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The University of Maine
Studies
No- 4
A Catalogue and Bibliography of the
Odonata (Dragon-flies) of Maine^
With an Annotated List of their Collectors
BY
FRANCES LEROY HARVEY
Professor of Natural History
ORONO, MAINE
August, )902
The University of Maine
Studies
No* 4
A Catalogue and Bibliography of the
Odonata (Dragon-flies) of Maine^
With an Annotated List of their Collectors
BY
FRANCES LEROY HARVEY
Professor of Natural History
ORONO, MAINE
August, J902
Most of the papers left by the late Professor Harvey have
been published in the Bnfomologica! A^ezvs. The local interest
attaching to this paper made its publication in Maine desirable.
The proof has been read by L. H. Harvey, of the class of 1901.
A CATALOGUE AND BIBLIOGRAPHY OF THE
ODONATA (DRAGON-FLIES) OF MAINE, WITH
AN ANNOTATED LIST OF THEIR COLLECTORS
SUBFAMILY I. CALOPTERYCxIN^.
I. CALOPTERYX LEACH.
T. Calopteryx maailata Beauv.
Maine ( Proc. B. S. N. H. 15:273; no authority, but
probably Packard) ; Manchester (Miss Wadsworth) ;
Orono, Bradley, S. Lagrange, Foxcroft, N. E. Carry
(Harvey) ; Farmington (Miss Furbish) ; Mt. Desert
(Bullock).
2. C. aequabilis Say.
Maine (Proc. B. S. N. H. 15: 274, no authority);
Orono, Bradley, S. Lagrange, Foxcroft (Harvey) ;
Manchester (Miss Wadsworth).
2. HET^^RINA IIAGEN.
3. Hetcerina Americana Fabr.
Norway (Smith) ; Bradley, 1897, Auburn, 1898 (Mer-
rill) ; Winslow, 1899 (Hitchings) ; Chemo Mills
(Harvey).
SUBFAMILY II. AGRIONIN^.
3. LESTES LEACH.
4. Lestes congener Hagen.
Orono, 1899; N. W. Carry, 1898 (Harvey) ; Manchester
1898 (Miss Wadsworth).
5. L. iinguiculatns Hagen.
Orono, N. W. Carry (Harvey) ; Manchester (Miss
Wadsworth) ; York Harbor (Moore).
6. L. iincatus Kirby.
Orono (Bartle & F. L. Harvey) ; Manchester (Miss
Wadsworth).
3
7- /,. disjunct us Selys.
Manchester (Miss Wadsworth) ; Orono, Fryeburg
(Harvey).
8. L. forcipatns Rambur.
Orono, Fryeburg (Harvey) ; Manchester (Miss Wads-
worth).
9. L. rectangularis Say.
Orono, Fryeburg (Harvey) ; Manchester (!Miss Wads-
worth).
10. L. vigilax Selys.
Manchester, (Miss Wadsworth) ; Fryeburg (Harvey).
II; L. ineqiialis Walsh.
Orono, Bradley (Harvey) ; Manchester (Miss Wads-
worth).
4. ARGIA RAMBIIR.
12. Ai'gia ptiirida Hagen.
Orono and many other localities (Harvey). Manchester
(]\Iiss Wadsworth).
13. A. violacea Hagen.
Orono, Bradley, etc. (Harvey) ; Manchester (Miss
Wadsworth; Mt. Desert (Bullock).
14. A. apicalis Say.
Orono, (Harvey).
5. KRYTHROMMA CHARPEXTlER.
15. Brythromma conditum Hagen,
Manchester (Miss W.) ; Bradley & Orono (Harvey).
6. NEHALKNNIA SELYS.
16. Nehalennia Irene, Hagen.
Maine (Packard) ; Manchester (Miss Wadsworth) ;
Orono, Fryeburg (Harvey); York Harbor (Moore).
17. N. posit a Hagen,
Orono, Penobscot River 1897 ( Bartle Harvey).
7. AMPHIAGRION SELYS.
18. Amphiagrion smichim Burmeister.
Maine (Packard); Manchester (Miss Wadsworth);
Orono (Bartle Harvey) ; Bradley (Harvey).
