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BUEEETINS
OF
AMERICAN
ave ONTOLOGY
VOEP EX Xl
1977-1978
Paleontological Research Institution
Ithaca, New York 14850
U::S: A:
IN MEMORIAM
Mrs. Kitti Westfall
1930-1977
Waldo Schmitt
1887-1977
John R. Sandidge
1897-1976
Axel A. Olsson
1889-1977
CONTENTS OF VOLUME LXXiII
Bulletin No. Pages
296. The Disparid Inadunate Superfamilies Homo-
crinacea and Cincinnaticrinacea (Echinoder-
mata:Crinoidea), Ordovician-Silurian, North
America
Byam arn ands He slas Strimip eyes sssceceeees 1-138
297. Some Paleocene and Eocene Barnacles (Cir-
ripedia) of Alabama
By Normans WeISDOLG \.c25.4..c0:.4.ceceecssaccesosese 139-166
298. The Archaediscidae of the Fraileys Facies (Mis-
sissippian) of Central Kentucky
By R. G. Browne, J. W. Baxter, and T. G.
ROW ONES ype ote reco coocse endian ceasten feces ees: 167-228
299. Scalpellid Barnacles (Cirripedia) of Florida and
of Surrounding Waters
By Norman’ Fe WEISDOTO) c.c...sccecc.<ssveecese-veceeses 229-312
300. Primary Types in the Stanford Paleontological
Type Collection
Bye sudithy Merry SMith) 2 .32.02054-ccksooantsesteacess te 313-552
Plates
19-21
22-25
INDEX
No separate index is included in the volume. Each number is
indexed separately. Contents of the volume are listed in the begin-
ning of the volume.
—\/
rs
BULLETINS tee!
OF LARVARD
AMERICAN |
PALEONTOLOGY
(Founded 1895)
Vol. 72
No. 296
THE DISPARID INADUNATE SUPERFAMILIES
HOMOCRINACEA AND CINCINNATICRINACEA
(ECHINODERMATA: CRINOIDEA),
ORDOVICIAN-SILURIAN, NORTH AMERICA
By
J. Warn anv H. L. Strimpte
1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
PALEONTOLOGICAL RESEARCH INSTITUTION
1976-1979
PRESIDEN Tocca eae seat ee pee oo eee oe eee ee ae 2h De, HArROLp E. VOKES
WIGE-E RESIDEN pies stores ee ee Ee ee i ee DUANE O. LERoy
SS RCR TARY te se erect ee ee I eee ee ee PHILIP C. WAKELEY
SURE ASURER 62 ae ee ne Wek eee ae oe ERNESTINE Q. WRIGHT
PURE CTOR eee ce eS See ee ee eee KATHERINE V. W. PALMER
ASSISTANT SECRETARY, ASSISTANT TREASURER ...........--ccse------eceeee-a-- REBECCA S. HARRIS
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REPRESENTATIVE AAAS CoUNCIL .......... Sie ry Sh ee ee Ste RicHarpD G. Oscoop, Jr.
Trustees
RutH G. Browne (1976-1979) KATHERINE V. W. PALMER (Life)
KENNETH E. CASTER (1975-1978) JoHN PojeTA, Jr. (1975-1978)
Joun L. CIsNE (1976-1977) K. NORMAN SACHS, JR. (1974-1977)
REBECCA S. Harris (Life) DANIEL B. Sass (1974-1977)
MarcareT B. HeEroy (1975-1978) Harotp E. Vokes (1975-1978)
DUANE O. LERoy (1974-1977) Puitie C. WAKELEY (1976-1979)
WiiiaAM A. OLIVER, JR. (1976-1979) ERNESTINE Q. WRIGHT (1976-1979)
AxeEL A. OLsson (Life)
BULLETINS OF AMERICAN PALEONTOLOGY
and
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Advisory Board
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AMERICAN
Pale ONTOEOGY
(Founded 1895)
Vol. 72
No. 296
THE DISPARID INADUNATE SUPERFAMILIES
HOMOCRINACEA AND CINCINNATICRINACEA
(ECHINODERMATA: CRINOIDEA),
ORDOVICIAN-SILURIAN, NORTH AMERICA
By
J. Warn anv H. L. Stripe
May 31, 1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
Library of Congress Card Number: 77-78618
Printed in the United States of America
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CONTENTS
Page
PNDSITAG | ses jeeter ence eer eer ae el lates Ne ey le a pe ea ye pe ee ir Mire 5
STECSET © GLUT GEL i eer nee ar rece ee ce a rere noe aire ere eres 5
PNG KETT CWV)] © CLT © Tt S eee asec ne ee a ee es Se ee a ee ee ee 7
TEGTD@RTIOP OED 1 erence ade OB ee ele el a arn aR A 7
SUTRAS TY Na a ee re ee ett ee ae eee 8
Rock-stratizraphic units in’ the Cincinnati areal 22-2 eee 17
sime-straticraphic units) im! the, Cimeimmnati areal oes acecere cee eee cree 20
@reanisms as indices of Cincinnatian ‘stages: 22 cc ec eececce cece 23
CONGO NTs eee eS ee ee te tae ee 23
(CROSS eke cere meee te ae ge A aE Pe ee Zep a a ea ei ea ae 24
BA CHI O POG Speen eee eee ce ne OTe Re RN OE ere ree ae eee 24
RROD. etter tee tee tap tee Se tt Nas A pk ee eee ai pec a A 24+
Mbnilobite ss eee. eae Sete pel fot hl ich te ia ee ete PRO el ona 24+
CORTE ae te ae eR eA et Ge 25
SAV SLEMatl Cm all COMCOLO Lice te se ee eee ee eee eee ee ee 25
Crinorde age linia GUA ae een ek leet eee ee ee ee 25
DD) 2S ya Ty 1 ca ere eee ere a KA. Sah Pe eee LUN ee eee ee eee 26
CinGiNM A tier iT St a eee Roe Os ere heen cr 35
TABLE
1. Classification of Cincinnaticrinacea and Homocrinacea ....between pp. 22, 23
TEXT-FIGURES
i mCities and ‘states referred to) iN text 2.5. 22-t-cesesccseceacnececcecncnset cos ee
2. Correlation of Ordovician and Silurian time-stratigraphic standards ....
3. Orton’s rock-stratigraphic classification, Cincinnati Group ......................
4 Cincinnatian: stratigraphy: 2. 2-s.cc.:0--. cesses io ae
Seronape Jof (Greek: Vases" 2iicccccccdz2cecescstcccses covscdvecceete sansa. stt eee
Geaely pessotearm branching... 2.cctcccecet cco toseoccsecseetsteeee ee ee eee
jeeelypesvof heterotomous) arm) branching, cee cee
SaNGINCINMALICTINUS VATIVIAChiQlts seco. ee
9. General basal plate shapes, cincinnaticrinaceans and homocrinaceans ..
10s Localities Of Cx Varturachialis, ere re eee
iipeOntogeny) ofG. varibrackialus Column eee ee
12. Ranges of cincinnaticrinacean and homocrinacean species .................-..----
13. Ontogenetic change in life habit of C. varibrachialus 0.000000 .0c0ceceee--e
14. Phylogeny of Cincinnaticrinacea and Homocrinacea ...................-.--...--------
Wier Usovomiocrintts. Lert ooo o2 5 oo cocosccostasacenseasesteentnnnt eee
16. Ulrich’s (1882) abnormal specimens of Heterocrinus oehanus ...............
Ae MORO Crimes DraUns 22.02 coc nccdec. cates csesmcsten nce eee
Sem ALOPOCTINUS -DTiS CUS, 5.
DOO MLO CHINUS PATUUS. (oc. oo ccassceaccsectoctecacsentia thinset ee eee
20. Exploded diagram of Ectenocrinus simplex. ..............--------000----0--00eeeeneeneenees
ZipmExplodedsdiagramiot Apodasmocrinus =
OMAP OAASHUO CUATEUS. | oo 2a 2 omen c codececnce led uaece tea se ee
23h bexocrinusleplon Rew snes Be Wis Sh aie ee a ee
DAE SPP CAULOCTITUG, Ly DUS. eh ease cscs ecccnscetceew te eee a
Ose dedalocrinussbellewtllensis: <-.ee cu eae ae eee
THE DISPARID INADUNATE SUPERFAMILIES
HOMOCRINACEA AND CINCINNATICRINACEA
(ECHINODERMATA: CRINOIDEA),
ORDOVICIAN-SILURIAN, NORTH AMERICA
J. Warn! and H. L. Strimple?
ABSTRACT
The discovery that Heterocrinus heterodactylus Hall, type species of
Heterocrinus Hall, 1847, is unrecognizable necessitates new names for crinoid
taxa formerly placed in Heterocrinus and in the superfamily Heterocrinacea.
The new genus Cincinnaticrinus is erected to accommodate the new species C.
varibrachialus. The new superfamily Cincinnaticrinacea essentially replaces the
Heterocrinacea (nom. trans. Ubaghs, 1953, ex Heterocrinidae Zittel, 1879).
Revision of this superfamily and the related Homocrinacea has enabled elimina-
tion of many superfluous taxa, the establishment of numerous lectotypes and
lectoparatypes, more accurate geographic and stratigraphic ranges for the re-
maining species, and consistent diagnoses and descriptions of well-established
taxa. The new family Cincinnaticrinidae (= Heterocrinidae Zittel, 1879) is
divided into two new subfamilies, the Cincinnaticrininae (including C. vari-
brachialus, n. sp., C. pentagonus (Ulrich), Dystactocrinus constrictus (Hall),
Isotomocrinus tenuis (Billings), J. minutus Kolata, Ohiocrinus laxus (Hall), and
O. brauni Ulrich, and Atopocrininae (A. priscus Lane). The family Homocrini-
dae Kirk, 1914, is also divided into two new subfamilies, the Homocrininae (for
Homocrinus parvus (Hall), Ectenocrinus simplex (Hall), E. geniculatus (UI-
rich), Apodasmocrinus punctatus (Brower and Veinus), A. daubei, n.g., n. sp.,
Ibexocrinus lepton Lane, and Sygcaulocrinus typus Ulrich) and Daedalocrininae
[containing only Daedalocrinus bellevillensis (Billings) ]. Possible phylogenies
and the paleoecology of the included species are discussed; it is concluded that
crinoids with lichenocrinid-type bases were probably effectively eleutherozoic.
INTRODUCTION
Current concepts of the disparid inadunate crinoid families
Heterocrinidae and Homocrinidae (elevated to superfamilies by
Ubaghs, 1953) date essentially from Ulrich (1925). The forthcoming
crinoid section of the Treatise on Invertebrate Paleontology will
include few changes: the Treatise will incorporate two new genera,
Atopocrinus (a heterocrinid) and Ibexocrinus (a homocrinid),
described by Lane (1970) and placed by him in those families; and
the Treatise will characterize the genus Heterocrinus as having iso-
tomous rather than heterotomous branching, as it was defined by UI-
rich (1925, p. 84) and others (Wachsmuth and Springer, 1886, p.
207; Wachsmuth, 1900, p. 152; Grabau and Shimer, 1910, p. 502;
Springer, 1911, p. 27; Springer, 1913, p. 212; Moore and Laudon,
1944, p. 149; and Warn, 1973, p. 12). Ulrich (1925) apparently
based his study on small numbers of specimens (for some species, on
from one to a few). Modern treatment, with examination of large
numbers of specimens and attention to intraspecific variation, ap-
pears necessary.
1. Western Exploration Division, Shell Oil Company, Houston, Texas. 2. Geology
Department, The University of Iowa, lowa City, Iowa.
6 BuLLeETIN 296
Additionally major nomenclatural changes are needed because
Heterocrinus heterodactylus (type species of Heterocrinus) is un-
recognizable and the name must be restricted to Hall’s (1847) type
specimens (from New York strata). Thus, new names must be given
to taxa from strata in and around Cincinnati, Ohio, formerly at-
tributed to Heterocrinus and H. heterodactylus (the authors choose
the new names Cincinnaticrinus and C. varibrachialus for these
taxa); and, because Heterocrinus is the type genus of the familial
taxa Heterocrinidae and Heterocrinacea, new names must also be ap-
plied to these taxa (designated respectively, Cincinnaticrinidae and
Cincinnaticrinacea herein).
Study of Cincinnaticrinacea and Homocrinacea has been par-
ticularly facilitated by taking advantage of the large collections of
these crinoids at the University of Cincinnati. Type species of six
(Heterocrinus, Atyphocrinus, Dystactocrinus, Ohiocrinus, Ecteno-
crinus, and Drymocrinus) of the 11 genera placed by Ulrich (1925)
in the Heterocrinidae and Homocrinidae have been reported to occur
in Cincinnatian strata in and around Cincinnati, Ohio, and they are.
well represented in existing collections. Of the remaining five genera,
two were first described from the Hull Limestone, Kirkfield, On-
tario, (Isotomocrinus and Daedalocrinus), one (Sygcaulocrinus)
from the Fort Atkinson Member of the Maquoketa Formation of
Iowa, one (Columbicrinus) from the Lebanon Limestone of Ten-
nessee, and one (Homocrinus) from the Rochester Shale of New
York. The Kopf Collection at Cincinnati is one of the finest North
American echinoderm collections and contains hundreds of Kirkfield
crinoids. Thus, large numbers of cincinnaticrinids and homocrinids
are housed in the University of Cincinnati Geology Museum and in
other Cincinnati area museums. Finally, large pockets of Cinctn-
naticrinus varibrachialus (type species of Cincinnaticrinus) and
Ectenocrinus simplex (type species of Ectenocrinus) have recently
been discovered in the area.
Most of this work has been drawn from Warn’s Ph.D. disserta-
tion (University of Cincinnati, 1974) entitled “The disparid crinoid
superfamilies Homocrinacea (Ord.-Sil.) and Cincinnaticrinacea
(Ord.)”. Incorporation of subsequently published data and interpre-
tations have been made by Strimple. The authors, however, concur
and share responsibility for the entire study.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 7
ACKNOWLEDGMENTS
The authors thank Kenneth E. Caster, University of Cincin-
nati, who allowed the senior author free use of his personal library
and gave advice on preparation of the manuscript. Thanks are also
due R. A. Davis, who offered suggestions on taxonomy and nomen-
clature; E. A. Dalvé, who draughted Text-figures 2 and 13; K. L.
Derstler, J. F. Gastright, and W. H. White for donation, or loan,
of specimens for study; and the following individuals and their re-
spective institutions for loan of specimens: R. L. Batten and N. D.
Newell, American Museum of Natural History; R. P. S. Jefferies and
H. G. Owen, British Museum (Natural History); E. S. Richardson,
Field Museum of Natural History; T. E. Bolton, Geological Survey
of Canada; J. H. Marak and J. K. Pope, Department of Geology,
Miami University; B. H. Kummel, Museum of Comparative Zoology,
Harvard University; B. M. Bell, New York State Museum; S. M.
Bergstrom, Orton Museum, Ohio State University; J. S. Lawless and
K. A. Waage, Peabody Museum of Natural History, Yale University;
John Monteith, Royal Ontario Museum; F. J. Collier, P. M. Kier,
and Thomas Phelan, United States National Museum; and R. A.
Davis, University of Cincinnati Geology Museum. We also thank
Mrs. Olive Daube, Sam Daube, and Jim Manton, manager of the
Daube Ranch Company, for allowing access to the exposure from
which Apodasmocrinus daubei, n.g., n. sp. is described. We are in-
debted to Mr. T. J. Frest, The University of Iowa, for assistance in
incorporating current data from published and unpublished studies
as well as for editing assistance. The final typescript was prepared
by Mrs. Jan Thein, The University of Iowa. Finally, a NDEA
summer research grant (1972) to the senior author made possible,
in part, necessary travel and purchase of supplies.
REPOSITORIES
Specimens referred to in this work are listed by catalogue num-
bers with the respository names abbreviated as follows:
AMNH American Museum of Natural History, New York, New York
BM(NH) British Museum (Natural History), London, England
CFM Field Museum of Natural History, Chicago, Illinois, (numbers pre-
ceded by UC denote specimens in the University of Chicago
Walker Museum Collection)
GSC Geological Survey of Canada, Ottawa, Ontario
HM Hunterian Museum (Geology), The University, Glasgow, Scotland
8 BULLETIN 296
MCZ Museum of Comparative Zoology, Harvard University, Cambridge,
Massachusetts
MU Geology Museum, Department of Geology, Miami University, Ox-
ford, Ohio
NYSM New York State Museum, Albany, New York
OM Orton Museum, Department of Geology, Ohio State University,
Columbus, Ohio
ROM Royal Ontario Museum, Toronto, Ontario
SUI Geology Department, The University of Iowa, Iowa City, Iowa
UCGM University of Cincinnati Geology Museum, Department of Geology,
Cincinnati, Ohio, (numbers preceded by K denote specimens in
the Kopf Collection)
UI Geology Department, University of Illinois, Urbana, Illinois
UM Geology Department, University of Minnesota, Minneapolis, Minne-
sota
USNM National Museum of Natural History, Smithsonian Institution,
Washington, D.C., (numbers preceded by S. denote specimens in
the Springer Collection)
UMMP University of Michigan, Museum of Paleontology, Ann Arbor,
Michigan
YMP Peabody Museum of Natural History, Yale University, New Haven,
Connecticut
STRATIGRAPHY
Cincinnaticrinacea and Homocrinacea range from Whiterockian
to Niagaran strata from western North America (Utah and
Wyoming) to eastern North America (New York and Quebec).
Various members of the two superfamilies occur in the Kanosh
Shale of Utah (Whiterockian); the Decorah Shale, St. Paul, Minne-
sota (Kirkfieldian); the Hull beds, Kirkfield, Ontario (Kirkfield-
ian); the Hull (Kirkfieldian?), Sherman Fall (Shermanian? ), and
Coburg (Edenian?) beds of Ottawa and Montreal; the Whetstone
Gulf Formation of northwestern New York (Edenian? ); the Shegui-
andah Formation of the Manitoulin Island area of Canada (Eden-
ian); the Edenian and Maysvillian (upper) portion of the Martins-
burg Formation of Maryland and southern Pennsylvania; the
Maquoketa Formation of Iowa (Richmondian?); and the Roches-
ter Shale near Lockport, New York (Niagaran),— Text-figure 1 is
a map that shows the location of cities and towns referred to in the
text. No attempt, beyond Text-figure 2 and references under the
respective occurrences of taxa, will be made to discuss these strata.
However, seven of the thirteen cincinnaticrinacean ard homocrina-
cean species recognized here occur in Cincinnatian strata in the
southwestern Ohio-southeastern Indiana-northern Kentucky area,
and Cincinnatian stratigraphy and stratigraphic nomenclature de-
serves attention.
9
WARN AND STRIMPLE
ORDOVICIAN-SILURIAN CRINOIDS
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Text-fig. 2. Correlation of Ordovician and Silurian time-stratigraphic
standards of North America and Europe and pertinent rock units of geographic
areas where crinoids studied herein occur. Areas with diagonal lines = strata
representing that time missing, blank areas = strata unexposed or considered
not pertinent by the authors. T = Tremadoc, and A = Arenig. Although all
rock units are time-transgressive, only those that have been demonstrated to be
so are so illustrated; others are bounded by straight lines. (From Twenhofel,
et al., 1954, with modification from Weiss and Sweet, 1964; Peck, 1966; Sweet
and Bergstrém, 1971.)
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE tl
HISTORICAL SURVEY OF CINCINNATIAN STRATIGRAPHY
In 1829, Vanuxem (p. 256) correlated Cincinnatian rocks of
Ohio and Kentucky (and perhaps Champlainian strata of Ken-
tucky) with strata at Trenton Falls, New York. Hall (1842, p. 61)
proposed age equivalence of shales at Newport, Kentucky, (directly
across the Ohio River from Cincinnati) with the New York Utica
and the underlying limestone exposed in the Ohio River only at low-
water with the Trenton of New York. In 1843, Hall referred strata
at Cincinnati to the Hudson River Group. The term “Hudson River”
was used by Mather, Emmons, Vanuxem, and other early New York
geologists mainly for rocks of Late Ordovician age exposed in the
Hudson River Valley, New York. Hall also correlated a body of
underlying strata (containing Triarthrus eatoni — T. becki of older
literature) with the New York Utica and the lowermost strata ex-
posed at Cincinnati with the New York Trenton (with reservation).
Thus, from 1843 to 1865 the name Hudson River Group was widely
used for strata at Cincinnati.
In 1859, Mather (p. 6) used the name “Cincinnati limestone” in
passing for strata at Cincinnati. First usage of Cincinnati as a strati-
graphic term, however, is usually attributed to Meek and Worthen
(1865, p. 155, and in reports on the geology of [linois — Worthen,
1866; Meek and Worthen, 1868), who suggested that the name
Cincinnati Group be substituted for Hudson River Group, because
the strata at Cincinnati and the New York Hudson River Group
are of different age. Orton (1873, p. 369) proposed that it is “proba-
ble that the lowermost beds of Cincinnati are the proper equivalent
of the Utica Slate [of New York].” Orton (1873) divided the Cin-
cinnati Group into five lithic units (in ascending order): Point
Pleasant beds; River Quarry beds; Middle or Eden shales; Hill
Quarry beds; and Lebanon beds.
The River Quarry beds, Eden shales, and Hill Quarry beds, all
named for strata exposed in the immediate vicinity of Cincinnati,
were lumped together by Orton as the Cincinnati beds. The name
Point Pleasant was used for strata exposed some distance upriver
from Cincinnati (the town Point Pleasant is situated about 25 miles,
about 40 km, upriver from downtown Cincinnati), while the name
Lebanon was applied to strata outcropping on top of the Ohio River
hills nowhere closer than 20 miles (about 32 km) from Cincinnati.
Orton defined the units as follows (refer to Text-fig. 3):
12 BuLLETIN 296
description
predominatly limestones (shales: limestone
somewhat higher than 1:1 in the lower portion
and somewhat lower than 1:1 in the higher
portion) lying between the highest stratum of
the Cincinnati hills and the lowest Silurian
(Brassfield) beds and outcropping no closer
than 20 miles (32 km) from Cincinnati
shale and limestone about equal (ratios of
less than 5 or 6:1 and approaching 1:1); out-
cropping just below the tops of the hills
at Cincinnati; extensively quarried
Eden
shales |(76m)
predominatly shale (shale to limestone ratios
of about 4:1 and up to 10:1); named for ex-
posures in Eden Park in Cincinnati
Cincinnati group
Cincinnati beds
4-8" (10-20 cm) thick limestones, commonly
with rippled surfaces, made up of crinoid
parts; sh:ls = 4:1; quarried in Cincinnati
Point
Pleasant
beds
thick (16-18''--41-46 cm), barren limestones
and shales outcropping in the north bank
of the Ohio River at Point Pleasant, Ohio
Text-fig. 3. Orton’s (1873) rock-stratigraphic classification of the Cincin-
nati Group. All ratios are shale to limestone.
1)
2
—
3)
4)
5)
6)
7)
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 13
Cincinnati Group — 750 to 800 feet (about 230 to 245 m) of
alternating beds of blue “clay” (shale or mudstone) and blue-
gray limestone outcropping in and around Cincinnati, Ohio, and
including the Point Pleasant beds, Cincinnati beds, and Lebanon
beds;
Point Pleasant beds — lowest 50 feet (about 15 m) of the Cin-
cinnati Group with lighter-colored, essentially barren limestones
and shales and thicker (16 to 18 inches — about 41 to 46 cm)
limestones than overlying beds and outcropping in the north
bank of the Ohio River about 25 miles (about 40 km) east of
Cincinnati;
Cincinnati beds — 425 to 450 feet (about 130 to 137 m) of the
Cincinnati Group beginning at low-water of the Ohio River in
Cincinnati and extending to the tops of the hills, having a shale
to limestone ratio of at least 5:3, and including the River
Quarry beds, Eden shales, and Hill Quarry beds;
River Quarry beds — 50 feet (about 15 m) of firm and compact
limestones (commonly with rippled surfaces) of about 4 to 8
inches (about 10 to 20 cm) thickness (but sometimes up to 2
feet — about 60 cm), made up almost entirely of crinoid col-
umns alternating with thicker shales, with a shale to limestone
ratio of 4:1 and quarried in the Cincinnati area;
Eden shales — 250 feet (about 76 m) of predominantly shale
(with a shale to limestone ratio of at least 4:1 and as high as
10:1) named for exposures in Eden Park;
Hill Quarry beds — 125 to 150 feet (about 38 to 46 m) of more
limy rock (with shale to limestone ratios of less than 5 or 6:1
and approaching 1:1) outcropping just below the tops of the
hills at Cincinnati and extensively quarried;
Lebanon beds — about 300 feet (about 92 m) of predominantly
limy rock (shale to limestone somewhat higher than 1:1 in the
lower portion and somewhat lower than 1:1 in the higher por-
tion) lying between the highest stratum of the Cincinnati hills
and the lowermost Silurian beds and outcropping no closer than
20 miles (about 32 km) from Cincinnati (with good exposures
at Madison and Richmond, Indiana, and Oxford and Lebanon,
Ohio. )
14 BULLETIN 296
Orton (1873) represents the first good lithostratigraphic study of
Cincinnatian strata at Cincinnati (for much of the classical period
of American geology, lithologies at Cincinnati were overshadowed
by the abundance of fossils in the strata).
U. P. James (1879) reconsidered correlation of Cincinnati and
New York strata. He reported that of 500 species in strata at Cin-
cinnati only about 100 occur in the Trenton, Utica, and Hudson
River of New York; of that 100, 65 are confined to the Trenton,
18 to Utica and Hudson River, and 17 are shared by all three. James
concluded that Trenton would be a better designation than pre-
viously used Hudson River but opted for Cincinnati Group because
of the obvious faunal dissimilarity of local strata to that of New
York.
In 1881, S. A. Miller (pp. 268-269; 283-287) presented a new
correlation of Cincinnati and New York strata: the River Quarry
beds with the upper part of the Trenton Group, the Eden shales
with the Utica Group; and the Hill Quarry and Lebanon beds with
the Hudson River Group. The names Trenton, Utica, and Hudson
River unfortunately came to be used in place of Orton’s lithic names.
In 1891, J. F. James recognized a problem that has only recently
(Weiss et al., 1965, pp. 18-19) been resolved. James compared the
beds exposed at Point Pleasant, Ohio (= Orton’s Point Pleasant
beds), with those exposed during low-water in the Ohio River at
Ludlow, Kentucky (== Orton’s River Quarry beds), and found no
difference. This is the earliest implication that Orton (1873) had
given different names to two bodies of rock (which Orton thought
were distinct) that are now known to be portions of one unit, the
Point Pleasant Formation.
Winchell and Ulrich (1897, pp. ci-cv) used the term “Cincin-
nati Period” for rocks occupying a position between Trenton (in-
cluding Point Pleasant beds) and the Silurian Brassfield Formation
and rejected associations of the Hill Quarry and Lebanon beds with
the New York Hudson River Group. Rather, they correlated the
Hill Quarry beds with the Lorraine Group of New York and Ontario.
They then replaced Hudson River with Lorraine, and, because
Lebanon was preoccupied (Safford, 1851, pp. 353-355, had used
Lebanon for part of the Stones River Group in Tennessee), re-
placed Lebanon with Richmond. Thus, the names Trenton, Utica,
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 15
Lorraine, and Richmond came to be used for units at Cincinnati
that were essentially equal to Orton’s Point Pleasant beds (and
River Quarry beds), Eden shales, Hill Quarry beds, and Lebanon
beds. Clarke and Schuchert (1899, pp. 876-877) dropped usage of
Hudson River even for New York strata in favor of Cincinnatian,
with the divisions Utica (= Orton’s Eden shales), Lorraine (=
Orton’s Hill Quarry beds), and Richmond (= Orton’s Lebanon
beds) for the North American Upper Ordovician, and the name
Hudson River finally ceased to be applied to strata at Cincinnati.
Nickles (1902, pp. 56-98) equated Orton’s Point Pleasant beds
and River Quarry beds and subdivided the Utica, Lorraine, and
Richmond at Cincinnati into a number of faunal zones (mainly on
the basis of maximum abundance of various species of brachiopods
and Bryozoa). Foerste (1905) discarded New York nomenclature
altogether for use in the Cincinnati area and divided strata at Cin-
cinnati into Point Pleasant beds (= Orton’s Point Pleasant and
River Quarry beds), Eden (== Orton’s Eden shales), Maysville (a
new name for Orton’s Hill Quarry beds and lowermost Lebanon
beds), and Richmond (= the remainder of the Lebanon beds).
Foerste described the Fulton beds as the lowermost 4 or 5 feet
(about 1.2 or 1.5 m) of shales of the Eden containing the trilobite
Triarthrus eatoni (Foerste called it T. beckt). Bassler (1906, pp. 8-
10) moved the Maysville-Richmond boundary to equal that of
Orton’s Hill Quarry-Lebanon boundary, correlated the Fulton beds
with the New York Utica, and gave geographical names of local
derivation to Nickles’ bryozoan zones, which he treated as members.
Foerste (1914a, p. 251) concluded that beds of the “Lorraine of
New York show much greater affinities with the .. . Lorraine. . .
of Quebec than with any part of the Cincinnatian . . . of Ohio... .”
Fenneman (1916), in anticipation of the Ulrich and Bassler USGS
Cincinnati Folio, used Cynthiana in place of Point Pleasant, divided
the Eden into Utica below and Latonia above, and used Nickles’
(1902) divisions (with Bassler’s 1906 names) of the Maysvillian
and Richmond. The Ulrich and Bassler Cincinnati Folio, intended as
the much needed standard for future work in the Cincinnati area,
was unfortunately never published (the USGS refused to accept
Ulrich and Bassler’s location of the Ordovician-Silurian boundary at
the base of the Richmond, and Ulrich and Bassler were unrelenting in
16 BULLETIN 296
their position, K. E. Caster, personal communication, October 1973).
In fact, the incomplete manuscript became lost for some time and
was discovered among Bassler’s effects after his death (in 1961).
The manuscript is available in the open file of the USGS library,
Washington, D.C.
In 1925, Wilmarth (p. 86) pointed out that the USGS was at
that time employing the term Cincinnatian Series with the same
limits as those given by Winchell and Ulrich (1897) and Clark and
Schuchert (1899). Caster, Dalvé, and Pope (1955, text-fig. 3) re-
stricted the name Eden to use as a stadial term (this had come to be
its common usage) and replaced Eden with Latonia as the lithic
name (Text-fig. 4). In 1959, Sweet, et al. (pp. 1030-1032) revived
Eden as a formational name; but Weiss and Sweet (1964) objected
to use of Eden as both a rock-stratigraphic unit and a time-strati-
graphic unit and replaced Eden Formation with Kope Formation
(they restricted the name Eden to use as a stadial division).
Weiss, et al. (1965) discussed Orton’s (1873) Point Pleasant,
River Quarry, and Eden beds. They concluded that Orton’s River
Quarry beds are not a different unit from Orton’s Point Pleasant
beds and that the entire mass of sub-Eden supra-Lexington lime-
stones and shales in the Ohio River Valley should be called Point
Pleasant (p. 19). Whether to use Cynthiana (with Point Pleasant
as a member) or the older term Point Pleasant as the name of this
formation was raised as a problem needing solution (p. 21). They,
however, used Eden as a lithic name, rejected Fulton as a rock unit
(they said that what earlier workers referred to as Fulton is really
the Triarthrus eatom zone), and rejected the rock names Bassler
(1906) had given to Nickles’ (1902) faunal zones for their biostrati-
graphic rather than lithostratigraphic nature (pp. 25-28).
Peck (1966) confronted with a Cincinnati stratigraphic nomen-
clature rife with lithic names for faunal units, practically began
anew in the Maysville, Kentucky, area. Peck, using Weiss and
Sweet’s (1964) Kope Formation and accepting the Fairview Forma-
tion as a valid lithic unit, defined two new units, the Grant Lake
Limestone (overlying the Fairview) and the overlying Bull Fork
Formation. In addition, at Maysville, Peck found the Preachers-
ville Member of the Drakes Formation (described by Weir, et al.,
1965), which apparently does not occur in the immediate vicinity
of Cincinnati or on the west side of the Cincinnati Arch.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE We
Anstey and Fowler (1969) opined that Eden should be retained
as a rock-stratigraphic name and that Kope should be disregarded.
Their reasoning was that Eden could no longer be used as a stadial
name because of overlap with the New York Trentonian and thus
was available for use as a rock name. Sweet and Bergstrém (1971),
after illustrating this overlap (they showed that the upper part of
the Trenton Group is the same age as Edenian and Maysvillian
strata in the Cincinnati area), rejected Trentonian and Coburgian
in favor of the older stadial names Edenian (Orton, 1873), Mays-
villian (Foerste, 1905), and Richmondian (Winchell and Ulrich,
1897). Because Eden is a valid stadial name, Sweet and Bergstrom
reinstated Kope as a rock name to avoid confusion of dual usage
of Eden (as both a stadial and formational name).
The present state of stratigraphic nomenclature in the Cincin-
nati, Ohio, area, as synthesized from the proceeding works, will be
summarized in the following section. It is essentially the nomencla-
ture used by the United States and Kentucky Geological Surveys
jointly mapping Middle and Upper Ordovician strata in Kentucky
and by the majority of Ohio and Kentucky students of Cincinnatian
stratigraphy:
ROCK-STRATIGRAPHIC UNITS IN THE CINCINNATI AREA
The name Cincinnati Group was first used by Meek and
Worthen (1865, p. 155) for blue-gray and gray limestones and
shales outcropping in and around Cincinnati, Ohio. Although the
fossils of the Cincinnati area had been the subject of considerable
study during the early and middle 1800's, the strata’s lithologies
and relationships were without detailed description until Orton
(1873). Orton divided the Cincinnati Group into five lithic units
(in ascending order): Point Pleasant beds (exposed at Point
Pleasant, Ohio), River Quarry beds (in and along the Ohio River
at Cincinnati), Eden shales (outcropping in Eden Park, Cincin-
nati), Hill Quarry beds (at the tops of the Cincinnati hills), and
Lebanon beds (exposed at Lebanon, Ohio). The Point Pleasant
beds and River Quarry beds have been shown (James, 1891; Nickles,
1902, pp. 56-58; Foerste, 1905, p. 151; Weiss, et al., 1965, p. 19) to
be parts of the same lithic unit, the Point Pleasant Formation. The
lithic unit named Eden shales by Orton [replaced by Kope Forma-
18 BULLETIN 296
tion (Weiss and Sweet, 1964)]; Fairview Formation (Bassler, 1906)
and Grant Lake Limestone (Peck, 1966) have essentially been sub-
stituted for Orton’s Hill Quarry beds; and most of Orton’s Lebanon
beds are now called the Bull Fork Formation (Peck, 1966). Beds
comprising the Point Pleasant Formation were removed from the
Cincinnati Group by Hall (1842, p. 61), Miller, et al. (1879, pp.
193-194), and Orton (1888, p. 5). Thus, the Cincinnati Group
presently contains the Kope Formation, Fairview Formation, Grant
Lake Limestone, and Bull Fork Formation. Stratigraphers tend not
to use the name Cincinnati as a rock-stratigraphic unit. Rather,
they reserve the name Cincinnati for time-stratigraphic nomencla-
ture and avoid dual usage.
The Point Pleasant Formation, named for strata at Point
Pleasant, Ohio, by Orton (1873), is the lowest unit exposed in the
Cincinnati region. According to Weiss, et al. (1965), it consists of
thin and medium-bedded, light to dark gray, fossiliferous, biogenic
limestones parted by gray shales and mudstones. Limestone and
shale each make up about 50% of the unit; the mean clastic ratio
(limestone: shale and mudstone) calculated for successive 0.9 m
units is 1.0. Thickness ranges from a few feet (about a meter) to
nearly 70 feet (about 21 m). The unit is believed to be Shermanian
(and partly Edenian upriver from Cincinnati) in the Ohio River
Valley.
The Kope Formation, named for exposures in Kope Hollow near
Levanna, Ohio, by Weiss and Sweet (1964), conformably overlies
the Point Pleasant Formation and consists of lenses and discon-
tinuous thin-bedded, gray and bluish gray, highly fossiliferous, bio-
genic limestones (up to about a foot thick — about 30 cm thick)
and thicker sequences of gray, bluish gray, and greenish gray, less
fossiliferous shales and mudstones. The Kope is made up of 75% (or
more) shale and mudstone and 25% (or less) limestone; the mean
clastic ratio for 0.9 m units is 3.25. Thickness ranges from 150 to
270 feet (about 46 to 82 m). The Kope is Edenian in the immediate
area of Cincinnati, but the upper part becomes Maysvillian to the
east and southwest away from Cincinnati. The Point Pleasant-Kope
boundary is gradational and can be observed with certainty only
where a number of feet (a few meters) of strata on either side of
the boundary are exposed. The contact is placed at the base of the
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 19
lowest series (at least 15 feet — about 5.0 m thick) of Kope
mudrocks that lie on the uppermost limestone of the Point Pleasant
Formation.
The Fairview Formation, named for exposures in and around
Fairview Park, Cincinnati, by Bassler (1906), conformably overlies
the Kope and, as described by Peck (1966), consists of alternating
sequences of thin to medium-bedded, gray, biogenic limestones and
partings and thin beds of gray mudstones and shales. The Fairview
is composed of 50 to 60% limestone and 40 to 50% mudstone and
shale; the mean clastic ratio calculated for 0.9 m intervals is 0.5.
Thickness ranges from about 80 to 115 feet (about 24 to 35 m).
The Fairview is essentially Maysvillian, but in some areas the lower-
most part is Edenian. The Kope-Fairview contact is somewhat
tenuous except in a few outcrops; it is marked at the base of the
first thick (over 8 inches or about 20 cm) limestone that is suc-
ceeded by limestone in significantly more abundance than mudrock
and that overlies the highest series (at least 1.5 feet, about 0.5 m)
of Kope mudrock. In some areas the Fairview becomes more limy
near the top, so that a fairly thick sequence must be observed to pick
the contact with certainty.
The Grant Lake Limestone, first described by Peck (1966) from
exposures along Kentucky Route 1449 northeast of Grant Lake near
Maysville, Kentucky, conformably overlies the Fairview and con-
sists of irregularly thin-bedded, rubbly, fossiliferous, gray lime-
stones alternating with irregular partings and thin beds of fossili-
ferous, gray shales and mudstones. The Grant Lake is made up of
70 to 90% limestone and has a thickness of 100 to 120 feet (about
30 to 37 m). The age of the Grant Lake in and around its type area
is Maysvillian but has not been established elsewhere with cer-
tainty. The Fairview-Grant Lake contact is placed at the base of
the lowest sequence of irregularly bedded argillaceous Grant Lake
Limestones; the boundary is often transitional, but even when not
transitional, it is inconspicuous.
The Bull Fork Formation, named by Peck (1966) for Bull Fork
Creek near Plumville (which is near Maysville), Kentucky, and
described from exposures along Kentucky Route 1443 near Spring-
dale (also near Maysville), Kentucky, conformably overlies the
Grant Lake Limestone. It is composed of alternating thin to medium-
20 BULLETIN 296
bedded, gray, bluish gray, and greenish gray, fossiliferous, sometimes
argillaceous limestones and gray and greenish gray, fossiliferous
shales and mudstones. Clastic ratios (shale and mudstone: lime-
stone) increase from about 1:4 near the base to 4:1 near the top.
The Richmondian Bull Fork is about 200 feet (about 61 m) thick in
its type area and thins southward. The Grant Lake-Bull Fork con-
tact is usually transitional and is placed at the base of the lowest
sequence of rubbly, argillaceous Grant Lake Limestones.
The Preachersville Member of the Drakes Formation was named
by Weir, et al. (1965) for outcrops along Kentucky Route 39 about
2 miles (about 3.3 km) southeast of Preachersville, Kentucky. In the
Ohio Valley the Preachersville occurs only along the east side of
the Cincinnati Arch (actually the Preachersville occurs around the
east side of the Arch from near Dayton, Ohio, in the north to south
of Lexington, Kentucky,) where it conformably overlies the Bull
Fork and consists of green and reddish purple, calcareous to dolo-
mitic, essentially barren mudstones and thin, gray to brown, essen-
tially barren, dolomitic limestones and dolomites. Mudstone com-
prises about 90% of the unit. Thickness in the Maysville area
ranges from 25 to 30 feet (about 8 to 9 m) and increases southward.
In the Ohio Valley, the Preachersville is apparently Richmondian.
The Preachersville and Bull Fork lithologies are transitional and the
Bull Fork-Preachersville contact is placed at the top of the highest
fossiliferous Bull Fork Limestone. The boundary between the
Preachersville and the overlying Silurian Brassfield Formation may
be conformable and transitional locally. It is placed at the base of
the lowest sequence of thicker bedded, brown Brassfield dolomites
and dolomitic limestones. In most areas, however, the Brassfield rests
unconformably on the Preachersville, the Whitewater (as used by
Gray, 1972), or the Bull Fork.
TIME-STRATIGRAPHIC UNITS IN THE CINCINNATI AREA
Since Winchell and Ulrich (1897) and Clarke and Schuchert
(1899), the Cincinnatian Series has been used as the North Ameri-
can Late Ordovician time-stratigraphic unit; and, since Foerste
(1905) and Cumings (1908), the names Edenian, Maysvillian, and
Richmondian have been used as Cincinnatian stadial divisions. Al-
though two Cincinnatian stages have names derived from localities
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE Zi
outside Ohio (Maysvillian was named for Maysville, Kentucky,
while Richmondian was named for Richmond, Indiana), according
to Sweet and Bergstrom (1971, pp. 614-616) all have their reference
sections in southwestern Ohio as established in Orton’s (1873) re-
port. The Edenian reference section is without doubt in southwestern
Ohio. Orton’s unit Eden shales (Edenian Stage) was named for
exposures in Eden Park, Cincinnati, Ohio. However, location of
Maysvillian and Richmondian reference sections in southwestern
Ohio is less demonstrable.
When Foerste (1905, p. 150) used the name Maysville, he did
so for strata at Cincinnati; but Foerste’s Maysville section may
have been at Maysville, Kentucky: “. . . the name Maysville is here
suggested for the strata at Cincinnati hitherto identified as Lorraine.
Along the railroad south of Maysville, Kentucky, from the first cut
a little over a mile from town to the overhead bridge a mile north
of Summit a magnificent series of exposures gives a complete sec-
tion of all the subdivisions of the Maysville division. . . .” The Rich-
mond reference section is a similar case. Winchell and Ulrich (1897,
p. cil) first used the name: “Resting on the Lorraine [around Cin-
cinnati, Ohio] there is a series of alternating thin bedded shales
and limestones and in some localities finally a sandstone, in all quite
350 feet thick in southwestern Ohio and southeastern Indiana. Al-
most the entire series is excellently exposed at Richmond, Indiana,
so that the name Richmond group which we propose to apply to the
series 1s eminently appropriate.* [with the following footnote from
the bottom of page ciii] Prof. Orton’s [1873] name “Lebanon”
would have been adopted had his name not been used before for a
division of the Trenton by Prof. Safford [1851]. The Richmond ex-
posures besides are larger and more characteristic of the group than
those near Lebanon, Ohio. As well, although Winchell and Ulrich’s
Richmond group is nearly equivalent to Orton’s Lebanon beds,
Foerste’s Maysville is markedly different from Orton’s Hill Quarry
beds. The difference, simplified, is that Orton included strata that
was later called Arnheim (a name used by Foerste, 1905, in place of
Nickles’ preoccupied Warren, Text-fig. 4) in the Richmond, while
Foerste agreed with Nickles in including the Arnheim in the Mays-
ville. Whatever the valid reference sections for the Maysvillian and
Richmondian Stages, reference sections have come to be exposures
22 BuLLETIN 296
SUBDIVISIONS BASED ON{ SUBDIVISIONS BASED
LITHOLOGY AND FOSSILS} ON LITHOLOGY
after Peck (1966) and |after Gray
Weiss & Sweet (i964) (1972)
after Caster, Dalve’ & Pope (1955)
m
[=
A
ae
(2)
Po)
2
|
WHITEWATER
Se |
SALUDA a
LOWER =
WHITEWATER
WHITEWATER
LIBERTY P
BLANCHESTER ==
CLARKSVILLE SE=====a =
FORT ANCIENT &
OREGONIA
SUNSET
MOUNT AUBURN E==
| MOUNT AUBURN eee
CORRYVILLE §
BELLEVUE
FAIRMOUNT
MOUNT HOPE
Mo MICKEN SSSS—=
———
SSS SSsSs5
SOUTHGATE E£ = SSS
=—_e j=]
SSS
==
a
ECONOMY SSS ==
os Td
=== =
Text-fig. 4. Cincinnatian stratigraphy of the Cincinnati area (from a Uni-
versity of Cincinnati Geology Museum display).
after Peck (1966)
DILLSBORO
MAQUOKETA GROUP
| EDENIAN. —s | MAYSVILLIAN RICHMONDIAN STAGES
SSF AIRVIEW
EDENIAN MAYSVILLIAN RICHMONDIAN STAGES
LATONIA FAIRVIEW [MCMILLAN |ARNHEIMIWAYNESVILLE|
if 8
Cincinnaticrinus
Dystactocrinus
(=Atyphocrinus )
x=
fo)
Isotomocrinus
ae!
co)
co)
(=)
Es
oO
aS
oO
Ohiocrinus He He
a
Sygcaulocrinus
=>)
o)
fe[ =e]
o)
pe} 2 ]ek
H =o a
oO
oO oO
o) oO oO
oO
oO
°
{o)
Gi
ae
a
<
2
re)
=
°
°
{e)
Ho
oO
°
x=
°
2)
oo
{e)
Daedalocrinus
Apodasmocrinus
ae
°
ro)
pee :
fe) fo)
°
te)
Table 1. Historical summary of the classification of members of the Cin-
cinnaticrinacea and Homocrinacea. Column headings are: 1) Wachsmuth and
Springer (1886), 2) Bather (1900), 3) Springer (1913), +) Jaekel (1918),
5) Ulrich (1925), 6) Bassler (1938), 7) Moore and Laudon (1943), 8) Moore
(1962), 9) Moore, Lane and Strimple in Moore and Strimple, 1973 and 10)
herein. Abbreviations for families are: He—Heterocrinidae, Ho—Homocrinidae,
Cy—Cyathocrinidae, De—Dendrocrinidae, and Ci—Cincinnaticrinidae.
Ho
=
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Pfeteleie fale el
fo) be pe
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 23
in southwestern Ohio, the bluffs along Clifton Avenue and in Belle-
vue and Fairview Parks, Cincinnati (in the case of Maysvillian),
and exposures in railroad and highway cuts around and some dis-
tance south of Lebanon, Ohio, (in the case of Richmondian).
The Cincinnatian stadial reference sections need study. Ranges
of species should be firmly established in the reference sections with
extrapolation away from Cincinnati; thus far, only conodonts have
received adequate modern biostratigraphic attention. The following
organisms have been used as indices for Cincinnatian stages. Al-
though the authors have chosen organisms that are considered most
trustworthy by modern workers, the list is at best a poor one
(perhaps except for conodonts).
ORGANISMS AS INDICES OF CINCINNATIAN STAGES
CONODONTS
According to Kohut and Sweet (1968, p. 1460), an association
typical of Edenian and older strata is Cyrtontodus flexuosus (Bran-
son and Mehl), Drepanodus suberectus (Branson and Mehl), Ozar-
kodina tenuis (Branson and Mehl), Phragmodus undatus (Branson
and Mehl), and Plectodina furcata furcata (Hinde). The combina-
tion of Ambalodus, Keislognathus, Sagittodontus, probably Priomo-
dus, and Scolopodus, and perhaps Eoligonodina (genera more char-
acteristic of the Anglo-Scandinavian-Appalachian province) marks
early Edenian time; the combination of Phragmodus undatus,
Dichognathus, and Belodina is late Edenian and early Maysvillian,
while that combination without Belodina is late Maysvillian or early
Richmondian (Sweet, et al., 1959, p. 1038). This significance of
Belodina was affirmed by Pulse and Sweet (1960, p. 245), who re-
ported that all strata with Belodina are Maysvillian or older. In
addition, Pulse and Sweet (1960, pp. 243-246) submitted that
Trichonodella angulata Sweet, Turco, Warner and Wilkie and T.
subundulata Sweet, Turco, Warner and Wilkie are not known from
rocks older than Edenian and that Prioniodina delecta (Stauffer)
and 7. tenuis (Branson and Mehl) are Edenian and Maysvillian.
According to Branson, et al. (1951, p. 4), Zygognathus, Rhipidog-
nathus, and abundance and variety of Paltodus species marks Rich-
mondian.
GRAPTOLITES
Graptolites fall short of the abundance and variety of most
24 BULLETIN 296
other groups in strata around Cincinnati. Climacograptus typicalis
Hall, long thought to be a good Edenian indicator, is now known
from both younger and older strata (Pulse and Sweet, 1960, p. 239;
Berry, 1960), although Orthograptus truncatus richmondensis
Ruedemann is apparently limited to Richmondian rock (Berry,
1966).
CORALS
No corals are known from Edenian or Maysvillian strata around
Cincinnati but a few corals have been found in the Kope Formation
at Newport, Kentucky, and are presently under study (Richard S.
Laub, personal communication, October 1973). Corals are abundant
in Richmondian strata; Browne (1964; 1965) reported that Favtstel-
la alveolata Goldfuss, Foerstephyllum vacuum (Foerste), Tetradium
approximatum Ulrich, Calapoecia huronensis Billings, Avwlacera,
Grewingkia rustica (Billings), G. divaricans (Nicholson), and Saf-
fordophyllum flower Browne are common in Richmondian strata.
In addition, Paleofavosites is Richmondian and younger.
BRACHIOPODS
Resserella emacerata (Hall) (? = Onniella) was reported (Cas-
ter, Dalvé, and Pope, 1955, text-fig. 3) to be Edenian; Platystro-
phia hopensis is Maysvillian (Weiss, et al., 1965, pp. 36-37); Rhyn-
chotrema dentatum (Hall), Leptaena richmondensis Foerste (? =
Kiaeromena), Resserella meeki (Miller) (? = Onniella), Stropho-
mena planumbona (Hall) (? = S. rugosa), and Lepidocyclus capax
(Conrad) are apparently Richmondian (Caster, Dalvé, and Pope,
1955, text-fig. 3).
BRYOZOA
Constellaria florida Ulrich and Escharopora falciformis (Nichol-
son) are reported to be Maysvillian (Caster, Dalvé, and Pope, 1955,
text-fig. 3; Weiss, et al., 1965, pp. 36-37).
TRILOBITES
Cryptolithus tesselatus Green appears to be early Maysvillian or
older (Sweet, et al., 1959; Pulse and Sweet, 1960), while Triarthrus
eatont (Hall) has been used as an index of earliest Edenian as well
as the nominate species of the faunal zone named Fulton by Foerste
(1905, p. 150; Weiss, et al., 1965, pp. 26-28), although Caster, Dalvé,
and Pope (1955, text-fig. 3; pl. 2, fig. 17) reported that T. eatoni
also occurs at a higher (younger) horizon.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 25
CRINOIDS
The common occurrence of crinoids in pockets makes them only
occasionally useful in correlation. However, the abundance of pockets
in strata around Cincinnati increases their value in local correlation,
where Ectenocrinus geniculatus (Ulrich) is earliest Edenian, Cin-
cinnaticrinus varibrachialus, n.sp. is Edenian and Maysvillian, Ohto-
crinus (although rare) is known only from Maysvillian strata, and
C. pentgonus (Ulrich) is Maysvillian and Richmondian. On a broad-
er scale, Cincinnaticrinus is relatively widespread (southwestern
Ohio, northern Kentucky, southeastern Indiana, northwestern New
York, southern Pennsylvania, and Maryland) and limited to Cin-
cinnatian strata, while Jsotomocrinus and Daedalocrinus, although
less widespread (Ontario, Quebec, New York, Illinois, Minnesota,
and possibly Tennessee for the former and Ontario for the latter),
are confined to late Champlainian strata (Kuirkfieldian to Sher-
manian).
SYSTEMATIC PALEONTOLOGY
Class CRINOIDEA Miller, 1821
Subclass INADUNATA Wachsmuth and Springer, 1885
Diagnosis. — Crinoids with plates of the dorsal cup joined firm-
ly together by close suture, with a subtegminal mouth, and with arms
free above the radials (hereafter abbreviated RR) or, in some mem-
bers, above the first primibrachials (IBrr,) or second primi-
brachials (IBrre).
Discussion. — The documented range of the Inadunata is from
the Ordovician to the Triassic although Sprinkle (1973, pp. 177-183,
pls. 42-43) described an apparent crinoid (Echmatocrinus; subclass
and order undetermined) from the Burgess Shale (Middle Cam-
brian) of British Columbia which may well be an inadunate. They
are abundant in Paleozoic strata, but only one family (Erisocrinidae)
occurs in strata later than Permian. Moore and Laudon (1943)
divided inadunates into two orders, the monocyclic Disparata
[equivalent to Bather’s (1899a) Monocyclica Inadunata] with 14
families, and the dicyclic Cladoidea [equivalent to Bather’s (18992)
Dicyclica Inadunata| with 39 families. Moore (1952) changed the
ordinal names Disparata and Cladoidea to Disparida and Cladida
26 BULLETIN 296
and elevated part of the Disparida, the Hybocrinidae, to ordinal
level. Knapp (1969) segregated cladids with downflaring IBB (infra-
basals )into his new inadunate order, Declinida; however, the order
has not been accepted by subsequent authors. Comprehensive and
relatively contemporary discussions of inadunates appear in Moore
and Laudon (1943, pp. 21-64) and Moore (1962).
Order DISPARIDA Moore and Laudon, 1943
(nom. correg. Moore, 1952, p. 613
ex Disparata Moore & Laudon, 1943, p. 24)
Diagnosis. —Monocyclic inadunates with conical cup and an
armlike anal series on or branching off the C ray.
Discussion. — Disparids are characterized by structural dis-
similarity among the five rays of individuals and among correspond-
ing rays of different families. Moore and Laudon (1943, pp. 24-29)
envisioned two general groupings: a homosynbathocrinid stock and
a hybocrinid stock. The hybocrinid stock, consisting of one family,
the Hybocrinidae, with a “bowl-shaped” (krateriform, 1.¢., shaped
like a Greek krater, Text-fig 5) dorsal cup with unbranched arms
distinctly narrower than the underlying RR, was made by Moore
(1952) into the new order Hybocrinida. The homo-synbathocrinid
stock, or Disparida as Moore (1952) viewed it, included the remain-
ing 13 monocyclic families and was characterized by a steeply coni-
cal lekythosiform to skyphosiform (Text-fig. 5) dorsal cup, an arm-
like anal sac on or branching off the C ray, and wide branched arms
that articulate along the entire distal edge of the RR. Disparids
range from Ordovician to Permian.
Members of four disparid families (the Cincinnaticrinidae,
Homocrinidae, Anomalocrinidae, and Jocrinidae) occur in Cincin-
natian strata in the Cincinnati, Ohio, area; but only the closely re-
lated Cincinnaticrinidae and Homocrinidae are discussed here. The
Iocrinidae, while in need of modern treatment, are only distantly re-
lated to other Cincinnatian disparid families. Nothing new can be
added to knowledge of the Anomalocrinidae at this time, and ano-
malocrinids are discussed only in passing. The Homocrinidae and
Anomalocrinidae do not closely resemble one another but the
Cincinnaticrinidae show similarities to both the Homocrinidae and
Lf
WARN AND STRIMPLE
ORDOVICIAN-SILURIAN CRINOIDS
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28 BULLETIN 296
the Anomalocrinidae in addition to the ordinal characters. Table 1
is a historical summary of classification of these crinoids.
Superfamily CINCINNATICRINACEA, new superfamily
Diagnosis. — Disparid inadunate crinoids with a conical dorsal
cup having undivided RR in three rays (A, B, and D rays) and
compound RR in two rays (C and E rays).
Description. — The cincinnaticrinacean dorsal cup has five sym-
metrically pentagonal, sub-hexagonal, or hexagonal BB (basals) of
nearly equal size and shape. Both the compound and the fused RR
are inverted pentagons (with slight modification in the C and D
rays of members of the Cincinnaticrininae); compound RR are
typically divided about equally into a pentagonal iR below and a
quadrilateral sR above. The five rays bifurcate isotomously to form
ten arms, after which branching is isotomous or alternately heteroto-
mous. No arm cover plates are known, and they may have been ab-
sent. The arms are commonly folded tightly together making ob-
servation of the food grooves difficult except in fortuitously broken
or disarticulated specimens, but scrutiny of exposed food grooves and
end-on examination of broken arms in numerous specimens has not
disclosed the existence of cover plates. In adults, at least, the column
is quinquepartite, with each columnal composed of five radially dis-
posed fused plates or pentameres.
Discussion.— The superfamily Cincinnaticrinacea is erected
essentially to replace Heterocrinacea [Zittel’s (1879) family elevated
to superfamilial status by Ubaghs, 1953], because the type genus of
the latter (Heterocrinus) is unrecognizable. Four previously
described genera, Dystactocrinus, Isotomocrinus, Ohiocrinus, and
Atopocrinus, are available for selection as type genus. Dystactocrinus
and Ohiocrinus are rejected because they are rare and their mor-
phology is not well known. Atopocrinus and Isotomocrinus are less
typical of the superfamily than Cincinnaticrinus (Atopocrinus has
a brachianal and multipinnulate Brr, while Zsotomocrinus is the
only completely isotomously branching member of a dominantly
heterotomously branching group). Colwmbicrinus is not adequately
preserved for consideration or identification in our judgement. Cin-
cinnaticrinus is selected because it is most typical of the superfamily
and most common and widespread of the five included genera.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE
29
Text-fig. 6. Types of arm branching. Each drawing (all stylized) repre-
sents one ray with an isotomous branch to form two arms, only one of which
is fully illustrated. a-isotomous, b-alternating heterotomous, c-exotomous; b and
c are forms of heterotomous arm branching.
30 BULLETIN 296
Zittel (1879, pp. 343, 358-359) included in his family Hetero-
crinidae Heterocrinus Hall, Graphiocrinus de Koninck, Erisocrinus
Meek and Worthen, Philocrinus de Koéninck, and Stemmatocrinus
Trautschold. These are forms with fairly simple, monocyclic or
dicyclic, dorsal cups with five BB (or five BB and five IBB) and
five RR supporting long, branched or simple, arms. Wachsmuth and
Springer (1886, pp. 127-128) removed dicyclic forms, leaving only
Heterocrinus, which they split into Stenocrinus Wachsmuth and
Springer (= Heterocrinus Hall), and Heterocrinus Hall, Wachsmuth
and Springer. They also placed Jocrinus Hall in the family. Bather
(1893, p. 35) added Anomalocrinus Meek and Worthen (Wachs-
muth and Springer, 1886, p. 135 had used Anomalocrinus as nomi-
nate genus of their new family Anomalocrinidae) and Herptocrinus
Salter (= Myelodactylus Hall).
Ulrich (1925) established the modern concept of the Hetero-
crinidae as monocyclic inadunates, generally with conical cup, having
two compound and three fused RR. He transferred Ectenocrinus to
the Homocrinidae (to which he added his new genera Daedalocrinus,
Drymocrinus, and Sygcaulocrinus), reinstated the Anomalocrinidae
with Anomalocrinus and his new genus Geraocrinus (the latter was
included with reservation), and removed Jocrinus. To the previously
established heterocrinid genera, Heterocrinus and Oliocrinus, Ulrich
added his new genera Atyphocrinus, Columbicrinus, Dystactocrinus,
and Isotomocrinus.
Bassler (1938, pp. 16-17) placed a number of other genera in
the family, but of these additions only the European genera Caleido-
crinus Waagen and Jahn and Ristnacrinus Opik were accepted as
heterocrinids by Moore and Laudon (1943, p. 31). Moore and Laudon
(1944, p. 149) included Lichenocrinus, an omnium gatherum for
multi-plated discoidal Ordovician crinoid bases containing, among
other things, the juvenile holdfast of Heterocrinus. Ramsbottom
(1961, p. 39) removed Caleidocrinus to the Iocrinidae, a move with
which Moore (1962, p. 39) agreed. Moore (1962, p. 35) transferred
Ristnacrinus to the Eustenocrinidae. Lane (1970, p. 14) expanded
the concept of the family somewhat with addition of his new genus
Atopocrinus: Atopocrinus became the only member of the Hetero-
crinidae with the anal series branching off the C ray IBr,, termed
brachianal by Moore, 1962.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 31
Although Ulrich’s (1925) concept of the Heterocrinidae is ac-
cepted by us, nomenclatural and taxonomic changes are needed.
This is evident from comments of other authors: Ramsbottom
(1961), for example, was unable to assign some supposed British
heterocrinids to a definite genus: “Following the brief revision of the
Ordovician Heterocrinidae given by Ulrich (1925) it is now diffi-
cult to determine generically many species which would formerly
have been assigned to Heterocrinus....” (op. cit., p. 10). It is evi-
dent that many cincinnaticrinacean genera and species need clearer
delineation. In this paper Heterocrinus is shown to be unrecogniza-
ble, and a new genus, Cincinnaticrinus, is erected to include the Cin-
cinnati area species formerly referred to Heterocrinus. Atyphocrinus
is considered a junior synonym of Dystactocrinus. Columbicrinus,
while exhibiting cincinnaticrinacean cup features, is unrecognizable
because the holotype of the type species, C. crassus, is an incomplete
specimen lacking most of the arms and all of the stem (PI. 2, figs.
6-7). The only known specimen is from the Lebanon Limestone of
central Tennessee. Thus, the new superfamily Cincinnaticrinacea
contains Cincinnaticrinus, n. gen., Atopocrinus Lane, 1970; Dystacto-
crinus Ulrich, 1925; Isotomocrinus Ulrich, 1925; and Ohtocrinus
Wachsmuth and Springer, 1886.
The Cincinnaticrinacea show some similarities to the Homo-
crinacea and Anomalocrinacea. Cincinnaticrinacea and Anomalo-
crinacea both have two compound RR (in the C and E rays) and
three fused RR (in the A, B, and D rays) and, except for Atopo-
crinus, similar placement of anal X. However, they have divergent
cup shapes (Cincinnaticrinacea have conical cups, while Anomalo-
crinacea have krateriform cups), dissimilar arms (Cincinnaticrinacea
have subcircular arms as wide as the underlying RR, while Anomalo-
crinids have nearly round arms significantly narrower than the un-
derlying RR), and different modes of arm branching. Cincinnati-
crinacea have isotomous and alternating heterotomous arms, while
Anomalocrinacea have endotomous and alternating endotomous-
exotomous arms (Text-fig. 5).
The arms of cincinnaticrinaceans are usually found folded tight-
ly together. This may have been due to a detrimental influx of sedi-
ment and consequent contraction of muscles during catastrophic
death or to relaxation of muscles with ligamental folding of the arms
42 BuLLETIN 296
after death. The former alternative is most probable: existing crin-
oids have muscles to close the arms and ligaments to open them
(Hyman, 1955, p. 60). Lane and Macurda (1975) confirmed the
existence of muscular articulations in one upper Paleozoic inadunate
(Aesiocrinus). However, some Paleozoic crinoids may have had
only ligaments in the arms (Van Sant, 1964, p. 40), probably for
closing them. Extension was initiated by the water-vascular system.
Cincinnaticrinacea and Homocrinacea are similar in cup shape,
arm size and shape, and placement of anal X (except for Atopo-
crinus), but Homocrinacea have three compound RR (in the B, C,
and E rays) and only two fused RR (in the A and D rays). The
homocrinids Ectenocrinus and Sygcaulocrinus have similar branching
(alternating heterotomous), but Ectenocrinus has a tripartite rather
than quinquepartite column. Daedalocrinus has a similar column but
dissimilar branching (endotomous as opposed to isotomous and al-
ternating heterotomous). Moore and Laudon (1943, p. 25) en-
visioned a closer affinity for heterocrinids (Cincinnaticrinacea) and
homocrinids (Homocrinacea) than for heterocrinids and anomalo-
crinids (Anomalocrinacea) and suggested that the Heterocrinidae
developed from the Homocrinidae or their immediate forerunners.
Whether one judges the cincinnaticrinacean-anomalocrinacean or
cincinnaticrinacean-homocrinacean relationship to be closer depends
largely on which characters (¢.g. cup and arm shape, branching,
number of fused versus compound RR, or column features) are as-
sumed to be of greatest evolutionary significance. The cladid Ot-
tawacrinacea are also similar to the Cincinnaticrinacea, but because
ottawacrinaceans are dicyclic, the two superfamilies are best regarded
as homeomorphs.
Cincinnaticrinaceans occur in Whiterockian to Richmondian
rocks of western, mideastern, and eastern North America. They have
been found throughout Cincinnatian strata in the tristate Ohio-
Kentucky-Indiana area (around Cincinnati); in Edenian rocks of
northwestern New York, southern Pennsylvania, and Maryland; in
Kirkfieldian rocks of Minnesota, Wisconsin, and Illinois; in Kirk-
fieldian to Shermanian strata of mideastern Canada; and in White-
rockian strata of Utah. Kolata (1975, 1976) reported cincinnati-
crinaceans in middle Upper Ordovician strata (? Maysvillian) of
Illinois and Wyoming. In addition, Ulrich (1925) reported hetero-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 33
crinids (cincinnaticrinaceans) from Black Riverian rocks of Ten-
nessee (Lebanon Limestone), Wisconsin (probably from the
Decorah Shale, apparently Kirkfieldian in large part), and Pennsyl-
vania (Text-fig. 1).
Ramsbottom (1961, p. 10, pl. 3, fig. 8; pl. 6, fig. 5) reported
heterocrinids from Ashgillian strata of Scotland and Ireland; how-
ever, the crinoids do not appear to us to be cincinnaticrinaceans,
and are not considered herein.
Family CINCINNATICRINIDAE, new family
Because this is the only family of the Cincinnaticrinacea, familial
characters are the same as for the superfamily. Two new subfamilies
are erected herein. The subfamily Cincinnaticrininae comprises those
forms which Ulrich (1925) included in his family Heterocrinidae:
the Cincinnati forms of Heterocrinus (1.e., Cincinnaticrinus), Dystac-
tocrinus, Isotomocrinus, and Olhiocrinus. The subfamily Atopo-
crininae is erected to accommodate Atopocrinus, which is morpho-
logically, and presumably phylogenetically, distinct from other
cincinnaticrinids.
Subfamily CINCINNATICRININAE, new subfamily
Diagnosis. — Cincinnaticrinidae with a lekythosiform (steeply
conical) dorsal cup; equal-sized compound RR (in the C and E
rays) somewhat taller than the equal-sized fused RR (in the A, B,
and D rays); and an armlike anal series resting on the truncated
left corner of the C ray sR.
Description. — Cincinnaticrininae with RR, both fused and
compound, that are taller than broad. The distal left corner of the
C ray sR and the distal right corner of the D ray R are truncated
to accommodate anal X (the first anal plate), which is an inverted
pentagon equal in size to or larger than adjacent Brr. In some species
anal X enters more deeply into the cup; the proximal point of the
pentagon reaches the line of junction of the sR and iR of the C ray.
The two or three (minimally) successive anal tube plates are quad-
rilateral and appear armlike. Thecal plates as preserved are usually
smooth, but different areas in various specimens are finely nodose,
so that all ossicles may have had nodose surfaces.
34 BULLETIN 296
Each of the five RR supports a series of quadrilateral [Brr.
IBr, is the largest; it articulates with the underlying R along its
entire proximal surface. The [Brr; are fixed, 1.e., united with the
RR by immobile (synarthal? ) suture, and functioned as part of the
calyx. The uppermost IBr in each ray is a pentagonal axillary,
bearing upon its upper sloping sides two equal-sized arms (to form
a total of ten arms) made up of IIBrr, all but the last quadrilateral.
The last is a pentagonal axillary. The number of Brr in each arm
division is variable in members of the Cincinnaticrininae, both
among different rays in single individuals and among the equiva-
lent rays in different individuals. Branching on and beyond the IIBr
axillaries varies among the genera of the Cincinnaticrininae but can
be a useful taxonomic discriminant.
The column is long (probably up to about a meter), but no
complete specimens have been found. Therefore, column length,
nature of the column away from the calyx, and nature of attach-
ment (if any in adults) are matters for conjecture. However, rela-
tively good evidence exists for the column and its ontogeny for
Cincinnaticrinus varibrachialus (new herein), and the column fea-
tures of other heterocrinids are probably similar. The column is
pentapartite and pentagonal, with the points of the pentagon dis-
posed radially, although pentagonality can be shrouded by secon-
dary overgrowth to produce a round appearance. The articular
surfaces of each columnal are petaloid, with five petal-shaped arti-
cular facets, one facet per pentamere. The axial canal is small but
conspicuous, star-shaped or pentagonal, with interradial points or
angles.
Discussion. — Distribution of the subfamily Cincinnaticrininae
is the same as for the superfamily Cincinnaticrinacea except for
deletion of Whiterockian strata in Utah.
In the following generic and specific synonymies, references
that duplicate earlier illustrations or descriptions are listed with the
earlier work from which the information was borrowed. For example,
Cumings (1908) and Bassler (1919) borrowed Meek’s (1873)
illustrations of Heterocrinus heterodactylus Hall for use in their
works. The illustrations in such references are listed under the orig-
inal source in the synonymy.
ORDOVICIAN-SILURIAN CrRiINoIDs: WARN AND STRIMPLE 35
Genus CINCINNATICRINUS, new genus
1866. Hetcrocrinus Hall, Hall, p. 41; Hall, 1872, p. 210 (partim) ; Meek, 1873,
p. 1 (partim); Wachsmuth & Springer, 1880, p. 68 (partim); Springer,
1S pe 275 Ulrich, 91925: ps 83eEritz,, 1925. jp; 10) (pariim)': Moore &
Laudon, 1944, p. 149; Moore, 1962, p. 13, text-fig. 5-3; Warn, 1973, p. 12
(partim).
1886. Stenocrinus Wachsmuth & Springer, p. 207 (partim).
Type species. — Cincinnaticrinus varibrachialus Warn and
Strimple, n. sp., from Edenian and Maysvillian strata of the Cin-
cinnati area, northwestern New York, northern Maryland, and
southern Pennsylvania.
Diagnosis. — Cincinnaticrininae with a short, straight anal tube
made up of three to five facing plates; ten arms exhibiting alter-
nating heterotomous branching; equidimensional (height = width)
pentagonal BB; and height of IBrr, less than three-fourths the
height of the fused RR.
Description. — Cincinnaticrinus has the features of the sub-
family Cincinnaticrininae with some generic additions. The IBrr;
are shaped like upright, truncated cones, while the [Brr2 are in-
verted, truncated cones. Thus, the junction of the [Brr; and IBrrz
forms a constriction in the crown that marks the position of the teg-
men, above which the arms become free. This constriction appears
to have been a plane of weakness that resulted in loss and occasional
regeneration of arms. The tegmen of Cincinnaticrinus (probably of
C. pentagonus) was described by Ulrich (1925, p. 84) as “.. . gently
convex, its middle on a plane with, or slightly beneath the top of,
the fixed primibrachs. It is composed of a large polygonal central
plate around which are many much smaller, loosely fitting plates.
The smaller plates arch over the arm furrows, at least three rows
being required to cover them. On the posterior side the small plates
of the tegmen pass, evidently without break or change, into the
anterior wall of the ventral sac.” Although Ulrich was correct in say-
ing that the tegmen is located just proximal to the distal edges of
the IBrr,; (the fixed Brr), tegmen morphology appears to be quite
different from what Ulrich described; it is more similar to the
description of Wachsmuth and Springer (1886, p. 207) as “. . . five
comparatively large interradial pieces enclosing a small oral
plate... .” The tegmen (Text-fig. 8; Pl. 3, figs. 4-5) is actually made
up of five relatively large, finely nodose (Ulrich evidently inter-
BuLLeETIN 296
36
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ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE bi
preted each node as a plate), interradial plates, or orals (00), that
have their outer edges curved down into the spaces between adyja-
cent [Brr, and their inner, adjoining edges upturned to form gabled
passageways over the subtegmenal portions of the ambulacra. Three
gabled passageways (one anterior, two lateral) radiate from a central
point (presumably over the mouth) to the food grooves of the A, C,
and D ray IBrr; the two lateral (C and D ray) passageways bi-
furcate near their distal ends and send off two posterior passageways
to the B and E ray IBrr;. The CD interray O is apparently porous
and served the function of a sieve plate; it gives off numerous small
plates from its outer edge that continue up the back of the XX.
Although the tegmen morphology is known only from Cincinnati-
crinus it is probably similar in other cincinnaticrinids.
The anal structure is armlike, apparently tubular throughout
its length, although Ulrich (1925, p. 91) reported that it had been
observed in only one of hundreds of specimens; this author has seen
a maximum of only five anal plates in any of over a thousand speci-
mens. It appears that the anal tube is made up of armlike series of
three to five facing plates (XX) backed by numerous small poly-
gonal plates given off from the tegmen. Whether or not the numerous
small plates that back the XX continue beyond, or distal to, the XX
(as in Ohtocrinus) is uncertain, although it is likely they do not.
The pattern of arm branching in Cincinnaticrinus, after initial
division of the five rays (isotomous as in all Cincinnaticrinacea) to
form ten arms, is alternating heterotomous. The first of the hetero-
tomous divisions (on the IIBr axillaries) produces a large arm as
the inner branch and a smaller arm, or armlet, away from the ray;
the second division (on the IIIBr axillaries) has the arm on the
outside and the armlet on the inside; the third has the arm on the
inside and the armlet on the outside. The armlets commonly remain
simple, but bifurcating armlets have been observed in a few speci-
mens.
The XX and Brr, after initial formation as tall, narrow rec-
tangles, grow faster laterally (marginally) than vertically (perra-
dially). Thus, young (small, calyx height of about 2.5 mm or less)
Cincinnaticrinus have tall XX and Brr, while older (larger, calyx
height of about 2.8 mm or more) Cincinnaticrinus have broad IBrr
but tall TVBrr and nearly square XX, with gradation in the Brr
38 BULLETIN 296
from broad to tall away from the dorsal cup. In young Cincinnati-
crinus, the arms are so narrow that they appear to be isotomously
branched (PI. 3, fig. 11). Arms, when initially formed, may really
be isotomously branched, but with ageing heterotomy becomes in-
creasingly distinct.
Sharply V-shaped grooves (food grooves) with narrow flattened
bottoms (PI. 3, fig. 7) extend down the inner surfaces of the Brr —
two converging to one at each axillary (PI. 3, fig. 7). These grooves
deepen gradually proximally, until they reach the RR, where they
shallow rapidly, after passing beneath the tegmen, and disappear
about one-fourth the way down the RR.
Occurrence. —Edenian to Richmondian. Cincinnaticrinus is
known from the Kope, Fairview, Grant Lake, and Bull Fork Forma-
tions of the Cincinnati, Ohio, area, from the Whetstone Gulf Forma-
tion of northwestern New York; and from the upper part of the
Martinsburg Formation of southern Pennsylvania and Maryland.
Discussion. — In 1847 James Hall erected the new genus Hetero-
crinus to include the three new species H. heterodactylus (p. 279),
H. simplex (p. 280), and H. ? gracilis (p. 280). Hall did not desig-
nate a type species, nor did he refer to any of the three species as
typical (he did, however, emphasize that H. gracilis deserved only
provisional placement under the genus). Hall had a concept of the
genus Heterocrinus that allowed considerable variation; this likely
was his reason in choosing heteros (Greek for different or changed )
for the name of the genus. In including the heterocrinid H. hetero-
dactylus with the homocrinid H. simplex (now type species of
Ectenocrinus), Hall created a problem that was to be a source of
confusion until Ulrich’s (1925) revision of the Heterocrinidae. Some
paleontologists embraced Hall’s concept and included forms with
three compound and two fused RR (e.g., Heterocrinus simplex) and
forms with two compound and three fused RR (e.g., H. heterodac-
tylus), while others limited the genus to forms like H. simplex. Con-
fusion over the type species compounded the problem.
In 1866 (pp. 4-6) Hall described three more species of Hetero-
crinus and compared one of them to H. simplex. Wachsmuth and
Springer (1880, p. 69) enumerated known species of Heterocrinus
and listed H. heterodactylus as the type species. Later (1886, pp.
205-208) these authors recognized that the differences between H.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 39
heterodactylus and H. simplex are greater than specific differences,
and they redefined Heterocrinus. Of the 11 species which they placed
in Heterocrinus in 1880, in 1885 they transferred H. heterodactylus,
along with four other species, to the new genus Stenocrinus with
H. heterodactylus as type species; they left H. simplex and one other
species in Heterocrinus (with H. simplex as type species); two species
were transferred to Calceocrinus and the remaining two species were
assigned to the new genus Ofiocrinus. Wachsmuth and Springer’s
ideas concerning Stenocrinus and Heterocrinus were not accepted by
S. A. Miller (1889), who erected the genus Ectenocrinus with H. sim-
plex as type species (p. 242) and listed H. heterodactylus as type
species of Heterocrinus (p. 252). Springer (1911) recognized that
Wachsmuth and he had mistakenly substituted Heterocrinus simplex
as the type species of Heterocrinus in place of H. heterodactylus, the
valid type species: “But through some misunderstanding of types the
name Heterocrinus was assigned by us [Wachsmuth and Springer,
1885] to the wrong set of species, H. heterodactylus being Hall’s
type of that genus; therefore, Stenocrinus must go into synonymy.
Heterocrinus must be retained for the H. heterodactylus group. . . .”
(Springer, 1911, p. 27).
Hall, although he did not designate a type species, may have felt
that H. simplex was typical of Heterocrinus (in succeeding descrip-
tions of new Heterocrinus species, he referred to H. simplex but not
to H. heterodactylus); but if, in naming H. heterodactylus, Hall had
in mind the concept of what we now call virtual tautonymy, he
probably considered H. heterodactylus as typical. Whatever may
have been Hall’s original views, they were not documented and are,
therefore, not pertinent. When Wachsmuth and Springer (1880),
possibly applying the convention of page priority or the tautonymic
concept (or perhaps having communicated with Hall), listed H.
heterodactylus as the type species of Heterocrinus, they established
H. heterodactylus as the type species by subsequent designation
(Warn, 1973, pp. 10-11). Problems, however, do not end here.
Although the genus Heterocrinus Hall, 1847 and two of its
species, H. heterodactylus Hall, 1847 (the valid type species of
Heterocrinus) and H. juvenis Hall, 1866, have come to be relatively
common names, Hall’s original descriptions, figures, and type ma-
terial do not make these taxa recognizable. Modern understanding
40 BULLETIN 296
of these taxa in large part dates from Meek (1873) and Ulrich
(1925), who figured new material and described it in detail. H.
juvemis is unrecognizable for reasons discussed under Cincinnati-
crinus pentagonus (conceptually similar to Meek’s, 1973, H.
juvems). H. heterodactylus, the valid type species of Heterocrinus,
is unrecognizable because neither Hall’s (1847) figures (and descrip-
tion) nor any type material (PI. 1, figs. 1-6) shows branching be-
yond the isotomous branches on the [Br axillaries. Meek (1873)
recognized this problem when he (p. 13) considered specimens from
around Cincinnati referred to H. heterodactylus by Hall (1847):
“This is the western form that has always been referred to H.
heterodactylus, of Hall; but as the original typical specimen of that
species did not show whether or not its arms bifurcate above the
division on the last primary radial, . . . its identity with that species
can scarcely be established beyond doubt.” The names Heterocrinus
and H. heterodactylus must at present be restricted to Hall’s (1847)
type material. The new name Cincinnaticrinus is used here for the
concept of Heterocrinus put forth by Meek (1873), Ulrich (1925),
and subsequent workers, 1.e., a monocyclic inadunate with three
fused and two compound RR, a short, straight, armlike anal tube,
and alternating heterotomous branching beyond the isotomous
branch on the [Br axillaries. Cincinnaticrinus varibrachialus, the
new name for Meek’s (1873) and subsequent workers’ concept of H.
heterodactylus, is discussed later.
Cincinnaticrinus appears to differ from Ohiocrinus mainly with
respect to the anal sac. Both Ohiocrinus and Cincinnaticrinus have
an anal tube that is an armlike branch of about four plates off the
C ray sR. In Cincinnaticrinus the XX are backed by small polygonal
plates given off from the tegmen to form a short narrow tube; the
four (or five) XX of Ohiocrinus have a backing of small polygonal
plates which extends away from the XX (rather than closing around
the back to form a tube) and beyond (distal to) the XX as a high,
inflated, polyplated coil with wide whorls. For a time the authors
thought the differences to be preservational (1.e., that Ohiocrinus
were well-preserved Cincinnaticrinus); but so many well-preserved
Cincinnaticrinus, all lacking coiled anal sacs, have been examined
that it now appears the two are distinct.
Cincinnaticrinus probably evolved from an earlier cincinnati-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 41
crinid. The step from Jsotomocrinus to Cincinnaticrinus is simply
one of making isotomous arms heterotomous (isotomous to heteroto-
mous is a common evolutionary trend in crinoids, Moore and
Laudon, 1943, p. 10) and shortening the anal tube somewhat.
Cincinnaticrinus and Ohiocrinus are certainly similar; it is easier
to derive Ohiocrinus from Cincinnaticrinus with coiling and elonga-
tion of the polyplated anal sac and elongation of the arms than
to do the reverse, although Ofiocrius could be an independent
offshoot from Jsotomocrinus (the arms of O. brawni are nearly iso-
tomous). Dystactocrinus is also like Cincinnaticrinus, from which
it probably evolved. Evolution of Cincinnaticrinus to Dystacto-
crinus requires only regularization of branching (constancy in
number of Brr in each division series in single specimens and among
different individuals is apparently an evolutionary endpoint in cin-
cinnaticrinids), broadening of BB, and enlargement of [Brry.
Cincinnaticrinus is known from thousands of specimens (all but
a handful from in and around Cincinnati, Ohio,) and is easily the
best known of cincinnaticrinids. Two species, C. varibrachialus (new
herein) and C. pentagonus (Ulrich), 1882, are recognized. Hetero-
crinus isodactylus Miller, 1875, may belong to Cincinnaticrinus but
must, at present, be restricted to its holotype. Heterocrinus 1sodac-
tylus Miller may be conspecific with Cincinnaticrinus pentagonus,
but Miller’s drawing and description are poor and hardly allow this
to be suggested with much authority.
There appears to be a trend in Cincinnaticrinus toward thicker
columns through time. C. varibrachialus has a column with a rela-
tively consistent width (proximal column diameter is about half
distal cup diameter) through Edenian and into Maysvillian time.
Maysvillian C. pentagonus (with columns having proximal column
diameter somewhat smaller than distal cup diameter), however, give
way to even broader columned forms in Richmondian time (Rich-
mondian C. pentagonus have columns with proximal diameter about
equal to distal cup diameter).
Cincinnaticrinus varibrachialus, new species Pls. 3-5; Text-fig. 8
1873. Heterocrinus heterodactylus Hall, Meek, p. 12, pl. 1, figs. 1a-b; Cumings,
1908, pl. 3, figs. 5, 5a; Bassler, 1919, pl. 53, figs. 5-6; Ulrich, 1925, p. 83,
text-fig. 3a; Moore & Laudon, 1944, pl. 52, fig. 11.
42 BULLETIN 296
Text-fig. 8. Cincinnaticrinus varibrachialus. a & b—tegmen; c—plate dia-
gram. a—oral view, a drawing of UCGM 40575L (X15); rays are lettered A,
B, C, D, and E, scoring is vertical on the RR, horizontal on the IBrn, and
diagonal on X; OO are unmarked (except, on the CD interray O, dots which
represent pores): b, crosssection of the OO (30) in the plane marked by the
two lateral lines in figure a; OO are upturned where they join, presumably
over the five ambulacra, and have their outer edges turned down between the
IBrri; the two arrows point to funnel-shaped (in cross-section) pores in the
CD interray O; c — exploded diagram of Cincinnaticrinus varibrachialus.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 43
1925. Heterocrinus difficilis Ulrich in Ruedeman, p. 76.
1973. Heterocrinus tenuis Billings, Warn, p. 10, pl. 1, figs. 2-19 (nox fig. 1).
Primary type material.—The holotype is here designated
UCGM 3871 (the specimen illustrated by Meek, 1873, pl. 1, figs.
la-b as Heterocrinus heterodactylus?). Paratypes are here desig-
nated UCGM 40497, 40500, 40502, 40531, 40555, 40556, and 405751.
All primary types are from Edenian strata in the Cincinnati, Ohio,
area.
Diagnosis. — Cincinnaticrinus with steeply conical (lekythosi-
form) cup and narrow column, so that in uncrushed specimens dis-
tal cup diameter is at least 1.4 times as great as proximal cup (or
proximal column) diameter.
Description—C. varibrachialus, in addition to generic and high-
er characters, has BB (Text-fig. 9) and RR that expand distally and
make the dorsal cup conical. This is more obvious in juveniles, which
have globular calyces. With growth, the angle formed by the edges
of the cup (in lateral view) decreases as the sides of the BB and
RR approach a parallel condition (compare Pl. 5, figs. 1-2 and
12-13). No new cup plates are added during ontogeny (that is,
during that part of the ontogeny that is known), and shapes and
relative size ratios of cup ossicles change little, other than widening
of the bottoms of the BB and RR. Thus, the smallest (youngest )
and largest (oldest) crinoids have dorsal cups that are nearly identi-
cal except for size.
Cincinnaticrinus varibrachialus, with two to seven [Brr (com-
monly three, four, or five), three to seven I]Brr (four or five is most
common), and four to six IIIBrr (commonly four or five), has arms
that are more variable than in any other cincinnaticrinid aside from
Tsotomocrinus tenuis. Ramules given off at heterotomous branches
usually remain simple and have more plates, but bifurcating ramules
have been observed in a few specimens. Warn (1973) described
small (smaller than adjacent Brr) doubly convex (marquise) plates
occurring in some specimens at various places in the arms (in the
IBr series most commonly between IBr; and IBro, PI. 3, fig. 3).
No complete Cincinnaticrinus varibrachialus specimens (hold-
fast, column, and crown or calyx) are known. However, that juve-
niles (and possibly adults) have an obscurely polyplated, inverted
+4 BULLETIN 296
b c
d &
Text-fig. 9. Five general basal plate shapes in cincinnaticrinaceans and
homocrinaceans.
a—distally expanding, symmetrically pentagonal (Cincinnaticrinus varibrachia-
lus, Isotomocrinus tenuis, Atopocrinus priscus, and Homocrinus parvus). b—
parallel-sided, symmetrically pentagonal (C. pentagonus, Ohiocrinus laxus—
also e in some members, O. brauni, Daedalocrinus bellevillensis, Ectenocrinus
simplex, E. geniculatus, and Sygcaulocrinus typus — in the case of the last
three, only the AB interray B). c—asymmetrically pentagonal with one steeply
sloping and one gently sloping upper side (S. typus — all but the AB interray
B). d—asymmetrically pentagonal with one steeply sloping and one horizontal
upper side (E. simplex and E. geniculatus — in both, all but the AB interray
B). e—symmetrically hexagonal (Dystactocrinus constrictus and occasionally
O. laxus.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 45
saucer-like (lichenocrinid) basal attachment is fairly certain (and
has been known for some time, see discussion). In collecting locali-
ties 1,2, and 4 (Text-figure 10) C. varibrachalus calyces and crowns
and lichenocrinid bases and columns have been found in abundant
association. As well, an ontogenetic sequence from lichenocrinid to
cincinnaticrinid column is evident among separate columns and in
single columns collected from these, and other, pockets.
Juvenile C. varibrachialus holdfasts (Pl. 5, figs. 15-16) are
roughly circular discs, usually attached to such foreign objects as
adult C. varibrachialus columns, other adult crinoid columns,
brachiopods, bryozoans, trilobites, pelecypods, or phosphate nodules.
They range in diameter from less than one mm to about five mm,
with most having a diameter of about two or two and one-half mm.
They have a convex, obscurely polyplated, upper wall (roof) and
a large flat plate as the lower wall (floor). When the inhabited sub-
strate is not flat, concomitant changes in shape occur, é.g., when en-
crusting crinoid columns they curl around the column (PI. 5, fig.
16). The lichenocrinid column protrudes from a central depression
or crater in the roof. Internally, five primary lamellae extend from
the periphery of the holdfast and meet at the center. These lamellae
rest upright on the floor. Second, third, and fourth order lamellae
are inserted serially between the primary lamellae and may or may
not reach the center. The lamellae (PI. 5, fig. 15) appear to support
the roof. (For detailed information on lichenocrinid bases in general,
affinities unknown, see Faber, 1929).
Juvenile Cincinnaticrinus varibrachialus columns (lichenocrinid
columns — PI. 3, figs. 3, 10) show an interesting morphologic (and
apparently ontogenetic) sequence. They are composed distally (z.e.,
adjacent to the base) of five vertical series of hexagonal plates with
the plates of each series alternating with laterally adjacent plates
(1.e., plates of adjacent series) to form zigzag sutures between series
(Text-fig. 11). These grade proximally into a section with alter-
nating plates that abut, so that a straight suture is formed between
series (Text-fig. 11). Farther proximally, abutting plates come to
lie in parallel planes to form circlets of five plates (Text-fig. 11).
Each circlet has one plate from each of the five series. Transition
to the cincinnaticrinid column occurs with gradation into columnals
by fusion of the five plates (pentameres). Thus, the most proximal
46 BULLETIN 296
INDIANA
KENTUCKY
@35
Text-fig. 10. Localities of Cincinnaticrinus varibrachialus collected in con-
nection with this study. Numbers refer to localities described in text.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 47
(youngest) part of the juvenile column (= distal and oldest of
adult ?) is round with equal-sized columnals (Text-fig. 11), each
made up of five pentameres.
The adult Cincinnaticrinus varibrachialus column is either
pentagonal proximally grading distally into terete, or round through-
out. The proximal pentagonal part can be made up of two sets of
columnals: larger, more rounded columnals and smaller, markedly
pentagonal columnals inserted between the larger columnals. Each
columnal is composed of five fused plates with each plate forming a
point in the pentagonal columnals. With secondary secretion of
stereom (seemingly a normal feature of column aging), the column
becomes round, but the former pentagonal column can be seen inside
the round sheath in cross-section.
Occurrence. —Edenian and Maysvillian from the Kope and
Fairview Formations around Cincinnati, Ohio; the Whetstone Gulf
Formation of northwestern New York; and the Martinsburg Forma-
tion in Maryland and southern Pennsylvania. Ruedemann (1925, p.
70) briefly described a Cincinnaticrinus from zone I of the Whet-
stone Gulf as Heterocrinus dtfficilis. This species was to be further
described later, but never was, by E. O. Ulrich: it is considered here
to be a junior synonym of C. varibrachialus. The authors have found
C. varibrachialus only in the Kope Formation, in which crinoids
are fairly common, weathering more easily out of the predominantly
shaley unit than from the more limy Fairview above. The dominant-
ly calcareous Fairview (“Hill Quarry beds”) is less propitious for
well-preserved cincinnaticrinids. However, during the 1800’s and
early 1900’s dozens of quarries were operating in the Cincinnati area
and more good Fairview exposures were available. Fortunately, area
museums have specimens from the “Hill Quarry beds” (Fairview
Formation) collected around the turn of the century.
Discussion. — Cincinnaticrinus varibrachtalus is erected to
house the taxon that Meek (1873) and Ulrich (1925) made known
as Heterocrinus heterodactylus. H. heterodactylus must be con-
sidered as unrecognizable, because Hall’s (1847) type material
(AMNH 1116/1, PI. 1, figs. 46; AMNH 1116/2, Pl. 1, fig. 3;
AMNH 1116/3, Pl. 1, figs. 1-2), illustrations (Hall, 1847, pl. 76,
figs. la-o), and description (Hall, 1847, p. 279) do not demonstrate
the nature of the arm branching.
48 BULLETIN 296
Au
==
Text-fig. 11. Ontogeny of the Cincinnaticrinus varibrachialus column.
a, b, c and d are portions of the juvenile column viewed progressively more
proximally; a is the most distal portion (i.e., nearest the holdfast) and is com-
posed of five vertical series of hexagonal plates with the plates of each series
alternating with laterally adjacent plates to form zigzag sutures between series;
these grade proximally into a section (b) with alternating plates that abut,
so that a straight suture is formed between series; farther proximally, abutting
plates come to lie in parallel planes to form circlets of five plates (c) still
farther proximally the circlets of five plates fuse to form pentapartite columnals
(d) e and f are portions of the adult column; the oldest portion of the adult
(f) is round with equal-sized columnals, which are apparently first secreted as
pentagonal columnals and become round with secondary secretion of stereom;
this grades proximally into a portion (e) with larger, more rounded columnals
(in some specimens an additional proximal portion is made up of equal-sized
pentagonal columnals).
a
2
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 49
The trivial name varibrachialus is chosen to describe the vari-
ability in number of Brr per division series that is characteristic of
the species. Originally the word “brachial” was used as an adjective
to denote arm ossicles (e.g., brachial plates or ossicles; primibrach-
ial plates or ossicles). More recently, the word “brachial” has come
to be widely used in an abbreviated sense as a noun to denote arm
ossicles (¢.g., brachials, primibrachials); and abbreviation is often
carried even further (e.g., brachs, primibrachs). The trivial name
varibrachialus, chosen for its descriptiveness, is somewhat awkward,
in that it represents latinization of an anglicized Latin word. How-
ever, it is chosen over the original Latin brachialis because the latter
would give the name too much breadth of meaning (varibrachtalis
would mean simply arm variation). The trivial name varibrachialus
is more appropriate to denote variation in number of Brr per divi-
sion series, as typifies the arms of this species.
Great variability in number of [Brr (and higher Br series), both
in single individuals and among different individuals, appears to be
a feature unique to cincinnaticrinids. Warn (1973) interpreted the
smaller marquise-shaped Brr as intercalates and judged intercalation
of Brr to sufficiently explain the brachial variability in Cincinnati-
crinus varibrachialus. In that paper, 61 different IBr arrangements
were reported from a single pocket of 72 crowns (specimens with all
five IBr series still intact), 116 partial crowns (one to four [Br
series), and 219 calyces (no complete IBr series). Similiar variation
and variation in additional populations has since been found. Kesling
and Strimple (1971) reported, in Eutaxocrinus wideneri (a flexible
crinoid), IBr and IIBr variation (considered mutation by Kesling
and Strimple) from a basic plan of two [Brr per ray and three or
four I[Brr per arm; in cincinnaticrinids, however, variation within
limits (eé.g., two to seven [Brr for Cincinnaticrinus varibrachialus )
seems to be the rule, rather than an exception.
Lichenocrinus was described by Hall (1866, p. 9) for what the
authors herein have referred to as lichenocrinid bases. Hall thought
these to be the “bodies” of parasitic crinoids because of their con-
sistent attachment to other organisms. Meek (1871, 1872b, 1872c)
and Sardeson (1899, p. 275) theorized that Lichenocrinus might
actually represent basal attachments of crinoids. Schuchert (1904,
p. 268) stated more definitely that Lichenocrinus are bases of
50 BULLETIN 296
fata pase len
Ashgill Richmondian
Maysvillian
te
Caradoc Shermanian
Io
SILURIAN
Cincinnatian
Kirkfieldian
ORDOVICIAN
Rocklandian
Black
Riveran
Ie tp
Llandeilo
Llanvirn
Champlainian
Chazyan
EA <8
Whiterockian
Arenig :
Canadian
Ea
Text-fig. 12. Ranges of cincinnaticrinacean and homocrinacean species.
European stages are on the left of the column; North American series and
stages are on the right. A—Ibexocrinus lepton, B—Altopocrinus priscus, C—
Apodasmocrinus daubei, A. punctatus; D—Isotomocrinus minutus, E—I, tenuis;
F—Daedalocrinus bellevillensis; G—Cincinnaticrinus wvartbrachialus; H—
Ecterocrinus simplex; I—E. geniculatus; J—Cincinnaticrinus pentagonus ; K—
Sygcaulocrinus typus; L—Dystactocrinus constrictus; M—Ohiocrinus laxus; O.
brauni; N—Ectenocrinus sp. indet.; O—Homocrinus parwus.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 51
crinoids. Springer (1917, p. 11) reported the affinity of Heterocrinus
and lichenocrinid bases but cited no evidence, “This curious disc-
like body [lichenocrinid base] . . . is now known to be the encrusting
root of a very small crinoid of the Heterocrinus type.” Perhaps
Springer was referring to material sent to him for description by an
amateur collector, George M. Austin, in 1903 (see below). Foerste
(1925, pp. 102-103) alluded to complete juvenile heterocrinids with
lichenocrinid bases and column in the USNM collection. Unfortu-
nately, Foerste could not discern the plate arrangement of the calyx,
and the authors have not been able to locate the specimens (perhaps
Foerste, too, was referring to the Austin material, see below).
Bassler (1928) indicated that George M. Austin, from Wilmington,
Ohio, had discovered evidence for the lichenocrinid-heterocrinid af-
finity in 1898 and had (in 1903) communicated his discovery to
Frank Springer for description, which Springer never did. Faber
(1929, pp. 455-456) reported that in 1898 he, G. Ashman, and A.
Albers found three tiny crinoids complete with lichenocrinid bases.
According to Faber, these specimens were never illustrated or
described and disappeared, along with part of Albers’ collection,
just before Albers’ death. Fenton (1929) discussed Austin’s 1898
material (USNM 89862a-f, the material that Springer was to have
described) in detail. Reexamination of this material confirms Fen-
ton’s observation that some heterocrinids (cincinnaticrinids) and
some lichenocrinids represent different parts of the same organism.
USNM 89862a-f consists of 20 lichenocrinid bases attached to
Rafinesquina, an Isotelus fragment, and trepostome bryozoan frag-
ments associated with lichenocrinid columns and three juvenile
Cincinnaticrinus sp. cf. C. pentagonus crowns from Richmondian
strata near Clarksville, Ohio. One of these crowns appears to have
been attached to one of the lichenocrinid columns; only a 0.8 mm
long furrow, presumably the result of loss of a portion of the column
from the slab, separates the crown from the column.
While earlier workers’ evidence, as well as recent observations
[herein and by Weaver (1976)] indicates a lichenocrinid base-
juvenile Cincinnaticrinus varibrachalus (and probably C. penta-
gonus) affinity, resolution of the matter of priority of Lichenocrinus
as a generic name is delayed until more information, hopefully from
BULLETIN 296
On
bo
discovery of complete specimens, is available. Considerable morpho-
logic variation in lichenocrinid bases suggest that juveniles of a
variety of Cincinnatian crinoids have lichenocrinid holdfasts. One
such association involving Jsotomocrinus has been discussed by
Kolata (1975, p. 27).
Miller (1874) described the axial changes in the Heterocrinus
heterodactylus (C. varibrachialus) column through much of its
length but said nothing of the lichenocrinid nature of the distal
(juvenile) column and base. Bather (1891, pp. 400-401, text-fig. 5;
1900, p. 89, text-fig. 3) described a similar distal-proximal columnar
gradation for the cladid Botryocrinus decadactylus from the Wen-
lock Limestone.
It appears that in adults new columnals are added both at the
base of the calyx and intercalated serially for a short distance dis-
tally between older columnals. Columnals are, in both cases, first
added as five discrete radial plates which fuse to form pentagonal .
columnals. Older columnals become round with secondary secretion
of stereom. Addition of new columnals could cease at some stage in
ontogeny; if this occurs all columnals eventually would become
round. Thus, most adult columnals have a proximal section made up
of pentagonal columnals, grading distally into a series of pentagonal
columnals alternating with larger rounded columnals, and finally
into a distal region of round columnals only. Some columns (of
younger crinoids?) are pentagonal for proportionally greater dis-
tances, while others (of older crinoids?) are round throughout their
observed length. This is in apparent agreement with “Jackson’s
law” (Jackson, 1896; 1899), which is essentially: In organisms pos-
sessing organs which grow by the serial addition of parts, the onto-
geny of the organ tends to rehearse its phylogeny. ‘
Warn (1974) described swellings, which he interpreted as myzo-
stome galls, in columns of Heterocrinus juvents (= Cincinnaticrinus
pentagonus, herein). The authors have seen similar galls in columns
of C. varibrachialus and Ectenocrinus simplex. These have recently
been reinterpreted as annelids (Phosphannulus) by Welch (1976).
The rarity of complete juvenile C. vartbrachialus is probably a
result of breakage of the fragile column during or after death, either
before burial or during exposure and subsequent collecting. However,
the association of lichenocrinid bases with juvenile C. varibrachialus
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 53
crowns is pervasive. No complete adult specimens (7.¢., none with
holdfasts) and no associations of large crowns and holdfasts are
known. This may be due to post-mortem transport and differential
deposition of crowns and proximal parts of the columns apart from
the holdfasts and distal parts of the columns as hypothesized by
Brett (1976) for supposed Caryocrinites roots. Conversely, con-
siderable evidence suggests that adults were eleutherozoic. The deli-
cate, attenuated “lichenocrinid” distal column would seem to have
been inadequate to support the adults upright during life and would
have easily broken in currents, if not autotomized as a matter of
course in normal development (Text-fig. 13). As well, occasionally
sections of Cincinnaticrinus columns are found with a single rounded,
apparently abraded, end. It is unlikely that delicate Cincinnaticrinus
crowns could be differentially transported for any great distance,
and it is likely significant that despite the genus’ abundance no as-
sociation of adult crowns and holdfasts has yet been observed. Thus
we believe that the genus was effectively eleutherozoic as an adult.
A similar conclusion was reached by Weaver (op. cit.).
Warn (1973, p. 13, table 1) noted that various species of
Heterocrinus described during the classical period of paleontology
were characterized as having different [Br arrangements. The brach-
ial arrangements of any of these species would fit into the normal
intraspecific variation of any large population of C. varibrachialus.
It is probable that H. exilis Hall, 1866, H. extgwus Meek, 1872a,
H. propinquus Meek, 1873, and C. varibrachialus (new herein) are
conspecific. However, type material for H. exiguus and H. propin-
quus has not been located; H. exilis is apparently based on a juvenile
of questionable affinity. Because of the unavailability or inadequacy
of type material we prefer to restrict these names to the types rather
than to synonymize them. Similar reasoning, plus the inadequate
nature of the existing figures and descriptions, precludes referring
this taxon (C. varibrachialus) to one of the earlier described species.
It is not surprising that such workers as Hall, Meek, and Ulrich
assumed intraspecific constancy in number of [Brr per ray. Vari-
ability in number of IBrr has seldom been documented for fossil
crinoids with the exception of cincinnaticrinids and one anomalous
population of Eutaxocrinus widenert (Kesling & Strimple, 1971).
Indeed, all Recent comatulids (non-stalked crinoids) have either two
54 BULLETIN 296
Cc
Text-fig. 13. Ontogenetic change in the life habit of Cincinnaticrinus vari-
brachialus.
a—attached juvenile with polyplated (lichenocrinid) column and holdfast. b—
attached adult with expanded, “adult” column proximal to the thin, juvenile
(lichenocrinid) column. c—adult breaks free (whether because of increased
current activity or autotomization is unknown) at the attenuated juvenile
column and thereafter lives unattached.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 55
or four IBrr per ray, although in Hyman’s (1955, p. 92) opinion,
the generic allocation of existing pentacrinites (stalked crinoids) is
in a state of confusion largely because of the use of number of Brr
in each division series as a taxobasis.
Cincinnaticrinus varibrachialus probably evolved from I[soto-
mocrinus tenuis (Billings) by reduction of the anal tube and by
transformation of isotomous to alternating heterotomous branching.
In other respects the two genera and species are similar. C. vart-
brachialus seems to have given rise to C. pentagonus. Such evolution
would have encompassed widening of the column and proximal calyx
and initiation of the trend toward regularization of branching, which
in cincinnaticrinids culminates in Dystactocrinus constrictus.
Cincinnaticrinus pentagonus (Ulrich), 1882 Pl. 6
1873. Heterocrinus juvenis Hall, Meek, p. 10, pl. 1, figs. 3a-c; Cumings, 1908,
pl. 3, figs. 3, 3a-b; Ulrich, 1925, text-fig. 4c; Warn, 1974, pl. 1, figs. 1, 9.
1882. Heterocrinus pentagonus Ulrich, p. 176, pl. 5, figs. 10, 10a.
Primary type material. — YPM 24801 and 24802 are syntypes
of H. pentagonus Ulrich, 1882. YPM 24801 is herein designated lec-
totype and YPM 24802 lectoparatype of H. pentagonus. Both are
Maysvillian and are from Cincinnati, Ohio.
Diagnosis. — Cincinnaticrinus with cylindrical dorsal cup and
wide column, so that in uncrushed specimens distal cup diameter
is less than 1.4 times as great as proximal cup (or proximal column)
diameter.
Description. —C. pentagonus has parallel-sided pentagonal.
BB (Text-fig. 8) and large, distally tapering IBrr, (although the
IBrr, are less than three-fourths as tall as the fused RR). The
dorsal cup is hardly wider than the proximal column, and the arms,
when folded (as is nearly always the case), continue nearly straight
from the calyx, so that crowns attached to sections of column are
not conspicuous features as in other cincinnaticrinids (and as in
most crinoids), where the crowns are obvious expansions at the ends
of the columns. As in Cincinnaticrinus varibrachialus, no new cup
plates are added during the known part of ontogeny, and the
smallest and largest C. pentagonus dorsal cups are nearly identical
but for size.
Brachial variability in C. pentagonus seems to be somewhat
56 BULLETIN 296
smaller than in C. varibrachialus, although it is less well known; a
peculiarity of C. pentagonus is that specimens are seldom found with
arms above the IBrr, the fixed IBrr. C. pentagonus seems to vary
little (three to five IBrr) around a basic plan of four [Brr per ray.
Variability beyond the IBr series is poorly known. The column of
C. pentagonus is like that of C. varibrachialus but broader and with
a greater propensity for roundness.
Occurrence. — Maysvillian and Richmondian. C. pentagonus is
known from the Fairview, Grant Lake, and Bull Fork Formations
of the Cincinnati, Ohio, area. Heterocrinus juvems Hall, Meek
(= C. pentagonus) was reported by Meek (1873, p. 12) from the
“... upper part of the Cincinnati group near Lebanon, Ohio” ( Rich-
mondian). Ulrich (1882, p. 176) described H. pentagonus “. . . from
the Cincinnati group at Cincinnati about 375 feet [about 115 m]
above low-water mark in the Ohio river”: the Fairview outcrops at
that elevation in Cincinnati.
Discussion. — The name Cincinnaticrinus pentagonus (n.
comb.) is applied to crinoids Meek (1873) called Heterocrinus
juvenms Hall, 1866. Just what Hall’s concept of H. zuvents was is
unfortunately unclear, in large part because of questionable type
material. Hall described H. juvenis in 1866 but did not illustrate
it until 1871. Whitfield and Hovey (1898, pp. 24-25) listed AMNH
1173/1 as the holotype of H. juvenis and the specimen figured by
Hall (1871, pl. 1, figs. 9-10; 1872, pl. 5, figs. 9-10). However, the
specimen which presently carries this American Museum number and
label is neither that figured by Hall nor that described by Whitfield
and Hovey as the type. The holotype was reported by Whitfield
and Hovey (1898, pp. 24-25) to have been a free calyx, and Hall’s
(1871; 1872) figures are two views of a free calyx, but AMNH
1173/1 is a specimen imbedded in a slab (pl. 2, fig. 5). Further,
Hall’s (1871, 1872) figures and (1866; 1872) description are of a
juvenile with diameters of distal calyx and proximal column nearly
equal (this feature is a specific character of C. pentagonus), while
AMNH 1173/1 is probably (the specimen is far from complete) a
juvenile C. varibrachialus with distal cup diameter nearly twice that
of the column (or proximal cup). Hall’s original specimen has
evidently been lost or misplaced.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE ST
Division of Cincinnaticrinus into two species may be somewhat
artificial, because there is convergence of distal cup diameter to
proximal cup (and proximal column) diameter ratios in the two.
Examination of targe numbers of both taxa has established the fol-
lowing: 1) young (small — cup height of about 2.5 mm or less)
Cincinnaticrinus pentagonus have ratios equal to about 1.2, while
young (similarly sized) C. varibrachialus have ratios of about 2.0;
2) with age, the column and proximal cup of both broaden relative
to the distal cup, but the column of C. varibrachialus widens propor-
tionally more than that of C. pentagonus so that with increasing
size (age) C. varibrachialus ratios become smaller faster; 3) Cincin-
naticrinus populations with ratios of about 0.9 to 1.3 appear to be
segregated from populations with ratios of about 1.5 to 2.2 (the
former group appears to be Maysvillian and Richmondian, while the
latter is Edenian and Maysvillian). Choice of 1.4 as the major dif-
ferentiating feature of the two species is somewhat arbitrary; this
figure was chosen because it is the number (expressed to the nearest
tenth) that falls closest to the midpoint between the highest ob-
served ratio (to the nearest tenth) in populations clustering around
1 and the lowest cbserved ratio (to the nearest tenth) in popula-
tions clustering around 2. Specific identification of individual speci-
mens of Cincinnaticrinus on this basis is frequently problematic, be-
cause it requires measurement of uncrushed (nearly round in oral
or aboral view) dorsal cups. Such preservation is uncommon, but
the problem is not without solution. In general, the distal cup 1s
flattened more than the proximal cup and column, with increase
(apparent) in ratios. One can, by averaging the shortest and longest
diameters in each of the two planes of measurement, convert an ap-
parent ratio (from a distorted specimen) to an approximation of the
“real” ratio that is probably close enough to be useful.
Cincinnaticrinus pentagonus likely evolved from C. varibrachia-
lus with broadening of the column (presumably a response to in-
creased current activity) and proximal cup and reduction in num-
ber of IBrr. A similar evolutionary trend may occur in Ectenocrinus
(see below). C. pentagonus probably gave rise to Dystactocrinus
constrictus with increased regularization in number of IBrr, broaden-
ing of BB, and enlargement of [Brry.
58 BULLETIN 296
Genus DYSTACTOCRINUS Ulrich, 1925
1925. Dystactocrinus Ulrich, p. 87; Moore & Laudon, 1943, p. 14, text-fig. 1;
Moore & Laudon, 1944, p. 149; Moore, 1962, p. 13, text-fig. 5—1a-b (1b
is a copy of Hall, 1871, pl. 1, fig. 13 as Heterocrinus constrictus).
1925. Atyphocrinus Ulrich, p. 85; Moore, 1962, p. 13, text-fig. 5—6a-c (6b-c
are from Ulrich, 1925, text-fig. 4a-b as Atyphocrinus corryvillensis).
Type species. — Heterocrinus constrictus Hall, 1871 from Mays-
villian strata at Cincinnati, by original designation of Ulrich (1925,
p- 87).
Diagnosis. — Cincinnaticrininae with an anal tube evidently like
that of Cincinnaticrinus; with ten arms exhibiting alternating hetero-
tomous branching; with I[Brr; large, nearly the same size as the
fused RR (width of IBrr; is nearly equal to that of the fused RR;
height is three-fourths or more the height of the fused RR); and
with distinctly hexagonal BB, noticeably broader than tall (Text-
fig. Ye).
Description. —In addition to familial and higher characters,
Dystactocrinus has markedly hexagonal BB, three-fourths or less as
tall as wide. In general, plates of the dorsal cup tend to be shorter
and broader than in other cincinnaticrinids. The IBrr; are large,
rectangular in plan view (actually, the IBrr; are tumescent and,
thus, are shaped like a barrel cut longitudinally in half), and about
the same size as the fused RR. A constriction occurs in the crown
in the plane of the distal ends of the IBrr,.
The arms are broader than in other cincinnaticrinids, while the
armlets are narrow (about the same as in other cincinnaticrinids),
so that the arm to armlet width ratio is high and is a striking feature
evident even from cursory examination. Dystactocrinus has only two
or three IBrr per ray and three or four I]Brr per arm, with armlets
beyond the IIBr axillaries branching off every third or fourth Br.
The proportionally small size of the armlets and the extent of regu-
larization of branching (not attained by other cincinnaticrinids )
gives the arms the near appearance of pinnulation. In reality, there
is gradation in crinoids from heterotomous branching to “pinnula-
tion,” with armlets in the former becoming pinnules in the latter.
Use of the term pinnulation, while descriptive in some cases, empha-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 59
sizes differences between some related forms which are actually
slight and clouds phylogentic relationships, ¢.g., armlets of Cincin-
naticrinus, with heterotomous branching, are certainly homologues of
“pinnules” in closely related Ectenocrinus, said to be pinnulated, and
the use of different terms for such similar branching is unfortunate.
The broadness of the arms of Dystactocrinus prohibits them
from being folded into a tight bundle (as in other cincinnaticrinids )
and causes the crown to be expanded distally. This distal crown
expansion emphasizes the constriction at the bottom of the free arms
(at the articulation of the IBrr,; and IBrr2) — thus, Hall’s specific
name. The column of Dystactocrinus is like that of Cincinnati-
crinus varibrachialus but with a greater tendency toward completely
round columnals.
Occurrence. — Maysvillian (?Kirkfieldian, Shermanian, or
Edenian, and Maysvillian). Dystactocrinus (monospecific) is known
from only a few specimens from the Fairview and Grant Lake Forma-
tions of Cincinnati and environs. Hall (1872, p. 211) described
D. constrictus (as Heterocrinus constrictus) from a single specimen
from limestone of the “Hudson-river group” at Cincinnati; Meek
(1873, p. 4) reported that Hall’s specimen had been found about
100 feet below the tops of the hills at Cincinnati and that another
species, H. compactus (a junior synonym of D. constrictus) occurs
at the same level (Fairview Formation). Ulrich (1925, p. 85)
described Atyphocrinus corryvillensis (a junior synonym of D. con-
strictus) from the Corryville member of the McMillan Formation at
Cincinnati (= Grant Lake Limestone).
Ulrich (1925, p. 88) alluded to two undescribed species of
Dystactocrinus, each represented by a single specimen, from older
strata, one from the “Trenton limestone” at Ottawa, Ontario,
(= Hull-Kirkfieldian, Sherman Fall-Shermanian, or Coburg beds-
Edenian?), and another from the “Cynthiana limestone” at West
Covington, Kentucky, (= Point Pleasant Formation). The Point
Pleasant at Cincinnati is Shermanian and the “Trenton limestone”
of New York and Canada has been shown to be Fdenian and Mays-
villian (Sweet and Bergstrém, 1971).
Discussion. — Dystactocrinus probably evolved from Cincin-
60 BULLETIN 296
naticrinus by enlargement of [Brr,, broadening and/or shortening
of BB, reduction in number of [Brr, and regularization of branching.
The genus gave rise to no known successors. The arm characters
suggest that Dystactocrinus is a cincinnaticrinacean homeomorph
of the homocrinacean genus Apodasmocrinus.
Dystactocrinus constrictus (Hall), 1871 1G of
1871. Heterocrinus constrictus Hall, pl. 1, figs. 13-14; Hall, 1872, p. 210, pl. 5,
figs. 13-14; Meek, 1973, p. 3, pl. 1, figs. 10a-b; Ulrich, 1925, p. 87, text-
fig. 6a; Moore, 1962, pl. 1, figs. la-b.
1873. Heterocrinus constrictus var. compactus Meek, p. 4, pl. 1, fig. 11.
1925. Atyphocrinus corryvillensis Ulrich, p. 85, text-figs. 4a-b.
1925. Dystactocrinus constrictus (Hall), Ulrich, p. 87, text-figs. 6b-e, p. 88.
1944. Dystactocrinus constrictus (Hall), Moore & Laudon, pl. 52, fig. 11.
Primary type material.— MCZ 2165 (Hall, 1871, pl. 1, figs.
13-14 and herein, PI. 7, figs. 5-7) from Maysvillian strata at Cincin-
nati is the holotype of H. constrictus Hall, 1871.
Because D. constrictus is at present the only known species of
Dystactocrinus, the specific diagnosis, description, and occurrence
are the same as for the genus.
Discussion. — D. constrictus, while having numerous features in
common with Cincinnaticrinus varibrachialus, differs from it in hav-
ing wider cup plates, a tendency to reduce [Brr from four or five
to two or three, arms which branch on every third or fourth IIIBr
and higher, and a marked contrast in width of the arms and armlets.
In the last three of these four respects, Dystactocrinus constrictus is
more like the Cincinnatian homocrinid Ectenocrinus simplex. It is,
however, certainly a cincinnaticrinid and shows closer relation to
Cincinnaticrinus pentagonus, which also tends, as compared to C.
varibrachialus, to broaden cup plates and to reduce [Brr from four
or five to three or four. In addition, both C. pentagonus and D. con-
strictus have broad columns. It appears that C. pentagonus gave
rise to D. constrictus.
Ohiocrinus exilis Foerste, 1914b (p. 125, pl. 1, fig. 7) has armlets
that are markedly smaller than the arms and may be conspecific
with D. constrictus, but the holotype (USNM 78718, pl. 2, fig. 2),
and only known specimen, consists only of arms, and assignment to
D. constrictus is uncertain.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 61
Genus ISOTOMOCRINUS Ulrich, 1925
1925. Isotomocrinus Ulrich, p. 86; Moore & Laudon, 1944, p. 149; Moore, 1962,
p. 13, text-fig. 5-2; Kolata, 1975, p. 26.
Type species. —Isotomocrinus typus Ulrich, 1925 by original
designation (1925, p. 87): this species is a junior synonym of Hetero-
crinus tenuis Billings, 1857.
Diagnosis. — Cincinnaticrininae with isotomous arm branching.
Description. —Isotomocrinus has the general features of the
subfamily and, aside from its isotomous arm branching and anal
sac, is like Cincinnaticrinus. The anal sac is tubular, as in Cincin-
naticrinus, but is somewhat broader and much longer and composed
of more facing plates (at least seven or eight). The sides of the dorsal
cup (in lateral view) form the largest angle of all Cincinnaticrininae;
this is a product of distally expanding BB (as in C. varibrachialus )
and RR. Jsotomocrinus has C and D ray RR that are wider than in
other cincinnaticrinids and wider than other RR (in the A, B, and E
rays) in single specimens.
The IBrr, taper more and the IBrrz expand less, if at all, than
in other cincinnaticrinids, which makes for proportionally narrower
arms. The arms are long with few branches. There are two to six [Brr
per ray and four to nine []Brr (and higher series) per arm. Brachial
variability seems to be similar to that in Cincinnaticrinus vari-
brachialus, both for individuals and for the genus (and species) in
general.
The column is pentaparitite with interradial pentameres and
with a pentagonal lumen having radially disposed points. The column
is pentagonal near the cup and becomes gradually more rounded
distally. There appear to be two sets of columnals of different size
alternating in position proximally, but only one size distally. Distal
columnals evidently become similar in size and shape with secondary
overgrowth.
Occurrence. — Kirkfieldian (? Blackriverian to Kirkfieldian,
Shermanian, or Edenian; Rocklandian to Kirkfieldian, Shermanian,
or Edenian; Kirkfieldian to Shermanian or FEdenian). [sotomo-
crinus is known from the Hull beds of Kirkfield, Ontario; Hull, Sher-
man Fall, and/or Coburg beds of Ottawa and Montreal; the “Tren-
ton limestone” at Trenton Falls, New York; the Decorah shale,
St. Paul, Minnesota, (UM 9274); and the Dunleith and Grand
BULLETIN 296
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ORDOVICIAN-SILURIAN CrINOIDS: WARN AND STRIMPLE 63
Detour Formations, Illinois. Billings (1857, p. 274; 1859, p. 50)
described Heterocrinus tenuis from the “Trenton limestone” of Ot-
tawa and Montreal (= Hull, Sherman Fall, and/or Coburg beds).
Springer (1911, p. 25) alluded to the same species in the Hull
crinoid beds at Kirkfield, Ontario. Wilson (1946, p. 32) listed it
from Hull (GSC localities 34 and 37), Sherman Fall (GSC locality
44), and Coburg (GSC localities 4, 9, 13, 38, 39, 52, and 53) beds of
Ontario and Quebec. Ulrich (1925, p. 87) alluded to two undescribed
species: one from “limestone of Black River age” of central Pennsyl-
vania (? = pre-Rocklandian Hatter or Hunter Limestones), another
from “Upper Black River” of Wisconsin (? = unnamed pre-Rock-
landian Limestones or the Kirkfieldian Decorah Shale). Jsotomo-
crinus is common in the Hull crinoid beds at Kirkfield Quarry, Kirk-
field, Ontario. A possible [sotomocrinus, briefly discussed by Brower
and Veinus (1974, pp. 20-21) under the heading “Jsotomocrinus,
n. sp.” was reported from Blackriverian rocks of Tennessee by those
authors. Because of the limited nature of available material this
occurrence will not be further considered in this paper, and the
reference is cited only for completeness.
Discussion. — The nature of the arm branching suggests that
Isotomocrinus may have been the progenitor of all Cincinnati-
crininae, and possibly of anomalocrinids as well. Evolution of [soto-
mocrinus to Cincinnaticrinus could occur with alteration of isoto-
mous to alternating heterotomous branching and shortening of the
anal tube. Jsotomucrinus may have arisen from Atopocrinus or
Ectenocrinus; alternatively [sotomocrinus and Atopocrinus or Isoto-
mocrinus and Ectenocrinus may share an as yet unknown common
ancestor.
Isotomocrinus tenuis (Billings), 1857 Pl. 8; Text-fig. 15
1857. Heterocrinus tenuis Billings, p. 273; Billings, 1859, p. 50, pl. 4, figs. 6a-b,
pl. 10, figs. 1a-c; Springer, 1911, p. 25; Jaekel, 1918, p. 85, text-fig. 79;
Wilson, 1946, p. 32; Warn, 1973, p. 10, pl. 1, fig. 1 (nom figs. 2-19).
1925. Isotomocrinus typus Ulrich, p. 87, text-figs. 5a-b; Moore & Laudon, 1944,
oll, Sah, saver, ail
1925. Heterocrinus juvenis Hall, Fritz, p. 10, text-fig. 7.
21971. Ectenocrinus, n. sp. Steele & Sinclair, pl. 16, figs. 10-11.
1975. Isotomocrinus tenuis (Billings), Kolata, p. 27.
Primary type material. — GSC 1438 (the only remaining of
Billing’s syntypes) was designated lectotype of H. tenuis by Wilson
64+ BULLETIN 296
a b
Text-fig. 15. Isotomocrinus tenuis.
USNM S.2077a-b are the primary types of J. tyfus Ulrich, 1925, which is a
junior synonym of Heterocrinus tenuis Billings, 1857. a—camera lucida drawing
of a CD interray view of USNM S.2077a; areas with question marks contain
what appear to be disarticulated anal backing plates; b—camera lucida draw-
ing of a CD interray view of USNM S.2077b.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 65
(1946, p. 32; Warn, 1973, pp. 11-12). GSC 1438 was adequately
illustrated by Warn (1973, pl. 1, fig. 1) but is illustrated again here
(Pl. 8, fig. 5). USNM S.2077a (Pl. 8, figs. 1, 4) is the holotype of
I. typus, a junior synonym of [sotomocrinus tenuis. USNM S.2077b
(PI. 8, figs. 2, 4) and USNM S.2077c (PI. 8, figs. 3, 4) are paratypes
of I. typus. GSC 1438 is from the Kirkfieldian Hull beds at Ottawa,
Ontario, while USNM S.2077a, b, and c are from the Kirkfieldian
crinoid beds at Kirkfield, Ontario.
The specific diagnosis and description of J. tenuis are essentially
identical to that of the genus, given above. Comparisons with I.
minutus Kolata, the only other known species, are made in the dis-
cussion of the latter. Occurrences of J. tenuis are coextensive with
those of the genus, excepting the Grand Detour Formation record.
Discussion. —I. tenuis is known from the Trenton Limestone
of Kirkfield and Ottawa, Ontario, and Montreal, Quebec, and from
the Buckhorn Member, Dunleith Formation, Illinois. Numerous
good specimens, mainly from Kirkfield, are housed in the collections
of: Royal Ontario Museum, Geological Survey of Canada, United
States National Museum, and University of Cincinnati Geological
Museum (in the Kopf Collection).
The specimen figured by Steele and Sinclair (1971, pl. 16, figs.
10-11) as a new species of Ectenocrinus appears to be a cincinnati-
crinid. It resembles both J. tenuts and Cincinnaticrinus vartbrachia-
lus but shows no arm branching, critical for differentiation between
these two species. Because its occurrence is more reconcilable with
TI. tenwis than with C. varibrachialus, the authors have tentatively
referred it to J. tenuis. The specimen has a more steeply conical cup
than in other /sotomocrinus tenuis specimens and a smaller anal X
than any other cincinnaticrinid.
Heterocrinus tenuis (Billings) was referred to Isotomocrinus by
Kolata (1975, p. 26), who regarded it and I. typus as separate
species. For the reasons stated above we prefer to subsume J. typus
into I. tenuts.
A good case can be made for evolution of Cincinnaticrinus vari-
brachialus from Isctomocrinus tenuis with reduction of the anal tube
and transformation of isotomous to alternating heterotomous branch-
ing. In fact, 7. tenuis probably gave rise, directly or indirectly, to all
other Cincinnaticrininae. Choosing a progenitor for [. tenuis is a
66 BULLETIN 296
greater problem. No known crinoid (other than a member of the
Cincinnaticrininae) exhibits a sufficiently obvious close morpho-
logical relationship with J. tenwis (including Atopocrinus, the only
older cincinnaticrinid) to merit consideration as a progenitor of J.
tenuis.
Isotomocrinus minutus Kolata, 1975
1975. Isotomocrinus minutus Kolata, pl. 4, fig. 4, text-figs. 4, p. 27.
Primary type material. — UI X-4886 is the holotype; two para-
types are UI X-4940 and UI X-491. All types are deposited in the
collections of the University of Illinois (UI).
Diagnosis. —Isotomocrinus with small steep-sided dorsal cup
and anal X deeply set within dorsal cup (Kolata, of. cit., p. 27).
Remarks. — This species has been well described by Kolata (op.
cit.) and will not be redescribed here. Pending further study the
species is accepted as valid, but the possibility remains that the
three specimens placed in J. minutus by Kolata are juvenile I. tenuis.
The steep-sided dorsal cup and more strongly pentagonal stem as
compared to adult /. tenuis could be immature features; juvenile
Cincinnaticrinus as noted above, exhibit similar morphology. The
ontogeny of Jsotomocrinus, however, is not so well known as that of
the related Cincinnaticrinus, so the possibility remains that J.
minutus is a valid species. The noted deeper penetration of the anal
X into the cup in J. minutus could also be a feature that changes
during ontogeny. All of the J. tenuis from the Dunleith, while oc-
curring in a different formation, are much larger. The finding of un-
doubted J. tenuis of similar size to J. minutus would be necessary,
in our opinion, to solidly establish the species.
Genus OHIOCRINUS Wachsmuth & Springer, 1886
1886. Ohiocrinus Wachsmuth & Springer, p. 208; Miller, 1889, p. 263; Wachs-
muth, 1900, p. 152; Springer, 1911, p. 27; Springer, 1913, p. 212; Ulrich,
1925, p. 90; Moore, 1962, p. 13, text-fig. 5-4a-d ((4a, b, d are from Ul-
rich, 1925, p. 90, text-figs. 7a-c).
Type species. — Heterocrinus laxus Hall, 1866 by original desig-
nation of Wachsmuth and Springer (1886, p. 208).
Diagnosis. — Cincinnaticrininae with spirally coiled anal sac and
ten arms exhibiting alternating heterotomous branching.
Description. — Like Cincinnaticrinus, Ohtocrinus has an anal
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 67
sac that is an armlike branch of four (or possibly five) facing plates
(XX) off the C ray sR, filled out by numerous small backing plates
proliferated from the tegmen. In Cincinnaticrinus the backing plates
close around the back of the XX to form a short, straight tube; but
in Ohtocrinus the backing plates extend away from, and beyond,
the XX as an inflated, polygonally polyplated, high-spired coil with
wide whorls (Text-fig. 17). In contrast to Wachsmuth and Springer’s
(1886, p. 208) description of the anal sac as composed of “. . . num-
erous hexagonal pieces, arranged alternately, and in longitudinal
rows,” the backing plates are polygonal (quadragonal, pentagonal,
hexagonal, or septagonal) and are apparently not arranged in
definite rows or circlets. In other respects (column morphology, B
and [Br shape, etc.) Ohiocrinus is like Cincinnaticrinus.
Occurrence. — Maysvillian. Ohiocrinus is known from the Fair-
view Formation from Cincinnati, Ohio, and Madison, Indiana.
Discussion. — For a time, the authors thought Cincinnaticrinus
and Ohiocrinus to be congeneric, for it seemed that Ohtocrinus
(specimens with spiral anal sacs) were simply Cincinnaticrinus with
preservation of the polyplated sac. This view was bolstered by
Wachsmuth and Springer’s (1886, p. 208) footnote to the descrip-
tion of their new genus Ohtocrinus:
“Ohiocrinus resembles Stenocrinus [a junior synonym of Hetero-
crinus but used for Meek’s, 1873 concept of Heterocrinus that
is herein called Cincinnaticrinus] very closely, and can only be
upheld by the form of the ventral tube. We [Wachsmuth and
Springer] never saw the appendage of Stenocrinus [Cincinnati-
crinus|, but Mr. S. A. Miller claims it to be distinct, and this
induced us to make the separation.”
However, so many beautifully preserved Cincinnaticrinus with the
anal sac ending as a short tube are now known that the two seem to
be distinct, as Miller postulated.
Ulrich (1925, p. 90) described Ohiocrinus as having a dorsal cup
structurally similar to those of Cincinnaticrinus and Dystactocrinus
but with great variation due to “breakage and irregular regeneration
of parts.” Ulrich did not know the repository of Hall’s holotype
(MCZ 2167) of Heterocrinus laxus, type species of Ohiocrinus, but
had at least six specmens (USNM 42304a-e and an unlocated speci-
men represented by Ulrich, 1925, text-figs. 9, 9a) which he had in
68 BULLETIN 296
1882 used as the basis for his new species H. oehanus and which
he thought were perhaps conspecific with H. laxus. For H. laxus,
Ulrich substituted (at least conceptually) H. oehanus as the type
species of Ohiocrinus. Three of the four best (of the six) syntypes
of H. oehanus are abnormal specimens (Text-figs. 16a-b-c), which
caused Ulrich to characterize Ohiocrinus as having great cup vari-
ability. The authors, after comparing the types of H. laxus and H.
oehanus, believe the two to be conspecific. H. oehanus is, then, a
junior synonym of HZ. laxus.
Ohiocrinus evolved from Cincinnaticrinus with elongation, in-
flation, and coiling of the tubelike anal sac and gave rise to no known
successors.
Ohiocrinus laxus (Hall), 1871 Pl. 9: Text-figs. 16a-c
1871. Heterocrinus laxus Hall, pl. 1, fig. 15; Hall, 1872, p. 211, pl. 5, fig. 15;
Meek, 1873, p. 5, pl. 1, fig. 12.
1882. Heterocrinus (Iocrinus) oehanus Ulrich, p. 175, pl. 5, figs. 9, 9a-c.
1925. Ohiocrinus laxus (Hall), Ulrich, p. 90, text-fig. 7a; Moore, 1962, pl. 1, fig.
5s
1925. Ohiocrinus oehanus (Ulrich), Ulrich, p. 90.
Primary type material.— MCZ 2167 (pl. 7, figs. 5-7) is the
holotype of H. laxus Hall, 1871.
Diagnosis. — Ohiocrinus with markedly heterotomous branch-
ing, 7.e., with arms strikingly broader than the armlets.
Description. —O. laxus has a distally expanding crown that
widens uniformly and long arms with numerous branches (six to ten
per arm). Each division series has three to seven Brr (four or five
is most common). Number of Brr per division series appears to be
variable, both among individuals and in different rays of the same
individual, but not to the extent as in Cincinnaticrinus varibrachia-
lus. Whereas a single specimen of C. varibrachialus might have as
many as four different numbers of IBrr in the five rays (e.g., UCGM
40500 with the following IBr arrangement: A-2, B-4, C-4, D-3, E-5),
a single specimen of Ohiocrinus laxus typically has fewer different
numbers of IBrr in five rays (e.g., MCZ 2167 with: A-5, B-4, C-4,
D-4, E-4). This diminished variability is apparently true also for
higher division series. Thus, while total intraspecific variation in
number of Brr per division series in O. laxus is as great as in Cin-
cinnaticrinus varibrachialus (species range of variation > individual
69
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE
)
laxus
nym of Ohiocrinus :
a—exploded diagram of USNM 42304a. b—exploded diagram of USNM
42304b. c—exploded diagram of USNM 42304c.
Text-fig. 16. Ulrich’s (1882) abnormal specimens
syno ‘TINt
(junior
70 BULLETIN 296
range of variation), variation in single individuals is greater in C.
varibrachialus. Armlets given off at the axillaries bifurcate two or
three times and appear to reach the tips of the arms.
Occurrence. — Maysvillian. O. laxus is known from at least
seven specimens from the Fairview Formation at Cincinnati. Hall
(1872, p. 211) described Heterocrinus laxus from the “Hudson-river
group” at Cincinnati. Ulrich (1882, p. 176) described H. oehanus, a
junior synonym of O. laxus from “on the hills back of Cincinnati,
Ohio, at an elevation of about 325 feet above low-water mark in the
Ohio river” (= Fairview Formation).
Discussion. —USNM 42304a (Pl. 9, fig. 9; Text-fig. 16a) is
herein designated lectotype and the specimen figured by Ulrich
(1882, pl. 5, figs. 9, 9a) and USNM 42304b-e (Text-figs. 16b-c)
lectoparatypes of H. ochanus Ulrich, 1882, a junior synonym of H.
laxus Hall, 1871, the type species of Ohiocrinus. The specimen repre-
sented by figures 9 and 9a has not been located, although a note, -
apparently in Ulrich’s handwriting, accompanying the USNM type
specimens reads: “Remainder of Oeh’s spms [specimens] are at
Yale.” However, search at Peabody Museum of Natural History,
Yale University (1973), failed to reveal other specimens.
O. laxus probably arose from Cincinnaticrinus varibrachialus by
elongation, inflation, and coiling of the anal sac and elongation of the
arms, either directly or with O. brawnt as intermediary.
O. laxus and Dystactocrinus constrictus (Hall) Ulrich, 1925
may be conspecific. Wachsmuth and Springer (1886, p. 208) placed
Heterocrinus constrictus Hall, 1872 (type of Dystactocrinus Ulrich,
1925) in their new genus Ohiocrinus (with H. laxus Hall, 1871 as
type species). O. laxus has broad arms and narrow armlets, as does
D. constrictus, and the BB are markedly hexagonal (as in D. con-
strictus). One specimen (UCGM 23048; Pl. 9, fig. 1) of O. laxus
particularly resembles D. constrictus. At present, however, the two
are considered distinct, for examination of specimens referred here
to D. constrictus has brought to light no spiral anal sac, although
the anal tube-bearing ray is visible on some specimens.
Ohiocrinus brauni Ulrich, 1925 Pl. 10; Text-fig. 17
1925. Ohiocrinus brauni Ulrich, p. 90, text-figs. 7b-c.
Primary type material. —USNM S.2082a (Ulrich’s text-fig. 7b)
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE fal
a 2 2p
Sacey an
\
6%
a Neots
Text-fig. 17. Ohtocrinus brauni.
a—camera lucida drawing of a CD interray view of USNM S. 2082b. b—the
same from a slightly different angle. c—camera lucida drawing of an E ray
view of USNM 6S. 2082a.
72 BULLETIN 296
and S.2082b (Ulrich’s text-fig. 7c) are syntypes of O. brawm. USNM
S.2082b (PI. 10, figs. 1-3; Text-fig. 17) is herein designated lectotype
and USNM S.2082a (Pl. 10, figs. 4-6; Text-fig. 17) lectoparatype
of O. brauni. Both are Maysvillian, from Madison, Indiana.
Diagnosis. — Ohtocrinus with nearly isotomous branching, 1.e.,
with the arms and armlets of about the same width.
Description. — O. brauni has arms with three or four branches
per arm and three to four (commonly four?) Brr per division series.
Br variability is apparently smaller than in O. laxus. Arms and arm-
lets are about the same size, but armlets (given off on alternate sides
beginning with the first abradially or away from the ray) continue
unbranched to the tips of the arms.
Occurrence. — Maysvillian. O. brawnt is known from only two
specimens from the Fairview Formation at Madison, Indiana.
Discussion. — O. braum, while differing from O. laxus, may not
really be distinct; it is conceivable that specimens referred to O.
brauni might be juveniles of O. laxus. This will remain uncertain,
however, until our knowledge of the ontogeny of O. laxus approxi-
mates that known for Cincinnaticrinus varibrachialus, Cincinnatt-
crinus pentagonus, and Ectenocrinus simplex. O. braum could have
arisen from C. varibrachialus by elongation, inflation, and coiling
of the anal sac, either directly or with O. laxus as intermediary.
Subfamily ATOPOCRININAE, new subfamily
Diagnosis. — Cincinnaticrinidae with a conical (less steeply
than in the Cincinnaticrininae) dorsal cup; with unequal-sized com-
pound RR in the C and E rays; the C ray R is somewhat shorter
and the E ray R somewhat taller than the nearly equal-sized fused
RR in the A, B, and D rays; the anal series is an armlike branch
off the C ray IBr; (termed brachianal by Moore, 1962).
Genus ATOPOCRINUS Lane, 1970
*1970. Atopocrinus Lane, p. 14.
Type species.— Atopocrinus priscus Lane, 1970 by original
designation (p. 14), Whiterockian of Utah.
*The generic name Atopocrinus was first used by Clark (1912) for an extant
comatulid crinoid. It was later used by Lane (1970) for an Ordovician in-
adunate from Utah. Lane (pers. comm., Mar. 22, 1977) proposed the substitute
name Othneiocrinus for the Ordovician form.
ORDOVICIAN-SILURIAN CrrInNoIDS: WARN AND STRIMPLE 73
Description. — Atopocrinus has equidimensional, pentagonal BB
that expand distally. The E ray sR extends nearly to the distal
margins of the A and B ray [Brrz. The C ray IBr; (brachianal) has
a truncated left shoulder to support anal X; succeeding [Brr are
narrower and rest on the remaining distal edge of the IBr;. There are
two arms in the A and B rays, but branching in the C, D, and E rays
is unknown. The A and B ray [Brr are as wide as the underlying RR
(the A and B ray IBrr are quite low rectangles), but the C and E
(and apparently D) ray IBrr are much narrower than the under-
lying RR: the C and E ray IBrr are low, nearly square rectangles.
The [Brr and one or two proximalmost II Brr lack armlets; the next
ten or so Brr have armlets given off from every Br on alternate
sides, with the first given off as an inner branch; succeeding Brr
have armlets given off one or both sides of each Br. Where there are
two armlets per Br, they are offset, indicating derivation from an
alternating heterotomous condition by fusion of two adjacent Brr.
Armlet facets on the oral surfaces of the arms are connected to the
ambulacral groove by oblique grooves that join the ambulacral
groove alternately (Text-fig. 18c). The stem is circular, pentapar-
tite with radial pentameres, and has a proximal portion that tapers
rapidly distally (as in members of the Homocrininae).
Occurrence. — Whiterockian. Atopocrinus is known from a sin-
gle specimen from the M zone (of Hintze, 1951) of the Kanosh Shale
near Ibex, Utah.
Discussion. — Among inadunates, branching of the anal series
off the third radial plate of the C ray, rather than off the first or
second, is a rarity. The only other known inadunate with this C ray
plate arrangement is Peniculocrinus Moore, 1962. However, Atopo-
crinus differs from Pentculocrinus in having compound RR in two
rays rather than in all five. Possession of an anal series as a branch
off the C ray IBr,; (branchianal) is a primitive feature and supports
the view that the anal series originated as a C ray arm branch that
came to be modified and incorporated into the calyx.
Atopocrinus’s branching in the distal portions of the arms is
unique among disparids and appears to have been derived from an
alternating heterotomous condition by fusion of adjacent Brr in sets
of two. Derivation from an ancestor with alternating heterotomous
arms is not only supported by food groove configuration of bipin-
74 BULLETIN 296
nulate Brr but also by individual arm ontogeny (unbranched —
alternating heterotomous — bipinnulate Brr).
Atopocrinus priscus Lane, 1970 Pl. 2; Text-fig. 18
1970. Atopocrinus priscus Lane, p. 15; p. 8, text-fig. 2f-j; p. 11, pl. 1, figs. 4-6.
Primary type material. — The holotype and only known speci-
men of A. priscus is USNM 165240.
Because A. priscus is presently the only known species of Atopo-
crinus, the specific diagnosis, description, and occurrence are the
same as for the genus.
Superfamily Homocrinacea Kirk, 1914
(nom. transl. Ubaghs, 1953 ex. Homocrinidae Kirk, 1914)
Diagnosis. — Disparid inadunate crinoids with a steeply conical
dorsal cup having undivided RR in two rays (in the A and D rays)
and compound RR in three rays (in the B, C, and E rays).
Description.— The homocrinacean dorsal cup has five sym- .
metrically or four asymmetrically and one symmetrically pentagonal
BB, about equal in size. The distal left corner of the C ray sR and
the distal right corner of the D ray R are truncated to accommodate
anal X, which is a branch off the C ray. Succeeding XX are quad-
rangular and backed by numerous small polygonal plates to form a
tubular (and armlike) anal sac (this is unknown in Jbexocrinus but
known to varying degree in other homocrinaceans). Each of the five
RR supports a series of quadrilateral [Brr. IBr; articulates with the
underlying R by immoble suture (and is thus fixed) along its entire
proximal surface.
Discussion. — Kirk (1914) erected the family Homocrinidae for
Homocrinus. Kirk was of the opinion that, while Homocrinus is re-
lated to the Heterocrinidae, especially to Ectenocrinus (this was,
of course, prior to Ulrich’s, 1925, transferral of Ectenocrinus to the
Homocrinidae), the Heterocrinidae could not house Homocrinus.
Jaekel (1918, p. 54), believing that Homocrinus was dicyclic, added
his new genera Nassoviocrinus, Jahnocrinus, and Ascocrinus (all
dicyclic and none closely related to Homocrinus). Ulrich (1925)
added Ectenocrinus Miller, 1889, and his new genera Drymocrinus,
Daedalocrinus, and Sygcaulocrinus. Drymocrinus is considered as a
junior synonym of Ectenocrinus. Lane (1970, p. 12) added his new
genus Ibexocrinus. Ubaghs (1953) elevated Kirk’s family Homo-
crinidae to superfamily Homocrinacea. The superfamily Homo-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 75
crinacea as now envisioned, then, contains Homocrinus, Daedalo-
crinus, Ectenocrinus, Apodasmocrinus, n.g., [bexocrinus, and Sygcau-
locrinus.
Text-fig. 18. Atopocrinus priscus.
All are of USNM 165420, after Lane (1970).
a—E ray view. b—C ray view. c—ventral view of arm. d—exploded diagram.
76 BuLLeETIN 296
Homocrinaceans are morphologically similar to cincinnati-
crinaceans; both typically have steeply conical cups, similar arm
size and shape, and similar placement of X (except for Atopo-
crinus); homocrinaceans have three compound RR (in the B, C, and
E rays) and only two fused RR (in the A and D rays); Ecteno-
crinus, Apodasmocrinus, Ibexocrinus, and Sygcaulocrinus have alter-
nating heterotomous branching as do the Cincinnaticrinacea (except
Isotomocrinus, which has isotomous branching).
Homocrinaceans occur in Whiterockian to Niagaran rocks of
western, central, mideastern, and eastern United States and mid-
eastern Canada. They have been found in Edenian and Maysvillian
strata in the tristate Ohio-Kentucky-Indiana area (around Cincin-
nati); in Shermanian and Edenian rocks of northwestern New York;
in the Edenian of southern Pennsylvania; in Kirkfieldian to Edenian
rocks of the Ottawa-St. Lawrence lowland of Canada; in Rich-
mondian strata of Iowa; in Whiterockian strata of Utah; in Maysvil-
lian strata of Wyoming; in Blackriverian rocks of Oklahoma, Ten-
nessee and Virginia; and in Niagaran rocks of New York.
Family HOMOCRINIDAE Kirk, 1914
Because this is the only family of the Homocrinacea, familial
characters are the same as for the superfamily. Two subfamilies are
envisioned here. The subfamily Homocrininae contains Homocrinus,
Ectenocrinus, Apodasmocrinus, Ibexocrinus, and Sygcaulocrinus. The
subfamily Daedalocrininae is erected to accommodate Daedalocrinus,
which is somewhat removed, morphologically and presumably phylo-
genetically, from other homocrinids.
Subfamily HOMOCRININAE Kirk, 1914
(nom. transl. ex Homocrinidae Kirk, 1914)
Diagnosis. — Homocrinidae with equal-sized compound RR (in
the B, C, and E rays) somewhat taller than the equal-sized fused
RR (in the A and D rays) and with a round column that tapers
rapidly distally just below the dorsal cup.
Description. — Members of the subfamily Homocrininae have
IBrr; that taper distally and IBrr2 that expand slightly distally. The
column is round with a pentagonal lumen and tapers rapidly distally
just below the calyx. [bexocrinus, however, is an exception to both
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE ai
statements; it has rectangular [Brr,; and IBrre, and the column of
Ibexocrinus, while round, tapers more gradually than in other Homo-
crininae.
Discussion. — Apparently, members of the Homocrininae have
a point of columnal generation at the base of the rapidly tapering
proximal portion of the column rather than at the base of the dorsal
cup, the common location of columnal addition (aside from distal
insertion ). It appears that a trend in Homocrininae is to incorporate
proximal columnals into the calyx. This feature is undeveloped in
Tbexocrinus, the oldest of the Homocrininae; well developed in
Ectenocrinus; and best developed in Sygcaulocrinus and Homocrinus,
the youngest of the Homocrininae.
Members of the Homocrininae occur in Whiterockian to
Niagaran rocks of western, central, mideastern, and eastern United
States and mideastern Canada. They have been found in FEdenian
and Maysvillian strata around Cincinnati; in Shermanian and
Edenian rocks of northwestern New York; in the Edenian of south-
ern Pennsylvania; in Edenian rocks of the Ottawa-St. Lawrence
lowland of Canada; in Richmondian strata of Iowa; in Maysvillian
rocks of Wyoming; in Whiterockian strata of Utah; and in Niagaran
rocks of New York.
Genus HOMOCRINUS Hall, 1852
1852. Homocrinus Hall, p. 185 (partim); Hall, 1859, p. 102 (partim); Miller,
1889, p. 255 (partim); Kirk, 1914, p. 476; Ulrich, 1925, p. 94; Moore &
Laudon, p. 145; Moore, 1962, p. 7, text-figs. 1-8; pp. 10, 11, text-figs. 3-4.
1880. Non Homocrinus Hall, Wachsmuth & Springer, p. 77, text-fig. 6; Wachs-
muth & Springer, 1886, p. 144; Bather, 1893, p. 101; Bather, 1900, p. 178;
Wachsmuth, 1900, p. 155; Slocom, 1907, p. 289; Springer, 1913, p. 217.
1900. Non Homocrinus Hall, Wachsmuth, p. 155.
Type species. — Homocrinus parvus Hall, 1852 by subsequent
designation of Meek and Worthen (1866, p. 182).
Diagnosis. — Homocrininae with tall (about twice as tall as
broad and about as tall as the RR), symmetrical, and similarly
shaped BB; with the five rays unbranched; and with proximal
columnals short, of about equal height.
Description. — Specimens belonging to the genus Homocrinus
are minute (height of the dorsal cup is less than two and one-half
millimeters — commonly about one and three-fourths millimeters).
The BB are tall, about one-half the height of the dorsal cup. The
78 BULLETIN 296
A and D ray RR are fused; the B, C, and E ray RR are slightly taller
and compound, divided into iRR and sRR of about equal size. The
anal structure beyond anal X is unknown but is presumably similar
to other homocrinids, cincinnaticrinids, and related forms (1.e., the
anal series is probably an armlike branch off the C ray).
According to Kirk (1914, p. 477), each of the five arms has a
food groove roofed over by an alternating biseries of tiny cover plates.
The [Brr; articulate along their entire proximal surfaces with the
underlying RR and are apparently fixed; they are shaped like in-
verted truncated cones and are shorter than succeeding Brr; the
IBrr; are about as broad as tall. Succeeding IBrr are about twice
or more as tall as broad and shorten somewhat distally; they are
wider at the articulations than at the middles of the plates.
The column tapers rapidly in a distal direction just below the
cup. Kirk (1914, pp. 477-478) related that just distal to the tapering
portion is an area in which two sizes of columnals alternate and
that this alternating portion grades distally into an area with colum-
nals of uniform size. Specimens examined in connection with this
study show a round column that gradually enlarges distally below
the rapidly tapering portion with all columnals observable approxi-
mately equal in size to their neighbors.
Occurrence. —Niagaran. Homocrinus is known from the
Rochester Shale around Lockport, New York, (according to Ringue-
berg, 1888, p. 269 from the top of the lower third of the Rochester
Shale).
Discussion. —Homocrinus was mistakenly thought to be di-
cyclic until Kirk’s (1914) restudy of Homocrinus. Thus, in the
synonymy, all pre-1914 references were to Homocrinus as being
dicyclic; those with partim were with H. parvus (monocyclic) as
type species, while those with non were with a dicyclic type species
(and with H. parvus at most only listed as an included species).
From 1914 to the present, references have been to Homocrinus as
monocyclic with H. parvus as type (and only) species.
Hall (1852, Paleontology of New York, vol. 2) included in his
new genus Homocrinus two new species, 7. parvus (p. 185, pl. 41,
figs. la-c) and H. cylindricus (p. 186, pl. 41, figs. 2a-c, 3a-c), and
two species described by Hall in the first volume of the Paleontology
of New York (1847), Poteriocrinus alternatus (p. 83) and P. gra-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 79
cilis (p. 84). In 1859a Hall added two new species (both dicyclic),
Homocrinus scoparius (p. 102, pl. 1, figs. 1-9) and H. probosctdialis
(p. 138, pl. 84, figs. 24-25).
Hall’s original description of Homocrinus is pertinent and will
be quoted in part (Hall, 1852, p. 185):
Crinoidea having the calyces composed of three series of simple plates,
each series consisting of five plates; sometimes one or more irregular
plates intercalated between the scapular or third series of plates on one
side; arms proceeding from the summit of the third series of plates,
simple or bifurcating, composed of a single series of plates, without
tentacula.
Hall evidently believed that all species he referred to his new
genus had similar plate configurations. The generic description is
clearly intended to apply to dicyclic crinoids (1.¢., with three princi-
pal series of plates; IBB, BB & RR of current usage) with one or
more anal plates (“irregular plates” of Hall, op. cit.) intruded into
the cup. This definition, while loose by modern standards, does
clearly apply to two species which were placed by Hall in Homo-
crinus (H. cylindricus, H. scoparius) but could not accommodate H.
parvus, which, as established by Kirk (of. cit.), is a monocyclic
crinoid with three compound radials. Hall’s description of H. parvus
was based on incomplete material which Hall assumed represented
a dicyclic species. He was only able to deciper a small part of the
calyx plate arrangement (Hall, pl. 41A, fig. 1d); nonetheless his
description assumes three complete circlets of plates (op. cit., p.
185).
In neither of his two papers on Homocrinus did Hall designate
a type species. This practice is characteristic of his earlier work. In
the second volume of the Paleontology of New York (1852), for
example, although numerous new genera (besides Homocrinus) are
described, none is explicitly given a type species. It seems likely that
Hall intended the first species in each to be the type, but this cannot
be demonstrated consistently from his own works. The so-called “first
species rule” (Stoll, et al., 1961, p. 71) is incorporated in the Code,
but only as a recommendation. The first subsequent designation of
a type for Homocrinus has been overlooked in the later literature
but is apparently valid. However, as later workers based their revi-
sions on the incorrect designation the history of the genus will be
reviewed briefly below.
80 BULLETIN 296
A discussion of the Devonian-Mississippian crinoid genus
“Poteriocrinus” (= Poteriocrinites, partim) by Meek & Worthen
(1866, p. 182) contains the following sentence:
Again, if this arrangement of the lowest anal plate excludes it from
Poteriocrinus, how can it, upon such a basis of classification, be referred
to Homocrinus?, the type of which (H. parvus) presents the marked dif-
ference of having the lowest anal piece resting directly down upon the
basal pieces, to say nothing of the wide differences, in the structure of
the arms.
This sentence would seem to qualify as a valid subsequent designa-
tion of a type species under Article 69a, paragraph i of the Code
(Stoll, et al., 1961, p. 69):
In the absence of a prior valid type-designation for a nominal genus,
an author is considered to have designated one of the originally in-
cluded nominal species as type species, if he states that it is the type (or
type-species). for whatever reason, right or wrong, and if it is clear
that he himself accepts it as the type-species.
