate Clays (lower part), sample 24154, Zao River.

Fics. 7-10. Dorsal, left, right and ventral views, carapace female, x 92. Holotype, Io. 4356.
Upper Chocolate Clays (lower part), sample 24154, Zao River.

Fics. 2, 4, 5, 6.

Stigmatocythere lumaria sp. nov.
Fic. 11. Anterior radial pore canals x 150, right valve female.

Paratype, lo. 3154. Upper
Chocolate Clays (upper part), sample 24174, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 37

PLATE 38

Stigmatocythere lumaria sp. nov.
Morphotype A

Fics. 1, 5, 6. Left, dorsal and ventral views, carapace male, x 92. Holotype, lo. 4354.
Upper Chocolate Clays (upper part), sample 3642, Rakhi Nala.

Fics. 2, 4, 7. External, internal and dorsal views, right valve male, x 92. Paratype,
lo. 3151. Upper Chocolate Clays (upper part), sample 3630, Rakhi Nala.

Fics. 3,8. External and dorsal views, right valve female, x 92. Paratype, lo. 4307. Upper
Chocolate Clays (upper part), sample 3630, Rakhi Nala.

Fic. 9. Dorsal view of hinge x 240, left valve female. Paratype, Io. 3152. Upper Choco-
late Clays (upper part), sample 24174, Zao River.

Fic. to. Dorsal view of hinge x 240, right valve female. Paratype, lo. 3154. Upper
Chocolate Clays (upper part), sample 24174, Zao River.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 38

PLATE 39

Stigmatocythere lumaria sp. nov.
Morphotype B
Figs. 1-4. Left, right, dorsal and ventral views, carapace male, x 92. Paratype, lo. 3150.
Upper Chocolate Clays (upper part), sample 3649, Rakhi Nala.
Fics. 5-8. Left, dorsal, ventral and right views, carapace female, x 92. Paratype, Io. 3153.
Upper Chocolate Clays (upper part), sample 3649, Rakhi Nala.

Morphotype A
Fic. 11. Anterior radial pore canals x 232, left valve male (juv.). Paratype, Io. 4306.
Upper Chocolate Clays (upper part), sample 24174, Zao River.

Trachyleberis (Trachyleberis) lobuculus sp. nov.
Fics. 9, 10. Left and dorsal views, carapace male, x 94. Paratype, lo. 4308. Upper
Rakhi Gaj Shales, sample 3163, Rakhi Nala.

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 39

a

:

PLATE 40
Trachyleberis (Trachyleberis) lobuculus sp. nov.
Fics. 1, 3. Left and dorsal views, carapace female, x 93. Holotype, lo. 4364. Upper
Rakhi Gaj Shales, sample 3166, Rakhi Nala.

Trachyleberis (Trachyleberis) bimammillata sp. nov.

Fics. 2, 8, 10. Left, right and dorsal views, carapace male, x 94. Holotype, lo. 4363.
Upper Chocolate Clays (lower part), sample 3613, Rakhi Nala.

Fic. 4. Dorsal view, left valve female, x 94. Paratype, Io. 3156. Upper Chocolate Clays
(lower part), sample 3611, Rakhi Nala.

Fic. 5. Dorsal view, right valve female, x 94. Paratype, lo. 3155. Upper Chocolate
Clays (lower part), sample 3614, Rakhi Nala.

Fic. 6. Dorsal view, carapace male, x 94. Paratype, lo. 3160. Upper Chocolate Clays
(lower part), sample 3613, Rakhi Nala.

Fic. 7. Dorsal view, carapace female, x 94. Paratype, lo. 3157. Upper Chocolate Clays
(lower part), sample 3613, Rakhi Nala.

Fic. 9. Left view, carapace female, x 94. Paratype, lo. 3159. Upper Chocolate Clays
(lower part) sample 3613, Rakhi Nala.

Fic. 11. Muscle scars x 172, fragment of left valve male. Paratype, lo. 3158. Upper
Chocolate Clays (lower part), sample 3613, Rakhi Nala.