8. ENALLAGMA CHARPENTlER.
19. Bnallagiua civile Hagen.
Orono (Harvey) ; Manchester (Miss Wadsworth) ; Y.-^rk
Harbor (Moore).
20. B. ehriuhi Hagen,
Manchester (Miss Wadsworth).
21. £. cyathigerum Charp., race annexum Hagen,
Manchester (Miss Wadsworth).
22. B. Calverti, Morse,
Orono 1897 (Florence Harvey) ; 1898 (Harvey).
23. B. Hageiii Walsh,
Orono, Fryebiirg (Harvey) ; Manchester (Miss Wads-
worth) ; Mt. Desert (Bullock) ; York Harbor
(Moore).
24. B. geminatum Kellicott,
Bradley, Chemo Mills (Harvey).
25. B. signatiDn Hagen,
Bradley (Harvey) ; Manchester Miss Wadsworth).
26. B. pollntum Hagen,
Chemo stream, Bradley (Harvey) ; ^lanchester (Miss
Wadsworth ) .
27. B. exsulans Hagen,
Manchester, 1891 (Miss Wadsworth) ; Orono, 1898
(Harvey).
Q. ISCHNURA CHARPENTlER.
28. Ischnura verticalis Say.
Maine. 1836 (Randall; Proc. B. S. N. H. 15:276)
Orono, Fryeburg (Harvey) ; Manchester (Miss Wads-
worth) ; Mt. Desert (Bullock) ; York Harbor
(Moore).
29. /. Ramburii Selys.
Manchester (Miss Wadsworth).
10. ANOMALAr,RION SELYS.
30. Anomalgrion hastatuni Say.
Maine. Hagen's synopsis p. yy (Packard). Has not
been re-discovered.
SUBFAMILY III. GOMPHINyE.
II. HAGENIUS SELYS.
31. Ha genius hreinstyhis Selys.
Bradley, Orono, Greenfield, N. E. Carry (Harvey) ;
Manchester (Miss Wadsworth).
12. OPIIIOGOMPHUS SELYS.
32. Ophiogomphiis rupuunlensis Walsh.
Maine (Packard, Proc. Acad. Nat. Sci. Phila. 1862, p,
388. Also by Morrison, teste Selys 1878).
33. O. aspersus Morse.
Russell Stream, Aug. 1899 (Harvey).
34. O. carohis Needham.
Orono 5 (Florence Harvey) J' (Ned Mansfield) ; Green-
field, Bradley (Harvey).
35. 0. mainensis Packard.
Maine (Packard. Proc. Ent. Soc. Phila. 1S63, p. 255).
36. 0. anomalus Harvey.
Orono (Harvey).
2i7. O. johannus Needham.
Chemo Mill, Bradley, 1899 (Harvey).
13. GOMPHUS LEACH.
38. GompJiiis iiccvius Hagen.
Chemo Stream, Bradley, a single $ 1890, (Harvey);
1897-9 Many males and a few females (F. L. & Bartle
Harvey).
39. G. parvulus. Selys.
Maine (Hag. Syn. Odon. of Am. p. 46, 1875 ' "o author-
ity) ; Manchester (Miss Wadsworth).
40. G. abbreviatus Hagen.
Orono (Harvey) ; Manchester (Miss Wadsworth, teste
P. P. Cahert).
41. G. brezis Selys.
Orono (Harvey).
42. G. exilis Selys.
Orono (Harvey) ; Manchester (Miss Wadsworth) ; Mt.
Desert (Bullock).
43- (j- spicatus Selys.
S. Lagrange. Orono, Oldtown (Harvey) ; Manchester
(Miss Wadsworth).
44. G. Scudderi Selys.
Russell stream near N. E. Carry, one male, 1897, many
males and females, Aug. 25-29, 1899 (Harvey).
14. DROMOGOMPHUS SELYS.
45. Dromogomphus spinosus, Selys.
Manchester (Miss Wadsworth) ; Old Hown, Pushaw
Pond, Aug. 16, 1898, a single J* (Harvey).
SUBFAMILY IV. CORDULEGASTERIN^..
15. cordule:gastkr leach.
46. Cordnlegaster mactilafns Selys,
Greenfield, Orono (Harvey) ; Manchester (Miss Wads-
worth).