In this paper, this is accepted as legitimate designation of the type
of Homocrinus.
Wachsmuth and Springer (1880, pp. 77-78) attempted to make
H. scoparius type species of Homocrinus:
The typical specimens which Hall used for description were most un-
satisfactory, that of H. parvus being evidently a very young individual,
while those of H. cylindricus are very imperfectly preserved. In Hall’s
corrected list of New York fossils he seems to have given up both
Dendrocrinus and Homocrinus, as he groups the species of both under
Poteriocrinus [no such reference has been located; indeed, Hall, 1859a,
p. 82, listed H. parvus and H. cylindricus in unaltered fashion]. In
1861 [1859b], however, he described two new species under Homocrinus,
from good specimens. They are not Poteriocrinus, for they have no pin-
nulae, nor Cyathocrinus, for they have an extra intercalated plate above
the basals: nor Dendrocrinus for that plate is not radial; but their af-
finities are the closest with the latter, with which they agree in all
principal characters. We [Wachsmuth and Springer] therefore regard
Homocrinus as a subgenus under Dendrocrinus [dicyclic] . . . with
Homocrinus scoparius Hall [dicyclic] as type... .
Designation of H. scoparius Hall, 1859 as type species of Homo-
crinus Hall, 1852 is not allowable under article 69a of the Code
(Stoll, et al., 1961, p. 69), for H. scoparius is not one of the (four)
originally included nominal species.
In 1889 Miller (p. 255) listed H. parvus as type species of
Homocrinus, possibly based on Meek and Worthen’s statement.
Later, Bather (1893, p. 101),) rejected H. parvus as type species:
There is certainly nothing in the description or figures of H. parvus to
show that it is congeneric with H. cylindricus, and it seems very doubt-
ful to what genus it belongs; it is therefore better to ignore this species,
at all events until it has been properly described, and not take it, as
Mr. S. A. Miller has done, for the type-species of the genus.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 81
Bather then suggested that H. cylindricus be considered type species
of Homocrinus, and referred other dicyclic species to the genus.
It is apparent that, until Kirk’s (1914) revision of Homocrinus,
the genus was considered to be dicyclic, largely because three dif-
ferent species (H. parvus, monocyclic; H. cylindricus, dicyclic; and
H. scoparius, dicyclic) were considered the type by various authors,
and because the monocyclic nature of H. parvus was not known (in
fact, 20 dicyclic species have been referred to Homocrinus, and only
one monocyclic species, H. parvus, has ever been included). Kirk,
applying the convention of page priority, chose H. parvus as type
species (apparently Kirk was not aware of Meek and Worthen’s
work and did not consider Miller’s 1889, listing of H. parvus as the
type species adequate, perhaps because no one after Miller, 1889,
and before Kirk, 1914, e.g., Bather, 1893, had either). He also cor-
rectly demonstrated for the first time the monocyclic nature of H.
parvus. He erected the new dicyclic genus Lastocrinus with H.
scoparius as type species for some of the dicyclic forms formerly re-
ferred to Homocrinus. Kirk berated some authors’ choice (¢.g.,
Wachsmuth and Springer’s, 1880) of a species not included among
those in the original description of the genus for the type and far-
sightedly argued for the need for rules in paleontology to restrict
“... the powers of subsequent writers in revising the original author’s
conception of the genus. . . .” (Kirk, 1914, p. 474). As will be seen
from the above it is doubtful that H. parvus really represents Hall’s
conception of the genus. Nonetheless Kirk’s emendation of the genus
is apparently technically justified. Kirk’s (1914) work caused H.
parvus to be accepted universally as the type species and put an end
to over fifty years of confusion on the nature of Homocrinus parvus.
The diminutiveness of H. parvus has led some crinoid specialists
(e.g., Wachsmuth and Springer, 1880) to view specimens attributed
to H. parvus as juveniles of some other species with a radically dif-
ferent adult form. However, many specimens, all tiny, have been
found; no gradation in morphology away from the common Homo-
crinus parvus form has been observed, and no morphologically rea-
sonable potential adult is known from the same strata. It appears
that adults of H. parvus are minute.
Homocrinus differs from other Homocrininae mainly in having
five unbranched arms and taller BB. As (Kirk, 1914, p. 479) sug-
82 BULLETIN 296
gested, Homocrinus would make a good ancestor for cincinnati-
crinids; it would also be a good ancestor for homocrinids. However,
its age (Niagaran) precludes its being anything but a successor to
known homocrinids and cincinnaticrinids. Possibly Homocrinus was
a precursor of haplocrinitids (Devonian), which have similar struc-
ture in the radial circlet and unbranched arms, and pisocrinids
(Silurian-Devonian), which have modified homocrinid cup struc-
ture and unbranched arms.
Homocrinus parvus Hall, 1852 Pl. 11; Text-fig. 19
1852. Homocrinus parvus Hall, p. 185, pl. 41, figs. 1a-f; Kirk, 1914, pl. 42, figs.
6-7; Meek & Worthen, 1866, p. 182; Kirk, 1914, p. 476, pl. 42, figs. 1-5,
8; Ulrich, 1925, p. 93, text-figs. 10a-b (mislabelled 10a-a); Moore &
Laudon, 1943, pl. 1, figs. 4a-b; Moore & Laudon, 1944, pl. 53, figs. 4a-e;
Springer, 1920, pl. 4, fig. 22 (an illustration of Lecanocrinus nitidus
with H. parvus entangled among its arms); Moore & Laudon, 1944, pl.
SYR vite TP
Primary type material. — AMNH 1705a, b, and c (all from the
Rochester Shale, Lockport, New York) are syntypes of H. parvus.
AMNH 1705a (PI. 11, figs. 1-2) is herein designated lectotype and
1705b and c (PI. 11, figs. 3-4) lectoparatypes of H. parvus Hall,
1852.
Because H. parvus is at present the only known species of
Homocrinus, the specific diagnosis, description, occurrence, and dis-
cussion are the same as for the genus.
Genus ECTENOCRINUS Miller, 1889
1847. Heterocrinus Hall, p. 278 (partim); d’Orbigny, 1850, p. 24 (partim) ;
Pictet, 1857, p. 329 (partim); Billings, 1857, p. 271; Billings, 1859, p.
48; Hall, 1866, p. 4; Hall, 1872, Op. 210 (partim); Meek, 1873, p. 1
(partim) ; Zittel, 1879, p. 358 (partim) ; Wachsmuth & Springer, 1880, p.
68 (partim); Wachsmuth & Springer, 1886, p. 205; Cumings, 1908, p.
713; Miller, 1889, p. 252 (partim); Bather, 1893, p. 25; Wachsmuth,
1900, p. 152; Jaekel, 1902, p. 1100; Grabau & Shimer, 1910, p. 50
(partim) ; Springer, 1913, p. 212; Jaekel, 1918, p. 86; Fritz, 1925, p. 10
(partim).
1886. Stenocrinus Wachsmuth & Springer, p. 207 (partim).
1889. Ectenocrinus Miller, p. 242; Bather, 1900, p. 146, fig. 58-3; Wachsmuth,
1900, p. 152; Cumings, 1908, p. 712; Springer, 1911, p. 26; Springer,
1913, p. 212 (partim) ; Slocum, 1924, p. 337; Ulrich, 1925, p. 94; Moore
& Laudon, 1943, p. 27, text-fig. 3; Moore & Laudon, 1944, p. 145; Moore,
1962, p. 7, text-figs. 1-6, p. 10.
1925. Drymocrinus Ulrich, 1925, p. 96; Moore & Laudon, 1944, p. 145; Moore,
1962, p. 10.
Type species. — Heterocrinus simplex Hall, 1847 by original
designation (Miller, 1889, p. 242).
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 83
Text-fig. 19. Homocrinus parvus.
a—A ray view (after Kirk, 1914); b—exploded diagram (atter Kirk, 1914).
Diagnosis. — Homocrininae with short (about half as tall as
broad) BB; one symmetrically pentagonal B (in the BC interray)
and four asymmetrically pentagonal BB; with five rays bifurcating
isotomously to form ten arms; and with proximal columnals short,
of about equal height.
Descnption. — Ectenocrinus has short, irregularly pentagonal
BB; BB that underlie a compound R and a simple R have one
sloping upper side (under the compound R) and one horizontal
upper side (under the fused R); a single B (in the BC interray)
underlies two compound RR (in the B and C rays) and has two
sloping upper sides. The compound RR (in the B, C, and E rays)
are inverted pentagons, divided into a taller sR and shorter 1R;
compound RR are slightly taller than fused RR, which are tall
rectangles. The distal left corner of the C ray sR and distal right
corner of the D ray R are truncated to accommodate the armlike
anal series. Anal X is an inverted, nearly parallel-sided, pentagon
that supports a series of rapidly tapering distally XX backed by
numerous small polygonal plates to form a tube.
84 BULLETIN 296
Ectenocrinus has two IBrr in each ray. IBr; is a low rectangle
nearly twice as broad as high; it articulates along its entire proximal
surface with the underlying R and is fixed. IBrz is a pentagonal
axillary supporting two arms (to form a total of ten arms) with
alternating heterotomous branching with the first armlets given off
away from the ray.
Occurrence. — Kirkfieldian or Shermanian to Richmondian.
Ectenocrinus is known from the Kope and Fairview Formations
(Edenian and Maysvillian at Cincinnati) in the Ohio-Kentucky-In-
diana tristate area; from the “Trenton limestone,’ Trenton Falls,
New York, Ottawa, Ontario, and Montreal, Quebec; the Maysvillian
of Wyoming; and from the Maquoketa Formation (Richmondian? )
of Iowa. In addition, Ulrich (1925, p. 95) reported a few specimens
from the Curdsville formation (Kirkfieldian?) of central Kentucky,
but this report requires verification.
Discussion. — Ectenocrinus is the genus that for over half a
century was confused with Heterocrinus (see discussion of Cincin-
naticrinus vartbrachialus). Two species, FE. simplex and E. genicula-
ius, are recognized. H. geniculatus is the type species of Drymocrinus
Ulrich, 1925, but the differences between H. geniculatus and E.
simplex appear to be specific rather than generic. Drymocrinus, then,
is a junior synonym of Ectenocrinus. Ectenocrinus may have been
the progenitor, directly or indirectly, of all homocrinids, although
Ibexocrinus or Daedalocrinus could have served this function.
Ectenocrinus simplex (Hall), 1847 Pls. 12-14; Text-fig. 20
1847. Heterocrinus simplex Hall, p. 280, pl. 76, figs. 2a-d; Cumings, 1908, p.
720.
1857. Heterocrinus simplex Hall, Billings, 1857, p. 271; Hall, 1871, pl. 1, figs.
11-12; Hall, 1872, p. 5, figs. 11-12: Meek, 1873, p. 7, pl. 1, figs. 4a-b,
5a-b; Cumings, 1908, p. 720, pl. 4, figs. 10, 10a; Grabau & Shimer, 1910,
p. 502, text-fig. 1814: Moore & Laudon, 1943, pl. 1, figs. 5a-b; Moore
& Laudon, 1944, pl. 53, fies. 8a-b; Moore, 1962, pl. 1, fig. 2a.
1859. Heterocrinus canadensis Billings, p. 48, pl. 4, figs. 5a-d.
1873. Heterocrinus simplex var. grandis Meek, p. 9, pl. 1, figs. 6a-b, 7a-c;
Grabau & Shimer, 1910, p. 502, text-fig. 1814; Moore, 1962, pl. 1, figs.
2b-c.
1909. Ectenocrinus canadensis (Billings), Wood, p. 22.
1914. Ectenocrinus grandis (Meek), Foerste, pv. 124, pl. 1, figs. 8a-d.
1924. Ectenocrinus raymondi, Slocom, p. 337, pl. 29, figs. 5-9; Thomas and
Ladd, 1926, p. 14, pl. 2, fig. 2.
1925. Ectenocrinus simplex (Hall), Ulrich, p. 95, text-fig. 11; Moore & Laudon,
1944, pl. 52, fig. 7.
85
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE
eT
OPANVADNVADUP AES
Text-fig. 20. Exploded diagram of Ectenocrinus simplex (after Ulrich,
1925).
86 BuLueTIN 296
Primary type material. — AMNH 656/2a, b, c, d, e, f, g, and h
are syntypes. AMNH 656/2a (Hall, 1847, pl. 76, figs. 2a and d;
herein Pl. 12, figs. 1-2) is herein designated lectotype and 656/2 b,
c, d, e, f, g and h lectoparatypes of Heterocrinus simplex Hall, 1847.
Diagnosis. — Ectenocrinus having straight arms made up of
numerous syzygial pairs, each pair composed of an armlet-bearing
epizygal above articulating syzygally below with a hypozygal. Cup
subconical, stem facet covering base.
Description. — E. simplex, as well as having the generic (Ecte-
nocrinus) features of two Brr in the [Br series, has two Brr in each
succeeding series. Diagonal sutures, alternating in direction of slope,
separate each division series, with an armlet (pinnule) given off at
the highest part of every second Br (hypozygal). The armlets are not
visible when the arms are folded tightly together, which is commonly
the case (presumably for the same reason that cincinnaticrinacean
arms are usually folded, see cincinnaticrinacean discussion). Young
(small) E. stmplex have tall Brr, while older (larger) individuals
have shorter Brr. Apparently Brr are first secreted as tall quad-
rilateral ossicles which then grow faster laterally than vertically
and so get proportionally shorter.
Distal to the rapidly tapering proximal part (a homocrininan
character), the column of Ectenocrinus simplex shows a columnar
gradation similar in some respects to that of Cincinnaticrinus vari-
brachialus. Just below the rapidly tapering portion of the column,
columnals are short and nearly equal in size to the few adjacent
columnals on either side. The column enlarges gradually distally, and
the section of equal-sized short columnals grades into a zone with
columnals of two different sizes: smaller (shorter and narrower)
columnals alternating with larger (taller and broader) columnals
CGPIE3 ais. 3)).
The column is tripartite with the trimeres of each columnal
disposed in the following manner: one occupies the EA and AB in-
terrays, another lies in the BC interray and the C ray, and the third
occupies the D ray and the DE interray (Text-fig. 20). Each trimere
is in optical continuity; therefore, the trimeres are apparently not
derived from a pentameric condition by fusion of two sets of two
plates (such derivation is obvious in the basal circlets of many
crinoids having only three BB as well as in nearly all blastoids). De-
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 87
rivation from a pentameric condition by fusion of two sets of two
pentameres is also precluded by the unique disposition of the inter-
trimeric sutures. One is interradial (in the CD interray) and two are
radial (in the B and E rays). In monocyclic crinoids with penta-
partite columns, interpetameric sutures are all interradial, whereas
in dicyclic and pseudomonocyclic crinoids with pentapartite columns,
all interpentameric sutures are radial (Warn, 1975). The axial canal
in Ectenocrinus simplex, however, is pentalobate with the lobes
directed interradially (Warn, 1975, text-fig. 3).
Occurrence. — Kirkfieldian or Shermanian to Richmondian. £.
simplex is known from the Kope and Fairview Formations around
Cincinnati; the Trenton Limestone around Ottawa and Montreal
and at Trenton Falls, New York; the Martinsburg Formation of
southern Pennsylvania; and the Maquoketa Formation at Clermont,
Towa. Hall (1847, p. 280) described Heterocrinus simplex from
“... the soft shaly portions of the Blue limestone of Ohio at Cin-
cinnati, equivalent in position to the Hudson-river group of New
York.” Billings (1857, pp. 271-273) described specimens he found
in the “Trenton limestone, Ottawa and Montreal” as H. canadensis,
which is now a junior synonym of E. simplex. Wood (1909, p. 23)
reported E. canadensis (Billings) from the “. . . lower part of Tren-
ton formation [at] Frankfort, Kentucky.” Slocom and Foerste (1924,
pp. 337-339) described E. raymondi, a junior synonym of E. simplex,
from the lower part of the Maquoketa Formation at Clermont, Iowa.
Additionally, numerous good specimens are known from the Kope
Formation around Cincinnati and the Martinsburg Formation of
southern Pennsylvania (especially from Swatara Gap).
Discussion. — Small Ectenocrinus simplex and_lichenocrinid
bases are a common association, and juvenile E. stmplex probably
have a lichenocrinid holdfast. Because Cincinnaticrinus varibracha-
lus and E. simplex usually occur together (in the Kope), nothing
definite can be said of the EF. simplex holdfast. In these occurrences,
however, there are holdfasts that differ from those that can probably
be referred to C. varibrachialus in two respects: they are somewhat
larger (with diameters of about four to five mm as opposed to two
to two and one-half mm) and the plates of the polyplated upper wall
are well demarcated (unlike the C. varibrachialus holdfast which
is obscurely plated).
88 BULLETIN 296
It appears that in adult E. simplex new columnals are added
at the base of the rapidly tapering proximal column and inter-
calated distally, for the smallest (cup height of 1.4 mm) individual
has a rapidly tapering portion, as do all others (the largest has a
cup height of 7.0 mm). A growth zone, similar to that at the base
of the cup in most disparids, some distance below the cup (at the
base of the rapidly tapering portion of the column) is a feature
common to homocrinids and apparently unique among disparids.
Evidently, a trend in homocrinids is to incorporate a few proximal
columnals into the calyx. The interradial lumen extensions suggest
that Ectenocrinus simplex is a true monocyclic crinoid, but the
strange trimeric distribution suggests both monocyclicism and pseu-
domonocyclism.
The taxonomic splitting of EZ. simplex (as shown in the syn-
onymy) was largely due to lack of awareness of population variation
during the classical period of paleontology. Billings (1857, pees
reported that his specimens were conspecific with Heterocrinus
simplex:
I had drawn up the description of our Canadian specimens as above,
under the impression that they were of a species different from that of
the Hudson River Group [H. simplex]. But having since seen Professor
Hall’s collection, I now believe that ours are identical. . . Should,
however, it hereafter be found that ours is different from the Hudson
River species, I beg that it may be called H. Canadensis. .
Hall (1847, p. 280) had incorrectly described the proximal part
of the column of H. simplex as pentagonal (it is round), and Billings
(1859, pp. 48-49) used the Canada specimens’ having round columns
as the differentium between Heterocrinus canadensis and H. simplex.
Meek (1873, pp. 9-10) described H. grandis as a subspecies of H.
simplex; the subspecies was reported to be larger than H. simplex
with shorter Brr than in H. canadensis. Slocom (1924, pp. 337-339)
described Ectenocrinus raymondi as like E. grandis but with shorter
Brr, more slender pinnules, and transverse grooves on the dorsal
sides of the arms. Size of the crown and height of Brr are poor
taxobases, for individuals grow larger and Brr grow faster laterally
than vertically, so that older individuals have proportionally shorter
Brr than younger individuals. The transverse grooves in the single
specimen (CFM UC24701) of E. raymondi may be a unique fea-
ture, but E. raymondi is considered to be conspecific with E. simplex.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 89
E. simplex probably gave rise to £. geniculatus with addition of
a third Br in each division series and geniculation of the arms: E£.
simplex may have given rise to Sygcaulocrinus typus with heighten-
ing of the BB, heightening and fusion of the three most proximal
columnals, and addition of Brr in each division series; and perhaps
E. simplex produced Homocrinus parvus with heightening of the
BB and elimination of branching.
Ectenocrinus geniculatus (Ulrich), 1879 Pl. 14-16
1879. Heterocrinus geniculatus Ulrich, p. 16, pl. 7, figs. 13, 13a-c.
1925. Drymocrinus geniculatus (Ulrich), Ulrich, p. 96, text-figs. 12a-b; Moore
& Laudon, 1944, pl. 52, fig. 7.
1925. Drymocrinus manitoulinensis Foerste, p. 101, pl. 7, figs. 7.
1925. Drymocrinus sp. Foerste, pl. 7, fig. 2
Primary type material.— The holotype (figured by Ulrich,
1879, pl. 7, fig. 13) is UCGM 36313. A natural mold of the holotype
is USNM 42219a. USNM 42219b, c, d, e, f, g, h, i, j, k, and 1 and
CFM UC8829 are paratypes. All are lowest Edenian at Cincinnati.
Diagnosis. — Ectenocrinus with geniculate (zigzag) arms and
two to four IIBr and higher (more commonly two or three than
four).
Description. — E. geniculatus has IBrr, that taper distally and
IBrr2 that expand slightly distally. The [Brr,; are shaped like up-
right, truncated cones. Thus, the junction of the [Brr; and [Brrs
forms a constriction in the crown that marks the position of the
tegmen, above which the arms become free. The IBr and higher
axillaries expand noticably distally. Whereas the armlets in E£.
stmplex are usually concealed when the arms are folded together
(which is usually the case), the armlets in Ectenocrinus geniculatus
are obvious in folded specimens for the zigzag nature of the arms
reveals them.
The column is round and expands gradually distally. Near the
cup, the columnals are short, but they become gradually taller
distally until they are nearly as tall as wide (PI. 15, fig. 6). Ulrich
(1925, p. 96) reported that the column is quinquepartite, but this
has not been verified. The nature of the column is difficult to deter-
mine from specimens in the type suite; the column is probably
pentapartite, as Ulrich said, but may be tripartite, as in E. simplex.
The axial canal is pentalobate with the five lobes directed inter-
radially.
90 BuLLETIN 296
Occurrence. — Edenian. At the base of the Kope Formation in
the immediate vicinity of Cincinnati; Sheguiandah Formation
northeast of Tamarack Point and at St. Hyacinthe in the Mani-
toulin Island (in Lake Huron) area of Canada.
Discussion. — Ulrich (1925, p. 96) described the column of
E. gemculatus as being cirrose and illustrated it (p. 96, text-fig. 12b)
as being profusely so. One of the paratypes (USNM 422191) has
numerous appendages that resemble cirri but are apparently broken
armlets lying along its column (none appear to be attached to the
column). No specimen examined for this report possesses either
cirri or attachment sites for cirri, and the column of Ectenocrinus
gemculatus is evidently not cirrose.
Ulrich (1925, p. 96) also described the anal sac as like that of
E. simplex but wider and with “. . . a series of thin quadrate plates
on either side of the median series.” Available evidence, however,
indicates that EF. geniculatus has an anal sac like that of E. stmplex
(a series of facing XX backed by numerous small polygonal plates).
E. gemculatus was probably a short-lived offshoot from E. stm-
plex that gave rise to no successors. Such evolution would have re-
quired only geniculation of the arms and slight increase in number
of Brr in the IIBr, and higher, division series.
Ectenocrinus sp. indet. Kolata, 1976
1976. Ectenocrinus sp. indet. Kolata, p. 447, pl. 1, figs. 6-7, text-fig. 2.
Primary type material. — A single speciemen, UI X-5184.
Diagnosis. — A species of Ectenocrinus with rotund, pyriform
dorsal cup that is wider than high; anal X arcuate, elongate; com-
pound radials (in B, C, & E rays) with inferradial and superradial
components about equal in height; arms unknown; axial canal of
column and preserved proximal columnals round; column tapering
rapidly distally.
Discussion. — Though the arms are lacking this crinoid was
probably correctly placed in Ectenocrinus by Kolata (1976). The cup
shape is unlike that of E. simplex or E. geniculatus and reminiscent
of that of Apodasmocrinus punctatus. Like the latter E. sp. indet.
has a round column that is apparently not tri- or pentapartite. How-
ever the distal taper of the column and relatively wide column facet
are more comparable to those of the better-known Ectenocrinus
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 91
species. The near-equal size of the inferradial-superradial pairs is a
feature characteristic of Ectenocrinus and far removed from the
strongly unequal compound RR of Apodasmocrinus.
Kolata (1976, p. 448) compared his species with an unnamed
species discussed by Ulrich (1925, p. 95) and cited by him as from
the Curdsville Limestone of Kentucky and the lower Trenton
crinoid beds near Kirkfield, Ontario. The authors have not located
Curdsville Ectenocrinus but the Trentonian specimens from Canada
appear to be E. simplex. Ulrich’s description does not seem to us
applicable to this species, because the dorsal cup of Ulrich’s crinoid
is “more slender, and tapers more gently and more regularly into
the expanding proximal part of the column [in comparison to £.
simplex|” (Ulrich, op. cit., p. 95). The cup of this species is stouter
than F. simplex and the rounded sides cause the change from cup to
column to appear more abrupt than in E. simplex or E. geniculatus,
but less so than in Apodasmocrinus punctatus.
Until better preserved and more complete material is en-
countered the species is best left unnamed, but the generic assign-
ment seems plausible. #. sp. indet. was probably a derivative of
E. simplex that retained primitive cup characters (i.e., size of infer-
and superradials) but had a more evolved column.
Genus Apodasmocrinus Warn and Strimple, new genus
Text-fig. 21
Type spectes.— Apodasmocrinus daubei Warn and Strimple,
1977 by original designation herein.
Diagnosis. — Homocrininae with barrel-shaped dorsal cup hav-
ing a moderate basal concavity; with superradials (in compound
rays )only slightly shorter than simple radials; column round except
in most proximal segment, heteromorphic, with large barrel-shaped
nodals and internodals; proximal columnals much narrower than cup
base; arms 10, apparently uniserial, arms constricted at distal end
of IBr,, expanding above; “pinnules” present, exact arrangement
unknown.
Description. — Crown long, slender, constricted at the summit
of primibrachs 1. Cup barrel-shaped, widest at mid-section of super-
radials or above mid-height of simple radials; base of cup broad,
planate with narrow columnar attachment area impressed into base,
forming a narrow but moderately deep basal concavity; compound
92 BuLLETIN 296
Text-fig. 21. Exploded diagram of Apodasmocrinus daubet.
radials in C, B and E rays (familial characteristic) with short infer-
radials (Text-figure 21); anal X small, resting mainly on diagonal |
left shoulder of C superradial but notching slightly into right
shoulder of D radial. Arms 10, uniserial, long and slender; proximal
end of primibrachs J fills distal faces of radials (or superradials)
but they taper sharply to become narrow at distal ends; axillary
primibrachs 2 expand rapidly distalward; arms do not taper ap-
preciably until well above mid-height. Column round except in
proximal segments which are reported to be composed of five penta-
meres in Apodasmocrinus punctatus; columnals barrel-shaped with
non-cirriferous nodals alternating with much smaller nodals.
Name. — Gr. apodasmos-divided, with reference to the divided
or compound radials.
Occurrence.—Middle Ordovician, Blackriverian; North America
(Va., Tenn., Okla.). The type species is from the Bromide Forma-
tion of Oklahoma. Brower & Veinus (1974) reported A. punctatus
from Benbolt Formation localities in southwestern Virginia and
northeastern Tennessee.
Discussion. — Apodasmocrinus has extremely narrow infer-
radials, perhaps indicating a trend toward eventual elimination,
rather than fusion, of compound RR in one ectenocrinid line. The
anal tube is not preserved on available material but may be much
like that of Ectenocrinus. The arms divide once on the axillary [Brre
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 93
and are pinnulate (bear armlets). Like those of Ectenocrinus the
armlets of Apodasmocrinus are not visible when the arms are tightly
folded. As the arms (main rami) appear completely uniserial and only
a few pinnules are preserved on the paratype of A. dawbez it is not
known with certainty whether the pinnules are alternate on each
brachial or arranged as in E. simplex. The narrowness of each arm
brachial suggests either that syzygial pairs are not developed in this
genus or, less likely, that formerly paired brachials have fused. The
strict uniseriality of the brachials indicates that the former is more
probable.
The barrel-shaped dorsal cup is unusual among Homocrinacea,
being most nearly paralleled in Sygcaulocrinus. The rounded base
and well-developed, though narrow, basal concavity are present
in both species of Apodasmocrinus. No other cincinnaticrinacean
posses both features. Nevertheless, the other features of the
genus individually are found in other Cincinnaticrinacea, and
separation even on a subfamilial level does not appear warranted
at this time. Constriction of the arms above the [Brr, in A. dauben
is reminiscent of a similar trend in Cincinnaticrinus and is also well
exemplified in Dystactocrinus constrictus. Development of penta-
meres involving only the proximal-most columnals in A. punctatus
(Brower and Veinus, 1974, pp. 18-19) indicates that the column is
morphologically advanced. The genus appears to be specialized in
most features (compound RR, column, [Br arm constriction, cup
shape) but retains some archaic features, e.g. uniserial main arms.
Occurrence. — Blackriverian, Benbolt Formation and Hogskin
Member of Lincolnshire Formation or Benbolt Formation, Tennes-
see, and Virginia, Mountain Lake Member, Bromide Formation,
Oklahoma.
Apodasmocrinus daubei Warn & Strimple, new species Text-fig. 22 a-c
Primary type material.— Holotype SUI 39593 (Repository,
University of Iowa); paratype USNM 164106.
Diagnosis. — Base of cup broad with large basals flexed upward
to form an appreciable portion of lateral sides of cup. Cup plates
tumid and smooth. Occasional pinnule-like ramules in distal por-
tions of arms.
Description. — Same as that of genus, except where noted (see
BuLLeETIN 296
94
w asl
Qo
22. Apodasmocrinus. a — CD interray view of A. daubei, para-
type USNM 164106. b — A ray view of A. daubei,
paratype USNM 164106.
Text-fig.
c — Holotype of A. daubei (SUI 39593); drawing centered on CD interray.
d — Drawing of holotype of A. punctatus (USNM 164097), CD interray view.
1974), pl. 2, cig, 2.
(
Adopted from Brower & Veinus
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 95
above and discussion sections under both Apodasmocrinus species).
Discussion. — A. daubei and A. punctatus are similar in overall
appearance but the punctate plate ornament of the latter and the
proportionately taller basals of A. dawbez readily distinguish the two
species. Additionally A. dawbei has a broader stem facet (compared
to maximum cup width) than does the Benbolt species, and its
column appears to lack a differentiated quinquepartite proximal
portion even in young specimens (¢.g., the holotype). A. punctatus
has basal plates with only the distal tips flexing out of the basal
plane which contributes to formation of a proportionately shorter
dorsal cup than that of A. daubei. In most other respects the two
species are remarkably similar.
Measurements in millimeters:
Holotype Paratype
SUI 39595 USNM 164106
Length of crown (excluding basals) — 31.4
Width of crown (at secundibrachs 2) — 6.7
Height of cup 4.0 —
Width of cup (maximum) 4.2 523
Width of cup (antero-posterior ) — 5x2.
Height of anal X L3 1.5
Width of anal X 12 1.4
Height of D radial 22 25
Width of D radial 2.3 2.6
Height of C’ superradial zl Wd}?
Width of C superradial jp) 2.8
Height of C inferradial 0.7 —
Length of C inferradial lateral sides 0.4 —
Name. — Particular mention is made here of the kindness of
Leon Daube who first allowed the junior author permission to col-
lect on his ranch and to the later cooperation of his heirs, Mrs. Olive
Daube and son Sam Daube, and to Jim Manton, manager of the
Daube Ranch Company. It is with this in mind that the presently
described species is named daubez in slight token of gratitude to
Leon Daube.
Occurrence. — “Platycystites zone,’ Mountain Lake Member,
Bromide Formation, Blackriveran, Middle Ordovician; West Branch
96 BULLETIN 296
of Sycamore Creek, Daube Ranch, Johnson County, Oklahoma (SW
1/4 SE 1/4 NW 1/4 sec. 27, T.3 S., R. 4 E.).
Apodasmocrinus punctatus (Brower & Veinus), 1974 Text-fig. 22 d
1974. Ectenocrinus punctatus Brower & Veinus, pp. 17-20, pl. 1, figs. 2-4; pl. 2,
figs. 1-6.
Primary type material. — Holotype, USNM 164097; paratypes,
USNM 164098-164105; UMMP 57521, 57522; MCZ 621.
Diagnosis.— A species of Apodasmocrinus with BB barely
visible in side view; cup short, broad, barrel-shaped in young indi-
viduals but somewhat quadrate appearing in mature specimens.
Column round, possibly proximally quinquepartite; distally strongly
heteromorphic; column facet narrow.
Discussion. — The original description by Brower and Venus
(op. cit., pp. 17-20) is complete and needs no supplementation.
Though no specimen with arms has been found, the cup shape,
narrow inferradials, and column features suggests referral to Apo-
dasmocrinus. The proportionately narrower stem facet of this species
is likely an advanced feature, as is the reduction in size and promi-
nence of the BB; however, the proximal portion of the column of A.
punctatus still is quinquepartite, indicating a closer relationship
for this species to its probable ectenocrinid ancestors. Possibly both
species are descended from a common ancestor which itself had
earlier diverged from Ectenocrinus. The most likely antecedent for
both Apodasmocrinus species is a form like Ectenocrinus sp. indet.
Kolata.
Genus IBEXOCRINUS Lane, 1970
1970. Ibexocrinus Lane, p. 12.
Type species. — Ibexocrinus lepton Lane, 1970 by original desig-
nation (p. 12).
Diagnosis. — Homocrininae with symmetrically pentagonal BB
about as tall as wide and equal in size; with five rays bifurcating
isotomously to form ten arms; and with proximal columnals narrow,
of about equal height.
Description. — Ibexocrinus has compound RR divided about
equally into i1RR and sRR. The anal tube, aside from the first two
XX, is unknown but is probably like that of Ectenocrinus. Each ray
apparently has two [Brr, with isotomous branching on the [Brro.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 97
IBr, is a low rectangle nearly twice as broad as high, articulating
with the underlying R along its entire distal surface. IBr2 is a
pentagonal axillary supporting two equal-sized arms. Succeeding
branching is alternating heterotomous with the first armlets on the
outside (or abradially). In subsequent arm divisions there are six
to nine I] Brr branchials in each division series. The stem is round
and expands only slightly proximally just below the dorsal cup; it is
pentapartite with radial pentameres.
Occurrence. — Whiterockian. [bexocrinus is known from a single
specimen from the M zone (of Hintze, 1951), Kanosh Shale, near
Ibex, Utah.
Ibexccrinus lepton Lane, 1970 Pl. 16, figs. 46; Text-fig. 23
1970. Ibexocrinus lepton Lane, p. 13; p. 8, text-figs. 2b-c; p. 11, pl. 1, fig. 1.
Primary type material. — The holotype and only known speci-
men of [bexocrinus lepton Lane, 1970 is USNM 165239.
Because J. lepton is presently the only known species of [bexo-
crinus, the specific diagnosis, description, occurrence, and discussion
are the same as for the genus.
Genus SYGCAULOCRINUS Ulrich, 1925
1925. Sygcaulocrinus Ulrich, p. 98; Moore & Laudon, 1944, p. 145; Moore,
1962, p. 11, text-figs. 3-7a-b (b is Ulrich, 1925, p. 93, text-fig. 10b).
Type species. —Sygcaulocrinus typus Ulrich, 1925, by original
designation (Ulrich, 1925, p. 99).
Diagnosis. — Homocrininae with tall BB, about one and one-
half times as tall as broad; with one symmetrically pentagonal B
(in the BC interray) and four asymmetrically pentagonal BB; with
five rays bifurcating isotomously to form ten arms; and with proxi-
mal columnals inflated, greatly taller than adjacent (more distal)
columnals.
Description. — Sygcaulocrinus has irregularly pentagonal BB;
those BB that underlie a compound R and a simple R have one
steeply sloping upper side (under the compound R) and one nearly
horizontal upper side (under the fused R). A single B (in the BC
interray) underlies two compound RR and has two sloping upper
sides. The compound RR (in the B, C, and E rays) are inverted
pentagons, divided into a taller sR and a shorter iR; compound RR
BULLETIN 296
98
MATT
al tmaun nastie
HT
oy
-fig. 23. Ibexocrinus lepton.
USNM 165249 (both after Lane, 1970).
A ray view.
b
a—CD interray view.
Text
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 99
are taller than fused RR. They are unlike Homocrinus and Ibexo-
crinus, which have the proximal points of the RR even (at the same
level of the cup) and the distal edges of the compound RR higher.
They are also unlike the compound RR of Ectenocrinus, which have
the distal edges higher and the proximal points lower than those of
the fused RR. The distal edges of all RR in Sygcaulocrinus are even,
and the proximal points of the compound RR are lower than the
proximal points of the simple RR. Only anal X of the anal series
is known; it is an inverted, distally tapering pentagon inserted into
the notch formed by the truncated shoulders of the C and D ray
RR.
Sygcaulocrinus has two [Brr in each ray. IBr; articulates along
its entire proximal surface with the underlying R and tapers some-
what distally. IBr2 is a pentagonal axillary supporting two arms.
Branching and number of Brr per division series beyond this isoto-
mous division is unknown except for USNM 89876 (the holotype
of S. typus).
The most distinctive feature of Sygcaulocrinus is the “ex-
ploded” nature of the proximal columnals. The most proximal
columnals (usually three) are wider and higher than distally adja-
cent columnals. As in other Homocrininae, this proximal portion
of the column tapers distally. The column is evidently round, al-
though it is unknown beyond (distal to) the first five or six most
proximal columnals.
Occurrence. — Richmondian. Maquoketa Formation from Fort
Atkinson, Iowa.
Discussion. — Ulrich (1925, pp. 98-99) described and illustrated
a number of features for Sygcaulocrinus that cannot be verified from
known specimens: 1) a tripartite column, 2) alternating hetero-
tomous branching, 3) three to six I[Brr and higher, and 4) a tiny
anal X lying in a similar-sized notch at the junction of the C and D
ray RR. The authors have been unable to establish the tripartite
nature of the column. Only the holotype exhibits branching or num-
ber of Brr beyond the [Br axillaries, but number of [[Brr and
branching pattern is difficult to determine from this specimen. How-
ever, anal X and the proximal column have been observed in a num-
ber of specimens. Anal X, and the notch formed by the truncated
corners of the C and D ray RR, seem to be larger than Ulrich re-
100 BULLETIN 296
ported (Strimple, 1974, p. 116). Ulrich probably described anal X as
minute because anal X of the holotype has been rotated and only
the northeast corner of the plate juts through sediment enclosing it.
Strimple (of. cit.) regarded this crinoid as a bottom-dweller which
autotomizes a portion of the column at some point during growth.
Syscaulocrinus typus Ulrich, 1925 Pl. 17; Text-fig. 24
1925. Sygcaulocrinus typus Ulrich, p. 90, text-figs. 10a-b (? mislabelled 10b-b) ;
Moore & Laudon, 1944, pl. 52, fig. 7.
1926. Ectenocrinus eclongatus Thomas & Ladd, p. 12, pl. 2, figs. 3-8, pl. 5, figs.
3-4.
Primary type material. — The holotype of S. typus Ulrich, 1925
is USNM 89876.
Because S. typus is presently the only known species of Sygcau-
locrinus, the specific diagnosis, description, occurrence, and discus-
sion are the same as for the genus.
Subfamily DAEDALOCRININAE, new subfamily
Diagnosis. — Homocrinidae with the dorsal cup made up of
strongly interlocking RR; with the compound RR somewhat taller
than the fused RR, except for the B ray R, which, although com-
pound, is the same height as the fused RR; with five rays bifurcating
isotomously to form ten arms, after which branching is endotomous;
and with a pentagonal column without a proximal tapering portion.
Because this is a monogeneric subfamily, other features are dis-
cussed under Daedalocrinus.
Genus DAEDALOCRINUS Ulrich, 1925
Text-fig. 25
1925. Daedalocrinus Ulrich, p. 97; Moore & Laudon, 1944, p. 145; Moore,
1962, p. 10.
Type species. — Daedalocrinus kirki Ulrich, 1925 by original
designation (p. 97); this species is considered a junior subjective
synonym of Heterocrinus bellevillensis Billings, 1883 herein.
Description. — Daedalocrinus has equi-dimensional, symmetri-
cally pentagonal BB of about equal size. The arms are long and have
numerous branches; each arm has as many as ten armlets. Armlets
are unbranched and extend to the arm tips. The genus has three to
five [Brr per ray (apparently four is most common). Like members
OrRDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 101
be
Text-fig. 24. Sygcaulocrinus typus.
a—exploded diagram (after Ulrich, 1925); b—camera lucida drawing of
USNM 89876, D ray view; c—D ray view (after Ulrich, 1925).
of the Cincinnaticrininae, number of Brr in each division series ap-
pears to be variable, both among different rays in single individuals
and among the same rays in different individuals. Branching beyond
the axillary [Brr is variable, with armlets given off anywhere from
every third to seventh Br; a single specimen might have an arm
with five I]Brr, three I]]Brr, and seven IVBrr.
The column, like that of members of the Cincinnaticrininae, is
pentapartite with radially disposed pentameres, has a pentagonal
102 BuLLeTIN 296
Text-fig. 25. Daedalocrinus bellevillensis.
a—exploded diagram of UCGM K.3669a; b—exploded diagram of UCGM
K.3669b; c—exploded diagram of the posterior side of the proximal] part of the
crown of USNM S.2141 (lectotype of Daedalocrinus kirki, a junior synonym
of Heterocrinus bellevillensis,
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE _ 103
lumen with interradial angles, and is pentagonal proximally, with
gradation distally from pentagonal to round.
Occurrence. — Kirkfieldian. Daedalocrinus is known from the
Hull crinoid beds of Belleville and Kirkfield, Ontario. Billings (1883,
p- 50) described Heterocrinus bellevillensis from the “Trenton lime-
stone” at Belleville, Ontario, (= Hull beds); Ulrich (1925, p. 97)
reported its, and another species’ (D. kirkt, considered a junior
synonym of D. bellevillensis), occurrence in the “Lower Trenton
crinoid beds,” Kirkfield, Ontario (= Hull crinoid beds at Kirkfield,
Ontario, where it is evidently fairly common).
Discussion. — Daedalocrinus was erected by Ulrich (1925, p.
97) for inadunates with a conical cup with three compound and two
fused RR and with ten arms branching endotomously. The latter
feature had previously been noted by Billings (1883, p. 50) for H.
bellevillensis. Ulrich (1925) placed Daedalocrinus in the Homo-
crinidae.
In some respects, Daedalocrinus resembles certain crinoids not
referable to the Homocrinidae. It has endotomous branching like
Geraocrinus, an anomalocrinid, and a column and variable number
of Brr per division series like members of the Cincinnaticrininae.
Springer (1911, p. 27) reported that Kirkfield material in the
United States National Museum collection makes it evident that
Heterocrinus bellevillensis has a convoluted anal sac, which would
confirm its referral (by Springer) to Ohiocrinus. Ulrich (1925, pp.
97-98), using the same material as Springer had, described the anal
sac as large and balloon-shaped. The authors have examined a num-
ber of well-preserved specimens in the Kopf collection (at the Uni-
versity of Cincinnati) and have perused the USNM cincinnaticrinids
and homocrinids but have not found evidence to corroborate
Springer’s or Ulrich’s observations. However, only one of the at least
12 syntypes of Daedalocrinus kirki has been located. A note (pos-
sibly in Springer’s or Ulrich’s handwriting) accompanying USNM
S.2141 lists 12 specimens collected from Kirkfield, Ontario, in 1905
by Edwin Kirk. However, M. W. Moodey added the comment (dated
March 16, 1934) that she “located only what is in this tray [USNM
S.2141].” Unless the missing syntypes or better topotype material
are discovered it seems best to maintain Daedalocrinus Ulrich with
the cited type species. Certainly the morphology of the existing
104 BuLLETIN 296
material confirms Ulrich’s descriptions of the cup, arms, and column,
and renders Springer’s referral unacceptable.
LOCALITIES
Localities of Cincinnaticrinus varibrachialus collected in connec-
tion with this study. Numbers refer to Text-figure 10. All were in
the Kope Formation. Crinoid remains in all but locality 2 were de-
posited as large ripples.
1—N39 06’, W84 24’ at the base of an old road cut on the east side of Elstun
Avenue, 75 yards (nearly 70 meters) south of Beechmont Avenue. Neither
the Kope-Fairview nor the Kope-Point Pleasant contact is visible. Fossil
content and elevation of the outcrop suggest occurrence in the Southgate
member (Text-fig. 4). Crinoids were found in a north-south trending de-
posit about 12 feet by 3 feet (nearly 4 meters by 1 meter) in areal extent
and about 1/2 inch to 4 inches (about 1 1/4 to 10 centimeters) thick. The
deposit was in mudstone and consisted mostly of column fragments, at least
88 tiny Cincinnaticrinus varibrachialus crowns and calyces, 7 tiny Ecteno-
crinus simplex crowns, 72 juvenile holdfasts, trilobites, brachiopods, gastro-
pods, trepostome Bryozoa, and small Mesopaleaster. The trilobites and star-
fish were apparently scavenging the dead crinoids before burial. Many
(all but the 95 cited above) of the crinoids are presently in the hands of
amateur collectors.
2—N39 10’, W84 34’ in the west bank of West Fork Creek 50 yards (about 46
meters) northwest of the intersection of Diehl Road and West Fork Road,
198 feet (about 60 meters) below the Kope-Fairview contact which is
visible along Shepherd Road just west of West Fork Road. Crinoids were
found in mudstone and occupied about one square yard (nearly one square
meter). Long (up to two feet, about 0.6 meters), unbroken columns lay with-
out consistent orientation. Eighteen Cimncinnaticrinus varibrachialus, 17
Ectenocrinus simplex, and 38 juvenile holdfasts were found.
3—N38 39’, W85 07’ near the top of a large outcrop at the southeast corner of
the Carrollton, Kentucky I-71 interchange, 30 to 50 feet (about 9 to 15
meters) below the Kope-Fairview contact. The contact is covered here, but
it is visible in outcrops along I-71 on the other side of the Kentucky River.
The west-northwest—east-southeast trending deposit was about 3 feet by 6
inches in area and 2 inches thick (about 92 * 15 5 centimeters) and con-
sisted of 17 C. varibrachialus, 2 E. simplex, and numerous columns.
4—N38 56’30’, W84 50’ in the creek bed of an unnamed tributary (of the Ohio
River) which flows through Rabbit Hash, Kentucky, 70 feet (about 21
meters) below the Kope-Fairview contact which occurs in the westernmost
fork of the tributary about 250 yards (about 80 meters) upstream from the
pocket. The pocket consisted of “knotted columns” striking approximately
east-west, 82 Cincinnaticrinus varibrachialus crowns and calyces, 54 Ecteno-
crinus simplex, and 21 juvenile holdfasts in mudstone, becoming more limy
northward, and finally grading into a biogenic limestone made up in large
part of discrete columnals.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 105
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PLATES
114 BULLETIN 296
EXPLANATION OF PLATE 1
Unrecognizable species.
Figure Page
1-2. Heterocrinus heterodactylus Hall 2000... 40, 47
Lectotype (designated herein) and latex cast of same, AMNH
1116/3, from Snake Hill, Saratoga County, N.Y., X1.7.
3-6. Heterocrinus heterodactylus Hall ...o.........000cccccccccceeeeeeeeees 40, 47
Lectoparatype, AMNH 1116/2, from Boonville, N.Y., three lecto-
types, AMNH 1116/1, from Pulaski, N.Y., X0.6.
7-S. Heterocrinus exilis Hall .....:.05<..c.00cc.c0c..00eqeossouenticere ee 53
Holotype, AMNH 1176/1, from Cincinnati, Ohio, view from CD
interray, X0.8 and 2.5.
PLATE 1
BULL. AMER. PALEONT., VOL. 72
PLATE 2
BULL. AMER. PALEONT., VOL. 72
SE eae
reser eS t.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE- 115
EXPLANATION OF PLATE 2
Unrecognizable species (figs. 4-8).
Figure Page
iS PATOpOCrINUS priscus Liane Bios ee. Ae En 74
Holotype, USNM 165240, from Ibex, Utah, E ray view; C ray
view photographed under ethanol, C ray view, X2.1.
APMHIGIGFOCHINUS .GHACIIES Flall uo... .coce..ctscceeesntesccenes cesdteene ocsenorocesonas 38
Latex cast of holotype, AMNH 1117, from Snake Hill, Saratoga
County, NeYen Xx 157:
FeeHeTerocrinUs jUVeNis Elall) 8 oo. scrcscccsccscsesoccacesccececenorezoessnsess 39, 56
Supposed holotype, AMNH 1173/1, purportedly — Lebanon,
Ohio, D ray view, 3.8.
Gee COlUMDBICKINUS) CrasSUS UIICI aren cee eee eneecete esas oeeeeseee sare 31
Holotype, USNM 89826, A ray view and CD interray view, from
Columbia, Tenn., X2.1.
SMEONIOCFINUS @XiliS: FOCTSUC. -....cciccccecconc codecs cetn sesteer ssisadoue sosgscegees 60
Holotype, USNM 78718, from Rogers Gap, Ky., 2.1.
116 BULLETIN 296
EXPLANATION OF PLATE 3
Cincinnaticrinus varibrachialus, n. sp.
Figure Page
1-2. Cincinnaticrinus varibrachialus, n. sp. oo... 41
Holotype, UCGM 3871, specimen figured by Meek (1873, pl. 1
figs. la-b) as Heterocrinus heterodactylus, from Kope Forma-
tion, Cincinnati, Ohio, C ray view, X0.8 and 2.5.
3) Gx waribrachialus: 2....4.03:0808s 5 orto eee 41
Figured specimen, MU 959a, from Kope Formation, Cincinnati,
Ohios <255:
4:5. GC. waribrachialus.¢....0-.-.205 See fons. aiscde teed. aoe 41
Paratype, UCGM 405751, from locality 4, AB interray view with
tegmen visible just over IBrr; as well as anal back plates on
anal X (see Text-fig. 8), X3.0 and oral view, 6.0.
6: 5C. varibrachialus |. <...5<::....sc2. etek dita egos... aoe 41
Illustrated specimen, UCGM 40580, from localicy - is cp interray
view, X3.8.
Hee Goavarl DFACHIAIUS Fe 5 eee oc eiceveaten’:
Illustrated specimen, UCGM 42674, from Kope oration New-
port, Ky., 3.4
Gee GC.owaribrachialyg) 2... 2.02. 2ciscy ss. ses.scsedoenseosscessese er 41
Illustrated specimen, UCGM 36287, from Krenton Falls, N.Y.,
D255
9. C. varibrachialus
Illustrated specimen, UCGM 40580, CD interray view, 11.5.
OD PCAC VANIDTACKIALUS. «...5:iisiccch< csc Bono tate sere ee 41
Illustrated specimen. UCGM 6562, Kope Formation, Rapid Run
Creek, Cincinnati, Ohio, X1.7.
Deco WAriDrMchialUs: «0: :.:.0sc:ceepeen ee eee, Bion. ne 41
Illustrated specimen, UCGM 2021a, Kope Formation, Cincinnati,
Ohio, A ray view, X2.5.
PLATE 3
BULL. AMER. PALEONT., VOL. 72
PLATE 4
BuLu. AMER. PALEONT., VOL. 72
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE~ 117
EXPLANATION OF PLATE 4
Paratypes of Cincinnaticrinus varibrachialus, n. sp.
from locality 5, all 2.5
Figure Page
1-10. Cincinnaticrinus varibrachialus, n. sp. ....................... ieee CL
Paratypes (1) UCGM 40555, C ray view; UCGM 40497, (2) C
ray view and (3) E ray view; (4) UCGM 40555, E ray view;
UCGM 40500, (5) H ray view and (6) CD interray view;
(7) UCGM 50502, E ray view; UCGM 40531, (8) A ray
view, (9) CD interray view photographed under ethanol, (10)
and E ray view.
118 BULLETIN 296
EXPLANATION OF PLATE 5
Cincinnaticrinus varibrachialus, n. sp. photographed under ethanol.
Figure Page
1-9. Cincinnaticrinus varibrachialus, n. sp. oo... 41
Illustrated specimens, (1-2), UCGM 40579, from locality 1 (see
Text-fig. 8), A ray view and D ray view; (3-4) UCGM 40580,
from locality 1, B ray view and DE interray view; (5-6),
UCGM, from locality 1, A ray view and D ray view; (7-8),
UCGM, from locality 1, A ray view and CD interray view;
(9), UCGM 40583 BU, from locality 1, B ray view, all 7.6.
10-13. C. varibrachialus, n. sp. ........ eT 41
Illustrated specimens, (10-11), UCGM 40568, from locality 4,
C ray view and E ray view; (12-13), UCGM 40569, from
locality 4, B ray view and DE interray view, all 4.7.
BULL. AMER. PALEONT., VOL. 72 PLATE 5
PLATE 6
S Magic - pie PAs He Pie & rh Ss. ch
eanreteitreriretts
oP ep?’ i
See ee,
sf i Te lew
BuLL. AMER. PALEONT., VOL. 72
2-4.
6-7.
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE- 119
EXPLANATION OF PLATE 6
Cincinnaticrinus pentagonus (Ulrich)
Cincinnaticrinus pentagonus (Ulrich) 2.00. 55
Lectotype of Heterocrinus pentagonus Ulrich, YPM 24801, from
Fairview Formation, Cincinnati, Ohio, EA interray view, X1.7.
CoO TAGOMUS: ccc eee ee rises ina-sdehogincng: sneer cane 55
Illustrated specimens, (1), UCGM 17626, from Grant Lake
Formation?, Lebanon, Ohio, BC interray view; (3), UCGM
6559a, from Bull Fork Formation, Westwood, Cincinnati,
Ohio, A ray view; (4) UCGM 11609, from Fairview Forma-
tion, Fairview Heights, Cincinnati, Ohio, CD interray view, all
SOIT:
GCEP DEMTAGONUS sss cccc see ee eee ot aoe cans ca os aos on eae enone 55
Lectoparatype, YPM 24892, from Fairview Formation, Cincin-
nati, Ohio, CD interray view, X1.7.
ME PUES RUE COTM S ods 0.05, Dae eee ease eee vi vote Sea donee cvs cov cena nee 55
Illustrated specimens, (6) UCGM 41501, from Bull Park Forma-
tion, Hueston Woods State Park, Ohio, X0.8; (7) UCGM
6450c, from Bull Park Formation, Clarksville, Ohio, 1.7.
120
Figure
5-7.
9.
BULLETIN 296
EXPLANATION OF PLATE 7
Dystactocrinus constrictus (Hall) ........20...-......ceeeecceeseser eet
Illustrated specimens, (1-2), UCGM 42675, from Cincimat
Ohio, B ray view and A ray view; (3) UCGM 6542, from
Cincinnati, Ohio, A ray view; (4), UCGM 6424, Cincinnati,
Ohio, all 1.3.
DivCOnstTrichus) eo oikoas nc sorted. See
Holotype, HMCZ 2165, Con ‘ake Formation, Cincinnati, Ohio,
B ray view, C ray view and DE interray view, X1.7.
Daiconstrictusy) 20 oe ee. eee eee
Illustrated specimen, UCGM 42676, Grant Lake Formation, Ft.
Mitchell, Ky., A ray view, X0.8.
BD; -COnsinrictus:: (See. 6.2 le ee
Holotype of Atyphocrinus corryvillensis (a junior synonym of
D. constrictus), Grant Lake Formation, Cincinnati, Ohio, X1.3;
USNM 89827.
PLATE 7
BuLu. AMER. PALEONT., VOL. 72
PLATE 8
BULL. AMER. PALEONT., VOL. 72
abe sais ,
2 dn yg ER AL
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 121
EXPLANATION OF PLATE 8
Figure Page
leelsotomocrinus teMmuis: (Billings) eae cscee seco ec esc de cee cdeeencs eves 63
Holotype, USNM S.2077a, Isotomocrinus typus Ulrich, a junior
synonym of J. tenuis, from Kirkfield, Ontario, D ray view,
4.7.
TAMER TER) LTISIa tera) Mie. A ee Wl net nem: SURE BPSD oa ok 63
Paratypes, (2) USNM S.2077b, cD interray view, (3) §S.2077c,
AB interray view, 4.7, (4) USNM S.2077a-c, X1.3, all J.
typus from Kirkfield, Ontario.
BIE GREMOIS I. 2. iS ces). 00.1. SED oR OD oe fees REA cos seen! 63
Lectotype, GSC 1438, from Ottawa, Ontario, EA interray view,
XZ.5.
Goiire tOMUNS | 1286). ie Se ree AS. epee aes seat. 0 See . 63
Illustrated specimens, UCGM K.42677, AB interray view, UCGM
K.42678, CD interray view, from Kirkfield, Ontario, NL. 3.
122 BULLETIN 296
EXPLANATION OF PLATE 9
Figure Page
1. Ohiocrinus laxus (Hall) ............... «do Seiiwisgac bbws sosek seca oss ee 68
Illustrated specimen, UCGM 23048, B ray view, 1.3.
DP OMIGCHINUS SD. . <<:..../-2-c.ceoe ee ecne ds eeeee scree eds onan <ce 67
Illustrated specimen, UCGM 6600, Pauviow Formation, Madison,
nde <1-3.
SAP Ol Laxusy (Ea reese eae soso ree eee aoe cena ones er 68
Illustrated specimen, USGM 6545a, Grant Lake Formation,
Orland Ave., Cincinnati, Ohio, CD interray view, 1.3, and
same X3.8.
5. 0. laxus (Hall)
Holotype, HMCZ 2167, CD interray view, Cincinnati, Ohio, 1.7.
6:82) OF laxus; (Halle ee Been ccssch src coors cas ase 68
Illustrated specimen, UCGM 6524a, Cincinnati, Ohio, (6) CD
interray view, X1.7, (7) same 3.8, showing spiral anal sac
between arms of C and D rays, (8) same, X3.8, showing spiral
anal sac between arms of 5 and A rays.
O10: “Oslaxus. (lall) 22506. ok ein inte a 68
Lectotype of Heterocrinus oehanus a junior synonym of O. laxus,
USNM 42304a, Fairview Formation, Cincinnati, Ohio, 1.3
and X2.5.
PLATE 9
BULL. AMER. PALEONT., VOL. 72
PNP. bei EY
PLATE 10
BULL. AMER. PALEONT., VOL. 72
——- aap PF
—
i ee roo
—"
er ine
posscitet Mame .
a ma .
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 123
EXPLANATION OF PLATE 10
Figure Page
Nese ONIOCHIMUS DraUriiy, WLPICH eegeee ce ose eas. Son cen oteeeerotecceeertateeseccs 70
Lectotype, USNM S.2082b, Fairview Romaation Madison, Ind.,
CD interray view, 3.8, photographed under ethanol, X6.8
and X13.6.
AS Os, LESTE TI ONS Os Vee en se cee Cee ace eee esa eee 70
Lectoparatype, USNM S.2082a, Fairview Formation, Neder:
Ind., E ray view, X3.8, photographed under ethanol, XK 6.8 and
LL:
124 BuLLETIN 296
EXPLANATION OF PLATE 11
Figure
1-2: Homocrinus parvus Hall ..................204..200-2.2
Holotype, AMNH 1705a, Lockport, N.Y., X1.7 and 3.0.
SA AD parves Hall 5. occcccccccccccsencen sez --vosessvessnaaoceceta-aee se eee
Paratypes, AMNH 1705b-c, Lockport, N.Y., X1.7 and X30.
B,D woes BEAU so. ccs bec ec cse apn etic ca haces Sete ees a
Illustrated specimen, UCGM K.36292, Lockport, N.Y., 1.7.
BULL. AMER. PALEONT., VOL. 72 PLATE
Oh yo tm, >
: Tre rrges re 0
¢
PLATE 12
BULL. AMER. PALEONT., VOL. 72
iy eos
$y ra. -
a> oe
Ria Si re siinwonga, 5”
’ cic A aii :
con * arena
ORDOVICIAN-SILURIAN CrrnoIDS: WARN AND STRIMPLE- 125
EXPLANATION OF PLATE 12
All figures magnified 1.3
Figure Page
ERE CTENOCHINUS. SIMMPIEX (eral) err es eee cece eic occ crack ceseenen anes 84
Holotype, AMNH 656/2a, Cincinnati, Ohio, A ray view and CD
interray view.
3-10. E. simplex (Hall) .......... oR Me Rene Meh) eno ae EN 84
Paratypes from Cincinnati, Ohio, (3), AMNH 656/2b, CO inter-
ray view; (4) AMNH 656/2c, C ray view; (5) AMNH
656/2d, DE interray view; (6) AMNH 656/2e, B ray view;
(7-8) AMNH 656/2f, CD interray view and DE interray
view; (9-10), AMNH 656/2g, BC interray view, CD interray
view.
t-i3-) Es simplex: (Hall): ..3.....cgjess0...cp ce corse ceecensecseo doo eS RRREE eb <A 84
Figured specimen, SUI 3770, Maquoketa Formation, Clermont,
Iowa, E ray view, A ray view, BC interray view.
ae Siri lexy (EVAL) sci... eae reese eee ae ce foc. be Bae tate te Soe oct esas Ree 84
Paratype, AMNH 656/21, Cincinnati, ‘Onion AB interray view.
TREN. LSM T=) 32( GE E11) a eae ne Pree poate ee eee cee 84
Holotype of Ectenocrinus raymondi Slocum, junior synonym of E£.
simplex, Maquoketa Formation, Clermont, Iowa, BC interray
view, E ray view and CD interray view.
126 BuLueTin 296
EXPLANATION OF PLATE 13
Figure Page
1-3. Ectenocrinus simplex (Hall) ...............0....:-:¢cc0esss aceon 84
Illustrated specimens, (1) UCGM 36281a, Trenton Falls, Nays.
CD interray view, X1.3; (2) UCGM 36281b, Trenton Falls,
N.Y., C ray view, X1.3; (3) UCGM 42679, x<0.4.
4-5. E. simplex (Hall) .. A OE isons 86
Illustrated specimen, ulstes cast, “UCGM. 42680, Martinsburg
Formation, Swatara Gap, Pennsylv ania, A ray view and CD
interray view, X1.3.
6. E:simplex:(iall) i: ee a ee. eee 84
Illustrated specimen, UCGM 42681a, locality 2, BC interray view,
Seek
7..-E.-simplex (lal): .ccis00.c.2 ee sees. eee eee 87
Illustrated specimen, latex cast, Derstler no. 1a, Martinsburg
Formation, Swatara Gap, Pennsylvania, BC interray view,
eels
$-9./-E. simplex(Hall) 2.2) 2taneian.t..5 266. eee 87
Illustrated specimen, latex cast, Derstler no. 1b, Martinsburg ;
Formation, Swatara Gap, Pennsylvania, E ray view showing
interior surfaces of B and C radials and E ray view, X1.3.
PLATE 13
BULL. AMER. PALEONT., VOL. 72
=.
;
LU
~
bladed dag
.
est
SS
ais
S33
Sa
¥
PLATE 14
BuLu. AMER. PALEONT., VOL. 72
>
mAs
RARE,
pc OOO TR he 6 OL
eeokemete VU 77 Fy
PUT PP gen
A
aay ba
SAGAN SLABS AP,
OrDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE- 127
EXPLANATION OF PLATE 14
All latex casts of Ectenocrinus simplex (Hall) from the Martinsburg Formation
Swatara Gap, Pennsylvania, except figure 8.
Figure Page
ilo,
9-10. Ectenocrinus simplex (Hali) ................... REARA oo ee PEM aR foe 89
(1) Derstler No. 5, B ray view, 1.7; (2) Derstler No. 11, D
ray view, 1.3; (3) Derstler No. 12, D ray view, X1.3; (4)
Derstler No. 16, interray view, 1.3; (5) Derstler No. 3b,
cray view, X1.3; (6) Derstler No. 3a, E ray view, X1.3; (7)
Derstler No. 5b, C ray view, X1.7; (9) Derstler No. 7a, D ray
view, 1.7; (10) Derstler No. 17, X1.3.
8. Ectenocrinus geniculatus (Ulrich) 2.000000. eee 89
Holotype, UCGM 36313, lowermost Kope Formation, Cincin-
nati, Ohio, AB interray, 1.3.
128 BULLETIN 296
EXPLANATION OF PLATE 15
All from lowermost Kope Formation, Cincinnati, Ohio.
Figure Page
1-2. Ectenocrinus geniculatus (Ulrich) <....................... 22222 89
Paratype, USNM 42219b, BC interray view 1.7 and X3.0.
3., E. geniculatus (Ulrich) ........5..0.2...2 a i ee 89
Latex cast of USNM 42219a which is a natural mold of the holo-
type (UCGM 36313), A ray view, X1.3.
4-6. E.-geniculatus (UITICN) 2..42.:..-cc.c.00000. deste ek oo eee 89
Paratypes, USNM 42219d, A ray view, *3, USNM 42219k, B ray
view, X1.7, and USNM 42219d, A ray view, 1.7.
PLATE 15
BULL. AMER. PALEONT., VOL. 72
PLATE 16
BULL. AMER. PALEONT., VOL. 72
es a. ¥} :
os eS mmr | | eee EEE
Ne Corman
as BAAg needs ry Ls
ee ene. RT
eae et gud NAD
aa
5 >
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE
EXPLANATION OF PLATE 16
ligure
1-3, 7. Ectenocrinus geniculatus (Ulrich) ................... Pee sess 2 le
Paratypes from the lowermost Kope Formation, Cincinnati, Ohio,
USNM 42219c, ?A ray view, X1.7, USNM 42219f, C ray view,
1.7, USNM 42219e, X3.0, and CFM UC8829, X<1.7.
#6. “ibexocrinus lepton Lane: ..2..2...2....2.06..0000..c0cscececcsececeseeess
Holotype, USNM 165239, from Ibex, Utah, CD interray view,
A ray view and CD interray view photographed under
ethanol, 2.1.
129
97
130 BULLETIN 296
EXPLANATION OF PLATE 17
All specimens from the Maquoketa Formation, Ft. Atkinson, Iowa.
Figure Page
i-2) “Sygeaulocrinus fypus: Ulrich: 02020... ..<-2:.0...-0-- 100
Holotype, USNM 89876, D ray view, X2.1 and 3.0.
3-4: /S. typus Ulrich: ...k.... es in tid. Avan. re 100
Lectotype, SUI 3771, of Ectenocrinus elongatus Thomas & Ladd,
a junior synonym of S. tyfus, A ray view and CD interray
view, X2.1.
pe TU eC) belo eR nr 100
Lectoparatypes of Ectenocrinus elongatus (see above), SUI 3772,
A ray view and E ray view, SUI 3774, A ray view and CD in-
terray view, X2.1.
O11 SS. Stypes ere ere hc kes 100
Illustrated specimens, SUI 37921, B ray view, SUI 37923, CD
interray view, and SUI 37922, DE interray view, X2.1.
PLATE 17
BULL. AMER. PALEONT., VOL. 72
PLATE 18
BuLL. AMER. PALEONT., VOL. 72
ORDOVICIAN-SILURIAN CRINOIDS: WARN AND STRIMPLE 131
EXPLANATION OF PLATE 18
All specimens from Kirkfield, Ontario.
Tigure
1-4,6. Daedalocrinus bellevillensis (Billings)
Illustrated specimens, UCGM K.42682, CD interray view, UCGM
K.42683, A ray view, ROM 619T, E ray view, UCGM
K.42684, CD interray view, and UCGM K.36696b, CD interray
view, X1.7.
Hee pelLevillensis: (Billings) 2. eect oats cece ee retest tee pitt _ 100
Lectotype of Daedalocrinus kirki Ulrich, a junior synonym of
D. bellevillensis, USNM S.2141, CD interray view 1.3 and
0.5.
INDEX
Note: Light face type refers to page numbers. Bold face type refers
to plate figures.
A
INGSIOCrINUS a ee 32
Albion Formation. ...... 10
alternatus,
Poteriocrinus .......... 78
alveolata, Favistella .. 24
Ambaloduswes 23
Anglo-Scandinavian-
Appalachian
PTOVINCE” 220.) tsccbse: 23
Anomalocrinus ........... 30
Apodasmocrinus ........ 5, 50, 60, 62,
75, '76, 91, 92
approximatum,
etradium se 24
angulata,
Trichonodella ........ 23
Arenig Stage .............. 10, 50
Arnheim Formation
(beds) tee. oes eee: A [a7
ASCOCTINUSs _.................- 7
Ashgillian Stage ........ 10, 33, 50
ATODOGEINUS) ....<..00-3225:: , 28, 30-33,
63, 66, '72-74,
76
AtyDOCEINUS! =.2..-: 6, 30, 31, af
AU AC Cate es ee 24
B
Bass Island beds ........ 10
becki, Triarthrus ........ eS
Belleville, Ont. ............ 9, 103
bellevillensis,
Daedalocrinus ....18 5, 50, 62, 100,
102, 131
Heterocrinus .......... 102, 103
Bellevue Member ...... 22
Bellevue Park
(Cinemnati)) =... 23
Belodinalee 23
Benbolt Formation .... 92, 93, 95
=pillings” sbeds ......... 10
iBisherspedss 4... 10
Black Riveran (Black-
riverian) Stage ........ 10, 33, 50, 61,
63, 76, 92, 93,
95
Blanchester Member .. 22
“Blue limestone” ........ 87
Boonville NiYe ce 114
Botryocrinus .............. 52
Brassfield Formation. 10, 14, 20
brauni, Ohiocrinus 10 _ 5, 44, 50,62,
70, 71,72
Bromide Formation .. 92, 93, 95
Buckhorn Member .... 65
Bull Fork Creek, Ky... 19
Bull Fork Formation.. 10, 16, 17, 18,
19, 20, 22, 38,
56, 119
Burgess Shale ............ 25
Cc
Calapoeciae ee 24
Caleeoerimus: 7.44)..." 39
Caleidocrinus .............. 30
Camp Nelson beds ..... 10
canadensis,
Ectenocrinus .......... 84, 87
Heterocrinus .......... 84, 87
Canadian Stage .......... 10, 50
capax, Lepidocyclus .. 24
Caradoc Stage ............ 10, 50
Carrollton Kyar 104
Caryocrinites .............. 53
Cataract beds. .....:..: 10
Cayugan Stage ............ 10, 50
Chambersburg beds .. 10
Champlainian Series. 10, 11, 25, 50
Chazyan Stage ............ 10, 50
Cincinnaticrinus ........ 5G) 25a:
31, 33, 34, 35,
37, 38, 40, 41,
43, 44, 45, 47,
48, 50, 51, 52,
53, 54, 55, 56,
57, 58, 59, 60,
61, 62, 63, 65,
66, 67, 68, 70,
72, 84, 85, 87
CimemnativArch 20
Cincinnati beds .......... 1s
Ber. 11, 1254:
17, 18
ah heen 11
Shee 6,8, So 1s 13,
14015 ay, 2
23, 24, 26, 38,
41, 43, 47, 55,
56, 59, 60, 67,
70, 84, 86, 89,
90, 114, 116,
119, 120, 122,
iPr t a Paz)
“Cincinnati Period” .. 14
“Cincinnati
Limestone”
Cincinnatian Series ..
Clarksville Member ..
Clarksville, Ohio
Clermont, Iowa
Clifton Avenue
(Cincinnati)
Climacograptus
Clinton beds
Cobleskill beds
Coburg beds
Columbia, Tenn. ........
INDEX
6, 8, 10, 11,
14, 15, 16, 20,
23, 26, 32, 50
22
119
9, 87, 125
8, 10, 17, 59,
61, 63
Columbicrinus ............ 6,28, 30; 31;
115
compactus,
Heterocrinus .......... 59
Constellaria —.............. 24
constrictus,
Dystatocrinus ........ 7 =+5, 44, 50, 55,
57, 58, 59, 60,
61, 70, 93, 120
Heterocrinus .......... 58, 59, 60, 70
Heterocrinus var.
COMpactus, sie... 60
corryvillensis,
Atypocrinus ............ 58, 59, 60
Corryville Member .... 22, 59
Covington, Ky ............ 9
Crab Orchard beds .... 10
crassus,
Columbicrinus ...... 2 31, 415
Cryptolithus) 922. 24
Curdsville (Limestone)
HOTMAtON os......-...-1- 84, 91
Gyathocrinus) 80
cylindricus,
HOMOCEINUS) 2. 78, 79, 80, 81
Cynthiana (beds or
limestone) ................ 15, 16, 59
Cyrionioduss 23
D
Daedalocrinus ............ 5565255305
32, 44, 50, 61,
74-76, 84, 100-
103, 131
Daube Ranch .............. 95, 96
daubei, Apodasmo-
CHINUS eee 5, 50, 91-95
Dayton Ohio —..2.- 20
decadactylus,
Botryocrinus ............ 52
Decorah Shale ............ 8, 10, 32, 61,
63
Decow beds
delecta, Prioniodina ..
Dendrocrinus
dentatum,
Rhynchotrema
Dichognathus
difficilis,
Heterocrinus
Dillsboro Formation.
Disparida
divaricans,
Grewingkia
Drakes Formation .....
Drepanodus
Drymocrinus
Dundee Member
Dunleith Formation...
Dystactocrinus
eatoni, Triarthus
Echmatocrinus
Economy Member
Ectenocrinus
Eden Formation
Eden Park
(Cincinnati)
Eden shales (beds) ....
Edenian Stage
Elkhorn Formation ....
elongatus,
Ectenocrinus
emacerata, Resserella
Engodine beds
Eoligonodina
Erisocrinus
Escharopora
Eutaxocrinus
exiguus, Heterocrinus
exilis, Heterocrinus 1
Ohiocrinus
133
24
10, 16, 20, 22
23
84, 89
5, 6, 28, 30,
31, 33, 41, 44,
50, 55, 57, 58,
59, 60, 61, 67,
70, 93, 120
11, 15, 16, 24
59, 60, 62, 63,
72, 74, 75, 76,
77, 82-93, 96,
98, 100, 127-
130
16, 17
13, 17,21
11-17, 21
8, 10, 16-25,
32, 35, 38, 41,
43, 47, 50, 57,
59, 61, 76, 77,
84, 89, 90
22
100, 130
2
53, 114
60, 115
INDEX
F
Fairmount Member .. 22
Fairview Formation .. 10, 16-19, 22,
38, 47, 56, 59,
67, 70, 72, 84,
86, 119, 122,
123
Fairview Heights
(Cinemnati) 119
Fairview Park
(Cmemnat) ye. 19, 23
falciformis,
Escharopora ............ 24
Mavistellaw ees: 24
flexuosus,
Cyrtoniodus) |... 23
florida, Constellaria .. 24
floweri,
Saffordophyllum .... 24
Foerstephyllum ......... 24
Fort Ancient beds .... 22
Fort Atkinson, Iowa .. 9, 99, 130
Fort Atkinson
Member 3.022... 6
Fort Loudon, Pa. ........ 9
Fort Mitchell, Ky. ...... 120
Frankfurt, Kentucky .. 9, 87
Hulton beds’... 15, 24
furcata furcata,
Pectodina 2.0.2... 23
G
geniculatus,
Drymocrinus .......... 89
Ecteno-
erinus __...14, 15,16 5, 25,44, 50;
62, 84, 89-91,
127, 128, 129
Heterocrinus .......... 84, 89
Geraocrinuss 2 30, 103
Graphiocrinus ............ 30
gracilis,
Heterocrinus ? ...... 2 38, 114
Poteriocrinus .......... 78, 79
Grand Detour
Formation =.........24: 61, 65
grandis, Ectenocrinus 84
Grant Lake, Ky. .......... 19
Grant Lake Limestone
(Formation) ............ 10, 16-20, 22,
38, 56, 59, 119,
120, 122
Grewingkia 3... 24
Guelph beds ................ 10
Hatter Limestone ...... 63
Herptocrinus .............. 30
Heterocrinus _.............. 5:6) 2600;
BUSBY Bul sit
38, 39, 40, 51,
53, 56, 63, 67,
82, 84, 103,
114-116, 122
heterodactylus,
Heterocrinus ........ 1 5,6, 34, 38-
40, 57, 51, 114,
116
Stenocrinus ............ 39
High Bridge beds ...... 10
Hill Quarry beds ........ inapasy yi.
21, 47
Hogskin Member ........ 93
HLOMOCEINUS eee 5, 6, 44, 74-
79, 82, 99, 124
hopensis,
Platystrophia .......... 24
Hudson River Group .. 11, 14, 15, 59,
70, 87, 88
Hueston Woods State
Park Ohi0) ee 119
Hull Limestone (beds) 6, 8, 10, 59,
61, 63, 65
Hunter Limestone ...... 63
huronensis,
Calapoecia _...2228 24
I
Ibexocrimus! 5, 50, 62, 74-
77, 84, 96, 98,
129
Ibex: cUtahie-.c. eee 9, 73, 96, 115,
129
LOCKINUS ee eee 30
ISOtelUS) oe ee 51
Isotomocrinus ............ 5, 6, 25, 28,
30, 33, 41, 52,
61, 63, 76, 121
J
JAaHnHOCEINUS, oe 74
Johnson City,
Okiahomavee ss ee 96
Juniete beds ................ 10
juvenis,
Heterocrinus ........ 2 39, 40, 52, 55,
K
Keanoshyshales
134
INDEX
Keislognathus ............ 23
Kiaeromena | «...c-..6...... 24
Kirkfieldian Stage ... 8, 10, 25, 32,
33, 50, 59, 61,
63, 65, 76, 84,
86, 103
Mackiields Onts =. 6, 8, 9, 61, 63,
65, 91, 103,
121
Kirkfield Quarry ........ 63
kirki, Daedalocrinus.. 100, 102, 103,
183
Kope Formation ........ 10, 16-19, 22,
24, 38, 47, 84,
86, 87, 90, 104,
116, 127
Kope Hollow (Ohio) .. 18
krateriform) 222. DAS PALS
KeyXIOTM eee. 7A
L
Lake Huron, Canada .. 90
Lasiocrinus
SCODALIUS a eee 81
Latonia Formation
(beds) Is}, 7)
laxus, Heterocrinus .. 66, 67, 68, 69,
70
....9_ 5, 44, 50, 62,
68, 69, 72, 122
See 11, 12, 13, 14,
laxus, Ohiocrinus
Lebanon beds
sai far ak
Lebanon Limestone .. 65333
Eebanon, Ohio ...........- ORS lial
23, 115, 119
Lecanocrinus
FOUL EKOUDIS! oc csesaccneeseenaaeene 82
lekythosiform ............ 26, 33, 43
Lepidocyclus capax .... 24
Leptaena
richmondensis ........ 24
lepton,
Ibexocrinus ........... 16 5, 50, 62, 129
Levanna, Ohio ............ 18
Lexington Formation.. 10
Rexine oa KY. ....:.:-.: 9, 20
Liberty Member ........ 22
Lichenocrinus
(lichenocrinid) ........ 5, 30, 45, 49,
51, 52, 53, 54,
87
Lincolnshire
HOrmatlOn i ...66s.:0:.:. 93
Llandeilo Stage .......... 10, 50
Llandovery Stage ...... 10, 50
Llanvirn Stage .......... 10, 50
Lockport-Guelph
MoOrmation eee ene
mockportiw Neyer... 8, 9, 78, 82,
124
Lorraine Group .......... 145 15) 21
Lower Whitewater
Member es niece: 22,
Tou dlOwWs ye este 9,14
Ludlow Stage .............. 10, 50
M
M zone ve AO Rar eee aeeeens CBee
Madisons indies... 9, 13, 67, 72,
122, 123
Manistique beds ........ 10
Manitoulin Island,
Canadas 8,9
manitoulinensis,
Drymocrinus .......... 89
Maquoketa Formation _ 6, 8, 10, 84,
87, 99, 125
Maquoketa Group ...... 22
Martinsburg
MOGMVAGION 2o.5scsce-c ee 8, 10, 38, 47,
Si 126127
Maysvillian Stage. ...... 8, 10, 17, 18,
19). 20; 215722,
23) 24, 25) 32;
35, 41, 47, 50,
55, 56, 57, 59,
60, 67, 70, 72,
76, 77, 84
Maysville Formation. 1522
Maysville, Ky. ............ 9, 16, 19, a
2
Meaford beds .............. 10
Medinon Stage. .............. 10, 50
meeki, Resserella ...... 24
Mercersburg beds ...... 10
Mesopaleaster ............ 104
Middle (Eden) shales.. Val
minutus,
Isotomocrinus ........ 5, 50, 62, 65,
MeMicken Member .... 22
McMillan Formation .. 22, 59
Montreal, Quebec ...... 8, 9, 61, 63,
84, 87
Mount Auburn
Member ....sccesccene 22
Mt. Hope Member ...... 22
Mountain Lake
Member (oe 93, 95
Myelodactylus ............ 3
Myzostome (galls) ...... 52
135
INDEX
N
Nassoviocrinus .......... 74
Newport, Ky.9.2--.... 9, 11, 24, 116
Niagaran Stage .......... 8, 10, 50, 76,
77, 82
°
ochanus,
Heterocrinus .......... 68, 69, 70, 122
OHIOCEINUS” "eee: 5, 6, 25, 28,
30, 33, 39, 40,
41, 44, 50, 62,
66, 67, 68, 69,
TkUS Cree Ee
122
Ohio River (Valley) .. 9, 11, 13, 14,
, 18, 20
QOnnrellay 24
Oregon beds ................. 10
Oregonia Member ...... 22
Orthograptus! 2... 24
Oswego) beds! ...2............ 10
Othneiocrinus ............ 72
Ottaway Ont 8, 9, 59, 61,
63, 65, 76; 77,
gcoy ee lPA
Oxtord, sOM0: 20.2; 9,13
Ozarkodinay ==...) 23
P
Paleofavosites ............ 24
Paltoduse +a 23
parvus,
Homocrinus ........ 11 5, 44, 50, 62,
77, 79, 80, 83,
89, 124
Peniculocrinus ............ 73
pentagonus, Cincin-
MaAGeEINUS eae 6 5, 25, 35, 40,
44, 50, 51, 52,
55, 56, 57, 60,
62, 72, 119
pentagonus,
Heterocrinus .......... 55, 119
Phialocrinus ................ 30
Phosphannulus ............ 52
Phragmodus).. 23
planumbona,
Strophomena ........... 24
Platycystites zone ...... 95
Platystrophia’ ....:..62 24
Plectodmiae. ee 23
Rlumville) Ky. ......-. 19
Point Pleasant beds.... 13, 14, 15, 16,
17.
Point Pleasant
Hormation, 32. 1G 2s:
19, 59
Point Pleasant,
OHIG 28. 28. oo. ae 9, 14, 17,18
Poteriocrinus — .......... 78, 80
Preachersville, Ky. .... 9, 20
Preachersville
Member 28%)...428 16, 20
Prioniodina 22... 23
Prioniodusy. 3. 23
priscus,
Atopocrinus .......... 2 5, 44, 50, 62,
72, 74, 75
proboscidialis .............. 79
propinquus,
Heterocrinus .......... 53
Pulaskisbeds 10
punctatus,
Apodasmocrinus .... 5,50, 90, 91,
92, 93, 94, 95,
96
punctatus,
Ectenocrinus .......... 96
Q
Queenston beds .......... 10
R
Rabbit Hatch, Ky. ...... 104
Rafinesquina .............. 51
Rapid Run Creek
(Cincinnati) .....= 116
raymondi,
Ectenocrinus .......... 84, 87, 88
Resserella. 24
Rhipidognathus ........... 23
Rhynchotrema ............ 24
richmondensis,
eptaena: ....cs eee 24
Richmondian Stage .... 8, 10, 14, 15,
20521 2a oo:
24, 25, 32, 38,
41, 50, 56, 57,
76, 77, 84, 86,
99
Richmond, Ind. .......... 9, 13, 21
Ristnacrinus ................ 30
River Quarry beds ...... bapa ls} Web
15, 16, 17
Rochester Shale .......... 6, 8, 10, ie
Rocklandian Stage .... 10, 50, 61
Rockland Formation .. 10
Rogers Gap, Ky. .......... 115
136
INDEX
rugosa, Strophomena.. 24
rustica, Grewingkia .. 24
Ss
Saffordophyllum ........ 24
Sarittodontus’ 2.0. 23
St. Edward beds ........ 10
St. Hyacinthe, Mani-
toulin Island ............ 90
St. Lawrence River
@owlands))-- ee a CA
st. Paul; Minn. ........... 8, 9, 61
Ine: DOS Aik. shee: 10
Saluda Member .......... 22
Saratoga County,
INES See ee Cage ene 114, 115
Secolopodus) =)... 23
scoparius,
Homocrinus ............ 79, 80, 81
scoparius,
TASIOCHINUS ....24....5 81
Sheguiandah
mormarion. 8, 10, 90
Sherman Fall beds .... 8, 10, 59, 61,
63
Shermanian Stage ...... 8, 10, 18, 25,
32, 50, 59, 61,
76, 77, 84, 86
simplex, Ecteno-
crinus ...... 12,13,14 5,6, 39, 44,
50, 52, 60, 62,
72, 84, 85, 86,
87, 88, 89, 90,
91, 93, 104,
125, 126
Heterocrinus .......... 38, 39, 82, 84,
86, 88
var. grandis,
Heterocrinus .......... 84, 88
Snake Hill, (N.Y.) ...... 114, 115
Southgate Member ..... 22, 104
sp., Drymocrinus ........ 89
sp. indet.,
Ectenocrinus .......... 50, 62, 63, 90,
91
sp., Ohiocrinus .......... 9 122
Springdale, Ky. .......... 19
Stemmatocrinus .......... 30
Stenocrinus ................ 30, 35, 39, 67,
82
Stones River Group .. 14
Strophomena ............... 24
suberectus,
Drepanodus ............. 248)
subundulata,
Trichonodella ........ 23
Summiteky ge 21
Sunset Member .......... 22
Swatara Gap, Pa. ........ 9, 87, 126,
127
Sygeaulocrinus .......... 5, 6, 30, 32,
74, 75, 76, 77,
93, 97, 99, 100,
101
T
Tamarack Point,
Manitoulin Island .. 90
tenuis,
Heterocrinus .......... 43, 61, 63, 65
Isotomocrinus ........ 8 5, 43, 44, 50,
55, 62, 63, 64,
65, 66, 121
Ozarkodina (2.3... 23
Trichonodella .......... 23
tesselatus,
Cryptolithus ............ 24
Metradiimers 24
TorontowOnts see.
Tremadoc Stage .......... 10, 50
Trentonian. 17
Trenton Falls, N.Y. .. 9, 11, 61, 84,
87, 116, 126
Trenton Group ............ Iba Save Pal
Trenton Limestone .... 59, 61, 63, 65,
84, 87, 91, 103
IAT TUSe eee eee 11, 15, 16, 24
Trichonodella .............. 7483}
truncatus richmonden-
sis, Orthograptus .... 24
typicalis,
Climacograptus ...... 24
typus,
Isotomocrinus ....... 63, 65, 121
Sygcaulocrinus ....17 5, 44, 50, 62,
89, 97, 98, 100,
130
ivrones beds 10
U
undatus, Phragmodus 23
Upper Whitewater ....
Utica Group
22
10, 11, 14, 15
vacuum,
Foerstephyllum ...... 24
137
INDEX
varibrachialus, Cincin-
naticrinus ...... ce at es a ae ls ae Se
41, 42, 43, 44,
45, 47, 48, 49,
50, 51, 52, 53,
54555) 565 50,
59, 60, 61, 65,
68, 70, 72, 76,
84, 86, 87, 104,
116, 117, 118
Warren (beds)
Wenlock Stage ............ 10, 50
West Branch of Syca-
more Creek, Okla. ..
West Covington, Ky..... 59
Westwood
(Cmeinnati) 119
Whetstone Gulf
ERormation - = 8, 10, 38, 47
Whiterockian Stage .. 8, 10, 32, 34,
50, 72, 73, 76,
77, 97
Whitewater
Hormation 22 20, 22
wideneri,
Eutaxocrinus .......... 49, 53
Wilmington, Ohio ...... 9
Z
Zygognathus ................. 23
138
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2- f LIBRARY
NOV 7 1977
BULLETINS
HARVARD
OF
AMERICAN
PALEONTOLOGY
(Founded 1895)
Vol. 72
No. 297
SOME PALEOCENE AND EOCENE
BARNACLES (CIRRIPEDIA) OF ALABAMA
By
Norman E. WEISBoRD
1977
Paleontological Research Institution
Ithaca, New York 14850 U.S.A.
PALEONTOLOGICAL RESEARCH INSTITUTION
1976-1979
PRESIDEN ce Pes cee een. eee eet ees es ee HAroLp E. VoKEs
WAICESPRESINEINM, scces se sees, ke Re ee ec ae en oe a ae Reine ent eS DUANE O. LERoy
SECRETARY pice sean fe ace aca ra) Coney Ae ea Reece | Ee ee a PHitip C. WAKELEY
ARICA SURE Weert crete et erence ee en No ee ee ERNESTINE Q. WRIGHT
DOP aah 9 V0) 2 eer eh A ae eee wn ee ee era KATHERINE V. W. PALMER
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ASSISTANT SECRETARY, ASSISTANT TREASURER ............-00---00ceeeseeeeeee REBECCA S. HARRIS
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BULLETINS
OF
AMERICAN
PALE ONLOLOGY
(Founded 1895)
Vol. 72
No. 297
SOME PALEOCENE AND EOCENE
BARNACLES (CIRRIPEDIA) OF ALABAMA
By
Norman E. WEISBORD
June 24, 1977
Paleontological Research Institution
Ithaca, New York 14850 U.S.A.
Library of Congress Card Number: 77-89951
Printed in the United States of America
Arnold Printing Corporation
Ithaca, N.Y.
CONTENTS
Page
JAA SOUNRGY TS GR ere eee ee eo re a ae 143
TICES NEYO OVO a me eG ee eS ee rere ee ee eo ee ee 143
Mocallatie swear dees tra tis ort al li ypu terse aca rs rasa eee eee eee 143
ENS NCTE ig EER ea aR Se ee er ee 144
IMESCEIp tlON POLS PCCIES mest ere seers eee oes ne ee ee ete 145
FREE O TC TICES Merete oe ate ee a a PO es 2 a RN pS EU eg ash 157
TEE GOS Gasset ed iad ee gm oe tap nies UN ee eh on Sas 5 ee en REPRE Re PAA en eA 161
SOME PALEOCENE AND EOCENE
BARNACLES (CIRRIPEDIA) OF ALABAMA
Norman E. WEISBOoRD
Department of Geology
The Florida State University
ABSTRACT
Four species are described and illustrated. Three of the taxa — Arcoscal-
pellum choctawensis, n. sp., Euscalpellum isneyensis, n. sp., and Balanus an-
tiquus (Meyer) are late Eocene in age, and one — Arcoscalpellum toulmini,
n. sp., is from middle Paleocene.
INTRODUCTION
The fossils described in this work, and the locality data per-
taining to them, were generously provided by Dr. Lyman D. Toul-
min of Florida State University, a colleague of mine in the Depart-
ment of Geology for nearly 20 years. Among many hundreds of
other taxa in the Toulmin collections (which were obtained during
many years of field mapping in the Southeastern Coastal Plain),
are four species of little-known barnacles from two localities in
Alabama: ACH-19, in the upper Eocene Yazoo Group, and ABu-5
in the Porters Creek Formation of Paleocene age. The three Yazoo
species are Arcoscalpellum choctawensis Weisbord, n. sp., Euscal-
pellum isneyensis Weisbord, n. sp., and Balanus antiquus (Meyer),
probably. The one Porters Creek species is Arcoscalpellum toulmint
Weisbord, n. sp., a form reminiscent of, but seemingly distinct from
Arcoscalpellum conradi (Gabb) found in the Paleocene Vincentown
Sand of New Jersey. The four species enumerated above from Ala-
bama have been deposited with the Paleontological Research Insti-
tution in Ithaca, N.Y.
LOCALITIES AND STRATIGRAPHY
ACH-19. The three Eocene species with the prefix ACH-19 were
collected between Silas and Isney, in Choctaw County, Alabama, on
U.S. Highway 84, about 4.0 and 4.2 miles west of Silas. The locality
lies in the NW 1/4 of Sec. 4, T 9 N, R 4 W, at approximately
31°46.5’N, 88°24’W, in a small outlier of the Yazoo “Clay”. Col-
lected 15 August 1966.
In Alabama (and parts of Mississippi) the Yazoo Group con-
sists of the following stratigraphic units, from bottom to top: the
North Creek Member, the Cocoa Sand, the Pachuta Marl, and at the
top, the Shubuta Member. The Yazoo sequence occupies the upper
three-fourths of the Jackson Group. The lower fourth of the Jackson
Group consists of the Moodys Branch Formation resting on the
144 BULLETIN 297
Scutella bed, which in Alabama represents the base of the Jack-
sonian. The Jackson Group correlates with the Bartonian and
Ludian Stages of England, the Bartonian the earlier of the two.
Thus the North Creek Member of the Yazoo, represented by the
ACH-19 barnacles is positioned in about the middle of the Jackson
Group which is upper Eocene in age.
ABu-5. The single Paleocene barnacle species, ABu-5, was
collected in Butler County, Alabama, Sec. 9, T 11 N, R 12 E, at ap-
proximately 31°56.5’N, 86°51’W. According to Toulmin’s notes the
specimens were obtained from a road cut on a paved road 1.4 miles
north of Wolf Creek and 3.0 miles north of Monterey, Butler County,
in the Porters Creek Formation. “The fossils are from the thin-
bedded zone of gray to brown calcareous sand and sandstones above
the lower clay.” Collected 11 February 1968.
The Porters Creek Formation is part of the Midway Group
which is Paleocene in age. In Alabama the Midway Group is made
up at the base of the Pine Barren Limestone followed above by the
McBryde Limestone, both limestones within the Clayton Formation.
The Clayton Formation is succeeded upward by the Porters Creek
Formation, the Matthews Landing Marl, the Oak Hill and Coal
Bluff Members of the Naheola Formation, and at the top of the
Midway, the Salt Mountain Limestone. The Midway group 1s cor-
related with three European stages, the Danian below, the Thanetian
in the middle, and the Sparnacian above.
Thus the Porters Creek Formation which is believed to extend
from the upper Danian to upper Thanetian in Alabama may be
considered as spanning the middle Paleocene in time.
ACKNOWLEDGMENTS
I wish to thank Frank H. Wind of Florida State University for
having taken and processed the photographs contained in this work.
I am also indebted to Charles W. Copeland of the Geological Sur-
vey of Alabama and Druid Wilson of the U.S. Geological Survey for
their help in trying to locate the type of Crucibulum antiquum
Meyer. The generic name of that species was later changed to
Balanus after the true classification was revealed by careful clean-
ing of the type specimen. It seems the Aldrich collection, which in-
cluded many of Meyer’s specimens, was originally deposited with
PALEOCENE-EOCENE BARNACLES: WEISBORD 145
Johns Hopkins University from which it was transferred a number
of years ago to the present stewardship of the U.S. Geological Sur-
vey at the National Museum of Natural History (U.S. National
Museum), Washington, D.C. Three or four of Meyer’s types are in
the Alabama Survey but Balanus antiquus (Meyer) is not among
them; neither is it in the Aldrich collection in Washington, D.C.,
and is thus presumed to be lost.
Katherine V. W. Palmer and her staff have been most helpful
in the editorial review of the work, and in the cataloguing of the
four species in the Paleontological Research Institution, Nos. 8205-
8219.
DESCRIPTION OF SPECIES
Class CIRRIPEDIA Burmeister, 1834
Order THORACICA Darwin, 1854
Suborder Lepadomorpha Pilsbry, 1916
Family SCALPELLIDAE Pilsbry, 1916
Arcoscalpellum (?) choctawensis Weisbord, n. sp. Pl. 19, figs. 9-12
The holotype is a right scutum (ACH-19c) broken off at the
apex and the basi-tergal angle, measuring 13.8 mm along the occlu-
dent margin and 7 mm in width across the basal margin. The valve
is thin and divided into unequl halves, the exterior of the tergal side
the narrower, flattish, and sloping, the larger and medial half sub-
regularly convex. Below the apical area there is a transverse depres-
sion across both halves of the exterior. The umbonal area is skewed,
and extending down from it to near the basi-tergal angle there is a
vague fold or bend demarcating the two sides of the exterior; at the
fold the numerous growth lineations form nearly a right angle, those
of the tergal side vertical, the ones of the middle part of the valve
horizontal, and those near the occludent margin turned down rather
sharply. In certain light and on the medial area of the exterior there
are seen very faint and thin longitudinal ridges extending from the
umbone toward the base. The upper and lower tergal margins are
straight and form an angle of approximately 143 degrees; the oc-
cludent margin is slightly wavy but nearly straight, and projects a
little below the base. In the interior of the scutum the apical area is
thickened and 6 mm in length, and nearer the occludent side of it
146 BULLETIN 297
there is a prominent furrow 4.5 mm in length and increasing in
width from .5 mm to 1.5 mm from top to bottom; adjacent to the
furrow on the lower occludent side is a slightly sunken, elongate
triangular area marked with fine longitudinal ridges which extend
upward into the elongated excavation under the apical area of the
exterior. The apical furrow leads below into a faint, rounded muscle
scar which is depressed below and nestled into the lower corner of
the apical area; bounding the occludent side of the apical furrow and
raised slightly above it is a thickened lamina curving around the
muscle scar and continuing down the thickened occludent margin
to near the base where it merges with the shell material. The in-
terior of the valve below the apical area is shallowly concave except
at the margins which are upturned, thus producing the appearance of
tumidity of the exterior.
The paratype is a right tergum (ACH-19cl) measuring 18 mm
in length and 11.75 mm in greatest width. The valve is thin, flat, and
rhomboidal-lanceolate in outline. The exterior of the carinal margin is
straight above, slightly concave below where it forms an angle of
about 122 degrees with the lower carinal margin; the upper and lower
occludent margins are nearly straight and form a rounded angle of
about 126 degrees with each other. The growth lineations of the
exterior are fine and numerous, and form an acute V at the apico-
basal demarcation line. In the interior of the valve the sides of the
apical area are widened to about 0.75 mm and diverge down from
the apex for about 5 mm, each side shallowly furrowed along the
middle. Just below the apex is a laminar ridge following the contour
of the apical area, one branch forming the outer rim of the apico-
carinal margin, the other running down the middle of the apico-oc-
cludent furrow; splaying off from the apex of this laminar ridge
are three short welded ridges, each of the outer ones forming the in-
ner margin of the apical furrows, and a medial one some 2.5 mm in
length terminating in a pointed spur at its lower end. The paratype
tergum is chalky and stained within.
Type locality. — ACH-19, about 4.0 and 4.2 miles west of Silas,
Alabama, on U.S. Highway 84, in NE 1/4 NW 1/4 Sec. 4, T 9N, R
4 W, at approximately 31°46.5’N, 88°24’ W, Choctaw County.
Formation. — Yazoo Group (North Creek Member). The north
PALEOCENE-EOCENE BARNACLES: WEISBORD 147
Creek Member lies above the Moodys Branch Formation and below
the Cocoa Sand in the Jackson Group of the upper Eocene.
Diagnosis. — The diagnostic features of the scutum are the
prominent elongated furrow of the apical area in the interior and the
slightly skewed umbone of the exterior. The tergum is characterized
by its rhomboid-lanceolate and flattened form and the unequal
areas of the exterior defined by the apico-basal ridge.
Comments. — This species is based on two complete valves, one
of them a scutum, the other a tergum. Both valves are thin-shelled,
occur within the same formation at the same locality, and are the
only two valves that are distinct from scores of others which repre-
sent another scalpellid species. Nevertheless I am not certain that the
two valves belong to the proposed new species, Arcoscalpellum
choctawensis. Therefore, the scutum is designated as the holotype
and the tergum as the questionable paratype of the new species.
Arcoscalpellum toulmini Weisbord, n. sp. Pl. 20, figs. 1-8
This species is based on four specimens — two scuta and two
terga — all presumed to belong to the same taxon.
The holotype is the left scutum (ABu-5al) which is 12 mm in
height from the apex to the basal margin, and 7.1 mm in width
across the base. The valve is trapezoidal, with a moderately pro-
nounced apico-basal fold which divides the exterior into unequal
halves — a smaller, flatly depressed tergal flank, and a broader,
convex occludent side. The occludent margin is evenly and slightly
convex, and projects a little at the basal angle; the basal margin is
nearly straight except at the basi-occludent angle where it swerves
downward; the carinal margin is straight, the tergal somewhat con-
cave. The apex is acute and turned a little toward the tergum. The
outer surface is scored faintly with concentric growth furrows be-
tween which are minute growth ridges, all of these contoured to
form a V at the apico-basal fold. Also there are faint longitudinal
radii on the larger half of the valve. In the interior of the left scutum
is a large roundish muscle-pit leading to a shallow medial depression
which broadens to the basal margin. The apical area, 5 mm in length,
is much thickened, its base rising vertically from the muscle-pit.
The occludent margin of the interior is also thickened, and there is
a fine incision or furrow running along the middle of it from near
148 BULLETIN 297
the base to near the apex. Extending downward from the apex there
are four crowded longitudinal ridges, the innermost one the highest
and extending along the occludent side, then swerving at the muscle-
pit, and continuing therefrom farther down to merge with the shell
substance of the margin; the other apical ridges are shorter and
bound an elongated triangular pit on the occludent side just below
the apex, and marked with six or seven oblique rugae. The tergal
side of the apical area is callused and smooth.
The paratype right scutum (ABu-5a2) is 11 mm in height and
about 5 mm in width across the base, and is thus proportionally
slightly narrower than the left scutum. It is similarly sculptured
on the outer surface except that in certain light longitudinal radii
on the more chalky surface are somewhat more distinct. The inner
surface is chalky and weathered and the triangular pit with the
oblique rugae of the left valve are not visible.
The paratype right tergum (ABu-5a3) is blue-gray in color
streaked with light gray, in contrast with the cream-colored or
whitish scuta; however, a fragment of another tergum (ABu-5a4),
similar to the paratype tergum, is also cream-colored, so that al-
though I do not know that the terga belong to A. toulmint, the
color difference of ABu-5a3 is no hindrance for considering it the
same.
The paratype tergum is thin, flat, elongate-subrhomboidal, and
broken or worn off at the apex and base. On the exterior there is
a faint apico-basal ridge or line of demarcation dividing the valve
into unequal halves, the carinal side the narrower, the valve shal-
lowly depressed on either side of the ridge. The numerous growth
lineations are fine and closely spaced, and form an acute V at the
apico-basal ridge. As seen in the interior, the carinal border is slightly
upturned, but the rest of the inner surface is shallowly concave. The
shorter apico-carinal margin slants at an angle of about 20° and the
longer apico-occludent margin at about 30° with reference to a ver-
tical axis; whether these margins meet to form a pointed apex, or
whether the apical area is truncate or blunt as on the two terga at
hand is not known. The likelihood is that the apex on a complete
tergum is moderately acute. Below the apex are two short longi-
tudinal ridges with a narrow depression between them. Both apical
margins are beveled, a little widened, inclined inward, and built up
PALEOCENE-EOCENE BARNACLES: WEISBORD 149
of exceedingly fine elongated ridges, the innermost of which appears
to be undercut by the depression of the inner surface of the valve.
The paratype tergum (ABu-5a3) is 10.75 mm in length and 6 mm
in greatest width but is broken off at the base and apex. Tergum
ABu-5a4 is a fragment about 7.5 mm in height, 4 mm in greatest
width.
Type locality. — ABu-5, Butler County, Alabama, Sec. 9, T 11
N, R 12 E, the approximate coordinates 31°56.5’N, 86°51’W. Ac-
cording to Toulmin’s notes the specimens were collected from a road
cut on a paved road 1.4 miles north of Wolf Creek and 3.0 miles
north of Monterey, Butler County. “The fossils are from the thin-
bedded zone of gray to brown calcareous sand and sandstone above
the lower clay.”
Formation. — Porters Creek (lower Member); Paleocene.
Comparisons. — The left scutum of A. toulmini resembles the
left scutum of A. conradi (Gabb) from the Vincentown Limesand
(Paleocene) of New Jersey, but among other differences the apico-
basal ridge of the exterior is less definite, and the apical area of the
interior far less elaborately sculptured and ridged on the occludent
side of A. conradi. These differences might be explained by the
greater weathering of the A. conradi scutum, but because the tergum
of A. conradi is not known, and the carina of A. toulmini is not
known, the two species are considered distinct on the observable dif-
ferences between the type left scuta of each.
Externally the tergum of Arcoscalpellum toulmini is remark-
ably similar to that of Arcoscalpellum bakeri Collins (1973) from
the Maestrichtian, Ripley Formation of Oktibbeha County, Missis-
sippi. Not observed on A. toulmini and present on the tergum of
A. bakeri is a faint groove “extending from the apex to near the base
of the scutal margin and between this and the apico-basal ridge ex-
tend several fine ridges.” Because the carina of A. toulmint is not
known and the scutum of A. bakeri not known the slight apparent
differences in the two terga plus the discrepant stratigraphic posi-
tions of the two taxa lead me to consider each a valid species. Car-
rying this a step farther, the carina of the Upper Cretaceous A. bakeri
is dissimilar from the carina of the Paleocene A. conradi. I suspect
that when the carina of A. toulmini is found that it too will be
unique to the species.
150 BULLETIN 297
Euscalpellum isneyensis Weisbord, n. sp. Pl. 19, figs. 1-8
This species is based on 106 valves, 59 of them scuta, 47 terga.
The holotype is a left scutum (ACH-19al1), measuring 15.5 mm in
length and 6.25 mm in greatest width; a paratype right scutum
(ACH-19a2) is 19.5 mm in length and 7.75 mm in greatest width.
The scutum is elongated, crescentic, and moderately tumid, the
length about 2-1/2 times the greatest width. The apex is acute and
beaklike, and is turned slightly toward the tergal margin. The apico-
basal ridge is narrow, curved, and well defined; diverging from it
on the paratype a short distance below the apex is a faint minor
fold continuing to the base where it is close to the apico-basal ridge;
between the two, the surface of the valve is flattish. The tergal
margin is gently concave at the upper fourth, convex below, with a
rounded angulation at about the lower third. Near the occludent
margin, which is convex, there is a fine ridge extending from the
apex toward the base where it plays out into a vague narrow rise;
the occludent side of the valve adjacent to this rise is narrow and
somewhat depressed as is the narrow tergal side from the apex to the
lower third of that side. The base is slightly concave on the occludent
side of the apico-basal ridge which itself terminates acutely. The ex-
terior of the scutum is strongly sculptured by growth markings which
form V’s at the apico-basal ridge and are sharply upturned along
the margins. The markings consist of elevated ridges with flattish
spaces between them, the interspaces themselves bearing micro-
scopic striae. The inner surface of the scutum, except for the up-
turned margins, is shallowly concave. The adductor muscle-pit is
large, subrounded, and nestled more or less centrally into the base
of the apical area. The apical area is thickened, about 6.5 mm in
length, and is marked by closely spaced fine ridges or striae diverging
from the apex down the widened margins, the ridges of the o¢cludent
margin playing out at the basi-occludent angle, those of the tergal
margin terminating opposite the top of the muscle pit; the rest of
the interior, including a central triangular area well below the apex
proper, is smooth. The interior of the right scutum (ACH-19a2) is
similar to that of the left scutum except that the apical ridges on
the occludent side are stronger than those on the tergal side.
The tergal valves vary considerably in outline, some being
elongate subtriangular (ACH-19a3), others subrhomboidal (ACH-
PALEOCENE-EOCENE BARNACLES: WEISBORD Sl
19a4), and some obtusely subpentagonal (ACH-19a5). A left tergal
valve (paratype ACH-19a3) measures 17.2 mm in length and 7.5
mm in greatest width; a right tergal valve (paratype ACH-19a5) is
19 mm in length and 10.75 mm in greatest width; paratype ACH-
19a4 is 16 mm in length and 8 mm in greatest width.
Both terga are flat, the left one, as exemplified by ACH-19a4,
subrhomboidal in outline, with an arcuate apico-basal ridge and a
subacute apex turned toward the carina. Externally the middle area
of the valves is slightly depressed. The occludent margin is convex,
moderately so above, less so below; the carinal margin is vertical
above, nearly straight to slightly concave from the lateral angle to
the base; the basal margin is nearly straight but becomes convex at
the basi-occludent angle. The exterior of the terga is sculptured by
prominent growth ridges and furrows, the principal ones thickening
at the intercepts of the apico-basal ridge, the furrows lined with
fine striae. There is a slight narrow depression along the upper half
of the occludent margin, bounded by a faint line of sculpture-
demarcation from the apex to the basi-occludent angle. The growth
markings form a V at the apico-basal ridge and swerve upward at
the sides. The inner surface of the terga is smooth and flat, and
showing through the calcification in lesser or greater degree are the
reflections of the principal growth ridges of the exterior. The apical
area is marked by fine striae diverging from the apex halfway down
the carinal margin and nearly the full length of the occludent
margin. In the interior of the right tergum (ACH-19a5) there is an
oval depression or hollow in the upper middle of the widened occlu-
dent margin. Unfigured paratype 8209 PRI.
Type locality. — ACH-19, Choctaw County, Alabama, about
4.0 and 4.2 miles west of Silas, on U.S. Highway 84, in NW 1/4 of
Sec. 4, T 9 N, R 4 W, at approximately 31°46.5’N, 88°24’W, in an
outlier of the Yazoo Group.
Formation. — Yazoo Group (North Creek Member); lower up-
per Eocene.
Diagnosis. — The scutum of this species is characterized by its
elongated crescentic form, the relatively long apical area, the sharp
apico-basal ridge, and the strong external markings. The tergum is
characterized by its subrhomboidal outline, the strong apico-basal
152 BuLLeTIN 297
ridge, the slight medial depression adjoining the ridge, and the
prominent external markings.
The tergum of this species somewhat resembles that of Luscal-
pellum eocenense (Meyer) (1895) from the middle Eocene Clai-
borne Group of Alabama, Mississippi, and Texas, but is differentiated
from FE. eocenense by its rhomboidal shape and strong apico-basal
ridge. The scuta of both these species, however, are distinct.
Suborder BALANOMORPHA Pilsbry, 1916
Family BALANIDAE Leach, 1817
Balanus antiquus (Meyer) Pl. 20, figs. 9-11; Pl. 21, figs. 1-9, 11
Crucibulum antiquum Meyer, 1886b, p. 68, pl. 1, fig. 11; 1887a, p. 55; Pilsbry,
1930, p. 433; Palmer, 1937, p. 149; Zullo. 1963, p. 133; Palmer and Brann,
1966, p. 616.
Balanus antiquus (Meyer), Meyer, 1887a, p. 55; Pilsbry, 1930, p. 433; Palmer,
1937, p. 149; Zullo, 1963, p. 133; Ross, 1965, p. 60; Palmer and Brann,
1966, p. 616.
Balanus aff. B. unguiformis J. de C. Sowerby, 1846, pl. 648, fig. 1; Withers,
1953, pp. 72, 91-92; Zullo, 1960, p. 21; Ross and Newman, 1967, pp. 4-7.
?Hesperibalanus antiquus (Meyer), Zullo, 1963, pp. 133, 207-208, text-fig. 10A.
Meyer’s original description of this species in 1886, under the
name of Crucibulum antiquum was the following:
CRUCIBULUM ANTIQUUM, n. sp. Pl. 1, fig. 11.
Subconical; margin oval, striate within; diaphragm entire; rhom-
boidal, close to the shell.
Locality. — Claiborne, Ala.
The surface of the single specimen is badly preserved. If I am
not mistaken it is the first Crucibulum found in the Old Tertiary
Formation.
In 1887, under Notes, p. 55, Meyer emended the generic desig-
nation of Crucibulum to Balanus with these statements.
The following mistake is to be corrected. I described a specimen
from Claiborne as ‘Crucibulum antiguum’ (Bull. 1, Geol. Surv. Ala.,
1886, p. 68, pl. 1, fig. 11). Having recently carefully cleaned the out-
side of this specimen it proved to be a Balanus with preserved oper-
culum.
Measurements of the type were not given, but if I judge the
scale next to Meyer’s figure 11 correctly, the carino-rostral length
at the base is about 12.5 mm, and the width across the base at its
widest about 10.5 mm, and this is somewhat smaller than the shell
of our ACH-19b.
The four individual specimens referred to B. antiquus (Meyer)
PALEOCENE-EOCENE BARNACLES: WEISBORD 153
consist of a nearly whole shell (ACH-19b) with an entire but half-
covered basis; a carinolateral compartment (ACH-19b1) with the
rim of the basis also preserved; another carinolateral (?) compart-
ment (ACH-19b2); and a nearly entire rostrum (ACH-19b3). No
opercula have been found, and there is little likelihood they are
present within the sandstone-filled orifice of ACH-19b which has
been dug into as far as possible.
ACH-19b has six compartments, is low conic, and is elon-
gate-oval around the basal margin. The specimen is slightly mashed,
and the rhomboid orifice is filled with sandstone, obscuring the
peritreme. The compartments are intact, and the exposed half of
the basis is well enough preserved to show that it is thick and cal-
careous, and seems to consist of small, closely spaced tubules radi-
ating from an off-centered nucleus; the bottom surface of the basis
is crossed by growth lineations following somewhat eccentrically the
contour of the basal margin of the shell. In plan view the rim of
the base is seen to consist of small quadrangular openings, each
one bounded by a short lamina or septum, the openings representing
the termini of the tubules of the basis. The aspect around the basal
rim is so similar to that of Meyer’s drawing of Balanus antiquus that
the specimens in the Toulmin collection described herein are believed
to represent the same species.
As viewed externally, the carina of ACH-19b is somewhat
concave in profile, the rostrum somewhat convex, the lateral com-
partments much the widest, and the carinolaterals much the nar-
rowest. The radii are comparatively wide at their widest, varying
from about 1 mm on the carinolaterals to 2 mm on the laterals. The
summits of the radii are very oblique and obscurely crenate, and
within, their sutural edges are strongly crenulate. The alae are rela-
tively broad and undulatory, varying in height from 2 mm to
about 3 mm.
ACH-19b has a carino-rostal length at the base of 14.3 mm,
a maximum width of 9 mm across the base, a height of about 9 mm
at the carinal end (broken at the apex), and a height of about 10.5
mm at the rostral end (broken at the apex). The sandstone-filled
orifice is approximately 9.5 mm in length and 7.8 mm in width. In-
dividual measurements of the compartments are tabulated below
in millimeters.
154 BULLETIN 297
Measurements of ACH-19b
Compartment Width at base Height
Carina 4.3 8.7
Carinolaterals De) 9.7
2.6 7.0
Laterals 8.5 11.6
9.0 8.5
Rostrum 6.5 10.5
The individual compartments from shells other than the ACH-
19b but believed to be representatives of Balanus antiquus (Meyer)
are described below.
ACH-19b1: Carinolateral compartment, height 10.6 mm, width
at base 6.6 mm. Radius broken away above. Sheath height about
3 mm. Number of longitudinal ribs in the interior about 26. Thick-
ness of basis at rim of compartment 0.6 mm.
The internal ribs are narrow and pronounced in about the lower
3 mm of the compartment; above that they are abruptly weaker and
continue so upward to near the base of the sheath. Below, each rib
merges into the basis where it becomes a septum in the narrow space
between the basis and inner surface of the paries. Thus looking down
on the base of the paries its periphery is seen to consist of small
quadrangular openings, each one bounded by a septum. The com-
partment is moderately convex and marked on the surface by
numerous fine growth lineations through which appear some faint
longitudinal radii which are reflections of the inner ribs.
The sheath occupies a little less than the upper third of the
compartment. On the radius side there is a pronounced transverse
furrow below which the sheath is hollowed out a little more than
on the opposite half which is undercut but slightly.
The ala is divided into unequal halves, the outer the smaller,
by a line of demarcation, at which the lines of growth form a
pronounced V.
ACH-19b2: This is externally smooth and is inferred to be ano-
ther carinolateral compartment, 8 mm in height and 5.2 in width
across the base (PI. 21, figs. 8, 9). There are about 28 internal ribs,
PALEOCENE-EOCENE BARNACLES: WEISBORD 155
a number of them alternating in size, but all of them pronounced and
confined to the lower 1.5 mm of the interior. The remainder of the
interior is smooth because of the thickened shell which covers the
weaker ribs underneath. The sheath is 4 mm in height and is under-
cut slightly the full width. The furrow on the radius side is even more
deeply excavated than in ACH-b1 but the ala of ACH-b2 does not
show the line of demarcation of ACH-b1.
ACH-19b3: This is a convex rostrum 10 mm in length and 8.2
mm in breadth across the base. The two radii are intact, each with
oblique and erose summits which are obscurely crenate, and each
with regularly, strong, and simply crenulate margins. The outer sur-
face is smooth with faint concentric and in places crinkly growth
lines on the paries, and nearly vertical growth lines on the radii. In
the interior there are about 47 narrow ribs, these most pronounced
near the base but continuing weakly to about the middle of the paries
where some play out and others persist to near the base of the sheath.
The sheath is about 3.4 mm in height and is slightly undercut the
full width. Diverging from the apex are two sharp laminar ridges
projecting slightly below the base of the sheath to form an inverted
V bounding the apical area, with the strong concentric striae of the
apical area and of the radii abutting each of the ridges. The interior
of the rostrum is white except for the tan apical area and radii,
whereas the exterior is tan and the radii alternating tan and white.
Diagnosis. — The distinguishing characters of the shell are its
thomboid aperture, the externally smooth parietes, the numerous
longitudinal ribs in the interior, the broad lateral compartments,
and the row of small quadrangular openings (of which there may be
as many as 180 on an adult specimen) around the basal rim of the
shell.
Type locality. — Claiborne (approximately 31°33’N, 87°31’W),
Monroe County, Alabama. The precise locality and stratigraphic
position of the fossils collected by Meyer at Claiborne are not known,
but the locality may well have been Claiborne Bluff or Claiborne
Landing on the Alabama River. The Bluff, from water level to top
is made up below of the Lisbon Formation overlain by the Gosport
Sand, both of the Claiborne Group of middle Eocene age; the Gos-
port is succeeded by the Moodys Branch and Yazoo Clay of the
Jackson Group (upper Eocene). The Yazoo Clay of the Bluff is
156 BULLETIN 297
equivalent to the North Creek Member of the Yazoo between Silas
and Isney in Choctaw County, the latter the collecting locality of the
Balanus antiquus (Meyer) of this paper.
Other localities. — “Middle Eocene, Gosport Sand, Claiborne
Landing, Alabama River”, as Hesperibalanus gosportensis Zullo (PI.
21, fig. 10). Zullo (1963, p. 133) stated that his H. gosportensis “is
probably the same as Balanus antiquus but the latter species is not
recognizably described.” Zullo (1963, pp. 207-208, text-fig. 10A)
figured a tergum of H. gosportensis, but not having seen Meyer’s
operculum or the shell to which it was attached, I cannot affirm
that H. gosportensis is the same as Balanus antiquus (Meyer),
though it may be.
Collections in the American Museum of Natural History from
the Claiborne area of Alabama are reported by Ross and Newman
(1967) as “Eocene Claiborne Beds, Claiborne, Alabama, Hall col-
lection” and “Gosport Sand, Middle Eocene, Claiborne Group, Clai-
borne Landing. Collectors Donald F. Squires and William Heaslip,
August 1955”. In these collections it is likely that the Balanus sp.
aff. B. unguiformis of Ross and Newman is equivalent to B. antiquus
(Meyer) from the type locality. As indicated by Ross and Newman,
the type of Balanus unguiformis J. de C. Sowerby (1846, pl. 648,
fig. 1), a taxon occurring in the upper middle Eocene and upper
Eocene of England, is not clearly established either by Sowerby’s
illustration or later description by Darwin (1854, pp. 296-298, pl.
8, figs. 8a, 8b). According to Darwin (1854, p. 297), there is indeed
a smooth-plated variety of B. wnguiformis, and its stratigraphic
position is equivalent to that of B. antiquus (Meyer). Nevertheless,
despite the unlikely but possible precedence of the name wnguiformts,
I prefer to relate the Choctaw County taxon to B. antiquus (Meyer),
because even as to size, the base of the shell looks like Meyer’s draw-
ing of B. antiquus and is close geographically and stratigraphically to
the Claiborne B. antiquus.
Mississippi: As “Balanus (B.) aff. ungutformis” reported from
the Jackson Group of Mississippi by Withers (1953, p. 72). A
locality in Mississippi was not cited by Withers, but I suspect it
might be west of Isney, Alabama, in the Yazoo terrain which ex-
tends into easternmost Mississippi from western Alabama.
PALEOCENE-EOCENE BARNACLES: WEISBORD 157
Florida: As Balanus sp. aff. B. unguiformis Sowerby in Ross and
Newman (1967). Limerock quarry about 200 yards south of the
Withlacoochee River, NE 1/4 Sec. 12, T 12 S, R 16 E, Citrus Coun-
ty. Inglis Limestone, Ocala Group, upper Eocene. Collectors Jackson
E. Lewis and Arnold Ross, May, 1965. The Inglis Limestone is con-
sidered to be in the lower part of the upper Eocene, or by some
geologists, in the middle Eocene. Thus the stratigraphic position is
about the same as for Balanus antiquus. Although the Florida speci-
mens are not well enough preserved, better material may indicate
that the taxon is conspecific with B. antiquus.
Geologic range. —- Upper middle Eocene to lower upper Eocene
in Mississippi, Alabama, and probably Florida.
Classtfication. — As stated by Ross and Newman, definite con-
clusions pertaining to the generic or subgeneric classification of the
taxon in question cannot yet be reached. And, as stated by Zullo, his
Hesperibalanus gosportensis cannot be assigned to Balanus antiquus
(Meyer) until the opercular valves of both are known. Unfor-
tunately I have been unable to track down the whereabouts of the
cleaned type specimen of Meyer to which the operculum is adherent
and which also might reveal the character of the shell itself.
REFERENCES
Bassindale, R.
1964. British barnacles. With keys and notes for the identification of
the species. Linnean Soc. London, Synopsis of the British Fauna,
pp. 1-68, figs. 1-16.
Burmeister, Carl Hermann Conrad
1834. Beitrége zur Naturgeschichte der Rankenfiisser (Cirripedia), Pp.
i-viii, 1-60, 2 pls., Berlin.
Cheetham, Alan H.
1963. Gooseneck barnacles in the Gulf Coast Tertiary. Jour. Paleont.,
vol. 37, No. 2, pp. 393-400, pl. 46, text-fig. 1.
Collins, Joseph Stephen Henry, and Mellen, Frederic Francis
1973. Cirripedes from the Upper Cretaceous of Alabama and Mississippi,
eastern Gulf region, U. §. A. I. Palaeontology, J. S. H. Collins. 1.
Geology, F. F. Mellen. British Mus. (Nat. Hist.), Bull. Geol., vol.
23, No. 6, pp. 351-388, pls. 1-5, text-figs. 1-5.
Darwin, Charles Robert
1851. A monograph on the sub-class Cirripedia, with figures of all the
species. The Lepadidae; or, pedunculated cirripedes. Ray Society,
London, pp. i-xi, 1-400, pls. 1-10, text-figs. 1-3 + 2 figs. p. 28.
1854. A monograph on the sub-class Cirripedia, with figures of all the
species. The Balanidae, (or sessile cirripedes); The Verrucidae,
etc., etc., etc. Ray Society, London, pp. i-viii, 1-684, pls. 1-30, text-
figs. 1-11.
158 BULLETIN 297
Gabb, William A.
1876. Note on a discovery of representatives of three orders of fossils
new to the Cretaceous Formation of North America. Acad. Nat.
Sci., Philadelphia, Proc., vol. 28, pp. 178-179, pls. 5, 17.
Hoek, Paulus Peronius Cato
1907. The Cirripedia of the Siboga-Expedition. A. Cirripedia Peduncu-
lata. Siboga-Expeditie, Uitkomsten, vol. 18, Mon. XXXIA, pp.
1-127, pls. 1-10.
1913. The ‘Cirripedia of the Siboga-Expedition. B. Cirripedia Sessilia.
Siboga-Expeditie, Uitkomsten, vol. 18, Mon. XXXIB, pp. 129-275,
pls. 11-27, text-figs. 1, 2.
Leach, William Elford
1817. Distribution systématique de la class Cirripédes. Jour. Phys., Chim.
et d’Hist. Nat., vol. 85, pp. 67-69.
Meyer, Otto
1885. The genealogy and the age of the species in the Southern Old
Tertiary. Amer. Jour. Sci., ser. 3, vol. 130, No. 175, art. X, pp. 60-
72, text-figs. a-c.
1886a. Observations on the Tertiary and Grand Gulf of Mississippi.
Amer. Jour. Sci., ser. 3, vol. 132, No. 187, art. III, pp. 20-25.
1886b. Contributions to "the Eocene paleontology ‘of Alabama and Missis-
sippi. Geol. Sur. Alabama, Bull. 1, pt. 2, pp. 63-85, pls. 1-3.
1887a. On invertebrates from the Eocene of. Mississippi and Alabama.
Acad. Nat. Sci., Philadelphia, Proc., vol. 39, pp. 51-56, pl. 3.
1887b. Beitrag zur Kenntniss der Fauna der Alttertiars von Mississippi
und Alabama. Senckenberg. Naturforsch. Gesell. Frankfort am
Main, Bericht, vol. 2, pp. 3-22, 2 pls.
Newman, William A., Zullo, Victor A., and Withers, T. H.
1969. Cirripedia. Treatise on Invertebrate Paleontology, Part R, Arthro-
poda 4, vol. 1, pp. R206-R295, figs. 80-119. Geol. Soc. America and
Univ. Kansas.
Newman, W. A., and Zullo, V. A.
1969. Addendum to Cirripedia. Treatise on Invertebrate Paleontology,
ere R, Arthropoda 4, vol. 2, p. 628. Geol. Soc. America and Univ.
Kansas.
Palmer, Katherine V. W.
1937. The Claibornian Scaphopoda, Gastropoda and dibranchiate Cepha-
lopoda of the southern United States. Bull. Amer. Paleont., vol. 7,
No. 32, Pt. 1, Text, pp. 1-548; Pt. 2, Plates, pp. 550-730, pls. 1-90.
Palmer, Katherine Vv. W., and Brann, Doris Cc.
1965. Catalogue of the Paleocene and Eocene Mollusca of the southern
and eastern United States. Part I. Pelecypoda, Amphineura,
Pteropoda, Scaphopoda, and Cephalopoda. Bull. Amer. Paleont.,
vol. 48, No. 218, pp. 1-466, pls. 1-3.
1966. Catalogue of the Paleocene and Eocene Mollusca of the southern
and eastern United States. Part II. Gastropoda (excluding Ptero-
poda, Pt. I, 1965). Bull. Amer. Paleont., vol. 48, No. 218, pp. 467-
1057, pls. 4-5.
Pilsbry, Henry Augustus
1897. Scalpellum and Balanus from Texas. Acad. Nat. Sci., Philadelphia,
Proc., vol. 49, pp. 332-333, fig. 1.
1907. The barnacles (Girripedia) contained in the collections of the U.S.
National Museum. U.S. Nat. Mus., Bull. 60, pp. i-x, 1-122, pls. 1-11,
text-figs. 1-36.
1916. The sessile barnacles (Cirripedia) contained in the collections of
the U.S. National Museum; including a monograph of the American
species. U.S. Nat. Mus., Bull. 93, pp. i-xi, 1-366, pls. 1-76, text-
figs 1-99.
PALEOCENE-EOCENE BARNACLES: WEISBORD 159
1930. Cirripedia (Balanus) from the Miocene of New Jersey. Acad. Nat.
Sci., Philadelphia, Proc., vol. 82, pp. 429-433, pls. 36-37.
Ross, Arnold
1965. A new cirriped from the Eocene of Georgia. Florida Acad. Sci.,
Quart. Jour., vol. 28, No. 1, pp. 59-67, figs. 1-2.
Ross, Arnold, and Newman, William A.
1967. Eocene Balanidae of Florida, including a new genus and species
with a unique plan of “turtle-barnacle” organization, Amer. Mus.
Novitates, No. 2288, pp. 1-21, figs. 1-7.
Sowerby, James de Carle
1846. The Mineral Conchology of Great Britain; or coloured figures and
descriptions of those remains of testaceous animals or shells which
have been preserved at various times and depths in the earth, Lon-
don, Pt. 113, pp. 57-80, pls. 644-648.
Toulmin, L. D., Lamoreaux, P. E., and Lanphere, C. R.
1951. Geology and ground-water resources of Choctaw County, Alabama.
Geol. Sur. Alabama, Spec. Rept. 21 and County Rept. 2, pp. i-ix,
1-197, pls. 1-11, text-figs. 1-23, tables 1-4.
Withers, Thomas Henry
1924. The fossil cirripedes of New Zealand, New Zealand Dept. Mines,
Geol. Sur. Branch, Palaeont. Bull. 10, pp. 1-47, figs. 1-8, pls. 1-8,
map.
1935. Catalogue of fossil Cirripedia in the Department of Geology. Vol.
2. Cretaceous. British Mus. (Nat. Hist.), pp. i-xili, 1-534, pls. 1-50,
text-figs. 1-64.
1953. Catalogue of fossil Cirripedia in the Department of Geology, Vol.
3. Tertiary. British Mus. (Nat. Hist.), pp. i-xv, 1-396, pls. 1-64,
text-figs. 1-105.
Zullo, Victor August
1960a. Eocene species of the genus Balanus (Cirripedia). Geol. Soc. Amer.,
Bull., vol. 71, No. 12, pt. 2, p. 2084. (Abstract).
1960b. Cenozoic Balanomorpha of the Pacific Coast of North America.
Master’s Thesis, University of California, Berkeley, pp. 1-147, pls.
1-9.
1963. Classification and phylogeny of the Balanomorpha (Cirripedia).
Doctoral Dissertation, University of California, Berkeley, pp. i-iv,
1-372, pls. 1-2, text-figs. 1-14. [This has been published in book
form by Xerox University Microfilms, P.O. Box 1346, Ann Arbor,
Michigan 48106. ]
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PLATES
162 BULLETIN 297
EXPLANATION OF PLATE 19
Figure Page
1-8. Evuscalpellum (?) isneyensis Weisbord, N. Sp. .........00...cccceeeeeeeeee 150
Figs. 1, 2. Interior and exterior of holotype scutum, No. 8205 PRI,
ACH-19al1. Natural size 15.5 mm X 6.25 mm.; figs. 3, 4. Ex-
terior and interior of paratype scutum, No. 8206 PRI, ACH-
19a2. Natural size 19.5 mm X 7.75 mm.; figs. 5, 6. Interior and
exterior of paratype tergum, No. 8207 PRI, ACH-19a4. Natural
size 16 mm X 8 mm; figs. 7, 8. Exterior and interior of para-
type tergum, No. 8208 PRI, ACH-19a5. Natural size 19 mm
< 10.75 mm.
9-12. Arcoscalpellum (?) choctawensis Weisbord, n. Sp. ..................0.. 145
Figs. 9, 10. Exterior and interior of holotype scutum, No. 8210
PRI, ACH-19c; Natural size 13.8 mm X 7 mm; figs. 11, 12.
Exterior and interior of paratype tergum, No. 8211 PRI, ACH-
19cl. Natural size 18 mm X 11.75 mm.
BULL. AMER. PALEONT., VOL. 72 PLATE 19
BULL. AMER. PALEONT., VOL. 72 PLATE 20
PALEOCENE-EOCENE BARNACLES: WEISBORD 163
EXPLANATION OF PLATE 20
Figure Page
1-8. Arcoscalpellum toulmini Weisbord, n. sp. ............0000ccccceeeeee 147
Figs. 1, 2. Exterior and interior of holotype scutum, No. 8212 PRI,
ABu-5al. Natural size 12 mm X 7.1 mm; figs. 3, 4. Exterior
and interior of paratype scutum, No. 8213 PRI, ABu-5a2.
Natural size 11 mm X 5 mn; figs. 5, 6. Exterior and interior
of paratype tergum, No. 8214 PRI, ABu-5a3. Natural size 10.75
mm X 6 mm; figs. 7, 8. Exterior and interior of partial ter-
gum, No. 8215 PRI, ABu-5a4. Natural size 7.5 mm xX 4 mm.
Sie, eBalanus wamtiiquus, (Meyer) o...0.c:06.lne ee Ag cece veces ccesens 152
Hypotype, No. 8216 PRI, ACH-19b. 9, 10. Lateral views of ex-
terior. Natural measurements; 14.3 mm in length at base; 9 mm
in width across base; 9 mm in height at carinal end; 10.5 mm
in height at rostral end. Fig. 11, view looking down on sand-
stone-filled orifice showing rhomboidal outline of peritreme.
Length about 9.5 mm; greatest width 7.8 mm.
164 BULLETIN 297
EXPLANATION OF PLATE 21
Figure Page
1-7. Balanus antiquus: (Meer) 2.22. .c.c...-ccc5.ece-ccepcessseees eee 152
Figs. 1-3, hypotype, No. 8216 PRI, ACH-19b. Fig. 1. Frontal view
of carinal end. Height about 9 mm; fig. 2. View of rostral end.
Height about 10.5 mm; fig. 3. View of base showing part of
basis and the termini of the inner tubules around the rim.
Length 14.3 mm; width 9 mm; figs. 4, 5. No. 8217 PRI, ACH-
19b3. Interior and exterior of rostrum. Height 10.5 mm; width
across base 6.5 mm; figs. 6, 7. No. 8218 PRI, ACH-19b1. In-
terior and exterior of carino lateral compartment. Height 8
mm; width across base 5.2 mm.
8,9. ? Balanus’ antiquus. (Meyer), ......0.:...:05...540..4. 48. 152
ACH-19b2, No. 8219 PRI. Interior and exterior of another carino-
lateral compartment. Height 8 mm; width across base 5.2 mm.
10. ? Hesperibalanus gosportensis Zullo 0oo0ooo.oooococccceccceeeeeeee 156
Holotype (?) tergum, 5. Natural size length about 4.3 mm;
maximum width about 2.5 mm. After Zullo, 1965, p. 133, who
stated that the tergum “is possibly the same as Balanus anti-
guus.” ;
11. Crucibulum antiquum Meyer ........0:5....0....0.0...c01.0.2 0... 152
Holotype = Balanus antiquus (Meyer). View of underside
(basis absent) showing “numerous striae” within, the termini
of the tubules around the basal rim, the opercular ? apparatus,
and the rhomboidal shape of the orifice from below. Natural
size about 12.5 mm length; 3.6 mm width. After Meyer. Com-
pare with figure 3 adjacent.
PLATE 21
BULL. AMER. PALEONT., VOL. 72
INDEX
NUMBER 297
Note: Light face figures refer to the page numbers. Bold face figures
refer to the plate numbers.
A
Aldrich collection ...... 144
antiquum,
Crucibulum <...... 21 152
antiquus,
iBalanusee ee 20,21 143, 152-157
Arcoscalpellum .......... 143, 147-149
B
bakeri,
Arcoscalpellum ...... 148, 147-149
PLAINES es castssscescsREne 143, 152-157
Bartonian Stage .......... 144
Butler County,
Alabama’ <...c:ccecnse~<ts 144, 149
€
Choctaw County,
PRVADAINIA® 4. cssceceessceess 143, 146, 151
choctawensis,
Arcoscalpellum 19 1438, 145-147
Citrus County, Florida 157
Claiborne, Alabama .. 152, 155, 156
Claiborne beds ............ 156
STUB Diese oeacoscce tet secasns 155
GHOUDT cencccsssosscocsseccss 152, 155
PAPACLET US bee. 5ccoccens 155, 156
Clayton Formation .... 144
Coal Bluff Member .. 144
COCOABSANG! merceeeceseoess 143, 147
conradi,
Arcoscalpellum ...... 149
D
Dantany Stage) cecccccc---s- 144
Darwin, Charles
ODOM? eset eorecnne 145, 156, 157
E
Eocene Epoch ............ 143, 144
eocenense,
Euscalpellum .......... 152
Euscalpellum .............. 148, 150-152
G
Geological Survey
PATA AMA av creccssseasseees 144, 145
Gosport Sand,
Alabama 22.60 <cssenseie 155, 156
gosportensis, Hesperi-
balanus? (i:csce 21 156, 157
H
Hall collection ............ 156
?Hesperibalanus ........ 156, 157
I
Inglis Limestone,
I Fig Ca 3 (0 (ee 157
Isney, Alabama .......... 143, 156
isneyensis,
Euscalpellum ...... 19 143, 150-152
J
Jackson Group ............. 143, 144, 155,
156
L
Lisbon Formation ...... 155
Ludiany Stage -2..2...... 144
M
McBryde Limestone .. 144
Maestrichtian Stage... 149
Matthews Landing
WE Well at sosasaene eee eee 144
Meyers Otto) cccccscccccssse 144, 158
Midway Group ............ 144
Monroe County,
Al aDAaMiay eeccsscccssecceces 155
Monterey, Alabama .... 144, 149
Moodys Branch
HIOEMALION: <...ccccsseezers 143, 147, 155
N
Naheola Formation .... 144
Newman, William A... 152, 156, 157,
158, 159
North Creek
Member | ..-..c.258-0c 143, 144, 146,
151, 156
ie}
Oak Hill Member ....... 144
OcalanGroupierccece 157
Oktibbeha County,
Mississippi .............. 149
165
INDEX
P U
RachutasMvarl sues 143 U.S. Geological
Paleocene Epoch ........ 143, 144, 149 SULVCY. 4cissvesseseetecvtee 144, 145
Pilsbry, Henry U.S. National Museum
AUISUIS UU Sieeeetceneseceees 145, 152, 158 (National Museum
Pine Barren of Natural History) 145
WoIMESCOME ees sereeseceec 144 unguiformis,
Porters Creek IBalanush cscce eee 156
HMOLMAtON ets ee 143, 144, 149
Vv
; Vincentown Sand ...... 143, 149
Ripley Formation ...... 149
TROSS9PATNOIG ee sccrssseeeeeee 152, 156, 157, W
159
Withers, Thomas
S 1a (2100 ai giererpey eric rroccc 152, 156, 158,
159
Salt Mountain Withlacoochee River,
PAMESIONE | 220.20ct8ecess0s 144 Rloridas ee 157
Scutella bed ................ 144 Wolf Creek, Alabama 144
Shubuta Member ........ 143 i
Silas, Alabama ............ 143, 146, 11, Y
56
Sparnacian Stage ........ 154 Vaz0e Clay. 143, 155
GrOUD Pisce. cece cosets 143, 146, 151,
7 156
Thanetian Stage ......... 144 7é
toulmini, Arcoscal-
pellumye ee 20 143,147-149 Zullo, Victor
Toulmin, Lyman D. .... 143, 144, 149, VAMOUSE ....shterseteee 152, 156, 157,
153, 159 158, 159
166
| ict, PER ee. | !
. Tnlears ea ous a.
LII.
LIT.
LIV.
LV.
LVI.
LVII.
LVIII.
LIX.
LX.
LXI.
LXII.
LXIII.
LXIV.
LXV.
LXVI.
LXVII.
LXVIII.
LXIX.
LXX.
LXXI.
LXXII.
(USES PER BALD IR We Noens) oe) fol (ee ae ee ee ee
New Zealand forams, Stromatoporoidea, Indo-Pacific, Mio-
cene-Pliocene California forams.
(Nos. 237-238). PMCS, Moy ge 1 U5) /4) ol Kis an a UC ae eee an eee
Venezuela Bryozoa, Kinderhookian Brachiopods.
(Nos. 239-245). CalObapovniyy EXO) M0) IG) tea ie teeretee te eedeeretee see tet eet
Dominican ostracodes, Lepidocyclina, mollusks.
(Nos. 246-247). CASI yo ov ege C0) 9) 0) fg ae salle te eee ae
Cenozoic corals, Trinidad Neogene mollusks.
(Nos. 248-254). CME Pepa 2A CRI al A telnet een Se
Forams, North Carolina fossils, coral types, Cenozoic
Echinoids, Cretaceous Radiolaria, Cymatiid gastropods
(Nos. 255-256). SO DDO Zi pisses oe ee easton ccsce
Jurassic ammonites.
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BWELETING V7. ar
HARVARD
OF tot a RIT Y,
AMERICAN
PALEONTOLOGY
(Founded 1895)
Vol. 72
No. 298
THE ARCHAEDISCIDAE OF THE FRAILEYS FACIES
(MISSISSIPPIAN) OF CENTRAL KENTUCKY
By
R. G. BrowneE, J. W. BAxTER, AND T. G. RoBERTs
1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
PALEONTOLOGICAL RESEARCH INSTITUTION
1976-1979
DP RE SUISEB Ty ose ce cea ee se Se) ee HAROLD E. VoOKESs
WICE=P RESIN Nye nee roe LIOR SN es PE ee ee DUANE O. LeRoy
SECRETARY (5 oor te en a a Bate See ee eee Puitie C. WAKELEY
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BOE TINS
OF
AMERICAN
PALEONTOLOGY
(Founded 1895)
Vol. 72
No. 298
THE ARCHAEDISCIDAE OF THE FRAILEYS FACIES
(MISSISSIPPIAN) OF CENTRAL KENTUCKY
By
R. G. Browne, J. W. BAXTER, AND T. G. RoBERTs
October 24, 1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
Library of Congress Card Number: 77-85464
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CONTENTS
Page
JAVON CLE ene er ee ee eee eee 171
VIRUS SEIU OY 171
AINE TORT a VES Cc 172
(SHEEN HES SE a) OP ae a a ON ne ee 0 ee 173
Searen ai eareey ad OV CUENGEYG FC GTOS NY cece cece a a ee 73
Criteria for classification of the Archaediscidae: 2.222.022 osoocc conc ececsesee 175
Genenicgs Crite rake ae ee ne eee eee ee ee ee 175
Subgenerigu@ritentay ces-ccc ee eee ee ee 176
SpEeciationmande Collen gg Group sis cee eee eee ee ree eee 177
BVO ET OM ay Cale Speech. ee eee eee eee 177
OG ae CLO ri gee ere eek ee | Bn eed ee ee 178
Systema ti cus aleontolOcy anes eee... SMa ene eee ee eee ee 178
Family Archaediscidae
Genusi Arch aedss cuge ites. .... 05s oo eee eee et RT cen ae 178
Subgenus Aichaedts cus) cc 31 Meena Sie ey ee 179
Subpenuis, Hiemtanceacdtscus: (exp tee eee eee eee ee eae 188
Subpemuss criewmiarcnaed: sci g aee 189
INE WwarsUD een Use iee.2. 08. ob Ri Bearer eaee eee Tee I ee 194
Genus, NOdOsanchaedts cus rc... .s eee ee 2 asco r h ek 195
Subgenus Vodasperddiscus | seas ee ee ee 195
Sabgenus=V courchacdiscus (ee wee ee Be ee 198
Subgenus Aster oarchacdts cis: se mere ee re 206
GGenUsieA Ti TVATCHOCAUS CIS). sa ccssit sete eer te eta Re sae acs et a erase aoe 212
Subgenus: AU je seis So eee eee a eee esceneettenccact 213
Subgeenusy 2 wh7s pio dts cts, ee ee ee ee 214
IRGL eT CN COS hee Beane rar eS es LA a ee a = els
LEED cee ee a ee ee a eee pk A eer a Sen Be 221
LINO OS cheer eee ee EE ep ee ye
ILLUSTRATIONS
Page
textaniouneslanCorrel atio my Ghia rts seer seetee erence, Sere ee eee nee es ancce easton tree 174
‘ables — "Classification of the Arehaediscidae si scccscceecrerseeeseeeeece eee ee 178
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THE ARCHAEDISCIDAE OF THE FRAILEYS FACIES
(MISSISSIPPIAN) OF CENTRAL KENTUCKY
R. G. BrowneE,* J. W. BAxTER,} AND T. G. RoBertst
ABSTRACT
Samples of washed shale collected from the Fraileys facies of the Big
Clifty Formation (Chesterian) in Central Kentucky revealed the presence of a
unique free-form foraminiferal fauna. A study made from thin sections of the
calcareous forms of this fauna was reported at the generic level by Browne and
Pohl (1973). The present report covers a study, from the same fauna, of forms
belonging to the family Archaediscidae. They are discussed at the specific
level.
Representatives of two subfamilies are recognized as Archaediscinae and
Ammarchaediscinae. The Archaediscinae are assigned to two genera — Archae-
discus and Nodosarchaediscus and the Ammarchaediscinae to one, Ammarchae-
discus.
The authors have availed themselves of the term subgenus to describe those
forms which they consider to be monogeneric because they represent morpho-
logical changes showing an evolutionary development in chronological sequence
and transitional forms exist.
Three subgenera are placed in the genus Archaediscus — Archaediscus,
?Hemiarchaediscus, and a new subgenus, described but unnamed. Three sub-
genera are placed in the genus Nodosarchaediscus — Nodasperodiscus, Neo-
archaediscus, and Asteroarchaediscus. Three subgenera are placed in the genus
Ammarchaediscus — Ammarchaediscus Tubispirodiscus, and A.
A total of 27 species are described, four of which are new. The original
descriptions are given. The geographic distribution and the stratigraphic range
are also recorded.
INTRODUCTION
The discovery of a prolific, free-form microfauna from the
Fraileys Shale facies of the Big Clifty Formation in central Kentucky
was revealed in a preliminary note in 1968 (Pohl, Browne, and
Chaplin). This excellently preserved faunule contains representatives
of 16 families and approximately 37 genera which include an un-
usual assortment of calcareous foraminifers. The stratigraphy of the
Fraileys Shale and the generic affiliations of the calcareous forms
were subsequently discussed in some detail (Browne and Pohl,
1973). The present authors are proceeding to systemize taxonomical-
ly related calcareous forms beginning, in this report, with the Archae-
discidae.
This paper is the outgrowth of a larger effort directed toward
the recognition of time-related taxa within the type Mississippian
area and adjacent portions of the Illinois Basin and the establish-
ment of criteria for their use in the biostratigraphic zonation of the
type Mississippian. This ongoing research is based on study of col-
*4007 Elfin Avenue, Louisville, Kentucky, 40207; +Illinois State Geological
Survey, Urbana, Illinois, 61801; {University of Kentucky, Lexington, Kentucky,
40506.
172 BULLETIN 298
lections that now comprise approximately 7,000 thin sections of type
and reference rock material and 9 free-form collections of varying
productivity.
The Fraileys fauna was discovered by Dr. E. R. Pohl of Horse
Cave, Kentucky, who recognized the biostratigraphic importance
of the calcareous Foraminifera and especially that of the Archae-
discidae. The senior author joined Dr. Pohl in the early phases of
the project. Baxter became involved later, first through consultation
on calcispheres (which are a conspicuous component of the Fraileys
fauna), and later when he joined the study of the wider aspects of
Mississippian biostratigraphy. Since Dr. Pohl’s death in 1973, and
the addition of Roberts, Pohl’s work has been continued in appre-
ciation of his early efforts and has been sustained by the enthusiasm
he engendered.
This report is based upon the examination of approximately
700 oriented thin sections cut from free-form archaediscids, washed
from shale samples. In most cases the orientation is axial and speci-
mens are sectioned to reveal the proloculus. External views of the
specimens, photographed prior to sectioning, were used for compara-
tive purposes. No external views are reproduced for this report.
Any study of calcareous Foraminifera and the Archaediscidae in
particular naturally reflects the enormous efforts of specialists work-
ing in Western Europe and the USSR. In the section on Systematic
Paleontology the reader is referred to existing translations of orig-
inal descriptions where such are available (as in Ellis and Messina,
1940-1964). Other translations, obtained during the course of our
studies, are given.
Credit for translation of Russian literature used in this report
is gratefully accorded to Dr. Leonard Latkovski, Professor Emeritus,
Department of Foreign Languages, Bellarmine College, Louisville,
Kentucky.
ACKNOWLEDGMENTS
We wish to especially acknowledge Professor Raphael Conil of
the Institute of Geology and Geography, University of Louvain,
Louvain- la Neuve, Belgium. His continued interest in our efforts
and his invaluable advice concerning some taxonomic assignments re-
ported here are appreciated. We are also indebted to Dr. M. V.
Vdovenko of the Institute of Geological Sciences, Academy of Sci-
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 473
ences, Ukrainian SSR, for her enlightening correspondence concern-
ing the comparison of our material with faunas of similar age in the
USSR. Finally, we acknowledge Dr. Paul Brenckle of Amoco Pro-
duction Company, Tulsa, Oklahoma. His review of our material and
critical review of this paper were most helpful.
STRATIGRAPHY
The archaediscids described herein were recovered from 11 feet
(3m) of grey-blue shale exposed for 200 feet in the road ditch and
bank on the west side of the Broadford Church Road, 200 feet south
of the junction with KY 1214 at Broadford, Grayson County, Ken-
tucky. The location is in the northwest quarter of section #11, K 42,
NJ 16-8, Evansville sheet of the Carter Coordinate System, Millers-
town Quadrangle, GQ-417, Kentucky (Browne and Pohl, 1973, p.
176). The stratigraphic details of the Broadford exposure are dis-
cussed by Pohl (im Browne and Pohl, 1973, pp. 175-190). We can
add little to this previous discussion beyond placing the present
stratigraphic classification in its historical perspective and showing
relationship with adjacent regions.
STRATIGRAPHIC CLASSIFICATION
The microfauna occurs in shale at the base of the Big Clifty
Sandstone of Hombergian (Middle Mississippian) age (Text-fig. 1).
The Big Clifty of central Kentucky is considered a formation by
many authors (Browne & Pohl, 1973; Schwalb, 1975) but as
a member of the Golconda Formation on the recent geologic map of
the Millerstown Quadrangle (Moore, 1965). The Big Clifty occupies
a position below the Haney Limestone and above the Beech Creek
Limestone and has a facies relationship with the Fraileys Shale to the
west (McFarlan, et al., 1955; Swann, 1963). The productive strata
at Broadford contains sparse stringers of crinoidal debris and occa-
sional limestone lenticles and thus resembles typical Fraileys Shale.
Pohl (in Browne and Pohl, 1973) referred this fauna to the Fraileys
“facies” of the Big Clifty Sandstone Formation. This sandstone at
one time was correlated with the Cypress Sandstone to the west and
was once called “Cypress” (Butts, 1917; McFarlan, 1943) but this
miscorrelation was corrected (Dana and Scobey, 1941; Swann and
Atherton, 1948) and the name “Big Clifty” (Norwood, 1876) re-
vived for the sandstone equivalent of the Fraileys.
BuLLETIN 298
174
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uBIdZ1aqMoyy AY} FO SYIOI FO UOljEoIFISse[D IYydess1j}e13g —"]T 9INBIZ-3xa 7,
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(0261) UMOJSIIIII | PJP! 4 1D09 MM JO 4apsog a |8Ia
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SIONIT 11
AMONLNAM
ARCHAEDISCIDAE: BROWNE, BAxTER, & ROBERTS 75
In his classification of Chesterian rocks, Swann (1963) assigned
the Fraileys, Haney, Hardinsburg, and Glen Dean to the Hom-
bergian Stage, derived from and roughly equivalent to the Homberg
Group of Weller and Sutton (1940). The stage differs from the group
in the exclusion of the Cypress Sandstone and Beech Creek Lime-
stone. As pointed out by Vincent (1975), the assignment to a time
stratigraphic unit is not entirely justified because the time equivalen-
cies of the boundaries involved have not been established. The simi-
larity of the microfauna of the Fraileys with that of the underlying
Beech Creek caused Browne and Pohl (1973) to question the ex-
clusion of the latter from the Hombergian Stage.
CRITERIA FOR CLASSIFICATION OF THE
ARCHAEDISCIDAE
The classification of the Archaediscidae followed here is essen-
tially that of Pirlet and Conil (1974) but in the application of the
system to our material we find that certain departures are either
required by laws of priority or seem advantageous. The family
Archaediscidae evolved from ancestral forms by the addition of a
clear, more or less radial wall on an ancestral, dark, microcrystalline
wall. The recognition of the subfamilies Archaediscinae, Ammarchae-
discinae, and Tournarchaediscinae (Table 1) is based upon charac-
teristics that appear to have been inherited from ancestral stock.
Thus Ammarchaediscinae probably inherit planispiral coiling from
Pseudoammodiscus and Archaediscinae and Tournarchaediscinae,
a variable plane of coiling from Brunsia and Brunsuna respectively.
The Tournarchaediscinae, not represented in our material, are further
characterized by the presence of pseudochambers.
The simple acquisition of a radial layer may in some Foramini-
fera (e.g. Tetrataxis) be of no more than specific importance. How-
ever, in the Archaediscidae it marks the beginning of profound
chronologically related evolutionary changes that logically lead to
the revised generic divisions of Pirlet and Conil (1974) and at vari-
ous stages of development to the establishment of subgenera. More
subtle evolutionary changes permit the recognition of stages within
individual species.
GENERIC CRITERIA
Archaediscidae of the Fraileys belong to the subfamilies Archae-
176 BuLLeETIN 298
discinae and Ammarchaediscinae. For each subfamily the recognition
of genera is based upon the presence or absence of occulusions in the
form of nodes and stellate central flarings or stellate central flaring.
Thus in axial thin sections Archaediscus and Ammarchaediscus are
characterized by free lumina throughout the test and Nodosarchae-
discus by occluded lumina. Similar occlusions are known among the
Ammarchaediscinae, (?Permodiscus Conil and Pirlet, in Pirlet and
Conil, 1974) but such forms are not present in the Fraileys material.