Trachyleberis (Acanthocythereis) procapsus sp. nov.
Fic. 12. Left view, carapace male, x 94. Holotype, Io. 4360. Upper Palaeocene, sample
460-1, Sor Range, 8 miles east of Quetta.
Fic. 13. Left view, carapace female, x 94. Paratype, lo. 3164. Upper Palaeocene, sample
460-j, Sor Range, 8 miles east of Quetta.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 40

PLATE 41

Trachyleberis (Acanthocythereis) procapsus sp. nov.
Fics. 1, 3. Right and dorsal views, carapace male, x 94. Holotype, lo. 4360. Upper
Palaeocene, sample 460-i, Sor Range, 8 miles east of Quetta.
Fic. 4. Dorsal view, carapace female, x 94. Paratype, Ilo. 3164. Upper Palaeocene,
sample 460-j], Sor Range, 8 miles east of Quetta.

Trachyleberis (Acanthocythereis) usitata sp. nov.
Fic. 2. Right view, carapace male, x 94. Holotype, lo. 4362. Gorge Beds, sample 3111,
Rakhi Nala.

Fics. 5,7. Left and right views, carapace female, x 94. Paratype, Io. 3161. Gorge Beds,
sample 3111, Rakhi Nala.

Trachyleberis (Acanthocythereis) pedigaster sp. nov.
Fics. 6, 8. Right and left views, carapace, x 70. Holotype, Io. 4358. Lower Rakhi Gaj
Shales, sample 3671, Rakhi Nala.

Trachyleberis (Acanthocythereis) postcornis sp. nov.
Fic. 9. Left view, carapace male, x 94. Holotype, Io. 4361. Lower Chocolate Clays,
sample 3499, Rakhi Nala.
Fic. 10. Left view, carapace female, x 94. Paratype, lo. 3162. Lower Chocolate Clays,
sample 3499, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 41

PLATE 42
Trachyleberis (Acanthocythereis) postcornis sp. nov.
Itc. 1. Dorsal view, carapace male, x 94. Holotype, Io. 4361. Lower Chocolate Clays,
sample 3499, Rakhi Nala.
Fic. 2. Dorsal view, carapace female, x 94. Paratype, Io. 3162. Lower Chocolate Clays,
sample 3499, Rakhi Nala.
Fics. 7, 10. 7, Dorsal view of hinge, x 232. 10, Internal view to show radial pore canals,

x 178. Right valve female. Paratype, Io. 3163. Lower Chocolate Clays, sample 3498
Rakhi Nala.

Trachyleberis (Acanthocythereis) decoris sp. nov.
Fics. 3-5. Dorsal, left and right views, carapace male, x 94. Holotype, Io. 4359. Upper
Chocolate Clays (upper part), sample 3640, Rakhi Nala.
Fics. 6, 8, 9. Dorsal, left and right views, carapace female, x 94. Paratype, Io. 4310.
Upper Chocolate Clays (upper part), sample 3640, Rakhi Nala.

Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9 PLATE 42

° 2? ee

5
~

Om —

OMI gH &

Bull. Br. Mus. nat. Hist. (Geol.) Suppl. 9

——_—
rididae in the Palagocene and

fo)

f the Rakhi

N

a

. 4 SS
Table 1. Stratigraphic frequency of the ostracod family

m

3200
3403

3405 1a
3407

3409

3415

3416

3417

3418
3419

3420
3421

3422
3423
3424
3426
3428
3429

3432

3434

3435

3438
3440
3443
3444
3445
3446
3447

3448
3450
3451
3452
3453
3454
3456
3457
3458
3459,
3460
3461
3462
3463
3464
3465
3466,
3467

3470

Samples with total no. of ostracods
Species _|5

13672

3671

3147
3152
3153

~
a

3158
3159

3160
3161

3162

n
Ge

3174

3175
3176

3177
3178

3179
3180
3182
3183
3184
31
3186
3187
3188

3197,
3198

3199
3401

So
=
ise}

3404

3410

[473

Alocopocythere rupina sp. nov.

Buntonia devexa sp nov.

Hermanites cracens sp.nov.

Trachyleberis (Acanthocythereis) usitata sp. nov.
Buntonia sp A

Occultocythereis sp. A

Trachyleberis (Acanthocythereis) Pedigaster sp. nov.

Alocopocythere abstracta sp. nov.

Trachyleberis ( Trachyleberis) lobuculus sp. nov.
Gyrocythere parvicarinata sp.nov.
Phalcocythere sentosa sp.nov.