47. C. Sayi Selys.
Portland (Packard, Hag. Syn. O. of A. p. 50, 1875).
48. C. diastatops Selys.
Manchester (Miss Wadsworth) ; Orono, Greenfield,
Bradley (Harvey).
49. C. (Taeniogaster) obliqiius, Say.
Orono, Me. (Packard, see Proc. B. S. N. H. 16: 356) ;
Manchester (Miss Wadsworth); Orono & Bradley
(Harvey).
SUBFAMILY V. AESCHIN^.
16. EPLAESHNA SELYS.
50. Epiaeschna heros Fabricius.
Maine (Proc. B. S. N. H. 15: 272, no authority) ; 1899
(Harvey).
17. BOYERL\ MCLACHLAN.
51. Boyeria z'inosa Sa.y.
Maine (Proc. B. S. N. H. 15:272, no authority);
Orono, Greenfield, N. E. Carry, etc. (Harvey) ; Man-
chester (Miss Wadsworth).
1 8. GOMPH.^SCHNA SELYS.
52. Gomphceschna furcillata Say.
Manchester (Miss Wadsworth).
19. BASI.^SCHNA SELYS.
53. Basiceschna Janata Say.
Orono, Bradley (Harvey) ; Manchester (Miss Wads-
worth).
20. .^SCHNA FABRTCIUS.
54. Mschna jiincea var rerticalis Hagen.
Maine (Smith? Hagen's Syn. Od. of A. p. 34) ; Man-
chester (Miss Wadsworth) ; Orono, Bradley (Harvey).
55. Ai. clepsydra Say.
Maine (Proc. B. S. N. H. 15: 271, 1873, no authority).
Orono, S. Lagrange in ponds, Bradley, Greenfield, etc.
(Harvey).
56. Ai constricia Say.
Maine (Smith? Synopsis Od. of A. Hagen p. 34) ; Man-
chester (j\Iiss Wadsworth) ; Orono, N. E. Carry,
Kineo, Fryeburg, etc. (Harvey).
21. AN AX LEACH.
57. Anax Junius Drury.
Orono (Harvey) ; Manchester (Miss Wadsworth).
SUBFAMILY VI. CORDULIN^.
22. DIDYMOPS RAMBUR.
58. Didymops transversa Say.
Manchester (Miss Wadsworth) ; Orono (Harvey).
23. MACROMIA RAMBUR.
59. Macromia illinoensis Walsh.
Manchester (Miss Wadsworth) ; Orono, Bradley (Har-
vey).
24. CORDULL\ LEACH.
60. Cordnlia shurtleHi vScudd.
Orono (Bartle Harvey) ; Bradley (Harvey).
8
25- EPICORDULIA SEIYS.
6i. Bpicordnlia princeps Hagen.
Manchester (Miss Wadsworth).
26. TETRAGONKtJRIA HAGEN.
62. Tetragonenria cynosura Say.
Maine (Hag. Syn. O. of A. 1875, P- ^O' "o authority) ;
Bradley (Harvey) ; Manchester (Miss Wadsworth).
63. T. spinigera Selys.
Manchester (Miss Wadsworth) ; Bradley (Harvey).
64. T. semiaqnca Bnrmeister.
Manchester (Miss Wadsworth) ; Mt. Desert (Bullock).
2y. NEUROCORDULIA SELYS.
65. Neurocordnlia uhleri Selys.
Orono (Packard, Hag. Syn. O. of A. 1875, p. 61) ; Orono
Harvey) ; Manchester (Miss Wadsworth).
66. A^. obsoleta Say.
Manchester, July 5th (Miss Wadsworth).
67. N. yamaskanensis Provancher.
Orono, 1808 r? (L. H. Harvey) : ^(Harvey), several in
1899 (Harvey).
28. SOMATOCHLORA SELYS.
68. Somatochlora elongata Scudd.
Coast of Maine at sea (Harvey).
69. S. elongata var. minor Calvert.
Over Penobscot River, Orono, Bradley (Harvey).
70. vS. linearis Hagen, ( Probably ) .
Manchester, July 27, 1897 (Miss Wadsworth).