SUBGENERIC CRITERIA
Classification at the subgeneric level is based on the recognition
of stages in the evolution of the wall structure and, for subgeneric
Nodosarchaediscus, of nature of the occlusions of the lumina.
Throughout the range of the Archaediscidae there is in each sub-
family a progressive diminution in the development of the ancestral
dark inner wall layer. Archaediscidae characterized by thick, dark,
microgranular inner layers are primitive forms (V1b in Belgium) not
present in the Fraileys. Representatives of Archaediscus and Am-
marchaediscus have reached a stage of evolution at which the inner
layer either ranges from both poorly developed to almost impercepti-
ble, as in advanced stages of both A. (Archaediscus) and Amm.
(subgenus A), or is totally absent as in Amm. (Tubispirodiscus).
We differ from Pirlet and Conil (1974) in our recognition of A.
(? Hemiarchaediscus) as a valid subgenus occupying a position paral-
lel to Amm. (Tubispirodiscus). A. (?Hemiarchaediscus) differs
from the original description of Hemiarchaediscus Miklukho-Maklay
(1957) in that the wall is a single radial layer lacking a dark in-
terior layer.
Nodosarchaediscus first appears in the Visean (V2b8) of Bel-
gium (Pirlet and Conil, 1974) and is known in the Harrodsburg
Limestone of Valmeyeran (Middle Mississippian) age in Kentucky,
(Baxter, Browne and Roberts, in press). Earliest forms, (Nodo-
sarchaediscus), with simple elevated nodes on the lumen floor, have
a dark inner layer that is no more than moderate in development,
and the importance of the inner layer is progressively diminished
in younger forms. Fraileys representatives include (Neoarchae-
discus) with central stellate flarings, (Nodasperodiscus) with nodes
and central flaring, and (Asteroarchaediscus) with closed lumina
throughout most of the test. We differ from Pirlet and Conil (1974)
in recognizing a priority for (Neoarchaediscus) over (Asperodiscus).
ARCHAEDISCIDAE: BROWNE, Baxter, & ROBERTS 177
Some uncertainty persists in the literature concerning the dif-
ferentiation between the subgenera Neoarchaediscus (Asperodiscus
of Conil), Nodasperodiscus and in some instances Asteroarchae-
discus. Our concept of (Neoarchaediscus) requires central, confused,
stellate coiling followed by at least 1% coils open and free of nodes.
In (Nodasperodiscus) stellate coiling is followed by final coils
in which the lumina are partially open (reduced by nodes) al-
though the ultimate coil may be completely free as in Nodaspero-
discus (Nod.) minimus. In (Asteroarchaediscus) the lumina
throughout the test are generally completely closed along irregular
crenulations but the final coil may be free or partially free as in
Nod. (Asteroarchaediscus) postrugosus.
SPECIATION AND COILING GROUPS
Where the recognition of the various coiling groups defined by
Pirlet and Conil (1974) is applicable our speciation is based upon
a combination of that feature and traditional biometric measure-
ments. Thus for Archaediscus we recognize groups of species with
aligned, stilus; oscillating, chernoussovensis; sigmoidal, karrert; im-
perfect sigmoidal, gigas; and initial sigmoidal, krestountkovi coiling.
In the description of the various species of this report where the
coiling pattern is applicable it is listed under the term — “Coiling”.
In the subgenus Asteroarchaediscus which represents the final stage
of evolution of the family the coiling pattern is somewhat zigzag and
normally little apparent. Therefore, the form of the test produced
by the coiling is substituted in this subgenus and is listed under the
term “test form” (e.g. — flat, lenticular, round).
EVOLUTIONARY STAGES
Beginning with the ancient Glomodiscus the coiling habit of the
Archaediscinae shows a marked evolutionary tendency to become
more evolute in character. This tendency is operative at the species
level and in Archaediscus is in company with and accomplished by
morphological changes that permit the recognition of evolutionary
stages that have chronologic value (Pirlet and Conil, 1974). We dif-
fer from Conil in that we prefer to consider these characteristics as
simply evolutionary stages (involutus, concavus, angulatus, evolutus,
and tenuis) rather than critera for subspecies.
178 BULLETIN 298
Table I — General Criteria for Classification of the Archaediscidae (from
Pirlet and Conii, 1974)
Genera
lumina nodes and
subfamilies free stellate flaring
Tubular Chamber coiling
smooth, not streptospiral
divided. Wall
porous Archaediscinae Archaediscus Nodosarchaediscus
coiling
planispiral
Ammarchaediscinae Ammarchaediscus to be named
Tubular chamber coiling
with pseudo- streptospiral
chambers. ;
Wall porous. Tournarchaediscinae Tournarchaediscus unknown
CORRELATION
The fauna is characterized by Archaediscus (Archaediscus) at
the angulatus stage, Archaediscus (?Hemiarchaediscus) approach-
ing the tenuis stage, and fairly abundant small, species of
the subgenus Nodosarchaediscus (Asteroarchaediscus): parvus,
rugosus, postrugosus, and syzranicus. The population also includes
Nodosarchaediscus (Nodasperodiscus), numerous Nodosarchaediscus
(Neoarchaediscus) and Ammarchaediscus (Tubispirodiscus) and
(subgenus A.). This assemblage, while close to the Namurian in age,
is in its overall aspect indicative of late V3c reported by Browne
and Pohl (1973).
SYSTEMATIC PALEONTOLOGY
Family Archaediscidae Cushman, 1928, emend. Conil and Pirlet, 1974
Fusulinina with a proloculus and coiled tubular chamber, usually not
divided, but may possess pseudo-chambers or polar septa. The first coils are
involute, except among very rare forms. A calcareous wall comprises a dark
internal microgranuiar layer, tending to disappear in the more evolved forms,
and a clear, more or less porous radial layer. (Pirlet and Conil, 1974, p. 252).
Subfamily Archaediscinae Cushman, 1928, emend. Conil and Pirlet, 1974
Archaediscidae without internal divisions into chambers or pseudocham-
bers; coiling streptospiral. Wall formed of a dark, microgranular internal and
external radial layer. The internal layer, pronounced in the primitive forms,
tends to disappear among those more evolved. (Pirlet and Conil, 1974, p. 254).
Genus Archaediscus Brady, 1873, emend. Conil and Pirlet, 1974
Type species: Archaediscus karrert Brady, 1873.
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 179
Diagnosis. — Archaediscinae possessing free lumina, without nodosities, or
stellate flaring. Internal dark layer pronounced to imperceptible. The external
radial layer developed in the first coils only or throughout the test. Coiling
involute to evolute (Pirlet and Conil, 1974, p. 254).
Subgenus Archaediscus Conil and Pirlet, 1974
Type species: Archaediscus karreri Brady, 1873.
Diagnosis. — Archaediscus with the dark internal layer moderately to
feebly developed, without lateral corner fillings and with the radial layer
completely enveloping all the coils except in the immediate vicinity of the aper-
ture. The more ancient forms are involute and the floors of the lumina are
convex; the evolved forms tend to become evolute and beginning with the
first coils their floors are concave. The walls of certain very evolved forms
tend to become very thin without any epaulets or covering. The floors of the
lumina then become convex again (Pirlet and Conil, 1974, p. 258).
Archaediscus (Archaediscus) cf. absimilis (Sosipatrova), 1962
Pl. 22, figs. 1-3
Planoarchaediscus absimilis Sosipatrova, 1962, pp. 58, 59, pl. 5, figs. 3, 4; Sosi-
patrova, 1966, pp. 24, 25, pl. 1, figs. 5, 6.
Holotype. — Institute of Geology of Arctic Regions, No. 716/14.
Original description.—The shell is of small dimensions, involute except
for the final evolute coil, with parallel lateral sides. The ratio of width to
diameter is 0.31-0.33. The diameter of the shell is 0.17-0.32 mm, the width
0.056-0.096 mm. The number of coils is three to four. The proloculus is spherical
and relatively large with a diameter of 32u. The whorls of the second tubular-
shaped chamber are freely and glomospirally wound with a displacement of
10-15° from the axial plane. The final coil is planispiral and flat. The height
of the opening in the last coil is 0.020 mm. The wall consists of an exterior,
bright, glassy radial type layer and an interior dark, granular layer. The thick-
ness of the wall is 6-8x.
The identifying characteristics of this species are:
1. asymmetric and involute coiling
2. small number of coils
3. rather large proloculus
Remarks and comparisons.— On the basis of coiling, this species, as here
described, is close to Planoarchaediscus abseus (?), n. sp. from which it is
distinguished by its smaller dimensions, smaller number of coils and large
proloculus.
Diagnosis. — Test small, discoidal with approximately plano-
parallel sides, broadly rounded periphery and slightly uneven sur-
face; coiling involute except for the final whorl with interior coils
tightly wound and final two to three approaching the planispiral
plane; layering of the wall which appears as parallel bands, extends
the length of the test except for the final evolute whorl; flat floored
lumina increase in size and breadth and at a rapid rate; wall is
bilayered with an exterior bright radial layer and a poorly developed
interior dark microgranular layer.
Measurements. — (Based on three specimens). Number of volu-
tions (based on two specimens): 5-6. Diameter: 185.00-262.50y.
180 BULLETIN 298
Width (based on two specimens): 72.50. Ratio W/D: 0.276-0.32.
Proloculus: 20u. Height of lumen last volution: 18.30-22.50y. Peri-
pheral wall thickness: 6.25-11.25p.
Coiling. — Aligned.
Cowling stage. — Angulatus.
Stratigraphic range. — USSR-Baschkirian (lower part of Maka-
rov horizon).
Remarks. — The distinguishing features of this species are the
almost flat plano-parallel sides, the broadly rounded wide outline of
the final whorl and the pronounced lateral thickening edging the
sides of the test. Sosipatrova’s 1962 and 1966 descriptions and il-
lustrations differ. The original description notes three to four coils,
a proloculus of 324, and wall thickness of 6-8u. The 1966 description
gives three and a half to five coils, a proloculus size of 24-25» and
wall thickness of 9-15. Our specimens approach one of Sosipatrova’s
original illustrations (1962, pl. 5, fig. 4) in which the parallel band-
ing characteristic of our forms is faintly discernible. However, the
proloculus is smaller and the coils are numerous in our specimens.
The wall thickness of our specimen encompasses the range of both
of Sosipatrova’s descriptions.
Archaediscus (Archaediscus) chernoussovensis Mamet, 1966 PI. 22, fig. 4
Archaediscus chernoussovensis Mamet subsp. angulatus Conil and Pirlet (in
Austin, Conil, Groessens, and Pirlet), 1974, pl. 3, figs. 14-15.
Diagnosis. — Test small, disc-shaped, becoming moderately con-
vex at center of test, periphery broadly rounded, surface uneven;
coiling is streptospiral throughout and oscillating, characterized by
two definite breaks in the pattern of deflection; concave to flat-
floored lumina increase gradually in size and the floors extend to
the wall of the succeeding coil forming prominent angular contacts
along the entire length of test and pseudo-stellate structure in the
center; wall has a well-developed porous layer and a poorly defined
inner dark layer.
Measurements. — (Based on one specimen). Number of volu-
tions: 5?. Diameter: 232.50. Width: 88. Ratio W/D: .382. Prolo-
culus: 17.50u. Height of lumen last volution: 26.25. Peripheral wall
thickness: 8.75y.
Coiling. — Oscillating.
Coiling stage. — Angulatus.
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 181
Stratigraphic range.— North America — Visean (V2)-early
Namurian, USSR-Visean, Belgium-Visean (V2-V3), France-Visean
(V3):
Remarks. — Mamet in Mamet, Choubert, and Hottinger (1966)
changed the name of Archaediscus karreri Brady of Rauzer-Cher-
noussova (1948a, p. 230, pl. 5, figs. 10, 11) to A. chernoussovensis.
The distinguishing feature of A. chernoussovensis, according to
Mamet, et al., 1966, is its oscillating mode of coiling. Pirlet and
Conil (1974, p. 259) excluded figure 10 as not conforming to Mamet’s
definitive diagnosis, thus leaving figure 11 as the type specimen.
Our form compares well with the type specimen except for a smaller
ratio of width to diameter and the pronounced angularity made by
the flat floors and their junction with the succeeding whorls. This
angularity represents an evolutionary trend in the archaediscids. Be-
cause Pirlet and Conil have not yet described the subspecies “angu-
latus,” the authors can not make positive identification.
Archaediscus (Archaediscus) conili Browne, n. sp. Pl. 22, figs. 5-7
Archaediscus aff. infantus Shlykova, Conil and Lys, 1964, pp. 116, 117, pl. 17,
fig. 319.
Holotype. — Raphael Conil.
Test lenticular and flattened on the sides. Coiling feebly oscil-
lating.
Whorls: Four.
Diameter: 130u. Width: 70u.
Ratio W/D: 0.53.
Description. — Small species with few whorls. Fibrous layer,
moderately developed, measuring about 25, in the axial part of test.
The internal dark layer is well developed. The proloculus measures
20. The profile is lightly deformed by the oscillations of the coiling.
Comments and differences. — Our form differs from the species
described in the USSR by proportionally larger lumina, proloculus
of greater size and smaller dimensions. We lack sufficient material,
however, to make a careful comparison with Shlykova’s species.
Diagnosis. — Test small, lenticular, surface smooth; first two
whorls involute with final whorl entirely free; coiling streptospiral in
the initial whorls; later coils oscillate about a plane as in Archae-
discus (Archaediscus) chernoussovensis; lumina with slightly convex
floors increase in size at a continuous rate and become progressively
182 BuLLETIN 298
broader in relation to height; wall is composed of two layers, a
fiberous outer layer and a poorly developed inner microgranular
dark layer.
Measurements. — (Based on four specimens). Number of volu-
tions (based on two specimens): 4-5. Diameter: 132.50-200u. Width:
66.25-75u. Ratio W/D: 0.375-0.50. Proloculus: 18.32-23.75y. Height
of lumen last volution: 12.50-17.50u. Peripheral wall thickness: 6.25-
8.75.
Coiling. — Oscillating.
Coiling stage. — Angulatus.
Stratigraphic range. — Belgium — Visean (V3). North America
— Visean (late V3C), this report.
Remarks. — Our specimens compare favorably with the form
described by Conil and Lys with its smaller size, manner of coiling,
larger proloculus, and thinner wall than those of Shlykova (1951).
Conil and Lys gave a ratio of width to diameter of 0.53 for their one
specimen, but the ratio of the specimen illustrated on plate 17, figure
319 is 0.42 which is within the size range of our forms.
The authors believe that neither Shlykova’s description of A.
infantus nor her illustrations bear resemblance to our forms.
We consider this form to be a new species which we are naming
Archaediscus conili in honor of Dr. Raphael Conil who first described
the form.
Archaediscus (Archaediscus) infantus Shlykova, 1951 Pl. 22, figs. 8, 9
Archaediscus infantus Shlykova, 1951, p. 172, pl. 6, figs. 4, 5, Grozdilova (in
Dain and Grozdilova), 1953, pp. 98, 99, pl. 3, figs. 6, 7.
Not Archaediscus aff. infantus Shlykova, Conil and Lys, 1964, pp. 116, 117, pl.
17, fig. 319.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
No. 2220.
Original description. —'The shell is involute, lentil shaped with nearly flat
parallel sides and narrowly rounded periphery. The ratio of width to the
diameter is 0.50-0.58. Coils number four to six, most frequently four to four
and a half. The dimensions are small; the width is equal to 0.10-0.16 mm, the
diameter 0.19-0.30 mm. The coiling: In the first three coils of the second chamber
the central plane of each consecutive coil is turned in respect to the previous
or preceding coil by an angle of 90 degrees so that, in section, the second coil
in circular form is seen to surround the first coil. In the final two to three
coils the central plane of each coil is slightly displaced in relation to the pre-
ceding one to the same side by 10 degrees - 15 degrees. Sometimes the central
plane of the final coil may be displaced in opposite direction to the common
direction of coiling of the exterior coils.
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 183
The height of the lumina varies from 13 to 30 in the last coils. They are
relatively wide with commonly slightly convex or flattened bases.
The wall is smooth with a thickness from 10-154.
Remarks. — The form of the test with its almost flat sides and the slightly
displaced coiling planes of the final whorls indicates this species is close to
Archaediscus krestounikovi Rauzer-Chernoussova, but it differs from the latter
in the manner of coiling of the interior coils, by smaller diameter of the test,
by the larger ratio of width to diameter, by the average lower clearances of
the chamber and by the thick wall.
Diagnosis. — Test small, lenticular with slightly convex sides,
surface smooth; initial whorls involute, becoming partially evolute
with the final whorl entirely free. Coiling streptospiral with first
three whorls tending to encircle the proloculus in axial section; ex-
terior whorls are arranged in sigmoidal fashion except for the final
whorl which departs in an opposite direction from the preceding
whorls at an angle of approximately 45 degrees; lumina with slightly
convex to flat floors increase progressively in size becoming broader
in relation to height; wall is composed of two layers; a fibrous outer
layer and an inner microgranular dark layer.
Measurements. — (Based on two specimens). Number of volu-
tions: 4?-5. Diameter: 150-172. Width: 78-82. Ratio W/D: 0.476-
0.52. Proloculus (based on one specimen): 11.50u. Height of lumen
last volution: 14y. Peripheral wall thickness: 7.00-8.50y.
Coiling. — Imperfect sigmoidal.
Coiling stage. — Angulatus.
Stratigraphic range. — USSR — Visean (late V3). Belgium —
Visean (late V3a).
Remarks.— Our specimens compare favorably with Shly-
kova’s type. They are slightly smaller in size but are similarly pro-
portioned and have a small proloculus.
Shlykova’s description referred to the central plane of the final
coil being “sometimes” displaced in opposite direction to the common
direction of coiling of the exterior coils. She also noted the difference
in the coiling manner of the interior coils from that of A. krestount-
kovt Rauzer-Chernoussova. Her illustrations show the displacement
of the final coil which we consider to be representative of imperfect
sigmoidal type of coiling (Pirlet and Conil, 1974).
The differences between this species and Archaediscus (Archae-
discus) comli, n. sp. are referred to under the description of the latter
species.
184 BULLETIN 298
Archaediscus (Archaediscus) krestovnikovi Rauzer-Chernoussova, 1948
Pl. 22.) 7185-7105 18:
Archaediscus krestounikovi Rauzer-Chernoussova, 1948b, pp. 10, 11, pl. 2, figs.
18-20; Bogush and Yuferev, 1962, pp. 202, 203, pl. 9, fig. 7; Mamet, 1973,
pl. 4, figs. 8, 11.
Not Archaediscus krestovnikovi Rauzer-Chernoussova, Shlykova, 1951, pl. 5,
figs. 8, 9; Brazhnikova and Vdovenko, 1973, pp. 232-234, pl. 37, figs. 15,
16: 19520:
Archaediscus krestovnikovi subsp. krestovnikovi Rauzer-Chernoussova, Conil
and Lys, 1968, pp. 510-512, text-fig. 2
Archaediscus krestovnikovi var. krestounikovi Rauzer-Chernoussova, Grozdilova
and Lebedeva (in Dain and Grozdilova), 1953, p. 95, pl. 2, figs. 17-19;
Bozorgnia, 1973, pp. 115, pl. 22, figs. 3, 4.
Not Archaediscus krestounikovi var. krestovnikovi Rauzer-Chernoussova, Conil
and Lys, 1964, pp. 120, 121, pl. 18, figs. 345-351.
Archaediscus krestounikovi Rauzer- Chernoussova forma typfica, Bogush and
Yuferev, 1966, pl. 11, fig. 13.
Archaediscus krestounikovi var. koktjubensis Rauzer-Chernoussova, 1948b, pp.
10, 11, pl. 3, figs. 1-3; Shlykova, 1951, pl. 5, fig. 11.
Archaediscus koktjubensis Rauzer- Chernoussova, Conil and Lys, 1964, pp. 119,
120, pl. 17, figs. 338-340; Mamet, 1973, pl. 4, figs. 1-7.
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, fig. 19, No. 2834/42.
Original description. — (Translated from the Russian in Ellis
and Messina, supplement No. 2, 1958).
Diagnosis. — Test small, lenticular with uneven surface, flat to
moderately convex sides and moderately rounded to slightly angled
periphery; mode of coiling changes markedly from aligned or with
slight oscillation of the outer whorls to sigmoidal in the inner whorls
which are involute and form thickened coalescing walls; the outer
whorls become evolute and approximately or completely planispiral;
lumina open and free, expanding at a rather rapid rate and changing
in shape from spherical to semi-lunular; wall composed of an outer,
clear, coarsely fibrous wall and an inner poorly developed thin, dark
microgranular layer.
Measurements. — (Based on two specimens). Number of volu-
tions: Six. Diameter: 172.50-192.50n. Width: 61.25-75.00u. Ratio
W/D: 0.375-0.40. Proloculus: not determinable. Height of lumen
last volution: 16.25-20.00u. Peripheral wall thickness: 6.25-7.50p.
Coiling. — Initial sigmoidal only.
Coiling stage. — Angulatus.
Stratigraphic range.— North America — Visean (V3) —
Namurian (Morrowan R?). USSR — late Visean (Tula-Serpukhov).
Belgium — Visean (V2b-V3b). Iran — Visean (V2-V3).
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 185
Remarks. — A number of authors have, in the past, incorrectly
assigned the species Archaediscus stilus Grozdilova and Lebedeva
(in Dain and Grozdilova, 1953) to this species. Conil and Lys
(1968), upon examination of the holotype, separated these two
species by their mode of coiling. They recognized the mode of coiling
of A. krestovnikovi to be that of the “variety” A. krestoumkouvt
koktjubensis, intermediate between oscillating and aligned and sig-
moidal only in the inner whorls.
Our specimens average approximately 3/4 the size of the mini-
mal dimensions given by Rauzer-Chernoussova for the type species.
The proportions, however, are the same as ours. Bozorgnia’s (1973)
forms, on the contrary, have a range with minimal dimensions about
equal to the maximum of the type and a proportionally greater ratio
of width to diameter.
Archaediscus (Archaediscus) miklukhomaklayi Browne, n. sp.
Pi. 22. figs. 12. 13
Hemigordius schlumbergeri (Howchin), Miklukho-Maklay, 1953, p. 129, pl. 6,
Teas Big
Not Cornuspira schlumbergi Howchin, 1895, pp. 195-196, pl. 10, figs. 1-3.
Holotype. — Repository not located.
Original description. — Shell lenticular, the first chamber is spheric, the
second tube type. Coiling initially archaediscid type, then flat spiral. The wall
is calcareous, brownish, sometimes dark. Diameter: 0.15-0.25 mm. Width: 0.05-
0.10 mm.
Diagnosis. — Test free, with the greatest thickness through the
axis of revolution; composed of a proloculus (not clearly defined in
our form) followed by a second tubular chamber which is initially
streptospirally coiled, then spiral with the final one to two whorls
evolute; lumina, in axial view, are open, semicircular in shape, in-
crease gradually in height and have flat-floored bases which extend
to the edges of the wall; wall is yellowish brown in color, composed
of a fibrous outer layer and a thin, little discernable, dark, micro-
granular layer. The fibrous layer tends to thicken toward the center
of the test.
Measurements. — (Based on two specimens). Number of volu-
tions: at least three. Diameter 140.00-187.50x. Width: 58.75-75.00x.
Ratio W/D: 0.40-0.42. Proloculus: Not determinable. Height of
lumen last volution: 11.25-13.75,. Peripheral wall thickness: 8.75-
i 25 pe
186 BULLETIN 298
Cotling. — Aligned.
Coting stage. — Angulatus.
Stratigraphic range. —USSR — Carboniferous. Australia —
Carboniferous.
Remarks. — Our two specimens appear to be identical to Mik-
lukho-Maklay’s 1953 species. Miklukho-Maklay considered her form
to be the same as that described by Howchin (1895) as Cornuspira
schlumbergt. Schubert (1908) erected the genus Hemigordius using
Howchin’s species C. schlumbergi as the type species. Cornuspira and
Hemigordius both belong to the family Fischerinidae and do not
possess radial walls. Miklukho-Maklay’s original description does
not mention radial walls and her illustration, though somewhat sug-
gestive of such walls, is not drawn with sufficient clarity. However,
she placed the genus Hemigordius in the family Archaediscidae which
she considered to have “indistinctly porous to coarsely porous walls.”
In this connection it is interesting to note that Hemigordius ulmert
Mikhailov, 1939 was selected by Miklukho-Maklay as the type
species for Propermodiscus, another archaediscid genus. She has,
however, described the latter genus as possessing radial walls.
The question arises as to whether Howchin’s C. schlumbergt is
an archaediscid. Without access to the type specimen this cannot
be definitely determined. From the original free-hand drawing of the
type specimen it is not possible to tell. In any event, Miklukho-
Maklay’s H. schlumbergeri cannot be the same as that described by
Howchin because that form is approximately three times the size
of Miklukho-Maklay’s specimens, has a smaller ratio of width to
diameter and has five volutions.
Both the size of our forms and the ratio of width to diameter
compare well with Miklukho-Maklay’s species.
While the authors believe a new species should not normally
be erected without a minimum of three specimens, in this case of
misidentification we are making an exception, naming the species
after the original author and using Miklukho-Maklay’s figured speci-
men as the holotype.
Archaediscus (Archaediscus) ex gr. moelleri Rauzer-Chernoussova, 1948
Pl. 22, fig. 14
Diagnosis. — Test small, disc-shaped with compressed nearly
parallel lateral sides, well-rounded periphery and relatively smooth
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 187
surface; coiling sigmoidal with only the final coil evolute; axial sec-
tion shows perfect sigmoidal coiling with the first one and a half
coils wound in a plane at 45 degrees to the axial plane and the final
two to three coils turned sharply to a 45 degree angle with the axial
plane in the opposite direction, making a 90 degree angle with the
plane of the initial coils; lumina dominantly flat-floored and semi-
lunular in shape, increasing progressively and markedly in size and
breadth in relation to height; wall has a well-developed clear radial
layer and poorly developed inner dark layer.
Measurements. — (Based on two specimens). Number of volu-
tions: four. Diameter: 146.87-160.00. Width: 18.75-23.00n. Ratio:
W/D .45-.478. Proloculus: 18.75-23.00. Height of lumen last volu-
tion: 15.62-16.00u. Peripheral wall thickness: 6.25-6.30p.
Coiling. — Sigmoidal.
Coiling stage. — Angulatus.
Stratigraphic range. — North America — Visean (V3-late V3c)
— Namurian. USSR — Visean (V3). Belgium — Visean (V3).
Germany — late Visean. Iran — Visean (V3b).
Remarks.— This species bears resemblance to Archaedsscus
pauxillus Shlykova, 1951 in sigmoidal coiling. However, in that
species the initial volution appears to be at 90 degrees to the suc-
ceeding one. The angle of deviation between these two volutions in
our two specimens varies from about 10° to 40°. The present form
is closer in size range to Archaediscus “var.” nana Rauzer-Chernous-
sova but differs in having sigmoidal coiling while A. nana is sigmoi-
dal only in the initial stage.
Our form has advanced to the typical angulatus stage in evo-
lutionary development. It is unfortunate that one of our two forms
was thinned so much it was partially destroyed and we have no
more criteria on which to base a specific diagnosis.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova, Mamet, 1973
Pl. 22, figs. 15-20
Archaediscus krestovnikovi var. pusillus Rauzer-Chernoussova, 1948a, p. 232,
pl. 16, figs. 4, 5; Grozdilova and Lebedeva (iz Dain and Grozdilova),
1953hep. 96, plo 3, figs: 35 4:
Archaediscus pusillus Rauzer-Chernoussova, Mamet, 1973, pl. 4, fig. 24.
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, No. 19.
Original description. — (Translated from the Russian in Ellis
and Messina, supplement No. 1, 1958.)
188 BuLLETIN 298
Diagnosis. — Test small with pronounced sutural depression outlining the
final whorl, parallel to moderately convex in outline in axial section with a
final evolute whorl appearing somewhat detached from the plane of symmetry;
initial three coils are involute and streptospirally wound and the final coils
evolute and slightly oscillating; completely open lumina increasing in size
have slightly convex to flat floors with epaulets extending on to the walls;
wall composed of two layers — a fibrous outer layer and a thin, dark micro-
granular layer.
Measurements. — (Based on 21 specimens). Number of volu-
tions: 3-1/2-5. Diameter: (based on 20 specimens): 158-225y. Ratio
W/D: 0.289-0.411. Width (based on 19 specimens): 58.00-81.25y.
Proloculus (based on 12 specimens): 16.25-25.50. Height of lumen
last volution: 16.66-27.60. Peripheral wall thickness (based on 19
specimens): 6.25-12.70n.
Coiling. — Oscillating.
Coiling stage. — Angulatus.
Stratigraphic range. — USSR — Visean (V3-Tula). France —
Visean (V3).
Remarks. — This species is one of the most abundant forms re-
covered from the fauna. The overall dimensions correspond remark-
ably well with those of the original description except for the fact
the proloculus attains larger dimensions. All of the specimens have
reached the angulatus stage but there seems to be some variation
in this feature.
We have adopted Mamet’s assignment of this form since we con-
sider it to be a definite species with chernoussovensis coiling. The
specimens do not have the characteristic type of coiling of the
species Archaediscus krestovmkovt Rauzer-Chernoussova.
Subgenus HEMIARCHAEDISCUS Miklukho-Maklay, 1957
Type species: Hemiarchaediscus planus Muiklukho-Maklay,
1957.
Original description, — Shell flat, lens shaped with slightly circular edges.
Test consists of a proloculus and a second pseudotubular chamber, glomerately
coiled at the beginning. The final coils are relatively freely coiled in a flat
spiral plane. The wall is bright, calcareous, distinctly porous with a clear dark
interior layer.
Among the representatives of this genus it is possible to see several new
species.
The stratigraphic range is from the beginning of the Visean to the end
of the Namurian. The geographic distribution covers Central Asia, Urals,
Kazakhstan and the European part of the USSR.
Remarks.— Specimens of this genus have been assigned by
other authors to several different genera — Planoarchaediscus
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 189
Miklukho-Maklay, Planospirodiscus Sosipatrova, Propermodiscus
Miklukho-Maklay, and Archaediscus Brady.
Hemiarchaediscus differs from Planoarchaediscus in being more
symmetrical and in possessing a clear, bright, radial wall. From
Planospirodiscus, it is differentiated by streptospiral coiling of the
initial coils and the high open, evolute final coils.
Miklukho-Maklay’s illustration of Hemigordius ulmert Mikhai-
lov, 1939, the type species of Propermodtscus, appears to belong to
the genus Archaediscus Brady, 1873 as emended by Conil and Pirlet
(1974, p. 254).
Most authors have equated Hemtarchaediscus with Archaedtscus
Brady, believing the tendency to lateral side thickenings is not a
criteria for generic designation. We believe this assignment is in-
correct.
We suspect that the wall of Hemiarchaediscus is, in fact, single-
layered and that other genera have been mistakenly assigned to this
genus. The possibility exists that petrographic relief along the inner
edge of the wall was mistaken for an inner layer. If such is the case
Hemiarchaediscus would represent a subgenus at a morphological
stage of evolution beyond (Archaediscus) equivalent to that attained
in Ammarchaediscus (Tubispirodiscus). If the inner layer is real,
Hemiarchaediscus is assignable to Archaediscus (Archaediscus) and
represents an advanced evolutionary stage. A definitive deposition of
this question cannot be achieved without access to the original type
specimens.
Subgenus 7HEMIARCHAEDISCUS Miklukho-Maklay, 1957
Type species: ?Hemiarchaediscus planus Miklukho-Maklay,
1957.
Original description. — Hemiarchaediscus planus Miklukho-
Maklay.
Holotype. — No. 16-23 Visean, Pamir.
Diagnosis. — The test is irregular, disc-shape. The first three to three and
a half coils are streptospirally wound with the final three to three and a half
coiled approximately in one plane.
Dimensions. — Diameter 0.32-0.46 mm (the holotype is 0.35 mm). Width
0.09-0.14 mm (the holotype is 0.12 mm).
Remarks.— The representatives of this genus have been referred to the
genus Propermodiscus Miklukho-Maklay 1953. Although they are similar to
Propermodiscus in the mode of coiling of the tubular chamber they cannot be
referred to that genus since they lack side thickenings. The manner of coiling
of the specimens appears more closely related to the genus Planoarchaediscus.
190 BULLETIN 298
Both have the same type of test structure. They are, however, distinct in wall
structure. The walls of Planoarchaediscus are thin and brownish with poorly
developed porosity. In Hemiarchaediscus they are bright and clear with well
developed porosity.
Diagnosis. — Archaediscus, discoidal to lenticular in shape,
initial coils involute with the final coils evolute and freely wound in
a flat, spiral plane with side thickenings confined to the initial
tangle; lumina open with flat floored bases in the final coils; wall is
a clear bright porous to poorly porous layer.
Remarks. — We are tentatively assigning to (?Hemiarchae-
discus) forms having a single radial layer but otherwise resembling
Miklukho-Maklay’s genus. If Hemiarchaediscus is indeed assignable
to Archaediscus (Archaediscus) our specimens and those placed in
synonymy with (?Hemiarchaediscus) represent a separate genus
that should be defined and named.
We have purposely given the original description of Miklukho-
Maklay’s type species H. planus because there is no mention made
of an inner dark layer although her description of the genus Hems-
archaediscus describes its presence.
Archaediscus (?Hemiarchaediscus) swanni Browne, n.sp. PI. 23, figs. 1-5
Holotype. —USNM (Nat. Mus. Nat. Hist.), No. 244590.
Diagnosis. —Test small with well-rounded periphery and mod-
erately convex sides marked by thickenings which increase toward
the center of the test; oscillation of coiling departs only slightly from
the plane of symmetry; inner coils involute with the final coil evo-
lute; open lumina increase markedly in size and breadth with each
coil changing in shape from nearly spheric to semi-lunular in out-
line as viewed in thin section; floors of the lumina dominantly
flat to moderately convex in shape with extensions on to the wall;
initial chamber small and spheric in form; wall is moderately thick,
clear, and fibrous without a dark inner layer.
Measurements. — (Based on 11 specimens). Number of volu-
tions (based on four specimens): 4-67. Diameter: 212.50-295.00u.
Width: 75-93n. Ratio W/D: 0.30-0.44. Proloculus (based on two
specimens): 13.75-15.00n. Height of lumen last volution: 16.00-
27.901. Peripheral wall thickness (based on ten specimens): 7.00-
11.25z.
Coiling. — Aligned.
Coiling stage. — Angulatus.
ARCHAEDISCIDAE: BROWNE, BAxTER, & ROBERTS 191
Stratigraphic range. — Unknown, Visean (late V3c), this report.
Remarks. — A. (?Hemiarchaediscus) swanm appears to be an
early form of the subgenus. The stage of evolution is somewhat
beyond A. (Archaediscus) but less advanced than A. (? Hemiarchae-
discus) stilus. It differs from the latter species in having fewer evo-
lute final coils and possessing marked side thickenings of the wall
that are indicative of its nearness to A. (Archaediscus). It also is
somewhat larger, has a lesser range of width to diameter and a
greater range in size of the proloculus. The mode of coiling is inter-
mediate to the gently oscillatory group stidus and true planispiral
forms assigned to Ammarchaediscus.
This species is named in honor of the late Dr. David H. Swann
in recognition of his contribution to Chesterian stratigraphy.
Archaediscus (?Hemiarchaediscus) cornuspiroides (Brazhnikova and Vdo-
venko, 1967) PE 23, 112.16
Archaediscus? cornuspiroides Brazhnikova and Vdovenko (in Brazhnikova,
et al.), 1967, pp. 162, 163, pl. 54, figs. 14-19; pl. 55, fig. 1.
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, No. 181.
Original description. —The test is small, strongly compressed along the
lateral margins with flat umbilici and is nearly disc shaped. The early coils
are involute and the last three to three and a half, sometimes four completely
evolute. The surface of all the coils is smooth. The ratio of the width to the
diameter ranges from 0.19-0.35. The range in dimensions of the diameter is
considerable, from 0.11-0.37, commonly from 0.15-0.26 mm. The width ranges
from 0.03-0.08 mm, commonly 0.05-0.07 mm. The number of coils is 5-7%. The
proloculus is large and spherical in shape with a diameter of 0.026-0.040 mm.
In the first two to three coils the coiling is in differing planes. The last three
to four coils are wound in a flat spiral. In saggital sections the flat spiral coil-
ing is quite apparent and shows a resemblance to the cornuspiroid genera. The
height of the tubular chamber shows a gradual increase which is sometimes
quite discernable in the last one or two coils. A height of 0.03-0.07 mm is at-
tained in the final coil. The wall, with clearly distinct outline, is thin, calcareous
and glassy (no porosity being visible). The maximum wall thickness of the
final coil is 0.01 mm. Variability is expressed by the considerable range in di-
mensions and in the displacement of the planes of coiling in the initial coils.
Comparison. — The wall structure and the manner of coiling of the final
coils distinguish these specimens from all known Archaediscidae. Because of
this they are tentatively referred to the genus Archaediscus.
Archaediscus(?) cornuspiroides, due to the characteristic coiling of the
final coils and its glassy, nonporous wall resembles the cornuspirids. It is
pertinent to note also that the structure of these forms is close to that of
Eosigmolina.
Diagnosis. — Test small, discoidal with plano-parallel sides, sur-
face marked by a lightly impressed suture of the final whorl; coiling
aligned; initial coils streptospirally wound and involute with the final
192 BULLETIN 298
two coils evolute, showing a maximum deviation of approximately
10 degrees from the axial plane; lumina with slightly convex floors
and semi-lunular outline increase progressively in size with a
marked increase from the involute to the evolute whorls; wall clear,
bright, and of approximately uniform thickness throughout. The
porosity is poorly developed and visible only at high magnifications.
Measurements. —(Based on one specimen). Number of volu-
tions: 5-6? Diameter: 187.50u. Width: 47.50u. Ratio W/D: 0.25.
Proloculus: 12.504. Height of lumen last volution: 18.75y. Peripheral
wall thickness: 6.25.
Coiling. — Aligned.
Coiling stage. — Angulatus approaching tenuis.
Stratigraphic range. — North America — Visean (V3c), this
report. USSR — late Visean — early Namurian of Dnieper —
Donetz Basin.
Remarks. — The authors believe this species should be tenta-
tively classified under the subgenus (?Hemiarchaediscus). The
original description of this form noted the “non-porous” wall as one
of the features by which it resembles the cornuspirids. However,
personal communication from Vdovenko confirms the presence of a
porous wall, noted only at magnifications of 180x and above. This
porosity is clearly visible in photographs sent by her. The tendency
toward a glassy wall of uniform thickness is an apparent evolutionary
development.
Archaediscus (?Hemiarchaediscus) stilus (Grozdilova and Lebedeva, 1953)
Pl. 23, figs. 7-14
Archaediscus stilus Grozdilova and Lebedeva (in Dain and Grozdilova), 1953,
(part) pp. 113, 114, pl. 4, fig. 20; Grozdilova and Lebedeva, 1954, pp.
61, 62, pl. 7, fig. 19; Vachard 1975, pp. 56, 57, pl. 8, figs. 2, 5.
Not Archaediscus stilus Grozdilova and Lebedeva, Bozorgnia 1973, pp. 112, 113,
pl. 17, fig. 6; pl. 19, figs. 11-13, pl. 22, fig. 17; Malpica, 1973, pl. 2, fig. 26.
Planoarchaediscus stilus (Grozdilova and Lebedeva), Sosipatrova, 1962, p. 58,
pl. 5, figs. 5, 6.
Planoarchaediscus stilus (Grozdilova and Lebedeva) forma compressa Bogush
and Yuferev, 1966, p. 160, pl. 11, fig. 7. :
Not Planoarchaediscus stilus (Grozdilova and Lebedeva) forma ftyfica Bogush
and Yuferev, 1966, p. 160, pl. 11, fig. 6.
Planoarchaediscus? stilus (Grozdilova and Lebedeva) forma magna, Bogush
and Yuferev, 1966, p. 160, pl. 11, fig. 8.
Archaediscus cf. ex gr. stilus (Grozdilova and Lebedeva) subspecies angulatus
Conil and Pirlet im Austin, Conil, Groessens, and Pirlet, 1974, pl. 3, figs.
9-12.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
No. 3191.
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 193
Original description. — The test is disc-shaped, elongated in axial section,
involute in the beginning whorls and evolute in the last 2-3. Periphery is round
and lateral sides are almost parallel. The surface of the test is smooth or slightly
dented. The ratio of the width to the diameter ranges from 0.32 : 1 to 0.45 : 1.
Dimensions (in mm).— Diameter 0.17-0.31, most commonly 0.23-0.31, the
width of the test 0.61-0.16 and most commonly 0.095-0.13. The proloculus is
spherical with a diameter ranging from 0.010-0.038 mm. The number of coils
is 5-6. The coiling of the tubular chamber is comparatively free, with gradual
increase in height with growth. The beginning coils are involute, wound in
differing planes (< 15-20 degrees to 40 degrees). The exterior 2-3 coils are
evolute and spirally flattened. ‘The height of the lumen of the final coil ranges
from 0.05-0.038 mm. The wall is glassy, radiant, finely porous, thin and not
pronounced in the early coils but tends to be somewhat thicker in the last
coils, changing from 0.007-0.019 mm. Due to the test shape and the type of
coiling this species bears a relationship to the group Archaediscus spirillinoides
Rauzer. Differences are observed in the wall structure which in the group
Archaediscus stilus here described is two layered, consisting of a well-developed
glassy, radiant layer and a less clear, inner dark layer. It is, likewise, distin-
guished by the large ratio of test inflation.
Diagnosis. — Test small to medium in size, ranging in shape
from flat, nearly plano-parallel, to moderately convex in outline, with
narrowly rounded periphery and slightly irregular to normally
smooth surface; interior coils streptospirally wound with the final
two to three coils evolute; layered thickening of the wall is apparent
on the sides of the test but is confined to the region of the area of
the involute coils only; lumina increase in size and shape at a marked
pace especially with the change from involute to evolute coiling, be-
ginning lumina semi-circular in shape with the later ones becoming
broader in relation to the height; wall is a single clear, porous, radial
layer which envelopes the test.
Measurements. — (Based on 23 specimens). Number of volu-
tions: (14 specimens) 5-6. Diameter: 153-290». Width: (21 speci-
mens) 44-954. Ratio W/D (based on 21 specimens): .30-.44. Pro-
loculus (based on eight specimens): 7.00-31.254. Height of lumen
last volution: 11.60-26.25y. Peripheral wall thickness: 5.00-11.60p.
Coiling. — Aligned.
Coiling stage. — Specimens show evolutionary stages from angu-
latus approaching tenuis.
Stratigraphic range. — North America — Visean (late V3c),
this report. USSR — Visean (V3) — Baschkirian. Morocco —
Visean (late V3c).
Remarks. — This species is one of the more abundant in the
fauna. It is characterized by its elongated shape in axial section
with its involute early whorls which possess lateral side thickenings
194 BULLETIN 298
and its free, evolute, dominantly planispiral final whorls without side
thickenings.
We elected to list only the holotype of A. stilus Grozdilova and
Lebedeva (im Dain and Grozdilova), 1953, plate 4, figure 20 in the
above synonymy because their other illustration on the same plate
(fig. 19) shows a pronounced dark inner layer. Grozdilova and Lebe-
deva (1954, pl. 7, fig. 19) refigured this holotype. The latter illustra-
tion shows what appears as a two-layered wall. The problem cannot
be resolved without access to the type specimens.
Our specimens show a series of progressive stages of evolutionary
development.
Archaediscus sp. [n. subgenus] Pll 25, Be 16
Figured specimen. — USNM (Nat. Mus. Nat. Hist.)
Diagnosis. — Test small, umbilicate in shape with the largest
diameter at extremities and the smallest through the axis of coiling,
surface uneven; coiling is aligned with the inner coils involute and
the outer two evolute; proloculus spherical, of moderate size; lumina
of nearly circular to semi-circular outline increase markedly in size
from center outward and have convex floors; a finely fibrous wall
of approximately equal thickness envelops all the whorls.
Measurements. — (Based on one specimen). Number of volu-
tions: 4?. Diameter: 178.75. Width: 47.50u. Ratio W/D: 0.266.
Proloculus: 15. Height of lumen last volution: 18.75. Peripheral
wall thickness: 10u.
Coiling. — Aligned.
Coiling stage. — Tenuis.
Stratigraphic range. — Unknown — Visean (late V3c), this re-
port.
Remarks. — Because the authors have only one specimen it is
not possible to adequately diagnose and define the limits required
for naming this new subgenus. We believe this subgenus represents
a final and rare stage of development of the genus Archaediscus. It
differs from the forms we have assigned to the subgenus ?Hemi-
archaediscus in the following respects:
1. test shape umbilicate with the smallest diameter through the
axis of the test
2. floors convex without shoulders or epaulets
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 195
3. lumina subcircular in outline throughout the test
4, radial wall of equal thickness well developed at low magni-
fications (single specimen)
This subgenus resembles Ammarchaediscus (subgenus A) sp.
Conil, 1974 (in Austin, Conil, Groessens, and Pirlet, 1974, pp. 116,
pl. 3, fig. 5) except that form is planispiral, completely evolute, and
still possesses evidence of an inner layer.
The form figured by Conil (in Austin, Conil, Groessens, and
Pirlet, 1974, pl. 3, figs. 1,3) as Archaediscus ex gr. stilus may belong
to our new subgenus. It will be noted we have assigned A. stilus to
the subgenus ?Hemtarchaediscus.
Genus NODOSARCHAEDISCUS Conil and Pirlet, 1974
Type species: Archaediscus maximum Grozdilova and Lebedeva,
1954,
Diagnosis. — Nodosarchaediscus characterized by the presence of nodes
angular nodosities in the ]umina or by a stellate central part formed by the first
whorls or by the combination of both. The internal dark layer is feeble to
imperceptible. The external radial layer comprises the large part of the test.
The coiling is normally evolute at least in the final coils. Floors are frequently
in the form of the letter W (Pirlet and Conil, 1974, p. 264).
Subgenus NODASPERODISCUS Conil and Pirlet, 1974
Type species: Archaediscus saleet Conil and Lys, 1964.
Diagnosis. — Nodosarchaediscus characterized by the presence of nodes
in addition to a stellate central part in which the coiling becomes confused due
to the occlusion of the lumina and the disappearance of the dark internal layer.
The dark internal layer which is feeble to imperceptible in the first coils is
completely covered throughout the test by the radial layer (Pirlet and Conil,
1974, p. 264).
Nodosarchaediscus (Nodasperodiscus) gregorii (Dain), 1953
Pl. 23, figs. 15, 16
Archaediscus gregorii var. gregorii Dain (in Dain and Grozdilova), 1953, p.
108, pl. 4, figs. 12, 13; Grozdilova and Lebedeva, 1954, p. 59, 60, pl. 7,
figss 12.13:
Planospirodiscus gregorii (Dain), Mamet, 1970, fig. 3 (chart) pl. 7, figs. 9, 10,
13, 14; Mamet, 1973, pl. 4, fig. 34.
Neoarchaediscus gregorii var. gregorit (Dain), Bozorgnia, 1973, pp. 135, 136,
pl. 30, figs. 7-9.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
No. 2640.
Original description. — The shell is disc-shaped with rounded periphery,
strongly compressed, producing parallel lateral sides.
Relation of the width to the diameter 0.30:1-0.40:1, Dimensions (mm).
Diameter of shell 0.28-0.38, the width of the shell 0.094-0.12, number of coils 5-6.
196 BULLETIN 298
The proloculus is spheric. It has a diameter of 0.019 to 0.029 mm. The
coiling of the tubular chamber in its initial stage lies in varying planes with a
gradual increase in the height of the coils — the first two to three coils in-
volute and the later ones strictly planospiral. The last three to four spiral flat
coils are strictly evolute. The lumina of the coils are narrow. They appear to
have an irregular outline, making a contour on the wall. The height of the
lumen of the last coil is 0.015 mm. The wall is thick, porous. Dain noted the
angularity of the walls of this species in his samples from the Donetz Basin.
It is expressed by the presence of three projections on the exterior side of the
peripheral part of the coil. The thickness of the wall of the last coil is 0.015
to 0.03 mm.
During the past four years this species has been discovered to be of wide
geographic distribution. The Ural samples which possess the same characteristics
and common dimensions differ from the Donetz-basin samples by free coiling
of the tubular chamber and a rather thin, porous wail. Specimens of Archae-
discus gregorii possess slight variations in dimensions, in the manner of coiling
of the tubular chamber (tight or more loosely wound) in test form and wall
thickness.
Diagnosis. — Test disc-shaped with plano-parallel sides and
rounded periphery; final two to three coils evolute and planispiral;
lumina increase moderately in size and are crescentic in shape due
to the fact they are dominantly filled with nodes which reduce the
fissural openings throughout; extensions of the floors of the lumina
extend across the walls of the outermost, evolute whorls; wall of
moderate thickness is composed of a fibrous layer and a very thin
dark microgranular layer.
Measurements. — (Based on two specimens). Number of volu-
tions: 4-5. Diameter: 220-280n. Width: 80-88. Ratio W/D: 0.31-
0.36. Proloculus (based on two specimens): 18.32-33.00n. Height of
lumen last volution: 19.46-20.60,. Peripheral wall thickness: 11.45-
12.60.
Coiling. — Aligned.
Coiling stage. — Angulatus.
Stratigraphic range.— North America — Visean (V3c)
Namurian. USSR — Visean (V3c) — Baschkirian. France — Visean
(V3b8). Iran — Visean (V3c) — early Namurian.
Remarks. — Our forms differ from Dain’s in being of slightly
smaller dimensions but having a similar width to diameter ratio.
They match more closely the dimensions given by Bozorgnia (1973).
This species is difficult to distinguish from (Nodasperodtscus) mint-
mus (Grozdilova and Lebedeva 1953) from which it differs by hav-
ing a larger ratio of width to diameter, a relatively larger proloculus,
lower lumina and a somewhat thicker wall. It differs from (Astero-
archaediscus) rugosus Rauzer-Chernoussova, 1948 in that the latter
ARCHAEDISCIDAE: BRowNE, BAXTER, & ROBERTS 197
by generic definition, except for a final coil, has occluded lumina, is
of wider dimensions and possesses a rugose outline. Assignment of
this species to Planospirodiscus Sosipatrova by Mamet (1970) is
inappropriate. It resembles that genus but Planospirodiscus taimiri-
cus, the type species, is planispiral and without stellate center-coil-
ing; we believe Planospirodiscus is a subgenus at an evolutionary
stage more advanced than ?Permodiscus Chernysheva of Pirlet and
Conil (1974, p. 280).
Nodosarchaediscus (Nodasperodiscus) minimus (Grozdilova and Lebedeva),
1953 Pl. 23, figs. 17-19
Archaediscus minimus Grozdilova and Lebedeva (in Dain and Grozdilova),
1953, pp. 111, 112, pl. 4, fig. 15.
Not Avchaediseis? minimus Grozdilova and Lebedeva, 1954, pp. 62, 63, pl. 7
fig. 16.
Asteroarchaediscus gregoriu (Dain) Brazhnikova, et al., 1967, pl. 21, fig. 4
Planospirodiscus minimus (Grozdilova and Lebedeva) Sosipatrova, 1962, pp.
64, 65, pl. 5, figs. 22-24; Mamet, 1970, pl. 7, figs. 15-18.
Neoarchaediscus incertus (Grozdilova and Lebedeva) Bozorgnia, 1973, pp. 130,
131, pl. 20, figs. 138-139.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
No. 3190.
Original description. — The shell is small, disc-shaped with widely turned
peripheral region and parallel lateral sides. The surface is smooth. The ratio
of width to diameter varies from 0.30:1 to 0.50:1 — the most frequently en-
countered ratio being 0.30:1 to 0.40:1. Dimensions (mm): diameter varies from
0.19 to 0.27; the width from 0.076 to 0.095. The number of coils two to five;
commonly three. The proloculus is spheric. In consideration of the small dimen-
sions of the species it is comparatively large with a diameter varying from
0.019 to 0.023 mm. The initial one and a half to two coils decline slightly from
the plane of symmetry. The final two to three coils are evolute flat and plani-
spiral. The lumina are relatively large with the clearly expressed outline of
a small arch. The height of the lumen of the last coil is 0.019 to 0.023 mm.
The wall is glassy, radial, and finely porous. The thickness of the wall about
equals the height of the lumen.
Archaediscus minimus, n. sp. because of the form of the test and the char-
acteristics of the external "coils approaches the genus Permodiscus from which
it differs by the initial coiling of the tubular chamber and the absence of
lateral thickenings.
Distribution. — This species is found in deposits of Baschkirian age on the
western slope of the Ural Mountains.
Diagnosis. — Test small, smooth to slightly rugose, flat-sided
with narrowly rounded periphery; initial coils tightly wound pro-
ducing flaring with the final three coils evolute and planispiral;
lumina, except for the final whorl, are almost completely filled with
chevron-shaped nodes which have reduced the fissural openings
to narrow slits; wall is a single fibrous layer.
198 BuLLeETIN 298
Measurements. — (Based on six specimens). Number of volu-
tions: 4544. Diameter (based on five specimens): 181.00-248.75y.
Width: 52.50-67.50n. Ratio W/D: 0.249-0.32. Proloculus (based on
two specimens): 9.16-11.00u. Height of lumen last volution: 11.60-
25.00. Peripheral wall thickness: 5-10p.
Coiling. — Aligned.
Coiling stage. — Angulatus.
Stratigraphic range. — North America — Visean (V3c) — early
Namurian. USSR — Baschkirian. Iran — Visean (V3c) — early
Namurian.
Remarks.— Our specimens approximate those of the authors.
They possess a slightly greater width than the given width although
the author’s illustrated figure 15 on plate 4 has the same ratio of
width to diameter as our forms the proportions of which match those
of Bozorgnia, 1973. The proloculus is slightly smaller than those of
the authors. This may be due to the limited number of our forms.
Grozdilova and Lebedeva’s illustration of the type is not clear
and not diagnostic enough to determine the stellate juvenarium.
However, the reference in their description to “the clearly expressed
outline of a small arch” in the lumen leads us to believe they were
describing nodes and not the crenulate outline of the occluded lumina
of the genus Asteroarchaediscus Miklukho-Maklay. Moreover, the
fact that our specimens match so closely the illustration given by the
other authors in the above synonymy and are from the same strati-
graphic level lends credence to our assignment.
Subgenus NEOARCHAEDISCUS Miklukho-Maklay, 1956
Type species: Archaediscus incertus Grozdilova and Lebedeva,
1954.
Original description.— The shells are flat — discus-shaped with more or
less parallel sides. The surface of the shell is smooth or somewhat uneven.
The beginning chamber is spheric. The second chamber, not divided, coiled at
the beginning (frequently with star-shaped structure), is followed by two or
three coils turned more or less in one plane and more freely. The wall is cal-
careous, bright, quite thick, glassy, finely porous, with an interior, thin dark
layer.
Diagnosis. — Nodosarchaediscus characterized by the confused and stellate
central coils. The terminal spires (at least one and a half to two) are free,
without nodes or traces of occlusion. The dark layer, thin or imperceptible,
is completely covered by the radial layer. (Pirlet and Conil, 1974, p. 268).
Remarks. — Pirlet and Conil (1974) erected the subgenus
Asperodiscus to include the genera Neoarchaediscus and Rugo-
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 199
sarchaediscus which they separated only at the specific level. The
authors agree that the two genera can be differentiated only on the
tendency of forms assigned to Rugosarchaediscus to be somewhat
more convex in shape and to have the terminal spires depart more
from the plane of symmetry. These characteristics are evolutionary
and intermediate forms exist.
We consider the genus Rugosarchaediscus Miklukho-Maklay,
1957, and the subgenus Asperodiscus Pirlet and Conil (1974) to
be junior synonyms of (Neoarchaediseus).
Noedosarchaediscus (Neoarchaediscus) cf. bykovensis Sosipatrova, 1966
PIS23 5 fis. 205) Pi 24 figs 2
Neoarchaediscus bykovensis Sosipatrova, 1966, pp. 19, 20, pl. 3, figs. 1, 2.
Holotype. — Institute of Geology of Arctic Regions, 659/3 By-
kov Canal, Tixin suite, collection of R. V. Solomina, 1959, specimen
396.
Original description.— The shell is lens-shaped, almost flat with parallel
lateral sides and rounded peripheral margin, involute in starting coils and
evolute in the last two. The ratio of width to diameter is 0.20-0.31. Diameter
0.24-0.32 mm. Width 0.060-0.075 mm. The number of coils 514-6.
The proloculus is small with a diameter of 0.019 mm. The tubular chamber
is tightly wound with small declination of the axis in the initial coils. The last
three coils are more open and planispiral. The height of the lumen in the final
coil is 0.22-0.029 mm, giving a height three times the wall thickness. The lumina
are semi-lunular in shape with slightly swollen bases.
The wall consists of two layers with the outer, glassy, radial layer pro-
nounced. In the initial coils, the stellate structure is well-displayed. The wall
thickness is 0.006-0.009 mm. The characteristic features of this species are:
1. The nearly spiral flat coiling.
2. Stellate structure of the wall in the initial coils.
3. The parallel or slightly swoilen sides of the test.
Comparison.— Our species bears the greatest resemblance to Neoarchae-
discus subplanus (Brazhnikova), known in the suite C4, from which they may
be separated by the thinner wall and the more convex sides of the test. In
manner of coiling of the tubular chamber N. bykovensis bears closer affinity to
Planospirodiscus minimus (Grozdilova and Lebedeva) from which it differs
in the stellate structure of the initial coils, the much greater height of the
lumina and the smaller proportion of width to diameter. The present species,
with its parallel sides and the spirally flattened coiling of its outer coils is
closer to Neoarchaediscus borealis Reitlinger from which it differs by having
a rather narrow test, high open lumina in the later coils, and a thick wall.
Distribution: Northern Kharaulakh. Baschkirian layer. The location is in the
Bykov canal, the upper part of the Tinosh suite. Collection of R. V. Solomena,
1959, Sample 396.
Diagnosis. — Test small, discoidal, with plano-parallel sides,
broadly rounded periphery and slightly uneven surface, initial coils
involute, tightly wound and displaying stellate structure; final two to
three evolute, flat and planispiral except for the final coil which may
200 BULLETIN 298
turn as much as 30° from the plane of symmetry; lumina of the in-
terior coils, often hard to detect due to the tight coiling, are small
and somewhat semicircular in shape; lumina of the outer evolute coils
change abruptly from tight coiling to free open planispiral coiling
and increase progressively in height and breadth, becoming semi-
Junular to somewhat quadrate in shape; wall bilayered with an outer,
clear radial layer and a thin, dark microgranular layer.
Measurements. — (Based on six specimens). Number of volu-
tions (based on three specimens): 5-7?. Diameter: 195-251y. Width:
57.50-70n. Ratio W/D: 0.22-.34. Proloculus (based on two speci-
mens): 14.00-17.50u. Height of lumen — Last volution: 18.75-30.00z.
Wall thickness: 5.00-7.50n.
Coiling. — Aligned.
Coiling stage. — Angulatus.
Stratigraphic range. — North America — Visean (late V3c),
this report. USSR — Baschkirian.
Remarks. — Sosipatrova’s description made no reference to any
declination of the final coil from the plane of symmetry nor do her
illustrations show it. One of our specimens (PI. 23, fig. 20) has a
projection extending out from the final coil, but this projection ap-
pears to be matrix and not part of a broken coil which lends credence
to the fact that slight declination of the final coil is not a specific
character.
This species bears close resemblance to Neoarchaedtscus incertus
(Grozdilova and Lebedeva) from which it is separated by a lesser
ratio of width to diameter, more evolute planispiral coiling of the
outer whorls and an abrupt change from involute to evolute coiling.
In our forms the outer planispiral coiling comprises from approxi-
mately two to three times the length of the test versus 1 to 1.5 times
that of those forms in this fauna which we have assigned to the
species incertus. Moreover, (N.) cf. bykovensis has a thinner wall
with a final lumen height about three times the wall thickness.
Nodosarchaediscus (Neoarchaediscus) incertus (Grozdilova and Lebedeva)
1954 Pl. 24, figs. 3-6
Archaediscus incertus Grozdilova and Lebedeva, 1954, pp. 60, 61, pl. 7, figs. 14,
iby
Neoarchaediscus incertus (Grozdilova and Lebedeva), Miklukho-Maklay, 1956,
p. 11; Grozdilova and Lebedeva, 1960, p. 98, pl. 11, fig. 11; Brenckle, 1973,
p. 63 (part) pl. figs. 16-19, 20?, 21?, 22-25; Vachard, 1975, pp. 58-60, pl.
8, figs. 1-3. -_
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 201
Not Neoarchaediscus incertus var. incertus (Grozdilova and Lebedeva), Conil
and Lys, 1964, p. 130, pl. 20, figs. 389-391.
Neoarchaediscus postrugosus (Reitlinger), Hewitt and Conil, 1969, p. 178 (part)
Not Le aaa incertus Bozorgnia, 1973, p. 130, pl. 30, figs. 1-6.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
Leningrad. No. 3686.
Original description. — (Translated from the Russian in Ellis
and Messina, supplement No. 1, 1964).
Diagnosis. — Test small, disc-shaped, usually with rugose sur-
face producing lightly serrated edges; periphery broadly rounded and
sides plano-parallel to moderately convex; initial coils streptospirally
wound with final one and a half to two coils evolute, more open with
less departure from the plane of symmetry; open lumina of the outer
whorls increase moderately in size, are of semilunular outline and
have slightly convex floors extending to the edges of the wall; wall
composed of an outer fibrous layer and a poorly developed, frequently
absent, inner layer.
Measurements. — (Based on ten specimens). Number of volu-
tions: 5-6?. Diameter: 192.50-273.75y. Width: (based on nine speci-
mens): 52.50-91.25. Ratio W/D: 0.275-0.388. Proloculus (based
on two specimens): 7.50-10.00u. Height of lumen last volution:
12.50-25.00u. Wall thickness: 7.50-11.25p.
Coiling. — Aligned.
Coiling stage. — Angulatus.
Stratigraphic range. — North American — Visean (V3bB-8) —
lower Namurian. USSR — Visean (late V3c) — Baschkirian. Moroc-
co — Visean (V3c) of Akerchi.
Remarks. — This species has been confused in the literature
with a number of similar species. This is perhaps due to the lack of
clearly defined illustrations by the original authors. It differs from
N. (Nodasperodiscus) gregori (Grozdilova and Lebedeva) primarily
in that the species gregoru has the lumina filled by nodes.
Bozorgnia (1973) identified specimens which we would assign
to N. (Nodasperodiscus) minimus (Grozdilova and Lebedeva) as
Neoarchaediscus incertus. The dimensions of Bozorgnia’s specimens
match those of Neoarchaediscus incertus but the lumina are, as he
described them, filled with nodes.
Our forms conform well to the original description given by
202 BULLETIN 298
Grozdilova and Lebedeva although the specimens are more uniform
in size.
Nodosarchaediscus (Neoarchaediscus) latispiralis (Grozdilova and
Lebedeva), 1953 Pl. 24, figs. 7-9
Archaediscus latispiralis Grozdilova and Lebedeva (in Dain and Grozdilova),
1953, pp. 102-103, pl. 3, fig. 17.
Neoarchaediscus latispiralis (Bogush and Yuferev), 1962, p. 207, pl. 9, fig. 16;
Bogush and Yuferev, 1966, p. 160, pl. 11, fig. 23.
Archaediscus aff. latispiralis Grozdilova and Lebedeva, Conil and Lys, 1964,
p. 122, pl. 18, fig. 360.
Archaediscus? (Rugosarchaediscus) latispiralis Grozdilova and Lebedeva,
Bozorgnia, 1973, p. 120, pl. 27, figs. 1-6.
Holotype. — All Union Petrol. Sci. Res. Geol. Prospect. Inst.,
No. 2292.
Original description.— The test is disc-shaped with broadly rounded peri-
phery and nearly parallel] sides, involute with the exception of the last coil. The
ratio of the width to the diameter is rather constant within the limited range
of 0.40:1-0.50:1. .
The exterior surface is predominantly smooth with only slight unevenness.
Dimensions (in mm): diameter of the test 0.28-0.36, Width 0.12-0.20, Number of
coils: 4-7.
The proloculus is spheric with a diameter of 0.029 mm. The tubular cham-
ber is very tightly coiled initially in planes which are displaced in respect to
the axis of coiling. The height of the lumen of the final coil ranges from
0.035-0.058 mm.
The wall is bright, consisting of two layers with the bright, glassy radial
layer more pronounced. The wall thickness of the final coil ranges from
0.015-0.022 mm.
This species belongs to the group Archaediscus baschkiricus Krestovnikovi
and Theodorovich. The distinguishing features of the species are as follows:
1. The presence of two clearly distinct growth stages — the initial globular
shape and a later more freely coiled planispiral one.
2. The unique form of the test. This species resembles Archaediscus basch-
kiricus Krest. and Theod. in its early stages of growth. Mature specimens
differ in the shape of the test as well as in the manner of coiling of the
final whorls.
Diagnosis. — Test small, discoidal to lenticular with well-
rounded periphery, surface somewhat uneven; coiling of tubular
chamber streptospiral with the initial coils involute and tightly
wound with stellate outline formed by small pointed nodes, final
one to one and a half coils evolute, more freely wound and open,
with less departure from the plane of symmetry; wall two layers with
an outer bright well-developed finely fibrous layer and a poorly de-
veloped inner dark layer.
Measurements. — (Based on six specimens). Number of volu-
tions: not determinable. Diameter: 122.50-198.75. Width: 62.50-
86.25. Ratio W/D: 0.43-0.51. Proloculus (based on one specimen):
26.66u. Height of lumen last volution (based on five specimens):
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 203
10.00-18.75y. Peripheral wall thickness (based on five specimens) :
8.75-11.25.
Coiling. — Oscillating.
Coiling stage. — Angulatus.
Stratigraphic range. — North America — Visean (late V3c),
this report. USSR — Visean (middle) dominantly Baschkirian.
Belgium — Visean (V3b8). Iran — Visean (V3b to V3c).
Remarks. — Our specimens resemble the type specimen in shape
and manner of coiling. The inner coils are so tightly wound that it is
difficult to accurately determine the number of volutions. We esti-
mate a range with a minimum of three and a maximum of five.
Specimens with the number of whorls in the lower part of this range
happen to be better preserved in our material and in axial sections
show a marked likeness to the axial section illustrated by Bogush and
Yuferev (1966, pl. 11, fig. 23). Likewise, our specimens are closer
in size range and lumen height to those of Bogush and Yuferev.
Bozorgnia’s specimens (1973) are probably the same species or a
subspecies. The dimensions are considerably larger than the type
specimen, the nodes of the early whorls more pronounced and
the final whorls more nearly planispiral.
Nodosarchaediscus (Neoarchaediscus) pohli, Browne, n. sp.
Pl. 24, figs. 10-12
Archaediscus (?) minimus Grozdilova and Lebedeva, 1954, pp. 62, 63, pl. 7, fig.
16.
Not Archaediscus minimus Grozdilova and Lebedeva, (im Dain and Grozdi-
lova), 1953, pp. 111, 112; pl. 4, fig. 15.
Not Planospirodiscus minimus (Grozdilova and Lebedeva), Sosipatrova, 1962,
pp. 64, 65, pl. 5, figs. 22-24.
Planospirodiscus sulcus (Grozdilova and Lebedeva), Brenckle, 1973, p. 65, pl.
9, figs. 35-37.
Holotype. —USNM (Nat. Mus. Nat. Hist.), No. 244618.
Diagnosis. — Test small with planoparallel to slightly convex
sides and well-rounded periphery; initial coils involute, tightly wound
and angled, forming a stellate outline; final coils more freely wound,
as many as four of which may be evolute; crenulated edges of arch-
shaped walls fill the lumina, except for the final two coils, which are
open and free; lumen height of final coil nearly double the wall thick-
ness; walls two layered with an outer thick, fibrous layer and a thin
dark microgranular layer.
204 BULLETIN 298
Measurements. — (Based on six specimens). Number of volu-
tions: 5-5%?. Diameter: 202.30-260.00u. Width: (based on five
specimens) 65-88. Ratio W/D: 0.29-0.33. Proloculus (based on two
specimens): 16-184. Height of lumen last volution (based on five
specimens): 15.00-20.60u. Peripheral wall thickness (based on five
specimens) : 6.90-11.40z.
Coiling. — Aligned.
Cotling stage. — Angulatus.
Stratigraphic range. — North America — Visean (late V3b) —
Baschkirian. USSR — Baschkirian.
Remarks. — The present species has been confused in the litera-
ture with Archaediscus minimus Grozdilova and Lebedeva (in Dain
and Grozdilova, 1953) which we now consider to belong to the sub-
genus (Nodasperodiscus). This is probably due to the somewhat
arch-shaped outline of the lumina of the planispiral coils. Our speci-
mens are of similar size range and proportion to Archaediscus?
minimus Grozdilova and Lebedeva, 1954.
We have noted a resemblance of this species to Planospirodiscus
sulcus in Brenckle (1973, p. 65, pl. 9, figs. 35-37). We do not concur
in Brenckle’s generic assignment of this species. We would assign it
to the subgenus Neoarchaediscus. P. sulcus has the outer whorls
free and open. An examination of the types shows what appears to
be stellate central coiling.
The final whorl of Nod. (Neoarchaediscus) pohli has a tendency
to inflate as does P. sulcus, but the lumina of the latter forms are
higher and the ratio of width to diameter is greater.
This species and those previously assigned conditionally to A.
minimus of authors can be readily separated at the subgeneric level.
The final two coils of the present species are open and free which
distinguishes (Neoarchaediscus). Grozdilova and Lebedeva (in Dain
and Grozdilova, 1953) described A. minimus as having a wall thick-
ness “about as high” as the height of the lumen.
This species is named in honor of the late Dr. E. R. Pohl who
collected the material on which this report is based.
Nodosarchaediscus (Neoarchaediscus) sp. Pl. 24, fig. 16
Figured specimen. —USNM (Nat. Mus. Nat. Hist.), No.
244623.
Diagnosis. — Test small with nearly flat, planoparallel sides and
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 205
angularly rounded periphery; surface rough; proloculus, hard to
distinguish, is surrounded by a tightly coiled tubular chamber, strep-
tospirally wound and initially showing stellate structure with the
final two coils becoming planispiral; lumina increase slowly in size
and become broader and more semicircular toward the peripheral
ends; floors of lumina are nearly flat in final whorls and they extend
to edges of the walls; wall is thick, clear and fibrous.
Measurements. — (Based on one specimen). Number of volu-
tions: five?. Diameter: 210u. Width: 75y. Ratio W/D: 0.36. Pro-
loculus: indeterminable. Height of lumen last volution: 11.45y.
Peripheral wall thickness: 11p.
Cotling. — Aligned.
Coiling stage. — Angulatus.
Stratigraphic range. —Unknown — Visean (late V3c), this
report.
Remarks. — This species in rough outline and size bears resem-
blance to Asteroarchaediscus rugosus (Rauzer-Chernoussova) from
which it is differentiated at the subgeneric level. We believe our
specimen to be (Neoarchaediscus) because the final two to two and
half coils are open, lack nodes, and are flat-floored.
Nodosarchaediscus (Neoarchaediscus) timanicus Reitlinger, 1949
Pl. 24, figs. 13-15
Archaediscus timanicus Reitlinger, 1949, p. 163, pl. 1, figs. 7a, b, c; Grozdilova
and Lebedeva, (iz Dain and Grozdilova), 1953, p. 109, pl. 3, figs. 18-20.
Neoarchaediscus timanicus (Reitlinger) Sosipatrova, 1962, p. 62, pl. 5, fig. 11;
Brazhnikova, et al., 1967, pl. 25, fig. 3; Vdovenko, 1968, pl. 2, fig. 25;
Hewitt and Conil, 1969, pl. 2, figs. 28, 29.
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, No. 3278/12.
Original descriftion. — The test is not large with flattened or poorly convex
sides and dull, rounded periphery. The last half of the tubular coil is evolute.
The dimensions of the diameter range from 0.11-0.25 mm. The width ranges
from 0.062-0.11 mm. The ratio of the width to the diameter varies from 0.37-
0.55. The proloculus is spherical in shape with an interior diameter of 18-31
microns. The tubular chamber has 5-6 coils, the first two to three of which oscil-
late sharply from the axis of coiling, with the final coils wound in a nearly
flat spiral. The lumina of the chamber are semi-lunular. The height of the
lumina of the first coils is equal to the thickness of the wall while the height
of the lumina of the last two to three coils exceeds the thickness of the wall.
The height of the lumen of the last coil is 24-31 microns. Angular projections
are formed by the bending of the early coils with their conjunction with the
wall. This feature gradually disappears in the outer coils. The interior coils,
due to the angularity of the coiling, are characteristic of the group Archaediscus
206 BuLLETIN 298
baschkiricus of similar stellate contour. The wall is bright, poorly radial and
thickened laterally. The thickness of the wall of the last coil is 6.5-12, sometimes
18 microns.
Comparison.— This form has the features characteristic of the group
Archaediscus baschkiricus and Archaediscus krestovnikovi. It is similar to the
first by the angularity of the coils and to the second by the well-defined lumina
of the beginning coil to the smooth one-two last coils. Evolution shows a ten-
dency to the diminishing of dimensions, thinning of the walls and larger to
smaller thickening of the test. The specimens from the upper part of the upper
Kayal horizon and the bottom of the Verey have average dimensions for the
diameter of 0.11-0.17 mm and for the wall thickness 6-9 microns.
Diagnosis. — Test small with slightly convex sides and rounded
peripheral margins; initial two to three coils tightly and strepto-
spirally wound producing serrated outlines; final coils approximate
the plane of symmetry with one to one and a half coils evolute;
lumina with semilunular outline and slightly convex floors increase
moderately in size; wall is clear and fibrous.
Measurements. — (Based on eight specimens). Number of volu-
tions (based on three specimens): 4-6?. Diameter: 170.00-218.75y.
Width: 60.00-87.50n. Ratio W/D: 0.33-0.41. Proloculus: (based on
three specimens) 15.00-18.60u. Height of lumen final volution: 13.75-
17.00. Peripheral wall thickness: 7.00-13.75y.
Coiling. — Slightly oscillating.
Coiling stage. — Angulatus.
Stratigraphic range.— North America Visean (late V3c) —
Namurian (Pennington). USSR — Baschkirian with isolated exam-
ples in early Vesean.
Remarks. — Our specimens compare favorably with Reitlinger’s.
The height of the lumina measures from a third to double the wall
thickness. Grozdilova (in Dain and Grozdilova, 1953) gave a lumen
height of double to three times the wall thickness. In the sharply
serrated edges of the wall outline of the initial coils the species re-
sembles (Neoarchaediscus) incertus. It differs from N. incertus by its
smaller size, less symmetrical form, a larger ratio of width to
diameter, a lesser lumen height, and a generally smaller proloculus.
Subgenus ASTEROARCHAEDISCUS Miklukho-Maklay, 1956
Type species: Archaediscus baschkiricus Krestovnikov and
Theodorovitch, 1936.
Original description. —'The tests assume differing shapes, generally with
somewhat uneven surface. The coiling is streptospiral, streptospiral to flat
spiral or only flat spiral. Due to the sharp turns of the tubular chamber the
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 207
contour presents a stellate outline. The height of the lumina is several times
smaller than the wall thickness between corresponding coils. The wall is thinly
porous. (Miklukho-Maklay, 1956, p. 10).
Diagnosis. — Nodosarchaediscus characterized by the almost total occlusion
of the lumina and the barely discernable to complete disappearance of the in-
ternal dark layers. Unless the enrollment is aligned it is difficult to determine.
Only the last coil may show a free lumen. The radial layers completely cover
all the coils and are well developed. (Pirlet and Conil, 1974, p. 270).
Nodosarchaediscus (Asteroarchaediscus) parvus (Rauzer-Chernoussova),
1948 Pl. 24, figs. 17-20
Archaediscus parvus Rauzer-Chernoussova, 1948a, p. 233, pl. 16, figs. 9-12;
Grozdilova (in Dain and Grozdilova), 1953, pp. 104, 105, pl. 4, fig. 6;
Malakova, 1956, p. 41, pl. 3, figs. 4, 5.
Neoarchaediscus parvus (Rauzer-Chernoussova), Miklukho-Maklay, 1956, p.
11; Mamet 1973, pl. 4, figs. 20, 21.
Neoarchaediscus cf. N. parvus (Rauzer-Chernoussova), Brenckle, 1973, pl. 9,
figs. 11-15.
Asteroarchaediscus parvus (Rauzer-Chernoussova), Vdovenko, 1968, pl. 2, fig.
ile
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, fig. 10.
Original description. — (Translated from the Russian in Ellis
and Messina, supplement No. 1, 1958).
Diagnosis. — Test small, discoidal, surface uneven which im-
parts an irregular or serrated outline to axial sections; sides slightly
convex to nearly parallel; periphery sharp and broadly rounded; in-
terior coils involute, streptospirally and tightly wound producing flar-
ing; final coils, one to two of which may be evolute, deviate only
slightly from the plane of symmetry; lumina with low fissural open-
ings except for the final coil, filled by irregular to chevron-shaped
nodes, formed by the crenulated walls of the coils; wall clear,
finely fibrous and moderately thick.
Measurements. — (Based on 12 specimens). Number of volu-
tion (based on seven specimens): 4-5. Diameter: 140.00-247.50y.
Width: 55.00-92.50. Ratio W/D: 0.335-0.40. Proloculus (based on
five specimens): 7.00-11.60u. Height of lumen last volution: (Based
on 10 specimens) 0.00-20.00u. Peripheral wall thickness: 7.00-18.75.
Test form. — Flat to lenticular.
Stratigraphic range. — North America — late Visean — Namu-
rian. USSR — Visean (V3c) Oka — Venev — early Serpukhovian.
France — Visean (V3b-V3c).
Remarks.— Our specimens match closely in size those of
Rauzer-Chernoussova’s type specimens with only two exceeding the
given dimensions. The ratio of width to diameter is similar. The
208 BULLETIN 298
lumen height is equal or less than the wall thickness except for the
final whorl. The author stated in describing this species “the cham-
ber lumen usually equals the wall thickness.” Her specimen figure
11, like some of ours, shows a high open lumen in the final coil. This
feature appears to be common to the subgenus Asteroarchaediscus.
The authors have doubts that the A. parvus regularis Sulei-
manoy, 1948 warrants the designation of a subspecies. Due to lack
of a sufficient number of forms in our material we can not verify
this. However, we do not consider the height of the frequently open
lumen of the final whorl diagnostic. The smoother surface of the
final whorls seems likewise, not to be a character confined to the
subspecies. Some of our specimens have less rough walls than others
in the final whorls.
Superficially this species resembles (Asteroarchaediscus) rugo-
sus (Rauzer-Chernoussova), 1948 due to shape and roughness of the
wall. (Ast.) rugosus has larger dimensions and is less symmetrical
in outline.
Nodosarchaediscus (Asteroarchaediscus) postrugosus (Reitlinger), 1949
Pl. 25, figs. 1-3
Archaediscus postrugosus Reitlinger, 1949, p. 162, pl. 1, figs. 10a, c; Grozdilova
in Dain and Grozdilova, 1953, pl. 4, figs. 9, 10.
Asteroarchaediscus postrugosus (Reitlinger) Brazhnikova, et al., 1967, pl. 20,
fig. 4 and pl. 21, figs. 2, 3; Aizenverg, Brazhnikova and Potievskaia, 1968,
pl. 26, fig. 9; Bozorgnia 1973, p. 138, pl. 30, figs. 10-13, 19.
Neoarchaediscus postrugosus (Reitlinger), Hewitt and Conil, 1969, p. 178, pl.
2, figs. 16-23.
Holotype. — Repository not given — probably Museum Inst.
Geol. Sci. Acad. Sci., USSR, Moscow No. 327/21.
Original description. —The shell is not large and has flat sides with full
rounded periphery. The final coil in typical specimens is evolute and situated
symmetrically in the plane of symmetry of the test. The dimensions of the
diameter range from 0.17 to 0.34 mm, commonly 0.22-0.29 mm. The width ranges
from 0.074-0.14 mm, usually 0.10-0.11 mm. The ratio of the width to the
diameter is 0.30-0.54 and most frequently 0.35-0.42. The interior coils are
densely wound in ball-shaped form. The lumina of the final coil are high
and several times greater than the height of the wall thickness. The height in
the final coil is 18-29 microns. Those of the penultimate whorl are much smaller,
the height less than the wall thickness. The wall of the interior coils is thick
while that of the final two is thin. Wall thickness varies from 6-15 microns.
Comparison.— This form is very characteristic with its interior tightly
wound coils similar to Archaediscus rugosus. However, the evolute, final coil
with its characteristic symmetry, evidenced in all specimens is the reason for
considering this form a subsequent stage in the development of Archaediscus
rugosus, From Archaediscus parvus Rauzer it differs by its larger dimensions,
its less regular and poorly defined coiling of the interior coils and its rather
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 209
thick wall. A similar and evidently convergent form which, however, belongs
to another genetic line appears to be Archaediscus parvus regularis var.
Suleimanov. Characteristic of the latter form are the high lumina in the two
to three final coils (equal to or exceeding wall thickness), the smooth wall of
the final coils (with this species only the final coil is smooth, the interior coils
having the characteristic angular protrusion of Archaediscus rugosus along
the periphery) and the more regular disposition of the interior coils.
Diagnosis. — Test small, lenticular, characteristically convex in
outline, although occasional specimens may have a flattened con-
tour, surface normally showing roughness along the sides caused by
the angularity of the involute whorls; interior coils involute with the
final one to two becoming evolute, all typically tightly wound except
for the final coil which is distinguished by its free symmetrical coil-
ing: lumen of the final whorl open and moderately high, remaining
ones almost completely closed by prominent nodes which present
a stellate outline; wall, without an apparent inner layer, is clear and
fibrous.
Measurements. — (Based on seven specimens). Number of
volutions (based on three specimens): 5-6?. Diameter: 212.50-
235.00n. Width: 63.75-77.50u. Ratio W/D: 0.28-0.33. Proloculus:
indeterminate. Height of lumen last volution: 18.30-25.50y. Peri-
pheral wall thickness (based on six specimens): 6.25-8.75y.
Test form. — Flat to normally lenticular.
Stratigraphic range. — North America — late Visean — Namu-
rian. USSR — late Visean — early Namurian and Baschkirian.
Morocco — early Namurian.
Remarks. — (Ast.) postrugosus is larger than (Ast.) rugosus,
generally more symmetrical, has higher open lumen, and a lesser wall
thickness in the final evolute coil.
Nodosarchaediscus (Asteroarchaediscus) rugosus (Rauzer-Chernoussova),
1948 Pl. 25, figs. 4-8
Archaediscus rugosus Rauzer-Chernoussova, 1948b, p. 11, pl. 3, figs. 4-6;
Grozdilova (im Dain and Grozdilova), 1953, pp. 103, 104, pl. 4, figs. 1-3.
Neoarchaediscus rugosus (Rauzer-Chernoussova), Sosipatrova, 1962, pp. 61, 62,
pl. 5, figs. 12-14.
Asteroarchaediscus rugosus (Rauzer-Chernoussova), Bogush and Yuferev, 1962,
p. 205, pl. 9, fig. 13; Brazhnikova, ef al., 1967, pl. 21, fig. 2; Popova, 1973,
p. 57, pl. 9, figs. 7, 8; Bozorgnia, 1973, pp. 137, 138, pl. 30, figs. 14-16, 18-20.
Asteroarchaediscus rugosimilis Brenckle, 1973, p. 62, pl. 9, figs. 7-10.
?Asteroarchaediscus gnomellus Brenckle, 1973, p. 62, pl. 9, figs. 2-6.
Not Asteroarchaediscus pustulus gnomellus Brenckle, Vachard, 1975, pp. 63, 64.
Holotype. — Museum Inst. Geol. Sci. Acad. Sci., USSR, Mos-
cow, fig. 5, No. 2834/49.
210 BULLETIN 298
Original description. — (Translated from Ellis and Messina,
supplement No. 2, 1958).
Diagnosis. — Test small, discoidal with a slightly rough surface
and broadly rounded periphery, coils streptospirally wound with the
initial coils more tightly wound than the final; involute except for
the final one to one and a half coils which may be evolute, showing
a tendency not to embrace the preceding coils; serrated nodes
formed by the walls of the subjacent coils fill the lumina, with the
exception of the final coil. The final coil may be partially or com-
pletely closed. In other instances, and not infrequently, it is open
and free. Wall consists of a moderately thick, fibrous layer, and a
poorly developed, sometimes absent inner dark layer.
Measurements. — (Based on 15 specimens). Number of volu-
tions (two specimens): 4-5. Diameter: 140.00-207.50». Width: 65-
90u. Ratio W/D: 0.39-0.56. Proloculus (based on five specimens) :
18.32-28.00n. Height of lumen last volution (based on 15 speci-
mens): 0.00-17.50,. Peripheral wall thickness (based on nine speci-
mens): 6.87-14.00z.
Test form. — Flat to lenticular.
Coiling stage. — Angulatus.
Stratigraphic range.— North America — Visean (V3) —
Namurian. USSR — Visean (V3) — early Namurian. Iran —
Visean (V3) — early Namurian.
Remarks. — The majority of our specimens are under the mini-
mum range limit given by Rauzer-Chernoussova. In all other re-
spects, including ratio of width to diameter they seem similar.
Rauzer-Chernoussova described her forms as having a lumen height
in the last whorls of 15-20u, rarely up to 25. The photographic
illustration of her holotype, figure 5, does not give the appearance
of having an open lumen. Her illustration of paratype, figure 4,
shows the lumen of the final coil, at least in the northern hemisphere,
to be only partially open. The remaining paratype, figure 6 (a sag-
gital section) shows an open final lumen.
Brenckle compared his Ast. rugosimilis to Archaediscus rugosus
Rauzer-Chernoussova. He differentiated them on the basis of the
greater height of the lumina in the last volutions of A. rugosus. He
attributed this greater height to the possibiliy that A. rugosus might
belong to Neoarchaediscus. In Brenckle’s description of Ast. rugo-
ARCHAEDISCIDAE: BROWNE, Baxter, & RoBERTS 211
similis, based on six specimens, he stated “a few lumina in the later
volution are slightly open but their height does not exceed the thick-
ness of the surrounding wall.”
Brenckle, likewise, observed in studying his species Astero-
archaediscus gnomellus that while most of his specimens have closed
lumina the final lumen in a few specimens was high.
In studying the various species of (Asteroarchaediscus) the
authors have noted that variability in height of the final lumen in
a species seems to be a feature pertinent to the subgenus and can be
noted in some other species than those mentioned above.
We believe (Asteroarchaediscus) rugosvmilis is a junior synonym
for (Ast.) rugosus. Brenckle’s illustrations of Ast. rugosimilis com-
pare well with Rauzer-Chernoussova’s holotype of A. rugosus.
Asteroarchaediscus gnomellus Brenckle was distinguished from
Ast. rugosimilis on the basis of its somewhat smaller size and greater
width to diameter ratio. We now believe (Ast.) gnomellus is prob-
ably also a junior synonym of (Ast.) rugosus. Our specimens en-
compass the dimensions and width to diameter ratio of both
Brenckle’s species Ast. rugosimilis and Ast. gnomellus. We have,
however, questioned the placement of Ast. gnomellus in the above
synonym because Brenckle now feels (personal communication) that
while Ast. rugostmilis may be synonymous with Ast. rugosus our
specimens are transitional between his species Ast. rugosimilis and
Ast. gnomellus.
Differences between this species and (Ast.) postrugosus are
given under the description of the latter species. (Ast.) rugosus dif-
fers from (Ast.) parvus in having nodes that are more sharply
serrated or angular. (Ast.) rugosus is also less symmetrical with the
coils deviating more from the plane of symmetry.
Nodosarchaediscus (Asteroarchaediscus) syzranicus (Chernysheva), ee
1eAls PAs, sales
Permodiscus syzranicus Chernysheva, 1948, p. 156, pl. 2, fig. 10.
Permodiscus syzranicus Chernysheva, Grozdilova and Lebedeva (in Dain and
Grozdilova) p. 114, pl. 4, fig. 21.
Holotype. — Not given, probably Geol. Res. Inst., Leningrad,
No. 2640.
Original description. — (Translated from the Russian in Ellis
and Messina, supplement No. 1, 1958).
212 BuLLETIN 298
Diagnosis. — Test lenticular with well-rounded periphery and
slightly convex sides; initial coils, deviating slightly from the plane
of symmetry, are tightly wound and involute while the final coil is
evolute; proloculus indeterminate in this single specimen; lumina are
low and filled with nodes except for the final coil which is open
and has a high lumen; nodes, formed by the crenulate walls
of the preceding coils are numerous, small, finely and sharply ser-
rated; wall, which thickens toward center of the test, is composed of
a clear, thinly porous layer, and a little perceptible dark layer.
Measurements. — (Based on one specimen). Number of volu-
tions: five?. Diameter: 197.50u. Width: 80% Ratio W/D: 0.40.
Proloculus: not determinable. Height of lumen last volution: 16.25.
Peripheral wall thickness: 11.25p.
Test form. — Flat to lenticular.
Stratigraphic range. — North America — Visean (late V3c),
this report. USSR — Visean (V3c).
Remarks. — This species was placed in the genus Permodiscus
by Chernysheva due apparently to the side thickenings of the wall
and the fact she considered the coiling to be planispiral. Because this
form has a barely detectable inner dark layer, is without lateral
corner fillings or contreforts (sensu Conil) and the lumina are filled
by nodes the authors consider the generic designation in error.
Moreover, the initial coils, though small, appear to deviate from the
planispiral plane.
The distinguishing feature of this species we consider to be the
extremely small, finely serrated nodes.
Subfamily AMMARCHAEDISCINAE Conil and Pirlet, 1974
Archaediscidae without internal division, with planispiral enrollment. Wall
similar to that of the Archaediscinae. (Pirlet and Conil, 1974, p. 271.)
Genus AMMARCHAEDISCUS Conil and Pirlet, 1974
Type species: Ammarchaediscus (Amm.) bozorgniae Conil and
Pirlet, 1974.
Diagnosis. — Ammarchaediscinae possessing free lumina, with-
out nodes or stellate structure. Dark internal layer very pro-
nounced to imperceptible; external radial layer developed in the axial
region only or throughout the test. Coiling involute to evolute. (Pir-
let and Conil, 1974, p. 271.)
ARCHAEDISCIDAE: BRowNE, BAXTER, & ROBERTS 213
Subgenus A Conil and Pirlet, 1974
Diagnosis. — Ammarchaediscus with the dark inner layer moderately to
feebly developed, without lateral corner fillings (semsu Conil). The exterior
radial layer completely envelopes all the coils. Enrollment involute and the
lumina cresent shaped in the primitive forms; enrollment more or less evolute
and the floors of the Jumina concave in the evolved forms. (Pirlet and Conil,
1974, p. 278).
Ammarchaediscus (subgenus A) sp. Pl. 25, fig. 10
Figured specimen. — USNM (Nat. Mus. Nat. Hist.), No.
244637.
Diagnosis. — Test small, discoidal with greatest thickening at
axis of coiling, surface slightly rough; coiling dominantly planispiral
with slight oscillation in one hemisphere; inner coils involute and
final two evolute; open lumina of semilunular shape increase pro-
gressively in size becoming both broader and higher; floors flat with
epaulets or extensions on the walls which are more pronounced on
the later whorls; outer fibrous wall, covering all the whorls, thickens
approximately three and a half times in width from margin of test
to the center where it appears fused by the involute whorls; dark
microgranular, inner wall present in inner whorls is little perceptible
in outer whorls.
Measurements. — (Based on one specimen). Number of volu-
tions: five?. Diameter: 271.25. Width: 105. Ratio W/D: 0.387.
Proloculus: not determinable. Height of lumen last volution: 23.75p.
Peripheral wall thickness: 8.75y.
Coiling form. — Planispiral.
Cowling stage. — Angulatus.
Straitigraphic range. — Unknown. North America — Visean —
(late V3c), this report.
Remarks. — This species of Ammarchaediscus is at an advanced
evolutionary stage of the genus Ammarchaediscus with the flat
floors which appear at the same horizon in the genus Archaediscus.
It is a good example of the angulatus stage.
This species belongs to Conil and Pirlet subgenus A (1974,
p. 278, pl. 3, figs. 37-40). Our forms give the appearance of being
the same as Permodiscus vetustus Chernysheva (1948, fig. 15)
and Grozdilova and Lebedeva (in Dain and Grozdilova, 1953, pl.
4, fig. 23). The latter specimen was designated as lectotype for the
species P. vetustus because Chernysheva, who described the species,
214 BuLLETIN 298
had never chosen a holotype. Unfortunately, the center coils are not
exposed in the specimen selected as holotype so it can not be used.
Subgenus TUBISPIRODISCUS Browne and Pohl, 1973
Type species: Tubispirodiscus simplissmus Browne and Pohl,
1973.
Diagnosis. — Test free, flattened, concave, discoidal with the narrowest
dimension through the axis of revolution; composed of a proloculus followed
by a freely coiled undivided chamber which is planispirally enrolled and entire-
ly evolute throughout; periphery well-rounded and surface somewhat uneven
with evident sutures; side thickenings absent; wall bright calcareous composed
of a single fibroradiate layer only; aperture a circular opening at the end of
the tube. (Browne and Pohl, 1973, p. 202).
Ammarchaediscus (Tubispirodiscus) simplissimus (Browne and Pohl), 1973
Pl. 25, figs. 11-13
Tubispirodiscus simplissimus Browne and Pohl, 1973, pp. 202, 203, pl. 25, figs.
10-12; pl. 26, figs. 1-3.
Holotype. — USNM (Nat. Mus. Nat. Hist.), No. 186634.
Coiling form. — Planispiral.
Coiling stage. — Evolutus.
Stratigraphic range. — North America — Visean (V3c).
Remarks. — Because these specimens are the same as those
published previously by the authors no description is given.
Ammarchaediscus (Tubispirodiscus) sp. Pl. 25, figs. 14, 15
Diagnosis. — Test small, surface somewhat uneven; _ initial
chamber followed by tubular chamber wound in a planispiral plane,
wall a single, bright, fibroradiate, moderately thick layer.
Measurements. — (Based on three specimens). Number of volu-
tions: 4-5%. Diameter: 230-255. Width: not determined. Ratio
W/D: not determined. Height of lumen last volution: 25.00-39.50y.
Peripheral wall thickness: 9.00-11.60p.
Coiling form. — Planispiral.
Coiling stage. — Not determined.
Stratigraphic range. — North America — Visean (late V3c),
this report.
Remarks. — Because the three sections which form the basis
for the above description are all sagittal sections it is presently im-
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 215
possible to give a complete diagnosis of this species. It differs from the
type specimen of (Tubispirodiscus) simplissimus Browne and Pohl,
in its larger size, having a final lumen height, and wall thickness
approximately double that of the type specimen.
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Nova Scotia and New Brunswick. Geol. Sur. Canada, Dept. of
Energy, Mines, Resources, paper 70-21, pp. 1-64, 19 pls., 7 figs.
1973. Microfacies viseens du Boulonnats (Nord, France). Revue de
Micropaleontologie, No. 2, pp. 101-124, 9 pls.
Mamet, B. L., Choubert, G. and Hottinger, L.
1966. Note sur le Carbonifere du jebel Quarkziz. Etude du passage du
Viseen au Namuricn d’apres les Foraminiferes. Notes et Mém.,
Sér. Geol. Maroc, No. 198, pp. 6-20.
Mikhailov, A. D.
1939. On the characteristics of the genera of Lower Carboniferous
Foraminifera. In Malivkin, S. F., ed., The Lower Carboniferous
deposits of the northwestern limb of the Moscow Basin. Leningrad,
Geol. Admin, Symposium (Sbornik), No. 3, pp. 47-62, 4 pls.
Miklukho-Maklay, A. D.
1953. On systematics of the family Archaediscidae. Ezhegodnik Vses.
Paleont. Obshch., vol. 14, Otdel. Ottisk, pp. 127-131, 6 pls.
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Vses. Naucho-Issledov. Geol. Inst. (VSEGEI), Min. Geol. i
Okhrana Nedr. SSSR, n.s., vol. 12, pp. 9-15.
1957. New data and the systematics and phylogeny of the Archaediscidae.
Vestnik Leningrad Univ., No. 24, ser. Geol. and Geogr., No. 4,
pp. 34-46, 4 text-figs.
Moore, F. B.
1965. Millerstown (Grayson, Hart, and Hardin Cos.). Areal Geologic
map GQ-417, U.S. Geol. Sur.
Norwood, J. G.
1876. Report on the geology of the regions adjacent to the Louisville,
Paducah, and Southwestern Railroad, with a section. Kentucky
Geol. Sur., ser. 2, vol. 1, pp. 355-448.
Petryk, A. L.
1971. A new Lower Carboniferous Archaediscus (Foraminifera) from
southwestern Alberta. Micropaleontology, vol. 17, No. 2, pp. 249-
252, 2 text-figs.
Pirlet, H., and Conil, R.
1974. L’evolution des Archaediscidae, visean. Bull., Soc. belge Geol.,
t. 82, fasc, 2, pp. 241-299, 3 pls., 5 figs.
218 BULLETIN 298
Pohl, E. R.
1970. Upper Mississippian deposits of southcentral Kentucky. Kentucky
Acad. Sci., Trans., vol. 31 (1-2), pp. 1-15.
Pohl, E. R., Browne, R. G., and Chaplin, J. R.
1968. Foraminifera of the Fraileys member (Upper Mississippian) of
Central Kentucky. Jour. Paleont., vol. 42, No. 2, pp. 581-582.
Popova, J. G., and Reitlinger, E. A.
1973. Foraminifera: in The stratigraphy and fauna of the Carboniferous
from the River Shartym (southern Urals). Kiev, Ord. Len. Gosud.
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1948a. Some new species of Foraminifera from the Lower Carboniferous
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1948b. Foraminifera from the Carboniferous deposits of central Kazakh-
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1948c. Certain new Lower Carboniferous fauna from the Syzransky dis-
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Reitlinger, E. A.
1949. An account of the smaller Foraminifera in the lower part of the
Middle Carboniferous in the central Ural and Kama regions. Akad.
Nauk S.S.S.R., Izv., Geol. Ser., No. 6, pp. 149-164.
Schubert, R. J.
1908. Beitrage zu einer naturlichen Systematik der Foraminiferen.
Neues Jahrb. Mineral. Geol. Palaont., Teil-Bd., 25, pp. 232-260, 1
pl.
Schwalb, H. R.
1975. Oil and Gas in Butler County, Kentucky. Kentucky Geol. Sur. Re-
ports, Ser. 10, RI 16.
Shaver, R. H., Burger, A. M., Gates, G. R., Gray, H. H., Hutchinson, H. C.,
Keller, S. J., Patton, J. B., Rexroad, C. B., Smith, N. M., Wayne, W. J.,
and Weir, C. E.
1970. Compendium of rock-unit stratigraphy in Indiana. Indiana Geol.
Sur., Bull. 43, 229 pp.
Shlykova, T. I.
1951. Foraminifera of the Visean and Namurian (Lower Carboniferous)
stages of the western part of the Moscow Basin. Vses. Neft.
Nauchno. Issledov. Geol.-Razved. Inst. (VNIGRI). Trudy, vol. 56,
pp. 109-178, pls.
Sosipatrova, G. P.
1962. Foraminifera of the Upper Paleozoic of the Taimyr. Sbornik Stat.
p. paleont. i biostrat., Nauchno-Issledov, Inst. Geol. Arkt., Min.
Geol. i. Okhrana Nedr. S.S.S.R., vol. 30, Leningrad, pp. 35-72, 5
pls.
1966. Foraminifera of the Tiksinski suite of the northern Kharaulakh
Range, Uchen. Zapis. Paleont. i Biostrat., Nauchno-Issledov, Inst.
Geol. Arkt., Min. Geol. S.S.S.R., vol. 11, Leningrad, pp. 6-32, 3 pls.
Suleimanoy, I. S.
1948. Report on some Foraminifera of the Lower Carboniferous from the
region of Sterlitamak. Akad. Nauk S.S.S.R., Inst. Geol. Nauk, vol.
62, Geol. ser., No. 19, pp. 244-245, 4 figs.
Swann, D. H.
1963. Classification of Genevievian and Chesterian (Late Mississippian)
rocks of Illinois. Illinois State Geol. Sur., Report of Investigation
216, pp. 91, pl. 1, 23 figs. glossary.
Swann, D. H. and Atherton, E.
1948. Subsurface correlations of lower Chester strata of the Eastern In-
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ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 219
Vachard, D.
1975. Etude Systematique des Algues et des Foraminiferes. In Recherch-
ers de micropaleontologiques in Le Paleozotque Superieur du
Maroc Central by Henri Termier, Genevieve Termier and Daniel
Vachard, Cahiers de Micropaleontologies 4-1975, pp. 1-98.
Vdovenko, M. V.
1968. On an unusual assemblage of Foraminifera from the upper Visean
succession of the Dnieper-Donetz Basin. Paleontol. Sbornik, No. 5,
Isdatel. L’vov Univ., pp. 3-7, 2 pls.
Vincent, J. W.
1975. Lithofacies and biofacies of the Haney Limestone (Mississippian)
Illinois, Indiana, and Kentucky. Kentucky Geol. Sur., ser. X Thesis
Ser. 4, 64 pp., 23 figs., 3 tables.
Weller, J. M., and Sutton, A. H
1940. Mississippian border of Eastern Interior Basin. American Assoc.
Pet. Geol., Bull. vol. 24, pp. 765-858.
vit _ eemeagn fh as7 Bad, ator Can A
2 tem, fe B | 7 “<
eh (i ; nes iw
oad a Lost, Wies
tees WL tel ae =, > : -
vag! ates it
fecal
i See tro + WS, thm SOMOS
PLATES
222
Figure
a
2.
10.
ine
12.
13.
14.
15.
16.
17.
18.
19.
20.
BuLLETIN 298
EXPLANATION OF PLATE 22
All X 140
Page
Archaediscus cf. absimilis (SOSipatrOva) ...........ssscssssscsssssssssssseasees 179
Axial section, USNM No. 244568.
Archaediscus cf. absimilis (SoOSipatrOova) ............:ccsssscessssrsesssescess 179
Axial section, USNM No. 244569.
Archaediscus Cf. absimilis (SOSipatrOVa) .............ssccessssesscssssessensees 179
Tangential section, USNM No. 244570.
Archaediscus (Archaediscus) chernoussovensis Mamet ............. 180
Axial section through proloculus, USNM No. 244571.
Archaediscus (Archaediscus) conili, 1. SDP. ..........ccsssccessscserscsenses wo LOL
Axial section through proloculus, paratype, USNM No. 244572.
Archaediscus (Archaediscus) conili, 1. SP. .........:csssccecescssreceee eed kel!
Axial section, paratype, USNM No. 244573.
Archaediscus (Archaediscus) conili, 1. SD. ........:ccssssccssccessceesessns 181
Axial section, paratype, USNM No. 244574.
Archaediscus (Archaediscus) infantus Shlykova_ ...........ccsseseees . 182
Axial section, USNM No. 244575.
Archaediscus (Archaediscus) infantus Shlykova ...........ccccsceesees . 182
Axial section through proloculus, USNM No. 244576.
Archaediscus (Archaediscus) krestovnkovi Rauzer-
CHELMOUSSOVA scicceccececedsasabcinecccs:cucucccccsesceesavonsevacceesestezessteetererteereaeee . 184
Axial section, USNM No. 244577.
Archaediscus (Archaediscus) krestovnikovi Rauzer-
CHEETOUSSOV A: iivadscsaccesese ste scsdosvoscaccavanscseatincotocevscanescucdessevssieaeeeeeeeaenen 184
Axial section, USNM No. 244578.
Archaediscus (Archaediscus) miklukhomaklayi, 0. SD. ...........0+ 185
Axial section, paratype, USNM No. 244579.
Archaediscus (Archaediscus) miklukhomaklayi, ND. SP. ..........000 . 185
Axial section, paratype, USNM No. 244580.
Archaediscus (Archaediscus) ex. gr. moelleri Rauzer-
GRETMOUSSOV As acissccccoseeeccncossinscccncontnwsasesnecbeucccnsestectesseerscecsosmaneoneneemntes 186
Axial section, through proloculus, USNM No. 244581.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244582.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244583.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244584.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244585.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244586.
Archaediscus (Archaediscus) pusillus Rauzer-Chernoussova ...... 187
Axial section, through proloculus, USNM No. 244587.
BULL. AMER. PALEONT., VOL. 72 PLATE 22
PLATE 23
BuLL. AMER. PALEONT., VOL. 72
2
s
- wg poh Nye rig i%
f £eBe +. * pet Br ‘
- Meg Sry" any -)
a eh
ig 3
Pe, ae
fad ts
on
Figure
7-14.
15.
16.
INA
18.
19.
20.
ARCHAEDISCIDAE: BRowNE, BAxtTer, & ROBERTS 223
EXPLANATION OF PLATE 23
Page
Archaediscus (?Hemiarchaediscus) swanni, 1. SP. ..........s:ssss0ec000 190
Axial section, paratype, USNM No. 244588.
Archaediscus (?Hemiarchaediscus) swanni, N. SDP. ............sssccseee0e 190
Axial section, paratype, USNM No. 244589.
Archaediscus (?Hemiarchaediscus) swanni, 0. SP. ..........scceecceeeeee 190
Axial section, holotype, USNM No. 244590.
Archaediscus (?Hemiarchaediscus) swanni, 0. SP. ..............sseee008 190
Axial section, through proloculus, USNM No. 244591.
Archaediscus (?Hemiarchaediscus) swanni, 0. SD. ..............sesceeseee 190
Axial section, paratype, USNM No. 244592.
Archaediscus (?Hemiarchaediscus) cornuspiroides Vdovenko .. 191
Axial section, through proloculus. USNM No. 244593.
Archaediscus (?Hemiarchaediscus) stilus Grozdilova and
LOY) yey 0 CEN,” a eben Fat PRIS SS Rents SRN A oad» Mier dat SAI ehcatie a A 192
All axial sections, showing progressive evolutionary tendencies
from the more typical angulatus stage of figure 7 to the near
tenuis stage of figure 14. 7. USNM No. 244594. 8. USNM No.
244595. 9. USNM No. 244596. 10. USNM 244597. 11. USNM No.
244598. 12. At this magnification radial wall apparent only in
central part of test-angulatus stage tending to tenuis
stage. USNM No. 244599. 13. At this magnification radial wall
apparent only in central part of test-angulatus stage but less
radial and tending more towards tenuis stage than figure
10. USNM No. 244600. 14. Near tenuis stage with wall of
approximately uniform thickness. Shows tendency of floors
to become convex, wall porosity little apparent. USNM No.
244601.
Nodosarchaediscus (Nodasperodiscus) gregorii Dain .................. 195
Axial section, through proloculus, USNM No. 244602.
Nodosarchaediscus (Nodasperodiscus) gregorii Dain ................. 195
Axial section, through proloculus, USNM No. 244603.
Nodosarchaediscus (Nodasperodiscus) minimus Grozdilova
ETOYS | 3] GE) 0270 CS) 172 WARS See R a Pe neni oe ditt rd Caen Re oe 197
Axial section, through proloculus, USNM No. 244604.
Nodosarchaediscus (Nodasperodiscus) minimus Grozdilova
RIM C OO osu sone tatevacennnil svstpakasdnonnesbesdaspetess ands aectcemeeexeeeeeatoosctees 197
Axial section, USNM No. 244605.
Nodosarchaediscus (Nodasperodiscus) minimus Grozdilova
ANGE WEDCACV AL siccsascstvewcsessesetescen ceees socekuescucbeoben tke osecasucessnoscassaodesssaevetes 197
Axial section, USNM No. 244606.
Nodosarchaediscus (Neoarchaediscus) bykovensis Sosipatrova .. 199
Axial section, through proloculus, USNM No. 244607.
224
Figure
18.
19.
20.
BULLETIN 298
EXPLANATION OF PLATE 24
All * 140
Page
Nodosarchaediscus (Neoarchaediscus) bykovensis Sosipatrova .. 199
Axial section, USNM No. 244608.
Nodosarchaediscus (Neoarchaediscus) bykovensis Sosipatrova .. 199
Axial section, through proloculus, USNM No. 244609.
Nodosarchaediscus (Neoarchaediscus) incertus Grozdilova
ANG “EQ DOGEVA. c.cc).cccccccsessscscccocssconsevaceteesescsuctcosscsccsostsosscostectteneeenee ecsooe OO
Axial section, USNM No. 244610.
Nodosarchaediscus (Neoarchaediscus) incertus Grozdilova
ANGE TE DCAOV | ype sccscckvasstscecesesvaeoeuacceccccosenveoscesncsscasaucssausieaveskuleuaceeeenteeeee 2
Axial section, USNM No. 244611.
Nodosarchaediscus (Neoarchaediscus) incerftus Grozdilova
ANG, PEDEM OVA. escB eh eeec ck ebe ch Rock hacks sce deka wececteccavucttosontect coterceeremeeent 200
Axial section, USNM No. 244612.
Nodosarchaediscus (Neoarchaediscus) incertus Grozdilova
ANG LEDCCO VAN warceeakocccccssccneccSteccvecessancuces Oise ehestdsdeaducecobdeceee tae eee 200
Axial section, USNM No. 244613.
Nodosarchaediscus (Neoarchaediscus) latispiralis Grozdilova
ANGE TE DOC EVA Metcescecerscccessccesccrecastaccotensces succsacscecccsctocsncatessetretcaeeteeetreen . 202
Axial section, USNM No. 244614.
Nodosarchaediscus (Neoarchaediscus) latispiralis Grozdilova
ANI CDC CEC Vals occ ccccsaccesuasideexaiuseencuntessccst ooucecenatasscanaveseneiteusreceoteeeteee 202
Axial section, USNM No. 244615.
Nodosarchaediscus (Neoarchaediscus) latispiralis Grozdilova
ANA: LEDCACV A) Wesivesscssoushcsdsikdesvede den studakes Gobo seneldouctdeadesel doaseveunmeeesememnaeee 202
Axial section, through proloculus, USNM No. 244616.
Nodosarchaediscus (Neoarchaediscus) pohli, n. SD. ............cescceceee 203
Axial section, through proloculus, paratype, USNM No. 244617.
Nodosarchaediscus (Neoarchaediscus) pohli, N. SD. ...........cceesceeeee 203
Axial section, through proloculus, holotype, USNM No. 244618.
Nodosarchaediscus (Neoarchaediscus) pohli, 1. SDP. ...........c.sceceeeee 203
Axial section, paratype, USNM No. 244619.
Nodosarchaediscus (Neoarchaediscus) timanicus Reitlinger ...... 205
Axial section, USNM No. 244620.
Nodosarchaediscus (Neoarchaediscus) timanicus Reitlinger ...... 205
Axial section, USNM No. 244621.
Nodosarchaediscus (Neoarchaediscus) timanicus Reitlinger ...... 205
Axial section, USNM No. 244622.
Nodosarchaediscus (Neoarchaediscus) SD. ..........cccccecsssscseesseceeeeseers 204
Axial section, USNM No. 244623.
Nodosarchaediscus (Asteroarchaediscus) parvus Rauzer-
CHELPNOUSSOV As «sc issesasivsctvversecctesissccsseveoccoanecesctcuouncacdtee Meee Create rere Reeneeee 207
Axial section, through proloculus, USNM No. 244624.
Nodosarchaediscus (Asteroarchaediscus) parvus Rauzer-
HET HOUMBSOVAS ..usccocecesseeescecncseetoecaacerne nereaudoaceeenetescen: cat ceTeett een 207
Axial section, USNM No. 244625.
Nodosarchaediscus (Asteroarchaediscus) parvus Rauzer-
RU EETIOUSSO Val er csunxsheoiass ckpsonensddonasncseastanainn seausaxapncnust tuctereteuakes a: eae aan 207
Axial section, USNM No. 244626.
Nodosarchaediscus (Asteroarchaediscus) parvus Rauzer-
WRETNOUSSOV ES fast oee\ snes consciacesnaasscttudsteoucneacwancavsususcceenmtereaeerceueeeee te aananE 207
Axial section, through proloculus, USNM No. 244627.
BULL. AMER. PALEONT., VOL. 72 PLATE 24
BULL. AMER. PALEONT., VOL. 72 PLATE 25
Figure
10.
dt.
12.
13.
14.
15.
16.
ARCHAEDISCIDAE: BROWNE, BAXTER, & ROBERTS 225
EXPLANATION OF PLATE 25
All & 140
Nodosarchaediscus (Asteroarchaediscus) postrugosus
ROTEL GI eter ee teen ee eect cesctect ahakcat ostecseonsensddessevasesevouaereuscassshencesde 208
Axial section, USNM No. 244628.
Nodosarchaediscus (Asteroarchaediscus) postrugosus
CRONIES OT vac sasscsessttueseseds sansbecsaecsesatdevsessvsseusivacecsosatessccusdeasaucsssacstvsdsdesbenes 208
Axial section, USNM No. 244629.
Nodosarchaediscus (Asteroarchaediscus) postrugosus
PERG NBL TNE Tasso crtee eee cei cece ive ceecenves cost sces onde avestanesseesesecsusvashesvescderusssoes 208
Axial section, USNM No. 244630.
Nodosarchaediscus (Asteroarchaediscus) rugosus Rauzer-
WU ECENOUSSOV dir tce cater ceteccarvees joceeacnonsticwass <i intuanadetedeedetensnsdasatececstotecceae 209
Axial section, through proloculus, USNM No. 244631.
Nodosarchaediscus (Asteroarchaediscus) rugosus Rauzer-
WHETMOUSSOV A: iis cxcsscheccccesecscosssececessesnectaetoucede ces destanobae cossvascdtentcctesesteses 209
Axial section, USNM No. 244632.
Nodosarchaediscus (Asteroarchaediscus) rugosus Rauzer-
GIERNOUSSOV AR bcccistecccoston ss sooth toss un hace astoee eect aree nal Sevan ete esasedevddeds sseséeanabe 209
Axial section, through proloculus, USNM No. 244633.
Nodosarchaediscus (Asteroarchaediscus) rugosus Rauzer-
(GITEVTIOUSSOVAY cc .ceiscescsecucccccccseasstesetecesestocssancccctavarsecaccctcevecctusteccasssesscees 209
Axial section, through proloculus, USNM No. 244634.
Nodosarchaediscus (Asteroarchaediscus) rugosus Rauzer-
DIVEST OUSS OVS esere cerca c ccs eceet tae aereret eases scausiaavavasessesecssccssssescoseeens 209
Axial section, USNM No. 244635.
Nodosarchaediscus (Asteroarchaediscus) syzranicus
GEE TY SCV Aid eerics cocca esa tai tccadvonsnsctvtoncarcecsensastecoscscaberercesedsbessapcopoaases 211
Axial section, USNM No. 244636.
Ammarchaediscus (Subgenus A) sp. Conil and Pirlet ................ 213
Axial section, USNM No. 244637.
Ammarchaediscus (Tubispirodiscus) simplissimus Browne
UIT OTM Serres ces setetocsdacie ave con toueus ce scusdusoetex tucsecouencsesdeaastestscewaevensctes 214
Axial section, through proloculus, USNM No. 186635.
Ammarchaediscus (Tubispirodiscus) simplissimus Browne
ATTA LO Halen eee osce metre cacecta caucus ence tecsuatececnanencutssseadseeassesacseudcecsvonsnqad 214
Sagittal section, through proloculus, USNM No. 186636.
Ammarchaediscus (Tubispirodiscus) simplissimus Browne
BTEC EO HL soos ce seseccczaceskeasedecnceacecvscaeenttonk cctanctcosoeuscdteconaversttstseodeccestacsccedvs 214
Sagittal section, through proloculus, USNM No. 186634.
Ammarchaediscus (Tubispirodiscus) Sp. .............ccccssssssecccessessseacees 214
Sagittal section, through proloculus, USNM No. 244638.
Ammarchaediscus (Tubispirodiscus) SP. ...........ccccessssssscccesssssssssers 214
Sagittal section, through proloculus, USNM No. 244639.
Archaediscus sp. [MEW SUDZENUS] .............scccsseccesrccessccenscesnreseseesees 194
Axial section, through proloculus, USNM No. 244640.
INDEX
Note: Light face type refers to page numbers. Bold face type refers
to plate figures.
A
abseus?
Planoarchaediscus .. 179
absimilis,
Archaediscus. ......... 179
Archaediscus (Archae-
GISCUS) a Chip cccevecezeze 22 179
Planoarchaediscus .. 179, 180
Ammarchaediscus ....... iizale ales, irs
191, 212, 213
Ammarchaediscus
(Tubispirodiscus) .... 178, 189
Ammarchaediscus
(Subgenus A) .......... 176
Archaediscus ............. iyAla ilefon INTE
178, 179, 189,
190, 191, 213
Archaediscus
(Archaediscus) ........ 176, 178, 191,
213
Archaediscus
(-Hemiarchaediscus) 176, 178, rey
Archaediscus
(new subgenus) ...... 194
AsperodiscuS ...........4 176, 198
Asteroarchaediscus .... 171, 176, 177,
178, 198, 206,
208, 211
B
BAS CHERATIAM ete vevecsscvsess 180, 193, 196,
197, 198, 200,
201, 202, 204,
206, 209
baschkiricus,
Archaediscus .......... 202, 206
Beech Creek
TeAMESTOME -cceeseeeseeees 173, 174, 175
Big Clifty Formation.. 171, 173, Eo
borealis,
Neoarchaediscus .... 199
bozorgniae,
Ammarchaediscus .. 212
IBSTOAGTOUG peereeerteceee teases 173
BrumnStae ero os iyi
ISCUNISTINI Me esteteeeceeece 175
bykovensis,
Neoarchaediscus .... 199
Nodosoarchaediscus
(Neoarchaediscus)
CER rrr sa 23 199, 200
Cc
Carboniferous ............ 186
chernoussovensis,
Archaediscus .......... 177, 181
Archaediscus
(Archaediscus) ....22 180, 181
Archaediscus subsp.
angulatus' ! .c.ccene.2: 180
conili,
Archaediscus
(Archaediscus) ..22 181, 183
cornuspiroides,
?Archaediscus ........ 191
Archaediscus (?Hemi-
archaediscus) ...... 23 191
Cypress Sandstone .... 173, 174, 175
E
Hosigmolinate ee 191
Frailey’s Shale ............ 171, 172, 4,
175, 176
PUSUIININA YS 2 os «cee 178
G
Glen Dean ...:cenases 174, 175
Glomodiscus ..............++ 177
gnomellus,
Asteroarchaediscus 209, 211
Golconda Formation .. 174
gregorii,
Archaediscus .......... 196
Asteroarchaediscus 197
Nodosarchaediscus
(Nodaspero-
GUISCUS) | do5cc- -cevssesseocs 23 195, 201
Planospirodiscus .... 195
gregorii “var.” gregorii,
Archaediscus .......... 195
Neoarchaediscus ..... 197, 201
H
Haney Limestone ....... 173, 174, 175
Efardinis DUM i yeccaseeceeneseee 174, 175
Hemiarchaediscus ...... 176, 188, cPH
?Hemiarchaediscus .... 171, 176, 190,
192, 195
Hemigordius _ ............+. 186
a vautetevenne 173, 174, 175
Hombergian
226
INDEX
incertus,
Archaediscus .......... 198, 200
Neoarchaediscus .... 197, 200, 201
Nodosarchaediscus
(Neoarchae-
GISCUS)) kesscccceeeecens 24 200
incertus “var.” incertus,
Neoarchaediscus .... 201, 206
infantus,
Archaediscus .......... 182
Archaediscus
(Archaediscus) ....22 182
Archaediscus aff. .... 181, 182
K
karreri,
Archaediscus .......... 177, 178, ate
8
koktjubensis,
Archaediscus .......... 184
koktjubensis subsp.
krestovnikovi,
Archaediscus .......... 184
krestovnikovi,
Archaediscus .......... 183, 184, 185,
188, 206
Archaediscus (Archae-
GISCUIS) Rte ene 22 184
krestovnikovi “var.”
koktjubensis,
Archaediscus .......... 184, 185
krestovnikovi “var.”
krestovnikovi,
Archaediscus .......... 184
krestovnikovi “var.”
pusillus,
Archaediscus .......... 187
krestovnikovi
forma typica,
Archaediscus .......... 184
L
latispiralis,
Archaediscus .......... 202
Archaediscus aff. .. 202
Archaediscus ?(Rugo-
archaediscus) .......... 202
Neoarchaediscus .... 202
Nodosarchaediscus
(Neoarchae-
DISCUS) Gesceesaicesestsce 24 177202
M
maximus,
Archaediscus .......... 195
mikluklo-maklayi,
Archaediscus
(Archaediscus) ....22 185
minimus,
Archaediscus .......... 197, 203, 204
Archaediscus (?) .... 197, 203, 204
Planospirodiscus .... 197, 199, 203
Nodosarchaediscus
(Nodaspero-
ISCUS) 9 occ cesecsteanee 23 196, 197, 201,
204
moelleri,
Archaediscus
EXOT iy secccccesernvecueess 22 186, 198
N
INEWETEBB ENE, Goceccsnnconconconde 178, 184, 187,
188, 192, 196,
198, 199, 206,
207, 209, 210
nana,
Archaediscus “var.” 187
Neoarchaediscus ........ U7 i. 198:
199, 204, 210
Nodasperodiscus ........ 171, 176, 1377,
178, 195, 204
Nodosarchaediscus .... 171, 176, 178,
195, 198, 207
Nodosarchaediscus
(Asteroarchaediscus) 176, 177
Nodosarchaediscus
(Neoarchaediscus) .. 176, 177, 178
Nodosarchaediscus
(Nodasperodiscus) .. 176, 177, 178
P
parvus,
Archaediscus .......... 178, 207, 208
Asteroarchaediscus 207, 211
Neoarchaediscus .. 207
Neoarchaediscus cf. 207
Nodosarchaediscus
(Asteroarchae-
GISCUS) pee erence: 24 178, 207
parvus regularis,
Archaediscus “var.” 209
pauxillus,
Archaediscus .......... 187
RETEMOGISCUS eee sresssscste 197, 212
?Permodiscus ...........0 176, 197
Planoarchaediscus ...... 188, 189, 190
Planospirodiscus ........ 197
planus,
Hemiarchaediscus .. 188, 189
pohli,
Nodosarchaediscus
(Neoarchae-
GISCUS) BSF. sscecc5-:3 24 203
postrugosus,
Archaediscus .......... 178, 208
Asteroarchaediscus 177, 178, 208
227
INDEX
Neoarchaediscus .... 208
Nodosarchaediscus
(Asteroarchae-
GISCUS)) Verteecccnsestess 25 177, 178, 208,
209, 211
Propermodiscus .......... 186, 189
Pseudoammodiscus .... avis)
pusillus,
Archaediscus .......... 187
Archaediscus
(Archaediscus) ....22 187
pustulus gnomellus,
Asteroarchaediscus ‘209
R
Rugosarchaediscus ..... 199
rugosimilis,
Asteroarchaediscus 209, 210, 211
rugosus,
Archaediscus .......... 178, 205, 208,
209, 210, 211
Asteroarchaediscus 196, 205, 209,
211
Neoarchaediscus .... 209, 210
Nodosarchaediscus
(Asteroarchae-
discus)
suesucesat cesusaes 25 178, 196, 209,
211
Ss
saleii,
Archaediscus .......... 195
schlumbergeri,
Hemigordius ............ 185
schlumbergi,
Cornuspira ce 185, 186
simplissimus,
Ammarchaediscus
(Tubispiro-
GISCUS) Pe. bP 25 214
Tubispirodiscus ...... 214, 215
sp.,
Ammarchaediscus
(Tubispiro-
GIS@US)! cect sccessseererce 25 214
sp.,
Nodosarchaediscus
(Neoarchae-
GISCUS) Meee 24 204
spirillinoides,
Archaediscus .......... 193
stilus,
Archaediscus .......... 177, 185, 192,
193, 194, 195
Archaediscus cf.
ex gr. stilus, subsp.
QNGWIACUS <2... cenncscnnes 192, 195
Archaediscus (?Hemi-
archaediscus) ...... 23 178, 191, 192
Planoarchaediscus .. 192
Planoarchaediscus? 190
Planoarchaediscus
forma compressa .... 192
Planoarchaediscus
formal typiCal..-c-.-caee 192
?Planoarchaediscus
forma magna ..........
Subgenus) Avene 25 176, 195, 213
subplanus,
Neoarchaediscus .... 199
sulcus,
Planospirodiscus .... 203
swanni,
Archaediscus (?Hemi-
archaediscus) ...... 23 190, 191
syzranicus, ;
Permodiscus ............ 211
Nodosarchaediscus
(Asteroarchae-
GISCUIS)» deeciecceedeceavs 25 178, 211
T
taimiricus,
Planospirodiscus .... 197
RECT ACARIS /ccccccceccccceeeetre 175
timanicus,
Archaediscus .......... 205
Neoarchaediscus .... 205
Nodosarchaediscus
(Neoarchae-
GISCUS) pete 25 205
Tournarchaediscus .... 178
Tubispirodiscus .......... 171, 214
U
ulmeri,
Hemigordius _.......... 186, 189
V
WETGAN Gicccccccssvencceutaee 206
vestustus,
Permodiscus ............ 213
VIUSEAN: -. cciicucdecteuasexcorsee 176, 181, 182,
183, 184, 187,
228
188, 191, 192,
193, 194, 196,
201, 202, 204,
207, 209, 210
<= 08 - 1p we
of bet Soe .. :
Wt 1568)
LII.
LIT.
LIV.
LV.
LVI.
LVII.
LVIII.
LIX.
LX.
LXI.
LXII.
LXIII.
LXIV.
LXV.
LXVI.
LXVII.
LXVIII.
LXIX.
LXX.
LXXI.
LXXII.
(ds ESN 8 8 6) ae Ey AT [UE cc Ti «| | ce ee ee Or
New Zealand forams, Stromatoporoidea, Indo-Pacific, Mio-
cene-Pliocene California forams.
(Nos. 237-238). AS SMD pet 5 wp Say care cese cares swessecceeetees cesecme seen toae
Venezuela Bryozoa, Kinderhookian Brachiopods.
(Nos. 239-245). EMU): Sop SUN 0) i eres pete ee eae amr ee ee en
Dominican ostracodes, Lepidocyclina, mollusks.
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(No. 291). PALAIS VY RRS We 0) (Fh eee er ter eran ee ree eee
Bryozoa.
(Nos. 292-294). AGA myn, AD DL Ss pesos see os eaccee sett octes + se weasueacoocccevions
Turrids, gastropods, forams.
(No. 295). EIPASAG) 0} oar ons) Ook) 0) fst goatee eee ci eer Sen ee
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2 OF
AMERICAN
PALEONTOLOGY
(Founded 1895)
Vol. 72
No. 299
SCALPELLID BARNACLES (CIRRIPEDIA) OF
FLORIDA AND OF SURROUNDING WATERS
By
Norman E. WEIsBorD
1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
PALEONTOLOGICAL RESEARCH INSTITUTION
1976-1979
PRESIDENT yiice tee eet ee een ee ete bY ale Ee Be Se eee HAROLD E. VoKESs
IMICE=PRESID PNT: sore eee ees oe teens erie 1 eee SEC Duane O. LeRoy
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Poke ON TOLOG y
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Vol. 72
No. 299
SCALPELLID BARNACLES (CIRRIPEDIA) OF
FLORIDA AND OF SURROUNDING WATERS
By
NorMan E. WEIsSBoRD
December 29, 1977
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
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CONTENTS
Page
ENDRURENE — eeetedsccecheconsecneee tee coecbe concaeeeccnedebadp eset teehee bere acP nae eee eee eee OS eee PEE 235
AVR fe Co CAIUT CG ro Ta se ee ree aSecnenereee reescencecrerce 235
PNGKMO Wl eG SMe Sy merece ocean ce eat ete atone nous cu cat as sncaesbcenct nas ceeseecescasesces 236
TURE CoE OLSEN cece a he ee ge aE a ee ee 236
ESCM HOME GL ASP COLES asec: sy cee Race ek a aca re ces eee n anon Pate anna Casa cesceceeene 237
TRS STRSTR En POR eS 290
TPS VRS. eee oe ae a ee Ne ae DE ee Se ee ee ee eee eee 291
SCALPELLID BARNACLES (CIRRIPEDIA) OF FLORIDA
AND OF SURROUNDING WATERS
Norman E. WEIsBorpD
Department of Geology
The Florida State University
ABSTRACT
This work is an annotated inventory of the barnacles of the family Scal-
pellidae occuring in and immediately off Florida as well as in the surrounding
waters of the Gulf of Mexico, the Caribbean Sea, and the Western Atlantic
Ocean. Some 39 species are described, and data are submitted on their geo-
graphic range, depth of water, habitat, and substrate. The type specimen of
each taxon described herein is illustrated from the original author’s portrayal
to facilitate comparison of all species as well as certain of those synonymized.
All of the scalpellids discussed are living; however, one of the species, Scal-
pellum gibbum Pilsbry, has also been found in the early Pliocene of Florida,
and another, Arcoscalpellum michelottianum (Seguenza) s.s., occurs in the
Pliocene of Italy and Sicily. It is believed that a number of species now known
from the waters surrounding Florida will be found off Florida itself, and that
a few more will eventually be discovered as fossils in the Southeastern Coastal
Plain of the United States.
INTRODUCTION
This paper is the second of a series dealing with the barnacles
of Florida and of the waters around it. The first of the series (Weis-
bord, 1975) dealt with the orders Acrothoracica and Rhizocephala,
and the present one is concerned with the family Scalpellidae.
Most of the information contained in this work has been culled
from published sources. Each species is described, and the type of
the species re-figured from the author’s own illustration. The type
locality, the geographic range, the habitat, and the substrate are
noted. Synonymies proposed by authors are listed, and it seems from
this study that some of the synonymized species may have to be re-
assigned to their original nomenclatural status.
Surrounding Florida are the Gulf of Mexico, the Caribbean Sea,
and the Western Atlantic Ocean. Many of the scalpellids from these
adjacent seas are described in this report even though their presence
in Florida has not yet been verified. Nevertheless a number of species
first recorded from the Gulf, Caribbean, or Western Atlantic have
later been identified in Florida and there is every reason to believe
that more of them will be in the future. Only one fossil scalpellid,
Scalpellum gibbum Pilsbry, has been reported from the mainland
of Florida, and this occurs in the Ecphora zone of the Jackson Bluff
Formation, in Leon County. The age of the Jackson Bluff Forma-
tion is late Miocene or early Pliocene.
The present report is in part an updated inventory of the scal-
pellid species reported within the region under consideration. The
generic classification of these species is based on the work of Withers
236 BuLLETIN 299
(1953) and of Newman, Zullo, and Withers (1969) in the Treatise
on Invertebrate Paleontology. One of the taxonomic difficulties I
have encountered is with the genus Scalpellum sensu stricto. Former-
ly, Scalpellum sensu lato included a host of species which today are
properly assigned to many different scalpellid genera including Scal-
pellum s.s. Unfortunately all of the species under the “old” use of
Scalpellum have not been sufficiently studied to determine whether
they should be retained in Scalpellum s.s. or reassigned to another
genus. In this work I employ the genus Scalpelluwm Leach s.s. a) for
examples meeting the modern diagnosis of the genus; b) for exam-
ples named Scalpellum by the original author and not yet changed
by later taxonomists; and c) for examples whose original generic
name of Scalpellwm is in doubt and which I have questioned, but
whose correct identification has not yet been established. Many of
the older scalpellid species discussed in the present report fall in the
last category, although I think eventually some of them will be
found to belong to genera other than Scalpellum s-s.
Briefly, Scalpellum s.s. of Leach consists generally of 14 plates,
usually wholly calcified. The carina is angularly bent, with the umbo
removed from the apex. The apical area and carinal side of the
scutum are extended and alate, with the umbo removed from the
apex, and the extended sides sometimes obscured by the overlapping
tergum and upper latus. The inframedian latus is large with the
umbo varying in position from middle to basal. As these external
characters are not always apparent, especially in the whole animal
with closely articulated valves or obscurative integument, careful
examination of both the outer plates and inner organs is necessary
for definitive identification.
ACKNOWLEDGMENTS
I am greatly indebted to former workers and contemporary
cirripedologists for the considerable knowledge contained in, and
imparted by their writings. I also wish to thank Katherine V. W.
Palmer of the Paleontological Research Institution for editing and
attending to matters relative to publication of this paper.
LIST OF SPECIES
Each species discussed in this report is listed in the tabulation
below irrespective of the synonymous status of some of them. The
latitudes and longitudes are predominantly those given by authors
but a few have been obtained from reference points measured to
FLORIDIAN CIRRIPEDIA: WEISBORD 237
scale in the mid-century edition of the “Times Atlas of the World.”
English and metric systems are used interchangeably in the body
of the report depending more or less on the first usage by the taxono-
mist, but the depths of water given in the following list are expressed
in meters. Considerable data contained in the records of the United
States Fish Commission steamer “Albatross” have been adopted
from the work of C. H. Townsend (1901).
DESCRIPTION OF SPECIES
Class CIRRIPEDIA Burmeister, 1834
Order THORACICA Darwin, 1854
Suborder LEPADOMORPHA Pilsbry, 1916
Family SCALPELLIDAE Pilsbry, 1916
Scalpellum (?) albatrossianum Pilsbry PIF 26; figs 1
Scalpellum tenue Annandale, 1905, p. 83; not of Hoek, 1883, p. 119, pl. 4,
figs. 20-21, fide Broch, 1953, p. 6.
Scalpellum albatrossianum Pilsbry, 1907, pp. 47, 54-55, fig. 19; Annandale,
1908, pl. 3, fig. 10; 1913, pp. 228, 229, 232; 1916a, pp. 128, 130, pl. 6, fig. 9;
1916b, p. 282; Stubbings, 1936, pp. 56, 57, 62, 64, 66, text-fig. 24; Nilsson-Cantell,
1938, pp. 7, 18; 1955, p. 218; Krtiger, 1940, p. 60; Broch, 1953, pp. 5, 6-7, 10,
12, 15, figs. 3a-c; Zullo, 1968, p. 211; Lakshmana Rao and Newman, 1972, p. 84;
Zevina, 1973a, pp. 847-848.
Not Scalpellum albatrossianum Nilsson-Cantell, 1926a, pp. 7-11, text-figs.
2a-j [= Scalpellum striolatum G. O. Sars, fide Broch, 1953, pp. 6, 7.]
The capitulum of the type is 10.5 mm in length and 5.3 mm in
width and is composed of 13 fully calcified plates separated by nar-
row chitinous sutures. The plates are distinctly marked with lines of
growth, and are covered with a thin, shortly and sparsely pilose
cuticle.
The tergum is triangular, its occludent margin arcuate, its
lateral margin convex below, slightly kinked at the apex of the
carina, the basal margin long and nearly straight. The scutum is
elongate, its carinal margin deeply sinuated for the apex of the upper
latus just below the tergo-lateral angle, which is extended in a nar-
row acute lobe; the baso-lateral angle is rounded and rests for a
short distance against the inframedian latus. The upper latus is ir-
regularly pentagonal in outline; the scutal margin is longest and
slightly concave, the tergal a little shorter and convex, the basal
margin short and in contact with the summit of the inframedian
latus; the umbo is terminal, projecting into a recess of the scutum.
The carina is strongly arched above, less so below; the umbo is
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240 BULLETIN 299
apical against but not between the terga; the roof is flat, bounded
by low narrow ribs; the sides are wide in the upper half, narrower,
and tapering in the lower.
There is no rostrum. The rostral latus is wider than high, tri-
angular in outline. The inframedian latus is suboblong with concave
margins, about three times longer than wide, with a slightly raised
mucro from which low ribs radiate to the angles of the plate; the
upper end is truncated. The carinal latus is about twice as wide as
high, with the umbo slightly projecting behind it at the lower fifth
of the carinal margin; the carinal margin is slightly concave above
the umbo, convex below it.
The peduncle is 3.5 mm in length, and is covered with rounded
imbricating scales, in nine rows of about eight scales each.
Type locality. — “Albatross” sta. 2226 (37°N, 71°54’W), 2045
fathoms, Globigerina ooze, bottom temperature 36.8°F, about 220
statute miles east of Newport News, Virginia. Among other localities
this is the nearest to Florida, which is some 7 degrees of latitude
farther south.
Other localities. — Bay of Bengal (9°34’N, 85°43’15”E); “In-
golf” sta. 10 (64°24’N, 28°50’W), 1484 meters, 3.5°C. bottom tem-
perature, about 200 statute miles west of Reykjavik, Iceland; Wan-
del, 1889 (65°36’N, 56°24’W), 349 fathoms, northwest of Godthaab,
Greenland.
Depths range from 349 fathoms (638 meters) in the North
Atlantic west of Greenland to 2045 fathoms (3742 meters) in the
Western Atlantic off Virginia. The geographic extremes are Green-
land in the north and west, and the Bay of Bengal in the south and
east.
Scalpellum (?) antillarum Pilsbry Pl. 26, fig. 2
Scalpellum antillarum Pilsbry, 1907, pp. 48, 61-62, fig. 24a-c; Henry, 1954,
p. 444; Zullo, 1968, p. 211.
Pilsbry’s description is recapitulated as follows:
The capitulum of the type is long oval, 11 mm in length and
5.7 mm in breadth, and is composed of 13 or 14 plates. The plates
are sculptured with widely spaced wrinkles conforming with growth
lines, and there are some extremely faint radial striae. The cuticle
is very thin and somewhat hairy on the carina, sutures, and peduncle.
FLORIDIAN CIRRIPEDIA: WEISBORD 241
The tergum is sharply triangular, with nearly straight margins
and an erect apex; shortly below the apex there is a sharp recess
in the carinal margin in which is nestled the apex of the carina. The
scutum is irregular in shape, the lower half wider than the upper;
the occludent margin is convex, the lateral margin weakly sigmoid,
and the basal margin straight; the apex is acuminate and a little
recurved, overlying the lower angle of the tergum. The carina is long
and evenly arched, with a length of 10 mm and the diameter at the
base 1.1 mm, extending between the carinal latera to the peduncle;
the roof is flat, bounded by acute angles, which toward the upper
part project a little forming narrow marginal ribs; the sides are
narrow.
The upper latus is obliquely spatulate, the basal angle directly
above the inframedian latus. The rostrum is represented by a linear
vestige almost concealed in the cuticle. The rostral latera are sub-
quadrate, the occludent and scutal margins straight, nearly equal,
and at right angles; the basal margin is not much more than half as
long as the scutal, and the lateral margin is weakly sigmoid. The
inframedian latus is small, narrowly triangular, about half the height
of the rostral latera; the umbo is at the obtuse apex. The carinal
latus is twice as high as wide, with four unequal sides, no two of
them parallel; the umbo is at the lower sixth of the straight carinal
margin, and projects slightly beyond the carina; the carinal margin
is longest, the basal and lateral margins short and nearly equal; the
upper angle is acute.
The peduncle is 4.5 mm in length, with about eight rows of
eight narrow, transversely lengthened scales each, the intervals be-
tween them hairy.
According to Pilsbry, the peculiar shape of the upper latus and
the long carina which passes between and entirely separates the
carinal latera, are the more conspicuous features of this species.
Type locality. — “Albatross” sta. 2384 (28°45’N, 88°15’30”W),
Gulf of Mexico, 940 fathoms (1719 meters), bottom temperature
39.6°F, bottom of brown and gray mud, about 135 statute miles
southwest of Pensacola, Florida, and 140 statute miles southeast
of New Orleans, Louisiana.
Remarks. — This species is one of several originally described
242 BULLETIN 299
under the old Scalpellum sensu lato that does not fit into the genus
Scalpellum sensu stricto as defined by modern taxonomists.
Scalpellum arietinum Pilsbry Pl. 26, figs. 3-5
Scalpellum arietinum Pilsbry, 1907, pp. 26, 43-45, fig. 13a-b; 1953, pp. 19-21,
fig. 4a-f, pl. 1, fig. 5; MacDonald, 1929, p. 531; Henry, 1954, p. 444; Hulings,
1961, p. 216; Wells, 1966, pp. 89-90; Zullo, 1968, p. 211.
The capitulum of the type is subtrapezoidal, and measures 11.6
mm in length and 7 mm in breadth. There are 14 fully calcified
plates, sculptured by growth lines and indistinct radial striae; the
occludent margins of the terga and scuta are straight and thus
determine the occludent border of the capitulum. The cuticle is thin
and inconspicuous, and nearly smooth except on the roof of the
carina where it is finely and shortly pilose and crossed by six to
eight transverse tufts of longer hairs.
The tergum is triangular and much longer than the scutum.
The scutum is about twice as long as wide, its lateral margin concave
near the tergolateral angle, convex below; the umbo 1s nearly termi-
nal, bent inward. The carina is moderately arched and extends a short
distance above the prominent umbo; it is 12 mm in length and its
diameter near the base is 1.7 mm; the roof is nearly flat, bounded
by distinct but obtuse angles, with a low rib on each side running
parallel with and near the angles; the lines of growth on the roof
are nearly U-shaped; the sides are flat and wide in the upper half,
delicately marked with fine longitudinal and radial striae. The basal
margin of the carina is deeply rounded, and there is a chitinous space
between the carina and the other plates.
The upper latus is pentagonal, the scutal and carinal margins
about equal and parallel, the tergal margin slightly longer; the basal
margin is shorter and the oblique lateral margin against the infra-
median plate still shorter; the umbo is not quite apical, the apex
beyond it obtuse, rounded, and white. The rostrum is small triangu-
lar, and with equal sides. The rostral latus is low, about five times
as long as high, somewhat narrower in front than at the lateral end,
and stands out in relief above the surface of the scutum. The infra-
median latus is about twice as high as wide and irregularly penta-
gonal; the umbo is elevated, and from it obtuse ridges radiate to the
two basal angles of the plate. The carinal latus is somewhat tri-
angular; its apical half projects free behind and below the carina,
FLorIDIAN CIRRIPEDIA: WEISBORD 243
flaring strongly outward and noticeably twisted; the apex is acute. In
dorsal view the carina is seen to extend between the hornlike latera
nearly to the peduncle, the spread from apex to apex of the two
carinal latera 6 mm in the type specimen. These laterally flaring,
hornlike carinal latera are the distinguishing character of the species.
Type locahty.— “Albatross” sta. 2405 (28°45’N, 85°02’W),
Gulf of Mexico, 30 fathoms (55 meters), gray sand and broken
coral, about 267 statute miles west off Homosassa, Florida.
Other localities —“Albatross” sta. 2315 (24°26’N, 81°48’15”W),
37 fathoms (68 meters), coral bottom, on spines of Cidaris tribu-
loides, about 10 statute miles south of Key West, Florida; “Triton”
sta. 484, off Palm Beach, Florida, on vermetid shell; Cape St.
George, St. George Island, Franklin County, Florida, on calico scal-
lop Aequipecten gibbus, 30-100 ft, 15 miles south of Alligator Point,
Franklin County, Florida.
Scalpellum carinatum Hoek Pl. 27, figs. 3, 4
Scalpellum carinatum Hoek, 1883, pp. 29, 31, 63, 67, 76-77, 102, pl. 3, figs.
7-8; Weltner, 1895, p. 289; 1897, p. 247; Murray, 1896, pp. 385, 397; Gruvel,
19 0Zayepeeo 9 1905) pn 50) figs 551920" pp. 20) ik pl, 7 hea 8s Bilsbry, 11907,
pp. 47, 53, figs. 18a, b; Barnard. 1925, pp. 3-4; Broch, 1953, pp. 7, 10, 12, 15.
Scalpellum imperfectum Pilsbry, 1907, pp. 70, 75-77, fig. 30, pl. 4, figs. 15-18
[fide Barnard, 1924, p. 47]; Barnard, 1925, pp. 3-4.
Hoek’s description of the type is summarized as follows:
The capitulum is 16 mm in length, is covered by a transparent
chitinous membrane, and consists of 14 smooth valves separated
by broad chitinous interspaces.
The tergum is large, triangular, and flattish, the apex recurved,
the carinal margin excavated and concave and divided into a small
superior and long inferior part. The scutum is elongated, two and a
half times longer than wide, its apex pointed, its basal margin
forming a right angle with the occludent margin but passing with a
rounded angle into the lateral margin. The lateral margin is slightly
hollowed out and is separated from the tergal margin by a smallish
shoulder. The umbo is at the uppermost point.
The carina is simply bowed, with a flat roof much increasing
in width from the upper to the lower end, and bordered on each side
by an indistinct ridge. The umbo is seated at the top of the roof a
short distance down from the apex. The part above the umbo is
formed by the upward prolongation of the sides of the valve.
244 BULLETIN 299
The upper latus is irregularly pentagonal, its upper half nar-
rowed, the lower half broad. The umbo is near but not at the apex.
The rostrum is elongated, extremely narrowed, enclosed between
the two rostral sides of the rostral latera. The rostral latera are con-
vex and fit into the inframedian latus which is wine-glass shaped with
a foot. The carinal latus is large and flat; the umbones of the two
valves almost touch each other under the middle of the carina and
project over the base of the carina.
The peduncle is about 6 mm in length, nearly cylindrical. The
scales are highly calcareous and white, placed in about seven longi-
tudinal rows, each row bearing four to six scales.
Type locality. — “Challenger” sta. 235, near the Island of Tris-
tan da Cunha (37°15’S, 12°30’W), depth 1000 fathoms (1829
meters), bottom of rock and shells.
Other localities. — “Challenger” sta. 2111 (35°0950’N,
74°57'40”W ), 938 fathoms (1700 meters), about 22 statute miles
east off Cape Hatteras, North Carolina. This is the nearest locality
to Florida. “Challenger” sta. 2731 (36°45’N, 75°28’W), 781 fathoms
(1428 meters), about 65 statute miles off Norfolk, Virginia, on
Arcoscalpellum velutinum (Hoek); “Ingolf” sta. 83 (62°25’N,
28°30’W), 1717 meters, 3.5°C bottom temperature in the North
Atlantic about 220 statute miles southwest off Reykjavik, Iceland;
“Prince de Monaco” Campagne 1895 (38°21’N, 37°41’W), Eastern
Atlantic, 2028 meters (1109 fathoms), brownish gray limy mud,
about 40 statute miles southwest of Lisbon, Portugal.
Scalpellum carinatum ranges from Iceland in the North Atlantic
to Tristan da Cunha in the South Atlantic, and occurs at depths
ranging from 1428 meters off Norfolk, Virginia, to 2028 meters off
Lisbon, Portugal.
Scalpellum (?) diceratum Pilsbry Pl. 27, figss, 6
Scalpellum diceratum Pilsbry, 1907, pp. 26, 45-46, figs. 14a, b; 1953, p. 21,
pl. 1, fig. 4; MacDonald, 1929, p. 532; Kriiger, 1940, p. 43, fig. 28d; Henry,
1954, p. 444; Ross, Cerame-Vivas, and McCloskey, 1964, p. 312; Cerame-Vivas
and Gray, 1966, p. 263; Zullo, 1968, p. 212; Bayer, Voss, and Robins, 1970,
p. A43.
Pilsbry’s description is summarized as follows:
The capitulum is subtrapezoidal, 13.5 mm in length, 7.8 mm in
width, and has a shape similar to that of S. arietinum Pilsbry. The
FLORIDIAN CIRRIPEDIA: WEISBORD 245
ventral margin is nearly straight, but with a low prominence in the
middle. The tergum is longer than the scutum, subpentagonal in
outline. The scutum has a terminal mucro, a gently convex occlu-
dent margin, and a slight indentation to receive the apex of the
upper latus.
The carina is strongly arched, 13.5 mm long and 2.8 mm wide
near the base, with an acute apical umbo which intrudes between
the terga. The roof is convex, the sides bicostate and very narrow,
the basal margin rounded. The upper latus is pentagonal, the scutal
margin the longest, and the tergal, carinal, and basal margins suc-
cessively shorter, the margin against the inframedian plate the
shortest and less than half the basal margin. The rostrum is small and
triangular. The rostral latus has two low ridges running from the
apex to the upper and lower angles of the plate. The inframedian
latus is narrow and triangular, with the apex strongly curved toward
the occludent border and overlying the baso-lateral angle of the
scutum near the baso-lateral angle of the inframedian latus. A low
rounded rib runs down each side. The umbo is apical. The carinal
latus is subtriangular, its apical half projecting free behind and down-
ward below the carina. The spread from tip to tip is 3.75 mm in the
type specimen.
The peduncle is 7 mm in length, clothed with rather large, nar-
row, transversely lengthened scales, in about eight rows of eight or
nine scales each.
According to Pilsbry, Scalpellum diceratum lives with and is re-
lated to Scalpellum arietinum Pilsbry, yet is distinct by the dif-
ferences in the shape of the carina and inframedian latus, and in the
terminal umbones of the inframedian, upper latera, and carina of
the two species.
Type locahty. — “Albatross” sta. 2319 (23°10'37”N,
82°20’06” W ), 143 fathoms, off Habana, Cuba, on crinoid arms.
Florida localities. — “Albatross” sta. 2405, (23°45’N, 85°02’W),
off West Florida in the Gulf of Mexico, 30 fathoms, about 267
statute miles west of Homosassa; off Palm Beach, on gorgonians,
hydroids, and echinoid spines, 30 to 80 fathoms; off Sombrero Key
Light; “Triton” sta. 1952, off Cape Florida, 100 fathoms (183
meters), University of Miami collection.
Other localities. — “Albatross” sta. 2324 (23°10'25”N,
246 BULLETIN 299
82°20’24”W), off Habana, Cuba, 33 fathoms, on Cidaris spine;
“Albatross” sta. 2317, Straits of Florida (24°25’45”N, 81°46’W),
45 fathoms, temperature 75°F, on spines of Cidaris; “Albatross”
sta. 2315 (24°26’N, 81°48’15”W), Straits of Florida, 37 fathoms
(68 meters); continental shelf off North Carolina, near Cape Hat-
teras, south of Diamond Shoals (34°56’N, 75°26’W), 46 fathoms
(84 meters); “Pillsbury” sta. P-372, in Caribbean Sea about 40
kilometers northwest of Covemias (9°24’N, 75°44’W), Colombia, 82-
100 meters (151-186 fathoms).
Scalpellum (?) diceratum Pilsbry is a relatively shallow-water
species ranging in depth from 30 to 186 fathoms. Geographically it
ranges from North Carolina in the north to Colombia in the south.
Remarks. — Although Scalpellum (?) diceratum Pilsbry lives
with Scalpellum arietinum Pilsbry the two are not related, the
former lacking the characters pertaining to the genus Scalpellum
sensu stricto.
Scalpellum gibbum Pilsbry Pl. 27, fig. 7; PL eee
Scalpellum gibbum Pilsbry, 1907, pp. 14, 17-18, figs. 4a, b; 1953, p. 19, text-
fig. 2; Henry, 1954, p. 444; Ross, Cerame-Vivas, and McCloskey, 1964, p. 312;
Ross, 1965, pp. 219-220, figs. 1A, B; Zullo, 1966, pp. 230, 231-232, figs. 2A, B;
eo 212; Cerame-Vivas and Gray, 1966, p. 263; Newman and Ross, 1971,
The capitulum of the type is subtetragonal, 7 mm in length
and 4 mm in breadth, with a slightly sinuous ventral margin and
an angularly bent carinal margin. It is composed of 14 fully calcified
plates which are faintly marked by growth lines and separated by
narrow chitinous sutures. The tergum is much longer than the
scutum, obtusely and narrowly triangular in shape, its occludent
margin slightly convex and strongly recurved at the summit; the
carinal margin is biconcave, short above, longer and gently curved
below. The scutum is twice as long as wide, its umbo at the upper
third of the occludent margin; the tergal margin is straight and
oblique; the lateral margin is angular, the basal margin slightly con-
cave. The carina is prominently angular near the middle. The roof
is convex, bounded by two lateral ribs, accompanied below by a
second arcuate rib on each side. The sides are wide, flat, and marked
by four or five wrinkles parallel with the growth lines.
The upper latus is rhomboidal, the umbo lying near the scutal
margin about midway between the basal and tergal borders. The
FLORIDIAN CIRRIPEDIA: WEISBORD 247
rostrum is narrow and parallel-sided, with the beaks of the rostral
latera meeting over it above the middle. The rostral latus is twice
as long as high, its umbo acute, the upper and basal margins parallel,
the lateral margin straight. The inframedian latus is convex, penta-
gonal and much larger than the other lower plates and fully equal to
the upper latus; the umbo is nearly central. The carinal latus is
claw-shaped, the umbo projecting below the carina; the basal and
lateral margins are about equal, the upper margin very short, the
carinal margin concave, with a low submarginal rib.