Occultocythereis peristicta sp.nov. morphotype A
Alocopocythere transcendens sp.nov
Occultocythereis spilota sp.nov.

Occultocythereis peristicta sp nov. morphotype B
Quadracythere (Hornibrookella) directa sp nov.
Occultocythereis peristicta sp.nov. morphotype C
Occultocythereis peristicta sp nov. morphotype D
Occultocyshereis peristicta sp nov. morphotype E
Anommatocythere laqueta sp. nov.
Echinocythereis ( Echinocythereis) elongata sp. nov.
Alocopocythere coarctata sp. nov
Alocopocythere longilinea sp. nov.
Stigmatocythere obliqua sp. nov.

Gyrocythere grandilaevis sp nov.
Phalcocythere dissenta sp. nov.

Shales

LRGS =Lower Rakhi Gaj

GB. = Gorge Beds

Ranikot

Palaeecene

1

_

wo

NiO) (O'S) RATE ODIs

NO OSTRACODS

20 25 18 37 6
42 -

38

wr

50
45

18

QB
'
i)
w

10 - = - - 211

Green and Nodular

85
9) 33|- - 8
Po) oo =

20

1 38 18 19

= 31 15°47) 2 1) = = -
Se hn 4 Js

Shales

= 220250 40 55

30 55 100
260i 72 7
= §

Uv 6

nNa a

425

alk,

3355 40 10 170 68 55 30

23

ih 1

Rubbly Limestones

13

40 90 175

ul

1

60 75 105 10

= 100131) =

= 2 4B

= S595 55> 9)
680 80 1 - - - — 18 300
25145 22 - -

i)
a
i
w
i)

4 i i
1 30 - - — 21013550 - 70

Shales with Alabaster

Gr 1H) VAY ZI

PAL.

{GB, LRGS Upper Rakhi Gaj Shales ——
RA ote

LOW ER

~

EQOCE WN E

J

35 48 35 24 17023080,30 1 480)

ja 4 A - enn +. ~- = .
t bab I Be a —~

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erm v4 rivootlvestie
es

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Pann ee Sm i
gene? _ *t
onion Aight a

ii
nyt

——

i &§
Pm ;

SRE eee eGo.) SUPP) 9 Table 2, Stratigraphic frequency of the ostracod family Trachyleberididae
in the Middle and Upper Eocene of the Rakhi Nala.

Samples with total no. of ostracods S=r284nnroSegtoxys ere
| el SANG) ot) dt Gai
1 Alocopocythere transcendens spnoy. eS Se 38}
2 Gyrocythere perfecta sp. nov. 5 15
3 Stigmatocythere portentum sp. nov. | 7
4 Trachyleberis (Acanthocythereis) postcornis sp. nov. 12 33
5 Anommatocythere confirmata sp. nov. 12 es ==s 5 9 6 i? 32
6 Echinocythereis (Scelidocythereis) rasilis sp. nov. 12 ao oe eoa ose 3 So 2
7 Hermanites scopus sp.nov. 1 SP Se Gs Gh lene 02
8 Occultocythereis indistincta sp. nov. 2 BS ee eS Se ee ee ea ee
9 Quadracythere (Hornibrookella) arcana(Lubimova & Guha) 2
10 Costa (Paracosta) compitalis sp. nov. fA W 4
11 Trachyleberis (Acanthocythereis) decoris sp nov. a Ogre B2 2W6 5 -=-se-=se e552 5 56
12 Actinocythereis ? quasibathonica sp.nov. fo) | 3 & 2 2 3
13. Trachyleberis (Trachyleberis) bimammillata sp. nov. y S) 4 252 6
14 Gyrocythere exaggerata sp. nov. Ki < 7 |
15 Hermanites palmatus sp. nov. @e a $3 1 1 1
16 Echinocythereis (Scelidocythereis) multibullata sp. nov. (= FE 4 = = 8 1
17 Stigmatocythere lumaria sp. nov. morphotype A w Vp) QPooR@Pht G> Feo Gh 6 1 BeBe = — 1 = = Bs
18 Costa (Paracosta) disintegrata sp.nov. 2) fe) 2 2
19 Patagonacythere?nidulus sp. nov. ro) S 145 - - - = 188 $604 628s 2-55 §85 22
20 Alocopocythere transversa sp.nov.morphotype A Zz z 3 9 § = = 12 2
21 Alocopocythere transversa sp.nov. morphotype B 358 see =o g
22 Alocopocythere transversa sp. nov. morphotype C Bt {=> bo so ji] Bez ||
23 Alocopocythere transversa sp. nov. morphotype D 74s so 22 if 3 WY
24 Alocopocythere transversa sp. nov. morphotype E 1 - 2 -- 11 120 21 63
25 Stigmatocythere lumaria sp.nov. morphotype B he a= 2 - 1412461 47
26 Alocopocythere transversa sp.nov morphotype F 1 -- 8 30- 35 2
27 Alocopocythere radiata sp. nov. Us
7 NW 2