71. S. forcipata Scudd.
Maine (Randall, 1836. Proc. B. S. N, H. 15:268);
Orono, Bradley (Harvey); Manchester (Miss Wads-
worth),
y2. S. sepfentrionalis Hagen.
Orono, 6-18-1898 (Bartle Harvey). Over swamp.
73. S. zvalskii Scudd.
Manchester (Miss Wadsworth) ; Orono (L. H. Har-
vey) ; Bradley (Harvey).
9
74- S. lepida. Selys.
Portland (Jones and probably Randall. Proc. B. S. N.
. H. 15: 270); Greenfield, Bradley, Orono (Harvey);
Manchester (Miss Wadsworth).
75. S. libera Selys.
Orono (Bartle Harvey) ; Greenfield (Harvey) ; Man-
chester (^liss Wadsworth).
SUBFAMILY VII. LIBELLULIN^.
29. LIBELLULA LINNE).
76. Libel hila exits fa Say.
Norway (Smith. Hag. Syn. O. of A. 1875 P- 69). ^lan-
chester (]Miss Wadsworth).
yy. ' L. incesta Hagen.
Manchester (Miss Wadsworth).
78. L. 4-maciilata Linne.
Manchester (Miss Vv^adsworth) ; Orono, Bradley (Har-
vey) ; Mt. Desert (Bnllock) Miller) (var. pranubila
Newman. Orono (Harvey).
79. L. pidchcUa Drnry.
Brmiswick (Proc. B. S. N. H. 16: 367) ; Orono, Bradley
(Harvey) ; Manchester (^liss Wadsworth).
80. L. semifasciata Burmeister.
Manchester (Miss Wadsworth) ; Orono, June 1898
(Harvey).
30. PLATHEMIS HAGKN.
81. Plathcmis irimaculata De Geer.
?vlaine (Hag. Syn. O. of A. 1875 P- ^7> "^ authority) ;
Orono, Bradley (Harvey) ; Manchested (Wiss W^ads-
worth) ; Mt. Desert (Bullock) ; Bar Harbor (Miller).
31. NANNOTHEMIS BRAUER.
82. Nannotheuiis bella IJhler.
Maine (Packard) : Mt. Desert (Bullock).
},2. CELITHEMIS HAGEN.
83. CeUtheniis ornata Rambur.
Manchester (Miss Wadsworth) ; Bradley (Bartle Har-
vey).
10
84. C. elisa Hagen.
Bradley (Harvey); Manchester (Miss Wadsworth) ;
Mt. Desert (Bullock).
33. LEUCORHINIA BRITTINGER.
85. Leucorhinia intacta Hagen.
Orono, Bradley (Harvey); Manchester (Miss Wads-
worth ) .
86. L. hiidsonica Selys.
Oldtown, Orono (Harvey).
87. L. proxima Calvert.
Manchester (Miss Wadsworth) ; Old Town, Orono
(Harvey).
88. L. frigida Hagen.
Bradlev (Harvev & Bar tie Harvey) ; Mt. Desert (Bul-
. lock).
89. L. glacialis.
Orono, Bradley, Chemo Stream (Harvey).
34. SYMPE:TRUM NEWMAN.
90. Sympctrum riibicundulum Say.
Maine (Proc. B. S. N. H. 15:267, no authority);
AFanchester (Miss Wadsworth) ; N. W. Carry, South
Lagrange, Greenfield, Bradley, Orono, Fryeburg, etc..
(Harvey) ; York Harbor (Moore).
Greenfield, Bradley, Orono, Fryeburg, etc. (Harvey) ;
York Harbor (Moore).
91. S. obstrusum Hagen.
York Harbor (Moore) ; Fryeburg (Harvey).
92. S. semicincttim Say.
Maine (Randall. Proc. B. S. N. H. 15: 267); Orono,
Bradley (Harvey) ; Manchester (Miss Wadsworth) ;
York Harbor (Moore).
93. S. vicinum Hagen.
Brunswick ( Packard Hag. Syn. O. of A. p. 80) ; ?^lan-
chester (Miss Wadsworth) ; Orono, Bradley, Frye-
burg (Harvey) ; York Harbor (Moore).