The peduncle is 2 mm in length. It is covered with large im-
bricating scales, in about 10 rows.
Type locality. —“Albatross” sta. 2388 (29°24’30”N, 88°01’W),
35 fathoms (64 meters), in Gulf of Mexico about 100 statute miles
south of Mobile, Alabama, and 40 statute miles southeast off the
forward edge of the delta of the Mississippi River.
Florida locality. — “Triton” sta. 441, off Palm Beach (26°4V/N,
80°02’W), 30 fathoms (55 meters).
Other localities. —South Carolina, “Miss Kim” sta. 12
(32°28.7'N, 78°47.1’W), 46% miles off Racoon Key, depth 64
meters, hard sand and shell bottom. North Carolina, near Cape
Hatteras, south of Diamond Shoals (35°01’N, 75°25’W), 30 fathoms;
off Cape Lookout (34°11’N, 76°08’W), 50 fathoms.
Fossil locality. — Ecphora Zone, Jackson Bluff Formation, Leon
County, Florida. Late Miocene—early Pliocene.
The geologic range of this species is Mio-Pliocene to Recent.
The geographic range is from North Carolina to Florida. The bathy-
metric range is 30-50 fathoms (55-91 meters).
Scalpellum (?) giganteum Gruvel Rig 28. ties 2.3
Scalpellum giganteum Gruvel, 1901b, pp. 153-156, pl. 17, figs. 1-8, 17; 1902a,
pot 1905, pp. 78-79) fig, 88; Pilsbry. 1907, pp. 25, 32-33, pl. 2, fig. 1, pl. 3,
fig. 1; Annandale, 1909-1910, p. 132; Bayer, Voss, and Robins, 1970, p. A43.
Gruvel’s original description is summarized as follows:
The capitulum of the type is flattish, 45 mm long and 32 mm
in breadth, and is composed of 14 calcified plates nearly completely
covered with a thick chitinous cuticle. In shape the capitulum is a
curvilinear, nearly isosceles triangle, the sides convex and the base
slightly concave. The plates are sculptured by prominent growth
striae which are clearly vestiges from the cuticle. The tergum 1s ir-
248 BULLETIN 299
regularly quadrangular. The scutum is nearly triangular, the lower
margin measuring not quite half the length of the plate. The carina
is regularly arched; the umbo is at the apex and does not project
between the terga; the dorsal margin is convex as are the lateral
margins; the basal margin is also strongly convex, the lower angle
blunt, not reaching the summit of the carino-lateral plates; the
roof of the carina enlarges gradually from the summit to the base.
The upper latus is irregularly quadrilateral. The rostrum is
small and oval, the lateral margins hidden, and the remainder en-
tirely masked by the cuticle. The rostral latus is elongated, narrow,
larger anteriorly than posteriorly. The inframedian latus is triangu-
lar, the lower margin longer than the posterior and the anterior;
the apex is directed toward the summit of the capitulum. The carino-
lateral latus is elongated, narrow, and with a backward slope; the
apex is strongly recurved upward and in front, and does not reach
the lower or external margin of the carina; the umbo is nearly at
the base.
The peduncle, which is more or less cylindrical, is 45 mm in
length and 15 mm in diameter. It consists of six longitudinal and
alternating rows of scales, with 10 to 12 transversely elongated
scales per row.
Type locality. — Coasts of Cuba, depth 500 fathoms.
Florida locality. — “Albatross” sta. 2658 (28°21’N, 78°37’W),
514 fathoms (940 meters), about 120 miles east off Cape Canaveral.
With Megalasma (Glyptelasma) gracilius Pilsbry.
Other localities. — “Albatross” sta. 2554 (39°48’30’N,
70°41’W), 455 fathoms (832 meters), on Scalpellum velutinum
Hoek, about 120 statute miles east off Surf City, New Jersey; West
Indies, on Atlantic cable, from capitulum of Scalpellum velutinum
Hoek; “Pillsbury” sta. 338 (9°58.3’N, 78°30.5’W ), 1836-1822 meters,
about 80 kilometers northwest off Punta San Blas, Panama, in the
Caribbean Sea; “Pillsbury” sta. 364 (9°28.7’N, 76°34.3’W to
9°20.2’N, 76°34.2’W), 933-961 meters, about 90 kilometers north-
west off Covefias, Colombia, in the Caribbean Sea; “Pillsbury” sta.
407 (9°02’N, 77°25.3’W to 9°02’N, 77°28.8’W), 1171-1239 meters,
about 40 kilometers east of Punta Mosquito, Panama, in the Carib-
bean Sea.
The recorded depths of S. (?) gigantewm range from 832 to
FLORIDIAN CIRRIPEDIA: WEISBORD 249
1822 meters. The known geographic range is from off Colombia,
South America, to off New Jersey, U.S.A., although Pilsbry (1907,
p. 33, pl. 3, fig. 1) mentioned the possibility of a large specimen
having been obtained on the “fishing banks” (off Newfoundland? ).
Scalpellum (?) gorgoniophilum Pilsbry Pl. 28, figs. 4a, b
Scalpellum gorgoniophilum Pilsbry, 1907, pp. 25, 33-34, fig. 7a, b; Zullo,
1968, p. 213.
Pilsbry’s description is summarized as follows:
The capitulum is subrectangular for about two-thirds its length,
triangular apically; the type is 9 mm in length, 5 mm in breadth.
The plates are lacking in hair and have no noticeable cuticle but
are sculptured by coarse growth lines and fine radial striae, with a
strong diagonal rib on the scutum and upper latus. The tergum is
triangular with gently convex occludent and basal margins. The
scutum is large, subtetragonal with parallel lateral margins, a slightly
concave tergal margin, and a slightly sinuous basal margin. The
carina is relatively short, strongly arched above, hardly convex
below, the apex reaching only to the middle of the carinal side of
the tergum. The length is 6.8 mm, diameter near the base 1.8 mm.
The roof is convex and radially striate with narrow ribs separating
it from the sides; the sides are wide and bear a sharply elevated
arcuate rib. The umbo 1s apical.
The upper latus is subtriangular, the lower margins conforming
with the margins of the adjoining plates. The carinal latus is penta-
gonal, curved like a scoop, with the apex projecting outward beyond
the carina. The inframedian latus is narrow, obliquely triangular,
tapering to the apex which curves toward the scutum and overlies
its baso-lateral angle; the umbo is apical. Three unequal faces abut
the upper and inframedian latera and the peduncle. Behind the
carina the two latera meet only at the base. The rostrum is com-
paratively large, in the shape of an isosceles triangle. The rostral
latus is low and wide, its surface divided by a diagonal rib.
The peduncle is stout and short, measuring about 2.8 mm in
length; it is covered with projecting scales in about eight deeply
interlocking rows of six or seven scales each. The peduncle is in-
conspicuously hairy.
The large size of the rostrum, the short carina, and the pro-
jecting apices beyond the carina of the carinal latera are the con-
spicuous characters of this species.
250 BULLETIN 299
Type locality. — “Albatross” sta. 2338 (23°10’40’N,
82°20'15” W ), off Habana, Cuba in 189 fathoms (346 meters), coral
bottom, on a gorgonian. The type locality lies in the Caribbean Sea
about 110 statute miles southwest of Key West, Florida.
So far as I have been able to determine, the type locality is the
only one recorded for this unique species. It is included in this work
because of the probability it eventually will be found in Florida
waters.
Scalpellum (?) gracilius Pilsbry Pl. 28, figs. 5a-c
Scalpellum gracilius Pilsbry, 1907, pp. 47, 51-53, figs. 17a-c; 1911, p. 173;
Weltner, 1922, pp. 96, 106; Zullo, 1968, p. 213.
Following is a resumé of Pilsbry’s original description:
The capitulum is oval, length 8 mm, breadth 3.3 mm, consisting
of 14 fully calcified plates covered with a thin smooth cuticle, and
separated by linear sutures. The plates have faint growth lines and
a few barely perceptible radial striae.
The tergum is larger than the scutum, triangular in outline,
with a slightly convex occludent margin, a straight basal margin,
and a weakly sigmoidal carinal margin; the apex is erect. The scu-
tum is longer than wide, the occludent margin convex above, slightly
concave near the base, the lateral margin slightly sinuous, the basal
margin convex passing into the lateral in a smooth curve; the apex
is a little incurved and acuminate. The carina, measuring 6.3 mm
in length and 1 mm at the base, is regularly curved, the apex
terminal. The roof is rounded, curving into wide sides; the basal
margin is convex. The growth lines of the roof curve deeply down-
ward (PI. 28, fig. 5c).
The upper latus is irregularly pentagonal, the margin against
the carinal ]atus concave, the apex subterminal. The rostrum is re-
duced to a linear rudiment separating the rostral latera along the
upper half of their contiguous borders. The rostral latus is sub-
triangular. The inframedian latus is narrowly oblong, contracted
slightly below the middle, the basal segment much smaller than the
upper. The carinal latus is long and narrow, the occludent margin
convex at the border of the upper latus, sinuous against the margin
of the inframedian latus; the umbo is close to but not at the base
of the plate and does not project beyond it. The two plates meet
in a short straight suture below the carina.
FLORIDIAN CIRRIPEDIA: WEISBORD 251
The peduncle is 1.8 mm in length, closely covered with large
transversely lengthened scales in six rows of about five scales each.
Type locality. — “Albatross” sta. 2678 (32°40’N, 76°40/30”W),
about 185 statute miles east off Folly Beach, South Carolina, 731
fathoms (1337 meters), bottom temperature 38.7°F.
Other localities. —“Albatross” sta. 2751 (16°54’N, 63°12’W),
687 fathoms, bottom temperature 40°F., blue Globigerina ooze,
about 20 statute miles southwest off Charlestown, Nevis, Lesser
Antilles.
Inasmuch as Florida lies between South Carolina and Nevis it
is anticipated that Scalpellum gracilius eventually will be discovered
also in Florida waters.
Scalpellum hendersoni Pilsbry Pl. 28, fig. 6
Scalpellum hendersoni Pilsbry 1911, pp. 172-173, fig. 1; Zullo, 1968, p. 213.
Pilsbry’s original description of the type is summarized as
follows:
The capitulum is subquadrate and swollen except for the upper
end which is compressed and triangular. The length is 5 mm, the
breadth 2.5 mm. The carinal margin is arched, the occludent margin
convex. The plates are fully calcified, with widely spaced growth
lines and, on the tergum, scutum, and upper latus, a few weak radial
striae. The tergum is triangular. The scutum is trapezoidal. The
carina, measuring 3.75 mm in length is arcuate, more so in upper
third where there is a space between the apical area and the margin
of the tergum. The roof of the carina is strongly convex, widening
rapidly toward the base which wedges between the carinal latera.
The intraparietes are narrow, bounded by a ridge, and visible only
in the upper part of the plate.
The upper latus is trapezoidal, with an apical umbo. The rostral
latera are triangular, obtuse at the rostral angle. There is no visible
rostrum. The inframedian plate is narrow and high, contracting per-
ceptibly at the lower fourth where the umbo is situated. The carinab
latera are large and irregular, with the umbo at the lower carinal
angle; the two latera meet in a short suture below the carina.
The peduncle is short and is covered with large scales in about
seven vertical rows.
Type locality. — Ten miles south of Key West, Florida, in 125
fathoms, on spines of a sea urchin, Dorocidarts, associated with the
barnacle Verruca alba Pilsbry.
252 BULLETIN 299
Scalpellum idioplax Pilsbry Pl. 29, figs. 2a-c
Scalpellum idioplax Pilsbry, 1907, pp. +7, 48-50, figs. 15a-c; Broch, 1924,
pp. 41, 45, 102; 1953, p. 5; Zullo, 1968, p. 213; Bayer, Voss, and Robins, 1970,
p. A43.
Pilsbry’s description is summarized as follows:
The capitulum is twice as long as wide (18 mm X 9 mm), con-
vex at the ventral and dorsal margins, subtruncate at the base, and
triangular at the apex. The cuticle is very thin and smooth. There
are 13 fully calcified plates, sculptured with unequal lines of growth
and fine, low radial striae.
The tergum is longer than the scutum, triangular, the occludent
margin slightly convex, the basal margin slightly concave centrally,
and the carinal margin weakly sigmoid. The scutum is longer than
wide, the lateral margin irregular, projecting in an angular lobe at
the upper lateral angle and deeply excavated below the lobe for the
reception of the apex of the upper latus. The apex is acuminate.
The carina, measuring 15 mm in length and 3 mm in width at the
base, is arched, more so near the terminal mucro than below. The
roof is flat, with bordering ribs. The growth striae of the roof are
convex upward. The sides are wide, regularly tapering toward the
base. The basal margin is slightly concave.
The upper latus is hexagonal-pyriform; the tergal and scutal
margins are long, the former hardly convex, the latter concave; the
carinal margin is short and straight, that against the carinal latus
also short. The carinal latus is twice as high as wide, irregularly
triangular, the occludent margin concave in the middle. The umbo
projects slightly beyond the carina near the base of the plate; the
carinal latera meet below the keel. The inframedian latus is com-
posed of a large upper segment and a small basal segment, the junc-
tion narrow. The rostral latus is squarish with straight margins, the
lower lateral corner rounded, and the ventral margins of the rostral
latera in contact.
The peduncle is 4.3 mm in length, with 10 rows of transversely
lengthened scales, about eight scales in a row.
Type locality. — “Albatross” sta. 2140 (17°36'10’N,
76°46'05”W), Caribbean Sea between Jamaica and Haiti, 966
fathoms (1767 meters), sand bottom.
Florida locality —“Albatross” sta. 2656 (27°58’30’N, 78°24’W),
FLORIDIAN CIRRIPEDIA: WEISBORD 253
572 fathoms (1046 meters), bottom temperature 41.2°F, about 135
statute miles east off Melbourne Beach.
Other localities. —“Pillsbury” sta. 338 (9°57.5’N, 78°31’W to
9°58.3’N, 78°30.5’W), 1836-1822 meters, Caribbean Sea, about 80
kilometers northeast off Punta San Blas, Panama; “Pillsbury” sta.
388 (10°16’N, 76°03’W to 10°10’N, 76°08’W) 824-1061 meters,
Caribbean Sea, about 70 kilometers southwest of Cartagena, Colom-
bia; “Pillsbury” sta. 407 (9°2’N, 77°25.3’W to 9°2’N, 77°28.8’W),
1171-1239 meters, Caribbean Sea, about 40 kilometers east off
Punta Mosquito, Panama.
Range and distribution. — The species has been reported from
off Melbourne, Florida, in the Western Atlantic to as far south as
northern Colombia, in depths ranging from 824 meters in the Carib-
bean to 1836 meters off Punta San Blas, Panama, in the Caribbean.
Scalpellum latidorsum (Pilsbry) Pi oleetieaG
Scalpellum regium latidorsum Pilsbry, 1907, pp. 25, 29-31, pl. 2, figs. 2, 3,
7, pl. IV, figs. 10, 11, 12, 14; Fowler, 1912, p. 500; Zullo, 1968, p. 214.
The capitulum of Scalpellum regium latidorsum varies from 48
to 60 mm in length and 31 to 38 mm in breadth. It is high domal
in outline, with a pointed apex, a moderately convex occludent
border, a more convex carinal border, and a slightly concave base.
The 14 plates, which abut or are close to one another, are covered
with a thin cuticle bearing few hairs, and are sculptured in the upper
whorls by a series of fine growth lines interspersed at intervals with
stronger growth lines. The tergum is pentagonal in outline, with
steep unequal apical margins and a pointed basal angle. The scutum
has longer apical margins than the tergum and the basal angle is
subrounded. The carina measures 43 mm in length and 9.5 mm in
width near the base, and has a flat roof with discrete, widely spaced,
V-shaped markings; laterally there are low ribs, and the sides widen
gradually toward the base.
The upper latus is subtriangular, with an acute slightly curved
apex, a convex tergal margin, a concave scutal margin, and a sub-
angular base. The rostrum is narrow and indistinctly visible through
the cuticle. The rostral latus is elongate, with subparallel scutal and
basal margins and a sharply convex occludent margin. The infra-
median latus is small and triangular, and there is a rooflike chitinous
254 BuLLeTIN 299
extension above its apex. The carinal latus has a long, strongly re-
curved umbo and a horn-shaped basal extension sculptured by fair-
ly numerous sinuous concentric striae; above the umbo and the upper
margin of the latus there is a chitinous extension of the plate.
The peduncle varies in length from 21 to 36 mm, and has large,
transversely lengthened scales thinned at their ends. There are seven
rows of about nine scales each.
Type locality. — “Albatross” sta. 222 (39°03’15”N,
70°50’45”W), 1537 fathoms, gray ooze, surface temperature 73°F,
bottom temperature 36.9°F, about 210 miles east off Cape May,
New Jersey.
Other localities —“Albatross” sta. 2042 (39°33’N, 68°2645”W),
1555 fathoms (2844 meters), Globigerina ooze, surface temperature
71°F, bottom temperature 38.5°F, about 310 statute miles east off
Atlantic City, New Jersey; “Albatross” sta. 2041 (39°22’50”N,
68°25’W), 1608 fathoms (3028 meters), Globigerina ooze, surface
temperature 72°F, bottom temperature 38°F, about 325 statute
miles east off Ocean City, New Jersey; “Albatross” sta. 2210
(39°37'45”"N, 71°18’45”W), 991 fathoms (1813 meters), surface
temperature 74°F, bottom temperature 38.1°F, Globigerina ooze,
about 170 statute miles east off Ocean City, New Jersey.
Inasmuch as the carina of this taxon is different from that of
Scalpellum regium regium Thomson (Pilsbry, 1907, pl. 2) and that
S. regiwm itself resembles other species to which the present taxon
might be allied, it is suggested that Pilsbry’s subspecies latidorsum
be given specific rank, that is Scalpellwm latidorsum (Pilsbry) to
replace Scalpellum regium latidorsum Pilsbry.
Scalpellum (?) longicarinatum Pilsbry Pl. 29, figs. 3a-c
Scalpellum longicarinatum Pilsbry, 1907, pp. 26, 37-39, figs. 9a-c; Broch,
1924, p. 39; 1953, pp. 4-5, 10, 12, 15, figs. la-d; Zullo, 1968, p. 214.
Pilsbry’s description is summarized as follows:
This is a strong, robust little species. The capitulum, which is
10 mm in length and 5.4 mm in breadth, is long-oval, widest in the
middle, with convex lateral margins, the ventral border less curved
than the dorsal. It is composed of 14 fully calcified plates marked
by emphatic concentric growth striae with prominent grooves at
intervals between them.
The tergum is obliquely elongated, a little longer than the scu-
FLoRIDIAN CIRRIPEDIA: WEISBORD 255
tum; the occludent and basal margins are slightly convex, the carinal
margin slightly concave near the summit but convex below. The
scutum is trapezoidal, the occludent margin convex, the apex
acuminate and recurved, the tergal margin concave, the carinal
margin convex, the basal margin nearly straight. The carina, 9.5
mm in length and 2 mm wide at the base, is evenly arched, with the
umbo apical. The roof is flat between strong bordering ribs. The
sides are wide and sulcate with deep growth lines. The basal margin
is convex.
The upper latus is quadrangular, the scutal and carinal margins
parallel, the carinal about half as long as the scutal; the umbo is
apical. The rostrum is small and triangular. The rostral latus is
trapezoidal with parallel upper and lower margins. The inframedian
latus is narrowly triangular, the base half the height; the umbo is
apical, and there is an inconspicuous triangular wing at the carinal
side of the apex. The carinal latus is irregularly pentagonal, as wide
as high, the carinal margin deeply concave; the umbones project
a little beyond the carina and are somewhat recurved. The portions
of the carinal latera seen in dorsal view are obliquely triangular, the
roof of the carina wedging narrowly between them to the peduncle.
The peduncle is 2 mm in length, closely covered with transverse-
ly lengthened scales, in about 18 rows of seven or eight scales each.
Type locality. — “Albatross” sta. 2668 (30°58’30’N, 79°38’W),
294 fathoms (538 meters), about 105 statute miles east off St.
Andrews Sound, Georgia.
Other localities. —‘“Albatross” sta. 2415 (30°44’N, 79°26’W),
440 fathoms (805 meters), about 120 statute miles east off mouth
of St. Mary’s River, between Georgia and Florida; “Albatross” sta.
2663 (29°39'N, 79°49°W), 421 fathoms (770 meters), about 63
statute miles east off Marineland, Florida; “Ingolf” sta. 92 (64°44’N,
32°52’W), between Iceland and Greenland, depth 1838 meters, bot-
tom temperature 1.4°C.
Scalpellum longicarinatum occurs in the Western Atlantic off
the east coast of Georgia and Florida, and was reported by Broch
(1953) in the North Atlantic between Iceland and Greenland.
Depths range from 294 to 1006 fathoms (538 to 1838 meters), the
shallowest off Georgia, U.S.A., the deepest west of Iceland.
Scalpellum (?) microceros MacDonald Pl. 29; tigot
Scalpellum microceros MacDonald, 1929, pp. 531-532, pl. 2, fig. 1.
BuLLeTIN 299
bo
Lea
nN
MacDonald’s description of the type and only specimen is sum-
marized as follows:
The capitulum is trapezoidal, 31 mm in length and 23 mm in
width. There are 14 well-calcified valves in close contact, covered
with a thin cuticle and sculptured by fine growth striae. The tergum
is longer than the scutum and somewhat lanceolate; the occludent
margin is straight, the carinal and scutal margins convex. The
scutum is strongly convex and twice as long as broad; the occludent
and lateral margins are subparallel and the umbo is apical. The
carina is well arched, with an acute apical umbo wedged between the
terga. The roof is flat and bordered with prominent ridges. The
sides are broad and of equal breadth throughout their length, and
the basal margin of the carina is almost straight.
The upper latus is pentagonal, the carinal and scutal margins
almost parallel, with two parallel shallow grooves along the scutal
margin; the lower half of the scutal margin overlaps the scutum, and
there is a low ridge running along the tergal margin. The carinal latus
is triangular, the apical half projecting upward and considerably
beyond the carina. The roof of the carina extends between the carinal
latera to the peduncle. The rostrum is small and triangular, and
overlaps the apices of the rostral latera. The rostral latus is linear,
about seven times as long as broad, with a shallow groove running
the length of the surface; the latus stands out prominently above
the surface of the scutum. The inframedian latus is small and tri-
angular, with the apex curved toward the occludent border. The
umbo is apical, and there is a low ridge along each side.
The peduncle is equal in length to the capitulum and is covered
with imbricated scales in about 28 rows.
Type locality. —MCZ collection (13°52’N, 61°7’W), just off
the west coast of St. Lucia, 278 fathoms (508 meters). St. Lucia
Island lies 1,250 statute miles southeast off Florida’s east coast.
Scalpellum (?) micrum Pilsbry Pl. 29, fig. 4
Scalpellum micrum Pilsbry, 1907, pp. 47, 57-58, fig. 21; Barnard, 1924, pp.
17, 46-47; Zullo, 1968, p. 24.
Pilsbry’s description of this species is as follows:
The capitulum, measuring 5 mm in length and 2.5 mm in
breadth, is oval, with the ventral and dorsal margins about equally
FLORIDIAN CIRRIPEDIA: WEISBORD 257
convex. There are 14 fully calcified plates separated by linear
sutures, the plates marked with faint lines of growth and a few
faint radial striae. There is no perceptible cuticle.
The tergum is triangular, the occludent and the scuto-lateral
margins convex, and the carinal margin sinuous, concave above,
somewhat convex below. The scutum is about twice as long as wide,
the occludent and lateral margins subparallel, the basal margin
nearly at right angles to them, the upper third of the occludent
margin bent backwards; the umbo is acute, terminal and recurved.
The carina is 3.2 mm in length, simply arched, with an apical mucro.
The roof is rounded, marked with transverse, arcuate lines of growth,
the sides narrow; the apex reaches to the upper third of the carinal
margin of the tergum; the base of the carina is rounded.
The upper latus is trapezoidal with straight margins, the apex
terminal at the scuto-tergal angle. The rostrum is well developed,
forming a band about one-fifth as wide as long, and slightly narrower
above the base; it extends the whole length of the adjacent latera.
The rostral latus is triangular, with the basal angle of the triangle
truncated. The inframedian latus is narrow and triangular, its
height equal to that of the rostral latus and about double the basal
width; the umbo is apical. The carinal latus is irregularly pentagonal,
with the upper lateral and carinal margins about equal and straight,
the subcarinal margin the longest and concave, the basal margin the
shortest; the umbo of the carinal latus projects angularly beyond
the carina. The two carinal latera meet below the carina in a straight
suture as far up as their umbones.
The peduncle is but 1.3 mm in length; it is covered with large
imbricating scales in five rows of five scales each.
Type locality. — “Albatross” sta. 2668 (30°58’30’N,
79°38’30”W ), 290 fathoms (530 meters), on a delicate hydroid, bot-
tom temperature 46.3°F, the bottom of gray sand with dead coral.
“Albatross” sta. 2668 is in the Western Atlantic about 105 statute
miles east off St. Andrews Sound, Georgia.
Other localities. — “Pieter Faure” sta. about 20 miles southeast
off East London (33°S, 27°54’E), South Africa, 400-450 fathoms
(732-823 meters).
Scalpellum (?) pentacrinarum Pilsbry Pl. 30, figs. la-c
Scalpellum pentacrinarum Pilsbry, 1907, pp. 47, 55-57, figs. 20a-c; Gruvel,
1909, p. 208; Zullo, 1968, p. 215; Newman and Ross, 1971, p. 51.
258 BULLETIN 299
Pilsbry’s description of the type is summarized as follows:
The capitulum of this small species is 8 mm in length and 3.7
mm in breadth. It is subtriangular in shape and is composed of 13
fully calcified plates, separated by linear sutures, and without per-
ceptible cuticle. The plates are marked with fine lines of growth,
and the scutum, tergum, and upper latus are marked by low radial
striae.
The tergum is triangular, with an erect apex. The scutum is
long and narrow, widest at the base where it is three-fourths the
width of the capitulum at that plate. The lateral margins of the
scutum are subparallel, converging slightly above, the occludent
margin hardly convex, the carinal margin straight. The straight basal
margin makes a right angle with the occludent margin.. A low nar-
row ridgelet runs from the acute apex of the scutum to the baso-
lateral margin. The carina is irregularly arched and unusually short,
measuring 5.2 mm in length and 1.2 mm in width near the base.
The roof is rounded, passing directly into the narrow sides, and is
marked with faint transverse arcuate growth lines. The base is
wedged triangularly between the carinal latera. The apex is terminal
and incurved but not inserted between the terga; it reaches only to
the lower fourth of the margin of the tergum.
The upper latus is wedge-shaped, with straight scutal and tergal
margins and a slightly convex carinal margin; the umbo is terminal
at the scuto-lateral angle. There is no rostrum but a lanceolate space
between the rostral latera. The rostral latus is quadrangular, at least
twice as wide as high, and is divided by a low diagonal riblet into
two unequal triangular parts; the rostral margin is concave, the
lateral somewhat irregular. The inframedian latus is narrow, sinuous,
and as high as the adjacent latera, its umbo at the acute apex. The
carinal latus is triangular, higher than wide, the apex curved toward
the inframedian latus. The two latera almost meet at the base below
the carina. The almost concrescent inframedian and carinal latera
are a distinguishing character of the species.
The peduncle, 3.7 mm in length, is covered with wide imbri-
cating scales in six rows of about 15 scales each. The scales of ad-
jacent rows interlock only a little.
Type locality. —“Albatross” stations 2319-2350 (23°10/37’N,
82°20’06’” W to 23°10’39’N, 82°20’21”W), off Habana, Cuba, on a
FLORIDIAN CIRRIPEDIA: WEISBORD 259
pinnule of Pentacrinus. Depths within the relatively small area en-
compassed by “Albatross” stations 2319 to 2350 vary from 33
fathoms (60 meters) to 279 fathoms (510 meters), the bottoms
mostly of coral with rare sand, and the recorded bottom tempera-
tures 58° to 79.1°F, the latter at the 33 fathom depth.
The Habana type locality lies about 110 statute miles south-
west of Key West, Florida.
Scalpellum (?) portoricanum Pilsbry Pl. 29, figs. 5a-c
Scalpellum (species?), Bigelow, 1901, p. 179. [Fide Pilsbry, 1907, p. 35.]
Scalpellum portoricanum Pilsbry, 1907, pp. 26, 35-36, figs. 8a-c; 1953, p. 19;
U.S. Naval Inst., 1967, p. 194; Broch, 1953, pp. 4, 9, 10; Zullo, 1968, p. 215;
Bayer, Voss, and Robins, 1970, p. A43.
Scalpellum (Scalpellum) portoricanum Pilsbry, Calman, 1918a, pp. 121-122.
Pilsbry’s description of the type and only specimen is sum-
marized as follows:
The capitulum, measuring 12 mm in length and 7.7 mm in
breadth, is rhombic-oblong, with a nearly straight occludent margin
and a more convex carinal margin. It is composed of 14 wholly
calcified plates which are covered with a thin and sparsely pilose
cuticle, and are marked weakly with lines of growth.
The tergum is larger than the scutum and has slightly convex
basal and carinal margins, straight occludent and lateral margins,
the latter shorter, and an erect apex. The scutum is trapezoidal,
more than twice as long as wide, the occludent and lateral margins
subparallel, the basal margin straight, and the umbo apical, not
projecting beyond the occludent outline. The carina, 11.3 mm long
and 2.2 mm wide at the base, is gently arched, its umbo apical,
against but hardly between the terga. The roof is flat, bounded by
low lateral ribs, and faintly marked with arcuate growth lines; the
sides are narrow and concave, the basal margin a little concave.
The upper latus is pentagonal, with a superior, terminal, and
acute apex. The rostrum is small and triangular. The rostral lJatus
is narrow, the scutal and basal margins more or less parallel. The
inframedian latus is triangular, the apex curving ventrad around the
end of the rostral latus and between the lower angles of the upper
latus and scutum. The carinal latus is irregularly triangular and
projects backward beyond the carina; the umbo is recurved and
flares outward in the shape of a subspiral horn.
260 BULLETIN 299
The peduncle is 7 mm in length, with about 13 rows of trans-
versely lengthened scales, about six scales in a row.
Type locality. — “Fish Hawk” sta. in Mayagiiez Harbor, Puerto
Rico (18°13’N, 67°09’W), depth between 25 and 76 fathoms (46-
129 meters).
Florida localities. — Off Palm Beach, 75 fathoms (137 meters),
at several “Triton” stations.
Other localities. —“Pillsbury” sta. 340 (9°13.5’N, 77°46’W),
307-366 meters, about 40 kilometers northeast of Sasardi Viejo,
Panama, Gulf of Darien; “Pillsbury” sta. 445 (9°02.3’N, 81°23.8’W),
342-346 meters, about 70 kilometers west of Belén, Gulf of Mosqui-
tos, Panama; C/S “Henry Holmes” sta. at 18°31’N, 66°19 W, 180
fathoms (329 meters), about 30 kilometers west-northwest of San
Juan, Puerto Rico.
Scalpellum (?) intonsum (Pilsbry) Pl. 29, figs. 6d, e
Scalpellum portoricanum intonsum Pilsbry, 1907, pp. 25, 36-37, figs. 8d, e;
Henry, 1954, p. 444; Zullo, 1968, p. 214.
Pilsbry described this taxon from three individuals recovered
with a large beam trawl in the Gulf of Mexico at Albatross station
2401. Concerning it he wrote:
They [the specimens] are smaller than the Porto Rican type, rather densely
hairy, and differ from typical S. portoricanum somewhat in shape. The capi-
tula measure 9.5, 9.7, and 7 mm. long. The occludent margin of the scutum
is distinctly convex, that of the tergum straight or even a trifle concave. The
summit is erect, not recurved. The inframedian latus is longer and narrower
than in S. portoricanum. The rostrum is narrower. The umbones of the carinal
latera project less and are situated higher. The valves are sculptured with
concentric grooves at subequal intervals. The two larger examples are evi-
dently adult. (Fig. 8d, e).
Type locahty. — “Albatross” sta. 2401 (28°38’30’N,
85°52’30”W), 142 fathoms (260 meters), Gulf of Mexico, about 85
statute miles west of Bayport, Florida, and about 108 statute miles
south of St. Andrews, Florida. The bottom is green mud and broken
shells.
Because the subspecies intonswm seems to me to be distinguish-
able from S. portoricanum s.s., and because S. portoricanum resem-
bles somewhat a number of other species, I am inclined to raise the
rank of intonswm to species.
Scalpellum pressum Pilsbry Pl. 30, figs. 2a, b
Scalpellum pressum Pilsbry, 1907, pp. 14, 23-24, figs. 6a, b; Broch, 1924,
pp. 22, 28, 29, 30 [= S. stroemii M. Sars, fide Broch, 1924, p. 28.]; Zullo, 1968,
p. 215 [= S. stroemti Sars, fide Zullo, p. 215.]
FLORIDIAN CiRRIPEDIA: WEISBORD 261
Pilsbry’s original description of S. presswm is summarized as
follows:
The capitulum, measuring 8 mm in length, 4 mm in breadth
and 1.8 mm in thickness, is compressed, is widest above the middle,
and tapers toward the base which is obliquely truncated. There are
14 plates, irregularly marked with concentric wrinkles and a few
weak radial striae. The occludent border is convex above, much less
so below.
The tergum has a convex occludent margin, a recurved apex,
and a carinal margin which is a little concave below the apex and
nearly straight where it is in contact with the carina. The scutum
is trapezoidal, with a slightly convex occludent margin which is
subparallel with the short lateral margin, a slightly concave tergal
margin, and a straight basal margin. The carina is moderately arched,
with the umbo projecting a little near the apex. The roof is convex
and marked by a few faint longitudinal striae. The parietes are
narrow, the intraparietes a little wider.
The upper latus is pentagonal, the umbo not quite terminal.
The rostrum is long, narrowly wedge-shaped, blunted at the pro-
jecting apex. The rostral latus is broadly triangular. The inframedian
latus is more than twice as high as wide, with the umbo on the rostral
margin below the middle. The scutal margin is slightly shorter than
that against the upper latus. The carinal latus is about twice as wide
as high, its umbo projecting slightly behind the base of the carina;
below it there is a nearly straight margin almost as long as the basal
margin and about one-third the length of the plate; the upper
margin is oblique. The dorsal margins of the two carinal latera
meet below the umbones in a straight suture.
The peduncle is about one-third to one-half the length of the
capitulum or about 3 mm. It is covered with eight rows of large im-
bricating scales, eight to ten scales in a row.
Type locality. —Le Have Bank (the center of which is about
43°02’N, 64°01’W), 300 fathoms (549 meters).
Georgta-Florida locality. — “Albatross” sta. 2668 (30°58’30’N,
79°38/30’W ), 294 fathoms (538 meters), about 105 statute miles
east off St. Andrews Sound, Georgia, bottom of gray sand and dead
coral, bottom temperature 46.3°F.
Other localities. — U.S. Fish Commission sta. 1124, off Martha’s
262 BULLETIN 299
Vineyard; “Albatross” sta. 2470 (44°47’N, 56°33’45”W), off Nova
Scotia, 224 fathoms (410 meters), bottom of gray mud, bottom
temperature 40.2°F.; “Albatross” sta. 2527 (41°59’N, 65°35’30”W),
off Georges Bank, 117 fathoms (214 meters), bottom of sand and
gravel.
In working up S. presswm, Pilsbry recognized its general simi-
larity to S. stroemu M. Sars. However, as the original of S. stroemu
was not illustrated by M. Sars, Pilsbry communicated with G. O.
Sars from whom he received two specimens with the notation that
“they may be regarded as typical”. One of these “typical” specimens
was figured by Pilsbry (1907, p. 22, pl. 1, figs. 6, 7) and may be com-
pared with the type of S. pressum on Plate 30, figure 2.
Concerning the relationship of S. pressum to the S. stroemit
complex, Pilsbry wrote as follows:
This species [S. pressum], which seems to be somewhat abundant off our
northeastern coast, resembles the form which Aurivillius has called S. septen-
trionale. It differs from that, however, by the narrower base of the capitulum,
the greater compression, and the position of the umbo of the carina, which is
much nearer the apex. The inframedian lateral plate is longer than in any of
the related forms, and the rostrum has the long and narrow shape figured
by Aurivillius for §. septentrionale and S. obesum. The capitulum of §. pressum
is more lengthened than that of S. stroemii, chiefly by reason of the elongation
of the plates of the lower whorl.
The latitudinal range of S. pressum is between 31° North and
45° North in the Western Atlantic Ocean.
Scalpellum prunulum Aurivillius Pl. 30, figs. 3, 4
Scalpellum prunulum Anurivillius, 1894a, p. 669; 1894b, pp. 62-64, pl. 5,
figs. 3-4; Gruvel, 1905, p. 63 [as Scalpellum primulum], fig. 70; Nilsson-Cantell,
1921, pp. 104, 205.
Aurivillius’ original German description of the exoskeleton is
translated as follows:
Capitulum with 14 plates. Carina gently arched. Rostrum rudimentary
covering only the posterior 1/3 of the rostral Jatera. Umbones of the scutum
and latera as in Sc. erosum, those of the inframedian lying at the apices.
Peduncle with 8 elongated rows, each provided with 5-6 moderately distant
scales, the outer ends of which are interspersed with adjacent scales.
The color of the specimens in alcohol is brownish yellow between the white
plates and scales.
Dimensions. Length of animal 6 mm., length of capitulum 4 mm, breadth
2.5 mm.
[Type] locality. Sea of the Antilles, off St. Martin [18°05’N, 63°05’W],
depth 350-600 meters.
Gruvel (1905) added that the 14 plates are slightly separated
and are covered by a thin and smooth cuticle. The umbo of the
FLoRIDIAN CIRRIPEDIA: WEISBORD 263
carina is at the apex. The umbones of the carinal latera are at the
base and project beyond the dorsal margin and above the carina. The
umbones of the inframedian latera are at the base.
Scalpellum (?) pteryges MacDonald Pl. 30, fig. 5
Scalpellum pteryges Macdonald, 1929, pp. 532-533, pl. 2, fig. 4.
The capitulum is trapezoidal, 23 mm in length and 15 mm in
breadth, with an approximately straight occludent border and a
markedly convex carinal border. There are 14 smooth plates sepa-
rated by narrow chitinous sutures and marked by fine, closely
spaced growth striae.
The tergum is large, subquadrangular with straight occludent
and basal margins, and very convex carinal margin. The scutum is
marked by a prominent ridge running from the acuminate apex to
the basal-lateral angle. The carina is profoundly arched, its acute
apical umbo intruded between the terga. The roof is slightly convex,
bordered by low ridges. The sides are moderately wide, tapering
toward the apex and having a shallow sulcus running the entire
length. The basal margin is deeply rounded.
The upper latus is pentagonal, with the scutal margin longest,
the tergal, carinal, basal, and that bordering on the inframedian latus
successively shorter. The umbo is acute and apical, and there is a
low ridge running from the apex to the carinal-basal angle. The
carinal latus is somewhat triangular, with the base in two parts:
a long upper lateral margin, and a shorter margin lying against the
inframedian latus. The apical portion of the carinal latus is con-
siderably curved downward, the apex itself being very acute. The
winglike latera, viewed dorsally, show the carina extending between
them almost to the peduncle. Toward the apex the latera appear
twisted outward. The rostral latus is raised slightly above the
scutum. The length of the valve is three times as long as wide. The
apex is beaked and a ridge runs from the apex to the basal-lateral
angle. The rostrum is small and triangular, the apex projecting
outward from the apices of the rostral latera. The inframedian latus
is triangular, higher than wide, the margins slightly raised, and the
apex curved toward the occludent border.
The peduncle has about 13 rows of scales and is 10 mm in
length.
Type locality. — “Enterprise” sta. (13°52’N, 67°7’W), off St.
Lucia, attached to cable at a depth of 278 fathoms (508 meters).
264 BULLETIN 299
Scalpellum (?) semisculptum Pilsbry Pl. 31, figs. Ta-c
Scalpellum semisculptum Pilsbry, 1907, pp. 48, 62-64, figs. 25a-c; Mac-
Donald, 1929, p. 535; Broch, 1953, pp. 4, 7. 10, 12, 15; Henry, 1954, p. 44; Zullo,
1968, p. 216.
Pilsbry’s description is summarized as follows:
The capitulum of the type, which is 16 mm in length, 7.7 mm in
breadth, is suboval, twice as long as wide, with the occludent and
carinal borders nearly equally convex. There are 13 fully calcified
plates joined by linear sutures, the plates marked with lines of
growth and fine radial striae, excepting the carinal latera which have
distinct radial riblets. The cuticle covering the capitulum is very
thin and smooth.
The tergum is large and triangular, the occludent margin
strongly arched, the acute summit somewhat recurved; the scutal
margin is a little longer than the occludent; the carinal margin is
convex except near the apex where it is concave. The scutum is
thomboidal, about twice as long as wide, with an acute apical umbo;
the occludent and lateral margins are subparallel and slightly convex;
the tergal margin is concave, the basal straight. The carina is 12.5
mm in length and 2.5 mm in diameter at the base. It is long and
regularly arched, its apical umbo at about the upper fourth of the
tergum. The roof is flat between low but robust bordering ribs;
the sides are narrow below, wider above; the basal margin is convex.
The upper latus is trapezoidal, with a concave scutal margin
and a short carinal margin; the tergal and basal margins are straight
and about equal in length. The umbo is terminal and there is a nar-
row rib extending from it to the baso-carinal angle. The plate is
finely and sharply striate radially. There is no rostrum, or merely
a sunken linear rudiment. The rostral latus is as high as wide, the
basal margin much shorter than the others; the umbo projects a
little at the upper occludent angle, and from it a narrow diagonal
rib runs to the lower lateral angle, the surface below this rib being
radially striate. The two rostral latera rise in a low welt at their
occludent margin. The inframedian latus is almost linear, curving
above slightly toward the the rostral border; the umbo is not visible
but is probably apical. The carinal latus is irregularly trapezoidal,
the obtuse umbo at about the lower third of the carinal margin
and not projecting beyond the carina. The plate is sculptured with
FLORIDIAN CIRRIPEDIA: WEISBORD 265
strong radial riblets. In dorsal view the carinal latera are seen to
be strongly tricostate, and meet in an irregular suture.
The peduncle is 3 mm in length, closely covered with large,
projecting, transversely lengthened scales, in about eight rows of
eight scales each.
Type locality. — “Albatross” sta. 2397 (28°42’N, 86°36’W),
280 fathoms (512 meters), gray mud, bottom temperature 46.1°F,
surface temperature 65°F, Gulf of Mexico, about 280 statute miles
west of Bayport, Florida, and due south of Destin, Florida.
Other localities. — Broch reported this species at “Ingolf” sta.
10 (64°24’N, 28°50’W), 1484 meters (807 fathoms), bottom tem-
perature 3.5°C, about 240 statute miles west of Reykjavik, Iceland.
Broch stated that although the single Icelandic specimen was
smaller (10 mm) than Pilsbry’s type (16 mm), it agreed well with
his description.
Scalpellum (?) sinuatum Pilsbry Pl. 30, figs. 7a-c
Scalpellum sinuatum Pilsbry, 1907, pp. 47, 50-51, figs. 16a-c; Fowler, 1912,
p. 500; Barnard, 1924, pp. 17, 40-43; Nilsson-Cantell, 1955, p. 219; Zullo, 1968,
‘i edit bvitum sp., cf. A. simuatum (Pilsbry), Newman and Ross, 1971, pp.
81-82, pl. 9 D, text-figs. 40A-H.
The capitulum of Pilsbry’s type is trapezoidal, about twice as
long as wide, and measuring 13.5 mm in length and 7 mm in width.
It is composed of 14 nearly smooth plates with no hairs.
The tergum is triangular in shape, with a convex occludent
margin, a slightly recurved apex, and a slight prominence on the
carinal margin just above the apex of the carina. The scutum has
a slightly convex occludent margin and a pointed lateral margin with
a broad excavation below it to accommodate the apex and carinal
margin of the upper altus. The carina is long and arcuate with an
apical umbo. The roof is flat between two moderate rounded ribs.
The sides are wide above, tapering to the base.
The upper latus is subtriangular with a deep notch in the lower
margin; the scutal margin is longer than the tergal and the mucro is
at the scuto-tergal angle. The rostrum is small and subtriangular,
lying between the umbones of the rostral latera. The rostral latus
is nearly as high as wide, its basal margin shorter than the scutal, its
lateral margin convex. The inframedian latus is wineglass-shaped, its
266 BULLETIN 299
upper margin concave and nestled into the upper latus; the umbo
is median, and the base of the plate is expanded. The carinal latus is
irregularly triangular, projecting a little beyond and above the
carina, the umbo slightly recurved at the base of the carina; the two
latera meet in a short suture below the carina.
The peduncle is 5 mm in length, with ten rows of large scales,
about six scales in a row.
According to Pilsbry, the adult Scalpellum sinuatum is notable
for the prominent notch in the lower margin of the upper latus
which is “unlike any known form of the same group”. Also charac-
teristic is the “very small, nodule-like rostrum visible only between
the apices of the rostral latera”.
Type locality. — “Albatross” sta. 2037 (38°53’N, 69°23’30’”N),
1731 fathoms (3166 meters), Globigerina ooze, bottom temperature
38°F, surface temperature 76°F, about 305 statute miles east of
mouth of Delaware Bay and 7 degrees of latitude north of Florida.
Other localities. —“Pieter Faure” sta. of 14 July 1903, in the
Eastern Atlantic, 40 miles south-southwest of Cape Point, South
Africa, 800-900 fathoms; “Pieter Faure” sta. of 19 August 1903, 43
miles nearly due west of Cape Point, 900-1000 fathoms (The stations
off Cape Point lie between 34 and 35 degrees South and 18 and 17
degrees East); “Eltanin” sta. 18 (58°15’N, 48°36’W), 3404-3422
meters, southwest off Kap Farvel, Greenland.
Barnard (1924), who identified S. sinwatum off Cape Point,
stated, “The identification of these specimens has caused me con-
siderable difficulty, and other workers may differ from my conclu-
sions”. Nilsson-Cantell (1955) did not list S. stnwatwm as occurring
off Cape Point but that may have been because he listed only those
species recovered at depths below 3000 meters whereas the greatest
depth recorded by Barnard was 1829 meters (1000 fathoms) off
Cape Point. Newman and Ross (1971) determined that certain
skeletal features of their Arcoscalpellum sp. from Greenland waters
are similar to those from off the Delaware coast, yet the absence of
the sinus in the basicarinal margin of the upper latus persuades them
that their Greenland taxon may be distinct from Pilsbry’s type of
S. sinuatum which is also nearly twice as large. It would thus seem
to this writer that the type locality of S. stnwatwm is as yet the only
one known for the species.
FLORIDIAN CIRRIPEDIA: WEISBORD 267
Arcoscalpellum aurivillii (Pilsbry) Ply 27, figs: laeb
Scalpellum aurivillii Pilsbry, 1907, pp. 48, 64-66, figs. 26a-b; Fowler, 1912,
p. 500; MacDonald, 1929, p. 535; Withers, 1953, pp. 9, 10, figs. 11a, b, as
Arcoscalpellum,; Broch, 1953, pp. 4, 7, 12, 15; Zullo, 1968, p. 211.
The capitulum of the type is rhombic-oblong, 15.3 mm in length,
7.5 mm in breadth, and is composed of 13 fully calcified plates,
separated by linear sutures, and marked with fine, irregular lines
of growth, and minute, inconspicuous radial striae.
The tergum is triangular with a convex occludent margin, a
straight scutal margin, a weakly sigmoidal carinal margin which is
concave above and convex below, and a slightly recurved umbo.
The scutum is longer than wide, its acute apex recurved within the
ventral border; the lateral margin is concave below the tergo-internal
angle, convex in the middle, and slightly recessed at the basal angle;
the basal margin is nearly straight. The carina is 13.5 mm in length,
2.2 mm in diameter at the base. It is simply arched, more strongly
so above, and its umbo is terminal; the roof is flat with distinct
bordering ribs; the sides are moderately developed near the umbo,
narrow elsewhere; the basal margin is straight, as are the lines of
growth across the roof.
The upper latus is trapezoidal, the scutal margin much the
longest and concave; the other margins are straight, the carinal the
shortest; the apex is produced in a small triangle above and beyond
the umbo, which is acute and marginal, on the scutal side. There is
no rostrum. The rostral latus is quadrangular, the ventral and scutal
borders straight, the basal short where it comes in contact at the
upper interior angle with the upper latus; the carinal margin is in
contact with the carinal latus, but the suture is more or less covered
by the extremely narrow inframedian latus, which overlies the borders
of the plates. The inframedian latus is narrowly triangular, the umbo
apical; it overlies the suture instead of occupying a space between the
rostral and carinal latera, and is often abnormal. The carinal latus is
twice as high as wide, quadrangular, the umbo at its lower third not
projecting beyond the carina; the basal and rostral margins are
subequal, straight, and at right angles; the carinal margin is nearly
straight, projecting a little in the lower third; the two latera meet
below the carina; from the umbo of the carinal latus a conical raised
and radially costulate area extends to the basal margin.
268 BuLLETIN 299
The peduncle is 5 mm in length. It is compactly covered with
narrow transverse scales in eight rows of about eight scales each.
Type locality. —“Albatross” sta. 2731 (36°45’W, 74°28’W),
781 fathoms (1428 meters), growing on Scalpellum velutinum
Hoek, about 90 statute miles east-southeast of Virginia Beach,
Virginia.
Other localities. —“Albatross” sta. 2728 (36°30’N, 74°33’W),
850 fathoms (1555 meters), gray ooze, about 83 statute miles east
off Currituck, North Carolina. This locality is the nearest one to
Florida which is some 6 degrees of latitude to the south; “Albatross”
sta. 2710 (40°06’N, 68°01’30” W ), 984 fathoms (1800 meters), green
mud, about 320 statute miles east off Point Pleasant, New Jersey;
“Albatross” sta. 2529 (41°03’30”N, 66°14’W), 662 fathoms (1211
meters), gray mud, bottom temperature 38.7°F, about 290 statute
miles east off Montauk Point, New York; U.S. Fish Commission sta.
1123, off Martha’s Vineyard; “Tjalfe” sta. 408 (64°14’N, 55°55’W),
839 meters (453 fathoms), northwest off Godthaab, Greenland.
Arcoscalpellum aurivilli (Pilsbry) is a Western Atlantic and
North Atlantic species ranging geographically from off the coast
of North Carolina in the south to Greenland in the north, and oc-
curring in waters with reported depths of 406 fathoms to 984
fathoms.
Arcoscalpellum aurivillii incertum (Pilsbry) Pl. 27. figs 2e
Scalpellum aurivillii incertum Pilsbry, 1907, p. 67, fig. 26c; Withers, 1953,
pp. 9, 10, fig. 11c, as Arcoscalpellum,; Zullo, 1968, p. 214.
Although this subspecies was recovered in the Northeast Pacific,
it is mentioned here because of its possible relationship to Scalpel-
lum aurivillu aurivilli Pilsbry which is a Western Atlantic species
and, therefore, within the purview of this work.
Pilsbry’s description of S. a. incertwm was as follows:
A single example (Cat. No. 32871, U.S.N.M.), evidently very closely related
to S. aurivillii, was found growing on the peduncle of one of a series of S.
regium var., said to be from Albatross Station 3342, off British Columbia, in
1,588 fathoms. Having been preserved probably in formaldehyde, the apices of
the valves are more or less eroded, especially those of the terga. Allowing for
this the length of the capitulum would be 24, breadth 13.5 mm; length of the
peduncle 7.5 mm. Length of the carina 22, diameter at base 3 mm. The plates
are pale cream-colored, smoothish, except for narrow, widely spaced growth-
arrest marks. On the roof of the carina the growth lines arch downwards. The
FLORIDIAN CIRRIPEDIA: WEISBORD 269
upper latus is larger than in S. aurivillii, its length being twice the breadth,
and its carinal margin is decidedly longer than in S. aurivillii. On the right
side of the capitulum there is no inframedian latus and no indication that there
ever was one, and on the left side only a small basal triangular plate; but the
absence of these plates may be due to the action of the formalin, though I can
not positively affirm that this is the case. The rostral latus is comparatively
lower and wider, its greatest height only half the width. No rostrum. In other
characters of the plates there is no important divergence from 8S. aurivillit, ex-
cept for size, which is much greater than that of any of the series of apparently
adult examples of that species. . .
Type locality.—“Albatross” sta. 3342 (52°3930’N, 132°38’W),
1588 fathoms (2904 meters) gray ooze and coarse sand, bottom
temperature 35.3°F, surface temperature 57°F; about 30 miles west
off Moresby Island, British Columbia, Canada.
Arcoscalpellum regina (Pilsbry) Pl. 30, fig. 6
Scalpellum regina Pilsbry, 1907, pp. 25, 31-32, pl. II, figs. 4-6; Calman,
1918a, pp. 112-113; Barnard, 1925, pp. 1, 2-3; U.S. Naval Inst., 1967, p. 194;
Henry, 1954, p. 444; Zullo, 1968, p. 216; Kaufmann, 1971, pp. 73-85, figs. 1-4.
Following is a summary of Pilsbry’s description:
The capitulum, measuring 43 mm in height and 34 mm in
breadth, is moderately compressed, high domal in outline, acuminate
at the apex, subtruncate at the contact with the peduncle. The
capitulum is covered with a densely and shortly pilose cuticle. There
are 14 plates separated by wide chitinous sutures in adults but in
contact in immature specimens. The plates are weakly sculptured
with widely spaced low wrinkles along the lines of growth.
The tergum is divided into two areas by a straight apico-basal
ridge, the carinal area about half as wide as the scutal. The scutum
is twice as long as wide, its occludent margin arcuate, its acuminate
apex a little recurved, its basal and lateral margins straight, and the
tergal margin straight immediately below the apex. The carina, 40
mm in length and 6 mm in diameter at the base, is gently arched,
separated from the scuta and latera by a wide chitinous space. The
umbo is terminal at the apex which intrudes slightly between the
scuta. The roof is slightly convex, marked with V-shaped lines of
growth, the sides narrow throughout, and the base wedged between
the carinal latera.
The upper latus is subpentagonal with slightly concave tergal
and scutal margins and subrounded carinal and basal margins; the
umbo is at the apex. The rostrum is small and triangular and
separates slightly the rostral latera. The rostral latus is low, the
270 BULLETIN 299
upper and lower margins parallel. The inframedian latus is small
and triangular with the basal margin the longest; the umbo is apical.
The carinal latus is irregular in shape. The convex posterior margins
project beyond the carina, and the two latera meet below it. The
umbo is elevated, acute, and curved toward the scutal margin. A
prominent ridge runs from the umbo to the scutal end of the plate
and there are two or three inconspicuous ridges to the basal margin.
The peduncle is 26 mm in length and is covered with large
scales clothed in a velvety cuticle. There are 10 rows of about 12
scales each in the figured type but more in old individuals.
Type locality. — “Albatross” sta. 2376 (29°03’N, 88°16’W),
Gulf of Mexico about 95 miles southeast of Pascagoula, Mississippi,
and southwest of Pensacola, Florida, in 324 fathoms (593 meters),
bottom of gray mud, bottom temperature 46.5°F.
Additional localities in Gulf of Mexico. — Provided by Henry A.
Spivey of Florida State University who obtained the data from Jack
Rudloe, Gulf Specimen Co., Panacea, Florida. Trawl between
29°32’N, 86°57’W and 29°25’N, 87°15’W, about 53 statute miles
south of Pensacola, Florida, 216-228 f.; trawl at 29°24’N, 87°12’W,
about 46 statute miles southeast of Gulf Beach, Florida, 230-248 f.;
trawl between 29°16N, 87°42’W and 29°25’N, 87°23’W, about 67
statute miles south of Orange Beach, Florida, 198-300 f.; “Eric
Wakefield” sta. at 29°07’N, 88°10’W, trawl, about 107 statute miles
southeast of Biloxi, Mississippi, 370 f.
Among a cluster of numerous specimens of Arcoscalpellum re-
gina attached to each other and collected in the Gulf of Mexico off
one or the other of the Florida locations mentioned above, there are
two measuring over 130 mm in length when fully extended. The
capitulum of one is about 55 mm in length and 45 mm in width,
and the capitulum of the other about 50 mm in length and 38 mm
in width. The maximum diameters of the peduncle are 30 mm and
28 mm, respectively.
Other localities. — Off the Caribbean coast of Colombia at the
following stations — “Oregon” sta. 4882 (10°16’N, 75°54’W), 30
km west of Isla Bari (10°10’/N, 75°36’W), 300 fathoms; “Oregon IT”
sta. 267 (11°12’N, 74°21’W), 14 km west of Santa Marta, 240
fathoms; “Oregon IT” sta. 268 (11°26’N, 74°14’°W), 21 km west of
Santa Marta, 280 fathoms; “Oregon II” sta. 287 (11°35’N,
FLORIDIAN CIRRIPEDIA: WEISBORD 271
73°26’W ), 57 km west of Riohacha, Colombia (11°34’N, 72°57’W),
250 fathoms; “Oregon II” sta. 288 (11°27’N, 73°42’W), 87 km
west-southwest of Riohacha, 220 fathoms; “Oregon II” sta. 289
(11°24’N, 73°47’W), 95 km west-southwest of Riohacha, 150
fathoms; Brazil — “Norseman” sta. (7°37’S, 34°26.5’W), 50-150
fathoms, 55 statute miles northeast off Pernambuco.
To judge from the localities at which this species is reported,
Arcoscalpellum regina (Pilsbry) has a latitudinal range of some 36
degrees and a longitudinal range of some 54 degrees, from the
northern Gulf of Mexico to the Western Atlantic off the bulge of
Brazil.
Arcoscalpellum regium (Thomson) Pl. 31, figs. 1-5
Scalpellum regium Thomson, 1878, vol. 2, pp. 11-14, figs. 2-3; Hoek, 1883,
pp. 22, 27, 29, 65, 96, 100, 104, 105, 106-109, 111, 122, 124, 126, pl. 4, figs. 3-5,
pl. 9, fig. 12, pl. 10, figs. 1-2; Aurivillius, 1894b, p. 89; Weltner, 1897, p. 249;
Gruvel,/ 1905: p: 77, figs: S6A-B; 1912, p: 25) 1920; pp. 30, 85, pl. I, fig: 75
Pilsbry, 1907, pp. 25, 28-29, pl. 3, figs. 4-5; Fowler, 1912, p. 499; Kriiger, 1940,
p. 225; Nilsson-Cantell, 1955, p. 219; Newman and Ross, 1971, p. 71.
Scalpellum regiwm Thomson was recovered with a trawl on
June 17, 1873 at a depth of 2850 fathoms, adhering to a concre-
tionary mass containing a large percentage of peroxide of manganese.
Thomson’s excellent description is repeated in full.
Scalpellum regium (Fig. 2) is one of the largest of the known living species
of the genus. The extreme length of a full-sized specimen of the female is 60
mm., of which 40 mm. are occupied by the capitulum and 20 mm. by the ped-
uncle. The capitulum is much compressed, 25 mm. in width from the occludent
margin of the scutum to the back of the carina. The valves are 14 in number;
they are thick and strong, with the lines of growth strongly marked, and they
fit very closely to one another, in most cases slightly overlapping. When living,
the capitulum is covered with a pale brown epidermis, with scattered hairs
of the same color.
The scuta are slightly convex, nearly once and a half as long as broad.
The upper angle is considerably prolonged upwards, and, as in most fossil
species, the centre of calcification is at the upper apex. A defined line runs
downwards and backwards from the apex to the angle between the lateral
and basal margins. The occludent margin is almost straight; there is no de-
pression for the adductor muscle, and there is no trace of notches or grooves
along the occludent margin for the reception of the males; the interior of the
valve is quite smooth. The terga are large, almost elliptical in shape, the centre
of calcification at the upper angle. The carina is a handsome plate, very uni-
formly arched, with the umbo placed at the apex; two lateral ridges and a
slight median ridge runs from the umbo to the basal margin; the lower part
of the valve widens out rapidly, and the whole is deeply concave. The rostrum,
as in Scalpellum vulgare, is very minute, entirely hidden during life by the in-
vesting membrane. The upper latera are triangular, the upper angle curving
rather gracefully forwards; the umbo of growth is apical.
The rostral latera are long transverse plates lying beneath the basal
margins of the scuta. The carinal latera are large and triangular, with the apex
272 BULLETIN 299
curved forward very much like the upper latera, and the infra-median latera
are very small, but in form and direction of growth nearly the same.
The peduncle is round jn section and strong, and covered with a felting
of light-brown hair. The scales of the peduncle are imbricated and remarkably
large, somewhat as in S. ornatum, Darwin. About three, or at most four, scales
pass entirely round the peduncle. The base of attachment is very small, the
lower part of the peduncle contracting rapidly. Some of the specimens taken
were attached to the lumps of clay and manganese concretions, but rather
feebly, and several of them were free, and showed no appearance of having
been attached. There is no doubt, however, that they had all been more or less
securely fixed, and had been pulled from their places of attachment by the
trawl. On one lump of clay there were one mature specimen and two or three
young ones, some of these only lately attached. The detailed anatomy of this
species will be given hereafter, but the structure of the soft parts is much the
same as in Scalpellum vulgare.
In two specimens dissected there was no trace of a testis or of an intromit-
tent organ, while the ovaries were well developed. I conclude, therefore, that
the large attached examples are females, corresponding, in this respect, with
the species otherwise almost nearly allied, S. ormatum.
In almost all the specimens which were procured by us, several males, in
number varying from five to nine, were attached within the occludent margin
of the scuta, not imbedded in the chitinous border of the valve, or even in any
way in contact with the shell, but in a fold of the body-sac quite free from the
valve. They were ranged in rows, sometimes stretching — as in one case
where there were seven males on one side — along the whole of the middle two-
thirds of the edge of the tergum.
The male of Scalpellum regium (Fig. 3) is the simplest in structure of these
parasitic males which have yet been observed. It is oval and sac-like, about 2
mm. in length by 9 mm. in extreme width. There is an opening at the upper
extremity which usually appears narrow, like a slit, and this is surrounded by
a dark, well-defined, slightly raised ring. The antennae are placed near the
posterior extremity of the sac, and resemble closely in form those of §. vulgare.
The whole of the sac, with the exception of a small bald patch near the point
of attachment, is covered with fine chitinous hairs arranged in transverse
rings. There is not the slightest rudiment of a valve, and I could detect no
trace of a jointed thorax, although several specimens were rendered very trans-
parent by boiling in caustic potash. There seems to be no oesophagus nor
stomach, and the whole of the posterior two-thirds of the body in the mature
specimens was filled with a lobulated mass of sperm-cells. Under the border of
the mantle of one female there were the dead and withered remains of five
males, and in most cases one or two of the males were not fully developed;
several appeared to be mature, and one or two were dead — empty, dark-
colored chitine sacs.
Type locality. — “Challenger” sta. 61 (34°54’N, 56°38’W),
2850 fathoms (5212 meters), bottom of gray ooze, temperature
1.5°C, about 840 statute miles south of Grand Bank, Newfoundland,
and 1140 miles east of Cape Lookout, North Carolina.
Other locahties.— “Challenger” sta. 63 (35°29N, 50°53’W),
2750 fathoms (5030 meters), bottom of gray ooze, about 1120 statute
miles east off Cape Hatteras, North Carolina, and southeast off
Cape Race, Newfoundland; “Albatross” sta. 2226 (37°N, 71°54’W),
2045 fathoms (3740 meters), bottom of Globigerina ooze, bottom
FLORIDIAN CIRRIPEDIA: WEISBORD 273
temperature 36.8°F, about 220 miles east of Newport News, Vir-
ginia, seated on a slender gorgonian stem and on a pebble, and at-
tached to Scalpellum albatrossianum Pilsbry. This locality is the one
nearest to Florida; “Albatross” sta. 2228 (37°25’N, 73°06’W), 1582
fathoms (2893 meters), bottom of brown mud, temperature 36.8°F,
235 miles east of Newport News, Virginia; “Albatross” sta. 2533
(40°16’30”N, 67°26’15”W), 828 fathoms (1514 meters), bottom
of brown ooze, temperature 38.7°F, about 340 statute miles east of
New York City and 240 statute miles southwest of Cape Sable, Nova
Scotia; “Albatross” sta. 2575 (41°07’N, 65°30’'W), 1710 fathoms
(3128 meters), bottom of gray ooze, temperature 37.1°F, about 220
statute miles east of Nantucket, Massachusetts, and about 150
statute miles south of Cape Sable, Nova Scotia; “Prince de Monaco
Cruise of 1910” (43°21’N, 10°02’W), 2779 fathoms (5083 meters),
northwest off Cape Finisterre (42°54’N, 9°16’W), Spain.
The latitudinal range of A. regiwm is from about 34° North to
43° North, the longitudinal from 10° West to 73° West. Depths
range from 828 to 2850 fathoms (1514 to 5212 meters).
Arcoscalpellum velutinum (Hoek) Pl. 32, tgs. i 2
This name is applied to a taxon variously identified by authors
as Scalpellum michelottianwm Seguenza (1876); Scalpellum velu-
tinum Hoek (1883); Scalpellum eximiwm Hoek (1883); Scalpellum
sordidum Aurivillius (1898); Scalpellum erectum Aurivillius (1898);
and Scalpellum alatum Gruvel (1900). My own feeling, based on
comparing the illustrations of the types (PI. 32) and on their original
descriptions, is that S. michelottianum, S. velutinum, and S. eximium
are distinct species, and that S. sordidum, S. erectum, and S. alatum
may perhaps be synonymous with one or the other of the six species
listed above. The synonymy proposed by authors is the following:
Scalpellum michelottianum Seguenza, 1876, pp. 381-386, 422, 423, 426, 427,
432, 464, 481, pl. 6, figs. 15-25, pl. 10, figs. 26, 26a; Alessandri, 1894, pp. 263-
265, pl. 1, figs. 6a-6m, pars; 1897, p. 47; 1906, pp. 251-252; Pilsbry, 1907, p.
32; Withers, 1953, as Arcoscalpellum, pp. 101, 225-229, pl. 37, figs. 1-10, pl.
64, fig. 4; Newman, Zullo, and Withers, 1969, as Arcoscalpellum, p. R277;
Newman and Ross, 1971, as Arcoscalpellum, p. 71, figs. 34A-J, pl. [X,B; Rao
and Newman, 1972, as Arcoscalpellum, pp. 76-80, figs. 5, 11A-B. [Plate 32,
figs. 5(15 to 25)].
Scalpellum velutinum Hoek, 1883, pp. 22, 25, 27, 31, 65, 96-99, 100, 104, 105,
126, pl. 4, figs. 10-11; pl. 9, figs. 7-9; 1914, p. 4; Weltner, 1895, p. 289; 1897, p.
251: 1922, pp. 75, 92, 94, 106, pl. 3, fig. 10; Murray, 1896, pp. 386, 397, 453;
Gruvel, 1902a, pp 31, 50, 52, 57, 136-137, pl. 2, figs. 3c 14; pl. 3, figs. 1, 27-31;
pl. 4, figs. 6, 11-22; 1902c, p. 523; 1905, pp. 73-74, fig. 83; 1912, p. 2; 1920, pp.
226 eS Me Son plede tics 8-1 Ope 7 ties 4 Annandale. 1905) p: 183)
1908, pl. 4. fig. 7; 1911, pp. 588, 589; zon 1913, pp. 228-229 (= S. annan-
274 BULLETIN 299
dalei Calman); 1916a (?), pp. 128-129, pl. 6, figs. 6-7; Pilsbry, 1907, pp. 25,
26-27, 64, 75, pl. 3, figs. 2-3; 1908, as Scalpellum (Arcoscalpellum), pp. 105,
109, figs. li, j; Fowler, 1912, p. 499; Calman, 1918a, pp. 108-109; Broch, 1924,
p. 39; 1953, p. 9; Barnard, 1925, pp. 1-2; Nilsson-Cantell, 1927, pp. 743-745,
text-fig. 1; 1928, p. 4; 1931la, pp. 1-2; 1938, pp. 8, 18, 21; Stubbings, 1936, pp.
2, 28, 29, 30, 67; 1967, p. 234; Kriiger, 1940, as Arcoscalpellum, pp. 46, 63, 113,
139, 141, 265, figs. 28a, 143a-c; Withers, 1953, as Arcoscalpellum, pp. 97, 196,
228; Tarasov and Zevina, 1957, p. 24, figs. 9, 11; Bassindale, 1964, p. 31, fig.
on p. 54; U.S. Naval Inst., 1967, p. 194; Zullo, 1968, as Arcoscalpellum, p. 213;
Newman, Zullo, and Withers, 1969, as Arcoscalpellum, p. R277; Newman and
Ross, 1971, as Arcoscalpellum, p. 73; Collins and Mellen, 1973, as Arcoscalpel-
lum, p. 363.
Scalpellum eximium Hoek, 1883, pp. 22, 25, 31, 98, 100-102, pl. 4, figs. 6-7;
pl. 9, figs. 10-11; Weltner, 1897, p. 247; Gruvel, 1905, p. 73; 1912, p. 2; Annan-
dale, 1913, p. 229; Barnard, 1925, p. 1; Withers, 1953, as synonymous with
Arcoscalpellum michelottianum (Seguenza), p. 225; Newman and Ross, 1971, p.
72. [Pl. 32, figs. 3-4.]
Scalpellum sordidum Aurivillius, 1898, pp. 190-191; Gruvel, 1905, p. 73;
1912, p. 2; 1920, pp. 27-28, pl. 1, fig. 15, as S. velutinum forma sordidum;
Barnard, 1925, p. 1; Withers, 1953, as synonymous with Arcoscalpellum miche-
lottianum (Seguenza), p. 225; Newman and Ross, 1971, p. 72.
Scalpellum erectum Aurivillius, 1898, p. 192; Gruvel, 1905, p. 73; 1920, pp.
27-28, as §. velutinum forma erectum; Withers, 1953, as synonymous with
Arcoscalpellum michelottianum (Seguenza), p. 225; Newman and Ross, 1971,
sr ae alatum Gruvel, 1900, p. 192; 1905, p. 73; 1912, p. 2; Barnard,
1925, p. 1; Withers, 1953, as synonymous with Arcoscalpellum michelottianum
(Seguenza), p. 225; Newman and Ross, 1971, p. 72.
EYPE, LOCALLEIES
The taxon Scalpellum michelottianum Seguenza is a fossil form
wholly reconstructed by Seguenza from numerous but discrete and
separated valves and scales first found near the town of Messina
(38°13’N, 15°33 E), in Sicily. These external components of the
species have since been discovered abundantly in the Plaisancian,
Zanclian, and Astian stages (lower to upper Pliocene) in Sicily and
Italy. In Sicily they are common in the Pliocene of Messina Province
at Salice, Soppo, Trapani, and Gravitelli; in Italy proper they have
been reported from Reggio (38°06’N, 15°39E) in the Province of
Calabria. To my knowledge S. michelottianwm has not been reported
living in the waters surrounding Italy.
The holotype of Arcoscalpellum velutinum (Hoek) was re-
covered at “Challenger” sta. 3 (37°2’N, 9°14’W), 900 fathoms, in
Globigerina ooze, off Cabo Sao Vicente, Portugal. The paratype was
recovered at “Challenger” sta. 335 (32°24’S, 13°5’W), 1425 fathoms
(2606 meters), bottom temperature 2.3°C, in Globigerina ooze,
about 270 statute miles north of Tristan da Cunha.
FLORIDIAN CrIRRIPEDIA: WEISBORD 275
The type of Scalpellum eximiwm Hoek was taken at “Chal-
lenger” sta. 135, between Nightingale Island (37°28’S, 12°32’W)
and Tristan da Cunha (37°15’S, 12°30’W). First sounding 1000
fathoms (1829 meters), shells and rock on bottom; second sounding
1100 fathoms (2012 meters).
The type of Scalpellum sordidwm Aurivillius was recovered
during the “Prince de Monaco Campagne 1887”, sta. 161
(46°04’40”N, 46°42’715”W), 1267 meters, soft gray mud, off New-
foundland.
The type of Scalpellum erectwm Aurivillius was recovered dur-
ing the “Prince de Monaco Campagne 1887”, sta. 227 (38°23’N,
28°26'37” W ), 1135 meters, bottom of rock, gravel, and broken shells,
near south coast of Pico, Azores.
The type of Scalpellum alatum Gruvel was recovered during
the “Campagne du Talisman” at Cap Cantin (32°33’N, 9°17’W),
Morocco.
In this work the name Arcoscalpellum velutinum (Hoek) has
preference for the reason that this species, according to Pilsbry,
occurs in the Western Atlantic along the east coast of North America
from Newfoundland southwestward to off South Carolina. A. velu-
tinum has not been reported from Florida, but inasmuch as it occurs
within two or three degrees of latitude south of South Carolina, it is
possible that A. velutinum will eventually be discovered in the deeper
waters off Florida’s east coast.
Hoek’s description of his Scalpellum velutinum is summarized
as follows:
The capitulum of the type is 33 mm in length, is covered by a
velvety hirsute membrane, and consists of 14 valves which touch
each other. The carina is gently arched, its flat roof widening from
the umbo to the base, its apex penetrating between the two terga.
The tergum is large and narrowish, the carinal and scutal margins
moderately convex, the occludent margin excavated, the umbo nar-
rowly rounded at the apex. The scutum is tumid, its length twice the
breadth, the apex of the umbo sharply pointed.
The upper latus is triangular, the basal margin slightly convex,
the scutal and tergal margins nearly equal, the umbo at the apex.
The rostrum is small and totally covered by membrane; it is tri-
angular in shape, with the apex separated from the two scuta by the
276 BULLETIN 299
umbones of the rostral latera which touch each other in front of
the rostrum. The rostral latus is broad and low, the basal margin
almost parallel with the scutal margin. The inframedian latus is small
and triangular.
The carinal latus is robust and irregular in shape; the carinal
margin is divided into an upper hollowed out portion to receive the
convex margin of the upper latus, and a lower convex portion be-
neath the middle of the carina; from the umbo arises a ledge which
divides the valves into a true lateral and carinal part. Between the
latter and the carina a distinct cavity or kind of bag is formed.
The peduncle is robust, nearly cylindrical, 12 mm in length. The
scales are covered by a membrane, the edges of the scales only being
calcareous. There are about 12 scales in each obliquely longitudinal
row, of which there are about 10.
Type locality. — “Challenger” sta. 3 (37°2’N, 9°14’°W), 900
fathoms (1646 meters), off Cabo Sao Vicente, Portugal.
Paratype. — “Challenger” sta. 335 (32°24’S, 13°5’W), about
270 statute miles north of Tristan da Cunha, depth 1425 fathoms
(2606 meters).
Range and distribution. — The taxon Arcoscalpellum velutinum
or its congeners is reported from the Atlantic Ocean, the Mediter-
ranean Sea, the Indian Ocean, off both the west and east coast of
Africa, the Gulf of Oman, and Indonesia. As noted by Nilsson-Cantell
(1927), however, the descriptions of the internal parts of many of
the synonymized species are lacking, and that in the absence of those
characters the identification of a taxon as Arcoscalpellum velutinum
may be suspect. Nevertheless A. velutinwm s. l. has been reported
from as far north as 72° in the Atlantic to as far south as the Ker-
guelen Archipelago (48°37’S). Depths range from 63 meters, to 3422
meters off the southern tip of Greenland. Details are as follows:
Northern and Western Atlantic: Off southern tip of Greenland,
“Eltanin” sta. 18 (58°15’N, 48°36’W), 3404-3422 meters; New-
foundland (46°04’40”N, 46°42715”W), 1267 meters, soft gray
mud; Numerous stations from Newfoundland southwestward to
South Carolina (consult Pilsbry, 1907, p. 27); “Eastward” sta. 7552
(33°39.5’N, 75°41’W), 301 meters, about 150 statute miles east off
Crescent Beach, South Carolina; “Albatross” sta. 2678 (32°40’N,
76°40/30’”W ), 371 fathoms (679 meters), about 285 statute miles
FLoRIDIAN CIRRIPEDIA: WEISBORD INET
east off Fort Sumter, South Carolina. This locality is the nearest
one to Florida.
Eastern and Southern Atlantic: Off Ireland (51°22’N, 12°W),
695-720 fathoms; Off Liston, Portugal (38°21’N, 9°41’37”W), 2028
meters, brownish gray limy mud; Off Cabo Sao Vicente (Cape St.
Vincent), Portugal (37°2’N, 9°14’W) 1615 meters; Gibraltar,
“Michael Sars” sta. 24 (35°34’N, 7°35’W), 1615 meters. Azores
(38°47’N, 30°16’W), 1331 meters; (38°26’N, 26°30’45”W), 1165
meters, argillaceous sand; “Michael Sars” sta. 53 (34°5YN, 33°1’/W),
2865 meters, about 275 statute miles southwest of Faial Island,
Azores. Canary Islands (29°06’30’N, 13°02’45”W), 1098 meters,
sandy clay; Fuerteventura (28°25’N, 14°W), 2000 meters. “Chal-
lenger” sta. 335 (32°24’S, 13°5’W, 1425 fathoms (2606 meters),
about 270 statute miles north of Tristan da Cunha; South Africa
(34°32’S, 17°49’E), about 40 statute miles southwest of Cape Town,
on water-logged pumice stone and phosphate nodule, depth 612
fathoms (1119 meters).
Mediterranean Sea: Monaco, 515-2028 meters.
West Africa: Morocco - Mogador (31°30’N, 9°48’W), 1050
meters; Sidi Moussa (33°N, 8°50°W); Cap Cantin (32°33’N,
9°17’W), 1350-1590 meters, as S. alatum; Cap Noun, 1255 meters,
as S. alatum; Spanish Sahara (Los Pilones (25°48’N, 14°40’W),
882 meters.
Gulf of Oman: 430 fathoms (786 meters).
East Africa: “Valdivia” sta. 257 (1°48.2’N, 45°42.5’E), depth
1644 meters, bottom temperature 4.6°C, just off Mogadishu,
Somalia; Gulf of Aden (12°20’N, 52°30’E), Socatra Island; “Col-
onia” sta. Aden-Zanzibar cable.
Indian Ocean: “Investigator” sta. 232 (7°17'30’N, 76°54’-
30”E), 40 fathoms (73 meters), off Trivandrum, India; Nicobar
Island, India (6°39’N, 93°12’E), 880 fathoms (1610 meters);
(6°12’N, 93°52’E), 600-1300 fathoms (1097-2378 meters).
Indonesia: “Recorder” sta. off south coast of Bali (8°46’S,
114°44’E), 400 fathoms (732 meters); “Patrol” sta. southeast of
Sumba (10°45’S, 120°50’E), 700 fathoms (1280 meters); “Patrol”
sta. south off Sumba (11°S, 121°30’E), 500 fathoms (914 meters);
“Patrol” sta. off Sawu (11°S, 122°E), 600 fathoms (1097 meters).
278 BuLLeETIN 299
The living Arcoscalpellum velutinum is a deep water species
which attaches itself to varying substrates, among them telegraph
cables, pumice stone, phosphatic nodules, and corals.
Arcoscalpellum vitreum (Hoek) Pi. 3a, i= 8
This species has also been known under several names: Scalpel-
lum talisman Gruvel, Scalpellum formosum Pilsbry, and Scalpellum
bellum Pilsbry, the last as replacement for Scalpellum formosum
Hoek.
The following synonymy is adopted from Newman and Ross
(1971) with a few additions.
Scalpellum vitreum Hoek, 1883, pp. 22, 35, 65, 115-116, pl. 5, fig. 14; Welt-
ner, 1897, p. 251; Gruvel, 1902b, p. 54; 1905, pp. 84-85, fig. 94; Pilsbry, 1907,
p. 60; Nilsson-Cantell, 1955, p. 219; Tarasov and Zevina, 1957, p. 142; Utinomi,
1958, pp. 283-286, figs. 1-2.
Scalpellum talismani Gruvel, 1900, pp. 193-194; 1902b, p. 86, pl. 2, figs. 3D,
6, 7; 1905, p. 86, fig. 96; 1920, p. 23 [see Pl. 33, fig. 2, this report.]; Nilsson-
Cantell, 1955, p. 219; Broch, 1953, p. 8, fig. 4; Zullo, 1968, p. 211.
Scalpellum formosum Pilsbry [not Hoek], 1907, pp. 47, 58-60, figs. 22a-c;
Weltner, 1922, pp. 95-96; Fowler, 1912, p. 500; Stubbings, 1936, pp. 55-56, text-
fig. 24, pars; Nilsson-Cantell, 1938, p. 21; 1955, p. 219.
Scalpellum bellum Pilsbry, 1908, p. 111 [mew name for S. formosum Pilsbry,
1907, zon Hoek, 1907]; Zullo, 1968, p. 211.
Scalpellum sp. cf. bellum Bayer, Voss, and Robins, 1970, p. A43.
Arcoscalpellum vitreum (Hoek), Newman and Ross, 1971, pp. 87-91, 195,
197, pl. 8 E, F, text-figs. 44-47.
Hoek’s original diagnosis was as follows. “Surface of the valves
smooth, not covered by membrane, beautifully striated. Valves thir-
teen. Carina simple, only slightly bowed, with the roof flat. Umbo
of the carina at the apex. Upper latus trapeziform. Infra-median
latus small, triangular. Other valves of the lower whorl well-
developed. Peduncle short.”
The capitulum of the type is elongate-ovate, 13.5 mm in length.
The tergum is slightly smaller than the scutum, with the carinal
margin considerably longer than that of the scutum. The scutum is
quadrilateral. The carina has a flat roof which is wider below than
above, and in the superior half the sides are distinctly furrowed.
The four margins of the upper latus are nearly straight. The rostral
latus is almost triangular. The inframedian latus has the umbo at
the superior extremity. The carinal latus is large and almost trapezi-
form; it has the umbo at one-fourth the total length from the in-
ferior extremity, and the part above the umbo is slightly excavated.
The peduncle is 3.5 mm in length and is covered with a membrane
FLORIDIAN CrIRRIPEDIA: WEISBORD 279
through which are visible imperfectly seven longitudinal rows, each
of them composed of about eight rather large scales.
Broch (1953) synonymized Scalpellum bellum Pilsbry with
Scalpellum talisman Gruvel, and stated that neither Gruvel nor
Pilsbry “mentioned a membraneous interspace between the strongly
and evenly arched carina and tergum-latus superiorus . . . probably
because this interspace is also lacking in the smaller specimen from
[Ingolf] St. 20.” This gap is clearly shown by Broch on what he
believed to be S. talisman. However, the gap is also at least sug-
gested on the drawing of the type S. vitrewm by Hoek, and is defi-
nitely portrayed on the sketches of S. vitreum by Newman and
Ross.
Type locahty.— “Challenger” sta. 237 (34°37’N, 140°32’E),
off Yeddo, Chiba Prefecture, Japan, depth 1875 fathoms (3429
meters), mud bottom, temperature 1.7°C. The species was also listed
(as Scalpellum formosum Pilsbry) by Weltner in 1922 from Sagami
Bay, Japan, at a depth of 366 meters.
Other localities. — “Eastward” sta. 7617 (33°58.7’N, 45°42’W),
about 125 statute miles east of Fort Fisher, South Carolina, depth
2280 meters. This Western Atlantic occurrence is the nearest one to
Florida; “Albatross” sta. 2205 (39°35’N, 71°18’45”W) south of
Martha’s Vineyard, depth 1073 fathoms (1963 meters), 38.1° bottom
temperature; “Albatross” sta. 2097 (37°56’20’N, 70°57’30”W),
1917 fathoms (3506 meters), Globigerina ooze; Type of S. formosum
Pilsbry, about 230 statute miles east of Chincoteague, Virginia; Cap
Ghir (30°40’N, 9°54’W), Morocco, depth 2125 meters; Golfe de
Gascogne, 4255 meters; Indian Ocean and Malay Archipelago (Nils-
son-Cantell, 1938); “Ingolf” sta. 20 (58°20’N, 40°48’W), southeast
off Cape Farewell, Greenland, 3192 meters, 1.5°C bottom tempera-
ture; “Eltanin” sta. 18 (58°15’N, 48°36’W), southwest off Cape
Farewell, Greenland, depth 3404 to 3422 meters; “Eltanin” sta. 791
(63°54’S, 83°03’W), 4531 meters, off Bryan Coast, Ellsworth Land,
Antarctica.
Arcoscalpellum vitreum and the species synonymized with it by
authors ranges in depth from 366 meters to 4531 meters, ranges
latitudinally from 58° north to 63° south, and occurs in the North-
west Pacific (Japan), the Malay Archipelago, the Indian Ocean, and
in the Eastern and Western Atlantic.
280 BULLETIN 299
Should the Scalpellum sp. cf. bellum of Bayer, Voss and Robins
(1970) be confirmed as one of the synonymous species under Arco-
scalpellum vitrewm, the range of A. vitrewm can be extended to in-
clude “Pillsbury” sta. 345 (9°59.6'N, 77°33’W to 10°11.5’N,
77°2V’W) at depths of 2434-3111 meters and “Pillsbury” sta. 346
(9°54.5’N, 77°03’W to 9°51’N, 76°58’W) at depths of 2983-2970
meters. These stations are in the Caribbean Sea about midway be-
tween Punta San Blas, Panama, and Cartagena, Colombia.
Calantica superba (Pilsbry) Pl. 33, figs. 3a-c
Scalpellum (Calantica) superbum Pilsbry, 1907, pp. 9, 11-13, figs. 3a-c.
Scalpellum [= Calantica] superbum Pilsbry, Zullo, 1968, p. 216.
The capitulum of the type from “Albatross” sta. 2669 is 46 mm
in length and 34 mm in width. The capituli of two specimens from
“Albatross” sta. 2415 measure 35 & 28 mm and 28 X 22 mn, re-
spectively. The capitulum of the type is somewhat triangular, wide
and thick at the base, and is composed of 13 strong white plates with-
out perceptible cuticle. The plates are sculptured with radiating
striae crossed by growth lines.
The tergum is in part concealed under the margins of the scuta
and carina, the visible part divided equally by a median ridge from
apex to base; the summit is erect, only a trifle recurved. The carina
is somewhat curved, its apex not inserted between the terga; the
roof is strongly carinate along the median line, sloping and sculptured
with radial striae on each side of the keel; the sides are narrow and
incurved, and at the base the roof is wide.
The rostrum is triangular, with an incurved apex and a strong
median longitudinal rib. The rostrolateral plate is obliquely triangu-
lar, with incurved apex, the surface sculptured with several coarse
low radial ribs, numerous fine radial striae, and curved coarse radial
growth wrinkles; the base of the plate overlies the adjacent bases
of the rostrum and inframedian latera. There is no subrostrum. The
median lateral plate is oblique, triangular, much wider than high,
its apex incurved and twisted; a strong flat-topped rib runs from
the apex to the basal margin which in the middle rests on the ped-
uncle. The carinal latus is oblique and triangular, its apex curved
under the apices of the inframedian latera. The surface of the carinal
latus is ribbed. The subcarina is triangular, usually asymmetrical,
and with an incurved apex.
FLORIDIAN CIRRIPEDIA: WEISBORD 281
The peduncle of the two specimens from “Albatross” sta. 2415
measures 12 and 15 mm respectively, in length. The peduncle is
covered with large, strongly imbricating white scales.
Type locahty.—“Albatross” sta. 2669 (31°0YN, 79°33’W),
352 fathoms (644 meters), gray sand and dead coral, bottom tem-
perature 43.7°F, about 105 statute miles east off Sea Island, Georgia.
Other localities. —“Albatross” sta. 2415 (30°44’N, 79°26’W),
440 fathoms (805 meters), bottom temperature 45.6°F, on branching
white coral, bottom of coral and coarse sand with shells and
Foraminiferida, about 120 statute miles east off mouth of St. Marys
River, boundary between Georgia and Florida.
Euscalpellum stratum (Aurivillius) Pl. 33, figs. 4-7
Scalpellum stratum Aurivillius, 1893, p. 132; 1894b, pp. 65-67, pl. 3, figs.
10-11, pl. 8, fig. 8; Weltner, 1897, p. 250; Gruvel, 1905, p. 58, fig. 62.
Scalpellum (Smilium) stratum (Aurivillius), Pilsbry, 1907, p. 13.
Euscalpellum stratum (Aurivillius), Kriiger, 1940, p. 82, fig. 84b1; Pilsbry,
1953 apps Zi-23, pl. 1, tigs 7 Wathers 1953.) ps 171:
Aurivillius’ original description in German is translated as
follows:
Diagnosis. Capitulum with 15 calicified plates. Carina simply
arched, the roof flattened below. Subcarina small, equilaterally
triangular. Rostrum extending to the muscle margin of the scuta,
convex, pointed below, widest near the middle where it is almost
angular. Rostrolatera triangular, measuring in length one third of
the rostrum. Laterals and inframedian quadrilateral, their scutal
margins about one third the length of the scutum. The umbo lies in
the lower angle of these plates.
The peduncle is crossed by 14 diagonal rows of about 14-15
thomboidal scales.
Color of specimens in alcohol. Light brown, on the capitulum
only between the margins of the plates, on the peduncle clearly
visible between the rows of scales.
Dimensions. Length of animal 9 mm. Length of capitulum 5.5
mm; breadth across rostrum — 3 mm.
Locality and occurrence. Sea of the Antilles, near Anguilla,
360-680 meters in depth. Various specimens (A. Goés). RM.
The above description was added to by Gruvel (1905) who
stated that the carina was rounded at the base; that the apices of
282 BULLETIN 299
the terga and scuta were pointed and nearly straight; that the infra-
laterals were not in immediate contact with the capitulum; and that
the peduncle was straight, uniform, and about half the length of the
capitulum.
Pilsbry (1953) further remarked that the Florida specimens
were covered with a thin transparent cuticle, and that the species
was separable from all other scalpellid barnacles by the long rostrum.
Type locality. — Near Anguilla (18°14’N, 63°05’W), Leeward
Islands, depth 360-680 meters.
Florida localities. —Off Palm Beach (26°41’N, 80°02’W) at
many “Triton” stations, between 50 and 100 fathoms, most abundant
at about 75 fathoms (137 meters). “Triton” sta. 378, off Boynton
Beach Inlet (26°32’N, 80°04’W), 50 fathoms (91 meters). Off Cape
Florida (southern tip of Key Biscayne), 100 fathoms (183 meters),
on Rochinia crassa.
Lithotrya dorsalis (Ellis and Solander) Pl. 34, figs. 1-3
Lepas dorsalis Ellis and Solander, 1786, p. 197, pl. 15, fig. 5.
Litholepas de Mont Serrat, Blainvilie, 1824, pl., fig. 5. [Fide Darwin, 1851,
pepsolel
Lithotrya dorsalis G. B. Sowerby I, 1822, unnumbered page; Gray, 1825,
p. 101; Darwin, 1851, pp. 335, 336, 341-343, 346, 347, 351-356, 363, pl. 8, figs.
1tian= -1c’; Chenu, 1858, p. 77; Hoek, 1883, p. 29; Weltner, 1897, p. 251; Bige-
low, 19005 p: 179): Grivel 1902a, pp. 249, 255, 259; 1902c, p. 524; 1905, pp.
98-99, fig. 108; 1907, pp. 162-163; Jennings, 1915, p. 287; Pilsbry, 1927, p. 27;
1953, p. 23, pl. 1, fig. 8; Nilsson-Cantell, 1931b, p. 105; 1933, pp. 504-505; 1939,
p. 3; Cannon, 1947, pp. 89, 92, 97; G. L. and N. A. Voss, 1955, pp. 212-213.
Lithotrya dorsalis (Ellis and Solander), Pilsbry, 1907, p. 6; Kriiger, 1940,
pp. 22, 72, 126, 129, 225, 338, 459, figs. 71, 74, 78a-c, 101a, 127; Newell, Im-
brie, Purdy, and Thurber, 1959, pp. 207, 211, fig 12; Newman, Zullo, and
Withers, 1969, p. R272, fig. 115.10; Southward, 1975, p. 3.
Ellis described this species as follows:
Abi, 1S.
Fig. 5. Lepas dorsalis, testa quinquevalvi corpus tegente basi squamosa,
valvulis lateralibus laevibus; dorsali rotundata transversum rugosa, stipite
squamuloso.
From Musquito shore.
The following is taken from Darwin’s description of the hard
parts.
The capitulum is half an inch in width and height; the entire
length of the animal with contracted peduncle is about an inch and
a half. The valves are dirty white, with the enveloping membrane
yellow.
The scuta are triangular, internally concave, with a small
roughened internal knob or tooth at the rostral angle of both valves;
FLORIDIAN CIRRIPEDIA: WEISBORD 283
the tergal margin is straight, overlapping about one third the entire
width of the terga.
The terga are irregularly oval, internally slightly concave, with
straight scutal margins, and with the lower part of the carinal
margin, immediately over the latera, slightly hollowed out. Ex-
teriorly, toward the bottom of the valves, a narrow ridge is exposed,
which runs down to the basal angle at about one third the entire
width of the valve, from the scutal margin.
The carina slightly overlaps the terga; it is internally concave,
generally with a large upper portion freely projecting, without any
central crest or ridge. The carina is nearly as wide as the middle
part of the terga, the inner growing or corium-covered surface, with
its basal margin, protuberant and arched. The dorsum of the carina
is marked by strong concentric growth ridges separated by wider
interspaces.
The rostrum is small and narrow, with deeply sinuous sides and
a rounded basal margin; in width the rostrum equals about two and
a half of the uppermost scales of the peduncle, and about half as wide
as the latera.
The latera are small, oblique, and parallel with the carinal
margin of the terga; the longer axis is equal to five of the uppermost
scales of the peduncle and to nearly half the width of the carina.
The peduncle varies in length, generally twice, but on one of
Darwin’s specimens thrice as long as the capitulum. The upper part
of the peduncle is as wide as the capitulum, the lower part attenu-
ated. The calcified scales in the uppermost whorl are only slightly
larger than those in the second whorl, and the scales in the suc-
ceeding three or four whorls are considerably larger than those be-
low, which latter gradually decrease in size until, low down on the
peduncle they are barely visible to the naked eye. In the lower part
they appear as calcareous beads, standing apart from each other —
smooth, translucent, and furnished with a conical fang. The upper
scales vary somewhat in outline, the most usual shape being triangu-
lar, with the lower margin arched and protuberant; this margin, in
the two or three upper whorls, is crenated with teeth which are first
conical and sharp but after molting become mere notches. The scales
in the uppermost whorls are nearly quadrilateral, the imbedded por-
tion or fang produced into a blunt rounded point.
284 BuLLeTIN 299
The basal calcareous cup of the peduncle is well developed, is
composed of swirling overlapping growth lamellae, and attains a
diameter of as much as half an inch. Before the cup is formed, there
is a column of small flat discs attached to the side of the burrow.
Type locality. — Northeast (Mosquitia) coast of Honduras.
Florida localities. — Sambo Shoals (south of western group of
Florida keys) where L. dorsalis occupies borings in corals or aeolian
rocks; Soldier Key in Biscayne Bay, in intertidal rocks.
Other localities. —Great Bahama Bank, intertidal shoals; San
Salvador; Cuba; Jamaica; Puerto Rico (Ensenada Honda, Culebra,
Aguadilla); Barbados; Venezuela (Rio Tocuyo); Netherlands An-
tilles: Curacao (Westpuntbaai; Caracasbaai); Bonaire (Kralen-
dijk); Klein Bonaire, east coast; Solomon Islands; Philippines;
Chagos Archipelago; Farquhar Atoll.
Neoscalpellum dichelopiax (Pilsbry) Pl. 34. figs. 4a-c
The names applied to this taxon by one author or another are
the following:
Scalpellum debile Aurivillius (1898); Scalpellum edwardsu
Gruvel (1900); Scalpellum dicheloplax Pilsbry (1907); Scalpellum
dicheloplax benthophila Pilsbry (1907); and Scalpellum alboranense
Gruvel (1920). Since Neoscalpellum dicheloplax (Pilsbry) is found
nearest to Florida it is selected as the taxon most appropriate for
this work. It is anticipated that the dictates of biogeography and
details of external and internal morphology will play a role in ulti-
mately determining which of the taxa should be combined and which
separated.
The synonymy proposed by authors is as follows:
Scalpellum debile Aurivillius, 1898, p. 189; 1938, p. 71; Gruvel, 1905, p. 27;
1920, pp. 31-32, 73, pl. 5, figs. 13-15, pl. 7, fig. 1; Nilsson-Cantell, 1955, p. 218;
Belloc, 1959, p. 2; Newman and Ross, 1971, pp. 96-99, text-figs. 49-50, as Neo-
scalpellum.
Scalpellum edwardsii Gruvel, 1900, p. 189; 1902b, p. 63, pl. 2, figs. 3B, 16;
1902c, p. 523; 1905, pp. 28-29, fig. 27; Newman and Ross, 1971, p. 96.
Scalpellum dicheloplax Pilsbry, 1907, pp. 70-73, fig. 28a-c; Fowler, 1912,
p. 500; Hoek, 1914, p. 4; Gruvel, 1920, p. 32, pl. 7, fig. 2; Withers, 1926,
p. 102; 1928, pp. xi, 13, fig. 36, as Neoscalpellum; Zullo, 1968, p. 212 and p. 214,
as Mesoscalpellum; Newman, Zullo, and Withers, 1969, p. R224, fig. 94-11, as
Mesoscalpellum; Newman and Ross, 1971, p. 96, as Neoscalpellum.
Scalpellum alboranense Gruvel, 1920, p. 33, pl. 5, figs. 4-6; Belloc, 1959,
p. 4; Newman and Ross, 1971, p. 96.
TYPE LOCALITIES
The type locality for each of the species enumerated above is
the following:
FLORIDIAN CIRRIPEDIA: WEISBORD 285
Scalpellum debile Aurivillius
“Princesse-Alice” sta. 749 (38°54’N, 23°27’W), 5005 meters,
between Lisbon, Portugal and Terceira Island, Azores.
Scalpellum edwardsu Gruvel
“Talisman” dragage 136 (near the Azores), 4255 meters.
Scalpellum dicheloplax Pilsbry
“Albatross” sta. 2711 (38°5YN, 70°07’W), 1544 fathoms (2824
meters), about 255 statute miles east off Cape May, New Jersey.
Nearest locality to Florida.
Scalpellum dicheloplax benthophila Pilsbry
“Albatross” sta. 2042 (39°33’N, 68°26’45”W), 1555 fathoms
(2844 meters), about 310 statute miles east off Atlantic City, New
Jersey.
Scalpellum alboranense Gruvel
“Prince Monaco” sta. 650 (36°54’N, 20°46’15’”W), 4400 meters,
white foraminiferal ooze, between Lisbon, Portugal and Ponta Del-
gado, Azores.
Pilsbry’s description of his Scalpellum dicheloplax is summarized
as follows:
The capitulum, measuring 44 mm in length and 31 mm in width,
is ovate and strongly compressed, and consists of 13 imperfectly cal-
cified plates, all of them biramose or V-shaped and with a smooth
cuticle.
The tergum is V-shaped, having a curved occludent branch
about twice as long as wide and a long, slender curved carinal branch
about twice the length of the occludent branch. The scutum has a
wide convex occludent segment and a narrow, curved calcified tergal
segment. The surface is sculptured by low, narrow, widely spaced
riblets which are parallel with the basal margin. The occludent
margin is strongly convex. The umbo is terminal at the apex and
overlies the base of the tergum. The carina is strongly arched, more
so above than below, and has a length of 42 mm and a diameter of
4.5 mm at the base. The umbo is turned inward but is not quite
terminal, a flattened continuation of the sides extending beyond it.
The roof is deeply channeled, with high bordering ribs. The sides are
of nearly equal width throughout.
The upper latus is broadly V-shaped in the upper calcified por-
tion, the two branches subequal, somewhat curved, the lower part
286 BULLETIN 299
of the broader half with two prongs. Another slender branch rises at
the apex and runs toward the tergum in a direction at right angles
to the tergal branch. There is no rostrum. The rostral latus is V-
shaped, the basal branch shorter and wider. The inframedian latus
has a calcified portion with a modified wineglass shape, being very
narrow below the middle and expanded at the base, the upper part
composed of two diverging branches, the one directed toward the
scutum the longer of the two. The umbo is at or below the lower
fourth in the adult stage but is higher in the young. The carinal
latus is broadly V-shaped, the carinal branch the larger and curved.
The umbones are recurved and project below and beyond the carina.
The peduncle is 24 mm in length and is clothed with large
strongly projecting scales in 7 rows of about 12 scales each.
Type locality. — “Albatross” sta. 2711 (38°59N, 70°07’W),
1554 fathoms (2842 meters), about 255 statute miles east of Cape
May, New Jersey. Although Florida lies nearer to the type locality
of this taxon than do the other four congeners, it is still remote, lying
8° of latitude to the south and 10° of longitude to the west.
Range and distribution.—In addition to the type localities
specified above, the following localities have been cited for the
Neoscalpellum dicheloplax group:
As Scalpellum dicheloplax: “Michael Sars” sta. 10 (45°26’N,
9°20”’W), 4700 meters, bottom of Globigerina ooze, attached to
stones, lying west of the Bay of Biscay; “Albatross” sta. 2221
(39°05’30”N, 70°44’30”W ), 2788 meters, and “Albatross” sta. 2222
(39°03/15”N, 70°50’45”W), 2810 meters, both stations about 205
statute miles east of Avalon, New Jersey.
As Scalpellum dicheloplax benthophila: “Chain 50” sta. 81
(34°41’N, 66°28’W), 5042 meters and “Atlantis II” sta. 93
(34°39N, 66°26’W), 5007 meters, both stations lying northwest of
Bermuda between Bermuda and Cape Hatteras, North Carolina.
As Scalpellum debile: “Prince Monaco” sta. 652 (46°56’N,
22°22’W ), 4261 meters, white foraminiferal ooze, east of Azores, be-
tween Ponta Delgado and Cabo Sao Vicente, Portugal.
As Scalpellum alboranense: “Prince Monaco” sta. 650 (38°54’N,
21°06’W), 5005 meters, white Globigerina ooze, east off Azores be-
tween Faial and Lisbon, Portugal.
FLoRIDIAN CIRRIPEDIA: WEISBORD 287
Recapitulating, the Scalpellum dicheloplax complex are deep
water taxa ranging in recorded depth from 2788 meters east of
Avalon, New Jersey, to 5042 meters between Cape Hatteras and:
Bermuda. Geographically the farthest north locality is the Bay of
Biscay in the Eastern Atlantic, and the farthest south is the Western
Atlantic between Cape Hatteras and Bermuda, or roughly between
46° north and 34° north. This is a fairly narrow latitudinal zone
of occurrence and lends support for those who advocate the uniting
of the S. dicheloplax complex in the Western Atlantic with the
earlier-named S. debile-edwardsu complex in the Eastern Atlantic.
Neoscalpellum dicheloplax benthophila (Pilsbry) Ri-7a4, fig. 5
Scalpellum dicheloplax benthophila Pilsbry, 1907, p. 73, fig. 28d; Zullo,
1968, p. 211; Newman and Ross, 1971, pp. 96, 97, 99.
Scalpellum dicheloplax bentophila [sic] Pilsbry, Gruvel, 1920, pl. 7, fig. 9;
Nilsson-Cantell, 1938, p. 7.
Scalpellum (Arcoscalpellum) dicheloplax Pilsbry, subsp. benthophila Pils-
bry, Weltner, 1922, p. 67.
Concerning the legitimacy of this subspecies, Pilsbry wrote the
following:
The capitulum is more lengthened than in S. dicheloplax, its length twice
the breadth. The carina is less arcuate with wider sides, and separated from
the tergum by a much narrower chitinous suture. The plates of the lower
whorl are completely calcified, and the inframedian lateral plate is narrower,
with central umbo. The scuta, terga, and upper lateral plates are V-shaped, with
comparatively shorter, wider branches than in S. dicheloplax.
Length of capitulum 15, width 7.5; length of peduncle, 4.5 mm.
The much more extensive calcification of the plates in the single example
of this subspecies, as compared with §. dichcloplax, may be due to youth; but
the narrower shape of the whole capitulum, the narrower inframedian latera,
and the reduction of the chitinous space between carina and tergum are
features which render it advisable te distinguish this form by name. It re-
quires comparison with specimens of S. dicheloplax of equally small size which
are unfortunately not yet in our possession.
The views of other authors concerning the possible synonymy
of the subspecies benthophila with S. dicheloplax Pilsbry and other
earlier-named species are given on the preceding pages under Neo-
scalpellum dicheloplax (Pilsbry).
Type locality. — The type locality of Neoscalpellum dicheloplax
benthophila is “Albatross” sta. 2042 (39°33’N, 68°26’45”W), 1555
fathoms (2844 meters), about 310 statute miles east off Atlantic
City, New Jersey, bottom of Globigerina ooze, bottom temperature
Bhshor| Cs
Other localittes.—“Chain 50” sta. 81 (34°39N, 66°26’W),
288 BULLETIN 299
5042 meters (2757 fathoms) and “Atlantis II” sta. 240 (34°39N,
66°26’W ), 5007 meters (2738 fathoms). These two stations are about
570 statute miles east off Cape Lookout, North Carolina, and lati-
tudinally are about 5° north of Florida waters; “Valdivia” sta. 240
(6°12.9’S, 41°17.3’E), 2959 meters (1618 fathoms), bottom tempera-
ture 2°C, about 140 statute miles northeast of Dar es Salaam;
“Michael Sars” sta. (45°26’N, 9°20’W), 4700 meters (2569 fath-
oms), about 140 statute miles north off La Corufia, Spain, in the
North Atlantic.
Summarizing, Neoscalpellum dicheloplax benthophia ( Pilsbry)
has been reported from the Western Atlantic, North Atlantic, and
Indian Ocean off the east coast of Africa, at depths ranging from
2844 meters to 5042 meters.
Mesoscalpellum imperfectum (Pilsbry) Pl. 33, figs. 8, 9; Pl. 34, fig. 6
Scalpellum imperfectum Pilsbry, 1907, pp. 70, 75-77, fig. 30, pl. 4, figs. 15-
18; Fowler, 1912, p. 500; Annandale, 1913, p. 233; Barnard, 1924, pp. 17, 47;
MacDonald, 1929, pp. 534, 537, pl. 2, fig. 3; Broch, 1953, pp. 9, 10, 12, 15, fig.
12; Stubbings, 1961, pp. 11-13, text-fig. 2; 1967, p. 234; Zullo, 1968, p. 213.
Scalpellum (Mesoscalpellum) imperfectum Pilsbry, 1908, p. 110.
Mesoscalpellum imperfectum (Pilsbry), Newman and Ross, 1971, p. 119,
text-fig. 62.
Pilsbry’s description of the skeletal elements is summarized as
follows:
The capitulum of the type is 29 mm in length, 20 mm in
breadth. It is composed of 13 valves, the upper ones imperfectly cal-
cified, joined by wide chitinous sutures. The occludent and carinal
margins are regularly convex, and the apex is obtuse. The cuticle is
thin and smooth, and the plates are weakly marked with growth
lines.
The tergum is shaped like an inverted V, the two branches some-
what curved, that along the occludent margin truncate at the apical
end, the carinal branch much longer and tapering to an acute lower
end. The apex is strongly recurved and acute, but a chitinous
border projects beyond it along the occludent margin, and there is
a small wing on the carinal side of the apex. The growth lines on
the carinal branch are deeply V-shaped. The scutum is triangular,
narrow above, with a small wing or triangular projection on the
lateral side of the apex. The umbo is apical. The carina, measuring
27 mm in length and 5 mm in width at the base, is widely separated
from the other plates and is abruptly bent at the umbo. The roof
FLORIDIAN CIRRIPEDIA: WEISBORD 289
is flat, bounded by angles or low ribs; the sides are moderately wide
in the middle, narrowing above and below; the base is almost
squarely truncate.
The upper latus is pyriform-polygonal, widely separated from
the other plates. The umbo is at the upper third of the plate. There
is no rostrum. The rostral latus is narrow, the scutal and basal
margins subparallel, the umbones at the upper front angle, and in
contact. The inframedian latus is narrow, tapering somewhat toward
the base which itself is expanded; the umbo is at the lower third.
The carinal latus has the shape of a horn, curved at the apex which
extends well beyond the carina.
The peduncle is 14 mm in length, composed of eight rows of
five scales each.
Type locahty.—“Albatross” sta. 2731 (36°45’N, 74°28’W),
781 fathoms, (1428 meters), attached to Arcoscalpellum velutinum
(Hoek), about 100 statute miles east of Norfolk, Virginia. This is the
locality nearest Florida.
Other locahties. —“Albatross” sta. 2741 (37°44’°N, 73°56’W),
852 fathoms (1558 meters), about 90 statute miles east off Cedar
Island, Virginia; “Albatross” sta. 2196 (39°35’N, 69°44’W), 1230
fathoms (2250 meters), on echinoderm spine, about 255 statute miles
east off Beach Haven, New Jersey; “Pieter Faure” sta., 38 miles
southwest off Cape Point, South Africa, at approximately 34°30’S,
17°45’E, 755 fathoms (1381 meters); “Ingolf” sta. 63 (62°40’/N,
19°05’W), off the south coast of Iceland; “Atlantide” sta. 120
(2°09°N, 9°27’E), 650-260 meters, about 45 statute miles west off
Equatorial Guinea; ? Galapagos Islands in the Eastern Pacific.
The recorded depth range of Mesoscalpellum imperfectum is 260
to 2250 meters (143 to 1230 fathoms). The farthest north record is
Iceland, the farthest south off the southern tip of Africa, thus occur-
ring in polar, temperate, and tropical latitudes of the Western and
Eastern Atlantic. Newman and Ross (1971) stated that Mac-
Donald’s Galapagos locality in the Eastern Pacific is in need of
confirmation.
Mesoscalpellum sp. Bayer, Voss, and Robins
Scalpellum (Mesoscalpellum), n. sp. Bayer, Voss, and Robins, 1970, p. A43.
This taxon, undescribed, was reported by the authors as oc-
290 BULLETIN 299
curring in the Gulf of Mosquitos, Panama, at “Pillsbury” sta. 447
(9°07.4’N, 81°07.4’°W to 9°04’N, 81°13.8’W), depths 664 to 681
meters. “Pillsbury” sta. 447 lies northwest of San Cristébal, Panama,
the coordinates of which are 8°52’N, 80°56’W.
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i-xx, 17-391, pls. I-XIV, wood cuts 1-106, vignettes pp. 72, 159, 322,
375; Vol. 2, pp. i-viii, 9-340, pls. XV-XLII, wood cuts 1-62,
vignettes pp. 48, 57, 171, 231.
Townsend, C. H.
1901. Dredging and other records of the steamer Albatross, with bib-
liography relative to the work of the vessel. U.S. Commiss. Fish
and Fisheries, Rept. U.S. Commissioner Fish and Fisheries for the
fiscal year ending June 30, 1900, pp. 387-562, pls. 1-7.
United States Naval Institute
1967. Marine fouling and its prevention. Prepared for Bureau of Ships,
Navy Department by Woods Hole Oceanographic Institution,
Woods Hole, Massachusetts. Annapolis, Md., 3rd printing July
1967, pp. i-ix, 1-338, figs. tables, pls. [Cirripedia, pp. 121-131, 194-
196.]
Utinomi, Huzio
1958. Studies on the cirripedian fauna of Japan. VII. Cirripeds from
Sagami Bay. Seto Marine Biol. Lab., Publ., vol. VI, No. 3, pp.
281-311, figs. 1-10.
Voss, Gilbert L., and Voss, Nancy A.
1955. An ecological survey of Soldier Key, Biscayne Bay, Florida. Bull.
Marine Sci. Gulf and Caribbean, vol. 5, No. 3, pp. 203-229.
Weisbord, Norman E.
1975. Cirripedia of Florida and surrounding waters (Acrothoracica and
Rhizocephala). Bull. Amer. Paleont., vol. 68, No. 290, pp. 168-233,
pls. 20-28.
Wells, Harry W.
1966. Barnacles of the northeasiern Gulf of Mexico. Florida Acad. Sci.,
Quart. Jour., vol. 29, No. 2, pp. 81-95.
Weltner, Wilhelm
1895. Die Cirripedien von Patagonien, Chile und Juan Fernandez. Arch.
f. Naturgesch., Jahrg. 61, vol. 1, No. 2, pp. 288-292.
1897. Verzeichnis der bisher beschreibenen recenten Cirripedienarten.
Mit Angabe der im berliner Museum vorhandenen Species und
threr Fundorte. Arch. f. Naturgesch., Jahrg. 63, vol. 1, No. 3, pp.
227-280.
1922. Cirripedia der Deutschen Tiefsee-Expedition. In Car] Chun, Wis-
senschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf
dem Damfer “Valdivia”, 1898-1899. Jena, vol. 23, No. 2, pp. 59-
112, text-figs. 1-3, pls. 2-4.
296 BULLETIN 299
Withers, Thomas Henry
1926. A new cirripede from the Upper Cretaceous of Western Australia.
Roy. Soc. Western Australia, Jour., vol. 12, No. 11, pp. 101-104,
text-fig. 2, pl. XI.
1928. Catalogue of fossil Cirripedia in the Department of Geology. Vol.
1. Triassic and Jurassic. British Museum (Natural History), pp.
1-xii, 1-154, pls. 1-12, text-figs. 1-93.
1953. Catalogue of fossil Cirripedia in the Depariment of Geology. Vol.
3. Tertiary. British Museum (Natural History), pp. 1-xv, 1-396,
pls. 1-64, text-figs. 1-105.
Zevina, G. B.
1973a. Scalpellidae (Cirripedia) from the Indian Ocean. I. Species of
subgenera Scalpellum and Arcoscalpellum of the genus Scalpellum.
Zool. Zhur., vol. 52, No. 6, pp. 842-848, figs. 1-19. [In Russian with
English summary. |
1973b. Scalpellidae (Cirripedia) from the Indian Ocean. 2. Species of the
subgenera Annandaleum, Mesoscalpellum and Neoscalpellum of
the genus Scalpellum. Zool. Zhur., vol. 52, No. 7, pp. 1000-1007,
figs. 1-27. [In Russian with English summary. |
Zullo, Victor August
1966. Thoracic Cirripedia from the continental shelf off South Carolina,
U.S.A. Crustaceana, vol. 11, pt. 3, pp. 229-244, figs. 1-7.
1968. Catalog of the Cirripedia named by Henry A. Pilsbry. Acad. Nat.
Sci., Philadeiphia, Proc., vol. 120, pp. 209-235, pls. 1-5.
FLorIDIAN CIRRIPEDIA: WEISBORD 297
PARES
298 BULLETIN 299
EXPLANATION OF PLATE 26
Figure Page
1. Scalpellum (?) albatrossianum PilSDIy ..............cccccsscsscceceeeseeeceeeeees 237
Holotype X 4. Dimensions: capitulum 10.5 mm X 5.3 mm; peduncle
length 3.5 mm; carina length 9 mm; diameter at base 1.7 mm.
2; scalpetlum) (2) antillaruml BilSbTy, cccccccccseeectsesecsecseeeesee ee 240
2a. Holotype X 3. Dimensions: capitulum 11 mm X 5.7 mm; carina
length 4.5 mm; diameter at base 1.1 mm. 2b. Details of rostrum.
2c. Details of carina.
3-0. ‘Scalpellum: arietinum PilsSDry «scccccccssscsccscs000s<sccsscosvccacessvsuceosseeceeenee 242
3a,b. Holotype. 3a. Lateral view X 3; 3b. Dorsal view X 4. Dimen-
sions: capitulum 11.6 mm X 7 mm; peduncle length 6 mm; carina
length 12 mm; diameter near base 1.7 mm. 4,5. Specimens from
off Palm Beach, Florida. 4. Lateral view of whole animal,
dimensions not given. 5a. Mandible; 5b. Maxilla. 5c. Protopo-
dite of cirrus VI, penis, and caudal appendage. 5d. Segments of
cirrus I. 5e. Segment of cirrus V. 5f. Palpus.
BULL. AMER. PALEONT., VOL. 72 PLATE 26
PLATE 27
BULL. AMER. PALEONT., VOL. 72
FLORIDIAN CIRRIPEDIA: WEISBORD 299
EXPLANATION. OF PLATE 27
Figure Page
i PArcoscalpellum aUrivillit CPUISDIY) Mic. siecceccexcccetessenseceseessassossrosestecese 267
la,b. Holotype, lateral views X 3. Dimensions: capitulum 15.3 mm
< 7.5 mm; peduncle length 5 mm; carina length 13.5 mm,
diameter at base 2.2 mm.
2. Arcoscalpellum aurivillii incertum (PilSbry) .............. ccc eeeeee 268
2c. Holotype X 2, lateral view. Dimensions: capitulum 24 mm X
13 mm; peduncle length 7.5 mm; carina length 22 mm, diameter
at base 3 mm.
Bat eeO Cal pellUMNCaniNatUligblOCk. cccc.cesccscesoer esse oete saeco nore casera oeneoaaee 243
Holotype X 3. 3. Lateral view. 4. Carinal view. Dimensions: cap-
itulum length 16 mm; peduncle length about 6 mm. From Hoek
(1883).
HaGweocalpellum) (?) GiceratUM Ev SDiY: cicccc-seccecccesescrercceeteeeece teens 244
5a,b. Lateral and carinal views of holotype X 3. Dimensions: cap-
itulum 13.5 mm X 7.8 mm; peduncle length 7 mm; carina length
13.5 mm, diameter near base 2.8 mm. 6. Specimen from off Palm
Beach, Florida. 6g. Third tooth and lower point of mandible. 6h.
Base of cirrus VI, penis, and caudal appendage.
igmocalpellum gibbum) Pilsbry:::20803.5.328 2805005. 6.8 onc hss ee ee ress 246
Holotype. 7a. Lateral view X 4.5. 7b. Detail of rostrum. Dimen-
sions: capitulum 7 mm X 4 mm; peduncle length 2 mm.
300 BULLETIN 299
EXPLANATION OF PLATE 28
Figure Page
f. Scalpellumgibbum Pusbry ...,.¢..:¢cc6.c56sectbess2hcb-oncescdnnsto ee 246
Hypotypes collected off South Carolina by Zullo (1966b). 1A
Adult. 1B. Juvenile.
2a. scalpellum)(?) giganteum (Gruvelieessi.ssrsers sc scssccceccssscesedoneeeeeeeee 247
2. Gruvel’s types from off Cuba, depth 500 fathoms, x about ¥%.
Dimensions: capitulum 45 mm X 32 mm; peduncle 45 mm X 15
mm. 3. Specimen from fishing banks off Maine? Dimensions:
capitulum 47 mm X 35 mm X 20 mm; peduncle length about 37
mm. After Pilsbry (1907).
4. Scalpellum (?) gorgoniophilum Pilsbry ...............cccccccssccecssseseesseseees 249
Holotype. 4a. Lateral view X 4. 4b. Detail of rostrum. Dimensions:
capitulum 9 mm X 5 mm; carina Jength 6.8 mm, diameter near
base 1.8 mm; peduncle length 2.8 mm.
D2, scaipellum (2) Qrachlis PHSDIY <..:.<:-0c0<-.ncscosseosccseseocsssecuee se eee 250
Holotype. 5a. Lateral view X 4. 5b. Details of rostrum. 5c. Details
of carina. Dimensions: capitulum 8 mm X 3.3 mm; carina length
6.3 mm, diameter at base 1 mm; peduncle length 18 mm.
6. (Scalpellum, henderson Piulsbry -2-).2).05..<<0<0kccceceece--c<--ctne-xcl ee 251
Holotype X 10. Dimensions: capitulum 5 mm X 2.5 mm; carina
length 3.75 mm.
BULL. AMER. PALEONT., VOL. 72 PLATE 28
PLATE 29
VOL. 72
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BULL. AMER. PALEONT.
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Pi
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FLoRIDIAN CIRRIPEDIA: WEISBORD
EXPLANATION OF PLATE 29
Scalpellum (?) microceros MacDonald. 2...............:s:cccccceeessssssseeesseees
Holotype. Capitulum 31 mm X 23 mm; peduncle about 32 mm in
length, width near base 14 mm.
Seal Pelluini- (HOP a PTSD Ys ies eR cack sale eiete ee sade lh dodhaaebesassnecses
Holotype. 2a. Lateral view X 2. 2b. Rostral view. 2c. Dorsal view.
Dimensions: capitulum 18 mm X 9 mm; peduncle length 4.3 mm;
carina length 15 mm, diameter at base 3 mm.
Scalpellum (?) longicarimatum PilSDry ...............ccccccesscccceeesssssesesees
Holotype X 5. 3a. Lateral view. 3b. Carina] view. 3c. Rostral view.
Dimensions: capitulum 10 mm X 5.4 mm; peduncle length 2 mm;
carina length 9.5 mm, diameter at base 2 mm.
ScalpellUm CA) mMICHUMPe MSDs oe cecccscecccectccescesecencencereeercccceerse tates
4. Lateral view of holotype x 8. 4b. Detail view of rostrum.
Dimensions: capitulum 5 mm X 2.5 mm; peduncle length 1.3 mm;
carina length 3.2 mm.
Scalpellum (?) portoricanum PilSDry ...............cccccccceeseeceeeeseeseeseeees
Holotype. 5a. Rostral view. 5b. Lateral view X 2.7 approximately.
5c. Carinal view. Dimensions: capitulum 12 mm X 7.7 mm;
peduncle length 7 mm; carina length 11.3 mm, diameter at base
2.2 mm.
Scalpellum (?) intonsum (PilSDIy) ..................cescccccceceseeesereccceeceeeeeees
Type. 6d. Detail of rostrum. 6e. Lateral view xX 4. Dimensions:
length of three different capitula 9.5 mm, 9.7 mm, and 7 mm.
301
302 BULLETIN 299
EXPLANATION OF PLATE 30
Figure Page
1. Scalpellum (?) pentacrinarum PilSDIy .................ccccsssscerccccssessnceee 257
Holotype. 1a. Lateral view 6. 1b. Dorsal view. 1c. Ventral view.
Dimensions: capitulum 8 mm X 3.7 mm; peduncle length 3.7
mm; carina length 5.2 mm, diameter near base 1.2 mm.
2. SCalBellUni, PressSUM PIISDILY <...isccccc....0..cséecsecnsseceseansnsnextereneeee omen 260
Holotype. 2a. Lateral view X 5. 2b. Ventral view. Dimensions:
capitulum 8 mm X 4 mm X 1.8 mm; peduncle length 3 mm.
3,4. Scalpellum prunulum AUurivillius ..............:.:s<saccscassececcsaeeddceoouseeeeeneen 262
Holotype. 3(3). Lateral view of capitulum. 4(4). Capitulum from
below. Dimensions: length of whole specimen 6 mm; capitulum 4
mm X 2.5 mm.
5s Scalpellum’ (2) pteryges MacDonald ..c..2:seccescessesscccceseeeeetee eee 263
Holotype. Dimensions: capitulum 23 * 15 mm; peduncle length 10
mm, about 13 rows of scales.
6. Arcoscalpellum regina (PIsDIry)) cccccsicsccccecceciesce-deecsacscssaccenostagenteeeeer 269
Holotype. 6(4). Diagrammatic section of carina. 6(5). Detail of
base of carina. 6(6). Lateral view. Dimensions: capitulum 43
mm X 30 mm; peduncle length 26 mm; carina length 40 mm,
diameter at base 6 mm.
fe scalpellum) (2) sinUatUml Ell slytiyaecers..cssccscccnersercessetrecceeeseteeeeecee eee 265
7a. Holotype, lateral view X 3. 7b. Rostral view. 7c. Topotype
>< 3. Dimensions of holotype: capitulum 13 mm X 7 mm; ped-
uncle length 5 mm.
PLATE 30
BULL. AMER. PALEONT., VOL. 72
BULL. AMER. PALEONT., VOL. 72
PLATE 31
perars i. :
SY
FLORIDIAN CIRRIPEDIA: WEISBORD 303
EXPLANATION OF PLATE 31
Figure Page
1-5. Arcoscalpellum regium (ThoMSoONn) .................cccseesccceeeeesssessseeeceesees 271
1. Holotype, female. Dimensions: jength 60 mm; capitulum 40 mm
xX 25 mm; peduncle length 20 mm; 1a, males lodged within edge
of scutum. 1(8). Diagrammatic section of carina, after Pilsbry
(1970). 2. Male of Arcoscalpellum regium X 20. 3(3). Lateral
view of young specimens, natural size, after Hoek (1883). 4(4).
Lateral view of fully grown specimen, natural size, after Hoek
(1883). 5(5). Carinal view of fully grown specimen, natural size,
after Hoek (1883).
Ga Seal pelluni latidorsGm CElSPLy.) 2.2..ss<...s00cccdecesecees ot-0-4setbes cteedeasesee 253
Holotype. 6(2). Dorsal view, natural size. 6(3). Lateral view,
natural size. Dimensions: length of 6(3) 70 mm; capitulum 42
mm X 28.5 mm; carina length of 6(2) 38 mm, diameter near
base 11 mm. 6(7). Diagrammatic section of carina.
7. Scalpellum (?) semisculptum Pilsbry .............ccccccccesscessceescsesscesesesees 264
7a. Holotype <X 3. 7b. Details of rostrum. 7c. Details of carina.
Dimensions: capitulum 16 mm X 7.7 mm; peduncle length 3 mm;
carina length 12.5 mm, diameter at base 2.5 mm.
304 BULLETIN 299
EXPLANATION OF PLATE 32
Figure Page
1,2. Arcoscalpellum velutinum (Hoek) ..................csscsccssssssccsssssssessesssens eB:
Holotype. 1(10). Lateral view, natural size. 2 (11). Carinal view,
natural size. Dimensions: capitulum length 33 mm; peduncle
length 12 mm.
3,4. Arcoscalpellum eximium (HOeK) «....20.....:..0.2..:0cheactssacaneshenentarstastns 273
Holotype. 3(6). Lateral view, natural size. 4(7). Carinal view
natural size. Dimensions: capitulum length about 43 mm; ped-
uncle length nearly 20 mm.
5. Arcoscalpellum michelottianum (SeguenZa) ...............ccceeeseeeeeeeeees 273
“Types”. All figures except 5(25) enlarged more or less.
§(15,15a). Inner and outer faces of scutum, from the upper
Zanclean of Salice, Messina Province. 5(16,16a). Inner and outer
faces of tergum from the same locality. 5(17,17a,17b). Carina
from the same location and terrane, shown in three positions.
5(17c). Section of same. 5(18,18a). Upper latus from same
locality and terrane, showing inner and outer faces. 5(19,19a).
Carinal latus found at Scoppo, near Messina, in the upper
Zanclean Marl, showing external and internal surfaces; 5(20).
Rostral latus showing outer surface, found in the upper Zanclean
of Salice. 5(21). Rostral latus showing inner surface, from the
upper marl of the Zanclean at Gravitelli near Messina.
5 (22,23,24). Scales of three diverse forms belonging to the ped-
uncle of the same species. Found in great abundance in the sand
of the upper Zanclean at Salice, together with a large number
of other valves. 5(25). Completely imaginative representation
and of natural size, of the mode of attachment of all the plates
of S. michelottianum, together with peduncular scales; or a
restoration of said species. (After Seguenza, 1876).
PLATE 32
BULL. AMER. PALEONT., VOL. 72
peepee TTT Meee ytd Bas tr fs Von OAT?
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PLATE 33
BuLu. AMER. PALEONT., VOL. 72
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Tae ge a
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Lat
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ATT
FLORIDIAN CIRRIPEDIA: WEISBORD 305
EXPLANATION OF PLATE 33
Figure Page
tee Arcoscalpellumi Vith@Uinig (HOCK) trees. cccccecsosscaccess ones sascscsesence coceccess sce 278
Holotype X 6, lateral view. Dimensions: capitulum length 13.5
mm; peduncle length 3.5 mm.
2. Arcoscalpellum talismani Gruvel .....................ccccccccccccscsssscessseesesseee 278
Holotype, redrawn by Gruvel (1905). Dimensions: capitulum 18
mm Xx 9 mm; peduncle length 3.5 mm, diameter 4.5 mm.
See Cal aiticd SUPCG Ba: CE TSI) rice cccsenstenssecces sss deoceacoesstesoesesitenesonctoverucscies 280
Holotype, natural size. 3a. Ventral view. 3b. Lateral view. 3c.
Dorsal view. Dimensions: capitulum 46 mm X 34 mm; length of
visible part of carina 38 mm, diameter 14 mm; peduncle short.
4-7. Euscalpellum stratum (AUuriVillius) .0.....00.........cccceseccsscceseesesseeeccceeeee 281
4. Lateral view of holotype. 5. Capitulum from below. 6. Anterior
maxilla. 7. Specimen from Palm Beach, Florida (Pilsbry, 1953).
7R. Front view of rostrum.
8,9. Mesoscalpellum imperfectum (PUISDIy) ..........0000cccccccceceeesceeceeseeeeeeee 288
8. Holotype xX 1.5, lateral view. Dimensions: capitulum 29 mm
x 20 mm; peduncle length 14 mm; carina length 27 mm, diam-
eter at base 5 mm. 9. Specimen from off Galapagos (MacDonald,
1929), lateral view, natural size.
306 BULLETIN 299
EXPLANATION OF PLATE 34
Figure
133, Lithotrya-dorsalis: (llis and ‘Solander)/e2c2..ccce ee ee
1.. Holotype. 2(1) - 2(1a’,1b’,1c’). Specimens figured by Darwin
(1851) and described as follows: 2(1). Rock bored in two direc-
tions, X 2. 2(1a’). Animal with basal cup adherent, xX 2.
2(1b’). Greatly enlarged, showing rostrum and rostral corners
of the two scuta, together with a small portion of the subjacent
membrane of the peduncle with its calcareous scales and in-
ferior crenated edges. 2(1c’). Basal calcareous cup, x 1.5. 3.
Specimen from Sambo Shoals, Florida, length 35 mm (Pilsbry,
1953).
Disregard figure 8.
4. Neoscalpellum dicheloplax (PilSbry) .0.....0.........cccssesccccsessseesseeeceseeeee
4a,b. Lateral and carinal views of holotype, natural size. Dimen-
sions: capitulum 44 mm X 31 mm; peduncle length 24 mm;
carina length 42 mm, diameter at base 4.5 mm; 4c, young indi-
vidual, natural size.
rc
5. Neoscalpellum dicheloplax benthophila (Pilsbry) ........................
5d. Holotype * 2. Dimensions: capitulum 15 mm X 7.5 mm; ped-
uncle length 4.5 mm.
6. Mesoscalpellum imperfectum (PIISDIy) ..............cccccecccececssecceeceeeeeeeee
6(15). Intermediate segment from sixth cirrus. 6(16). Terminal
appendage. 6(17). Mandible. 6(18). Lower tooth and lower point
of same more enlarged.
PLATE 34
BULL. AMER. PALEONT., VOL. 72
pa A
> eee
ae
5 7, ha
rea
-
INDEX
NUMBER 299
Note: Light face type refers to page numbers. Bold face type refers
to plate numbers.
A
Aden-Zanzibar cable .. 277
Aequipecten gibbus .. 243
JNDGUON sonia ua eeerrrn er 288, 276
Aguadilla,
PUeTLO RICO sesass<cc2.2 284
alatum, Scalpellum .... 273, 274, 277
albatrossianum,
?Scalpellum ............ 237, 238, 240
alboranense,
Scalpellum’ <22.2.:2:..0<. 284
Alligator Point,
TEOTRIG Ey aa eacca none 243
Anguilla, Leeward
TISTENTIVGIS) vcscactnareceeocbee: 282
PATA GATCEIC@ As a ccreebtrssceseses 279
antillarum,
Zscalpellum ......c.<2 238, 240
Arcoscalpellum
MUTATION eeeeeeseses ees 32 239, 274, 275
michelottianum ..32 235, 239, 273,
274
ME OUND te tees a 30 239, 269
OSU teeeccooscceevacsec 31 239, 271
Ciersinuatun es. 265
GalanS mn aeeeeccceees ee 33 239, 278
velutinum. ............ 32 239, 244, 248,
268, 273, 274,
289
VATE UIs ocececsse: 33 239, 278
arietinum,
Scealpellum .......... 26 238, 246
ASHIANE STAGE! .c...<ccs-s.- 274
Atlantic City,
New Jersey .............. 254, 285, 287
aurivillii, Arco-
scalpellum .............. 239, 268
aurivillii incertum,
Areoscalpellum ...... 239, 268
Avalon, New Jersey .. 286, 287
Azores) Islands) <..ca..- PALSY, PAD, PATH
285
B
Bali, Island of,
IMGOMESTA™ css ce-ccstesec OATH
Barbados) essere 284
Bayport, Florida ........ 260, 265
Bay of Bengal, India .. 240
Bays Of BISCAY e-.-e 286, 287
Beach Haven,
New Jersey .............. 289
Belén, Gulf of
Mosquitos, Panama.. 260
bellum, Scalpellum .... 278, 279
Bermudas cee 286, 287
Biloxi, Mississippi ...... 270
Biscayne Bay, Florida 284
Bonaire, Netherlands
ATEN CS eases tavte ces. 284
Boynton Beach Inlet,
TIOTEIGED cocsocnsnossenseeetec 282
183 Al Ne a 272
Bryan Coast,
ANCA CtIC aes eee 279
Cc
Cabo Sao Vicente,
Portigalaeeses ee 274, 276, 277,
286
Cap Cantin, Morocco .. Patel
Cap Noun, Morocco .... 277
Cape
Canaveral, Florida .. 248
Finisterre, Spain .... 273
May, New Jersey .... 254, 285
Farewell, Greenland 279
Florida, Florida ...... 245, 282
Hatteras, North
Carolina. sc.:-secee 244, 246, 247
272, 286, 287
Lookout, North
Carolina: he: ee. 247, 272, 288
Point, South Africa.. 266, 289
Race, New-
foundland .......c:.:..-. 272
Sable, Nova Scotia.. 275
St. George, Florida.. 243
Cape Town,
South Africa ............ 277
Calantica
SUperbale..ee 33 239, 280
Canary Islands ............ 277
Caracasbaai,
CuracaOie ee 285
Caribbean Sea ............ 235, 248, 250,
253, 270
carinatum,
Scealpellum .......... 27 238, 243
Cartagena, Colombia .. 253
Cedar Island,
Mireiniante. 2%. 2.......0; 289
Chagos Archipelago .. 284
Charlestown, Nevis .... 251
Chincoteague,
WAPI Acct 278
307
INDEX
Cidaris tribuloides ...... 243
Colombiae eee 270, 271
Covenas, Colombia .... 246, 248
Crescent Beach,
South Carolina ...... 276
Cubase ees 284
Culebra, Puerto Rico.. 284
Curacao, Netherlands
PATTIE GS errr css 284
Currituck, North
Carolinawee te ee 268
D
Dar es Salaam ............ 288
debile, Scalpellum .... 284
Delaware Bay .............. 266
Destin, Florida ............ 265
Diamond Shoals,
North Carolina ........ 247
diceratum, ?Scal-
pelle ee 27 238, 244
dicheloplax,
Neoscalpellum ....34 239, 284
dicheloplax bentho-
phila, Neoscalpel-
UI 2 116.0 pet eee oe ee re ie 34 239, 284, 287
dorsalis, epas' <....c5-., 282
dorsalis,
aithotryai yess. 34 239, 282
E
East London,
SouthvAtricaye 256
Eastern Atlantic
Ocean. hese 244, 266, 287,
289
Eastern Pacific
OCCAN Seeks 289
Ecphora Zone .............. 235, 247
edwardsii,
Sealpellum .............. 284
Ellsworthland,
PN CUETO), cerppcasconccene 279
Ensenada Honda,
IPUCELO RICO ene 284
Equatorial Guinea ...... 289
erectum, Scalpellum .. 273, 274, 275
Euscalpellum
Stratum) -220-22--- 33 239, 281
eximium, Arcoscal-
ELUM. oii eeeeeeees: 32 239, 273, 274,
275
F
Faial Island, Azores .. 277, 286
Farquhar’ Atoll’ 2:2... 284
Folly Beach,
South Carolina ........ 251
formosum,
Sealpellum <.......:...- 278
Fort Fisher,
South Carolina ........ 279
Sumter, South
Carolinawes sere: 277
G
Galapagos Islands ...... 289
Georgia” 2a eee 2505 201, 201.
281
gibbum, Scalpel-
Ue 27,28 235, 238, 246
gibbus, Aequipecten .. 243
Gibraltar yee. 277
giganteum, ?Scalpel-
WWM cies cee 2 238, 247
Godthaab, Greenland.. 240, 268
Golfe de Gascogne,
TANCE) ees 279
gorgoniophilum,
?Scalpellum ........ 28 238, 249
gracilius,
?Scalpellum ........ 28 238, 250
Megalasma
(Glyptelasma) .......... 248
Grand Bank,
Newfoundland ........ 272
Gravitelli, Sicily .......... 274
Great Bahama Bank .. 284
Greenland. 42.7. 240, 268, 276,
279
Gulf Beach, Florida .. 270
Gulf of
A GON see eee 277
Darien 22. 260
MEXICON ie eee 235, 241, 243,
245, 247, 260,
265, 270, 271
Mosquitos, Panama.. 260, 290
H
Ealbanass Culpaliereceree ese 245, 246, 258
hendersoni,
Scalpellum .......... 28 238, 251
Homosassa, Florida .... 243, 245
HEONGUGAS) fee eee 284
I
Neeland)3 20.3. eee 240, 244, 265,
289
idioplax,
Sealpellum ........... 29 238, 252
imperfectum, Neoscal-
pellum’ .<....:..:: 33, 34 239, 288
308
INDEX
imperfectum,
Scallpe lita) <itc.cscese 243
1OaYG WE rs saan eoeeereeeeeeee 277
Indian Ocean .............. 276, 279, 288
UINGOMES Tatere see rees sneezes ZO 208)
intonsum,
?Scalpellum ........ 29 238, 260
Ibe ENaC | aha eee FAT
USL ap BAT Ueicc.vsssesssssewsses 270
Mtalivaeeene tier ec cectes 274
J
Jackson Bluff
HOnMaAtlOMees ese eceeeee 235, 247
AJEUTE NER) ceed BaAsasaneecoreer 284
K
Kap Farvel,
Greenland) ve.ccssseceees- 266
Kerguelen
Archipelago ............ 276
Key
Biscayne, Florida .. 282
West, Florida .......... 243, 250, 251,
259
Klein Bonaire,
Netherlands
PATIENTS peeeecn teeesee 284
Kralendijk, Bonaire .. 284
L
La Coruna, Spain ...... 288
latidorsum,
Sealpellum. .....:....: 31 238, 253
Le Have Bank ............ 261
Leon County, Florida.. 235, 247
Lepas dorsalis ............ 282
Lisbon, Portugal ........ 244, 277, 285,
286
Lithotrya
@MOTSANS “a.kcc2es0200295 34 239, 282
longicarinatum,
?Scealpellum ........ 29 238, 254
Los Pilones,
Spanish Sahara ...... 277
M
Malay Archipelago .... 279
Marineland, Florida .. 255
Martha’s Vineyard .... 262, 268, 279
Massachusetts ............. 273
Mayagiiez,
Puerto TRICO 2225!.345: 260
Mediterranean Sea .... PATA CALITL
Megalasma (Glypte-
lasma) gracilius ...... 248
Melbourne Beach,
Ploriday 6:.28 22! 253
Mesoscalpellum imper-
fECtuIMiye..-.-.. 33, 34 239, 288
Mesoscalpellum, n. sp. 289
Messina Province,
Sicilliype Macestese orcessss 274
michelottianum, Arco-
scalpellum .......... 32 235, 239, 274,
239
microceros,
?Scalpellum ........ 29 238, 255
micrum,
?Scalpellum ........ 29 238, 256
Mississippi River...... .. 247
Mississippi,
Stategotes.\ ccs ee 270
Mobile, Alabama ........ 247
Mogadishu, Somalia .. 277
MONACO) seer essecs eter 277
Montauk Point,
ING W WORK eee eesccccees 268
Moresby Island,
Canadas ee 269
MOROCCO esse eens PALSY PACT
Mosquitia Coast,
ELOMGUTAS ese 284
N
Nantucket,
Massachusetts ........ 273
Neoscalpellum
dicheloplax .......... 34 239, 284
dicheloplax
benthophila ........34 238, 284, 287
Netherlands Antilles.. 284
Newfoundland ............ 249, 272, 275,
276
INGW2 JiGTSCY, cc.ccccccecesccess 248, 254, 268,
285, 286, 287,
289
New Orleans,
Lousianay |] 241
Newport News,
WARTEABIUEY ceresntereosnsnonece 240, 273
INE Wwe vork. =.ce see 268
Nicobar Island, India.. 277
Nightingale Island . 275
Norfolk, Virginia ........ 244, 289
North Atlantic Ocean 268, 288
North Carolina...) 244, 246, 247,
268, 272, 286,
287
INN SGOWE eapsstoxes 273
Oo
obesum, Scalpellum .. 262
Ocean City,
New Jersey .............. 254
309
INDEX
Oman, Gulf Of, ...:5.s.00 276, 277
Orange Beach,
LUG TEKG Re Sees Acetate 270
P
Palm Beach, Florida .. 243, 245, 247,
260
Palmer,
Katherine V. W. .... 230
Panacea, Florida ........ 270
[eR ITETTIEy Ceoecos NCO eaCereE 248, 253, 260,
290
Pascagoula,
WOSSISSIPDL socc.csc050552 270
Pensacola, Florida ...... 241, 270
pentacrinarum,
?Sealpellum ........ 30 238, 257
Pernambuco, Brazil .... PAL (2
Philippine Islands ...... 284
Ercan vAZOLeS) sicccesscescece 275
Plaisancian Stage ...... 274
Point Pleasant,
New Jersey ............+- 268
Ponta Delgado,
FAZOTES® | Be ricsccesssstascie 285, 286
portoricanum,
?Scalpellum ........ 29 238, 259
POTUICAN Gear cccwweseeceneses 274, 276, 277,
286
pressum,
Scalpellum, .....-.:.:. 30 238, 260
prunulum,
Sealpellum .......... 30 238, 262
pteryges,
?Scalpellum ........ 30 238, 263
IPM EELOVRACO et eeeeeesesence=es 284
Punta Mosquito,
IPAM aM Aer eee 248, 253
Punta San Blas,
IPAM wo cccscs cccessssees 248, 253
R
Racoon Key,
South Carolina ...... 247
Regssio, Waly. -...-.-:...... 274
regina,
Arcoscalpellum ..30 239, 269
regium,
Arcoscalpellum ..31 239, 271
Reykjavik, Iceland .... 240, 244, 265
Riohacha, Colombia .. 271
Rio Tocuyo,
WEnezZUGla «...:..05..2... 284
S
Sagami Bay, Japan .... 279
St. Andrews, Florida .. 260
St. Andrews Sound,
Georgia -:c.85
St lucia island!
St. Mary’s River,
Georgia-Florida ......
Salices tala cee
Sambo Shoals,
OTIGAL cc.seeesce eee
Panama 3.2.
San Juan,
Puerto vRicor..e
Sane Salvador ee
Santa Marta,
Colombpiaye ee
Sasardi Viejo,
Panama ets. -eeeee
Sawu, Indonesia ..........
Sealpeliim 2)
Sealpellum
alatims 2 eee
255, 257, 261
25
299, 281
274
284, 307
290
260
284
270
260
277
236
273, 274, 277
albatrossianum ....26 237, 238, 240,
alboranense ............
antellarimal ee-c.eee 26
arietinum!) =2..2 26
ora auuii pes pesssscox 27
aurivillii
INCcertumMe + eee 27
aurivillii
bellume.co... ee
GarinatwM oes 27
debpilet 2 2. aw
diceratumees 27
edwacdsiliy
erectum ©.....2545-455
fOrMOSUM) 2.
Feallo|oybb000 cpceneeee 27, 28
giganteum ............ 28
gorgoniophilum ..28
sTracwius) se 28
hendersoni .......... 28
1diOplax: 4 o/s 29
imperfectum ............
NCCT UUM scene 27
intonsum) 2......... 29
latidorsum) 22 31
longicarinatum ....29
ODESUNDDE Feeeccsetecest
microceros .......... 29
MITC RUT eevsces eeeeees 29
pentacrinarum ....30
portoricanum ....... 29
DECSSUMN seeeececrscseaee 30
pranulum) <....68 30
DLCLYVEES) ce esecesesanee 30
semisculptum ...... 31
septentrionale ........
273
284
238, 240
238, 242, 246
239, 267
239, 268
278, 279
273, 274, 275
278
235, 238, 246
238, 247
238, 249
238, 250
238, 251
238, 252
243
239, 268
238, 260
238, 253
238, 254
262
238, 255
238, 256
238, 257
238, 259
238, 260
238, 262
238, 263
239, 264
262
INDEX
SUT ALUM aeeeeesseereee 30 239, 265
SOMGUGUIN eee eeerreeeee 273, 274, 275
(SPECIES?) vecceecesssse-ee 259
SUrIOlAtUIM seeeeeeeees ee 23
SETO CMM eee reece 260, 262
LEMUC Ween 237
Sea Island, Georgia .... 281
semisculptum,
?Scealpellum ........ 31 239, 264
septentrionale,
Scalpellumy 262
SHC et eee 274
Sidi Moussa, Morocco.. aT
cf. sinuatum,
Arcoscalpellum ...... 265
sinuatum,
?Sealpellum ........ 30 239, 265
Socatra Island ............ PAT
Soldier Key, Florida.. 284
Solomon Islands ........ 284
SOMA meee eee PAM |
Sombrero Key Light,
LOGIC Ame eee 245
sordidum,
Sealpellum) 2... 273, 274, 275
SOPPOw SICHly, 2. 274
SON Ataarel eae PAT
South Atlantic Ocean.. 277
South Carolinas... 247, 251, 275,
276, 277, 279
Sas ee oe cokes teases 288
Straits of Florida ........ 246
stratum,
Euscalpellum ...... 33 239, 281
stroemii, Scalpellum .. 260, 262
Sumba, Island of,
INGONESIA: ...56<c055-000025 277
superba,
Calanticay.-.. 33 239, 280
Surf City,
New Jersey .............. 248
1
talismani,
Arcoscalpellum ..33 239, 278
tenue, Scalpellum ...... 2a
Terceira Island,
INT ODOS mre eee ecssorans 285
Trapani, Sicily ............ 274
tribuloides, Cidaris .... 243
Tristan da Cunha ...... 244, 274, 275,
276, 277
Trivandrum, India ...... 277
Vv
velutinum,
Arcoscalpellum 32 239, 244, 273,
274
Venezuela 284
\WATHEAUINE) asssonaceecosccocaaneong 240, 244, 273,
279, 289
vitreum,
Arcoscalpellum ..33 239, 278
WwW
Western Atlantic
OCCAMB Ri viscstsecteeess 235, 240, 253
257, 262, 268,
PHL, PASTS OA).
287, 289
Westpuntbaai,
Curacao ee 284
iY
Yeddo, Japan .............. 279
Z
Zanclian Stage ............ 274
cx 1 wirwetu
4 (reat mt
hiieel
’ > irae
7 ) : I,
+
nie
LXVI.
LXVII.
LXVIII.
LXTX.
LXX.
LXXI.
LXXiII.
(Nos. 233, 236). SIRIA AD DEA SUD ISie decsecotecene tasasvesiceecrsuccccemcetas tases
New Zealand forams, Stromatoporoidea, Indo-Pacific, Mio-
cene-Pliocene California forams.
(Nos. 237-238). AR BD es 4D) DSs, (cet Seccreascusctstecctee sneak tse teencoseceson
Venezuela Bryozoa, Kinderhookian Brachiopods.
(Nos. 239-245). BIRT seen OD Sey eee n teense re esau nc eee regen
Dominican ostracodes, Lepidocyclina, mollusks.
(Nos. 246-247). GST DD ay OO) Seperate ee eee ceeaecasaneasesecescusaorcnes
Cenozoic corals, Trinidad Neogene mollusks.
(Nos. 248-254). SD joyee poly cae ee at ee aren eee
Forams, North Carolina fossils, coral types, Cenozoic
Echinoids, Cretaceous Radiolaria, Cymatiid gastropods
(Nos. 255-256). CPST oa) VAN) 0) Klee eee ee ia a ea
Jurassic ammonites.
(Nos. 257-262). HOS joe KS eV crete a ee eee
Cretaceous Radiolaria and Forams, Pacific Silicoflagellates,
North American Cystoidea, Cyclonema, Vasum.
(No. 263). CAT BR 0 oe Seer he CALe Ree pete tte Ree Nia ei ee
Bibliography of Cenozoic Echinoidea.
(Nos. 264-267). BB SUID Dis OS DIS s cesresteree trees cot ete i ce acccedaieeesae
Radiolaria, cirripeds, Bryozoa, palynology.
(Nos. 268-270). BOS SP Pe. Sill Pisce ees cere eats
Mollusks, Murex catalogue, Cretaceous Radiolaria.
(Nos. 271-274). U7 /G\ V0) one, Oe i 0) esta Ap
Trace fossils, ammonoids, Silicoflagellates, microfauna.
(Nos. 275-277). VAD Foy ets SNe 10) Rita eee
Chitinozoa, Spumellariina, Mexican Ammonites
(Nos. 278-281). BO joy ele AbLL TO. cece acetone es rte eer
Palynology, corals, echinoderms, Foraminifera, and crinoids.
(No. 282). FTP) 0) 0 ee) 0) bc mnie 95 Ree Sere ern
Ostracode Symposium.
(Nos. 283-286). CA Tah oye Be Pug 0) Cf Soe pe ee Oe ne eee RS rem
Crinoids, gastropods, corals, ostracodes.
(No. 287). PUY: 40) ee O00) ON ise eee ee sete ec eet
Misc. Paleozoic
(Nos. 288-290). DREN Tas 0) SpA 190) 9) Ce ee ean ee eee
Paracrinoidea, ostracodes, cirripeds.
(No. 291). PHA RS 3} SEAR} Way 0) (Mme ee ot a ace enue Pe
Bryozoa.
(Nos. 292-294). re OR ol ofr, O-7RM oC raetie oe otae a ee nae Ree. ee ten
Turrids, gastropods, forams.
(No. 295). 31/9) SDD sya SO pm Dl Sap rect sees ss cece cw eeet cceeeshacacctateos
Paleocene Nigeria
(Nos. 296-298). PP SN Of BRE PASO 0) (a eee ee ek iene Re a eR
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BULLETINS Sage pra ol
OF MAR 2 4 197
AMERICAN &eN2e3iry
(Founded 1895)
Vol. 72
No. 300
PRIMARY TYPES IN THE STANFORD
PALEONTOLOGICAL TYPE COLLECTION
By
Jupita Terry SMITH
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
PALEONTOLOGICAL RESEARCH INSTITUTION
1976-1979
PRES TUE Ny Paya ea ee eRe er a eee LS HAROLD E. VoKES
VACKE=PRESIDEIN Tyner, eens Bede ea ee een er ae pee ee DUANE O. LERoy
SECRETAR Vere 2 opto eee rene ae BY ee ne ee eS Puitrie C. WAKELEY
PIC REA SURE Rye rents ee eae eee ee he re ERNESTINE Q. WRIGHT
DIRECTOR ee rt ae ee -KATHERINE V. W. PALMER
VASSTS TANIA) TRIE CL) gene ccee a a a ...PETER R. Hoover
ASSISTANT SECRETARY, ASSISTANT TREASURER ..........--cece--se-e-ceeeeeeeees REBECCA S. HARRIS
MC OUINSE Era ee gt ee ARMAND L, ADAMS
REPRESENTATIVE AAAS COUNCIL .........- RED SS Cee, Be eee -RIcHARD G. Oscoop, Jr.
Trustees
RutH G. Browne (1976-1979) KATHERINE V. W. PALMER (Life)
KENNETH E. CASTER (1975-1978) Joun Pojeta, Jr. (1975-1978)
Joun L. CIsNE (1976-1977) K. NorMAN SACHS, JR. (1974-1977)
ResBecca S. Harris (Life) DANIEL B. Sass (1974-1977)
MarcareT B. HERoy (1975-1978) HAROLD E. VoKEs (1975-1978)
Duane O. LERoy (1974-1977) PuHiLie C. WAKELEY (1976-1979)
Wu.1aM A. OLIVER, JR. (1976-1979) ERNESTINE Q. WRIGHT (1976-1979)
BULLETINS OF AMERICAN PALEONTOLOGY
and
PALAEONTOGRAPHICA AMERICANA
KATHERINE V. W. PALMER, Editor
PETER R. HOOVER Doris C. BRANN
Assistant Assistant
Advisory Board
KENNETH E. CASTER HANS KUGLER
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Pelee INS
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PAEEONTOLOGY
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Vol. 72
No. 300
PRIMARY TYPES IN THE STANFORD
PALEONTOLOGICAL TYPE COLLECTION
By
JupirH Terry SMITH
March 14, 1978
Paleontological Research Institution
Ithaca, New York 14850, U.S.A.