28 Costa (Paracosta) declivis sp. nov.
W_M.B = White Marl Band

Upper Chocolate Clays Lower part Upper Chocolate Clays Upper part Pellatispira Beds
_—e y= = pper ee a
LOWER KIR T H AR (<ekirthar sensu stricto of Eames ) ale UPPER KIRTHAR (Tapti of Eames)

Lee, _ ts
=

LEC = Lower Chocolate Clays

NED Ds EE OnGsEaNie cE WPRP ER le @Cle IN le

PL= Platy Limestone

, Verve. © =

34 von Qazines) 206 {atsyarlf strosa)

‘D - . wONy sete fy: oft |
A " vn ¥ Piiam (aiviectf wobibbs2 )
r : GR tha vorlad niasea
| Wert ad af snike ink mmarity,

f brid Apvoriidutd anes. (olinlacndinot)’s stot

te

von .g2 2ilntiame> (alzoz io an. ~
& lve ge Fe4eb RidteAnSodins citedglyiaat 5.
| a re von ge | A> eete Ae write
Lyon oe Dipilidamomme (Ay

«= F
ia

| 3
| Woe OFD! UdLM (2isneMysobisse ) ane
. A ecviona TOT) VOR 2 Drm ack d
ie 4 a >. vero2 oimgeinialb tot
i | “von G2 aE FE
= A sgyvloriqom von de Ovary Port —
|) Be 8 sqydordnom voraz oerevetiont, sad} ie
.2 agyior qo Von 92 peieventht: : )
; 7 sqyoAmon vor Ge pemvenayw orsetiy
3 20yredeRomh von. Ga darqvenmn? oorliyo
2 aq orgie Von a2 onorrwi ees
7 sqvorigiomn \oh.G2 Se1svenod’ sitet
a. | Yon .42 dipibm wudivstad

7 . von qe mila (ofedore a
ie | sede + lieper _fayecolai¢ Via Mab
———e e , 7 - om Sawer

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a ae
43. 4

v=

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©

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Bull. By. Mus. nat. Hist. (Geol.) Suppl. 9

Table 3. Stratigraphic frequency of the ostracod family Trachyleberididae in the Eocene of the Zao River.

Samples with total no. of ostracods
Species
Anommatocythere laqueta sp, nov.

Phalcocythere dissenta sp. nov.

Stigmatocythere obliqua sp. nov.

Alocopocythere transcendens sp. nov.

Gyrocythere mitigata sp. nov.

Hermanites palmatus sp. nov.

Trachyleberis (Acanthocythereis)postcornis sp.nov
Anommatocythere confirmata sp.nov.
Echinocythereis (Scelidocythereis) rasilis sp. nov.
Gyrocythere exaggerata sp nov.

Hermanites scopus sp nov

Quadracythere (Hornibrookella) sp.A.
Stigmatocythere calia sp. nov.

Actinocythereis ? quasibathonica sp. nov.
Trachyleberis (Trachyleberis) bimammillata sp.nov.
Patagonacythere?nidulus sp.nov.

Echinocythereis (Scelidocythereis) multibullata sp.nov.

Stigmatocythere delineata sp.nov

Trachylebens (Acanthocythereis) decoris sp. nov.
Alocopocythere transversa sp.nov.morphotype A.
Alocopocythere transversa sp nov. morphotype B.
Stigmatocythere lumaria sp.nov. morphotype A
Alocopocythere transversa sp.nov.morphotype E.
Alocopocythere transversa sp. nov. morphotype C.
Bradleya ? voraginosa sp.nov.

Echinocythereis (Scelidocythereis) sparsa sp.nov.
Phalcocythere spinosa sp. nov.