94. S. costifenim Uhler.
Maine (Hag. Syn. O. of A. p. 81, no authority) : Orono,
Bradley (Harvey); Manchester (Miss Wadsworth);
York Harbor (Moore).
TI
BIBLIOGRAPHY.
BULLOCK, DAVID JAYNE.
I. Notes on Mt. Desert Dragon-flies. Ent. News, 2: 93.
1891.
CULVERT, PHILIP P.
1. Notes on N. Am. Odonata. Trans. Am. Ent. Soc. 17:
33-40. 1890. Maine species collected by Miss
Wadsworth at Manchester are considered.
2. Data on the Distribution of Dragon-flies (Odonata) i.
Ent. News, 5 : October, 1894. Mentions papers on
Odonates by Miss Wadsworth, and Mr. D. J. Bullock,
F. L. Harvey, besides giving lists of species collected
at Mt. Desert by Bullock, and at York Harbor by J. P.
Moore.
3. Catalogue of the Odonata of the Vicinity of Phila., Pa.
Trans. Am. Ent. Soc. 20: 1893. Maine species men-
tioned for geographical distribution.
HAGEN, HERMAN A.
1. Synopsis of the Neuroptera of North America, Smith-
sonian Institution, Washington, D. C. 1861. Mentions
species collected in Maine by Prof. A. S. Packard.
2. On the Neuroptera in Harris' Collection. Proc. Bost.
Soc. Nat. Hist, 15: 1873. Ten species from Maine
are mentioned, three of which are credited to John Witt
Randall, one to K. T. Jones, and the others are without
authority.
3. Odonate Fauna of Georgia. Proc. Bost. Soc. Nat. Hist.,
16: 1874. Two Maine species mentioned, one credited
to Packard and the other without authority.
4. Synopsis of the Odonata of America. Proc. Bost. Soc.
Nat. Hist. 18: 20-96, 1875. Twenty-one species
are mentioned including sev^eral recorded in previous
papers. For several species only the State is given and
for all no authority. Orono, Brunswick, Norway, and
Portland are localities mentioned and would suggest
Packard, Smith, Jones, Morse, and probably Randall
as the collectors.
12
5- Synopsis of the Odonata of North America No. I.
Psyche, 5: 241. 1889. Two Maine species mentioned.
6. Synopsis of the Odonate Genus Leucorhinia Britt. Trans.
Am. Ent. Soc. 22: 229-230. 1890. Two Maine species
mentioned in notes by Mr. Calvert.
7. Notes and Descriptions of some North American Libellu-
Hna. Psyche 5 : 383. 1890. One species from Maine
mentioned as collected by Smith or L. Agassiz.
HARVKY, FRANCIS LEIROY.
1. A contribution to the Odonata of Maine. Ent. News, 2:
50, 73. 1891.
2. Contributions to the Odonata of Maine — II. Ent. News,
3 : 91, 116. 1892.
3. Contributions to the Odonata of Maine — III. Ent. News,
9:59,85. 1898.
4. Contributions to the Odonata of Maine — IV. Ent. News,
12: 1901.
KIRBY, w. F.
I. A Synonymic Catalogue of Odonata or Dragon-flies, Lon-
don, 1890. Mentions some species from Maine.
WADS WORTH, MISS MATTlE.
1. List of the Dragon-flies (Odonata) taken at Manchester,
Kennebec Co., Me., in 1888 and 1889. Ent. News.
1 : 36, 55. 1890.
2. Additions and Corrections to the above. Ent. News, 2:
II. 1891.
3. Second Additions and corrections to the above. Ent.
News 3 : 8. 1892.
4. Third Additions to the above. Ent. News 9: iii. 1898.
5. Fourth Addition to the above. Ent. News, ix, p. iii,
1898.
WALSH, BENJAMIN D.
I. Observations on certain N. A. Neuroptera by H. Hagen,
M. D. Translated from the original French MS and
published by permission of the author with notes and
descriptions of about twenty new N. A. species of
Pseudoneuroptera. Proc. Ent. Soc. Phila., 2: 107-
2^2. 1863 Ophiogomphns Mainensis, Packard des-
cribed p. 255.
13
Collectors of AJaiue Odonatcs.