Library of Congress Card Number: 78-51977
Printed in the United States of America
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CONTENTS
Page
DENSE G ULE CO 0 GS ee ee 317
UES FB VA ON WET a 0) aA Pee ee 317
Arrangement of the Stanford University Paleontological Type
Colle etic rime eer eet aio en rN EEE PSR VRE 2 318
Type collection records, card files and ledgers ._....................-0--o--cc-sessesseeesese 318
Special strengths) and) historical) perspective: <22sc2ccc-cccecctocencsesesee scene ecesenscecece 320
Winexpectedmty pesm amd whol Gili pg ieee eee ena ee 320
FTV; OEY) DCS accor eco ea ne ees ae A eS ne eee 322
Other Stantord Winiversity, dsyper Collections seer eee 323
JENS TERR AL (SS VSENC) ssc eer A a Ee ee a eee 323
Catalogue mannan ge riie i tyes ee a ae 324
Rrepanationvotatiencatalop icy see eee ee ee 324
DEPOSIT OTIC, peace eco notes eae rnin S, eae eens Cee A estat, oe 325
Catalocucgofaprimany,sty pes ese < esos ae eee 325
INT OTS Cage seen ses Ce EE ee eS 5 a ete OE 325
1 EAS) KER i 0X0) 0 Sere a ek RN ea ert 325
We phial o pod aly eas a i ae ee a ere 374
Scapho pod ay corsets Sate tee) hee ee ee ee er 8 eee 398
(GASET Op OG lpr c scene ene ene ere ree et a aaa eee east ee ee 398
Poly placophoram (eAnnp lime iia) aces 459
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PRIMARY TYPES IN THE STANFORD
PALEONTOLOGICAL TYPE COLLECTION
JupirH Terry SMITH
INTRODUCTION
The Stanford University Paleontological Type Collection is part
of an important national, as well as West Coast resource of Holo-
cene and fossil, primarily invertebrate, specimens that are housed
with an extensive library of systematic publications in the Depart-
ment of Geology, Stanford University, Stanford, California. The
University acquired its first general collections between 1892 and
1895 at the instigation of James Perrin Smith, Assistant Professor
of Mineralogy and Paleontology. Although the nearby California
Academy of Sciences collections were severely damaged in the earth-
quake and fire of 1906, the Stanford collections remained unharmed
and intact. Acquisitions increased steadily, and by the mid 1920's
the collection included large suites of irreplaceable material from
localities no longer accessible, such as many drawers of Pleistocene
specimens collected from Deadman Island, off San Pedro, California,
before it was destroyed in the mid 1900’s. Several drawers of Paris
Basin material was received from A. E. M. Cossmann in exchange
for California specimens collected by Delos Arnold. Large population
samples, primarily from the California Coast Ranges and Trans-
verse Ranges, were contributed by Stanford Summer Geology classes
from the late 1890’s to the present.
THE TYPE COLLECTION
The Stanford University Department of Geology became a
recognized repository for type specimens in 1924 when four graduate
students, Eric Knight Jordan, Leo George Hertlein, Albert B. Rea-
gan, and Colin H. Crickmay, set up a register and numbering sys-
tem. Many holotypes and paratypes described before then remained
in the general collections until 1940, when A. Myra Keen became the
curator and undertook an exhaustive search to isolate unrecognized
type material. Those who had some charge of the collections over
the years included Carl H. Beal (1911-1915), Ida Shepard Oldroyd
1As of March 9, 1977, the type and general collections were transferred from
Stanford University to the Department of Geology of the California Academy
of Sciences, Golden Gate Park. San Francisco, California 94118. The types
are now stored in the type collection room, and the general collection is to be
integrated with the holdings of the Academy. No Stanford numbers are to
be changed and type specimens should be cited as SUPTC numbers now in
the collections of the California Academy of Sciences.
318 BuL.etin 300
(1917-1940), and A. Myra Keen (1936 until retirement in 1970;
curator emeritus 1970-).
ARRANGEMENT OF THE STANFORD UNIVERSITY
PALEONTOLOGICAL TYPE COLLECTION
Type specimens are arranged in order of numerical accession in
locked cases in the Department of Geology. They include 6,435
primary and secondary types as of June 30, 1976. Microfossils are
numbered sequentially with the megafossils but housed separately.
Type specimens bear numbers 1 to 1000 and 5000 to 10,345; num-
bers 1001-4999 were set aside for the general collections and never
used for types.
Type specimens are marked with the symbols shown in Text-
fig. 1, which were recommended by Howell (1929, p. 7). They may
also bear Stanford University locality numbers on a spot of white
paint or a general collection accession number on yellow paint. Some-
times these numbers are written directly on the specimens. Numbers
on turquoise paint identify specimens formerly in the collection of
Hubert G. Schenck.
Published references to Stanford material should be to Stanford
University Paleontological Type Collection (SUPTC) numbers.
LSJU, sometimes abbreviated to SU, numbers refer to general col-
lection accession numbers and in some cases to Stanford locality
numbers.
TYPE COLLECTION RECORDS, CARD FILES AND LEDGERS
This publication lists only primary types: holotypes, paratypes,
syntypes, lectotypes, plastoholotypes and plastosyntypes. Four com-
plete listings of all holdings are kept with the type collection: three
sets of file cards and a ledger in four volumes. Duplicate suites of
4 x 6 inch cards are arranged in two series, one systematically and
one alphabetically by genus. Each card gives page and figure refer-
ences for the type specimen, its locality, age, and formation data
where applicable. For cross reference, specific names are indexed
alphabetically on 3 x 5 inch cards. Complete bibliographic reference
cards are arranged alphabetically by author and include annotations
of which type numbers are described therein. Many of the original
references are represented in the Conchological Library reprint col-
STANFORD UNIVERSITY Types: SMITH 319
lection, housed near the types; others are in the journals in the Uni-
versity’s Branner Geology and Falconer Biology Libraries.
The ledger in four volumes, begun in 1924, records only type
numbers and names of taxa. Locality data are detailed in two mega-
faunal ledgers and the Micropaleontology Locality Book of M-num-
bers. Megafaunal and microfossil localities were recorded together
in the megafaunal ledger from 1923 to 1936. Type collection and
locality ledgers and Harold Hannibal’s handwritten “N.P. book” of
North Pacific localities were microfilmed in January, 1969. Addi-
tional sources of locality data are the Stanford Summer Geology
notebooks and a book of Ralph Arnold’s California localities, the
“C book,” reconstructed from specimen labels by A. Myra Keen.
KINDS_ OF
TYPE SPECIMENS MARKS COLOR
HOLOTYPE
S<
ao SYNTYPE Oo
7
s PARATYPE Se GOLD
SS
see hoa Gee
MEEa
NEOTYPE <<
Sy san > [cast] [ca] a [ca]
©
HYPOTYPE
S (PLESIOTYPE) /\ SILVER
c
g TOPOTYPE (1) WHITE
Text-figure 1. Type specimen marks and symbols.
320 BULLETIN 300
SPECIAL STRENGTHS AND HISTORICAL PERSPECTIVE
Type holdings are greatest in Cenozoic mollusks, especially
gastropods and pelecypods, and Foraminifera. The majority of
specimens are from the North Pacific, tropical eastern Pacific, and
the California marine Tertiary. From 1903 to the 1960’s a number
of large important monographs were published based on material
in the Stanford Paleontological Type Collection. These included com-
prehensive systematic studies reflecting the research interests and
course offerings of professors James Perrin Smith, Siemon W. Muller,
and N. J. Silberling (Paleozoic and Mesozoic mollusks), Hubert G.
Schenck (Foraminifera and Tertiary mollusks), Hans Thalmann,
Joseph J. Graham and James C. Ingle (Foraminifera), A. Myra
Keen (Tertiary and Quaternary mollusks), and W. R. Evitt (or-
ganic microfossils ).
Many of the Stanford paleontologists, especially Siemon W.
Muller and James Perrin Smith, acquired plaster casts of compara-
tive material for their studies and placed these specimens in the type
collection. The 483 plastoholotypes and plastosyntypes in the Stan-
ford collection constitute an important resource for those taxa whose
originals may be lost or are deposited in foreign repositories that do
not lend primary specimens. Many of the plastoholotypes listed here
are accompanied by plastoparatypes, although the latter are omitted
from the catalogue. Table 1 lists authors, papers and numbers of
plastoholotypes in the Stanford Paleontological Type Collection.
UNEXPECTED TYPES AND HOLDINGS
Students of Clark and Arnold’s “Fauna of the Sooke Formation,
Vancouver Island” (1923) will find all but four of the holotypes and
most of the paratypes published with University of California
(UCMP) numbers in the Stanford University Type Collection. The
paper was in press before Arnold, who had financed the field work
and intended the material for Stanford, had the specimens trans-
ferred from the University of California and renumbered. Many of
the new species were described from SU loc. NP 129, sometimes
cited as between Muir and Coal Creeks, elsewhere as between Muir
and Kirby Creeks. Coal Creek is an older name for Kirby Creek,
the current name.
Three holotypes, 1 syntype and 43 paratypes described by W. H.
Dall were deposited in the Stanford University Type Collection in
STANFORD UNIVERSITY Types: SMITH 321
exchanges arranged by Ida Oldroyd. Many of these are Holocene
species from the North Pacific and the Galapagos Islands, useful to
West Coast workers and more readily available than the holotypes
in the U.S. National Museum, Washington, D.C. The holotypes are
Sigaretus oldroydu Dall, 1897c; Drila empyrosia Dall, 1899a; and
Atrina oldroydu Dall, 1901a. The syntype is Venericardia hadra
Dall, 1903b, from Florida, and there is a neotype, Lasaea subviridis
Dall, 1899b designated by Keen (1938).
TABLE 1—PLASTOHOLOTYPES AND PLASTOSYNTYPES
Author Subject No. of Types
Burckhardt, 1903 Jurassic cephalopods, Chile and Argentina 7
Dickerson, 1914 Paleocene mollusks, California 7
Goldfuss, 1836 Cretaceous pelecypods, Germany 6
Hyatt and Smith, 1905 ‘Triassic cephalopods, Inyo Mts. 45
Kittl, 1912 Triassic pelecypods, Austria 9
Merriam, 1941 Fossil Turritellas 21
Oppel, 1862 Jurassic mollusks, Tibet and Germany 9
Popenoe, 1937 Cretaceous mollusks, Santa Ana Mts. 30
Reeside, 1927b Cephalopods, western U.S. 12
Smith, 1914 Triassic cephalopods, Humboldt Range 48
Smith, 1927 Triassic mollusks, northern California 34
Smith, 1932 Triassic cephalopods, Inyo Mts. 46
Waagen, 1895 Triassic cephalopods, Salt Range 15
Ninety-six other papers are represented by five or fewer plastoholotypes, of
which perhaps the most unexpected are Diener (1895, 1903, 1907), Gabb (1864,
1866), Jimbo (1894), Mantell (1822), Matsumoto (1942, 1955a, 1955b, 1956),
Noetling (1880), Pavlow (1891), Schluter (1867), Spath (1921), Steiger
(1914), Uhlig (1910), Waagen (1867), Wachsmuth and Springer (1890).
Five or fewer plastoholotypes: Anderson, 1902; Arnold, 1903, 1906, 1908a;
Billings, 1859, 1860, 1861, 1863, 1865; Clark, 1915, 1918, 1925, 1932, 1938; Clark
and Anderson, 1928; Clark and Arnold, 11923) Clark and Woodford, IS YA/3
Conrad, 1858; Diener, 1895, 1903, 1907; Durham, 1944, 1950, 1957; Fenton and
Fenton, 1938; Gabb, 1864, 1866; Hanna, 1927; Heinz, 1928, 1934; Jenkins,
1913; Jimbo, 1894; Keen, 1954; Keen and Campbell, 1964; Loel and Corey,
1932; Mantell, 1922: Marwick 1944; Matsumoto, 1942, 1955a, 1955b, 1956;
McLearn, 1931, 1933b; Meek, 1864a, 1864b, 1876, 1877; Meek and Hayden,
1856, 1859, 1861, 1863, 1865; Nelson, 19251: Nicol, 1945; ‘Noetling, 1880; Nom-
land, 1916b, 1917a, 1917b; Olsson, 1944: Owen, 1852; "Parker, 1949; Pavlow,
1891; Pilsbry and Olsson, 1941; Reeside, 1927a; Reinhart, 1937a, 1937b; Rivers,
1913; Schenck, 1936; Schluter, 1867; Shattuck, 1903; Shimizu, 1930; Smith,
1904; Spath, 1921; Stanton, 1895, 1920; Steiger, 1914; Stephenson, 1923: Tile
mann, 1917; Turner, 1936; Uhlig, 1910; Vincent, 1913: Vokes, 1935, 1939;
Waagen, 1867; Wachsmuth and Springer, 1890; Wade, 1926; Wagner and
Schilling, 1923; Walcott, 1884; Waterfall, 1929; Weaver, 1905; Wheeler, 1939;
Whiteaves, 1884, 1893; Woodring, 1938; Yabe, 1904; Vane and Shimizu, 1921.
BuL.etin 300
os)
bo
bo
Seven holotypes, three syntypes and one paratype described by
Diener (1914, 1916) were purchased from the Palaeontologische
Institut, Weiner Universitat, by Professor James Perrin Smith. They
were discovered in a Stanford attic in the 1930’s along with 10
plastoholotypes, accounting for 21 of Diener’s ammonite types from
the Triassic of the New Siberian Islands, Madagascar, and the Hima-
layas. Two types of Quenstedt (1885) may have been acquired at
the same time — the holotype of Ammonites psilonotus plicatus and
a syntype of Ammonites laqueus from the Jurassic of Germany.
Although most of the California Cretaceous and Tertiary speci-
mens described by Gabb (1864) are at the Philadelphia Academy of
Natural Sciences, the Harvard Museum of Comparative Zoology, or
the University of California Museum of Paleontology, two paratypes
are in the Stanford University Paleontological Type Collection:
Aporrhais californica Gabb, 1864, Cretaceous of the Siskiyou Moun-
tains of California. Other types which are at Stanford, while all the
other material described in the same papers is elsewhere, are the
crinoid Actinocrinus arnoldi Wachsmuth and Springer, 1890 from
Marshall Co., Iowa, and the upper Paleozoic pelecypod Chaenomya
maria Worthen, 1882 from Shawnee Co., Kansas.
Most of the type specimens are marine taxa, although many
fresh water and land mollusks are represented. They include taxa
described by Hemphill (1876-1901, 119 types); Henderson (1913-
1935, 22 types); Pilsbry, (1891-1940, 28 paratypes, 34 coauthored
types); Hannibal (1912b, 36 types); Dall (1896, 1900a, 1917c);
Berry (1930a, 1932, 1937, 1938b, 1940a); Fred Baker (1914); and
Frank Baker (1939).
HYPOTYPES
Some of the most important holdings are the hypotypes or
figured specimens, which are not treated here because of space limi-
tations. These include large numbers of well-preserved specimens
illustrated in the monographs of Grant and Gale (1931, 81 types),
Miller (1947, 27 types of Tertiary nautiloids of the Americas) and
Matsumoto (1959, 74 upper Cretaceous ammonites of California).
Specimens illustrated in two editions of “Sea Shells of Tropical
West America” by A. Myra Keen (1958, 1971) remaian in the gen-
eral collection. A silver dot identifies these shells.
STANFORD UNIVERSITY Types: SMITH 323
OTHER STANFORD UNIVERSITY TYPE COLLECTIONS
Over the years workers at Stanford University in departments
other than Geology described new species that were kept in separate
collections at the Hopkins Marine Station, Pacific Grove, and in the
Stanford Natural History Museum of Systematic Biology on the
main campus. These taxa included new species of mollusks, mam-
mals, Holocene and some fossil fish whose types might be expected
in the paleontological type collection but which were never part of it.
The Stanford Natural History Museum type ledgers began in
1939 and were closed in 1963 when the material, including David
Starr Jordan’s fish collection, was moved to the California Academy
of Sciences in San Francisco. Fossil fish went to the Department of
Geology, insects to the Department of Entomology. In 1971 the
types at Hopkins Marine station were transferred to the Academy’s
Department of Invertebrate Zoology. These included wet-preserved
cephalopods described by S. S. Berry (1908-1910) and hundreds
opisthobranch gastropod types described by MacFarland (1966)
(James Carlton, personal communication, 1972). In 1976, the Dud-
ley Herbarium also was moved to the Academy.
Some of the transferred types are described or listed, in some
cases with other California Academy of Sciences holdings, in the
following: MacFarland (1966, opisthobranchs); Mayer (1949, mam-
mals in the type collection of Stanford Natural History Museum);
Jordan (1896-1900, 4 volumes on “The Fishes of Middle and North
America”); Bohlke, 1953 (type specimens of Recent fishes); Firby
(1972, fossil fish).
ACKNOWLEDGMENTS
In assembling the information catalogued here, I was greatly
assisted by A. Myra Keen, whose careful records and personal com-
munications helped solve many problems of dates and references,
and whose curatorial work over the past 36 years made the Stan-
ford University Paleontological Type Collection the valuable re-
source it now is. My work was encouraged and helped by present
and former members of the Paleontological Committee of the De-
partment of Geology: W. R. Evitt, James C. Ingle, Warren O. Addi-
cott, Norman J. Silberling, Carole §. Hickman, and A. Myra Keen.
324 BuLteTINn 300
Joseph H. Peck of the University of California Museum of Paleon-
tology consulted on missing type specimens and LouElla R. Saul,
University of California, provided information on some of the plasto-
holotypes. Text-fig. 1 was drafted by Natalie Miller, U.S. Geological
Survey, Menlo Park, and the manuscript was read by Warren O.
Addicott and A. Myra Keen. Publication funds were provided by
the Pacific Section of the Society of Economic Paleontologists and
Mineralogists and by the Department of Geology, Stanford Uni-
versity.
CATALOGUE ARRANGEMENT
Holocene and fossil taxa are listed alphabetically by species or
subspecies under Phylum, or, in the cases of Mollusca and Crustacea,
Class. The format is as follows: .
SUPTC
no. species, Genus (Subgenus) : Author Kind of type
or subspecies, Genus (Subgenus) species: Author
Author, date, page, plate, figure:
Type locality. [supplementary data, e.g., current quadrangle]
Age, Formation, if a fossil: [supplementary information ],
1 Subsequent references are given only if the illustrations are much improved
or if the original paper is not readily accessible.
2 Locality is for the particular specimen. Early workers occasionally designated
paratypes from localities other than the holotype locality.
3 Age and formation are those given in the original description; many have
been reassigned by later workers and should be verified with more recent
publications. The formations “Chico” and “Horsetown”’ were used widely
for Cretaceous rocks in California and are especially untrustworthy.
4 Supplementary information includes currently accepted age and formation
assignments where these were readily available, data for missing specimens,
and repositories of plastoholotypes represented in the Stanford type collection.
Repositories are those from which the casts were received and may not reflect
the present repositories of the types. Nomenclatural annotations are beyond
the scope of this paper, but can be found in Keen and Bentson (1944), Keen
(1971) and other systematic reviews.
PREPARATION OF THE CATALOGUE
About two thirds of the catalogue was compiled from systematic index
cards, the remaining third directly from specimen labels and original references.
The author checked megafaunal lists against the specimens in 1974-1976. Micro-
fossi] types were inspected by Elizabeth Watson in 1940-1941 and by Marjorie
Korringa in 1965, since which time there have been few acquisitions of these
groups. Most of the original references were consulted and publication dates
verified.
STANFORD UNIVERSITY Types: SMITH 325
DEPOSITORIES
ANSP Academy of Natural Sciences of Philadelphia
BM(NH) _ British Museum of Natural History
CAS California Academy of Sciences
CIT California Institute of Technology
Geol. Surv. Canada Geological Survey of Canada
LACMNH Los Angeles County Museum of Natural History
LSJU (or SU) Leland Stanford Junior University
N.Z. Geol. Surv. New Zealand Geological Survey
MCZ or Mus. Comp. Zool. Harvard Museum of Comparative Zoology
SUPTC Stanford University Paleontological Type Collections
UCLA University of California at Los Angeles
UCMP University of California at Berkeley
USGS or U.S. Geol. Surv. United States Geological Survey
USNM or U.S. Nat. Mus. United States National Museum
I.G.P.S. Institute Geology and Paleontology, Tohoku Univ., Sendai, Japan
CATALOGUE OF PRIMARY TYPES
PELECYPODA
7616 acutiplicatus, Pecten: Meek Plastoholotype
Meek, 1864b, p. 46, pl. 8, fig. 3
Plumas Co., Calif.; Genessee Valley area
Jurassic [cast received from Museum of Comparative Zoology |
154 aequilateralis, Spisula: Waring Holotype
Waring, 1917, p. 80, pl. 14, fig. 8
Ventura Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm
5345 africana, Fossularca: Newton Paratype
Newton, 1922, p. 68
Southern Nigeria, Africa; Ameki, Omobialla district
Upper Eocene, upper Lutetian
6061 agulhasensis, Arca (Acar): Thiele Paratype
Thiele and Jaechel, 1931, p. 177
Cape Agulhas, near Cape of Gocd Hope, South Africa
7971 alargada, Anadara (Anadara): Marks Paratypes
Marks, 1951, p. 56
SW Ecuador; Zacachiin corehole, depth 710-720’
Miocene, Subibaja Fm
6062 alaskana, Halobia: Smith Holotype
Smith, 1927, p. 113, pl. 100, fig. 5
SE Alaska; Gravina Island, Thompson Cove
Upper Triassic
5514 alaskana, Monotis: Smith Plastoholotype
Smith, 1927, p. 119, pl. 101, fig. 1
Copper River region, Alaska; Mill Creek, near forks
Upper Triassic [holotype USNM 74193]
44 aletes, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925a, p. 8, pl. 2, fig. 4
Baja California, Mexico; Rancho Refugio, N of San Jose del Cabo
SU loc. 50
Upper Miocene or Lower Pliocene
326
45
6943
5231
9932
8504
8504a
559
75360
7560a
134
299
5137
8734
8735
8500
8501
BuLLetin 300
aletes, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925a, p. 8, pl. 2, fig. 1
Baja California, Mexico; Rancho Refugio, SU loc. 50
Upper Miocene or Lower Pliocene
aleutica, Mysella: Dall Paratype
Dall, 1899b, p. 892
Aleutian Islands, Alaska; Kyska Harbor
alkiensis, Leda: Clark Holotype
Clark, 1925, p. 76, pl. 8, figs. 7, 10
Seattle, Wash.; R.R. cuts between Argo and Georgetown stations SU
loc. NP 49
Oligocene, Blakeley Fm
americana, Daonella: Smith Plastoholotype
Smith, 1914, p. 145, pl. 49, figs. 4, 5
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74362]
americana, Leptomya: Keen Holotype
Keen, 1958a, p. 246, pl. 30, fig. 10; pl. 31, figs. 3, 6
Panama; San Miguel Bay, E side of Punta Alegre ;
americana, Leptomya: Keen Paratype
Keen, 1958a, p. 246, pl. 30, fig. 9; pl. 31, fig. 5
Panama; San Miguel Bay, E side of Punta Alegre
anahuacensis, Immanitas: Palmer Paratype
Palmer, 1928a, p. 30, pl. 4, fig. 1
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
anahuacensis, Immanitas: Palmer Paratypes
Palmer, 1928a, p. 30
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
andersoni, Pecten (Plagioctenium): Arnold Paratypes
Arnold, 1906, p. 82, pl. 26, fig. 6
Santa Clara Co., Calif.; near Stanford University, Frenchmens Tower
Miocene, Temblor Fm
andersonianum, Sphaerium (Amesoda): Hannibal Holotype
Hannibal, 1912b, p. 132, pl. 6, fig. 11. Also in Taylor and Smith, 1971,
fig. 6
Badland Hill, Oregon; 1 mile E of Sand Hollow
Pliocene, Idaho Lake Beds [Grassy Mountain Fm, fide Taylor and
Smith]
angermanni, Ostrea: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 621, pl. 17, figs. 3, 6
Baja California, Mexico; trail from Arroyo Mesquital to La Purisima,
in Turritella bed above San Gregorio Lagoon SU loc. 59
Miocene, Isidro Fm
_annulatus, Inoceramus: Goldfuss Plastosyntypes
Goldfuss, 1826, p. 114, pl. 110, figs. 7a, 7b
Westphalia, Germany
Cretaceous [casts of Goldfuss specimens 675b, 675c from BM(NH) ]
anomioides, Plicatula: Keen Holotype
Keen, 1958a, p. 241, pl. 31, figs. 4, 7, 8. Also im Keen, 1971, p. 94, fig.
206
Sonora, Mexico; Guaymas
anomioides, Plicatula: Keen Paratype
Keen, 1958a, p. 241
Sonora, Mexico; Guaymas
619
7995
364
526
9736
5908
5908a
7273
5213
5214
8103
STANFORD. UNIVERSITY Types: SMITH 327
aragonia, Venericardia planicosta: Arnold and Hannibal Neotype
Arnold and Hannibal, 1914, p. 907. Illustrated in Waring, 1915, pl. 1,
fig. 22. Designated neotype of Venericardia (Leuroactis) aragonia by
Stewart, 1930, p. 170
Umpqua Valley, Ore.
Upper Eocene, Umpqua Fm [= holotype of Venericardia planicosta
ionense Waring, 1915]
argentea, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 25, pl. 1, figs. 10-14
Fresno, Co., Calif.; Tumey Hills Qd, Sec. 29, T 15 S$, R 12 E SU loc.
2073
Paleocene, Lodo Fm
arnoldi, Pecten (Lyropecten): Aguerrevere Holotype
Aguerrevere, 1925, p. 51, pl. 5
Sucre, Venezuela; 1.75 miles E of Cumana Castle
Miocene?
auburyi, Pecten (Pecten): Arnold Paratypes
Arnold, 1906, p. 94
Los Angeles Co., Calif.; Puente Hills, 1 mile E of Chandler Wells
Pliocene
aurora, Semele: Tursch and Pierret Holotype
Tursch and Pierret, 1964, p. 35, figs. 1, 2
Off Rio de Janeiro, Brazil, 30 fms, on sand
austini, Leda: Oldroyd Holotype
Oldroyd, 1935, p. 14, fig. 2
British Columbia, Canada; near Nanaimo, off Neck Point
austini, Leda: Oldroyd Paratype
Oldroyd, 1935, p. 14
British Columbia, Canada; near Nanaimo, off Neck Point
baileyi, Solen gravidus: Loel and Corey Plastoholotype
Loel and Corey, 1932, p. 230, pl. 44, fig. 5
Ventura Co., Calif.; South Mountain UCMP loc. A-244
Miocene, Vaqueios Fm_ [holotype UCMP 31831]
bainbridgensis, Cochlodesma: Clark Holotype
Clark 1925> p> 86, ple 13% fig. 3
Bainbridge Island, Wash.; beach between S side of entrance to
Blakeley Harbor and Restoration Point SU loc. NP 103
Oligocene, Blakeley Fm
bainbridgensis, Cochlodesma: Clark Paratype
Clark, 1925, p. 86, pl. 13, fig. 4
Bainbridge Island, Wash.; beach between S side of entrance to
Blakeley Harbor and Kestoration Point SU loc. NP 103
Oligocene, Blakeley Fm
balesi, Arca (Barbatia): Pilsbry and McLean Paratype
Pilsbry and McLean, 1939, p. 1
Missouri Key, Fla.
balesi, Asthenothaerus: Rehder Paratype
Rehder, 1943, p. 189
Missouri Key, Fla.
bardwelli, Macrocallista (Paradione): Clench and McLean
Clench and McLean, 1936, p. 202 Paratype
Australia ;
baughmani, Anadara: Hertlein Paratype
Hertlein, 1951b, p. 487
SE of Port Aransas, Texas; in 40 fms
507
55
56
6947
10337
35
9910
10323
10327
6532
7527
7527a
7527b
BuLLeTIN 300
beali, Mactra: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 80. Illustrated in Wiedey, 1929b, pl. 1,
fig. 3
Alameda Co., Calif.; Pleasanton Qd
Miocene, Monterey Fm
beali, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925a, p. 10, pl. 2, fig. 3
Baja California, Mexico; Arroyo Fortuna, N of San Jose del Cabo SU
loc. 44
Pliocene?
beali, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925a, p. 10, pl. 5, fig. 8
Baja California, Mexico; 2 mi. NW of arroyo near La Palma, N of
San Jose del Cabo SU loc. 64
Pliocene ?
beckii, Liocyma: Dall Paratype
Dall, 1870, p. 257
Eastern Siberia; Plover Bay, Bering Strait
bella, Cymbophora: Saul ' Paratypes
Saul, 1974, p. 1089
Butte Co., Calif.; Cherokee Qd, near Pentz, conglomerate beds 1400'S,
600’W of NE cor. Sec. 36, T 21 N, R 3 E, UCLA loc. 4340
Cretaceous, early Campanian, Chico Fm
bellilamellatus, Pecten (Chlamys): Arnold Holotype
Arnold, 1906, p. 108, pl. 41, figs. 6, 6a, 7, 7a
San Diego Co., Calif.; Pacific Beach
Pliocene, San Diego Fm
beringiana, Myophoria: Smith Plastoholotype
Smith, 1927, p. 109, pl. 101, fig. 3
SE Alaska; Gravina Island
Upper Triassic [holotype USNM 74194]
beta, Lima (Acesta): Popenoe Plastoholotype
Popenoe, 1937, p. 382, pl. 45, fig. 5
Santa Ana Mts., Calif.; CIT loc. 1069
Cretaceous, Turonian [holotype UCLA 40619]
bifurcatus, Brachidontes: Popenoe Plastoholotype
Popenoe, 1937, p. 383, pl. 46, fig. 2
Santa Ana Mts., Calif.; CIT loc. 974
Cretaceous, Campanian [holotype UCLA 40622]
binakayanensis, Arca: Faustino Paratypes
Faustino, 1932, p. 545
Manila Bay, Philippines; Paranaque, Rizal
birchi, Nucula (Ennucula): Keen Holotype
Keen, 1943, p. 41, pl. 3, fig. 12
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4 Sec.
6, T 29 S,R30£E SU loc. 2121
Miocene, Temblor Fm, Round Mountain
birchi, Nucula (Ennucula): Keen Paratype
Keen, 1943, p. 41, pl. 3, figs. 9, 11
Kern Co., Calif.; Caliente Qd, SW 1/4 Sec. 6, T 29 S, R 30 E. SU
loc. 2121
Miocene, Temblor Fm, Round Mountain
birchi, Nucula (Ennucula): Keen Paratype
Keen, 1943, p. 41, pl. 3, fig. 10
Kern Co., Calif.; Caliente Qd, SW 1/4 Sec. 6, T 29 S, R 30 E. SU loc.
2121
Miocene, Temblor Fm, Round Mountain
6026
6026a
9908
59
9739
9740
9741
220
432
8334
160
7876
7781
358
356
8696
STANFORD UNIVERSITY Types: SMITH 329
bisenensis, Anadara (Anadara): Schenck and Reinhart Holotype
Schenck and Reinhart, 1938, p. 44, pl. 4, figs. 2a, 2b, 2e; pl. 5, figs. 1a,
fey id
Inland Sea, Japan; Bisen, Okayama Prefecture
bisenensis, Anadara (Anadara): Schenck and Reinhart Paratype
Schenck and Reinhart, 1938, p. 44
Inland Sea, Japan; Bisen, Okayama Prefecture
blackburnei, Lima: Smith Plastoholotype
Smith, 1927, p. 122, pl. 103, fig. 11
Alaska, Copeland Creek
Upper Triassic [holotype USNM 74216]
blancoensis, Acila: Howe Holotype
Howe, 1922, p. 95, pl. 9, fig. 3
Cape Blanco, Ore. SU loc. NP 26
Pliocene, Empire *m
borealis, Aligena (Odontogena): Cowan Holotype
Cowan, 1964, p. 108, pl. 20, figs. 1, 2
Georgia Strait, British Columbia, Canada; 190 fms
borealis, Aligena (Odontogena): Cowan Paratypes
Cowan, 1964, p. 108
Georgia Strait, British Columbia, Canada; 190 fms
bosei, Pecten (Pecten): Hanna and Hertlein Paratypes
Hanna and Hertlein, 1927, p. 154
Baja California, Mexico; canyon inland 1/2 mile from Santa Antonita
Point CAS loc. 795
Upper Pliocene
bowersi, Pecten (Lyropecten): Arnold Paratype
Arnold, 1906, p. 70, pl. 12, fig. 2
Ventura Co., Calif.; Santa Monica Mts.
Lower Miocene
bramkampi, Aequipecten circularis: Durham Plastoholotype
Durham, 1950, p. 63, pl. 9, figs. 4, 8
Imperial Co., Calif.; NW side Carrizo Mountain UCMP loc. A1268
Lower Pliocene, Imperial Fm [holotype UCMP 30035]
branneri, Crassatellites: Waring Holotype
Waring, 1914, p. 782. Illustrated in Waring, 1917, p. 74, pl. 14, fig. 17
Ventura Co., Calif.; Calabasas Qd, Simi Hills, Martinez area
“Lower Eocene,” Martinez Fm_ [Paleocene]
branneri, Glycymeris: Arnold Plastoholotype
Arnold, 1908a, p. 377, pl. 34, fig. 1
San Mateo Co., Calif.; Mindego Creek, 1 mile above Alpine Creek
Arnold loc. 12
Upper Oligocene to lower Miocene
branneri, Mulinia: Dall Paratype
Dall, 1901b, p. 145
Mamanguape, Brazil
branneri, Pecten (Chlamys): Arnold Holotype
Arnold, 1906, p. 55, pl. 3, fig. 9 (cast of external mold)
Santa Clara Co., Calif.; near Stanford University, Tuff Hill
Lower Miocene, Vaqueros Fm
branneri, Pecten (Chlamys): Arnold Paratypes
Arnold, 1906, p. 55, pl. 3, figs. 10, 11 (casts of fragments of molds)
Santa Clara Co., Calif.; near Stanford University, Tuff Hill
Lower Miocene, Vaqueros Fm
branneri, Trigonia: Anderson Holotype
Anderson, 1958, p. 112, pl. 17, fig. 5
Siskiyou Co., Calif.; rocky gulch 2.5 miles SW of Hornbrook
Upper Cretaceous
330
9911
9870
7864a
7864b
7864¢c
7248
5368
8843
8444
436
6732
BuLLeTIN 300
brockensis, Myophoria: Smith Plastoholotype
Smith, 1927, p. 110, pl. 96, figs. 25, 26
Shasta Co., Calif.; quarry SW end Brock Mt., between Squaw Creek
and Pitt River
Upper Triassic, Hosselkus Ls [holotype USNM 74173]
brooksi, Halobia: Smith Plastoholotype
Smith, 1927, p. 114, pl. 99, fig. 7
Chitina region, Alaska; W bank, Roadhouse Creek, 2 miles from
Kuskulana River USGS loc. 8153
Upper Triassic [holotype USNM]
budaense, Cardium (Granocardium): Shattuck Plastosyntype
Shattuck, 1903, p. 25, pl. 13, fig. 2
Near Austin, Texas
Cretaceous, Buda Ls
budaense, Cardium (Granocardium): Shattuck Plastosyntype
Shattuck, 1903. p. 25, pl. 13, fig. 3
near Austin, Texas
Cretaceous, Buda Ls
budaense, Cardium (Granocardium): Shattuck Plastosyntype
Shattuck, 1903, p. 25, pl. 13, fig. 4
near Austin, Texas
Cretaceous, Buda Ls
buwaldi, Petricola: Clark Plastoholotype
Clark, 1915, p. 471, pl. 60, fig. 6
Contra Costa Co., Calif.; SE of Walnut Creek UCMP loc. 1942
Miocene, San Pablo Fm eee UCMP 11657]
cahillensis, Leda: Arnold Holotype
Arnold, 1908a, p. 375, pl. 34, fig. 9
San Mateo Co., Calif. 2 miles W of Woodside on road to Kings Mt.
House
Lower Miocene, Vaqueros Fm_ [Arnold’s specimen 1065]
calaverasensis, Pecten (Patinopecten) haywardensis: Hall
Holotype
Hall, 1958, p. 51, pl. 2, fig. 2
Alameda Co., Calif.; La Costa Valley Qd, NW 1/4 NW 1/4 See. 11,
T5S,R1£E SU loc. 3245
Middle Miocene, Oursan Fm
calaverasensis, Pecten (Patinopecten) haywardensis: Hail
Paratype
Hall, 1958, p. 51, pl. 3, fig. 4
Alameda Co., Calif.; La Costa Valley Qd. NW 1/4, NW 1/4 Sec. 11,
T5S,R1E SU loc. 3245
Middle Miocene, Oursan Fm
calaverasensis, Pecten (Patinopecten) haywardensis: Hall
Paratype
Hall, 1958, p. 51, pl. 4, fig. 3
Alameda Co., Calif.; La Costa Valley Qd, NW 1/4 NW 1/4 Sec. 11,
T5S,R1£E SW loc. 3245
Middle Miocene, Oursan Fm
calcarea, Arca (Arca) trilineata: Grant and Gale Holotype
Grant and Gale, 1931, p. 140, pl. 2, figs. 6a, 6b
San Diego Co., Calif.; San Diego well
Middle Pliocene
calkinsi, Pecten (Chlamys): Arnold Paratypes
Arnold, 1906, p. 51
Ventura Co., Calif.; N side of Sisar Valley
Eocene, ‘“Tejo:"* Fm
68
125
127
126
53
54
54a
131
8042
8733
5249
5177
11
9714
30
STANFORD UNIVERSITY TyrrEs: SMITH 331
calli, Pecten (Plagioctenium): Hertlein Holotype
Hertlein, 1925a, p. 17, pl. 4, fig. 6
Baja California, Mexico; first arroyo E of Santiago SU loc. 53
Miocene?
calli, Pecten (Plagioctenium): Hertlein Paratypes
Hertlein, 1925a, p. 17, pl. 4, figs. 5, 7 (type 125)
Baja California, Mexico; Scammon Lagoon Qd, W side Elephant
Mesa _ SU loc. 60
Miocene? [paratype 125 = holotype of Pecten (Plagioctenium)
diminutivus Hertlein and Jordan]
calli, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 17
Baja California, Mexico; Arroyo Fortuna at Arroyo Refugio near
San Jose del Cabo SU loc. 63
Miocene?
callidus, Pecten (Plagioctenium): Hertlein Holotype
Hertlein, 1925a, p. 22, pl. 5, figs. 1, 5
Baja California, Mexico; Cedros Island SU loc. 116
Pliocene, Salada Fm
callidus, Pecten (Plagioctenium): Hertlein Paratypes
Hertlein, 1925a, p. 22, pl. 5, figs. 3 (type 54), 6 (type 54a)
Baja California, Mexico; Cedros Island SU loc. 116
Pliocene, Salada Fm
callidus, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 22
Baja California, Mexico; Scammon Lagoon Qd, mouth of big arroyo
NW of Elephant Mesa_ SU loc. 48
Pliocene, Salada Fm
canoa, Glycymeris: Pilsbry and Olsson Plastoholotype
Pilsbry and Olsson, 1941, p. 54, pl. 13, figs. 2, 2a
Punta Blanca, Ecuador
Pliocene, Canoa Fm_ [holotype ANSP 13669]
cardissoides, Inceramus: Goldfuss Plastoholotype
Goldfuss, 1826, p. 112, pl. 110, figs. 2a, 2b
Westphalia, Germany
Cretaceous [holotype 672, from BM(NH) ]
carmanahensis, Limopsis: Clark Holotype
Clark, 1925, p. 80, pl. 22, fig. 8
Vancouver Island, British Columbia, Canada; in sea cliff ca. 3 miles
W of Carmanah Point SU loc. NP 141
Oligocene
carmanahensis, Limopsis: Clark Paratype
Clark, 1925, p. 80
Vancouver Island, British Columbia, Canada; in sea cliff ca. 3 miles
W of Carmanah Point SU loc. NP 141
Oligocene
carrizoensis, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 59, pl. 4, figs. 1, 1a, 1b
San Diego Co., Calif.; Alverson Canyon
“Miocene,” Carrizo Fm
caryonautes, Transennella: Berry Holotype
Berry, 1963, p. 141. Illustrated in Keen, 1971, p. 166, fig. 391
Sinaloa, Mexico; near Mazatlan
catalinae, Pecten (Lyropecten) estrellanus: Arnold Holotype
Arnold, 1906, p. 76, pl. 20, figs. 3, 3a
Los Angeles Co., Calif.; Santa Catalina Island, near isthmus
Upper Miocene
So
Oo
bo
561
5816
31
3la
9903
5375
5394
399
8722
6048
6050
265
8087
24
BuLLeETIN 300
catalinae, Pecten (Lyropecten) estrellanus: Arnold Paratype
Arnold, 1906, p. 76, pl. 20, fig. 4
Los Angeles Co., Calif.; Santa Catalina Island, near isthmus
Upper Miocene
catherinae, Sphaerium?: Hannibal Holotype
Hannibal, 1912b, p. 132, pl. 7, fig. 20. Also in Taylor and Smith, 1971,
figs. 1, 5, 8 (as Pisidium)
Near Hawthorne, Nevada; hill on Belmont stage road
“Eocene” [Miocene, fide Taylor and Smith, 1971]
cerritensis, Pecten (Chlamys) latiauritus: Arnold Holotype
Arnold, 1906, p. 129, pl. 46, fig. 6
Los Angeles Co., Calif.; San Pedro
Pleistocene, upper San Pedro Fm
cerritensis, Pecten (Chlamys) latiauritus: Arnold Paratype
Arnold, 1906, p. 129, pl. 46, fig. 7
Los Angeles Co., Calif.; San Pedro
Pleistocene, upper San Pedro Fm
ceruleus, Pecten (Entolium): Smith Plastoholotype
Smith, 1927, p. 121, pl. 95, fig. 13
Baker Co., Ore.; Martin Bridge, Eagle River
Upper Triassic [holotype USNM 74158]
chehalisensis, Malletia: Arnold Paratype
Arnold, 1908a, p. 365. Ilustrated in Arnold, 1909, illust. 2, fig. 32
Santa Cruz Co., Calif.; Kings Creek, % mile above its confluence with
San Lorenzo River
Oligocene, San Lorenzo Fm [Arnold’s specimen 1062]
chehalisensis, Malletia: Arnold Paratype
Arnold, 1908a, p. 365
Santa Cruz Co., Calif.; Kings Creek, % mile above its confluence
with San Lorenzo River
Oligocene, San Lorenzo Fm
chicoensis, Isocardia: Waring Holotype
Waring, 1917, p. 62, pl. 8, fig. 3
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon, N of Simi fault
Upper Cretaceous, Chico Fm
circularis, Venus?: Meek and Hayden Plastoholotype
Meek and Hayden, 1856, p. 272. Illustrated in Meek, 1876, p. 190, pl.
17, fig. 8a (as Thetis?)
Valley Co., Montana; mouth of Milk River
Cretaceous, Bear Paw Fm
cistula, Lasaea: Keen Holotype
Keen, 1938, pp. 25-26, pl. 2, figs. 7-9
San Mateo Co., Calif.; Half Moon Bay, Moss Beach
cistula, Lasaea: Keen Paratype
Keen, 1938, pp. 25-26
San Mateo Co., Calif.; Half Moon Bay, Moss Beach
clallamensis, Solen: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 152, pl. 20, fig. 4
Clallam Bay, Wash.; sea cliffs 1.5 miles W of West Clallam SU loc.
NP 88
Oligocene? Sooke Fm
clarionense, Cardium (Laevicardium): Hertlein and Strong
Paratype
Hertlein and Strong, 1947a, p. 144
Gulf of California; Santa Inez Bay
clarkensis, Pecten: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 68. Illustrated in Wiedey, 1929b, pl. 2,
fig. 3
San Jose Qd, Calif.; 2.5 miles NE of Milpitas
Middle Cretaceous, Horsetown Fm
5096
29
15
14
210
10315
10265
395
STANFORD UNIVERSITY Types: SMITH 3515)
clivi, Bayleoidea: Palmer Paratype
Palmer, 1928a, p. 38
Vera Cruz, Mexico; Escamela Hill, Orizaba
Cretaceous
coalingaensis, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 97, pl. 4, figs. 4, 4a
Fresno Co., Calif.; near Coalinga
Pliocene
coani, Tellina: Keen Holotype
Keen, 19715 ps 21d, figs S12
Baja California, Mexico; Candelero Bay, near La Paz
cognata, Lutricola: Pilsbry and Vanatta Syntype
Pilsbry and Vanatta, 1902, p. 556. I'lustrated in Keen, 1971 p. 225, fig.
557 left (as Florimetis)
Galapagos Islands; Albemarle Island, Tagus Cove
columbianum, Pecten: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 139, pl. 23, fig. 1
Vancouver Island, British Cclumbia, Canada; Jordan River, 2 miles
W of Sherringham Point, sea cliffs at mouth of Fossil Creek SU loc.
NP 130
Upper Oligocene or Lower Miocene, Sooke Fm
condoni, Pecten (Amusium): Hertlein Holotype
Hertlein, 1925b, p. 41, pl. 4, figs. 8, 9
Ventura Co., Calif.; Piru Qd, E of Timber Canyon, Sec. 29, T 4 N,
R 20 W_ SU loc. NP 244
Pleistocene, Saugus Fm
condoni, Pecten (Amusium): Hertlein Paratypes
Hertlein, 1925b, p. 41
West Wishkah, Wash.; at Dam 35 SU loc. 148
Miocene, Montesano Fm
cooperi, Pecten (Plagioctenium): Arnold Holotype
Arnold, 1906, p. 124, pl. 49, fig. 2
San Diego Co., Calif.; Pacific Beach
Pliocene, San Diego Fm [Renamed Pecten invalidus by Hanna, 1924,
pats
cooperi, Pecten (Plagioctenium): Arnold Paratype
Arnold, 1906, p. 124, pl. 49, fig. 3
San Diego Co., Calif.; Pacific Beach
Pliocene, San Diego Fm
cooperi, Pecten (Plagioctenium): Arnold Paratype
Arnold, 1906, p. 124, pl. 49, fig. 4
San Diego Co., Calif.; Pacific Beach
Pliocene, San Diego Fm
cor, Trinacria: Popenoe Plastoholotype
Popenoe, 1937, p. 45, figs. 1, 3
Santa Ana Mts., Calif.; CIT loc. 974
Cretaceous, Campanian [holotype UCLA 40615]
Cerbicula n. sp.: Addicott Holotype
Addicott, 1976, p. 107, pl. 3, fig. 15
SW Wash.; SU loc. NP 220
Upper Miocene, Wishkahan stage
cordata, Macrocallista: Waring Holotype
Waring, 1917, p. 62, pl. 8, fig. 1
Ventura Co., Calif.; Bell’s Canyon, N of Simi fault
Upper Cretaceous, Chico Fm
334
9872
28
8337
7567
10320
826
7566
550
393
394
8056
8731
BuLLeETIN 300
cordillerana, Halobia: Smith Plastoholotype
Smith, 1927, p. 114, pl. 99, fig. 2
Alaska; S bank of Yukon River, 1 mile above Nation River USGS
loc. 8897, bed 86
Upper Triassic [holotype USNM]
cornwalli, Pecten (Chlamys): Clark and Arnold Holotype
Clark and Arnold, 1923, p. 140, pl. 25, fig. 1
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Kirby Creeks, W of Otter Point SU loc.
NP 129
Upper Oligocene or Lower Miocene, Sooke Fm
cornwalli, Pecten (Chlamys): Clark and Arnold Paratype
Clark and Arnold, 1923, p. 140
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Kirby Creeks, W of Otter Point SU loc.
NP 129
Upper Oligocene or lower Miocene, Sooke Fm
coronadosensis, Protothaca?: Durham Plastoholotype
Durham, 1950, p. 86, pl. 22, figs. 2, 9, 11
Coronado Island, Gulf of California UCMP loc. A3549 ~
Upper Pliocene [holotype UCMP 32596]
corrugata, Tepeyacia: Palmer Paratype
Palmer, 1928a, p. 46
Vera Cruz, Mexico; Orizaba
Cretaceous, Turonian
corrugatus, Clisocolus: Popenoe Plastoholotype
Popenoe, 1937, p. 390-391, pl. 47, figs. 9, 10, 12
Santa Ana Mts., Calif.; CIT loc. 302
Upper Cretaceous, Turonian, Ladd Fm, Baker Mbr [holotype UCLA
40646]
corteziana, Glycymeris: Dall Paratype
Dall, 1916, p. 402
Cortez Bank, off southern Calif. US Bur. Fish. loc. 2518
costata, Caprinuloidea: Palmer Paratype
Palmer, 1928a, p. 62, pl. 11, fig. 2
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
costata, Caprinuloidea: Palmer Paratype
Palmer, 1928a, p. 62
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
cowperi, Pecten (Propeamusium): Waring Syntype
Waring, 1917, p. 63, pl. 7, fig. 2
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon. N of Simi fault
Upper Cretaceous, Chico Fm
cowperi, Pecten (Propeamusium): Waring Syntype
Waring, 1917, p. 63, pl. 7, fig. 1
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon. N of Simi fault
Upper Cretaceous, Chico Fm
craneana, Semele: Hertlein and Strong Paratype
Hertlein and Strong, 1949b, p. 241
S end of Gulf of California; Arena Bank, in 50 fms
crippsi, Inoceramus: Mantell Plastoholotype
Mantell, 1822, p. 133, pl. 27, fig. 11
Sussex, England; Offham, near Lewes
Cretaceous, Cenomanian, Chalk Marl [holotype BM(NH) 5893]
36
37
37a
37b
37¢
37d
94
9712
8808
8338
7305
10037
10037a
5281
7865
7865a
STANFORD UNIVERSITY TypEs: SMITH 335
cristobalensis, Pecten (Plagioctenium) Hertlein Holotype
Hertlein, 1925a, p. 19, pl. 3, figs. 2, 3
Baja California, Mexico; San Cristobal Bay Qd, 3 miles SE of Turtle
Bay, SU loc. 49
Pliocene, Salada Fm, uppermost beds
cristobalensis, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 19, pl. 3, fig. 1
Baja California, Mexico; San Cristobal Bay Qd, 3 miles SE of Turtle
Bay SU loc. 49
Pliocene, Salada Fm, uppermost beds
cristobalensis, Pecten (Plagioctenium): Hertlein Paratypes
Hertlein, 1925a, p. 19
Baja California, Mexico; San Cristobal Bay Qd, 3 miles SE of Turtle
Bay SU loc. 49
Pliocene, Salada Fm, uppermost beds
cristobalensis, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 19
Baja California, Mexico; Scammon Lagoon Qd, arroyo NW of Ele-
phant Mesa_ SU loc. 48
Pliocene, Salada Fm
cristulata: Tellidorella: Berry Holotype
Berry, 1963, p. 140. Illustrated in Keen, 1971, p. 106, fig. 236
Sonora, Mexico; off Puerto Libertad, in 40 fms
crockeri, Solen: Hertlein and Strong Paratype
Hertlein and Strong, 1950, pp. 225-226
Gulf of Fonseca, Nicaragua; Monypenny Point, lat. 13° 03’N., long.
87° 30’W, in 5-16 fms
crooki, Pitar (Lamelliconcha) Clark and Anderson Plastoholotype
Clark and Anderson, 1938, p. 946, pl. 1, figs. 4, 5
Yuba Co., Calif.; E of Marysville UCMP loc. A-1889
Upper Eocene, Wheatland Fm_ [holotype UCMP 11217]
crooki, Tellina: Nelson Plastoholotype
Nelson, 1925, p. 415, pl. 53, fig. 4
Ventura Co., Calif.; S of Simi Valley UCMP loc. 3776
Eocene, “Martinez” Fm _ [holetype UCMP 30523]
cultrata, Amerycina: Keen Holotype
Keen, 1971, p. 135, fig. 310
Near La Paz, Baja California, Mexico; off Isla Partida, Espiritu
Santo Id., in 5-33 m
cultrata, Amerycina: Keen Paratype
Keen, 1971, p. 135
Near La Paz, Baja California, Mexico; off Isla Partida, Espiritu
Santo Id., in 5-33 m
cumshewensis, Parallelodon (Nanonavis): Reinhart
Plastolectotype
Reinhart, 1937a, p. 173. Lectotype illustrated by Whiteaves, 1884, p.
235, pl. 31, figs. 8a, 8b [as Grammatodon inornatus Meek and Hayden]
Queen Charlotte Islands, British Columbia, Canada; N shore of Cum-
shewa Inlet
Middle Cretaceous [lectotype, selected by Reinhart, 1937, is Geol.
Surv. Canada specimen 4915]
cyclia, Adontorhina: Berry Holotype
Berry, 1947b, p. 260, pl. 26, figs. 1, 2
San Pedro, Calif.; Hilltop Quarry
Pleistocene, Lomita Marl
cyclia, Adontorhina: Berry Paratype
Berry, 1947b, p. 260
San Pedro, Calif.; Hilltop Quarry
Pleistocene, Lomita Marl
336
7295
7296
8584
6946
100
100a
100b
225
227
228
229
8328
5238
9877
BuLetin 300
cylindrica, Psammobia (?): Dickerson Plastosyntype
Dickerson, 1914, p. 139, pl. 12, fig. 2a
Lake Co.. Calif.; near Lower Lake UCMP loc. 780
Eocene, Martinez Fm_ [syntype UCMP 11678]
cylindrica, Psammobia (?): Dickerson Plastosyntype
Dickerson, 1914, p. 139, pl. 12, fig. 2b
Lake Co., Calif.; near Lower Lake UCMP loc. 780
Eocene, Martinez Fm [syntype UCMP 11677]
cylista, Botula: Berry Holotype
Berry, 1959, p. 107. Illustrated in Keen, 1971, p. 74, fig. 155 (lower)
Sinaloa, Mexico; Mazatlan, Las Gaviotas Beach
cymata, Psephidia: Dall Paratype
Dall, 1913, p. 593
Baja California. Mexico; San Bartolomé [Turtle Bay]
dallasi, Pecten (Plagioctenium): Jordan and Hertlein Paratypes
Jordan and Hertlein, 1926a, p. 213
Baja California, Mexico; canyons 1 or 2 miles from San Antonio
Point CAS loc. 795
Upper Pliocene, Salada Fm? ‘
dalli, Myadesma: Clark Holotype
Clark, 1922, p. 117, pl. 14, figs. 3a, 3b
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
dalli, Myadesma: Clark Paratype
Clark, 1922, p. 117, pl. 13, fig. 6
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
dalli, Myadesma: Clark Paratype
Clark, 1922, p. 117, pl. 13, fig. 2
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
dalli, Myadesma: Clark Paratype
Clark, 1922, p. 117, pl. 13, fig. 4
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
dalli, Nucula (Acila): Arnold Paratypes
Arnold, 1908a, p. 364
Santa Cruz Co., Calif.; Santa Cruz sheet, Big Basin, Blooms Creek,
SW 1/4 Sec. 9, T 9S,R3 W _ Arnold loc. C-415
Oligocene, San Lorenzo Fm
dalli, Solemya: Clark Holotype
Clark, 1925, p. 73, pl. 22, fig. 3
Twin, Wash.; sea cliffs W of West Twin River SU loc. NP 120
Oligocene, Blakeley Fm
dalliana, Halobia: Smith Plastoholotype
Smith, 1927, p. 115, pl. 98, fig. 5
Keku Islet No. 1, Admiralty Island, Alaska; Herring Bay USGS loc.
10196
Upper Triassic, upper Karnic [holotype USNM]
7979
7970
32
32a
10322
7250
7246
411
9294
9295
9916
5480
5481
5482
192
STANFORD UNIVERSITY Tyres: SMITH 337
dauleana, Chione (Chionopsis): Marks Paratypes
Marks, 1951, p. 81
SW Ecuador; Daule Basin, near Pedro Carbo
Middle Miocene, Daule Fm
dauleana, Noetia: Marks Paratypes
Marks, 1951, p. 52
SW Ecuador; Manabi Province, E of village of Calceta
Middle Miocene, Daule Fm
delosi, Pecten (Chlamys) latiauritus: Arnold Holotype
Arnold, 1906, p. 130, pl. 46, figs. 9, 9a
Los Angeles Co., Calif.; Deadman Island
Pleistocene, San Pedro Fm
delosi, Pecten (Chlamys) latiauritus: Arnold Paratype
Arnold, 1906, p. 130, pl. 46, figs. 10, 10a
Los Angeles Co., Calif.; Deadman Island
Pleistocene, San Pedro Fm
delta, Isocardia: Popenoe Plastoholotype
Popenoe, 1937, p. 389, pl. 47, figs. 7, 8
Santa Ana Mts.. Calif.
Cretaceous, Turonian [holotype UCLA 40643]
diabloensis, Chione: Clark Plastoholotype
Clark, 1915, p. 468, pl. 58, fig. 4
Contra Costa Co., Calif.; E of town of Walnut Creek UCMP Joc.
1492
Miocene, San Pablo Fm_ [holotype UCMP 12325]
diabloensis, Tellina: Clark Plastoholotype
Clark, 1915, p. 471, pl. 61, fig. 5
Contra Costa Co., Calif.; SE of Walnut Creek UCMP loc. 1478
Miocene, San Pablo Fm [holotype UCMP 11531]
diabloensis, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 27, pl. 5, figs. 5-7
Contra Costa Co., Calif.; Brentwood, Marsh Creek
Eocene, Meganos Fm, D Mbr
dibbleei, Meretrix: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 203, pl. 34, figs. 8, 9
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek SU loc. 2217
Eocene, Sacate-Gaviota Fm
dibbleei, Meretrix: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 203, pl. 35, figs. 1, 2
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek SU loc. 2217
Eocene, Sacate-Gaviota Fm
digglesi, Cardiamorpha?: Smith Plastoholotype
Smith, 1927, p. 111, pl. 94, fig. 8
Shasta Co., Calif.; old quarry SW end Brock Mt., between Squaw
Creek and Pit River
Upper Triassic, Hosselkus Ls [holotype USNM 74141]
digona, Monotis: Kittl Plastosyntypes
Kittl, 1912, p. 174, pl. 10, figs. 16 (type 5480), 17 (type 5481), 18
(type 5482)
Austria; Siriuskogel, Ischal
Upper Triassic, Noric
dilatata, Corbula: Waring Holotype
Waring, 1917, p. 92, pl. 15, fig. 2
Ventura Co., Calif.; McCray Wells
Eocene, Tejon Fm [Renamed Corbula complicata by Hanna, 1924,
p. 163]
125
127
7309
8005
9285
9713
322
9960
7315
6000a
6000b
5994
6514
5198
BuL.etin 300
diminutivus, Pecten (Plagioctenium): Hertlein and Jordan
Holotype
Hertlein and Jordan, 1927, p. 623. Illustrated in Hertlein, 1925a, p. 16,
pl. 4, figs. 5, 7 (as Pecten (Plagioctenium) calli Hertlein, 1925)
Baja California, Mexico; Scammon Lagoon Qd, W side Elephant Mesa
SU loc. 60
Miocene, Isidro Fm
diminutivus, Pecten (Plagioctenium): Hertlein and Jordan
Paratype
Hertlein and Jordan, 1927, p. 623
Baja California, Mexico; Scammon Lagoon Qd, W side Elephant Mesa
SU loc. 60
Miocene, Isidro Fm
domenginensis, Tellina: Vokes Plastoholotype
Vokes, 1939, p. 91, pl. 14, fig. 14
Fresno Co., Calif.; Domengine Creek UCMP loc. 3315
Eocene, Domengine Fm_ [holotype UCMP 15694]
durhami, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 23, pl. 7, figs. 1, 2 :
Ventura Co., Calif.; 1.5 miles W of Vickers Hot Spring
Lower Eocene, Juncal Fm
effingeri, Lucina (Here): Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 201, pl. 33, fig. 9
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek
Eocene, Sacate-Gaviota Fm
electilis, Crenimargo: Berry Holotype
Berry, 1963, p. 140. Illustrated in Keen, 1971, p. 133, fig. 304
Colima, Mexico; Playa las Hadas, 5 miles N of Manzanillo
elegans, Septifer: Waring Paratype
Waring, 1917, p. 79, pl. 14, fig. 2
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
elimata, Macoma: Dunnhill and Coan Paratype
Dunnhill and Coan, 1968, pp. 1-9
Near Victoria, British Columbia, Canada; N of Moresby Island, fine
silty sd
eoundulata, Gari: Vokes Plastoholotype
Vokes, 1939, p. 93, pl. 14, fig. 23
Fresno Co., Calif.; N of Domengine Creek UCMP loc. A-820
Eocene [holotype UCMP 15707]
equilateralis, Arca (Cucullaea): Meek Plastosyntypes
Meek, 1864a, pp. 39-40. Illustrated in Meek, 1876b, p. 357, pl. 2, figs.
6, 6a
Vancouver Island, British Columbia, Canada; Nanaimo?
Cretaceous [syntypes USNM 12386]
equilateralis, Mactra (Mactrotoma) californica: Clark
Plastoholotype
Clark, 1932, p. 819, pl. 14, fig. 8
Alaska; near Yakataga River, head of Oil Creek UCMP loc. 3870
Upper Oligocene, Poul Creek Fm_ [holotype UCMP 30390]
etheringtoni, Loxocardium: Effinger Paratype
Effinger, 1938, p. 370
Lewis Co., Wash.; on Cowlitz River, Sec. 25, T 11 N, R 2 W
Lower Oligocene, Gries Ranch Fm
eugenensis, Mulinia: Clark Plastoholotype
Clark, 1925, p. 104, pl. 14, fig. 2
Lane Co., Ore.; 3 miles S of Eugene
Oligocene [holotype UCMP 30372]
8505
8304
7282
16
17
10059
8089
6524
5138
8051
794
10342
6222
STANFORD UNIversity |yprres: SMITH 339
euterpes, Pecten (Leptopecten): Berry Holotype
Berry, 1957, p. 75. Illustrated in Keen, 1971, p. 91, fig. 197
Off Acapulco, Mexico, 6-10 fms
ezoense, Nemocardium (Arctopratulum): Takeda Paratypes
Takeda, 1953, p. 82
Hokkaido, Japan; Kushiro Province, along Koikatahbrokachoro
River, upper course of Charo River, Shiranuka-gun
Upper Oligocene, Poronai Fm
fausta, Semele: Nomland Plastoholotype
Nomland, 1917a, p. 233, pl. 9, figs. 3a, 3b
Fresno Co., Calif.; Zapato Creek UCMP loc. 2991
Pliocene, Etchegoin Fm [holotype UCMP 11102]
fernandoensis, Pecten (Pseudamusium) vancouverensis: Hertlein
Holotype
Hertlein, 1925b, p. 43, pl. 4, fig. 7
Ventura Co., Calif.; 1.5 miles N of Ventura on Ventura River, 1/4
mile S of Taylor Well No.1 SU loc. 155
Lower Pliocene, lower Fernando Fm
fernandoensis, Pecten (Pseudamusium) vancouverensis: Hertlein
Paratype
Hertlein, 1925b, p. 43, pl. 4, fig. 6
Long Beach, Calif.; drill core, 2800’ deep, about 4500’ NW of Signal
Hill, 500’ E of Orange Ave., 750’ N of Willow St.
Lower Pliocene
fitchi, Penitella: Turner Paratype
Turner, 1955, p. 71-74
Baja California, Mexico; San Bartolomé [Turtle Bay]
fonsecana, Mactra (Micromactra): Hertlein and Strong Paratype
Hertlein and Strong, 1950, p. 232
Gulf of Fonseca, Nicaragua; Potosi and Monypenny Point
forma, Chama sinuosa: Pilsbry and McGinty Paratype
Pilsbry and McGinty, 1938, p. 76
Palm Beach Co., Fla.; rock reef S of Boynton Inlet
freudenbergi, Ostrea: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 622, pl. 17, fig. 9; pl. 18, fig. 4
Baja California, Mexico; above San Gregorio Lagoon, on trail from
Arroyo Mesquital to La Purisima, in Turritella bed SU loc. 59
Miocene, Isidro Fm
frizzelli, Pitar (Lamelliconcha): Hertlein and Strong Paratype
Hertlein and Strong, 1948, p. 176
Gulf of California, near Gorda Banks, 50 fms
frustra, Spisula pittsburgensis: Tegland Paratype
Tegland, 1933, p. 121, pl. 9, fig. 12
Puget Sound, Wash.; Bainbridge Island, beach between S side of en-
trance to Blakeley Harbor and Restoration Point SU loc. NP 103
Upper Oligocene, Blakeley Fm
gabbiana, Mactra: Anderson Plastoholotype
Anderson, 1902, p. 74, pl. 7, fig. 156. Also in Saul, 1974, pp. 1084-1087,
pl. 1, fig. 16; pl. 2, fig. 10; pl. 3, fig. 17 [as Cymbophora gabbiana
(Anderson) ]
Siskiyou Co., Calif.; Henley and Willow Creek
Cretaceous, Turonian, “lower Chico beds” Hornbrook Fm
galapagensis, Lima: Pilsbry and Vanatta Syntype
Pilsbry and Vanatta, 1902, p. 556
Galapagos; Albemarle Island, Tagus Cove
340
10314
5907
5907a
5223
59224
5225
9385
7573
556
9882
9917
6135
9834
553
7568
6582
BuLLeETIN 300
gamma, Crassatella: Popenoe Plastoholotype
Popenoe, 1937, p. 388, pl. 46, figs. 13, 14
Santa Ana Mts., Calif.; CIT loc. 1069
Cretaceous, Turonian, Ladd Fm _ [holotype UCLA 40636]
gardneri, Yoldia: Oldroyd Holotype
Oldroyd, 1935, p. 14, fig. 1
Vancouver Island, British Columbia, Canada; Pender Harbor, Gard-
ner Bay
gardneri, Yoldia: Oldroyd Paratype
Oldroyd, 1935, p. 14
Vancouver Island, British Columbia, Canada; Pender Harbor, Gard-
ner Bay
gastonensis, Tivela: Clark Holotype
Clark, 1925,-p. 93; pi. 19: fic: 1
Gaston, Ore.; county quarry, Scroggins Canyon SU loc. NP 295
Oligocene, lower Astoria Ss
gastonensis, Tivela: Clark Paratype
Clark31925-993,p iho ii 2
Gaston, Ore.; county quarry, Scroggins Canyon SU loc. NP 295
Oligocene, lower Astoria Ss
gastonensis, Tivela: Clark Paratype
Clark 1925 p293; (pl19; fics 3
Gaston, Ore.; county quarry, Scroggins Canyon SU loc. NP 295
Oligocene, lower Astoria Ss
gayi, Semele: Arnold Paratype
Arnold, 1908a, p. 360
San Mateo Co., Calif.; between headwaters of San Lorenzo River and
Pescadero Creek
Eocene
gherzii, Agria: Palmer Syntype
Palmer, 1928a, p. 78, pl. 15, figs. 4, 5
Colima, Mexico; Paso del Ric
Cretaceous, Cenomanian
gherzii, Agria: Palmer Paratype
Palmer, 1928a, p. 78
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
gigantea, Halobia: Smith Plastoholotype
Smith, 1927, p. 116, pl. 93, fig. 6
Shasta Co., Calif.; E side Brock Mt., Bear Cove
Upper Triassic, Juvavites beds of Bear Cove [holotype USNM]
gleimi, Cardinia: Smith Plastoholotype
Smith, 1927, p. 110, pl. 96, fig. 7
Shasta Co., Calif.; N fork Squaw Creek, 3 miles N of Kellys Ranch
Upper Triassic, Hosselkus Ls [holotype USNM 74165]
globula, Sphenia: Dall Paratype
Dall, 1919b, p. 370. Illustrated in Schenck, 1945, p. 519, pl. 67, figs. 5-8
Bolinas Bay, Calif.
goesi, Kellyella: Odhner Paratypes
Odhner, 1960, p. 397
Off St. Martin, West Indies, 300 fms
gracilis, Caprinuloidea perfecta: Palmer Paratypes
Palmer, 1928a, p. 60
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
granti, Pseudochama: Strong Paratypes
Strong, 1934, p. 137
Orange Co., Calif.; dredged off Corona del Mar
6940
7260
9914
5578
6513
8295
8296
8297
5443
8052
5338
8359
STANFORD University Types: SMITH 341
granulata, Pandora: Dall Paratype
Dall, 1915b, p. 449. Illustrated in Keen, 1971, p. 289, fig. 739 below
Off La Paz, Baja California, Mexico: Gulf of California
gravidus, Solen: Clark Plastoholotype
Clark, 1918, p. 156, pl. 10, fig. 7
Contra Costa Co., Calif.; SW of Walnut Creek UCMP loc. 1131
Oligocene, San Ramon Fm_ [holotype UCMP 11138]
gravinaensis, Cassianella: Smith Plastoholotype
Smith, 1927, p. 112, pl. 101, figs. 4, 5
Gravina Island, SE Alaska
Upper Triassic [holotype USNM 74195]
gregaria, Horipleura: Palmer Paratypes
Palmer, 1928a, p. 49
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
gregoryi, Avicula: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 69. Illustrated in Wiedey, 1929b, pl. 1,
fig. 1
Alameda Co., Calif.; Tesla Qd, 1.5 miles § 10° W of Carnegie
Middle Cretaceous, Horsetown Fm
grewingki, Mya (Arenomya): Makiyama Paratype
Makiyama, 1934, p. 156
Cape Maly, Matchgar coast, Russian Sachalin
“Oligocene,” Asagaian Fm
griesensis, Ostrea: Effinger Paratype
Effinger, 1938, p. 368
Lewis Co., Wash.; on Cowlitz River, Sec. 25, T 11 N, R2 W
Oligocene, Gries Ranch Fm
griphus, Nemocardium (Arctopratulum): Keen Holotype
Keen, 1954, p. 318, pl. 29, figs. 14, 17
Grays Harbor Co., Wash.; middle fork Wishkah River, 14 miles N of
Aberdeen SU loc. NP 243
Middle Miocene, Astoria Fm
griphus, Nemocardium (Arctopratulum): Keen Paratype
Keen, 1954, p. 318, pl. 29, fig. 12, text fig. 4
Grays Harbor Co., Wash.; middle fork Wishkah River, 14 miles N of
Aberdeen SU loc. NP 243
Middle Miocene, Astoria Fm
griphus, Nemocardium (Arctopratuium): Keen Paratype
Keen, 1954, p. 318, text fig. 3
Grays Harbor Co., Wash.; middle fork Wishkah River, 14 miles N of
Aberdeen SU loc. NP 243
Middle Miocene, Astoria Fm
guadalupensis, Glycymeris: Strong Paratype
Strong, 1938, p. 213
Off Guadalupe Island, Mexico; 9-15 fms
guatulcoensis, Chione: Hertlein and Strong Paratypes
Hertlein and Strong, 1948, p. 182
Off Port Guatulco, Mexico; 15° 44’ 28” N, 96° 07’ 51” W, 7 fms
guineensis, Nucula: Thiele Paratype
Thiele, 1931, p. 193
Gulf of Guinea; lat. 3° 10’ N, long. 5° 28’ W, 1139 fms
hadra, Venericardia: Dall Syntypes
Dall, 1903b, p. 1429
Calhoun Co., Fla.; Chipola River, 1 mile below Bailey’s Ferry
“Oligocene,” in riverbank above white Is bed
41
95
8516
514
8302
8303
234
5248
5246
230
231
BuLLETIN 300
hakei, Pecten (Plagioctenium): Hertlein Holotype
Hertlein, 1925a, p. 18, pl. 4, fig. 1
Baja California, Mexico; Turtle Bay SU loc. 47
Pliocene, Salada Fm
hakei, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 18, pl. 4, fig. 3
Baja California, Mexico; Turtle Bay SU loc. 47
Pliocene, Salada Fm
hakei, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 18
Baja California, Mexico; Ballenas Bay Qd, N edge of tilted mesa ca.
5 miles N of Abreojos Point SU lec. 46
Pliocene, Salada Fm
hancocki, Lithophaga (Leiosolenus): Soot-Ryen Paratype
Soot-Ryen, 1955, p. 102. Illustrated in Keen, 1971, p. 68, fig. 141
Galapagos: Isla Onslow, N of Isla Floreana (Charles Island)
hannai, Anomia: Wiedey Holotype
Wiedey, 1929c, p. 280, pl. 21, fig. 1
Monterey Co., Calif.; Val Celico, W of Pleyto SU loc. 449
Lower Miocene, Vaqueros Fm
hannibali, Clinocardium: Keen Holotype
Keen, 1954, p. 324, pl. 29, fig. 16
Aberdeen, Wash.; Chehalis and Summit Streets SU loc. NP 235
Mio-Pliocene, Montesano Fm
hannibali, Clinocardium: Keen Paratype
Keen, 1954, p. 324, text fig. 9
Aberdeen, Wash.; Chehalis and Summit Streets SU loc. NP 235
Mio-Pliocene, Montesano Fm
hannibali, Mytilus: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 142, pl. 16, fig. 3
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, sea cliffs at mouth of Fossil Creek SU loc.
NP 130
Oligocene? Sooke Fm
hannibali, Nucula: Clark Holotype
Clark 1925) pe'/3; ple Sy tig. 2
W of Gettysburg, Wash.; sea cliffs at mouth of Duncan Creek SU
loc. NP 90
Oligocene, Blakeley Fm
hannibali, Paphia staleyi: Howe Holotype
Howe, 1922, p. 98, pl. 10, figs. 1, 4
Scotia, Calif.; Eel River valley between Scotia and Nanning Switch
SU loc. NP 82, Arnold loc. C-13
Pliocene, Wildcat Fm
hannibali, Phacoides (Lucinoma): Clark Holotype
Clark, 1925, p. 89, pl. 22, figs. 2, 4
Wash.; bluff on Chehalis River below Porter SU loc. NP 53
Oligocene, Porter Fm
hannibali, Spisula (Hemimactra): Clark and Arnold Holotype
Clark and Arnold, 1923, p. 153, pl. 19, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; W of Otter Point, sea
cliffs between mouths of Muir and Coal Creeks SU loc. NP 129
Oligocene, Sooke Fm
hannibali, Spisula (Hemimactra): Clark and Arnold Paratype
Clark and Arnold, 1923, p. 153, pl. 19, fig. 4
Vancouver Island, British Columbia, Canada; W of Otter Point, sea
cliffs between mouths of Muir and Coal Creeks SU loc. NP 129
Oligocene, Sooke Fm
5253
408
5832
505
48
5475
5476
5477
5343
5344
19
522
fod
SranForp UNIVERSITY Tyres: SMITH 343
hannibali, Venericardia (Cyclocardia): Clark Holotype
Clark, 1925, p. 88, pl. 19, figs. 6, 7
3/4 mile above Porter, Wash.; shaly ss bluffs along Porter Creek
SU loc. NP 54
Middle Oligocene, Sooke Fm
harfordus, Pecten (Camptonectes): Davis Syntype
Davis, 1913, p. 456, fig. 6
San Luis Obispo Co., Calif.; 6 miles N of Port Harford
Jurassic, San Luis Fm
harfordus, Pecten (Camptonectes): Davis Syntype
Davis, 1913, p. 456, figs. 3, 5
San Luis Obispo Co., Calif.; 6 miles N of Port Harford
Jurassic, San Luis Fm
haroldiana, Gonidea angulata: Dall Paratype
Dall, 1908a, p. 500. Illustrated in Hannibal, 1912a, fig. 4 and Hanni-
bal, 1912b, pl. 6, fig. 10
Coyote Creek, near San Jose, Calif.
harrigani, Pholadomya: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 77. Illustrated in Wiedey, 1929b, pl. 1,
fig. 5
Alameda Co., Calif.; Tesla Qd, Altamont, black shale in Western
Pacific R.R. cut
Upper Cretaceous, Chico Fm
hartmanni, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925a, p. 8, pl. 1, figs. 4, 6
Baja California, Mexico; Arroyo Mesquital SU loc. 54
Lower Pliocene?
haveri, Monotis: Kittl Plastosyntypes
Kitt], 1912, p. 171, pl. 10, figs. 7 (type 5475), 8 (type 5476), 9 (type
5477)
Upper Austria; Rossmoos bei Goisern
Upper Triassic, Noric [syntypes at Geolog.-Paleont. Abtlg. Naturh.
Staats—Museum, Wien]
hawleyi, Arca (Arca): Reinhart Holotype
Reinhart, 1943, p. 21, pl. 2, figs. 20, 22
Santa Barbara Co., Calif.; Lompoc Qd, E side of Nojoqui Creek, 3
miles N of Gaviota Pass SU loc. 834
Eocene, Gaviota Fm
hawleyi, Arca (Arca): Reinhart Paratype
Reinhart, 1943, p. 21
Santa Barbara Co., Calif.; Lompoc Qd, E side of Nojoqui Creek, 3
miles N of Gaviota Pass SU loc. 834
Eocene, Gaviota Fm
hawleyi, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925b, p. 40, pl. 4, fig. 5
Santa Barbara Co., Calif.; Santa Inez Mts.
Miocene, Vaqueros Fm
hawleyi, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925b, p. 40, pl. 4, fig. 4
Santa Barbara Co., Calif.; Santa Inez Mts.
Miocene, Vaqueros Fm
healeyi, Pecten (Patinopecten): Arnold Paratype
Arnold, 1906, p. 103, pl. 37, fig. 2
San Mateo Co., Calif.; San Gregorio
Pliocene, Purisima Fm
heimi, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925a, p. 9, pl. 1, fig. 3
Baja California, Mexico; S part of Arroyo San Gregorio SU loc. 65
Pliocene ?
47
8083
455
6531
452
5160
8340
9516
7792
20
21
22
9912
10
BuLuetin 300
heimi, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925a, p. 9, pl. 1, fig. 3
Baja California, Mexico; § part of Arroyo San Gregorio SU loc. 65
Pliocene?
helgolandicus, Mimoceramus: Heinz Plastoholotype
Heinz, 1934, p. 728, pl. 61, fig. 2, text fig. 1
North Sea; Helgoland Island
Cretaceous, Campanian [holotype at Geologisches Staatinstut, Ham-
burg]
hemphilli, Gonidea: Hannibal Holotype
Hannibal, 1912b, p. 128, pl. 7, fig. 19. Also in Taylor and Smith, 1971,
figs. 38, 39
Berkeley Hills, Calif.; Telegraph Canyon, water tunnel
Miocene [Pliocene, fide Taylor and Smith, 1971, p. 310]
hemphillii, Asthenothaerus: Dall Paratypes
Dall, 1886, p. 308
Marco, Fla., 2 fms
herrei, Margaritana: Hannibal Holotype
Hannibal, 1912b, p. 121, pl. 7, fig. 17. Also in Taylor and Smith, 1971,
figs. 12, 14 (as “Margaritifera’’)
Tesla, Calif.; 1/4 mile above Carnegie Pottery Plaint, in cut along
Western Pacific R.R., Corral Hollow
Eocene
hertleini, Pteria: Wiedey Holotype
Wiedey, 1928, p. 133, pl. 21, fig. 1
Monterey Co., Calif.; Los Vaqueros Valley SU loc. 200
Miocene, Vaqueros Fm _ [Wiedey’s specimen 434]
hertleini, Semele: Durham Plastoholotype
Durham, 1950, p. 90, pl. 24, fig. 6; pl. 25, fig. 7
Gulf of California; Coronado Island UCMP loc. A 3548
Pleistocene [holotype UCMP 30367]
hesperius, Pitar (Lamelliconcha): Berry Holotype
Berry, 1960, p. 115. Illustrated in Keen, 1971, p. 174, fig. 415
Near Mazatlan, Mexico
hilli, Cardita: Willett Paratype
Willett, 1944a, p. 19
Orange Co., Calif.; mesa at head of Newport Bay
Upper Pleistocene
hodgei, Pecten (Chlamys): Hertlein Holotype
Hertlein, 1925b, p. 42, pl. 4, fig. 2
Coalinga area, Calif.; Sec. 20,T 19 S,R15E
Miocene, Santa Margarita Fm
hodgei, Pecten (Chlamys): Hertlein Paratype
Hertlein, 1925b, p. 42, pl. 4, fig. 1
Coalinga area, Calif.; Sec. 20,T 19 S,R15E
Miocene, Santa Margarita Fm
hodgei, Pecten (Chlamys): Hertlein Paratype
Hertlein, 1925b, p. 42
Coalinga area, Calif.; Sec. 20,T 19S, R15E
Miocene, Santa Margarita Fm
humboldtensis, Myophoria: Smith Plastoholotype
Smith, 1927, p. 110, pl. 96, fig. 7
Humboldt Range, Nevada; Mulberry Canyon
Upper Triassic, Star Peak Fm [holotype USNM 74174]
hydei, Pecten (Chlamys) sespeensis: Arnold Holotype
Arnold, 1906, p. 69, pl. 5, figs. 3a-3c
Monterey Co., Calif.; Lynch’s Mt.
Lower Miocene, Vaqueros Fm
8629
5833
5162
7854
5168
8339
424
8253
5101
5348
5349
814
STANFORD UNIVERSITY Tyres: SMITH 345
hyphalopilema, Anadara (Scapharca): Campbell Holotype
Campbell, 1962, p. 152, figs. 2, 4, 5, 7, 8
Guaymas, Mexico; near Cabo Haro, off Catalina Bay, 18-20 fms
idahoense, Pisidium: Roper Paratypes
Roper, 1890, p. 85
Old Mission, Idaho
impavida, Arca: Wiedey Holotype
Wiedey, 1928, p. 130, pl. 14, figs. 2, 3
Kern Co., Calif.; Barker’s Ranch SU loc. 442
Middle Miocene, Temblor Fm_ [Wiedey’s specimen 436]
impolita, Diplodonta: Berry Holotype
Berry, 1953b, p. 409, pl. 28, figs. 3, 4
Forrester Island, Alaska; 15 fms
inequalis, Clementia: Wiedey Paratype
Wiedey, 1928, p. 146, pl. 18, fig. 5
Ventura Co., Calif.; Santa Paula Qd, SW 1/4 NW 1/4 Sec. 22, T 3 N,
R 21 W, from the abrupt terminus of South Mt.. along Santa Clara
River SU loc. 406
Lower Miocene, Vaqueros Fm [Wiedey’s specimen 426]
inezana, Plicatula: Durham Plastoholotype
Durham, 1950, p. 68, pl. 13, fig. 6
Baja California, Mexico; Santa Inez Bay UCMP loc. A3584
Pleistocene [holotype UCMP 15532]
inezana, Spondylus: Wiedey Paratype
Wiedey, 1928, p. 139
Ventura Co., Calif.; Calabasas sheet, head of Wiley Canyon [Piru
Qd]
Miccene, Vaqueros Fm
infelix, Hiata: Zetek and McLean Paratypes
Zetek and McLean, 1936, p. 110
Balboa, Canal Zone
inflata, Radiolites: Palmer Paratype
Palmer, 1928a, p. 83, pl. 17, fig. 4
Jalisco, Mexico; Huescalapa
Cretaceous, “Turonian”
inornatus, Grammatodon: (Meek and Hayden) Plastosyntype
Meek and Hayden, 1865, p. 90, pl. 3, figs. 9a, 9c
Wyoming; SW base of Black Hills
Jurassic, Sundance Fm [syntype USNM 201 = Arca (Cucullaea)
inornata Meek and Hayden, 1858, type species of Grammatodon Meek
and Hayden, 1860]
inornatus, Grammatodon: (Meek and Hayden) Plastosyntype
Meek and Hayden, 1865, p. 90, pl. 3, fig. 9b
Wyoming: SW base of Black Hills
Jurassic, Sundance Fm [syntype USNM 201 = Arca (Cucullaea)
inornata Meek and Hayden, 1858, type species of Grammatodon Meek
and Hayden, 1860]
insignis, Schizodus: Drake Holotype
Drake, 1898, p. 406, pl. 9, fig. 7
McDermitt, Okla.; 5 miles E of town
Permian
invalidus, Pecten: Hanna
Hanna, 1924, p. 177. [Renaming of Pecten (Plagioctenium) cooperi
Arnold, 1906]
San Diego, Calif.; Pacific Beach
Pliocene, San Diego Fm
346
619
5825
10056
9919
421
5218
7992
9301
9302
9303
5819
128
BuLvetin 300
ionense, Venericardia planicosta: Waring Holotype
Waring, 1914, p. 785. Illustrated in Waring, 1917, p. 96, pl. 11, fig. 1
Umpqua Valley, Oregon
Eocene, Umpqua Fm [= neotype of Venericardia (Leuroactis)
aragonia Arnold and Hannibal, 1914, designated by Stewart, 1930,
p. 170]
irisans, Anodontites: Marshall Paratypes
Marshall, 1926, p. 10
Venezuela
jamesi, Nuttallia: Roth and Guruswami-Naidu Paratype
Roth and Guruswami-Naidu, 1974, p. 143
Sonoma Co., Calif.; Sebastopol Qd, road cut N side of River Rd., .02
miles N of Trentcn CAS loc. 54164
Pliocene, Merced Fm
jenkinsi, Cardita: Smith Plastoholotype
Smith, 1927, p. 111, pl. 96, fig. 2
Shasta Co., Calif.; N fork Squaw Creek, 3 miles N of Kellys Ranch
Upper Triassic, Hosselkus Ls [holotype USNM 74160]
jordani, Pteria: Wiedey Paratype
Wiedey, 1928, p. 134, pl. 15, fig. 3 ;
Los Angeles Co., Calif.; Dry Canyon, 2 miles S§ of Calabasas
Middle Miocene, Temblor Fm
kamakawaensis, Tellina: Clark Holotype
Clark, 1925, p. 95, pl. 12; fig. 13
Skamokawa, Wash.; along Skamokawa River, above big bend, 1 mile
E of junction of main and middle forks SU loc. NP 272
Oligocene, Lincoln Fm, ss bluffs
keenae, Venericardia (Glyptoactis): Verastegui Holotype
Verastegui, 1953, p. 41, pl. 1, figs. 1-5
Fresno Co., Calif.; Panoche Qd, Sec. 29, T 15 S, R 12 E [Tumey Hills
Qd] opposite jct. of Panoche and Silver Creeks SU loc. 2073
Paleocene, Lodo Fm
keepi, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 60, pl. 5, fig. 1; pl. 6, figs. 1, la
San Diego Co., Calif.; Carrizo Creek area
Miocene
kelleyi, Pitar: Weaver and Kleinpell — Holotype
Weaver and Kleinpell, 1963, p. 204, pl. 35, fig. 11
Santa Barbara Co., Calif.; Goleta Qd, near Las Yegas Canyon
UCMP loc. B6983
Eocene, Gaviota Fm
kelleyi, Pitar: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 204, pl. 35, fig. 10
Santa Barbara Co., Calif.; Goleta Qd, near Las Yegas Canyon
UCMP loc. B6979
Eocene, Gaviota Fm
kelleyi, Pitar: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 204, pl. 36, fig. 1
Santa Barbara Co., Calif.; Goleta Qd, near Las Yegas Canyon
UCMP loc. B6933
Eocene, Gaviota Fm
kelseyi, Diplodon: Baker Paratype
Baker, 1914, p. 665
Rio Jamauchim, Brazil
kernensis, Pecten (Patinopecten): Hertlein Holotype
Hertlein, 1925b, p. 40, pl. 4, fig. 3
Kern Co., Calif.; Pyramid Hill, 3 miles NW of mouth of Kern River
Canyon SU loc. 150 [Rio Bravo Ranch Qd, T 28 S, R 29 E]
Miocene, Monterey Fm
5166
515
5159
5998
6311
5548
8426
8426a
9304
9305
9306
5412
5194
6938
STANFORD UNIveRsiry Tyres: SMITH 347
kernensis, Pholadomya: Wiedey Holotype
Wiedey, 1928, p. 141, pl. 17, figs. 1, 2
Kern Co., Calif.; N of Poso Creek, SW 1/4 SE 1/4 Sec. 12, T 27 S,
R 28 E SU loc. 438
Middle Miocene, Temblor Fm _ [Wiedey’s specimen 437]
kewi, Mytilus: Wiedey Holotype
Wiedey, 1929c, p. 281, pl. 31, fig. 2 [renamed Mytilus loeli by Grant,
1930, p. 419]
Monterey Co., Calif.; Los Vaqueros Valley SU loc. 100
Lower Miocene, Vaqueros Fm
kewi, Mytilus: Wiedey Paratype
Wiedey, 1929c, p. 281 [renamed Mytilus loeli by Grant, 1930, p. 419]
Monterey Co., Calif.; Los Vaqueros Valley
Lower Miocene, Vaqueros Fm
kewi, Tellina: Dickerson Plastoholotype
Dickerson, 1914, p. 138, pl. 12, fig. 1
Lake Co., Calif.; near Lower Lake UCMP loc. 784
Lower Eocene, Martinez Fm [holotype UCMP 11718]
kincaidi, Pecten hindsii: Oldroyd Holotype
Oldroyd, 1920, p. 135, pl. 4, figs. 3, 4
Puget Sound, Wash.
kiyonoi, Arca: Makiyama Paratype
Makiyama, 193la, p. 269, 273
Kyushu, Japan; Hakata Bay, in mud_ [cited as specimen VI]
landanensis, Venericardia: Vincent Plastosyntypes
Vincent, 1913, p. 29, pl. 3, figs. 5 (type 8426), 6 (type 8426a)
Portuguese West Africa; Falaise de Landana, Cabinda
“Paleocene” [syntypes RG 174, RG 110, at Mus. R. Congo Belge? ]
lascrucensis, Pitar: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 204, pl. 36, fig. 2
Santa Barbara Co., Calif.; Lompoc Qd, San Julian Ranch UCMP
loc. A 940
Eocene-Oligocene, upper Gaviota Fm
lascrucensis, Pitar: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 204, pl. 36, fig. 5
Santa Barbara Co., Calif.; Lompoc Qd, San Julian Ranch UCMP
loc. A 940
Eocene-Oligocene, upper Gaviota Fm
lascrucensis, Pitar: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 204, pl. 36, fig. 3
Santa Barbara Co., Calif.; Lompoc Qd, El Jaro at Yridisis Creek
loc. 2906b
Eocene, Middle Gaviota Fm
leana, Trigonia: Gabb Plastosyntype
Gabb, 1877, p. 312, pl. 31, fig. 362. Also in Gabb, 1864, p. 190, pl. 25,
fig. 178 (as T. gibboniana Lea?) [See Stewart, 1930, pp. 92-93]
Near Martinez, California (fig. 178) ; Jacksonville, Ore.
Cretaceous
lecontei, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 98, pl. 33, figs. 4, 4a, 4b
Off Baja California, Mexico; Cedros Island
Pliocene, Salada Fm?
lewisi, Pinna: Waring Holotype
Waring, 1917, p. 94, pl. 15, fig. 24
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
limata, Leda hamata: Dall Paratype
Dall, 1916, p. 397
Off Santa Rosa Island, California; 53 fms
8023
8741
519
515
5425
407
9711
167
9906
5167
5583
7258
8021
BULLETIN 300
lisa, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 39, pl. 21, figs. 1, 2
Lewis Co., Wash.; bluffs along Olequa Creek at Old Ainslee Mill,
ieN, Re2 Ww:
Upper Eocene, Cowlitz Fm
lobatus, Inoceramus: Munster in Goldfuss Plastoholotype
Goldfuss, 1836, p. 113, pl. 110, fig. 3
Westphalia, Germany
Cretaceous [holotype 673 BM (NH) ]
loeli, Amiantis (?): Wiedey Holotype
Wiedey, 1929c, p. 288, pl. 32, fig. 2; pl. 33, fig. 3
San Mateo Co., Calif.; Searsville Road roadcut near Stanford Uni-
versity SU loc. 450
Middle Miocene, Monterey Fm
loeli, Mytilus: Grant Holotype
Grant, 1930, p. 419. [mew name for Mytilus kewi Wiedey, 1929c]
Monterey Co., Calif.; Los Vaqueros Valley SU loc. 100
Lower Miocene, Vaqueros Fm
lorenzanum, Cardium cooperi: Arnold Paratype
Arnold, 1908a, p. 366
Santa Cruz Co., Calif.; E branch, N Fork, Waddell Creek, Big Basin
Oligocene, San Lorenzo Fm
lucianus, Inoceramus: Davis Holotype
Davis, 1913, p. 455, fig. 2
Monterey Co., Calif.; 4 miles N of Slate’s Springs
Jurassic, Franciscan Fm
lunaris, Pecten: Berry Holotype
Berry, 1963, p. 139. Illustrated in Keen, 1971, p. 85, fig. 176
Sonora, Mexico; off Morro Colorado, 30-45 fms
maccrayi, Glycimeris [sic]): Waring Holotype
Waring, 1917, p. 93, pl. 15, fig. 1
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
madisonensis, Posidonia: Smith Plastoholotype
Smith, 1927, p. 112, pl. 94, fig. 12
Shasta Co., Calif.; NW end Brock Mt. between Squaw Creek and Pit
River
Upper Triassic, Hosselkus Ls_ [holotype USNM 74144]
margaritana, Dosinia: Wiedey Paratype
Wiedey, 1928, p. 145, pl. 18, fig. 2
San Luis Obispo Co., Calif.; 4 miles E of La Panza, S side of low
ridge forming N wall of canyon through which McKittrick-La Panza
road passes SU loc. 436
Lower Miocene, Vaqueros Fm _ [Wiedey’s specimen 425]
maria, Chaenomya: Worthen Holotype
Worthen, 1882, pv. 39. Illustrated in Worthen, 1883, p. 319, figs. 1a, 1b
Shawnee Co., Kansas; Plowboy
Pennsylvanian, Upper Coal Measures
markleyensis, Mactra (Spisula): Clark Plastoholotype
Clark, 1938, p. 699, pl. 2, fig. 6
Solano Co., Calif.; Napa Qd, S of Putah Creek UCMP loc. A 1297
Eocene, Markley Fm [holotype UCMP 30852]
marksi, Venericardia (Glyptoactis): Verastegui Holotype
Verastegui, 1953, p. 44, pl. 19, figs. 2-4
Kern Co., Calif.; E side Live Oak Canyon SU loc. 183
Upper Eocene, Tejon Fm
7303
9915
8011
450
7526
6560
6941
7787
433
8586
7850
398
39
8029
STANFORD UNIVERSITY I yPEs: SMITH 349
martinezensis, Teliina: Weaver Plastoholotype
Weaver, 1905, p. 115, pl. 12, fig. 3
Contra Costa Co., Calif.; S of Martinez UCMP loc. 337
“Eocene,” Martinez Fm [Paleocene] [holotype UCMP 11816]
martini, Lima: Smith Plastoholotype
Smith, 1927, p. 122, pl. 101, fig. 11
Alaska; S bank of Yukon River opposite Nation River
Upper Triassic [holotype USNM 74200]
mcmastersi, Venericardia (Glyptoactis): Verastegui Holotype
Verastegui, 1953, p. 42, pl. 13, figs. 2, 3
San Diego Co., Calif.; San Clemente Canyon
Middle Eocene, La Jolla Fm
meeki, Corneocyclas: Hannibal Holotype & Paratypes
Hannibal, 1912b, p. 135, pl. 6, fig. 12. Also in Taylor and Smith, 1971,
fig. 2 (as Sphaerium)
Near Hawthorne, Nevada; hill on Belmont Stage road
Eocene [Miocene, Esmeralda Fm, fide Taylor and Smith, 1971, p.
310]
menuda, Lucinisca: Keen Holotype
Keen, 1943, p. 40, pl. 3, figs. 15, 16
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4 Sec.
6,T 29S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
meridionalis, Chione: Oldroyd Paratype
Oldroyd, 1921, p. 93, pl. 4, fig. 4
Peru
meridionalis, Miodontiscus: Dail Paratype
Dall, 1916, p. 408
San Diego Co., Calif.; off Point Loma 67-78 fms
meropsis, Tellina (Moerella): Dall Paratypes
Dall, 1900b, p. 317
San Diego, Calif.
merriami, Pecten (Pecten): Arnold Neotype
Arnold, 1906, p. 99. Holotype presumed destroyed in San Francisco
fire, 1906, in California State Mining Bureau collections. Neotype
selected by Grant and Gale, 1931, p. 195
Ventura Co., Calif.; San Felician Creek, near Piru
Pliocene
mexicanum, Galeomma (Lepirodes?): Berry Holotype
Berry, 1959, p. 108. Illustrated in Keen, 1971, p. 135, fig. 308 (as
Tryphomyax)
Gulf of California, San Luis Gonzaga Bay, 3-4 fms
microsperma, Nucula (Ennucula): Berry Holotype
Berry, 1947b, p. 258, pl. 26, fig. 2
San Pedro, Calif.; near Second and Pacific Streets
Pleistocene, Lomita Fm
milthoidea, Dosinia: Waring Holotype
Waring, 1917, p. 60, pl. 8, fig. 5
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon, N. of Simi fault
Upper Cretaceous, Chico Fm
modulatus, Pecten (Lyropecten): Hertlein Holotype
Hertlein, 1925a, p. 11, pl. 3, fig. 6
Baja California, Mexico; Scammon Lagoon Qd, mesa W of Mesa de
las Auras SU loc. 43
Pliocene, Salada Fm
montereyensis, Cardita (Cyclocardia) ventricosa:
Smith and Gordon Paratypes
Smith and Gordon, 1948, p. 213
Off Monterey, Calif., 70 fms
350
10319a
10319b
10319¢
166
516
9308
9309
7994
971
5100
7852
7851
9499
10189
8059
5914
BuLueTIN 300
moorei, Cyprimeria: Popenoe Plastosyntypes
Popenoe, 1937, p. 391, pl. 48, fig. 1 (type 10319), fig. 2 (type 10319a)
Santa Ana Mts., Calif.; CIT loc. 92
Cretaceous, Turonian [syntypes UCLA 40648, 40649, 40650]
morani, Cucullaea: Waring Holotype
Waring, 1914, p. 784. Ilustrated in Waring, 1917, pl. 14, figs. 12, 13
Ventura Co., Calif.; 1.5 miles E of McCray Wells
Eocene, Tejon Fm
morani, Dosinia: Wiedey Holotype
Wiedey, 1929c, p. 281, pl. 31, fig. 3
San Luis Obispo Co., Calif.; Canyon de Piedra, 4 miles E of San
Luis Obispo
Lower Miocene, Vaqueros Fm
mulinoidus, Pitar: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 205, pl. 36, fig. 7
Santa Barbara Co., Calif.; El Jaro at Yridisis Creek loc. 2907
Eocene, middle Gaviota Fm
mulinoidus, Pitar: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 205, pl. 36, figs. 6, 10 :
Santa Barbara Co., Calif.; El Jaro at Yridisis Creek loc. 2907
Eocene, middle Gaviota Fm
mulleri, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 20, pl. 1, figs. 6-9
Fresno Co., Calif.; Panoche Qd [Tumey Hills Qd], Sec. 29, T 15 S,
R12 E_ SU loc. 2073
Paleocene, Lodo Fm
muitirugosus, Pecten (Chlamys): Gale Paratype
Gale, 1928, p. 92. Illustrated zx Grant and Gale, 1931, p. 159, pl. 11,
figs. 5a, 5b
San Pedro, Calif.
multitubifera, Caprinuloidea: Palmer Paratype
Palmer, 1928a, p. 61
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
myrae, Ensis: Berry Holotype
Berry, 1953a, p. 398, pl. 29, figs. 5, 6, text fig. 4
San Pedro Bay, Calif.; near Terminal Island
myrae, Ensis: Berry Paratype
Berry, 1953a, p. 398, text fig. 3
San Pedro Bay, Calif.; near Terminal Island
myrae, Periploma (Halistrepta): Rogers Holotype
Rogers, 1962, p. 235, figs. 1, 2. Illustrated in Keen, 1971, p. 295, fig.
257
Gulf of California; off Loreto, near Carmen Island, 15-25 fms
Mytilus, n. sp. aff. M. tichanovitchi Makiyama: Addicott
Holotype
Addicott, 1976, p. 101, pl. 1, fig. 6
Clallam Co., Wash.; Clallam Bay, seacliffs eastward from Slip Point
for 1/2 mile. SU loc. NP 89
Lower Miocene, Clallam Fm, Pillarian stage
nakamurai, Katelysia (Nipponomarcia): Ikebe Paratypes
Ikebe, 1941, p. 50
Shiga Prefecture, Japan; Sendani, Yamanouchi-mura, Koga-gun
Middle Miocene, Ayugawa group
nana, Cuspidaria: Oldroyd Holotype
Oldroyd, 1918b, p. 28. Illustrated in Oldroyd, 1925, p. 99, pl. 13, figs.
8,9
Monterey, Calif.
9909
118
5203
431
66
87
88
72
73
STANFORD UNIVERSITY Types: SMITH 55h
nana, Myoconcha: Smith Plastoholotype
Smith, 1927, p. 111, pl. 94, figs. 10, 11
Shasta Co., Calif.; old quarry SW end Brock Mt. between Squaw
Creek and Pit River
Upper Triassic, Hosselkus Ls [holotype USNM 74141]
nanaimensis, Pholadomya: Reagan Holotype
Reagan, 1924, p. 185, pl. 20, fig. 7
Vancouver Island, British Columbia, Canada; near Nanaimo
Upper Cretaceous
nelsoni, Nucula (Acila): Clark Holotype
Clark, 1925, p. 74, pl. 8, fig. 1
Wash.; 3/4 mile W of Gettysburg in shaly ss sea cliffs at mouth of
Duncan Creek SU loc. NP 90
Oligocene, Blakeley Fm
nevadanus, Pecten Conrad: Grant and Gale Neotype
Conrad, 1856, p. 329, pl. 8, fig. 7. Neotype designated by Grant and
Gale, 1931, p. 189, pl. 7, figs. 2a-2c, as type of Vertipecten Grant and
Gale. [Specimen is P¢cten bowersi Arnold, not Pecten nevadanus Con-
rad |
McKittrick district? Santa Monica Mts. ?
Middle Miocene
newcombei, Mulinia: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 153, pl. 16, figs. la, 1b
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NE 129
Oligocene? Sooke Fm
newcombei, Mulinia: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 153, pl. 15, fig. 2
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene? Sooke Fm
newcombei, Mulinia: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 153, pl. 15, fig. 3
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene? Sooke Fm
newcombei, Mulinia: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 153, pl. 15, figs. 4a, 4b
Vancouver Island, British Columbia, Canada; Socke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene? Sooke Fm
newcombei, Pododesmus: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 141, pl. 21, fig. 4
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, sea cliffs at mouth of Fossil Creek SU loc.
NP 130
Oligocene? Sooke Fm
newcombei, Pododesmus: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 141, pl. 21, fig. 6
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, sea cliffs at mouth of Fossil Creek SU loc.
NP 130
Oligocene? Sooke Fm
352
92
8054
6049
6051
5950
6570
6570a
5220
5346
6533
6508
5913
121
BULLETIN 300
newcombei, Pododesmus: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 141, pl. 21, fig. 3
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, sea cliffs at mouth of Fossil Creek SU loc.
NP 130
Oligocene? Sooke Fm
nicoyana, Tellina (Scissula): Hertlein and Strong Paratype
Hertlein and Strong, 1949a, p. 85
Gulf of Nicoya, Costa Rica; off Ballenas Bay, 35 fms
nipponica, Lasaea: Keen Holotype
Keen, 1938, pp. 26-27, figs. 1a, 1b
NE Matsusima, Japan; Watanoha, Rikuzen
nipponica, Lasaea: Keen Paratype
Keen, 1938, pp. 26-27
NE Matsusima, Japan; Watanoha, Rikuzen
nodosus, Vermetus: Oldroyd Holotype
Oldroyd, T. S., 1921a, p. 116, pl. 5, fig. 10
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm [=a burrow lining of a Holocene
teredid pelecypod, teste Keen, 1976]
nuculiformis, Crassinella: Berry Holotype
Berry, 1940b, p. 149, pl. 17, fig. 1
San Pedro, Calif.; W side of Gaffey Street cut
Pleistocene
nuculiformis, Crassinella: Berry Paratype
Berry, 1940b, p. 149
San Pedro, Calif.; W side of Gaffey Street cut
Pleistocene
nutteri, Pecten (Chlamys): Arnold Holotype
Arnold, 1906, p. 67, pl. 11, fig. 3
San Mateo Co., Calif.; S of mouth of San Gregorio Creek
Pliocene, Purisima Fm
oakvillensis, Lima (Radula): Clark Holotype
Clark, 1925, p. 84, pl. 15, fig. 1
Wash.; 1 mile W of Oakville, in lower tuffaceous conglomerate beds
immediately overlying basalt at quarry on N.P. R.R. SU loc. NP 109
Oligocene, Lincoln Fm
obliqua, Protarca: Stephenson Plastoholotype
Stephenson, 1923, p. 104, pl. 19, fig. 3
Greene Co., N.C.; Snow Hill
Cretaceous, Black Creek Fm, Snow Hill Mbr
okawensis, Nuculopsis (Palaeonucula?): Schenck Paratype
Schenck, 1939, p. 23
Illinois; 1.4 miles NE of Ruma
Mississippian, lower Okaw Ls
oldroydi, Corbicula: Clark Paratype
Clark, 1938, p. 698
Napa Qd, Calif.; Brink Ranch, 2 miles S of Putah Creek
Upper Eocene, Markley Fm
oldroydii, Atrina: Dall Holotype
Dall, 1901a, p. 143. Illustrated in Dall, 1921, pl. 2, figs. 4-6
Los Angeles Co., Calif.; San Pedro Bay, 25 fms
oldroydii, Avicula: Reagan Holotype
Reagan, 1924, p. 186, pl. 20, fig. 1
Vancouver Island, British Columbia, Canada; near Nanaimo SU loc.
ila b7/
Upper Cretaceous
7978
5251
7773
5991
6590
6052
9888
29
201
191
8009
9890
5468
Cn
W
STANFORD UNIVERSITY Types: SMITH 3
olssoni, Megapitaria: Marks Paratype
Marks, 1951, p. 79
SW Ecuador; NE of Progreso
Middle Miocene, Progreso Fm
olympiana, Yoldia: Clark Holotype
Clark, 1925, ps 77, pla 9. fig. 9
Twin, Wash.; sea cliffs W of Twin River SU loc. NP 120
Oligocene, Blakeley Fm
onestae, Integricardium (Onestia): McLearn Plastoholotype
McLearn, 1933b, p. 152, pl. 2, fig. 10
Alberta, Canada; E bank Athabasca River, 3 miles below Brule rapids
Cretaceous, Clearwater Fm [holotype at Natl. Mus. Canada]
ooides, Tellina: Gabb Plastoholotype or Plastosyntype
Gabb, 1864, p. 157, pl. 22, fig. 135a
Butte Co.. Calif.; Pence’s Ranch
Cretaceous [holotype UCMP 31437]
operculiformis, Pecten: Gabb Plastoholotype
Gabb, 1864, p. 201, pl. 26, fig. 188. Cited as lectotype by Stewart,
1930, p. 120
Shasta Co., Calif.; possibly from Huling Creek
Cretaceous [holotype UCMP 31446]
oregonensis, Crassinella: Keen Holotype
Keen, 1938, p. 31, pl. 2, figs. 11, 12
Coos Bay, Ore.; South Slough at highway bridge, 1-2 fms
oregonensis, Halobia: Smith Plastoholotype
Smith, 1927, p. 117, pl. 95, fig. 1
Baker Co., Ore.; Martins Bridge
Upper Triassic, upper Karnic, Eagle River Fm [holotype USNM]
oregonensis, Pecten: Howe Holotype
Howe, 1922, p. 98, pl. 11, fig. 1
Coos Bay, Ore.; SU loc. NP 44
Pliocene, Empire Fm
oregonensis, Pecten: Howe Paratype
Howe, 1922, p. 98, pl. 12, fig. 2
Coos Bay, Ore.; SU loc. NP 44
Pliocene, Empire Fm
oregonensis, Pecten: Howe Paratype
Howe, 1922, p. 98, pl. 11, fig. 2 _
Grays Harbor Co., Wash.; N of mouth of Raft River, Taholah SU
loc. NP 82
Pliocene, Quillayute Fm
oregonensis, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 25, pl. 9, figs. 7-9
Douglas Co., Ore.; Roseburg Qd, Little River bluffs at jct. with North
Umpqua River near Glide
Lower Eocene, Umpqua Fm
ornatissima, Halobia: Smith Plastoholotype
Smith, 1927, p. 117, pl. 94, fig. 4
Shasta Co., Calif.; W side Brock Mt. between Squaw Creek and Pit
River
Upper Triassic, Hosselkus Ls, upper horizon, Juvavites subzone of
Tropites subbullatus zone
ovalis, Posidonia: Kittl Plastoholotype
Katt 1912) 5ps-29; ple ties 15
Pelponnes or Dalmatia; Kurkuli
Middle Jurassic, Humphresianum zone [holotype in Naturh. Staats-
mus. Wien]
9913
6937
513
7293
6944
150
8335
7793
6518
6939
8060
5820
6945
42
BuLLeETIN 300
overbecki, Pleurophorus: Smith Plastoholotype
Smith, 1927, p. 111, pl. 101, fig. 15
Alaska; S bank of Yukon River opposite Nation River
Upper Triassic [holotype USNM 74203]
pacifica, Malletia: Dall Paratype
Dall, 1897a, p. 11
Off Pt. Conception, Calif.; 278 fms, USBF Sta. 3198
pacifica, Mesodesma: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 79. Illustrated in Clark, 1922, p. 118, pl.
13, fig. 5 (as Myadesma)
Alameda Co., Calif.; Pleasanton Qd, Alameda Creek, 1.5 miles S of
Welch Creek, 1/5 mile § of Calaveras fault
Miocene, Monterey Fm
packardi, Tellina: Dickerson Plastoholotype
Dickerson, 1914, p. 137, pl. 11, fig. 11
Lake Co., Calif.; Lower Lake UCMP loc. 784
Eocene, Martinez Fm [holotype UCMP 11739]
panamensis, Protocardia: Dall Paratype
Dall, 1908b, p. 415 .
Panama Bay, 182 fms
parsonsi, Miltha: Waring Holotype
Waring, 1917, p. 78, pl. 12, fig. 13
Ventura Co., Calif.; Martinez area in the Simi Hills
Lower Eocene, Martinez Fm
peabodyi, Chione (Chione) californiensis: Parker Plastoholotype
Parker, 1949, p. 581, pl. 90, fig. 1
Ventura Co., Calif.; N of Springville
Pleistocene [holotype UCMP]
pectunculoides, Peruarca: Olsson Plastoholotype
Olsson, 1944, p. 33, pl. 3, figs. 6, 7
Paita region, Peru; near La Tortuga
Cretaceous, Maestrichtian, Radiolite Ss Baculites zone [holotype PRI
No. 4817]
pembertoni, Inoceramus: Waring Holotype
Waring, 1917, p. 61, pl. 7, figs. 7, 8
Los Angeles Co., Calif.; S of Santa Monica Mts.
Upper Cretaceous, Chico Fm
penderi, Leda: Dall and Bartsch Paratype
Dall and Bartsch, 1910, p. 9
Vancouver Island, British Columbia, Canada; Barkley Sound
pentodon, Limopsis: Aguayo and Borro Paratype
Aguayo and Borro, 1946b, p. 48
Matanzas, Cuba; Barranco E of Rio Canimar
Upper Miocene, Yumuri Fm
peraltum, Pisidium: Sterki Paratypes
Sterki, 1900, p. 5
Benzie Co., Mich.; Crystal Lake
perambilis, Cardium (Fulvia): Dall Paratype
Dall, 1881, p. 132
Off Barbados, 100 fms
percarus, Pecten (Aequipecten): Hertlein Holotype
Hertlein, 1925a, p. 13, pl. 2, figs. 2, 5
Baja California, Mexico; Scammon Lagoon Qd, mouth of large arroyo
NW of Elephant Mesa SU loc. 48
Pliocene, Salada Fm
8582
5163
552
7571
5529
5098
7564
7569
551
8732
303
502
13
STANFORD UNIVERSITY TyYPEs: SMITH 355
percarus, Pecten (Aequipecten): Hertlein Paratypes
Hertlein, 1925a, p. 13
Baja California, Mexico; Scammon Lagoon Qd, mouth of large arroyo
NW of Elephant Mesa _ SU loc. 48
Pliocene, Salada Fm
percarus, Pecten (Aequipecten): Hertlein Paratype
Hertlein, 1925a, p. 13
Baja California, Mexico; Turtle Bay CAS loc. 930
Pliocene, Salada Fm
percrassa, Nucula: Conrad Plastoholotype
Conrad, 1858, p. 327, pl. 35, fig. 4
Mississippi; Owl Creek, 3 miles N of Ripley
Upper Cretaceous, Ripley Fm [types at ANSP No. 16710]
perdisparis, Arca: Wiedey Paratype
Wiedey, 1928, p. 131, pl. 14, fig. 1. Also iz Reinhart, 1943, p. 72, pl. 10,
fig. 8
Monterey Co., Calif.; 3/4 mile SW of Zayante Station, Santa Cruz
Mts. SU loc. 443
Middle Miocene, Monterey Fm _ [Wiedey’s specimen 433]
perfecta, Caprinuloidea: Palmer Paratypes
Palmer, 1928a, p. 59
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
perforata, Radiolites: Palmer Paratype
Palmer, 1928a, p. 81, pl. 16, fig. 11
Jalisco, Mexico; Huescalapa
Cretaceous, Turonian
perforata, Radiolites: Palmer Paratype
Palmer, 1928a, p. 81, pl. 14, figs. 6, 7
Jalisco, Mexico; Huescalapa
Cretaceous, Turonian
perforata, Radiolites: Palmer Paratypes
Palmer, 1928a, p. 81, pl. 16, fig. 9 (type 7569)
Jalisco, Mexico; Huescalapa
Cretaceous, Turonian
pernoides, Inoceramus: Goldfuss Plastoholotype
Goldfuss, 1826, p. 109, pl. 109, fig. 3
Westphalia, Germany
Cretaceous [cast of Goldfuss specimen 665 BM(NH) ]
perrini, Lima: Waring Holotype
Waring, 1914, p. 782. Illustrated zz Waring, 1917, p. 76, pl. 10, figs.
i 2
Ventura Co., Calif.; Simi Hills, Martinez area, Calabasas sheet
Lower Eocene, Martinez Fm
perrini, Ostrea titan: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 80. Illustrated in Wiedey, 1929b, pl. 3,
fig. 1
Alameda Co., Calif.; Pleasanton Qd
Middle Miocene, Briones Fm
perrini, Pecten (Lyropecten): Arnold Holotype
Arnold, 1906, p. 80, pl. 14, figs. 1, 1a; pl. 15, fig. 1
San Luis Obispo Co., Calif.; [Cayucos Qd] between Morro and Toro
Creeks
Miocene, Vaqueros Fm
perrini, Spondylus: Wiedey Paratype
Wiedey, 1928, p. 138
Ventura Co., Calif.; Calabasas sheet, Wiley Canyon [Piru Qd]
Miocene, Vaqueros Fm
7292
8697
5485
6138
9515
921
9904
5202
5239
977
8066
8287
8288
8289
BuL_eTIN 300
perrini, Tellina: Dickerson Plastoholotype
Dickerson, 1914, p. 137, pl. 11, fig. 8
Lake Co., Calif.; Lower Lake UCMP loc. 784
Eocene, Martinez Fm_ [holotype UCMP 11716]
perrinsmithi, Trigonia: Anderson Holotype
Anderson, 1958, p. 110, pl. 2, fig. 7
Shasta Co., Calif.; Horsetown
Upper Cretaceous, Horsetown Fm
peruanus, Pecten: Tilmann Plastoholotype
Tilmann, 1917, pp. 673-674, pl. 24, fig. 5
Peru; Chilingote, E] Tingo, Utcubamba-Tal
Lower Jurassic, Arietenzone, Psiloceras beds [holotype probably at
der Sammlung des Geologisch-palaontologischen Instituts der Univer-
sitat Bonn]
phenax, Musculus: Dall Paratype
Dall, 1915a, p. 138. Illustrated iz Schenck, 1945, p. 519, pl. 67, figs.
27-30
Bering Sea; Pribiloff Islands, St. George
phoebe, Pegmapex: Berry Holotype
Berry, 1960, p. 115. Illustrated 77 Keen, 1971, p. 131, fig. 300
Sinaloa, Mexico; Las Gaviotas Beach, Mazatlan
piedraensis, Platyodon: Wiedey Holotype
Wiedey, 1929c, p. 289, pl. 33, fig. 2
San Luis Obispo Co., Calif.; head of Canyon de Piedra, ca. 5 miles E
of San Luis Obispo SU loc. 441
Lower Miocene, Vaqueros Fm
pittensis, Pecten (Entolium): Smith Plastoholotype
Smith, 1927, p. 121, pl. 7, fig: 5
Shasta Co., Calif.; Brock Mt
Upper Triassic, Hosselkus Ls [holotype USNM 73947]
pittsburgensis, Spisula: Clark Holotype
Clark, 1925, p. 101, pl. 17, figs. 2, 4
Ore.; bluffs along Nehalem River near old Pittsburg mill below
Vernonia
Oligocene, Pittsburg Bluff Fm [specimen published as LSJU No. 53]
pittsburgensis, Tellina: Clark Holotype
Glarky9255 p. 9S pls da tigs
Ore.; ss bluffs along Nehalem River near old Pittsburg mill below
Vernonia SU loc. NP 5
Oligocene, Pittsburg Bluff Fm
planiuscula, Macoma: Grant and Gale Holotype
Grant and Gale, 1931, p. 372, pl. 14, figs. 11a, 11b; pl. 20, figs. 8a, 8b
Bering Sea, off Alaska; Nunivak Island
planiuscula, Macoma: Grant and Gale Paratype
Grant and Gale, 1931, p. 372
Bering Sea, off Alaska; Nunivak Island
pomeyroli, Granocardium (Ethmocardium): Keen Holotype
Keen, 1954, p. 314, pl. 29, fig. 4
New Caledonia, area of Momea tribe
Upper Cretaceous
pomeyroli, Granocardium (Ethmecardium): Keen Paratype
Keen, 1954, p. 314, pl. 29, fig. 3
New Caledonia; area of Momea tribe
Upper Cretaceous
pomeyroli, Granocardium (Ethmocardium): Keen Paratype
Keen, 1954, p. 314, pl. 29, fig. 2
New Caledonia; area of Momea tribe
Upper Cretaceous
8290
8291
8292
8293
8294
10336
5222
7808
8298
7262
38
89
6960
6961
6962
6963
STANFORD UNIVERSITY TyprEs: SMITH 357
pomeyroli, Granocardium (Ethmocardium): Keen Paratype
Keen, 1954, p. 314, text figs. 1, 2
New Caledonia; area of Momea tribe
Upper Cretaceous
pomeyroli, Granocardium (Ethmocardium): Keen Paratypes
Keen, 1954, p. 314
New Caledonia; area of Momea tribe
Upper Cretaceous
popenoei, Cymbophora: Saul Paratypes
Saul, 1974, p. 1087
Santa Ana Mts., Calif.; Corona sheet, SW slope of ridge between
Aliso and Santiago Creek, 1650’ N 38° E of Pankratz Ranch house,
4800’ S 18° W of dam 1/4 mile above mouth of Harding Canyon
CIT loc. 974
Cretaceous, late Campanian, Williams Fm, Pleasants Ss Mbr
porterensis, Modiolus: Clark Holotype
Clark, 1925, p. 85, pl. 9, fig. 11
Wash.; marly tuffs at old log dam on Porter Creek, 1.5 miles above
Porter SU loc. NP 51
Oligocene, Lincoln Fm
portusregii, Pecten (Plagioctenium) gibbus: Grau Paratype
Grau, 1952a, p. 17. Grau, 1952b, p. 69 (mew name for P.g. carolinensts,
preoccupied)
Off South Carolina; 2 miles off Port Royal, 80’
praeblandum, Clinocardium: Keen Plastoholotype
Keen, 1954, p. 321, pl. 29, fig. 6
Contra Costa Co., Calif.; W end of Las Trampas Ridge
Upper Miocene, Briones Fm _ [holotype UCMP 14836]
praecuta, Tellina: Clark Plastoholotype
Clark e1908'tpr 1535 plei2) trons
Contra Costa Co., Calif.; Sobrante Ridge UCMP loc. 14
Oligocene, San Ramon Fm_ [holotype UCMP 11166]
pretiosus, Pecten (Lyropecten): Hertlein Holotype
Hertlein, 1925a, p. 12, pl. 3, fig. 4
Baja California, Mexico; Turritella bed above San Gregorio Lagoon,
on the trail from Arroyo Mesquital to La Purisima SU loc. 59
Miocene, Isidro Fm
pretiosus, Pecten (Lyropecten): Hertlein Paratype
Hertlein, 1925a, p. 12, pl. 2, fig. 6
Baja California, Mexico; La Purisima cliffs on San Ramon River
SU loc. 57
Miocene, Isidro Fm
princeps, Acila (Truncacila): Schenck Holotype
Schenck, 1943, p. 63, pl. 8, figs. 4, 6, 7, 8
Merced Co., Calif.; Sec. 12, T 12 S,R10E SU loc. 2372
Upper Cretaceous, Moreno Fm
princeps, Acila (Truncacila): Schenck Paratype
Schenck, 1943, p. 63, pl. 8, fig. 2
Merced Co., Calif.; Sec. 12, T 12 S,R10E SU loc. 2372
Upper Cretaceous, Moreno Fm
princeps, Acila (Truncacila): Schenck Paratype
Schenck, 1943, p. 63, pl. 8, figs. 1, 3
Merced Co., Calif.; Sec. 12,T 12 S,R10E SU loc. 2372
Upper Cretaceous, Moreno Fm
princeps, Acila (Truncacila): Schenck Paratype
Schenck, 1943, p. 63
Merced Co., Calif.; Sec. 12, T 12 S,R10E SU loc. 2372
Upper Cretaceous, Moreno Fm
358
8301
420
7972
8086
7379
6235
5114
6312
6231
6924
7872
7871
BuLLETIN 300
pristinum, Clinocardium: Keen Plastoholotype
Keen, 1954, p. 322, pl. 29, fig. 15
Contra Costa Co., Calif.; Concord Qd, Shell Ridge
Upper Miocene, Neroly Fm? [holotype UCMP]
procumbens, Arca: Wiedey Holotype
Wiedey, 1928, p. 132, pl. 13, fig. 11. Also zz Reinhart, 1943, p. 54, pl.
5, fig. 2 (as Anadara)
Lincoln Co., Ore.; 5 miles N of Yaquina Head SU loc. 444
Miocene
progresoensis, Pecten (Aequipecten): Marks Paratypes
Marks, 1951, p. 60
SW Ecuador; about 6 miles NE of Progreso
Middle Miocene, Progreso Fm
prosperi, Glibertia: Van der Meulen Paratype
Van der Meulen, 1951, pp. 49, 53
The Netherlands; beach sand near Rittham, Zeeland Province
Pliocene, reworked [cited as paratype II]
pseudoillota, Barbatia (Fugleria): Reinhart Plastoholotype
Reinhart, 1937b, p. 184, pl. 28, figs. 6, 9, 10 -
Santa Barbara Co., Calif.; Fugler Point
Pliocene [holctype CIT 1383 = now LACMNH 4075]
pugetensis, Lyonsia: Dall Paratype
Dall, 1913, p. 595
Wash.; coast N of Queets River
pugetensis, Nucula (Acila): Clark Holotype
Clark, 1925, p. 75, pl. 8, fig. 4
Bainbridge Island, Wash.; Bean Point SU loc. NP 205
Oligocene, Blakeley Fm [Clark’s specimen No. 5]
pugetensis, Pecten islandicus: Oldroyd Holotype
Oldroyd, I. S., 1920, p. 136, pl. 4, figs. 5, 6. Also zz Oldroyd, I. S.,
1925, p. 55, pl. 12, figs. 4, 5
Puget Sound, Wash.; off San Juan Island
puntarenensis, Mytilus (Hormomya): Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932a, p. 104, Illustrated iz Keen, 1971, p. 61, fig.
121, lower left (as Brachidontes)
Puntarenas, Costa Rica
purisimaensis, Pecten (Patinopecten): Arnold Holotype
Arnold, 1906, p. 105, pl. 34, fig. 3
San Mateo Co., Calif.; N of mouth of Pescadero Creek
Pliocene, Purisima Fm
pygmaeus, Musculus: Glynn Holotype
Glynn, 1964, pp. 121-128, pl. 23, figs. 1a, 1b
Pacific Grove, Calif.; near Hopkins Marine Station
pygmaeus, Musculus: Glynn Paratype
Glynn, 1964, pp. 121-128
Pacific Grove, Calif.; near Hopkins Marine Station
quadrata, Palaeocardita: Trechmann Paratypes
Trechmann, 1918, p. 212
New Zealand; Nugget Point, Otago
Triassic, Carnic
redondoensis, Aligena: Burch Paratypes
Bureh, 2, 1941; sp. 50
Los Angeles Co., Calif.; off Redondo Beach, 75 fms
redondoensis, Cardita: Burch Paratype
Burch, J. Q., 1945, p. 32
Los Angeles Co., Calif.; off Redondo Beach, 100 fms, mud bottom
redondoensis, Nuculana penderi: Burch Paratypes
Burch, J. Q., 1945, p. 10
Los Angeles Co., Calif.; off Redondo Beach, 25 fms, gravel bottom
49
50
10328
10329
7306
6232
5143
5144
34
5483
557
7563
7563a
STANFORD UNIVERSITY Types: SMITH 359
refugioensis, Pecten (Pecten): Hertlein Holotype
Hertlein, 1925a, p. 7, pl. 1, fig. 2
Baja California, Mexico; Rancho Refugio, N of San Jose del Cabo
SU loc. 50
Upper Miocene or lower Pliocene
refugioensis, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925a, p. 7, pl. 5, fig. 9
Baja California, Mexico; Rancho Refugio, N of San Jose del Cabo
SU loc. 50
Upper Miocene or lower Pliocene
refugioensis, Pecten (Pecten): Hertlein Paratype
Hertlein, 1925a, p. 7
Baja California, Mexico; Arroyo Fortuna, N of San Jose del Cabo
SU loc. 44
Upper Miocene or lower Pliocene
regina, Calva: Popenoe Plastosyntype
Popenoe, 1937, p. 395, pl. 48, figs. 6, 13
Santa Ana Mts., Calif.; CIT loc. 1164
Cretaceous, Turonian, Ladd Fm, Baker Mbr_ [syntype UCLA 40660]
regina, Calva: Popenoe Plastosyntype
Popenoe, 1937, p. 395, pl. 48, figs. 7, 14
Santa Ana Mts., Calif.; CIT loc. 1164
Cretaceous, Turonian, Ladd Fm, Baker Mbr_ [syntype UCLA 40661]
remondii, Tellina: Gabb Plastoholotype
Gabb, 1864, p. 156, pl. 22, fig. 132
Contra Costa Co., Calif.; Cochran’s, E of Mt. Diablo UCMP loc. 138
“Cretaceous,” [Eocene, Meganos Fm] [holotype UCMP 314511]
rhypis, Pandora (Kennerlia): Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932a, p. 105
Gulf of Fonseca, El Salvador; La Unicn
richthofeni, Chione: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 619, pl. 17, figs. 7, 8
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
richthofeni, Chione: Hertlein and Jordan Paratype
Hertlein and Jordan, 1927, p. 619, pl. 17, fig. 4
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
riversi, Pecten (Propeamusium): Arnold Holotype
Arnold, 1906, p. 126, pl. 44, fig. 8
Los Angeles Co., Calif.; Santa Monica Canyon
“Pliocene”
riversi, Pecten (Propeamusium): Arnold Paratype
Arnold, 1906, p. 126, pl. 44, fig. 9
Los Angeles Co., Calif.; Santa Monica Canyon
“Pliocene”
robusta, Posidonia wengensis: Kittl Plastoholotype
Kitt], 1912, p. 18, pl. 1, fig. 12
Austria; Pederoa, Abteital
Middle Triassic, Wengener Schichten [holotype in Naturh. Staats-
mus. Wien]
robusta, Radiolites: Palmer Paratypes
Palmer, 1928a, p. 80
Jalisco, Mexico; Huescalapa
Cretaceous, Turonian
360
8506
457
7856
8740
7362
7562a
6250
7853
7968
410
9930
5377
223
224
7367
BULLETIN 300
rogersi, Lithophaga (Labis) attenuata: Berry Holotype
Berry, 1957, p. 76. Illustrated iz Keen, 1971, p. 68, fig. 140
Sonora, Mexico; Cholla Cove, Bahia de Adair
rogersi, Sphaerium: Hannibal Holotype
Hannibal, 1912b, p. 131, pl. 7, fig. 21. Also ix Taylor and Smith, 1971,
fics dts, 5
Tesla Qd, Calif.; 1/4 mile above Carnegie Pottery, Corral Hollow
Eocene
rostae, Barbatia (Acar): Berry Holotype
Berry, 1954a, p. 67. Illustrated zz Keen, 1971, p. 40, fig. 72
Sinaloa, Mexico; Mazatlan
rostratus, Inoceramus: Goldfuss Plastoholotype
Goldfuss, 1836, p. 110, pl. 115, fig. 3
Westphalia, Germany
Cretaceous [Goldfuss holotype 667 BM(NH) ]
rotunda, Immanitas: Palmer Paratypes
Palmer, 1928a, p. 32
Colima, Mexico; Paso de] Rio
Cretaceous, Cenomanian :
rugosa, Nucula: Odhner Paratypes
Odhner, 1919, p. 23
Tamatave, Madagascar
sacculifer, Volsella: Berry Holotype
Berry, 1953b, p. 407, pl. 28, figs. 1, 2
San Pedro Harbor, California
saibana, Nuculana (Saccella): Marks Paratype
Marks, 1951, p. 48
SW Ecuador; Zacachin corehole, 890-900’ depth
Miocene, Subibaja Fm
salazari, Monopieura: Palmer Paratype
Palmer, 1928a, p. 45, pl. 7, figs. 2, 3
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
sanctaeanae, Daonella: Smith Plastoholotype
Smith, 1914, p. 145, pl. 50, fig. 12, as sanctae-anae
Orange Co., Calif.; Santa Ana Mts., Silverado Canyon
Middle Triassic [holotype USNM 74365]
sanctaecrucis, Periploma: Arnold Holotype
Arnold, 1908a, p. 382, pl. 35, fig. 8. Also im Arnold, 1909, Illus. 2,
fig. 53
Santa Clara Co., Calif.; 2.5 miles SSW of Mayfield, E side Madera
Creek
Upper Miocene [Arnold’s specimen No. 1074]
sanjuanensis, Pecten (Pseudamusium) vancouverensis:
Clark and Arnold Holotype
Clark and Arnold, 1923, p. 140, pl. 16, fig. 5
Vancouver Island, British Columbia, Canada; Port San Juan, sea
cliffs 1/4 mile E of Providence Cove SU loc. NP 133
Oligocene? Sooke Fm
sanjuanensis, Pecten (Pseudamusium) vancouverensis:
Clark and Arnold Paratype
Clark and Arnold, 1923, p. 140, pl. 16, fig. 6
Vancouver Island, British Columbia, Canada; Port San Juan, sea
cliffs 1/4 mile E of Providence Cove SU loc. NP 133
Oligocene? Sooke Fm
santaclarana, Arca (Anadara): Loel and Corey Paratype
Ventura Co., Calif.; ridge W of mouth of Wiley Canyon UCMP loc.
A-252
Lower Miocene, Vaqueros Fm
360
361
7377
6571
6571a
5165
616
7880
7881
7882
7883
7884
7885
7887
7886
7888
STANFORD UNIVERSITY Tyres: SMITH 361
santaecruzensis, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 54, pl. 3, fig. 13
Santa Cruz Co., Calif.; Twobar Creek
Oligocene, San Lorenzo Fm
santaecruzensis, Pecten (Pecten): Arnold Paratype
Arnold, 1906, p. 54, pl. 3, fig. 12
Santa Cruz Co., Calif.; Bear Creek
Oligocene, San Lorenzo Fm
santamariensis, Arca (Arca): Reinhart Plastoholotype
Reinhart, 1937b, p. 183, pl. 28, figs. 4, 5, 7, 8, 11
Santa Barbara Co., Calif.; Fugler Point
Pliocene [holotype CIT 1381, now LACMNH 4072]
scarificata, Tivela: Berry Holotype
Berry, 1940b, p. 151, pl. 17, fig. 5
San Pedro, Calif.; NW corner of Beacon and Second Streets
Pleistocene
scarificata, Tivela: Berry Paratypes
Berry, 1940b, p. 151
San Pedro, Calif.; NW corner of Beacon and Second Streets
Pleistocene
schencki, Cardium: Wiedey Paratype
Wiedey, 1928, p. 143, pl. 17, fig. 4
Los Angeles Co., Calif.; Santa Monica Mts., Dry Canyon, 2 miles §
of Calabasas SU loc. 425
Middle Miocene, Temblor Fm_ [Wiedeys’ No. 431]
schencki, Chione: Loel and Corey Holotype
Loel and Corey, 1932, p. 224, pl. 42, fig. 5
San Luis Obispo Co., Calif.; Corral de] Piedra Creek
Lower Miocene, Vaqueros Fm
schencki, Glycymeris: Nicol Holotype
Nicol, 1947, p. 349, pl. 50, figs. 5, 6
Panama Canal Zone; 9° 18’ N, 79° 55’ + 200’ W_ SU loc. 2654
Miocene, Gatun Fm
schencki, Glycymeris: Nicol Paratype
Nicol, 1947, p. 349
Panama Canal Zone; 9° 16’ +4700’ N, 79° 54’ + 5800’ W SU loc.
2653
Miocene, Gatun Fm
schencki, Glycymeris: Nicol Paratype
Nicol, 1947, p. 349
Colon Province, Republic of Panama; 9° 21’ + 5000’ N, 79° 50’ +
1000’ W_ SU loc. 2656
Miocene, Gatun Fm
schencki, Glycymeris: Nicol Paratype
Nicol, 1947, p. 349, pl. 50, fig. 3
Panama Canal Zone; 9° 18’ N, 79° 55’ + 200’ W_ SU loc. 2654
Miocene, Gatun Fm
schencki, Glycymeris: Nicol Paratypes
Nicol, 1947, p. 349
Panama Canal Zone; 9° 18’ N, 79° 55’ + 200’ W_ SU loc. 2654
Miocene, Gatun Fm :
schencki, Glycymeris: Nicol Paratype
Nicol, 1947, p. 349, pl. 50, figs. 2, 4
Panama Canal Zone; 9° 18’ N, 79° 55’ + 200’ W_ SU loc. 2654
Miocene, Gatun Fm
schencki, Glycymeris: Nicol Paratype
Nicol, 1947, p. 349, pl. 50, fig. 1
Panama Canal Zone; 9° 18’ N, 79° 55’ + 200’ W_ SU loc. 2654
Miocene, Gatun Fm
362
789
790
8003
7279
7955
7284
7559
9895
215
7314
8001
8002
9518
BuLLeETIN 300
schencki, Thracia: Clark ex Tegland Ms Paratype
Clark, 1932, p. 808. Illustrated in Tegland, 1933, p. 112, pl. 6, fig. 8
Puget Sound, Wash.; beach between S side of entrance to Blakeley
Harbor and Restoration Point, Bainbridge Island SU loc. NP 103
Upper Oligocene, Blakeley Fm
schencki, Thracia: Clark ex Tegland Ms Paratype
Clark, 1932, p. 808. Illustrated in Tegland, 1933, pp. 112-113, pl. 6,
fig. 9
Puget Sound, Wash.; beach between S side entrance to Blakeley
Harbor and Restoration Point, Bainbridge Island SU loc. NP 103
Upper Oligocene, Blakeley Fm
schencki, Venericardia (Leuroactis): Verastegui Holotype
Verastegui, 1953, p. 50, pl. 4, figs. 6-8
Ventura Co., Calif.; Camulos Qd, Simi Hills, 2 miles NE of Simi Peak
Lower Eocene, Santa Susana Shale
scrippsensis, Donax: Hanna Plastoholotype
Hanna, 1927, p. 293, pl. 40, figs. 1, 12
San Diego Co., Calif.; Scripps Institution UCMP loc. 5089
Eocene, La Jolla Fm [holotype UCMP 30992]
secticostata, Glycymeris: Nicol Plastoholotype
Nicol, 1945, p. 623, pl. 85, fig. 3
Costa Rica; E Grape Point Creek
Miocene, Gatun Fm
semiplicata, Chione: Nomland Plastoholotype
Nomland, 1917b, p. 305, pl. 15, figs. 2a, 2b
Fresno Co., Calif.; near Coalinga UCMP loc. 2283
Miocene, Santa Margarita Fm [holotype UCMP 11318]
septata, Caprinuloidea: Palmer Paratype
Palmer, 1928a, p. 62, pl. 11, fig. 1
Jalisco, Mexico; Soyatlan de Adentro
Cretaceous, Cenomanian
septentrionalis, Halobia: Smith Plastoholotype
Smith, 1927, p. 118, pl. 98, fig. 1
Alaska; Keku Islet No. 1, Admiralty Island, Herring Bay USGS loc.
10196
Upper Triassic, lower Noric or upper Karnic [holotype USNM]
sespeensis, Pecten (Chlamys): Arnold Plastoholotype
Arnold, 1906, p. 69. pl. 8, fig. 3
Ventura Co., Calif.; Sespe Canyon
Miocene [holotype USNM]
sheridani, Macoma: Vokes Plastoholotype
Vokes, 1939, p. 92, pl. 14, fig. 21
San Benito Co., Calif.; Vallecitos UCMP loc. A-1154
Eocene, Domengine [holotype UCMP 15703]
simiana, Venericardia (Venericor): Verastegui Holotype
Verastegui, 1953, p. 47, pl. 4, figs. 2-4
Ventura Co., Calif.; Calabasas Qd, 1/2 mile NE of Hill 2150, Simi
Hills
Paleocene
simiana, Venericardia (Venericor): Verastegui Paratype
Verastegui, 1953, p. 47, pl. 4, fig. 1
Ventura Co., Calif.; Calabasas Qd, 1/2 mile NE of Hill 2150, Simi
Hills
Paleocene
singularis, Orobitella (lsorobitella): Keen Holotype
Keen, 1962, p. 323, figs. 4a-4c, 5a, 5b
Baja California del Norte, Mexico; Bahia de San Quintin, on mud
flats
7378
9829
508
8071
5205
5206
7281
64
212
70
235
STANFORD UNIVERSITY Types: SMITH 363
sisquocensis, Arca (Arca): Reinhart Plastoholotype
Reinhart, 1937b, p. 182, pl. 28, figs. 1-3
Santa Barbara Co., Calif.; Fugler Point
Pliocene [holotype CIT 1382 now LACMNH 4073]
sloati, Siliqua: Hertlein Paratype
Hertlein, 1961, p. 14
Point Bonita, Calif.
smithii, Panopea: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 79. Illustrated in Wiedey, 1929b, pl. 2,
fig
Alameda Co., Calif.; Tesla Qd, cut opposite R.R. crossing, Corral
Hollow. Arnold loc. C-141
Upper Eocene, ‘Tejon’ Fm
smithii, Panopea: Hall and Ambrose Paratype
Hall and Ambrose, 1916, p. 79
Alameda Co., Calif.; Tesla Qd, cut opposite RR crossing, Corral
Hollow
Upper Eocene, “Tejon” Fm
snohomishensis, Panope: Clark Holotype
Clark, 1925, p. 105, pl. 10, fig. 1
Opposite Snohomish, Wash.; ss on Fiddlers Bluffs, along Snohomish
River SU loc. NP 146
Oligocene, Lincoln Fm
snohomishensis, Panope: Clark Paratype
Clark, 1925, p. 105, pl. 11, fig. 2
Opposite Snohomish, Wash.; ss on Fiddlers Bluffs, along Snohomish
River SU loc. NP 146
Oligocene, Lincoln Fm
soledadensis, Tellina: Hanna Plastosyntype
Hanna, 1927, p. 291, pl. 42, fig. 2
San Diego Co., Calif.; Tecolote Creek UCMP loc. 5091
Eocene, La Jolla Fm [syntype UCMP 31369]
sookensis, Cardium: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 145, pl. 22, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Cardium: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 145, pl. 22, fig. 2
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Macoma: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 151, pl. 25, fig. 3
Vancouver Island, British Columbia, Canada; Sooke, ss and cgl on
sea cliffs between mouths of Muir and Kirby Creeks, W of Otter
Point SU loc. NP 129
Oligocene, Sooke Fm
sookensis, Modiolus: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 143, pl. 26, fig. 2
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouth of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
364
236
290
9907
8065
7974
12
8454
8453
8455
5178
9179
5180
5316
BuLLeETIN 300
sookensis, Modiolus: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 143, pl. 26, fig. 4
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Ostrea: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 138, pl. 17, fig. 2
Vancouver Island, British Columbia, Canada; Jordan River, sea cliffs
at mouth of Fossil Creek, 2 miles W of Sherringham Point SU loc.
NP 130
Oligocene, Sooke Fm
soperi, Avicula: Smith Plastoholotype
Smith, 1927, p. 112, pl. 96, fig. 9
Shasta Co., Calif.; N fork Squaw Creek, 3 miles N of Kellys Ranch
Upper Triassic, Hosselkus Ls [holotype USNM 74166]
spectri, Macoma (Psammacoma) panamensis: Hertlein and Strong
Paratype
Hertlein and Strong, 1949a, p. 91
Gulf of California, Mexico; Arena Bank, 45 fms
stainforthi, Anodontia: Marks Paratype
Marks, 1951, p. 69
SW Ecuador; §S of Progreso
Middle Miocene, upper Progresso Fm
stanfordensis, Pecten (Propeamusium): Arnold Holotype
Arnold, 1906, p. 91, pl. 23, fig. 4
Santa Clara Co., Calif.; Burke Ranch, 3 miles S of Stanford University
Miocene, Vaqueros Fm
stanfordia, Tivela: Hall Holotype
Hall, 1958, p. 53, pl. 6, figs. 3-5
Alameda Co., Calif.; La Costa Valley Qd, NE 1/4 Sec. 11, T 5S,R1E
SU loc. 3244
Upper Miocene, Briones Fm
stanfordia, Tivela: Hall Paratype
Hall, 1958, p. 53, pl. 6, figs. 1, 2
Alameda Co., Calif.; La Costa Valley Qd, NE 1/4 Sec. 11,T5S,R1E
Upper Miocene, Briones Fm
stanfordia, Tivela: Hall Paratype
Hall, 1958, p. 53, pl. 6, figs. 6, 7
Alameda Co., Calif.; La Costa Valley Qd, NE 1/4 Sec. 11, T 5S,R1E
Upper Miocene, Briones Fm
stantoni, Macrocallista: Waring Holotype
Waring, 1917, p. 77, pl. 14, fig. 6
Ventura Co., Calif.; Martinez area, Simi Hills SU loc. 2695
Lower Eocene, Martinez Fm
stantoni, Macrocallista: Waring Paratype
Waring, 1917, p. 77
Ventura Co., Calif.; Martinez area, Simi Hills SU loc. 2695
Lower Eocene, Martinez Fm
stantoni, Macrocallista: Waring Paratype
Waring, 1917, p. 77, pl. 14, fig. 1
Ventura Co., Calif.; Martinez area, Simi Hills SU loc. 2695
Lower Eocene, Martinez Fm
strongi, Arca (Barbatia): Loel and Corey Syntype
Loel and Corey, 1932, p. 183
Orange Co., Calif.; San Joaquin Hills, 2.5 miles N of Laguna Beach
UCMP loc. A-527
Lower Miocene, Vaqueros Fm
51
52
198
7969
61
6053
9265
9266
9267
9268
120
120a
120b
STANFORD UNIVERSITY TypEs: SMITH 365
Subdolus, Pecten (Plagioctenium): Hertlein Holotype
Hertlein, 1925a, p. 20, pl. 5, figs. 4, 7
San Diego Co., Calif.; Pacific Beach SU loc. 115
Pliocene, San Diego Fm
subdolus, Pecten (Plagioctenium): Hertlein Paratype
Hertlem), 19254, p. 20s ple 55 tis) 2
San Diego Co., Calif.; Pacific Beach SU loc. 115
Pliocene, San Diego Fm
subdolus, Pecten (Plagioctenium): Hertlein Paratype
Hertlein, 1925a, p. 20
Off Baja California, Mexico; Cedros Island SU loc. 116
Pliocene
subibajana, Nuculana (Saccella): Marks Paratypes
Marks, 1951, p. 50
SW Ecuador; Zacachtin corehole, 500-510’ depth
Miocene, Subibaja Fm
subimpressa, Leda: Howe Holotype
Howe, 1922, p. 97, pl. 10, fig. 3
Coos Bay, Ore. SU loc. NP 36
Pliocene, Empire Fm
subviridis, Lasaea rubra: Dall ex Carpenter Ms Neotype
Dall, 1899b, p. 881. Neotype selected by Keen, 1938, p. 29, pl. 2, figs.
1-3
Baja California, Mexico; San Martin Island
subyneziana, Pecten (Vertipecten) yneziana:
Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 198, pl. 31, fig. 3
Santa Barbara Co., Calif.; Camino Cielo, UCMP loc. B-6940
Eocene, “Coldwater” Ss
subyneziana, Pecten (Vertipecten) yneziana:
Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 198, pl. 31, fig. 5
Santa Barbara Co., Calif.; Camino Cielo UCMP loc. B-6940
Eocene, “Coldwater” Ss
subyneziana, Pecten (Vertipecten) yneziana:
Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 198, pl. 31, fig. 7
Santa Barbara Co., Calif.; Camino Cielo UCMP loc. B-6940
Eocene, ‘‘Coldwater” Ss
subyneziana, Pecten (Vertipecten) yneziana:
Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 198, pl. 31, fig. 2
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek UCMP loc.
B-6963
Eocene, Sacate-Gaviota Fm
suciensis, Thracia: Reagan Syntype
Reagan, 1924, p. 183, pl. 20, fig. 3
Puget Sound, Wash.; Sucia Islands
Upper Cretaceous, upper Chico Fm
suciensis, Thracia: Reagan Snytype
Reagan, 1924, p. 183, pl. 20, fig. 4
Puget Sound, Wash.; Sucia Islands
Upper Cretaceous, upper Chico Fm
suciensis, Thracia: Reagan Syntype
Reagan, 1924, p. 183, pl. 20, fig. 5
Puget Sound, Wash.; Sucia Islands
Upper Cretaceous, upper Chico Fm
366
143
5362
8004
5132
9899
8336
7996
7997
7918
5999
189
5188
5189
5190
BuLtetTin 300
superioris, Cardita: Waring Holotype
Waring, 1917, p. 91
Ventura Co., Calif.; McCray Wells SU loc. 8
Eocene, Tejon Fm [= Schedocardia brewerii (Gabb), teste Keen,
1949]
supramontereyensis, Yoldia: Arnold Holotype
Arnold, 1908a, p. 382, pl. 35, fig. 9. Also im Arnold, 1909, Illus. 2,
fig. 56
Santa Clara Co., Calif.; 2.5 miles S of Mayfield, “Tusk Gully” near
road
Upper Miocene [Arnold’s No. 1067]
susanaensis, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 22, pl. 5, figs. 1-4
Ventura Co., Calif.; Camulos Qd, McCray Wells, Oil Canyon
Lower Eocene, Santa Susana Shale
swartsi, Glycimeris [sic.|: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 620, pl. 17, fig. 2
Baja California, Mexico; Scammon Lagoon Qd, W side of Elephant
Mesa _ SU loc. 60 :
Miocene, Isidro Fm
symmetrica, Halobia: Smith Plastoholotype
Smith, 1927, p. 119, pl. 98, fig. 7
Alaska; Keku Islet No. 1, Admiralty Island, Herring Bay
Upper Triassic [holotype USNM 74182]
taberi, Chione (Chione) undatella: Parker Plastoholotype
Parker, 1949, p. 582, pl. 90, figs. 2, 4, 9
Gulf of California; loc. 2897 [holotype UCMP]
taliaferroi, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 38, pl. 1, fig. 15
San Luis Obispo Co., Calif.; Adelaida Qd. NW 1/4 NE 1/4 Sec. 30,
T 25 S,R10E, S of Williams Ranch on the Nacimiento River
Paleocene, Dip Creek Fm
taliaferroi, Venericardia (Pacificor): Verastegui Paratype
Verastegui, 1953, p. 38, pl. 1, fig. 16
San Luis Obispo Co., Calif.; Adelaida Qd. NW 1/4 NE 1/4 See. 30,
T 25 S,R10E, S of Williams Ranch on the Nacimiento River
Paleocene, Dip Creek Fm
tayloriana, Ostrea: Gabb Plastoholotype
Gabb, 1866, p. 34, pl. 12, figs. 60, 60a
San Marcos Pass, near Santa Barbara, Calif.
“Miocene” [holotype UCMP 12005]
tehamaensis, Arca: Stanton Plastoholotype
Stanton, 1895, p. 18, pl. 6, fig. 8. Also iz Reinhart, 1937, p. 174 [as
Parallelodon? (Gilbertwhitea?) tehamaensis (Stanton) ]
Tehama Co., Calif.; 5 miles N of Paskenta, Shelton’s Ranch
Cretaceous? upper Knoxville Fm [holotype USNM 23044]
tejonensis, Isocardia: Waring Holotype
Waring, 1914, p. 784. Illustrated in Waring, 1917, p. 93, pl. 15, fig. 14
Ventura Co., Calif.; Camulos Qd, 1.5 miles E of McCray Wells SU
loc. 2696
Upper Eocene, Tejon Fm [Llajas Fm, fide Keen and Bentson, 1944,
p. 54]
tejonensis, Isocardia: Waring Paratypes
Waring, 1914, p. 784
Ventura Co., Calif.; Camulos Qd. 1.5 miles E of McCray Wells
SU loc. 2696
Upper Eocene, Tejon Fm [Llajas Fm, fide Keen and Bentson, 1944,
p. 54]
6001
6002
7973
7975
7976
1977
5099
5134
5209
STANFORD UNIVERSITY Tyres: SMITH 367
teltschenensis, Daonella: Kittl Plastoholotype
Kitt], 1912, p. 33, pl. 1, fig. 18
Austria; Feuerkogel (Teltschen) Aussia
Upper Triassic, Karnic [holotype at Naturh. Staatsmus. Wien]
textrina, Arca: Stanton Plastosyntype
Stanton, 1895, p. 14, pl. 6, fig. 7. Also im Reinhart, 1937a, p. 175 [as
Nemodon? textrina (Stanton) ]
Tehama Co., Calif.; Cottonwood Creek, Cold Fork, near Stephenson’s
Cretaceous, “upper Knoxville Fm’ [syntype USNM 23045]
textrina, Arca: Stanton Plastosyntype
Stanton, 1895, p. 14, pl. 6, fig. 6. Also zz Reinhart, 1937a, p. 175
Tehama Co., Calif.; Cottonwood Creek, Cold Fork, near Stephenson’s
Cretaceous, “upper Knoxville Fm” [syntype USNM 23045]
thalmanni, Cavilucina (Pegophysema): Marks Paratype
Marks, 1951, p. 68
SW Ecuador; N of Pajan, Daule Basin
Middle Miocene, Daule Fm
thompsoni, Pitar (Lamelliconcha): Marks Holotype
Marks, 1951, p. 74, pl. 4, fig. 7
Republic of Panama; 6 miles E of Colon, on Roosevelt-Boyd Trans-
isthmian Highway SU loc. 2611
Miocene, lower Gatun Fm
thompsoni, Pitar (Lamelliconcha): Marks Paratype
Marks, 1951, p. 74, pl. 4, fig. 6
Republic of Panama; 6 miles E of Colon, on Roosevelt-Boyd Trans-
isthmian Highway SU loc. 2611
Miocene, lower Gatun Fm
thompsoni, Pitar (Lamelliconcha): Marks Paratype
Marks, 1951, p. 74
Republic of Panama; 6 miles E of Colon, on Roosevelt-Boyd Trans-
isthmian Highway SU loc. 2611
Miocene, lower Gatun Fm
tolmani, Pecten: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 82. Illustrated ix Wiedey, 1929b, p. 23,
plat, figs Z
Alameda Co., Calif.; Pleasanton Qd, Sunol, mouth of Welch Creek
Middle Miocene? Briones Fm?
topangaensis, Anadara (Anadara): Reinhart Paratypes
Reinhart, 1943, p. 53
Los Angeles Co., Calif.; Santa Monica Mts., Sec. 36, T 1 N, R15 W
Miocene, Topanga Fm
totiseptata, Sabinia: Palmer Paratype
Palmer, 1928a, p. 73
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
toulai, Sanguinolaria: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 625, pl. 20, fig. 2
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
townsendensis, Sanguinolaria (Nuttalina): Clark Holotype
Clark) 1925. ps 97,pli 185 fiz: 7
Townsend Bay, Wash.; ss sea cliffs between Classens Wharf and ship
canal estuary SU loc. NP 125
Oligocene, Lincoln Fm
368
5200
5201
5208
5207
454
453
9805
5815a
397
7307
430
430a
363
BuLLeETIN 300
townsendensis, Solen (Plectosolen): Clark Holotype
Clark, 1925, p. 97, pl. 22, fig. 10
Skamokawa, Wash.; ss bluffs along Skamokawa River above big
bend, 1 mile E of jet. of main and middle forks SU loc. NP 272
Oligocene, Lincoln Fm
townsendensis, Solen (Plectosolen): Clark Paratype
Clark 1925: p: 97, pl; 22) fie: 7
Skamokawa, Wash.; ss bluffs along Skamokawa River above big
bend, 1 mile E of jct. of main and middle forks SU loc. NP 272
Oligocene, Lincoln Fm
townsendensis, Tellina: Clark Holotype
Clark, 1925, p. 94, pl. 12, fig. 12
Oregon; Grays River, in tuffaceous ss in R.R. cut on logging road
up Fossil Creek, 3 miles above jct. with Grays River SU loc. NP 278
Oligocene, Lincoln Fm
townsendensis, Tellina: Clark Paratype
Clark, 1925, p. 94, pl. 12, fig. 11
Townsend Bay, Wash.; from sea cliffs between Classens Wharf and
ship canal estuary SU loc. NP 125 ;
Oligocene, Lincoln Fm
transpacifica, Unio: Hannibal Holotype
Hannibal, 1912b, p. 123, pl. 7, fig. 18a. Also iz Taylor and Smith,
1971, figs. 3, 4 (as Plesielliptio)
Wash.; Olequa Creek, at shoals, 1.5 miles above Little Falls
Eocene [late Eocene, Cowlitz Fm, fide Taylor and Smith, 1971, p.