Quadracythere (Hornibrookella) subquadra sp.nov.
Alocopocythere transversa sp.nov. morphotype D.
Alocopocythere transversa sp.nov. morphotype F
Alocopocythere radiata sp. nov.

WM.B = White Marl Band

LCC. = Lower Chocolate Clays

Paiie = Platy Limestone

S WA_= Shales with Alabaster

14 2 50 35

81
ER ae
fo a = 2
3.1 16 24 3
6 4 ti 3 =
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i 1
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fo} (eo)
[e) (e)
a

8
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(=Kirthar s.s.ofEames) UPPER KIRTHAR (- Tapti of Eames)

UPPER] iE OlG E INSE

GHAZI LOWER K | R T
E

H
Lr EOC MIDDLE (0)

AR
(C fe

56 1

16 —

7

==5 § Ai = il WS hea
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Qo = =O S84
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a ce en
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23 10

4 10-

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Bull, Br. Mus. nat. Hist. (Geol.) Suppl. 9

Rakhi Nala Stratigraphic Section, measured by S.L.Rieb and sampled by D.D.Bayliss. 1957.

SAMPLE

NUMBERS

———r
aoa nd

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‘n0D

LITHOLOGY

SCALE

THICKNESS

KHIRTHAR

FORMATION

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RANIKOT

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LEGEND

Sond

Sondstone

Conglomeratic sandstone

- =} Silt

Siltstone

. .| Siltstreaks

= Mudstone

= | Shole

— | Argillaceous formation

= | Shole or clay lenses

~ ~| Marl

Coquina or fossiliferous limestone
Argillaceous limestone
Calcareous streaks or lenses

Gypsum

ABBREVIATIONS
Cov. Covered
S? Siwalik

Pab sst.

Pab Sandstone

SCALE: Feet and Metres.
0:5 inches = 100 feet,
42 cms. = 100 metres.

N.B. Sample numbers for every fifth

sample unless otherwise indicated.

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A

A LIST OF SUPPLEMENTS
TO THE GEOLOGICAL SERIES
OF THE BULLETIN OF
THE BRITISH MUSEUM (NATURAL HISTORY)

1. Cox, L. E, Jurassic Bivalvia and Gastropoda from Tanganyika and Kenya.
Pp. 213; 30 Plates; 2 Text-figures. 1965. {6.

2. Et-Naccar, Z. R. Stratigraphy and Planktonic Foraminifera of the Upper
Cretaceous—Lower Tertiary Succession in the Esna-Idfu Region, Nile Valley,
Egypt, U.A.R. Pp. 291; 23 Plates; 18 Text-figures. 1966. £10.

3. Davey, R. J., Downie, C., SARGEANT, W. A. S. & WitLiaMs, G. L. Studies on
Mesozoic and Cainozoic Dinoflagellate Cysts. Pp. 248; 28 Plates, 64 Text-
figures. 1966. {7.

3. APPENDIX. DAVEY, R. J., DowniE, C., SARGEANT, W. A. S. & Witiiams, G. L.
Appendix to Studies on Mesozoic and Cainozoic Dinoflagellate Cysts. Pp. 24.
1969. 16s. és

4. Extiott, G. F. Permian to Palaeocene Calcareous Algae (Dasycladaceae) of the — t%
Middle East. Pp. 111; 24 Plates, 17 Text-figures. 1968. {£5 2s. 6d.

5. Ruopes, F.H.T., Austin, R.L.& Druce, E.C. British Avonian (Carboniferous)
Conodont faunas, and their value in local and continental correlation. Pp. 315;
31 Plates, 92 Text-figures. 1969. {11. .

6. Cuitps, A. Upper Jurassic Rhynchonellid Brachiopods from Northwestern
Europe. Pp. 119; 12 Plates, 40 Text-figures. 1969. {£4 I5s. .

7. Goopy,P.C. The relationships of certain Upper Cretaceous Teleosts with special —_
reference to the Myctophoids Pp. 255; 102 Text-figures. 1969. {£6 Ios.

8. OwEN, H. G. Middle Albian Stratigraphy in the aaa Basin. Pp.164;
3 Plates; 52 Text-figures. 1971. {6.

PRINTEDINGREATBRITAINBYALDEN & MOWBRAYLTDATTHEALDEN PRESS,OXFORD |

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