Randall, Dr. John Witt. Earliest collector of insects in ]Maine,
1836-40. Specimens sent to Thomas W. Harris and credited
to Randall in his MS catalogue. He collected coleoptera mostly,
but also insects of other orders, inchiding a few Odonates. He
published a paper ''Description of New Coleoptera from Maine."
(Bost. Jour. Nat. Hist. A'^ol. 2.)
Packard, Prof. Alphens Spring. Began gathering insects in
Maine when only 16 years old; collected during his college
course 1858-61, and until 1864 at Brunswick mostly. Also at
Orono and in Northern Maine while on the Geological Survey,
August and September 1861. Collected in all orders. Uhler
named his Odonates. Specimens went first to the Peabody
Academy of Science and afterwards to the Cambridge Museum.
Author of many articles bearing upon the Insects of Maine.
Morse, Prof. Bdzvard Sylvester. Collected in the vicinity of
Portland, Maine from 1856-62 in all orders. Specimens went
to Museum of Comparative Zoology, Cambridge, Mass.
Smith, Prof. S. I. Made large collections in all orders at
Norway, Maine. His specimens are now in the Museum of
Comparative Zoology, Cambridge, Mass. Prof. Smith is the
author of a paper, **Orthoptera of Maine." published in the Proc.
Portland Soc'y Nat. Hist. Vol. I.
Agassis, Prof. Louis. Is credited with an Odonate collected
at Bethel, Maine.
Verrill, Prof. A. B. May have collected insects in Elaine at
Norway between 1858-61 ; if so, they probably went to the Cam-
bridge Museum.
Jones, Thomas K., Portland. An amateur collector particu-
larly of beetles. He donated a fine collection of ]\Iaine beetles
to the Portland Society of Natural History. The present secre-
tary writes, "They have gone the way of all insect collections
not having daily care." Odonates were collected somewhat and
the specimens, we think, went to the Cambridge Museum, as
Mr. Jones is mentioned by Dr. Hagen. Mr. Jones is a sign
14
painter by trade and has collected insects many years. Has been
an active member of the Portland Society of Natural History
and once its secretary.
Morrison, H. K. Is credited with collecting Ophioa^omphns
rapinsidensis in Maine. (»See Selys Quatriemes Additions au
Synopsis des Gomphines, Brussels, 1878: 50.)
Fernald, Prof, C. H. Orono and vicinity, while connected
with the Maine State College from 1871-86. Specimens in col-
lection of University of Maine, as reported by the present writer
in Entomological News, are without collector's name, but we
presume were collected by Prof. Fernald or his pupils.
Wadsworth, Mattie B. Manchester, Kennebec Co. from 1890
to date. Results published in Entomological News. (See Bib-
liography).
Harvey, Prof. F . L. Orono, Me. and various localities in
the Penobscot valley 1886- 1899. Results published in Entomo-
logical News, (see Bibliography.)
Moore, Dr. J. Percy. Instructor of Zoology, University of
Pennsylvania, Philadelphia. Collected at York Harbor, Maine,
in summer of 1890. Specimens given to Mr. P. P. Calvert and
reported by him in Ent. News. 5: 242. 1894. (See Biblio-
graphy).
Bullock, David Jaync. Lives {1901) in Philadelphia, Pa.
Collected in vicinity of Bar Harbor, in 1890. Specimens reported
by collector.
Specimens collected in 1891 were reported by Mr. P. P. Cal-
vert in Ent. News, 5 : 242, 1894. (See Bibliography.)
Collections made by the following persons have been examined
by the writer and reported by him : Mr. and Mr^ Anson Allen,
Orono; Miss Kate Furbish, Rangeley region; IV^r. E. D. Mer-
rill, Auburn, Monmouth and Orono ; P. L. Ricker, Westbrook ;
LeRoy H. Harvey, Bartle F. Harvey, Miss Florence Harvey,
and Willis E. Harvey, Orono, Bradley, Greenfield, and other
localities in the Penobscot valley.
15
We know of Maine Odonates in the collections of other Ento-
mologists of the state, but have not examined them critically.
Miss Mattie Wadsworth, David Jayne Bullock and the writer
are the only collectors of Odonates of Maine v^ho have published
their results.
16
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