309]
transpacifica, Unio: Hannibal Paratype
Hannibal, 1912b, p. 123, pl. 7, fig. 18b. Also ix Taylor and Smith,
1971, figs. 7, 10 (as Plesielliptio)
Wash.; Olequa Creek, at shoals, 1.5 miles above Little Falls
Eocene [late Eocene, Cowlitz Fm, fide Taylor and Smith, 1971, p.
309]
tremperi, Corneocyclas: Hannibal Holotype
Hannibal, 1912b, p. 137, pl. 7, fig. 22. Also im Taylor and Herring-
ton, 1962, pl. 28, figs. 1, 2 (as Pisidium)
San Bernardino Mts., Calif.; Bluff Lake Cienaga
tremperi, Corneocyclas: Hannibal Paratype
Hannibal, 1912b, p. 137
San Bernardino Mts., Calif.; Bluff Lake Cienaga
triangulatus, Crassatellites: Waring Holotype
Waring, 1917, p. 59, pl. 9, fig. 1
Los Angeles Co., Calif.; Calabasas sheet, S of Santa Monica Mts.
Cretaceous, Chico Fm
truncata, Tapes: Gabb Plastoholotype
Gabb, 1866, p. 25, pl. 7, fig. 44
San Benito Co., Calif.; Griswold’s “Monterey”
Miocene, T'emblor Fm [holotype UCMP 12335]
turneri, Pecten (Patinopecten): Arnold Holotype
Arnold, 1906, p. 106, pl. 35, fig. 2
Marin Co., Calif.; near Tomales Bay in Arroyo San Antonio
Pliocene
turneri, Pecten (Patinopecten): Arnold Paratype
Arnold, 1906, p. 106, pl. 35, fig. 3
Marin Co., Calif.; near Tomales Bay in Arroyo San Antonio
Pliocene
turneri, Pecten (Patinopecten): Arnold Paratype
Arnold, 1906, p. 106, pl. 34, fig. 4
Marin Co., Calif.; near Tomales Bay in Arroyo San Antonio
Pliocene
5236
5235
9243
5446
7298
831
8585
517
6003
246
63
211
STANFORD UNIVERSITY Types: SMITH 369
twinensis, Kellia ?: Clark Holotype
Clark, 1925, p. 90, pl. 18, fig. 8
Twin, Wash.; sea cliffs W of West Twin River for a distance of 3/4
mile SU loc. NP 120
Oligocene, Blakeley Fm
twinensis, Macoma: Clark Holotype
Clark, 1925, p. 96, pl. 12, fig. 7
Townsend Bay, Wash.; Port Hadlock, Help-Me-Jack Rock SU loc.
NB A127
Oligocene
twinensis, Spisula: Clark Holotype
Clark, 1925, p. 103, pl. 16, fig. 6
Twin, Wash.; sea cliffs W of West Twin River for a distance of 3/4
mile SU loc. NP 120
Oligocene, Blakeley Fm
umnaka, Cardita: Willett Paratype
Willett, 1932, p. 87
Umnak Island, Alaska
umpquaensis, Gari hornii: Turner Plastoholotype
Turner, 1938, p. 62, pl. 7, fig. 11
Douglas Co., Ore.; Little River UCMP loc. A-662
Eocene, Umpqua Fm_ [holotype UCMP 33149]
undulata, Pleuromya (?): Davis Holotype
Davis, 1913, p. 454, text fig. 4
Monterey Co., Calif.; Slates Hot Springs
“Jurassic,” “Franciscan” Fm
ursipes, Spondylus: Berry Holotype
Berry, 1959, p. 107. Illustrated iz Keen, 1971, p. 98, fig. 213
Baja California, Mexico; Isla Angel de la Guarda, Puerto Refugio
valentinei, Chione: Wiedey Holotype
Wiedey, 1929c, p. 284, pi. 31, fig. 4
Santa Clara Co., Calif.; 2 miles S of Mayfield SU loc. 448
Miocene, Temblor Fm?
vancouverensis, Arca: Meek Plastoholotype
Meek, 1864a, p. 40. Illustrated 7x Meek, 1876, p. 356, pl. 3, figs. 5, 5a.
Also iz Reinhart, 1937a, p. 171, pl. 27, fig. 4 [as Parallelodon
(Nanonavis) vancouverensis (Meek) |
Vancouver Island, British Columbia, Canada; Comox
Cretaceous [holotype USNM 12398]
vancouverensis, Chione: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 147, pl. 20, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Glycimeris [sic.]: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 137, pl. 27, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Glycimeris [sic.]: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 137, pl. 27, fig. 5
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
262
289
264
263
59226
9405
520
9215
5216
5217
BuL_eTIN 300
vancouverensis, Metis: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 150, pl. 22, fig. 3
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Semele: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 151, pi. 27, fig. 4
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Tellina: Clark and Arnold Plastoholotype
Clark and Arnold, 1923, p. 149
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouth of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129 (= CAS loc. 231)
Oligocene, Sooke Fm_ [holotype CAS 599]
vancouverensis, Tellina: Clark and Arnold - Paratype
Clark and Arnold, 1923, p. 149, pl. 22, fig. 5
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
vanwinkleae, Pecten: Clark Holotype
Clark, 19255 (p82) pl. 15, fig: 2
Wash.; ss bluffs along Porter Creek, 3/4 mile above Porter SU loc.
NP 54
Lower Oligocene, Lincoln Fm
vaquerosensis, Cardium (Trachycardium): Arnold Paratype
Arnold, 1908a, p. 378
San Mateo Co.. Calif.; Mindego Creek, 1 mile above Alpine Creek
Lower Miocene, Vaqueros Fm
vaquerosensis, Tivela (?): Wiedey Holotype
Wiedey, 1929c, p. 288, pl. 33, fig. 1
Monterey Co., Calif.; Los Vaqueros Valley, type section of Vaqueros
Fm _ SU loc. 200
Lower Miocene, Vaqueros Fm
vaughani, Pecten (Lyropecten): Arnold Holotype
Arnold, 1906, p. 81, pl. 23, figs. 3, 3a, 3b
Ventura Co., Calif.; Ojai Valley
Lower Miocene
veneriformis, Spisula: Clark Holotype
Clark, 1925, p. 103, pl. 16, fig. 3
Oregon coast W of Coos Bay; sea cliffs at Tunnel Point SU loc.
NP 42
Oligocene, Lincoln Fm
veneriformis, Spisula: Clark Paratype
Clark, 1925, p. 103, pl. 16, fig. 1
Wash.; bluffs along Porter Creek, 1/4 to 1 mile above old log dam
at Porter SU loc. NP 56
Oligocene, Lincoln Fm
veneriformis, Spisula: Clark Paratype
Clark, 1925, p. 103, pl. 16, fig. 2
Porter, Wash.; ss cut on Lytle logging R.R. near top of ridge 1 mile
above switch SU loc. NP 55
Oligocene, Lincoln Fm
8333
159
6942
8601
518
26
5581
164
7561
7565
8016
STANFORD UNIVERSITY TyPEs: SMITH 371
venturaensis, Pecten (Chlamys): Waterfall Plastoholotype
Waterfall, 1929, p. 84, pl. 6, fig. 4
Ventura Co., Calif.; E center Sec. 21, T 3 N, R 21 W
Pliocene, Pico Fm [holotype UCMP 31416]
venturensis, Venericardia planicosta: Waring Holotype
Waring, 1915, map folio fig. 12. Also in Waring, 1917, p. 80, pl. 11,
figs. 6, 7
Ventura Co., Calif.; Calabasas sheet, 3 miles NE of Simi Peak SU
loc. 2697
Lower Eocene, Martinez Fm
vernicosa, Astarte: Dall Paratype
Dall, 1903a, p. 948
Icy Cape, Alaska; 15 fms
vespertina, Ostrea: Conrad Plastolectotype
Conrad, 1854, p. 300. Lectotype selected by Woodring, 1938, p. 43, pl.
8, figs. 3, 8
Calif.; “near San Diego” [probably Carrizo Creek fide Woodring,
1938]
“Miocene” [probably Pliocene] [lectotype ANSP 13366]
vickeryi, Chione: Wiedey Holotype
Wiedey, 1929c, p. 286, pl. 32, fig. 4
Santa Clara Co., Calif.; E of San Jose, Alum Rock Canyon, 500 yds
upstream from the falls. SU loc. 451
Middle Miocene, upper Monterey Fm
vickeryi, Pecten (Lyropecten): Trask Holotype
Trask, 1922, p. 148, pl. 4, fig. 1
Alameda Co., Calif.; Pleasanton Qd, vicinity of McGuire Peaks
Miocene, Briones Fm
vigilia, Acila (Acila) divaricata: Schenck Plastoholotype
Schenck, 1936, p. 101, pl. 17, figs. 1-6
Japan; off S coast of Yesso [Hokkaido], 175 fms Albatross Sta. 5038
[holotype USNM 406502]
virginalis, Opis: Waring Holotype
Waring, 1917. p. 78, pl. 14, fig. 4
Venture Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm
vivari, Sabinia: Palmer Paratype
Palmer, 1928a, p. 74, pl. 14, fig. 4
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
vivari, Sabinia: Palmer Paratype
Palmer, 1928a, p. 74, pl. 13, fig. 4
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
vivari, Sabinia: Palmer Paratypes
Palmer, 1928a, p. 74
Colima, Mexico; Paso del Rio
Cretaceous, Cenomanian
vogdesi, Pecten (Pecten): Arnold Holotype
Arnold, 1906, p. 100, pl. 33, fig. 1
Los Angeles Co., Calif.; San Pedro
Pleistocene, San Pedro Fm
vokesi, Venericardia (Leuroactis): Verastegui Paratype
Verastegui, 1953, p. 61, pl. 14, fig. 3
Kings Co., Calif.; Cholame Qd, Reef Ridge sheet, SW cor. Sec. 17,
T 23 S,R17E, 1/2 mile E of Big Tar Canyon
Eocene, Avenal Fm
372
7280
816
5230
5232
5233
5234
5340
8024
8460
8461
7877
7878
7879
BuLLeTIN 300
vorbei, Tellina: Hanna Plastoholotype
Hanna, 1927, p. 292, pl. 40, fig. 16
San Diego Co., Calif.; Soledad Canyon UCMP loc. 5074
Eocene, La Jolla Fm_ [holotype UCMP 30984]
wairarapaensis, Glycimeris [sic.] (Grandaxinea): Powell
Powell, 1938, p. 158 Paratype
New Zealand; Castle Point, SE coast of North Island
Pliocene, Nukumaruan stage
washingtonensis, Mytilus: Clark Holotype
Clark 1925. ps 85; pl. 9) fiz.
Freshwater Bay, Wash.; point E of old shingle warehouse SU loc.
NP 155
Oligocene
washingtoniana, Corbis: Clark Holotype
Clark, 1925, p. 90, pl. 20, figs. 2, 3
Port Townsend, Wash.; sandy shales in sea cliffs, S shore of Mystery
Inlet, Scow Bay SU loc. NP 126
Oligocene, Keasey Fm
washingtoniana, Corbis: Clark ; Paratype
Clark, 1925, p. 90, pl. 20, fig. 1
Port Townsend, Wash.; sea cliffs on S shore of Mystery Inlet, Scow
Bay
Oligocene, Keasey Fm
washingtoniana, Corbis: Clark Paratype
Clark, 1925, p. 90, pl. 20, fig. 4
Port Townsend, Wash.; sea cliffs on S shore of Mystery Inlet, Scow
Bay
Oligocene, Keasey Fm
waylandi, Anadara: Cox Paratype
Cox, 1927, p. 34
East Africa; Ras Tungwe, Pemba Island
Lower Miocene
weaveri, Venericardia (Pacificor): Verastegui Holotype
Verastegui, 1953, p. 31, pl. 21, figs. 3, 4
Wash.; 1.25 miles NW of Vader on SE bank of Stillwater Creek
Upper Eocene, Cowlitz Fm
welchensis, Ventricolaria: Hall Holotype
Hall, 1958, p. 54, pl. 7, figs. 3, 4
Contra Costa Co., Calif.; 1 mile NE of Hercules SU loc. 3255
Upper Miocene, Cierbo Fm
welchensis, Ventricolaria: Hall Paratype
Hall, 1958) p. 54, pl. 7, fir. 5
Alameda Co., Calif.; La Costa Valley Qd. NE 1/4 Sec. 1,T5S,R1E
SU loc. 3239
Upper Miocene, Briones Fm
whaleyi, Glycimeris [sic.]: Nicol Paratype
Nicol, 1947, p. 347, pl. 50, fig. 7
Fresno Co., Calif.; near Arroyo Ciervo, Sec. 36, T 16 S, R 13 E,
2000’ N, 400’ W of SE corner of section, 800’ S of point where the
first Temblor “reef” crosses Arroyo Ciervo
Miocene, Temblor Fm?
whaleyi, Glycimeris [sic.|: Nicol Paratypes
Nicol, 1947, p. 347
Fresno Co., Calif.; near Arroyo Ciervo, 2000’ N, 400’ W of SE cor.
Sec. 36, T 16 S,R 13 E
Miocene, Temblor Fm?
5280
5280a
6234
9517
5228
92952
7810 (T)
5204
386
386a
386b
9905
8053
10302
8084
8057
STANFORD UNIveRsITY Tyres: SMITH 373
whiteavesi, Parallelodon (Nanonavis): Reinhart Plastosyntypes
Reinhart, 1937a, p. 172. Illustrated iz Whiteaves, 1879, pl. 19, figs.
1, la (as Nemodon vancouverensis Meek)
Vancouver Island, British Columbia, Canada; Blunden Point
Cretaceous [syntypes Geol. Surv. Canada 5684, 5684a ]
willetti, Astarte: Dall Paratypes
Dall, 1903a, p. 948
Forrester Island, Alaska; 50 fms
williamsi, Mactra (Mactra): Berry Holotype
Berry, 1960, p. 116. Illustrated zz Keen, 1971, p. 202, fig. 486
Off La Libertad, Ecuador; 10 fms
willipaensis, Trinacria: Clark Holotype
Clark, 1925, p. 81, pl. 9, figs. 5, 10
N of Holcomb, Wash.; ss bluffs along Willipa River SU loc. NP 253
Oligocene, Keasey Fm
willipaensis, Trinacria: Clark Paratype
Clark, 1925, p. 81, pl. 9, fig. 8
N of Holcomb, Wash.; ss bluffs along Willipa River SU loc. NP 253
Oligocene, Keasey Fm
woodsi, Ethmocardium: Marwick Plastoholotype
Marwick, 1944, p. 259, pl. 36, fig. 21
New Zealand; Selwyn Rapids, Canterbury
Upper Cretaceous, Piripauan stage, upper Senonian [holotype at
N.Z. Geol. Surv.]
yaquinensis, Mulinia (?): Clark Holotype
Clark, 1925, p. 105, pl. 17, fig. 1
Yaquina, Ore.; ss in sea cliffs along Yaquina Bay SU loc. NP 306
Oligocene
youngi, Cucullaea: Waring Holotype
Waring, 1917, p. 59, pl. 8, fig. 12
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon, N of Simi fault
Upper Cretaceous, Chico Fm
youngi, Cucullaea: Waring Paratypes
Waring, 1917, p. 59
Ventura Co., Calif.; Calabasas sheet, Bell’s Canyon, N of Simi fault
Upper Cretaceous, Chico Fm
yukonensis, Pecten (Entolium): Smith Plastoholotype
Smith, 1927, p. 122, pl. 101, fig. 9
Alaska; S bank Yukon River opposite Nation River
Upper Triassic [holotype USNM 74199]
zacae, Tellina (Tellinella): Hertlein and Strong Paratype
Hertlein and Strong, 1949a, p. 65
Gulf of California; Arena Bank, 35 fms
zeta, Flaventia: Popenoe Plastoholotype
Popenoe, 1937, p. 393, pl. 48, fig. 9
Santa Ana Mts., Calif.; CIT loc. 1068
Cretaceous, Turonian [holotype UCLA 40654]
zeltbergensis, Inoceramus humboldti: Heinz Plastoholotype
Heinz, 1928, p. 35, pl. 3, fig. 1
Hanover, Germany; Zeltberg bei Liineberg
Upper Cretaceous, u. 1. Emscher Fm [holotype at Geologisches
Staatinstitut, Hamburg]
zeteki, Mytilopsis: Hertlein and Hanna Paratypes
Hertlein and Hanna, 1949, p. 15
Panama Canal Zone; Miraflores Locks
374
8900
7620
9429
9028
8684
9006
6503
9091
6471
6469
5436
5435
BuLLETIN 300
CEPHALOPODA
acutus, Aspenites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 96, pl. 3, figs. 1, 2
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75249]
adicrus, Ammonites: Waagen Plastoholotype
Waagen, 1867, p. 591, pl. 25, figs. 1a, 1b
Schwaben, Germany; Gingen, Vilsthale
Jurassic, Dogger [cast from Pal. Mus. Wien]
alexandrae, Gymnites: Smith Paratype
Smith, 1914, p. 52, pl. 25, fig. 1
West Humboldt Range, Nevada; Fossil Hill, S American Canyon
SU loc. 1780
Middle Triassic, Star Peak Fm
alexandrae, Gymnites: Smith Plastoholotype
Smith, 1914, p. 53, pl. 26, figs. 1, 2
West Humboldt Range, Nevada; Fossil Hill, between Troy Canyon
and § fork of American Canyon ;
Middle Triassic, Star Peak Fm_ [holotype USNM 74300]
allani, Gastroplites: McLearn Plastoholotype
McLearn, 1931, p. 5, pl. 1, fig. 10
Alberta, Canada; Peace River, 20 miles below Cadotte River
Lower Cretaceous, Peace River Ss_ [holotype Geol. Surv. Canada
6337]
alternans, Acrochordiceras: Smith Plastoholotype
Smith, 1914, p. 38, pl. 32, figs. 15-17
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74326]
alternecostatus, Perisphinctes: Steiger Plastoholotype
Steiger, 1914, p. 483, pl. 104, figs. 1a, 1b
Himalaya Mts.
Upper Jurassic, upper Malm, Spiti Shale
altilis, Ceratites: Smith Plastoholotype
Smith, 1914, p. 83, pl. 67, figs. 19-21
West Humboldt Range, Nevada; Fossil Hill, S fork of American
Canyon
Middle Triassic [holotype USNM 74394]
ambiensis, Paranorites: Waagen Plastoholotype
Waagen, 1895, p. 158, pl. 22, fig. 1
Punjab, India; Salt Range, Amb (Stachella beds)
Triassic, Ceratite [holotype Palaeont. Inst. Wiener Univ. 4041]
ammonoides, Proptychites: Waagen Plastosyntype
Waagen, 1895, p. 171, pl. 17, fig. 1
Punjab, India; Salt Range, W of Khoora
Triassic, Ceratite [syntype Palaeont. Inst. Wiener Univ. 4236]
andersoni, Arcestes: Hyatt and Smith Holotype
Hyatt and Smith, 1905, p. 74, pl. 56, figs. 1-3
West Humboldt Range, Nevada; Muttleberry Canyon, 8 miles SE of
Lovelock
Upper Triassic
andersoni, Arcestes: Hyatt and Smith Paratype
Hyatt and Smith, 1905, p. 74, pl. 56, figs. 4-6
West Humboldt Range, Nevada; Muttleberry Canyon, 8 miles SE of
Lovelock
Upper Triassic
5497
8685
8928
8929
8930
8931
8931la
8945
9083
7597
5494
5488
8757
8804
8795
8890
8932
STANFORD UNIVERSITY TyrrEs: SMITH 375
andinus, Macrocephalites: Burckhardt Plastoholotype
Burckhardt, 1903, p. 33, pl. 3, figs. 10-12
Chile; Comisaria Lonquimay, Rio Colorado
Jurassic, lower Callovian
anguinus, Gastroplites: McLearn Plastoholotype
McLearn, 1931, p. 5, pl. 1, fig. 11
Alberta, Canada; Peace River, 8 miles below Cadotte River
Lower Cretaceous, Peace River Ss_ [holotype Geol. Sur. Canada 6338]
angulatus, Cordillerites: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 110, pl. 2, figs. 1-3 (type 8928), 4, 5 (type
8929), 6 (type 8930) ; pl. 68, figs. 1-3 (type 8931a), 4-7 (type 8931)
Aspen Ridge, Idaho; Wood Canyon, 9 miles NE of Soda Springs
Lower Triassic, Meckoceras zone [syntypes USNM 75247, 75347,
75300]
apostolicus, Celtites: Smith Plastoholotype
Smith, 1932, p. 104, pl. 48, figs. 1, 2
Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74989]
applanatus, Ceratites: Smith Plastoholotype
Smith, 1914, p. 80, pl. 53, figs. 9-11
West Humboldt Range, Nevada; Fossil Hill, S fork of American
Canyon
Middle Triassic [holotype USNM 74372]
aquilaensis, Scaphites: Reeside Plastoholotype
Reeside, 1927b, p. 25, pl. 19, figs. 1-5
Fergus Co., Montana; Willow Creek, 6 miles above Ft. Maginnis-
Junction City road
Upper Cretaceous, Eagle Ss_ [holotype USNM 73348 |
araucanus, Macrocephalites: Burckhardt Plastoholotype
Burckhardt, 1903, p. 30, pl. 3, figs. 1-3
Chile; Comisaria Lonquimay, Rio Colorado
Jurassic, lower Callovian
argentina, Witchellia: Burckhardt Plastoholotype
Burckhardt, 1903, p. 17, pl. 1, figs. 15-17
Argentina: Mendoza Province, Cerro Puchén
Jurassic, lower Dogger
arnoldi, Paralecanites: Hyatt and Smith Plastoholotype
Hyatt and Smith. 1905, p. 136, pl. 64, figs. 1-4
Idaho; Aspen Ridge, Wood Canyon
Lower Triassic, Meekoceras zone [holotype USNM 75295]
arthaberi, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 56, pl. 32, figs. 26-28
Bear Lake Co., Idaho; NE end of Bear Lake, 1 mile NE of Hot
Springs
Lower Triassic, Meekoceras zone [holotype USNM 74973]
aspenensis, Flemingites: Smith Plastoholotype
Smith, 1932, p. 52, pl. 23, figs. 6-8
SE Idaho; 5 miles E of Grays Lake
Lower Triassic, Meekoceras zone [holotype USNM 74919]
attenuatus, Dalmatites: Smith Plastoholotype
Smith, 1932, p. 81, pl. 57, figs. 11-13
Idaho; Paris Canyon, 1.5 miles W of Paris
Lower Triassic, Columbites zone [holotype USNM 75023]
austini, Prosphingites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 72, pl. 7, figs. 1-4
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75256]
376
618
8796
7591
9105
8918
8730
9035
9082
8866
7607
9045
8867
6497
BuLietin 300
Baculites sp., of Baculites anceps group: Nomland and Schenck
“Holotype”
Nomland and Schenck, 1932, fig. 4
Monterey Co., Calif.; Slate’s Hot Springs, on sea coast NE 1/4 See. 9,
T21S,R3 E SU loc. 929
Cretaceous
bannockensis, Flemingites: Smith Plastoholotype
Smith, 1932, p. 52, pl. 23, figs. 18-20
SE Idaho; Aspen Mts., Slug Creek, 14 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74922]
bassleri, Desmoscaphites: Reeside Plastoholotype
Reeside, 1927b, p. 16, pl. 21, fig. 17
San Juan Co., New Mexico; just W of Hogback Mt. and 1 mile N of
Shiprock-Farmington Rd.
Upper Cretaceous, Mancos Shale (280’ below top) [holotype USNM
73358 |
beecheri, Ceratites: Smith Plastoholotype
Smith, 1914, p. 94, pl. 43, figs. 15-17
West Humboldt Range, Nevada; Fossil Hill, S fork of American
Canyon :
Middle Triassic [holotype USNM 74349]
bicarinatus, Lanceolites: Smith Plastoholotype
Smith, 1932, p. 90, pl. 55, figs. 1-3
Elko Co., Nevada; 70 miles S of Wells
Lower Triassic, Meekoceras zone [holotype USNM 75013]
bispinosum, Trachyceras (Trachyceras): Johnston Paratype
Johnston, 1941, p. 487 (cited as No. 3)
New Pass Range, Nevada
Upper Triassic, Star Peak Fm
bittneri, Xenodiscus: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 123, pl. 20, figs. 5-7
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [holotype USNM 74460]
bonaevistae, Dinarites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 162, pl. 60, figs. 1-4, as bonae-vistae
West Humboldt Range, Nevada; Buena Vista Canyon
Middle Triassic [holotype USNM 74383]
bonnevillense, Dagnoceras: Smith Plastoholotype
Smith, 1932, p. 65, pl. 29, figs. 9-11
Idaho; Wood Canyon, 9 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74949]
brevis, Scaphites nodosus: Meek Plastoholotype
Meek, 1876, p. 426, pl. 25, figs. la-1c
Montana; Yellowstone River near Miles City
Upper Cretaceous, Pierre Shale [holotype USNM 367]
breweri, Eutomoceras: Smith Plastoholotype
Smith, 1914, p. 61, pl. 28, figs. 1-4
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74312]
bridgesi, Dagnoceras: Smith Plastoholotype
Smith, 1932, p. 65, pl. 31, figs. 1-3
Idaho; Slug Creek, 14 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74956]
broilii, Perisphinctes (Virgatosphinctes): Uhlig Plastoholotype
Uhlig, 1910, p. 336, pl. 91, fig. 1 (reversed)
Himalaya Mts., India; Shangra Laptel, Gnari-Khorsum
Upper Jurassic, Spiti Shale, Chidamu beds
9617
9617a
9617b
9032
8688
5610
5611
8935
6485
8537
8933
8915
8916
8917
8925
6468
STANFORD UNIVERSITY TYPES: SMITH 377
californicum, Delepinoceras: Gordon Paratypes
Gordon, 1964, p. A19, pl. 2, figs. 10 (type 9167a), 15-17 (type 9167)
eye Co., Calif.; Panamint Range, Cottonwood Mts., near Rest Spring
Upper Mississippian, Perdido Fm
calli, Gymnites: Smith Plastoholotype
Smith, 1914, p. 53, pl. 26, fig. 1
West ‘Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74306]
canadensis, Hoplites: Whiteaves Plastoholotype
Whiteaves, 1893, p. 118, pl. 11, figs. 3-5
Alberta, Canada; Peace River, 20 miles below Cadotte River
Lower Cretaceous, Peace River Ss [holotype Geol. Surv. Canada
7430]
carbonarius, Bactrites: Smith Holotype
Smith, 1903, p. 31, pl. 6, fig. 9
Independence Co., Arkansas; near Moorfield, on O. P. Goodwin
farm
Carboniferous, Fayetteville Fm, St. Louis-Chester stage
carbonarius, Bactrites: Smith Paratype
Smith, 1903, p. 31, pl. 6, figs. 10, 11
Independence Co., Arkansas: near Moorfield, on O. P. Goodwin
farm
Carboniferous, Fayetteville Fm, St. Louis-Chester stage
carpenteri, Owenites: Smith Plastoholotype
Smith, 1932, p. 100, pl. 54, figs. 31-32
Inyo Co., Calif.; Inyo Range, Union Wash, 15 miles SE of Inde-
pendence
Lower Triassic, Owenites subzone [holotype USNM 75012]
cautleyi, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 279, pl. 78, fig. 1 (fig. inverted)
Tibet; Laptel, Gnari-Khorsum
Jurassic, upper Malm, Spiti Shale
chicoensis, Baculites: Trask Neotype
Matsumoto, 1959a, p. 145, pl. 36, fig. 2, text fig. 60
Butte Co., Calif.; E bank of Chico Creek SU loc. 2609
Upper Grefaceous, Chico Fm
columbianus, Paranannites: Smith Plastoholotype
Smith, 1932, p. 99, pl. 32, figs. 11-13
Idaho; Wood Canyon, 9 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74968]
compactus, Lanceolites: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 113, pl. 5, figs. 7, 8 (type 8916); pl. 78,
figs. 9-11 (type 8917)
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [syntypes USNM 75252, 75254,
75281]
compressa, Ussuria: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 89, pl. 3, figs. 6, 7
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, ” Meekoceras zone [holotype USNM 75250]
compressus, Flemingites: Waagen Plastoholotype
Waagen, 1895, p. 202, pl. 15, fig. 1; pl. 16, fig. 1
Punjab, India; Koofri, Salt Range [east from Paleont. Inst. Wiener
Univ. ]
Triassic, Ceratite
378
8749
8948
8761
9115
8856
7598
10015
10016
9108
6492
9067
7594
BuLtetTIn 300
compressus, Marshallites: Matsumoto Plastoholotype
Matsumoto, 1955a, pp. 123-124, pl. VIII, figs. 1a, 1b
Hokkaido, Japan; Teshio Province, Abishinai Valley, loc. T608, bed
IIb
Cretaceous, Paleogyliakian [cast from Dept. of Geology, Kyushu
Univ., specimen GK-H-2751 = GT-I-3231]
consanguineus, Columbites: Smith Plastoholotype
Smith, 1932, p. 106, pl. 46, figs. 1, 2
Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74983]
cordilleranus, Xenodiscus: Smith Plastoholotype
Smith, 1932, p. 43, pl. 24, figs. 21-23
Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74926]
cornatus, Ceratites: Smith Plastoholotype
Smith, 1914, p. 98, pl. 62, figs. 1-4
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74387]
corrugata, Meekoceras mushbachanum: Smith Plastoholotype
Smith, 1932, p. 61, pl. 38, fig. 1
Idaho; NE end of Bear Lake, 1 mile NE of Hot Springs
Lower Triassic, Meekoceras zone [holotype USNM 74980]
costatus, Scaphites aquilaensis: Reeside Plastoholotype
Reeside, 1927b, p. 25, pl. 19, figs. 10-13
Park Co., Wyoming; Sec. 25, T 58 N, R 100 W
Upper Cretaceous, Telegraph Creek Fm, Elk Basin Ss mbr_ [holotype
USNM 73351]
costula, Fontannesia: Imlay Paratype
Imlay, 1973, p. 57, pl. 4, figs. 22-24
Grant Co., Ore.; SW 1/4, NE 1/4 Sec. 29, T 18 S, R 26 E, 600’ S of
head of gully draining SSW from North Ammonite Hill
Middle Jurassic, Snowshoe Fm, near top of lower 1/3 of Weberg Mbr,
Bajocian stage
costula, Fontannesia: Imlay Paratype
Imlay, 1973, p. 57, pl. 4, figs. 18-20
Crook Co., Ore.; near Wade Butte, a little W of center of Sec. 24,
ApelSES) Ro24: E
Middle Jurassic, Bajocian stage Snowshoe Fm, Weberg mbr
crassicornu, Ceratites: Smith Plastoholotype
Smith, 1914, p. 95, pl. 43, figs. 11, 12
West Humboldt Range, Nevada; Fossil Hill, S fork of American
Canyon
Middle Triassic [holotype USNM 74348]
crassicostatus, Ceratites (Hollandites) japonicus: Shimizu
Plastoholotype
Shimizu, 1930, p. 66, pl. 24, fig. 2
Oshika-gun, Japan; Inai, Inai-mura
Triassic
crassus, Lecanites: Smith Plastoholotype
Smith, 1914, p. 66, pl. 89, figs. 1, 2
West Humboldt Range, Nevada; S fork American Canyon
Middle Triassic (holotype USNM 74424]
crassus, Scaphites hippocrepis: Reeside Plastoholotype
Reeside, 1927b, p. 23, pl. 17, figs. 8-13
Sheridan Co., Wyoming; 2 miles W of Parkman, SW 1/4 Sec. 33,
TSS Nae or WW
Upper Cretaceous, Steele Shale [holotype USNM 73336]
8805
8562
8808
9048
8488
6464
6476
10007
10012
10013
10009
10010
10011
10014
10008
STANFORD UNIVERSITY Typrs: SMITH 379
cristatum, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 56, pl. 34, figs. 1-3
Caribou Co., Idaho; 5 miles E of Grays Lake
Lower Triassic, Meekoceras zone [holotype USNM 74974]
cumshewaense, Haploceras: Whiteaves Plastoholotype
Whiteaves, 1884, p. 208, pl. 24, fig. 1
Queen Charlotte Islands, Canada; N shore Cumshewa Inlet
Cretaceous, Haida Fm_ [holotype Geol. Surv. Canada 4973]
curticostatum, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 56, pl. 48, figs. 21-22.
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74990]
dalli, Eutomoceras (Halilucites): Smith Plastoholotype
Smith, 1914, p. 65, pl. 29, figs. 1-4
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74314]
damesi, Desmoceras: Jimbo Plastolectotype
Jimbo, 1894, p. 26, pl. 1, fig. 2. Specimen selected as lectotype of
Damesites by Matsumoto, 1954a, p. 267
Hokkaido, Japan; Chiptaushibets, Tumbets River, Kitami Province,
about 68 km from river mouth
Cretaceous [lectotype Kyushu Univ. GT-I-91]
damesii, Ammonites (Acrochordiceras): Noetling Plastoholotype
Noetling, 1880, p. 334, pl. 15, figs. 1a-1c
Silesia, Germany; Gross-Hartmannsdorf (Schlesien)
Triassic, Wellenkalk [holotype in Geol.-Palaont. Mus., Berlin]
declivis, Kingites: Waagen Plastoholotype
Waagen, 1895, p. 233, pl. 26, fig. 2
Punjab, India; Virgal, Salt Range
Triassic, Ceratite Marl [holotype at Paieont. Inst. Wiener Univ.]
delicatum, Asthenoceras: Imlay Paratypes
Imlay, 1973, pp. 55-56, pl. 3, figs. 12 (type 10012), 13 (type 10013),
32 (type 10007)
Grant Co., Ore.; Delintment Lake 15’ Qd, NE 1/4, SW 1/4 SW 1/4
Sec. 29, T 18 S, R 26 E
Middle Jurassic, Bajocian stage, Snowshoe Fm, Warm Springs Mbr
delicatum, Asthenoceras: Imlay Paratypes
Imlay, 1973, pp. 55-56, pl. 3, figs. 15 (type 10010); pl. 4, figs. 4 (type
10009), 3 (type 10011)
Grant Co., Ore.; SE cor. NE 1/4, NE 1/4 Sec. 19, T 18 S, R 26 E,
from spur projecting into SE end of small valley ESE of Weberg
Ranch house
Middle Jurassic, Bajocian stage, Snowshoe Fm, Weberg Mbr (near
top)
delicatum, Asthenoceras: Imlay Paratype
Imlay, 1973, pp. 55-56, pl. 3, fig. 14
Grant Co., Ore.; Delintment Lake 15’ Qd, NW 1/4, SE 1/4 NE 1/4
Sec. 30, T 18 S, R 26 E, from calcareous ss on W slope of hil] 1000’
E of old Washburn place
Middle Jurassic, Bajocian stage, Snowshoe Fm, Warm Springs Mbr,
basal bed
delicatum, Asthenoceras: Imlay Paratype
Imlay, 1973, pp. 55-56, pl. 3, fig. 21
Grant Co., Ore.; Delintment Lake 15’ Qd, SE 1/4, NW 1/4 Sec. 29,
T 18 S, R 26 E. Bulldozer cut on divide SW of jct between road to
Boundary Spring and Suplee-Izee Road
Middle Jurassic, Bajocian, Snowshoe Fm, Warm Springs Mbr, near
base
9079
8726
5413
9853
9852
8963
8781
6475
6500
5872
9050
BuLLeETIN 300
denseplicatum, Lytoceras: Jimbo Plastoholotype
Jimbo, 1894, p. 36, pl. 7, fig. 1
Hokkaido, Japan; Bache Ekimomaanoro
Cretaceous [holotype Kyushu Univ. GT-I-118]
desertorum, Anasibirites: Smith Plastoholotype
Smith, 1932, p. 71, pl. 51, figs. 7, 8
Inyo Co., Calif.; Union Wash, Inyo Range, 15 miles SE of Inde-
pendence
Lower Triassic, Meekoceras zone [holotype USNM 74998]
desertorum, Dinarites: Smith Plastoholotype
Smith, 1914, p. 69, pl. 89, figs. 3, 4
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74425]
desertorum, Metahedenstroemia?: Johnston Paratype
Johnston, 1941, p. 460 (cited as no. 3)
New Pass Range, Nevada; South Canyon
Upper Triassic, Star Peak Fm
devasena, Ceratites: Diener Plastoholotype
Diener, 1907, p. 55, pl. 4, fig. 4 ;
Himalaya Mts.; NNW of Kaga, Spiti
Triassic, Muschelkalk [holotype Paleont. Inst. Wiener Uniy. 4077]
dickinsoni, Leptaleoceras: Imlay Holotype
Imlay, 1968, p. C 32, pl. 6, figs. 7, 9-11
Grant Co., Ore.; Izee Qd, in concretions on E slope Pole Canyon, NW
1/4, SW 1/4 Sec. 35, T 17 S, R 27 E about 75’ above andesite flow
Upper Lower Jurassic, Nicely Shale
dickinsoni, Leptaleoceras: Imlay Paratype
Imlay, 1968, p. C 32, pl. 6, fig. 8
Grant Co., Ore.; Izee Qd, NW 1/4, SW 1/4 Sec. 35, T 17S, R27 E
Upper Lower Jurassic, Nicely Shale
dieneri, Nannites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 79, pl. 7, figs. 10-13
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75257]
dieneri, Ophiceras: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 118, pl. 8, figs. 16-18
SE Idaho; Aspen Mts., Wood Canyon
Lower Triassic, Meekoceras zone [holotype USNM 75260]
discus, Ambites: Waagen Plastosyntype
Waagen, 1895, p. 152, pl. 21, fig. 5
Punjab, India; Salt Range, Amb
Triassic, Ceratite Marls [syntype at Palaeont. Inst. Wiener Univ.]
divergens, Aulacosphinctes: Steiger Plastoholotype
Steiger, 1914, p. 464, pl. 101, figs. 3a-3c
Himalaya Mts.; Shangra, Gnari-Khorsum
Upper Jurassic, upper Malm, Spiti Shale
douvillei, Xenodiscus: Diener Holotype
Diener, 1914, p. 918, pl. 1, fig. 1. Also iz Tozer, 1969, p. 361, pl. 16,
figs. a-d (as Paratirolites)
Madagascar
Lower Triassic
dunni, Eutomoceras: Smith Plastoholotype
Smith, 1904, p. 381, pl. 43, fig. 11; pl. 44, fig. 4
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74310]
8811
9119
8838
9017
6478
8743
9041
9008
6491
6479
8991
8699
6473
9095
STANFORD UNIVERSITY TyPEs: SMITH 381
elkoense, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 56, pl. 55, figs. 14-16
Es Co., Nevada; Ruby Range, Cottonwood Canyon, 70 miles § of
Wells
Lower Triassic, Meekoceras zone [holotype USNM 75015]
emmonsi, Ceratites: Smith Plastoholotype
Smith, 1914, p. 98, pl. 60, figs. 13-15
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74382]
evansi, Meekoceras (Koninckites): Smith Plastoholotype
Smith, 1932, p. 60, pl. 35, figs. 1-3
Idaho; E of Hot Springs, NE of Bear Lake
Lower Triassic, Meekoceras zone [holotype USNM 74975]
evansi, Ptychites: Smith Plastoholotype
Smith, 1914, p. 47, pl. 21, fig. 3
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74295]
falcatum, Meekoceras: Waagen Plastoholotype
Waagen, 1895, p. 242, pl. 36, fig. 4
Punjab, India; Salt Range, Amb
aan Middle Ceratite [holotype Palaeont. Inst. Wiener Univ.
4024
fascicostatum, Pachydiscus: Yabe and Shimizu Plastoholotype
Yabe and Shimizu, 1921, p. 57, pl. 9, fig. 2
Hokkaido, Japan; Abeshinai Valley, Teshio Province
Cretaceous, Santonian, upper Urakawan [cast of holotype GT-I-386
from Kyushu Univ., specimen in Tokyo Univ. ]
fittingensis, Hungarites: Smith Plastoholotype
Smith, 1914, p. 58, pl. 90, figs. 5-7
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74431]
foltzense, Acrochordiceras: Smith Plastoholotype
Smith, 1914, p. 39, pl. 32, figs. 13, 14
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74325]
frequens, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 295, pl. 87, fig. 1
Tibet; Shangra, E of Puling, Gnari-Khorsum
Jurassic, upper Malm, Spiti Shale
frequens, Gyronites: Waagen Plastosyntype
Waagen, 1895, p. 292, pl. 37, fig. 1
Punjab, India; Khoora, Salt Range
Triassic, lower Ceratite [syntype Palaeont. Inst. Wiener Univ. 4037]
gabbi, Celtites: Smith Plastosyntype
Smith, 1914, p. 34, pl. 20, figs. 9, 10
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [syntype USNM 74290]
georgianum, Canadoceras: Anderson Holotype
Anderson, 1958, p. 234, pl. 32, figs. 3, 3a
Straits of Georgia, B.C., Canada; Sucia Islands
Cretaceous
gigas, Koninckites: Waagen Plastoholotype
Waagen, 1895, p. 266, pl. 31, fig. 2
Punjab, India; Salt Range, Choa
Triassic, Ceratite [holotype Palaeont. Inst. Wiener Univ. 4044]
gilberti, Ceratites: Smith Plastoholotype
Smith, 1914, p. 84, pl. 98, figs. 1-3
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74353]
8763
8803
524
525
6489
9111
7609
560
8491
8891
9015
5415
9010
9010a
BuLLeETIN 300
gilberti, Xenodiscus: Smith Plastoholotype
Smith, 1932, p. 43, pl. 24, figs. 1-3
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74923]
gracilis, Flemingites russelli: Smith Plastoholotype
Smith, 1932, p. 53-54, pl. 23, figs. 1-3
SE Idaho; Slug Creek, Aspen Mts., 14 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74918]
grandior, Aturia angustata: Schenck Holotype
Schenck, 1931, p. 462, pls. 73, 74
Wash.; bluffs on Vance’s Creek, 2.5 miles above jct. with Skokomish
River, 13 miles above Union Canyon NP loc. 207
Middle Oligocene
grandior, Aturia angustata: Schenck Paratype
Schenck, 1931, p. 462, pls. 75, 76
Wash.; Port Townsend NP loc. 125
Middle Oligocene
greppini, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 154 :
Trimbach, Switzerland; between Olten and Hauenstein tunnel
Jurassic, Callovian
groteanus, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 283, pl. 80, fig. 4
Tibet; Spiti Province
Jurassic, upper Malm, Spiti Shale
haguei, Ceratites: Smith Plastoholotype
Smith, 1914, p. 97, pl. 42, figs. 1, 2
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74347]
halli, Ammonites: Meek and Hayden Plastoholotype
Meek and Hayden, 1856, p. 70. Illustrated iz Meek, 1876, p. 458, pl.
24, figs. 3a-3c
Montana; Missouri River, 150’ above mouth of Milk River
Cretaceous, Bearpaw Shale [holotype USNM 384]
hallidayi, Nautilus: Waring Holotype
Waring, 1914, p. 783. Illustrated in Waring, 1917, pl. 13, fig. 13
Ventura Co., Calif.; Simi Hills
Eocene, Martinez Fm
haradai, Pachydiscus: Jimbo Plastoholotype
Jimbo, 1894, p. 29, pl. 2, fig. 2
Hokkaido, Japan; Abeshinai, Teshio Province
Cretaceous, Teshio Fm [holotype Kyushu Univ. GT-I-100]
harti, Tirolites: Smith Plastoholotype
Smith, 1932, p. 83, pl. 57, figs. 9, 10
Bear Lake Co., Idaho; Paris Canyon, 1.5 miles W of Paris
Lower Triassic, Columbites zone [holotype USNM 75022]
hartzelli, Arcestes (Proarcestes): Smith Plastoholotype
Smith, 1914, p. 43, pl. 93, figs. 17, 18
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74438]
hatschekii, Ceratites (Haydenites): Diener Plastoholotype
Diener, 1907, p. 72, pl. 6, fig. 1
Himalaya Mts.; NNW of Kaga, Spiti
Triassic, Muschelkalk [holotype Palaeont. Inst. Wiener Univ. 4120]
haugi, Popanoceras (Parapopanoceras): Hyatt and Smith
Hyatt and Smith, 1905, p. 71, pl. 76, figs. 1-4 Plastosyntypes
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [syntypes USNM 74280]
5489
8868
5499
8862
9122
9123
9124
9125
9126
8995
8996
6462
6463
8877
5503
10017
7611
8907
8764
9009
STANFORD UNIVERSITY Types: SMITH 383
hauthali, Harpoceras: Burckhardt Plastoholotype
Burckhardt, 1903, p. 16, pl. 1, figs. 18-20
Argentina; Mendoza Province, Cerro Puchén
Jurassic, Lower Dogger
haydeni, Dagnoceras: Smith Plastoholotype
Smith, 1932, p. 66, pl. 29, figs. 1-3
Idaho; E of Hot Springs, NE end of Bear Lake
Lower Triassic, Meekoceras zone [holotype USNM 74948]
hidimba, Ceratites: Diener Plastoholotype
Diener, 1895, p. 13, pl. 3, figs. 1a-1c
Himalaya Mts., Tibet, Tsang Tsok Li
Triassic, Muschelkalk [holotype Palaeont. Inst. Wiener Univ. 4088]
hooveri, Aspidites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 153, pl. 17, figs. 1-3
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75268]
humboldtensis, Ceratites: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 170, pl. 7, figs. 1-13
West Humboldt Range, Nevada; Troy Canyon area
Middle Triassic [syntypes USNM 74375]
humboldtensis, Columbites: Smith Plastosyntypes
Smith, 1914, p. 36, pl. 20, figs. 26-28; pl. 87, figs. 1-3
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [syntypes USNM 74416]
hyatti, Acrochordiceras: Meek Plastosyntypes
Meek, 1877, p. 124, pl. 11, figs. 5 (type 6462), 5a (type 6463)
Nevada; New Pass, Desatoya Mts.
Triassic [syntypes USNM 12514]
hyatti, Hedenstroemia: Smith Plastoholotype
Smith, 1932, p. 78, pl. 27, figs. 13-15
SE Idaho; 5 miles E of Grays Lake
Lower Triassic, Meekoceras zone [holotype USNM 74938]
insignis, Cyclolobus: Diener Plastoholotype
Diener, 1903, p. 164, pl. 6, fig. 5
Himalaya Mts.; Lilang, Spiti
Permian, Kuling Shale [holotype Palaeont. Inst. Wiener Univ.]
intermedia, Fontannesia: Imlay Paratype
Imlay, 1973, pp. 57-58, pl. 4, figs. 8, 9
Grant Co., Ore.; SE cor. NE 1/4, NE 1/4 Sec. 19, T 18 S, R 26 E
Middle Jurassic, Bajocian stage, Snowshoe Fm, Weberg Mbr (near
top)
intermedius, Scaphites conradi: Meek Plastoholotype
Meek, 1876, p. 433, pl. 34, figs. 3a-3c
S. Dakota; Moreau River
Cretaceous, Fox Hills Fm [holotype USNM 408]
intermontanum, Pseudosageceras: Hyatt and Smith
Plastoholotype
Hyatt and Smith, 1905, p. 99, pl. 4, figs. 1-3
Idaho; Aspen Ridge, Wocd Canyon
Lower Triassic, Meekoceras zone [holotype USNM 75251]
intermontanus, Xenodiscus: Smith Plastoholotype
Smith, 1932, p. 44, pl. 24, figs. 10, 11
Idaho; Slug Creek, 14 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74924]
inyoense, Acrochordiceras: Smith Plastoholotype
Smith, 1914, p. 40, pl. 34, figs. 11, 12
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [holotype USNM 74330]
384
9157
9159
9160
9158
9158b
9158c
9158d
8777
8427
7618
9500
9132
8489
8698
9101
8687
6474
BuLLeETIN 300
inyoense, Cravenoceras: Gordon Holotype
Gordon, 1964, p. Al4, pl. 3, figs. 1, 2, text fig. 4£
Inyo Co., Calif.; Cottonwood Mts., near Rest Spring SU loc. 2776
Upper Mississippian, Perdido Fm
inyoense, Cravenoceras: Gordon Paratypes
Gordon, 1964, p. Al4, pl. 2, figs. 5, 6 (type 9159)
Inyo Co., Calif.; Cottonwood Mts., near Rest Spring SU loc. 2776
Upper Mississippian, Perdido Fm
inyoense, Cravenoceras: Gordon Paratypes
Gordon, 1964, p. Al4, pl. 3, figs. 6-9 (type 9158b), 10-13 (type
9158c), 18-20 (type 9158d)
Inyo Co., Calif.; Cottonwood Mts., near Rest Spring SU loc. 2776
Upper Mississippian, Perdido Fm
jacksoni, Meekoceras (Prionolobus): Hyatt and Smith
Plastoholotype
Hyatt and Smith, 1905, p. 151, pl. 62, figs. 11, 12
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 75292]
japonicum, Desmoceras dawsoni: Yabe Plastoholotype
Yabe, 1904, p. 35, pl. 5, figs. 3, 4
Hokkaido, Japan
Cretaceous [holotype Kyushu Univ. GT-I-260]
jugifer, Ammonites: Waagen Plastoholotype
Waagen, 1867, p. 596, pl. 26, figs. 1a, 1b
Schwaben, Germany; Gingen, Vilsthale
Jurassic, Dogger [cast donated by Palaeont. Mus. Wien]
kamadeva, Ceratites: Diener Plastoholotype
Diener, 1895, p. 24, pl. 5, fig. 1
Himalaya Mts.; Shalshal cliff near Rimkin Pairar
Triassic, Muschelkalk [holotype Palaeont. Inst. Wiener Univ. 4082]
karpinskyi, Ceratites: Smith Plastoholotype
Smith, 1914, p. 100, pl. 44, figs. 4-6
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74351]
kawanoi, Desmoceras: Jimbo Plastoholotype
Jimbo, 1894, p. 28, pl. 1, fig. 7
Hokkaido, Japan; Tshashikoto, Ikandai
Cretaceous [holotype Kyushu Univ. GT-I-98]
kernense, Didymoceras: Anderson Holotype
Anderson, 1958, p. 196, pl. 65, figs. 1, 2
Kern Co., Calif.; Honolulu Consolidated Oil Company Well, T 32 S,
R 24 E, depth 2450’
Cretaceous
kingi, Ceratites: Smith Plastoholotype
Smith, 1914, p. 85, pl. 41, figs. 1-3
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74352]
kingi, Gastroplites: McLearn Plastoholotype
McLearn, 1931, p. 5, pl. 1, fig. 9
Alberta, Canada; S side Peace River, just above mouth of Deep Creek
Lower Cretaceous, Peace River Ss [holotype Geol. Surv. Canada
6340]
kingianus, Aspidites: Waagen Plastoholotype
Waagen, 1895, p. 225, pl. 32, fig. 1; pl. 33, fig. 1
Punjab, India; Virgal, Salt Range
Triassic, Ceratite [holotype Palaeont. Inst. Wiener Univ. 4043]
6466
8758
8893
8937
8482
8878
8428
5504
9054
8972
8721
7606
8487
7600
STANFORD UNIVERSITY Types: SMITH 385
kingianus, Sibirites: Waagen Plastosyntype
Waagen, 1895, p. 168, pl. 18, fig. 1
Punjab, India; Chidroo, Salt Range
Triassic, Ceratite [syntype Palaeont. Inst. Wiener Univ. 4064]
knechti, Lecanites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 138, pl. 9, figs. 11-13
Inyo Co., Calif.; Inyo Mts., Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75264]
knighti, Tirolites: Smith Plastoholotype
Smith, 1932, p. 84, pl. 57, figs. 1, 2
Bear Lake Co., Idaho; Paris Canyon, 1.5 miles W of Paris
Lower Triassic, Columbites zone [holotype USNM 75020]
koeneni, Owenites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 83, pl. 10, figs. 1-4
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75261]
kossmati, Canadoceras: Matsumoto Plastoholotype
Matsumoto, 1954a, p. 295, pl. 13, fig. 1
Hokkaido, Japan; N of Chiptauchibets River, Tumbets, Kitami Pro-
vince
Cretaceous [holotype Kyushu Univ. GT-I-381]
kessmati, Hedenstroemia: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 101, pl. 67, figs. 3-7
Idaho; Aspen Ridge, Wood Canyon
Lower Triassic, Meekoceras zone [holotype USNM 75298]
kotoi, Ammonites: Yabe Plastoholotype
Yabe, 1904, p. 26, pl. 6, figs. 3, 4
Hokkaido, Japan
Cretaceous [holotype Kyushu Univ. GT-I-254]
kraffti, Cyclolobus (Krafftoceras): Diener Plastoholotype
Diener, 1903, p. 165, pl. 6, figs. 9a-9c
Himalaya Mts.; Lilang, Spiti
Permian, Kuling Shale [holotype Paleont. Inst. Wiener Univ. ]
lahontanum, Eutomoceras: Smith Plastoholotype
Smith, 1914, p. 63, pl. 28, figs. 8-11
West Humboidt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74313]
lahontanus, Tropigastrites: Smith Plastoholotype
Smith, 1914, p. 28, pl. 19, figs. 14, 15
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74288]
laqueus, Ammonites: Quenstedt Syntype
Quenstedt, 1885, p. 18, pl. 1, fig. 15
Quedlinburg, Germany
Lower Jurassic, Lias
larvaeformis, Scaphites: Meek and Hayden Plastoholotype
Meek and Hayden, 1858, p. 58. Illustrated in Meek, 1876, p. 418, pl. 6,
figs. 6a-6c
S. Dakota; E base of Black Hills
Cretaceous, Carlile Shale (lower part) [holotype USNM 229]
laticarinatus, Damesites: Saito and Matsumoto Plastoholotype
Saito and Matsumoto, 1956, p. 192, text fig. 1
Hokkaido, Japan; Ikushumbets River, Ishikari Province
Cretaceous, Cenomanian [holotype Kyushu Univ. GT-T-3245]
leei, Scaphites: Reeside Plastoholotype
Reeside, 1927b, p. 26, pl. 20, figs. 17-22
Santa Fe Co., New Mexico; 1 mile S of Waldo
Upper Cretaceous, Mancos Shale, uppermost part [holotype USNM
73354]
386
6498
7602
8951
8874
6499
8975
9854
9165
9166
9166a
5873
5874
5487
8773
7899
BuLtetin 300
lemoinei, Perisphinctes (Virgatosphinctes): Uhlig Plastoholotype
Uhlig, 1910, p. 343, pl. 92, fig. 1 (fig. reversed)
Himalaya Mts.; Tibet, Shangra, Gnari-Khorsum
Upper Jurassic, Spiti Shale
levis, Scaphites: Reeside Plastoholotype
Reeside, 1927b, p. 26, pl. 20, figs. 7-12
Park Co., Wyoming; Sec. 25, T 58 N, R 100 W
Upper Cretaceous, Telegraph Creek Fm, Elk Basin Ss Mbr [holotype
USNM 73353]
ligatus, Columbites: Smith Plastoholotype
Smith, 1932, p. 106, pl. 47, figs. 1-3
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74985]
lindgreni, Anasibirites: Smith Plastoholotype
Smith, 1932, p. 73, pl. 53, figs. 13-15
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Owenites subzone [holotype USNM 75008]
aff. lorioli, n. sp., Aulacosphinctes: Steiger Plastoholotype
Steiger, 1914, p. 460, pl. 101, fig. 1 :
Himalaya Mts., Shangra, Gnari-Khorsum
Upper Jurassic, Spiti Shale
louderbacki, Sibyllites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 58, pl. 74, figs. 10-12
West Humboldt Range, Nevada; N of Troy Canyon
Middle Triassic [holotype USNM 74400]
lupheri, Arieticeras: Imlay Paratype
Imlay, 1968, pp. C34-C35, pl. 4, fig. 15
Grant Co., Ore.; Delintment Lake Qd, SW 1/4 Sec. 28, T 18 S, R 26E
Upper Lower Jurassic, Nicely Shale
macallisteri, Anthracoceras: Gordon Holotype
Gordon, 1964, p. A18, pl. 4, figs. 1-3, text fig. 8
Inyo Co., Calif.; Panamint Range, Cottonwood Mts., near Rest Spring
Upper Mississippian, Perdido Fm
macallisteri, Anthracoceras: Gordon Paratypes
Gordon, 1964, p. A18, pl. 4, figs. 7-9 (type 9166)
Inyo Co., Calif.; Panamint Range, Cottonwood Mts., near Rest Spring
Upper Mississippian, Perdido Fm
madagascariensis, Aspidites: Diener Holotype
Diener, 1914, p. 914, pl. 1, fig. 2
Madagascar
Lower Triassic
madagascariensis, Aspidites: Diener Paratype
Diener, 1914, p. 914, pl. 1, fig. 3
Madagascar
Lower Triassic
malarguense, Harpoceras: Burckhardt Plastoholotype
Burckhardt, 1903, p. 12, pl. 1, figs. 9, 10
Argentina; Mendoza Province, Cerro Puchén
Lower Jurassic, Upper Lias
marcoui, Xenodiscus: Hyatt and Smith Plastosyntype
Hyatt and Smith, 1905, p. 116, pl. 7, fig. 26
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [syntype USNM 75258]
marksi, Eutrephoceras: Miller Holotype
Miller, 1947, p. 33, pl. 20, figs. 1, 2
Kern Co., Calif.; Reed Canyon, elev. 2350’, NW cor. Tejon Qd
Eocene, Tejon Fm
9168
9169
7617
8825
7588
6488
5417
7608
8027
8028
8613
9039
5901
5902
7599
STANFORD UNIVERSITY TyrEs: SMITH 387
masoni, Dombarocanites: Gordon Holotype
Gordon, 1964, p. A21, pl. 4, figs. 4-6, text fig. 10a
Inyo Range, Calif.; 2.25 miles N of Cerro Gordo mine
Upper Mississippian, Chainman Fm
masoni, Dombarocanites: Gordon Paratype
Gordon, 1964, p. A21, pl. 4, fig. 14
Inyo Range, Calif.; 2.25 miles N of Cerro Gordo mine
Upper Mississippian, Chainman Fm
mesacanthus, Ammonites: Waagen Plastoholotype
Waagen, 1867, p. 594, pl. 28, figs. 1a, 1b
Schwaben, Germany; Gingen, Vilsthale
Jurassic, Dogger
micromphalus, Meekeceras: Smith Plastoholotype
Smith, 1932, p. 58, pl. 49, figs. 5-8
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74992]
moreauensis, Ammonites: Owen Plastoholotype
Owen, 1852, p. 579, pl. 8, fig. 7
Fox Hills, S. Dakota
Cretaceous, Fox Hills Ss_ [holotype USNM 20244]
morikeanus, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 281, pl. 80, fig. 2
Tibet; Ki, Spiti Province
Jurassic, Spiti Shale
moorei, Ceratites (Hollandites): Diener Plastoholotype
Diener, 1907, p. 65, pl. 8, fig. 1
Himalaya Mts., Muth, Spiti
Triassic, Muschelkalk Fm_ [holotype Palaeont. Inst. Wiener Univ.
4089]
mullananus, Ammonites ?: Meek and Hayden Plastoholotype
Meek and Hayden, 1863, p. 23. Illustrated in Meek, 1876, p. 607, pl.
8, figs. la-1c
Near Fort Benton, Montana; Chippewa Point
Cretaceous, Colorado Shale (upper part) [holotype USNM 1924]
mulleri, Choanoteuthis: Fischer Holotype
Fischer, 1951, p. 387, pl. 1, figs. 1-3; pl. 2, figs. 1, 2
Mineral Co., Nevada; Gabbs Valley Range, 4 miles E of Luning
Hawthorne Qd_ SU loc. 781
Triassic, Gabbs Fm_ [2 sections of a single specimen]
mulleri, Sonneratia: Anderson Paratype
Anderson, 1938, p. 195, pl. 54, fig. 3
Shasta Co., Calif.; Hulen Creek
Late Cretaceous, Horsetown Fm
multicameratus, Xenodiscus: Smith Plastoholotype
Smith, 1914, p. 57, pl. 34, fig. 5
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [holotype USNM 74329]
nalikanta, Meekoceras: Diener Plastosyntype
Diener, 1895, p. 45, pl. 9, figs. 5a, 5b
Himalaya Mts.; Shalshal cliff near Rimkin Paiar
Triassic, Muschelkalk [syntype Palaeont. Inst. Wiener Univ. 4030]
nanda, Meekoceras: Diener Plastoholotype
Diener, 1895, p. 48, pl. 9, fig. 8
Himalaya Mts.; Shalshal cliff near Rimkin Paiar
Triassic, Muschelkalk [holotype Palaeont. Inst. Wiener Univ. 4098]
nanus, Scaphites aquilaensis: Reeside Plastoholotype
Reeside, 1927b, p. 26, pl. 19, figs. 14-19
Park Co., Wyoming; Sec. 25, T 58 N, R 100 W
Upper Cretaceous, Telegraph Creek Fm, Elk Basin Ss Mbr_ [holotype
USNM 73352]
388
7587
8774
8857
8725
5877
5608
5609
5495
8875
7592
9068
5414
8978
8906
9096
BuLLeETIN 300
nebrascensis, Ammonites: Owen Plastosyntype
Owen, 1852, p. 577, pl. 8, figs. 3, 3a
Fox Hills, S. Dakota
Cretaceous, Fox Hills Fm [syntype USNM 20242]
nevadanus, Xenodiscus (Xenaspis): Smith Plastoholotype
Smith, 1932, p. 47, pl. 56, figs. 1, 2
Elko Co., Nevada; 70 miles S of Wells
Lower Triassic, Meekoceras zone [holotype USNM 75019]
newberryi, Meekoceras (Konickites): Smith Plastoholotype
Smith, 1932, p. 62, pl. 53, figs. 1-3
Inyo Co., Calif.; Union Wash, 15 miles SE of Independence
Lower Triassic, Owenites subzone [holotype USNM 75005]
newpassense, Hannaoceras: Johnston Paratype
Johnston, 1941, p. 454 (cited as No. 4)
Lander Co., Nevada; New Pass Range SU loc. 730
Upper Triassic, Star Peak Fm
newsomi, Goniatites: Smith Holotype
Smith, 1903, p. 78, pl. 17, figs. 2-5
Batesville, Arkansas
Lower Carboniferous, Fayetteville Shale
newsomi, Paralegoceras: Smith Holotype
Smith, 1903, p. 101, pl. 12, figs. 4-9
Conway Co., Arkansas; Morrillton, N 1/2 Sec. 17, T 5 N, R 16 W
Lower Carboniferous [5609 is inner whorl of 5608]
noetlingi, Macrocephalites: Burckhardt Plastoholotype
Burckhardt, 1903, p. 31, pl. 3, figs. 5, 6
Chile; Comisaria Lonquimay, Rio Colorado
Jurassic, Callovian
noetlingi, Sibirites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 49, pl. 9, figs. 1-3
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75262]
novimexicanus, Desmoscaphites: Reeside Plastoholotype
Reeside, 1927b, p. 17, pl. 11, figs. 1-4
Santa Fe Co., New Mexico; 1 mile E of head of Canyon del Yeso
tae Cretaceous, Mancos Shale, uppermost part [holotype USNM
73312 -
nudus, Lecanites: Smith Plastoholotype
Smith, 1914, p. 66, pl. 98, figs. 8, 9
West Humbolt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74456]
oberhummeri, Ceratites (Salterites): Diener Plastoholotype
Diener, 1907, p. 70, pl. 5, fig. 1
Himalaya Mts., Muth, Spiti
Triassic, Muschelkalk [holotype Palaeont. Inst. Wiener Univ. 4081]
obliterans, Tropigastrites: Smith Plastoholotype
Smith, 1914, p. 30, pl. 87, figs. 27-32
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74418]
obtusus, Aspenites: Smith Plastoholotype
Smith, 1932, p. 86, pl. 31, figs. 8-10
Idaho; E of Hot Springs, NE end of Bear Lake
Lower Triassic, Meekoceras zone [holotype USNM 74958]
occidentalis, Ceratites: Smith Plastoholotype
Smith, 1914, p. 84, pl. 44, figs. 21, 22
West Humboldt Range, Nevada; Fossil Hill, § fork American Canyon
Middle Triassic [holotype USNM 74352]
8921
9855
7892
7893
7894
7895
6753
8954
8882
8883
8884
8885
8886
8887
8888
8889
8742
9078
5899
5900
STANFORD UNIVERSITY TypEs: SMITH 389
occidentalis, Ussuria: Smith Plastoholotype
Smith, 1932, p. 91, pl. 27, figs. 8-10
Idaho; Wood Canyon, 9 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [holotype USNM 74937]
ochocoense, Protogrammoceras ?: Imlay Paratype
Imlay, 1968, p. C40, pl. 6, fig. 26
Grant Co., Ore.; Izee Qd, concretions on E slope Pole Canyon, NW
1/4, SW 1/4 Sec. 36, T 17 S,R 27 E
Lower Jurassic, Nicely Shale, about 75’ above andesite flow
olssoni, Aturoidea: Miller Holotype
Miller, 1947, p. 73, pl. 51, figs. 1, 2; pl. 52, fig. 1; pl. 53, figs. 3, 4
Peru; 2 miles N, 3 miles E of Punta Parinas
Eocene, Salina Fm
olssoni, Aturoidea: Miller Paratype
Miller, 1947, p. 73, pl. 54, figs. 1, 2
Peru; 2 miles N, 3 miles E of Punta Parinas
Eocene, Salina Fm
olssoni, Aturoidea: Miller Paratypes
Miller, 1947, p. 73, pl. 92, figs. 3-5 (type 7894)
Peru; 2 miles N, 3 miles E of Punta Parinas
Eocene, Salina Fm
ornatum, Yokoyamaoceras: Matsumoto Plastoholotype
Matsumoto, 1956, p. 183, pl. 16, fig. 3
Hokkaido, Japan; Abeshinai Valley, Teshio Province
Upper Cretaceous [holotype Kyushu Univ., GK-H-5210]
ornatus, Columbites: Smith Plastoholotype
Smith, 1932, p. 107, pl. 46, figs. 14, 15
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74984]
oweni, Inyoites: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 134, pl. 6, figs. 1 (type 8882), 3, 4 (type
8883), 6 (type 8884)
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [syntypes USNM 75255]
oweni, Inyoites: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 134, pl. 69, figs. 1 (type 8885), 2, 3 (type
8886), 4-6 (type 8887), 7-9 (type 8888)
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [syntypes USNM 75301]
oweni, Inyoites: Hyatt and Smith Plastosyntype
Hyatt and Smith, 1905, p. 134, pl. 40, figs. 1-8
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [syntype USNM 75280]
Pacifica, Mesopuzosia: Matsumoto Plastoholotype
Matsumoto, 1954b, p. 82, pl. 15, fig. 1
Hokkaido, Japan; Ishikari Province, Shiyubari Valley
Cretaceous, Saku Fm_ [holotype Kyushu Univ. GK-H-1257]
pacificus, Tirolites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 159, pl. 21, figs. 14, 15
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [holotype USNM 74461]
packardi, Tritropidoceras: Schenk Holotype
Schenk, 1935, p. 402, pl. 17, figs. 1, 2, 13
Grant Co., Ore.; 10 miles E of Suplee, 1.5 miles S of Bailey Ranch
Upper Triassic, upper Karnic
packardi, Tritropidoceras: Schenk Paratype
Schenk, 1935, p. 402, pl. 17, figs. 5-12, 14, 15, 16b-16f
Grant Co., Ore.; 10 miles E of Suplee, 1.5 miles § of Bailey Ranch
Upper Triassic, upper Karnic
8956
8783
9044
9069
7601
8718a
8718b
8718¢c
8826
9163
9164
8895
5625
8829
8830
9140
8492
BuLLETIN 300
parisianus, Columbites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 51, pl. 1, figs. 9-11
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 75246]
parvum, Ophiceras: Smith Plastoholotype
Smith, 1932, p. 49, pl. 54, figs. 25-27
Inyo Co., Calif.; Inyo Range, 15 miles SE of Independence, Union
Wash
Lower Triassic, Owenites subzone [holotype USNM 75011]
parvus, Dalmatites: Smith Plastoholotype
Smith, 1914, p. 60, pl. 30, figs. 1, 2
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74317]
parvus, Lecanites: Smith Plastoholotype
Smith, 1914, p. 66, pl. 30, figs. 25, 26
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74320]
parvus, Scaphites leei: Reeside Plastoholotype
Reeside, 1927b, p. 27, pl. 21, figs. 8-14 5
Sandoval Co., New Mexico; 3/4 mile N of Copper City
Upper Cretaceous, Mesaverde Fm (near base) [holotype USNM
73356]
patagiosus, Ammonites: Schluter Plastosyntypes
Schluter, 1867, p. 22, pl. 4, figs. 4, 5
Coesfeld, Germany [type species of Patagiosites Spath, 1953]
Cretaceous [syntypes Geol.-Pal. Inst. Univ. Bonn]
patelliforme, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 58, pl. 28, figs. 21-23
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74945]
paucinodum, Eumorphoceras: Gordon Holotype
Gordon, 1964, p. A17, pl. 2, figs. 7-9, text fig. 7
Inyo Co., Calif.; Panamint Range, Cottonwood Mts., near Rest Spring
Upper Mississippian, Perdido Fm
paucinodum, Eumorphoceras: Gordon Paratype
Gordon, 1964, p. A17
Inyo Co., Calif.; Panamint Range, Cottonwood Mts., near Rest Spring
Upper Mississippian, Perdido Fm
pealei, Tirolites: Smith Plastoholotype
Smith, 1932, p. 84, pl. 57, figs. 5, 6
Bear Lake Co., Idaho; Paris Canyon, 1.5 miles W of Paris
Lower Triassic, Columbites zone [holotype USNM 75021]
perrini, Scaphites: Anderson Holotype
Anderson, 1902, p. 114, pl. 2, figs. 71-73
Jackson Co., Ore.; Phoenix, near Smith’s ranch
Upper Cretaceous, lower Chico Fm
pilatum, Meekoceras: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 144, pl. 63, figs. 3-9
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Meekoceras zone [syntypes USNM 75294]
pilatus, Ceratites: Smith Plastoholotype
Smith, 1914, p. 102, pl. 89, figs. 10-13
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74427]
planulatiforme, Desmoceras: Jimbo Plastoholotype
Jimbo, 1894, p. 27, pl. 1, fig. 4. Also iz Matsumoto, 1954, p. 96, pl. 20,
fig. 1 (as type of Jimboiceras)
Hokkaido, Japan; Obirashibets, Teshio Province
Cretaceous [holotype Kyushu Univ. GT-I-94]
7605
8997
8720
8992
8979
6482
8745
7595
9081
7614
6481
9104
7589
STANFORD UNIVERSITY TyrEs: SMITH 391
plenus, Scaphites nodosus: Meek and Hayden Plastoholotype
Meek and Hayden, 1860, p. 177. Illustrated in Meek, 1876, p. 429, pl.
26, figs. la-1c
Montana; Yellowstone River near Miles City
Cretaceous, Pierre Shale [holotype USNM 364]
plicatulus, Columbites: Smith Plastoholotype
Smith, 1914, p. 37, pl. 20, figs. 15, 16
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74291]
plicatus, Ammonites psilonotus: Quenstedt Holotype
Quenstedt, 1885, p. 14, pl. 1, fig. 9
Nellingen, Germany
Jurassic, Lias
polygyratus, Celtites: Smith Plastoholotype
Smith, 1914, p. 35, pl. 20, figs. 1, 2
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74289]
powelli, Tropigastrites: Smith Plastoholotype
Smith, 1914, p. 31, pl. 97, figs. 1-4
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74451]
psilogyrus, Lecanites: Waagen Plastoholotype
Waagen, 1895, p. 280, pl. 39, fig. 5
Punjab, India; Khoora, Salt Range
Triassic, lower Ceratite Ss [holotype Palaeont. Inst. Wiener Univ.
4006]
pusillus, Menuites: Matsumoto Plastoholotype
Matsumoto, 1955b, p. 165, pl. 32, fig. 1
Hokkaido, Japan; Hidaka Province, Ikandai, Urakawa area
Cretaceous, upper Yezo group
pusillus, Scaphites hippocrepis: Reeside Plastoholotype
Reeside, 1927b, p. 23, pl. 17, figs. 1-5
Park Co., Wyoming; Sec. 25, T 58 N, R 100 W
Upper Cretaceous, Telegraph Creek Fm, Elk Basin Ss Mbr_ [holotype
USNM 73334]
pygmaeus, Dinarites: Smith Plastoholotype
Smith, 1914, p. 70, pl. 89, figs. 8, 9
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74426]
quadrangularis, Scaphites nodosus: Meek Plastoholotype
Meek, 1876, p. 428, pl. 25, figs. 3a-3c
S. Dakota; Cheyenne River
Cretaceous, Pierre Shale [holotype USNM 366]
radiosum, Meekoceras: Waagen Plastoholotype
Waagen, 1895, p. 257, pl. 36, fig. 2
Punjab, India; Chitta-wan, Salt Range
Triassic, lower Ceratite Ss [holotype Palaeont. Inst. Wiener Univ.
4036]
rectangularis, Ceratites: Smith Plastoholotype
Smith, 1914, p. 85, pl. 41, figs. 14, 15
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74345]
reesidei, Scaphites: Wade Plastoholotype
Wade, 1926, p. 183, pl. 61, figs. 3-6
McNairy Co., Tenn.; Coon Creek, Dave Weeks’ farm
Upper Cretaceous, Ripley Fm, Coon Creek tongue [holotype USNM
73112]
6505
511
8700
9034
8746
8984
9034
6480
8944
8800
185
5614
BuLLeTIN 300
regalis, Hoplites: Pavlow Plastosyntypes
Pavlow, 1891, p. 102, pl. 17, figs. 1, 2
Speeton, England
Cretaceous, Neocomian
regiforme, Pinacoceras: Diener Holotype
Diener, 1916, p. 450, pl. 1, figs. 6a, 6b
Siberia; New Siberian Islands, at the head of Balykatch River,
Kotelny Island
Upper Triassic, Noric
rehmanni, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 153, pl. 48, figs. 1a-1c
Baden, Germany; Geisingen, Donaueschingen
Jurassic, Malm, Callovian
rogersi, Sonneratia: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 69. Illustrated in Wiedey, 1929, pl. 1, fig. 7
Alameda Co., Calif.; Tesla Qd, 3/4 mile S of Carnegie, Corral Hollow
Middle Cretaceous, Horsetown Fm
roguense, Cunningtoniceras: Anderson Holotype
Anderson, 1958, p. 246, pl. 15, figs. 1, la .
Jackson Co., Ore.; “Forty nine” mine
Cretaceous
rosenbergi, Gymnites (Anagymnites): Smith Plastoholotype
Smith, 1914, p. 54, pl. 26, figs. 2, 3
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74307]
rotalinoides, Pachydiscus: Yabe Plastolectotype
Matsumoto, 1955b, p. 169, pl. 34, figs. la-1c
Hokkaido, Japan; Urakawa, Hidaka Province
Cretaceous [cast of I.G.P.S. 54438, designated lectotype of Uraka-
wites |
rothpletzi, Tropigastrites: Smith Plastoholotype
Smith, 1914, p. 31, pl. 19, figs. 1-3
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74286]
rotuloides, Ceratites: Smith Plastoholotype
Smith, 1914, p. 80, pl. 47, figs. 1-3
West Humboldt Range, Nevada; N fork Cottonwood Canyon
Middle Triassic [holotype USNM 74356]
rotundatus, Prionolobus: Waagen Plastosyntype
Waagen, 1895, p. 310, pl. 34, fig. 1
Punjab, India; Virgal, Salt Range
Triassic, Ceratite Marl [syntype Paleont. Inst. Wiener Uniy. 4034]
rotundus, Proteusites: Smith Plastoholotype
Smith, 1932, p. 102, pl. 53, figs. 5, 6
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Owenites subzone [holotype USNM 75006]
russelli, Flemingites: Smith Plastoholotype
Smith, 1904, p. 378, pl. 42, fig. 5; pl. 43, figs. 5, 6
Idaho; 9 miles NE of Soda Springs, Wood Canyon
Lower Triassic, Meekoceras zone [holotype USNM 75302]
sanctaemonicae, Placenticeras: Waring Holotype
Waring, 1915, fig. 11. Also in Waring, 1917, p. 70, pl. 9, fig. 21
Los Angeles Co., Calif.; 4 miles NW of Santa Monica
Upper Cretaceous, Chico Fm
sanctijohanis, Eumorphoceras: Wiedey Holotype
Wiedey, 1929a, p. 323, figs. 1-4, 6
Greene Co., Iowa; Bussey’s Coal Bank, NE 1/4 Sec. 30, T 32 N,
R 29 W
Pennsylvanian, Lower Coal Measures
9615
8833
8592
8593
8594
8595
8576
6490
9011
123
5607
99
5621
5622
10028
STANForRD UNIvERsITY TypEs: SMITH 393
sanctijohanis, Eumorphoceras: Wiedey Paratype
Wiedey, 1929a, p. 323, figs. 5, 7
Greene Co., Iowa; Bussey’s Coal Bank, NE 1/4 Sec. 30, T 32 N,
R 29 W
Pennsylvanian, Lower Coal Measures
sanctorum, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 59, pl. 49, figs. 1, 2
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74991]
schencki, Baculites: Matsumoto Paratypes
Matsumoto, 1959a, p. 113, text figs. 22 (type 8593), 25 (type 8592)
Yolo Co., Calif.; Rumsey Hills, Sec. 19, T 12 N, R 3 W_ SU loc. 2004
Upper Cretaceous, Funks Shale
schencki, Baculites: Matsumoto Paratypes
Matsumoto, 1959a, p. 113
Yolo Co., Calif.; Rumsey Hills, Sec. 30, T 12 N, R 3 W_ SU loc. 2001
Upper Cretaceous, Funks Shale
schencki, Baculites: Matsumoto Paratype
Matsumoto, 1959a, p. 113, text fig. 24
Fresno Co., Calif.; Panoche Qd, Sec. 28, T 14 S,R11E SU loc. 3315
Upper Cretaceous, Panoche Fm
schenki, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 286, pl. 81, fig. 4
Tibet; Shangra, E of Puling, Gnari-Khorsum
Jurassic, Spiti Shale
septentrionalis, Megaphyllites: Smith Plastoholotype
Smith, 1914, p. 42, pl. 21, figs. 4, 5
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74296]
shastense, Acanthoceras: Reagan Holotype
Reagan, 1924, p. 179, pl. 18, fig. 1. Also zz Anderson, 1958, p. 242, pl.
20, figs. 1, 2
Shasta Co., Calif.; Cottonwood Creek SU loc. 121
Cretaceous, Horsetown Fm
siebenthali, Pronorites: Smith Holotype
Smith, 1903, p. 47, pl. 11, figs. 5-7
Scott Co., Arkansas; SE 1/4 SE 1/4 Sec. 4, T 1 N, R 28 W
Upper Carboniferous
silviesi, Uptonia: Hertlein Holotype
Hertlein, 1925b, p. 39, pl. 3, figs. 1, 2, 5
Harney Co., Ore.; Sec. 7, T 20 S, R 30 E, 18 miles N of Burns SU
loc. 27
Middle Lower Jurassic, Hardgrave Ss
simondsi, Shumardites: Smith Holotype
Smith, 1903, p. 135, pl. 3, figs. 11-13
Young Co., Texas; Salt Creek, Graham
Pennsylvanian, Cisco Fm
simondsi, Shumardites: Smith Paratype
Smith, 1903, p. 135
Young Co., Texas; Salt Creek, Graham
Pennsylvanian, Cisco Fm
sparsicostatum, Docidoceras: Imlay Holotype
Imlay, 1973, p. 79, pl. 37, figs. 5-7
Grant Co., Ore.; SW 1/4 SW 1/4 Sec. 27, T 18 S, R 26 E. Float along
irrigation ditch 400’ W of Freeman Creek, 1300’ S of Beaver Creek
Middle Jurassic, Bajocian stage, Snowshoe Fm, Weberg Mbr
394
8999
610
8786
8787
8686
6486
8683
7590
7596
8719
8897
8858
6752a
BuLLETIN 300
spencei, Columbites: Smith Plastoholotype
Smith, 1914, p. 36, pl. 70, figs. 1, 2
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 75309]
spencei, Ophiceras: Hyatt and Smith Paratype
Hyatt and Smith, 1905, p. 119
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic
spencei, Ophiceras: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 119, pl. 62, figs. 1-7
Bear Lake Co., Idaho; 1 mile W of Paris
Lower Triassic [syntypes USNM 75291]
spiekeri, Gastroplites: McLearn Plastoholotype
McLearn, 1931, p. 5, pl. 2, fig. 2
Alberta, Canada; Peace River, 8 miles below Cadotte River
Lower Cretaceous, Peace River Ss [holotype Geol. Surv. Canada
6339 |
stanleyi, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 282, pl. 79, fig. 1 ;
Tibet; Laptel, Gnari-Khorsum
Jurassic, upper Malm, Spiti Shale
stantoni, Gastroplites: McLearn Plastoholotype
McLearn, 1931, p. 5, pl. 1, fig. 4
Alberta, Canada; W bank of Peace River, about 15 miles below
Cadotte River mouth
Lower Cretaceous, Peace River Ss [holotype Geol. Surv. Canada
6336]
stantoni, Scaphites ventricosus: Reeside Plastoholotype
Reeside, 1927a, p. 7, pl. 4, figs. 9, 10
Park Co., Montana; Devil’s Slide, Cinnabar Mt.
Upper Cretaceous, Colorado Mt. Shale (upper part) [holotype
USNM 18817]
“stantoni,” Scaphites: Reeside Plastoholotype
Reeside, 1927b, p. 23, pl. 17, figs. 16-21
Fergus Co., Montana; Willow Creek, 6 miles above old Fort Maginnis-
Junction City road
Upper Cretaceous, Eagle Ss_ [holotype USNM 73338]
stansoni, Sonneratia: Anderson Plastoholotype
Anderson, 1902, p. 105, pl. 3, figs. 91-92; pl. 10, fig. 198
Shasta Co., Calif.; near Horsetown
Lower Cretaceous, Horsetown Fm [= type species of Coloboceras
Crickmay, 1927, p. 511: holotype UCMP]
strongi, Danubites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 165, pl. 9, figs. 4-6
Inyo Co., Calif.; Union Wash, Inyo Range
Lower Triassic, Meekoceras zone [holotype USNM 75263]
strongi, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 62, pl. 52, figs. 12-14
Inyo Co., Calif.; Inyo Range, Union Wash, 15 miles SE of Inde-
pendence
Lower Triassic, Owenites subzone [holotype USNM 75003]
subcostatus, Tragodesmoceroides: Matsumoto Plastoholotype
Matsumoto, 1942, p. 25, fig. 1d. Also iz Matsumoto, 1954, p. 263, pl. 4,
fig. 1
Hokkaido, Japan; Abeshinai district, Teshio Province 7
Cretaceous, Turonian (Neogyliakian) Saku Fm_ [holotype Kyushu
Univ. GT-I-3087]
8788
8839
aU
912
8881
7593
9087
6504
5862
390
STANFORD UNIVERSITY Tyres: SMITH 395
subquadratum, Ophiceras: Smith Plastoholotype
Smith, 1932, p. 50, pl. 54, figs. 18-20
Inyo Co., Calif.; Inyo Range, Union Wash, 15 miles SE of Inde-
pendence
Lower Triassic, Owenites subzone [holotype USNM 75010]
superbus, Aspidites: Waagen Plastoholotype
Waagen, 1895, p. 218, pl. 23, fig. 1; pl. 24, fig. 1
Punjab, India; Salt Range, Chidroo
Triassic, upper Ceratite Ss ? [holotype Palaeont. Inst. Wiener Univ.
4042]
sylvanum, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 59, pl. 33, figs. 1-3
Idaho; 5 miles E of Grays Lake
Lower Triassic, Meekoceras zone [holotype USNM 74970]
tarpeyi, Xenodiscus: Smith Plastoholotype
Smith, 1932, p. 45, pl. 25, figs. 4-6
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74929]
tehamaensis, Schloenbachia: Reagan Holotype
Reagan, 1924, p. 182, pl. 19, fig 3
Tehama Co., Calif.; 30 miles W of Red Bluff
Upper Cretaceous, Chico Fm
templetoni, Schloenbachia: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 78. Illustrated iz Wiedey, 1929b, pl. 2,
fig. 4
Alameda Co., Calif.; Tesla Qd, Western Pacific R.R. cut between
Altamont and Greenway
Upper Cretaceous, upper Chico Fm
tenuis, Clypites: Hyatt and Smith Plastosyntype
Hyatt and Smith, 1905, p. 103, pl. 1, figs. 4-6
Idaho; Wood Canyon, 9 miles NE of Soda Springs
Lower Triassic, Meekoceras zone [syntype USNM 75245]
tenuis, Scaphites hippocrepis: Reeside Plastoholotype
Reeside, 1927b, p. 23, pl. 16, figs. 12, 13
Carbon Co., Wyoming; near Mahoney Ranch, Sec. 7, T 26 N, R 88 W
Upper Cretaceous, Steele Shale (1728’ above base) [holotype USNM
73331]
tenuispiralis, Ceratites: Smith Plastoholotype
Smith, 1914, p. 81, pl. 46, figs. 17-19
West Humboldt Range, Nevada; Fossil Hill, S fork American Canyon
Middle Triassic [holotype USNM 74355]
theodorii, Ammonites: Oppel Plastoholotype
Oppel, 1862, p. 280, pl. 83, figs. 2a, 2b
Tibet; Laptel, Gnari-Khorsum
Jurassic, upper Malm, Spiti Shale
tolli, Cladiscites: Diener Holotype
Diener, 1916, p. 455, pl. 1, figs. 1a, 1b
New Siberian Islands; Kotelny Island, at the head of Balykatch River
Upper Triassic, Noric
toulai, Xenodiscus: Smith Plastoholotype
Smith, 1932, p. 45, pl. 53, figs. 9-12
Inyo Co., Calif.; Inyo Range, Union Wash, 15 miles SE of Inde-
pendence
Lower Triassic, Owenites subzone [holotype USNM 75007]
transitionale, Hauvericeras: Waring Holotype
Waring, 1917, p. 69, pl. 9, fig. 15
Ventura Co., Calif.; Chico area in Bells Canyon, N of Simi fault
Upper Cretaceous, Chico Fm
8988
8860
6465
8897
7603
5493
7612
9071
8771
8922
8923
8941
7610
9088
BuLLetTIN 300
trojanus, Tropigastrites: Smith Plastoholotype
Smith, 1914, p. 65, pl. 29, figs. 1-4
West Humboldt Range, Nevada; Fossil Hill
Middle Triassic [holotype USNM 74281]
tuberculatum, Meekoceras: Smith Plastoholotype
Smith, 1932, p. 62, pl. 50, figs. 1-3
Inyo Co., Calif.; Union Wash, 15 miles SE of Independence
wLower Triassic, Meckoceras zone [holotype USNM 74995]
tuberculatus, Prionites: Waagen Plastoholotype
Waagen, 1895, p. 58, pl. 5, fig. 2
Punjab, India; Salt Range, Chidroo
Triassic, Ceratite [holotype Palaeont. Inst. Wiener Univ. 4038]
ursensis, Celtites: Smith Plastoholotype
Smith, 1932, p. 104, pl. 47, figs. 11, 12
Bear Lake Co., Idaho; Paris Canyon, 1 mile W of Paris
Lower Triassic, Columbites zone [holotype USNM 74987]
ventricosus, Scaphites: Meek and Hayden Plastoholotype
Meek and Hayden, 1863, p. 22. Illustrated in Meek, 1876, p. 425, pl. 6,
figs. 7a, 7b j
Montana; Chippewa Point, near Fort Benton
Cretaceous, Colorado Shale (upper part) [holotype USNM 1903]
vergarensis, Macrocephalites: Burckhardt Plastoholotype
Burckhardt, 1903, p. 29, pl. 2, figs. 18-20; pl. 3, fig. 4
Chile; Vergara, Rio Teno
Jurassic, Bathonian
vermiformis, Scaphites: Meek and Hayden Plastoholotype
Meek and Hayden, 1863, p. 22. Illustrated iz Meek, 1876, p. 423, pl.
6, figs. 4a, 4b
Montana; Chippewa Point, near Fort Benton
Cretaceous, Colorado Shale (upper part) [holotype USNM 1902]
vogdesi, Lecanites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 139, pl. 60, figs. 12-15
West Humboldt Range, Nevada; between Troy Canyon and S fork
American Canyon
Middle Triassic [holotype USNM 74385]
waageni, Meekoceras (Prionolobus): Hyatt and Smith
Plastosyntype
Hyatt and Smith, 1905, p. 150, pl. 77, figs. 3-5
Inyo Co., Calif.; Inyo Range, Union Wash, 1.5 mi E of Union Spring
Lower Triassic, Meekoceras zone [syntypes USNM 75278]
waageni, Ussuria: Hyatt and Smith Plastosyntypes
Hyatt and Smith, 1905, p. 90, pl. 66, figs. 4-6 (type 8922); pl. 67, figs.
1, 2 (type 8923)
SE Idaho; Wood Canyon, Aspen Ridge
Lower Triassic, Meekoceras zone [syntype USNM 75297]
walcoftti, Proptychites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 85, pl. 19, figs. 1-3
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 75270]
warreni, Scaphites: Meek and Hayden Plastoholotype
Meek and Hayden, 1860, p. 177. Illustrated iz Meek, 1876, p. 420, pl.
6, fig. 5
S. Dakota; southern base of Black Hills
Cretaceous, Carlisle Shale (upper part) [holotype USNM 225]
weaveri, Ceratites: Smith Plastoholotype
Smith, 1914, p. 82, pl. 98, figs. 4-7
Desatoya Mts., Nevada; New Pass
Middle Triassic [holotype USNM 74455]
10026
5898
5616
5617
59620
5618
9619
8321
9089
8927
7801
119
9043
8429
8939
STANFORD UNiversity Types: SMITH 397
webergi, Pelekodites: Imlay Paratype
Imlay, 1973, pp. 73-74, pl. 34, fig. 19
Grant Co., Ore.; SE 1/4 SW 1/4 Sec. 29, T 18 S, R 26 E, S slope
pyramidal hill directly S of South Ammonite Hill
Middle Jurassic, Bajocian, Snowshoe Fm, Weberg Mbr, 100’ below top
welleri, Gonioloboceras: Smith Lectotype
Smith, 1903, p. 125, pl. 21, figs. 3, 4 (designated lectotype by Elias,
1938, p. 94, 97, pl. 19, figs. 1a, 1b)
Young Co., Texas; W of Marr’s Hill, Graham
Pennsylvanian, Graham I'm, Cisco group
welleri, Gonioloboceras: Smith Paralectotype
Smith, 1903, p. 125, pl. 21, figs. 1, 2 (designated lectotype by Elias,
1938, p. 94, pl. 19, figs. 3a, 3c)
Young Co., Texas; W of Marr’s Hill, Graham
Pennsylvanian, Graham Fm, Cisco group
welleri, Gonioloboceras: Smith Paratypes
Smith, 1903, p. 125, pl. 21, fig. 3 (type 5617)
Young Co., Texas; W of Marr’s Hill, Graham
Pennsylvanian, Graham Fm, Cisco group
welleri, Gonioloboceras: Smith Paratypes
Smith, 1903, p. 125, pl. 20, figs. 9-11 (type 5618). Also zm Elias, 1938,
p. 94, 97, pl. 19, figs. 4a, 4b (type 5618); pl. 19, figs. 2a, 2b (type
5619)
Young Co., Texas; W of Marr’s Hill, Graham
Pennsylvanian, Graham Fm, Cisco group
wilkinsoni, Engonoceras: Packard Holotype
Packard, 1956, pp. 399-401, fig. 1
Wheeler Co., Ore.; Mitchell Qd, SW 1/4, SE 1/4 Sec. 29, T 11 N,
Re22ER
Cretaceous
williamsi, Ceratifes: Smith Plastoholotype
Smith, 1914, p. 82, pl. 47, figs. 11-14
West Humboldt Range, Nevada; N fork Cottonwood Canyon
Middle Triassic [holotype USNM 74358]
woodini, Sturia: Smith Plastoholotype
Smith, 1932, p. 94, pl. 51, figs. 5, 6
Inyo Co., Calif.; Inyo Range, Union Wash
Lower Triassic, Meekoceras zone [holotype USNM 74997]
woodsi, Mortoniceras: Spath Plastoholotype
Spath, 1921, p. 232, pl. 21, fig. 1
Zululand, South Africa; Umkwelane Hill, Umfolozi
Cretaceous [holotype in South African Museum]
wyomingensis, Metoicoceras: Reagan Holotype
Reagan, 1924, p. 181, pl. 19, figs. 1, 2
Big Horn, Wyoming; Salt Creek region
Cretaceous, Colorado Fm
yatesi, Hungarites: Hyatt and Smith Plastoholotype
Hyatt and Smith, 1905, p. 129, pl. 30, figs. 1-4
Inyo Co., Calif.; Inyo Range, Union Wash
Middle Triassic [holotype USNM 74292]
yokoyamai, Gaudryceras: Yabe Plastoholotype
Yabe, 1904, p. 36, pl. 6, fig. 1
Hokkaido, Japan
Cretaceous [holotype Kyushu Univ. GT-I-197]
zitteli, Owenites: Smith Plastoholotype
Smith, 1932, p. 100, pl. 52, figs. 1-3
Inyo Co., Calif.; Inyo Range, Union Wash, 15 miles SE of Inde-
pendence
Lower Triassic, Owenites subzone [holotype USNM 75000]
398
6397
6399
9507
6528
10285
7089
8710
8069
BuLLETIN 300
SCAPHOPODA
hannai, Dentalina: Baker Paratype
Baker, 1925, p. 84
Gulf of California; off South Coronado Island, 10-18 fms
vallicolens, Dentalium: Raymond Paratype
Raymond, 1904, p. 123. Illustrated in Oldroyd, 1927, p. 13, pl. 1, fig. 2
Santa Monica Bay, Calif.; off Redondo, 145 fms
vallicolens, Dentalium: Raymond Paratypes
Raymond, 1904, p. 123
Santa Monica Bay, Calif.; off Redondo, 145 fms
GASTROPODA
ablita, Epiginella: Laseron Paratype
Laseron, 1957, p. 291
Queensland, Australia; Rocky Island
abreojosensis, Melanella (Melanella): Bartsch - Paratypes
Bartsch, 1917, p. 315
Baja California, Mexico; Point Abreojos
acclivicosta, Bellaspira: McLean and Poorman Paratype
McLean and Poorman, 1970, pp. 6-8
Sonora, Mexico; 1 km S of the E point at entrance to Bahia San
Carlos, 27° 56’ N, 111° 03’ W, 15-20 fms, rock and shell substrate
acerva, Uberella: Laws Paratype
Laws, 1933, p. 321
Blue Cliffs, South Canterbury, New Zealand
Lower Miocene, White Rock River Fm
acutapex, “Acmaea”: Berry Holotype
Berry, 1960, p. 117. Illustrated in Keen, 1971, p. 323, fig. 45 (as
Collisella)
Sonora, Mexico; Punta Cholla, W of Puerto Penasco
adelae, Cancellaria reticulata: Pilsbry Paratype
Pilsbry, 1940, p. 54
Little Duck Key, Fla.
adusta, Arena: McLean Paratype
McLean, 1970c, p. 123
Baja California, Mexico; cove adjoining W sides of Isla Partida and
Espiritu Santo Islands, 24° 25’ N, 110° 25’ W. LACM Sta. 66-28
hermit crab specimen from approx. low water line
aedificata, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 90, pl. 17, fig. 11
Contra Costa Co., Calif.; Carquinez Qd, Vine Station UCMP loc.
1421
Eocene, “Domengine”’ Fm _ [holotype UCMP 33988]
agna, Tectonatica: Woodring Paratypes
Woodring, 1957, p. 88
Panama Canal Zone; Panama R.R. cut, 3500’ SE of Gatun Station
Miocene, middle Gatun Fm
agnesae, Micrarionta: Kanakoff Paratypes
Kanakoff, 1950, p. 85
San Clemente Island, Calif.; China Point
Upper Pleistocene?
8612
6206
6226
9987
8075
8076
7908
9994
6260
7134
8568
8348
5145
5124
STANFORD UNIVERSITY Typrs: SMITH 399
aguilerai, Tylostoma: Alencaster de Cserna Paratypes
Alencaster de Cserna, 1956, p. 24
Puebla, Mexico; San Juan Raya
Cretaceous, San Juan Raya Fm
albemarlensis, Bulimulus (Naesiotus): Dall Paratypes
Dall, 1917c, p. 377
Albermarle Island, Galapagos; near Villamil, 2300-3300’, on grass
and bushes
albemarlensis, Drillia: Pilsbry and Vanatta Syntypes
Pilsbry and Vanatta, 1902, p. 558
Albemarle Island, Galapagos; Tagus Cove
albicarinata, Littorina: McLean Paratypes
McLean, 1970, p. 127
Concepcion Bay, Baja California, Mexico; El Requeson, 26° 38’ N,
111° 50’ W
alfi, Helminthoglypta: Taylor Holotype
Taylor, 1954, p. 76, pl. 20, figs. 30-32
San Bernardino Co., Calif.; Barstow Hills, voleanic ash stratum in
NW cor. Rainbow Basin
Upper Miocene, Barstow Fm
alfi, Helminthoglypta: Taylor Paratypes
Taylor, 1954, p. 76
San Bernardino Co., Calif.; Barstow Hills, volcanic ash stratum in
NW cor. Rainbow Basin
Upper Miocene, Barstow Fm
allyni, Ammonitella yatesi: Chace Paratypes
Chace, 1951, p. 122
Fresno Co., Calif.; near Boyden’s Cave, Kings Canyon
allyni, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970a, p. 298
Tres Marias Islands, W Mexico; off Maria Madre Island, 5-10 fms
almo, Strombiformis: Bartsch Paratypes
Bartsch, 1917, p. 342
Off San Pedro, Calif., in deep water
altacorona, Turritella inezana: Loel and Corey Plastoholotype
Loel and Corey, 1932, p. 256, pl. 57, fig. 6. Also in Merriam, 1941,
p. 109, pl. 25, fig. 4
Santa Barbara Co., Calif.; western Santa Ynez Mts. UCMP loc.
A-602
Lower Miocene, Vaqueros Fm _ [holotype UCMP 31676]
altatae, Olivella (Olivella): Burch and Campbell Paratypes
Burch and Campbell, 1963, p. 123
Sinaloa, Mexico; Altata, in sand at low tide
altispira, Neptunea: Gabb Plastoholotype
Gabb, 1866, p. 44, pl. 14, fig. 2. Also zm Stewart, 1927, p. 395, pl. 31,
fig. 6.
Humboldt Co., Calif.; Eagle Prairie
Pliocene [holotype ANSP 4322]
amandusi, Cypraea: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 628, pl. 18, fig. 1; pl. 19, fig. 1
Baja California, Mexico; San Ignacio Arroyo, 8 kms W of San Ignacio
SU loc. 66
Miocene, Isidro Fm
amandusi, Cypraea: Hertlein and Jordan Paratype
Hertlein and Jordan, 1927, p. 628
Baja California, Mexico; San Ignacio Arroyo, 8 kms W of San Ignacio
Miocene, Isidro Fm
10070
8715
10043
7805
8509
8509a
7539
195
196
8704
6563
BuLLeTIN 300
amara, Nicema: Woodring Paratypes
Woodring, 1964, p. 268
Panama; Transisthmian Highway, lat. 9° 21’ + 1100 feet N, long.
79° 49° WSU loc. 2611 = USGS loc. 16912
Middle Miocene, lower Gatun Fm
ame, Dirocerithium: Woodring Paratype
Woodring, 1959, p. 175
Panama Canal Zone; Rio Casaya area USGS loc. 17166
Middle Eocene, Gatuncillo Fm
amictoideum, Cymatium (Gutturnium): Keen Holotype
Keen, 1971, p. 505, fig. 954
Panama Bay, off NW end San Jose Island, 27-55 m
amputatus, Homorus (Subulona): Pilsbry Paratypes
Pilsbry, 1919, p. 118
Medje, Belgian Congo
anactor, Turritella: Berry Holotype
Berry, 1957, p. 78. Ilustrated im Keen, 1971, p. 392, fig. 433
Baja California, Mexico; 12 miles N of San Felipe
anactor, Turritella: Berry _ Paratype
Berry, 1957; p. 78
Baja California, Mexico; 12 miles N of San Felipe
anchuela, Mitrella (Mitrella): Keen Holotype
Keen, 1943, p. 48, pl. 4, fig. 12
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29 S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
andersoni, Lyria: Waring Holotype
Waring, 1917, p. 97, pl. 15, fig. 12
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
andersoni, Lyria: Waring Paratype
Waring, 1917, p. 97, pl. 15, fig. 12
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
andersoniana, Lioplax: Hannibal Holotype
Hannibal, 1912b, p. 196, pl. 8, fig. 33. Also iz Taylor and Smith,
1971, figs. 32, 33 (as Campeloma)
Tesla, Calif.; 1/4 mile above Carnegie Pottery, Corral Hollow
Eocene
andrium, Teinostoma (Aepystoma): Woodring Paratype
Woodring, 1957, p. 70
Panama; highway 1.7 km NW of Sabanita
Miocene, Gatun Fm
anebus, Solariorbis (Haplorbis) hyptius: Woodring Paratype
Woodring, 1957, p. 75
Canal Zone; N end of third locks excavation SU loc. 2654
Miocene, upper Gatun Fm
angelena, Helminthoglypta tudiculata: Berry Paratype
Berry, 1938a, p. 21
Redlands, Calif.; NE side of lower Timoteo Canyon
angelensis, Solenosteira: Carson Holotype
Carson, 1925, p. 32, pl. 1, figs. 3, 5
Los Angeles Co., Calif.; Puente Hills, mouth of Brea Canyon
Lower Pliocene, Fernando Fm
angelica, Acanthina: Oldroyd Holotype
Oldroyd, 1918a, p. 26. Illustrated im Keen, 1971, p. 552, fig. 1082
Gulf of California; Redondo Bay, Angel Island
6140
7964
6142
8530
7079
10296
599
8093
8090
6581
6178
9737
6534
7621
STANFORD UNIVERSITY Types: SMITH 401
angelina, Olivella biplicata: Oldroyd Holotype
Oldroyd, T. S., 1921b, p. 119, pl. 5, fig. 6. Also im Oldroyd, I. S., 1927,
p. 161, pl. 26, figs. 17, 17a
San Pedro, Calif.
angelina, Olivella biplicata: Oldroyd Paratype
Oldroyd, T. S., 1921b, p. 119
San Pedro, Calif.
angigyra, Ashmunella levettei: Pilsbry Paratypes
Pilsbry, 1905, p. 240
Huachuca Mts., Arizona; Ramsey Canyon, near Ft. Huachuca
angosturana, Cancellaria (Hertleinia): Marks Paratype
Marks, 1949, p. 463, pl. 78, fig. 2
Ecuador; Angostura Cave, Santiago River, Esmeraldas Province
Miocene, Angostura Fm
angulata, Ashmunella: Pilsbry Paratypes
Pilsbry, 1905, p. 244
Chiricahua Mts., Arizona; Cave Creek
anitae, Nassarina (Zanassarina): Campbell Holotype
Campbell, 1961b, p. 26, pl. 5, fig. 4. Also iz Keen, 1971, p. 596, fig.
1253
Guaymas, Mexico; off Cabo Haro, 30 fms
applini, Turritella: Hanna Plastoholotype
Hanna, 1927, p. 307, pl. 49, fig. 4. Also in Merriam, 1941, p. 16, fig. 5
(as Turritella uvasana applini)
San Diego Co., Calif.; La Jolla Qd UCMP loc. 3993
Eocene, La Jolla Fm [holotype UCMP 30971]
approximatus, Bulimulus: Dall Paratype
Dall, 1900a, p. 90
Galapagos; Hood Island
apta, Galeodea: Tegland Paratype
Tegland, 1931, p. 415, pl. 64, figs. 1, 2
Wash.; sea cliffs % to 3 miles E of Twin SU loc. NP 122
Oligocene, Twin Rivers Fm
arenaense, Bittium (Lirobittium): Hertlein and Strong Paratypes
Hertlein and Strong, 1951a, p. 107
Gulf of California; Arena Bank, 23° 32’ N, 109° 25’ W, 45 fms
arenensis, Cymatosyrinx: Hertlein and Strong Paratype
Hertlein and Strong, 195la, p. 76
Gulf of California; near Arena Bank, 23° 32’ N, 109° 27’ W, 55 fms
aresta, Margarites (Lirularia): Berry Paratype
Berry, 1941, p. 13
San Pedro, Calif.; upper sands at Hilltop Quarry
Lower Pleistocene, Lomita Fm
argus, Sonorella: Edson Paratype
Edson, 1912, p. 37
Inyo Co., Calif.; Iron Cap Mine, Argus Range
arnaldoi, Epitonium (Epitonium): Tursch and Pierret Holotype
Tursch and Pierret, 1964, p. 36, fig. 4
Rio de Janeiro, Brazil; off Punta de Juatinga, 23° 22’ S, 48° 28’ W,
50 m
arnoldi, Melanella (Balcis): Bartsch Paratype
Bartsch, 1917, p. 322
San Pedro, Calif.; Deadman’s Island, Sand Rock
Pleistocene, San Pedro Fm
arnoldi, Turcicula: Durham Holotype
Durham, 1944, p. 153, pl. 15, fig. 10
Port Townsend, Wash.; Scow Bay, S shore of Mystery Inlet SU Joc.
NP 126
Middle Oligocene, Marrowstone Shale
7622
8515
304
305
10284
6043
8328
8329
BuLtetIn 300
arnoldi, Turcicula: Durham Paratype
Durham, 1944, p. 153
Port Townsend, Wash.; Scow Bay, S shore of Mystery Inlet SU loc.
NP 126
Middle Oligocene, Marrowstone Shale
artia, Pleuroliria: Berry Holotype
Berry, 1957, p. 82. Illustrated zz Keen, 1971, p. 708, fig. 1648 [as
Polystira oxytropis (Sowerby, 1834) ]
Gulf of California; off Angel de la Guarda Island, 67 fms
ashleyi, Cantharus: Carson Holotype
Carson, 1925, p. 31, pl. 1, figs. 6, 7
Los Angeles Co., Calif.; San Fernando, near tunnel
Pliocene, Fernando Fm
ashleyi, Cantharus: Carson Paratype
Carson, 1925, p. 31
Los Angeles Co., Calif.; Camulos sheet, Gavin Canyon
Pliocene, Fernando Fm
ashleyi, Lirofusus: Arnold Paratypes
Arnold, 1908a, p. 372 .
Santa Cruz Co., Calif.; San Lorenzo River, 3 miles above Boulder
Creek
Oligocene, San Lorenzo Fm
ashmuni, Polygyra: Dall Paratypes
Dall, 1897b, p. 342
Bland, New Mexico
aureola, Pyrene: Howard Paratypes
Howard, 1963a, p. 2. [= Pyrene aureomexicana Howard, 1963b]
Sonora, Mexico; Puerto Penasco, Norse Beach
avalonensis, Helix: Hemphill Paratypes
Hemphill, 1911, p. 104
Santa Catalina Island, Calif.
avawatzica, Micrarionta (Eremarionta): Berry Paratypes
Berry, 1930c, p. 190
San Bernardino Co., Calif.; Avawatz Mts., 5 miles S of Cave Spring
avenosooki, Margarites: MacGinitie Paratype
MacGinitie, 1959, p. 77, pl. 1, fig. 8
About 4 miles off Point Barrow, Alaska, 70 fms
azteca, Nerinea: Alencaster de Cserna Paratype
Alencaster de Cserna, 1956, p. 37
Puebla, Mexico; San Juan Raya
Lower Cretaceous, San Juan Raya Fm
badia, Agladrillia: McLean and Poorman Paratype
McLean and Poorman, 1971, p. 94, fig. 11
Galapagos; off § coast Isla Santa Cruz, 0° 47’ S, 90° 21’ W, 170-200 m
bakeri, Gundlachia: Pilsbry in Baker Paratype
Pilsbry im Baker, Fred, 1914, p. 670
Paria, Brazil
baldwini, Pleurotomaria (Entemnotrochus?): Hickman Paratype
Hickman, 1976b, p. 1095-1096
Polk Co., Ore.; Dallas Qd, SW 1/4 Sec. 25, T 7 S, R 6 W, Ellendale
Basalt Quarry SU loc. 3221 = UCMP A4753
Early Eocene, Siletz River volcanics
baldwini, Pleurotomaria (Entemnotrochus?): Hickman Paratype
Hickman, 1976b, p. 1095-1096, pl. 2, figs. 6, 7
Polk Co., Ore.; Dallas Qd, SW 1/4 Sec. 25, T 7 S, R 6 W, Ellendale
Basalt Quarry SU loc. 3221 = UCMP A4753
Early Eocene, Siletz River volcanics
8329a
6592
8341
6154
9923
8664
115
6526
8062
8064
9990
STANFORD UNIVERSITY Tyres: SMITH 403
baldwini, Pleurotomaria (Entemnotrochus?): Hickman Paratype
Hickman, 1976b, p. 1095-1096, pl. 2, fig. 5
Polk Co., Ore.; Dallas Qd, SW 1/4 Sec. 25, T 7 S, R 6 W, Ellendale
Basalt Quarry SU loc. 3221 = UCMP A4753
Early Eocene, Siletz River volcanics
bandera, Persicula: Coan and Roth Paratypes
Coan and Roth, 1965, p. 67
Jalisco, Mexico; Banderas Bay
baratariae, Corambella: Harry Paratypes
Harry, 1953, pp. 1-9
Lower Barataria Bay, La., in oyster beds
barbata, Oreohelix: Pilsbry Paratypes
Pilsbry, 1905, p. 279
Chiricahua Mts., Arizona; Cave Creek Canyon
bassetti, Protorcula: Smith Plastoholotype
Smith, 1927, p. 109, pl. 101, fig. 7
SE Alaska; Gravina Island, N arm Threemile Cove
Upper Triassic [holotype USNM 74197]
baxteriana, Monadenia fidelis: Talmadge Paratype
Talmadge, 1954, p. 52
Curry Co., Ore.; Sisters Rocks
beali, Conus: Carson Holotype
Carson, 1926, p. 49, pl. 1, fig. 2
Orange Co., Calif.; Puente Hills
Pliocene, Fernando Fm
beali, Marginella: McGinty Paratypes
McGinty, 1940, p. 63
Off Lake Worth, Fla., 84 fms
beaui, Tricolia affinis: Robertson Paratypes
Robertson, 1958, p. 265
Barbados; Bathsheba
beebei, Trophon (Boreotrophon): Hertlein and Strong Paratypes
Hertlein and Strong, 1947b, p. 79
Gulf of California; Gorda Banks, S end of the gulf
bellamaris, Neosimnia: Berry Holotype
Berry, 1946b, p. 191, fig. 1, as bella-maris
Off entrance to San Diego Bay, Calif.; 18 fms
bellamaris, Neosimnia: Berry Paratype
Berry, 1946b, p. 191, as bella-maris
Off entrance to San Diego Bay, Calif.; 18 fms
belvederica, Berthelinia (Edenttellina) chloris: Keen and Smith
Paratype
Keen and Smith, 1961, pp. 53-54
Baja California, Mexico; Puerto Ballandra, Candelero Bay, Espiritu
Santo Island, off La Paz
bermudezi, Cyclostremiscus: Aguayo and Borro Paratype
Aguayo and Borro, 1946a, p. 10
Matanzas, Cuba; Barranco E of Rio Canimar
Upper Miocene, Yumuri Fm
bermudezi, Mecoliotia: Clench and Aguayo Paratype
Clench and Aguayo, 1936, p. 92
Matanzas, Cuba; near mouth of Rio Canimar
Upper Miocene [erroneously cited as Pleistocene]
berryi, Cantharus (Gemophos): McLean Paratypes
McLean, 1970a, p. 314
Jalisco, Mexico; Banderas Bay, off La Cruz, in 10-15 fms
404
9752
8622
10204
10205
8681
9749
8656
9174
6243
8358
6517
6045
5157
5893
193
8025
BULLETIN 300
berryi, Homalopoma: McLean Paratypes
McLean, 1964, p. 132
San Pedro, Calif.; on bluff E of 22nd St.
Pleistocene, Timms Point Fm
bicarinata, Clathrodrillia (Carinodrillia): Shasky Holotype
Shasky, 1961, p. 21, pl. 4, fig. 10
Gulf of California; off Isla Espiritu Santo, 45-90 fms
biconica, Comitas (Boreocomitas): Hickman Paratype
Hickman, 1976, p. 44-46, pl. 2, fig. 6
NW Ore. SU loc. Holman 46
Eocene, Cowlitz Fm
biconica, Comitas (Boreocomitas): Hickman Paratype
Hickman, 1976, p. 44-46
NW Ore. SU loc. Holman 46
Eocene, Cowlitz Fm
biconica, Siphonalia declivis: Makiyama Paratypes
Makiyama, 1941, p. 85
Shizuoka Prefecture, Japan; Tonbe, near Kakegawa
Pliocene, Nango Fm :
bicostata, Lirularia: McLean Paratype
McLean, 1954, p. 129
Gulf of California; off N side of Middle Coronado Island, 15 m
bifasciata, Nassa perpinguis: Berry Holotype
Berry, 1908, p. 39
San Pedro, Calif.
billeeana, Scalina: DuShane and Bratcher Paratype
DuShane and Bratcher, 1965, p. 160
Gulf of California; SW end of Cerralbo Island, 8-10’
binneyanum, Glyptostoma pilsbryanum: Berry Paratype
Berry, 1938c, p. 56
Los Angeles Co., Calif.; Dominguez Hills
blakeana, Pyrgulopsis: Taylor Paratypes
Taylor, 1950, p. 30
Imperial Co., Calif.; Salton Sea, shore by Fish Springs
Upper Pleistocene
boninensis, Patella: Pilsbry Paratype
Pilsbry, 1891, p. 79
Bonin Islands, Japan; Ogasawa
bormanni, Epitonium (Nitidoscala) tinctum: Strong Paratypes
Strong, 1941, p. 47
San Diego Co., Calif.; Mission Bay
bosei, Turritela: Hertlein and Jordan Syntype
Hertlein and Jordan, 1927, p. 634, pl. 21, fig. 1
Baja California, Mexico; San Ignacio Arroyo, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
bosei, Turritella: Hertlein and Jordan Syntype
Hertlein and Jordan, 1927, p. 634, pl. 21, fig. 2. Also in Merriam,
1941p. 114, pl. 29, fig. 3 (as Turritella ocoyana bosei)
Baja California, Mexico; San Ignacio Arroyo, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm [middle Miocene, fide Merriam, 1941]
boundeyi, Bathytoma: Waring Holotype
Waring, 1917, p. 81
Ventura Co., Calif.; Simi Hills, Calabasas sheet SU loc. 2695
Eocene, Martinez Fm
brandi, Amnicola: Drake Paratypes
Drake, 1953, p. 27
Chihuahua, Mexico; Las Palomas, Distrito Galeana
510
6188
6190
216
7546
7546a
7546b
116
306
307
8098
8036
5152
STANFORD UNIVERSITY TyYPEs: SMITH 405
branneri, Cerithium: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 70. Illustrated im Wiedey, 1929b, p. 25,
pl. 1, fig. 6. [Renamed Cerithium ? teslaensis Hanna by Hanna, 1924,
p. 162]
Alameda Co., Calif.; 1 mile N 20° W of Tesla and Corral Hollow
Upper Cretaceous, middle Chico Fm
branneri, Drymaeus: Baker Paratypes
Baker, 1914, p. 637
Matto Grosso, Brazil; Madeira-Mamore R.R., 292 km above Porto
Velho
branneri, Odontostomus (Cyclodontina): Dall Paratype
Dall, 1909b, p. 363
Bahia, Brazil; Rio San Francisco, Serra do Mulato
branneri, Searlesia: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 159, pl. 30, figs. 3a, 3b
Vancouver Island, British Columbia, Canada; W of Otter Point, Sooke
SU loc. NP 129
ligocene, Sooke Fm
bravoensis, Turbonilla (Pyrgiscus): Keen Holotype
Keen, 1943, p. 51, pl. 4, fig. 26
Kern Co., Calif.; Caliente Qd in small gully near center SW 1/4
See; 6, 1). 29'S;, R 30°E, SU loc: 2121
Miocene, Temblor Fm, Round Mountain Silt
bravoensis, Turbonilla (Pyrgiscus): Keen Paratype
Keen, 1943, p. 51, pl. 4, fig. 27
Kern Co., Calif.; Caliente Qd, near center SW 1/4 Sec. 6, T 29 S,
R30 E
Miocene, Temblor Fm, Round Mountain Silt
bravoensis, Turbonilla (Pyrgiscus): Keen Paratype
Keen, 1943, p. 51, pl. 4, fig. 20
Kern Co., Calif.; Caliente Qd, near center SW 1/4 Sec. 6, T 29 S,
R30E
Miocene, Temblor Fm, Round Mountain Silt
breaensis, Astrea: Carson Holotype
Carson, 1926, p. 57, pl. 4, figs. 3, 4
Orange Co., Calif.; Puente Hills, at mouth of Brea Canyon
Lower Pliocene, Fernando Fm
breaensis, Cantharus: Carson Holotype
Carson, 1925, p. 31, pl. 1, fig. 2
Los Angeles Co., Calif.; Puente Hills, at mouth of Brea Canyon
Lower Pliocene, Fernando Fm
breaensis, Cantharus: Carson Paratype
Carson, 1925, p. 31
Los Angeles Co., Calif.; Camulos Qd
Lower Pliocene, Fernando Fm
bristolae, Calotrophon: Hertlein and Strong Paratype
Hertlein and Strong, 1951a, p. 87
Gulf of California; Gorda Banks, lat. 23° 01’ N, long. 109° 29’ W,
60 fms
burchi, Calyptraea: Smith and Gordon Paratypes
Smith and Gordon, 1948, p. 227
Monterey Bay, Calif.; off Del Monte, 15 fms
burkhardti, Terebra: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 632, pl. 21, fig. 6
Baja California, Mexico; San Ignacio Arroyo, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
6218
5774a
59774b
9500
9504
266
6569
5320
6446
8627
6272
8343
8344
5853
BULLETIN 300
buttoni, Cypraea undata: Oldroyd Holotype
Oldroyd, 1916, p. 107. [Renamed Palmadusta diluculum virginalis by
Schilder and Schilder, 1938, p. 160]
Fiji Islands
buttoni, Stagnicola proxima: Henderson ex Baker Ms_ Holotype
Henderson, 1934b, pl. 14, fig. 4 center. Described in Baker, 1934b,
p. 18 [as S. palustris buttoni|
Near Salt Lake City, Utah
buttoni, Stagnicola proxima: Henderson ex Baker Ms__ Paratype
Henderson, 1934b, pl. 14, fig. 4 left. Described zz Baker, 1934b, p.
18 [as Stagnicola palustris buttoni]
Near Salt Lake City, Utah
buttoni, Stagnicola proxima: Henderson ex Baker Ms_ Paratype
Henderson, 1934b, pl. 14, fig. 4 right. Described im Baker, 1934b, p.
18 [as Stagnicola palustris buttoni|
Near Salt Lake City, Utah
californiana, Rimula: Berry Holotype
Berry, 1964, p. 147
Santa Catalina Island, Calif.; Long Point, NE bay, 9-25 fms
californianus, Melampus olivaceous: Berry Holotype
Berry, 1964, p. 153
San Diego Co., Calif.; Pacific Beach, N shore of Mission Bay
californica, Aporrhais: Gabb Paratypes
Gabb, 1864, p. 128
Siskiyou Mts., Calif.
Cretaceous
californica, Oreohelix: Berry Paratypes
Berry, 1931c, p. 115
NE San Bernardino Co., Calif.; at 7500’ on W slope of Clark Mt.
californica, Strepsidura: Arnold Paratype
Arnold, 1908a, p. 370
Santa Cruz Co., Calif.; Kings Creek, 1/2 mile above San Lorenzo
River
Oligocene, San Lorenzo Fm
californicum, Sinum: Oldroyd Holotype
Oldroyd, I. S., 1917, p. 13. Also iz Oldroyd, I. S., 1927, p. 130, pl. 92,
figs. 13, 14
San Pedro, Calif.
californicus, Megomphix: Smith Paratypes
Smith, 1960, pp. 1-3
Trinity Co., Calif.; Natural Bridge Cave
californicus, Pleurobranchus: Dall Syntypes
Dall, 1900c, p. 92
San Pedro, Calif.
californicus, Velates: Vokes Plastosyntype
Vokes, 1935, p. 384, pl. 26, figs. 3, 5
Simi Valley, Calif. UCMP loc. 3792
Eocene, lower Llajas Fm_ [syntype UCMP 15482]
californicus, Velates: Vokes Plastosyntype
Vokes, 1935, p. 384, pl. 26, fig. 4
Simi Valley, Calif. UCMP loc. 3792
Eocene, Jower Llajas Fm [syntype UCMP 15483]
californiense, Helisoma tenue: Baker Holotype
Baker, 1934a, p. 140
Santa Clara Co., Calif.; San Jose, Guadalupe Creek
472
Ti91
6593
6042
6217
7965
5826
9716
6258
6270
6196
STANFORD UNIversITy Types: SMITH 407
calli, Valvata: Hannibal Holotype
Hannibal, 1910, p. 107. Illustrated ix Taylor and Smith, 1971, figs. 47,
48, 51, 52
Near Summer Lake, Ore.
Quaternary, upper Lahontan [Pliocene, probably Blancan, fide Taylor
and Smith]
callidina, Monadenia fidelis: Berry Paratype
Berry, 1940a, p. 13
Del Norte Co., Calif.; S side of Klamath River, near mouth
callidinus, Muricanthus: Berry Holotype
Berry, 1958a, p. 84. Illustrated iz Keen, 1971, p. 523, fig. 1000 (left)
Costa Rica; Bahia Culebra
callinepius, Micrarionta (Eremarionta): Berry Paratypes
Berry, 1930b, p. 544
San Diego Co., Calif.; S slope Santa Rosa Mts., E of mouth of Rock-
house Canyon
callista, Thyca (Bessomia): Berry Paratype
Berry, 1959, p. 110
Sonora, Mexico; Bahia San Carlos, near Guaymas, 3-4 fms
calodinota, Mitra (Tiara): Berry Holotype
Berry, 1960, p. 121. Illustrated im Keen, 1971, p. 644, fig. 1435
Gulf of Nicoya, Costa Rica; off Islas Tortugas
campbelli, Trigonostoma: Shasky Holotype
Shasky, 1961, p. 20, pl. 4, fig. 5
Sonora, Mexico; off Cabo Haro, 30-50 fms
capitanea, Hanetia: Berry Holotype
Berry, 1957, p. 80. Illustrated zm Keen, 1971, p. 563, fig. 1118
Baja California, Mexico; about 8 miles N of San Felipe
caribaea, Rissoella (Phycodrosus): Rehder Paratypes
Rehder, 1943, p. 194
Bonefish Key, Fla.
carmelensis, Skenea: Smith and Gordon Paratype
Smith and Gordon, 1948, p. 239
Carmel Bay, Calif.; 25 fms
Carmen, Bulimulus: Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932b, p. 50
Baja California, Mexico; Salinas Bay, Carmen Island
caroli, Opisthosiphon: Aguayo Paratypes
Aguayo, 1932a, p. 94
Cuba; Loma de la Caridad, Holguin, Oriente
casicalva, Cancellaria: Marks Paratype
Marks, 1949, p. 464
Ecuador; near Jerusalém, Guayas Province
Middle Miocene, Daule Fm
castanea, Chilina: Marshall Paratypes
Marshall, 1924, p. 2
Chubut Province, Argentina; Rio Corcavado
castellum, Crucibulum: Berry Holotype
Berry, 1963, p. 143. Illustrated im Keen, 1971, p. 465, fig. 828 (above)
Guerrero, Mexico; off Acapulco, 6-10 fms
catalinensis, Melanella (Balcis): Bartsch Paratype
Bartsch, 1917, p. 329
Off San Pedro, Calif.; in deep water
catalinensis, Odostoma (Chrysallida): Bartsch Paratypes
Bartsch, 1927, p. 17
Santa Catalina Island, Calif.; Isthmus Cove
catalinensis, Selenites duranti: Hemphill in Binney Paratypes
Hemphill iz Binney, 1890, p. 221
Santa Catalina Island, Calif.
408
6452
6453
6454
6929
6929a
10046
10334
10334a
10334b
10334¢
477
6555
5834
6577
7071
6573
BuL.etTIn 300
catalinensis, Trophon: Oldroyd Holotype
Oldroyd, I. S., 1927, p. 69, pl. 34, figs. 1, 2
Off San Pedro, Calif.; 25 fms
catalinensis, Trophon: Oldroyd Paratype
Oldroyd, I. S., 1927, p. 69, pl. 34, fig. 4
Off San Pedro, Calif.; 25 fms
catalinensis, Trophon: Oldroyd Paratype
Oldroyd, I. S., 1927, p. 69, pl. 34, fig. 5
Off San Pedro, Calif.; 25 fms
catalinensis, Trophon: Oldroyd Paratypes
Oldroyd, I. S., 1927, p. 69, pl. 34, fig. 3
Off San Pedro, Calif.; 25 fms
catalinensis, Trophon: Oldroyd Paratypes
Oldroyd, I. S. 1927, p. 69
Off San Pedro, Calif.; 25-30 fms
caulerpae, Mitrella: Keen Paratype
Keen, 1971, p. 590, fig. 1232
Baja California, Mexico; Puerto Ballandra, about 10 miles NE of
La Paz, in sand among holdfasts of the green alga Caulerpa
cavagnaroi, Naesiotus: Smith Paratype
Smith, 1972, pp. 12-17
Galapagos; Isla Santa Cruz, near top of Mt. Crocker CAS loc,
27538
cavagnaroi, Naesiotus: Smith Paratype
Smith, 1972, pp. 12-17
Galapagos; Isla Santa Cruz, 2 miles W of Mt. Crocker on ground
under small trees CAS loc. 43333
cavagnaroi, Naesiotus: Smith Paratype
Smith, 1972, pp. 12-17
Galapagos; Isla Santa Cruz, ca. 7 km NE of Santa Rosa, Scalesia
zone CAS loc. 40303
cavagnaroi, Naesiotus: Smith Paratype
Smith, 1972, pp. 12-17
Galapagos; Isla Santa Cruz, top of Mt. Crocker, 2900’ elev., sedge
fern zone CAS loc. 27537
cerritensis, Ocinebra lurida: Arnold Paratype
Arnold, 1903, p. 258
Los Angeles Co., Calif.; Los Cerritos, Long Beach
Pleistocene, upper San Pedro Fm
chaceana, Monadenia: Berry Paratype
Berry, 1940a, p. 9
Siskiyou Co., Calif.: Badger Mts., W side of Shasta Canyon
chacei, Goniobasis: Henderson Paratypes
Henderson, 1935, p. 2
Del Norte Co., Calif.; near Crescent City
chacei, Moniliopsis: Berry Paratype
Berry, 1941, p. 6
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene, Lomita Fm
chaneyi, Turritella: Merriam Plastoholotype
Merriam, 1941, p. 71, pl. 6, fig. 8
Santa Clara Co., Calif.; Pacheco Pass region UCMP loc. 10043
Upper Cretaceous, upper Moreno Fm_ [holotype UCMP 33954]
charybdis, Verticumbo: Berry Holotype
Berry, 1940b, p. 154, pl. 17, figs. 6, 7
San Pedro, Calif.; alley S of Second St. and E of Pacific St.
Lower Pleistocene, San Pedro Fm
6573a
7086
10065
6166
6155
9523
6152
6194
197
5948
5018
7219
9724
9725
8354
8354a
STANFORD UNIversItTy TyprEs: SMITH 409
charybdis, Verticumbo: Berry Paratype
Berry, 1940b, p. 154
San Pedro, Calif.; alley S of Second St. and E of Pacific St.
Lower Pleistocene, San Pedro Fm
chehalisensis, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 94, pl. 16, fig. 13
Grays Harbor Co., Wash.; near Balch UCMP loc. 7170
Eocene, Cowlitz Fm [holotype UCMP 33891]
cheloma, Cymia (Cymia): Woodring Paratype
Woodring, 1959, pp. 223-224
Panama; N side Transisthmian Highway, knoll ca. 30 m N of high-
way, 1.2 km NW of Sabanita SU loc. 2611 = USGS loc. 16912
Middle Miocene, lower Gatun Fm
chiricahuana, Holospira: Pilsbry Paratypes
Pilsbry, 1905, p. 219
Fort Bowie, Arizona; Chiricahua Mts.
chiricahuana, Oreohelix (Radiocentrum): Pilsbry Paratypes
Pilsbry, 1905, p. 283
Chiricahua Mts., Arizona; Cave Creek Canyon
civitella, Odostomia (Evalea): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 32
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
clappi, Oreohelix: Ferriss Paratypes
Ferriss, 1904, p. 53
Chiricahua Mts., Arizona; Cave Creek Canyon
clappi, Punctum: Pilsbry Paratypes
Pilsbry, 1898, p. 133
Seattle, Wash.
clarki, Ancistrolepis: Tegland Paratype
Tegland, 1933, p. 3, pl. 12, fig. 17
Twin, Wash.; sea cliffs W of Twin River, for a distance of 3/4 mile
SU loc. NP 120
Oligocene, Twin River Fm
clarki, Epitonium: Oldroyd Holotype
Oldroyd, T. S., 1921a, p. 115, pl. 5, fig. 13
Los Angeles Co., Calif.; Santa Monica
Pleistocene, upper San Pedro Fm
clarki, Epitonium: Oldroyd Paratype
Oldroyd, T. S., 1921a, pv. 115
Los Angeles Co., Calif.; Santa Monica
Pleistocene, upper San Pedro Fm
clarki, Turritella: Dickerson Plastoholotype
Dickerson, 1914, p. 142, pl. 13, fig. 8. Also iz Merriam, 1941, p. 128,
pl. 39, fig. 6 (as Mesalia)
Contra Costa Co., Calif.; Stewartville UCMP loc. 1540
“Eocene,” Martinez Fm _ [holotype UCMP 11936]
clarki, Typhis (Typhisopsis): Keen and Campbell Holotype
Keen and Campbell, 1964, p. 48, figs. 15, 19. Also im Keen, 1971, p. 540,
fig. 1050
Panama Bay; Venado Island, intertidally at -3.0’ tide
clarki, Typhis (Typhisopsis): Keen and Campbell Paratype
Keen and Campbell, 1964, p. 48, fig. 23
Panama Bay; Venado Island, intertidally at -3.0’ tide
clarkiana, Bathytoma: Rivers Plastosyntypes
Rivers, 1913, p. 29, illust. opp. p. 29
San Pedro, Calif.
Upper Pleistocene
410
8257
10299
Pe
137
5831
5806
5807
3829
400
7538
9715
6245
8347
BuLLeETIN 300
clavella, Balcis (Balcis): Berry Paratype
Berry, 1954b, p. 259
Santa Monica, Calif; Long Wharf Canyon
Upper Pleistocene
clementina, Pupa: Sterki Syntype
Sterki, 1890, p. 44
San Clemente Island, Calif.
cocosensis, Vertigo: Dall Paratype
Dall, 1900a, p. 98
Cocos Island, Costa Rica
coei, Crepidula: Berry Holotype
Berry, 1950, p. 35. Illustrated iz McLean, 1969, pp. 35-36, fig. 18.3
Orange Co., Calif.; SE of Seal Beach
collisella, Turbonilla (Pyrgolampros): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 25
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
collomi, Thais (Nucella): Carson Holotype
Carson, 1926, p. 57, pl. 4, fig. 2 :
Santa Barbara Co., Calif.; 1/2 mile N of Schuman in R.R. cut, Santa
Maria district
Lower Pliocene, Fernando Fm
collomi, Thais (Nucella): Carson Paratype
Carson, 1926, p. 57, pl. 4, fig. 1
Santa Barbara Co., Calif.; 1/2 mile N of Schuman in R.R. cut, Santa
Maria district
Lower Pliocene, Fernando Fm
columbiana, Fluminicola: Pilsbry Paratypes
Pilsbry, 1899a, p. 125. [species attributed to Hemphill by some authors,
but Pilsbry is correct]
Columbia River, near Wallula, Wash.
columbiana, Physa: Hemphill Syntypes
Hemphill, 1890, p. 27
Astoria, Ore.; Columbia River
compacta, Cochliopa: Pilsbry Paratypes
Pilsbry, 1910, p. 99
San Luis Potosi, Mexico; Choy River at cave 3 miles S of Las Palmas
compressus, Gyrodes: Waring Holotype
Waring, 1917, p. 67, pl. 9, fig. 6
Calabasas sheet, Calif.; near Ventura-Los Angeles Co. line, N of Simi
fault
Upper Cretaceous, Chico Fm
conchita, Balcis: Keen Holotype
Keen, 1943, p. 43, pl. 4, fig. 5
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4 Sec.
6, T 29 S,R 30 E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
concreta, “Acmaea”: Berry Holotype
Berry, 1963, p. 142. [= Collisella stanfordiana (Berry, 1957), fide
Keen, 1971, p. 325]
Baja California, Mexico; Punta San Felipe
consors, Helminthoglypta dupetithouarsi: Berry Paratype
Berry, 1938a, p. 18
Monterey Co., Calif.; S slope San Juan grade, 8 miles NE of Salinas
constantiae, Diodora: Kanakoff Paratypes
Kanakoff, 1953, pp. 67-70
Wilmington, Calif.; E bank Bermont Ave., 450’ S of SE cor. Sepulveda
Blvd.
Upper Pleistocene, Palos Verdes Sand
644
5815
5814
426
5830
464
247
8645
9744
9830
102838
5824
6515
STANFORD UNIVERSITY [TyPEs: SMITH 411
contignata, Ficus (Trophosycon) ocoyana: Grant and Gale
Paratype
Grant and Gale, 1931, p. 749, pl. 30, fig. 1
“Middle California” [central Calif., perhaps vicinity of Coalinga]
Lower Pliocene, Jacalitos Fm
cooperi, Lymnaea: Hannibal Holotype
Hannibal, 1912b, p. 143, pl. 6, fig. 13a. Also im Taylor and Smith,
1971, p. 312, figs. 36, 37 (as Fossaria)
Santa Cruz Mts., Calif.; spring at Wright’s [NW 1/4 Sec. 23, T 9 S,
R 1 W, in Santa Clara Co., fide Taylor and Smith]
cooperi, Lymnaea: Hannibal Paratype
Hannibal, 1912b, p. 143, pl. 6, fig. 13b. Also ix Taylor and Smith,
1971, p. 312, fig. 40 (as Fossaria)
Santa Cruz Mts., Calif.; spring at Wright’s, Santa Clara Co.
cooperi, Pleurotoma (Dolichotoma): Arnold Plastoholotype
Arnold, 1903, p. 203, pl. 7, fig. 3. Also in Grant and Gale, 1931, p.
499, pl. 25, fig. 3 (as Surculites (Megasurcula) carpenterianus var.
cooperi)
Off San Pedro, Calif.; Deadman Island
Pleistocene, upper San Pedro Fm_ [holotype USNM; plastoholotype
never received at SU]
coquillensis, Goniobasis: Henderson Paratypes
Henderson, 1935, p. 2
Coquille River drainage, Ore.
cordillerana, Heliosoma: Hannibal Holotype
Hannibal, 1912b, p. 161, pl. 6, fig. 16; pl. 8, fig. 34. Also in Taylor
and Smith, 1971, figs. 57, 58, 60, 61 (as Vorticifex)
Nevada; hill near Hawthorne, Belmont stage road
Eocene [late Miocene to early Pliocene, Esmeralda Fm, fide Taylor
and Smith, 1971, p. 313]
cornwalli, Thais: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 162, pl. 31, fig. 1
Vancouver Island, British Columbia, Canada; Jordan River, sea
cliffs at mouth of Fossil Creek, 2 miles W of Sherringham Point
SU loc. NP 130
Oligocene, Sooke Fm
coronadoensis, Macrarene: Stohler Paratype
Stohler, 1959, p. 439
Gulf of California: Coronado Islands, North Island, 150’
cortezi, Crassispira (Striospira): Shasky and Campbell Holotype
Shasky and Campbell, 1964, p. 119, pl. 22, fig. 16
Sonora, Mexico; NW of Bahia Saladita, Guaymas, 10-15 m
cortezi, Sinum: Burch and Burch Paratype
Burch and Burch, 1964, pp. 109-110
Off West Mexico; between Mazatlan and Altata, 15 fms. Taken by
shrimp trawlers
corteziana, Tegula (Agathistoma): McLean Paratypes
McLean, 1970c, p. 119
Sonora, Mexico; S side Cabo Tepoca, 30° 16’ N, 112° 30’ W, mid
intertidal LACM sta. 67-19
costata, Parapholyx: Stearns Paratypes
Stearns, 1901, p. 291 [species not of Hemphill as cited by some
authors, fide Henderson, 1929, p. 81]
The Dalles, Ore.; Columbia River
cowlitzensis, Turbella: Effinger Paratypes
Effinger, 1938, p. 379
Lewis Co.. Wash.; on Cowlitz River, Sec. 25, T 11 N, R 2 W
Lower Oligocene, Gries Ranch Fm
412
139
8350
9745
7859
8097
6511
6163
8671
6205
6179
6180
9502
427
9719
6159
8655
6148
BuLLeTINn 300
crassa, Cancellaria: Waring Holotype
Waring, 1917, p. 66, pl. 9, fig. 5. [Renamed Cancellaria simiana by
Hanna, 1924, p. 160]
Near Ventura-Los Angeles Co. line, in Chico area of Bell’s Canyon,
N of Simi fault; Calabasas sheet
Cretaceous, upper Chico Fm
crassa, Cancellaria: Nomland Plastoholotype
Nomland, 1917a, p. 237, pl. 12, figs. 7, 7a
Fresno Co., Calif.; near Coalinga, N bank of Waltham Creek
Middle Pliocene, Etchegoin Fm [holotype UCMP 11098]
crebriforma, Clathurella (Lioglyphostoma): Shasky and Campbell
Holotype
Shasky and Campbell, 1964, p. 119, pl. 22, fig. 20. Also im Keen, 1971,
p. 761, fig. 1843
Sonora, Mexico; NW of Bahia Saladita, Guaymas, 7-10 m
crispatissima, Ocenebra: Berry Holotype
Berry, 1953b, p. 414, pl. 28, fig. 6
Santa Catalina Island, Calif.; off Isthmus Cove, 33 fms
crockeri, Strombinoturris: Hertlein and Strong Paratype
Hertlein and Strong, 1951b, p. 84
Gulf of California; Arena Bank, 33-35 fms
crooki, Molopophorus: Clark Paratype
Clark, 1938, p. 715
Napa Qd, Calif.; Brink Ranch, 2 miles § of Putah Creek
Upper Eocene, Markley Fm
crotalina, Helminthoglypta: Berry Paratypes
Berry, 1928, p. 276
Mojave Desert, Calif.; N end Granite Mts., Sidewinder Mine
cruenta, Tricolia affinis: Robertson Paratype
Robertson, 1958, p. 267
Sao Paulo, Brazil; Bahia de Flamengo, Ubatuba
cucullinus, Bulimulus (Naesiotus): Dall Paratypes
Dall, 1917c, p. 377
Galapagos; Hood Island, 380’ elev., under stones
cuestana, Epiphragmophora dupetithouarsi: Edson Paratypes
Edson, 1912, p. 37
Santa Lucia Mts., Calif.; Cuesta Pass
cunninghamae, Trialatella: Berry Holotype
Berry, 1964, p. 149. Illustrated iz Keen, 1971, p. 529, fig. 1019 (as
A spella)
Sonora, Mexico; Puerto San Carlos, 15-35 fms
curta, Pleurotoma (Borsonia) bartschi: Arnold Plastoholotype
Arnold, 1903, p. 201, pl. 5, fig. 7
Los Angeles Co., Calif.; Deadman Island
Pleistocene, San Pedro Fm [holotype USNM; plastoholotype never
received at SU]
cymatilis, Olivella (Dactylidiella): Berry Holotype
Berry, 1963, p. 146. Illustrated zz Keen, 1971, p. 629, fig. 1388
Baja California, Mexico; Magdalena Bay
dakani, Oreohelix hendersoni: Henderson Paratypes
Henderson, 1913, p. 38
Colorado; 2 miles up Elk Creek from Newcastle
danai, Terebra (Strioterebrum): Berry Holotype
Berry, 1958b, p. 96. Illustrated in McLean, 1969, p. 52, fig. 28.3
San Pedro, Calif.
danielsi, Ashmunella: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915b, p. 34
Socorro Co., New Mexico; Cave Spring Canyon, R 19 W, lat. 33° 27’
7982
9961
8753
9180
9181
7988
8031
8473
5870
5854
10286
10333
7907
5823
10292
STANFORD UNIversIry Tyres: SMITH 413
daulechica, Strombina: Marks Paratypes
Marks, 1951, p. 112
SW Ecuador; Daule Basin, near Jerusalém
Middle Miocene, Daule Fm
decorata, Puncturella: Cowan and McLean Paratype
Cowan and McLean, 1968, p. 105
Off W coast Queen Charlotte Islands, British Columbia; 53° 21.3’ N,
133° 04.1’ W, 193 m
decoris, Phyllonotus peratus: Keen Holotype
Keen, 1960, p. 107, pl. 10, figs. 4, 5
W Mexico coast near the Guatemalan border, 15 fms
delaguerrae, Turritella schencki: Weaver and Kleinpell
Holotype
Weaver and Kleinpell, 1963, p. 184, pl. 23, fig. 5
Santa Barbara Co., Calif.; W of San Marcos Pass
Eocene, “Coldwater” Ss
delaguerrae, Turritella schencki: Weaver and Kleinpell
Paratype
Weaver and Kleinpell, 1963, p. 184, pl. 23, fig. 6
Santa Barbara Co., Calif.; W of San Marcos Pass
Eocene, ‘Coldwater’ Ss
delgada, Fusiturricula: Marks Paratypes
Marks, 1951, p. 127
SW Ecuador; near Las Masas, Progreso Basin
Lower Miocene, Subibaja Fm
delmontensis, Balcis: Smith and Gordon Paratype
Smith and Gordon, 1948, p. 219
Monterey Bay, Calif.; off Del Monte, 10 fms
delorae, Ceratostoma: Hall Holotype
Hall, 1958, p. 57, pl. 10, figs. 1-3. Also im Hall, 1959, p. 430, pl. 63,
figs. 8-10
Alameda Co., Calif.; NW 1/4 Sec. 11, T 5 S, R 1 E, Alameda Creek
SU loc. 3245
Middle Miocene, Oursan Ss
depressa, Polygyra columbiana: Pilsbry and Henderson Holotype
Pilsbry and Henderson, 1936, p. 134, pl. 7, fig. 2
The Dalles, Ore. [retained at Univ. Colorado Museum Zoological
Coll. as holotype 22519 of Polygyra mullani depressa “Hemphill’’]
depressum, Helisoma occidentale: Baker Paratypes
Baker, 1934a, p. 140
Klamath Lake, Ore.
deroyae, Fissurella (Cremides): McLean Paratype
McLean, 1970c, p. 118
Galapagos; Santa Cruz Island, Academy Bay, 0° 45’ S, 90° 20’ W,
on surf exposed rocks at low tide
deroyi, Naesiotus: Smith Paratype
Smith, 1972, pp. 9-12
Galapagos; NW side Isla Santa Cruz, 870’ elev., on thorn bushes
devexa, Episcynia: Keen Holotype
Keen, 1946, p. 9, pl. 1, figs. 1-4
Santa Barbara Co., Calif.; Santa Cruz Island, Scorpion Harbor, 2-3
fms
diagonalis, Parapholyx effusa: Henderson Paratypes
Henderson, 1929, p. 82
Crater Lake, Ore.
diantha, Tricolia: McLean Paratypes
McLean, 1970c, pp. 125-126
Galapagos; Albemarle (Isabela) Island, E of S end, 0° 55’ S, 90° 30’
W, 60 fms, R/V Velero III bottom sample 450 (not live taken)
414
163
0952
9751
9513
6149
7104
9993
458
7534
9204
9205
BuLtetin 300
dickersoni, Elimia: Clark Paratype
Clark, 1938, p. 707
Napa Qd, Calif.; Pleasant Creek, 1-2 miles S of Putah Creek
Upper Eocene, Markley Fm
dickersoni, Sinum: Waring Holotype
Waring, 1917, p. 86, pl. 14, fig. 10
Ventura Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm
diegensis, Clathrodrillia: Oldroyd Paratypes
Oldroyd, T. S., 1921a, p. 115
San Diego Co., Calif.; Pacific Beach
Upper Pleistocene
diegensis, Macrarene: McLean Paratypes
McLean, 1964, p. 131
San Diego Co., Calif.; Sec. 8, T 19S, R 2 W
Pliocene, San Diego Fm
diegensis, Olivella boetica: Oldroyd Holotype
Oldroyd, T. S., 1921b, p. 118, pl. 5, fig. 2
San Diego, Calif.
dineana, Lymnaea: Taylor Paratypes
Taylor, 1957, p. 659, text-fig. 1, figs. 1-3
Navajo Co., Arizona; White Cone Peak, Sec. 12, T 25 N, R 21 E
Middle Pliocene, Bidahochi Fm
directa, Mitra: Berry Holotype
Berry, 1960, p. 120. Illustrated in Keen, 1971, p. 644, fig. 1436 (as
Subcancilla)
Sonora, Mexico; off Cabo Haro, Guaymas, 30-50 fms
dispar, Ashmunella danielsi: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915b, p. 41
Socorro Co., New Mexico; Little Whitewater Canyon, Mogollon Mts.
diversilineata, Turritella: Merriam Plastoholotype
Merriam, J. C., 1897, p. 65. Illustrated iz Clark, 1918, p. 170, pl. 22,
fig. 5. Also in Merriam, C. W., 1941, p. 103, pl. 20, fig. 1
Vancouver Island, British Columbia, Canada; Carmanah Point
Oligocene, Blakeley Fm [Sooke Fm] [holotype UCMP 11224]
dorothyae, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970a, p. 297
Off San Jose Point, Guatemala; on black sands, 7-11 fms
drakei, Pachychilus: Hannibal Holotype
Hannibal, 1912b, p. 183, pl. 8, fig. 26. Also zz Taylor and Smith, 1971,
figs. 41, 42
Wash.; Olequa Creek, below Little Falls
Eocene [late Eocene, Cowlitz Fm, fide Taylor and Smith, 1971, p.
311]
durhami, Ferminoscala: Keen Holotype
Keen, 1943, p. 46, pl. 4, fig. 31
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4 Sec.
6, I 29S, R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
durhami, Trichotropis (?): Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 188, pl. 25, fig. 4
Santa Barbara Co., Calif.; Nojoqui Creek, 1200’ above Gaviota Can-
yon SU loc. 2908
Eocene-Oligocene, Gaviota Fm
durhami, Trichotropis (?): Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 188, pl. 25, fig. 5
Santa Barbara Co., Calif.; Gaviota Pass UCMP loc. B-7001
Eocene-Oligocene, Gaviota Fm
9206
8261
79
80
6203
9936
7540
6200
109
5848
310
311
5909
7857
8599
STANFORD UNIVERSITY TypPEs: SMITH 415
durhami, Trichotropis (?): Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 188, pl. 25, fig. 3
Santa Barbara Co., Calif.; near Las Cruces UCMP loc. B-6999
Eocene-Oligocene, Gaviota Fm
ebriconus, Balcis (Vitreolina): Berry Paratype
Berry, 1954b, p. 265
San Pedro, Calif.; Hilltop Quarry
Pleistocene, Lomita
egberti, Phalium (Bezoardica): Schenck Holotype
Schenck, 1926, p. 80, pl. 13, fig. 7
Port Discovery, Wash.; sea cliffs 1/4 mile N of old Woodman Wharf
SU loc. NP 148
Oligocene, Lincoln Fm ?
egberti, Phalium (Bezoardica): Schenck Paratype
Schenck, 1926, p. 80
Port Discovery, Wash.; sea cliffs 1/4 mile N of old Woodman Wharf
SU loc. NP 148
Oligocene, Lincoln Fm ?
elaeodes, Bulimulus (Naesiotus): Dall Paratypes
Dall, 1917c, p. 376
Galapagos; Albemarle Island, Banks Bay, at 1500-2300’ elev.
eleanorae, Lucapinella: McLean Paratype
McLean, 1967, p. 350
Jalisco, Mexico; off La Cruz, N shore of Banderas Bay, 20° 44’ N,
105° 29’ W, from cobble bottom, 10 fms
electilis, Moniliopsis: Keen Holotype
Keen, 1943, p. 49, pl. 4, fig. 15
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29 S, R30 E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
elegans, Helix intercisa: Hemphill Paratypes
Hemphill, 1891, p. 330
San Clemente Island, Calif.
elodiae, Cancellaria: Carson Holotype
Carson, 1926, p. 49, pl. 1, fig. 1
Santa Barbara Co., Calif.; Fugler’s Point
Lower Pliocene, Fernando Fm
elrodi, Stagnicola: Baker and Henderson Paratypes
Baker and Henderson, 1933, p. 30
Montana; W shore of Flathead Lake
elsmerensis, Cantharus: Carson Holotype
Carson, 1925, p. 32, pl. 1, fig. 4
Ventura Co., Calif.; Holser Canyon, Piru Valley
Lower Pliocene, Fernando Fm
elsmerensis, Cantharus: Carson Paratype
Carson, 1925, p. 32
Ventura Co., Calif.; Elsmere Canyon, near the forks
Lower Pliocene, Fernando Fm
empyrosia, Drillia: Dall Holotype
Dall, 1899a, p. 127. Illustrated iz Dall, 1902, p. 516, pl. 39, fig. 5
San Pedro, Calif.; 20-50 fms
encopendema, Turveria: Berry Holotype
Berry, 1956b, p. 356, fig. 2. Also zz Keen, 1971, p. 451, fig. 762
Sonora, Mexico; Cholla Cove, Bahia de Adair
englerti, Pisania: Hertlein Paratype
Hertlein, 1960, p. 19
Easter Island
416
5512
7133
8701
6738
9732
6144
7074
8061
6739
194
8640
8032
594
595
BULLETIN 300
epiphanea, Turbonilla (Mormula): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 28
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
equistriata, Turritella inezana: Merriam Plastoholotype
Merriam, 1941, p. 109, pl. 25, fig. 10
Ventura Co., Calif.; probably Ojai Valley
Lower Miocene, Vaqueros Fm [holotype UCMP 33985]
eremum, Calliostoma (Leiotrochus): Woodring Paratype
Woodring, 1957, p. 63
Panama Canal Zone; 1 mile N of Gatun Lake SU loc. 2653
Miocene, Gatun Fm
eritrichius, Mesodon (megasoma, subsp.?): Berry Paratypes
Berry, 1939, p. 56
Butte Co., Calif.; Table Bluff Light
erythrostigma, Siphonochelus (Siphonochelus):
Keen and Campbell Holotype
Keen and Campbell, 1964, p. 51, pl. 10, figs. 27, 31, 35
Queensland, Australia; near Brisbane, Moreton Bay,-12 miles off
Moreton Lighthouse, approx. 51 m
esuritor, Ashmunella: Pilsbry Paratypes
Pilsbry, 1905, p. 249
Chiricahua Mts., Arizona
etheringtoni, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 94, pl. 15, fig. 14
Ventura Co., Calif.; Simi Valley UCMP loc. 7003
Eocene, “Domengine Fm” [holotype UCMP 33875]
euglyptus, Cyclostremiscus: Aguayo and Borro Paratype
Aguayo and Borro, 1946a, p. 9
Cuba; Barranco E of Rio Canimar, Matanzas
Upper Miocene, Yumuri Fm
euthales, Mesodon megasoma: Berry Paratype
Berry, 1939, p. 60
Del Norte Co., Calif.; Chaffay Ranch, 7 miles above mouth of
Klamath River
evoluta, Tornatina: Waring Holotype
Waring, 1917, p. 99, pl. 15, fig. 8
Ventura Co., Calif.; McCray Wells
Upper Eocene, Tejon Fm
eyerdami, Beringius: Smith Paratype
Smith, 1959, p. 5
Off Cape Flattery, Wash.; about 40 miles offshore, 100 fms
fackenthallae, Turbonilla (Turbonilla): Smith and Gordon
Paratype
Smith and Gordon, 1948, p. 220
Monterey Bay, Calif.; off Del Monte, 20-30 fms
fax (?), Galeodea: Tegland Paratype
Tegland, 1931, p. 412, pl. 59, fig. 5
Townsend’s Bay, Wash.; sea cliffs between Classen’s wharf and ship
canal estuary SU loc. NP 125
Lower Oligocene
fax (?), Galeodea: Tegland Paratype
Tegland, 1931, p. 412, pl. 59, fig. 4
Townsend’s Bay, Wash.; sea cliffs between Classen’s Wharf and ship
canal estuary SU loc. NP 125
Lower Oligocene
10044
10044a
9726
9726a
9726b
9726c
10289
6199
106
136
6143
8102
6170
8623
8653
5524
5827
STANFORD UNIVERSITY Types: SMITH 417
fayae, Anachis (Costoanachis): Keen Paratype
Keen, 1971, p. 579, fig. 1178
Sonora, Mexico; Guaymas
fayae, Anachis (Costoanachis): Keen Paratypes
Keen, 1971, p. 579
Sonora, Mexico; Guaymas
fayae, Pterotyphis (Tripterotyphis): Keen and Campbell
Paratype
Keen and Campbell, 1964, p. 54, pl. 11, fig. 40. Also im Keen, 1971, p.
542, fig. 1057
Jalisco, Mexico; Barra de Navidad
fayae, Pterotyphis (Tripterotyphis): Keen and Campbell
Paratypes
Keen and Campbell, 1964, p. 54
Jalisco, Mexico; Barra de Navidad
fayae, Pterotyphis (Tripterotyphis): Keen and Campbell
Plastoholotype
Keen and Campbell, 1964, p. 54, pl. 11, fig. 44
Jalisco, Mexico; Barra de Navidad [holotype Santa Barbara Mus.
Nat. Hist. 15999]
felipensis, Tegula (Agathistoma): McLean Paratypes
McLean, 1970c, p. 121
Baja California del Norte, Mexico; Punta San Felipe, 31° 02’ N, 114°
49’ W, among small rocks at low tide
feralis, Helix: Hemphill Paratypes
Hemphill, 1901, p. 121
San Nicholas Island, Calif.
fergusoni, Cancellaria: Carson Holotype
Carson, 1926, p. 53, pl. 1, fig. 8
Ventura Co., Calif.; Barlow’s Ranch
Pliocene, upper San Pedro Fm
fergusoni, Cancellaria: Carson Paratype
Carson, 1926, p. 53, pl. 1, fig. 7
Santa Barbara Co., Calif.; Fugler’s Point
Lower Pliocene, Fernando Fm
ferrissi, Ashmunella: Pilsbry Paratypes
Pilsbry, 1905, p. 247
Chiricahua Mts., Arizona; Cave Creek Canyon
ferrissi, Holospira: Pilsbry Paratypes
Pilsbry, 1905, p. 215
Huachuca Mts., Arizona; Manilla mine
ferrissi, Sonorella: Pilsbry Paratypes
Pilsbry iz Pilsbry and Ferriss, 1915a, p. 368
Dragoon Mts., Arizona
filiareginae, Vexillum regina: Cate Holotype
Cate, J., 1961, p. 80, pl. 18, figs. 6a, 6b; pl. 19, fig. 6; pl. 20, fig. 1
Philippine Islands; Cape Melville, Balabac
fitchi, Terebra (Strioterebrum): Berry Holotype
Berry, 1958a, p. 89. [= Terebra tiarella Deshayes, fide Keen, 1971, p.
684]
Baja California, Mexico; Vahia Santa Maria, Isla Magdalena
fitella, Odostomia (Evalea): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 33
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
flammulina, Chilina: Marshall Paratypes
Marshall, 1924, p. 3
Chubut, Argentina; Rio Fitaleufu, 43° 9’ S, 71° 35’ W
418
8652
8610
7790
5949
6173
6150
6309
6310
7207
9988
7961
6434
8254
10287
6580
BuL.eTINn 300
fletcherae, Olivella: Berry Holotype
Berry, 1958a, p. 85. Il!ustrated in Keen, 1971, p. 628, fig. 1378
Sonora, Mexico; Cholla Cove, Bahia de Adair
floresi, Craginia: Alencaster de Cserna Paratypes
Alencaster de Cserna, 1956, p. 33
Mexico; San Juan Raya
Lower Cretaceous, San Juan Raya Fm
floridanus, Microcochus: Rehder Paratypes
Rehder, 1943, p. 193
Missouri Key, Fla.
fossilis, Conus californicus: Oldroyd Holotype
Oldroyd, T. S., 1921a, p. 116, pl. 5, fig. 9
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, San Pedro Fm
fossor, Holospira ferrissi: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915a, p. 387
Mule Mt., Arizona; 2 miles E of Warren
fragilis, Ashmunella tetrodon: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1917, p. 89
Black Range, New Mexico; Cave Creek, near Hillsboro’
fraseri, Tritonalia: Oldroyd Holotype
Oldroyd, I. S., 1920, p. 135, pl. 4, figs. 1, 2. Also zz Oldroyd, I. S.,
1927, p. 25, pl. 30, figs. 11, 1la
Vancouver Island, British Columbia, Canada; Brandon Island, Depar-
ture Bay, Nanaimo
fraseri, Tritonalia: Oldroyd Paratype
Oldroyd, I. S., 1920, p. 135
Vancouver Island, British Columbia, Canada; Nanaimo, Brandon
Island, Departure Bay
freya, Turritella: Nomland Plastoholotype
Nomland, 1917b, p. 312, pl. 19, fig. 2. Also im Merriam, 1941, p. 124,
pl. 37, fig. 14
Fresno Co., Calif.; NE of Coalinga UCMP loc. 2283
Miocene, Santa Margarita Fm [holotype UCMP 11313]
frisbeyae, Vermicularia: McLean Paratype
McLean, 1970a, p. 311
Colima, Mexico; Manzanillo, 19° 03’ N, 104° 20’ W, off the lighthouse,
30-40 fms
frizzelli, Cancellaria (Bivetiella): Marks Paratype
Marks, 1949, p. 462
Ecuador; near Jerusalém, Guayas Province
Middle Miocene, Daule Fm
fucana, Olivella biplicata: Oldroyd Holotype
Oldroyd, T. S., 1921, p. 118, pl. 5, fig. 4. Also in Oldroyd, I. S., 1927,
pl. 26, figs. 23, 23a
Straits of Juan da Fuca, near Cape Flattery, Wash.
galapagensis, Cypraea (Trivia): Melvill Syntypes
Melvill, 1900, p. 208, text figs.
Galapagos Islands; Albemarle Island
galapagensis, Mirachelus: McLean Paratype
McLean, 1970c, p. 118
Galapagos; Isabela Island, off Canal Bolivar, near Tagus Cove,
0° 16’ S, 91° 22’ W, 40-55 fms
galeana, Mitromorpha: Berry Paratypes
Berry, 1941, p. 12
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene, Lomita Fm
9718
8751
7550
5516
9508
9985
6267
5008
8081
8082
9925
6198
6251
STANFORD UNIVERSITY Tyres: SMITH 419
gatesi, Solenosteira: Berry Holotype
Berry, 1963, p. 144. Illustrated in Keen, 1917, p. 563, fig. 1120, left
Sinaloa, Mexico; NW of Mazatlan, 15 fms
ghanaense, Dendropoma: Keen and Morton Holotype
Keen and Morton, 1960, p. 48, pl. 4, figs. 7, 8
Ghana, West Africa; about 10 miles W of Takoradi
gluma, Volvulella: Keen Holotype
Keen, 1943, p. 54, pl. 4, fig. 10
Kern Co., Calif.; Caliente Qd, Barker’s Ranch, 1000’ S, 600’ W of NE
cor Sec. 5, T 29 S,R29E_ SU loc. 2641
Miocene, Temblor Fm, Round Mountain Silt or uppermost Olcese Sand
gnomon, Hastula: Keen Holotype
Keen, 1943, p. 47, pl. 4, fig. 11
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4 Sec.
6, T 29S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt (lowermost part)
gomphina, Odostomia (Chrysallida): Oldroyd Paratype
Oldroyd, T. S., 1924, p. 29
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
goodmani, “Acmaea”: Berry Holotype
Berry, 1960, p. 117. [= Collisella stanfordiana (Berry, 1957), fide
Keen, 1971, p. 325]
Baja California, Mexico; 1 mile N of Puertecitos
gordanum, Calliostoma: McLean Paratype
McLean, 1970b, p. 422-423
Baja California, Mexico; Gorda Bank, 70 fms
gouldi, Turbonilla (Pyrgolampros): Dall and Bartsch Paratypes
Dall and Bartsch, 1909, p. 66
San Pedro, Calif.
gracilior, Daphnella aspera: Hemphill in Tryon Lectotype
Hemphill iz Tryon, 1884, p. 317, pl. 25, fig. 62. Lectotype designated
by Grant and Gale, 1931, p. 597, pl. 25, fig. 22 [as Mangelia (Mitro-
morpha) gracilior (Hemphill iz Tryon) ]
Monterey, Calif.
gracilis, Decipifus: McLean Holotype
McLean, 1959, p. 10, pl. 4, fig. 1. Also im Keen, 1971, p. 587, fig. 1222
Sonora, Mexico; Bocochibampo Bay, Guaymas
gracilis, Decipifus: McLean Paratype
McLean, 1959, p. 10. Also iz Keen, 1971, p. 587, fig. 1222
Sonora, Mexico; Bocochibampo Bay, Guaymas
gravinaensis, Purpurina: Smith Plastoholotype
Smith, 1927, p. 109, pl. 101, fig. 6
SE Alaska; Gravina Island
Upper Triassic [holotype USNM 74196]
grippi, Epiphragmophora tudiculata: Pilsbry Paratypes
Pilsbry, 1913, p. 49
Santee, Calif.; 18 miles from San Diego
grippi, Leptothyra: Dall Paratype
Dall, 1911, p. 25
San Diego, Calif.; 100-150 fms, in harbor
grippi, Melanella (Balcis): Bartsch Paratypes
Bartsch, 1917, p. 327
San Pedro, Calif.
gruveli, Marginella: Bavay Paratypes
Bavay in Dautzenburg, 1912, p. 24
Angola, West Africa; Bai de Mossamedes, 15-20 m
420
8508
9837
10300
7989
9986
6156
6255
110
135
9995
6157
5846
5847
8034
69
5131
BuLLeETIN 300
guadalupeana, Astraea: Berry Holotype
Berry, 1957, p. 77 [= Astraea (Pomaulax) gibberosa (Dillwyn, 1817),
fide Keen, 1971, p. 355]
Baja California, Mexico; § end Guadalupe Island, 26.5 fms
guadalupensis, Haliotis fulgens: Talmadge Paratype
Talmadge, 1964, p. 375
Baja California, Mexico; Morro Sur, Guadalupe Island
guadelupiana, Epiphragmophora: Dall Paratype ?
Dall, 1900a, p. 101
Mexico; Guadalupe Island
guayasensis, Megasurcula: Marks Paratype
Marks, 1951, p. 132
SW Ecuador; S of Las Masas
Lower Miocene, Subibaja Fm
guttata, Arene: McLean Paratypes
McLean, 1970a, p. 310-311
Galapagos; Santa Cruz Island, Academy Bay, under rocks in tide
pools
hachetana, Oreohelix (Radiocentrum): Pilsbry - Paratypes
Pilsbry, 1915, p. 330
New Mexico; summit of Hacheta Grande Mt.
halia, Melanella (Balcis): Bartsch Paratype
Bartsch, 1917, p. 322
Baja California, Mexico; Point Abreojos
hamlini, Cancellaria: Carson Holotype
Carson, 1926, p. 51, pl. 1, fig. 6
Los Angeles Co., Calif.; Elsmere Canyon
Lower Pliocene, Fernando Fm
hamlini, Cancellaria: Carson Paratype
Carson, 1926, p. 51, pl. 1, fig. 4
Los Angeles Co., Calif.; Elsmere Canyon
Lower Pliocene, Fernando Fm
hancocki, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970a, p. 299
Santa Elena Bay, Ecuador; off La Libertad, 2° 08’ 20” S, 81° 0’ 15”
W, 8-10 fms, on rocks with gorgonids
handi, Oreohelix: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1918. p. 94
Lincoln Co., Nevada; Charleston Mt., 30 miles N of Las Vegas
hannai, Lanx: Walker Holotype
Walker, 1925, p. 6, pl. 3, figs. 1, 3
Shasta Co., Calif.; Baird, McCloud River
hannai, Lanx: Walker Paratypes
Walker, 1925, p. 6
Shasta Co., Calif.; Baird, McCloud River
hannai, Rissoina: Smith and Gordon Paratypes
Smith and Gorden, 1948, p. 226
Carmel Bay, Calif.; 25 fms
hannibali, Acmaea: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 171, pl. 38, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; Port San Juan, sea
cliffs 1/4 mile E of Providence Cove SU loc. NP 133
Oligocene, Sooke Fm, basal ss and cgl
hannibali, Calliostoma: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 608, pl. 21, fig. 9
Baja California, Mexico; San Ignacio Arroyo, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
129
240
241
157
158
6183
5146
5147
5148
5149
5150
5151
7984
6520
113
5813
5378
5139
STANFORD UNIVERSITY TyPEs: SMITH 421
hannibali, Chrysodomus: Hertlein Holotype
Hertlein, 1925b, p. 42, pl. 3, fig. 4
Montesano, Wash.; 8 miles up Sylvia Creek SU loc. 152 = NP 220
Miocene, Montesano Fm
hannibali, Fusinus: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 158, pl. 30, fig. 2
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
hannibali, Fusinus: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 158, pl. 30, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
hannibali, Lyria: Waring Syntype
Waring, 1917, p. 84, pl. 12, fig. 3
Ventura Co., Calif.; Simi Hills, Martinez area, Calabasas sheet
Lower Eocene, Martinez Fm
hannibali, Lyria: Waring Syntype
Waring, 1917, p. 84, pl. 12, fig. 2
Ventura Co., Calif.; Simi Hills, Martinez area, Calabasas sheet
Lower Eocene, Martinez Fm
hapla, Polygyra: Berry Paratypes
Berry, 1933, p. 14
Butte Co., Calif.; Butte Creek Canyon, near Chico
hartmanni, Macron: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 629, pl. 18, fig. 2; pl. 21, fig. 5
Baja California, Mexico; San Ignacio Arroyo, 8 km W of San
ignacio SU loc. 66
Miocene, Isidro Fm
hartmanni, Macron: Hertlein and Jordan Paratypes
Hertlein and Jordan, 1927, p. 269
Baja California, Mexico; San Ignacio Arroyo, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
haughti, Phos: Marks Paratypes
Marks, 1951, p. 114
Ecuador; Daule Basin, near Jerusalém
Middle Miocene, Daule Fm
haullevillei, Clathurella: Dautzenberg Paratypes
Dautzenberg, 1912, p. 14
West Africa; Guinea coast, off wharf at Tamara
hawleyi, Chrysodomus: Carson Holotype
Carson, 1926, p. 55, pl. 2, fig. 3
4 miles W of Santa Barbara, Calif.
Upper Pliocene, San Pedro Fm [Santa Barbara Fm]
heathi, Doryssa: Pilsbry Paratype
Pilsbry im Baker, 1914, p. 653
Rio Jary, Brazil; Sao Antonio do Cachoeira
hecoxi, Fusus: Arnold Paratype
Arnold, 1908a, p. 371
Santa Cruz Co., Calif.; 5.5 miles above town of Boulder Creek
Oligocene, San Lorenzo Fm
heimi, Cymia: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 622, pl. 18, fig. 5
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
422
5140
5141
5142
10047
10297
9835
9899
5856
6253
8033
9996
9708
6141
8047
8092
Bu.tetin 300
heimi, Cymia: Hertlein and Jordan Paratypes
Hertlein and Jordan, 1927, p. 622
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
helenae, Nassarina (Cigclirina): Keen Holotype
Keen, 1971, p. 594, fig. 1247
Sonora, Mexico; off Guaymas, 45 m
helleri, Endodonta: Dall Paratype
Dall, 1900a, p. 93
Galapagos; Isabela Island, Iguana Cove, 2000’ elev.
hemphilli, Goniobasis: Henderson Paratypes
Henderson, 1935, p. 96
Portland, Ore.
hemphilli, Helisoma: Baker and Henderson Holotype
Baker and Henderson im Baker, Frank, 1934a, p. 141
San Francisco, Calif.; Mountain Lake
hemphilli, Helisoma: Baker and Henderson Paratypes
Baker and Henderson im Baker, Frank, 1934a, p. 141 3
San Francisco, Calif.; Mountain Lake
hemphilli, Melanella (Melanelia): Bartsch Paratypes
Bartsch, 1917, p. 313
Baja California, Mexico; Point Abreojos
hemphilli, Stagnicola: Henderson ex Baker Ms Holotype
Henderson, 1934b, pl. 14, fig. 7, right. Described in Baker, 1934b, p. 19
Utah Co., Utah; near Salt Lake City
hemphilli, Stagnicola: Henderson ex Baker Ms Paratype
Henderson, 1934b, pl. 14, fig. 7, left
Utah Co., Utah; near Salt Lake City
hemphilli, Strombiformis: Bartsch Paratypes
Bartsch, 1917, p. 344
Baja California, Mexico; Point Abreojos
hemphilli, Tegula: Oldroyd Paratypes
Oldroyd, T. S., 1921a, p. 115
San Diego, Calif.; Pacific Beach
hendersoni: Polygyra mullani: Pilsbry Paratypes
Pilsbry, 1928, p. 178
The Dalles, Ore.
hertleini, Rissoella: Smith and Gordon Paratype
Smith and Gordon, 1948, p. 225
Monterey Bay, Calif.; off Cabrillo Point, 10 fms
hertleini, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970b, pp. 1-2
Galapagos; Santa Cruz Island, Academy Bay, 3.5-5.5 fms
hesperina: Blasicrura coxeni: Schilder and Summers Paratype
Shilder and Summers, 1963, p. 68
Talesea, New Britain
heterodonta, Ashmunella levettei: Pilsbry Paratypes
Pilsbry, 1905, p. 241
Huachuca Mts., Arizona
hewitti, Ampullella: Hanna and Hertlein Paratypes
Hanna and Hertlein, 1949, p. 393
Kern Co., Calif.; Sec. 18, T 29 S,R20E CAS loc. 32388A
Middle Eocene, Domengine Fm
hilli, Crockerella: Hertlein and Strong Paratype
Hertlein and Strong, 1951a, p. 79
Gulf of California; Santa Inez Bay, 26° 52’ N, 111° 53’ W, 4-13 fms
5511
6262
6263
8256
8063
6216
531
8693
6040
6244
8707
6249
5517
7861
8357
5776
5776a
5776b
5776c
STANFORD UNIVERSITY Types: SMITH 423
himerta, Turbonilla (Pyrgiscus): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 27
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
hipolitensis, Niso: Bartsch Paratypes
Bartsch, 1917, p. 350
San Diego, Calif. (type 6262) ; Baja California, Mexico, San Hipolito
Point (type 6263)
hoffmeyeri, Terebra (Strioterebrum): Abbott Paratypes
Abbott, 1952, p. 78
Philippines; Luzon Island, Pasay Beach, Manila Bay
hoffi, Cyclostremiscus (Bathyspira): Aguayo and Borro
Paratype
Aguayo and Borro, 1946b, p. 44
Cuba; Barranco E of Rio Canimar, Matanzas
Upper Miocene, Yumuri Fm
holguinense, Opisthosiphon aguilerianum: Aguayo Paratypes
Aguayo, 1932a, p. 93
Oriente, Cuba; Cerro San Juan, Sao Arriba, Holguin
hooveri, Mangilia: Arnold Paratype
Arnold, 1903, p. 212
Los Angeles Co., Calif.; San Pedro
Pleistocene, upper San Pedro Fm
howardae, Nassarius: Chace Paratypes
Chace, 1958b, p. 333
Baja California, Mexico; Almejas Beach, 5 miles N of San Felipe
huachucana, “Pyramidula” strigosa: Pilsbry Paratypes
Pilsbry, 1902, p. 511
Huachuca Mts., Arizona
humboldtica, Helminthoglypta arrosa: Berry Paratypes
Berry, 1938a, p. 17
Humboldt Co., Calif.; near Bridge Creek Camp, S of Scotia
hyptius, Solariorbis (Haplorbis) hyptius: Woodring Paratypes
Woodring, 1957, p. 75
Panama Canal Zone; R.R. 3500’ SE of Gatun Station
Miocene, middle Gatun Fm
idae, Mitra: Melvill Paratypes
Melvill, 1893, p. 140
San Diego Co., Calif.; Point Loma
idae, Turbonilla (Pyrgolampros): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 26
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
idahoensis, Lymnaea: Henderson Paratypes
Henderson, 1931, p. 75
Idaho; Little Salmon River, 16 miles N of New Meadows, on rocks
in a mountain stream
imminens, Pyrgulopsis: Taylor Paratypes
Taylor, 1950, p. 28
Imperial Co., Calif.; shore of Salton Sea, by Fish Springs
Upper Pleistocene
impedita, Stagnicola: Henderson ex Baker Ms Holotype
Henderson, 1934b, pl. 14, fig. 3 left. Described in Baker, 1934b, p. 20
Cache Co., Utah; near Logan
impedita, Stagnicola: Henderson ex Baker Ms Paratypes
Henderson, 1934b, pl. 14, fig. 3, right. Also im Baker, 1934b, p. 20
Cache Co., Utah; near Logan
424
6922
5191
9727
9728
8260
9512
6424
6147
7226
7849
7849a
8356
6213
10279
10326
6214
BuL.etIn 300
imperialis, Chrysodomus: Dall Paratype
Dall, 1909a, p. 42, pl. 18, fig. 1
Santa Cruz Qd, Calif.; near headwaters of Alpine Creek Arnold’s
loc. 6 = C-306
Pliocene, Purisima Fm
imperialis, Rapana: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, pp. 631-632, pl. 20, fig. 1
Baja California, Mexico; San Ramon River, La Purisima cliffs
SUiMlocs 57
Lower Miocene, Isidro Fm
imperialis, Typhis (Typhina): Keen and Campbell Paratype
Keen and Campbell, 1964, p. 46, fig. 4
Off Tosa, Japan; approx. 200 m
imperialis, Typhis (Typhina): Keen and Campbell Plastoholotype
Keen and Campbell, 1964, p. 46, figs. 1-3
Off Tosa, Japan; 200 m_ [holotype in Kyoto, Japan, private coll. of
Mr. Akimbumi Teramachi |]
incallida, Balcis (Vitreolina): Berry Paratype
Berry, 1954b, p. 264 :
San Pedro, Calif.; Hilltop quarry
Lower Pleistocene, Lomita
incompta, Coralliophila: Berry Holotype
Berry, 1960, p. 119
Gulf of California; Angel de la Guarda Island, 20 miles off Puerto
Refugio
indisputabilis, Alectrion mendicus: Oldroyd “Holotype”
Oldroyd, I. S., 1927, pl. 26, fig. 4, no description. [= a variant of
Alectrion mendicus, teste Keen, 1976]
San Diego, Calif.
inermis, Ashmunella tetrodon: Pilsbry and Ferriss Paratype
Pilsbry and Ferriss, 1915b, p. 33
Socorro Co., New Mexico; Mogollon Mts., Dry Creek Canyon
infera, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 90, pl. 40, fig. 4
Ventura Co., Calif.; Simi Valley, Las Llajas Canyon UCMP Joc.
A-994
Eocene, Llajas Fm_ [holotype UCMP 33993]
infima, Assiminea: Berry Holotype
Berry, 1947a, p. 5, text fig. 1
Inyo Co., Calif.; Death Valley, Badwater, elev. -279.6’
infima, Assiminea: Berry Paratypes
Berry, 1947a, p. 5
Inyo Co., Calif.; Death Valley, Badwater, elev. -279.6’
infirma, Baroginella: Laseron Paratypes
Laseron, 1957, p. 305
Torres Strait, Australia; Murray Island, 5-8 fms
inglesi, Helminthoglypta: Berry Paratype
Berry, 1938b, p. 43
Kern Co., Calif.; Horse Meadows, trail to Sunday Peak
insalli, Terebra (Triplostephanus): Bratcher and Burch
Paratype
Bratcher and Burch, 1967, p. 7
Red Sea, coast of Israel; Eilat, Gulf of Aqaba (Akabar)
iota, Turritella: Popenoe Plastoholotype
Popenoe, 1937, p. 401, pl. 49, fig. 8
Orange Co., Calif.; Corona sheet CIT loc. 984
Cretaceous, Turonian [holotype UCLA 40673]
isabella, Helminthoglypta: Berry Paratypes
Berry, 1938b, p. 42
Kern Co., Calif.; 2 miles E of Isabella
7542
9521
6044
8106
9997
6568
10294
6552
5809
8502
8503
8255
130
10042
7099
497
STANFORD UNIVERSITY Types: SMITH 425
ischnon, Olivella: Keen Holotype
Keen, 1943, p. 50, pl. 4, figs. 3, 4
Kern Co., Calif.; Caliente Qd, near Barker’s Ranch, SE 1/4 Sec. 5,
T 29 S,R29E_ SU loc. 2641
Miocene, Temblor Fm, basal Round Mountain Silt or uppermost
Olcese Sand
ithea, Odostomia (Evalea): Oldroyd Paratype
Oldroyd, T. S., 1924, p. 31
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
jacksonensis, Lymnaea: Baker Paratypes
Baker, 1907, p. 52
Jackson Lake, Wyoming
jacoquea, Turritella (?): Jenkins Plastosyntypes
Jenkins, 1913, p. 451, pl. 20, fig. 7. Also ix Maury, 1934, p. 150, pl.
14, fig. 4 [as Cerithium (?)]
Brazil; Jacoca, 4 km SW of Ceara-Mirim, Rio Grande do Norte
Eocene? or Upper Cretaceous
jacquelinae, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970b, pp. 2-5
Galapagos; Santa Cruz Island, Academy Bay, ca. 10 fms
jaegeri, Oreohelix handi: Berry Paratypes
Berry, 1931c, p. 118
Charleston Mts., Nevada; ridge W of Griffith’s Hotel, 7500’ elev.
jaliscoensis, Calliclava: McLean and Poorman Paratype
McLean and Poorman, 1917, p. 90
Jalisco, Mexico; Tenacatita Bay, 19° 17’ N, 104° 50’ W, 20-40 fms
janesburgensis, Turricula: Stanton Plastoholotype
Stanton, 1920, p. 45, pl. 9, figs. 2a, 2b
North Dakota; Cannonball River near Janesburg
Cretaceous, Cannonball Fm_ [holotype USNM 32447]
jaryensis, Doryssa transversa: Pilsbry Paratype
Pilsbry in Baker, 1914, p. 649
Rio Jary, Brazil; Sao Antonio da Cachoeira
jayana, Cancellaria (Narona): Keen Holotype
Keen, 1958a, p. 249, pl. 30, fig. 5. Also iz Keen, 1971, p. 651, fig. 1461
Panama Bay; 1 mile off Canal entrance, 10 fms
jayana, Cancellaria (Narona): Keen Paratype
Keen, 1958a, p. 249
Panama Bay; 1 mile off Canal entrance, 10 fms
jekylli, Entodina: Baker Paratype
Baker, 1914, p. 630
Brazil; Camp 39, M. & M. R.R., 284 km above Porto Velho
jordani, Buccinum: Hertlein Holotype
Hertlein, 1925b, p. 41, pl. 3, fig. 3
Montesano, Wash.; 8 miles up Sylvia Creek SU loc. 152 = NP 220
Miocene, Montesano Fm
judithae, Liocerithium: Keen Holotype
Keen, 1971, p. 411, fig. 517
Gulf of California; Angel de la Guarda Island
jUliana, Turritella variata: Merriam Plastoholotype
Merriam, 1941, p. 99, pl. 19, fig. 10
Santa Ynez Mts., Calif.; San Julian Ranch UCMP loc. A-312
Lower Oligocene, Gaviota Fm [holotype UCMP 33912]
keaseyense, Epitonium (Boreoscala): Durham Holotype
Durham, 1937, p. 498, pl. 57, fig. 17
Ore.; 3/4 mile W of Strassel SU loc. NP 292
Oligocene, Keasey Fm
426
6516
7915
7916
8035
10061
10062
10063
6268
5849
5850
9922
6714
6715
8659
8674
8677
5364
BuLLETIN 300
keenae, “Alvania” (Willettia): Gordon Holotype
Gordon, 1939, p. 31
San Mateo Co., Calif.; Moss Beach, among boulders
keenae, Ocenebra: Bormann Holotype
Bormann, 1946, p. 40, pl. 4, fig. 17
Los Angeles Co., Calif.; White’s Point
keenae, Ocenebra: Bormann Paratype
Bormann, 1946, p. 40, pl. 4, fig. 18
Los Angeles Co., Calif.; White’s Point
keenae, Rissoina: Smith and Gordon Paratype
Smith and Gordon, 1948, p. 227
Monterey Bay, Calif.; off Point Pinos, 5-15 fms
keenae, Septa (Monoplex) parthenopea: Beu Paratype
Beu, 1970, p. 233, pl. 2, figs. 6, 8
Mazatlan, Mexico; taken by shrimp dredger
keenae, Septa (Monoplex) parthenopea: Beu Paratype
Beu, 1970, p. 233, pl. 2, fig. 9
Sonora, Mexico; off Guaymas, taken by shrimp boats
keenae, Septa (Monoplex) parthenopea: Beu Paratype
Beu, 1970, p. 233, pl. 3, fig. 17
Galapagos; Albemarle Island, Tagus Cove
kincaidi, Turbonilla (Strioturbonilla): Bartsch Paratypes
Bartsch, 1921, p. 33
Puget Sound, Wash.; Dogfish Bay
klamathensis, Lanx (Walkerola): Hannibal Holotype
Hannibal, 1912b, p. 149, pl. 8, fig. 25a
Upper Klamath Lake, Ore.; Government Irrigation Dam
klamathensis, Lanx (Walkerola): Hannibal Paratype
Hannibal, 1912b, p. 149, pl. 8, fig. 25b
Upper Klamath Lake, Ore.; Government Irrigation Dam
klamathensis, Worthenia: Smith Plastoholotype
Smith, 1927, p. 108, pl. 96, fig. 3
Shasta Co., Calif.; N fork Squaw Creek, 3 miles N of Kellys Ranch
Upper Triassic, Hosselkus Ls [holotype USNM 74161]
knechti, Margarita optabilis: Arnold Paratypes
Arnold, 1903, p. 332
San Pedro, Calif.
Pleistocene, lower San Pedro Fm
kochi, Calliostoma dubium: Pallary Paratypes
Pallary, 1902b, p. 26
Tanger, Morocco [Tangier |
kurodai, Bittium: Makiyama Paratype
Makiyama, 1927, p. 66
Japan; Honohasi, Shizuoka Prefecture
Pliocene, Dainiti
kurodai, Nassarius (Hinia): Makiyama Paratype
Makiyama, 1927, p. 121
Shizuoka Prefecture, Japan; Dainiti
Pliocene, Dainiti
lahondaensis, Chlorostoma stantoni: Arnold Holotype
Arnold, 1908a, p. 388, pl. 36, fig. 2. Also in Arnold, 1909, illust. 2,
fig. 63
San Mateo Co., Calif.; Pescadero Creek just above mouth of Jones
Gulch, 3 miles S of La Honda
Upper Miocene, lower Purisima Fm [Pliocene] [Arnold’s specimen
1079]
5365
6229
7548
7549
5851
7985
7804
200
7545
6574a
STANFORD UNIVERSITY Tyres: SMITH 427
lahondaensis, Chlorostoma stantoni: Arnold Paratype
Arnold, 1908a, p. 388
San Mateo Co., Calif.; Pescadero Creek just above mouth of Jones
Gulch
Upper Miocene, lower Purisima Fm [Pliocene |
lalage, Mitrella: Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932a, p. 70
Mazatlan, Mexico
lampada, Typhis (Talityphis): Keen Holotype
Keen, 1943, p. 53, pl. 3, figs. 14, 19, 23
Kern Co., Calif.; Caliente Qd, center SW 1/4 Sec. 6, T 29 S, R 30 E,
in small gully SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
lampada, Typhis (Talityphis): Keen Paratype
Keen, 1943, p. 53
Kern Co., Calif.; Caliente Qd, center SW 1/4 Sec. 6, T 29 S, R 30 E,
in small gully SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
lancides, Fisherola: Hannibal Holotype
Hannibal, 1912b, p. 152, pl. 8, fig. 35a. Also ix Taylor and Smith,
1971, fig. 34
Snake River, Wash.
lancides, Fisherola: Hannibal Paratype
Hannibal, 1912b, p. 152
Snake River, Wash.
landesi, Ancistrolepis: Tegland Holotype
Tegland, 1933, p. 132, pl. 13, fig. 2
Puget Sound, Wash.; Bainbridge Island, beach between S side of
entrance to Blakeley Harbor and Restoration Point SU loc. NP 103
Upper Oligcecene, Blakeley Fm
landesi, Tritiaria (Antillophos): Marks Paratypes
Marks, 1951, p. 115
SW Ecuador; Las Masas area, NE Progreso Basin
Lower Miocene, Subibaja Fm
langi, Homorus (Subulona): Pilsbry Paratypes
Pilsbry, 1919, p. 115
Zambi, Belgian Congo, Africa
lawsoni, Pachychilus: Hannibal Holotype
Hannibal, 1912b, p. 183, pl. 8, fig. 23. Also im Taylor and Smith,
1971, figs. 43, 44 (as Lymnaea)
Alameda Co., Calif.; Berkeley Hills, near Bald Peak
Miocene, Contra Costa Lake beds
lens, Teinostoma (Teinostoma?): Keen Holotype
Keen, 1943, p. 51, pl. 4, figs. 7-9
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec: 6) 2 29'S, R30) ‘SU lees 2121
Miocene, Temblor Fm, Round Mountain Silt
leonina, Monadenia fidelis: Berry Paratype
Berry, 1937, p. 30
Siskiyou Co., Calif.; Beaver Creek, 1 mile above mouth
lepisma, Acmaea: Berry Holotype
Berry, 1940b, p. 155, pl. 17, figs. 3, 4
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene
lepisma, Acmaea: Berry Paratypes
Berry, 1940b, p. 155
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene
428
6227
8654
10220
10221
10222
114
138
6228
8713
6551
7532
8065
7101
6936
BuLtetin 300
leucocyma: Drillia: Dall Paratypes
Dall, 1883, p. 328
Key West, Fla.
leucostephes, Hertleinella: Berry Holotype
Berry, 1958b, p. 95. Illustrated iz Keen, 1971, p. 530, fig. 1023 left
Baja California, Mexico; E side Cedros Island
levis, Clivuloturris: Hickman Holotype
Hickman, 1976a, p. 78-79, pl. 6, figs. 10, 11
W. central Wash. SU loc. NP 50
Oligocene, Lincoln Creek Fm
levis, Clivuloturris: Hickman Paratype
Hickman, 1976a, pp. 78-79, pl. 6, fig. 6
W central Wash. SU loc. NP 50
Oligocene, Lincoln Creek Fm
levis, Clivuloturris: Hickman Paratype
Hickman, 1976a, pp. 78-79
W central Wash. SU loc. NP 50
Oligocene, Lincoln Creek Fm
lewisii, Gyrineum: Carson - Holotype
Carson, 1926, p. 53, pl. 2, fig. 1. Also zm Smith, 1970, p. 504, pl. 47,
fig. 8 [as Mediargo mediocris (Dall) ]
Santa Barbara Co. Calif.; Santa Maria District, Fugler’s Point
Lower Pliocene, Fernando Fm
lewisii, Gyrineum: Carson Paratype
Carson, 1926, p. 53, pl. 2, fig. 2. Also im Smith, 1970, p. 504, pl. 47,
fig. 4 [as Mediargo mediocris (Dall) ]
Santa Barbara Co., Calif.; Santa Maria District, Fugler’s Point
Lower Pliocene, Fernando Fm
limonitella, Drillia: Dall Paratypes
Dall, 1884, p. 329
Cedar Keys, Fla.
listrota, Turritella: Woodring Paratype
Woodring, 1959, p. 160
Canal Zone; Barro Colorado Island
Upper Oligocene, Bohio Fm
lloydi, Turris: Stanton Plastoholotype
Stanton, 1920, p. 45, pl. 8, fig. 16
North Dakota; Cannonball River, 7 miles S of Leith
Cretaceous, Cannonball Fm [holotype USNM 32445]
loismartinae, Cylichna?: Keen Holotype
Keen, 1943, p. 44, pl. 4, figs. 16, 18
Kern Co., Calif.; Caliente Qd, near Kern River, center SW 1/4 Sec.
6, T 29 S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
iombardii, Allogena: Smith Paratypes
Smith, 1943, p. 545
Idaho Co., Idaho; Meadow Creek, 1.5 miles S of Selway Falls
lorenzana, Turritella: Wagner and Schilling Plastoholotype
Wagner and Schilling, 1923, p. 257, pl. 50, fig. 11. Also in Merriam,
1941, p. 99, pl. 19, fig. 12
Kern Co., Calif.; near San Emigdio Canyon UCMP loc. 3217
Oligocene, Pleito Fm [holotype UCMP 11424]
louderbacki, Turris: Dickerson Plastosyntype
Dickerson, 1914, p. 147, pl. 16, fig. 9b
Contra Costa Co., Calif.; Mt. Diablo Qd, 1 mile S of Stewartville
UCMP loc. 1540
Eocene, Martinez Fm [syntype UCMP 11698]
7806
9501
6207
7869
152
5184
7866
6525
10280
10045
6256
8511
6161
10283
6168
5773
STANFORD UNIVERSITY TyYPEs: SMITH 429
lowei, Subulina: Pilsbry Paratypes
Pilsbry, 1919, p. 141
Africa; Belgian Congo
lunaris, Lunaia: Berry Holotype
Berry, 1964, p. 148. Illustrated iz Keen, 1971, p. 477, fig. 869 [as
Natica (Lunaia) |
Sonora, Mexico
lycodus, Bulimulus (Naesiotus): Dall Paratypes
Dall, 1917c, p. 379
Galapagos; Indefatigable Island
lyra, Scissurella: Berry Paratype
Berry, 1947b, p. 268
San Pedro, Calif.; near Second and Pacific Streets
Lower Pleistocene, Lomita
maccreadyi, Turritella: Waring Holotype
Waring, 1914, p. 783. Illustrated iz Waring, 1917, p. 87, pl. 12, fig. 10
Ventura Co., Calif.; Martinez area, Simi Hills, 3 miles NE of Simi
Peak
Lower Eocene, Martinez Fm_ [Paleocene]
maccreadyi, Turritella: Waring Paratype
Waring, 1914, p. 783
Ventura Co., Calif.; Martinez area, Simi Hills, 3 miles NE of Simi
Peak
Lower Eocene, Martinez Fm_ [Paleocene]
macfarlandi, Antiplanes: Berry Paratype
Berry, 1947b, p. 262
San Pedro, Calif.; Hilltop Quarry
Pleistocene, Lomita Fm
macgintyi, Murex: Smith Paratype
Smith, 1938, p. 88
Florida; Clewiston, Lake Okeechobee
Pliocene
macleani, Coralliophila: Shasky Paratype
Shasky, 1970, pp. 189-190
Sonora, Mexico; Guaymas, Saladita Bay, 27° 53’ 15” N, 110° 59’ W,
3-4 m on bases of white gorgonid sea whips
macleani, Decipifus: Keen Paratypes
Keen, 1971, p. 588
Baja California del Norte, Mexico; Puertecitos, intertidal
macra, Melanella (Balcis): Bartsch Paratypes
Bartsch, 1917, p. 326
Vancouver Island, British Columbia, Canada; Nanaimo, False Nar-
rows
macrospira, Hanetia: Berry Holotype
Berry, 1957, p. 79. Illustrated iz Keen, 1971, p. 563, fig. 1121 (as
Solenosteira)
Baja California, Mexico; about 8 miles N of San Felipe
maculata, Oreohelix: Henderson Paratypes
Henderson, 1921, p. 15
Northern Wyoming; White Creek Canyon
maesae, Maesiella: McLean and Poorman Paratype
McLean and Poorman, 1971, pp. 101-102
Sonora, Mexico; Guaymas, 1 mile S of Puerto San Carlos, 17 fms
magazinensis, Polygyra edentata: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1907, p. 545
Logan Co., Ark.; Magazine Mt.
magister, Stagnicola palustris: Henderson ex Baker Ms Holotype
Henderson, 1934b, pl. 14, fig. 1 left. Described iz Baker, 1934b, p. 17
Modoc Co., Calif.; E shore Rhett (Tule) Lake
430
9773a
5440
8755
9522
8752
7190
6230
8666
7547
7547a
6510
8096
6554
7990
5395
5396
5397
5398
5399
BuLLeETIN 300
magister, Stagnicola palustris: Henderson ex Baker MS Paratype
Henderson, 1934b, pl. 14, fig. 1, right. Baker, 1934b, p. 17
Modoc Co., Calif.; E shore, Rhett (Tule) Lake
magna, Lirularia: Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 36
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
mamillatum, Stephopoma: Morton and Keen Paratype
Morton and Keen, 1960, p. 28, pl. 1, fig. 2
West Africa, off Gorée, Senegal; 27 fms
manca, Odostomia (Evalea): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 32
Los Angeles, Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
marchadi, Dendropoma: Keen and Morton Paratypes
Keen and Morton, 1960, p. 37, pl. 2, fig. 3
Senegal, West Africa; Gorée
margaritana, Turritella: Nomland Plastoholotype
Nomland, 1917b, p. 312, pl. 20, fig. 5. Also in Merriam, 1941, p. 120,
pl. 34, fig. 10
San Luis Obispo Co., Calif.; UCMP loc. 1706
Upper Miocene, Santa Margarita Fm [holotype UCMP 11312]
mariamadrae, Tegula mariana: Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932a, p. 85
Gulf of California; Tres Marias Islands, Isla Maria Madre
mariposa, Monadenia (Corynadenia) hillebrandi: Smith Paratype
Smith, 1957, p. 24
Mariposa Co., Calif.; McLean Cave
mariposa, Turbonilla (Pyrgolampros): Keen Holotype
Keen, 1943, p. 52, pl. 4, fig. 19
Kern Co., Calif.; Caliente Qd, small gully near center SW 1/4 Sec.
6, T 29S,R30E_ SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
mariposa, Turbonilla (Pyrgolampros): Keen Paratype
Keen, 1943, p. 52, pl. 4, fig. 25
Kern Co., Calif.; Caliente Qd, small gully near center SW 1/4 Sec.
60 2935S S0,E SUiloe: 2121
Miocene, Temblor Fm, Round Mountain Silt
markleyensis, Pseudoliva: Clark Paratype
Clark, 1938, p. 710
Napa Qd, Calif.; Brink Ranch, 2 miles S of Putah Creek
Upper Eocene, Markley Fm
marksi, Strombina: Hertlein and Strong Paratype
Hertlein and Strong, 1951a, p. 84
Gulf of California; near Arena Bank, 23° 29’ 30” N, 109° 25’ 30” W,
45 fms
marmarotis, Monadenia: Berry Paratypes
Berry, 1940a, p. 3
Siskiyou Co., Calif.; Marble Valley, near Ranger Station
masasensis, Conus (Leptoconus): Marks Paratypes
Marks, 1951, p. 139
Guayas Province, Ecuador; near Las Masas
Lower Miocene, Subibaja Fm
mateoensis, Patella: Arnold Paratypes
Arnold, 1908a, p. 362
San Mateo Co., Calif.; ridge between San Lorenzo River and Pesca-
dero Creek
Eocene, Martinez Fm ?
9400
5401
9402
5403
10278a
9856
9738
9742
6167
10041
7802
7031
6565
6151
9750
8591
10281
5929
STANFORD UNIVERSITY TyPEs: SMITH 431
mateoensis, Patella: Arnold Paratypes
Arnold, 1908a, p. 362
San Mateo Co., Calif.; ridge between San Lorenzo River and Pesca-
dero Creek
Eocene, Martinez Fm ?
matthewsi, Ancilla: Burch and Burch Paratype
Burch, J. Q., and Burch, R. L., 1967a, pp. 81-82
Off Fortaleza, Ceara, Brazil; from digestive tract of toad fish
Amphichthys cryptocentrus (Val., 1837)
matthewsi, Marginella (Prunum): van Mol and Tursch Holotype
van Mol and Tursch, 1967, pp. 196-197, fig. 1
Off Fortaleza, Ceara, Brazil; from stomachs of fishes known locally
as “pacamao”, 20 fms
mauryi, Epitonium (Epitonium): Tursch and Pierret Holotype
Tursch and Pierret, 1964, p. 36, fig. 5
Rio de Janeiro, Brazil; off Punta de Juatinga 23° 22’ S, 48° 28’ W,
50 m
mcleani, Calliostoma: Shasky and Campbell Holotype
Shasky and Campbell, 1964, p. 117, pl. 22, figs. 21, 24. Also in Keen,
1971, p. 334, fig. 86
Sonora, Mexico; Guaymas, NW of Bahia Saladita, 2-15 m
media, Holospira bilamellata: Pilsbry Paratypes
Pilsbry, 1915, p. 339
Hacheta Grande Mts., New Mexico; Sheridan Canyon
medialis, Episcynia: Keen Holotype
Keen, 1971, p. 381, fig. 352
Guaymas, Mexico; off Cabo Haro, 18 m
medjensis, Limicolaria laeta: Pilsbry Paratypes
Pilsbry, 1919, p. 97
Africa; Medje, Belgian Congo
meganosensis, Turritella: Clark and Woodford Plastoholotype
Clark and Woodford, 1927, p. 119, pl. 21, fig. 2. Also im Merriam,
1941, p. 75, pl. 8, fig. 3
Contra Costa Co., Calif.; Mt. Diablo area UCMP loc. 3159
Eocene, Meganos Fm_ [holotype UCMP 12445]
melanopylon, Micrarionta (Eremarionta): Berry Paratypes
Berry, 1930c, p. 187
San Bernardino Co., Calif.; W side Black Canyon, 9 miles N of
Hinkley
mendax, Ashmunella: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1917, p. 92
New Mexico; Gallina Canyon, Black Range
mendella, Tegula (Agathistoma): McLean Paratype
McLean, 1964, p. 131
San Diego Co., Calif.; Mission Bay, depths to 10’
mendozana, Hanetia: Berry Holotype
Berry, 1959, p. 111 [= Solenosteira mendozana (Berry), fide Keen,
1971, p. 563]
Baja California, Mexico; Magdalena Bay, 10-25 fms
mendozana, Strombina (Cotonopsis): Shasky Paratype
Shasky, 1970, p. 194
Gulf of Fonseca, El] Salvador; 15° 57’ N, 95° 32’ W, 33-73 m
menzola, Odostomia (Amaura): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 33
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
432
496
10298
6433
6566
8079
8080
6164
8588
7578
6171
8714
165
8105
8660
6219
BuLLeETIN 300
merriami, Drillia: Arnold Plastoholotype
Arnold, 1903, p. 207, pl. 8, fig. 7
Los Angeles Co., Calif.; Deadman Island
Pleistocene, lower San Pedro Fm [holotype USNM]
mertensi, Leptinaria: Dall Paratype
Dall, 1900a, p. 97
Cocos Island, Costa Rica
mexicana, Olivella boetica: Oldroyd Holotype
Oldroyd, T. S., 1921b, p. 118, pl. 5, fig. 3. Also zz Oldroyd, I. S., 1927,
pr 163" pl Z6,nics: ale Zilla
Baja California, Mexico; Scammons Lagoon
micrometalleus, Micrarionta (Eremarionta): Berry Paratypes
Berry, 1930c, p. 189
Kern Co., Calif.; 3.5 miles S of Petrified Forest, Elpaso Range
micromphalus, Menetus?: Taylor Paratype
Taylor, 1954, p. 74, pl. 20, figs. 7-9
San Bernardino Co., Calif.; W end Barstow Hills in “Lake Bed
Horizon’, canyon next S from Pirie Canyon, middle of SE 1/4 See.
1 Seen eNE eRe VV.
Upper Miocene, Barstow Fm ?
micromphalus, Menetus?: Taylor Paratype
Taylor, 1954, p. 74
San Bernardino Co., Calif.; W end Barstow Hills, middle of SE 1/4
Sec. 15, T 11 N, R2 W
Upper Miocene, Barstow Fm ? [also = paratype of Planorbis mo-
javensis Hannibal, 1912b, p. 157]
millepalmarum, Micrarionta (Erenarionta): Berry Paratypes
Berry, 1930b, p. 543, as mille-palmarum
Riverside, Calif.; Thousand Palms
milleri, Lucapinella: Berry Holotype
Berry, 1959, p. 109
Baja California, Mexico; Puertecitos
milleri, Trigonostoma: Burch Paratype
Burch, 1949, p. 3. Illustrated 72 Keen, 1971, p. 656, fig. 1480
Costa Rica; Tambor, near Puntarenas
millestriata, Holospira: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915a, p. 380
Dragoon Mts., Arizona
mimeticum, Cerithium (Thericium): Woodring Paratypes
Woodring, 1959, p. 171
Panama Canal Zone; Barro Colorado Island
Upper Oligocene, upper Bohio Fm
miranda, Cyclostrema: Bartsch Paratypes
Bartsch, 1911la, p. 230
San Pedro, Calif. [Moore, 1969, pp. 169-170 points out that the
locality is erroneous and that the specimens are Tornus subcarinatus
(Montagu, 1803), a European taxon |
mirimense, Cerithium (?): Jenkins Syntypes
Jenkins, 1913, p. 450, pl. 20, fig. 8. Also iz Maury, 1934, p. 150, pl.
14, fig. 3
Rio Grande de Norte, Brazil; near Itapasaroca
“RKocene”’ [specimens missing, 1936]
miscowichi, Ocinebra: Pallary Paratype
Pallary, 1906, p. 3
Mogador, Morocco
mitchelli, Acmaea striata: Oldroyd Paratypes
Oldroyd, 1933, p. 205
Philippine Islands; southern Luzon
6220
8621
9777
5777a
8077
8078
5460
6210
5158
6451
6265
9503
STANFORD UNIVERSITY TYPES: SMITH 433
mitchelli, Nerita: Oldroyd Paratypes
Oldroyd, I. S., 1933, p. 205
Philippine Islands
mitriformis, Arielia: Shasky Holotype
Shasky, 1961, p. 20, pl. 4, figs. 7-9. Also in Keen, 1971, p. 741, fig.
1769 (as Mitrolumna)
Gulf of California; off Isla Espiritu Santo, 40-90 fms
modoci, Fluminicola: Hannibal Holotype
Hannibal, 1912b, p. 187, pl. 8, fig. 30. Also zz Taylor and Smith, 1971,
figs. 16, 21 [as Lithoglyphus turbiniformis (Tryon, 1865) ]
S end of Goose Lake, Calif.; Fletcher’s Spring
modoci, Fluminicola: Hannibal Paratype
Hannibal, 1912b, p. 187
S end of Goose Lake, Calif.; Fletcher’s Spring
mohaveana, Lymnaea: Taylor Holotype
Taylor, 1954, p. 73, pl. 20, figs. 1, 2
San Bernardino Co., Calif.; “Lake bed horizon” in canyon next S
from Pirie Canyon, W end of Barstow Hills, middle of SE 1/4 Sec.
15,T11N,R2W
Upper Miocene, Barstow Fm
mohaveana, Lymnaea: Taylor Paratype
Taylor, 1954, p. 73
San Bernardino Co., Calif.; W end Barstow Hills, middle of SE 1/4
Sec. 15, IT 11 N, R 2 W
Upper Miocene, Barstow Fm
mojavensis, Planorbis: Hannibal Paratype
Hannibal, 1912b, p. 157
San Bernardino Co., Calif.; near Barstow, Mojave Desert
Upper Miocene, Barstow Fm [= paratype 8080 Menetus (?) microm-
phalus Taylor, 1954]
monotaenius, Bulimulus (Naesiotus) nux: Dall and Ochsner
Paratypes
Dall and Ochsner, 1928, p. 157
Galapagos; Charles Island
montereyana, Turritella: Wiedey Syntype
Wiedey, 1928, p. 123, pl. 21, fig. 2
Monterey Co., Calif.; Bryson Qd, 1.5 miles S of San Antonio River
SU loc. 447
Middle Miocene, Monterey Fm
montereyensis, Astraea inaequalis: Oldroyd Holotype
Oldroyd, I. S., 1927, p. 165, pl. 108, figs. 5, 6
Monterey, Calif.
montereyensis, Odostomia (Chrysallida): Dall and Bartsch
Paratype
Dall and Bartsch, 1907, p. 516
Monterey, Calif.; 12 fms
montereyensis, Retusa (Sulcularia): Smith and Gordon Paratype
Smith and Gordon, 1948, p. 217
Monterey Bay, Calif.; off Del Monte, 8-15 fms
morani, Astraea: Loel and Corey Holotype
Loel and Corey, 1932, p. 271, pl. 64, figs. 6a, 6b
San Luis Obispo Co., Calif.; Corral de Piedra Creek
Lower Miocene, Vaqueros Fm
morani, Zalophancylus: Hannibal Holotype
See Chordata
mousleyi, Melampus: Berry Holotype
Berry, 1964, p. 152. Illustrated iz Keen, 1971, pp. 844-846, fig. 2399
Sonora, Mexico; Cholla Cove, Bahia Adair, upper estero
434
10064
269
270
6172
7047
7860
6224
6146
8589
8529
9991
8756
8756a
8678
8675
5508
Bu t.etin 300
mucronata, Primovula: Azuma and Cate Paratype
Azuma and Cate, 1971, p. 264
Kirimeaski, Kii, Japan; 25 fms
muirensis, Antiplanes: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 157, pl. 30, fig. 6
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
muirensis, Antiplanes: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 157, pl. 30, fig. 4
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Muir and Coal Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
mularis, Holospira arizonensis: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915a, p. 386
Mule Mts., Arizona; Escabrosa Ridge
mulleri, Turritella andersoni: Merriam Plastoholotype
Merriam, 1941, p. 80, pl. 11, fig. 2
Ventura Co., Calif.; 1.5 miles W of Vickers Hot Springs UCMP loc.
A-1414
Middle Eocene [holotype UCMP 15297]
murrha, Agaronia: Berry Paratype
Berry, 1953b, p. 417
Corinto, Nicaragua
mutata, Cerithidea: Pilsbry and Vanatta Syntypes
Pilsbry and Vanatta, 1902, p. 558
Galapagos; Albemarle Island, Tagus Cove
mutator, Ashmunella tetrodon: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915b, p. 31
Socorro Co., New Mexico; Mogollon Mts., Dry Creek Canyon
myrae, Nomaeopelta: Berry Holotype
Berry, 1959, p. 109. Illustrated iz Keen, 1971, p. 327, fig. 56b
Sinaloa, Mexico; Mazatlan, Las Gaviotas Beach
myrae, Trivia (Pusula): Campbell Holotype
Campbell, 1961b, p. 25, pl. 5, figs. 1-3. Also im Keen, 1971, p. 487, fig.
907
Baja California, Mexico; off Loreto, 25 fms
myrakeenae, Aspella (? Dermomurex): Emerson and D’Attilio
Paratype
Emerson and D’ Attilio, 1970, pp. 89-92
Nayarit, Mexico; Banderas Bay, intertidal under rocks 22° 44’ N,
105° 29’ W
myrakeenae, Stephopoma: Olsson and McGinty Paratypes
Olsson and McGinty, 1958, p. 35
Panama; Atlantic coast, near Bocas del Toro
nakamurai, Thais: Makiyama Paratype
Makiyama, 1927, p. 128
Shizuoka Prefecture, Japan; Dainiti
Pliocene, Dainiti
nakamurai, Uromitra: Makiyama Paratype
Makiyama, 1927, p. 78
Shizuoka Prefecture, Japan; Tennoyama
Pliocene, Dainiti
nanella, Marginella jewettii: Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 24
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
379
8108
8346
7065
274
6241
108
5409
8571
199
STANFORD UNIVERSITY Types: SMITH 435
nanus, Strombus raninus: Bales Paratypes
Bales, 1942, p. 19
Lake Worth, Fla.
natalensis, Turritella: Jenkins Syntypes
Jenkins, 1913, p. 451, pl. 20, figs. 6, 6a
Brazil; Rio Grande do Norte, near Itapasaroca
Eocene? [Cretaceous, fide Maury, 1934, pp. 126, 143]
natlandi, Hemitoma: Durham Plastoholotype
Durham, 1950, p. 132, pl. 28, figs. 7, 8
Gulf of California; Coronado Island UCMP loc. A-3548
Pleistocene [holotype UCMP 30474]
nelsonensis, Patella (?): Trechmann Paratype
Trechmann, 1918, p. 185
New Zealand; Nelson District, Eighty-Eight Valley
Triassic, Kaihiku
neopleura, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 93, pl. 15, fig. 6
Kern Co., Calif.; Tejon Qd, Liveoak Canyon UCMP loc. 7182
Eocene, Tejon Fm_ [holotype UCMP 33873 |
nepos, Helix intercisa: Hemphill Paratypes
Hemphill, 1891, p. 330
San Clemente Island, Calif.
nevadensis, Oreohelix: Berry Paratypes
Berry, 1932, p. 60
White Pine Co., Nevada; Shell Creek Mts., Cleve Creek, elev. 8100’
nevadensis, Parapholyx effusa: Henderson Holotype
Henderson, 1934a, p. 91, pl. 9, fig. 6, second from left
Winnemucca Lake, Nevada
nevadensis, Parapholyx effusa: Henderson Paratypes
Henderson, 1934a, p. 91, pl. 9, fig. 6 except second from left
Winnemucca Lake, Nevada
newcombei, Cerithidea: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 163, pl. 31, figs. 4a, 4b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Sooke SU loc. NP 129
Oligocene, Sooke Fm
newcombei, Cerithidea: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 163, pl. 31, fig. 5
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Sooke SU loc. NP 129
Oligocene, Sooke Fm
newcombiana, Paludinella: Hemphill Syntypes
Hemphill, 1876, p. 49
Humboldt Bay, Calif.
newhallensis, Cancellaria: Carson Holotype
Carson, 1926, p. 56, pl. 3, fig. 3
Los Angeles Co., Calif.; Elsmere Canyon
Lower Pliocene
newsomi, Pleurotoma: Arnold Plastoholotype
Arnold, 1908a, p. 368, pl. 33, fig. 2
Santa Cruz Co., Calif.; Boulder Creek, 2.25 miles N of Eagle Rock
Oligocene, San Lorenzo Fm [holotype USNM]
‘ninfae, Terebra (Strioterebrum): Campbell Paratype
Campbell, 1961b, p. 27
Chiapas, Mexico; about 30 miles N of Guatemalan border
nipponensis, Cylichna: Nomura and Hatai Paratype
Nomura and Hatai, 1940, p. 72
Aomori-Ken, NE Honsyu, Japan; off Kyuroku-Shima
436
9731
9730
5950
7182
6160
8259
6248
5817
5448
5449
6208
7157
8717
5892
6264
6410a
6410¢
6410b
BULLETIN 300
nipponensis, Siphonochelus (Siphonochelus): Keen and Campbell
Plastoholotype
Keen and Campbell, 1964, p. 50, pl. 10, fig. 25
Off Tosa, Japan, 200+ m_ [holotype in Kyoto, Japan, private collec-
tion of Mr. Akibumi Teramachi]
nipponensis, Siphonochelus (Siphonochelus): Keen and Campbell]
Paratype
Keen and Campbell, 1964, p. 50
Off Tosa, Japan; 200+ m
nodosus, Vermetus: Oldroyd Holotype
See Pelecypoda
nova, Turritella: Nomland Plastoholotype
Nomland, 1916b, p. 208, pl. 11, fig. 3. Also zz Merriam, 1941, p. 119,
pl. 34, fig. 7 (as Turritella cooperi nova)
Fresno Co., Calif.; Waltham Canyon UCMP loc. 2533
Pliocene, Jacalitos Fm [holotype UCMP 12060]
obscura, Oreohelix cooperi: Henderson Paratypes
Henderson, 1918, p. 46
White Creek Canyon, Wyoming :
obstipa, Balcis (Vitreolina): Berry Paratype
Berry, 1954b, p. 262
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene, Lomita Fm.
occidentalis, Eulimella: Hemphill Paratypes
Hemphill, 1894, p. 395
San Diego, Calif.
occidentalis, Limnaea stagnalis: Hemphill Paratypes
Hemphill, 1890, p. 26
Whatcom Co., Wash.; Whatcom Lake, Bellingham
ochromphalus, Monadenia fidelis: Berry Paratypes
Berry, 1937, p. 28
Siskiyou Co., Calif.; Etna Creek, 2.5 miles above Etna
ochsneri, Helicina (Idesa): Dall Paratypes
Dall, 1917c, p. 382
Galapagos; Albemarle Island, Cowley Mt.
ocoyana, Turritella: Conrad Plastoneotype
Conrad, 1855, p. 329, pl. 8, figs. 73a, 73b. Neotype designated by
Merriam, 1941, p. 112, pl. 29, fig. 5
Kern Co., Calif.; Poso Creek, Kern River region UCMP loc. 2713
Middle Miocene, Temblor Fm [neotype UCMP 31641]
oeciscus, Hemisinus (Longiverena): Woodring Paratype
Woodring, 1959, p. 157
Panama Canal Zone; Barro Colorado Island
Upper Oligocene, upper Bohio Fm
oldroydae, Chilina: Marshall Paratype
Marshall, 1924, p. 4
Chubut Province, Argentina; Lake Fetalafquen, Andes
oldroydae, Diastoma: Bartsch Paratype
Bartsch, 1911b, p. 583
San Pedro, Calif.
oldroydi, Acteocina: Dall Paratypes
Dall, 1925, p. 25
Vancouver Island, British Columbia, Canada; Nanaimo, Departure
Bay
oldroydi, Acteocina: Dall Paratype
Dall, 1925, p. 25. Also in Oldroyd, 1927, p. 28, pl. 2, fig. 4
Vancouver Island, British Columbia, Canada; Nanaimo, Departure
Bay
6455
6456
6457
429
6252
104
6447
459
5828
7624
8351
6269
8058
10203
6184
STANFORD UNIVERSITY Types: SMITH 437
oldroydi, Coralliophila: Oldroyd Holotype
Oldroyd, I. S., 1929, p. 98, pl. 5, figs. 1, 2. Also iz McLean, 1969, p.
44, fig. 23.4 (as Latiaxis)
Catalina Island, Calif.; off Isthmus, Bird Rock
oldroydi, Coralliophila: Oldroyd Paratype
Oldroyd, I. S., 1929, p. 98, pl. 5, fig. 4
Catalina Island, Calif.; off Isthmus, Bird Rock
oldroydi, Coralliophila: Oldroyd Paratype
Oldroyd, I. S., 1929, p. 98, pl. 5, fig. 3 [not fig. 4 as stated]
Galapagos; Indefatigable Island
oldroydi, Mangilia: Arnold Holotype
Arnold, 1903, p. 213, pl. 6, fig. 16
Los Angeles Co., Calif.; Deadman Island
Pleistocene, lower San Pedro Fm
oldroydi, Melanella (Melanella): Bartsch Paratypes
Bartsch, 1917, p. 309
San Pedro, Calif.
oldroydia, Cancellaria: Carson Holotype
Carson, 1926, p. 51, pl. 1, fig. 5
San Mateo Co., Calif.; near mouth of Purisima Creek
oldroydii, Sigaretus: Dall Holotype
Dall, 1897c, p. 85. Illustrated zz Dall, 1921, pl. 14, figs. 1, 3 (as
Eunaticina). Also in Oldroyd, I. S., 1927, pl. 92, figs. 11, 1la
Off Catalina Island, Calif.; in deep water
olequaensis, Ambloxus: Hannibal Holotype
Hannibal, 1912b, p. 178, pl. 8, fig. 27. Also in Taylor and Smith, 1971,
figs. 22, 23 (as Juga)
Wash.; Olequa Creek, 2 miles N of Little Falls
Eocene [Late Eocene, Cowlitz Fm, fide Taylor and Smith, 1971, p.
311]
olivacea, Chilina: Marshall Paratypes
Marshall, 1924, p. 4
Chubut, Argentina; 43° 20’ S, 71° 30’ W, Rio Corcovado
olympicensis, Perse: Durham Paratype
Durham, 1944, p. 174
Jeferson Co., Wash.; Point Nill, Port Discovery SU loc. NP 151
Lower Oligocene, Quimper Ss
olympicensis, Turritella: Durham Plastoholotype
Durham, 1944, p. 163, pl. 17, fig. 1
Jefferson Co., Wash.; UCMP loc. A-3702
Lower Oligocene, Quimper Ss_ [holotype UCMP 35318]
onealensis, Cerithiopsis: Bartsch Paratypes
Bartsch, 1921, p. 35
Puget Sound, Wash.; off O’Neal Island, 20 fms
onoyamai, Bittium: Oinomikado and Ikebe Paratypes
Oinomikado and Ikebe, 1939, p. 105
Toyama Prefecture, Japan; Tagawa, Konade-mura, Nishi Tonami-
gun
Upper Pliocene, Tagawa beds
oregonensis, Comitas (Boreocomitas): Hickman Paratype
Hickman, 1976, p. 43, pl. 2, fig. 14
Ore. SU loc. H 40
Oligocene, Keasey Fm, upper mbr
oria, Polygyra columbiana: Berry Paratypes
Berry, 1933, p. 15
Eldorado Co., Calif.; § fork American River Canyon near Riverton
438
6215
6181
7859
8095
10325
7848
7848a
10060
6192
8073
8074
10332
7000
107
7780
7797
BuLteTIin 300
orina, Helminthoglypta: Berry Paratypes
Berry, 1938b, p. 41
Kern Co., Calif.; near summit of Breckinridge Mt.
orotis, Vitrea: Berry Paratype
Berry, 1930a, p. 113
San Diego Co., Calif.; Palomar Mts., E of Palomar P.O., 5000’ elev.
orthosymmetra, Turritella: Berry Paratype
Berry, 1953b, p. 412
Santa Catalina Island, Calif.; off Pebbly Beach, 50 fms
osborni, Aesopus: Hertlein and Strong Paratypes
Hertlein and Strong, 1951a, p. 83
Off Port Guatulco, Mexico; 15° 44’ 28” N, 96° 07’ 51” W, 7 fms
ossa, Turritella: Popenoe Plastoholotype
Popenoe, 1937, p. 401, pl. 49, fig. 6
Santa Ana Mts., Calif.; CIT loc. 1066
Cretaceous, Campanian [holotype UCLA 40674]
ovuliformis, Pedicularia californica: Berry Holotype
Berry, 1946c, p. 3, fig. 1
Santa Catalina Island, Calif.; Farnsworth Bank, 42 m
ovuliformis, Pedicularia californica: Berry Paratype
Berry, 1946c, p. 3
Santa Catalina Island, Calif.; Farnsworth Bank, 42 m
oweni, Haliotis corrugata: Talmadge Paratype
Talmadge, 1966, p. 1
Guadalupe Island, off Baja California, Mexico; 20’
ozarkensis, Omphalina fuliginosa: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1907, p. 562
Logan Co., Ark.; Petit Jean Mts.
pachyostracon, Craterarion: Taylor Holotype
Taylor, 1954, p. 75, pl. 20, figs. 18-20
San Bernardino Co., Calif.; canyon S of Pirie Canyon, SE 1/4 Sec.
Gy AP UBB ING Le OY
Upper Miocene, Barstow Fm
pachyostracon, Craterarion: Taylor Paratypes
Taylor, 1954, p. 75
San Bernardino Co., Calif.; canyon S of Pirie Canyon, SE 1/4 Sec.
1 IP SIT, RB AY
Upper Miocene, Barstow Fin
packardi,.,Ampullina: Popenoe Plastoholotype
Popenoe, 1937, p. 399, pl. 49, figs. 4, 5
Santa Ana Mts., Calif.; CIT loc. 1054
Cretaceous, Campanian [holotype UCLA 40667]
packardi, Turritella: Merriam Plastoholotype
Merriam, 1941, p. 66, pl. 3, fig. 6
Orange Co., Calif.; Santa Ana Mts. UCMP loc. A-810
Upper Cretaceous, “Chico” Fm _ [holotype UCMP 15362]
palmeri, Cancellaria: Carson Holotype
Carson, 1926, p. 55, pl. 2, fig. 4
Santa Cruz Co., Calif.; bluffs above beach E of hotel at Capitola
Lower Pliocene, Purisima Fm
Paparyensis, Segmentina: Baker Paratype
Baker, Fred, 1914, p. 662
Brazil; mouth of main affluent of Papary Lake
pareximia, Actaeopyramis: Nomura Paratypes
Nomura, 1936, p. 19
NE Honsyu, Japan; Siogama Bay
10293
8514
6438
5387
8682
5509
5506
7196
503
6254
10058
7981
8353
10048
6209
STANFORD UNIVERSITY [TyPEs: SMITH 439
parkeri, Turritella: McLean Paratypes
McLean, 1970c, p. 127
Baja California, Mexico; Bahia de la Paz, W of Espiritu Santo
Island, 24° 24.3’ - 24° 25.6’ N,
110° 23.7’ - 110° 25.5’ W, 45-65 fms
parthenia, Pleuroliria: Berry Holotype
Berry, 1957, p. 81. Illustrated in Keen, 1958b, p. 477, fig. 912
Gulf of Nicoya, Costa Rica; off Isla Tortugas, 10 fms
parva, Olivella biplicata: Oldroyd Holotype
Oldroyd, T. S., 1921, p. 119, pl. 5, fig. 7
Baja California, Mexico; Point Abreojos
Patella n. sp. b: Arnold Syntypes
Arnold, 1908a, p. 362
San Mateo Co., Calif.; ridge between San Lorenzo River and Pesca-
dero Creek
Eocene, Martinez Fm ?
paucilirata, Siphonalia: Makiyama Paratype
Makiyama, 1941, p. 88
Chiba-ken, Japan; coast at Sasage
Lower Pleistocene, Kanozan Fm
pecora, Turbonilla (Strioturbonilla): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 24
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pablo Fm
pedroensis, Acteocina: Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 23
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
pedroensis, Turritella: Applin Plastoholotype
Applin MS in Merriam, 1941, p. 121, pl. 35, fig. 5
Los Angeles Co., Calif.; Timms Point, San Pedro UCMP loc. 7102
Pleistocene [holotype UCMP 15236]
pembertoni, Ataphrus: Hall and Ambrose Holotype
Hall and Ambrose, 1916, p. 70. Illustrated in Wiedey, 1929b, pl. 1,
fig. 7
Alameda Co., Calif.; Tesla Qd, Jordan Ranch, Arroyo del Valle
Upper Cretaceous, lower Chico Fm
peninsularis, Melanella (Balcis): Bartsch Paratypes
Bartsch, 1917, p. 320
Baja California, Mexico; San Hipolite Point and Pt. Abreojos
pentedesmium, Cirsotrema: Berry Holotype
Berry, 1963, p. 143. Illustrated in Keen, 1971, p. 428, fig. 634 left
[as Epitonium (Cirsotrema) vulpinum (Hinds, 1844) ]
Guaymas, Mexico; San Carlos, 15-30 fms
pequenita, Strombina: Marks Paratype
Marks, 1951, p. 111
SW Ecuador; Zacachun corehole, 80-90’
Lower Miocene, upper Subibaja Fm
perangulatus, Murex: Nomland Plastoholotype
Nomland, 1916b, p. 206, pl. 11, figs. 1a, 1b
Fresno Co., Calif.; Coalinga-Priest Valley Road
Lower Pliocene [holotype UCMP 10257]
perata, Nassarina (Cigclirina): Keen Paratype
Keen, 1971, p. 594, fig. 1248
Chiapas, Mexico; Puerto Videra, 37-45 m
perchloris, Bulimulus (Naesiotus) nux: Dall and Ochsner
Paratypes
Dall and Ochsner, 1928, p. 156
Galapagos; Charles Island
440
6195
8716
5386
8496
213
102
103
103a
7763
277
6996
6204
8498
BuL.etin 300
percostata, Polygyra dorfeuilliana: Pilsbry Paratypes
Pilsbry, 1899b, p. 37
Arkansas; Red River, near Texarkana
pericallum, Bittium: Woodring Paratype
Woodring, 1959, p. 179
Panama Canal Zone; Mount Hope Cemetery area
Middle Miocene, upper Gatun Fm
perissolaxoides, Pleurotoma: Arnold Paratype
Arnold, 1908a, p. 368
Santa Cruz Co., Calif.; San Lorenzo River, 3.75 miles above Boulder
Creek
Oligocene, San Lorenzo Fm
perplexa, Aspella: Keen Holotype
Keen, 1958a, p. 248, pl. 30, fig. 11. Also im Keen, 1971, p. 527, fig.
1014 left [as Aspella (Dermomurex) indentata (Carpenter, 1857) ]
Perlas Islands, Panama
perrini, Acanthina: Trask Holotype
Trask, 1922, p. 157, pl. 8, figs. 1a, 1b
6 miles S of Livermore, Calif.
Miocene, Briones Fm
perrini, Acanthina: Trask Paratype
sirask 92250 poe 517,
6 miles S of Livermore, Calif.
Miocene, Briones Fm
perrini, Cancellaria: Carson Holotype
Carson, 1926, p. 56, pl. 3, fig. 4
Santa Barbara Co., Calif.; Santa Maria District, Fugler’s Point
Lower Pliocene, Fernando Fm
perrini, Cancellaria: Carson Paratype
Carson, 1926, p. 56
Los Angeles Co., Calif.; Elsmere Canyon
Lower Pliocene, Fernando Fm
perrini, Cancellaria: Carson Paratype
Carson, 1926, p. 56
Santa Barbara Co., Calif.; Santa Maria District, Fugler’s Point
Lower Pliocene, Fernando Fm
perrini, Fissurella: Arnold Paratype
Arnold, 1908a, p. 362
San Mateo Co., Calif.; ridge between headwaters of San Lorenzo
River and Pescadero Creek, Santa Cruz Qd_ SU loc. 2694
“Eocene, Martinez Fm”
perrini, Rapana: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 161, pl. 31, fig. 7
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
perrini, Turritella chicoensis: Merriam Plastoholotype
Merriam, 1941, p. 66, pl. 2, fig. 3
Corona Qd, Calif.; near Santiago Canyon UCMP loc. 2154
Upper Cretaceous, “Chico” Fm_ [holotype UCMP 15366]
perrus, Bulimulus (Naesiotus): Dall Paratypes
Dall, 1917c, p. 376
Galapagos; Narborough Island, 2000-4500’ elev.
personatum, Crucibulum: Keen Holotype
Keen, 1958a, p. 247, pl. 30, figs. 6, 8. Also in Keen, 1971, p. 463, fig.
824
Pacific coast of Panama
8499
418
7130
5896
5891
6575
6211
460
5821
10290
6153
153
481
5420
STANFORD UNIVERSITY Tyres: SMITH 44]
personatum, Crucibulum: Keen Paratype
Keen, 1958a, p. 247, pl. 30, fig. 7
Pacific coast of Panama
pertumida, Turritella inezana: Wiedey Paratype
Wiedey, 1928, p. 119, pl. 12, fig. 6
San Luis Obispo Co., Calif.; Canal de Pietro, or Corral de Piedra,
5 miles E of San Luis Obispo
Miocene, Vaqueros Fm [= hypotype 418, Turritella inezana Waring,
1915, fig. 28]
pervulgata, Turritella inezana: Merriam Plastoholotype
Merriam, 1941, p. 108, pl. 25, fig. 11
Ventura Co., Calif.; Ojai Valley UCMP loc. A-330
Lower Miocene, Vaqueros Fm_ [holotype UCMP 31686]
pescaderoensis, Turritella: Arnold Holotype
Arnold, 1908a, p. 358, pl. 31, fig. 7. Also in Merriam, 1941, p. 66, pl.
Ze hige 5
San Mateo Co., Calif.; 2.5 miles N of Bolsa Point, 1 mile S of Arroyo
de los Frijoles
Upper Cretaceous, Chico Fm
pescaderoensis, Turritella: Arnold Paratype
Arnold, 1908a, p. 358
San Mateo Co., Calif.; 2.5 miles N of Bolsa Point, 1 mile S of Arroyo
de los Frijoles
Upper Cretaceous, Chico Fm
petrothauma, Astraea (Pomaulax): Berry Paratype
Berry, 1940b, p. 156
San Pedro, Calif.; Hilltop Quarry
Lower Pleistocene
Phlegonis, Bulimulus (Naesiotus) ustulatus: Dall and Ochsner
Paratypes
Dall and Ochsner, 1928, p. 160
Galapagos; Charles Island, 1650’ elev.
physispira, Brannerillus: Hannibal Holotype
Hannibal, 1912b, p. 191, pl. 8, fig. 28. Also in Taylor and Smith,
1971, figs. 49, 53
Kettleman Hills, Calif.; gulch S of Medallion One Canyon, E flank of
Hills
Pliocene [Upper Pliocene, Tulare Fm, basal part, fide Taylor and
Smith, 1971, p. 313]
picta, Cochliopa: Pilsbry Paratypes
Pilsbry, 1910, p. 100
San Luis Potosi, Mexico; Coy River, near Tampamolon
picta, Tegula (Agathistoma): McLean Paratype
McLean, 1970c, p. 121-122
W of Manta, Ecuador; 0° 56’ 43” S, 80° 44’ 43” W, on exposed reef
at low tide R/V Velero III Sta. 403-35
pilsbryi, Oreohelix: Ferriss Paratypes
Ferriss, 1917, p. 102
Black Range, New Mexico; Chloride (Mineral Creek)
plectatus, Ficus: Waring Holotype
Waring, 1917, p. 83, pl. 12, fig. 8
Ventura Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm_ [Paleocene]
pleistocenensis, Eupleura muriciformis: Arnold Holotype
Arnold, 1903, p. 249, pl. 9, fig. 16
Los Angeles Co., Calif.; San Pedro, lumber yard
Pleistocene, upper San Pedro Fm
portolaensis, Fusus: Arnold Paratype
Arnold, 1908a, p. 385. Illustrated zz Arnold, 1909, Illust. 2, fig. 68
San Mateo Co., Calif.; 1/2 mile SW of Portola on Sausal Creek
Upper Miocene, Purisima Fm [Arnold’s specimen 1080]
442
537
9735
7986
7784
6182
10067a
10067b
10067¢
8669
9722
7868
7868a
7867
7867a
8587
8605
5447
9175
BuLLeETIN 300
praecursor, Columbella solidula: Arnold Plastoholotype
Arnold, 1903, p. 236, pl. 10, fig. 4
Los Angeles Co., Calif.; San Pedro, lumber yard
Pleistocene, upper San Pedro Fm _ [holotype USNM]
precursor, Typhis (Talityphis): Keen and Campbell
Plastoholotype
Keen and Campbell, 1964, p. 49, pl. 9, figs. 14, 18
Atlantico, Colombia; 6 km W of Puerto Columbia UC loc. S-8012
Upper Oligocene, Las Perdices Shale [Holotype UCMP No. 15083]
predistortus, Cantharus (Triumphis): Marks Paratype
Marks, 1951, p. 117
SW Ecuador; Daule Basin
Middle Miocene, Daule Fm
profundorum, Oreohelix yavapai: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1911, p. 182
Grand Canyon, Arizona; Specimen Cove, Bass Trail
pronotis, Monadenia fidelis: Berry Paratypes
Berry, 1931la, p. 122
Del Norte Co., Calif.; Point St. George, near Crescent City
protera, Hanetia dalli: Woodring ’ Paratypes
Woodring, 1964, p. 257
Panama; Transisthmian Highway, 9° 21’ + 335 m N, 79° 49° W
SU loc. 2611 = USGS loc. 16912
Middle Miocene, Gatun Fm, lower part
pterocladica, Tricolia affinis: Robertson Paratypes
Robertson, 1958, p. 264
Boynton Beach, Fla.
puertoricensis, Typhis (Talityphis): Warmke Holotype
Warmke, 1964, p. 1, pl. 1, figs. 3, 4
Puerto Rico; off Punta Cadena, N of Mayaguez, 33 fms
punctocostata, Puncturella: Berry Holotype
Berry, 1947b, p. 265, pl. 26, figs. 7-9
San Pedro, Calif.; near Second and Pacific Streets
Lower Pleistocene, Lomita Fm
punctocostata, Puncturella: Berry Paratype
Berry, 1947b, p. 265
San Pedro, Calif.; near Second and Pacific Streets
Lower Pleistocene, Lomita Fm
punctulum, Mistostigma: Berry Holotype
Berry, 1947b, p. 264, pl. 27, fig. 5
Santa Barbara, Calif.; Bath House Cliff
Upper Pliocene, Santa Barbara Fm
punctulum, Mistostigma: Berry Paratype
Berry, 1947b, p. 264
Santa Barbara, Calif.; Bath House Cliff
Upper Pliocene, Santa Barbara Fm
pusilla, Diodora: Berry Holotype
Berry, 1959, p. 109. Illustrated im Keen, 1971, p. 316, fig. 22
Guerrero, Mexico; off Acapulco, 6-10 fms
pycna, Olivella: Berry Holotype
Berry, 1935, p. 262, fig. 1
Marin Co., Calif.; Bolinas Bay, 3-4 fms
pycna, Olivella: Berry Paratype
Berry, 1935, p. 262
Marin Co.. Calif.; Bolinas Bay, 3-4 fms
quadrangulata, Nerita: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 183, pl. 23, fig. 1
Santa Barbara Co., Calif.; N of Gaviota Pass UCMP loc. B-6962
Eocene-Oligocene, Refugian stage, Gaviota Fm
9175a
9175b
9175¢
7625
6591
9579
498
5811
8567
428
7006
161
146
596
597
598
STANFORD UNIVERSITY TypEs: SMITH 443
quadrangulata, Nerita: Weaver and Kleinpell Paratypes
Weaver and Kleinpell, 1963, p. 183
Santa Barbara Co., Calif.; N of Gaviota Pass UCMP loc. B-6962
Eocene-Oligocene, Refugian stage, Gaviota Fm
quimperensis, Perse olympicensis: Durham Paratype
Durham, 1944, p. 175, pl. 16, fig. 6
Jefferson Co., Wash.; Point Nill, Port Discovery SU loc. NP 151
Oligocene, Quimper Ss
ralphi, Puncturella: Berry Holotype
Berry, 1947b, p. 267, pl. 26, figs. 4-6
San Pedro, Calif.; near Second and Pacific Streets
Pleistocene, Lomita Fm
redondoensis, Pseudomelatoma semiinflata: Burch Paratype
Burchh ays) 1938, p. 21
Redondo Beach, Calif.; 25 fms
refulleri, Epitonium (Boreoscale) condoni: Durham Holotype
Durham, 1937, p. 497, pl. 57. fig. 3
Washington-Columbia Co., line, Nehalem River, near Vernonia, Ore.
SU loc. NP 1
Oligocene, Keasey Fm
regina, Doryssa rex: Pilsbry Paratype
Pilsbry in Baker, Fred, 1914, p. 651
Rio Jary, Brazil; Sao Antonio do Cachoeira
rejecta, Oliva: Burch and Burch Paratype
Burch and Burch, 1962, p. 166
Baja California, Mexico; La Paz, on tide flats
renaudi, Drillia: Arnold Plastoholotype
Arnold, 1903, p. 208, pl. 8, fig. 5
Los Angeles Co., Calif.; Deadman Island
Pleistocene, lower San Pedro Fm_ [holotype USNM]
renodata, Turritella pachecoensis: Merriam Plastoholotype
Merriam, 1941, p. 69, pl. 4, fig. 8
Simi Valley, Calif.; SE 1/4 Sec. 23, T 2 N, R 18 W UCMP loc.
ST
Paleocene, Martinez Fm [holotype UCMP 15315]
reticulata, Pseudoliva: Waring Holotype
Waring, 1914, p. 783. Also in Waring, 1917, p. 86, pl. 12, fig. 4 (as
Pseudoliva howardi Dickerson)
Ventura Co., Calif.; Martinez area of Simi Hills
Lower Eocene, Martinez Fm_ [Paleocene ]
reversa, Turritella: Waring Holotype
Waring, 1917, p. 88, pl. 12, fig. 15. Also i2 Merriam, 1941, p. 74,
DG Ay artes, 7
Ventura Co., Calif.; Calabasas sheet, Simi Hills
Eocene, Martinez Fm _ [Paleocene]
rex, Galeodea: Tegland Paratype
Tegland, 1931, p. 413, pl. 62, figs. 2, 3
Bainbridge Island, Wash.; Puget Sound, beach between S side of
entrance to Blakeley Harbor and Restoration Point SU loc. NP 103
Oligocene, Blakeley Fm
rex, Galeodea: Tegland Paratype
Tegland, 1931, p. 413, pl. 62, fig. 5
Bainbridge Island, Wash.; Puget Sound, beach between S side of
entrance to Blakeley Harbor and Restoration Point SU loc. NP 103
Oligocene, Blakeley Fm
rex, Galeodea: Tegland Paratype
Tegland, 1931, p. 413, pl. 62, fig. 4
Bainbridge Island, Wash.; Puget Sound, beach between S side of
entrance to Blakeley Harbor and Restoration Point SU loc. NP 103
Oligocene, Blakeley Fm
444
7803
8712
8094
6247
147
148
6185
7991
6176
6949
6225
7531
402
646
8644
6212
BuLLETIN 300
rhodacme, Achatina schweinfurthi: Pilsbry Paratype
Pilsbry, 1919, p. 74
Africa; Stanleyville, Belgian Congo
rhytodes, Turritella gatunensis: Woodring Paratypes
Woodring, 1957, p. 109
Canal Zone; W side Rio Chagres, NW of Gatun Dam
Miocene, middle Gatun Fm
ritteri, Anachis: Hertlein and Strong Paratypes
Hertlein and Strong, 1951a, p. 82
Off Port Guatulco, Mexico; 15° 44’ 28” N, 96° 07’ 51” W, 7 fms
ritteri, Trivia: Raymond Paratypes
Raymond, 1903, p. 85
Off San Pedro, Calif.; 50 fms
robustus, Gyrodes: Waring Syntype
Waring, 1917, p. 84, pl. 13, fig. 11
Ventura Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm_ [Paleocene]
robustus, Gyrodes: Waring Syntype
Waring, 1917, p. 84, pl. 13, fig. 12 :
Ventura Co., Calif.; Martinez area, Simi Hills
Lower Eocene, Martinez Fm_ [Paleocene]
rochai, Bulimulus (Rhinus): Baker Paratypes
Baker, 1914, p. 636
Ceara, Brazil; Ceara-Mirim
roigi, Conus (Leptoconus): Marks Paratype
Marks, 1951, p. 140
Ecuador; near Las Masas, Guayas Province
Lower Miocene, Subibaja Fm
rooseveltiana, Sonorella: Berry Paratypes
Berry, 1917a, p. 14
Gila Co., Arizona; Roosevelt
rosea, Nucella (?): Dall Paratype
Dall, 1872, p. 270
Simeonoff Island, Shumagin group, Alaska
roseobasis, Drillia: Pilsbry and Vanatta Syntype
Pilsbry and Vanatta, 1902, p. 558
Galapagos; Albemarle Island, Tagus Cove
rotundomontana, Chrysallida: Keen Holotype
Keen, 1943, p. 43, pl. 4, fig. 28
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
rotundus, Pugnellus: Waring Holotype
Waring, 1917, p. 67, pl. 9, fig. 10
near Ventura-Los Angeles Co. line, Calif.; Calabasas sheet, S of
Santa Monica Mts.
Upper Cretaceous, Chico Fm
ruginodosa, Ficus (Trophosycon) ocoyana: Grant and Gale
Paratype
Grant and Gale, 1931, p. 746, pl. 30, fig. 5
Los Angeles Co., Calif.; Elsmere Canyon
Lower Pliocene
rupicollina, Astraea (Uvanilla): Stohler Paratype
Stohler, 1959, p. 434
Baja California, Mexico; 8 miles SE of South Coronado Island, 70’
saccharodytes, Helminthoglypta proles: Berry Paratypes
Berry, 1938b, p. 46
Tulare Co., Calif.; Sugar Loaf Mt., 6000’ elev.
6197
8665
451
796
5418
5419
101
6625
5527
6191
9190
9191
9192
5369
STANFORD University Types: SMITH 445
salmonensis, Helicodiscus fimbriatus: Hemphill Paratypes
Hemphill zz Binney, 1890, p. 220
Salmon River, Idaho
salmonensis, Monadenia fidelis: Talmadge Paratype
Talmadge, 1954, p. 54
Siskiyou Co., Calif.; Wooley Creek, Salmon River
sanctaeclarae, Carinifex: Hannibal Holotype
Hannibal, 1909, p. 40. Illustrated in Hannibal, 1912b, p. 163, pl. 6, fig.
14a (as Pompholyx). Also in Taylor and Smith, 1971, figs. 54-56, 59
(as Helisoma)
Santa Clara Co., Santa Cruz Mts., Calif.; Los Gatos, near Is quarry
Pliocene, Santa Clara Lake Beds
sanctaecrucis, Fusus: Arnold Holotype
Arnold, 1908a, p. 372, pl. 33, fig. 3. Also zz Arnold, 1909, fig. 19, and
in Tegland, 1933, p. 166, pl. 12, fig. 2 (as Fusinus)
Santa Cruz Co., Calif.; Bear Creek, 4 miles above San Lorenzo River
Oligocene, San Lorenzo Fm
sanctaecrucis, Fusus: Arnold Paratypes
Arnold, 1908a, p. 372
Santa Cruz Co., Calif.; Bear Creek, 4 miles above San Lorenzo River
Oligocene, San Lorenzo Fm
sanctaemariae, Cancellaria: Carson Holotype
Carson, 1926, p. 57, pl. 3, fig. 5, as sanctae-mariae
Santa Barbara Co., Calif.; Santa Maria District, Fugler’s Point
Lower Pliocene, Fernando Fm
sanctijosephi, Lymnaea cubensis: Hannibal Holotype
Hannibal in Keep, 1911, p. 309, pl. 3, fig. 6, as sancti-josephi. Also
in Taylor and Smith, 1971, fig. 35 [as Bakerilymnaea bulimoides (Lea,
1841) ]
Santa Clara Co., Calif.; Calabasas Slough between Alviso and
Lawrence
sanesia, Odostomia (Amaura): Oldroyd Paratype
Oldroyd, T. S., 1924, p. 34
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
sanmarcosensis, Bulimulus: Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932b, p. 49
Gulf of California; San Marcos Island
sanmarcosensis, Turritella variata: Weaver and Kleinpell
Holotype
Weaver and Kleinpell, 1963, p. 185, pl. 24, fig. 4
Santa Barbara Co., Calif.; Camino Cielo UCMP loc. B-6940
Eocene, “Coldwater” Fm
sanmarcosensis, Turritella variata: Weaver and Kleinpell
Paratype
Weaver and Kleinpell, 1963, p. 185, pl. 24, fig. 2
Santa Barbara Co., Calif.; Camino Cielo UCMP loc. B-6940
Eocene, “Coldwater” Fm
sanmarcosensis, Turritella variata: Weaver and Kleinpell
Paratype
Weaver and Kleinpell, 1963, p. 185, pl. 24, fig. 3
Santa Barbara Co., Calif.; Camino Cielo UCMP loc. B-6940
Eocene, “Coldwater” Fm
santacruzana, Agasoma: Arnold Holotype
Arnold, 1908a, p. 379, pl. 34, fig. 7. Also iz Arnold, 1909, Illus. 2, fig.
44.
San Mateo Co., Calif.; 1/2 mile NNE of N end of Searsville Lake, on
hill N of road
Lower Miocene, Vaqueros Fm [Arnold’s specimen 1072]
446
9422
7126
7962
10335
0729
9730
7544
7544a
7544b
9519
6576
9237
9238
7783
9210
BuLLetIN 300
santacruzana, Turcicula: Arnold Paratypes
Arnold, 1908a, p. 373
pam Cruz Co., Calif.; San Lorenzo River, 3 miles above Boulder
Cree
Oligocene, San Lorenzo Fm
santana, Turritella inezana: Loel and Corey Plastoholotype
Loel and Corey, 1932, p. 259, pl. 59, fig. 13
eee Co., Calif.; Santa Ana Mts., W end Plano Trabuco UCMP
oc. 6128
Lower Miocene, Vaqueros Fm_ [holotype UCMP 31691]
santiagensis, Cancellaria (Bivetiella): Marks Paratype
Marks, 1949, p. 462
Ecuador; Esmeraldas Province, Angostura Cave, Santiago River
Lower Miocene, Angostura Fm
scalesiana, Naesiotus: Smith Paratype
Smith, A. G., 1972, pp. 17-19
Galapagos; Isla Santa Cruz, Harneman Farm, Scalesia Zone CAS
loc. 40021
scalpta, Streptacis: Knight _ Paratypes
Knight, 1931, p. 12
St. Louis Co., Missouri; St. Louis
Pennsylvanian, Harriet Fm, top of Labette Shale
scandix, Syrnola, Keen Holotype
Keen, 1943, p. 50, pl. 4, fig. 29
Kern Co., Calif.; Caliente Qd, small gully near center SW 1/4 Sec.
6, T 29S,R30E_ SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
scandix, Syrnola: Keen Paratype
Keen, 1943, p. 50, pl. 4, fig. 24
Kern Co., Calif.; Caliente Qd, small gully near center SW 1/4 Sec.
6, T 29 S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
scandix, Syrnola: Keen Paratype
Keen, 1943, p. 50, pl. 4, fig. 30
Kern Co., Calif.; Caliente Qd, small gully near center SW 1/4 Sec.
6, T 29S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
scelera, Odostomia (Chrysallida): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 30
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
schencki, Actaeon (Microglyphis): Berry Paratype
Berry, 1941, pp. 3-4
Los Angeles Co., Calif.; San Pedro, Hilltop Quarry
Lower Pleistocene, Lomita
schencki, Conus: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 194, pl. 27, fig. 8
Santa Barbara Co., Calif.; Canada de Santa Anita SU loc. 2091
Eocene, middle Gaviota Fm
schencki, Conus: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 194, pl. 27, fig. 9
Santa Barbara Co., Calif.; Canada de Santa Anita SU loc. 2091
Eocene, middle Gaviota Fm
schencki, Epitonium (Boreoscala) keaseyense: Durham Holotype
Durham, 1937, p. 498, pl. 57, fig. 14
Washington Co., Ore.; 3 miles S of Timber, Sec. 3, T 2N, RR 5 W
Oligocene, Keasey Fm
schencki, Galeodea: Weaver and Kleinpell Holotype
Weaver and Kleinpell, 1963, p. 189, pl. 25, fig. 16
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek UCMP loc.
B-6933
Eocene-Oligocene, Sacate-Gaviota Fm
9211
9723
7623
5711
7043
6983
6983a
7779
8628
6556
8345
7040
7038
8692
8513
8519
STANFORD UNIVERSITY TYPES: SMITH 447
schencki, Galeodea: Weaver and Kleinpell Paratype
Weaver and Kleinpell, 1963, p. 189, pl. 25, fig. 15
Santa Barbara Co., Calif.; Lompoc Qd, Nojoqui Creek UCMP loc.
B-6933
Eocene-Oligocene, Sacate-Gaviota Fm
schencki, Laevityphis (Laevityphis): Keen and Campbell
Holotype
Keen and Campbell, 1964, p. 53, pl. 9, figs. 16, 20
Atlantico, Colombia; Puerto Colombia
Upper Oligocene, Las Perdices Fm
schencki, Olequahia: Durham Holotype
Durham, 1944, p. 168, pl. 15, fig. 15
Washington Co., Ore.; Sec. 8, T 3 N, R4 W
Oligocene, Keasey Fm
schencki, Pleurotomaria (Entemnotrochus?): Hickman Holotype
Hickman, 1976a, p. 1095, pl. 1, figs. 1, 2, 11
Polk Co., Ore.; Dallas Qd, nr line between Secs. 11 & 12, T 8S, R 6
W; Oregon Portland Cement Company Quarry, 1-1/2 mi SW of Dal-
las SU loc. 1111
Eocene, Yamhill Fm, Rickreall Ls mbr
schencki, Turritella: Merriam Plastoholotype
Merriam, 1941, p. 81, pl. 10, fig. 10
Kern Co., Calif.; Tecuya Creek UCMP loc. A-1399
Upper Eocene, Tejon Fm [holotype UCMP 33945]
schencki, Turritella: Merriam Paratypes
Merriam, 1941, p. 81
Kern Co., Calif.; Tecuya Creek UCMP loc. A-1399
Upper Eocene, Tejon Fm
schereri, Helicina: Baker Paratypes
Baker, Fred, 1914, p. 625
Ceara-Mirim, Brazil
schilderiana, Cypraea tigris: Cate Paratype
Cate, 1961, p. 108
Oahu, Hawaii; Koko Head
scottiana, Monadenia fidelis: Berry Paratypes
Berry, 1940a, p. 11
Siskiyou Co., Calif.; Kelsey Creek, Scott River
scrippsae, Hemitoma: Durham Plastoholotype
Durham, 1950, p. 133, pl. 28, figs. 9, 14
Gulf of California; Carmen Island, Marquer Bay UCMP loc. A-3520
Upper Pliocene, Marquer Fm_ [holotype UCMP 30363]
scrippsensis, Turritella: Hanna Plastoholotype
Hanna, 1927, p. 308, pl. 49, fig. 10. Also im Merriam, 1941, p. 81, pl.
Ch aaver, 115)
San Diego Co., Calif.; La Jolla Qd UCMP loc. 5085
Eocene, Rose Canyon Fm_ [holotype UCMP 30904]
secondaria, Turritella andersoni lawsoni: Merriam
Plastoholotype
Merriam, 1941, p. 78, pl. 9, fig. 9
Ventura Co., Calif.; Llajas Canyon, Simi Valley UCMP loc. 7004
Eocene, “Domengine” Fm [holotype UCMP 33998]
seftoni, Ocenebra: Chace Paratype
Chace, 1958a, p. 331
Baja California, Mexico; Guadalupe Island, Melpomene Cove, 40
fms
semiusta, Mitra: Berry Holotype
Berry, 1957, p. 80
Santa Barbara Co., Calif.; off Point Conception, 15 m
sericeus, Capulus: Burch and Burch Holotype
Burch and Burch, 1961, p. 19, pl. 2, figs. 1, 2. Also im Keen, 1971,
p. 467, fig. 832
Sonora, Mexico; off Cabo Haro, Guaymas, 100 fms
448
6266
9514
8590
6175
7798
9921
5845
6233
112
141
9992
10278
6174
140
6523
6054
6055
BuL.etin 300
serrae, Turbonilla (Strioturbonilla): Dall and Bartsch Paratype
Dall and Bartsch, 1907, p. 497
Monterey, Calif.; 12 fms
sharonae, Lamellaria: Willett Paratype
Willett, 1939, p. 123
Anaheim Bay, Calif.
shaskyi, Cantharus: Berry Holotype
Berry, 1959, p. 111
Sonora, Mexico; probably S of Guaymas, from shrimp boats
shasta, Polygyra columbiana: Berry Paratype
Berry, 1921, p. 37
Shasta Co., Calif.; La Moine
shataii, Menestho (Menestho): Nomura Paratypes
Nomura, 1936, p. 36, as s-hataii
NE Honsyu, Japan; Siogama Bay
sheehani, Patella: Smith Plastoholotype
Smith, 1927, p. 108, pl. 96, figs. 28, 29
Shasta Co., Calif.; N fork Squaw Creek, 3 miles N of Kellys Ranch
Upper Triassic, Hosselkus Ls [holotype USNM 74175]
shepardi, Zonites: Hemphill Syntypes
Hemphill iz Binney, 1892, p. 167
Santa Catalina Island, Calif.; Avalon
shepardiana, Graphis: Dall Paratypes
Dall, 1919b, p. 342
San Pedro, Calif.; foot of Ash Street
shumanensis, Thais (Nucella): Carson Holotype
Carson, 1926, p. 56, pl. 3, fig. 1
Ventura Co. Calif.; Santa Maria District 1/2 mile N of Schuman, in
R.R. cut
Lower Pliocene, Fernando Fm
shumanensis, Thais (Nucella): Carson Paratype
Carson, 1926, p. 56, pl. 3, fig. 2
Ventura Co., Calif.; Santa Maria District, 1/2 mile N of Schuman
Lower Pliocene, Fernando Fm
shyana, Terebra: Bratcher and Burch Paratype
Bratcher and Burch, 1970a, p. 295
Colima, Mexico; off Manzanillo, 17-40 fms
shyorum, Epitonium (Epitonium): DuShane and McLean
Paratype
DuShane and McLean, 1968, p. 2
Colima, Mexico; Manzanillo, 12-13 fms
sierrana, Polygyra: Berry Paratype
Berry, 1921, p. 36
Siskiyou Co., Calif.; 2 miles N of Weed
simiensis, Turritella: Waring Holotype
Waring, 1917, p. 88, pl. 14, fig. 15. Also zz Merriam, 1941, p. 67, pl.
5, fig. 4 (as T. pachecoensis Stanton)
Ventura Co., Calif.; Martinez area, Simi Hills
Eocene, Martinez Fm_ [Paleocene]
sirius, Siphonaria: Pilsbry Paratypes
Pilsbry, 1894, p. 9
Sagami, Japan
skogsbergi, Turbonilla (Pyrgolampros): Strong Holotype
Strong, 1937, p. 54, pl. 4, fig. 3a
Monterey Bay, Calif.; 5 miles N of Monterey, 28 fms
skogsbergi, Turbonilla (Pyrgolampros): Strong Paratype
Strong, 1937, p. 54, pl. 4, fig. 3b
Monterey Bay, Calif.; 5 miles N of Monterey, 28 fms
6558
10295
6223
8100
9510
67
252
253
260
261
232
233
71
STANFORD UNIVERSITY TyPEs: SMITH 449
smithiana, Monadenia fidelis: Berry Paratype
Berry, 1940a, p. 14
Del Norte Co., Calif.; 3 miles below Hiouchi, N side Smith River
snodgrassi, Bulimulus: Dall Paratype
Dall, 1900a, p. 90
Galapagos; Hood Island
snodgrassi, Chlorostoma: Pilsbry and Vanatta Syntypes
Pilsbry and Vanatta, 1902, p. 557
Galapagos; Albemarle Island, Iguana Cove
socorroensis, Latirus: Hertlein and Strong Paratype
Hertlein and Strong, 1951b, p. 76
Clarion Island, off West Mexico
sonorana, Bursa californica: Berry Holotype
Berry, 1960, p. 118. Illustrated 7m Keen, 1971, p. 509, fig. 967
Sonora, Mexico; near Guaymas, from shrimp boats
sookensis, Acmaea mitra: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 171, pl. 35, figs. 2a, 2b
Vancouver Island, British Columbia, Canada: sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Calyptraea: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 168, pl. 36, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Calyptraea: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 168, pl. 36, fig. 2
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creek, Sooke SU loc. NP 129
Oligocene, Sooke Fm
sookensis, Crepidula: Clark and Arnold Holotype
Clark and Arnld, 1923, p. 166, pl. 35, figs. 5a, 5b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, Sooke SU loc. NP 129
Oligocene, Sooke Fm
sookensis, Crepidula: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 166, pl. 32, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, Sooke SU loc. NP 129
Oligocene, Sooke Fm
sookensis, Gadinia reticulata: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 157, pl. 35, fig. 3
Vancouver Island, British Columbia, Canada; Jordan River, sea
cliffs at mouth of Fossil] Creek, 2 miles W of Sherringham Point
SU loc. NP 130
Oligocene, Sooke Fm
sookensis, Gadinia reticulata: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 157, pl. 35, fig. 4
Vancouver Island, British Columbia, Canada; Jordan River, sea cliffs
at mouth of Fossil Creek, 2 miles W of Sherringham Point SU loc.
NP 130
Oligocene, Sooke Fm
sookensis, Goniobasis: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 164, pl. 32, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Kirby Creeks, Sooke
Oligocene, Sooke Fm
450
271
245
5347
8702
9743
8569
8668
197
7862
425
5380
BuL.etin 300
sookensis, Littorina: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 165, pl. 37, figs. 4a, 4b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
sookensis, Polinices (Ampullina?): Clark and Arnold Holotype
Clark and Arnold, 1923, p. 170, pl. 33, figs. 4a, 4b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
sorenseni, Haliotis: Bartsch Paratype
Bartsch, 1940, p. 50
Santa Barbara Co., Calif.; just S of Point Conception, 10 fms
spectabilis, Coronaria: Trechmann Paratypes
Trechmann, 1918, p. 187
New Zealand; Otago, Nugget Point (type 8048) ; Wairoa Gorge, Nel-
son (type 8049) ;
Triassic, Karnic
spelaea, ?Vitrea subrupicola: Dall Paralectotypes
Dall, 1895, pp. 27-28. Paralectotypes designated by Smith, 1957, p. 30
[referred to Pristiloma subrupicola spelaeum (Dall) ]
Calaveras Co., Calif.; Cave City
spermatia, Teinostoma (Idioraphe): Woodring Paratypes
Woodring, 1957, p. 69
Canal Zone; 3500’ SE of Gatun R.R. station
Miocene, Gatun Fm
sphoni, Mitra (Strigatella): Shasky and Campbell Holotype
Shasky and Campbell, 1964, p. 118, pl. 22, figs. 13, 14. Also im Keen,
1971, p. 642, fig. 1428
Sonora, Mexico; NW of Bahia Saladita, Guaymas, 2-15 m
sphoni, Olivella (Olivella): Burch and Campbell Paratype
Burch and Campbell, 1963, p. 124
Guaymas, Mexico; Bocochibampo Bay, 20 m
spirellum, Speleodiscoides: Smith Paratypes
Smith, 1957, p. 34
Amador Co., Cali.; Violin Cave, S fork Dry Creek
spissa, Olivella: Waring Holotype
Waring, 1917, p. 85, pl. 12, fig. 7
Ventura Co., Calif.; Simi Hills
Eocene, Martinez Fm [Paleocene]
squamulifer, Trophon: Gabb Plastoholotype
Gabb, 1866, p. 44. Illustrated zz Bormann, 1946, pl. 4, fig. 13 (as
Ocenebra)
Santa Barbara, Calif.
Pleistocene, Santa Barbara Fm [holotype UCMP 15459]
stanfordensis, Agasoma: Arnold Holotype
Arnold, 1908a, p. 384, pl. 35, fig. 5
Santa Clara Co., Calif.; Tusk Gully, 2.5 miles S of Mayfield
Upper Miocene [Arnold’s specimen 1087]
stanfordensis, Fusus (Priscofusus?): Arnold Holotype
Arnold, 1908a, p. 383, pl. 35, fig. 7. Also in Arnold, 1909, Illus. 2, fig.
55
Santa Clara Co., Calif.; near Frenchman’s Tower, on hill between
Tusk Gully and Madera Creek, 2.5 miles SSW of Mayfield
Upper Miocene, Temblor Fm_ [Arnold’s specimen 1081]
8507
5371
5372
5812
5133
6242
8705
8709
403
8570
7983
7081
6461
8663
532
STANFORD University Types: SMITH 451
stanfordiana, “Acmaea”: Berry Holotype
Berry, 1957, p. 76. Illustrated ix Keen, 1971, p. 325, fig. 51 (as Col-
lisella)
Sonora, Mexico; Pelican Point
stantoni, Chrysodomus: Arnold Paratype
Arnold, 1908a, p. 386, pl. 37, fig. 4. Also zz Arnold, 1909, Illus. 2,
fig. 65
San Mateo Co., Calif.; 7/8 mile E of Anto Nuevo Point
Pliocene, upper Purisima Fm [Arnold’s specimen 1088]
stantoni, Chrysodomus: Arnold Paratype
Arnold, 1908a, p. 386
San Mateo Co., Calif.; 7/8 mile E of Ano Nuevo Point
Pliocene, upper Purisima Fm
starksi, Doryssa: Pilsbry Paratype
Pilsbry iz Baker, Fred, 1914, p. 652
Rio Iriri, Brazil
starri, Crassispira: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 626, pl. 21, fig. 7
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
stearnsi, Ocinebra: Hemphill Paratypes
Hemphill, 1911, p. 100
Monterey, Calif.
stemonium, Teinostoma (Pseudorotella): Woodring Paratypes
Woodring, 1957, p. 71
Canal Zone; highway 1.6 km NE of boundary SU loc. 2656
Miocene, Gatun Fm
stenopa, Natica (Naticarius): Woodring Paratype
Woodring, 1957, p. 85
Canal Zone; Mount Hope, W side of Panama R.R.
Miocene, upper Gatun Fm
stephensae, Amphithalamus: Bartsch Paratypes
Bartsch, 1927, p. 27
Baja California, Mexico; Magdalena Bay
steveni, Olivella (Olivella): Burch and Campbell Paratype
Burch and Campbell, 1963, p. 125
Baja California, Mexico; 2 miles S of Aguachale
stevensoni, Anachis (Costoanachis): Marks Paratypes
Marks, 1951, p. 112
Ecuador; Zacachun corehole, 80-90’ depth, Progreso area
Lower Miocene, Subibaja Fm
stewarti, Turritella uvasana: Merriam Plastoholotype
Merriam, 1941, p. 95, pl. 16, fig. 7
Cowlitz Co., Wash.; Coal Creek UCMP loc. 7167
Eocene, Cowlitz Fm [holotype UCMP 33888]
stimpsoni, Truncatella: Stearns Paratypes
Stearns, 1872, p. 249
San Diego, Calif.; False Bay [Mission Bay]
stocki, Nassarius: Kanakoff Paratypes
Kanakoff, 1956, p. 110
Los Angeles Co., Calif.; 1/2 mile S of Humphreys R.R. station
Pliocene, Pico Fm
stokesi, Paludestrina: Arnold Paratype
Arnold, 1903, p. 305
Los Angeles Co., Calif.; San Pedro
Pleistocene, upper San Pedro Fm
9498
7980
8091
6271
8706
9920
9717
7796
7858
7858a
6189
7966
6187
6186
5810
6257
5910
BuLLeTIN 300
striata, Kogomea: Laseron Paratypes
Laseron, 1957, p. 294
Queensland, Australia; Swain Reef, Michmaelmas Cay
striatocostata, Strombina: Marks Paratypes
Marks, 1951, p. 110
SW Ecuador; Daule Basin, near Pedro Carbo
Middle Miocene, Daule Fm
strohbeeni, Cymatosyrinx: Hertlein and Strong Paratype
Hertlein and Strong, 1951b, p. 77
Baja California, Mexico; off Cape San Lucas
strongi, Odostomia (Evalea): Bartsch Paratypes
Bartsch, 1927, p. 19
Santa Catalina Island, Calif.; Isthmus Cove
strongylus, Solariorbis (Solariorbis): Woodring Paratypes
Woodring, 1957, p. 75
Canal Zone; highway 1.6 km NE of boundary SU loc. 2656
Miocene, lower Gatun Fm
stuarti, Patella: Smith Plastoholotype
Smith, 1927, p. 108, pl. 91, fig. 18 :
Shasta Co., Calif.; Bear Cove, Brock Mt., between Squaw Creek and
Pit River
Upper Triassic, Hosselkus Ls [holotype USNM 74149]
subactum, Crucibulum: Berry Holotype
Berry, 1963, p. 144. Illustrated iz Keen, 1971, p. 465, fig. 831
Sinaloa, Mexico; off Teacapan, 25-35 fms
subcinctella, Syrnela (Syrnola): Nomura Paratypes
Nomura, 1936, p. 15
NE Honsyu, Japan; Siogama Bay
succinea, Lacuna: Berry Holotype
Berry, 1953b, p. 411, fig. 4
San Pedro, Calif.
succinea, Lacuna: Berry Paratype
Berry, 1953b, p. 141
San Pedro, Calif.
suprapunctatus, Drymaeus linostoma: Baker Paratype
Baker, Fred, 1914, p. 638
Matto Grosso, Brazil, Madeira-Mamoré R.R., 284 km above Porto
Velho
sursalta, Cancellaria (Cancellaria): Marks Paratype
Marks, 1949, p. 461
Ecuador; Guayas Province, Zacachun corehole, 140’-150’ depth
Lower Miocene
suturalis, Bulimulus (Rhinus) rochai: Baker Paratypes
Baker, Fred, 1914, p. 637
Ceara-Mirim, Ceara, Brazil
taipuensis, Bulimulus (Rhinus) rochai: Baker Paratype
Baker, 1914, p. 636
Taipu, Brazil; 46 km from Natal
tapajozensis, Doryssa transversa: Pilsbry Paratype
Pilsbry iz Baker, Fred, 1914, p. 649
Rio Tapajoz, Brazil
taravali, Melanella (Balcis): Bartsch Paratypes
Bartsch, 1917, p. 328
Baja California, Mexico; Point Abreojos
taylori, Tegula pulligo: Oldroyd Holotype
Oldroyd, I. S., 1925, p. 171, pl. 20, figs. 1, 2. Also iz Oldroyd, I. S.,
1927, p. 179, pl. 91, figs. 3, 6
Off N end Vancouver Island, British Columbia, Canada; Hope Island
5911
10277
7052
7533
419
8043
6041
9729
8258
5520
510
6145
8672
8750
8750a
STANFORD UNIVERSITY J yPEs: SMITH 453
taylori, Tegula pulligo: Oldroyd Paratype
Oldroyd, I. S., 1925, p. 171
Off N end Vancouver Island, British Columbia, Canada; Hope Island
tehuanarum, Amaea (Scalina): DuShane and McLean Paratype
DuShane and McLean, 1968, pp. 4-6
Gulf of Tehuantepec, Mexico; 15° 58’ N, 95° 00’ W, 33-38 fms
tejonensis, Turritella: Merriam Plastoholotype
Merriam, 1941, p. 81, pl. 11, fig. 7
Kern Co., Calif.; Grapevine Canyon UCMP loc. 452
Eocene, Tejon Fm _ [holotype UCMP 15190]
temblorensis, Cylichna: Keen Holotype
Keen, 1943, p. 44, pl. 4, figs. 13, 14
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29S, R30 E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
temblorensis, Turritella: Wiedey Paratype
Wiedey, 1928, p. 122, pl. 11, fig. 9
Los Angeles Co., Calif.; Santa Monica Mts., in small canyon trending
W from head of Dry Canyon, at base of E-W divide, 2 miles § of
Calabasas
Middle Miocene, Temblor Fm
tenuissima, Volvulella: Willett Paratypes
Willett, 1944b, p. 71
Los Angeles Co., Calif.; off Redondo Beach, 75 fms
tenuistriata, Urocoptis: Aguayo Paratypes
Aguayo, 1932b, p. 96
Madruga, Havana, Cuba
teramachii, Typhis (Typhina) Keen and Campbell Plastoholotype
Keen and Campbell, 1964, p. 48, pl. 8, figs. 10-11
Off Kii, Japan, 100 m_ [holotype in private collection of Mr. Akibumi
Teramachi, Kyoto, Japan]
tersa, Balcis (Balcis): Berry Paratype
Berry, 1954b, p. 261
Los Angeles Co., Calif.; San Pedro, Hilltop Quarry
Lower Pleistocene, Lomita Fm.
tersa, Odostomia (Evalea): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 31
Los Angeles Co., Calif.; San Pedro, Nob Hill Cut
Pleistocene
teslaensis, Cerithium ?: Hanna Holotype
Hanna, 1924, p. 162. Illustrated in Wiedey, 1929b, p. 25, pl. 1, fig. 6
(as Cerithium branneri Hall and Ambrose) [Renamed by Hanna;
originally described as Cerithium branneri Hall and Ambrose, 1916,
p. 70]
Alameda Co., Calif.; 1 mile N 20° W of Tesla and Corral Hollow
Upper Cretaceous, middle Chico Fm
tetrodon, Ashmunella: Pilsbry and Ferriss Paratypes
Pilsbry and Ferriss, 1915b, p. 15
Socorro Co., New Mexico; Mogollon Mts., Dry Creek Canyon
thalassicola, Tricolia: Robertson Paratypes
Robertson, 1958, p. 271
Great Abaco Island, Bahamas; North Point, Elbow Cay
tholia, Dendropoma: Keen and Morton Holotype
Keen and Morton, 1960, p. 41, pl. 3, figs. 4, 5
Mozambique, East Africa; Inhaca Island, Lorenzo Marques
tholia, Dendropoma: Keen and Morton Paratypes
Keen and Morton, 1960, p. 41, pl. 3, fig. 6
Mozambique, East Africa; Inhaca Island, Lorenzo Marques
454
10068
10069
5528
8658
8657
142
8099
6158
7166
8680
8673
8679
5367
9511
6578
6578a
8703
BuLLetTIN 300
thompsoni, Eupleura: Woodring Paratypes
Woodring, 1959, pp. 218-220
Panama; Transisthmian Highway, 9° 21’ + 335 m N, 79° 49° W SU
loc. 2611 = USGS loc. 16912
Middle Miocene, Gatun Fm, lower middle part
timessa, Odostomia (Amaura): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 35
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
tingitana, Gibbula: Pallary Paratypes
Pallary, 1902a, p. 315
Tanger, Morocco, on stones [Tangier]
tingitana, Nassa: Pallary Paratype
Pallary, 1901, p. 226
Tanger, Morocco, 12-21 m_ [Tangier]
titan, Trachytriton: Waring Holotype
Waring, 1917, p. 87, pl. 14, fig. 18
Ventura Co., Calif.; Martinez area, Simi Hills, Calabasas sheet
Lower Eocene, Martinez Fm [Paleocene]
togatum, Epitonium (Cirsotrema): Hertlein and Strong Paratype
Hertlein and Strong, 1951b, p. 89
Near Manzanillo, Mexico; 19° 04’ N, 104° 22’ W, 30 fms
tooelensis, Oreohelix strigosa depressa: Henderson and Daniels
Paratypes
Henderson and Daniels, 1916, p. 323
6 miles NE of Tooele, Utah
topangensis, Turritella ocoyana: Merriam Plastoholotype
Merriam, 1941, p. 115, pl. 30, fig. 1
Santa Monica Mts., Calif.; Malibu Canyon, Mesa Peak UCMP loc.
A-556
Miocene, Topanga Fm_ [holotype UCMP 31648]
totomiensis, Siphonalia tonohamaensis: Makiyama Paratypes
Makiyama, 1941, p. 80
Shizuoka Prefecture, Japan; Ugari, near Fukuroi
Pliocene, Hosoya Fm
totomiensis, Thiara: Makiyama Paratype
Makiyama, 1927, p. 66
Shizuoka Prefecture, Japan; Dainiti
Pliocene, Dainitian
totomiensis, Turris (Gemmula): Makiyama Paratypes
Makiyama, 1931b, p. 46
Pliocene, Hosoya (Kakegawa)
trancosana, Thais: Arnold Holotype
Arnold, 1908a, p. 388, pl. 36, fig. 3. Also zz Arnold, 1909, Illus. 2,
fig. 74
Santa Clara Co., Calif.; 2.5 miles SSW of Stanford University; ditch
between Felt Lake and Los Trancos Creek
Upper Pliocene, Merced Fm [Arnold’s specimen 1082]
tricoronis, Murex (Murex): Berry Holotype
Berry, 1960, p. 119. Illustrated in Keen, 1971, p. 514, fig. 978
Baja California, Mexico; 1 mile off Cedros Village, Cedros Island;
40 fms
tridesmia, Clathurella (Glyphostoma): Berry Paratypes
Berry, 1941, p. 8
Los Angeles Co., Calif.; San Pedro, Hilltop Quarry
Lower Pleistocene, Lomita Fm
trochalum, Teinostoma (Idoraphe) angulatum: Woodring
Paratypes
Woodring, 1957, p. 70
Canal Zone; 1.6 km NE of boundary on highway SU loc. 2656
Miocene, Gatun Fm
5526
10282
5901
5902
7238
5903
8510
9509
7756
7797
7757a
254
STANFORD UNIVERSITY TyrEs: SMITH 455
trochilia, Odostomia (Amaura): Oldroyd Paratypes
Oldroyd, T. S., 1924, p. 34
Los Angeles Co., Calif.; San Pedro, Nob Hill cut
Pleistocene, lower San Pedro Fm
tumida, Drillia (Drillia):; McLean and Poorman Paratype
McLean and Poorman, 1971, p. 97
Jalisco, Mexico; Banderas Bay, 20° 40’ N, 105° 25’ W, 20-40 fms
turneri, Viviparus (Callina): Hannibal Syntype
Hannibal, 1912b, p. 193, pl. 8, fig. 31
Silver Peak Range, Nevada
“Eocene,” Truckee beds [upper Miocene-lower Pliocene, Esmeralda
Fm, fide Taylor and Smith, 1971, p. 311]
turneri, Viviparus (Callina): Hannibal Paratype
Hannibal, 1912b, p. 193. Also zx Taylor and Smith, 1971, figs. 28, 29
(as Bellamya)
Silver Peak Range, Nevada
“Eocene,” Truckee beds [upper Miocene-lower Pliocene, Esmeralda
Fm fide Taylor and Smith, 1971, p. 311]
Turritella sp. B: Schenck and Keen Holotype
Schenck and Keen, 1940, pl. 27, figs. 5, 6
Ventura Co., Calif.; 1 mile SE of Matilija, on S wall of Kennedy
Canyon, 1150’ contour, 1400’ due W of Ventura River
Eocene, “Coldwater” Fm
turveri, Haliotis fulgens: Bartsch Paratype
Bartsch, 1942, p. 57
Baja California, Mexico; Magdalena Bay
tyrianthina, Acanthina: Berry Holotype
Berry, 1957, p. 78. Illustrated im Keen, 1971, p. 552, fig. 1085
Baja California, Mexico; Magdalena Bay, Man-of-War Cove
tyrianthina, Neosimnia vidleri: Berry Holotype
Berry, 1960, p. 118
Sonora, Mexico; Puerto Penasco, Cholla Cove
usanium, Serratocerithium: Compton Holotype
Compton, 1944, p. 466, pl. 78, figs. 3, 6
Los Angeles Co., Calif.; ridge E of Santa Ynez Canyon, 4 miles E of
road in canyon, along fire road just W of top of ridge. Santa Monica
Mts. SU loc. 2691
Paleocene, Martinez Fm
usanium, Serratocerithium: Compton Paratype
Compton, 1944, p. 466, pl. 78, fig. 5
Los Angeles Co., Calif.; Santa Monica Mts., ridge E of Santa Ynez
Canyon, 4 miles E of road in canyon, along fire road just W of top of
ridge
Paleocene, Martinez Fm
usanium, Serratocerithium: Compton Paratype
Compton, 1944, p. 466
Los Angeles Co., Calif.; Santa Monica Mts., ridge E of Santa Ynez
Canyon, 4 miles E of road in canyon, along fire road just W of top of
ridge
Paleocene, Martinez Fm
vancouverensis, Acmaea persona: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 172, pl. 35, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU Joc.
NP 129
Oligocene, Sooke Fm
456
5912a
5912b
284
244
279
272
293
5838
BULLETIN 300
vancouverensis, Acteon punctocoelata: Oldroyd Syntype
Oldroyd, 1927, p. 25, pl. 1, fig. 19
Vancouver Island, British Columbia, Canada; Nanaimo, Departure
Bay, off Brandon Island, 10-15 fms
vancouverensis, Acteon punctocoelata: Oldroyd Syntype
Oldroyd, I. S., 1927, p. 25, pl. 1, fig. 20. Also in Oldroyd, I. S., 1924,
pl. 1, fig. 9 (as Acteon punctocoelata)
Vancover Island, British Columbia, Canada; Nanaimo, Departure
Bay, Brandon Island, 10-15 fms
vancouverensis, Bursa: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 163, pl. 37, figs. 1a, 1b. Also iz Smith,
1970, pl. 48, fig. 3 [as Mediargo mathewsonii (Gabb) ]
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Calyptraea (Galerus) mammillaris:
Clark and Arnold Holotype
Clark and Arnold, 1923, p. 167, pl. 36, figs. 3a, 3b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Leptothyra: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 173, pl. 37, figs. 3a, 3b
Vancouver Island, British Columbia, Canada; Jordan River, sea
cliffs at mouth of Fossil Creek, 2 miles W of Sherringham Point
SU loc. NP 130
Oligocene, Sooke Fm
vancouverensis, Megathura: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 173, pl. 34, figs. 3a, 3b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Polinices (Neverita) recluziana:
Clark and Arnold Holotype
Clark and Arnold, 1923, p. 169, pl. 33, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
vancouverensis, Stagnicola bulimoides: Baker Holotype
Baker, 1939, p. 144
Vancouver Island, British Columbia, Canada; hospital at Nanaimo
vancouverensis, Stagnicola bulimoides: Baker Paratypes
Baker, 1939, p. 144
Vancouver Island, British Columbia, Canada; hospital at Nanaimo
vaquerosensis, Purpura: Arnold Holotype
Arnold, 1907c, p. 427, pl. 52, figs. la, 1b
Monterey Co., Calif.; Lynch Mt.
Lower Miocene, Vaqueros Fm
velascoensis, Emarginula: Shasky Holotype
Shasky, 1961, p. 18, pl. 4, figs. 1-3. Also im Keen, 1971, p. 310, fig. 7
Gulf of California; off Isla Monserrate, 40-80 fms
verrucosa, Tegula (Agathistoma): McLean Paratype
McLean, 1970b, pp. 122-123
Canal Zone, Panama; Palo Seco, 8° 55’ N, 79° 34’ W, rocky intertidal
237
238
239
9506
8662
6169
9720
792
8327
5818
7785
462
6527
7530
STANFORD UNIVERSITY TYPES: SMITH 457
victoriana, Acmaea: Clark and Arnold Holotype
Clark and Arnold, 1923, p. 172, pl. 34, figs. 1a, 1b
Vancouver Island, British Columbia, Canada; sea cliffs 1-1.5 miles W
of Owens Point, Port San Juan SU loc. NP 134
Oligocene, Sooke Fm
victoriana, Acmaea: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 172, pl. 34, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; sea cliffs 1-1.5 miles
W of Owens Point, Port San Juan SU loc. NP 134
Oligocene, Sooke Fm
victoriana, Acmaea: Clark and Arnold Paratype
Clark and Arnold, 1923, p. 172, pl. 34, fig. 4
Vancouver Island, British Columbia, Canada; sea cliffs 1-1.5 miles
W of Owens Point, Port San Juan SU loc. NP 134
Oligocene, Sooke Fm
viridicolor, Cypraea cernica: Cate Holotype
Cate, 1962, pp. 175-177, pl. 40, fig. 1
Western Australia; Northwest Cape, Viaming Head
viadimiri, Kelletia: Kanakoff Paratypes
Kanakoff, 1954, pp. 114-117
Los Angeles Co., Calif.; 1/2 mile S of Humphreys R.R. station
Pliocene, Pico Fm
walcottiana, Sonorella: Bartsch Paratypes
Bartsch, 1903, p. 103
San Diego Co., Calif.; Palm Springs [= Sonorella wolcottiana,
Bartsch em.]
walkeri, Knefastia: Berry Holotype
Berry, 1958a, p. 87. Illustrated iz Keen, 1958b, p. 447, figs. 728a
Gulf of California; off Puerto Refugio, Angel de la Guarda Island
wardi, Leucosyrinx clallamensis: Tegland Paratype
Tegland, 1933, p. 124, pl. 10, fig. 8
Bainbridge Island, Wash.; beach between S side of entrance to Blake-
ley Harbor and Restoration Point SU loc. NP 103
Upper Oligocene, Blakeley Fm
warrenae, Megalacron tabarensis: Clench and Turner Paratypes
Clench and Turner, 1964, p. 43
Off New Ireland, Tanga Group; Boang Island, Bismarck Archi-
pelago
wasatchensis, Lymnaea stagnalis: Baker Paratypes
Baker, Frank, 1911, p. 152
Near Salt Lake City, Utah
wasatchensis, Patula strigosa: Hemphill Paratypes
Hemphill zz Binney, 1886, p. 34
Wasatch Mts., near Ogden, Utah; among quartzite boulders, 4500’
elev.
washingtonianus, Viviparus: Hannibal Holotype
Hannibal, 1912b, p. 194, pl. 8, fig. 32. Also im Taylor and Smith,
1971, figs. 26, 30 (as Bellamya)
Wash.; Olequa Creek, 2 miles N of Little Falls
Eocene [Late Eocene, Cowlitz Fm, fide Taylor and Smith, 1971, p.
311]
watermani, Olivella: McGinty Paratype
McGinty, 1940, p. 6
Off Palm Beach, Fla.; 80 fms
watsonae, Anachis: Keen Holotype
Keen, 1943, p. 42, pl. 4, figs. 1, 2
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29 S, R30 E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
458
8661
6512
7535
473
9710
9989
461
465
466
8604
7786
5125
5126
5127
5128
5129
5130
BuLLeTIN 300
weyersi, Cerithidea (Aphanistylus): Dautzenberg Paratype
Dautzenberg, 1899, p. 8
W coast of Sumatra, near Indrapoera River
wheatlandensis, Siphonalia bicarinata: Clark and Anderson
Paratype
Clark and Anderson, 1938, p. 952
Yuba Co., Calif.; Dry Creek, 6 miles NE of Wheatland
Upper Eocene-Lower Oligocene, Wheatland Fm
whitei, Ferminoscala: Keen Holotype
Keen, 1943, p. 46, pl. 4, figs. 32, 33
Kern Co., Calif.; Caliente Qd, in small gully near center SW 1/4
Sec. 6, T 29 S,R30E SU loc. 2121
Miocene, Temblor Fm, Round Mountain Silt
whitei, Valvata: Hannibal Holotype
Hannibal, 1910, p. 107. Illustrated in Taylor and Smith, 1971, figs. 45,
46, 50
Oregon, near Summer Lake
Quaternary, Upper Lahontan [Pliocene, probably Blancan, fide Tay-
lor and Smith, 1971] ;
willetti, Antiplanes (Rectiplanes): Berry Holotype
Berry, 1953b, p. 419, pl. 29, fig. 2
Alaska; Forrester Island, 50 fms
willetti, Turritella: McLean Paratype
McLean, 1970a, p. 312
Colima, Mexico; Manzanillo, Santiago Bay, 19° 06’ N, 104° 23’ W,
7-12 fms
williamsi, Pyrgulopsis: Hannibal Holotype
Hannibal, 1912b, p. 189, pl. 8, fig. 29a. Also im Taylor and Smith,
1971, figs. 24, 25
San Joaquin Valley, Calif.; Lost Hills, Martin and Dudleys Oilwell
[SE 1/4 Sec. 32, T 26 S, R 21 E]
Pliocene [San Joaquin Fm, fide Taylor and Smith, 1971, p. 312]
williamsi, Pyrgulopsis: Hannibal Paratype
Hannibal, 1912b, p. 189, pl. 8, fig. 29b
San Joaquin Valley, Calif.; Lost Hills, SE 1/4 Sec. 32, T 26 S, R 21E
Pliocene [San Joaquin Fm, fide Taylor and Smith, 1971, p. 312]
williamsi, Pyrgulopsis: Hannibal Paratype
Hannibal, 1912b, p. 189, pl. 8, fig. 29c
San Joaquin Valley, Calif.; Lost Hills, SE 1/4 Sec. 32, T 26 S, R 21 E
Pliocene [San Joaquin Fm, fide Taylor and Smith, 1971, p. 312]
williamsi, Woodbridgea: Berry Holotype
Berry, 1953b, p. 422, fig. 8
Baja California, Mexico; off Cedros Village, Cedros Island, 25 fms
winslowae, Arena: Pilsbry and Lowe Paratypes
Pilsbry and Lowe, 1932a, p. 86
Taboga Island, Panama
wittichi, Thais: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 633, pl. 18, fig. 3
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
wittichi, Thais: Hertlein and Jordan Paratypes
Hertlein and Jordan, 1927, p. 633
Baja California, Mexico; Arroyo San Ignacio, 8 km SW of San
Ignacio SU loc. 66
Miocene, Isidro Fm
5894
5135
6169
6579
8520
8711
8676
5836
7807
8262
6572
6236
8650
8646
7847
STANFORD UNIVERSITY [TyPES: SMITH 459
wittichi, Turritella: Hertlein and Jordan Holotype
Hertlein and Jordan, 1927, p. 635, pl. 21, fig. 3. Also im Merriam,
1941, p. 114, pl. 29, fig. 1
Baja California, Mexico; on trail from Arroyo Mesquital to La Puri-
sima, in Turritella bed above San Gregorio Lagoon SU loc. 59
Miocene, Isidro Fm
wittichi, Turritella: Hertlein and Jordan Paratype
Hertlein and Jordan, 1927, p. 635
Baja California, Mexico; on trail from Arroyo Mesquital to La Puri-
sima, in Turritella bed above San Gregorio Lagoon SU loc. 59
Miocene, Isidro Fm
wolcottiana, Sonorella: Bartsch Paratypes
Bartsch, 1903, p. 103
San Diego Co., Calif.; Palm Springs [emended from S. walcottiana]
woodfordi, Mitromorpha barbarensis: Berry Paratype
Berry, 1941, p. 10
Los Angeles Co., Calif.; San Pedro, Hilltop Quarry
Lower Pleistocene, Lomita Fm
xavieri, Colubraria: Campbell Holotype
Campbell, 1961a, p. 141, pl. 10, figs. 7, 8. Also im Keen, 1971, p. 512,
fig. 974
Sonora, Mexico; Guaymas, 2 miles W of Cabo Haro, 100 fms
xena, Neverita (Glossaulax) reclusiana: Woodring Paratypes
Woodring, 1957, p. 92
Canal Zone; highway 1.7 km SW of Sabanita SU loc. 2611
Miocene, lower Gatun Fm
yokoyamai, Asthenotoma: Makiyama Paratypes
Makiyama, 1927, p. 95
Shizuoka Prefecture, Japan; Dainiti
Pliocene, Dainiti /m
yrekaensis, Goniobasis: Henderson Paratypes
Henderson, 1935, p. 97
Shasta River, below Yreka, Calif.; 4 miles above river mouth
zambiensis, Thapsia: Pilsbry Paratypes
Pilsbry, 1919, p. 237
Zambi, Belgian Congo
zeteki, Epitonium: Dall Paratype
Dall, 1917b, p. 486. Illustrated iz Keen, 1971, p. 428, fig. 632 left
Near Panama City, Panama
zizyphus, Clavus (Crassispira): Berry Paratype
Berry, 1940b, p. 152
Los Angeles Co., Calif.; San Pedro, Hilltop Quarry
Lower Pleistocene
POLYPLACOPHORA (AMPHINEURA)
californiensis, Ischnochiton (Lepidozona): Berry Paratype
Berry, 1931b, p. 255
San Diego Co., Calif.; near Scripps Institution, La Jolla
circumsenta, Stenoplax: Berry Paratype
Berry, 1956a, p. 72
Baja California, Mexico; Scammon Lagoon, W of Isla Concha
crossota, Nuttallina: Berry Paratype
Berry, 1956a, p. 71. Illustrated iz Keen, 1958, p. 528, fig. 49
Sonora, Mexico; W end of Puerto Penasco
heathiana, Stenoplax (Stenoradsia): Berry Holotype
Berry, 1946a, p. 161, pl. 4, fig. 8
Monterey Co., Calif.; Pacific Grove, shoreline
281
6240
6239
6238
6273
8647
8648
6237
818
819
815
BuLtetTin 300
isoglypta, Stenoplax: Berry Paratype
Berry, 1956a, p. 72
Isabel Island, Peru
keepiana, Lepidochitona: Berry Paratype
Berry, 1948, p. 14
lioplax, Oligochiton: Berry Holotype
Orange Co., Calif.; Newport
Berry, 1922, p. 431, pl. 1, figs. 5, 6
Vancouver Island, British Columbia, Canada; sea cliffs between
mouths of Muir and Coal Creeks, W of Otter Point, Sooke SU loc.
NP 129
Oligocene, Sooke Fm
lioplax, Oligochiton: Berry Paratype
Berry, 1922, p. 431, pl. 1, figs. 3, 4
Vancouver Island, British Columbia, Canada; Sooke, sea cliffs be-
tween mouths of Coal and Muir Creeks, W of Otter Point SU loc.
NP 129
Oligocene, Sooke Fm
oldroydi, Lepidopleurus (Leptochiton): Dall ~ Paratypes
Dall, 1919a, p. 560
San Pedro, Calif.
percrassus, Lepidopleurus (Oldroydia): Dall Paratypes
Dall, 1894, p. 90
Santa Barbara Channel, off San Pedro, Calif.; 75 fms
semiliratus: Dendrochiton: Berry Holotype
Berry, 1927, p. 160, pl. 13, figs. 1, 2
Vancouver Island, British Columbia, Canada; Nanaimo, Departure
Bay
semiliratus, Dendrochiton: Berry Paratype
Berry, 1927, p. 160
Vancouver Island, British Columbia, Canada; Nanaimo, Departure
Bay
sonorana, Stenoplax (Maugerella) conspicua: Berry Paratypes
Berry, 1956a, p. 73. Illustrated iz Keen, 1958, p. 528, fig. 47
Sonora, Mexico; W end Puerto Penasco
subtilis, Lepidozona: Berry Paratypes
Berry, 1956a, p. 74. Illustrated in Keen, 1958, p. 526, fig. 42
Sonora, Mexico; W end Puerto Penasco
willetti, Ischnochiton (Lepidozona): Berry Paratypes
Berry, 1917b, p. 236
Forrester Island, Alaska
BRACHIOPODA
adairensis, Productus (Marginifera): Drake Syntype
Drake, 1898, p. 402, pl. 9, figs. 1, 3
Adair, Okla.; 5 miles SE, 7 miles E of town
Carboniferous, Boston Fm
adairensis, Productus (Marginifera): Drake Syntype
Drake, 1898, p. 402, pl. 9, fig. 2
Adair, Okla.; 5 miles SE, 7 miles E of town
Carboniferous, Boston Fm
cherokeensis, Productus: Drake Holotype
Drake, 1898, p. 404, pl. 9, figs. 4, 5
Adair, Okla.; 5 miles SE of town
Carboniferous, Boston Fm
9929
7776
5860
5857
9926
9928
144
5376
242
243
7778
5858
5859
STANFORD UNIVERSITY TYPES: SMITH 461
hamiltonense, Dielasma: Smith Plastoholotype
Smith, 1927, p. 123, pl. 102, figs. 14-16
Kupreanof Island, Alaska; Hamilton Bay
Upper Triassic [holotype USNM 74208]
hannibali, Discinisca cumingii: Hertlein and Grant Holotype
Hertlein and Grant, 1944, p. 29, pl. 16, figs. 7, 8, 11
Oak Bay, Wash.; between Port Townsend and Port Ludlow SU loc,
NP 128
Oligocene, Lincoln Fm
laevis, Rhynchonella wollossowitschii: Diener Holotype
Diener, 1924, p. 14, pl. 1, figs. 12a-12d
New Siberian Islands; Koteleny Island, at head of Balyktach River
Upper Triassic, Noric
lata, Rhynchonella wollossowitschii: Diener Holotype
Diener, 1924, p. 14, pl. 1, figs. 11a-11d
New Siberian Islands; Koteleny Island, head of Balyktach River
Upper Triassic, Noric
pittensis, Spiriferina: Smith Plastoholotype
Smith, 1927, p. 124, pl. 95, fig. 10
Shasta Co., Calif.; Brock Mt.
Upper Triassic, Hosselkus Ls [holotype USNM 74156]
richardsoni, Rhynchonella: Smith Plastoholotype
Smith, 1927, p. 123, pl. 96, figs. 19-21
Shasta Co., Calif.; old quarry SW end of Brock Mt. between Squaw
Creek and Pit River
Upper Triassic, Hosseikus Ls [holotype USNM 74171]
simiensis, Kingena: Waring Holotype
Waring, 1917, p. 73, pl. 12, fig. 11
Ventura Co., Calif.; Simi Hills, Martinez area
Lower Eocene, Martinez Fm
smithi, Terebratalia: Arnold Holotype
Arnold, 1903, p. 93, pl. 17, fig. 9
Los Angeles Co., Calif.; Deadman Island, off San Pedro
Pleistocene, San Pedro Fm
sookensis, Terebratella (?): Clark and Arnold Holotype
Clark and Arnold, 1923, p. 176, pl. 36, figs. 5a, 5b
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, at mouth of Fossil Creek SU loc. NP 130
Oligocene, Sooke Fm
sookensis, Terebratella (?): Clark and Arnold Paratype
Clark and Arnold, 1923, p. 176, pl. 36, fig. 4
Vancouver Island, British Columbia, Canada; Jordan River, 2 miles
W of Sherringham Point, at mouth of Fossi] Creek SU loc. NP 130
Oligocene, Sooke Fm
washingtonensis, Gryphus: Hertlein and Grant Holotype
Hertlein and Grant, 1944, p. 93, pl. 16, figs. 13, 14, 16
Grays Harbor Co., Wash.; R.R. cuts E of Balch SU loc. NP 57
Eocene, Cowlitz Fm
wollossowitschii, Rhynchonella: Diener Syntype
Diener, 1924, p. 14, pl. 1, figs. 10a-10d
New Siberian Islands; Koteleny Island, head of Balyktach River
Triassic
wollossowitschii, Rhynchonella: Diener Syntype
Diener, 1924, p. 14, pl. 1, figs. 9a-9d
New Siberian Islands; Koteleny Island, head of Balyktach River
Triassic
462
9927
5169
5374
6628
5601
5062
777a
777b
5286
5287
5287a
6612
BuLLetin 300
yukonensis, Spiriferina: Smith
Smith, 1927, p. 124, pl. 101, figs. 13, 14
Plastoholotype
S bank Yukon River, opposite Nation River
Upper Triassic
[holotype USNM 74202]
ARTHROPODA (EXCEPT OSTRACODA)
alaskensis, Portunites: Rathbun
Rathbun, 1926, p. 72, pl. 22, fig. 3
Pacific Co., Wash.; N of Holcomb, bluffs along Willapa River
loc. NP 253
Oligocene, Keasey Fm
antennatus, Archaeopus: Rathbun
Rathbun, 1908, p. 347, pl. 47, figs. 4, 5, 6
Paratype
SU
Paratype
San Mateo Co., Calif.; Bolsa Point, 1 mile N of Pigeon Point
Upper Cretaceous, Chico Fm
apollo, Cheirurus: Billings
Plastoholotype
Billings, 1860, p. 322, fig. 28. Also im Billings, 1865, fig. 397 (as
stay sp.”)
Quebec, Canada; Pt. Lévis
Middle Ordovician, Beekmantown Fm
5380]
bainbridgensis, Cancer: Rathbun
Rathbun, 1926, p. 60, pl. 16, fig. 2
Bainbridge Island, Wash.; Bean Point
Upper Oligocene, Blakeley Fm
bainbridgensis, Cancer: Rathbun
Rathbun, 1926, p. 60, pl. 16, fig. 3
Bainbridge Island, Wash.; Bean Point
Upper Oligocene, Blakeley Fm
bairdensis, Proteus: Wheeler
Wheeler, 1935, p. 49, pl. 6, figs. 1-3
Shasta Co., Calif.; Redding Qd, SW
R4W _ SU loc. 1041
Carboniferous, Baird Ls
bairdensis, Proteus: Wheeler
Wheeler, 1935, p. 49
Shasta Co., Calif.; Redding Qd, SW
R4W
Carboniferous, Baird Fm
bandonensis, Callianassa: Rathbun
Rathbun, 1926, p. 118, pl. 27, figs. 5, 6
Coos Co., Ore.; S of mouth of Five
NP 38
Oligocene
bandonensis, Callianassa: Rathbun
Rathbun, 1926, p. 118, pl. 27, fig. 8
Coos Co., Ore.; S of mouth of Five
NP 38
Oligocene
bandonensis, Callianassa: Rathbun
Rathbun, 1926, p. 118, pl. 27, fig. 7
Coos Co., Ore.; S of mouth of Five
NP 38
Oligocene
barrandei, Amphion: Billings
Billings, 1865, p. 288, fig. 277b
Newfoundland, Canada; Cow Head
[holotype Geol. Surv. Canada
Holotype
SU loc. NP 205
Paratype
SU loc. NP 205
Holotype
1/4 SE 1/4 Sec. 14, T 34 N,
Paratype
1/4 SE 1/4 Sec. 14, T 34 N,
Holotype
Mile Creek, Bandon SU loc.
Paratype
Mile Creek, Bandon SU loc.
Paratype
Mile Creek, Bandon SU loc.
Plastosyntype
Ordovician, Quebec group [syntype Geol. Surv. Canada 682b]
6613
6987
8305
9321
5321
9319
6619
6621
5176
6629
5077
5175
9273
STANFORD UNIVERSITY TyPEs: SMITH 463
barrandei, Amphion: Billings Plastosyntype
Billings, 1865, p. 288, fig. 277a
Newfoundland, Canada; Cow Head
Ordovician, Quebec group [syntype Geol. Surv. Canada 681b]
beattyana, Parapilekia: Holliday Paratype
Holliday, 1942, p. 475, pl. 73, fig. 4
Nye Co., Nevada; Furnace Creek Qd, 1 mile SE of Beatty, gully in
Meikeljohn Peak on road to Telluride Canyon SU loc. 2204
Lower Ordovician
bifida, Acanthopyge (Mephiarges): Edgell Holotype
Edgell, 1955, p. 138, pl. 14, figs. 1, 3-8
New South Wales, Australia; Goodradigbee Valley, 4 miles SE of
Burrinjuck Dam, 4 miles NNE of Wee Jasper Village
Middle Devonian, Wee Jasper Ls
brucei, Blepharipoda: Rathbun Holotype
Rathbun, 1926, p. 126, pl. 28, fig. 11
Jefferson Co., Wash.; Classens Wharf, Townsend Bay SU loc. NP
125
Oligocene, Lincoln Fm
brucei, Blepharipoda: Rathbun Paratype
Rathbun, 1926, p. 126, pl. 28, fig. 10
Jefferson Co., Wash.; Classens Wharf, Townsend Bay SU loc. NP
125
Oligocene, Lincoln Fm
brucei, Blepharipoda: Rathbun Paratype
Rathbun, 1926, p. 126
Jefferson Co., Wash.; Classens Wharf, Townsend Bay SU loc. NP
125
Oligocene, Lincoln Fm
canadensis, Amphion: Billings Plastosyntypes
Billings, 1859, p. 381, figs. 12a, 12b
Quebec; Mingan Islands
Middle Ordovician, Chazyan, Mingan Fm [syntypes at Geol. Sury.
Canada]
carmanahensis, Pilumnoplax: Rathbun Holotype
Rathbun, 1926, p. 38, pl. 9, figs. 1-4
Vancouver Island, British Columbia, Canada; sea cliffs for a distance
of 3 miles W of Carmanah Point SU loc. NP 141
Oligocene
cayleyi, Amphion: Billings Plastoholotype
Billings, 1863, p. 239, fig. 277. Also zm Billings, 1865, p. 413, fig. 398
(as Amphion sp?)
Quebec; Pt. Lévis
Middle Ordovician, Lévis Fm _ [holotype Geol. Surv. Canada 825]
conwayensis, Griffithides: Wheeler Holotype
Wheeler, 1935, p. 53, pl. 6, figs. 4, 5
Conway Co., Ark.; near center NW 1/4 Sec. 17, T 5 N,R 16 W SU
loc. 1040
Pennsylvanian, Atoka Fm [new name for Phillipsia (Griffithides)
ornata Vogdes, 1895]
hannibalanus, Pilumnoplax: Rathbun Holotype
Rathbun, 1926, p. 39, pl. 10, figs. 1, 2
Nehalem Bay, Ore.; cut on Tillamook branch of Southern Pacific
R.R., 1 mile E of Wheeler SU loc. NP 229
Middle Oligocene?
hannibalanus, Pilumnoplax: Rathbun Paratype
Rathbun, 1926, p. 39, pl. 10, fig. 3
W of Neah Bay, Wash.; sea cliffs at Koitlah Point SU loc. NP 167
Middle Oligocene?
5951
6626
9258
6986
5070
6611
5077
10301
5063
5064
5065a
5065b
BuLuetin 300°
idae, Mesorhoea: Rathbun Paratype
Rathbun, 1926, p. 27
NE of San Pedro, Calif.; Nob Hill
Pleistocene, San Pedro Fm
julius, Amphion: Billings Plastoholotype
Billings, 1865, p. 290, fig. 279
Newfoundland; Cow Head
Middle Ordovician, Quebec group [holotype Geol. Surv. Canada
680]
marcusana, Mursia: Rathbun Paratype
Rathbun, 1926, p. 82, pl. 19, fig. 7
Puget Sound, Wash.; Alki Point, near Seattle SU loc. NP 48
Upper Oligocene, Blakeley Fm
marginatus, Ectenonotus: Holliday Paratype
Holliday, 1942, p. 476, pl. 73, fig. 3
Nye Co., Nevada; Furnace Creek Qd, 1 mile SE of Beatty SU loc.
2205
Lower Ordovician
naselensis, Eumorphocorystes: Rathbun Holotype
Rathbun, 1926, p. 100, pl. 24, figs. 9, 10
Pacific Co., Wash.; bluffs along Nasel River near mouth of Salmon
Creek SU loc. NP 281
Oligocene, Lincoln Fm
nevadensis, Amphion: Walcott Plastoholotype
Walcott, 1884, p. 94, pl. 12, fig. 13
Eureka district, Nevada
Ordovician, Pogonip Fm [holotype USNM 24645]
nosoniensis, Griffithides: Wheeler Holotype
Wheeler, 1935, p. 51, pl. 6, figs. 6, 7
Shasta Co., Calif.; Redding Qd, NE 1/4 SW 1/4 Sec. 24, T 34. N, R
4 W_ SU loc. 1034
Permian, Nosoni Fm
ornata, Phillipsia (Griffithides): Vogdes Holotype
Vogdes, 1895, pp. 589-591, text fig. [renamed Griffithides conwayensis
by Wheeler, 1935, p. 53]
Conway Co., Ark.; Sec. 17,T 5N,R16 W_ SU loc. 1040
Pennsylvanian, Atoka Fm
pleistocenica, Randallia: Rathbun Paratype
Rathbun, 1926, p. 77
San Pedro, Calif.; Nob Hill
Pleistocene
porterensis, Callianassa: Rathbun Holotype
Rathbun, 1926, p. 119, pl. 28, fig. 4
Wash.; bluff on Chehalis River below Porter SU loc. NP 53
Oligocene, Lincoln Fm
porterensis, Callianassa: Rathbun Paratype
Rathbun, 1926, p. 119
Wash.; bluff on Chehalis River below Porter SU loc. NP 53
Oligocene, Lincoln Fm
porterensis, Callianassa: Rathbun Paratype
Rathbun, 1926, p. 119, pl. 28, fig. 3 (as paratype C)
Yaquina Bay, Ore.; cut on C and E. R.R. between Rocky Point and
Oysterville SU loc. NP 15
Oligocene, Lincoln Fm
porterensis, Callianassa: Rathbun Paratype
Rathbun, 1926, p. 119, pl. 28, fig. 1 (as paratype D)
Yaquina Bay, Ore.; cut on C and E. R.R. between Rocky Point and
Oysterville SU loc. NP 15
Oligocene, Lincoln Fm
6610
5322
5172a
5172b
5066
6830
6831
6835
6839
6840
STANFORD UNIVERSITY TyPEs: SMITH
salteri, Amphion: Billings
Billings, 1861, p. 322, fig. 6
465
Plastoholotype
Grenville Co., Ontario, Canada; “Philipsburg,” Oxford Township
Lower Ordovician, Beekmantown Fm _ [holotype Geol. Surv. Canada
515]
triangulum, Portunites: Rathbun
Paratype
Rathbun, 1926, p. 68, pl. 17, figs. 3, 4
Lewis Co., Wash.; Chehalis River, near mouth of Lincoln Creek
SU loc. NP 211
Oligocene, Lincoln Fm
twinensis, Callianassa: Rathbun
Rathbun, 1926, p. 117, pl. 27, fig. 2
Clallam Co., Wash.; W of Twin
SU loc. NP 120
Oligocene, Blakeley Fm
twinensis, Callianassa: Rathbun
Rathbun, 1926, p. 117, pl. 27, fig. 3
Clallam Co., Wash.; W of Twin
SU loc. NP 120
Oligocene, Blakeley Fm
twinensis, Callianassa: Rathbun
Rathbun, 1926, p. 117
Clallam Co., Wash.; W of Twin
SU loc. NP 120
Oligocene, Blakeley Fm
twinensis, Callianassa: Rathbun
Holotype
River for a distance of 3/4 mile
Paratype
River for a distance of 3/4 mile
Paratype
River for a distance of 3/4 mile
Paratype
Rathbun, 1926, p. 117, pl. 27, fig. 3 (as paratype D)
Wahkiakum Co., Wash.; bluffs on Gray’s River
Oligocene, Blakeley Fm
willapensis, Ranidina: Rathbun
SU loc. NP 274
Holotype
Rathbun, 1926, p. 99, pl. 21. figs. 4, 5
Thurston Co., Wash.; bluffs along Willapa River N of Holcomb
SU loc. NP 253
Middle Oligocene, Keasey Fm
OSTRACODA
beaconensis, Cytheridea: LeRoy
Holotype
LeRoy, 1943, p. 359, pl. 58, figs. 21-24
Wilmington Qd., Calif.; San Pedro,
Pleistocene, San Pedro Fm
beaconensis, Cytheridea: LeRoy
LeRoy, 1943, p. 359, pl. 58, fig. 25
Wilmington Qd., Calif.; San Pedro,
Pleistocene, San Pedro Fm
californica, Cytherelloidea: LeRoy
Beacon and Second Streets
Paratype
Beacon and Second Streets
Holotype
LeRoy, 1943, p. 357, pl. 58, figs. 32-35
Wilmington Qd., Calif.; San Pedro, 7.2 inches N, 2.05 inches E of SW
corner of sheet
Pleistocene, Lomita Fm
californiensis, Hemicythere?: LeRoy
Holotype
LeRoy, 1943, p. 366, pl. 61, figs. 29-31
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
californiensis, Hemicythere?: LeRoy
Paratype
LeRoy, 1943, p. 366, pl. 61, figs. 32, 33
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
466
6846
6841
6842
6807
6808
6800
6779
6809
6776
6777
6851
6853
6804
6805
6806
BuLLETIN 300
californiensis, Hemicythere?: LeRoy Paratype
LeRoy, 1943, p. 366, pl. 62, figs. 5, 6
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
californiensis, Hemicythere?: LeRoy Paratypes
LeRoy, 1943, p. 366
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
corrugata, Leguminocythereis: LeRoy Holotype
LeRoy, 1943, p. 372, pl. 59, figs. 7-10
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
corrugata, Leguminocythereis: LeRoy Paratype
LeRoy, 1943, p. 372, pl. 59, figs. 11, 12
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
corrugata, Leguminocythereis: LeRoy Paratype
LeRoy, 1943, p. 372, pl. 62, figs. 7, 8
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
corrugata, Leguminocythereis: LeRoy Paratypes
LeRoy, 1943, p. 372
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
delreyensis, Basslerites: LeRoy Holotype
LeRoy, 1943, p. 368, pl. 59, figs. 23-26
Los Angeles Co., Calif.; Venice Qd, 6.2 inches S, 3.9 in W of NE
corner of map, on Lincoln Blvd
Pleistocene
delreyensis, Basslerites: LeRoy Paratypes
LeRoy, 1943, p. 368, pl. 59, fig. 27
Los Angeles Co., Calif.; Venice Qd, 6.2 inches S, 3.9 inches W of NW
corner of map, on Lincoln Blvd
Pleistocene
delreyensis, Basslerites: LeRoy Paratype
LeRoy, 1943, p. 368, pl. 62, figs. 21, 22
Los Angeles Co., Calif.; Venice Qd, 6.2 inches S, 3.9 inches W of NE
corner of map, on Lincoln Blvd
Pleistocene
delreyensis, Basslerites: LeRoy Paratype
LeRoy, 1943, p. 368
Los Angeles Co., Calif.; Venice Qd, 6.2 inches S, 3.9 inches W of NE
corner of map, on Lincoln Blvd
Pleistocene
diegoensis, Cythereis: LeRoy Holotype
LeRoy, 1943, p. 369, pl. 58, figs. 26-29
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
diegoensis, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 369, pl. 58, figs. 30, 31
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
diegoensis, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 369
San Diego, Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
6781
6775
6849
6782
6829
6788
6789
6778
6792
6845
6815
6816
6810
STANFORD UNIVERSITY Types: SMITH 467
driveri, Brachycythere: LeRoy Holotype
LeRoy, 1943, p. 361, pl. 61, figs. 6-8
Santa Barbara, Calif.; Bathhouse Beach
“Pliocene,” Santa Barbara Fm
driveri, Brachycythere: LeRoy Paratype
LeRoy, 1943, p. 361, pl. 61, figs. 9, 10
Santa Barbara, Calif.; Bathhouse Beach
“Pliocene,” Santa Barbara Fm
driveri, Brachycythere: LeRoy Paratype
LeRoy, 1943, p. 361, pl. 62, figs. 17, 18
Santa Barbara, Calif.; Bathhouse Beach
“Pliocene,” Santa Barbara Fm
driveri, Brachycythere: LeRoy Paratype
LeRoy, 1943, p. 361
Santa Barbara, Calif.; Bathhouse Beach
‘Pliocene,” Santa Barbara Fm
elongata, Bythocypris: LeRoy Holotype
LeRoy, 1943, p. 358, pl. 59, figs. 13-16
Wilmington Qd, Calif.; San Pedro, Second St., 7.2 inches N, 2.05 inches
E of SW corner of sheet
Pleistocene, Lomita Fm
fragilis, Caudites: LeRoy Holotype
LeRoy, 1943, p. 372, pl. 60, figs. 10-12
Wilmington Qd, Calif.; San Pedro, Second Street, 100’ E of Pacific
Ave
Pleistocene, Lomita Fm
fragilis, Caudites: LeRoy Paratype
LeRoy, 1943, p. 372, pl. 60, fig. 13
Wilmington Qd., Calif.; San Pedro, Second Street, 100’ E of Pacific
Ave
Pleistocene, Lomita Fm
granti, Paracytheridea: LeRoy Syntype
LeRoy, 1943, p. 361, pl. 61, figs. 11, 12, 14
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pleistocene, San Diego Fm
granti, Paracytheridea: LeRoy Syntype
LeRoy, 1943, p. 361, pl. 61, fig. 13
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
granti, Paracytheridea: LeRoy Syntype
LeRoy, 1943, p. 361, pl. 62, figs. 3, 4
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
hispida, Hemicythere? californiensis: LeRoy Holotype
LeRoy, 1943, p. 367, pl. 60, figs. 1-3
Santa Barbara, Calif.; Bathhouse Beach
“Pliocene,” Santa Barbara Fm
hispida, Hemicythere? californiensis: LeRoy Paratype
LeRoy, 1943, p. 367, pl. 60, fig. 4
Santa Barbara, Calif.; Bathhouse Beach
“Pliocene,” Santa Barbara Fm
holmani, Archicythereis: LeRoy Holotype
LeRoy, 1943, p. 371, pl. 58, figs. 1-4
Orange Co., Calif.; Newport Lagoon, Tustin Qd, 2.95 inches N, 1.08
inches E of SW corner of map
Upper Pliocene
468
6852
6801
6802
6803
6848
6797
6798
6843
6799
6836
6837
6838
6774
BULLETIN 300
holmani, Archicythereis: LeRoy Paratype
LeRoy, 1943, p. 371, pl. 62, figs. 23, 24
Orange Co., Calif.; Newport Lagoon, Tustin Qd, 2.95 inches N, 1.08
inches E of SW corner of map
Upper Pliocene
jollaensis, Hemicythere: LeRoy Holotype
LeRoy, 1943, p. 365, pl. 59, figs. 28-31
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
jollaensis, Hemicythere: LeRoy Paratype
LeRoy, 1943, p. 365, pl. 59, figs. 32, 33
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
jollaensis, Hemicythere: LeRoy Paratype
LeRoy, 1943, p. 365, text-fig. q
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
jollaensis, Hemicythere: LeRoy Paratype
LeRoy, 1943, p. 365, pl. 62, figs. 15, 16
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
kewi, Cythereis: LeRoy Holotype
LeRoy, 1943, p. 369, pl. 60, figs. 24-26
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,’ Santa Barbara Fm
kewi, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 369, pl. 60, fig. 27
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
kewi, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 369, pl. 62, figs. 9, 10
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
kewi, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 369, text-fig. d
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,’ Santa Barbara Fm
lenticulata, Loxoconcha: LeRoy Holotype
LeRoy, 1943, p. 360, pl. 60, figs. 19-23
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
lenticulata, Loxoconcha: LeRoy Paratype
LeRoy, 1943, p. 360, text fig. f
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
lenticulata, Loxoconcha: LeRoy Paratype
LeRoy, 1943, p. 360, text-fig. g
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
lenticulata, Loxoconcha: LeRoy Paratype
LeRoy, 1943, p. 360, pl. 61, figs. 34-36
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
6847
6783
6784
6785
5922
6823
6824
6825
6780
6812
6813
6814
STANFORD UNIVERSITY TyYPEs: SMITH 469
lenticulata, Loxoconcha: LeRoy Paratype
LeRoy, 1943, p. 360, pl. 62, figs. 13, 14
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
lincolnensis, Brachycythere: LeRoy Holotype
LeRoy, 1943, p. 364, pl. 61, figs. 1-3
Los Angeles Co., Calif.; Lincoln Blvd., Venice; Venice Qd, 6.2 inches
S, 3.9 inches W of NE corner of sheet
Pleistocene
lincolnensis, Brachycythere: LeRoy Paratype
LeRoy, 1943, p. 364, pl. 61, figs. 4, 5
Los Angeles Co., Calif.; Lincoln Blvd., Venice; Venice Qd, 6.2 inches
S, 3.9 inches W of NE corner of sheet
Pleistocene
lincolnensis, Brachycythere: LeRoy Paratype
LeRoy, 1943, p. 364, pl. 62, figs. 1, 2
Los Angeles Co., Calif.; Lincoln Blvd., Venice; Venice Qd, 6.2 inches
S, 3.9 inches W of NE corner of sheet
Pleistocene
martini, Brachycythere: Murray and Hussey Paratype
Murray and Hussey, 1942, p. 177
Louisiana; S side of lower Dodson Rd, Winn Parish; NE 1/4, SW 1/4,
SW 1/4 Sec. 28, T 13 N,R3 W
Eocene, Cook Mountain Fm
microreticulata, Cythereis: LeRoy Holotype
LeRoy, 1943, p. 370, pl. 59, figs. 17-20
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
microreticulata, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 370, pl. 59, figs. 21, 22
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,’ Santa Barbara Fm
microreticulata, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 370, text fig. n
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
minutum, Cytheropteron: LeRoy Holotype
LeRoy, 1943a, p. 361, pl. 60, figs. 28-30. [LeRoy, 1943b, p. 629, re-
named it Cytheropteron pacificum |
Los Angeles Co., Calif.; LaHabra Qd, 8.55 inches S, 4.25 inches E
of NW corner of map
Pliocene? [specimen missing, July, 1976]
newportensis, Archicythereis: LeRoy Syntype
LeRoy, 1943, p. 372, pl. 58, figs. 7, 8
Orange Co., Calif.; Newport Lagoon, Tustin Qd, 2.95 inches N, 1.08
inches E of SW corner of sheet
Upper Pliocene
newportensis, Archicythereis: LeRoy Syntype
LeRoy, 1943, p. 372, pl. 58, figs. 5, 6
Orange Co., Calif.; Newport Lagoon, Tustin Qd, 2.95 inches N, 1.08
inches E of SW corner of sheet
Upper Pliocene
newportensis, Archicythereis: LeRoy Syntype
LeRoy, 1943, p. 372, text fig. b
Orange Co., Calif.; Newport Lagoon, Tustin Qd, 2.95 inches N, 1.08
inches E of SW corner of sheet
Upper Pliocene
470
6769
6770
6786
6787
6826
6827
6828
6817
6818
6850
6832
6833
6834
BULLETIN 300
pacificus, Paracypris: LeRoy Holotype
LeRoy, 1943, p. 358, pl. 61, figs. 15-17 and text fig. z
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
pacificus, Paracypris: LeRoy Paratype
LeRoy, 1943, p. 358, pl. 61, fig. 18
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
palosensis, Hemicythere: LeRoy Holotype
LeRoy, 1943, p. 365, pl. 60, figs. 14-16 and text fig. c
Los Angeles Co., Calif.; San Pedro, Wilmington Qd, on Second Street,
100’ E of Pacific Ave
Pleistocene, Lomita Fm
palosensis, Hemicythere: LeRoy Paratype
LeRoy, 1943, p. 365, pl. 60, figs. 17, 18
Los Angeles Co., Calif.; San Pedro, Wilmington Qd, on Second Street,
100’ E of Pacific Ave ;
Pleistocene, Lomita Fm
pedroensis, Cytheridea?: LeRoy Holotype
LeRoy, 1943, p. 359, pl. 58, figs. 15-18
Los Angeles Co., Calif.; San Pedro, Wilmington Qd, Beacon and
Second Streets
Pleistocene, San Pedro Fm
pedroensis, Cytheridea?: LeRoy Paratype
LeRoy, 1943, p. 359, pl. 58, figs. 19, 20
Los Angeles Co., Calif.; San Pedro, Wilmington Qd, Beacon and
Second Streets
Pleistocene, San Pedro Fm
pedroensis, Cytheridea?: LeRoy Paratype
LeRoy, 1943, p. 359, text-fig. t
Los Angeles Co., Calif.; San Pedro, Wilmington Qd, Beacon and
Second Streets
Pleistocene, San Pedro Fm
pennata, Cythereis: LeRoy Holotype
LeRoy, 1943, p. 370, pl. 59, figs. 34-37
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
pennata, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 370, text-fig. h
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
pennata, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 370, pl. 62, figs. 19, 20
San Diego Co., Calif.; Pacific Beach, La Jolla Qd
Pliocene, San Diego Fm
schencki, Cythereis: LeRoy Holotype
LeRoy, 1943, p. 371, pl. 58, figs. 9-12
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
schencki, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 371, pl. 58, figs. 13, 14
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
schencki, Cythereis: LeRoy Paratype
LeRoy, 1943, p. 371, text-fig. u
Santa Barbara, Calif.; Bathhouse Beach
“Upper Pliocene,” Santa Barbara Fm
6819
6820
6821
6822
6795
6796
6771
6772
6773
7330a
7330b
STANFORD UNIversIty TyrEs: SMITH 471
schumannensis, Brachycythere lincolnensis: LeRoy Holotype
LeRoy, 1943, p. 364, pl. 59, figs. 1-4
Santa Barbara Co., Calif.; Guadalupe Qd, in R.R. cut just N of
Schumann
Pliocene, Foxen Mudstone
schumannensis, Brachycythere lincolnensis: LeRoy Paratype
LeRoy, 1943, p. 364, pl. 59, figs. 5, 6
Santa Barbara Co., Calif.; Guadalupe Qd, in R.R. cut just N of
Schumann
Pliocene, Foxen Mudstone
schumannensis, Brachycythere lincolnensis: LeRoy Paratype
LeRoy, 1943, p. 364, text-fig. i
Santa Barbara Co., Calif.; Guadalupe Qd, in R.R. cut just N of
Schumann
Pliocene, Foxen Mudstone
schumannensis, Brachycythere lincolnensis: LeRoy Paratype
LeRoy, 1943, p. 364, text-fig. j
Santa Barbara Co., Calif.; Guadalupe Qd, in R.R. cut just N of
Schumann
Pliocene, Foxen Mudstone
simiensis, Pyricythereis: LeRoy Holotype
LeRoy, 1943, p. 368, pl. 61, figs. 24-26
Ventura Co., Calif.; Piru Qd, Happy Canyon, N side of Simi Valley
Pliocene, “San Diego” Fm
simiensis, Pyricythereis: LeRoy Paratype
LeRoy, 1943, p. 368, pl. 61, figs. 27, 28, text fig. e
Ventura Co., Calif.; Piru Qd, Happy Canyon, N side of Simi Valley
Pliocene, “San Diego” Fm
verdesensis, Bairdia: LeRoy Holotype
LeRoy, 1943, p. 358, pl. 60, figs. 5-7
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
verdesensis, Bairdia: LeRoy Paratype
LeRoy, 1943, p. 358, pl. 60, figs. 8, 9
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
verdesensis, Bairdia: LeRoy Paratype
Los Angeles Co., Calif.; Deadman Island, Wilmington Qd, 4.72 inches
N, 4.6 inches E of SE corner of sheet
Pleistocene, Timms Point Fm
FORAMINIFERA
acris, Schwagerina pavilionensis: Thompson and Wheeler
Syntypes
Thompson and Wheeler, 1942, p. 707, pl. 105, figs. 1, 2
Southern British Columbia, Canada; Marble Canyon
Permian, Marble Canyon Ls
aculeata, Schwagerina: Thompson and Hazzard Holotype
Thompson and Hazzard, 1946, p. 45, pl. 12, fig. 5
San Bernardino Co., Calif.; E front of Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
aculeata, Schwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 45, pl. 12, figs. 1, 4, 6, 7, 8
San Bernardino Co., Calif.; Providence Mts., S of Gilroy Mine, 1.5
miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
7708
753
6134
cell 36
7824
7825
to
7833
5633
5632
$396
710
711
712
7752
BuLLETIN 300
aculeata, Schwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 45, pl. 12, fig. 2
San Bernardino Co., Calif., along ridge and main divide, Providence
Mts., about 1.25 miles W of mouth of large canyon just N of Gilroy
Mine
Permian, Bird Spring Fm
aculeata, Schwagerina: Thompson and Hazzard Paratype
Thompson and Hazzard, 1946, p. 45, pl. 12, fig. 3
San Bernardino Co., Calif., along ridge and main divide, Providence
Mts., about 1.25 miles W of mouth of large canyon just N of Gilroy
Mine
Permian, Bird Spring Fm
advena, Bolivina: Cushman Paratype
Cushman, 1925c, p. 29
San Luis Obispo Co., Calif., Sec. 24, T 28S, R14E
Miocene, Monterey Shale
aequa, Globorotalia crassata: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 12
Trinidad, British W. I., Soldado Rock
Paleocene, Soldado Fm
afghanensis, Polydiexodina: Thompson Holotype
Thompson, 1946, p. 150, pl. 26, fig. 2
Shibar Pass, Afghanistan, on road from Kabul to Bamian, ca. 7 km
W of summit
Permian, Bamian Ls
afghanensis, Polydiexodina: Thompson Paratypes
Thompson, 1946, p. 150, pl. 26, figs. 3-5; pl. 24, figs. 1-6
Shibar Pass, Afghanistan, on road from Kabul to Bamian, ca. 7 km
W of summit
Permian, Bamian Ls
alazanensis, Bolivina: Cushman Paratype
Cushman, 1926a, p. 82
Veracruz, Mexico, km post 20.15, Tampico-Panuco R.R.
Eocene-Oligocene?, Alazan Clay
aliformis, Bolivina mexicana: Cushman Paratype
Cushman, 1926a, p. 82
Veracruz, Mexico; Rio Buena Vista, .5 km S 25° E from Tumbadero
Hacienda House
Eocene-Oligocene?, Alazan Clay
almgreni, Lenticulina: Martin Holotype
Martin, Lewis, 1964, p. 65, pl. 6, figs. 1a, 1b
Merced Co., Calif., Laguna Seca Creek, Laguna Seca Hills, 7 miles
N of Little Panoche Creek
Upper Cretaceous, Panoche Fm
alticostatus, Robulus mexicanus: Cushman and Barksdale
Holotype
Cushman and Barksdale, 1930, p. 63, pl. 11, fig. 7
Contra Costa Co., Calif., Carquinez Qd., on Shell Co. property 1.5
miles E of Arroyo del Hambre, 1.1 miles S of Bull’s Head Point
Eocene, upper Martinez Fm [Paleocene]
alticostatus, Robulus mexicanus: Cushman and Barksdale
Paratypes
Cushman and Barksdale, 1930, p. 63, pl. 11, figs. 4-6
Contra Costa Co., Calif., Carquinez Qd., on Shell Co. property 1.5
miles E of Arroyo del Hambre, 1.1 miles S of Bull’s Head Point
Eocene, upper Martinez Fm _ [Paleocene]
angulata, Entosolenia marginata: Uchio Paratype
Uchio, 1951a, p. 38
Chiba-ken, Japan; Tomiya, Takeoka-mura, Kimitsu-gun
Pliocene, Tomiya Fm
586
9801
6132
cell 37
7732
7733
7734
7749
6561
6133
cell 13
8112
947
7517
9465
5466
7446
STANFORD UNIVERSITY TyrEs: SMITH 473
angulostriata, Quinqueloculina: Cushman and Valentine
Paratype
Cushman and Valentine, 1930, p. 12, as angulo-striata
Channel Islands off southern California
appressa, Valvulineria californica: Cushman Paratypes
Cushman, 1926d, p. 60
San Luis Obispo Co., Calif., Sec. 24, T 28 S, R14E
Miocene, Monterey Shale
arbenzi, Planularia: Cushman and Renz Paratype
Cushman and Renz, p. 13
Falcon, Venezuela; Pozon, 17.7 km SE of Pueblo Jacura, District
Acosta.
Miocene, Agua Salada (Zone II)
arta, Pseudoschwagerina: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 49, pl. 18, figs. 1-3
San Bernardino Co., Calif., along ridge and summit of main divide,
Providence Mts., ca. 1.25 miles W of mouth of large canyon just N
of Gilroy Mine
Permian, Bird Spring Fm
asanoi, Cassidulina: Uchio Paratype
Uchio, 1950, p. 190, text-fig. 13
Chiba-ken, Japan; Otaniki, Tsuchimutsu-mura, Chosei-gun
upper Pliocene
aster, Asterocyclina: Woodring Paratypes
Woodring, 1930, p. 152
Santa Barbara Co., Calif., Canada de los Sauces
Eocene
attenuata, Uvigerina auberiana: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 21
Falcon, Venezuela; Isidro, 35.0 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada (Zone III)
australis, Bolivinoides decorata: Edgell Holotype
Edgell, 1954, p. 71, pl. 13, fig. 6
Northwest Australia, C.Y. Creek, W flank of Giralia Anticline
Upper Cretaceous
baggi, Planulina: Kleinpell Holotype
Kleinpell, 1938, p. 349, pl. VIII, figs. 14a, 14b, 14c
Monterey Co., Calif., Reliz Canyon
Miocene, Salinas Shale
baileyi, Cibicides: Beck Holotype
Beck, 1943, p. 611, pl. 109, figs. 7-9
Lewis Co., Wash., W bank of Cowlitz River, 1.5 miles E of Vader,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2 W
Eocene, Cowlitz Fm
bandyi, Rotalia: Martin Holotype
Martin, Lewis, 1964, p. 94, pl. 12, figs. 10a, 10b, 10c
Fresno Co., Calif.; Moreno Gulch, Panoche Hills, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
barbarense, Elphidium fax: Nicol Holotype
Nicol, 1944, p. 178, pl. 29, figs. 10, 12
Santa Barbara, California
Pleistocene, Santa Barbara Fm
barksdalei, Astacolus: Beck Holotype
Beck, 1943, p. 597, pl. 104, fig. 17
Lewis Co., Wash.; E 1/2, SE 1/4 Sec. 28, T 11 N, R 2 W; 1.5 miles
E of Vader, on W bank of Cowlitz River
Eocene, Cowlitz Fm
474
6133
cell 14
6132
cell 19
756
7048
836
6881
7392
6084
758
6132
cell 28
745
BuLLeTIN 300
basicordata, Uvigerina gallowayi: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 21
Falcon, Venezuela; Tocuyo, 18.7 km S of San Juan de los Cayos,
District Acosta
Miocene, Agua Salada (Zone II)
basispinosa, Marginulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 13
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada (Zone IV)
beali, Cristellaria: Cushman Paratype
Cushman, 1925b, p. 25
San Luis Obispo Co., Calif.; Sec. 24, T 28S, R14E
Miocene, Monterey Shale
beatus, Cibicides: Martin Holotype
Martin, Lois, 1943, p. 30, pl. 8, figs. 6a-6c
Fresno Co., Calif.; Lodo Gulch, Tumey Hills Qd
Eocene, Lodo Fm
belridgensis, Nonion: Barbat and Johnson Holotype
Barbat and Johnson, 1934, p. 11, pl. 1, fig. 8
Kern Co., Calif.; McKittrick Qd, Sec. 30. 1128.8; R 21 E; Ohio Oil
Coz Bearstate No. 23, Belridge field
Miocene, Reef Ridge Shale
belridgensis, Nonion: Barbat and Johnson Paratype
Barbat and Johnson, 1934, p. 11, pl. 1, fig. 9
Kern Co., Calif.; McKittrick Qd, Sec. 30, T 28 S, R 21 E; Ohio Oil
Co., Bearstate No. 23, Belridge field
Miocene, Reef Ridge Shale
biserialis, Dyocibicides: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 30
Channel Islands, off southern California
biserialis, Siphonides: Feray Paratype
Feray, 1941, p. 175
Smithville, Texas; S bank of the Colorado River
Middle Eocene, Weches Fm, Claiborne Group
bleeckeri, Bulimina: Hedberg Paratypes
Hedberg, 1937, p. 675
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
bradburyi, Bulimina: Martin Paratype
Martin, Lois, 1943, p. 19, pl. 6, figs. 4a, 4b
Fresno Co., Calif.; Panoche Qd, Lodo Gulch [Tumey Hills Qd]
Eocene, Lodo Fm
bramlettei, Bolivina: Kleinpell Paratype
Kleinpell, 1938, p. 67, pl. 21, figs. 10, 11
Los Angeles Co., Calif.; Palos Verdes Hills
Miocene, Valmonte Diatomite
brevior, Bolivina: Cushman Paratype
Cushman, 1925c, p. 31
San Luis Obispo Co., Calif.; Sec. 24,T 28 S, R14E
Miocene, Monterey Shale
brevis, Textularia miocenica: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 9
Falcon, Venezuela; Tocuyo, 18.7 km S of San Juan de los Cayos,
District Acosta
Lower Miocene, Agua Salada Fm (Zone II)
byramensis, Pulvinulina: Cushman Paratype
Cushman, 1922, p. 99, pl. XXII, figs. 4, 5
Byram, Miss.
Oligocene, Byram Marl
5796
5629
570
571
072
073
574
5975
576
O77
578
7349
STANFORD UNIVERSITY TYPES: SMITH
californica, Buliminella: Cushman
Cushman, 1925c, p. 33
San Luis Obispo Co., Calif.; Sec. 24, T 28 S,R14E
Miocene, Monterey Shale
californica, Cassidulina: Cushman and Hughes
Cushman and Hughes, 1925, p. 12
San Pedro, Calif.; Timms Point
SU loc. 2024
“Pliocene,” Timms Point Fm _ [Pleistocene]
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 27, figs. 4, 6
475
Paratype
Paratypes
Paratype
Santa Clara Co., Calif.; New Almaden Qd, .25 miles NE of Guada-
lupe quicksilver mine; S 72° W from Pioneer School, S 44° E from
Lone Hill
Eocene, “Tejon” Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 28, figs. 2, 5
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, “Tejon” Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 28, fig. 4
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, ‘Tejon’ Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 28, fig. 3
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, “Tejon” Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 29, fig. 1
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, “Tejon” Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 29, fig. 3
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, ‘Tejon’ Fm
californica, Discocyclina: Schenck
Schenck, 1929, p. 224, pl. 30, fig. 3
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, “Tejon” Fm
californica, Discocyclina: Schenck
Qd,
Qd,
Qd,
Qd,
Qd,
Qd,
Schenck, 1929, p. 224, pl. 30, fig. 3 (type 577)
Santa Clara Co., Calif.; New Almaden
lupe quicksilver mine
Eocene, “Tejon” Fm
Qd,
25
25
25
25
25
25
25
miles NE
miles NE
miles NE
miles NE
miles NE
miles NE
miles NE
Paratype
of Guada-
Paratype
of Guada-
Paratype
of Guada-
Paratype
of Guada-
Paratype
of Guada-
Paratype
of Guada-
Paratypes
of Guada-
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Schenck and Childs, 1942, p. 17, pl. 2, fig. 4; pl. 3, fig. 4
San Luis Obispo Co., Calif.; Adelaida Qd, near BM 836, center Sec.
7, AP PAS BERTI 185 SW thee. 1al55
Oligocene, Vaqueros Fm
Holotype
476
7358
7359
7360
7365
7356
7350
7359
7353
7354
73951
7352
7357
7361
7363
7362
7364
6100
5800
9331a
7685
7686
7687
7688
7689
7633
7634
7636
BULLETIN 300
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Paratypes
Schenck and Childs, 1942, p. 17, pl. 1, figs. 1 (type 7358), 2 (type
7365), 3 (type 7359), 4 (type 7360)
San Luis Obispo Co., Calif.; Adelaida Qd, center Sec. 7, T 26 S, R 10
B.S U) Toe. b055
Oligocene, Vaqueros Fm
californica, Lepidocyclina (Lepidecyclina): Schenck and Childs
Paratype
Schenck and Childs, 1942, p. 17, pl. 2, fig. 1; pl. 3, fig. 1
San Luis Obispo Co., Calif.; Adelaida Qd, Sec. 7, T 26S, R 10E
Oligocene, Vaqueros Fm
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Paratype
Schenck and Childs, 1942, p. 17, pl. 2, fig. 2; pl. 3, fig. 6
San Luis Obispo Co., Calif.; Adelaida Qd, Sec. 7, T 26 S,R10E
Oligocene, Vaqueros Fm
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Paratype
Schenck and Childs, 1942, p. 17, pl. 2, fig. 3; pl. 3, fig. 9
San Luis Obispo Co., Calif.; Adelaida Qd, Sec. 7, T 26 $,R10E
Oligocene, Vaqueros Fm
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Paratypes
Schenck and Childs, 1942, p. 17, pl. 3, figs. 2 (type 7353), 3 (type
7354), 5 (type 7351), 7 (type 7352), 8 (type 7357)
San Luis Obispo Co., Calif.; Adelaida Qd, Sec. 7, T 26S, R10 E
Oligocene, Vaqueros Fm
californica, Lepidocyclina (Lepidocyclina): Schenck and Childs
Paratypes
Schenck and Childs, 1942, p. 17, pl. 4, figs. 1, 5 (type 7361), figs. 2,
6 (type 7363), 3 (type 7362), 4 (type 7364)
San Luis Obispo Co., Calif.; Adelaida Qd, Sec. 7, T 26S,R 10E
Oligocene, Vaqueros Fm
californica, Suggrunda: Kleinpell Holotype
Kleinpell, 1938, p. 287, pl. 18, figs. 8-10
Contra Costa Co., Calif.; San Pablo Creek
Miocene, Tice Shale
californica, Valvulineria: Cushman Paratype
Cushman, 1926d, p. 60
San Luis Obispo Co., Calif.; Sec. 24, T 28 S,R 14 E
Miocene, Monterey Shale
californicus, Bathysiphon: Martin Holotype
Martin, Lewis, 1964, p. 43, pl. 1, figs. 2a, 2b
Fresno Co., Calif.; eastern Panoche Hills Martin loc. MG 247
Cretaceous, Panoche group, Uhalde Shale [missing; no record that
specimen was received at SU]
californicus, Triticites: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 42, pl. 10, figs. 10 (type 7685), 11
(type 7686), 12 (type 7687), 13 (type 7688), 14 (type 7689)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
calx, Parafusulina? Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 29, pl. 4, figs. 4 (type 7634), 5
(type 7633), 6 (type 7636)
Shasta Co., Calif.; Redding Qd, NE 1/4 SE 1/4 Sec. 23, T 34 N, R 4
W, crest of Is ridge S of Potter Ck, elev. 1660’
Permian, McCloud Fm
7635
7636
5022
5023
5024
5797
9393
863
6883
6882
6887
6884
6869
6132
cell 17
7750
STANFORD University Tyres: SMITH 477
calx, Parafusulina?: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 29, pl. 6, figs. 4 (type 7637), 5 (type
7635)
Shasta Co., Calif.; Redding Qd, NE 1/4 SE 1/4 Sec. 23, T 34 N, R 4
W, crest of Is ridge S of Potter Ck, elev. 1660’
Permian, McCloud Fm
californiensis, Anomalina: Cushman and Hobson Paratypes
Cushman and Hobson, 1935, p. 64
Santa Cruz Co., Calif.; Santa Cruz Qd, Bear Creek SU loc. 1102
Oligocene, San Lorenzo Fm
californiensis, Anomalina: Cushman and Hobson Paratype
Cushman and Hobson, 1935, p. 64
Santa Cruz Co., Calif.; Santa Cruz Qd, Kings Creek, SU loc. 1103
Oligocene, San Lorenzo Fm [missing since 1940-41 ]
californiensis, Virgulina: Cushman Paratypes
Cushman, 1925c, p. 32
San Luis Obispo Co., Calif.; Sec. 24, T 28 S, R14 E
Miocene, Monterey Shale
campbelli, Marginulina: Martin Holotype
Martin, Lewis, 1964, p. 64, pl. 5, figs. 11a, 11b
Merced Co., Calif.; Laguna Seca Hills, Laguna Seca Ck, 7 miles N
of Little Panoche Ck
Upper Cretaceous, Panoche Fm
cancriformis, Baggina: Kleinpell Holotype
Kleinpell, 1938, p. 324, pl. IX, fig. 24a-24c
Monterey Co., Calif.; Reliz Canyon
Miocene, Salinas Shale
capayana, Uvigerina pygmaea: Hedberg Paratypes
Hedberg, 1937, p. 677
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
carapitana, Bolivina aenariensis: Hedberg Paratypes
Hedberg, 1937, p. 676
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
carapitana, Cassidulina: Hedberg Paratype
Hedberg, 1937, p. 680
Anzoategui, Venezuela: District Libertad
Oligocene, Carapita Fm
carapitana, Uvigerina: Hedberg Paratypes
Hedberg, 1937, p. 667
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
carapitanus, Bathysiphon: Hedberg Paratype
Hedberg, 1937, p. 665
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
caribbeana, Nodosaria raphanistrum: Hedberg Paratypes
Hedberg, 1937, p. 671
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
carinata, Clavulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 8
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
carinata, Entosolenia circulocosta: Uchio Paratype
Uchio, 1951a, p. 37 (as circulo-costacarinata)
Chiba-ken, Japan; Hoonji, Nishi-mura, Chosei-gun
Pleistocene, Chonan Fm
478
6132
cell 26
9432
6132
cell 6
7396
9434
6088
6635
6636
6637
7939
7940
746
6132
cell 15
BuLtetTIn 300
carinatum, Haplophragmoides: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 2
Falcon, Venezuela; Curamichate, 17.6 km W of San Juan de los
Cayos, District Acosta
Miocene, Agua Salada Fm (Zone III)
caryi, Praeglobotruncana: Martin Holotype
Martin, 1964, p. 78, pl. 9, fig. 3a-3c
Eastern Panoche Hills, Calif.; Martin loc. MG 578
[missing; no record of type having been received at SU]
caudriae, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 19
Falcon, Venezuela; Pozon, 27 km E of Pueblo Jacura, District Acosta
Lower Miocene, Lower Agua Salada Fm (Zone II)
childsi, Gyroidina: Martin Holotype
Martin, Lois, 1943, p. 22, pl. 6, figs. 6a-6c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd]
Eocene, Lodo Fm
churchi, Globotruncana: Martin Holotype
Martin, Lewis, 1964, p. 79, pl. 9, figs. 5a-5c ;
Fresno Co., Calif.; eastern Panoche Hills Martin loc. MG 544
Cretaceous [missing; no record of specimen being received at SU]
cienegaensis, Nodogenerina: Kleinpell Holotype
Kleinpell, 1938, p. 244, pl. 6, fig. 4
Kern Co., Calif.; Bitter Creek
Oligocene, Maricopa Shale
colei, Dentalina: Cushman and Dusenbury Paratype
Cushman and Dusenbury, 1934, p. 54, pl. 7, fig. 10
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles S
38° W of BM 394
Eocene, Poway Conglomerate
colei, Dentalina: Cushman and Dusenbury Paratype
Cushman and Dusenbury, 1934, p. 54, pl. 7, fig. 11
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles S
38° W of BM 394
Eocene, Poway Conglomerate
colei, Dentalina: Cushman and Dusenbury Paratype
Cushman and Dusenbury, 1934, p. 54, pl. 7, fig. 12
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles S
38° W of BM 394
Eocene, Poway Conglomerate
collyra, Haplophragmoides: Nauss Holotype
Nauss, 1947, p. 337, pl. 49, fig. 2a, 2b
Alberta, Canada; Clonmel Well No. 1, Legal subdivision 1, Sec. 32,
T 55, R 20, W 4th meridian, depth 1765-1788’
Upper Cretaceous, Lloydminster Shale
collyra, Haplophragmoides: Nauss Paratype
Nauss, 1947, p. 337, pl. 49, fig. 5
Alberta, Canada; Clonmel Well No. 1, Legal subdivision 1, Sec. 32,
T 55, R 20, W 4th meridian, depth 1765-1788’
Upper Cretaceous, Lloydminster Shale
columbiensis, Pulvinulina: Cushman Paratype
Cushman, 1925d, p. 43, pl. 7, fig. la-c
Queen Charlotte Sound, British Columbia, Canada, in 20 fms
compressa, Cibicides floridanus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 26
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
5464
7692
7690
7691
7693
760
721
7418
7416
5937
7491
7430
STANFORD UNIVERSITY Types: SMITH 479
concinnum, Elphidium: Nicol Holotype
Nicol, 1944, p. 179, pl. 29, figs. 5, 6
Baja California, Mexico; San Quintin
concisa, Dunbarinella: Thompson and Hazzard Holotype
Thompson and Hazzard, 1946, pl. 42, pl. 11, fig. 9
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchells’ Caverns
Permian, Bird Spring Fm
concisa, Dunbarinella: Thompson and Hazzard Paratype
Thompson and Hazzard, 1946, p. 42, pl. 11, figs. 8, 11
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Springs Fm
concisa, Dunbarinella: Thompson and Hazzard Paratype
Thompson and Hazzard, 1946, p. 42, pl. 11, fig. 10
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
concisa, Dunbarinella: Thompson and Hazzard Paratype
Thompson and Hazzard, 1946, p. 42, pl. 11, fig. 12
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
conica, Bolivina: Cushman Paratype
Cushman, 1925c, p. 30
San Luis Obispo Co., Calif.; Sec. 24, T 28 S,R 14E
Miocene, Monterey Shale
conscripta, Lagena isabella: Cushman and Barksdale Holotype
Cushman and Barksdale, 1930, p. 65, pl. 12, fig. 4
Contra Costa Co., Calif.; Carquinez Qd, on Shell Co. property 1.5
miles E of mouth of Arroyo del Hambre, 1.1 miles S of Bull’s Head
Point, 2.35 miles N 61° W of Vine Hill
Eocene, upper Martinez Fm [missing, ca. 1940-1941] [Paleocene]
contorta, Karreriella: Beck Holotype
Beck, 1943, p. 592, pl. 98, figs. 4, 5
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2 W_ [SU loc. M-335]
Eocene, Cowlitz Fm
coombsi, Ammodiscus: Beck Holotype
Beck, 1943, p. 591, pl. 98, fig. 1
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
coralliformis, Ferayina: Frizzell Paratype
Frizzell, 1949, pp. 483-486, figs. 1-3
Bastrop Co., Texas; Smithville, S bank of Colorado River, about
0.1 mile W of bridge
Middle Eocene, Weches Fm, Claiborne Grp
cowlitzensis, Bulimina ovata: Beck Holotype
Beck, 1943, p. 605, pl. 107, fig. 22
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
cowlitzensis, Biloculina: Beck Holotype
Beck, 1943, p. 594, pl. 101. figs. 6, 7
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
480
7435
7465
7506
6132
cell 34
5020
5021
6886
7389
7950
938
855
7751
7393
BuLtetiIn 300
cowlitzensis, Robulus propinquus: Beck Holotype
Beck, 1943, p. 595, pl. 104, figs. 6, 12
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
cowlitzensis, Saracenaria mackini: Beck Holotype
Beck, 1943, p. 600, pl. 106, figs. 18, 19
Lewis Co., Wash. ; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R 2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
cowlitzensis, Siphonina claibornensis: Beck Holotype
Beck, 1943, p. 608, pl. 108, figs. 16, 18
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ [SU loc. M-335]
Eocene, Cowlitz Fm
crassa, Liebusella pozoensis: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 10
Falcon, Venezuela; Isidro, 35.7 km E of Puerto Piritu, District Zamura
Lower Miocene, Agua Salada Fm (Zone II)
crassipunctata, Cassidulina: Cushman and Hobson Paratype
Cushman and Hobson, 1935, p. 63
Santa Cruz Co., Calif. ; Bear Creek SU loc. 987
Oligocene, San Lorenzo Fm
crassipunctata, Cassidulina: Cushman and Hobson Paratype
Cushman and Hobson, 1935, p. 63
Santa Cruz Co., Calif.; on Kings Creek near San Lorenzo River, Sec.
18, T9S,R2W _ SU loc. 1103
Oligocene, San Lorenzo Fm
crebbsi, Eponides: Hedberg Paratypes
Hedberg, 1937, p. 679
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
crowleyi, Lagena: Martin Holotype
Martin, Lois, 1943, p. 18, pl. 5, figs. 5a, 5b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
cummingensis, “Verneuilina”: Nauss Holotype
Nauss, 1947, p. 341, pl. 49, fig. 4
Alberta, Canada; NW Mannville Well No. 1, Legal subdivision 1, Sec.
18, T 50, R 8 W, 4th meridian. Depth 2152-2162’
Lower Cretaceous, Mannville Fm, Cummings Mbr
cuneata, Bolivina tumida: Kleinpell Holotype
Kleinpell, 1938, p. 285, pl. XIV, figs. 9a, 9b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
cuneiformis, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 270, pl. IX, fig. 3
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
cushmani, Entosolenia marginata: Uchio Paratype
Uchio, 1951a, p. 37
Chiba-ken, Japan; Tomiya, Takeoka-mura, Kimitsu-gun
Middle Pliocene, Tomiya Fm
debilis, Bulimina: Martin Holotype
Martin, Lois, 1943, p. 20, pl. 6, figs. la-1c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
7399
6681
7388
928
6676
683
713
909
919
7456
7456a
7932
STANFORD UNIVERSITY TYPES: SMITH 481
decepta, Globigerina: Martin Holotype
Martin, Lois, 1943, p. 24, pl. 7, figs. 2a-2c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
decepta, Nodosaria: Bagg Holotype
Bagg, 1912, p. 55, pl. 16, fig. 1
San Pedro, Calif.; Timms Point [SU loc. 2024]
“Pliocene”
deliciae, Nodosaria: Martin Holotype
Martin, Lois, 1943, p. 17, pl. 6, figs. 3a, 3b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
delmonteensis, Bulimina montereyana: Kleinpell Holotype
Kleinpell, 1938, p. 255, pl. XVI, fig. 9
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
dentaliformis, Lagena: Bagg Syntypes
Bagg, 1912, p. 45, pl. 13, figs. 1a-2b
San Pedro, California; Timms Point SU loc. 2024
“Pliocene”
dubia, Buliminella: Barbat and Johnson Holotype
Barbat and Johnson, 1934, p. 13, pl. 1, fig. 14
Kern Co., Calif.; McKittrick Qd, Sec. 30, T 28 S, R 21 E; Ohio Oil
Co. Bearstate No. 23, Belridge field SU loc. 696
Miocene, Reef Ridge Shale
dubia, Buliminella: Barbat and Johnson Paratypes
Barbat and Johnson, 1934, p. 13, pl. 1, fig. 15
Kern Co., Calif.; McKittrick Qd, Sec. 30, T 28 S, R 21 E
Miocene, Reef Ridge Shale
dubia, Planularia: Kleinpell Holotype
Kleinpell, 1938, p. 207, pl. XIII, fig. 4
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
dunlapi, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 271, pl. XV, fig. 2
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
dusenburyi, Dentalina: Beck Holotype
Beck, 1943, p. 599, pl. 105, fig. 23
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W _ SU loc. M-335
Eocene, Cowlitz Fm
dusenburyi, Dentalina: Beck Paratype
Beck, 1943, p. 599, pl. 105, fig. 20
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
elkensis, Bolivina: Nauss Holotype
Nauss, 1947, p. 334, pl. 48, figs. 7a, 7b
Alberta, Canada; Imperial Core Test No. 73, in Elk Point, Legal sub-
division 9, Sec. 1, T 57, R 7 W, 4th meridian, depth 285-290’, 360’
above base of formation
Upper Cretaceous, Lea Park Shale
482
722
7409
6132
cell 13
6953
6953a
6078
5933
7911
5018
5019
761
5463
6133
cell 9
5462
7934
BULLETIN 300
eocenica, Spiroplectoides: Cushman and Barksdale Holotype
Cushman and Barksdale, 1930, p. 66, pl. 12, figs. 5a, 5b
Contra Costa Co., Calif.; Carquinez Qd, .9 miles S 78° W of Hill 187,
E of town of Martinez SU loc. 327
Eocene, Martinez Fm [Paleocene]
eponidiformis, Cibicides: Martin Holotype
Martin, Lois, 1943, p. 30, pl. 6, figs. 7a-7c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
erecta, Cassidulinoides: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 25
Falcon, Venezuela; core from Aguide Well No. 1, depth 111’, 2.5 km
S of Pueblo Aguide, District Acosta
Middle Miocene ?, upper Agua Salada Fm
ervinensis, Dunbarinella: Thompson Syntypes
Thompson, 1942, p. 419
Osage Co., Okla.; old quarry on N side of Highway 11, 3.7 miles W
of river bridge at Pawhuska ,
Pennsylvanian, Deer Creek Fm, Ervine Creek Mbr
estorffi, Nodosaria: Kleinpell Holotype
Kleinpell, 1938, p. 217, pl. 4, fig. 21
Kern Co., Calif.; Carneros Spring, W side of county
Oligocene, Temblor Fm
estorffi, Nodosaria: Kleinpell Paratype
Kleinpell, 1938, p. 217, pl. 6, fig. 5
Kern Co., Calif.; Carneros Spring, W side of county
Oligocene, Temblor Fm
etigoense, Elphidium: Husezima and Maruhasi Paratype
Husezima and Maruhasi, 1944, p. 392
Niigata-ken, Japan; Kashiwazaki Oil Field, Well No. 2, depth 94.8-
110.5 m
Pliocene, upper Haizume Fm
evolutus, Cibicides pasudoungerianus: Cushman and Hobson
Paratypes
Cushman and Hobson, 1935, p. 64
Santa Cruz Co., Calif.; Bear Creek, SU loc. 987 (type 5018); SU loc.
1102, Sec. 21, T 9 S, R 2 W (type 5019)
Oligocene, San Lorenzo Fm
excolata, Guembelina: Cushman Paratype
Cushman, 1926c, p. 20
San Luis Potosi, Mexico; near Coco, on Tampico R.R.
Cretaceous, Mendez Shale
excubitor, Elphidium: Nicol Holotype
Nicol, 1944, p. 178, pl. 29, figs. 4, 8
Sonora, Mexico; Punta Penasco, lat. 31° 21’ N
falconensis, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 3
Lara, Venezuela; central Falcon, 16.5 km N of Siquisique, District
Urdaneta
Lower Miocene, Agua Salada Fm (Zone II)
fax, Elphidium fax: Nicol Holotype
Nicol, 1944, p. 177, pl. 29, figs. 3, 11
Clallam Co., Wash.; Dallas Bank, Straits of Juan de Fuca
fax, Epistomina: Nauss Holotype
Nauss, 1947, p. 335, pl. 48, fig. 16
Alberta, Canada; Vermilata Frankview Well. No. 1. Legal subdivision
16, Sec. 28, T 50, R 5 W, 4th meridian, depth 660-670’, 180-190’ above
base of fm
Upper Cretaceous, Lea Park Shale
7935
6722
6723
6724
6725
7410
6133
cell 15
7411
9436
6115
7817
7818
7819
7822
7823
7820
7821
6092
590
STANFORD UNIVERSITY TYPEs: SMITH 483
fax, Epistomina: Nauss Paratype
Nauss, 1947, p. 335, pl. 48, fig. 15
Alberta, Canada; Vermilata Frankview Well No. 1, Sec. 28, T 50,
R 5 W, 4th meridian, depth 660-670’, 180-190’ above base of fm
Upper Cretaceous, Lea Park Shale
fax, Schwagerina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 27, pl. 1, figs. 1 (type 6723), 2 (type
6724), 3 (type 6722), 4 (type 6725)
Shasta Co., Calif.; Redding Qd, W side of limestone hogback, NW 1/4,
NE 1/4 Sec. 15, T 33 N, R 4 W. Elev. 1400’ SU loc. 774
Permian, McCloud Fm
felix, Cibicides: Martin Holotype
Martin, Lois, 1943, p. 31, pl. 8, figs. 7a-7c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
flexilis, Valvulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 7
Falcon, Venezuela, Aguide, 3.85 km SE of Pueblo Aguide, District
Acosta
Lower Miocene, Agua Salada Fm (Zone II)
fortunatus, Cibicides: Martin Holotype
Martin, Lois, 1943, p. 31, pl. 8, figs. 5a-5c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
fresnoensis, Globotruncana: Martin Holotype
Martin, Lewis, 1964, p. 80, pl. 9, figs. 8a-8d
Fresno Co., Calif.; eastern Panoche Hills, Martin loc. MG 574
Cretaceous, Panoche Group, upper Marlife Shale [missing; no record
of type having been deposited at SU]
frizzelli, Eponides: Kleinpell Holotype
Kleinpell, 1938, p. 318, pl. 2, figs. 15, 16
Santa Barbara Co., Calif.; near Gaviota Pass SU loc. 1436
Oligocene, “Sespe” Fm
furoni, Schwagerina: Thompson Holotype
Thompson, 1946, p. 147, pl. 24, fig. 7
Afghanistan; Shibar Pass, on road from Kabul to Bamian, about 7
km W of summit SU loc. 2612
Permian, Bamian ls
furoni, Schwagerina: Thompson Paratypes
Thompson, 1946, p. 147, pl. 23, figs. 1 (type 7818), 2 (type 7819), 3
(type 7822), 4 (type 7823) ; pl. 24, fig. 10 (tye 7822)
Afghanistan; Shibar Pass, on road from Kabul to Bamian, about 7
km W of summit SU loc. 2612
Permian, Bamian ls
furoni, Schwagerina: Thompson Paratypes
Thompson, 1946, p. 147, pl. 24, figs. 8 (type 7821), 9 (type 7820)
Afghanistan; Shibar Pass, on road from Kabul to Bamian, about 7
km W of summit SU loc. 2612
Permian, Bamian Is
galliheri, Bulimina: Kleinpell Holotype
Kleinpell, 1938, p. 253, pl. 17, figs. 2, 5
Monterey Co., Calif.; Monterey Qd, 1 mile N of Carmel SU loc. 333
Miocene, Monterey Shale
gallowayi, Cibicides: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 30
Channel Islands, off southern California
484
689
691
692
6130
7427
749
906
925
9435
7650
7646
7647
7648
7649
8315
BULLETIN 300
galvestonensis, Elphidium gunteri: Kornfeld Syntype
Kornfield, 1931, p. 87, pl. 15, figs. 3a, 3b
Galveston Co., Texas; E end of Galveston Island in beach sand
galvestonensis, Elphidium gunteri: Kornfeld Syntype
Kornfeld, 1931, p. 87, pl. 15, figs. 1a, 1b
Galveston Co., Texas; E end of Galveston Island in beach sand
galvestonensis, Elphidium gunteri: Kornfeld Syntype
Kornfeld, 1931, p. 87, pl. 15, figs. 2a, 2b
Galveston Co., Texas; E end of Galveston Island in beach sand
galvinensis, Cassidulina: Cushman and Frizzell Paratype
Cushman and Frizzell, 1940, p. 43
Lewis Co., Wash.; R.R. cuts .25 miles N of Galvin, SW 1/4 Sec. 28,
T15N,R3 W SU loc. 1167
Oligocene, Lincoln Fm
gilboei, Triloculina: Beck Holotype
Beck, 1943, p. 594, pl. 101, figs. 1-3
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ SU loc. M-335
Eocene, Cowlitz Fm
glabrata, Ancmalina: Cushman Paratype
Cushman, 1924, p. 39, pl. 12, figs. 5-7
Pago Pago, Samoa, in 50 fms
globosa, Baggina robusta: Kleinpell Holotype
Kleinpell, 1938, p. 326, pl. XIII, figs. 2a-2c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
globula, Pullenia miocenica: Kleinpell Holotype
Kleinpell, 1938, p. 340, pl. XVI, figs. 2a, 2b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
goudkoffi, Globotruncana: Martin Holotype
Martin, Lewis, 1964, p. 80, pl. 10, figs. la-1c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
gracilis, Fusulina: Meek Neotype
Meek, 1864b, p. 4, pl. 2, figs. 1-1c. Neotype designated by Thomp-
son and Wheeler, 1946, p. 31, pl. 1, fig. 10 [as Parafusulina]
Shasta Co., Calif.; Redding Qd, W side of limestone hogback, NE
1/4, NE 1/4 Sec. 10, T 33 N, R 4 W, elev. 1660’
Permian, McCloud Fm
gracilis, Fusulina: Meek Paraneotypes
Meek, 1864, p. 4, illustrated by Thompson and Wheeler, 1946, p. 31,
pl. 1, figs. 6, 7 [as Parafusulina]
Shasta Co., Calif.; Redding Qd, W side of limestone hogback, NE
1/4, NE 1/4 Sec. 10, T 33 N, R 4 W, elev. 1660’
Permian, McCloud Fm
gracilis, Fusulina: Meek Paraneotypes
Meek, 1864b, p. 4, illustrated by Thompson and Wheeler, 1946, p. 31,
Plow. eties: 8) 9
Shasta Co., Calif.; Redding Qd, W side of limestone hogback, NE
1/4, NE 1/4 Sec. 10, T 33 N, R 4 W, elev. 1660’
Permian, McCloud Fm
grahami, Eofabiania: Kiipper Holotype
Kipper, 1955, p. 136, pl. 19, fig. 4; text fig. 1, fig. 1
Santa Clara Co., Calif.; New Almaden Qd, near Old Guadalupe
quicksilver mine SU loc. 309
Middle Eocene
6132
cell 35
6872
6079
60380
7500
7811
7812
7813
7814
7815
7816
STANFORD UNIVERSITY Types: SMITH 485
grahami, Eofabiania: Kupper Paratypes
Kiipper, 1955, p. 136, text fig. 1, pl. 19, figs. 1 (type 8317), 2, 3 (type
8318), 5 (type $316), 6, 7 (type 8319)
Santa Clara Co., Calif.; New Almaden Qd, near old Guadalupe
quicksilver mine SU loc. 309
Middle Eocene
grahami, Gyroidinoides: Martin Holotype
Martin, Lewis, 1964, p, 95, pl. 13, figs. la-1c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
grenadana, Textularia: Hedberg Paratype
Hedberg, 1937, p. 667
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
grimsdalei, Lingulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 14
Falcon, Venezuela; Tocuyo, 17.7 km § of San Juan de los Cayos,
District Acosta
Upper Oligocene, Agua Salada Fm (Zone I)
halconi, Heterostomella (?): Hedberg Paratype
Hedberg, 1937, p. 667
Anzoategui, Venezuela; District Libertad
Oligocene, Carapite Fm
hamilli, Nodosaria: Kleinpell Holotype
Kleinpell, 1938, p. 218, pl. 4, fig. 5
Kern Co., Calif.; Bitter Creek
Oligocene, Vaqueros Fm
hamilli, Nodosaria: Kleinpell Paratype
Kleinpell, 1938, p. 218, pl. 4, fig. 4
Kern Co., Calif.; Bitter Creek
Oligocene, Vaqueros Fm
hannai, Angulogerina: Beck Holotype
Beck, 1943, p. 607, pl. 108, figs. 26, 28
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2 W_ SU loc. M-335
Eocene, Cowlitz Fm
hannai, Uvigerina: Kleinpell Holotype
Kleinpell, 1938, p. 294
Monterey Co., Calif.; Monterey Qd, 108.7 mm E, 105 mm N of inter-
section of lat. 36° 30’, long. 121° 55’ on map, 25-40’ from top of hill
SU loc. 336
Miocene, Monterey Shale, type locality, upper Mohnian Stage
haydeni, Yangchienia: Thompson Holotype
Thompson, 1946, p. 146, pl. 23, fig. 6
Afghanistan; SW of summit of Shibar Pass, 7 km on road from Kabul
to Bamian SU loc. 2612
Permian, Bamian Ls
haydeni, Yangchienia: Thompson Paratypes
Thompson, 1946, p. 146, pl. 23, figs. 1, 11 (type 7812), 8 (type 7813),
5 (type 7814)
Afghanistan; Shibar Pass, on road from Kabul to Bamian, about 7
km W of summit SU loc. 2612
Permian, Bamian Ls
haydeni, Yangchienia: Thompson Paratypes
Thompson, 1946, p. 146, pl. 23, figs. 9 (type 7816), 10 (type 7815)
Afghanistan; Shibar Pass, on road from Kabul to Bamian, about 7
km W of summit SU loc. 2612
Permian, Bamian Ls
486
7936
6132
cell 25
6134
cell 39
7495
6132
cell 12
412
417
663
71927
7914
9349
BULLETIN 300
hectori, Gaudryina: Nauss Holotype
Nauss, 1947, p. 335, pl. 48, figs. 6a, 6b
Alberta, Canada; NW Mannville Well No. 1 in Legal subdivision 1,
Sec. 18, T 50, R 8 W, 4th meridian, depth 1806-1813’, 20-25’ above
base of formaticn
Upper Cretaceous, Lloydminster Shale
hedbergi, Robulus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 10
Falcon, Venezuela; Tocuyo, 18.0 km S of San Juan de los Cayos,
District Acosta
Upper Oligocene, Agua Salada Fm (Zone I)
herberti, Trifarina: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 9
Trinidad, B.W.I.; Soldado Rock
Paleocene, Soldado Fm
hobsoni, Virgulina: Beck Holotype
Beck, 1943, p. 606, pl. 107, figs. 6, 10
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
horizontalis, Cassidulina subglobosa: Cushman and Renz
Paratype
Cushman and Renz, 1941, p. 26
Falcon, Venezuela; Isidro, 35.7 km E of Pueblo Piritu, District Zamura
Middle Miocene, upper Agua Salada Fm (Zone III)
hughesi, Elphidium: Cushman and Grant Holotype
Cushman and Grant, 1927, p. 75, pl. 7, fig. 1
Monterey Co., Calif.; Pine Valley, N 1/2 Sec. 12, T 21S,R 10E
Lower Pliocene, Pancho Rico Fm
hughesi, Elphidium: Cushman and Grant Paratypes
Cushman and Grant, 1927, p. 75
Monterey Co., Calif.; Pine Valley, N 1/2 Sec. 12, T 21$,R 10E
Lower Pliocene, Pancho Rico Fm
hughesi, Robulus: Kleinpell Holotype
Kleinpell, 1938, p. 198, pl. 7, figs. 18a, 18b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
humei, Ammobaculites: Nauss Holotype
Nauss, 1947, p. 333, pl. 48, fig. 1
Alberta, Canada; NW Mannville Well No. 1 in Legal subdivision 1,
Sec. 18, T 50, R 8 W, 4th meridian
Lower Cretaceous, Mannville Fm, Cummings Mbr
ikebei, Angulogerina: Husezima and Maruhasi Paratype
Husezima and Maruhasi, 1944, p. 396
Niigata-ken, Japan; Kashiwazaki Oil Field, well no. 1, 200 m
Pliocene, lower Haizume Fm
illingi, Angulogerina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 21
Falcon, Venezuela; Tocuyo, 16.9 km § of San Juan de los Cayos,
District Acosta
Miocene, Agua Salada Fm (Zone IIT)
imbricata, Bolivina: Cushman Paratype
Cushman, 1925c, p. 31
San Luis Obispo Co., Calif.; Sec. 24, T 28S, R14E
Miocene, Monterey Shale
impensus, Haplophragmoides: Martin Holotype
Martin, Lewis, 1964, p. 48, pl. 2, figs. 3a, 3b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
9349a
9350
9350a
6132
cell 8
6666
6132
cell 29
6089
6132
cell 3
6132
cell 14
6132
cell 27
6133
cell 5
6133
cell 12
STANFORD UNIVERSITY Tyres: SMITH 487
impensus, Haplophragmoides: Martin Paratype
Martin, Lewis, 1964, p. 48, pl. 2, figs. 4, 4b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
incognatus, Haplophragmoides: Martin Holotype
Martin, Lewis, 1964, p. 49, pl. 2, figs. 6a, 6b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
incognatus, Haplophragmoides: Martin Paratype
Martin, Lewis, 1964, p. 49, pl. 2, figs. 7a, 7b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
inconspicua, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 18
Falcon, Venezuela; Aguide, 2.5 km S of Pueblo Aguide, core from
Aguide well no. 1, 1646’, District Acosta
Miocene ?, Agua Salada Fm (Zone II?)
involuta, Valvulineria: Cushman and Dusenbury Holotype
Cushman and Dusenbury, 1934, p. 63, pl. 8, figs. 12a-12c
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles
S 38° W of BM 394 SU loc. 1150
Eocene, Poway Conglomerate
irregularis, Gaudryina jacksonensis: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 6
Falcon, Venezuela; Araurima, 11.6 km SE of Pueblo Jacura, District
Acosta
Upper Oligocene, Agua Salada Fm (Zone [)
irregularis, Nodogenerina: Kleinpell Holotype
Kleinpell, 1938, p. 245, pl. 17, fig. 12
Santa Barbara Co., Calif.; near Naples
Miocene, Monterey Shale
isidroensis, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 17
Falcon, Venezuela; Isidro, 33.75 km E of Pueblo Piritu, District
Zamura
Miocene, Agua Salada Fm (Zone IV)
isidroensis, Cibicides: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 26
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
isidroensis, Dentalina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 15
Falcon, Venezuela; Isidro, 34.4 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone III)
isidroensis, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 4
Falcon, Venezuela; Pozon, 21.1 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
isidroensis, Uvigerina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 20
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
6133
cell 18
6132
cell 39
9486
9451
6107
6108
7406
7407
7385
BULLETIN 300
jacksonensis, Discorbis: Cushman and Applin Paratype
Cushman and Applin, 1926, p. 178, pl. 9, figs. 8, 9
San Augustine Co., Texas; Bridge Ck, 1.5 miles above Angelina
River
Upper Eocene, Jackson group
jacuraensis, Vulvulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 5
Falcon, Venezuela; 11.65 km SE of Pueblo Jacura, District Acosta
Lower Miocene, Agua Salada Fm (Zone II)
japonica, Pseudoeponides: Uchio Paratype
Uchio, 1950, p. 190, text fig. 16
Chosei-gun, Chiba Prefecture, Japan; in fine sand along Otaki-
Chonan Highway, beside a bridge 200 m SE of Primary School,
Satsubo, Nishi-mura
Upper Pliocene, Kakinokidae Fm
jarvisi, Hastigerinella: Cushman Paratype
Cushman, 1930, p. 18
Trinidad, B.W.I.; 17.25 miles out on Cunapo Southern Road
Middle Eocene, Navet Fm
jarvisi, Pulvinulinella: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 24
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone II)
jarvisi, Valvulineria: Martin Holotype
Martin, Lewis, 1964, p. 103, pl. 15, figs. 4a-4c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
joaquinensis, Bulimina: Martin Holotype
Martin, Lewis, 1964, p. 87, pl. 11, figs. 5a, 5b, 6a, 6b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
joaquinensis, Uvigerina: Kleinpell Holotype
Kleinpell, 1938, p. 296, pl. 17, figs. 6, 10
Kern Co., Calif.; Chico Martinez Creek
Miocene, Monterey Shale
joaquinensis, Uvigerina: Kleinpell Paratype
Kleinpell, 1938, p. 296, pl. 17, fig. 11
Kern Co., Calif.; Chico Martinez Creek
Miocene, Monterey Shale
judas, Anomalina: Martin Holotype
Martin, Lois, 1943, p. 28, pl. 7, figs. 4a-4c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hill Qd] SU
loc. M-74
Eocene, Lodo Fm
keenae, Anomalina: Martin Holotype
Martin, Lois, 1943, p. 29, pl. 7, figs. 5a-5c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
kelleyi, Vaginulinopsis, Martin Holotype
Martin, Lois, 1943, p. 15, pl. 5 figs. 8a-8c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
8113
5794
7434
7497
6129
6134
cell 32
6132
cell 16
6133
cell 1
6133
cell 10
5920
6133
cell 7
987
6133
cell 17
STANFORD UNIVERSITY Types: SMITH 489
kentuckyensis, Hyperammina: Conkin Paratype
Conkin, 1954, p. 166
Jefferson Co., Kentucky; Mitchell Kill, SW part of the county
Mississippian, Floyds Knob Fm
kernensis, Nonion incisum: Kleinpell Holotype
Kleinpell, 1938, p. 232
Kern Co., Calif.; Carneros Springs SU loc. 675
Miocene, Temblor Fm
kincaidi, Robulus: Beck Holotype
Beck, 1943, p. 595, pl. 102, figs. 1, 7
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2 W_ SU loc. M-335
Eocene, Cowlitz Fm
kleinpelli, Bolivina: Beck Holotype
Beck, 1943, p. 606, pl. 107, fig. 39
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R 2 W SU loc. M-335
Eocene, Cowlitz Fm
kleinpelli, Eponides: Cushman and Frizzell Paratype
Cushman and Frizzell, 1940, p. 42
Lewis Co., Wash.; on R.R. .25 miles N of Galvin, SW 1/4 Sec. 27, T
15 N,R 3 W_ SU loc. 1167
Oligocene, Lincoln Fm
kugleri, Bulimina: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 9
Trinidad B.W.I.; Soldado Rock
Paleocene, Soldado Fm
kugleri, Cibicides: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 27
Falcon, Venezuela; Pozon, 20.8 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone III)
kugleri, Siphogenerina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 22
Falcon, Venezuela; Pozon, 20.2 km SE of Pueblo Jacura, District
Acosta
Lower Miocene, Agua Salada Fm (Zone II)
kugleri, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 5
Falcon, Venezuela; Aguide, 12.3 km S of Pueblo Aguide, District
Acosta
Lower Miocene, Agua Salada Fm (Zone II)
laimingi, Plectofrondicularia miocenica: Kleinpell Holotype
Kleinpell, 1938, p. 241
Ventura Co., Calif.; Los Sauces Creek
Oligocene, Rincon Shale, middle mbr, Lower Saucesian stage
[specimen missing since 1942]
lalickeri, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 3
Falcon, Venezuela; Aguide, 2.5 km S of Pueblo Aguide, District
Acosta; from well core at 237’ depth
Lower Miocene, Agua Salada Fm (Zone II)
lata, Nonionella: Cushman and Valentine Paratype
Cushman and Valentine, p. 20
Channel Islands off southern California
lehneri, Globorotalia: Cushman and Jarvis Paratype
Cushman and Jarvis, 1929, p. 17
Trinidad, B.W.I.; source of Moruga River
Middle Eocene, Navet Fm
490
6133
cell 16
6730
6731
6133
cell 6
7431
7453
5630
7943
9333
6132
cell 11
7398
7384
7395
BuLLeETIN 300
lehneri, Hantkenina: Cushman and Jarvis Paratype
Cushman and Jarvis, 1929, p. 16
Trinidad, B.W.I.; source of Moruga River
Middle Eocene, Navet Fm
lepida, Schwagerina: Schwager Syntypes
Schwager im Richthofen, 1883, p. 138. Also zz Schenck and Thompson,
1940, p. 588
Hupei Province, China; right bank of Yangtze River, opposite Ki-
tschou
Permian
leuzingeri, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 3
Falcon, Venezuela; Isidro, 34.0 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
lewisensis, Cornuspira: Beck Holotype
Beck, 1934, p. 594, pl. 101, figs. 4, 5
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ SU loc. M-335
Eocene, Cowlitz Fm
lewisensis, Vaginulinopsis saundersi: Beck Holotype
Beck, 1943, p. 598, pl. 105, figs. 3, 13
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ SU loc. M-335
Eocene, Cowlitz Fm
limbata, Cassidulina: Cushman and Hughes Paratype
Cushman and Hughes, 1925, p. 12
San Pedro, Calif.; Timms Point SU loc. 2024
“Pliocene,” Timms Point Fm
linki, Haplophragmoides: Nauss Holotype
Nauss, 1947, p. 339, pl. 49, figs. 7a, 7b
Alberta, Canada; Dina Omega Well No. 1, legal subdivision 14, Sec.
9, T 45, R 1 W, 4th meridian, depth 1478-1488’, 17-27’ above base of
fm
Upper Cretaceous, Lloydminster Shale
Ilanadoensis, Psammosiphonella: Martin Holotype
Martin, Lewis, 1964, p. 43, pl. 1, figs. 4a, 4b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch surface section,
Martin loc. MG 574
Cretaceous, Panoche Group, upper Marlife Shale [missing; no record
of type having been received at SU]
lobata, Valvulineria inaequalis: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 23
Falcon, Venezuela; Pozon, 20.3 km SE of Pueblo Jacura, District
Acosta
Lower Miocene, lower Agua Salada Fm (Zone II)
lodoensis, Eponides: Martin Holotype
Martin, Lois, 1943, pl. 6, figs. 8a-8c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
lodoensis, Palmula henbesti: Martin Holotype
Martin, Lois, 1943, p. 15, pl. 9, figs. la 1b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
lodoensis, Uvigerina: Martin Holotype
Martin, Lois, 1943, p. 21, pl. 6, figs. 2a, 2b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
7382
7937
841
7464
6950
763
7402
726
7682
STANFORD UNIVERSITY Types: SMITH 491
lodoensis, Zeauvigerina: Martin Holotype
Martin, Lois, 1943, p. 21, pl. 5, figs. 1a, 1b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
loetterli, Globigerina: Nauss Holotype
Nauss, 1947, p. 336, pl. 49, figs. 1la-11c
Alberta, Canada; Clonmel Well No. 1, Legal subdivision 1, Sec. 32,
T 55, R 20 W, 4th meridian, W of Vermilion area; depth 1875-1885’,
485-495’ below top of fm
Upper Cretaceous Lloydminster Shale
luciana, Planularia: Kleinpell Holotype
Kleinpell, 1938, p. 207, pl. IX, figs. 25a, 25b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
mackini, Saracenaria: Beck Holotype
Beek, 1943, p. 600, pl. 106, figs. 1, 5
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
marblensis, Millerella: Thompson Paratypes
Thompson, 1942, p. 405
Marble Falls, Texas; near water level, 150 yds downstream from
bridge
Pennsylvanian, Marble Falls Ls
marginata, Bolivina: Cushman Paratype
Cushman, 1925c, p. 30
San Luis Obispo Co., Calif.; Sec. 24, T 28 S, R 14 E
Miocene, Monterey Shale
marksi, Globorotalia: Martin Holotype
Martin, Lois, 1943, p. 26, pl. 8, figs. 1a-1c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
martinezensis, Cibicides: Cushman and Barksdale Holotype
Cushman and Barksdale, 1930, p. 68, pl. 12, figs. 9a-9c
Contra Costa Co., Calif.; Carquinez Qd, on Shell Co. property 1.5
miles E of mouth of Arroyo del Hambre, 1.1 mile S of Bull’s Head
Point and 2.35 miles N 61° W of Vine Hill SU loc. 322
Eocene, upper Martinez Fm_ [Paleocene ]
masoni, Schubertella: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 41, pl. 13, figs. 7, 8 (type 7682), 10
(type 7684), 9 (type 7683)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
matagordana, Nonion depressula: Kornfeld Holotype
Kornfeld, 1931, p. 87, pl. 13, figs. 2a, 2b
Matagorda Co., Texas; Gulf of Mexico, in beach sand
mauryae, Cancris: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 11
Trinidad, B.W.I.; Soldado Rock
Paleocene, Soldado Fm
mayori, Calcarina: Cushman Paratype
Cushman, 1924, p. 44, pl. 14, figs. 4-7
Pago Pago, Samoa, in 18 fms
492
7520
9795
6132
cell 20
5634
7942
7423
9466
7331a
7331b
7331¢c
7331d
6907
BuLLeETIN 300
memastersi, Cibicides: Beck Holotype
Beck, 1943, p. 612, pl. 109, figs. 2, 4, 15
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
mediocostata, Nonionina: Cushman Paratype
Cushman, 1926b, p. 89, pl. 13, figs. la-1c, as medio-costata
San Luis Obispo Co., Calif.; Sec. 24, T 28 S, R14E
Miocene, Monterey Shale
melvilli, Robulus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 12
Falcon, Venezuela; Isidro, 33.5 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
mexicana, Bolivina: Cushman Paratypes
Cushman, 1926a, p. 8i
Vera Cruz, Mexico; Panuco-Tampico R.R., between km posts 21 and
22
Eocene-Oligocene ?, Alazan Clay
mexicana, Haplophragmoides canariensis: Kornfeld Neotype
Kornfeld, 1931, p. 83, pl. 13, figs. 4a-4c
Cameron Parish, Louisiana: E of Calcasieu Pass, in beach sand
mexicanum, Elphidium incertum: Kornfeld Syntype
Kornfeld, 1931, p. 89, pl. 16, figs. 2a, 2b
Galveston Co., Texas; E end of Galveston Island, in beach sand
mexicanum, Elphidium incertum: Kornfeld Syntype
Kornfeld, 1931, p. 89, pl. 16, figs. 1a, 1b
Galveston Co., Texas; E end of Galveston Island, in beach sand
milleri, Quinqueloculina: Beck Holotype
Beck, 1943, p. 593, pl. 99, figs. 8, 9, 10
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
minor, Haplophragmoides gigas: Nauss Holotype
Nauss, 1947, p. 338, pl. 49, figs. 10a, 10b
Alberta, Canada; NW Mannville Well No. 1 in Legal subdivision 1,
Sec. 18, T 50, R 8 W, 4th meridian, depth 2173-2183’, 5-51’ above base
of member
Lower Cretaceous, Mannville Fm, Cummings mbr
minuta, Quinqueloculina: Beck Holotype
Beck, 1943, p. 593, pl. 99, figs. 5-7
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W _ SU loc. M-335
Eocene, Cowlitz Fm
minuta, Rotalia: Martin Holotype
Martin, Lewis, 1964, p. 94, pl. 12, figs. 1la-11c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
minuta, Yabeina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1942, p. 707, pl. 106, figs. 6, 8 (type 7331a),
7 (type 7331b), 9 (type 7331c), 10 (type 7331d)
Marble Canyon, southern British Columbia, Canada
Permian, Marble Canyon Ls
miocenica, Cristellaria: Chapman Holotype
Chapman, 1900, p. 250, pl. 30, figs. 1, la
Santa Clara Co., Calif.; from a well
Miocene
9412
7701
7702
7703
7704
7705
6071
7743
893
5976
5977
5978
6720
6122
Hel)
7720
T7121
7722
STANFORD UNIVERSITY TYPES: SMITH 493
miocenica, Pullenia: Kleinpell Holotype
Kleinpell, 1938, p. 338, pl. 14, fig. 6
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
mirabilis, Planularia: Martin Holotype
Martin, Lewis, 1964, p. 71, pl. 7, figs. 5a-5c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, + miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
modica, Schwagerina: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 44, pl. 11, figs. 1 (type 7701), 2
(type 7702), 3, 6 (type 7703)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
modica, Schwagerina: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 44, pl. 11, figs. 4, 5 (type 7704), 7
(type 7705)
San Bernardino Co., Calif.; E front, Providence Mts., 1.5 miles N
of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
mohnensis, Robulus: Kleinpell Holotype
Kleinpell, 1938, p. 200, pl. 18, figs. 1, 2
Los Angeles Co., Calif.; Mohn Springs
Miocene, Modelo Shale
momiyamensis, Elphidiella: Uchio Paratype
Uchio, 1951b, p. 372, pl. 5. figs. 7a, 7b
Tochigi Prefecture, Japan; in cliff facing Tobu electric R.R., 600 m
NW of Momiyama Station
Miocene, Kanuma Fm
montereyana, Bulimina: Kleinpell Holotype
Kleinpell, 1938, p. 254, pl. XII, fig. 13
Monterey Co., Caiif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
montis, Neofusulinella: Thompson and Wheeler Holotype
Thompson and Wheeler, 1946, p. 26, pl. 2, figs. 7, 8
Shasta Co., Calif.; Redding Qd, crest of limestone ridge S of Potter
Creek, elev. 1675’, NE 1/4, SE 1/4 Sec. 23, T 34 N, R 4 W. SU loc.
757
Permian, McCloud Fm
montis, Neofusulinella: Thompson and Wheeler Paratypes
Thompson and Wheeler, 1946, p. 26, pl. 2, figs. 5 (type 5977), 6 (type
5978), 9 (type 6720)
Shasta Co., Calif.; Redding Qd, crest of limestone ridge S of Potter
Creek, elev. 1675’, NE 1/4 SE 1/4 Sec. 23, T 34 N, R 4 W. SU loc.
7 sy7
Permian, McCloud Fm
moorei, Pulienia: Kleinpell Holotype
Kieinpell, 1938, p. 340, pl. 18, figs. 11, 16
Santa Barbara Co., Calif.; near Naples
Miocene, Monterey Shale
multispira, Schwagerina?: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 46, pl. 15, figs. 1 (type 7719), 2
(type 7720), 3 (type 7721), 4 (type 7722)
San Bernardino Co., Calif.; along ridge and summit of main divide,
Providence Mts., ca. 1.25 miles W of mouth of large canyon opening
just N of Gilroy Mine
Permian, Bird Spring Fm
494
7748
7521
7403
6951
7401
7912
7400
6890
7661
7662
7663
7664
7665
7666
7667
7668
7669
7670
5786
BuLueTIN 300
nakamurai, Cassidulina: Uchio Paratype
Uchio, 1950, p. 190, text fig. 14
Chiba-ken, Japan; S entrance of Odoroa Tunnel, Kamitaki-mura,
Isumi-gun
Upper Pliocene
natlandi, Cibicides: Beck Holotype
Beck, 1943, p. 612, pl. 109, figs. 2, 4, 15
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2 W_ SU loc. M-335
Eocene, Cowlitz Fm
naussi, Globorotalia: Martin Holotype
Martin, Lois, 1943, p. 26, pl. 8, figs. 3a-3c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
needhami, Pseudostaffella: Thompson Syntype
Thompson, 1942, p. 411
NW side of Mud Springs Mt., New Mexico; W end of Whiskey
Canyon
Pennsylvanian, Magdalena Fm, Bend Series
nicoli, Globorotalia: Martin Holotype
Martin, Lois, 1943, p. 27, pl. 7, figs. 3a-3c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
niigataensis, Eponides: Husezima and Maruhasi Paratype
Husezima and Maruhasi, 1944, p. 398
Niigata-ken, Japan; Kashiwazaki Oil Field, well no. 1, depth 94.8-
110.5 m
Pliocene, upper Haizume Fm
nitida, Globigerina: Martin Holotype
Martin, Lois, 1943, p. 25, pl. 7, figs. la-1c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
nolani, Anomalina: Hedberg Paratypes
Hedberg, 1937, p. 681
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
nosonensis, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 33
Shasta Co., Calif.; Redding Qd, SW 1/4, SE 1/4 Sec. 22, T 33 N,
R 4 W, elev. 180’ above base of Nosoni Fm_ SU loc. 2577
Permian, Nosoni Fm
nosonensis, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 33, pl. 7, figs. 1 (type 7663), 2 (type
7664), 3 (type 7665), 4 (type 7666)
Shasta Co., Calif.; Redding Qd, SW 1/4, SE 1/4 Sec. 22, T 33 N,
R4W _ SU loc. 2577
Permian, Nosoni Fm
nosonensis, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 33, pl. 7, figs. 5, 9 (type 7667), 6
(type 7668), 7 (type 7669), 8, 10 (type 7670)
Shasta Co., Calif.; Redding Qd, SW 1/4, SE 1/4 Sec. 22, T 33 N,
R4W _ SU loc. 2577
Permian, Nosoni Fm
nuciformis, Siphogenerina: Kleinpell Holotype
Kleinpell, 1938, p. 303, pl. 15, fig. 10
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
9787
6879
6132
cell 21
5798
703
712
9366
9366a
9492
5972
9973
5974
5975
7522
STANFORD UNIVERSITY TyPEs: SMITH 495’
nuciformis, Siphogenerina: Kleinpell Paratype
Kleinpell, 1938, p. 303, pl. 14, fig. 12
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
nuttalli, Nodosaria: Hedberg Paratypes
Hedberg, 1937, p. 673
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
nuttalli, Robulus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 11
Falcon, Venezuela; Isidro, 35 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone III)
obesa, Uvigerinella: Cushman Paratype
Cushman, 1926d, p. 59
San Luis Obispo Co., Calif.; Sec. 24, T 28 S,R14E
Miocene, Monterey Shale
obliqua, Bolivina: Barbat and Johnson Holotype
Barbat and Johnson, 1934, p. 15, pl. 1, fig. 20
Kings Co., Calif.; Coalinga Qd, Kettleman Hills, Sec. 35, T 21 S,
R 17 E, Associated Oil Co., Whepley No. 1
Miocene, Reef Ridge Shale
obliqua, Bolivina: Barbat and Johnson Paratype
Barbat and Johnson, 1934, p. 15
Kings Co., Calif.; Coalinga Qd, Kettleman Hills, Sec. 35, T 21 S,
R 17 E, Associated Oil Co., Whepley No. 1
Miocene, Reef Ridge Shale
obscura, Eggerella: Martin Holotype
Martin, Lewis, 1964, p. 55, pl. 3 figs. 10a, 10b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panocke Fm
obscura, Eggerella: Martin Paratype
Martin, Lewis, 1964, p. 55, pl. 3 fig. 11
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
occidentalis, Anomalina: Martin Holotype
Martin, Lewis, 1964, p. 105, pl. 16, figs. 3a-3c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
occidentalis, Neofusulinella: Thompson and Wheeler Holotype
Thompson and Wheeler, 1946, p. 25, pl. 2, fig. 2
Shasta Co., Calif.; Redding Qd, “Bass Ranch,” SE 1/4, SE 1/4 Sec.
15, T 33 N, R 4 W, elevation 1100’ SU loc. 775
Permian, McCloud Fm
occidentalis, Neofusulinella: Thompson and Wheeler Paratypes
Thompson and Wheeler, 1946, p. 25, pl. 2, figs. 1 (type 5975), 3 (type
5974), 4 (type 5973)
Shasta Co., Calif.; Redding Qd, “Bass Ranch,” SE 1/4, SE 1/4 Sec.
15, T 33 N, R 4 W, elevation 1100’ SU loc. 775
Permian, McCloud Fm
olequaensis, Cibicides natlandi: Beck Holotype
Beck, 1943, p. 612, pl. 109, figs. 3, 20, 22
Lewis Co., Wash.; 1.5 miles E of Vader, E 1/2, SE 1/4 Sec. 28, T 11
N, R 2 W, on W bank of Cowlitz River SU loc. M-335
Eocene, Cowlitz Fm
496
7420
6874
754
5802
7741
7742
5939
7417
582
7333a
7333e
7333b
7333¢
7333d
945
6889
6878
BULLETIN 300
olequaensis, Quinqueloculina goodspeedi: Beck Holotype
Beck, 1943, p. 592, pl. 99, figs. 3, 4
Lewis Co., Wash.; 1.5 miles E of Vader, E 1/2, SE 1/4 Sec. 28, T 11
N, R 2 W, on W bank of Cowlitz River SU loc. M-335
Eocene, Cowlitz Fm
orinocoensis, Sigmoilina: Hedberg Paratype
Hedberg, 1937, p. 669
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
ornata, Bolivina advena: Cushman Paratype
Cushman, 1925c, p. 29
San Luis Obispo Co., Calif.; Sec. 24, T 28 S, R14 E
Miocene, Menterey Shale
ornata, Valvulineria: Cushman Paratypes
Cushman, 1926d, p. 61
San Luis Obispo Co., Calif.; Sec. 24, T 28S, R14E
Miocene, Monterey Shale
otukai, Vaginulina: Uchio Paratype
Uchio, 1951b, p. 370, pl. 5, figs. 4a-4e
Tochigi Prefecture, Japan; in cliff facing Tobu electric R.R., 600 m
NW of Momiyama Station
Miocene, Kanuma Fm
ozawai, Elphidium: Uchio Paratype
Uchio, 1951b, p. 372, pl. 5, figs. 11a, 11b
Tochigi Prefecture, Japan; in bluish grey sandstone of “Terayama
group” at Hachimanyama, Ozo, Utsunomiya City
Miocene, Kanuma Fm
ozawai, Pseudodoliolina: Yabe and Hanzawa Paratype
Yabe and Hanzawa, 1932, p. 40-43
Gifu Prefecture, Japan; Akasaka, Nino Province
pacifica, Cyclammina: Beck Holotype
Beck, 1943, p. 591, pl. 98, figs. 2, 3
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
pacificus, Ammodiscus: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 7
Channel Islands, off southern California
packardi, Yabeina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1942, p. 710, pl. 106, fig. 4 (type 7333a); pl.
108, fig. 4 (type 7333e)
Jefferson Co., Ore.; near base of Gray Butte, near Madras
Permian
packardi, Yabeina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1942, p. 710, pl. 107, figs. 2 (type 7333b), 3
(type 7333c), 4 (type 7333d)
Jefferson Co., Ore.; near Madras, near base of Gray Butte
Permian
panzana, Cassidulina: Kleinpell Holotype
Kleinpell, 1938, p. 335, pl. 8, figs. 9a, 9b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
pariana, Anomalina: Hedberg Paratypes
Hedberg, 1937, p. 681
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
pariana, Nodosaria: Hedberg Paratypes
Hedberg, 1937, p. 672
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
6870
6132
cell 31
859
6132
cell 38
7329a
7329b
7329¢c
7329d
9459
7424
671
9461
7910
7714
7715
7716
Teele
7718
6132
cell 5
STANFORD UNIVERSITY TyPEs: SMITH 497
parianus, Ammodiscus: Hedberg Paratype
Hedberg, 1937, p. 666
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
parva, Gyroidina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 23
Falcon, Venezuela; Pozon, 18.5 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
parva, Uvigerinella californica: Kleinpell Holotype
Kleinpell, 1938, p. 289, pl. 9, fig. 14
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
paucicostata, Pseudoglandulina gallowayi: Cushman and Renz
Paratype
Cushman and Renz, 1941, p. 16
Falcon, Venezuela; Isidro, 34.5 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone III)
pavilionensis, Schwagerina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1942, p. 706, pl. 105, figs. 3, 4, 5, 6
British Columbia, Canada; Marble Canyon, near Lillooet
Permian, Marble Canyon Ls
paynei, Bolivinoides: Martin Holotype
Martin, Lewis, 1964, p. 90, pl. 12, figs. la-1c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
paynei, Quinqueloculina: Beck Holotype
Beck, 1943, p. 593, pl. 98, figs. 6, 7, 8
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
perrini, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 278, pl. 7, figs. 4a, 4b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
pingue, Elphidium fax: Nicol Holotype
Nicol, 1944, p. 177, pl. 29, figs. 1, 2
Monterey Co., Calif.; Mussel Point, Monterey Bay
planum, Elphidium: Husezima and Maruhasi Paratype
Husezima and Maruhasi, 1944, p. 392
Niigata-ken, Japan; Kashiwazaki Oil Field, well No. 1, depth 60 m
Pliocene, upper Haizume Fm
plena, Schwagerina aculeata: Thompson and Hazzard Syntypes
ieee and Hazzard, 1946, p. 46, pl. 13, figs. 1 (type 7714), 2
(7715
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
plena, Schwagerina aculeata: Thompson and Hazzard Syntypes
Thompson and Hazzard, 1946, p. 46, pl. 13, figs. 3 (type 7716), 4, 6
(type 7717), 5 (type 7718)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
pozoensis, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 16
Falcon, Venezuela; Pozon, 18.35 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
498
6132
cell 33
6133
cell 3
6133
cell 8
9398
8323
5938
7698
7694
7695
7696
7697
7699
7700
849
BuLuetin 300
pozoensis, Liebusella: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 9
Falcon, Venezula; Pozon, 17.7 km SE of Pueblo Jacura, District
Zamura
Lower Miocene, Agua Salada Fm (Zone II)
pozoensis, Siphonina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 24
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
pozoensis, Textularia: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 4
Falcon, Venezuela; Pozon, 21.1 km SE of Pueblo Jacura, District
Acosta
Middle Miocene, Agua Salada Fm (Zone VI)
praeconvergens, Lenticulina: Martin Holotype
Martin, Lewis, 1964, p. 66, pl. 6, figs. 3a, 3b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
primitiva, Globotruncana (Praeglobotruncana) renzi: Kupper
Holotype
Kupper, 1956, p. 43, pl. 8, fig. 2
Colusa Co., Calif.; Lodoga Qd, 325’ S, 500’ W of NE cor. Sec. 8, T
17N,R 4W
Upper Cretaceous
primus, Eponides: Martin Holotype
Martin, Lois, 1943, p. 23, pl. 9, figs. 4a-4c
Fresno Co., Calif.; Panoche Qd, Lodo Gulch [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
proteus, Manorella: Grice Paratypes
Grice, 1948, pp. 222-224, figs. 1, 3, 4, 5
Travis Co., Texas; Austin-Manor Highway, bridge over Little Wal-
nut Creek
Cretaceous, Austin Chalk
providens, Schwagerina: Thompson and Hazzard Holotype
Thompson and Hazzard, 1946, p. 43, pl. 14, fig. 1
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
providens, Schwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 43, pl. 14, figs. 4 (type 7695), 5 (type
7696), 6 (type 7694)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
providens, Schwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 43, pl. 14, figs. 2, 7, 8 (type 7697),
3 (type 7699), 9 (type 7700)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
pseudoaffinis, Bulimina: Kleinpell Holotype
Kleinpell, 1938, p. 257, pl. 9, fig. 9
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
9407
6096
5060
6561
7913
97799
5627
6097
7383
9399
660
931
5799
STANFORD UNIVERSITY TyPEs: SMITH 499
pseudoligostegius, Robulus: Martin Holotype
Martin, Lewis, 1964, p. 69, pl. 6, figs. 12a-12c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
pseudospissa, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 279, pl. 21, fig. 6
Los Angeles Co., Calif.; Mohn Springs
Miocene, Modelo Shale
psila, Discocyclina: Woodring Neotype
Woodring, 1930, p. 148, pl. 3, fig. 4
Santa Barbara Co., Calif.; Lompoc Qd, about 5 miles NW of Point
Conception lighthouse SU loc. 356
Upper Eocene
psila, Discocyclina: Woodring Paratype
Woodring, 1930, p. 148
Santa Barbara Co., Calif.; Lompoc Qd, Canada de los Sauces SU
loc. 167
Upper Eocene
pulchella, Epistominella: Husezima and Maruhasi Paratype
Husezima and Maruhasi, 1944, p. 398
Niigata-ken, Japan; Kashiwazaki Oil Field, well No. 1, depth 221.5-
222 m
Pliocene, lower Haizume Fm
putahensis, Giobotruncana: Takayanagi Holotype
Takayanagi, 1965, p. 221, pl. 27, figs. 2a-2c
Yolo Co., Calif.; Putah Creek, hole No. 5A, Sec. A, 70’
Upper Cretaceous, Forbes Fm
quadrata, Cassidulina subglobosa: Cushman and Hughes
Paratype
Cushman and Hughes, 1925, p. 15
Los Angeles Co., Calif.; Palos Verdes Hills, Lomita Quarry SU loc.
1125
Pleistocene, San Pedro Fm
rankini, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 290, pl. 22, figs. 4, 9
Los Angeles Co., Calif.; near Girard
Miocene, Modelo Diatomite
rectiangula, Gaudryina (Siphogaudryina): Martin Holotype
Martin, Lois, 1943, p. 14, pl. 5, figs. 4a, 4b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
rectovalis, Lenticulina; Martin Holotype
Martin, Lewis, 1964, p. 66, pl. 6, figs. 4a, 4b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
reedi, Robulus: Kleinpell Holotype
Kleinpell, 1938, p. 201, pl. 7, figs. 23a, 23b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
reedi, Robulus: Kleinpell Paratype
Kleinpell, 1938, p. 201, pl. 8, figs. 5
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
reedi, Siphogenerina: Cushman Paratype
Cushman, 1925a, p. 3
San Luis Obispo Co., Calif.; Sec. 24,T 28S,R14E
Miocene, Monterey Shale
500
6132
cell 2
840
959
5016
942
7404
7387
762
7381
873
7628
7629
7630
BULLETIN 300
regularis, Bolivina floridana: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 17
Falcon, Venezuela; Isidro, 33.5 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
relizensis, Cibicides: Kleinpell Holotype
Kleinpell, 1938, p. 355, pl. VII, figs. 15a-15c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
relizensis, Lenticulina: Kleinpell Holotype
Kleinpell, 1938, p. 205, pl. 10, figs. 6a, 6b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
relizensis, Pulvinulinella: Kleinpell Holotype
Kleinpell, 1938, p. 329, pl. X, figs. 10a-10c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale [specimen missing, ca. 1940-1941]
reliziana, Gyroidina: Kleinpell Holotype
Kleinpell, 1938, p. 315, pl. X, figs. lla, 11b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
rex, Globorotalia: Martin Holotype
Martin, Lois, 1943, p. 27, pl. 8, figs. 2a-2c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
rex, Vaginulinopsis saundersi: Martin Holotype
Martin, Lois, 1943, p. 17, pl. 9, figs. 2a, 2b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
rhomboidea, Bolivina: Cushman Paratype
Cushman, 1926b, p. 19
San Luis Potosi, Mexico; Tamuin River, 5 km SE of Guerrero
Cretaceous, Mendez Shale
richardi, Spiroplectammina: Martin Holotype
Martin, Lois, 1943, p. 14, pl. 5, figs. 3a, 3b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
robusta, Baggina: Kleinpell Holotype
Kleinpell, 1938, p. 325, pl. XI, figs. 8a-8c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
robusta, Fusulina: Meek Neotype
Meek, 1864b, pp. 3-4, pl. 2, figs. 3a-3c. Neotype designated by Thomp-
son and Wheeler, 1946, pp. 28-29, pl. 3, fig. 1 [as Pseudoschwagerina
robusta (Meek) |
Shasta Co., Calif.; Redding Qd, W side of Is hogback, NW 1/4, NE
1/4 Sec. 15, T 33 N, R 4 W, elev. 1400’ SU loc. 774
Permian, McCloud Ls
robusta, Fusulina: Meek Paraneotypes
Meek, 1864b, pp. 3-4. Paraneotypes illustrated by Thompson and
Wheeler, 1946, pp. 28-29, pl. 6, figs. 7 (type 7629), 6 (type 7630)
Shasta Co., Calif.; Redding Qd, W side of Is hogback, NW 1/4, NE
1/4 Sec. 15, T 33 N, R 4 W, elev. 1400’ SU loc. 774
Permian, McCloud Ls
7631
7632
585
7726
7723
7724
7729
9479
7944
7945
6132
cell 10
6133
cell 4
908
918
854
STANFORD UNIVERSITY TyPEs: SMITH 501
robusta, Fusulina: Meek Paraneotypes
Meek, 1864b, pp. 3-4. Paraneotypes illustrated by Thompson and
Wheeler, 1946, pp. 28-29, pl. 3, figs. 2 (type 7631), 3 (type 7632)
Shasta Co., Calif.; Redding Qd, W side of Is knoll in SE 1/4, SE 1/4
Sec. 15, T 33 N, R 4 W, elev. 1100’ SU loc. 775
Permian, McCloud Ls
robustior, Massilina: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 8
Channel Islands, off southern California
roeseleri, Pseudoschwagerina: Thompson and Hazzard Holotype
Thompson and Hazzard, 1946, p. 47, pl. 17, fig. 4
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
roeseleri, Pseudoschwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 47, pl. 17, figs. 5 (type 7725), 6
(type 7723); pl. 18, fig. 4 (type 7724)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
rosaceus, Globorotalites: Martin Holotype
Martin, Lewis, 1964, p. 99, pl. 14, figs. 5a-5c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
rota, Haplophragmoides: Nauss Holotype
Nauss, 1947, p. 339, pl. 49, figs. la, 1b
Alberta, Canada; Imperial core test No. 14, Legal subdivision 5, Sec.
7, T 51, R 8 W, 4th meridian, depth 55-60’, 10-15’ above base of
member
Upper Cretaceous, Belly River Fm, Grizzly Bear tongue
rota, Haplophragmoides: Nauss Paratype
Nauss, 1947, p. 339, pl. 49, figs. 3a, 3b
Alberta, Canada; Imperial core test No. 2, depth 190-200’, Vermilion
area
Upper Cretaceous, Belly River Fm, Grizzly Bear tongue
rudderi, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 19
Falcon, Venezuela; Pozon, 18.6 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
rutschi, Sphaeroidinella: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 25
Falcon, Venezuela; Isidro, 33.5 km E of Pueblo Piritu, District
Zamura
Miocene, Agua Salada Fm (Zone IV)
salinasensis, Anomalina: Kleinpell Holotype
Kleinpell, 1938, p. 347, pl. XIII, figs. 1a-1c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
salinasensis, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 280, pl. 15, fig. 3
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
salinasensis, Bolivina: Kleinpell Paratype
Kleinpell, 1938, p. 280, pl. 9, fig. 6
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
502
6090
7835
7836
7837
7838
7839
7840
7841
7842
7843
7844
7492
6659
9401
962
926
5025
583
6091
BuLLeTIn 300
sanctaecrucis, Nodogenerina: Kleinpell Holotype
Kleinpell, 1938, p. 246, pl. 4, fig. 22
Santa Cruz Co., Calif.; Santa Cruz Mts. SU loc. 1162
Oligocene, Vaqueros Fm
schencki, Afghanella: Thompson Holotype
Thompson, 1946, p. 153, pl. 25, fig. 2
Afghanistan; Shibar Pass, ca. 7 km W of summit on Kabul to Bamian
road
Permian, Bamian Ls
schencki, Afghanella: Thompson Paratypes
Thompson, 1946, p. 153, pl. 25, figs. 1 (type 7836), 4 (type 7838), 5
(type 7837), 7 (type 7839), 8 (type 7840)
Afghanistan; Shibar Pass, ca. 7 km W of Summit on Kabul to Bamian
road
Permian, Bamian Ls
schencki, Afghanella: Thompson Paratypes
Thompson, 1946, p. 153, pl. 25, figs. 9 (type 7841), 10 (type 7842),
11 (type 7843), 12 (type 7844) .
Afghanistan; Shibar Pass, ca. 7 km W of Summit on Kabul to Bamian
road
Permian, Bamian Ls
schencki, Bulimina: Beck Holotype
Beck, 1943, p. 605, pl. 107, figs. 28, 33
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
Bd/25SEr/4)Seer 285 ay LN eRe aw,
Eocene, Cowlitz Fm
schencki, Elphidium: Cushman and Dusenbury Holotype
Cushman and Dusenbury, 1934, p. 60, pl. 8, figs. 8a, 8b
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles S$
38° W of BM 394 SU loc. 1150
Eocene, Poway Conglomerate
schencki, Lenticulina: Martin Holotype
Martin, Lewis, 1964, p. 67, pl. 6, figs. 6a, 6b
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
schencki, Nonion: Kleinpell Holotype
Kleinpell, 1938, p. 235
Monterey Co., Calif.; diatomite quarry SU loc. 662
Miocene, Monterey Shale
schencki, Nonion: Kleinpell Paratype
Kleinpell, 1938, p. 235, pl. XVI, figs. 11a, 11b
Monterey Co., Calif.; Reliz Canyon SU loc. 691, sample Fi, 3110’
above Vaqueros Ss
Miocene, Monterey Shale
schencki, Saracenaria: Cushman and Hobson Paratype
Cushman and Hobson, 1935, p. 57
Santa Cruz Co., Calif.; Santa Cruz Qd, Bear Creek SU loc. 1102
Oligocene, San Lorenzo Fm
schencki, Textularia: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 8
Channel Islands, off southern California
semihispida, Buliminella: Kleinpell Holotype
Kleinpell, 1938, p. 250, pl. 20, figs. 8, 15, 16
Santa Barbara Co., Calif.; near Naples
Miocene, Monterey Shale
6132
cell 22
6880
6133
cell 2
6673
6674
6067
6132
cell 4
7482
910
6658
6109
6110
6134
cell 34
STANFORD UNIVERSITY TyPEs: SMITH 503
senni, Robulus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 12
Falcon, Venezuela; Pozon, 18.9 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
senni, Saracenaria: Hedberg Paratype
Hedberg, 1937, p. 674
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
senni, Siphogenerina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 22
Falcon, Venezuela; Isidro, 34.9 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone III)
sesquistriata, Lagena: Bagg Holotype
Bagg, 1912, p. 50, pl. 13, figs. 13, 14b
San Pedro, Calif.; Timms Point SU loc. 2024
“Pliocene”
sesquistriata, Lagena: Bagg Paratype
Bagg, 1912, p. 50, pl. 13, figs. 12, 14a
San Pedro, Calif.; Timms Point SU loc. 2024
“Pliocene”
shivelyi, Textularia: Kleinpell Holotype
Kleinpell, 1938, p. 190, pl. 1, figs. 5, 9
Santa Barbara Co., Calif.; near Gaviota Pass SU loc. 1436
Oligocene, “Sespe” Fm
simplex, Bolivina interjuncta: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 20
Falcon, Venezuela; Pozon, 21.1 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
sinuata, Globuiina minuta: Beck Holotype
Beck, 1943, p. 603, pl. 106, fig. 13
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W _ SU loc. M-335
Eocene, Cowlitz Fm
smiieyi, Robulus: Kleinpell Holotype
Kleinpell, 1938, p. 202, pl. XV, figs. 14a, 14b
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
smithi, Elphidium: Cushman and Dusenbury Holotype
Cushman and Dusenbury, 1934, p. 61, pl. 8, figs. 7a, 7b
San Diego Co., Calif.; La Jolla Qd, Murray Canyon, 1 1/8 miles S
38° W of BM 394 SU loc. 1150
Eocene, Poway Conglomerate
smithi, Siphogenerina: Kleinpell Holotype
Kleinpell, 1938, p. 304, pl. VI, fig. 1
Santa Barbara Co., Calif.; Elwood Field, Doty well No. 4
Oligocene, Rincon Shale
smithi, Siphogenerina: Kleinpell Paratype
Kleinpell, 1938, p. 304, pl. VI, fig. 2
Santa Barbara Co., Calif.; Elwood Field, Doty well No. 4
Oligocene, Temblor Shale
soldadoensis, Discorbis midwayensis: Cushman and Renz
Paratype
Cushman and Renz, 1942, p. 10
Trinidad, B.W.I.; Soldado Rock
Paleocene, Soldado Fm
504
6134
cell 38
7929
7948
588
6952
7946
5941
6132
cell 36
6113
755
686
684
BuLLETIN 300
soldadoensis, Nonionella: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 7
Trinidad, B.W.1; Soldado Rock
Paleocene, Soldado Fm
solis, Anomalina: Nauss Holotype
Nauss, 1947, p. 333, pl. 49, figs. 9a-9c
Alberta, Canada; Imperial Core test No. 82, Legal subdivision 16,
Sec. 24, T 56, R 7 W, 4th meridian, depth 420-425’
Upper Cretaceous, Lea Park Shale
sphaera, Quinqueloculina: Nauss Holotype
Nauss, 1947, p. 340, pl. 48, figs. 14a-14c
Alberta, Canada; Vermilata Frankview Well No. 1, Legal subdivision
16, Sec. 28, T 50, R 5 W, 4th meridian, 630-640’, 220’ above base of
fm
Upper Cretaceous, Lea Park Shale
spinatum, Elphidium: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 21
Channel Islands off southern California
spiveyi, Waeringella: Thompson Syntype
Thompson, 1942, p. 414
SE of Graham, Texas; Herron Bend, Brazos River
Pennsylvanian, Salem School Ls
sproulei, Miliammina: Nauss Holotype
Nauss, 1947, p. 339, pl. 48, figs. 13a, 13b
Alberta, Canada; NW Mannville well No. 1, Legal subdivision 1, Sec.
18, T 50, R 8 W, 4th meridian, depth 2152-2162’. Vermillion area
Lower Cretaceous, Mannville Fm, Cummings Mbr
sproulei, Miliammina: Nauss Paratype
Nauss, 1947, p. 339
Alberta, Canada; NW Mannville well No. 1, Legal subdivision 1, Sec.
18, T 50, R 8 W, 4th meridian, depth 2152-2162’. Vermillion area
Lower Cretaceous, Mannville Fm, Cummings Mbr
stainforthi, Nodosaria: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 15
Falcon, Venezuela; Isidro, 33.75 km E of Pueblo Piritu, District
Zamura
Miocene, Agua Salada Fm (Zone IV)
striatella, Bolivina advena: Cushman Paratype
Cushman, 1925c, p. 30
San Luis Obispo Co., Calif.; Sec. 24, T 28S, R14 E
Miocene, Monterey Shale
subcasitasensis, Valvulineria casitasensis: Kleinpell Holotype
Kleinpell, 1938, p. 311, pl. 2, figs. 3, 4, 14
Santa Barbara Co., Calif.; near Gaviota Pass SU loc. 1436
Oligocene, “Sespe” Fm
subfusiformis, Buliminella: Cushman Paratype
Cushman, 1925c, p. 33
San Luis Obispo Co., Calif.; Sec. 24, T 28 §,R14E
Miocene, Monterey Shale
subplana, Virgulina: Barbat and Johnson Holotype
Barbat and Johnson, 1934, p. 14, pl. 1, fig. 17
Kings Co., Calif.; Associated Oil Co., Whepley No. 1, Kettleman
Hills; Coalinga Qd, Sec. 35, T 21S,R17E SU loc. 697
Miocene, lower Reef Ridge Shale
subplana, Virgulina: Barbat and Johnson Paratype
Barbat and Johnson, 1934, p. 14, pl. 1, fig. 16
Kings Co., Calif.; Coalinga Qd, Sec. 35, T 21 S, R 17 E, Associated Oil
Co., Whepley No. 1, Kettleman Hills SU loc. 697
Upper Miocene, lower Reef Ridge Shale
6132
cell 23
6132
cell 9
6132
cell 18
7930
7930a
7746
6132
cell 30
6098
7744
7753
5631
5628
STANFORD UNIVERSITY Types: SMITH 505
superba, Marginulina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 14
Falcon, Venezuela; Pozon, 18.4 km SE of Pueblo Jacura, District
Acosta
Miocene, Agua Salada Fm (Zone IV)
suteri, Bolivina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 18
Falcon, Venezuela; Tocuyo, 15.7 km E of San Juan de los Cayos,
District Acosta
Miocene, Agua Salada Fm (Zone IV)
suteri, Robulus: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 10
Falcon, Venezuela; Isidro, 33.2 km E of Pueblo Piritu, District Zamura
Miocene, Agua Salada Fm (Zone IV)
talaria, Anomalina: Nauss Holotype
Nauss, 1947, p. 334, pl. 48, figs. lla-11c
Alberta, Canada; Imperial Core Test No. 27, Legal subdivision 1,
Sec. 4, T 54, R 8 W, 4th meridian, depth 260-270’
Upper Cretaceous, Lea Park Shale
talaria, Anomalina: Nauss Paratype
Nauss, 1947, p. 334, pl. 48, figs. 12a-12c
Alberta, Canada; Imperial Core Test No. 27, Legal subdivision 1,
Sec. 4, T 54, R 8 W, 4th meridian, depth 260-270’
Upper Cretaceous, Lea Park Shale
tanaii, Eponides: Uchio Paratype
Uchio, 1951b, p. 376, figs. 8a-8c, 9a-Yc
Tochigi Prefecture, Japan; in cliff facing Tobu electric R.R., 600
m NW of Momiyama Station
Miocene, Kanuma Fm
thalmanni, Gaudryina: Cushman and Renz Paratype
Cushman and Renz, 1941, p. 7
Falcon, Venezuela; Pozon, 27 km SE of Pueblo Jacura, District
Acosta
Lower Miocene, Agua Salada Fm (Zone II)
ticensis, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 284, pl. 18, figs. 6, 7
Contra Costa Co., Calif.; San Pablo Creek
Miocene, Tice Shale
tochigiensis, Rotalia: Uchio Paratype
Uchio, 1951b, p. 374, pl. 5, figs. 1a-1c
Tochigi Prefecture, Japan; in cliff facing Tobu electric R.R., 600
m NW of Momiyama Station
Miocene, Kanuma Fm
tomiyensis, Cassidulina: Uchio Paratype
Uchio, 195la, p. 40
Chiba-ken, Japan; Tomiya, Takeoka-mura, Kimitsu-gun
Pliocene, Tomiya Fm
tortuesa, Cassidulina: Cushman and Hughes Paratype
Cushman and Hughes, 1925, p. 14
San Pedro, Calif.; Timms Point SU loc. 2024
“Pliocene,” Timms Point Fm
translucens, Cassidulina: Cushman and Hughes Paratype
Cushman and Hughes, 1925, p. 15
Los Angeles Co., Calif.; Palos Verdes Hills, Lomita Quarry SU loc.
1125
Pleistocene, San Pedro Fm
6691
6134
cell 35
6134
cell 37
9477a
9477b
589
7638
7639
7640
7641
7642
7643
7644
7645
7928
5940
7730
7727
7728
7729
7731
BuLLeTiIn 300
trilocularia, Polymorphina: Bagg Holotype
Bagg, 1912, p. 75, pl. 20, figs. 15, 17 (?)
San Pedro, Calif.; Timms Point SU loc. 2024
“Pliocene”
trinitatensis, Discorbis midwayensis: Cushman and Renz
Paratype
Cushman and Renz, 1942, p. 10
Trinidad, B.W.I.; Soldado Rock
Paleocene, Soldado Fm
trinitatensis, Gumbelina: Cushman and Renz Paratype
Cushman and Renz, 1942, p. 8
Trinidad, B.W.I.; Soldado Rock
Paleocene, Soldado Fm
turbinata, Gavelinella: Martin Holotype
Martin, Lewis, 1964, p. 99, pl. 14, figs. 2a-2c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Moreno Fm
turbinata, Gavelinella: Martin Paratype
Martin, Lewis, 1964, p. 99, pl. 14, figs. 3a-3c
Fresno Co., Calif.; Panoche Hills, Sec. 6, T 15 S, R12 E
Upper Cretaceous, Moreno Fm
turbinata, Rotalia: Cushman and Valentine Paratype
Cushman and Valentine, 1930, p. 25
Channel Islands off southern California
turgida, Parafusulina?: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 30, pl. 4, fig. 1, pl. 5, fig. 6 (type
7639); pl. 4, fig. 2 (type 7641); pl. 5, figs. 1 (type 7640), 2 (type
7638)
Shasta Co., Calif.; Redding Qd, crest of Is ridge S of Potter Creek,
NE 1/4, SE 1/4 Sec. 23, T 34 N, R 4 W, elev. 1675’ SU loc. 757
Permian, McCloud Fm, middle part
turgida, Parafusulina?: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 30, pl. 5, figs. 3 (type 7642), 4 (type
7643), 5 (type 7644) ; pl. 4, fig. 3 (type 7645)
Shasta Co., Calif.; Redding Qd, crest of Is ridge S of Potter Creek,
NE 1/4, SE 1/4 Sec. 23, T 34 N, R 4 W, elev. 1675’ SU loc. 757
Permian, McCloud I'm, middle part
tyrrelli, Ammobaculites: Nauss Holotype
Nauss, 1947, pp. 333-334, pl. 48, fig. 2
Alberta, Canada; Dina Omega Well No. 1, Legal subdivision 14, Sec.
9, T 45, R 1 W, 4th meridian, depth 1478-1488’ Vermillion area
Upper Cretaceous, Lloydminster Shale
tyrrelli, Ammobaculites: Nauss Paratype
Nauss, 1947, pp. 333-334
Alberta, Canada; Dina Omega Well No. 1, Legal subdivision 14, Sec.
9, T 45, R 1 W, 4th meridian, depth 1478-1488’ Vermillion area
Upper Cretaceous, Lloydminster Shale
uber, Pseudoschwagerina: Thompson and Hazzard Holotype
Thompson and Hazzard, 1946, p. 48, pl. 17, fig. 1
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
uber, Pseudoschwagerina: Thompson and Hazzard Paratypes
Thompson and Hazzard, 1946, p. 48, pl. 14, figs. 10 (type 7731), 11
(type 7729) ; pl. 17, figs. 2 (type 7727), 3 (type 7728)
San Bernardino Co., Calif.; E front, Providence Mts., S of Gilroy
Mine, 1.5 miles N of end of road to Mitchell’s Caverns
Permian, Bird Spring Fm
9362
7469
9496
6888
6875
6885
7933
7386
7651
7652
7653
7654
7655
7656
7657
7658
7659
7660
STANFORD UNIVERSITY TyPEs: SMITH 507
uvigerinaeformis, Bermudezina: Martin Holotype
Martin, Lewis, 1964, p. 53, pl. 3, figs. 6a-6c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch
Upper Cretaceous, Moreno Fm
[ Missing; no record that specimen was received at SU]
vaderensis, Frondicularia: Beck Holotype
Beck, 1943, p. 601, pl. 107, fig. 18
Lewis Co., Wash.; 1.5 miles E of Vader, W bank of Cowlitz River,
E 172, SE 1/4 Sec...28, T 11.N, KR 2 W SU loc. M-335
Eocene, Cowlitz Fm
validus, Cibicidoides: Martin Holotype
Martin, Lewis, 1964, p. 107, pl. 16, figs. 5a-5c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
venezuelana, Giobigerina: Hedberg Paratypes
Hedberg, 1937, p. 681
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
venezuelana, Planularia: Hedberg Paratype
Hedberg, 1937, p. 670
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
venezuelana, Rzehakina: Hedberg Paratype
Hedberg, 1937, p. 669
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
venezuelana, Valvulineria: Hedberg Paratypes
Hedberg, 1937, p. 678
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
venusae, Bulimina: Nauss Holotype
Nauss, 1947, p. 334, pl. 48, fig. 10
Alberta, Canada; Imperial core test no. 7, Legal subdivision 11, Sec.
8, T 51, R 7 W, 4th meridian, depth 125’, 15’ above base of mbr
Upper Cretaceous, Belly River Fm, Vanesti tongue
verruculosa, Vaginulinopsis, Martin Holotype
Martin, Lois, 1943, p. 16, pl. 5, figs. 6a, 6b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
virga, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 32, pl. 6, figs. 1 (type 7652), 2 (type
7653), 3 (type 7651)
Shasta Co., Calif.; Redding Qd, SW 1/4, SW 1/4 Sec. 2, T 33 N,
R 4 W, elev. 1600’
Permian, Noscni Fm
virga, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 32, pl. 9, figs. 1 (type 7654), 2 (type
7655), 3 (type 7656), 4 (type 7657)
Shasta Co., Calif.; Redding Qd, SW 1/4, SW 1/4 Sec. 2, T 33 N,
R 4 W, elev. 1600’
Permian, Nosoni Fm
virga, Parafusulina: Thompson and Wheeler Syntypes
Thompson and Wheeler, 1946, p. 32, pl. 9, figs. 5 (type 7658), 6 (type
7659), 7 (type 7660)
Shasta Co., Calif.; Redding Qd, SW 1/4, SW 1/4 Sec. 2, T 33 N,R
4 W, elev. 1600’
Permian, Nosoni Fm
508
7931
6876
7444
7489
7390
7436
9493
7394
7412
7391
7425
6854
BuLLETIN 300
vitta, Bathysiphon: Nauss Holotype
Nauss, 1947, p. 334, pl. 48, fig. 4
Alberta, Canada; Imperial Core Test No. 83, Legal subdivision 4,
Sec. 4, T 56, R 5 W, 4th meridian, depth 265-270’, about 270’ above
base of fm
Upper Cretaceous, Lea Park Shale
wallacei, Marginulina: Hedberg Paratype
Hedberg, 1937, p. 670
Anzoategui, Venezuela; District Libertad
Oligocene, Carapita Fm
washingtonensis, Lenticulina: Beck Holotype
Beck, 1943, p. 597, pl. 104, figs. 18, 21
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
washingtonensis, Robertina: Beck Holotype
Beck, 1943, p. 604, pl. 107, figs. 17, 19
Lewis Co., Wash.; 1.5 miles E of Vader on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T11N,R2W_ SU loc. M-335
Eocene, Cowlitz Fm
watsonae, Lagena: Martin Holotype
Martin, Lois, 1943, p. 18, pl. 15, figs. 7a, 7b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Gd] SU
loc. M-74
Eocene, Lodo Fm
weaveri, Robulus: Beck Holotype
Beck, 1943, p. 595, pl. 103, figs. 3, 8
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N, R2W_ SU loc. M-335
Eocene, Cowlitz Fm
whitei, Anomalina: Martin Holotype
Martin, Lewis, 1964, p. 106, pl. 16, figs. 4a-4c
Fresno Co., Calif.; Panoche Hills, Moreno Gulch, 4 miles SE of Little
Panoche Creek
Upper Cretaceous, Panoche Fm
whitei, Bulimina: Martin Holotype
Martin, Lois, 1943, p. 20, pl. 6, figs. 5a, 5b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
whitei, Cibicides: Martin Holotype
Martin, Lois, 1943, p. 32, pl. 8, figs. 4a-4c
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
whitei, Plectofrondicularia: Martin Holotype
Martin, Lois, 1943, p. 19, pl. 5, figs. 2a, 2b
Fresno Co., Calif.; Lodo Gulch, Panoche Qd [Tumey Hills Qd] SU
loc. M-74
Eocene, Lodo Fm
whitei, Quinqueloculina: Beck Holotype
Beck, 1943, p. 593, pl. 99, figs. 11, 12, 13
Lewis Co., Wash.; 1.5 miles E of Vader, on W bank of Cowlitz River,
E 1/2, SE 1/4 Sec. 28, T 11 N,R2W_ SU loc. M-335
Eocene, Cowlitz Fm
whitei, Siphogenerinoides: Church Paratypes
Church, 1941, p. 182
Fresno Co., Calif.; Panoche Hills, near center of Sec. 6, T 15S,R12E
Upper Cretaceous, Moreno Shale
944
678
9937
6099
6087
7593
8524
8922
7374
6767
5210
5211
STANFORD UNIVERSITY [TyPrEs: SMITH 509
williami, Cassidulina: Kleinpell Holotype
Kleinpell, 1938, p. 337, pl. XIV, fig. 5
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
williami, Valvulineria: Kleinpell Holotype
Kleinpell, 1938, p. 315, pl. VII, figs. 14a-14c
Monterey Co., Calif.; Reliz Canyon SU loc. 691
Miocene, Salinas Shale
yabei, Parafusulina: Hanzawa Paratypes
Hanzawa, 1942, p. 127-128
Ago-gun, Totigi Prefecture, Japan; Tomuro, Miyosi-mura
Permian
yneziana, Bolivina: Kleinpell Holotype
Kleinpell, 1938, p. 286, pl. 2, fig. 8
Santa Barbara Co., Calif.; near Gaviota Pass SU loc. 1436
Oligocene, “Sespe” Fm
ynezianum, Nonion: Kleinpell Holotype
Kleinpell, 1938, p. 237, pl. II, figs. 1, 2
Santa Barbara Co., Calif.; near Gaviota Pass
Oligocene, “Sespe” Fm
COELENTERATA
browni, Astrangia: Palmer Paratype
Palmer, 1928b, p. 27
Oaxaca, Mexico; 4 miles W of Puerto Angel
confluens, Heliophyllum obconicum: Fenton and Fenton
Plastoholotype
Fenton and Fenton, 1938, p. 221, pl. 21, fig. 1
New York, near East Bethany
Devonian, Moscow Fm [holotype 37765 from Walker Museum, now
probably in the Field Museum Nat. Hist., Chicago]
decorosum, Heliophyllum: Fenton and Fenton Plastoholotype
Fenton and Fenton, 1938, p. 216, pl. 18, figs. 7, 8
Leicester, N.Y.; Little Beard Creek
Devonian, Moscow Fm [cast from Carnegie Museum sections 6754,
6755 |
fresnoense, Flabellum: Durham Holotype
Durham, 1943, p. 197, pl. 32, figs. 2, 3
Fresno Co., Calif.; SU loc. M-49, Cheney Well No. 1, 5800’
Upper Cretaceous
hannibali, Coenocyathus?: Durham Holotype
Durham, 1942, p. 93, pl. 17, fig. 14, text fig. 1
Mason Co., Wash.; NP loc. 207, T 21 N, R 5 W;; bluffs on Vance’s
Ck 2.5 miles above jet. with Skokomish River, 13 miles above Union
City
Lower Oligocene
hannibali, Dendrophyllia: Nomland Syntype
Nomland, 1916a, p. 67, pl. 6, figs. 1, 2
Grays Harbor Co., Wash.; NP icc. 51, bluffs at old log dam on
Porter Ck, 1.5 miles above Porter
Oligocene, Linceln Fm
hannibali, Dendrophyllia: Nomland Syntype
Nomland, 1916a, p. 67, pl. 6, fig. 3
Grays Harbor Co., Wash.; NP loc. 51, bluffs at old log dam on
Porter Ck, 1.5 miles above Porter
Oligocene, Lincoln Fm
510
362
9505a
7594
5905
7956
7954
221
8523
6765
6546
9260
9860a
9863
9864
BuLueTIN 300
hyatti, Astrocoenia: Wells Paratype
Wells, 1942, p. 1
Wyoming; 3 miles W of Cody, bank of Shoshone River
Jurassic, Sundance Fm
hypatiae, Multithecopora: Wilson Paratype
Wilson, 1963, p. 158, pl. 21, figs. 3, 4; pl. 22, figs. 1-3, 7
White Pine Co., Nevada; near Lund. SU loc. 3474
Middle Pennsylvanian, Ely Fm
mexicana, Cycloseris: Durham Paratypes
Durham, 1947, p. 24
Gulf of California; Amortajada Bay, in La Paz Bay, Carmen Island
oldroydi, Dendrophyllia: Oldroyd ex Faustino MS Syntype
Oldroyd, I. S., 1925, pl. 49, fig. 7 (part of the type colony). Described
by Faustino, 1931, pp. 286-287, pl. 1
Sunken Valley, Calif.; between San Pedro and Redondo, 200 fms
palmata, Pocillopora: Palmer Syntype
Palmer, 1928), p. 31, pl. 2, fig. 3; pl. 3, fig. 1
Oaxaca, Mexice; Puerto Angel harbor ,
quaylei, Cyathoceras: Durham Paratype
Durham, 1947, p. 32
Monterey Co., Calif.; off Point Sur, 160 fms
radcliffi, Sidastrea: Faustino Holotype
Faustino, 1931, p. 285, pl. 1, fig. 1
Ventura Co., Calif.; Camulos Qd, near Simi Peak
Lower Eocene, Martinez Fm_ [Paleocene]
teres, Heliophyllum obconicum: Fenton and Fenton
Plastoholotype
Fenton and Fenton, 1938, p. 222, pl. 19, fig. 6
Western N.Y., near Le Roy
Devonian, Moscow Fm [cast of 2 sections, Carnegie Museum Nos.
6868, 6864]
townsendensis, Trochocyathus: Durham Holotype
Durham, 1942, p. 90, pl. 15, fig. 6
Jefferson Co., Wash.; NP loc. 148, sea cliffs .25 miles N of old Wood-
man Wharf, Port Discovery; NE 1/4 Sec. 8, T 29 N, R1 W
Lower Oligocene, Quimper
whitei, Deltocyathus: Durham Holotype
Durham, 1943, p. 200, pl. 32, figs. 13, 16, text fig. 1
Fresno Co., Calif.; SU loc. 2073, Tumey Hills Qd, ject of Silver and
Panoche Creeks
Paleocene, Lodo Fm
PORIFERA
astoma, Polytholosia: Seilacher Holotype
Seilacher, 1962, p. 758, pl. 3, figs. 1, 2
Cedar Mts., Nevada
Triassic, Luning Fm., Karnic
astoma, Polytholosia: Seilacher Paratype
Seilacher, 1962, p. 758, pl. 3, figs. 3, 4, 5
Cedar Mts., Nevada
Triassic, Luning Fm., Karnic
cylindrica, Polytholosia cylindrica: Seilacher Holotype
Seilacher, 1962, p. 758, 764, pl. 5, fig. 1
Mineral Co., Nevada; Dunlap Canyon, Pilot Mts.
Triassic, lower Luning Fm, Karnic
cylindrica, Polytholosia cylindrica: Seilacher Paratypes
Seilacher, 1962, p. 758, 764, pl. 5, figs. 2-5; pl. 6, fig. 1
Mineral Co., Nevada; Dunlap Canyon, Pilot Mts.
Triassic, lower Luning Fm, Karnic
6756
6755
9865
9865a
6754
9861
9862
7795
8329
5170
5171
9411
STANFORD UNIVERSITY TyPEs: SMITH 511
ellipticus, Receptaculites: Walcott Plastoholotype
Walcott, 1884, p. 67, pl. 11, fig. 12
Eureka district, Nevada; Goodwin Canyon, White Mt.
Ordovician, Pogonip Fm_ [holotype USNM 24548]
elongatus, Receptaculites: Walcott Plastoholotype
Walcott, 1884, p. 66
White Pine district, Nevada; Treasure City
Ordovician, Pogonip Fm [holotype USNM 24635]
expansum, Ascosymplegma: Seilacher Holotype
Seilacher, 1962, p. 759, 768, pl. 8, figs. 2, 3, 4
Mineral Co., Nevada; Cinnabar Canyon, near Mina
Triassic, Luning, Karnic
expansum, Ascosymplegma: Seilacher Paratype
Seilacher, 1962, p. 759, 768, pl. 8, fig. 1
Mineral Co., Nevada; Cinnabar Canyon, near Mina
Triassic, Luning, Karnic
mammillaris, Receptaculites: Walcott Plastoholotype
Walcott, 1884, p. 65, pl. 11, fig. 11
Eureka district, Nevada; Goodwin Canyon, White Mt.
Ordovician, upper Pogonip Fm_ [holotype USNM 24636]
polystoma, Polytholosia: Seilacher Holotype
Seilacher, 1962, p. 758, 762, pl. 4, fig. 1
Augusta Mt., Nevada
Triassic, Karnic, Winnemucca Fm
polystoma, Polytholosia: Seilacher Paratypes
Seilacher, 1962, p. 758, 762, pl. 4, figs. 2, 4, 5
Augusta Mt., Nevada
Triassic, Karnic, Winnemucca Fm
ECHINODERMATA
alaskense, Echinarachnius: Durham Plastoholotype
Durham, 1957, p. 628, pl. 72, figs. 6, 8
Lituya Bay, Alaska; SE shore Cenotaph Island
Pliocene [holotype USNM 562073 fide Durham]
arnoldi, Actinocrinus: Wachsmuth and Springer Holotype
Wachsmuth and Springer, 1890, p. 168, pl. 17, fig. 10
Marshall Co., Iowa; Le Grand
Lower Carboniferous, Kinderhook group [crinoid ]
bahiaensis, Orthopsis: Machado-Brito Paratype
Machado-Brito, 1964, p. 6, pl. 2, fig. 1
Bahia, Brazil; Boipeba Island, Camamu area
Cretaceous, Algodones Fm
blancoensis, Scutella: Kew Syntype
Kew, 1920, p. 64, pl. 11, figs. 1b, 1c
Cape Blanco, Ore.; SU loc. NP 26
Oligocene, “San Lorenzo” Fm
blancoensis, Scutella: Kew Syntype
Kew, 1920, p. 64, pl. 11, fig. 1a
Cape Blanco, Ore.; SU loc. NP 26
Oligocene, “San Lorenzo” Fm
branneri, Cidaris: Arnold Holotype
Arnold, 1908a, p. 363, pl. 33, fig. 5
Santa Cruz Co., Calif.; Bear Creek, 4 miles above San Lorenzo River
Oligocene, San Lorenzo Fm_ [Arnold’s No. 1056]
51Z
7370
579
389
5393
5388
5389
5390
5391
5392
5164
7809
7952
7762
10271
5373
57
Bu.tetin 300
inezana, Encope grandis: Durham Paratype
Durham, 1950, p. 45
Baja California, Mexico; Santa Jnez Point, 10 miles N of Mulege SU
loc. 805
Pleistocene
lovenioides, Megapetalus: Clark Holotype
Clark, 1929, p. 260, pl. 31, figs. 1-6
Ventura Co., Calif.; Santa Paula Qd, E of Coche Canyon on divide
between Coche and Sulphur Canyons, 75 yds W of Crest and on top of
lateral ridge SU loc. 667
Upper Miocene, Santa Margarita Fm?
lymani, Amphiura: Waring Holotype
Waring, 1917, p. 58, pl. 9, fig. 13
Ventura Co., Calif.; Chice area, Bell’s Canyon, N of Simi fault
Upper Cretaceous, “Chico” Fm
merriami, Cidaris: Arnold Plastoholotype
Arnold, 1908a, p. 359, pl. 32, fig. 8
San Mateo Co., Calif.; between headwaters of San Lorenzo River and
Pescadero Creek, SE 1/4 Sec. 23, T 8 S, R 3 W, just W of Santa Cruz
Co. line
“Eocene” [Oligocene or Miocene, fide Keen and Bentson, 1944, p. 231]
[holotype USNM 165438]
merriami, Cidaris: Arnold Paratypes
Arnold, 1908a, p. 359
San Mateo Co., Calif.; between headwaters of San Lorenzo River and
Pescadero Creek, SE 1/4 Sec. 23, T 8S,R 3 W
“Eocene” [Oligocene or Miocene, fide Keen and Bentson, 1944, p. 231]
newcombei, Scutella: Kew Holotype
Kew, 1920, p. 73, pl. 8, figs. 2a, 2b
Vancouver Island, British Columbia, Canada; Jordan River, 1/2 mile
E of Slide Hill Telegraph Station SU loc. NP 131
Oligocene, Sooke Fm
newcombei, Scutella: Kew Paratype
Kew, 1920, p. 73
Vancouver Island, British Columbia, Canada; Jordan River, 1/2 mile
E of Slide Hill Telegraph Station SU loc. NP 131
Oligocene, Sooke Fm
nipponicus, Astrodapsis: Nisiyama Paratypes
Nisiyama, 1948, p. 602
Iwate-ken, Japan; Ninohe-gun, 150 m E of bridge of Kita-Fukuoka
Mio-Pliocene, Suenomatsuyama Fm
nobilis, Megistocrinus: Wachsmuth and Springer Plastoholotype
Wachsmuth and Springer, 1890, p. 169, pl. 16, fig. 6
Le Grand, Iowa
Mississippian, Kinderhook group
perrini, Scutella: Weaver Holotype
Weaver, 1908, p. 273, pl. 22, fig. 2
Fresno Co., Calif.; vicinity of Coalinga
“Miocene” [probably Pliocene]
sanctaecrucis, Amphiura: Arnold Holotype
Arnold, 1908b, p. 404, pl. 40, figs. 1, 2. Also in Arnold, 1909, illus. 2,
fig. 59
Santa Cruz Co., Calif.; 6 miles NNE of Santa Cruz, hills immediately
SE of Scott Valley
Upper Miocene, upper Santa Margarita Fm [Arnold’s No. 1078]
semigibbosus, Dendraster (Calaster) oregonensis: Howe Holotype
Howe, 1922, p. 102, pl. 7, fig. 3
Cape Blanco, Ore. SU loc. NP 27
Pliocene, Empire Fm
7371
482
5118
5119
3878
6005
6006
6006a
5546
STANFORD UNIVERSITY TyPEs: SMITH 513
sverdrupi, Encope: Durham Paratype
Durham, 1950, p. 48
Baja California, Mexico; Santa Inez Point, 10 miles N of Mulege
SU loc. 805
Pliocene
tapinus, Spatangus: Schenck Paratype
Schenck, 1928, p. 198, pl. 24, fig. 2
Ventura Co., Calif.; Santa Paula Qd, Timber Canyon SU loc. 277
Upper Eocene, Tejon Fm
CHORDATA
californicus, Desmostylus: Hay Holotype
Hay, 1923, p. 106. Illustrated iz Hannibal, 1922, pp. 238-240, pl. 12,
figs. 8, 9
San Jose Qd, Calif.; between Monument Peak and Milpitas-Calaveras
Miocene, San Pablo Fm [type includes fragments and worn second
or third molar]
californicus, Desmostylus: Hay Paratype
Hay, 1923, p. 106, Illustrated zz Hannibal, 1922, pp. 238-240, pl. 12,
rasee, Y/
San Jose Qd, Calif.; between Monument Peak and Milpitas-Calaveras
Miocene, San Pablo Fm
californicus, Desmostylus: Hay Paratypes
Hay, 1923, p. 106
San Jose Qd, Calif.; between Monument Peak and Milpitas-Calaveras
Miocene, San Pablo Fm
morani, Zalophancylus: Hannibal Holotype
Hannibal, 1912b, p. 152, pl. 6, fig. 15. Also im Hanna, 1925, p. 18-19
Bad Land Hills, Ore.; 1 mile E of Sand Hollow
Pliocene, Idaho Lake beds [originally described as a mollusk, recog-
nized as a fish vertebra by Hanna, 1925]
nevadanus, Helicoprion: Wheeler Plastoholotype
Wheeler, 1939, p. 109, fig. 3
Lovelock Qd, Nevada; SE 1/4, SW 1/4 Sec. 16, T 28 N, R 34 E
Anthracolithic [late Paleozoic] Rochester Fm _ [holotype 1001 Univ.
Nevada MacKay Mus. Paleontol. |
pacificus, Shastasaurus: Merriam Syntype
Merriam, 1895, p. 57, fig. 1. Also iz Merriam, 1902, p. 102, pl. 14,
figs
Shasta Co., Calif.
Triassic
pacificus, Shastasaurus: Merriam Syntype
Merriam, 1895, p. 57, fig. 2. Also in Merriam, 1908, p 143, pl. 17,
fig. 3
Shasta Co., Calif.
Triassic
pacificus, Shastasaurus: Merriam Syntype
Merriam, 1895, p. 57
Shasta Co., Calif.
Triassic
perrini, Thalattosaurus: Merriam Holotype
Merriam, 1905, p. 36, pl. 4, fig. 3; pl. 7, fig. 6
Shasta Co., Calif.; Smith Cove, near Squaw Creek
Triassic, Hosselkus Ls
514 BuL_eTIN 300
5547 perrini, Thalattosaurus: Merriam Syntype
Merriam, 1905, p 36, pl. 5, fig. 3
Shasta Co., Calif.; Smith Cove, near Squaw Creek
Triassic, Hosselkus Ls
5879 sierrensis, Helicoprion: Wheeler Plastoholotype
Wheeler, 1939, p. 112, fig. 4
Plumas Co., Calif.; Downieville Qd, SE 1/4, SE 1/4 Sec. 22, T 22 N,
R12E
Anthracolithic [late Paleozoic] [holotype 1002 Univ. Nevada Mackay
Mus. Paleontol.]
PLANTS; ORGANIC AND SILICEOUS MICROFOSSILS
9939 californicum, Plataninium: Page Holotype
Page, 1968, p. 169-170, figs. 4-6 .
Stanislaus Co, Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
1/4 Sec. 32, T 5S,R7E
Upper Cretaceous, Panoche Fm [angiosperm wood]
9944 caudatum, Tunisphaeridium: Deunff and Evitt Paratypes (4)
Deunff and Evitt, 1968, p. 4, pl. 2, fig. 4 (R 21.5, + 8.6); fig. 8 (R
128% 1°32); fies 10,11 (R 1:65 =F" 7.2)= figs 13 (Re eee
Rochester, New York; gorge of Genesee River
Middle Silurian Clinton group, Maplewood Shale _ [acritarch]
9945 caudatum, Tunisphaeridium: Deunff and Evitt Paratypes (2)
Deunff and Evitt, 1968, p. 4, pl. 2, fig. 6 (R 25.9, + 4.9); fig. 7 (R
25 320-4 1219)
Rochester, New York; gorge of Genesee River
Middle Silurian, Clinton group, Maplewood Shale _ [acritarch]
9948 caudatum, Tunisphaeridium: Deunff and Evitt Paratypes (3)
Deunff and Evitt, 1968, p. 4, pl. 2, fig. 2 (R 17.3, + 8.8); fig. 12 (R
25.6, + 5.8); fig. 14 (R 19.2, 416.3)
Rochester, New York; gorge of Genesee River
Middle Silurian, Clinton group, Maplewood Shale [acritarch]
9998 cocculoides, Lardizabaloxylon: Page Holotype
Page, 1970, p. 1139, figs. 1, 2, 8
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
1/4 Sec. 32, 758, R 7E
Upper Cretaceous, Panoche Fm [angiosperm wood ]
9944 concentricum, Tunisphaeridium: Deunff and Evitt Paratype
Deunff and Evitt, 1968, p. 3, pl. 1, fig. 7 (R 11.7, + 7.8)
Rochester, New York; gorge of Genesee River
Middle Silurian, Clinton group, Maplewood Shale [acritarch]
9948 concentricum, Tunisphaeridium: Deunff and Evitt Paratypes (7)
Deunff and Evitt, 1968, p. 3, pl. 1, fig. 3 (R 6.1, + 11.9); figs. 4, 10
(R 11.5, -- 30); fig. 5 (R 24.3, + 31); fig. 8. (R 59) 4 eee
9 (R 24.0, + 7.9); fig. 11 (L 6.3, + 6.7) ; fig. 12 (R 7.0, + 4.5)
Rochester, New York; gorge of Genesee River
Middle Silurian, Clinton group, Maplewood Shale _ [acritarch]
10100 = cretacea, Margariella: Page Paratype
Page, 1973, pp. 572-574, figs. 11, 13
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Del Puerto Canyon, SE
1/4, SW 1/4 Sec. 20, T 5S, R7E
Upper Cretaceous, Panoche Fm_ [conifer]
10077 +=cretacea, Margariella: Page Holotype
Page, 1973, pp. 572-574, figs. 1-9, 15
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Del Puerto Canyon, SE
1/4, SW 1/4 Sec. 20, T 58,R7E
Upper Cretaceous, Panoche Fm_ [conifer]
10000
10001
10079
10080
10081
10082
8360
8361
8362
8363
STANFORD UNIVERSITY Tyres: SMITH 515
cretacea, Riboidoxylon: Page Holotype
Page, 1970, pp. 1141-1142, figs. 7, 9-11
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Del Puerto Canyon, SE
1/4, SW 1/4 Sec. 20, T 5S,R7E
Upper Cretaceous, Panoche Fm [angiosperm wood]
eupomatioides, Mulleroxylon: Page Holotype
Page, 1970, p. 1143, figs. 12-14
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
1/4 Sec. 32,17 5 S,R7E
Upper Cretaceous, Panoche Fm_ [angiosperm wood |
exilimurum, Inversidinium: McLean Holotype
McLean, D., 1973, p. 730, pl. 90, figs. 1, 2 (R 26.0, + 12.5), slide
CV 53
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W, bluffs along S bank of Aquia Creek, 1/2 mile SE of Md.-Va.
Monument No. 37
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
exilimurum, Inversidinium: McLean Paratype
McLean, D., 1973, p. 730, pl. 90, figs. 3, 6 (R 14.2, + 4.3) Slide CV
18
Prince Georges Co., Md.; Anacostia, Md.-D.C. Qd, 38° 45’ 10” N,
76° 59’ 15” W, 1/2 mile W of Friendly, Md
Upper Paleocene, Aquia Fm
exilimurum, Inversidinium: McLean Paratype
McLean, D., 1973, p. 730, pl. 90, figs. 4, 5 (R 19.0, + 12.0) Slide CV
75
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W, bluffs along S bank Aquia Creek, ca. 1/2 mile SE of Md.-Va.
Monument No. 37
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
exilimurum, Inversidinium: McLean Paratype
McLean, D., 1973, p. 730, pl. 90, figs. 7-9 (R 24.3, + 1.5) Slide CV 87
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W, bluffs along S bank Aquia Creek, ca. 1/2 mile SE of Md.-Va.
Monument No. 37
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
hannae, Cyclotella: Kanaya Holotype
Kanaya, 1957, p. 82, pl. 3, fig. 10
Contra Costa Co., Calif.; Byron Qd, 2.8 miles W of town of Byron
SU loc. M-611.7
Eocene, Kellogg Shale [diatom]
hannae, Cyclotella: Kanaya Paratype
Kanaya, 1957, p. 82, pl. 3, fig. 11
Contra Costa Co., Calif.; Byron Qd, 2.8 miles W of town of Byron
SU loc. M-611.7
Eocene, Kellogg Shale [diatom]
hannae, Cyclotella: Kanaya Paratype
Kanaya, 1957, p. 82, pl. 3, fig. 12
Contra Costa Co., Calif.; Byron Qd, 2.8 miles W of town of Byron
SU loc. M-611.7
Eocene, Kellogg Shale [diatom]
hannae, Cyclotella: Kanaya Paratype
Kanaya, 1957, p. 82, pl. 3, fig. 13
Contra Costa Co., Calif.; Byron Qd, 2.8 miles W of town of Byron
SU loc. M-611.7
Eocene, Kellogg Shale [diatom]
516
7904.2
7905
7906
7904.3
7904.4
7904.5
7904.6
7904.7
7904.11
7904.1
7904.8
7904.9
7904.10
5057
BuLLETIN 300
keenae, Permopora: Elias Holotype
Elias, 1947, pp. 53-54, pl. 18, figs. 1, 11
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress; NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54
Cottle Co., Texas; NW cor. Buckle L Ranch, NW cor. Sec. 655, 6
miles S, 4 miles W of Childress
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54
Cottle Co., Texas; SW cor. Buckle L Ranch, SW cor. Sec. 661, 13
miles S, 4 miles W of Childress
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, figs. 5, 8, 9
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress; NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, figs. 8, 10
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, fig. 6
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, figs. 1, 3
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, figs. 1, 2, 4
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenae, Permopora: Elias Paratype
Elias, 1947, pp. 53-54, pl. 18, fig. 7
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments |
keenae, Permopora: Elias Paratypes
Elias, 1947, pp. 53-54
Childress Co., Texas; ca. 2.5 miles N 60° E of Childress, NW 1/4
Sec. 325, 200 yds. NE of No. 1 G. R. Cooper Well
Late Permian, Childress Dolomite [plant fragments]
keenani, Archaeolithothamnium: Howe Holotype
Howe, 1934, p. 513, pl. 54, fig. A. Also in Keenan, 1932, pl. 4, fig. 5
Santa Barbara Co., Calif.; Santa Ynez Qd, just NW of right angle
in stream on W bank of E fork Cachuma Creek, T 3 N, R 28 W. 4/5
miles W, 3/5 miles S of intersection of 34° 40’ N, 119° 50’ W SU
loc. 1106
Eocene, Sierra Blanca Ls_ [marine alga]
806
10073
8425
9567
810
807
805
9999
811
804
808
10002
STANFORD UNIVERSITY Types: SMITH 517
Lacuma sp.: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 92, pl. 35, fig. 1
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
laminosum, Lithothamnium: Howe Holotype
Howe, 1934, p. 513, pl. 55, fig. A
Santa Barbara Co., Calif.; Santa Ynez Qd, Indian Creek, at its inter-
section with Is beds 4 miles S of Big Pine Mt. SU loc. 930
Eocene, Sierra Blanca Ls_ [marine alga]
mentitum, Hystrichokolpoma: McLean Holotype
McLean, D., 1974, p. 67, pl. 8, figs. 1-5 (R 4.0, + 11.4) Slide CV 25
Prince Georges Co., Md.; Anacostia, Md.-D.C. Qd, 38° 45’ 10” N,
76° 59’ 15” W, 1/2 mile W of Friendly, Md.
Upper Paleocene, Aquia Fm_ [dinoflagellate]
nevadensis, Lyonothamnoxylon: Page Holotype
Page, 1964, pp. 257-266, 10 figs.
Esmeralda Co., Nevada; David Mt. Qd, Fish Lake Valley, 3/4 mile
S of hill 6061, T1N,R35E
Lower Pliocene [5567 is matrix from which holotype came]
oregona, Ilex: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 80, pl. 22, fig. 5
Lane Co., Ore.; 9 miles S of Goshen, E side Pacific Highway SU
loc. 36
Eocene-Oligocene, Goshen Fm
oregona, Symplocos: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 93, pl. 37, fig. 5
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
oregona, Tetracera: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 87, pl. 31, fig. 5
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
oregona, Tetracera: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 87, pl. 31, fig. 7
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
ostryopsoides: Carpinoxylon: Page Holotype
Page, 1970, p. 1139-1141, figs. 3-6
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
1/4See..32, TK SeSeRG7 E
Upper Cretaceous, Panoche Fm_ [angiosperm wood ]
ovalis, Siparuna: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 71, pl. 15, fig. 4
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
ovoidea, Ocotea: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 75, pl. 20, fig. 3
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
ovoidea, Ocotea: Chaney and Sanborn Paratype
Chaney and Sanborn, 1933, p. 75, pl. 20, fig. 1
Lane Co., Ore.; 9 miles S of Goshen SU loc. 36
Eocene-Oligocene, Goshen Fm
panochensis, Magnolioxylon: Page Holotype
Page, 1970, p. 1143, figs. 15-17
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
W/45Secs 52, ly 5) Ss Ro 7
Upper Cretaceous, Panoche Fm [angiosperm wood]
518
6855
9948
9938
5639
5641
10055
10052
10053
10054
5638
122
BuLtetin 300
panochetris, Tetracentronites: Page Holotype
Page, 1968, pp. 170-172, figs. 7-9
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Del Puerto Canyon, SE
1/4, SW 1/4 Sec. 20, T 5S, R7E
Upper Cretaceous, Panoche Fm [angiosperm wood]
parvum, Tunisphaeridium: Deunff and Evitt Holotype
Deunff and Evitt, 1968, p. 3, pl. 2, fig. 15 (R 5.3, + 10.7); pl. 2, fig.
18 (L 1.9, + 12.1)
Rochester, New York; gorge of Genesee River
Middle Silurian, Clinton group, Maplewood Shale _ [acritarch]
platanoides, Plataninium: Page Holotype
Page, 1968, pp. 168-169, figs. 1-3
Stanislaus Co., Calif.; Patterson 7 1/2’ Qd, Black Gulch, NE 1/4, SE
1/4 Sec. 32, T5S,R7E
Upper Cretaceous, Panoche Fm
schenckii, Mesophyllum: Howe Holotype
Howe, 1934, p. 512, pl. 52, fig. E
Santa Barbara Co., Calif.; Santa Ynez Qd, where Indian Creek inter-
sects ls beds 4 miles S of Big Pine Mt. SU loc. 930
Eocene, Sierra Blanca Ls
schenckii, Mesophylilum: Howe Paratype
Howe, 1934, p. 512, pl. 52, figs. A, B
Santa Barbara Co., Calif.; Santa Ynez Qd, where Indian Creek inter-
sects ls beds 4 miles S of Big Pine Mt. SU loc. 930
Eocene, Sierra Blanca Ls
schenckii, Mesophyllum: Howe Paratype
Howe, 1934, p. 512, pl. 52, tig. C
Santa Barbara Co., Calif.; Santa Ynez Qd, where Indian Creek inter-
sects Is beds 4 miles S of Big Pine Mt. SU loc. 930
Eocene, Sierra Blanca Ls
septatum, Cladopyxidium: McLean Holotype
McLean, D., 1972, p. 862, pl. 1, figs. 5-8 (R 18.2, + 9.9) Slide CV 83
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
septatum, Cladopyxidium: McLean Paratype
McLean, D., 1972, p. 862, pl. 1, fig. 11( R 29.3, + 2.7) Slide CV 28
Prince Georges Co., Md.; Anacostia, Md.-D.C. Qd, 38° 45’ 10” N,
76° 59’ 15” W, 1/2 mile W of Friendly, Md.
Upper Paleocene, Aquia Fm_ [dinoflagellate]
septatum, Cladopyxidium: McLean Paratype
McLean, D., 1972, p. 862, pl. 1, fig. 12 (R 19.2, + 2.0) Slide CV 41
Prince Georges Co., Md.; Anacostia, Md.-D.C. Qd, 38° 45’ 10” N,
76° 59’ 15” W, 1/2 mile W of Friendly, Md.
Upper Paleocene, Aquia Fm, lowermost part
septatum, Cladopyxidium: McLean Paratype
McLean, D., 1972, p. 862, pl. 1, figs. 1-3 (R 27.1, + 1.0) Slide CV 42
Prince Georges Co., Md.; Anacostia, Md.-D.C. Qd, 38° 45’ 10” N,
76° 59’ 15” W, 1/2 mile W of Friendly, Md.
Upper Paleocene, Aquia Fm, lowermost part
sierrablancae, Lithophyllum: Howe Holotype
Howe, 1934, p. 514, pl. 56, fig. A, as sterra-blancae
Santa Barbara Co., Calif.; Santa Ynez Qd, where Indian Creek inter-
sects ls beds 4 miles § of Big Pine Mt. SU loc. 930
Eocene, Sierra Blanca Ls_ [marine alga]
steamboatea, Nilsonia: Reagan Holotype
Reagan, 1925, p. 141, fig. 1c
18 miles W of Ganado, Arizona; Steamboat Canyon
Cretaceous, Dakota Ss_ [leaf]
STANFORD UNIVERSITY | yPEs: SMITH 519
10075 tumescens, Hystrichokolpoma: McLean Holotype
McLean, 1974, p. 66, pl. 8, figs. 7, 8 (R 12.5, + 4.7) Slide CW 14
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W, bluffs along § bank Aquia Creek, ca. 1/2 mile SE of Md.-Va.
Monument No. 37
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
10074 tumescens, Hystrichokolpoma: McLean Paratype
McLean, D., 1974, p. 66, pl. 8, fig. 6 (R 21.7, + 16.4) Slide CW 50
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 22’ 15” N, 77° 17’
50” W, bluffs along § bank Aquia Creek, ca. 1/2 mile SE of Md.-Va.
Monument No. 37
Upper Paleocene, Aquia Fm, type section [dinoflagellate]
10076 tumescens, Hystrichokolpoma: McLean Paratype
McLean, D., 1974, p. 66, pl. 8, fig. 9 (R 3.7, + 8.7) Slide CW 63
Stafford Co., Va.; Passapatanzy, Va.-Md. Qd, 38° 20’ 35” N, 77° 17’
17” W, bluffs along S bank Potomac Creek, from .05-.15 mile W of
Md.-Va. Monument No. 35
Upper Paleocene, Aquia Fm_ [dinoflagellate]
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22
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1940a (Mar.) Nine new snails of the genus Monadenia from California.
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1948 (Feb.). Two misunderstood West American chitons. Leaflets in
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1860. On some new species of fossils from the limestone near Point
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1861. On some of the rocks and fossils occurring near Phillipsburgh,
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524 BULLETIN 300
1962 (April). New species of Oliva from west Mexico. Nautilus, vol.
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1899a
1899b
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1908a
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1913
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LI.
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LXVII.
LXVIII.
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